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<document id="72D41413235BE35E10BAC90297059E08" ID-DOI="10.11646/zootaxa.5501.4.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="13745322" ID-ZooBank="C4509093-8845-45A6-AB97-3710B300830C" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726050899385" checkinUser="plazi" docAuthor="Makarchenko, Eugenyi A., Semenchenko, Alexander A., Palatov, Dmitry M. &amp; Lisanovskaya, Elena A." docDate="2024" docId="03A387EEFF963B0F21D7232FFB0FFF44" docLanguage="en" docName="zootaxa.5501.4.2.pdf" docOrigin="Zootaxa 5501 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5501.4.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thalassomya paraskevae Makarchenko, Semenchenko et Palatov 2024, sp. nov." docType="treatment" docUuid="ED222392-FC41-44C4-BCA1-0E74C1AB7640" docUuidSource="ZooBank" docVersion="2" lastPageNumber="528" masterDocId="FF9AFF96FF973B0B21402011FFA2FFBD" masterDocTitle="Morphological description and DNA barcoding of Thalassomya paraskevae sp. nov. (Diptera: Chironomidae: Telmatogetoninae) from coast of the Black Sea" masterLastPageNumber="530" masterPageNumber="524" pageNumber="525" updateTime="1726051295610" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C7C01B3BCB5AF1B5741E8AAC26FFAC38">Morphological description and DNA barcoding of Thalassomya paraskevae sp. nov. (Diptera: Chironomidae: Telmatogetoninae) from coast of the Black Sea</mods:title>
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<mods:namePart id="FDD644E54FCC92590B7F216382835367">Makarchenko, Eugenyi A.</mods:namePart>
<mods:affiliation id="D010FB164DB91CC1B82957008CDF4819">Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far East Branch of the Russian Academy of Sciences, 100 let Vladivostoku 159, 690022 Vladivostok, Russia.</mods:affiliation>
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<mods:namePart id="9E2C59CEB0CB5F8F9ACFFC0F08FA2D89">Semenchenko, Alexander A.</mods:namePart>
<mods:affiliation id="D8535C4C3E963B163A9696EBDE9CF2E6">Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far East Branch of the Russian Academy of Sciences, 100 let Vladivostoku 159, 690022 Vladivostok, Russia.</mods:affiliation>
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<mods:namePart id="73146642C88CAC86857779D2016295EA">Palatov, Dmitry M.</mods:namePart>
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<mods:namePart id="1FCD99E050411EBA254AD24ACF187CDF">Lisanovskaya, Elena A.</mods:namePart>
<mods:affiliation id="2E2C909A919080A00D94B5F7CE22C0DF">Far Eastern Federal University, Institute of the World Ocean, 10 Ajax Bay, Russky Island, 690922 Vladivostok, Russia.</mods:affiliation>
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<mods:date id="EB9E945055C73CF710F130AEBF792F5A">2024</mods:date>
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<emphasis id="B97EEAEAFF963B0A21D7232FFE67FCE4" bold="true" box="[151,453,830,857]" italics="true" pageId="1" pageNumber="525">Thalassomya paraskevae</emphasis>
Makarchenko, Semenchenko
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Palatov
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,
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http://zoobank.org/NomenclaturalActs/
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(
<figureCitation id="13312A7DFF963B0A21DF2396FF4BFC1C" box="[159,233,903,929]" captionStart="FIGURES 18" captionStartId="2.[152,270,1792,1816]" captionTargetBox="[151,1435,179,1766]" captionTargetId="figure-18@2.[151,1435,179,1766]" captionTargetPageId="2" captionText="FIGURES 18. Adult male of Thalassomya paraskevae sp. nov. 1, head; 2, antenna; 3, wing; 4, claws of mid leg; 5, claws of hind leg; 6, endomeres and phallapodemes; 78, hypopygium, in dorsal view; 9, part of hypopygium, in ventral view; 10, dorsobasal lobe of gonocoxite; 11, gonostylus." figureDoi="http://doi.org/10.5281/zenodo.13745324" httpUri="https://zenodo.org/record/13745324/files/figure.png" pageId="1" pageNumber="525">Figs. 1</figureCitation>
11)
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<paragraph id="8BB536F8FF963B0A21D723DEFDA1FBC4" blockId="1.[151,1437,974,1253]" pageId="1" pageNumber="525">
<emphasis id="B97EEAEAFF963B0A21D723DEFEE7FC55" bold="true" box="[151,325,974,1001]" pageId="1" pageNumber="525">Type material.</emphasis>
<materialsCitation id="3B623CA5FF963B0A200D23DEFBEDFBB0" collectingDate="2024-01" collectorName="D. Palatov." latitude="43.08334" location="Gudautra District" longLatPrecision="1" longitude="40.81668" municipality="New Athos Town" pageId="1" pageNumber="525" specimenCount="1" typeStatus="holotype">
<typeStatus id="54B1885AFF963B0A200D23DEFE1AFC54" box="[333,440,975,1001]" pageId="1" pageNumber="525" type="holotype">Holotype</typeStatus>
, adult male, ABKHAZIA:
<location id="8ED56023FF963B0A23A723DEFC0CFC54" LSID="urn:lsid:plazi:treatment:03A387EEFF963B0F21D7232FFB0FFF44:8ED56023FF963B0A23A723DEFC0CFC54" box="[743,942,975,1001]" latitude="43.08334" longLatPrecision="1" longitude="40.81668" municipality="New Athos Town" name="Gudautra District" pageId="1" pageNumber="525">Gudautra District</location>
,
<collectingMunicipality id="6BD1AC82FF963B0A22F923DEFB22FC54" box="[953,1152,975,1001]" pageId="1" pageNumber="525">New Athos Town</collectingMunicipality>
, breakwater near the pier on the beach of the village, 01.0
<date id="FFB41038FF963B0A234923E2FDF2FBB0" box="[521,592,1011,1037]" pageId="1" pageNumber="525" value="2024-01">
<collectingDate id="EFF0E9D0FF963B0A234923E2FDF2FBB0" box="[521,592,1011,1037]" pageId="1" pageNumber="525" value="2024-01">I.2024</collectingDate>
</date>
,
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,
<geoCoordinate id="EE3E503FFF963B0A224023E2FC30FBB0" box="[768,914,1011,1037]" degrees="40.816681" direction="east" orientation="longitude" pageId="1" pageNumber="525" precision="1" value="40.81668">40.816681 E</geoCoordinate>
, leg.
<collectorName id="26FF532EFF963B0A229123E2FBEDFBB0" box="[977,1103,1011,1037]" pageId="1" pageNumber="525">D. Palatov.</collectorName>
</materialsCitation>
<materialsCitation id="3B623CA5FF963B0A251623E2FE3BFB8C" collectingDate="2024-01" collectorName="D. Palatov." latitude="43.08334" location="Gudautra District" longLatPrecision="1" longitude="40.81668" municipality="New Athos Town" pageId="1" pageNumber="525" specimenCount="6" specimenCount-adult="6" typeStatus="paratype">
<typeStatus id="54B1885AFF963B0A251623E2FB65FBB0" box="[1110,1223,1011,1037]" pageId="1" pageNumber="525" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D0CFD71FF963B0A259323E2FA83FBB0" box="[1235,1313,1011,1037]" count="6" pageId="1" pageNumber="525" type="adult">6 adult</specimenCount>
males, the same data as holotype
</materialsCitation>
;
<materialsCitation id="3B623CA5FF963B0A20E62406FDA1FBC4" collectingDate="2024-04-25" collectorName="D. Palatov." country="Russia" county="Territory" latitude="44.132805" location="Cape Kadosh" longLatPrecision="1" longitude="39.026054" municipality="Tuapse District" pageId="1" pageNumber="525" specimenCount="3" specimenCount-adult="3" stateProvince="Krasnodar" typeStatus="paratype">
<specimenCount id="9D0CFD71FF963B0A20E62406FE57FB8C" box="[422,501,1047,1073]" count="3" pageId="1" pageNumber="525" type="adult">3 adult</specimenCount>
males,
<collectingCountry id="F31D7668FF963B0A23102406FD1AFB8C" box="[592,696,1047,1073]" name="Russia" pageId="1" pageNumber="525">RUSSIA</collectingCountry>
:
<collectingRegion id="49CEF81AFF963B0A23852406FC98FB8C" box="[709,826,1047,1073]" country="Russia" name="Krasnodar" pageId="1" pageNumber="525">Krasnodar</collectingRegion>
<collectingCounty id="62D44E74FF963B0A22022406FC04FB8C" box="[834,934,1047,1073]" pageId="1" pageNumber="525">Territory</collectingCounty>
,
<collectingMunicipality id="6BD1AC82FF963B0A22F32406FBC1FB8C" box="[947,1123,1047,1073]" pageId="1" pageNumber="525">Tuapse District</collectingMunicipality>
,
<location id="8ED56023FF963B0A25312406FAA8FB8C" LSID="urn:lsid:plazi:treatment:03A387EEFF963B0F21D7232FFB0FFF44:8ED56023FF963B0A25312406FAA8FB8C" box="[1137,1290,1047,1073]" country="Russia" county="Territory" latitude="44.132805" longLatPrecision="1" longitude="39.026054" municipality="Tuapse District" name="Cape Kadosh" pageId="1" pageNumber="525" stateProvince="Krasnodar">Cape Kadosh</location>
cliffs on the
<location id="8ED56023FF963B0A21D7242AFEA5FBE8" LSID="urn:lsid:plazi:treatment:03A387EEFF963B0F21D7232FFB0FFF44:8ED56023FF963B0A21D7242AFEA5FBE8" box="[151,263,1083,1109]" country="Russia" county="Territory" latitude="44.132805" longLatPrecision="1" longitude="39.026054" municipality="Tuapse District" name="Black Sea" pageId="1" pageNumber="525" stateProvince="Krasnodar">Black Sea</location>
coast, near the village of
<location id="8ED56023FF963B0A2364242AFDC1FBE8" LSID="urn:lsid:plazi:treatment:03A387EEFF963B0F21D7232FFB0FFF44:8ED56023FF963B0A2364242AFDC1FBE8" box="[548,611,1083,1109]" country="Russia" county="Territory" latitude="44.132805" longLatPrecision="1" longitude="39.026054" municipality="Tuapse District" name="Agoy" pageId="1" pageNumber="525" stateProvince="Krasnodar">Agoy</location>
, south of the mouth of the river of the same name,
<date id="FFB41038FF963B0A25DA242BFAB9FBE8" box="[1178,1307,1082,1109]" pageId="1" pageNumber="525" value="2024-04-25">
<collectingDate id="EFF0E9D0FF963B0A25DA242BFAB9FBE8" box="[1178,1307,1082,1109]" pageId="1" pageNumber="525" value="2024-04-25">25.04.2024</collectingDate>
</date>
,
<geoCoordinate id="EE3E503FFF963B0A2465242AFF0CFBC4" degrees="44.132806" direction="north" orientation="latitude" pageId="1" pageNumber="525" precision="1" value="44.132805">44.132806 N</geoCoordinate>
,
<geoCoordinate id="EE3E503FFF963B0A21F9244FFEE8FBC4" box="[185,330,1118,1145]" degrees="39.026056" direction="east" orientation="longitude" pageId="1" pageNumber="525" precision="1" value="39.026054">39.026056 E</geoCoordinate>
, leg.
<collectorName id="26FF532EFF963B0A20C7244EFDA1FBC4" box="[391,515,1119,1145]" pageId="1" pageNumber="525">D. Palatov.</collectorName>
</materialsCitation>
</paragraph>
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<subSubSection id="C3106573FF963B0A21872493FD2BFB58" pageId="1" pageNumber="525" type="etymology">
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<emphasis id="B97EEAEAFF963B0A21872493FE06FB21" bold="true" box="[199,420,1154,1180]" pageId="1" pageNumber="525">Derivatio nominis.</emphasis>
The species is named in honour of Bulgarian chironomidologist Dr. Paraskeva Mikhailova (Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, Sofia) who studied
<taxonomicName id="4C0A4D7BFF963B0A244824B6FEB0FB58" authorityName="Schiner" authorityYear="1856" class="Insecta" family="Chironomidae" genus="Thalassomya" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="525" phylum="Arthropoda" rank="species" species="frauenfeldi">
<emphasis id="B97EEAEAFF963B0A244824B6FEB0FB58" italics="true" pageId="1" pageNumber="525">Thalassomya frauenfeldi</emphasis>
</taxonomicName>
from Bulgarias Black Sea coast.
</paragraph>
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<subSubSection id="C3106573FF963B0821D72523FE4EFC26" lastPageId="3" lastPageNumber="527" pageId="1" pageNumber="525" type="description">
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<emphasis id="B97EEAEAFF963B0A21D72523FE8AFAF1" bold="true" box="[151,296,1330,1356]" pageId="1" pageNumber="525">Description</emphasis>
</paragraph>
<paragraph id="8BB536F8FF963B0A21D7256BFAD0FA28" blockId="1.[151,1437,1402,2041]" box="[151,1394,1402,1429]" pageId="1" pageNumber="525">
<emphasis id="B97EEAEAFF963B0A21D7256BFEB4FA28" bold="true" box="[151,278,1402,1429]" italics="true" pageId="1" pageNumber="525">Adult male</emphasis>
(n = 5, except when otherwise stated). Total length
<quantity id="4CF29B1DFF963B0A221B256AFC47FA28" box="[859,997,1403,1429]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" metricValueMax="4.2" metricValueMin="3.4" pageId="1" pageNumber="525" unit="mm" value="3.8" valueMax="4.2" valueMin="3.4">3.44.2 mm</quantity>
. Total length/wing length 0.981.0.
</paragraph>
<paragraph id="8BB536F8FF963B0A21D7258EFC8BFA04" blockId="1.[151,1437,1402,2041]" box="[151,809,1439,1465]" pageId="1" pageNumber="525">Coloration. Dark brown. Wings grey, with brownish veins.</paragraph>
<paragraph id="8BB536F8FF963B0A218725D2FB96F940" blockId="1.[151,1437,1402,2041]" pageId="1" pageNumber="525">
Head (
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). Eyes bare, rounded, without dorsomedial projections. Temporal setae strong and numerous— 75108. Clypeus with 4958 strong setae. Antenna with scape covered
<emphasis id="B97EEAEAFF963B0A22F625F6FC72F9BD" box="[950,976,1511,1536]" italics="true" pageId="1" pageNumber="525">ca</emphasis>
30 strong setae and 6 flagellomeres with reduced plume of setae (
<figureCitation id="13312A7DFF963B0A20EF261AFE57F998" box="[431,501,1547,1573]" captionStart="FIGURES 18" captionStartId="2.[152,270,1792,1816]" captionTargetBox="[151,1435,179,1766]" captionTargetId="figure-18@2.[151,1435,179,1766]" captionTargetPageId="2" captionText="FIGURES 18. Adult male of Thalassomya paraskevae sp. nov. 1, head; 2, antenna; 3, wing; 4, claws of mid leg; 5, claws of hind leg; 6, endomeres and phallapodemes; 78, hypopygium, in dorsal view; 9, part of hypopygium, in ventral view; 10, dorsobasal lobe of gonocoxite; 11, gonostylus." figureDoi="http://doi.org/10.5281/zenodo.13745324" httpUri="https://zenodo.org/record/13745324/files/figure.png" pageId="1" pageNumber="525">Fig. 2</figureCitation>
); number and length of these setae on 15 flagellomeres respectively: 56 (4468 μm), 4 (6068 μm), 4 (4452 μm), 4 (4856 μm), 4 (4468 μm); terminal flagellomere with 4 setae, 4860 μm long in basal part and with 1 subapical setae, 3240 μm long. Length of 16 flagellomeres (μm): 96112, 5256, 4452, 4048, 4044, 96120; terminal flagellomere with a distinct nipple-like apex. AR 0.350.39. Palpomere length (μm): 4042, 3244, 8088, 136188, 208252. Palpomere 1 without setae, palpomeres 23 with setae 6896 μm long, palpomeres 45 with setae 2432 μm long; palpomere
<quantity id="4CF29B1DFF963B0A220226AEFCCFF964" box="[834,877,1727,1753]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="1" pageNumber="525" unit="in" value="3.0">3 in</quantity>
distal part with sensilla capitata with diameter 16 μm. Head width/palpal length 1.121.26. Antennal length/palpal length 0.740.75.
</paragraph>
<paragraph id="8BB536F8FF963B0A21872716FF75F830" blockId="1.[151,1437,1402,2041]" pageId="1" pageNumber="525">Thorax. Antepronotum with 3740 ventrolateral setae, 148256 μm long. Acrostichals 3032, 104128 μm long, in anterior part in 1 row, in posterior part in 2 rows; dorsocentrals 2339, 156 μm long; humerals 56, 100104 μm long; prealars 1621, 100112 μm long (in 2 groups). Scutellum with 3034 setae. Preepisternum without setae.</paragraph>
<paragraph id="8BB536F8FF963B0A21872786FF72F844" blockId="1.[151,1437,1402,2041]" pageId="1" pageNumber="525">
Wing (
<figureCitation id="13312A7DFF963B0A20542786FEF8F80C" box="[276,346,1943,1969]" captionStart="FIGURES 18" captionStartId="2.[152,270,1792,1816]" captionTargetBox="[151,1435,179,1766]" captionTargetId="figure-18@2.[151,1435,179,1766]" captionTargetPageId="2" captionText="FIGURES 18. Adult male of Thalassomya paraskevae sp. nov. 1, head; 2, antenna; 3, wing; 4, claws of mid leg; 5, claws of hind leg; 6, endomeres and phallapodemes; 78, hypopygium, in dorsal view; 9, part of hypopygium, in ventral view; 10, dorsobasal lobe of gonocoxite; 11, gonostylus." figureDoi="http://doi.org/10.5281/zenodo.13745324" httpUri="https://zenodo.org/record/13745324/files/figure.png" pageId="1" pageNumber="525">Fig. 3</figureCitation>
). Length
<quantity id="4CF29B1DFF963B0A20872786FDCFF80C" box="[455,621,1943,1969]" metricMagnitude="-3" metricUnit="m" metricValue="3.76" metricValueMax="4.08" metricValueMin="3.44" pageId="1" pageNumber="525" unit="mm" value="3.76" valueMax="4.08" valueMin="3.44">3.444.08 mm</quantity>
, width
<quantity id="4CF29B1DFF963B0A23812786FCCAF80C" box="[705,872,1943,1969]" metricMagnitude="-3" metricUnit="m" metricValue="1.18" metricValueMax="1.28" metricValueMin="1.08" pageId="1" pageNumber="525" unit="mm" value="1.18" valueMax="1.28" valueMin="1.08">1.081.28 mm</quantity>
. R
<subScript id="178E34BDFF963B0A22C627B8FC3DF80A" attach="left" box="[902,927,1961,1975]" fontSize="6" pageId="1" pageNumber="525">2+3</subScript>
absent. R and R
<subScript id="178E34BDFF963B0A251B27B8FBC1F80A" attach="left" box="[1115,1123,1961,1975]" fontSize="6" pageId="1" pageNumber="525">1</subScript>
with 6268 setae, R
<subScript id="178E34BDFF963B0A240C27B8FAC4F80A" attach="left" box="[1356,1382,1961,1975]" fontSize="6" pageId="1" pageNumber="525">4+5</subScript>
with 3538 setae. fCu well beyond base of r-m. Anal lobe rounded-angular. Squama with 3145 setae, 100120 μm long.
</paragraph>
<caption id="DF756670FF953B0921D82711FDCDF8DD" ID-DOI="http://doi.org/10.5281/zenodo.13745324" ID-Zenodo-Dep="13745324" httpUri="https://zenodo.org/record/13745324/files/figure.png" pageId="2" pageNumber="526" startId="2.[152,270,1792,1816]" targetBox="[151,1435,179,1766]" targetPageId="2" targetType="figure">
<paragraph id="8BB536F8FF953B0921D82711FDCDF8DD" blockId="2.[151,1437,1792,1888]" pageId="2" pageNumber="526">
<emphasis id="B97EEAEAFF953B0921D82711FEE0F8A5" bold="true" box="[152,322,1792,1816]" pageId="2" pageNumber="526">FIGURES 18.</emphasis>
Adult male of
<taxonomicName id="4C0A4D7BFF953B0920A02711FD43F8A5" authorityName="Makarchenko &amp; Semenchenko &amp; Palatov &amp; Lisanovskaya" authorityYear="2024" box="[480,737,1792,1816]" class="Insecta" family="Chironomidae" genus="Thalassomya" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="526" phylum="Arthropoda" rank="species" species="paraskevae" status="sp. nov.">
<emphasis id="B97EEAEAFF953B0920A02711FD43F8A5" box="[480,737,1792,1816]" italics="true" pageId="2" pageNumber="526">Thalassomya paraskevae</emphasis>
</taxonomicName>
<emphasis id="B97EEAEAFF953B0923A92711FCF4F8A5" bold="true" box="[745,854,1792,1816]" pageId="2" pageNumber="526">
<taxonomicNameLabel id="A24D5791FF953B0923A92711FC99F8A5" box="[745,827,1792,1816]" pageId="2" pageNumber="526" rank="species">sp. nov.</taxonomicNameLabel>
1,
</emphasis>
head;
<emphasis id="B97EEAEAFF953B0922DD2711FC0DF8A5" bold="true" box="[925,943,1792,1816]" pageId="2" pageNumber="526">2,</emphasis>
antenna;
<emphasis id="B97EEAEAFF953B0925552711FB8AF8A5" bold="true" box="[1045,1064,1792,1816]" pageId="2" pageNumber="526">3,</emphasis>
wing;
<emphasis id="B97EEAEAFF953B0925312711FB26F8A5" bold="true" box="[1137,1156,1792,1816]" pageId="2" pageNumber="526">4,</emphasis>
claws of mid leg;
<emphasis id="B97EEAEAFF953B0924082711FAF9F8A5" bold="true" box="[1352,1371,1792,1816]" pageId="2" pageNumber="526">5,</emphasis>
claws of hind leg;
<emphasis id="B97EEAEAFF953B0920572735FE88F881" bold="true" box="[279,298,1828,1852]" pageId="2" pageNumber="526">6,</emphasis>
endomeres and phallapodemes;
<emphasis id="B97EEAEAFF953B09233A2735FD04F881" bold="true" box="[634,678,1828,1852]" pageId="2" pageNumber="526">78,</emphasis>
hypopygium, in dorsal view;
<emphasis id="B97EEAEAFF953B09229E2735FC53F881" bold="true" box="[990,1009,1828,1852]" pageId="2" pageNumber="526">9,</emphasis>
part of hypopygium, in ventral view;
<emphasis id="B97EEAEAFF953B09243D2735FA3EF881" bold="true" box="[1405,1436,1828,1852]" pageId="2" pageNumber="526">10,</emphasis>
dorsobasal lobe of gonocoxite;
<emphasis id="B97EEAEAFF953B0920952759FE51F8DD" bold="true" box="[469,499,1864,1888]" pageId="2" pageNumber="526">11,</emphasis>
gonostylus.
</paragraph>
</caption>
<paragraph id="8BB536F8FF943B0821872086FC3BFF44" blockId="3.[151,1437,151,249]" pageId="3" pageNumber="527">
Legs. Spur of front tibia 80100 µm long. Spurs of mid tibia 7680 µm and 96120 µm long. Spurs of hind tibia 92 µm and 92100 µm long. Anterior tarsal claws on front and middle legs pectinate at tip, other claws sharp (
<figureCitation id="13312A7DFF943B08242420AAFF64FF44" captionStart="FIGURES 18" captionStartId="2.[152,270,1792,1816]" captionTargetBox="[151,1435,179,1766]" captionTargetId="figure-18@2.[151,1435,179,1766]" captionTargetPageId="2" captionText="FIGURES 18. Adult male of Thalassomya paraskevae sp. nov. 1, head; 2, antenna; 3, wing; 4, claws of mid leg; 5, claws of hind leg; 6, endomeres and phallapodemes; 78, hypopygium, in dorsal view; 9, part of hypopygium, in ventral view; 10, dorsobasal lobe of gonocoxite; 11, gonostylus." figureDoi="http://doi.org/10.5281/zenodo.13745324" httpUri="https://zenodo.org/record/13745324/files/figure.png" pageId="3" pageNumber="527">Figs. 45</figureCitation>
). Length (μm) and proportions of leg segments are as in
<tableCitation id="C6880343FF943B08227F20CEFC31FF44" box="[831,915,223,249]" captionStart="TABLE 1" captionStartId="3.[151,239,332,356]" captionText="TABLE 1. Lengths (in µm) and proportions of leg segments of Thalassomya paraskevae sp. nov., male (n=4)" pageId="3" pageNumber="527">Table 1</tableCitation>
.
</paragraph>
<caption id="DF756670FF943B0821D7215DFA89FED8" box="[151,1323,332,357]" pageId="3" pageNumber="527" startId="3.[151,239,332,356]" targetType="table">
<paragraph id="8BB536F8FF943B0821D7215DFA89FED8" blockId="3.[151,1323,332,357]" box="[151,1323,332,357]" pageId="3" pageNumber="527">
<emphasis id="B97EEAEAFF943B0821D7215DFEA8FED9" bold="true" box="[151,266,332,356]" pageId="3" pageNumber="527">TABLE 1.</emphasis>
Lengths (in µm) and proportions of leg segments of
<taxonomicName id="4C0A4D7BFF943B08227F215DFBEEFED9" authorityName="Makarchenko &amp; Semenchenko &amp; Palatov &amp; Lisanovskaya" authorityYear="2024" box="[831,1100,332,356]" class="Insecta" family="Chironomidae" genus="Thalassomya" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="527" phylum="Arthropoda" rank="species" species="paraskevae" status="sp. nov.">
<emphasis id="B97EEAEAFF943B08227F215DFBEEFED9" box="[831,1100,332,356]" italics="true" pageId="3" pageNumber="527">Thalassomya paraskevae</emphasis>
</taxonomicName>
<emphasis id="B97EEAEAFF943B082513215CFB0AFED8" bold="true" box="[1107,1192,333,357]" pageId="3" pageNumber="527">
<taxonomicNameLabel id="A24D5791FF943B082513215CFB0AFED8" box="[1107,1192,333,357]" pageId="3" pageNumber="527" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male (n=4)
</paragraph>
</caption>
<paragraph id="8BB536F8FF943B0821BD218AFA2BFD90" pageId="3" pageNumber="527">
<table id="F90AC458FF94C4F421E6218AFA2BFD8F" box="[166,1417,411,562]" gridcols="12" gridrows="4" pageId="3" pageNumber="527">
<tr id="353A34BAFF94C4F421E6218AFA2BFE08" box="[166,1417,411,437]" gridrow="0" pageId="3" pageNumber="527" rowspan-0="1">
<th id="76EB5DC6FF94C4F42195218AFE95FE08" box="[213,311,411,437]" gridcol="1" gridrow="0" pageId="3" pageNumber="527">fe</th>
<th id="76EB5DC6FF94C4F42013218AFE17FE08" box="[339,437,411,437]" gridcol="2" gridrow="0" pageId="3" pageNumber="527">ti</th>
<th id="76EB5DC6FF94C4F42091218AFD91FE08" box="[465,563,411,437]" gridcol="3" gridrow="0" pageId="3" pageNumber="527">ta1</th>
<th id="76EB5DC6FF94C4F4230D218AFD3BFE08" box="[589,665,411,437]" gridcol="4" gridrow="0" pageId="3" pageNumber="527">ta2</th>
<th id="76EB5DC6FF94C4F423F2218AFD5CFE08" box="[690,766,411,437]" gridcol="5" gridrow="0" pageId="3" pageNumber="527">ta3</th>
<th id="76EB5DC6FF94C4F42257218AFCC0FE08" box="[791,866,411,437]" gridcol="6" gridrow="0" pageId="3" pageNumber="527">ta4</th>
<th id="76EB5DC6FF94C4F4223E218AFC6BFE08" box="[894,969,411,437]" gridcol="7" gridrow="0" pageId="3" pageNumber="527">ta5</th>
<th id="76EB5DC6FF94C4F422AA218AFBE3FE08" box="[1002,1089,411,437]" gridcol="8" gridrow="0" pageId="3" pageNumber="527">LR</th>
<th id="76EB5DC6FF94C4F42521218AFB1AFE08" box="[1121,1208,411,437]" gridcol="9" gridrow="0" pageId="3" pageNumber="527">BV</th>
<th id="76EB5DC6FF94C4F42593218AFA88FE08" box="[1235,1322,411,437]" gridcol="10" gridrow="0" pageId="3" pageNumber="527">SV</th>
<th id="76EB5DC6FF94C4F42408218AFA2BFE08" box="[1352,1417,411,437]" gridcol="11" gridrow="0" pageId="3" pageNumber="527">BR</th>
</tr>
<tr id="353A34BAFF94C4F421E621D2FA2BFE63" box="[166,1417,451,478]" gridrow="1" pageId="3" pageNumber="527">
<th id="76EB5DC6FF94C4F421E621D2FF18FE63" box="[166,186,451,478]" gridcol="0" gridrow="1" pageId="3" pageNumber="527">P 1</th>
<td id="76EB5DC6FF94C4F4219521D2FE95FE63" box="[213,311,451,478]" gridcol="1" gridrow="1" pageId="3" pageNumber="527">1960-2080</td>
<td id="76EB5DC6FF94C4F4201321D2FE17FE63" box="[339,437,451,478]" gridcol="2" gridrow="1" pageId="3" pageNumber="527">2240-2360</td>
<td id="76EB5DC6FF94C4F4209121D2FD91FE63" box="[465,563,451,478]" gridcol="3" gridrow="1" pageId="3" pageNumber="527">1240-1320</td>
<td id="76EB5DC6FF94C4F4230D21D2FD3BFE63" box="[589,665,451,478]" gridcol="4" gridrow="1" pageId="3" pageNumber="527">508-525</td>
<td id="76EB5DC6FF94C4F423F221D2FD5CFE63" box="[690,766,451,478]" gridcol="5" gridrow="1" pageId="3" pageNumber="527">295-328</td>
<td id="76EB5DC6FF94C4F4225721D2FCC0FE63" box="[791,866,451,478]" gridcol="6" gridrow="1" pageId="3" pageNumber="527">98-120</td>
<td id="76EB5DC6FF94C4F4223E21D2FC6BFE63" box="[894,969,451,478]" gridcol="7" gridrow="1" pageId="3" pageNumber="527">197-213</td>
<td id="76EB5DC6FF94C4F422AA21D2FBE3FE63" box="[1002,1089,451,478]" gridcol="8" gridrow="1" pageId="3" pageNumber="527">0.45-0.55</td>
<td id="76EB5DC6FF94C4F4252121D2FB1AFE63" box="[1121,1208,451,478]" gridcol="9" gridrow="1" pageId="3" pageNumber="527">4.69-4.99</td>
<td id="76EB5DC6FF94C4F4259321D2FA88FE63" box="[1235,1322,451,478]" gridcol="10" gridrow="1" pageId="3" pageNumber="527">3.39-3.81</td>
<td id="76EB5DC6FF94C4F4240821D2FA2BFE63" box="[1352,1417,451,478]" gridcol="11" gridrow="1" pageId="3" pageNumber="527">2.0-2.2</td>
</tr>
<tr id="353A34BAFF94C4F421E621FCFA2BFDB5" box="[166,1417,493,520]" gridrow="2" pageId="3" pageNumber="527">
<th id="76EB5DC6FF94C4F421E621FCFF18FDB5" box="[166,186,493,520]" gridcol="0" gridrow="2" pageId="3" pageNumber="527">P2</th>
<td id="76EB5DC6FF94C4F4219521FCFE95FDB5" box="[213,311,493,520]" gridcol="1" gridrow="2" pageId="3" pageNumber="527">2680-2920</td>
<td id="76EB5DC6FF94C4F4201321FCFE17FDB5" box="[339,437,493,520]" gridcol="2" gridrow="2" pageId="3" pageNumber="527">2640-2720</td>
<td id="76EB5DC6FF94C4F4209121FCFD91FDB5" box="[465,563,493,520]" gridcol="3" gridrow="2" pageId="3" pageNumber="527">1000-1040</td>
<td id="76EB5DC6FF94C4F4230D21FCFD3BFDB5" box="[589,665,493,520]" gridcol="4" gridrow="2" pageId="3" pageNumber="527">360-410</td>
<td id="76EB5DC6FF94C4F423F221FCFD5CFDB5" box="[690,766,493,520]" gridcol="5" gridrow="2" pageId="3" pageNumber="527">262-280</td>
<td id="76EB5DC6FF94C4F4225721FCFCC0FDB5" box="[791,866,493,520]" gridcol="6" gridrow="2" pageId="3" pageNumber="527">98-120</td>
<td id="76EB5DC6FF94C4F4223E21FCFC6BFDB5" box="[894,969,493,520]" gridcol="7" gridrow="2" pageId="3" pageNumber="527">160-213</td>
<td id="76EB5DC6FF94C4F422AA21FCFBE3FDB5" box="[1002,1089,493,520]" gridcol="8" gridrow="2" pageId="3" pageNumber="527">0.37-0.39</td>
<td id="76EB5DC6FF94C4F4252121FCFB1AFDB5" box="[1121,1208,493,520]" gridcol="9" gridrow="2" pageId="3" pageNumber="527">6.48-6.96</td>
<td id="76EB5DC6FF94C4F4259321FCFA88FDB5" box="[1235,1322,493,520]" gridcol="10" gridrow="2" pageId="3" pageNumber="527">5.15-5.64</td>
<td id="76EB5DC6FF94C4F4240821FCFA2BFDB5" box="[1352,1417,493,520]" gridcol="11" gridrow="2" pageId="3" pageNumber="527">2.0-2.1</td>
</tr>
<tr id="353A34BAFF94C4F421E62206FA2BFD8F" box="[166,1417,535,562]" gridrow="3" pageId="3" pageNumber="527">
<th id="76EB5DC6FF94C4F421E62206FF18FD8F" box="[166,186,535,562]" gridcol="0" gridrow="3" pageId="3" pageNumber="527">P3</th>
<td id="76EB5DC6FF94C4F421952206FE95FD8F" box="[213,311,535,562]" gridcol="1" gridrow="3" pageId="3" pageNumber="527">2800-3000</td>
<td id="76EB5DC6FF94C4F420132206FE17FD8F" box="[339,437,535,562]" gridcol="2" gridrow="3" pageId="3" pageNumber="527">3080-3160</td>
<td id="76EB5DC6FF94C4F420912206FD91FD8F" box="[465,563,535,562]" gridcol="3" gridrow="3" pageId="3" pageNumber="527">1360-1440</td>
<td id="76EB5DC6FF94C4F4230D2206FD3BFD8F" box="[589,665,535,562]" gridcol="4" gridrow="3" pageId="3" pageNumber="527">600-705</td>
<td id="76EB5DC6FF94C4F423F22206FD5CFD8F" box="[690,766,535,562]" gridcol="5" gridrow="3" pageId="3" pageNumber="527">360-377</td>
<td id="76EB5DC6FF94C4F422572206FCC0FD8F" box="[791,866,535,562]" gridcol="6" gridrow="3" pageId="3" pageNumber="527">115-120</td>
<td id="76EB5DC6FF94C4F4223E2206FC6BFD8F" box="[894,969,535,562]" gridcol="7" gridrow="3" pageId="3" pageNumber="527">213-230</td>
<td id="76EB5DC6FF94C4F422AA2206FBE3FD8F" box="[1002,1089,535,562]" gridcol="8" gridrow="3" pageId="3" pageNumber="527">0.43-0.48</td>
<td id="76EB5DC6FF94C4F425212206FB1AFD8F" box="[1121,1208,535,562]" gridcol="9" gridrow="3" pageId="3" pageNumber="527">5.28-5.72</td>
<td id="76EB5DC6FF94C4F425932206FA88FD8F" box="[1235,1322,535,562]" gridcol="10" gridrow="3" pageId="3" pageNumber="527">4.17-4.38</td>
<td id="76EB5DC6FF94C4F424082206FA2BFD8F" box="[1352,1417,535,562]" gridcol="11" gridrow="3" pageId="3" pageNumber="527">2.2-2.5</td>
</tr>
</table>
</paragraph>
<paragraph id="8BB536F8FF943B0821872294FDB0FCCA" blockId="3.[151,1437,645,2040]" pageId="3" pageNumber="527">
Hypopygium (
<figureCitation id="13312A7DFF943B08202D2294FE1DFD22" box="[365,447,645,671]" captionStart="FIGURES 18" captionStartId="2.[152,270,1792,1816]" captionTargetBox="[151,1435,179,1766]" captionTargetId="figure-18@2.[151,1435,179,1766]" captionTargetPageId="2" captionText="FIGURES 18. Adult male of Thalassomya paraskevae sp. nov. 1, head; 2, antenna; 3, wing; 4, claws of mid leg; 5, claws of hind leg; 6, endomeres and phallapodemes; 78, hypopygium, in dorsal view; 9, part of hypopygium, in ventral view; 10, dorsobasal lobe of gonocoxite; 11, gonostylus." figureDoi="http://doi.org/10.5281/zenodo.13745324" httpUri="https://zenodo.org/record/13745324/files/figure.png" pageId="3" pageNumber="527">Figs. 7</figureCitation>
11). Tergite IX reduced, with 2529 setae in two groups. Laterosternite IX with 10 11 setae. Gonocoxite 304308 µm long, broad at base, rounded laterally, abruptly narrowed on distal half; dorsobasal lobe triangle rounded or rounded, with 45 setae 3240 µm long and in subapical part with group of very short setae; on ventral side at base with a patch of strong and long (112128 µm) setae. Gonostylus 144156 µm long, broadest medially, tapering apically to 2 setae, 2444 µm long, with some setae 7288 µm along the outer edge. Endomere yellow and with setae, 68 µm long in distal part. Phallapodeme dark brown, 184188 µm long. HR 1.932.17.
</paragraph>
<paragraph id="8BB536F8FF943B0821872390FE4EFC26" blockId="3.[151,1437,645,2040]" box="[199,492,896,923]" pageId="3" pageNumber="527">
<emphasis id="B97EEAEAFF943B0821872390FEA6FC26" bold="true" box="[199,260,897,923]" italics="true" pageId="3" pageNumber="527">Pupa</emphasis>
and
<emphasis id="B97EEAEAFF943B08207A2391FED7FC26" bold="true" box="[314,373,896,923]" italics="true" pageId="3" pageNumber="527">larva</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3106573FF943B08218723B4FBE6F8DA" pageId="3" pageNumber="527" type="diagnosis">
<paragraph id="8BB536F8FF943B08218723B4FEADFAD2" blockId="3.[151,1437,645,2040]" pageId="3" pageNumber="527">
<emphasis id="B97EEAEAFF943B08218723B4FE5BFC02" bold="true" box="[199,505,933,959]" pageId="3" pageNumber="527">Diagnosis and comments.</emphasis>
Adult male total length
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. Eyes bare, without dorsomedial projections. Temporal setae strong and numerous. Clypeus with 4958 strong setae. Antenna with 6 flagellomeres and reduced plume of setae; terminal flagellomere with a distinct nipple-like apex; AR 0.350.39. Palpomere
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distal part with sensilla capitata. Head width/palpal length 1.121.26. Antennal length/palpal length 0.740.75. Antepronotum with 3740 ventrolateral setae. Acrostichals 3032, in anterior part in 1 row, in posterior part in 2 rows; dorsocentrals 2339; humerals 56; prealars 1621 (in 2 groups). Scutellum with 3034 setae. Preepisternum without setae. Wing length
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, R
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absent. Anal lobe rounded-angular. Squama with 3145 setae. Anterior tarsal claws on front and middle legs pectinate at tip, other claws sharp. LR
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0.450.55, BV
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4.694.99, SV
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3.393.81. Tergite IX reduced, with 2529 setae in two groups. Gonocoxite broad at base, rounded laterally, abruptly narrowed on distal half; dorsobasal lobe triangle rounded or rounded, with 45 setae and in subapical part with group of very short setae; on ventral side at base with a patch of strong and long setae. Gonostylus broadest medially, tapering apically to 2 setae, with some setae along outer edge. Endomere yellow and with setae in distal part. HR 1.932.17.
</paragraph>
<paragraph id="8BB536F8FF943B0821872568FBE6F8DA" blockId="3.[151,1437,645,2040]" pageId="3" pageNumber="527">
As noted above, the new species is most closely related to
<taxonomicName id="4C0A4D7BFF943B082223256BFC58FA2E" authorityName="Schiner" authorityYear="1856" box="[867,1018,1401,1427]" class="Insecta" family="Chironomidae" genus="Thalassomya" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="527" phylum="Arthropoda" rank="species" species="frauenfeldi">
<emphasis id="B97EEAEAFF943B082223256BFC58FA2E" box="[867,1018,1401,1427]" italics="true" pageId="3" pageNumber="527">T. frauenfeldi</emphasis>
</taxonomicName>
and is poorly distinguished from the latter in morphological features. The matter is further complicated by the fact that
<taxonomicName id="4C0A4D7BFF943B082518258FFB53FA0A" authorityName="Schiner" authorityYear="1856" box="[1112,1265,1437,1463]" class="Insecta" family="Chironomidae" genus="Thalassomya" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="527" phylum="Arthropoda" rank="species" species="frauenfeldi">
<emphasis id="B97EEAEAFF943B082518258FFB53FA0A" box="[1112,1265,1437,1463]" italics="true" pageId="3" pageNumber="527">T. frauenfeldi</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="EF9B4B09FF943B0821FB25D0FEC6FA61" author="Schiner, J. B." box="[187,356,1473,1500]" pageId="3" pageNumber="527" pagination="215 - 224" refId="ref4207" refString="Schiner, J. B. (1856) Anmerkung zu dem im Bande V, pag. 13 dieser Verhandlungen abgedruckten aufsatze Frauenfeld's Beitrag zur Insekten-Geschichte. Verhandlungen des zoologisch-botanischen Vereins in Wien, 6, 215 - 224." type="journal article" year="1856">Schiner (1856)</bibRefCitation>
from adult female and for comparison we can use only the description by
<bibRefCitation id="EF9B4B09FF943B0825F625D0FAE9FA66" author="Wirth, W. W." box="[1206,1355,1473,1499]" pageId="3" pageNumber="527" pagination="117 - 139" refId="ref4410" refString="Wirth, W. W. (1947) Notes on the genus Thalassomyia Schiner, with description of two new species (Diptera: Tendipedidae). Proceedings of the Hawaiian entomological Society, 13, 117 - 139." type="journal article" year="1947">Wirth (1947)</bibRefCitation>
, which is based on an adult male, a pupa and a larva from San Remo in
<collectingCountry id="F31D7668FF943B0822C825F4FC1FFA42" box="[904,957,1509,1535]" name="Italy" pageId="3" pageNumber="527">Italy</collectingCountry>
. This description is not complete enough. However, it can be noted that structure of the male hypopygium of these two species do not differ as well as value of AR, wing venation, shape of terminal flagellomere and some other features. There are differences in the number of ventrolateral setae on the antepronotum and the values of LR, BV and SV. Thus, a male
<taxonomicName id="4C0A4D7BFF943B0825DC2643FA99F9D6" authorityName="Makarchenko &amp; Semenchenko &amp; Palatov &amp; Lisanovskaya" authorityYear="2024" box="[1180,1339,1617,1644]" class="Insecta" family="Chironomidae" genus="Thalassomya" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="527" phylum="Arthropoda" rank="species" species="paraskevae" status="sp. nov.">
<emphasis id="B97EEAEAFF943B0825DC2643FB0EF9D6" box="[1180,1196,1618,1643]" italics="true" pageId="3" pageNumber="527">T</emphasis>
.
<emphasis id="B97EEAEAFF943B0825FB2640FA99F9D6" box="[1211,1339,1617,1643]" italics="true" pageId="3" pageNumber="527">paraskevae</emphasis>
</taxonomicName>
<emphasis id="B97EEAEAFF943B0824022640FA3EF9D6" bold="true" box="[1346,1436,1617,1643]" pageId="3" pageNumber="527">
<taxonomicNameLabel id="A24D5791FF943B0824022640FA3EF9D6" box="[1346,1436,1617,1643]" pageId="3" pageNumber="527" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
with 3740 ventrolateral setae on the antepronotum, LR
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0.450.55, BV
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4.694.99, SV
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3.393.81, while male
<taxonomicName id="4C0A4D7BFF943B0821D7268BFE8DF90E" authorityName="Schiner" authorityYear="1856" box="[151,303,1689,1715]" class="Insecta" family="Chironomidae" genus="Thalassomya" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="527" phylum="Arthropoda" rank="species" species="frauenfeldi">
<emphasis id="B97EEAEAFF943B0821D7268BFE8DF90E" box="[151,303,1689,1715]" italics="true" pageId="3" pageNumber="527">T. frauenfeldi</emphasis>
</taxonomicName>
with 810 ventrolateral setae, LR
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0.57, BV
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4.53, SV
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3.25. It is also possible that there will be a different number of acrostichals, dorsocentrals and other setae of mesonotum but unfortunately there is no such data in the description of
<taxonomicName id="4C0A4D7BFF943B0820F126F3FD40F946" authority="(Wirth 1947)" baseAuthorityName="Wirth" baseAuthorityYear="1947" box="[433,738,1761,1787]" class="Insecta" family="Chironomidae" genus="Thalassomya" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="527" phylum="Arthropoda" rank="species" species="frauenfeldi">
<emphasis id="B97EEAEAFF943B0820F126F3FDE5F946" box="[433,583,1761,1787]" italics="true" pageId="3" pageNumber="527">T. frauenfeldi</emphasis>
(
<bibRefCitation id="EF9B4B09FF943B08231526F0FD7BF946" author="Wirth, W. W." box="[597,729,1761,1787]" pageId="3" pageNumber="527" pagination="117 - 139" refId="ref4410" refString="Wirth, W. W. (1947) Notes on the genus Thalassomyia Schiner, with description of two new species (Diptera: Tendipedidae). Proceedings of the Hawaiian entomological Society, 13, 117 - 139." type="journal article" year="1947">Wirth 1947</bibRefCitation>
)
</taxonomicName>
. Most likely, we are dealing with cryptic species, which can be reliably distinguished only by molecular methods. The use of this method will also allow in the future to analyze the species of
<taxonomicName id="4C0A4D7BFF943B08204D2738FE03F8FE" authorityName="Makarchenko &amp; Semenchenko &amp; Palatov &amp; Lisanovskaya" authorityYear="2024" box="[269,417,1833,1859]" class="Insecta" family="Chironomidae" genus="Thalassomya" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="527" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEAEAFF943B08204D2738FE03F8FE" box="[269,417,1833,1859]" italics="true" pageId="3" pageNumber="527">Thalassomya</emphasis>
</taxonomicName>
from the Bulgarian Black Sea coast, which was recorded as
<taxonomicName id="4C0A4D7BFF943B082506273BFB7EF8FE" baseAuthorityName="Wirth" baseAuthorityYear="1947" box="[1094,1244,1833,1859]" class="Insecta" family="Chironomidae" genus="Thalassomya" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="527" phylum="Arthropoda" rank="species" species="frauenfeldi">
<emphasis id="B97EEAEAFF943B082506273BFB7EF8FE" box="[1094,1244,1833,1859]" italics="true" pageId="3" pageNumber="527">T. frauenfeldi</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF9B4B09FF943B0825AA2738FA35F8FE" author="Valkanov, A." box="[1258,1431,1833,1859]" pageId="3" pageNumber="527" pagination="103 - 112" refId="ref4327" refString="Valkanov, A. (1949) Thalassomyia frauenfeldi Schiner ot Cherno More. Trudove na morska biologichna stantsiya v Varna, 14, 103 - 112. [in Bulgarian]" type="journal article" year="1949">Valkanov 1949</bibRefCitation>
;
<bibRefCitation id="EF9B4B09FF943B0821D7275CFE44F8DA" author="Valkanov, A. &amp; Michailova, P." box="[151,486,1869,1895]" pageId="3" pageNumber="527" pagination="5 - 16" refId="ref4359" refString="Valkanov, A. &amp; Michailova, P. (1974) Untersuchungen uber den Karyotypus und Chromosomen Polymorphismus bei Thalassomyia frauenfeldi Schiner (Diptera, Chironomidae) von der bulgarischen Schwarzmeerkuste. Bulletin de l'Institut de Zoologie et Musee Academie Bulgare des Science, 40, 5 - 16." type="journal article" year="1974">Valkanov &amp; Michailova 1974</bibRefCitation>
;
<bibRefCitation id="EF9B4B09FF943B0820B2275CFD10F8DA" author="Michailova, P." box="[498,690,1869,1895]" pageId="3" pageNumber="527" pagination="267 - 269" refId="ref4003" refString="Michailova, P. (1976) An additional B chromosome in Thalassomyia frauenfeldi Schiner (Chironomidae, Diptera) from Bulgaria's Black Sea coast. Doklady bolgarskoi akademii nauk, 29 (2), 267 - 269." type="journal article" year="1976">Michailova 1976</bibRefCitation>
) and may be
<taxonomicName id="4C0A4D7BFF943B082209275FFC46F8DA" authorityName="Makarchenko &amp; Semenchenko &amp; Palatov &amp; Lisanovskaya" authorityYear="2024" box="[841,996,1869,1895]" class="Insecta" family="Chironomidae" genus="Thalassomya" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="527" phylum="Arthropoda" rank="species" species="paraskevae" status="sp. nov.">
<emphasis id="B97EEAEAFF943B082209275FFC46F8DA" box="[841,996,1869,1895]" italics="true" pageId="3" pageNumber="527">T. paraskevae</emphasis>
</taxonomicName>
<emphasis id="B97EEAEAFF943B0822AB275CFBE6F8DA" bold="true" box="[1003,1092,1869,1895]" pageId="3" pageNumber="527">
<taxonomicNameLabel id="A24D5791FF943B0822AB275CFBE6F8DA" box="[1003,1092,1869,1895]" pageId="3" pageNumber="527" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3106573FF943B0F21872760FF15FF68" lastPageId="4" lastPageNumber="528" pageId="3" pageNumber="527" type="biology_ecology">
<paragraph id="8BB536F8FF943B0F21872760FF15FF68" blockId="3.[151,1437,645,2040]" lastBlockId="4.[151,1436,151,249]" lastPageId="4" lastPageNumber="528" pageId="3" pageNumber="527">
<emphasis id="B97EEAEAFF943B0821872760FE8EF836" bold="true" box="[199,300,1905,1931]" pageId="3" pageNumber="527">Ecology.</emphasis>
Like other species of
<taxonomicName id="4C0A4D7BFF943B08235E2760FD10F836" authorityName="Makarchenko &amp; Semenchenko &amp; Palatov &amp; Lisanovskaya" authorityYear="2024" box="[542,690,1905,1931]" class="Insecta" family="Chironomidae" genus="Thalassomya" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="527" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEAEAFF943B08235E2760FD10F836" box="[542,690,1905,1931]" italics="true" pageId="3" pageNumber="527">Thalassomya</emphasis>
</taxonomicName>
, the new species is associated with algal fouling on rocks or similar artificial structures such as breakwaters located along the coastal area. Adult individuals actively moved along the rocks at the waterline, in the splash zone, occasionally flying from one place to another. The highest locomotor activity of imago was observed during daytime hours, correlated with maximum air temperature. However, the activity of some individuals had been also noted at night. The species cohabits with
<taxonomicName id="4C0A4D7BFF933B0F25172086FAB5FF0C" box="[1111,1303,151,177]" class="Insecta" family="Chironomidae" genus="Telmatogeton" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="528" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B97EEAEAFF933B0F25172086FB4CFF0C" box="[1111,1262,151,177]" italics="true" pageId="4" pageNumber="528">Telmatogeton</emphasis>
sp.
</taxonomicName>
and
<taxonomicName id="4C0A4D7BFF933B0F24102086FF15FF68" class="Insecta" family="Chironomidae" genus="Clunio" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="528" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B97EEAEAFF933B0F24102086FA3EFF0C" box="[1360,1436,151,177]" italics="true" pageId="4" pageNumber="528">Clunio</emphasis>
sp.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3106573FF933B0F218720CFFB0FFF44" box="[199,1197,222,249]" pageId="4" pageNumber="528" type="distribution">
<paragraph id="8BB536F8FF933B0F218720CFFB0FFF44" blockId="4.[151,1436,151,249]" box="[199,1197,222,249]" pageId="4" pageNumber="528">
<emphasis id="B97EEAEAFF933B0F218720CFFEC3FF45" bold="true" box="[199,353,222,248]" pageId="4" pageNumber="528">Distribution.</emphasis>
Known only from the
<typeStatus id="54B1885AFF933B0F232020CEFD32FF44" box="[608,656,223,249]" pageId="4" pageNumber="528">type</typeStatus>
locality in coast of the Black Sea (
<figureCitation id="13312A7DFF933B0F255720CEFB03FF44" box="[1047,1185,222,249]" captionStart="FIGURES 1215" captionStartId="4.[152,270,1036,1060]" captionTargetBox="[151,1435,273,1013]" captionTargetId="figure-52@4.[151,1435,273,1013]" captionTargetPageId="4" captionText="FIGURES 1215. Habitats of Thalassomya paraskevae sp. nov. on the coast of the Black Sea near New Athos (Abkhazia) (12 13, type locality) and Cape Kadosh in the vicinity of Tuapse (Krasnodar Territory, Russia) (1415). Arrows indicate collection points." figureDoi="http://doi.org/10.5281/zenodo.13745326" httpUri="https://zenodo.org/record/13745326/files/figure.png" pageId="4" pageNumber="528">Figs. 1215</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<taxonomicName id="4C05A8A9FF90FFFCFF17F93AFD93BB70" authority="(Hampe, 1850)" baseAuthorityName="Hampe" baseAuthorityYear="1850" box="[151,609,1771,1798]" class="Insecta" family="Eucnemidae" genus="Clypeorhagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="565" phylum="Arthropoda" rank="species" species="clypeatus">
<emphasis id="B9710F38FF90FFFCFF17F93AFD93BB70" bold="true" box="[151,609,1771,1798]" pageId="1" pageNumber="565">
<emphasis id="B9710F38FF90FFFCFF17F93AFE5BBB73" bold="true" box="[151,425,1771,1798]" italics="true" pageId="1" pageNumber="565">Clypeorhagus clypeatus</emphasis>
(Hampe, 1850)
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Distribution in
<collectingCountry id="F31293BAFF90FFFCFED8F8DEFE7DBB5C" box="[344,399,1807,1833]" name="Iran" pageId="1" pageNumber="565">Iran</collectingCountry>
.
</emphasis>
<collectingRegion id="49C11DC8FF90FFFCFE1FF8DEFDF1BB5C" box="[415,515,1807,1833]" country="Iran" name="Golestan" pageId="1" pageNumber="565">Golestan</collectingRegion>
(
<bibRefCitation id="EF94AEDBFF90FFFCFD95F8DEFCE9BB5C" author="Farashiani, M. E. &amp; Varandi, H. B. &amp; Kazerani, F. &amp; Yarmand, H. &amp; Babaee, M. &amp; Thorn, S. &amp; Lange, F. &amp; Rafiei-Jahed, R. &amp; Muller, J. &amp; Amini, S." box="[533,795,1807,1834]" pageId="1" pageNumber="565" pagination="1063 - 1120" refId="ref4245" refString="Farashiani, M. E., Varandi, H. B., Kazerani, F., Yarmand, H., Babaee, M., Thorn, S., Lange, F., Rafiei-Jahed, R., Muller, J. &amp; Amini, S. (2022) A preliminary checklist of saproxylic beetles (Coleoptera) in the Hyrcanian forests of Iran, with distributional data. Check List, 18 (5), 1063 - 1120. https: // doi. org / 10.15560 / 18.5.1053" type="journal article" year="2022">
Farashiani
<emphasis id="B9710F38FF90FFFCFD17F8C1FD3FBB5C" box="[663,717,1807,1833]" italics="true" pageId="1" pageNumber="565">et al</emphasis>
. 2022
</bibRefCitation>
),
<collectingRegion id="49C11DC8FF90FFFCFCB3F8DEFC4FBB5C" box="[819,957,1807,1833]" country="Iran" name="Mazandaran" pageId="1" pageNumber="565">Mazandaran</collectingRegion>
(
<bibRefCitation id="EF94AEDBFF90FFFCFC50F8DEFB22BB5C" author="Jarzabek-Muller, A." box="[976,1232,1807,1833]" pageId="1" pageNumber="565" pagination="119 - 125" refId="ref4747" refString="Jarzabek-Muller, A. (2019) Neue Nachweise von Schienen- und Schnellkafern (Coleoptera: Eucnemidae, Elateridae) fur die palaarktische Fauna sowie eine Anmerkung zur Verbreitung von Hemicrepidius koenigi (Schwarz, 1897). Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen, 71, 119 - 125." type="journal article" year="2019">Jarzabek-Müller 2019</bibRefCitation>
;
<bibRefCitation id="EF94AEDBFF90FFFCFB60F8DEFEF6BB38" author="Farashiani, M. E. &amp; Varandi, H. B. &amp; Kazerani, F. &amp; Yarmand, H. &amp; Babaee, M. &amp; Thorn, S. &amp; Lange, F. &amp; Rafiei-Jahed, R. &amp; Muller, J. &amp; Amini, S." pageId="1" pageNumber="565" pagination="1063 - 1120" refId="ref4245" refString="Farashiani, M. E., Varandi, H. B., Kazerani, F., Yarmand, H., Babaee, M., Thorn, S., Lange, F., Rafiei-Jahed, R., Muller, J. &amp; Amini, S. (2022) A preliminary checklist of saproxylic beetles (Coleoptera) in the Hyrcanian forests of Iran, with distributional data. Check List, 18 (5), 1063 - 1120. https: // doi. org / 10.15560 / 18.5.1053" type="journal article" year="2022">
Farashiani
<emphasis id="B9710F38FF90FFFCFAE0F8C1FA67BB5C" box="[1376,1429,1807,1833]" italics="true" pageId="1" pageNumber="565">et al</emphasis>
. 2022
</bibRefCitation>
).
</paragraph>
<paragraph id="8BBAD32AFF90FFFCFF17F886FE91BBA8" blockId="1.[151,1436,1663,2013]" pageId="1" pageNumber="565">
<emphasis id="B9710F38FF90FFFCFF17F886FE60BB04" bold="true" box="[151,402,1879,1905]" pageId="1" pageNumber="565">General distribution.</emphasis>
<collectingCountry id="F31293BAFF90FFFCFE18F886FDF3BB04" box="[408,513,1879,1905]" name="Germany" pageId="1" pageNumber="565">Germany</collectingCountry>
,
<collectingCountry id="F31293BAFF90FFFCFD8BF889FD9CBB07" box="[523,622,1880,1906]" name="Hungary" pageId="1" pageNumber="565">Hungary</collectingCountry>
,
<collectingCountry id="F31293BAFF90FFFCFDF7F886FD33BB04" box="[631,705,1879,1905]" name="Latvia" pageId="1" pageNumber="565">Latvia</collectingCountry>
,
<collectingCountry id="F31293BAFF90FFFCFD4CF886FCEFBB04" box="[716,797,1879,1905]" name="Poland" pageId="1" pageNumber="565">Poland</collectingCountry>
,
<collectingCountry id="F31293BAFF90FFFCFCA6F886FC7CBB04" box="[806,910,1879,1905]" name="Romania" pageId="1" pageNumber="565">Romania</collectingCountry>
,
<collectingCountry id="F31293BAFF90FFFCFC18F886FC11BB04" box="[920,995,1879,1905]" name="Russia" pageId="1" pageNumber="565">Russia</collectingCountry>
(Central European Territory),
<collectingCountry id="F31293BAFF90FFFCFAB3F886FA6ABB04" box="[1331,1432,1879,1905]" name="Slovakia" pageId="1" pageNumber="565">Slovakia</collectingCountry>
,
<collectingCountry id="F31293BAFF90FFFCFF47F8AAFED3BBE0" box="[199,289,1915,1941]" name="Ukraine" pageId="1" pageNumber="565">Ukraine</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF90FFFCFEB3F8AAFE38BBE0" author="Muona, J." box="[307,458,1915,1941]" pageId="1" pageNumber="565" refId="ref5680" refString="Muona, J. (2007) Family Eucnemidae Eschscholtz, 1829. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 4. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, pp. 81 - 87. https: // doi. org / 10.1163 / 9789004260894 _ 004" type="book" year="2007">Muona 2007</bibRefCitation>
),
<collectingCountry id="F31293BAFF90FFFCFE61F8AAFDDEBBE0" box="[481,556,1915,1941]" name="Russia" pageId="1" pageNumber="565">Russia</collectingCountry>
(South European Territory) (
<bibRefCitation id="EF94AEDBFF90FFFCFCFEF8AAFB9EBBE0" author="Nikitsky, N. V. &amp; Bibin, A. R." box="[894,1132,1915,1942]" pageId="1" pageNumber="565" refId="ref6282" refString="Nikitsky, N. V., Bibin, A. R. &amp; Dolgin М. М. (2008) Ksilofil'nye zhestkokrylye Kavkazskogo gosudarstvennogo prirodnogo biosfernogo zapovednika i sopredel'nykh territorij [Xylophilous beetles (Coleoptera) of the Caucasian State Biospheric Natural Reserve and adjacent territories]. Institute of Biology of Komi center of science of the Ural branch of the Russian Academy of sciences, Syktyvkar, 452 рр. [in Russian]" type="journal volume" year="2008">
Nikitsky
<emphasis id="B9710F38FF90FFFCFC6AF8ADFBEDBBE0" box="[1002,1055,1915,1941]" italics="true" pageId="1" pageNumber="565">et al</emphasis>
. 2008
</bibRefCitation>
),
<collectingCountry id="F31293BAFF90FFFCFB02F8AAFA6EBBE0" box="[1154,1436,1915,1941]" name="Bosnia and Herzegovina" pageId="1" pageNumber="565">Bosnia and Herzegovina</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF90FFFCFF4EF84EFE5CBBCC" author="Mertlik, J. &amp; Jenis, I. &amp; Zbuzek, B." box="[206,430,1951,1978]" pageId="1" pageNumber="565" pagination="1 - 6" refId="ref5539" refString="Mertlik, J., Jenis, I. &amp; Zbuzek, B. (2009) New records on the distribution of some species of the family Melasidae (Coleoptera) - II. Elateridarium, 3, 1 - 6." type="journal article" year="2009">
Mertlik
<emphasis id="B9710F38FF90FFFCFEAEF871FE93BBCC" box="[302,353,1951,1977]" italics="true" pageId="1" pageNumber="565">et al</emphasis>
. 2009
</bibRefCitation>
),
<collectingCountry id="F31293BAFF90FFFCFE44F84EFDEABBCC" box="[452,536,1951,1977]" name="Estonia" pageId="1" pageNumber="565">Estonia</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF90FFFCFDA9F84EFD5ABBCC" author="Suda, I." box="[553,680,1951,1977]" pageId="1" pageNumber="565" pagination="98 - 114" refId="ref7238" refString="Suda, I. (2009) Metsamardikate (Coleoptera) uued liigid Eestis [New woodland beetle species (Coleoptera) in Estonian fauna]. Forestry Studies, 50, 98 - 114. https: // doi. org / 10.2478 / v 10132 - 011 - 0071 - 0" type="journal article" year="2009">Süda 2009</bibRefCitation>
),
<collectingCountry id="F31293BAFF90FFFCFD3EF84EFCE6BBCC" box="[702,788,1951,1977]" name="Belarus" pageId="1" pageNumber="565">Belarus</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF90FFFCFCA4F84EFB29BBCC" author="Solodovnikov, I. A. &amp; Solodovnikova, S. V." box="[804,1243,1951,1977]" pageId="1" pageNumber="565" pagination="62 - 71" refId="ref7171" refString="Solodovnikov, I. A. &amp; Solodovnikova, S. V. (2019) New and rare species of beetles (Coleoptera) in Belarusian Lake District (Poozeriye) and in the Republic of Belarus. Part 8. Viesnik Viciebskaha dziarzaunaha univiersiteta [Bulletin of Vitebsk State University], 1 (102), 62 - 71. [in Russian]" type="journal article" year="2019">Solodovnikov &amp; Solodovnikova 2019</bibRefCitation>
),
<collectingCountry id="F31293BAFF90FFFCFB71F871FAECBBCF" box="[1265,1310,1952,1978]" name="Iran" pageId="1" pageNumber="565">Iran</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF90FFFCFAAFF84EFEA5BBA8" author="Jarzabek-Muller, A." pageId="1" pageNumber="565" pagination="119 - 125" refId="ref4747" refString="Jarzabek-Muller, A. (2019) Neue Nachweise von Schienen- und Schnellkafern (Coleoptera: Eucnemidae, Elateridae) fur die palaarktische Fauna sowie eine Anmerkung zur Verbreitung von Hemicrepidius koenigi (Schwarz, 1897). Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen, 71, 119 - 125." type="journal article" year="2019">Jarzabek-Müller 2019</bibRefCitation>
).
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View file

@ -0,0 +1,98 @@
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<mods:title id="841641BB9393790ED1133E797AE463C1">A checklist of Eucnemidae (Coleoptera: Elateroidea) from Iran</mods:title>
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<taxonomicName id="4C05A8A9FF90FFFCFF17FA3EFD90BA7C" authority="Roubal, 1912" authorityName="Roubal" authorityYear="1912" box="[151,610,1519,1546]" class="Insecta" family="Eucnemidae" genus="Dromaeolus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="565" phylum="Arthropoda" rank="species" species="gracilicornis">
<emphasis id="B9710F38FF90FFFCFF17FA3EFD90BA7C" bold="true" box="[151,610,1519,1546]" pageId="1" pageNumber="565">
<emphasis id="B9710F38FF90FFFCFF17FA3EFE4CBA7F" bold="true" box="[151,446,1519,1546]" italics="true" pageId="1" pageNumber="565">Dromaeolus gracilicornis</emphasis>
<bibRefCitation id="EF94AEDBFF90FFFCFE45FA3EFD90BA7C" author="Roubal, J." box="[453,610,1519,1545]" pageId="1" pageNumber="565" pagination="124 - 126" refId="ref6901" refString="Roubal, J. (1912) Koleopterologicke vysledky me cesty na Kavkaz v cervenci r. 1910. Quid novi de Coleopterorum Caucasi ad occidentem vergentis fauna in meo itinere Julio mense anni 1910 suscepto cognoverim. VI. Krasnaja Poljana. Eucnemidae. Casopis Ceske Spolecnosti Entomologicke, 9, 124 - 126." type="journal article" year="1912">Roubal, 1912</bibRefCitation>
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<paragraph id="8BBAD32AFF90FFFCFF17F9C2FCC2BA58" blockId="1.[151,1300,1411,1618]" box="[151,816,1555,1582]" pageId="1" pageNumber="565">
<emphasis id="B9710F38FF90FFFCFF17F9C2FE79BA58" bold="true" box="[151,395,1555,1581]" pageId="1" pageNumber="565">
Distribution in
<collectingCountry id="F31293BAFF90FFFCFECFF9C2FE74BA58" box="[335,390,1555,1581]" name="Iran" pageId="1" pageNumber="565">Iran</collectingCountry>
.
</emphasis>
<collectingRegion id="49C11DC8FF90FFFCFE12F9C2FDEEBA58" box="[402,540,1555,1581]" country="Iran" name="Mazandaran" pageId="1" pageNumber="565">Mazandaran</collectingRegion>
(
<bibRefCitation id="EF94AEDBFF90FFFCFDAAF9C2FCD6BA58" author="Farashiani, M. E. &amp; Varandi, H. B. &amp; Kazerani, F. &amp; Yarmand, H. &amp; Babaee, M. &amp; Thorn, S. &amp; Lange, F. &amp; Rafiei-Jahed, R. &amp; Muller, J. &amp; Amini, S." box="[554,804,1555,1582]" pageId="1" pageNumber="565" pagination="1063 - 1120" refId="ref4245" refString="Farashiani, M. E., Varandi, H. B., Kazerani, F., Yarmand, H., Babaee, M., Thorn, S., Lange, F., Rafiei-Jahed, R., Muller, J. &amp; Amini, S. (2022) A preliminary checklist of saproxylic beetles (Coleoptera) in the Hyrcanian forests of Iran, with distributional data. Check List, 18 (5), 1063 - 1120. https: // doi. org / 10.15560 / 18.5.1053" type="journal article" year="2022">
Farashiani
<emphasis id="B9710F38FF90FFFCFD28F9C5FD2BBA58" box="[680,729,1555,1581]" italics="true" pageId="1" pageNumber="565">et al</emphasis>
. 2022
</bibRefCitation>
).
</paragraph>
<paragraph id="8BBAD32AFF90FFFCFF17F9E6FAE6BA24" blockId="1.[151,1300,1411,1618]" box="[151,1300,1591,1618]" pageId="1" pageNumber="565">
<emphasis id="B9710F38FF90FFFCFF17F9E6FE61BA24" bold="true" box="[151,403,1591,1617]" pageId="1" pageNumber="565">General distribution.</emphasis>
<collectingCountry id="F31293BAFF90FFFCFE1AF9E6FE17BA24" box="[410,485,1591,1617]" name="Russia" pageId="1" pageNumber="565">Russia</collectingCountry>
(South European Territory) (
<bibRefCitation id="EF94AEDBFF90FFFCFCAAF9E6FC32BA24" author="Roubal, J." box="[810,960,1591,1617]" pageId="1" pageNumber="565" pagination="124 - 126" refId="ref6901" refString="Roubal, J. (1912) Koleopterologicke vysledky me cesty na Kavkaz v cervenci r. 1910. Quid novi de Coleopterorum Caucasi ad occidentem vergentis fauna in meo itinere Julio mense anni 1910 suscepto cognoverim. VI. Krasnaja Poljana. Eucnemidae. Casopis Ceske Spolecnosti Entomologicke, 9, 124 - 126." type="journal article" year="1912">Roubal 1912</bibRefCitation>
),
<collectingCountry id="F31293BAFF90FFFCFC53F9E9FBF2BA27" box="[979,1024,1592,1618]" name="Iran" pageId="1" pageNumber="565">Iran</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF90FFFCFB8FF9E6FAFABA24" author="Farashiani, M. E. &amp; Varandi, H. B. &amp; Kazerani, F. &amp; Yarmand, H. &amp; Babaee, M. &amp; Thorn, S. &amp; Lange, F. &amp; Rafiei-Jahed, R. &amp; Muller, J. &amp; Amini, S." box="[1039,1288,1591,1618]" pageId="1" pageNumber="565" pagination="1063 - 1120" refId="ref4245" refString="Farashiani, M. E., Varandi, H. B., Kazerani, F., Yarmand, H., Babaee, M., Thorn, S., Lange, F., Rafiei-Jahed, R., Muller, J. &amp; Amini, S. (2022) A preliminary checklist of saproxylic beetles (Coleoptera) in the Hyrcanian forests of Iran, with distributional data. Check List, 18 (5), 1063 - 1120. https: // doi. org / 10.15560 / 18.5.1053" type="journal article" year="2022">
Farashiani
<emphasis id="B9710F38FF90FFFCFB0CF9E9FB4CBA24" box="[1164,1214,1591,1617]" italics="true" pageId="1" pageNumber="565">et al</emphasis>
. 2022
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,139 @@
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<mods:title id="9E5EFFF133F723816148E917358B3DD9">A checklist of Eucnemidae (Coleoptera: Elateroidea) from Iran</mods:title>
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<paragraph id="8BBAD32AFF92FFFEFF17F881FDB1BB1F" blockId="3.[151,1436,1764,2042]" box="[151,579,1872,1899]" pageId="3" pageNumber="567">
<heading id="D0F26446FF92FFFEFF17F881FDB1BB1F" bold="true" box="[151,579,1872,1899]" fontSize="11" level="1" pageId="3" pageNumber="567" reason="1">
<taxonomicName id="4C05A8A9FF92FFFEFF17F881FDB1BB1F" authority="P. H. Lucas, 1846" authorityName="P. H. Lucas" authorityYear="1846" box="[151,579,1872,1899]" class="Insecta" family="Eucnemidae" genus="Anelastes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="567" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="B9710F38FF92FFFEFF17F881FDB1BB1F" bold="true" box="[151,579,1872,1899]" pageId="3" pageNumber="567">
<emphasis id="B9710F38FF92FFFEFF17F881FE87BB1E" bold="true" box="[151,373,1872,1899]" italics="true" pageId="3" pageNumber="567">Anelastes barbarus</emphasis>
P.H. Lucas, 1846
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<subSubSection id="C31F80A1FF92FFFEFF17F8A5FE8BBB8F" pageId="3" pageNumber="567" type="distribution">
<paragraph id="8BBAD32AFF92FFFEFF17F8A5FA6EBBC7" blockId="3.[151,1436,1764,2042]" pageId="3" pageNumber="567">
<emphasis id="B9710F38FF92FFFEFF17F8A5FE79BBFB" bold="true" box="[151,395,1908,1934]" pageId="3" pageNumber="567">
Distribution in
<collectingCountry id="F31293BAFF92FFFEFECFF8A5FE74BBFB" box="[335,390,1908,1934]" name="Iran" pageId="3" pageNumber="567">Iran</collectingCountry>
.
</emphasis>
<collectingRegion id="49C11DC8FF92FFFEFE12F8A5FE04BBFB" box="[402,502,1908,1934]" country="Iran" name="Golestan" pageId="3" pageNumber="567">Golestan</collectingRegion>
(
<bibRefCitation id="EF94AEDBFF92FFFEFD84F8A5FD3FBBFA" author="Bonvouloir, H. A. de" box="[516,717,1908,1935]" pageId="3" pageNumber="567" pagination="561 - 907" refId="ref3640" refString="Bonvouloir, H. A. de (1875) Monographie de la famille des eucnemides. 4 e Cahier. L. Buquet, Societe Entomologique de France, Paris, 907 pp., 42 pls. [1871 - 1875, pp. 561 - 907, pls. 37 - 42] https: // doi. org / 10.5962 / bhl. title. 48404" type="book chapter" year="1875">Bonvouloir, 1875</bibRefCitation>
, as
<taxonomicName id="4C05A8A9FF92FFFEFD76F8A5FBD7BBFA" authority="LeConte, 1852" authorityName="LeConte" authorityYear="1852" box="[758,1061,1908,1935]" class="Insecta" family="Eucnemidae" genus="Anelastes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="567" phylum="Arthropoda" rank="species" species="latreillei">
<emphasis id="B9710F38FF92FFFEFD76F8A5FC86BBFB" box="[758,884,1908,1934]" italics="true" pageId="3" pageNumber="567">A. latreillei</emphasis>
LeConte, 1852
</taxonomicName>
;
<bibRefCitation id="EF94AEDBFF92FFFEFBB1F8A5FB49BBFB" author="Lucht, W." box="[1073,1211,1908,1934]" pageId="3" pageNumber="567" pagination="133 - 140" refId="ref5151" refString="Lucht, W. (1990) Erstnachweis von Anelastes barbarus Luc. in Sudosteuropa (Coleoptera: Eucnemidae: Melasinae). Mitteilungen des Internationalen Entomologischen Vereins, 15 (3 - 4), 133 - 140." type="book chapter" year="1990">Lucht, 1990</bibRefCitation>
;
<bibRefCitation id="EF94AEDBFF92FFFEFB47F8A5FA9DBBFB" author="Kovalev, A. V." box="[1223,1391,1908,1934]" pageId="3" pageNumber="567" pagination="97 - 119" refId="ref4951" refString="Kovalev, A. V. (2019) A review of the genus Anelastes Kirby, 1819 (Coleoptera: Eucnemidae) of the Palaearctic fauna. Zootaxa, 4683 (1), 97 - 119. https: // doi. org / 10.11646 / zootaxa. 4683.1.5" type="journal article" year="2019">Kovalev, 2019</bibRefCitation>
).
<emphasis id="B9710F38FF92FFFEFF17F849FE68BBC7" bold="true" box="[151,410,1944,1970]" pageId="3" pageNumber="567">General distribution.</emphasis>
Algeria, Croatia, France, Greece, Iran, Italy, Libya, Morocco, Spain, Tunisia, Turkey,
</paragraph>
<paragraph id="8BBAD32AFF92FFFEFF47F86DFA6EBBA3" blockId="3.[151,1436,1764,2042]" box="[199,1436,1980,2007]" pageId="3" pageNumber="567">
<collectingCountry id="F31293BAFF92FFFEFF47F86DFE90BBA3" box="[199,354,1980,2006]" name="Turkmenistan" pageId="3" pageNumber="567">Turkmenistan</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF92FFFEFEEEF86DFDF2BBA3" author="Muona, J." box="[366,512,1980,2006]" pageId="3" pageNumber="567" refId="ref5680" refString="Muona, J. (2007) Family Eucnemidae Eschscholtz, 1829. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 4. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, pp. 81 - 87. https: // doi. org / 10.1163 / 9789004260894 _ 004" type="book" year="2007">Muona 2007</bibRefCitation>
), Canary Islands (Tenerife) (
<bibRefCitation id="EF94AEDBFF92FFFEFCB9F86DFC31BBA2" author="Cobos, A." box="[825,963,1980,2007]" pageId="3" pageNumber="567" pagination="27 - 30" refId="ref4103" refString="Cobos, A. (1965) Comentarios sobre Anelastes barbarus Lucas y su rehabilitacion especifica (Col. Eucnemidae). Eos, Revista Espanola de Entomologia, 41 (1), 27 - 30." type="journal article" year="1965">Cobos 1965</bibRefCitation>
),
<collectingCountry id="F31293BAFF92FFFEFC55F86DFBA1BBA3" box="[981,1107,1980,2006]" name="Uzbekistan" pageId="3" pageNumber="567">Uzbekistan</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF92FFFEFBE0F86DFAF0BBA3" author="Kovalev, A. V." box="[1120,1282,1980,2006]" pageId="3" pageNumber="567" pagination="97 - 119" refId="ref4951" refString="Kovalev, A. V. (2019) A review of the genus Anelastes Kirby, 1819 (Coleoptera: Eucnemidae) of the Palaearctic fauna. Zootaxa, 4683 (1), 97 - 119. https: // doi. org / 10.11646 / zootaxa. 4683.1.5" type="journal article" year="2019">Kovalev 2019</bibRefCitation>
),
<collectingCountry id="F31293BAFF92FFFEFA93F86DFA6EBBA3" box="[1299,1436,1980,2006]" name="Serbia" pageId="3" pageNumber="567">Montenegro</collectingCountry>
</paragraph>
<paragraph id="8BBAD32AFF92FFFEFF47F831FE8BBB8F" blockId="3.[151,1436,1764,2042]" box="[199,377,2016,2042]" pageId="3" pageNumber="567">
(
<bibRefCitation id="EF94AEDBFF92FFFEFF4EF831FE9FBB8F" author="Nemeth, T." box="[206,365,2016,2042]" pageId="3" pageNumber="567" pagination="31 - 33" refId="ref6168" refString="Nemeth, T. (2023) The mysterious Anelastes barbarus P. H. Lucas, 1846 from Montenegro (Coleoptera: Eucnemidae). Folia Entomologica Hungarica, 84, 31 - 33. https: // doi. org / 10.17112 / FoliaEntHung. 2023.84.31" type="journal article" year="2023">Németh 2023</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31F80A1FF95FFF9FF17FF46FE53BC8C" pageId="4" pageNumber="568" type="discussion">
<paragraph id="8BBAD32AFF95FFF9FF17FF46FE53BC8C" blockId="4.[151,1437,151,249]" pageId="4" pageNumber="568">
<emphasis id="B9710F38FF95FFF9FF17FF46FEF2BCC4" bold="true" box="[151,256,151,177]" pageId="4" pageNumber="568">Remark.</emphasis>
All citations of
<taxonomicName id="4C05A8A9FF95FFF9FE30FF46FD68BCC4" authority="Kirby, 1819" authorityName="Kirby" authorityYear="1819" box="[432,666,151,177]" class="Insecta" family="Eucnemidae" genus="Anelastes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="568" phylum="Arthropoda" rank="species" species="drurii">
<emphasis id="B9710F38FF95FFF9FE30FF46FDFDBCC4" box="[432,527,151,177]" italics="true" pageId="4" pageNumber="568">A. drurii</emphasis>
Kirby, 1819
</taxonomicName>
(
<taxonomicName id="4C05A8A9FF95FFF9FD29FF46FCFFBCC4" authorityName="Kirby" authorityYear="1818" box="[681,781,151,177]" class="Insecta" family="Eucnemidae" genus="Anelastes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="568" phylum="Arthropoda" rank="species" species="druryi">
<emphasis id="B9710F38FF95FFF9FD29FF46FCFFBCC4" box="[681,781,151,177]" italics="true" pageId="4" pageNumber="568">A. druryi</emphasis>
</taxonomicName>
auct.) from “
<collectingCountry id="F31293BAFF95FFF9FC1FFF46FC1ABCC4" box="[927,1000,151,177]" name="Iran" pageId="4" pageNumber="568">Persia</collectingCountry>
” (
<bibRefCitation id="EF94AEDBFF95FFF9FB80FF46FB71BCC4" author="Bedel, L." box="[1024,1155,151,177]" pageId="4" pageNumber="568" pagination="223 - 228" refId="ref3518" refString="Bedel, L. (1904) Liste des principales especes de Coleopteres recueillies par le Mis de Segonzac. In: Mis de Segonzac, Voyages au Maroc (1899 - 1901). L'Abeille, Journal d'Entomologie, 30, pp. 223 - 228." type="journal article" year="1904">Bedel 1904</bibRefCitation>
;
<bibRefCitation id="EF94AEDBFF95FFF9FB0EFF46FAC5BCC4" author="Jakobson, G. G." box="[1166,1335,151,177]" pageId="4" pageNumber="568" pagination="76 - 83" refId="ref4677" refString="Jakobson, G. G. (1913) Vypusk 10. In: Zhuki Rossii i Zapadnoy Evropy. Rukovodstvo k opredeleniyu zhukov. [Beetles of Russia and Western Europe. A guide to identification of beetles]. Fascicle 10. A. F. Devrien, St. Petersburg, pp. 721 - 864, pls. 76 - 83. [in Russian]" type="book chapter" year="1913">Jakobson 1913</bibRefCitation>
;
<bibRefCitation id="EF94AEDBFF95FFF9FAC2FF46FEF0BCA0" author="Winkler, A." pageId="4" pageNumber="568" pagination="617 - 619" refId="ref7531" refString="Winkler, A. (1925) Eucnemidae. In: Winkler, A. (Ed.), Catalogus Coleopterorum regionis palaearcticae. I. A. Caraboidea, B. Palpicornia, Staphylinoidea, C. Diversicornia. Pars 5. A. Winkler, Wien, pp. 617 - 619." type="book chapter" year="1925">Winkler 1925</bibRefCitation>
;
<bibRefCitation id="EF94AEDBFF95FFF9FE8CFF6AFE38BCA0" author="Schenkling, S." box="[268,458,187,213]" pageId="4" pageNumber="568" refId="ref7011" refString="Schenkling, S. (1928) Coleopterorum Catalogus. Pars 96. Melasidae. W. Junk, Berlin, 110 pp." type="book" year="1928">Schenkling 1928</bibRefCitation>
;
<bibRefCitation id="EF94AEDBFF95FFF9FE54FF6AFDABBCA0" author="Cobos, A." box="[468,601,186,213]" pageId="4" pageNumber="568" pagination="77 - 82" refId="ref4070" refString="Cobos, A. (1959) Materiales para el conocimiento de los Eucnemidae y Throscidae (Coleoptera) ibericos. Miscelanea Zoologica, 1 (2), 77 - 82." type="journal article" year="1959">Cobos 1959</bibRefCitation>
) appear to be based on the Bonvouloir's record of
<taxonomicName id="4C05A8A9FF95FFF9FBFCFF6AFB0ABCA0" authorityName="LeConte" authorityYear="1852" box="[1148,1272,187,213]" class="Insecta" family="Eucnemidae" genus="Anelastes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="568" phylum="Arthropoda" rank="species" species="latreillei">
<emphasis id="B9710F38FF95FFF9FBFCFF6AFB0ABCA0" box="[1148,1272,187,213]" italics="true" pageId="4" pageNumber="568">A. latreillei</emphasis>
</taxonomicName>
from Astrabad (currently Gorgan).
</paragraph>
</subSubSection>
</treatment>
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@ -0,0 +1,188 @@
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<mods:affiliation id="05B61CC1011EEE3750CC44C479226DED">All-Russian Institute of Plant Protection of the Russian Academy of Sciences, Pushkin, St. Petersburg, 196608 Russia. &amp; Biological Faculty, M. V. Lomonosov Moscow State University, Moscow, 119234 Russia.</mods:affiliation>
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<mods:namePart id="E936409D7390E16280193C13D87980D1">Ghahari, Hassan</mods:namePart>
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<emphasis id="B9710F38FF92FFFEFF17FC1EFE7ABF9F" bold="true" box="[151,392,975,1002]" italics="true" pageId="3" pageNumber="567">Melasis buprestoides</emphasis>
(Linnaeus, 1760)
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Distribution in
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.
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<collectingRegion id="49C11DC8FF92FFFEFE12FC22FE0AB878" box="[402,504,1011,1037]" country="Iran" name="Golestan" pageId="3" pageNumber="567">Golestan</collectingRegion>
,
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Farashiani
<emphasis id="B9710F38FF92FFFEFD5DFC25FCFCB878" box="[733,782,1011,1037]" italics="true" pageId="3" pageNumber="567">et al</emphasis>
. 2022
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),
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(
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;
<bibRefCitation id="EF94AEDBFF92FFFEFA8CFC22FEDAB844" author="Farashiani, M. E. &amp; Varandi, H. B. &amp; Kazerani, F. &amp; Yarmand, H. &amp; Babaee, M. &amp; Thorn, S. &amp; Lange, F. &amp; Rafiei-Jahed, R. &amp; Muller, J. &amp; Amini, S." pageId="3" pageNumber="567" pagination="1063 - 1120" refId="ref4245" refString="Farashiani, M. E., Varandi, H. B., Kazerani, F., Yarmand, H., Babaee, M., Thorn, S., Lange, F., Rafiei-Jahed, R., Muller, J. &amp; Amini, S. (2022) A preliminary checklist of saproxylic beetles (Coleoptera) in the Hyrcanian forests of Iran, with distributional data. Check List, 18 (5), 1063 - 1120. https: // doi. org / 10.15560 / 18.5.1053" type="journal article" year="2022">
Farashiani
<emphasis id="B9710F38FF92FFFEFA08FC25FF2FB844" italics="true" pageId="3" pageNumber="567">et al</emphasis>
. 2022
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).
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<emphasis id="B9710F38FF92FFFEFF17FBEAFE6FB820" bold="true" box="[151,413,1083,1109]" pageId="3" pageNumber="567">General distribution.</emphasis>
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,
<collectingCountry id="F31293BAFF92FFFEFD96FBEAFD99B820" box="[534,619,1083,1109]" name="Austria" pageId="3" pageNumber="567">Austria</collectingCountry>
, Azerbaijan,
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,
<collectingCountry id="F31293BAFF92FFFEFCFEFBEAFC13B820" box="[894,993,1083,1109]" name="Belgium" pageId="3" pageNumber="567">Belgium</collectingCountry>
,
<collectingCountry id="F31293BAFF92FFFEFC76FBEAFAD2B820" box="[1014,1312,1083,1109]" name="Bosnia and Herzegovina" pageId="3" pageNumber="567">Bosnia and Herzegovina</collectingCountry>
,
<collectingCountry id="F31293BAFF92FFFEFAB5FBEAFA6AB820" box="[1333,1432,1083,1109]" name="Bulgaria" pageId="3" pageNumber="567">Bulgaria</collectingCountry>
,
<collectingCountry id="F31293BAFF92FFFEFF47FB8EFEEEB80C" box="[199,284,1119,1145]" name="Croatia" pageId="3" pageNumber="567">Croatia</collectingCountry>
,
<collectingCountry id="F31293BAFF92FFFEFEA8FB8EFE2CB80C" box="[296,478,1119,1145]" name="Czech Republic" pageId="3" pageNumber="567">Czech Republic</collectingCountry>
,
<collectingCountry id="F31293BAFF92FFFEFE6AFB8EFDA7B80C" box="[490,597,1119,1145]" name="Denmark" pageId="3" pageNumber="567">Denmark</collectingCountry>
,
<collectingCountry id="F31293BAFF92FFFEFDE1FB8EFD4BB80C" box="[609,697,1119,1145]" name="Finland" pageId="3" pageNumber="567">Finland</collectingCountry>
,
<collectingCountry id="F31293BAFF92FFFEFD45FBB1FCE6B80F" box="[709,788,1120,1146]" name="France" pageId="3" pageNumber="567">France</collectingCountry>
,
<collectingCountry id="F31293BAFF92FFFEFCA0FB8EFC7BB80C" box="[800,905,1119,1145]" name="Germany" pageId="3" pageNumber="567">Germany</collectingCountry>
,
<collectingCountry id="F31293BAFF92FFFEFC16FB8EFBDFB80C" box="[918,1069,1119,1145]" name="United Kingdom" pageId="3" pageNumber="567">Great Britain</collectingCountry>
,
<collectingCountry id="F31293BAFF92FFFEFBB8FB8EFB78B80C" box="[1080,1162,1119,1145]" name="Greece" pageId="3" pageNumber="567">Greece</collectingCountry>
,
<collectingCountry id="F31293BAFF92FFFEFB17FBB1FB08B80F" box="[1175,1274,1120,1146]" name="Hungary" pageId="3" pageNumber="567">Hungary</collectingCountry>
,
<collectingCountry id="F31293BAFF92FFFEFA86FB8EFAAAB80C" box="[1286,1368,1119,1145]" name="Ireland" pageId="3" pageNumber="567">Ireland</collectingCountry>
,
<collectingCountry id="F31293BAFF92FFFEFAE4FB8EFA6AB80C" box="[1380,1432,1119,1145]" name="Italy" pageId="3" pageNumber="567">Italy</collectingCountry>
,
<collectingCountry id="F31293BAFF92FFFEFF47FB52FEA8B8E8" box="[199,346,1155,1181]" name="Luxembourg" pageId="3" pageNumber="567">Luxembourg</collectingCountry>
,
<collectingCountry id="F31293BAFF92FFFEFEE6FB52FE02B8E8" box="[358,496,1155,1181]" name="Netherlands" pageId="3" pageNumber="567">Netherlands</collectingCountry>
, North Macedonia,
<collectingCountry id="F31293BAFF92FFFEFD52FB52FCDFB8E8" box="[722,813,1155,1181]" name="Norway" pageId="3" pageNumber="567">Norway</collectingCountry>
,
<collectingCountry id="F31293BAFF92FFFEFCB9FB52FC78B8E8" box="[825,906,1155,1181]" name="Poland" pageId="3" pageNumber="567">Poland</collectingCountry>
,
<collectingCountry id="F31293BAFF92FFFEFC16FB52FC0CB8E8" box="[918,1022,1155,1181]" name="Romania" pageId="3" pageNumber="567">Romania</collectingCountry>
,
<collectingCountry id="F31293BAFF92FFFEFB8AFB52FBA7B8E8" box="[1034,1109,1155,1181]" name="Russia" pageId="3" pageNumber="567">Russia</collectingCountry>
(Central European Territory, South European Territory, “Siberia”),
<collectingCountry id="F31293BAFF92FFFEFDECFB76FC87B8B4" box="[620,885,1191,1217]" name="Serbia" pageId="3" pageNumber="567">Serbia and Montenegro</collectingCountry>
,
<collectingCountry id="F31293BAFF92FFFEFCFEFB76FC11B8B4" box="[894,995,1191,1217]" name="Slovakia" pageId="3" pageNumber="567">Slovakia</collectingCountry>
,
<collectingCountry id="F31293BAFF92FFFEFC6EFB76FBA1B8B4" box="[1006,1107,1191,1217]" name="Slovenia" pageId="3" pageNumber="567">Slovenia</collectingCountry>
,
<collectingCountry id="F31293BAFF92FFFEFBDDFB76FB52B8B4" box="[1117,1184,1191,1217]" name="Spain" pageId="3" pageNumber="567">Spain</collectingCountry>
,
<collectingCountry id="F31293BAFF92FFFEFB2AFB76FAF7B8B4" box="[1194,1285,1191,1217]" name="Sweden" pageId="3" pageNumber="567">Sweden</collectingCountry>
,
<collectingCountry id="F31293BAFF92FFFEFA8FFB76FA6AB8B4" box="[1295,1432,1191,1217]" name="Switzerland" pageId="3" pageNumber="567">Switzerland</collectingCountry>
,
<collectingCountry id="F31293BAFF92FFFEFF47FB1AFEEAB890" box="[199,280,1227,1253]" name="Turkey" pageId="3" pageNumber="567">Turkey</collectingCountry>
,
<collectingCountry id="F31293BAFF92FFFEFEA1FB1AFE89B890" box="[289,379,1227,1253]" name="Ukraine" pageId="3" pageNumber="567">Ukraine</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF92FFFEFE08FB1AFDE9B890" author="Muona, J." box="[392,539,1227,1253]" pageId="3" pageNumber="567" refId="ref5680" refString="Muona, J. (2007) Family Eucnemidae Eschscholtz, 1829. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 4. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, pp. 81 - 87. https: // doi. org / 10.1163 / 9789004260894 _ 004" type="book" year="2007">Muona 2007</bibRefCitation>
), Georgia (
<bibRefCitation id="EF94AEDBFF92FFFEFD13FB1AFCD7B890" author="Horion, A." box="[659,805,1226,1253]" pageId="3" pageNumber="567" refId="ref4641" refString="Horion, A. (1953) Faunistik der Mitteleuropaischen Kafer. Bd. 3. Malacodermata, Sternoxia (Elateridae bis Throscidae). Eigenverlag Museum G. Frey, Munchen, 340 pp." type="book" year="1953">Horion 1953</bibRefCitation>
),
<collectingCountry id="F31293BAFF92FFFEFCB8FB1AFC65B890" box="[824,919,1227,1253]" name="Portugal" pageId="3" pageNumber="567">Portugal</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF92FFFEFC24FB1AFBDCB890" author="Cobos, A." box="[932,1070,1226,1253]" pageId="3" pageNumber="567" pagination="77 - 82" refId="ref4070" refString="Cobos, A. (1959) Materiales para el conocimiento de los Eucnemidae y Throscidae (Coleoptera) ibericos. Miscelanea Zoologica, 1 (2), 77 - 82." type="journal article" year="1959">Cobos 1959</bibRefCitation>
),
<collectingCountry id="F31293BAFF92FFFEFBC0FB1AFB59B890" box="[1088,1195,1227,1253]" name="Lithuania" pageId="3" pageNumber="567">Lithuania</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF92FFFEFB38FB1AFA62B890" author="Ferenca, R. &amp; Ivinskis, P. &amp; Tamutis, V." box="[1208,1424,1227,1253]" pageId="3" pageNumber="567" pagination="11 - 21" refId="ref4343" refString="Ferenca, R., Ivinskis, P. &amp; Tamutis, V. (2006) New and rare for Lithuania species of beetles (Coleoptera). New and Rare for Lithuania Insect Species. Records and Descriptions, 17, 11 - 21." type="journal article" year="2006">
Ferenca
<emphasis id="B9710F38FF92FFFEFA98FB1AFABAB890" box="[1304,1352,1227,1253]" italics="true" pageId="3" pageNumber="567">et al</emphasis>
. 2006
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),
<collectingCountry id="F31293BAFF92FFFEFF47FB3FFEDEB97D" box="[199,300,1262,1288]" name="Moldova" pageId="3" pageNumber="567">Moldova</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF92FFFEFEBAFB3FFDC5B97D" author="Derjanschi, V. &amp; Baban, E. &amp; Calestru, L. &amp; Stahi, N. &amp; Tugulea, C." box="[314,567,1262,1289]" pageId="3" pageNumber="567" refId="ref4141" refString="Derjanschi, V., Baban, E., Calestru, L., Stahi, N. &amp; Tugulea, C. (2016) Catalogue of the N. Zubowsky entomological collection. Bons Offices, ChiSinau, 296 pp. https: // doi. org / 10.53937 / 9789975871488" type="book" year="2016">
Derjanschi
<emphasis id="B9710F38FF92FFFEFE3CFB3EFE1CB97D" box="[444,494,1262,1288]" italics="true" pageId="3" pageNumber="567">et al</emphasis>
. 2016
</bibRefCitation>
),
<collectingCountry id="F31293BAFF92FFFEFDCBFB3EFD8AB97C" box="[587,632,1263,1289]" name="Iran" pageId="3" pageNumber="567">Iran</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF92FFFEFD06FB3FFC76B97C" author="Jarzabek-Muller, A." box="[646,900,1262,1289]" pageId="3" pageNumber="567" pagination="119 - 125" refId="ref4747" refString="Jarzabek-Muller, A. (2019) Neue Nachweise von Schienen- und Schnellkafern (Coleoptera: Eucnemidae, Elateridae) fur die palaarktische Fauna sowie eine Anmerkung zur Verbreitung von Hemicrepidius koenigi (Schwarz, 1897). Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen, 71, 119 - 125." type="journal article" year="2019">Jarzabek-Müller 2019</bibRefCitation>
).
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View file

@ -0,0 +1,148 @@
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Distribution in
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.
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Farashiani
<emphasis id="B9710F38FF92FFFEFDEAFD04FD69BE9B" box="[618,667,724,750]" italics="true" pageId="3" pageNumber="567">et al</emphasis>
. 2022
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),
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(
<bibRefCitation id="EF94AEDBFF92FFFEFC11FD05FB7EBE9A" author="Jarzabek-Muller, A." box="[913,1164,724,751]" pageId="3" pageNumber="567" pagination="119 - 125" refId="ref4747" refString="Jarzabek-Muller, A. (2019) Neue Nachweise von Schienen- und Schnellkafern (Coleoptera: Eucnemidae, Elateridae) fur die palaarktische Fauna sowie eine Anmerkung zur Verbreitung von Hemicrepidius koenigi (Schwarz, 1897). Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen, 71, 119 - 125." type="journal article" year="2019">Jarzabek-Müller 2019</bibRefCitation>
;
<bibRefCitation id="EF94AEDBFF92FFFEFB18FD05FA62BE9A" author="Farashiani, M. E. &amp; Varandi, H. B. &amp; Kazerani, F. &amp; Yarmand, H. &amp; Babaee, M. &amp; Thorn, S. &amp; Lange, F. &amp; Rafiei-Jahed, R. &amp; Muller, J. &amp; Amini, S." box="[1176,1424,724,751]" pageId="3" pageNumber="567" pagination="1063 - 1120" refId="ref4245" refString="Farashiani, M. E., Varandi, H. B., Kazerani, F., Yarmand, H., Babaee, M., Thorn, S., Lange, F., Rafiei-Jahed, R., Muller, J. &amp; Amini, S. (2022) A preliminary checklist of saproxylic beetles (Coleoptera) in the Hyrcanian forests of Iran, with distributional data. Check List, 18 (5), 1063 - 1120. https: // doi. org / 10.15560 / 18.5.1053" type="journal article" year="2022">
Farashiani
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. 2022
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).
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<emphasis id="B9710F38FF92FFFEFF17FD29FE63BF67" bold="true" box="[151,401,760,786]" pageId="3" pageNumber="567">General distribution.</emphasis>
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,Azerbaijan,
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,
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,
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,
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,
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,
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,
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(South European Territory),
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,
<collectingCountry id="F31293BAFF92FFFEFD44FCCDFCE6BF43" box="[708,788,796,822]" name="Turkey" pageId="3" pageNumber="567">Turkey</collectingCountry>
(
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), Georgia (
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),
<collectingCountry id="F31293BAFF92FFFEFB4AFCCDFADEBF43" box="[1226,1324,796,822]" name="Slovakia" pageId="3" pageNumber="567">Slovakia</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF92FFFEFABAFCCDFEF0BF2F" author="Smatana, I." pageId="3" pageNumber="567" refId="ref7123" refString="Smatana, I. (1996) Faunistic records. Entomological Problems, 27 (2), 148." type="journal volume" year="1996">Smatana 1996</bibRefCitation>
; Mertlik 2008),
<collectingCountry id="F31293BAFF92FFFEFE3EFC91FE0CBF2F" box="[446,510,832,858]" name="Spain" pageId="3" pageNumber="567">Spain</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF92FFFEFD8EFC91FCBFBF2F" author="Recalde Irurzun, J. I. &amp; Perez-Moreno, I. &amp; San Martin, A. F." box="[526,845,832,859]" pageId="3" pageNumber="567" pagination="397 - 401" refId="ref6757" refString="Recalde Irurzun, J. I., Perez-Moreno, I. &amp; San Martin, A. F. (2007) Crepidophorus mutilatus (Rosenhauer 1847), Aulonothroscus laticollis (Ribinsky 1897) e Isoriphis nigriceps (Mannerheim 1823): tres destacables Elateroidea de distribucion discontinua, nuevos para la fauna iberica (Coleoptera: Elateridae, Throscidae, Eucnemidae). Boletin Sociedad Entomologica Aragonesa, 41, 397 - 401." type="journal article" year="2007">
Recalde Irurzun
<emphasis id="B9710F38FF92FFFEFD4EFC90FCF3BF2F" box="[718,769,832,858]" italics="true" pageId="3" pageNumber="567">et al</emphasis>
. 2007
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),
<collectingCountry id="F31293BAFF92FFFEFCE2FC91FC4EBF2F" box="[866,956,832,858]" name="Ukraine" pageId="3" pageNumber="567">Ukraine</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF92FFFEFC4DFC91FB44BF2E" author="Chumak, M. V. &amp; Mateleshko, O. Y. &amp; Chumak, V. O. &amp; Varyvoda, M. V. &amp; Hrytsiuk, I. V. &amp; Zamoroka, A. M. &amp; Mirutenko, V. V. &amp; Nazarenko, V. I. &amp; Nikulina, T. V. &amp; Petrenko, A. A. &amp; Rizun, V. B. &amp; Seredyuk, G. V. &amp; Sergi, T. I. &amp; Tymochko, V. B. &amp; Turys, E. V. &amp; Yanytsky, T. P." box="[973,1206,832,859]" pageId="3" pageNumber="567" pagination="5 - 11" refId="ref3905" refString="Chumak, M. V., Mateleshko, O. Y., Chumak, V. O., Varyvoda, M. V., Hrytsiuk, I. V., Zamoroka, A. M., Mirutenko, V. V., Nazarenko, V. I., Nikulina, T. V., Petrenko, A. A., Rizun, V. B. Seredyuk, G. V., Sergi, T. I., Tymochko, V. B., Turys, E. V. &amp; Yanytsky, T. P. (2015) Taxonomic composition of the saproxylic Coleoptera (Arthropoda: Insecta) fauna in Uholka division of Carpathian Biosphere Reserve. Naukovij Visnik Uzhgorodskogo Universitetu [Scientific Bulletin of the Uzhgorod University], Seria Biologia [Series Biology], 38 - 39, 5 - 11. [in Ukrainian]" type="book chapter" year="2015">
Chumak
<emphasis id="B9710F38FF92FFFEFBB7FC90FB98BF2F" box="[1079,1130,832,858]" italics="true" pageId="3" pageNumber="567">et al</emphasis>
. 2015
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),
<collectingCountry id="F31293BAFF92FFFEFB4BFC91FAEBBF2F" box="[1227,1305,832,858]" name="Poland" pageId="3" pageNumber="567">Poland</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF92FFFEFAA9FC91FE68BF0B" author="Szoltys, H. &amp; Taszakowski, A." pageId="3" pageNumber="567" pagination="161 - 165" refId="ref7355" refString="Szoltys, H. &amp; Taszakowski, A. (2017) Isorhipis nigriceps (Mannerheim, 1823) - nowy dla fauny Polski gatunek chrzaszcza (Coleoptera: Eucnemidae). Acta entomologica silesiana, 25, 161 - 165." type="journal article" year="2017">Szołtys &amp; Taszakowski 2017</bibRefCitation>
),
<collectingCountry id="F31293BAFF92FFFEFE2DFCB4FE28BF0A" box="[429,474,869,895]" name="Iran" pageId="3" pageNumber="567">Iran</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF92FFFEFE69FCB5FD14BF0B" author="Jarzabek-Muller, A." box="[489,742,868,894]" pageId="3" pageNumber="567" pagination="119 - 125" refId="ref4747" refString="Jarzabek-Muller, A. (2019) Neue Nachweise von Schienen- und Schnellkafern (Coleoptera: Eucnemidae, Elateridae) fur die palaarktische Fauna sowie eine Anmerkung zur Verbreitung von Hemicrepidius koenigi (Schwarz, 1897). Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen, 71, 119 - 125." type="journal article" year="2019">Jarzabek-Müller 2019</bibRefCitation>
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View file

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<taxonomicName id="4C05A8A9FF92FFFEFF17FF0FFDB1BC8D" authority="(Laporte, 1835)" baseAuthorityName="Laporte" baseAuthorityYear="1835" box="[151,579,222,249]" class="Insecta" family="Eucnemidae" genus="Isorhipis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="567" phylum="Arthropoda" rank="species" species="melasoides">
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<emphasis id="B9710F38FF92FFFEFF17FF0FFE71BC8C" bold="true" box="[151,387,222,249]" italics="true" pageId="3" pageNumber="567">Isorhipis melasoides</emphasis>
(Laporte, 1835)
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</paragraph>
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<subSubSection id="C31F80A1FF92FFFEFF17FED3FEC1BEF6" pageId="3" pageNumber="567" type="distribution">
<paragraph id="8BBAD32AFF92FFFEFF17FED3FB86BD35" blockId="3.[151,1436,150,643]" pageId="3" pageNumber="567">
<emphasis id="B9710F38FF92FFFEFF17FED3FE60BD69" bold="true" box="[151,402,258,284]" pageId="3" pageNumber="567">
Distribution in
<collectingCountry id="F31293BAFF92FFFEFED7FED3FE7CBD69" box="[343,398,258,284]" name="Iran" pageId="3" pageNumber="567">Iran</collectingCountry>
.
</emphasis>
<collectingRegion id="49C11DC8FF92FFFEFE1DFED3FDF3BD69" box="[413,513,258,284]" country="Iran" name="Golestan" pageId="3" pageNumber="567">Golestan</collectingRegion>
(
<bibRefCitation id="EF94AEDBFF92FFFEFD93FED3FCDDBD69" author="Rafiei-Jahed, R. &amp; Kavousi, M. R. &amp; Farashiani, M. E. &amp; Sagheb-Talebi, K. &amp; Muller, J. &amp; Babanezhad, M." box="[531,815,258,285]" pageId="3" pageNumber="567" pagination="94 - 109" refId="ref6581" refString="Rafiei-Jahed, R., Kavousi, M. R., Farashiani, M. E., Sagheb-Talebi, K., Muller, J. &amp; Babanezhad, M. (2021) A comparison of the efficiency of sampling methods beetles (Insecta: Coleoptera) Shast-kalateh beech forest, Golestan province. Iranian Journal of Forest and Range Protection Research, 19 (1), 94 - 109. [in Persian, English summary]" type="journal article" year="2021">
Rafiei-Jahed
<emphasis id="B9710F38FF92FFFEFD2DFED2FD10BD69" box="[685,738,258,284]" italics="true" pageId="3" pageNumber="567">et al</emphasis>
. 2021
</bibRefCitation>
, under the family
<taxonomicName id="4C05A8A9FF92FFFEFB8AFED3FB8DBD69" authorityName="Leach" authorityYear="1815" box="[1034,1151,258,284]" class="Insecta" family="Elateridae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="567" phylum="Arthropoda" rank="family">Elateridae</taxonomicName>
;
<bibRefCitation id="EF94AEDBFF92FFFEFB0EFED3FA62BD69" author="Farashiani, M. E. &amp; Varandi, H. B. &amp; Kazerani, F. &amp; Yarmand, H. &amp; Babaee, M. &amp; Thorn, S. &amp; Lange, F. &amp; Rafiei-Jahed, R. &amp; Muller, J. &amp; Amini, S." box="[1166,1424,258,285]" pageId="3" pageNumber="567" pagination="1063 - 1120" refId="ref4245" refString="Farashiani, M. E., Varandi, H. B., Kazerani, F., Yarmand, H., Babaee, M., Thorn, S., Lange, F., Rafiei-Jahed, R., Muller, J. &amp; Amini, S. (2022) A preliminary checklist of saproxylic beetles (Coleoptera) in the Hyrcanian forests of Iran, with distributional data. Check List, 18 (5), 1063 - 1120. https: // doi. org / 10.15560 / 18.5.1053" type="journal article" year="2022">
Farashiani
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. 2022
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),
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(present paper),
<collectingRegion id="49C11DC8FF92FFFEFE51FEF7FDA9BD35" box="[465,603,294,320]" country="Iran" name="Mazandaran" pageId="3" pageNumber="567">Mazandaran</collectingRegion>
(
<bibRefCitation id="EF94AEDBFF92FFFEFDE9FEF7FC97BD35" author="Jarzabek-Muller, A." box="[617,869,294,320]" pageId="3" pageNumber="567" pagination="119 - 125" refId="ref4747" refString="Jarzabek-Muller, A. (2019) Neue Nachweise von Schienen- und Schnellkafern (Coleoptera: Eucnemidae, Elateridae) fur die palaarktische Fauna sowie eine Anmerkung zur Verbreitung von Hemicrepidius koenigi (Schwarz, 1897). Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen, 71, 119 - 125." type="journal article" year="2019">Jarzabek-Müller 2019</bibRefCitation>
;
<bibRefCitation id="EF94AEDBFF92FFFEFCF1FEF7FB9ABD35" author="Farashiani, M. E. &amp; Varandi, H. B. &amp; Kazerani, F. &amp; Yarmand, H. &amp; Babaee, M. &amp; Thorn, S. &amp; Lange, F. &amp; Rafiei-Jahed, R. &amp; Muller, J. &amp; Amini, S." box="[881,1128,294,321]" pageId="3" pageNumber="567" pagination="1063 - 1120" refId="ref4245" refString="Farashiani, M. E., Varandi, H. B., Kazerani, F., Yarmand, H., Babaee, M., Thorn, S., Lange, F., Rafiei-Jahed, R., Muller, J. &amp; Amini, S. (2022) A preliminary checklist of saproxylic beetles (Coleoptera) in the Hyrcanian forests of Iran, with distributional data. Check List, 18 (5), 1063 - 1120. https: // doi. org / 10.15560 / 18.5.1053" type="journal article" year="2022">
Farashiani
<emphasis id="B9710F38FF92FFFEFC6CFEF6FBEFBD35" box="[1004,1053,294,320]" italics="true" pageId="3" pageNumber="567">et al</emphasis>
. 2022
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).
</paragraph>
<paragraph id="8BBAD32AFF92FFFEFF17FE9BFC37BDFD" blockId="3.[151,1436,150,643]" pageId="3" pageNumber="567">
<emphasis id="B9710F38FF92FFFEFF17FE9BFE4EBD11" bold="true" box="[151,444,330,356]" pageId="3" pageNumber="567">New material examined.</emphasis>
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<collectingRegion id="49C11DC8FF92FFFEFE43FE9BFD8FBD11" box="[451,637,330,356]" country="Iran" name="Gilan" pageId="3" pageNumber="567">Guilan province</collectingRegion>
,
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,
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, 78 (
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the modern Gregorian calendar)
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<materialsCitation id="3B6DD977FF92FFFEFAC5FE9BFC33BDFD" collectingDate="1916-04" collectorName="B. S. Iljin" location="Russian Academy of Sciences" municipality="Zoological Institute of Russian Academy" pageId="3" pageNumber="567" specimenCount="1" stateProvince="Guilan province">
<date id="FFBBF5EAFF92FFFEFAC5FE9BFA6ABD11" box="[1349,1432,330,356]" pageId="3" pageNumber="567" value="1916-04">
<collectingDate id="EFFF0C02FF92FFFEFAC5FE9BFA6ABD11" box="[1349,1432,330,356]" pageId="3" pageNumber="567" value="1916-04">iv.1916</collectingDate>
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,
<collectorName id="26F0B6FCFF92FFFEFF47FEBFFEDFBDFD" box="[199,301,366,392]" pageId="3" pageNumber="567">B.S. Iljin</collectorName>
leg.,
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Zoological Institute of
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of Sciences
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.
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<paragraph id="8BBAD32AFF92FFFEFF17FE40FEC1BEF6" blockId="3.[151,1436,150,643]" pageId="3" pageNumber="567">
<emphasis id="B9710F38FF92FFFEFF17FE40FE65BDDE" bold="true" box="[151,407,401,427]" pageId="3" pageNumber="567">General distribution.</emphasis>
<collectingCountry id="F31293BAFF92FFFEFE21FE43FE04BDD9" box="[417,502,402,428]" name="Austria" pageId="3" pageNumber="567">Austria</collectingCountry>
,
<collectingCountry id="F31293BAFF92FFFEFD85FE43FCD1BDD9" box="[517,803,402,428]" name="Bosnia and Herzegovina" pageId="3" pageNumber="567">Bosnia and Herzegovina</collectingCountry>
,
<collectingCountry id="F31293BAFF92FFFEFCB2FE43FC67BDD9" box="[818,917,402,428]" name="Bulgaria" pageId="3" pageNumber="567">Bulgaria</collectingCountry>
,
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,
<collectingCountry id="F31293BAFF92FFFEFB89FE43FB30BDD9" box="[1033,1218,402,428]" name="Czech Republic" pageId="3" pageNumber="567">Czech Republic</collectingCountry>
,
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,
<collectingCountry id="F31293BAFF92FFFEFAAFFE43FA6ABDD9" box="[1327,1432,402,428]" name="Germany" pageId="3" pageNumber="567">Germany</collectingCountry>
,
<collectingCountry id="F31293BAFF92FFFEFF47FE67FED8BDA5" box="[199,298,438,464]" name="Hungary" pageId="3" pageNumber="567">Hungary</collectingCountry>
,
<collectingCountry id="F31293BAFF92FFFEFEBCFE64FE82BDBA" box="[316,368,437,463]" name="Italy" pageId="3" pageNumber="567">Italy</collectingCountry>
,
<collectingCountry id="F31293BAFF92FFFEFE02FE64FE21BDBA" box="[386,467,437,463]" name="Poland" pageId="3" pageNumber="567">Poland</collectingCountry>
,
<collectingCountry id="F31293BAFF92FFFEFE64FE64FDBEBDBA" box="[484,588,437,463]" name="Romania" pageId="3" pageNumber="567">Romania</collectingCountry>
,
<collectingCountry id="F31293BAFF92FFFEFDDEFE64FD5BBDBA" box="[606,681,437,463]" name="Russia" pageId="3" pageNumber="567">Russia</collectingCountry>
(Central European Territory),
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,
<collectingCountry id="F31293BAFF92FFFEFB10FE64FB12BDBA" box="[1168,1248,437,463]" name="Turkey" pageId="3" pageNumber="567">Turkey</collectingCountry>
(
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), Azerbaijan (
<bibRefCitation id="EF94AEDBFF92FFFEFECFFE08FE16BD81" author="Koenig, E." box="[335,484,473,500]" pageId="3" pageNumber="567" pagination="339 - 403" refId="ref4797" refString="Koenig, E. (1899) Coleoptera Caucasica. In: Radde, G. (Ed.), Die Sammlungen des Kaukasischen Museums. Band 1. Zoologie. Typographie der Kanzelei des Landeschefs, Tiflis, pp. 339 - 403." type="book chapter" year="1899">Koenig 1899</bibRefCitation>
;
<bibRefCitation id="EF94AEDBFF92FFFEFE6EFE08FD3BBD81" author="Varshalovitch, A. A." box="[494,713,473,500]" pageId="3" pageNumber="567" refId="ref7444" refString="Varshalovitch, A. A. (1935) Novyj v SSSR vreditel' drevesiny - bukovyj buril'shchik (Jsorrhipis melasoides Lap. - Cast.) [Jsorrhipis melasoides Lap. - Gast. A New Pest of Wood in the USSR]. Zashchita rastenij, 1, 147. [in Russian]" type="journal volume" year="1935">Varshalovitch 1935</bibRefCitation>
),
<collectingCountry id="F31293BAFF92FFFEFD5CFE08FCC4BD86" box="[732,822,473,499]" name="Ukraine" pageId="3" pageNumber="567">Ukraine</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF92FFFEFCC3FE08FC25BD86" author="Roubal, J." box="[835,983,473,499]" pageId="3" pageNumber="567" refId="ref6957" refString="Roubal, J. (1936) Katalog Coleopter (Brouku) Slovenska a Podkarpatske Rusi na zaklade bionomickem a zoogeografickem a spolu systematicky doplnek Ganglbauerovych &quot; Die Kafer Mitteleuropa &quot; a Reitterovy &quot; Fauna germanica &quot;. Dil II. Ucena Spolecnost Safarikova v Bratislave, Bratislava, viii + 434 pp." type="book" year="1936">Roubal 1936</bibRefCitation>
;
<bibRefCitation id="EF94AEDBFF92FFFEFC61FE08FB54BD81" author="Mateleshko, A." box="[993,1190,473,500]" pageId="3" pageNumber="567" pagination="155 - 163" refId="ref5269" refString="Mateleshko, A. (2009) New records of beetles (Insecta, Coleoptera) from the Ukrainian Carpathians region. Naukovij Visnik Uzhgorodskogo Universitetu [Scientific Bulletin of the Uzhgorod University], Seria Biologia [Series Biology], 25, 155 - 163. [in Ukrainian]" type="journal article" year="2009">Mateleshko 2009</bibRefCitation>
),
<collectingCountry id="F31293BAFF92FFFEFB38FE08FACCBD86" box="[1208,1342,473,499]" name="Switzerland" pageId="3" pageNumber="567">Switzerland</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF92FFFEFACBFE08FEF0BE6D" author="Horion, A." pageId="3" pageNumber="567" refId="ref4641" refString="Horion, A. (1953) Faunistik der Mitteleuropaischen Kafer. Bd. 3. Malacodermata, Sternoxia (Elateridae bis Throscidae). Eigenverlag Museum G. Frey, Munchen, 340 pp." type="book" year="1953">Horion 1953</bibRefCitation>
;
<bibRefCitation id="EF94AEDBFF92FFFEFE91FE2CFDC3BE62" author="Chittaro, Y. &amp; Blanc, M." box="[273,561,509,535]" pageId="3" pageNumber="567" pagination="91 - 114" refId="ref3864" refString="Chittaro, Y. &amp; Blanc, M. (2012) Liste commentee des Cerophytidae, Elateridae, Eucnemidae et Throscidae (Coleoptera) de Suisse. Bulletin de la Societe Entomologique Suisse, 85, 91 - 114." type="journal article" year="2012">Chittaro and Blanc 2012</bibRefCitation>
),
<collectingCountry id="F31293BAFF92FFFEFDC8FE2CFD61BE62" box="[584,659,509,535]" name="Russia" pageId="3" pageNumber="567">Russia</collectingCountry>
(South European Territory) (
<bibRefCitation id="EF94AEDBFF92FFFEFC65FE2CFB7DBE62" author="Mamaev, B. M." box="[997,1167,509,535]" pageId="3" pageNumber="567" pagination="136 - 155" refId="ref5191" refString="Mamaev, B. M. (1976) Morfologicheskie tipy lichinok zhukov-drevoedov (Coleoptera, Eucnemidae) i ikh evolyutsionnoe znachenie [Morphological types of eucnemid larvae (Coleoptera, Eucnemidae) and their evolutionary significance]. In: Mamaev, B. M. (Ed.), Evolyutsionnaya Morfologiya Lichinok Nasekomykh [Evolutionary Morphology of Insect Larvae]. Nauka, Moscow, pp. 136 - 155. [in Russian]" type="book chapter" year="1976">Mamaev 1976</bibRefCitation>
),
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(
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),
<collectingCountry id="F31293BAFF92FFFEFE96FDF0FEA4BE4E" box="[278,342,545,571]" name="Spain" pageId="3" pageNumber="567">Spain</collectingCountry>
(
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),
<collectingCountry id="F31293BAFF92FFFEFDEFFDF0FD4CBE4E" box="[623,702,545,571]" name="Greece" pageId="3" pageNumber="567">Greece</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF92FFFEFD4DFDF0FC56BE4E" author="Mertlik, J. &amp; Jenis, I. &amp; Zbuzek, B." box="[717,932,545,572]" pageId="3" pageNumber="567" pagination="1 - 6" refId="ref5539" refString="Mertlik, J., Jenis, I. &amp; Zbuzek, B. (2009) New records on the distribution of some species of the family Melasidae (Coleoptera) - II. Elateridarium, 3, 1 - 6." type="journal article" year="2009">
Mertlik
<emphasis id="B9710F38FF92FFFEFCA9FDF3FCA9BE4E" box="[809,859,545,571]" italics="true" pageId="3" pageNumber="567">et al</emphasis>
. 2009
</bibRefCitation>
),
<collectingCountry id="F31293BAFF92FFFEFC37FDF0FBD0BE4E" box="[951,1058,545,571]" name="Lithuania" pageId="3" pageNumber="567">Lithuania</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF92FFFEFBB1FDF0FAB9BE4E" author="Ferenca, R. &amp; Tamutis, V." box="[1073,1355,545,571]" pageId="3" pageNumber="567" pagination="32 - 39" refId="ref4390" refString="Ferenca, R. &amp; Tamutis, V. 2009. Data on seventeen beetle (Coleoptera) species new for Lithuanian fauna. New and Rare for Lithuania Insect Species. Records and Descriptions, 21, 32 - 39." type="book chapter" year="2009">Ferenca &amp; Tamutis 2009</bibRefCitation>
),
<collectingCountry id="F31293BAFF92FFFEFADEFDF0FEE7BE2A" name="United Kingdom" pageId="3" pageNumber="567">Great Britain</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF92FFFEFEA4FD94FE0FBE2A" author="Mendel, H. &amp; Jeffery, P. &amp; Pledger, M. J." box="[292,509,581,608]" pageId="3" pageNumber="567" pagination="41 - 43" refId="ref5321" refString="Mendel, H., Jeffery, P. &amp; Pledger, M. J. (2011) Isorhipis melasoides (Laporte, 1835) (Eucnemidae) breeding and probably established in the British Isles. The Coleopterist, 20 (2), 41 - 43." type="journal article" year="2011">
Mendel
<emphasis id="B9710F38FF92FFFEFE02FD97FE46BE2A" box="[386,436,581,607]" italics="true" pageId="3" pageNumber="567">et al</emphasis>
. 2011
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),
<collectingCountry id="F31293BAFF92FFFEFD91FD94FDABBE2A" box="[529,601,581,607]" name="Serbia" pageId="3" pageNumber="567">Serbia</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF92FFFEFDE8FD94FCB4BE2A" author="Nemeth, T. &amp; Laus, B. &amp; Tallosi, B." box="[616,838,581,608]" pageId="3" pageNumber="567" pagination="47 - 56" refId="ref6216" refString="Nemeth, T., Laus, B. &amp; Tallosi, B. (2017) New distributional data on Elateroidea for Albania, Bulgaria, Croatia, Greece, Macedonia, Montenegro and Serbia (Coleoptera: Elateridae, Eucnemidae). Folia Entomologica Hungarica, 78, 47 - 56. https: // doi. org / 10.17112 / FoliaEntHung. 2017.78.47" type="journal article" year="2017">
Németh
<emphasis id="B9710F38FF92FFFEFD4AFD97FD0EBE2A" box="[714,764,581,607]" italics="true" pageId="3" pageNumber="567">et al</emphasis>
. 2017
</bibRefCitation>
),
<collectingCountry id="F31293BAFF92FFFEFCDAFD97FC75BE15" box="[858,903,582,608]" name="Iran" pageId="3" pageNumber="567">Iran</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF92FFFEFC16FD94FB61BE2A" author="Jarzabek-Muller, A." box="[918,1171,581,607]" pageId="3" pageNumber="567" pagination="119 - 125" refId="ref4747" refString="Jarzabek-Muller, A. (2019) Neue Nachweise von Schienen- und Schnellkafern (Coleoptera: Eucnemidae, Elateridae) fur die palaarktische Fauna sowie eine Anmerkung zur Verbreitung von Hemicrepidius koenigi (Schwarz, 1897). Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen, 71, 119 - 125." type="journal article" year="2019">Jarzabek-Müller 2019</bibRefCitation>
),
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Troukens
<emphasis id="B9710F38FF92FFFEFA08FD97FF2FBEF6" italics="true" pageId="3" pageNumber="567">et al</emphasis>
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Distribution in
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Farashiani
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Farashiani
<emphasis id="B9710F38FF93FFFFFAE3F86AFA67BBA0" box="[1379,1429,1979,2005]" italics="true" pageId="2" pageNumber="566">et al</emphasis>
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Distribution in
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East Azarbaijan (
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<emphasis id="B9710F38FF93FFFFFD3DF9DAFD1CBA50" box="[701,750,1547,1573]" italics="true" pageId="2" pageNumber="566">et al</emphasis>
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<emphasis id="B9710F38FF93FFFFFF17F9FFFE6ABA3D" bold="true" box="[151,408,1582,1608]" pageId="2" pageNumber="566">General distribution.</emphasis>
<collectingCountry id="F31293BAFF93FFFFFE22F9FEFE05BA3C" box="[418,503,1583,1609]" name="Austria" pageId="2" pageNumber="566">Austria</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFD86F9FEFDACBA3C" box="[518,606,1583,1609]" name="Belarus" pageId="2" pageNumber="566">Belarus</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFDEEF9FEFC7FBA3C" box="[622,909,1583,1609]" name="Bosnia and Herzegovina" pageId="2" pageNumber="566">Bosnia and Herzegovina</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFC1DF9FEFC00BA3C" box="[925,1010,1583,1609]" name="Croatia" pageId="2" pageNumber="566">Croatia</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFB82F9FEFBABBA3C" box="[1026,1113,1583,1609]" name="Estonia" pageId="2" pageNumber="566">Estonia</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFBE8F9FEFB32BA3C" box="[1128,1216,1583,1609]" name="Finland" pageId="2" pageNumber="566">Finland</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFB50F9FEFAEDBA3C" box="[1232,1311,1583,1609]" name="France" pageId="2" pageNumber="566">France</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFAAFF9FEFA6ABA3C" box="[1327,1432,1583,1609]" name="Germany" pageId="2" pageNumber="566">Germany</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFF47F982FEEBBA18" box="[199,281,1619,1645]" name="Greece" pageId="2" pageNumber="566">Greece</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFEA2F982FE77BA18" box="[290,389,1619,1645]" name="Hungary" pageId="2" pageNumber="566">Hungary</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFE0DF982FE33BA18" box="[397,449,1619,1645]" name="Italy" pageId="2" pageNumber="566">Italy</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFE4AF982FDE6BA18" box="[458,532,1619,1645]" name="Latvia" pageId="2" pageNumber="566">Latvia</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFD9DF982FD7FBA18" box="[541,653,1619,1645]" name="Mongolia" pageId="2" pageNumber="566">Mongolia</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFD16F982FD15BA18" box="[662,743,1619,1645]" name="Poland" pageId="2" pageNumber="566">Poland</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFD6FF982FCA5BA18" box="[751,855,1619,1645]" name="Romania" pageId="2" pageNumber="566">Romania</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFCE0F982FC59BA18" box="[864,939,1619,1645]" name="Russia" pageId="2" pageNumber="566">Russia</collectingCountry>
(Central European Territory, North European Territory, West Siberia),
<collectingCountry id="F31293BAFF93FFFFFE5DF9A6FDB0BAE4" box="[477,578,1655,1681]" name="Slovakia" pageId="2" pageNumber="566">Slovakia</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFDCDF9A6FD54BAE4" box="[589,678,1655,1681]" name="Sweden" pageId="2" pageNumber="566">Sweden</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF93FFFFFD35F9A6FCB8BAE4" author="Muona, J." box="[693,842,1655,1681]" pageId="2" pageNumber="566" refId="ref5680" refString="Muona, J. (2007) Family Eucnemidae Eschscholtz, 1829. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 4. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, pp. 81 - 87. https: // doi. org / 10.1163 / 9789004260894 _ 004" type="book" year="2007">Muona 2007</bibRefCitation>
),
<collectingCountry id="F31293BAFF93FFFFFCDEF9A6FC4ABAE4" box="[862,952,1655,1681]" name="Norway" pageId="2" pageNumber="566">Norway</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF93FFFFFC48F9A6FB79BAE4" author="Silfverberg, H." box="[968,1163,1655,1681]" pageId="2" pageNumber="566" pagination="67 - 95" refId="ref7143" refString="Silfverberg, H. (2002) Further additions and corrections to Enumeratio Coleopterorum Fennoscandiae, Daniae et Baltiae. Sahlbergia, 7, 67 - 95." type="journal article" year="2002">Silfverberg 2002</bibRefCitation>
),
<collectingCountry id="F31293BAFF93FFFFFB1FF9A6FB0BBAE4" box="[1183,1273,1655,1681]" name="Ukraine" pageId="2" pageNumber="566">Ukraine</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF93FFFFFA88F9A6FEF1BAC0" author="Drogvalenko, A. N." pageId="2" pageNumber="566" pagination="9 - 19" refId="ref4195" refString="Drogvalenko, A. N. (2002) The new and rare species of beetles (Insecta: Coleoptera) for fauna of Ukraine. Information 2. The Kharkov Entomological Society Gazette, 9 (1 - 2), 9 - 19. [in Russian]" type="journal article" year="2002">Drogvalenko 2002</bibRefCitation>
),
<collectingCountry id="F31293BAFF93FFFFFE97F94AFE90BAC0" box="[279,354,1691,1717]" name="Russia" pageId="2" pageNumber="566">Russia</collectingCountry>
(South European Territory) (
<bibRefCitation id="EF94AEDBFF93FFFFFD2AF94AFC52BAC0" author="Negrobov, S. O. &amp; Tsurikov, M. N. &amp; Logvinovsky, V. D. &amp; Fomichev, A. I. &amp; Prokin, A. A. &amp; Gilmutdinov, K. S." box="[682,928,1690,1717]" pageId="2" pageNumber="566" pagination="534 - 673" refId="ref6072" refString="Negrobov, S. O., Tsurikov, M. N., Logvinovsky, V. D., Fomichev, A. I., Prokin, A. A. &amp; Gilmutdinov, K. S. (2005) Otryad Coleoptera [Order Coleoptera]. In: Negrobov, O. P. (Ed.), Kadastr bespozvonochnykh zhivotnykh Voronezhskoj oblasti [Cadastre of invertebrate animals of the Voronezh region]. Voronezh State University, Voronezh, pp. 534 - 673. [in Russian]" type="book chapter" year="2005">
Negrobov
<emphasis id="B9710F38FF93FFFFFCA3F94AFCA7BAC0" box="[803,853,1691,1717]" italics="true" pageId="2" pageNumber="566">et al</emphasis>
. 2005
</bibRefCitation>
),
<collectingCountry id="F31293BAFF93FFFFFC34F94AFC0CBAC0" box="[948,1022,1691,1717]" name="Serbia" pageId="2" pageNumber="566">Serbia</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFB89F94AFBABBAC0" box="[1033,1113,1691,1717]" name="Turkey" pageId="2" pageNumber="566">Turkey</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF93FFFFFBE8F94AFAB0BAC0" author="Mertlik, J. &amp; Jenis, I. &amp; Zbuzek, B." box="[1128,1346,1691,1717]" pageId="2" pageNumber="566" pagination="92 - 96" refId="ref5498" refString="Mertlik, J., Jenis, I. &amp; Zbuzek, B. (2007) New records on the distribution of some species of the family Eucnemidae (Coleoptera). Elateridarium, 1, 92 - 96." type="journal article" year="2007">
Mertlik
<emphasis id="B9710F38FF93FFFFFB46F94AFB0ABAC0" box="[1222,1272,1691,1717]" italics="true" pageId="2" pageNumber="566">et al</emphasis>
. 2007
</bibRefCitation>
),
<collectingCountry id="F31293BAFF93FFFFFAD6F94AFEDEBAAC" name="Czech Republic" pageId="2" pageNumber="566">Czech Republic</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF93FFFFFEBDF96EFE3ABAAC" author="Vavra, J. C." box="[317,456,1727,1753]" pageId="2" pageNumber="566" pagination="303 - 304" refId="ref7499" refString="Vavra, J. C. (2008) Faunistic Records from the Czech Republic - 270. Coleoptera: Eucnemidae, Nitidulidae. Klapalekiana, 44, 303 - 304." type="journal article" year="2008">Vávra 2008</bibRefCitation>
),
<collectingCountry id="F31293BAFF93FFFFFE5EF96EFDBBBAAC" box="[478,585,1727,1753]" name="Lithuania" pageId="2" pageNumber="566">Lithuania</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF93FFFFFDDAF96EFC6FBAAC" author="Ostrauskas, H. &amp; Ferenca, R." box="[602,925,1727,1753]" pageId="2" pageNumber="566" pagination="41 - 46" refId="ref6485" refString="Ostrauskas, H. &amp; Ferenca, R. (2010) Beetles (Coleoptera) caught in traps baited with pheromones for Dendroctonus rufipennis (Kirby) (Curculionidae: Scolytinae) in Lithuania. Ekologija, 56 (1 - 2), 41 - 46. https: // doi. org / 10.2478 / v 10055 - 010 - 0006 - 8" type="journal article" year="2010">Ostrauskas &amp; Ferenca 2010</bibRefCitation>
),
<collectingCountry id="F31293BAFF93FFFFFC33F96EFC01BAAC" box="[947,1011,1727,1753]" name="Spain" pageId="2" pageNumber="566">Spain</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF93FFFFFB84F96EFAF3BAAC" author="Recalde Irurzun, J. I." box="[1028,1281,1727,1753]" pageId="2" pageNumber="566" pagination="195 - 196" refId="ref6713" refString="Recalde Irurzun, J. I. (2012) Rhacopus sahlbergi (Mannerheim, 1823): un nuevo eucnemido para la fauna iberica (Coleoptera: Elateroidea: Eucnemidae). Boletin Sociedad Entomologica Aragonesa, 51, 195 - 196." type="journal article" year="2012">Recalde Irurzun 2012</bibRefCitation>
),
<collectingCountry id="F31293BAFF93FFFFFA96F96EFA6EBAAC" box="[1302,1436,1727,1753]" name="Switzerland" pageId="2" pageNumber="566">Switzerland</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF93FFFFFF4EF932FE26BA88" author="Chittaro, Y. &amp; Blanc, M." box="[206,468,1763,1789]" pageId="2" pageNumber="566" pagination="91 - 114" refId="ref3864" refString="Chittaro, Y. &amp; Blanc, M. (2012) Liste commentee des Cerophytidae, Elateridae, Eucnemidae et Throscidae (Coleoptera) de Suisse. Bulletin de la Societe Entomologique Suisse, 85, 91 - 114." type="journal article" year="2012">Chittaro &amp; Blanc 2012</bibRefCitation>
),
<collectingCountry id="F31293BAFF93FFFFFE67F932FDB5BA88" box="[487,583,1763,1789]" name="Bulgaria" pageId="2" pageNumber="566">Bulgaria</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF93FFFFFDD6F932FCC1BA88" author="Nemeth, T. &amp; Laus, B. &amp; Tallosi, B." box="[598,819,1763,1789]" pageId="2" pageNumber="566" pagination="47 - 56" refId="ref6216" refString="Nemeth, T., Laus, B. &amp; Tallosi, B. (2017) New distributional data on Elateroidea for Albania, Bulgaria, Croatia, Greece, Macedonia, Montenegro and Serbia (Coleoptera: Elateridae, Eucnemidae). Folia Entomologica Hungarica, 78, 47 - 56. https: // doi. org / 10.17112 / FoliaEntHung. 2017.78.47" type="journal article" year="2017">
Németh
<emphasis id="B9710F38FF93FFFFFD38F932FD1BBA88" box="[696,745,1763,1789]" italics="true" pageId="2" pageNumber="566">et al</emphasis>
. 2017
</bibRefCitation>
),
<collectingCountry id="F31293BAFF93FFFFFCC6F932FC81BA88" box="[838,883,1763,1789]" name="Iran" pageId="2" pageNumber="566">Iran</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF93FFFFFC02F932FB98BA88" author="Bunalski, M. &amp; Navaiian, M. &amp; Hosseini Boldaji, S. A. &amp; Hava, J. &amp; Angelini, F. &amp; Sakenin, H. &amp; Samin, N." box="[898,1130,1763,1789]" pageId="2" pageNumber="566" pagination="114 - 121" refId="ref3756" refString="Bunalski, M., Navaiian, M., Hosseini Boldaji, S. A., Hava, J., Angelini, F., Sakenin, H. &amp; Samin, N. (2023) A faunistic study of Coleoptera of Iran with nine new country records. Wiadomosci Entomologiczne, 42, 114 - 121." type="journal article" year="2023">
Bunalski
<emphasis id="B9710F38FF93FFFFFC6EF932FBEDBA88" box="[1006,1055,1763,1789]" italics="true" pageId="2" pageNumber="566">et al</emphasis>
. 2023
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,153 @@
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<paragraph id="8BBAD32AFF93FFFFFF17FB8FFDC9B80D" blockId="2.[151,1437,1082,1469]" box="[151,571,1118,1145]" pageId="2" pageNumber="566">
<heading id="D0F26446FF93FFFFFF17FB8FFDC9B80D" bold="true" box="[151,571,1118,1145]" fontSize="11" level="1" pageId="2" pageNumber="566" reason="1">
<taxonomicName id="4C05A8A9FF93FFFFFF17FB8FFDC9B80D" authority="(Olexa, 1962)" baseAuthorityName="Olexa" baseAuthorityYear="1962" box="[151,571,1118,1145]" class="Insecta" family="Eucnemidae" genus="Rhacopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="566" phylum="Arthropoda" rank="species" species="bodemeyeri">
<emphasis id="B9710F38FF93FFFFFF17FB8FFDC9B80D" bold="true" box="[151,571,1118,1145]" pageId="2" pageNumber="566">
<emphasis id="B9710F38FF93FFFFFF17FB8FFE67B80C" bold="true" box="[151,405,1118,1145]" italics="true" pageId="2" pageNumber="566">Rhacopus bodemeyeri</emphasis>
(
<bibRefCitation id="EF94AEDBFF93FFFFFE24FB8FFDC6B80D" author="Olexa, A." box="[420,564,1118,1144]" pageId="2" pageNumber="566" pagination="43 - 44" refId="ref6360" refString="Olexa, A. (1962) Dirrhagus ferrugineus Reitt. et D. bodemeyeri n. sp. (Col. Eucnemidae). Bulletin de la Societe Entomologique de Mulhouse, 1962, 43 - 44." type="journal article" year="1962">Olexa, 1962</bibRefCitation>
)
</emphasis>
</taxonomicName>
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</paragraph>
</subSubSection>
<subSubSection id="C31F80A1FF93FFFFFF17FB53FDA4B8B4" pageId="2" pageNumber="566" type="distribution">
<paragraph id="8BBAD32AFF93FFFFFF17FB53FC36B8E8" blockId="2.[151,1437,1082,1469]" box="[151,964,1154,1181]" pageId="2" pageNumber="566">
<emphasis id="B9710F38FF93FFFFFF17FB53FE79B8E8" bold="true" box="[151,395,1154,1181]" pageId="2" pageNumber="566">
Distribution in
<collectingCountry id="F31293BAFF93FFFFFECFFB52FE74B8E8" box="[335,390,1155,1181]" name="Iran" pageId="2" pageNumber="566">Iran</collectingCountry>
.
