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<mods:title id="4D4A0D151B25FBE10943EE3FBC596A4F">Parapaulipalpina lobata, new species, the first species of Parapaulipalpina Gnaspini, 1996 (Coleoptera: Leiodidae: Cholevinae: Ptomaphagini) from Brazil</mods:title>
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Key for the species of the genus
<taxonomicName id="4C254D203B150873FDC5B6F1FC26F9E2" ID-CoL="8XNMT" authority="Gnaspini, 1996" authorityName="Gnaspini" authorityYear="1996" box="[551,958,1533,1560]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="130" phylum="Arthropoda" rank="genus">
<emphasis id="B951EAB13B150873FDC5B6F1FD63F9ED" bold="true" box="[551,763,1533,1560]" italics="true" pageId="1" pageNumber="130">Parapaulipalpina</emphasis>
<bibRefCitation id="EFB44B523B150873FCE0B6F1FC26F9E2" author="Gnaspini, P." box="[770,958,1533,1559]" pageId="1" pageNumber="130" pagination="509 - 556" refId="ref4390" refString="Gnaspini, P. (1996) Phylogenetic analysis of the tribe Ptomaphagini, with description of new Neotropical genera and species (Coleoptera, Leiodidae, Cholevinae, Ptomaphagini). Papeis Avulsos de Zoologia, 39, 509 - 556, errata. https: // doi. org / 10.11606 / 0031 - 1049.1994.39. p 509 - 556" type="journal article" year="1996">Gnaspini, 1996</bibRefCitation>
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<paragraph id="8B9A36A33B150873FF75B54EFA04F9AD" blockId="1.[151,1437,1602,1947]" box="[151,1436,1602,1624]" pageId="1" pageNumber="130">
<keyLead id="30D495963B150873FF75B54EFA04F9AD" box="[151,1436,1602,1624]" pageId="1" pageNumber="130">
1a Head with strongly reduced eyes; flagellum of the aedeagus expanded apically;
<collectingCountry id="F33276333B150873FC03B54EFBD8F9AD" box="[993,1088,1602,1624]" name="Colombia" pageId="1" pageNumber="130">Colombia</collectingCountry>
......
<taxonomicName id="4C254D203B150873FB76B54FFA04F9AD" ID-CoL="4D7CV" authority="(Jeannel, 1936)" baseAuthorityName="Jeannel" baseAuthorityYear="1936" box="[1172,1436,1602,1624]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="130" phylum="Arthropoda" rank="species" species="filicornis">
<emphasis id="B951EAB13B150873FB76B54FFA9BF9A2" box="[1172,1283,1602,1624]" italics="true" pageId="1" pageNumber="130">P. filicornis</emphasis>
(
<bibRefCitation id="EFB44B523B150873FAEDB54EFA0DF9AD" author="Jeannel, R." box="[1295,1429,1602,1624]" pageId="1" pageNumber="130" pagination="1 - 433" refId="ref4903" refString="Jeannel, R. (1936) Monographie des Catopidae. Memoires du Museum National d'Histoire Naturelle, New Series, 1, 1 - 433." type="journal article" year="1936">Jeannel, 1936</bibRefCitation>
)
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<keyLead id="30D495963B150873FF75B56CFA04F980" box="[151,1436,1632,1654]" pageId="1" pageNumber="130">1b Eyes not strongly reduced; flagellum of the aedeagus not expanded apically...................................... 2</keyLead>
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2a Elytra truncate; male protarsomeres slender;
<collectingCountry id="F33276333B150873FD73B571FD25F967" box="[657,701,1661,1682]" name="Peru" pageId="1" pageNumber="130">Peru</collectingCountry>
....................................
<taxonomicName id="4C254D203B150873FB9DB572FA05F967" ID-CoL="6HLC6" authority="(Salgado, 2013)" baseAuthorityName="Salgado" baseAuthorityYear="2013" box="[1151,1437,1661,1683]" class="Insecta" family="Leiodidae" genus="Excelsiorella" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="130" phylum="Arthropoda" rank="species" species="tambopata">
<emphasis id="B951EAB13B150873FB9DB572FB65F967" box="[1151,1277,1661,1683]" italics="true" pageId="1" pageNumber="130">P. tambopata</emphasis>
(
<bibRefCitation id="EFB44B523B150873FAE8B571FA0DF967" author="Salgado" box="[1290,1429,1661,1683]" pageId="1" pageNumber="130" pagination="79 - 92" refId="ref5558" refString="Salgado *, J. M. (2013) Cholevinae de Peru (Coleoptera, Leiodidae): especies nuevas y nuevos datos. Boletin de la Sociedad Entomologica Aragonesa, 52, 79 - 92." type="journal article" year="2013">Salgado, 2013</bibRefCitation>
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<keyLead id="30D495963B150873FF75B596FA04F95A" box="[151,1436,1690,1712]" pageId="1" pageNumber="130">2b Elytra apices normally rounded; male protarsomeres normally expanded laterally.................................. 3</keyLead>
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3a Eyes slightly reduced; posterior margin of male metafemur straight; male metatibia largely widened and flattened internally close to its apex;
<collectingCountry id="F33276333B150873FE6BB5D9FE5CF91E" box="[393,452,1749,1771]" name="Brazil" pageId="1" pageNumber="130">Brazil</collectingCountry>
...................................................................
<taxonomicName id="4C254D203B150873FB17B5DAFAD1F91F" authorityName="Gnaspini &amp; Moraes &amp; Gomyde" authorityYear="2024" box="[1269,1353,1749,1771]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="130" phylum="Arthropoda" rank="species" species="lobata" status="sp. nov.">
<emphasis id="B951EAB13B150873FB17B5DAFAD1F91F" box="[1269,1353,1749,1771]" italics="true" pageId="1" pageNumber="130">P. lobata</emphasis>
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<keyLead id="30D495963B150873FF75B5FEFA04F8D0" pageId="1" pageNumber="130">3b Eyes normal; posterior margin of male metafemur bearing a tubercle medially; male metatibia curved inwards medially and not expanded apically.................................................................................... 4</keyLead>
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4a Last palpomere much longer than previous one; aedeagus concave ventrally, deeper about one third from base;
<collectingCountry id="F33276333B150873FAECB421FAA4F8B7" box="[1294,1340,1837,1858]" name="Peru" pageId="1" pageNumber="130">Peru</collectingCountry>
,
<collectingCountry id="F33276333B150873FAA4B421FA0EF8B6" box="[1350,1430,1837,1859]" name="Ecuador" pageId="1" pageNumber="130">Ecuador</collectingCountry>
..............................................................................
<taxonomicName id="4C254D203B150873FB79B447FA04F895" ID-CoL="4D7CW" authority="Salgado, 2005" authorityName="Salgado" authorityYear="2005" box="[1179,1436,1866,1888]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="130" phylum="Arthropoda" rank="species" species="giachinoi">
<emphasis id="B951EAB13B150873FB79B447FA95F8AA" box="[1179,1293,1866,1888]" italics="true" pageId="1" pageNumber="130">P. giachinoi</emphasis>
<bibRefCitation id="EFB44B523B150873FAF1B446FA04F895" author="Salgado" box="[1299,1436,1866,1888]" pageId="1" pageNumber="130" pagination="963 - 982" refId="ref5398" refString="Salgado *, J. M. (2005) New species of Leiodidae (Coleoptera) and new records from the Neotropical Region. Revue Suisse de Zoologie, 112, 963 - 982. https: // doi. org / 10.5962 / bhl. part. 80333" type="journal article" year="2005">Salgado, 2005</bibRefCitation>
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<keyLead id="30D495963B150873FF75B464FA04F86E" pageId="1" pageNumber="130">
4b Last palpomere as long as previous one; aedeagus concave ventrally, deeper medially, and convexly produced near apex;
<collectingCountry id="F33276333B150873FF00B489FEDDF86E" box="[226,325,1925,1947]" name="Venezuela" pageId="1" pageNumber="130">Venezuela</collectingCountry>
.......................................................................
<taxonomicName id="4C254D203B150873FB46B48AFA04F86E" ID-CoL="762WC" authority="Gnaspini, 1996" authorityName="Gnaspini" authorityYear="1996" box="[1188,1436,1925,1947]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="130" phylum="Arthropoda" rank="species" species="dentata">
<emphasis id="B951EAB13B150873FB46B48AFA9CF86F" box="[1188,1284,1925,1947]" italics="true" pageId="1" pageNumber="130">P. dentata</emphasis>
<bibRefCitation id="EFB44B523B150873FAE8B489FA04F86E" author="Gnaspini, P." box="[1290,1436,1925,1947]" pageId="1" pageNumber="130" pagination="509 - 556" refId="ref4390" refString="Gnaspini, P. (1996) Phylogenetic analysis of the tribe Ptomaphagini, with description of new Neotropical genera and species (Coleoptera, Leiodidae, Cholevinae, Ptomaphagini). Papeis Avulsos de Zoologia, 39, 509 - 556, errata. https: // doi. org / 10.11606 / 0031 - 1049.1994.39. p 509 - 556" type="journal article" year="1996">Gnaspini, 1996</bibRefCitation>
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<mods:url id="67FFB58CDE7C8388F29755731D9E4B2D">http://dx.doi.org/10.11646/zootaxa.5506.1.9</mods:url>
</mods:location>
<mods:classification id="DB175CCA76FB69FD9057BE2BDB970EAD">journal article</mods:classification>
<mods:identifier id="40325E7D5507DB4E76C73D58F66901EF" type="DOI">10.11646/zootaxa.5506.1.9</mods:identifier>
<mods:identifier id="704BCD872F7D69C844A8D7E7311193E8" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D04F554C451F1D028F5868AB93176AA8" type="ZooBank">24C421CD-0CF8-4422-8692-0086B4C671B9</mods:identifier>
</mods:mods>
<treatment id="038C87B53B160874FF75B39BFC34FAA5" LSID="urn:lsid:zoobank.org:act:B140C9C4-372B-4ACE-A029-F5EDFB62C895" httpUri="http://treatment.plazi.org/id/038C87B53B160874FF75B39BFC34FAA5" lastPageId="6" lastPageNumber="135" pageId="2" pageNumber="131">
<subSubSection id="C33F65283B160870FF75B39BFC49FF23" pageId="2" pageNumber="131" type="nomenclature">
<paragraph id="8B9A36A33B160870FF75B39BFC49FF23" blockId="2.[151,980,151,250]" pageId="2" pageNumber="131">
<heading id="D0D281CF3B160870FF75B39BFC4CFF44" bold="true" box="[151,980,151,178]" fontSize="11" level="1" pageId="2" pageNumber="131" reason="1">
<emphasis id="B951EAB13B160870FF75B39BFC4CFF44" bold="true" box="[151,980,151,178]" pageId="2" pageNumber="131">
<taxonomicName id="4C254D203B160870FF75B39BFCAFFF44" authority="Gnaspini, Moraes &amp; Gomyde" authorityName="Gnaspini, Moraes &amp; Gomyde" authorityYear="2024" box="[151,823,151,178]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="lobata" status="sp. nov.">
<emphasis id="B951EAB13B160870FF75B39BFE25FF47" bold="true" box="[151,445,151,178]" italics="true" pageId="2" pageNumber="131">Parapaulipalpina lobata</emphasis>
Gnaspini, Moraes &amp; Gomyde
</taxonomicName>
,
<taxonomicNameLabel id="A26257CA3B160870FCA6B394FC4CFF44" box="[836,980,151,178]" pageId="2" pageNumber="131" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
<uri id="FFB43AA13B160870FF75B3B7FC49FF23" box="[151,977,187,214]" pageId="2" pageNumber="131">
urn:lsid:zoobank.org:act:
<uuid id="FF830C763B160870FE4BB3B7FC49FF23" box="[425,977,187,214]" pageId="2" pageNumber="131">B140C9C4-372B-4ACE-A029-F5EDFB62C895</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C33F65283B160870FF75B3ECFE8AFF0C" box="[151,274,223,250]" pageId="2" pageNumber="131" type="description">
<paragraph id="8B9A36A33B160870FF75B3ECFE8AFF0C" blockId="2.[151,980,151,250]" box="[151,274,223,250]" pageId="2" pageNumber="131">
(
<figureCitation id="131E2A263B160870FF7DB3ECFE91FF0C" box="[159,265,223,250]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="2.[152,255,1189,1213]" captionStartId-1="2.[152,255,1819,1843]" captionStartId-2="3.[152,255,1116,1140]" captionStartId-3="3.[152,255,1940,1964]" captionStartId-4="4.[152,255,1324,1348]" captionStartId-5="4.[152,255,1844,1868]" captionTargetBox-0="[453,1135,526,1166]" captionTargetBox-1="[151,1436,1274,1796]" captionTargetBox-2="[158,1430,181,1092]" captionTargetBox-3="[296,1291,1309,1917]" captionTargetBox-4="[151,1436,181,1296]" captionTargetBox-5="[151,1436,1409,1820]" captionTargetId-0="figure-129@2.[453,1135,526,1166]" captionTargetId-1="figure-154@2.[151,1436,1274,1796]" captionTargetId-2="figure-17@3.[158,1430,181,1092]" captionTargetId-3="figure-131@3.[296,1291,1309,1917]" captionTargetId-4="figure-28@4.[151,1436,181,1301]" captionTargetId-5="figure-57@4.[151,1436,1409,1820]" captionTargetPageId-0="2" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="3" captionTargetPageId-4="4" captionTargetPageId-5="4" captionText-0="FIGURE 1. Parapaulipalpina lobata Gnaspini, Moraes &amp; Gomyde, new species, male paratype. Habitus, dorsal (A) and left lateral (b) views." captionText-1="FIGURE 2. Parapaulipalpina lobata Gnaspini, Moraes &amp; Gomyde, new species, male holotype. Head, dorsal (A) and left lateral (B) views." captionText-2="FIGURE 3. Parapaulipalpina lobata Gnaspini, Moraes &amp; Gomyde, new species, male holotype. A, protarsus, ventral view (detail of tenent setae is shown in B); CE, apex of protarsus in ventral (C), left lateral (D), and dorsal (E) views; F, hindleg (detail of apex of tibia and base of tarsus is shown in G); H, apex of metatibia, ventral view. est = empodial setae; mp = medial projection on the distal margin of the terminal tarsomere; mpp = pair of medial projections on the distal margin of the terminal tarsomere;?sdc = small dorsal spines close to the base of tarsal claw; slc = spines near base of tarsal claw." captionText-3="FIGURE 4. Parapaulipalpina lobata Gnaspini, Moraes &amp; Gomyde, new species. A, male holotype genital segment; B, C, male holotype aedeagus, ventral (B) and right lateral (C) views; D, female spermatheca. All figures to the same scale. fl = flagellum." captionText-4="FIGURE 5. Parapaulipalpina lobata Gnaspini, Moraes &amp; Gomyde, new species, male paratype in MZSP.A rotational view of the aedeagus, around its longitudinal axis." captionText-5="FIGURE 6. Parapaulipalpina lobata Gnaspini, Moraes &amp; Gomyde, new species, male paratype in MZSP.Apex of aedeagus in dorsal (A), left lateral (B), frontal (C), and ventral (D) views. do = dorsal opening; fl = flagellum; ll = left lobe; par = paralobe of the right lobe; rl = right lobe; tb = tubercles on the internal margin of the ventral opening." pageId="2" pageNumber="131">Figs. 16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33F65283B160874FF75B22BFC34FAA5" lastPageId="6" lastPageNumber="135" pageId="2" pageNumber="131" type="materials_examined">
<paragraph id="8B9A36A33B160870FF75B22BFD55FE03" blockId="2.[151,1436,295,502]" pageId="2" pageNumber="131">
<emphasis id="B951EAB13B160870FF75B22BFE1AFEB4" bold="true" box="[151,386,295,321]" pageId="2" pageNumber="131">Material examined.</emphasis>
<materialsCitation id="3B4D3CFE3B160870FE68B224FCAFFE93" collectingDate="1985-06" collectionCode="CMNC, FIT" collectorName="N. Degallier" country="Brazil" latitude="-3.7666667" location="Tucurui" longLatPrecision="1307" longitude="-49.666668" pageId="2" pageNumber="131" specimenCount="1" stateProvince="Para" typeStatus="holotype">
<typeStatus id="549E88013B160870FE68B224FE6DFEB7" box="[394,501,296,322]" pageId="2" pageNumber="131" type="holotype">Holotype</typeStatus>
, male (
<collectionCode id="ED34AE663B160870FDABB224FD3AFEB7" box="[585,674,296,322]" pageId="2" pageNumber="131">CMNC</collectionCode>
). Type locality and data:
<collectingCountry id="F33276333B160870FC23B224FB92FEB7" box="[961,1034,296,322]" name="Brazil" pageId="2" pageNumber="131">Brazil</collectingCountry>
:
<collectingRegion id="49E1F8413B160870FBF4B224FBD0FEB7" box="[1046,1096,296,322]" country="Brazil" name="Para" pageId="2" pageNumber="131">Pará</collectingRegion>
[State]:
<location id="8EFA60783B160870FB49B224FA9EFEB7" LSID="urn:lsid:plazi:treatment:038C87B53B160874FF75B39BFC34FAA5:8EFA60783B160870FB49B224FA9EFEB7" box="[1195,1286,296,322]" country="Brazil" latitude="-3.7666667" longLatPrecision="1307" longitude="-49.666668" name="Tucurui" pageId="2" pageNumber="131" stateProvince="Para">Tucuruí</location>
;
<date id="FF9B10633B160870FAF0B22BFAF3FEB7" box="[1298,1387,295,322]" pageId="2" pageNumber="131" value="1985-06">
<collectingDate id="EFDFE98B3B160870FAF0B22BFAF3FEB7" box="[1298,1387,295,322]" pageId="2" pageNumber="131" value="1985-06">vi.1985</collectingDate>
</date>
;
<geoCoordinate id="EE1150643B160870FA9AB224FF44FE93" degrees="49" direction="west" minutes="40" orientation="longitude" pageId="2" pageNumber="131" precision="925" value="-49.666668">49º 40 W</geoCoordinate>
<geoCoordinate id="EE1150643B160870FF01B240FEA7FE93" box="[227,319,331,358]" degrees="3" direction="south" minutes="46" orientation="latitude" pageId="2" pageNumber="131" precision="925" value="-3.7666667">3º 46 S</geoCoordinate>
;
<collectionCode id="ED34AE663B160870FEAEB240FEEFFE93" box="[332,375,332,358]" pageId="2" pageNumber="131">FIT</collectionCode>
, meat &amp; human dung;
<collectorName id="26D053753B160870FD98B240FC9DFE93" box="[634,773,332,358]" pageId="2" pageNumber="131">N. Degallier</collectorName>
col.
</materialsCitation>
<materialsCitation id="3B4D3CFE3B160870FCDCB240FD51FE03" collectingDate="1985-05" collectingDateMax="1985-07" collectingDateMin="1985-05" collectionCode="CMNC, IPEAN, FIT" collectorName="Belem &amp; N. Degallier &amp; Ipean &amp; de Pesquisa" country="Brazil" county="Empresa Brasileira de Pesquisa Agropecuaria" latitude="-3.7666667" location="Note" longLatPrecision="1307" longitude="-49.666668" municipality="Amazonia Oriental'" pageId="2" pageNumber="131" specimenCode="MZSP 61161" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Para" typeStatus="paratype">
<typeStatus id="549E88013B160870FCDCB240FC33FE93" box="[830,939,332,358]" pageId="2" pageNumber="131" type="paratype">Paratypes</typeStatus>
(
<collectionCode id="ED34AE663B160870FC59B240FB8AFE93" box="[955,1042,332,358]" pageId="2" pageNumber="131">CMNC</collectionCode>
, except when noted):
<specimenCount id="9D23FD2A3B160870FAE8B240FACEFE93" box="[1290,1366,332,358]" count="1" pageId="2" pageNumber="131" type="male">1 male</specimenCount>
and
<specimenCount id="9D23FD2A3B160870FA6CB240FF7BFE7F" count="1" pageId="2" pageNumber="131" type="female">1 female</specimenCount>
with same data;
<collectorName id="26D053753B160870FE43B27CFE75FE7F" box="[417,493,368,394]" pageId="2" pageNumber="131">Belém</collectorName>
,
<collectionCode id="ED34AE663B160870FE1AB27CFDD5FE7F" box="[504,589,368,394]" pageId="2" pageNumber="131">IPEAN</collectionCode>
,
<date id="FF9B10633B160870FDBAB263FD3EFE7F" box="[600,678,367,394]" pageId="2" pageNumber="131" value="1985-05">
<collectingDate id="EFDFE98B3B160870FDBAB263FD3EFE7F" box="[600,678,367,394]" pageId="2" pageNumber="131" value="1985-05">v.1985</collectingDate>
</date>
, meat,
<collectionCode id="ED34AE663B160870FD14B27CFCB9FE7F" box="[758,801,368,394]" pageId="2" pageNumber="131">FIT</collectionCode>
,
<collectorName id="26D053753B160870FCCEB27CFC20FE7F" box="[812,952,368,394]" pageId="2" pageNumber="131">N. Degallier</collectorName>
,
<specimenCount id="9D23FD2A3B160870FC21B27CFBABFE7F" box="[963,1075,368,394]" count="2" pageId="2" pageNumber="131" type="female">2 females</specimenCount>
;
<collectorName id="26D053753B160870FBDFB27CFBE6FE7F" box="[1085,1150,368,394]" pageId="2" pageNumber="131">Ipean</collectorName>
,
<collectorName id="26D053753B160870FB68B27CFB4EFE7F" box="[1162,1238,368,394]" pageId="2" pageNumber="131">Belém</collectorName>
,
<date id="FF9B10633B160870FB03B263FAA7FE7F" box="[1249,1343,367,394]" pageId="2" pageNumber="131" value="1985-07">
<collectingDate id="EFDFE98B3B160870FB03B263FAA7FE7F" box="[1249,1343,367,394]" pageId="2" pageNumber="131" value="1985-07">vii.1985</collectingDate>
</date>
,
<collectionCode id="ED34AE663B160870FAA9B27CFAEEFE7F" box="[1355,1398,368,394]" pageId="2" pageNumber="131">FIT</collectionCode>
,
<collectorName id="26D053753B160870FA63B27CFE99FE5B" pageId="2" pageNumber="131">N. Degallier</collectorName>
,
<specimenCount id="9D23FD2A3B160870FEEFB298FEC1FE5B" box="[269,345,404,430]" count="1" pageId="2" pageNumber="131" type="male">1 male</specimenCount>
and
<specimenCount id="9D23FD2A3B160870FE73B298FE6BFE5B" box="[401,499,404,430]" count="1" pageId="2" pageNumber="131" type="female">1 female</specimenCount>
(
<specimenCode id="DB839ED83B160870FDE0B298FD01FE58" box="[514,665,403,430]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="131" type="Museum">MZSP 61161</specimenCode>
and 61162). [
<location id="8EFA60783B160870FCD7B298FCEAFE5B" LSID="urn:lsid:plazi:treatment:038C87B53B160874FF75B39BFC34FAA5:8EFA60783B160870FCD7B298FCEAFE5B" box="[821,882,404,430]" county="Empresa Brasileira de Pesquisa Agropecuaria" municipality="Amazonia Oriental'" name="Note" pageId="2" pageNumber="131">Note</location>
:
<collectionCode id="ED34AE663B160870FC9FB298FC57FE5B" box="[893,975,404,430]" pageId="2" pageNumber="131">IPEAN</collectionCode>
(
<location id="8EFA60783B160870FC3DB298FB52FE5B" LSID="urn:lsid:plazi:treatment:038C87B53B160874FF75B39BFC34FAA5:8EFA60783B160870FC3DB298FB52FE5B" box="[991,1226,404,430]" county="Empresa Brasileira de Pesquisa Agropecuaria" municipality="Amazonia Oriental'" name="Instituto de Pesquisa" pageId="2" pageNumber="131">
Instituto
<collectorName id="26D053753B160870FBA4B298FB52FE5B" box="[1094,1226,404,430]" pageId="2" pageNumber="131">de Pesquisa</collectorName>
</location>
e
<location id="8EFA60783B160870FB04B298FE3CFE27" LSID="urn:lsid:plazi:treatment:038C87B53B160874FF75B39BFC34FAA5:8EFA60783B160870FB04B298FE3CFE27" county="Empresa Brasileira de Pesquisa Agropecuaria" municipality="Amazonia Oriental'" name="Experimentacao Agropecuaria do Norte" pageId="2" pageNumber="131">Experimentação Agropecuária do Norte</location>
) is presently in EMBRAPA
<collectingMunicipality id="6BFEACD93B160870FD1FB2B4FC47FE27" box="[765,991,440,466]" pageId="2" pageNumber="131">Amazônia Oriental</collectingMunicipality>
(EMBRAPA =
<collectingCounty id="62FB4E2F3B160870FB7FB2B4FE02FE03" pageId="2" pageNumber="131">Empresa Brasileira de Pesquisa Agropecuária</collectingCounty>
)—see
<bibRefCitation id="EFB44B523B160870FE05B2D0FD59FE03" author="EMBRAPA" box="[487,705,476,502]" pageId="2" pageNumber="131" refId="ref4308" refString="EMBRAPA (2024). Embrapa Amazonia Oriental. Digital resource. Available from: https: // www. embrapa. br / amazonia-oriental / historia (accessed 8 April 2024) [in Portuguese]" type="url" year="2024">EMBRAPA (2024)</bibRefCitation>
]
</materialsCitation>
.
</paragraph>
<caption id="DF5A662B3B160870FF7AB7A9FEDBFB14" pageId="2" pageNumber="131" startId="2.[152,255,1189,1213]" targetBox="[453,1135,526,1166]" targetPageId="2" targetType="figure">
<paragraph id="8B9A36A33B160870FF7AB7A9FEDBFB14" blockId="2.[151,1436,1189,1249]" pageId="2" pageNumber="131">
<emphasis id="B951EAB13B160870FF7AB7A9FE81FB48" bold="true" box="[152,281,1189,1213]" pageId="2" pageNumber="131">FIGURE 1.</emphasis>
<taxonomicName id="4C254D203B160870FEC2B7A9FCCCFB48" authority="Gnaspini, Moraes &amp; Gomyde" authorityName="Gnaspini, Moraes &amp; Gomyde" authorityYear="2024" box="[288,852,1189,1214]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="lobata" status="sp. nov.">
<emphasis id="B951EAB13B160870FEC2B7A9FD84FB48" box="[288,540,1189,1213]" italics="true" pageId="2" pageNumber="131">Parapaulipalpina lobata</emphasis>
Gnaspini, Moraes &amp; Gomyde
</taxonomicName>
,
<taxonomicNameLabel id="A26257CA3B160870FCBDB7AAFC44FB48" box="[863,988,1189,1213]" pageId="2" pageNumber="131" rank="species">new species</taxonomicNameLabel>
, male paratype. Habitus, dorsal (A) and left lateral (b) views.
</paragraph>
</caption>
<caption id="DF5A662B3B160870FF7AB417FEDFF8A2" pageId="2" pageNumber="131" startId="2.[152,255,1819,1843]" targetBox="[151,1436,1274,1796]" targetPageId="2" targetType="figure">
<paragraph id="8B9A36A33B160870FF7AB417FEDFF8A2" blockId="2.[151,1436,1819,1879]" pageId="2" pageNumber="131">
<emphasis id="B951EAB13B160870FF7AB417FE83F8C6" bold="true" box="[152,283,1819,1843]" pageId="2" pageNumber="131">FIGURE 2.</emphasis>
<taxonomicName id="4C254D203B160870FEC1B417FCC6F8C6" authority="Gnaspini, Moraes &amp; Gomyde" authorityName="Gnaspini, Moraes &amp; Gomyde" authorityYear="2024" box="[291,862,1819,1843]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="lobata" status="sp. nov.">
<emphasis id="B951EAB13B160870FEC1B417FDB9F8C6" box="[291,545,1819,1843]" italics="true" pageId="2" pageNumber="131">Parapaulipalpina lobata</emphasis>
Gnaspini, Moraes &amp; Gomyde
</taxonomicName>
,
<taxonomicNameLabel id="A26257CA3B160870FC89B417FC71F8C6" box="[875,1001,1819,1843]" pageId="2" pageNumber="131" rank="species">new species</taxonomicNameLabel>
, male holotype. Head, dorsal (A) and left lateral (B) views.
</paragraph>
</caption>
<caption id="DF5A662B3B170871FF7AB750FB53FAF1" pageId="3" pageNumber="132" startId="3.[152,255,1116,1140]" targetBox="[158,1430,181,1092]" targetPageId="3" targetType="figure">
<paragraph id="8B9A36A33B170871FF7AB750FB53FAF1" blockId="3.[151,1436,1116,1284]" pageId="3" pageNumber="132">
<emphasis id="B951EAB13B170871FF7AB750FE82FB81" bold="true" box="[152,282,1116,1140]" pageId="3" pageNumber="132">FIGURE 3.</emphasis>
<taxonomicName id="4C254D203B170871FEC0B750FCC4FB81" authority="Gnaspini, Moraes &amp; Gomyde" authorityName="Gnaspini, Moraes &amp; Gomyde" authorityYear="2024" box="[290,860,1116,1140]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="132" phylum="Arthropoda" rank="species" species="lobata" status="sp. nov.">
<emphasis id="B951EAB13B170871FEC0B750FDB8FB81" box="[290,544,1116,1140]" italics="true" pageId="3" pageNumber="132">Parapaulipalpina lobata</emphasis>
Gnaspini, Moraes &amp; Gomyde
</taxonomicName>
,
<taxonomicNameLabel id="A26257CA3B170871FC85B751FC7EFB81" box="[871,998,1116,1140]" pageId="3" pageNumber="132" rank="species">new species</taxonomicNameLabel>
, male holotype. A, protarsus, ventral view (detail of tenent setae is shown in B); CE, apex of protarsus in ventral (C), left lateral (D), and dorsal (E) views; F, hindleg (detail of apex of tibia and base of tarsus is shown in G); H, apex of metatibia, ventral view. est = empodial setae; mp = medial projection on the distal margin of the terminal tarsomere; mpp = pair of medial projections on the distal margin of the terminal tarsomere;?sdc = small dorsal spines close to the base of tarsal claw; slc = spines near base of tarsal claw.
</paragraph>
</caption>
<caption id="DF5A662B3B170871FF7AB498FE98F801" pageId="3" pageNumber="132" startId="3.[152,255,1940,1964]" targetBox="[296,1291,1309,1917]" targetPageId="3" targetType="figure">
<paragraph id="8B9A36A33B170871FF7AB498FE98F801" blockId="3.[151,1436,1940,2036]" pageId="3" pageNumber="132">
<emphasis id="B951EAB13B170871FF7AB498FE83F859" bold="true" box="[152,283,1940,1964]" pageId="3" pageNumber="132">FIGURE 4.</emphasis>
<taxonomicName id="4C254D203B170871FEC6B498FCF8F859" authority="Gnaspini, Moraes &amp; Gomyde" authorityName="Gnaspini, Moraes &amp; Gomyde" authorityYear="2024" box="[292,864,1940,1964]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="132" phylum="Arthropoda" rank="species" species="lobata" status="sp. nov.">
<emphasis id="B951EAB13B170871FEC6B498FDBAF859" box="[292,546,1940,1964]" italics="true" pageId="3" pageNumber="132">Parapaulipalpina lobata</emphasis>
Gnaspini, Moraes &amp; Gomyde
</taxonomicName>
,
<taxonomicNameLabel id="A26257CA3B170871FC8EB499FC73F859" box="[876,1003,1940,1964]" pageId="3" pageNumber="132" rank="species">new species</taxonomicNameLabel>
. A, male holotype genital segment; B, C, male holotype aedeagus, ventral (B) and right lateral (C) views; D, female spermatheca. All figures to the same scale. fl = flagellum.
</paragraph>
</caption>
<caption id="DF5A662B3B100876FF7AB620FDC9FA9C" pageId="4" pageNumber="133" startId="4.[152,255,1324,1348]" targetBox="[151,1436,181,1296]" targetPageId="4" targetType="figure">
<paragraph id="8B9A36A33B100876FF7AB620FDC9FA9C" blockId="4.[151,1436,1324,1385]" pageId="4" pageNumber="133">
<emphasis id="B951EAB13B100876FF7AB620FE80FAB0" bold="true" box="[152,280,1324,1349]" pageId="4" pageNumber="133">FIGURE 5.</emphasis>
<taxonomicName id="4C254D203B100876FEFCB621FCD3FAB0" authority="Gnaspini, Moraes &amp; Gomyde" authorityName="Gnaspini, Moraes &amp; Gomyde" authorityYear="2024" box="[286,843,1325,1349]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="133" phylum="Arthropoda" rank="species" species="lobata" status="sp. nov.">
<emphasis id="B951EAB13B100876FEFCB621FD81FAB0" box="[286,537,1325,1349]" italics="true" pageId="4" pageNumber="133">Parapaulipalpina lobata</emphasis>
Gnaspini, Moraes &amp; Gomyde
</taxonomicName>
,
<taxonomicNameLabel id="A26257CA3B100876FCB7B621FC49FAB0" box="[853,977,1325,1349]" pageId="4" pageNumber="133" rank="species">new species</taxonomicNameLabel>
, male paratype in MZSP. A rotational view of the aedeagus, around its longitudinal axis.
</paragraph>
</caption>
<caption id="DF5A662B3B100876FF7AB438FBD5F861" pageId="4" pageNumber="133" startId="4.[152,255,1844,1868]" targetBox="[151,1436,1409,1820]" targetPageId="4" targetType="figure">
<paragraph id="8B9A36A33B100876FF7AB438FBD5F861" blockId="4.[151,1437,1844,1940]" pageId="4" pageNumber="133">
<emphasis id="B951EAB13B100876FF7AB438FE8FF8B9" bold="true" box="[152,279,1844,1868]" pageId="4" pageNumber="133">FIGURE 6.</emphasis>
<taxonomicName id="4C254D203B100876FEFFB438FCD2F8B9" authority="Gnaspini, Moraes &amp; Gomyde" authorityName="Gnaspini, Moraes &amp; Gomyde" authorityYear="2024" box="[285,842,1844,1868]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="133" phylum="Arthropoda" rank="species" species="lobata" status="sp. nov.">
<emphasis id="B951EAB13B100876FEFFB438FD80F8B9" box="[285,536,1844,1868]" italics="true" pageId="4" pageNumber="133">Parapaulipalpina lobata</emphasis>
Gnaspini, Moraes &amp; Gomyde
</taxonomicName>
,
<taxonomicNameLabel id="A26257CA3B100876FCB1B438FC57F8B9" box="[851,975,1844,1868]" pageId="4" pageNumber="133" rank="species">new species</taxonomicNameLabel>
, male paratype in MZSP. Apex of aedeagus in dorsal (A), left lateral (B), frontal (C), and ventral (D) views. do = dorsal opening; fl = flagellum; ll = left lobe; par = paralobe of the right lobe; rl = right lobe; tb = tubercles on the internal margin of the ventral opening.
</paragraph>
</caption>
<paragraph id="8B9A36A33B110877FF25B39BFC80FF20" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="134">
Measurements: Length:
<quantity id="4CDD9B463B110877FE35B39AFDA3FF44" box="[471,571,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="5" pageNumber="134" unit="mm" value="1.05">1.05 mm</quantity>
(
<typeStatus id="549E88013B110877FDABB39BFD29FF44" box="[585,689,151,177]" pageId="5" pageNumber="134" type="holotype">holotype</typeStatus>
),
<quantity id="4CDD9B463B110877FD26B39BFCD2FF44" box="[708,842,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="5" pageNumber="134" unit="mm" value="1.2" valueMax="1.3" valueMin="1.1">1.11.3 mm</quantity>
(other males),
<quantity id="4CDD9B463B110877FC11B39BFBE2FF44" box="[1011,1146,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="1.2999999999999998" metricValueMax="1.4" metricValueMin="1.2" pageId="5" pageNumber="134" unit="mm" value="1.2999999999999998" valueMax="1.4" valueMin="1.2">1.21.4 mm</quantity>
(females); width:
<quantity id="4CDD9B463B110877FAA5B39BFA05FF44" box="[1351,1437,151,177]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="134" unit="mm" value="0.6">0.6 mm</quantity>
(
<typeStatus id="549E88013B110877FF7DB3B7FE9FFF20" box="[159,263,187,213]" pageId="5" pageNumber="134" type="holotype">holotype</typeStatus>
),
<quantity id="4CDD9B463B110877FEF8B3B7FEE8FF20" box="[282,368,187,213]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="134" unit="mm" value="0.8">0.8 mm</quantity>
(other males),
<quantity id="4CDD9B463B110877FDFBB3B7FD38FF20" box="[537,672,187,213]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="5" pageNumber="134" unit="mm" value="0.85" valueMax="0.9" valueMin="0.8">0.80.9 mm</quantity>
(females).
</paragraph>
<paragraph id="8B9A36A33B110877FF25B3D3FA93FBC0" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="134">
<emphasis id="B951EAB13B110877FF25B3D3FE68FF0D" box="[199,496,223,249]" italics="true" pageId="5" pageNumber="134">Diagnosis and Description</emphasis>
. General characteristics as listed above. Body ovoid, convex; color dark redish brown (
<figureCitation id="131E2A263B110877FF7DB20FFF79FEE8" box="[159,225,259,285]" captionStart="FIGURE 1" captionStartId="2.[152,255,1189,1213]" captionTargetBox="[453,1135,526,1166]" captionTargetId="figure-129@2.[453,1135,526,1166]" captionTargetPageId="2" captionText="FIGURE 1. Parapaulipalpina lobata Gnaspini, Moraes &amp; Gomyde, new species, male paratype. Habitus, dorsal (A) and left lateral (b) views." pageId="5" pageNumber="134">Fig. 1</figureCitation>
). Winged [metathoracic wing fully developed]. Head longer than high in lateral view (
<figureCitation id="131E2A263B110877FB53B20FFA91FEE8" box="[1201,1289,259,285]" captionStart="FIGURE 2" captionStartId="2.[152,255,1819,1843]" captionTargetBox="[151,1436,1274,1796]" captionTargetId="figure-154@2.[151,1436,1274,1796]" captionTargetPageId="2" captionText="FIGURE 2. Parapaulipalpina lobata Gnaspini, Moraes &amp; Gomyde, new species, male holotype. Head, dorsal (A) and left lateral (B) views." pageId="5" pageNumber="134">Fig. 2B</figureCitation>
); integument punctuated, punctures not organized in strigae (
<figureCitation id="131E2A263B110877FD49B22BFCA8FEB4" box="[683,816,295,321]" captionStart="FIGURE 2" captionStartId="2.[152,255,1819,1843]" captionTargetBox="[151,1436,1274,1796]" captionTargetId="figure-154@2.[151,1436,1274,1796]" captionTargetPageId="2" captionText="FIGURE 2. Parapaulipalpina lobata Gnaspini, Moraes &amp; Gomyde, new species, male holotype. Head, dorsal (A) and left lateral (B) views." pageId="5" pageNumber="134">Figs. 2A, B</figureCitation>
); eyes slightly reduced (
<figureCitation id="131E2A263B110877FBA0B22BFB02FEB4" box="[1090,1178,295,321]" captionStart="FIGURE 2" captionStartId="2.[152,255,1819,1843]" captionTargetBox="[151,1436,1274,1796]" captionTargetId="figure-154@2.[151,1436,1274,1796]" captionTargetPageId="2" captionText="FIGURE 2. Parapaulipalpina lobata Gnaspini, Moraes &amp; Gomyde, new species, male holotype. Head, dorsal (A) and left lateral (B) views." pageId="5" pageNumber="134">Fig. 2B</figureCitation>
). Pronotum transverse, widest at base; posterior angles acute; integument with [light] transverse strigae. Elytra together 1.25 times as long as wide, with transverse, diagonal, strigae. All legs with a crown of flat and equal sized spines around the tibia apex (e.g.,
<figureCitation id="131E2A263B110877FF34B29FFEB6FE58" box="[214,302,403,429]" captionStart="FIGURE 3" captionStartId="3.[152,255,1116,1140]" captionTargetBox="[158,1430,181,1092]" captionTargetId="figure-17@3.[158,1430,181,1092]" captionTargetPageId="3" captionText="FIGURE 3. Parapaulipalpina lobata Gnaspini, Moraes &amp; Gomyde, new species, male holotype. A, protarsus, ventral view (detail of tenent setae is shown in B); CE, apex of protarsus in ventral (C), left lateral (D), and dorsal (E) views; F, hindleg (detail of apex of tibia and base of tarsus is shown in G); H, apex of metatibia, ventral view. est = empodial setae; mp = medial projection on the distal margin of the terminal tarsomere; mpp = pair of medial projections on the distal margin of the terminal tarsomere;?sdc = small dorsal spines close to the base of tarsal claw; slc = spines near base of tarsal claw." pageId="5" pageNumber="134">Fig. 3G</figureCitation>
). First
<specimenCount id="9D23FD2A3B110877FE99B29FFE7DFE58" box="[379,485,403,429]" count="4" pageId="5" pageNumber="134" type="male">four male</specimenCount>
protarsomeres expanded, with elongate discoidal tenent setae ventrally (
<figureCitation id="131E2A263B110877FAE9B29FFA16FE58" box="[1291,1422,403,429]" captionStart="FIGURE 3" captionStartId="3.[152,255,1116,1140]" captionTargetBox="[158,1430,181,1092]" captionTargetId="figure-17@3.[158,1430,181,1092]" captionTargetPageId="3" captionText="FIGURE 3. Parapaulipalpina lobata Gnaspini, Moraes &amp; Gomyde, new species, male holotype. A, protarsus, ventral view (detail of tenent setae is shown in B); CE, apex of protarsus in ventral (C), left lateral (D), and dorsal (E) views; F, hindleg (detail of apex of tibia and base of tarsus is shown in G); H, apex of metatibia, ventral view. est = empodial setae; mp = medial projection on the distal margin of the terminal tarsomere; mpp = pair of medial projections on the distal margin of the terminal tarsomere;?sdc = small dorsal spines close to the base of tarsal claw; slc = spines near base of tarsal claw." pageId="5" pageNumber="134">Figs. 3A, B</figureCitation>
). Mesotibia curved inwards in both sexes. Metatibia straight; in males it is largely widened and flattened internally close to its apex (
<figureCitation id="131E2A263B110877FE83B2D7FE7DFE00" box="[353,485,475,501]" captionStart="FIGURE 3" captionStartId="3.[152,255,1116,1140]" captionTargetBox="[158,1430,181,1092]" captionTargetId="figure-17@3.[158,1430,181,1092]" captionTargetPageId="3" captionText="FIGURE 3. Parapaulipalpina lobata Gnaspini, Moraes &amp; Gomyde, new species, male holotype. A, protarsus, ventral view (detail of tenent setae is shown in B); CE, apex of protarsus in ventral (C), left lateral (D), and dorsal (E) views; F, hindleg (detail of apex of tibia and base of tarsus is shown in G); H, apex of metatibia, ventral view. est = empodial setae; mp = medial projection on the distal margin of the terminal tarsomere; mpp = pair of medial projections on the distal margin of the terminal tarsomere;?sdc = small dorsal spines close to the base of tarsal claw; slc = spines near base of tarsal claw." pageId="5" pageNumber="134">Figs. 3F, G</figureCitation>
). Tarsi of all legs (in both sexes) bearing a pair of empodial setae with asymmetric length, with different proportions on different legs (
<figureCitation id="131E2A263B110877FD13B2F3FCE4FDEC" box="[753,892,511,537]" captionStart="FIGURE 3" captionStartId="3.[152,255,1116,1140]" captionTargetBox="[158,1430,181,1092]" captionTargetId="figure-17@3.[158,1430,181,1092]" captionTargetPageId="3" captionText="FIGURE 3. Parapaulipalpina lobata Gnaspini, Moraes &amp; Gomyde, new species, male holotype. A, protarsus, ventral view (detail of tenent setae is shown in B); CE, apex of protarsus in ventral (C), left lateral (D), and dorsal (E) views; F, hindleg (detail of apex of tibia and base of tarsus is shown in G); H, apex of metatibia, ventral view. est = empodial setae; mp = medial projection on the distal margin of the terminal tarsomere; mpp = pair of medial projections on the distal margin of the terminal tarsomere;?sdc = small dorsal spines close to the base of tarsal claw; slc = spines near base of tarsal claw." pageId="5" pageNumber="134">Figs. 3C, H</figureCitation>
, est); distal margin of the terminal tarsomere with a pair of medial projections (
<figureCitation id="131E2A263B110877FDFFB12FFDEEFDC8" box="[541,630,547,573]" captionStart="FIGURE 3" captionStartId="3.[152,255,1116,1140]" captionTargetBox="[158,1430,181,1092]" captionTargetId="figure-17@3.[158,1430,181,1092]" captionTargetPageId="3" captionText="FIGURE 3. Parapaulipalpina lobata Gnaspini, Moraes &amp; Gomyde, new species, male holotype. A, protarsus, ventral view (detail of tenent setae is shown in B); CE, apex of protarsus in ventral (C), left lateral (D), and dorsal (E) views; F, hindleg (detail of apex of tibia and base of tarsus is shown in G); H, apex of metatibia, ventral view. est = empodial setae; mp = medial projection on the distal margin of the terminal tarsomere; mpp = pair of medial projections on the distal margin of the terminal tarsomere;?sdc = small dorsal spines close to the base of tarsal claw; slc = spines near base of tarsal claw." pageId="5" pageNumber="134">Fig. 3C</figureCitation>
, mpp), except on the hindtarsi, in which the projection is simple and triangular (
<figureCitation id="131E2A263B110877FEF7B14BFEF7FD94" box="[277,367,583,609]" captionStart="FIGURE 3" captionStartId="3.[152,255,1116,1140]" captionTargetBox="[158,1430,181,1092]" captionTargetId="figure-17@3.[158,1430,181,1092]" captionTargetPageId="3" captionText="FIGURE 3. Parapaulipalpina lobata Gnaspini, Moraes &amp; Gomyde, new species, male holotype. A, protarsus, ventral view (detail of tenent setae is shown in B); CE, apex of protarsus in ventral (C), left lateral (D), and dorsal (E) views; F, hindleg (detail of apex of tibia and base of tarsus is shown in G); H, apex of metatibia, ventral view. est = empodial setae; mp = medial projection on the distal margin of the terminal tarsomere; mpp = pair of medial projections on the distal margin of the terminal tarsomere;?sdc = small dorsal spines close to the base of tarsal claw; slc = spines near base of tarsal claw." pageId="5" pageNumber="134">Fig. 3H</figureCitation>
, mp); claws with a few long and acuminate spines at their dorsolateral margin (
<figureCitation id="131E2A263B110877FAF3B14BFA0FFD94" box="[1297,1431,583,609]" captionStart="FIGURE 3" captionStartId="3.[152,255,1116,1140]" captionTargetBox="[158,1430,181,1092]" captionTargetId="figure-17@3.[158,1430,181,1092]" captionTargetPageId="3" captionText="FIGURE 3. Parapaulipalpina lobata Gnaspini, Moraes &amp; Gomyde, new species, male holotype. A, protarsus, ventral view (detail of tenent setae is shown in B); CE, apex of protarsus in ventral (C), left lateral (D), and dorsal (E) views; F, hindleg (detail of apex of tibia and base of tarsus is shown in G); H, apex of metatibia, ventral view. est = empodial setae; mp = medial projection on the distal margin of the terminal tarsomere; mpp = pair of medial projections on the distal margin of the terminal tarsomere;?sdc = small dorsal spines close to the base of tarsal claw; slc = spines near base of tarsal claw." pageId="5" pageNumber="134">Figs. 3D, E</figureCitation>
, slc); the typical small dorsal spines close to the base of the claws of
<taxonomicName id="4C254D203B110877FC7BB167FBADFD70" authorityName="Jeannel" authorityYear="1911" box="[921,1077,619,645]" class="Insecta" family="Leiodidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="134" phylum="Arthropoda" rank="tribe" tribe="Ptomaphagini">Ptomaphagini</taxonomicName>
are not clear in the pictures here analysed and may be represented by the microtrichia observed around the base of the claws as in
<figureCitation id="131E2A263B110877FB01B183FAAFFD5C" box="[1251,1335,655,681]" captionStart="FIGURE 3" captionStartId="3.[152,255,1116,1140]" captionTargetBox="[158,1430,181,1092]" captionTargetId="figure-17@3.[158,1430,181,1092]" captionTargetPageId="3" captionText="FIGURE 3. Parapaulipalpina lobata Gnaspini, Moraes &amp; Gomyde, new species, male holotype. A, protarsus, ventral view (detail of tenent setae is shown in B); CE, apex of protarsus in ventral (C), left lateral (D), and dorsal (E) views; F, hindleg (detail of apex of tibia and base of tarsus is shown in G); H, apex of metatibia, ventral view. est = empodial setae; mp = medial projection on the distal margin of the terminal tarsomere; mpp = pair of medial projections on the distal margin of the terminal tarsomere;?sdc = small dorsal spines close to the base of tarsal claw; slc = spines near base of tarsal claw." pageId="5" pageNumber="134">Fig. 3E</figureCitation>
(?sdc). Male genital segment round, with spiculum gastrale straight and long (
<figureCitation id="131E2A263B110877FC2AB1BFFBBEFD38" box="[968,1062,691,717]" captionStart="FIGURE 4" captionStartId="3.[152,255,1940,1964]" captionTargetBox="[296,1291,1309,1917]" captionTargetId="figure-131@3.[296,1291,1309,1917]" captionTargetPageId="3" captionText="FIGURE 4. Parapaulipalpina lobata Gnaspini, Moraes &amp; Gomyde, new species. A, male holotype genital segment; B, C, male holotype aedeagus, ventral (B) and right lateral (C) views; D, female spermatheca. All figures to the same scale. fl = flagellum." pageId="5" pageNumber="134">Fig. 4A</figureCitation>
). Aedeagus elongate (~4.2 times as long as wide, in dorsal view), with sides subparallel (
<figureCitation id="131E2A263B110877FCF3B1DBFCCBFD04" box="[785,851,726,753]" captionStart="FIGURE 5" captionStartId="4.[152,255,1324,1348]" captionTargetBox="[151,1436,181,1296]" captionTargetId="figure-28@4.[151,1436,181,1301]" captionTargetPageId="4" captionText="FIGURE 5. Parapaulipalpina lobata Gnaspini, Moraes &amp; Gomyde, new species, male paratype in MZSP.A rotational view of the aedeagus, around its longitudinal axis." pageId="5" pageNumber="134">Fig. 5</figureCitation>
)—widest near base and slightly narrowing towards apex (
<figureCitation id="131E2A263B110877FF3EB1F7FEAEFCE0" box="[220,310,762,789]" captionStart="FIGURE 5" captionStartId="4.[152,255,1324,1348]" captionTargetBox="[151,1436,181,1296]" captionTargetId="figure-28@4.[151,1436,181,1301]" captionTargetPageId="4" captionText="FIGURE 5. Parapaulipalpina lobata Gnaspini, Moraes &amp; Gomyde, new species, male paratype in MZSP.A rotational view of the aedeagus, around its longitudinal axis." pageId="5" pageNumber="134">Fig. 5A</figureCitation>
), somewhat S-shaped in dorsal view (
<figureCitation id="131E2A263B110877FD06B1F7FCDFFCE0" box="[740,839,763,789]" captionStart="FIGURE 4" captionStartId="3.[152,255,1940,1964]" captionTargetBox="[296,1291,1309,1917]" captionTargetId="figure-131@3.[296,1291,1309,1917]" captionTargetPageId="3" captionText="FIGURE 4. Parapaulipalpina lobata Gnaspini, Moraes &amp; Gomyde, new species. A, male holotype genital segment; B, C, male holotype aedeagus, ventral (B) and right lateral (C) views; D, female spermatheca. All figures to the same scale. fl = flagellum." pageId="5" pageNumber="134">Figs. 4B</figureCitation>
,
<figureCitation id="131E2A263B110877FCB6B1F6FCE0FCE0" box="[852,888,762,789]" captionStart="FIGURE 5" captionStartId="4.[152,255,1324,1348]" captionTargetBox="[151,1436,181,1296]" captionTargetId="figure-28@4.[151,1436,181,1301]" captionTargetPageId="4" captionText="FIGURE 5. Parapaulipalpina lobata Gnaspini, Moraes &amp; Gomyde, new species, male paratype in MZSP.A rotational view of the aedeagus, around its longitudinal axis." pageId="5" pageNumber="134">5A</figureCitation>
), with a small, subelliptical dorsal opening (
<figureCitation id="131E2A263B110877FA8DB1F7FF24FCCC" captionStart="FIGURE 6" captionStartId="4.[152,255,1844,1868]" captionTargetBox="[151,1436,1409,1820]" captionTargetId="figure-57@4.[151,1436,1409,1820]" captionTargetPageId="4" captionText="FIGURE 6. Parapaulipalpina lobata Gnaspini, Moraes &amp; Gomyde, new species, male paratype in MZSP.Apex of aedeagus in dorsal (A), left lateral (B), frontal (C), and ventral (D) views. do = dorsal opening; fl = flagellum; ll = left lobe; par = paralobe of the right lobe; rl = right lobe; tb = tubercles on the internal margin of the ventral opening." pageId="5" pageNumber="134">Fig. 6A</figureCitation>
, do); curved ventrad (i.e., concave—concavity/total height = ~52% [
<figureCitation id="131E2A263B110877FC35B013FBFCFCCC" box="[983,1124,798,825]" captionStart="FIGURE 5" captionStartId="4.[152,255,1324,1348]" captionTargetBox="[151,1436,181,1296]" captionTargetId="figure-28@4.[151,1436,181,1301]" captionTargetPageId="4" captionText="FIGURE 5. Parapaulipalpina lobata Gnaspini, Moraes &amp; Gomyde, new species, male paratype in MZSP.A rotational view of the aedeagus, around its longitudinal axis." pageId="5" pageNumber="134">Figs. 5E, M</figureCitation>
]), with the axis of the basal opening pointing about 45º downwards in relation to the sagittal plane of the aedeagus); the paralobe of the right lobe elongate and curved ventrally (
<figureCitation id="131E2A263B110877FDC1B06BFD04FC74" box="[547,668,871,897]" captionStart="FIGURE 6" captionStartId="4.[152,255,1844,1868]" captionTargetBox="[151,1436,1409,1820]" captionTargetId="figure-57@4.[151,1436,1409,1820]" captionTargetPageId="4" captionText="FIGURE 6. Parapaulipalpina lobata Gnaspini, Moraes &amp; Gomyde, new species, male paratype in MZSP.Apex of aedeagus in dorsal (A), left lateral (B), frontal (C), and ventral (D) views. do = dorsal opening; fl = flagellum; ll = left lobe; par = paralobe of the right lobe; rl = right lobe; tb = tubercles on the internal margin of the ventral opening." pageId="5" pageNumber="134">Fig. 6AC</figureCitation>
, par); left lobe with acute apex in lateral view (
<figureCitation id="131E2A263B110877FB56B06BFA92FC74" box="[1204,1290,871,897]" captionStart="FIGURE 6" captionStartId="4.[152,255,1844,1868]" captionTargetBox="[151,1436,1409,1820]" captionTargetId="figure-57@4.[151,1436,1409,1820]" captionTargetPageId="4" captionText="FIGURE 6. Parapaulipalpina lobata Gnaspini, Moraes &amp; Gomyde, new species, male paratype in MZSP.Apex of aedeagus in dorsal (A), left lateral (B), frontal (C), and ventral (D) views. do = dorsal opening; fl = flagellum; ll = left lobe; par = paralobe of the right lobe; rl = right lobe; tb = tubercles on the internal margin of the ventral opening." pageId="5" pageNumber="134">Fig. 6B</figureCitation>
, ll). Lateral margins of the ventral opening with a series of tubercles which might represent sensory structures (
<figureCitation id="131E2A263B110877FB1DB087FAC0FC50" box="[1279,1368,907,933]" captionStart="FIGURE 6" captionStartId="4.[152,255,1844,1868]" captionTargetBox="[151,1436,1409,1820]" captionTargetId="figure-57@4.[151,1436,1409,1820]" captionTargetPageId="4" captionText="FIGURE 6. Parapaulipalpina lobata Gnaspini, Moraes &amp; Gomyde, new species, male paratype in MZSP.Apex of aedeagus in dorsal (A), left lateral (B), frontal (C), and ventral (D) views. do = dorsal opening; fl = flagellum; ll = left lobe; par = paralobe of the right lobe; rl = right lobe; tb = tubercles on the internal margin of the ventral opening." pageId="5" pageNumber="134">Fig. 6D</figureCitation>
, tb). Ventral face of aedeagus with a series of subparallel ridges-and-sulci (
<figureCitation id="131E2A263B110877FC4EB0A3FC69FC3C" box="[940,1009,942,969]" captionStart="FIGURE 5" captionStartId="4.[152,255,1324,1348]" captionTargetBox="[151,1436,181,1296]" captionTargetId="figure-28@4.[151,1436,181,1301]" captionTargetPageId="4" captionText="FIGURE 5. Parapaulipalpina lobata Gnaspini, Moraes &amp; Gomyde, new species, male paratype in MZSP.A rotational view of the aedeagus, around its longitudinal axis." pageId="5" pageNumber="134">Fig. 5</figureCitation>
). Flagellum (of the endophallus) thin, sinuate, shorter (about 2/
<quantity id="4CDD9B463B110877FE4EB0DFFE4DFC18" box="[428,469,979,1005]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="5" pageNumber="134" unit="in" value="3.0">3 in</quantity>
length) than aedeagus (
<figureCitation id="131E2A263B110877FD39B0DFFCA4FC18" box="[731,828,979,1005]" captionStart="FIGURE 4" captionStartId="3.[152,255,1940,1964]" captionTargetBox="[296,1291,1309,1917]" captionTargetId="figure-131@3.[296,1291,1309,1917]" captionTargetPageId="3" captionText="FIGURE 4. Parapaulipalpina lobata Gnaspini, Moraes &amp; Gomyde, new species. A, male holotype genital segment; B, C, male holotype aedeagus, ventral (B) and right lateral (C) views; D, female spermatheca. All figures to the same scale. fl = flagellum." pageId="5" pageNumber="134">Figs. 4B</figureCitation>
,
<figureCitation id="131E2A263B110877FCA5B0DFFCF4FC18" box="[839,876,979,1005]" captionStart="FIGURE 6" captionStartId="4.[152,255,1844,1868]" captionTargetBox="[151,1436,1409,1820]" captionTargetId="figure-57@4.[151,1436,1409,1820]" captionTargetPageId="4" captionText="FIGURE 6. Parapaulipalpina lobata Gnaspini, Moraes &amp; Gomyde, new species, male paratype in MZSP.Apex of aedeagus in dorsal (A), left lateral (B), frontal (C), and ventral (D) views. do = dorsal opening; fl = flagellum; ll = left lobe; par = paralobe of the right lobe; rl = right lobe; tb = tubercles on the internal margin of the ventral opening." pageId="5" pageNumber="134">6D</figureCitation>
, fl). Parameres present, flat and thin, fused to the body of the aedegus except for their apical portion (
<figureCitation id="131E2A263B110877FD36B0FBFCC9FBE4" box="[724,849,1014,1041]" captionStart="FIGURE 5" captionStartId="4.[152,255,1324,1348]" captionTargetBox="[151,1436,181,1296]" captionTargetId="figure-28@4.[151,1436,181,1301]" captionTargetPageId="4" captionText="FIGURE 5. Parapaulipalpina lobata Gnaspini, Moraes &amp; Gomyde, new species, male paratype in MZSP.A rotational view of the aedeagus, around its longitudinal axis." pageId="5" pageNumber="134">Figs. 5E, L</figureCitation>
) [Note: the aedeagus is covered with debris, and it is not possible to observe the number of setae on the apex of the parameres and apex of aedeagal lobes].