</emphasis>
<collectingRegion id="49C11DC8FF93FFFFFE11FB52FE2CB8E8" box="[401,478,1155,1181]" country="Iran" name="Tehran" pageId="2" pageNumber="566">Tehran</collectingRegion>
(
<typeStatus id="54BE6D88FF93FFFFFE6DFB52FDD4B8E8" box="[493,550,1155,1181]" pageId="2" pageNumber="566">Type</typeStatus>
locality: Mt.
<collectingRegion id="49C11DC8FF93FFFFFD3FFB52FCEBB8E8" box="[703,793,1155,1181]" country="Iran" name="Alborz" pageId="2" pageNumber="566">Elbourz</collectingRegion>
) (
<bibRefCitation id="EF94AEDBFF93FFFFFCAFFB52FC45B8E8" author="Olexa, A." box="[815,951,1155,1181]" pageId="2" pageNumber="566" pagination="43 - 44" refId="ref6360" refString="Olexa, A. (1962) Dirrhagus ferrugineus Reitt. et D. bodemeyeri n. sp. (Col. Eucnemidae). Bulletin de la Societe Entomologique de Mulhouse, 1962, 43 - 44." type="journal article" year="1962">Olexa 1962</bibRefCitation>
).
</paragraph>
<paragraph id="8BBAD32AFF93FFFFFF17FB77FDA4B8B4" blockId="2.[151,1437,1082,1469]" box="[151,598,1190,1217]" pageId="2" pageNumber="566">
<emphasis id="B9710F38FF93FFFFFF17FB77FE61B8B5" bold="true" box="[151,403,1190,1216]" pageId="2" pageNumber="566">General distribution.</emphasis>
Endemic to
<collectingCountry id="F31293BAFF93FFFFFDA2FB76FDA1B8B4" box="[546,595,1191,1217]" name="Iran" pageId="2" pageNumber="566">Iran</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C31F80A1FF93FFFFFF17FB1AFD46B9C8" pageId="2" pageNumber="566" type="discussion">
<paragraph id="8BBAD32AFF93FFFFFF17FB1AFD46B9C8" blockId="2.[151,1437,1082,1469]" pageId="2" pageNumber="566">
<emphasis id="B9710F38FF93FFFFFF17FB1AFEF2B890" bold="true" box="[151,256,1227,1253]" pageId="2" pageNumber="566">Remark.</emphasis>
The identity and generic level placement of this taxon remains uncertain.
<bibRefCitation id="EF94AEDBFF93FFFFFBCCFB1AFB10B890" author="Olexa, A." box="[1100,1250,1226,1253]" pageId="2" pageNumber="566" pagination="155 - 164" refId="ref6446" refString="Olexa, A. (1975) Reclassification of the genus Dirhagus and related genera (Coleoptera - Eucnemidae). Entomologische Blatter fur Biologie und Systematik der Kafer, 71 (3), 155 - 164." type="journal article" year="1975">Olexa (1975)</bibRefCitation>
proposed a new subgenus
<taxonomicName id="4C05A8A9FF93FFFFFEB9FB3EFD7BB97C" authority="Olexa, 1975" authorityName="Olexa" authorityYear="1975" box="[313,649,1262,1289]" class="Insecta" family="Eucnemidae" genus="Pseudorhacopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="566" phylum="Arthropoda" rank="genus">
<emphasis id="B9710F38FF93FFFFFEB9FB3EFE01B97C" box="[313,499,1263,1289]" italics="true" pageId="2" pageNumber="566">Pseudorhacopus</emphasis>
<bibRefCitation id="EF94AEDBFF93FFFFFE7DFB3EFD7BB97C" author="Olexa, A." box="[509,649,1262,1289]" pageId="2" pageNumber="566" pagination="155 - 164" refId="ref6446" refString="Olexa, A. (1975) Reclassification of the genus Dirhagus and related genera (Coleoptera - Eucnemidae). Entomologische Blatter fur Biologie und Systematik der Kafer, 71 (3), 155 - 164." type="journal article" year="1975">Olexa, 1975</bibRefCitation>
</taxonomicName>
within the genus
<taxonomicName id="4C05A8A9FF93FFFFFCDCFB3EFC38B97C" authorityName="Hampe" authorityYear="1855" box="[860,970,1263,1289]" class="Insecta" family="Eucnemidae" genus="Rhacopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="566" phylum="Arthropoda" rank="genus">
<emphasis id="B9710F38FF93FFFFFCDCFB3EFC38B97C" box="[860,970,1263,1289]" italics="true" pageId="2" pageNumber="566">Rhacopus</emphasis>
</taxonomicName>
for
<emphasis id="B9710F38FF93FFFFFC7FFB3EFB81B97C" box="[1023,1139,1263,1289]" italics="true" pageId="2" pageNumber="566">Dirrhagus</emphasis>
<taxonomicName id="4C05A8A9FF93FFFFFBFDFB3EFA6EB97C" authority="Reitter, 1889" authorityName="Reitter" authorityYear="1889" baseAuthorityName="Reitter" baseAuthorityYear="1889" box="[1149,1436,1263,1289]" class="Insecta" family="Eucnemidae" genus="Rhacopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="566" phylum="Arthropoda" rank="species" species="ferrugineus">
<emphasis id="B9710F38FF93FFFFFBFDFB3EFB0CB97C" box="[1149,1278,1263,1289]" italics="true" pageId="2" pageNumber="566">ferrugineus</emphasis>
<bibRefCitation id="EF94AEDBFF93FFFFFA88FB3EFA6EB97C" author="Reitter, E." box="[1288,1436,1263,1289]" pageId="2" pageNumber="566" pagination="273 - 288" refId="ref6834" refString="Reitter, E. (1889) Neue Coleopteren aus Europa, den angrenzenden Landern und Sibirien, mit Bemerkungen uber bekannte Arten. Siebenter Theil. Deutsche entomologische Zeitschrift, 1889, 273 - 288." type="journal article" year="1889">Reitter, 1889</bibRefCitation>
</taxonomicName>
(
<typeStatus id="54BE6D88FF93FFFFFF4EFAC2FEF2B958" box="[206,256,1299,1325]" pageId="2" pageNumber="566">type</typeStatus>
species) and
<emphasis id="B9710F38FF93FFFFFE21FAC2FDE7B958" box="[417,533,1299,1325]" italics="true" pageId="2" pageNumber="566">Dirrhagus</emphasis>
<taxonomicName id="4C05A8A9FF93FFFFFD9FFAC2FCC9B958" authority="Olexa, 1962" authorityName="Olexa" authorityYear="1962" box="[543,827,1299,1325]" class="Insecta" family="Eucnemidae" genus="Rhacopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="566" phylum="Arthropoda" rank="species" species="bodemeyeri">
<emphasis id="B9710F38FF93FFFFFD9FFAC2FD50B958" box="[543,674,1299,1325]" italics="true" pageId="2" pageNumber="566">bodemeyeri</emphasis>
<bibRefCitation id="EF94AEDBFF93FFFFFD2CFAC2FCC9B958" author="Olexa, A." box="[684,827,1299,1325]" pageId="2" pageNumber="566" pagination="43 - 44" refId="ref6360" refString="Olexa, A. (1962) Dirrhagus ferrugineus Reitt. et D. bodemeyeri n. sp. (Col. Eucnemidae). Bulletin de la Societe Entomologique de Mulhouse, 1962, 43 - 44." type="journal article" year="1962">Olexa, 1962</bibRefCitation>
</taxonomicName>
. Subsequently, the
<typeStatus id="54BE6D88FF93FFFFFB9BFAC2FBB9B958" box="[1051,1099,1299,1325]" pageId="2" pageNumber="566">type</typeStatus>
species of the subgenus was transferred to
<taxonomicName id="4C05A8A9FF93FFFFFEE4FAE6FD39B924" authority="Fleutiaux, 1935" authorityName="Fleutiaux" authorityYear="1935" box="[356,715,1334,1361]" class="Insecta" family="Eucnemidae" genus="Dirrhagofarsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="566" phylum="Arthropoda" rank="genus">
<emphasis id="B9710F38FF93FFFFFEE4FAE6FDE0B924" box="[356,530,1335,1361]" italics="true" pageId="2" pageNumber="566">Dirrhagofarsus</emphasis>
Fleutiaux, 1935
</taxonomicName>
, and the name
<taxonomicName id="4C05A8A9FF93FFFFFCF1FAE6FBD9B924" authorityName="Olexa" authorityYear="1975" box="[881,1067,1335,1361]" class="Insecta" family="Eucnemidae" genus="Pseudorhacopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="566" phylum="Arthropoda" rank="genus">
<emphasis id="B9710F38FF93FFFFFCF1FAE6FBD9B924" box="[881,1067,1335,1361]" italics="true" pageId="2" pageNumber="566">Pseudorhacopus</emphasis>
</taxonomicName>
was synonymized with the latter (
<bibRefCitation id="EF94AEDBFF93FFFFFF4EFA8AFE87B900" author="Kovalev, A. V." box="[206,373,1371,1397]" pageId="2" pageNumber="566" pagination="277 - 290" refId="ref4896" refString="Kovalev, A. V. (2016) Two new species of Microrhagus from the Russian Far East with notes on some Palaearctic Dirhagini (Coleoptera: Eucnemidae). Zoosystematica Rossica, 25 (2), 277 - 290. https: // doi. org / 10.31610 / zsr / 2016.25.2.277" type="journal article" year="2016">Kovalev 2016</bibRefCitation>
). It is likely that
<taxonomicName id="4C05A8A9FF93FFFFFDBAFA8AFD13B900" authorityName="Olexa" authorityYear="1962" box="[570,737,1371,1397]" class="Insecta" family="Eucnemidae" genus="Rhacopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="566" phylum="Arthropoda" rank="species" species="bodemeyeri">
<emphasis id="B9710F38FF93FFFFFDBAFA8AFD13B900" box="[570,737,1371,1397]" italics="true" pageId="2" pageNumber="566">D. bodemeyeri</emphasis>
</taxonomicName>
may also belong to
<taxonomicName id="4C05A8A9FF93FFFFFC4DFA8AFB89B900" authorityName="Fleutiaux" authorityYear="1935" box="[973,1147,1371,1397]" class="Insecta" family="Eucnemidae" genus="Dirrhagofarsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="566" phylum="Arthropoda" rank="genus">
<emphasis id="B9710F38FF93FFFFFC4DFA8AFB89B900" box="[973,1147,1371,1397]" italics="true" pageId="2" pageNumber="566">Dirrhagofarsus</emphasis>
</taxonomicName>
(see also
<bibRefCitation id="EF94AEDBFF93FFFFFB6FFA8AFA6AB900" author="Kovalev, A. V." box="[1263,1432,1371,1397]" pageId="2" pageNumber="566" pagination="277 - 290" refId="ref4896" refString="Kovalev, A. V. (2016) Two new species of Microrhagus from the Russian Far East with notes on some Palaearctic Dirhagini (Coleoptera: Eucnemidae). Zoosystematica Rossica, 25 (2), 277 - 290. https: // doi. org / 10.31610 / zsr / 2016.25.2.277" type="journal article" year="2016">Kovalev, 2016</bibRefCitation>
; Muona, 2023), but here we maintain the classification proposed by
<bibRefCitation id="EF94AEDBFF93FFFFFC3CFAAEFBA3B9EC" author="Olexa, A." box="[956,1105,1406,1433]" pageId="2" pageNumber="566" pagination="155 - 164" refId="ref6446" refString="Olexa, A. (1975) Reclassification of the genus Dirhagus and related genera (Coleoptera - Eucnemidae). Entomologische Blatter fur Biologie und Systematik der Kafer, 71 (3), 155 - 164." type="journal article" year="1975">Olexa (1975)</bibRefCitation>
until examination of the
<typeStatus id="54BE6D88FF93FFFFFAECFAAEFA6EB9EC" box="[1388,1436,1407,1433]" pageId="2" pageNumber="566">type</typeStatus>
material of
<taxonomicName id="4C05A8A9FF93FFFFFEC8FA72FE1FB9C8" authorityName="Olexa" authorityYear="1962" box="[328,493,1443,1469]" class="Insecta" family="Eucnemidae" genus="Rhacopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="566" phylum="Arthropoda" rank="species" species="bodemeyeri">
<emphasis id="B9710F38FF93FFFFFEC8FA72FE1FB9C8" box="[328,493,1443,1469]" italics="true" pageId="2" pageNumber="566">D. bodemeyeri</emphasis>
</taxonomicName>
is made possible.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,184 @@
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<paragraph id="8BBAD32AFF93FFFFFF17FD2BFD88BF61" blockId="2.[151,1437,726,1041]" box="[151,634,762,789]" pageId="2" pageNumber="566">
<heading id="D0F26446FF93FFFFFF17FD2BFD88BF61" bold="true" box="[151,634,762,789]" fontSize="11" level="1" pageId="2" pageNumber="566" reason="1">
<taxonomicName id="4C05A8A9FF93FFFFFF17FD2BFD88BF61" authority="(Fabricius, 1792)" baseAuthorityName="Fabricius" baseAuthorityYear="1792" box="[151,634,762,789]" class="Insecta" family="Eucnemidae" genus="Microrhagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="566" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B9710F38FF93FFFFFF17FD2BFD88BF61" bold="true" box="[151,634,762,789]" pageId="2" pageNumber="566">
<emphasis id="B9710F38FF93FFFFFF17FD2BFE5ABF60" bold="true" box="[151,424,762,789]" italics="true" pageId="2" pageNumber="566">Microrhagus pygmaeus</emphasis>
(Fabricius, 1792)
</emphasis>
</taxonomicName>
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</paragraph>
</subSubSection>
<subSubSection id="C31F80A1FF93FFFFFF17FCCFFB64B864" pageId="2" pageNumber="566" type="distribution">
<paragraph id="8BBAD32AFF93FFFFFF17FCCFFBC4BF28" blockId="2.[151,1437,726,1041]" pageId="2" pageNumber="566">
<emphasis id="B9710F38FF93FFFFFF17FCCFFE7EBF4C" bold="true" box="[151,396,798,825]" pageId="2" pageNumber="566">
Distribution in
<collectingCountry id="F31293BAFF93FFFFFED0FCCEFE75BF4C" box="[336,391,799,825]" name="Iran" pageId="2" pageNumber="566">Iran</collectingCountry>
.
</emphasis>
East Azarbaijan (
<bibRefCitation id="EF94AEDBFF93FFFFFDD3FCCEFCCEBF4C" author="Bunalski, M. &amp; Navaiian, M. &amp; Hosseini Boldaji, S. A. &amp; Hava, J. &amp; Angelini, F. &amp; Sakenin, H. &amp; Samin, N." box="[595,828,799,825]" pageId="2" pageNumber="566" pagination="114 - 121" refId="ref3756" refString="Bunalski, M., Navaiian, M., Hosseini Boldaji, S. A., Hava, J., Angelini, F., Sakenin, H. &amp; Samin, N. (2023) A faunistic study of Coleoptera of Iran with nine new country records. Wiadomosci Entomologiczne, 42, 114 - 121." type="journal article" year="2023">
Bunalski
<emphasis id="B9710F38FF93FFFFFD40FCCEFD03BF4C" box="[704,753,799,825]" italics="true" pageId="2" pageNumber="566">et al</emphasis>
. 2023
</bibRefCitation>
),
<collectingRegion id="49C11DC8FF93FFFFFCCFFCCEFC2BBF4C" box="[847,985,799,825]" country="Iran" name="Mazandaran" pageId="2" pageNumber="566">Mazandaran</collectingRegion>
(
<bibRefCitation id="EF94AEDBFF93FFFFFC69FCCEFB13BF4C" author="Farashiani, M. E. &amp; Varandi, H. B. &amp; Kazerani, F. &amp; Yarmand, H. &amp; Babaee, M. &amp; Thorn, S. &amp; Lange, F. &amp; Rafiei-Jahed, R. &amp; Muller, J. &amp; Amini, S." box="[1001,1249,799,825]" pageId="2" pageNumber="566" pagination="1063 - 1120" refId="ref4245" refString="Farashiani, M. E., Varandi, H. B., Kazerani, F., Yarmand, H., Babaee, M., Thorn, S., Lange, F., Rafiei-Jahed, R., Muller, J. &amp; Amini, S. (2022) A preliminary checklist of saproxylic beetles (Coleoptera) in the Hyrcanian forests of Iran, with distributional data. Check List, 18 (5), 1063 - 1120. https: // doi. org / 10.15560 / 18.5.1053" type="journal article" year="2022">
Farashiani
<emphasis id="B9710F38FF93FFFFFBE6FCCEFB6ABF4C" box="[1126,1176,799,825]" italics="true" pageId="2" pageNumber="566">et al</emphasis>
. 2022
</bibRefCitation>
, as
<taxonomicName id="4C05A8A9FF93FFFFFA8CFCCEFE5BBF28" authority="(Olexa, 1963)" baseAuthorityName="Olexa" baseAuthorityYear="1963" class="Insecta" family="Eucnemidae" genus="Microrhagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="566" phylum="Arthropoda" rank="species" species="palmi">
<emphasis id="B9710F38FF93FFFFFA8CFCCEFEF5BF28" italics="true" pageId="2" pageNumber="566">Microrhagus palmi</emphasis>
(Olexa, 1963)
</taxonomicName>
currently considered as junior synonym of
<taxonomicName id="4C05A8A9FF93FFFFFC11FC92FBD4BF29" baseAuthorityName="Fabricius" baseAuthorityYear="1792" box="[913,1062,835,860]" class="Insecta" family="Eucnemidae" genus="Microrhagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="566" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B9710F38FF93FFFFFC11FC92FBD4BF29" box="[913,1062,835,860]" italics="true" pageId="2" pageNumber="566">M. pygmaeus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BBAD32AFF93FFFFFF17FCB7FB64B864" blockId="2.[151,1437,726,1041]" pageId="2" pageNumber="566">
<emphasis id="B9710F38FF93FFFFFF17FCB7FE67BFF5" bold="true" box="[151,405,870,896]" pageId="2" pageNumber="566">General distribution.</emphasis>
<collectingCountry id="F31293BAFF93FFFFFE1DFCB6FE07BFF4" box="[413,501,871,897]" name="Belarus" pageId="2" pageNumber="566">Belarus</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFD82FCB6FD97BFF4" box="[514,613,871,897]" name="Belgium" pageId="2" pageNumber="566">Belgium</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFDF2FCB6FC79BFF4" box="[626,907,871,897]" name="Bosnia and Herzegovina" pageId="2" pageNumber="566">Bosnia and Herzegovina</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFC18FCB6FC09BFF4" box="[920,1019,871,897]" name="Bulgaria" pageId="2" pageNumber="566">Bulgaria</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFB88FCB6FBAFBFF4" box="[1032,1117,871,897]" name="Croatia" pageId="2" pageNumber="566">Croatia</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFBEAFCB6FAD2BFF4" box="[1130,1312,871,897]" name="Czech Republic" pageId="2" pageNumber="566">Czech Republic</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFAADFCB6FA6ABFF4" box="[1325,1432,871,897]" name="Denmark" pageId="2" pageNumber="566">Denmark</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFF47FC5AFEECBFD0" box="[199,286,907,933]" name="Estonia" pageId="2" pageNumber="566">Estonia</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFEADFC5AFE77BFD0" box="[301,389,907,933]" name="Finland" pageId="2" pageNumber="566">Finland</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFE15FC5AFE16BFD0" box="[405,484,907,933]" name="France" pageId="2" pageNumber="566">France</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFE73FC5AFDAEBFD0" box="[499,604,907,933]" name="Germany" pageId="2" pageNumber="566">Germany</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFDECFC5AFCF5BFD0" box="[620,775,907,933]" name="United Kingdom" pageId="2" pageNumber="566">Great Britain</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFC96FC5AFC8BBFD0" box="[790,889,907,933]" name="Hungary" pageId="2" pageNumber="566">Hungary</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFC08FC5AFC4EBFD0" box="[904,956,907,933]" name="Italy" pageId="2" pageNumber="566">Italy</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFC4CFC5AFBE4BFD0" box="[972,1046,907,933]" name="Latvia" pageId="2" pageNumber="566">Latvia</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFBA6FC5AFB66BFD0" box="[1062,1172,907,933]" name="Lithuania" pageId="2" pageNumber="566">Lithuania</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFB23FC5AFADFBFD0" box="[1187,1325,907,933]" name="Netherlands" pageId="2" pageNumber="566">Netherlands</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFABDFC5AFA6ABFD0" box="[1341,1432,907,933]" name="Norway" pageId="2" pageNumber="566">Norway</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFF47FC7EFEEABFBC" box="[199,280,943,969]" name="Poland" pageId="2" pageNumber="566">Poland</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFEA0FC7EFE7ABFBC" box="[288,392,943,969]" name="Romania" pageId="2" pageNumber="566">Romania</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFE10FC7EFE29BFBC" box="[400,475,943,969]" name="Russia" pageId="2" pageNumber="566">Russia</collectingCountry>
(Central European Territory, North European Territory),
<collectingCountry id="F31293BAFF93FFFFFBC6FC7EFB59BFBC" box="[1094,1195,943,969]" name="Slovakia" pageId="2" pageNumber="566">Slovakia</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFB33FC7EFAFCBFBC" box="[1203,1294,943,969]" name="Sweden" pageId="2" pageNumber="566">Sweden</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFA96FC7EFA6EBFBC" box="[1302,1436,943,969]" name="Switzerland" pageId="2" pageNumber="566">Switzerland</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF93FFFFFF4EFC02FE94BF98" author="Muona, J." box="[206,358,979,1005]" pageId="2" pageNumber="566" refId="ref5680" refString="Muona, J. (2007) Family Eucnemidae Eschscholtz, 1829. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 4. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, pp. 81 - 87. https: // doi. org / 10.1163 / 9789004260894 _ 004" type="book" year="2007">Muona 2007</bibRefCitation>
),
<collectingCountry id="F31293BAFF93FFFFFEFCFC02FE35BF98" box="[380,455,979,1005]" name="Russia" pageId="2" pageNumber="566">Russia</collectingCountry>
(South European Territory) (
<bibRefCitation id="EF94AEDBFF93FFFFFC98FC02FC43BF98" author="Roubal, J." box="[792,945,979,1005]" pageId="2" pageNumber="566" pagination="124 - 126" refId="ref6901" refString="Roubal, J. (1912) Koleopterologicke vysledky me cesty na Kavkaz v cervenci r. 1910. Quid novi de Coleopterorum Caucasi ad occidentem vergentis fauna in meo itinere Julio mense anni 1910 suscepto cognoverim. VI. Krasnaja Poljana. Eucnemidae. Casopis Ceske Spolecnosti Entomologicke, 9, 124 - 126." type="journal article" year="1912">Roubal 1912</bibRefCitation>
),
<collectingCountry id="F31293BAFF93FFFFFC46FC02FBEFBF98" box="[966,1053,979,1005]" name="Algeria" pageId="2" pageNumber="566">Algeria</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFBA9FC02FB8CBF98" box="[1065,1150,979,1005]" name="Austria" pageId="2" pageNumber="566">Austria</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFB0DFC02FB15BF98" box="[1165,1255,979,1005]" name="Ukraine" pageId="2" pageNumber="566">Ukraine</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF93FFFFFB79FC02FA7DBF98" author="Horion, A." box="[1273,1423,978,1005]" pageId="2" pageNumber="566" refId="ref4641" refString="Horion, A. (1953) Faunistik der Mitteleuropaischen Kafer. Bd. 3. Malacodermata, Sternoxia (Elateridae bis Throscidae). Eigenverlag Museum G. Frey, Munchen, 340 pp." type="book" year="1953">Horion 1953</bibRefCitation>
),
<collectingCountry id="F31293BAFF93FFFFFF47FC26FEF5B864" box="[199,263,1015,1041]" name="Spain" pageId="2" pageNumber="566">Spain</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF93FFFFFE95FC26FDFFB864" author="Recalde Irurzun, J. I." box="[277,525,1015,1041]" pageId="2" pageNumber="566" pagination="233 - 252" refId="ref6663" refString="Recalde Irurzun, J. I. (2008) Elementos para el conocimiento de los eucnemidos del norte de Espana y actualizacion del catalogo de especies ibericas (Coleoptera: Elateroidea: Eucnemidae). Heteropterus Revista de Entomologia, 8 (2), 233 - 252." type="journal article" year="2008">Recalde Irurzun 2008</bibRefCitation>
),
<collectingCountry id="F31293BAFF93FFFFFDA1FC26FDAEB864" box="[545,604,1015,1041]" name="Syria" pageId="2" pageNumber="566">Syria</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF93FFFFFDEAFC26FCB0B864" author="Mertlik, J. &amp; Jenis, I. &amp; Zbuzek, B." box="[618,834,1015,1041]" pageId="2" pageNumber="566" pagination="1 - 6" refId="ref5539" refString="Mertlik, J., Jenis, I. &amp; Zbuzek, B. (2009) New records on the distribution of some species of the family Melasidae (Coleoptera) - II. Elateridarium, 3, 1 - 6." type="journal article" year="2009">
Mertlik
<emphasis id="B9710F38FF93FFFFFD47FC26FD0AB864" box="[711,760,1015,1041]" italics="true" pageId="2" pageNumber="566">et al</emphasis>
. 2009
</bibRefCitation>
),
<collectingCountry id="F31293BAFF93FFFFFCD5FC26FC70B864" box="[853,898,1015,1041]" name="Iran" pageId="2" pageNumber="566">Iran</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF93FFFFFC11FC26FB79B864" author="Farashiani, M. E. &amp; Varandi, H. B. &amp; Kazerani, F. &amp; Yarmand, H. &amp; Babaee, M. &amp; Thorn, S. &amp; Lange, F. &amp; Rafiei-Jahed, R. &amp; Muller, J. &amp; Amini, S." box="[913,1163,1015,1041]" pageId="2" pageNumber="566" pagination="1063 - 1120" refId="ref4245" refString="Farashiani, M. E., Varandi, H. B., Kazerani, F., Yarmand, H., Babaee, M., Thorn, S., Lange, F., Rafiei-Jahed, R., Muller, J. &amp; Amini, S. (2022) A preliminary checklist of saproxylic beetles (Coleoptera) in the Hyrcanian forests of Iran, with distributional data. Check List, 18 (5), 1063 - 1120. https: // doi. org / 10.15560 / 18.5.1053" type="journal article" year="2022">
Farashiani
<emphasis id="B9710F38FF93FFFFFB8FFC26FBB2B864" box="[1039,1088,1015,1041]" italics="true" pageId="2" pageNumber="566">et al</emphasis>
. 2022
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,159 @@
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<heading id="D0F26446FF93FFFFFF17FE66FD52BDA5" bold="true" box="[151,672,438,465]" fontSize="11" level="1" pageId="2" pageNumber="566" reason="1">
<taxonomicName id="4C05A8A9FF93FFFFFF17FE66FD52BDA5" authority="(Piller &amp; Mitterpacher, 1783)" baseAuthorityName="Piller &amp; Mitterpacher" baseAuthorityYear="1783" box="[151,672,438,465]" class="Insecta" family="Eucnemidae" genus="Farsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="566" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="B9710F38FF93FFFFFF17FE66FD52BDA5" bold="true" box="[151,672,438,465]" pageId="2" pageNumber="566">
<emphasis id="B9710F38FF93FFFFFF17FE66FECFBDA4" bold="true" box="[151,317,438,465]" italics="true" pageId="2" pageNumber="566">Farsus dubius</emphasis>
(Piller &amp; Mitterpacher, 1783)
</emphasis>
</taxonomicName>
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</paragraph>
</subSubSection>
<subSubSection id="C31F80A1FF93FFFFFF17FE0BFE30BEDC" pageId="2" pageNumber="566" type="distribution">
<paragraph id="8BBAD32AFF93FFFFFF17FE0BFCB9BD80" blockId="2.[151,1436,402,681]" box="[151,843,474,501]" pageId="2" pageNumber="566">
<emphasis id="B9710F38FF93FFFFFF17FE0BFE79BD80" bold="true" box="[151,395,474,501]" pageId="2" pageNumber="566">
Distribution in
<collectingCountry id="F31293BAFF93FFFFFECFFE0AFE74BD80" box="[335,390,475,501]" name="Iran" pageId="2" pageNumber="566">Iran</collectingCountry>
.
</emphasis>
West Azarbaijan (
<bibRefCitation id="EF94AEDBFF93FFFFFDD7FE0AFCB2BD80" author="Bunalski, M. &amp; Navaiian, M. &amp; Hosseini Boldaji, S. A. &amp; Hava, J. &amp; Angelini, F. &amp; Sakenin, H. &amp; Samin, N." box="[599,832,475,501]" pageId="2" pageNumber="566" pagination="114 - 121" refId="ref3756" refString="Bunalski, M., Navaiian, M., Hosseini Boldaji, S. A., Hava, J., Angelini, F., Sakenin, H. &amp; Samin, N. (2023) A faunistic study of Coleoptera of Iran with nine new country records. Wiadomosci Entomologiczne, 42, 114 - 121." type="journal article" year="2023">
Bunalski
<emphasis id="B9710F38FF93FFFFFD44FE0AFD07BD80" box="[708,757,475,501]" italics="true" pageId="2" pageNumber="566">et al</emphasis>
. 2023
</bibRefCitation>
).
</paragraph>
<paragraph id="8BBAD32AFF93FFFFFF17FE2FFE30BEDC" blockId="2.[151,1436,402,681]" pageId="2" pageNumber="566">
<emphasis id="B9710F38FF93FFFFFF17FE2FFE63BE6D" bold="true" box="[151,401,510,536]" pageId="2" pageNumber="566">General distribution.</emphasis>
<collectingCountry id="F31293BAFF93FFFFFE13FE2EFE1ABE6C" box="[403,488,511,537]" name="Austria" pageId="2" pageNumber="566">Austria</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFE71FE2EFD51BE6C" box="[497,675,511,537]" name="Czech Republic" pageId="2" pageNumber="566">Czech Republic</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFD2CFE2EFD09BE6C" box="[684,763,511,537]" name="France" pageId="2" pageNumber="566">France</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFC83FE2EFC94BE6C" box="[771,870,511,537]" name="Hungary" pageId="2" pageNumber="566">Hungary</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFCEFFE2EFC51BE6C" box="[879,931,511,537]" name="Italy" pageId="2" pageNumber="566">Italy</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFC2BFE2EFBE1BE6C" box="[939,1043,511,537]" name="Romania" pageId="2" pageNumber="566">Romania</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFB9CFE2EFB95BE6C" box="[1052,1127,511,537]" name="Russia" pageId="2" pageNumber="566">Russia</collectingCountry>
(South European Territory),
<collectingCountry id="F31293BAFF93FFFFFF47FDF2FE27BE48" box="[199,469,547,573]" name="Serbia" pageId="2" pageNumber="566">Serbia and Montenegro</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFE62FDF2FDD7BE48" box="[482,549,547,573]" name="Spain" pageId="2" pageNumber="566">Spain</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFDB2FDF2FD9FBE48" box="[562,621,547,573]" name="Syria" pageId="2" pageNumber="566">Syria</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF93FFFFFDFEFDF2FCE6BE48" author="Muona, J." box="[638,788,547,573]" pageId="2" pageNumber="566" refId="ref5680" refString="Muona, J. (2007) Family Eucnemidae Eschscholtz, 1829. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 4. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, pp. 81 - 87. https: // doi. org / 10.1163 / 9789004260894 _ 004" type="book" year="2007">Muona 2007</bibRefCitation>
),
<collectingCountry id="F31293BAFF93FFFFFCAAFDF2FBB7BE48" box="[810,1093,547,573]" name="Bosnia and Herzegovina" pageId="2" pageNumber="566">Bosnia and Herzegovina</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFBD2FDF2FB56BE48" box="[1106,1188,547,573]" name="Greece" pageId="2" pageNumber="566">Greece</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFB31FDF2FAC8BE48" box="[1201,1338,547,573]" name="Serbia" pageId="2" pageNumber="566">Montenegro</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF93FFFFFACBFDF2FEF0BE14" author="Horion, A." pageId="2" pageNumber="566" refId="ref4641" refString="Horion, A. (1953) Faunistik der Mitteleuropaischen Kafer. Bd. 3. Malacodermata, Sternoxia (Elateridae bis Throscidae). Eigenverlag Museum G. Frey, Munchen, 340 pp." type="book" year="1953">Horion 1953</bibRefCitation>
),
<collectingCountry id="F31293BAFF93FFFFFE98FD96FE9ABE14" box="[280,360,583,609]" name="Turkey" pageId="2" pageNumber="566">Turkey</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF93FFFFFEF8FD96FDFFBE14" author="Horion, A." box="[376,525,582,609]" pageId="2" pageNumber="566" refId="ref4641" refString="Horion, A. (1953) Faunistik der Mitteleuropaischen Kafer. Bd. 3. Malacodermata, Sternoxia (Elateridae bis Throscidae). Eigenverlag Museum G. Frey, Munchen, 340 pp." type="book" year="1953">Horion 1953</bibRefCitation>
;
<bibRefCitation id="EF94AEDBFF93FFFFFD9BFD96FCE9BE14" author="Mertlik, J. &amp; Platia, G." box="[539,795,583,609]" pageId="2" pageNumber="566" pagination="1 - 40" refId="ref5582" refString="Mertlik, J. &amp; Platia, G. (2008) Catalogue of the family Cebrionidae, Elateridae, Lissomidae, Melasidae and Throscidae (Coleoptera) from Turkey. Elateridarium, 2, 1 - 40." type="journal article" year="2008">Mertlik &amp; Platia 2008</bibRefCitation>
),
<collectingCountry id="F31293BAFF93FFFFFCB0FD96FC6ABE14" box="[816,920,583,609]" name="Morocco" pageId="2" pageNumber="566">Morocco</collectingCountry>
,
<collectingCountry id="F31293BAFF93FFFFFC25FD96FBF6BE14" box="[933,1028,583,609]" name="Portugal" pageId="2" pageNumber="566">Portugal</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF93FFFFFB95FD96FB51BE14" author="Cobos, A." box="[1045,1187,582,609]" pageId="2" pageNumber="566" pagination="77 - 82" refId="ref4070" refString="Cobos, A. (1959) Materiales para el conocimiento de los Eucnemidae y Throscidae (Coleoptera) ibericos. Miscelanea Zoologica, 1 (2), 77 - 82." type="journal article" year="1959">Cobos 1959</bibRefCitation>
),
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Mertlik
<emphasis id="B9710F38FF93FFFFFA08FD96FF2FBEF0" italics="true" pageId="2" pageNumber="566">et al</emphasis>
. 2009
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),
<collectingCountry id="F31293BAFF93FFFFFEBDFDBAFE64BEF0" box="[317,406,619,645]" name="Albania" pageId="2" pageNumber="566">Albania</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF93FFFFFE27FDBAFDB0BEF0" author="Mertlik, J." box="[423,578,619,645]" pageId="2" pageNumber="566" pagination="55 - 58" refId="ref5468" refString="Mertlik, J. (2011) Nove udaje o rozsireni tri druhu z celedi Lissomidae a Melasidae (Coleoptera). Elateridarium, 5, 55 - 58." type="journal article" year="2011">Mertlik 2011</bibRefCitation>
),
<collectingCountry id="F31293BAFF93FFFFFDD7FDBAFD2FBEF0" box="[599,733,619,645]" name="Switzerland" pageId="2" pageNumber="566">Switzerland</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF93FFFFFD6EFDBAFC0EBEF0" author="Chittaro, Y. &amp; Blanc, M." box="[750,1020,619,645]" pageId="2" pageNumber="566" pagination="91 - 114" refId="ref3864" refString="Chittaro, Y. &amp; Blanc, M. (2012) Liste commentee des Cerophytidae, Elateridae, Eucnemidae et Throscidae (Coleoptera) de Suisse. Bulletin de la Societe Entomologique Suisse, 85, 91 - 114." type="journal article" year="2012">Chittaro &amp; Blanc 2012</bibRefCitation>
),
<collectingCountry id="F31293BAFF93FFFFFB92FDBAFB96BEF0" box="[1042,1124,619,645]" name="Croatia" pageId="2" pageNumber="566">Croatia</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF93FFFFFBF5FDBAFAABBEF0" author="Nemeth, T. &amp; Laus, B. &amp; Tallosi, B." box="[1141,1369,619,645]" pageId="2" pageNumber="566" pagination="47 - 56" refId="ref6216" refString="Nemeth, T., Laus, B. &amp; Tallosi, B. (2017) New distributional data on Elateroidea for Albania, Bulgaria, Croatia, Greece, Macedonia, Montenegro and Serbia (Coleoptera: Elateridae, Eucnemidae). Folia Entomologica Hungarica, 78, 47 - 56. https: // doi. org / 10.17112 / FoliaEntHung. 2017.78.47" type="journal article" year="2017">
Németh
<emphasis id="B9710F38FF93FFFFFB59FDBAFAE6BEF0" box="[1241,1300,619,645]" italics="true" pageId="2" pageNumber="566">et al.</emphasis>
2017
</bibRefCitation>
),
<collectingCountry id="F31293BAFF93FFFFFAEFFDBAFA6EBEF0" box="[1391,1436,619,645]" name="Iran" pageId="2" pageNumber="566">Iran</collectingCountry>
(
<bibRefCitation id="EF94AEDBFF93FFFFFF4EFD5EFE45BEDC" author="Bunalski, M. &amp; Navaiian, M. &amp; Hosseini Boldaji, S. A. &amp; Hava, J. &amp; Angelini, F. &amp; Sakenin, H. &amp; Samin, N." box="[206,439,655,681]" pageId="2" pageNumber="566" pagination="114 - 121" refId="ref3756" refString="Bunalski, M., Navaiian, M., Hosseini Boldaji, S. A., Hava, J., Angelini, F., Sakenin, H. &amp; Samin, N. (2023) A faunistic study of Coleoptera of Iran with nine new country records. Wiadomosci Entomologiczne, 42, 114 - 121." type="journal article" year="2023">
Bunalski
<emphasis id="B9710F38FF93FFFFFEBBFD5EFE9EBEDC" box="[315,364,655,681]" italics="true" pageId="2" pageNumber="566">et al</emphasis>
. 2023
</bibRefCitation>
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View file

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<emphasis id="B9710F38FF93FFFFFF17FF6BFE2ABCA1" bold="true" box="[151,472,186,213]" italics="true" pageId="2" pageNumber="566">Dirrhagofarsus ferrugineus</emphasis>
(
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Distribution in
<collectingCountry id="F31293BAFF93FFFFFED5FF0EFE7EBC8C" box="[341,396,223,249]" name="Iran" pageId="2" pageNumber="566">Iran</collectingCountry>
.
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<collectingRegion id="49C11DC8FF93FFFFFE1BFF0EFE0DBC8C" box="[411,511,223,249]" country="Iran" name="Golestan" pageId="2" pageNumber="566">Golestan</collectingRegion>
(
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Farashiani
<emphasis id="B9710F38FF93FFFFFD11FF0EFD34BC8C" box="[657,710,223,249]" italics="true" pageId="2" pageNumber="566">et al</emphasis>
. 2022
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(
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Farashiani
<emphasis id="B9710F38FF93FFFFFAE1FF0EFA67BC8C" box="[1377,1429,223,249]" italics="true" pageId="2" pageNumber="566">et al</emphasis>
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<emphasis id="B9710F38FF93FFFFFEABFED2FDD0BD68" box="[299,546,259,285]" italics="true" pageId="2" pageNumber="566">Rhacopus ferrugineus</emphasis>
(
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<emphasis id="B9710F38FF93FFFFFF17FEF7FE64BD35" bold="true" box="[151,406,294,320]" pageId="2" pageNumber="566">General distribution.</emphasis>
<collectingCountry id="F31293BAFF93FFFFFE1DFEF6FDEABD34" box="[413,536,295,321]" name="Azerbaijan" pageId="2" pageNumber="566">Azerbaijan</collectingCountry>
(
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),
<collectingCountry id="F31293BAFF93FFFFFD58FEF6FCFABD34" box="[728,776,295,321]" name="Iran" pageId="2" pageNumber="566">Iran</collectingCountry>
,?
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(
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),
<collectingCountry id="F31293BAFF93FFFFFB09FEF6FB26BD34" box="[1161,1236,295,321]" name="Russia" pageId="2" pageNumber="566">Russia</collectingCountry>
(South European Territory) (
<bibRefCitation id="EF94AEDBFF93FFFFFEC2FE9AFE34BD10" author="Bibin, A. R." box="[322,454,331,357]" pageId="2" pageNumber="566" pagination="1099 - 1101" refId="ref3566" refString="Bibin, A. R. (2020) To the fauna of the false click beetles (Coleoptera, Eucnemidae) of Daghestan, Caucasus. Zoologicheskij zhurnal, 99 (9), 999 - 1001. [in Russian; English translation: Entomological Review, 100 (8), 1099 - 1101 (2021 (2020 ))] https: // doi. org / 10.1134 / S 0013873820080060" type="journal article" year="2020">Bibin 2020</bibRefCitation>
).
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View file

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Distribution in
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(
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Bunalski
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(South European Territory) (
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(
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;
<bibRefCitation id="EF94AEDBFF95FFF9FF47FE0AFEAFBD80" author="Zaitzev, Ph. A." box="[199,349,474,501]" pageId="4" pageNumber="568" pagination="239 - 245" refId="ref7587" refString="Zaitzev, Ph. A. (1951) Semejstvo Phylloceridae v faune Zakavkaz'ya (Coleoptera) [Family Phylloceridae in the fauna of Transcaucasia (Coleoptera)]. Trudy Zoologicheskogo instituta Akademii nauk Gruzinskoj SSR, 10, 239 - 245. [in Russian]" type="journal article" year="1951">Zaitzev 1951</bibRefCitation>
),
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(
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),
<collectingCountry id="F31293BAFF95FFF9FDF3FE0AFD3FBD80" box="[627,717,475,501]" name="Georgia" pageId="4" pageNumber="568">Georgia</collectingCountry>
(
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),
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Bunalski
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<figureCitation id="1320578EF368FFB9FEEB0382AC615FAE" box="[285,356,1415,1441]" captionStart="FIGURES 4446" captionStartId="8.[151,264,683,707]" captionTargetBox="[175,1414,240,656]" captionTargetId="figure-62@8.[170,1418,237,660]" captionTargetPageId="8" captionText="FIGURES 4446. Paramritodus triangulus. 44, Genital capsule, lateral view; 45. Genital capsule, dorsal view; 46, Aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13745541" httpUri="https://zenodo.org/record/13745541/files/figure.png" pageId="4" pageNumber="587">4446</figureCitation>
)
</paragraph>
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<paragraph id="8BA44B0BF368FFB9FF6103C1AE275FD2" blockId="4.[151,802,1476,1501]" box="[151,802,1476,1501]" pageId="4" pageNumber="587">
<treatmentCitationGroup id="AB0B6C25F368FFB9FF6103C1AE275FD2" box="[151,802,1476,1501]" pageId="4" pageNumber="587">
<treatmentCitation id="0ABA6D1AF368FFB9FF6103C1AFA35FD3" author="Xue, Q. Q. &amp; Zhang, Y. L." box="[151,678,1476,1501]" page="1457" pageId="4" pageNumber="587" year="2020">
<taxonomicName id="4C1B3088F368FFB9FF6103C1AFA35FD3" ID-CoL="8JXJK" authority="Xue &amp; Zhang, 2020: 1457" authorityName="Xue &amp; Zhang, 2020:" authorityPageNumber="1457" authorityYear="1457" box="[151,678,1476,1501]" class="Insecta" family="Cicadellidae" genus="Paramritodus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="587" phylum="Arthropoda" rank="species" species="triangulus">
<emphasis id="B96F9719F368FFB9FF6103C1AC975FD3" box="[151,402,1476,1500]" italics="true" pageId="4" pageNumber="587">Paramritodus triangulus</emphasis>
<bibRefCitation id="EF8A36FAF368FFB9FE6F03C0AFA35FD3" author="Xue, Q. Q. &amp; Zhang, Y. L." box="[409,678,1476,1501]" pageId="4" pageNumber="587" pagination="1438 - 1463" refId="ref4537" refString="Xue, Q. Q. &amp; Zhang, Y. L. (2020) Phylogeny and revision of the Oriental leafhopper genus Amritodus (Hemiptera: Cicadellidae: Idiocerini). Zoological Journal of the Linnean Society, 189 (4), 1438 - 1463. https: // doi. org / 10.1093 / zoolinnean / zlz 129" type="journal article" year="2020">Xue &amp; Zhang, 2020: 1457</bibRefCitation>
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, figs 10, 19.
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</paragraph>
</subSubSection>
<subSubSection id="C3011880F368FFB9FF610001A9AE5CF9" pageId="4" pageNumber="587" type="diagnosis">
<paragraph id="8BA44B0BF368FFB9FF610001AF7E5C69" blockId="4.[151,1437,1540,1963]" pageId="4" pageNumber="587">
<emphasis id="B96F9719F368FFB9FF610001AC175C11" bold="true" box="[151,274,1540,1566]" pageId="4" pageNumber="587">Diagnosis.</emphasis>
Crown (
<figureCitation id="1320578EF368FFB9FE8E0001ACC65C10" box="[376,451,1540,1567]" captionStart="FIGURES 16" captionStartId="2.[151,264,1309,1333]" captionTargetBox="[350,1238,164,1299]" captionTargetId="figure-37@2.[350,1238,164,1299]" captionTargetPageId="2" captionText="FIGURES 16. Paramritodus dorsal habitus, male. 1, P. introflexus; 2, P. pistacious; 3, P. podocarpus; 4, P. spatiosus; 5, P. triangulus; 6, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745523" httpUri="https://zenodo.org/record/13745523/files/figure.png" pageId="4" pageNumber="587">Figs 5</figureCitation>
,
<figureCitation id="1320578EF368FFB9FE270001ACE95C11" box="[465,492,1540,1566]" captionStart="FIGURES 712" captionStartId="3.[151,264,1022,1046]" captionTargetBox="[348,1240,309,999]" captionTargetId="figure-65@3.[348,1240,309,999]" captionTargetPageId="3" captionText="FIGURES 712. Facial aspect of Paramritodus, male. 7, P. introflexus; 8, P. pistacious; 9, P. podocarpus; 10, P. spatiosus; 11, P. triangulus; 12, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745525" httpUri="https://zenodo.org/record/13745525/files/figure.png" pageId="4" pageNumber="587">11</figureCitation>
) and face (
<figureCitation id="1320578EF368FFB9FD860001AFC75C11" box="[624,706,1540,1566]" captionStart="FIGURES 712" captionStartId="3.[151,264,1022,1046]" captionTargetBox="[348,1240,309,999]" captionTargetId="figure-65@3.[348,1240,309,999]" captionTargetPageId="3" captionText="FIGURES 712. Facial aspect of Paramritodus, male. 7, P. introflexus; 8, P. pistacious; 9, P. podocarpus; 10, P. spatiosus; 11, P. triangulus; 12, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745525" httpUri="https://zenodo.org/record/13745525/files/figure.png" pageId="4" pageNumber="587">Fig. 11</figureCitation>
) pale brown. Frontclyeal and clypellus yellow brownish, genae and lora (
<figureCitation id="1320578EF368FFB9FEF7002DAC555C4D" box="[257,336,1576,1602]" captionStart="FIGURES 712" captionStartId="3.[151,264,1022,1046]" captionTargetBox="[348,1240,309,999]" captionTargetId="figure-65@3.[348,1240,309,999]" captionTargetPageId="3" captionText="FIGURES 712. Facial aspect of Paramritodus, male. 7, P. introflexus; 8, P. pistacious; 9, P. podocarpus; 10, P. spatiosus; 11, P. triangulus; 12, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745525" httpUri="https://zenodo.org/record/13745525/files/figure.png" pageId="4" pageNumber="587">Fig. 11</figureCitation>
) pale brown. Pronotum (
<figureCitation id="1320578EF368FFB9FD9E002DAFAE5C4C" box="[616,683,1576,1603]" captionStart="FIGURES 16" captionStartId="2.[151,264,1309,1333]" captionTargetBox="[350,1238,164,1299]" captionTargetId="figure-37@2.[350,1238,164,1299]" captionTargetPageId="2" captionText="FIGURES 16. Paramritodus dorsal habitus, male. 1, P. introflexus; 2, P. pistacious; 3, P. podocarpus; 4, P. spatiosus; 5, P. triangulus; 6, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745523" httpUri="https://zenodo.org/record/13745523/files/figure.png" pageId="4" pageNumber="587">Fig. 5</figureCitation>
), mesonotum and scutellum pale brown. Forewings (
<figureCitation id="1320578EF368FFB9FB08002DA8455C4C" box="[1278,1344,1576,1603]" captionStart="FIGURES 16" captionStartId="2.[151,264,1309,1333]" captionTargetBox="[350,1238,164,1299]" captionTargetId="figure-37@2.[350,1238,164,1299]" captionTargetPageId="2" captionText="FIGURES 16. Paramritodus dorsal habitus, male. 1, P. introflexus; 2, P. pistacious; 3, P. podocarpus; 4, P. spatiosus; 5, P. triangulus; 6, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745523" httpUri="https://zenodo.org/record/13745523/files/figure.png" pageId="4" pageNumber="587">Fig. 5</figureCitation>
) yellow brown except the base of claval area white.
</paragraph>
<paragraph id="8BA44B0BF368FFB9FF310075A9AE5CF9" blockId="4.[151,1437,1540,1963]" pageId="4" pageNumber="587">
Pygofer (
<figureCitation id="1320578EF368FFB9FEC70075AC815C85" box="[305,388,1648,1674]" captionStart="FIGURES 4446" captionStartId="8.[151,264,683,707]" captionTargetBox="[175,1414,240,656]" captionTargetId="figure-62@8.[170,1418,237,660]" captionTargetPageId="8" captionText="FIGURES 4446. Paramritodus triangulus. 44, Genital capsule, lateral view; 45. Genital capsule, dorsal view; 46, Aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13745541" httpUri="https://zenodo.org/record/13745541/files/figure.png" pageId="4" pageNumber="587">Fig. 44</figureCitation>
) obviously triangular, apex of pygofer hook-like; pygofer inner process not reaching caudal margin; anal tube surpassing pygofer lobe dorsal margin and enlarged terminally. Length of subgenital plates (
<figureCitation id="1320578EF368FFB9FA990091ADB15CDD" captionStart="FIGURES 4446" captionStartId="8.[151,264,683,707]" captionTargetBox="[175,1414,240,656]" captionTargetId="figure-62@8.[170,1418,237,660]" captionTargetPageId="8" captionText="FIGURES 4446. Paramritodus triangulus. 44, Genital capsule, lateral view; 45. Genital capsule, dorsal view; 46, Aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13745541" httpUri="https://zenodo.org/record/13745541/files/figure.png" pageId="4" pageNumber="587">Fig. 44</figureCitation>
) approximate to pygofer. Aedeagal shaft (
<figureCitation id="1320578EF368FFB9FD6100BDAFEE5CDD" box="[663,747,1720,1746]" captionStart="FIGURES 4446" captionStartId="8.[151,264,683,707]" captionTargetBox="[175,1414,240,656]" captionTargetId="figure-62@8.[170,1418,237,660]" captionTargetPageId="8" captionText="FIGURES 4446. Paramritodus triangulus. 44, Genital capsule, lateral view; 45. Genital capsule, dorsal view; 46, Aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13745541" httpUri="https://zenodo.org/record/13745541/files/figure.png" pageId="4" pageNumber="587">Fig. 46</figureCitation>
) laterally slightly curved dorsally and tapered to apex, shaft with basal pair of slender processes on ventral surface; gonopore on ventral surface medially.
</paragraph>
</subSubSection>
<subSubSection id="C3011880F368FFB9FF310105ADF95D31" pageId="4" pageNumber="587" type="materials_examined">
<paragraph id="8BA44B0BF368FFB9FF310105ADF95D31" blockId="4.[151,1437,1540,1963]" pageId="4" pageNumber="587">
<emphasis id="B96F9719F368FFB9FF310105ACB35D15" bold="true" box="[199,438,1792,1818]" pageId="4" pageNumber="587">Material examined.</emphasis>
<materialsCitation id="3B734156F368FFB9FE370105ADFD5D31" collectingDate="2009-05-28" collectionCode="IMNU" collectorName="Bin Zhang" country="China" location="Mt. Gaoligongshan" pageId="4" pageNumber="587" specimenCount="23" specimenCount-female="13" specimenCount-male="10" stateProvince="Yunnan">
<collectingCountry id="F30C0B9BF368FFB9FE370105AF0D5D15" box="[449,520,1792,1818]" name="China" pageId="4" pageNumber="587">China</collectingCountry>
:
<specimenCount id="9D1D8082F368FFB9FDEE0105AF505D16" box="[536,597,1792,1818]" count="10" pageId="4" pageNumber="587" type="male">10 ♂</specimenCount>
,
<specimenCount id="9D1D8082F368FFB9FD9E0105AFA05D14" box="[616,677,1792,1819]" count="13" pageId="4" pageNumber="587" type="female">13 ♀</specimenCount>
,
<collectingRegion id="49DF85E9F368FFB9FD410104AE535D14" box="[695,854,1793,1819]" country="China" name="Yunnan" pageId="4" pageNumber="587">Yunnan Prov.</collectingRegion>
,
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,
<date id="FFA56DCBF368FFB9FBA00105A9D65D15" box="[1110,1235,1792,1818]" pageId="4" pageNumber="587" value="2009-05-28">
<collectingDate id="EFE19423F368FFB9FBA00105A9D65D15" box="[1110,1235,1792,1818]" pageId="4" pageNumber="587" value="2009-05-28">28.v. 2009</collectingDate>
</date>
, coll.
<collectorName id="26EE2EDDF368FFB9FAD60105A8985D15" box="[1312,1437,1792,1818]" pageId="4" pageNumber="587">Bin Zhang</collectorName>
(
<collectionCode id="ED0AD3CEF368FFB9FF690121ADF55D31" box="[159,240,1828,1854]" pageId="4" pageNumber="587">IMNU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3011880F368FFB9FF31014DAF245D6D" box="[199,545,1864,1890]" pageId="4" pageNumber="587" type="distribution">
<paragraph id="8BA44B0BF368FFB9FF31014DAF245D6D" blockId="4.[151,1437,1540,1963]" box="[199,545,1864,1890]" pageId="4" pageNumber="587">
<emphasis id="B96F9719F368FFB9FF31014DAC645D6D" bold="true" box="[199,353,1864,1890]" pageId="4" pageNumber="587">Distribution.</emphasis>
<collectingCountry id="F30C0B9BF368FFB9FE91014DACAF5D6D" box="[359,426,1864,1890]" name="China" pageId="4" pageNumber="587">China</collectingCountry>
(
<collectingRegion id="49DF85E9F368FFB9FE4F014DAF105D6D" box="[441,533,1864,1890]" country="China" name="Yunnan" pageId="4" pageNumber="587">Yunnan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C3011880F368FFB9FF310169AEE35DA5" pageId="4" pageNumber="587" type="discussion">
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<emphasis id="B96F9719F368FFB9FF310169AC3E5D89" bold="true" box="[199,315,1900,1926]" pageId="4" pageNumber="587">Remarks.</emphasis>
This species is similar to
<taxonomicName id="4C1B3088F368FFB9FDA00168AFE25D89" authorityName="Xue &amp; Zhang, 2020:" authorityYear="1451" box="[598,743,1900,1926]" class="Insecta" family="Cicadellidae" genus="Paramritodus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="587" phylum="Arthropoda" rank="species" species="introflexus">
<emphasis id="B96F9719F368FFB9FDA00168AFE25D89" box="[598,743,1900,1926]" italics="true" pageId="4" pageNumber="587">P. introflexus</emphasis>
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, but can be distinguished from the latter by the shorter pygofer inner process and the aedeagal shaft longer and hardly concave subapically.
</paragraph>
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</treatment>
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View file

@ -0,0 +1,274 @@
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<mods:title id="16600FA1812ACD2C3505ED13AE096D1B">Review of Chinese species of genus Paramritodus Xue &amp; Zhang (Hemiptera: Cicadellidae) with description of a new species</mods:title>
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<heading id="D0ECFC67F368FFB9FF610410AF625820" bold="true" box="[151,615,533,560]" fontSize="11" level="1" pageId="4" pageNumber="587" reason="1">
<taxonomicName id="4C1B3088F368FFB9FF610410AF625820" ID-CoL="8JXJJ" authority="Xue &amp; Zhang" authorityName="Xue &amp; Zhang, 2020:" authorityYear="1455" box="[151,615,533,560]" class="Insecta" family="Cicadellidae" genus="Paramritodus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="587" phylum="Arthropoda" rank="species" species="spatiosus">
<emphasis id="B96F9719F368FFB9FF610410AF625820" bold="true" box="[151,615,533,560]" pageId="4" pageNumber="587">
<emphasis id="B96F9719F368FFB9FF610410ACB15820" bold="true" box="[151,436,533,560]" italics="true" pageId="4" pageNumber="587">Paramritodus spatiosus</emphasis>
Xue &amp; Zhang
</emphasis>
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</heading>
</paragraph>
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<subSubSection id="C3011880F368FFB9FF61043CAC68585B" box="[151,365,569,596]" pageId="4" pageNumber="587" type="description">
<paragraph id="8BA44B0BF368FFB9FF61043CAC68585B" blockId="4.[151,615,533,596]" box="[151,365,569,596]" pageId="4" pageNumber="587">
(
<figureCitation id="1320578EF368FFB9FF69043CADED585C" box="[159,232,569,595]" captionStart="FIGURES 16" captionStartId="2.[151,264,1309,1333]" captionTargetBox="[350,1238,164,1299]" captionTargetId="figure-37@2.[350,1238,164,1299]" captionTargetPageId="2" captionText="FIGURES 16. Paramritodus dorsal habitus, male. 1, P. introflexus; 2, P. pistacious; 3, P. podocarpus; 4, P. spatiosus; 5, P. triangulus; 6, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745523" httpUri="https://zenodo.org/record/13745523/files/figure.png" pageId="4" pageNumber="587">Figs 4</figureCitation>
,
<figureCitation id="1320578EF368FFB9FF02043CAC17585C" box="[244,274,569,595]" captionStart="FIGURES 712" captionStartId="3.[151,264,1022,1046]" captionTargetBox="[348,1240,309,999]" captionTargetId="figure-65@3.[348,1240,309,999]" captionTargetPageId="3" captionText="FIGURES 712. Facial aspect of Paramritodus, male. 7, P. introflexus; 8, P. pistacious; 9, P. podocarpus; 10, P. spatiosus; 11, P. triangulus; 12, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745525" httpUri="https://zenodo.org/record/13745525/files/figure.png" pageId="4" pageNumber="587">10</figureCitation>
,
<figureCitation id="1320578EF368FFB9FEE8043CAC60585B" box="[286,357,569,596]" captionStart="FIGURES 3543" captionStartId="7.[151,264,1476,1500]" captionTargetBox="[158,1424,189,1444]" captionTargetId="figure-17@7.[151,1436,181,1452]" captionTargetPageId="7" captionText="FIGURES 3543. Paramritodus spatiosus Xue &amp; Zhang. 35, Dorsal and ventral abdominal apodemes, dorsal view; 36, Genital capsule, lateral view; 37, Aedeagus, lateral view; 38, Aedeagus, ventral view; 39, Connective lateral view; 40, Connective ventral view; 41, Style, lateral view; 42, Female genital, ventral view; 43, Female genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13745537" httpUri="https://zenodo.org/record/13745537/files/figure.png" pageId="4" pageNumber="587">3543</figureCitation>
)
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<treatmentCitationGroup id="AB0B6C25F368FFB9FF61047BAE175899" box="[151,786,638,662]" pageId="4" pageNumber="587">
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<emphasis id="B96F9719F368FFB9FF61047BAC8D589A" box="[151,392,638,662]" italics="true" pageId="4" pageNumber="587">Paramritodus spatiosus</emphasis>
<bibRefCitation id="EF8A36FAF368FFB9FE79047BAF995899" author="Xue, Q. Q. &amp; Zhang, Y. L." box="[399,668,638,662]" pageId="4" pageNumber="587" pagination="1438 - 1463" refId="ref4537" refString="Xue, Q. Q. &amp; Zhang, Y. L. (2020) Phylogeny and revision of the Oriental leafhopper genus Amritodus (Hemiptera: Cicadellidae: Idiocerini). Zoological Journal of the Linnean Society, 189 (4), 1438 - 1463. https: // doi. org / 10.1093 / zoolinnean / zlz 129" type="journal article" year="2020">Xue &amp; Zhang, 2020: 1455</bibRefCitation>
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, figs. 9, 18.
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</paragraph>
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<subSubSection id="C3011880F368FFB9FF6104C1AF6959B9" pageId="4" pageNumber="587" type="diagnosis">
<paragraph id="8BA44B0BF368FFB9FF6104C1AFB55928" blockId="4.[151,1437,708,1310]" pageId="4" pageNumber="587">
<emphasis id="B96F9719F368FFB9FF6104C1AC1758D1" bold="true" box="[151,274,708,734]" pageId="4" pageNumber="587">Diagnosis.</emphasis>
Crown (
<figureCitation id="1320578EF368FFB9FE8404C1ACBE58D1" box="[370,443,708,734]" captionStart="FIGURES 16" captionStartId="2.[151,264,1309,1333]" captionTargetBox="[350,1238,164,1299]" captionTargetId="figure-37@2.[350,1238,164,1299]" captionTargetPageId="2" captionText="FIGURES 16. Paramritodus dorsal habitus, male. 1, P. introflexus; 2, P. pistacious; 3, P. podocarpus; 4, P. spatiosus; 5, P. triangulus; 6, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745523" httpUri="https://zenodo.org/record/13745523/files/figure.png" pageId="4" pageNumber="587">Figs 4</figureCitation>
,
<figureCitation id="1320578EF368FFB9FE3004C1ACE658D1" box="[454,483,708,734]" captionStart="FIGURES 712" captionStartId="3.[151,264,1022,1046]" captionTargetBox="[348,1240,309,999]" captionTargetId="figure-65@3.[348,1240,309,999]" captionTargetPageId="3" captionText="FIGURES 712. Facial aspect of Paramritodus, male. 7, P. introflexus; 8, P. pistacious; 9, P. podocarpus; 10, P. spatiosus; 11, P. triangulus; 12, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745525" httpUri="https://zenodo.org/record/13745525/files/figure.png" pageId="4" pageNumber="587">10</figureCitation>
) with pair of black spots. Pronotum (
<figureCitation id="1320578EF368FFB9FC7704C1AEC158D1" box="[897,964,708,734]" captionStart="FIGURES 16" captionStartId="2.[151,264,1309,1333]" captionTargetBox="[350,1238,164,1299]" captionTargetId="figure-37@2.[350,1238,164,1299]" captionTargetPageId="2" captionText="FIGURES 16. Paramritodus dorsal habitus, male. 1, P. introflexus; 2, P. pistacious; 3, P. podocarpus; 4, P. spatiosus; 5, P. triangulus; 6, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745523" httpUri="https://zenodo.org/record/13745523/files/figure.png" pageId="4" pageNumber="587">Fig. 4</figureCitation>
) with pair of triangular black markings on anterior margin, mesonotum (
<figureCitation id="1320578EF368FFB9FE1504EDAF23590D" box="[483,550,744,770]" captionStart="FIGURES 16" captionStartId="2.[151,264,1309,1333]" captionTargetBox="[350,1238,164,1299]" captionTargetId="figure-37@2.[350,1238,164,1299]" captionTargetPageId="2" captionText="FIGURES 16. Paramritodus dorsal habitus, male. 1, P. introflexus; 2, P. pistacious; 3, P. podocarpus; 4, P. spatiosus; 5, P. triangulus; 6, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745523" httpUri="https://zenodo.org/record/13745523/files/figure.png" pageId="4" pageNumber="587">Fig. 4</figureCitation>
) yellowish with basal triangles. Gena (
<figureCitation id="1320578EF368FFB9FC2C04EDA92E590D" box="[986,1067,744,770]" captionStart="FIGURES 712" captionStartId="3.[151,264,1022,1046]" captionTargetBox="[348,1240,309,999]" captionTargetId="figure-65@3.[348,1240,309,999]" captionTargetPageId="3" captionText="FIGURES 712. Facial aspect of Paramritodus, male. 7, P. introflexus; 8, P. pistacious; 9, P. podocarpus; 10, P. spatiosus; 11, P. triangulus; 12, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745525" httpUri="https://zenodo.org/record/13745525/files/figure.png" pageId="4" pageNumber="587">Fig. 10</figureCitation>
) pale yellowish. Forewings (
<figureCitation id="1320578EF368FFB9FA9904EDADA35929" captionStart="FIGURES 16" captionStartId="2.[151,264,1309,1333]" captionTargetBox="[350,1238,164,1299]" captionTargetId="figure-37@2.[350,1238,164,1299]" captionTargetPageId="2" captionText="FIGURES 16. Paramritodus dorsal habitus, male. 1, P. introflexus; 2, P. pistacious; 3, P. podocarpus; 4, P. spatiosus; 5, P. triangulus; 6, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745523" httpUri="https://zenodo.org/record/13745523/files/figure.png" pageId="4" pageNumber="587">Fig. 4</figureCitation>
) yellowish brown except clavus yellow green.
</paragraph>
<paragraph id="8BA44B0BF368FFB9FF310535AF6959B9" blockId="4.[151,1437,708,1310]" pageId="4" pageNumber="587">
Pygofer (
<figureCitation id="1320578EF368FFB9FEC40535AC835945" box="[306,390,816,842]" captionStart="FIGURES 3543" captionStartId="7.[151,264,1476,1500]" captionTargetBox="[158,1424,189,1444]" captionTargetId="figure-17@7.[151,1436,181,1452]" captionTargetPageId="7" captionText="FIGURES 3543. Paramritodus spatiosus Xue &amp; Zhang. 35, Dorsal and ventral abdominal apodemes, dorsal view; 36, Genital capsule, lateral view; 37, Aedeagus, lateral view; 38, Aedeagus, ventral view; 39, Connective lateral view; 40, Connective ventral view; 41, Style, lateral view; 42, Female genital, ventral view; 43, Female genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13745537" httpUri="https://zenodo.org/record/13745537/files/figure.png" pageId="4" pageNumber="587">Fig. 36</figureCitation>
) inner process concave dorsally and not reaching caudal margin in lateral view. Aadeagus (
<figureCitation id="1320578EF368FFB9FF690551AC1A5961" box="[159,287,852,878]" captionStart="FIGURES 3543" captionStartId="7.[151,264,1476,1500]" captionTargetBox="[158,1424,189,1444]" captionTargetId="figure-17@7.[151,1436,181,1452]" captionTargetPageId="7" captionText="FIGURES 3543. Paramritodus spatiosus Xue &amp; Zhang. 35, Dorsal and ventral abdominal apodemes, dorsal view; 36, Genital capsule, lateral view; 37, Aedeagus, lateral view; 38, Aedeagus, ventral view; 39, Connective lateral view; 40, Connective ventral view; 41, Style, lateral view; 42, Female genital, ventral view; 43, Female genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13745537" httpUri="https://zenodo.org/record/13745537/files/figure.png" pageId="4" pageNumber="587">Figs 3738</figureCitation>
) with preatrium broadened, process longer than half length of shaft, dorsal apodeme poorly developed; aedeagal shaft laterally obviously curved dorsally, and with elongate process on each side above preatrium. Gonopore subapical. Style (
<figureCitation id="1320578EF368FFB9FEAE0599ACAC59B9" box="[344,425,924,950]" captionStart="FIGURES 3543" captionStartId="7.[151,264,1476,1500]" captionTargetBox="[158,1424,189,1444]" captionTargetId="figure-17@7.[151,1436,181,1452]" captionTargetPageId="7" captionText="FIGURES 3543. Paramritodus spatiosus Xue &amp; Zhang. 35, Dorsal and ventral abdominal apodemes, dorsal view; 36, Genital capsule, lateral view; 37, Aedeagus, lateral view; 38, Aedeagus, ventral view; 39, Connective lateral view; 40, Connective ventral view; 41, Style, lateral view; 42, Female genital, ventral view; 43, Female genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13745537" httpUri="https://zenodo.org/record/13745537/files/figure.png" pageId="4" pageNumber="587">Fig. 41</figureCitation>
) curved dorsally.
</paragraph>
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<emphasis id="B96F9719F368FFB9FF3105C5ACB559D5" bold="true" box="[199,432,960,986]" pageId="4" pageNumber="587">Material examined.</emphasis>
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,
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<elevation id="0036AC38F368FFB9FF070255AC4D5E64" box="[241,328,1104,1131]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="4" pageNumber="587" unit="m" value="1700.0">1700 m</elevation>
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,
<date id="FFA56DCBF368FFB9FEA20255ACDB5E64" box="[340,478,1104,1131]" pageId="4" pageNumber="587" value="2009-08-15">
<collectingDate id="EFE19423F368FFB9FEA20255ACDB5E64" box="[340,478,1104,1131]" pageId="4" pageNumber="587" value="2009-08-15">15.viii.2009</collectingDate>
</date>
, coll.
<collectorName id="26EE2EDDF368FFB9FDD40255AF9C5E65" box="[546,665,1104,1130]" pageId="4" pageNumber="587">Bin Zhang</collectorName>
(
<collectionCode id="ED0AD3CEF368FFB9FD510255AFFD5E65" box="[679,760,1104,1130]" pageId="4" pageNumber="587">IMNU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3011880F368FFB9FF310271AF245E81" box="[199,545,1140,1166]" pageId="4" pageNumber="587" type="distribution">
<paragraph id="8BA44B0BF368FFB9FF310271AF245E81" blockId="4.[151,1437,708,1310]" box="[199,545,1140,1166]" pageId="4" pageNumber="587">
<emphasis id="B96F9719F368FFB9FF310271AC645E81" bold="true" box="[199,353,1140,1166]" pageId="4" pageNumber="587">Distribution.</emphasis>
<collectingCountry id="F30C0B9BF368FFB9FE910271ACAF5E81" box="[359,426,1140,1166]" name="China" pageId="4" pageNumber="587">China</collectingCountry>
(
<collectingRegion id="49DF85E9F368FFB9FE4F0271AF105E81" box="[441,533,1140,1166]" country="China" name="Yunnan" pageId="4" pageNumber="587">Yunnan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C3011880F368FFB9FF31029DAC0A5F11" pageId="4" pageNumber="587" type="discussion">
<paragraph id="8BA44B0BF368FFB9FF31029DAC0A5F11" blockId="4.[151,1437,708,1310]" pageId="4" pageNumber="587">
<emphasis id="B96F9719F368FFB9FF31029DAC3E5EBD" bold="true" box="[199,315,1176,1202]" pageId="4" pageNumber="587">Remarks.</emphasis>
This species has a pair of processes on the aedeagal preatrium as in
<taxonomicName id="4C1B3088F368FFB9FBA8029CA9F15EBD" authorityName="Xue &amp; Zhang, 2020:" authorityYear="1451" box="[1118,1268,1176,1202]" class="Insecta" family="Cicadellidae" genus="Paramritodus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="587" phylum="Arthropoda" rank="species" species="introflexus">
<emphasis id="B96F9719F368FFB9FBA8029CA9F15EBD" box="[1118,1268,1176,1202]" italics="true" pageId="4" pageNumber="587">P. introflexus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1B3088F368FFB9FAF2029CA8905EBD" authorityName="Xue &amp; Zhang, 2020:" authorityYear="1457" box="[1284,1429,1176,1202]" class="Insecta" family="Cicadellidae" genus="Paramritodus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="587" phylum="Arthropoda" rank="species" species="triangulus">
<emphasis id="B96F9719F368FFB9FAF2029CA8905EBD" box="[1284,1429,1176,1202]" italics="true" pageId="4" pageNumber="587">P. triangulus</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C1B3088F368FFB9FF3E02B8AC8E5ED9" box="[200,395,1212,1238]" class="Insecta" family="Cicadellidae" genus="Paramritodus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="587" phylum="Arthropoda" rank="species" species="concavicaudus">
<emphasis id="B96F9719F368FFB9FF3E02B8AC8E5ED9" box="[200,395,1212,1238]" italics="true" pageId="4" pageNumber="587">P. concavicaudus</emphasis>
</taxonomicName>
. It differs from
<taxonomicName id="4C1B3088F368FFB9FDB202B8AFDD5ED9" authorityName="Xue &amp; Zhang, 2020:" authorityYear="1451" box="[580,728,1212,1238]" class="Insecta" family="Cicadellidae" genus="Paramritodus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="587" phylum="Arthropoda" rank="species" species="introflexus">
<emphasis id="B96F9719F368FFB9FDB202B8AFDD5ED9" box="[580,728,1212,1238]" italics="true" pageId="4" pageNumber="587">P. introflexus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1B3088F368FFB9FCE602B8AEA55ED9" authorityName="Xue &amp; Zhang, 2020:" authorityYear="1457" box="[784,928,1212,1238]" class="Insecta" family="Cicadellidae" genus="Paramritodus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="587" phylum="Arthropoda" rank="species" species="triangulus">
<emphasis id="B96F9719F368FFB9FCE602B8AEA55ED9" box="[784,928,1212,1238]" italics="true" pageId="4" pageNumber="587">P. triangulus</emphasis>
</taxonomicName>
in having the aedeagal processes longer and the gonopore near the subapex, and from
<taxonomicName id="4C1B3088F368FFB9FD8E02E4AE3E5EF5" box="[632,827,1248,1274]" class="Insecta" family="Cicadellidae" genus="Paramritodus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="587" phylum="Arthropoda" rank="species" species="concavicaudus">
<emphasis id="B96F9719F368FFB9FD8E02E4AE3E5EF5" box="[632,827,1248,1274]" italics="true" pageId="4" pageNumber="587">P. concavicaudus</emphasis>
</taxonomicName>
in having the style lacking a bifucate process on the apophysis.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,232 @@
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<mods:title id="6EB390CB95A5BAA0E0DBF6719CFE5DF7">Review of Chinese species of genus Paramritodus Xue &amp; Zhang (Hemiptera: Cicadellidae) with description of a new species</mods:title>
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<subSubSection id="C3011880F36BFFBAFF610073AFB35C9E" box="[151,694,1654,1681]" pageId="7" pageNumber="590" type="nomenclature">
<paragraph id="8BA44B0BF36BFFBAFF610073AFB35C9E" blockId="7.[151,694,1654,1717]" box="[151,694,1654,1681]" pageId="7" pageNumber="590">
<heading id="D0ECFC67F36BFFBAFF610073AFB35C9E" bold="true" box="[151,694,1654,1681]" fontSize="11" level="1" pageId="7" pageNumber="590" reason="1">
<emphasis id="B96F9719F36BFFBAFF610073AFB35C9E" bold="true" box="[151,694,1654,1681]" pageId="7" pageNumber="590">
<taxonomicName id="4C1B3088F36BFFBAFF610073AF545C9E" authority="Zhang" authorityName="Zhang" authorityYear="2024" box="[151,593,1654,1681]" class="Insecta" family="Cicadellidae" genus="Paramritodus" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="590" phylum="Arthropoda" rank="species" species="concavicaudus" status="sp. nov.">
<emphasis id="B96F9719F36BFFBAFF610073ACFD5C9E" bold="true" box="[151,504,1654,1681]" italics="true" pageId="7" pageNumber="590">Paramritodus concavicaudus</emphasis>
Zhang
</taxonomicName>
<taxonomicNameLabel id="A25C2A62F36BFFBAFDAE0072AFB35C9E" box="[600,694,1655,1681]" pageId="7" pageNumber="590" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C3011880F36BFFBAFF61009FAC685CBA" box="[151,365,1690,1717]" pageId="7" pageNumber="590" type="description">
<paragraph id="8BA44B0BF36BFFBAFF61009FAC685CBA" blockId="7.[151,694,1654,1717]" box="[151,365,1690,1717]" pageId="7" pageNumber="590">
(
<figureCitation id="1320578EF36BFFBAFF69009FADED5CBB" box="[159,232,1690,1716]" captionStart="FIGURES 16" captionStartId="2.[151,264,1309,1333]" captionTargetBox="[350,1238,164,1299]" captionTargetId="figure-37@2.[350,1238,164,1299]" captionTargetPageId="2" captionText="FIGURES 16. Paramritodus dorsal habitus, male. 1, P. introflexus; 2, P. pistacious; 3, P. podocarpus; 4, P. spatiosus; 5, P. triangulus; 6, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745523" httpUri="https://zenodo.org/record/13745523/files/figure.png" pageId="7" pageNumber="590">Figs 6</figureCitation>
,
<figureCitation id="1320578EF36BFFBAFF02009EAC175CBA" box="[244,274,1691,1717]" captionStart="FIGURES 712" captionStartId="3.[151,264,1022,1046]" captionTargetBox="[348,1240,309,999]" captionTargetId="figure-65@3.[348,1240,309,999]" captionTargetPageId="3" captionText="FIGURES 712. Facial aspect of Paramritodus, male. 7, P. introflexus; 8, P. pistacious; 9, P. podocarpus; 10, P. spatiosus; 11, P. triangulus; 12, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745525" httpUri="https://zenodo.org/record/13745525/files/figure.png" pageId="7" pageNumber="590">12</figureCitation>
,
<figureCitation id="1320578EF36BFFBAFEE8009FAC605CBA" box="[286,357,1690,1717]" captionStart="FIGURES 4757" captionStartId="8.[151,264,1898,1922]" captionTargetBox="[176,1413,776,1871]" captionTargetId="figure-84@8.[170,1417,767,1874]" captionTargetPageId="8" captionText="FIGURES 4757. Paramritodus concavicaudus Zhang sp. nov. 47, Dorsal and ventral abdominal apodemes, dorsal view; 48, Genital capsule, lateral view; 49, Genital capsule, dorsal view; 50, Aedeagus, lateral view; 51, Aedeagus, ventral view; 52, Connective ventral view; 53, Connective lateral view; 54, Style, lateral view; 55, Female genital, ventral view; 56, Female seventh sternite, dorsal view; 57, Female genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13745543" httpUri="https://zenodo.org/record/13745543/files/figure.png" pageId="7" pageNumber="590">4757</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3011880F36BFFB4FF6100E7AEF95B12" lastPageId="9" lastPageNumber="592" pageId="7" pageNumber="590" type="diagnosis">
<paragraph id="8BA44B0BF36BFFBAFF6100E7A9E45D4B" blockId="7.[151,1436,1762,2041]" pageId="7" pageNumber="590">
<emphasis id="B96F9719F36BFFBAFF6100E7AC175CF3" bold="true" box="[151,274,1762,1788]" pageId="7" pageNumber="590">Diagnosis.</emphasis>
<emphasis id="B96F9719F36BFFBAFEE800E7AC5A5CF3" box="[286,351,1762,1788]" italics="true" pageId="7" pageNumber="590">Male.</emphasis>
Body yellowish brown (
<figureCitation id="1320578EF36BFFBAFD7D00E7AFD05CF3" box="[651,725,1762,1788]" captionStart="FIGURES 16" captionStartId="2.[151,264,1309,1333]" captionTargetBox="[350,1238,164,1299]" captionTargetId="figure-37@2.[350,1238,164,1299]" captionTargetPageId="2" captionText="FIGURES 16. Paramritodus dorsal habitus, male. 1, P. introflexus; 2, P. pistacious; 3, P. podocarpus; 4, P. spatiosus; 5, P. triangulus; 6, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745523" httpUri="https://zenodo.org/record/13745523/files/figure.png" pageId="7" pageNumber="590">Fig. 6</figureCitation>
). Upper part of face, genae and lora (
<figureCitation id="1320578EF36BFFBAFB5000E7A9F85CF2" box="[1190,1277,1762,1789]" captionStart="FIGURES 712" captionStartId="3.[151,264,1022,1046]" captionTargetBox="[348,1240,309,999]" captionTargetId="figure-65@3.[348,1240,309,999]" captionTargetPageId="3" captionText="FIGURES 712. Facial aspect of Paramritodus, male. 7, P. introflexus; 8, P. pistacious; 9, P. podocarpus; 10, P. spatiosus; 11, P. triangulus; 12, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745525" httpUri="https://zenodo.org/record/13745525/files/figure.png" pageId="7" pageNumber="590">Fig. 12</figureCitation>
) pale brown, frontoclypeus and anteclypeus yellowish brown. Pronotum (
<figureCitation id="1320578EF36BFFBAFCB40103AE825D2F" box="[834,903,1798,1824]" captionStart="FIGURES 16" captionStartId="2.[151,264,1309,1333]" captionTargetBox="[350,1238,164,1299]" captionTargetId="figure-37@2.[350,1238,164,1299]" captionTargetPageId="2" captionText="FIGURES 16. Paramritodus dorsal habitus, male. 1, P. introflexus; 2, P. pistacious; 3, P. podocarpus; 4, P. spatiosus; 5, P. triangulus; 6, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745523" httpUri="https://zenodo.org/record/13745523/files/figure.png" pageId="7" pageNumber="590">Fig. 6</figureCitation>
) with triangular mark, small, black on anterior margin, mesonotum with basal triangles mark black. Forewings (
<figureCitation id="1320578EF36BFFBAFC99012FAEB65D4B" box="[879,947,1834,1860]" captionStart="FIGURES 16" captionStartId="2.[151,264,1309,1333]" captionTargetBox="[350,1238,164,1299]" captionTargetId="figure-37@2.[350,1238,164,1299]" captionTargetPageId="2" captionText="FIGURES 16. Paramritodus dorsal habitus, male. 1, P. introflexus; 2, P. pistacious; 3, P. podocarpus; 4, P. spatiosus; 5, P. triangulus; 6, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745523" httpUri="https://zenodo.org/record/13745523/files/figure.png" pageId="7" pageNumber="590">Fig. 6</figureCitation>
) brown, veins dark brown.
</paragraph>
<paragraph id="8BA44B0BF36BFFB5FF31014BAF475ADA" blockId="7.[151,1436,1762,2041]" lastBlockId="8.[151,1436,150,213]" lastPageId="8" lastPageNumber="591" pageId="7" pageNumber="590">
<emphasis id="B96F9719F36BFFBAFF31014BAC685D67" box="[199,365,1870,1896]" italics="true" pageId="7" pageNumber="590">Male genitalia</emphasis>
. Pygofer (
<figureCitation id="1320578EF36BFFBAFE13014BAF625D66" box="[485,615,1870,1897]" captionStart="FIGURES 4757" captionStartId="8.[151,264,1898,1922]" captionTargetBox="[176,1413,776,1871]" captionTargetId="figure-84@8.[170,1417,767,1874]" captionTargetPageId="8" captionText="FIGURES 4757. Paramritodus concavicaudus Zhang sp. nov. 47, Dorsal and ventral abdominal apodemes, dorsal view; 48, Genital capsule, lateral view; 49, Genital capsule, dorsal view; 50, Aedeagus, lateral view; 51, Aedeagus, ventral view; 52, Connective ventral view; 53, Connective lateral view; 54, Style, lateral view; 55, Female genital, ventral view; 56, Female seventh sternite, dorsal view; 57, Female genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13745543" httpUri="https://zenodo.org/record/13745543/files/figure.png" pageId="7" pageNumber="590">Figs 4849</figureCitation>
) with ventral margin strongly concave dorsally; inner process S-shaped, not reaching caudal margin; anal tube surpassing pygofer lobe ventral margin and hook-like terminally. Subgenital plates distinctly shorter than pygofer.Aadeagus (
<figureCitation id="1320578EF36BFFBAFD5A0193AE2F5DBE" box="[684,810,1942,1969]" captionStart="FIGURES 4757" captionStartId="8.[151,264,1898,1922]" captionTargetBox="[176,1413,776,1871]" captionTargetId="figure-84@8.[170,1417,767,1874]" captionTargetPageId="8" captionText="FIGURES 4757. Paramritodus concavicaudus Zhang sp. nov. 47, Dorsal and ventral abdominal apodemes, dorsal view; 48, Genital capsule, lateral view; 49, Genital capsule, dorsal view; 50, Aedeagus, lateral view; 51, Aedeagus, ventral view; 52, Connective ventral view; 53, Connective lateral view; 54, Style, lateral view; 55, Female genital, ventral view; 56, Female seventh sternite, dorsal view; 57, Female genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13745543" httpUri="https://zenodo.org/record/13745543/files/figure.png" pageId="7" pageNumber="590">Figs 5051</figureCitation>
) with developed dorsal apodeme and elongate preatrium; aedeagal shaft slightly curved dorsally and tapered to apex, with elongate process on each side above preatrium, gonopore subapical on ventral surface. Connective (
<figureCitation id="1320578EF36BFFBAFD0601DBAE765DF6" box="[752,883,2014,2041]" captionStart="FIGURES 4757" captionStartId="8.[151,264,1898,1922]" captionTargetBox="[176,1413,776,1871]" captionTargetId="figure-84@8.[170,1417,767,1874]" captionTargetPageId="8" captionText="FIGURES 4757. Paramritodus concavicaudus Zhang sp. nov. 47, Dorsal and ventral abdominal apodemes, dorsal view; 48, Genital capsule, lateral view; 49, Genital capsule, dorsal view; 50, Aedeagus, lateral view; 51, Aedeagus, ventral view; 52, Connective ventral view; 53, Connective lateral view; 54, Style, lateral view; 55, Female genital, ventral view; 56, Female seventh sternite, dorsal view; 57, Female genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13745543" httpUri="https://zenodo.org/record/13745543/files/figure.png" pageId="7" pageNumber="590">Figs 5253</figureCitation>
) as in generic diagnosis. Style (
<figureCitation id="1320578EF36BFFBAFB1F01DBA83B5DF6" box="[1257,1342,2014,2041]" captionStart="FIGURES 4757" captionStartId="8.[151,264,1898,1922]" captionTargetBox="[176,1413,776,1871]" captionTargetId="figure-84@8.[170,1417,767,1874]" captionTargetPageId="8" captionText="FIGURES 4757. Paramritodus concavicaudus Zhang sp. nov. 47, Dorsal and ventral abdominal apodemes, dorsal view; 48, Genital capsule, lateral view; 49, Genital capsule, dorsal view; 50, Aedeagus, lateral view; 51, Aedeagus, ventral view; 52, Connective ventral view; 53, Connective lateral view; 54, Style, lateral view; 55, Female genital, ventral view; 56, Female seventh sternite, dorsal view; 57, Female genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13745543" httpUri="https://zenodo.org/record/13745543/files/figure.png" pageId="7" pageNumber="590">Fig. 54</figureCitation>
) curved dorsally at nearly 45° angle, apophysis of uniform width to apex fork-like, ventral margin with a small number of setae, dorsal margin with 1416 teeth.
</paragraph>
<caption id="DF641B83F364FFB5FF6104AEAC1158E8" ID-DOI="http://doi.org/10.5281/zenodo.13745541" ID-Zenodo-Dep="13745541" httpUri="https://zenodo.org/record/13745541/files/figure.png" pageId="8" pageNumber="591" startId="8.[151,264,683,707]" targetBox="[175,1414,240,656]" targetPageId="8" targetType="figure">
<paragraph id="8BA44B0BF364FFB5FF6104AEAC1158E8" blockId="8.[151,1436,683,743]" pageId="8" pageNumber="591">
<emphasis id="B96F9719F364FFB5FF6104AEAC5058CC" bold="true" box="[151,341,683,707]" pageId="8" pageNumber="591">FIGURES 4446.</emphasis>
<taxonomicName id="4C1B3088F364FFB5FEAD04AEAF5358CC" authorityName="Xue &amp; Zhang, 2020:" authorityYear="1457" box="[347,598,683,707]" class="Insecta" family="Cicadellidae" genus="Paramritodus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="591" phylum="Arthropoda" rank="species" species="triangulus">
<emphasis id="B96F9719F364FFB5FEAD04AEAF5358CC" box="[347,598,683,707]" italics="true" pageId="8" pageNumber="591">Paramritodus triangulus</emphasis>
</taxonomicName>
. 44, Genital capsule, lateral view; 45. Genital capsule, dorsal view; 46, Aedeagus, lateral view.
</paragraph>
</caption>
<caption id="DF641B83F364FFB5FF61016FAE595DE1" ID-DOI="http://doi.org/10.5281/zenodo.13745543" ID-Zenodo-Dep="13745543" httpUri="https://zenodo.org/record/13745543/files/figure.png" pageId="8" pageNumber="591" startId="8.[151,264,1898,1922]" targetBox="[176,1413,776,1871]" targetPageId="8" targetType="figure">
<paragraph id="8BA44B0BF364FFB5FF61016FAE595DE1" blockId="8.[151,1437,1898,2030]" pageId="8" pageNumber="591">
<emphasis id="B96F9719F364FFB5FF61016FAC525D8D" bold="true" box="[151,343,1898,1922]" pageId="8" pageNumber="591">FIGURES 4757.</emphasis>
<taxonomicName id="4C1B3088F364FFB5FE96016FAFDD5D8D" authority="Zhang" authorityName="Zhang" authorityYear="2024" box="[352,728,1898,1922]" class="Insecta" family="Cicadellidae" genus="Paramritodus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="591" phylum="Arthropoda" rank="species" species="concavicaudus" status="sp. nov.">
<emphasis id="B96F9719F364FFB5FE96016FAF8B5D8D" box="[352,654,1898,1922]" italics="true" pageId="8" pageNumber="591">Paramritodus concavicaudus</emphasis>
Zhang
</taxonomicName>
<emphasis id="B96F9719F364FFB5FD17016FAE305D8D" bold="true" box="[737,821,1898,1922]" pageId="8" pageNumber="591">
<taxonomicNameLabel id="A25C2A62F364FFB5FD17016FAE305D8D" box="[737,821,1898,1922]" pageId="8" pageNumber="591" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
47, Dorsal and ventral abdominal apodemes, dorsal view; 48, Genital capsule, lateral view; 49, Genital capsule, dorsal view; 50, Aedeagus, lateral view; 51, Aedeagus, ventral view; 52, Connective ventral view; 53, Connective lateral view; 54, Style, lateral view; 55, Female genital, ventral view; 56, Female seventh sternite, dorsal view; 57, Female genital capsule, lateral view.
</paragraph>
</caption>
<paragraph id="8BA44B0BF365FFB4FF310692AFE75ADA" blockId="9.[151,1437,151,573]" pageId="9" pageNumber="592">
<emphasis id="B96F9719F365FFB4FF310692AC1E5ABE" box="[199,283,151,177]" italics="true" pageId="9" pageNumber="592">Female</emphasis>
. Similar in overall coloration to male, but face slightly darker. Crown yellowish brown, with pale brown maculae on either side of median line closer to eyes.
</paragraph>
<paragraph id="8BA44B0BF365FFB4FF3106DAAEF95B12" blockId="9.[151,1437,151,573]" pageId="9" pageNumber="592">
<emphasis id="B96F9719F365FFB4FF3106DAAC8E5AF6" box="[199,395,223,249]" italics="true" pageId="9" pageNumber="592">Female genitalia</emphasis>
. Seventh sternite (
<figureCitation id="1320578EF365FFB4FD9A06DAAFC75AF6" box="[620,706,222,249]" captionStart="FIGURES 4757" captionStartId="8.[151,264,1898,1922]" captionTargetBox="[176,1413,776,1871]" captionTargetId="figure-84@8.[170,1417,767,1874]" captionTargetPageId="8" captionText="FIGURES 4757. Paramritodus concavicaudus Zhang sp. nov. 47, Dorsal and ventral abdominal apodemes, dorsal view; 48, Genital capsule, lateral view; 49, Genital capsule, dorsal view; 50, Aedeagus, lateral view; 51, Aedeagus, ventral view; 52, Connective ventral view; 53, Connective lateral view; 54, Style, lateral view; 55, Female genital, ventral view; 56, Female seventh sternite, dorsal view; 57, Female genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13745543" httpUri="https://zenodo.org/record/13745543/files/figure.png" pageId="9" pageNumber="592">Fig. 56</figureCitation>
) concave in dorsal view. Second valvulae (
<figureCitation id="1320578EF365FFB4FB3A06DAA8265AF6" box="[1228,1315,222,249]" captionStart="FIGURES 4757" captionStartId="8.[151,264,1898,1922]" captionTargetBox="[176,1413,776,1871]" captionTargetId="figure-84@8.[170,1417,767,1874]" captionTargetPageId="8" captionText="FIGURES 4757. Paramritodus concavicaudus Zhang sp. nov. 47, Dorsal and ventral abdominal apodemes, dorsal view; 48, Genital capsule, lateral view; 49, Genital capsule, dorsal view; 50, Aedeagus, lateral view; 51, Aedeagus, ventral view; 52, Connective ventral view; 53, Connective lateral view; 54, Style, lateral view; 55, Female genital, ventral view; 56, Female seventh sternite, dorsal view; 57, Female genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13745543" httpUri="https://zenodo.org/record/13745543/files/figure.png" pageId="9" pageNumber="592">Fig. 57</figureCitation>
) regularly broadened in lateral view, bearing approximately 9 teeth along dorsal margin.
</paragraph>
</subSubSection>
<subSubSection id="C3011880F365FFB4FF310722AE465B4E" box="[199,835,294,321]" pageId="9" pageNumber="592" type="description">
<paragraph id="8BA44B0BF365FFB4FF310722AE465B4E" blockId="9.[151,1437,151,573]" box="[199,835,294,321]" pageId="9" pageNumber="592">
<emphasis id="B96F9719F365FFB4FF310722AC7B5B4E" bold="true" box="[199,382,295,321]" pageId="9" pageNumber="592">Measurements.</emphasis>
Male: 4.34.4mm, Female: 4.24.5mm.
</paragraph>
</subSubSection>
<subSubSection id="C3011880F365FFB4FF31074FAFDD5BA2" pageId="9" pageNumber="592" type="materials_examined">
<paragraph id="8BA44B0BF365FFB4FF31074FAFDD5BA2" blockId="9.[151,1437,151,573]" pageId="9" pageNumber="592">
<emphasis id="B96F9719F365FFB4FF31074FACB65B6B" bold="true" box="[199,435,330,356]" pageId="9" pageNumber="592">Material examined.</emphasis>
<materialsCitation id="3B734156F365FFB4FE4A074EAFD15BA2" collectingDate="2021-07-26" collectingDateMax="2022-07-20" collectingDateMin="2021-07-26" collectionCode="IMNU" collectorName="Wei Wang &amp; Qiuyi Zhu" country="China" county="Chongzuo City" location="Mt. Damingshan" municipality="Longzhou" pageId="9" pageNumber="592" specimenCount="9" specimenCount-female="5" specimenCount-male="4" stateProvince="Guangxi" typeStatus="holotype">
Type.
<typeStatus id="54A0F5A9F365FFB4FDF2074EAF745B6A" box="[516,625,331,357]" pageId="9" pageNumber="592" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D1D8082F365FFB4FD88074EAFA95B6A" box="[638,684,331,357]" count="1" pageId="9" pageNumber="592" type="male">1 ♂</specimenCount>
,
<collectingCountry id="F30C0B9BF365FFB4FD4D074EAE125B6A" box="[699,791,331,357]" name="China" pageId="9" pageNumber="592">CHINA</collectingCountry>
,
<collectingRegion id="49DF85E9F365FFB4FCD2074EAECE5B6A" box="[804,971,331,357]" country="China" name="Guangxi" pageId="9" pageNumber="592">Guangxi Prov.</collectingRegion>
,
<collectingCounty id="62C53387F365FFB4FC2E074EA9835B6A" box="[984,1158,331,357]" pageId="9" pageNumber="592">Chongzuo City</collectingCounty>
,
<collectingMunicipality id="6BC0D171F365FFB4FB62074EA80D5B6A" box="[1172,1288,331,357]" pageId="9" pageNumber="592">Longzhou</collectingMunicipality>
,
<date id="FFA56DCBF365FFB4FAE0074EA8925B6A" box="[1302,1431,331,357]" pageId="9" pageNumber="592" value="2021-07-26">
<collectingDate id="EFE19423F365FFB4FAE0074EA8925B6A" box="[1302,1431,331,357]" pageId="9" pageNumber="592" value="2021-07-26">26.vii.2021</collectingDate>
</date>
, coll.
<collectorName id="26EE2EDDF365FFB4FF39076AAC465B86" box="[207,323,367,393]" pageId="9" pageNumber="592">Wei Wang</collectorName>
(
<collectionCode id="ED0AD3CEF365FFB4FEA7076AACA75B86" box="[337,418,367,393]" pageId="9" pageNumber="592">IMNU</collectionCode>
).
<typeStatus id="54A0F5A9F365FFB4FE43076AAF235B86" box="[437,550,367,393]" pageId="9" pageNumber="592" type="paratype">Paratypes</typeStatus>
:
<collectingRegion id="49DF85E9F365FFB4FDC6076AAFD05B86" box="[560,725,367,393]" country="China" name="Guangxi" pageId="9" pageNumber="592">Guangxi Prov.</collectingRegion>
,
<specimenCount id="9D1D8082F365FFB4FD16076AAE095B86" box="[736,780,367,393]" count="2" pageId="9" pageNumber="592" type="male">2 ♂</specimenCount>
,
<specimenCount id="9D1D8082F365FFB4FCEE076AAE475B86" box="[792,834,367,393]" count="4" pageId="9" pageNumber="592" type="female">4 ♀</specimenCount>
, data same as holotype;
<specimenCount id="9D1D8082F365FFB4FBA3076AA9845B86" box="[1109,1153,367,393]" count="1" pageId="9" pageNumber="592" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D1D8082F365FFB4FB7B076AA9B25B86" box="[1165,1207,367,393]" count="1" pageId="9" pageNumber="592" type="female">1 ♀</specimenCount>
,
<collectingRegion id="49DF85E9F365FFB4FB33076AA86F5B86" box="[1221,1386,367,393]" country="China" name="Guangxi" pageId="9" pageNumber="592">Guangxi Prov.</collectingRegion>
,
<location id="8EC41DD0F365FFB4FA82076AAC225BA2" LSID="urn:lsid:plazi:treatment:03B2FA1DF36BFFB4FF610073A9865832:8EC41DD0F365FFB4FA82076AAC225BA2" country="China" county="Chongzuo City" municipality="Longzhou" name="Mt. Damingshan" pageId="9" pageNumber="592" stateProvince="Guangxi">Mt. Damingshan</location>
,
<date id="FFA56DCBF365FFB4FEC50796ACB05BA2" box="[307,437,403,429]" pageId="9" pageNumber="592" value="2022-07-20">
<collectingDate id="EFE19423F365FFB4FEC50796ACB05BA2" box="[307,437,403,429]" pageId="9" pageNumber="592" value="2022-07-20">20.vii.2022</collectingDate>
</date>
, coll.
<collectorName id="26EE2EDDF365FFB4FE0E0796AF695BA2" box="[504,620,403,429]" pageId="9" pageNumber="592">Qiuyi Zhu</collectorName>
(
<collectionCode id="ED0AD3CEF365FFB4FD8D0796AFC95BA2" box="[635,716,403,429]" pageId="9" pageNumber="592">IMNU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3011880F365FFB4FF3107B3AF245BDE" box="[199,545,438,465]" pageId="9" pageNumber="592" type="distribution">
<paragraph id="8BA44B0BF365FFB4FF3107B3AF245BDE" blockId="9.[151,1437,151,573]" box="[199,545,438,465]" pageId="9" pageNumber="592">
<emphasis id="B96F9719F365FFB4FF3107B3AC645BDF" bold="true" box="[199,353,438,464]" pageId="9" pageNumber="592">Distribution.</emphasis>
<collectingCountry id="F30C0B9BF365FFB4FE9107B2ACAB5BDE" box="[359,430,439,465]" name="China" pageId="9" pageNumber="592">China</collectingCountry>
:
<collectingRegion id="49DF85E9F365FFB4FE4F07B2AF185BDE" box="[441,541,439,465]" country="China" name="Guangxi" pageId="9" pageNumber="592">Guangxi</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C3011880F365FFB4FF3107DEA9205816" pageId="9" pageNumber="592" type="discussion">
<paragraph id="8BA44B0BF365FFB4FF3107DEA9205816" blockId="9.[151,1437,151,573]" pageId="9" pageNumber="592">
<emphasis id="B96F9719F365FFB4FF3107DEAC3E5BFA" bold="true" box="[199,315,475,501]" pageId="9" pageNumber="592">Remarks.</emphasis>
This species can be distinguished by the following features: pygofer lobe ventral margin concave, apex of style fork-like, and aedeagus with gonopore subapical on ventral surface.
</paragraph>
</subSubSection>
<subSubSection id="C3011880F365FFB4FF310426A9865832" box="[199,1155,547,573]" pageId="9" pageNumber="592" type="etymology">
<paragraph id="8BA44B0BF365FFB4FF310426A9865832" blockId="9.[151,1437,151,573]" box="[199,1155,547,573]" pageId="9" pageNumber="592">
<emphasis id="B96F9719F365FFB4FF310426AC4B5832" bold="true" box="[199,334,547,573]" pageId="9" pageNumber="592">Etymology.</emphasis>
The specific epithet refers to the deeply concave pygofer ventral margin.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,173 @@
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<mods:title id="F778B1D845F5F65715073759802FC054">Review of Chinese species of genus Paramritodus Xue &amp; Zhang (Hemiptera: Cicadellidae) with description of a new species</mods:title>
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<paragraph id="8BA44B0BF36DFFBCFF610331AF795F40" blockId="1.[151,636,1332,1395]" box="[151,636,1332,1359]" pageId="1" pageNumber="584">
<heading id="D0ECFC67F36DFFBCFF610331AF795F40" bold="true" box="[151,636,1332,1359]" fontSize="11" level="1" pageId="1" pageNumber="584" reason="1">
<taxonomicName id="4C1B3088F36DFFBCFF610331AF795F40" ID-CoL="8JXJF" authority="Xue &amp; Zhang" authorityName="Xue &amp; Zhang, 2020:" authorityYear="1451" box="[151,636,1332,1359]" class="Insecta" family="Cicadellidae" genus="Paramritodus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="584" phylum="Arthropoda" rank="species" species="introflexus">
<emphasis id="B96F9719F36DFFBCFF610331AF795F40" bold="true" box="[151,636,1332,1359]" pageId="1" pageNumber="584">
<emphasis id="B96F9719F36DFFBCFF610331ACCC5F40" bold="true" box="[151,457,1332,1359]" italics="true" pageId="1" pageNumber="584">Paramritodus introflexus</emphasis>
Xue &amp; Zhang
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<paragraph id="8BA44B0BF36DFFBCFF61035DAC5A5F7C" blockId="1.[151,636,1332,1395]" box="[151,351,1368,1395]" pageId="1" pageNumber="584">
(
<figureCitation id="1320578EF36DFFBCFF69035DADE35F7C" box="[159,230,1368,1395]" captionStart="FIGURES 16" captionStartId="2.[151,264,1309,1333]" captionTargetBox="[350,1238,164,1299]" captionTargetId="figure-37@2.[350,1238,164,1299]" captionTargetPageId="2" captionText="FIGURES 16. Paramritodus dorsal habitus, male. 1, P. introflexus; 2, P. pistacious; 3, P. podocarpus; 4, P. spatiosus; 5, P. triangulus; 6, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745523" httpUri="https://zenodo.org/record/13745523/files/figure.png" pageId="1" pageNumber="584">Figs 1</figureCitation>
,
<figureCitation id="1320578EF36DFFBCFF02035CAC015F7C" box="[244,260,1369,1395]" captionStart="FIGURES 712" captionStartId="3.[151,264,1022,1046]" captionTargetBox="[348,1240,309,999]" captionTargetId="figure-65@3.[348,1240,309,999]" captionTargetPageId="3" captionText="FIGURES 712. Facial aspect of Paramritodus, male. 7, P. introflexus; 8, P. pistacious; 9, P. podocarpus; 10, P. spatiosus; 11, P. triangulus; 12, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745525" httpUri="https://zenodo.org/record/13745525/files/figure.png" pageId="1" pageNumber="584">7</figureCitation>
,
<figureCitation id="1320578EF36DFFBCFEE6035DAC535F7C" box="[272,342,1368,1395]" captionStart="FIGURES 1315" captionStartId="3.[151,264,1623,1647]" captionTargetBox="[156,1431,1148,1597]" captionTargetId="figure-99@3.[152,1436,1143,1600]" captionTargetPageId="3" captionText="FIGURES 1315. Paramritodus introflexus. 13, Genital capsule, lateral view; 14. Genital capsule, dorsal view; 15, Aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13745529" httpUri="https://zenodo.org/record/13745529/files/figure.png" pageId="1" pageNumber="584">1315</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3011880F36DFFBCFF610398AE1C5FBA" box="[151,793,1437,1461]" pageId="1" pageNumber="584" type="reference_group">
<paragraph id="8BA44B0BF36DFFBCFF610398AE1C5FBA" blockId="1.[151,793,1437,1461]" box="[151,793,1437,1461]" pageId="1" pageNumber="584">
<treatmentCitationGroup id="AB0B6C25F36DFFBCFF610398AE1C5FBA" box="[151,793,1437,1461]" pageId="1" pageNumber="584">
<treatmentCitation id="0ABA6D1AF36DFFBCFF610398AFAC5FBA" author="Xue, Q. Q. &amp; Zhang, Y. L." box="[151,681,1437,1461]" page="1451" pageId="1" pageNumber="584" year="2020">
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<emphasis id="B96F9719F36DFFBCFF610398AC905FBA" box="[151,405,1437,1461]" italics="true" pageId="1" pageNumber="584">Paramritodus introflexus</emphasis>
<bibRefCitation id="EF8A36FAF36DFFBCFE6A0398AFAC5FBA" author="Xue, Q. Q. &amp; Zhang, Y. L." box="[412,681,1437,1461]" pageId="1" pageNumber="584" pagination="1438 - 1463" refId="ref4537" refString="Xue, Q. Q. &amp; Zhang, Y. L. (2020) Phylogeny and revision of the Oriental leafhopper genus Amritodus (Hemiptera: Cicadellidae: Idiocerini). Zoological Journal of the Linnean Society, 189 (4), 1438 - 1463. https: // doi. org / 10.1093 / zoolinnean / zlz 129" type="journal article" year="2020">Xue &amp; Zhang, 2020: 1451</bibRefCitation>
</taxonomicName>
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, figs 7, 16.
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</paragraph>
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<subSubSection id="C3011880F36DFFBCFF6103E6AE975CDA" pageId="1" pageNumber="584" type="diagnosis">
<paragraph id="8BA44B0BF36DFFBCFF6103E6AC545C4A" blockId="1.[151,1437,1507,1966]" pageId="1" pageNumber="584">
<emphasis id="B96F9719F36DFFBCFF6103E6AC175FF2" bold="true" box="[151,274,1507,1533]" pageId="1" pageNumber="584">Diagnosis.</emphasis>
Crown (
<figureCitation id="1320578EF36DFFBCFE8403E6ACBD5FF2" box="[370,440,1507,1533]" captionStart="FIGURES 16" captionStartId="2.[151,264,1309,1333]" captionTargetBox="[350,1238,164,1299]" captionTargetId="figure-37@2.[350,1238,164,1299]" captionTargetPageId="2" captionText="FIGURES 16. Paramritodus dorsal habitus, male. 1, P. introflexus; 2, P. pistacious; 3, P. podocarpus; 4, P. spatiosus; 5, P. triangulus; 6, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745523" httpUri="https://zenodo.org/record/13745523/files/figure.png" pageId="1" pageNumber="584">Figs 1</figureCitation>
,
<figureCitation id="1320578EF36DFFBCFE3303E6ACD15FF2" box="[453,468,1507,1533]" captionStart="FIGURES 712" captionStartId="3.[151,264,1022,1046]" captionTargetBox="[348,1240,309,999]" captionTargetId="figure-65@3.[348,1240,309,999]" captionTargetPageId="3" captionText="FIGURES 712. Facial aspect of Paramritodus, male. 7, P. introflexus; 8, P. pistacious; 9, P. podocarpus; 10, P. spatiosus; 11, P. triangulus; 12, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745525" httpUri="https://zenodo.org/record/13745525/files/figure.png" pageId="1" pageNumber="584">7</figureCitation>
) yellow; gena white and lora yellowish. Pronotum (
<figureCitation id="1320578EF36DFFBCFBE503E6A9575FF2" box="[1043,1106,1507,1533]" captionStart="FIGURES 16" captionStartId="2.[151,264,1309,1333]" captionTargetBox="[350,1238,164,1299]" captionTargetId="figure-37@2.[350,1238,164,1299]" captionTargetPageId="2" captionText="FIGURES 16. Paramritodus dorsal habitus, male. 1, P. introflexus; 2, P. pistacious; 3, P. podocarpus; 4, P. spatiosus; 5, P. triangulus; 6, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745523" httpUri="https://zenodo.org/record/13745523/files/figure.png" pageId="1" pageNumber="584">Fig. 1</figureCitation>
) yellow greenish with pair of triangular black markings on anterior margin. Mesonotum (
<figureCitation id="1320578EF36DFFBCFCDB0002AE685C2E" box="[813,877,1543,1569]" captionStart="FIGURES 16" captionStartId="2.[151,264,1309,1333]" captionTargetBox="[350,1238,164,1299]" captionTargetId="figure-37@2.[350,1238,164,1299]" captionTargetPageId="2" captionText="FIGURES 16. Paramritodus dorsal habitus, male. 1, P. introflexus; 2, P. pistacious; 3, P. podocarpus; 4, P. spatiosus; 5, P. triangulus; 6, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745523" httpUri="https://zenodo.org/record/13745523/files/figure.png" pageId="1" pageNumber="584">Fig. 1</figureCitation>
) yellow greenish, basal triangles black, scutellum yellow greenish.
</paragraph>
<paragraph id="8BA44B0BF36DFFBCFF31004AAE975CDA" blockId="1.[151,1437,1507,1966]" pageId="1" pageNumber="584">
Pygofer (
<figureCitation id="1320578EF36DFFBCFEC7004AACB25C66" box="[305,439,1615,1641]" captionStart="FIGURES 1315" captionStartId="3.[151,264,1623,1647]" captionTargetBox="[156,1431,1148,1597]" captionTargetId="figure-99@3.[152,1436,1143,1600]" captionTargetPageId="3" captionText="FIGURES 1315. Paramritodus introflexus. 13, Genital capsule, lateral view; 14. Genital capsule, dorsal view; 15, Aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13745529" httpUri="https://zenodo.org/record/13745529/files/figure.png" pageId="1" pageNumber="584">Figs 13, 14</figureCitation>
) narrow basally, gradually broadened, narrow and rounded terminally; inner processes curved laterally and exceeding the caudal margin slightly; anal tube surpassing pygofer lobe dorsal margin and enlarged terminally. Aedeagal shaft (
<figureCitation id="1320578EF36DFFBCFDB40092AF925CBD" box="[578,663,1687,1714]" captionStart="FIGURES 1315" captionStartId="3.[151,264,1623,1647]" captionTargetBox="[156,1431,1148,1597]" captionTargetId="figure-99@3.[152,1436,1143,1600]" captionTargetPageId="3" captionText="FIGURES 1315. Paramritodus introflexus. 13, Genital capsule, lateral view; 14. Genital capsule, dorsal view; 15, Aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13745529" httpUri="https://zenodo.org/record/13745529/files/figure.png" pageId="1" pageNumber="584">Fig. 15</figureCitation>
) obviously concave subapically in lateral view, with basal pair of slender processes on ventral surface; gonopore near middle of shaft.
</paragraph>
</subSubSection>
<subSubSection id="C3011880F36DFFBCFF3100DAADF95D12" pageId="1" pageNumber="584" type="materials_examined">
<paragraph id="8BA44B0BF36DFFBCFF3100DAADF95D12" blockId="1.[151,1437,1507,1966]" pageId="1" pageNumber="584">
<emphasis id="B96F9719F36DFFBCFF3100DAACB35CF6" bold="true" box="[199,438,1759,1785]" pageId="1" pageNumber="584">Material examined.</emphasis>
<materialsCitation id="3B734156F36DFFBCFE3500DAADFD5D12" collectingDate="2020-07-10" collectionCode="IMNU" collectorName="Bin Zhang" country="China" location="Mt. Gaoligongshan" pageId="1" pageNumber="584" specimenCount="19" specimenCount-female="11" specimenCount-male="8" stateProvince="Yunnan">
<collectingCountry id="F30C0B9BF36DFFBCFE3500DAAF0F5CF6" box="[451,522,1759,1785]" name="China" pageId="1" pageNumber="584">China</collectingCountry>
:
<specimenCount id="9D1D8082F36DFFBCFDEC00DAAF495CF5" box="[538,588,1759,1786]" count="8" pageId="1" pageNumber="584" type="male">8 ♂</specimenCount>
,
<specimenCount id="9D1D8082F36DFFBCFDA800DAAF9F5CF5" box="[606,666,1759,1786]" count="11" pageId="1" pageNumber="584" type="female">11 ♀</specimenCount>
,
<collectingRegion id="49DF85E9F36DFFBCFD5B00E5AE485CF5" box="[685,845,1760,1786]" country="China" name="Yunnan" pageId="1" pageNumber="584">Yunnan Prov.</collectingRegion>
,
<location id="8EC41DD0F36DFFBCFCAB00E5A9385CF6" LSID="urn:lsid:plazi:treatment:03B2FA1DF36DFFBCFF610331AF4D5DA1:8EC41DD0F36DFFBCFCAB00E5A9385CF6" box="[861,1085,1759,1786]" country="China" name="Mt. Gaoligongshan" pageId="1" pageNumber="584" stateProvince="Yunnan">Mt. Gaoligongshan</location>
,
<date id="FFA56DCBF36DFFBCFBB900DAA9D45CF6" box="[1103,1233,1759,1785]" pageId="1" pageNumber="584" value="2020-07-10">
<collectingDate id="EFE19423F36DFFBCFBB900DAA9D45CF6" box="[1103,1233,1759,1785]" pageId="1" pageNumber="584" value="2020-07-10">10.vii.2020</collectingDate>
</date>
, coll.
<collectorName id="26EE2EDDF36DFFBCFAD600DAA8985CF6" box="[1312,1437,1759,1785]" pageId="1" pageNumber="584">Bin Zhang</collectorName>
(
<collectionCode id="ED0AD3CEF36DFFBCFF690106ADF55D12" box="[159,240,1795,1821]" pageId="1" pageNumber="584">IMNU</collectionCode>
)
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.
</paragraph>
</subSubSection>
<subSubSection id="C3011880F36DFFBCFF310122AF245D4E" box="[199,545,1831,1857]" pageId="1" pageNumber="584" type="distribution">
<paragraph id="8BA44B0BF36DFFBCFF310122AF245D4E" blockId="1.[151,1437,1507,1966]" box="[199,545,1831,1857]" pageId="1" pageNumber="584">
<emphasis id="B96F9719F36DFFBCFF310122AC645D4E" bold="true" box="[199,353,1831,1857]" pageId="1" pageNumber="584">Distribution.</emphasis>
<collectingCountry id="F30C0B9BF36DFFBCFE910122ACAF5D4E" box="[359,426,1831,1857]" name="China" pageId="1" pageNumber="584">China</collectingCountry>
(
<collectingRegion id="49DF85E9F36DFFBCFE4F0122AF105D4E" box="[441,533,1831,1857]" country="China" name="Yunnan" pageId="1" pageNumber="584">Yunnan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C3011880F36DFFBCFF31014EAF4D5DA1" pageId="1" pageNumber="584" type="discussion">
<paragraph id="8BA44B0BF36DFFBCFF31014EAF4D5DA1" blockId="1.[151,1437,1507,1966]" pageId="1" pageNumber="584">
<emphasis id="B96F9719F36DFFBCFF31014EAC3E5D6A" bold="true" box="[199,315,1867,1893]" pageId="1" pageNumber="584">Remarks.</emphasis>
This species is similar to
<taxonomicName id="4C1B3088F36DFFBCFDA90149AFE85D6A" authorityName="Xue &amp; Zhang, 2020:" authorityYear="1457" box="[607,749,1867,1893]" class="Insecta" family="Cicadellidae" genus="Paramritodus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="584" phylum="Arthropoda" rank="species" species="triangulus">
<emphasis id="B96F9719F36DFFBCFDA90149AFE85D6A" box="[607,749,1867,1893]" italics="true" pageId="1" pageNumber="584">P. triangulus</emphasis>
</taxonomicName>
, but can be distinguished from the latter by the inner process of the pygofer slightly surpassing the caudal margin, the aedeagal shaft obviously concave subapically and aedeagal processes shorter than in
<taxonomicName id="4C1B3088F36DFFBCFE440191AF445DA2" authorityName="Xue &amp; Zhang, 2020:" authorityYear="1457" box="[434,577,1939,1965]" class="Insecta" family="Cicadellidae" genus="Paramritodus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="584" phylum="Arthropoda" rank="species" species="triangulus">
<emphasis id="B96F9719F36DFFBCFE440191AF445DA2" box="[434,577,1939,1965]" italics="true" pageId="1" pageNumber="584">P. triangulus</emphasis>
</taxonomicName>
.
</paragraph>
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</treatment>
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View file

@ -0,0 +1,290 @@
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<mods:title id="754CBC8E0A754ECDAA2FE932A50C5828">Review of Chinese species of genus Paramritodus Xue &amp; Zhang (Hemiptera: Cicadellidae) with description of a new species</mods:title>
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<mods:date id="88F62A2972713DD89C376C6FFED0F5D4">2024</mods:date>
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<paragraph id="8BA44B0BF36EFFBFFF6103AEAE445FCA" blockId="2.[151,833,1451,1514]" box="[151,833,1451,1478]" pageId="2" pageNumber="585">
<heading id="D0ECFC67F36EFFBFFF6103AEAE445FCA" bold="true" box="[151,833,1451,1478]" fontSize="11" level="1" pageId="2" pageNumber="585" reason="1">
<taxonomicName id="4C1B3088F36EFFBFFF6103AEAE445FCA" ID-CoL="8JXJG" authority="Huang &amp; Maldonado Capriles" authorityName="Huang &amp; Maldonado Capriles" box="[151,833,1451,1478]" class="Insecta" family="Cicadellidae" genus="Paramritodus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="585" phylum="Arthropoda" rank="species" species="pistacious">
<emphasis id="B96F9719F36EFFBFFF6103AEAE445FCA" bold="true" box="[151,833,1451,1478]" pageId="2" pageNumber="585">
<emphasis id="B96F9719F36EFFBFFF6103AEACBB5FC9" bold="true" box="[151,446,1451,1478]" italics="true" pageId="2" pageNumber="585">Paramritodus pistacious</emphasis>
Huang &amp; Maldonado Capriles
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C3011880F36EFFBFFF6103D5AC5A5FE5" box="[151,351,1487,1514]" pageId="2" pageNumber="585" type="description">
<paragraph id="8BA44B0BF36EFFBFFF6103D5AC5A5FE5" blockId="2.[151,833,1451,1514]" box="[151,351,1487,1514]" pageId="2" pageNumber="585">
(
<figureCitation id="1320578EF36EFFBFFF6903D5ADED5FE5" box="[159,232,1488,1514]" captionStart="FIGURES 16" captionStartId="2.[151,264,1309,1333]" captionTargetBox="[350,1238,164,1299]" captionTargetId="figure-37@2.[350,1238,164,1299]" captionTargetPageId="2" captionText="FIGURES 16. Paramritodus dorsal habitus, male. 1, P. introflexus; 2, P. pistacious; 3, P. podocarpus; 4, P. spatiosus; 5, P. triangulus; 6, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745523" httpUri="https://zenodo.org/record/13745523/files/figure.png" pageId="2" pageNumber="585">Figs 2</figureCitation>
,
<figureCitation id="1320578EF36EFFBFFF0203D5AC065FE5" box="[244,259,1488,1514]" captionStart="FIGURES 712" captionStartId="3.[151,264,1022,1046]" captionTargetBox="[348,1240,309,999]" captionTargetId="figure-65@3.[348,1240,309,999]" captionTargetPageId="3" captionText="FIGURES 712. Facial aspect of Paramritodus, male. 7, P. introflexus; 8, P. pistacious; 9, P. podocarpus; 10, P. spatiosus; 11, P. triangulus; 12, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745525" httpUri="https://zenodo.org/record/13745525/files/figure.png" pageId="2" pageNumber="585">8</figureCitation>
,
<figureCitation id="1320578EF36EFFBFFEE603CAAC525FE5" box="[272,343,1487,1514]" captionStart="FIGURES 1625" captionStartId="5.[151,264,1909,1933]" captionTargetBox="[178,1410,189,1874]" captionTargetId="figure-17@5.[169,1419,181,1884]" captionTargetPageId="5" captionText="FIGURES 1625. Paramritodus pistacious. 16, Dorsal and ventral abdominal apodemes, dorsal view; 17, Genital capsule, lateral view; 18. Genital capsule, dorsal view; 19, Aedeagus, lateral view; 20, Aedeagus, ventral view; 21, Connective lateral view; 22, Connective ventral view; 23, Style, lateral view; 24, Female genital, ventral view; 25. Female genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13745531" httpUri="https://zenodo.org/record/13745531/files/figure.png" pageId="2" pageNumber="585">1625</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3011880F36EFFBFFF610011AEBF5C44" pageId="2" pageNumber="585" type="reference_group">
<paragraph id="8BA44B0BF36EFFBFFF610011A9995C23" blockId="2.[151,1180,1556,1611]" box="[151,1180,1556,1581]" pageId="2" pageNumber="585">
<taxonomicName id="4C1B3088F36EFFBFFF610011AE045C23" ID-CoL="8HV58" authority="Huang &amp; Maldonado Capriles, 1992: 2" authorityName="Huang &amp; Maldonado Capriles" authorityPageNumber="2" authorityYear="1992" box="[151,769,1556,1581]" class="Insecta" family="Cicadellidae" genus="Amritodus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="585" phylum="Arthropoda" rank="species" species="pistacious">
<emphasis id="B96F9719F36EFFBFFF610011AC6B5C23" box="[151,366,1556,1580]" italics="true" pageId="2" pageNumber="585">Amritodus pistacious</emphasis>
Huang &amp; Maldonado Capriles, 1992: 2
</taxonomicName>
, fig. 1.
<typeStatus id="54A0F5A9F36EFFBFFCB80011AEA95C23" box="[846,940,1556,1580]" pageId="2" pageNumber="585" type="holotype">Holotype</typeStatus>
<specimenCount id="9D1D8082F36EFFBFFC450010AECD5C23" box="[947,968,1557,1580]" count="1" pageId="2" pageNumber="585" type="male"></specimenCount>
, (NMNS, examined)
</paragraph>
<paragraph id="8BA44B0BF36EFFBFFF610036AEBF5C44" blockId="2.[151,1180,1556,1611]" box="[151,954,1587,1611]" pageId="2" pageNumber="585">
<treatmentCitationGroup id="AB0B6C25F36EFFBFFF610036AEBF5C44" box="[151,954,1587,1611]" pageId="2" pageNumber="585">
<treatmentCitation id="0ABA6D1AF36EFFBFFF610036AFDF5C44" author="Cai, P. &amp; He, J. H. &amp; Gu, X. L." box="[151,730,1587,1611]" page="199" pageId="2" pageNumber="585" year="2001">
<taxonomicName id="4C1B3088F36EFFBFFF610036AFDF5C44" ID-CoL="8HV56" authority="Cai &amp; He, Cai et al., 2001: 199" authorityName="Cai &amp; He, Cai" authorityPageNumber="199" authorityYear="2001" box="[151,730,1587,1611]" class="Insecta" family="Cicadellidae" genus="Amritodus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="585" phylum="Arthropoda" rank="species" species="flavocapitatus">
<emphasis id="B96F9719F36EFFBFFF610036AC925C44" box="[151,407,1587,1611]" italics="true" pageId="2" pageNumber="585">Amritodus flavocapitatus</emphasis>
<bibRefCitation id="EF8A36FAF36EFFBFFE6B0036AFDF5C44" author="Cai, P. &amp; He, J. H. &amp; Gu, X. L." box="[413,730,1587,1611]" pageId="2" pageNumber="585" pagination="185 - 218" refId="ref3968" refString="Cai, P., He, J. H. &amp; Gu, X. L. (2001) Homoptera, Cicadellidae. In: Wu, H. &amp; Pan, C. W. (Eds.), Insects of Tianmushan National Nature Reserve. Science Press, Beijing, pp. 185 - 218." type="book chapter" year="2001">Cai &amp; He, Cai et al., 2001: 199</bibRefCitation>
</taxonomicName>
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, figs 5461.
<emphasis id="B96F9719F36EFFBFFCAB0036AEBF5C44" bold="true" box="[861,954,1587,1611]" pageId="2" pageNumber="585">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
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<subSubSection id="C3011880F36EFFBFFF61007CAECB5D2C" pageId="2" pageNumber="585" type="diagnosis">
<paragraph id="8BA44B0BF36EFFBFFF61007CAFC45CB8" blockId="2.[151,1437,1657,2043]" pageId="2" pageNumber="585">
<emphasis id="B96F9719F36EFFBFFF61007CAC175C9C" bold="true" box="[151,274,1657,1683]" pageId="2" pageNumber="585">Diagnosis.</emphasis>
Crown and face (
<figureCitation id="1320578EF36EFFBFFE23007CAF185C9C" box="[469,541,1657,1683]" captionStart="FIGURES 16" captionStartId="2.[151,264,1309,1333]" captionTargetBox="[350,1238,164,1299]" captionTargetId="figure-37@2.[350,1238,164,1299]" captionTargetPageId="2" captionText="FIGURES 16. Paramritodus dorsal habitus, male. 1, P. introflexus; 2, P. pistacious; 3, P. podocarpus; 4, P. spatiosus; 5, P. triangulus; 6, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745523" httpUri="https://zenodo.org/record/13745523/files/figure.png" pageId="2" pageNumber="585">Figs 2</figureCitation>
,
<figureCitation id="1320578EF36EFFBFFDDE007CAF335C9C" box="[552,566,1657,1683]" captionStart="FIGURES 712" captionStartId="3.[151,264,1022,1046]" captionTargetBox="[348,1240,309,999]" captionTargetId="figure-65@3.[348,1240,309,999]" captionTargetPageId="3" captionText="FIGURES 712. Facial aspect of Paramritodus, male. 7, P. introflexus; 8, P. pistacious; 9, P. podocarpus; 10, P. spatiosus; 11, P. triangulus; 12, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745525" httpUri="https://zenodo.org/record/13745525/files/figure.png" pageId="2" pageNumber="585">8</figureCitation>
) yellowish brown, without black spots. Pronotum, mesoscutum and scutellum (
<figureCitation id="1320578EF36EFFBFFF690098ADE65CB8" box="[159,227,1693,1719]" captionStart="FIGURES 16" captionStartId="2.[151,264,1309,1333]" captionTargetBox="[350,1238,164,1299]" captionTargetId="figure-37@2.[350,1238,164,1299]" captionTargetPageId="2" captionText="FIGURES 16. Paramritodus dorsal habitus, male. 1, P. introflexus; 2, P. pistacious; 3, P. podocarpus; 4, P. spatiosus; 5, P. triangulus; 6, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745523" httpUri="https://zenodo.org/record/13745523/files/figure.png" pageId="2" pageNumber="585">Fig. 2</figureCitation>
) ocher yellow lacking the black markings.
</paragraph>
<paragraph id="8BA44B0BF36EFFBFFF3100C4AECB5D2C" blockId="2.[151,1437,1657,2043]" pageId="2" pageNumber="585">
Pygofer (
<figureCitation id="1320578EF36EFFBFFEC400C4AC835CD4" box="[306,390,1729,1755]" captionStart="FIGURES 1625" captionStartId="5.[151,264,1909,1933]" captionTargetBox="[178,1410,189,1874]" captionTargetId="figure-17@5.[169,1419,181,1884]" captionTargetPageId="5" captionText="FIGURES 1625. Paramritodus pistacious. 16, Dorsal and ventral abdominal apodemes, dorsal view; 17, Genital capsule, lateral view; 18. Genital capsule, dorsal view; 19, Aedeagus, lateral view; 20, Aedeagus, ventral view; 21, Connective lateral view; 22, Connective ventral view; 23, Style, lateral view; 24, Female genital, ventral view; 25. Female genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13745531" httpUri="https://zenodo.org/record/13745531/files/figure.png" pageId="2" pageNumber="585">Fig. 17</figureCitation>
) with elongate caudal lobe; inner processes short and curved apically, not reaching caudal margin, apex hook-shaped. Aedeagal shaft (
<figureCitation id="1320578EF36EFFBFFD8900E0AFFB5CF0" box="[639,766,1765,1791]" captionStart="FIGURES 1625" captionStartId="5.[151,264,1909,1933]" captionTargetBox="[178,1410,189,1874]" captionTargetId="figure-17@5.[169,1419,181,1884]" captionTargetPageId="5" captionText="FIGURES 1625. Paramritodus pistacious. 16, Dorsal and ventral abdominal apodemes, dorsal view; 17, Genital capsule, lateral view; 18. Genital capsule, dorsal view; 19, Aedeagus, lateral view; 20, Aedeagus, ventral view; 21, Connective lateral view; 22, Connective ventral view; 23, Style, lateral view; 24, Female genital, ventral view; 25. Female genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13745531" httpUri="https://zenodo.org/record/13745531/files/figure.png" pageId="2" pageNumber="585">Figs 19, 20</figureCitation>
) with subapex curved dorsally in lateral view, without basal pair of processes; dorsal apodeme poorly developed; gonopore subapical.
</paragraph>
</subSubSection>
<subSubSection id="C3011880F36EFFBFFF310128AC585DF4" pageId="2" pageNumber="585" type="materials_examined">
<paragraph id="8BA44B0BF36EFFBFFF310128AC585DF4" blockId="2.[151,1437,1657,2043]" pageId="2" pageNumber="585">
<emphasis id="B96F9719F36EFFBFFF310128ACB45D48" bold="true" box="[199,433,1837,1863]" pageId="2" pageNumber="585">Material examined.</emphasis>
<typeStatus id="54A0F5A9F36EFFBFFE4F0128AF245D48" box="[441,545,1837,1863]" pageId="2" pageNumber="585" type="holotype">Holotype</typeStatus>
<specimenCount id="9D1D8082F36EFFBFFDDF012BAF455D47" box="[553,576,1838,1864]" count="1" pageId="2" pageNumber="585" type="male"></specimenCount>
,
<specimenCount id="9D1D8082F36EFFBFFDBB0128AF7D5D47" box="[589,632,1837,1864]" count="5" pageId="2" pageNumber="585" type="female">5 ♀</specimenCount>
<materialsCitation id="3B734156F36EFFBFFD76012BAC865D63" collectingDate="1987-07-15" collectionCode="NMNS" collectorName="C. T. Yang" country="China" location="Province" pageId="2" pageNumber="585" specimenCount="1" specimenCount-male="1" stateProvince="Taichung" typeStatus="paratype">
<typeStatus id="54A0F5A9F36EFFBFFD76012BAFF55D47" box="[640,752,1838,1864]" pageId="2" pageNumber="585" type="paratype">Paratypes</typeStatus>
,
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, Taiwan
<location id="8EC41DD0F36EFFBFFC4B0128A9265D48" LSID="urn:lsid:plazi:treatment:03B2FA1DF36EFFBEFF6103AEAE755B12:8EC41DD0F36EFFBFFC4B0128A9265D48" box="[957,1059,1837,1863]" country="China" name="Province" pageId="2" pageNumber="585" stateProvince="Taichung">Province</location>
,
<collectingRegion id="49DF85E9F36EFFBFFBD90128A99F5D48" box="[1071,1178,1837,1863]" country="Taiwan" name="Taichung" pageId="2" pageNumber="585">Taizhong</collectingRegion>
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<collectingDate id="EFE19423F36EFFBFFB530128A82D5D47" box="[1189,1320,1837,1864]" pageId="2" pageNumber="585" value="1987-07-15">15.vii.1987</collectingDate>
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, coll.
<collectorName id="26EE2EDDF36EFFBFFA9A0128ADD45D63" pageId="2" pageNumber="585">C.T. Yang</collectorName>
(
<collectionCode id="ED0AD3CEF36EFFBFFF120154AC3E5D64" box="[228,315,1873,1899]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="2" pageNumber="585">NMNS</collectionCode>
);
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,
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<collectingCountry id="F30C0B9BF36EFFBFFE650154ACEA5D64" box="[403,495,1873,1899]" name="China" pageId="2" pageNumber="585">CHINA</collectingCountry>
, Taiwan
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,
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<collectingMunicipality id="6BC0D171F36EFFBFFCB50154AE715D64" box="[835,884,1873,1899]" pageId="2" pageNumber="585">City</collectingMunicipality>
,
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,
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,
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,
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<collectingDate id="EFE19423F36EFFBFFAFA0154A89D5D64" box="[1292,1432,1873,1899]" pageId="2" pageNumber="585" value="1981-05-06" valueMax="1981-05-11" valueMin="1981-05-06">611.v.1981</collectingDate>
</date>
, coll.
<collectorName id="26EE2EDDF36EFFBFFF260170AC375D80" box="[208,306,1909,1935]" pageId="2" pageNumber="585">K.S. Lin</collectorName>
&amp;
<collectorName id="26EE2EDDF36EFFBFFEAE0170ACB25D80" box="[344,439,1909,1935]" pageId="2" pageNumber="585">S.C. Lin</collectorName>
(
<collectionCode id="ED0AD3CEF36EFFBFFE3E0170AF1A5D80" box="[456,543,1909,1935]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="2" pageNumber="585">NMNS</collectionCode>
);
<specimenCount id="9D1D8082F36EFFBFFDC50170AF645D9F" box="[563,609,1909,1936]" count="2" pageId="2" pageNumber="585" type="male">2 ♂</specimenCount>
,
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,
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,
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<collectingMunicipality id="6BC0D171F36EFFBFFC130170A9135D80" box="[997,1046,1909,1935]" pageId="2" pageNumber="585">City</collectingMunicipality>
,
<collectingRegion id="49DF85E9F36EFFBFFBD50173A9D45D80" box="[1059,1233,1909,1936]" country="Taiwan" name="Nantou" pageId="2" pageNumber="585">Nantou County</collectingRegion>
,
<location id="8EC41DD0F36EFFBFFB2B0170A8375D80" LSID="urn:lsid:plazi:treatment:03B2FA1DF36EFFBEFF6103AEAE755B12:8EC41DD0F36EFFBFFB2B0170A8375D80" box="[1245,1330,1909,1935]" country="Taiwan" county="Province" municipality="City" name="Lushan" pageId="2" pageNumber="585" stateProvince="Taichung">Lushan</location>
,
<quantity id="4CE3E6EEF36EFFBFFAC90170A8925D9F" box="[1343,1431,1909,1936]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="585" unit="m" value="1000.0">
<elevation id="0036AC38F36EFFBFFAC90170A8925D9F" box="[1343,1431,1909,1936]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="585" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
<date id="FFA56DCBF36EFFBFFF61019CAC375DBC" box="[151,306,1945,1971]" pageId="2" pageNumber="585" value="1980-05-27" valueMax="1980-05-31" valueMin="1980-05-27">
<collectingDate id="EFE19423F36EFFBFFF61019CAC375DBC" box="[151,306,1945,1971]" pageId="2" pageNumber="585" value="1980-05-27" valueMax="1980-05-31" valueMin="1980-05-27">2731.v.1980</collectingDate>
</date>
, coll.
<collectorName id="26EE2EDDF36EFFBFFE8F019CACDF5DBC" box="[377,474,1945,1971]" pageId="2" pageNumber="585">K.S. Lin</collectorName>
&amp;
<collectorName id="26EE2EDDF36EFFBFFDF7019FAF735DBC" box="[513,630,1945,1972]" pageId="2" pageNumber="585">L.Y. Chou</collectorName>
(
<collectionCode id="ED0AD3CEF36EFFBFFD70019CAFD85DBC" box="[646,733,1945,1971]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="2" pageNumber="585">NMNS</collectionCode>
);
<specimenCount id="9D1D8082F36EFFBFFD04019CAE255DBB" box="[754,800,1945,1972]" count="1" pageId="2" pageNumber="585" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D1D8082F36EFFBFFCD8019CAE5C5DBB" box="[814,857,1945,1972]" count="3" pageId="2" pageNumber="585" type="female">3 ♀</specimenCount>
</materialsCitation>
,
<materialsCitation id="3B734156F36EFFBFFC9E019CAC5C5DF4" collectingDate="1983-05-03" collectingDateMax="2008-06-15" collectingDateMin="1983-05-03" collectionCode="NMNS, IMNU" collectorName="K. C. Chou &amp; C. C. Pan &amp; Wei Liu" country="Taiwan" county="Province" elevation="800" location="Henan Province" municipality="Paling" pageId="2" pageNumber="585" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Taoyuan" typeStatus="paratype">
<collectingCountry id="F30C0B9BF36EFFBFFC9E019CAEBF5DBC" box="[872,954,1945,1971]" name="Taiwan" pageId="2" pageNumber="585">Taiwan</collectingCountry>
<collectingCounty id="62C53387F36EFFBFFC35019CA92C5DBC" box="[963,1065,1945,1971]" pageId="2" pageNumber="585">Province</collectingCounty>
,
<collectingRegion id="49DF85E9F36EFFBFFBC3019FA9F75DBC" box="[1077,1266,1945,1972]" country="Taiwan" name="Taoyuan" pageId="2" pageNumber="585">Taoyuan County</collectingRegion>
,
<collectingMunicipality id="6BC0D171F36EFFBFFB08019CA84D5DBC" box="[1278,1352,1945,1971]" pageId="2" pageNumber="585">Paling</collectingMunicipality>
,
<quantity id="4CE3E6EEF36EFFBFFAA3019CA89D5DBC" box="[1365,1432,1945,1971]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="585" unit="m" value="800.0">
<elevation id="0036AC38F36EFFBFFAA3019CA89D5DBC" box="[1365,1432,1945,1971]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="585" unit="m" value="800.0">800m</elevation>
</quantity>
,
<date id="FFA56DCBF36EFFBFFF6101B8AC135DD7" box="[151,278,1981,2008]" pageId="2" pageNumber="585" value="1983-05-03" valueMax="1983-05-05" valueMin="1983-05-03">
<collectingDate id="EFE19423F36EFFBFFF6101B8AC135DD7" box="[151,278,1981,2008]" pageId="2" pageNumber="585" value="1983-05-03" valueMax="1983-05-05" valueMin="1983-05-03">35.v.1983</collectingDate>
</date>
, coll.
<collectorName id="26EE2EDDF36EFFBFFEAB01B8ACDD5DD8" box="[349,472,1981,2007]" pageId="2" pageNumber="585">K.C. Chou</collectorName>
&amp;
<collectorName id="26EE2EDDF36EFFBFFE0901B8AF635DD7" box="[511,614,1981,2008]" pageId="2" pageNumber="585">C.C. Pan</collectorName>
(
<collectionCode id="ED0AD3CEF36EFFBFFD8001B8AFC85DD8" box="[630,717,1981,2007]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="2" pageNumber="585">NMNS</collectionCode>
);
<specimenCount id="9D1D8082F36EFFBFFD1501B8AE145DD7" box="[739,785,1981,2008]" count="2" pageId="2" pageNumber="585" type="male">2 ♂</specimenCount>
,
<specimenCount id="9D1D8082F36EFFBFFCE801B8AE4F5DD7" box="[798,842,1981,2008]" count="3" pageId="2" pageNumber="585" type="female">3 ♀</specimenCount>
,
<location id="8EC41DD0F36EFFBFFCAC01BBA9175DD8" LSID="urn:lsid:plazi:treatment:03B2FA1DF36EFFBEFF6103AEAE755B12:8EC41DD0F36EFFBFFCAC01BBA9175DD8" box="[858,1042,1981,2008]" country="Taiwan" county="Province" municipality="Paling" name="Henan Province" pageId="2" pageNumber="585" stateProvince="Taoyuan">Henan Province</location>
, Huixian County,
<date id="FFA56DCBF36EFFBFFB1501B8A85B5DD7" box="[1251,1374,1981,2008]" pageId="2" pageNumber="585" value="2008-06-15">
<collectingDate id="EFE19423F36EFFBFFB1501B8A85B5DD7" box="[1251,1374,1981,2008]" pageId="2" pageNumber="585" value="2008-06-15">15.vi.2008</collectingDate>
</date>
, coll.
<collectorName id="26EE2EDDF36EFFBFFF6101E4ADF75DF4" box="[151,242,2017,2043]" pageId="2" pageNumber="585">Wei Liu</collectorName>
(
<collectionCode id="ED0AD3CEF36EFFBFFEF601E4AC545DF4" box="[256,337,2017,2043]" pageId="2" pageNumber="585">IMNU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3011880F36FFFBEFF310693AF705ABE" box="[199,629,150,177]" pageId="3" pageNumber="586" type="distribution">
<paragraph id="8BA44B0BF36FFFBEFF310693AF705ABE" blockId="3.[151,1436,150,285]" box="[199,629,150,177]" pageId="3" pageNumber="586">
<emphasis id="B96F9719F36FFFBEFF310693AC645ABF" bold="true" box="[199,353,150,176]" pageId="3" pageNumber="586">Distribution.</emphasis>
<collectingCountry id="F30C0B9BF36FFFBEFE910692ACAF5ABE" box="[359,426,151,177]" name="China" pageId="3" pageNumber="586">China</collectingCountry>
(
<collectingCountry id="F30C0B9BF36FFFBEFE4F0692AF155ABE" box="[441,528,151,177]" name="Taiwan" pageId="3" pageNumber="586">Taiwan</collectingCountry>
, Henan).
</paragraph>
</subSubSection>
<subSubSection id="C3011880F36FFFBEFF3106BEAE755B12" pageId="3" pageNumber="586" type="discussion">
<paragraph id="8BA44B0BF36FFFBEFF3106BEAE755B12" blockId="3.[151,1436,150,285]" pageId="3" pageNumber="586">
<emphasis id="B96F9719F36FFFBEFF3106BEAC3E5ADA" bold="true" box="[199,315,187,213]" pageId="3" pageNumber="586">Remarks.</emphasis>
We examined the
<typeStatus id="54A0F5A9F36FFFBEFDE306BEAF405ADA" box="[533,581,187,213]" pageId="3" pageNumber="586">type</typeStatus>
material and clearly showed that
<taxonomicName id="4C1B3088F36FFFBEFC3806BEA9895ADA" box="[974,1164,187,213]" class="Insecta" family="Cicadellidae" genus="Amritodus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="586" phylum="Arthropoda" rank="species" species="flavocapitatus">
<emphasis id="B96F9719F36FFFBEFC3806BEA9895ADA" box="[974,1164,187,213]" italics="true" pageId="3" pageNumber="586">P. flavocapitatus</emphasis>
</taxonomicName>
should be considered a junior synonym of
<taxonomicName id="4C1B3088F36FFFBEFE8606DAACFA5AF7" authorityName="Huang &amp; Maldonado Capriles" authorityYear="1992" box="[368,511,223,248]" class="Insecta" family="Cicadellidae" genus="Amritodus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="586" phylum="Arthropoda" rank="species" species="pistacious">
<emphasis id="B96F9719F36FFFBEFE8606DAACFA5AF7" box="[368,511,223,248]" italics="true" pageId="3" pageNumber="586">P. pistacious</emphasis>
</taxonomicName>
. This species can be distinguished from the other species of
<taxonomicName id="4C1B3088F36FFFBEFB4406DAA84B5AF6" authorityName="Xue &amp; Zhang, 2020:" authorityYear="1449" box="[1202,1358,223,249]" class="Insecta" family="Cicadellidae" genus="Paramritodus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="586" phylum="Arthropoda" rank="genus">
<emphasis id="B96F9719F36FFFBEFB4406DAA84B5AF6" box="[1202,1358,223,249]" italics="true" pageId="3" pageNumber="586">Paramritodus</emphasis>
</taxonomicName>
by the elongate pygofer with short inner process and hook-shaped apex.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,196 @@
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<mods:title id="D944D90E177A51F456FFB447244B605E">Review of Chinese species of genus Paramritodus Xue &amp; Zhang (Hemiptera: Cicadellidae) with description of a new species</mods:title>
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<mods:affiliation id="3375CEED4CA22974F740626441AF8312">College of Life Sciences &amp; Technology, Inner Mongolia Normal University, Hohhot 010022, China &amp; Harrow Beijing City Campus No. 5, 4 th Block Anzhen Xili Chaoyang District Beijing 100029 China</mods:affiliation>
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<mods:affiliation id="74A20C70A73DB9ECCD98A1D9B893C6D7">College of Life Sciences &amp; Technology, Inner Mongolia Normal University, Hohhot 010022, China &amp; Key Laboratory of Biodiversity conservation and Sustainable utilization for College and University of Inner Mongolia Autonomous Region, Hohhot 010022, China &amp; Center for Applied Mathematics Inner Mongolia, Hohhot 010022, China</mods:affiliation>
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<treatment id="03B2FA1DF36FFFB9FF6100E0AF0C5BDE" ID-DOI="http://doi.org/10.5281/zenodo.13745539" ID-Zenodo-Dep="13745539" LSID="urn:lsid:plazi:treatment:03B2FA1DF36FFFB9FF6100E0AF0C5BDE" httpUri="http://treatment.plazi.org/id/03B2FA1DF36FFFB9FF6100E0AF0C5BDE" lastPageId="4" lastPageNumber="587" pageId="3" pageNumber="586">
<subSubSection id="C3011880F36FFFBEFF6100E0AF685CF0" box="[151,621,1765,1792]" pageId="3" pageNumber="586" type="nomenclature">
<paragraph id="8BA44B0BF36FFFBEFF6100E0AF685CF0" blockId="3.[151,621,1765,1828]" box="[151,621,1765,1792]" pageId="3" pageNumber="586">
<heading id="D0ECFC67F36FFFBEFF6100E0AF685CF0" bold="true" box="[151,621,1765,1792]" fontSize="11" level="1" pageId="3" pageNumber="586" reason="1">
<taxonomicName id="4C1B3088F36FFFBEFF6100E0AF685CF0" ID-CoL="8JXJH" authority="Zhang &amp; Li" authorityName="Zhang &amp; Li" box="[151,621,1765,1792]" class="Insecta" family="Cicadellidae" genus="Paramritodus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="586" phylum="Arthropoda" rank="species" species="podocarpus">
<emphasis id="B96F9719F36FFFBEFF6100E0AF685CF0" bold="true" box="[151,621,1765,1792]" pageId="3" pageNumber="586">
<emphasis id="B96F9719F36FFFBEFF6100E0ACD45D0F" bold="true" box="[151,465,1765,1792]" italics="true" pageId="3" pageNumber="586">Paramritodus podocarpus</emphasis>
Zhang &amp; Li
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3011880F36FFFBEFF61010FAC5A5D2B" box="[151,351,1802,1828]" pageId="3" pageNumber="586" type="description">
<paragraph id="8BA44B0BF36FFFBEFF61010FAC5A5D2B" blockId="3.[151,621,1765,1828]" box="[151,351,1802,1828]" pageId="3" pageNumber="586">
(
<figureCitation id="1320578EF36FFFBEFF69010FADE25D2B" box="[159,231,1802,1828]" captionStart="FIGURES 16" captionStartId="2.[151,264,1309,1333]" captionTargetBox="[350,1238,164,1299]" captionTargetId="figure-37@2.[350,1238,164,1299]" captionTargetPageId="2" captionText="FIGURES 16. Paramritodus dorsal habitus, male. 1, P. introflexus; 2, P. pistacious; 3, P. podocarpus; 4, P. spatiosus; 5, P. triangulus; 6, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745523" httpUri="https://zenodo.org/record/13745523/files/figure.png" pageId="3" pageNumber="586">Figs 3</figureCitation>
,
<figureCitation id="1320578EF36FFFBEFF02010FAC015D2B" box="[244,260,1802,1828]" captionStart="FIGURES 712" captionStartId="3.[151,264,1022,1046]" captionTargetBox="[348,1240,309,999]" captionTargetId="figure-65@3.[348,1240,309,999]" captionTargetPageId="3" captionText="FIGURES 712. Facial aspect of Paramritodus, male. 7, P. introflexus; 8, P. pistacious; 9, P. podocarpus; 10, P. spatiosus; 11, P. triangulus; 12, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745525" httpUri="https://zenodo.org/record/13745525/files/figure.png" pageId="3" pageNumber="586">9</figureCitation>
,
<figureCitation id="1320578EF36FFFBEFEE6010FAC525D2B" box="[272,343,1802,1828]" captionStart="FIGURES 2634" captionStartId="6.[151,264,1835,1859]" captionTargetBox="[162,1426,188,1803]" captionTargetId="figure-37@6.[151,1436,181,1811]" captionTargetPageId="6" captionText="FIGURES 2634. Paramritodus podocarpus (Zhang &amp; Li). 26, Dorsal and ventral abdominal apodemes, dorsal view; 27, Genital capsule, lateral view; 28,Aedeagus, lateral view; 29,Aedeagus, ventral view; 30, Connective lateral view; 31, Connective ventral view; 32, Style, lateral view; 33, Female genital, ventral view; 34, Female genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13745535" httpUri="https://zenodo.org/record/13745535/files/figure.png" pageId="3" pageNumber="586">2634</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3011880F36FFFBEFF61014BAFD05D69" box="[151,725,1870,1894]" pageId="3" pageNumber="586" type="reference_group">
<paragraph id="8BA44B0BF36FFFBEFF61014BAFD05D69" blockId="3.[151,725,1870,1894]" box="[151,725,1870,1894]" pageId="3" pageNumber="586">
<treatmentCitationGroup id="AB0B6C25F36FFFBEFF61014BAFD05D69" box="[151,725,1870,1894]" pageId="3" pageNumber="586">
<treatmentCitation id="0ABA6D1AF36FFFBEFF61014BAF745D69" author="Zhang, B. &amp; Li, Z. Z." box="[151,625,1870,1894]" page="730" pageId="3" pageNumber="586" year="2010">
<taxonomicName id="4C1B3088F36FFFBEFF61014BAF745D69" ID-CoL="8JPC6" authority="Zhang &amp; Li, 2010: 730" authorityName="Zhang &amp; Li" authorityPageNumber="730" authorityYear="2010" box="[151,625,1870,1894]" class="Insecta" family="Cicadellidae" genus="Amritodus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="586" phylum="Arthropoda" rank="species" species="podocarpus">
<emphasis id="B96F9719F36FFFBEFF61014BAC7A5D69" box="[151,383,1870,1894]" italics="true" pageId="3" pageNumber="586">Amritodus podocarpus</emphasis>
<bibRefCitation id="EF8A36FAF36FFFBEFE73014BAF745D69" author="Zhang, B. &amp; Li, Z. Z." box="[389,625,1870,1894]" pageId="3" pageNumber="586" pagination="730 - 732" refId="ref4599" refString="Zhang, B. &amp; Li, Z. Z. (2010) Amritodus podocarpus, a new species of the genus Amritodus Anufriev breeding on Podocarpus macrophyllus (Hemiptera, Cicadellidae). Acta Zootaxonomica Sinica, 35, 730 - 732." type="journal article" year="2010">Zhang &amp; Li, 2010: 730</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 19.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3011880F36FFFB9FF610191A8975AF6" lastPageId="4" lastPageNumber="587" pageId="3" pageNumber="586" type="diagnosis">
<paragraph id="8BA44B0BF36FFFBEFF610191ADE35DF9" blockId="3.[151,1437,1940,2038]" pageId="3" pageNumber="586">
<emphasis id="B96F9719F36FFFBEFF610191AC175DA1" bold="true" box="[151,274,1940,1966]" pageId="3" pageNumber="586">Diagnosis.</emphasis>
Crown (
<figureCitation id="1320578EF36FFFBEFE8A0191ACCD5DA1" box="[380,456,1940,1966]" captionStart="FIGURES 16" captionStartId="2.[151,264,1309,1333]" captionTargetBox="[350,1238,164,1299]" captionTargetId="figure-37@2.[350,1238,164,1299]" captionTargetPageId="2" captionText="FIGURES 16. Paramritodus dorsal habitus, male. 1, P. introflexus; 2, P. pistacious; 3, P. podocarpus; 4, P. spatiosus; 5, P. triangulus; 6, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745523" httpUri="https://zenodo.org/record/13745523/files/figure.png" pageId="3" pageNumber="586">Figs 3</figureCitation>
,
<figureCitation id="1320578EF36FFFBEFE2F0191ACED5DA1" box="[473,488,1940,1966]" captionStart="FIGURES 712" captionStartId="3.[151,264,1022,1046]" captionTargetBox="[348,1240,309,999]" captionTargetId="figure-65@3.[348,1240,309,999]" captionTargetPageId="3" captionText="FIGURES 712. Facial aspect of Paramritodus, male. 7, P. introflexus; 8, P. pistacious; 9, P. podocarpus; 10, P. spatiosus; 11, P. triangulus; 12, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745525" httpUri="https://zenodo.org/record/13745525/files/figure.png" pageId="3" pageNumber="586">9</figureCitation>
) yellow, without black spot. Mesonotum (
<figureCitation id="1320578EF36FFFBEFC2A0191A9265DA1" box="[988,1059,1940,1966]" captionStart="FIGURES 16" captionStartId="2.[151,264,1309,1333]" captionTargetBox="[350,1238,164,1299]" captionTargetId="figure-37@2.[350,1238,164,1299]" captionTargetPageId="2" captionText="FIGURES 16. Paramritodus dorsal habitus, male. 1, P. introflexus; 2, P. pistacious; 3, P. podocarpus; 4, P. spatiosus; 5, P. triangulus; 6, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745523" httpUri="https://zenodo.org/record/13745523/files/figure.png" pageId="3" pageNumber="586">Fig. 3</figureCitation>
) yellow without basal triangles. Frontoclypeus, anteclypeus, gena, and lora (
<figureCitation id="1320578EF36FFFBEFD7101BDAFCE5DDD" box="[647,715,1976,2002]" captionStart="FIGURES 712" captionStartId="3.[151,264,1022,1046]" captionTargetBox="[348,1240,309,999]" captionTargetId="figure-65@3.[348,1240,309,999]" captionTargetPageId="3" captionText="FIGURES 712. Facial aspect of Paramritodus, male. 7, P. introflexus; 8, P. pistacious; 9, P. podocarpus; 10, P. spatiosus; 11, P. triangulus; 12, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745525" httpUri="https://zenodo.org/record/13745525/files/figure.png" pageId="3" pageNumber="586">Fig. 9</figureCitation>
) pale yellow. Forewings (
<figureCitation id="1320578EF36FFFBEFC1901BDA9375DDD" box="[1007,1074,1976,2002]" captionStart="FIGURES 16" captionStartId="2.[151,264,1309,1333]" captionTargetBox="[350,1238,164,1299]" captionTargetId="figure-37@2.[350,1238,164,1299]" captionTargetPageId="2" captionText="FIGURES 16. Paramritodus dorsal habitus, male. 1, P. introflexus; 2, P. pistacious; 3, P. podocarpus; 4, P. spatiosus; 5, P. triangulus; 6, P. concavicaudus." figureDoi="http://doi.org/10.5281/zenodo.13745523" httpUri="https://zenodo.org/record/13745523/files/figure.png" pageId="3" pageNumber="586">Fig. 3</figureCitation>
) gray brown except clavus dark brown.
</paragraph>
<paragraph id="8BA44B0BF368FFB9FF310692A8975AF6" blockId="4.[151,1436,151,465]" pageId="4" pageNumber="587">
Pygofer (
<figureCitation id="1320578EF368FFB9FEC60692AC875ABE" box="[304,386,151,177]" captionStart="FIGURES 2634" captionStartId="6.[151,264,1835,1859]" captionTargetBox="[162,1426,188,1803]" captionTargetId="figure-37@6.[151,1436,181,1811]" captionTargetPageId="6" captionText="FIGURES 2634. Paramritodus podocarpus (Zhang &amp; Li). 26, Dorsal and ventral abdominal apodemes, dorsal view; 27, Genital capsule, lateral view; 28,Aedeagus, lateral view; 29,Aedeagus, ventral view; 30, Connective lateral view; 31, Connective ventral view; 32, Style, lateral view; 33, Female genital, ventral view; 34, Female genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13745535" httpUri="https://zenodo.org/record/13745535/files/figure.png" pageId="4" pageNumber="587">Fig. 27</figureCitation>
) inner process not reaching caudal margin. Subgenital plates (
<figureCitation id="1320578EF368FFB9FBB20692A9935ABE" box="[1092,1174,151,177]" captionStart="FIGURES 2634" captionStartId="6.[151,264,1835,1859]" captionTargetBox="[162,1426,188,1803]" captionTargetId="figure-37@6.[151,1436,181,1811]" captionTargetPageId="6" captionText="FIGURES 2634. Paramritodus podocarpus (Zhang &amp; Li). 26, Dorsal and ventral abdominal apodemes, dorsal view; 27, Genital capsule, lateral view; 28,Aedeagus, lateral view; 29,Aedeagus, ventral view; 30, Connective lateral view; 31, Connective ventral view; 32, Style, lateral view; 33, Female genital, ventral view; 34, Female genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13745535" httpUri="https://zenodo.org/record/13745535/files/figure.png" pageId="4" pageNumber="587">Fig. 27</figureCitation>
) distinctly shorter than pygofer. Aedeagus (
<figureCitation id="1320578EF368FFB9FE8306BEACF15ADA" box="[373,500,187,213]" captionStart="FIGURES 2634" captionStartId="6.[151,264,1835,1859]" captionTargetBox="[162,1426,188,1803]" captionTargetId="figure-37@6.[151,1436,181,1811]" captionTargetPageId="6" captionText="FIGURES 2634. Paramritodus podocarpus (Zhang &amp; Li). 26, Dorsal and ventral abdominal apodemes, dorsal view; 27, Genital capsule, lateral view; 28,Aedeagus, lateral view; 29,Aedeagus, ventral view; 30, Connective lateral view; 31, Connective ventral view; 32, Style, lateral view; 33, Female genital, ventral view; 34, Female genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13745535" httpUri="https://zenodo.org/record/13745535/files/figure.png" pageId="4" pageNumber="587">Figs 28, 29</figureCitation>
) without process and dorsal apodeme poorly developed, aedeagal preatrium arcuate in lateral view, gonopore near middle of shaft. Style (
<figureCitation id="1320578EF368FFB9FD1A06DAAE385AF6" box="[748,829,223,249]" captionStart="FIGURES 2634" captionStartId="6.[151,264,1835,1859]" captionTargetBox="[162,1426,188,1803]" captionTargetId="figure-37@6.[151,1436,181,1811]" captionTargetPageId="6" captionText="FIGURES 2634. Paramritodus podocarpus (Zhang &amp; Li). 26, Dorsal and ventral abdominal apodemes, dorsal view; 27, Genital capsule, lateral view; 28,Aedeagus, lateral view; 29,Aedeagus, ventral view; 30, Connective lateral view; 31, Connective ventral view; 32, Style, lateral view; 33, Female genital, ventral view; 34, Female genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13745535" httpUri="https://zenodo.org/record/13745535/files/figure.png" pageId="4" pageNumber="587">Fig. 32</figureCitation>
) subapical extended. Connective (
<figureCitation id="1320578EF368FFB9FB4906DAA83A5AF6" box="[1215,1343,223,249]" captionStart="FIGURES 2634" captionStartId="6.[151,264,1835,1859]" captionTargetBox="[162,1426,188,1803]" captionTargetId="figure-37@6.[151,1436,181,1811]" captionTargetPageId="6" captionText="FIGURES 2634. Paramritodus podocarpus (Zhang &amp; Li). 26, Dorsal and ventral abdominal apodemes, dorsal view; 27, Genital capsule, lateral view; 28,Aedeagus, lateral view; 29,Aedeagus, ventral view; 30, Connective lateral view; 31, Connective ventral view; 32, Style, lateral view; 33, Female genital, ventral view; 34, Female genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13745535" httpUri="https://zenodo.org/record/13745535/files/figure.png" pageId="4" pageNumber="587">Figs 3031</figureCitation>
) small.
</paragraph>
</subSubSection>
<subSubSection id="C3011880F368FFB9FF310707A88B5B4E" pageId="4" pageNumber="587" type="materials_examined">
<paragraph id="8BA44B0BF368FFB9FF310707A88B5B4E" blockId="4.[151,1436,151,465]" pageId="4" pageNumber="587">
<emphasis id="B96F9719F368FFB9FF310707ACAA5B13" bold="true" box="[199,431,258,284]" pageId="4" pageNumber="587">Material examined.</emphasis>
<materialsCitation id="3B734156F368FFB9FE430706AF5D5B4E" collectingDate="2006-06-14" collectionCode="IMNU" collectorName="Bin Zhang" country="China" elevation="500" location="Maolan National Nature Reserve" municipality="Tongren City" pageId="4" pageNumber="587" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Guizhou">
<specimenCount id="9D1D8082F368FFB9FE430706ACDA5B12" box="[437,479,259,285]" count="2" pageId="4" pageNumber="587" type="male">2 ♂</specimenCount>
,
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,
<collectingCountry id="F30C0B9BF368FFB9FDE90706AF7E5B12" box="[543,635,259,285]" name="China" pageId="4" pageNumber="587">CHINA</collectingCountry>
,
<collectingRegion id="49DF85E9F368FFB9FD730706AE545B12" box="[645,849,259,285]" country="China" name="Guizhou" pageId="4" pageNumber="587">Guizhou Province</collectingRegion>
,
<collectingMunicipality id="6BC0D171F368FFB9FCAD0706AEEB5B12" box="[859,1006,259,285]" pageId="4" pageNumber="587">Tongren City</collectingMunicipality>
,
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<elevation id="0036AC38F368FFB9FA840707ADAA5B4E" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="587" unit="m" value="500.0">500 m</elevation>
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,
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<collectingDate id="EFE19423F368FFB9FF4D0722AC305B4E" box="[187,309,295,321]" pageId="4" pageNumber="587" value="2006-06-14">14.vi.2006</collectingDate>
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, coll.
<collectorName id="26EE2EDDF368FFB9FE8F0722ACF55B4E" box="[377,496,295,321]" pageId="4" pageNumber="587">Bin Zhang</collectorName>
(
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</materialsCitation>
;
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<specimenCount id="9D1D8082F368FFB9FD950722AF8A5B4E" box="[611,655,295,321]" count="1" pageId="4" pageNumber="587" type="male">1 ♂</specimenCount>
,
<collectingRegion id="49DF85E9F368FFB9FD6D0722AE6D5B4E" box="[667,872,295,321]" country="China" name="Guizhou" pageId="4" pageNumber="587">Guizhou Province</collectingRegion>
,
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,
<date id="FFA56DCBF368FFB9FBF30722A96D5B4E" box="[1029,1128,295,321]" pageId="4" pageNumber="587" value="2006-05-06">
<collectingDate id="EFE19423F368FFB9FBF30722A96D5B4E" box="[1029,1128,295,321]" pageId="4" pageNumber="587" value="2006-05-06">6.v.2006</collectingDate>
</date>
, coll.
<collectorName id="26EE2EDDF368FFB9FB5D0722A8265B4E" box="[1195,1315,295,321]" pageId="4" pageNumber="587">Bin Zhang</collectorName>
(
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)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3011880F368FFB9FF31074FAF295B6A" box="[199,556,330,357]" pageId="4" pageNumber="587" type="distribution">
<paragraph id="8BA44B0BF368FFB9FF31074FAF295B6A" blockId="4.[151,1436,151,465]" box="[199,556,330,357]" pageId="4" pageNumber="587">
<emphasis id="B96F9719F368FFB9FF31074FAC645B6B" bold="true" box="[199,353,330,356]" pageId="4" pageNumber="587">Distribution.</emphasis>
<collectingCountry id="F30C0B9BF368FFB9FE91074EACAF5B6A" box="[359,426,331,357]" name="China" pageId="4" pageNumber="587">China</collectingCountry>
(
<collectingRegion id="49DF85E9F368FFB9FE4F074EAF255B6A" box="[441,544,331,357]" country="China" name="Guizhou" pageId="4" pageNumber="587">Guizhou</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C3011880F368FFB9FF31076AAF0C5BDE" pageId="4" pageNumber="587" type="discussion">
<paragraph id="8BA44B0BF368FFB9FF31076AAF0C5BDE" blockId="4.[151,1436,151,465]" pageId="4" pageNumber="587">
<emphasis id="B96F9719F368FFB9FF31076AAC3E5B86" bold="true" box="[199,315,367,393]" pageId="4" pageNumber="587">Remarks.</emphasis>
This species can be distinguished by the following features: crown without pair of black spots on either side of midline; style medially concave and extended subapically; aedeagal shaft slender and elongate, and preatrium arcuate in lateral view.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,107 @@
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View file

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<emphasis id="B909EAA9FFA8FF9DFF2AFE40E9E812D1" bold="true" box="[151,437,427,454]" italics="true" pageId="3" pageNumber="545">Eustrophopsis bombinus</emphasis>
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<emphasis id="B909EAA9FFA8FF9DFF2AFE1BE9DA111F" box="[151,391,496,520]" italics="true" pageId="3" pageNumber="545">Eustrophinus bombinus</emphasis>
Seidlitz, 1848: 441
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.
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<emphasis id="B909EAA9FFA8FF9DFF2AFDE4E9CC1130" box="[151,401,527,551]" italics="true" pageId="3" pageNumber="545">Eustrophopsis bombinus</emphasis>
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<bibRefCitation id="EFEC4B4AFFA8FF9DFD2DFDE4EB081130" author="Champion, G. C." box="[656,853,527,551]" pageId="3" pageNumber="545" pagination="1 - 10" refId="ref4747" refString="Champion, G. C. (1916) Notes on Melandryidae (2). The Entomologist's Monthly Magazine, 52, 1 - 10. https: // doi. org / 10.5962 / bhl. part. 23928" type="journal article" year="1916">Champion, 1916: 6</bibRefCitation>
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Distr.:
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—state of
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Distr.:
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<emphasis id="B909EAA9FFA9FF9CFF2AF918E9E61419" bold="true" box="[151,443,1779,1806]" italics="true" pageId="2" pageNumber="544">Eustrophopsis bilunulata</emphasis>
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<emphasis id="B909EAA9FFA9FF9CFF2AF8D3E9CB1447" box="[151,406,1848,1872]" italics="true" pageId="2" pageNumber="544">Eustrophopsis bilunulata</emphasis>
<bibRefCitation id="EFEC4B4AFFA9FF9CFE21F8D3EA781446" author="Pic, M." box="[412,549,1848,1873]" pageId="2" pageNumber="544" pagination="93 - 94" refId="ref5867" refString="Pic, M. (1912) Coleopteres exotiques nouveaux ou peu connus. Echange, 28, 93 - 94." type="journal article" year="1912">Pic, 1912: 93</bibRefCitation>
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;
<treatmentCitation id="0ADC10AAFFA9FF9CFD8DF8D3EAA81446" author="Champion, G. C." box="[560,757,1848,1873]" page="2" pageId="2" pageNumber="544" year="1916">
<bibRefCitation id="EFEC4B4AFFA9FF9CFD8DF8D3EAA81446" author="Champion, G. C." box="[560,757,1848,1873]" pageId="2" pageNumber="544" pagination="1 - 10" refId="ref4747" refString="Champion, G. C. (1916) Notes on Melandryidae (2). The Entomologist's Monthly Magazine, 52, 1 - 10. https: // doi. org / 10.5962 / bhl. part. 23928" type="journal article" year="1916">Champion, 1916: 2</bibRefCitation>
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;
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<bibRefCitation id="EFEC4B4AFFA9FF9CFCBDF8D3EBAD1446" author="Blackwelder, R. E." box="[768,1008,1848,1873]" pageId="2" pageNumber="544" pagination="551 - 763" refId="ref4580" refString="Blackwelder, R. E. (1945) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Bulletin of the United States National Museum, 185, 551 - 763. https: // doi. org / 10.5479 / si. 03629236.185.3" type="journal article" year="1945">Blackwelder 1945: 494</bibRefCitation>
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Distr.:
<collectingCountry id="F36A762BFFA9FF9CFF5EF892E97A1484" box="[227,295,1913,1939]" name="Brazil" pageId="2" pageNumber="544">Brazil</collectingCountry>
—states of Amazonas and
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.
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<emphasis id="B909EAA9FFA9FF9CFF2AFB25E9AE17FE" bold="true" box="[151,499,1230,1257]" italics="true" pageId="2" pageNumber="544">Eustrophopsis bicoloricornis</emphasis>
(
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)
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<taxonomicName id="4C7D4D38FFA9FF9CFF2AFAF8EA1E163C" authority="Pic, 1923: 23" authorityName="Pic" authorityPageNumber="23" authorityYear="1923" box="[151,579,1299,1323]" class="Insecta" family="Tetratomidae" genus="Eustrophinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="544" phylum="Arthropoda" rank="species" species="bicoloricornis">
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<bibRefCitation id="EFEC4B4AFFA9FF9CFE07FAF8EA1E163C" author="Pic, M." box="[442,579,1299,1323]" pageId="2" pageNumber="544" pagination="3 - 32" refId="ref5889" refString="Pic, M. (1923) Nouveautes diverses.. Melanges exotico-entomologiques, 40, 3 - 32." type="journal article" year="1923">Pic, 1923: 23</bibRefCitation>
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;
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<bibRefCitation id="EFEC4B4AFFA9FF9CFDF0FAF8EB60163C" author="Blackwelder, R. E." box="[589,829,1299,1323]" pageId="2" pageNumber="544" pagination="551 - 763" refId="ref4580" refString="Blackwelder, R. E. (1945) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Bulletin of the United States National Museum, 185, 551 - 763. https: // doi. org / 10.5479 / si. 03629236.185.3" type="journal article" year="1945">Blackwelder 1945: 495</bibRefCitation>
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Distr.: recorded for “
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” in the original description, it is unclear if it refers to the Brazilian state of
<collectingRegion id="49B9F859FFA9FF9CFA9BFABFEDC11679" box="[1318,1436,1364,1390]" country="Peru" name="Amazonas" pageId="2" pageNumber="544">Amazonas</collectingRegion>
or the Amazon Rainforest.
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<document id="1766369451A19FC8B6401DA86FFC17A1" ID-DOI="10.11646/zootaxa.5501.4.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="13745413" ID-ZooBank="81A58D39-1C56-421F-B988-FCF8088A94FC" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726051020352" checkinUser="plazi" docAuthor="Begha, Bruno Piotrovski, Souza, Isabella Magalhães De &amp; Oliveira, Sarah Siqueira" docDate="2024" docId="03D487ADFFADFF96FF2AFD81ECE1102E" docLanguage="en" docName="zootaxa.5501.4.4.pdf" docOrigin="Zootaxa 5501 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5501.4.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eustrophopsis laurae Begha, Souza &amp; Oliveira, 2024, sp. nov." docType="treatment" docUuid="D811574E-A968-4FEE-91C5-FD48AA7D3FDF" docUuidSource="ZooBank" docVersion="2" lastPageNumber="550" masterDocId="FFEDFFD5FFABFF9EFFBDFFEBE85D1317" masterDocTitle="Polypore fungus beetles (Coleoptera: Tetratomidae) in a remnant of semi-deciduous forest of the Brazilian Cerrado: checklist and the description of a new species of Eustrophopsis Champion, 1889" masterLastPageNumber="552" masterPageNumber="542" pageNumber="548" updateTime="1726051541809" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C93888AFD9C00EA5C8BBC7D66FD03897">Polypore fungus beetles (Coleoptera: Tetratomidae) in a remnant of semi-deciduous forest of the Brazilian Cerrado: checklist and the description of a new species of Eustrophopsis Champion, 1889</mods:title>
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<mods:namePart id="D89420250BF812F5818F919D16CF8880">Begha, Bruno Piotrovski</mods:namePart>
<mods:affiliation id="26B926D42D2350A5984144E4C076069C">Universidade Federal de Goiás (UFG), Instituto de Ciências Biológicas (ICB), Departamento de Ecologia (DECOL), Goiânia, Goiás, Brazil.</mods:affiliation>
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<mods:namePart id="881D4EE19A0062639D3064E224DCC48F">Souza, Isabella Magalhães De</mods:namePart>
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<mods:affiliation id="1C1E3574E1913531CC6546AD81D5B23D">Universidade Federal de Goiás (UFG), Instituto de Ciências Biológicas (ICB), Departamento de Ecologia (DECOL), Goiânia, Goiás, Brazil.</mods:affiliation>
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<mods:affiliation id="9BFCD201589190A6720EDC14E51743A8">Universidade Federal de Goiás (UFG), Instituto de Ciências Biológicas (ICB), Departamento de Ecologia (DECOL), Goiânia, Goiás, Brazil.</mods:affiliation>
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<paragraph id="8BC236BBFFADFF98FF2AFD81EA9B1193" blockId="6.[151,1194,618,717]" box="[151,710,618,645]" pageId="6" pageNumber="548">
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<taxonomicName id="4C7D4D38FFADFF98FF2AFD81E9C41192" authorityName="Begha &amp; Souza &amp; Oliveira" authorityYear="2024" box="[151,409,618,645]" class="Insecta" family="Tetratomidae" genus="Eustrophopsis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="548" phylum="Arthropoda" rank="species" species="laurae" status="sp. nov.">
<emphasis id="B909EAA9FFADFF98FF2AFD81E9C41192" bold="true" box="[151,409,618,645]" italics="true" pageId="6" pageNumber="548">Eustrophopsis laurae</emphasis>
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<taxonomicNameLabel id="A23A57D2FFADFF98FE1CFD80E9A31192" box="[417,510,619,645]" pageId="6" pageNumber="548" rank="species">sp. nov.</taxonomicNameLabel>
Begha &amp; Souza
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<paragraph id="8BC236BBFFADFF98FF2AFD58E96411DA" blockId="6.[151,1194,618,717]" box="[151,313,691,717]" pageId="6" pageNumber="548">
(
<figureCitation id="13462A3EFFADFF98FF22FD58E8B511DA" box="[159,232,691,717]" captionStart="FIGURE 2" captionStartId="5.[151,250,1587,1611]" captionTargetBox="[165,1423,725,1564]" captionTargetId="figure-169@5.[165,1423,725,1564]" captionTargetPageId="5" captionText="FIGURE 2. Eustrophopsis bilunulata Pic, 1912, male, sampled in Malaise trap in the Parque Estadual Altamiro de Moura Pacheco, Goianápolis, Goiás, Brazil. (A) Dorsal view; (B) Ventral view." figureDoi="http://doi.org/10.5281/zenodo.13745419" httpUri="https://zenodo.org/record/13745419/files/figure.png" pageId="6" pageNumber="548">Figs 2</figureCitation>
,
<figureCitation id="13462A3EFFADFF98FF49FD58E95E11DA" box="[244,259,691,717]" captionStart="FIGURE 3" captionStartId="6.[152,255,1929,1953]" captionTargetBox="[220,1357,921,1874]" captionTargetId="figure-265@6.[202,1386,921,1906]" captionTargetPageId="6" captionText="FIGURE 3. Eustrophopsis laurae sp. nov., female, holotype sampled in Malaise trap in the Parque Estadual Altamiro de Moura Pacheco, Goianápolis, Goiás, Brazil. (A) Dorsal view; (B) Ventral view." figureDoi="http://doi.org/10.5281/zenodo.13745421" httpUri="https://zenodo.org/record/13745421/files/figure.png" pageId="6" pageNumber="548">3</figureCitation>
,
<figureCitation id="13462A3EFFADFF98FEADFD58E96F11DA" box="[272,306,691,717]" captionStart="FIGURE 4" captionStartId="7.[152,255,1512,1536]" captionTargetBox="[341,1245,185,1475]" captionTargetId="figure-29@7.[341,1245,181,1488]" captionTargetPageId="7" captionText="FIGURE 4. Illustrations of genitalia of Eustrophopsis species. (4) Median lobe of E. bilunulata Pic, 1912; (4) Ovipositor of E. laurae sp. nov. CXT, coxites; STL, stylus; VLV, vulva." figureDoi="http://doi.org/10.5281/zenodo.13745423" httpUri="https://zenodo.org/record/13745423/files/figure.png" pageId="6" pageNumber="548">4B</figureCitation>
)
</paragraph>
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<subSubSection id="C3676530FFADFF96FF2AFD11E88912C6" lastPageId="8" lastPageNumber="550" pageId="6" pageNumber="548" type="diagnosis">
<paragraph id="8BC236BBFFADFF98FF2AFD11E9EF1096" blockId="6.[151,1437,762,897]" pageId="6" pageNumber="548">
<emphasis id="B909EAA9FFADFF98FF2AFD11E9491003" bold="true" box="[151,276,762,788]" pageId="6" pageNumber="548">Diagnosis:</emphasis>
Body length of nearly one centimeter. Overall body honey colored, edges of pronotum and elytra with slightly lighter tone in dorsal view, pronotum with well-defined lighter edge and elytropleura also yellow. Body completely covered by many fine setae. Eyes large, nearly contiguous dorsally. Prosternal truncated, nearly reaching the apex of the procoxae.
</paragraph>
<caption id="DF026633FFADFF98FF25F862EB2F14D2" ID-DOI="http://doi.org/10.5281/zenodo.13745421" ID-Zenodo-Dep="13745421" httpUri="https://zenodo.org/record/13745421/files/figure.png" pageId="6" pageNumber="548" startId="6.[152,255,1929,1953]" targetBox="[220,1357,921,1874]" targetPageId="6" targetType="figure">
<paragraph id="8BC236BBFFADFF98FF25F862EB2F14D2" blockId="6.[151,1437,1929,1989]" pageId="6" pageNumber="548">
<emphasis id="B909EAA9FFADFF98FF25F862E94A14B6" bold="true" box="[152,279,1929,1953]" pageId="6" pageNumber="548">FIGURE 3.</emphasis>
<taxonomicName id="4C7D4D38FFADFF98FEA1F862E9A914B6" authorityName="Begha &amp; Souza &amp; Oliveira" authorityYear="2024" box="[284,500,1929,1953]" class="Insecta" family="Tetratomidae" genus="Eustrophopsis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="548" phylum="Arthropoda" rank="species" species="laurae" status="sp. nov.">
<emphasis id="B909EAA9FFADFF98FEA1F862E9A914B6" box="[284,500,1929,1953]" italics="true" pageId="6" pageNumber="548">Eustrophopsis laurae</emphasis>
</taxonomicName>
<emphasis id="B909EAA9FFADFF98FE44F862EA1514B6" bold="true" box="[505,584,1929,1953]" pageId="6" pageNumber="548">
<taxonomicNameLabel id="A23A57D2FFADFF98FE44F862EA1514B6" box="[505,584,1929,1953]" pageId="6" pageNumber="548" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female, holotype sampled in Malaise trap in the Parque Estadual Altamiro de Moura Pacheco, Goianápolis, Goiás, Brazil.
<emphasis id="B909EAA9FFADFF98FDAFF846EA6814D2" bold="true" box="[530,565,1965,1989]" pageId="6" pageNumber="548">(A)</emphasis>
Dorsal view;
<emphasis id="B909EAA9FFADFF98FD79F846EABB14D2" bold="true" box="[708,742,1965,1989]" pageId="6" pageNumber="548">(B)</emphasis>
Ventral view.
</paragraph>
</caption>
<caption id="DF026633FFACFF99FF25FA03EA9B1533" ID-DOI="http://doi.org/10.5281/zenodo.13745423" ID-Zenodo-Dep="13745423" httpUri="https://zenodo.org/record/13745423/files/figure.png" pageId="7" pageNumber="549" startId="7.[152,255,1512,1536]" targetBox="[341,1245,185,1475]" targetPageId="7" targetType="figure">
<paragraph id="8BC236BBFFACFF99FF25FA03EA9B1533" blockId="7.[151,1437,1512,1572]" pageId="7" pageNumber="549">
<emphasis id="B909EAA9FFACFF99FF25FA03E9451517" bold="true" box="[152,280,1512,1536]" pageId="7" pageNumber="549">FIGURE 4.</emphasis>
Illustrations of genitalia of
<taxonomicName id="4C7D4D38FFACFF99FD8DFA03EB4A1517" box="[560,791,1512,1536]" class="Insecta" family="Tetratomidae" genus="Eustrophopsis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="549" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B909EAA9FFACFF99FD8DFA03EA9D1517" box="[560,704,1512,1536]" italics="true" pageId="7" pageNumber="549">Eustrophopsis</emphasis>
species.
</taxonomicName>
<emphasis id="B909EAA9FFACFF99FCA1FA03EB641517" bold="true" box="[796,825,1512,1536]" pageId="7" pageNumber="549">(4)</emphasis>
Median lobe of
<taxonomicName id="4C7D4D38FFACFF99FC62FA02EC911517" authority="Pic, 1912" authorityName="Pic" authorityYear="1912" box="[991,1228,1512,1536]" class="Insecta" family="Tetratomidae" genus="Eustrophopsis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="549" phylum="Arthropoda" rank="species" species="bilunulata">
<emphasis id="B909EAA9FFACFF99FC62FA02EC3E1517" box="[991,1123,1512,1536]" italics="true" pageId="7" pageNumber="549">E. bilunulata</emphasis>
<bibRefCitation id="EFEC4B4AFFACFF99FBD4FA03EC911517" author="Pic, M." box="[1129,1228,1512,1536]" pageId="7" pageNumber="549" pagination="93 - 94" refId="ref5867" refString="Pic, M. (1912) Coleopteres exotiques nouveaux ou peu connus. Echange, 28, 93 - 94." type="journal article" year="1912">Pic, 1912</bibRefCitation>
</taxonomicName>
;
<emphasis id="B909EAA9FFACFF99FB6BFA03ECAE1517" bold="true" box="[1238,1267,1512,1536]" pageId="7" pageNumber="549">(4)</emphasis>
Ovipositor of
<taxonomicName id="4C7D4D38FFACFF99FA3BFA02E8841533" authorityName="Begha &amp; Souza &amp; Oliveira" authorityYear="2024" class="Insecta" family="Tetratomidae" genus="Eustrophopsis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="549" phylum="Arthropoda" rank="species" species="laurae" status="sp. nov.">
<emphasis id="B909EAA9FFACFF99FA3BFA02E8841533" italics="true" pageId="7" pageNumber="549">E. laurae</emphasis>
</taxonomicName>
<emphasis id="B909EAA9FFACFF99FF5DF9E7E96D1533" bold="true" box="[224,304,1548,1572]" pageId="7" pageNumber="549">
<taxonomicNameLabel id="A23A57D2FFACFF99FF5DF9E7E96D1533" box="[224,304,1548,1572]" pageId="7" pageNumber="549" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
CXT, coxites; STL, stylus; VLV, vulva.
</paragraph>
</caption>
<paragraph id="8BC236BBFFACFF99FF7AF99DEB4B1587" blockId="7.[151,1437,1654,2041]" box="[199,790,1654,1681]" pageId="7" pageNumber="549">
<emphasis id="B909EAA9FFACFF99FF7AF99DE97B1587" bold="true" box="[199,294,1654,1680]" pageId="7" pageNumber="549">Length:</emphasis>
8.939.87 mm (based on three individuals).
</paragraph>
<paragraph id="8BC236BBFFACFF99FF7AF971EC311436" blockId="7.[151,1437,1654,2041]" pageId="7" pageNumber="549">
<emphasis id="B909EAA9FFACFF99FF7AF971E94D15A3" bold="true" box="[199,272,1690,1716]" pageId="7" pageNumber="549">Head:</emphasis>
testaceous. Eyes large, covering most of the head (ocular index 3,902), deeply emarginated around antennal insertions, nearly contiguous dorsally; antennomeres IIV distinctly narrow, antennomeres VX distinctly widened and ventrally flattened; with annular antennal sensilla around entire circumference of antennomeres; clypeus inflated, uniform punctuation; labrum rounded; region of the gena light orange.
</paragraph>
<paragraph id="8BC236BBFFACFF96FF7AF8C1EA4F120A" blockId="7.[151,1437,1654,2041]" lastBlockId="8.[151,1437,151,825]" lastPageId="8" lastPageNumber="550" pageId="7" pageNumber="549">
<emphasis id="B909EAA9FFACFF99FF7AF8C1E9161453" bold="true" box="[199,331,1834,1860]" pageId="7" pageNumber="549">Prothorax:</emphasis>
pronotum with testaceous disc and lighter margins; prosternum black to dark brown, outer margins of the pronotum light orange posterior margin of the pronotum bisinuated, median portion deeply sinuated, rounded; prothoracic episterna with distinct, transverse suture; prosternal process truncated, nearly reaching the apex of the procoxae. Prothoracic episterna each with distinct, transverse suture.
<emphasis id="B909EAA9FFACFF99FC51F87DED0814A7" bold="true" box="[1004,1365,1942,1968]" pageId="7" pageNumber="549">Mesothorax and Metathorax:</emphasis>
black to dark brown; mesosternum short and keeled; metasternum with well-marked median suture.
<emphasis id="B909EAA9FFACFF99FB76F851ED7D14C3" bold="true" box="[1227,1312,1978,2004]" pageId="7" pageNumber="549">Elytra:</emphasis>
testaceous with lighter margins; elytropleura light orange, wider at the humeral region and tapering towards the apex; elytra with rows of punctures.
<emphasis id="B909EAA9FFA3FF96FE1AFF7CE9B513A6" bold="true" box="[423,488,151,177]" pageId="8" pageNumber="550">Legs:</emphasis>
all legs similar; metathoracic legs longer than meso- or prothoracic legs; outer faces of meso- and metatibiae with numerous oblique, comb-like ridges; tarsal formula 5-5-4, all tarsi cylindrical, with first tarsomere being the longest, the other subequal in length; metatarsomere I subequal in length to the other four metatarsomeres; all claws simple.
</paragraph>
<paragraph id="8BC236BBFFA3FF96FF7AFECDE88912C6" blockId="8.[151,1437,151,825]" pageId="8" pageNumber="550">
<emphasis id="B909EAA9FFA3FF96FF7AFECDE9181257" bold="true" box="[199,325,294,320]" pageId="8" pageNumber="550">Abdomen:</emphasis>
ventrite I dark brown to black, median portion testaceous; ventrites IIIV curved, testaceous with inner margin dark brown, ventrite V testaceous.
<emphasis id="B909EAA9FFA3FF96FD09FEA0EC5F1273" bold="true" box="[692,1026,330,357]" pageId="8" pageNumber="550">
Female ovipositor (
<figureCitation id="13462A3EFFA3FF96FC24FEA1EBA91273" box="[921,1012,330,356]" captionStart="FIGURE 4" captionStartId="7.[152,255,1512,1536]" captionTargetBox="[341,1245,185,1475]" captionTargetId="figure-29@7.[341,1245,181,1488]" captionTargetPageId="7" captionText="FIGURE 4. Illustrations of genitalia of Eustrophopsis species. (4) Median lobe of E. bilunulata Pic, 1912; (4) Ovipositor of E. laurae sp. nov. CXT, coxites; STL, stylus; VLV, vulva." figureDoi="http://doi.org/10.5281/zenodo.13745423" httpUri="https://zenodo.org/record/13745423/files/figure.png" pageId="8" pageNumber="550">Fig. 4B</figureCitation>
):
</emphasis>
membranous, gonocoxites parallels, inner portions with spike-like projection, with apex bearing many fine setae; apical portion of vulva tapered, sclerotized portion parallel, spike-like; stylus with rounded apex slightly wider than base, bearing fine setae at apex.
</paragraph>
</subSubSection>
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, GO,
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. (
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<typeStatus id="54C68819FFA3FF96FF7AFDACE9691176" box="[199,308,583,609]" pageId="8" pageNumber="550" type="paratype">Paratype</typeStatus>
material:
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, GO,
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,
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, Malaise Mata—
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Trilha
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-
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<collectingDate id="EF87E993FFA3FF96FB67FD64ED0A11BE" box="[1242,1367,655,681]" pageId="8" pageNumber="550" value="2023-03-21">21.iii.2023</collectingDate>
</date>
;
<collectorName id="2688536DFFA3FF96FADFFD64E97311DA" pageId="8" pageNumber="550">Cols. Oliveira, S.S.</collectorName>
e equipe cols. (
<collectionCode id="ED6CAE7EFFA3FF96FE60FD58EA7011DA" box="[477,557,691,717]" pageId="8" pageNumber="550">ZUFG</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3676530FFA3FF96FF7AFD3CEAF01002" pageId="8" pageNumber="550" type="etymology">
<paragraph id="8BC236BBFFA3FF96FF7AFD3CEAF01002" blockId="8.[151,1437,151,825]" pageId="8" pageNumber="550">
<emphasis id="B909EAA9FFA3FF96FF7AFD3CE90F11E6" bold="true" box="[199,338,727,753]" pageId="8" pageNumber="550">Etymology:</emphasis>
the specific epithet is a patronym honoring Dr. Laura Rocha Prado, for her important contributions to the taxonomy and systematics of
<taxonomicName id="4C7D4D38FFA3FF96FD97FD10EAF41002" authorityName="Linnaeus" authorityYear="1758" box="[554,681,763,789]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="8" pageNumber="550" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3676530FFA3FF96FF7AFCF5ECE1102E" box="[199,1212,798,825]" pageId="8" pageNumber="550" type="distribution">
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<emphasis id="B909EAA9FFA3FF96FF7AFCF5E93E102F" bold="true" box="[199,355,798,824]" pageId="8" pageNumber="550">Distribution:</emphasis>
<collectingCountry id="F36A762BFFA3FF96FED7FCF4E9EC102E" box="[362,433,799,825]" name="Brazil" pageId="8" pageNumber="550">Brazil</collectingCountry>
,
<collectingRegion id="49B9F859FFA3FF96FE01FCF4EA5D102E" box="[444,512,799,825]" country="Brazil" name="Goias" pageId="8" pageNumber="550">Goiás</collectingRegion>
, Goianópolis, Parque Altamiro de Moura Pacheco (this work).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,257 @@
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<mods:title id="DEB2AE2855D8F900A2CDF7B5ECBE91AB">Polypore fungus beetles (Coleoptera: Tetratomidae) in a remnant of semi-deciduous forest of the Brazilian Cerrado: checklist and the description of a new species of Eustrophopsis Champion, 1889</mods:title>
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<treatment id="03D487ADFFAEFF98FF2AFE6CEA24110E" LSID="urn:lsid:plazi:treatment:03D487ADFFAEFF98FF2AFE6CEA24110E" httpUri="http://treatment.plazi.org/id/03D487ADFFAEFF98FF2AFE6CEA24110E" lastPageId="6" lastPageNumber="548" pageId="5" pageNumber="547">
<subSubSection id="C3676530FFAEFF9BFF2AFE6CEA7012B5" box="[151,557,391,418]" pageId="5" pageNumber="547" type="nomenclature">
<paragraph id="8BC236BBFFAEFF9BFF2AFE6CEA7012B5" blockId="5.[151,557,391,454]" box="[151,557,391,418]" pageId="5" pageNumber="547">
<heading id="D08A81D7FFAEFF9BFF2AFE6CEA7012B5" bold="true" box="[151,557,391,418]" fontSize="11" level="1" pageId="5" pageNumber="547" reason="1">
<taxonomicName id="4C7D4D38FFAEFF9BFF2AFE6CEA7012B5" authority="Pic, 1912" authorityName="Pic" authorityYear="1912" box="[151,557,391,418]" class="Insecta" family="Tetratomidae" genus="Eustrophopsis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="547" phylum="Arthropoda" rank="species" species="bilunulata">
<emphasis id="B909EAA9FFAEFF9BFF2AFE6CEA7012B5" bold="true" box="[151,557,391,418]" pageId="5" pageNumber="547">
<emphasis id="B909EAA9FFAEFF9BFF2AFE6CE9E612B5" bold="true" box="[151,443,391,418]" italics="true" pageId="5" pageNumber="547">Eustrophopsis bilunulata</emphasis>
<bibRefCitation id="EFEC4B4AFFAEFF9BFE7FFE63EA7012B5" author="Pic, M." box="[450,557,392,418]" pageId="5" pageNumber="547" pagination="93 - 94" refId="ref5867" refString="Pic, M. (1912) Coleopteres exotiques nouveaux ou peu connus. Echange, 28, 93 - 94." type="journal article" year="1912">Pic, 1912</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3676530FFAEFF9BFF2AFE47E94212D1" box="[151,287,428,454]" pageId="5" pageNumber="547" type="description">
<paragraph id="8BC236BBFFAEFF9BFF2AFE47E94212D1" blockId="5.[151,557,391,454]" box="[151,287,428,454]" pageId="5" pageNumber="547">
(
<figureCitation id="13462A3EFFAEFF9BFF22FE47E8B512D1" box="[159,232,428,454]" captionStart="FIGURE 2" captionStartId="5.[151,250,1587,1611]" captionTargetBox="[165,1423,725,1564]" captionTargetId="figure-169@5.[165,1423,725,1564]" captionTargetPageId="5" captionText="FIGURE 2. Eustrophopsis bilunulata Pic, 1912, male, sampled in Malaise trap in the Parque Estadual Altamiro de Moura Pacheco, Goianápolis, Goiás, Brazil. (A) Dorsal view; (B) Ventral view." figureDoi="http://doi.org/10.5281/zenodo.13745419" httpUri="https://zenodo.org/record/13745419/files/figure.png" pageId="5" pageNumber="547">Figs 2</figureCitation>
,
<figureCitation id="13462A3EFFAEFF9BFF49FE47E94A12D1" box="[244,279,428,454]" captionStart="FIGURE 4" captionStartId="7.[152,255,1512,1536]" captionTargetBox="[341,1245,185,1475]" captionTargetId="figure-29@7.[341,1245,181,1488]" captionTargetPageId="7" captionText="FIGURE 4. Illustrations of genitalia of Eustrophopsis species. (4) Median lobe of E. bilunulata Pic, 1912; (4) Ovipositor of E. laurae sp. nov. CXT, coxites; STL, stylus; VLV, vulva." figureDoi="http://doi.org/10.5281/zenodo.13745423" httpUri="https://zenodo.org/record/13745423/files/figure.png" pageId="5" pageNumber="547">4A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3676530FFAEFF9BFF2AFE1BEB5E111E" box="[151,771,496,521]" pageId="5" pageNumber="547" type="reference_group">
<paragraph id="8BC236BBFFAEFF9BFF2AFE1BEB5E111E" blockId="5.[151,771,496,521]" box="[151,771,496,521]" pageId="5" pageNumber="547">
<treatmentCitationGroup id="AB6D1195FFAEFF9BFF2AFE1BEB5E111E" box="[151,771,496,521]" pageId="5" pageNumber="547">
<treatmentCitation id="0ADC10AAFFAEFF9BFF2AFE1BEA72111E" author="Pic, M." box="[151,559,496,521]" page="93" pageId="5" pageNumber="547" year="1912">
<taxonomicName id="4C7D4D38FFAEFF9BFF2AFE1BEA72111E" authority="Pic, 1912: 93" authorityName="Pic" authorityPageNumber="93" authorityYear="1912" box="[151,559,496,521]" class="Insecta" family="Tetratomidae" genus="Eustrophopsis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="547" phylum="Arthropoda" rank="species" species="bilunulatus">
<emphasis id="B909EAA9FFAEFF9BFF2AFE1BE9FD111F" box="[151,416,496,520]" italics="true" pageId="5" pageNumber="547">Eustrophopsis bilunulatus</emphasis>
<bibRefCitation id="EFEC4B4AFFAEFF9BFE1BFE1BEA72111E" author="Pic, M." box="[422,559,496,521]" pageId="5" pageNumber="547" pagination="93 - 94" refId="ref5867" refString="Pic, M. (1912) Coleopteres exotiques nouveaux ou peu connus. Echange, 28, 93 - 94." type="journal article" year="1912">Pic, 1912: 93</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0ADC10AAFFAEFF9BFD87FE1BEAA2111E" author="Champion, G. C." box="[570,767,496,521]" page="3" pageId="5" pageNumber="547" year="1916">
<bibRefCitation id="EFEC4B4AFFAEFF9BFD87FE1BEAA2111E" author="Champion, G. C." box="[570,767,496,521]" pageId="5" pageNumber="547" pagination="1 - 10" refId="ref4747" refString="Champion, G. C. (1916) Notes on Melandryidae (2). The Entomologist's Monthly Magazine, 52, 1 - 10. https: // doi. org / 10.5962 / bhl. part. 23928" type="journal article" year="1916">Champion, 1916: 3</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3676530FFAEFF9BFF2AFDDDEB0E1492" pageId="5" pageNumber="547" type="diagnosis">
<paragraph id="8BC236BBFFAEFF9BFF2AFDDDEA8F11AA" blockId="5.[151,1437,566,701]" pageId="5" pageNumber="547">
<emphasis id="B909EAA9FFAEFF9BFF2AFDDDE9491147" bold="true" box="[151,276,566,592]" pageId="5" pageNumber="547">Diagnosis:</emphasis>
Body length of nearly half a centimeter. Overall body brown to testaceous, covered by many fine setae. Eyes nearly contiguous dorsally; antennomeres IIV and the tip of antennomere XI yellow, antennomeres VX dark brown; clypeus inflated. Apex of the prosternal process bifurcated. The most noticeable feature is the two black rounded maculae in the upper portion of the elytra.
</paragraph>
<caption id="DF026633FFAEFF9BFF2AF9D8EB2F1578" ID-DOI="http://doi.org/10.5281/zenodo.13745419" ID-Zenodo-Dep="13745419" httpUri="https://zenodo.org/record/13745419/files/figure.png" pageId="5" pageNumber="547" startId="5.[151,250,1587,1611]" targetBox="[165,1423,725,1564]" targetPageId="5" targetType="figure">
<paragraph id="8BC236BBFFAEFF9BFF2AF9D8EB2F1578" blockId="5.[151,1437,1587,1647]" pageId="5" pageNumber="547">
<emphasis id="B909EAA9FFAEFF9BFF2AF9D8E94B155C" bold="true" box="[151,278,1587,1611]" pageId="5" pageNumber="547">FIGURE 2.</emphasis>
<taxonomicName id="4C7D4D38FFAEFF9BFEA2F9D8EAD3155B" authority="Pic, 1912" authorityName="Pic" authorityYear="1912" box="[287,654,1587,1612]" class="Insecta" family="Tetratomidae" genus="Eustrophopsis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="547" phylum="Arthropoda" rank="species" species="bilunulata">
<emphasis id="B909EAA9FFAEFF9BFEA2F9D8EA7D155C" box="[287,544,1587,1611]" italics="true" pageId="5" pageNumber="547">Eustrophopsis bilunulata</emphasis>
<bibRefCitation id="EFEC4B4AFFAEFF9BFD94F9D8EAD3155B" author="Pic, M." box="[553,654,1587,1612]" pageId="5" pageNumber="547" pagination="93 - 94" refId="ref5867" refString="Pic, M. (1912) Coleopteres exotiques nouveaux ou peu connus. Echange, 28, 93 - 94." type="journal article" year="1912">Pic, 1912</bibRefCitation>
</taxonomicName>
, male, sampled in Malaise trap in the Parque Estadual Altamiro de Moura Pacheco, Goianápolis, Goiás, Brazil.
<emphasis id="B909EAA9FFAEFF9BFDAFF9BCEA681578" bold="true" box="[530,565,1623,1647]" pageId="5" pageNumber="547">(A)</emphasis>
Dorsal view;
<emphasis id="B909EAA9FFAEFF9BFD79F9BCEABB1578" bold="true" box="[708,742,1623,1647]" pageId="5" pageNumber="547">(B)</emphasis>
Ventral view.
</paragraph>
</caption>
<paragraph id="8BC236BBFFAEFF9BFF7AF95CEB4B15C6" blockId="5.[151,1437,1719,2033]" box="[199,790,1719,1746]" pageId="5" pageNumber="547">
<emphasis id="B909EAA9FFAEFF9BFF7AF95CE97B15C6" bold="true" box="[199,294,1719,1745]" pageId="5" pageNumber="547">Length:</emphasis>
5.716.30 mm (based on three individuals).
</paragraph>
<paragraph id="8BC236BBFFAEFF9BFF7AF930EA2D140E" blockId="5.[151,1437,1719,2033]" pageId="5" pageNumber="547">
<emphasis id="B909EAA9FFAEFF9BFF7AF930EACC15E2" bold="true" box="[199,657,1755,1781]" pageId="5" pageNumber="547">
Male aedeagus median lobe (
<figureCitation id="13462A3EFFAEFF9BFD99F930EADF15E2" box="[548,642,1755,1781]" captionStart="FIGURE 4" captionStartId="7.[152,255,1512,1536]" captionTargetBox="[341,1245,185,1475]" captionTargetId="figure-29@7.[341,1245,181,1488]" captionTargetPageId="7" captionText="FIGURE 4. Illustrations of genitalia of Eustrophopsis species. (4) Median lobe of E. bilunulata Pic, 1912; (4) Ovipositor of E. laurae sp. nov. CXT, coxites; STL, stylus; VLV, vulva." figureDoi="http://doi.org/10.5281/zenodo.13745423" httpUri="https://zenodo.org/record/13745423/files/figure.png" pageId="5" pageNumber="547">Fig. 4A</figureCitation>
):
</emphasis>
slender, overall shape oval, with apical segment slightly longer than basal segment, with distal apex spear-like,
</paragraph>
<paragraph id="8BC236BBFFAEFF9BFF7AF8C8EB0E1492" blockId="5.[151,1437,1719,2033]" pageId="5" pageNumber="547">
<emphasis id="B909EAA9FFAEFF9BFF7AF8C8E9F6142A" bold="true" box="[199,427,1827,1853]" pageId="5" pageNumber="547">Studied specimens:</emphasis>
<materialsCitation id="3B153CE6FFAEFF9BFE0DF8C8E8911491" collectingDate="2022-06-29" collectingDateMax="2022-07-02" collectingDateMin="2022-06-29" collectionCode="R" collectorName="Cols. Oliveira, S. S. &amp; Lopes, W." country="Brazil" county="Malaise Mata - Trilha do Tamandua" elevation="880" latitude="-16.55325" location="Goianapolis" longLatPrecision="1" longitude="-49.13151" municipality="Parque Altamiro de Moura" pageId="5" pageNumber="547" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<specimenCount id="9D7BFD32FFAEFF9BFE0DF8C8E9861429" box="[432,475,1827,1854]" count="2" pageId="5" pageNumber="547" type="male">2 ♂</specimenCount>
,
<specimenCount id="9D7BFD32FFAEFF9BFE58F8C8EA5B142A" box="[485,518,1827,1853]" count="1" pageId="5" pageNumber="547" type="female">1 ♀</specimenCount>
<collectingCountry id="F36A762BFFAEFF9BFDBBF8C8EA28142A" box="[518,629,1827,1853]" name="Brazil" pageId="5" pageNumber="547">BRAZIL</collectingCountry>
, GO,
<location id="8EA26060FFAEFF9BFD0EF8C8EB60142A" LSID="urn:lsid:plazi:treatment:03D487ADFFAEFF98FF2AFE6CEA24110E:8EA26060FFAEFF9BFD0EF8C8EB60142A" box="[691,829,1827,1853]" country="Brazil" county="Malaise Mata - Trilha do Tamandua" latitude="-16.55325" longLatPrecision="1" longitude="-49.13151" municipality="Parque Altamiro de Moura" name="Goianapolis" pageId="5" pageNumber="547">Goianápolis</location>
,
<collectingMunicipality id="6BA6ACC1FFAEFF9BFCFAF8CFEC331429" box="[839,1134,1827,1854]" pageId="5" pageNumber="547">Parque Altamiro de Moura</collectingMunicipality>
Pacheco-PEAMP;
<collectingCounty id="62A34E37FFAEFF9BFAF9F8C8E9901476" pageId="5" pageNumber="547">Malaise Mata—Trilha do Tamanduá</collectingCounty>
<geoCoordinate id="EE49507CFFAEFF9BFE68F8ACEA081475" box="[469,597,1863,1890]" degrees="16,553250" direction="south" orientation="latitude" pageId="5" pageNumber="547" precision="1" value="-16.55325">-16,553250</geoCoordinate>
<geoCoordinate id="EE49507CFFAEFF9BFDE1F8ACEA801475" box="[604,733,1863,1890]" degrees="49,131511" direction="west" orientation="longitude" pageId="5" pageNumber="547" precision="1" value="-49.13151">-49,131511</geoCoordinate>
<quantity id="4C859B5EFFAEFF9BFD47F8ACEB671475" box="[762,826,1863,1890]" metricMagnitude="2" metricUnit="m" metricValue="8.8" pageId="5" pageNumber="547" unit="m" value="880.0">
<elevation id="0050D188FFAEFF9BFD47F8ACEB671475" box="[762,826,1863,1890]" metricMagnitude="2" metricUnit="m" metricValue="8.8" pageId="5" pageNumber="547" unit="m" value="880.0">880m</elevation>
</quantity>
;
<date id="FFC3107BFFAEFF9BFCFBF8ACEBE21476" box="[838,959,1863,1889]" pageId="5" pageNumber="547" value="2022-06-29">
<collectingDate id="EF87E993FFAEFF9BFCFBF8ACEBE21476" box="[838,959,1863,1889]" pageId="5" pageNumber="547" value="2022-06-29">29.vi.2022</collectingDate>
</date>
-
<date id="FFC3107BFFAEFF9BFC7AF8ACEC171476" box="[967,1098,1863,1889]" pageId="5" pageNumber="547" value="2022-07-02">
<collectingDate id="EF87E993FFAEFF9BFC7AF8ACEC171476" box="[967,1098,1863,1889]" pageId="5" pageNumber="547" value="2022-07-02">02.vii.2022</collectingDate>
</date>
;
<collectorName id="2688536DFFAEFF9BFBE8F8ACED701476" box="[1109,1325,1863,1889]" pageId="5" pageNumber="547">Cols. Oliveira, S.S.</collectorName>
&amp;
<collectorName id="2688536DFFAEFF9BFAECF8A3E8E51491" pageId="5" pageNumber="547">Lopes, W.</collectorName>
<collectionCode id="ED6CAE7EFFAEFF9BFF05F887E8911491" box="[184,204,1900,1926]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="547">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B153CE6FFAEFF9BFF6AF880EA771492" box="[215,554,1899,1925]" collectionCode="MZUSP" country="Brazil" pageId="5" pageNumber="547" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="9D7BFD32FFAEFF9BFF6AF880E8A21492" box="[215,255,1899,1925]" count="1" pageId="5" pageNumber="547" type="male">1 ♂</specimenCount>
and
<specimenCount id="9D7BFD32FFAEFF9BFE8DF880E90F1492" box="[304,338,1899,1925]" count="1" pageId="5" pageNumber="547" type="female">1 ♀</specimenCount>
housed in
<collectionCode id="ED6CAE7EFFAEFF9BFE70F880EA771492" box="[461,554,1899,1925]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="5" pageNumber="547">MZUSP</collectionCode>
</materialsCitation>
, and
<materialsCitation id="3B153CE6FFAEFF9BFDD8F880EB121492" box="[613,847,1899,1925]" collectionCode="ZUFG" country="Brazil" pageId="5" pageNumber="547" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D7BFD32FFAEFF9BFDD8F880EAD11492" box="[613,652,1899,1925]" count="1" pageId="5" pageNumber="547" type="male">1 ♂</specimenCount>
housed in
<collectionCode id="ED6CAE7EFFAEFF9BFCBFF880EB121492" box="[770,847,1899,1925]" pageId="5" pageNumber="547">ZUFG</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3676530FFAEFF98FF7AF864EB3B12C6" lastPageId="6" lastPageNumber="548" pageId="5" pageNumber="547" type="discussion">
<paragraph id="8BC236BBFFAEFF98FF7AF864EB3B12C6" blockId="5.[151,1437,1719,2033]" lastBlockId="6.[151,1437,151,537]" lastPageId="6" lastPageNumber="548" pageId="5" pageNumber="547">
<emphasis id="B909EAA9FFAEFF9BFF7AF864E96014BE" bold="true" box="[199,317,1935,1961]" pageId="5" pageNumber="547">Remarks:</emphasis>
No photos or illustrations have previously been made for this species. This species appears very similar to
<taxonomicName id="4C7D4D38FFAEFF9BFEBAF85FEA7214DA" authority="(Pic, 1912)" baseAuthorityName="Pic" baseAuthorityYear="1912" box="[263,559,1971,1997]" class="Insecta" family="Tetratomidae" genus="Eustrophopsis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="547" phylum="Arthropoda" rank="species" species="bimaculatus">
<emphasis id="B909EAA9FFAEFF9BFEBAF85FE9F314DA" box="[263,430,1971,1997]" italics="true" pageId="5" pageNumber="547">E. bimaculatus</emphasis>
(
<bibRefCitation id="EFEC4B4AFFAEFF9BFE01F858EA7A14DA" author="Pic, M." box="[444,551,1971,1997]" pageId="5" pageNumber="547" pagination="93 - 94" refId="ref5867" refString="Pic, M. (1912) Coleopteres exotiques nouveaux ou peu connus. Echange, 28, 93 - 94." type="journal article" year="1912">Pic, 1912</bibRefCitation>
)
</taxonomicName>
due to the two black elytral maculae. The main subtle difference is the position of the black maculae, the maculae in
<taxonomicName id="4C7D4D38FFAEFF9BFDF9F833EAB314E6" baseAuthorityName="Pic" baseAuthorityYear="1912" box="[580,750,2007,2033]" class="Insecta" family="Tetratomidae" genus="Eustrophopsis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="547" phylum="Arthropoda" rank="species" species="bimaculatus">
<emphasis id="B909EAA9FFAEFF9BFDF9F833EAB314E6" box="[580,750,2007,2033]" italics="true" pageId="5" pageNumber="547">E. bimaculatus</emphasis>
</taxonomicName>
is placed in the median portion of the elytron, while that of
<taxonomicName id="4C7D4D38FFADFF98FF2AFF7CE97713A6" authorityName="Pic" authorityYear="1912" box="[151,298,151,177]" class="Insecta" family="Tetratomidae" genus="Eustrophopsis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="548" phylum="Arthropoda" rank="species" species="bilunulata">
<emphasis id="B909EAA9FFADFF98FF2AFF7CE97713A6" box="[151,298,151,177]" italics="true" pageId="6" pageNumber="548">E. bilunulata</emphasis>
</taxonomicName>
is placed in the upper portion of the elytron, which is pointed out in the original description. Another difference is the geographical occurrence,
<taxonomicName id="4C7D4D38FFADFF98FDC7FF50EB5313C2" authorityName="Pic" authorityYear="1912" box="[634,782,187,213]" class="Insecta" family="Tetratomidae" genus="Eustrophopsis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="548" phylum="Arthropoda" rank="species" species="bilunulata">
<emphasis id="B909EAA9FFADFF98FDC7FF50EB5313C2" box="[634,782,187,213]" italics="true" pageId="6" pageNumber="548">E. bilunulata</emphasis>
</taxonomicName>
was first described from Jataí,
<collectingRegion id="49B9F859FFADFF98FBCBFF50ECE713C2" box="[1142,1210,187,213]" country="Brazil" name="Goias" pageId="6" pageNumber="548">Goiás</collectingRegion>
, within the Cerrado (
<bibRefCitation id="EFEC4B4AFFADFF98FF22FF34E95213EE" author="Pic, M." box="[159,271,223,249]" pageId="6" pageNumber="548" pagination="93 - 94" refId="ref5867" refString="Pic, M. (1912) Coleopteres exotiques nouveaux ou peu connus. Echange, 28, 93 - 94." type="journal article" year="1912">Pic, 1912</bibRefCitation>
), while
<taxonomicName id="4C7D4D38FFADFF98FED7FF34EA7C13EE" box="[362,545,223,249]" class="Insecta" family="Tetratomidae" genus="Eustrophopsis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="548" phylum="Arthropoda" rank="species" species="bimaculataus">
<emphasis id="B909EAA9FFADFF98FED7FF34EA7C13EE" box="[362,545,223,249]" italics="true" pageId="6" pageNumber="548">E. bimaculataus</emphasis>
</taxonomicName>
from Itatiaia,
<collectingRegion id="49B9F859FFADFF98FD78FF34EB3113EE" box="[709,876,223,249]" country="Brazil" name="Rio de Janeiro" pageId="6" pageNumber="548">Rio de Janeiro</collectingRegion>
, and there are also records for
<collectingRegion id="49B9F859FFADFF98FB76FF34ED3113EE" box="[1227,1388,223,249]" country="Brazil" name="Espirito Santo" pageId="6" pageNumber="548">Espírito Santo</collectingRegion>
and
<collectingRegion id="49B9F859FFADFF98FF2AFEE8E91C120A" box="[151,321,259,285]" country="Brazil" name="Santa Catarina" pageId="6" pageNumber="548">Santa Catarina</collectingRegion>
, all within the Atlantic Forest area (
<bibRefCitation id="EFEC4B4AFFADFF98FD5EFEE8EB08120A" author="Pic, M." box="[739,853,259,285]" pageId="6" pageNumber="548" pagination="93 - 94" refId="ref5867" refString="Pic, M. (1912) Coleopteres exotiques nouveaux ou peu connus. Echange, 28, 93 - 94." type="journal article" year="1912">Pic, 1912</bibRefCitation>
). Other more recent records are also known for
<taxonomicName id="4C7D4D38FFADFF98FA39FEE8E97D1256" baseAuthorityName="Pic" baseAuthorityYear="1912" class="Insecta" family="Tetratomidae" genus="Eustrophopsis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="548" phylum="Arthropoda" rank="species" species="bimaculatus">
<emphasis id="B909EAA9FFADFF98FA39FEE8E97D1256" italics="true" pageId="6" pageNumber="548">E. bimaculatus</emphasis>
</taxonomicName>
in the same biome (iNaturalist 2024). Whether these are just geographical variations of the same species, or two truly distinct taxa remains to be seen, as a proper dissection of both male and female, as well as a complete
<typeStatus id="54C68819FFADFF98FEBBFE84E96B129E" box="[262,310,367,393]" pageId="6" pageNumber="548">type</typeStatus>
revision, would be required. Furthermore,
<taxonomicName id="4C7D4D38FFADFF98FCA2FE84EBEF129E" authorityName="Pic" authorityYear="1912" box="[799,946,367,393]" class="Insecta" family="Tetratomidae" genus="Eustrophopsis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="548" phylum="Arthropoda" rank="species" species="bilunulata">
<emphasis id="B909EAA9FFADFF98FCA2FE84EBEF129E" box="[799,946,367,393]" italics="true" pageId="6" pageNumber="548">E. bilunulata</emphasis>
</taxonomicName>
has a record for the Amazon Rainforest, in Tefé,
<collectingRegion id="49B9F859FFADFF98FF6BFE78E91112BA" box="[214,332,403,429]" country="Peru" name="Amazonas" pageId="6" pageNumber="548">Amazonas</collectingRegion>
(
<bibRefCitation id="EFEC4B4AFFADFF98FEE6FE78EA4112BA" author="Champion, G. C." box="[347,540,403,429]" pageId="6" pageNumber="548" pagination="1 - 10" refId="ref4747" refString="Champion, G. C. (1916) Notes on Melandryidae (2). The Entomologist's Monthly Magazine, 52, 1 - 10. https: // doi. org / 10.5962 / bhl. part. 23928" type="journal article" year="1916">Champion, 1916</bibRefCitation>
), but given the distance and dissimilarity of the environment, we are not sure if this is truly a record for
<taxonomicName id="4C7D4D38FFADFF98FE1BFE5CEA6512C6" authorityName="Pic" authorityYear="1912" box="[422,568,439,465]" class="Insecta" family="Tetratomidae" genus="Eustrophopsis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="548" phylum="Arthropoda" rank="species" species="bilunulata">
<emphasis id="B909EAA9FFADFF98FE1BFE5CEA6512C6" box="[422,568,439,465]" italics="true" pageId="6" pageNumber="548">E. bilunulata</emphasis>
</taxonomicName>
or another similar species.
</paragraph>
</subSubSection>
<subSubSection id="C3676530FFADFF98FF7AFE31EA24110E" pageId="6" pageNumber="548" type="distribution">
<paragraph id="8BC236BBFFADFF98FF7AFE31EA24110E" blockId="6.[151,1437,151,537]" pageId="6" pageNumber="548">
<emphasis id="B909EAA9FFADFF98FF7AFE31E93E12E3" bold="true" box="[199,355,474,500]" pageId="6" pageNumber="548">Distribution:</emphasis>
<collectingCountry id="F36A762BFFADFF98FED5FE30E9F212E2" box="[360,431,475,501]" name="Brazil" pageId="6" pageNumber="548">Brazil</collectingCountry>
,
<collectingRegion id="49B9F859FFADFF98FE04FE30E9A012E2" box="[441,509,475,501]" country="Brazil" name="Goias" pageId="6" pageNumber="548">Goiás</collectingRegion>
, Goianópolis, Parque Altamiro de Moura Pacheco (this work), and Jataí (
<bibRefCitation id="EFEC4B4AFFADFF98FA9FFE30EDCD12E2" author="Pic, M." box="[1314,1424,475,501]" pageId="6" pageNumber="548" pagination="93 - 94" refId="ref5867" refString="Pic, M. (1912) Coleopteres exotiques nouveaux ou peu connus. Echange, 28, 93 - 94." type="journal article" year="1912">Pic, 1912</bibRefCitation>
).
<collectingCountry id="F36A762BFFADFF98FF2AFE14E883110E" box="[151,222,511,537]" name="Brazil" pageId="6" pageNumber="548">Brazil</collectingCountry>
, Amazonas, Tefé (
<bibRefCitation id="EFEC4B4AFFADFF98FE11FE14EA31110E" author="Champion, G. C." box="[428,620,511,537]" pageId="6" pageNumber="548" pagination="1 - 10" refId="ref4747" refString="Champion, G. C. (1916) Notes on Melandryidae (2). The Entomologist's Monthly Magazine, 52, 1 - 10. https: // doi. org / 10.5962 / bhl. part. 23928" type="journal article" year="1916">Champion, 1916</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,111 @@
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<emphasis id="B909EAA9FFAFFF9AFF2AF9C8E99A1529" bold="true" box="[151,455,1571,1598]" italics="true" pageId="4" pageNumber="546">Eustrophopsis subtestacea</emphasis>
(
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)
</emphasis>
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</paragraph>
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<emphasis id="B909EAA9FFAFFF9AFF2AF983E9C71597" box="[151,410,1640,1664]" italics="true" pageId="4" pageNumber="546">Eustrophinus subtestacea</emphasis>
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.
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</paragraph>
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<taxonomicName id="4C7D4D38FFAFFF9AFF2AF96CE9BE1588" authority="Refs." authorityName="Refs." box="[151,483,1671,1695]" class="Insecta" family="Tetratomidae" genus="Eustrophopsis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="546" phylum="Arthropoda" rank="species" species="subtestacea">
<emphasis id="B909EAA9FFAFFF9AFF2AF96CE9F91588" box="[151,420,1671,1695]" italics="true" pageId="4" pageNumber="546">Eustrophopsis subtestacea</emphasis>
Refs.
</taxonomicName>
:
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</treatmentCitation>
.
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Distr.:
<collectingCountry id="F36A762BFFAFFF9AFF5EF92CE97A15F6" box="[227,295,1735,1761]" name="Brazil" pageId="4" pageNumber="546">Brazil</collectingCountry>
—state of
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<emphasis id="B909EAA9FFAFFF9AFF2AFA9CE92D1699" box="[151,368,1399,1423]" italics="true" pageId="4" pageNumber="546">Eustrophopsis striata</emphasis>
<bibRefCitation id="EFEC4B4AFFAFFF9AFECBFA9CEA361699" author="Blackwelder, R. E." box="[374,619,1398,1423]" pageId="4" pageNumber="546" pagination="551 - 763" refId="ref4580" refString="Blackwelder, R. E. (1945) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Bulletin of the United States National Museum, 185, 551 - 763. https: // doi. org / 10.5479 / si. 03629236.185.3" type="journal article" year="1945">Blackwelder, 1945: 495</bibRefCitation>
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;
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<bibRefCitation id="EFEC4B4AFFAFFF9AFDC8FA9CEB671699" author="Champion, G. C." box="[629,826,1398,1423]" pageId="4" pageNumber="546" pagination="1 - 10" refId="ref4747" refString="Champion, G. C. (1916) Notes on Melandryidae (2). The Entomologist's Monthly Magazine, 52, 1 - 10. https: // doi. org / 10.5962 / bhl. part. 23928" type="journal article" year="1916">Champion, 1916: 5</bibRefCitation>
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Distr.:
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—state of
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Diagnosis (adapted from
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and
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:
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Eyes large, covering most of the head, deeply emarginated around antennal insertions, nearly contiguous dorsally. Antennomeres IIV distinctly narrow, antennomeres VX distinctly widened and flattened, with annular antennal sensilla around the entire circumference of antennomeres. Sexually dimorphic antennomeres, males with distinctive sensilla on antennomeres VX. Prosternal process separating procoxa. Prothoracic episterna with distinct, transverse suture. Front legs separated by the prothoracic process. Outer faces of meso- and metatibiae with numerous oblique, comb-like ridges. Tarsal formula 5-5-4, with the first tarsomere being the longest, the other subequal in length; metatarsomere I subequal in length to the other four metatarsomeres.
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<mods:title id="A6E7D6FFFE6FE2AA89870095E693503E">Contribution to the genus Clypeocaenis Soldán, 1978 (Ephemeroptera: Caenidae) from Lao Peoples Democratic Republic with a key to Oriental species</mods:title>
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<mods:namePart id="3F14FD30C1F2EE01C4BE9E004293A6F1">Vannachak, Viengkhone</mods:namePart>
<mods:affiliation id="08E28E4D2A9A46953E2CA92863AAC62C">Department of Biology, Faculty of Science, Khon Kaen University, Khon Kaen 40002, Thailand</mods:affiliation>
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<mods:namePart id="15A890CAC95C13BBBFFEC7CFF7D6754B">Malzacher, Peter</mods:namePart>
<mods:affiliation id="8183F460DD1B8251E261CCC5ED472065">Friedrich-Ebert-Strasse 63, 71638 Ludwigsburg, Germany</mods:affiliation>
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<mods:namePart id="951EF01779414A31D64149394F37AEF3">Sangpradub, Narumon</mods:namePart>
<mods:affiliation id="F6451ACB28F495D4DA1EB93930A2D0A9">Department of Biology, Faculty of Science, Khon Kaen University, Khon Kaen 40002, Thailand &amp; Center of Excellence in Biodiversity, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:date id="7F3FDBF8E7CA8C5CFBB2F12013A18EF6">2024</mods:date>
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<mods:number id="4FAE5C49437ED060C07E7109EF9ED946">5501</mods:number>
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<taxonomicName id="4C5C9282FFA0FFC2FF6DFF0AFDDEFF53" ID-CoL="5ZCPL" authority="Soldan, 1978" authorityName="Soldan" authorityYear="1978" box="[151,623,150,178]" class="Insecta" family="Caenidae" genus="Clypeocaenis" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="multisetosa">
<emphasis id="B9283513FFA0FFC2FF6DFF0AFE76FF50" bold="true" box="[151,455,151,178]" italics="true" pageId="13" pageNumber="514">Clypeocaenis multisetosa</emphasis>
<bibRefCitation id="EFCD94F0FFA0FFC2FE34FF0BFDDEFF53" author="Soldan, T." box="[462,623,150,177]" pageId="13" pageNumber="514" pagination="119 - 129" refId="ref10571" refString="Soldan, T. (1978) New genera and species of Caenidae (Ephemeroptera) from Iran, India and Australia. Acta Entomologica Bohemoslovaca, 75, 119 - 129." type="journal article" year="1978">Soldán, 1978</bibRefCitation>
</taxonomicName>
(
<figureCitation id="1367F584FFA0FFC2FD84FF0AFD4DFF53" box="[638,764,151,177]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStart-2="FIGURE 10" captionStart-3="FIGURE 11" captionStartId-0="12.[151,250,1776,1800]" captionStartId-1="14.[151,250,1776,1800]" captionStartId-2="15.[151,251,1467,1491]" captionStartId-3="16.[151,253,1280,1304]" captionTargetBox-0="[158,1430,181,1752]" captionTargetBox-1="[158,1430,179,1750]" captionTargetBox-2="[158,1430,199,1444]" captionTargetBox-3="[151,1435,199,1256]" captionTargetId-0="figure-26@12.[158,1430,181,1752]" captionTargetId-1="figure-26@14.[158,1430,179,1750]" captionTargetId-2="figure-18@15.[158,1430,199,1444]" captionTargetId-3="figure-26@16.[151,1435,199,1256]" captionTargetPageId-0="12" captionTargetPageId-1="14" captionTargetPageId-2="15" captionTargetPageId-3="16" captionText-0="FIGURE 8. Clypeocaenis oligosetosa Soldán, 1983 (A, B) and Clypeocaenis multisetosa Soldán, 1978 (C, D). A, C. High magnification view of microtrichia on the dorsal surface of operculate gills. B, D. Microtrichia band on the ventral side of operculate gills. Scale bars: A = 20 μm; C = 50 μm; B, D = 100 μm." captionText-1="FIGURE 9. Clypeocaenis multisetosa Soldán, 1978, collected from Xe Nou River, Xe Bang Hieng River, and outlet Stream of Nam Xouang Resurvoir in Lao PDR, A. Head capsule. B. Labrum. C. Mandibles. D. Maxillae. E. Hypopharynx. F. Labium. Scale bars: A, D (left maxillary palp) = 500 μm; B, C (right mandible), E = 200 μm; C (left mandible) = 300 μm; D, F (right maxillary palp) = 400 μm." captionText-2="FIGURE 10. Clypeocaenis multisetosa Soldán, 1978, from Xe Nou River, Xe Bang Hieng River, and outlet Stream of Nam Xouang Resurvoir, Lao PDR, A. Forefemur. B. Mid femur. C. Hind femur. D. Foreclaw. E. Mid claw. F. Hind claw. Scale bars: A, B = 500 μm; C = 400 μm; DF = 100 μm." captionText-3="FIGURE 11. Clypeocaenis multisetosa Soldán, 1978, from Xe Nou River, Xe Bang Hieng River, and outlet Stream of Nam Xouang Resurvoir in Lao PDR, A. Abdominal terga IIVII. B. Operculate gill. C, D. Eggs. Scale bars: A, B = 500 μm; C, D = 100 μm." figureDoi-0="http://doi.org/10.5281/zenodo.13745381" figureDoi-1="http://doi.org/10.5281/zenodo.13745385" figureDoi-2="http://doi.org/10.5281/zenodo.13745389" figureDoi-3="http://doi.org/10.5281/zenodo.13745391" httpUri-0="https://zenodo.org/record/13745381/files/figure.png" httpUri-1="https://zenodo.org/record/13745385/files/figure.png" httpUri-2="https://zenodo.org/record/13745389/files/figure.png" httpUri-3="https://zenodo.org/record/13745391/files/figure.png" pageId="13" pageNumber="514">Figs. 811</figureCitation>
)
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<taxonomicName id="4C5C9282FFA0FFC2FF6DFF41FDE8FF16" ID-CoL="5ZCPL" authority="Soldan, 1978: 122" authorityName="Soldan" authorityPageNumber="122" authorityYear="1978" box="[151,601,220,244]" class="Insecta" family="Caenidae" genus="Clypeocaenis" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="multisetosa">
<emphasis id="B9283513FFA0FFC2FF6DFF41FE28FF16" box="[151,409,220,244]" italics="true" pageId="13" pageNumber="514">Clypeocaenis multisetosa</emphasis>
<bibRefCitation id="EFCD94F0FFA0FFC2FE5AFF41FDE8FF16" author="Soldan, T." box="[416,601,220,244]" pageId="13" pageNumber="514" pagination="119 - 129" refId="ref10571" refString="Soldan, T. (1978) New genera and species of Caenidae (Ephemeroptera) from Iran, India and Australia. Acta Entomologica Bohemoslovaca, 75, 119 - 129." type="journal article" year="1978">Soldán, 1978: 122</bibRefCitation>
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(orig. desc., Figs 2024).
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<emphasis id="B9283513FFA0FFC2FF6DFEBCFE3AFEDE" bold="true" box="[151,395,289,316]" italics="true" pageId="13" pageNumber="514">Materials examined</emphasis>
.
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larvae (THNHM-I-0002978000029781).
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, outlet
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larvae (
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PDR,
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,
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,
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a.s.l.
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, deposited in NUoL-AIC
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and
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larvae (
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/
<collectingCountry id="F34BA991FFA0FFC2FC77FE2FFC78FE2E" box="[909,969,434,460]" name="Laos" pageId="13" pageNumber="514">Laos</collectingCountry>
.007)
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,
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,
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,
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a.s.l.
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,
<date id="FFE2CFC1FFA0FFC2FBD6FE4BFB07FE12" box="[1068,1206,470,496]" pageId="13" pageNumber="514" value="2019-11-24">
<collectingDate id="EFA63629FFA0FFC2FBD6FE4BFB07FE12" box="[1068,1206,470,496]" pageId="13" pageNumber="514" value="2019-11-24">24.XI.2019</collectingDate>
</date>
, deposited in KKU-AIC, all
<collectionCode id="ED4D71C4FFA0FFC2FF04FE67FEA5FDF6" box="[254,276,506,532]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="13" pageNumber="514" type="Museum">V</collectionCode>
.
<collectorName id="26A98CD7FFA0FFC2FEE4FE67FE2EFDF6" box="[286,415,506,532]" pageId="13" pageNumber="514">Vannachak</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="8BE3E901FFA0FFC2FF3DFD83FBEFFDBE" blockId="13.[151,1437,289,1504]" pageId="13" pageNumber="514">
<emphasis id="B9283513FFA0FFC2FF3DFD83FEB0FDDA" bold="true" box="[199,257,542,568]" pageId="13" pageNumber="514">Egg:</emphasis>
Oval-shaped, with 134143 μm in length and 98100 μm in width. Very flat cap-shaped epithema. Chorion finely porous, both micropyle and sperm guide not visible (
<figureCitation id="1367F584FFA0FFC2FC3BFDDFFBE3FDBE" box="[961,1106,578,604]" captionStart="FIGURE 11" captionStartId="16.[151,253,1280,1304]" captionTargetBox="[151,1435,199,1256]" captionTargetId="figure-26@16.[151,1435,199,1256]" captionTargetPageId="16" captionText="FIGURE 11. Clypeocaenis multisetosa Soldán, 1978, from Xe Nou River, Xe Bang Hieng River, and outlet Stream of Nam Xouang Resurvoir in Lao PDR, A. Abdominal terga IIVII. B. Operculate gill. C, D. Eggs. Scale bars: A, B = 500 μm; C, D = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13745391" httpUri="https://zenodo.org/record/13745391/files/figure.png" pageId="13" pageNumber="514">Fig. 11C, D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C346BA8AFFA0FFC2FF3DFDFBFA9BFB7E" pageId="13" pageNumber="514" type="discussion">
<paragraph id="8BE3E901FFA0FFC2FF3DFDFBFA9BFB7E" blockId="13.[151,1437,289,1504]" pageId="13" pageNumber="514">
<emphasis id="B9283513FFA0FFC2FF3DFDFBFEF1FD62" bold="true" box="[199,320,614,640]" pageId="13" pageNumber="514">Remarks.</emphasis>
<taxonomicName id="4C5C9282FFA0FFC2FEB2FDFBFDC7FD62" authorityName="Soldan" authorityYear="1978" box="[328,630,614,640]" class="Insecta" family="Caenidae" genus="Clypeocaenis" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="multisetosa">
<emphasis id="B9283513FFA0FFC2FEB2FDFBFDC7FD62" box="[328,630,614,640]" italics="true" pageId="13" pageNumber="514">Clypeocaenis multisetosa</emphasis>
</taxonomicName>
was initially reported from
<collectingCountry id="F34BA991FFA0FFC2FC38FDFBFC4EFD62" box="[962,1023,614,640]" name="India" pageId="13" pageNumber="514">India</collectingCountry>
by Soldán in 1978. The specimens from Lao PDR were identified as
<taxonomicName id="4C5C9282FFA0FFC2FDA8FD17FCBAFD46" authorityName="Soldan" authorityYear="1978" box="[594,779,650,676]" class="Insecta" family="Caenidae" genus="Clypeocaenis" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="multisetosa">
<emphasis id="B9283513FFA0FFC2FDA8FD17FCBAFD46" box="[594,779,650,676]" italics="true" pageId="13" pageNumber="514">Cl. multisetosa</emphasis>
</taxonomicName>
based on a combination of morphological features, including the number and arrangement of setae on the clypeal protrusion with less than 20 long setae (
<figureCitation id="1367F584FFA0FFC2FA97FD33FF0FFD0E" captionStart="FIGURE 9" captionStartId="14.[151,250,1776,1800]" captionTargetBox="[158,1430,179,1750]" captionTargetId="figure-26@14.[158,1430,179,1750]" captionTargetPageId="14" captionText="FIGURE 9. Clypeocaenis multisetosa Soldán, 1978, collected from Xe Nou River, Xe Bang Hieng River, and outlet Stream of Nam Xouang Resurvoir in Lao PDR, A. Head capsule. B. Labrum. C. Mandibles. D. Maxillae. E. Hypopharynx. F. Labium. Scale bars: A, D (left maxillary palp) = 500 μm; B, C (right mandible), E = 200 μm; C (left mandible) = 300 μm; D, F (right maxillary palp) = 400 μm." figureDoi="http://doi.org/10.5281/zenodo.13745385" httpUri="https://zenodo.org/record/13745385/files/figure.png" pageId="13" pageNumber="514">Fig. 9A</figureCitation>
), maxillary palp two segmented, labial palp with second segment short and irregularly trapezoidal, and third segment elongated (
<figureCitation id="1367F584FFA0FFC2FE36FD6BFDFEFCF2" box="[460,591,758,784]" captionStart="FIGURE 9" captionStartId="14.[151,250,1776,1800]" captionTargetBox="[158,1430,179,1750]" captionTargetId="figure-26@14.[158,1430,179,1750]" captionTargetPageId="14" captionText="FIGURE 9. Clypeocaenis multisetosa Soldán, 1978, collected from Xe Nou River, Xe Bang Hieng River, and outlet Stream of Nam Xouang Resurvoir in Lao PDR, A. Head capsule. B. Labrum. C. Mandibles. D. Maxillae. E. Hypopharynx. F. Labium. Scale bars: A, D (left maxillary palp) = 500 μm; B, C (right mandible), E = 200 μm; C (left mandible) = 300 μm; D, F (right maxillary palp) = 400 μm." figureDoi="http://doi.org/10.5281/zenodo.13745385" httpUri="https://zenodo.org/record/13745385/files/figure.png" pageId="13" pageNumber="514">Fig. 9D, F</figureCitation>
). Distinctive transverse row of short stout setae on the dorsal surface of the forefemur, marginal setation patterns on the mid and hind femora (
<figureCitation id="1367F584FFA0FFC2FBEAFC87FB13FCD6" box="[1040,1186,794,820]" captionStart="FIGURE 10" captionStartId="15.[151,251,1467,1491]" captionTargetBox="[158,1430,199,1444]" captionTargetId="figure-18@15.[158,1430,199,1444]" captionTargetPageId="15" captionText="FIGURE 10. Clypeocaenis multisetosa Soldán, 1978, from Xe Nou River, Xe Bang Hieng River, and outlet Stream of Nam Xouang Resurvoir, Lao PDR, A. Forefemur. B. Mid femur. C. Hind femur. D. Foreclaw. E. Mid claw. F. Hind claw. Scale bars: A, B = 500 μm; C = 400 μm; DF = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13745389" httpUri="https://zenodo.org/record/13745389/files/figure.png" pageId="13" pageNumber="514">Fig. 10AC</figureCitation>
), median projections and spines on tergum II, and the characteristic posterior margin process of terga VIII and IX, as well as the structure of the operculate gill (
<figureCitation id="1367F584FFA0FFC2FDF2FCFFFD24FC9E" box="[520,661,866,892]" captionStart="FIGURE 11" captionStartId="16.[151,253,1280,1304]" captionTargetBox="[151,1435,199,1256]" captionTargetId="figure-26@16.[151,1435,199,1256]" captionTargetPageId="16" captionText="FIGURE 11. Clypeocaenis multisetosa Soldán, 1978, from Xe Nou River, Xe Bang Hieng River, and outlet Stream of Nam Xouang Resurvoir in Lao PDR, A. Abdominal terga IIVII. B. Operculate gill. C, D. Eggs. Scale bars: A, B = 500 μm; C, D = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13745391" httpUri="https://zenodo.org/record/13745391/files/figure.png" pageId="13" pageNumber="514">Fig. 11A, B</figureCitation>
). The specimens from Lao PDR exhibit some variations compared to the original description by
<bibRefCitation id="EFCD94F0FFA0FFC2FE0EFC1BFD2EFC42" author="Soldan, T." box="[500,671,902,928]" pageId="13" pageNumber="514" pagination="119 - 129" refId="ref10571" refString="Soldan, T. (1978) New genera and species of Caenidae (Ephemeroptera) from Iran, India and Australia. Acta Entomologica Bohemoslovaca, 75, 119 - 129." type="journal article" year="1978">Soldán (1978)</bibRefCitation>
, such as the presence of 67 setae on the operculate gills medial ridge and 45 denticles on the tarsal claw, in contrast to the 34 setae on the medial ridge and 68 denticles on the tarsal claw as reported by
<bibRefCitation id="EFCD94F0FFA0FFC2FDE1FC53FD76FC0A" author="Soldan, T." box="[539,711,974,1000]" pageId="13" pageNumber="514" pagination="119 - 129" refId="ref10571" refString="Soldan, T. (1978) New genera and species of Caenidae (Ephemeroptera) from Iran, India and Australia. Acta Entomologica Bohemoslovaca, 75, 119 - 129." type="journal article" year="1978">Soldán (1978)</bibRefCitation>
. Additionally, shield-shaped microtrichia on the body surface and operculate gills, resembling a spider web with a smooth margin, were observed in
<taxonomicName id="4C5C9282FFA0FFC2FB52FC6FFAEAFBEE" authorityName="Soldan" authorityYear="1978" box="[1192,1371,1010,1036]" class="Insecta" family="Caenidae" genus="Clypeocaenis" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="multisetosa">
<emphasis id="B9283513FFA0FFC2FB52FC6FFAEAFBEE" box="[1192,1371,1010,1036]" italics="true" pageId="13" pageNumber="514">Cl. multisetosa</emphasis>
</taxonomicName>
(
<figureCitation id="1367F584FFA0FFC2FA96FC6FFF0CFBD2" captionStart="FIGURE 8" captionStartId="12.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-26@12.[158,1430,181,1752]" captionTargetPageId="12" captionText="FIGURE 8. Clypeocaenis oligosetosa Soldán, 1983 (A, B) and Clypeocaenis multisetosa Soldán, 1978 (C, D). A, C. High magnification view of microtrichia on the dorsal surface of operculate gills. B, D. Microtrichia band on the ventral side of operculate gills. Scale bars: A = 20 μm; C = 50 μm; B, D = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13745381" httpUri="https://zenodo.org/record/13745381/files/figure.png" pageId="13" pageNumber="514">Fig. 8C</figureCitation>
), differing from the characteristics of
<taxonomicName id="4C5C9282FFA0FFC2FD85FB8BFC80FBD2" authorityName="Soldan" authorityYear="1983" box="[639,817,1046,1072]" class="Insecta" family="Caenidae" genus="Clypeocaenis" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="oligosetosa">
<emphasis id="B9283513FFA0FFC2FD85FB8BFC80FBD2" box="[639,817,1046,1072]" italics="true" pageId="13" pageNumber="514">Cl. oligosetosa</emphasis>
</taxonomicName>
(
<figureCitation id="1367F584FFA0FFC2FCBBFB8BFC11FBD2" box="[833,928,1046,1072]" captionStart="FIGURE 8" captionStartId="12.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-26@12.[158,1430,181,1752]" captionTargetPageId="12" captionText="FIGURE 8. Clypeocaenis oligosetosa Soldán, 1983 (A, B) and Clypeocaenis multisetosa Soldán, 1978 (C, D). A, C. High magnification view of microtrichia on the dorsal surface of operculate gills. B, D. Microtrichia band on the ventral side of operculate gills. Scale bars: A = 20 μm; C = 50 μm; B, D = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13745381" httpUri="https://zenodo.org/record/13745381/files/figure.png" pageId="13" pageNumber="514">Fig. 8A</figureCitation>
). The microtrichia band on the ventral side of the operculate gills follows closely along the lateral margin, terminating well before the gills hind margin. This band comprises clusters of 2 or 3 spines and single spines, primarily located on the inner margin of the band (
<figureCitation id="1367F584FFA0FFC2FF1BFB1FFEFAFB7E" box="[225,331,1154,1180]" captionStart="FIGURE 8" captionStartId="12.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-26@12.[158,1430,181,1752]" captionTargetPageId="12" captionText="FIGURE 8. Clypeocaenis oligosetosa Soldán, 1983 (A, B) and Clypeocaenis multisetosa Soldán, 1978 (C, D). A, C. High magnification view of microtrichia on the dorsal surface of operculate gills. B, D. Microtrichia band on the ventral side of operculate gills. Scale bars: A = 20 μm; C = 50 μm; B, D = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13745381" httpUri="https://zenodo.org/record/13745381/files/figure.png" pageId="13" pageNumber="514">Figs. 8D</figureCitation>
,
<figureCitation id="1367F584FFA0FFC2FEA2FB1FFE3DFB7E" box="[344,396,1154,1180]" captionStart="FIGURE 12" captionStartId="18.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-26@18.[158,1430,181,1752]" captionTargetPageId="18" captionText="FIGURE 12. Microtrichia bands on the ventral side of the operculate gill, as previously documented. A. Clypeocaenis soldani. B. Clypeocaenis multisetosa. C. Clypeocaenis oligosetosa. D. Clypeocaenis kaveri. E. Clypeocaenis afrosetosa. F. Clypeocaenis laosensis sp. nov. G. Mekongellina sangpradubae." figureDoi="http://doi.org/10.5281/zenodo.13745395" httpUri="https://zenodo.org/record/13745395/files/figure.png" pageId="13" pageNumber="514">12B</figureCitation>
). Eggs extracted from mature female larvae are documented for the first time.
</paragraph>
</subSubSection>
<subSubSection id="C346BA8AFFA0FFC2FF3DFB38FE65FA02" pageId="13" pageNumber="514" type="distribution">
<paragraph id="8BE3E901FFA0FFC2FF3DFB38FE65FA02" blockId="13.[151,1437,289,1504]" pageId="13" pageNumber="514">
<emphasis id="B9283513FFA0FFC2FF3DFB38FE4DFB22" bold="true" box="[199,508,1189,1216]" pageId="13" pageNumber="514">
<emphasis id="B9283513FFA0FFC2FF3DFB38FE45FB22" bold="true" box="[199,500,1189,1216]" italics="true" pageId="13" pageNumber="514">Distribution and habitats</emphasis>
.
</emphasis>
<taxonomicName id="4C5C9282FFA0FFC2FDFBFB3BFC9DFB22" authorityName="Soldan" authorityYear="1978" box="[513,812,1190,1216]" class="Insecta" family="Caenidae" genus="Clypeocaenis" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="multisetosa">
<emphasis id="B9283513FFA0FFC2FDFBFB3BFC9DFB22" box="[513,812,1190,1216]" italics="true" pageId="13" pageNumber="514">Clypeocaenis multisetosa</emphasis>
</taxonomicName>
specimens were documented in the
<collectingRegion id="499827E3FFA0FFC2FB33FB3BFA8BFB22" box="[1225,1338,1190,1216]" country="Laos" name="Viangchan" pageId="13" pageNumber="514">Vientiane</collectingRegion>
Capital,
<collectingRegion id="499827E3FFA0FFC2FF6DFB57FE8EFB06" box="[151,319,1226,1252]" country="Laos" name="Khammouan" pageId="13" pageNumber="514">Khammouane</collectingRegion>
, and
<collectingRegion id="499827E3FFA0FFC2FE85FB57FDA7FB06" box="[383,534,1226,1252]" country="Laos" name="Savannakhet" pageId="13" pageNumber="514">Savannakhet</collectingRegion>
Provinces of the Lao PDR (
<figureCitation id="1367F584FFA0FFC2FC96FB57FB81FB06" box="[876,1072,1226,1252]" captionStart="FIGURE 13" captionStartId="19.[151,251,1776,1800]" captionTargetBox="[158,1430,179,1750]" captionTargetId="figure-18@19.[158,1430,179,1750]" captionTargetPageId="19" captionText="FIGURE 13. Habitats of the genus Clypeocaenis Soldán, 1978 in Lao PDR, A. Xe Nou River, Xebangfai District, Khammouane Province. B. Tha Hao Stream, Xaybury District, Savannakhet Province. C. Xe Bang Hieng River, Songkhone District, Savannakhet Province. D. Outlet Stream of Nam Xouang Reservoir, Naxaythong District, Vientiane Capital." figureDoi="http://doi.org/10.5281/zenodo.13745397" httpUri="https://zenodo.org/record/13745397/files/figure.png" pageId="13" pageNumber="514">Figs. 13A, C, D</figureCitation>
,
<figureCitation id="1367F584FFA0FFC2FBC7FB57FBEFFB06" box="[1085,1118,1226,1252]" captionStart="FIGURE 14" captionStartId="20.[151,250,1091,1115]" captionTargetBox="[160,1430,185,1064]" captionTargetId="figure-26@20.[158,1430,181,1067]" captionTargetPageId="20" captionText="FIGURE 14. Geographic distribution of the Clypeocaenis laosensis sp. nov. (circle), C. oligosetosa (star); C. multisetosa (triangle)." figureDoi="http://doi.org/10.5281/zenodo.13745401" httpUri="https://zenodo.org/record/13745401/files/figure.png" pageId="13" pageNumber="514">14</figureCitation>
). This species was located in the littoral zones of the Xe Nou River, Xe Bang Hieng River, and the outlet stream of the Nam Xouang Reservoir. The ecological characteristics of the Xe Nou and Xe Bang Hieng Rivers have been previously mentioned. The outlet stream of the Nam Xouang Reservoir meanders through a village. Surrounding areas comprise residential settlements, agricultural fields, gardens, and areas designated for livestock activities. The water velocity within this stream is noted to be low (less than
<quantity id="4CA444E4FFA0FFC2FC4BFAE3FC48FA7A" box="[945,1017,1406,1433]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="514" unit="m" value="0.5">0.5 m</quantity>
/s). The substrate primarily consists of bedrock, interspersed with boulders, sand, silt, and accumulations of detritus leaf litter, providing a diverse habitat for
<taxonomicName id="4C5C9282FFA0FFC2FEE0FA5BFE7CFA02" authorityName="Soldan" authorityYear="1978" box="[282,461,1478,1504]" class="Insecta" family="Caenidae" genus="Clypeocaenis" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="multisetosa">
<emphasis id="B9283513FFA0FFC2FEE0FA5BFE7CFA02" box="[282,461,1478,1504]" italics="true" pageId="13" pageNumber="514">Cl. multisetosa</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:start id="168A56B369C51A4AB451D8D254284443">501</mods:start>
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</mods:extent>
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</mods:relatedItem>
<mods:location id="0992EDCA74DCC2CA9E2574A76FC6F916">
<mods:url id="93B992DCED853C0A23F7838D6FD9152D">http://dx.doi.org/10.11646/zootaxa.5501.4.1</mods:url>
</mods:location>
<mods:classification id="AB916AC27397D5F8700491A5E62B78F1">journal article</mods:classification>
<mods:identifier id="CFA14830CBF8DF8548E683EE3E891EDD" type="DOI">10.11646/zootaxa.5501.4.1</mods:identifier>
<mods:identifier id="D09F32C71F69E1AD7A40D3D200A28D45" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="C87126D52C4E9F946EE55FC411B9643B" type="Zenodo-Dep">13745355</mods:identifier>
<mods:identifier id="A7801868EFC236CD0B83F8477D4E23A1" type="ZooBank">29E9284E-6F5C-46E5-BA25-B54F820BE0C0</mods:identifier>
</mods:mods>
<treatment id="03F55817FFAFFFC6FF6DFF0AFD53FD13" ID-DOI="http://doi.org/10.5281/zenodo.13745361" ID-Zenodo-Dep="13745361" LSID="urn:lsid:plazi:treatment:03F55817FFAFFFC6FF6DFF0AFD53FD13" httpUri="http://treatment.plazi.org/id/03F55817FFAFFFC6FF6DFF0AFD53FD13" lastPageId="9" lastPageNumber="510" pageId="2" pageNumber="503">
<subSubSection id="C346BA8AFFAFFFCDFF6DFF0AFB35FF53" box="[151,1156,151,178]" pageId="2" pageNumber="503" type="nomenclature">
<paragraph id="8BE3E901FFAFFFCDFF6DFF0AFB35FF53" blockId="2.[151,1156,151,178]" box="[151,1156,151,178]" pageId="2" pageNumber="503">
<heading id="D0AB5E6DFFAFFFCDFF6DFF0AFB35FF53" bold="true" box="[151,1156,151,178]" fontSize="11" level="1" pageId="2" pageNumber="503" reason="1">
<emphasis id="B9283513FFAFFFCDFF6DFF0AFB35FF53" bold="true" box="[151,1156,151,178]" pageId="2" pageNumber="503">
<taxonomicName id="4C5C9282FFAFFFCDFF6DFF0AFC24FF53" authority="Vannachak, Malzacher &amp; Sangpradub" authorityName="Vannachak, Malzacher &amp; Sangpradub" authorityYear="2024" box="[151,917,151,178]" class="Insecta" family="Caenidae" genus="Clypeocaenis" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="laosensis" status="sp. nov.">
<emphasis id="B9283513FFAFFFCDFF6DFF0AFE1AFF50" bold="true" box="[151,427,151,178]" italics="true" pageId="2" pageNumber="503">Clypeocaenis laosensis</emphasis>
Vannachak, Malzacher &amp; Sangpradub
</taxonomicName>
<taxonomicNameLabel id="A21B8868FFAFFFCDFC66FF0AFC4BFF53" box="[924,1018,151,177]" pageId="2" pageNumber="503" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="1367F584FFAFFFCDFBF3FF0AFBCBFF53" box="[1033,1146,151,177]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="3.[151,250,2008,2032]" captionStartId-1="4.[151,250,1776,1800]" captionStartId-2="5.[151,250,1776,1800]" captionStartId-3="6.[151,250,1776,1800]" captionStartId-4="7.[151,250,1776,1800]" captionTargetBox-0="[261,1327,626,1982]" captionTargetBox-1="[158,1430,181,1752]" captionTargetBox-2="[158,1430,181,1752]" captionTargetBox-3="[158,1430,181,1752]" captionTargetBox-4="[158,1430,181,1752]" captionTargetId-0="figure-232@3.[261,1327,626,1982]" captionTargetId-1="figure-26@4.[158,1430,181,1752]" captionTargetId-2="figure-18@5.[158,1430,181,1752]" captionTargetId-3="figure-26@6.[158,1430,181,1752]" captionTargetId-4="figure-18@7.[158,1430,181,1752]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionText-0="FIGURE 1. Clypeocaenis laosensis sp. nov., habitus. A. Female larva. B. Male larva. Scale bars: 1 mm." captionText-1="FIGURE 2. Clypeocaenis laosensis sp. nov., A. Head capsule, viewed dorsally. B. Labrum. C. Mandibles. D. Maxillae. E. Hypopharynx. F. Labium. Scale bars: A = 500 μm; BD = 200 μm; E = 100 μm; F = 300 μm." captionText-2="FIGURE 3. Clypeocaenis laosensis sp. nov., A. Foreleg. B. High magnification view setae of transverse row forefemur. C. Forecoxa. D. Foreclaw. E. Mid leg. F. Mid claw. G. Hind leg. H. Hind claw. Scale bars: A, E, G = 500 μm; B = 300 μm; C = 200 μm; D, F, H = 100 μm." captionText-3="FIGURE 4. Clypeocaesni laosensis sp. nov., A. Abdominal terga VIX. B. Tergum II. C. Sternum IX. D. Operculate gill, dorsal view. E. Higher magnification of the operculate gill, dorsal view. F. Spatulate setae on the medial ridge. G. Operculate gill, viewed from the ventral side. Scale bars: A, D, G = 500 μm; B = 100 μm; C, F = 200 μm; E = 50 μm." captionText-4="FIGURE 5. Clpeocaenis laosensis sp. nov., A. Gills III, IV. B. Egg, arrow indicate the micropyle. C. Epithema. D. Detail of the micropyle. Scale bars: A = 500 μm; B = 100 μm; C = 40 μm; D = 30 μm." figureDoi-0="http://doi.org/10.5281/zenodo.13745357" figureDoi-1="http://doi.org/10.5281/zenodo.13745363" figureDoi-2="http://doi.org/10.5281/zenodo.13745365" figureDoi-3="http://doi.org/10.5281/zenodo.13745367" figureDoi-4="http://doi.org/10.5281/zenodo.13745371" httpUri-0="https://zenodo.org/record/13745357/files/figure.png" httpUri-1="https://zenodo.org/record/13745363/files/figure.png" httpUri-2="https://zenodo.org/record/13745365/files/figure.png" httpUri-3="https://zenodo.org/record/13745367/files/figure.png" httpUri-4="https://zenodo.org/record/13745371/files/figure.png" pageId="2" pageNumber="503">Figs. 15</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C346BA8AFFAFFFCCFF6DFF46FE2BFDBE" lastPageId="3" lastPageNumber="504" pageId="2" pageNumber="503" type="materials_examined">
<paragraph id="8BE3E901FFAFFFCDFF6DFF46FDDFFEDF" blockId="2.[151,1437,219,2024]" pageId="2" pageNumber="503">
<emphasis id="B9283513FFAFFFCDFF6DFF46FE32FF17" bold="true" box="[151,387,219,246]" pageId="2" pageNumber="503">
<emphasis id="B9283513FFAFFFCDFF6DFF46FECDFF14" bold="true" box="[151,380,219,246]" italics="true" pageId="2" pageNumber="503">Materials examined</emphasis>
.
</emphasis>
<materialsCitation id="3B34E35CFFAFFFCDFE73FF46FDDFFEDF" collectingDate="2022-01-06" collectionCode="V" collectorName="Vannachak &amp; Natural History Museum" country="Thailand" county="Xe Nou River" elevation="154" latitude="17.068213" location="Male" longLatPrecision="1" longitude="105.070465" municipality="Xebangfai District" pageId="2" pageNumber="503" specimenCount="1" typeStatus="holotype">
<emphasis id="B9283513FFAFFFCDFE73FF46FE47FF17" bold="true" box="[393,502,219,245]" pageId="2" pageNumber="503">
<typeStatus id="54E757A3FFAFFFCDFE73FF46FE47FF17" box="[393,502,219,245]" pageId="2" pageNumber="503" type="holotype">Holotype</typeStatus>
</emphasis>
(in ethanol):
<location id="8E83BFDAFFAFFFCDFD70FF46FD75FF17" LSID="urn:lsid:plazi:treatment:03F55817FFAFFFC6FF6DFF0AFD53FD13:8E83BFDAFFAFFFCDFD70FF46FD75FF17" box="[650,708,219,245]" country="Thailand" county="Xe Nou River" latitude="17.068213" longLatPrecision="1" longitude="105.070465" municipality="Xebangfai District" name="Male" pageId="2" pageNumber="503">Male</location>
larva (THNHM-I-00029764),
<location id="8E83BFDAFFAFFFCDFBE0FF41FBF7FF14" LSID="urn:lsid:plazi:treatment:03F55817FFAFFFC6FF6DFF0AFD53FD13:8E83BFDAFFAFFFCDFBE0FF41FBF7FF14" box="[1050,1094,220,246]" country="Thailand" county="Xe Nou River" latitude="17.068213" longLatPrecision="1" longitude="105.070465" municipality="Xebangfai District" name="Lao" pageId="2" pageNumber="503">Lao</location>
PDR,
<location id="8E83BFDAFFAFFFCDFB75FF46FA29FF17" LSID="urn:lsid:plazi:treatment:03F55817FFAFFFC6FF6DFF0AFD53FD13:8E83BFDAFFAFFFCDFB75FF46FA29FF17" box="[1167,1432,219,245]" country="Thailand" county="Xe Nou River" latitude="17.068213" longLatPrecision="1" longitude="105.070465" municipality="Xebangfai District" name="Khammouane Province" pageId="2" pageNumber="503">Khammouane Province</location>
,
<collectingMunicipality id="6B87737BFFAFFFCDFF6DFF62FED8FEFB" box="[151,361,255,281]" pageId="2" pageNumber="503">Xebangfai District</collectingMunicipality>
,
<collectingCounty id="6282918DFFAFFFCDFE88FE9DFDBFFEFB" box="[370,526,255,282]" pageId="2" pageNumber="503">Xe Nou River</collectingCounty>
,
<geoCoordinate id="EE688FC6FFAFFFCDFDE2FF62FD1AFEF8" box="[536,683,255,282]" degrees="17" direction="north" minutes="4" orientation="latitude" pageId="2" pageNumber="503" precision="1" seconds="5.56" value="17.068213">17°45.56”N</geoCoordinate>
,
<geoCoordinate id="EE688FC6FFAFFFCDFD4FFF62FCD4FEF8" box="[693,869,255,282]" degrees="105" direction="east" minutes="4" orientation="longitude" pageId="2" pageNumber="503" precision="1" seconds="13.68" value="105.070465">105°413.68E</geoCoordinate>
,
<elevation id="00710E32FFAFFFCDFC95FF62FC41FEFB" box="[879,1008,255,282]" metricMagnitude="2" metricUnit="m" metricValue="1.54" pageId="2" pageNumber="503" unit="m" value="154.0">
<quantity id="4CA444E4FFAFFFCDFC95FF62FC04FEF8" box="[879,949,255,282]" metricMagnitude="2" metricUnit="m" metricValue="1.54" pageId="2" pageNumber="503" unit="m" value="154.0">154 m</quantity>
a.s.l.
</elevation>
,
<date id="FFE2CFC1FFAFFFCDFC01FF62FBEAFEFB" box="[1019,1115,255,281]" pageId="2" pageNumber="503" value="2022-01-06">
<collectingDate id="EFA63629FFAFFFCDFC01FF62FBEAFEFB" box="[1019,1115,255,281]" pageId="2" pageNumber="503" value="2022-01-06">6.I.2022</collectingDate>
</date>
,
<collectionCode id="ED4D71C4FFAFFFCDFB9FFE9DFBC8FEF8" box="[1125,1145,256,282]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="503" type="Museum">V</collectionCode>
.
<collectorName id="26A98CD7FFAFFFCDFB7BFF62FB4DFEFB" box="[1153,1276,255,281]" pageId="2" pageNumber="503">Vannachak</collectorName>
leg, deposited in the
<collectorName id="26A98CD7FFAFFFCDFF27FEBEFE49FEDC" box="[221,504,291,318]" pageId="2" pageNumber="503">Natural History Museum</collectorName>
,
<collectingCountry id="F34BA991FFAFFFCDFDF9FEBEFDDBFEDF" box="[515,618,291,317]" name="Thailand" pageId="2" pageNumber="503">Thailand</collectingCountry>
.
</materialsCitation>
</paragraph>
<paragraph id="8BE3E901FFAFFFCDFF3DFEDAFDAEFE4B" blockId="2.[151,1437,219,2024]" pageId="2" pageNumber="503">
<materialsCitation id="3B34E35CFFAFFFCDFF3DFEDAFDAAFE4B" collectingDate="2022-01-06" collectionCode="V" collectorName="Vannachak &amp; Natural History Museum" country="Thailand" county="Xe Nou River" elevation="154" latitude="17.068213" location="Natural History Museum" longLatPrecision="1" longitude="105.070465" municipality="Cl" pageId="2" pageNumber="503" specimenCount="18" specimenCount-female="5" specimenCount-male="13" typeStatus="paratype">
<emphasis id="B9283513FFAFFFCDFF3DFEDAFE8EFE83" bold="true" box="[199,319,327,353]" pageId="2" pageNumber="503">
<typeStatus id="54E757A3FFAFFFCDFF3DFEDAFE8EFE83" box="[199,319,327,353]" pageId="2" pageNumber="503" type="paratype">Paratypes</typeStatus>
</emphasis>
(in ethanol):
<specimenCount id="9D5A2288FFAFFFCDFE23FEDAFD95FE83" box="[473,548,327,353]" count="3" pageId="2" pageNumber="503" type="male">3 male</specimenCount>
and
<specimenCount id="9D5A2288FFAFFFCDFDA6FEDAFD0FFE83" box="[604,702,327,353]" count="2" pageId="2" pageNumber="503" type="female">2 female</specimenCount>
larvae (THNHM-I-0002976500029769) same data as holotype, deposited in the
<location id="8E83BFDAFFAFFFCDFEA6FEF6FDCEFE64" LSID="urn:lsid:plazi:treatment:03F55817FFAFFFC6FF6DFF0AFD53FD13:8E83BFDAFFAFFFCDFEA6FEF6FDCEFE64" box="[348,639,363,390]" country="Thailand" municipality="Cl" name="Natural History Museum" pageId="2" pageNumber="503">Natural History Museum</location>
,
<collectingCountry id="F34BA991FFAFFFCDFD74FEF6FD44FE67" box="[654,757,363,389]" name="Thailand" pageId="2" pageNumber="503">Thailand</collectingCountry>
.
<specimenCount id="9D5A2288FFAFFFCDFCF9FEF6FCD1FE67" box="[771,864,363,389]" count="10" pageId="2" pageNumber="503" type="male">10 male</specimenCount>
and
<specimenCount id="9D5A2288FFAFFFCDFC65FEF6FBB5FE67" box="[927,1028,363,389]" count="3" pageId="2" pageNumber="503" type="female">3 female</specimenCount>
larvae (
<collectingMunicipality id="6B87737BFFAFFFCDFB9DFEF6FB33FE67" box="[1127,1154,363,389]" pageId="2" pageNumber="503">Cl</collectingMunicipality>
/
<collectingCountry id="F34BA991FFAFFFCDFB71FEF6FB75FE67" box="[1163,1220,363,389]" name="Laos" pageId="2" pageNumber="503">Laos</collectingCountry>
.002) same data as
<typeStatus id="54E757A3FFAFFFCDFF6DFE12FF4DFE4B" box="[151,252,399,425]" pageId="2" pageNumber="503" type="holotype">holotype</typeStatus>
, deposited in NUoL-AIC
</materialsCitation>
.
</paragraph>
<paragraph id="8BE3E901FFAFFFCDFF3DFE2EFB35FDDB" blockId="2.[151,1437,219,2024]" pageId="2" pageNumber="503">
<emphasis id="B9283513FFAFFFCDFF3DFE2EFE32FE2F" bold="true" box="[199,387,435,461]" pageId="2" pageNumber="503">Other materials</emphasis>
:
<materialsCitation id="3B34E35CFFAFFFCDFE6AFE2EFB4EFE13" collectingDate="2019-11-23" country="Laos" county="Cl" elevation="140" latitude="16.865131" location="Xaybury District" longLatPrecision="1" longitude="104.90105" municipality="Lao" pageId="2" pageNumber="503" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Savannakhet">
<specimenCount id="9D5A2288FFAFFFCDFE6AFE2EFE68FE2F" box="[400,473,435,461]" count="1" pageId="2" pageNumber="503" type="male">1 male</specimenCount>
and
<specimenCount id="9D5A2288FFAFFFCDFDF7FE2EFDDDFE2F" box="[525,620,435,461]" count="2" pageId="2" pageNumber="503" type="female">2 female</specimenCount>
larvae (
<collectingCounty id="6282918DFFAFFFCDFD39FE2EFD6EFE2F" box="[707,735,435,461]" pageId="2" pageNumber="503">Cl</collectingCounty>
/
<collectingCountry id="F34BA991FFAFFFCDFD12FE2EFC93FE2F" box="[744,802,435,461]" name="Laos" pageId="2" pageNumber="503">Laos</collectingCountry>
.003),
<collectingMunicipality id="6B87737BFFAFFFCDFC99FE29FC3EFE2C" box="[867,911,436,462]" pageId="2" pageNumber="503">Lao</collectingMunicipality>
PDR,
<collectingRegion id="499827E3FFAFFFCDFC2CFE2EFB61FE2F" box="[982,1232,435,461]" country="Laos" name="Savannakhet" pageId="2" pageNumber="503">Savannakhet Province</collectingRegion>
,
<location id="8E83BFDAFFAFFFCDFB20FE2EFA29FE2F" LSID="urn:lsid:plazi:treatment:03F55817FFAFFFC6FF6DFF0AFD53FD13:8E83BFDAFFAFFFCDFB20FE2EFA29FE2F" box="[1242,1432,435,461]" country="Laos" county="Cl" latitude="16.865131" longLatPrecision="1" longitude="104.90105" municipality="Lao" name="Xaybury District" pageId="2" pageNumber="503" stateProvince="Savannakhet">Xaybury District</location>
,
<location id="8E83BFDAFFAFFFCDFF6DFE4AFEE2FE13" LSID="urn:lsid:plazi:treatment:03F55817FFAFFFC6FF6DFF0AFD53FD13:8E83BFDAFFAFFFCDFF6DFE4AFEE2FE13" box="[151,339,471,498]" country="Laos" county="Cl" latitude="16.865131" longLatPrecision="1" longitude="104.90105" municipality="Lao" name="Tha Hao Stream" pageId="2" pageNumber="503" stateProvince="Savannakhet">Tha Hao Stream</location>
,
<geoCoordinate id="EE688FC6FFAFFFCDFEA5FE4AFDBFFE10" box="[351,526,471,498]" degrees="16" direction="north" minutes="51" orientation="latitude" pageId="2" pageNumber="503" precision="1" seconds="54.47" value="16.865131">16°5154.47”N</geoCoordinate>
,
<geoCoordinate id="EE688FC6FFAFFFCDFDE1FE4AFD77FE10" box="[539,710,471,498]" degrees="104" direction="east" minutes="54" orientation="longitude" pageId="2" pageNumber="503" precision="1" seconds="3.77" value="104.90105">104°543.77”E</geoCoordinate>
,
<elevation id="00710E32FFAFFFCDFD29FE4AFCE9FE13" box="[723,856,471,498]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="2" pageNumber="503" unit="m" value="140.0">
<quantity id="4CA444E4FFAFFFCDFD29FE4AFCABFE10" box="[723,794,471,498]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="2" pageNumber="503" unit="m" value="140.0">140 m</quantity>
a.s.l.
</elevation>
,
<date id="FFE2CFC1FFAFFFCDFC9FFE4AFC59FE13" box="[869,1000,471,497]" pageId="2" pageNumber="503" value="2019-11-23">
<collectingDate id="EFA63629FFAFFFCDFC9FFE4AFC59FE13" box="[869,1000,471,497]" pageId="2" pageNumber="503" value="2019-11-23">23.XI.2019</collectingDate>
</date>
, deposited in KKU-AIC
</materialsCitation>
.
<materialsCitation id="3B34E35CFFAFFFCDFAF0FE4AFB35FDDB" collectingDate="2019-11-24" collectionCode="V" collectorName="Vannachak" country="Laos" county="Cl" elevation="126" latitude="16.097618" location="Xe Bang Hieng River" longLatPrecision="1" longitude="105.37587" municipality="Songkhone District" pageId="2" pageNumber="503" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Savannakhet">
<specimenCount id="9D5A2288FFAFFFCDFAF0FE4AFADDFE13" box="[1290,1388,471,497]" count="4" pageId="2" pageNumber="503" type="female">4 female</specimenCount>
and
<specimenCount id="9D5A2288FFAFFFCDFF6DFE66FF55FDF7" box="[151,228,507,533]" count="2" pageId="2" pageNumber="503" type="male">2 male</specimenCount>
larvae (
<collectingCounty id="6282918DFFAFFFCDFEBEFE66FED1FDF7" box="[324,352,507,533]" pageId="2" pageNumber="503">Cl</collectingCounty>
/
<collectingCountry id="F34BA991FFAFFFCDFE92FE66FE13FDF7" box="[360,418,507,533]" name="Laos" pageId="2" pageNumber="503">Laos</collectingCountry>
.004),
<collectingRegion id="499827E3FFAFFFCDFE12FE66FD56FDF7" box="[488,743,507,533]" country="Laos" name="Savannakhet" pageId="2" pageNumber="503">Savannakhet Province</collectingRegion>
,
<collectingMunicipality id="6B87737BFFAFFFCDFD0EFE66FC62FDF7" box="[756,979,507,533]" pageId="2" pageNumber="503">Songkhone District</collectingMunicipality>
,
<location id="8E83BFDAFFAFFFCDFC1AFE61FB6DFDF7" LSID="urn:lsid:plazi:treatment:03F55817FFAFFFC6FF6DFF0AFD53FD13:8E83BFDAFFAFFFCDFC1AFE61FB6DFDF7" box="[992,1244,507,534]" country="Laos" county="Cl" latitude="16.097618" longLatPrecision="1" longitude="105.37587" municipality="Songkhone District" name="Xe Bang Hieng River" pageId="2" pageNumber="503" stateProvince="Savannakhet">Xe Bang Hieng River</location>
,
<geoCoordinate id="EE688FC6FFAFFFCDFB13FE66FA26FDF4" box="[1257,1431,507,534]" degrees="16" direction="north" minutes="05" orientation="latitude" pageId="2" pageNumber="503" precision="1" seconds="51.42" value="16.097618">16°0551.42”N</geoCoordinate>
,
<geoCoordinate id="EE688FC6FFAFFFCDFF6DFD82FEE0FDD8" box="[151,337,543,570]" degrees="105" direction="east" minutes="22" orientation="longitude" pageId="2" pageNumber="503" precision="1" seconds="33.13" value="105.37587">105°2233.13”E</geoCoordinate>
,
<elevation id="00710E32FFAFFFCDFEA6FD82FE51FDDB" box="[348,480,543,570]" metricMagnitude="2" metricUnit="m" metricValue="1.26" pageId="2" pageNumber="503" unit="m" value="126.0">
<quantity id="4CA444E4FFAFFFCDFEA6FD82FE12FDD8" box="[348,419,543,570]" metricMagnitude="2" metricUnit="m" metricValue="1.26" pageId="2" pageNumber="503" unit="m" value="126.0">126 m</quantity>
a.s.l.