</paragraph>
<paragraph id="8B9A36A33B110877FF25B733FE79FB88" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="134">
Female description. Same as male, except for the slender protarsus, and non-expanded metatibia. Spermatheca coiled with 2-turns (
<figureCitation id="131E2A263B110877FE98B76FFE4DFB88" box="[378,469,1123,1149]" captionStart="FIGURE 4" captionStartId="3.[152,255,1940,1964]" captionTargetBox="[296,1291,1309,1917]" captionTargetId="figure-131@3.[296,1291,1309,1917]" captionTargetPageId="3" captionText="FIGURE 4. Parapaulipalpina lobata Gnaspini, Moraes &amp; Gomyde, new species. A, male holotype genital segment; B, C, male holotype aedeagus, ventral (B) and right lateral (C) views; D, female spermatheca. All figures to the same scale. fl = flagellum." pageId="5" pageNumber="134">Fig. 4D</figureCitation>
).
</paragraph>
<paragraph id="8B9A36A33B110877FF25B78BFE17FB30" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="134">
<emphasis id="B951EAB13B110877FF25B78BFEA7FB54" box="[199,319,1159,1185]" italics="true" pageId="5" pageNumber="134">Etymology</emphasis>
. The name is given in reference to the wide and flat lateral projection at the apex of the metatibiae (from Latin, “lobed”).
</paragraph>
<paragraph id="8B9A36A33B110877FF25B7C3FDB2FB1C" blockId="5.[151,1437,150,2013]" box="[199,554,1231,1257]" pageId="5" pageNumber="134">
<emphasis id="B951EAB13B110877FF25B7C3FEC0FB1C" box="[199,344,1231,1257]" italics="true" pageId="5" pageNumber="134">Distribution.</emphasis>
<collectingCountry id="F33276333B110877FEBCB7C3FE3FFB1C" box="[350,423,1231,1257]" name="Brazil" pageId="5" pageNumber="134">Brazil</collectingCountry>
:
<collectingRegion id="49E1F8413B110877FE53B7C3FDBEFB1C" box="[433,550,1231,1257]" country="Brazil" name="Para" pageId="5" pageNumber="134">Pará State</collectingRegion>
.
</paragraph>
<paragraph id="8B9A36A33B110877FF25B7FFFCDFF980" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="134">
<emphasis id="B951EAB13B110877FF25B7FFFE2AFAF8" box="[199,434,1267,1293]" italics="true" pageId="5" pageNumber="134">Taxonomic Remarks.</emphasis>
The genus
<taxonomicName id="4C254D203B110877FDD0B7FFFD61FAF8" authorityName="Gnaspini" authorityYear="1996" box="[562,761,1267,1293]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="B951EAB13B110877FDD0B7FFFD61FAF8" box="[562,761,1267,1293]" italics="true" pageId="5" pageNumber="134">Parapaulipalpina</emphasis>
</taxonomicName>
previously included only four species, restricted to northern South America (see
<bibRefCitation id="EFB44B523B110877FE62B61BFDDAFAC4" author="Peck, S. B. &amp; Gnaspini, P. &amp; Newton, A. F." box="[384,578,1303,1329]" pageId="5" pageNumber="134" pagination="1 - 114" refId="ref5287" refString="Peck, S. B., Gnaspini, P. &amp; Newton, A. F. (2020) Updated catalog and generic keys of the Leiodidae (Insecta: Coleoptera) of the Neotropical region (&quot; Latin America &quot;: Mexico, the West Indies, and Central and South America). Zootaxa, 4741 (1), 1 - 114. https: // doi. org / 10.11646 / zootaxa. 4741.1.1" type="journal article" year="2020">
Peck
<emphasis id="B951EAB13B110877FE22B61BFE62FAC4" box="[448,506,1303,1329]" italics="true" pageId="5" pageNumber="134">et al.</emphasis>
, 2020
</bibRefCitation>
and
<figureCitation id="131E2A263B110877FD9FB61BFD58FAC4" box="[637,704,1303,1329]" captionStart="FIGURE 7" captionStartId="6.[152,255,867,891]" captionTargetBox="[471,1117,345,844]" captionTargetId="figure-111@6.[471,1117,345,844]" captionTargetPageId="6" captionText="FIGURE 7. Distribution map of the species of Parapaulipalpina (plotted on Google Maps map—available at https://www. google.com/maps). D = P. dentata Gnaspini, 1996; F = P. filicornis (Jeannel, 1936) (the grey ellipse indicates that this is not a precise locality); G = P. giachinoi Salgado, 2005; L = P. lobata Gnaspini, Moraes &amp; Gomyde, sp. nov.; T = P. tambopata (Salgado, 2013)." pageId="5" pageNumber="134">Fig. 7</figureCitation>
)—
<taxonomicName id="4C254D203B110877FD04B61BFB96FAC4" authority="Gnaspini, 1996" authorityName="Gnaspini" authorityYear="1996" box="[742,1038,1303,1329]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="134" phylum="Arthropoda" rank="species" species="dentata">
<emphasis id="B951EAB13B110877FD04B61BFCC0FAC4" box="[742,856,1303,1329]" italics="true" pageId="5" pageNumber="134">P. dentata</emphasis>
<bibRefCitation id="EFB44B523B110877FC83B61BFB96FAC4" author="Gnaspini, P." box="[865,1038,1303,1329]" pageId="5" pageNumber="134" pagination="509 - 556" refId="ref4390" refString="Gnaspini, P. (1996) Phylogenetic analysis of the tribe Ptomaphagini, with description of new Neotropical genera and species (Coleoptera, Leiodidae, Cholevinae, Ptomaphagini). Papeis Avulsos de Zoologia, 39, 509 - 556, errata. https: // doi. org / 10.11606 / 0031 - 1049.1994.39. p 509 - 556" type="journal article" year="1996">Gnaspini, 1996</bibRefCitation>
</taxonomicName>
(
<typeStatus id="549E88013B110877FBFDB61BFBC9FAC4" box="[1055,1105,1303,1329]" pageId="5" pageNumber="134">type</typeStatus>
species), from
<collectingCountry id="F33276333B110877FAE6B61BFAE3FAC4" box="[1284,1403,1303,1329]" name="Venezuela" pageId="5" pageNumber="134">Venezuela</collectingCountry>
;
<taxonomicName id="4C254D203B110877FA6AB61BFE2AFAA0" authority="(Jeannel, 1936)" baseAuthorityName="Jeannel" baseAuthorityYear="1936" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="134" phylum="Arthropoda" rank="species" species="filicornis">
<emphasis id="B951EAB13B110877FA6AB61BFF64FAA0" italics="true" pageId="5" pageNumber="134">P. filicornis</emphasis>
(
<bibRefCitation id="EFB44B523B110877FEEEB637FE32FAA0" author="Jeannel, R." box="[268,426,1339,1365]" pageId="5" pageNumber="134" pagination="1 - 433" refId="ref4903" refString="Jeannel, R. (1936) Monographie des Catopidae. Memoires du Museum National d'Histoire Naturelle, New Series, 1, 1 - 433." type="journal article" year="1936">Jeannel, 1936</bibRefCitation>
)
</taxonomicName>
(transferred from
<taxonomicName id="4C254D203B110877FD66B637FD69FAA0" authorityName="Portevin" authorityYear="1907" box="[644,753,1339,1365]" class="Insecta" family="Leiodidae" genus="Adelopsis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="B951EAB13B110877FD66B637FD69FAA0" box="[644,753,1339,1365]" italics="true" pageId="5" pageNumber="134">Adelopsis</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFB44B523B110877FCFFB637FC52FAA0" author="Gnaspini, P." box="[797,970,1339,1365]" pageId="5" pageNumber="134" pagination="509 - 556" refId="ref4390" refString="Gnaspini, P. (1996) Phylogenetic analysis of the tribe Ptomaphagini, with description of new Neotropical genera and species (Coleoptera, Leiodidae, Cholevinae, Ptomaphagini). Papeis Avulsos de Zoologia, 39, 509 - 556, errata. https: // doi. org / 10.11606 / 0031 - 1049.1994.39. p 509 - 556" type="journal article" year="1996">Gnaspini, 1996</bibRefCitation>
based solely on fig.
<quantity id="4CDD9B463B110877FB5AB636FB6AFAA0" box="[1208,1266,1338,1365]" metricMagnitude="0" metricUnit="m" metricValue="2.159" pageId="5" pageNumber="134" unit="in" value="85.0">85 in</quantity>
<bibRefCitation id="EFB44B523B110877FB18B637FA00FAA0" author="Jeannel, R." box="[1274,1432,1339,1365]" pageId="5" pageNumber="134" pagination="1 - 433" refId="ref4903" refString="Jeannel, R. (1936) Monographie des Catopidae. Memoires du Museum National d'Histoire Naturelle, New Series, 1, 1 - 433." type="journal article" year="1936">Jeannel, 1936</bibRefCitation>
, because the male genitalia was not available for study at that time, and the external features were redescribed in
<bibRefCitation id="EFB44B523B110877FF75B68FFE03FA68" author="Gnaspini, P. &amp; Peck, S. B." box="[151,411,1411,1437]" pageId="5" pageNumber="134" pagination="1 - 62" refId="ref4599" refString="Gnaspini, P. &amp; Peck, S. B. (2019) Redescription of the ' older Adelopsis ' species (Coleoptera: Leiodidae: Cholevinae: Ptomaphagini) based on the analysis of type specimens. Zootaxa, 4696 (1), 1 - 62. https: // doi. org / 10.11646 / zootaxa. 4696.1.1" type="journal article" year="2019">Gnaspini &amp; Peck, 2019</bibRefCitation>
—therefore, a comparison involving aedeagal morphology can not be fully done presently), from
<collectingCountry id="F33276333B110877FF34B6ABFEDCFA34" box="[214,324,1447,1473]" name="Colombia" pageId="5" pageNumber="134">Colombia</collectingCountry>
(no precise locality);
<taxonomicName id="4C254D203B110877FDA0B6ABFCEEFA34" authority="Salgado, 2005" authorityName="Salgado" authorityYear="2005" box="[578,886,1446,1473]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="134" phylum="Arthropoda" rank="species" species="giachinoi">
<emphasis id="B951EAB13B110877FDA0B6ABFD51FA34" box="[578,713,1447,1473]" italics="true" pageId="5" pageNumber="134">P. giachinoi</emphasis>
<bibRefCitation id="EFB44B523B110877FD33B6ABFCEEFA34" author="Salgado" box="[721,886,1446,1473]" pageId="5" pageNumber="134" pagination="963 - 982" refId="ref5398" refString="Salgado *, J. M. (2005) New species of Leiodidae (Coleoptera) and new records from the Neotropical Region. Revue Suisse de Zoologie, 112, 963 - 982. https: // doi. org / 10.5962 / bhl. part. 80333" type="journal article" year="2005">Salgado, 2005</bibRefCitation>
</taxonomicName>
, from
<collectingCountry id="F33276333B110877FC20B6ABFC6DFA34" box="[962,1013,1447,1473]" name="Peru" pageId="5" pageNumber="134">Peru</collectingCountry>
(and additional record in
<collectingCountry id="F33276333B110877FAC2B6ABFAE5FA34" box="[1312,1405,1447,1473]" name="Ecuador" pageId="5" pageNumber="134">Ecuador</collectingCountry>
in
<bibRefCitation id="EFB44B523B110877FF75B6C7FEA7FA10" author="Salgado" box="[151,319,1483,1509]" pageId="5" pageNumber="134" pagination="205 - 229" refId="ref5509" refString="Salgado *, J. M. (2010) Descripcion de una nueva especie del genero Paulipalpina y de tres nuevas species del genero Adelopsis de Ecuador (Coleoptera, Leiodidae, Cholevinae). Nouvelle Revue d'Entomologie, New Series, 26, 205 - 229." type="journal article" year="2010">Salgado, 2010</bibRefCitation>
); and
<taxonomicName id="4C254D203B110877FE65B6C7FD45FA10" authority="(Salgado, 2013)" baseAuthorityName="Salgado" baseAuthorityYear="2013" box="[391,733,1483,1509]" class="Insecta" family="Leiodidae" genus="Excelsiorella" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="134" phylum="Arthropoda" rank="species" species="tambopata">
<emphasis id="B951EAB13B110877FE65B6C7FD86FA10" box="[391,542,1483,1509]" italics="true" pageId="5" pageNumber="134">P. tambopata</emphasis>
(
<bibRefCitation id="EFB44B523B110877FDCDB6C7FD4DFA10" author="Salgado" box="[559,725,1483,1509]" pageId="5" pageNumber="134" pagination="79 - 92" refId="ref5558" refString="Salgado *, J. M. (2013) Cholevinae de Peru (Coleoptera, Leiodidae): especies nuevas y nuevos datos. Boletin de la Sociedad Entomologica Aragonesa, 52, 79 - 92." type="journal article" year="2013">Salgado, 2013</bibRefCitation>
)
</taxonomicName>
(transferred from
<taxonomicName id="4C254D203B110877FC51B6C7FB24FA10" authorityName="Salgado" authorityYear="2013" box="[947,1212,1483,1509]" class="Insecta" family="Leiodidae" genus="Excelsiorella" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="134" phylum="Arthropoda" rank="subGenus" subGenus="Viruana">
<emphasis id="B951EAB13B110877FC51B6C7FBDEFA10" box="[947,1094,1483,1509]" italics="true" pageId="5" pageNumber="134">Excelsiorella</emphasis>
(
<emphasis id="B951EAB13B110877FBBBB6C7FB2BFA11" box="[1113,1203,1483,1508]" italics="true" pageId="5" pageNumber="134">Viruana</emphasis>
)
</taxonomicName>
by
<bibRefCitation id="EFB44B523B110877FB0EB6C7FF57F9FC" author="Gnaspini, P. &amp; Antunes-Carvalho, C. &amp; Peck, S. B." pageId="5" pageNumber="134" pagination="272 - 274" refId="ref4539" refString="Gnaspini, P., Antunes-Carvalho, C. &amp; Peck, S. B. (2016) Two new synonymies at the generic level in Neotropical Ptomaphagini (Coleoptera, Leiodidae, Cholevinae). Zootaxa, 4132 (2), 272 - 274. https: // doi. org / 10.11646 / zootaxa. 4132.2.7" type="journal article" year="2016">
Gnaspini
<emphasis id="B951EAB13B110877FAB8B6C7FA0EFA10" box="[1370,1430,1483,1509]" italics="true" pageId="5" pageNumber="134">et al.</emphasis>
, 2016
</bibRefCitation>
based on the drawings and discussion in the original description), from
<collectingCountry id="F33276333B110877FC1BB6E3FBA8F9FC" box="[1017,1072,1519,1545]" name="Peru" pageId="5" pageNumber="134">Peru</collectingCountry>
. Recently,
<bibRefCitation id="EFB44B523B110877FB49B6E3FA04F9FC" author="Gomyde, E. C. &amp; de Medeiros, B. A. S. &amp; Gnaspini, P." box="[1195,1436,1519,1545]" pageId="5" pageNumber="134" refId="ref4766" refString="Gomyde, E. C., de Medeiros, B. A. S. &amp; Gnaspini, P. (2024) The paralobe: a new diagnostic and synapomorphic character for the genera Paulipalpina Gnaspini &amp; Peck, 1996 and Parapaulipalpina Gnaspini, 1996 (Leiodidae: Cholevinae: Ptomaphagini). Journal of Morphology, 285, e 21693. https: // doi. org / 10.1002 / jmor. 21693" type="journal volume" year="2024">
Gomyde
<emphasis id="B951EAB13B110877FAF0B6E3FAD3F9FC" box="[1298,1355,1519,1545]" italics="true" pageId="5" pageNumber="134">et al.</emphasis>
(2024)
</bibRefCitation>
described a new feature (the paralobe of the aedeagus), considered to be synapomorphic for
<taxonomicName id="4C254D203B110877FB47B51FFAF4F9D8" authorityName="Gnaspini" authorityYear="1996" box="[1189,1388,1555,1581]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="B951EAB13B110877FB47B51FFAF4F9D8" box="[1189,1388,1555,1581]" italics="true" pageId="5" pageNumber="134">Parapaulipalpina</emphasis>
</taxonomicName>
and
<taxonomicName id="4C254D203B110877FF75B53BFDA0F9A4" authority="Gnaspini &amp; Peck, 1996" authorityName="Gnaspini &amp; Peck" authorityYear="1996" box="[151,568,1591,1617]" class="Insecta" family="Leiodidae" genus="Paulipalpina" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="B951EAB13B110877FF75B53BFEB1F9A4" box="[151,297,1591,1617]" italics="true" pageId="5" pageNumber="134">Paulipalpina</emphasis>
Gnaspini &amp; Peck, 1996
</taxonomicName>
, examining specimens of
<taxonomicName id="4C254D203B110877FCBBB53BFC50F9A4" authorityName="Gnaspini" authorityYear="1996" box="[857,968,1591,1617]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="134" phylum="Arthropoda" rank="species" species="dentata">
<emphasis id="B951EAB13B110877FCBBB53BFC50F9A4" box="[857,968,1591,1617]" italics="true" pageId="5" pageNumber="134">P. dentata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D203B110877FC34B53BFBF1F9A4" baseAuthorityName="Salgado" baseAuthorityYear="2013" box="[982,1129,1591,1617]" class="Insecta" family="Leiodidae" genus="Excelsiorella" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="134" phylum="Arthropoda" rank="species" species="tambopata">
<emphasis id="B951EAB13B110877FC34B53BFBF1F9A4" box="[982,1129,1591,1617]" italics="true" pageId="5" pageNumber="134">P. tambopata</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C254D203B110877FB44B53BFA91F9A4" authorityName="Gnaspini &amp; Moraes &amp; Gomyde" authorityYear="2024" box="[1190,1289,1591,1617]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="134" phylum="Arthropoda" rank="species" species="lobata" status="sp. nov.">
<emphasis id="B951EAB13B110877FB44B53BFA91F9A4" box="[1190,1289,1591,1617]" italics="true" pageId="5" pageNumber="134">P. lobata</emphasis>
</taxonomicName>
<emphasis id="B951EAB13B110877FAEDB53BFAF0F9A4" bold="true" box="[1295,1384,1591,1617]" pageId="5" pageNumber="134">
<taxonomicNameLabel id="A26257CA3B110877FAEDB53BFAF0F9A4" box="[1295,1384,1591,1617]" pageId="5" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(see their fig. 6), reinforcing their placement in
<taxonomicName id="4C254D203B110877FD9AB557FCA7F980" authorityName="Gnaspini" authorityYear="1996" box="[632,831,1627,1653]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="B951EAB13B110877FD9AB557FCA7F980" box="[632,831,1627,1653]" italics="true" pageId="5" pageNumber="134">Parapaulipalpina</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9A36A33B110874FF25B573FDE3FEB4" blockId="5.[151,1437,150,2013]" lastBlockId="6.[151,1437,151,321]" lastPageId="6" lastPageNumber="135" pageId="5" pageNumber="134">
<taxonomicName id="4C254D203B110877FF25B573FE46F96C" authorityName="Gnaspini &amp; Moraes &amp; Gomyde" authorityYear="2024" box="[199,478,1663,1689]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="134" phylum="Arthropoda" rank="species" species="lobata" status="sp. nov.">
<emphasis id="B951EAB13B110877FF25B573FE46F96C" box="[199,478,1663,1689]" italics="true" pageId="5" pageNumber="134">Parapaulipalpina lobata</emphasis>
</taxonomicName>
<emphasis id="B951EAB13B110877FE07B573FDA7F96C" bold="true" box="[485,575,1663,1689]" pageId="5" pageNumber="134">
<taxonomicNameLabel id="A26257CA3B110877FE07B573FDA7F96C" box="[485,575,1663,1689]" pageId="5" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is the first species recorded in
<collectingCountry id="F33276333B110877FC7FB573FC7CF96C" box="[925,996,1663,1689]" name="Brazil" pageId="5" pageNumber="134">Brazil</collectingCountry>
, enlarging the distribution of the genus (see
<figureCitation id="131E2A263B110877FF2EB5AFFE88F948" box="[204,272,1699,1725]" captionStart="FIGURE 7" captionStartId="6.[152,255,867,891]" captionTargetBox="[471,1117,345,844]" captionTargetId="figure-111@6.[471,1117,345,844]" captionTargetPageId="6" captionText="FIGURE 7. Distribution map of the species of Parapaulipalpina (plotted on Google Maps map—available at https://www. google.com/maps). D = P. dentata Gnaspini, 1996; F = P. filicornis (Jeannel, 1936) (the grey ellipse indicates that this is not a precise locality); G = P. giachinoi Salgado, 2005; L = P. lobata Gnaspini, Moraes &amp; Gomyde, sp. nov.; T = P. tambopata (Salgado, 2013)." pageId="5" pageNumber="134">Fig. 7</figureCitation>
) and the first one studied under SEM (because of that, many characters can not be presently discussed among all species). Unfortunately, the specimen examined with SEM was covered with debris, not allowing observation of details; because of the small number of specimens, which were fragile for handling, we were not able to produce additional images.
<taxonomicName id="4C254D203B110877FE13B403FC91F8DC" authorityName="Gnaspini &amp; Moraes &amp; Gomyde" authorityYear="2024" box="[497,777,1807,1833]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="134" phylum="Arthropoda" rank="species" species="lobata" status="sp. nov.">
<emphasis id="B951EAB13B110877FE13B403FC91F8DC" box="[497,777,1807,1833]" italics="true" pageId="5" pageNumber="134">Parapaulipalpina lobata</emphasis>
</taxonomicName>
<emphasis id="B951EAB13B110877FCF3B403FCF5F8DC" bold="true" box="[785,877,1807,1833]" pageId="5" pageNumber="134">
<taxonomicNameLabel id="A26257CA3B110877FCF3B403FCF5F8DC" box="[785,877,1807,1833]" pageId="5" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be promptly recognized by the widened and flattened apex of the male metatibia, which is unique (maybe in the tribe as a whole) and diagnostic for the species. It is the smallest species recorded so far in the genus. The aedeagus of
<taxonomicName id="4C254D203B110877FC26B45BFBB1F884" authorityName="Gnaspini &amp; Moraes &amp; Gomyde" authorityYear="2024" box="[964,1065,1879,1905]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="134" phylum="Arthropoda" rank="species" species="lobata" status="sp. nov.">
<emphasis id="B951EAB13B110877FC26B45BFBB1F884" box="[964,1065,1879,1905]" italics="true" pageId="5" pageNumber="134">P. lobata</emphasis>
</taxonomicName>
<emphasis id="B951EAB13B110877FBD3B45BFB14F884" bold="true" box="[1073,1164,1879,1905]" pageId="5" pageNumber="134">
<taxonomicNameLabel id="A26257CA3B110877FBD3B45BFB14F884" box="[1073,1164,1879,1905]" pageId="5" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is concave ventrally, as in the other species of the genus, but, differently from them, there is a strong convex curvature near the apex. The flagellum of the aedeagus seems to be less elongate than the pattern described for the genus, being longer than the aedeagus in the other species known so far. However, the flagellum is connected to a strong basal piece, as it occurs in the other known species (based either on the descriptions and/or figures)—the aedeagus of
<taxonomicName id="4C254D203B120874FB4DB39BFAB7FF44" baseAuthorityName="Jeannel" baseAuthorityYear="1936" box="[1199,1327,151,177]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="species" species="filicornis">
<emphasis id="B951EAB13B120874FB4DB39BFAB7FF44" box="[1199,1327,151,177]" italics="true" pageId="6" pageNumber="135">P. filicornis</emphasis>
</taxonomicName>
is known only for the drawing in
<bibRefCitation id="EFB44B523B120874FE4FB3B7FDC0FF20" author="Jeannel, R." box="[429,600,187,213]" pageId="6" pageNumber="135" pagination="1 - 433" refId="ref4903" refString="Jeannel, R. (1936) Monographie des Catopidae. Memoires du Museum National d'Histoire Naturelle, New Series, 1, 1 - 433." type="journal article" year="1936">Jeannel (1936)</bibRefCitation>
, which does not allow a proper recognition of this feature; however, the drawing does show a seemingly unique feature of that species, which is an expansion near the apex of the flagellum.
<taxonomicName id="4C254D203B120874FF75B20FFE35FEE8" authorityName="Gnaspini &amp; Moraes &amp; Gomyde" authorityYear="2024" box="[151,429,259,285]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="species" species="lobata" status="sp. nov.">
<emphasis id="B951EAB13B120874FF75B20FFE35FEE8" box="[151,429,259,285]" italics="true" pageId="6" pageNumber="135">Parapaulipalpina lobata</emphasis>
</taxonomicName>
<emphasis id="B951EAB13B120874FE51B20FFD93FEE8" bold="true" box="[435,523,259,285]" pageId="6" pageNumber="135">
<taxonomicNameLabel id="A26257CA3B120874FE51B20FFD93FEE8" box="[435,523,259,285]" pageId="6" pageNumber="135" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
seems to bear a slightly reduced eye, but not as strongly reduced as in
<taxonomicName id="4C254D203B120874FAFEB20FFA04FEE8" baseAuthorityName="Jeannel" baseAuthorityYear="1936" box="[1308,1436,259,285]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="species" species="filicornis">
<emphasis id="B951EAB13B120874FAFEB20FFA04FEE8" box="[1308,1436,259,285]" italics="true" pageId="6" pageNumber="135">P. filicornis</emphasis>
</taxonomicName>
(another diagnostic feature of that species).
</paragraph>
<caption id="DF5A662B3B120874FF7AB06FFED8FC1D" pageId="6" pageNumber="135" startId="6.[152,255,867,891]" targetBox="[471,1117,345,844]" targetPageId="6" targetType="figure">
<paragraph id="8B9A36A33B120874FF7AB06FFED8FC1D" blockId="6.[151,1437,867,1000]" pageId="6" pageNumber="135">
<emphasis id="B951EAB13B120874FF7AB06FFE81FC89" bold="true" box="[152,281,867,892]" pageId="6" pageNumber="135">FIGURE 7.</emphasis>
Distribution map of the species of
<taxonomicName id="4C254D203B120874FD64B068FCA2FC89" authorityName="Gnaspini" authorityYear="1996" box="[646,826,868,892]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis id="B951EAB13B120874FD64B068FCA2FC89" box="[646,826,868,892]" italics="true" pageId="6" pageNumber="135">Parapaulipalpina</emphasis>
</taxonomicName>
(plotted on Google Maps map—available at https://www. google.com/maps). D =
<taxonomicName id="4C254D203B120874FE73B084FD07FC55" authority="Gnaspini, 1996" authorityName="Gnaspini" authorityYear="1996" box="[401,671,904,928]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="species" species="dentata">
<emphasis id="B951EAB13B120874FE73B084FE60FC55" box="[401,504,904,928]" italics="true" pageId="6" pageNumber="135">P. dentata</emphasis>
<bibRefCitation id="EFB44B523B120874FE1DB084FD07FC55" author="Gnaspini, P." box="[511,671,904,928]" pageId="6" pageNumber="135" pagination="509 - 556" refId="ref4390" refString="Gnaspini, P. (1996) Phylogenetic analysis of the tribe Ptomaphagini, with description of new Neotropical genera and species (Coleoptera, Leiodidae, Cholevinae, Ptomaphagini). Papeis Avulsos de Zoologia, 39, 509 - 556, errata. https: // doi. org / 10.11606 / 0031 - 1049.1994.39. p 509 - 556" type="journal article" year="1996">Gnaspini, 1996</bibRefCitation>
</taxonomicName>
; F =
<taxonomicName id="4C254D203B120874FD34B084FC69FC55" authority="(Jeannel, 1936)" baseAuthorityName="Jeannel" baseAuthorityYear="1936" box="[726,1009,904,928]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="species" species="filicornis">
<emphasis id="B951EAB13B120874FD34B084FCD5FC55" box="[726,845,904,928]" italics="true" pageId="6" pageNumber="135">P. filicornis</emphasis>
(
<bibRefCitation id="EFB44B523B120874FCB9B084FC72FC55" author="Jeannel, R." box="[859,1002,904,928]" pageId="6" pageNumber="135" pagination="1 - 433" refId="ref4903" refString="Jeannel, R. (1936) Monographie des Catopidae. Memoires du Museum National d'Histoire Naturelle, New Series, 1, 1 - 433." type="journal article" year="1936">Jeannel, 1936</bibRefCitation>
)
</taxonomicName>
(the grey ellipse indicates that this is not a precise locality); G =
<taxonomicName id="4C254D203B120874FE6DB0A0FD30FC36" authority="Salgado, 2005" authorityName="Salgado" authorityYear="2005" box="[399,680,939,964]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="species" species="giachinoi">
<emphasis id="B951EAB13B120874FE6DB0A0FD92FC31" box="[399,522,940,964]" italics="true" pageId="6" pageNumber="135">P. giachinoi</emphasis>
<bibRefCitation id="EFB44B523B120874FDF0B0A0FD30FC36" author="Salgado" box="[530,680,939,964]" pageId="6" pageNumber="135" pagination="963 - 982" refId="ref5398" refString="Salgado *, J. M. (2005) New species of Leiodidae (Coleoptera) and new records from the Neotropical Region. Revue Suisse de Zoologie, 112, 963 - 982. https: // doi. org / 10.5962 / bhl. part. 80333" type="journal article" year="2005">Salgado, 2005</bibRefCitation>
</taxonomicName>
; L =
<taxonomicName id="4C254D203B120874FD00B0A0FBE2FC31" authority="Gnaspini, Moraes &amp; Gomyde" authorityName="Gnaspini, Moraes &amp; Gomyde" authorityYear="2024" box="[738,1146,940,964]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="species" species="lobata" status="sp. nov.">
<emphasis id="B951EAB13B120874FD00B0A0FCA5FC31" box="[738,829,940,964]" italics="true" pageId="6" pageNumber="135">P. lobata</emphasis>
Gnaspini, Moraes &amp; Gomyde
</taxonomicName>
,
<emphasis id="B951EAB13B120874FB64B0A0FB40FC31" bold="true" box="[1158,1240,940,964]" pageId="6" pageNumber="135">
<taxonomicNameLabel id="A26257CA3B120874FB64B0A0FB40FC31" box="[1158,1240,940,964]" pageId="6" pageNumber="135" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; T =
<taxonomicName id="4C254D203B120874FAF7B0A0FEA4FC1D" authority="(Salgado, 2013)" baseAuthorityName="Salgado" baseAuthorityYear="2013" class="Insecta" family="Leiodidae" genus="Excelsiorella" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="species" species="tambopata">
<emphasis id="B951EAB13B120874FAF7B0A0FA04FC31" box="[1301,1436,940,964]" italics="true" pageId="6" pageNumber="135">P. tambopata</emphasis>
(
<bibRefCitation id="EFB44B523B120874FF7CB0DCFEADFC1D" author="Salgado" box="[158,309,976,1000]" pageId="6" pageNumber="135" pagination="79 - 92" refId="ref5558" refString="Salgado *, J. M. (2013) Cholevinae de Peru (Coleoptera, Leiodidae): especies nuevas y nuevos datos. Boletin de la Sociedad Entomologica Aragonesa, 52, 79 - 92." type="journal article" year="2013">Salgado, 2013</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8B9A36A33B120874FF25B736FCB3FB35" blockId="6.[151,1437,1082,1360]" pageId="6" pageNumber="135">
<taxonomicName id="4C254D203B120874FF25B736FE76FBA1" authorityName="Gnaspini" authorityYear="1996" box="[199,494,1082,1108]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="species" species="dentata">
<emphasis id="B951EAB13B120874FF25B736FE76FBA1" box="[199,494,1082,1108]" italics="true" pageId="6" pageNumber="135">Parapaulipalpina dentata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C254D203B120874FDCDB737FD22FBA1" authorityName="Salgado" authorityYear="2005" box="[559,698,1082,1108]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="species" species="giachinoi">
<emphasis id="B951EAB13B120874FDCDB737FD22FBA1" box="[559,698,1082,1108]" italics="true" pageId="6" pageNumber="135">P. giachinoi</emphasis>
</taxonomicName>
have a tubercle medially on the posterior margin of the male metafemur, which was not observed in
<taxonomicName id="4C254D203B120874FDADB753FD2BFB8D" authorityName="Gnaspini &amp; Moraes &amp; Gomyde" authorityYear="2024" box="[591,691,1118,1144]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="species" species="lobata" status="sp. nov.">
<emphasis id="B951EAB13B120874FDADB753FD2BFB8D" box="[591,691,1118,1144]" italics="true" pageId="6" pageNumber="135">P. lobata</emphasis>
</taxonomicName>
<emphasis id="B951EAB13B120874FD5BB752FC8BFB8D" bold="true" box="[697,787,1118,1144]" pageId="6" pageNumber="135">
<taxonomicNameLabel id="A26257CA3B120874FD5BB752FC8BFB8D" box="[697,787,1118,1144]" pageId="6" pageNumber="135" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and not studied in the other species, but their aedeagi are different.
<taxonomicName id="4C254D203B120874FEE5B78EFDD6FB69" baseAuthorityName="Salgado" baseAuthorityYear="2013" box="[263,590,1154,1180]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="species" species="tambopata">
<emphasis id="B951EAB13B120874FEE5B78EFDD6FB69" box="[263,590,1154,1180]" italics="true" pageId="6" pageNumber="135">Parapaulipalpina tambopata</emphasis>
</taxonomicName>
can be diagnostically recognized by the truncate elytra and non-expanded male protarsomeres (according to the original description).
</paragraph>
<paragraph id="8B9A36A33B120874FF25B7C6FC34FAA5" blockId="6.[151,1437,1082,1360]" pageId="6" pageNumber="135">
<taxonomicName id="4C254D203B120874FF25B7C6FE64FB11" baseAuthorityName="Jeannel" baseAuthorityYear="1936" box="[199,508,1226,1252]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="species" species="filicornis">
<emphasis id="B951EAB13B120874FF25B7C6FE64FB11" box="[199,508,1226,1252]" italics="true" pageId="6" pageNumber="135">Parapaulipalpina filicornis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D203B120874FDE9B7C7FD38FB11" baseAuthorityName="Salgado" baseAuthorityYear="2013" box="[523,672,1226,1252]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="species" species="tambopata">
<emphasis id="B951EAB13B120874FDE9B7C7FD38FB11" box="[523,672,1226,1252]" italics="true" pageId="6" pageNumber="135">P. tambopata</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C254D203B120874FD02B7C7FCFEFB11" authorityName="Salgado" authorityYear="2005" box="[736,870,1226,1252]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="species" species="giachinoi">
<emphasis id="B951EAB13B120874FD02B7C7FCFEFB11" box="[736,870,1226,1252]" italics="true" pageId="6" pageNumber="135">P. giachinoi</emphasis>
</taxonomicName>
are known only from their male
<typeStatus id="549E88013B120874FB03B7C6FAD6FB11" box="[1249,1358,1226,1252]" pageId="6" pageNumber="135" type="holotype">holotypes</typeStatus>
(and a subsequently added male of the latter, from another country). Therefore, the spermatheca of the female is unkown for those species. The spermatheca of
<taxonomicName id="4C254D203B120874FDBAB61FFD26FAD9" authorityName="Gnaspini &amp; Moraes &amp; Gomyde" authorityYear="2024" box="[600,702,1298,1324]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="species" species="lobata" status="sp. nov.">
<emphasis id="B951EAB13B120874FDBAB61FFD26FAD9" box="[600,702,1298,1324]" italics="true" pageId="6" pageNumber="135">P. lobata</emphasis>
</taxonomicName>
<emphasis id="B951EAB13B120874FD2AB61EFCBDFAD9" bold="true" box="[712,805,1298,1324]" pageId="6" pageNumber="135">
<taxonomicNameLabel id="A26257CA3B120874FD2AB61EFCBDFAD9" box="[712,805,1298,1324]" pageId="6" pageNumber="135" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resembles that of
<taxonomicName id="4C254D203B120874FBE2B61FFBEBFAD9" authorityName="Gnaspini" authorityYear="1996" box="[1024,1139,1298,1324]" class="Insecta" family="Leiodidae" genus="Parapaulipalpina" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="species" species="dentata">
<emphasis id="B951EAB13B120874FBE2B61FFBEBFAD9" box="[1024,1139,1298,1324]" italics="true" pageId="6" pageNumber="135">P. dentata</emphasis>
</taxonomicName>
, by having a 2-turns coil followed by a somewhat long, curved duct ending with an apical bulb.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,214 @@
<document id="DF8B81D3B01B3F7422AF831AEF9E717D" ID-DOI="10.11646/zootaxa.5506.2.2" ID-ISSN="1175-5326" ID-ZooBank="03F52CEE-CECD-4FE5-93B9-8CC9F31FCCA4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726053331936" checkinUser="plazi" docAuthor="Tan, Ming Kai, Tumbrinck, Josef, Rivera, Rodrin R. &amp; Nuñeza, Olga Macas" docDate="2024" docId="03B3F80FA350FFA1FF46D5EFFA36328D" docLanguage="en" docName="zootaxa.5506.2.2.pdf" docOrigin="Zootaxa 5506 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5506.2.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trusmaditetrigini Storozhenko 2023" docType="key" docVersion="1" lastPageNumber="197" masterDocId="FF8A8077A353FFA2FFD1D578FFAA312C" masterDocTitle="A new genus and a new species of Cladonotinae (Orthoptera: Tetrigidae) from Mindanao, Philippines" masterLastPageNumber="204" masterPageNumber="194" pageNumber="197" updateTime="1726053447841" updateUser="GgImagineBatch" zenodo-license-document="CLOSED">
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<mods:title id="2152ED66FA71EC7066FA48E3C258255B">A new genus and a new species of Cladonotinae (Orthoptera: Tetrigidae) from Mindanao, Philippines</mods:title>
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<mods:namePart id="BC296B51BEB941235DFBA596BA8575FD">Tan, Ming Kai</mods:namePart>
<mods:affiliation id="926DCC19D6CC2B86A93686D3E840858E">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore</mods:affiliation>
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<mods:namePart id="98A915823C9A1D7BB6C7DF0054135801">Tumbrinck, Josef</mods:namePart>
<mods:affiliation id="12A6F078C01645248C11BC89EE8418BB">Auf der Hees 1, D- 41849 Wassenberg, Germany</mods:affiliation>
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<mods:namePart id="41A8C3B4C614D6F5223AAF8C060A1065">Rivera, Rodrin R.</mods:namePart>
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<mods:affiliation id="5B55C3779A89ACDE26CDFE35E5CEC467">Department of Biological Sciences, College of Science and Mathematics, Mindanao State University-Iligan Institute of Technology, Tibanga, Iligan City 9200, Philippines</mods:affiliation>
<mods:nameIdentifier id="FA94DA4EB22764D2F86FB18CFF372719" type="email">rodrin.rivera@nmsc.edu.ph</mods:nameIdentifier>
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<mods:namePart id="958255B8AD5D104A4ED92203881AE645">Nuñeza, Olga Macas</mods:namePart>
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<emphasis id="B96E950BA350FFA1FF46D5EFFC01319D" bold="true" box="[151,939,151,177]" pageId="3" pageNumber="197">
Key to genera of the tribe
<taxonomicName id="4C1A329AA350FFA1FE0FD5EFFC01319D" ID-CoL="BST3Z" authority="Storozhenko, 2023" authorityName="Storozhenko" authorityYear="2023" box="[478,939,151,177]" class="Insecta" family="Tetrigidae" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="197" phylum="Arthropoda" rank="tribe" tribe="Trusmaditetrigini">Trusmaditetrigini Storozhenko, 2023</taxonomicName>
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Parts of the key are modified from
<bibRefCitation id="EF8B34E8A350FFA1FDCED5A7FD5A31D5" author="Tumbrinck, J." box="[543,752,223,249]" pageId="3" pageNumber="197" pagination="350 - 396" refId="ref5999" refString="Tumbrinck, J. (2014) Taxonomic revision of the Cladonotinae (Orthoptera: Tetrigidae) from the islands of South-East Asia and from Australia, with general remarks to the classification and morphology of the Tetrigidae and descriptions of new genera and species from New Guinea and New Caledonia. In: Telnov, D. (Ed.), Biodiversity, Biogeography and Nature Conservation in Wallacea and New Guinea, II, pp. 350 - 396." type="book chapter" year="2014">Tumbrinck (2014)</bibRefCitation>
.
</paragraph>
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<subSubSection id="C3001A92A350FFA1FF46D45CFA36328D" pageId="3" pageNumber="197" type="key">
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<paragraph id="8BA54919A350FFA1FF46D45CFA363015" blockId="3.[151,1436,292,343]" box="[151,1436,292,314]" pageId="3" pageNumber="197">
<keyLead id="30EBEA2CA350FFA1FF46D45CFA363015" box="[151,1436,292,314]" pageId="3" pageNumber="197">
1. Distribution: East of Wallace Line: New
<collectingCountry id="F30D0989A350FFA1FDBBD45CFD183016" box="[618,690,292,314]" name="Guinea-Bissau" pageId="3" pageNumber="197">Guinea</collectingCountry>
,
<collectingCountry id="F30D0989A350FFA1FD6AD45DFCFA3016" box="[699,848,292,314]" name="New Caledonia" pageId="3" pageNumber="197">New Caledonia</collectingCountry>
and
<collectingCountry id="F30D0989A350FFA1FCACD45CFC7F3016" box="[893,981,292,314]" name="Australia" pageId="3" pageNumber="197">Australia</collectingCountry>
.................................... 2
</keyLead>
</paragraph>
<paragraph id="8BA54919A350FFA1FF46D43AFA36307B" blockId="3.[151,1436,292,343]" box="[151,1436,321,343]" pageId="3" pageNumber="197">
<keyLead id="30EBEA2CA350FFA1FF46D43AFA36307B" box="[151,1436,321,343]" pageId="3" pageNumber="197">
-. Distribution: West of Wallace Line: Borneo,
<collectingCountry id="F30D0989A350FFA1FD5ED439FD51307B" box="[655,763,321,343]" name="Philippines" pageId="3" pageNumber="197">Philippines</collectingCountry>
....................................................... 6
</keyLead>
</paragraph>
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<keyStep id="30EE51BCA350FFA1FF46D404FA363083" pageId="3" pageNumber="197">
<paragraph id="8BA54919A350FFA1FF46D404FA3630BD" blockId="3.[151,1436,380,431]" box="[151,1436,380,402]" pageId="3" pageNumber="197">
<keyLead id="30EBEA2CA350FFA1FF46D404FA3630BD" box="[151,1436,380,402]" pageId="3" pageNumber="197">2. Pronotum and body conspicuously flattened, hind femur more or less inclined with lateral curved lappets............... 3</keyLead>
</paragraph>
<paragraph id="8BA54919A350FFA1FF46D4E2FA363083" blockId="3.[151,1436,380,431]" box="[151,1436,409,431]" pageId="3" pageNumber="197">
<keyLead id="30EBEA2CA350FFA1FF46D4E2FA363083" box="[151,1436,409,431]" pageId="3" pageNumber="197">-. Pronotum and body not conspicuously flattened............................................................. 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EE51BCA350FFA1FF46D4ACFA36332A" pageId="3" pageNumber="197">
<paragraph id="8BA54919A350FFA1FF46D4ACFA3730C5" blockId="3.[151,1437,468,519]" box="[151,1437,468,490]" pageId="3" pageNumber="197">
<keyLead id="30EBEA2CA350FFA1FF46D4ACFA3730C5" box="[151,1437,468,490]" pageId="3" pageNumber="197">
3. Brachypronotal, pronotum length&gt;
<quantity id="4CE2E4FCA350FFA1FDEDD4ACFDDC30C5" box="[572,630,468,490]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="197" unit="mm" value="5.0">5 mm</quantity>
; internal lateral carina present. Distribution:
<collectingCountry id="F30D0989A350FFA1FBF5D4ACFB1230C6" box="[1060,1208,468,490]" name="New Caledonia" pageId="3" pageNumber="197">New Caledonia</collectingCountry>
.....
<taxonomicName id="4C1A329AA350FFA1FAD3D4ACFA3730C5" ID-CoL="8NRNZ" authorityName="Hancock" authorityYear="1907" box="[1282,1437,468,489]" class="Insecta" family="Tetrigidae" genus="Eurymorphopus" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B96E950BA350FFA1FAD3D4ACFA3730C5" box="[1282,1437,468,489]" italics="true" pageId="3" pageNumber="197">Eurymorphopus</emphasis>
</taxonomicName>
</keyLead>
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<paragraph id="8BA54919A350FFA1FF46D48AFA36332A" blockId="3.[151,1437,468,519]" box="[151,1436,497,519]" pageId="3" pageNumber="197">
<keyLead id="30EBEA2CA350FFA1FF46D48AFA36332A" box="[151,1436,497,519]" pageId="3" pageNumber="197">
-. Micropronotal, pronotum length &lt;
<quantity id="4CE2E4FCA350FFA1FDE0D489FDC1332B" box="[561,619,497,519]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="197" unit="mm" value="5.0">5 mm</quantity>
; internal lateral carina absent. Distribution: New
<collectingCountry id="F30D0989A350FFA1FB94D489FB20332B" box="[1093,1162,497,519]" name="Guinea-Bissau" pageId="3" pageNumber="197">Guinea</collectingCountry>
.............
<taxonomicName id="4C1A329AA350FFA1FAE2D489FA36332A" ID-CoL="8NXNH" authorityName="Tumbrinck" authorityYear="2014" box="[1331,1436,497,518]" class="Insecta" family="Tetrigidae" genus="Planotettix" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B96E950BA350FFA1FAE2D489FA36332A" box="[1331,1436,497,518]" italics="true" pageId="3" pageNumber="197">Planotettix</emphasis>
</taxonomicName>
</keyLead>
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<keyStep id="30EE51BCA350FFA1FF46D754FA363373" pageId="3" pageNumber="197">
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<keyLead id="30EBEA2CA350FFA1FF46D754FA36336D" box="[151,1436,556,578]" pageId="3" pageNumber="197">
4. Head exserted over the pronotum, with globular projecting eyes. Distribution: New
<collectingCountry id="F30D0989A350FFA1FBD7D754FBE1336E" box="[1030,1099,556,578]" name="Guinea-Bissau" pageId="3" pageNumber="197">Guinea</collectingCountry>
................
<taxonomicName id="4C1A329AA350FFA1FAC5D754FA36336D" ID-CoL="8NSTN" authorityName="Tumbrinck" authorityYear="2014" box="[1300,1436,556,577]" class="Insecta" family="Tetrigidae" genus="Ichikawatettix" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B96E950BA350FFA1FAC5D754FA36336D" box="[1300,1436,556,577]" italics="true" pageId="3" pageNumber="197">Ichikawatettix</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA54919A350FFA1FF46D732FA363373" blockId="3.[151,1436,556,607]" box="[151,1436,585,607]" pageId="3" pageNumber="197">
<keyLead id="30EBEA2CA350FFA1FF46D732FA363373" box="[151,1436,585,607]" pageId="3" pageNumber="197">-. Head not exserted. Eyes not conspicuously globular.......................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EE51BCA350FFA1FF46D7FCFA36339B" pageId="3" pageNumber="197">
<paragraph id="8BA54919A350FFA1FF46D7FCFA3633B6" blockId="3.[151,1436,644,695]" box="[151,1436,644,666]" pageId="3" pageNumber="197">
<keyLead id="30EBEA2CA350FFA1FF46D7FCFA3633B6" box="[151,1436,644,666]" pageId="3" pageNumber="197">
5. Median carina reaches the posterior margin of the pronotum. Distribution: New
<collectingCountry id="F30D0989A350FFA1FC32D7FCFB8233B6" box="[995,1064,644,666]" name="Guinea-Bissau" pageId="3" pageNumber="197">Guinea</collectingCountry>
...................
<taxonomicName id="4C1A329AA350FFA1FAC6D7FDFA3633B6" ID-CoL="8NQRH" authorityName="Tumbrinck" authorityYear="2014" box="[1303,1436,645,666]" class="Insecta" family="Tetrigidae" genus="Devriesetettix" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B96E950BA350FFA1FAC6D7FDFA3633B6" box="[1303,1436,645,666]" italics="true" pageId="3" pageNumber="197">Devriesetettix</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA54919A350FFA1FF46D7DAFA36339B" blockId="3.[151,1436,644,695]" box="[151,1436,673,695]" pageId="3" pageNumber="197">
<keyLead id="30EBEA2CA350FFA1FF46D7DAFA36339B" box="[151,1436,673,695]" pageId="3" pageNumber="197">
-. Median carina does not reach the posterior margin of the pronotum. Distribution:
<collectingCountry id="F30D0989A350FFA1FC20D7D9FBE3339B" box="[1009,1097,673,695]" name="Australia" pageId="3" pageNumber="197">Australia</collectingCountry>
..................
<taxonomicName id="4C1A329AA350FFA1FACED7DAFA36339B" ID-CoL="8NZXX" authorityName="Rehn" authorityYear="1952" box="[1311,1436,674,695]" class="Insecta" family="Tetrigidae" genus="Tepperotettix" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B96E950BA350FFA1FACED7DAFA36339B" box="[1311,1436,674,695]" italics="true" pageId="3" pageNumber="197">Tepperotettix</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EE51BCA350FFA1FF46D7A4FA363223" pageId="3" pageNumber="197">
<paragraph id="8BA54919A350FFA1FF46D7A4FA3633DD" blockId="3.[151,1436,732,783]" box="[151,1436,732,754]" pageId="3" pageNumber="197">
<keyLead id="30EBEA2CA350FFA1FF46D7A4FA3633DD" box="[151,1436,732,754]" pageId="3" pageNumber="197">
6. Pronotum more raised, more rugose and femora with carinae more undulated...................
<taxonomicName id="4C1A329AA350FFA1FB7DD7A5FA9633DE" authorityName="Tan &amp; Tumbrinck &amp; Rivera &amp; Nuñeza" authorityYear="2024" box="[1196,1340,733,754]" class="Insecta" family="Tetrigidae" genus="Fartmanntettix" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="197" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96E950BA350FFA1FB7DD7A5FA9633DE" box="[1196,1340,733,754]" italics="true" pageId="3" pageNumber="197">Fartmanntettix</emphasis>
</taxonomicName>
<emphasis id="B96E950BA350FFA1FA93D7A4FA3633DD" bold="true" box="[1346,1436,732,753]" pageId="3" pageNumber="197">
<taxonomicNameLabel id="A25D2870A350FFA1FA93D7A4FA3633DD" box="[1346,1436,732,753]" pageId="3" pageNumber="197" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BA54919A350FFA1FF46D782FA363223" blockId="3.[151,1436,732,783]" box="[151,1436,761,783]" pageId="3" pageNumber="197">
<keyLead id="30EBEA2CA350FFA1FF46D782FA363223" box="[151,1436,761,783]" pageId="3" pageNumber="197">-. Pronotum more flattened, less rugose and femora less undulated................................................ 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EE51BCA350FFA1FF46D64CFA36328D" pageId="3" pageNumber="197">
<paragraph id="8BA54919A350FFA1FF46D64CFA36324A" blockId="3.[151,1437,820,930]" pageId="3" pageNumber="197">
<keyLead id="30EBEA2CA350FFA1FF46D64CFA36324A" pageId="3" pageNumber="197">
7. Vertex wider than an eye. Pronotum with apex narrowly rounded. Winged. Fore femora distinctly elongated. Distribution: Borneo..................................................................................
<taxonomicName id="4C1A329AA350FFA1FADED629FA36324A" ID-CoL="BST9Y" authorityName="Storozhenko" authorityYear="2023" box="[1295,1436,849,870]" class="Insecta" family="Tetrigidae" genus="Trusmaditetrix" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B96E950BA350FFA1FADED629FA36324A" box="[1295,1436,849,870]" italics="true" pageId="3" pageNumber="197">Trusmaditetrix</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA54919A350FFA1FF46D617FA36328D" blockId="3.[151,1437,820,930]" pageId="3" pageNumber="197">
<keyLead id="30EBEA2CA350FFA1FF46D617FA36328D" pageId="3" pageNumber="197">
-. Vertex as wide as an eye. Pronotum with apex distinctly bilobate. Lack wings. Fore femora shorter. Distribution:
<collectingCountry id="F30D0989A350FFA1FAF1D617FA2632A9" box="[1312,1420,879,901]" name="Philippines" pageId="3" pageNumber="197">Philippines</collectingCountry>
.......................................................................................
<taxonomicName id="4C1A329AA350FFA1FB27D6F4FA36328D" ID-CoL="CF64T" authorityName="Patano, Skejo, Amoroso &amp; Kasalo" authorityYear="2024" box="[1270,1436,908,929]" class="Insecta" family="Tetrigidae" genus="Storozhenkotettix" higherTaxonomySource="CoL" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B96E950BA350FFA1FB27D6F4FA36328D" box="[1270,1436,908,929]" italics="true" pageId="3" pageNumber="197">Storozhenkotettix</emphasis>
</taxonomicName>
</keyLead>
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<mods:title id="250FCB0D67998A2B5022B77152B804EE">A new genus and a new species of Cladonotinae (Orthoptera: Tetrigidae) from Mindanao, Philippines</mods:title>
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<mods:namePart id="BD5B3AC439F4289348FCDF8A3181FB45">Rivera, Rodrin R.</mods:namePart>
<mods:nameIdentifier id="1ADD588D8E94080ADAB648C4199AEA23" type="ORCID">0000-0003-3019-0258</mods:nameIdentifier>
<mods:affiliation id="3D9649F1A4F63569FA013F4ABBB63551">Department of Biological Sciences, College of Science and Mathematics, Mindanao State University-Iligan Institute of Technology, Tibanga, Iligan City 9200, Philippines</mods:affiliation>
<mods:nameIdentifier id="9EFC1538728A78AF6EA72942AF796575" type="email">rodrin.rivera@nmsc.edu.ph</mods:nameIdentifier>
</mods:name>
<mods:name id="C7FF90663D808025B28EDC060ECBA609" type="personal">
<mods:role id="A4FAAB247718269A69F93064B83A9AF3">
<mods:roleTerm id="01601A6188D0310AD23FCC7DE1762F2B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6A1337C2331001AD31EB9F0E91D3ACBF">Nuñeza, Olga Macas</mods:namePart>
<mods:nameIdentifier id="EF8034728058ED4AF38D695504C951F7" type="ORCID">0000-0002-7102-9674</mods:nameIdentifier>
<mods:affiliation id="82E97403FC17B73A3AC07DCB24DDA271">Department of Biological Sciences, College of Science and Mathematics, Mindanao State University-Iligan Institute of Technology, Tibanga, Iligan City 9200, Philippines</mods:affiliation>
<mods:nameIdentifier id="F474015BD09D379471218B704774E66E" type="email">olga.nuneza@g.msuiit.edu.ph</mods:nameIdentifier>
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<mods:typeOfResource id="E9EC94E5ABE4564D7F3FE6681261A5B1">text</mods:typeOfResource>
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<mods:title id="DA47C227F01D4DCDCA080FCB7D49EA91">Zootaxa</mods:title>
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<mods:part id="B8AF6EACCC39D6FAC5068FAF9D38A3C9">
<mods:date id="A21F85AFC5CFD77ECA6F3561433F52E1">2024</mods:date>
<mods:detail id="8A35A3E63B18A8C37613AB48B59B3643" type="pubDate">
<mods:number id="EB1A402DD10549CA416161546C8F1D5C">2024-09-06</mods:number>
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<mods:detail id="BB748614118BFC6D2A568522F186F8FC" type="volume">
<mods:number id="113966FDAD43E6EB8BACE762CA9A7249">5506</mods:number>
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<mods:number id="A9E09C2B60B013BCBEA972CACFAAF654">2</mods:number>
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<mods:url id="C1D1705C952C3F2C3603246DFE25ADC6">http://dx.doi.org/10.11646/zootaxa.5506.2.2</mods:url>
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<mods:classification id="F6ED186106F5B7AC6F24E37F7C45A692">journal article</mods:classification>
<mods:identifier id="DB2794342234FBEC3CA9EF52B08A011C" type="DOI">10.11646/zootaxa.5506.2.2</mods:identifier>
<mods:identifier id="64CAD61E6D2B4C730347B36AEEF6650E" type="ISSN">1175-5326</mods:identifier>
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<subSubSection id="C3001A92A350FFA1FF46D68CFCA73523" box="[151,781,1012,1039]" pageId="3" pageNumber="197" type="nomenclature">
<paragraph id="8BA54919A350FFA1FF46D68CFCA73523" blockId="3.[151,781,1012,1039]" box="[151,781,1012,1039]" pageId="3" pageNumber="197">
<heading id="D0EDFE75A350FFA1FF46D68CFCA73523" bold="true" box="[151,781,1012,1039]" fontSize="11" level="1" pageId="3" pageNumber="197" reason="1">
<emphasis id="B96E950BA350FFA1FF46D68CFCA73523" bold="true" box="[151,781,1012,1039]" pageId="3" pageNumber="197">
Genus
<taxonomicName id="4C1A329AA350FFA1FF3FD68CFD393523" authority="Tumbrinck &amp; Tan" authorityName="Tumbrinck &amp; Tan" authorityYear="2024" box="[238,659,1012,1039]" class="Insecta" family="Tetrigidae" genus="Fartmanntettix" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="197" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96E950BA350FFA1FF3FD68CFE0C3522" bold="true" box="[238,422,1012,1038]" italics="true" pageId="3" pageNumber="197">Fartmanntettix</emphasis>
Tumbrinck &amp; Tan
</taxonomicName>
,
<taxonomicNameLabel id="A25D2870A350FFA1FD4ED68DFCA73523" box="[671,781,1013,1039]" pageId="3" pageNumber="197" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3001A92A350FFA1FF46D141FB8D357D" box="[151,1063,1081,1105]" pageId="3" pageNumber="197" type="type_taxon">
<paragraph id="8BA54919A350FFA1FF46D141FB8D357D" blockId="3.[151,1063,1081,1105]" box="[151,1063,1081,1105]" pageId="3" pageNumber="197">
<emphasis id="B96E950BA350FFA1FF46D141FE8C357D" bold="true" box="[151,294,1081,1105]" pageId="3" pageNumber="197">
<typeStatus id="54A1F7BBA350FFA1FF46D141FF66357D" box="[151,204,1081,1105]" pageId="3" pageNumber="197">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C1A329AA350FFA1FEFDD142FD46357D" authority="Tan &amp; Tumbrinck" authorityName="Tan &amp; Tumbrinck" authorityYear="2024" box="[300,748,1081,1105]" class="Insecta" family="Tetrigidae" genus="Fartmanntettix" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="197" phylum="Arthropoda" rank="species" species="undulatus" status="gen. et sp. nov.">
<emphasis id="B96E950BA350FFA1FEFDD142FD84357D" box="[300,558,1081,1105]" italics="true" pageId="3" pageNumber="197">Fartmanntettix undulatus</emphasis>
Tan &amp; Tumbrinck
</taxonomicName>
,
<emphasis id="B96E950BA350FFA1FD27D141FC39357D" bold="true" box="[758,915,1081,1105]" pageId="3" pageNumber="197">
<taxonomicNameLabel id="A25D2870A350FFA1FD27D141FC39357D" box="[758,915,1081,1105]" pageId="3" pageNumber="197" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, by monotypy
</paragraph>
</subSubSection>
<subSubSection id="C3001A92A350FFA1FF46D107FDCC3741" pageId="3" pageNumber="197" type="diagnosis">
<paragraph id="8BA54919A350FFA1FF46D107FC273766" blockId="3.[151,1437,1151,2042]" pageId="3" pageNumber="197">
<emphasis id="B96E950BA350FFA1FF46D107FEB835B5" bold="true" box="[151,274,1151,1177]" pageId="3" pageNumber="197">Diagnosis.</emphasis>
Fairly stout habitus that is neither distinctly compressed dorso-ventrally nor rounded. Vertex is wider than the width of one eye, flat or slightly concave; middle carina is distinct; lateral carinae slightly elevated to form low horns; apex of vertex surpassing the front of eyes. Bifurcation of frontal costa in line with middle of eye. Facial carinae of the frontal costa are considerably widened and enclose a broadened scutellum. Scutellum is less wide than the scapus. Antennae long, segments elongated and flattened but not foliaceous. Eyes usually not elevated above pronotum. Lateral ocelli situated at level with the middle of eyes; medial ocellus located below the lower margin of the eyes. Pronotum rugose and not surpassing abdominal apex and apices of posterior femora. Pronotal disk generally flattened with undulated carinae: median carina is prominent throughout its length and undulated, most prominent at anterior half and highest between prozona and interhumeral carinae. Median carina is also flanked by a network of irregular carinae: prozona and interhumeral carinae roundly raised. Angle of ventral sinus between lateral shoulder and lateral lobe deep and rounded. Lower margin of lateral lobe of pronotum bluntly triangular with obtuse angle. Apterous. Anterior and middle femora with dorsal margins having well-developed carinae. First segment of the tarsus on the posterior legs is as long as 3
<superScript id="7C6FE451A350FFA1FCC2D357FC8B3711" attach="left" box="[787,801,1583,1597]" fontSize="6" pageId="3" pageNumber="197">rd</superScript>
segment.