</elevation>
,
<date id="FFE2CFC1FFAFFFCDFE16FD82FDDFFDDB" box="[492,622,543,569]" pageId="2" pageNumber="503" value="2019-11-24">
<collectingDate id="EFA63629FFAFFFCDFE16FD82FDDFFDDB" box="[492,622,543,569]" pageId="2" pageNumber="503" value="2019-11-24">24.XI.2019</collectingDate>
</date>
, deposited in NUoL-AIC, all
<collectionCode id="ED4D71C4FFAFFFCDFC40FDBDFC7FFDD8" box="[954,974,544,570]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="503" type="Museum">V</collectionCode>
.
<collectorName id="26A98CD7FFAFFFCDFC22FD82FBE2FDDB" box="[984,1107,543,569]" pageId="2" pageNumber="503">Vannachak</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="8BE3E901FFAFFFCCFF3DFDDEFB4CFE2F" blockId="2.[151,1437,219,2024]" lastBlockId="3.[151,1437,151,605]" lastPageId="3" lastPageNumber="504" pageId="2" pageNumber="503">
<emphasis id="B9283513FFAFFFCDFF3DFDDEFE39FDBF" bold="true" box="[199,392,579,605]" pageId="2" pageNumber="503">Mature Nymph.</emphasis>
Body length (excl. cerci):
<quantity id="4CA444E4FFAFFFCDFD49FDDFFC8BFDBF" box="[691,826,578,605]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="2" pageNumber="503" unit="mm" value="3.0" valueMax="3.5" valueMin="2.5">2.53.5 mm</quantity>
(males) and
<quantity id="4CA444E4FFAFFFCDFC31FDDEFBE3FDBF" box="[971,1106,579,605]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="3.7" metricValueMin="3.3" pageId="2" pageNumber="503" unit="mm" value="3.5" valueMax="3.7" valueMin="3.3">3.33.7 mm</quantity>
(females); cerci
<quantity id="4CA444E4FFAFFFCDFAF4FDDFFA29FDBF" box="[1294,1432,578,605]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="2" pageNumber="503" unit="mm" value="1.35" valueMax="1.5" valueMin="1.2">1.21.5 mm</quantity>
; antennae
<quantity id="4CA444E4FFAFFFCDFEF9FDFBFE3EFD63" box="[259,399,614,641]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="2" pageNumber="503" unit="mm" value="1.0" valueMax="1.1" valueMin="0.9">0.91.1 mm</quantity>
. General coloration light yellow to pale yellow. Epidermal pigmentation: vertex in females with a transverse dark brown band between the lateral ocelli. Pronotum with a distinctive pattern as in
<figureCitation id="1367F584FFAFFFCDFAAAFD14FA27FD46" box="[1360,1430,649,676]" captionStart="FIGURE 1" captionStartId="3.[151,250,2008,2032]" captionTargetBox="[261,1327,626,1982]" captionTargetId="figure-232@3.[261,1327,626,1982]" captionTargetPageId="3" captionText="FIGURE 1. Clypeocaenis laosensis sp. nov., habitus. A. Female larva. B. Male larva. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13745357" httpUri="https://zenodo.org/record/13745357/files/figure.png" pageId="2" pageNumber="503">Fig. 1</figureCitation>
. Females with a dark brown, butterfly-like pattern between the wing pads. Abdominal terga I,
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, and
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IX in both sexes with broad transverse dark brown bands, contrasting to the light foundational coloration (
<figureCitation id="1367F584FFAFFFCDFB3EFD4DFA8FFD09" box="[1220,1342,720,747]" captionStart="FIGURE 1" captionStartId="3.[151,250,2008,2032]" captionTargetBox="[261,1327,626,1982]" captionTargetId="figure-232@3.[261,1327,626,1982]" captionTargetPageId="3" captionText="FIGURE 1. Clypeocaenis laosensis sp. nov., habitus. A. Female larva. B. Male larva. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13745357" httpUri="https://zenodo.org/record/13745357/files/figure.png" pageId="2" pageNumber="503">Fig. 1A, B</figureCitation>
).
<emphasis id="B9283513FFAFFFCDFAA9FD4CFA2DFD09" bold="true" box="[1363,1436,721,747]" pageId="2" pageNumber="503">Head:</emphasis>
Head length
<quantity id="4CA444E4FFAFFFCDFED9FD69FECBFCEC" box="[291,378,756,783]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="503" unit="mm" value="0.5">0.5 mm</quantity>
, width
<quantity id="4CA444E4FFAFFFCDFE30FD69FD91FCEC" box="[458,544,756,782]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="503" unit="mm" value="1.8">1.8 mm</quantity>
, light yellow color. Compound eyes black; ocelli with the apical half black, basal half whitish. Antennae transparent; pedicel with 23 short setae distally on inner margin, 3 shorter setae on ventral surface. Pedicels length three times of scape, one and a half times of antennal segment III. Clypeus protruded, with 2 moderately long setae, longer than pedicel (
<figureCitation id="1367F584FFAFFFCDFD1FFCC3FCF3FC9A" box="[741,834,862,888]" captionStart="FIGURE 2" captionStartId="4.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-26@4.[158,1430,181,1752]" captionTargetPageId="4" captionText="FIGURE 2. Clypeocaenis laosensis sp. nov., A. Head capsule, viewed dorsally. B. Labrum. C. Mandibles. D. Maxillae. E. Hypopharynx. F. Labium. Scale bars: A = 500 μm; BD = 200 μm; E = 100 μm; F = 300 μm." figureDoi="http://doi.org/10.5281/zenodo.13745363" httpUri="https://zenodo.org/record/13745363/files/figure.png" pageId="2" pageNumber="503">Fig. 2A</figureCitation>
). Labrum three times wider than long, with anterior margin and semicircular curved lateral margins apical half bearing bipinnate setae. The dorsum presents long, fine frayed setae and long, thin setae, with a few long, thin setae scattered near basal half (
<figureCitation id="1367F584FFAFFFCDFB8DFC38FB60FC5D" box="[1143,1233,933,959]" captionStart="FIGURE 2" captionStartId="4.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-26@4.[158,1430,181,1752]" captionTargetPageId="4" captionText="FIGURE 2. Clypeocaenis laosensis sp. nov., A. Head capsule, viewed dorsally. B. Labrum. C. Mandibles. D. Maxillae. E. Hypopharynx. F. Labium. Scale bars: A = 500 μm; BD = 200 μm; E = 100 μm; F = 300 μm." figureDoi="http://doi.org/10.5281/zenodo.13745363" httpUri="https://zenodo.org/record/13745363/files/figure.png" pageId="2" pageNumber="503">Fig. 2B</figureCitation>
). Outer incisor of mandibles with five denticles, situated below molar with 45 long, thin setae. The dorso-lateral surface with two curved transverse rows of long setae; 812 setae on basal row and 5 setae on topical row (
<figureCitation id="1367F584FFAFFFCDFB60FC71FB42FBE4" box="[1178,1267,1004,1030]" captionStart="FIGURE 2" captionStartId="4.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-26@4.[158,1430,181,1752]" captionTargetPageId="4" captionText="FIGURE 2. Clypeocaenis laosensis sp. nov., A. Head capsule, viewed dorsally. B. Labrum. C. Mandibles. D. Maxillae. E. Hypopharynx. F. Labium. Scale bars: A = 500 μm; BD = 200 μm; E = 100 μm; F = 300 μm." figureDoi="http://doi.org/10.5281/zenodo.13745363" httpUri="https://zenodo.org/record/13745363/files/figure.png" pageId="2" pageNumber="503">Fig. 2C</figureCitation>
). Maxillae and labium with setation patterns (
<figureCitation id="1367F584FFAFFFCDFE09FB8DFDDFFBC8" box="[499,622,1040,1066]" captionStart="FIGURE 2" captionStartId="4.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-26@4.[158,1430,181,1752]" captionTargetPageId="4" captionText="FIGURE 2. Clypeocaenis laosensis sp. nov., A. Head capsule, viewed dorsally. B. Labrum. C. Mandibles. D. Maxillae. E. Hypopharynx. F. Labium. Scale bars: A = 500 μm; BD = 200 μm; E = 100 μm; F = 300 μm." figureDoi="http://doi.org/10.5281/zenodo.13745363" httpUri="https://zenodo.org/record/13745363/files/figure.png" pageId="2" pageNumber="503">Fig. 2D, F</figureCitation>
). Maxillary palp segment I narrow, about half as wide as galea-lacinia. Base of labial palp segment III not significantly broad. Hypopharynxs superlingua with stout setae and linguas apical margin with thin setae at the apical margin (
<figureCitation id="1367F584FFAFFFCDFD37FBCBFC95FB93" box="[717,804,1110,1137]" captionStart="FIGURE 2" captionStartId="4.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-26@4.[158,1430,181,1752]" captionTargetPageId="4" captionText="FIGURE 2. Clypeocaenis laosensis sp. nov., A. Head capsule, viewed dorsally. B. Labrum. C. Mandibles. D. Maxillae. E. Hypopharynx. F. Labium. Scale bars: A = 500 μm; BD = 200 μm; E = 100 μm; F = 300 μm." figureDoi="http://doi.org/10.5281/zenodo.13745363" httpUri="https://zenodo.org/record/13745363/files/figure.png" pageId="2" pageNumber="503">Fig. 2E</figureCitation>
). Three segmented labial palps; segment III more or less sickle-shaped, with bluntly pointed tip, as long as segments I and
<collectionCode id="ED4D71C4FFAFFFCDFC82FBE6FC3AFB77" box="[888,907,1147,1173]" pageId="2" pageNumber="503">II</collectionCode>
together, segment III 4.24.5 times of segment
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. (
<figureCitation id="1367F584FFAFFFCDFF45FB00FEA2FB5A" box="[191,275,1181,1208]" captionStart="FIGURE 2" captionStartId="4.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-26@4.[158,1430,181,1752]" captionTargetPageId="4" captionText="FIGURE 2. Clypeocaenis laosensis sp. nov., A. Head capsule, viewed dorsally. B. Labrum. C. Mandibles. D. Maxillae. E. Hypopharynx. F. Labium. Scale bars: A = 500 μm; BD = 200 μm; E = 100 μm; F = 300 μm." figureDoi="http://doi.org/10.5281/zenodo.13745363" httpUri="https://zenodo.org/record/13745363/files/figure.png" pageId="2" pageNumber="503">Fig. 2F</figureCitation>
).
<emphasis id="B9283513FFAFFFCDFEDCFB03FE39FB5A" bold="true" box="[294,392,1182,1208]" pageId="2" pageNumber="503">Thorax:</emphasis>
Pronotum slightly convex sides with anterior protrusion (
<figureCitation id="1367F584FFAFFFCDFBFDFB00FBC6FB5A" box="[1031,1143,1181,1208]" captionStart="FIGURE 1" captionStartId="3.[151,250,2008,2032]" captionTargetBox="[261,1327,626,1982]" captionTargetId="figure-232@3.[261,1327,626,1982]" captionTargetPageId="3" captionText="FIGURE 1. Clypeocaenis laosensis sp. nov., habitus. A. Female larva. B. Male larva. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13745357" httpUri="https://zenodo.org/record/13745357/files/figure.png" pageId="2" pageNumber="503">Fig.1A, B</figureCitation>
). Mesonotum light yellow coloration. Legs: light yellow to whitish, cover with knobs, interspersed with very delicate microtrichia, each with a few needle-shaped branches (pine-needle like,
<figureCitation id="1367F584FFAFFFCDFD39FB79FCAAFB1D" box="[707,795,1252,1279]" captionStart="FIGURE 4" captionStartId="6.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-26@6.[158,1430,181,1752]" captionTargetPageId="6" captionText="FIGURE 4. Clypeocaesni laosensis sp. nov., A. Abdominal terga VIX. B. Tergum II. C. Sternum IX. D. Operculate gill, dorsal view. E. Higher magnification of the operculate gill, dorsal view. F. Spatulate setae on the medial ridge. G. Operculate gill, viewed from the ventral side. Scale bars: A, D, G = 500 μm; B = 100 μm; C, F = 200 μm; E = 50 μm." figureDoi="http://doi.org/10.5281/zenodo.13745367" httpUri="https://zenodo.org/record/13745367/files/figure.png" pageId="2" pageNumber="503">Fig. 4E</figureCitation>
). Foreleg: length of femur/ tibia/ tarsus/ claw viz., 0.52/ 0.42/ 0.41/
<quantity id="4CA444E4FFAFFFCDFEE2FA95FE31FAC0" box="[280,384,1288,1314]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="503" unit="mm" value="0.16">0.16 mm</quantity>
. Coxal processes sickle-shaped with a denticulated surface featuring thin setae and a serrated margin. Trochanter with one regular and another irregular row of short setae (
<figureCitation id="1367F584FFAFFFCDFBDFFAB6FB30FAA7" box="[1061,1153,1323,1349]" captionStart="FIGURE 3" captionStartId="5.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-18@5.[158,1430,181,1752]" captionTargetPageId="5" captionText="FIGURE 3. Clypeocaenis laosensis sp. nov., A. Foreleg. B. High magnification view setae of transverse row forefemur. C. Forecoxa. D. Foreclaw. E. Mid leg. F. Mid claw. G. Hind leg. H. Hind claw. Scale bars: A, E, G = 500 μm; B = 300 μm; C = 200 μm; D, F, H = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13745365" httpUri="https://zenodo.org/record/13745365/files/figure.png" pageId="2" pageNumber="503">Fig. 3C</figureCitation>
). Femur with three long filtering setae at the middle of the inner margin, short setae on the outer margins proximal half, and approximately ten large, broadly spatulate bristles arranged in a transverse row on the distal halfs dorsal surface. Area apical to transverse row with denticulated and scattered with long, pine-needle-like setae (
<figureCitation id="1367F584FFAFFFCDFBD3FA0BFB1EFA52" box="[1065,1199,1430,1456]" captionStart="FIGURE 3" captionStartId="5.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-18@5.[158,1430,181,1752]" captionTargetPageId="5" captionText="FIGURE 3. Clypeocaenis laosensis sp. nov., A. Foreleg. B. High magnification view setae of transverse row forefemur. C. Forecoxa. D. Foreclaw. E. Mid leg. F. Mid claw. G. Hind leg. H. Hind claw. Scale bars: A, E, G = 500 μm; B = 300 μm; C = 200 μm; D, F, H = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13745365" httpUri="https://zenodo.org/record/13745365/files/figure.png" pageId="2" pageNumber="503">Figs. 3A, B</figureCitation>
). Dorsal side of tibia with two transverse rows of very long filtering setae, extending longitudinally on the ventral side. Tarsus with a row of long filtering setae, inner margin with eight stout setae, outer margin with scattered, thin, hair-like setae. Claw long, slender, apically bent, with 35 denticles, smallest basal (
<figureCitation id="1367F584FFAFFFCDFC8EF99DFC60F9F8" box="[884,977,1536,1562]" captionStart="FIGURE 3" captionStartId="5.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-18@5.[158,1430,181,1752]" captionTargetPageId="5" captionText="FIGURE 3. Clypeocaenis laosensis sp. nov., A. Foreleg. B. High magnification view setae of transverse row forefemur. C. Forecoxa. D. Foreclaw. E. Mid leg. F. Mid claw. G. Hind leg. H. Hind claw. Scale bars: A, E, G = 500 μm; B = 300 μm; C = 200 μm; D, F, H = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13745365" httpUri="https://zenodo.org/record/13745365/files/figure.png" pageId="2" pageNumber="503">Fig. 3D</figureCitation>
). Midleg: length of femur/ tibia/ tarsus/ claw viz., 0.47/ 0.29/ 0.33/
<quantity id="4CA444E4FFAFFFCDFE2BF9BEFD88F9DC" box="[465,569,1571,1598]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="503" unit="mm" value="0.16">0.16 mm</quantity>
. Coxa and trochanter similar to foreleg. Femur with rows of spatulate setae, strongly narrowed at base on outer margins, microscopic setae on inner margin, proximal half of dorsal surface with scattered, short spatulate setae. Inner margin of tibia with a row of 812 stout setae and submarginal row of spatulate setae, outer margins with a few hair-like setae. Inner margin of tarsus with of seven stout setae, outer margin with scattered hair-like setae (
<figureCitation id="1367F584FFAFFFCDFE57F92CFDB5F92E" box="[429,516,1713,1740]" captionStart="FIGURE 3" captionStartId="5.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-18@5.[158,1430,181,1752]" captionTargetPageId="5" captionText="FIGURE 3. Clypeocaenis laosensis sp. nov., A. Foreleg. B. High magnification view setae of transverse row forefemur. C. Forecoxa. D. Foreclaw. E. Mid leg. F. Mid claw. G. Hind leg. H. Hind claw. Scale bars: A, E, G = 500 μm; B = 300 μm; C = 200 μm; D, F, H = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13745365" httpUri="https://zenodo.org/record/13745365/files/figure.png" pageId="2" pageNumber="503">Fig. 3E</figureCitation>
). Claw long, slightly bent apically, with 4 denticles increasing in size distally (
<figureCitation id="1367F584FFAFFFCDFA95F92CFF09F90D" captionStart="FIGURE 3" captionStartId="5.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-18@5.[158,1430,181,1752]" captionTargetPageId="5" captionText="FIGURE 3. Clypeocaenis laosensis sp. nov., A. Foreleg. B. High magnification view setae of transverse row forefemur. C. Forecoxa. D. Foreclaw. E. Mid leg. F. Mid claw. G. Hind leg. H. Hind claw. Scale bars: A, E, G = 500 μm; B = 300 μm; C = 200 μm; D, F, H = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13745365" httpUri="https://zenodo.org/record/13745365/files/figure.png" pageId="2" pageNumber="503">Fig. 3F</figureCitation>
). Hindleg: lengths of femur/ tibia/ tarsus/ claw viz., 0.50/ 0.30/ 0.28/
<quantity id="4CA444E4FFAFFFCDFC36F948FB82F90D" box="[972,1075,1749,1775]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="503" unit="mm" value="0.11">0.11 mm</quantity>
. Coxa similar to foreleg. Femur outer margin with a row of spatulate setae, inner margins proximal half with short spatulate setae. Tibia with a row of 1315 stout setae on inner margin, outer and submarginal inner margins with spatulate setae. Tarsus with a row of 810 stout setae on inner margin, outer margins with a few hair-like setae (
<figureCitation id="1367F584FFAFFFCDFC1FF8A2FB8EF8BB" box="[997,1087,1855,1881]" captionStart="FIGURE 3" captionStartId="5.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-18@5.[158,1430,181,1752]" captionTargetPageId="5" captionText="FIGURE 3. Clypeocaenis laosensis sp. nov., A. Foreleg. B. High magnification view setae of transverse row forefemur. C. Forecoxa. D. Foreclaw. E. Mid leg. F. Mid claw. G. Hind leg. H. Hind claw. Scale bars: A, E, G = 500 μm; B = 300 μm; C = 200 μm; D, F, H = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13745365" httpUri="https://zenodo.org/record/13745365/files/figure.png" pageId="2" pageNumber="503">Fig. 3G</figureCitation>
). Claw, similar to midleg, with 45 denticles, basal denticle minuscule (
<figureCitation id="1367F584FFAFFFCDFDA1F8FEFD04F89F" box="[603,693,1891,1917]" captionStart="FIGURE 3" captionStartId="5.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-18@5.[158,1430,181,1752]" captionTargetPageId="5" captionText="FIGURE 3. Clypeocaenis laosensis sp. nov., A. Foreleg. B. High magnification view setae of transverse row forefemur. C. Forecoxa. D. Foreclaw. E. Mid leg. F. Mid claw. G. Hind leg. H. Hind claw. Scale bars: A, E, G = 500 μm; B = 300 μm; C = 200 μm; D, F, H = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13745365" httpUri="https://zenodo.org/record/13745365/files/figure.png" pageId="2" pageNumber="503">Fig. 3H</figureCitation>
).
<emphasis id="B9283513FFAFFFCDFD32F8FEFCF7F89F" bold="true" box="[712,838,1891,1917]" pageId="2" pageNumber="503">Abdomen:</emphasis>
Terga I,
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, and
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IX with marked broad transverse dark brown bands. Posterolateral processes short, slight elongated segment
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(
<figureCitation id="1367F584FFAFFFCDFBF8F81BFBEDF842" box="[1026,1116,1926,1952]" captionStart="FIGURE 4" captionStartId="6.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-26@6.[158,1430,181,1752]" captionTargetPageId="6" captionText="FIGURE 4. Clypeocaesni laosensis sp. nov., A. Abdominal terga VIX. B. Tergum II. C. Sternum IX. D. Operculate gill, dorsal view. E. Higher magnification of the operculate gill, dorsal view. F. Spatulate setae on the medial ridge. G. Operculate gill, viewed from the ventral side. Scale bars: A, D, G = 500 μm; B = 100 μm; C, F = 200 μm; E = 50 μm." figureDoi="http://doi.org/10.5281/zenodo.13745367" httpUri="https://zenodo.org/record/13745367/files/figure.png" pageId="2" pageNumber="503">Fig. 4A</figureCitation>
). Tergum
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with a triangular posteromedian process, pronounced denticulation at posterior margin, surface with scales (
<figureCitation id="1367F584FFAFFFCDFB6CF837FB41F826" box="[1174,1264,1962,1988]" captionStart="FIGURE 4" captionStartId="6.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-26@6.[158,1430,181,1752]" captionTargetPageId="6" captionText="FIGURE 4. Clypeocaesni laosensis sp. nov., A. Abdominal terga VIX. B. Tergum II. C. Sternum IX. D. Operculate gill, dorsal view. E. Higher magnification of the operculate gill, dorsal view. F. Spatulate setae on the medial ridge. G. Operculate gill, viewed from the ventral side. Scale bars: A, D, G = 500 μm; B = 100 μm; C, F = 200 μm; E = 50 μm." figureDoi="http://doi.org/10.5281/zenodo.13745367" httpUri="https://zenodo.org/record/13745367/files/figure.png" pageId="2" pageNumber="503">Fig. 4B</figureCitation>
). Terga
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IX with strongly denticulate posterior margin, very small spatulate setae on tergum
<collectionCode id="ED4D71C4FFAFFFCDFBE0F853FBF0F80A" box="[1050,1089,1998,2024]" pageId="2" pageNumber="503">VII</collectionCode>
(
<figureCitation id="1367F584FFAFFFCDFBAAF850FB1AF805" box="[1104,1195,1997,2023]" captionStart="FIGURE 4" captionStartId="6.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-26@6.[158,1430,181,1752]" captionTargetPageId="6" captionText="FIGURE 4. Clypeocaesni laosensis sp. nov., A. Abdominal terga VIX. B. Tergum II. C. Sternum IX. D. Operculate gill, dorsal view. E. Higher magnification of the operculate gill, dorsal view. F. Spatulate setae on the medial ridge. G. Operculate gill, viewed from the ventral side. Scale bars: A, D, G = 500 μm; B = 100 μm; C, F = 200 μm; E = 50 μm." figureDoi="http://doi.org/10.5281/zenodo.13745367" httpUri="https://zenodo.org/record/13745367/files/figure.png" pageId="2" pageNumber="503">Fig. 4A</figureCitation>
). Sternum IX slightly concave hind margin (
<figureCitation id="1367F584FFAEFFCCFE68FF0AFE5BFF53" box="[402,490,151,177]" captionStart="FIGURE 4" captionStartId="6.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-26@6.[158,1430,181,1752]" captionTargetPageId="6" captionText="FIGURE 4. Clypeocaesni laosensis sp. nov., A. Abdominal terga VIX. B. Tergum II. C. Sternum IX. D. Operculate gill, dorsal view. E. Higher magnification of the operculate gill, dorsal view. F. Spatulate setae on the medial ridge. G. Operculate gill, viewed from the ventral side. Scale bars: A, D, G = 500 μm; B = 100 μm; C, F = 200 μm; E = 50 μm." figureDoi="http://doi.org/10.5281/zenodo.13745367" httpUri="https://zenodo.org/record/13745367/files/figure.png" pageId="3" pageNumber="504">Fig. 4C</figureCitation>
), shagreen field on dorsal surface with 4 irregular rows of small denticles parallel to the hind margin. Gill I short, two segments, length one-third of gill cover. Operculate gill quadrate, with the dorsal side covered with knobs and scattered long, leaf-like, pine-needle-shaped microtrichia as described in the section on the thorax (
<figureCitation id="1367F584FFAEFFCCFEC2FE9CFE21FEF9" box="[312,400,257,283]" captionStart="FIGURE 4" captionStartId="6.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-26@6.[158,1430,181,1752]" captionTargetPageId="6" captionText="FIGURE 4. Clypeocaesni laosensis sp. nov., A. Abdominal terga VIX. B. Tergum II. C. Sternum IX. D. Operculate gill, dorsal view. E. Higher magnification of the operculate gill, dorsal view. F. Spatulate setae on the medial ridge. G. Operculate gill, viewed from the ventral side. Scale bars: A, D, G = 500 μm; B = 100 μm; C, F = 200 μm; E = 50 μm." figureDoi="http://doi.org/10.5281/zenodo.13745367" httpUri="https://zenodo.org/record/13745367/files/figure.png" pageId="3" pageNumber="504">Fig. 4E</figureCitation>
). A strongly developed medial ridge forms an elevated keel, basally with 810 stout spatulate setae (
<figureCitation id="1367F584FFAEFFCCFF18FEB9FE8AFEDD" box="[226,315,292,319]" captionStart="FIGURE 4" captionStartId="6.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-26@6.[158,1430,181,1752]" captionTargetPageId="6" captionText="FIGURE 4. Clypeocaesni laosensis sp. nov., A. Abdominal terga VIX. B. Tergum II. C. Sternum IX. D. Operculate gill, dorsal view. E. Higher magnification of the operculate gill, dorsal view. F. Spatulate setae on the medial ridge. G. Operculate gill, viewed from the ventral side. Scale bars: A, D, G = 500 μm; B = 100 μm; C, F = 200 μm; E = 50 μm." figureDoi="http://doi.org/10.5281/zenodo.13745367" httpUri="https://zenodo.org/record/13745367/files/figure.png" pageId="3" pageNumber="504">Fig. 4F</figureCitation>
). The outer ridge narrow and inconspicuous, featuring a row of very small denticles. The lateral margin adorned with short setae, the inner margin with moderate-length thin bipinnate setae, and the posterior margin with very short, broadly spatulate setae (
<figureCitation id="1367F584FFAEFFCCFD46FEF6FC8AFE64" box="[700,827,363,390]" captionStart="FIGURE 4" captionStartId="6.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-26@6.[158,1430,181,1752]" captionTargetPageId="6" captionText="FIGURE 4. Clypeocaesni laosensis sp. nov., A. Abdominal terga VIX. B. Tergum II. C. Sternum IX. D. Operculate gill, dorsal view. E. Higher magnification of the operculate gill, dorsal view. F. Spatulate setae on the medial ridge. G. Operculate gill, viewed from the ventral side. Scale bars: A, D, G = 500 μm; B = 100 μm; C, F = 200 μm; E = 50 μm." figureDoi="http://doi.org/10.5281/zenodo.13745367" httpUri="https://zenodo.org/record/13745367/files/figure.png" pageId="3" pageNumber="504">Fig. 4D, G</figureCitation>
). The microtrichia band on the ventral side of the gill cover absent (
<figureCitation id="1367F584FFAEFFCCFECBFE12FE25FE4B" box="[305,404,399,425]" captionStart="FIGURE 4" captionStartId="6.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-26@6.[158,1430,181,1752]" captionTargetPageId="6" captionText="FIGURE 4. Clypeocaesni laosensis sp. nov., A. Abdominal terga VIX. B. Tergum II. C. Sternum IX. D. Operculate gill, dorsal view. E. Higher magnification of the operculate gill, dorsal view. F. Spatulate setae on the medial ridge. G. Operculate gill, viewed from the ventral side. Scale bars: A, D, G = 500 μm; B = 100 μm; C, F = 200 μm; E = 50 μm." figureDoi="http://doi.org/10.5281/zenodo.13745367" httpUri="https://zenodo.org/record/13745367/files/figure.png" pageId="3" pageNumber="504">Figs. 4G</figureCitation>
,
<figureCitation id="1367F584FFAEFFCCFE64FE12FE7FFE4B" box="[414,462,399,425]" captionStart="FIGURE 12" captionStartId="18.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-26@18.[158,1430,181,1752]" captionTargetPageId="18" captionText="FIGURE 12. Microtrichia bands on the ventral side of the operculate gill, as previously documented. A. Clypeocaenis soldani. B. Clypeocaenis multisetosa. C. Clypeocaenis oligosetosa. D. Clypeocaenis kaveri. E. Clypeocaenis afrosetosa. F. Clypeocaenis laosensis sp. nov. G. Mekongellina sangpradubae." figureDoi="http://doi.org/10.5281/zenodo.13745395" httpUri="https://zenodo.org/record/13745395/files/figure.png" pageId="3" pageNumber="504">12F</figureCitation>
). Gill III possesses a single filament with three branches and 3035 filaments, each with simple or bifurcated branches (
<figureCitation id="1367F584FFAEFFCCFE09FE2FFDFCFE2F" box="[499,589,434,461]" captionStart="FIGURE 5" captionStartId="7.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-18@7.[158,1430,181,1752]" captionTargetPageId="7" captionText="FIGURE 5. Clpeocaenis laosensis sp. nov., A. Gills III, IV. B. Egg, arrow indicate the micropyle. C. Epithema. D. Detail of the micropyle. Scale bars: A = 500 μm; B = 100 μm; C = 40 μm; D = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.13745371" httpUri="https://zenodo.org/record/13745371/files/figure.png" pageId="3" pageNumber="504">Fig. 5A</figureCitation>
). The cerci translucent with whorls of setae on each segment.
</paragraph>
<paragraph id="8BE3E901FFAEFFCCFF3DFE4BFAAAFDDA" blockId="3.[151,1437,151,605]" pageId="3" pageNumber="504">
<emphasis id="B9283513FFAEFFCCFF3DFE4BFF4EFE12" bold="true" box="[199,255,470,496]" pageId="3" pageNumber="504">Egg:</emphasis>
Oval shape (
<figureCitation id="1367F584FFAEFFCCFE6BFE4BFE59FE13" box="[401,488,470,497]" captionStart="FIGURE 5" captionStartId="7.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-18@7.[158,1430,181,1752]" captionTargetPageId="7" captionText="FIGURE 5. Clpeocaenis laosensis sp. nov., A. Gills III, IV. B. Egg, arrow indicate the micropyle. C. Epithema. D. Detail of the micropyle. Scale bars: A = 500 μm; B = 100 μm; C = 40 μm; D = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.13745371" httpUri="https://zenodo.org/record/13745371/files/figure.png" pageId="3" pageNumber="504">Fig. 5B</figureCitation>
), length 121133 μm, 92100 μm width. Chorion smooth with a single epithema that forms a cap consisting of fine threads, terminating in numerous very small knobs (
<figureCitation id="1367F584FFAEFFCCFBF0FE67FBD1FDF7" box="[1034,1120,506,533]" captionStart="FIGURE 5" captionStartId="7.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-18@7.[158,1430,181,1752]" captionTargetPageId="7" captionText="FIGURE 5. Clpeocaenis laosensis sp. nov., A. Gills III, IV. B. Egg, arrow indicate the micropyle. C. Epithema. D. Detail of the micropyle. Scale bars: A = 500 μm; B = 100 μm; C = 40 μm; D = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.13745371" httpUri="https://zenodo.org/record/13745371/files/figure.png" pageId="3" pageNumber="504">Fig.5C</figureCitation>
). Positioned subequatorially, micropyle moderate length, mouth not broadened and encircled by a circular sperm-guide (
<figureCitation id="1367F584FFAEFFCCFB69FD83FABFFDDA" box="[1171,1294,542,569]" captionStart="FIGURE 5" captionStartId="7.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-18@7.[158,1430,181,1752]" captionTargetPageId="7" captionText="FIGURE 5. Clpeocaenis laosensis sp. nov., A. Gills III, IV. B. Egg, arrow indicate the micropyle. C. Epithema. D. Detail of the micropyle. Scale bars: A = 500 μm; B = 100 μm; C = 40 μm; D = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.13745371" httpUri="https://zenodo.org/record/13745371/files/figure.png" pageId="3" pageNumber="504">Fig. 5B, D</figureCitation>
).
</paragraph>
<paragraph id="8BE3E901FFAEFFCCFF3DFDDEFE2BFDBE" blockId="3.[151,1437,151,605]" box="[199,410,578,605]" pageId="3" pageNumber="504">
<emphasis id="B9283513FFAEFFCCFF3DFDDEFEACFDBF" bold="true" box="[199,285,579,605]" pageId="3" pageNumber="504">Imago:</emphasis>
Unknown.
</paragraph>
</subSubSection>
<caption id="DF23B989FFAEFFCCFF6DF845FB0BF812" ID-DOI="http://doi.org/10.5281/zenodo.13745357" ID-Zenodo-Dep="13745357" box="[151,1210,2008,2033]" httpUri="https://zenodo.org/record/13745357/files/figure.png" pageId="3" pageNumber="504" startId="3.[151,250,2008,2032]" targetBox="[261,1327,626,1982]" targetPageId="3" targetType="figure">
<paragraph id="8BE3E901FFAEFFCCFF6DF845FB0BF812" blockId="3.[151,1210,2008,2033]" box="[151,1210,2008,2033]" pageId="3" pageNumber="504">
<emphasis id="B9283513FFAEFFCCFF6DF845FEA2F812" bold="true" box="[151,275,2008,2032]" pageId="3" pageNumber="504">FIGURE 1.</emphasis>
<taxonomicName id="4C5C9282FFAEFFCCFEE3F845FDB4F812" authorityName="Vannachak &amp; Malzacher &amp; Sangpradub" authorityYear="2024" box="[281,517,2008,2032]" class="Insecta" family="Caenidae" genus="Clypeocaenis" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="laosensis" status="sp. nov.">
<emphasis id="B9283513FFAEFFCCFEE3F845FDB4F812" box="[281,517,2008,2032]" italics="true" pageId="3" pageNumber="504">Clypeocaenis laosensis</emphasis>
</taxonomicName>
<emphasis id="B9283513FFAEFFCCFDF1F844FDEDF813" bold="true" box="[523,604,2009,2033]" pageId="3" pageNumber="504">
<taxonomicNameLabel id="A21B8868FFAEFFCCFDF1F844FDEDF813" box="[523,604,2009,2033]" pageId="3" pageNumber="504" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, habitus.
<emphasis id="B9283513FFAEFFCCFD44F845FD67F812" bold="true" box="[702,726,2008,2032]" pageId="3" pageNumber="504">A.</emphasis>
Female larva.
<emphasis id="B9283513FFAEFFCCFC97F844FC35F813" bold="true" box="[877,900,2009,2033]" pageId="3" pageNumber="504">B.</emphasis>
Male larva. Scale bars: 1 mm.
</paragraph>
</caption>
<caption id="DF23B989FFA9FFCBFF6DF96DFBF9F8CF" ID-DOI="http://doi.org/10.5281/zenodo.13745363" ID-Zenodo-Dep="13745363" httpUri="https://zenodo.org/record/13745363/files/figure.png" pageId="4" pageNumber="505" startId="4.[151,250,1776,1800]" targetBox="[158,1430,181,1752]" targetPageId="4" targetType="figure">
<paragraph id="8BE3E901FFA9FFCBFF6DF96DFBF9F8CF" blockId="4.[151,1436,1776,1837]" pageId="4" pageNumber="505">
<emphasis id="B9283513FFA9FFCBFF6DF96DFEA4F8EA" bold="true" box="[151,277,1776,1800]" pageId="4" pageNumber="505">FIGURE 2.</emphasis>
<taxonomicName id="4C5C9282FFA9FFCBFEE4F96DFDBDF8EA" authorityName="Vannachak &amp; Malzacher &amp; Sangpradub" authorityYear="2024" box="[286,524,1776,1800]" class="Insecta" family="Caenidae" genus="Clypeocaenis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="laosensis" status="sp. nov.">
<emphasis id="B9283513FFA9FFCBFEE4F96DFDBDF8EA" box="[286,524,1776,1800]" italics="true" pageId="4" pageNumber="505">Clypeocaenis laosensis</emphasis>
</taxonomicName>
<emphasis id="B9283513FFA9FFCBFDEEF96DFDD6F8EA" bold="true" box="[532,615,1776,1800]" pageId="4" pageNumber="505">
<taxonomicNameLabel id="A21B8868FFA9FFCBFDEEF96DFDD6F8EA" box="[532,615,1776,1800]" pageId="4" pageNumber="505" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="B9283513FFA9FFCBFD8CF96DFD3EF8EA" bold="true" box="[630,655,1776,1800]" pageId="4" pageNumber="505">A.</emphasis>
Head capsule, viewed dorsally.
<emphasis id="B9283513FFA9FFCBFC1BF96DFC49F8EA" bold="true" box="[993,1016,1776,1800]" pageId="4" pageNumber="505">B.</emphasis>
Labrum.
<emphasis id="B9283513FFA9FFCBFB9AF96DFBC9F8EA" bold="true" box="[1120,1144,1776,1800]" pageId="4" pageNumber="505">C.</emphasis>
Mandibles.
<emphasis id="B9283513FFA9FFCBFB01F96DFAA5F8EA" bold="true" box="[1275,1300,1776,1800]" pageId="4" pageNumber="505">D.</emphasis>
Maxillae.
<emphasis id="B9283513FFA9FFCBFA7FF96DFA2DF8EA" bold="true" box="[1413,1436,1776,1800]" pageId="4" pageNumber="505">E.</emphasis>
Hypopharynx.
<emphasis id="B9283513FFA9FFCBFED5F889FEF3F8CE" bold="true" box="[303,322,1812,1836]" pageId="4" pageNumber="505">F.</emphasis>
Labium. Scale bars: A = 500 μm; BD = 200 μm; E = 100 μm; F = 300 μm.
</paragraph>
</caption>
<caption id="DF23B989FFA8FFCAFF6DF96DFE1DF8B3" ID-DOI="http://doi.org/10.5281/zenodo.13745365" ID-Zenodo-Dep="13745365" httpUri="https://zenodo.org/record/13745365/files/figure.png" pageId="5" pageNumber="506" startId="5.[151,250,1776,1800]" targetBox="[158,1430,181,1752]" targetPageId="5" targetType="figure">
<paragraph id="8BE3E901FFA8FFCAFF6DF96DFE1DF8B3" blockId="5.[151,1436,1776,1873]" pageId="5" pageNumber="506">
<emphasis id="B9283513FFA8FFCAFF6DF96DFEA4F8EA" bold="true" box="[151,277,1776,1800]" pageId="5" pageNumber="506">FIGURE 3.</emphasis>
<taxonomicName id="4C5C9282FFA8FFCAFEE7F96DFDBAF8EA" authorityName="Vannachak &amp; Malzacher &amp; Sangpradub" authorityYear="2024" box="[285,523,1776,1800]" class="Insecta" family="Caenidae" genus="Clypeocaenis" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="laosensis" status="sp. nov.">
<emphasis id="B9283513FFA8FFCAFEE7F96DFDBAF8EA" box="[285,523,1776,1800]" italics="true" pageId="5" pageNumber="506">Clypeocaenis laosensis</emphasis>
</taxonomicName>
<emphasis id="B9283513FFA8FFCAFDE9F96DFDD7F8EA" bold="true" box="[531,614,1776,1800]" pageId="5" pageNumber="506">
<taxonomicNameLabel id="A21B8868FFA8FFCAFDE9F96DFDD7F8EA" box="[531,614,1776,1800]" pageId="5" pageNumber="506" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="B9283513FFA8FFCAFD8FF96DFD3CF8EA" bold="true" box="[629,653,1776,1800]" pageId="5" pageNumber="506">A.</emphasis>
Foreleg.
<emphasis id="B9283513FFA8FFCAFD0BF96DFCB8F8EA" bold="true" box="[753,777,1776,1800]" pageId="5" pageNumber="506">B.</emphasis>
High magnification view setae of transverse row forefemur.
<emphasis id="B9283513FFA8FFCAFA7EF96DFA2DF8EA" bold="true" box="[1412,1436,1776,1800]" pageId="5" pageNumber="506">C.</emphasis>
Forecoxa.
<emphasis id="B9283513FFA8FFCAFEF9F889FEADF8CE" bold="true" box="[259,284,1812,1836]" pageId="5" pageNumber="506">D.</emphasis>
Foreclaw.
<emphasis id="B9283513FFA8FFCAFE77F889FE15F8CE" bold="true" box="[397,420,1812,1836]" pageId="5" pageNumber="506">E.</emphasis>
Mid leg.
<emphasis id="B9283513FFA8FFCAFDF3F889FDADF8CE" bold="true" box="[521,540,1812,1836]" pageId="5" pageNumber="506">F.</emphasis>
Mid claw.
<emphasis id="B9283513FFA8FFCAFD6BF889FD1AF8CE" bold="true" box="[657,683,1812,1836]" pageId="5" pageNumber="506">G.</emphasis>
Hind leg.
<emphasis id="B9283513FFA8FFCAFCE2F889FC83F8CE" bold="true" box="[792,818,1812,1836]" pageId="5" pageNumber="506">H.</emphasis>
Hind claw. Scale bars: A, E, G = 500 μm; B = 300 μm; C = 200 μm; D, F, H = 100 μm.
</paragraph>
</caption>
<caption id="DF23B989FFABFFC9FF6DF96DFB2BF8B3" ID-DOI="http://doi.org/10.5281/zenodo.13745367" ID-Zenodo-Dep="13745367" httpUri="https://zenodo.org/record/13745367/files/figure.png" pageId="6" pageNumber="507" startId="6.[151,250,1776,1800]" targetBox="[158,1430,181,1752]" targetPageId="6" targetType="figure">
<paragraph id="8BE3E901FFABFFC9FF6DF96DFB2BF8B3" blockId="6.[151,1436,1776,1873]" pageId="6" pageNumber="507">
<emphasis id="B9283513FFABFFC9FF6DF96DFEA3F8EA" bold="true" box="[151,274,1776,1800]" pageId="6" pageNumber="507">FIGURE 4.</emphasis>
<taxonomicName id="4C5C9282FFABFFC9FEEDF96DFDB3F8EA" authorityName="Vannachak &amp; Malzacher &amp; Sangpradub" authorityYear="2024" box="[279,514,1776,1800]" class="Insecta" family="Caenidae" genus="Clypeocaesni" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="507" phylum="Arthropoda" rank="species" species="laosensis" status="sp. nov.">
<emphasis id="B9283513FFABFFC9FEEDF96DFDB3F8EA" box="[279,514,1776,1800]" italics="true" pageId="6" pageNumber="507">Clypeocaesni laosensis</emphasis>
</taxonomicName>
<emphasis id="B9283513FFABFFC9FDFDF96DFDE7F8EA" bold="true" box="[519,598,1776,1800]" pageId="6" pageNumber="507">
<taxonomicNameLabel id="A21B8868FFABFFC9FDFDF96DFDE7F8EA" box="[519,598,1776,1800]" pageId="6" pageNumber="507" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="B9283513FFABFFC9FD98F96DFDCAF8EA" bold="true" box="[610,635,1776,1800]" pageId="6" pageNumber="507">A.</emphasis>
Abdominal terga VIX.
<emphasis id="B9283513FFABFFC9FC8EF96DFC3AF8EA" bold="true" box="[884,907,1776,1800]" pageId="6" pageNumber="507">B.</emphasis>
Tergum II.
<emphasis id="B9283513FFABFFC9FC05F96DFBA9F8EA" bold="true" box="[1023,1048,1776,1800]" pageId="6" pageNumber="507">C.</emphasis>
Sternum IX.
<emphasis id="B9283513FFABFFC9FB65F96DFB06F8EA" bold="true" box="[1183,1207,1776,1800]" pageId="6" pageNumber="507">D.</emphasis>
Operculate gill, dorsal view.
<emphasis id="B9283513FFABFFC9FF2FF889FF5CF8CE" bold="true" box="[213,237,1812,1836]" pageId="6" pageNumber="507">E.</emphasis>
Higher magnification of the operculate gill, dorsal view.
<emphasis id="B9283513FFABFFC9FCBEF889FCE6F8CE" bold="true" box="[836,855,1812,1836]" pageId="6" pageNumber="507">F.</emphasis>
Spatulate setae on the medial ridge.
<emphasis id="B9283513FFABFFC9FB20F889FB45F8CE" bold="true" box="[1242,1268,1812,1836]" pageId="6" pageNumber="507">G.</emphasis>
Operculate gill, viewed from the ventral side. Scale bars: A, D, G = 500 μm; B = 100 μm; C, F = 200 μm; E = 50 μm.
</paragraph>
</caption>
<caption id="DF23B989FFAAFFC8FF6DF96DFCCAF8CF" ID-DOI="http://doi.org/10.5281/zenodo.13745371" ID-Zenodo-Dep="13745371" httpUri="https://zenodo.org/record/13745371/files/figure.png" pageId="7" pageNumber="508" startId="7.[151,250,1776,1800]" targetBox="[158,1430,181,1752]" targetPageId="7" targetType="figure">
<paragraph id="8BE3E901FFAAFFC8FF6DF96DFCCAF8CF" blockId="7.[151,1436,1776,1837]" pageId="7" pageNumber="508">
<emphasis id="B9283513FFAAFFC8FF6DF96DFEA3F8EA" bold="true" box="[151,274,1776,1800]" pageId="7" pageNumber="508">FIGURE 5.</emphasis>
<taxonomicName id="4C5C9282FFAAFFC8FEE2F96DFE49F8EA" authorityName="Vannachak &amp; Malzacher &amp; Sangpradub" authorityYear="2024" box="[280,504,1776,1800]" class="Insecta" family="Caenidae" genus="Clpeocaenis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="laosensis" status="sp. nov.">
<emphasis id="B9283513FFAAFFC8FEE2F96DFE49F8EA" box="[280,504,1776,1800]" italics="true" pageId="7" pageNumber="508">Clpeocaenis laosensis</emphasis>
</taxonomicName>
<emphasis id="B9283513FFAAFFC8FE07F96DFDFCF8EA" bold="true" box="[509,589,1776,1800]" pageId="7" pageNumber="508">
<taxonomicNameLabel id="A21B8868FFAAFFC8FE07F96DFDFCF8EA" box="[509,589,1776,1800]" pageId="7" pageNumber="508" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="B9283513FFAAFFC8FDA3F96DFDC3F8EA" bold="true" box="[601,626,1776,1800]" pageId="7" pageNumber="508">A.</emphasis>
Gills III, IV.
<emphasis id="B9283513FFAAFFC8FD0DF96DFCBFF8EA" bold="true" box="[759,782,1776,1800]" pageId="7" pageNumber="508">B.</emphasis>
Egg, arrow indicate the micropyle.
<emphasis id="B9283513FFAAFFC8FB8CF96DFB3FF8EA" bold="true" box="[1142,1166,1776,1800]" pageId="7" pageNumber="508">C.</emphasis>
Epithema.
<emphasis id="B9283513FFAAFFC8FAFBF96DFAABF8EA" bold="true" box="[1281,1306,1776,1800]" pageId="7" pageNumber="508">D.</emphasis>
Detail of the micropyle. Scale bars: A = 500 μm; B = 100 μm; C = 40 μm; D = 30 μm.
</paragraph>
</caption>
<subSubSection id="C346BA8AFFAAFFC7FF3DF8E3FD5AFDDF" lastPageId="8" lastPageNumber="509" pageId="7" pageNumber="508" type="diagnosis">
<paragraph id="8BE3E901FFAAFFC7FF3DF8E3FD5AFDDF" blockId="7.[151,1436,1918,2017]" lastBlockId="8.[151,1437,151,573]" lastPageId="8" lastPageNumber="509" pageId="7" pageNumber="508">
<emphasis id="B9283513FFAAFFC8FF3DF8E3FEF3F87A" bold="true" box="[199,322,1918,1944]" pageId="7" pageNumber="508">Diagnosis.</emphasis>
<taxonomicName id="4C5C9282FFAAFFC8FEB1F8E3FDE3F87A" authorityName="Vannachak &amp; Malzacher &amp; Sangpradub" authorityYear="2024" box="[331,594,1918,1944]" class="Insecta" family="Caenidae" genus="Clypeocaenis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="laosensis" status="sp. nov.">
<emphasis id="B9283513FFAAFFC8FEB1F8E3FDE3F87A" box="[331,594,1918,1944]" italics="true" pageId="7" pageNumber="508">Clypeocaenis laosensis</emphasis>
</taxonomicName>
<emphasis id="B9283513FFAAFFC8FDA1F8E3FD06F87A" bold="true" box="[603,695,1918,1944]" pageId="7" pageNumber="508">
<taxonomicNameLabel id="A21B8868FFAAFFC8FDA1F8E3FD06F87A" box="[603,695,1918,1944]" pageId="7" pageNumber="508" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
exhibits distinctive morphological characteristics, and is closely related to
<taxonomicName id="4C5C9282FFAAFFC8FEF8F83FFCF9F85E" authority="Balasubramanian &amp; Muthukatturaja, 2021" authorityName="Balasubramanian &amp; Muthukatturaja" authorityYear="2021" box="[258,840,1954,1980]" class="Insecta" family="Caenidae" genus="Clypeocaenis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="kaveri">
<emphasis id="B9283513FFAAFFC8FEF8F83FFEDEF85E" box="[258,367,1954,1980]" italics="true" pageId="7" pageNumber="508">Cl. kaveri</emphasis>
<bibRefCitation id="EFCD94F0FFAAFFC8FE89F83FFCF9F85E" author="Balasubramanian, C. &amp; Muthukatturaja, M." box="[371,840,1954,1980]" pageId="7" pageNumber="508" pagination="377 - 388" refId="ref9722" refString="Balasubramanian, C. &amp; Muthukatturaja, M. (2021) Two additional new species of Clypeocaenis Soldan, 1978 (Ephemeroptera: Caenidae) from the Western Ghats of Peninsular India. Zootaxa, 4915 (3), 377 - 388. https: // doi. org / 10.11646 / zootaxa. 4915.3.6" type="journal article" year="2021">Balasubramanian &amp; Muthukatturaja, 2021</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C5C9282FFAAFFC8FC84F83FFF7EF802" authority="Balasubramanian &amp; Muthukatturaja, 2020" authorityName="Balasubramanian &amp; Muthukatturaja" authorityYear="2020" class="Insecta" family="Caenidae" genus="Clypeocaenis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="gayathri">
<emphasis id="B9283513FFAAFFC8FC84F83FFBB2F85E" box="[894,1027,1954,1980]" italics="true" pageId="7" pageNumber="508">Cl. gayathri</emphasis>
Balasubramanian &amp;
<bibRefCitation id="EFCD94F0FFAAFFC8FB13F83FFF7EF802" author="Muthukatturaja, M. &amp; Balasubramanian, C. &amp; Murugan, A." pageId="7" pageNumber="508" pagination="361 - 370" refId="ref10400" refString="Muthukatturaja, M., Balasubramanian, C. &amp; Murugan, A. (2020) Two new species of Clypeocaenis (Ephemeroptera: Caenidae) from Western Ghats, Southern India. Zootaxa, 4722 (4), 361 - 370. https: // doi. org / 10.11646 / zootaxa. 4722.4.5" type="journal article" year="2020">Muthukatturaja, 2020</bibRefCitation>
</taxonomicName>
(
<tableCitation id="C6DEDCBAFFAAFFC8FF1AF85BFE86F802" box="[224,311,1990,2016]" captionStart="TABLE 1" captionStartId="8.[152,240,620,644]" captionText="TABLE 1. Comparative morphological characteristics of Clypeocaenis laosensis sp. nov. larvae and closely related species." pageId="7" pageNumber="508">Table 1</tableCitation>
). The following combination of features distinguishes this new species from its congeners: i) the pedicel is one and a half times the length of the third antennal segment (
<figureCitation id="1367F584FFA5FFC7FC21FF0AFB86FF53" box="[987,1079,151,177]" captionStart="FIGURE 2" captionStartId="4.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-26@4.[158,1430,181,1752]" captionTargetPageId="4" captionText="FIGURE 2. Clypeocaenis laosensis sp. nov., A. Head capsule, viewed dorsally. B. Labrum. C. Mandibles. D. Maxillae. E. Hypopharynx. F. Labium. Scale bars: A = 500 μm; BD = 200 μm; E = 100 μm; F = 300 μm." figureDoi="http://doi.org/10.5281/zenodo.13745363" httpUri="https://zenodo.org/record/13745363/files/figure.png" pageId="8" pageNumber="509">Fig. 2A</figureCitation>
); ii) below the molar, there are 45 thin setae; iii) the forefemur, at the middle of the inner margin, is equipped with three long filtering setae; the outer margin features short, simple setae, and the dorsal surface presents a transverse row of large, broadly spatulate setae (
<figureCitation id="1367F584FFA5FFC7FF1AFE9EFEEDFEFF" box="[224,348,259,285]" captionStart="FIGURE 3" captionStartId="5.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-18@5.[158,1430,181,1752]" captionTargetPageId="5" captionText="FIGURE 3. Clypeocaenis laosensis sp. nov., A. Foreleg. B. High magnification view setae of transverse row forefemur. C. Forecoxa. D. Foreclaw. E. Mid leg. F. Mid claw. G. Hind leg. H. Hind claw. Scale bars: A, E, G = 500 μm; B = 300 μm; C = 200 μm; D, F, H = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13745365" httpUri="https://zenodo.org/record/13745365/files/figure.png" pageId="8" pageNumber="509">Fig. 3A, B</figureCitation>
); iv) the mid and hind femora are adorned with short, marginal bristles that are basally strongly narrowed (
<figureCitation id="1367F584FFA5FFC7FEE9FEBAFE20FEA3" box="[275,401,295,321]" captionStart="FIGURE 3" captionStartId="5.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-18@5.[158,1430,181,1752]" captionTargetPageId="5" captionText="FIGURE 3. Clypeocaenis laosensis sp. nov., A. Foreleg. B. High magnification view setae of transverse row forefemur. C. Forecoxa. D. Foreclaw. E. Mid leg. F. Mid claw. G. Hind leg. H. Hind claw. Scale bars: A, E, G = 500 μm; B = 300 μm; C = 200 μm; D, F, H = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13745365" httpUri="https://zenodo.org/record/13745365/files/figure.png" pageId="8" pageNumber="509">Fig. 3E, G</figureCitation>
); v) tarsal claws are long, slender, and slightly bent apically, with 35 denticles (
<figureCitation id="1367F584FFA5FFC7FAC6FEBAFF7CFE87" captionStart="FIGURE 3" captionStartId="5.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-18@5.[158,1430,181,1752]" captionTargetPageId="5" captionText="FIGURE 3. Clypeocaenis laosensis sp. nov., A. Foreleg. B. High magnification view setae of transverse row forefemur. C. Forecoxa. D. Foreclaw. E. Mid leg. F. Mid claw. G. Hind leg. H. Hind claw. Scale bars: A, E, G = 500 μm; B = 300 μm; C = 200 μm; D, F, H = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13745365" httpUri="https://zenodo.org/record/13745365/files/figure.png" pageId="8" pageNumber="509">Fig. 3D, F, H</figureCitation>
); vi) the posterior margin of abdominal tergum VII is strongly denticulate and bears small spatulate setae, whereas terga VIIIIX are only denticulated (
<figureCitation id="1367F584FFA5FFC7FD6DFEF2FD42FE6B" box="[663,755,367,393]" captionStart="FIGURE 4" captionStartId="6.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-26@6.[158,1430,181,1752]" captionTargetPageId="6" captionText="FIGURE 4. Clypeocaesni laosensis sp. nov., A. Abdominal terga VIX. B. Tergum II. C. Sternum IX. D. Operculate gill, dorsal view. E. Higher magnification of the operculate gill, dorsal view. F. Spatulate setae on the medial ridge. G. Operculate gill, viewed from the ventral side. Scale bars: A, D, G = 500 μm; B = 100 μm; C, F = 200 μm; E = 50 μm." figureDoi="http://doi.org/10.5281/zenodo.13745367" httpUri="https://zenodo.org/record/13745367/files/figure.png" pageId="8" pageNumber="509">Fig. 4A</figureCitation>
); vii) the operculate gill features short setae along the lateral margin, thin bipinnate setae along the inner margin, and very short spatulate setae along the posterior margin (
<figureCitation id="1367F584FFA5FFC7FA95FE0EFF51FE33" captionStart="FIGURE 4" captionStartId="6.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-26@6.[158,1430,181,1752]" captionTargetPageId="6" captionText="FIGURE 4. Clypeocaesni laosensis sp. nov., A. Abdominal terga VIX. B. Tergum II. C. Sternum IX. D. Operculate gill, dorsal view. E. Higher magnification of the operculate gill, dorsal view. F. Spatulate setae on the medial ridge. G. Operculate gill, viewed from the ventral side. Scale bars: A, D, G = 500 μm; B = 100 μm; C, F = 200 μm; E = 50 μm." figureDoi="http://doi.org/10.5281/zenodo.13745367" httpUri="https://zenodo.org/record/13745367/files/figure.png" pageId="8" pageNumber="509">Fig. 4D, G</figureCitation>
); the middle ridge is notably developed with 810 stout spatulate setae (
<figureCitation id="1367F584FFA5FFC7FBF7FE2AFBD2FE33" box="[1037,1123,439,465]" captionStart="FIGURE 4" captionStartId="6.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-26@6.[158,1430,181,1752]" captionTargetPageId="6" captionText="FIGURE 4. Clypeocaesni laosensis sp. nov., A. Abdominal terga VIX. B. Tergum II. C. Sternum IX. D. Operculate gill, dorsal view. E. Higher magnification of the operculate gill, dorsal view. F. Spatulate setae on the medial ridge. G. Operculate gill, viewed from the ventral side. Scale bars: A, D, G = 500 μm; B = 100 μm; C, F = 200 μm; E = 50 μm." figureDoi="http://doi.org/10.5281/zenodo.13745367" httpUri="https://zenodo.org/record/13745367/files/figure.png" pageId="8" pageNumber="509">Fig. 4F</figureCitation>
); viii) the operculate gill on the ventral side lacks a microtrichia band (
<figureCitation id="1367F584FFA5FFC7FD89FE46FD66FE17" box="[627,727,475,501]" captionStart="FIGURE 4" captionStartId="6.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-26@6.[158,1430,181,1752]" captionTargetPageId="6" captionText="FIGURE 4. Clypeocaesni laosensis sp. nov., A. Abdominal terga VIX. B. Tergum II. C. Sternum IX. D. Operculate gill, dorsal view. E. Higher magnification of the operculate gill, dorsal view. F. Spatulate setae on the medial ridge. G. Operculate gill, viewed from the ventral side. Scale bars: A, D, G = 500 μm; B = 100 μm; C, F = 200 μm; E = 50 μm." figureDoi="http://doi.org/10.5281/zenodo.13745367" httpUri="https://zenodo.org/record/13745367/files/figure.png" pageId="8" pageNumber="509">Figs. 4G</figureCitation>
,
<figureCitation id="1367F584FFA5FFC7FD19FE46FCA3FE17" box="[739,786,475,501]" captionStart="FIGURE 12" captionStartId="18.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-26@18.[158,1430,181,1752]" captionTargetPageId="18" captionText="FIGURE 12. Microtrichia bands on the ventral side of the operculate gill, as previously documented. A. Clypeocaenis soldani. B. Clypeocaenis multisetosa. C. Clypeocaenis oligosetosa. D. Clypeocaenis kaveri. E. Clypeocaenis afrosetosa. F. Clypeocaenis laosensis sp. nov. G. Mekongellina sangpradubae." figureDoi="http://doi.org/10.5281/zenodo.13745395" httpUri="https://zenodo.org/record/13745395/files/figure.png" pageId="8" pageNumber="509">12F</figureCitation>
); ix) the eggs epithema is cap-shaped, constituted by fine threads terminating in numerous small knobs; the micropyle is of moderate length, the mouth is not broadened, and it is encircled by a circular sperm-guide (
<figureCitation id="1367F584FFA5FFC7FD99FDBEFD6EFDDF" box="[611,735,546,573]" captionStart="FIGURE 5" captionStartId="7.[151,250,1776,1800]" captionTargetBox="[158,1430,181,1752]" captionTargetId="figure-18@7.[158,1430,181,1752]" captionTargetPageId="7" captionText="FIGURE 5. Clpeocaenis laosensis sp. nov., A. Gills III, IV. B. Egg, arrow indicate the micropyle. C. Epithema. D. Detail of the micropyle. Scale bars: A = 500 μm; B = 100 μm; C = 40 μm; D = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.13745371" httpUri="https://zenodo.org/record/13745371/files/figure.png" pageId="8" pageNumber="509">Fig. 5BD</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF23B989FFA5FFC7FF62FDF1FA36FD66" box="[152,1415,620,645]" pageId="8" pageNumber="509" startId="8.[152,240,620,644]" targetType="table">
<paragraph id="8BE3E901FFA5FFC7FF62FDF1FA36FD66" blockId="8.[152,1415,620,645]" box="[152,1415,620,645]" pageId="8" pageNumber="509">
<emphasis id="B9283513FFA5FFC7FF62FDF1FEB8FD66" bold="true" box="[152,265,620,644]" pageId="8" pageNumber="509">TABLE 1.</emphasis>
Comparative morphological characteristics of
<taxonomicName id="4C5C9282FFA5FFC7FD1FFDF1FC60FD66" authorityName="Vannachak &amp; Malzacher &amp; Sangpradub" authorityYear="2024" box="[741,977,620,644]" class="Insecta" family="Caenidae" genus="Clypeocaenis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="509" phylum="Arthropoda" rank="species" species="laosensis" status="sp. nov.">
<emphasis id="B9283513FFA5FFC7FD1FFDF1FC60FD66" box="[741,977,620,644]" italics="true" pageId="8" pageNumber="509">Clypeocaenis laosensis</emphasis>
</taxonomicName>
<emphasis id="B9283513FFA5FFC7FC2DFDF0FB99FD67" bold="true" box="[983,1064,621,645]" pageId="8" pageNumber="509">
<taxonomicNameLabel id="A21B8868FFA5FFC7FC2DFDF0FB99FD67" box="[983,1064,621,645]" pageId="8" pageNumber="509" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
larvae and closely related species.
</paragraph>
</caption>
<paragraph id="8BE3E901FFA5FFC7FF65FD01FB5AF86F" pageId="8" pageNumber="509">
<table id="F95C1BA1FFA50030FF65FD01FA3EF832" box="[159,1423,668,2000]" gridcols="4" gridrows="15" pageId="8" pageNumber="509">
<tr id="356CEB43FFA50030FF65FD01FA3EFD52" box="[159,1423,668,688]" gridrow="0" pageId="8" pageNumber="509">
<th id="76BD823FFFA50030FF65FD01FDB7FD52" box="[159,518,668,688]" gridcol="0" gridrow="0" pageId="8" pageNumber="509">
<emphasis id="B9283513FFA5FFC7FF65FD01FEB4FD4D" bold="true" box="[159,261,668,687]" pageId="8" pageNumber="509">Characters</emphasis>
</th>
<th id="76BD823FFFA50030FDE5FD01FC87FD52" box="[543,822,668,688]" gridcol="1" gridrow="0" pageId="8" pageNumber="509">
<taxonomicName id="4C5C9282FFA5FFC7FDE5FD00FD30FD4D" authorityName="Balasubramanian &amp; Muthukatturaja" authorityYear="2020" box="[543,641,668,688]" class="Insecta" family="Caenidae" genus="Clypeocaenis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="509" phylum="Arthropoda" rank="species" species="gayathri">
<emphasis id="B9283513FFA5FFC7FDE5FD00FD30FD4D" bold="true" box="[543,641,668,688]" italics="true" pageId="8" pageNumber="509">
C.
<emphasis id="B9283513FFA5FFC7FDC2FD01FD30FD4D" bold="true" box="[568,641,668,687]" italics="true" pageId="8" pageNumber="509">gayathri</emphasis>
</emphasis>
</taxonomicName>
</th>
<th id="76BD823FFFA50030FCB3FD01FBD8FD52" box="[841,1129,668,688]" gridcol="2" gridrow="0" pageId="8" pageNumber="509">
<taxonomicName id="4C5C9282FFA5FFC7FCB3FD00FC28FD4D" authorityName="Balasubramanian &amp; Muthukatturaja" authorityYear="2021" box="[841,921,668,688]" class="Insecta" family="Caenidae" genus="Clypeocaenis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="509" phylum="Arthropoda" rank="species" species="kaveri">
<emphasis id="B9283513FFA5FFC7FCB3FD00FC28FD4D" bold="true" box="[841,921,668,688]" pageId="8" pageNumber="509">
<emphasis id="B9283513FFA5FFC7FCB3FD00FCE6FD52" bold="true" box="[841,855,669,688]" italics="true" pageId="8" pageNumber="509">C</emphasis>
.
<emphasis id="B9283513FFA5FFC7FC98FD01FC28FD4D" bold="true" box="[866,921,668,687]" italics="true" pageId="8" pageNumber="509">kaveri</emphasis>
</emphasis>
</taxonomicName>
</th>
<th id="76BD823FFFA50030FB83FD01FA3EFD52" box="[1145,1423,668,688]" gridcol="3" gridrow="0" pageId="8" pageNumber="509">
<emphasis id="B9283513FFA5FFC7FB83FD00FA9AFD52" bold="true" box="[1145,1323,668,688]" pageId="8" pageNumber="509">
<taxonomicName id="4C5C9282FFA5FFC7FB83FD00FB50FD4D" authorityName="Vannachak &amp; Malzacher &amp; Sangpradub" authorityYear="2024" box="[1145,1249,668,688]" class="Insecta" family="Caenidae" genus="Clypeocaenis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="509" phylum="Arthropoda" rank="species" species="laosensis" status="sp. nov.">
<emphasis id="B9283513FFA5FFC7FB83FD00FB36FD52" bold="true" box="[1145,1159,669,688]" italics="true" pageId="8" pageNumber="509">C</emphasis>
.
<emphasis id="B9283513FFA5FFC7FB68FD01FB50FD4D" bold="true" box="[1170,1249,668,687]" italics="true" pageId="8" pageNumber="509">laosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B8868FFA5FFC7FB1DFD00FA9AFD52" box="[1255,1323,669,688]" pageId="8" pageNumber="509" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
</tr>
<tr id="356CEB43FFA50030FF65FD4BFA3EFD08" box="[159,1423,726,746]" gridrow="1" pageId="8" pageNumber="509">
<th id="76BD823FFFA50030FF65FD4BFDB7FD08" box="[159,518,726,746]" gridcol="0" gridrow="1" pageId="8" pageNumber="509">
<emphasis id="B9283513FFA5FFC7FF65FD4BFE06FD0B" bold="true" box="[159,439,726,745]" pageId="8" pageNumber="509">Length of pedicel: segment III</emphasis>
</th>
<td id="76BD823FFFA50030FDE5FD4BFC87FD08" box="[543,822,726,746]" gridcol="1" gridrow="1" pageId="8" pageNumber="509">2.5: 1</td>
<td id="76BD823FFFA50030FCB3FD4BFBD8FD08" box="[841,1129,726,746]" gridcol="2" gridrow="1" pageId="8" pageNumber="509">1.5: 1</td>
<td id="76BD823FFFA50030FB83FD4BFA3EFD08" box="[1145,1423,726,746]" gridcol="3" gridrow="1" pageId="8" pageNumber="509">1.5: 1</td>
</tr>
<tr id="356CEB43FFA50030FF65FC88FA3EFCCB" box="[159,1423,789,809]" gridrow="2" pageId="8" pageNumber="509">
<th id="76BD823FFFA50030FF65FC88FDB7FCCB" box="[159,518,789,809]" gridcol="0" gridrow="2" pageId="8" pageNumber="509">
<emphasis id="B9283513FFA5FFC7FF65FC88FEF7FCCA" bold="true" box="[159,326,789,808]" pageId="8" pageNumber="509">Setae below molar</emphasis>
</th>
<td id="76BD823FFFA50030FDE5FC88FC87FCCB" box="[543,822,789,809]" gridcol="1" gridrow="2" pageId="8" pageNumber="509">6 setae</td>
<td id="76BD823FFFA50030FCB3FC88FBD8FCCB" box="[841,1129,789,809]" gridcol="2" gridrow="2" pageId="8" pageNumber="509">7 setae</td>
<td id="76BD823FFFA50030FB83FC88FA3EFCCB" box="[1145,1423,789,809]" gridcol="3" gridrow="2" pageId="8" pageNumber="509">45 setae</td>
</tr>
<tr id="356CEB43FFA50030FF65FCC9FA3EFC49" box="[159,1423,852,939]" gridrow="3" pageId="8" pageNumber="509">
<th id="76BD823FFFA50030FF65FCC9FDB7FC49" box="[159,518,852,939]" gridcol="0" gridrow="3" pageId="8" pageNumber="509">
<emphasis id="B9283513FFA5FFC7FF65FCC9FE1BFC6B" bold="true" pageId="8" pageNumber="509">Forefemur setae: inner margin; outer margin; transverse row</emphasis>
</th>
<td id="76BD823FFFA50030FDE5FCC9FC87FC49" box="[543,822,852,939]" gridcol="1" gridrow="3" pageId="8" pageNumber="509">4 long setae; stout and bifid setae; bifid setae</td>
<td id="76BD823FFFA50030FCB3FCC9FBD8FC49" box="[841,1129,852,939]" gridcol="2" gridrow="3" pageId="8" pageNumber="509">4 long setae; both outer margin and transverse row with bifid setae</td>
<td id="76BD823FFFA50030FB83FCC9FA3EFC49" box="[1145,1423,852,939]" gridcol="3" gridrow="3" pageId="8" pageNumber="509">3 long setae; simple setae; large broadly spatulate setae</td>
</tr>
<tr id="356CEB43FFA50030FF65FC4BFA3EFC08" box="[159,1423,982,1002]" gridrow="4" pageId="8" pageNumber="509">
<th id="76BD823FFFA50030FF65FC4BFDB7FC08" box="[159,518,982,1002]" gridcol="0" gridrow="4" pageId="8" pageNumber="509">
<emphasis id="B9283513FFA5FFC7FF65FC4BFF69FC0B" bold="true" box="[159,216,982,1001]" pageId="8" pageNumber="509">Coxae</emphasis>
</th>
<td id="76BD823FFFA50030FDE5FC4BFC87FC08" box="[543,822,982,1002]" gridcol="1" gridrow="4" pageId="8" pageNumber="509">enlarged</td>
<td id="76BD823FFFA50030FCB3FC4BFBD8FC08" box="[841,1129,982,1002]" gridcol="2" gridrow="4" pageId="8" pageNumber="509">not enlarged</td>
<td id="76BD823FFFA50030FB83FC4BFA3EFC08" box="[1145,1423,982,1002]" gridcol="3" gridrow="4" pageId="8" pageNumber="509">not enlarged</td>
</tr>
<tr id="356CEB43FFA50030FF65FB88FA3EFBA8" box="[159,1423,1045,1098]" gridrow="5" pageId="8" pageNumber="509">
<th id="76BD823FFFA50030FF65FB88FDB7FBA8" box="[159,518,1045,1098]" gridcol="0" gridrow="5" pageId="8" pageNumber="509">
<emphasis id="B9283513FFA5FFC7FF65FB88FEAEFBA8" bold="true" pageId="8" pageNumber="509">Mid femur setae-shaped: outer margin; dorsal surface</emphasis>
</th>
<td id="76BD823FFFA50030FDE5FB88FC87FBA8" box="[543,822,1045,1098]" gridcol="1" gridrow="5" pageId="8" pageNumber="509">stout setae; bifid</td>
<td id="76BD823FFFA50030FCB3FB88FBD8FBA8" box="[841,1129,1045,1098]" gridcol="2" gridrow="5" pageId="8" pageNumber="509">both with bifid</td>
<td id="76BD823FFFA50030FB83FB88FA3EFBA8" box="[1145,1423,1045,1098]" gridcol="3" gridrow="5" pageId="8" pageNumber="509">both with spatulate</td>
</tr>
<tr id="356CEB43FFA50030FF65FBE8FA3EFB6B" box="[159,1423,1141,1161]" gridrow="6" pageId="8" pageNumber="509">
<th id="76BD823FFFA50030FF65FBE8FDB7FB6B" box="[159,518,1141,1161]" gridcol="0" gridrow="6" pageId="8" pageNumber="509">
<emphasis id="B9283513FFA5FFC7FF65FBE8FE6AFB6B" bold="true" box="[159,475,1141,1161]" pageId="8" pageNumber="509">Hind femur marginal setae-shaped</emphasis>
</th>
<td id="76BD823FFFA50030FDE5FBE8FC87FB6B" box="[543,822,1141,1161]" gridcol="1" gridrow="6" pageId="8" pageNumber="509">stout setae</td>
<td id="76BD823FFFA50030FCB3FBE8FBD8FB6B" box="[841,1129,1141,1161]" gridcol="2" gridrow="6" pageId="8" pageNumber="509">stout bifid setae</td>
<td id="76BD823FFFA50030FB83FBE8FA3EFB6B" box="[1145,1423,1141,1161]" gridcol="3" gridrow="6" pageId="8" pageNumber="509">spatulate setae</td>
</tr>
<tr id="356CEB43FFA50030FF65FB28FA3EFB2A" box="[159,1423,1205,1224]" gridrow="7" pageId="8" pageNumber="509">
<th id="76BD823FFFA50030FF65FB28FDB7FB2A" box="[159,518,1205,1224]" gridcol="0" gridrow="7" pageId="8" pageNumber="509">
<emphasis id="B9283513FFA5FFC7FF65FB28FDB7FB2A" bold="true" box="[159,518,1205,1224]" pageId="8" pageNumber="509">Number of denticles of the tarsal claws;</emphasis>
</th>
<td id="76BD823FFFA50030FDE5FB28FC87FB2A" box="[543,822,1205,1224]" gridcol="1" gridrow="7" pageId="8" pageNumber="509">910</td>
<td id="76BD823FFFA50030FCB3FB28FBD8FB2A" box="[841,1129,1205,1224]" gridcol="2" gridrow="7" pageId="8" pageNumber="509">68</td>
<td id="76BD823FFFA50030FB83FB28FA3EFB2A" box="[1145,1423,1205,1224]" gridcol="3" gridrow="7" pageId="8" pageNumber="509">35</td>
</tr>
<tr id="356CEB43FFA50030FF65FB69FA3EFAEA" box="[159,1423,1268,1288]" gridrow="8" pageId="8" pageNumber="509">
<th id="76BD823FFFA50030FF65FB69FDB7FAEA" box="[159,518,1268,1288]" gridcol="0" gridrow="8" pageId="8" pageNumber="509">
<emphasis id="B9283513FFA5FFC7FF65FB69FEE6FAE5" bold="true" box="[159,343,1268,1287]" pageId="8" pageNumber="509">Fore and hind claws</emphasis>
</th>
<td id="76BD823FFFA50030FDE5FB69FC87FAEA" box="[543,822,1268,1288]" gridcol="1" gridrow="8" pageId="8" pageNumber="509">shortened, strong bowed</td>
<td id="76BD823FFFA50030FCB3FB69FBD8FAEA" box="[841,1129,1268,1288]" gridcol="2" gridrow="8" pageId="8" pageNumber="509">not shortened, slightly bowed</td>
<td id="76BD823FFFA50030FB83FB69FA3EFAEA" box="[1145,1423,1268,1288]" gridcol="3" gridrow="8" pageId="8" pageNumber="509">not shortened, slightly bowed</td>
</tr>
<tr id="356CEB43FFA50030FF65FAAEFA3EFA8A" box="[159,1423,1331,1384]" gridrow="9" pageId="8" pageNumber="509">
<th id="76BD823FFFA50030FF65FAAEFDB7FA8A" box="[159,518,1331,1384]" gridcol="0" gridrow="9" pageId="8" pageNumber="509">
<emphasis id="B9283513FFA5FFC7FF65FAAEFF0EFA8A" bold="true" pageId="8" pageNumber="509">Posterior margin of abdominal tergum VII</emphasis>
</th>
<td id="76BD823FFFA50030FDE5FAAEFC87FA8A" box="[543,822,1331,1384]" gridcol="1" gridrow="9" pageId="8" pageNumber="509">spatulate, bifid and pinnate setae</td>
<td id="76BD823FFFA50030FCB3FAAEFBD8FA8A" box="[841,1129,1331,1384]" gridcol="2" gridrow="9" pageId="8" pageNumber="509">bifid setae</td>
<td id="76BD823FFFA50030FB83FAAEFA3EFA8A" box="[1145,1423,1331,1384]" gridcol="3" gridrow="9" pageId="8" pageNumber="509">strongly denticulate, small spatulate setae</td>
</tr>
<tr id="356CEB43FFA50030FF65FA09FA3EF9E9" box="[159,1423,1428,1547]" gridrow="10" pageId="8" pageNumber="509">
<th id="76BD823FFFA50030FF65FA09FDB7F9E9" box="[159,518,1428,1547]" gridcol="0" gridrow="10" pageId="8" pageNumber="509">
<emphasis id="B9283513FFA5FFC7FF65FA09FF63F9E9" bold="true" pageId="8" pageNumber="509">Operculate gill setae on posterior margin; basal medial ridge; microtrichia band on ventral surface of gill</emphasis>
</th>
<td id="76BD823FFFA50030FDE5FA09FC87F9E9" box="[543,822,1428,1547]" gridcol="1" gridrow="10" pageId="8" pageNumber="509">row of short setae; 6 stout setae; not known</td>
<td id="76BD823FFFA50030FCB3FA09FBD8F9E9" box="[841,1129,1428,1547]" gridcol="2" gridrow="10" pageId="8" pageNumber="509">small bifid setae; 9 bifid setae; 23 irregular row of cluster 26 setae</td>
<td id="76BD823FFFA50030FB83FA09FA3EF9E9" box="[1145,1423,1428,1547]" gridcol="3" gridrow="10" pageId="8" pageNumber="509">short broadly spatulate setae; 810 spatulate setae; without microtrichia band</td>
</tr>
<tr id="356CEB43FFA50030FF65F9ABFA3EF98E" box="[159,1423,1590,1644]" gridrow="11" pageId="8" pageNumber="509">
<th id="76BD823FFFA50030FF65F9ABFDB7F98E" box="[159,518,1590,1644]" gridcol="0" gridrow="11" pageId="8" pageNumber="509">
<emphasis id="B9283513FFA5FFC7FF65F9ABFEACF989" bold="true" pageId="8" pageNumber="509">Hind margin of sternum IX and shagreen field</emphasis>
</th>
<td id="76BD823FFFA50030FDE5F9ABFC87F98E" box="[543,822,1590,1644]" gridcol="1" gridrow="11" pageId="8" pageNumber="509">not known</td>
<td id="76BD823FFFA50030FCB3F9ABFBD8F98E" box="[841,1129,1590,1644]" gridcol="2" gridrow="11" pageId="8" pageNumber="509">broadly round with 56 irregular rows of small denticles</td>
<td id="76BD823FFFA50030FB83F9ABFA3EF98E" box="[1145,1423,1590,1644]" gridcol="3" gridrow="11" pageId="8" pageNumber="509">slightly concave with 4 irregular rows of small denticles</td>
</tr>
<tr id="356CEB43FFA50030FF65F90AFA3EF8EC" box="[159,1423,1687,1806]" gridrow="12" pageId="8" pageNumber="509">
<th id="76BD823FFFA50030FF65F90AFDB7F8EC" box="[159,518,1687,1806]" gridcol="0" gridrow="12" pageId="8" pageNumber="509">
<emphasis id="B9283513FFA5FFC7FF65F90AFF7AF948" bold="true" box="[159,203,1687,1706]" pageId="8" pageNumber="509">Eggs</emphasis>
</th>
<td id="76BD823FFFA50030FDE5F90AFC87F8EC" box="[543,822,1687,1806]" gridcol="1" gridrow="12" pageId="8" pageNumber="509">not known</td>
<td id="76BD823FFFA50030FCB3F90AFBD8F8EC" box="[841,1129,1687,1806]" gridcol="2" gridrow="12" pageId="8" pageNumber="509">oval-shape; chorion smooth; epithema flat; micropyle short, mouth slightly broadened; sperm- guide invisible</td>
<td id="76BD823FFFA50030FB83F90AFA3EF8EC" box="[1145,1423,1687,1806]" gridcol="3" gridrow="12" pageId="8" pageNumber="509">oval-shape; chorion smooth; epithema cap-shaped; micropyle long, mouth not broadened; sperm-guide circular</td>
</tr>
<tr id="356CEB43FFA50030FF65F8A7FA3EF8AC" box="[159,1423,1850,1870]" gridrow="13" pageId="8" pageNumber="509">
<th id="76BD823FFFA50030FF65F8A7FDB7F8AC" box="[159,518,1850,1870]" gridcol="0" gridrow="13" pageId="8" pageNumber="509">
<emphasis id="B9283513FFA5FFC7FF65F8A7FEBFF8AF" bold="true" box="[159,270,1850,1869]" pageId="8" pageNumber="509">Distribution</emphasis>
</th>
<td id="76BD823FFFA50030FDE5F8A7FC87F8AC" box="[543,822,1850,1870]" gridcol="1" gridrow="13" pageId="8" pageNumber="509">India</td>
<td id="76BD823FFFA50030FCB3F8A7FBD8F8AC" box="[841,1129,1850,1870]" gridcol="2" gridrow="13" pageId="8" pageNumber="509">India</td>
<td id="76BD823FFFA50030FB83F8A7FA3EF8AC" box="[1145,1423,1850,1870]" gridcol="3" gridrow="13" pageId="8" pageNumber="509">Lao PDR</td>
</tr>
<tr id="356CEB43FFA50030FF65F8E4FA3EF832" box="[159,1423,1913,2000]" gridrow="14" pageId="8" pageNumber="509">
<th id="76BD823FFFA50030FF65F8E4FDB7F832" box="[159,518,1913,2000]" gridcol="0" gridrow="14" pageId="8" pageNumber="509">
<emphasis id="B9283513FFA5FFC7FF65F8E4FEB3F86E" bold="true" box="[159,258,1913,1932]" pageId="8" pageNumber="509">References</emphasis>
</th>
<td id="76BD823FFFA50030FDE5F8E4FC87F832" box="[543,822,1913,2000]" gridcol="1" gridrow="14" pageId="8" pageNumber="509">
<bibRefCitation id="EFCD94F0FFA5FFC7FDE5F8E7FCA0F86F" author="Muthukatturaja, M. &amp; Balasubramanian, C. &amp; Murugan, A." box="[543,785,1914,1933]" pageId="8" pageNumber="509" pagination="361 - 370" refId="ref10400" refString="Muthukatturaja, M., Balasubramanian, C. &amp; Murugan, A. (2020) Two new species of Clypeocaenis (Ephemeroptera: Caenidae) from Western Ghats, Southern India. Zootaxa, 4722 (4), 361 - 370. https: // doi. org / 10.11646 / zootaxa. 4722.4.5" type="journal article" year="2020">
Muthukatturaja
<emphasis id="B9283513FFA5FFC7FD52F8E7FD7CF86F" box="[680,717,1914,1933]" italics="true" pageId="8" pageNumber="509">et al</emphasis>
. (2020)
</bibRefCitation>
</td>
<td id="76BD823FFFA50030FCB3F8E4FBD8F832" box="[841,1129,1913,2000]" gridcol="2" gridrow="14" pageId="8" pageNumber="509">
<bibRefCitation id="EFCD94F0FFA5FFC7FCB3F8E7FBBCF84C" author="Balasubramanian, C. &amp; Muthukatturaja, M." pageId="8" pageNumber="509" pagination="377 - 388" refId="ref9722" refString="Balasubramanian, C. &amp; Muthukatturaja, M. (2021) Two additional new species of Clypeocaenis Soldan, 1978 (Ephemeroptera: Caenidae) from the Western Ghats of Peninsular India. Zootaxa, 4915 (3), 377 - 388. https: // doi. org / 10.11646 / zootaxa. 4915.3.6" type="journal article" year="2021">Balasubramanian &amp; Muthukatturaja (2021)</bibRefCitation>
;
<bibRefCitation id="EFCD94F0FFA5FFC7FCB3F821FBA0F832" author="Srinivasan, P. &amp; Sivaruban, T. &amp; Barathy, S. &amp; Isack, R." box="[841,1041,1980,2000]" pageId="8" pageNumber="509" pagination="39 - 75" refId="ref10643" refString="Srinivasan, P., Sivaruban, T., Barathy, S. &amp; Isack, R. (2023) Contribution to the knowledge of family Caenidae (Insecta: Ephemeroptera) from Madurai District, Tamil Nadu. Zootaxa, 5258 (1), 39 - 75. https: // doi. org / 10.11646 / zootaxa. 5258.1.2" type="journal article" year="2023">
Srinivasan
<emphasis id="B9283513FFA5FFC7FC52F820FC7FF82D" box="[936,974,1980,1999]" italics="true" pageId="8" pageNumber="509">et al</emphasis>
. (2023)
</bibRefCitation>
</td>
<td id="76BD823FFFA50030FB83F8E4FA3EF832" box="[1145,1423,1913,2000]" gridcol="3" gridrow="14" pageId="8" pageNumber="509">Present study</td>
</tr>
</table>
</paragraph>
<subSubSection id="C346BA8AFFA4FFC6FF3DFF0BFC55FF37" pageId="9" pageNumber="510" type="etymology">
<paragraph id="8BE3E901FFA4FFC6FF3DFF0BFC55FF37" blockId="9.[151,1437,150,753]" pageId="9" pageNumber="510">
<emphasis id="B9283513FFA4FFC6FF3DFF0BFEF9FF53" bold="true" box="[199,328,150,177]" italics="true" pageId="9" pageNumber="510">Etymology.</emphasis>
The specific epithet
<taxonomicName id="4C5C9282FFA4FFC6FDCDFF0AFD2FFF53" authorityName="Vannachak &amp; Malzacher &amp; Sangpradub" authorityYear="2024" box="[567,670,151,177]" class="Insecta" family="Caenidae" genus="Clypeocaenis" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="510" phylum="Arthropoda" rank="species" species="laosensis">
<emphasis id="B9283513FFA4FFC6FDCDFF0AFD2FFF53" box="[567,670,151,177]" italics="true" pageId="9" pageNumber="510">laosensis</emphasis>
</taxonomicName>
is derived from the country of collection,
<collectingCountry id="F34BA991FFA4FFC6FB85FF0AFF4DFF37" name="Laos" pageId="9" pageNumber="510">Lao Peoples Democratic Republic</collectingCountry>
(Lao PDR), indicating the geographical origin of the new species.
</paragraph>
</subSubSection>
<subSubSection id="C346BA8AFFA4FFC6FF3DFF43FD53FD13" pageId="9" pageNumber="510" type="distribution">
<paragraph id="8BE3E901FFA4FFC6FF3DFF43FD53FD13" blockId="9.[151,1437,150,753]" pageId="9" pageNumber="510">
<emphasis id="B9283513FFA4FFC6FF3DFF43FE49FF1B" bold="true" box="[199,504,222,249]" pageId="9" pageNumber="510">
<emphasis id="B9283513FFA4FFC6FF3DFF43FE40FF1B" bold="true" box="[199,497,222,249]" italics="true" pageId="9" pageNumber="510">Distribution and Habitat</emphasis>
.
</emphasis>
<taxonomicName id="4C5C9282FFA4FFC6FDFCFF42FCA3FF1B" authorityName="Vannachak &amp; Malzacher &amp; Sangpradub" authorityYear="2024" box="[518,786,223,249]" class="Insecta" family="Caenidae" genus="Clypeocaenis" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="510" phylum="Arthropoda" rank="species" species="laosensis" status="sp. nov.">
<emphasis id="B9283513FFA4FFC6FDFCFF42FCA3FF1B" box="[518,786,223,249]" italics="true" pageId="9" pageNumber="510">Clypeocaenis laosensis</emphasis>
</taxonomicName>
<emphasis id="B9283513FFA4FFC6FCE5FF42FCCEFF1B" bold="true" box="[799,895,223,249]" pageId="9" pageNumber="510">
<taxonomicNameLabel id="A21B8868FFA4FFC6FCE5FF42FCCEFF1B" box="[799,895,223,249]" pageId="9" pageNumber="510" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has been collected in the
<collectingRegion id="499827E3FFA4FFC6FB33FF42FAD7FF1B" box="[1225,1382,223,249]" country="Laos" name="Khammouan" pageId="9" pageNumber="510">Khammouane</collectingRegion>
and
<collectingRegion id="499827E3FFA4FFC6FF6DFE9EFE97FEFF" box="[151,294,259,285]" country="Laos" name="Savannakhet" pageId="9" pageNumber="510">Savannakhet</collectingRegion>
Provinces of the Lao PDR (
<figureCitation id="1367F584FFA4FFC6FD8EFE9EFD7BFEFF" box="[628,714,259,285]" captionStart="FIGURE 14" captionStartId="20.[151,250,1091,1115]" captionTargetBox="[160,1430,185,1064]" captionTargetId="figure-26@20.[158,1430,181,1067]" captionTargetPageId="20" captionText="FIGURE 14. Geographic distribution of the Clypeocaenis laosensis sp. nov. (circle), C. oligosetosa (star); C. multisetosa (triangle)." figureDoi="http://doi.org/10.5281/zenodo.13745401" httpUri="https://zenodo.org/record/13745401/files/figure.png" pageId="9" pageNumber="510">Fig. 14</figureCitation>
). Specimens were collected from stream and rivers situated in the central and southern regions of Lao PDR, within an elevation range of
<quantity id="4CA444E4FFA4FFC6FC01FEBBFBCCFEA3" box="[1019,1149,294,321]" metricMagnitude="2" metricUnit="m" metricValue="1.4" metricValueMax="1.54" metricValueMin="1.26" pageId="9" pageNumber="510" unit="m" value="140.0" valueMax="154.0" valueMin="126.0">126154 m</quantity>
above sea level. The Xe Nou River (
<figureCitation id="1367F584FFA4FFC6FEE4FED6FE34FE87" box="[286,389,331,357]" captionStart="FIGURE 13" captionStartId="19.[151,251,1776,1800]" captionTargetBox="[158,1430,179,1750]" captionTargetId="figure-18@19.[158,1430,179,1750]" captionTargetPageId="19" captionText="FIGURE 13. Habitats of the genus Clypeocaenis Soldán, 1978 in Lao PDR, A. Xe Nou River, Xebangfai District, Khammouane Province. B. Tha Hao Stream, Xaybury District, Savannakhet Province. C. Xe Bang Hieng River, Songkhone District, Savannakhet Province. D. Outlet Stream of Nam Xouang Reservoir, Naxaythong District, Vientiane Capital." figureDoi="http://doi.org/10.5281/zenodo.13745397" httpUri="https://zenodo.org/record/13745397/files/figure.png" pageId="9" pageNumber="510">Fig. 13A</figureCitation>
) meanders through a village setting, bordered on one side by agricultural lands and forested areas. The riverbed primarily consists of bedrock and boulders, complemented by patches of gravel and sand. The water current exhibits a moderate to fast flow, particularly intensifying in the rivers central channel. The Tha Hao Stream (
<figureCitation id="1367F584FFA4FFC6FF0FFE2AFEE9FE33" box="[245,344,439,465]" captionStart="FIGURE 13" captionStartId="19.[151,251,1776,1800]" captionTargetBox="[158,1430,179,1750]" captionTargetId="figure-18@19.[158,1430,179,1750]" captionTargetPageId="19" captionText="FIGURE 13. Habitats of the genus Clypeocaenis Soldán, 1978 in Lao PDR, A. Xe Nou River, Xebangfai District, Khammouane Province. B. Tha Hao Stream, Xaybury District, Savannakhet Province. C. Xe Bang Hieng River, Songkhone District, Savannakhet Province. D. Outlet Stream of Nam Xouang Reservoir, Naxaythong District, Vientiane Capital." figureDoi="http://doi.org/10.5281/zenodo.13745397" httpUri="https://zenodo.org/record/13745397/files/figure.png" pageId="9" pageNumber="510">Fig. 13B</figureCitation>
) flows adjacent to a small village, encircled by agricultural fields and a stretch of forest. Its current is characterized as moderately fast. The streambeds composition largely includes bedrock, boulders, and sand, interspersed with snags and leaf litter, providing a diverse substrate. Bordering the Xe Bang Hieng River (
<figureCitation id="1367F584FFA4FFC6FAD0FE62FA3EFDFB" box="[1322,1423,511,537]" captionStart="FIGURE 13" captionStartId="19.[151,251,1776,1800]" captionTargetBox="[158,1430,179,1750]" captionTargetId="figure-18@19.[158,1430,179,1750]" captionTargetPageId="19" captionText="FIGURE 13. Habitats of the genus Clypeocaenis Soldán, 1978 in Lao PDR, A. Xe Nou River, Xebangfai District, Khammouane Province. B. Tha Hao Stream, Xaybury District, Savannakhet Province. C. Xe Bang Hieng River, Songkhone District, Savannakhet Province. D. Outlet Stream of Nam Xouang Reservoir, Naxaythong District, Vientiane Capital." figureDoi="http://doi.org/10.5281/zenodo.13745397" httpUri="https://zenodo.org/record/13745397/files/figure.png" pageId="9" pageNumber="510">Fig. 13C</figureCitation>
), residential dwellings and vegetable gardens are prevalent. The riverbed here is distinguished by a foundation of bedrock, boulders, and remnants of concrete from old bridges, with minor sections of gravel, sand, mud, large wood debris (snags), and plant litter. This river features a moderate to fast current, especially intensifying in the rivers central channel. Larvae of this newly described species were predominantly found in the littoral zones, which are defined by their boulder, gravel, and sand substrates, accompanied by slower current velocities, providing a specific ecological niche for
<taxonomicName id="4C5C9282FFA4FFC6FE87FD4AFD30FD13" authorityName="Vannachak &amp; Malzacher &amp; Sangpradub" authorityYear="2024" box="[381,641,727,753]" class="Insecta" family="Caenidae" genus="Clypeocaenis" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="510" phylum="Arthropoda" rank="species" species="laosensis" status="sp. nov.">
<emphasis id="B9283513FFA4FFC6FE87FD4AFD30FD13" box="[381,641,727,753]" italics="true" pageId="9" pageNumber="510">Clypeocaenis laosensis</emphasis>
</taxonomicName>
<emphasis id="B9283513FFA4FFC6FD72FD4AFD53FD13" bold="true" box="[648,738,727,753]" pageId="9" pageNumber="510">
<taxonomicNameLabel id="A21B8868FFA4FFC6FD72FD4AFD53FD13" box="[648,738,727,753]" pageId="9" pageNumber="510" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="F495695B209DEA4D72E54E6D91E99E82" ID-DOI="10.11646/zootaxa.5501.4.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="13745467" ID-ZooBank="E8E3071A-9082-42FA-BFE7-DED7B2359B05" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726051241689" checkinUser="plazi" docAuthor="Breedy, Odalisca, Murillo-Cruz, Catalina &amp; Cortés, Jorge" docDate="2024" docId="602701104846B408FF44F9E2E9E28ADF" docLanguage="en" docName="zootaxa.5501.4.7.pdf" docOrigin="Zootaxa 5501 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5501.4.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Heterogorgia abdita Breedy, Murillo-Cruz &amp; Cortés, 2024, sp. nov." docType="treatment" docUuid="E4BEA3A4-3221-4C9D-A6D8-3DCFF5D820EE" docUuidSource="ZooBank" docVersion="2" lastPageNumber="579" masterDocId="9C1E79684844B40EFFD3FFFEEB798A26" masterDocTitle="A new species of Heterogorgia Verrill, 1868 (Octocorallia, Malacalcyonacea) from a mesophotic fishing shoal in the eastern Pacific" masterLastPageNumber="582" masterPageNumber="573" pageNumber="575" updateTime="1726051649801" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="27B6A9A73284F6682F8E094859983AB1">A new species of Heterogorgia Verrill, 1868 (Octocorallia, Malacalcyonacea) from a mesophotic fishing shoal in the eastern Pacific</mods:title>
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<mods:namePart id="353A8DF6676FC7C761792ED605AA033C">Breedy, Odalisca</mods:namePart>
<mods:affiliation id="2F6E015EAB2A54F89D68161A0397AA3E">Museo de Zoología, Centro de Investigación en Biodiversidad y Ecología Tropical, Universidad de Costa Rica, P. O. Box 11501 - 2060, San José, Costa Rica, and the Smithsonian Tropical Research Institute, Republic of Panama &amp; Centro de Investigación en Estructuras Microscópicas, Universidad de Costa Rica, San José, Costa Rica &amp; Centro de Investigación en Ciencias del Mar y Limnología, Universidad de Costa Rica, San José, Costa Rica</mods:affiliation>
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<mods:namePart id="CD2AE4C36574877170DF0356F417CEE8">Murillo-Cruz, Catalina</mods:namePart>
<mods:affiliation id="CBAA3BB703CC9FD5EF1C3B1750700251">Centro de Investigación en Estructuras Microscópicas, Universidad de Costa Rica, San José, Costa Rica</mods:affiliation>
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<mods:namePart id="0D1F9919868CE43E4C4B48ADCAEAD615">Cortés, Jorge</mods:namePart>
<mods:affiliation id="ADF0FAC414D578D487673492EA590693">Centro de Investigación en Ciencias del Mar y Limnología, Universidad de Costa Rica, San José, Costa Rica</mods:affiliation>
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<mods:date id="02F4D18EB65413E6401C70CDA3236AA5">2024</mods:date>
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<heading id="B379076A4846B40CFF44F9E2EA898C10" bold="true" box="[151,496,1563,1590]" fontSize="11" level="1" pageId="2" pageNumber="575" reason="1">
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<taxonomicName id="2F8ECB854846B40CFF44F9E2EAF28C10" authorityName="Breedy &amp; Murillo-Cruz &amp; Cortés" authorityYear="2024" box="[151,395,1563,1590]" class="Anthozoa" family="Plexauridae" genus="Heterogorgia" kingdom="Animalia" order="Alcyonacea" pageId="2" pageNumber="575" phylum="Cnidaria" rank="species" species="abdita" status="sp. nov.">
<emphasis id="DAFA6C144846B40CFF44F9E2EAF28C10" bold="true" box="[151,395,1563,1590]" italics="true" pageId="2" pageNumber="575">Heterogorgia abdita</emphasis>
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urn:lsid:zoobank.org:act:
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<figureCitation id="70B5AC834846B40CFF44F99AEB858C58" box="[151,252,1636,1662]" captionStart="FIGURE 1" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[219,1367,181,1919]" captionTargetId="figure-17@3.[219,1367,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. Heterogorgia abdita sp. nov., holotype. MZUCR 4004. (A) Holotype, (B) SEM image of mineralized fibers of the axis; (C) tentacular rods." figureDoi="http://doi.org/10.5281/zenodo.13745469" httpUri="https://zenodo.org/record/13745469/files/figure.png" pageId="2" pageNumber="575">Figures 1</figureCitation>
4
</paragraph>
<paragraph id="E831B0064846B40CFF44F952EA0B8CE0" blockId="2.[151,1437,1708,2023]" box="[151,370,1708,1734]" pageId="2" pageNumber="575">Material examined.</paragraph>
<paragraph id="E831B0064846B40CFF14F92EE8908D28" blockId="2.[151,1437,1708,2023]" pageId="2" pageNumber="575">
<materialsCitation id="58E6BA5B4846B40CFF14F92EE8908D28" collectingDate="2016-12-06" collectionCode="MZUCR" collectorName="La Cruz &amp; G. Ampie &amp; M. Lara &amp; O. Breedy" country="Saint Helena, Ascension and Tristan da Cunha" county="Guanacaste" elevation="45" latitude="10.951" location="Bajo Tigre" longLatPrecision="9" longitude="-85.9056" municipality="La Cruz" pageId="2" pageNumber="575" specimenCode="MZUCR 4004" specimenCount="1" typeStatus="holotype">
<typeStatus id="37350EA44846B40CFF14F92EEA4A8CCC" box="[199,307,1744,1770]" pageId="2" pageNumber="575" type="holotype">Holotype</typeStatus>
.
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, ethanol preserved,
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,
<collectingCountry id="9099F0964846B40CFCBFF92EE8838CCC" box="[876,1018,1744,1770]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="2" pageNumber="575">Santa Elena</collectingCountry>
,
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<collectingMunicipality id="08552A7C4846B40CFBD9F92FEF158CCC" box="[1034,1132,1744,1771]" pageId="2" pageNumber="575">La Cruz</collectingMunicipality>
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,
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,
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,
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,
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,
<collectorName id="457BD5D04846B40CFE5CF90AEA868D28" box="[399,511,1780,1806]" pageId="2" pageNumber="575">G. Ampié</collectorName>
,
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and
<collectorName id="457BD5D04846B40CFD48F90AE8768D28" box="[667,783,1780,1806]" pageId="2" pageNumber="575">O. Breedy</collectorName>
,
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<collectingDate id="8C746F2E4846B40CFCC9F90AE89C8D28" box="[794,997,1780,1806]" pageId="2" pageNumber="575" value="2016-12-06">6 December 2016</collectingDate>
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.
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</paragraph>
<paragraph id="E831B0064846B40CFF14F8E6E9788D70" blockId="2.[151,1437,1708,2023]" pageId="2" pageNumber="575">Sequences have been deposited in GenBank under accession numbers PP864465 (28S rDNA), PP873181 (mtMutS) and PP944718 (COI).</paragraph>
<paragraph id="E831B0064846B40CFF14F89FE9B68D5C" blockId="2.[151,1437,1708,2023]" box="[199,719,1888,1915]" pageId="2" pageNumber="575">
<materialsCitation id="58E6BA5B4846B40CFF14F89FE9B28D5C" box="[199,715,1888,1915]" collectingDate="2016-12-06" collectionCode="MZUCR" collectorName="La Cruz &amp; G. Ampie &amp; M. Lara &amp; O. Breedy" country="Saint Helena, Ascension and Tristan da Cunha" county="Guanacaste" elevation="45" latitude="10.951" location="Bajo Tigre" longLatPrecision="9" longitude="-85.9056" municipality="La Cruz" pageId="2" pageNumber="575" specimenCode="MZUCR 4005" specimenCount="1" typeStatus="paratype">
<typeStatus id="37350EA44846B40CFF14F89FEA548D5D" box="[199,301,1889,1915]" pageId="2" pageNumber="575" type="paratype">Paratype</typeStatus>
.
<specimenCode id="B828187D4846B40CFEE4F89EEAA48D5D" box="[311,477,1888,1915]" collectionCode="MZUCR" pageId="2" pageNumber="575">MZUCR 4005</specimenCode>
, data as the holotype
</materialsCitation>
.
</paragraph>
<paragraph id="E831B0064846B40CFF14F87AE9788DE4" blockId="2.[151,1437,1708,2023]" pageId="2" pageNumber="575">Sequences have been deposited in GenBank under accession numbers PP864466 (28S rDNA), PP873182 (mtMutS) and PP944719 (COI).</paragraph>
<paragraph id="E831B0064846B40CFF14F833EF918DC1" blockId="2.[151,1437,1708,2023]" box="[199,1256,1996,2023]" pageId="2" pageNumber="575">
<materialsCitation id="58E6BA5B4846B40CFF14F833EF918DC1" box="[199,1256,1996,2023]" collectorName="La Cruz" country="Costa Rica" elevation="45" location="Guanacaste" pageId="2" pageNumber="575" specimenCount="45" stateProvince="Guanacaste">
Type locality. Bajo Tigre, Santa Elena,
<collectorName id="457BD5D04846B40CFDADF833E9A28DC0" box="[638,731,1996,2023]" pageId="2" pageNumber="575">La Cruz</collectorName>
,
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, Pacific Ocean,
<collectingCountry id="9099F0964846B40CFBF3F832EFE48DC0" box="[1056,1181,1996,2022]" name="Costa Rica" pageId="2" pageNumber="575">Costa Rica</collectingCountry>
,
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<quantity id="2F761DE34846B40CFB7BF832EF9D8DC1" box="[1192,1252,1996,2023]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="2" pageNumber="575" unit="m" value="45.0">
<elevation id="63A357354846B40CFB7BF832EF9D8DC1" box="[1192,1252,1996,2023]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="2" pageNumber="575" unit="m" value="45.0">45 m</elevation>
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.
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</paragraph>
<caption id="BCF1E08E4847B40DFF4BF869EAEA8DF5" ID-DOI="http://doi.org/10.5281/zenodo.13745469" ID-Zenodo-Dep="13745469" httpUri="https://zenodo.org/record/13745469/files/figure.png" pageId="3" pageNumber="576" startId="3.[152,255,1943,1967]" targetBox="[219,1367,181,1919]" targetPageId="3" targetType="figure">
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<emphasis id="DAFA6C144847B40DFF4BF869EA618D89" bold="true" box="[152,280,1943,1967]" pageId="3" pageNumber="576">FIGURE 1.</emphasis>
<taxonomicName id="2F8ECB854847B40DFECDF866EA958D89" authorityName="Breedy &amp; Murillo-Cruz &amp; Cortés" authorityYear="2024" box="[286,492,1943,1967]" class="Anthozoa" family="Plexauridae" genus="Heterogorgia" kingdom="Animalia" order="Alcyonacea" pageId="3" pageNumber="576" phylum="Cnidaria" rank="species" species="abdita" status="sp. nov.">
<emphasis id="DAFA6C144847B40DFECDF866EA958D89" box="[286,492,1943,1967]" italics="true" pageId="3" pageNumber="576">Heterogorgia abdita</emphasis>
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<taxonomicNameLabel id="C1C9D16F4847B40DFE21F866E93B8D96" box="[498,578,1944,1968]" pageId="3" pageNumber="576" rank="species">sp. nov.</taxonomicNameLabel>
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, holotype. MZUCR 4004. (A) Holotype, (B) SEM image of mineralized fibers of the axis; (C) tentacular rods.
</paragraph>
</caption>
<caption id="BCF1E08E4840B40AFF4BF9FAE81D8C66" ID-DOI="http://doi.org/10.5281/zenodo.13745475" ID-Zenodo-Dep="13745475" httpUri="https://zenodo.org/record/13745475/files/figure.png" pageId="4" pageNumber="577" startId="4.[152,255,1540,1564]" targetBox="[151,1436,181,1516]" targetPageId="4" targetType="figure">
<paragraph id="E831B0064840B40AFF4BF9FAE81D8C66" blockId="4.[151,1436,1540,1600]" pageId="4" pageNumber="577">
<emphasis id="DAFA6C144840B40AFF4BF9FAEA628C3A" bold="true" box="[152,283,1540,1564]" pageId="4" pageNumber="577">FIGURE 2.</emphasis>
<taxonomicName id="2F8ECB854840B40AFEF0F9FBEA8C8C3A" authorityName="Breedy &amp; Murillo-Cruz &amp; Cortés" authorityYear="2024" box="[291,501,1540,1564]" class="Anthozoa" family="Plexauridae" genus="Heterogorgia" kingdom="Animalia" order="Alcyonacea" pageId="4" pageNumber="577" phylum="Cnidaria" rank="species" species="abdita" status="sp. nov.">
<emphasis id="DAFA6C144840B40AFEF0F9FBEA8C8C3A" box="[291,501,1540,1564]" italics="true" pageId="4" pageNumber="577">Heterogorgia abdita</emphasis>
</taxonomicName>
<emphasis id="DAFA6C144840B40AFE2EF9FAE9298C3A" bold="true" box="[509,592,1540,1564]" pageId="4" pageNumber="577">
<taxonomicNameLabel id="C1C9D16F4840B40AFE2EF9FAE9298C3A" box="[509,592,1540,1564]" pageId="4" pageNumber="577" rank="species">sp. nov.</taxonomicNameLabel>
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, holotype, MZUCR 4004, sclerites: (A) collaret spindles; (B) point spindles; (C) thorns; (D-F) spindles; (G) club-like spindles; (H) crosses; (I) radiates.
</paragraph>
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<paragraph id="E831B0064840B40AFF44F96CEA518C8A" blockId="4.[151,296,1682,1708]" box="[151,296,1682,1708]" pageId="4" pageNumber="577">
<emphasis id="DAFA6C144840B40AFF44F96CEA518C8A" bold="true" box="[151,296,1682,1708]" pageId="4" pageNumber="577">Description</emphasis>
</paragraph>
<paragraph id="E831B0064840B40BFF44F924EE2E8847" blockId="4.[151,1437,1754,2033]" lastBlockId="5.[151,1437,151,969]" lastPageId="5" lastPageNumber="578" pageId="4" pageNumber="577">
The
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is a bushy colony irregularly branched, about
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tall and
<quantity id="2F761DE34840B40AFC1DF925EF7D8CD3" box="[974,1028,1755,1781]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="577" unit="cm" value="7.0">7 cm</quantity>
wide. The stem,
<quantity id="2F761DE34840B40AFB6CF924EE608CD3" box="[1215,1305,1754,1781]" metricMagnitude="-1" metricUnit="m" metricValue="1.15" pageId="4" pageNumber="577" unit="mm" value="115.0">115 mm</quantity>
long, arises from an encrusting holdfast,
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in diameter. The stem,
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diameter, subdivides three times close to the base, with subsequent branching up to 7 times. Branches are
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in diameter and split irregularly several times in different planes (
<figureCitation id="70B5AC834840B40AFEA0F8B8EAB58D46" box="[371,460,1862,1888]" captionStart="FIGURE 1" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[219,1367,181,1919]" captionTargetId="figure-17@3.[219,1367,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. Heterogorgia abdita sp. nov., holotype. MZUCR 4004. (A) Holotype, (B) SEM image of mineralized fibers of the axis; (C) tentacular rods." figureDoi="http://doi.org/10.5281/zenodo.13745469" httpUri="https://zenodo.org/record/13745469/files/figure.png" pageId="4" pageNumber="577">Fig. 1A</figureCitation>
). The thinner branchlets are
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diameter. The axis is horny with a chambered central core with filaments coated with hydroxylapatite (
<figureCitation id="70B5AC834840B40AFCC4F894E8178DA2" box="[791,878,1898,1924]" captionStart="FIGURE 1" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[219,1367,181,1919]" captionTargetId="figure-17@3.[219,1367,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. Heterogorgia abdita sp. nov., holotype. MZUCR 4004. (A) Holotype, (B) SEM image of mineralized fibers of the axis; (C) tentacular rods." figureDoi="http://doi.org/10.5281/zenodo.13745469" httpUri="https://zenodo.org/record/13745469/files/figure.png" pageId="4" pageNumber="577">Fig. 1B</figureCitation>
). The polyps are totally or partially retracted into prominent, dome-shaped calyces,
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height and about
<quantity id="2F761DE34840B40AFC99F870E8E68D8F" box="[842,927,1934,1961]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="4" pageNumber="577" unit="mm" value="1.8">1.8 mm</quantity>
in diameter (
<figureCitation id="70B5AC834840B40AFBE1F870EFF38D8E" box="[1074,1162,1934,1960]" captionStart="FIGURE 1" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[219,1367,181,1919]" captionTargetId="figure-17@3.[219,1367,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. Heterogorgia abdita sp. nov., holotype. MZUCR 4004. (A) Holotype, (B) SEM image of mineralized fibers of the axis; (C) tentacular rods." figureDoi="http://doi.org/10.5281/zenodo.13745469" httpUri="https://zenodo.org/record/13745469/files/figure.png" pageId="4" pageNumber="577">Fig. 1A</figureCitation>
). Calyces are irregularly distributed, not very close together,
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apart (about 6 calyces/cm, n=30) and are more separated at the base of the colony and closer at the tips where they are in clusters of 24, bud-like (
<figureCitation id="70B5AC834840B40AFC22F828EF308DD6" box="[1009,1097,2006,2032]" captionStart="FIGURE 1" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[219,1367,181,1919]" captionTargetId="figure-17@3.[219,1367,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. Heterogorgia abdita sp. nov., holotype. MZUCR 4004. (A) Holotype, (B) SEM image of mineralized fibers of the axis; (C) tentacular rods." figureDoi="http://doi.org/10.5281/zenodo.13745469" httpUri="https://zenodo.org/record/13745469/files/figure.png" pageId="4" pageNumber="577">Fig. 1A</figureCitation>
). They are sparsely distributed on the holdfast (
<figureCitation id="70B5AC834841B40BFE9CFF69EADE8A97" box="[335,423,151,177]" captionStart="FIGURE 1" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[219,1367,181,1919]" captionTargetId="figure-17@3.[219,1367,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. Heterogorgia abdita sp. nov., holotype. MZUCR 4004. (A) Holotype, (B) SEM image of mineralized fibers of the axis; (C) tentacular rods." figureDoi="http://doi.org/10.5281/zenodo.13745469" httpUri="https://zenodo.org/record/13745469/files/figure.png" pageId="5" pageNumber="578">Fig. 1B</figureCitation>
). Polyps are preserved totally or partially retracted; they are whitish. Polyp tentacles bear tuberculate rods,
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long and
<quantity id="2F761DE34841B40BFDBBFF45E8618AF3" box="[616,792,187,213]" metricMagnitude="-5" metricUnit="m" metricValue="2.3" metricValueMax="3.0" metricValueMin="1.6" pageId="5" pageNumber="578" unit="mm" value="0.023" valueMax="0.03" valueMin="0.016">0.0160.03 mm</quantity>
wide (
<figureCitation id="70B5AC834841B40BFCB7FF45E8C28AF3" box="[868,955,187,213]" captionStart="FIGURE 1" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[219,1367,181,1919]" captionTargetId="figure-17@3.[219,1367,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. Heterogorgia abdita sp. nov., holotype. MZUCR 4004. (A) Holotype, (B) SEM image of mineralized fibers of the axis; (C) tentacular rods." figureDoi="http://doi.org/10.5281/zenodo.13745469" httpUri="https://zenodo.org/record/13745469/files/figure.png" pageId="5" pageNumber="578">Fig. 1C</figureCitation>
). The anthocodial armature is composed of points of two to three pairs of spindles arranged
<emphasis id="DAFA6C144841B40BFD7DFF21E85E8ADF" box="[686,807,223,249]" italics="true" pageId="5" pageNumber="578">en chevron</emphasis>
, measuring
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long and
<quantity id="2F761DE34841B40BFB6DFF20EE198ADF" box="[1214,1376,222,249]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="5" pageNumber="578" unit="mm" value="0.04" valueMax="0.05" valueMin="0.03">0.030.05 mm</quantity>
wide (
<figureCitation id="70B5AC834841B40BFF4CFEFDEB808B3B" box="[159,249,259,285]" captionStart="FIGURE 2" captionStartId="4.[152,255,1540,1564]" captionTargetBox="[151,1436,181,1516]" captionTargetId="figure-24@4.[151,1436,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Heterogorgia abdita sp. nov., holotype, MZUCR 4004, sclerites: (A) collaret spindles; (B) point spindles; (C) thorns; (D-F) spindles; (G) club-like spindles; (H) crosses; (I) radiates." figureDoi="http://doi.org/10.5281/zenodo.13745475" httpUri="https://zenodo.org/record/13745475/files/figure.png" pageId="5" pageNumber="578">Fig. 2A</figureCitation>
). The point sclerites are bent or straight, some with a spiny distal end, and with sparse spines on the surface. The collaret is composed of two rows of spiny spindles,
<quantity id="2F761DE34841B40BFCC6FED9E8C08B67" box="[789,953,295,321]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="5" pageNumber="578" unit="mm" value="0.25" valueMax="0.3" valueMin="0.2">0.200.30 mm</quantity>
long and
<quantity id="2F761DE34841B40BFBFAFED8EF908B67" box="[1065,1257,294,321]" metricMagnitude="-5" metricUnit="m" metricValue="3" metricValueMax="3.5" metricValueMin="2.5" pageId="5" pageNumber="578" unit="mm" value="0.03" valueMax="0.035" valueMin="0.025">
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mm
</quantity>
wide (
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). The calicular rim bears one whorl of thorns,
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long and
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mm
</quantity>
wide, the thorns shafts bearing short spines (
<figureCitation id="70B5AC834841B40BFE59FE91EA9A8BAF" box="[394,483,367,393]" captionStart="FIGURE 2" captionStartId="4.[152,255,1540,1564]" captionTargetBox="[151,1436,181,1516]" captionTargetId="figure-24@4.[151,1436,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Heterogorgia abdita sp. nov., holotype, MZUCR 4004, sclerites: (A) collaret spindles; (B) point spindles; (C) thorns; (D-F) spindles; (G) club-like spindles; (H) crosses; (I) radiates." figureDoi="http://doi.org/10.5281/zenodo.13745475" httpUri="https://zenodo.org/record/13745475/files/figure.png" pageId="5" pageNumber="578">Fig. 2C</figureCitation>
), and club-like sclerites,
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long and
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mm
</quantity>
wide, with warty tubercles and bifurcated or oval heads (
<figureCitation id="70B5AC834841B40BFD8AFE6DE9CD8B8B" box="[601,692,403,429]" captionStart="FIGURE 2" captionStartId="4.[152,255,1540,1564]" captionTargetBox="[151,1436,181,1516]" captionTargetId="figure-24@4.[151,1436,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Heterogorgia abdita sp. nov., holotype, MZUCR 4004, sclerites: (A) collaret spindles; (B) point spindles; (C) thorns; (D-F) spindles; (G) club-like spindles; (H) crosses; (I) radiates." figureDoi="http://doi.org/10.5281/zenodo.13745475" httpUri="https://zenodo.org/record/13745475/files/figure.png" pageId="5" pageNumber="578">Fig. 2G</figureCitation>
). The coenenchymal sclerites comprise various types of spindles: short and wide, with expanded warty tubercles,
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long and
<quantity id="2F761DE34841B40BFC12FE48EFF98BF7" box="[961,1152,438,465]" metricMagnitude="-5" metricUnit="m" metricValue="7.25" metricValueMax="8.5" metricValueMin="6.0" pageId="5" pageNumber="578" unit="mm" value="0.0725" valueMax="0.085" valueMin="0.06">
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mm
</quantity>
wide (
<figureCitation id="70B5AC834841B40BFB1EFE49EE5F8BF7" box="[1229,1318,439,465]" captionStart="FIGURE 2" captionStartId="4.[152,255,1540,1564]" captionTargetBox="[151,1436,181,1516]" captionTargetId="figure-24@4.[151,1436,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Heterogorgia abdita sp. nov., holotype, MZUCR 4004, sclerites: (A) collaret spindles; (B) point spindles; (C) thorns; (D-F) spindles; (G) club-like spindles; (H) crosses; (I) radiates." figureDoi="http://doi.org/10.5281/zenodo.13745475" httpUri="https://zenodo.org/record/13745475/files/figure.png" pageId="5" pageNumber="578">Fig. 2D</figureCitation>
), and with mostly blunt ends. Spindles are long, straight or bent,
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long and
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mm
</quantity>
wide, with pointed or irregular ends, one end acute and the other blunt (
<figureCitation id="70B5AC834841B40BFD0BFE01E856883F" box="[728,815,511,537]" captionStart="FIGURE 2" captionStartId="4.[152,255,1540,1564]" captionTargetBox="[151,1436,181,1516]" captionTargetId="figure-24@4.[151,1436,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Heterogorgia abdita sp. nov., holotype, MZUCR 4004, sclerites: (A) collaret spindles; (B) point spindles; (C) thorns; (D-F) spindles; (G) club-like spindles; (H) crosses; (I) radiates." figureDoi="http://doi.org/10.5281/zenodo.13745475" httpUri="https://zenodo.org/record/13745475/files/figure.png" pageId="5" pageNumber="578">Fig. 2E</figureCitation>
); and irregularly branched spindles,
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long and
<quantity id="2F761DE34841B40BFF14FDDDEA12881B" box="[199,363,547,573]" metricMagnitude="-5" metricUnit="m" metricValue="7" metricValueMax="8.0" metricValueMin="6.0" pageId="5" pageNumber="578" unit="mm" value="0.07" valueMax="0.08" valueMin="0.06">0.060.08 mm</quantity>
wide (
<figureCitation id="70B5AC834841B40BFE6BFDDDE977881B" box="[440,526,547,573]" captionStart="FIGURE 2" captionStartId="4.[152,255,1540,1564]" captionTargetBox="[151,1436,181,1516]" captionTargetId="figure-24@4.[151,1436,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Heterogorgia abdita sp. nov., holotype, MZUCR 4004, sclerites: (A) collaret spindles; (B) point spindles; (C) thorns; (D-F) spindles; (G) club-like spindles; (H) crosses; (I) radiates." figureDoi="http://doi.org/10.5281/zenodo.13745475" httpUri="https://zenodo.org/record/13745475/files/figure.png" pageId="5" pageNumber="578">Fig. 2F</figureCitation>
). The inner coenenchyme contain small tuberculate radiates,
<quantity id="2F761DE34841B40BFB13FDDDEE1A881B" box="[1216,1379,547,573]" metricMagnitude="-5" metricUnit="m" metricValue="6.5" metricValueMax="7" metricValueMin="6.0" pageId="5" pageNumber="578" unit="mm" value="0.065" valueMax="0.07" valueMin="0.06">0.060.07 mm</quantity>
long and
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wide (
<figureCitation id="70B5AC834841B40BFE65FDB9E97C8847" box="[438,517,583,609]" captionStart="FIGURE 2" captionStartId="4.[152,255,1540,1564]" captionTargetBox="[151,1436,181,1516]" captionTargetId="figure-24@4.[151,1436,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Heterogorgia abdita sp. nov., holotype, MZUCR 4004, sclerites: (A) collaret spindles; (B) point spindles; (C) thorns; (D-F) spindles; (G) club-like spindles; (H) crosses; (I) radiates." figureDoi="http://doi.org/10.5281/zenodo.13745475" httpUri="https://zenodo.org/record/13745475/files/figure.png" pageId="5" pageNumber="578">Fig. 2I</figureCitation>
); and warty crosses,
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long and
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wide (
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).
</paragraph>
<paragraph id="E831B0064841B40BFF14FD95EF3888A3" blockId="5.[151,1437,151,969]" box="[199,1089,619,645]" pageId="5" pageNumber="578">
The colour of the colony is white to pale yellow preserved in ethanol (
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).
</paragraph>
<paragraph id="E831B0064841B40BFF14FD71EFAA88EB" blockId="5.[151,1437,151,969]" pageId="5" pageNumber="578">
Variability. The
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is a
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colony branching up to four times.A small branching hydroid and a polychaete tube is attached to the colony holdfast. All other characteristics are as described for the
<typeStatus id="37350EA44841B40BFBBAFD4DEFB688EB" box="[1129,1231,691,717]" pageId="5" pageNumber="578" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="E831B0064841B40BFF14FD29E9E589EF" blockId="5.[151,1437,151,969]" pageId="5" pageNumber="578">
Habitat. The colonies were found in small clusters on large rocks mingled with branching hydroids, sponges, and bryozoans; alongside other octocoral species that are more conspicuous:
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<emphasis id="DAFA6C144841B40BFC06FD05EF108932" box="[981,1129,763,788]" italics="true" pageId="5" pageNumber="578">H. verrucosa</emphasis>
</taxonomicName>
,
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<emphasis id="DAFA6C144841B40BFBABFD05EE3F8933" box="[1144,1350,763,789]" italics="true" pageId="5" pageNumber="578">Eugorgia daniana</emphasis>
<bibRefCitation id="8C1FCDF74841B40BFA9DFD05EBAB891F" author="Verrill, A. E." pageId="5" pageNumber="578" pagination="377 - 478" refId="ref5991" refString="Verrill, A. E. (1868) Notes on Radiata in the Museum of Yale College, number 6: review of the corals and polyps of the west coast of America. Transactions of the Connecticut Academy of Arts and Sciences, 2 nd Edition, 1, 377 - 478." type="journal article" year="1868">Verrill, 1868</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="2F8ECB854841B40BFF0DFCE1E929891F" authority="Verrill, 1869" authorityName="Verrill" authorityYear="1869" box="[222,592,799,825]" class="Anthozoa" family="Plexauridae" genus="Muricea" kingdom="Animalia" order="Alcyonacea" pageId="5" pageNumber="578" phylum="Cnidaria" rank="species" species="squarrosa">
<emphasis id="DAFA6C144841B40BFF0DFCE1EACF891E" box="[222,438,799,824]" italics="true" pageId="5" pageNumber="578">Muricea squarrosa</emphasis>
<bibRefCitation id="8C1FCDF74841B40BFE6DFCE1E929891F" author="Verrill, A. E." box="[446,592,799,825]" pageId="5" pageNumber="578" pagination="423 - 502" refId="ref6046" refString="Verrill, A. E. (1869) Notes on Radiata in the Museum of Yale College, Number 6: Review of the corals and polyps of the West Coast of America. Transactions of the Connecticut Academy of Arts and Sciences, 2 nd Edition, 1869, 423 - 502." type="journal article" year="1869">Verrill, 1869</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="2F8ECB854841B40BFD8FFCE1E8B9891F" authority="Verrill, 1869" authorityName="Verrill" authorityYear="1869" box="[604,960,799,825]" class="Anthozoa" family="Plexauridae" genus="Muricea" kingdom="Animalia" order="Alcyonacea" pageId="5" pageNumber="578" phylum="Cnidaria" rank="species" species="fruticosa">
<emphasis id="DAFA6C144841B40BFD8FFCE1E85E891F" box="[604,807,799,825]" italics="true" pageId="5" pageNumber="578">Muricea fruticosa</emphasis>
<bibRefCitation id="8C1FCDF74841B40BFCFCFCE1E8B9891F" author="Verrill, A. E." box="[815,960,799,825]" pageId="5" pageNumber="578" pagination="423 - 502" refId="ref6046" refString="Verrill, A. E. (1869) Notes on Radiata in the Museum of Yale College, Number 6: Review of the corals and polyps of the West Coast of America. Transactions of the Connecticut Academy of Arts and Sciences, 2 nd Edition, 1869, 423 - 502." type="journal article" year="1869">Verrill, 1869</bibRefCitation>
</taxonomicName>
,
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<emphasis id="DAFA6C144841B40BFC1EFCE1EF5D891E" box="[973,1060,799,824]" italics="true" pageId="5" pageNumber="578">P. senta</emphasis>
</taxonomicName>
,
<taxonomicName id="2F8ECB854841B40BFBE7FCE1EF9A891F" authorityName="Breedy &amp; Guzman" authorityYear="2018" box="[1076,1251,799,825]" class="Anthozoa" family="Gorgoniidae" genus="Adelogorgia" kingdom="Animalia" order="Alcyonacea" pageId="5" pageNumber="578" phylum="Cnidaria" rank="species" species="osculabunda">
<emphasis id="DAFA6C144841B40BFBE7FCE1EF9A891F" box="[1076,1251,799,825]" italics="true" pageId="5" pageNumber="578">A. osculabunda</emphasis>
</taxonomicName>
, an undescribed species of
<taxonomicName id="2F8ECB854841B40BFED9FCBDE93E897B" authority="Verrill, 1868" authorityName="Verrill" authorityYear="1868" box="[266,583,835,861]" class="Anthozoa" family="Plexauridae" genus="Psammogorgia" kingdom="Animalia" order="Alcyonacea" pageId="5" pageNumber="578" phylum="Cnidaria" rank="genus">
<emphasis id="DAFA6C144841B40BFED9FCBDEACD897A" box="[266,436,835,860]" italics="true" pageId="5" pageNumber="578">Psammogorgia</emphasis>
<bibRefCitation id="8C1FCDF74841B40BFE6AFCBDE93E897B" author="Verrill, A. E." box="[441,583,835,861]" pageId="5" pageNumber="578" pagination="377 - 478" refId="ref5991" refString="Verrill, A. E. (1868) Notes on Radiata in the Museum of Yale College, number 6: review of the corals and polyps of the west coast of America. Transactions of the Connecticut Academy of Arts and Sciences, 2 nd Edition, 1, 377 - 478." type="journal article" year="1868">Verrill, 1868</bibRefCitation>
</taxonomicName>
, and the black corals
<taxonomicName id="2F8ECB854841B40BFCE3FCBDEF94897B" authority="(Verrill, 1869)" baseAuthorityName="Verrill" baseAuthorityYear="1869" box="[816,1261,835,861]" class="Anthozoa" family="Myriopathidae" genus="Myriopathes" kingdom="Animalia" order="Antipatharia" pageId="5" pageNumber="578" phylum="Cnidaria" rank="species" species="panamensis">
<emphasis id="DAFA6C144841B40BFCE3FCBDEF3E897A" box="[816,1095,835,861]" italics="true" pageId="5" pageNumber="578">Myriopathes panamensis</emphasis>
(
<bibRefCitation id="8C1FCDF74841B40BFB87FCBDEF9C897B" author="Verrill, A. E." box="[1108,1253,835,861]" pageId="5" pageNumber="578" pagination="423 - 502" refId="ref6046" refString="Verrill, A. E. (1869) Notes on Radiata in the Museum of Yale College, Number 6: Review of the corals and polyps of the West Coast of America. Transactions of the Connecticut Academy of Arts and Sciences, 2 nd Edition, 1869, 423 - 502." type="journal article" year="1869">Verrill, 1869</bibRefCitation>
)
</taxonomicName>
, and
<taxonomicName id="2F8ECB854841B40BFAF7FCBDEA8289A7" authority="Diechmann, 1941" authorityName="Diechmann" authorityYear="1941" class="Anthozoa" family="Antipathidae" genus="Antipathes" kingdom="Animalia" order="Antipatharia" pageId="5" pageNumber="578" phylum="Cnidaria" rank="species" species="galapagensis">
<emphasis id="DAFA6C144841B40BFAF7FCBDEA5289A7" italics="true" pageId="5" pageNumber="578">Antipathes galapagensis</emphasis>
<bibRefCitation id="8C1FCDF74841B40BFEE1FC99EA8289A7" author="Diechmann, E." box="[306,507,871,897]" pageId="5" pageNumber="578" pagination="1 - 17" refId="ref5263" refString="Diechmann, E. (1941) Coelenterates collected on the Presidential Cruise of 1938. Smithsonian Miscellaneous Collections, 99 (10), 1 - 17." type="journal article" year="1941">Diechmann, 1941</bibRefCitation>
</taxonomicName>
, the latter appearing in solitary colonies close to the spot of octocorals, but making forests deeper in the shoal (Matamoros-Calderón
<emphasis id="DAFA6C144841B40BFD69FC75E9818983" box="[698,760,907,933]" italics="true" pageId="5" pageNumber="578">et. al.</emphasis>
, 2021). It was not possible to photograph the colonies
<emphasis id="DAFA6C144841B40BFA82FC75EEEC8982" box="[1361,1429,907,932]" italics="true" pageId="5" pageNumber="578">in situ</emphasis>
, thus, in
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, we show its interesting habitat.
</paragraph>
<caption id="BCF1E08E4841B40BFF4BF8ADEA928DA9" ID-DOI="http://doi.org/10.5281/zenodo.13745477" ID-Zenodo-Dep="13745477" httpUri="https://zenodo.org/record/13745477/files/figure.png" pageId="5" pageNumber="578" startId="5.[152,255,1875,1899]" targetBox="[151,1436,993,1852]" targetPageId="5" targetType="figure">
<paragraph id="E831B0064841B40BFF4BF8ADEA928DA9" blockId="5.[151,1436,1875,1936]" pageId="5" pageNumber="578">
<emphasis id="DAFA6C144841B40BFF4BF8ADEA618D4D" bold="true" box="[152,280,1875,1899]" pageId="5" pageNumber="578">FIGURE 3.</emphasis>
View of Bajo Tigre, 45 m in depth showing the habitat of
<taxonomicName id="2F8ECB854841B40BFCB1F8AAEF488D4D" authorityName="Breedy &amp; Murillo-Cruz &amp; Cortés" authorityYear="2024" box="[866,1073,1875,1899]" class="Anthozoa" family="Plexauridae" genus="Heterogorgia" kingdom="Animalia" order="Alcyonacea" pageId="5" pageNumber="578" phylum="Cnidaria" rank="species" species="abdita" status="sp. nov.">
<emphasis id="DAFA6C144841B40BFCB1F8AAEF488D4D" box="[866,1073,1875,1899]" italics="true" pageId="5" pageNumber="578">Heterogorgia abdita</emphasis>
</taxonomicName>
<emphasis id="DAFA6C144841B40BFBE5F8AAEFFF8D4A" bold="true" box="[1078,1158,1876,1900]" pageId="5" pageNumber="578">
<taxonomicNameLabel id="C1C9D16F4841B40BFBE5F8AAEFFF8D4A" box="[1078,1158,1876,1900]" pageId="5" pageNumber="578" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
. Photograph from video by Steven Lara and Beatriz Naranjo.
</paragraph>
</caption>
<paragraph id="E831B0064842B408FF14FF69E99F8A97" blockId="6.[151,1436,151,249]" box="[199,742,151,177]" pageId="6" pageNumber="579">
Distribution. Only known from the
<typeStatus id="37350EA44842B408FD86FF69E9FC8A97" box="[597,645,151,177]" pageId="6" pageNumber="579">type</typeStatus>
locality.
</paragraph>
<paragraph id="E831B0064842B408FF14FF45E9E28ADF" blockId="6.[151,1436,151,249]" pageId="6" pageNumber="579">
Etymology.
<emphasis id="DAFA6C144842B408FE82FF45EADE8AF3" box="[337,423,187,213]" italics="true" pageId="6" pageNumber="579">Abditus</emphasis>
(L), meaning hidden, concealed. The name of the coral alludes to its cryptic way of life, mingled with other small branching creatures.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,153 @@
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<mods:title id="CCF174A78ED30AFB6F34FC494E004502">Review of the genus Diduga (Lepidoptera, Erebidae, Arctiinae) of Vietnam, with a new species and two newly recorded species</mods:title>
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<paragraph id="F27EF62231181130A0A8FA2AFD17FA3C" blockId="7.[151,706,1446,1509]" box="[151,706,1446,1473]" pageId="7" pageNumber="560">
<heading id="A936414E31181130A0A8FA2AFD17FA3C" bold="true" box="[151,706,1446,1473]" fontSize="11" level="1" pageId="7" pageNumber="560" reason="1">
<taxonomicName id="35C18DA131181130A0A8FA2AFD17FA3C" ID-CoL="BXPFB" authority="Bayarsaikhan &amp; Bae, 2023" authorityName="Bayarsaikhan &amp; Bae" authorityYear="2023" box="[151,706,1446,1473]" class="Insecta" family="Erebidae" genus="Diduga" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="560" phylum="Arthropoda" rank="species" species="robdevosi">
<emphasis id="C0B52A3031181130A0A8FA2AFD17FA3C" bold="true" box="[151,706,1446,1473]" pageId="7" pageNumber="560">
<emphasis id="C0B52A3031181130A0A8FA2AFEB9FA3D" bold="true" box="[151,364,1446,1473]" italics="true" pageId="7" pageNumber="560">Diduga robdevosi</emphasis>
Bayarsaikhan &amp; Bae, 2023
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BADBA5A931181130A0A8FA47FAF3F9DB" pageId="7" pageNumber="560" type="description">
<paragraph id="F27EF62231181130A0A8FA47FEAFFA19" blockId="7.[151,706,1446,1509]" box="[151,378,1482,1509]" pageId="7" pageNumber="560">
(
<figureCitation id="6AFAEAA731181130A0A0FA47FECAFA19" box="[159,287,1483,1509]" captionStart="FIGURES 68" captionStartId="3.[151,264,1653,1677]" captionTargetBox="[151,1435,453,1630]" captionTargetId="figure-117@3.[151,1435,453,1630]" captionTargetPageId="3" captionText="FIGURES 68. Adults of the Diduga from Vietnam. 6a. D. laocai Bayarsaikhan &amp; Heppner, sp. nov., holotype, male. 6b. diito, paratype, female. 7a. D. robdevosi Bayarsaikhan &amp; Bae, male. 7b. ditto, female. 8. D. quinquicornuta Bayarsaikhan &amp; Bae, male." figureDoi="http://doi.org/10.5281/zenodo.13745449" httpUri="https://zenodo.org/record/13745449/files/figure.png" pageId="7" pageNumber="560">Figs 7a, 7b</figureCitation>
,
<figureCitation id="6AFAEAA731181130A115FA46FE9DFA19" box="[298,328,1482,1509]" captionStart="FIGURES 915" captionStartId="4.[151,264,1901,1925]" captionTargetBox="[160,1427,319,1868]" captionTargetId="figure-69@4.[151,1436,309,1878]" captionTargetPageId="4" captionText="FIGURES 915. Male genitalia of the Diduga from Vietnam. 9. D. alternota Bucsek, UB-12951Vietnam. 10. D. bayartogtokhi Bayarsaikhan&amp; Bae, UB-12958Vietnam.11.D.hanoiensis Bayarsaikhan&amp; Bae, UB-12976Vietnam.12.D.hastataBayarsaikhan &amp; Bae, UB-12966Vietnam. 13. D. nigridentata Bayarsaikhan &amp; Bae, UB-12957Vietnam. 14. D. quinquicornuta Bayarsaikhan &amp; Bae, UB-12962Vietnam. 15. D. robdevosi Bayarsaikhan &amp; Bae, UB-12970Vietnam." figureDoi="http://doi.org/10.5281/zenodo.13745453" httpUri="https://zenodo.org/record/13745453/files/figure.png" pageId="7" pageNumber="560">15</figureCitation>
,
<figureCitation id="6AFAEAA731181130A16BFA47FEA4FA19" box="[340,369,1483,1509]" captionStart="FIGURES 1822" captionStartId="8.[151,264,1908,1932]" captionTargetBox="[161,1429,571,1875]" captionTargetId="figure-157@8.[151,1436,561,1885]" captionTargetPageId="8" captionText="FIGURES 1822. Female genitalia of the Diduga from Vietnam. 18. D. alternota Bucsek, UB-12981Vietnam. 19. D. bayartogtokhi Bayarsaikhan &amp;Bae, UB-12965Vietnam.20.D.hanoiensis Bayarsaikhan &amp;Bae, UB-12969Vietnam.21.D.laocai Bayarsaikhan &amp; Heppner, sp. nov., paratype, UB-12956Vietnam. 22. D. robdevosi Bayarsaikhan &amp; Bae, UB-12971Vietnam." figureDoi="http://doi.org/10.5281/zenodo.13745459" httpUri="https://zenodo.org/record/13745459/files/figure.png" pageId="7" pageNumber="560">22</figureCitation>
)
</paragraph>
<paragraph id="F27EF62231181130A0A8F983FAF3F9DB" blockId="7.[151,1318,1551,1575]" box="[151,1318,1551,1575]" pageId="7" pageNumber="560">
<taxonomicName id="35C18DA131181130A0A8F983FDB3F9DB" authority="Bayarsaikhan &amp; Bae, 2023" authorityName="Bayarsaikhan &amp; Bae" authorityYear="2023" box="[151,614,1551,1575]" class="Insecta" family="Erebidae" genus="Diduga" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="560" phylum="Arthropoda" rank="species" species="robdevosi">
<emphasis id="C0B52A3031181130A0A8F983FE98F9DB" box="[151,333,1551,1575]" italics="true" pageId="7" pageNumber="560">Diduga robdevosi</emphasis>
Bayarsaikhan &amp; Bae, 2023
</taxonomicName>
,
<emphasis id="C0B52A3031181130A24FF99CFD6EF9DB" box="[624,699,1552,1575]" italics="true" pageId="7" pageNumber="560">Forests</emphasis>
2023, 14: 889. TL:
<collectingCountry id="8AD6B6B231181130A3B9F99CFC62F9DB" box="[902,951,1552,1575]" name="Laos" pageId="7" pageNumber="560">Laos</collectingCountry>
(
<collectingRegion id="300538C031181130A3FAF983FB9CF9DB" box="[965,1097,1551,1575]" country="Laos" name="Bolikhamxai" pageId="7" pageNumber="560">Bolikhamsai</collectingRegion>
, Tad Leuk Waterfall).
</paragraph>
</subSubSection>
<subSubSection id="BADBA5A931181130A0A8F9D9FF2CF927" pageId="7" pageNumber="560" type="distribution">
<paragraph id="F27EF62231181130A0A8F9D9FD38F993" blockId="7.[151,1436,1621,1792]" box="[151,749,1621,1648]" pageId="7" pageNumber="560">
<emphasis id="C0B52A3031181130A0A8F9D9FEE4F993" bold="true" box="[151,305,1621,1647]" pageId="7" pageNumber="560">Distribution.</emphasis>
<collectingCountry id="8AD6B6B231181130A108F9D9FE43F993" box="[311,406,1621,1647]" name="Vietnam" pageId="7" pageNumber="560">Vietnam</collectingCountry>
(new record),
<collectingCountry id="8AD6B6B231181130A205F9DAFDA6F98C" box="[570,627,1622,1648]" name="Laos" pageId="7" pageNumber="560">Laos</collectingCountry>
,
<collectingCountry id="8AD6B6B231181130A240F9D9FD3CF993" box="[639,745,1621,1647]" name="Malaysia" pageId="7" pageNumber="560">Malaysia</collectingCountry>
.
</paragraph>
<paragraph id="F27EF62231181130A0F8F9F5FF2CF927" blockId="7.[151,1436,1621,1792]" pageId="7" pageNumber="560">
<emphasis id="C0B52A3031181130A0F8F9F5FDE0F96F" bold="true" box="[199,565,1657,1683]" pageId="7" pageNumber="560">Additional material examined.</emphasis>
<materialsCitation id="42A9FC7F31181130A202F9F5FB82F94B" collectingDate="2008-06-21" collectingDateMax="2008-06-26" collectingDateMin="2008-06-21" collectionCode="UB" collectorName="Ba Be N. P. &amp; J. B. Heppner" country="Vietnam" elevation="240" location="Bac Kan" pageId="7" pageNumber="560" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Bac Kan">
<specimenCount id="E4C73DAB31181130A202F9F5FDB4F96F" box="[573,609,1657,1683]" count="1" pageId="7" pageNumber="560" type="male">1♂</specimenCount>
,
<specimenCount id="E4C73DAB31181130A251F9F5FD44F96F" box="[622,657,1657,1683]" count="1" pageId="7" pageNumber="560" type="female">1♀</specimenCount>
,
<collectingCountry id="8AD6B6B231181130A2A0F9F5FCD4F96F" box="[671,769,1657,1683]" name="Vietnam" pageId="7" pageNumber="560">Vietnam</collectingCountry>
,
<collectingRegion id="300538C031181130A332F9F6FCA4F968" box="[781,881,1658,1684]" country="Vietnam" name="Bac Kan" pageId="7" pageNumber="560">Bac Kan</collectingRegion>
,
<collectorName id="5F3493F431181130A342F9F6FC2CF968" box="[893,1017,1658,1684]" pageId="7" pageNumber="560">Ba Be N.P.</collectorName>
(lodge),
<quantity id="35395BC731181130A45FF9F5FB77F96F" box="[1120,1186,1657,1683]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="7" pageNumber="560" unit="m" value="240.0">
<elevation id="79EC111131181130A45FF9F5FB77F96F" box="[1120,1186,1657,1683]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="7" pageNumber="560" unit="m" value="240.0">240m</elevation>
</quantity>
,
<date id="867FD0E231181130A491F9F5FAB0F96F" box="[1198,1381,1657,1684]" pageId="7" pageNumber="560" value="2008-06-21" valueMax="2008-06-26" valueMin="2008-06-21">
<collectingDate id="963B290A31181130A491F9F5FAB0F96F" box="[1198,1381,1657,1684]" pageId="7" pageNumber="560" value="2008-06-21" valueMax="2008-06-26" valueMin="2008-06-21">2126 Jun 2008</collectingDate>
</date>
,
<collectorName id="5F3493F431181130A54EF9F6FF2CF944" pageId="7" pageNumber="560">J.B. Heppner</collectorName>
, adults no. FLMNH-MGCL1174185, 1174186 and genitalia slides no.
<collectionCode id="94D06EE731181130A40FF911FB82F94B" box="[1072,1111,1693,1719]" country="France" name="Laboratoire de Biostratigraphie" pageId="7" pageNumber="560">UB</collectionCode>
</materialsCitation>
˗12970, 12971 (deposited in MGCL).
</paragraph>
</subSubSection>
<subSubSection id="BADBA5A931181130A0F8F969FBA3F903" box="[199,1142,1765,1792]" pageId="7" pageNumber="560" type="discussion">
<paragraph id="F27EF62231181130A0F8F969FBA3F903" blockId="7.[151,1436,1621,1792]" box="[199,1142,1765,1792]" pageId="7" pageNumber="560">
<emphasis id="C0B52A3031181130A0F8F969FEEEF903" bold="true" box="[199,315,1765,1791]" pageId="7" pageNumber="560">Remarks.</emphasis>
This species is herein newly recorded from
<collectingCountry id="8AD6B6B231181130A315F969FC5CF903" box="[810,905,1765,1791]" name="Vietnam" pageId="7" pageNumber="560">Vietnam</collectingCountry>
(
<collectingRegion id="300538C031181130A3A7F969FBBFF903" box="[920,1130,1765,1792]" country="Vietnam" name="Bac Kan" pageId="7" pageNumber="560">Bac Kan Province</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,229 @@
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<mods:title id="954CEA7EEEFDB1E1D6E94E81137320D2">Review of the genus Diduga (Lepidoptera, Erebidae, Arctiinae) of Vietnam, with a new species and two newly recorded species</mods:title>
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<mods:namePart id="4A2A03F443CF8909A46B5D03A852D791">Kwon, Hyung Wook</mods:namePart>
<mods:affiliation id="D60370DC7B62544F62BEE58FF83D2659">Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. &amp; Bio ˗ Resource and Environmental Center, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea.</mods:affiliation>
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<mods:affiliation id="F39BEBD31C77DB44B14646D2E509B8A6">Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. &amp; Bio ˗ Resource and Environmental Center, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. &amp; Convergence Research Center for Insect Vectors, Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, Songdo ˗ dong, Incheon, 22012, Republic of Korea.</mods:affiliation>
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<heading id="A936414E31181130A0A8F8DDFCDEF897" bold="true" box="[151,779,1873,1900]" fontSize="11" level="1" pageId="7" pageNumber="560" reason="1">
<taxonomicName id="35C18DA131181130A0A8F8DDFCDEF897" ID-CoL="BHR9V" authority="Bayarsaikhan &amp; Bae, 2019" authorityName="Bayarsaikhan &amp; Bae" authorityYear="2019" box="[151,779,1873,1900]" class="Insecta" family="Erebidae" genus="Diduga" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="560" phylum="Arthropoda" rank="species" species="quinquicornuta">
<emphasis id="C0B52A3031181130A0A8F8DDFCDEF897" bold="true" box="[151,779,1873,1900]" pageId="7" pageNumber="560">
<emphasis id="C0B52A3031181130A0A8F8DDFE60F897" bold="true" box="[151,437,1873,1900]" italics="true" pageId="7" pageNumber="560">Diduga quinquicornuta</emphasis>
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<paragraph id="F27EF62231181130A0A8F8F9FECFF873" blockId="7.[151,779,1873,1935]" box="[151,282,1909,1935]" pageId="7" pageNumber="560">
(
<figureCitation id="6AFAEAA731181130A0A0F8F9FF32F873" box="[159,231,1909,1935]" captionStart="FIGURES 68" captionStartId="3.[151,264,1653,1677]" captionTargetBox="[151,1435,453,1630]" captionTargetId="figure-117@3.[151,1435,453,1630]" captionTargetPageId="3" captionText="FIGURES 68. Adults of the Diduga from Vietnam. 6a. D. laocai Bayarsaikhan &amp; Heppner, sp. nov., holotype, male. 6b. diito, paratype, female. 7a. D. robdevosi Bayarsaikhan &amp; Bae, male. 7b. ditto, female. 8. D. quinquicornuta Bayarsaikhan &amp; Bae, male." figureDoi="http://doi.org/10.5281/zenodo.13745449" httpUri="https://zenodo.org/record/13745449/files/figure.png" pageId="7" pageNumber="560">Figs 8</figureCitation>
,
<figureCitation id="6AFAEAA731181130A0CBF8F9FEC4F873" box="[244,273,1909,1935]" captionStart="FIGURES 915" captionStartId="4.[151,264,1901,1925]" captionTargetBox="[160,1427,319,1868]" captionTargetId="figure-69@4.[151,1436,309,1878]" captionTargetPageId="4" captionText="FIGURES 915. Male genitalia of the Diduga from Vietnam. 9. D. alternota Bucsek, UB-12951Vietnam. 10. D. bayartogtokhi Bayarsaikhan&amp; Bae, UB-12958Vietnam.11.D.hanoiensis Bayarsaikhan&amp; Bae, UB-12976Vietnam.12.D.hastataBayarsaikhan &amp; Bae, UB-12966Vietnam. 13. D. nigridentata Bayarsaikhan &amp; Bae, UB-12957Vietnam. 14. D. quinquicornuta Bayarsaikhan &amp; Bae, UB-12962Vietnam. 15. D. robdevosi Bayarsaikhan &amp; Bae, UB-12970Vietnam." figureDoi="http://doi.org/10.5281/zenodo.13745453" httpUri="https://zenodo.org/record/13745453/files/figure.png" pageId="7" pageNumber="560">14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="BADBA5A931181130A0A8F836FA85F82E" box="[151,1360,1978,2002]" pageId="7" pageNumber="560" type="reference_group">
<paragraph id="F27EF62231181130A0A8F836FA85F82E" blockId="7.[151,1360,1978,2002]" box="[151,1360,1978,2002]" pageId="7" pageNumber="560">
<taxonomicName id="35C18DA131181130A0A8F836FD4AF82E" ID-CoL="BHR9V" authority="Bayarsaikhan &amp; Bae, 2019" authorityName="Bayarsaikhan &amp; Bae" authorityYear="2019" box="[151,671,1978,2002]" class="Insecta" family="Erebidae" genus="Diduga" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="560" phylum="Arthropoda" rank="species" species="quinquicornuta">
<emphasis id="C0B52A3031181130A0A8F836FE53F82D" box="[151,390,1978,2002]" italics="true" pageId="7" pageNumber="560">Diduga quinquicornuta</emphasis>
<bibRefCitation id="96508BD331181130A1B3F836FD4AF82E" author="Bayarsaikhan, U. &amp; Bae, Y. S." box="[396,671,1978,2002]" pageId="7" pageNumber="560" pagination="365 - 376" refId="ref4259" refString="Bayarsaikhan, U. &amp; Bae, Y. S. (2019) Four new and one newly recorded species of Diduga Moore, [1887] (Lepidoptera, Erebidae, Arctiinae) from Vietnam, with redescription of the little known species Diduga haematomiformis van Eecke, 1920. Zootaxa, 4624 (3), 365 - 376. https: // doi. org / 10.11646 / zootaxa. 4624.3.5" type="journal article" year="2019">Bayarsaikhan &amp; Bae, 2019</bibRefCitation>
</taxonomicName>
,
<emphasis id="C0B52A3031181130A295F837FD28F82E" box="[682,765,1979,2002]" italics="true" pageId="7" pageNumber="560">Zootaxa</emphasis>
4624 (3): 370, figs. 6, 12, 18. TL:
<collectingCountry id="8AD6B6B231181130A462F836FB66F82E" box="[1117,1203,1978,2002]" name="Vietnam" pageId="7" pageNumber="560">Vietnam</collectingCountry>
(
<collectingRegion id="300538C031181130A4FFF836FAD7F82E" box="[1216,1282,1978,2002]" country="Vietnam" name="Ha Noi" pageId="7" pageNumber="560">Hanoi</collectingRegion>
, BaVi).
</paragraph>
</subSubSection>
<subSubSection id="BADBA5A93117113FA0A8FF1AFDDAFE51" pageId="8" pageNumber="561" type="distribution">
<paragraph id="F27EF6223117113FA0A8FF1AFE48FF4D" blockId="8.[151,1437,150,501]" box="[151,413,150,177]" pageId="8" pageNumber="561">
<emphasis id="C0B52A303117113FA0A8FF1AFEE4FF4C" bold="true" box="[151,305,150,176]" pageId="8" pageNumber="561">Distribution.</emphasis>
<collectingCountry id="8AD6B6B23117113FA108FF1BFE4FFF4D" box="[311,410,151,177]" name="Vietnam" pageId="8" pageNumber="561">Vietnam</collectingCountry>
.
</paragraph>
<paragraph id="F27EF6223117113FA0F8FF36FDDAFE51" blockId="8.[151,1437,150,501]" pageId="8" pageNumber="561">
<emphasis id="C0B52A303117113FA0F8FF36FDE2FF28" bold="true" box="[199,567,186,212]" pageId="8" pageNumber="561">Additional material examined.</emphasis>
<materialsCitation id="42A9FC7F3117113FA200FF37FB8DFF05" collectingDate="2010-07-19" collectingDateMax="2010-07-20" collectingDateMin="2010-07-19" collectionCode="UB" collectorName="J. B. Heppner" country="Vietnam" county="Prov." elevation="1080" location="Pk." municipality="Ba Vi Natl." pageId="8" pageNumber="561" specimenCount="2" specimenCount-male="2">
<specimenCount id="E4C73DAB3117113FA200FF37FDB0FF29" box="[575,613,187,213]" count="2" pageId="8" pageNumber="561" type="male">2♂</specimenCount>
,
<collectingCountry id="8AD6B6B23117113FA24EFF37FD06FF29" box="[625,723,187,213]" name="Vietnam" pageId="8" pageNumber="561">Vietnam</collectingCountry>
, Hatay
<collectingCounty id="1B1F8EAE3117113FA314FF37FCBDFF29" box="[811,872,187,213]" pageId="8" pageNumber="561">Prov.</collectingCounty>
,
<collectingMunicipality id="121A6C583117113FA34BFF37FC2DFF29" box="[884,1016,187,213]" pageId="8" pageNumber="561">Ba Vi Natl.</collectingMunicipality>
,
<location id="F71EA0F93117113FA43AFF37FBF9FF29" LSID="urn:lsid:plazi:treatment:7A6847343118113FA0A8F8DDFB5DFE09:F71EA0F93117113FA43AFF37FBF9FF29" box="[1029,1068,187,213]" country="Vietnam" county="Prov." municipality="Ba Vi Natl." name="Pk." pageId="8" pageNumber="561">Pk.</location>
,
<quantity id="35395BC73117113FA406FF37FB5CFF29" box="[1081,1161,187,213]" metricMagnitude="3" metricUnit="m" metricValue="1.08" pageId="8" pageNumber="561" unit="m" value="1080.0">
<elevation id="79EC11113117113FA406FF37FB5CFF29" box="[1081,1161,187,213]" metricMagnitude="3" metricUnit="m" metricValue="1.08" pageId="8" pageNumber="561" unit="m" value="1080.0">1080m</elevation>
</quantity>
,
<date id="867FD0E23117113FA4A9FF37FA9CFF29" box="[1174,1353,187,213]" pageId="8" pageNumber="561" value="2010-07-19" valueMax="2010-07-20" valueMin="2010-07-19">
<collectingDate id="963B290A3117113FA4A9FF37FA9CFF29" box="[1174,1353,187,213]" pageId="8" pageNumber="561" value="2010-07-19" valueMax="2010-07-20" valueMin="2010-07-19">
1920 Jul
<quantity id="35395BC73117113FA531FF37FA9CFF29" box="[1294,1353,187,213]" metricMagnitude="3" metricUnit="m" metricValue="2.01093" pageId="8" pageNumber="561" unit="m" value="2010.93">2010</quantity>
</collectingDate>
</date>
, 930m,
<collectorName id="5F3493F43117113FA0A8FF53FEFEFF05" box="[151,299,223,249]" pageId="8" pageNumber="561">J.B. Heppner</collectorName>
, adults no. FLMNH-MGCL1174130, 1174134 and genitalia slide no.
<collectionCode id="94D06EE73117113FA40FFF52FB8DFF05" box="[1072,1112,222,249]" country="France" name="Laboratoire de Biostratigraphie" pageId="8" pageNumber="561">UB</collectionCode>
</materialsCitation>
˗12959, 12963;
<materialsCitation id="42A9FC7F3117113FA538FF53FE17FEBD" collectingDate="2010-07-19" collectingDateMax="2010-07-23" collectingDateMin="2010-07-19" collectionCode="UB" collectorName="J. B. Heppner &amp; Y. - S. Bae" country="Vietnam" elevation="455" location="Ba Vi Natl. Pk." municipality="Prov." pageId="8" pageNumber="561" specimenCount="1" specimenCount-male="1">
<specimenCount id="E4C73DAB3117113FA538FF53FAFEFF05" box="[1287,1323,223,249]" count="1" pageId="8" pageNumber="561" type="male">1♂</specimenCount>
,
<collectingCountry id="8AD6B6B23117113FA509FF53FA4DFF05" box="[1334,1432,223,249]" name="Vietnam" pageId="8" pageNumber="561">Vietnam</collectingCountry>
, Hatay
<collectingMunicipality id="121A6C583117113FA0DEFE8FFECBFEE1" box="[225,286,259,285]" pageId="8" pageNumber="561">Prov.</collectingMunicipality>
,
<location id="F71EA0F93117113FA116FE8FFE00FEE1" LSID="urn:lsid:plazi:treatment:7A6847343118113FA0A8F8DDFB5DFE09:F71EA0F93117113FA116FE8FFE00FEE1" box="[297,469,259,285]" country="Vietnam" municipality="Prov." name="Ba Vi Natl. Pk." pageId="8" pageNumber="561">Ba Vi Natl. Pk.</location>
,
<quantity id="35395BC73117113FA1DFFE8EFDF6FEE1" box="[480,547,258,285]" metricMagnitude="2" metricUnit="m" metricValue="4.55" pageId="8" pageNumber="561" unit="m" value="455.0">
<elevation id="79EC11113117113FA1DFFE8EFDF6FEE1" box="[480,547,258,285]" metricMagnitude="2" metricUnit="m" metricValue="4.55" pageId="8" pageNumber="561" unit="m" value="455.0">455m</elevation>
</quantity>
,
<date id="867FD0E23117113FA211FE8FFD0BFEE1" box="[558,734,259,285]" pageId="8" pageNumber="561" value="2010-07-19" valueMax="2010-07-23" valueMin="2010-07-19">
<collectingDate id="963B290A3117113FA211FE8FFD0BFEE1" box="[558,734,259,285]" pageId="8" pageNumber="561" value="2010-07-19" valueMax="2010-07-23" valueMin="2010-07-19">1923 Jul 2010</collectingDate>
</date>
,
<collectorName id="5F3493F43117113FA2D6FE8FFCAEFEE1" box="[745,891,259,285]" pageId="8" pageNumber="561">J.B. Heppner</collectorName>
&amp;
<collectorName id="5F3493F43117113FA3A1FE8FFBDEFEE1" box="[926,1035,259,285]" pageId="8" pageNumber="561">Y.-S. Bae</collectorName>
, adult no. FLMNH-MGCL1174193 and genitalia slide no.
<collectionCode id="94D06EE73117113FA1A5FEABFE17FEBD" box="[410,450,295,321]" country="France" name="Laboratoire de Biostratigraphie" pageId="8" pageNumber="561">UB</collectionCode>
</materialsCitation>
˗12977;
<materialsCitation id="42A9FC7F3117113FA21EFEABFBF0FE99" collectingDate="2010-07-22" collectionCode="UB" collectorName="J. B. Heppner &amp; Y. - S. Bae" country="Vietnam" elevation="1080" location="Ba Vi Natl. Pk." municipality="Prov." pageId="8" pageNumber="561" specimenCount="1" specimenCount-male="1">
<specimenCount id="E4C73DAB3117113FA21EFEABFD90FEBD" box="[545,581,295,321]" count="1" pageId="8" pageNumber="561" type="male">1♂</specimenCount>
,
<collectingCountry id="8AD6B6B23117113FA26CFEABFD60FEBD" box="[595,693,295,321]" name="Vietnam" pageId="8" pageNumber="561">Vietnam</collectingCountry>
, Hatay
<collectingMunicipality id="121A6C583117113FA332FEABFC9FFEBD" box="[781,842,295,321]" pageId="8" pageNumber="561">Prov.</collectingMunicipality>
,
<location id="F71EA0F93117113FA368FEABFBDDFEBD" LSID="urn:lsid:plazi:treatment:7A6847343118113FA0A8F8DDFB5DFE09:F71EA0F93117113FA368FEABFBDDFEBD" box="[855,1032,295,321]" country="Vietnam" municipality="Prov." name="Ba Vi Natl. Pk." pageId="8" pageNumber="561">Ba Vi Natl. Pk.</location>
,
<quantity id="35395BC73117113FA42AFEABFBB0FEBD" box="[1045,1125,295,321]" metricMagnitude="3" metricUnit="m" metricValue="1.08" pageId="8" pageNumber="561" unit="m" value="1080.0">
<elevation id="79EC11113117113FA42AFEABFBB0FEBD" box="[1045,1125,295,321]" metricMagnitude="3" metricUnit="m" metricValue="1.08" pageId="8" pageNumber="561" unit="m" value="1080.0">1080m</elevation>
</quantity>
,
<date id="867FD0E23117113FA44DFEABFB2EFEBD" box="[1138,1275,295,321]" pageId="8" pageNumber="561" value="2010-07-22">
<collectingDate id="963B290A3117113FA44DFEABFB2EFEBD" box="[1138,1275,295,321]" pageId="8" pageNumber="561" value="2010-07-22">22 Jul 2010</collectingDate>
</date>
,
<collectorName id="5F3493F43117113FA537FEABFA49FEBD" box="[1288,1436,295,321]" pageId="8" pageNumber="561">J.B. Heppner</collectorName>
&amp;
<collectorName id="5F3493F43117113FA085FEC7FEFBFE99" box="[186,302,331,357]" pageId="8" pageNumber="561">Y.-S. Bae</collectorName>
, adult no. FLMNH-MGCL1174195 and genitalia slide no.
<collectionCode id="94D06EE73117113FA3C2FEC7FBF0FE99" box="[1021,1061,331,357]" country="France" name="Laboratoire de Biostratigraphie" pageId="8" pageNumber="561">UB</collectionCode>
</materialsCitation>
˗12979;
<materialsCitation id="42A9FC7F3117113FA4B6FEC7FF6BFE51" collectingDate="2016-08-09" collectingDateMax="2016-08-11" collectingDateMin="2016-08-09" collectionCode="UB" collectorName="J. B. Heppner" country="Vietnam" location="MuongKhuong" pageId="8" pageNumber="561" specimenCount="1" specimenCount-female="1" stateProvince="Lao Cai">
<specimenCount id="E4C73DAB3117113FA4B6FEC7FB79FE99" box="[1161,1196,331,357]" count="1" pageId="8" pageNumber="561" type="female">1♀</specimenCount>
,
<collectingCountry id="8AD6B6B23117113FA4FEFEC7FAF6FE99" box="[1217,1315,331,357]" name="Vietnam" pageId="8" pageNumber="561">Vietnam</collectingCountry>
,
<collectingRegion id="300538C03117113FA50AFEC7FA4DFE99" box="[1333,1432,331,357]" country="Vietnam" name="Lao Cai" pageId="8" pageNumber="561">Lao Cai</collectingRegion>
,
<location id="F71EA0F93117113FA0A8FEE3FE90FE75" LSID="urn:lsid:plazi:treatment:7A6847343118113FA0A8F8DDFB5DFE09:F71EA0F93117113FA0A8FEE3FE90FE75" box="[151,325,367,393]" country="Vietnam" name="MuongKhuong" pageId="8" pageNumber="561" stateProvince="Lao Cai">MuongKhuong</location>
,
<date id="867FD0E23117113FA16CFEE3FDD2FE75" box="[339,519,367,393]" pageId="8" pageNumber="561" value="2016-08-09" valueMax="2016-08-11" valueMin="2016-08-09">
<collectingDate id="963B290A3117113FA16CFEE3FDD2FE75" box="[339,519,367,393]" pageId="8" pageNumber="561" value="2016-08-09" valueMax="2016-08-11" valueMin="2016-08-09">
911 Aug
<quantity id="35395BC73117113FA1F3FEE3FDD2FE75" box="[460,519,367,393]" metricMagnitude="3" metricUnit="m" metricValue="2.0169249999999996" pageId="8" pageNumber="561" unit="m" value="2016.925">2016</quantity>
</collectingDate>
</date>
, 925m,
<collectorName id="5F3493F43117113FA25AFEE3FD29FE75" box="[613,764,367,393]" pageId="8" pageNumber="561">J.B. Heppner</collectorName>
, adult no. FLMNH-MGCL1174133 and genitalia slide no.
<collectionCode id="94D06EE73117113FA0A8FE1FFF6BFE51" box="[151,190,403,429]" country="France" name="Laboratoire de Biostratigraphie" pageId="8" pageNumber="561">UB</collectionCode>
</materialsCitation>
˗12962 (deposited in MGCL).
</paragraph>
</subSubSection>
<subSubSection id="BADBA5A93117113FA0F8FE3BFB5DFE09" pageId="8" pageNumber="561" type="discussion">
<paragraph id="F27EF6223117113FA0F8FE3BFB5DFE09" blockId="8.[151,1437,150,501]" pageId="8" pageNumber="561">
<emphasis id="C0B52A303117113FA0F8FE3BFEEEFE2D" bold="true" box="[199,315,439,465]" pageId="8" pageNumber="561">Remarks.</emphasis>
This species was reported from
<collectingCountry id="8AD6B6B23117113FA2F2FE3BFCE3FE2D" box="[717,822,439,465]" name="Vietnam" pageId="8" pageNumber="561">Vietnam</collectingCountry>
(
<collectingCountry id="8AD6B6B23117113FA375FE3BFC42FE2D" box="[842,919,439,465]" name="Turkey" pageId="8" pageNumber="561">Hatay</collectingCountry>
(
<collectingRegion id="300538C03117113FA395FE3BFC2FFE2D" box="[938,1018,439,465]" country="Vietnam" name="Ha Noi" pageId="8" pageNumber="561">Hanoi</collectingRegion>
) Province) by
<bibRefCitation id="96508BD33117113FA483FE3BFEFDFE09" author="Bayarsaikhan, U. &amp; Bae, Y. S." pageId="8" pageNumber="561" pagination="365 - 376" refId="ref4259" refString="Bayarsaikhan, U. &amp; Bae, Y. S. (2019) Four new and one newly recorded species of Diduga Moore, [1887] (Lepidoptera, Erebidae, Arctiinae) from Vietnam, with redescription of the little known species Diduga haematomiformis van Eecke, 1920. Zootaxa, 4624 (3), 365 - 376. https: // doi. org / 10.11646 / zootaxa. 4624.3.5" type="journal article" year="2019">Bayarsaikhan and Bae (2019)</bibRefCitation>
. In this study, we report it for
<collectingRegion id="300538C03117113FA244FE57FCEAFE09" box="[635,831,475,501]" country="Vietnam" name="Lao Cai" pageId="8" pageNumber="561">Lao Cai Province</collectingRegion>
of
<collectingCountry id="8AD6B6B23117113FA35BFE57FC16FE09" box="[868,963,475,501]" name="Vietnam" pageId="8" pageNumber="561">Vietnam</collectingCountry>
for the first time.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,271 @@
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<mods:title id="23C20F843C2A75F70E07314DDAD629BB">Review of the genus Diduga (Lepidoptera, Erebidae, Arctiinae) of Vietnam, with a new species and two newly recorded species</mods:title>
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<mods:role id="EC7C81DB6765F15CE0B623040AED6914">
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<mods:namePart id="191D1BD838FE66C6183429F265E40E99">Bayarsaikhan, Ulziijargal</mods:namePart>
<mods:affiliation id="1D7FAD1721C4D4D0BAFCE6CD90ABD212">Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. &amp; Bio ˗ Resource and Environmental Center, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea.</mods:affiliation>
</mods:name>
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<mods:namePart id="4473AD62FB364C0EA7DEEF4C8B62FAA7">Heppner, John B.</mods:namePart>
<mods:affiliation id="E3967FEB64220DCA2070B4616EB8D023">McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Gainesville, Florida, USA.</mods:affiliation>
</mods:name>
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<mods:role id="BC21267770D16CEC70F34CF4A55B0EAC">
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<mods:namePart id="76AF49492C94C127C86FD4412360027E">Kwon, Hyung Wook</mods:namePart>
<mods:affiliation id="19B0CE8DDBF1257BF4C48709AE5AAEA4">Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. &amp; Bio ˗ Resource and Environmental Center, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea.</mods:affiliation>
</mods:name>
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<mods:namePart id="DDB6BD9326B497341DF3613EE598A421">Bae, Yang-Seop</mods:namePart>
<mods:nameIdentifier id="8D1857A5360F5A3DED38CEBB288A46CB" type="ORCID">0000-0001-7356-5633</mods:nameIdentifier>
<mods:affiliation id="4114AED226C8C97099C368AED231F6F1">Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. &amp; Bio ˗ Resource and Environmental Center, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. &amp; Convergence Research Center for Insect Vectors, Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, Songdo ˗ dong, Incheon, 22012, Republic of Korea.</mods:affiliation>
<mods:nameIdentifier id="8AC8348E315AD10A05781ED391CA48F6" type="email">baeys@inu.ac.kr</mods:nameIdentifier>
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<mods:title id="AAD4A103F809A027171B225D83E7EE98">Zootaxa</mods:title>
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<mods:part id="31ED2950B0066E1A7D7C713D962E57A7">
<mods:date id="E28B05C480BA97CD35819C898A185B73">2024</mods:date>
<mods:detail id="532E51150B66A3B17DF90CD1EB50528A" type="pubDate">
<mods:number id="49125F10559C6F374817C2E99BC23DF9">2024-09-03</mods:number>
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<mods:number id="86E33385A0C610362C24F04529AFE218">5501</mods:number>
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<mods:url id="0CAD7490381438C34A7DAA35C8DEA87A">http://dx.doi.org/10.11646/zootaxa.5501.4.5</mods:url>
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<mods:identifier id="DD06A7375E1157561761214BE4D86902" type="DOI">10.11646/zootaxa.5501.4.5</mods:identifier>
<mods:identifier id="6AEA19129E3DF41F550076AD15C0895A" type="ISSN">1175-5326</mods:identifier>
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<subSubSection id="BADBA5A9311A1132A0A8F95DFD22F910" box="[151,759,1745,1772]" pageId="5" pageNumber="558" type="nomenclature">
<paragraph id="F27EF622311A1132A0A8F95DFD22F910" blockId="5.[151,1067,1745,1845]" box="[151,759,1745,1772]" pageId="5" pageNumber="558">
<heading id="A936414E311A1132A0A8F95DFD22F910" bold="true" box="[151,759,1745,1772]" fontSize="11" level="1" pageId="5" pageNumber="558" reason="1">
<emphasis id="C0B52A30311A1132A0A8F95DFD22F910" bold="true" box="[151,759,1745,1772]" pageId="5" pageNumber="558">
<taxonomicName id="35C18DA1311A1132A0A8F95DFD59F910" authority="Bayarsaikhan &amp; Heppner" authorityName="Bayarsaikhan &amp; Heppner &amp; Kwon &amp; Bae" authorityYear="2024" box="[151,652,1745,1772]" class="Insecta" family="Erebidae" genus="Diduga" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="558" phylum="Arthropoda" rank="species" species="laocai" status="sp. nov.">
<emphasis id="C0B52A30311A1132A0A8F95DFE94F910" bold="true" box="[151,321,1745,1772]" italics="true" pageId="5" pageNumber="558">Diduga laocai</emphasis>
Bayarsaikhan &amp; Heppner
</taxonomicName>
,
<taxonomicNameLabel id="DB86974B311A1132A2A6F95EFD22F910" box="[665,759,1746,1772]" pageId="5" pageNumber="558" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BADBA5A9311A1132A0A8F97AFBFEF8C9" pageId="5" pageNumber="558" type="description">
<paragraph id="F27EF622311A1132A0A8F97AFEAFF8EC" blockId="5.[151,1067,1745,1845]" box="[151,378,1782,1808]" pageId="5" pageNumber="558">
(
<figureCitation id="6AFAEAA7311A1132A0A0F97AFECAF8EC" box="[159,287,1782,1808]" captionStart="FIGURES 68" captionStartId="3.[151,264,1653,1677]" captionTargetBox="[151,1435,453,1630]" captionTargetId="figure-117@3.[151,1435,453,1630]" captionTargetPageId="3" captionText="FIGURES 68. Adults of the Diduga from Vietnam. 6a. D. laocai Bayarsaikhan &amp; Heppner, sp. nov., holotype, male. 6b. diito, paratype, female. 7a. D. robdevosi Bayarsaikhan &amp; Bae, male. 7b. ditto, female. 8. D. quinquicornuta Bayarsaikhan &amp; Bae, male." figureDoi="http://doi.org/10.5281/zenodo.13745449" httpUri="https://zenodo.org/record/13745449/files/figure.png" pageId="5" pageNumber="558">Figs 6a, 6b</figureCitation>
,
<figureCitation id="6AFAEAA7311A1132A115F97AFE9DF8EC" box="[298,328,1782,1808]" captionStart="FIGURES 1617" captionStartId="6.[151,264,1843,1867]" captionTargetBox="[161,1425,190,1809]" captionTargetId="figure-24@6.[151,1436,181,1819]" captionTargetPageId="6" captionText="FIGURES 1617. Male genitalia of the new Diduga species from Vietnam and compared species. 16. D. laocai Bayarsaikhan &amp; Heppner, sp. nov., holotype, UB-12955Vietnam. 17. D. mucronata Bayarsaikhan &amp; Bae, holotype, INU- 10172Laos (after Bayarsaikhan et al. 2021)." figureDoi="http://doi.org/10.5281/zenodo.13745455" httpUri="https://zenodo.org/record/13745455/files/figure.png" pageId="5" pageNumber="558">16</figureCitation>
,
<figureCitation id="6AFAEAA7311A1132A16BF97AFEA4F8EC" box="[340,369,1782,1808]" captionStart="FIGURES 1822" captionStartId="8.[151,264,1908,1932]" captionTargetBox="[161,1429,571,1875]" captionTargetId="figure-157@8.[151,1436,561,1885]" captionTargetPageId="8" captionText="FIGURES 1822. Female genitalia of the Diduga from Vietnam. 18. D. alternota Bucsek, UB-12981Vietnam. 19. D. bayartogtokhi Bayarsaikhan &amp;Bae, UB-12965Vietnam.20.D.hanoiensis Bayarsaikhan &amp;Bae, UB-12969Vietnam.21.D.laocai Bayarsaikhan &amp; Heppner, sp. nov., paratype, UB-12956Vietnam. 22. D. robdevosi Bayarsaikhan &amp; Bae, UB-12971Vietnam." figureDoi="http://doi.org/10.5281/zenodo.13745459" httpUri="https://zenodo.org/record/13745459/files/figure.png" pageId="5" pageNumber="558">21</figureCitation>
)
</paragraph>
<paragraph id="F27EF622311A1132A0A8F896FBFEF8C9" blockId="5.[151,1067,1745,1845]" box="[151,1067,1818,1845]" pageId="5" pageNumber="558">
LSID:
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urn:lsid:zoobank.org:pub:
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</paragraph>
</subSubSection>
<subSubSection id="BADBA5A9311A1132A0A8F8EEFCE9F814" pageId="5" pageNumber="558" type="materials_examined">
<paragraph id="F27EF622311A1132A0A8F8EEFCE9F814" blockId="5.[151,1436,1889,2025]" pageId="5" pageNumber="558">
<emphasis id="C0B52A30311A1132A0A8F8EEFE46F887" bold="true" box="[151,403,1889,1916]" pageId="5" pageNumber="558">Type materials (n=4).</emphasis>
<materialsCitation id="42A9FC7F311A1132A1A7F8EEFAC7F85D" collectingDate="2011-05-17" collectingDateMax="2011-05-18" collectingDateMin="2011-05-17" collectionCode="MGCL, UB" collectorName="J. B. Heppner" country="Vietnam" location="Lao Cai Prov." pageId="5" pageNumber="558" specimenCount="1" specimenCount-male="1" stateProvince="Lao Cai" typeStatus="holotype">
<emphasis id="C0B52A30311A1132A1A7F8EEFDDBF880" bold="true" box="[408,526,1890,1916]" pageId="5" pageNumber="558">
<typeStatus id="2D7A4880311A1132A1A7F8EEFDDCF880" box="[408,521,1890,1916]" pageId="5" pageNumber="558" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="E4C73DAB311A1132A22CF8EEFDFFF880" box="[531,554,1890,1916]" count="1" pageId="5" pageNumber="558" type="male"></specimenCount>
,
<collectingCountry id="8AD6B6B2311A1132A20BF8EEFD43F880" box="[564,662,1890,1916]" name="Vietnam" pageId="5" pageNumber="558">Vietnam</collectingCountry>
,
<collectingRegion id="300538C0311A1132A2A0F8EEFCECF880" box="[671,825,1890,1916]" country="Vietnam" name="Lao Cai" pageId="5" pageNumber="558">Lao Cai Prov.</collectingRegion>
, Tram Ton Pass For. Sta.,
<date id="867FD0E2311A1132A46FF8EEFADBF880" box="[1104,1294,1890,1916]" pageId="5" pageNumber="558" value="2011-05-17" valueMax="2011-05-18" valueMin="2011-05-17">
<collectingDate id="963B290A311A1132A46FF8EEFADBF880" box="[1104,1294,1890,1916]" pageId="5" pageNumber="558" value="2011-05-17" valueMax="2011-05-18" valueMin="2011-05-17">
1718 May
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</date>
, 1950m,
<collectorName id="5F3493F4311A1132A54EF8EEFF2CF85C" pageId="5" pageNumber="558">J.B. Heppner</collectorName>
, adult no. FLMNH-MGCL1174125 (deposited in
<collectionCode id="94D06EE7311A1132A312F80AFC55F85C" box="[813,896,1926,1952]" pageId="5" pageNumber="558">MGCL</collectionCode>
) and genitalia slides no.
<collectionCode id="94D06EE7311A1132A4A6F80AFB14F85D" box="[1177,1217,1926,1953]" country="France" name="Laboratoire de Biostratigraphie" pageId="5" pageNumber="558">UB</collectionCode>
˗12955
</materialsCitation>
.
<materialsCitation id="42A9FC7F311A1132A522F80AFCECF814" collectingDate="2011-05-17" collectingDateMax="2011-05-18" collectingDateMin="2011-05-17" collectionCode="MGCL" collectorName="J. B. Heppner" country="Vietnam" location="Lao Cai Prov." pageId="5" pageNumber="558" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Lao Cai" typeStatus="paratype">
<emphasis id="C0B52A30311A1132A522F80AFA49F85C" bold="true" box="[1309,1436,1926,1952]" pageId="5" pageNumber="558">
<typeStatus id="2D7A4880311A1132A522F80AFA42F85C" box="[1309,1431,1926,1952]" pageId="5" pageNumber="558" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="E4C73DAB311A1132A0A8F826FF68F838" box="[151,189,1962,1988]" count="2" pageId="5" pageNumber="558" type="male">2♂</specimenCount>
,
<specimenCount id="E4C73DAB311A1132A0F3F826FF3AF838" box="[204,239,1962,1988]" count="1" pageId="5" pageNumber="558" type="female">1♀</specimenCount>
, same data as holotype, adults no. FLMNH-MGCL1174123-male, 1174124-male, 1174126-female and genitalia slides no. UB-12956-female (deposited in
<collectionCode id="94D06EE7311A1132A2E4F842FCE4F814" box="[731,817,1998,2024]" pageId="5" pageNumber="558">MGCL</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="A6BEA6AA31191131A0A8F8BFFE76F868" ID-DOI="http://doi.org/10.5281/zenodo.13745455" ID-Zenodo-Dep="13745455" httpUri="https://zenodo.org/record/13745455/files/figure.png" pageId="6" pageNumber="559" startId="6.[151,264,1843,1867]" targetBox="[161,1425,190,1809]" targetPageId="6" targetType="figure">
<paragraph id="F27EF62231191131A0A8F8BFFE76F868" blockId="6.[151,1436,1843,1940]" pageId="6" pageNumber="559">
<emphasis id="C0B52A3031191131A0A8F8BFFE81F8B7" bold="true" box="[151,340,1843,1867]" pageId="6" pageNumber="559">FIGURES 1617.</emphasis>
Male genitalia of the new
<taxonomicName id="35C18DA131191131A25AF8BFFCD4F8B7" box="[613,769,1843,1867]" pageId="6" pageNumber="559">
<emphasis id="C0B52A3031191131A25AF8BFFD64F8B7" box="[613,689,1843,1867]" italics="true" pageId="6" pageNumber="559">Diduga</emphasis>
species
</taxonomicName>
from Vietnam and compared species. 16.
<taxonomicName id="35C18DA131191131A492F8B8FEDEF893" authority="Bayarsaikhan &amp; Heppner" authorityName="Bayarsaikhan &amp; Heppner &amp; Kwon &amp; Bae" authorityYear="2024" class="Insecta" family="Erebidae" genus="Diduga" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="559" phylum="Arthropoda" rank="species" species="laocai" status="sp. nov.">
<emphasis id="C0B52A3031191131A492F8B8FADEF8B7" box="[1197,1291,1843,1867]" italics="true" pageId="6" pageNumber="559">D. laocai</emphasis>
Bayarsaikhan &amp; Heppner
</taxonomicName>
,
<emphasis id="C0B52A3031191131A128F8D4FEBDF88C" bold="true" box="[279,360,1880,1904]" pageId="6" pageNumber="559">
<taxonomicNameLabel id="DB86974B31191131A128F8D4FEBDF88C" box="[279,360,1880,1904]" pageId="6" pageNumber="559" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, UB-12955Vietnam. 17.
<taxonomicName id="35C18DA131191131A2ECF8D4FBEBF893" authority="Bayarsaikhan &amp; Bae" authorityName="Bayarsaikhan &amp; Bae" authorityYear="2021" box="[723,1086,1879,1904]" class="Insecta" family="Erebidae" genus="Diduga" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="559" phylum="Arthropoda" rank="species" species="mucronata">
<emphasis id="C0B52A3031191131A2ECF8D4FCB5F893" box="[723,864,1880,1903]" italics="true" pageId="6" pageNumber="559">D. mucronata</emphasis>
Bayarsaikhan &amp; Bae
</taxonomicName>
, holotype, INU- 10172Laos (after
<bibRefCitation id="96508BD331191131A0A8F8F7FE42F868" author="Bayarsaikhan, U. &amp; Cha, Y. B. &amp; Lee, T. G. &amp; Jang, C. M. &amp; Kwon, H. W. &amp; Bae, Y. S." box="[151,407,1915,1940]" pageId="6" pageNumber="559" pagination="131 - 142" refId="ref4484" refString="Bayarsaikhan, U., Cha, Y. B., Lee, T. G., Jang, C. M., Kwon, H. W. &amp; Bae, Y. S. (2021) Two new species and four new records of Diduga (Lepidoptera, Erebidae, Arctiinae) from Laos. Zootaxa, 4970 (1), 131 - 142. https: // doi. org / 10.11646 / zootaxa. 4970.1.5" type="journal article" year="2021">
Bayarsaikhan
<emphasis id="C0B52A3031191131A118F8F0FE8EF86F" box="[295,347,1915,1939]" italics="true" pageId="6" pageNumber="559">et al.</emphasis>
2021
</bibRefCitation>
).
</paragraph>
</caption>
<subSubSection id="BADBA5A931181130A0F8FF1AFC31FD79" pageId="7" pageNumber="560" type="diagnosis">
<paragraph id="F27EF62231181130A0F8FF1AFB79FDE5" blockId="7.[151,1437,150,1365]" pageId="7" pageNumber="560">
<emphasis id="C0B52A3031181130A0F8FF1AFE97FF4C" bold="true" box="[199,322,150,176]" pageId="7" pageNumber="560">Diagnosis.</emphasis>
By the wing pattern of this species, it is hardly distinguishable from many others of the genus
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<emphasis id="C0B52A3031181130A0A8FF37FF3EFF29" box="[151,235,187,213]" italics="true" pageId="7" pageNumber="560">Diduga</emphasis>
</taxonomicName>
, but the shape of the forewing of the new species is most similar to
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<emphasis id="C0B52A3031181130A3E8FF37FB7CFF28" box="[983,1193,187,213]" italics="true" pageId="7" pageNumber="560">Diduga mucronata</emphasis>
<bibRefCitation id="96508BD331181130A490FF37FF1AFF05" author="Bayarsaikhan, U. &amp; Cha, Y. B. &amp; Lee, T. G. &amp; Jang, C. M. &amp; Kwon, H. W. &amp; Bae, Y. S." pageId="7" pageNumber="560" pagination="131 - 142" refId="ref4484" refString="Bayarsaikhan, U., Cha, Y. B., Lee, T. G., Jang, C. M., Kwon, H. W. &amp; Bae, Y. S. (2021) Two new species and four new records of Diduga (Lepidoptera, Erebidae, Arctiinae) from Laos. Zootaxa, 4970 (1), 131 - 142. https: // doi. org / 10.11646 / zootaxa. 4970.1.5" type="journal article" year="2021">Bayarsaikhan &amp; Bae, 2021</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="96508BD331181130A0E1FF53FDE4FF05" author="Bayarsaikhan, U. &amp; Cha, Y. B. &amp; Lee, T. G. &amp; Jang, C. M. &amp; Kwon, H. W. &amp; Bae, Y. S." box="[222,561,223,249]" pageId="7" pageNumber="560" pagination="131 - 142" refId="ref4484" refString="Bayarsaikhan, U., Cha, Y. B., Lee, T. G., Jang, C. M., Kwon, H. W. &amp; Bae, Y. S. (2021) Two new species and four new records of Diduga (Lepidoptera, Erebidae, Arctiinae) from Laos. Zootaxa, 4970 (1), 131 - 142. https: // doi. org / 10.11646 / zootaxa. 4970.1.5" type="journal article" year="2021">
Bayarsaikhan
<emphasis id="C0B52A3031181130A140FF53FE62FF05" box="[383,439,223,249]" italics="true" pageId="7" pageNumber="560">et al.</emphasis>
2021: 134
</bibRefCitation>
, figs. 2a, 2b); both species are larger than other members of the genus
<taxonomicName id="35C18DA131181130A577FF53FA49FF05" authorityName="Moore" authorityYear="1887" box="[1352,1436,223,249]" class="Insecta" family="Erebidae" genus="Diduga" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="560" phylum="Arthropoda" rank="genus">
<emphasis id="C0B52A3031181130A577FF53FA49FF05" box="[1352,1436,223,249]" italics="true" pageId="7" pageNumber="560">Diduga</emphasis>
</taxonomicName>
(length of forewing longer than
<quantity id="35395BC731181130A1C8FE8FFDE3FEE1" box="[503,566,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="560" unit="mm" value="6.0">6 mm</quantity>
in the male) and forewing has a strongly angled apex and terminal area weakly covered by dark brown scales. In the male genitalia, both species have an apically bent uncus with broadened basal area, and a stout valva with a large weakly sclerotized, rounded spur in basal area of costal margin. However,
<taxonomicName id="35C18DA131181130A5BEFEC7FF08FE75" authorityName="Bayarsaikhan &amp; Heppner &amp; Kwon &amp; Bae" authorityYear="2024" class="Insecta" family="Erebidae" genus="Diduga" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="560" phylum="Arthropoda" rank="species" species="laocai" status="sp. nov.">
<emphasis id="C0B52A3031181130A5BEFEC7FF08FE75" italics="true" pageId="7" pageNumber="560">D. laocai</emphasis>
</taxonomicName>
<emphasis id="C0B52A3031181130A0D9FEE3FE97FE75" bold="true" box="[230,322,367,393]" pageId="7" pageNumber="560">
<taxonomicNameLabel id="DB86974B31181130A0D9FEE3FE97FE75" box="[230,322,367,393]" pageId="7" pageNumber="560" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
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<emphasis id="C0B52A3031181130A1E4FEE3FDA2FE74" box="[475,631,367,392]" italics="true" pageId="7" pageNumber="560">D. mucronata</emphasis>
</taxonomicName>
, in the male genitalia by having a larger, asymmetric apical spine and edged by a strongly sclerotized rounded spur in waved apical area of valva, and a strongly sclerotized arched juxta, and a large spine in the aedeagus vesica. In
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<emphasis id="C0B52A3031181130A2A4FE3BFCEDFE2C" box="[667,824,439,464]" italics="true" pageId="7" pageNumber="560">D. mucronata</emphasis>
</taxonomicName>
, valva has an asymmetric small apical spine in right valva (about 34 times shorter than new species), and edged membranous narrowly, and juxta broadly truncate, and aedeagus vesica with six differently sized spine-shaped cornuti in the male genitalia (
<figureCitation id="6AFAEAA731181130A46DFE73FB4BFDE5" box="[1106,1182,511,537]" captionStart="FIGURES 1617" captionStartId="6.[151,264,1843,1867]" captionTargetBox="[161,1425,190,1809]" captionTargetId="figure-24@6.[151,1436,181,1819]" captionTargetPageId="6" captionText="FIGURES 1617. Male genitalia of the new Diduga species from Vietnam and compared species. 16. D. laocai Bayarsaikhan &amp; Heppner, sp. nov., holotype, UB-12955Vietnam. 17. D. mucronata Bayarsaikhan &amp; Bae, holotype, INU- 10172Laos (after Bayarsaikhan et al. 2021)." figureDoi="http://doi.org/10.5281/zenodo.13745455" httpUri="https://zenodo.org/record/13745455/files/figure.png" pageId="7" pageNumber="560">Fig 17</figureCitation>
).
</paragraph>
<paragraph id="F27EF62231181130A0F8FDAFFC31FD79" blockId="7.[151,1437,150,1365]" pageId="7" pageNumber="560">
In female genitalia, ostium plate is strongly sclerotized and narrowly U-shaped, and ovoid corpus bursa has a large strongly sclerotized plate in cervix of bursa and whole of bursa is covered by short strong spines; and this distinguishes this species well from all the other known females of
<emphasis id="C0B52A3031181130A3B6FDE7FC31FD79" box="[905,996,619,645]" italics="true" pageId="7" pageNumber="560">
<taxonomicName id="35C18DA131181130A3B6FDE7FC34FD79" authorityName="Moore" authorityYear="1887" box="[905,993,619,645]" class="Insecta" family="Erebidae" genus="Diduga" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="560" phylum="Arthropoda" rank="genus">Diduga</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="BADBA5A931181130A0F8FD02FB02FB15" pageId="7" pageNumber="560" type="description">
<paragraph id="F27EF62231181130A0F8FD02FB02FB15" blockId="7.[151,1437,150,1365]" pageId="7" pageNumber="560">
<emphasis id="C0B52A3031181130A0F8FD02FE8CFD54" bold="true" box="[199,345,654,680]" pageId="7" pageNumber="560">Description.</emphasis>
<emphasis id="C0B52A3031181130A15BFD03FE74FD55" box="[356,417,655,681]" italics="true" pageId="7" pageNumber="560">Adult</emphasis>
(
<figureCitation id="6AFAEAA731181130A18CFD03FDB5FD55" box="[435,608,655,681]" captionStart="FIGURES 68" captionStartId="3.[151,264,1653,1677]" captionTargetBox="[151,1435,453,1630]" captionTargetId="figure-117@3.[151,1435,453,1630]" captionTargetPageId="3" captionText="FIGURES 68. Adults of the Diduga from Vietnam. 6a. D. laocai Bayarsaikhan &amp; Heppner, sp. nov., holotype, male. 6b. diito, paratype, female. 7a. D. robdevosi Bayarsaikhan &amp; Bae, male. 7b. ditto, female. 8. D. quinquicornuta Bayarsaikhan &amp; Bae, male." figureDoi="http://doi.org/10.5281/zenodo.13745449" httpUri="https://zenodo.org/record/13745449/files/figure.png" pageId="7" pageNumber="560">Figures 6a, 6b</figureCitation>
). Length of forewing
<quantity id="35395BC731181130A358FD03FC1DFD55" box="[871,968,655,681]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="7" pageNumber="560" unit="mm" value="8.5" valueMax="9.0" valueMin="8.0">89 mm</quantity>
in both sexes. Head and patagium pale yellowish. Proximal end of tegula pale yellowish and distal end dark brownish. Thorax dark brownish. Forewing ground color dark brownish, with pale yellowish, broad costal and terminal margin, it edged by golden yellowish line, except slender dark line in basal area of costal margin; with small, dark brown dots in costal border; cilia golden yellowish. Hindwing ground color pale brown; cilia pale brown. Abdomen brown, with deep yellow anal tuft and pale brown ventral side in male and female.
<emphasis id="C0B52A3031181130A2D4FCCFFC45FCA1" box="[747,912,835,861]" italics="true" pageId="7" pageNumber="560">Male genitalia</emphasis>
(
<figureCitation id="6AFAEAA731181130A3A1FCCFFBC5FCA1" box="[926,1040,835,861]" captionStart="FIGURES 1617" captionStartId="6.[151,264,1843,1867]" captionTargetBox="[161,1425,190,1809]" captionTargetId="figure-24@6.[151,1436,181,1819]" captionTargetPageId="6" captionText="FIGURES 1617. Male genitalia of the new Diduga species from Vietnam and compared species. 16. D. laocai Bayarsaikhan &amp; Heppner, sp. nov., holotype, UB-12955Vietnam. 17. D. mucronata Bayarsaikhan &amp; Bae, holotype, INU- 10172Laos (after Bayarsaikhan et al. 2021)." figureDoi="http://doi.org/10.5281/zenodo.13745455" httpUri="https://zenodo.org/record/13745455/files/figure.png" pageId="7" pageNumber="560">Figure 16</figureCitation>
). Uncus stout, tapering, apical area weakly bent, basal area broadened, with a bundle of spines ventrally, almost same length with tegumen. Anal tube membranous and scaphium weakly sclerotized. Tegumen triangular, weakly sclerotized. Valvae asymmetric, stout, weakly elongate and weakly sclerotized, with a strongly sclerotized, pointed, spine-like apical process, left apex of valva narrower than right; a large basal harpe weakly sclerotized; waved costal margin of valva strongly sclerotized, with a rounded, membranous apical process; apical saccular process strongly sclerotized and roundly edged. Juxta weakly sclerotized, arched. Vinculum short, weakly sclerotized; saccus broadly “U”˗shaped. Aedeagus stout, weakly sclerotized, and vesica covered with spinules and a large spine shaped cornuti and a sclerotized plate.
<emphasis id="C0B52A3031181130A0E4FBEFFE4CFB81" box="[219,409,1123,1149]" italics="true" pageId="7" pageNumber="560">Female genitalia</emphasis>
(
<figureCitation id="6AFAEAA731181130A198FBEFFDC2FB81" box="[423,535,1123,1149]" captionStart="FIGURES 1822" captionStartId="8.[151,264,1908,1932]" captionTargetBox="[161,1429,571,1875]" captionTargetId="figure-157@8.[151,1436,561,1885]" captionTargetPageId="8" captionText="FIGURES 1822. Female genitalia of the Diduga from Vietnam. 18. D. alternota Bucsek, UB-12981Vietnam. 19. D. bayartogtokhi Bayarsaikhan &amp;Bae, UB-12965Vietnam.20.D.hanoiensis Bayarsaikhan &amp;Bae, UB-12969Vietnam.21.D.laocai Bayarsaikhan &amp; Heppner, sp. nov., paratype, UB-12956Vietnam. 22. D. robdevosi Bayarsaikhan &amp; Bae, UB-12971Vietnam." figureDoi="http://doi.org/10.5281/zenodo.13745459" httpUri="https://zenodo.org/record/13745459/files/figure.png" pageId="7" pageNumber="560">Figure 21</figureCitation>
). Papillae anales weakly covered with setae; both apophyses slender and anterior apophysis shorter than posterior apophysis; lamella postvaginalis sclerotized; ostium plate strongly sclerotized, narrowly U-shaped. Ductus bursae membranous, short, broadly tubular; corpus bursae ovoid, covered by short, strong spines whole bursae, and a large strongly sclerotized plate covered in cervix of bursa area.
</paragraph>
</subSubSection>
<subSubSection id="BADBA5A931181130A0F8FB7EFD7EFAF1" box="[199,683,1266,1293]" pageId="7" pageNumber="560" type="distribution">
<paragraph id="F27EF62231181130A0F8FB7EFD7EFAF1" blockId="7.[151,1437,150,1365]" box="[199,683,1266,1293]" pageId="7" pageNumber="560">
<emphasis id="C0B52A3031181130A0F8FB7EFEB4FAF0" bold="true" box="[199,353,1266,1292]" pageId="7" pageNumber="560">Distribution.</emphasis>
<collectingCountry id="8AD6B6B231181130A158FB7FFE13FAF1" box="[359,454,1267,1293]" name="Vietnam" pageId="7" pageNumber="560">Vietnam</collectingCountry>
(
<collectingRegion id="300538C031181130A1EAFB7FFD75FAF1" box="[469,672,1267,1293]" country="Vietnam" name="Lao Cai" pageId="7" pageNumber="560">Lao Cai Province</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="BADBA5A931181130A0F8FA9BFD67FAA9" pageId="7" pageNumber="560" type="etymology">
<paragraph id="F27EF62231181130A0F8FA9BFD67FAA9" blockId="7.[151,1437,150,1365]" pageId="7" pageNumber="560">
<emphasis id="C0B52A3031181130A0F8FA9BFE9BFACD" bold="true" box="[199,334,1303,1329]" pageId="7" pageNumber="560">Etymology.</emphasis>
The species name refers to the
<typeStatus id="2D7A488031181130A2A0FA9BFD1AFACD" box="[671,719,1303,1329]" pageId="7" pageNumber="560">type</typeStatus>
locality,
<collectingRegion id="300538C031181130A30DFA9BFC21FACD" box="[818,1012,1303,1329]" country="Vietnam" name="Lao Cai" pageId="7" pageNumber="560">Lao Cai Province</collectingRegion>
, a mountainous region in northwestern
<collectingCountry id="8AD6B6B231181130A0A8FAB7FF23FAA9" box="[151,246,1339,1365]" name="Vietnam" pageId="7" pageNumber="560">Vietnam</collectingCountry>
bordering the
<collectingRegion id="300538C031181130A1A6FAB7FD8EFAA9" box="[409,603,1339,1365]" country="China" name="Yunnan" pageId="7" pageNumber="560">Yunnan Province</collectingRegion>
,
<collectingCountry id="8AD6B6B231181130A258FAB7FD7BFAA9" box="[615,686,1339,1365]" name="China" pageId="7" pageNumber="560">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,220 @@
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<mods:title id="151EB8A6C1E071CD665B9805AE0BF06B">Review of the genus Diduga (Lepidoptera, Erebidae, Arctiinae) of Vietnam, with a new species and two newly recorded species</mods:title>
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<taxonomicName id="35C18DA1311A1132A0A8FB0CFD37FB66" ID-CoL="BHR9T" authority="Bayarsaikhan &amp; Bae, 2019" authorityName="Bayarsaikhan &amp; Bae" authorityYear="2019" box="[151,738,1152,1179]" class="Insecta" family="Erebidae" genus="Diduga" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="558" phylum="Arthropoda" rank="species" species="nigridentata">
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<emphasis id="C0B52A30311A1132A0A8FB0CFE59FB67" bold="true" box="[151,396,1152,1179]" italics="true" pageId="5" pageNumber="558">Diduga nigridentata</emphasis>
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(
<figureCitation id="6AFAEAA7311A1132A0A0FB28FF3DFB43" box="[159,232,1188,1215]" captionStart="FIGURES 15" captionStartId="2.[151,264,1840,1864]" captionTargetBox="[212,1398,179,1762]" captionTargetId="figure-24@2.[151,1436,179,1814]" captionTargetPageId="2" captionText="FIGURES 15. Adults of the Diduga from Vietnam. 1a. D. alternota Bucsek, male. 1b. ditto, female. 2a. D. bayartogtokhi Bayarsaikhan &amp; Bae, male. 2b. ditto, female. 3a. D. hanoiensis Bayarsaikhan &amp; Bae, male. 3b. ditto, female. 4. D. hastata Bayarsaikhan &amp; Bae, male. 5. D. nigridentata Bayarsaikhan &amp; Bae, male." figureDoi="http://doi.org/10.5281/zenodo.13745445" httpUri="https://zenodo.org/record/13745445/files/figure.png" pageId="5" pageNumber="558">Figs 5</figureCitation>
,
<figureCitation id="6AFAEAA7311A1132A0CBFB29FEC5FB43" box="[244,272,1189,1215]" captionStart="FIGURES 915" captionStartId="4.[151,264,1901,1925]" captionTargetBox="[160,1427,319,1868]" captionTargetId="figure-69@4.[151,1436,309,1878]" captionTargetPageId="4" captionText="FIGURES 915. Male genitalia of the Diduga from Vietnam. 9. D. alternota Bucsek, UB-12951Vietnam. 10. D. bayartogtokhi Bayarsaikhan&amp; Bae, UB-12958Vietnam.11.D.hanoiensis Bayarsaikhan&amp; Bae, UB-12976Vietnam.12.D.hastataBayarsaikhan &amp; Bae, UB-12966Vietnam. 13. D. nigridentata Bayarsaikhan &amp; Bae, UB-12957Vietnam. 14. D. quinquicornuta Bayarsaikhan &amp; Bae, UB-12962Vietnam. 15. D. robdevosi Bayarsaikhan &amp; Bae, UB-12970Vietnam." figureDoi="http://doi.org/10.5281/zenodo.13745453" httpUri="https://zenodo.org/record/13745453/files/figure.png" pageId="5" pageNumber="558">13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="BADBA5A9311A1132A0A8FB65FADAFAFD" box="[151,1295,1257,1281]" pageId="5" pageNumber="558" type="reference_group">
<paragraph id="F27EF622311A1132A0A8FB65FADAFAFD" blockId="5.[151,1295,1257,1281]" box="[151,1295,1257,1281]" pageId="5" pageNumber="558">
<taxonomicName id="35C18DA1311A1132A0A8FB65FD55FAFD" ID-CoL="BHR9T" authority="Bayarsaikhan &amp; Bae, 2019" authorityName="Bayarsaikhan &amp; Bae" authorityYear="2019" box="[151,640,1257,1281]" class="Insecta" family="Erebidae" genus="Diduga" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="558" phylum="Arthropoda" rank="species" species="nigridentata">
<emphasis id="C0B52A30311A1132A0A8FB65FEB2FAFD" box="[151,359,1257,1281]" italics="true" pageId="5" pageNumber="558">Diduga nigridentata</emphasis>
<bibRefCitation id="96508BD3311A1132A152FB65FD55FAFD" author="Bayarsaikhan, U. &amp; Bae, Y. S." box="[365,640,1257,1281]" pageId="5" pageNumber="558" pagination="365 - 376" refId="ref4259" refString="Bayarsaikhan, U. &amp; Bae, Y. S. (2019) Four new and one newly recorded species of Diduga Moore, [1887] (Lepidoptera, Erebidae, Arctiinae) from Vietnam, with redescription of the little known species Diduga haematomiformis van Eecke, 1920. Zootaxa, 4624 (3), 365 - 376. https: // doi. org / 10.11646 / zootaxa. 4624.3.5" type="journal article" year="2019">Bayarsaikhan &amp; Bae, 2019</bibRefCitation>
</taxonomicName>
,
<emphasis id="C0B52A30311A1132A2B4FB66FD0BFAFD" box="[651,734,1258,1281]" italics="true" pageId="5" pageNumber="558">Zootaxa</emphasis>
4624 (3): 373, figs. 7, 13. TL:
<collectingCountry id="8AD6B6B2311A1132A427FB65FBBBFAFD" box="[1048,1134,1257,1281]" name="Vietnam" pageId="5" pageNumber="558">Vietnam</collectingCountry>
(
<collectingCountry id="8AD6B6B2311A1132A443FB65FB14FAFD" box="[1148,1217,1257,1281]" name="Turkey" pageId="5" pageNumber="558">Hatay</collectingCountry>
, BaVi).
</paragraph>
</subSubSection>
<subSubSection id="BADBA5A9311A1132A0A8FAA3FB4CF9DD" pageId="5" pageNumber="558" type="distribution">
<paragraph id="F27EF622311A1132A0A8FAA3FE48FAB5" blockId="5.[151,1437,1327,1677]" box="[151,413,1327,1353]" pageId="5" pageNumber="558">
<emphasis id="C0B52A30311A1132A0A8FAA3FEE4FAB5" bold="true" box="[151,305,1327,1353]" pageId="5" pageNumber="558">Distribution.</emphasis>
<collectingCountry id="8AD6B6B2311A1132A108FAA3FE4FFAB5" box="[311,410,1327,1353]" name="Vietnam" pageId="5" pageNumber="558">Vietnam</collectingCountry>
.
</paragraph>
<paragraph id="F27EF622311A1132A0F8FADFFB4CF9DD" blockId="5.[151,1437,1327,1677]" pageId="5" pageNumber="558">
<emphasis id="C0B52A30311A1132A0F8FADFFDE3FA91" bold="true" box="[199,566,1363,1389]" pageId="5" pageNumber="558">Additional material examined.</emphasis>
<materialsCitation id="42A9FC7F311A1132A201FADFFC0FFA6D" collectingDate="2009-05-04" collectingDateMax="2009-05-07" collectingDateMin="2009-05-04" collectionCode="UB" collectorName="Cuc Phuong N. P. &amp; J. B. Heppner" country="Vietnam" location="Mac Lk." pageId="5" pageNumber="558" specimenCount="1" specimenCount-male="1" stateProvince="Ninh Binh">
<specimenCount id="E4C73DAB311A1132A201FADFFDB7FA91" box="[574,610,1363,1389]" count="1" pageId="5" pageNumber="558" type="male">1♂</specimenCount>
,
<collectingCountry id="8AD6B6B2311A1132A250FADFFD04FA91" box="[623,721,1363,1389]" name="Vietnam" pageId="5" pageNumber="558">Vietnam</collectingCountry>
,
<collectingRegion id="300538C0311A1132A2E2FADFFC83FA91" box="[733,854,1363,1389]" country="Vietnam" name="Ninh Binh" pageId="5" pageNumber="558">Ninh Binh</collectingRegion>
,
<collectorName id="5F3493F4311A1132A35DFADFFBF3FA92" box="[866,1062,1363,1390]" pageId="5" pageNumber="558">Cuc Phuong N.P.</collectorName>
,
<location id="F71EA0F9311A1132A40DFAD8FB40FA91" LSID="urn:lsid:plazi:treatment:7A684734311A1132A0A8FB0CFEFBF971:F71EA0F9311A1132A40DFAD8FB40FA91" box="[1074,1173,1363,1390]" country="Vietnam" name="Mac Lk." pageId="5" pageNumber="558" stateProvince="Ninh Binh">Mac Lk.</location>
,
<date id="867FD0E2311A1132A49EFADFFA9CFA91" box="[1185,1353,1363,1390]" pageId="5" pageNumber="558" value="2009-05-04" valueMax="2009-05-07" valueMin="2009-05-04">
<collectingDate id="963B290A311A1132A49EFADFFA9CFA91" box="[1185,1353,1363,1390]" pageId="5" pageNumber="558" value="2009-05-04" valueMax="2009-05-07" valueMin="2009-05-04">
47 May
<quantity id="35395BC7311A1132A531FADFFA9CFA91" box="[1294,1353,1363,1389]" metricMagnitude="3" metricUnit="m" metricValue="2.009155" pageId="5" pageNumber="558" unit="m" value="2009.155">2009</quantity>
</collectingDate>
</date>
, 155m,
<collectorName id="5F3493F4311A1132A0A8FAF4FEFFFA6E" box="[151,298,1400,1426]" pageId="5" pageNumber="558">J.B. Heppner</collectorName>
, adult no. FLMNH-MGCL1174188 and genitalia slide no.
<collectionCode id="94D06EE7311A1132A38DFAFBFC0FFA6D" box="[946,986,1399,1425]" country="France" name="Laboratoire de Biostratigraphie" pageId="5" pageNumber="558">UB</collectionCode>
</materialsCitation>
˗12973;
<materialsCitation id="42A9FC7F311A1132A40AFAFBFA93FA49" collectingDate="2010-07-19" collectingDateMax="2010-07-23" collectingDateMin="2010-07-19" collectionCode="UB" collectorName="Prov. &amp; Ba Vi Natl. Pk. &amp; Heppner" country="Vietnam" elevation="455" location="Vietnam" pageId="5" pageNumber="558" specimenCount="1" specimenCount-male="1">
<specimenCount id="E4C73DAB311A1132A40AFAFBFB8FFA6D" box="[1077,1114,1399,1425]" count="1" pageId="5" pageNumber="558" type="male">1♂</specimenCount>
,
<collectingCountry id="8AD6B6B2311A1132A45BFAFBFB13FA6D" box="[1124,1222,1399,1425]" name="Vietnam" pageId="5" pageNumber="558">Vietnam</collectingCountry>
, Hatay
<collectorName id="5F3493F4311A1132A527FAF4FA80FA6E" box="[1304,1365,1400,1426]" pageId="5" pageNumber="558">Prov.</collectorName>
,
<collectorName id="5F3493F4311A1132A561FAF4FF2EFA49" pageId="5" pageNumber="558">Ba Vi Natl. Pk.</collectorName>
,
<quantity id="35395BC7311A1132A13BFA17FE92FA4A" box="[260,327,1435,1462]" metricMagnitude="2" metricUnit="m" metricValue="4.55" pageId="5" pageNumber="558" unit="m" value="455.0">
<elevation id="79EC1111311A1132A13BFA17FE92FA4A" box="[260,327,1435,1462]" metricMagnitude="2" metricUnit="m" metricValue="4.55" pageId="5" pageNumber="558" unit="m" value="455.0">455m</elevation>
</quantity>
,
<date id="867FD0E2311A1132A16FFA17FE29FA49" box="[336,508,1435,1461]" pageId="5" pageNumber="558" value="2010-07-19" valueMax="2010-07-23" valueMin="2010-07-19">
<collectingDate id="963B290A311A1132A16FFA17FE29FA49" box="[336,508,1435,1461]" pageId="5" pageNumber="558" value="2010-07-19" valueMax="2010-07-23" valueMin="2010-07-19">1923 Jul 2010</collectingDate>
</date>
, J.B.
<collectorName id="5F3493F4311A1132A209FA10FD4DFA4A" box="[566,664,1436,1462]" pageId="5" pageNumber="558">Heppner</collectorName>
, adult no. FLMNH-MGCL1174194 and genitalia slide no.
<collectionCode id="94D06EE7311A1132A521FA17FA93FA49" box="[1310,1350,1435,1461]" country="France" name="Laboratoire de Biostratigraphie" pageId="5" pageNumber="558">UB</collectionCode>
</materialsCitation>
˗12978;
<materialsCitation id="42A9FC7F311A1132A0A8FA33FE45FA01" collectingDate="2014-10-10" collectingDateMax="2014-10-12" collectingDateMin="2014-10-10" collectionCode="UB" collectorName="J. B. Heppner" country="Vietnam" location="Tam Dao" pageId="5" pageNumber="558" specimenCount="1" specimenCount-male="1" stateProvince="Vinh Phuc">
<specimenCount id="E4C73DAB311A1132A0A8FA33FF69FA25" box="[151,188,1471,1497]" count="1" pageId="5" pageNumber="558" type="male">1♂</specimenCount>
,
<collectingCountry id="8AD6B6B2311A1132A0F7FA33FEFFFA25" box="[200,298,1471,1497]" name="Vietnam" pageId="5" pageNumber="558">Vietnam</collectingCountry>
,
<collectingRegion id="300538C0311A1132A10AFA33FE7BFA25" box="[309,430,1471,1497]" country="Vietnam" name="Vinh Phuc" pageId="5" pageNumber="558">Vinh Phuc</collectingRegion>
,
<location id="F71EA0F9311A1132A185FA4CFDF1FA26" LSID="urn:lsid:plazi:treatment:7A684734311A1132A0A8FB0CFEFBF971:F71EA0F9311A1132A185FA4CFDF1FA26" box="[442,548,1472,1498]" country="Vietnam" name="Tam Dao" pageId="5" pageNumber="558" stateProvince="Vinh Phuc">Tam Dao</location>
,
<date id="867FD0E2311A1132A20FFA33FD3CFA25" box="[560,745,1471,1497]" pageId="5" pageNumber="558" value="2014-10-10" valueMax="2014-10-12" valueMin="2014-10-10">
<collectingDate id="963B290A311A1132A20FFA33FD3CFA25" box="[560,745,1471,1497]" pageId="5" pageNumber="558" value="2014-10-10" valueMax="2014-10-12" valueMin="2014-10-10">
1012 Oct
<quantity id="35395BC7311A1132A291FA33FD3CFA25" box="[686,745,1471,1497]" metricMagnitude="3" metricUnit="m" metricValue="2.01493" pageId="5" pageNumber="558" unit="m" value="2014.93">2014</quantity>
</collectingDate>
</date>
, 930m,
<collectorName id="5F3493F4311A1132A37CFA4CFC0DFA26" box="[835,984,1472,1498]" pageId="5" pageNumber="558">J.B. Heppner</collectorName>
, adult no. FLMNH-MGCL1174135 and genitalia slide no.
<collectionCode id="94D06EE7311A1132A157FA6FFE45FA01" box="[360,400,1507,1533]" country="France" name="Laboratoire de Biostratigraphie" pageId="5" pageNumber="558">UB</collectionCode>
</materialsCitation>
˗12964;
<materialsCitation id="42A9FC7F311A1132A1D1FA6FFC9DF9DE" collectingDate="2017-07-14" collectingDateMax="2017-07-18" collectingDateMin="2017-07-14" collectionCode="UB" collectorName="J. B. Heppner" country="Vietnam" location="Truong Xuan Lodge" municipality="Hagiang" pageId="5" pageNumber="558" specimenCount="1" specimenCount-female="1">
<specimenCount id="E4C73DAB311A1132A1D1FA6FFDC4FA01" box="[494,529,1507,1533]" count="1" pageId="5" pageNumber="558" type="female">1♀</specimenCount>
,
<collectingCountry id="8AD6B6B2311A1132A21FFA6FFD57FA01" box="[544,642,1507,1533]" name="Vietnam" pageId="5" pageNumber="558">Vietnam</collectingCountry>
,
<collectingMunicipality id="121A6C58311A1132A2B1FA6FFD38FA01" box="[654,749,1507,1533]" pageId="5" pageNumber="558">Hagiang</collectingMunicipality>
(
<location id="F71EA0F9311A1132A2C3FA6FFC33FA01" LSID="urn:lsid:plazi:treatment:7A684734311A1132A0A8FB0CFEFBF971:F71EA0F9311A1132A2C3FA6FFC33FA01" box="[764,998,1507,1534]" country="Vietnam" municipality="Hagiang" name="Truong Xuan Lodge" pageId="5" pageNumber="558">Truong Xuan Lodge</location>
)
<date id="867FD0E2311A1132A3C9FA6FFB7DFA01" box="[1014,1192,1507,1533]" pageId="5" pageNumber="558" value="2017-07-14" valueMax="2017-07-18" valueMin="2017-07-14">
<collectingDate id="963B290A311A1132A3C9FA6FFB7DFA01" box="[1014,1192,1507,1533]" pageId="5" pageNumber="558" value="2017-07-14" valueMax="2017-07-18" valueMin="2017-07-14">
1418 Jul
<quantity id="35395BC7311A1132A452FA6FFB7DFA01" box="[1133,1192,1507,1533]" metricMagnitude="3" metricUnit="m" metricValue="2.0171099999999997" pageId="5" pageNumber="558" unit="m" value="2017.11">2017</quantity>
</collectingDate>
</date>
, 110m,
<collectorName id="5F3493F4311A1132A53DFA68FA4DFA02" box="[1282,1432,1508,1534]" pageId="5" pageNumber="558">J.B. Heppner</collectorName>
, adult no. FLMNH-MGCL1174127 and genitalia slide no.
<collectionCode id="94D06EE7311A1132A31EF98BFC9DF9DE" box="[801,840,1543,1570]" country="France" name="Laboratoire de Biostratigraphie" pageId="5" pageNumber="558">UB</collectionCode>
</materialsCitation>
˗12957 (deposited in MGCL).
</paragraph>
</subSubSection>
<subSubSection id="BADBA5A9311A1132A0F8F9A7FEFBF971" pageId="5" pageNumber="558" type="discussion">
<paragraph id="F27EF622311A1132A0F8F9A7FEFBF971" blockId="5.[151,1437,1327,1677]" pageId="5" pageNumber="558">
<emphasis id="C0B52A30311A1132A0F8F9A7FEEEF9B9" bold="true" box="[199,315,1579,1605]" pageId="5" pageNumber="558">Remarks.</emphasis>
This species was reported from
<collectingCountry id="8AD6B6B2311A1132A2D8F9A7FC85F9B9" box="[743,848,1579,1605]" name="Vietnam" pageId="5" pageNumber="558">Vietnam</collectingCountry>
(
<collectingCountry id="8AD6B6B2311A1132A358F9A7FC61F9B9" box="[871,948,1579,1605]" name="Turkey" pageId="5" pageNumber="558">Hatay</collectingCountry>
(Haoi) and
<collectingRegion id="300538C0311A1132A460F9A7FB3DF9B9" box="[1119,1256,1579,1605]" country="Vietnam" name="Vinh Phuc" pageId="5" pageNumber="558">Vinh Phuc</collectingRegion>
Provinces) by
<bibRefCitation id="96508BD3311A1132A0A8F9C3FDC6F995" author="Bayarsaikhan, U. &amp; Bae, Y. S." box="[151,531,1615,1642]" pageId="5" pageNumber="558" pagination="365 - 376" refId="ref4259" refString="Bayarsaikhan, U. &amp; Bae, Y. S. (2019) Four new and one newly recorded species of Diduga Moore, [1887] (Lepidoptera, Erebidae, Arctiinae) from Vietnam, with redescription of the little known species Diduga haematomiformis van Eecke, 1920. Zootaxa, 4624 (3), 365 - 376. https: // doi. org / 10.11646 / zootaxa. 4624.3.5" type="journal article" year="2019">Bayarsaikhan and Bae (2019)</bibRefCitation>
. In this study, we report it for Hagiang and
<collectingRegion id="300538C0311A1132A43DF9C3FBADF995" box="[1026,1144,1615,1641]" country="Vietnam" name="Ninh Binh" pageId="5" pageNumber="558">Ninh Binh</collectingRegion>
Provinces of
<collectingCountry id="8AD6B6B2311A1132A52AF9C3FAA1F995" box="[1301,1396,1615,1641]" name="Vietnam" pageId="5" pageNumber="558">Vietnam</collectingCountry>
for the first time.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,178 @@
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<mods:title id="EDE82C10E0F872E25646DC2F91476698">Review of the genus Diduga (Lepidoptera, Erebidae, Arctiinae) of Vietnam, with a new species and two newly recorded species</mods:title>
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<mods:namePart id="936C5B603E844EFFA52833452530A0F6">Bayarsaikhan, Ulziijargal</mods:namePart>
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<mods:affiliation id="898FF472905151493F6E65F793B881FF">Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. &amp; Bio ˗ Resource and Environmental Center, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. &amp; Convergence Research Center for Insect Vectors, Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, Songdo ˗ dong, Incheon, 22012, Republic of Korea.</mods:affiliation>
<mods:nameIdentifier id="556CA6F48ECD3B34EC12F55CEB99C065" type="email">baeys@inu.ac.kr</mods:nameIdentifier>
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<heading id="A936414E311A1132A0A8FD2CFD73FD46" bold="true" box="[151,678,672,699]" fontSize="11" level="1" pageId="5" pageNumber="558" reason="1">
<taxonomicName id="35C18DA1311A1132A0A8FD2CFD73FD46" ID-CoL="BHR9P" authority="Bayarsaikhan &amp; Bae, 2021" authorityName="Bayarsaikhan &amp; Bae" authorityYear="2021" box="[151,678,672,699]" class="Insecta" family="Erebidae" genus="Diduga" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="558" phylum="Arthropoda" rank="species" species="hastata">
<emphasis id="C0B52A30311A1132A0A8FD2CFD73FD46" bold="true" box="[151,678,672,699]" pageId="5" pageNumber="558">
<emphasis id="C0B52A30311A1132A0A8FD2CFE85FD47" bold="true" box="[151,336,672,699]" italics="true" pageId="5" pageNumber="558">Diduga hastata</emphasis>
<bibRefCitation id="96508BD3311A1132A167FD2CFD73FD46" author="Bayarsaikhan, U. &amp; Cha, Y. B. &amp; Lee, T. G. &amp; Jang, C. M. &amp; Kwon, H. W. &amp; Bae, Y. S." box="[344,678,672,699]" pageId="5" pageNumber="558" pagination="131 - 142" refId="ref4484" refString="Bayarsaikhan, U., Cha, Y. B., Lee, T. G., Jang, C. M., Kwon, H. W. &amp; Bae, Y. S. (2021) Two new species and four new records of Diduga (Lepidoptera, Erebidae, Arctiinae) from Laos. Zootaxa, 4970 (1), 131 - 142. https: // doi. org / 10.11646 / zootaxa. 4970.1.5" type="journal article" year="2021">Bayarsaikhan &amp; Bae, 2021</bibRefCitation>
</emphasis>
</taxonomicName>
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</paragraph>
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<paragraph id="F27EF622311A1132A0A8FD48FECFFD23" blockId="5.[151,678,672,735]" box="[151,282,708,735]" pageId="5" pageNumber="558">
(
<figureCitation id="6AFAEAA7311A1132A0A0FD48FF3DFD23" box="[159,232,708,735]" captionStart="FIGURES 15" captionStartId="2.[151,264,1840,1864]" captionTargetBox="[212,1398,179,1762]" captionTargetId="figure-24@2.[151,1436,179,1814]" captionTargetPageId="2" captionText="FIGURES 15. Adults of the Diduga from Vietnam. 1a. D. alternota Bucsek, male. 1b. ditto, female. 2a. D. bayartogtokhi Bayarsaikhan &amp; Bae, male. 2b. ditto, female. 3a. D. hanoiensis Bayarsaikhan &amp; Bae, male. 3b. ditto, female. 4. D. hastata Bayarsaikhan &amp; Bae, male. 5. D. nigridentata Bayarsaikhan &amp; Bae, male." figureDoi="http://doi.org/10.5281/zenodo.13745445" httpUri="https://zenodo.org/record/13745445/files/figure.png" pageId="5" pageNumber="558">Figs 4</figureCitation>
,
<figureCitation id="6AFAEAA7311A1132A0CBFD49FEC4FD23" box="[244,273,709,735]" captionStart="FIGURES 915" captionStartId="4.[151,264,1901,1925]" captionTargetBox="[160,1427,319,1868]" captionTargetId="figure-69@4.[151,1436,309,1878]" captionTargetPageId="4" captionText="FIGURES 915. Male genitalia of the Diduga from Vietnam. 9. D. alternota Bucsek, UB-12951Vietnam. 10. D. bayartogtokhi Bayarsaikhan&amp; Bae, UB-12958Vietnam.11.D.hanoiensis Bayarsaikhan&amp; Bae, UB-12976Vietnam.12.D.hastataBayarsaikhan &amp; Bae, UB-12966Vietnam. 13. D. nigridentata Bayarsaikhan &amp; Bae, UB-12957Vietnam. 14. D. quinquicornuta Bayarsaikhan &amp; Bae, UB-12962Vietnam. 15. D. robdevosi Bayarsaikhan &amp; Bae, UB-12970Vietnam." figureDoi="http://doi.org/10.5281/zenodo.13745453" httpUri="https://zenodo.org/record/13745453/files/figure.png" pageId="5" pageNumber="558">12</figureCitation>
)
</paragraph>
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<paragraph id="F27EF622311A1132A0A8FC85FEB6FCBC" blockId="5.[151,1436,777,832]" pageId="5" pageNumber="558">
<taxonomicName id="35C18DA1311A1132A0A8FC85FD98FCDD" ID-CoL="BHR9P" authority="Bayarsaikhan &amp; Bae, 2021" authorityName="Bayarsaikhan &amp; Bae" authorityYear="2021" box="[151,589,777,801]" class="Insecta" family="Erebidae" genus="Diduga" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="558" phylum="Arthropoda" rank="species" species="hastata">
<emphasis id="C0B52A30311A1132A0A8FC85FEE1FCDD" box="[151,308,777,801]" italics="true" pageId="5" pageNumber="558">Diduga hastata</emphasis>
<bibRefCitation id="96508BD3311A1132A105FC85FD98FCDD" author="Bayarsaikhan, U. &amp; Cha, Y. B. &amp; Lee, T. G. &amp; Jang, C. M. &amp; Kwon, H. W. &amp; Bae, Y. S." box="[314,589,777,801]" pageId="5" pageNumber="558" pagination="131 - 142" refId="ref4484" refString="Bayarsaikhan, U., Cha, Y. B., Lee, T. G., Jang, C. M., Kwon, H. W. &amp; Bae, Y. S. (2021) Two new species and four new records of Diduga (Lepidoptera, Erebidae, Arctiinae) from Laos. Zootaxa, 4970 (1), 131 - 142. https: // doi. org / 10.11646 / zootaxa. 4970.1.5" type="journal article" year="2021">Bayarsaikhan &amp; Bae, 2021</bibRefCitation>
</taxonomicName>
,
<emphasis id="C0B52A30311A1132A268FC86FD7FFCDD" box="[599,682,778,801]" italics="true" pageId="5" pageNumber="558">Zootaxa</emphasis>
4970 (1): 132, figs. 1, 11, 19. TL:
<collectingCountry id="8AD6B6B2311A1132A437FC85FBECFCDC" box="[1032,1081,777,800]" name="Laos" pageId="5" pageNumber="558">Laos</collectingCountry>
(
<collectingRegion id="300538C0311A1132A478FC85FB1EFCDD" box="[1095,1227,777,801]" country="Laos" name="Bolikhamxai" pageId="5" pageNumber="558">Bolikhamsai</collectingRegion>
, Phou Khao Khouay National Park).
</paragraph>
</subSubSection>
<subSubSection id="BADBA5A9311A1132A0A8FCE1FDADFC74" box="[151,632,877,904]" pageId="5" pageNumber="558" type="distribution">
<paragraph id="F27EF622311A1132A0A8FCE1FDADFC74" blockId="5.[151,1437,877,1084]" box="[151,632,877,904]" pageId="5" pageNumber="558">
<emphasis id="C0B52A30311A1132A0A8FCE1FEE4FC7B" bold="true" box="[151,305,877,903]" pageId="5" pageNumber="558">Distribution.</emphasis>
<collectingCountry id="8AD6B6B2311A1132A108FCE2FE43FC74" box="[311,406,878,904]" name="Vietnam" pageId="5" pageNumber="558">Vietnam</collectingCountry>
(new record),
<collectingCountry id="8AD6B6B2311A1132A205FCE2FDA1FC74" box="[570,628,878,904]" name="Laos" pageId="5" pageNumber="558">Laos</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="BADBA5A9311A1132A0F8FC1DFB3BFBE4" pageId="5" pageNumber="558" type="materials_examined">
<paragraph id="F27EF622311A1132A0F8FC1DFB3BFBE4" blockId="5.[151,1437,877,1084]" pageId="5" pageNumber="558">
<emphasis id="C0B52A30311A1132A0F8FC1DFE65FC57" bold="true" box="[199,432,913,939]" pageId="5" pageNumber="558">Material examined.</emphasis>
<materialsCitation id="42A9FC7F311A1132A189FC1EFCDAFC2C" collectingDate="2010-08-01" collectingDateMax="2010-08-06" collectingDateMin="2010-08-01" collectionCode="NW, UB" collectorName="J. B. Heppner" country="Vietnam" elevation="970" location="Tam Dao" pageId="5" pageNumber="558" specimenCount="1" specimenCount-male="1" stateProvince="Vinh Phuc">
<specimenCount id="E4C73DAB311A1132A189FC1EFE0FFC50" box="[438,474,914,940]" count="1" pageId="5" pageNumber="558" type="male">1♂</specimenCount>
,
<collectingCountry id="8AD6B6B2311A1132A1D9FC1EFD9DFC50" box="[486,584,914,940]" name="Vietnam" pageId="5" pageNumber="558">Vietnam</collectingCountry>
,
<collectingRegion id="300538C0311A1132A26EFC1EFCDFFC50" box="[593,778,914,940]" country="Vietnam" name="Vinh Phuc" pageId="5" pageNumber="558">Vinh Phuc Prov.</collectingRegion>
,
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(
<collectionCode id="94D06EE7311A1132A3B7FC1EFC69FC50" box="[904,956,914,940]" pageId="5" pageNumber="558">NW</collectionCode>
),
<quantity id="35395BC7311A1132A3F1FC1EFBC4FC50" box="[974,1041,914,940]" metricMagnitude="2" metricUnit="m" metricValue="9.7" pageId="5" pageNumber="558" unit="m" value="970.0">
<elevation id="79EC1111311A1132A3F1FC1EFBC4FC50" box="[974,1041,914,940]" metricMagnitude="2" metricUnit="m" metricValue="9.7" pageId="5" pageNumber="558" unit="m" value="970.0">970m</elevation>
</quantity>
,
<date id="867FD0E2311A1132A424FC1EFB6EFC50" box="[1051,1211,914,940]" pageId="5" pageNumber="558" value="2010-08-01" valueMax="2010-08-06" valueMin="2010-08-01">
<collectingDate id="963B290A311A1132A424FC1EFB6EFC50" box="[1051,1211,914,940]" pageId="5" pageNumber="558" value="2010-08-01" valueMax="2010-08-06" valueMin="2010-08-01">16 Aug 2010</collectingDate>
</date>
,
<collectorName id="5F3493F4311A1132A4F9FC1EFA8FFC50" box="[1222,1370,914,940]" pageId="5" pageNumber="558">J.B. Heppner</collectorName>
, adult no. FLMNH-MGCL1174189 and genitalia slide no.
<collectionCode id="94D06EE7311A1132A2D8FC3AFCDAFC2C" box="[743,783,950,976]" country="France" name="Laboratoire de Biostratigraphie" pageId="5" pageNumber="558">UB</collectionCode>
</materialsCitation>
˗12988;
<materialsCitation id="42A9FC7F311A1132A352FC3AFAD3FC08" collectingDate="2016-08-18" collectingDateMax="2016-08-19" collectingDateMin="2016-08-18" collectionCode="UB" collectorName="J. B. Heppner" country="Vietnam" location="Truong Xuan Lodge" municipality="Hagiang" pageId="5" pageNumber="558" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="E4C73DAB311A1132A352FC3AFC44FC2C" box="[877,913,950,976]" count="1" pageId="5" pageNumber="558" type="male">1♂</specimenCount>
,
<specimenCount id="E4C73DAB311A1132A3A0FC3AFC17FC2C" box="[927,962,950,976]" count="1" pageId="5" pageNumber="558" type="female">1♀</specimenCount>
,
<collectingCountry id="8AD6B6B2311A1132A3EFFC3AFBE7FC2C" box="[976,1074,950,976]" name="Vietnam" pageId="5" pageNumber="558">Vietnam</collectingCountry>
,
<collectingMunicipality id="121A6C58311A1132A401FC3AFB48FC2C" box="[1086,1181,950,976]" pageId="5" pageNumber="558">Hagiang</collectingMunicipality>
(
<location id="F71EA0F9311A1132A493FC3AFA41FC2C" LSID="urn:lsid:plazi:treatment:7A684734311A1132A0A8FD2CFBA1FBC0:F71EA0F9311A1132A493FC3AFA41FC2C" box="[1196,1428,950,976]" country="Vietnam" municipality="Hagiang" name="Truong Xuan Lodge" pageId="5" pageNumber="558">Truong Xuan Lodge</location>
)
<date id="867FD0E2311A1132A0A8FC56FE83FC08" box="[151,342,986,1012]" pageId="5" pageNumber="558" value="2016-08-18" valueMax="2016-08-19" valueMin="2016-08-18">
<collectingDate id="963B290A311A1132A0A8FC56FE83FC08" box="[151,342,986,1012]" pageId="5" pageNumber="558" value="2016-08-18" valueMax="2016-08-19" valueMin="2016-08-18">
1819 Aug
<quantity id="35395BC7311A1132A124FC56FE83FC08" box="[283,342,986,1012]" metricMagnitude="3" metricUnit="m" metricValue="2.01611" pageId="5" pageNumber="558" unit="m" value="2016.11">2016</quantity>
</collectingDate>
</date>
, 110m,
<collectorName id="5F3493F4311A1132A190FC56FD90FC08" box="[431,581,986,1012]" pageId="5" pageNumber="558">J.B. Heppner</collectorName>
, adult no. FLMNH-MGCL1174138 and genitalia slide no.
<collectionCode id="94D06EE7311A1132A4E0FC56FAD3FC08" box="[1247,1286,986,1012]" country="France" name="Laboratoire de Biostratigraphie" pageId="5" pageNumber="558">UB</collectionCode>
</materialsCitation>
˗12966-male, adult no. FLMNH-MGCL1174132 and genitalia slide no.
<collectionCode id="94D06EE7311A1132A31EFC72FC9DFBE4" box="[801,840,1022,1048]" country="France" name="Laboratoire de Biostratigraphie" pageId="5" pageNumber="558">UB</collectionCode>
˗12961-female (deposited in
<collectionCode id="94D06EE7311A1132A4B3FC72FB37FBE4" box="[1164,1250,1022,1048]" pageId="5" pageNumber="558">MGCL</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="BADBA5A9311A1132A0F8FBAEFBA1FBC0" box="[199,1140,1058,1084]" pageId="5" pageNumber="558" type="discussion">
<paragraph id="F27EF622311A1132A0F8FBAEFBA1FBC0" blockId="5.[151,1437,877,1084]" box="[199,1140,1058,1084]" pageId="5" pageNumber="558">
<emphasis id="C0B52A30311A1132A0F8FBAEFEEEFBC0" bold="true" box="[199,315,1058,1084]" pageId="5" pageNumber="558">Remarks.</emphasis>
This species is herein newly recorded from
<collectingCountry id="8AD6B6B2311A1132A315FBAEFC5CFBC0" box="[810,905,1058,1084]" name="Vietnam" pageId="5" pageNumber="558">Vietnam</collectingCountry>
(Hagiang Province).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,206 @@
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<mods:title id="8D3E241CF0F3E83B3C642644F7B65890">Review of the genus Diduga (Lepidoptera, Erebidae, Arctiinae) of Vietnam, with a new species and two newly recorded species</mods:title>
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<mods:namePart id="F41264798C5A24BF27139855BA1BDE87">Bayarsaikhan, Ulziijargal</mods:namePart>
<mods:affiliation id="2F17CCC6681A5603FD9B6F3F0C38E8CB">Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. &amp; Bio ˗ Resource and Environmental Center, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea.</mods:affiliation>
</mods:name>
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<mods:namePart id="924FD65649F3EFB405FF828E10C53C83">Heppner, John B.</mods:namePart>
<mods:affiliation id="805DC169CDCAC731D611F59E2EC1143A">McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Gainesville, Florida, USA.</mods:affiliation>
</mods:name>
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<mods:namePart id="160596CC06A8C64B131912145C263575">Kwon, Hyung Wook</mods:namePart>
<mods:affiliation id="71720034D52D27119AEDB299826868F8">Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. &amp; Bio ˗ Resource and Environmental Center, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea.</mods:affiliation>
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</mods:role>
<mods:namePart id="BB81AD410E58E24D520A0E210E30AC8F">Bae, Yang-Seop</mods:namePart>
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<mods:affiliation id="830B88077AC943824A6672CFFF8C6688">Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. &amp; Bio ˗ Resource and Environmental Center, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. &amp; Convergence Research Center for Insect Vectors, Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, Songdo ˗ dong, Incheon, 22012, Republic of Korea.</mods:affiliation>
<mods:nameIdentifier id="F8BBCEE87574042EFEDD2F7715685819" type="email">baeys@inu.ac.kr</mods:nameIdentifier>
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<treatment id="7A684734311A1132A0A8FF1BFA90FDA0" ID-DOI="http://doi.org/10.5281/zenodo.13745457" ID-Zenodo-Dep="13745457" LSID="urn:lsid:plazi:treatment:7A684734311A1132A0A8FF1BFA90FDA0" httpUri="http://treatment.plazi.org/id/7A684734311A1132A0A8FF1BFA90FDA0" lastPageNumber="558" pageId="5" pageNumber="558">
<subSubSection id="BADBA5A9311A1132A0A8FF1BFD05FF4D" box="[151,720,151,178]" pageId="5" pageNumber="558" type="nomenclature">
<paragraph id="F27EF622311A1132A0A8FF1BFD05FF4D" blockId="5.[151,720,151,213]" box="[151,720,151,178]" pageId="5" pageNumber="558">
<heading id="A936414E311A1132A0A8FF1BFD05FF4D" bold="true" box="[151,720,151,178]" fontSize="11" level="1" pageId="5" pageNumber="558" reason="1">
<taxonomicName id="35C18DA1311A1132A0A8FF1BFD05FF4D" ID-CoL="BHR9N" authority="Bayarsaikhan &amp; Bae, 2019" authorityName="Bayarsaikhan &amp; Bae" authorityYear="2019" box="[151,720,151,178]" class="Insecta" family="Erebidae" genus="Diduga" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="558" phylum="Arthropoda" rank="species" species="hanoiensis">
<emphasis id="C0B52A30311A1132A0A8FF1BFD05FF4D" bold="true" box="[151,720,151,178]" pageId="5" pageNumber="558">
<emphasis id="C0B52A30311A1132A0A8FF1BFEAFFF4E" bold="true" box="[151,378,151,178]" italics="true" pageId="5" pageNumber="558">Diduga hanoiensis</emphasis>
<bibRefCitation id="96508BD3311A1132A1BDFF1BFD05FF4D" author="Bayarsaikhan, U. &amp; Bae, Y. S." box="[386,720,151,177]" pageId="5" pageNumber="558" pagination="365 - 376" refId="ref4259" refString="Bayarsaikhan, U. &amp; Bae, Y. S. (2019) Four new and one newly recorded species of Diduga Moore, [1887] (Lepidoptera, Erebidae, Arctiinae) from Vietnam, with redescription of the little known species Diduga haematomiformis van Eecke, 1920. Zootaxa, 4624 (3), 365 - 376. https: // doi. org / 10.11646 / zootaxa. 4624.3.5" type="journal article" year="2019">Bayarsaikhan &amp; Bae, 2019</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BADBA5A9311A1132A0A8FF37FEACFF29" box="[151,377,187,213]" pageId="5" pageNumber="558" type="description">
<paragraph id="F27EF622311A1132A0A8FF37FEACFF29" blockId="5.[151,720,151,213]" box="[151,377,187,213]" pageId="5" pageNumber="558">
(
<figureCitation id="6AFAEAA7311A1132A0A0FF37FECBFF29" box="[159,286,187,213]" captionStart="FIGURES 15" captionStartId="2.[151,264,1840,1864]" captionTargetBox="[212,1398,179,1762]" captionTargetId="figure-24@2.[151,1436,179,1814]" captionTargetPageId="2" captionText="FIGURES 15. Adults of the Diduga from Vietnam. 1a. D. alternota Bucsek, male. 1b. ditto, female. 2a. D. bayartogtokhi Bayarsaikhan &amp; Bae, male. 2b. ditto, female. 3a. D. hanoiensis Bayarsaikhan &amp; Bae, male. 3b. ditto, female. 4. D. hastata Bayarsaikhan &amp; Bae, male. 5. D. nigridentata Bayarsaikhan &amp; Bae, male." figureDoi="http://doi.org/10.5281/zenodo.13745445" httpUri="https://zenodo.org/record/13745445/files/figure.png" pageId="5" pageNumber="558">Figs 3a, 3b</figureCitation>
,
<figureCitation id="6AFAEAA7311A1132A115FF37FE92FF29" box="[298,327,187,213]" captionStart="FIGURES 915" captionStartId="4.[151,264,1901,1925]" captionTargetBox="[160,1427,319,1868]" captionTargetId="figure-69@4.[151,1436,309,1878]" captionTargetPageId="4" captionText="FIGURES 915. Male genitalia of the Diduga from Vietnam. 9. D. alternota Bucsek, UB-12951Vietnam. 10. D. bayartogtokhi Bayarsaikhan&amp; Bae, UB-12958Vietnam.11.D.hanoiensis Bayarsaikhan&amp; Bae, UB-12976Vietnam.12.D.hastataBayarsaikhan &amp; Bae, UB-12966Vietnam. 13. D. nigridentata Bayarsaikhan &amp; Bae, UB-12957Vietnam. 14. D. quinquicornuta Bayarsaikhan &amp; Bae, UB-12962Vietnam. 15. D. robdevosi Bayarsaikhan &amp; Bae, UB-12970Vietnam." figureDoi="http://doi.org/10.5281/zenodo.13745453" httpUri="https://zenodo.org/record/13745453/files/figure.png" pageId="5" pageNumber="558">11</figureCitation>
,
<figureCitation id="6AFAEAA7311A1132A16CFF37FEA5FF29" box="[339,368,187,213]" captionStart="FIGURES 1822" captionStartId="8.[151,264,1908,1932]" captionTargetBox="[161,1429,571,1875]" captionTargetId="figure-157@8.[151,1436,561,1885]" captionTargetPageId="8" captionText="FIGURES 1822. Female genitalia of the Diduga from Vietnam. 18. D. alternota Bucsek, UB-12981Vietnam. 19. D. bayartogtokhi Bayarsaikhan &amp;Bae, UB-12965Vietnam.20.D.hanoiensis Bayarsaikhan &amp;Bae, UB-12969Vietnam.21.D.laocai Bayarsaikhan &amp; Heppner, sp. nov., paratype, UB-12956Vietnam. 22. D. robdevosi Bayarsaikhan &amp; Bae, UB-12971Vietnam." figureDoi="http://doi.org/10.5281/zenodo.13745459" httpUri="https://zenodo.org/record/13745459/files/figure.png" pageId="5" pageNumber="558">20</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="BADBA5A9311A1132A0A8FE8CFA90FEE4" box="[151,1349,255,280]" pageId="5" pageNumber="558" type="reference_group">
<paragraph id="F27EF622311A1132A0A8FE8CFA90FEE4" blockId="5.[151,1349,255,280]" box="[151,1349,255,280]" pageId="5" pageNumber="558">
<taxonomicName id="35C18DA1311A1132A0A8FE8CFD44FEE4" ID-CoL="BHR9K" authority="Bayarsaikhan &amp; Bae, 2019" authorityName="Bayarsaikhan &amp; Bae" authorityYear="2019" box="[151,657,256,280]" class="Insecta" family="Erebidae" genus="Diduga" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="558" phylum="Arthropoda" rank="species" species="bayartogtokhi">
<emphasis id="C0B52A30311A1132A0A8FE8CFEADFEE4" box="[151,376,256,280]" italics="true" pageId="5" pageNumber="558">Diduga bayartogtokhi</emphasis>
<bibRefCitation id="96508BD3311A1132A141FE8CFD44FEE4" author="Bayarsaikhan, U. &amp; Bae, Y. S." box="[382,657,256,280]" pageId="5" pageNumber="558" pagination="365 - 376" refId="ref4259" refString="Bayarsaikhan, U. &amp; Bae, Y. S. (2019) Four new and one newly recorded species of Diduga Moore, [1887] (Lepidoptera, Erebidae, Arctiinae) from Vietnam, with redescription of the little known species Diduga haematomiformis van Eecke, 1920. Zootaxa, 4624 (3), 365 - 376. https: // doi. org / 10.11646 / zootaxa. 4624.3.5" type="journal article" year="2019">Bayarsaikhan &amp; Bae, 2019</bibRefCitation>
</taxonomicName>
,
<emphasis id="C0B52A30311A1132A2A3FE8CFD3AFEEB" box="[668,751,256,279]" italics="true" pageId="5" pageNumber="558">Zootaxa</emphasis>
4624 (3): 368, figs. 4, 10, 16. TL:
<collectingCountry id="8AD6B6B2311A1132A470FE8CFB70FEE4" box="[1103,1189,256,280]" name="Vietnam" pageId="5" pageNumber="558">Vietnam</collectingCountry>
(
<collectingCountry id="8AD6B6B2311A1132A48CFE8CFB2DFEE4" box="[1203,1272,256,280]" name="Turkey" pageId="5" pageNumber="558">Hatay</collectingCountry>
, BaVi).
</paragraph>
</subSubSection>
<subSubSection id="BADBA5A9311A1132A0A8FEC9FB8EFDE8" pageId="5" pageNumber="558" type="distribution">
<paragraph id="F27EF622311A1132A0A8FEC9FE48FE9C" blockId="5.[151,1436,325,604]" box="[151,413,325,352]" pageId="5" pageNumber="558">
<emphasis id="C0B52A30311A1132A0A8FEC9FEE4FEA3" bold="true" box="[151,305,325,351]" pageId="5" pageNumber="558">Distribution.</emphasis>
<collectingCountry id="8AD6B6B2311A1132A108FECAFE4FFE9C" box="[311,410,326,352]" name="Vietnam" pageId="5" pageNumber="558">Vietnam</collectingCountry>
.
</paragraph>
<paragraph id="F27EF622311A1132A0F8FEE5FB8EFDE8" blockId="5.[151,1436,325,604]" pageId="5" pageNumber="558">
<emphasis id="C0B52A30311A1132A0F8FEE5FDE9FE7F" bold="true" box="[199,572,361,387]" pageId="5" pageNumber="558">Additional material examined.</emphasis>
<materialsCitation id="42A9FC7F311A1132A278FEE6FC0EFE54" collectingDate="2008-06-21" collectingDateMax="2008-06-26" collectingDateMin="2008-06-21" collectionCode="UB" collectorName="Ba Be N. P. &amp; J. B. Heppner" country="Vietnam" elevation="240" location="Bac Kan" pageId="5" pageNumber="558" specimenCount="1" specimenCount-female="1" stateProvince="Bac Kan">
<specimenCount id="E4C73DAB311A1132A278FEE6FDBCFE78" box="[583,617,362,388]" count="1" pageId="5" pageNumber="558" type="female">1♀</specimenCount>
,
<collectingCountry id="8AD6B6B2311A1132A244FEE6FD08FE78" box="[635,733,362,388]" name="Vietnam" pageId="5" pageNumber="558">Vietnam</collectingCountry>
,
<collectingRegion id="300538C0311A1132A2D4FEE6FC86FE78" box="[747,851,362,388]" country="Vietnam" name="Bac Kan" pageId="5" pageNumber="558">Bac Kan</collectingRegion>
,
<collectorName id="5F3493F4311A1132A35DFEE6FC30FE78" box="[866,997,362,388]" pageId="5" pageNumber="558">Ba Be N.P.</collectorName>
(lodge),
<quantity id="35395BC7311A1132A46CFEE6FB40FE78" box="[1107,1173,362,388]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="5" pageNumber="558" unit="m" value="240.0">
<elevation id="79EC1111311A1132A46CFEE6FB40FE78" box="[1107,1173,362,388]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="5" pageNumber="558" unit="m" value="240.0">240m</elevation>
</quantity>
,
<date id="867FD0E2311A1132A49BFEE6FAB7FE78" box="[1188,1378,362,388]" pageId="5" pageNumber="558" value="2008-06-21" valueMax="2008-06-26" valueMin="2008-06-21">
<collectingDate id="963B290A311A1132A49BFEE6FAB7FE78" box="[1188,1378,362,388]" pageId="5" pageNumber="558" value="2008-06-21" valueMax="2008-06-26" valueMin="2008-06-21">2126 Jun 2008</collectingDate>
</date>
,
<collectorName id="5F3493F4311A1132A54EFEE6FF2CFE54" pageId="5" pageNumber="558">J.B. Heppner</collectorName>
, adult no. FLMNH-MGCL1174184 and genitalia slide no.
<collectionCode id="94D06EE7311A1132A38CFE02FC0EFE54" box="[947,987,398,424]" country="France" name="Laboratoire de Biostratigraphie" pageId="5" pageNumber="558">UB</collectionCode>
</materialsCitation>
˗12969;
<materialsCitation id="42A9FC7F311A1132A403FE02FF3DFE0C" collectingDate="2009-05-04" collectingDateMax="2009-05-07" collectingDateMin="2009-05-04" collectionCode="UB" collectorName="Cuc Phuong N. P. &amp; J. B. Heppner" country="Vietnam" location="Mac Lk." pageId="5" pageNumber="558" specimenCount="1" specimenCount-female="1" stateProvince="Ninh Binh">
<specimenCount id="E4C73DAB311A1132A403FE02FB8AFE54" box="[1084,1119,398,424]" count="1" pageId="5" pageNumber="558" type="female">1♀</specimenCount>
,
<collectingCountry id="8AD6B6B2311A1132A44EFE02FB06FE54" box="[1137,1235,398,424]" name="Vietnam" pageId="5" pageNumber="558">Vietnam</collectingCountry>
,
<collectingRegion id="300538C0311A1132A4DDFE02FA8AFE54" box="[1250,1375,398,424]" country="Vietnam" name="Ninh Binh" pageId="5" pageNumber="558">Ninh Binh</collectingRegion>
,
<collectorName id="5F3493F4311A1132A550FE02FEF0FE30" pageId="5" pageNumber="558">Cuc Phuong N.P.</collectorName>
,
<location id="F71EA0F9311A1132A10EFE3EFE46FE30" LSID="urn:lsid:plazi:treatment:7A684734311A1132A0A8FF1BFA90FDA0:F71EA0F9311A1132A10EFE3EFE46FE30" box="[305,403,434,460]" country="Vietnam" name="Mac Lk." pageId="5" pageNumber="558" stateProvince="Ninh Binh">Mac Lk.</location>
,
<date id="867FD0E2311A1132A1A0FE3EFD93FE30" box="[415,582,434,460]" pageId="5" pageNumber="558" value="2009-05-04" valueMax="2009-05-07" valueMin="2009-05-04">
<collectingDate id="963B290A311A1132A1A0FE3EFD93FE30" box="[415,582,434,460]" pageId="5" pageNumber="558" value="2009-05-04" valueMax="2009-05-07" valueMin="2009-05-04">
47 May
<quantity id="35395BC7311A1132A234FE3EFD93FE30" box="[523,582,434,460]" metricMagnitude="3" metricUnit="m" metricValue="2.009155" pageId="5" pageNumber="558" unit="m" value="2009.155">2009</quantity>
</collectingDate>
</date>
, 155m,
<collectorName id="5F3493F4311A1132A29FFE3EFCE0FE30" box="[672,821,434,460]" pageId="5" pageNumber="558">J.B. Heppner</collectorName>
, adult no. FLMNH-MGCL1174187 and genitalia slide no.
<collectionCode id="94D06EE7311A1132A0FFFE5AFF3DFE0C" box="[192,232,470,496]" country="France" name="Laboratoire de Biostratigraphie" pageId="5" pageNumber="558">UB</collectionCode>
</materialsCitation>
˗12972;
<materialsCitation id="42A9FC7F311A1132A17BFE5AFC8BFDE8" collectingDate="2010-07-19" collectingDateMax="2010-07-23" collectingDateMin="2010-07-19" collectionCode="UB" collectorName="J. B. Heppner" country="Vietnam" county="Prov." elevation="455" location="Ba Vi Natl Pk." municipality="Ba Vi Natl Pk." pageId="5" pageNumber="558" specimenCount="3" specimenCount-male="3">
<specimenCount id="E4C73DAB311A1132A17BFE5AFEBFFE0C" box="[324,362,470,496]" count="3" pageId="5" pageNumber="558" type="male">3♂</specimenCount>
,
<collectingCountry id="8AD6B6B2311A1132A14BFE5AFE03FE0C" box="[372,470,470,496]" name="Vietnam" pageId="5" pageNumber="558">Vietnam</collectingCountry>
, Hatay
<collectingCounty id="1B1F8EAE311A1132A217FE5AFDB3FE0C" box="[552,614,470,496]" pageId="5" pageNumber="558">Prov.</collectingCounty>
,
<location id="F71EA0F9311A1132A24FFE5AFCC7FE0C" LSID="urn:lsid:plazi:treatment:7A684734311A1132A0A8FF1BFA90FDA0:F71EA0F9311A1132A24FFE5AFCC7FE0C" box="[624,786,470,496]" country="Vietnam" county="Prov." municipality="Ba Vi Natl Pk." name="Ba Vi Natl Pk." pageId="5" pageNumber="558">
<collectingMunicipality id="121A6C58311A1132A24FFE5AFCC7FE0C" box="[624,786,470,496]" pageId="5" pageNumber="558">Ba Vi Natl Pk.</collectingMunicipality>
</location>
,
<quantity id="35395BC7311A1132A323FE5AFC8AFE0D" box="[796,863,470,497]" metricMagnitude="2" metricUnit="m" metricValue="4.55" pageId="5" pageNumber="558" unit="m" value="455.0">
<elevation id="79EC1111311A1132A323FE5AFC8AFE0D" box="[796,863,470,497]" metricMagnitude="2" metricUnit="m" metricValue="4.55" pageId="5" pageNumber="558" unit="m" value="455.0">455m</elevation>
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,
<date id="867FD0E2311A1132A356FE5AFBC2FE0C" box="[873,1047,470,496]" pageId="5" pageNumber="558" value="2010-07-19" valueMax="2010-07-23" valueMin="2010-07-19">
<collectingDate id="963B290A311A1132A356FE5AFBC2FE0C" box="[873,1047,470,496]" pageId="5" pageNumber="558" value="2010-07-19" valueMax="2010-07-23" valueMin="2010-07-19">1923 Jul 2010</collectingDate>
</date>
,
<collectorName id="5F3493F4311A1132A41EFE5AFB60FE0C" box="[1057,1205,470,496]" pageId="5" pageNumber="558">J.B. Heppner</collectorName>
, adults no. FLMNH-MGCL1174190, 1174191, 1174192 and genitalia slides no.
<collectionCode id="94D06EE7311A1132A309FE76FC8BFDE8" box="[822,862,506,532]" country="France" name="Laboratoire de Biostratigraphie" pageId="5" pageNumber="558">UB</collectionCode>
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˗12974, 12975, 12976.
</paragraph>
</subSubSection>
<subSubSection id="BADBA5A9311A1132A0F8FD92FA90FDA0" pageId="5" pageNumber="558" type="discussion">
<paragraph id="F27EF622311A1132A0F8FD92FA90FDA0" blockId="5.[151,1436,325,604]" pageId="5" pageNumber="558">
<emphasis id="C0B52A30311A1132A0F8FD92FEEEFDC4" bold="true" box="[199,315,542,568]" pageId="5" pageNumber="558">Remarks.</emphasis>
This species was reported from
<collectingCountry id="8AD6B6B2311A1132A2F2FD92FCE3FDC4" box="[717,822,542,568]" name="Vietnam" pageId="5" pageNumber="558">Vietnam</collectingCountry>
(
<collectingCountry id="8AD6B6B2311A1132A375FD92FC42FDC4" box="[842,919,542,568]" name="Turkey" pageId="5" pageNumber="558">Hatay</collectingCountry>
(
<collectingRegion id="300538C0311A1132A395FD92FC2FFDC4" box="[938,1018,542,568]" country="Vietnam" name="Ha Noi" pageId="5" pageNumber="558">Hanoi</collectingRegion>
) Province) by
<bibRefCitation id="96508BD3311A1132A483FD92FEFDFDA0" author="Bayarsaikhan, U. &amp; Bae, Y. S." pageId="5" pageNumber="558" pagination="365 - 376" refId="ref4259" refString="Bayarsaikhan, U. &amp; Bae, Y. S. (2019) Four new and one newly recorded species of Diduga Moore, [1887] (Lepidoptera, Erebidae, Arctiinae) from Vietnam, with redescription of the little known species Diduga haematomiformis van Eecke, 1920. Zootaxa, 4624 (3), 365 - 376. https: // doi. org / 10.11646 / zootaxa. 4624.3.5" type="journal article" year="2019">Bayarsaikhan and Bae (2019)</bibRefCitation>
. In this study, we report it for
<collectingRegion id="300538C0311A1132A246FDCEFD0EFDA0" box="[633,731,578,604]" country="Vietnam" name="Bac Kan" pageId="5" pageNumber="558">Bac Kan</collectingRegion>
and
<collectingRegion id="300538C0311A1132A32EFDCEFC53FDA0" box="[785,902,578,604]" country="Vietnam" name="Ninh Binh" pageId="5" pageNumber="558">Ninh Binh</collectingRegion>
Provinces of
<collectingCountry id="8AD6B6B2311A1132A41EFDCEFB55FDA0" box="[1057,1152,578,604]" name="Vietnam" pageId="5" pageNumber="558">Vietnam</collectingCountry>
for the first time.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="9B1108F1953A3DE911DAACAA046D637E">Review of the genus Diduga (Lepidoptera, Erebidae, Arctiinae) of Vietnam, with a new species and two newly recorded species</mods:title>
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<emphasis id="C0B52A30311C1134A0A8F8ABFE5BF8BE" bold="true" box="[151,398,1831,1858]" italics="true" pageId="3" pageNumber="556">Diduga cucphuonga</emphasis>
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15 (3): 237, figs. 18, 37. TL:
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NP).
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<emphasis id="C0B52A30311C1134A0A8F85DFEE4F817" bold="true" box="[151,305,2001,2027]" pageId="3" pageNumber="556">Distribution.</emphasis>
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,
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,
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<collectingDate id="963B290A311B1133A1B6FF36FDF6FF29" box="[393,547,186,213]" pageId="4" pageNumber="557" value="2012-12-03" valueMax="2012-12-05" valueMin="2012-12-03">35.XII 2012</collectingDate>
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,
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.
<collectorName id="5F3493F4311B1133A273FF37FD6FFF29" box="[588,698,187,213]" pageId="4" pageNumber="557">Zolotuhin</collectorName>
leg. (after
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).
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<emphasis id="C0B52A30311B1133A0F8FF53FEEEFF05" bold="true" box="[199,315,223,249]" pageId="4" pageNumber="557">Remarks.</emphasis>
This species was reported from
<collectingCountry id="8AD6B6B2311B1133A2DCFF53FC99FF05" box="[739,844,223,249]" name="Vietnam" pageId="4" pageNumber="557">Vietnam</collectingCountry>
(
<collectingRegion id="300538C0311B1133A35DFF53FBB4FF05" box="[866,1121,223,249]" country="Vietnam" name="Ninh Binh" pageId="4" pageNumber="557">Ninh Binh Province</collectingRegion>
) by
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.
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</treatment>
</document>

View file

@ -0,0 +1,241 @@
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<mods:affiliation id="42DC326C642870FBB588765E8E9997B7">Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. &amp; Bio ˗ Resource and Environmental Center, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea.</mods:affiliation>
</mods:name>
<mods:name id="693788F6E1369B1CA4D0BBC14C8FCF3C" type="personal">
<mods:role id="0347DD695DA5E1FF8047D446DBD70E46">
<mods:roleTerm id="4561EFE9837289FEFE6640DD1DCAB560">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CA34BA102209092F444F8DBADD530795">Heppner, John B.</mods:namePart>
<mods:affiliation id="4554B614E9653F855AE591A70705AF98">McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Gainesville, Florida, USA.</mods:affiliation>
</mods:name>
<mods:name id="4570E8C5996CE3221143271E59D11599" type="personal">
<mods:role id="532C769D024D84B045913104C6211C3B">
<mods:roleTerm id="B3A0994D56BC7EBBEB795E21D96C7E4F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="18024097BD56BA630DC0A7B4849CD239">Kwon, Hyung Wook</mods:namePart>
<mods:affiliation id="8E008660C28B72E876C0F8B443CD2B37">Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. &amp; Bio ˗ Resource and Environmental Center, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea.</mods:affiliation>
</mods:name>
<mods:name id="16BD633848C908E1A1CDC49F2E446BE6" type="personal">
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<mods:namePart id="4D83025943141571EF0B9D941E77E27B">Bae, Yang-Seop</mods:namePart>
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<mods:affiliation id="895C53C4D6A9258988C6ABCA978F69C8">Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. &amp; Bio ˗ Resource and Environmental Center, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. &amp; Convergence Research Center for Insect Vectors, Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, Songdo ˗ dong, Incheon, 22012, Republic of Korea.</mods:affiliation>
<mods:nameIdentifier id="BAAB58B538A2B13A93EF0970562398EC" type="email">baeys@inu.ac.kr</mods:nameIdentifier>
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<mods:title id="E3C41614DABD42077D6F9CC62560D7E6">Zootaxa</mods:title>
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<mods:date id="1E4EACAEE54ECF8074C15222536144F6">2024</mods:date>
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<mods:number id="028C488696DC7B72B09855B87B33DCE1">5501</mods:number>
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<treatment id="7A684734311E1134A0A8F89FFAA5FE75" ID-DOI="http://doi.org/10.5281/zenodo.13745447" ID-Zenodo-Dep="13745447" LSID="urn:lsid:plazi:treatment:7A684734311E1134A0A8F89FFAA5FE75" httpUri="http://treatment.plazi.org/id/7A684734311E1134A0A8F89FFAA5FE75" lastPageId="3" lastPageNumber="556" pageId="1" pageNumber="554">
<subSubSection id="BADBA5A9311E1136A0A8F89FFD23F8D1" box="[151,758,1811,1838]" pageId="1" pageNumber="554" type="nomenclature">
<paragraph id="F27EF622311E1136A0A8F89FFD23F8D1" blockId="1.[151,758,1811,1873]" box="[151,758,1811,1838]" pageId="1" pageNumber="554">
<heading id="A936414E311E1136A0A8F89FFD23F8D1" bold="true" box="[151,758,1811,1838]" fontSize="11" level="1" pageId="1" pageNumber="554" reason="1">
<taxonomicName id="35C18DA1311E1136A0A8F89FFD23F8D1" ID-CoL="BHR9K" authority="Bayarsaikhan &amp; Bae, 2019" authorityName="Bayarsaikhan &amp; Bae" authorityYear="2019" box="[151,758,1811,1838]" class="Insecta" family="Erebidae" genus="Diduga" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="bayartogtokhi">
<emphasis id="C0B52A30311E1136A0A8F89FFD23F8D1" bold="true" box="[151,758,1811,1838]" pageId="1" pageNumber="554">
<emphasis id="C0B52A30311E1136A0A8F89FFE75F8D2" bold="true" box="[151,416,1811,1838]" italics="true" pageId="1" pageNumber="554">Diduga bayartogtokhi</emphasis>
<bibRefCitation id="96508BD3311E1136A198F89FFD23F8D1" author="Bayarsaikhan, U. &amp; Bae, Y. S." box="[423,758,1811,1837]" pageId="1" pageNumber="554" pagination="365 - 376" refId="ref4259" refString="Bayarsaikhan, U. &amp; Bae, Y. S. (2019) Four new and one newly recorded species of Diduga Moore, [1887] (Lepidoptera, Erebidae, Arctiinae) from Vietnam, with redescription of the little known species Diduga haematomiformis van Eecke, 1920. Zootaxa, 4624 (3), 365 - 376. https: // doi. org / 10.11646 / zootaxa. 4624.3.5" type="journal article" year="2019">Bayarsaikhan &amp; Bae, 2019</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BADBA5A9311E1136A0A8F8BBFEAFF8AD" box="[151,378,1847,1873]" pageId="1" pageNumber="554" type="description">
<paragraph id="F27EF622311E1136A0A8F8BBFEAFF8AD" blockId="1.[151,758,1811,1873]" box="[151,378,1847,1873]" pageId="1" pageNumber="554">
(
<figureCitation id="6AFAEAA7311E1136A0A0F8BBFECAF8AD" box="[159,287,1847,1873]" captionStart="FIGURES 15" captionStartId="2.[151,264,1840,1864]" captionTargetBox="[212,1398,179,1762]" captionTargetId="figure-24@2.[151,1436,179,1814]" captionTargetPageId="2" captionText="FIGURES 15. Adults of the Diduga from Vietnam. 1a. D. alternota Bucsek, male. 1b. ditto, female. 2a. D. bayartogtokhi Bayarsaikhan &amp; Bae, male. 2b. ditto, female. 3a. D. hanoiensis Bayarsaikhan &amp; Bae, male. 3b. ditto, female. 4. D. hastata Bayarsaikhan &amp; Bae, male. 5. D. nigridentata Bayarsaikhan &amp; Bae, male." figureDoi="http://doi.org/10.5281/zenodo.13745445" httpUri="https://zenodo.org/record/13745445/files/figure.png" pageId="1" pageNumber="554">Figs 2a, 2b</figureCitation>
,
<figureCitation id="6AFAEAA7311E1136A115F8BBFE9DF8AD" box="[298,328,1847,1873]" captionStart="FIGURES 915" captionStartId="4.[151,264,1901,1925]" captionTargetBox="[160,1427,319,1868]" captionTargetId="figure-69@4.[151,1436,309,1878]" captionTargetPageId="4" captionText="FIGURES 915. Male genitalia of the Diduga from Vietnam. 9. D. alternota Bucsek, UB-12951Vietnam. 10. D. bayartogtokhi Bayarsaikhan&amp; Bae, UB-12958Vietnam.11.D.hanoiensis Bayarsaikhan&amp; Bae, UB-12976Vietnam.12.D.hastataBayarsaikhan &amp; Bae, UB-12966Vietnam. 13. D. nigridentata Bayarsaikhan &amp; Bae, UB-12957Vietnam. 14. D. quinquicornuta Bayarsaikhan &amp; Bae, UB-12962Vietnam. 15. D. robdevosi Bayarsaikhan &amp; Bae, UB-12970Vietnam." figureDoi="http://doi.org/10.5281/zenodo.13745453" httpUri="https://zenodo.org/record/13745453/files/figure.png" pageId="1" pageNumber="554">10</figureCitation>
,
<figureCitation id="6AFAEAA7311E1136A16BF8BBFEA4F8AD" box="[340,369,1847,1873]" captionStart="FIGURES 1822" captionStartId="8.[151,264,1908,1932]" captionTargetBox="[161,1429,571,1875]" captionTargetId="figure-157@8.[151,1436,561,1885]" captionTargetPageId="8" captionText="FIGURES 1822. Female genitalia of the Diduga from Vietnam. 18. D. alternota Bucsek, UB-12981Vietnam. 19. D. bayartogtokhi Bayarsaikhan &amp;Bae, UB-12965Vietnam.20.D.hanoiensis Bayarsaikhan &amp;Bae, UB-12969Vietnam.21.D.laocai Bayarsaikhan &amp; Heppner, sp. nov., paratype, UB-12956Vietnam. 22. D. robdevosi Bayarsaikhan &amp; Bae, UB-12971Vietnam." figureDoi="http://doi.org/10.5281/zenodo.13745459" httpUri="https://zenodo.org/record/13745459/files/figure.png" pageId="1" pageNumber="554">19</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="BADBA5A9311E1136A0A8F8F0FA43F868" box="[151,1430,1915,1940]" pageId="1" pageNumber="554" type="reference_group">
<paragraph id="F27EF622311E1136A0A8F8F0FA43F868" blockId="1.[151,1430,1915,1940]" box="[151,1430,1915,1940]" pageId="1" pageNumber="554">
<taxonomicName id="35C18DA1311E1136A0A8F8F0FD44F868" ID-CoL="BHR9K" authority="Bayarsaikhan &amp; Bae, 2019" authorityName="Bayarsaikhan &amp; Bae" authorityYear="2019" box="[151,657,1916,1940]" class="Insecta" family="Erebidae" genus="Diduga" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="bayartogtokhi">
<emphasis id="C0B52A30311E1136A0A8F8F0FEADF868" box="[151,376,1916,1940]" italics="true" pageId="1" pageNumber="554">Diduga bayartogtokhi</emphasis>
<bibRefCitation id="96508BD3311E1136A141F8F0FD44F868" author="Bayarsaikhan, U. &amp; Bae, Y. S." box="[382,657,1916,1940]" pageId="1" pageNumber="554" pagination="365 - 376" refId="ref4259" refString="Bayarsaikhan, U. &amp; Bae, Y. S. (2019) Four new and one newly recorded species of Diduga Moore, [1887] (Lepidoptera, Erebidae, Arctiinae) from Vietnam, with redescription of the little known species Diduga haematomiformis van Eecke, 1920. Zootaxa, 4624 (3), 365 - 376. https: // doi. org / 10.11646 / zootaxa. 4624.3.5" type="journal article" year="2019">Bayarsaikhan &amp; Bae, 2019</bibRefCitation>
</taxonomicName>
,
<emphasis id="C0B52A30311E1136A2A3F8F0FD3AF86F" box="[668,751,1916,1939]" italics="true" pageId="1" pageNumber="554">Zootaxa</emphasis>
4624 (3): 368, figs. 5, 11, 17. TL:
<collectingCountry id="8AD6B6B2311E1136A471F8F0FB71F868" box="[1102,1188,1916,1940]" name="Vietnam" pageId="1" pageNumber="554">Vietnam</collectingCountry>
(
<collectingRegion id="300538C0311E1136A48EF8F0FACAF868" box="[1201,1311,1916,1940]" country="Vietnam" name="Vinh Phuc" pageId="1" pageNumber="554">Vinh Phuc</collectingRegion>
, Tam Dao).
</paragraph>
</subSubSection>
<subSubSection id="BADBA5A9311E1134A0A8F84DFDDAFEBD" lastPageId="3" lastPageNumber="556" pageId="1" pageNumber="554" type="distribution">
<paragraph id="F27EF622311E1136A0A8F84DFE48F820" blockId="1.[151,413,1985,2012]" box="[151,413,1985,2012]" pageId="1" pageNumber="554">
<emphasis id="C0B52A30311E1136A0A8F84DFEE4F827" bold="true" box="[151,305,1985,2011]" pageId="1" pageNumber="554">Distribution.</emphasis>
<collectingCountry id="8AD6B6B2311E1136A108F84EFE4FF820" box="[311,410,1986,2012]" name="Vietnam" pageId="1" pageNumber="554">Vietnam</collectingCountry>
.
</paragraph>
<caption id="A6BEA6AA311D1135A0A8F8BCFC53F86C" ID-DOI="http://doi.org/10.5281/zenodo.13745445" ID-Zenodo-Dep="13745445" httpUri="https://zenodo.org/record/13745445/files/figure.png" pageId="2" pageNumber="555" startId="2.[151,264,1840,1864]" targetBox="[212,1398,179,1762]" targetPageId="2" targetType="figure">
<paragraph id="F27EF622311D1135A0A8F8BCFC53F86C" blockId="2.[151,1437,1840,1937]" pageId="2" pageNumber="555">
<emphasis id="C0B52A30311D1135A0A8F8BCFEE8F8B4" bold="true" box="[151,317,1840,1864]" pageId="2" pageNumber="555">FIGURES 15.</emphasis>
Adults of the
<taxonomicName id="35C18DA1311D1135A1EAF8BCFDF4F8B4" authorityName="Moore" authorityYear="1887" box="[469,545,1840,1864]" class="Insecta" family="Erebidae" genus="Diduga" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="555" phylum="Arthropoda" rank="genus">
<emphasis id="C0B52A30311D1135A1EAF8BCFDF4F8B4" box="[469,545,1840,1864]" italics="true" pageId="2" pageNumber="555">Diduga</emphasis>
</taxonomicName>
from Vietnam. 1a.
<taxonomicName id="35C18DA1311D1135A2D2F8BDFC14F8B4" authority="Bucsek" authorityName="Bucsek" authorityYear="2014" box="[749,961,1840,1864]" class="Insecta" family="Erebidae" genus="Diduga" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="555" phylum="Arthropoda" rank="species" species="alternota">
<emphasis id="C0B52A30311D1135A2D2F8BDFCB9F8B4" box="[749,876,1840,1864]" italics="true" pageId="2" pageNumber="555">D. alternota</emphasis>
Bucsek
</taxonomicName>
, male. 1b.
<emphasis id="C0B52A30311D1135A40AF8BCFBB6F8B4" box="[1077,1123,1840,1864]" italics="true" pageId="2" pageNumber="555">ditto</emphasis>
, female. 2a.
<taxonomicName id="35C18DA1311D1135A4D2F8BDFEA5F890" authority="Bayarsaikhan &amp; Bae" authorityName="Bayarsaikhan &amp; Bae" authorityYear="2019" class="Insecta" family="Erebidae" genus="Diduga" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="555" phylum="Arthropoda" rank="species" species="bayartogtokhi">
<emphasis id="C0B52A30311D1135A4D2F8BDFA49F8B4" box="[1261,1436,1840,1864]" italics="true" pageId="2" pageNumber="555">D. bayartogtokhi</emphasis>
Bayarsaikhan &amp; Bae
</taxonomicName>
, male. 2b.
<emphasis id="C0B52A30311D1135A1D9F8D8FDC1F890" box="[486,532,1876,1900]" italics="true" pageId="2" pageNumber="555">ditto</emphasis>
, female. 3a.