</paragraph>
<paragraph id="8BA54919A350FFA1FF16D32BFDCC3741" blockId="3.[151,1437,1151,2042]" box="[199,614,1619,1646]" pageId="3" pageNumber="197">
<emphasis id="B96E950BA350FFA1FF16D32BFE383741" bold="true" box="[199,402,1619,1645]" pageId="3" pageNumber="197">Included species.</emphasis>
<typeStatus id="54A1F7BBA350FFA1FE49D32CFE7A3742" box="[408,464,1620,1646]" pageId="3" pageNumber="197">Type</typeStatus>
species only.
</paragraph>
</subSubSection>
<subSubSection id="C3001A92A350FFA6FF16D30FFE173049" lastPageId="4" lastPageNumber="198" pageId="3" pageNumber="197" type="discussion">
<paragraph id="8BA54919A350FFA1FF16D30FFF5A360D" blockId="3.[151,1437,1151,2042]" pageId="3" pageNumber="197">
<emphasis id="B96E950BA350FFA1FF16D30FFE9137BD" bold="true" box="[199,315,1655,1681]" pageId="3" pageNumber="197">Remarks.</emphasis>
We consider this new genus under the subfamily
<taxonomicName id="4C1A329AA350FFA1FC4ED30FFB9E37BD" authorityName="Bolivar" authorityYear="1887" box="[927,1076,1655,1681]" class="Insecta" family="Tetrigidae" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="197" phylum="Arthropoda" rank="subFamily" subFamily="Cladonotinae">Cladonotinae</taxonomicName>
by these characters following
<bibRefCitation id="EF8B34E8A350FFA1FF46D3E3FEC23799" author="Tumbrinck, J." box="[151,360,1691,1717]" pageId="3" pageNumber="197" pagination="350 - 396" refId="ref5999" refString="Tumbrinck, J. (2014) Taxonomic revision of the Cladonotinae (Orthoptera: Tetrigidae) from the islands of South-East Asia and from Australia, with general remarks to the classification and morphology of the Tetrigidae and descriptions of new genera and species from New Guinea and New Caledonia. In: Telnov, D. (Ed.), Biodiversity, Biogeography and Nature Conservation in Wallacea and New Guinea, II, pp. 350 - 396." type="book chapter" year="2014">Tumbrinck (2014)</bibRefCitation>
: the medial ocellus located below the lower margin of the eyes; the fascial carinae of the frontal costa considerably widened and enclose a broadened scutellum (in this genus the scutellum is however less wide than the scapus, atypical in this subfamily); dorsal margins of the anterior and middle femora with well-developed carinae.
</paragraph>
<paragraph id="8BA54919A350FFA1FF16D253FE5336FA" blockId="3.[151,1437,1151,2042]" pageId="3" pageNumber="197">
<emphasis id="B96E950BA350FFA1FF16D253FEC83669" bold="true" box="[199,354,1835,1861]" pageId="3" pageNumber="197">Comparison.</emphasis>
This new genus is similar to
<taxonomicName id="4C1A329AA350FFA1FD6AD253FB953669" authority="Storozhenko, 2023" authorityName="Storozhenko" authorityYear="2023" box="[699,1087,1835,1861]" class="Insecta" family="Tetrigidae" genus="Trusmaditetrix" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B96E950BA350FFA1FD6AD253FCCA3669" box="[699,864,1835,1861]" italics="true" pageId="3" pageNumber="197">Trusmaditetrix</emphasis>
Storozhenko, 2023
</taxonomicName>
from Borneo by the generally flattened pronotum and widened vertex and the anterior and middle femora having undulating dorsal and ventral margins and the relative length of the 1
<superScript id="7C6FE451A350FFA1FD84D20BFDCA36AD" attach="left" box="[597,608,1907,1921]" fontSize="6" pageId="3" pageNumber="197">st</superScript>
and 3
<superScript id="7C6FE451A350FFA1FD77D20BFD1E36AD" attach="left" box="[678,692,1907,1921]" fontSize="6" pageId="3" pageNumber="197">rd</superScript>
segments of the posterior tarsus; but differs by the carinae on the pronotum more distinctly undulated and raised, forming numerous small peaks, and at apex faintly bifurcated and not surpassing abdominal apex.
</paragraph>
<paragraph id="8BA54919A350FFA6FF16D2A7FE173049" blockId="3.[151,1437,1151,2042]" lastBlockId="4.[151,1437,151,465]" lastPageId="4" lastPageNumber="198" pageId="3" pageNumber="197">
The new genus also differs from
<taxonomicName id="4C1A329AA350FFA1FDE1D2A7FB1F36D5" authority="Patano, Skejo, Amoroso &amp; Kasalo, 2024" authorityName="Patano, Skejo, Amoroso &amp; Kasalo" authorityYear="2024" box="[560,1205,2015,2042]" class="Insecta" family="Tetrigidae" genus="Storozhenkotettix" higherTaxonomySource="CoL" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B96E950BA350FFA1FDE1D2A7FD5B36D5" box="[560,753,2015,2041]" italics="true" pageId="3" pageNumber="197">Storozhenkotettix</emphasis>
<bibRefCitation id="EF8B34E8A350FFA1FD26D298FB1F36D5" author="Patano, R. R., Jr. &amp; Skejo, J. &amp; Amoroso, V. B. &amp; Kasalo, N." box="[759,1205,2015,2042]" pageId="3" pageNumber="197" pagination="577 - 586" refId="ref5287" refString="Patano, R. R., Jr., Skejo, J., Amoroso, V. B. &amp; Kasalo, N. (2024) Storozhenkotettix, a new genus of Trusmaditetrigini (Orthoptera: Tetrigidae) from Mindanao, with notes on Bolivaritettix. Zootaxa, 5453 (4), 577 - 586. https: // doi. org / 10.11646 / zootaxa. 5453.4.8" type="journal article" year="2024">Patano, Skejo, Amoroso &amp; Kasalo, 2024</bibRefCitation>
</taxonomicName>
from the
<collectingCountry id="F30D0989A350FFA1FACFD2A7FA3636D5" box="[1310,1436,2015,2041]" name="Philippines" pageId="3" pageNumber="197">Philippines</collectingCountry>
by the vertex distinctly wider than an eye, the pronotum more rugose and at apex only faintly bifurcated and the femora with carinae more undulated; from
<taxonomicName id="4C1A329AA357FFA6FD61D5C3FC5D31F9" authority="Tumbrinck, 2014" authorityName="Tumbrinck" authorityYear="2014" box="[688,1015,187,213]" class="Insecta" family="Tetrigidae" genus="Planotettix" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="198" phylum="Arthropoda" rank="genus">
<emphasis id="B96E950BA357FFA6FD61D5C3FC8131F9" box="[688,811,187,213]" italics="true" pageId="4" pageNumber="198">Planotettix</emphasis>
<bibRefCitation id="EF8B34E8A357FFA6FCE5D5C3FC5D31F9" author="Tumbrinck, J." box="[820,1015,187,213]" pageId="4" pageNumber="198" pagination="350 - 396" refId="ref5999" refString="Tumbrinck, J. (2014) Taxonomic revision of the Cladonotinae (Orthoptera: Tetrigidae) from the islands of South-East Asia and from Australia, with general remarks to the classification and morphology of the Tetrigidae and descriptions of new genera and species from New Guinea and New Caledonia. In: Telnov, D. (Ed.), Biodiversity, Biogeography and Nature Conservation in Wallacea and New Guinea, II, pp. 350 - 396." type="book chapter" year="2014">Tumbrinck, 2014</bibRefCitation>
</taxonomicName>
from New
<collectingCountry id="F30D0989A357FFA6FBACD5C3FB6431F9" box="[1149,1230,187,213]" name="Guinea-Bissau" pageId="4" pageNumber="198">Guinea</collectingCountry>
by the larger size (prontoum length&gt;
<quantity id="4CE2E4FCA357FFA6FEBED5A6FE1831D5" box="[367,434,222,249]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="198" unit="mm" value="5.0">5 mm</quantity>
), less dorso-ventrally flattened habitus and the legs having less strongly undulated carinae; from
<taxonomicName id="4C1A329AA357FFA6FF04D47BFD903031" authority="Tumbrinck, 2014" authorityName="Tumbrinck" authorityYear="2014" box="[213,570,259,285]" class="Insecta" family="Tetrigidae" genus="Devriesetettix" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="198" phylum="Arthropoda" rank="genus">
<emphasis id="B96E950BA357FFA6FF04D47BFEDB3030" box="[213,369,259,284]" italics="true" pageId="4" pageNumber="198">Devriesetettix</emphasis>
<bibRefCitation id="EF8B34E8A357FFA6FEA9D47BFD903031" author="Tumbrinck, J." box="[376,570,259,285]" pageId="4" pageNumber="198" pagination="350 - 396" refId="ref5999" refString="Tumbrinck, J. (2014) Taxonomic revision of the Cladonotinae (Orthoptera: Tetrigidae) from the islands of South-East Asia and from Australia, with general remarks to the classification and morphology of the Tetrigidae and descriptions of new genera and species from New Guinea and New Caledonia. In: Telnov, D. (Ed.), Biodiversity, Biogeography and Nature Conservation in Wallacea and New Guinea, II, pp. 350 - 396." type="book chapter" year="2014">Tumbrinck, 2014</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C1A329AA357FFA6FDA3D47BFC1C3031" authority="Tumbrinck, 2014" authorityName="Tumbrinck" authorityYear="2014" box="[626,950,259,285]" class="Insecta" family="Tetrigidae" genus="Planotettix" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="198" phylum="Arthropoda" rank="genus">
<emphasis id="B96E950BA357FFA6FDA3D47BFD473031" box="[626,749,259,285]" italics="true" pageId="4" pageNumber="198">Planotettix</emphasis>
<bibRefCitation id="EF8B34E8A357FFA6FD25D47BFC1C3031" author="Tumbrinck, J." box="[756,950,259,285]" pageId="4" pageNumber="198" pagination="350 - 396" refId="ref5999" refString="Tumbrinck, J. (2014) Taxonomic revision of the Cladonotinae (Orthoptera: Tetrigidae) from the islands of South-East Asia and from Australia, with general remarks to the classification and morphology of the Tetrigidae and descriptions of new genera and species from New Guinea and New Caledonia. In: Telnov, D. (Ed.), Biodiversity, Biogeography and Nature Conservation in Wallacea and New Guinea, II, pp. 350 - 396." type="book chapter" year="2014">Tumbrinck, 2014</bibRefCitation>
</taxonomicName>
from New
<collectingCountry id="F30D0989A357FFA6FBE9D47BFB233031" box="[1080,1161,259,285]" name="Guinea-Bissau" pageId="4" pageNumber="198">Guinea</collectingCountry>
and
<taxonomicName id="4C1A329AA357FFA6FB10D47BFF65306D" authority="Rehn, 1952" authorityName="Rehn" authorityYear="1952" class="Insecta" family="Tetrigidae" genus="Tepperotettix" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="198" phylum="Arthropoda" rank="genus">
<emphasis id="B96E950BA357FFA6FB10D47BFAF93030" box="[1217,1363,259,284]" italics="true" pageId="4" pageNumber="198">Tepperotettix</emphasis>
Rehn, 1952
</taxonomicName>
from
<collectingCountry id="F30D0989A357FFA6FEC6D45FFED4306D" box="[279,382,295,321]" name="Australia" pageId="4" pageNumber="198">Australia</collectingCountry>
by its vertex, the apex of its pronotal process narrow, shape of median carina, and the legs distinctly more undulated.
</paragraph>
</subSubSection>
<subSubSection id="C3001A92A357FFA6FF16D417FA3330FD" pageId="4" pageNumber="198" type="etymology">
<paragraph id="8BA54919A357FFA6FF16D417FF5B3081" blockId="4.[151,1437,151,465]" pageId="4" pageNumber="198">
<emphasis id="B96E950BA357FFA6FF16D417FEE430A5" bold="true" box="[199,334,367,393]" pageId="4" pageNumber="198">Etymology.</emphasis>
The new genus is named after Thomas Fartmann, former president of the German Orthopterological Society.
</paragraph>
<paragraph id="8BA54919A357FFA6FF16D4CFFA3330FD" blockId="4.[151,1437,151,465]" box="[199,1433,439,465]" pageId="4" pageNumber="198">The suffix “tettix” refers to the Ancient Greek word for “grasshopper”, making the name of masculine gender.</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="E349C35B16EB47E26BACA6DA393BE6E1">A new genus and a new species of Cladonotinae (Orthoptera: Tetrigidae) from Mindanao, Philippines</mods:title>
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<mods:namePart id="47272642F236465025CBA424C6B0B42F">Tan, Ming Kai</mods:namePart>
<mods:affiliation id="9460FC489FDFC9A643E8065B0459AC31">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore</mods:affiliation>
</mods:name>
<mods:name id="CE9A66A0168B5799436856FA68B96135" type="personal">
<mods:role id="FAD16821F9D720E1D8165F92C18A30CE">
<mods:roleTerm id="72D1DB5B23A8116672C286C26ED5D014">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="949DA6ADC2F575ACA9136C6DFB7C0ECE">Tumbrinck, Josef</mods:namePart>
<mods:affiliation id="593975F95ADF9D3A6B76E528187DA7CB">Auf der Hees 1, D- 41849 Wassenberg, Germany</mods:affiliation>
</mods:name>
<mods:name id="A3FF9170BD045F2A8A8DA9F692FC0A12" type="personal">
<mods:role id="5DDA1E33C63B3D153385DAA50D6D11CB">
<mods:roleTerm id="BA1032BB992E5FEBB92522773C71DB1E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5478AE4A4E88D1843598E7DA28293457">Rivera, Rodrin R.</mods:namePart>
<mods:nameIdentifier id="333EAFE0969E24F04F2471617B2AB814" type="ORCID">0000-0003-3019-0258</mods:nameIdentifier>
<mods:affiliation id="189DDE6394D436D7B0685341384C4355">Department of Biological Sciences, College of Science and Mathematics, Mindanao State University-Iligan Institute of Technology, Tibanga, Iligan City 9200, Philippines</mods:affiliation>
<mods:nameIdentifier id="FB2771025125CE6592D2CA0448F24E9F" type="email">rodrin.rivera@nmsc.edu.ph</mods:nameIdentifier>
</mods:name>
<mods:name id="ED850C01787EAE9C104CE031359DAC9A" type="personal">
<mods:role id="38D11372C8EB170BCF7A5E1C02DD53D7">
<mods:roleTerm id="EEE70A8C72BCCA3D03E68C2E9CE424A2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C450F3A3EC196F5BAA07C13174670F13">Nuñeza, Olga Macas</mods:namePart>
<mods:nameIdentifier id="93AF69F030B48BE05CF98F9CEF8498BF" type="ORCID">0000-0002-7102-9674</mods:nameIdentifier>
<mods:affiliation id="86C85BB22B906157224D25CD19CF84B6">Department of Biological Sciences, College of Science and Mathematics, Mindanao State University-Iligan Institute of Technology, Tibanga, Iligan City 9200, Philippines</mods:affiliation>
<mods:nameIdentifier id="8391E04F150BB2740E5E1EB98EDA4BBE" type="email">olga.nuneza@g.msuiit.edu.ph</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="25D9A383E39FD216B1F29183F2B825E9">text</mods:typeOfResource>
<mods:relatedItem id="4085C3C78A0665F3D827A7F26C62A1FD" type="host">
<mods:titleInfo id="9B67CF70A1C0F5FE39FB64CE7E082A4D">
<mods:title id="6526C1494D20B8D672D88C88479961F7">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="9AD07A8C90560CB5B000D3C0B81BF8A4">
<mods:date id="78F73D8C1D2FB3C44ACC908B72AB6E45">2024</mods:date>
<mods:detail id="E481197B917139C4BC022C8834264365" type="pubDate">
<mods:number id="9B870D0A740DE06C1CC37F2B56E13E0C">2024-09-06</mods:number>
</mods:detail>
<mods:detail id="E8EBE2DFD1970EF2B7E208CBA9EDBE4C" type="volume">
<mods:number id="877D701934BA889A0487C05931D198EC">5506</mods:number>
</mods:detail>
<mods:detail id="5BE2EA58DB8D9D4DC183C6D090B491DB" type="issue">
<mods:number id="AE89801277101E7ECEBEDCBEC45A5B53">2</mods:number>
</mods:detail>
<mods:extent id="99623B16762921EFD08BE77FA07DF1D0" unit="page">
<mods:start id="6F347A92D9F183219EE22D05EC594853">194</mods:start>
<mods:end id="FCACFF01D523A36D00C62E3C527C7B8C">204</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="A59897C904FE201B3A53317B62292DE2">
<mods:url id="8AD8FF78E5F0150182369F87B9E14175">http://dx.doi.org/10.11646/zootaxa.5506.2.2</mods:url>
</mods:location>
<mods:classification id="17FF58E342D2096E89984F0412E9E093">journal article</mods:classification>
<mods:identifier id="91BA0C9CC19AD46ADFBABB804DD6A840" type="DOI">10.11646/zootaxa.5506.2.2</mods:identifier>
<mods:identifier id="805B7B7C5CBB9CA220BDEE5CD07ED3BF" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="1C3918378CA297788D6791367DCB7B3F" type="ZooBank">03F52CEE-CECD-4FE5-93B9-8CC9F31FCCA4</mods:identifier>
</mods:mods>
<treatment id="03B3F80FA357FFA4FF46D75BFBF836B9" LSID="urn:lsid:plazi:treatment:03B3F80FA357FFA4FF46D75BFBF836B9" httpUri="http://treatment.plazi.org/id/03B3F80FA357FFA4FF46D75BFBF836B9" lastPageId="6" lastPageNumber="200" pageId="4" pageNumber="198">
<subSubSection id="C3001A92A357FFA6FF46D75BFC8D3311" box="[151,807,546,573]" pageId="4" pageNumber="198" type="nomenclature">
<paragraph id="8BA54919A357FFA6FF46D75BFC8D3311" blockId="4.[151,807,546,609]" box="[151,807,546,573]" pageId="4" pageNumber="198">
<heading id="D0EDFE75A357FFA6FF46D75BFC8D3311" bold="true" box="[151,807,546,573]" fontSize="11" level="1" pageId="4" pageNumber="198" reason="1">
<emphasis id="B96E950BA357FFA6FF46D75BFC8D3311" bold="true" box="[151,807,546,573]" pageId="4" pageNumber="198">
<taxonomicName id="4C1A329AA357FFA6FF46D75BFD173310" authority="Tan &amp; Tumbrinck" authorityName="Tan &amp; Tumbrinck" authorityYear="2024" box="[151,701,546,573]" class="Insecta" family="Tetrigidae" genus="Fartmanntettix" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="198" phylum="Arthropoda" rank="species" species="undulatus" status="sp. nov.">
<emphasis id="B96E950BA357FFA6FF46D75BFE7B3311" bold="true" box="[151,465,546,573]" italics="true" pageId="4" pageNumber="198">Fartmanntettix undulatus</emphasis>
Tan &amp; Tumbrinck
</taxonomicName>
,
<taxonomicNameLabel id="A25D2870A357FFA6FD1BD75BFC8D3311" box="[714,807,547,573]" pageId="4" pageNumber="198" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3001A92A357FFA6FF46D73FFEA1334D" box="[151,267,582,609]" pageId="4" pageNumber="198" type="description">
<paragraph id="8BA54919A357FFA6FF46D73FFEA1334D" blockId="4.[151,807,546,609]" box="[151,267,582,609]" pageId="4" pageNumber="198">
(
<figureCitation id="1321559CA357FFA6FF4ED73FFEA9334D" box="[159,259,582,609]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="4.[151,250,1803,1827]" captionStartId-1="5.[151,250,1458,1482]" captionStartId-2="7.[151,250,1899,1923]" captionStartId-3="8.[151,250,1908,1932]" captionTargetBox-0="[169,1417,813,1779]" captionTargetBox-1="[162,1425,194,1434]" captionTargetBox-2="[164,1422,181,1875]" captionTargetBox-3="[192,1396,181,1881]" captionTargetId-0="figure-196@4.[151,1436,813,1779]" captionTargetId-1="figure-21@5.[151,1436,181,1434]" captionTargetId-2="figure-21@7.[164,1423,181,1875]" captionTargetId-3="figure-17@8.[192,1396,181,1885]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="7" captionTargetPageId-3="8" captionText-0="FIGURE 2. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♂ holotype: habitus in dorsal (A) and lateral (B) views. Scale bars: 1 mm." captionText-1="FIGURE 3. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♀ allotype: habitus in dorsal (A) and lateral (B) views. Scale bars: 1 mm." captionText-2="FIGURE 4. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♂ holotype: face in anterior view (A); head and anterior half of pronotum in dorsal (B) and lateral (C) views; posterior half of pronotum in dorsal view (D); anterior (E) and middle (F) legs in external views; posterior femur in external (G) and dorsal (H) views; posterior tibia in external view (I). Scale bars: 1 mm." captionText-3="FIGURE 5. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♀ allotype: face in anterior view (A); head and anterior half of pronotum in dorsal (B) and lateral (C) views; anterior (D) and middle (E) legs in external views; posterior femur in dorsal (F) and external (G) views; posterior tibia in external view (H); abdominal apex in lateral (I) and ventral (J) views. Scale bars: 1 mm." pageId="4" pageNumber="198">Figs 25</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3001A92A357FFA6FF46D7F6FC1536D9" pageId="4" pageNumber="198" type="materials_examined">
<paragraph id="8BA54919A357FFA6FF46D7F6FD643239" blockId="4.[151,1436,654,789]" pageId="4" pageNumber="198">
<emphasis id="B96E950BA357FFA6FF46D7F6FE293384" bold="true" box="[151,387,654,680]" pageId="4" pageNumber="198">Material examined.</emphasis>
<materialsCitation id="3B724344A357FFA6FE5AD7F7FD643239" collectingDate="2024-01-09" collectionCode="PNM" collectorName="Abaca &amp; M. K. Tan" country="Philippines" county="Mindanao Island" elevation="8" latitude="8.30801" location="Lake Duminagat Barangay" longLatPrecision="1" longitude="123.6172" municipality="Don Victoriano Chiongbian" pageId="4" pageNumber="198" specimenCount="1" specimenCount-male="1" stateProvince="Misamis Occidental" typeStatus="holotype">
<emphasis id="B96E950BA357FFA6FE5AD7F7FDAB3385" bold="true" box="[395,513,655,681]" pageId="4" pageNumber="198">
<typeStatus id="54A1F7BBA357FFA6FE5AD7F7FE563385" box="[395,508,655,681]" pageId="4" pageNumber="198" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectingCountry id="F30D0989A357FFA6FDDBD7F7FD193385" box="[522,691,655,681]" name="Philippines" pageId="4" pageNumber="198">PHILIPPINES</collectingCountry>
:
<collectingCounty id="62C43195A357FFA6FD11D7F7FC293385" box="[704,899,655,681]" pageId="4" pageNumber="198">Mindanao Island</collectingCounty>
:
<collectingRegion id="49DE87FBA357FFA6FC5ED7F7FB4B3385" box="[911,1249,655,681]" country="Philippines" name="Misamis Occidental" pageId="4" pageNumber="198">Misamis Occidental province</collectingRegion>
:
<collectingMunicipality id="6BC1D363A357FFA6FB3FD7F7FEB433E1" pageId="4" pageNumber="198">Don Victoriano Chiongbian</collectingMunicipality>
:
<location id="8EC51FC2A357FFA6FEFDD7CBFDF533E1" LSID="urn:lsid:plazi:treatment:03B3F80FA357FFA4FF46D75BFBF836B9:8EC51FC2A357FFA6FEFDD7CBFDF533E1" box="[300,607,691,717]" country="Philippines" county="Mindanao Island" latitude="8.30801" longLatPrecision="1" longitude="123.6172" municipality="Don Victoriano Chiongbian" name="Lake Duminagat Barangay" pageId="4" pageNumber="198" stateProvince="Misamis Occidental">Lake Duminagat Barangay</location>
<specimenCount id="9D1C8290A357FFA6FDACD7CBFD3F33E1" box="[637,661,691,717]" count="1" pageId="4" pageNumber="198" type="male"></specimenCount>
;
<location id="8EC51FC2A357FFA6FD75D7CBFBBA33E1" LSID="urn:lsid:plazi:treatment:03B3F80FA357FFA4FF46D75BFBF836B9:8EC51FC2A357FFA6FD75D7CBFBBA33E1" box="[676,1040,691,717]" country="Philippines" county="Mindanao Island" latitude="8.30801" longLatPrecision="1" longitude="123.6172" municipality="Don Victoriano Chiongbian" name="Mount Malindang Natural Park" pageId="4" pageNumber="198" stateProvince="Misamis Occidental">Mount Malindang Natural Park</location>
, abandoned
<location id="8EC51FC2A357FFA6FB71D7CBFB4233E1" LSID="urn:lsid:plazi:treatment:03B3F80FA357FFA4FF46D75BFBF836B9:8EC51FC2A357FFA6FB71D7CBFB4233E1" box="[1184,1256,691,717]" country="Philippines" county="Mindanao Island" latitude="8.30801" longLatPrecision="1" longitude="123.6172" municipality="Don Victoriano Chiongbian" name="Abaca" pageId="4" pageNumber="198" stateProvince="Misamis Occidental">Abacá</location>
plantation near
<location id="8EC51FC2A357FFA6FF46D7AFFEB133DD" LSID="urn:lsid:plazi:treatment:03B3F80FA357FFA4FF46D75BFBF836B9:8EC51FC2A357FFA6FF46D7AFFEB133DD" box="[151,283,727,753]" country="Philippines" county="Mindanao Island" latitude="8.30801" longLatPrecision="1" longitude="123.6172" municipality="Don Victoriano Chiongbian" name="Lake Linao" pageId="4" pageNumber="198" stateProvince="Misamis Occidental">Lake Linao</location>
;
<geoCoordinate id="EE2E2FDEA357FFA6FEF6D7AFFE3233DD" box="[295,408,727,753]" degrees="8.30801" direction="north" orientation="latitude" pageId="4" pageNumber="198" precision="1" value="8.30801">N8.30801</geoCoordinate>
,
<geoCoordinate id="EE2E2FDEA357FFA6FE75D7AFFD8533DD" box="[420,559,727,753]" degrees="123.61720" direction="east" orientation="longitude" pageId="4" pageNumber="198" precision="1" value="123.6172">E123.61720</geoCoordinate>
, 1571.0±
<quantity id="4CE2E4FCA357FFA6FD48D7AEFD7433DD" box="[665,734,726,753]" metricMagnitude="0" metricUnit="m" metricValue="8.1" pageId="4" pageNumber="198" unit="m" value="8.1">
<elevation id="0037AE2AA357FFA6FD48D7AEFD7433DD" box="[665,734,726,753]" metricMagnitude="0" metricUnit="m" metricValue="8.1" pageId="4" pageNumber="198" unit="m" value="8.1">8.1 m</elevation>
</quantity>
.a.s.l;
<date id="FFA46FD9A357FFA6FCCBD7AFFC6233DD" box="[794,968,727,753]" pageId="4" pageNumber="198" value="2024-01-09">
<collectingDate id="EFE09631A357FFA6FCCBD7AFFC6233DD" box="[794,968,727,753]" pageId="4" pageNumber="198" value="2024-01-09">9 January 2024</collectingDate>
</date>
, 10h00; among wet decaying
<collectorName id="26EF2CCFA357FFA6FAC7D7AFFAF433DD" box="[1302,1374,727,753]" pageId="4" pageNumber="198">Abacá</collectorName>
plant on the ground; coll.
<collectorName id="26EF2CCFA357FFA6FEABD783FE423239" box="[378,488,763,789]" pageId="4" pageNumber="198">M.K. Tan</collectorName>
; MIN.24.24 (
<collectionCode id="ED0BD1DCA357FFA6FDAED783FD683239" box="[639,706,763,789]" country="Philippines" httpUri="http://grbio.org/cool/mxaf-c5e2" name="Philippine National Museum" pageId="4" pageNumber="198">PNM</collectionCode>
).
</materialsCitation>
</paragraph>
<caption id="DF651991A357FFA6FF46D273FE3F366B" pageId="4" pageNumber="198" startId="4.[151,250,1803,1827]" targetBox="[169,1417,813,1779]" targetPageId="4" targetType="figure">
<paragraph id="8BA54919A357FFA6FF46D273FE3F366B" blockId="4.[151,1436,1803,1863]" pageId="4" pageNumber="198">
<emphasis id="B96E950BA357FFA6FF46D273FEBF360F" bold="true" box="[151,277,1803,1827]" pageId="4" pageNumber="198">FIGURE 2.</emphasis>
<taxonomicName id="4C1A329AA357FFA6FECFD274FD4D360F" authority="Tan &amp; Tumbrinck" authorityName="Tan &amp; Tumbrinck" authorityYear="2024" box="[286,743,1803,1827]" class="Insecta" family="Tetrigidae" genus="Fartmanntettix" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="198" phylum="Arthropoda" rank="species" species="undulatus" status="gen. et sp. nov.">
<emphasis id="B96E950BA357FFA6FECFD274FD89360F" box="[286,547,1803,1827]" italics="true" pageId="4" pageNumber="198">Fartmanntettix undulatus</emphasis>
Tan &amp; Tumbrinck
</taxonomicName>
,
<emphasis id="B96E950BA357FFA6FD25D273FC32360F" bold="true" box="[756,920,1803,1827]" pageId="4" pageNumber="198">
<taxonomicNameLabel id="A25D2870A357FFA6FD25D273FC32360F" box="[756,920,1803,1827]" pageId="4" pageNumber="198" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
♂ holotype: habitus in dorsal (A) and lateral (B) views. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="8BA54919A357FFA6FF16D2EBFC1536D9" blockId="4.[151,1436,1938,2038]" pageId="4" pageNumber="198">
<materialsCitation id="3B724344A357FFA6FF16D2EBFC1136D9" collectingDate="2024-01-09" collectionCode="ZRC" collectorName="Abaca &amp; M. K. Tan" country="Philippines" county="Mindanao Island" elevation="8" latitude="8.30801" location="Lake Duminagat Barangay" longLatPrecision="1" longitude="123.6172" municipality="Don Victoriano Chiongbian" pageId="4" pageNumber="198" specimenCount="1" specimenCount-female="1" stateProvince="Misamis Occidental" typeStatus="paratype">
<emphasis id="B96E950BA357FFA6FF16D2EBFE6A3680" bold="true" box="[199,448,1938,1965]" pageId="4" pageNumber="198">
<typeStatus id="54A1F7BBA357FFA6FF16D2EBFE9E3681" box="[199,308,1939,1965]" pageId="4" pageNumber="198" type="paratype">Paratype</typeStatus>
(
<typeStatus id="54A1F7BBA357FFA6FE9CD2EAFE193680" box="[333,435,1938,1964]" pageId="4" pageNumber="198" type="allotype">allotype</typeStatus>
):
</emphasis>
<collectingCountry id="F30D0989A357FFA6FE00D2EBFDD03681" box="[465,634,1939,1965]" name="Philippines" pageId="4" pageNumber="198">PHILIPPINES</collectingCountry>
:
<collectingCounty id="62C43195A357FFA6FD5ED2EBFCF03681" box="[655,858,1939,1965]" pageId="4" pageNumber="198">Mindanao Island</collectingCounty>
:
<collectingRegion id="49DE87FBA357FFA6FCBED2EBFB7B3681" box="[879,1233,1939,1965]" country="Philippines" name="Misamis Occidental" pageId="4" pageNumber="198">Misamis Occidental province</collectingRegion>
:
<collectingMunicipality id="6BC1D363A357FFA6FB34D2EBFEB436FD" pageId="4" pageNumber="198">Don Victoriano Chiongbian</collectingMunicipality>
:
<location id="8EC51FC2A357FFA6FEFCD2CFFDCE36FD" LSID="urn:lsid:plazi:treatment:03B3F80FA357FFA4FF46D75BFBF836B9:8EC51FC2A357FFA6FEFCD2CFFDCE36FD" box="[301,612,1975,2001]" country="Philippines" county="Mindanao Island" municipality="Don Victoriano Chiongbian" name="Lake Duminagat Barangay" pageId="4" pageNumber="198" stateProvince="Misamis Occidental">Lake Duminagat Barangay</location>
<specimenCount id="9D1C8290A357FFA6FD57D2CFFD0036FD" box="[646,682,1975,2001]" count="1" pageId="4" pageNumber="198" type="female">1♀</specimenCount>
; same locality as holotype;
<date id="FFA46FD9A357FFA6FC2AD2CFFB1836FD" box="[1019,1202,1975,2001]" pageId="4" pageNumber="198" value="2024-01-09">
<collectingDate id="EFE09631A357FFA6FC2AD2CFFB1836FD" box="[1019,1202,1975,2001]" pageId="4" pageNumber="198" value="2024-01-09">9 January 2024</collectingDate>
</date>
, 10h00; among wet decaying
<collectorName id="26EF2CCFA357FFA6FED3D2A3FEE036D9" box="[258,330,2011,2037]" pageId="4" pageNumber="198">Abacá</collectorName>
plant on the ground; coll.
<collectorName id="26EF2CCFA357FFA6FDA3D2A3FD4A36D9" box="[626,736,2011,2037]" pageId="4" pageNumber="198">M.K. Tan</collectorName>
; MIN.24.25 (
<collectionCode id="ED0BD1DCA357FFA6FCA6D2A3FC1936D9" box="[887,947,2011,2037]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="198">ZRC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF651991A356FFA7FF46D0CAFEE434C2" pageId="5" pageNumber="199" startId="5.[151,250,1458,1482]" targetBox="[162,1425,194,1434]" targetPageId="5" targetType="figure">
<paragraph id="8BA54919A356FFA7FF46D0CAFEE434C2" blockId="5.[151,1437,1458,1519]" pageId="5" pageNumber="199">
<emphasis id="B96E950BA356FFA7FF46D0CAFEB834E6" bold="true" box="[151,274,1458,1482]" pageId="5" pageNumber="199">FIGURE 3.</emphasis>
<taxonomicName id="4C1A329AA356FFA7FEC6D0CBFD7934E6" authority="Tan &amp; Tumbrinck" authorityName="Tan &amp; Tumbrinck" authorityYear="2024" box="[279,723,1458,1483]" class="Insecta" family="Tetrigidae" genus="Fartmanntettix" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="199" phylum="Arthropoda" rank="species" species="undulatus" status="gen. et sp. nov.">
<emphasis id="B96E950BA356FFA7FEC6D0CBFDB334E6" box="[279,537,1458,1482]" italics="true" pageId="5" pageNumber="199">Fartmanntettix undulatus</emphasis>
Tan &amp; Tumbrinck
</taxonomicName>
,
<emphasis id="B96E950BA356FFA7FD0DD0CBFCDC34E7" bold="true" box="[732,886,1459,1483]" pageId="5" pageNumber="199">
<taxonomicNameLabel id="A25D2870A356FFA7FD0DD0CBFCDC34E7" box="[732,886,1459,1483]" pageId="5" pageNumber="199" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
♀ allotype: habitus in dorsal (A) and lateral (B) views. Scale bars: 1 mm.
</paragraph>
</caption>
<subSubSection id="C3001A92A356FFA7FF16D338FD273777" box="[199,653,1600,1627]" pageId="5" pageNumber="199" type="diagnosis">
<paragraph id="8BA54919A356FFA7FF16D338FD273777" blockId="5.[151,1436,1600,2023]" box="[199,653,1600,1627]" pageId="5" pageNumber="199">
<emphasis id="B96E950BA356FFA7FF16D338FEE83776" bold="true" box="[199,322,1600,1626]" pageId="5" pageNumber="199">Diagnosis.</emphasis>
Refer to the genus diagnosis.
</paragraph>
</subSubSection>
<subSubSection id="C3001A92A356FFA7FF16D31DFB0E3753" box="[199,1188,1637,1663]" pageId="5" pageNumber="199" type="etymology">
<paragraph id="8BA54919A356FFA7FF16D31DFB0E3753" blockId="5.[151,1436,1600,2023]" box="[199,1188,1637,1663]" pageId="5" pageNumber="199">
<emphasis id="B96E950BA356FFA7FF16D31DFEE43753" bold="true" box="[199,334,1637,1663]" pageId="5" pageNumber="199">Etymology.</emphasis>
The species name refers to undulating carinae on the pronotum and femora.
</paragraph>
</subSubSection>
<subSubSection id="C3001A92A356FFA4FF16D3F0FDAE3759" lastPageId="6" lastPageNumber="200" pageId="5" pageNumber="199" type="description">
<paragraph id="8BA54919A356FFA7FF16D3F0FD6637EB" blockId="5.[151,1436,1600,2023]" pageId="5" pageNumber="199">
<emphasis id="B96E950BA356FFA7FF16D3F0FEF3378E" bold="true" box="[199,345,1672,1698]" pageId="5" pageNumber="199">Description.</emphasis>
Habitus as shown in
<figureCitation id="1321559CA356FFA7FD95D3F1FD21378F" box="[580,651,1673,1699]" captionStart="FIGURE 2" captionStartId="4.[151,250,1803,1827]" captionTargetBox="[169,1417,813,1779]" captionTargetId="figure-196@4.[151,1436,813,1779]" captionTargetPageId="4" captionText="FIGURE 2. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♂ holotype: habitus in dorsal (A) and lateral (B) views. Scale bars: 1 mm." pageId="5" pageNumber="199">Figs 2</figureCitation>
,
<figureCitation id="1321559CA356FFA7FD47D3F1FD0C378F" box="[662,678,1673,1699]" captionStart="FIGURE 3" captionStartId="5.[151,250,1458,1482]" captionTargetBox="[162,1425,194,1434]" captionTargetId="figure-21@5.[151,1436,181,1434]" captionTargetPageId="5" captionText="FIGURE 3. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♀ allotype: habitus in dorsal (A) and lateral (B) views. Scale bars: 1 mm." pageId="5" pageNumber="199">3</figureCitation>
. Different shades of brown, ranging from pale brown to dark brown, well camouflaged against leaf litter on the ground.
</paragraph>
<paragraph id="8BA54919A356FFA4FF16D3A9FF653031" blockId="5.[151,1436,1600,2023]" lastBlockId="6.[151,1437,150,1941]" lastPageId="6" lastPageNumber="200" pageId="5" pageNumber="199">
Head: In frontal view (
<figureCitation id="1321559CA356FFA7FE12D3A9FD8A37C7" box="[451,544,1745,1771]" captionStart="FIGURE 4" captionStartId="7.[151,250,1899,1923]" captionTargetBox="[164,1422,181,1875]" captionTargetId="figure-21@7.[164,1423,181,1875]" captionTargetPageId="7" captionText="FIGURE 4. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♂ holotype: face in anterior view (A); head and anterior half of pronotum in dorsal (B) and lateral (C) views; posterior half of pronotum in dorsal view (D); anterior (E) and middle (F) legs in external views; posterior femur in external (G) and dorsal (H) views; posterior tibia in external view (I). Scale bars: 1 mm." pageId="5" pageNumber="199">Figs 4A</figureCitation>
,
<figureCitation id="1321559CA356FFA7FDFAD3A8FDFB37C7" box="[555,593,1744,1771]" captionStart="FIGURE 5" captionStartId="8.[151,250,1908,1932]" captionTargetBox="[192,1396,181,1881]" captionTargetId="figure-17@8.[192,1396,181,1885]" captionTargetPageId="8" captionText="FIGURE 5. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♀ allotype: face in anterior view (A); head and anterior half of pronotum in dorsal (B) and lateral (C) views; anterior (D) and middle (E) legs in external views; posterior femur in dorsal (F) and external (G) views; posterior tibia in external view (H); abdominal apex in lateral (I) and ventral (J) views. Scale bars: 1 mm." pageId="5" pageNumber="199">5A</figureCitation>
): antennal groove inserted at lower margin of eyes, less than half a diameter of antennal groove below lower margin of eyes. Fastigium convex, carinae forming short horns. Face with numerous tiny white nodules. Frontal costa as long as half eye height in anterior view. Bifurcation of frontal costa in line with middle of eye; at widest between antennal groove; narrow ventrally just dorsal of median ocellus. Facial carinae slightly curved. Compound eyes hemispheric, not exerted above vertex. Lateral ocelli above antennal groove, at middle of eye just before bifurcation of frontal costa. In dorsal view (
<figureCitation id="1321559CA356FFA7FC7BD2FDFBAC36B3" box="[938,1030,1925,1951]" captionStart="FIGURE 4" captionStartId="7.[151,250,1899,1923]" captionTargetBox="[164,1422,181,1875]" captionTargetId="figure-21@7.[164,1423,181,1875]" captionTargetPageId="7" captionText="FIGURE 4. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♂ holotype: face in anterior view (A); head and anterior half of pronotum in dorsal (B) and lateral (C) views; posterior half of pronotum in dorsal view (D); anterior (E) and middle (F) legs in external views; posterior femur in external (G) and dorsal (H) views; posterior tibia in external view (I). Scale bars: 1 mm." pageId="5" pageNumber="199">Figs 4B</figureCitation>
,
<figureCitation id="1321559CA356FFA7FBC2D2FCFB9D36B3" box="[1043,1079,1924,1951]" captionStart="FIGURE 5" captionStartId="8.[151,250,1908,1932]" captionTargetBox="[192,1396,181,1881]" captionTargetId="figure-17@8.[192,1396,181,1885]" captionTargetPageId="8" captionText="FIGURE 5. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♀ allotype: face in anterior view (A); head and anterior half of pronotum in dorsal (B) and lateral (C) views; anterior (D) and middle (E) legs in external views; posterior femur in dorsal (F) and external (G) views; posterior tibia in external view (H); abdominal apex in lateral (I) and ventral (J) views. Scale bars: 1 mm." pageId="5" pageNumber="199">5B</figureCitation>
): apex of fastigium in line with frontal margin of eyes, emarginated at middle. Vertex 1.51.7 times wider than eye width. Lateral and median carinae of fastigium distinct, not surpassing apex of fastigium. In lateral view (
<figureCitation id="1321559CA356FFA7FC29D2B5FBF936CB" box="[1016,1107,1997,2023]" captionStart="FIGURE 4" captionStartId="7.[151,250,1899,1923]" captionTargetBox="[164,1422,181,1875]" captionTargetId="figure-21@7.[164,1423,181,1875]" captionTargetPageId="7" captionText="FIGURE 4. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♂ holotype: face in anterior view (A); head and anterior half of pronotum in dorsal (B) and lateral (C) views; posterior half of pronotum in dorsal view (D); anterior (E) and middle (F) legs in external views; posterior femur in external (G) and dorsal (H) views; posterior tibia in external view (I). Scale bars: 1 mm." pageId="5" pageNumber="199">Figs 4C</figureCitation>
,
<figureCitation id="1321559CA356FFA7FB8CD2B4FB2B36CB" box="[1117,1153,1996,2023]" captionStart="FIGURE 5" captionStartId="8.[151,250,1908,1932]" captionTargetBox="[192,1396,181,1881]" captionTargetId="figure-17@8.[192,1396,181,1885]" captionTargetPageId="8" captionText="FIGURE 5. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♀ allotype: face in anterior view (A); head and anterior half of pronotum in dorsal (B) and lateral (C) views; anterior (D) and middle (E) legs in external views; posterior femur in dorsal (F) and external (G) views; posterior tibia in external view (H); abdominal apex in lateral (I) and ventral (J) views. Scale bars: 1 mm." pageId="5" pageNumber="199">5C</figureCitation>
): frontal costa arched and well protruding in front of eyes. Antenna of female, excluding scapus and pedicel, with 11 segments. 8
<superScript id="7C6FE451A355FFA4FAFBD5EEFA9D3188" attach="left" box="[1322,1335,150,164]" fontSize="6" pageId="6" pageNumber="200">th</superScript>
segment significantly wider over the entire length than other segments. Mouthparts mostly brown to grey with small palecoloured nodules. Maxillary palps pale coloured with tint of grey; with segments stout and apical segment somewhat oval.
</paragraph>
<paragraph id="8BA54919A355FFA4FF16D45FFE3733E1" blockId="6.[151,1437,150,1941]" pageId="6" pageNumber="200">
Pronotum: Pronotal process rugose, particularly at anterior half; reaching apex of posterior leg; 1.82.0 times longer than wide (pronotal lateral lobe width). In dorsal view (
<figureCitation id="1321559CA355FFA4FC8CD433FC113049" box="[861,955,331,357]" captionStart="FIGURE 2" captionStartId="4.[151,250,1803,1827]" captionTargetBox="[169,1417,813,1779]" captionTargetId="figure-196@4.[151,1436,813,1779]" captionTargetPageId="4" captionText="FIGURE 2. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♂ holotype: habitus in dorsal (A) and lateral (B) views. Scale bars: 1 mm." pageId="6" pageNumber="200">Figs 2A</figureCitation>
,
<figureCitation id="1321559CA355FFA4FC19D433FC453049" box="[968,1007,331,357]" captionStart="FIGURE 3" captionStartId="5.[151,250,1458,1482]" captionTargetBox="[162,1425,194,1434]" captionTargetId="figure-21@5.[151,1436,181,1434]" captionTargetPageId="5" captionText="FIGURE 3. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♀ allotype: habitus in dorsal (A) and lateral (B) views. Scale bars: 1 mm." pageId="6" pageNumber="200">3A</figureCitation>
): Anterior margin of prozona straight, without processes pointing anteriorly, at middle faintly emarginated. Median carina distinct throughout length of pronotum, flanked by a network of irregular carinae. Prozona carina distinct, more or less parallel; interhumeral carina distinct, faintly curved, diverging posteriorly. Posterior half of pronotal process rugose and punctuated. Apex narrow and subacute, with small emargination. Lower margin of lateral lobe of pronotum bluntly triangular with obtuse angle. Apical end of pronotal disc acute (
<figureCitation id="1321559CA355FFA4FD7CD487FCAD3335" box="[685,775,511,537]" captionStart="FIGURE 4" captionStartId="7.[151,250,1899,1923]" captionTargetBox="[164,1422,181,1875]" captionTargetId="figure-21@7.[164,1423,181,1875]" captionTargetPageId="7" captionText="FIGURE 4. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♂ holotype: face in anterior view (A); head and anterior half of pronotum in dorsal (B) and lateral (C) views; posterior half of pronotum in dorsal view (D); anterior (E) and middle (F) legs in external views; posterior femur in external (G) and dorsal (H) views; posterior tibia in external view (I). Scale bars: 1 mm." pageId="6" pageNumber="200">Fig. 4D</figureCitation>
). In lateral view (
<figureCitation id="1321559CA355FFA4FC16D487FB883335" box="[967,1058,511,537]" captionStart="FIGURE 2" captionStartId="4.[151,250,1803,1827]" captionTargetBox="[169,1417,813,1779]" captionTargetId="figure-196@4.[151,1436,813,1779]" captionTargetPageId="4" captionText="FIGURE 2. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♂ holotype: habitus in dorsal (A) and lateral (B) views. Scale bars: 1 mm." pageId="6" pageNumber="200">Figs 2B</figureCitation>
,
<figureCitation id="1321559CA355FFA4FBFCD487FBF83335" box="[1069,1106,511,537]" captionStart="FIGURE 3" captionStartId="5.[151,250,1458,1482]" captionTargetBox="[162,1425,194,1434]" captionTargetId="figure-21@5.[151,1436,181,1434]" captionTargetPageId="5" captionText="FIGURE 3. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♀ allotype: habitus in dorsal (A) and lateral (B) views. Scale bars: 1 mm." pageId="6" pageNumber="200">3B</figureCitation>
): with median carina strongly and irregularly undulated, most prominent at anterior half and highest between prozona and interhumeral carinae; prozona and interhumeral carinae roundly raised (
<figureCitation id="1321559CA355FFA4FD1CD73FFC8D334D" box="[717,807,583,609]" captionStart="FIGURE 4" captionStartId="7.[151,250,1899,1923]" captionTargetBox="[164,1422,181,1875]" captionTargetId="figure-21@7.[164,1423,181,1875]" captionTargetPageId="7" captionText="FIGURE 4. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♂ holotype: face in anterior view (A); head and anterior half of pronotum in dorsal (B) and lateral (C) views; posterior half of pronotum in dorsal view (D); anterior (E) and middle (F) legs in external views; posterior femur in external (G) and dorsal (H) views; posterior tibia in external view (I). Scale bars: 1 mm." pageId="6" pageNumber="200">Fig. 4C</figureCitation>
). At posterior half with two pairs of metamedian carina distinctly raised, the anterior ones higher than the posterior ones but not as high as interhumeral carina. Infrascapular area very broad, external lateral carina angularly sinuous. Angle of ventral sinus between lateral shoulder and lateral lobe deep and rounded.
</paragraph>
<paragraph id="8BA54919A355FFA4FF16D7AFFC0D358D" blockId="6.[151,1437,150,1941]" pageId="6" pageNumber="200">
Legs: Anterior and middle legs generally dark-brown coloured. Anterior femur with dorsal and ventral carinae undulated; dorsal carina forming two low nodules; ventral carina slightly distal of middle having triangular nodule with white apex and slightly basal of middle with nodule smaller and less distinct (
<figureCitation id="1321559CA355FFA4FBE1D667FB203215" box="[1072,1162,799,825]" captionStart="FIGURE 4" captionStartId="7.[151,250,1899,1923]" captionTargetBox="[164,1422,181,1875]" captionTargetId="figure-21@7.[164,1423,181,1875]" captionTargetPageId="7" captionText="FIGURE 4. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♂ holotype: face in anterior view (A); head and anterior half of pronotum in dorsal (B) and lateral (C) views; posterior half of pronotum in dorsal view (D); anterior (E) and middle (F) legs in external views; posterior femur in external (G) and dorsal (H) views; posterior tibia in external view (I). Scale bars: 1 mm." pageId="6" pageNumber="200">Figs 4E</figureCitation>
,
<figureCitation id="1321559CA355FFA4FB44D666FB103215" box="[1173,1210,798,825]" captionStart="FIGURE 5" captionStartId="8.[151,250,1908,1932]" captionTargetBox="[192,1396,181,1881]" captionTargetId="figure-17@8.[192,1396,181,1885]" captionTargetPageId="8" captionText="FIGURE 5. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♀ allotype: face in anterior view (A); head and anterior half of pronotum in dorsal (B) and lateral (C) views; anterior (D) and middle (E) legs in external views; posterior femur in dorsal (F) and external (G) views; posterior tibia in external view (H); abdominal apex in lateral (I) and ventral (J) views. Scale bars: 1 mm." pageId="6" pageNumber="200">5D</figureCitation>
). Anterior tibia dark coloured with three pale rings: one at base, large one at middle, small one just before apex (
<figureCitation id="1321559CA355FFA4FB78D63BFAAF3271" box="[1193,1285,835,861]" captionStart="FIGURE 4" captionStartId="7.[151,250,1899,1923]" captionTargetBox="[164,1422,181,1875]" captionTargetId="figure-21@7.[164,1423,181,1875]" captionTargetPageId="7" captionText="FIGURE 4. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♂ holotype: face in anterior view (A); head and anterior half of pronotum in dorsal (B) and lateral (C) views; posterior half of pronotum in dorsal view (D); anterior (E) and middle (F) legs in external views; posterior femur in external (G) and dorsal (H) views; posterior tibia in external view (I). Scale bars: 1 mm." pageId="6" pageNumber="200">Figs 4E</figureCitation>
,
<figureCitation id="1321559CA355FFA4FAC0D63AFA9C3271" box="[1297,1334,834,861]" captionStart="FIGURE 5" captionStartId="8.[151,250,1908,1932]" captionTargetBox="[192,1396,181,1881]" captionTargetId="figure-17@8.[192,1396,181,1885]" captionTargetPageId="8" captionText="FIGURE 5. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♀ allotype: face in anterior view (A); head and anterior half of pronotum in dorsal (B) and lateral (C) views; anterior (D) and middle (E) legs in external views; posterior femur in dorsal (F) and external (G) views; posterior tibia in external view (H); abdominal apex in lateral (I) and ventral (J) views. Scale bars: 1 mm." pageId="6" pageNumber="200">5D</figureCitation>
). Middle femur with dorsal and ventral carinae also undulating, forming each three weak peaks (
<figureCitation id="1321559CA355FFA4FBA4D61FFB6732AD" box="[1141,1229,871,897]" captionStart="FIGURE 4" captionStartId="7.[151,250,1899,1923]" captionTargetBox="[164,1422,181,1875]" captionTargetId="figure-21@7.[164,1423,181,1875]" captionTargetPageId="7" captionText="FIGURE 4. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♂ holotype: face in anterior view (A); head and anterior half of pronotum in dorsal (B) and lateral (C) views; posterior half of pronotum in dorsal view (D); anterior (E) and middle (F) legs in external views; posterior femur in external (G) and dorsal (H) views; posterior tibia in external view (I). Scale bars: 1 mm." pageId="6" pageNumber="200">Figs 4F</figureCitation>
,
<figureCitation id="1321559CA355FFA4FB0BD61EFB5632AD" box="[1242,1276,870,897]" captionStart="FIGURE 5" captionStartId="8.[151,250,1908,1932]" captionTargetBox="[192,1396,181,1881]" captionTargetId="figure-17@8.[192,1396,181,1885]" captionTargetPageId="8" captionText="FIGURE 5. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♀ allotype: face in anterior view (A); head and anterior half of pronotum in dorsal (B) and lateral (C) views; anterior (D) and middle (E) legs in external views; posterior femur in dorsal (F) and external (G) views; posterior tibia in external view (H); abdominal apex in lateral (I) and ventral (J) views. Scale bars: 1 mm." pageId="6" pageNumber="200">5E</figureCitation>
). Middle tibia not different from anterior tibiae (
<figureCitation id="1321559CA355FFA4FDDFD6F3FDCE3289" box="[526,612,907,933]" captionStart="FIGURE 4" captionStartId="7.[151,250,1899,1923]" captionTargetBox="[164,1422,181,1875]" captionTargetId="figure-21@7.[164,1423,181,1875]" captionTargetPageId="7" captionText="FIGURE 4. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♂ holotype: face in anterior view (A); head and anterior half of pronotum in dorsal (B) and lateral (C) views; posterior half of pronotum in dorsal view (D); anterior (E) and middle (F) legs in external views; posterior femur in external (G) and dorsal (H) views; posterior tibia in external view (I). Scale bars: 1 mm." pageId="6" pageNumber="200">Figs 4F</figureCitation>
,
<figureCitation id="1321559CA355FFA4FDBED6F2FD3B3289" box="[623,657,906,933]" captionStart="FIGURE 5" captionStartId="8.[151,250,1908,1932]" captionTargetBox="[192,1396,181,1881]" captionTargetId="figure-17@8.[192,1396,181,1885]" captionTargetPageId="8" captionText="FIGURE 5. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♀ allotype: face in anterior view (A); head and anterior half of pronotum in dorsal (B) and lateral (C) views; anterior (D) and middle (E) legs in external views; posterior femur in dorsal (F) and external (G) views; posterior tibia in external view (H); abdominal apex in lateral (I) and ventral (J) views. Scale bars: 1 mm." pageId="6" pageNumber="200">5E</figureCitation>
). Posterior femur about 2.52.6 times longer than wide, about 1.11.2 times longer than posterior tibia. Dorsal margin with clear undulating carinae (
<figureCitation id="1321559CA355FFA4FC2AD6D7FBFD32E5" box="[1019,1111,943,969]" captionStart="FIGURE 4" captionStartId="7.[151,250,1899,1923]" captionTargetBox="[164,1422,181,1875]" captionTargetId="figure-21@7.[164,1423,181,1875]" captionTargetPageId="7" captionText="FIGURE 4. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♂ holotype: face in anterior view (A); head and anterior half of pronotum in dorsal (B) and lateral (C) views; posterior half of pronotum in dorsal view (D); anterior (E) and middle (F) legs in external views; posterior femur in external (G) and dorsal (H) views; posterior tibia in external view (I). Scale bars: 1 mm." pageId="6" pageNumber="200">Figs 4G</figureCitation>
,
<figureCitation id="1321559CA355FFA4FBB3D6D6FB2232E5" box="[1122,1160,942,969]" captionStart="FIGURE 5" captionStartId="8.[151,250,1908,1932]" captionTargetBox="[192,1396,181,1881]" captionTargetId="figure-17@8.[192,1396,181,1885]" captionTargetPageId="8" captionText="FIGURE 5. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♀ allotype: face in anterior view (A); head and anterior half of pronotum in dorsal (B) and lateral (C) views; anterior (D) and middle (E) legs in external views; posterior femur in dorsal (F) and external (G) views; posterior tibia in external view (H); abdominal apex in lateral (I) and ventral (J) views. Scale bars: 1 mm." pageId="6" pageNumber="200">5G</figureCitation>
); in dorsal view showing three nodules, largest one at middle, smallest one slightly after middle (
<figureCitation id="1321559CA355FFA4FC6ED6ABFBB732C1" box="[959,1053,979,1005]" captionStart="FIGURE 4" captionStartId="7.[151,250,1899,1923]" captionTargetBox="[164,1422,181,1875]" captionTargetId="figure-21@7.[164,1423,181,1875]" captionTargetPageId="7" captionText="FIGURE 4. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♂ holotype: face in anterior view (A); head and anterior half of pronotum in dorsal (B) and lateral (C) views; posterior half of pronotum in dorsal view (D); anterior (E) and middle (F) legs in external views; posterior femur in external (G) and dorsal (H) views; posterior tibia in external view (I). Scale bars: 1 mm." pageId="6" pageNumber="200">Figs 4H</figureCitation>
,
<figureCitation id="1321559CA355FFA4FBF8D6AAFBE132C1" box="[1065,1099,978,1005]" captionStart="FIGURE 5" captionStartId="8.[151,250,1908,1932]" captionTargetBox="[192,1396,181,1881]" captionTargetId="figure-17@8.[192,1396,181,1885]" captionTargetPageId="8" captionText="FIGURE 5. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♀ allotype: face in anterior view (A); head and anterior half of pronotum in dorsal (B) and lateral (C) views; anterior (D) and middle (E) legs in external views; posterior femur in dorsal (F) and external (G) views; posterior tibia in external view (H); abdominal apex in lateral (I) and ventral (J) views. Scale bars: 1 mm." pageId="6" pageNumber="200">5F</figureCitation>
). Antegenicular and genicular teeth distinct, triangular; at apex antegenicular tooth rounded, genicular teeth acute (
<figureCitation id="1321559CA355FFA4FB91D68FFB37353D" box="[1088,1181,1015,1041]" captionStart="FIGURE 4" captionStartId="7.[151,250,1899,1923]" captionTargetBox="[164,1422,181,1875]" captionTargetId="figure-21@7.[164,1423,181,1875]" captionTargetPageId="7" captionText="FIGURE 4. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♂ holotype: face in anterior view (A); head and anterior half of pronotum in dorsal (B) and lateral (C) views; posterior half of pronotum in dorsal view (D); anterior (E) and middle (F) legs in external views; posterior femur in external (G) and dorsal (H) views; posterior tibia in external view (I). Scale bars: 1 mm." pageId="6" pageNumber="200">Figs 4G</figureCitation>
,
<figureCitation id="1321559CA355FFA4FB79D68EFB64353D" box="[1192,1230,1014,1041]" captionStart="FIGURE 5" captionStartId="8.[151,250,1908,1932]" captionTargetBox="[192,1396,181,1881]" captionTargetId="figure-17@8.[192,1396,181,1885]" captionTargetPageId="8" captionText="FIGURE 5. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♀ allotype: face in anterior view (A); head and anterior half of pronotum in dorsal (B) and lateral (C) views; anterior (D) and middle (E) legs in external views; posterior femur in dorsal (F) and external (G) views; posterior tibia in external view (H); abdominal apex in lateral (I) and ventral (J) views. Scale bars: 1 mm." pageId="6" pageNumber="200">5G</figureCitation>
). Posterior knee at apex nodular (
<figureCitation id="1321559CA355FFA4FEE7D163FE383519" box="[310,402,1051,1077]" captionStart="FIGURE 4" captionStartId="7.[151,250,1899,1923]" captionTargetBox="[164,1422,181,1875]" captionTargetId="figure-21@7.[164,1423,181,1875]" captionTargetPageId="7" captionText="FIGURE 4. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♂ holotype: face in anterior view (A); head and anterior half of pronotum in dorsal (B) and lateral (C) views; posterior half of pronotum in dorsal view (D); anterior (E) and middle (F) legs in external views; posterior femur in external (G) and dorsal (H) views; posterior tibia in external view (I). Scale bars: 1 mm." pageId="6" pageNumber="200">Figs 4G</figureCitation>
,
<figureCitation id="1321559CA355FFA4FE4CD162FE693519" box="[413,451,1050,1077]" captionStart="FIGURE 5" captionStartId="8.[151,250,1908,1932]" captionTargetBox="[192,1396,181,1881]" captionTargetId="figure-17@8.[192,1396,181,1885]" captionTargetPageId="8" captionText="FIGURE 5. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♀ allotype: face in anterior view (A); head and anterior half of pronotum in dorsal (B) and lateral (C) views; anterior (D) and middle (E) legs in external views; posterior femur in dorsal (F) and external (G) views; posterior tibia in external view (H); abdominal apex in lateral (I) and ventral (J) views. Scale bars: 1 mm." pageId="6" pageNumber="200">5G</figureCitation>
). Posterior tibia at base dark coloured, becoming slightly lighter in colouration; with five spines on each lateral margin, with numerous spinules between these large dorsal spines (
<figureCitation id="1321559CA355FFA4FBAAD147FB673575" box="[1147,1229,1087,1113]" captionStart="FIGURE 4" captionStartId="7.[151,250,1899,1923]" captionTargetBox="[164,1422,181,1875]" captionTargetId="figure-21@7.[164,1423,181,1875]" captionTargetPageId="7" captionText="FIGURE 4. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♂ holotype: face in anterior view (A); head and anterior half of pronotum in dorsal (B) and lateral (C) views; posterior half of pronotum in dorsal view (D); anterior (E) and middle (F) legs in external views; posterior femur in external (G) and dorsal (H) views; posterior tibia in external view (I). Scale bars: 1 mm." pageId="6" pageNumber="200">Figs 4I</figureCitation>
,
<figureCitation id="1321559CA355FFA4FB09D146FB543575" box="[1240,1278,1086,1113]" captionStart="FIGURE 5" captionStartId="8.[151,250,1908,1932]" captionTargetBox="[192,1396,181,1881]" captionTargetId="figure-17@8.[192,1396,181,1885]" captionTargetPageId="8" captionText="FIGURE 5. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♀ allotype: face in anterior view (A); head and anterior half of pronotum in dorsal (B) and lateral (C) views; anterior (D) and middle (E) legs in external views; posterior femur in dorsal (F) and external (G) views; posterior tibia in external view (H); abdominal apex in lateral (I) and ventral (J) views. Scale bars: 1 mm." pageId="6" pageNumber="200">5H</figureCitation>
). Basal article of posterior tarsus generally pale coloured, 0.91.2 times as long as apical article; middle article dark coloured and very short; apical article white, except apex being black (
<figureCitation id="1321559CA355FFA4FCC7D1FFFCC3358D" box="[790,873,1159,1185]" captionStart="FIGURE 4" captionStartId="7.[151,250,1899,1923]" captionTargetBox="[164,1422,181,1875]" captionTargetId="figure-21@7.[164,1423,181,1875]" captionTargetPageId="7" captionText="FIGURE 4. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♂ holotype: face in anterior view (A); head and anterior half of pronotum in dorsal (B) and lateral (C) views; posterior half of pronotum in dorsal view (D); anterior (E) and middle (F) legs in external views; posterior femur in external (G) and dorsal (H) views; posterior tibia in external view (I). Scale bars: 1 mm." pageId="6" pageNumber="200">Figs 4I</figureCitation>
,
<figureCitation id="1321559CA355FFA4FCA5D1FEFC31358D" box="[884,923,1158,1185]" captionStart="FIGURE 5" captionStartId="8.[151,250,1908,1932]" captionTargetBox="[192,1396,181,1881]" captionTargetId="figure-17@8.[192,1396,181,1885]" captionTargetPageId="8" captionText="FIGURE 5. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♀ allotype: face in anterior view (A); head and anterior half of pronotum in dorsal (B) and lateral (C) views; anterior (D) and middle (E) legs in external views; posterior femur in dorsal (F) and external (G) views; posterior tibia in external view (H); abdominal apex in lateral (I) and ventral (J) views. Scale bars: 1 mm." pageId="6" pageNumber="200">5H</figureCitation>
).