<taxonomicName id="35C18DA1311D1135A2A1F8D9FBDBF890" authority="Bayarsaikhan &amp; Bae" authorityName="Bayarsaikhan &amp; Bae" authorityYear="2019" box="[670,1038,1876,1901]" class="Insecta" family="Erebidae" genus="Diduga" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="555" phylum="Arthropoda" rank="species" species="hanoiensis">
<emphasis id="C0B52A30311D1135A2A1F8D9FCFEF890" box="[670,811,1876,1900]" italics="true" pageId="2" pageNumber="555">D. hanoiensis</emphasis>
Bayarsaikhan &amp; Bae
</taxonomicName>
, male. 3b.
<emphasis id="C0B52A30311D1135A4BCF8D8FB64F890" box="[1155,1201,1876,1900]" italics="true" pageId="2" pageNumber="555">ditto</emphasis>
, female. 4.
<taxonomicName id="35C18DA1311D1135A50FF8D9FEBEF86C" authority="Bayarsaikhan &amp; Bae" authorityName="Bayarsaikhan &amp; Bae" authorityYear="2021" class="Insecta" family="Erebidae" genus="Diduga" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="555" phylum="Arthropoda" rank="species" species="hastata">
<emphasis id="C0B52A30311D1135A50FF8D9FA48F890" box="[1328,1437,1876,1900]" italics="true" pageId="2" pageNumber="555">D. hastata</emphasis>
Bayarsaikhan &amp; Bae
</taxonomicName>
, male. 5.
<taxonomicName id="35C18DA1311D1135A1F2F8F5FC91F86C" authority="Bayarsaikhan &amp; Bae" authorityName="Bayarsaikhan &amp; Bae" authorityYear="2019" box="[461,836,1912,1937]" class="Insecta" family="Erebidae" genus="Diduga" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="555" phylum="Arthropoda" rank="species" species="nigridentata">
<emphasis id="C0B52A30311D1135A1F2F8F5FDBCF86C" box="[461,617,1912,1936]" italics="true" pageId="2" pageNumber="555">D. nigridentata</emphasis>
Bayarsaikhan &amp; Bae
</taxonomicName>
, male.
</paragraph>
</caption>
<paragraph id="F27EF622311C1134A0F8FF1AFDDAFEBD" blockId="3.[151,1437,150,393]" pageId="3" pageNumber="556">
<emphasis id="C0B52A30311C1134A0F8FF1AFDE1FF4C" bold="true" box="[199,564,150,176]" pageId="3" pageNumber="556">Additional material examined.</emphasis>
<materialsCitation id="42A9FC7F311C1134A204FF1BFCD9FF29" collectingDate="2014-10-06" collectingDateMax="2014-10-09" collectingDateMin="2014-10-06" collectionCode="UB" collectorName="J. B. Heppner" country="Vietnam" location="Tam Dao" pageId="3" pageNumber="556" specimenCount="1" specimenCount-male="1" stateProvince="Vinh Phuc">
<specimenCount id="E4C73DAB311C1134A204FF1BFD8AFF4D" box="[571,607,151,177]" count="1" pageId="3" pageNumber="556" type="male">1♂</specimenCount>
,
<collectingCountry id="8AD6B6B2311C1134A253FF1BFD1BFF4D" box="[620,718,151,177]" name="Vietnam" pageId="3" pageNumber="556">Vietnam</collectingCountry>
,
<collectingRegion id="300538C0311C1134A2E7FF1BFC85FF4D" box="[728,848,151,177]" country="Vietnam" name="Vinh Phuc" pageId="3" pageNumber="556">Vinh Phuc</collectingRegion>
,
<location id="F71EA0F9311C1134A364FF1BFC11FF4D" LSID="urn:lsid:plazi:treatment:7A684734311E1134A0A8F89FFAA5FE75:F71EA0F9311C1134A364FF1BFC11FF4D" box="[859,964,151,177]" country="Vietnam" name="Tam Dao" pageId="3" pageNumber="556" stateProvince="Vinh Phuc">Tam Dao</location>
,
<date id="867FD0E2311C1134A3EFFF1BFBBEFF4D" box="[976,1131,151,177]" pageId="3" pageNumber="556" value="2014-10-06" valueMax="2014-10-09" valueMin="2014-10-06">
<collectingDate id="963B290A311C1134A3EFFF1BFBBEFF4D" box="[976,1131,151,177]" pageId="3" pageNumber="556" value="2014-10-06" valueMax="2014-10-09" valueMin="2014-10-06">
69 Oct
<quantity id="35395BC7311C1134A40FFF1BFBBEFF4D" box="[1072,1131,151,177]" metricMagnitude="3" metricUnit="m" metricValue="2.01493" pageId="3" pageNumber="556" unit="m" value="2014.93">2014</quantity>
</collectingDate>
</date>
, 930m,
<collectorName id="5F3493F4311C1134A4FBFF1BFA8CFF4D" box="[1220,1369,151,177]" pageId="3" pageNumber="556">J.B. Heppner</collectorName>
, adult no. FLMNH-MGCL1174128 and genitalia slide no.
<collectionCode id="94D06EE7311C1134A2DBFF36FCD9FF29" box="[740,780,186,213]" country="France" name="Laboratoire de Biostratigraphie" pageId="3" pageNumber="556">UB</collectionCode>
</materialsCitation>
˗12958;
<materialsCitation id="42A9FC7F311C1134A356FF37FBA5FF05" collectingDate="2016-08-09" collectingDateMax="2016-08-11" collectingDateMin="2016-08-09" collectionCode="UB" collectorName="J. B. Heppner" country="Vietnam" location="Muong Khuong" pageId="3" pageNumber="556" specimenCount="1" specimenCount-female="1" stateProvince="Lao Cai">
<specimenCount id="E4C73DAB311C1134A356FF37FC59FF29" box="[873,908,187,213]" count="1" pageId="3" pageNumber="556" type="female">1♀</specimenCount>
,
<collectingCountry id="8AD6B6B2311C1134A3A5FF37FC29FF29" box="[922,1020,187,213]" name="Vietnam" pageId="3" pageNumber="556">Vietnam</collectingCountry>
,
<collectingRegion id="300538C0311C1134A438FF37FBB6FF29" box="[1031,1123,187,213]" country="Vietnam" name="Lao Cai" pageId="3" pageNumber="556">Lao Cai</collectingRegion>
,
<location id="F71EA0F9311C1134A450FF37FAF1FF29" LSID="urn:lsid:plazi:treatment:7A684734311E1134A0A8F89FFAA5FE75:F71EA0F9311C1134A450FF37FAF1FF29" box="[1135,1316,187,213]" country="Vietnam" name="Muong Khuong" pageId="3" pageNumber="556" stateProvince="Lao Cai">Muong Khuong</location>
,
<date id="867FD0E2311C1134A510FF37FF07FF05" pageId="3" pageNumber="556" value="2016-08-09" valueMax="2016-08-11" valueMin="2016-08-09">
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911 Aug
<quantity id="35395BC7311C1134A0A8FF53FF07FF05" box="[151,210,223,249]" metricMagnitude="3" metricUnit="m" metricValue="2.0169249999999996" pageId="3" pageNumber="556" unit="m" value="2016.925">2016</quantity>
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</date>
, 925m,
<collectorName id="5F3493F4311C1134A116FF53FE69FF05" box="[297,444,223,249]" pageId="3" pageNumber="556">J.B. Heppner</collectorName>
, adult no. FLMNH-MGCL1174136 and genitalia slide no.
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</materialsCitation>
˗12965;
<materialsCitation id="42A9FC7F311C1134A4F3FF53FF6BFEBD" collectingDate="2016-08-22" collectingDateMax="2016-08-27" collectingDateMin="2016-08-22" collectionCode="UB" collectorName="J. B. Heppner" country="Vietnam" location="Tam Dao" pageId="3" pageNumber="556" specimenCount="1" specimenCount-female="1" stateProvince="Vinh Phuc">
<specimenCount id="E4C73DAB311C1134A4F3FF53FB3AFF05" box="[1228,1263,223,249]" count="1" pageId="3" pageNumber="556" type="female">1♀</specimenCount>
,
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,
<collectingRegion id="300538C0311C1134A559FF53FF07FEE1" country="Vietnam" name="Vinh Phuc" pageId="3" pageNumber="556">Vinh Phuc</collectingRegion>
,
<location id="F71EA0F9311C1134A0E1FE8FFE9FFEE1" LSID="urn:lsid:plazi:treatment:7A684734311E1134A0A8F89FFAA5FE75:F71EA0F9311C1134A0E1FE8FFE9FFEE1" box="[222,330,259,285]" country="Vietnam" name="Tam Dao" pageId="3" pageNumber="556" stateProvince="Vinh Phuc">Tam Dao</location>
,
<date id="867FD0E2311C1134A168FE8FFDCDFEE1" box="[343,536,259,285]" pageId="3" pageNumber="556" value="2016-08-22" valueMax="2016-08-27" valueMin="2016-08-22">
<collectingDate id="963B290A311C1134A168FE8FFDCDFEE1" box="[343,536,259,285]" pageId="3" pageNumber="556" value="2016-08-22" valueMax="2016-08-27" valueMin="2016-08-22">
2227 Aug
<quantity id="35395BC7311C1134A1E2FE8FFDCDFEE1" box="[477,536,259,285]" metricMagnitude="3" metricUnit="m" metricValue="2.01693" pageId="3" pageNumber="556" unit="m" value="2016.93">2016</quantity>
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, 930m,
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, adult no. FLMNHMGCL1174129 and genitalia slide no.
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˗12960 (deposited in MGCL).
</paragraph>
</subSubSection>
<subSubSection id="BADBA5A9311C1134A0F8FEC7FAA5FE75" pageId="3" pageNumber="556" type="discussion">
<paragraph id="F27EF622311C1134A0F8FEC7FAA5FE75" blockId="3.[151,1437,150,393]" pageId="3" pageNumber="556">
<emphasis id="C0B52A30311C1134A0F8FEC7FEEEFE99" bold="true" box="[199,315,331,357]" pageId="3" pageNumber="556">Remarks.</emphasis>
This species was reported from
<collectingCountry id="8AD6B6B2311C1134A2E6FEC7FC97FE99" box="[729,834,331,357]" name="Vietnam" pageId="3" pageNumber="556">Vietnam</collectingCountry>
(
<collectingRegion id="300538C0311C1134A368FEC7FC35FE99" box="[855,992,331,357]" country="Vietnam" name="Vinh Phuc" pageId="3" pageNumber="556">Vinh Phuc</collectingRegion>
and
<collectingCountry id="8AD6B6B2311C1134A419FEC7FBA5FE99" box="[1062,1136,331,357]" name="Turkey" pageId="3" pageNumber="556">Hatay</collectingCountry>
(
<collectingRegion id="300538C0311C1134A4B9FEC7FB03FE99" box="[1158,1238,331,357]" country="Vietnam" name="Ha Noi" pageId="3" pageNumber="556">Hanoi</collectingRegion>
) Provinces) by
<bibRefCitation id="96508BD3311C1134A0A8FEE3FDC5FE75" author="Bayarsaikhan, U. &amp; Bae, Y. S." box="[151,528,367,393]" pageId="3" pageNumber="556" pagination="365 - 376" refId="ref4259" refString="Bayarsaikhan, U. &amp; Bae, Y. S. (2019) Four new and one newly recorded species of Diduga Moore, [1887] (Lepidoptera, Erebidae, Arctiinae) from Vietnam, with redescription of the little known species Diduga haematomiformis van Eecke, 1920. Zootaxa, 4624 (3), 365 - 376. https: // doi. org / 10.11646 / zootaxa. 4624.3.5" type="journal article" year="2019">Bayarsaikhan and Bae (2019)</bibRefCitation>
. In this study, we report it for
<collectingRegion id="300538C0311C1134A35CFEE3FBFDFE75" box="[867,1064,367,393]" country="Vietnam" name="Lao Cai" pageId="3" pageNumber="556">Lao Cai Province</collectingRegion>
of
<collectingCountry id="8AD6B6B2311C1134A473FEE3FB7EFE75" box="[1100,1195,367,393]" name="Vietnam" pageId="3" pageNumber="556">Vietnam</collectingCountry>
for the first time.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,606 @@
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<mods:title id="CEBC9E7C0D35163F1D4C1189D7A97ED1">An endemic species flock of Candonidae Kaufmann, 1900 (Crustacea, Ostracoda) from wells in Benin (Africa), with the description of a new subfamily, a new genus and five new species</mods:title>
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<emphasis id="40C26A3E4507FF8EFF411E84FE5928D0" bold="true" box="[151,471,1482,1509]" italics="true" pageId="50" pageNumber="51">Benincandona trapezoides</emphasis>
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Hotèkpo &amp; Martens
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(
<figureCitation id="EA8DAAA94507FF8EFF491D5DFECA2B18" box="[159,324,1555,1581]" captionStart-0="FIGURE 32" captionStart-1="FIGURE 33" captionStart-2="FIGURE 34" captionStart-3="FIGURE 35" captionStart-4="FIGURE 36" captionStart-5="FIGURE 37" captionStart-6="FIGURE 38" captionStart-7="FIGURE 39" captionStartId-0="51.[151,250,1943,1967]" captionStartId-1="52.[151,250,1520,1544]" captionStartId-2="54.[151,250,1520,1544]" captionStartId-3="55.[151,250,1943,1967]" captionStartId-4="56.[151,250,1943,1967]" captionStartId-5="57.[151,250,1981,2005]" captionStartId-6="58.[151,250,1943,1967]" captionStartId-7="59.[151,250,1960,1984]" captionTargetBox-0="[179,1408,181,1920]" captionTargetBox-1="[151,1436,181,1496]" captionTargetBox-2="[151,1436,181,1496]" captionTargetBox-3="[186,1401,185,1915]" captionTargetBox-4="[166,1418,185,1915]" captionTargetBox-5="[223,1388,222,1895]" captionTargetBox-6="[189,1403,188,1912]" captionTargetBox-7="[175,1409,202,1932]" captionTargetId-0="figure-17@51.[179,1408,181,1920]" captionTargetId-1="figure-31@52.[151,1436,181,1496]" captionTargetId-2="figure-31@54.[151,1436,181,1496]" captionTargetId-3="figure-17@55.[182,1405,181,1919]" captionTargetId-4="figure-31@56.[166,1422,181,1919]" captionTargetId-5="figure-17@57.[166,1422,181,1957]" captionTargetId-6="figure-31@58.[179,1408,181,1919]" captionTargetId-7="figure-17@59.[175,1413,198,1936]" captionTargetPageId-0="51" captionTargetPageId-1="52" captionTargetPageId-2="54" captionTargetPageId-3="55" captionTargetPageId-4="56" captionTargetPageId-5="57" captionTargetPageId-6="58" captionTargetPageId-7="59" captionText-0="FIGURE 32. Benincandona trapezoides gen. et spec. nov., valves (AC, GK ♂; DF ♀). AF, H. CpRl (AC, H. INV.159276, DF. INV.1592 INV.159279). G, I, J. INV.159279. CpV. K. INV.159277. CpD. Scale bars. 1 (B, E, G, K) = 500 µm; 2 (A, C, D, F, I) = 200 µm, 3 (H, J) = 40 µm." captionText-1="FIGURE 33. Benincandona trapezoides gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159263). B, E, F, H. RVi (INV.159263). G, H. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (GH) = 50 µm." captionText-2="FIGURE 34. Benincandona trapezoides gen. et spec. nov., valves (♀). A, C, D, G. LVi (INV.159264). B, E, F, H. RVi (INV.159264). G, H. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (GH) = 50 µm." captionText-3="FIGURE 35. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. A2 (INV.159280). B. A1 (INV.159265). C. MdCoxa (INV.159263). Scale bar = 50 µm." captionText-4="FIGURE 36. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Mx1, respiratory plate (INV.159263). B. Mx1 (Chaetotaxy incomplete) (INV.159263). C. MdPalp (INV.159263). Scale bar. 1 (A), 2 (C), 3 (B) = 50 µm." captionText-5="FIGURE 37. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. T2 (INV.159263). B. T3 (INV.159263). C. CR (INV.159280). D. CRAtt (INV.159280). Scale bar = 50 µm." captionText-6="FIGURE 38. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159263). BD. Hp (INV.159263). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." captionText-7="FIGURE 39. Benincandona trapezoides gen. et spec. nov., limbs (AB ♀ and CE ♂). A. T1, Lpp (INV.159280). B. T1, Rpp (INV.159280). C. A2 (INV.159264, INV.159269). D. T1 (INV.159268). E. CR (INV.159264). Scale bar = 50 µm." pageId="50" pageNumber="51">
Figures 32
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,
<figureCitation id="EA8DAAA94507FF8EFE861D5DFE2C2B18" box="[336,418,1555,1581]" captionStart="FIGURE 41" captionStartId="61.[151,250,1113,1137]" captionTargetBox="[151,1436,181,1089]" captionTargetId="figure-17@61.[151,1436,181,1089]" captionTargetPageId="61" captionText="FIGURE 41. Benincandona gen. et spec. nov., CpRl. (A, C, E, F, G, I, K, N, O, Q ♂ and B, D, H, J, L, M, P, R ♀). AD. B. martini. EF. B. ibikounlei. GJ. B. sakitii. KN. B. cebios. OR. B. trapezoides. Scale bar = 500 µm." pageId="50" pageNumber="51">41OR</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74507FF8EFF411D15FB332BAC" box="[151,1213,1627,1689]" pageId="50" pageNumber="51" type="etymology">
<paragraph id="7209B62C4507FF8EFF411D15FE962B40" blockId="50.[151,1213,1627,1689]" box="[151,280,1627,1653]" pageId="50" pageNumber="51">
<emphasis id="40C26A3E4507FF8EFF411D15FE962B40" bold="true" box="[151,280,1627,1653]" pageId="50" pageNumber="51">Etymology</emphasis>
</paragraph>
<paragraph id="7209B62C4507FF8EFF411D31FB332BAC" blockId="50.[151,1213,1627,1689]" box="[151,1213,1663,1689]" pageId="50" pageNumber="51">The species is named after the shape of the carapace in lateral view, which is largely trapezoid.</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74507FF8EFF411D8EFBA12ACF" pageId="50" pageNumber="51" type="diagnosis">
<paragraph id="7209B62C4507FF8EFF411D8EFE842BEF" blockId="50.[151,1437,1728,2043]" box="[151,266,1728,1754]" pageId="50" pageNumber="51">
<emphasis id="40C26A3E4507FF8EFF411D8EFE842BEF" bold="true" box="[151,266,1728,1754]" pageId="50" pageNumber="51">Diagnosis</emphasis>
</paragraph>
<paragraph id="7209B62C4507FF8EFF411DAAFBA12ACF" blockId="50.[151,1437,1728,2043]" pageId="50" pageNumber="51">
CpRl relatively high (L/H = 2.0), dorsal margin straight over
<emphasis id="40C26A3E4507FF8EFCEA1DABFCFB2BCB" box="[828,885,1765,1790]" italics="true" pageId="50" pageNumber="51">circa</emphasis>
one third of the entire length, with greatest height in the middle; LV dorsally extending beyond RV over the entire length of the straight part of the dorsal margin, anterior margin broadly rounded, posterior margin running straight towards the ventral side, the latter almost straight, valve surface set with ridges in most of the central part. CpD and CpV relatively wide, with greatest width in the middle and lateral sides straight in the middle, anterior extremity sharply and posterior side more bluntly pointed. A2 with seta X long, reaching well beyond the tip of the aesthetasc Y, the latter distally not bifurcated. Second palp segment of Mx1 with L
<emphasis id="40C26A3E4507FF8EFE721CF3FE3E2AE3" box="[420,432,1981,2006]" italics="true" pageId="50" pageNumber="51">c</emphasis>
. 1.5x basal W. T3 with seta d2 absent. Hp with ls and ms with overlapping positions, lobe ls broad and rounded, extending beyond lobe ms with more than half of its length.
</paragraph>
</subSubSection>
<caption id="26C9E6A44506FF8FFF411CD9FD892ACD" pageId="51" pageNumber="52" startId="51.[151,250,1943,1967]" targetBox="[179,1408,181,1920]" targetPageId="51" targetType="figure">
<paragraph id="7209B62C4506FF8FFF411CD9FD892ACD" blockId="51.[151,1436,1943,2040]" pageId="51" pageNumber="52">
<emphasis id="40C26A3E4506FF8FFF411CD9FE912A9A" bold="true" box="[151,287,1943,1967]" pageId="51" pageNumber="52">FIGURE 32.</emphasis>
<taxonomicName id="B5B6CDAF4506FF8FFEF21CD6FDBF2A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[292,561,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="trapezoides" status="gen. et spec. nov.">
<emphasis id="40C26A3E4506FF8FFEF21CD6FDBF2A85" box="[292,561,1944,1968]" italics="true" pageId="51" pageNumber="52">Benincandona trapezoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454506FF8FFDE01CD6FD6A2A9A" box="[566,740,1944,1968]" pageId="51" pageNumber="52" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, valves (AC, GK ♂; DF ♀).
<emphasis id="40C26A3E4506FF8FFBF01CD9FBFC2A85" bold="true" box="[1062,1138,1943,1968]" pageId="51" pageNumber="52">AF, H</emphasis>
. CpRl (AC, H. INV.159276, DF. INV.1592 INV.159279).
<emphasis id="40C26A3E4506FF8FFE1D1CF5FD9E2AE1" bold="true" box="[459,528,1979,2004]" pageId="51" pageNumber="52">G, I, J</emphasis>
. INV.159279. CpV.
<emphasis id="40C26A3E4506FF8FFD371CF2FD7B2AE1" bold="true" box="[737,757,1980,2004]" pageId="51" pageNumber="52">K</emphasis>
. INV.159277. CpD. Scale bars. 1 (B, E, G, K) = 500 µm; 2 (A, C, D, F, I) = 200 µm, 3 (H, J) = 40 µm.
</paragraph>
</caption>
<caption id="26C9E6A44501FF88FF411EBEFBFC2B18" pageId="52" pageNumber="53" startId="52.[151,250,1520,1544]" targetBox="[151,1436,181,1496]" targetPageId="52" targetType="figure">
<paragraph id="7209B62C4501FF88FF411EBEFBFC2B18" blockId="52.[151,1437,1520,1581]" pageId="52" pageNumber="53">
<emphasis id="40C26A3E4501FF88FF411EBEFEAA2B3D" bold="true" box="[151,292,1520,1544]" pageId="52" pageNumber="53">FIGURE 33.</emphasis>
<taxonomicName id="B5B6CDAF4501FF88FEF81EBEFDCE2B3D" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[302,576,1520,1544]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="trapezoides" status="gen. et spec. nov.">
<emphasis id="40C26A3E4501FF88FEF81EBEFDCE2B3D" box="[302,576,1520,1544]" italics="true" pageId="52" pageNumber="53">Benincandona trapezoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454501FF88FD9C1EBFFC862B3D" box="[586,776,1521,1545]" pageId="52" pageNumber="53" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, valves (♂).
<emphasis id="40C26A3E4501FF88FC431EBEFB9C2B3D" bold="true" box="[917,1042,1520,1544]" pageId="52" pageNumber="53">A, C, D, G</emphasis>
. LVi (INV.159263).
<emphasis id="40C26A3E4501FF88FB261EBEFAEB2B3D" bold="true" box="[1264,1381,1520,1544]" pageId="52" pageNumber="53">B, E, F, H</emphasis>
. RVi (INV.159263).
<emphasis id="40C26A3E4501FF88FEE71D5AFEE52B19" bold="true" box="[305,363,1556,1580]" pageId="52" pageNumber="53">G, H.</emphasis>
CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (GH) = 50 µm.
</paragraph>
</caption>
<subSubSection id="3AACE5A74501FF89FF411D30FD7D2C98" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53" type="materials_examined">
<paragraph id="7209B62C4501FF88FF411D30FEB02BAD" blockId="52.[151,318,1662,1688]" box="[151,318,1662,1688]" pageId="52" pageNumber="53">
<emphasis id="40C26A3E4501FF88FF411D30FEB02BAD" bold="true" box="[151,318,1662,1688]" pageId="52" pageNumber="53">Type material</emphasis>
</paragraph>
<paragraph id="7209B62C4501FF88FF411D88FE8A2BD5" blockId="52.[151,1282,1734,1797]" box="[151,260,1734,1760]" pageId="52" pageNumber="53">
<emphasis id="40C26A3E4501FF88FF411D88FE8A2BD5" bold="true" box="[151,260,1734,1760]" pageId="52" pageNumber="53">
<typeStatus id="AD0D088E4501FF88FF411D88FE8A2BD5" box="[151,260,1734,1760]" pageId="52" pageNumber="53" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7209B62C4501FF88FF411DA4FA8C2A30" blockId="52.[151,1282,1734,1797]" box="[151,1282,1770,1797]" pageId="52" pageNumber="53">
<materialsCitation id="C2DEBC714501FF88FF411DA4FA8C2A30" box="[151,1282,1770,1797]" collectionCode="INV" pageId="52" pageNumber="53" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="64B07DA54501FF88FF411DA4FF4D2A30" box="[151,195,1770,1797]" count="1" pageId="52" pageNumber="53" type="male">1 ♂</specimenCount>
, with soft parts dissected in glycerine in a sealed slide, and with valves stored dry (
<collectionCode id="14A72EE94501FF88FBB01DA4FB182A31" box="[1126,1174,1770,1796]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="52" pageNumber="53" type="Herbarium">INV</collectionCode>
.159263).
</materialsCitation>
</paragraph>
<paragraph id="7209B62C4501FF88FF411C7CFF722A79" blockId="52.[151,736,1842,1905]" box="[151,252,1842,1868]" pageId="52" pageNumber="53">
<emphasis id="40C26A3E4501FF88FF411C7CFF722A79" bold="true" box="[151,252,1842,1868]" pageId="52" pageNumber="53">
<typeStatus id="AD0D088E4501FF88FF411C7CFF722A79" box="[151,252,1842,1868]" pageId="52" pageNumber="53" type="allotype">Allotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7209B62C4501FF88FF411C18FD6E2A44" blockId="52.[151,736,1842,1905]" box="[151,736,1878,1905]" pageId="52" pageNumber="53">
<materialsCitation id="C2DEBC714501FF88FF411C18FD6E2A44" box="[151,736,1878,1905]" collectionCode="INV" pageId="52" pageNumber="53" specimenCount="1" specimenCount-female="1" typeStatus="allotype">
<specimenCount id="64B07DA54501FF88FF411C18FF4F2A44" box="[151,193,1878,1905]" count="1" pageId="52" pageNumber="53" type="female">1 ♀</specimenCount>
, dissected and stored as the male (
<collectionCode id="14A72EE94501FF88FD921C18FDF72A44" box="[580,633,1878,1905]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="52" pageNumber="53" type="Herbarium">INV</collectionCode>
.159264).
</materialsCitation>
</paragraph>
<paragraph id="7209B62C4501FF88FF411CD0FE812A8D" blockId="52.[151,1436,1950,2013]" box="[151,271,1950,1976]" pageId="52" pageNumber="53">
<emphasis id="40C26A3E4501FF88FF411CD0FE812A8D" bold="true" box="[151,271,1950,1976]" pageId="52" pageNumber="53">
<typeStatus id="AD0D088E4501FF88FF411CD0FE812A8D" box="[151,271,1950,1976]" pageId="52" pageNumber="53" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7209B62C4501FF89FF411C8CFC282DCC" blockId="52.[151,1436,1950,2013]" lastBlockId="53.[151,1436,150,249]" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">
<materialsCitation id="C2DEBC714501FF88FF411C8CFC442AE8" box="[151,970,1986,2013]" collectionCode="INV" pageId="52" pageNumber="53" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="64B07DA54501FF88FF411C8CFF572AE8" box="[151,217,1986,2013]" count="2" pageId="52" pageNumber="53" type="male">2 ♂♂</specimenCount>
dissected and stored as the
<typeStatus id="AD0D088E4501FF88FDF21C8CFD082AE9" box="[548,646,1986,2012]" pageId="52" pageNumber="53" type="holotype">holotype</typeStatus>
(
<collectionCode id="14A72EE94501FF88FD4E1C8CFD422AE8" box="[664,716,1986,2013]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="52" pageNumber="53" type="Herbarium">INV</collectionCode>
.159265, INV.159266)
</materialsCitation>
;
<materialsCitation id="C2DEBC714501FF89FC0F1C8CFEA32DE0" collectionCode="INV" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53" specimenCount="9" specimenCount-female="9" typeStatus="paratype">
<specimenCount id="64B07DA54501FF88FC0F1C8CFB952AE8" box="[985,1051,1986,2013]" count="9" pageId="52" pageNumber="53" type="female">9 ♀♀</specimenCount>
dissected and stored as the male (
<collectionCode id="14A72EE94500FF89FF491BD8FF5D2D84" box="[159,211,150,177]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="53" pageNumber="54" type="Herbarium">INV</collectionCode>
.159267, INV.159268, INV.159269, INV.159270, INV.159271, INV.159272, INV.159273, INV.159274, INV.159275)
</materialsCitation>
;
<materialsCitation id="C2DEBC714500FF89FEEE1BF5FA162DE0" box="[312,1432,186,213]" collectionCode="INV" pageId="53" pageNumber="54" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="64B07DA54500FF89FEEE1BF5FEF92DE0" box="[312,375,187,213]" count="3" pageId="53" pageNumber="54" type="male">3 ♂♂</specimenCount>
carapaces stored dry in micropalaeontological slides (
<collectionCode id="14A72EE94500FF89FC051BF4FB892DE0" box="[979,1031,186,213]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="53" pageNumber="54" type="Herbarium">INV</collectionCode>
.159276, INV.159277, INV.159278)
</materialsCitation>
;
<materialsCitation id="C2DEBC714500FF89FF411B91FC2C2DCC" box="[151,930,222,249]" collectionCode="INV" pageId="53" pageNumber="54" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="64B07DA54500FF89FF411B91FF4F2DCC" box="[151,193,223,249]" count="1" pageId="53" pageNumber="54" type="female">1 ♀</specimenCount>
carapace stored dry in micropalaeontological slide (
<collectionCode id="14A72EE94500FF89FCDD1B90FCCE2DCC" box="[779,832,222,249]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="53" pageNumber="54" type="Herbarium">INV</collectionCode>
.159279)
</materialsCitation>
.
</paragraph>
<paragraph id="7209B62C4500FF89FF411A69FEBE2C75" blockId="53.[151,1436,294,429]" box="[151,304,294,321]" pageId="53" pageNumber="54">
<emphasis id="40C26A3E4500FF89FF411A69FEBE2C75" bold="true" box="[151,304,294,321]" pageId="53" pageNumber="54">Type locality</emphasis>
</paragraph>
<paragraph id="7209B62C4500FF89FF411A05FD7D2C98" blockId="53.[151,1436,294,429]" pageId="53" pageNumber="54">
<materialsCitation id="C2DEBC714500FF89FF411A05FE2D2CBC" country="Benin" location="Parakou District" pageId="53" pageNumber="54" specimenCount="1" stateProvince="Borgou">
<collectingCountry id="0AA1F6BC4500FF89FF411A05FF542C50" box="[151,218,331,357]" name="Benin" pageId="53" pageNumber="54">Benin</collectingCountry>
,
<collectingRegion id="B07278CE4500FF89FF3E1A05FE442C50" box="[232,458,331,357]" country="Benin" name="Borgou" pageId="53" pageNumber="54">Borgou Department</collectingRegion>
,
<location id="7769E0F74500FF89FE001A05FD1E2C50" LSID="urn:lsid:plazi:treatment:FA1F073A4507FF82FF411E84FB1329F0:7769E0F74500FF89FE001A05FD1E2C50" box="[470,656,331,357]" country="Benin" name="Parakou District" pageId="53" pageNumber="54" stateProvince="Borgou">Parakou District</location>
, town Bapérou located in the Okpara catchment area (tributary of the
<collectingRegion id="B07278CE4500FF89FF411A21FF662CBC" box="[151,232,367,393]" country="Benin" name="Oueme" pageId="53" pageNumber="54">Ouémé</collectingRegion>
catchment area)
</materialsCitation>
, coordinates:
<geoCoordinate id="1782D0EB4500FF89FDE81A21FD4A2CBC" box="[574,708,367,393]" degrees="9" direction="north" minutes="16.380" orientation="latitude" pageId="53" pageNumber="54" precision="1" value="9.273">9°16.380N</geoCoordinate>
,
<geoCoordinate id="1782D0EB4500FF89FD191A21FCD82CBC" box="[719,854,367,393]" degrees="2" direction="east" minutes="34.913" orientation="longitude" pageId="53" pageNumber="54" precision="1" value="2.5818834">2°34.913 E</geoCoordinate>
, Leg.: S.J. Hotèkpo and M. Lagnika collected on
<date id="060890EC4500FF89FA561A20FEC22C98" pageId="53" pageNumber="54" value="2022-11-05">05 November 2022</date>
(Locality code A
<quantity id="B54E1BC94500FF89FDC21ADDFDB52C98" box="[532,571,403,429]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="53" pageNumber="54" unit="in" value="1.0">1 in</quantity>
<tableCitation id="3F3483974500FF89FD971ADDFD102C98" box="[577,670,403,429]" captionStart="TABLE 1" captionStartId="5.[152,244,929,955]" captionText="TABLE 1. Localities where Benincandona gen. nov. sp. nov. were collected, coordinates and hydro-morphological characteristics. In all stations, the water was clear and was used for both drinking water and household purposes by the riparian villages." pageId="53" pageNumber="54">Tables 1</tableCitation>
and
<tableCitation id="3F3483974500FF89FD031ADDFD682C98" box="[725,742,403,429]" captionStart="TABLE 2" captionStartId="6.[152,244,1634,1658]" captionText="TABLE 2. Measurements of physical and chemical water parameters and variables at the time of well fauna collection. WT= water temperature (°C), EC = electrical conductivity (µS cm-1), DO = dissolved oxygen (mg L-1). NM= no measurement." pageId="53" pageNumber="54">2</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74500FF82FF411A94FD9E2FF8" lastPageId="62" lastPageNumber="63" pageId="53" pageNumber="54" type="description">
<paragraph id="7209B62C4500FF89FF411A94FE0F2CC0" blockId="53.[151,1437,474,1905]" box="[151,385,474,501]" pageId="53" pageNumber="54">
<emphasis id="40C26A3E4500FF89FF411A94FE0F2CC0" bold="true" box="[151,385,474,501]" pageId="53" pageNumber="54">Description of male</emphasis>
</paragraph>
<paragraph id="7209B62C4500FF89FF411AB1FCD82FF8" blockId="53.[151,1437,474,1905]" pageId="53" pageNumber="54">
CpRl (
<figureCitation id="EA8DAAA94500FF89FF361AB1FE0D2F2C" box="[224,387,511,537]" captionStart="FIGURE 32" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure-17@51.[179,1408,181,1920]" captionTargetPageId="51" captionText="FIGURE 32. Benincandona trapezoides gen. et spec. nov., valves (AC, GK ♂; DF ♀). AF, H. CpRl (AC, H. INV.159276, DF. INV.1592 INV.159279). G, I, J. INV.159279. CpV. K. INV.159277. CpD. Scale bars. 1 (B, E, G, K) = 500 µm; 2 (A, C, D, F, I) = 200 µm, 3 (H, J) = 40 µm." pageId="53" pageNumber="54">Figure 32AC</figureCitation>
) with trapezoidal shape (L/H ratio=2.0); dorsal margin straight in the middle 1/3, with greatest height in the middle, anterior margin broadly rounded, posterior margin running straight towards ventral margin, the latter almost straight; LV overlapping RV along anterior, ventral and posterior sides, dorsally strongly overlapping RV along the entire length of the straight part; valve surface set with ridges in the middle part of the valves (
<figureCitation id="EA8DAAA94500FF89FA871925FF6B2F9C" captionStart="FIGURE 32" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure-17@51.[179,1408,181,1920]" captionTargetPageId="51" captionText="FIGURE 32. Benincandona trapezoides gen. et spec. nov., valves (AC, GK ♂; DF ♀). AF, H. CpRl (AC, H. INV.159276, DF. INV.1592 INV.159279). G, I, J. INV.159279. CpV. K. INV.159277. CpD. Scale bars. 1 (B, E, G, K) = 500 µm; 2 (A, C, D, F, I) = 200 µm, 3 (H, J) = 40 µm." pageId="53" pageNumber="54">Figure 32HJ</figureCitation>
). CpD and CpV (
<figureCitation id="EA8DAAA94500FF89FE7019C1FDC32F9C" box="[422,589,655,681]" captionStart="FIGURE 32" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure-17@51.[179,1408,181,1920]" captionTargetPageId="51" captionText="FIGURE 32. Benincandona trapezoides gen. et spec. nov., valves (AC, GK ♂; DF ♀). AF, H. CpRl (AC, H. INV.159276, DF. INV.1592 INV.159279). G, I, J. INV.159279. CpV. K. INV.159277. CpD. Scale bars. 1 (B, E, G, K) = 500 µm; 2 (A, C, D, F, I) = 200 µm, 3 (H, J) = 40 µm." pageId="53" pageNumber="54">Figure 32G, K</figureCitation>
) relatively wide, pointed sharply anteriorly and bluntly posteriorly, greatest width in the middle, lateral margins straight in the middle part.
</paragraph>
<paragraph id="7209B62C4500FF89FF111999FB412E0C" blockId="53.[151,1437,474,1905]" pageId="53" pageNumber="54">
LVi (
<figureCitation id="EA8DAAA94500FF89FF281999FE6F2FC4" box="[254,481,727,753]" captionStart="FIGURE 33" captionStartId="52.[151,250,1520,1544]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-31@52.[151,1436,181,1496]" captionTargetPageId="52" captionText="FIGURE 33. Benincandona trapezoides gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159263). B, E, F, H. RVi (INV.159263). G, H. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (GH) = 50 µm." pageId="53" pageNumber="54">Figure 33A, C, D, G</figureCitation>
) elongated and sub-triangular, dorsal margin straight in the middle third, with greatest height in the middle, anterior margin broadly rounded; posterior margin sloping towards ventral margin, the latter almost straight; ventral il without antero-ventral tooth. CMS (
<figureCitation id="EA8DAAA94500FF89FC9F1851FC452E0C" box="[841,971,799,825]" captionStart="FIGURE 33" captionStartId="52.[151,250,1520,1544]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-31@52.[151,1436,181,1496]" captionTargetPageId="52" captionText="FIGURE 33. Benincandona trapezoides gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159263). B, E, F, H. RVi (INV.159263). G, H. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (GH) = 50 µm." pageId="53" pageNumber="54">Figure 33G</figureCitation>
) as in the
<typeStatus id="AD0D088E4500FF89FB961851FBFE2E0C" box="[1088,1136,799,825]" pageId="53" pageNumber="54">type</typeStatus>
species.
</paragraph>
<paragraph id="7209B62C4500FF89FF11180DFBF02E90" blockId="53.[151,1437,474,1905]" pageId="53" pageNumber="54">
RVi (
<figureCitation id="EA8DAAA94500FF89FED4180DFE682E68" box="[258,486,835,861]" captionStart="FIGURE 33" captionStartId="52.[151,250,1520,1544]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-31@52.[151,1436,181,1496]" captionTargetPageId="52" captionText="FIGURE 33. Benincandona trapezoides gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159263). B, E, F, H. RVi (INV.159263). G, H. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (GH) = 50 µm." pageId="53" pageNumber="54">Figure 33B, E, F, H</figureCitation>
) elongated, with dorsal margin straight in the middle third and with greatest height in the middle, anterior margin broadly rounded; posterior margin sloping towards ventral margin, the latter almost straight; ventral il without antero-ventral tooth. CMS (
<figureCitation id="EA8DAAA94500FF89FD2118C5FCF42E90" box="[759,890,907,933]" captionStart="FIGURE 33" captionStartId="52.[151,250,1520,1544]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-31@52.[151,1436,181,1496]" captionTargetPageId="52" captionText="FIGURE 33. Benincandona trapezoides gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159263). B, E, F, H. RVi (INV.159263). G, H. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (GH) = 50 µm." pageId="53" pageNumber="54">Figure 33H</figureCitation>
) as in the
<typeStatus id="AD0D088E4500FF89FC3818C5FB902E90" box="[1006,1054,907,933]" pageId="53" pageNumber="54">type</typeStatus>
species.
</paragraph>
<paragraph id="7209B62C4500FF89FF1118E1FB0A2838" blockId="53.[151,1437,474,1905]" pageId="53" pageNumber="54">
A1 (
<figureCitation id="EA8DAAA94500FF89FF2D18E1FEF12EFC" box="[251,383,942,969]" captionStart="FIGURE 35" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[186,1401,185,1915]" captionTargetId="figure-17@55.[182,1405,181,1919]" captionTargetPageId="55" captionText="FIGURE 35. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. A2 (INV.159280). B. A1 (INV.159265). C. MdCoxa (INV.159263). Scale bar = 50 µm." pageId="53" pageNumber="54">Figure 35B</figureCitation>
) with seven segments, one basal and six distal. Basal segment with two sub-apical, ventral setae, one long and one shorter, the latter about half of the length of the longer one, and two shorter setae on the mid-dorsal side (one not illustrated here), situated some distance from each other. First distal segment about 2.5 times as long as the basal width, with one short dorso-subapical seta. Second distal segment with about the same length/width ratio as the previous segment, set with one short dorso-apical seta. Third distal segment also with about the same length/width ratio, carrying one short apical setae on each side. Fourth distal segment about three times as long as wide, and set with one short ventro-apical seta and two long dorso-apical. Fifth distal segment about five time as long as wide, with two long dorso-apical setae and one short ventro-apical seta clearly exceeding the next segment. Sixth distal segment narrow, at least five times as long as wide, with four apical structures: one aesthetasc Ya of intermediate length, two long and one short setae (about one third the length of Ya).
</paragraph>
<paragraph id="7209B62C4500FF89FF111E59FA9A2A1C" blockId="53.[151,1437,474,1905]" pageId="53" pageNumber="54">
A2 (
<figureCitation id="EA8DAAA94500FF89FF2D1E59FE0D2804" box="[251,387,1302,1329]" captionStart="FIGURE 35" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[186,1401,185,1915]" captionTargetId="figure-17@55.[182,1405,181,1919]" captionTargetPageId="55" captionText="FIGURE 35. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. A2 (INV.159280). B. A1 (INV.159265). C. MdCoxa (INV.159263). Scale bar = 50 µm." pageId="53" pageNumber="54">Figure 35A</figureCitation>
) with basal segment (first protopodite) broad and rounded, mid-ventrally with one seta of medium length, subapically with two setae of unequal length, proximal one
<emphasis id="40C26A3E4500FF89FBD41E75FB9B2861" box="[1026,1045,1339,1364]" italics="true" pageId="53" pageNumber="54">c.</emphasis>
four times the length of the more apical one. Second segment (second protopodite) large and
<emphasis id="40C26A3E4500FF89FCFA1E11FCB1284D" box="[812,831,1375,1400]" italics="true" pageId="53" pageNumber="54">c.</emphasis>
twice as long as basal width, set with one long ventro-subapical seta X. No remnant of exopodite visible. Endopodite consisting of four segments (penultimate segment divided). First endopodal segment large and stout, five times as long as basal width, carrying one long and narrow mid-ventral aesthetasc Y, and two ventro-apical setae, one short and one large and stout, hirsute in its distal 4/5. Second endopodal segment (segment 2A)
<emphasis id="40C26A3E4500FF89FDAD1EA1FD002B3D" box="[635,654,1519,1544]" italics="true" pageId="53" pageNumber="54">c.</emphasis>
twice as long as wide, carrying one stout dorso-apical plumose seta and three t-setae; seta t1 long, narrow and plumose, accompanied by a slender aesthetasc y1; t2 and t3 setae hyper-developed into male bristles, t2 ventro-apical, t3 centro-apical. Third endopodal segment (segment 2B) smaller than previous segment and about 1.5x as long as wide, set with a complex (and sexually dimorphic) apical chaetotaxy (
<figureCitation id="EA8DAAA94500FF89FF491D31FE9F2BAC" box="[159,273,1663,1689]" captionStart="FIGURE 40" captionStartId="60.[151,250,1516,1540]" captionTargetBox="[151,1435,181,1492]" captionTargetId="figure-31@60.[151,1435,181,1492]" captionTargetPageId="60" captionText="FIGURE 40. Diagrams of apical views of the A2 of Benincandona, showing position of (sub-) terminal claws, setae and aesthetascs of the terminal (small circle) and penultimate segment (large circle). For explanations of codes, see Material and Methods (Abbreviations used in text and figures)." pageId="53" pageNumber="54">Figure 40</figureCitation>
): ventro-apically with one slender aesthetasc y2 and claw G3 transformed into one short seta about equal length as y2; centrally with one short plumose seta z3; dorso-apically, z1 claw-like about 1.5x as long as penultimate segment. Claw G2 and G1 about 2.5 times as long as z1; seta z2 transformed into long claw, reaching tip of G1. Fourth, terminal, endopodal segment with one long claw (Gm), one short claw (GM), a long aesthetasc y3 with shorter accompanying seta, with fused part short, and a seta g (slightly exceeding tip of aesthetasc y3).
</paragraph>
<paragraph id="7209B62C4500FF89FF111C7DFAA52A44" blockId="53.[151,1437,474,1905]" pageId="53" pageNumber="54">
MdCoxa (
<figureCitation id="EA8DAAA94500FF89FEED1C7DFE4E2A78" box="[315,448,1842,1869]" captionStart="FIGURE 35" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[186,1401,185,1915]" captionTargetId="figure-17@55.[182,1405,181,1919]" captionTargetPageId="55" captionText="FIGURE 35. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. A2 (INV.159280). B. A1 (INV.159265). C. MdCoxa (INV.159263). Scale bar = 50 µm." pageId="53" pageNumber="54">Figure 35C</figureCitation>
) elongated, proximally sharply pointed, medially widened, distally with an obliquely widened end, set with a series (67) of strong teeth, interspaced with setae; subapically with a short seta.
</paragraph>
<caption id="26C9E6A44503FF8AFF411EBEFBFC2B18" pageId="54" pageNumber="55" startId="54.[151,250,1520,1544]" targetBox="[151,1436,181,1496]" targetPageId="54" targetType="figure">
<paragraph id="7209B62C4503FF8AFF411EBEFBFC2B18" blockId="54.[151,1437,1520,1581]" pageId="54" pageNumber="55">
<emphasis id="40C26A3E4503FF8AFF411EBEFEAA2B3D" bold="true" box="[151,292,1520,1544]" pageId="54" pageNumber="55">FIGURE 34.</emphasis>
<taxonomicName id="B5B6CDAF4503FF8AFEF81EBEFDCE2B3D" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[302,576,1520,1544]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="trapezoides" status="gen. et spec. nov.">
<emphasis id="40C26A3E4503FF8AFEF81EBEFDCE2B3D" box="[302,576,1520,1544]" italics="true" pageId="54" pageNumber="55">Benincandona trapezoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454503FF8AFD9C1EBFFC862B3D" box="[586,776,1521,1545]" pageId="54" pageNumber="55" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, valves (♀).
<emphasis id="40C26A3E4503FF8AFC431EBEFB9C2B3D" bold="true" box="[917,1042,1520,1544]" pageId="54" pageNumber="55">A, C, D, G</emphasis>
. LVi (INV.159264).
<emphasis id="40C26A3E4503FF8AFB261EBEFAEB2B3D" bold="true" box="[1264,1381,1520,1544]" pageId="54" pageNumber="55">B, E, F, H</emphasis>
. RVi (INV.159264).
<emphasis id="40C26A3E4503FF8AFEE71D5AFEE52B19" bold="true" box="[305,363,1556,1580]" pageId="54" pageNumber="55">G, H.</emphasis>
CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (GH) = 50 µm.
</paragraph>
</caption>
<paragraph id="7209B62C4503FF8AFF111D30FEF82A8D" blockId="54.[151,1437,1662,1976]" pageId="54" pageNumber="55">
MdPalp (
<figureCitation id="EA8DAAA94503FF8AFEF91D30FE3C2BAD" box="[303,434,1662,1688]" captionStart="FIGURE 36" captionStartId="56.[151,250,1943,1967]" captionTargetBox="[166,1418,185,1915]" captionTargetId="figure-31@56.[166,1422,181,1919]" captionTargetPageId="56" captionText="FIGURE 36. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Mx1, respiratory plate (INV.159263). B. Mx1 (Chaetotaxy incomplete) (INV.159263). C. MdPalp (INV.159263). Scale bar. 1 (A), 2 (C), 3 (B) = 50 µm." pageId="54" pageNumber="55">Figure 36C</figureCitation>
) with four segments. Basal segment ventro-subapically carrying two hirsute setae (S1 &amp; S2), S1 long, reaching tip of terminal segment and S2 short and stout, length about one fourth of the length of S1; in between with the short flagellum-like alpha seta (thin and as short as S2), and one smooth and medium size (slightly shorter than S1) more proximal seta. Second segment of sub-triangular shape, dorso-apically with two setae of unequal length; ventro-apically with a group of four long setae, one of these being the beta-seta. Third segment with two unequal dorso-subapical setae, two mid-dorsal setae, one of which being the gamma seta, and ventro-apically with two setae of sub-equal length. Terminal segment sub-rectangular, apically set with one large claw, fused with the segment over its entire distal margin, accompanied by one smooth lateral seta on each side, tips slightly shorter than the fused claw.
</paragraph>
<caption id="26C9E6A44502FF8BFF411CD9FE7C2AE1" pageId="55" pageNumber="56" startId="55.[151,250,1943,1967]" targetBox="[186,1401,185,1915]" targetPageId="55" targetType="figure">
<paragraph id="7209B62C4502FF8BFF411CD9FE7C2AE1" blockId="55.[151,1437,1943,2004]" pageId="55" pageNumber="56">
<emphasis id="40C26A3E4502FF8BFF411CD9FEAE2A9A" bold="true" box="[151,288,1943,1967]" pageId="55" pageNumber="56">FIGURE 35.</emphasis>
<taxonomicName id="B5B6CDAF4502FF8BFEF11CD6FDBB2A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[295,565,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="trapezoides" status="gen. et spec. nov.">
<emphasis id="40C26A3E4502FF8BFEF11CD6FDBB2A85" box="[295,565,1944,1968]" italics="true" pageId="55" pageNumber="56">Benincandona trapezoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454502FF8BFDEA1CD6FD612A9A" box="[572,751,1944,1968]" pageId="55" pageNumber="56" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E4502FF8BFCBF1CD9FCF52A9A" bold="true" box="[873,891,1943,1967]" pageId="55" pageNumber="56">A</emphasis>
. A2 (INV.159280).
<emphasis id="40C26A3E4502FF8BFB911CD6FBD62A85" bold="true" box="[1095,1112,1944,1968]" pageId="55" pageNumber="56">B</emphasis>
. A1 (INV.159265).
<emphasis id="40C26A3E4502FF8BFAF21CD9FAB32A9A" bold="true" box="[1316,1341,1943,1967]" pageId="55" pageNumber="56">C.</emphasis>
MdCoxa (INV.159263). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A4450DFF84FF411CD9FB492AE1" pageId="56" pageNumber="57" startId="56.[151,250,1943,1967]" targetBox="[166,1418,185,1915]" targetPageId="56" targetType="figure">
<paragraph id="7209B62C450DFF84FF411CD9FB492AE1" blockId="56.[151,1436,1943,2004]" pageId="56" pageNumber="57">
<emphasis id="40C26A3E450DFF84FF411CD9FEAB2A9A" bold="true" box="[151,293,1943,1967]" pageId="56" pageNumber="57">FIGURE 36.</emphasis>
<taxonomicName id="B5B6CDAF450DFF84FEF91CD6FDCC2A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[303,578,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="trapezoides" status="gen. et spec. nov.">
<emphasis id="40C26A3E450DFF84FEF91CD6FDCC2A85" box="[303,578,1944,1968]" italics="true" pageId="56" pageNumber="57">Benincandona trapezoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745450DFF84FD9A1CD6FC822A9A" box="[588,780,1944,1968]" pageId="56" pageNumber="57" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E450DFF84FC441CD9FC2A2A9A" bold="true" box="[914,932,1943,1967]" pageId="56" pageNumber="57">A</emphasis>
. Mx1, respiratory plate (INV.159263).
<emphasis id="40C26A3E450DFF84FA9C1CD6FAD52A85" bold="true" box="[1354,1371,1944,1968]" pageId="56" pageNumber="57">B</emphasis>
. Mx1 (Chaetotaxy incomplete) (INV.159263).
<emphasis id="40C26A3E450DFF84FDE41CF5FDCA2AE6" bold="true" box="[562,580,1979,2003]" pageId="56" pageNumber="57">C</emphasis>
. MdPalp (INV.159263). Scale bar. 1 (A), 2 (C), 3 (B) = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A4450CFF85FF411CF3FD7D2ACF" pageId="57" pageNumber="58" startId="57.[151,250,1981,2005]" targetBox="[223,1388,222,1895]" targetPageId="57" targetType="figure">
<paragraph id="7209B62C450CFF85FF411CF3FD7D2ACF" blockId="57.[151,1436,1981,2042]" pageId="57" pageNumber="58">
<emphasis id="40C26A3E450CFF85FF411CF3FEAA2AE0" bold="true" box="[151,292,1981,2005]" pageId="57" pageNumber="58">FIGURE 37.</emphasis>
<taxonomicName id="B5B6CDAF450CFF85FEF81CF3FDCE2AE0" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[302,576,1981,2005]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="trapezoides" status="gen. et spec. nov.">
<emphasis id="40C26A3E450CFF85FEF81CF3FDCE2AE0" box="[302,576,1981,2005]" italics="true" pageId="57" pageNumber="58">Benincandona trapezoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745450CFF85FD9C1CF0FC892AE0" box="[586,775,1982,2006]" pageId="57" pageNumber="58" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E450CFF85FC5A1CF3FC102AE0" bold="true" box="[908,926,1981,2005]" pageId="57" pageNumber="58">A</emphasis>
. T2 (INV.159263).
<emphasis id="40C26A3E450CFF85FBA41CF0FB0D2AE3" bold="true" box="[1138,1155,1982,2006]" pageId="57" pageNumber="58">B</emphasis>
. T3 (INV.159263).
<emphasis id="40C26A3E450CFF85FA811CF3FAE72AE0" bold="true" box="[1367,1385,1981,2005]" pageId="57" pageNumber="58">C</emphasis>
. CR (INV.159280).
<emphasis id="40C26A3E450CFF85FEE71CACFECD2ACF" bold="true" box="[305,323,2018,2042]" pageId="57" pageNumber="58">D</emphasis>
. CRAtt (INV.159280). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A4450FFF86FF411CD9FDB32ACD" pageId="58" pageNumber="59" startId="58.[151,250,1943,1967]" targetBox="[189,1403,188,1912]" targetPageId="58" targetType="figure">
<paragraph id="7209B62C450FFF86FF411CD9FDB32ACD" blockId="58.[151,1437,1943,2040]" pageId="58" pageNumber="59">
<emphasis id="40C26A3E450FFF86FF411CD9FEAE2A9A" bold="true" box="[151,288,1943,1967]" pageId="58" pageNumber="59">FIGURE 38.</emphasis>
<taxonomicName id="B5B6CDAF450FFF86FEF01CD9FDBD2A9A" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[294,563,1943,1967]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="trapezoides" status="gen. et spec. nov.">
<emphasis id="40C26A3E450FFF86FEF01CD9FDBD2A9A" box="[294,563,1943,1967]" italics="true" pageId="58" pageNumber="59">Benincandona trapezoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745450FFF86FDEF1CD6FD642A9A" box="[569,746,1944,1968]" pageId="58" pageNumber="59" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E450FFF86FCB41CD9FCFA2A9A" bold="true" box="[866,884,1943,1967]" pageId="58" pageNumber="59">A</emphasis>
. Zenker organ (INV.159263).
<emphasis id="40C26A3E450FFF86FB711CD6FB592A85" bold="true" box="[1191,1239,1944,1968]" pageId="58" pageNumber="59">BD</emphasis>
. Hp (INV.159263).
<emphasis id="40C26A3E450FFF86FF411CF2FF262AE1" bold="true" box="[151,168,1980,2004]" pageId="58" pageNumber="59">B</emphasis>
. External view (drawing).
<emphasis id="40C26A3E450FFF86FE141CF5FE5A2AE6" bold="true" box="[450,468,1979,2003]" pageId="58" pageNumber="59">C</emphasis>
. Internal view (photograph without interference contrast).
<emphasis id="40C26A3E450FFF86FBEE1CF2FBC42AE1" bold="true" box="[1080,1098,1980,2004]" pageId="58" pageNumber="59">D</emphasis>
. Internal view (photograph with interference contrast). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A4450EFF87FF411CE6FA9F2AD0" pageId="59" pageNumber="60" startId="59.[151,250,1960,1984]" targetBox="[175,1409,202,1932]" targetPageId="59" targetType="figure">
<paragraph id="7209B62C450EFF87FF411CE6FA9F2AD0" blockId="59.[151,1436,1960,2021]" pageId="59" pageNumber="60">
<emphasis id="40C26A3E450EFF87FF411CE6FEAE2AF4" bold="true" box="[151,288,1960,1985]" pageId="59" pageNumber="60">FIGURE 39.</emphasis>
<taxonomicName id="B5B6CDAF450EFF87FEF11CE7FDBB2AF4" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[295,565,1961,1985]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="trapezoides" status="gen. et spec. nov.">
<emphasis id="40C26A3E450EFF87FEF11CE7FDBB2AF4" box="[295,565,1961,1985]" italics="true" pageId="59" pageNumber="60">Benincandona trapezoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745450EFF87FDEA1CE7FD612AF5" box="[572,751,1961,1985]" pageId="59" pageNumber="60" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (AB ♀ and CE ♂).
<emphasis id="40C26A3E450EFF87FBCF1CE6FBA52AF5" bold="true" box="[1049,1067,1960,1984]" pageId="59" pageNumber="60">A</emphasis>
. T1, Lpp (INV.159280).
<emphasis id="40C26A3E450EFF87FAFD1CE7FAB22AF4" bold="true" box="[1323,1340,1961,1985]" pageId="59" pageNumber="60">B</emphasis>
. T1, Rpp (INV.159280).
<emphasis id="40C26A3E450EFF87FEE71C82FECD2AD1" bold="true" box="[305,323,1996,2020]" pageId="59" pageNumber="60">C</emphasis>
. A2 (INV.159264, INV.159269).
<emphasis id="40C26A3E450EFF87FD401C83FD262AD0" bold="true" box="[662,680,1997,2021]" pageId="59" pageNumber="60">D</emphasis>
. T1 (INV.159268).
<emphasis id="40C26A3E450EFF87FCA71C83FC0C2AD0" bold="true" box="[881,898,1997,2021]" pageId="59" pageNumber="60">E</emphasis>
. CR (INV.159264). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A44509FF80FF411EA2FD1F2B79" pageId="60" pageNumber="61" startId="60.[151,250,1516,1540]" targetBox="[151,1435,181,1492]" targetPageId="60" targetType="figure">
<paragraph id="7209B62C4509FF80FF411EA2FD1F2B79" blockId="60.[151,1437,1516,1612]" pageId="60" pageNumber="61">
<emphasis id="40C26A3E4509FF80FF411EA2FEAD2B31" bold="true" box="[151,291,1516,1540]" pageId="60" pageNumber="61">FIGURE 40.</emphasis>
Diagrams of apical views of the A2 of
<taxonomicName id="B5B6CDAF4509FF80FD071EA2FCED2B31" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[721,867,1516,1540]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="40C26A3E4509FF80FD071EA2FCED2B31" box="[721,867,1516,1540]" italics="true" pageId="60" pageNumber="61">Benincandona</emphasis>
</taxonomicName>
, showing position of (sub-) terminal claws, setae and aesthetascs of the terminal (small circle) and penultimate segment (large circle). For explanations of codes, see Material and Methods (Abbreviations used in text and figures).
</paragraph>
</caption>
<paragraph id="7209B62C4509FF80FF111DD0FCE42AA4" blockId="60.[151,1437,1694,1937]" pageId="60" pageNumber="61">
Mx1 (
<figureCitation id="EA8DAAA94509FF80FED91DD0FE322B8D" box="[271,444,1694,1720]" captionStart="FIGURE 36" captionStartId="56.[151,250,1943,1967]" captionTargetBox="[166,1418,185,1915]" captionTargetId="figure-31@56.[166,1422,181,1919]" captionTargetPageId="56" captionText="FIGURE 36. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Mx1, respiratory plate (INV.159263). B. Mx1 (Chaetotaxy incomplete) (INV.159263). C. MdPalp (INV.159263). Scale bar. 1 (A), 2 (C), 3 (B) = 50 µm." pageId="60" pageNumber="61">Figure 36A, B</figureCitation>
) with a basal (protopodite) part carrying a large respiratory plate (exopodite), three endites and a two-segmented palp (endopodite). Respiratory plate (
<figureCitation id="EA8DAAA94509FF80FC5C1D8CFB802BE9" box="[906,1038,1730,1756]" captionStart="FIGURE 36" captionStartId="56.[151,250,1943,1967]" captionTargetBox="[166,1418,185,1915]" captionTargetId="figure-31@56.[166,1422,181,1919]" captionTargetPageId="56" captionText="FIGURE 36. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Mx1, respiratory plate (INV.159263). B. Mx1 (Chaetotaxy incomplete) (INV.159263). C. MdPalp (INV.159263). Scale bar. 1 (A), 2 (C), 3 (B) = 50 µm." pageId="60" pageNumber="61">Figure 36A</figureCitation>
) elongated, carrying
<emphasis id="40C26A3E4509FF80FB291D8DFA9C2BE9" box="[1279,1298,1731,1756]" italics="true" pageId="60" pageNumber="61">c.</emphasis>
18 plumose respiratory rays, two short distal setae and a group of shorter proximal setae (not illustrated). Palp (
<figureCitation id="EA8DAAA94509FF80FAC61DA8FA1A2A35" box="[1296,1428,1766,1792]" captionStart="FIGURE 36" captionStartId="56.[151,250,1943,1967]" captionTargetBox="[166,1418,185,1915]" captionTargetId="figure-31@56.[166,1422,181,1919]" captionTargetPageId="56" captionText="FIGURE 36. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Mx1, respiratory plate (INV.159263). B. Mx1 (Chaetotaxy incomplete) (INV.159263). C. MdPalp (INV.159263). Scale bar. 1 (A), 2 (C), 3 (B) = 50 µm." pageId="60" pageNumber="61">Figure 36B</figureCitation>
) with first segment carrying four long dorso-apical setae; terminal segment short and rectangular, carrying four distal setae, some slightly claw-like; chaetotaxy of three endites incompletely illustrated: third endite with two strong teeth carrying lateral spines (
<emphasis id="40C26A3E4509FF80FE0F1C1CFDE92A59" box="[473,615,1874,1900]" italics="true" pageId="60" pageNumber="61">Zahnborsten</emphasis>
), basally with one lateral seta; sideways directed bristles near base of first endite, one long and one short,
<emphasis id="40C26A3E4509FF80FE211C39FD842AA5" box="[503,522,1911,1936]" italics="true" pageId="60" pageNumber="61">c.</emphasis>
half the length of the long one.
</paragraph>
<caption id="26C9E6A44508FF81FF411F17FB2529A3" pageId="61" pageNumber="62" startId="61.[151,250,1113,1137]" targetBox="[151,1436,181,1089]" targetPageId="61" targetType="figure">
<paragraph id="7209B62C4508FF81FF411F17FB2529A3" blockId="61.[151,1437,1113,1174]" pageId="61" pageNumber="62">
<emphasis id="40C26A3E4508FF81FF411F17FEAF2944" bold="true" box="[151,289,1113,1137]" pageId="61" pageNumber="62">FIGURE 41.</emphasis>
<taxonomicName id="B5B6CDAF4508FF81FEFE1F17FE342944" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[296,442,1113,1137]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="61" pageNumber="62" phylum="Arthropoda" rank="genus" status="gen. et spec. nov.">
<emphasis id="40C26A3E4508FF81FEFE1F17FE342944" box="[296,442,1113,1137]" italics="true" pageId="61" pageNumber="62">Benincandona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454508FF81FE141F14FDF82944" box="[450,630,1114,1138]" pageId="61" pageNumber="62" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, CpRl. (A, C, E, F, G, I, K, N, O, Q ♂ and B, D, H, J, L, M, P, R ♀).
<emphasis id="40C26A3E4508FF81FA9E1F17FAF72944" bold="true" box="[1352,1401,1113,1137]" pageId="61" pageNumber="62">AD</emphasis>
.
<emphasis id="40C26A3E4508FF81FA501F14FF6729A0" italics="true" pageId="61" pageNumber="62">
<taxonomicName id="B5B6CDAF4508FF81FA501F14FF6B29A0" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="martini">B. martini</taxonomicName>
.
</emphasis>
<emphasis id="40C26A3E4508FF81FF381F30FE9229A3" bold="true" box="[238,284,1150,1174]" pageId="61" pageNumber="62">EF</emphasis>
.
<emphasis id="40C26A3E4508FF81FEFE1F30FE3F29A0" box="[296,433,1149,1173]" italics="true" pageId="61" pageNumber="62">
<taxonomicName id="B5B6CDAF4508FF81FEFE1F30FE2329A0" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[296,429,1149,1173]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="ibikounlei">B. ibikounlei</taxonomicName>
.
</emphasis>
<emphasis id="40C26A3E4508FF81FE601F33FE6D29A3" bold="true" box="[438,483,1149,1174]" pageId="61" pageNumber="62">GJ</emphasis>
.
<emphasis id="40C26A3E4508FF81FE261F30FDDF29A0" box="[496,593,1149,1173]" italics="true" pageId="61" pageNumber="62">
<taxonomicName id="B5B6CDAF4508FF81FE261F30FDC329A0" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[496,589,1149,1173]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="sakitii">B. sakitii</taxonomicName>
.
</emphasis>
<emphasis id="40C26A3E4508FF81FD811F30FD0729A3" bold="true" box="[599,649,1150,1174]" pageId="61" pageNumber="62">KN</emphasis>
.
<emphasis id="40C26A3E4508FF81FD401F30FD7429A0" box="[662,762,1149,1173]" italics="true" pageId="61" pageNumber="62">
<taxonomicName id="B5B6CDAF4508FF81FD401F30FD7829A0" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[662,758,1149,1173]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="cebios">B. cebios</taxonomicName>
.
</emphasis>
<emphasis id="40C26A3E4508FF81FCD61F33FCBC29A3" bold="true" box="[768,818,1149,1174]" pageId="61" pageNumber="62">OR</emphasis>
.
<emphasis id="40C26A3E4508FF81FCE91F30FC5929A0" box="[831,983,1149,1173]" italics="true" pageId="61" pageNumber="62">
<taxonomicName id="B5B6CDAF4508FF81FCE91F30FC5D29A0" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[831,979,1149,1173]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="trapezoides">B. trapezoides</taxonomicName>
.
</emphasis>
Scale bar = 500 µm.
</paragraph>
</caption>
<paragraph id="7209B62C4508FF81FF111FA6FDA328A7" blockId="61.[151,1437,1256,2038]" pageId="61" pageNumber="62">
T1 (
<figureCitation id="EA8DAAA94508FF81FF201FA6FE152837" box="[246,411,1256,1282]" captionStart="FIGURE 39" captionStartId="59.[151,250,1960,1984]" captionTargetBox="[175,1409,202,1932]" captionTargetId="figure-17@59.[175,1413,198,1936]" captionTargetPageId="59" captionText="FIGURE 39. Benincandona trapezoides gen. et spec. nov., limbs (AB ♀ and CE ♂). A. T1, Lpp (INV.159280). B. T1, Rpp (INV.159280). C. A2 (INV.159264, INV.159269). D. T1 (INV.159268). E. CR (INV.159264). Scale bar = 50 µm." pageId="61" pageNumber="62">Figure 39A, B</figureCitation>
) consisting of basal part (protopodite), carrying respiratory plates (exopodites not shown). Protopodite (
<figureCitation id="EA8DAAA94508FF81FEFB1E42FE3C2813" box="[301,434,1292,1318]" captionStart="FIGURE 39" captionStartId="59.[151,250,1960,1984]" captionTargetBox="[175,1409,202,1932]" captionTargetId="figure-17@59.[175,1413,198,1936]" captionTargetPageId="59" captionText="FIGURE 39. Benincandona trapezoides gen. et spec. nov., limbs (AB ♀ and CE ♂). A. T1, Lpp (INV.159280). B. T1, Rpp (INV.159280). C. A2 (INV.159264, INV.159269). D. T1 (INV.159268). E. CR (INV.159264). Scale bar = 50 µm." pageId="61" pageNumber="62">Figure 39B</figureCitation>
) distally set with almost 10 plumose setae of different morphology and length; more proximally set with d, d and b setae. Prehensile palps asymmetrical Rpp (
<figureCitation id="EA8DAAA94508FF81FBD11E7EFB03287F" box="[1031,1165,1328,1354]" captionStart="FIGURE 39" captionStartId="59.[151,250,1960,1984]" captionTargetBox="[175,1409,202,1932]" captionTargetId="figure-17@59.[175,1413,198,1936]" captionTargetPageId="59" captionText="FIGURE 39. Benincandona trapezoides gen. et spec. nov., limbs (AB ♀ and CE ♂). A. T1, Lpp (INV.159280). B. T1, Rpp (INV.159280). C. A2 (INV.159264, INV.159269). D. T1 (INV.159268). E. CR (INV.159264). Scale bar = 50 µm." pageId="61" pageNumber="62">Figure 39B</figureCitation>
) slightly more swollen than Lpp (
<figureCitation id="EA8DAAA94508FF81FEDB1E1AFE1B285B" box="[269,405,1364,1390]" captionStart="FIGURE 39" captionStartId="59.[151,250,1960,1984]" captionTargetBox="[175,1409,202,1932]" captionTargetId="figure-17@59.[175,1413,198,1936]" captionTargetPageId="59" captionText="FIGURE 39. Benincandona trapezoides gen. et spec. nov., limbs (AB ♀ and CE ♂). A. T1, Lpp (INV.159280). B. T1, Rpp (INV.159280). C. A2 (INV.159264, INV.159269). D. T1 (INV.159268). E. CR (INV.159264). Scale bar = 50 µm." pageId="61" pageNumber="62">Figure 39A</figureCitation>
); one-segmented, distally sigmoid hook-like, both palps set with a short sensory organ and proximally with two subequal setae.
</paragraph>
<paragraph id="7209B62C4508FF81FF111ED2FB4E2B5F" blockId="61.[151,1437,1256,2038]" pageId="61" pageNumber="62">
T2 (
<figureCitation id="EA8DAAA94508FF81FF231ED2FEF62883" box="[245,376,1436,1462]" captionStart="FIGURE 37" captionStartId="57.[151,250,1981,2005]" captionTargetBox="[223,1388,222,1895]" captionTargetId="figure-17@57.[166,1422,181,1957]" captionTargetPageId="57" captionText="FIGURE 37. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. T2 (INV.159263). B. T3 (INV.159263). C. CR (INV.159280). D. CRAtt (INV.159280). Scale bar = 50 µm." pageId="61" pageNumber="62">Figure 37A</figureCitation>
) with 4-segmented endopodite (penultimate segment divided), generally elongated. First basal segment without setae d1 and d2. First segment of endopodite elongated (length more than four times basal width), without seta. Second segment (segment 2A) with length
<emphasis id="40C26A3E4508FF81FCF81EAAFCCF28C8" box="[814,833,1508,1533]" italics="true" pageId="61" pageNumber="62">c.</emphasis>
4x width with one short ventro-apical seta f. Third segment (segment 2B) about the same length/width ratio as the previous segment, set with one ventro-subapical seta g. Terminal segment (segment 3) almost as long as width, apically with two short setae (h1 and h3), h1 (ventral) and twice the length of h3 (dorsal), and a long and slender claw (h2), distally set with short setulae.
</paragraph>
<paragraph id="7209B62C4508FF81FF111D3AFF642A77" blockId="61.[151,1437,1256,2038]" pageId="61" pageNumber="62">
T3 (
<figureCitation id="EA8DAAA94508FF81FF231D3AFEF82BBB" box="[245,374,1652,1678]" captionStart="FIGURE 37" captionStartId="57.[151,250,1981,2005]" captionTargetBox="[223,1388,222,1895]" captionTargetId="figure-17@57.[166,1422,181,1957]" captionTargetPageId="57" captionText="FIGURE 37. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. T2 (INV.159263). B. T3 (INV.159263). C. CR (INV.159280). D. CRAtt (INV.159280). Scale bar = 50 µm." pageId="61" pageNumber="62">Figure 37B</figureCitation>
) with first segment carrying only one subapical seta d1, of intermediate length. Second (knee-) segment without seta d2. Third segment about four times as long as wide, not carrying any seta. Fourth segment narrower and at least 8x basal width and carrying one short subapical seta g. Terminal segment well-separated from penultimate segment, about twice as long as basal width and carrying three setae: one long and reflexed (h3), almost reaching knee-segment and two ventrally directed and curved setae h1 and h2, the later slightly exceeding of the former.
</paragraph>
<paragraph id="7209B62C4508FF81FF111C02FC862A9B" blockId="61.[151,1437,1256,2038]" pageId="61" pageNumber="62">
CR (
<figureCitation id="EA8DAAA94508FF81FF2C1C02FEF52A53" box="[250,379,1868,1894]" captionStart="FIGURE 37" captionStartId="57.[151,250,1981,2005]" captionTargetBox="[223,1388,222,1895]" captionTargetId="figure-17@57.[166,1422,181,1957]" captionTargetPageId="57" captionText="FIGURE 37. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. T2 (INV.159263). B. T3 (INV.159263). C. CR (INV.159280). D. CRAtt (INV.159280). Scale bar = 50 µm." pageId="61" pageNumber="62">Figure 37C</figureCitation>
) with stout ramus; claw Ga well-developed, stout with length slightly exceeding half the length of ramus, setae Sp, Sa and claw Gp reduced to small setae; seta Sp situated about at one-fifth length of ramus from distal end, Sa dorso-subapical and Gp ventro-subapical.
</paragraph>
<paragraph id="7209B62C4508FF81FF111CF6FA802AE7" blockId="61.[151,1437,1256,2038]" box="[199,1294,1976,2002]" pageId="61" pageNumber="62">
CRAtt (
<figureCitation id="EA8DAAA94508FF81FEC91CF6FE2C2AE7" box="[287,418,1976,2002]" captionStart="FIGURE 37" captionStartId="57.[151,250,1981,2005]" captionTargetBox="[223,1388,222,1895]" captionTargetId="figure-17@57.[166,1422,181,1957]" captionTargetPageId="57" captionText="FIGURE 37. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. T2 (INV.159263). B. T3 (INV.159263). C. CR (INV.159280). D. CRAtt (INV.159280). Scale bar = 50 µm." pageId="61" pageNumber="62">Figure 37D</figureCitation>
) long and stout, distally bifurcated, subapically with additional lateral branch.
</paragraph>
<paragraph id="7209B62C4508FF82FF111C92FD4E2DE0" blockId="61.[151,1437,1256,2038]" lastBlockId="62.[151,1436,151,249]" lastPageId="62" lastPageNumber="63" pageId="61" pageNumber="62">
Hp (
<figureCitation id="EA8DAAA94508FF81FF211C92FE142AC3" box="[247,410,2012,2038]" captionStart="FIGURE 38" captionStartId="58.[151,250,1943,1967]" captionTargetBox="[189,1403,188,1912]" captionTargetId="figure-31@58.[179,1408,181,1919]" captionTargetPageId="58" captionText="FIGURE 38. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159263). BD. Hp (INV.159263). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." pageId="61" pageNumber="62">Figure 38BD</figureCitation>
) with robust and rounded lobe ls, reaching well beyond the asymmetrically rounded lobe ms, both lobes overlapping, not diverging; lobe h shorter and sub-rectangular; internal anatomy (
<figureCitation id="EA8DAAA9450BFF82FB611BD9FAD52D84" box="[1207,1371,151,177]" captionStart="FIGURE 38" captionStartId="58.[151,250,1943,1967]" captionTargetBox="[189,1403,188,1912]" captionTargetId="figure-31@58.[179,1408,181,1919]" captionTargetPageId="58" captionText="FIGURE 38. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159263). BD. Hp (INV.159263). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." pageId="62" pageNumber="63">Figure 38C, D</figureCitation>
) with postlabyrinthal spermiduct with one circular coil.
</paragraph>
<paragraph id="7209B62C450BFF82FF111B91FC382DCC" blockId="62.[151,1436,151,249]" box="[199,950,223,249]" pageId="62" pageNumber="63">
Zenker organ (
<figureCitation id="EA8DAAA9450BFF82FEBD1B91FE602DCC" box="[363,494,223,249]" captionStart="FIGURE 38" captionStartId="58.[151,250,1943,1967]" captionTargetBox="[189,1403,188,1912]" captionTargetId="figure-31@58.[179,1408,181,1919]" captionTargetPageId="58" captionText="FIGURE 38. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159263). BD. Hp (INV.159263). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." pageId="62" pageNumber="63">Figure 38A</figureCitation>
) elongated with
<emphasis id="40C26A3E450BFF82FD7C1B91FD332DCD" box="[682,701,223,248]" italics="true" pageId="62" pageNumber="63">c.</emphasis>
seven spinous whirls.
</paragraph>
<paragraph id="7209B62C450BFF82FF411A68FE182C75" blockId="62.[151,1437,294,825]" box="[151,406,294,320]" pageId="62" pageNumber="63">
<emphasis id="40C26A3E450BFF82FF411A68FE182C75" bold="true" box="[151,406,294,320]" pageId="62" pageNumber="63">Description of female</emphasis>
</paragraph>
<paragraph id="7209B62C450BFF82FF411A05FB2E2C50" blockId="62.[151,1437,294,825]" box="[151,1184,331,357]" pageId="62" pageNumber="63">
Cp and valves (
<figureCitation id="EA8DAAA9450BFF82FE921A05FE7E2C50" box="[324,496,331,357]" captionStart="FIGURE 32" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure-17@51.[179,1408,181,1920]" captionTargetPageId="51" captionText="FIGURE 32. Benincandona trapezoides gen. et spec. nov., valves (AC, GK ♂; DF ♀). AF, H. CpRl (AC, H. INV.159276, DF. INV.1592 INV.159279). G, I, J. INV.159279. CpV. K. INV.159277. CpD. Scale bars. 1 (B, E, G, K) = 500 µm; 2 (A, C, D, F, I) = 200 µm, 3 (H, J) = 40 µm." pageId="62" pageNumber="63">Figures 32DF</figureCitation>
,
<figureCitation id="EA8DAAA9450BFF82FE2D1A05FDC02C50" box="[507,590,331,357]" captionStart="FIGURE 34" captionStartId="54.[151,250,1520,1544]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-31@54.[151,1436,181,1496]" captionTargetPageId="54" captionText="FIGURE 34. Benincandona trapezoides gen. et spec. nov., valves (♀). A, C, D, G. LVi (INV.159264). B, E, F, H. RVi (INV.159264). G, H. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (GH) = 50 µm." pageId="62" pageNumber="63">34AH</figureCitation>
) as in the male, without obvious sexual dimorphism.
</paragraph>
<paragraph id="7209B62C450BFF82FF111A21FB072CBC" blockId="62.[151,1437,294,825]" box="[199,1161,367,393]" pageId="62" pageNumber="63">
Limbs (
<figureCitation id="EA8DAAA9450BFF82FECB1A21FE312CBC" box="[285,447,367,393]" captionStart="FIGURE 39" captionStartId="59.[151,250,1960,1984]" captionTargetBox="[175,1409,202,1932]" captionTargetId="figure-17@59.[175,1413,198,1936]" captionTargetPageId="59" captionText="FIGURE 39. Benincandona trapezoides gen. et spec. nov., limbs (AB ♀ and CE ♂). A. T1, Lpp (INV.159280). B. T1, Rpp (INV.159280). C. A2 (INV.159264, INV.159269). D. T1 (INV.159268). E. CR (INV.159264). Scale bar = 50 µm." pageId="62" pageNumber="63">Figure 39CE</figureCitation>
) largely as in the male, but with sexually dimorphic A2 and T1.
</paragraph>
<paragraph id="7209B62C450BFF82FF111ADDFDCA2F08" blockId="62.[151,1437,294,825]" pageId="62" pageNumber="63">
A2 (
<figureCitation id="EA8DAAA9450BFF82FF2F1ADDFEF22C98" box="[249,380,403,429]" captionStart="FIGURE 39" captionStartId="59.[151,250,1960,1984]" captionTargetBox="[175,1409,202,1932]" captionTargetId="figure-17@59.[175,1413,198,1936]" captionTargetPageId="59" captionText="FIGURE 39. Benincandona trapezoides gen. et spec. nov., limbs (AB ♀ and CE ♂). A. T1, Lpp (INV.159280). B. T1, Rpp (INV.159280). C. A2 (INV.159264, INV.159269). D. T1 (INV.159268). E. CR (INV.159264). Scale bar = 50 µm." pageId="62" pageNumber="63">Figure 39C</figureCitation>
) with second endopodal segment undivided, with t1-seta (long, narrow and plumose) and t2- seta (thin and smooth); apical chaetotaxy (
<figureCitation id="EA8DAAA9450BFF82FDA91AF9FD7C2CE4" box="[639,754,439,465]" captionStart="FIGURE 40" captionStartId="60.[151,250,1516,1540]" captionTargetBox="[151,1435,181,1492]" captionTargetId="figure-31@60.[151,1435,181,1492]" captionTargetPageId="60" captionText="FIGURE 40. Diagrams of apical views of the A2 of Benincandona, showing position of (sub-) terminal claws, setae and aesthetascs of the terminal (small circle) and penultimate segment (large circle). For explanations of codes, see Material and Methods (Abbreviations used in text and figures)." pageId="62" pageNumber="63">Figure 40</figureCitation>
) with z1claw-like, z2 and z3 being both setae, z1 about 3x as long as z2, and z3 slightly shorter than z2; claws G1, G2, G3 and GM all reaching about the same point; seta g and aesthetasc y3 as in the male and the tip of Gm slightly longer than the tip of y3. Small claw Gm in female homologous to large claw in the male.
</paragraph>
<paragraph id="7209B62C450BFF82FF111909FB952F9C" blockId="62.[151,1437,294,825]" pageId="62" pageNumber="63">
T1 (
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) with basal part with
<emphasis id="40C26A3E450BFF82FDB01909FDF42F55" box="[614,634,583,608]" italics="true" pageId="62" pageNumber="63">c.</emphasis>
10 distal setae of unequal length and morphology, setae d and b present, with long additional seta (d) present close to seta d; respiratory plate represented by two long rays. Endopodite set with a short and narrow palp, with three subequal but short apical setae (h1h3).
</paragraph>
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<emphasis id="40C26A3E450BFF82FF1119FDFE0E2FF8" bold="true" box="[199,384,691,717]" pageId="62" pageNumber="63">Measurements:</emphasis>
See
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.
</paragraph>
</subSubSection>
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<emphasis id="40C26A3E450BFF82FF111998FE5B2FC5" bold="true" box="[199,469,726,752]" pageId="62" pageNumber="63">Differential Diagnosis:</emphasis>
See
<tableCitation id="3F348397450BFF82FDDC1999FDEE2FC4" box="[522,608,727,753]" captionStart="TABLE 4" captionStartId="20.[152,245,1665,1689]" captionText="TABLE 4. Differential character states between the five new species of Benincandona gen. nov." pageId="62" pageNumber="63">Table 4</tableCitation>
.
</paragraph>
</subSubSection>
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<emphasis id="40C26A3E450BFF82FF1119B5FE612E21" bold="true" box="[199,495,762,789]" pageId="62" pageNumber="63">Ecology and distribution</emphasis>
</paragraph>
<paragraph id="7209B62C450BFF82FF111851FC432E0C" blockId="62.[151,1437,294,825]" box="[199,973,799,825]" pageId="62" pageNumber="63">
This subterranean species was thus far only found in its
<typeStatus id="AD0D088E450BFF82FCEA1851FCE22E0C" box="[828,876,799,825]" pageId="62" pageNumber="63">type</typeStatus>
locality.
</paragraph>
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<emphasis id="40C26A3E450BFF82FF4118C4FD6D2E91" bold="true" box="[151,739,906,933]" pageId="62" pageNumber="63">
Other candonid species from the wells in
<collectingCountry id="0AA1F6BC450BFF82FD4C18C4FD6D2E91" box="[666,739,906,932]" name="Benin" pageId="62" pageNumber="63">Benin</collectingCountry>
</emphasis>
</paragraph>
<paragraph id="7209B62C450BFF82FF4118E1FE152EFC" blockId="62.[151,739,906,969]" box="[151,411,943,969]" pageId="62" pageNumber="63">
(
<figureCitation id="EA8DAAA9450BFF82FF4918E1FE952EFC" box="[159,283,943,969]" captionStart="FIGURE 42" captionStartId="64.[151,250,1943,1967]" captionTargetBox="[163,1424,181,1920]" captionTargetId="figure-31@64.[163,1424,181,1920]" captionTargetPageId="64" captionText="FIGURE 42. Scanning electron microscope illustration of fifteen of thirty further undescribed candonid species, (CpRl). Numbered: AO. Scale bar (for all individuals) = 500 µm." pageId="62" pageNumber="63">Figures 42</figureCitation>
,
<figureCitation id="EA8DAAA9450BFF82FEF018E1FED52EFC" box="[294,347,943,969]" captionStart="FIGURE 43" captionStartId="65.[151,250,1943,1967]" captionTargetBox="[160,1427,181,1920]" captionTargetId="figure-17@65.[160,1427,181,1920]" captionTargetPageId="65" captionText="FIGURE 43. Scanning electron microscope illustration of fifteen of thirty further undescribed candonid species, (CpRl). (Continued PAD). Scale bar (for all individuals) = 500 µm." pageId="62" pageNumber="63">43 A</figureCitation>
<collectingRegion id="B07278CE450BFF82FEBC18E1FE1D2EFC" box="[362,403,943,969]" country="Serbia" name="Ada" pageId="62" pageNumber="63">AD</collectingRegion>
)
</paragraph>
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Thirty further candonid species, in addition to the five new species described above, have been found in the samples from the wells in
<collectingCountry id="0AA1F6BC450BFF82FE8F1F55FE2E2900" box="[345,416,1051,1077]" name="Benin" pageId="62" pageNumber="63">Benin</collectingCountry>
. Most of these are also (sub-) triangular in lateral view, with strong (
<figureCitation id="EA8DAAA9450BFF82FB441F55FA852900" box="[1170,1291,1051,1077]" captionStart="FIGURE 42" captionStartId="64.[151,250,1943,1967]" captionTargetBox="[163,1424,181,1920]" captionTargetId="figure-31@64.[163,1424,181,1920]" captionTargetPageId="64" captionText="FIGURE 42. Scanning electron microscope illustration of fifteen of thirty further undescribed candonid species, (CpRl). Numbered: AO. Scale bar (for all individuals) = 500 µm." pageId="62" pageNumber="63">Figures 42</figureCitation>
/43E, N, AB) or weak (
<figureCitation id="EA8DAAA9450BFF82FF291F71FEF6296C" box="[255,376,1087,1113]" captionStart="FIGURE 42" captionStartId="64.[151,250,1943,1967]" captionTargetBox="[163,1424,181,1920]" captionTargetId="figure-31@64.[163,1424,181,1920]" captionTargetPageId="64" captionText="FIGURE 42. Scanning electron microscope illustration of fifteen of thirty further undescribed candonid species, (CpRl). Numbered: AO. Scale bar (for all individuals) = 500 µm." pageId="62" pageNumber="63">Figures 42</figureCitation>
/43K, M, Z) external lateral ridges; others have different shapes (
<figureCitation id="EA8DAAA9450BFF82FB9B1F71FB4B296C" box="[1101,1221,1087,1113]" captionStart="FIGURE 42" captionStartId="64.[151,250,1943,1967]" captionTargetBox="[163,1424,181,1920]" captionTargetId="figure-31@64.[163,1424,181,1920]" captionTargetPageId="64" captionText="FIGURE 42. Scanning electron microscope illustration of fifteen of thirty further undescribed candonid species, (CpRl). Numbered: AO. Scale bar (for all individuals) = 500 µm." pageId="62" pageNumber="63">Figures 42</figureCitation>
/43F, G, O, V, AC). Some species (
<figureCitation id="EA8DAAA9450BFF82FEE81F2DFD892948" box="[318,519,1123,1149]" captionStart="FIGURE 42" captionStartId="64.[151,250,1943,1967]" captionTargetBox="[163,1424,181,1920]" captionTargetId="figure-31@64.[163,1424,181,1920]" captionTargetPageId="64" captionText="FIGURE 42. Scanning electron microscope illustration of fifteen of thirty further undescribed candonid species, (CpRl). Numbered: AO. Scale bar (for all individuals) = 500 µm." pageId="62" pageNumber="63">Figure 42A, D, N</figureCitation>
) are much larger than most, and some are much smaller (
<figureCitation id="EA8DAAA9450BFF82FB5A1F2DFA882948" box="[1164,1286,1123,1149]" captionStart="FIGURE 42" captionStartId="64.[151,250,1943,1967]" captionTargetBox="[163,1424,181,1920]" captionTargetId="figure-31@64.[163,1424,181,1920]" captionTargetPageId="64" captionText="FIGURE 42. Scanning electron microscope illustration of fifteen of thirty further undescribed candonid species, (CpRl). Numbered: AO. Scale bar (for all individuals) = 500 µm." pageId="62" pageNumber="63">Figures 42</figureCitation>
/43O, X, AA, AD). Based on preliminary analyses of the soft parts of these species, it is at present most likely that all of these species belong to
<taxonomicName id="B5B6CDAF450BFF82FE891FE5FD6529F0" authority="Hotekpo &amp; Martens" authorityName="Hotekpo &amp; Martens" box="[351,747,1195,1221]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="62" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="40C26A3E450BFF82FE891FE5FD8F29F0" box="[351,513,1195,1221]" italics="true" pageId="62" pageNumber="63">Benincandona</emphasis>
Hotèkpo &amp; Martens
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, but this requires further confirmation.
</paragraph>
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</treatment>
</document>

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<mods:title id="ACA403E50B2C9DF0F1BD0CBD28B463C4">An endemic species flock of Candonidae Kaufmann, 1900 (Crustacea, Ostracoda) from wells in Benin (Africa), with the description of a new subfamily, a new genus and five new species</mods:title>
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<emphasis id="40C26A3E451DFF94FF411AB0FE152F2C" bold="true" box="[151,411,510,537]" italics="true" pageId="40" pageNumber="41">Benincandona cebios</emphasis>
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Hotèkpo &amp; Martens
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<paragraph id="7209B62C451DFF94FF411909FE252F54" blockId="40.[151,985,510,609]" box="[151,427,583,609]" pageId="40" pageNumber="41">
(
<figureCitation id="EA8DAAA9451DFF94FF491909FECA2F54" box="[159,324,583,609]" captionStart-0="FIGURE 24" captionStart-1="FIGURE 25" captionStart-2="FIGURE 26" captionStart-3="FIGURE 27" captionStart-4="FIGURE 28" captionStart-5="FIGURE 29" captionStart-6="FIGURE 30" captionStart-7="FIGURE 31" captionStartId-0="41.[151,250,1944,1968]" captionStartId-1="42.[151,250,1673,1697]" captionStartId-2="44.[151,250,1673,1697]" captionStartId-3="45.[151,250,1943,1967]" captionStartId-4="46.[151,250,1943,1967]" captionStartId-5="47.[151,250,1981,2005]" captionStartId-6="48.[151,250,1943,1967]" captionStartId-7="49.[151,250,1943,1967]" captionTargetBox-0="[170,1417,181,1921]" captionTargetBox-1="[151,1436,181,1649]" captionTargetBox-2="[151,1436,181,1649]" captionTargetBox-3="[173,1414,183,1915]" captionTargetBox-4="[188,1401,185,1915]" captionTargetBox-5="[197,1421,196,1942]" captionTargetBox-6="[194,1393,191,1910]" captionTargetBox-7="[184,1406,184,1915]" captionTargetId-0="figure-17@41.[170,1417,181,1921]" captionTargetId-1="figure-31@42.[151,1436,181,1649]" captionTargetId-2="figure-31@44.[151,1436,181,1649]" captionTargetId-3="figure-17@45.[171,1417,181,1919]" captionTargetId-4="figure-31@46.[184,1403,181,1919]" captionTargetId-5="figure-17@47.[166,1422,181,1957]" captionTargetId-6="figure-31@48.[184,1403,181,1920]" captionTargetId-7="figure-17@49.[180,1407,181,1919]" captionTargetPageId-0="41" captionTargetPageId-1="42" captionTargetPageId-2="44" captionTargetPageId-3="45" captionTargetPageId-4="46" captionTargetPageId-5="47" captionTargetPageId-6="48" captionTargetPageId-7="49" captionText-0="FIGURE 24. Benincandona cebios gen. et spec. nov., valves (AC, JK ♂; DI ♀). AI. CpRl (AC. INV.159255, DI. INV.159258). J. INV.159256. CpD. K. INV.159257. CpV. Scale bars. 1 (B, E, J, K) = 500 µm; 2 (A, C, D, F, H, I) = 200 µm." captionText-1="FIGURE 25. Benincandona cebios gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159244). B, E, F. RVi (INV.159244). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." captionText-2="FIGURE 26. Benincandona cebios gen. et spec. nov., valves (♀). A, C, D, G. LVi (INV.159253). B, E, F. RVi (INV.159253). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." captionText-3="FIGURE 27. Benincandona cebios gen. et spec. nov., limbs (♂). A. A2 (INV.159244). B. A1 (INV.159244). C. MdCoxa (INV.159248). Scale bar = 50 µm." captionText-4="FIGURE 28. Benincandona cebios gen. et spec. nov., limbs (♂). A. Mx1-palp (INV.159244). B. Mx1, respiratory plate (INV.159248). C. MdPalp (INV.159244). Scale bars. 1 (B), 2 (A), 3 (C) = 50 µm." captionText-5="FIGURE 29. Benincandona cebios gen. et spec. nov., limbs (♂). A. T3 (INV.159244). B. T2 (INV.159244). C. CRAtt (INV.159244). D. CR (INV.159244). Scale bar = 50 µm." captionText-6="FIGURE 30. Benincandona cebios gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159247). BD. Hp (INV.159244). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." captionText-7="FIGURE 31. Benincandona cebios gen. et spec. nov., limbs (AB ♂ and CE ♀). C. A2 (INV.159245). D. CR (INV.159245). E. T1 (INV.159245). A. T1, Lpp (INV.159244). B. T1, Rpp (INV.159244). Scale bar = 50 µm." pageId="40" pageNumber="41">Figures 2431</figureCitation>
,
<figureCitation id="EA8DAAA9451DFF94FE861909FE2D2F54" box="[336,419,583,609]" captionStart="FIGURE 41" captionStartId="61.[151,250,1113,1137]" captionTargetBox="[151,1436,181,1089]" captionTargetId="figure-17@61.[151,1436,181,1089]" captionTargetPageId="61" captionText="FIGURE 41. Benincandona gen. et spec. nov., CpRl. (A, C, E, F, G, I, K, N, O, Q ♂ and B, D, H, J, L, M, P, R ♀). AD. B. martini. EF. B. ibikounlei. GJ. B. sakitii. KN. B. cebios. OR. B. trapezoides. Scale bar = 500 µm." pageId="40" pageNumber="41">41KN</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A7451DFF94FF4119C1FC1F2E20" pageId="40" pageNumber="41" type="etymology">
<paragraph id="7209B62C451DFF94FF4119C1FE962F9C" blockId="40.[151,1436,655,789]" box="[151,280,655,681]" pageId="40" pageNumber="41">
<emphasis id="40C26A3E451DFF94FF4119C1FE962F9C" bold="true" box="[151,280,655,681]" pageId="40" pageNumber="41">Etymology</emphasis>
</paragraph>
<paragraph id="7209B62C451DFF94FF4119FDFC1F2E20" blockId="40.[151,1436,655,789]" pageId="40" pageNumber="41">
The species is named after CEBioS, the Belgian project “Capacities for Biodiversity and Sustainable Development”, which funded various field trips in
<collectingCountry id="0AA1F6BC451DFF94FDC91999FDEB2FC4" box="[543,613,727,753]" name="Benin" pageId="40" pageNumber="41">Benin</collectingCountry>
, the first visits of SJH to Brussels, and provided a microscope for work in
<collectingCountry id="0AA1F6BC451DFF94FF4119B5FF542E20" box="[151,218,763,789]" name="Benin" pageId="40" pageNumber="41">Benin</collectingCountry>
through the GTI program, the “Global Taxonomic Initiative”.
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A7451DFF94FF41180CFD732948" pageId="40" pageNumber="41" type="diagnosis">
<paragraph id="7209B62C451DFF94FF41180CFE842E69" blockId="40.[151,1437,834,1149]" box="[151,266,834,860]" pageId="40" pageNumber="41">
<emphasis id="40C26A3E451DFF94FF41180CFE842E69" bold="true" box="[151,266,834,860]" pageId="40" pageNumber="41">Diagnosis</emphasis>
</paragraph>
<paragraph id="7209B62C451DFF94FF411829FD732948" blockId="40.[151,1437,834,1149]" pageId="40" pageNumber="41">
CpRl elongated (L/H = 2.2), sub-triangular, with greatest width slightly behind the middle; LV dorsally extending beyond RV, anterior margin bluntly pointed and with anteroventral part straight, posterior margin asymmetrically rounded towards the ventral side, valve surface smooth in the anterior and posterior part, with slight ridges in the middle section. CpD and CpV narrow, with greatest width in the middle and evenly rounded lateral sides, posterior extremities bluntly rounded compared with anterior end pointed. A2 with seta X short, just reaching base of aesthetasc Y. This aesthetasc Y distally asymmetrically bifurcated. Second palp segment of Mx1 with L
<emphasis id="40C26A3E451DFF94FA5F1F55FA1B2901" box="[1417,1429,1051,1076]" italics="true" pageId="40" pageNumber="41">c</emphasis>
. 2.1x basal W. T3 with seta d2 short. Hp with ls and ms with overlapping positions, lobe ls broad, distally rounded, extending beyond lobe ms with
<emphasis id="40C26A3E451DFF94FE281F2DFDB92949" box="[510,567,1123,1148]" italics="true" pageId="40" pageNumber="41">circa</emphasis>
half of its length.
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A7451DFF94FF411FE5FC5C2AA0" pageId="40" pageNumber="41" type="materials_examined">
<paragraph id="7209B62C451DFF94FF411FE5FEB029F1" blockId="40.[151,318,1194,1221]" box="[151,318,1194,1221]" pageId="40" pageNumber="41">
<emphasis id="40C26A3E451DFF94FF411FE5FEB029F1" bold="true" box="[151,318,1194,1221]" pageId="40" pageNumber="41">Type material</emphasis>
</paragraph>
<paragraph id="7209B62C451DFF94FF411FBDFE8A2838" blockId="40.[151,1282,1267,1329]" box="[151,260,1267,1293]" pageId="40" pageNumber="41">
<emphasis id="40C26A3E451DFF94FF411FBDFE8A2838" bold="true" box="[151,260,1267,1293]" pageId="40" pageNumber="41">
<typeStatus id="AD0D088E451DFF94FF411FBDFE8A2838" box="[151,260,1267,1293]" pageId="40" pageNumber="41" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7209B62C451DFF94FF411E59FA8C2804" blockId="40.[151,1282,1267,1329]" box="[151,1282,1302,1329]" pageId="40" pageNumber="41">
<materialsCitation id="C2DEBC71451DFF94FF411E59FA8C2804" box="[151,1282,1302,1329]" collectionCode="INV" pageId="40" pageNumber="41" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="64B07DA5451DFF94FF411E59FF4D2804" box="[151,195,1303,1329]" count="1" pageId="40" pageNumber="41" type="male">1 ♂</specimenCount>
, with soft parts dissected in glycerine in a sealed slide, and with valves stored dry (
<collectionCode id="14A72EE9451DFF94FBB01E59FB182804" box="[1126,1174,1303,1329]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="40" pageNumber="41" type="Herbarium">INV</collectionCode>
.159244).
</materialsCitation>
</paragraph>
<paragraph id="7209B62C451DFF94FF411E10FF72284D" blockId="40.[151,736,1374,1437]" box="[151,252,1374,1400]" pageId="40" pageNumber="41">
<emphasis id="40C26A3E451DFF94FF411E10FF72284D" bold="true" box="[151,252,1374,1400]" pageId="40" pageNumber="41">
<typeStatus id="AD0D088E451DFF94FF411E10FF72284D" box="[151,252,1374,1400]" pageId="40" pageNumber="41" type="allotype">Allotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7209B62C451DFF94FF411ECDFD6E28A8" blockId="40.[151,736,1374,1437]" box="[151,736,1410,1437]" pageId="40" pageNumber="41">
<materialsCitation id="C2DEBC71451DFF94FF411ECDFD6E28A8" box="[151,736,1410,1437]" collectionCode="INV" pageId="40" pageNumber="41" specimenCount="1" specimenCount-female="1" typeStatus="allotype">
<specimenCount id="64B07DA5451DFF94FF411ECDFF4F28A8" box="[151,193,1411,1437]" count="1" pageId="40" pageNumber="41" type="female">1 ♀</specimenCount>
, dissected and stored as the male (
<collectionCode id="14A72EE9451DFF94FD931ECCFDF428A8" box="[581,634,1410,1437]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="40" pageNumber="41" type="Herbarium">INV</collectionCode>
.159245).
</materialsCitation>
</paragraph>
<paragraph id="7209B62C451DFF94FF411E85FE8128D0" blockId="40.[151,1436,1483,1653]" box="[151,271,1483,1509]" pageId="40" pageNumber="41">
<emphasis id="40C26A3E451DFF94FF411E85FE8128D0" bold="true" box="[151,271,1483,1509]" pageId="40" pageNumber="41">
<typeStatus id="AD0D088E451DFF94FF411E85FE8128D0" box="[151,271,1483,1509]" pageId="40" pageNumber="41" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7209B62C451DFF94FF411EA0FDF62B40" blockId="40.[151,1436,1483,1653]" pageId="40" pageNumber="41">
<materialsCitation id="C2DEBC71451DFF94FF411EA0FA162B3C" box="[151,1432,1518,1545]" collectionCode="INV" pageId="40" pageNumber="41" specimenCount="5" specimenCount-male="5" typeStatus="paratype">
<specimenCount id="64B07DA5451DFF94FF411EA0FF572B3C" box="[151,217,1518,1545]" count="5" pageId="40" pageNumber="41" type="male">5 ♂♂</specimenCount>
dissected and stored as the
<typeStatus id="AD0D088E451DFF94FDF51EA1FD0B2B3C" box="[547,645,1519,1545]" pageId="40" pageNumber="41" type="holotype">holotype</typeStatus>
(
<collectionCode id="14A72EE9451DFF94FD411EA0FD452B3C" box="[663,715,1518,1545]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="40" pageNumber="41" type="Herbarium">INV</collectionCode>
.159246, INV.159247, INV.159248, INV.159249, INV.159250)
</materialsCitation>
;
<materialsCitation id="C2DEBC71451DFF94FF411D5DFB5A2B18" box="[151,1236,1554,1581]" collectionCode="INV" pageId="40" pageNumber="41" specimenCount="4" specimenCount-female="4" typeStatus="paratype">
<specimenCount id="64B07DA5451DFF94FF411D5DFF542B18" box="[151,218,1555,1581]" count="4" pageId="40" pageNumber="41" type="female">4 ♀♀</specimenCount>
dissected and stored as the male (
<collectionCode id="14A72EE9451DFF94FDBB1D5CFD2C2B18" box="[621,674,1554,1581]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="40" pageNumber="41" type="Herbarium">INV</collectionCode>
.159251, INV.159252, INV.159253, INV.159254)
</materialsCitation>
;
<materialsCitation id="C2DEBC71451DFF94FB351D5DFBD92B64" collectionCode="INV" pageId="40" pageNumber="41" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="64B07DA5451DFF94FB351D5DFAAB2B18" box="[1251,1317,1555,1581]" count="3" pageId="40" pageNumber="41" type="male">3 ♂♂</specimenCount>
carapaces stored dry in micropalaeontological slides (
<collectionCode id="14A72EE9451DFF94FD5D1D78FD312B64" box="[651,703,1590,1617]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="40" pageNumber="41" type="Herbarium">INV</collectionCode>
.159255, INV.159256, INV.159257)
</materialsCitation>
;
<materialsCitation id="C2DEBC71451DFF94FBB31D79FDFA2B40" collectionCode="INV" pageId="40" pageNumber="41" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="64B07DA5451DFF94FBB31D79FB1C2B64" box="[1125,1170,1591,1617]" count="1" pageId="40" pageNumber="41" type="female">1 ♀</specimenCount>
carapace stored dry in micropalaeontological slide (
<collectionCode id="14A72EE9451DFF94FE0A1D14FD9F2B40" box="[476,529,1626,1653]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="40" pageNumber="41" type="Herbarium">INV</collectionCode>
.159258)
</materialsCitation>
.
</paragraph>
<paragraph id="7209B62C451DFF94FF411DEDFEBE2B89" blockId="40.[151,1436,1698,1833]" box="[151,304,1698,1725]" pageId="40" pageNumber="41">
<emphasis id="40C26A3E451DFF94FF411DEDFEBE2B89" bold="true" box="[151,304,1698,1725]" pageId="40" pageNumber="41">Type locality</emphasis>
</paragraph>
<paragraph id="7209B62C451DFF94FF411D89FBAC2A1C" blockId="40.[151,1436,1698,1833]" pageId="40" pageNumber="41">
<materialsCitation id="C2DEBC71451DFF94FF411D89FE652A30" country="Benin" location="Tchaourou District" pageId="40" pageNumber="41" specimenCount="1" stateProvince="Borgou">
<collectingCountry id="0AA1F6BC451DFF94FF411D89FF542BD4" box="[151,218,1735,1761]" name="Benin" pageId="40" pageNumber="41">Benin</collectingCountry>
<emphasis id="40C26A3E451DFF94FF371D89FF652BD4" bold="true" box="[225,235,1735,1761]" pageId="40" pageNumber="41"></emphasis>
<collectingRegion id="B07278CE451DFF94FF271D89FE5C2BD4" box="[241,466,1735,1761]" country="Benin" name="Borgou" pageId="40" pageNumber="41">Borgou Department</collectingRegion>
,
<location id="7769E0F7451DFF94FE0A1D89FD3F2BD4" LSID="urn:lsid:plazi:treatment:FA1F073A451DFF8EFF411AB0FEE8284C:7769E0F7451DFF94FE0A1D89FD3F2BD4" box="[476,689,1735,1761]" country="Benin" name="Tchaourou District" pageId="40" pageNumber="41" stateProvince="Borgou">Tchaourou District</location>
, town Badékparou, located in the Okpara catchment area (tributary of the
<collectingRegion id="B07278CE451DFF94FF091DA5FEBE2A30" box="[223,304,1771,1797]" country="Benin" name="Oueme" pageId="40" pageNumber="41">Ouémé</collectingRegion>
catchment area)
</materialsCitation>
, coordinates:
<geoCoordinate id="1782D0EB451DFF94FD501DA5FC832A30" box="[646,781,1771,1797]" degrees="9" direction="north" minutes="13.297" orientation="latitude" pageId="40" pageNumber="41" precision="1" value="9.221617">9°13.297N</geoCoordinate>
,
<geoCoordinate id="1782D0EB451DFF94FCCE1DA4FC152A30" box="[792,923,1770,1797]" degrees="2" direction="east" minutes="33.745" orientation="longitude" pageId="40" pageNumber="41" precision="1" value="2.5624166">2°33.745E</geoCoordinate>
, Leg.: S.J. Hotèkpo and M. Lagnika collected on
<date id="060890EC451DFF94FF6C1C41FDCB2A1C" box="[186,581,1807,1833]" pageId="40" pageNumber="41" value="2022-10-11" valueMax="2022-11-06" valueMin="2022-10-11">11 October and 06 November 2022</date>
(Locality code BEN065 in
<tableCitation id="3F348397451DFF94FCAD1C41FC432A1C" box="[891,973,1807,1833]" captionStart="TABLE 1" captionStartId="5.[152,244,929,955]" captionText="TABLE 1. Localities where Benincandona gen. nov. sp. nov. were collected, coordinates and hydro-morphological characteristics. In all stations, the water was clear and was used for both drinking water and household purposes by the riparian villages." pageId="40" pageNumber="41">Table 1</tableCitation>
and
<tableCitation id="3F348397451DFF94FBD21C41FB9B2A1C" box="[1028,1045,1807,1833]" captionStart="TABLE 2" captionStartId="6.[152,244,1634,1658]" captionText="TABLE 2. Measurements of physical and chemical water parameters and variables at the time of well fauna collection. WT= water temperature (°C), EC = electrical conductivity (µS cm-1), DO = dissolved oxygen (mg L-1). NM= no measurement." pageId="40" pageNumber="41">2</tableCitation>
).
</paragraph>
<paragraph id="7209B62C451DFF94FF411C18FE6E2A45" blockId="40.[151,978,1878,1941]" box="[151,480,1878,1904]" pageId="40" pageNumber="41">
<emphasis id="40C26A3E451DFF94FF411C18FE6E2A45" bold="true" box="[151,480,1878,1904]" pageId="40" pageNumber="41">Other material investigated</emphasis>
</paragraph>
<paragraph id="7209B62C451DFF94FF411C35FC5C2AA0" blockId="40.[151,978,1878,1941]" box="[151,978,1915,1941]" pageId="40" pageNumber="41">
See
<tableCitation id="3F348397451DFF94FF101C35FE972AA0" box="[198,281,1915,1941]" captionStart="TABLE 1" captionStartId="5.[152,244,929,955]" captionText="TABLE 1. Localities where Benincandona gen. nov. sp. nov. were collected, coordinates and hydro-morphological characteristics. In all stations, the water was clear and was used for both drinking water and household purposes by the riparian villages." pageId="40" pageNumber="41">Table 1</tableCitation>
for a list of localities where the present species was collected.
</paragraph>
</subSubSection>
<caption id="26C9E6A4451CFF95FF411CD6FA052AE0" pageId="41" pageNumber="42" startId="41.[151,250,1944,1968]" targetBox="[170,1417,181,1921]" targetPageId="41" targetType="figure">
<paragraph id="7209B62C451CFF95FF411CD6FA052AE0" blockId="41.[151,1437,1944,2005]" pageId="41" pageNumber="42">
<emphasis id="40C26A3E451CFF95FF411CD6FEAD2A85" bold="true" box="[151,291,1944,1968]" pageId="41" pageNumber="42">FIGURE 24.</emphasis>
<taxonomicName id="B5B6CDAF451CFF95FEFB1CD7FD842A84" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[301,522,1945,1969]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="cebios" status="gen. et spec. nov.">
<emphasis id="40C26A3E451CFF95FEFB1CD7FD842A84" box="[301,522,1945,1969]" italics="true" pageId="41" pageNumber="42">Benincandona cebios</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745451CFF95FDC51CD7FD412A85" box="[531,719,1945,1969]" pageId="41" pageNumber="42" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, valves (AC, JK ♂; DI ♀).
<emphasis id="40C26A3E451CFF95FBF41CD6FBC42A85" bold="true" box="[1058,1098,1944,1968]" pageId="41" pageNumber="42">AI</emphasis>
. CpRl (AC. INV.159255, DI. INV.159258).
<emphasis id="40C26A3E451CFF95FEFE1CF3FEBB2AE0" bold="true" box="[296,309,1981,2005]" pageId="41" pageNumber="42">J</emphasis>
. INV.159256. CpD.
<emphasis id="40C26A3E451CFF95FDD11CF3FD952AE0" bold="true" box="[519,539,1981,2005]" pageId="41" pageNumber="42">K</emphasis>
. INV.159257. CpV. Scale bars. 1 (B, E, J, K) = 500 µm; 2 (A, C, D, F, H, I) = 200 µm.
</paragraph>
</caption>
<caption id="26C9E6A4451FFF96FF411DC7FC172BF3" pageId="42" pageNumber="43" startId="42.[151,250,1673,1697]" targetBox="[151,1436,181,1649]" targetPageId="42" targetType="figure">
<paragraph id="7209B62C451FFF96FF411DC7FC172BF3" blockId="42.[151,1437,1673,1734]" pageId="42" pageNumber="43">
<emphasis id="40C26A3E451FFF96FF411DC7FEAE2B94" bold="true" box="[151,288,1673,1697]" pageId="42" pageNumber="43">FIGURE 25.</emphasis>
<taxonomicName id="B5B6CDAF451FFF96FEF11DC7FD8F2B94" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[295,513,1673,1697]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="cebios" status="gen. et spec. nov.">
<emphasis id="40C26A3E451FFF96FEF11DC7FD8F2B94" box="[295,513,1673,1697]" italics="true" pageId="42" pageNumber="43">Benincandona cebios</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745451FFF96FDDE1DC4FD352B94" box="[520,699,1674,1698]" pageId="42" pageNumber="43" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, valves (♂).
<emphasis id="40C26A3E451FFF96FCEB1DC7FC3E2B94" bold="true" box="[829,944,1673,1698]" pageId="42" pageNumber="43">A, C, D, G</emphasis>
. LVi (INV.159244).
<emphasis id="40C26A3E451FFF96FB551DC4FB402B97" bold="true" box="[1155,1230,1674,1698]" pageId="42" pageNumber="43">B, E, F</emphasis>
. RVi (INV.159244).
<emphasis id="40C26A3E451FFF96FF411DE3FF3F2BF3" bold="true" box="[151,177,1709,1734]" pageId="42" pageNumber="43">G.</emphasis>
CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm.
</paragraph>
</caption>
<subSubSection id="3AACE5A7451FFF8EFF411C59FD9E29DC" lastPageId="50" lastPageNumber="51" pageId="42" pageNumber="43" type="description">
<paragraph id="7209B62C451FFF96FF411C59FE0F2A07" blockId="42.[151,1436,1815,2022]" box="[151,385,1815,1842]" pageId="42" pageNumber="43">
<emphasis id="40C26A3E451FFF96FF411C59FE0F2A07" bold="true" box="[151,385,1815,1842]" pageId="42" pageNumber="43">Description of male</emphasis>
</paragraph>
<paragraph id="7209B62C451FFF97FF411C72FDA32DE0" blockId="42.[151,1436,1815,2022]" lastBlockId="43.[151,1437,151,2013]" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
CpRl (
<figureCitation id="EA8DAAA9451FFF96FF351C72FE042A63" box="[227,394,1852,1878]" captionStart="FIGURE 24" captionStartId="41.[151,250,1944,1968]" captionTargetBox="[170,1417,181,1921]" captionTargetId="figure-17@41.[170,1417,181,1921]" captionTargetPageId="41" captionText="FIGURE 24. Benincandona cebios gen. et spec. nov., valves (AC, JK ♂; DI ♀). AI. CpRl (AC. INV.159255, DI. INV.159258). J. INV.159256. CpD. K. INV.159257. CpV. Scale bars. 1 (B, E, J, K) = 500 µm; 2 (A, C, D, F, H, I) = 200 µm." pageId="42" pageNumber="43">Figure 24AC</figureCitation>
) elongated (L/H ratio=2.0) and sub-triangular, dorsal margin bluntly pointed, with greatest height slightly behind the middle, anterior part of dorsal margin sloping almost straight to anterior margin, passing into it with a blunt angle, anteroventral part of anterior margin straight, posterior part of dorsal margin sloping to ventral margin, the latter almost straight; LV overlapping RV along anterior, ventral and posterior sides, dorsally strongly overlapping RV; valve surface smooth in the anterior and posterior parts, with slight ridges in the middle section. CpD and CpV (
<figureCitation id="EA8DAAA9451EFF97FE7E1BD9FDC62D84" box="[424,584,151,177]" captionStart="FIGURE 24" captionStartId="41.[151,250,1944,1968]" captionTargetBox="[170,1417,181,1921]" captionTargetId="figure-17@41.[170,1417,181,1921]" captionTargetPageId="41" captionText="FIGURE 24. Benincandona cebios gen. et spec. nov., valves (AC, JK ♂; DI ♀). AI. CpRl (AC. INV.159255, DI. INV.159258). J. INV.159256. CpD. K. INV.159257. CpV. Scale bars. 1 (B, E, J, K) = 500 µm; 2 (A, C, D, F, H, I) = 200 µm." pageId="43" pageNumber="44">Figure 24J, K</figureCitation>
) narrow and pointed on both anterior and posterior sides, greatest width in the middle, lateral margins rounded.
</paragraph>
<paragraph id="7209B62C451EFF97FF111B91FEA92CBC" blockId="43.[151,1437,151,2013]" pageId="43" pageNumber="44">
LVi (
<figureCitation id="EA8DAAA9451EFF97FED71B91FE7F2DCC" box="[257,497,222,249]" captionStart="FIGURE 25" captionStartId="42.[151,250,1673,1697]" captionTargetBox="[151,1436,181,1649]" captionTargetId="figure-31@42.[151,1436,181,1649]" captionTargetPageId="42" captionText="FIGURE 25. Benincandona cebios gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159244). B, E, F. RVi (INV.159244). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." pageId="43" pageNumber="44">Figure 25A, C, D, G</figureCitation>
) elongated and sub-triangular, dorsal margin bluntly pointed, with greatest height slightly behind the middle, anterior part of dorsal margin sloping almost straight to anterior margin, passing into it with a blunt angle, anteroventral part of anterior margin straight; posterior part of dorsal margin sloping towards ventral margin, the latter almost straight; one ventral il without antero-ventral tooth. CMS (
<figureCitation id="EA8DAAA9451EFF97FB711A05FAA42C50" box="[1191,1322,330,357]" captionStart="FIGURE 25" captionStartId="42.[151,250,1673,1697]" captionTargetBox="[151,1436,181,1649]" captionTargetId="figure-31@42.[151,1436,181,1649]" captionTargetPageId="42" captionText="FIGURE 25. Benincandona cebios gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159244). B, E, F. RVi (INV.159244). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." pageId="43" pageNumber="44">Figure 25G</figureCitation>
) as in the
<typeStatus id="AD0D088E451EFF97FF411A21FF492CBC" box="[151,199,367,393]" pageId="43" pageNumber="44">type</typeStatus>
species.
</paragraph>
<paragraph id="7209B62C451EFF97FF111ADDFC882F2C" blockId="43.[151,1437,151,2013]" pageId="43" pageNumber="44">
RVi (
<figureCitation id="EA8DAAA9451EFF97FED41ADDFE4A2C98" box="[258,452,402,429]" captionStart="FIGURE 25" captionStartId="42.[151,250,1673,1697]" captionTargetBox="[151,1436,181,1649]" captionTargetId="figure-31@42.[151,1436,181,1649]" captionTargetPageId="42" captionText="FIGURE 25. Benincandona cebios gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159244). B, E, F. RVi (INV.159244). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." pageId="43" pageNumber="44">Figure 25B, E, F</figureCitation>
) elongated, with dorsal margin partly straight and with greatest height well behind the middle, anterior part of dorsal margin sloping almost straight to anterior margin, passing into it with a blunt angle and with anteroventral part straight, posterior part of dorsal margin sloping towards ventral margin, the latter almost straight; il running submarginally along ventral margin.
</paragraph>
<paragraph id="7209B62C451EFF97FF11196DFC8B2E90" blockId="43.[151,1437,151,2013]" pageId="43" pageNumber="44">
A1 (
<figureCitation id="EA8DAAA9451EFF97FF2C196DFEF02F08" box="[250,382,547,573]" captionStart="FIGURE 27" captionStartId="45.[151,250,1943,1967]" captionTargetBox="[173,1414,183,1915]" captionTargetId="figure-17@45.[171,1417,181,1919]" captionTargetPageId="45" captionText="FIGURE 27. Benincandona cebios gen. et spec. nov., limbs (♂). A. A2 (INV.159244). B. A1 (INV.159244). C. MdCoxa (INV.159248). Scale bar = 50 µm." pageId="43" pageNumber="44">Figure 27B</figureCitation>
) consisting of seven articulated segments, one basal and six distal. Basal segment with two ventro-subapical setae, one long and one shorter, the latter about one third of the length of the longer seta, and two shorter setae on the mid-dorsal side, situated some distance from each other. First distal segment about three times as long as the basal width, with one short dorso-subapical seta. Second distal segment with about the same length/ width ratio as the previous segment, set with one short dorso-apical seta. Third distal segment also with about the same length/width ratio as the previous segment, carrying one short apical setae on each side. Fourth distal segment more than three times as long as wide, and set with two short setae as on the previous segment. Fifth distal segment more than five times as long as wide, set with one short ventro-apical seta and two long dorso-apical plumose setae. Sixth distal segment narrow, more than four times as long as wide, with four apical structures: one aesthetasc Ya of intermediate length, two long but unequal length plumose setae, about twice and 1.5x the length of Ya respectively, and one short plumose seta, about half the length of Ya.
</paragraph>
<paragraph id="7209B62C451EFF97FF1118E1FA9A28D0" blockId="43.[151,1437,151,2013]" pageId="43" pageNumber="44">
A2 (
<figureCitation id="EA8DAAA9451EFF97FF2D18E1FE0C2EFC" box="[251,386,943,969]" captionStart="FIGURE 27" captionStartId="45.[151,250,1943,1967]" captionTargetBox="[173,1414,183,1915]" captionTargetId="figure-17@45.[171,1417,181,1919]" captionTargetPageId="45" captionText="FIGURE 27. Benincandona cebios gen. et spec. nov., limbs (♂). A. A2 (INV.159244). B. A1 (INV.159244). C. MdCoxa (INV.159248). Scale bar = 50 µm." pageId="43" pageNumber="44">Figure 27A</figureCitation>
) with basal segment (first protopodite) broad and rounded, mid-ventrally with one seta of medium length, subapically with two setae of unequal length, proximal one
<emphasis id="40C26A3E451EFF97FBD2189DFB992ED9" box="[1028,1047,979,1004]" italics="true" pageId="43" pageNumber="44">c.</emphasis>
five times the length of the more apical one (plumose). Second segment (second protopodite) large and
<emphasis id="40C26A3E451EFF97FC1E18B9FC522925" box="[968,988,1015,1040]" italics="true" pageId="43" pageNumber="44">c.</emphasis>
twice as long as basal width, set with one short and plumose ventro-subapical seta X. No remnant of exopodite visible. Endopodite consisting of four segments (penultimate segment divided). First endopodal segment large and stout,
<emphasis id="40C26A3E451EFF97FB911F71FBD4296D" box="[1095,1114,1087,1112]" italics="true" pageId="43" pageNumber="44">c.</emphasis>
three times as long as basal width, carrying one long mid-ventral aesthetasc Y (tip clearly exceeding half of segment) and two ventro-apical setae, one short and one large and stout, hirsute in its distal 4/5. Second endopodal segment (segment 2A)
<emphasis id="40C26A3E451EFF97FA931FC9FAD62995" box="[1349,1368,1159,1184]" italics="true" pageId="43" pageNumber="44">c.</emphasis>
twice as long as wide, carrying one stout dorso-apical plumose seta and three t-setae; seta t1 long, narrow and plumose, accompanied by a long and slender, but well-developed aesthetasc y1; setae t2 and t3 respectively ventro and centro-apical, hyper-developed into male bristles. Third endopodal segment (segment 2B) smaller than previous segment and about 1.5x as long as wide, set with a complex (and sexually dimorphic) apical chaetotaxy (
<figureCitation id="EA8DAAA9451EFF97FB101E59FAB72804" box="[1222,1337,1303,1329]" captionStart="FIGURE 40" captionStartId="60.[151,250,1516,1540]" captionTargetBox="[151,1435,181,1492]" captionTargetId="figure-31@60.[151,1435,181,1492]" captionTargetPageId="60" captionText="FIGURE 40. Diagrams of apical views of the A2 of Benincandona, showing position of (sub-) terminal claws, setae and aesthetascs of the terminal (small circle) and penultimate segment (large circle). For explanations of codes, see Material and Methods (Abbreviations used in text and figures)." pageId="43" pageNumber="44">Figure 40</figureCitation>
): ventro-apically with one slender aesthetasc y2 and claw G3 transformed into short seta of almost equal length; centrally with one short plumose seta z3; dorso-apically, z1 claw-like, length about 1.7x as long as penultimate segment. Claw G2 about 2.5 times as long as z1 with G1 slightly shorter; z2 seta transformed into long claw, reaching tip of G1. Fourth, terminal segment (segment 3) with one long claw (Gm), one short claw (GM), a long aesthetasc y3 with shorter accompanying seta, with fused part short, and a seta g (slightly exceeding tip of aesthetasc y3).
</paragraph>
<paragraph id="7209B62C451EFF97FF111EA1FAA52B18" blockId="43.[151,1437,151,2013]" pageId="43" pageNumber="44">
MdCoxa (
<figureCitation id="EA8DAAA9451EFF97FEED1EA1FE4E2B3C" box="[315,448,1519,1545]" captionStart="FIGURE 27" captionStartId="45.[151,250,1943,1967]" captionTargetBox="[173,1414,183,1915]" captionTargetId="figure-17@45.[171,1417,181,1919]" captionTargetPageId="45" captionText="FIGURE 27. Benincandona cebios gen. et spec. nov., limbs (♂). A. A2 (INV.159244). B. A1 (INV.159244). C. MdCoxa (INV.159248). Scale bar = 50 µm." pageId="43" pageNumber="44">Figure 27C</figureCitation>
) elongated, proximally sharply pointed, medially widened, distally with an obliquely widened end, set with a series (67) of strong teeth, interspaced with setae; subapically with a short seta.
</paragraph>
<paragraph id="7209B62C451EFF97FF111D79FAEA2A78" blockId="43.[151,1437,151,2013]" pageId="43" pageNumber="44">
MdPalp (
<figureCitation id="EA8DAAA9451EFF97FEF91D79FE3F2B64" box="[303,433,1591,1617]" captionStart="FIGURE 28" captionStartId="46.[151,250,1943,1967]" captionTargetBox="[188,1401,185,1915]" captionTargetId="figure-31@46.[184,1403,181,1919]" captionTargetPageId="46" captionText="FIGURE 28. Benincandona cebios gen. et spec. nov., limbs (♂). A. Mx1-palp (INV.159244). B. Mx1, respiratory plate (INV.159248). C. MdPalp (INV.159244). Scale bars. 1 (B), 2 (A), 3 (C) = 50 µm." pageId="43" pageNumber="44">Figure 28C</figureCitation>
) with four segments. Basal segment ventro-apically with two hirsute setae (S1 &amp; S2); S1 long, reaching tip of terminal segment and S2 short and stout, length about one fifth of the length of S1; in between with the short flagellum-like alpha seta (thin and about twice the length of S2), and one smooth and medium size, more proximal seta. Second segment of sub-triangular shape, dorso-apically with two setae of unequal length; ventro-apically with a group of four long setae, one of these being the beta-seta. Third segment with two unequal dorso-subapical setae, two mid-dorsal setae, one of which being the gamma seta, and ventro-apically with one seta of medium length. Terminal segment sub-rectangular, apically set with one large claw, fused with the segment over its entire distal margin, accompanied by one smooth lateral seta on each side, slightly shorter than fused claw.
</paragraph>
<paragraph id="7209B62C451EFF97FF111C19FF6B2AE8" blockId="43.[151,1437,151,2013]" pageId="43" pageNumber="44">
Mx1 (
<figureCitation id="EA8DAAA9451EFF97FEDC1C19FE232A44" box="[266,429,1879,1905]" captionStart="FIGURE 28" captionStartId="46.[151,250,1943,1967]" captionTargetBox="[188,1401,185,1915]" captionTargetId="figure-31@46.[184,1403,181,1919]" captionTargetPageId="46" captionText="FIGURE 28. Benincandona cebios gen. et spec. nov., limbs (♂). A. Mx1-palp (INV.159244). B. Mx1, respiratory plate (INV.159248). C. MdPalp (INV.159244). Scale bars. 1 (B), 2 (A), 3 (C) = 50 µm." pageId="43" pageNumber="44">Figure 28A, B</figureCitation>
) with first palp segment carrying four long dorso-apical setae; terminal segment short and rectangular, carrying four distal setae, some slightly claw-like. Respiratory plate (
<figureCitation id="EA8DAAA9451EFF97FBE61C35FB3D2AA0" box="[1072,1203,1915,1941]" captionStart="FIGURE 28" captionStartId="46.[151,250,1943,1967]" captionTargetBox="[188,1401,185,1915]" captionTargetId="figure-31@46.[184,1403,181,1919]" captionTargetPageId="46" captionText="FIGURE 28. Benincandona cebios gen. et spec. nov., limbs (♂). A. Mx1-palp (INV.159244). B. Mx1, respiratory plate (INV.159248). C. MdPalp (INV.159244). Scale bars. 1 (B), 2 (A), 3 (C) = 50 µm." pageId="43" pageNumber="44">Figure 28B</figureCitation>
) elongated, carrying
<emphasis id="40C26A3E451EFF97FF411CD1FF242A8D" box="[151,170,1951,1976]" italics="true" pageId="43" pageNumber="44">c.</emphasis>
16 plumose respiratory rays, two short distal setae and a group of about seven shorter proximal setae of unequal length.
</paragraph>
<caption id="26C9E6A44519FF90FF411DC7FC172BF3" pageId="44" pageNumber="45" startId="44.[151,250,1673,1697]" targetBox="[151,1436,181,1649]" targetPageId="44" targetType="figure">
<paragraph id="7209B62C4519FF90FF411DC7FC172BF3" blockId="44.[151,1437,1673,1734]" pageId="44" pageNumber="45">
<emphasis id="40C26A3E4519FF90FF411DC7FEAE2B94" bold="true" box="[151,288,1673,1697]" pageId="44" pageNumber="45">FIGURE 26.</emphasis>
<taxonomicName id="B5B6CDAF4519FF90FEF11DC7FD8F2B94" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[295,513,1673,1697]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="cebios" status="gen. et spec. nov.">
<emphasis id="40C26A3E4519FF90FEF11DC7FD8F2B94" box="[295,513,1673,1697]" italics="true" pageId="44" pageNumber="45">Benincandona cebios</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454519FF90FDDE1DC4FD352B94" box="[520,699,1674,1698]" pageId="44" pageNumber="45" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, valves (♀).
<emphasis id="40C26A3E4519FF90FCEB1DC7FC3E2B94" bold="true" box="[829,944,1673,1698]" pageId="44" pageNumber="45">A, C, D, G</emphasis>
. LVi (INV.159253).
<emphasis id="40C26A3E4519FF90FB551DC4FB402B97" bold="true" box="[1155,1230,1674,1698]" pageId="44" pageNumber="45">B, E, F</emphasis>
. RVi (INV.159253).
<emphasis id="40C26A3E4519FF90FF411DE3FF3F2BF3" bold="true" box="[151,177,1709,1734]" pageId="44" pageNumber="45">G.</emphasis>
CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm.
</paragraph>
</caption>
<paragraph id="7209B62C4519FF90FF111C56FB2B2AF7" blockId="44.[151,1436,1816,1986]" pageId="44" pageNumber="45">
T1 (
<figureCitation id="EA8DAAA94519FF90FF231C56FE142A07" box="[245,410,1816,1842]" captionStart="FIGURE 31" captionStartId="49.[151,250,1943,1967]" captionTargetBox="[184,1406,184,1915]" captionTargetId="figure-17@49.[180,1407,181,1919]" captionTargetPageId="49" captionText="FIGURE 31. Benincandona cebios gen. et spec. nov., limbs (AB ♂ and CE ♀). C. A2 (INV.159245). D. CR (INV.159245). E. T1 (INV.159245). A. T1, Lpp (INV.159244). B. T1, Rpp (INV.159244). Scale bar = 50 µm." pageId="44" pageNumber="45">Figure 31A, B</figureCitation>
) consisting of basal part (protopodite), carrying respiratory plates (exopodites) represented by three plumose setae, two short and one medium length. Protopodite (
<figureCitation id="EA8DAAA94519FF90FC011C72FBD42A63" box="[983,1114,1852,1878]" captionStart="FIGURE 31" captionStartId="49.[151,250,1943,1967]" captionTargetBox="[184,1406,184,1915]" captionTargetId="figure-17@49.[180,1407,181,1919]" captionTargetPageId="49" captionText="FIGURE 31. Benincandona cebios gen. et spec. nov., limbs (AB ♂ and CE ♀). C. A2 (INV.159245). D. CR (INV.159245). E. T1 (INV.159245). A. T1, Lpp (INV.159244). B. T1, Rpp (INV.159244). Scale bar = 50 µm." pageId="44" pageNumber="45">Figure 31B</figureCitation>
) distally set with almost 10 plumose setae of different morphology and length; more proximally set with d, d and b setae. Prehensile palps asymmetrical with Rpp (
<figureCitation id="EA8DAAA94519FF90FE711CCAFDA92AAB" box="[423,551,1924,1950]" captionStart="FIGURE 31" captionStartId="49.[151,250,1943,1967]" captionTargetBox="[184,1406,184,1915]" captionTargetId="figure-17@49.[180,1407,181,1919]" captionTargetPageId="49" captionText="FIGURE 31. Benincandona cebios gen. et spec. nov., limbs (AB ♂ and CE ♀). C. A2 (INV.159245). D. CR (INV.159245). E. T1 (INV.159245). A. T1, Lpp (INV.159244). B. T1, Rpp (INV.159244). Scale bar = 50 µm." pageId="44" pageNumber="45">Figure 31B</figureCitation>
) slightly more swollen than Lpp (
<figureCitation id="EA8DAAA94519FF90FC4A1CCAFBAE2AAB" box="[924,1056,1924,1950]" captionStart="FIGURE 31" captionStartId="49.[151,250,1943,1967]" captionTargetBox="[184,1406,184,1915]" captionTargetId="figure-17@49.[180,1407,181,1919]" captionTargetPageId="49" captionText="FIGURE 31. Benincandona cebios gen. et spec. nov., limbs (AB ♂ and CE ♀). C. A2 (INV.159245). D. CR (INV.159245). E. T1 (INV.159245). A. T1, Lpp (INV.159244). B. T1, Rpp (INV.159244). Scale bar = 50 µm." pageId="44" pageNumber="45">Figure 31A</figureCitation>
); one-segmented, distally sigmoid hook-like, both palps set with a short sensory organ and proximally with two subequal setae.
</paragraph>
<caption id="26C9E6A44518FF91FF411CD9FE7C2AE1" pageId="45" pageNumber="46" startId="45.[151,250,1943,1967]" targetBox="[173,1414,183,1915]" targetPageId="45" targetType="figure">
<paragraph id="7209B62C4518FF91FF411CD9FE7C2AE1" blockId="45.[151,1437,1943,2004]" pageId="45" pageNumber="46">
<emphasis id="40C26A3E4518FF91FF411CD9FEAA2A9A" bold="true" box="[151,292,1943,1967]" pageId="45" pageNumber="46">FIGURE 27.</emphasis>
<taxonomicName id="B5B6CDAF4518FF91FEFB1CD6FD842A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[301,522,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="cebios" status="gen. et spec. nov.">
<emphasis id="40C26A3E4518FF91FEFB1CD6FD842A85" box="[301,522,1944,1968]" italics="true" pageId="45" pageNumber="46">Benincandona cebios</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454518FF91FDC21CD6FD5E2A9A" box="[532,720,1944,1968]" pageId="45" pageNumber="46" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E4518FF91FC821CD9FCE82A9A" bold="true" box="[852,870,1943,1967]" pageId="45" pageNumber="46">A</emphasis>
. A2 (INV.159244).
<emphasis id="40C26A3E4518FF91FBED1CD6FBC22A85" bold="true" box="[1083,1100,1944,1968]" pageId="45" pageNumber="46">B</emphasis>
. A1 (INV.159244).
<emphasis id="40C26A3E4518FF91FAF71CD9FABD2A9A" bold="true" box="[1313,1331,1943,1967]" pageId="45" pageNumber="46">C</emphasis>
. MdCoxa (INV.159248). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A4451BFF92FF411CD9FC5F2AE1" pageId="46" pageNumber="47" startId="46.[151,250,1943,1967]" targetBox="[188,1401,185,1915]" targetPageId="46" targetType="figure">
<paragraph id="7209B62C451BFF92FF411CD9FC5F2AE1" blockId="46.[151,1436,1943,2004]" pageId="46" pageNumber="47">
<emphasis id="40C26A3E451BFF92FF411CD9FEAB2A9A" bold="true" box="[151,293,1943,1967]" pageId="46" pageNumber="47">FIGURE 28.</emphasis>
<taxonomicName id="B5B6CDAF451BFF92FEF91CD6FD802A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[303,526,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="cebios" status="gen. et spec. nov.">
<emphasis id="40C26A3E451BFF92FEF91CD6FD802A85" box="[303,526,1944,1968]" italics="true" pageId="46" pageNumber="47">Benincandona cebios</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745451BFF92FDCE1CD6FD562A9A" box="[536,728,1944,1968]" pageId="46" pageNumber="47" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E451BFF92FC881CD9FCFE2A9A" bold="true" box="[862,880,1943,1967]" pageId="46" pageNumber="47">A</emphasis>
. Mx1-palp (INV.159244).