</paragraph>
<paragraph id="8BA54919A355FFA4FF16D1D3FB5135E9" blockId="6.[151,1437,150,1941]" box="[199,1275,1195,1221]" pageId="6" pageNumber="200">Abdomen: Tergites and sternites typically dark coloured with some sparse patches of pale spots.</paragraph>
<paragraph id="8BA54919A355FFA4FF16D1B7FDAB3421" blockId="6.[151,1437,150,1941]" pageId="6" pageNumber="200">
<emphasis id="B96E950BA355FFA4FF16D1B7FEA135C5" bold="true" box="[199,267,1231,1257]" pageId="6" pageNumber="200">Male.</emphasis>
Area between prozona carinae distinctly cream-coloured, colouration extending laterally into lateral lobe anterior of sulcus (
<figureCitation id="1321559CA355FFA4FEB9D18BFE5F3421" box="[360,501,1267,1293]" captionStart="FIGURE 4" captionStartId="7.[151,250,1899,1923]" captionTargetBox="[164,1422,181,1875]" captionTargetId="figure-21@7.[164,1423,181,1875]" captionTargetPageId="7" captionText="FIGURE 4. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♂ holotype: face in anterior view (A); head and anterior half of pronotum in dorsal (B) and lateral (C) views; posterior half of pronotum in dorsal view (D); anterior (E) and middle (F) legs in external views; posterior femur in external (G) and dorsal (H) views; posterior tibia in external view (I). Scale bars: 1 mm." pageId="6" pageNumber="200">Figs 4B, 4C</figureCitation>
).
</paragraph>
<paragraph id="8BA54919A355FFA4FF16D06FFC823455" blockId="6.[151,1437,150,1941]" pageId="6" pageNumber="200">
<emphasis id="B96E950BA355FFA4FF16D06FFE69341C" bold="true" box="[199,451,1302,1329]" pageId="6" pageNumber="200">
Female (
<figureCitation id="1321559CA355FFA4FEFED06EFE1F341C" box="[303,437,1302,1329]" captionStart="FIGURE 5" captionStartId="8.[151,250,1908,1932]" captionTargetBox="[192,1396,181,1881]" captionTargetId="figure-17@8.[192,1396,181,1885]" captionTargetPageId="8" captionText="FIGURE 5. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♀ allotype: face in anterior view (A); head and anterior half of pronotum in dorsal (B) and lateral (C) views; anterior (D) and middle (E) legs in external views; posterior femur in dorsal (F) and external (G) views; posterior tibia in external view (H); abdominal apex in lateral (I) and ventral (J) views. Scale bars: 1 mm." pageId="6" pageNumber="200">Figs 5I, 5J</figureCitation>
).
</emphasis>
Cercus tiny (
<figureCitation id="1321559CA355FFA4FDB0D06FFD18341D" box="[609,690,1302,1329]" captionStart="FIGURE 5" captionStartId="8.[151,250,1908,1932]" captionTargetBox="[192,1396,181,1881]" captionTargetId="figure-17@8.[192,1396,181,1885]" captionTargetPageId="8" captionText="FIGURE 5. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♀ allotype: face in anterior view (A); head and anterior half of pronotum in dorsal (B) and lateral (C) views; anterior (D) and middle (E) legs in external views; posterior femur in dorsal (F) and external (G) views; posterior tibia in external view (H); abdominal apex in lateral (I) and ventral (J) views. Scale bars: 1 mm." pageId="6" pageNumber="200">Fig. 5I</figureCitation>
). Subgenital plate elongated, at middle with distinct longitudinal carina, broadest at middle, gently tapering into broadly truncated apex; at middle of apex with rounded lobule and some setae (
<figureCitation id="1321559CA355FFA4FEF0D027FEDB3455" box="[289,369,1374,1401]" captionStart="FIGURE 5" captionStartId="8.[151,250,1908,1932]" captionTargetBox="[192,1396,181,1881]" captionTargetId="figure-17@8.[192,1396,181,1885]" captionTargetPageId="8" captionText="FIGURE 5. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♀ allotype: face in anterior view (A); head and anterior half of pronotum in dorsal (B) and lateral (C) views; anterior (D) and middle (E) legs in external views; posterior femur in dorsal (F) and external (G) views; posterior tibia in external view (H); abdominal apex in lateral (I) and ventral (J) views. Scale bars: 1 mm." pageId="6" pageNumber="200">Fig. 5J</figureCitation>
). Ovipositor long and slender (
<figureCitation id="1321559CA355FFA4FD1DD027FCB13455" box="[716,795,1374,1401]" captionStart="FIGURE 5" captionStartId="8.[151,250,1908,1932]" captionTargetBox="[192,1396,181,1881]" captionTargetId="figure-17@8.[192,1396,181,1885]" captionTargetPageId="8" captionText="FIGURE 5. Fartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. ♀ allotype: face in anterior view (A); head and anterior half of pronotum in dorsal (B) and lateral (C) views; anterior (D) and middle (E) legs in external views; posterior femur in dorsal (F) and external (G) views; posterior tibia in external view (H); abdominal apex in lateral (I) and ventral (J) views. Scale bars: 1 mm." pageId="6" pageNumber="200">Fig. 5I</figureCitation>
).
</paragraph>
<paragraph id="8BA54919A355FFA4FF16D0FBFBA234C9" blockId="6.[151,1437,150,1941]" pageId="6" pageNumber="200">
<emphasis id="B96E950BA355FFA4FF16D0FBFE4234B0" bold="true" box="[199,488,1410,1437]" pageId="6" pageNumber="200">Measurements (in mm).</emphasis>
<specimenCount id="9D1C8290A355FFA4FE20D0FBFDAC34B1" box="[497,518,1411,1437]" count="1" pageId="6" pageNumber="200" type="male"></specimenCount>
<typeStatus id="54A1F7BBA355FFA4FDDED0FBFDDF34B1" box="[527,629,1411,1437]" pageId="6" pageNumber="200" type="holotype">holotype</typeStatus>
: BL = 7.9; VW = 0.80; EW = 0.53; SW = 0.21; MAL =?; MAW =?; 1
<superScript id="7C6FE451A355FFA4FF74D0DFFF7634ED" attach="right" box="[165,220,1447,1473]" fontSize="6" pageId="6" pageNumber="200">stAW</superScript>
= 0.27; PL = 6.9; PW = 3.8;
<collectingCountry id="F30D0989A355FFA4FDCDD0DFFDEA34ED" box="[540,576,1447,1473]" name="Philippines" pageId="6" pageNumber="200">PH</collectingCountry>
= 2.3; IAH = 0.4; PAW = 2.1; PPW = 0.2; FFL = 2.0; FFW = 0.6; MFL = 2.1; MFW = 0.6; HFL = 4.5; HFW = 1.8; HTL = 3.8; 1
<superScript id="7C6FE451A355FFA4FD1CD0B3FD7234F5" attach="left" box="[717,728,1483,1497]" fontSize="6" pageId="6" pageNumber="200">st</superScript>
TaL = 0.69; 3
<superScript id="7C6FE451A355FFA4FCA4D0B2FC2934F4" attach="left" box="[885,899,1482,1496]" fontSize="6" pageId="6" pageNumber="200">rd</superScript>
TaL = 0.73.
</paragraph>
<paragraph id="8BA54919A355FFA4FF16D097FDAE3759" blockId="6.[151,1437,150,1941]" pageId="6" pageNumber="200">
<materialsCitation id="3B724344A355FFA4FF16D097FAFE3725" box="[199,1364,1518,1545]" collectionCode="PL" pageId="6" pageNumber="200" specimenCount="1" specimenCount-female="1" typeStatus="allotype">
<specimenCount id="9D1C8290A355FFA4FF16D097FF763725" box="[199,220,1519,1545]" count="1" pageId="6" pageNumber="200" type="female"></specimenCount>
<typeStatus id="54A1F7BBA355FFA4FF32D097FEE83725" box="[227,322,1519,1545]" pageId="6" pageNumber="200" type="allotype">allotype</typeStatus>
: BL = 8.7; VW = 0.85; EW = 0.51; SW = 0.24; MAL = 0.50; MAW = 0.07; 1
<superScript id="7C6FE451A355FFA4FB70D097FB723725" attach="right" box="[1185,1240,1519,1545]" fontSize="6" pageId="6" pageNumber="200">stAW</superScript>
= 0.28;
<collectionCode id="ED0BD1DCA355FFA4FAE5D097FAFE3725" box="[1332,1364,1519,1545]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="6" pageNumber="200" type="Museum">PL</collectionCode>
</materialsCitation>
= 7.8; PW =
<materialsCitation id="3B724344A355FFA4FF31D36BFB543701" box="[224,1278,1555,1581]" collectionCode="MFW" country="Philippines" location="PH" pageId="6" pageNumber="200" specimenCount="1" typeStatus="allotype">
4.0;
<collectingCountry id="F30D0989A355FFA4FEC2D36BFE9D3701" box="[275,311,1555,1581]" name="Philippines" pageId="6" pageNumber="200">PH</collectingCountry>
= 2.4; IAH = 0.4; PAW = 2.2; PPW = 0.3; FFL = 2.2; FFW = 0.6; MFL = 2.4;
<collectionCode id="ED0BD1DCA355FFA4FB6DD36BFB543701" box="[1212,1278,1555,1581]" country="0" httpUri="http://grbio.org/cool/j70n-nrhw" name="Museum Freriks" pageId="6" pageNumber="200">MFW</collectionCode>
</materialsCitation>
= 0.6; HFL = 5.0; HFW = 1.9; HTL = 4.4; 1
<superScript id="7C6FE451A355FFA4FE20D34FFE563769" attach="left" box="[497,508,1591,1605]" fontSize="6" pageId="6" pageNumber="200">st</superScript>
TaL = 0.86; 3
<superScript id="7C6FE451A355FFA4FD4FD34EFD063768" attach="left" box="[670,684,1590,1604]" fontSize="6" pageId="6" pageNumber="200">rd</superScript>
TaL = 0.74; SGPL = 2.25; SGPW = 1.56; OVDL = 2.7; OVDW = 0.6; OVVL = 2.8; OVVW = 0.4.
</paragraph>
</subSubSection>
<subSubSection id="C3001A92A355FFA4FF16D307FE2D3605" pageId="6" pageNumber="200" type="biology_ecology">
<paragraph id="8BA54919A355FFA4FF16D307FE2D3605" blockId="6.[151,1437,150,1941]" pageId="6" pageNumber="200">
<emphasis id="B96E950BA355FFA4FF16D307FE8637B5" bold="true" box="[199,300,1663,1689]" pageId="6" pageNumber="200">Ecology.</emphasis>
The species was found dwelling and well-camouflaged among wet decaying Abacá (
<taxonomicName id="4C1A329AA355FFA4FB21D307FA2B37B5" box="[1264,1409,1663,1689]" class="Liliopsida" family="Musaceae" genus="Musa" kingdom="Plantae" order="Zingiberales" pageId="6" pageNumber="200" phylum="Tracheophyta" rank="species" species="textilis">
<emphasis id="B96E950BA355FFA4FB21D307FA2B37B5" box="[1264,1409,1663,1689]" italics="true" pageId="6" pageNumber="200">Musa textilis</emphasis>
</taxonomicName>
, a banana species endemic to the
<collectingCountry id="F30D0989A355FFA4FE3FD3DBFDC43791" box="[494,622,1699,1725]" name="Philippines" pageId="6" pageNumber="200">Philippines</collectingCountry>
) stem and leaves on the ground among abandoned Abacá plantation (
<figureCitation id="1321559CA355FFA4FABED3DBFF0D37CD" captionStart="FIGURE6" captionStartId="9.[151,250,1058,1082]" captionTargetBox="[151,1436,181,1034]" captionTargetId="figure-21@9.[151,1436,181,1034]" captionTargetPageId="9" captionText="FIGURE6. AbandonedAbacáplantation nearLakeLinao (A) andwetdecayingAbacástemsandleaves (B) whereFartmanntettix undulatus Tan &amp; Tumbrinck, gen. et sp. nov. were found." pageId="6" pageNumber="200">Fig. 6</figureCitation>
). The species can also be found to coexist with other
<taxonomicName id="4C1A329AA355FFA4FD27D3BFFCC037CD" box="[758,874,1735,1761]" class="Insecta" family="Tetrigidae" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="200" phylum="Arthropoda" rank="family">Tetrigidae</taxonomicName>
, including unknown members of
<taxonomicName id="4C1A329AA355FFA4FB0CD3BFFEEA3629" authority="Gunther, 1937" authorityName="Gunther" authorityYear="1937" class="Insecta" family="Tetrigidae" genus="Pseudoparatettix" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="200" phylum="Arthropoda" rank="genus">
<emphasis id="B96E950BA355FFA4FB0CD3BFFA3637CD" box="[1245,1436,1735,1761]" italics="true" pageId="6" pageNumber="200">Pseudoparatettix</emphasis>
Günther, 1937
</taxonomicName>
,
<taxonomicName id="4C1A329AA355FFA4FE80D393FE753629" authority="Bolivar, 1887" authorityName="Bolivar" authorityYear="1887" box="[337,479,1771,1797]" class="Insecta" family="Tetrigidae" genus="Systolederus" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="200" phylum="Arthropoda" rank="genus">
<emphasis id="B96E950BA355FFA4FE80D393FE753629" box="[337,479,1771,1797]" italics="true" pageId="6" pageNumber="200">Systolederus</emphasis>
</taxonomicName>
? Bolívar, 1887,
<taxonomicName id="4C1A329AA355FFA4FD78D393FC873629" authority="Hancock, 1909" authorityName="Hancock" authorityYear="1909" box="[681,813,1771,1797]" class="Insecta" family="Tetrigidae" genus="Thoradonta" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="200" phylum="Arthropoda" rank="genus">
<emphasis id="B96E950BA355FFA4FD78D393FC873629" box="[681,813,1771,1797]" italics="true" pageId="6" pageNumber="200">Thoradonta</emphasis>
</taxonomicName>
? Hancock, 1909,
<taxonomicName id="4C1A329AA355FFA4FBD9D393FB5D3629" authority="Bolivar, 1887" authorityName="Bolivar" authorityYear="1887" box="[1032,1271,1771,1797]" class="Insecta" family="Tetrigidae" genus="Xistra" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="200" phylum="Arthropoda" rank="genus">
<emphasis id="B96E950BA355FFA4FBD9D393FBE63628" box="[1032,1100,1771,1796]" italics="true" pageId="6" pageNumber="200">Xistra</emphasis>
Bolívar, 1887
</taxonomicName>
and
<taxonomicName id="4C1A329AA355FFA4FAE8D393FE293605" authority="Stal, 1877" authorityName="Stal" authorityYear="1877" class="Insecta" family="Tetrigidae" genus="Diotarus" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="200" phylum="Arthropoda" rank="species" species="verrucifer">
<emphasis id="B96E950BA355FFA4FAE8D393FEAD3605" italics="true" pageId="6" pageNumber="200">Diotarus verrucifer</emphasis>
Stål, 1877
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3001A92A355FFA4FF16D24BFC97365D" pageId="6" pageNumber="200" type="materials_examined">
<paragraph id="8BA54919A355FFA4FF16D24BFC97365D" blockId="6.[151,1437,150,1941]" pageId="6" pageNumber="200">
<emphasis id="B96E950BA355FFA4FF16D24BFEC23660" bold="true" box="[199,360,1842,1869]" pageId="6" pageNumber="200">Type locality.</emphasis>
<collectingCountry id="F30D0989A355FFA4FEA0D24BFDBC3661" box="[369,534,1843,1869]" name="Philippines" pageId="6" pageNumber="200">PHILIPPINES</collectingCountry>
(Mindanao Island:
<collectingRegion id="49DE87FBA355FFA4FD28D24BFBE63661" box="[761,1100,1843,1869]" country="Philippines" name="Misamis Occidental" pageId="6" pageNumber="200">Misamis Occidental province</collectingRegion>
: Don Victoriano Chiongbian: Lake Duminagat Barangay: Mount Malindang Natural Park)
</paragraph>
</subSubSection>
<subSubSection id="C3001A92A355FFA4FF16D202FBF836B9" box="[199,1106,1914,1941]" pageId="6" pageNumber="200" type="distribution">
<paragraph id="8BA54919A355FFA4FF16D202FBF836B9" blockId="6.[151,1437,150,1941]" box="[199,1106,1914,1941]" pageId="6" pageNumber="200">
<emphasis id="B96E950BA355FFA4FF16D202FECB36B8" bold="true" box="[199,353,1914,1940]" pageId="6" pageNumber="200">Distribution.</emphasis>
<collectingCountry id="F30D0989A355FFA4FEB6D203FDA636B9" box="[359,524,1915,1941]" name="Philippines" pageId="6" pageNumber="200">PHILIPPINES</collectingCountry>
(Mindanao Island: Mount Malindang Natural Park)
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="D0474CB22A916B02E50BB8010FFF2A0F">Turtles and tortoises of Syria: Diversity, distribution, and conservation</mods:title>
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<mods:name id="FF7DEE20E94E1ADA4C9EEACCCE0FFDAA" type="personal">
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<mods:namePart id="9D9B6312B89C96CEAFB62FD2DC107789">Aidek, Ahmad E.</mods:namePart>
<mods:affiliation id="AA58443449C9971E4A65365D5159BEF4">General Commission for Scientific Agricultural Research, Deir ez-Zor, Syria</mods:affiliation>
</mods:name>
<mods:name id="7B5E35E938BBE216C81CA5FA0A1CACD8" type="personal">
<mods:role id="DD14502B09E80DB8255B50B0F89DC8F7">
<mods:roleTerm id="14399B0E51C45665FD0573857717008E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="04C73901A10BE555328136F38467ACD3">Saad, Adib</mods:namePart>
<mods:affiliation id="86C21B6CA3F69BAF4F30DD8605B3D55D">Al-Manara University, Latakia, Syria</mods:affiliation>
</mods:name>
<mods:name id="4158EF6B887AE788CFC52FD4F246FE47" type="personal">
<mods:role id="4EFC2D9F5417C761EC34868CE754254B">
<mods:roleTerm id="DF4DDD4AADD767D93B3E8C6601580C38">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="05F9A835C57DB6AFF0E4730C013D926F">Jablonski, Daniel</mods:namePart>
<mods:affiliation id="E7FC52196D0DDD535D38DF5CB382AA4B">Department of Zoology, Comenius University in Bratislava, Ilkovičova 6, Mlynská dolina, 842 15 Bratislava, Slovakia</mods:affiliation>
</mods:name>
<mods:name id="82D683832E20D493BF5042A2480E6AAA" type="personal">
<mods:role id="5BA11D6C20C02A377667AF90C96AD7F3">
<mods:roleTerm id="7E25F80A8DADA08019647B292A81434D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5651047FF24786F1034522E2723789F2">Esterbauer, Hans</mods:namePart>
<mods:affiliation id="18B24677D09A8ACE713D02A0646511B9">Johann-Puch-Strasse 27 / III / 5, 4400 Steyr, Austria</mods:affiliation>
</mods:name>
<mods:name id="A0D6FFDDD220453665855E6CA8101B57" type="personal">
<mods:role id="E6B2B62F2849FBD1200F24EAD2E9E5EE">
<mods:roleTerm id="6E716E1BC1EEE05D24BC9908F2682AE3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="60539773E11F9AAF74BAC25C231FAC14">Fritz, Uwe</mods:namePart>
<mods:affiliation id="4878A8A1D9010FE6D7211D2D8703E9EC">Museum of Zoology, Senckenberg Dresden, A. B. Meyer Building, 01109 Dresden, Germany</mods:affiliation>
</mods:name>
<mods:typeOfResource id="D8B5561B3FB028E0CB00842473CE99EE">text</mods:typeOfResource>
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</mods:titleInfo>
<mods:part id="1751313AC10ABAC8BB8D90CB43CFE076">
<mods:date id="4739D5BE00C941677AFC999FAA479CF1">2024</mods:date>
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<mods:number id="FF64E8DD2DB409292B8953E61374B8B4">2024-09-06</mods:number>
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<mods:number id="3FB9898807143A79447729FD86ABD586">5506</mods:number>
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<mods:start id="A1F7A50119DD96534348190BDC1DDBAB">151</mods:start>
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</mods:location>
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</mods:mods>
<treatment id="03B9F52AC722FF8304B62DBDF108F875" LSID="urn:lsid:plazi:treatment:03B9F52AC722FF8304B62DBDF108F875" httpUri="http://treatment.plazi.org/id/03B9F52AC722FF8304B62DBDF108F875" lastPageNumber="158" pageId="7" pageNumber="158">
<subSubSection id="C30A17B7C722FF8304B62DBDF5F9FAA0" pageId="7" pageNumber="158" type="nomenclature">
<paragraph id="8BAF443CC722FF8304B62DBDF62BFB5A" blockId="7.[151,547,1237,1301]" box="[151,547,1237,1264]" pageId="7" pageNumber="158">
<heading id="D0E7F350C722FF8304B62DBDF62BFB5A" bold="true" box="[151,547,1237,1264]" fontSize="11" level="1" pageId="7" pageNumber="158" reason="1">
<taxonomicName id="4C103FBFC722FF8304B62DBDF62BFB5A" ID-CoL="69BHK" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[151,547,1237,1264]" class="Testudines" family="Cheloniidae" genus="Caretta" kingdom="Animalia" order="Cryptodira" pageId="7" pageNumber="158" phylum="Chordata" rank="species" species="caretta">
<emphasis id="B964982EC722FF8304B62DBDF62BFB5A" bold="true" box="[151,547,1237,1264]" pageId="7" pageNumber="158">
<emphasis id="B964982EC722FF8304B62DBDF543FB45" bold="true" box="[151,331,1238,1264]" italics="true" pageId="7" pageNumber="158">Caretta caretta</emphasis>
(Linnaeus, 1758)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BAF443CC722FF8304B62D91F5F9FAA0" blockId="7.[151,547,1237,1301]" box="[151,497,1274,1301]" pageId="7" pageNumber="158">
Loggerhead Sea Turtle (
<figureCitation id="132B58B9C722FF8305872D91F5E0FAA0" box="[422,488,1274,1301]" captionStart="FIGURE 5" captionStartId="8.[151,250,961,985]" captionTargetBox="[227,1361,181,937]" captionTargetId="figure-21@8.[227,1361,181,937]" captionTargetPageId="8" captionText="FIGURE 5. Loggerhead sea turtle, Caretta caretta off Latakia Beach (photo: Alan F. Rees/ARCHELON)." pageId="7" pageNumber="158">Fig. 5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30A17B7C722FF8304B62C2AF118FAE9" box="[151,1296,1345,1372]" pageId="7" pageNumber="158" type="description">
<paragraph id="8BAF443CC722FF8304B62C2AF118FAE9" blockId="7.[151,1436,1345,1984]" box="[151,1296,1345,1372]" pageId="7" pageNumber="158">
<emphasis id="B964982EC722FF8304B62C2AF5C9FAEE" bold="true" box="[151,449,1345,1372]" pageId="7" pageNumber="158">IUCN Red List Category</emphasis>
: Vulnerable (VU A2b); Mediterranean subpopulation: Least Concern (LC).
</paragraph>
</subSubSection>
<subSubSection id="C30A17B7C722FF8304E62C0DF70FF981" pageId="7" pageNumber="158" type="materials_examined">
<paragraph id="8BAF443CC722FF8304E62C0DF16AFA10" blockId="7.[151,1436,1345,1984]" pageId="7" pageNumber="158">
<emphasis id="B964982EC722FF8304E62C0DF56CFACA" bold="true" box="[199,356,1381,1408]" pageId="7" pageNumber="158">
<typeStatus id="54ABFA9EC722FF8304E62C0DF50AFA35" box="[199,258,1382,1408]" pageId="7" pageNumber="158">Type</typeStatus>
locality
</emphasis>
: “Insulas Americanas.” Restricted to “Mari Mediterraneo, Atlantico” by
<bibRefCitation id="EF8139CDC722FF8300982C0DF17FFA35" author="Schoepff, J. D." box="[1209,1399,1382,1408]" pageId="7" pageNumber="158" refId="ref19177" refString="Schoepff, J. D. (1793) Historia Testudinum Iconibus Illustrata. Fasciculus III et IV. Joannes Jacobus Palm, Erlangen, pp. 33 - 80, pls. XI - XVI + XVIIB - XX." type="book" year="1793">Schoepff (1793)</bibRefCitation>
; to “
<collectingCountry id="F30704ACC722FF8304802CE1F502FA11" box="[161,266,1418,1444]" name="Bermuda" pageId="7" pageNumber="158">Bermuda</collectingCountry>
Islands” by
<bibRefCitation id="EF8139CDC722FF8305B62CE1F695FA10" author="Smith, H. M. &amp; Taylor, E. H." box="[407,669,1418,1445]" pageId="7" pageNumber="158" pagination="313 - 380" refId="ref19489" refString="Smith, H. M. &amp; Taylor, E. H. (1950 a) Type localities of Mexican reptiles and amphibians. University of Kansas Science Bulletin, 33, 313 - 380." type="journal article" year="1950">
<collectingRegion id="49D48ADEC722FF8305B62CE1F5D2FA11" box="[407,474,1418,1444]" country="Bermuda" name="Smith's" pageId="7" pageNumber="158">Smith</collectingRegion>
&amp; Taylor (1950a
</bibRefCitation>
,
<bibRefCitation id="EF8139CDC722FF8306882CE1F6FBFA10" author="Smith, H. M. &amp; Taylor, E. H." box="[681,755,1418,1445]" pageId="7" pageNumber="158" pagination="1 - 253" refId="ref19526" refString="Smith, H. M. &amp; Taylor, E. H. (1950 b) An annotated checklist and key to the reptiles of Mexico exclusive of the snakes. United States National Museum Bulletin, 199, 1 - 253. https: // doi. org / 10.5479 / si. 03629236.199" type="journal article" year="1950">1950b</bibRefCitation>
); and to “
<collectingRegion id="49D48ADEC722FF83077C2CE1F7B8FA11" box="[861,944,1418,1444]" country="Bahamas" name="Bimini" pageId="7" pageNumber="158">Bimini</collectingRegion>
, British
<collectingCountry id="F30704ACC722FF83002F2CE1F07FFA11" box="[1038,1143,1418,1444]" name="Bahamas" pageId="7" pageNumber="158">Bahamas</collectingCountry>
” by
<bibRefCitation id="EF8139CDC722FF83008D2CE1F156FA10" author="Schmidt, K. P." box="[1196,1374,1418,1445]" pageId="7" pageNumber="158" refId="ref19143" refString="Schmidt, K. P. (1953) A Check List of North American Amphibians and Reptiles. Sixth Edition. University of Chicago Press, Chicago, Illinois, 280 pp." type="book" year="1953">Schmidt (1953)</bibRefCitation>
.
</paragraph>
<paragraph id="8BAF443CC722FF8304E62CC5F60FFA7D" blockId="7.[151,1436,1345,1984]" box="[199,519,1453,1480]" pageId="7" pageNumber="158">
<emphasis id="B964982EC722FF8304E62CC5F58AFA72" bold="true" box="[199,386,1453,1480]" pageId="7" pageNumber="158">
<typeStatus id="54ABFA9EC722FF8304E62CC5F50AFA7D" box="[199,258,1454,1480]" pageId="7" pageNumber="158">Type</typeStatus>
specimens
</emphasis>
: Unknown.
</paragraph>
<paragraph id="8BAF443CC722FF8304E62CBAF6A5FA59" blockId="7.[151,1436,1345,1984]" box="[199,685,1489,1516]" pageId="7" pageNumber="158">
<emphasis id="B964982EC722FF8304E62CBAF54EFA5E" bold="true" box="[199,326,1489,1515]" pageId="7" pageNumber="158">Chorotype</emphasis>
: Circumglobal (Cosmopolitan).
</paragraph>
<paragraph id="8BAF443CC722FF8304E62C9EF194F9A4" blockId="7.[151,1436,1345,1984]" box="[199,1436,1525,1553]" pageId="7" pageNumber="158">
<emphasis id="B964982EC722FF8304E62C9EF5DBF9BA" bold="true" box="[199,467,1525,1552]" pageId="7" pageNumber="158">
First records for
<collectingCountry id="F30704ACC722FF8305B22C9EF5DBF9BA" box="[403,467,1525,1551]" name="Syria" pageId="7" pageNumber="158">Syria</collectingCountry>
</emphasis>
: The rocky coasts of Syria (
<bibRefCitation id="EF8139CDC722FF83072F2C9DF79EF9A5" author="Lortet, L." box="[782,918,1526,1552]" pageId="7" pageNumber="158" pagination="99 - 194" refId="ref16560" refString="Lortet, L. (1883) Poissons et reptiles du Lac de Tiberiade et de quelques autres parties de la Syrie. Archives du Museum d'histoire naturelle de Lyon, 3, 99 - 194. https: // doi. org / 10.3406 / mhnly. 1883.941" type="journal article" year="1883">Lortet 1883</bibRefCitation>
), between
<collectingRegion id="49D48ADEC722FF83002E2C9DF06BF9A5" box="[1039,1123,1526,1552]" country="Syria" name="Al Ladhiqiyah" pageId="7" pageNumber="158">Latakia</collectingRegion>
and Jabla (
<bibRefCitation id="EF8139CDC722FF8300C02C9DF198F9A4" author="Kasparek, M." box="[1249,1424,1526,1553]" pageId="7" pageNumber="158" pagination="51 - 62" refId="ref16224" refString="Kasparek, M. (1995) The nesting of marine turtles on the coast of Syria. Zoology in the Middle East, 11, 51 - 62. https: // doi. org / 10.1080 / 09397140.1995.10637671" type="journal article" year="1995">Kasparek 1995</bibRefCitation>
).
</paragraph>
<paragraph id="8BAF443CC722FF8304E62F72F70FF981" blockId="7.[151,1436,1345,1984]" box="[199,775,1561,1588]" pageId="7" pageNumber="158">
<emphasis id="B964982EC722FF8304E62F72F5C6F981" bold="true" box="[199,462,1561,1588]" pageId="7" pageNumber="158">Citizen science record</emphasis>
:
<collectingRegion id="49D48ADEC722FF8305FD2F71F638F981" box="[476,560,1562,1588]" country="Syria" name="Al Ladhiqiyah" pageId="7" pageNumber="158">Latakia</collectingRegion>
(iNaturalist 2023).
</paragraph>
</subSubSection>
<subSubSection id="C30A17B7C722FF8304E62F56F16CF8E1" pageId="7" pageNumber="158" type="distribution">
<paragraph id="8BAF443CC722FF8304E62F56F16CF8E1" blockId="7.[151,1436,1345,1984]" pageId="7" pageNumber="158">
<emphasis id="B964982EC722FF8304E62F56F5B7F9E2" bold="true" box="[199,447,1597,1623]" pageId="7" pageNumber="158">
Distribution in
<collectingCountry id="F30704ACC722FF83055E2F56F5B7F9E2" box="[383,447,1597,1623]" name="Syria" pageId="7" pageNumber="158">Syria</collectingCountry>
</emphasis>
: Foraging and nesting in Syria.
<taxonomicName id="4C103FBFC722FF8307082F55F7DAF9ED" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[809,978,1598,1624]" class="Testudines" family="Cheloniidae" genus="Caretta" kingdom="Animalia" order="Cryptodira" pageId="7" pageNumber="158" phylum="Chordata" rank="species" species="caretta">
<emphasis id="B964982EC722FF8307082F55F7DAF9ED" box="[809,978,1598,1624]" italics="true" pageId="7" pageNumber="158">Caretta caretta</emphasis>
</taxonomicName>
is found throughout the year in different size classes in marine waters off the coast. Some dead turtles were also seen on at the beaches of ash-Shati Al-Azraq, Ugarit, Banias, Shati Ishtar, Shati Al-Dubbat, Al-Khrab, and Ras Al-Baseet. Nesting has been reported for sandy beaches in or near (1) Ras Al-Basit, (2) Um et-Toyour, (3) Wadi Kandil, (4)
<collectingRegion id="49D48ADEC722FF8307C12FC1F03CF971" box="[992,1076,1706,1732]" country="Syria" name="Al Ladhiqiyah" pageId="7" pageNumber="158">Latakia</collectingRegion>
beach (Shkaifat and Sanawbar), (5) Banias, (6) south of
<collectingRegion id="49D48ADEC722FF8305802FA5F5FDF95D" box="[417,501,1742,1768]" country="Syria" name="Tartus" pageId="7" pageNumber="158">Tartous</collectingRegion>
(
<bibRefCitation id="EF8139CDC722FF8306222FA5F6C0F95D" author="Rees, A. F. &amp; Jony, M. &amp; Margaritoulis, D. &amp; Godley, B. J." box="[515,712,1742,1768]" pageId="7" pageNumber="158" pagination="49 - 54" refId="ref17283" refString="Rees, A. F., Jony, M., Margaritoulis, D. &amp; Godley, B. J. (2008 a) Satellite tracking of a green turtle, Chelonia mydas, from Syria further highlights importance of North Africa for Mediterranean turtles. Zoology in the Middle East, 45, 49 - 54. https: // doi. org / 10.1080 / 09397140.2008.10638306" type="journal article" year="2008">
Rees
<emphasis id="B964982EC722FF8306632FA4F672F95D" box="[578,634,1742,1768]" italics="true" pageId="7" pageNumber="158">et al.</emphasis>
2008
<bibRefCitation id="EF8139CDC722FF83069D2FA5F6C0F95D" author="Rees, A. F. &amp; Saad, A. &amp; Jony, M." box="[700,712,1742,1768]" pageId="7" pageNumber="158" pagination="456 - 459" refId="ref17483" refString="Rees, A. F., Saad, A. &amp; Jony, M. (2008 b) Discovery of a regionally important green turtle Chelonia mydas rookery in Syria. Oryx, 42, 456 - 459. https: // doi. org / 10.1017 / S 0030605308000926" type="journal article" year="2008">a</bibRefCitation>
</bibRefCitation>
, 2008b, 2010;
<bibRefCitation id="EF8139CDC722FF83074C2FA5F02CF95D" author="Saad, A. &amp; Rees, A. &amp; Sabour, W. &amp; Othman, A. &amp; Genaidi, C. &amp; Alkhateeb, N." box="[877,1060,1742,1768]" pageId="7" pageNumber="158" refId="ref18975" refString="Saad, A., Rees, A., Sabour, W., Othman, A., Genaidi, C. &amp; Alkhateeb, N. (2010) Six years of survey (2004 - 2009) on reproduction of sea turtles on Shkaifat beach, Syrian coast (proposed as protected area). Rapports et proces-verbaux des reunions Commission internationale pour l'exploration scientifique de la Mer Mediterranee, 39, 651." type="journal volume" year="2010">
Saad
<emphasis id="B964982EC722FF83078A2FA4F7EBF95D" box="[939,995,1742,1768]" italics="true" pageId="7" pageNumber="158">et al.</emphasis>
2010
</bibRefCitation>
, 2020; see also
<bibRefCitation id="EF8139CDC722FF8300F52FA5F194F95D" author="Casale, P. &amp; Broderick, A. C. &amp; Caminas, J. A. &amp; Cardona, L. &amp; Carreras, C. &amp; Demetropoulos, A. &amp; Fuller, W. J. &amp; Godley, B. J. &amp; Hochscheid, S. &amp; Kaska, Y. &amp; Lazar, B. &amp; Margaritoulis, D. &amp; Panagopoulou, A. &amp; Rees, A. F. &amp; Tomas, J. &amp; Turkozan, O." box="[1236,1436,1742,1768]" pageId="7" pageNumber="158" pagination="229 - 267" refId="ref13981" refString="Casale, P., Broderick, A. C., Caminas, J. A., Cardona, L., Carreras, C., Demetropoulos, A., Fuller, W. J., Godley, B. J., Hochscheid, S., Kaska, Y., Lazar, B., Margaritoulis, D., Panagopoulou, A., Rees, A. F., Tomas, J. &amp; Turkozan, O. (2018) Mediterranean Sea turtles: Current knowledge and priorities for conservation and research. Endangered Species Research, 36, 229 - 267. https: // doi. org / 10.3354 / esr 00901" type="journal article" year="2018">
Casale
<emphasis id="B964982EC722FF8301072FA4F156F95D" box="[1318,1374,1742,1768]" italics="true" pageId="7" pageNumber="158">et al.</emphasis>
2018
</bibRefCitation>
and our
<figureCitation id="132B58B9C722FF8304D02F99F53BF8B9" box="[241,307,1778,1804]" captionStart="FIGURE 6" captionStartId="9.[151,250,1389,1413]" captionTargetBox="[161,1429,191,1356]" captionTargetId="figure-17@9.[151,1436,181,1365]" captionTargetPageId="9" captionText="FIGURE 6. Nesting sites of Caretta caretta in Syria. Black circles: previous records, blue circles: recently confirmed previous records." pageId="7" pageNumber="158">Fig. 6</figureCitation>
). Although the coast is mainly sandy, most beaches have a rocky shelf at the edge of the sea, rendering them unsuitable for nesting. There are 12 records off the coast (
<bibRefCitation id="EF8139CDC722FF8307422E7DF03AF885" author="Halpin, P. N. &amp; Read, A. J. &amp; Fujioka, E. &amp; Best, B. D. &amp; Donnelly, B. &amp; Hazen, L. J. &amp; Kot, C. &amp; Urian, K. &amp; LaBrecque, E. &amp; Dimatteo, A. &amp; Cleary, J. &amp; Good, C. &amp; Crowder, L. B. &amp; Hyrenbach, K. D." box="[867,1074,1814,1840]" pageId="7" pageNumber="158" pagination="104 - 115" refId="ref15484" refString="Halpin, P. N., Read, A. J., Fujioka, E., Best, B. D., Donnelly, B., Hazen, L. J., Kot, C., Urian, K., LaBrecque, E., Dimatteo, A., Cleary, J., Good, C., Crowder, L. B. &amp; Hyrenbach, K. D. (2009) OBIS-SEAMAP: The world data center for marine mammal, sea bird, and sea turtle distributions. Oceanography, 22, 104 - 115. https: // doi. org / 10.5670 / oceanog. 2009.42" type="journal article" year="2009">
Halpin
<emphasis id="B964982EC722FF8307992E7CF7F9F885" box="[952,1009,1814,1840]" italics="true" pageId="7" pageNumber="158">et al.</emphasis>
2009
</bibRefCitation>
) and
<collectingRegion id="49D48ADEC722FF8300502E7DF0CDF885" box="[1137,1221,1814,1840]" country="Syria" name="Al Ladhiqiyah" pageId="7" pageNumber="158">Latakia</collectingRegion>
(iNaturalist 2023), respectively. Syrian
<taxonomicName id="4C103FBFC722FF83055A2E51F5E1F8E1" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[379,489,1850,1876]" class="Testudines" family="Cheloniidae" genus="Caretta" kingdom="Animalia" order="Cryptodira" pageId="7" pageNumber="158" phylum="Chordata" rank="species" species="caretta">
<emphasis id="B964982EC722FF83055A2E51F5E1F8E1" box="[379,489,1850,1876]" italics="true" pageId="7" pageNumber="158">C. caretta</emphasis>
</taxonomicName>
belong to the Mediterranean Regional Management Unit (
<bibRefCitation id="EF8139CDC722FF83005B2E51F150F8E1" author="Wallace, B. P. &amp; Posnik, Z. A. &amp; Hurley, B. J. &amp; DiMatteo, A. D. &amp; Bandimere, A. &amp; Rodriguez, I. &amp; Maxwell, S. M. &amp; Meyer, L. &amp; Brenner, H. &amp; Jensen, M. P. &amp; LaCasella, E. &amp; Shamblin, B. M. &amp; Abreu-Grobois, F. A. &amp; Stewart, K. R. &amp; Dutton, P. H. &amp; Barrios-Garrido, H. &amp; Dalleau, M. &amp; Dell'Amico, F. &amp; Eckert, K. L. &amp; FitzSimmons, N. N. &amp; Garcia-Cruz, M. &amp; Hays, G. C. &amp; Kelez, S. &amp; Lagueux, C. J. &amp; Madden Hof, C. A. &amp; Marco, A. &amp; Martins, S. L. T. &amp; Mobaraki, A. &amp; Mortimer, J. A. &amp; Nel, R. &amp; Phillott, A. D. &amp; Pilcher, N. J. &amp; Putman, N. F. &amp; Rees, A. F. &amp; Rguez-Baron, J. M. &amp; Seminoff, J. A. &amp; Swaminathan, A. &amp; Turkozan, O. &amp; Vargas, S. M. &amp; Vernet, P. D. &amp; Vilaca, S. &amp; Whiting, S. D. &amp; Hutchinson, B. J. &amp; Casale, P. &amp; Mast, R. B." box="[1146,1368,1850,1876]" pageId="7" pageNumber="158" pagination="209 - 223" refId="ref21120" refString="Wallace, B. P., Posnik, Z. A., Hurley, B. J., DiMatteo, A. D., Bandimere, A., Rodriguez, I., Maxwell, S. M., Meyer, L., Brenner, H., Jensen, M. P., LaCasella, E., Shamblin, B. M., Abreu-Grobois, F. A., Stewart, K. R., Dutton, P. H., Barrios-Garrido, H., Dalleau, M., Dell'Amico, F., Eckert, K. L., FitzSimmons, N. N., Garcia-Cruz, M., Hays, G. C., Kelez, S., Lagueux, C. J., Madden Hof, C. A., Marco, A., Martins, S. L. T., Mobaraki, A., Mortimer, J. A., Nel, R., Phillott, A. D., Pilcher, N. J., Putman, N. F., Rees, A. F., Rguez-Baron, J. M., Seminoff, J. A., Swaminathan, A., Turkozan, O., Vargas, S. M., Vernet, P. D., Vilaca, S., Whiting, S. D., Hutchinson, B. J., Casale, P. &amp; Mast, R. B. (2023) Marine turtle regional management units 2.0: An updated framework for conservation and research of wide-ranging megafauna species. Endangered Species Research, 52, 209 - 223. https: // doi. org / 10.3354 / ESR 01243" type="journal article" year="2023">
Wallace
<emphasis id="B964982EC722FF8300FD2E50F105F8E1" box="[1244,1293,1850,1876]" italics="true" pageId="7" pageNumber="158">et al</emphasis>
. 2023
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30A17B7C722FF8304E62E35F108F875" pageId="7" pageNumber="158" type="discussion">
<paragraph id="8BAF443CC722FF8304E62E35F063F829" blockId="7.[151,1436,1345,1984]" pageId="7" pageNumber="158">
<emphasis id="B964982EC722FF8304E62E35F53CF8CD" bold="true" box="[199,308,1886,1912]" pageId="7" pageNumber="158">Remarks</emphasis>
: There are 14 putative records from
<collectingCountry id="F30704ACC722FF8306CA2E35F72EF8CD" box="[747,806,1886,1912]" name="Syria" pageId="7" pageNumber="158">Syria</collectingCountry>
on GBIF, but these are situated either in neighboring countries or the geographical coordinates refer to inland cities in
<collectingCountry id="F30704ACC722FF8307502EE9F7A4F829" box="[881,940,1922,1948]" name="Syria" pageId="7" pageNumber="158">Syria</collectingCountry>
far from the sea.
</paragraph>
<paragraph id="8BAF443CC722FF8304E62ECDF108F875" blockId="7.[151,1436,1345,1984]" box="[199,1280,1958,1984]" pageId="7" pageNumber="158">
Syrian
<taxonomicName id="4C103FBFC722FF8305362ECDF58EF875" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[279,390,1958,1984]" class="Testudines" family="Cheloniidae" genus="Caretta" kingdom="Animalia" order="Cryptodira" pageId="7" pageNumber="158" phylum="Chordata" rank="species" species="caretta">
<emphasis id="B964982EC722FF8305362ECDF58EF875" box="[279,390,1958,1984]" italics="true" pageId="7" pageNumber="158">C. caretta</emphasis>
</taxonomicName>
belong to the Mediterranean Regional Management Unit (
<bibRefCitation id="EF8139CDC722FF8300372ECDF0FCF875" author="Wallace, B. P. &amp; Posnik, Z. A. &amp; Hurley, B. J. &amp; DiMatteo, A. D. &amp; Bandimere, A. &amp; Rodriguez, I. &amp; Maxwell, S. M. &amp; Meyer, L. &amp; Brenner, H. &amp; Jensen, M. P. &amp; LaCasella, E. &amp; Shamblin, B. M. &amp; Abreu-Grobois, F. A. &amp; Stewart, K. R. &amp; Dutton, P. H. &amp; Barrios-Garrido, H. &amp; Dalleau, M. &amp; Dell'Amico, F. &amp; Eckert, K. L. &amp; FitzSimmons, N. N. &amp; Garcia-Cruz, M. &amp; Hays, G. C. &amp; Kelez, S. &amp; Lagueux, C. J. &amp; Madden Hof, C. A. &amp; Marco, A. &amp; Martins, S. L. T. &amp; Mobaraki, A. &amp; Mortimer, J. A. &amp; Nel, R. &amp; Phillott, A. D. &amp; Pilcher, N. J. &amp; Putman, N. F. &amp; Rees, A. F. &amp; Rguez-Baron, J. M. &amp; Seminoff, J. A. &amp; Swaminathan, A. &amp; Turkozan, O. &amp; Vargas, S. M. &amp; Vernet, P. D. &amp; Vilaca, S. &amp; Whiting, S. D. &amp; Hutchinson, B. J. &amp; Casale, P. &amp; Mast, R. B." box="[1046,1268,1958,1984]" pageId="7" pageNumber="158" pagination="209 - 223" refId="ref21120" refString="Wallace, B. P., Posnik, Z. A., Hurley, B. J., DiMatteo, A. D., Bandimere, A., Rodriguez, I., Maxwell, S. M., Meyer, L., Brenner, H., Jensen, M. P., LaCasella, E., Shamblin, B. M., Abreu-Grobois, F. A., Stewart, K. R., Dutton, P. H., Barrios-Garrido, H., Dalleau, M., Dell'Amico, F., Eckert, K. L., FitzSimmons, N. N., Garcia-Cruz, M., Hays, G. C., Kelez, S., Lagueux, C. J., Madden Hof, C. A., Marco, A., Martins, S. L. T., Mobaraki, A., Mortimer, J. A., Nel, R., Phillott, A. D., Pilcher, N. J., Putman, N. F., Rees, A. F., Rguez-Baron, J. M., Seminoff, J. A., Swaminathan, A., Turkozan, O., Vargas, S. M., Vernet, P. D., Vilaca, S., Whiting, S. D., Hutchinson, B. J., Casale, P. &amp; Mast, R. B. (2023) Marine turtle regional management units 2.0: An updated framework for conservation and research of wide-ranging megafauna species. Endangered Species Research, 52, 209 - 223. https: // doi. org / 10.3354 / ESR 01243" type="journal article" year="2023">
Wallace
<emphasis id="B964982EC722FF8300592ECCF0A2F875" box="[1144,1194,1958,1984]" italics="true" pageId="7" pageNumber="158">et al</emphasis>
. 2023
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,364 @@
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<mods:title id="BDB7966603CB591BCF207D9C1AD0EA01">Turtles and tortoises of Syria: Diversity, distribution, and conservation</mods:title>
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<paragraph id="8BAF443CC729FF8804B62F2CF632F9D4" blockId="12.[151,626,1607,1669]" box="[151,570,1607,1633]" pageId="12" pageNumber="163">
<heading id="D0E7F350C729FF8804B62F2CF632F9D4" bold="true" box="[151,570,1607,1633]" fontSize="11" level="1" pageId="12" pageNumber="163" reason="1">
<taxonomicName id="4C103FBFC729FF8804B62F2CF632F9D4" ID-CoL="3YFW8" authority="(Gmelin, 1774)" baseAuthorityName="Gmelin" baseAuthorityYear="1774" box="[151,570,1607,1633]" class="Testudines" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Cryptodira" pageId="12" pageNumber="163" phylum="Chordata" rank="species" species="caspica">
<emphasis id="B964982EC729FF8804B62F2CF632F9D4" bold="true" box="[151,570,1607,1633]" pageId="12" pageNumber="163">
<emphasis id="B964982EC729FF8804B62F2CF571F9D4" bold="true" box="[151,377,1607,1633]" italics="true" pageId="12" pageNumber="163">Mauremys caspica</emphasis>
(
<bibRefCitation id="EF8139CDC729FF8805A92F2CF63AF9D4" author="Gmelin, S. G." box="[392,562,1607,1633]" pageId="12" pageNumber="163" refId="ref15314" refString="Gmelin, S. G. (1774) Reise durch Russland zur Untersuchung der drey Natur-Reiche. Dritter Theil. Reise durch das nordliche Persien, in den Jahren 1770, 1771, bis im April 1772. Kayserliche Academie der Wissenschaften, St. Petersburg, 508 pp." type="book" year="1774">Gmelin, 1774</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BAF443CC729FF8804B62F00F67AF930" blockId="12.[151,626,1607,1669]" box="[151,626,1643,1669]" pageId="12" pageNumber="163">
Caspian Turtle, Caspian Terrapin (
<figureCitation id="132B58B9C729FF8806392F00F661F930" box="[536,617,1643,1669]" captionStart="FIGURE 12" captionStartId="14.[151,250,1064,1088]" captionTargetBox="[151,1436,181,1041]" captionTargetId="figure-21@14.[151,1436,181,1041]" captionTargetPageId="14" captionText="FIGURE 12. Caspian terrapins, Mauremys caspica. (A) Muhasan, Euphrates (photo: Ahmad Aidek); (B) Ayyash, Euphrates (photo: Ahmad Aidek); (C) Deir ez-Zor, Euphrates (photo: Abu Al-Abbas); (D) Nahr Al-Sajoor (photo: Zuhair Amr)." pageId="12" pageNumber="163">Fig. 12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30A17B7C729FF8804B62FD8F6BFF978" box="[151,695,1715,1742]" pageId="12" pageNumber="163" type="description">
<paragraph id="8BAF443CC729FF8804B62FD8F6BFF978" blockId="12.[151,1436,1715,1922]" box="[151,695,1715,1742]" pageId="12" pageNumber="163">
<emphasis id="B964982EC729FF8804B62FD8F5C9F978" bold="true" box="[151,449,1715,1741]" pageId="12" pageNumber="163">IUCN Red List Category</emphasis>
: Least Concern (LC).
</paragraph>
</subSubSection>
<subSubSection id="C30A17B7C729FF8A04E62FBCF4FBFAD4" lastPageId="14" lastPageNumber="165" pageId="12" pageNumber="163" type="materials_examined">
<paragraph id="8BAF443CC729FF8804E62FBCF026F944" blockId="12.[151,1436,1715,1922]" box="[199,1070,1751,1778]" pageId="12" pageNumber="163">
<materialsCitation id="3B784E61C729FF8804E62FBCF022F944" box="[199,1066,1751,1778]" country="Azerbaijan" location="Azerbaijan" pageId="12" pageNumber="163" specimenCount="1" typeStatus="holotype">
<emphasis id="B964982EC729FF8804E62FBCF568F944" bold="true" box="[199,352,1751,1777]" pageId="12" pageNumber="163">
<typeStatus id="54ABFA9EC729FF8804E62FBCF50AF944" box="[199,258,1751,1777]" pageId="12" pageNumber="163">Type</typeStatus>
locality
</emphasis>
: “Schmachie, Bach Pusahat” [Pirsagat, Shemakha],
<collectingCountry id="F30704ACC729FF88078A2FBCF022F944" box="[939,1066,1751,1777]" name="Azerbaijan" pageId="12" pageNumber="163">Azerbaijan</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8BAF443CC729FF8804E62F90F562F88C" blockId="12.[151,1436,1715,1922]" pageId="12" pageNumber="163">
<emphasis id="B964982EC729FF8804E62F90F57EF8A0" bold="true" box="[199,374,1787,1813]" pageId="12" pageNumber="163">Type specimen</emphasis>
: Not located, specimen figured in
<bibRefCitation id="EF8139CDC729FF8806D62F90F79EF8A0" author="Gmelin, S. G." box="[759,918,1787,1813]" pageId="12" pageNumber="163" refId="ref15314" refString="Gmelin, S. G. (1774) Reise durch Russland zur Untersuchung der drey Natur-Reiche. Dritter Theil. Reise durch das nordliche Persien, in den Jahren 1770, 1771, bis im April 1772. Kayserliche Academie der Wissenschaften, St. Petersburg, 508 pp." type="book" year="1774">Gmelin (1774</bibRefCitation>
: plates 10, 11) designated as
<typeStatus id="54ABFA9EC729FF8800F92F90F137F8A0" box="[1240,1343,1787,1813]" pageId="12" pageNumber="163" type="lectotype">lectotype</typeStatus>
by
<bibRefCitation id="EF8139CDC729FF8801462F90F56EF88C" author="Fritz, U. &amp; Wischuf, T." pageId="12" pageNumber="163" pagination="223 - 260" refId="ref15001" refString="Fritz, U. &amp; Wischuf, T. (1997) Zur Systematik westasiatisch - sudosteuropaischer Bachschildkroten (Gattung Mauremys) (Reptilia: Testudines: Bataguridae). Zoologische Abhandlungen, 49, 223 - 260." type="journal article" year="1997">Fritz &amp; Wischuf (1997)</bibRefCitation>
.
</paragraph>
<paragraph id="8BAF443CC729FF8804E62E28F651F8E8" blockId="12.[151,1436,1715,1922]" box="[199,601,1859,1886]" pageId="12" pageNumber="163">
<emphasis id="B964982EC729FF8804E62E28F54EF8E8" bold="true" box="[199,326,1859,1885]" pageId="12" pageNumber="163">Chorotype</emphasis>
: Turano-Mediterranean.