<emphasis id="40C26A3E451BFF92FB591CD6FB2E2A85" bold="true" box="[1167,1184,1944,1968]" pageId="46" pageNumber="47">B</emphasis>
. Mx1, respiratory plate (INV.159248).
<emphasis id="40C26A3E451BFF92FEE71CF5FECD2AE6" bold="true" box="[305,323,1979,2003]" pageId="46" pageNumber="47">C</emphasis>
. MdPalp (INV.159244). Scale bars. 1 (B), 2 (A), 3 (C) = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A4451AFF93FF411CF3FD5D2ACF" pageId="47" pageNumber="48" startId="47.[151,250,1981,2005]" targetBox="[197,1421,196,1942]" targetPageId="47" targetType="figure">
<paragraph id="7209B62C451AFF93FF411CF3FD5D2ACF" blockId="47.[151,1436,1981,2042]" pageId="47" pageNumber="48">
<emphasis id="40C26A3E451AFF93FF411CF3FEAB2AE0" bold="true" box="[151,293,1981,2005]" pageId="47" pageNumber="48">FIGURE 29.</emphasis>
<taxonomicName id="B5B6CDAF451AFF93FEE61CF3FD812AE0" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[304,527,1981,2005]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="cebios" status="gen. et spec. nov.">
<emphasis id="40C26A3E451AFF93FEE61CF3FD812AE0" box="[304,527,1981,2005]" italics="true" pageId="47" pageNumber="48">Benincandona cebios</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745451AFF93FDCC1CF0FD552AE0" box="[538,731,1982,2006]" pageId="47" pageNumber="48" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E451AFF93FCB51CF3FCFB2AE0" bold="true" box="[867,885,1981,2005]" pageId="47" pageNumber="48">A</emphasis>
. T3 (INV.159244).
<emphasis id="40C26A3E451AFF93FB9B1CF0FBD02AE3" bold="true" box="[1101,1118,1982,2006]" pageId="47" pageNumber="48">B</emphasis>
. T2 (INV.159244).
<emphasis id="40C26A3E451AFF93FAE01CF3FAC62AE0" bold="true" box="[1334,1352,1981,2005]" pageId="47" pageNumber="48">C</emphasis>
. CRAtt (INV.159244).
<emphasis id="40C26A3E451AFF93FEE71CACFECD2ACF" bold="true" box="[305,323,2018,2042]" pageId="47" pageNumber="48">D</emphasis>
. CR (INV.159244). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A44505FF8CFF411CD9FDB32ACD" pageId="48" pageNumber="49" startId="48.[151,250,1943,1967]" targetBox="[194,1393,191,1910]" targetPageId="48" targetType="figure">
<paragraph id="7209B62C4505FF8CFF411CD9FDB32ACD" blockId="48.[151,1437,1943,2040]" pageId="48" pageNumber="49">
<emphasis id="40C26A3E4505FF8CFF411CD9FEAD2A9A" bold="true" box="[151,291,1943,1967]" pageId="48" pageNumber="49">FIGURE 30.</emphasis>
<taxonomicName id="B5B6CDAF4505FF8CFEFA1CD6FD872A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[300,521,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="cebios" status="gen. et spec. nov.">
<emphasis id="40C26A3E4505FF8CFEFA1CD6FD872A85" box="[300,521,1944,1968]" italics="true" pageId="48" pageNumber="49">Benincandona cebios</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454505FF8CFDC41CD6FD432A9A" box="[530,717,1944,1968]" pageId="48" pageNumber="49" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E4505FF8CFC981CD9FCEE2A9A" bold="true" box="[846,864,1943,1967]" pageId="48" pageNumber="49">A</emphasis>
. Zenker organ (INV.159247).
<emphasis id="40C26A3E4505FF8CFB771CD6FB5F2A85" bold="true" box="[1185,1233,1944,1968]" pageId="48" pageNumber="49">BD</emphasis>
. Hp (INV.159244).
<emphasis id="40C26A3E4505FF8CFF411CF2FF262AE1" bold="true" box="[151,168,1980,2004]" pageId="48" pageNumber="49">B</emphasis>
. External view (drawing).
<emphasis id="40C26A3E4505FF8CFE141CF5FE5A2AE6" bold="true" box="[450,468,1979,2003]" pageId="48" pageNumber="49">C</emphasis>
. Internal view (photograph without interference contrast).
<emphasis id="40C26A3E4505FF8CFBEE1CF2FBC42AE1" bold="true" box="[1080,1098,1980,2004]" pageId="48" pageNumber="49">D</emphasis>
. Internal view (photograph with interference contrast). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A44504FF8DFF411CD9FBDC2AE1" pageId="49" pageNumber="50" startId="49.[151,250,1943,1967]" targetBox="[184,1406,184,1915]" targetPageId="49" targetType="figure">
<paragraph id="7209B62C4504FF8DFF411CD9FBDC2AE1" blockId="49.[151,1437,1943,2004]" pageId="49" pageNumber="50">
<emphasis id="40C26A3E4504FF8DFF411CD9FEAE2A9A" bold="true" box="[151,288,1943,1967]" pageId="49" pageNumber="50">FIGURE 31.</emphasis>
<taxonomicName id="B5B6CDAF4504FF8DFEF11CD6FD8F2A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[295,513,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="cebios" status="gen. et spec. nov.">
<emphasis id="40C26A3E4504FF8DFEF11CD6FD8F2A85" box="[295,513,1944,1968]" italics="true" pageId="49" pageNumber="50">Benincandona cebios</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454504FF8DFDD11CD6FD342A9A" box="[519,698,1944,1968]" pageId="49" pageNumber="50" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (AB ♂ and CE ♀).
<emphasis id="40C26A3E4504FF8DFC351CD9FC7B2A9A" bold="true" box="[995,1013,1943,1967]" pageId="49" pageNumber="50">C</emphasis>
. A2 (INV.159245).
<emphasis id="40C26A3E4504FF8DFB171CD6FB5D2A85" bold="true" box="[1217,1235,1944,1968]" pageId="49" pageNumber="50">D</emphasis>
. CR (INV.159245).
<emphasis id="40C26A3E4504FF8DFF411CF2FF262AE1" bold="true" box="[151,168,1980,2004]" pageId="49" pageNumber="50">E</emphasis>
. T1 (INV.159245).
<emphasis id="40C26A3E4504FF8DFEA61CF5FE0C2AE6" bold="true" box="[368,386,1979,2003]" pageId="49" pageNumber="50">A</emphasis>
. T1, Lpp (INV.159244).
<emphasis id="40C26A3E4504FF8DFD561CF2FD1F2AE1" bold="true" box="[640,657,1980,2004]" pageId="49" pageNumber="50">B</emphasis>
. T1, Rpp (INV.159244). Scale bar = 50 µm.
</paragraph>
</caption>
<paragraph id="7209B62C4507FF8EFF111BD9FC692C50" blockId="50.[151,1437,151,1401]" pageId="50" pageNumber="51">
T2 (
<figureCitation id="EA8DAAA94507FF8EFF231BD9FEF92D84" box="[245,375,151,177]" captionStart="FIGURE 29" captionStartId="47.[151,250,1981,2005]" captionTargetBox="[197,1421,196,1942]" captionTargetId="figure-17@47.[166,1422,181,1957]" captionTargetPageId="47" captionText="FIGURE 29. Benincandona cebios gen. et spec. nov., limbs (♂). A. T3 (INV.159244). B. T2 (INV.159244). C. CRAtt (INV.159244). D. CR (INV.159244). Scale bar = 50 µm." pageId="50" pageNumber="51">Figure 29B</figureCitation>
) with 4-segmented endopodite (penultimate segment divided), generally elongated. First basal segment without setae d1 and d2. First segment of endopodite elongated (length almost 4x the basal width), without seta. Second endopodal segment (segment 2A), with length
<emphasis id="40C26A3E4507FF8EFCEC1B91FCC32DCD" box="[826,845,223,248]" italics="true" pageId="50" pageNumber="51">c.</emphasis>
3x basal width, set with one short ventro-apical seta f. Third endopodal segment (segment 2B) of about the same length/width ratio as the previous segment, ventro-subapically set with a g-seta. Terminal segment about 1.5 times as long as basal width, apically with two short setae (h1 and h3) and a long and slender claw (h2), distally set with short setulae.
</paragraph>
<paragraph id="7209B62C4507FF8EFF111A21FF642F08" blockId="50.[151,1437,151,1401]" pageId="50" pageNumber="51">
T3 (
<figureCitation id="EA8DAAA94507FF8EFF211A21FEF02CBC" box="[247,382,367,393]" captionStart="FIGURE 29" captionStartId="47.[151,250,1981,2005]" captionTargetBox="[197,1421,196,1942]" captionTargetId="figure-17@47.[166,1422,181,1957]" captionTargetPageId="47" captionText="FIGURE 29. Benincandona cebios gen. et spec. nov., limbs (♂). A. T3 (INV.159244). B. T2 (INV.159244). C. CRAtt (INV.159244). D. CR (INV.159244). Scale bar = 50 µm." pageId="50" pageNumber="51">Figure 29A</figureCitation>
) with first segment carrying only one subapical seta d1, of medium length. Second (knee-) segment with short seta d2. Third segment about 5x as long as wide, not carrying any seta. Fourth segment narrower and at least 5x basal width, carrying one short subapical seta g. Terminal segment well-separated from penultimate segment, length about twice as long as basal width, carrying three setae: one long and reflexed (h3), almost reaching knee-segment and two short ventrally directed and curved setae h1 &amp; h2, the latter slightly exceeding tip of the former.
</paragraph>
<paragraph id="7209B62C4507FF8EFF111909FC862F9C" blockId="50.[151,1437,151,1401]" pageId="50" pageNumber="51">
CR (
<figureCitation id="EA8DAAA94507FF8EFF2C1909FEF32F54" box="[250,381,583,609]" captionStart="FIGURE 29" captionStartId="47.[151,250,1981,2005]" captionTargetBox="[197,1421,196,1942]" captionTargetId="figure-17@47.[166,1422,181,1957]" captionTargetPageId="47" captionText="FIGURE 29. Benincandona cebios gen. et spec. nov., limbs (♂). A. T3 (INV.159244). B. T2 (INV.159244). C. CRAtt (INV.159244). D. CR (INV.159244). Scale bar = 50 µm." pageId="50" pageNumber="51">Figure 29D</figureCitation>
) with stout ramus; claw Ga well-developed, stout with length slightly exceeding half the length of ramus, setae Sp, Sa and claw Gp reduced to small setae; seta Sp situated about at one-fifth length of ramus from distal end, Sa dorso-subapical and Gp ventro-subapical.
</paragraph>
<paragraph id="7209B62C4507FF8EFF1119FDFA832FF8" blockId="50.[151,1437,151,1401]" box="[199,1293,691,717]" pageId="50" pageNumber="51">
CRAtt (
<figureCitation id="EA8DAAA94507FF8EFEC919FDFE2F2FF8" box="[287,417,691,717]" captionStart="FIGURE 29" captionStartId="47.[151,250,1981,2005]" captionTargetBox="[197,1421,196,1942]" captionTargetId="figure-17@47.[166,1422,181,1957]" captionTargetPageId="47" captionText="FIGURE 29. Benincandona cebios gen. et spec. nov., limbs (♂). A. T3 (INV.159244). B. T2 (INV.159244). C. CRAtt (INV.159244). D. CR (INV.159244). Scale bar = 50 µm." pageId="50" pageNumber="51">Figure 29C</figureCitation>
) long and stout, distally bifurcated, subapically with additional lateral branch.
</paragraph>
<paragraph id="7209B62C4507FF8EFF111999FD772E0C" blockId="50.[151,1437,151,1401]" pageId="50" pageNumber="51">
Hp (
<figureCitation id="EA8DAAA94507FF8EFF2C1999FE2E2FC4" box="[250,416,727,753]" captionStart="FIGURE 30" captionStartId="48.[151,250,1943,1967]" captionTargetBox="[194,1393,191,1910]" captionTargetId="figure-31@48.[184,1403,181,1920]" captionTargetPageId="48" captionText="FIGURE 30. Benincandona cebios gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159247). BD. Hp (INV.159244). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." pageId="50" pageNumber="51">Figure 30BD</figureCitation>
) with robust lobe ls, apically rounded, reaching well beyond symmetrically rounded lobe ms, both lobes overlapping, not diverging; lobe h shorter and sub-rectangular; internal anatomy (
<figureCitation id="EA8DAAA94507FF8EFB3819B5FA1A2E20" box="[1262,1428,763,789]" captionStart="FIGURE 30" captionStartId="48.[151,250,1943,1967]" captionTargetBox="[194,1393,191,1910]" captionTargetId="figure-31@48.[184,1403,181,1920]" captionTargetPageId="48" captionText="FIGURE 30. Benincandona cebios gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159247). BD. Hp (INV.159244). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." pageId="50" pageNumber="51">Figure 30C, D</figureCitation>
) with postlabyrinthal spermiduct with one circular coil.
</paragraph>
<paragraph id="7209B62C4507FF8EFF11180DFBDD2E68" blockId="50.[151,1437,151,1401]" box="[199,1107,835,861]" pageId="50" pageNumber="51">
Zenker organ (
<figureCitation id="EA8DAAA94507FF8EFEBD180DFE612E68" box="[363,495,835,861]" captionStart="FIGURE 30" captionStartId="48.[151,250,1943,1967]" captionTargetBox="[194,1393,191,1910]" captionTargetId="figure-31@48.[184,1403,181,1920]" captionTargetPageId="48" captionText="FIGURE 30. Benincandona cebios gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159247). BD. Hp (INV.159244). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." pageId="50" pageNumber="51">Figure 30A</figureCitation>
) long and slightly broad, with
<emphasis id="40C26A3E4507FF8EFC91180DFCD52E69" box="[839,859,835,860]" italics="true" pageId="50" pageNumber="51">c.</emphasis>
seven spinous whirls.
</paragraph>
<paragraph id="7209B62C4507FF8EFF111828FE482EB5" blockId="50.[151,1437,151,1401]" box="[199,454,870,896]" pageId="50" pageNumber="51">
<emphasis id="40C26A3E4507FF8EFF111828FE482EB5" bold="true" box="[199,454,870,896]" pageId="50" pageNumber="51">Description of female</emphasis>
</paragraph>
<paragraph id="7209B62C4507FF8EFF1118C5FB422E90" blockId="50.[151,1437,151,1401]" box="[199,1228,907,933]" pageId="50" pageNumber="51">
Cp and valves (
<figureCitation id="EA8DAAA94507FF8EFEA218C5FD952E90" box="[372,539,907,933]" captionStart="FIGURE 24" captionStartId="41.[151,250,1944,1968]" captionTargetBox="[170,1417,181,1921]" captionTargetId="figure-17@41.[170,1417,181,1921]" captionTargetPageId="41" captionText="FIGURE 24. Benincandona cebios gen. et spec. nov., valves (AC, JK ♂; DI ♀). AI. CpRl (AC. INV.159255, DI. INV.159258). J. INV.159256. CpD. K. INV.159257. CpV. Scale bars. 1 (B, E, J, K) = 500 µm; 2 (A, C, D, F, H, I) = 200 µm." pageId="50" pageNumber="51">Figures 24DI</figureCitation>
,
<figureCitation id="EA8DAAA94507FF8EFDF118C5FDF72E90" box="[551,633,907,933]" captionStart="FIGURE 26" captionStartId="44.[151,250,1673,1697]" captionTargetBox="[151,1436,181,1649]" captionTargetId="figure-31@44.[151,1436,181,1649]" captionTargetPageId="44" captionText="FIGURE 26. Benincandona cebios gen. et spec. nov., valves (♀). A, C, D, G. LVi (INV.159253). B, E, F. RVi (INV.159253). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." pageId="50" pageNumber="51">26AG</figureCitation>
) as in the male, without obvious sexual dimorphism.
</paragraph>
<paragraph id="7209B62C4507FF8EFF1118E1FB072EFC" blockId="50.[151,1437,151,1401]" box="[199,1161,943,969]" pageId="50" pageNumber="51">
Limbs (
<figureCitation id="EA8DAAA94507FF8EFECB18E1FE302EFC" box="[285,446,943,969]" captionStart="FIGURE 31" captionStartId="49.[151,250,1943,1967]" captionTargetBox="[184,1406,184,1915]" captionTargetId="figure-17@49.[180,1407,181,1919]" captionTargetPageId="49" captionText="FIGURE 31. Benincandona cebios gen. et spec. nov., limbs (AB ♂ and CE ♀). C. A2 (INV.159245). D. CR (INV.159245). E. T1 (INV.159245). A. T1, Lpp (INV.159244). B. T1, Rpp (INV.159244). Scale bar = 50 µm." pageId="50" pageNumber="51">Figure 31CE</figureCitation>
) largely as in the male, but with sexually dimorphic A2 and T1.
</paragraph>
<paragraph id="7209B62C4507FF8EFF11189DFB53296C" blockId="50.[151,1437,151,1401]" pageId="50" pageNumber="51">
A2 (
<figureCitation id="EA8DAAA94507FF8EFF2F189DFEF22ED8" box="[249,380,979,1005]" captionStart="FIGURE 31" captionStartId="49.[151,250,1943,1967]" captionTargetBox="[184,1406,184,1915]" captionTargetId="figure-17@49.[180,1407,181,1919]" captionTargetPageId="49" captionText="FIGURE 31. Benincandona cebios gen. et spec. nov., limbs (AB ♂ and CE ♀). C. A2 (INV.159245). D. CR (INV.159245). E. T1 (INV.159245). A. T1, Lpp (INV.159244). B. T1, Rpp (INV.159244). Scale bar = 50 µm." pageId="50" pageNumber="51">Figure 31C</figureCitation>
) with second endopodal segment undivided, with t1-seta (long, narrow and plumose) and t2- seta (thin and smooth); apical chaetotaxy (
<figureCitation id="EA8DAAA94507FF8EFDA418B9FD6C2924" box="[626,738,1015,1041]" captionStart="FIGURE 40" captionStartId="60.[151,250,1516,1540]" captionTargetBox="[151,1435,181,1492]" captionTargetId="figure-31@60.[151,1435,181,1492]" captionTargetPageId="60" captionText="FIGURE 40. Diagrams of apical views of the A2 of Benincandona, showing position of (sub-) terminal claws, setae and aesthetascs of the terminal (small circle) and penultimate segment (large circle). For explanations of codes, see Material and Methods (Abbreviations used in text and figures)." pageId="50" pageNumber="51">Figure 40</figureCitation>
) with z1 claw-like, z2 and z3 being both setae, z1 about twice as long as z2, and z3 slightly shorter than z2; claws G1, G2, G3 and GM all reaching about the same point; seta g and aesthetasc y3 as in the male. Small claw Gm in female homologous to large claw in the male.
</paragraph>
<paragraph id="7209B62C4507FF8EFF111F2DFBDF29F0" blockId="50.[151,1437,151,1401]" pageId="50" pageNumber="51">
T1 (
<figureCitation id="EA8DAAA94507FF8EFF221F2DFEFA2948" box="[244,372,1123,1149]" captionStart="FIGURE 31" captionStartId="49.[151,250,1943,1967]" captionTargetBox="[184,1406,184,1915]" captionTargetId="figure-17@49.[180,1407,181,1919]" captionTargetPageId="49" captionText="FIGURE 31. Benincandona cebios gen. et spec. nov., limbs (AB ♂ and CE ♀). C. A2 (INV.159245). D. CR (INV.159245). E. T1 (INV.159245). A. T1, Lpp (INV.159244). B. T1, Rpp (INV.159244). Scale bar = 50 µm." pageId="50" pageNumber="51">Figure 31E</figureCitation>
) with basal part with
<emphasis id="40C26A3E4507FF8EFDB21F2DFDF62949" box="[612,632,1123,1148]" italics="true" pageId="50" pageNumber="51">c.</emphasis>
10 distal setae of unequal length and morphology, setae d and b present, with long additional seta (d) present close to seta d; respiratory plate represented by two long and one short rays. Endopodite with a broad palp, carrying three subequal but short apical setae (h1h3).
</paragraph>
<paragraph id="7209B62C4507FF8EFF111F81FD9E29DC" blockId="50.[151,1437,151,1401]" box="[199,528,1231,1257]" pageId="50" pageNumber="51">
<emphasis id="40C26A3E4507FF8EFF111F81FE0E29DC" bold="true" box="[199,384,1231,1257]" pageId="50" pageNumber="51">Measurements:</emphasis>
See
<tableCitation id="3F3483974507FF8EFE601F81FD8429DC" box="[438,522,1231,1257]" captionStart="TABLE 3" captionStartId="19.[152,244,1592,1616]" captionText="TABLE 3. Measurements of illustrated specimens of Benincandona gen. et spec. nov., using SEM." pageId="50" pageNumber="51">Table 3</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74507FF8EFF111FBCFDEA2838" box="[199,612,1266,1293]" pageId="50" pageNumber="51" type="diagnosis">
<paragraph id="7209B62C4507FF8EFF111FBCFDEA2838" blockId="50.[151,1437,151,1401]" box="[199,612,1266,1293]" pageId="50" pageNumber="51">
<emphasis id="40C26A3E4507FF8EFF111FBCFE5B2839" bold="true" box="[199,469,1266,1292]" pageId="50" pageNumber="51">Differential Diagnosis:</emphasis>
See
<tableCitation id="3F3483974507FF8EFDDC1FBDFDEE2838" box="[522,608,1267,1293]" captionStart="TABLE 4" captionStartId="20.[152,245,1665,1689]" captionText="TABLE 4. Differential character states between the five new species of Benincandona gen. nov." pageId="50" pageNumber="51">Table 4</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74507FF8EFF111E59FEE8284C" pageId="50" pageNumber="51" type="biology_ecology">
<paragraph id="7209B62C4507FF8EFF111E59FE612805" blockId="50.[151,1437,151,1401]" box="[199,495,1302,1329]" pageId="50" pageNumber="51">
<emphasis id="40C26A3E4507FF8EFF111E59FE612805" bold="true" box="[199,495,1302,1329]" pageId="50" pageNumber="51">Ecology and distribution</emphasis>
</paragraph>
<paragraph id="7209B62C4507FF8EFF111E75FEE8284C" blockId="50.[151,1437,151,1401]" pageId="50" pageNumber="51">
This subterranean species is presently known from its
<typeStatus id="AD0D088E4507FF8EFCFE1E75FCD62860" box="[808,856,1339,1365]" pageId="50" pageNumber="51">type</typeStatus>
locality and from a second, closely situated station, only (see
<tableCitation id="3F3483974507FF8EFED21E11FEDB284C" box="[260,341,1375,1401]" captionStart="TABLE 1" captionStartId="5.[152,244,929,955]" captionText="TABLE 1. Localities where Benincandona gen. nov. sp. nov. were collected, coordinates and hydro-morphological characteristics. In all stations, the water was clear and was used for both drinking water and household purposes by the riparian villages." pageId="50" pageNumber="51">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="C9591F3676E6CC347AEEAAFD4FD850CA" ID-DOI="10.11646/zootaxa.5503.1.1" ID-ISSN="1175-5334" ID-ZooBank="7385E4AA-D7AD-4E79-A05C-684319E0843B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726051594773" checkinUser="plazi" docAuthor="Hotèkpo, Sourou Joseph, Schön, Isa &amp; Martens, Koen" docDate="2024" docId="FA1F073A4520FFA1FF41190AFB072994" docLanguage="en" docName="zootaxa.5503.1.1.pdf" docOrigin="Zootaxa 5503 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5503.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Benincandona ibikounlei Hotèkpo, Schön &amp; Martens, 2024, gen. et spec. nov." docType="treatment" docVersion="1" lastPageNumber="30" masterDocId="06267F424535FFBCFFD61B4EFF8E2D35" masterDocTitle="An endemic species flock of Candonidae Kaufmann, 1900 (Crustacea, Ostracoda) from wells in Benin (Africa), with the description of a new subfamily, a new genus and five new species" masterLastPageNumber="72" masterPageNumber="1" pageNumber="22" updateTime="1726051738210" updateUser="GgImagineBatch">
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<mods:title id="F4A5C34D890A7CC3606D137ED0E0596E">An endemic species flock of Candonidae Kaufmann, 1900 (Crustacea, Ostracoda) from wells in Benin (Africa), with the description of a new subfamily, a new genus and five new species</mods:title>
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<mods:namePart id="7A7D910B20BE6C16FF9C13C9F6FACE68">Hotèkpo, Sourou Joseph</mods:namePart>
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<mods:affiliation id="D177CD0EE82ABBAD178EF2BF6578D064">Centre de Recherche pour la lutte contre les Maladies Infectieuses Tropicales (CReMIT / TIDRC), Université dAbomey-Calavi, Bénin jshotekpo @ gmail. com; https: // orcid. org / 0000 - 0001 - 9091 - 433 X &amp; Royal Belgian Institute of Natural Sciences, Freshwater Biology, Vautierstraat 29, B- 1000 Brussels, Belgium &amp; University of Hasselt, Research Group Zoology, Agoralaan Building D, B- 3590 Diepenbeek, Belgium ischoen @ naturalsciences. be; https: // orcid. org / 0000 - 0001 - 9269 - 6487</mods:affiliation>
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<mods:namePart id="678B603F492EB848F3F59A7CD4640B56">Schön, Isa</mods:namePart>
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<mods:affiliation id="B76DA3B65063E8F8076574034E09039B">Royal Belgian Institute of Natural Sciences, Freshwater Biology, Vautierstraat 29, B- 1000 Brussels, Belgium &amp; University of Hasselt, Research Group Zoology, Agoralaan Building D, B- 3590 Diepenbeek, Belgium ischoen @ naturalsciences. be; https: // orcid. org / 0000 - 0001 - 9269 - 6487</mods:affiliation>
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<mods:namePart id="8002A68533615FDF3878931081550E81">Martens, Koen</mods:namePart>
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<mods:affiliation id="4D3EDAA4BA7E6C1B95B4F534B4C4B206">Royal Belgian Institute of Natural Sciences, Freshwater Biology, Vautierstraat 29, B- 1000 Brussels, Belgium &amp; Ghent University, Dept. Biology, K. L. Ledeganckstraat 35, B- 9000 Ghent, Belgium kmartens @ naturalsciences. be; https: // orcid. org / 0000 - 0001 - 8680 - 973 X * Corresponding author</mods:affiliation>
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<taxonomicName id="B5B6CDAF4520FFA9FF41190AFE442F6A" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[151,458,580,607]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="ibikounlei" status="gen. et spec. nov.">
<emphasis id="40C26A3E4520FFA9FF41190AFE442F6A" bold="true" box="[151,458,580,607]" italics="true" pageId="21" pageNumber="22">Benincandona ibikounlei</emphasis>
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Hotèkpo &amp; Martens
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<paragraph id="7209B62C4520FFA9FF4119C3FE2D2F92" blockId="21.[151,980,580,680]" box="[151,419,653,680]" pageId="21" pageNumber="22">
(
<figureCitation id="EA8DAAA94520FFA9FF4919C3FECA2F9D" box="[159,324,653,680]" captionStart-0="FIGURE 10" captionStart-1="FIGURE 11" captionStart-2="FIGURE 12" captionStart-3="FIGURE 13" captionStart-4="FIGURE 14" captionStart-5="FIGURE 15" captionStartId-0="21.[151,250,1855,1879]" captionStartId-1="22.[151,250,1943,1967]" captionStartId-2="25.[151,250,1981,2005]" captionStartId-3="26.[151,250,1943,1967]" captionStartId-4="27.[151,250,1943,1967]" captionStartId-5="28.[151,250,1943,1967]" captionTargetBox-0="[151,1436,883,1832]" captionTargetBox-1="[183,1405,185,1915]" captionTargetBox-2="[166,1422,183,1955]" captionTargetBox-3="[191,1395,185,1916]" captionTargetBox-4="[229,1359,191,1909]" captionTargetBox-5="[176,1407,181,1915]" captionTargetId-0="figure-159@21.[151,1436,883,1832]" captionTargetId-1="figure-31@22.[179,1407,181,1919]" captionTargetId-2="figure-17@25.[166,1422,181,1957]" captionTargetId-3="figure-31@26.[187,1399,181,1919]" captionTargetId-4="figure-17@27.[219,1369,181,1919]" captionTargetId-5="figure-31@28.[176,1411,181,1919]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionTargetPageId-2="25" captionTargetPageId-3="26" captionTargetPageId-4="27" captionTargetPageId-5="28" captionText-0="FIGURE 10. Benincandona ibikounlei gen. et spec. nov., valves. A. INV.159222. CpRl. B. INV.159222. CpV. C. INV.159223. CpD. D. INV.159224. LVi. E. INV.159224. RVi. Scale bar = 500 µm." captionText-1="FIGURE 11. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. A2 (INV.159219, INV.159217, INV.159218). B. A1 (INV.159219, INV.159218). C, D. MdCoxa, (INV.159218). Scale bar = 50 µm." captionText-2="FIGURE 12. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. Mx1 (Chaetotaxy incomplete) (INV.159218). B. Mx1, respiratory plate (INV.159218). C. MdPalp (INV.159218). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." captionText-3="FIGURE 13. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. CR (INV.159219). B. CRAtt (INV.159219). C. T3 (INV.159219). D. T2 (INV.159217). Scale bar = 50 µm." captionText-4="FIGURE 14. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159219). BD. Hp (INV.159219). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." captionText-5="FIGURE 15. Benincandona ibikounlei gen. et spec. nov., limbs (AB ♂ and CE ♀). A. T1, Lpp (INV.159219). B. T1, Rpp (INV.159219). C. A2 (INV.159216). D. CR (INV.159216). E. T1 (INV.159221). Scale bar = 50 µm." pageId="21" pageNumber="22">
Figures 10
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,
<figureCitation id="EA8DAAA94520FFA9FE8619C3FE122F92" box="[336,412,653,679]" captionStart="FIGURE 41" captionStartId="61.[151,250,1113,1137]" captionTargetBox="[151,1436,181,1089]" captionTargetId="figure-17@61.[151,1436,181,1089]" captionTargetPageId="61" captionText="FIGURE 41. Benincandona gen. et spec. nov., CpRl. (A, C, E, F, G, I, K, N, O, Q ♂ and B, D, H, J, L, M, P, R ♀). AD. B. martini. EF. B. ibikounlei. GJ. B. sakitii. KN. B. cebios. OR. B. trapezoides. Scale bar = 500 µm." pageId="21" pageNumber="22">41E, F</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74520FFA9FF41199BFEB92E6E" pageId="21" pageNumber="22" type="etymology">
<paragraph id="7209B62C4520FFA9FF41199BFE962FDA" blockId="21.[151,1436,725,859]" box="[151,280,725,751]" pageId="21" pageNumber="22">
<emphasis id="40C26A3E4520FFA9FF41199BFE962FDA" bold="true" box="[151,280,725,751]" pageId="21" pageNumber="22">Etymology</emphasis>
</paragraph>
<paragraph id="7209B62C4520FFA9FF4119B7FEB92E6E" blockId="21.[151,1436,725,859]" pageId="21" pageNumber="22">
The species is named after Prof. Moudachirou Ibikounlé (Université dAbomey Calavi,
<collectingCountry id="0AA1F6BC4520FFA9FB5E19B7FB432E26" box="[1160,1229,761,787]" name="Benin" pageId="21" pageNumber="22">Bénin</collectingCountry>
) in recognition of his contribution to the advancement of scientific research in
<collectingCountry id="0AA1F6BC4520FFA9FCEF1853FCF12E02" box="[825,895,797,823]" name="Benin" pageId="21" pageNumber="22">Benin</collectingCountry>
, and in gratitude for his continued support to the career of SJH.
</paragraph>
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<paragraph id="7209B62C4520FFA9FF411C71FCDB2A49" blockId="21.[151,1437,1855,1916]" pageId="21" pageNumber="22">
<emphasis id="40C26A3E4520FFA9FF411C71FEAE2A62" bold="true" box="[151,288,1855,1879]" pageId="21" pageNumber="22">FIGURE 10.</emphasis>
<taxonomicName id="B5B6CDAF4520FFA9FEF01C71FDAD2A62" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[294,547,1855,1879]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="ibikounlei" status="gen. et spec. nov.">
<emphasis id="40C26A3E4520FFA9FEF01C71FDAD2A62" box="[294,547,1855,1879]" italics="true" pageId="21" pageNumber="22">Benincandona ibikounlei</emphasis>
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<taxonomicNameLabel id="5BF1D7454520FFA9FDFC1C0EFD552A62" box="[554,731,1856,1880]" pageId="21" pageNumber="22" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, valves.
<emphasis id="40C26A3E4520FFA9FCE41C71FCCA2A62" bold="true" box="[818,836,1855,1879]" pageId="21" pageNumber="22">A</emphasis>
. INV.159222. CpRl.
<emphasis id="40C26A3E4520FFA9FBCA1C71FBA32A62" bold="true" box="[1052,1069,1855,1879]" pageId="21" pageNumber="22">B</emphasis>
. INV.159222. CpV.
<emphasis id="40C26A3E4520FFA9FB2D1C71FA832A62" bold="true" box="[1275,1293,1855,1879]" pageId="21" pageNumber="22">C</emphasis>
. INV.159223. CpD.
<emphasis id="40C26A3E4520FFA9FF021C2DFF682A4E" bold="true" box="[212,230,1891,1915]" pageId="21" pageNumber="22">D</emphasis>
. INV.159224. LVi.
<emphasis id="40C26A3E4520FFA9FE7B1C2DFE302A4E" bold="true" box="[429,446,1891,1915]" pageId="21" pageNumber="22">E</emphasis>
. INV.159224. RVi. Scale bar = 500 µm.
</paragraph>
</caption>
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<paragraph id="7209B62C4523FFAAFF411CD9FC362AE1" blockId="22.[151,1436,1943,2004]" pageId="22" pageNumber="23">
<emphasis id="40C26A3E4523FFAAFF411CD9FEAE2A9A" bold="true" box="[151,288,1943,1967]" pageId="22" pageNumber="23">FIGURE 11.</emphasis>
<taxonomicName id="B5B6CDAF4523FFAAFEFF1CD6FDA62A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[297,552,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="ibikounlei" status="gen. et spec. nov.">
<emphasis id="40C26A3E4523FFAAFEFF1CD6FDA62A85" box="[297,552,1944,1968]" italics="true" pageId="22" pageNumber="23">Benincandona ibikounlei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454523FFAAFDE71CD6FD662A9A" box="[561,744,1944,1968]" pageId="22" pageNumber="23" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E4523FFAAFCB11CD9FCF72A9A" bold="true" box="[871,889,1943,1967]" pageId="22" pageNumber="23">A</emphasis>
. A2 (INV.159219, INV.159217, INV.159218).
<emphasis id="40C26A3E4523FFAAFA891CD6FAFE2A85" bold="true" box="[1375,1392,1944,1968]" pageId="22" pageNumber="23">B</emphasis>
. A1 (INV.159219, INV.159218).
<emphasis id="40C26A3E4523FFAAFE6C1CF5FE652AE1" bold="true" box="[442,491,1979,2004]" pageId="22" pageNumber="23">C, D</emphasis>
. MdCoxa, (INV.159218). Scale bar = 50 µm.
</paragraph>
</caption>
<subSubSection id="3AACE5A74522FFABFF411BD8FD132CBC" pageId="23" pageNumber="24" type="diagnosis">
<paragraph id="7209B62C4522FFABFF411BD8FE842D85" blockId="23.[151,1436,150,393]" box="[151,266,150,176]" pageId="23" pageNumber="24">
<emphasis id="40C26A3E4522FFABFF411BD8FE842D85" bold="true" box="[151,266,150,176]" pageId="23" pageNumber="24">Diagnosis</emphasis>
</paragraph>
<paragraph id="7209B62C4522FFABFF411BF5FD132CBC" blockId="23.[151,1436,150,393]" pageId="23" pageNumber="24">CpRl elongated (L/H = 2.4), sub-triangular, with greatest height slightly in front of the middle; LV dorsally extending beyond RV, anterior and posterior margins evenly rounded, valve surface smooth. CpD and CpV narrow, with greatest width in the middle and angular lateral sides, anterior and posterior extremities pointed. A2 with seta X short, not reaching base of aesthetasc Y. Second palp segment of Mx1 with basal W almost equalling L. T3 with seta d2 absent. Hp with ls and ms with slightly diverging positions, narrower lobe ls on Hp with skewed rectangular distal margin, extending well beyond lobe ms.</paragraph>
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<paragraph id="7209B62C4522FFABFF411AF9FEB02CE5" blockId="23.[151,318,438,465]" box="[151,318,438,465]" pageId="23" pageNumber="24">
<emphasis id="40C26A3E4522FFABFF411AF9FEB02CE5" bold="true" box="[151,318,438,465]" pageId="23" pageNumber="24">Type material</emphasis>
</paragraph>
<paragraph id="7209B62C4522FFABFF411AB1FE8A2F2C" blockId="23.[151,1394,511,573]" box="[151,260,511,537]" pageId="23" pageNumber="24">
<emphasis id="40C26A3E4522FFABFF411AB1FE8A2F2C" bold="true" box="[151,260,511,537]" pageId="23" pageNumber="24">
<typeStatus id="AD0D088E4522FFABFF411AB1FE8A2F2C" box="[151,260,511,537]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7209B62C4522FFABFF41196DFAFC2F08" blockId="23.[151,1394,511,573]" box="[151,1394,546,573]" pageId="23" pageNumber="24">
<materialsCitation id="C2DEBC714522FFABFF41196DFAFC2F08" box="[151,1394,546,573]" collectionCode="INV" pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="64B07DA54522FFABFF41196DFF4D2F08" box="[151,195,547,573]" count="1" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
, specimen with soft parts dissected in glycerine in a sealed slide, and with valves stored dry (
<collectionCode id="14A72EE94522FFABFB01196DFA892F08" box="[1239,1287,547,573]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="23" pageNumber="24" type="Herbarium">INV</collectionCode>
.159215).
</materialsCitation>
</paragraph>
<paragraph id="7209B62C4522FFABFF411924FF722FB1" blockId="23.[151,1436,618,717]" box="[151,252,618,644]" pageId="23" pageNumber="24">
<emphasis id="40C26A3E4522FFABFF411924FF722FB1" bold="true" box="[151,252,618,644]" pageId="23" pageNumber="24">
<typeStatus id="AD0D088E4522FFABFF411924FF722FB1" box="[151,252,618,644]" pageId="23" pageNumber="24" type="allotype">Allotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7209B62C4522FFABFF4119C1FEE92FF8" blockId="23.[151,1436,618,717]" pageId="23" pageNumber="24">
<materialsCitation id="C2DEBC714522FFABFF4119C1FEE92FF8" collectionCode="INV" pageId="23" pageNumber="24" specimenCount="1" specimenCount-female="1" typeStatus="allotype">
<specimenCount id="64B07DA54522FFABFF4119C1FF4F2F9C" box="[151,193,655,681]" count="1" pageId="23" pageNumber="24" type="female">1 ♀</specimenCount>
, soft part dissected as the male (
<collectionCode id="14A72EE94522FFABFDFA19C0FDEF2F9C" box="[556,609,654,681]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="23" pageNumber="24" type="Herbarium">INV</collectionCode>
.159216). Valves lost: the decalcified valves could not be preserved during the manipulations.
</materialsCitation>
</paragraph>
<paragraph id="7209B62C4522FFABFF4119B5FE812E20" blockId="23.[151,1436,763,933]" box="[151,271,763,789]" pageId="23" pageNumber="24">
<emphasis id="40C26A3E4522FFABFF4119B5FE812E20" bold="true" box="[151,271,763,789]" pageId="23" pageNumber="24">
<typeStatus id="AD0D088E4522FFABFF4119B5FE812E20" box="[151,271,763,789]" pageId="23" pageNumber="24" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7209B62C4522FFABFF411851FEBF2E68" blockId="23.[151,1436,763,933]" pageId="23" pageNumber="24">
<materialsCitation id="C2DEBC714522FFABFF411851FC4A2E0C" box="[151,964,798,825]" collectionCode="INV" pageId="23" pageNumber="24" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="64B07DA54522FFABFF411851FF582E0C" box="[151,214,799,825]" count="3" pageId="23" pageNumber="24" type="male">3 ♂♂</specimenCount>
dissected as the
<typeStatus id="AD0D088E4522FFABFE591851FE7F2E0C" box="[399,497,799,825]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
(
<collectionCode id="14A72EE94522FFABFE291850FDBD2E0C" box="[511,563,798,825]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="23" pageNumber="24" type="Herbarium">INV</collectionCode>
.159217, INV.159218, INV.159219)
</materialsCitation>
;
<materialsCitation id="C2DEBC714522FFABFC181851FEA32E68" collectionCode="INV" pageId="23" pageNumber="24" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="64B07DA54522FFABFC181851FB832E0C" box="[974,1037,799,825]" count="2" pageId="23" pageNumber="24" type="female">2 ♀♀</specimenCount>
dissected as the male (
<collectionCode id="14A72EE94522FFABFADC1850FAB02E0C" box="[1290,1342,798,825]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="23" pageNumber="24" type="Herbarium">INV</collectionCode>
.159220, INV.159221)
</materialsCitation>
.
</paragraph>
<paragraph id="7209B62C4522FFABFF111829FE872E90" blockId="23.[151,1436,763,933]" pageId="23" pageNumber="24">
The
<typeStatus id="AD0D088E4522FFABFF291829FEEF2EB4" box="[255,353,871,897]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
male comprises both valves and dissected soft parts, but no illustrations are based on this specimen.
</paragraph>
<paragraph id="7209B62C4522FFABFF41189DFEBE2ED9" blockId="23.[151,1436,978,1113]" box="[151,304,978,1005]" pageId="23" pageNumber="24">
<emphasis id="40C26A3E4522FFABFF41189DFEBE2ED9" bold="true" box="[151,304,978,1005]" pageId="23" pageNumber="24">Type locality</emphasis>
</paragraph>
<paragraph id="7209B62C4522FFABFF4118B9FC4E296C" blockId="23.[151,1436,978,1113]" pageId="23" pageNumber="24">
<materialsCitation id="C2DEBC714522FFABFF4118B9FC32296C" collectingDate="2016-08-09" collectorName="Leg. &amp; S. J. Hotekpo &amp; M. Lagnika &amp; P. Martin" country="Benin" county="Colline Department" latitude="7.8160834" location="Tchetti" longLatPrecision="1" longitude="1.67185" municipality="Savalou District" pageId="23" pageNumber="24" specimenCount="1" stateProvince="Zou">
<collectingCountry id="0AA1F6BC4522FFABFF4118B9FF542924" box="[151,218,1015,1041]" name="Benin" pageId="23" pageNumber="24">Benin</collectingCountry>
,
<collectingCounty id="9B68CEA04522FFABFF3C18B9FE422924" box="[234,460,1015,1041]" pageId="23" pageNumber="24">Colline Department</collectingCounty>
,
<collectingMunicipality id="926D2C564522FFABFE0F18B9FD1C2924" box="[473,658,1015,1041]" pageId="23" pageNumber="24">Savalou District</collectingMunicipality>
, town of
<location id="7769E0F74522FFABFD2918B9FCC12924" LSID="urn:lsid:plazi:treatment:FA1F073A4520FFA1FF41190AFB072994:7769E0F74522FFABFD2918B9FCC12924" box="[767,847,1015,1041]" country="Benin" county="Colline Department" latitude="7.8160834" longLatPrecision="1" longitude="1.67185" municipality="Savalou District" name="Tchetti" pageId="23" pageNumber="24" stateProvince="Zou">Tchetti</location>
, located in the
<collectingRegion id="B07278CE4522FFABFC2918B9FBA22924" box="[1023,1068,1015,1041]" country="Benin" name="Zou" pageId="23" pageNumber="24">Zou</collectingRegion>
catchment area (tributary of the
<collectingRegion id="B07278CE4522FFABFF411F55FF662900" box="[151,232,1051,1077]" country="Benin" name="Oueme" pageId="23" pageNumber="24">Ouémé</collectingRegion>
catchment area), coordinates:
<geoCoordinate id="1782D0EB4522FFABFD9A1F55FD532900" box="[588,733,1050,1077]" degrees="7" direction="north" minutes="48.965" orientation="latitude" pageId="23" pageNumber="24" precision="1" value="7.8160834">7° 48.965N</geoCoordinate>
,
<geoCoordinate id="1782D0EB4522FFABFD3D1F55FCF92900" box="[747,887,1051,1077]" degrees="1" direction="east" minutes="40.311" orientation="longitude" pageId="23" pageNumber="24" precision="1" value="1.67185">1° 40.311E</geoCoordinate>
,
<collectorName id="DF43D3FA4522FFABFC501F55FC332900" box="[902,957,1051,1077]" pageId="23" pageNumber="24">Leg.</collectorName>
:
<collectorName id="DF43D3FA4522FFABFC1D1F55FBEF2900" box="[971,1121,1051,1077]" pageId="23" pageNumber="24">S.J. Hotèkpo</collectorName>
,
<collectorName id="DF43D3FA4522FFABFBB91F55FB782900" box="[1135,1270,1051,1077]" pageId="23" pageNumber="24">M. Lagnika</collectorName>
and
<collectorName id="DF43D3FA4522FFABFAE41F55FA122900" box="[1330,1436,1051,1077]" pageId="23" pageNumber="24">P. Martin</collectorName>
collected on
<date id="060890EC4522FFABFEF01F71FE56296C" box="[294,472,1087,1113]" pageId="23" pageNumber="24" value="2016-08-09">
<collectingDate id="164C69044522FFABFEF01F71FE56296C" box="[294,472,1087,1113]" pageId="23" pageNumber="24" value="2016-08-09">09 August 2016</collectingDate>
</date>
(Locality code BEN072 in
<tableCitation id="3F3483974522FFABFCDB1F71FCE5296C" box="[781,875,1087,1113]" captionStart="TABLE 1" captionStartId="5.[152,244,929,955]" captionText="TABLE 1. Localities where Benincandona gen. nov. sp. nov. were collected, coordinates and hydro-morphological characteristics. In all stations, the water was clear and was used for both drinking water and household purposes by the riparian villages." pageId="23" pageNumber="24">Tables 1</tableCitation>
and
<tableCitation id="3F3483974522FFABFC771F71FC3C296C" box="[929,946,1087,1113]" captionStart="TABLE 2" captionStartId="6.[152,244,1634,1658]" captionText="TABLE 2. Measurements of physical and chemical water parameters and variables at the time of well fauna collection. WT= water temperature (°C), EC = electrical conductivity (µS cm-1), DO = dissolved oxygen (mg L-1). NM= no measurement." pageId="23" pageNumber="24">2</tableCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74522FFA1FF411FC8FD9E2900" lastPageId="29" lastPageNumber="30" pageId="23" pageNumber="24" type="description">
<paragraph id="7209B62C4522FFABFF411FC8FE0F2994" blockId="23.[151,1437,1158,2013]" box="[151,385,1158,1185]" pageId="23" pageNumber="24">
<emphasis id="40C26A3E4522FFABFF411FC8FE0F2994" bold="true" box="[151,385,1158,1185]" pageId="23" pageNumber="24">Description of male</emphasis>
</paragraph>
<paragraph id="7209B62C4522FFABFF411FE5FC9F28A8" blockId="23.[151,1437,1158,2013]" pageId="23" pageNumber="24">
CpRl (
<figureCitation id="EA8DAAA94522FFABFF331FE5FEE229F0" box="[229,364,1195,1221]" captionStart="FIGURE 10" captionStartId="21.[151,250,1855,1879]" captionTargetBox="[151,1436,883,1832]" captionTargetId="figure-159@21.[151,1436,883,1832]" captionTargetPageId="21" captionText="FIGURE 10. Benincandona ibikounlei gen. et spec. nov., valves. A. INV.159222. CpRl. B. INV.159222. CpV. C. INV.159223. CpD. D. INV.159224. LVi. E. INV.159224. RVi. Scale bar = 500 µm." pageId="23" pageNumber="24">Figure 10A</figureCitation>
) elongated (L/H ratio = 2.4) and sub-triangular, with anterior margin slightly more broadly rounded than posterior margin, dorsal margin bluntly pointed, with greatest height slightly in front of the middle, anterior part sloping almost straight to anterior margin, posterior part sloping almost straight to posterior margin, ventral margin almost straight; LV slightly overlapping RV along anterior, ventral and posterior sides, dorsally strongly overlapping RV; valves externally smooth. CpV (
<figureCitation id="EA8DAAA94522FFABFCF11E75FC272860" box="[807,937,1339,1365]" captionStart="FIGURE 10" captionStartId="21.[151,250,1855,1879]" captionTargetBox="[151,1436,883,1832]" captionTargetId="figure-159@21.[151,1436,883,1832]" captionTargetPageId="21" captionText="FIGURE 10. Benincandona ibikounlei gen. et spec. nov., valves. A. INV.159222. CpRl. B. INV.159222. CpV. C. INV.159223. CpD. D. INV.159224. LVi. E. INV.159224. RVi. Scale bar = 500 µm." pageId="23" pageNumber="24">Figure 10B</figureCitation>
) and CpD (
<figureCitation id="EA8DAAA94522FFABFBF81E75FB3F2860" box="[1070,1201,1339,1365]" captionStart="FIGURE 10" captionStartId="21.[151,250,1855,1879]" captionTargetBox="[151,1436,883,1832]" captionTargetId="figure-159@21.[151,1436,883,1832]" captionTargetPageId="21" captionText="FIGURE 10. Benincandona ibikounlei gen. et spec. nov., valves. A. INV.159222. CpRl. B. INV.159222. CpV. C. INV.159223. CpD. D. INV.159224. LVi. E. INV.159224. RVi. Scale bar = 500 µm." pageId="23" pageNumber="24">Figure 10C</figureCitation>
) narrow and pointed on both anterior and posterior sides, greatest width in the middle, lateral margins angular in the middle; CpV with LV forming a pronounced overlap over RV in the centre.
</paragraph>
<paragraph id="7209B62C4522FFABFF111EE9FBEA2B18" blockId="23.[151,1437,1158,2013]" pageId="23" pageNumber="24">
LVi (
<figureCitation id="EA8DAAA94522FFABFF291EE9FE0A28F4" box="[255,388,1447,1473]" captionStart="FIGURE 10" captionStartId="21.[151,250,1855,1879]" captionTargetBox="[151,1436,883,1832]" captionTargetId="figure-159@21.[151,1436,883,1832]" captionTargetPageId="21" captionText="FIGURE 10. Benincandona ibikounlei gen. et spec. nov., valves. A. INV.159222. CpRl. B. INV.159222. CpV. C. INV.159223. CpD. D. INV.159224. LVi. E. INV.159224. RVi. Scale bar = 500 µm." pageId="23" pageNumber="24">Figure 10D</figureCitation>
) elongated and triangular, with anterior margin slightly more broadly rounded than posterior margin, dorsal margin bluntly pointed, with greatest height in the middle showing an overlap, anterior part sloping almost straight to anterior margin, posterior part sloping almost straight to posterior margin, ventral margin almost straight, with a slight inward curl in the middle; one ventral il and antero-ventral tooth.
</paragraph>
<paragraph id="7209B62C4522FFABFF111D79FE062B88" blockId="23.[151,1437,1158,2013]" pageId="23" pageNumber="24">
RVi (
<figureCitation id="EA8DAAA94522FFABFED71D79FE0C2B64" box="[257,386,1591,1617]" captionStart="FIGURE 10" captionStartId="21.[151,250,1855,1879]" captionTargetBox="[151,1436,883,1832]" captionTargetId="figure-159@21.[151,1436,883,1832]" captionTargetPageId="21" captionText="FIGURE 10. Benincandona ibikounlei gen. et spec. nov., valves. A. INV.159222. CpRl. B. INV.159222. CpV. C. INV.159223. CpD. D. INV.159224. LVi. E. INV.159224. RVi. Scale bar = 500 µm." pageId="23" pageNumber="24">Figure 10E</figureCitation>
) elongated, with anterior margin slightly more broadly rounded than posterior margin, dorsal margin greatest height situated about 1/3 from the rear, anterior part sloping almost straight to anterior margin, posterior part sloping almost straight to posterior margin, ventral margin almost straight; il running submarginally along ventral margin.
</paragraph>
<paragraph id="7209B62C4522FFA4FF111D89FE0D2C74" blockId="23.[151,1437,1158,2013]" lastBlockId="24.[151,1437,151,1905]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
A1 (
<figureCitation id="EA8DAAA94522FFABFF2E1D89FEF62BD4" box="[248,376,1735,1761]" captionStart="FIGURE 11" captionStartId="22.[151,250,1943,1967]" captionTargetBox="[183,1405,185,1915]" captionTargetId="figure-31@22.[179,1407,181,1919]" captionTargetPageId="22" captionText="FIGURE 11. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. A2 (INV.159219, INV.159217, INV.159218). B. A1 (INV.159219, INV.159218). C, D. MdCoxa, (INV.159218). Scale bar = 50 µm." pageId="23" pageNumber="24">Figure 11B</figureCitation>
) consisting of seven articulated segments, one basal and six distal. Basal segment large with
<emphasis id="40C26A3E4522FFABFA5F1D89FA122BD5" box="[1417,1436,1735,1760]" italics="true" pageId="23" pageNumber="24">c.</emphasis>
2.5 times as long as basal width, supporting two ventro-subapical setae, more proximal about three times as long as second one; and two setae on the dorsal side situated some distance from each other with the more proximal one slightly longer than the second one. First distal segment length about twice the basal width, with one short dorso-apical seta, no ventral apical seta present. Next (second distal segment) about twice as long as wide, with one dorso-apical seta. Third distal segment slightly longer than the previous one, with one short apical seta on each side. Fourth distal segment also slightly longer than the previous one (more than three time as long as wide) with one short ventro-apical seta and one long dorso-apical seta, reaching beyond tip of sixth distal segment with almost half the total length. Fifth distal segment more than four times as long as wide, with two long dorso-apical setae (almost five times longer than the length of the segment) and one short ventro-apical seta slightly exceeding the middle of the next segment. Sixth distal (terminal) segment, narrow,
<emphasis id="40C26A3E452DFFA4FCFC1B91FCB32DCD" box="[810,829,223,248]" italics="true" pageId="24" pageNumber="25">c.</emphasis>
five times as long as wide, bearing one short and two longer setae and one aesthetasc (Ya). Both long setae about 2.5x the length of Ya, length of the short one is about half the length of Ya.
</paragraph>
<paragraph id="7209B62C452DFFA4FF111A05FBED2E68" blockId="24.[151,1437,151,1905]" pageId="24" pageNumber="25">
A2 (
<figureCitation id="EA8DAAA9452DFFA4FF2E1A05FEF52C50" box="[248,379,331,357]" captionStart="FIGURE 11" captionStartId="22.[151,250,1943,1967]" captionTargetBox="[183,1405,185,1915]" captionTargetId="figure-31@22.[179,1407,181,1919]" captionTargetPageId="22" captionText="FIGURE 11. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. A2 (INV.159219, INV.159217, INV.159218). B. A1 (INV.159219, INV.159218). C, D. MdCoxa, (INV.159218). Scale bar = 50 µm." pageId="24" pageNumber="25">Figure 11A</figureCitation>
) with a basal segment broad and rounded, mid-ventrally with one seta of medium length, two subapical setae of unequal length, proximal one
<emphasis id="40C26A3E452DFFA4FD671A21FD4A2CBD" box="[689,708,367,392]" italics="true" pageId="24" pageNumber="25">c.</emphasis>
twice the length of the more apical one. Second segment (second protopodite) large and
<emphasis id="40C26A3E452DFFA4FE4D1ADDFE202C99" box="[411,430,403,428]" italics="true" pageId="24" pageNumber="25">c.</emphasis>
twice as long as basal width, set with one short ventro-subapical seta X. No remnant of exopodite visible. Endopodite consisting of four segments with the penultimate segment divided. First endopodal segment long
<emphasis id="40C26A3E452DFFA4FEEF1A95FEC22CC1" box="[313,332,475,500]" italics="true" pageId="24" pageNumber="25">c.</emphasis>
five times as long as basal width, carrying one long mid-ventral aesthetasc (Y) (reaching tip of segment) and two ventro-apical setae, one short and one long, hirsute in its distal 4/5. Second endopodal segment (segment 2A)
<emphasis id="40C26A3E452DFFA4FEEF196DFEC32F09" box="[313,333,547,572]" italics="true" pageId="24" pageNumber="25">c.</emphasis>
twice as long as wide, carrying one stout dorso-apical seta and three t-setae; seta t1 long, narrow and plumose, accompanied by a long and slender, but well-developed aesthetasc y1; setae t2 (ventro-apical) and t3 (centro-apical) hyper-developed into male bristles. Third endopodal segment (segment 2B) smaller than previous segment and almost as long as wide, set with a complex (and sexually dimorphic) apical chaetotaxy (
<figureCitation id="EA8DAAA9452DFFA4FACA19C1FA1E2F9C" box="[1308,1424,655,681]" captionStart="FIGURE 40" captionStartId="60.[151,250,1516,1540]" captionTargetBox="[151,1435,181,1492]" captionTargetId="figure-31@60.[151,1435,181,1492]" captionTargetPageId="60" captionText="FIGURE 40. Diagrams of apical views of the A2 of Benincandona, showing position of (sub-) terminal claws, setae and aesthetascs of the terminal (small circle) and penultimate segment (large circle). For explanations of codes, see Material and Methods (Abbreviations used in text and figures)." pageId="24" pageNumber="25">Figure 40</figureCitation>
): ventro-apically with one short and slender aesthetasc y2 and the claw G3 transformed into short seta of almost equal length; centrally with one short seta z3; dorso-apically, with z1 claw-like, length twice as long as penultimate segment. Claw G2 about 2.5 times as long as z1; z2 seta transformed into long claw, slightly shorter than G1. Fourth endopodal, terminal, segment with one long claw (Gm), one short claw (GM), a long aesthetasc y3 with shorter accompanying seta, with fused part short, and seta g slightly longer than aesthetasc y3.
</paragraph>
<paragraph id="7209B62C452DFFA4FF111829FC302EFC" blockId="24.[151,1437,151,1905]" pageId="24" pageNumber="25">
MdCoxa (
<figureCitation id="EA8DAAA9452DFFA4FEEE1829FE522EB4" box="[312,476,871,897]" captionStart="FIGURE 11" captionStartId="22.[151,250,1943,1967]" captionTargetBox="[183,1405,185,1915]" captionTargetId="figure-31@22.[179,1407,181,1919]" captionTargetPageId="22" captionText="FIGURE 11. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. A2 (INV.159219, INV.159217, INV.159218). B. A1 (INV.159219, INV.159218). C, D. MdCoxa, (INV.159218). Scale bar = 50 µm." pageId="24" pageNumber="25">Figure 11CD</figureCitation>
) elongated, proximally sharply pointed, medially widened, distally with an obliquely widened end, set with a series (67) of strong teeth, interspaced with setae; subapically with a short seta. Remark: asymmetry illustrated possibly resulting from different position in slide.
</paragraph>
<paragraph id="7209B62C452DFFA4FF11189DFAEE29DC" blockId="24.[151,1437,151,1905]" pageId="24" pageNumber="25">
MdPalp (
<figureCitation id="EA8DAAA9452DFFA4FEE7189DFE3B2ED8" box="[305,437,979,1005]" captionStart="FIGURE 12" captionStartId="25.[151,250,1981,2005]" captionTargetBox="[166,1422,183,1955]" captionTargetId="figure-17@25.[166,1422,181,1957]" captionTargetPageId="25" captionText="FIGURE 12. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. Mx1 (Chaetotaxy incomplete) (INV.159218). B. Mx1, respiratory plate (INV.159218). C. MdPalp (INV.159218). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." pageId="24" pageNumber="25">Figure 12C</figureCitation>
) consisting of four segments. Basal segment ventro-apically with two hirsute setae, S1 (long, reaching tip of terminal segment) and S2 (short and stout, length about one third of the length of S1), in between with the short flagellum-like alpha seta (thin and about as long as S2), and one smooth and medium length, more proximal seta. Second segment of sub-triangular shape, dorso-apically with two setae of unequal length, ventro-apically with a group of four long setae, one of these being the beta-seta. Third segment with two unequal dorso-subapical setae, two mid-dorsal setae, one of which being the gamma seta and ventro-apically with 2 setae of sub-equal length. Terminal segment sub-rectangular, apically set with one large claw, fused with the segment over its entire distal margin, accompanied by one smooth lateral seta on each side, about as long as the fused claw.
</paragraph>
<paragraph id="7209B62C452DFFA4FF111FBDFC2028D0" blockId="24.[151,1437,151,1905]" pageId="24" pageNumber="25">
Mx1 (
<figureCitation id="EA8DAAA9452DFFA4FED91FBDFE322838" box="[271,444,1267,1293]" captionStart="FIGURE 12" captionStartId="25.[151,250,1981,2005]" captionTargetBox="[166,1422,183,1955]" captionTargetId="figure-17@25.[166,1422,181,1957]" captionTargetPageId="25" captionText="FIGURE 12. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. Mx1 (Chaetotaxy incomplete) (INV.159218). B. Mx1, respiratory plate (INV.159218). C. MdPalp (INV.159218). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." pageId="24" pageNumber="25">Figure 12A, B</figureCitation>
) with a basal (protopodite) part carrying a large respiratory plate (exopodite), three endites and a two-segmented palp (endopodite). Respiratory plate (
<figureCitation id="EA8DAAA9452DFFA4FC5D1E59FB802804" box="[907,1038,1303,1329]" captionStart="FIGURE 12" captionStartId="25.[151,250,1981,2005]" captionTargetBox="[166,1422,183,1955]" captionTargetId="figure-17@25.[166,1422,181,1957]" captionTargetPageId="25" captionText="FIGURE 12. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. Mx1 (Chaetotaxy incomplete) (INV.159218). B. Mx1, respiratory plate (INV.159218). C. MdPalp (INV.159218). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." pageId="24" pageNumber="25">Figure 12B</figureCitation>
) elongated, carrying
<emphasis id="40C26A3E452DFFA4FB281E59FA9C2805" box="[1278,1298,1303,1328]" italics="true" pageId="24" pageNumber="25">c.</emphasis>
13 plumose respiratory rays, two short distal setae and a group of four shorter proximal setae of unequal length. Palp (
<figureCitation id="EA8DAAA9452DFFA4FAE31E75FA1A2860" box="[1333,1428,1339,1365]" captionStart="FIGURE 12" captionStartId="25.[151,250,1981,2005]" captionTargetBox="[166,1422,183,1955]" captionTargetId="figure-17@25.[166,1422,181,1957]" captionTargetPageId="25" captionText="FIGURE 12. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. Mx1 (Chaetotaxy incomplete) (INV.159218). B. Mx1, respiratory plate (INV.159218). C. MdPalp (INV.159218). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." pageId="24" pageNumber="25">Fig 12A</figureCitation>
) with first segment carrying four long dorso-apical setae; terminal segment short and rectangular, carrying also 4 distal setae, some slightly claw-like; chaetotaxy of three endites incompletely illustrated: third endite with two strong teeth carrying lateral spines (
<emphasis id="40C26A3E452DFFA4FDF11EE9FD3028F4" box="[551,702,1447,1473]" italics="true" pageId="24" pageNumber="25">Zahnborsten)</emphasis>
, basally with one lateral seta; sideways directed bristles near base of first endite one long and one short
<emphasis id="40C26A3E452DFFA4FDED1E85FDC028D1" box="[571,590,1483,1508]" italics="true" pageId="24" pageNumber="25">c.</emphasis>
half the length of the long one.
</paragraph>
<paragraph id="7209B62C452DFFA4FF111EA1FF582BAC" blockId="24.[151,1437,151,1905]" pageId="24" pageNumber="25">
T1 (
<figureCitation id="EA8DAAA9452DFFA4FF251EA1FE1A2B3C" box="[243,404,1518,1545]" captionStart="FIGURE 15" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[176,1407,181,1915]" captionTargetId="figure-31@28.[176,1411,181,1919]" captionTargetPageId="28" captionText="FIGURE 15. Benincandona ibikounlei gen. et spec. nov., limbs (AB ♂ and CE ♀). A. T1, Lpp (INV.159219). B. T1, Rpp (INV.159219). C. A2 (INV.159216). D. CR (INV.159216). E. T1 (INV.159221). Scale bar = 50 µm." pageId="24" pageNumber="25">Figure 15A, B</figureCitation>
) basal part (protopodite) carrying respiratory plates (exopodites, not shown), distally set with 4+7 setae of different morphology and length; more proximally with setae d and b of medium length. Prehensile palps (
<figureCitation id="EA8DAAA9452DFFA4FF351D79FE052B64" box="[227,395,1590,1617]" captionStart="FIGURE 15" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[176,1407,181,1915]" captionTargetId="figure-31@28.[176,1411,181,1919]" captionTargetPageId="28" captionText="FIGURE 15. Benincandona ibikounlei gen. et spec. nov., limbs (AB ♂ and CE ♀). A. T1, Lpp (INV.159219). B. T1, Rpp (INV.159219). C. A2 (INV.159216). D. CR (INV.159216). E. T1 (INV.159221). Scale bar = 50 µm." pageId="24" pageNumber="25">Figure 15A, B</figureCitation>
) one-segmented, distally sigmoid hook-like, both palps set with a short sensory organ and proximally with two subequal setae (also sensory organs); Lpp (
<figureCitation id="EA8DAAA9452DFFA4FCA11D15FC722B40" box="[887,1020,1626,1653]" captionStart="FIGURE 15" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[176,1407,181,1915]" captionTargetId="figure-31@28.[176,1411,181,1919]" captionTargetPageId="28" captionText="FIGURE 15. Benincandona ibikounlei gen. et spec. nov., limbs (AB ♂ and CE ♀). A. T1, Lpp (INV.159219). B. T1, Rpp (INV.159219). C. A2 (INV.159216). D. CR (INV.159216). E. T1 (INV.159221). Scale bar = 50 µm." pageId="24" pageNumber="25">Figure 15A</figureCitation>
) slightly narrower than Rpp (
<figureCitation id="EA8DAAA9452DFFA4FA871D15FF472BAC" captionStart="FIGURE 15" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[176,1407,181,1915]" captionTargetId="figure-31@28.[176,1411,181,1919]" captionTargetPageId="28" captionText="FIGURE 15. Benincandona ibikounlei gen. et spec. nov., limbs (AB ♂ and CE ♀). A. T1, Lpp (INV.159219). B. T1, Rpp (INV.159219). C. A2 (INV.159216). D. CR (INV.159216). E. T1 (INV.159221). Scale bar = 50 µm." pageId="24" pageNumber="25">Figure 15B</figureCitation>
).
</paragraph>
<paragraph id="7209B62C452DFFA4FF111DEDFB5C2A44" blockId="24.[151,1437,151,1905]" pageId="24" pageNumber="25">
T2 (
<figureCitation id="EA8DAAA9452DFFA4FF231DEDFEF72B88" box="[245,377,1699,1725]" captionStart="FIGURE 13" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[191,1395,185,1916]" captionTargetId="figure-31@26.[187,1399,181,1919]" captionTargetPageId="26" captionText="FIGURE 13. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. CR (INV.159219). B. CRAtt (INV.159219). C. T3 (INV.159219). D. T2 (INV.159217). Scale bar = 50 µm." pageId="24" pageNumber="25">Figure 13D</figureCitation>
) with 4-segmented endopodite (penultimate segment divided), elongated, without basal setae. Setae d1 and d2 absent. First endopodal segment without any setae, length
<emphasis id="40C26A3E452DFFA4FC2B1D89FB9E2BD5" box="[1021,1040,1735,1760]" italics="true" pageId="24" pageNumber="25">c.</emphasis>
five times as long as basal width. Second endopodal segment (2A) set with one short ventro-apical seta f, length
<emphasis id="40C26A3E452DFFA4FBC21DA5FBA92A31" box="[1044,1063,1771,1796]" italics="true" pageId="24" pageNumber="25">c.</emphasis>
3x basal width. Third endopodal segment (2B) of about the same length/width ratio as the previous segment and with one ventro-subapical seta g, reaching about 1/3 of following segment. Terminal segment about 1.5 times as long as basal width, apically two short setae, h1 ventral and h3 dorsal; and a long slender claw (h2), distally set with short setulae.
</paragraph>
<caption id="26C9E6A4452CFFA5FF411CF3FBFF2ACF" pageId="25" pageNumber="26" startId="25.[151,250,1981,2005]" targetBox="[166,1422,183,1955]" targetPageId="25" targetType="figure">
<paragraph id="7209B62C452CFFA5FF411CF3FBFF2ACF" blockId="25.[151,1436,1981,2042]" pageId="25" pageNumber="26">
<emphasis id="40C26A3E452CFFA5FF411CF3FEAE2AE0" bold="true" box="[151,288,1981,2005]" pageId="25" pageNumber="26">FIGURE 12.</emphasis>
<taxonomicName id="B5B6CDAF452CFFA5FEF11CF3FDAA2AE0" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[295,548,1981,2005]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="ibikounlei" status="gen. et spec. nov.">
<emphasis id="40C26A3E452CFFA5FEF11CF3FDAA2AE0" box="[295,548,1981,2005]" italics="true" pageId="25" pageNumber="26">Benincandona ibikounlei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745452CFFA5FDFD1CF0FD532AE0" box="[555,733,1982,2006]" pageId="25" pageNumber="26" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E452CFFA5FC811CF3FCE72AE0" bold="true" box="[855,873,1981,2005]" pageId="25" pageNumber="26">A</emphasis>
. Mx1 (Chaetotaxy incomplete) (INV.159218).
<emphasis id="40C26A3E452CFFA5FA9E1CF0FAD72AE3" bold="true" box="[1352,1369,1982,2006]" pageId="25" pageNumber="26">B</emphasis>
. Mx1, respiratory plate (INV.159218).
<emphasis id="40C26A3E452CFFA5FE0A1CAFFE602ACC" bold="true" box="[476,494,2017,2041]" pageId="25" pageNumber="26">C</emphasis>
. MdPalp (INV.159218). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A4452FFFA6FF411CD9FD432AE1" pageId="26" pageNumber="27" startId="26.[151,250,1943,1967]" targetBox="[191,1395,185,1916]" targetPageId="26" targetType="figure">
<paragraph id="7209B62C452FFFA6FF411CD9FD432AE1" blockId="26.[151,1436,1943,2004]" pageId="26" pageNumber="27">
<emphasis id="40C26A3E452FFFA6FF411CD9FEAC2A9A" bold="true" box="[151,290,1943,1967]" pageId="26" pageNumber="27">FIGURE 13.</emphasis>
<taxonomicName id="B5B6CDAF452FFFA6FEFD1CD6FDA22A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[299,556,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="ibikounlei" status="gen. et spec. nov.">
<emphasis id="40C26A3E452FFFA6FEFD1CD6FDA22A85" box="[299,556,1944,1968]" italics="true" pageId="26" pageNumber="27">Benincandona ibikounlei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745452FFFA6FDE31CD6FD602A9A" box="[565,750,1944,1968]" pageId="26" pageNumber="27" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E452FFFA6FCB91CD9FC0F2A9A" bold="true" box="[879,897,1943,1967]" pageId="26" pageNumber="27">A</emphasis>
. CR (INV.159219).
<emphasis id="40C26A3E452FFFA6FB811CD6FBE62A85" bold="true" box="[1111,1128,1944,1968]" pageId="26" pageNumber="27">B</emphasis>
. CRAtt (INV.159219).
<emphasis id="40C26A3E452FFFA6FA891CD9FAFF2A9A" bold="true" box="[1375,1393,1943,1967]" pageId="26" pageNumber="27">C</emphasis>
. T3 (INV.159219).
<emphasis id="40C26A3E452FFFA6FEE71CF2FECD2AE1" bold="true" box="[305,323,1980,2004]" pageId="26" pageNumber="27">D</emphasis>
. T2 (INV.159217). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A4452EFFA7FF411CD9FDB32ACD" pageId="27" pageNumber="28" startId="27.[151,250,1943,1967]" targetBox="[229,1359,191,1909]" targetPageId="27" targetType="figure">
<paragraph id="7209B62C452EFFA7FF411CD9FDB32ACD" blockId="27.[151,1437,1943,2040]" pageId="27" pageNumber="28">
<emphasis id="40C26A3E452EFFA7FF411CD9FEAF2A9A" bold="true" box="[151,289,1943,1967]" pageId="27" pageNumber="28">FIGURE 14.</emphasis>
<taxonomicName id="B5B6CDAF452EFFA7FEFE1CD9FDA82A9A" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[296,550,1943,1967]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="ibikounlei" status="gen. et spec. nov.">
<emphasis id="40C26A3E452EFFA7FEFE1CD9FDA82A9A" box="[296,550,1943,1967]" italics="true" pageId="27" pageNumber="28">Benincandona ibikounlei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745452EFFA7FDFB1CD6FD6F2A9A" box="[557,737,1944,1968]" pageId="27" pageNumber="28" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E452EFFA7FC8A1CD9FCE02A9A" bold="true" box="[860,878,1943,1967]" pageId="27" pageNumber="28">A</emphasis>
. Zenker organ (INV.159219).
<emphasis id="40C26A3E452EFFA7FB701CD9FB5B2A9A" bold="true" box="[1190,1237,1943,1967]" pageId="27" pageNumber="28">BD</emphasis>
. Hp (INV.159219).
<emphasis id="40C26A3E452EFFA7FF411CF5FF262AE6" bold="true" box="[151,168,1979,2003]" pageId="27" pageNumber="28">B</emphasis>
. External view (drawing).
<emphasis id="40C26A3E452EFFA7FE141CF5FE5A2AE6" bold="true" box="[450,468,1979,2003]" pageId="27" pageNumber="28">C</emphasis>
. Internal view (photograph without interference contrast).
<emphasis id="40C26A3E452EFFA7FBEE1CF5FBC42AE6" bold="true" box="[1080,1098,1979,2003]" pageId="27" pageNumber="28">D</emphasis>
. Internal view (photograph with interference contrast). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A44529FFA0FF411CD9FB072AE1" pageId="28" pageNumber="29" startId="28.[151,250,1943,1967]" targetBox="[176,1407,181,1915]" targetPageId="28" targetType="figure">
<paragraph id="7209B62C4529FFA0FF411CD9FB072AE1" blockId="28.[151,1436,1943,2004]" pageId="28" pageNumber="29">
<emphasis id="40C26A3E4529FFA0FF411CD9FEAF2A9A" bold="true" box="[151,289,1943,1967]" pageId="28" pageNumber="29">FIGURE 15.</emphasis>
<taxonomicName id="B5B6CDAF4529FFA0FEFF1CD6FDA62A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[297,552,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="ibikounlei" status="gen. et spec. nov.">
<emphasis id="40C26A3E4529FFA0FEFF1CD6FDA62A85" box="[297,552,1944,1968]" italics="true" pageId="28" pageNumber="29">Benincandona ibikounlei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454529FFA0FDE61CD6FD6B2A9A" box="[560,741,1944,1968]" pageId="28" pageNumber="29" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (AB ♂ and CE ♀).
<emphasis id="40C26A3E4529FFA0FBC21CD9FBA82A9A" bold="true" box="[1044,1062,1943,1967]" pageId="28" pageNumber="29">A</emphasis>
. T1, Lpp (INV.159219).
<emphasis id="40C26A3E4529FFA0FAFC1CD6FAB52A85" bold="true" box="[1322,1339,1944,1968]" pageId="28" pageNumber="29">B</emphasis>
. T1, Rpp (INV.159219).
<emphasis id="40C26A3E4529FFA0FEE71CF5FECD2AE6" bold="true" box="[305,323,1979,2003]" pageId="28" pageNumber="29">C</emphasis>
. A2 (INV.159216).
<emphasis id="40C26A3E4529FFA0FDDB1CF2FD912AE1" bold="true" box="[525,543,1980,2004]" pageId="28" pageNumber="29">D</emphasis>
. CR (INV.159216).
<emphasis id="40C26A3E4529FFA0FD381CF2FD712AE1" bold="true" box="[750,767,1980,2004]" pageId="28" pageNumber="29">E</emphasis>
. T1 (INV.159221). Scale bar = 50 µm.
</paragraph>
</caption>
<paragraph id="7209B62C4528FFA1FF111BD9FE902C50" blockId="29.[151,1437,151,645]" pageId="29" pageNumber="30">
T3 (
<figureCitation id="EA8DAAA94528FFA1FF231BD9FEF82D84" box="[245,374,151,177]" captionStart="FIGURE 13" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[191,1395,185,1916]" captionTargetId="figure-31@26.[187,1399,181,1919]" captionTargetPageId="26" captionText="FIGURE 13. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. CR (INV.159219). B. CRAtt (INV.159219). C. T3 (INV.159219). D. T2 (INV.159217). Scale bar = 50 µm." pageId="29" pageNumber="30">Figure 13C</figureCitation>
) first segment with only one medium length seta d1. Second (knee-) segment without any seta. Third segment about 5x as long as wide, not carrying any seta. Fourth segment narrower,
<emphasis id="40C26A3E4528FFA1FB701BF5FB372DE1" box="[1190,1209,187,212]" italics="true" pageId="29" pageNumber="30">c.</emphasis>
7x basal width and slightly longer than the previous segment, carrying one short subapical seta g. Terminal segment well-separated from penultimate segment, L
<emphasis id="40C26A3E4528FFA1FE241A4DFD8B2C29" box="[498,517,259,284]" italics="true" pageId="29" pageNumber="30">c.</emphasis>
1.5x basal width and carrying three setae: one long and reflexed (h3), clearly exceeding half of the third segment and two ventrally directed short setae h1 and h2, the former
<emphasis id="40C26A3E4528FFA1FB0B1A69FB7E2C75" box="[1245,1264,295,320]" italics="true" pageId="29" pageNumber="30">c.</emphasis>
half the length of the latter.
</paragraph>
<paragraph id="7209B62C4528FFA1FF111A21FB082C98" blockId="29.[151,1437,151,645]" pageId="29" pageNumber="30">
CR (
<figureCitation id="EA8DAAA94528FFA1FF2D1A21FEF32CBC" box="[251,381,367,393]" captionStart="FIGURE 13" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[191,1395,185,1916]" captionTargetId="figure-31@26.[187,1399,181,1919]" captionTargetPageId="26" captionText="FIGURE 13. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. CR (INV.159219). B. CRAtt (INV.159219). C. T3 (INV.159219). D. T2 (INV.159217). Scale bar = 50 µm." pageId="29" pageNumber="30">Figure 13A</figureCitation>
) with ramus stout; claw Ga well-developed, stout with length ratio ramus/claw Ga = 1.5, setae Sp (posterior seta), Sa (anterior seta) and claw Gp (posterior claw) reduced to small setae.
</paragraph>
<paragraph id="7209B62C4528FFA1FF111AF9FABB2CE4" blockId="29.[151,1437,151,645]" box="[199,1333,439,465]" pageId="29" pageNumber="30">
CRAtt (
<figureCitation id="EA8DAAA94528FFA1FEC91AF9FE2F2CE4" box="[287,417,439,465]" captionStart="FIGURE 13" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[191,1395,185,1916]" captionTargetId="figure-31@26.[187,1399,181,1919]" captionTargetPageId="26" captionText="FIGURE 13. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. CR (INV.159219). B. CRAtt (INV.159219). C. T3 (INV.159219). D. T2 (INV.159217). Scale bar = 50 µm." pageId="29" pageNumber="30">Figure 13B</figureCitation>
) long and stout, distally bifurcated and subapically with additional lateral branch.
</paragraph>
<paragraph id="7209B62C4528FFA1FF111A95FD772F08" blockId="29.[151,1437,151,645]" pageId="29" pageNumber="30">
Hp (
<figureCitation id="EA8DAAA94528FFA1FF2F1A95FE102CC0" box="[249,414,475,501]" captionStart="FIGURE 14" captionStartId="27.[151,250,1943,1967]" captionTargetBox="[229,1359,191,1909]" captionTargetId="figure-17@27.[219,1369,181,1919]" captionTargetPageId="27" captionText="FIGURE 14. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159219). BD. Hp (INV.159219). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." pageId="29" pageNumber="30">Figure 14BD</figureCitation>
) of medium size, with (ls) relatively narrow, with skewed rectangular distal part, reaching well beyond the asymmetrically rounded ms; lobe h shorter and sub-rectangular; internal anatomy (
<figureCitation id="EA8DAAA94528FFA1FB241AB1FA1A2F2C" box="[1266,1428,511,537]" captionStart="FIGURE 14" captionStartId="27.[151,250,1943,1967]" captionTargetBox="[229,1359,191,1909]" captionTargetId="figure-17@27.[219,1369,181,1919]" captionTargetPageId="27" captionText="FIGURE 14. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159219). BD. Hp (INV.159219). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." pageId="29" pageNumber="30">Figure 14C, D</figureCitation>
) with postlabyrinthal spermiduct with one circular coil.
</paragraph>
<paragraph id="7209B62C4528FFA1FF111909FCDD2FB0" blockId="29.[151,1437,151,645]" pageId="29" pageNumber="30">
Zenker organ (
<figureCitation id="EA8DAAA94528FFA1FEA61909FE782F54" box="[368,502,583,609]" captionStart="FIGURE 14" captionStartId="27.[151,250,1943,1967]" captionTargetBox="[229,1359,191,1909]" captionTargetId="figure-17@27.[219,1369,181,1919]" captionTargetPageId="27" captionText="FIGURE 14. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159219). BD. Hp (INV.159219). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." pageId="29" pageNumber="30">Figure 14A</figureCitation>
) with
<emphasis id="40C26A3E4528FFA1FD951909FDD92F55" box="[579,599,583,608]" italics="true" pageId="29" pageNumber="30">c.</emphasis>
seven spinous whirls, relatively long and with an elongated central tube joining posterior funnel-shaped end plate to anterior end plate.
</paragraph>
<paragraph id="7209B62C4528FFA1FF4119FCFE182FF9" blockId="29.[151,1436,690,1185]" box="[151,406,690,716]" pageId="29" pageNumber="30">
<emphasis id="40C26A3E4528FFA1FF4119FCFE182FF9" bold="true" box="[151,406,690,716]" pageId="29" pageNumber="30">Description of female</emphasis>
</paragraph>
<paragraph id="7209B62C4528FFA1FF411999FB6F2FC4" blockId="29.[151,1436,690,1185]" box="[151,1249,727,753]" pageId="29" pageNumber="30">Cp and valves most likely as in the male, without obvious sexual dimorphism (see remark above).</paragraph>
<paragraph id="7209B62C4528FFA1FF1119B5FB072E20" blockId="29.[151,1436,690,1185]" box="[199,1161,762,789]" pageId="29" pageNumber="30">
Limbs (
<figureCitation id="EA8DAAA94528FFA1FECB19B5FE302E20" box="[285,446,762,789]" captionStart="FIGURE 15" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[176,1407,181,1915]" captionTargetId="figure-31@28.[176,1411,181,1919]" captionTargetPageId="28" captionText="FIGURE 15. Benincandona ibikounlei gen. et spec. nov., limbs (AB ♂ and CE ♀). A. T1, Lpp (INV.159219). B. T1, Rpp (INV.159219). C. A2 (INV.159216). D. CR (INV.159216). E. T1 (INV.159221). Scale bar = 50 µm." pageId="29" pageNumber="30">Figure 15CE</figureCitation>
) largely as in the male, but with sexually dimorphic A2 and T1.
</paragraph>
<paragraph id="7209B62C4528FFA1FF111851FC232E90" blockId="29.[151,1436,690,1185]" pageId="29" pageNumber="30">
A2 (
<figureCitation id="EA8DAAA94528FFA1FF2F1851FEF22E0C" box="[249,380,798,825]" captionStart="FIGURE 15" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[176,1407,181,1915]" captionTargetId="figure-31@28.[176,1411,181,1919]" captionTargetPageId="28" captionText="FIGURE 15. Benincandona ibikounlei gen. et spec. nov., limbs (AB ♂ and CE ♀). A. T1, Lpp (INV.159219). B. T1, Rpp (INV.159219). C. A2 (INV.159216). D. CR (INV.159216). E. T1 (INV.159221). Scale bar = 50 µm." pageId="29" pageNumber="30">Figure 15C</figureCitation>
) with second endopodal segment undivided, with two t-setae, t1 (long, narrow and plumose) and t2 (thin and smooth); apical chaetotaxy (
<figureCitation id="EA8DAAA94528FFA1FD47180DFC8F2E68" box="[657,769,835,861]" captionStart="FIGURE 40" captionStartId="60.[151,250,1516,1540]" captionTargetBox="[151,1435,181,1492]" captionTargetId="figure-31@60.[151,1435,181,1492]" captionTargetPageId="60" captionText="FIGURE 40. Diagrams of apical views of the A2 of Benincandona, showing position of (sub-) terminal claws, setae and aesthetascs of the terminal (small circle) and penultimate segment (large circle). For explanations of codes, see Material and Methods (Abbreviations used in text and figures)." pageId="29" pageNumber="30">Figure 40</figureCitation>
) with z1 claw-like, z2 and z3 being both setae, z1 twice as long as z2 and z3; claws G1, G2, G3 and GM all reaching to about the same point; seta g and aesthetasc y3 as in the male. Small claw Gm in female homologous to large claw in the male.
</paragraph>
<paragraph id="7209B62C4528FFA1FF1118E1FCFA2924" blockId="29.[151,1436,690,1185]" pageId="29" pageNumber="30">
T1 (
<figureCitation id="EA8DAAA94528FFA1FF2E18E1FEF52EFC" box="[248,379,942,969]" captionStart="FIGURE 15" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[176,1407,181,1915]" captionTargetId="figure-31@28.[176,1411,181,1919]" captionTargetPageId="28" captionText="FIGURE 15. Benincandona ibikounlei gen. et spec. nov., limbs (AB ♂ and CE ♀). A. T1, Lpp (INV.159219). B. T1, Rpp (INV.159219). C. A2 (INV.159216). D. CR (INV.159216). E. T1 (INV.159221). Scale bar = 50 µm." pageId="29" pageNumber="30">Figure 15E</figureCitation>
) with basal part with
<emphasis id="40C26A3E4528FFA1FDA918E1FD1C2EFD" box="[639,658,943,968]" italics="true" pageId="29" pageNumber="30">c.</emphasis>
10 distal setae of unequal length, setae b and d present, with long additional seta (d) present close to seta d; respiratory plate hidden on the dissected specimens. Endopodite a broad and short palp, with three subequal but short apical setae (h1-h3).
</paragraph>
<paragraph id="7209B62C4528FFA1FF111F55FD9E2900" blockId="29.[151,1436,690,1185]" box="[199,528,1051,1077]" pageId="29" pageNumber="30">
<emphasis id="40C26A3E4528FFA1FF111F55FE0E2900" bold="true" box="[199,384,1051,1077]" pageId="29" pageNumber="30">Measurements:</emphasis>
See
<tableCitation id="3F3483974528FFA1FE601F55FD842900" box="[438,522,1051,1077]" captionStart="TABLE 3" captionStartId="19.[152,244,1592,1616]" captionText="TABLE 3. Measurements of illustrated specimens of Benincandona gen. et spec. nov., using SEM." pageId="29" pageNumber="30">Table 3</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74528FFA1FF111F70FDEA296C" box="[199,612,1086,1113]" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="7209B62C4528FFA1FF111F70FDEA296C" blockId="29.[151,1436,690,1185]" box="[199,612,1086,1113]" pageId="29" pageNumber="30">
<emphasis id="40C26A3E4528FFA1FF111F70FE5B296D" bold="true" box="[199,469,1086,1112]" pageId="29" pageNumber="30">Differential Diagnosis:</emphasis>
See
<tableCitation id="3F3483974528FFA1FDDC1F71FDEE296C" box="[522,608,1087,1113]" captionStart="TABLE 4" captionStartId="20.[152,245,1665,1689]" captionText="TABLE 4. Differential character states between the five new species of Benincandona gen. nov." pageId="29" pageNumber="30">Table 4</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74528FFA1FF111F2DFB072994" pageId="29" pageNumber="30" type="biology_ecology">
<paragraph id="7209B62C4528FFA1FF111F2DFE612949" blockId="29.[151,1436,690,1185]" box="[199,495,1122,1149]" pageId="29" pageNumber="30">
<emphasis id="40C26A3E4528FFA1FF111F2DFE612949" bold="true" box="[199,495,1122,1149]" pageId="29" pageNumber="30">Ecology and distribution</emphasis>
</paragraph>
<paragraph id="7209B62C4528FFA1FF111FC9FB072994" blockId="29.[151,1436,690,1185]" box="[199,1161,1159,1185]" pageId="29" pageNumber="30">
This subterranean species is presently known from its
<typeStatus id="AD0D088E4528FFA1FCF11FC9FCD92994" box="[807,855,1159,1185]" pageId="29" pageNumber="30">type</typeStatus>
locality only (see
<tableCitation id="3F3483974528FFA1FBF01FC9FBF62994" box="[1062,1144,1159,1185]" captionStart="TABLE 1" captionStartId="5.[152,244,929,955]" captionText="TABLE 1. Localities where Benincandona gen. nov. sp. nov. were collected, coordinates and hydro-morphological characteristics. In all stations, the water was clear and was used for both drinking water and household purposes by the riparian villages." pageId="29" pageNumber="30">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,519 @@
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<mods:title id="E8DC3C9BB380F01983B76F4F04BC17E8">An endemic species flock of Candonidae Kaufmann, 1900 (Crustacea, Ostracoda) from wells in Benin (Africa), with the description of a new subfamily, a new genus and five new species</mods:title>
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<mods:namePart id="355D8BC12C8E209632FBED134354AAFA">Hotèkpo, Sourou Joseph</mods:namePart>
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<mods:affiliation id="2E7056142F919CBD10743945687018BB">Centre de Recherche pour la lutte contre les Maladies Infectieuses Tropicales (CReMIT / TIDRC), Université dAbomey-Calavi, Bénin jshotekpo @ gmail. com; https: // orcid. org / 0000 - 0001 - 9091 - 433 X &amp; Royal Belgian Institute of Natural Sciences, Freshwater Biology, Vautierstraat 29, B- 1000 Brussels, Belgium &amp; University of Hasselt, Research Group Zoology, Agoralaan Building D, B- 3590 Diepenbeek, Belgium ischoen @ naturalsciences. be; https: // orcid. org / 0000 - 0001 - 9269 - 6487</mods:affiliation>
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<mods:namePart id="EE7B131E97EEAEDE52029649FC7F4F2F">Schön, Isa</mods:namePart>
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<mods:namePart id="1088AC7095505FB6AF77CDA391DE013E">Martens, Koen</mods:namePart>
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<mods:affiliation id="42520D1326FE68C099CCD9B9E0251F81">Royal Belgian Institute of Natural Sciences, Freshwater Biology, Vautierstraat 29, B- 1000 Brussels, Belgium &amp; Ghent University, Dept. Biology, K. L. Ledeganckstraat 35, B- 9000 Ghent, Belgium kmartens @ naturalsciences. be; https: // orcid. org / 0000 - 0001 - 8680 - 973 X * Corresponding author</mods:affiliation>
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<paragraph id="7209B62C4528FFA1FF411FBCFC5D2804" blockId="29.[151,979,1266,1365]" pageId="29" pageNumber="30">
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<taxonomicName id="B5B6CDAF4528FFA1FF411FBCFE172838" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[151,409,1266,1293]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="sakitii" status="gen. et spec. nov.">
<emphasis id="40C26A3E4528FFA1FF411FBCFE172838" bold="true" box="[151,409,1266,1293]" italics="true" pageId="29" pageNumber="30">Benincandona sakitii</emphasis>
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<taxonomicNameLabel id="5BF1D7454528FFA1FE771FBDFDFF2838" box="[417,625,1267,1293]" pageId="29" pageNumber="30" rank="species">gen. et spec. nov.</taxonomicNameLabel>
Hotèkpo &amp; Martens
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<paragraph id="7209B62C4528FFA1FF411E75FE2C2860" blockId="29.[151,979,1266,1365]" box="[151,418,1339,1365]" pageId="29" pageNumber="30">
(
<figureCitation id="EA8DAAA94528FFA1FF491E75FECA2860" box="[159,324,1339,1365]" captionStart-0="FIGURE 16" captionStart-1="FIGURE 17" captionStart-2="FIGURE 18" captionStart-3="FIGURE 19" captionStart-4="FIGURE 20" captionStart-5="FIGURE 21" captionStart-6="FIGURE 22" captionStart-7="FIGURE 23" captionStartId-0="30.[151,250,1942,1966]" captionStartId-1="31.[151,250,1630,1654]" captionStartId-2="33.[151,250,1798,1822]" captionStartId-3="34.[151,250,1943,1967]" captionStartId-4="35.[151,250,1943,1967]" captionStartId-5="37.[151,250,1981,2005]" captionStartId-6="38.[151,250,1943,1967]" captionStartId-7="39.[151,250,1943,1967]" captionTargetBox-0="[169,1418,181,1919]" captionTargetBox-1="[151,1436,181,1606]" captionTargetBox-2="[151,1436,345,1775]" captionTargetBox-3="[156,1433,185,1915]" captionTargetBox-4="[185,1402,185,1915]" captionTargetBox-5="[177,1420,212,1949]" captionTargetBox-6="[195,1394,191,1912]" captionTargetBox-7="[173,1413,185,1915]" captionTargetId-0="figure-31@30.[169,1418,181,1919]" captionTargetId-1="figure-17@31.[151,1436,181,1606]" captionTargetId-2="figure-109@33.[151,1436,345,1775]" captionTargetId-3="figure-31@34.[152,1436,181,1919]" captionTargetId-4="figure-17@35.[181,1406,181,1919]" captionTargetId-5="figure-17@37.[166,1422,181,1957]" captionTargetId-6="figure-31@38.[185,1402,181,1919]" captionTargetId-7="figure-17@39.[170,1417,181,1919]" captionTargetPageId-0="30" captionTargetPageId-1="31" captionTargetPageId-2="33" captionTargetPageId-3="34" captionTargetPageId-4="35" captionTargetPageId-5="37" captionTargetPageId-6="38" captionTargetPageId-7="39" captionText-0="FIGURE 16. Benincandona sakitii gen. et spec. nov., valves (AC, G, IK ♂; DF, H ♀). AH. CpRl, (AC, G: INV.159235, DF, H: INV.159237). IK. INV.159236, CpD. Scale bars. 1 (B, E, I) = 500 µm; 2 (A, C, D, F, G, H, J, K) = 200 µm." captionText-1="FIGURE 17. Benincandona sakitii gen. et spec. nov., valves (♂). A, C, D, G. LVi (A, C, D. INV.159225; G. INV.159227). B, E, F. RVi (INV.159225). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." captionText-2="FIGURE 18. Benincandona sakitii gen. et spec. nov., valves (♀). A, C, D. LVi (A, C, D. INV.159226). B, E, F, G. RVi (INV.159226). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." captionText-3="FIGURE 19. Benincandona sakitii gen. et spec. nov., limbs (♂). A. A2 (INV.159225). B. A1 (INV.159225). C, D. MdCoxa (INV.159228). Scale bar = 50 µm." captionText-4="FIGURE 20. Benincandona sakitii gen. et spec. nov., limbs (♂). A. Mx1-palp (INV.159225). B. Mx1, respiratory plate (INV.159227). C. MdPalp (INV.159227). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." captionText-5="FIGURE 21. Benincandona sakitii et spec. nov., limbs (♂). A. T3 (INV.159225). B. T2 (INV.159225). C. CR (INV.159225). D. CRAtt (INV.15922). Scale bar = 50 µm." captionText-6="FIGURE 22. Benincandona sakitii gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159227). BD. Hp (INV.159225). B. External view (drawing). C. Internal view (photograph with interference contrast). D. Internal view (photograph without interference contrast). Scale bar = 50 µm." captionText-7="FIGURE 23. Benincandona sakitii gen. et spec. nov., limbs (A, B ♂ and CE ♀). A. T1, Lpp (INV.159228). B. T1, Rpp (INV.159228). C. A2 (INV.159226, INV.159231). D. CR (INV.159232). E. T1 (INV.159226). Scale bar = 50 µm." pageId="29" pageNumber="30">Figures 1623</figureCitation>
,
<figureCitation id="EA8DAAA94528FFA1FE861E75FE142860" box="[336,410,1339,1365]" captionStart="FIGURE 41" captionStartId="61.[151,250,1113,1137]" captionTargetBox="[151,1436,181,1089]" captionTargetId="figure-17@61.[151,1436,181,1089]" captionTargetPageId="61" captionText="FIGURE 41. Benincandona gen. et spec. nov., CpRl. (A, C, E, F, G, I, K, N, O, Q ♂ and B, D, H, J, L, M, P, R ♀). AD. B. martini. EF. B. ibikounlei. GJ. B. sakitii. KN. B. cebios. OR. B. trapezoides. Scale bar = 500 µm." pageId="29" pageNumber="30">41GJ</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74528FFA1FF411ECDFB6728D0" pageId="29" pageNumber="30" type="etymology">
<paragraph id="7209B62C4528FFA1FF411ECDFE9628A8" blockId="29.[151,1436,1411,1509]" box="[151,280,1411,1437]" pageId="29" pageNumber="30">
<emphasis id="40C26A3E4528FFA1FF411ECDFE9628A8" bold="true" box="[151,280,1411,1437]" pageId="29" pageNumber="30">Etymology</emphasis>
</paragraph>
<paragraph id="7209B62C4528FFA1FF411EE9FB6728D0" blockId="29.[151,1436,1411,1509]" pageId="29" pageNumber="30">
The species is named after Prof. Nestor G. Sakiti who initiated, in collaboration with Prof. Claude Boutin, subterranean biology research in
<collectingCountry id="0AA1F6BC4528FFA1FDDD1E85FDDF28D0" box="[523,593,1483,1509]" name="Benin" pageId="29" pageNumber="30">Benin</collectingCountry>
, in gratitude for his continued support to the career of SJH.
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74528FFA1FF411D5CFDE92A44" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="7209B62C4528FFA1FF411D5CFE842B19" blockId="29.[151,1437,1554,1905]" box="[151,266,1554,1580]" pageId="29" pageNumber="30">
<emphasis id="40C26A3E4528FFA1FF411D5CFE842B19" bold="true" box="[151,266,1554,1580]" pageId="29" pageNumber="30">Diagnosis</emphasis>
</paragraph>
<paragraph id="7209B62C4528FFA1FF411D79FDE92A44" blockId="29.[151,1437,1554,1905]" pageId="29" pageNumber="30">
CpRl elongated (L/H = 2.0), sub-triangular, with greatest height slightly behind the middle; LV dorsally extending beyond RV, anterior margin bluntly pointed and with anteroventral part almost straight, posterior margin asymmetrically rounded towards the almost straight ventral side, valve surface set with polygonal ridges in midventral and central part. CpD and CpV narrow, with greatest width in the middle and evenly rounded lateral sides, anterior margin sharply and posterior margin bluntly pointed. A2 with seta X (medium length) reaching beyond middle of aesthetasc Y in males, beyond tip of aesthetasc Y in females (long). This aesthetasc Y distally asymmetrically bifurcated. Second palp segment of Mx1 with L
<emphasis id="40C26A3E4528FFA1FD7B1C41FD372A1D" box="[685,697,1807,1832]" italics="true" pageId="29" pageNumber="30">c</emphasis>
. 1.5x basal W. T3 with seta d2 about 2/3 of L of seta d1. Hp with ls and ms with overlapping positions, narrower lobe ls on Hp with skewed rectangular distal margin, extending beyond lobe ms with more than half of its length.
</paragraph>
</subSubSection>
<caption id="26C9E6A4452BFFA2FF411CD8FAB22AE6" pageId="30" pageNumber="31" startId="30.[151,250,1942,1966]" targetBox="[169,1418,181,1919]" targetPageId="30" targetType="figure">
<paragraph id="7209B62C452BFFA2FF411CD8FAB22AE6" blockId="30.[151,1436,1942,2003]" pageId="30" pageNumber="31">
<emphasis id="40C26A3E452BFFA2FF411CD8FEAE2A9B" bold="true" box="[151,288,1942,1966]" pageId="30" pageNumber="31">FIGURE 16.</emphasis>
<taxonomicName id="B5B6CDAF452BFFA2FEF01CD9FE732A9A" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[294,509,1943,1967]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="sakitii" status="gen. et spec. nov.">
<emphasis id="40C26A3E452BFFA2FEF01CD9FE732A9A" box="[294,509,1943,1967]" italics="true" pageId="30" pageNumber="31">Benincandona sakitii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745452BFFA2FDD21CD9FD382A9B" box="[516,694,1943,1967]" pageId="30" pageNumber="31" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, valves (AC, G, IK ♂; DF, H ♀).
<emphasis id="40C26A3E452BFFA2FBE51CD8FBEB2A9B" bold="true" box="[1075,1125,1942,1966]" pageId="30" pageNumber="31">AH</emphasis>
. CpRl, (AC, G: INV.159235, DF, H: INV.159237).
<emphasis id="40C26A3E452BFFA2FE561CF5FE242AE6" bold="true" box="[384,426,1979,2003]" pageId="30" pageNumber="31">IK</emphasis>
. INV.159236, CpD. Scale bars. 1 (B, E, I) = 500 µm; 2 (A, C, D, F, G, H, J, K) = 200 µm.
</paragraph>
</caption>
<caption id="26C9E6A4452AFFA3FF411D10FB172BAE" pageId="31" pageNumber="32" startId="31.[151,250,1630,1654]" targetBox="[151,1436,181,1606]" targetPageId="31" targetType="figure">
<paragraph id="7209B62C452AFFA3FF411D10FB172BAE" blockId="31.[151,1436,1630,1691]" pageId="31" pageNumber="32">
<emphasis id="40C26A3E452AFFA3FF411D10FEAE2B43" bold="true" box="[151,288,1630,1654]" pageId="31" pageNumber="32">FIGURE 17.</emphasis>
<taxonomicName id="B5B6CDAF452AFFA3FEF11D10FE702B43" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[295,510,1630,1654]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="sakitii" status="gen. et spec. nov.">
<emphasis id="40C26A3E452AFFA3FEF11D10FE702B43" box="[295,510,1630,1654]" italics="true" pageId="31" pageNumber="32">Benincandona sakitii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745452AFFA3FDD31D11FD392B43" box="[517,695,1631,1655]" pageId="31" pageNumber="32" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, valves (♂).
<emphasis id="40C26A3E452AFFA3FCEC1D10FC222B43" bold="true" box="[826,940,1630,1655]" pageId="31" pageNumber="32">A, C, D, G</emphasis>
. LVi (A, C, D. INV.159225; G. INV.159227).
<emphasis id="40C26A3E452AFFA3FA531D11FF4A2BAE" bold="true" pageId="31" pageNumber="32">B, E, F</emphasis>
. RVi (INV.159225).
<emphasis id="40C26A3E452AFFA3FE4E1DCCFE3F2BAE" bold="true" box="[408,433,1666,1691]" pageId="31" pageNumber="32">G.</emphasis>
CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm.
</paragraph>
</caption>
<subSubSection id="3AACE5A7452AFF9CFF411DA3FC5C2F08" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32" type="materials_examined">
<paragraph id="7209B62C452AFFA3FF411DA3FEB02A33" blockId="31.[151,318,1772,1799]" box="[151,318,1772,1799]" pageId="31" pageNumber="32">
<emphasis id="40C26A3E452AFFA3FF411DA3FEB02A33" bold="true" box="[151,318,1772,1799]" pageId="31" pageNumber="32">Type material</emphasis>
</paragraph>
<paragraph id="7209B62C452AFFA3FF411C7BFE8A2A7A" blockId="31.[151,1282,1845,1907]" box="[151,260,1845,1871]" pageId="31" pageNumber="32">
<emphasis id="40C26A3E452AFFA3FF411C7BFE8A2A7A" bold="true" box="[151,260,1845,1871]" pageId="31" pageNumber="32">
<typeStatus id="AD0D088E452AFFA3FF411C7BFE8A2A7A" box="[151,260,1845,1871]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7209B62C452AFFA3FF411C17FA8C2A46" blockId="31.[151,1282,1845,1907]" box="[151,1282,1880,1907]" pageId="31" pageNumber="32">
<materialsCitation id="C2DEBC71452AFFA3FF411C17FA8C2A46" box="[151,1282,1880,1907]" collectionCode="INV" pageId="31" pageNumber="32" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="64B07DA5452AFFA3FF411C17FF4D2A46" box="[151,195,1881,1907]" count="1" pageId="31" pageNumber="32" type="male">1 ♂</specimenCount>
, with soft parts dissected in glycerine in a sealed slide, and with valves stored dry (
<collectionCode id="14A72EE9452AFFA3FBB01C17FB182A46" box="[1126,1174,1881,1907]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="31" pageNumber="32" type="Herbarium">INV</collectionCode>
.159225).
</materialsCitation>
</paragraph>
<paragraph id="7209B62C452AFFA3FF411CEEFF722A8F" blockId="31.[151,736,1952,2015]" box="[151,252,1952,1978]" pageId="31" pageNumber="32">
<emphasis id="40C26A3E452AFFA3FF411CEEFF722A8F" bold="true" box="[151,252,1952,1978]" pageId="31" pageNumber="32">
<typeStatus id="AD0D088E452AFFA3FF411CEEFF722A8F" box="[151,252,1952,1978]" pageId="31" pageNumber="32" type="allotype">Allotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7209B62C452AFFA3FF411C8BFD6E2AEA" blockId="31.[151,736,1952,2015]" box="[151,736,1988,2015]" pageId="31" pageNumber="32">
<materialsCitation id="C2DEBC71452AFFA3FF411C8BFD6E2AEA" box="[151,736,1988,2015]" collectionCode="INV" pageId="31" pageNumber="32" specimenCount="1" specimenCount-female="1" typeStatus="allotype">
<specimenCount id="64B07DA5452AFFA3FF411C8BFF4F2AEA" box="[151,193,1989,2015]" count="1" pageId="31" pageNumber="32" type="female">1 ♀</specimenCount>
, dissected and stored as the male (
<collectionCode id="14A72EE9452AFFA3FD921C8AFDF72AEA" box="[580,633,1988,2015]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="31" pageNumber="32" type="Herbarium">INV</collectionCode>
.159226).
</materialsCitation>
</paragraph>
<paragraph id="7209B62C4515FF9CFF411BD9FE812D84" blockId="32.[151,1436,151,285]" box="[151,271,151,177]" pageId="32" pageNumber="33">
<emphasis id="40C26A3E4515FF9CFF411BD9FE812D84" bold="true" box="[151,271,151,177]" pageId="32" pageNumber="33">
<typeStatus id="AD0D088E4515FF9CFF411BD9FE812D84" box="[151,271,151,177]" pageId="32" pageNumber="33" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7209B62C4515FF9CFF411BF5FADB2C28" blockId="32.[151,1436,151,285]" pageId="32" pageNumber="33">
<materialsCitation id="C2DEBC714515FF9CFF411BF5FBD12DE0" box="[151,1119,186,213]" collectionCode="INV" pageId="32" pageNumber="33" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="64B07DA54515FF9CFF411BF5FF542DE0" box="[151,218,187,213]" count="3" pageId="32" pageNumber="33" type="male">3 ♂♂</specimenCount>
, dissected and stored as the
<typeStatus id="AD0D088E4515FF9CFDF51BF5FD0B2DE0" box="[547,645,187,213]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
(
<collectionCode id="14A72EE94515FF9CFD431BF4FD472DE0" box="[661,713,186,213]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="32" pageNumber="33" type="Herbarium">INV</collectionCode>
.159227, INV.159228, INV.159229)
</materialsCitation>
;
<materialsCitation id="C2DEBC714515FF9CFBBB1BF5FD7D2DCC" collectionCode="INV" pageId="32" pageNumber="33" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<specimenCount id="64B07DA54515FF9CFBBB1BF5FB202DE0" box="[1133,1198,187,213]" count="3" pageId="32" pageNumber="33" type="female">3 ♀♀</specimenCount>
dissected and stored as the male (
<collectionCode id="14A72EE94515FF9CFEFF1B90FED32DCC" box="[297,349,222,249]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="32" pageNumber="33" type="Herbarium">INV</collectionCode>
.159230, INV.159231, INV.159232)
</materialsCitation>
;
<materialsCitation id="C2DEBC714515FF9CFCD61B91FDCD2C28" collectionCode="JSH" pageId="32" pageNumber="33" specimenCode="JSH0344" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="64B07DA54515FF9CFCD61B91FCCF2DCC" box="[768,833,223,249]" count="3" pageId="32" pageNumber="33" type="male">3 ♂♂</specimenCount>
carapaces stored dry in micropalaeontological slides (
<specimenCode id="22101E574515FF9CFF491A4DFE842C28" box="[159,266,259,285]" collectionCode="JSH" pageId="32" pageNumber="33">JSH0344</specimenCode>
, INV.159235, INV.159236)
</materialsCitation>
;
<materialsCitation id="C2DEBC714515FF9CFD981A4DFADB2C28" box="[590,1365,258,285]" collectionCode="INV" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="64B07DA54515FF9CFD981A4DFDF62C28" box="[590,632,259,285]" count="1" pageId="32" pageNumber="33" type="female">1 ♀</specimenCount>
carapace stored dry in micropalaeontological slide (
<collectionCode id="14A72EE94515FF9CFB171A4CFB7B2C28" box="[1217,1269,258,285]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="32" pageNumber="33" type="Herbarium">INV</collectionCode>
.159227)
</materialsCitation>
</paragraph>
<paragraph id="7209B62C4515FF9CFF411A05FEBE2C51" blockId="32.[151,1436,330,465]" box="[151,304,330,357]" pageId="32" pageNumber="33">
<emphasis id="40C26A3E4515FF9CFF411A05FEBE2C51" bold="true" box="[151,304,330,357]" pageId="32" pageNumber="33">Type locality</emphasis>
</paragraph>
<paragraph id="7209B62C4515FF9CFF411A21FCD32CE4" blockId="32.[151,1436,330,465]" pageId="32" pageNumber="33">
<materialsCitation id="C2DEBC714515FF9CFF411A21FE632C98" country="Benin" location="Tchaourou District" pageId="32" pageNumber="33" specimenCount="1" stateProvince="Borgou">
<collectingCountry id="0AA1F6BC4515FF9CFF411A21FF542CBC" box="[151,218,367,393]" name="Benin" pageId="32" pageNumber="33">Benin</collectingCountry>
,
<collectingRegion id="B07278CE4515FF9CFF311A21FE492CBC" box="[231,455,367,393]" country="Benin" name="Borgou" pageId="32" pageNumber="33">Borgou Department</collectingRegion>
,
<location id="7769E0F74515FF9CFE061A21FD2A2CBC" LSID="urn:lsid:plazi:treatment:FA1F073A4528FF94FF411FBCFEA02C98:7769E0F74515FF9CFE061A21FD2A2CBC" box="[464,676,367,393]" country="Benin" name="Tchaourou District" pageId="32" pageNumber="33" stateProvince="Borgou">Tchaourou District</location>
, town of Badékparou located in the Okpara catchment area (tributary of the
<collectingRegion id="B07278CE4515FF9CFF361ADDFEBF2C98" box="[224,305,403,429]" country="Benin" name="Oueme" pageId="32" pageNumber="33">Ouémé</collectingRegion>
catchment area)
</materialsCitation>
, coordinates:
<geoCoordinate id="1782D0EB4515FF9CFD5F1ADDFC832C98" box="[649,781,403,429]" degrees="9" direction="north" minutes="13.314" orientation="latitude" pageId="32" pageNumber="33" precision="1" value="9.2219">9°13.314N</geoCoordinate>
<geoCoordinate id="1782D0EB4515FF9CFCC21ADDFC162C98" box="[788,920,403,429]" degrees="2" direction="east" minutes="33.790" orientation="longitude" pageId="32" pageNumber="33" precision="1" value="2.5631666">2°33.790E</geoCoordinate>
, Leg.: S.J. Hotèkpo and M. Lagnika collected on
<date id="060890EC4515FF9CFF6C1AF9FEFB2CE4" box="[186,373,439,465]" pageId="32" pageNumber="33" value="2022-10-11">11 October 2022</date>
(Locality code BEN066 in
<tableCitation id="3F3483974515FF9CFD7D1AF9FC872CE4" box="[683,777,439,465]" captionStart="TABLE 1" captionStartId="5.[152,244,929,955]" captionText="TABLE 1. Localities where Benincandona gen. nov. sp. nov. were collected, coordinates and hydro-morphological characteristics. In all stations, the water was clear and was used for both drinking water and household purposes by the riparian villages." pageId="32" pageNumber="33">Tables 1</tableCitation>
and
<tableCitation id="3F3483974515FF9CFCE91AF9FCDE2CE4" box="[831,848,439,465]" captionStart="TABLE 2" captionStartId="6.[152,244,1634,1658]" captionText="TABLE 2. Measurements of physical and chemical water parameters and variables at the time of well fauna collection. WT= water temperature (°C), EC = electrical conductivity (µS cm-1), DO = dissolved oxygen (mg L-1). NM= no measurement." pageId="32" pageNumber="33">2</tableCitation>
).
</paragraph>
<paragraph id="7209B62C4515FF9CFF411AB0FE6E2F2D" blockId="32.[151,978,510,573]" box="[151,480,510,536]" pageId="32" pageNumber="33">
<emphasis id="40C26A3E4515FF9CFF411AB0FE6E2F2D" bold="true" box="[151,480,510,536]" pageId="32" pageNumber="33">Other material investigated</emphasis>
</paragraph>
<paragraph id="7209B62C4515FF9CFF41196DFC5C2F08" blockId="32.[151,978,510,573]" box="[151,978,547,573]" pageId="32" pageNumber="33">
See
<tableCitation id="3F3483974515FF9CFF10196DFE972F08" box="[198,281,547,573]" captionStart="TABLE 2" captionStartId="6.[152,244,1634,1658]" captionText="TABLE 2. Measurements of physical and chemical water parameters and variables at the time of well fauna collection. WT= water temperature (°C), EC = electrical conductivity (µS cm-1), DO = dissolved oxygen (mg L-1). NM= no measurement." pageId="32" pageNumber="33">Table 2</tableCitation>
for a list of localities where the present species was collected.
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74515FF94FF411924FD862C28" lastPageId="40" lastPageNumber="41" pageId="32" pageNumber="33" type="description">
<paragraph id="7209B62C4515FF9CFF411924FE0F2FB0" blockId="32.[151,1437,618,2013]" box="[151,385,618,645]" pageId="32" pageNumber="33">
<emphasis id="40C26A3E4515FF9CFF411924FE0F2FB0" bold="true" box="[151,385,618,645]" pageId="32" pageNumber="33">Description of male</emphasis>
</paragraph>
<paragraph id="7209B62C4515FF9CFF4119C1FDC42EB4" blockId="32.[151,1437,618,2013]" pageId="32" pageNumber="33">
CpRl (
<figureCitation id="EA8DAAA94515FF9CFF3619C1FE2C2F9C" box="[224,418,655,681]" captionStart="FIGURE 16" captionStartId="30.[151,250,1942,1966]" captionTargetBox="[169,1418,181,1919]" captionTargetId="figure-31@30.[169,1418,181,1919]" captionTargetPageId="30" captionText="FIGURE 16. Benincandona sakitii gen. et spec. nov., valves (AC, G, IK ♂; DF, H ♀). AH. CpRl, (AC, G: INV.159235, DF, H: INV.159237). IK. INV.159236, CpD. Scale bars. 1 (B, E, I) = 500 µm; 2 (A, C, D, F, G, H, J, K) = 200 µm." pageId="32" pageNumber="33">Figure 16AC, G</figureCitation>
) elongated (L/H ratio = 2.0) and sub-triangular, dorsal margin bluntly pointed, with greatest height behind the middle, anterior part of dorsal margin sloping almost straight to anterior margin, passing into it with a blunt angle, anteroventral part of anterior margin almost straight, posterior part of dorsal margin sloping towards ventral margin, the latter almost straight; LV overlapping RV along anterior, ventral and posterior sides, dorsally strongly overlapping RV; valve surface set with polygonal ridges in midventral and central part. CpD (
<figureCitation id="EA8DAAA94515FF9CFF49180DFEB62E68" box="[159,312,835,861]" captionStart="FIGURE 16" captionStartId="30.[151,250,1942,1966]" captionTargetBox="[169,1418,181,1919]" captionTargetId="figure-31@30.[169,1418,181,1919]" captionTargetPageId="30" captionText="FIGURE 16. Benincandona sakitii gen. et spec. nov., valves (AC, G, IK ♂; DF, H ♀). AH. CpRl, (AC, G: INV.159235, DF, H: INV.159237). IK. INV.159236, CpD. Scale bars. 1 (B, E, I) = 500 µm; 2 (A, C, D, F, G, H, J, K) = 200 µm." pageId="32" pageNumber="33">Figure 16IK</figureCitation>
) narrow and sharply pointed at anterior margin, more bluntly pointed at posterior side, greatest width in the middle, lateral margins rounded.
</paragraph>
<paragraph id="7209B62C4515FF9CFF1118C5FF7E2900" blockId="32.[151,1437,618,2013]" pageId="32" pageNumber="33">
LVi (
<figureCitation id="EA8DAAA94515FF9CFED418C5FE762E90" box="[258,504,907,933]" captionStart="FIGURE 17" captionStartId="31.[151,250,1630,1654]" captionTargetBox="[151,1436,181,1606]" captionTargetId="figure-17@31.[151,1436,181,1606]" captionTargetPageId="31" captionText="FIGURE 17. Benincandona sakitii gen. et spec. nov., valves (♂). A, C, D, G. LVi (A, C, D. INV.159225; G. INV.159227). B, E, F. RVi (INV.159225). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." pageId="32" pageNumber="33">Figure 17A, C, D, G</figureCitation>
) elongated and sub-triangular, dorsal margin bluntly pointed in the centre, with greatest height behind the middle, anterior part of dorsal margin sloping almost straight to anterior margin, passing into it with a blunt angle, anteroventral part almost straight, posterior part of dorsal margin sloping towards ventral margin, the latter almost straight; one ventral il without antero-ventral tooth. CMS (
<figureCitation id="EA8DAAA94515FF9CFBB318B9FB642924" box="[1125,1258,1015,1041]" captionStart="FIGURE 17" captionStartId="31.[151,250,1630,1654]" captionTargetBox="[151,1436,181,1606]" captionTargetId="figure-17@31.[151,1436,181,1606]" captionTargetPageId="31" captionText="FIGURE 17. Benincandona sakitii gen. et spec. nov., valves (♂). A, C, D, G. LVi (A, C, D. INV.159225; G. INV.159227). B, E, F. RVi (INV.159225). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." pageId="32" pageNumber="33">Figure 17G</figureCitation>
) as in the
<typeStatus id="AD0D088E4515FF9CFABA18B9FA122924" box="[1388,1436,1015,1041]" pageId="32" pageNumber="33">type</typeStatus>
species.
</paragraph>
<paragraph id="7209B62C4515FF9CFF111F71FE0629F0" blockId="32.[151,1437,618,2013]" pageId="32" pageNumber="33">
RVi (
<figureCitation id="EA8DAAA94515FF9CFED41F71FE4B296C" box="[258,453,1087,1113]" captionStart="FIGURE 17" captionStartId="31.[151,250,1630,1654]" captionTargetBox="[151,1436,181,1606]" captionTargetId="figure-17@31.[151,1436,181,1606]" captionTargetPageId="31" captionText="FIGURE 17. Benincandona sakitii gen. et spec. nov., valves (♂). A, C, D, G. LVi (A, C, D. INV.159225; G. INV.159227). B, E, F. RVi (INV.159225). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." pageId="32" pageNumber="33">Figure 17B, E, F</figureCitation>
) elongated, dorsal margin with greatest height well behind the middle, anterior part of dorsal margin sloping almost straight to anterior margin, passing into it with a blunt angle, anteroventral part straight, posterior part of dorsal margin sloping towards ventral margin, the latter almost straight; il running submarginally along ventral margin.
</paragraph>
<paragraph id="7209B62C4515FF9CFF111F81FB4C2B64" blockId="32.[151,1437,618,2013]" pageId="32" pageNumber="33">
A1 (
<figureCitation id="EA8DAAA94515FF9CFF211F81FEF929DC" box="[247,375,1231,1257]" captionStart="FIGURE 19" captionStartId="34.[151,250,1943,1967]" captionTargetBox="[156,1433,185,1915]" captionTargetId="figure-31@34.[152,1436,181,1919]" captionTargetPageId="34" captionText="FIGURE 19. Benincandona sakitii gen. et spec. nov., limbs (♂). A. A2 (INV.159225). B. A1 (INV.159225). C, D. MdCoxa (INV.159228). Scale bar = 50 µm." pageId="32" pageNumber="33">Figure 19B</figureCitation>
) consisting of seven articulated segments, one basal and six distal. Basal segment
<emphasis id="40C26A3E4515FF9CFADE1F81FA9529DD" box="[1288,1307,1231,1256]" italics="true" pageId="32" pageNumber="33">c.</emphasis>
three times as long as basal width, supporting two long ventro-subapical setae with the more apical slightly shorter than the other one; and two medium length setae on the dorsal side (one illustrated here), situated some distance from each other. First distal segment about twice the basal width, with one short dorso-apical seta. Second distal segment about the same length/width ratio as the previous segment with one very small dorso-apical seta. Third distal segment about three times as long as basal width, set with one very small apical seta in each side. Fourth distal segment about twice the basal width, with one short ventro-apical seta and one medium length dorso-apical seta, reaching beyond tip of sixth segment with more than the half the total length. Fifth distal segment about four times as long as wide, with two long dorso-apical setae and one short ventro-apical seta clearly exceeding the next segment. Sixth distal segment narrow,
<emphasis id="40C26A3E4515FF9CFE801D5DFEE72B19" box="[342,361,1555,1580]" italics="true" pageId="32" pageNumber="33">c.</emphasis>
four times as long as wide, with four apical structures: one aesthetasc Ya of intermediate length, two long setae about twice the length of Ya and one short seta, about one third the length of Ya.
</paragraph>
<paragraph id="7209B62C4515FF9DFF111D15FE042C74" blockId="32.[151,1437,618,2013]" lastBlockId="33.[151,1437,151,321]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
A2 (
<figureCitation id="EA8DAAA94515FF9CFF2C1D15FEF12B40" box="[250,383,1627,1653]" captionStart="FIGURE 19" captionStartId="34.[151,250,1943,1967]" captionTargetBox="[156,1433,185,1915]" captionTargetId="figure-31@34.[152,1436,181,1919]" captionTargetPageId="34" captionText="FIGURE 19. Benincandona sakitii gen. et spec. nov., limbs (♂). A. A2 (INV.159225). B. A1 (INV.159225). C, D. MdCoxa (INV.159228). Scale bar = 50 µm." pageId="32" pageNumber="33">Figure 19A</figureCitation>
) with a basal segment (first protopodite) broad and rounded, mid-ventrally with one seta of medium length, two subapical setae of unequal length, proximal one
<emphasis id="40C26A3E4515FF9CFC621D31FC492BAD" box="[948,967,1663,1688]" italics="true" pageId="32" pageNumber="33">c.</emphasis>
twice the length of the more apical one. Second protopodite large and stout,
<emphasis id="40C26A3E4515FF9CFDF91DEDFDCC2B89" box="[559,578,1699,1724]" italics="true" pageId="32" pageNumber="33">c.</emphasis>
twice as long as basal width, set with one long ventro-subapical seta X. No remnant of exopodite visible. Endopodite consisting of four segments with the penultimate segment divided. First endopodal segment large, more than five times as long as basal width, carrying one long aesthetasc Y (distally asymmetrically bifurcated) on the mid-ventral side (slightly under tip of segment) and two ventro-apical setae, one short and one large, hirsute in its distal 4/5. Second endopodal segment (segment 2A)
<emphasis id="40C26A3E4515FF9CFB5D1C7DFB102A79" box="[1163,1182,1843,1868]" italics="true" pageId="32" pageNumber="33">c.</emphasis>
twice as long as wide, carrying one stout dorso-apical seta and three t-setae; seta t1 long, narrow and plumose, accompanied by a long and slender aesthetasc y1; setae t2 and t3 hyper-developed into male bristles, t2 ventro-apical, t3 centro-apical. Third endopodal segment (segment 2B) smaller than previous segment and almost as long as wide, set with a complex (and sexually dimorphic) apical chaetotaxy (
<figureCitation id="EA8DAAA94515FF9CFD451C8DFC862AE8" box="[659,776,1987,2013]" captionStart="FIGURE 40" captionStartId="60.[151,250,1516,1540]" captionTargetBox="[151,1435,181,1492]" captionTargetId="figure-31@60.[151,1435,181,1492]" captionTargetPageId="60" captionText="FIGURE 40. Diagrams of apical views of the A2 of Benincandona, showing position of (sub-) terminal claws, setae and aesthetascs of the terminal (small circle) and penultimate segment (large circle). For explanations of codes, see Material and Methods (Abbreviations used in text and figures)." pageId="32" pageNumber="33">Figure 40</figureCitation>
): ventro-apically with one slender aesthetasc y2 and claw G3 transformed into short seta of almost equal length; centrally with one short seta z3; dorso-apically with z1 claw-like, length about twice as long as penultimate segment. Claw G2 and G1 about 2.5 times as long as z1, with z2 seta transformed into long claw, reaching tip of G2. Fourth, terminal, endopodal segment with one long claw (Gm), one short claw (GM), a long aesthetasc y3 with shorter accompanying seta, with fused part short, and seta g almost as long as aesthetasc y3.
</paragraph>
<caption id="26C9E6A44514FF9DFF411C48FBBD2A76" pageId="33" pageNumber="34" startId="33.[151,250,1798,1822]" targetBox="[151,1436,345,1775]" targetPageId="33" targetType="figure">
<paragraph id="7209B62C4514FF9DFF411C48FBBD2A76" blockId="33.[151,1437,1798,1859]" pageId="33" pageNumber="34">
<emphasis id="40C26A3E4514FF9DFF411C48FEAD2A2B" bold="true" box="[151,291,1798,1822]" pageId="33" pageNumber="34">FIGURE 18.</emphasis>
<taxonomicName id="B5B6CDAF4514FF9DFEFB1C49FD892A2A" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[301,519,1799,1823]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="sakitii" status="gen. et spec. nov.">
<emphasis id="40C26A3E4514FF9DFEFB1C49FD892A2A" box="[301,519,1799,1823]" italics="true" pageId="33" pageNumber="34">Benincandona sakitii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454514FF9DFDC71C49FD432A2B" box="[529,717,1799,1823]" pageId="33" pageNumber="34" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, valves (♀).
<emphasis id="40C26A3E4514FF9DFC8F1C48FC3E2A2A" bold="true" box="[857,944,1798,1823]" pageId="33" pageNumber="34">A, C, D</emphasis>
. LVi (A, C, D. INV.159226).
<emphasis id="40C26A3E4514FF9DFB241C49FAE82A2B" bold="true" box="[1266,1382,1798,1823]" pageId="33" pageNumber="34">B, E, F, G</emphasis>
. RVi (INV.159226).
<emphasis id="40C26A3E4514FF9DFEE71C64FEC52A76" bold="true" box="[305,331,1834,1859]" pageId="33" pageNumber="34">G.</emphasis>
CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A44517FF9EFF411CD9FE7C2AE1" pageId="34" pageNumber="35" startId="34.[151,250,1943,1967]" targetBox="[156,1433,185,1915]" targetPageId="34" targetType="figure">
<paragraph id="7209B62C4517FF9EFF411CD9FE7C2AE1" blockId="34.[151,1437,1943,2004]" pageId="34" pageNumber="35">
<emphasis id="40C26A3E4517FF9EFF411CD9FEAC2A9A" bold="true" box="[151,290,1943,1967]" pageId="34" pageNumber="35">FIGURE 19.</emphasis>
<taxonomicName id="B5B6CDAF4517FF9EFEFC1CD6FD8C2A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[298,514,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="sakitii" status="gen. et spec. nov.">
<emphasis id="40C26A3E4517FF9EFEFC1CD6FD8C2A85" box="[298,514,1944,1968]" italics="true" pageId="34" pageNumber="35">Benincandona sakitii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454517FF9EFDDC1CD6FD4F2A9A" box="[522,705,1944,1968]" pageId="34" pageNumber="35" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E4517FF9EFCE91CD9FCDF2A9A" bold="true" box="[831,849,1943,1967]" pageId="34" pageNumber="35">A</emphasis>
. A2 (INV.159225).
<emphasis id="40C26A3E4517FF9EFBF71CD6FBBC2A85" bold="true" box="[1057,1074,1944,1968]" pageId="34" pageNumber="35">B</emphasis>
. A1 (INV.159225).
<emphasis id="40C26A3E4517FF9EFAD41CD9FABB2A85" bold="true" box="[1282,1333,1943,1968]" pageId="34" pageNumber="35">C, D</emphasis>
. MdCoxa (INV.159228). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A44516FF9FFF411CD9FC482AE1" pageId="35" pageNumber="36" startId="35.[151,250,1943,1967]" targetBox="[185,1402,185,1915]" targetPageId="35" targetType="figure">
<paragraph id="7209B62C4516FF9FFF411CD9FC482AE1" blockId="35.[151,1436,1943,2004]" pageId="35" pageNumber="36">
<emphasis id="40C26A3E4516FF9FFF411CD9FEAB2A9A" bold="true" box="[151,293,1943,1967]" pageId="35" pageNumber="36">FIGURE 20.</emphasis>
<taxonomicName id="B5B6CDAF4516FF9FFEE61CD6FD852A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[304,523,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="sakitii" status="gen. et spec. nov.">
<emphasis id="40C26A3E4516FF9FFEE61CD6FD852A85" box="[304,523,1944,1968]" italics="true" pageId="35" pageNumber="36">Benincandona sakitii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454516FF9FFDC01CD6FD582A9A" box="[534,726,1944,1968]" pageId="35" pageNumber="36" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E4516FF9FFC8B1CD9FCE12A9A" bold="true" box="[861,879,1943,1967]" pageId="35" pageNumber="36">A</emphasis>
. Mx1-palp (INV.159225).
<emphasis id="40C26A3E4516FF9FFB581CD6FB112A85" bold="true" box="[1166,1183,1944,1968]" pageId="35" pageNumber="36">B</emphasis>
. Mx1, respiratory plate (INV.159227).
<emphasis id="40C26A3E4516FF9FFEE71CF5FECD2AE6" bold="true" box="[305,323,1979,2003]" pageId="35" pageNumber="36">C</emphasis>
. MdPalp (INV.159227). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm.
</paragraph>
</caption>
<paragraph id="7209B62C4511FF98FF111BD9FC472DCC" blockId="36.[151,1437,151,1689]" pageId="36" pageNumber="37">
MdCoxa (
<figureCitation id="EA8DAAA94511FF98FEEE1BD9FE532D84" box="[312,477,151,177]" captionStart="FIGURE 19" captionStartId="34.[151,250,1943,1967]" captionTargetBox="[156,1433,185,1915]" captionTargetId="figure-31@34.[152,1436,181,1919]" captionTargetPageId="34" captionText="FIGURE 19. Benincandona sakitii gen. et spec. nov., limbs (♂). A. A2 (INV.159225). B. A1 (INV.159225). C, D. MdCoxa (INV.159228). Scale bar = 50 µm." pageId="36" pageNumber="37">Figure 19CD</figureCitation>
) elongated, proximally sharply pointed, medially widened, distally with an obliquely widened end, set with a series (67) of strong teeth, interspaced with setae; subapically with a short seta.
<emphasis id="40C26A3E4511FF98FAEB1BF5FA1A2DE0" box="[1341,1428,187,213]" italics="true" pageId="36" pageNumber="37">Remark</emphasis>
: asymmetry illustrated possibly resulting from different positions in slide.
</paragraph>
<paragraph id="7209B62C4511FF98FF111A4DFAAB2F2C" blockId="36.[151,1437,151,1689]" pageId="36" pageNumber="37">
MdPalp (
<figureCitation id="EA8DAAA94511FF98FEE61A4DFE3D2C28" box="[304,435,259,285]" captionStart="FIGURE 20" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[185,1402,185,1915]" captionTargetId="figure-17@35.[181,1406,181,1919]" captionTargetPageId="35" captionText="FIGURE 20. Benincandona sakitii gen. et spec. nov., limbs (♂). A. Mx1-palp (INV.159225). B. Mx1, respiratory plate (INV.159227). C. MdPalp (INV.159227). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." pageId="36" pageNumber="37">Figure 20C</figureCitation>
) consisting of four segments. Basal segment ventro-apically with two hirsute setae: one long (S1), reaching tip of terminal segment and one short and stout (S2), length about one fifth of the length of S1; in between with a short flagellum-like alpha seta (thin and about 1.5x length of S2); more proximally, one smooth, medium-length seta. Second segment of sub-triangular shape, dorso-apically with two setae of unequal length, ventro-apically with a group of four setae (long and unequal size). Third segment with two unequal dorso-subapical setae, two mid-dorsal setae, one of which being the gamma seta, and ventro-apically with three setae of sub-equal length. Terminal segment sub-rectangular, apically set with one large claw, fused with the segment over its entire distal margin, accompanied by one smooth lateral seta on each side, slightly shorter than the fused claw.
</paragraph>
<paragraph id="7209B62C4511FF98FF11196DFF6B2F9C" blockId="36.[151,1437,151,1689]" pageId="36" pageNumber="37">
Mx1 (
<figureCitation id="EA8DAAA94511FF98FEDB196DFE382F08" box="[269,438,547,573]" captionStart="FIGURE 20" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[185,1402,185,1915]" captionTargetId="figure-17@35.[181,1406,181,1919]" captionTargetPageId="35" captionText="FIGURE 20. Benincandona sakitii gen. et spec. nov., limbs (♂). A. Mx1-palp (INV.159225). B. Mx1, respiratory plate (INV.159227). C. MdPalp (INV.159227). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." pageId="36" pageNumber="37">Figure 20A, B</figureCitation>
) palp with first segment carrying four long dorso-apical setae; terminal segment short and rectangular, carrying also four distal setae, some slightly claw-like. Respiratory plate (
<figureCitation id="EA8DAAA94511FF98FB401909FA962F54" box="[1174,1304,583,609]" captionStart="FIGURE 20" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[185,1402,185,1915]" captionTargetId="figure-17@35.[181,1406,181,1919]" captionTargetPageId="35" captionText="FIGURE 20. Benincandona sakitii gen. et spec. nov., limbs (♂). A. Mx1-palp (INV.159225). B. Mx1, respiratory plate (INV.159227). C. MdPalp (INV.159227). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." pageId="36" pageNumber="37">Figure 20B</figureCitation>
) elongated, carrying
<emphasis id="40C26A3E4511FF98FF2D1925FE802FB1" box="[251,270,619,644]" italics="true" pageId="36" pageNumber="37">c.</emphasis>
15 plumose respiratory rays, two short distal setae and a group of four shorter proximal setae of unequal length.