</paragraph>
<paragraph id="8BAF443CC729FF8804E62E0CF030F834" blockId="12.[151,1436,1715,1922]" box="[199,1080,1895,1922]" pageId="12" pageNumber="163">
<emphasis id="B964982EC729FF8804E62E0CF583F834" bold="true" box="[199,395,1895,1921]" pageId="12" pageNumber="163">First records for</emphasis>
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<emphasis id="B964982EC729FF8805B22E0CF5DBF834" bold="true" box="[403,467,1895,1921]" pageId="12" pageNumber="163">
<collectingCountry id="F30704ACC729FF8805B22E0CF5DBF834" box="[403,467,1895,1921]" name="Syria" pageId="12" pageNumber="163">Syria</collectingCountry>
</emphasis>
: Maskanah,
<collectingRegion id="49D48ADEC729FF8806432E03F6C3F837" box="[610,715,1896,1922]" country="Syria" name="Ar Raqqah" pageId="12" pageNumber="163">ar-Raqqa</collectingRegion>
,
<collectingRegion id="49D48ADEC729FF8806F72E0CF752F837" box="[726,858,1895,1922]" country="Syria" name="Dayr az Zawr" pageId="12" pageNumber="163">Deir ez-Zor</collectingRegion>
(
<bibRefCitation id="EF8139CDC729FF8807492E0CF023F834" author="Siebenrock, F." box="[872,1067,1895,1921]" pageId="12" pageNumber="163" pagination="171 - 225" refId="ref19218" refString="Siebenrock, F. (1913) Wissenschaftliche Ergebnisse der Expedition nach Mesopotamien, 1910. Schildkroten aus Syrien und Mesopotamien. Annalen des Naturhistorischen Museums Wien, 27, 171 - 225." type="journal article" year="1913">Siebenrock 1913</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
<caption id="DF6F14B4C728FF8904B62C07F17BFA31" box="[151,1395,1388,1412]" pageId="13" pageNumber="164" startId="13.[151,250,1388,1412]" targetBox="[161,1429,191,1357]" targetPageId="13" targetType="figure">
<paragraph id="8BAF443CC728FF8904B62C07F17BFA31" blockId="13.[151,1395,1388,1412]" box="[151,1395,1388,1412]" pageId="13" pageNumber="164">
<emphasis id="B964982EC728FF8904B62C07F516FA31" bold="true" box="[151,286,1388,1412]" pageId="13" pageNumber="164">FIGURE 11.</emphasis>
Records of
<taxonomicName id="4C103FBFC728FF8905B82C06F683FA31" box="[409,651,1388,1412]" class="Testudines" family="Emydidae" genus="Emys" kingdom="Animalia" order="Cryptodira" pageId="13" pageNumber="164" phylum="Chordata" rank="subSpecies" species="orbicularis" subSpecies="eiselti">
<emphasis id="B964982EC728FF8905B82C06F683FA31" box="[409,651,1388,1412]" italics="true" pageId="13" pageNumber="164">Emys orbicularis eiselti</emphasis>
</taxonomicName>
in Syria. Black circle: vouchered previous record, red circle: new record.
</paragraph>
</caption>
<paragraph id="8BAF443CC728FF8904E62CBDF7F7F935" blockId="13.[151,1437,1494,1773]" pageId="13" pageNumber="164">
<emphasis id="B964982EC728FF8904E62CBDF589FA45" bold="true" box="[199,385,1494,1520]" pageId="13" pageNumber="164">Literature data</emphasis>
: Maskanah,
<collectingRegion id="49D48ADEC728FF8906352CBCF675FA44" box="[532,637,1495,1521]" country="Syria" name="Ar Raqqah" pageId="13" pageNumber="164">ar-Raqqa</collectingRegion>
,
<collectingRegion id="49D48ADEC728FF8906AB2CBDF718FA44" box="[650,784,1494,1521]" country="Syria" name="Dayr az Zawr" pageId="13" pageNumber="164">Deir ez-Zor</collectingRegion>
(
<bibRefCitation id="EF8139CDC728FF8907012CBDF7EBFA45" author="Siebenrock, F." box="[800,995,1494,1520]" pageId="13" pageNumber="164" pagination="171 - 225" refId="ref19218" refString="Siebenrock, F. (1913) Wissenschaftliche Ergebnisse der Expedition nach Mesopotamien, 1910. Schildkroten aus Syrien und Mesopotamien. Annalen des Naturhistorischen Museums Wien, 27, 171 - 225." type="journal article" year="1913">Siebenrock 1913</bibRefCitation>
), Tell Al-Budeiry (
<bibRefCitation id="EF8139CDC728FF89009A2CBDF4DAF9A1" author="Krupp, F. &amp; Schneider, W." pageId="13" pageNumber="164" pagination="69 - 85" refId="ref16487" refString="Krupp, F. &amp; Schneider, W. (1991) Bestandserfassung der rezenten Fauna im Bereich des Nahr al-Habur. In: Kuhne, H. (Ed.), Die rezente Umwelt von Tall Seh Hamad und Daten zur Umweltrekonstruktion der assyrischen Stadt Dur-Katlimmu (Berichte der Ausgrabung von Tall Seh Hamad / Dur-Katlimmu 1). D. Reimer, Berlin, pp. 69 - 85." type="book chapter" year="1991">Krupp &amp; Schneider 1991</bibRefCitation>
), Ras Al-Ayn (N Al
<collectingRegion id="49D48ADEC728FF89058D2C91F60AF9A1" box="[428,514,1530,1556]" country="Syria" name="Al Hasakah" pageId="13" pageNumber="164">Hasaka</collectingRegion>
;
<bibRefCitation id="EF8139CDC728FF89062A2C90F69BF9A1" author="Krupp, F." box="[523,659,1530,1557]" pageId="13" pageNumber="164" pagination="26 - 33" refId="ref16464" refString="Krupp, F. (1992) The karst springs of Ras al-Ain. Aqua geographia, 1, 26 - 33." type="journal article" year="1992">Krupp 1992</bibRefCitation>
),
<quantity id="4CE8E9D9C728FF8906842C91F6D4F9A1" box="[677,732,1530,1557]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="164" unit="km" value="5.0">5 km</quantity>
N as-Suwar, Buhayrat Al-Asad (
<bibRefCitation id="EF8139CDC728FF8900632C91F4DBF98D" author="Lymberakis, R. &amp; Kalionzopoulou, A." pageId="13" pageNumber="164" pagination="33 - 39" refId="ref16609" refString="Lymberakis, R. &amp; Kalionzopoulou, A. (2003) Additions to the herpetofauna of Syria. Zoology in the Middle East, 29, 33 - 39. https: // doi. org / 10.1080 / 09397140.2003.10637967" type="journal article" year="2003">Lymberakis &amp; Kalionzopoulou 2003</bibRefCitation>
), Wadi Samum, Buhayrat Al-Khatuniyah, Wadi Furati, Tell Gunaidiya,
<collectingRegion id="49D48ADEC728FF8907C72F74F047F98C" box="[998,1103,1567,1593]" country="Syria" name="Ar Raqqah" pageId="13" pageNumber="164">ar-Raqqa</collectingRegion>
,
<collectingRegion id="49D48ADEC728FF8900792F75F0D3F98C" box="[1112,1243,1566,1593]" country="Syria" name="Dayr az Zawr" pageId="13" pageNumber="164">Deir ez-Zor</collectingRegion>
(
<bibRefCitation id="EF8139CDC728FF8900C62F75F4DBF9E9" author="Fritz, U. &amp; Wischuf, T." pageId="13" pageNumber="164" pagination="223 - 260" refId="ref15001" refString="Fritz, U. &amp; Wischuf, T. (1997) Zur Systematik westasiatisch - sudosteuropaischer Bachschildkroten (Gattung Mauremys) (Reptilia: Testudines: Bataguridae). Zoologische Abhandlungen, 49, 223 - 260." type="journal article" year="1997">Fritz &amp; Wischuf 1997</bibRefCitation>
), Nahr Al-Sajoor (
<bibRefCitation id="EF8139CDC728FF8905802F29F650F9E9" author="Amr, Z. &amp; Shehab, A. &amp; Abu Baker, M." box="[417,600,1602,1628]" pageId="13" pageNumber="164" pagination="21 - 26" refId="ref13174" refString="Amr, Z., Shehab, A. &amp; Abu Baker, M. (2007) Recent observations on the herpetofauna of Syria with notes on trade in reptiles. Herpetozoa, 20, 21 - 26." type="journal article" year="2007">
Amr
<emphasis id="B964982EC728FF8905FC2F28F61DF9E9" box="[477,533,1602,1628]" italics="true" pageId="13" pageNumber="164">et al.</emphasis>
2007
</bibRefCitation>
), Haweijt Katee (
<bibRefCitation id="EF8139CDC728FF8907052F29F7A5F9E9" author="Aidek, E. A." box="[804,941,1602,1628]" pageId="13" pageNumber="164" refId="ref12916" refString="Aidek, E. A. (2010) A Guide to the Biodiversity of Deir ez - Zor Area. Al-Quds House for Sciences, Damascus, 112 pp." type="book" year="2010">Aidek 2010</bibRefCitation>
), widespread along the Euphrates (
<bibRefCitation id="EF8139CDC728FF8901152F29F532F935" author="Murdoch, D. A. &amp; Aidek, A." pageId="13" pageNumber="164" pagination="152 - 176" refId="ref16892" refString="Murdoch, D. A. &amp; Aidek, A. (2012) Birding sites of the OSME Region 8 - The birds of the lower Syrian Euphrates. Sandgrouse, 34, 152 - 176." type="journal article" year="2012">Murdoch &amp; Aidek 2012</bibRefCitation>
), Tell Sheikh Hamad (
<bibRefCitation id="EF8139CDC728FF8906122F0DF6E3F935" author="Fritz, U. &amp; Ayaz, D. &amp; Buschbom, J. &amp; Kami, H. G. &amp; Mazanaeva, L. F. &amp; Aloufi, A. A. &amp; Auer, M. &amp; Rifai, L. &amp; Silic, T. &amp; Hundsdorfer, A. K." box="[563,747,1638,1664]" pageId="13" pageNumber="164" pagination="527 - 540" refId="ref14605" refString="Fritz, U., Ayaz, D., Buschbom, J., Kami, H. G., Mazanaeva, L. F., Aloufi, A. A., Auer, M., Rifai, L., Silic, T. &amp; Hundsdorfer, A. K. (2008) Go East: Phylogeographies of Mauremys caspica and M. rivulata - Discordance of morphology, mitochondrial and nuclear genomic markers and rare hybridization. Journal of Evolutionary Biology, 21, 527 - 540. https: // doi. org / 10.1111 / j. 1420 - 9101.2007.01485. x" type="journal article" year="2008">
Fritz
<emphasis id="B964982EC728FF8906502F0CF6A1F935" box="[625,681,1638,1664]" italics="true" pageId="13" pageNumber="164">et al.</emphasis>
2008
</bibRefCitation>
;
<bibRefCitation id="EF8139CDC728FF8906D72F0DF7FBF935" author="Vamberger, M. &amp; Stuckas, H. &amp; Ayaz, D. &amp; Gracia, E. &amp; Aloufi, A. A. &amp; Els, J. &amp; Mazanaeva, L. F. &amp; Kami, H. G. &amp; Fritz, U." box="[758,1011,1638,1664]" pageId="13" pageNumber="164" pagination="77 - 85" refId="ref20717" refString="Vamberger, M., Stuckas, H., Ayaz, D., Gracia, E., Aloufi, A. A., Els, J., Mazanaeva, L. F., Kami, H. G. &amp; Fritz, U. (2013) Conservation genetics and phylogeography of the poorly known Middle Eastern terrapin Mauremys caspica (Testudines: Geoemydidae). Organisms, Diversity &amp; Evolution, 13, 77 - 85. https: // doi. org / 10.1007 / s 13127 - 012 - 0102 - 6" type="journal article" year="2013">
Vamberger
<emphasis id="B964982EC728FF8907562F0CF7B8F935" box="[887,944,1638,1664]" italics="true" pageId="13" pageNumber="164">et al.</emphasis>
2013
</bibRefCitation>
).
</paragraph>
<paragraph id="8BAF443CC728FF8904E62FE1F6BAF97D" blockId="13.[151,1437,1494,1773]" pageId="13" pageNumber="164">
<emphasis id="B964982EC728FF8904E62FE1F5B9F911" bold="true" box="[199,433,1674,1700]" pageId="13" pageNumber="164">Museum specimens</emphasis>
:
<collectingCountry id="F30704ACC728FF8905E02FE1F5F4F911" box="[449,508,1674,1700]" name="Syria" pageId="13" pageNumber="164">Syria</collectingCountry>
(
<collectionCode id="ED01DCF9C728FF89062D2FE1F657F911" box="[524,607,1674,1700]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="13" pageNumber="164" type="Museum">FMNH</collectionCode>
73491, 73493, 73494),
<collectingRegion id="49D48ADEC728FF8907512FE1F7CAF911" box="[880,962,1674,1700]" country="Syria" name="Halab" pageId="13" pageNumber="164">Aleppo</collectingRegion>
—Euphrates (
<collectionCode id="ED01DCF9C728FF89007F2FE1F0BDF911" box="[1118,1205,1674,1700]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="164">MNHN</collectionCode>
1881.397),
<collectingRegion id="49D48ADEC728FF8901612FE1F4B7F97C" country="Syria" name="Dayr az Zawr" pageId="13" pageNumber="164">Deir ez-Zor</collectingRegion>
(
<collectionCode id="ED01DCF9C728FF8904EF2FC5F51FF97D" box="[206,279,1710,1736]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="13" pageNumber="164">NMW</collectionCode>
29475:12, 29634:12, 29662:13).
</paragraph>
<paragraph id="8BAF443CC728FF8904E62FB9F766F959" blockId="13.[151,1437,1494,1773]" box="[199,878,1746,1773]" pageId="13" pageNumber="164">
<emphasis id="B964982EC728FF8904E62FB9F5C6F959" bold="true" box="[199,462,1746,1772]" pageId="13" pageNumber="164">Citizen science record</emphasis>
:
<collectingRegion id="49D48ADEC728FF8905FD2FB9F668F958" box="[476,608,1746,1773]" country="Syria" name="Dayr az Zawr" pageId="13" pageNumber="164">Deir ez-Zor</collectingRegion>
(observation.org 2023).
</paragraph>
<caption id="DF6F14B4C72BFF8A04B62D43F131FBD0" pageId="14" pageNumber="165" startId="14.[151,250,1064,1088]" targetBox="[151,1436,181,1041]" targetPageId="14" targetType="figure">
<paragraph id="8BAF443CC72BFF8A04B62D43F131FBD0" blockId="14.[151,1436,1064,1125]" pageId="14" pageNumber="165">
<emphasis id="B964982EC72BFF8A04B62D43F52AFBF5" bold="true" box="[151,290,1064,1088]" pageId="14" pageNumber="165">FIGURE 12.</emphasis>
Caspian terrapins,
<taxonomicName id="4C103FBFC72BFF8A05CA2D42F6A2FBF5" baseAuthorityName="Gmelin" baseAuthorityYear="1774" box="[491,682,1065,1088]" class="Testudines" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Cryptodira" pageId="14" pageNumber="165" phylum="Chordata" rank="species" species="caspica">
<emphasis id="B964982EC72BFF8A05CA2D42F6A2FBF5" box="[491,682,1065,1088]" italics="true" pageId="14" pageNumber="165">Mauremys caspica</emphasis>
</taxonomicName>
. (
<emphasis id="B964982EC72BFF8A06E32D43F6DCFBF5" bold="true" box="[706,724,1064,1088]" pageId="14" pageNumber="165">A</emphasis>
) Muhasan, Euphrates (photo: Ahmad Aidek); (
<emphasis id="B964982EC72BFF8A009C2D42F0C6FBF4" bold="true" box="[1213,1230,1065,1089]" pageId="14" pageNumber="165">B</emphasis>
) Ayyash, Euphrates (photo: Ahmad Aidek); (
<emphasis id="B964982EC72BFF8A05B02D27F5ABFBD1" bold="true" box="[401,419,1100,1124]" pageId="14" pageNumber="165">C</emphasis>
) Deir ez-Zor, Euphrates (photo: Abu Al-Abbas); (
<emphasis id="B964982EC72BFF8A07812D26F7BAFBD0" bold="true" box="[928,946,1101,1125]" pageId="14" pageNumber="165">D</emphasis>
) Nahr Al-Sajoor (photo: Zuhair Amr).
</paragraph>
</caption>
<paragraph id="8BAF443CC72BFF8A04E62DDCF4FBFAD4" blockId="14.[151,1437,1207,1881]" pageId="14" pageNumber="165">
<emphasis id="B964982EC72BFF8A04E62DDCF577FB64" bold="true" box="[199,383,1207,1233]" pageId="14" pageNumber="165">Recent records</emphasis>
: Abu Hardoub, Abu Kamal, Ain Arous, aj-Jawadiyeh, Al-Ashara, Ali Baljeyeh, Al-Boliel,
<collectingRegion id="49D48ADEC72BFF8A04B62DB0F51AFB40" box="[151,274,1243,1269]" country="Syria" name="Al Hasakah" pageId="14" pageNumber="165">Al-Hasaka</collectingRegion>
, Al-Kasra, Al-Mayadin, Al-Mriiyah, Al-Qalteh, Al-Qamishli (Nahr Jaqjaq), Al-Qouriyah, Al-Qsupy, Al-Rheiat, Al-Shafa, Al-Shmeitiyeh, Ayyash, Buqruss, Buhayrat Al-Khatuniyah,
<collectingRegion id="49D48ADEC72BFF8A00142D94F0B3FAAC" box="[1077,1211,1279,1305]" country="Syria" name="Dayr az Zawr" pageId="14" pageNumber="165">Deir ez-Zor</collectingRegion>
, Ghandura, Hajeen, Hatla, Hazeimeh, Jarablus, Maadan, Muhasan,
<collectingRegion id="49D48ADEC72BFF8A06892C48F719FA88" box="[680,785,1315,1341]" country="Syria" name="Ar Raqqah" pageId="14" pageNumber="165">ar-Raqqa</collectingRegion>
, Rmeilah, Sbeikhan, Tabqah, Tell Tartab, Theiban, Tokhar Sagheir.
</paragraph>
</subSubSection>
<subSubSection id="C30A17B7C72BFF8A04E62C00F09DFA1C" pageId="14" pageNumber="165" type="distribution">
<paragraph id="8BAF443CC72BFF8A04E62C00F09DFA1C" blockId="14.[151,1437,1207,1881]" pageId="14" pageNumber="165">
<emphasis id="B964982EC72BFF8A04E62C00F5C8FA31" bold="true" box="[199,448,1386,1413]" pageId="14" pageNumber="165">
Distribution in
<collectingCountry id="F30704ACC72BFF8A05A12C01F5C8FA31" box="[384,448,1386,1412]" name="Syria" pageId="14" pageNumber="165">Syria</collectingCountry>
</emphasis>
: Common and widespread along the Euphrates and its tributaries, Al-Sajoor, Al-Balikh, and Al-Khabour and in all suitable ponds in the northeast of Syrian Mesopotamia (
<figureCitation id="132B58B9C72BFF8A00152CE4F080FA1C" box="[1076,1160,1423,1449]" captionStart="FIGURE 13" captionStartId="15.[151,250,1385,1409]" captionTargetBox="[161,1429,191,1357]" captionTargetId="figure-17@15.[151,1436,181,1361]" captionTargetPageId="15" captionText="FIGURE 13. Records of Mauremys caspica in Syria. Black circles: previous records, blue circles: recently confirmed previous records, red circles: new records." pageId="14" pageNumber="165">Fig. 13</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30A17B7C72BFF8A04E62CD8F628F8EC" pageId="14" pageNumber="165" type="discussion">
<paragraph id="8BAF443CC72BFF8A04E62CD8F68CF8A4" blockId="14.[151,1437,1207,1881]" pageId="14" pageNumber="165">
<emphasis id="B964982EC72BFF8A04E62CD8F53CFA78" bold="true" box="[199,308,1459,1485]" pageId="14" pageNumber="165">Remarks</emphasis>
:
<taxonomicName id="4C103FBFC72BFF8A05642CDFF610FA78" baseAuthorityName="Gmelin" baseAuthorityYear="1774" box="[325,536,1460,1485]" class="Testudines" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Cryptodira" pageId="14" pageNumber="165" phylum="Chordata" rank="species" species="caspica">
<emphasis id="B964982EC72BFF8A05642CDFF610FA78" box="[325,536,1460,1485]" italics="true" pageId="14" pageNumber="165">Mauremys caspica</emphasis>
</taxonomicName>
is a widespread monotypic species, ranging from Central
<collectingCountry id="F30704ACC72BFF8A00902CD8F119FA78" box="[1201,1297,1459,1485]" name="Turkey" pageId="14" pageNumber="165">Anatolia</collectingCountry>
through the interior of the Near and Middle East to
<collectingCountry id="F30704ACC72BFF8A06742CBCF68DFA44" box="[597,645,1495,1521]" name="Iran" pageId="14" pageNumber="165">Iran</collectingCountry>
,
<collectingCountry id="F30704ACC72BFF8A06B12CBCF726FA44" box="[656,814,1495,1521]" name="Turkmenistan" pageId="14" pageNumber="165">Turkmenistan</collectingCountry>
, and
<collectingCountry id="F30704ACC72BFF8A07482CBCF7F3FA44" box="[873,1019,1495,1521]" name="Saudi Arabia" pageId="14" pageNumber="165">Saudi Arabia</collectingCountry>
(
<bibRefCitation id="EF8139CDC72BFF8A002B2CBCF0A8FA44" author="TTWG &amp; Turtle Taxonomy Working Group" box="[1034,1184,1495,1521]" pageId="14" pageNumber="165" pagination="1 - 472" refId="ref20400" refString="TTWG [Turtle Taxonomy Working Group] (2021) Turtles of the World: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status (9 th Ed.). Chelonian Research Monographs, 8, 1 - 472. https: // doi. org / 10.3854 / crm. 8. checklist. atlas. v 9.2021" type="journal article" year="2021">TTWG 2021</bibRefCitation>
). The three previously recognized subspecies (
<emphasis id="B964982EC72BFF8A05812C97F63DF9A0" box="[416,565,1532,1557]" italics="true" pageId="14" pageNumber="165">
M. c.
<taxonomicName id="4C103FBFC72BFF8A05FE2C97F63DF9A0" baseAuthorityName="Gmelin" baseAuthorityYear="1774" box="[479,565,1532,1557]" class="Testudines" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Cryptodira" pageId="14" pageNumber="165" phylum="Chordata" rank="species" species="caspica">caspica</taxonomicName>
</emphasis>
,
<emphasis id="B964982EC72BFF8A06632C97F709F9A0" box="[578,769,1531,1557]" italics="true" pageId="14" pageNumber="165">M. c. siebenrocki</emphasis>
,
<emphasis id="B964982EC72BFF8A072F2C97F7FEF9A0" box="[782,1014,1531,1557]" italics="true" pageId="14" pageNumber="165">M. c. ventrimaculata</emphasis>
) have been synonymized (
<bibRefCitation id="EF8139CDC72BFF8A01012C90F51FF98C" author="Vamberger, M. &amp; Stuckas, H. &amp; Ayaz, D. &amp; Gracia, E. &amp; Aloufi, A. A. &amp; Els, J. &amp; Mazanaeva, L. F. &amp; Kami, H. G. &amp; Fritz, U." pageId="14" pageNumber="165" pagination="77 - 85" refId="ref20717" refString="Vamberger, M., Stuckas, H., Ayaz, D., Gracia, E., Aloufi, A. A., Els, J., Mazanaeva, L. F., Kami, H. G. &amp; Fritz, U. (2013) Conservation genetics and phylogeography of the poorly known Middle Eastern terrapin Mauremys caspica (Testudines: Geoemydidae). Organisms, Diversity &amp; Evolution, 13, 77 - 85. https: // doi. org / 10.1007 / s 13127 - 012 - 0102 - 6" type="journal article" year="2013">
Vamberger
<emphasis id="B964982EC72BFF8A04B62F4BF4DAF98C" box="[151,210,1567,1593]" italics="true" pageId="14" pageNumber="165">et al.</emphasis>
2013
</bibRefCitation>
).
<taxonomicName id="4C103FBFC72BFF8A050D2F4BF608F98C" baseAuthorityName="Gmelin" baseAuthorityYear="1774" box="[300,512,1568,1593]" class="Testudines" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Cryptodira" pageId="14" pageNumber="165" phylum="Chordata" rank="species" species="caspica">
<emphasis id="B964982EC72BFF8A050D2F4BF608F98C" box="[300,512,1568,1593]" italics="true" pageId="14" pageNumber="165">Mauremys caspica</emphasis>
</taxonomicName>
sporadically hybridizes with its sister species
<taxonomicName id="4C103FBFC72BFF8A00342F4BF09DF98C" box="[1045,1173,1567,1593]" class="Testudines" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Cryptodira" pageId="14" pageNumber="165" phylum="Chordata" rank="species" species="rivulata">
<emphasis id="B964982EC72BFF8A00342F4BF09DF98C" box="[1045,1173,1567,1593]" italics="true" pageId="14" pageNumber="165">M. rivulata</emphasis>
</taxonomicName>
in the Levant (
<bibRefCitation id="EF8139CDC72BFF8A01662F74F53EF9E8" author="Fritz, U. &amp; Wischuf, T." pageId="14" pageNumber="165" pagination="223 - 260" refId="ref15001" refString="Fritz, U. &amp; Wischuf, T. (1997) Zur Systematik westasiatisch - sudosteuropaischer Bachschildkroten (Gattung Mauremys) (Reptilia: Testudines: Bataguridae). Zoologische Abhandlungen, 49, 223 - 260." type="journal article" year="1997">Fritz &amp; Wischuf 1997</bibRefCitation>
;
<bibRefCitation id="EF8139CDC72BFF8A05602F28F5FCF9E8" author="Fritz, U. &amp; Ayaz, D. &amp; Buschbom, J. &amp; Kami, H. G. &amp; Mazanaeva, L. F. &amp; Aloufi, A. A. &amp; Auer, M. &amp; Rifai, L. &amp; Silic, T. &amp; Hundsdorfer, A. K." box="[321,500,1603,1629]" pageId="14" pageNumber="165" pagination="527 - 540" refId="ref14605" refString="Fritz, U., Ayaz, D., Buschbom, J., Kami, H. G., Mazanaeva, L. F., Aloufi, A. A., Auer, M., Rifai, L., Silic, T. &amp; Hundsdorfer, A. K. (2008) Go East: Phylogeographies of Mauremys caspica and M. rivulata - Discordance of morphology, mitochondrial and nuclear genomic markers and rare hybridization. Journal of Evolutionary Biology, 21, 527 - 540. https: // doi. org / 10.1111 / j. 1420 - 9101.2007.01485. x" type="journal article" year="2008">
Fritz
<emphasis id="B964982EC72BFF8A055D2F2FF5BBF9E8" box="[380,435,1603,1629]" italics="true" pageId="14" pageNumber="165">et al.</emphasis>
2008
</bibRefCitation>
;
<bibRefCitation id="EF8139CDC72BFF8A05DF2F28F6F1F9E8" author="Vamberger, M. &amp; Stuckas, H. &amp; Vargas-Ramirez, M. &amp; Kehlmaier, C. &amp; Ayaz, D. &amp; Aloufi, A. A. &amp; Lymberakis, P. &amp; Siroky, P. &amp; Fritz, U." box="[510,761,1603,1629]" pageId="14" pageNumber="165" pagination="401 - 413" refId="ref20888" refString="Vamberger, M., Stuckas, H., Vargas-Ramirez, M., Kehlmaier, C., Ayaz, D., Aloufi, A. A., Lymberakis, P., Siroky, P. &amp; Fritz, U. (2017) Unexpected hybridization patterns in Near Eastern terrapins (Mauremys caspica, M. rivulata) indicate ancient gene flow across the Fertile Crescent. Zoologica Scripta, 46, 401 - 413. https: // doi. org / 10.1111 / zsc. 12227" type="journal article" year="2017">
Vamberger
<emphasis id="B964982EC72BFF8A065E2F2FF6BEF9E8" box="[639,694,1603,1629]" italics="true" pageId="14" pageNumber="165">et al.</emphasis>
2017
</bibRefCitation>
).
<taxonomicName id="4C103FBFC72BFF8A072B2F2FF7D3F9E8" baseAuthorityName="Gmelin" baseAuthorityYear="1774" box="[778,987,1604,1629]" class="Testudines" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Cryptodira" pageId="14" pageNumber="165" phylum="Chordata" rank="species" species="caspica">
<emphasis id="B964982EC72BFF8A072B2F2FF7D3F9E8" box="[778,987,1604,1629]" italics="true" pageId="14" pageNumber="165">Mauremys caspica</emphasis>
</taxonomicName>
is an inland species whereas
<taxonomicName id="4C103FBFC72BFF8A013E2F2FF195F9E8" box="[1311,1437,1603,1629]" class="Testudines" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Cryptodira" pageId="14" pageNumber="165" phylum="Chordata" rank="species" species="rivulata">
<emphasis id="B964982EC72BFF8A013E2F2FF195F9E8" box="[1311,1437,1603,1629]" italics="true" pageId="14" pageNumber="165">M. rivulata</emphasis>
</taxonomicName>
occurs in Mediterranean coastal habitats (
<bibRefCitation id="EF8139CDC72BFF8A065F2F0CF78DF934" author="Fritz, U. &amp; Wischuf, T." box="[638,901,1639,1665]" pageId="14" pageNumber="165" pagination="223 - 260" refId="ref15001" refString="Fritz, U. &amp; Wischuf, T. (1997) Zur Systematik westasiatisch - sudosteuropaischer Bachschildkroten (Gattung Mauremys) (Reptilia: Testudines: Bataguridae). Zoologische Abhandlungen, 49, 223 - 260." type="journal article" year="1997">Fritz &amp; Wischuf 1997</bibRefCitation>
). In the southern Levant, the local
<taxonomicName id="4C103FBFC72BFF8A01062F03F194F934" box="[1319,1436,1640,1665]" class="Testudines" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Cryptodira" pageId="14" pageNumber="165" phylum="Chordata" rank="genus">
<emphasis id="B964982EC72BFF8A01062F03F194F934" box="[1319,1436,1640,1665]" italics="true" pageId="14" pageNumber="165">Mauremys</emphasis>
</taxonomicName>
populations resemble
<taxonomicName id="4C103FBFC72BFF8A05AC2FE7F604F910" box="[397,524,1675,1701]" class="Testudines" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Cryptodira" pageId="14" pageNumber="165" phylum="Chordata" rank="species" species="rivulata">
<emphasis id="B964982EC72BFF8A05AC2FE7F604F910" box="[397,524,1675,1701]" italics="true" pageId="14" pageNumber="165">M. rivulata</emphasis>
</taxonomicName>
but represent a hybrid swarm with genetic signatures of
<taxonomicName id="4C103FBFC72BFF8A00A82FE7F10DF910" baseAuthorityName="Gmelin" baseAuthorityYear="1774" box="[1161,1285,1676,1701]" class="Testudines" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Cryptodira" pageId="14" pageNumber="165" phylum="Chordata" rank="species" species="caspica">
<emphasis id="B964982EC72BFF8A00A82FE7F10DF910" box="[1161,1285,1676,1701]" italics="true" pageId="14" pageNumber="165">M. caspica</emphasis>
</taxonomicName>
, even though the geographically closest populations of
<taxonomicName id="4C103FBFC72BFF8A06562FDBF6FDF97C" baseAuthorityName="Gmelin" baseAuthorityYear="1774" box="[631,757,1712,1737]" class="Testudines" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Cryptodira" pageId="14" pageNumber="165" phylum="Chordata" rank="species" species="caspica">
<emphasis id="B964982EC72BFF8A06562FDBF6FDF97C" box="[631,757,1712,1737]" italics="true" pageId="14" pageNumber="165">M. caspica</emphasis>
</taxonomicName>
are widely separated. Gene flow between the two species, resulting in the current hybrid swarm, was facilitated during more humid climate episodes of the last glaciation and the early Holocene (
<bibRefCitation id="EF8139CDC72BFF8A05582F9CF670F8A4" author="Vamberger, M. &amp; Stuckas, H. &amp; Vargas-Ramirez, M. &amp; Kehlmaier, C. &amp; Ayaz, D. &amp; Aloufi, A. A. &amp; Lymberakis, P. &amp; Siroky, P. &amp; Fritz, U." box="[377,632,1783,1809]" pageId="14" pageNumber="165" pagination="401 - 413" refId="ref20888" refString="Vamberger, M., Stuckas, H., Vargas-Ramirez, M., Kehlmaier, C., Ayaz, D., Aloufi, A. A., Lymberakis, P., Siroky, P. &amp; Fritz, U. (2017) Unexpected hybridization patterns in Near Eastern terrapins (Mauremys caspica, M. rivulata) indicate ancient gene flow across the Fertile Crescent. Zoologica Scripta, 46, 401 - 413. https: // doi. org / 10.1111 / zsc. 12227" type="journal article" year="2017">
Vamberger
<emphasis id="B964982EC72BFF8A05DD2F93F63DF8A4" box="[508,565,1783,1809]" italics="true" pageId="14" pageNumber="165">et al.</emphasis>
2017
</bibRefCitation>
).
</paragraph>
<paragraph id="8BAF443CC72BFF8A04E62E70F628F8EC" blockId="14.[151,1437,1207,1881]" pageId="14" pageNumber="165">
On GBIF, there are 27 records for
<taxonomicName id="4C103FBFC72BFF8A06142E77F6A5F880" baseAuthorityName="Gmelin" baseAuthorityYear="1774" box="[565,685,1820,1845]" class="Testudines" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Cryptodira" pageId="14" pageNumber="165" phylum="Chordata" rank="species" species="caspica">
<emphasis id="B964982EC72BFF8A06142E77F6A5F880" box="[565,685,1820,1845]" italics="true" pageId="14" pageNumber="165">M.caspica</emphasis>
</taxonomicName>
from
<collectingCountry id="F30704ACC72BFF8A06CD2E70F723F880" box="[748,811,1819,1845]" name="Syria" pageId="14" pageNumber="165">Syria</collectingCountry>
.Among them, 18 are actually from countries neighboring
<collectingCountry id="F30704ACC72BFF8A04B62E54F4DDF8EC" box="[151,213,1855,1881]" name="Syria" pageId="14" pageNumber="165">Syria</collectingCountry>
, and five refer to
<taxonomicName id="4C103FBFC72BFF8A05BA2E2BF611F8EC" box="[411,537,1855,1881]" class="Testudines" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Cryptodira" pageId="14" pageNumber="165" phylum="Chordata" rank="species" species="rivulata">
<emphasis id="B964982EC72BFF8A05BA2E2BF611F8EC" box="[411,537,1855,1881]" italics="true" pageId="14" pageNumber="165">M. rivulata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,239 @@
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<heading id="D0E7F350C72DFF8C04B62D40F63AFBF0" bold="true" box="[151,562,1067,1094]" fontSize="11" level="1" pageId="8" pageNumber="159" reason="1">
<taxonomicName id="4C103FBFC72DFF8C04B62D40F63AFBF0" ID-CoL="TVGD" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[151,562,1067,1094]" class="Testudines" family="Cheloniidae" genus="Chelonia" kingdom="Animalia" order="Cryptodira" pageId="8" pageNumber="159" phylum="Chordata" rank="species" species="mydas">
<emphasis id="B964982EC72DFF8C04B62D40F63AFBF0" bold="true" box="[151,562,1067,1094]" pageId="8" pageNumber="159">
<emphasis id="B964982EC72DFF8C04B62D40F552FBF3" bold="true" box="[151,346,1067,1094]" italics="true" pageId="8" pageNumber="159">Chelonia mydas</emphasis>
(Linnaeus, 1758)
</emphasis>
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<paragraph id="8BAF443CC72DFF8C04B62D24F645FBDC" blockId="8.[151,589,1067,1129]" box="[151,589,1103,1129]" pageId="8" pageNumber="159">
Green Turtle, Green Sea Turtle (
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)
</paragraph>
</subSubSection>
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<emphasis id="B964982EC72DFF8C04B62DFCF5C9FB04" bold="true" box="[151,449,1175,1201]" pageId="8" pageNumber="159">IUCN Red List Category</emphasis>
: Endangered (EN A2bd); Mediterranean subpopulation: not assessed.
</paragraph>
</subSubSection>
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<paragraph id="8BAF443CC72DFF8C04E62DD0F4E1FB4C" blockId="8.[151,1437,1175,1850]" pageId="8" pageNumber="159">
<materialsCitation id="3B784E61C72DFF8C04E62DD0F4EEFB4C" country="Ascension Island" location="Restricted" pageId="8" pageNumber="159" specimenCount="1" typeStatus="holotype">
<emphasis id="B964982EC72DFF8C04E62DD0F56AFB60" bold="true" box="[199,354,1211,1237]" pageId="8" pageNumber="159">
<typeStatus id="54ABFA9EC72DFF8C04E62DD0F50AFB60" box="[199,258,1211,1237]" pageId="8" pageNumber="159">Type</typeStatus>
locality
</emphasis>
: “Insulas Pelagi: insulam Adscensionis.”
<location id="8ECF12E7C72DFF8C07632DD0F7BCFB60" LSID="urn:lsid:plazi:treatment:03B9F52AC72DFF8C04B62D40F779F88C:8ECF12E7C72DFF8C07632DD0F7BCFB60" box="[834,948,1211,1237]" country="Ascension Island" name="Restricted" pageId="8" pageNumber="159">Restricted</location>
to
<collectingCountry id="F30704ACC72DFF8C07FA2DD0F095FB60" box="[987,1181,1211,1237]" name="Ascension Island" pageId="8" pageNumber="159">Ascension Island</collectingCountry>
by
<bibRefCitation id="EF8139CDC72DFF8C00ED2DD7F4EEFB4C" author="Mertens, R. &amp; Muller, L." pageId="8" pageNumber="159" pagination="1 - 62" refId="ref16694" refString="Mertens, R. &amp; Muller, L. (1928) Liste der Amphibien und Reptilien Europas. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 41, 1 - 62." type="journal article" year="1928">Mertens &amp; Müller (1928)</bibRefCitation>
</materialsCitation>
.
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<paragraph id="8BAF443CC72DFF8C04E62C68F585FAF7" blockId="8.[151,1437,1175,1850]" pageId="8" pageNumber="159">
<emphasis id="B964982EC72DFF8C04E62C68F58CFAA8" bold="true" box="[199,388,1283,1309]" pageId="8" pageNumber="159">Type specimens</emphasis>
:
<materialsCitation id="3B784E61C72DFF8C05B42C68F581FAF7" county="Uppsala University" location="Uppsala Zoological Type Collection" municipality="Museum of Evolution" pageId="8" pageNumber="159" specimenCount="1" typeStatus="syntype">
<location id="8ECF12E7C72DFF8C05B42C68F73CFAA8" LSID="urn:lsid:plazi:treatment:03B9F52AC72DFF8C04B62D40F779F88C:8ECF12E7C72DFF8C05B42C68F73CFAA8" box="[405,820,1283,1310]" county="Uppsala University" municipality="Museum of Evolution" name="Uppsala Zoological Type Collection" pageId="8" pageNumber="159">Uppsala Zoological Type Collection</location>
,
<collectingMunicipality id="6BCBDE46C72DFF8C07602C6FF035FAA8" box="[833,1085,1283,1310]" pageId="8" pageNumber="159">Museum of Evolution</collectingMunicipality>
,
<collectingCounty id="62CE3CB0C72DFF8C006B2C68F12EFAA8" box="[1098,1318,1283,1309]" pageId="8" pageNumber="159">Uppsala University</collectingCounty>
, UPSZTY 19, 26, 231,
<typeStatus id="54ABFA9EC72DFF8C05022C43F581FAF7" box="[291,393,1320,1346]" pageId="8" pageNumber="159" type="syntype">syntypes</typeStatus>
</materialsCitation>
.
</paragraph>
<paragraph id="8BAF443CC72DFF8C04E62C20F6A5FAD0" blockId="8.[151,1437,1175,1850]" box="[199,685,1355,1382]" pageId="8" pageNumber="159">
<emphasis id="B964982EC72DFF8C04E62C20F54EFAD0" bold="true" box="[199,326,1355,1381]" pageId="8" pageNumber="159">Chorotype</emphasis>
: Circumglobal (Cosmopolitan).
</paragraph>
<paragraph id="8BAF443CC72DFF8C04E62C04F7C5FA3F" blockId="8.[151,1437,1175,1850]" box="[199,973,1391,1418]" pageId="8" pageNumber="159">
<emphasis id="B964982EC72DFF8C04E62C04F5C0FA3C" bold="true" box="[199,456,1391,1417]" pageId="8" pageNumber="159">
First record for
<collectingCountry id="F30704ACC72DFF8C05A92C04F5C0FA3C" box="[392,456,1391,1417]" name="Syria" pageId="8" pageNumber="159">Syria</collectingCountry>
</emphasis>
: Between
<collectingRegion id="49D48ADEC72DFF8C061E2C04F69BFA3C" box="[575,659,1391,1417]" country="Syria" name="Al Ladhiqiyah" pageId="8" pageNumber="159">Latakia</collectingRegion>
and Jabla (
<bibRefCitation id="EF8139CDC72DFF8C07332C04F7C8FA3F" author="Kasparek, M." box="[786,960,1391,1418]" pageId="8" pageNumber="159" pagination="51 - 62" refId="ref16224" refString="Kasparek, M. (1995) The nesting of marine turtles on the coast of Syria. Zoology in the Middle East, 11, 51 - 62. https: // doi. org / 10.1080 / 09397140.1995.10637671" type="journal article" year="1995">Kasparek 1995</bibRefCitation>
).
</paragraph>
<paragraph id="8BAF443CC72DFF8C04E62CF8F75AFA18" blockId="8.[151,1437,1175,1850]" box="[199,850,1427,1454]" pageId="8" pageNumber="159">
<emphasis id="B964982EC72DFF8C04E62CF8F5C6FA18" bold="true" box="[199,462,1427,1453]" pageId="8" pageNumber="159">Citizen science record</emphasis>
: Ras Al-Baseet (iNaturalist 2023).
</paragraph>
</subSubSection>
<subSubSection id="C30A17B7C72DFF8C04E62CDCF56FF930" pageId="8" pageNumber="159" type="distribution">
<paragraph id="8BAF443CC72DFF8C04E62CDCF56FF930" blockId="8.[151,1437,1175,1850]" pageId="8" pageNumber="159">
<emphasis id="B964982EC72DFF8C04E62CDCF5B6FA64" bold="true" box="[199,446,1463,1489]" pageId="8" pageNumber="159">
Distribution in
<collectingCountry id="F30704ACC72DFF8C055F2CDCF5B6FA64" box="[382,446,1463,1489]" name="Syria" pageId="8" pageNumber="159">Syria</collectingCountry>
</emphasis>
: Foraging and nesting in Syria.
<taxonomicName id="4C103FBFC72DFF8C07052CDCF7D1FA64" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[804,985,1463,1489]" class="Testudines" family="Cheloniidae" genus="Chelonia" kingdom="Animalia" order="Cryptodira" pageId="8" pageNumber="159" phylum="Chordata" rank="species" species="mydas">
<emphasis id="B964982EC72DFF8C07052CDCF7D1FA64" box="[804,985,1463,1489]" italics="true" pageId="8" pageNumber="159">Chelonia mydas</emphasis>
</taxonomicName>
is found off the Syrian coast in different ages throughout the year. Some dead turtles were also seen at the beaches of Ras Al-Baseet, Jabla, Shati Afamia, Al-Awaj. Nesting has been reported for many sites on the sandy beaches at (1) Ras Al-Basit, (2) Um et-Toyour, (3) Wadi Kandil, (4)
<collectingRegion id="49D48ADEC72DFF8C05A02F48F5DDF988" box="[385,469,1571,1597]" country="Syria" name="Al Ladhiqiyah" pageId="8" pageNumber="159">Latakia</collectingRegion>
beach (Shkaifat and Sanawbar), (5) Banias (
<bibRefCitation id="EF8139CDC72DFF8C07E82F48F087F988" author="Rees, A. F. &amp; Saad, A. &amp; Jony, M." box="[969,1167,1571,1597]" pageId="8" pageNumber="159" pagination="456 - 459" refId="ref17483" refString="Rees, A. F., Saad, A. &amp; Jony, M. (2008 b) Discovery of a regionally important green turtle Chelonia mydas rookery in Syria. Oryx, 42, 456 - 459. https: // doi. org / 10.1017 / S 0030605308000926" type="journal article" year="2008">
Rees
<emphasis id="B964982EC72DFF8C00292F4FF048F988" box="[1032,1088,1571,1597]" italics="true" pageId="8" pageNumber="159">et al.</emphasis>
2008b
</bibRefCitation>
, 2009, 2010;
<bibRefCitation id="EF8139CDC72DFF8C01062F48F4DAF9D4" author="Saad, A. &amp; Rees, A. &amp; Sabour, W. &amp; Othman, A. &amp; Genaidi, C. &amp; Alkhateeb, N." pageId="8" pageNumber="159" refId="ref18975" refString="Saad, A., Rees, A., Sabour, W., Othman, A., Genaidi, C. &amp; Alkhateeb, N. (2010) Six years of survey (2004 - 2009) on reproduction of sea turtles on Shkaifat beach, Syrian coast (proposed as protected area). Rapports et proces-verbaux des reunions Commission internationale pour l'exploration scientifique de la Mer Mediterranee, 39, 651." type="journal volume" year="2010">
Saad
<emphasis id="B964982EC72DFF8C01452F4FF195F988" box="[1380,1437,1571,1597]" italics="true" pageId="8" pageNumber="159">et al.</emphasis>
2010
</bibRefCitation>
, 2020;
<bibRefCitation id="EF8139CDC72DFF8C05032F2CF592F9D4" author="Saad, A." box="[290,410,1607,1633]" pageId="8" pageNumber="159" pagination="108 - 110" refId="ref18593" refString="Saad, A. (2012) Importance of Lattakia Beach (Syria) as nesting area for marine turtles: Results of seven years of field survey. Scholarly Journal of Agricultural Science, 2, 108 - 110." type="journal article" year="2012">Saad 2012</bibRefCitation>
;
<figureCitation id="132B58B9C72DFF8C05822F2CF5EDF9D4" box="[419,485,1607,1633]" captionStart="FIGURE 8" captionStartId="10.[151,250,1390,1414]" captionTargetBox="[161,1429,191,1356]" captionTargetId="figure-21@10.[151,1436,181,1366]" captionTargetPageId="10" captionText="FIGURE 8. Nesting sites of Chelonia mydas in Syria. Black circles: previous records, blue circle: recently confirmed previous record." pageId="8" pageNumber="159">Fig. 8</figureCitation>
). There are four records in the sea off
<collectingRegion id="49D48ADEC72DFF8C07A62F2CF7D3F9D4" box="[903,987,1607,1633]" country="Syria" name="Al Ladhiqiyah" pageId="8" pageNumber="159">Latakia</collectingRegion>
(
<bibRefCitation id="EF8139CDC72DFF8C07C82F2CF0BDF9D4" author="Halpin, P. N. &amp; Read, A. J. &amp; Fujioka, E. &amp; Best, B. D. &amp; Donnelly, B. &amp; Hazen, L. J. &amp; Kot, C. &amp; Urian, K. &amp; LaBrecque, E. &amp; Dimatteo, A. &amp; Cleary, J. &amp; Good, C. &amp; Crowder, L. B. &amp; Hyrenbach, K. D." box="[1001,1205,1607,1633]" pageId="8" pageNumber="159" pagination="104 - 115" refId="ref15484" refString="Halpin, P. N., Read, A. J., Fujioka, E., Best, B. D., Donnelly, B., Hazen, L. J., Kot, C., Urian, K., LaBrecque, E., Dimatteo, A., Cleary, J., Good, C., Crowder, L. B. &amp; Hyrenbach, K. D. (2009) OBIS-SEAMAP: The world data center for marine mammal, sea bird, and sea turtle distributions. Oceanography, 22, 104 - 115. https: // doi. org / 10.5670 / oceanog. 2009.42" type="journal article" year="2009">
Halpin
<emphasis id="B964982EC72DFF8C001D2F23F07BF9D4" box="[1084,1139,1607,1633]" italics="true" pageId="8" pageNumber="159">et al.</emphasis>
2009
</bibRefCitation>
), one at Ras Al-Basit (iNaturalist 2023).
</paragraph>
</subSubSection>
<subSubSection id="C30A17B7C72DFF8C04E62FE4F779F88C" pageId="8" pageNumber="159" type="discussion">
<paragraph id="8BAF443CC72DFF8C04E62FE4F779F88C" blockId="8.[151,1437,1175,1850]" pageId="8" pageNumber="159">
<emphasis id="B964982EC72DFF8C04E62FE4F53CF91C" bold="true" box="[199,308,1679,1705]" pageId="8" pageNumber="159">Remarks</emphasis>
:
<collectingCountry id="F30704ACC72DFF8C05632FE4F575F91C" box="[322,381,1679,1705]" name="Syria" pageId="8" pageNumber="159">Syria</collectingCountry>
is considered the third-most important country for
<taxonomicName id="4C103FBFC72DFF8C07902FE4F011F91C" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[945,1049,1679,1705]" class="Testudines" family="Cheloniidae" genus="Chelonia" kingdom="Animalia" order="Cryptodira" pageId="8" pageNumber="159" phylum="Chordata" rank="species" species="mydas">
<emphasis id="B964982EC72DFF8C07902FE4F011F91C" box="[945,1049,1679,1705]" italics="true" pageId="8" pageNumber="159">C. mydas</emphasis>
</taxonomicName>
nesting in the Mediterranean, after
<collectingCountry id="F30704ACC72DFF8C04B62FD8F4F8F978" box="[151,240,1715,1741]" name="Turkey" pageId="8" pageNumber="159">Türkiye</collectingCountry>
and
<collectingCountry id="F30704ACC72DFF8C05072FD8F57FF978" box="[294,375,1715,1741]" name="Cyprus" pageId="8" pageNumber="159">Cyprus</collectingCountry>
(
<bibRefCitation id="EF8139CDC72DFF8C05A42FD8F637F978" author="Rees, A. F. &amp; Saad, A. &amp; Jony, M." box="[389,575,1715,1741]" pageId="8" pageNumber="159" pagination="233 - 243" refId="ref17609" refString="Rees, A. F., Saad, A. &amp; Jony, M. (2010) Syria. In: Casale, P. &amp; Margaritoulis, D. (Eds.), Sea Turtles in the Mediterranean: Distribution, Threats and Conservation Priorities. IUCN, Gland, pp. 233 - 243." type="book chapter" year="2010">
Rees
<emphasis id="B964982EC72DFF8C05E52FDFF5F4F978" box="[452,508,1715,1741]" italics="true" pageId="8" pageNumber="159">et al.</emphasis>
2010
</bibRefCitation>
;
<bibRefCitation id="EF8139CDC72DFF8C06682FD8F71FF978" author="Casale, P. &amp; Broderick, A. C. &amp; Caminas, J. A. &amp; Cardona, L. &amp; Carreras, C. &amp; Demetropoulos, A. &amp; Fuller, W. J. &amp; Godley, B. J. &amp; Hochscheid, S. &amp; Kaska, Y. &amp; Lazar, B. &amp; Margaritoulis, D. &amp; Panagopoulou, A. &amp; Rees, A. F. &amp; Tomas, J. &amp; Turkozan, O." box="[585,791,1715,1741]" pageId="8" pageNumber="159" pagination="229 - 267" refId="ref13981" refString="Casale, P., Broderick, A. C., Caminas, J. A., Cardona, L., Carreras, C., Demetropoulos, A., Fuller, W. J., Godley, B. J., Hochscheid, S., Kaska, Y., Lazar, B., Margaritoulis, D., Panagopoulou, A., Rees, A. F., Tomas, J. &amp; Turkozan, O. (2018) Mediterranean Sea turtles: Current knowledge and priorities for conservation and research. Endangered Species Research, 36, 229 - 267. https: // doi. org / 10.3354 / esr 00901" type="journal article" year="2018">
Casale
<emphasis id="B964982EC72DFF8C06BA2FDFF6DBF978" box="[667,723,1715,1741]" italics="true" pageId="8" pageNumber="159">et al.</emphasis>
2018
</bibRefCitation>
). More than 85 turtles stranded during 2005 on the Syrian coast, the majority were green turtles (
<bibRefCitation id="EF8139CDC72DFF8C06632FBCF6F3F944" author="Saad, A. &amp; Ali, A. &amp; Darwich, A." box="[578,763,1751,1777]" pageId="8" pageNumber="159" refId="ref18900" refString="Saad, A., Ali, A. &amp; Darwich, A. (2006) Marine turtle nesting survey, Syria 2005. In: Frick, M., Panagopoulou, A., Rees, A. F. &amp; Williams, K. (Eds.), Book of Abstracts of 26 th Annual Symposium on Sea Turtles. Sea Turtle Society, Athens, p. 320." type="journal volume" year="2006">
Saad
<emphasis id="B964982EC72DFF8C06A02FB3F6B1F944" box="[641,697,1751,1777]" italics="true" pageId="8" pageNumber="159">et al.</emphasis>
2006
</bibRefCitation>
;
<bibRefCitation id="EF8139CDC72DFF8C07242FBCF7DEF944" author="Saad, A. &amp; Rees, A." box="[773,982,1751,1778]" pageId="8" pageNumber="159" pagination="274 - 286" refId="ref18634" refString="Saad, A. &amp; Rees, A. (2020) Syria. In: Casale, P., Hochscheid, S., Kaska, Y. &amp; Panagopoulou, A. (Eds.), Sea Turtles in the Mediterranean region. IUCN-SSC Marine Turtle Specialist Group Regional Report, 2020, 274 - 286." type="journal article" year="2020">Saad &amp; Rees 2020</bibRefCitation>
,
<bibRefCitation id="EF8139CDC72DFF8C07C02FBCF016F944" author="Saad, A. &amp; Rees, A. F." box="[993,1054,1751,1777]" pageId="8" pageNumber="159" pagination="449 - 461" refId="ref18695" refString="Saad, A. &amp; Rees, A. F. (2022) Syria. In: Casale, P., Hochscheid, S., Jribi, I., Kaska, Y. &amp; Panagopoulou, A. (Eds.), Sea turtles in the Mediterranean region. 2021 IUCN-SSC Marine Turtle Specialist Group Regional Report, 2021, pp. 449 - 461." type="journal article" year="2022">2022</bibRefCitation>
). Morphological malformations in green turtle embryos and hatchlings were studied at Latakia Beach (
<bibRefCitation id="EF8139CDC72DFF8C07A72F90F035F8A0" author="Rees, A. &amp; Jony, M. &amp; Saad, A. &amp; Andrea, D. &amp; Phillott, A. D." box="[902,1085,1787,1813]" pageId="8" pageNumber="159" pagination="12 - 15" refId="ref17354" refString="Rees, A., Jony, M., Saad, A., Andrea, D. &amp; Phillott, A. D. (2021) Malformations in green turtle embryos and hatchlings in Syria. Marine Turtle Newsletter, 164, 12 - 15." type="journal article" year="2021">
Rees
<emphasis id="B964982EC72DFF8C07E52F97F7F3F8A0" box="[964,1019,1787,1813]" italics="true" pageId="8" pageNumber="159">et al.</emphasis>
2021
</bibRefCitation>
). Syrian
<taxonomicName id="4C103FBFC72DFF8C00BE2F90F10EF8A0" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1183,1286,1787,1813]" class="Testudines" family="Cheloniidae" genus="Chelonia" kingdom="Animalia" order="Cryptodira" pageId="8" pageNumber="159" phylum="Chordata" rank="species" species="mydas">
<emphasis id="B964982EC72DFF8C00BE2F90F10EF8A0" box="[1183,1286,1787,1813]" italics="true" pageId="8" pageNumber="159">C. mydas</emphasis>
</taxonomicName>
belong to the Mediterranean Regional Management Unit (
<bibRefCitation id="EF8139CDC72DFF8C06A62E74F76DF88C" author="Wallace, B. P. &amp; Posnik, Z. A. &amp; Hurley, B. J. &amp; DiMatteo, A. D. &amp; Bandimere, A. &amp; Rodriguez, I. &amp; Maxwell, S. M. &amp; Meyer, L. &amp; Brenner, H. &amp; Jensen, M. P. &amp; LaCasella, E. &amp; Shamblin, B. M. &amp; Abreu-Grobois, F. A. &amp; Stewart, K. R. &amp; Dutton, P. H. &amp; Barrios-Garrido, H. &amp; Dalleau, M. &amp; Dell'Amico, F. &amp; Eckert, K. L. &amp; FitzSimmons, N. N. &amp; Garcia-Cruz, M. &amp; Hays, G. C. &amp; Kelez, S. &amp; Lagueux, C. J. &amp; Madden Hof, C. A. &amp; Marco, A. &amp; Martins, S. L. T. &amp; Mobaraki, A. &amp; Mortimer, J. A. &amp; Nel, R. &amp; Phillott, A. D. &amp; Pilcher, N. J. &amp; Putman, N. F. &amp; Rees, A. F. &amp; Rguez-Baron, J. M. &amp; Seminoff, J. A. &amp; Swaminathan, A. &amp; Turkozan, O. &amp; Vargas, S. M. &amp; Vernet, P. D. &amp; Vilaca, S. &amp; Whiting, S. D. &amp; Hutchinson, B. J. &amp; Casale, P. &amp; Mast, R. B." box="[647,869,1823,1850]" pageId="8" pageNumber="159" pagination="209 - 223" refId="ref21120" refString="Wallace, B. P., Posnik, Z. A., Hurley, B. J., DiMatteo, A. D., Bandimere, A., Rodriguez, I., Maxwell, S. M., Meyer, L., Brenner, H., Jensen, M. P., LaCasella, E., Shamblin, B. M., Abreu-Grobois, F. A., Stewart, K. R., Dutton, P. H., Barrios-Garrido, H., Dalleau, M., Dell'Amico, F., Eckert, K. L., FitzSimmons, N. N., Garcia-Cruz, M., Hays, G. C., Kelez, S., Lagueux, C. J., Madden Hof, C. A., Marco, A., Martins, S. L. T., Mobaraki, A., Mortimer, J. A., Nel, R., Phillott, A. D., Pilcher, N. J., Putman, N. F., Rees, A. F., Rguez-Baron, J. M., Seminoff, J. A., Swaminathan, A., Turkozan, O., Vargas, S. M., Vernet, P. D., Vilaca, S., Whiting, S. D., Hutchinson, B. J., Casale, P. &amp; Mast, R. B. (2023) Marine turtle regional management units 2.0: An updated framework for conservation and research of wide-ranging megafauna species. Endangered Species Research, 52, 209 - 223. https: // doi. org / 10.3354 / ESR 01243" type="journal article" year="2023">
Wallace
<emphasis id="B964982EC72DFF8C06C82E4BF712F88C" box="[745,794,1823,1849]" italics="true" pageId="8" pageNumber="159">et al</emphasis>
. 2023
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,261 @@
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<mods:mods id="CEA82A871367EB58629C5DDDDAEDA02C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1540DD7238CC824DA22AB4FB981F6030">
<mods:title id="1DD088F73F870B8226271FDA28CCE3AA">Turtles and tortoises of Syria: Diversity, distribution, and conservation</mods:title>
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<mods:name id="E4B367A57F33E008E3BC263F1E616B81" type="personal">
<mods:role id="C93EC6407C16A2800119AF88AA04DE5A">
<mods:roleTerm id="37175FAFB6A2DC3996C4A3F3CCB3DDC3">Author</mods:roleTerm>
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<mods:namePart id="1D2E6182751D7571C58DCB3F3E3613D0">Aidek, Ahmad E.</mods:namePart>
<mods:affiliation id="08019B027AEE1C476EF95057838B3B6E">General Commission for Scientific Agricultural Research, Deir ez-Zor, Syria</mods:affiliation>
</mods:name>
<mods:name id="0713DE0ADD32D51E142CEAD76D8BC0C3" type="personal">
<mods:role id="DFAE861EA89CBE881D7F3087C0E55A4B">
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<mods:namePart id="9AA3651D354949C51F0217FD2D97AE1E">Saad, Adib</mods:namePart>
<mods:affiliation id="5F1A31F805CBFC518076CC998A06229D">Al-Manara University, Latakia, Syria</mods:affiliation>
</mods:name>
<mods:name id="89A60ED23114AB5B8477A3D85FDF5B4A" type="personal">
<mods:role id="889C1B7F49440DBBEC5E47959F81551D">
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<mods:namePart id="79D51333CAD858A733EE7F9908B65EB6">Jablonski, Daniel</mods:namePart>
<mods:affiliation id="FEE1601AEAB0EC07FB5888121DAD07CA">Department of Zoology, Comenius University in Bratislava, Ilkovičova 6, Mlynská dolina, 842 15 Bratislava, Slovakia</mods:affiliation>
</mods:name>
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<mods:namePart id="3AE1F8399698A861F9DD19C916972A47">Esterbauer, Hans</mods:namePart>
<mods:affiliation id="F29A2197FC8A892061D10254B4111A9B">Johann-Puch-Strasse 27 / III / 5, 4400 Steyr, Austria</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="79E94A44D58B0E3FCD9CDA4C8DF871B1">Fritz, Uwe</mods:namePart>
<mods:affiliation id="5AE4BE11F3101EFA8E3134A22A0F8FC7">Museum of Zoology, Senckenberg Dresden, A. B. Meyer Building, 01109 Dresden, Germany</mods:affiliation>
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<mods:date id="A10DF5C47B2D3040EE23C9EAC7B4A65B">2024</mods:date>
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</mods:mods>
<treatment id="03B9F52AC72EFF8804B62FD8F194FA18" LSID="urn:lsid:plazi:treatment:03B9F52AC72EFF8804B62FD8F194FA18" httpUri="http://treatment.plazi.org/id/03B9F52AC72EFF8804B62FD8F194FA18" lastPageId="12" lastPageNumber="163" pageId="11" pageNumber="162">
<subSubSection id="C30A17B7C72EFF8F04B62FD8F5D6F944" pageId="11" pageNumber="162" type="nomenclature">
<paragraph id="8BAF443CC72EFF8F04B62FD8F7ACF978" blockId="11.[151,932,1715,1777]" box="[151,932,1715,1742]" pageId="11" pageNumber="162">
<heading id="D0E7F350C72EFF8F04B62FD8F7ACF978" bold="true" box="[151,932,1715,1742]" fontSize="11" level="1" pageId="11" pageNumber="162" reason="1">
<taxonomicName id="4C103FBFC72EFF8F04B62FD8F7ACF978" ID-CoL="7GRRH" authority="Fritz, Baran, Budak &amp; Amthauer, 1998" authorityName="Fritz, Baran, Budak &amp; Amthauer" authorityYear="1998" box="[151,932,1715,1742]" class="Testudines" family="Emydidae" genus="Emys" kingdom="Animalia" order="Cryptodira" pageId="11" pageNumber="162" phylum="Chordata" rank="subSpecies" species="orbicularis" subSpecies="eiselti">
<emphasis id="B964982EC72EFF8F04B62FD8F7ACF978" bold="true" box="[151,932,1715,1742]" pageId="11" pageNumber="162">
<emphasis id="B964982EC72EFF8F04B62FD8F5BBF97B" bold="true" box="[151,435,1715,1742]" italics="true" pageId="11" pageNumber="162">Emys orbicularis eiselti</emphasis>
<bibRefCitation id="EF8139CDC72EFF8F059B2FD8F7ACF978" author="Fritz, U. &amp; Baran, I. &amp; Budak, A. &amp; Amthauer, E." box="[442,932,1715,1741]" pageId="11" pageNumber="162" pagination="103 - 121" refId="ref14717" refString="Fritz, U., Baran, I., Budak, A. &amp; Amthauer, E. (1998) Some notes on the morphology of Emys orbicularis in Anatolia, especially on E. o. luteofusca and E. o. colchica, with the description of a new subspecies from southeastern Turkey. Mertensiella, 10, 103 - 121." type="journal article" year="1998">Fritz, Baran, Budak &amp; Amthauer, 1998</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BAF443CC72EFF8F04B62FBCF5D6F944" blockId="11.[151,932,1715,1777]" box="[151,478,1751,1777]" pageId="11" pageNumber="162">
Eiselts Pond Turtle (
<figureCitation id="132B58B9C72EFF8F05A22FBCF5DDF944" box="[387,469,1751,1777]" captionStart="FIGURE 10" captionStartId="12.[151,250,961,985]" captionTargetBox="[227,1361,181,937]" captionTargetId="figure-21@12.[227,1361,181,937]" captionTargetPageId="12" captionText="FIGURE 10. Eiselts pond turtle, Emys orbicularis eiselti, Antakya, Türkiye (photo: Yusuf Bayrakcı)." pageId="11" pageNumber="162">Fig. 10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30A17B7C72EFF8F04B62E74F7F4F88C" box="[151,1020,1823,1850]" pageId="11" pageNumber="162" type="description">
<paragraph id="8BAF443CC72EFF8F04B62E74F7F4F88C" blockId="11.[151,1436,1823,2030]" box="[151,1020,1823,1850]" pageId="11" pageNumber="162">
<emphasis id="B964982EC72EFF8F04B62E74F6B8F88C" bold="true" box="[151,688,1823,1849]" pageId="11" pageNumber="162">
Provisional Red List Category (
<bibRefCitation id="EF8139CDC72EFF8F062E2E74F6A1F88C" author="TTWG &amp; Turtle Taxonomy Working Group" box="[527,681,1823,1849]" pageId="11" pageNumber="162" pagination="1 - 472" refId="ref20400" refString="TTWG [Turtle Taxonomy Working Group] (2021) Turtles of the World: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status (9 th Ed.). Chelonian Research Monographs, 8, 1 - 472. https: // doi. org / 10.3854 / crm. 8. checklist. atlas. v 9.2021" type="journal article" year="2021">TTWG 2021</bibRefCitation>
)
</emphasis>
: Critically Endangered (CR).
</paragraph>
</subSubSection>
<subSubSection id="C30A17B7C72EFF8804E62E28F659FB60" lastPageId="12" lastPageNumber="163" pageId="11" pageNumber="162" type="materials_examined">
<paragraph id="8BAF443CC72EFF8F04E62E28F795F8E8" blockId="11.[151,1436,1823,2030]" box="[199,925,1859,1886]" pageId="11" pageNumber="162">
<materialsCitation id="3B784E61C72EFF8F04E62E28F791F8E8" box="[199,921,1859,1886]" country="Turkey" location="Vilayet" pageId="11" pageNumber="162" specimenCount="1" stateProvince="Gaziantep" typeStatus="holotype">
<emphasis id="B964982EC72EFF8F04E62E28F568F8E8" bold="true" box="[199,352,1859,1885]" pageId="11" pageNumber="162">
<typeStatus id="54ABFA9EC72EFF8F04E62E28F50AF8E8" box="[199,258,1859,1885]" pageId="11" pageNumber="162">Type</typeStatus>
locality
</emphasis>
:
<locationDeviation id="9A1D5D1EC72EFF8F054E2E28F654F8E8" box="[367,604,1859,1886]" location="FevzipaSa" pageId="11" pageNumber="162">
<quantity id="4CE8E9D9C72EFF8F054E2E28F5BEF8E8" box="[367,438,1859,1885]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="11" pageNumber="162" unit="km" value="14.0">14 km</quantity>
NE FevzipaŞa
</locationDeviation>
,
<location id="8ECF12E7C72EFF8F06472E28F6BFF8E8" LSID="urn:lsid:plazi:treatment:03B9F52AC72EFF8804B62FD8F194FA18:8ECF12E7C72EFF8F06472E28F6BFF8E8" box="[614,695,1859,1885]" country="Turkey" name="Vilayet" pageId="11" pageNumber="162" stateProvince="Gaziantep">Vilayet</location>
<collectingRegion id="49D48ADEC72EFF8F069F2E28F73AF8E8" box="[702,818,1859,1885]" country="Turkey" name="Gaziantep" pageId="11" pageNumber="162">Gaziantep</collectingRegion>
,
<collectingCountry id="F30704ACC72EFF8F071C2E28F791F8E8" box="[829,921,1859,1885]" name="Turkey" pageId="11" pageNumber="162">Türkiye</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8BAF443CC72EFF8F04E62E0CF75CF87C" blockId="11.[151,1436,1823,2030]" pageId="11" pageNumber="162">
<emphasis id="B964982EC72EFF8F04E62E0CF576F834" bold="true" box="[199,382,1895,1921]" pageId="11" pageNumber="162">Type specimens</emphasis>
:
<materialsCitation id="3B784E61C72EFF8F05AB2E0CF79EF837" box="[394,918,1895,1922]" collectionCode="NMW" location="Naturhistorisches Museum" pageId="11" pageNumber="162" specimenCode="NMW18551" specimenCount="1" stateProvince="Wien">
<location id="8ECF12E7C72EFF8F05AB2E0CF6BBF837" LSID="urn:lsid:plazi:treatment:03B9F52AC72EFF8804B62FD8F194FA18:8ECF12E7C72EFF8F05AB2E0CF6BBF837" box="[394,691,1895,1922]" name="Naturhistorisches Museum" pageId="11" pageNumber="162" stateProvince="Wien">Naturhistorisches Museum</location>
<collectingRegion id="49D48ADEC72EFF8F06972E0CF6FCF834" box="[694,756,1895,1921]" country="Austria" name="Wien" pageId="11" pageNumber="162">Wien</collectingRegion>
,
<specimenCode id="DBB6EC47C72EFF8F06DD2E03F79EF837" box="[764,918,1895,1922]" collectionCode="NMW" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="162">NMW18551</specimenCode>
</materialsCitation>
:
<materialsCitation id="3B784E61C72EFF8F07BB2E0CF758F87C" collectionCode="NMW, ZDEU, MTD" county="Naturhistorisches Museum" location="Ege University" municipality="Zoological Department" pageId="11" pageNumber="162" specimenCode="NMW 18551, ZDEU 40, MTD 40468" specimenCount="3" stateProvince="Wien" typeStatus="holotype">
1,
<typeStatus id="54ABFA9EC72EFF8F078D2E0CF01AF834" box="[940,1042,1895,1921]" pageId="11" pageNumber="162" type="holotype">holotype</typeStatus>
;
<specimenCount id="9D168FB5C72EFF8F00382E0CF09EF837" box="[1049,1174,1895,1922]" count="3" pageId="11" pageNumber="162" type="generic" typeStatus="paratype">3 paratypes</specimenCount>
in the
<collectingCounty id="62CE3CB0C72EFF8F00F92E0CF4F1F813" pageId="11" pageNumber="162">Naturhistorisches Museum</collectingCounty>
<collectingRegion id="49D48ADEC72EFF8F05212EE0F547F810" box="[256,335,1931,1957]" country="Austria" name="Wien" pageId="11" pageNumber="162">Vienna</collectingRegion>
(
<specimenCode id="DBB6EC47C72EFF8F057C2EE0F5F2F813" box="[349,506,1931,1958]" collectionCode="NMW" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="162">NMW 18551</specimenCode>
:2), the
<collectingMunicipality id="6BCBDE46C72EFF8F066B2EE0F747F813" box="[586,847,1931,1958]" pageId="11" pageNumber="162">Zoological Department</collectingMunicipality>
of the
<location id="8ECF12E7C72EFF8F07BF2EE7F04FF810" LSID="urn:lsid:plazi:treatment:03B9F52AC72EFF8804B62FD8F194FA18:8ECF12E7C72EFF8F07BF2EE7F04FF810" box="[926,1095,1931,1958]" county="Naturhistorisches Museum" municipality="Zoological Department" name="Ege University" pageId="11" pageNumber="162" stateProvince="Wien">Ege University</location>
,
<collectingRegion id="49D48ADEC72EFF8F00722EE0F098F810" box="[1107,1168,1931,1957]" country="Turkey" name="Izmir" pageId="11" pageNumber="162">Izmir</collectingRegion>
(
<specimenCode id="DBB6EC47C72EFF8F00BF2EE0F119F810" box="[1182,1297,1931,1957]" collectionCode="ZDEU" country="Turkey" name="Zoology Department, Ege University" pageId="11" pageNumber="162">ZDEU 40</specimenCode>
/1972:1) and the
<location id="8ECF12E7C72EFF8F04E12EDBF5A0F87C" LSID="urn:lsid:plazi:treatment:03B9F52AC72EFF8804B62FD8F194FA18:8ECF12E7C72EFF8F04E12EDBF5A0F87C" box="[192,424,1967,1994]" county="Naturhistorisches Museum" municipality="Zoological Department" name="Museum of Zoology" pageId="11" pageNumber="162" stateProvince="Wien">Museum of Zoology</location>
,
<location id="8ECF12E7C72EFF8F05922EC4F6AFF87C" LSID="urn:lsid:plazi:treatment:03B9F52AC72EFF8804B62FD8F194FA18:8ECF12E7C72EFF8F05922EC4F6AFF87C" box="[435,679,1967,1993]" county="Naturhistorisches Museum" municipality="Zoological Department" name="Senckenberg Dresden" pageId="11" pageNumber="162" stateProvince="Wien">Senckenberg Dresden</location>
(
<specimenCode id="DBB6EC47C72EFF8F06972EC4F740F87C" box="[694,840,1967,1993]" collectionCode="MTD" country="Germany" httpUri="http://grbio.org/cool/4j3t-w40g" name="Museum of Zoology Senckenberg Dresden" pageId="11" pageNumber="162">MTD 40468</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BAF443CC72EFF8F04E62EB8F6C5F858" blockId="11.[151,1436,1823,2030]" box="[199,717,2003,2030]" pageId="11" pageNumber="162">
<emphasis id="B964982EC72EFF8F04E62EB8F54EF858" bold="true" box="[199,326,2003,2029]" pageId="11" pageNumber="162">Chorotype</emphasis>
: Turano-European-Mediterranean.
</paragraph>
<caption id="DF6F14B4C729FF8804B62AAAF0AAFC6C" box="[151,1186,961,985]" pageId="12" pageNumber="163" startId="12.[151,250,961,985]" targetBox="[227,1361,181,937]" targetPageId="12" targetType="figure">
<paragraph id="8BAF443CC729FF8804B62AAAF0AAFC6C" blockId="12.[151,1186,961,985]" box="[151,1186,961,985]" pageId="12" pageNumber="163">
<emphasis id="B964982EC729FF8804B62AAAF528FC6C" bold="true" box="[151,288,961,985]" pageId="12" pageNumber="163">FIGURE 10.</emphasis>
Eiselts pond turtle,
<taxonomicName id="4C103FBFC729FF8805D32AA9F74EFC6C" authority=", Antakya" authorityName="Antakya" box="[498,838,961,985]" class="Testudines" family="Emydidae" genus="Emys" kingdom="Animalia" order="Cryptodira" pageId="12" pageNumber="163" phylum="Chordata" rank="subSpecies" species="orbicularis" subSpecies="eiselti">
<emphasis id="B964982EC729FF8805D32AA9F6EAFC6C" box="[498,738,961,985]" italics="true" pageId="12" pageNumber="163">Emys orbicularis eiselti</emphasis>
, Antakya
</taxonomicName>
, Türkiye (photo: Yusuf Bayrakcı).
</paragraph>
</caption>
<paragraph id="8BAF443CC729FF8804E62D40F048FBDC" blockId="12.[151,1437,1067,1454]" pageId="12" pageNumber="163">
<emphasis id="B964982EC729FF8804E62D40F58DFBF0" bold="true" box="[199,389,1067,1093]" pageId="12" pageNumber="163">First record for</emphasis>
<materialsCitation id="3B784E61C729FF8805AE2D40F034FBDC" collectingDate="1980-03-25" collectionCode="MTD" country="Syria" county="Senckenberg Dresden" location="Between Al-Kareem" municipality="Museum of Zoology" pageId="12" pageNumber="163" specimenCode="MTD 39470" specimenCount="1">
<emphasis id="B964982EC729FF8805AE2D40F5C7FBF0" bold="true" box="[399,463,1067,1093]" pageId="12" pageNumber="163">
<collectingCountry id="F30704ACC729FF8805AE2D40F5C7FBF0" box="[399,463,1067,1093]" name="Syria" pageId="12" pageNumber="163">Syria</collectingCountry>
</emphasis>
:
<location id="8ECF12E7C729FF8805C12D47F6CFFBF0" LSID="urn:lsid:plazi:treatment:03B9F52AC72EFF8804B62FD8F194FA18:8ECF12E7C729FF8805C12D47F6CFFBF0" box="[480,711,1067,1094]" country="Syria" county="Senckenberg Dresden" municipality="Museum of Zoology" name="Between Al-Kareem" pageId="12" pageNumber="163">Between Al-Kareem</location>
and
<location id="8ECF12E7C729FF8807232D40F79DFBF0" LSID="urn:lsid:plazi:treatment:03B9F52AC72EFF8804B62FD8F194FA18:8ECF12E7C729FF8807232D40F79DFBF0" box="[770,917,1067,1093]" country="Syria" county="Senckenberg Dresden" municipality="Museum of Zoology" name="Bab Al-Taqa" pageId="12" pageNumber="163">Bab Al-Taqa</location>
, collected
<date id="FFAE62FCC729FF8800302D40F0C9FBF0" box="[1041,1217,1067,1094]" pageId="12" pageNumber="163" value="1980-03-25">
<collectingDate id="EFEA9B14C729FF8800302D40F0C9FBF0" box="[1041,1217,1067,1094]" pageId="12" pageNumber="163" value="1980-03-25">25 March 1980</collectingDate>
</date>
(
<bibRefCitation id="EF8139CDC729FF8800F32D40F19FFBF0" author="Kinzelbach, R." box="[1234,1431,1067,1093]" pageId="12" pageNumber="163" pagination="385 - 419" refId="ref16358" refString="Kinzelbach, R. (1988) Die Europaische Sumpfschildkrote (Emys orbicularis) im Einzugsgebiet des Rheins. Zeitschrift fur Angewandte Zoologie, 75, 385 - 419." type="journal article" year="1988">Kinzelbach 1988</bibRefCitation>
; voucher specimen in the
<collectingMunicipality id="6BCBDE46C729FF88058E2D3BF69EFBDC" box="[431,662,1103,1130]" pageId="12" pageNumber="163">Museum of Zoology</collectingMunicipality>
,
<collectingCounty id="62CE3CB0C729FF8806802D24F791FBDC" box="[673,921,1103,1129]" pageId="12" pageNumber="163">Senckenberg Dresden</collectingCounty>
,
<specimenCode id="DBB6EC47C729FF8807852D3BF03CFBDC" box="[932,1076,1103,1130]" collectionCode="MTD" country="Germany" httpUri="http://grbio.org/cool/4j3t-w40g" name="Museum of Zoology Senckenberg Dresden" pageId="12" pageNumber="163">MTD 39470</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BAF443CC729FF8804E62D18F659FB60" blockId="12.[151,1437,1067,1454]" pageId="12" pageNumber="163">
<emphasis id="B964982EC729FF8804E62D18F579FB38" bold="true" box="[199,369,1139,1165]" pageId="12" pageNumber="163">Recent record</emphasis>
: East of Jourin. One individual was found dead east of Jourin near the Orontes River in 1997. The shell was preserved in a large and valuable collection of other specimens and photographs (Aidek collection), but all was lost during the war in
<collectingCountry id="F30704ACC729FF88062F2DD0F645FB60" box="[526,589,1211,1237]" name="Syria" pageId="12" pageNumber="163">Syria</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C30A17B7C729FF8804E62DB4F104FB4C" box="[199,1292,1247,1274]" pageId="12" pageNumber="163" type="distribution">
<paragraph id="8BAF443CC729FF8804E62DB4F104FB4C" blockId="12.[151,1437,1067,1454]" box="[199,1292,1247,1274]" pageId="12" pageNumber="163">
<emphasis id="B964982EC729FF8804E62DB4F5B7FB4C" bold="true" box="[199,447,1247,1273]" pageId="12" pageNumber="163">
Distribution in
<collectingCountry id="F30704ACC729FF88055E2DB4F5B7FB4C" box="[383,447,1247,1273]" name="Syria" pageId="12" pageNumber="163">Syria</collectingCountry>
</emphasis>
: Very rare. Recorded only in the middle and upper Orontes Basin (
<figureCitation id="132B58B9C729FF88008C2DB4F0F6FB4C" box="[1197,1278,1247,1273]" captionStart="FIGURE 11" captionStartId="13.[151,250,1388,1412]" captionTargetBox="[161,1429,191,1357]" captionTargetId="figure-17@13.[151,1436,181,1364]" captionTargetPageId="13" captionText="FIGURE 11. Records of Emys orbicularis eiselti in Syria. Black circle: vouchered previous record, red circle: new record." pageId="12" pageNumber="163">Fig. 11</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30A17B7C729FF8804E62C68F194FA18" pageId="12" pageNumber="163" type="discussion">
<paragraph id="8BAF443CC729FF8804E62C68F194FA18" blockId="12.[151,1437,1067,1454]" pageId="12" pageNumber="163">
<emphasis id="B964982EC729FF8804E62C68F53CFAA8" bold="true" box="[199,308,1283,1309]" pageId="12" pageNumber="163">Remarks</emphasis>
: This subspecies was described from
<collectingRegion id="49D48ADEC729FF8806D02C68F7D9FAA8" box="[753,977,1283,1309]" country="Turkey" name="Gaziantep" pageId="12" pageNumber="163">Gaziantep Province</collectingRegion>
(
<collectingCountry id="F30704ACC729FF8807C22C68F04AFAA8" box="[995,1090,1283,1309]" name="Turkey" pageId="12" pageNumber="163">Türkiye</collectingCountry>
), based on only four museum specimens collected at two different sites in 1966 and 1972 (
<bibRefCitation id="EF8139CDC729FF88070E2C4CF7EBFAF4" author="Fritz, U. &amp; Baran, I. &amp; Budak, A. &amp; Amthauer, E." box="[815,995,1319,1345]" pageId="12" pageNumber="163" pagination="103 - 121" refId="ref14717" refString="Fritz, U., Baran, I., Budak, A. &amp; Amthauer, E. (1998) Some notes on the morphology of Emys orbicularis in Anatolia, especially on E. o. luteofusca and E. o. colchica, with the description of a new subspecies from southeastern Turkey. Mertensiella, 10, 103 - 121." type="journal article" year="1998">
Fritz
<emphasis id="B964982EC729FF88074A2C43F7AAFAF4" box="[875,930,1319,1345]" italics="true" pageId="12" pageNumber="163">et al.</emphasis>
1998
</bibRefCitation>
). Despite many field surveys from
<quantity id="4CE8E9D9C729FF8801452C4CF502FAD0" metricMagnitude="1" metricUnit="m" metricValue="5.1053999999999995" metricValueMax="5.12826" metricValueMin="5.08254" pageId="12" pageNumber="163" unit="in" value="2010.0" valueMax="2019.0" valueMin="2001.0">2001 to 2019 in</quantity>
the
<collectingRegion id="49D48ADEC729FF88051A2C20F5F4FAD0" box="[315,508,1355,1381]" country="Turkey" name="Gaziantep" pageId="12" pageNumber="163">Gaziantep region</collectingRegion>
and east of the Amanos Mountains,
<collectingCountry id="F30704ACC729FF8807B92C20F7FCFAD0" box="[920,1012,1355,1381]" name="Turkey" pageId="12" pageNumber="163">Türkiye</collectingCountry>
, only
<specimenCount id="9D168FB5C729FF8800192C20F0D8FAD0" box="[1080,1232,1355,1381]" count="18" pageId="12" pageNumber="163" type="generic">18 specimens</specimenCount>
were found at the mouth of the Orontes River (
<collectingRegion id="49D48ADEC729FF8805FE2C04F699FA3C" box="[479,657,1391,1417]" country="Turkey" name="Hatay" pageId="12" pageNumber="163">Hatay Province</collectingRegion>
,
<collectingCountry id="F30704ACC729FF8806BC2C04F6F1FA3C" box="[669,761,1391,1417]" name="Turkey" pageId="12" pageNumber="163">Türkiye</collectingCountry>
;
<bibRefCitation id="EF8139CDC729FF8807252C04F7CAFA3C" author="Ayaz, D. &amp; Bayrakci, Y. &amp; Cicek, K. &amp; Ihlow, F. &amp; Tok, C. V. &amp; Fritz, U." box="[772,962,1391,1417]" pageId="12" pageNumber="163" pagination="222 - 230" refId="ref13246" refString="Ayaz, D., Bayrakci, Y., Cicek, K., Ihlow, F., Tok, C. V. &amp; Fritz, U. (2021) On the brink of extinction: Results of a 20 - year quest for Eiselt's pond turtle (Emys orbicularis eiselti) in southeastern Turkey. Chelonian Conservation and Biology, 20, 222 - 230. https: // doi. org / 10.2744 / CCB- 1505.1" type="journal article" year="2021">
Ayaz
<emphasis id="B964982EC729FF8807642C1BF776FA3C" box="[837,894,1391,1417]" italics="true" pageId="12" pageNumber="163">et al.</emphasis>
2021
</bibRefCitation>
). Fieldwork in the Jourin area in Al-Ghab Plain,
<collectingCountry id="F30704ACC729FF8804FE2CF8F515FA18" box="[223,285,1427,1453]" name="Syria" pageId="12" pageNumber="163">Syria</collectingCountry>
, in
<date id="FFAE62FCC729FF8805622CF8F5B5FA18" box="[323,445,1427,1453]" pageId="12" pageNumber="163" value="2009-04">April 2009</date>
failed to find any additional
<emphasis id="B964982EC729FF8807232CFFF788FA18" box="[770,896,1427,1453]" italics="true" pageId="12" pageNumber="163">E. o. eiselti</emphasis>
, while many
<taxonomicName id="4C103FBFC729FF8800382CFFF0E6FA18" box="[1049,1262,1427,1453]" class="Testudines" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Cryptodira" pageId="12" pageNumber="163" phylum="Chordata" rank="species" species="rivulata">
<emphasis id="B964982EC729FF8800382CFFF0E6FA18" box="[1049,1262,1427,1453]" italics="true" pageId="12" pageNumber="163">Mauremys rivulata</emphasis>
</taxonomicName>
were recorded.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,187 @@
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<mods:title id="6548CED76B5A4A1474CA5641912A7DE2">Turtles and tortoises of Syria: Diversity, distribution, and conservation</mods:title>
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<mods:namePart id="77D0962CE996D2937D47EB4CE2AEA29B">Aidek, Ahmad E.</mods:namePart>
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<mods:affiliation id="10FA0035F1280267AD48FCE4C33D691B">Al-Manara University, Latakia, Syria</mods:affiliation>
</mods:name>
<mods:name id="8A07CC30D3A9FFBE69B1E0FD4630A92C" type="personal">
<mods:role id="43D043DDF5A12EC6863C04CB5EB60D8D">
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<mods:namePart id="53E04740DE76F520825F8E28A8229E5F">Jablonski, Daniel</mods:namePart>
<mods:affiliation id="B26958AC4C1B1194DA0AD26FE762968E">Department of Zoology, Comenius University in Bratislava, Ilkovičova 6, Mlynská dolina, 842 15 Bratislava, Slovakia</mods:affiliation>
</mods:name>
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<mods:affiliation id="817F782DA078C1B815BD3666441187FD">Johann-Puch-Strasse 27 / III / 5, 4400 Steyr, Austria</mods:affiliation>
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</mods:role>
<mods:namePart id="B1AB4F0641462B1373E1A18BC216AF1E">Fritz, Uwe</mods:namePart>
<mods:affiliation id="242D1E1BF0ABD7E197925FE77EE2CA91">Museum of Zoology, Senckenberg Dresden, A. B. Meyer Building, 01109 Dresden, Germany</mods:affiliation>
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<treatment id="03B9F52AC72FFF8F04B62F2FF12EFB04" LSID="urn:lsid:plazi:treatment:03B9F52AC72FFF8F04B62F2FF12EFB04" httpUri="http://treatment.plazi.org/id/03B9F52AC72FFF8F04B62F2FF12EFB04" lastPageId="11" lastPageNumber="162" pageId="10" pageNumber="161">
<subSubSection id="C30A17B7C72FFF8E04B62F2FF687F937" pageId="10" pageNumber="161" type="nomenclature">
<paragraph id="8BAF443CC72FFF8E04B62F2FF666F9EB" blockId="10.[151,655,1604,1666]" box="[151,622,1604,1631]" pageId="10" pageNumber="161">
<heading id="D0E7F350C72FFF8E04B62F2FF666F9EB" bold="true" box="[151,622,1604,1631]" fontSize="11" level="1" pageId="10" pageNumber="161" reason="1">
<taxonomicName id="4C103FBFC72FFF8E04B62F2FF666F9EB" ID-CoL="34XB9" authority="(Vandelli, 1761)" baseAuthorityName="Vandelli" baseAuthorityYear="1761" box="[151,622,1604,1631]" class="Testudines" family="Dermochelyidae" genus="Dermochelys" kingdom="Animalia" order="Cryptodira" pageId="10" pageNumber="161" phylum="Chordata" rank="species" species="coriacea">
<emphasis id="B964982EC72FFF8E04B62F2FF666F9EB" bold="true" box="[151,622,1604,1631]" pageId="10" pageNumber="161">
<emphasis id="B964982EC72FFF8E04B62F2FF5ACF9EB" bold="true" box="[151,420,1604,1631]" italics="true" pageId="10" pageNumber="161">Dermochelys coriacea</emphasis>
(Vandelli, 1761)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BAF443CC72FFF8E04B62F03F687F937" blockId="10.[151,655,1604,1666]" box="[151,655,1640,1666]" pageId="10" pageNumber="161">
Leatherback, Leatherback Sea Turtle (
<figureCitation id="132B58B9C72FFF8E06622F03F68EF937" box="[579,646,1640,1666]" captionStart="FIGURE 9" captionStartId="11.[151,250,961,985]" captionTargetBox="[227,1361,181,937]" captionTargetId="figure-17@11.[227,1361,181,937]" captionTargetPageId="11" captionText="FIGURE 9. Leatherback sea turtle, Dermochelys coriacea, Port of Jabla (photo: Adib Saad)." pageId="10" pageNumber="161">Fig. 9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30A17B7C72FFF8E04B62FDBF09EF97F" box="[151,1174,1712,1739]" pageId="10" pageNumber="161" type="description">
<paragraph id="8BAF443CC72FFF8E04B62FDBF09EF97F" blockId="10.[151,1437,1712,1954]" box="[151,1174,1712,1739]" pageId="10" pageNumber="161">
<emphasis id="B964982EC72FFF8E04B62FDBF5C2F97F" bold="true" box="[151,458,1712,1738]" pageId="10" pageNumber="161">IUCN Red List Category:</emphasis>
Vulnerable (VU A2bd; no Mediterranean breeding population).
</paragraph>
</subSubSection>
<subSubSection id="C30A17B7C72FFF8F04E62FBFF732FBDC" lastPageId="11" lastPageNumber="162" pageId="10" pageNumber="161" type="materials_examined">
<paragraph id="8BAF443CC72FFF8E04E62FBFF595F8EF" blockId="10.[151,1437,1712,1954]" pageId="10" pageNumber="161">
<emphasis id="B964982EC72FFF8E04E62FBFF564F95B" bold="true" box="[199,364,1748,1774]" pageId="10" pageNumber="161">
<typeStatus id="54ABFA9EC72FFF8E04E62FBFF50AF95B" box="[199,258,1748,1774]" pageId="10" pageNumber="161">Type</typeStatus>
locality:
</emphasis>
“Maris Tyrrheni oram in agro Laurentiano” [
<collectingCountry id="F30704ACC72FFF8E07572FBFF7B9F95B" box="[886,945,1748,1774]" name="Italy" pageId="10" pageNumber="161">Italy</collectingCountry>
]. Restricted to “Palermo,
<collectingRegion id="49D48ADEC72FFF8E00FB2FBFF116F95B" box="[1242,1310,1748,1774]" country="Italy" name="Sicilia" pageId="10" pageNumber="161">Sicily</collectingRegion>
” [
<collectingCountry id="F30704ACC72FFF8E01192FBFF178F95B" box="[1336,1392,1748,1774]" name="Italy" pageId="10" pageNumber="161">Italy</collectingCountry>
] by
<bibRefCitation id="EF8139CDC72FFF8E04B62F93F5AFF8A6" author="Smith, H. M. &amp; Taylor, E. H." box="[151,423,1784,1811]" pageId="10" pageNumber="161" pagination="313 - 380" refId="ref19489" refString="Smith, H. M. &amp; Taylor, E. H. (1950 a) Type localities of Mexican reptiles and amphibians. University of Kansas Science Bulletin, 33, 313 - 380." type="journal article" year="1950">Smith &amp; Taylor (1950a)</bibRefCitation>
; to “la côte romain (
<collectingCountry id="F30704ACC72FFF8E06AA2F93F6C2F8A7" box="[651,714,1784,1810]" name="Italy" pageId="10" pageNumber="161">Italie</collectingCountry>
), Mer Tyrrhénienne, Méditerranée occidentale” (
<collectingCountry id="F30704ACC72FFF8E00CC2F93F12BF8A7" box="[1261,1315,1784,1810]" name="Italy" pageId="10" pageNumber="161">Italy</collectingCountry>
) by
<bibRefCitation id="EF8139CDC72FFF8E01742F92F54AF883" author="Fretey, J. &amp; Bour, R." pageId="10" pageNumber="161" pagination="193 - 205" refId="ref14558" refString="Fretey, J. &amp; Bour, R. (1980) Redecouverte du type de Dermochelys coriacea (Vandelli) (Testudinata, Dermochelyidae). Bollettino di Zoologia, 47, 193 - 205. https: // doi. org / 10.1080 / 11250008009440336" type="journal article" year="1980">Fretey &amp; Bour (1980)</bibRefCitation>
; and to “Laurentum, between Lido di Ostia and Tor Paterno, shore of the Tyrrhenian Sea,
<collectingCountry id="F30704ACC72FFF8E01182E77F167F883" box="[1337,1391,1820,1846]" name="Italy" pageId="10" pageNumber="161">Italy</collectingCountry>
” by
<bibRefCitation id="EF8139CDC72FFF8E04B62E2AF590F8EF" author="Bour, R. &amp; Dubois, A." box="[151,408,1856,1883]" pageId="10" pageNumber="161" pagination="356 - 361" refId="ref13827" refString="Bour, R. &amp; Dubois, A. (1984) Nomenclatural availability of Testudo coriacea Vandelli, 1761: A case against a rigid application of the rules to old, well-known zoological works. Journal of Herpetology, 17, 356 - 361. https: // doi. org / 10.2307 / 1563588" type="journal article" year="1984">Bour &amp; Dubois (1984)</bibRefCitation>
.
</paragraph>
<paragraph id="8BAF443CC72FFF8E04E62E0FF041F8CB" blockId="10.[151,1437,1712,1954]" box="[199,1097,1892,1919]" pageId="10" pageNumber="161">
<emphasis id="B964982EC72FFF8E04E62E0FF589F8CB" bold="true" box="[199,385,1892,1918]" pageId="10" pageNumber="161">Type specimen:</emphasis>
<materialsCitation id="3B784E61C72FFF8E05A92E0EF04EF8CB" box="[392,1094,1892,1919]" location="Museo di Zoologia Pavia" pageId="10" pageNumber="161" specimenCount="1" typeStatus="holotype">
<location id="8ECF12E7C72FFF8E05A92E0EF6ADF8CB" LSID="urn:lsid:plazi:treatment:03B9F52AC72FFF8F04B62F2FF12EFB04:8ECF12E7C72FFF8E05A92E0EF6ADF8CB" box="[392,677,1892,1919]" name="Museo di Zoologia Pavia" pageId="10" pageNumber="161">Museo di Zoologia Pavia</location>
, specimen without number,
<typeStatus id="54ABFA9EC72FFF8E07C12E0FF04EF8CB" box="[992,1094,1892,1918]" pageId="10" pageNumber="161" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
<paragraph id="8BAF443CC72FFF8E04E62EE3F6A7F817" blockId="10.[151,1437,1712,1954]" box="[199,687,1928,1954]" pageId="10" pageNumber="161">
<emphasis id="B964982EC72FFF8E04E62EE3F558F817" bold="true" box="[199,336,1928,1954]" pageId="10" pageNumber="161">Chorotype:</emphasis>
Circumglobal (Cosmopolitan).
</paragraph>
<caption id="DF6F14B4C72EFF8F04B62AAAF04DFC6C" box="[151,1093,961,985]" pageId="11" pageNumber="162" startId="11.[151,250,961,985]" targetBox="[227,1361,181,937]" targetPageId="11" targetType="figure">
<paragraph id="8BAF443CC72EFF8F04B62AAAF04DFC6C" blockId="11.[151,1093,961,985]" box="[151,1093,961,985]" pageId="11" pageNumber="162">
<emphasis id="B964982EC72EFF8F04B62AAAF51BFC6C" bold="true" box="[151,275,961,985]" pageId="11" pageNumber="162">FIGURE 9.</emphasis>
Leatherback sea turtle,
<taxonomicName id="4C103FBFC72EFF8F06272AAAF729FC6D" authority=", Port" authorityName="Port" box="[518,801,961,985]" class="Testudines" family="Dermochelyidae" genus="Dermochelys" kingdom="Animalia" order="Cryptodira" pageId="11" pageNumber="162" phylum="Chordata" rank="species" species="coriacea">
<emphasis id="B964982EC72EFF8F06272AAAF6E3FC6C" box="[518,747,961,985]" italics="true" pageId="11" pageNumber="162">Dermochelys coriacea</emphasis>
, Port
</taxonomicName>
of Jabla (photo: Adib Saad).
</paragraph>
</caption>
<paragraph id="8BAF443CC72EFF8F04E62D40F732FBDC" blockId="11.[151,1436,1067,1202]" pageId="11" pageNumber="162">
<emphasis id="B964982EC72EFF8F04E62D40F5E8FBF0" bold="true" box="[199,480,1067,1093]" pageId="11" pageNumber="162">
First records for
<collectingCountry id="F30704ACC72EFF8F05B92D40F5D4FBF0" box="[408,476,1067,1093]" name="Syria" pageId="11" pageNumber="162">Syria</collectingCountry>
:
</emphasis>
At about
<quantity id="4CE8E9D9C72EFF8F06742D40F6ACFBF0" box="[597,676,1067,1094]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="11" pageNumber="162" unit="km" value="2.5">2.5 km</quantity>
offshore Jabla town (
<bibRefCitation id="EF8139CDC72EFF8F07BC2D40F053FBF0" author="Rees, A. F. &amp; Saad, A. &amp; Jony, M." box="[925,1115,1067,1093]" pageId="11" pageNumber="162" refId="ref17403" refString="Rees, A. F., Saad, A. &amp; Jony, M. (2004) First record of a leatherback turtle in Syria. Marine Turtle Newsletter, 106, 13." type="journal volume" year="2004">
Rees
<emphasis id="B964982EC72EFF8F07FC2D47F01FFBF0" box="[989,1047,1067,1093]" italics="true" pageId="11" pageNumber="162">et al.</emphasis>
2004
</bibRefCitation>
), another record was off the Al-Kabeer River, Al-Bassah, Snaoubar (
<bibRefCitation id="EF8139CDC72EFF8F06782D24F726FBDC" author="Jony, M. &amp; Rees, F. A." box="[601,814,1103,1130]" pageId="11" pageNumber="162" pagination="11 - 13" refId="ref16161" refString="Jony, M. &amp; Rees, F. A. (2022) Additional records of leatherback turtles (Dermochelys coriacea) from Syria. MedTurtle Bulletin, 2, 11 - 13." type="journal article" year="2022">Jony &amp; Rees 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30A17B7C72EFF8F04E62D18F0BDFB38" box="[199,1205,1139,1166]" pageId="11" pageNumber="162" type="distribution">
<paragraph id="8BAF443CC72EFF8F04E62D18F0BDFB38" blockId="11.[151,1436,1067,1202]" box="[199,1205,1139,1166]" pageId="11" pageNumber="162">
<emphasis id="B964982EC72EFF8F04E62D18F5C0FB38" bold="true" box="[199,456,1139,1165]" pageId="11" pageNumber="162">
Distribution in
<collectingCountry id="F30704ACC72EFF8F055E2D18F5CBFB38" box="[383,451,1139,1165]" name="Syria" pageId="11" pageNumber="162">Syria</collectingCountry>
:
</emphasis>
The species is foraging in Syrian waters; nesting is not confirmed.
</paragraph>
</subSubSection>
<subSubSection id="C30A17B7C72EFF8F04E62DFCF12EFB04" box="[199,1318,1175,1202]" pageId="11" pageNumber="162" type="discussion">
<paragraph id="8BAF443CC72EFF8F04E62DFCF12EFB04" blockId="11.[151,1436,1067,1202]" box="[199,1318,1175,1202]" pageId="11" pageNumber="162">
<emphasis id="B964982EC72EFF8F04E62DFCF535FB04" bold="true" box="[199,317,1175,1201]" pageId="11" pageNumber="162">Remarks:</emphasis>
We follow
<bibRefCitation id="EF8139CDC72EFF8F05E12DFCF652FB04" author="Fretey, T." box="[448,602,1175,1201]" pageId="11" pageNumber="162" pagination="46 - 69" refId="ref14518" refString="Fretey, T. (2022) The authorship and date of the familial nomen Dermochelyidae (Chelonii). Bionomina, 31, 46 - 69. https: // doi. org / 10.11646 / bionomina. 31.1.3" type="journal article" year="2022">Frétey (2022)</bibRefCitation>
and accept
<bibRefCitation id="EF8139CDC72EFF8F06FF2DF3F76EFB04" author="Baur, G." box="[734,870,1175,1202]" pageId="11" pageNumber="162" pagination="44 - 45" refId="ref13393" refString="Baur, G. (1888) Dermochelys, Dermatochelys oder Sphargis. Zoologischer Anzeiger, 11, 44 - 45." type="journal article" year="1888">Baur (1888)</bibRefCitation>
as author of the name
<taxonomicName id="4C103FBFC72EFF8F00442DFCF12AFB04" authorityName="Baur" authorityYear="1888" box="[1125,1314,1175,1201]" class="Testudines" family="Dermochelyidae" kingdom="Animalia" order="Cryptodira" pageId="11" pageNumber="162" phylum="Chordata" rank="family">Dermochelyidae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,572 @@
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<mods:title id="6ED8A5CB4EADEC470CB76C21120D149F">Turtles and tortoises of Syria: Diversity, distribution, and conservation</mods:title>
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<mods:name id="CCFB87599F8FF8D3D487B83EAEBB73FB" type="personal">
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<mods:namePart id="1A2B9C292E50750DCF80A4FFB5464A51">Aidek, Ahmad E.</mods:namePart>
<mods:affiliation id="A22049216CCB79403C059085F907B18C">General Commission for Scientific Agricultural Research, Deir ez-Zor, Syria</mods:affiliation>
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<mods:namePart id="D2A15EC8F6532D1D31071F2626D6F873">Saad, Adib</mods:namePart>
<mods:affiliation id="041F4F07C07D9A88DFCF0D4BBA79F2BC">Al-Manara University, Latakia, Syria</mods:affiliation>
</mods:name>
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<mods:affiliation id="5FC6344C5E0D7E2579D9F7978AC6BD5B">Department of Zoology, Comenius University in Bratislava, Ilkovičova 6, Mlynská dolina, 842 15 Bratislava, Slovakia</mods:affiliation>
</mods:name>
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<mods:namePart id="456DD53642A5CFF55827C99229288B3C">Esterbauer, Hans</mods:namePart>
<mods:affiliation id="E075EA34BF6B4444047D155F2E183FE8">Johann-Puch-Strasse 27 / III / 5, 4400 Steyr, Austria</mods:affiliation>
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</mods:role>
<mods:namePart id="2DD2616A4A84E9F3BC9F18F3EE594F20">Fritz, Uwe</mods:namePart>
<mods:affiliation id="79E52895849506A577E88ED8484B50F4">Museum of Zoology, Senckenberg Dresden, A. B. Meyer Building, 01109 Dresden, Germany</mods:affiliation>
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<treatment id="03B9F52AC731FF9304B62F9CF61AF998" LSID="urn:lsid:plazi:treatment:03B9F52AC731FF9304B62F9CF61AF998" httpUri="http://treatment.plazi.org/id/03B9F52AC731FF9304B62F9CF61AF998" lastPageId="23" lastPageNumber="174" pageId="20" pageNumber="171">
<subSubSection id="C30A17B7C731FF9004B62F9CF6E0F883" pageId="20" pageNumber="171" type="nomenclature">
<paragraph id="8BAF443CC731FF9004B62F9CF679F8A4" blockId="20.[151,744,1782,1846]" box="[151,625,1782,1810]" pageId="20" pageNumber="171">
<heading id="D0E7F350C731FF9004B62F9CF679F8A4" bold="true" box="[151,625,1782,1810]" fontSize="11" level="1" pageId="20" pageNumber="171" reason="1">
<taxonomicName id="4C103FBFC731FF9004B62F9CF679F8A4" ID-CoL="5F7F2" authority="Forskal, 1775" authorityName="Forskal" authorityYear="1775" box="[151,625,1782,1810]" class="Testudines" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Cryptodira" pageId="20" pageNumber="171" phylum="Chordata" rank="subSpecies" species="graeca" subSpecies="terrestris">
<emphasis id="B964982EC731FF9004B62F9CF679F8A4" bold="true" box="[151,625,1782,1810]" pageId="20" pageNumber="171">
<emphasis id="B964982EC731FF9004B62F9CF5C9F8A7" bold="true" box="[151,449,1783,1810]" italics="true" pageId="20" pageNumber="171">Testudo graeca terrestris</emphasis>
<bibRefCitation id="EF8139CDC731FF9005E92F9DF679F8A4" author="Forskal, P." box="[456,625,1782,1809]" pageId="20" pageNumber="171" refId="ref14464" refString="Forskal, P. (1775) Descriptiones Animalium: Avium, Amphibiorum, Piscium, Insectorum, Vermium; quae in Itinere Orientali Observavit. Post mortem auctoris edidit Carsten Niebuhr. Moller, Copenhagen, 164 pp. https: // doi. org / 10.5962 / bhl. title. 2154" type="book" year="1775">Forskål, 1775</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BAF443CC731FF9004B62E77F6E0F883" blockId="20.[151,744,1782,1846]" box="[151,744,1820,1846]" pageId="20" pageNumber="171">
Mesopotamian Tortoise, Levantine Tortoise (
<figureCitation id="132B58B9C731FF9006AE2E77F6E8F883" box="[655,736,1820,1846]" captionStart="FIGURE 20" captionStartId="21.[151,250,1732,1756]" captionTargetBox="[151,1435,849,1709]" captionTargetId="figure-266@21.[151,1435,849,1709]" captionTargetPageId="21" captionText="FIGURE 20. Coloration of Testudo graeca terrestris. (A) Al-Hardana, north of Mesopotamia (photo: Mohammed Al-Karraf); (B) Maqta Hajar, northeast Aleppo (photo: Ahmad Aidek); (C) Majdalun Al-Bustan, coastal region (photo: Ali Ali); (D) Al- Bassah, coastal region (photo: Ali Alloush); (E) Sadd Al-Kafat, southeast Hamah (photo: Ehab Jbr); (F) as-Sabbourah, west of Damascus (photo: Wael Almatni); (G) Masil Al-Fawwar, southwestern Syria (photo: Hans Esterbauer); (H) hatchling, Shahba, Jabal Al-Arab (photo: Ahmad Aidek); (I) hatchling, Tell Sheihan, Jabal Al-Arab (photo: Wael Almatni)." pageId="20" pageNumber="171">Fig. 20</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30A17B7C731FF9004B62E36F796F8C2" box="[151,926,1884,1912]" pageId="20" pageNumber="171" type="description">
<paragraph id="8BAF443CC731FF9004B62E36F796F8C2" blockId="20.[151,1437,1884,2020]" box="[151,926,1884,1912]" pageId="20" pageNumber="171">
<emphasis id="B964982EC731FF9004B62E36F680F8C2" bold="true" box="[151,648,1884,1911]" pageId="20" pageNumber="171">
IUCN Red List Category (
<taxonomicName id="4C103FBFC731FF9005F02E37F688F8C2" box="[465,640,1884,1911]" class="Testudines" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Cryptodira" pageId="20" pageNumber="171" phylum="Chordata" rank="species" species="graeca">
<emphasis id="B964982EC731FF9005F02E37F688F8C2" bold="true" box="[465,640,1884,1911]" italics="true" pageId="20" pageNumber="171">Testudo graeca</emphasis>
</taxonomicName>
)
</emphasis>
: Vulnerable (VU A1cd).
</paragraph>
</subSubSection>
<subSubSection id="C30A17B7C731FF9304E62EEAF736FE18" lastPageId="23" lastPageNumber="174" pageId="20" pageNumber="171" type="materials_examined">
<paragraph id="8BAF443CC731FF9104E62EEAF4E8FF60" blockId="20.[151,1437,1884,2020]" lastBlockId="21.[151,1437,151,825]" lastPageId="21" lastPageNumber="172" pageId="20" pageNumber="171">
<emphasis id="B964982EC731FF9004E62EEAF569F82E" bold="true" box="[199,353,1921,1947]" pageId="20" pageNumber="171">Type locality</emphasis>
: “Lohajae ..&amp;.. Kahirae..&amp;.. Aleppo..&amp;..
<collectingCountry id="F30704ACC731FF90071B2EEAF792F82E" box="[826,922,1921,1947]" name="Lebanon" pageId="20" pageNumber="171">Libanon</collectingCountry>
” [Al Luhayyah ..&amp;.. Cario ..&amp;.. Aleppo ..&amp;.. Mount
<collectingCountry id="F30704ACC731FF9004CF2ECEF55FF80A" box="[238,343,1957,1983]" name="Lebanon" pageId="20" pageNumber="171">Lebanon</collectingCountry>
]. Restricted to Arabien [Arabia] by
<bibRefCitation id="EF8139CDC731FF9007242ECEF7CEF875" author="Wermuth, H." box="[773,966,1957,1984]" pageId="20" pageNumber="171" pagination="399 - 423" refId="ref21446" refString="Wermuth, H. (1956) Versuch der Deutung einiger bisher ubersehener Schildkroten-Namen. Zoologische Beitrage, Neue Folge, 2, 399 - 423." type="journal article" year="1956">Wermuth (1956)</bibRefCitation>
, to “Libanon-Gebirge,
<collectingCountry id="F30704ACC731FF9000FB2ECEF113F80A" box="[1242,1307,1957,1983]" name="Israel" pageId="20" pageNumber="171">Israel</collectingCountry>
” [
<collectingCountry id="F30704ACC731FF9001162ECEF194F80A" box="[1335,1436,1957,1983]" name="Lebanon" pageId="20" pageNumber="171">Lebanon</collectingCountry>
Mountains,
<collectingCountry id="F30704ACC731FF90053D2EA2F555F856" box="[284,349,1993,2019]" name="Israel" pageId="20" pageNumber="171">Israel</collectingCountry>
] by
<bibRefCitation id="EF8139CDC731FF9005AA2EA2F64DF851" author="Wermuth, H." box="[395,581,1993,2020]" pageId="20" pageNumber="171" pagination="149 - 153" refId="ref21473" refString="Wermuth, H. (1958) Status und Nomenklatur der Maurischen Landschildkrote, Testudo graeca, in SW-Asien und NO-Afrika. Senckenbergiana biologica, 39, 149 - 153." type="journal article" year="1958">Wermuth (1958)</bibRefCitation>
, to “environs dAlep (=
<collectingRegion id="49D48ADEC731FF9007732EA2F791F856" box="[850,921,1993,2019]" country="Syria" name="Halab" pageId="20" pageNumber="171">Halab</collectingRegion>
),
<collectingCountry id="F30704ACC731FF90078A2EA2F7E0F856" box="[939,1000,1993,2019]" name="Syria" pageId="20" pageNumber="171">Syrie</collectingCountry>
” [
<collectingCountry id="F30704ACC731FF9007DE2EA2F037F856" box="[1023,1087,1993,2019]" name="Syria" pageId="20" pageNumber="171">Syria</collectingCountry>
] by
<bibRefCitation id="EF8139CDC731FF90004F2EA1F0F1F856" author="Bour, R." box="[1134,1273,1993,2020]" pageId="20" pageNumber="171" pagination="13 - 19" refId="ref13799" refString="Bour, R. (1989) Caracteres diagnostiques offerts par le crane des tortues terrestres du genre Testudo. Mesogee, 48, 13 - 19." type="journal article" year="1989">Bour (1989)</bibRefCitation>
, and validly to “
<collectingRegion id="49D48ADEC730FF91048029FCF4FEFF04" box="[161,246,151,177]" country="Syria" name="Halab" pageId="21" pageNumber="172">Aleppo</collectingRegion>
(
<collectingRegion id="49D48ADEC730FF91052729FCF548FF04" box="[262,320,151,177]" country="Syria" name="Halab" pageId="21" pageNumber="172">Alep</collectingRegion>
,
<collectingRegion id="49D48ADEC730FF91056D29FCF59AFF04" box="[332,402,151,177]" country="Syria" name="Halab" pageId="21" pageNumber="172">Halab</collectingRegion>
;
<geoCoordinate id="EE2422FBC730FF9105BE29FCF60AFF04" box="[415,514,151,177]" degrees="36" direction="north" minutes="12" orientation="latitude" pageId="21" pageNumber="172" precision="925" value="36.2">36°12N</geoCoordinate>
,
<geoCoordinate id="EE2422FBC730FF91062F29FCF667FF04" box="[526,623,151,177]" degrees="37" direction="east" minutes="09" orientation="longitude" pageId="21" pageNumber="172" precision="925" value="37.15">37°09E</geoCoordinate>
),
<collectingCountry id="F30704ACC730FF9106A529FCF6B7FF04" box="[644,703,151,177]" name="Syria" pageId="21" pageNumber="172">Syria</collectingCountry>
(
<collectingCountry id="F30704ACC730FF9106EE29FCF7C1FF04" box="[719,969,151,177]" name="Syria" pageId="21" pageNumber="172">Syrian Arab Republic</collectingCountry>
)” by
<typeStatus id="54ABFA9EC730FF91002729FCF057FF04" box="[1030,1119,151,177]" pageId="21" pageNumber="172" type="neotype">neotype</typeStatus>
designation (
<bibRefCitation id="EF8139CDC730FF9100D629FCF4DCFF60" author="Perala, J. &amp; Bour, R." pageId="21" pageNumber="172" pagination="114 - 119" refId="ref16958" refString="Perala, J. &amp; Bour, R. (2004) Neotype of Testudo terrestris Forsskal, 1775 (Testudines, Testudinidae). Asiatic Herpetological Research, 10, 114 - 119." type="journal article" year="2004">Perälä &amp; Bour 2004</bibRefCitation>
).
</paragraph>
<paragraph id="8BAF443CC730FF9104E629B4F5F2FEA8" blockId="21.[151,1437,151,825]" pageId="21" pageNumber="172">
<emphasis id="B964982EC730FF9104E629B4F575FF4D" bold="true" box="[199,381,222,249]" pageId="21" pageNumber="172">Type specimen</emphasis>
:
<materialsCitation id="3B784E61C730FF9105B329B4F5ECFEA8" collectionCode="NMW" location="Naturhistorisches Museum" pageId="21" pageNumber="172" specimenCode="NMW 18674" specimenCount="2" stateProvince="Wien" typeStatus="syntype">
Original
<typeStatus id="54ABFA9EC730FF9105DC29B4F657FF4C" box="[509,607,223,249]" pageId="21" pageNumber="172" type="syntype">syntypes</typeStatus>
lost;
<typeStatus id="54ABFA9EC730FF91068829B4F70AFF4C" box="[681,770,223,249]" pageId="21" pageNumber="172" type="neotype">neotype</typeStatus>
designated by
<bibRefCitation id="EF8139CDC730FF91079C29B4F0CFFF4C" author="Perala, J. &amp; Bour, R." box="[957,1223,223,249]" pageId="21" pageNumber="172" pagination="114 - 119" refId="ref16958" refString="Perala, J. &amp; Bour, R. (2004) Neotype of Testudo terrestris Forsskal, 1775 (Testudines, Testudinidae). Asiatic Herpetological Research, 10, 114 - 119." type="journal article" year="2004">Perälä &amp; Bour (2004)</bibRefCitation>
;
<location id="8ECF12E7C730FF9100F929B4F4F1FEA8" LSID="urn:lsid:plazi:treatment:03B9F52AC731FF9304B62F9CF61AF998:8ECF12E7C730FF9100F929B4F4F1FEA8" name="Naturhistorisches Museum" pageId="21" pageNumber="172" stateProvince="Wien">Naturhistorisches Museum</location>
<collectingRegion id="49D48ADEC730FF9105212868F536FEA8" box="[256,318,259,285]" country="Austria" name="Wien" pageId="21" pageNumber="172">Wien</collectingRegion>
,
<specimenCode id="DBB6EC47C730FF9105682868F5ECFEA8" box="[329,484,259,285]" collectionCode="NMW" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="21" pageNumber="172">NMW 18674</specimenCode>
</materialsCitation>
:2.
</paragraph>
<paragraph id="8BAF443CC730FF9104E6284DF651FEF4" blockId="21.[151,1437,151,825]" box="[199,601,294,321]" pageId="21" pageNumber="172">
<emphasis id="B964982EC730FF9104E6284DF54EFEF5" bold="true" box="[199,326,294,320]" pageId="21" pageNumber="172">Chorotype</emphasis>
: Turano-Mediterranean.