</paragraph>
<paragraph id="7209B62C4511FF98FF1119FDFD712E68" blockId="36.[151,1437,151,1689]" pageId="36" pageNumber="37">
T1 (
<figureCitation id="EA8DAAA94511FF98FF2119FDFE132FF8" box="[247,413,691,717]" captionStart="FIGURE 23" captionStartId="39.[151,250,1943,1967]" captionTargetBox="[173,1413,185,1915]" captionTargetId="figure-17@39.[170,1417,181,1919]" captionTargetPageId="39" captionText="FIGURE 23. Benincandona sakitii gen. et spec. nov., limbs (A, B ♂ and CE ♀). A. T1, Lpp (INV.159228). B. T1, Rpp (INV.159228). C. A2 (INV.159226, INV.159231). D. CR (INV.159232). E. T1 (INV.159226). Scale bar = 50 µm." pageId="36" pageNumber="37">Figure 23A, B</figureCitation>
) with basal part (protopodite) carrying respiratory plates (exopodites not shown), distally set with 3+7 setae of different morphology and length including two plumose; more proximally set with d and d setae, length of the setae d being 1.5x the length of d. Prehensile palps clearly asymmetrical with Rpp (
<figureCitation id="EA8DAAA94511FF98FAC519B5FA1A2E20" box="[1299,1428,763,789]" captionStart="FIGURE 23" captionStartId="39.[151,250,1943,1967]" captionTargetBox="[173,1413,185,1915]" captionTargetId="figure-17@39.[170,1417,181,1919]" captionTargetPageId="39" captionText="FIGURE 23. Benincandona sakitii gen. et spec. nov., limbs (A, B ♂ and CE ♀). A. T1, Lpp (INV.159228). B. T1, Rpp (INV.159228). C. A2 (INV.159226, INV.159231). D. CR (INV.159232). E. T1 (INV.159226). Scale bar = 50 µm." pageId="36" pageNumber="37">Figure 23B</figureCitation>
) slightly more swollen than Lpp (
<figureCitation id="EA8DAAA94511FF98FE281851FD0D2E0C" box="[510,643,799,825]" captionStart="FIGURE 23" captionStartId="39.[151,250,1943,1967]" captionTargetBox="[173,1413,185,1915]" captionTargetId="figure-17@39.[170,1417,181,1919]" captionTargetPageId="39" captionText="FIGURE 23. Benincandona sakitii gen. et spec. nov., limbs (A, B ♂ and CE ♀). A. T1, Lpp (INV.159228). B. T1, Rpp (INV.159228). C. A2 (INV.159226, INV.159231). D. CR (INV.159232). E. T1 (INV.159226). Scale bar = 50 µm." pageId="36" pageNumber="37">Figure 23A</figureCitation>
); one-segmented, distally sigmoid hook-like, both palps set with a short sensory organ and proximally with two subequal setae.
</paragraph>
<paragraph id="7209B62C4511FF98FF111829FAFB2900" blockId="36.[151,1437,151,1689]" pageId="36" pageNumber="37">
T2 (
<figureCitation id="EA8DAAA94511FF98FF251829FEFC2EB4" box="[243,370,871,897]" captionStart="FIGURE 21" captionStartId="37.[151,250,1981,2005]" captionTargetBox="[177,1420,212,1949]" captionTargetId="figure-17@37.[166,1422,181,1957]" captionTargetPageId="37" captionText="FIGURE 21. Benincandona sakitii et spec. nov., limbs (♂). A. T3 (INV.159225). B. T2 (INV.159225). C. CR (INV.159225). D. CRAtt (INV.15922). Scale bar = 50 µm." pageId="36" pageNumber="37">Figure 21B</figureCitation>
) with 4-segmented endopodite (penultimate segment divided), elongated, without basal setae (d1 and d2). First endopodal segment without any setae, length
<emphasis id="40C26A3E4511FF98FCE318C5FCC62E91" box="[821,840,907,932]" italics="true" pageId="36" pageNumber="37">c.</emphasis>
four times as long as basal width. Second endopodal segment with length
<emphasis id="40C26A3E4511FF98FE5418E1FE1B2EFD" box="[386,405,943,968]" italics="true" pageId="36" pageNumber="37">c.</emphasis>
4x basal width, set with one short ventro-apical seta f. Third endopodal segment with about the same length/width ratio as the previous segment and with one ventro-subapical seta g, about 1/3 the length of following segment. Terminal segment about 1.5 times as long as basal width, apically two short setae, h1 ventral and h3 (length being the half of the length of h1) dorsal and a long slender claw (h2), distally set with short setulae.
</paragraph>
<paragraph id="7209B62C4511FF98FF111F71FE1B2838" blockId="36.[151,1437,151,1689]" pageId="36" pageNumber="37">
T3 (
<figureCitation id="EA8DAAA94511FF98FF231F71FEF6296C" box="[245,376,1087,1113]" captionStart="FIGURE 21" captionStartId="37.[151,250,1981,2005]" captionTargetBox="[177,1420,212,1949]" captionTargetId="figure-17@37.[166,1422,181,1957]" captionTargetPageId="37" captionText="FIGURE 21. Benincandona sakitii et spec. nov., limbs (♂). A. T3 (INV.159225). B. T2 (INV.159225). C. CR (INV.159225). D. CRAtt (INV.15922). Scale bar = 50 µm." pageId="36" pageNumber="37">Figure 21A</figureCitation>
) with first segment carrying only one medium length seta d1. Second (knee-) segment set with d2-seta. Third segment about 4x as long as wide, not carrying any seta. Fourth segment narrower,
<emphasis id="40C26A3E4511FF98FB0F1F2DFB622949" box="[1241,1260,1123,1148]" italics="true" pageId="36" pageNumber="37">c.</emphasis>
7x basal width, slightly longer than the previous segment, carrying one short subapical seta g. Terminal segment well-separated from penultimate segment, length about twice basal width and carrying three setae: one long and reflexed (h3), tip clearly reaching base of the third segment, and two ventrally directed and curved setae h1 &amp; h2, the former
<emphasis id="40C26A3E4511FF98FA801F81FAE729DD" box="[1366,1385,1231,1256]" italics="true" pageId="36" pageNumber="37">c.</emphasis>
half the length of the latter.
</paragraph>
<paragraph id="7209B62C4511FF98FF111E59FEB9284C" blockId="36.[151,1437,151,1689]" pageId="36" pageNumber="37">
CR (
<figureCitation id="EA8DAAA94511FF98FF2D1E59FEF32804" box="[251,381,1303,1329]" captionStart="FIGURE 21" captionStartId="37.[151,250,1981,2005]" captionTargetBox="[177,1420,212,1949]" captionTargetId="figure-17@37.[166,1422,181,1957]" captionTargetPageId="37" captionText="FIGURE 21. Benincandona sakitii et spec. nov., limbs (♂). A. T3 (INV.159225). B. T2 (INV.159225). C. CR (INV.159225). D. CRAtt (INV.15922). Scale bar = 50 µm." pageId="36" pageNumber="37">Figure 21C</figureCitation>
) with stout ramus; claw Ga sigmoid, well-developed and stout, length ratio of ramus and claw Ga approximately 1.6; setae Sp (ventro-subapical), Sa (dorso-apical) and claw Gp (first ventro-subapical) reduced to small setae.
</paragraph>
<paragraph id="7209B62C4511FF98FF111ECDFE1D28F4" blockId="36.[151,1437,151,1689]" pageId="36" pageNumber="37">
CRAtt (
<figureCitation id="EA8DAAA94511FF98FEC91ECDFE2D28A8" box="[287,419,1411,1437]" captionStart="FIGURE 21" captionStartId="37.[151,250,1981,2005]" captionTargetBox="[177,1420,212,1949]" captionTargetId="figure-17@37.[166,1422,181,1957]" captionTargetPageId="37" captionText="FIGURE 21. Benincandona sakitii et spec. nov., limbs (♂). A. T3 (INV.159225). B. T2 (INV.159225). C. CR (INV.159225). D. CRAtt (INV.15922). Scale bar = 50 µm." pageId="36" pageNumber="37">Figure 21D</figureCitation>
) long and stout, distally bifurcated and subapically with additional lateral branch; rounded fist-shaped apical end.
</paragraph>
<paragraph id="7209B62C4511FF98FF111E85FE032B64" blockId="36.[151,1437,151,1689]" pageId="36" pageNumber="37">
Hp (
<figureCitation id="EA8DAAA94511FF98FF2A1E85FE2A28D0" box="[252,420,1483,1509]" captionStart="FIGURE 22" captionStartId="38.[151,250,1943,1967]" captionTargetBox="[195,1394,191,1912]" captionTargetId="figure-31@38.[185,1402,181,1919]" captionTargetPageId="38" captionText="FIGURE 22. Benincandona sakitii gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159227). BD. Hp (INV.159225). B. External view (drawing). C. Internal view (photograph with interference contrast). D. Internal view (photograph without interference contrast). Scale bar = 50 µm." pageId="36" pageNumber="37">Figure 22BD</figureCitation>
) with elongated lobe ls, apically rounded and with a pronounced hump-shaped ventral protrusion, reaching well beyond asymmetrically rounded medium shield (ms), both lobes overlapping, not diverging; lobe h shorter and sub-rectangular; internal anatomy (
<figureCitation id="EA8DAAA94511FF98FC571D5DFBA82B18" box="[897,1062,1555,1581]" captionStart="FIGURE 22" captionStartId="38.[151,250,1943,1967]" captionTargetBox="[195,1394,191,1912]" captionTargetId="figure-31@38.[185,1402,181,1919]" captionTargetPageId="38" captionText="FIGURE 22. Benincandona sakitii gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159227). BD. Hp (INV.159225). B. External view (drawing). C. Internal view (photograph with interference contrast). D. Internal view (photograph without interference contrast). Scale bar = 50 µm." pageId="36" pageNumber="37">Figure 22C, D</figureCitation>
) with postlabyrinthal spermiduct with one circular coil.
</paragraph>
<paragraph id="7209B62C4511FF98FF111D15FD732BAC" blockId="36.[151,1437,151,1689]" pageId="36" pageNumber="37">
Zenker organ (
<figureCitation id="EA8DAAA94511FF98FEA41D15FE772B40" box="[370,505,1627,1653]" captionStart="FIGURE 22" captionStartId="38.[151,250,1943,1967]" captionTargetBox="[195,1394,191,1912]" captionTargetId="figure-31@38.[185,1402,181,1919]" captionTargetPageId="38" captionText="FIGURE 22. Benincandona sakitii gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159227). BD. Hp (INV.159225). B. External view (drawing). C. Internal view (photograph with interference contrast). D. Internal view (photograph without interference contrast). Scale bar = 50 µm." pageId="36" pageNumber="37">Figure 22A</figureCitation>
) elongated, with
<emphasis id="40C26A3E4511FF98FD131D15FD562B41" box="[709,728,1627,1652]" italics="true" pageId="36" pageNumber="37">c.</emphasis>
seven spinous whirls, with an elongated central tube joining posterior funnel-shaped end plate to anterior end plate.
</paragraph>
<paragraph id="7209B62C4511FF98FF411D88FE182BD5" blockId="36.[151,1436,1734,1977]" box="[151,406,1734,1760]" pageId="36" pageNumber="37">
<emphasis id="40C26A3E4511FF98FF411D88FE182BD5" bold="true" box="[151,406,1734,1760]" pageId="36" pageNumber="37">Description of female</emphasis>
</paragraph>
<paragraph id="7209B62C4511FF98FF411DA5FB4D2A30" blockId="36.[151,1436,1734,1977]" box="[151,1219,1771,1797]" pageId="36" pageNumber="37">
Cp and valves (
<figureCitation id="EA8DAAA94511FF98FE921DA5FD9C2A30" box="[324,530,1771,1797]" captionStart="FIGURE 16" captionStartId="30.[151,250,1942,1966]" captionTargetBox="[169,1418,181,1919]" captionTargetId="figure-31@30.[169,1418,181,1919]" captionTargetPageId="30" captionText="FIGURE 16. Benincandona sakitii gen. et spec. nov., valves (AC, G, IK ♂; DF, H ♀). AH. CpRl, (AC, G: INV.159235, DF, H: INV.159237). IK. INV.159236, CpD. Scale bars. 1 (B, E, I) = 500 µm; 2 (A, C, D, F, G, H, J, K) = 200 µm." pageId="36" pageNumber="37">Figures 16DF, H</figureCitation>
;
<figureCitation id="EA8DAAA94511FF98FDC81DA5FDFE2A30" box="[542,624,1771,1797]" captionStart="FIGURE 18" captionStartId="33.[151,250,1798,1822]" captionTargetBox="[151,1436,345,1775]" captionTargetId="figure-109@33.[151,1436,345,1775]" captionTargetPageId="33" captionText="FIGURE 18. Benincandona sakitii gen. et spec. nov., valves (♀). A, C, D. LVi (A, C, D. INV.159226). B, E, F, G. RVi (INV.159226). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." pageId="36" pageNumber="37">18AG</figureCitation>
) as in the male, without obvious sexual dimorphism.
</paragraph>
<paragraph id="7209B62C4511FF98FF111C41FB072A1C" blockId="36.[151,1436,1734,1977]" box="[199,1161,1807,1833]" pageId="36" pageNumber="37">
Limbs (
<figureCitation id="EA8DAAA94511FF98FECB1C41FE312A1C" box="[285,447,1807,1833]" captionStart="FIGURE 23" captionStartId="39.[151,250,1943,1967]" captionTargetBox="[173,1413,185,1915]" captionTargetId="figure-17@39.[170,1417,181,1919]" captionTargetPageId="39" captionText="FIGURE 23. Benincandona sakitii gen. et spec. nov., limbs (A, B ♂ and CE ♀). A. T1, Lpp (INV.159228). B. T1, Rpp (INV.159228). C. A2 (INV.159226, INV.159231). D. CR (INV.159232). E. T1 (INV.159226). Scale bar = 50 µm." pageId="36" pageNumber="37">Figure 23CE</figureCitation>
) largely as in the male, but with sexually dimorphic A2 and T1.
</paragraph>
<paragraph id="7209B62C4511FF98FF111C7DFC232A8C" blockId="36.[151,1436,1734,1977]" pageId="36" pageNumber="37">
A2 (
<figureCitation id="EA8DAAA94511FF98FF2F1C7DFEF22A78" box="[249,380,1843,1869]" captionStart="FIGURE 23" captionStartId="39.[151,250,1943,1967]" captionTargetBox="[173,1413,185,1915]" captionTargetId="figure-17@39.[170,1417,181,1919]" captionTargetPageId="39" captionText="FIGURE 23. Benincandona sakitii gen. et spec. nov., limbs (A, B ♂ and CE ♀). A. T1, Lpp (INV.159228). B. T1, Rpp (INV.159228). C. A2 (INV.159226, INV.159231). D. CR (INV.159232). E. T1 (INV.159226). Scale bar = 50 µm." pageId="36" pageNumber="37">Figure 23C</figureCitation>
) with second endopodal segment undivided, with two t-setae, t1 (long, narrow and plumose) and t2 (thin and smooth); apical chaetotaxy (
<figureCitation id="EA8DAAA94511FF98FD451C19FC8A2A44" box="[659,772,1879,1905]" captionStart="FIGURE 40" captionStartId="60.[151,250,1516,1540]" captionTargetBox="[151,1435,181,1492]" captionTargetId="figure-31@60.[151,1435,181,1492]" captionTargetPageId="60" captionText="FIGURE 40. Diagrams of apical views of the A2 of Benincandona, showing position of (sub-) terminal claws, setae and aesthetascs of the terminal (small circle) and penultimate segment (large circle). For explanations of codes, see Material and Methods (Abbreviations used in text and figures)." pageId="36" pageNumber="37">Figure 40</figureCitation>
) with z1claw-like, z2 and z3 being both setae, z1 twice as long as z2 and z3; claws G1, G2, G3 and GM all reaching about the same point; seta g and aesthetasc y3 as in the male. Small claw Gm in female homologous to large claw in the male.
</paragraph>
<caption id="26C9E6A44510FF99FF411CF3FDA02ACF" pageId="37" pageNumber="38" startId="37.[151,250,1981,2005]" targetBox="[177,1420,212,1949]" targetPageId="37" targetType="figure">
<paragraph id="7209B62C4510FF99FF411CF3FDA02ACF" blockId="37.[151,1436,1981,2042]" pageId="37" pageNumber="38">
<emphasis id="40C26A3E4510FF99FF411CF3FE912AE0" bold="true" box="[151,287,1981,2005]" pageId="37" pageNumber="38">FIGURE 21.</emphasis>
<taxonomicName id="B5B6CDAF4510FF99FEF31CF3FE752AE0" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[293,507,1981,2005]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="sakitii">
<emphasis id="40C26A3E4510FF99FEF31CF3FE752AE0" box="[293,507,1981,2005]" italics="true" pageId="37" pageNumber="38">Benincandona sakitii</emphasis>
</taxonomicName>
et spec. nov., limbs (♂).
<emphasis id="40C26A3E4510FF99FD231CF3FC892AE0" bold="true" box="[757,775,1981,2005]" pageId="37" pageNumber="38">A</emphasis>
. T3 (INV.159225).
<emphasis id="40C26A3E4510FF99FC181CF0FC512AE3" bold="true" box="[974,991,1982,2006]" pageId="37" pageNumber="38">B</emphasis>
. T2 (INV.159225).
<emphasis id="40C26A3E4510FF99FB731CF3FB392AE0" bold="true" box="[1189,1207,1981,2005]" pageId="37" pageNumber="38">C</emphasis>
. CR (INV.159225).
<emphasis id="40C26A3E4510FF99FA521CF0FA182AE3" bold="true" box="[1412,1430,1982,2006]" pageId="37" pageNumber="38">D</emphasis>
. CRAtt (INV.15922). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A44513FF9AFF411CD9FDB32ACD" pageId="38" pageNumber="39" startId="38.[151,250,1943,1967]" targetBox="[195,1394,191,1912]" targetPageId="38" targetType="figure">
<paragraph id="7209B62C4513FF9AFF411CD9FDB32ACD" blockId="38.[151,1437,1943,2040]" pageId="38" pageNumber="39">
<emphasis id="40C26A3E4513FF9AFF411CD9FEAD2A9A" bold="true" box="[151,291,1943,1967]" pageId="38" pageNumber="39">FIGURE 22.</emphasis>
<taxonomicName id="B5B6CDAF4513FF9AFEFB1CD9FD882A9A" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[301,518,1943,1967]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="sakitii" status="gen. et spec. nov.">
<emphasis id="40C26A3E4513FF9AFEFB1CD9FD882A9A" box="[301,518,1943,1967]" italics="true" pageId="38" pageNumber="39">Benincandona sakitii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454513FF9AFDC61CD6FD452A9A" box="[528,715,1944,1968]" pageId="38" pageNumber="39" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E4513FF9AFC9B1CD9FCD12A9A" bold="true" box="[845,863,1943,1967]" pageId="38" pageNumber="39">A</emphasis>
. Zenker organ (INV.159227).
<emphasis id="40C26A3E4513FF9AFB771CD6FB5E2A85" bold="true" box="[1185,1232,1944,1968]" pageId="38" pageNumber="39">BD</emphasis>
. Hp (INV.159225).
<emphasis id="40C26A3E4513FF9AFF411CF2FF262AE1" bold="true" box="[151,168,1980,2004]" pageId="38" pageNumber="39">B</emphasis>
. External view (drawing).
<emphasis id="40C26A3E4513FF9AFE141CF5FE5A2AE6" bold="true" box="[450,468,1979,2003]" pageId="38" pageNumber="39">C</emphasis>
. Internal view (photograph with interference contrast).
<emphasis id="40C26A3E4513FF9AFBCE1CF2FBA42AE1" bold="true" box="[1048,1066,1980,2004]" pageId="38" pageNumber="39">D</emphasis>
. Internal view (photograph without interference contrast). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A44512FF9BFF411CD9FA9F2AE1" pageId="39" pageNumber="40" startId="39.[151,250,1943,1967]" targetBox="[173,1413,185,1915]" targetPageId="39" targetType="figure">
<paragraph id="7209B62C4512FF9BFF411CD9FA9F2AE1" blockId="39.[151,1436,1943,2004]" pageId="39" pageNumber="40">
<emphasis id="40C26A3E4512FF9BFF411CD9FEAD2A9A" bold="true" box="[151,291,1943,1967]" pageId="39" pageNumber="40">FIGURE 23.</emphasis>
<taxonomicName id="B5B6CDAF4512FF9BFEFA1CD6FD882A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[300,518,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="sakitii" status="gen. et spec. nov.">
<emphasis id="40C26A3E4512FF9BFEFA1CD6FD882A85" box="[300,518,1944,1968]" italics="true" pageId="39" pageNumber="40">Benincandona sakitii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454512FF9BFDD91CD6FD442A9A" box="[527,714,1944,1968]" pageId="39" pageNumber="40" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (A, B ♂ and CE ♀).
<emphasis id="40C26A3E4512FF9BFBDF1CD9FB952A9A" bold="true" box="[1033,1051,1943,1967]" pageId="39" pageNumber="40">A</emphasis>
. T1, Lpp (INV.159228).
<emphasis id="40C26A3E4512FF9BFAF01CD6FAB92A85" bold="true" box="[1318,1335,1944,1968]" pageId="39" pageNumber="40">B</emphasis>
. T1, Rpp (INV.159228).
<emphasis id="40C26A3E4512FF9BFEE71CF5FECD2AE6" bold="true" box="[305,323,1979,2003]" pageId="39" pageNumber="40">C</emphasis>
. A2 (INV.159226, INV.159231).
<emphasis id="40C26A3E4512FF9BFD401CF2FD262AE1" bold="true" box="[662,680,1980,2004]" pageId="39" pageNumber="40">D</emphasis>
. CR (INV.159232).
<emphasis id="40C26A3E4512FF9BFCA11CF2FC062AE1" bold="true" box="[887,904,1980,2004]" pageId="39" pageNumber="40">E</emphasis>
. T1 (INV.159226). Scale bar = 50 µm.
</paragraph>
</caption>
<paragraph id="7209B62C451DFF94FF111BD9FC112DCC" blockId="40.[151,1436,151,429]" pageId="40" pageNumber="41">
T1 (
<figureCitation id="EA8DAAA9451DFF94FF231BD9FEF82D84" box="[245,374,151,177]" captionStart="FIGURE 23" captionStartId="39.[151,250,1943,1967]" captionTargetBox="[173,1413,185,1915]" captionTargetId="figure-17@39.[170,1417,181,1919]" captionTargetPageId="39" captionText="FIGURE 23. Benincandona sakitii gen. et spec. nov., limbs (A, B ♂ and CE ♀). A. T1, Lpp (INV.159228). B. T1, Rpp (INV.159228). C. A2 (INV.159226, INV.159231). D. CR (INV.159232). E. T1 (INV.159226). Scale bar = 50 µm." pageId="40" pageNumber="41">Figure 23E</figureCitation>
) with basal part with
<emphasis id="40C26A3E451DFF94FDBB1BD9FD0E2D85" box="[621,640,151,176]" italics="true" pageId="40" pageNumber="41">c.</emphasis>
10 distal setae of unequal length; d, d and b setae present; respiratory plate reduced to about three rays, two short, one medium size about twice as long as others. Endopodite with a broad and short palp, set with three subequal but short apical setae (h1h3).
</paragraph>
<paragraph id="7209B62C451DFF94FF111A4DFD862C28" blockId="40.[151,1436,151,429]" box="[199,520,259,285]" pageId="40" pageNumber="41">
<emphasis id="40C26A3E451DFF94FF111A4DFE0E2C28" bold="true" box="[199,384,259,285]" pageId="40" pageNumber="41">Measurements:</emphasis>
See
<tableCitation id="3F348397451DFF94FE601A4DFD862C28" box="[438,520,259,285]" captionStart="TABLE 3" captionStartId="19.[152,244,1592,1616]" captionText="TABLE 3. Measurements of illustrated specimens of Benincandona gen. et spec. nov., using SEM." pageId="40" pageNumber="41">Table 3</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A7451DFF94FF111A68FDD32C74" box="[199,605,294,321]" pageId="40" pageNumber="41" type="diagnosis">
<paragraph id="7209B62C451DFF94FF111A68FDD32C74" blockId="40.[151,1436,151,429]" box="[199,605,294,321]" pageId="40" pageNumber="41">
<emphasis id="40C26A3E451DFF94FF111A68FE5B2C75" bold="true" box="[199,469,294,320]" pageId="40" pageNumber="41">Differential Diagnosis:</emphasis>
See
<tableCitation id="3F348397451DFF94FDDC1A69FDD32C74" box="[522,605,295,321]" captionStart="TABLE 4" captionStartId="20.[152,245,1665,1689]" captionText="TABLE 4. Differential character states between the five new species of Benincandona gen. nov." pageId="40" pageNumber="41">Table 4</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A7451DFF94FF111A05FEA02C98" pageId="40" pageNumber="41" type="biology_ecology">
<paragraph id="7209B62C451DFF94FF111A05FE612C51" blockId="40.[151,1436,151,429]" box="[199,495,330,357]" pageId="40" pageNumber="41">
<emphasis id="40C26A3E451DFF94FF111A05FE612C51" bold="true" box="[199,495,330,357]" pageId="40" pageNumber="41">Ecology and distribution</emphasis>
</paragraph>
<paragraph id="7209B62C451DFF94FF111A21FEA02C98" blockId="40.[151,1436,151,429]" pageId="40" pageNumber="41">
This subterranean species is presently known from its
<typeStatus id="AD0D088E451DFF94FCFC1A21FCD42CBC" box="[810,858,367,393]" pageId="40" pageNumber="41">type</typeStatus>
locality and a second, closely situated station, only (see
<tableCitation id="3F348397451DFF94FF1D1ADDFEAF2C98" box="[203,289,403,429]" captionStart="TABLE 2" captionStartId="6.[152,244,1634,1658]" captionText="TABLE 2. Measurements of physical and chemical water parameters and variables at the time of well fauna collection. WT= water temperature (°C), EC = electrical conductivity (µS cm-1), DO = dissolved oxygen (mg L-1). NM= no measurement." pageId="40" pageNumber="41">Table 2</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="39A39419F87B1A8612B1FA980191609B" ID-DOI="10.11646/zootaxa.5503.1.1" ID-ISSN="1175-5334" ID-ZooBank="7385E4AA-D7AD-4E79-A05C-684319E0843B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726051594773" checkinUser="plazi" docAuthor="Hotèkpo, Sourou Joseph, Schön, Isa &amp; Martens, Koen" docDate="2024" docId="FA1F073A453DFFB5FF411E74FC6F2A50" docLanguage="en" docName="zootaxa.5503.1.1.pdf" docOrigin="Zootaxa 5503 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5503.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Benincandoninae Hotèkpo &amp; Schön &amp; Martens 2024, subfam. nov." docType="treatment" docUuid="0D13483E-A799-40BC-8382-0B62AA61DEE4" docUuidSource="ZooBank" docVersion="1" lastPageNumber="10" masterDocId="06267F424535FFBCFFD61B4EFF8E2D35" masterDocTitle="An endemic species flock of Candonidae Kaufmann, 1900 (Crustacea, Ostracoda) from wells in Benin (Africa), with the description of a new subfamily, a new genus and five new species" masterLastPageNumber="72" masterPageNumber="1" pageNumber="9" updateTime="1726051738210" updateUser="GgImagineBatch">
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<mods:title id="13280BAFB0ED69F5758D1EF7104CFA79">An endemic species flock of Candonidae Kaufmann, 1900 (Crustacea, Ostracoda) from wells in Benin (Africa), with the description of a new subfamily, a new genus and five new species</mods:title>
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<mods:namePart id="78C4943CC6350CE0C543E3E3273FBBF9">Hotèkpo, Sourou Joseph</mods:namePart>
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<mods:affiliation id="EB197509940B553C6EE1171E285BD38E">Centre de Recherche pour la lutte contre les Maladies Infectieuses Tropicales (CReMIT / TIDRC), Université dAbomey-Calavi, Bénin jshotekpo @ gmail. com; https: // orcid. org / 0000 - 0001 - 9091 - 433 X &amp; Royal Belgian Institute of Natural Sciences, Freshwater Biology, Vautierstraat 29, B- 1000 Brussels, Belgium &amp; University of Hasselt, Research Group Zoology, Agoralaan Building D, B- 3590 Diepenbeek, Belgium ischoen @ naturalsciences. be; https: // orcid. org / 0000 - 0001 - 9269 - 6487</mods:affiliation>
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<mods:namePart id="00494C7D52625320DF2C136794A87093">Schön, Isa</mods:namePart>
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<mods:affiliation id="69185319D9171ECFF93BA2CD0E677274">Royal Belgian Institute of Natural Sciences, Freshwater Biology, Vautierstraat 29, B- 1000 Brussels, Belgium &amp; University of Hasselt, Research Group Zoology, Agoralaan Building D, B- 3590 Diepenbeek, Belgium ischoen @ naturalsciences. be; https: // orcid. org / 0000 - 0001 - 9269 - 6487</mods:affiliation>
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<mods:namePart id="A45DE0746F7512F5268C483760A1BB5D">Martens, Koen</mods:namePart>
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<mods:affiliation id="84078FECEB392E0212C1C6452C832D0A">Royal Belgian Institute of Natural Sciences, Freshwater Biology, Vautierstraat 29, B- 1000 Brussels, Belgium &amp; Ghent University, Dept. Biology, K. L. Ledeganckstraat 35, B- 9000 Ghent, Belgium kmartens @ naturalsciences. be; https: // orcid. org / 0000 - 0001 - 8680 - 973 X * Corresponding author</mods:affiliation>
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<mods:date id="414F9680EAB61567CC0CECB4B91208CF">2024</mods:date>
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<heading id="29410140453DFFB4FF411E74FC2C2860" bold="true" box="[151,930,1338,1365]" fontSize="11" level="2" pageId="8" pageNumber="9" reason="3">
<emphasis id="40C26A3E453DFFB4FF411E74FC2C2860" bold="true" box="[151,930,1338,1365]" pageId="8" pageNumber="9">
Subfamily
<taxonomicName id="B5B6CDAF453DFFB4FEC91E74FE762861" authority="Hotekpo &amp; Martens" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[287,504,1338,1364]" class="Ostracoda" family="Candonidae" kingdom="Animalia" order="Podocopida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="subFamily" status="subfam. nov." subFamily="Benincandoninae">Benincandoninae</taxonomicName>
<taxonomicNameLabel id="5BF1D745453DFFB4FE291E74FD102860" box="[511,670,1338,1365]" pageId="8" pageNumber="9" rank="subFamily">subfam. nov.</taxonomicNameLabel>
Hotèkpo &amp; Martens
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genus:
<taxonomicName id="B5B6CDAF453DFFB4FEF81EE8FE5628F4" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[302,472,1446,1473]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="40C26A3E453DFFB4FEF81EE8FE5628F4" bold="true" box="[302,472,1446,1473]" italics="true" pageId="8" pageNumber="9">Benincandona</emphasis>
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<taxonomicNameLabel id="5BF1D745453DFFB4FE081EE9FDC928F4" box="[478,583,1447,1473]" pageId="8" pageNumber="9" rank="genus">gen. nov.</taxonomicNameLabel>
Hotèkpo &amp; Martens
</emphasis>
</paragraph>
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<subSubSection id="3AACE5A7453DFFB5FF411D5CFC452B14" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="7209B62C453DFFB4FF411D5CFE9E2B19" blockId="8.[151,272,1554,1580]" box="[151,272,1554,1580]" pageId="8" pageNumber="9">
<emphasis id="40C26A3E453DFFB4FF411D5CFE9E2B19" bold="true" box="[151,272,1554,1580]" pageId="8" pageNumber="9">Diagnosis</emphasis>
</paragraph>
<paragraph id="7209B62C453DFFB4FF411D15FDBE2AA0" blockId="8.[151,1437,1627,1941]" pageId="8" pageNumber="9">Cp and valves whitish, opaque, in general laterally flattened; in some species with external linear ornamentation. LV overlapping RV on all sides, but especially dorsally. Both valves without obvious marginal structures, only with ventral remnants of an il. A1 seven-segmented, without fused segments, but with reduced chaetotaxy on some segments. A2 with aesthetasc Y long and slender, sometimes distally bifurcated. MdPalp with seta s2 much shorter than seta s1; one large end claw fused with terminal segment. Mx1-palp with terminal segment quadrate to rectangular. T1 with setae d and d subequal to unequal in length; male prehensile palps subequal. T2 without d1, d2 and e setae, setae f and g present. T3 with seta dp absent, seta d1 long, seta d2 short, long or absent. CR with well-developed ramus, one claw Ga and three small setae (Gp, Sa and Sp). CRatt simple and linear, with few short side branches. Hp with three distal lobes.</paragraph>
<caption id="26C9E6A4453CFFB5FF411F16FDF129A1" pageId="9" pageNumber="10" startId="9.[151,250,1112,1136]" targetBox="[152,1436,181,1088]" targetPageId="9" targetType="figure">
<paragraph id="7209B62C453CFFB5FF411F16FDF129A1" blockId="9.[151,1436,1112,1173]" pageId="9" pageNumber="10">
<emphasis id="40C26A3E453CFFB5FF411F16FE982945" bold="true" box="[151,278,1112,1136]" pageId="9" pageNumber="10">FIGURE 2.</emphasis>
Map of the Ouémé catchment (and its position in Benin — right inlets), indicating localities where species of
<taxonomicName id="B5B6CDAF453CFFB5FF411F32FEA729A1" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[151,297,1148,1172]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus" status="gen. et spec. nov.">
<emphasis id="40C26A3E453CFFB5FF411F32FEA729A1" box="[151,297,1148,1172]" italics="true" pageId="9" pageNumber="10">Benincandona</emphasis>
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<taxonomicNameLabel id="5BF1D745453CFFB5FEE61F33FE5129A1" box="[304,479,1149,1173]" pageId="9" pageNumber="10" rank="species">gen. et spec. nov.</taxonomicNameLabel>
were collected.
</paragraph>
</caption>
<paragraph id="7209B62C453CFFB5FF411FA8FE2D2835" blockId="9.[151,419,1254,1280]" box="[151,419,1254,1280]" pageId="9" pageNumber="10">
<emphasis id="40C26A3E453CFFB5FF411FA8FE2D2835" bold="true" box="[151,419,1254,1280]" pageId="9" pageNumber="10">Differential diagnosis</emphasis>
</paragraph>
<paragraph id="7209B62C453CFFB5FF411E61FF7728A4" blockId="9.[151,1436,1327,1425]" pageId="9" pageNumber="10">
The new subfamily can be distinguished from the other subfamilies in the
<taxonomicName id="B5B6CDAF453CFFB5FC181E61FBDA287C" authorityName="Kaufmann" authorityYear="1900" box="[974,1108,1327,1353]" class="Ostracoda" family="Candonidae" kingdom="Animalia" order="Podocopida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="family">Candonidae</taxonomicName>
by a unique set of characters, including the seven-segmented A1 and the CR with a single distal claw. See Discussion for further details and analysis.
</paragraph>
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Genus
<taxonomicName id="B5B6CDAF453CFFB5FF381EACFE2E28C8" authority="Hotekpo &amp; Martens" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[238,416,1506,1533]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="40C26A3E453CFFB5FF381EACFE2E28C8" bold="true" box="[238,416,1506,1533]" italics="true" pageId="9" pageNumber="10">Benincandona</emphasis>
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<taxonomicNameLabel id="5BF1D745453CFFB5FE711EADFD9B28C8" box="[423,533,1507,1533]" pageId="9" pageNumber="10" rank="genus">gen. nov.</taxonomicNameLabel>
Hotèkpo &amp; Martens
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<paragraph id="7209B62C453CFFB5FF411D01FB232B5C" blockId="9.[151,1436,1614,1893]" box="[151,1197,1614,1641]" pageId="9" pageNumber="10">
<emphasis id="40C26A3E453CFFB5FF411D01FEBB2B5D" bold="true" box="[151,309,1614,1641]" pageId="9" pageNumber="10">
<typeStatus id="AD0D088E453CFFB5FF411D01FF5C2B5C" box="[151,210,1615,1641]" pageId="9" pageNumber="10">Type</typeStatus>
species:
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<taxonomicName id="B5B6CDAF453CFFB5FEEA1D01FDB82B5D" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[316,566,1615,1641]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="martini" status="gen. et spec. nov.">
<emphasis id="40C26A3E453CFFB5FEEA1D01FDB82B5D" box="[316,566,1615,1641]" italics="true" pageId="9" pageNumber="10">Benincandona martini</emphasis>
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<taxonomicNameLabel id="5BF1D745453CFFB5FDE81D01FD712B5C" box="[574,767,1615,1641]" pageId="9" pageNumber="10" rank="species">gen. et spec. nov.</taxonomicNameLabel>
Hotèkpo &amp; Martens
<emphasis id="40C26A3E453CFFB5FC3A1D00FC7B2B5D" bold="true" box="[1004,1013,1614,1640]" pageId="9" pageNumber="10">(</emphasis>
here designated)
</paragraph>
<paragraph id="7209B62C453CFFB5FF111D3CFAF52B84" blockId="9.[151,1436,1614,1893]" pageId="9" pageNumber="10">
<emphasis id="40C26A3E453CFFB5FF111D3CFEFA2BB9" bold="true" box="[199,372,1650,1676]" pageId="9" pageNumber="10">Other species:</emphasis>
<taxonomicName id="B5B6CDAF453CFFB5FEAB1D3DFD812BB8" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[381,527,1651,1677]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ibikounlei" status="gen. et spec. nov">
<emphasis id="40C26A3E453CFFB5FEAB1D3DFD812BB8" box="[381,527,1651,1677]" italics="true" pageId="9" pageNumber="10">B. ibikounlei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745453CFFB5FDCE1D3DFD522BB8" box="[536,732,1651,1677]" pageId="9" pageNumber="10" rank="species">gen. et spec. nov</taxonomicNameLabel>
Hotèkpo &amp; Martens,
<taxonomicName id="B5B6CDAF453CFFB5FC0D1D3DFBCE2BB8" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[987,1088,1651,1677]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sakitii" status="gen. et spec. nov.">
<emphasis id="40C26A3E453CFFB5FC0D1D3DFBCE2BB8" box="[987,1088,1651,1677]" italics="true" pageId="9" pageNumber="10">B. sakitii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745453CFFB5FB9D1D3DFA9D2BB8" box="[1099,1299,1651,1677]" pageId="9" pageNumber="10" rank="species">gen. et spec. nov.</taxonomicNameLabel>
Hotèkpo &amp; Martens,
<taxonomicName id="B5B6CDAF453CFFB5FED71DD9FEE62B84" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[257,360,1687,1713]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="cebios" status="gen. et spec. nov.">
<emphasis id="40C26A3E453CFFB5FED71DD9FEE62B84" box="[257,360,1687,1713]" italics="true" pageId="9" pageNumber="10">B. cebios</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745453CFFB5FEB91DD9FDBE2B84" box="[367,560,1687,1713]" pageId="9" pageNumber="10" rank="species">gen. et spec. nov.</taxonomicNameLabel>
Hotèkpo &amp; Martens,
<taxonomicName id="B5B6CDAF453CFFB5FCF21DD9FC4A2B84" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[804,964,1687,1713]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="trapezoides" status="gen. et spec. nov.">
<emphasis id="40C26A3E453CFFB5FCF21DD9FC4A2B84" box="[804,964,1687,1713]" italics="true" pageId="9" pageNumber="10">B. trapezoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745453CFFB5FC1D1DD9FB022B84" box="[971,1164,1687,1713]" pageId="9" pageNumber="10" rank="species">gen. et spec. nov.</taxonomicNameLabel>
Hotèkpo &amp; Martens.
</paragraph>
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<subSubSection id="3AACE5A7453CFFB5FF111DF5FE9E2A28" pageId="9" pageNumber="10" type="etymology">
<paragraph id="7209B62C453CFFB5FF111DF5FE9E2A28" blockId="9.[151,1436,1614,1893]" pageId="9" pageNumber="10">
<emphasis id="40C26A3E453CFFB5FF111DF5FEDC2BE0" bold="true" box="[199,338,1723,1749]" pageId="9" pageNumber="10">Etymology:</emphasis>
The prefix of the genus name is derived from the country from where the genus is described,
<collectingCountry id="0AA1F6BC453CFFB5FF411D91FF542BCC" box="[151,218,1759,1785]" name="Benin" pageId="9" pageNumber="10">Benin</collectingCountry>
in West Africa. The suffix is derived from the genus
<taxonomicName id="B5B6CDAF453CFFB5FC961D91FC292BCC" box="[832,935,1759,1785]" class="Ostracoda" family="Candonidae" genus="Candona" kingdom="Animalia" order="Podocopida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="40C26A3E453CFFB5FC961D91FC292BCC" box="[832,935,1759,1785]" italics="true" pageId="9" pageNumber="10">Candona</emphasis>
</taxonomicName>
, thus indicating the taxonomic affinity with this genus.
</paragraph>
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<paragraph id="7209B62C453CFFB5FF111C68FD6D2A74" blockId="9.[151,1436,1614,1893]" box="[199,739,1830,1857]" pageId="9" pageNumber="10">
<emphasis id="40C26A3E453CFFB5FF111C68FECA2A75" bold="true" box="[199,324,1830,1856]" pageId="9" pageNumber="10">Diagnosis:</emphasis>
As for the subfamily, see discussion.
</paragraph>
<paragraph id="7209B62C453CFFB5FF111C04FC6F2A50" blockId="9.[151,1436,1614,1893]" box="[199,993,1866,1893]" pageId="9" pageNumber="10">
<emphasis id="40C26A3E453CFFB5FF111C04FE5B2A51" bold="true" box="[199,469,1866,1892]" pageId="9" pageNumber="10">Differential Diagnosis:</emphasis>
The new subfamily is at present monogeneric.
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<mods:title id="F09BBAEAB105CB39E41C8097174221C9">An endemic species flock of Candonidae Kaufmann, 1900 (Crustacea, Ostracoda) from wells in Benin (Africa), with the description of a new subfamily, a new genus and five new species</mods:title>
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<mods:namePart id="A001626DE895000E268F9DADC410D049">Hotèkpo, Sourou Joseph</mods:namePart>
<mods:nameIdentifier id="876B3DD7156308FD95EA95C56324B3D3" type="ORCID">0000-0001-9091-433X</mods:nameIdentifier>
<mods:affiliation id="6B6006E5A0B28DD0D3DDCB5B2360C22C">Centre de Recherche pour la lutte contre les Maladies Infectieuses Tropicales (CReMIT / TIDRC), Université dAbomey-Calavi, Bénin jshotekpo @ gmail. com; https: // orcid. org / 0000 - 0001 - 9091 - 433 X &amp; Royal Belgian Institute of Natural Sciences, Freshwater Biology, Vautierstraat 29, B- 1000 Brussels, Belgium &amp; University of Hasselt, Research Group Zoology, Agoralaan Building D, B- 3590 Diepenbeek, Belgium ischoen @ naturalsciences. be; https: // orcid. org / 0000 - 0001 - 9269 - 6487</mods:affiliation>
<mods:nameIdentifier id="6CB5DE6711073A1B626BE0DEC123D4DB" type="email">jshotekpo@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="853AA93946A68D4222BB230B8FE90664" type="personal">
<mods:role id="32CEC4CDE0829EB4CBBD571485EE2358">
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</mods:role>
<mods:namePart id="A5C37B732E417FCA09AB27695A6FC88D">Schön, Isa</mods:namePart>
<mods:nameIdentifier id="4C3FE4EFB4EE49D6A53A8CEA3B0CBDD3" type="ORCID">0000-0001-9269-6487</mods:nameIdentifier>
<mods:affiliation id="5AA8587E1AA60BE72AD0A4D672ABD249">Royal Belgian Institute of Natural Sciences, Freshwater Biology, Vautierstraat 29, B- 1000 Brussels, Belgium &amp; University of Hasselt, Research Group Zoology, Agoralaan Building D, B- 3590 Diepenbeek, Belgium ischoen @ naturalsciences. be; https: // orcid. org / 0000 - 0001 - 9269 - 6487</mods:affiliation>
<mods:nameIdentifier id="14B882B5CF0BEAFF591A931C6A278827" type="email">ischoen@naturalsciences.be</mods:nameIdentifier>
</mods:name>
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<mods:role id="E7FE9CB445E1ED5429BE3ECBE4A13CE2">
<mods:roleTerm id="845672A6AB1F68497136D31B97DB1107">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="66B6DB7BAA0E0FA2885342D68FEC9FE2">Martens, Koen</mods:namePart>
<mods:nameIdentifier id="D7D1A32C44C3B9A584F7A0BE86ADF5D5" type="ORCID">0000-0001-8680-973X</mods:nameIdentifier>
<mods:affiliation id="FD7BB2A679A6D6D7965C8A996CD8041A">Royal Belgian Institute of Natural Sciences, Freshwater Biology, Vautierstraat 29, B- 1000 Brussels, Belgium &amp; Ghent University, Dept. Biology, K. L. Ledeganckstraat 35, B- 9000 Ghent, Belgium kmartens @ naturalsciences. be; https: // orcid. org / 0000 - 0001 - 8680 - 973 X * Corresponding author</mods:affiliation>
<mods:nameIdentifier id="EE0054258A544FEBB8989C664A2B3922" type="email">kmartens@naturalsciences.be</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="694E7FF01BAAFE6FE08CA64CAAD47294">text</mods:typeOfResource>
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<mods:title id="D4F308B673F1AE8080B8670827C9A345">Zootaxa</mods:title>
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<mods:part id="3537066349040E80312BEF3FF6A00431">
<mods:date id="230B722C51F73865E65B6FA443156247">2024</mods:date>
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<mods:number id="9509CFAC9A1FF7B122EE3F7B8CB7DA1E">2024-09-04</mods:number>
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<mods:detail id="72722DAE91AF38F80F7DB3732BA01BF3" type="volume">
<mods:number id="A88C5D0FFA611454666BECF8A958E36D">5503</mods:number>
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<mods:number id="210A69A8D9B04C5D2EB107B898178A4A">1</mods:number>
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<mods:url id="F62D6BD4EB1F94213156517A83AC3B27">http://dx.doi.org/10.11646/zootaxa.5503.1.1</mods:url>
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<mods:identifier id="49AA743A0B837442E528259684BA65B8" type="DOI">10.11646/zootaxa.5503.1.1</mods:identifier>
<mods:identifier id="9BD4B52C2DD4588A5945AA1C05607D27" type="ISSN">1175-5334</mods:identifier>
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<treatment id="FA1F073A453FFFAFFF411BD9FA1E2AD6" LSID="urn:lsid:plazi:treatment:FA1F073A453FFFAFFF411BD9FA1E2AD6" httpUri="http://treatment.plazi.org/id/FA1F073A453FFFAFFF411BD9FA1E2AD6" lastPageId="19" lastPageNumber="20" pageId="10" pageNumber="11">
<subSubSection id="3AACE5A7453FFFB6FF411BD9FC4E2DE3" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="7209B62C453FFFB6FF411BD9FC4E2DE3" blockId="10.[151,960,150,249]" pageId="10" pageNumber="11">
<heading id="29410140453FFFB6FF411BD9FC082D84" bold="true" box="[151,902,150,178]" fontSize="11" level="2" pageId="10" pageNumber="11" reason="3">
<emphasis id="40C26A3E453FFFB6FF411BD9FC082D84" bold="true" box="[151,902,150,178]" pageId="10" pageNumber="11">
<taxonomicName id="B5B6CDAF453FFFB6FF411BD9FE242D84" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[151,426,151,178]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="martini" status="gen. et spec. nov.">
<emphasis id="40C26A3E453FFFB6FF411BD9FE242D84" bold="true" box="[151,426,151,178]" italics="true" pageId="10" pageNumber="11">Benincandona martini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745453FFFB6FE671BD9FD0C2D84" box="[433,642,151,177]" pageId="10" pageNumber="11" rank="species">gen. et spec. nov.</taxonomicNameLabel>
Hotèkpo &amp; Martens
</emphasis>
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
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<subSubSection id="3AACE5A7453FFFB6FF411B91FE012DCC" box="[151,399,223,249]" pageId="10" pageNumber="11" type="description">
<paragraph id="7209B62C453FFFB6FF411B91FE012DCC" blockId="10.[151,960,150,249]" box="[151,399,223,249]" pageId="10" pageNumber="11">
(
<figureCitation id="EA8DAAA9453FFFB6FF491B91FEA92DCC" box="[159,295,223,249]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStart-4="FIGURE 7" captionStart-5="FIGURE 8" captionStart-6="FIGURE 9" captionStartId-0="11.[151,250,1933,1957]" captionStartId-1="12.[151,250,1944,1968]" captionStartId-2="13.[151,250,1943,1967]" captionStartId-3="14.[151,250,1981,2005]" captionStartId-4="16.[151,250,1981,2005]" captionStartId-5="17.[151,250,1943,1967]" captionStartId-6="18.[151,250,1943,1967]" captionTargetBox-0="[172,1415,181,1910]" captionTargetBox-1="[157,1430,181,1920]" captionTargetBox-2="[170,1415,185,1915]" captionTargetBox-3="[169,1422,182,1942]" captionTargetBox-4="[186,1404,198,1941]" captionTargetBox-5="[196,1391,191,1914]" captionTargetBox-6="[172,1415,185,1915]" captionTargetId-0="figure-17@11.[172,1415,178,1910]" captionTargetId-1="figure-31@12.[157,1430,181,1920]" captionTargetId-2="figure-17@13.[167,1419,181,1919]" captionTargetId-3="figure-31@14.[166,1422,181,1957]" captionTargetId-4="figure-31@16.[166,1422,181,1957]" captionTargetId-5="figure-17@17.[186,1401,181,1919]" captionTargetId-6="figure-31@18.[168,1419,181,1919]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionTargetPageId-3="14" captionTargetPageId-4="16" captionTargetPageId-5="17" captionTargetPageId-6="18" captionText-0="FIGURE 3. Benincandona martini gen. et spec. nov., valves (A, C, EH, J ♂; B, D, I ♀). AD, I. CpRl, (A, C. INV.159208; B, D, I. INV.159211). E. INV.159209. CpD. F. INV.159210. CpV. G. INV.159200. LVi. H. INV.159200. RVi. J. INV.159200. CMS. Scale bars. 1 (A, B, E, F, G, H) = 500 µm; 2 (C, D) = 200 µm; 3 (I, J) = 50 µm." captionText-1="FIGURE 4. Benincandona martini gen. et spec. nov., valves (AD: INV.159202 ♂ and EH. INV.159201 ♀) A, E. LVi. B, F. RVi, C, D, G, H. CMS. Scale bars. 1 (A, B, E, F) = 500 µm; 2 (C, D, G, H) = 50 µm." captionText-2="FIGURE 5. Benincandona martini gen. et spec. nov., limbs (♂). A. A1 (INV.159203). B, C. MdCoxa, (INV.159202). D. A2 (INV.159200 INV.159203). Scale bar = 50 µm." captionText-3="FIGURE 6. Benincandona martini gen. et spec. nov., limbs (♂). A. Mx1 (Chaetotaxy incomplete) (INV.159200). B. Mx1, respiratory plate (INV.159202). C. MdPalp (INV.159204 INV.159203). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." captionText-4="FIGURE 7. Benincandona martini gen. et spec. nov., limbs (♂). A. T2 (INV.159200). B. CR (INV.159200). C. CRatt (INV.159200). D. T3 (INV.159200). Scale bar = 50 µm." captionText-5="FIGURE 8. Benincandona martini gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159200). BD. Hp (INV.159200). B. External view (drawing). C. Internal view (photograph with interference contrast). D. Internal view (photograph without interference contrast). Scale bar = 50 µm." captionText-6="FIGURE 9. Benincandona martini gen. et spec. nov., limbs (AB ♂ and CE ♀). A. T1, Lpp (INV.159202). B. T1, Rpp (INV.159202). C. A2 (INV.159205, INV.159207). D. CR (INV.159205). E. T1 (INV.159200). Scale bar = 50 µm." pageId="10" pageNumber="11">Figures 39</figureCitation>
,
<figureCitation id="EA8DAAA9453FFFB6FEE21B91FE092DCC" box="[308,391,223,249]" captionStart="FIGURE 41" captionStartId="61.[151,250,1113,1137]" captionTargetBox="[151,1436,181,1089]" captionTargetId="figure-17@61.[151,1436,181,1089]" captionTargetPageId="61" captionText="FIGURE 41. Benincandona gen. et spec. nov., CpRl. (A, C, E, F, G, I, K, N, O, Q ♂ and B, D, H, J, L, M, P, R ♀). AD. B. martini. EF. B. ibikounlei. GJ. B. sakitii. KN. B. cebios. OR. B. trapezoides. Scale bar = 500 µm." pageId="10" pageNumber="11">41AD</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A7453FFFB6FF411A69FBD22C98" pageId="10" pageNumber="11" type="etymology">
<paragraph id="7209B62C453FFFB6FF411A69FE962C74" blockId="10.[151,1436,295,430]" box="[151,280,295,321]" pageId="10" pageNumber="11">
<emphasis id="40C26A3E453FFFB6FF411A69FE962C74" bold="true" box="[151,280,295,321]" pageId="10" pageNumber="11">Etymology</emphasis>
</paragraph>
<paragraph id="7209B62C453FFFB6FF411A05FBD22C98" blockId="10.[151,1436,295,430]" pageId="10" pageNumber="11">
The species is named after Dr Patrick Martin (RBINS,
<collectingRegion id="B07278CE453FFFB6FCDD1A05FCE32C50" box="[779,877,331,357]" country="Belgium" name="Region de Bruxelles-Capitale" pageId="10" pageNumber="11">Brussels</collectingRegion>
,
<collectingCountry id="0AA1F6BC453FFFB6FCAF1A05FC552C50" box="[889,987,331,357]" name="Belgium" pageId="10" pageNumber="11">Belgium</collectingCountry>
) in acknowledgement of his significant contributions to groundwater biodiversity research in
<collectingCountry id="0AA1F6BC453FFFB6FD2A1A21FCCC2CBC" box="[764,834,367,393]" name="Benin" pageId="10" pageNumber="11">Benin</collectingCountry>
, and with gratitude for his continuous support of the career of SJH and in recognition of decades of collegiality with IS and KM at RBINS.
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A7453FFFB6FF411A9AFE552FF2" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="7209B62C453FFFB6FF411A9AFE842CDB" blockId="10.[151,1437,468,711]" box="[151,266,468,494]" pageId="10" pageNumber="11">
<emphasis id="40C26A3E453FFFB6FF411A9AFE842CDB" bold="true" box="[151,266,468,494]" pageId="10" pageNumber="11">Diagnosis</emphasis>
</paragraph>
<paragraph id="7209B62C453FFFB6FF411AB6FE552FF2" blockId="10.[151,1437,468,711]" pageId="10" pageNumber="11">CpRl elongated (L/H = 2.0), sub-triangular, with greatest height in the middle; LV dorsally extending beyond RV, anterior and posterior margins evenly rounded, surface with ridges and polygonal ornamentation at the mid-ventral side. CpD and CpV narrow, with greatest width in the middle, lateral sides evenly rounded, anterior and posterior extremities pointed. A2 with seta X long, reaching beyond tip of aesthetasc Y. Second palp segment of Mx1 with basal W almost equalling L. T3 with short seta d2. Hp with ls and ms with diverging positions, ls relatively short, reaching slightly beyond ms.</paragraph>
</subSubSection>
<subSubSection id="3AACE5A7453FFFB6FF4119A0FF742864" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="7209B62C453FFFB6FF4119A0FEB02E32" blockId="10.[151,318,749,776]" box="[151,318,749,776]" pageId="10" pageNumber="11">
<emphasis id="40C26A3E453FFFB6FF4119A0FEB02E32" bold="true" box="[151,318,749,776]" pageId="10" pageNumber="11">Type material</emphasis>
</paragraph>
<paragraph id="7209B62C453FFFB6FF411861FE8A2E7C" blockId="10.[151,1282,815,878]" box="[151,260,815,841]" pageId="10" pageNumber="11">
<emphasis id="40C26A3E453FFFB6FF411861FE8A2E7C" bold="true" box="[151,260,815,841]" pageId="10" pageNumber="11">
<typeStatus id="AD0D088E453FFFB6FF411861FE8A2E7C" box="[151,260,815,841]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7209B62C453FFFB6FF41181DFA8C2E5B" blockId="10.[151,1282,815,878]" box="[151,1282,851,878]" pageId="10" pageNumber="11">
<materialsCitation id="C2DEBC71453FFFB6FF41181DFA8C2E5B" box="[151,1282,851,878]" collectionCode="INV" pageId="10" pageNumber="11" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="64B07DA5453FFFB6FF41181DFF4D2E5B" box="[151,195,851,878]" count="1" pageId="10" pageNumber="11" type="male">1 ♂</specimenCount>
, with soft parts dissected in glycerine in a sealed slide, and with valves stored dry (
<collectionCode id="14A72EE9453FFFB6FBB0181DFB182E58" box="[1126,1174,851,877]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="10" pageNumber="11" type="Herbarium">INV</collectionCode>
.159200).
</materialsCitation>
</paragraph>
<paragraph id="7209B62C453FFFB6FF4118DAFF722E9B" blockId="10.[151,780,916,979]" box="[151,252,916,942]" pageId="10" pageNumber="11">
<emphasis id="40C26A3E453FFFB6FF4118DAFF722E9B" bold="true" box="[151,252,916,942]" pageId="10" pageNumber="11">
<typeStatus id="AD0D088E453FFFB6FF4118DAFF722E9B" box="[151,252,916,942]" pageId="10" pageNumber="11" type="allotype">Allotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7209B62C453FFFB6FF4118F7FC822EE6" blockId="10.[151,780,916,979]" box="[151,780,952,979]" pageId="10" pageNumber="11">
<materialsCitation id="C2DEBC71453FFFB6FF4118F7FC822EE6" box="[151,780,952,979]" collectionCode="INV" pageId="10" pageNumber="11" specimenCount="1" specimenCount-female="1" typeStatus="allotype">
<specimenCount id="64B07DA5453FFFB6FF4118F7FF4F2EE6" box="[151,193,953,979]" count="1" pageId="10" pageNumber="11" type="female">1 ♀</specimenCount>
, dissected and stored as the
<typeStatus id="AD0D088E453FFFB6FE2918F6FDEF2EE7" box="[511,609,952,978]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
(
<collectionCode id="14A72EE9453FFFB6FDA618F6FD282EE6" box="[624,678,952,979]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="10" pageNumber="11" type="Herbarium">INV</collectionCode>
.159201).
</materialsCitation>
</paragraph>
<paragraph id="7209B62C453FFFB6FF4118B4FE812921" blockId="10.[151,1436,1018,1189]" box="[151,271,1018,1044]" pageId="10" pageNumber="11">
<emphasis id="40C26A3E453FFFB6FF4118B4FE812921" bold="true" box="[151,271,1018,1044]" pageId="10" pageNumber="11">
<typeStatus id="AD0D088E453FFFB6FF4118B4FE812921" box="[151,271,1018,1044]" pageId="10" pageNumber="11" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7209B62C453FFFB6FF411F50FDF92990" blockId="10.[151,1436,1018,1189]" pageId="10" pageNumber="11">
<materialsCitation id="C2DEBC71453FFFB6FF411F50FCC1290C" box="[151,847,1054,1081]" collectionCode="INV" pageId="10" pageNumber="11" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="64B07DA5453FFFB6FF411F50FF45290D" box="[151,203,1054,1080]" count="1" pageId="10" pageNumber="11" type="male">1 ♂</specimenCount>
dissected and stored as the
<typeStatus id="AD0D088E453FFFB6FDEB1F50FD11290D" box="[573,671,1054,1080]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
(
<collectionCode id="14A72EE9453FFFB6FD611F50FD62290C" box="[695,748,1054,1081]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="10" pageNumber="11" type="Herbarium">INV</collectionCode>
.159202)
</materialsCitation>
;
<materialsCitation id="C2DEBC71453FFFB6FCB21F50FEA32968" collectionCode="INV" pageId="10" pageNumber="11" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="64B07DA5453FFFB6FCB21F50FC23290D" box="[868,941,1054,1080]" count="2" pageId="10" pageNumber="11" type="male">2 ♂♂</specimenCount>
dissected as the
<typeStatus id="AD0D088E453FFFB6FB461F50FB7C290D" box="[1168,1266,1054,1080]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
(
<collectionCode id="14A72EE9453FFFB6FADC1F50FAB0290C" box="[1290,1342,1054,1081]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="10" pageNumber="11" type="Herbarium">INV</collectionCode>
.159203, INV.159204)
</materialsCitation>
;
<materialsCitation id="C2DEBC71453FFFB6FEE91F0CFBF72968" box="[319,1145,1090,1117]" collectionCode="INV" pageId="10" pageNumber="11" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<specimenCount id="64B07DA5453FFFB6FEE91F0CFE0A2969" box="[319,388,1090,1116]" count="3" pageId="10" pageNumber="11" type="female">3 ♀♀</specimenCount>
dissected as the male (
<collectionCode id="14A72EE9453FFFB6FD701F0CFD542968" box="[678,730,1090,1117]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="10" pageNumber="11" type="Herbarium">INV</collectionCode>
.159205, INV.159206, INV.159207)
</materialsCitation>
;
<materialsCitation id="C2DEBC71453FFFB6FB5D1F0CFBBE29B4" collectionCode="INV" pageId="10" pageNumber="11" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="64B07DA5453FFFB6FB5D1F0CFB5E2969" box="[1163,1232,1090,1116]" count="3" pageId="10" pageNumber="11" type="male">3 ♂♂</specimenCount>
carapaces stored dry in micropalaeontological slides (
<collectionCode id="14A72EE9453FFFB6FD811F28FD0529B4" box="[599,651,1126,1153]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="10" pageNumber="11" type="Herbarium">INV</collectionCode>
.159208, INV.159209, INV.159210)
</materialsCitation>
;
<materialsCitation id="C2DEBC71453FFFB6FB931F28FDFD2990" collectionCode="INV" pageId="10" pageNumber="11" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="64B07DA5453FFFB6FB931F28FBF629B5" box="[1093,1144,1126,1152]" count="1" pageId="10" pageNumber="11" type="female">1 ♀</specimenCount>
carapace stored dry in micropalaeontological slide (
<collectionCode id="14A72EE9453FFFB6FE0A1FC4FD9F2990" box="[476,529,1162,1189]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="10" pageNumber="11" type="Herbarium">INV</collectionCode>
.159211)
</materialsCitation>
.
</paragraph>
<paragraph id="7209B62C453FFFB6FF411F85FEBE29D0" blockId="10.[151,1437,1227,1361]" box="[151,304,1227,1253]" pageId="10" pageNumber="11">
<emphasis id="40C26A3E453FFFB6FF411F85FEBE29D0" bold="true" box="[151,304,1227,1253]" pageId="10" pageNumber="11">Type locality</emphasis>
</paragraph>
<paragraph id="7209B62C453FFFB6FF411FA1FF742864" blockId="10.[151,1437,1227,1361]" pageId="10" pageNumber="11">
<materialsCitation id="C2DEBC71453FFFB6FF411FA1FB89283C" box="[151,1031,1263,1290]" country="Benin" location="Adjohoun District" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Oueme">
<collectingCountry id="0AA1F6BC453FFFB6FF411FA1FF54283C" box="[151,218,1263,1289]" name="Benin" pageId="10" pageNumber="11">Benin</collectingCountry>
,
<collectingRegion id="B07278CE453FFFB6FF301FA1FE4C283F" box="[230,450,1263,1290]" country="Benin" name="Oueme" pageId="10" pageNumber="11">Ouémé Department</collectingRegion>
,
<location id="7769E0F7453FFFB6FE1F1FA1FD1B283C" LSID="urn:lsid:plazi:treatment:FA1F073A453FFFAFFF411BD9FA1E2AD6:7769E0F7453FFFB6FE1F1FA1FD1B283C" box="[457,661,1263,1289]" country="Benin" name="Adjohoun District" pageId="10" pageNumber="11" stateProvince="Oueme">Adjohoun District</location>
, town of Azowlissé, located in the
</materialsCitation>
<materialsCitation id="C2DEBC71453FFFB6FBDA1FA1FF782864" collectingDate="2016-08-27" collectingDateMax="2016-08-29" collectingDateMin="2016-08-27" collectorName="Leg. &amp; S. J. Hotekpo" country="Benin" latitude="6.66605" location="Oueme" longLatPrecision="1" longitude="2.4920833" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Oueme">
<collectingRegion id="B07278CE453FFFB6FBDA1FA1FBD3283C" box="[1036,1117,1263,1289]" country="Benin" name="Oueme" pageId="10" pageNumber="11">Ouémé</collectingRegion>
catchment area, coordinates:
<geoCoordinate id="1782D0EB453FFFB6FF411E5DFEAA2818" box="[151,292,1299,1325]" degrees="6" direction="north" minutes="39.963" orientation="latitude" pageId="10" pageNumber="11" precision="1" value="6.66605">6° 39.963N</geoCoordinate>
,
<geoCoordinate id="1782D0EB453FFFB6FEF91E5DFE36281B" box="[303,440,1299,1326]" degrees="2" direction="east" minutes="29.525" orientation="longitude" pageId="10" pageNumber="11" precision="1" value="2.4920833">2° 29.525E</geoCoordinate>
,
<collectorName id="DF43D3FA453FFFB6FE151E5AFE74281B" box="[451,506,1300,1326]" pageId="10" pageNumber="11">Leg.</collectorName>
:
<collectorName id="DF43D3FA453FFFB6FDD21E5DFD182818" box="[516,662,1299,1325]" pageId="10" pageNumber="11">S.J. Hotèkpo</collectorName>
, collected on
<date id="060890EC453FFFB6FCF91E5DFBBE2818" box="[815,1072,1299,1325]" pageId="10" pageNumber="11" value="2016-08-27" valueMax="2016-08-29" valueMin="2016-08-27">
<collectingDate id="164C6904453FFFB6FCF91E5DFBBE2818" box="[815,1072,1299,1325]" pageId="10" pageNumber="11" value="2016-08-27" valueMax="2016-08-29" valueMin="2016-08-27">27 and 29 August 2016</collectingDate>
</date>
(Locality code BVS
<quantity id="B54E1BC9453FFFB6FAC11E5DFAC02818" box="[1303,1358,1299,1325]" metricMagnitude="-1" metricUnit="m" metricValue="4.826" pageId="10" pageNumber="11" unit="in" value="19.0">19 in</quantity>
<tableCitation id="3F348397453FFFB6FA851E5DFF2B2864" captionStart="TABLE 1" captionStartId="5.[152,244,929,955]" captionText="TABLE 1. Localities where Benincandona gen. nov. sp. nov. were collected, coordinates and hydro-morphological characteristics. In all stations, the water was clear and was used for both drinking water and household purposes by the riparian villages." pageId="10" pageNumber="11">Tables 1</tableCitation>
and
<tableCitation id="3F348397453FFFB6FF0A1E79FF632864" box="[220,237,1335,1361]" captionStart="TABLE 2" captionStartId="6.[152,244,1634,1658]" captionText="TABLE 2. Measurements of physical and chemical water parameters and variables at the time of well fauna collection. WT= water temperature (°C), EC = electrical conductivity (µS cm-1), DO = dissolved oxygen (mg L-1). NM= no measurement." pageId="10" pageNumber="11">2</tableCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A7453FFFAFFF411E36FD8A28A8" lastPageId="19" lastPageNumber="20" pageId="10" pageNumber="11" type="description">
<paragraph id="7209B62C453FFFB6FF411E36FE0F28A6" blockId="10.[151,1437,1400,2039]" box="[151,385,1400,1427]" pageId="10" pageNumber="11">
<emphasis id="40C26A3E453FFFB6FF411E36FE0F28A6" bold="true" box="[151,385,1400,1427]" pageId="10" pageNumber="11">Description of male</emphasis>
</paragraph>
<paragraph id="7209B62C453FFFB6FF411ED2FB592BBA" blockId="10.[151,1437,1400,2039]" pageId="10" pageNumber="11">
CpRl (
<figureCitation id="EA8DAAA9453FFFB6FF091ED2FEFC2882" box="[223,370,1436,1463]" captionStart="FIGURE 3" captionStartId="11.[151,250,1933,1957]" captionTargetBox="[172,1415,181,1910]" captionTargetId="figure-17@11.[172,1415,178,1910]" captionTargetPageId="11" captionText="FIGURE 3. Benincandona martini gen. et spec. nov., valves (A, C, EH, J ♂; B, D, I ♀). AD, I. CpRl, (A, C. INV.159208; B, D, I. INV.159211). E. INV.159209. CpD. F. INV.159210. CpV. G. INV.159200. LVi. H. INV.159200. RVi. J. INV.159200. CMS. Scale bars. 1 (A, B, E, F, G, H) = 500 µm; 2 (C, D) = 200 µm; 3 (I, J) = 50 µm." pageId="10" pageNumber="11">Figure 3A, C</figureCitation>
) elongated and sub-triangular, with anterior margin slightly more broadly rounded than posterior margin, dorsal margin bluntly pointed, with greatest height in the middle, anterior part sloping almost straight to anterior margin, posterior part sloping almost straight to posterior margin, ventral margin almost straight; LV slightly overlapping RV along anterior, ventral and posterior sides, dorsally strongly overlapping RV; external ornamentation consisting of linear ridges (
<figureCitation id="EA8DAAA9453FFFB6FE061D62FDE42B72" box="[464,618,1580,1607]" captionStart="FIGURE 3" captionStartId="11.[151,250,1933,1957]" captionTargetBox="[172,1415,181,1910]" captionTargetId="figure-17@11.[172,1415,178,1910]" captionTargetPageId="11" captionText="FIGURE 3. Benincandona martini gen. et spec. nov., valves (A, C, EH, J ♂; B, D, I ♀). AD, I. CpRl, (A, C. INV.159208; B, D, I. INV.159211). E. INV.159209. CpD. F. INV.159210. CpV. G. INV.159200. LVi. H. INV.159200. RVi. J. INV.159200. CMS. Scale bars. 1 (A, B, E, F, G, H) = 500 µm; 2 (C, D) = 200 µm; 3 (I, J) = 50 µm." pageId="10" pageNumber="11">Figure 3AD</figureCitation>
) and polygonal structures (
<figureCitation id="EA8DAAA9453FFFB6FC741D62FB802B72" box="[930,1038,1580,1607]" captionStart="FIGURE 3" captionStartId="11.[151,250,1933,1957]" captionTargetBox="[172,1415,181,1910]" captionTargetId="figure-17@11.[172,1415,178,1910]" captionTargetPageId="11" captionText="FIGURE 3. Benincandona martini gen. et spec. nov., valves (A, C, EH, J ♂; B, D, I ♀). AD, I. CpRl, (A, C. INV.159208; B, D, I. INV.159211). E. INV.159209. CpD. F. INV.159210. CpV. G. INV.159200. LVi. H. INV.159200. RVi. J. INV.159200. CMS. Scale bars. 1 (A, B, E, F, G, H) = 500 µm; 2 (C, D) = 200 µm; 3 (I, J) = 50 µm." pageId="10" pageNumber="11">Figure 3I</figureCitation>
) in mid-ventral position of valves. CpD (
<figureCitation id="EA8DAAA9453FFFB6FF0C1D1EFEC52B5E" box="[218,331,1616,1643]" captionStart="FIGURE 3" captionStartId="11.[151,250,1933,1957]" captionTargetBox="[172,1415,181,1910]" captionTargetId="figure-17@11.[172,1415,178,1910]" captionTargetPageId="11" captionText="FIGURE 3. Benincandona martini gen. et spec. nov., valves (A, C, EH, J ♂; B, D, I ♀). AD, I. CpRl, (A, C. INV.159208; B, D, I. INV.159211). E. INV.159209. CpD. F. INV.159210. CpV. G. INV.159200. LVi. H. INV.159200. RVi. J. INV.159200. CMS. Scale bars. 1 (A, B, E, F, G, H) = 500 µm; 2 (C, D) = 200 µm; 3 (I, J) = 50 µm." pageId="10" pageNumber="11">Figure 3E</figureCitation>
) and CpV (
<figureCitation id="EA8DAAA9453FFFB6FE1C1D1EFDB42B5E" box="[458,570,1616,1643]" captionStart="FIGURE 3" captionStartId="11.[151,250,1933,1957]" captionTargetBox="[172,1415,181,1910]" captionTargetId="figure-17@11.[172,1415,178,1910]" captionTargetPageId="11" captionText="FIGURE 3. Benincandona martini gen. et spec. nov., valves (A, C, EH, J ♂; B, D, I ♀). AD, I. CpRl, (A, C. INV.159208; B, D, I. INV.159211). E. INV.159209. CpD. F. INV.159210. CpV. G. INV.159200. LVi. H. INV.159200. RVi. J. INV.159200. CMS. Scale bars. 1 (A, B, E, F, G, H) = 500 µm; 2 (C, D) = 200 µm; 3 (I, J) = 50 µm." pageId="10" pageNumber="11">Figure 3F</figureCitation>
) narrow and pointed on both anterior and posterior sides, greatest width in the middle, lateral margins evenly rounded; LV forming a pronounced overlap over RV in the centre.
</paragraph>
<paragraph id="7209B62C453FFFB6FF111DD6FB8C2A77" blockId="10.[151,1437,1400,2039]" pageId="10" pageNumber="11">
LVi (
<figureCitation id="EA8DAAA9453FFFB6FED71DD6FE0D2B86" box="[257,387,1688,1715]" captionStart="FIGURE 3" captionStartId="11.[151,250,1933,1957]" captionTargetBox="[172,1415,181,1910]" captionTargetId="figure-17@11.[172,1415,178,1910]" captionTargetPageId="11" captionText="FIGURE 3. Benincandona martini gen. et spec. nov., valves (A, C, EH, J ♂; B, D, I ♀). AD, I. CpRl, (A, C. INV.159208; B, D, I. INV.159211). E. INV.159209. CpD. F. INV.159210. CpV. G. INV.159200. LVi. H. INV.159200. RVi. J. INV.159200. CMS. Scale bars. 1 (A, B, E, F, G, H) = 500 µm; 2 (C, D) = 200 µm; 3 (I, J) = 50 µm." pageId="10" pageNumber="11">Figures 3G</figureCitation>
,
<figureCitation id="EA8DAAA9453FFFB6FE421DD7FE552B86" box="[404,475,1689,1715]" captionStart="FIGURE 4" captionStartId="12.[151,250,1944,1968]" captionTargetBox="[157,1430,181,1920]" captionTargetId="figure-31@12.[157,1430,181,1920]" captionTargetPageId="12" captionText="FIGURE 4. Benincandona martini gen. et spec. nov., valves (AD: INV.159202 ♂ and EH. INV.159201 ♀) A, E. LVi. B, F. RVi, C, D, G, H. CMS. Scale bars. 1 (A, B, E, F) = 500 µm; 2 (C, D, G, H) = 50 µm." pageId="10" pageNumber="11">4A, C</figureCitation>
) elongated and triangular, with anterior margin slightly more broadly rounded than posterior margin, dorsal margin bluntly pointed, with greatest height in the middle, overlap present at that site, anterior part sloping almost straight to anterior margin, posterior part sloping almost straight to posterior margin, ventral margin almost straight, with a slight inward curl in the middle; CMS (
<figureCitation id="EA8DAAA9453FFFB6FC261C4AFBED2A2A" box="[1008,1123,1796,1823]" captionStart="FIGURE 4" captionStartId="12.[151,250,1944,1968]" captionTargetBox="[157,1430,181,1920]" captionTargetId="figure-31@12.[157,1430,181,1920]" captionTargetPageId="12" captionText="FIGURE 4. Benincandona martini gen. et spec. nov., valves (AD: INV.159202 ♂ and EH. INV.159201 ♀) A, E. LVi. B, F. RVi, C, D, G, H. CMS. Scale bars. 1 (A, B, E, F) = 500 µm; 2 (C, D, G, H) = 50 µm." pageId="10" pageNumber="11">Figure 4C</figureCitation>
) consisting of six scars: one large dorsal, three anterior and two posterior ones; one short antero-ventral il.
</paragraph>
<paragraph id="7209B62C453FFFB6FF111C02FCDE2AC3" blockId="10.[151,1437,1400,2039]" pageId="10" pageNumber="11">
RVi (
<figureCitation id="EA8DAAA9453FFFB6FF291C02FE182A52" box="[255,406,1868,1895]" captionStart="FIGURE 3" captionStartId="11.[151,250,1933,1957]" captionTargetBox="[172,1415,181,1910]" captionTargetId="figure-17@11.[172,1415,178,1910]" captionTargetPageId="11" captionText="FIGURE 3. Benincandona martini gen. et spec. nov., valves (A, C, EH, J ♂; B, D, I ♀). AD, I. CpRl, (A, C. INV.159208; B, D, I. INV.159211). E. INV.159209. CpD. F. INV.159210. CpV. G. INV.159200. LVi. H. INV.159200. RVi. J. INV.159200. CMS. Scale bars. 1 (A, B, E, F, G, H) = 500 µm; 2 (C, D) = 200 µm; 3 (I, J) = 50 µm." pageId="10" pageNumber="11">Figures 3H, J</figureCitation>
,
<figureCitation id="EA8DAAA9453FFFB6FE771C03FE6D2A52" box="[417,483,1869,1895]" captionStart="FIGURE 4" captionStartId="12.[151,250,1944,1968]" captionTargetBox="[157,1430,181,1920]" captionTargetId="figure-31@12.[157,1430,181,1920]" captionTargetPageId="12" captionText="FIGURE 4. Benincandona martini gen. et spec. nov., valves (AD: INV.159202 ♂ and EH. INV.159201 ♀) A, E. LVi. B, F. RVi, C, D, G, H. CMS. Scale bars. 1 (A, B, E, F) = 500 µm; 2 (C, D, G, H) = 50 µm." pageId="10" pageNumber="11">4B, D</figureCitation>
) elongated, with anterior margin slightly more broadly rounded than posterior margin, dorsal margin greatest with height situated about 1/3 from the rear, anterior part sloping almost straight to anterior margin but with a blunt angle in the middle, posterior part sloping almost straight to posterior margin, ventral margin almost straight. CMS (
<figureCitation id="EA8DAAA9453FFFB6FE441CF6FD862AE6" box="[402,520,1976,2003]" captionStart="FIGURE 3" captionStartId="11.[151,250,1933,1957]" captionTargetBox="[172,1415,181,1910]" captionTargetId="figure-17@11.[172,1415,178,1910]" captionTargetPageId="11" captionText="FIGURE 3. Benincandona martini gen. et spec. nov., valves (A, C, EH, J ♂; B, D, I ♀). AD, I. CpRl, (A, C. INV.159208; B, D, I. INV.159211). E. INV.159209. CpD. F. INV.159210. CpV. G. INV.159200. LVi. H. INV.159200. RVi. J. INV.159200. CMS. Scale bars. 1 (A, B, E, F, G, H) = 500 µm; 2 (C, D) = 200 µm; 3 (I, J) = 50 µm." pageId="10" pageNumber="11">Figures 3J</figureCitation>
,
<figureCitation id="EA8DAAA9453FFFB6FDC51CF7FDB92AE6" box="[531,567,1977,2003]" captionStart="FIGURE 4" captionStartId="12.[151,250,1944,1968]" captionTargetBox="[157,1430,181,1920]" captionTargetId="figure-31@12.[157,1430,181,1920]" captionTargetPageId="12" captionText="FIGURE 4. Benincandona martini gen. et spec. nov., valves (AD: INV.159202 ♂ and EH. INV.159201 ♀) A, E. LVi. B, F. RVi, C, D, G, H. CMS. Scale bars. 1 (A, B, E, F) = 500 µm; 2 (C, D, G, H) = 50 µm." pageId="10" pageNumber="11">4D</figureCitation>
) consisting of six scars: one large dorsal (1), three anterior (24) and two (5,6) posterior ones; il running submarginally along ventral margin.
</paragraph>
<caption id="26C9E6A4453EFFB7FF411CC3FC722ADB" pageId="11" pageNumber="12" startId="11.[151,250,1933,1957]" targetBox="[172,1415,181,1910]" targetPageId="11" targetType="figure">
<paragraph id="7209B62C453EFFB7FF411CC3FC722ADB" blockId="11.[151,1437,1933,2030]" pageId="11" pageNumber="12">
<emphasis id="40C26A3E453EFFB7FF411CC3FE802A90" bold="true" box="[151,270,1933,1957]" pageId="11" pageNumber="12">FIGURE 3</emphasis>
.
<taxonomicName id="B5B6CDAF453EFFB7FECD1CC3FD8E2A90" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[283,512,1933,1957]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="martini" status="gen. et spec. nov.">
<emphasis id="40C26A3E453EFFB7FECD1CC3FD8E2A90" box="[283,512,1933,1957]" italics="true" pageId="11" pageNumber="12">Benincandona martini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745453EFFB7FDD11CC0FD352A90" box="[519,699,1934,1958]" pageId="11" pageNumber="12" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, valves (A, C, EH, J ♂; B, D, I ♀).
<emphasis id="40C26A3E453EFFB7FBEF1CC3FB0C2A93" bold="true" box="[1081,1154,1933,1958]" pageId="11" pageNumber="12">AD, I</emphasis>
. CpRl, (A, C. INV.159208; B, D, I. INV.159211).
<emphasis id="40C26A3E453EFFB7FEAB1CFCFE002AFF" bold="true" box="[381,398,1970,1994]" pageId="11" pageNumber="12">E</emphasis>
. INV.159209. CpD.
<emphasis id="40C26A3E453EFFB7FDB41CFCFDFF2AFF" bold="true" box="[610,625,1970,1994]" pageId="11" pageNumber="12">F</emphasis>
. INV.159210. CpV.
<emphasis id="40C26A3E453EFFB7FC951CFFFCD92AFC" bold="true" box="[835,855,1969,1993]" pageId="11" pageNumber="12">G</emphasis>
. INV.159200. LVi.
<emphasis id="40C26A3E453EFFB7FBF61CFCFBBA2AFF" bold="true" box="[1056,1076,1970,1994]" pageId="11" pageNumber="12">H</emphasis>
. INV.159200. RVi.
<emphasis id="40C26A3E453EFFB7FAD61CFCFA832AFF" bold="true" box="[1280,1293,1970,1994]" pageId="11" pageNumber="12">J</emphasis>
. INV.159200. CMS. Scale bars. 1 (A, B, E, F, G, H) = 500 µm; 2 (C, D) = 200 µm; 3 (I, J) = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A44539FFB0FF411CD6FC792AE0" pageId="12" pageNumber="13" startId="12.[151,250,1944,1968]" targetBox="[157,1430,181,1920]" targetPageId="12" targetType="figure">
<paragraph id="7209B62C4539FFB0FF411CD6FC792AE0" blockId="12.[151,1436,1944,2005]" pageId="12" pageNumber="13">
<emphasis id="40C26A3E4539FFB0FF411CD6FE9D2A85" bold="true" box="[151,275,1944,1968]" pageId="12" pageNumber="13">FIGURE 4.</emphasis>
<taxonomicName id="B5B6CDAF4539FFB0FECC1CD6FE702A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[282,510,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="martini" status="gen. et spec. nov.">
<emphasis id="40C26A3E4539FFB0FECC1CD6FE702A85" box="[282,510,1944,1968]" italics="true" pageId="12" pageNumber="13">Benincandona martini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454539FFB0FDD31CD7FD392A85" box="[517,695,1945,1969]" pageId="12" pageNumber="13" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, valves (AD: INV.159202 ♂ and EH. INV.159201 ♀)
<emphasis id="40C26A3E4539FFB0FB2F1CD6FAA72A85" bold="true" box="[1273,1321,1944,1968]" pageId="12" pageNumber="13">A, E</emphasis>
. LVi.
<emphasis id="40C26A3E4539FFB0FABE1CD6FA1B2A85" bold="true" box="[1384,1429,1944,1968]" pageId="12" pageNumber="13">B, F</emphasis>
. RVi,
<emphasis id="40C26A3E4539FFB0FF1D1CF2FEB32AE1" bold="true" box="[203,317,1980,2004]" pageId="12" pageNumber="13">C, D, G, H</emphasis>
. CMS. Scale bars. 1 (A, B, E, F) = 500 µm; 2 (C, D, G, H) = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A44538FFB1FF411CD9FD0F2AE1" pageId="13" pageNumber="14" startId="13.[151,250,1943,1967]" targetBox="[170,1415,185,1915]" targetPageId="13" targetType="figure">
<paragraph id="7209B62C4538FFB1FF411CD9FD0F2AE1" blockId="13.[151,1436,1943,2004]" pageId="13" pageNumber="14">
<emphasis id="40C26A3E4538FFB1FF411CD9FE9A2A85" bold="true" box="[151,276,1943,1968]" pageId="13" pageNumber="14">FIGURE 5.</emphasis>
<taxonomicName id="B5B6CDAF4538FFB1FECA1CD6FD8F2A9A" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[284,513,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="martini" status="gen. et spec. nov.">
<emphasis id="40C26A3E4538FFB1FECA1CD6FD8F2A9A" box="[284,513,1944,1968]" italics="true" pageId="13" pageNumber="14">Benincandona martini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454538FFB1FDDF1CD6FD312A9A" box="[521,703,1944,1968]" pageId="13" pageNumber="14" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E4538FFB1FCEA1CD9FCC02A9A" bold="true" box="[828,846,1943,1967]" pageId="13" pageNumber="14">A</emphasis>
. A1 (INV.159203).
<emphasis id="40C26A3E4538FFB1FBCB1CD6FBC02A9A" bold="true" box="[1053,1102,1943,1968]" pageId="13" pageNumber="14">B, C</emphasis>
. MdCoxa, (INV.159202).
<emphasis id="40C26A3E4538FFB1FA881CD6FAFE2A85" bold="true" box="[1374,1392,1944,1968]" pageId="13" pageNumber="14">D</emphasis>
. A2 (INV.159200 INV.159203). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A4453BFFB2FF411CF3FA8E2ACF" pageId="14" pageNumber="15" startId="14.[151,250,1981,2005]" targetBox="[169,1422,182,1942]" targetPageId="14" targetType="figure">
<paragraph id="7209B62C453BFFB2FF411CF3FA8E2ACF" blockId="14.[151,1436,1981,2042]" pageId="14" pageNumber="15">
<emphasis id="40C26A3E453BFFB2FF411CF3FE982AE0" bold="true" box="[151,278,1981,2005]" pageId="14" pageNumber="15">FIGURE 6.</emphasis>
<taxonomicName id="B5B6CDAF453BFFB2FEC91CF3FD8B2AE0" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[287,517,1981,2005]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="martini" status="gen. et spec. nov.">
<emphasis id="40C26A3E453BFFB2FEC91CF3FD8B2AE0" box="[287,517,1981,2005]" italics="true" pageId="14" pageNumber="15">Benincandona martini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745453BFFB2FDD81CF0FD492AE0" box="[526,711,1982,2006]" pageId="14" pageNumber="15" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E453BFFB2FC9E1CF3FCD42AE0" bold="true" box="[840,858,1981,2005]" pageId="14" pageNumber="15">A</emphasis>
. Mx1 (Chaetotaxy incomplete) (INV.159200).
<emphasis id="40C26A3E453BFFB2FA901CF0FAD92AE3" bold="true" box="[1350,1367,1982,2006]" pageId="14" pageNumber="15">B</emphasis>
. Mx1, respiratory plate (INV.159202).
<emphasis id="40C26A3E453BFFB2FE0A1CAFFE602ACC" bold="true" box="[476,494,2017,2041]" pageId="14" pageNumber="15">C</emphasis>
. MdPalp (INV.159204 INV.159203). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm.
</paragraph>
</caption>
<paragraph id="7209B62C453AFFB3FF111BD9FCD62F2C" blockId="15.[151,1437,150,1869]" pageId="15" pageNumber="16">
A1 (
<figureCitation id="EA8DAAA9453AFFB3FF2E1BD9FEE32D84" box="[248,365,150,177]" captionStart="FIGURE 5" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[170,1415,185,1915]" captionTargetId="figure-17@13.[167,1419,181,1919]" captionTargetPageId="13" captionText="FIGURE 5. Benincandona martini gen. et spec. nov., limbs (♂). A. A1 (INV.159203). B, C. MdCoxa, (INV.159202). D. A2 (INV.159200 INV.159203). Scale bar = 50 µm." pageId="15" pageNumber="16">Figure 5A</figureCitation>
) consisting of seven articulated segments, one basal and six distal. Basal segment
<emphasis id="40C26A3E453AFFB3FAD01BD9FA942D85" box="[1286,1306,151,176]" italics="true" pageId="15" pageNumber="16">c.</emphasis>
three times as long as basal width, with two sub-apical, ventral setae, one long and one shorter, the latter about half of the length of the longer seta; and two shorter setae on the dorsal side, situated some distance from each other (one sub-apical and one more proximal). First distal segment slightly longer than basal width, with one short dorso-apical seta. Second distal segment about twice as long as wide, set with one short dorso-apical seta. Third distal segment with about the same length/width ratio as the second segment, carrying one short apical seta on each side. Fourth distal segment about three times as long as wide, with one ventro-apical seta, and one long dorso-apical seta, reaching beyond tip of sixth segment with almost one third of total length. Fifth distal segment four times as long as wide, set with two long dorso-apical setae and one short ventro-apical seta. Sixth distal segment narrow,
<emphasis id="40C26A3E453AFFB3FB261AF9FA8D2CE5" box="[1264,1283,439,464]" italics="true" pageId="15" pageNumber="16">c.</emphasis>
four times as long as wide, with four apical structures: one aesthetasc Ya of intermediate length, two long setae more than twice the length of Ya and one short seta, about half the length of Ya.
</paragraph>
<paragraph id="7209B62C453AFFB3FF11196DFCE5296C" blockId="15.[151,1437,150,1869]" pageId="15" pageNumber="16">
A2 (
<figureCitation id="EA8DAAA9453AFFB3FF20196DFEE72F08" box="[246,361,546,573]" captionStart="FIGURE 5" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[170,1415,185,1915]" captionTargetId="figure-17@13.[167,1419,181,1919]" captionTargetPageId="13" captionText="FIGURE 5. Benincandona martini gen. et spec. nov., limbs (♂). A. A1 (INV.159203). B, C. MdCoxa, (INV.159202). D. A2 (INV.159200 INV.159203). Scale bar = 50 µm." pageId="15" pageNumber="16">Figure 5D</figureCitation>
) with basal segment (first protopodite) broad and rounded, mid-ventrally with one seta of medium length, subapically with two setae of unequal length, proximal one
<emphasis id="40C26A3E453AFFB3FCAC1909FC032F55" box="[890,909,583,608]" italics="true" pageId="15" pageNumber="16">c.</emphasis>
twice the length of the more apical one. Second segment (second protopodite) large and
<emphasis id="40C26A3E453AFFB3FD8A1925FDE12FB1" box="[604,623,619,644]" italics="true" pageId="15" pageNumber="16">c.</emphasis>
twice as long as basal width, set with one long ventro-subapical seta X, reaching tip of next segment (first endopodal segment). No remnant of exopodite visible. Endopodite consisting of four segments (penultimate segment divided). First endopodal segment large and stout,
<emphasis id="40C26A3E453AFFB3FBB319FDFBFF2FF9" box="[1125,1137,691,716]" italics="true" pageId="15" pageNumber="16">c</emphasis>
four times as long as basal width, carrying one long and narrow mid-ventral aesthetasc Y (reaching tip of segment) and two ventro-subapical setae, one short and one large and stout, hirsute in its distal 4/5. Second endopodal segment (segment 2A)
<emphasis id="40C26A3E453AFFB3FAFF19B5FAB22E21" box="[1321,1340,763,788]" italics="true" pageId="15" pageNumber="16">c.</emphasis>
twice as long as wide, carrying one stout dorso-apical seta and three t-setae; seta t1 long, narrow and plumose, accompanied by a long and slender, but well-developed aesthetasc y1; setae t2 and t3 hyper-developed into male bristles, t2 ventro-apical, t3 centro-apical. Third endopodal segment (segment 2B) smaller than previous segment and about as long as wide, set with a complex (and sexually dimorphic) apical chaetotaxy (
<figureCitation id="EA8DAAA9453AFFB3FBD518C5FBF82E90" box="[1027,1142,907,933]" captionStart="FIGURE 40" captionStartId="60.[151,250,1516,1540]" captionTargetBox="[151,1435,181,1492]" captionTargetId="figure-31@60.[151,1435,181,1492]" captionTargetPageId="60" captionText="FIGURE 40. Diagrams of apical views of the A2 of Benincandona, showing position of (sub-) terminal claws, setae and aesthetascs of the terminal (small circle) and penultimate segment (large circle). For explanations of codes, see Material and Methods (Abbreviations used in text and figures)." pageId="15" pageNumber="16">Figure 40</figureCitation>
): ventro-apically with one short and slender aesthetasc y2 and the claw G3 transformed into short seta of equal length; centrally with one short seta z3; dorso-apically with the seta z1 transformed into claw, length about twice as long as penultimate segment. Claw G1 slightly shorter than G2; seta z2 transformed into long claw, slightly shorter than G1. Fourth, terminal segment with one long claw (Gm), one short claw (GM), a long aesthetasc y3 with shorter accompanying seta, with fused part short, and a long seta g slightly shorter than claw Gm.
</paragraph>
<paragraph id="7209B62C453AFFB3FF111F2DFBDC29F0" blockId="15.[151,1437,150,1869]" pageId="15" pageNumber="16">
MdCoxa (
<figureCitation id="EA8DAAA9453AFFB3FEEF1F2DFE5F2948" box="[313,465,1122,1149]" captionStart="FIGURE 5" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[170,1415,185,1915]" captionTargetId="figure-17@13.[167,1419,181,1919]" captionTargetPageId="13" captionText="FIGURE 5. Benincandona martini gen. et spec. nov., limbs (♂). A. A1 (INV.159203). B, C. MdCoxa, (INV.159202). D. A2 (INV.159200 INV.159203). Scale bar = 50 µm." pageId="15" pageNumber="16">Figure 5B, C</figureCitation>
) elongated, proximally sharply pointed, medially widened, distally with an obliquely widened end, set with a series (67) of strong teeth, interspaced with setae; subapically with a short seta.
<emphasis id="40C26A3E453AFFB3FAEB1FC9FA1A2994" box="[1341,1428,1159,1185]" italics="true" pageId="15" pageNumber="16">Remark</emphasis>
: asymmetry illustrated in
<figureCitation id="EA8DAAA9453AFFB3FE781FE5FDAE29F0" box="[430,544,1194,1221]" captionStart="FIGURE 5" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[170,1415,185,1915]" captionTargetId="figure-17@13.[167,1419,181,1919]" captionTargetPageId="13" captionText="FIGURE 5. Benincandona martini gen. et spec. nov., limbs (♂). A. A1 (INV.159203). B, C. MdCoxa, (INV.159202). D. A2 (INV.159200 INV.159203). Scale bar = 50 µm." pageId="15" pageNumber="16">Figure 5C</figureCitation>
possibly resulting from different position in slide.
</paragraph>
<paragraph id="7209B62C453AFFB3FF111F81FE092B64" blockId="15.[151,1437,150,1869]" pageId="15" pageNumber="16">
MdPalp (
<figureCitation id="EA8DAAA9453AFFB3FEF81F81FE2F29DC" box="[302,417,1231,1257]" captionStart="FIGURE 6" captionStartId="14.[151,250,1981,2005]" captionTargetBox="[169,1422,182,1942]" captionTargetId="figure-31@14.[166,1422,181,1957]" captionTargetPageId="14" captionText="FIGURE 6. Benincandona martini gen. et spec. nov., limbs (♂). A. Mx1 (Chaetotaxy incomplete) (INV.159200). B. Mx1, respiratory plate (INV.159202). C. MdPalp (INV.159204 INV.159203). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." pageId="15" pageNumber="16">Figure 6C</figureCitation>
) set at about one third of the tip of the coxa and consisting of four segments. Basal segment dorsally carrying the respiratory plate, set with six distal rays (four long, two short), subapically with one long and two short rays, proximally with one long ray (this chaetotaxy approximate); ventro-subapically with one long hirsute seta (S1) and more apically S2 hirsute seta, shorter and stout, length about one third of the length of S1; in between with the short flagellum-like alpha seta, and one smooth and short, more proximal seta. Second segment of sub-triangular shape, dorso-apically with two setae of unequal length, ventro-apically with a group of four long setae, one of these being the beta-seta. Third segment with two unequal dorso-subapical setae, two mid-dorsal setae, one of which being the gamma seta and ventro-apically with two setae of sub-equal length. Terminal segment sub-rectangular, apically set with one large claw, fused with the segment over its entire distal margin, accompanied by one smooth lateral seta on each side, length of dorsal one (slightly shorter than the fused claw) about 1.5 times length of ventral one.
</paragraph>
<paragraph id="7209B62C453AFFB3FF111D15FCB12A78" blockId="15.[151,1437,150,1869]" pageId="15" pageNumber="16">
Mx1 (
<figureCitation id="EA8DAAA9453AFFB3FEDD1D15FE2E2B40" box="[267,416,1627,1653]" captionStart="FIGURE 6" captionStartId="14.[151,250,1981,2005]" captionTargetBox="[169,1422,182,1942]" captionTargetId="figure-31@14.[166,1422,181,1957]" captionTargetPageId="14" captionText="FIGURE 6. Benincandona martini gen. et spec. nov., limbs (♂). A. Mx1 (Chaetotaxy incomplete) (INV.159200). B. Mx1, respiratory plate (INV.159202). C. MdPalp (INV.159204 INV.159203). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." pageId="15" pageNumber="16">Figure 6A, B</figureCitation>
) with a basal (protopodite) part carrying a large respiratory plate (exopodite), three endites and a two-segmented palp (endopodite). Palp (
<figureCitation id="EA8DAAA9453AFFB3FD771D31FC992BAC" box="[673,791,1663,1689]" captionStart="FIGURE 6" captionStartId="14.[151,250,1981,2005]" captionTargetBox="[169,1422,182,1942]" captionTargetId="figure-31@14.[166,1422,181,1957]" captionTargetPageId="14" captionText="FIGURE 6. Benincandona martini gen. et spec. nov., limbs (♂). A. Mx1 (Chaetotaxy incomplete) (INV.159200). B. Mx1, respiratory plate (INV.159202). C. MdPalp (INV.159204 INV.159203). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." pageId="15" pageNumber="16">Figure 6A</figureCitation>
) with first segment carrying four long dorso-apical setae; terminal segment short and rectangular, carrying four distal setae, some slightly claw-like; chaetotaxy of three endites incompletely illustrated: third endite with two strong teeth carrying lateral spines (
<emphasis id="40C26A3E453AFFB3FB701D89FAB22BD4" box="[1190,1340,1735,1761]" italics="true" pageId="15" pageNumber="16">Zahnborsten)</emphasis>
, basally with one lateral seta; sideways directed bristles near base of first endite one long and one short
<emphasis id="40C26A3E453AFFB3FB141DA5FB5B2A31" box="[1218,1237,1771,1796]" italics="true" pageId="15" pageNumber="16">c.</emphasis>
half the length of the long one. Respiratory plate (
<figureCitation id="EA8DAAA9453AFFB3FE2F1C41FDE22A1C" box="[505,620,1807,1833]" captionStart="FIGURE 6" captionStartId="14.[151,250,1981,2005]" captionTargetBox="[169,1422,182,1942]" captionTargetId="figure-31@14.[166,1422,181,1957]" captionTargetPageId="14" captionText="FIGURE 6. Benincandona martini gen. et spec. nov., limbs (♂). A. Mx1 (Chaetotaxy incomplete) (INV.159200). B. Mx1, respiratory plate (INV.159202). C. MdPalp (INV.159204 INV.159203). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." pageId="15" pageNumber="16">Figure 6B</figureCitation>
) elongated, carrying
<emphasis id="40C26A3E453AFFB3FC8E1C41FCE52A1D" box="[856,875,1807,1832]" italics="true" pageId="15" pageNumber="16">c.</emphasis>
15 plumose respiratory rays, two short distal setae and a group of five shorter proximal setae of unequal length.
</paragraph>
<caption id="26C9E6A44525FFACFF411CF3FD432ACF" pageId="16" pageNumber="17" startId="16.[151,250,1981,2005]" targetBox="[186,1404,198,1941]" targetPageId="16" targetType="figure">
<paragraph id="7209B62C4525FFACFF411CF3FD432ACF" blockId="16.[151,1436,1981,2042]" pageId="16" pageNumber="17">
<emphasis id="40C26A3E4525FFACFF411CF3FE962AE3" bold="true" box="[151,280,1981,2006]" pageId="16" pageNumber="17">FIGURE 7.</emphasis>
<taxonomicName id="B5B6CDAF4525FFACFEF21CF3FD832AE0" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[292,525,1981,2005]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="martini" status="gen. et spec. nov.">
<emphasis id="40C26A3E4525FFACFEF21CF3FD832AE0" box="[292,525,1981,2005]" italics="true" pageId="16" pageNumber="17">Benincandona martini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454525FFACFDCE1CF0FD542AE0" box="[536,730,1982,2006]" pageId="16" pageNumber="17" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E4525FFACFCB51CF3FCFB2AE0" bold="true" box="[867,885,1981,2005]" pageId="16" pageNumber="17">A</emphasis>
. T2 (INV.159200).
<emphasis id="40C26A3E4525FFACFB9B1CF0FBD02AE3" bold="true" box="[1101,1118,1982,2006]" pageId="16" pageNumber="17">B</emphasis>
. CR (INV.159200).
<emphasis id="40C26A3E4525FFACFAEB1CF3FAC12AE0" bold="true" box="[1341,1359,1981,2005]" pageId="16" pageNumber="17">C</emphasis>
. CRatt (INV.159200).
<emphasis id="40C26A3E4525FFACFEE71CACFECD2ACF" bold="true" box="[305,323,2018,2042]" pageId="16" pageNumber="17">D</emphasis>
. T3 (INV.159200). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A44524FFADFF411CD9FDB32ACD" pageId="17" pageNumber="18" startId="17.[151,250,1943,1967]" targetBox="[196,1391,191,1914]" targetPageId="17" targetType="figure">
<paragraph id="7209B62C4524FFADFF411CD9FDB32ACD" blockId="17.[151,1437,1943,2040]" pageId="17" pageNumber="18">
<emphasis id="40C26A3E4524FFADFF411CD9FE982A9A" bold="true" box="[151,278,1943,1967]" pageId="17" pageNumber="18">FIGURE 8.</emphasis>
<taxonomicName id="B5B6CDAF4524FFADFEF61CD6FD892A9A" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[288,519,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="martini" status="gen. et spec. nov.">
<emphasis id="40C26A3E4524FFADFEF61CD6FD892A9A" box="[288,519,1944,1968]" italics="true" pageId="17" pageNumber="18">Benincandona martini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454524FFADFDC61CD6FD452A9A" box="[528,715,1944,1968]" pageId="17" pageNumber="18" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E4524FFADFC9B1CD9FCD12A9A" bold="true" box="[845,863,1943,1967]" pageId="17" pageNumber="18">A</emphasis>
. Zenker organ (INV.159200).
<emphasis id="40C26A3E4524FFADFB771CD6FB5E2A85" bold="true" box="[1185,1232,1944,1968]" pageId="17" pageNumber="18">BD</emphasis>
. Hp (INV.159200).
<emphasis id="40C26A3E4524FFADFF411CF2FF262AE1" bold="true" box="[151,168,1980,2004]" pageId="17" pageNumber="18">B</emphasis>
. External view (drawing).
<emphasis id="40C26A3E4524FFADFE141CF5FE5A2AE6" bold="true" box="[450,468,1979,2003]" pageId="17" pageNumber="18">C</emphasis>
. Internal view (photograph with interference contrast).
<emphasis id="40C26A3E4524FFADFBCE1CF2FBA42AE1" bold="true" box="[1048,1066,1980,2004]" pageId="17" pageNumber="18">D</emphasis>
. Internal view (photograph without interference contrast). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A44527FFAEFF411CD9FA9F2AE1" pageId="18" pageNumber="19" startId="18.[151,250,1943,1967]" targetBox="[172,1415,185,1915]" targetPageId="18" targetType="figure">
<paragraph id="7209B62C4527FFAEFF411CD9FA9F2AE1" blockId="18.[151,1436,1943,2004]" pageId="18" pageNumber="19">
<emphasis id="40C26A3E4527FFAEFF411CD9FE982A9A" bold="true" box="[151,278,1943,1967]" pageId="18" pageNumber="19">FIGURE 9.</emphasis>
<taxonomicName id="B5B6CDAF4527FFAEFEF61CD6FD892A9A" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[288,519,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="martini" status="gen. et spec. nov.">
<emphasis id="40C26A3E4527FFAEFEF61CD6FD892A9A" box="[288,519,1944,1968]" italics="true" pageId="18" pageNumber="19">Benincandona martini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454527FFAEFDC61CD6FD452A9A" box="[528,715,1944,1968]" pageId="18" pageNumber="19" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (AB ♂ and CE ♀).
<emphasis id="40C26A3E4527FFAEFBDE1CD9FB942A9A" bold="true" box="[1032,1050,1943,1967]" pageId="18" pageNumber="19">A</emphasis>
. T1, Lpp (INV.159202).
<emphasis id="40C26A3E4527FFAEFAF01CD6FAB92A85" bold="true" box="[1318,1335,1944,1968]" pageId="18" pageNumber="19">B</emphasis>
. T1, Rpp (INV.159202).
<emphasis id="40C26A3E4527FFAEFEE71CF5FECD2AE6" bold="true" box="[305,323,1979,2003]" pageId="18" pageNumber="19">C</emphasis>
. A2 (INV.159205, INV.159207).
<emphasis id="40C26A3E4527FFAEFD401CF2FD262AE1" bold="true" box="[662,680,1980,2004]" pageId="18" pageNumber="19">D</emphasis>
. CR (INV.159205).
<emphasis id="40C26A3E4527FFAEFCA11CF2FC062AE1" bold="true" box="[887,904,1980,2004]" pageId="18" pageNumber="19">E</emphasis>
. T1 (INV.159200). Scale bar = 50 µm.
</paragraph>
</caption>
<paragraph id="7209B62C4526FFAFFF111BD9FD062C50" blockId="19.[151,1437,151,1005]" pageId="19" pageNumber="20">
T1 (
<figureCitation id="EA8DAAA94526FFAFFF201BD9FE012D84" box="[246,399,151,177]" captionStart="FIGURE 9" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[172,1415,185,1915]" captionTargetId="figure-31@18.[168,1419,181,1919]" captionTargetPageId="18" captionText="FIGURE 9. Benincandona martini gen. et spec. nov., limbs (AB ♂ and CE ♀). A. T1, Lpp (INV.159202). B. T1, Rpp (INV.159202). C. A2 (INV.159205, INV.159207). D. CR (INV.159205). E. T1 (INV.159200). Scale bar = 50 µm." pageId="19" pageNumber="20">Figure 9A, B</figureCitation>
) consisting of basal part (protopodite), carrying respiratory plates (exopodites) represented here by one long and one short plumose setae, and slightly asymmetrical prehensile palps (endopodites). Protopodite (
<figureCitation id="EA8DAAA94526FFAFFF491B91FE9E2DCC" box="[159,272,223,249]" captionStart="FIGURE 9" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[172,1415,185,1915]" captionTargetId="figure-31@18.[168,1419,181,1919]" captionTargetPageId="18" captionText="FIGURE 9. Benincandona martini gen. et spec. nov., limbs (AB ♂ and CE ♀). A. T1, Lpp (INV.159202). B. T1, Rpp (INV.159202). C. A2 (INV.159205, INV.159207). D. CR (INV.159205). E. T1 (INV.159200). Scale bar = 50 µm." pageId="19" pageNumber="20">Figure 9B</figureCitation>
) distally set with 5+2 setae of different morphology and length (not illustrated); more proximally carrying d, d and b setae of medium length. Prehensile palps (
<figureCitation id="EA8DAAA94526FFAFFD341A4DFCFB2C28" box="[738,885,259,285]" captionStart="FIGURE 9" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[172,1415,185,1915]" captionTargetId="figure-31@18.[168,1419,181,1919]" captionTargetPageId="18" captionText="FIGURE 9. Benincandona martini gen. et spec. nov., limbs (AB ♂ and CE ♀). A. T1, Lpp (INV.159202). B. T1, Rpp (INV.159202). C. A2 (INV.159205, INV.159207). D. CR (INV.159205). E. T1 (INV.159200). Scale bar = 50 µm." pageId="19" pageNumber="20">Figure 9A, B</figureCitation>
) one-segmented, distally sigmoid hook-like, both palps apically set with a short sensory organ, proximally with two subequal setae (also sensory organs); Lpp (
<figureCitation id="EA8DAAA94526FFAFFA871A69FF352C50" captionStart="FIGURE 9" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[172,1415,185,1915]" captionTargetId="figure-31@18.[168,1419,181,1919]" captionTargetPageId="18" captionText="FIGURE 9. Benincandona martini gen. et spec. nov., limbs (AB ♂ and CE ♀). A. T1, Lpp (INV.159202). B. T1, Rpp (INV.159202). C. A2 (INV.159205, INV.159207). D. CR (INV.159205). E. T1 (INV.159200). Scale bar = 50 µm." pageId="19" pageNumber="20">Figure 9A</figureCitation>
) slightly narrower than Rpp (
<figureCitation id="EA8DAAA94526FFAFFDD31A05FDF52C50" box="[517,635,331,357]" captionStart="FIGURE 9" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[172,1415,185,1915]" captionTargetId="figure-31@18.[168,1419,181,1919]" captionTargetPageId="18" captionText="FIGURE 9. Benincandona martini gen. et spec. nov., limbs (AB ♂ and CE ♀). A. T1, Lpp (INV.159202). B. T1, Rpp (INV.159202). C. A2 (INV.159205, INV.159207). D. CR (INV.159205). E. T1 (INV.159200). Scale bar = 50 µm." pageId="19" pageNumber="20">Figure 9B</figureCitation>
).
</paragraph>
<paragraph id="7209B62C4526FFAFFF111A21FCD12F08" blockId="19.[151,1437,151,1005]" pageId="19" pageNumber="20">
T2 (
<figureCitation id="EA8DAAA94526FFAFFF201A21FEE32CBC" box="[246,365,367,393]" captionStart="FIGURE 7" captionStartId="16.[151,250,1981,2005]" captionTargetBox="[186,1404,198,1941]" captionTargetId="figure-31@16.[166,1422,181,1957]" captionTargetPageId="16" captionText="FIGURE 7. Benincandona martini gen. et spec. nov., limbs (♂). A. T2 (INV.159200). B. CR (INV.159200). C. CRatt (INV.159200). D. T3 (INV.159200). Scale bar = 50 µm." pageId="19" pageNumber="20">Figure 7A</figureCitation>
) with 4-segmented endopodite (penultimate segment divided), generally elongated. First basal segment and knee-segment without setae d1 and d2. First segment of endopodite elongated (length at least 4x the basal width), without setae. Second endopodal segment with length
<emphasis id="40C26A3E4526FFAFFC451AF9FC292CE5" box="[915,935,439,464]" italics="true" pageId="19" pageNumber="20">c.</emphasis>
3x basal width, with one short ventro-apical seta f. Third endopodal segment of about the same length/width ratio as the previous segment and with one ventro-subapical seta g. Terminal segment about 1.5 times as long as basal width, apically with two short setae (h1 and h3) and a long and slender claw (h2), distally set with short setulae.
</paragraph>
<paragraph id="7209B62C4526FFAFFF111909FBDB2FC4" blockId="19.[151,1437,151,1005]" pageId="19" pageNumber="20">
T3 (
<figureCitation id="EA8DAAA94526FFAFFF211909FEE12F54" box="[247,367,583,609]" captionStart="FIGURE 7" captionStartId="16.[151,250,1981,2005]" captionTargetBox="[186,1404,198,1941]" captionTargetId="figure-31@16.[166,1422,181,1957]" captionTargetPageId="16" captionText="FIGURE 7. Benincandona martini gen. et spec. nov., limbs (♂). A. T2 (INV.159200). B. CR (INV.159200). C. CRatt (INV.159200). D. T3 (INV.159200). Scale bar = 50 µm." pageId="19" pageNumber="20">Figure 7D</figureCitation>
) with first segment carrying only one long subapical seta d1. Second (knee-) segment with a short seta d2. Third segment about 5x as long as wide, not carrying any setae. Fourth segment narrower, but still
<emphasis id="40C26A3E4526FFAFFA5F1925FA122FB1" box="[1417,1436,619,644]" italics="true" pageId="19" pageNumber="20">c.</emphasis>
5x basal width and carrying one short subapical seta g. Terminal segment well-separated from penultimate segment, L
<emphasis id="40C26A3E4526FFAFFF7919FDFF4C2FF9" box="[175,194,691,716]" italics="true" pageId="19" pageNumber="20">c.</emphasis>
1.5x basal width and carrying three setae: one long and reflexed (h3), almost reaching knee-segment and two ventrally directed and curved setae h1 &amp; h2, the former
<emphasis id="40C26A3E4526FFAFFCDD1999FC902FC5" box="[779,798,727,752]" italics="true" pageId="19" pageNumber="20">c.</emphasis>
half the length of the latter.
</paragraph>
<paragraph id="7209B62C4526FFAFFF1119B5FC852E0C" blockId="19.[151,1437,151,1005]" pageId="19" pageNumber="20">
CR (
<figureCitation id="EA8DAAA94526FFAFFF2819B5FEFD2E20" box="[254,371,763,789]" captionStart="FIGURE 7" captionStartId="16.[151,250,1981,2005]" captionTargetBox="[186,1404,198,1941]" captionTargetId="figure-31@16.[166,1422,181,1957]" captionTargetPageId="16" captionText="FIGURE 7. Benincandona martini gen. et spec. nov., limbs (♂). A. T2 (INV.159200). B. CR (INV.159200). C. CRatt (INV.159200). D. T3 (INV.159200). Scale bar = 50 µm." pageId="19" pageNumber="20">Figure 7B</figureCitation>
) with ramus stout; claw Ga well-developed, stout and reaching widely beyond half L of the ramus, setae Sp, Sa and claw Gp reduced to small setae.
</paragraph>
<paragraph id="7209B62C4526FFAFFF11180DFCA92E68" blockId="19.[151,1437,151,1005]" box="[199,807,835,861]" pageId="19" pageNumber="20">
CRAtt (
<figureCitation id="EA8DAAA94526FFAFFEC9180DFE1D2E68" box="[287,403,835,861]" captionStart="FIGURE 7" captionStartId="16.[151,250,1981,2005]" captionTargetBox="[186,1404,198,1941]" captionTargetId="figure-31@16.[166,1422,181,1957]" captionTargetPageId="16" captionText="FIGURE 7. Benincandona martini gen. et spec. nov., limbs (♂). A. T2 (INV.159200). B. CR (INV.159200). C. CRatt (INV.159200). D. T3 (INV.159200). Scale bar = 50 µm." pageId="19" pageNumber="20">Figure 7C</figureCitation>
) short and stout, distally bifurcated.
</paragraph>
<paragraph id="7209B62C4526FFAFFF111829FD4E2EFC" blockId="19.[151,1437,151,1005]" pageId="19" pageNumber="20">
Hp (
<figureCitation id="EA8DAAA94526FFAFFF2F1829FE1E2EB4" box="[249,400,871,897]" captionStart="FIGURE 8" captionStartId="17.[151,250,1943,1967]" captionTargetBox="[196,1391,191,1914]" captionTargetId="figure-17@17.[186,1401,181,1919]" captionTargetPageId="17" captionText="FIGURE 8. Benincandona martini gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159200). BD. Hp (INV.159200). B. External view (drawing). C. Internal view (photograph with interference contrast). D. Internal view (photograph without interference contrast). Scale bar = 50 µm." pageId="19" pageNumber="20">Figure 8BD</figureCitation>
) relatively small, with bluntly pointed ls, reaching slightly beyond asymmetrically rounded ms; lobe h shorter and sub-rectangular, ls and ms with diverging positions; internal anatomy (
<figureCitation id="EA8DAAA94526FFAFFB1218C5FAD52E90" box="[1220,1371,907,933]" captionStart="FIGURE 8" captionStartId="17.[151,250,1943,1967]" captionTargetBox="[196,1391,191,1914]" captionTargetId="figure-17@17.[186,1401,181,1919]" captionTargetPageId="17" captionText="FIGURE 8. Benincandona martini gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159200). BD. Hp (INV.159200). B. External view (drawing). C. Internal view (photograph with interference contrast). D. Internal view (photograph without interference contrast). Scale bar = 50 µm." pageId="19" pageNumber="20">Figure 8C, D</figureCitation>
) with postlabyrinthal spermiduct with one circular coil.
</paragraph>
<paragraph id="7209B62C4526FFAFFF11189DFB892ED8" blockId="19.[151,1437,151,1005]" box="[199,1031,979,1005]" pageId="19" pageNumber="20">
Zenker organ (
<figureCitation id="EA8DAAA94526FFAFFEBD189DFE6F2ED8" box="[363,481,979,1005]" captionStart="FIGURE 8" captionStartId="17.[151,250,1943,1967]" captionTargetBox="[196,1391,191,1914]" captionTargetId="figure-17@17.[186,1401,181,1919]" captionTargetPageId="17" captionText="FIGURE 8. Benincandona martini gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159200). BD. Hp (INV.159200). B. External view (drawing). C. Internal view (photograph with interference contrast). D. Internal view (photograph without interference contrast). Scale bar = 50 µm." pageId="19" pageNumber="20">Figure 8A</figureCitation>
) short and broad, with eight-nine spinous whirls.
</paragraph>
<paragraph id="7209B62C4526FFAFFF411F54FE182901" blockId="19.[151,1436,1050,1545]" box="[151,406,1050,1076]" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFF411F54FE182901" bold="true" box="[151,406,1050,1076]" pageId="19" pageNumber="20">Description of female</emphasis>
</paragraph>
<paragraph id="7209B62C4526FFAFFF411F71FB10296C" blockId="19.[151,1436,1050,1545]" box="[151,1182,1087,1113]" pageId="19" pageNumber="20">
Cp and valves as in the male, without obvious sexual dimorphism (
<figureCitation id="EA8DAAA94526FFAFFC501F71FBCE296C" box="[902,1088,1087,1113]" captionStart="FIGURE 3" captionStartId="11.[151,250,1933,1957]" captionTargetBox="[172,1415,181,1910]" captionTargetId="figure-17@11.[172,1415,178,1910]" captionTargetPageId="11" captionText="FIGURE 3. Benincandona martini gen. et spec. nov., valves (A, C, EH, J ♂; B, D, I ♀). AD, I. CpRl, (A, C. INV.159208; B, D, I. INV.159211). E. INV.159209. CpD. F. INV.159210. CpV. G. INV.159200. LVi. H. INV.159200. RVi. J. INV.159200. CMS. Scale bars. 1 (A, B, E, F, G, H) = 500 µm; 2 (C, D) = 200 µm; 3 (I, J) = 50 µm." pageId="19" pageNumber="20">Figures 3B, D, I</figureCitation>
;
<figureCitation id="EA8DAAA94526FFAFFB9A1F71FB1C296C" box="[1100,1170,1087,1113]" captionStart="FIGURE 4" captionStartId="12.[151,250,1944,1968]" captionTargetBox="[157,1430,181,1920]" captionTargetId="figure-31@12.[157,1430,181,1920]" captionTargetPageId="12" captionText="FIGURE 4. Benincandona martini gen. et spec. nov., valves (AD: INV.159202 ♂ and EH. INV.159201 ♀) A, E. LVi. B, F. RVi, C, D, G, H. CMS. Scale bars. 1 (A, B, E, F) = 500 µm; 2 (C, D, G, H) = 50 µm." pageId="19" pageNumber="20">4EH</figureCitation>
).
</paragraph>
<paragraph id="7209B62C4526FFAFFF111F2DFBF52948" blockId="19.[151,1436,1050,1545]" box="[199,1147,1123,1149]" pageId="19" pageNumber="20">
Limbs (
<figureCitation id="EA8DAAA94526FFAFFECB1F2DFE3E2948" box="[285,432,1123,1149]" captionStart="FIGURE 9" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[172,1415,185,1915]" captionTargetId="figure-31@18.[168,1419,181,1919]" captionTargetPageId="18" captionText="FIGURE 9. Benincandona martini gen. et spec. nov., limbs (AB ♂ and CE ♀). A. T1, Lpp (INV.159202). B. T1, Rpp (INV.159202). C. A2 (INV.159205, INV.159207). D. CR (INV.159205). E. T1 (INV.159200). Scale bar = 50 µm." pageId="19" pageNumber="20">Figure 9CE</figureCitation>
) largely as in the male, but with sexually dimorphic A2 and T1.
</paragraph>
<paragraph id="7209B62C4526FFAFFF111FC9FC232838" blockId="19.[151,1436,1050,1545]" pageId="19" pageNumber="20">
A2 (
<figureCitation id="EA8DAAA94526FFAFFF2F1FC9FEE12994" box="[249,367,1159,1185]" captionStart="FIGURE 9" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[172,1415,185,1915]" captionTargetId="figure-31@18.[168,1419,181,1919]" captionTargetPageId="18" captionText="FIGURE 9. Benincandona martini gen. et spec. nov., limbs (AB ♂ and CE ♀). A. T1, Lpp (INV.159202). B. T1, Rpp (INV.159202). C. A2 (INV.159205, INV.159207). D. CR (INV.159205). E. T1 (INV.159200). Scale bar = 50 µm." pageId="19" pageNumber="20">Figure 9C</figureCitation>
) with second endopodal segment undivided, with two t-setae (one long and plumose (t1), one smooth short and thin (t2)); apical chaetotaxy (
<figureCitation id="EA8DAAA94526FFAFFD771FE5FC9E29F0" box="[673,784,1195,1221]" captionStart="FIGURE 40" captionStartId="60.[151,250,1516,1540]" captionTargetBox="[151,1435,181,1492]" captionTargetId="figure-31@60.[151,1435,181,1492]" captionTargetPageId="60" captionText="FIGURE 40. Diagrams of apical views of the A2 of Benincandona, showing position of (sub-) terminal claws, setae and aesthetascs of the terminal (small circle) and penultimate segment (large circle). For explanations of codes, see Material and Methods (Abbreviations used in text and figures)." pageId="19" pageNumber="20">Figure 40</figureCitation>
) with z1 claw-like, z2 and z3 being both setae, z1 twice as long as z2 and z3; claws G1, G2, G3 and GM all reaching to about the same point; seta g and aesthetasc y3 as in the male. Small claw Gm in female homologous to large claw in the male.
</paragraph>
<paragraph id="7209B62C4526FFAFFF111E59FC31284C" blockId="19.[151,1436,1050,1545]" pageId="19" pageNumber="20">
T1 (
<figureCitation id="EA8DAAA94526FFAFFF2F1E59FEE12804" box="[249,367,1303,1329]" captionStart="FIGURE 9" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[172,1415,185,1915]" captionTargetId="figure-31@18.[168,1419,181,1919]" captionTargetPageId="18" captionText="FIGURE 9. Benincandona martini gen. et spec. nov., limbs (AB ♂ and CE ♀). A. T1, Lpp (INV.159202). B. T1, Rpp (INV.159202). C. A2 (INV.159205, INV.159207). D. CR (INV.159205). E. T1 (INV.159200). Scale bar = 50 µm." pageId="19" pageNumber="20">Figure 9E</figureCitation>
) with basal part with
<emphasis id="40C26A3E4526FFAFFDA01E59FD072805" box="[630,649,1303,1328]" italics="true" pageId="19" pageNumber="20">c.</emphasis>
10 distal setae of unequal length, setae d and b present, with long additional seta (d) present close to seta d; respiratory plate reduced to about three short rays. Endopodite set with a broad and short palp, with three subequal but short apical setae (h1h3).
</paragraph>
<paragraph id="7209B62C4526FFAFFF111ECDFD8A28A8" blockId="19.[151,1436,1050,1545]" box="[199,516,1411,1437]" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFF111ECDFE0E28A8" bold="true" box="[199,384,1411,1437]" pageId="19" pageNumber="20">Measurements:</emphasis>
see
<tableCitation id="3F3483974526FFAFFE671ECDFD8A28A8" box="[433,516,1411,1437]" captionStart="TABLE 3" captionStartId="19.[152,244,1592,1616]" captionText="TABLE 3. Measurements of illustrated specimens of Benincandona gen. et spec. nov., using SEM." pageId="19" pageNumber="20">Table 3</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74526FFAFFF111EE8FDD328F4" box="[199,605,1446,1473]" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="7209B62C4526FFAFFF111EE8FDD328F4" blockId="19.[151,1436,1050,1545]" box="[199,605,1446,1473]" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFF111EE8FE5B28F5" bold="true" box="[199,469,1446,1472]" pageId="19" pageNumber="20">Differential Diagnosis:</emphasis>
See
<tableCitation id="3F3483974526FFAFFDDC1EE9FDD328F4" box="[522,605,1447,1473]" captionStart="TABLE 4" captionStartId="20.[152,245,1665,1689]" captionText="TABLE 4. Differential character states between the five new species of Benincandona gen. nov." pageId="19" pageNumber="20">Table 4</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74526FFAFFF111E85FA1E2AD6" pageId="19" pageNumber="20" type="biology_ecology">
<paragraph id="7209B62C4526FFAFFF111E85FE6128D1" blockId="19.[151,1436,1050,1545]" box="[199,495,1482,1509]" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFF111E85FE6128D1" bold="true" box="[199,495,1482,1509]" pageId="19" pageNumber="20">Ecology and distribution</emphasis>
</paragraph>
<paragraph id="7209B62C4526FFAFFF111EA1FBD02B3C" blockId="19.[151,1436,1050,1545]" box="[199,1118,1519,1545]" pageId="19" pageNumber="20">
This subterranean species is presently only known from its
<typeStatus id="AD0D088E4526FFAFFCB61EA1FC1E2B3C" box="[864,912,1519,1545]" pageId="19" pageNumber="20">type</typeStatus>
locality (
<tableCitation id="3F3483974526FFAFFC2C1EA1FBC32B3C" box="[1018,1101,1519,1545]" captionStart="TABLE 1" captionStartId="5.[152,244,929,955]" captionText="TABLE 1. Localities where Benincandona gen. nov. sp. nov. were collected, coordinates and hydro-morphological characteristics. In all stations, the water was clear and was used for both drinking water and household purposes by the riparian villages." pageId="19" pageNumber="20">Table 1</tableCitation>
).
</paragraph>
<caption id="26C9E6A44526FFAFFF4E1D76FB332B64" box="[152,1213,1592,1617]" pageId="19" pageNumber="20" startId="19.[152,244,1592,1616]" targetType="table">
<paragraph id="7209B62C4526FFAFFF4E1D76FB332B64" blockId="19.[152,1213,1592,1617]" box="[152,1213,1592,1617]" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFF4E1D76FE812B65" bold="true" box="[152,271,1592,1616]" pageId="19" pageNumber="20">TABLE 3.</emphasis>
Measurements of illustrated specimens of
<taxonomicName id="B5B6CDAF4526FFAFFD0F1D76FCFD2B65" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[729,883,1592,1616]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus" status="gen. et spec. nov.">
<emphasis id="40C26A3E4526FFAFFD0F1D76FCFD2B65" box="[729,883,1592,1616]" italics="true" pageId="19" pageNumber="20">Benincandona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454526FFAFFCAF1D77FBBA2B65" box="[889,1076,1593,1617]" pageId="19" pageNumber="20" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, using SEM.
</paragraph>
</caption>
<paragraph id="7209B62C4526FFAFFE8C1D2EFAF62AA7" pageId="19" pageNumber="20">
<table id="00B6448C45260043FF751D2EFA0B2AA7" box="[163,1413,1632,1938]" gridcols="7" gridrows="8" pageId="19" pageNumber="20">
<tr id="CC86B46E45260043FF751D2EFA0B2B4D" box="[163,1413,1632,1656]" gridrow="0" pageId="19" pageNumber="20">
<th id="8F57DD1245260043FF751D2EFE312B4D" box="[163,447,1632,1656]" gridcol="0" gridrow="0" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFE8C1D2EFE272B4D" bold="true" box="[346,425,1632,1656]" pageId="19" pageNumber="20">Species</emphasis>
</th>
<th id="8F57DD1245260043FD4D1D2EFD4F2B4D" box="[667,705,1632,1656]" gridcol="1" gridrow="0" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFD4D1D2EFD4F2B4D" bold="true" box="[667,705,1632,1656]" pageId="19" pageNumber="20">Sex</emphasis>
</th>
<th id="8F57DD1245260043FCD51D2EFCF12B4D" box="[771,895,1632,1656]" gridcol="2" gridrow="0" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFCD91D2EFCFD2B4D" bold="true" box="[783,883,1632,1656]" pageId="19" pageNumber="20">specimen</emphasis>
</th>
<th id="8F57DD1245260043FC611D2EFC7E2B4D" box="[951,1008,1632,1656]" gridcol="3" gridrow="0" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFC611D2EFC7E2B4D" bold="true" box="[951,1008,1632,1656]" pageId="19" pageNumber="20">V/Cp</emphasis>
</th>
<th id="8F57DD1245260043FBE71D2EFBF52B4D" box="[1073,1147,1632,1656]" gridcol="4" gridrow="0" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFBE71D2EFBF52B4D" bold="true" box="[1073,1147,1632,1656]" pageId="19" pageNumber="20">L (µm)</emphasis>
</th>
<th id="8F57DD1245260043FB641D2EFA8F2B4D" box="[1202,1281,1632,1656]" gridcol="5" gridrow="0" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFB641D2EFA8F2B4D" bold="true" box="[1202,1281,1632,1656]" pageId="19" pageNumber="20">H (µm)</emphasis>
</th>
<th id="8F57DD1245260043FAE41D2EFA0B2B4D" box="[1330,1413,1632,1656]" gridcol="6" gridrow="0" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFAE41D2EFA0B2B4D" bold="true" box="[1330,1413,1632,1656]" pageId="19" pageNumber="20">W (µm)</emphasis>
</th>
</tr>
<tr id="CC86B46E45260043FF751DC6FA0B2B94" box="[163,1413,1672,1697]" gridrow="1" pageId="19" pageNumber="20" rowspan-6="1">
<th id="8F57DD1245260043FF751DC6FE312B94" box="[163,447,1672,1697]" gridcol="0" gridrow="1" pageId="19" pageNumber="20">
<taxonomicName id="B5B6CDAF4526FFAFFF751DC7FE842B95" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[163,266,1673,1696]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="martini" status="gen. et spec. nov.">
<emphasis id="40C26A3E4526FFAFFF751DC7FE842B95" box="[163,266,1673,1696]" italics="true" pageId="19" pageNumber="20">B. martini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454526FFAFFEC61DC7FE312B95" box="[272,447,1673,1697]" pageId="19" pageNumber="20" rank="species">gen. et spec. nov.</taxonomicNameLabel>
</th>
<td id="8F57DD1245260043FD4D1DC6FD4F2B94" box="[667,705,1672,1697]" gridcol="1" gridrow="1" pageId="19" pageNumber="20"></td>
<td id="8F57DD1245260043FCD51DC6FCF12B94" box="[771,895,1672,1697]" gridcol="2" gridrow="1" pageId="19" pageNumber="20">INV.159200</td>
<td id="8F57DD1245260043FC611DC6FC7E2B94" box="[951,1008,1672,1697]" gridcol="3" gridrow="1" pageId="19" pageNumber="20">LVi</td>
<td id="8F57DD1245260043FBE71DC6FBF52B94" box="[1073,1147,1672,1697]" gridcol="4" gridrow="1" pageId="19" pageNumber="20">706.7</td>
<td id="8F57DD1245260043FB641DC6FA8F2B94" box="[1202,1281,1672,1697]" gridcol="5" gridrow="1" pageId="19" pageNumber="20">331.7</td>
</tr>
<tr id="CC86B46E45260043FF751DFFFA0B2BFC" box="[163,1413,1713,1737]" gridrow="2" pageId="19" pageNumber="20" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-6="1">
<td id="8F57DD1245260043FC611DFFFC7E2BFC" box="[951,1008,1713,1737]" gridcol="3" gridrow="2" pageId="19" pageNumber="20">RVi</td>
<td id="8F57DD1245260043FBE71DFFFBF52BFC" box="[1073,1147,1713,1737]" gridcol="4" gridrow="2" pageId="19" pageNumber="20">697.9</td>
<td id="8F57DD1245260043FB641DFFFA8F2BFC" box="[1202,1281,1713,1737]" gridcol="5" gridrow="2" pageId="19" pageNumber="20">310.7</td>
</tr>
<tr id="CC86B46E45260043FF751D97FA0B2BC4" box="[163,1413,1753,1777]" gridrow="3" pageId="19" pageNumber="20" rowspan-0="1" rowspan-1="1" rowspan-6="1">
<td id="8F57DD1245260043FCD51D97FCF12BC4" box="[771,895,1753,1777]" gridcol="2" gridrow="3" pageId="19" pageNumber="20">INV.159202</td>
<td id="8F57DD1245260043FC611D97FC7E2BC4" box="[951,1008,1753,1777]" gridcol="3" gridrow="3" pageId="19" pageNumber="20">LVi</td>
<td id="8F57DD1245260043FBE71D97FBF52BC4" box="[1073,1147,1753,1777]" gridcol="4" gridrow="3" pageId="19" pageNumber="20">667.5</td>
<td id="8F57DD1245260043FB641D97FA8F2BC4" box="[1202,1281,1753,1777]" gridcol="5" gridrow="3" pageId="19" pageNumber="20">317.5</td>
</tr>
<tr id="CC86B46E45260043FF751C4FFA0B2A2C" box="[163,1413,1793,1817]" gridrow="4" pageId="19" pageNumber="20" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-6="1">
<td id="8F57DD1245260043FC611C4FFC7E2A2C" box="[951,1008,1793,1817]" gridcol="3" gridrow="4" pageId="19" pageNumber="20">RVi</td>
<td id="8F57DD1245260043FBE71C4FFBF52A2C" box="[1073,1147,1793,1817]" gridcol="4" gridrow="4" pageId="19" pageNumber="20">656.7</td>
<td id="8F57DD1245260043FB641C4FFA8F2A2C" box="[1202,1281,1793,1817]" gridcol="5" gridrow="4" pageId="19" pageNumber="20">295.0</td>
</tr>
<tr id="CC86B46E45260043FF751C67FA0B2A77" box="[163,1413,1833,1858]" gridrow="5" pageId="19" pageNumber="20" rowspan-0="1" rowspan-1="1" rowspan-6="1">
<td id="8F57DD1245260043FCD51C67FCF12A77" box="[771,895,1833,1858]" gridcol="2" gridrow="5" pageId="19" pageNumber="20">INV.159208</td>
<td id="8F57DD1245260043FC611C67FC7E2A77" box="[951,1008,1833,1858]" gridcol="3" gridrow="5" pageId="19" pageNumber="20">CpRl</td>
<td id="8F57DD1245260043FBE71C67FBF52A77" box="[1073,1147,1833,1858]" gridcol="4" gridrow="5" pageId="19" pageNumber="20">692.9</td>
<td id="8F57DD1245260043FB641C67FA8F2A77" box="[1202,1281,1833,1858]" gridcol="5" gridrow="5" pageId="19" pageNumber="20">334.3</td>
</tr>
<tr id="CC86B46E45260043FF751C1CFA0B2A5F" box="[163,1413,1874,1898]" gridrow="6" pageId="19" pageNumber="20" rowspan-0="1" rowspan-1="1" rowspan-5="1">
<td id="8F57DD1245260043FCD51C1CFCF12A5F" box="[771,895,1874,1898]" gridcol="2" gridrow="6" pageId="19" pageNumber="20">INV.159209</td>
<td id="8F57DD1245260043FC611C1CFC7E2A5F" box="[951,1008,1874,1898]" gridcol="3" gridrow="6" pageId="19" pageNumber="20">CpD</td>
<td id="8F57DD1245260043FBE71C1CFBF52A5F" box="[1073,1147,1874,1898]" gridcol="4" gridrow="6" pageId="19" pageNumber="20">698.6</td>
<td id="8F57DD1245260043FAE41C1CFA0B2A5F" box="[1330,1413,1874,1898]" gridcol="6" gridrow="6" pageId="19" pageNumber="20">185.0</td>
</tr>
<tr id="CC86B46E45260043FF751C34FA0B2AA7" box="[163,1413,1914,1938]" gridrow="7" pageId="19" pageNumber="20" rowspan-0="1" rowspan-1="1" rowspan-5="1">
<td id="8F57DD1245260043FCD51C34FCF12AA7" box="[771,895,1914,1938]" gridcol="2" gridrow="7" pageId="19" pageNumber="20">INV.159210</td>
<td id="8F57DD1245260043FC611C34FC7E2AA7" box="[951,1008,1914,1938]" gridcol="3" gridrow="7" pageId="19" pageNumber="20">CpV</td>
<td id="8F57DD1245260043FBE71C34FBF52AA7" box="[1073,1147,1914,1938]" gridcol="4" gridrow="7" pageId="19" pageNumber="20">679.3</td>
<td id="8F57DD1245260043FAE41C34FA0B2AA7" box="[1330,1413,1914,1938]" gridcol="6" gridrow="7" pageId="19" pageNumber="20">190.7</td>
</tr>
</table>
</paragraph>
<paragraph id="7209B62C4526FFAFFB8F1C85FA1E2AD6" blockId="19.[1113,1424,1995,2019]" box="[1113,1424,1995,2019]" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFB8F1C85FA1E2AD6" box="[1113,1424,1995,2019]" italics="true" pageId="19" pageNumber="20">......continued on the next page</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>