</paragraph>
<paragraph id="8BAF443CC730FF9104E62821F740FED0" blockId="21.[151,1437,151,825]" box="[199,840,330,357]" pageId="21" pageNumber="172">
<emphasis id="B964982EC730FF9104E62821F583FED1" bold="true" box="[199,395,330,357]" pageId="21" pageNumber="172">First records for</emphasis>
<materialsCitation id="3B784E61C730FF9105B22821F74CFED0" box="[403,836,330,357]" country="Syria" location="Latakia" pageId="21" pageNumber="172" specimenCount="1" stateProvince="Al Ladhiqiyah">
<emphasis id="B964982EC730FF9105B22821F5DBFED1" bold="true" box="[403,467,330,356]" pageId="21" pageNumber="172">
<collectingCountry id="F30704ACC730FF9105B22821F5DBFED1" box="[403,467,330,356]" name="Syria" pageId="21" pageNumber="172">Syria</collectingCountry>
</emphasis>
:
<collectingRegion id="49D48ADEC730FF9105C02820F630FED0" box="[481,568,331,357]" country="Syria" name="Al Ladhiqiyah" pageId="21" pageNumber="172">Latakia</collectingRegion>
,
<collectingRegion id="49D48ADEC730FF9106632820F69CFED0" box="[578,660,331,357]" country="Syria" name="Halab" pageId="21" pageNumber="172">Aleppo</collectingRegion>
(
<bibRefCitation id="EF8139CDC730FF9106832821F733FED0" author="Forskal, P." box="[674,827,330,357]" pageId="21" pageNumber="172" refId="ref14464" refString="Forskal, P. (1775) Descriptiones Animalium: Avium, Amphibiorum, Piscium, Insectorum, Vermium; quae in Itinere Orientali Observavit. Post mortem auctoris edidit Carsten Niebuhr. Moller, Copenhagen, 164 pp. https: // doi. org / 10.5962 / bhl. title. 2154" type="book" year="1775">Forskål 1775</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BAF443CC730FF9104E62805F06BFC8C" blockId="21.[151,1437,151,825]" pageId="21" pageNumber="172">
<emphasis id="B964982EC730FF9104E62805F58BFE3C" bold="true" box="[199,387,366,393]" pageId="21" pageNumber="172">Literature data</emphasis>
:
<collectingRegion id="49D48ADEC730FF9105B52804F5EEFE3C" box="[404,486,367,393]" country="Syria" name="Halab" pageId="21" pageNumber="172">Aleppo</collectingRegion>
(
<bibRefCitation id="EF8139CDC730FF9105D82804F6AAFE3C" author="Boettger, O." box="[505,674,367,393]" pageId="21" pageNumber="172" pagination="132 - 219" refId="ref13583" refString="Boettger, O. (1880) Die Reptilien und Amphibien von Syrien, Palaestina und Cypern. Berichte der Senckenbergischen Naturforschenden Gesellschaft, 1879 - 1880, 132 - 219." type="book chapter" year="1880">Boettger 1880</bibRefCitation>
),
<collectingRegion id="49D48ADEC730FF9106962804F704FE3C" box="[695,780,367,393]" country="Syria" name="Halab" pageId="21" pageNumber="172">Aleppo</collectingRegion>
,
<collectingRegion id="49D48ADEC730FF91073D2804F76AFE3C" box="[796,866,367,393]" country="Syria" name="Hims" pageId="21" pageNumber="172">Homs</collectingRegion>
,
<collectingRegion id="49D48ADEC730FF9107512804F7CCFE3C" box="[880,964,367,393]" country="Syria" name="Hamah" pageId="21" pageNumber="172">Hamah</collectingRegion>
, Palmyra, near
<collectingRegion id="49D48ADEC730FF91005C2804F0F8FE3C" box="[1149,1264,367,393]" country="Syria" name="Dimashq" pageId="21" pageNumber="172">Damascus</collectingRegion>
(
<bibRefCitation id="EF8139CDC730FF9101222804F187FE3C" author="Lortet, L." box="[1283,1423,367,393]" pageId="21" pageNumber="172" pagination="99 - 194" refId="ref16560" refString="Lortet, L. (1883) Poissons et reptiles du Lac de Tiberiade et de quelques autres parties de la Syrie. Archives du Museum d'histoire naturelle de Lyon, 3, 99 - 194. https: // doi. org / 10.3406 / mhnly. 1883.941" type="journal article" year="1883">Lortet 1883</bibRefCitation>
),
<collectingRegion id="49D48ADEC730FF9104B628F8F4E1FE18" box="[151,233,403,429]" country="Syria" name="Halab" pageId="21" pageNumber="172">Aleppo</collectingRegion>
(
<bibRefCitation id="EF8139CDC730FF9104D628F8F5B0FE18" author="Siebenrock, F." box="[247,440,403,429]" pageId="21" pageNumber="172" pagination="171 - 225" refId="ref19218" refString="Siebenrock, F. (1913) Wissenschaftliche Ergebnisse der Expedition nach Mesopotamien, 1910. Schildkroten aus Syrien und Mesopotamien. Annalen des Naturhistorischen Museums Wien, 27, 171 - 225." type="journal article" year="1913">Siebenrock 1913</bibRefCitation>
),
<collectingRegion id="49D48ADEC730FF9105EA28F8F606FE18" box="[459,526,403,429]" country="Syria" name="Hims" pageId="21" pageNumber="172">Homs</collectingRegion>
Lake (
<bibRefCitation id="EF8139CDC730FF91067828F8F707FE18" author="Boulenger, G. A." box="[601,783,403,429]" pageId="21" pageNumber="172" pagination="41 - 55" refId="ref13716" refString="Boulenger, G. A. (1923) Etude sur les Batraciens et les Reptiles rapportes par M. Henri Gadeau de Kerville de son voyage zoologique en Syrie (avril-juin 1908). In: Pellegrin, J. (Ed.), Voyage zoologique d'Henri Gadeau de Kerville en Syrie (avril-juin 1908). Tome Quatrieme et Dernier. J. - B. Bailliere, Paris, pp. 41 - 55, pl. vi." type="book chapter" year="1923">Boulenger 1923</bibRefCitation>
)
<collectingRegion id="49D48ADEC730FF91073A28F8F778FE18" box="[795,880,403,429]" country="Syria" name="Halab" pageId="21" pageNumber="172">Aleppo</collectingRegion>
,
<collectingRegion id="49D48ADEC730FF91075B28F8F7C6FE18" box="[890,974,403,429]" country="Syria" name="Tartus" pageId="21" pageNumber="172">Tartous</collectingRegion>
(
<bibRefCitation id="EF8139CDC730FF9107FA28F8F078FE18" author="Werner, F." box="[987,1136,403,429]" pageId="21" pageNumber="172" pagination="130 - 141" refId="ref21504" refString="Werner, F. (1919) Ergebnisse einer mit Unterstutzung der Kais. Akademie der Wissenschaften in Wien ausgefuhrten Forschungsreise von weiland Prof. Dr. Franz Tolg. I. Reptilien und Amphibien aus dem Amanus-Gebirge. Archiv fur Naturgeschichte Berlin, 85 A / 8, 130 - 141." type="journal article" year="1919">Werner 1919</bibRefCitation>
,
<bibRefCitation id="EF8139CDC730FF91005828F9F0BCFE18" author="Werner, F." box="[1145,1204,402,429]" pageId="21" pageNumber="172" pagination="225 - 227" refId="ref21557" refString="Werner, F. (1935) Im Auto durch Syrien. Blatter fur Aquarien- und Terrarienkunde, 46, 225 - 227." type="journal article" year="1935">1935</bibRefCitation>
), between
<collectingRegion id="49D48ADEC730FF91010A28F8F166FE18" box="[1323,1390,403,429]" country="Syria" name="Hims" pageId="21" pageNumber="172">Homs</collectingRegion>
and T-4 (Al-Furuklus;
<bibRefCitation id="EF8139CDC730FF91054428DCF60DFE64" author="Schmidt, K. P." box="[357,517,439,465]" pageId="21" pageNumber="172" pagination="49 - 92" refId="ref19098" refString="Schmidt, K. P. (1939) Reptiles and amphibians from southwestern Asia. Zoological Series, Field Museum of Natural History, 24, 49 - 92. https: // doi. org / 10.5962 / bhl. title. 3841" type="journal article" year="1939">Schmidt 1939</bibRefCitation>
), Golan (
<bibRefCitation id="EF8139CDC730FF91064A28DCF76FFE64" author="Werner, Y. L. &amp; Avital, E." box="[619,871,439,465]" pageId="21" pageNumber="172" pagination="192 - 193" refId="ref21612" refString="Werner, Y. L. &amp; Avital, E. (1980) The herpetofauna of Mt. Hermon and its altitudinal distribution. Israel Journal of Zoology, 29, 192 - 193." type="journal article" year="1980">Werner &amp; Avital 1980</bibRefCitation>
;
<bibRefCitation id="EF8139CDC730FF91075328DCF076FE64" author="Berger-Dell'mour, V. H." box="[882,1150,439,465]" pageId="21" pageNumber="172" pagination="59 - 67" refId="ref13415" refString="Berger-Dell'mour, V. H. (1986) Zur Herpetofauna des Golan. Annalen des Naturhistorischen Museums in Wien, 87 B, 59 - 67." type="journal article" year="1986">Berger-Dellmour 1986</bibRefCitation>
),
<collectingRegion id="49D48ADEC730FF9100B128DCF0EDFE64" box="[1168,1253,439,465]" country="Syria" name="Halab" pageId="21" pageNumber="172">Aleppo</collectingRegion>
,
<collectingRegion id="49D48ADEC730FF9100D128DCF14FFE64" box="[1264,1351,439,465]" country="Syria" name="Al Ladhiqiyah" pageId="21" pageNumber="172">Latakia</collectingRegion>
,
<collectingRegion id="49D48ADEC730FF91017328DCF190FE64" box="[1362,1432,439,465]" country="Syria" name="Hims" pageId="21" pageNumber="172">Homs</collectingRegion>
,
<collectingRegion id="49D48ADEC730FF9104B628B0F4E3FE40" box="[151,235,475,501]" country="Syria" name="Hamah" pageId="21" pageNumber="172">Hamah</collectingRegion>
,
<collectingRegion id="49D48ADEC730FF9104D628B0F565FE40" box="[247,365,475,501]" country="Syria" name="Dimashq" pageId="21" pageNumber="172">Damascus</collectingRegion>
, Al-Qunaitra, Al-Hamidiyah (N Al-Qunaitra), Khan Arnabah (= Khan Uraynibah), Al-Kab, N and NW Taranjah, NW and W Hadar, S Arnah, W Majdal Shams (= Megdel Shams;
<bibRefCitation id="EF8139CDC730FF9100592894F149FDAC" author="Esterbauer, H." box="[1144,1345,510,537]" pageId="21" pageNumber="172" pagination="23 - 34" refId="ref14303" refString="Esterbauer, H. (1985 a) Zur Herpetofauna Sudwestsyriens. Herpetofauna, 7 (38), 23 - 34." type="journal article" year="1985">Esterbauer 1985a</bibRefCitation>
,
<bibRefCitation id="EF8139CDC730FF91016E2895F190FDAC" author="Esterbauer, H." box="[1359,1432,510,537]" pageId="21" pageNumber="172" pagination="389 - 392" refId="ref14326" refString="Esterbauer, H. (1985 b) Okologische und verhaltensbiologische Beobachtungen an der Maurischen Landschildkrote, Testudo graeca terrestris Forskal, 1775 in Sudwestsyrien. Aquarien-Terrarien, 32, 389 - 392." type="journal article" year="1985">1985b</bibRefCitation>
,
<bibRefCitation id="EF8139CDC730FF9104B62B48F4DBFD88" author="Esterbauer, H." box="[151,211,547,573]" pageId="21" pageNumber="172" pagination="21 - 54" refId="ref14360" refString="Esterbauer, H. (1992) Die Herpetofauna des ostlichen Golan und Hermongebietes. Funde und Bemerkungen zur Systematik und Okologie. Zoology in the Middle East, 7, 21 - 54. https: // doi. org / 10.1080 / 09397140.1992.10637623" type="journal article" year="1992">1992</bibRefCitation>
), SE Jabal Al-Arab (Dzabal ad Druz;
<bibRefCitation id="EF8139CDC730FF9106512B48F6F3FD88" author="Vesely, M." box="[624,763,547,573]" pageId="21" pageNumber="172" pagination="24 - 25" refId="ref21002" refString="Vesely, M. (1991) O herpetofaune Syrie. Akvarium-Terarium, 34, 24 - 25." type="journal article" year="1991">Veselý 1991</bibRefCitation>
), Golan and Mount Hermon (
<bibRefCitation id="EF8139CDC730FF9100612B48F13DFD88" author="Sivan, N. &amp; Werner, Y. L." box="[1088,1333,547,573]" pageId="21" pageNumber="172" pagination="193 - 211" refId="ref19439" refString="Sivan, N. &amp; Werner, Y. L. (1992) Survey of the reptiles of the Golan plateau and Mt. Hermon. Israel Journal of Ecology and Evolution, 37, 193 - 211. https: // doi. org / 10.1080 / 00212210.1992.10688655" type="journal article" year="1992">Sivan &amp; Werner 1992</bibRefCitation>
), Mesiaf, Slenfeh, Al-Barah (
<bibRefCitation id="EF8139CDC730FF9105502B2CF6B0FDD4" author="Bischoff, W. &amp; Schmidtler, J. F." box="[369,696,583,609]" pageId="21" pageNumber="172" pagination="4 - 22" refId="ref13554" refString="Bischoff, W. &amp; Schmidtler, J. F. (1994) Ergebnisse zweier Lacertiden-Exkursionen nach Syrien. Die Eidechse, 5, 4 - 22." type="journal article" year="1994">Bischoff &amp; Schmidtler 1994</bibRefCitation>
), Seidnaya, Mesiaf, Jabal Al-Arab (= Jebel ed Drouz),
<collectingRegion id="49D48ADEC730FF9101042B2CF195FDD4" box="[1317,1437,583,609]" country="Syria" name="Al Hasakah" pageId="21" pageNumber="172">Al-Hasaka</collectingRegion>
(
<bibRefCitation id="EF8139CDC730FF9104BE2B00F55EFD30" author="Fritz, U. &amp; Bischoff, W. &amp; Martens, H. &amp; Schmidtler, J. F." box="[159,342,619,645]" pageId="21" pageNumber="172" pagination="5 - 14" refId="ref14783" refString="Fritz, U., Bischoff, W., Martens, H. &amp; Schmidtler, J. F. (1996) Variabilitat syrischer Landschildkroten (Testudo graeca) sowie zur Systematik und Zoogeographie im Nahen Osten und in Nordafrika. Herpetofauna, 18 (104), 5 - 14." type="journal article" year="1996">
Fritz
<emphasis id="B964982EC730FF9104FD2B00F51CFD30" box="[220,276,619,645]" italics="true" pageId="21" pageNumber="172">et al.</emphasis>
1996
</bibRefCitation>
), Jabal Al-Arab (= Djebel Druz;
<bibRefCitation id="EF8139CDC730FF9106E22B00F7ACFD30" author="Bischoff, W. &amp; in den Bosch, H. A. J. &amp; Schmidtler, J. F." box="[707,932,619,645]" pageId="21" pageNumber="172" pagination="1 - 9" refId="ref13506" refString="Bischoff, W., in den Bosch, H. A. J. &amp; Schmidtler, J. F. (1998) Mit dem Finger auf der Landkarte entdeckt. Eidechsen auf dem Djebel Druz. Die Eidechse, 9, 1 - 9." type="journal article" year="1998">
Bischoff
<emphasis id="B964982EC730FF91070B2B00F76AFD30" box="[810,866,619,645]" italics="true" pageId="21" pageNumber="172">et al.</emphasis>
1998
</bibRefCitation>
),
<quantity id="4CE8E9D9C730FF9107962B01F7F8FD30" box="[951,1008,618,645]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="21" pageNumber="172" unit="km" value="5.0">5 km</quantity>
north of Sweida (
<bibRefCitation id="EF8139CDC730FF9100972B00F4DBFD1C" author="Moravec, J. &amp; Orszagh, I." pageId="21" pageNumber="172" pagination="181 - 182" refId="ref16858" refString="Moravec, J. &amp; Orszagh, I. (1998) Testudo graeca terrestris als Wirt von Leptoconops irritans (Diptera: Ceratopogonidae). Salamandra, 34, 181 - 182." type="journal article" year="1998">Moravec &amp; Orszagh 1998</bibRefCitation>
),
<collectingRegion id="49D48ADEC730FF9104C82BE4F53DFD1C" box="[233,309,655,681]" country="Saudi Arabia" name="Najran" pageId="21" pageNumber="172">Najran</collectingRegion>
(
<bibRefCitation id="EF8139CDC730FF9105672BE4F6E7FD1C" author="Lymberakis, R. &amp; Kalionzopoulou, A." box="[326,751,655,681]" pageId="21" pageNumber="172" pagination="33 - 39" refId="ref16609" refString="Lymberakis, R. &amp; Kalionzopoulou, A. (2003) Additions to the herpetofauna of Syria. Zoology in the Middle East, 29, 33 - 39. https: // doi. org / 10.1080 / 09397140.2003.10637967" type="journal article" year="2003">Lymberakis &amp; Kalionzopoulou 2003</bibRefCitation>
), Al-Badia near Palmyra, Fir-Cedar Reserve (
<bibRefCitation id="EF8139CDC730FF9100DD2BE4F4DBFD78" author="Sindaco, R. &amp; Serra, G. &amp; Menegon, M." pageId="21" pageNumber="172" pagination="29 - 38" refId="ref19307" refString="Sindaco, R., Serra, G. &amp; Menegon, M. (2006) New data on the Syrian herpetofauna, with a newly recorded species of snake. Zoology in the Middle East, 37, 29 - 38. https: // doi. org / 10.1080 / 09397140.2006.10638146" type="journal article" year="2006">
Sindaco
<emphasis id="B964982EC730FF9101432BE4F195FD1C" box="[1378,1437,655,681]" italics="true" pageId="21" pageNumber="172">et al.</emphasis>
2006
</bibRefCitation>
), Palmyra (
<bibRefCitation id="EF8139CDC730FF9105772BD8F61CFD78" author="Fritz, U. &amp; Bischoff, W. &amp; Martens, H. &amp; Schmidtler, J. F." box="[342,532,691,717]" pageId="21" pageNumber="172" pagination="5 - 14" refId="ref14783" refString="Fritz, U., Bischoff, W., Martens, H. &amp; Schmidtler, J. F. (1996) Variabilitat syrischer Landschildkroten (Testudo graeca) sowie zur Systematik und Zoogeographie im Nahen Osten und in Nordafrika. Herpetofauna, 18 (104), 5 - 14." type="journal article" year="1996">
Fritz
<emphasis id="B964982EC730FF9105B42BD8F5C0FD78" box="[405,456,691,717]" italics="true" pageId="21" pageNumber="172">et al</emphasis>
. 1996
</bibRefCitation>
), Tahonat Al-Halawa, Slenfeh (
<bibRefCitation id="EF8139CDC730FF91075B2BD8F03EFD78" author="Amr, Z. &amp; Shehab, A. &amp; Abu Baker, M." box="[890,1078,691,717]" pageId="21" pageNumber="172" pagination="21 - 26" refId="ref13174" refString="Amr, Z., Shehab, A. &amp; Abu Baker, M. (2007) Recent observations on the herpetofauna of Syria with notes on trade in reptiles. Herpetozoa, 20, 21 - 26." type="journal article" year="2007">
Amr
<emphasis id="B964982EC730FF9107992BD8F7FAFD78" box="[952,1010,691,717]" italics="true" pageId="21" pageNumber="172">et al.</emphasis>
2007
</bibRefCitation>
), Sweida, Qanawat, Rashiedeh, Seidnaya, Maalula, Mesiaf, Ain Al-Beidah, Jourin, Qalat Samaan (
<bibRefCitation id="EF8139CDC730FF9107A82BB3F050FD44" author="Siroky, P. &amp; Jandzik, D. &amp; Mikulicek, P. &amp; Moravec, J. &amp; Orszagh, I." box="[905,1112,727,755]" pageId="21" pageNumber="172" pagination="485 - 489" refId="ref19361" refString="Siroky, P., Jandzik, D., Mikulicek, P., Moravec, J. &amp; Orszagh, I. (2007) Leptoconops bezzii (Diptera: Ceratopogonidae) parasitizing tortoises Testudo graeca (Testudines: Testudinidae) in mountain ranges of Lebanon and western Syria. Parasitology Research, 101, 485 - 489. https: // doi. org / 10.1007 / s 00436 - 007 - 0490 - 2" type="journal article" year="2007">
Široký
<emphasis id="B964982EC730FF9107FC2BBCF01DFD44" box="[989,1045,727,753]" italics="true" pageId="21" pageNumber="172">et al.</emphasis>
2007
</bibRefCitation>
), Maalula, Seidnaya, Ain Al- Beida, Qalat Merza, Mesiaf, Al-Kafr, Sweida, between Sweida and Salah, Dar Tazah, Qalat Samaan (
<bibRefCitation id="EF8139CDC730FF9101072B90F4DBFC8C" author="Fritz, U. &amp; Hundsdorfer, A. K. &amp; Siroky, P. &amp; Auer, M. &amp; Kami, H. &amp; Lehmann, J. &amp; Mazanaeva, L. F. &amp; Turkozan, O. &amp; Wink, M." pageId="21" pageNumber="172" pagination="97 - 121" refId="ref14837" refString="Fritz, U., Hundsdorfer, A. K., Siroky, P., Auer, M., Kami, H., Lehmann, J., Mazanaeva, L. F., Turkozan, O. &amp; Wink, M. (2007) Phenotypic plasticity leads to incongruence between morphology-based taxonomy and genetic differentiation in western Palaearctic tortoises (Testudo graeca complex; Testudines, Testudinidae). Amphibia-Reptilia, 28, 97 - 121. https: // doi. org / 10.1163 / 156853807779799135" type="journal article" year="2007">
Fritz
<emphasis id="B964982EC730FF9101452B90F195FCA0" box="[1380,1437,763,789]" italics="true" pageId="21" pageNumber="172">et al.</emphasis>
2007
</bibRefCitation>
), Al-Qerdaha, Al-Boudi, Dhahr Drekeish, Jabla, Al-Bassah (
<bibRefCitation id="EF8139CDC730FF9107562A74F05EFC8C" author="Mhanna, H. &amp; AlMasri, A. &amp; Ibrahim, N." box="[887,1110,799,825]" pageId="21" pageNumber="172" pagination="40 - 46" refId="ref16725" refString="Mhanna, H., AlMasri, A. &amp; Ibrahim, N. (2021) The morphology and size of erythrocytes of six reptiles species [sic] from the Syrian coastal region. Scientific Reports in Life Sciences, 2, 40 - 46. https: // doi. org / 10.22034 / srls. 2021.247948" type="journal article" year="2021">
Mhanna
<emphasis id="B964982EC730FF9107FA2A74F01BFC8C" box="[987,1043,799,825]" italics="true" pageId="21" pageNumber="172">et al.</emphasis>
2021
</bibRefCitation>
).
</paragraph>
<caption id="DF6F14B4C730FF9104B62FAFF0B2F8D8" pageId="21" pageNumber="172" startId="21.[151,250,1732,1756]" targetBox="[151,1435,849,1709]" targetPageId="21" targetType="figure">
<paragraph id="8BAF443CC730FF9104B62FAFF0B2F8D8" blockId="21.[151,1436,1732,1901]" pageId="21" pageNumber="172">
<emphasis id="B964982EC730FF9104B62FAFF528F969" bold="true" box="[151,288,1732,1756]" pageId="21" pageNumber="172">FIGURE 20.</emphasis>
Coloration of
<taxonomicName id="4C103FBFC730FF9105992FAEF6BEF969" authorityName="Forskal" authorityYear="1775" box="[440,694,1733,1757]" class="Testudines" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Cryptodira" pageId="21" pageNumber="172" phylum="Chordata" rank="subSpecies" species="graeca" subSpecies="terrestris">
<emphasis id="B964982EC730FF9105992FAEF6BEF969" box="[440,694,1733,1757]" italics="true" pageId="21" pageNumber="172">Testudo graeca terrestris</emphasis>
</taxonomicName>
. (
<emphasis id="B964982EC730FF9106ED2FAFF6D6F969" bold="true" box="[716,734,1732,1756]" pageId="21" pageNumber="172">A</emphasis>
) Al-Hardana, north of Mesopotamia (photo: Mohammed Al-Karraf); (
<emphasis id="B964982EC730FF9104812F82F4B9F8B4" bold="true" box="[160,177,1769,1793]" pageId="21" pageNumber="172">B</emphasis>
) Maqta Hajar, northeast Aleppo (photo: Ahmad Aidek); (
<emphasis id="B964982EC730FF91072B2F83F714F8B5" bold="true" box="[778,796,1768,1792]" pageId="21" pageNumber="172">C</emphasis>
) Majdalun Al-Bustan, coastal region (photo: Ali Ali); (
<emphasis id="B964982EC730FF9101782F82F163F8B4" bold="true" box="[1369,1387,1769,1793]" pageId="21" pageNumber="172">D</emphasis>
) AlBassah, coastal region (photo: Ali Alloush); (
<emphasis id="B964982EC730FF9106432E66F67BF890" bold="true" box="[610,627,1805,1829]" pageId="21" pageNumber="172">E</emphasis>
) Sadd Al-Kafat, southeast Hamah (photo: Ehab Jbr); (
<emphasis id="B964982EC730FF9100BC2E66F0A4F890" bold="true" box="[1181,1196,1805,1829]" pageId="21" pageNumber="172">F</emphasis>
) as-Sabbourah, west of Damascus (photo: Wael Almatni); (
<emphasis id="B964982EC730FF9106242E5BF611F8FD" bold="true" box="[517,537,1840,1864]" pageId="21" pageNumber="172">G</emphasis>
) Masil Al-Fawwar, southwestern Syria (photo: Hans Esterbauer); (
<emphasis id="B964982EC730FF91009E2E5AF0DBF8FC" bold="true" box="[1215,1235,1841,1865]" pageId="21" pageNumber="172">H</emphasis>
) hatchling, Shahba, Jabal Al-Arab (photo: Ahmad Aidek); (
<emphasis id="B964982EC730FF9106042E3EF627F8D8" bold="true" box="[549,559,1877,1901]" pageId="21" pageNumber="172">I</emphasis>
) hatchling, Tell Sheihan, Jabal Al-Arab (photo: Wael Almatni).
</paragraph>
</caption>
<caption id="DF6F14B4C733FF9204B62C07F64DFA1D" pageId="22" pageNumber="173" startId="22.[151,250,1388,1412]" targetBox="[161,1429,191,1357]" targetPageId="22" targetType="figure">
<paragraph id="8BAF443CC733FF9204B62C07F64DFA1D" blockId="22.[151,1436,1388,1448]" pageId="22" pageNumber="173">
<emphasis id="B964982EC733FF9204B62C07F529FA31" bold="true" box="[151,289,1388,1412]" pageId="22" pageNumber="173">FIGURE 21.</emphasis>
Records of
<taxonomicName id="4C103FBFC733FF9205832C07F6AAFA31" authorityName="Forskal" authorityYear="1775" box="[418,674,1388,1412]" class="Testudines" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Cryptodira" pageId="22" pageNumber="173" phylum="Chordata" rank="subSpecies" species="graeca" subSpecies="terrestris">
<emphasis id="B964982EC733FF9205832C07F6AAFA31" box="[418,674,1388,1412]" italics="true" pageId="22" pageNumber="173">Testudo graeca terrestris</emphasis>
</taxonomicName>
in Syria. Black circles: previous records, blue circles: recently confirmed previous records, red circles: new records.
</paragraph>
</caption>
<paragraph id="8BAF443CC733FF9204E62C91F572F880" blockId="22.[151,1437,1530,2025]" pageId="22" pageNumber="173">
<emphasis id="B964982EC733FF9204E62C91F5B8F9A1" bold="true" box="[199,432,1530,1556]" pageId="22" pageNumber="173">Museum specimens</emphasis>
:
<collectingCountry id="F30704ACC733FF9205E12C91F5F3F9A1" box="[448,507,1530,1556]" name="Syria" pageId="22" pageNumber="173">Syria</collectingCountry>
(
<collectionCode id="ED01DCF9C733FF92062A2C91F656F9A1" box="[523,606,1530,1556]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="22" pageNumber="173" type="Museum">FMNH</collectionCode>
19715, 1846936,
<collectionCode id="ED01DCF9C733FF92070D2C90F78AF9A0" box="[812,898,1531,1557]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="22" pageNumber="173">MNHN</collectionCode>
1883.640,
<collectionCode id="ED01DCF9C733FF9200202C91F031F9A1" box="[1025,1081,1530,1556]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="22" pageNumber="173">SMF</collectionCode>
55201, 7821, 47365,
<collectionCode id="ED01DCF9C733FF9201132C90F194F9A0" box="[1330,1436,1531,1557]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="22" pageNumber="173" type="Museum">NHMUK</collectionCode>
1976.437, 1976.441), northern
<collectingCountry id="F30704ACC733FF92062D2F75F64FF98D" box="[524,583,1566,1592]" name="Syria" pageId="22" pageNumber="173">Syria</collectingCountry>
(
<collectionCode id="ED01DCF9C733FF92067C2F75F6AEF98D" box="[605,678,1566,1592]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="22" pageNumber="173">NMW</collectionCode>
34350:1),
<collectingRegion id="49D48ADEC733FF92070F2F75F788F98D" box="[814,896,1566,1592]" country="Syria" name="Halab" pageId="22" pageNumber="173">Aleppo</collectingRegion>
(
<collectionCode id="ED01DCF9C733FF9207B62F75F7E6F98D" box="[919,1006,1566,1592]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="22" pageNumber="173">MNHN</collectionCode>
1988.6790, 1988.6965,
<collectionCode id="ED01DCF9C733FF9201372F75F146F98D" box="[1302,1358,1566,1592]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="22" pageNumber="173">SMF</collectionCode>
7830,
<collectionCode id="ED01DCF9C733FF9204B62F28F4D6F9E8" box="[151,222,1603,1629]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="22" pageNumber="173">NMW</collectionCode>
1908:1, 18672:12, 18673:12, 30094:12, 18674:12), Naseeb (
<collectionCode id="ED01DCF9C733FF9207CB2F29F03BF9E9" box="[1002,1075,1602,1628]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="22" pageNumber="173">NMW</collectionCode>
18520:1),
<collectingRegion id="49D48ADEC733FF9200992F28F123F9E8" box="[1208,1323,1603,1629]" country="Syria" name="Dimashq" pageId="22" pageNumber="173">Damascus</collectingRegion>
(
<collectionCode id="ED01DCF9C733FF9201612F29F191F9E9" box="[1344,1433,1602,1628]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="173" type="Museum">RMNH</collectionCode>
. RENA.14045), near
<collectingRegion id="49D48ADEC733FF9205A02F0CF5FCF934" box="[385,500,1639,1665]" country="Syria" name="Dimashq" pageId="22" pageNumber="173">Damascus</collectingRegion>
(
<collectionCode id="ED01DCF9C733FF9206252F0DF64FF935" box="[516,583,1638,1664]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34797" lsid="urn:lsid:biocol.org:col:34797" name="Illinois Natural History Survey" pageId="22" pageNumber="173" type="Other type of research institution/biorepository">INHS</collectionCode>
10289),
<quantity id="4CE8E9D9C733FF92068C2F0DF6FEF935" box="[685,758,1638,1664]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="22" pageNumber="173" unit="km" value="10.0">10 km</quantity>
E
<collectingRegion id="49D48ADEC733FF9207362F0DF793F934" box="[791,923,1638,1665]" country="Syria" name="Dayr az Zawr" pageId="22" pageNumber="173">Deir ez-Zor</collectingRegion>
(
<collectionCode id="ED01DCF9C733FF92078A2F0DF7F3F935" box="[939,1019,1638,1664]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="22" pageNumber="173" type="Museum">ZFMK</collectionCode>
8665686659),
<quantity id="4CE8E9D9C733FF9200942F0DF0E7F935" box="[1205,1263,1638,1664]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="22" pageNumber="173" unit="km" value="1.0">1 km</quantity>
N Atil (
<collectionCode id="ED01DCF9C733FF92016C2F0DF195F935" box="[1357,1437,1638,1664]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="22" pageNumber="173" type="Museum">ZFMK</collectionCode>
64939), ruins of Apamea (
<collectionCode id="ED01DCF9C733FF9205E82FE1F611F911" box="[457,537,1674,1700]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="22" pageNumber="173" type="Museum">ZFMK</collectionCode>
30537),
<collectingRegion id="49D48ADEC733FF9206A22FE0F6CEF910" box="[643,710,1675,1701]" country="Syria" name="Hims" pageId="22" pageNumber="173">Homs</collectingRegion>
(
<collectionCode id="ED01DCF9C733FF9206F92FE1F74CF911" box="[728,836,1674,1700]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="22" pageNumber="173" type="Museum">NHMUK</collectionCode>
1909.4.20.1, 1970.1860),
<collectingRegion id="49D48ADEC733FF9200552FE1F0E4F911" box="[1140,1260,1674,1700]" country="Syria" name="Al Hasakah" pageId="22" pageNumber="173">Al-Hasaka</collectingRegion>
(
<collectionCode id="ED01DCF9C733FF9200DF2FE1F136F911" box="[1278,1342,1674,1700]" country="Germany" httpUri="http://grbio.org/cool/4j3t-w40g" name="Museum of Zoology Senckenberg Dresden" pageId="22" pageNumber="173">MTD</collectionCode>
32111), Jabal Al-Arab (
<collectionCode id="ED01DCF9C733FF9205652FC5F58CF97D" box="[324,388,1710,1736]" country="Germany" httpUri="http://grbio.org/cool/4j3t-w40g" name="Museum of Zoology Senckenberg Dresden" pageId="22" pageNumber="173">MTD</collectionCode>
47365), Camp Fanar, Golan (
<collectionCode id="ED01DCF9C733FF9206F62FC5F728F97D" box="[727,800,1710,1736]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="22" pageNumber="173">NMW</collectionCode>
30859:1), Golan (
<collectionCode id="ED01DCF9C733FF9207D12FC5F031F97D" box="[1008,1081,1710,1736]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="22" pageNumber="173">NMW</collectionCode>
26789:13, 26810:1, 33203:1 21, 33204:1, 34347:1,
<collectionCode id="ED01DCF9C733FF9205BB2FB8F5C5F958" box="[410,461,1747,1773]" pageId="22" pageNumber="173">HUJ</collectionCode>
120719, 120732, 12073620739, 120762), Wadi Daraa (= Wadidra;
<collectionCode id="ED01DCF9C733FF9200C12FB8F12FF958" box="[1248,1319,1747,1773]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="22" pageNumber="173">NMW</collectionCode>
34348:1), Mt. Hermon (
<collectionCode id="ED01DCF9C733FF9205112F9DF571F8A5" box="[304,377,1782,1808]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="22" pageNumber="173">NMW</collectionCode>
34349:1), Slenfeh (
<collectionCode id="ED01DCF9C733FF9206792F9DF6BBF8A5" box="[600,691,1782,1808]" pageId="22" pageNumber="173">JUSTM</collectionCode>
0202), Tahonat Al-Halawa (
<collectionCode id="ED01DCF9C733FF9207D32F9DF045F8A5" box="[1010,1101,1782,1808]" pageId="22" pageNumber="173">JUSTM</collectionCode>
201), E Al-Rashiedeh (
<collectionCode id="ED01DCF9C733FF9201752F9DF194F8A5" box="[1364,1436,1782,1808]" pageId="22" pageNumber="173">NMP-</collectionCode>
P6V 34915, 70504).
</paragraph>
<paragraph id="8BAF443CC733FF9304E62E55F76EFF60" blockId="22.[151,1437,1530,2025]" lastBlockId="23.[151,1437,151,1581]" lastPageId="23" lastPageNumber="174" pageId="22" pageNumber="173">
<emphasis id="B964982EC733FF9204E62E55F5D7F8ED" bold="true" box="[199,479,1854,1880]" pageId="22" pageNumber="173">Citizen science records</emphasis>
: Ad-Difleh, Ain Aaisheh, Ain at-Teineh, Ain ej-Jajeh, Al-Breikhiyeh, Al-Jbeiliyeh, Al-Khaldiyeh, Al-Kneisat, Al-Maliyeh, Al-Marqab, Al-Mzeiraa, Al-Qallouf, Al-Sqeilbiyeh, ash-Shabtaliyah, ash- Shamiyeh, ash-Sheikh Badr, as-Sarsakiyah, as-Sarsakiyah, as-Sukkariyeh, at-Tabrah, at-Teiha, Balluran, Baqu, Bashbatah,Al-Hasriyeh, Beit Al-Jabal, Beit Nafleh, Beit Shouhr, Burj Islam, Burmanah, Fadrah, Ghmam, Habbabah, Haffah, Hammam Al-Qarahleh, Jneinat Raslan, Jubb Al-Amlas, Jubb Ramlah, Kafr Sneif, Kafriyeh ash-Sharqiyeh, Kersana, Kherbat Abu Hamdan, Kherbat aj-Jouziyeh, Kherbat Al-Fars, NabAl-Ghaqah, Nab Al-Sakher, Palmyra, Safita, Salma, Sarai, Squbeen, Um et-Toyour (iNaturalist 2023).
</paragraph>
<paragraph id="8BAF443CC732FF9304E629B4F736FE18" blockId="23.[151,1437,151,1581]" pageId="23" pageNumber="174">
<emphasis id="B964982EC732FF9304E629B4F577FF4C" bold="true" box="[199,383,223,249]" pageId="23" pageNumber="174">Recent records</emphasis>
: Abu Qubays,
<quantity id="4CE8E9D9C732FF93061729B4F67CFF4C" box="[566,628,223,249]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="23" pageNumber="174" unit="km" value="2.0">2 km</quantity>
N Urman, Ain Al-Feijeh, aj-Jumrkiyeh, Al-Amriyeh, Al-Badrosiyah, Al-Bahlouliyah, Al-Bath, Al-Bassah, Al-Dali, Al-Furunlok, Al-Haffah, Al-Hardana, Al-Jabboul, Al-Kadmous, Al-Mazraa, Al-Mismiyeh, Al-Qerdaha, Al-Tamri, Al-Wadia, Azaz, Banias, Bennesh, Binjaru, Drekeish,
<collectingRegion id="49D48ADEC732FF930173284CF190FEF4" box="[1362,1432,295,321]" country="Syria" name="Hims" pageId="23" pageNumber="174">Homs</collectingRegion>
, Jaiboul, Jabla, Jadduh—Salamiyah, Jariz, Kherbat Al-Hamam, Kherbat Hazzour,
<collectingRegion id="49D48ADEC732FF9300682820F0A8FED0" box="[1097,1184,331,357]" country="Syria" name="Al Ladhiqiyah" pageId="23" pageNumber="174">Latakia</collectingRegion>
, Majdalun Al-Bustan, Maqta Hajar, Muhasan, Qiroun, Safita, Sahnaya, Salamiyah, Salhab, Sadd Al-Kafat, Shahba, Souran, Sweida, Tell Sheihan, Tnakha, Tseil,
<collectingRegion id="49D48ADEC732FF93058528F8F5F3FE18" box="[420,507,403,429]" country="Syria" name="Tartus" pageId="23" pageNumber="174">Tartous</collectingRegion>
, W of Yafour, Wadi Barada.
</paragraph>
</subSubSection>
<subSubSection id="C30A17B7C732FF9304E628DDF7F3FB70" pageId="23" pageNumber="174" type="distribution">
<paragraph id="8BAF443CC732FF9304E628DDF733FD88" blockId="23.[151,1437,151,1581]" pageId="23" pageNumber="174">
<emphasis id="B964982EC732FF9304E628DDF5B7FE65" bold="true" box="[199,447,438,464]" pageId="23" pageNumber="174">
Distribution in
<collectingCountry id="F30704ACC732FF93055E28DDF5B7FE65" box="[383,447,438,464]" name="Syria" pageId="23" pageNumber="174">Syria</collectingCountry>
</emphasis>
: Common. The distribution of
<emphasis id="B964982EC732FF93073F28DCF7B4FE65" box="[798,956,439,464]" italics="true" pageId="23" pageNumber="174">
T. g.
<taxonomicName id="4C103FBFC732FF93077728DCF7B4FE65" box="[854,956,439,464]" class="Testudines" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Cryptodira" pageId="23" pageNumber="174" phylum="Chordata" rank="species" species="terrestris">terrestris</taxonomicName>
</emphasis>
extends from east of
<collectingRegion id="49D48ADEC732FF93008C28DCF0F7FE64" box="[1197,1279,439,465]" country="Syria" name="Halab" pageId="23" pageNumber="174">Aleppo</collectingRegion>
westwards. It occurs throughout the coastal region, extending east beyond the Orontes Basin, then south into the
<collectingRegion id="49D48ADEC732FF9300CD28B0F157FE40" box="[1260,1375,475,501]" country="Syria" name="Dimashq" pageId="23" pageNumber="174">Damascus</collectingRegion>
area, the Lebanon Mountains, the southwestern region of Jabal Al-Arab, and the Golan and Mount Hermon, with few records from the northeast near the Turkish border (
<figureCitation id="132B58B9C732FF9306FB2B48F726FD88" box="[730,814,547,573]" captionStart="FIGURE 21" captionStartId="22.[151,250,1388,1412]" captionTargetBox="[161,1429,191,1357]" captionTargetId="figure-21@22.[151,1436,181,1364]" captionTargetPageId="22" captionText="FIGURE 21. Records of Testudo graeca terrestris in Syria. Black circles: previous records, blue circles: recently confirmed previous records, red circles: new records." pageId="23" pageNumber="174">Fig. 21</figureCitation>
).
</paragraph>
<paragraph id="8BAF443CC732FF9304E62B2CF644FC10" blockId="23.[151,1437,151,1581]" pageId="23" pageNumber="174">
<bibRefCitation id="EF8139CDC732FF9304E62B2CF557FDD4" author="Lortet, L." box="[199,351,583,609]" pageId="23" pageNumber="174" pagination="99 - 194" refId="ref16560" refString="Lortet, L. (1883) Poissons et reptiles du Lac de Tiberiade et de quelques autres parties de la Syrie. Archives du Museum d'histoire naturelle de Lyon, 3, 99 - 194. https: // doi. org / 10.3406 / mhnly. 1883.941" type="journal article" year="1883">Lortet (1883)</bibRefCitation>
mentioned that tortoises were common in the region of Palmyra (also mentioned in
<bibRefCitation id="EF8139CDC732FF9301042B2CF4DBFD30" author="Fritz, U. &amp; Bischoff, W. &amp; Martens, H. &amp; Schmidtler, J. F." pageId="23" pageNumber="174" pagination="5 - 14" refId="ref14783" refString="Fritz, U., Bischoff, W., Martens, H. &amp; Schmidtler, J. F. (1996) Variabilitat syrischer Landschildkroten (Testudo graeca) sowie zur Systematik und Zoogeographie im Nahen Osten und in Nordafrika. Herpetofauna, 18 (104), 5 - 14." type="journal article" year="1996">
Fritz
<emphasis id="B964982EC732FF9301422B2CF19DFDD4" box="[1379,1429,583,609]" italics="true" pageId="23" pageNumber="174">et al</emphasis>
. 1996
</bibRefCitation>
). This was the only record for this region for a long time, despite field surveys conducted by many researchers. Also, the senior author of this study did not record any tortoises during dozens of field trips to Palmyra and the Al-Badia between 2006 and 2013. However, it must be remembered that Lortet reported tortoises for this area 150 years ago, and the habitat has certainly changed since then. Thus, it cannot be excluded that
<taxonomicName id="4C103FBFC732FF93006C2BBCF0BDFD45" box="[1101,1205,727,752]" class="Testudines" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Cryptodira" pageId="23" pageNumber="174" phylum="Chordata" rank="species" species="graeca">
<emphasis id="B964982EC732FF93006C2BBCF0BDFD45" box="[1101,1205,727,752]" italics="true" pageId="23" pageNumber="174">T. graeca</emphasis>
</taxonomicName>
once occurred there, but has become extremely rare or disappeared since
<bibRefCitation id="EF8139CDC732FF9306CC2B90F79FFCA0" author="Lortet, L." box="[749,919,763,789]" pageId="23" pageNumber="174" pagination="99 - 194" refId="ref16560" refString="Lortet, L. (1883) Poissons et reptiles du Lac de Tiberiade et de quelques autres parties de la Syrie. Archives du Museum d'histoire naturelle de Lyon, 3, 99 - 194. https: // doi. org / 10.3406 / mhnly. 1883.941" type="journal article" year="1883">Lortets (1883)</bibRefCitation>
observations. Indeed, there is a photographic record of a tortoise from the Al-Badia taken between 1996 and 1999 (
<bibRefCitation id="EF8139CDC732FF9307BA2A74F07FFC8C" author="Sindaco, R. &amp; Serra, G. &amp; Menegon, M." box="[923,1143,799,825]" pageId="23" pageNumber="174" pagination="29 - 38" refId="ref19307" refString="Sindaco, R., Serra, G. &amp; Menegon, M. (2006) New data on the Syrian herpetofauna, with a newly recorded species of snake. Zoology in the Middle East, 37, 29 - 38. https: // doi. org / 10.1080 / 09397140.2006.10638146" type="journal article" year="2006">
Sindaco
<emphasis id="B964982EC732FF9307DC2A74F03DFC8C" box="[1021,1077,799,825]" italics="true" pageId="23" pageNumber="174">et al.</emphasis>
2006
</bibRefCitation>
), and a photo of a juvenile from Palmyra was published on iNaturalist in 2018. However, the latter photo shows that the colors and texture of the shell are similar to tortoises from the coastal region of
<collectingCountry id="F30704ACC732FF9307012A0CF756FC34" box="[800,862,871,897]" name="Syria" pageId="23" pageNumber="174">Syria</collectingCountry>
, rather than the morphotypes found in arid habitats, suggestive of an introduced individual.
</paragraph>
<paragraph id="8BAF443CC732FF9304E62AC4F151FB80" blockId="23.[151,1437,151,1581]" pageId="23" pageNumber="174">
<materialsCitation id="3B784E61C732FF9304E62AC4F15DFB80" collectingDate="1991-08" collectingDateMax="2020-06" collectingDateMin="1991-08" collectionCode="MTD" country="Syria" county="In" location="Turkish" municipality="One" pageId="23" pageNumber="174" specimenCode="MTD 32111" specimenCount="1" stateProvince="Al Hasakah">
<collectingCounty id="62CE3CB0C732FF9304E62AC4F4D6FC7C" box="[199,222,943,969]" pageId="23" pageNumber="174">In</collectingCounty>
northeastern
<collectingCountry id="F30704ACC732FF9305582AC4F5BFFC7C" box="[377,439,943,969]" name="Syria" pageId="23" pageNumber="174">Syria</collectingCountry>
, tortoises are very rare.
<collectingMunicipality id="6BCBDE46C732FF9306E82AC4F6F0FC7C" box="[713,760,943,969]" pageId="23" pageNumber="174">One</collectingMunicipality>
individual was found in
<date id="FFAE62FCC732FF9300322AC4F0ACFC7C" box="[1043,1188,943,969]" pageId="23" pageNumber="174" value="1991-08">
<collectingDate id="EFEA9B14C732FF9300322AC4F0ACFC7C" box="[1043,1188,943,969]" pageId="23" pageNumber="174" value="1991-08">August 1991</collectingDate>
</date>
near
<collectingRegion id="49D48ADEC732FF9300C22AC4F153FC7C" box="[1251,1371,943,969]" country="Syria" name="Al Hasakah" pageId="23" pageNumber="174">Al-Hasaka</collectingRegion>
close to the
<location id="8ECF12E7C732FF9304FD2AB8F53AFC58" LSID="urn:lsid:plazi:treatment:03B9F52AC731FF9304B62F9CF61AF998:8ECF12E7C732FF9304FD2AB8F53AFC58" box="[220,306,979,1005]" country="Syria" county="In" municipality="One" name="Turkish" pageId="23" pageNumber="174" stateProvince="Al Hasakah">Turkish</location>
border (
<specimenCode id="DBB6EC47C732FF9305B02AB8F616FC58" box="[401,542,979,1005]" collectionCode="MTD" country="Germany" httpUri="http://grbio.org/cool/4j3t-w40g" name="Museum of Zoology Senckenberg Dresden" pageId="23" pageNumber="174">MTD 32111</specimenCode>
, see also
<bibRefCitation id="EF8139CDC732FF9306A92AB8F736FC58" author="Fritz, U. &amp; Bischoff, W. &amp; Martens, H. &amp; Schmidtler, J. F." box="[648,830,979,1005]" pageId="23" pageNumber="174" pagination="5 - 14" refId="ref14783" refString="Fritz, U., Bischoff, W., Martens, H. &amp; Schmidtler, J. F. (1996) Variabilitat syrischer Landschildkroten (Testudo graeca) sowie zur Systematik und Zoogeographie im Nahen Osten und in Nordafrika. Herpetofauna, 18 (104), 5 - 14." type="journal article" year="1996">
Fritz
<emphasis id="B964982EC732FF9306E22AB8F6FCFC58" box="[707,756,979,1005]" italics="true" pageId="23" pageNumber="174">et al</emphasis>
. 1996
</bibRefCitation>
), and another specimen was found in
<date id="FFAE62FCC732FF9300C22AB8F15DFC58" box="[1251,1365,979,1005]" pageId="23" pageNumber="174" value="2020-06">
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June
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</collectingDate>
</date>
in
<location id="8ECF12E7C732FF9301562AB8F4F1FBA4" LSID="urn:lsid:plazi:treatment:03B9F52AC731FF9304B62F9CF61AF998:8ECF12E7C732FF9301562AB8F4F1FBA4" country="Syria" county="In" municipality="One" name="Al-Hardana" pageId="23" pageNumber="174" stateProvince="Al Hasakah">Al-Hardana</location>
, also near the Turkish border (
<figureCitation id="132B58B9C732FF9306672A9CF6A6FBA4" box="[582,686,1015,1041]" captionStart="FIGURE 20" captionStartId="21.[151,250,1732,1756]" captionTargetBox="[151,1435,849,1709]" captionTargetId="figure-266@21.[151,1435,849,1709]" captionTargetPageId="21" captionText="FIGURE 20. Coloration of Testudo graeca terrestris. (A) Al-Hardana, north of Mesopotamia (photo: Mohammed Al-Karraf); (B) Maqta Hajar, northeast Aleppo (photo: Ahmad Aidek); (C) Majdalun Al-Bustan, coastal region (photo: Ali Ali); (D) Al- Bassah, coastal region (photo: Ali Alloush); (E) Sadd Al-Kafat, southeast Hamah (photo: Ehab Jbr); (F) as-Sabbourah, west of Damascus (photo: Wael Almatni); (G) Masil Al-Fawwar, southwestern Syria (photo: Hans Esterbauer); (H) hatchling, Shahba, Jabal Al-Arab (photo: Ahmad Aidek); (I) hatchling, Tell Sheihan, Jabal Al-Arab (photo: Wael Almatni)." pageId="23" pageNumber="174">Fig. 20A</figureCitation>
). During dozens of field trips throughout the area between 2006 and 2023, no other tortoises were found, but locals confirmed that they are present but rare (Aidek pers. observ.)
</materialsCitation>
.
</paragraph>
<paragraph id="8BAF443CC732FF9304E62D54F7F3FB70" blockId="23.[151,1437,151,1581]" pageId="23" pageNumber="174">
There are also few records from the Euphrates region. In 1999, four tortoises were obtained from
<quantity id="4CE8E9D9C732FF9301272D54F144FBEC" box="[1286,1356,1087,1113]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="174" unit="km" value="10.0">10 km</quantity>
east of
<collectingRegion id="49D48ADEC732FF9304B62D08F514FBC8" box="[151,284,1123,1149]" country="Syria" name="Dayr az Zawr" pageId="23" pageNumber="174">Deir ez-Zor</collectingRegion>
(ZFMK 8665686659). These specimens are probably escaped or captive tortoises. This is suggested by the “locality names” of the ZFMK specimens, as these are names of people, not of sites. Two tortoises were also seen on agricultural land in Muhasan,
<quantity id="4CE8E9D9C732FF9306652DC0F683FB70" box="[580,651,1195,1221]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="23" pageNumber="174" unit="km" value="20.0">20 km</quantity>
east of
<collectingRegion id="49D48ADEC732FF9306C32DC0F76FFB70" box="[738,871,1195,1221]" country="Syria" name="Dayr az Zawr" pageId="23" pageNumber="174">Deir ez-Zor</collectingRegion>
, in fall 2021.
</paragraph>
</subSubSection>
<subSubSection id="C30A17B7C732FF9304E62DA4F61AF998" pageId="23" pageNumber="174" type="discussion">
<paragraph id="8BAF443CC732FF9304E62DA4F4E1FA74" blockId="23.[151,1437,151,1581]" pageId="23" pageNumber="174">
<emphasis id="B964982EC732FF9304E62DA4F53CFB5C" bold="true" box="[199,308,1231,1257]" pageId="23" pageNumber="174">Remarks</emphasis>
: Syrian tortoises are characterized by a great variety of coloration. The primary color of the shell varies from yellowish beige with various shades of light and dark brown, orange and gray to very dark
<typeStatus id="54ABFA9EC732FF9301092D98F16BFAB8" box="[1320,1379,1267,1293]" pageId="23" pageNumber="174">types</typeStatus>
with a primary black shell coloration. Tortoises from the humid coastal region are generally characterized by a shiny shell and lighter colors compared to tortoises from the
<collectingCountry id="F30704ACC732FF9307242C50F76FFAE0" box="[773,871,1339,1365]" name="Lebanon" pageId="23" pageNumber="174">Lebanon</collectingCountry>
Mountains and the semi-arid area east of
<collectingRegion id="49D48ADEC732FF9301622C50F191FAE0" box="[1347,1433,1339,1365]" country="Syria" name="Halab" pageId="23" pageNumber="174">Aleppo</collectingRegion>
. Tortoises from the black basalt regions (Jabal Al-Arab) are generally black or dark gray and dull in color (
<figureCitation id="132B58B9C732FF93014E2C34F4BEFA28" captionStart="FIGURE 20" captionStartId="21.[151,250,1732,1756]" captionTargetBox="[151,1435,849,1709]" captionTargetId="figure-266@21.[151,1435,849,1709]" captionTargetPageId="21" captionText="FIGURE 20. Coloration of Testudo graeca terrestris. (A) Al-Hardana, north of Mesopotamia (photo: Mohammed Al-Karraf); (B) Maqta Hajar, northeast Aleppo (photo: Ahmad Aidek); (C) Majdalun Al-Bustan, coastal region (photo: Ali Ali); (D) Al- Bassah, coastal region (photo: Ali Alloush); (E) Sadd Al-Kafat, southeast Hamah (photo: Ehab Jbr); (F) as-Sabbourah, west of Damascus (photo: Wael Almatni); (G) Masil Al-Fawwar, southwestern Syria (photo: Hans Esterbauer); (H) hatchling, Shahba, Jabal Al-Arab (photo: Ahmad Aidek); (I) hatchling, Tell Sheihan, Jabal Al-Arab (photo: Wael Almatni)." pageId="23" pageNumber="174">Fig. 20</figureCitation>
; see also
<bibRefCitation id="EF8139CDC732FF9305052CE8F5D4FA28" author="Fritz, U. &amp; Bischoff, W. &amp; Martens, H. &amp; Schmidtler, J. F." box="[292,476,1411,1437]" pageId="23" pageNumber="174" pagination="5 - 14" refId="ref14783" refString="Fritz, U., Bischoff, W., Martens, H. &amp; Schmidtler, J. F. (1996) Variabilitat syrischer Landschildkroten (Testudo graeca) sowie zur Systematik und Zoogeographie im Nahen Osten und in Nordafrika. Herpetofauna, 18 (104), 5 - 14." type="journal article" year="1996">
Fritz
<emphasis id="B964982EC732FF9305402CE8F592FA28" box="[353,410,1411,1437]" italics="true" pageId="23" pageNumber="174">et al.</emphasis>
1996
</bibRefCitation>
,
<bibRefCitation id="EF8139CDC732FF9305C92CE8F62DFA28" author="Fritz, U. &amp; Hundsdorfer, A. K. &amp; Siroky, P. &amp; Auer, M. &amp; Kami, H. &amp; Lehmann, J. &amp; Mazanaeva, L. F. &amp; Turkozan, O. &amp; Wink, M." box="[488,549,1411,1437]" pageId="23" pageNumber="174" pagination="97 - 121" refId="ref14837" refString="Fritz, U., Hundsdorfer, A. K., Siroky, P., Auer, M., Kami, H., Lehmann, J., Mazanaeva, L. F., Turkozan, O. &amp; Wink, M. (2007) Phenotypic plasticity leads to incongruence between morphology-based taxonomy and genetic differentiation in western Palaearctic tortoises (Testudo graeca complex; Testudines, Testudinidae). Amphibia-Reptilia, 28, 97 - 121. https: // doi. org / 10.1163 / 156853807779799135" type="journal article" year="2007">2007</bibRefCitation>
). Variation beyond
<collectingCountry id="F30704ACC732FF9307202CE8F734FA28" box="[769,828,1411,1437]" name="Syria" pageId="23" pageNumber="174">Syria</collectingCountry>
is discussed in
<bibRefCitation id="EF8139CDC732FF9307D12CE8F0B0FA28" author="Fritz, U. &amp; Hundsdorfer, A. K. &amp; Siroky, P. &amp; Auer, M. &amp; Kami, H. &amp; Lehmann, J. &amp; Mazanaeva, L. F. &amp; Turkozan, O. &amp; Wink, M." box="[1008,1208,1411,1437]" pageId="23" pageNumber="174" pagination="97 - 121" refId="ref14837" refString="Fritz, U., Hundsdorfer, A. K., Siroky, P., Auer, M., Kami, H., Lehmann, J., Mazanaeva, L. F., Turkozan, O. &amp; Wink, M. (2007) Phenotypic plasticity leads to incongruence between morphology-based taxonomy and genetic differentiation in western Palaearctic tortoises (Testudo graeca complex; Testudines, Testudinidae). Amphibia-Reptilia, 28, 97 - 121. https: // doi. org / 10.1163 / 156853807779799135" type="journal article" year="2007">
Fritz
<emphasis id="B964982EC732FF93000C2CE8F06EFA28" box="[1069,1126,1411,1437]" italics="true" pageId="23" pageNumber="174">et al.</emphasis>
(2007)
</bibRefCitation>
and
<bibRefCitation id="EF8139CDC732FF9300D12CE8F4EDFA74" author="Turkozan, O. &amp; Javanbakht, H. &amp; Mazanaeva, L. &amp; Meiri, S. &amp; Kornilev, Y. V. &amp; Tzoras, E. &amp; Popgeorgiev, G. &amp; Shanas, U. &amp; Escoriza, D." pageId="23" pageNumber="174" refId="ref20465" refString="Turkozan, O., Javanbakht, H., Mazanaeva, L., Meiri, S., Kornilev, Y. V., Tzoras, E., Popgeorgiev, G., Shanas, U. &amp; Escoriza, D. (2023) Testudo graeca Linnaeus 1758 (eastern subspecies clades: Testudo g. armeniaca, Testudo g. buxtoni, Testudo g. ibera, Testudo g. terrestris, Testudo g. zarudnyi) - Armenian tortoise, Zagros tortoise, Anatolian tortoise, Levantine tortoise, Kerman tortoise. In: Rhodin, A. G. J., Iverson, J. B., van Dijk, P. P., Stanford, C. B., Goode, E. V., Buhlmann, K. A. &amp; Mittermeier, R. A. (Eds.), Conservation biology of freshwater turtles and tortoises: A compilation project of the IUCN / SSC Tortoise and Freshwater Turtle Specialist Group. Chelonian Research Monographs, 5, 120.1 - 120.33. https: // doi. org / 10.3854 / crm. 5.120. eastern. graeca. v 1.2023" type="journal volume" year="2023">
Türkozan
<emphasis id="B964982EC732FF9301422CE8F195FA28" box="[1379,1437,1411,1437]" italics="true" pageId="23" pageNumber="174">et al.</emphasis>
(2023)
</bibRefCitation>
.
</paragraph>
<paragraph id="8BAF443CC732FF9304E62CA0F61AF998" blockId="23.[151,1437,151,1581]" pageId="23" pageNumber="174">
There are 117 records from
<collectingCountry id="F30704ACC732FF9305DB2CA0F63DFA50" box="[506,565,1483,1509]" name="Syria" pageId="23" pageNumber="174">Syria</collectingCountry>
on GBIF. However, of these supposedly Syrian records, 10 are not from
<collectingCountry id="F30704ACC732FF93017B2CA0F191FA50" box="[1370,1433,1483,1509]" name="Syria" pageId="23" pageNumber="174">Syria</collectingCountry>
. Another 76 records are on iNaturalist (2023). Many of these have incorrect coordinates and are in the sea, some are duplicates and refer to other sites.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,206 @@
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<taxonomicName id="4C103FBFC736FF9704B62BD8F639FD79" ID-CoL="58WKP" authority="(Forskal, 1775)" baseAuthorityName="Forskal" baseAuthorityYear="1775" box="[151,561,689,717]" class="Testudines" family="Trionychidae" genus="Trionyx" kingdom="Animalia" order="Cryptodira" pageId="19" pageNumber="170" phylum="Chordata" rank="species" species="triunguis">
<emphasis id="B964982EC736FF9704B62BD8F639FD79" bold="true" box="[151,561,689,717]" pageId="19" pageNumber="170">
<emphasis id="B964982EC736FF9704B62BD8F565FD78" bold="true" box="[151,365,691,717]" italics="true" pageId="19" pageNumber="170">Trionyx triunguis</emphasis>
(
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)
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<paragraph id="8BAF443CC736FF9704B62BBCF6F5FD44" blockId="19.[151,765,689,753]" box="[151,765,727,753]" pageId="19" pageNumber="170">
African Softshell Turtle, Nile Softshell Turtle (
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)
</paragraph>
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<subSubSection id="C30A17B7C736FF9704B62A75F538FCE8" pageId="19" pageNumber="170" type="description">
<paragraph id="8BAF443CC736FF9704B62A75F538FCE8" blockId="19.[151,1437,798,1509]" pageId="19" pageNumber="170">
<emphasis id="B964982EC736FF9704B62A75F5B7FC8D" bold="true" box="[151,447,798,825]" pageId="19" pageNumber="170">IUCN Red List Category</emphasis>
: Vulnerable (VU A4bcd); Mediterranean subpopulation Critically Endangered (CR C2a;
<bibRefCitation id="EF8139CDC736FF9704B62A28F520FCE8" author="TTWG &amp; Turtle Taxonomy Working Group" box="[151,296,835,861]" pageId="19" pageNumber="170" pagination="1 - 472" refId="ref20400" refString="TTWG [Turtle Taxonomy Working Group] (2021) Turtles of the World: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status (9 th Ed.). Chelonian Research Monographs, 8, 1 - 472. https: // doi. org / 10.3854 / crm. 8. checklist. atlas. v 9.2021" type="journal article" year="2021">TTWG 2021</bibRefCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30A17B7C736FF9704E62A0CF5C2FAE0" pageId="19" pageNumber="170" type="materials_examined">
<paragraph id="8BAF443CC736FF9704E62A0CF6A8FC34" blockId="19.[151,1437,798,1509]" box="[199,672,870,897]" pageId="19" pageNumber="170">
<materialsCitation id="3B784E61C736FF9704E62A0CF694FC34" box="[199,668,870,897]" country="Egypt" location="Nile River" pageId="19" pageNumber="170" specimenCount="1" typeStatus="holotype">
<emphasis id="B964982EC736FF9704E62A0CF568FC35" bold="true" box="[199,352,870,897]" pageId="19" pageNumber="170">
<typeStatus id="54ABFA9EC736FF9704E62A0CF50AFC34" box="[199,258,871,897]" pageId="19" pageNumber="170">Type</typeStatus>
locality
</emphasis>
: “Nilo” [
<location id="8ECF12E7C736FF9705E62A0CF649FC34" LSID="urn:lsid:plazi:treatment:03B9F52AC736FF9704B62BD8F054FA50:8ECF12E7C736FF9705E62A0CF649FC34" box="[455,577,871,897]" country="Egypt" name="Nile River" pageId="19" pageNumber="170">Nile River</location>
,
<collectingCountry id="F30704ACC736FF97066D2A0CF69DFC34" box="[588,661,871,897]" name="Egypt" pageId="19" pageNumber="170">Egypt</collectingCountry>
]
</materialsCitation>
.
</paragraph>
<paragraph id="8BAF443CC736FF9704E62AE0F777FC7C" blockId="19.[151,1437,798,1509]" pageId="19" pageNumber="170">
<emphasis id="B964982EC736FF9704E62AE0F57EFC11" bold="true" box="[199,374,906,933]" pageId="19" pageNumber="170">Type specimen</emphasis>
:
<materialsCitation id="3B784E61C736FF9705A22AE0F773FC7C" collectionCode="MNHN" pageId="19" pageNumber="170" specimenCode="MNHN 4147" specimenCount="2" typeStatus="lectotype">
Muséum National dHistoire naturelle, Paris,
<specimenCode id="DBB6EC47C736FF97075D2AE0F01AFC10" box="[892,1042,907,933]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="19" pageNumber="170">MNHN 4147</specimenCode>
,
<typeStatus id="54ABFA9EC736FF97003D2AE0F08BFC10" box="[1052,1155,907,933]" pageId="19" pageNumber="170" type="lectotype">lectotype</typeStatus>
designated by
<bibRefCitation id="EF8139CDC736FF9701092AE0F4EEFC7C" author="Bour, R. &amp; Dubois, A. &amp; Webb, R. G." pageId="19" pageNumber="170" pagination="73 - 92" refId="ref13884" refString="Bour, R., Dubois, A. &amp; Webb, R. G. (1995) Types of recent trionychid turtles in the Museum national d'Histoire naturelle, Paris. Dumerilia, 2, 73 - 92." type="journal article" year="1995">
Bour
<emphasis id="B964982EC736FF9701442AE0F195FC10" box="[1381,1437,907,933]" italics="true" pageId="19" pageNumber="170">et al.</emphasis>
(1995)
</bibRefCitation>
. There are
<specimenCount id="9D168FB5C736FF9705402AC4F63FFC7C" box="[353,567,943,969]" count="2" pageId="19" pageNumber="170" type="generic" typeStatus="paralectotype">two paralectotypes</specimenCount>
,
<collectionCode id="ED01DCF9C736FF9706632AC4F690FC7C" box="[578,664,943,969]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="19" pageNumber="170">MNHN</collectionCode>
A.5186 and A.5227
</materialsCitation>
.
</paragraph>
<paragraph id="8BAF443CC736FF9704E62AB9F686FC58" blockId="19.[151,1437,798,1509]" box="[199,654,978,1005]" pageId="19" pageNumber="170">
<emphasis id="B964982EC736FF9704E62AB9F54EFC59" bold="true" box="[199,326,978,1004]" pageId="19" pageNumber="170">Chorotype</emphasis>
: Afrotropico-Mediterranean.
</paragraph>
<paragraph id="8BAF443CC736FF9704E62A9DF6F7FB80" blockId="19.[151,1437,798,1509]" pageId="19" pageNumber="170">
<emphasis id="B964982EC736FF9704E62A9DF5C3FBA5" bold="true" box="[199,459,1014,1041]" pageId="19" pageNumber="170">
First record for
<collectingCountry id="F30704ACC736FF9705AA2A9DF5C3FBA5" box="[395,459,1014,1040]" name="Syria" pageId="19" pageNumber="170">Syria</collectingCountry>
</emphasis>
: Nahr Al-Kabir Al-Janoubi, dating back to the “
<emphasis id="B964982EC736FF9707D12A9CF0B1FBA4" box="[1008,1209,1015,1041]" italics="true" pageId="19" pageNumber="170">Naturalis historia</emphasis>
” by Pliny the Elder (Gaius Secundus Plinius 7779, reprinted 18551857).
</paragraph>
<paragraph id="8BAF443CC736FF9704E62D55F653FBC8" blockId="19.[151,1437,798,1509]" pageId="19" pageNumber="170">
<emphasis id="B964982EC736FF9704E62D55F577FBEC" bold="true" box="[199,383,1086,1113]" pageId="19" pageNumber="170">Literature data</emphasis>
: Middle Orontes River in Ain Taqa in the Ghab Valley (
<bibRefCitation id="EF8139CDC736FF9707D72D54F0B3FBEC" author="Kinzelbach, R." box="[1014,1211,1087,1113]" pageId="19" pageNumber="170" pagination="83 - 87" refId="ref16304" refString="Kinzelbach, R. (1986) Recent records of the Nile soft-shelled turtle, Trionyx triunguis, and the Euphrates soft-shelled turtle, Trionyx euphraticus, in the Middle East. Zoology in the Middle East, 1, 83 - 87. https: // doi. org / 10.1080 / 09397140.1986.10637519" type="journal article" year="1986">Kinzelbach 1986</bibRefCitation>
), Nahr Al-Kabir Al-Janoubi (
<bibRefCitation id="EF8139CDC736FF9704DC2D08F647FBC8" author="Kasparek, M. &amp; Kinzelbach, R." box="[253,591,1123,1149]" pageId="19" pageNumber="170" pagination="139 - 159" refId="ref16264" refString="Kasparek, M. &amp; Kinzelbach, R. (1991) Distribution and bionomics of the Nile soft-shelled turtle, Trionyx triunguis, in the Eastern Mediterranean. Zeitschrift fur Angewandte Zoologie, 78, 139 - 159." type="journal article" year="1991">Kasparek &amp; Kinzelbach 1991</bibRefCitation>
).
</paragraph>
<paragraph id="8BAF443CC736FF9704E62DECF5C2FAE0" blockId="19.[151,1437,798,1509]" pageId="19" pageNumber="170">
<emphasis id="B964982EC736FF9704E62DECF574FB14" bold="true" box="[199,380,1159,1185]" pageId="19" pageNumber="170">Recent records</emphasis>
: An individual was swimming in the Nahr Al-Kabir Al-Janobi River on the Syrian-Lebanese border, on
<date id="FFAE62FCC736FF9705372DC0F5C4FB70" box="[278,460,1195,1221]" pageId="19" pageNumber="170" value="2004-08-04">August 4, 2004</date>
, during an afternoon field survey (Saad &amp; Rees unpubl.). Another individual in the village of Al-Qabou was transferred on
<date id="FFAE62FCC736FF9706732DA4F6E1FB5C" box="[594,745,1230,1257]" pageId="19" pageNumber="170" value="2005-07-16">July 16, 2005</date>
from a spring to a freshwater pool near Sanwbar Beach (Saad unpubl.). There are two records from the upper Orontes River. One
<taxonomicName id="4C103FBFC736FF9707B82D98F013FAB9" baseAuthorityName="Forskal" baseAuthorityYear="1775" box="[921,1051,1267,1292]" class="Testudines" family="Trionychidae" genus="Trionyx" kingdom="Animalia" order="Cryptodira" pageId="19" pageNumber="170" phylum="Chordata" rank="species" species="triunguis">
<emphasis id="B964982EC736FF9707B82D98F013FAB9" box="[921,1051,1267,1292]" italics="true" pageId="19" pageNumber="170">T. triunguis</emphasis>
</taxonomicName>
was caught by a fisherman in Al-Alani on
<date id="FFAE62FCC736FF9705202C7CF5CEFA84" box="[257,454,1302,1329]" pageId="19" pageNumber="170" value="2020-10-15">October 15, 2020</date>
(
<figureCitation id="132B58B9C736FF9705F72C7CF634FA84" box="[470,572,1303,1329]" captionStart="FIGURE 18" captionStartId="19.[151,250,1982,2006]" captionTargetBox="[151,1436,1533,1958]" captionTargetId="figure-464@19.[151,1436,1533,1958]" captionTargetPageId="19" captionText="FIGURE 18. Nile Softshell Turtle, Trionyx triunguis. (A) Young individual, southern coast of Lebanon (photo: Rami Khashab); (B) adult individual, caught and killed by a fisherman, Al-Alani, upper Orontes (photo: H. Baniasi)." pageId="19" pageNumber="170">Fig. 18B</figureCitation>
), and another one was found dead on the riverbank in Al-Hamziyeh on
<date id="FFAE62FCC736FF9701482C7CF4F1FAE0" pageId="19" pageNumber="170" value="2022-05-17">May 17, 2022</date>
(Baniasi unpubl.).
</paragraph>
</subSubSection>
<subSubSection id="C30A17B7C736FF9704E62C35F116FACC" box="[199,1310,1374,1401]" pageId="19" pageNumber="170" type="distribution">
<paragraph id="8BAF443CC736FF9704E62C35F116FACC" blockId="19.[151,1437,798,1509]" box="[199,1310,1374,1401]" pageId="19" pageNumber="170">
<emphasis id="B964982EC736FF9704E62C35F5B7FACD" bold="true" box="[199,447,1374,1400]" pageId="19" pageNumber="170">
Distribution in
<collectingCountry id="F30704ACC736FF97055E2C35F5B7FACD" box="[383,447,1374,1400]" name="Syria" pageId="19" pageNumber="170">Syria</collectingCountry>
</emphasis>
: Restricted to the Nahr Al-Kabir Al-Janoubi and the Orontes River (
<figureCitation id="132B58B9C736FF97009C2C34F118FACC" box="[1213,1296,1375,1401]" captionStart="FIGURE 19" captionStartId="20.[151,250,1389,1413]" captionTargetBox="[161,1429,191,1356]" captionTargetId="figure-21@20.[151,1436,181,1365]" captionTargetPageId="20" captionText="FIGURE 19. Records of Trionyx triunguis in Syria. Black circle: previous record, blue circle: recently confirmed previous record, red circles: new records." pageId="19" pageNumber="170">Fig. 19</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30A17B7C736FF9704E62CE8F054FA50" pageId="19" pageNumber="170" type="discussion">
<paragraph id="8BAF443CC736FF9704E62CE8F054FA50" blockId="19.[151,1437,798,1509]" pageId="19" pageNumber="170">
<emphasis id="B964982EC736FF9704E62CE8F53CFA28" bold="true" box="[199,308,1411,1437]" pageId="19" pageNumber="170">Remarks</emphasis>
:
<bibRefCitation id="EF8139CDC736FF9705622CE8F601FA28" author="Fritz, U. &amp; Tolley, K. A. &amp; Vamberger, M. &amp; Ihlow, F." box="[323,521,1411,1437]" pageId="19" pageNumber="170" pagination="951 - 969" refId="ref14931" refString="Fritz, U., Tolley, K. A., Vamberger, M. &amp; Ihlow, F. (2022) Phylogeny and phylogeography of chelonians from sub-Saharan Africa - A review of current knowledge in tribute to Margaretha D. Hofmeyr. Vertebrate Zoology, 72, 951 - 969. https: // doi. org / 10.3897 / vz. 72. e 95681" type="journal article" year="2022">
Fritz
<emphasis id="B964982EC736FF97055E2CE8F5B8FA28" box="[383,432,1411,1437]" italics="true" pageId="19" pageNumber="170">et al</emphasis>
. (2022)
</bibRefCitation>
reviewed several studies providing anecdotal information on the phylogeography of
<taxonomicName id="4C103FBFC736FF9704962CCCF532FA75" baseAuthorityName="Forskal" baseAuthorityYear="1775" box="[183,314,1447,1472]" class="Testudines" family="Trionychidae" genus="Trionyx" kingdom="Animalia" order="Cryptodira" pageId="19" pageNumber="170" phylum="Chordata" rank="species" species="triunguis">
<emphasis id="B964982EC736FF9704962CCCF532FA75" box="[183,314,1447,1472]" italics="true" pageId="19" pageNumber="170">T. triunguis</emphasis>
</taxonomicName>
. Mediterranean populations, presumably including those from
<collectingCountry id="F30704ACC736FF9700282CCCF04FFA74" box="[1033,1095,1447,1473]" name="Syria" pageId="19" pageNumber="170">Syria</collectingCountry>
, represent a single population connected across the sea, while those from sub-Saharan Africa appear to be divergent.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,319 @@
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<emphasis id="B91D952CFFB23A1F95F4FB78FD1531F0" bold="true" box="[151,663,1247,1274]" pageId="5" pageNumber="232">
<emphasis id="B91D952CFFB23A1F95F4FB78FE6731F0" bold="true" box="[151,485,1247,1274]" italics="true" pageId="5" pageNumber="232">Labidodemas semperianum</emphasis>
<bibRefCitation id="EFF834CFFFB23A1F948FFB78FD1531F0" author="Selenka, E." box="[492,663,1247,1273]" pageId="5" pageNumber="232" pagination="291 - 374" refId="ref8895" refString="Selenka, E. (1867) Beitrage zur anatomie und systematik der holothurien. Zeitschrift fur wissenschaftliche Zoologie, 17 (2), 291 - 374." type="journal article" year="1867">Selenka, 1867</bibRefCitation>
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<paragraph id="8BD6493EFFB23A1F95F4FAA4FEAC3017" blockId="5.[151,663,1247,1310]" box="[151,302,1283,1310]" pageId="5" pageNumber="232">
(
<figureCitation id="135255BBFFB23A1F95FCFAA4FEA43017" box="[159,294,1283,1310]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStart-3="FIGURE 8" captionStartId-0="6.[151,250,923,947]" captionStartId-1="6.[151,250,1976,2000]" captionStartId-2="7.[151,250,1473,1497]" captionStartId-3="8.[151,250,1094,1118]" captionTargetBox-0="[257,1329,181,899]" captionTargetBox-1="[264,1322,1007,1952]" captionTargetBox-2="[257,1329,181,1449]" captionTargetBox-3="[265,1322,181,1069]" captionTargetId-0="figure-25@6.[257,1329,178,899]" captionTargetId-1="figure-49@6.[264,1322,1004,1954]" captionTargetId-2="figure-17@7.[257,1329,181,1449]" captionTargetId-3="figure-25@8.[265,1322,181,1069]" captionTargetPageId-0="6" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionText-0="FIGURE 5. Labidodemas semperianum Selenka, 1867. MZB Hol.385 (68x19 mm in ethanol). Specimen photograph.A: dorsal; B: ventral; C: calcareous ring." captionText-1="FIGURE 6. Labidodemas semperianum Selenka, 1867. MZB Hol.385. Ossicles dorsal body wall, A: tables; B: pseudo-button. Ossicles ventral body wall, C: tables; D: buttons." captionText-2="FIGURE 7. Labidodemas semperianum Selenka, 1867. MZB Hol.385. Ossicles dorsal papillae, A: tables; B: buttons, pseudobuttons and rods; C: plate. Ossicles ventral tube feet, D: tables; E: endplate." captionText-3="FIGURE 8. Labidodemas semperianum Selenka, 1867. MZB Hol.385. A: tentacle rods; B: ossicles of anal papillae." pageId="5" pageNumber="232">Figures 58</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3731AB5FFB23A1F95F4FAE6FE473071" pageId="5" pageNumber="232" type="reference_group">
<paragraph id="8BD6493EFFB23A1F95F4FAE6FE473071" blockId="5.[151,1436,1345,1400]" pageId="5" pageNumber="232">
<treatmentCitationGroup id="AB796E10FFB23A1F95F4FAE6FE473071" pageId="5" pageNumber="232">
<treatmentCitation id="0AC86F2FFFB23A1F95F4FAE6FDF03050" author="Selenka, E." box="[151,626,1345,1369]" page="309" pageId="5" pageNumber="232" year="1867">
<taxonomicName id="4C6932BDFFB23A1F95F4FAE6FDF03050" ID-CoL="3RJJM" authority="Selenka, 1867: 309" authorityName="Selenka" authorityPageNumber="309" authorityYear="1867" box="[151,626,1345,1369]" class="Holothuroidea" family="Holothuriidae" genus="Labidodemas" kingdom="Animalia" order="Holothuriida" pageId="5" pageNumber="232" phylum="Echinodermata" rank="species" species="semperianum">
<emphasis id="B91D952CFFB23A1F95F4FAE6FE2E3050" box="[151,428,1345,1369]" italics="true" pageId="5" pageNumber="232">Labidodemas semperianum</emphasis>
<bibRefCitation id="EFF834CFFFB23A1F94D2FAE6FDF03050" author="Selenka, E." box="[433,626,1345,1369]" pageId="5" pageNumber="232" pagination="291 - 374" refId="ref8895" refString="Selenka, E. (1867) Beitrage zur anatomie und systematik der holothurien. Zeitschrift fur wissenschaftliche Zoologie, 17 (2), 291 - 374." type="journal article" year="1867">Selenka, 1867: 309</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. 17, figs. 13;
<treatmentCitation id="0AC86F2FFFB23A1F967CFAE6FB993050" author="Massin, C. &amp; Samyn, Y. &amp; Thandar, A. S." box="[799,1051,1345,1369]" page="1833" pageId="5" pageNumber="232" year="2004">
<bibRefCitation id="EFF834CFFFB23A1F967CFAE6FB993050" author="Massin, C. &amp; Samyn, Y. &amp; Thandar, A. S." box="[799,1051,1345,1369]" pageId="5" pageNumber="232" pagination="1811 - 1847" refId="ref8451" refString="Massin, C., Samyn, Y. &amp; Thandar, A. S. (2004) The genus Labidodemas (Holothuroidea: Aspidochirotida) revisited, with description of three new species and with re-positioning of Holothuria (Irenothuria) maccullochi Deichmann, 1958. Journal of Natural History, 38, 1811 - 1847. https: // doi. org / 10.1080 / 0022293031000156268" type="journal article" year="2004">
Massin
<emphasis id="B91D952CFFB23A1F960FFAE5FC153050" box="[876,919,1345,1369]" italics="true" pageId="5" pageNumber="232">et al</emphasis>
., 2004: 1833
</bibRefCitation>
</treatmentCitation>
, figs. 9AJ, 10AH, map 6 (synonymy and records 19702003).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3731AB5FFB23A1295F4FA07FA9131E8" lastPageId="8" lastPageNumber="235" pageId="5" pageNumber="232" type="materials_examined">
<paragraph id="8BD6493EFFB23A1F95F4FA07FD8A330B" blockId="5.[151,1437,1439,2041]" pageId="5" pageNumber="232">
<emphasis id="B91D952CFFB23A1F95F4FA07FEC330B0" bold="true" box="[151,321,1439,1466]" pageId="5" pageNumber="232">Type material</emphasis>
.
<typeStatus id="54D2F79CFFB23A1F9430FA07FE4130B3" box="[339,451,1440,1466]" pageId="5" pageNumber="232" type="lectotype">Lectotype</typeStatus>
MCZ 736 (the dissected specimen,
<quantity id="4C91E4DBFFB23A1F960FFA38FC3D30B3" box="[876,959,1439,1466]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" pageId="5" pageNumber="232" unit="mm" value="75.0">75 mm</quantity>
long),
<typeStatus id="54D2F79CFFB23A1F9175FA38FB2F30B0" box="[1046,1197,1439,1465]" pageId="5" pageNumber="232" type="paralectotype">paralectotype</typeStatus>
MCZ 736 (the nondissected specimen,
<quantity id="4C91E4DBFFB23A1F941FFA63FE4930D7" box="[380,459,1476,1502]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="232" unit="mm" value="90.0">90 mm</quantity>
long) (
<bibRefCitation id="EFF834CFFFB23A1F9778FA64FD7230D7" author="Massin, C. &amp; Samyn, Y. &amp; Thandar, A. S." box="[539,752,1475,1502]" pageId="5" pageNumber="232" pagination="1811 - 1847" refId="ref8451" refString="Massin, C., Samyn, Y. &amp; Thandar, A. S. (2004) The genus Labidodemas (Holothuroidea: Aspidochirotida) revisited, with description of three new species and with re-positioning of Holothuria (Irenothuria) maccullochi Deichmann, 1958. Journal of Natural History, 38, 1811 - 1847. https: // doi. org / 10.1080 / 0022293031000156268" type="journal article" year="2004">
Massin
<emphasis id="B91D952CFFB23A1F9717FA63FD2730D4" box="[628,677,1475,1501]" italics="true" pageId="5" pageNumber="232">et al</emphasis>
. 2004
</bibRefCitation>
). The
<typeStatus id="54D2F79CFFB23A1F9657FA63FC1230D7" box="[820,912,1476,1502]" pageId="5" pageNumber="232" type="syntype">Syntype</typeStatus>
<collectionCode id="ED78D1FBFFB23A1F96F5FA63FC5630D7" box="[918,980,1476,1502]" country="Germany" httpUri="http://grbio.org/cool/gs4h-1fee" name="Zoologischen Museums Greifswald" pageId="5" pageNumber="232">ZMG</collectionCode>
(no number given by
<bibRefCitation id="EFF834CFFFB23A1F91A8FA64FAE930D4" author="Selenka, E." box="[1227,1387,1475,1501]" pageId="5" pageNumber="232" pagination="291 - 374" refId="ref8895" refString="Selenka, E. (1867) Beitrage zur anatomie und systematik der holothurien. Zeitschrift fur wissenschaftliche Zoologie, 17 (2), 291 - 374." type="journal article" year="1867">Selenka, 1867</bibRefCitation>
) are untraceable (
<bibRefCitation id="EFF834CFFFB23A1F9444FA40FE7E330B" author="Massin, C. &amp; Samyn, Y. &amp; Thandar, A. S." box="[295,508,1511,1538]" pageId="5" pageNumber="232" pagination="1811 - 1847" refId="ref8451" refString="Massin, C., Samyn, Y. &amp; Thandar, A. S. (2004) The genus Labidodemas (Holothuroidea: Aspidochirotida) revisited, with description of three new species and with re-positioning of Holothuria (Irenothuria) maccullochi Deichmann, 1958. Journal of Natural History, 38, 1811 - 1847. https: // doi. org / 10.1080 / 0022293031000156268" type="journal article" year="2004">
Massin
<emphasis id="B91D952CFFB23A1F94E3FA4FFE3B3308" box="[384,441,1511,1537]" italics="true" pageId="5" pageNumber="232">et al.</emphasis>
2004
</bibRefCitation>
).
</paragraph>
<paragraph id="8BD6493EFFB23A1F95A4F9ABFD163343" blockId="5.[151,1437,1439,2041]" pageId="5" pageNumber="232">
<emphasis id="B91D952CFFB23A1F95A4F9ABFEE1332C" bold="true" box="[199,355,1547,1574]" pageId="5" pageNumber="232">Type locality</emphasis>
.
<materialsCitation id="3B014363FFB23A1F9417F9ACFD123343" county="Society Islands" location="Formerly Hawaiian Islands" municipality="Sandwich Islands" pageId="5" pageNumber="232" specimenCount="1" typeStatus="lectotype">
<location id="8EB61FE5FFB23A1F9417F9ACFD29332C" LSID="urn:lsid:plazi:treatment:03C0F828FFB23A1295F4FB78FCB83308:8EB61FE5FFB23A1F9417F9ACFD29332C" box="[372,683,1547,1573]" county="Society Islands" municipality="Sandwich Islands" name="Formerly Hawaiian Islands" pageId="5" pageNumber="232">Formerly Hawaiian Islands</location>
(as
<collectingMunicipality id="6BB2D344FFB23A1F9783F9ACFC29332C" box="[736,939,1547,1573]" pageId="5" pageNumber="232">Sandwich Islands</collectingMunicipality>
), but now
<collectingCounty id="62B731B2FFB23A1F9149F9ACFB55332C" box="[1066,1239,1547,1573]" pageId="5" pageNumber="232">Society Islands</collectingCounty>
according to the designation of
<typeStatus id="54D2F79CFFB23A1F945DF988FE273340" box="[318,421,1583,1609]" pageId="5" pageNumber="232" type="lectotype">lectotype</typeStatus>
(
<bibRefCitation id="EFF834CFFFB23A1F94D0F988FD0A3343" author="Massin, C. &amp; Samyn, Y. &amp; Thandar, A. S." box="[435,648,1583,1610]" pageId="5" pageNumber="232" pagination="1811 - 1847" refId="ref8451" refString="Massin, C., Samyn, Y. &amp; Thandar, A. S. (2004) The genus Labidodemas (Holothuroidea: Aspidochirotida) revisited, with description of three new species and with re-positioning of Holothuria (Irenothuria) maccullochi Deichmann, 1958. Journal of Natural History, 38, 1811 - 1847. https: // doi. org / 10.1080 / 0022293031000156268" type="journal article" year="2004">
Massin
<emphasis id="B91D952CFFB23A1F976FF997FDBC3340" box="[524,574,1583,1609]" italics="true" pageId="5" pageNumber="232">et al</emphasis>
. 2004
</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BD6493EFFB23A1F95A4F9F4FBF1339B" blockId="5.[151,1437,1439,2041]" pageId="5" pageNumber="232">
<emphasis id="B91D952CFFB23A1F95A4F9F4FE2A3364" bold="true" box="[199,424,1619,1645]" pageId="5" pageNumber="232">Material examined</emphasis>
.
<materialsCitation id="3B014363FFB23A1F94D6F9F3FC703398" collectionCode="MZB" county="Batulawang" elevation="5" location="Tim Teripang" municipality="Situbondo" pageId="5" pageNumber="232" specimenCount="1" stateProvince="Jawa Timur">
<collectionCode id="ED78D1FBFFB23A1F94D6F9F3FE703367" box="[437,498,1620,1646]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="5" pageNumber="232" type="Museum">MZB</collectionCode>
Hol.385 (
<specimenCount id="9D6F82B7FFB23A1F9702F9F3FD7C3364" box="[609,766,1619,1646]" count="1" pageId="5" pageNumber="232" type="generic">one specimen</specimenCount>
), 68x
<quantity id="4C91E4DBFFB23A1F965FF9F4FC0A3367" box="[828,904,1619,1646]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="5" pageNumber="232" unit="mm" value="19.0">19 mm</quantity>
in ethanol, collected in
<collectingCounty id="62B731B2FFB23A1F91ECF9F4FA9A3364" box="[1167,1304,1619,1645]" pageId="5" pageNumber="232">Batulawang</collectingCounty>
,
<collectingMunicipality id="6BB2D344FFB23A1F9041F9F4FA1A3364" box="[1314,1432,1619,1645]" pageId="5" pageNumber="232">Situbondo</collectingMunicipality>
,
<collectingRegion id="49AD87DCFFB23A1F95F4F9DFFE9F3398" box="[151,285,1655,1682]" country="Indonesia" name="Jawa Timur" pageId="5" pageNumber="232">Jawa Timur</collectingRegion>
, by diving, found under a rock,
<elevation id="0044AE0DFFB23A1F97E1F9D0FD773398" box="[642,757,1655,1682]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="232" unit="m" value="5.0">
<quantity id="4C91E4DBFFB23A1F97E1F9D0FD2F339B" box="[642,685,1655,1682]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="232" unit="m" value="5.0">5 m</quantity>
depth
</elevation>
, coll. by
<location id="8EB61FE5FFB23A1F9638F9D0FC703398" LSID="urn:lsid:plazi:treatment:03C0F828FFB23A1295F4FB78FCB83308:8EB61FE5FFB23A1F9638F9D0FC703398" box="[859,1010,1655,1681]" county="Batulawang" municipality="Situbondo" name="Tim Teripang" pageId="5" pageNumber="232" stateProvince="Jawa Timur">Tim Teripang</location>
</materialsCitation>
<materialsCitation id="3B014363FFB23A1F969AF9DFFBED339B" box="[1017,1135,1656,1682]" collectionCode="LIPI" pageId="5" pageNumber="232" specimenCode="LIPI 2018" specimenCount="1" stateProvince="Jawa Timur">
<specimenCode id="DBCFE145FFB23A1F969AF9DFFBED339B" box="[1017,1135,1656,1682]" collectionCode="LIPI" pageId="5" pageNumber="232">LIPI 2018</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BD6493EFFB23A1F95A4F93BFAC7322B" blockId="5.[151,1437,1439,2041]" pageId="5" pageNumber="232">
<emphasis id="B91D952CFFB23A1F95A4F93BFE3933BC" bold="true" box="[199,443,1691,1718]" pageId="5" pageNumber="232">External description</emphasis>
. Body cylindrical; colour in ethanol white uniformly (
<figureCitation id="135255BBFFB23A1F916CF93CFB1233BF" box="[1039,1168,1691,1718]" captionStart="FIGURE 5" captionStartId="6.[151,250,923,947]" captionTargetBox="[257,1329,181,899]" captionTargetId="figure-25@6.[257,1329,178,899]" captionTargetPageId="6" captionText="FIGURE 5. Labidodemas semperianum Selenka, 1867. MZB Hol.385 (68x19 mm in ethanol). Specimen photograph.A: dorsal; B: ventral; C: calcareous ring." pageId="5" pageNumber="232">Figs. 5A, B</figureCitation>
). Body soft to the touch,
<quantity id="4C91E4DBFFB23A1F95F4F967FF7433D3" box="[151,246,1728,1754]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="5" pageNumber="232" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">12 mm</quantity>
thick. Mouth and anus terminal; mouth brown, surrounded by 16 potato-digitate tentacles. Anal papillae present, not calcified. Dorsal papillae small, conical shape, yellow, scattered over the surface. Ventral tube feet are very long (
<quantity id="4C91E4DBFFB23A1F9471F8AFFED5322B" box="[274,343,1800,1826]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="232" unit="mm" value="3.0">3 mm</quantity>
), white to yellowish, arranged in three rows in the ambulacral areas with a zigzag pattern.
</paragraph>
<paragraph id="8BD6493EFFB23A1F95A4F88BFD0132F0" blockId="5.[151,1437,1439,2041]" pageId="5" pageNumber="232">
<emphasis id="B91D952CFFB23A1F95A4F88BFE34324C" bold="true" box="[199,438,1835,1862]" pageId="5" pageNumber="232">Internal description</emphasis>
. Calcareous ring small (
<figureCitation id="135255BBFFB23A1F97A5F88CFC9F324F" box="[710,797,1835,1862]" captionStart="FIGURE 5" captionStartId="6.[151,250,923,947]" captionTargetBox="[257,1329,181,899]" captionTargetId="figure-25@6.[257,1329,178,899]" captionTargetPageId="6" captionText="FIGURE 5. Labidodemas semperianum Selenka, 1867. MZB Hol.385 (68x19 mm in ethanol). Specimen photograph.A: dorsal; B: ventral; C: calcareous ring." pageId="5" pageNumber="232">Fig. 5C</figureCitation>
), with radial pieces twice wider than interradial pieces (
<quantity id="4C91E4DBFFB23A1F90EEF88BFF413263" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="232" unit="mm" value="2.0">2 mm</quantity>
wide and
<quantity id="4C91E4DBFFB23A1F945AF8F7FEF93263" box="[313,379,1872,1898]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="232" unit="mm" value="2.0">2 mm</quantity>
high). Cuvierian tubules present, not expelled during collection, adhesive, bulbous and thick, brownish, branched, distal part of the tube with spherules, more than 10 attachment sites. Tentacle ampullae present, short. Single Polian vesicle, sac-like. Stone canal not observed. Gonad one tuft, branched. Respiratory trees arise from a single stalk. Longitudinal muscles flat and completely attached to the body wall. Inner body wall whitish, intestine filled with gravel and coral rubble.
</paragraph>
<caption id="DF1619B6FFB13A1C95F4FC3CFE4B36DE" pageId="6" pageNumber="233" startId="6.[151,250,923,947]" targetBox="[257,1329,181,899]" targetPageId="6" targetType="figure">
<paragraph id="8BD6493EFFB13A1C95F4FC3CFE4B36DE" blockId="6.[151,1436,923,983]" pageId="6" pageNumber="233">
<emphasis id="B91D952CFFB13A1C95F4FC3CFE9036BA" bold="true" box="[151,274,923,947]" pageId="6" pageNumber="233">FIGURE 5.</emphasis>
<taxonomicName id="4C6932BDFFB13A1C9474FC3CFD4736BA" authority="Selenka, 1867" authorityName="Selenka" authorityYear="1867" box="[279,709,923,947]" class="Holothuroidea" family="Holothuriidae" genus="Labidodemas" kingdom="Animalia" order="Holothuriida" pageId="6" pageNumber="233" phylum="Echinodermata" rank="species" species="semperianum">
<emphasis id="B91D952CFFB13A1C9474FC3CFDAF36BA" box="[279,557,923,947]" italics="true" pageId="6" pageNumber="233">Labidodemas semperianum</emphasis>
<bibRefCitation id="EFF834CFFFB13A1C9750FC3CFD4736BA" author="Selenka, E." box="[563,709,923,947]" pageId="6" pageNumber="233" pagination="291 - 374" refId="ref8895" refString="Selenka, E. (1867) Beitrage zur anatomie und systematik der holothurien. Zeitschrift fur wissenschaftliche Zoologie, 17 (2), 291 - 374." type="journal article" year="1867">Selenka, 1867</bibRefCitation>
</taxonomicName>
. MZB Hol.385 (68x19 mm in ethanol). Specimen photograph. A: dorsal; B: ventral; C: calcareous ring.
</paragraph>
</caption>
<caption id="DF1619B6FFB13A1C95F4F81FFD0A32FD" pageId="6" pageNumber="233" startId="6.[151,250,1976,2000]" targetBox="[264,1322,1007,1952]" targetPageId="6" targetType="figure">
<paragraph id="8BD6493EFFB13A1C95F4F81FFD0A32FD" blockId="6.[151,1437,1976,2036]" pageId="6" pageNumber="233">
<emphasis id="B91D952CFFB13A1C95F4F81FFE9132D9" bold="true" box="[151,275,1976,2000]" pageId="6" pageNumber="233">FIGURE 6.</emphasis>
<taxonomicName id="4C6932BDFFB13A1C947AF81FFD4832D9" authority="Selenka, 1867" authorityName="Selenka" authorityYear="1867" box="[281,714,1976,2000]" class="Holothuroidea" family="Holothuriidae" genus="Labidodemas" kingdom="Animalia" order="Holothuriida" pageId="6" pageNumber="233" phylum="Echinodermata" rank="species" species="semperianum">
<emphasis id="B91D952CFFB13A1C947AF81FFDB232D9" box="[281,560,1976,2000]" italics="true" pageId="6" pageNumber="233">Labidodemas semperianum</emphasis>
<bibRefCitation id="EFF834CFFFB13A1C9754F81FFD4832D9" author="Selenka, E." box="[567,714,1976,2000]" pageId="6" pageNumber="233" pagination="291 - 374" refId="ref8895" refString="Selenka, E. (1867) Beitrage zur anatomie und systematik der holothurien. Zeitschrift fur wissenschaftliche Zoologie, 17 (2), 291 - 374." type="journal article" year="1867">Selenka, 1867</bibRefCitation>
</taxonomicName>
. MZB Hol.385. Ossicles dorsal body wall, A: tables; B: pseudo-button. Ossicles ventral body wall, C: tables; D: buttons.
</paragraph>
</caption>
<caption id="DF1619B6FFB03A1D95F4FA66FC1430F4" pageId="7" pageNumber="234" startId="7.[151,250,1473,1497]" targetBox="[257,1329,181,1449]" targetPageId="7" targetType="figure">
<paragraph id="8BD6493EFFB03A1D95F4FA66FC1430F4" blockId="7.[151,1436,1473,1534]" pageId="7" pageNumber="234">
<emphasis id="B91D952CFFB03A1D95F4FA66FE9130D3" bold="true" box="[151,275,1473,1498]" pageId="7" pageNumber="234">FIGURE 7.</emphasis>
<taxonomicName id="4C6932BDFFB03A1D9479FA66FD4930D0" authority="Selenka, 1867" authorityName="Selenka" authorityYear="1867" box="[282,715,1473,1497]" class="Holothuroidea" family="Holothuriidae" genus="Labidodemas" kingdom="Animalia" order="Holothuriida" pageId="7" pageNumber="234" phylum="Echinodermata" rank="species" species="semperianum">
<emphasis id="B91D952CFFB03A1D9479FA66FDB330D0" box="[282,561,1473,1497]" italics="true" pageId="7" pageNumber="234">Labidodemas semperianum</emphasis>
<bibRefCitation id="EFF834CFFFB03A1D975BFA66FD4930D0" author="Selenka, E." box="[568,715,1473,1497]" pageId="7" pageNumber="234" pagination="291 - 374" refId="ref8895" refString="Selenka, E. (1867) Beitrage zur anatomie und systematik der holothurien. Zeitschrift fur wissenschaftliche Zoologie, 17 (2), 291 - 374." type="journal article" year="1867">Selenka, 1867</bibRefCitation>
</taxonomicName>
. MZB Hol.385. Ossicles dorsal papillae, A: tables; B: buttons, pseudobuttons and rods; C: plate. Ossicles ventral tube feet, D: tables; E: endplate.
</paragraph>
</caption>
<paragraph id="8BD6493EFFB03A1D95A4F9E8FA9233DF" blockId="7.[151,1437,1615,2038]" pageId="7" pageNumber="234">
<emphasis id="B91D952CFFB03A1D95A4F9E8FEA73360" bold="true" box="[199,293,1615,1641]" pageId="7" pageNumber="234">Ossicles</emphasis>
. Dorsal body wall with tables and buttons; tables disc 6080 µm across, disc rims mostly spinous, perforated by 46 central holes; table spire low (2030 µm), ending in four to six long spines, some of them bifurcated distally; table crown generally larger than table disc (
<figureCitation id="135255BBFFB03A1D9780F93FFCBF33BB" box="[739,829,1688,1714]" captionStart="FIGURE 6" captionStartId="6.[151,250,1976,2000]" captionTargetBox="[264,1322,1007,1952]" captionTargetId="figure-49@6.[264,1322,1004,1954]" captionTargetPageId="6" captionText="FIGURE 6. Labidodemas semperianum Selenka, 1867. MZB Hol.385. Ossicles dorsal body wall, A: tables; B: pseudo-button. Ossicles ventral body wall, C: tables; D: buttons." pageId="7" pageNumber="234">Fig. 6A</figureCitation>
). Pseudo-buttons only a few; 6080 µm long; knobbed (
<figureCitation id="135255BBFFB03A1D95FCF91BFF7533DF" box="[159,247,1724,1750]" captionStart="FIGURE 6" captionStartId="6.[151,250,1976,2000]" captionTargetBox="[264,1322,1007,1952]" captionTargetId="figure-49@6.[264,1322,1004,1954]" captionTargetPageId="6" captionText="FIGURE 6. Labidodemas semperianum Selenka, 1867. MZB Hol.385. Ossicles dorsal body wall, A: tables; B: pseudo-button. Ossicles ventral body wall, C: tables; D: buttons." pageId="7" pageNumber="234">Fig. 6B</figureCitation>
). Ventral body wall holds tables and buttons (
<figureCitation id="135255BBFFB03A1D9792F91BFCF033DF" box="[753,882,1724,1750]" captionStart="FIGURE 6" captionStartId="6.[151,250,1976,2000]" captionTargetBox="[264,1322,1007,1952]" captionTargetId="figure-49@6.[264,1322,1004,1954]" captionTargetPageId="6" captionText="FIGURE 6. Labidodemas semperianum Selenka, 1867. MZB Hol.385. Ossicles dorsal body wall, A: tables; B: pseudo-button. Ossicles ventral body wall, C: tables; D: buttons." pageId="7" pageNumber="234">Figs. 6C, D</figureCitation>
) similar to those in dorsal body wall.
</paragraph>
<paragraph id="8BD6493EFFB03A1D95A4F947FF4532FF" blockId="7.[151,1437,1615,2038]" pageId="7" pageNumber="234">
Dorsal papillae with tables, buttons, pseudo-buttons, rods and plates; tables disc 5590 µm across, disc rims spinous; perforated by four to five central holes and a variable number of peripheral holes; spire low, single crossbeam, ending in four to six long spines with distal bifurcation; table crown wider than table disc (
<figureCitation id="135255BBFFB03A1D91B4F88FFAB3324B" box="[1239,1329,1832,1858]" captionStart="FIGURE 7" captionStartId="7.[151,250,1473,1497]" captionTargetBox="[257,1329,181,1449]" captionTargetId="figure-17@7.[257,1329,181,1449]" captionTargetPageId="7" captionText="FIGURE 7. Labidodemas semperianum Selenka, 1867. MZB Hol.385. Ossicles dorsal papillae, A: tables; B: buttons, pseudobuttons and rods; C: plate. Ossicles ventral tube feet, D: tables; E: endplate." pageId="7" pageNumber="234">Fig. 7A</figureCitation>
). Buttons 4050 µm long, sometimes with knobbed; pseudo-buttons 60100 µm long; rods 5090 µm long (
<figureCitation id="135255BBFFB03A1D9192F8EBFACB326F" box="[1265,1353,1868,1894]" captionStart="FIGURE 7" captionStartId="7.[151,250,1473,1497]" captionTargetBox="[257,1329,181,1449]" captionTargetId="figure-17@7.[257,1329,181,1449]" captionTargetPageId="7" captionText="FIGURE 7. Labidodemas semperianum Selenka, 1867. MZB Hol.385. Ossicles dorsal papillae, A: tables; B: buttons, pseudobuttons and rods; C: plate. Ossicles ventral tube feet, D: tables; E: endplate." pageId="7" pageNumber="234">Fig. 7B</figureCitation>
); small plate observed in heaps with rods and buttons, 100 µm across (
<figureCitation id="135255BBFFB03A1D9630F8D7FC293283" box="[851,939,1904,1930]" captionStart="FIGURE 7" captionStartId="7.[151,250,1473,1497]" captionTargetBox="[257,1329,181,1449]" captionTargetId="figure-17@7.[257,1329,181,1449]" captionTargetPageId="7" captionText="FIGURE 7. Labidodemas semperianum Selenka, 1867. MZB Hol.385. Ossicles dorsal papillae, A: tables; B: buttons, pseudobuttons and rods; C: plate. Ossicles ventral tube feet, D: tables; E: endplate." pageId="7" pageNumber="234">Fig. 7C</figureCitation>
). Ventral tube feet only with tables and large end plates. Tables 6080 µm across, with four to five central holes, crown sometimes with only three spines, tables reduced to the (spiny) disc or with a reduced crown also observed (
<figureCitation id="135255BBFFB03A1D96E3F81FFC5B32DB" box="[896,985,1976,2002]" captionStart="FIGURE 7" captionStartId="7.[151,250,1473,1497]" captionTargetBox="[257,1329,181,1449]" captionTargetId="figure-17@7.[257,1329,181,1449]" captionTargetPageId="7" captionText="FIGURE 7. Labidodemas semperianum Selenka, 1867. MZB Hol.385. Ossicles dorsal papillae, A: tables; B: buttons, pseudobuttons and rods; C: plate. Ossicles ventral tube feet, D: tables; E: endplate." pageId="7" pageNumber="234">Fig. 7D</figureCitation>
). Endplate reaching 500 µm across (
<figureCitation id="135255BBFFB03A1D900CF81FFF3832FF" captionStart="FIGURE 7" captionStartId="7.[151,250,1473,1497]" captionTargetBox="[257,1329,181,1449]" captionTargetId="figure-17@7.[257,1329,181,1449]" captionTargetPageId="7" captionText="FIGURE 7. Labidodemas semperianum Selenka, 1867. MZB Hol.385. Ossicles dorsal papillae, A: tables; B: buttons, pseudobuttons and rods; C: plate. Ossicles ventral tube feet, D: tables; E: endplate." pageId="7" pageNumber="234">Fig. 7E</figureCitation>
).
</paragraph>
<caption id="DF1619B6FFBF3A1295F4FBE1FAB13157" box="[151,1331,1094,1118]" pageId="8" pageNumber="235" startId="8.[151,250,1094,1118]" targetBox="[265,1322,181,1069]" targetPageId="8" targetType="figure">
<paragraph id="8BD6493EFFBF3A1295F4FBE1FAB13157" blockId="8.[151,1331,1094,1118]" box="[151,1331,1094,1118]" pageId="8" pageNumber="235">
<emphasis id="B91D952CFFBF3A1295F4FBE1FE913157" bold="true" box="[151,275,1094,1118]" pageId="8" pageNumber="235">FIGURE 8.</emphasis>
<taxonomicName id="4C6932BDFFBF3A12947AFBE1FD483157" authority="Selenka, 1867" authorityName="Selenka" authorityYear="1867" box="[281,714,1094,1118]" class="Holothuroidea" family="Holothuriidae" genus="Labidodemas" kingdom="Animalia" order="Holothuriida" pageId="8" pageNumber="235" phylum="Echinodermata" rank="species" species="semperianum">
<emphasis id="B91D952CFFBF3A12947AFBE1FDB23157" box="[281,560,1094,1118]" italics="true" pageId="8" pageNumber="235">Labidodemas semperianum</emphasis>
<bibRefCitation id="EFF834CFFFBF3A129754FBE1FD483157" author="Selenka, E." box="[567,714,1094,1118]" pageId="8" pageNumber="235" pagination="291 - 374" refId="ref8895" refString="Selenka, E. (1867) Beitrage zur anatomie und systematik der holothurien. Zeitschrift fur wissenschaftliche Zoologie, 17 (2), 291 - 374." type="journal article" year="1867">Selenka, 1867</bibRefCitation>
</taxonomicName>
. MZB Hol.385. A: tentacle rods; B: ossicles of anal papillae.
</paragraph>
</caption>
<paragraph id="8BD6493EFFBF3A1295A4FB04FA9131E8" blockId="8.[151,1437,1187,1537]" pageId="8" pageNumber="235">
Tentacles with only rods, thin, smooth or spiny, 3060 µm long (
<figureCitation id="135255BBFFBF3A1296FAFB04FC7131B4" box="[921,1011,1187,1213]" captionStart="FIGURE 8" captionStartId="8.[151,250,1094,1118]" captionTargetBox="[265,1322,181,1069]" captionTargetId="figure-25@8.[265,1322,181,1069]" captionTargetPageId="8" captionText="FIGURE 8. Labidodemas semperianum Selenka, 1867. MZB Hol.385. A: tentacle rods; B: ossicles of anal papillae." pageId="8" pageNumber="235">Fig. 8A</figureCitation>
). Ossicles of anal papillae consist of a mixture of table discs, 4050 µm across, irregular rods and reduced buttons, 3045 µm long (
<figureCitation id="135255BBFFBF3A1291CDFB60FA8431E8" box="[1198,1286,1223,1249]" captionStart="FIGURE 8" captionStartId="8.[151,250,1094,1118]" captionTargetBox="[265,1322,181,1069]" captionTargetId="figure-25@8.[265,1322,181,1069]" captionTargetPageId="8" captionText="FIGURE 8. Labidodemas semperianum Selenka, 1867. MZB Hol.385. A: tentacle rods; B: ossicles of anal papillae." pageId="8" pageNumber="235">Fig. 8B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3731AB5FFBF3A1295A4FB4CFEAA3078" pageId="8" pageNumber="235" type="discussion">
<paragraph id="8BD6493EFFBF3A1295A4FB4CFEAA3078" blockId="8.[151,1437,1187,1537]" pageId="8" pageNumber="235">
<emphasis id="B91D952CFFBF3A1295A4FB4CFEB6300C" bold="true" box="[199,308,1259,1285]" pageId="8" pageNumber="235">Remarks</emphasis>
. The specimen examined in the present study possesses Cuvierian tubules with more than 10 attachment sites. Such had never been observed before in this species. On the other hand,
<taxonomicName id="4C6932BDFFBF3A1291FEFAB7FAD13020" authorityName="Selenka" authorityYear="1867" box="[1181,1363,1296,1321]" class="Holothuroidea" family="Holothuriidae" genus="Labidodemas" kingdom="Animalia" order="Holothuriida" pageId="8" pageNumber="235" phylum="Echinodermata" rank="species" species="semperianum">
<emphasis id="B91D952CFFBF3A1291FEFAB7FAD13020" box="[1181,1363,1296,1321]" italics="true" pageId="8" pageNumber="235">L. semperianum</emphasis>
</taxonomicName>
in the present study has few buttons and pseudo-buttons in the body wall and dorsal papillae, similar to the study of
<bibRefCitation id="EFF834CFFFBF3A129028FA94FEA63078" author="Massin, C. &amp; Samyn, Y. &amp; Thandar, A. S." pageId="8" pageNumber="235" pagination="1811 - 1847" refId="ref8451" refString="Massin, C., Samyn, Y. &amp; Thandar, A. S. (2004) The genus Labidodemas (Holothuroidea: Aspidochirotida) revisited, with description of three new species and with re-positioning of Holothuria (Irenothuria) maccullochi Deichmann, 1958. Journal of Natural History, 38, 1811 - 1847. https: // doi. org / 10.1080 / 0022293031000156268" type="journal article" year="2004">
Massin
<emphasis id="B91D952CFFBF3A1295F4FAFFFF4A3078" box="[151,200,1367,1393]" italics="true" pageId="8" pageNumber="235">et al</emphasis>
. (2004)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3731AB5FFBF3A1295A4FADCFCB83308" pageId="8" pageNumber="235" type="biology_ecology">
<paragraph id="8BD6493EFFBF3A1295A4FADCFB60309C" blockId="8.[151,1437,1187,1537]" box="[199,1250,1403,1429]" pageId="8" pageNumber="235">
<emphasis id="B91D952CFFBF3A1295A4FADCFEA4309C" bold="true" box="[199,294,1403,1429]" pageId="8" pageNumber="235">Ecology</emphasis>
. Found in shallow waters, less than
<quantity id="4C91E4DBFFBF3A1297DEFADCFD74309C" box="[701,758,1403,1429]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="235" unit="m" value="30.0">30 m</quantity>
depth (
<bibRefCitation id="EFF834CFFFBF3A12962AFADCFB87309C" author="Lane, D. J. W. &amp; Marsh, L. M. &amp; VandenSpiegel, D. &amp; Rowe, F. W. E." box="[841,1029,1403,1429]" pageId="8" pageNumber="235" pagination="459 - 493" refId="ref8229" refString="Lane, D. J. W., Marsh, L. M., VandenSpiegel, D. &amp; Rowe, F. W. E. (2000) Echinoderm Fauna of the South China Sea: an Inventory and Analysis of Distribution Patterns. Raffles Bulletin of Zoology, 48, 459 - 493." type="journal article" year="2000">
Lane
<emphasis id="B91D952CFFBF3A1296E9FADBFC39309C" box="[906,955,1403,1429]" italics="true" pageId="8" pageNumber="235">et al</emphasis>
. 2000
</bibRefCitation>
), under coral rocks.
</paragraph>
<paragraph id="8BD6493EFFBF3A1295A4FA38FD8930B0" blockId="8.[151,1437,1187,1537]" box="[199,523,1439,1465]" pageId="8" pageNumber="235">
<emphasis id="B91D952CFFBF3A1295A4FA38FED930B0" bold="true" box="[199,347,1439,1465]" pageId="8" pageNumber="235">Depth range</emphasis>
.
<quantity id="4C91E4DBFFBF3A129409FA38FE3D30B0" box="[362,447,1439,1465]" metricMagnitude="1" metricUnit="m" metricValue="1.55" metricValueMax="3.0" metricValueMin="0.1" pageId="8" pageNumber="235" unit="m" value="15.5" valueMax="30.0" valueMin="1.0">130 m</quantity>
depth.
</paragraph>
<paragraph id="8BD6493EFFBF3A1295A4FA64FCB83308" blockId="8.[151,1437,1187,1537]" pageId="8" pageNumber="235">
<emphasis id="B91D952CFFBF3A1295A4FA64FE6830D4" bold="true" box="[199,490,1475,1501]" pageId="8" pageNumber="235">Geographic distribution</emphasis>
. The following new locality is added here since
<bibRefCitation id="EFF834CFFFBF3A129174FA64FAC230D7" author="Massin, C. &amp; Samyn, Y. &amp; Thandar, A. S." box="[1047,1344,1475,1502]" pageId="8" pageNumber="235" pagination="1811 - 1847" refId="ref8451" refString="Massin, C., Samyn, Y. &amp; Thandar, A. S. (2004) The genus Labidodemas (Holothuroidea: Aspidochirotida) revisited, with description of three new species and with re-positioning of Holothuria (Irenothuria) maccullochi Deichmann, 1958. Journal of Natural History, 38, 1811 - 1847. https: // doi. org / 10.1080 / 0022293031000156268" type="journal article" year="2004">
Massin
<emphasis id="B91D952CFFBF3A129112FA63FB2630D4" box="[1137,1188,1475,1501]" italics="true" pageId="8" pageNumber="235">et al</emphasis>
. (2004: 1835
</bibRefCitation>
, map 6) publication: Situbondo-East
<collectingRegion id="49AD87DCFFBF3A1294BBFA40FD8E3308" box="[472,524,1511,1537]" country="Indonesia" name="Jawa Timur" pageId="8" pageNumber="235">Java</collectingRegion>
,
<collectingCountry id="F37E09AEFFBF3A129774FA40FD063308" box="[535,644,1511,1537]" name="Indonesia" pageId="8" pageNumber="235">Indonesia</collectingCountry>
(present study).
</paragraph>
</subSubSection>
</treatment>
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<paragraph id="8BD6493EFFB53A1895F4FE80FDF23448" blockId="2.[151,624,295,357]" box="[151,624,295,322]" pageId="2" pageNumber="229">
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<taxonomicName id="4C6932BDFFB53A1895F4FE80FDF23448" ID-CoL="3RJJK" authority="(Ludwig, 1875)" authorityName="Kudingareng-Sulawesi" baseAuthorityName="Ludwig" baseAuthorityYear="1875" box="[151,624,295,322]" class="Holothuroidea" family="Holothuriidae" genus="Labidodemas" kingdom="Animalia" order="Holothuriida" pageId="2" pageNumber="229" phylum="Echinodermata" rank="species" species="rugosum">
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<emphasis id="B91D952CFFB53A1895F4FE80FE2E3448" bold="true" box="[151,428,295,322]" italics="true" pageId="2" pageNumber="229">Labidodemas rugosum</emphasis>
(
<bibRefCitation id="EFF834CFFFB53A1894D8FE80FDE53448" author="Ludwig, H." box="[443,615,295,321]" pageId="2" pageNumber="229" pagination="77 - 118" refId="ref8343" refString="Ludwig, H. (1875) Beitrage zur Kenntniss der Holothurien. Arbeiten aus dem Zoologisch-Zootomischen Institut in Wurzburg, 2, 77 - 118." type="journal article" year="1875">Ludwig, 1875</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3731AB5FFB53A1895F4FEECFEAD346C" box="[151,303,331,357]" pageId="2" pageNumber="229" type="description">
<paragraph id="8BD6493EFFB53A1895F4FEECFEAD346C" blockId="2.[151,624,295,357]" box="[151,303,331,357]" pageId="2" pageNumber="229">
(
<figureCitation id="135255BBFFB53A1895FCFEECFEA5346C" box="[159,295,331,357]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="2.[151,250,1752,1776]" captionStartId-1="3.[151,250,1564,1588]" captionStartId-2="4.[151,250,1696,1720]" captionTargetBox-0="[257,1329,889,1728]" captionTargetBox-1="[257,1329,181,1540]" captionTargetBox-2="[257,1329,181,1672]" captionTargetId-0="figure-190@2.[257,1329,889,1728]" captionTargetId-1="figure-17@3.[257,1329,181,1540]" captionTargetId-2="figure-25@4.[257,1329,181,1672]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 2. Labidodemas rugosum (Ludwig, 1875). Live specimen photograph, MZB Hol.427 (70x15 mm in ethanol). A: dorsal; B. ventral; C: illustration of calcareous ring." captionText-1="FIGURE 3. Labidodemas rugosum (Ludwig, 1875). A, C, D, and E, MZB Hol.427; B, MZB Hol.390. Dorsal body wall, A: tables; B: buttons. Ventral body wall, C: tables; D: buttons. E: tentacle rods." captionText-2="FIGURE 4. Labidodemas rugosum (Ludwig, 1875). MZB Hol.427. Dorsal papillae, A: rod-like buttons; B: rods; C: buttons; D: tables. Ventral tube feet, E: tables; F: buttons; G: buttons-plate; H: plates." pageId="2" pageNumber="229">Figures 24</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3731AB5FFB53A1895F4FE37FD3434CF" pageId="2" pageNumber="229" type="reference_group">
<paragraph id="8BD6493EFFB53A1895F4FE37FC1E34A1" blockId="2.[151,1437,399,577]" box="[151,924,399,424]" pageId="2" pageNumber="229">
<treatmentCitationGroup id="AB796E10FFB53A1895F4FE37FC1E34A1" box="[151,924,399,424]" pageId="2" pageNumber="229">
<treatmentCitation id="0AC86F2FFFB53A1895F4FE37FD9F34A1" author="Ludwig, H." box="[151,541,399,424]" page="110" pageId="2" pageNumber="229" year="1875">
<taxonomicName id="4C6932BDFFB53A1895F4FE37FD9F34A1" ID-CoL="3MB4Y" authority="Ludwig, 1875: 110" authorityName="Ludwig" authorityPageNumber="110" authorityYear="1875" box="[151,541,399,424]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Holothuriida" pageId="2" pageNumber="229" phylum="Echinodermata" rank="species" species="rugosa">
<emphasis id="B91D952CFFB53A1895F4FE37FED634AE" box="[151,340,400,424]" italics="true" pageId="2" pageNumber="229">Holothuria rugosa</emphasis>
<bibRefCitation id="EFF834CFFFB53A189438FE37FD9F34A1" author="Ludwig, H." box="[347,541,399,424]" pageId="2" pageNumber="229" pagination="77 - 118" refId="ref8343" refString="Ludwig, H. (1875) Beitrage zur Kenntniss der Holothurien. Arbeiten aus dem Zoologisch-Zootomischen Institut in Wurzburg, 2, 77 - 118." type="journal article" year="1875">Ludwig, 1875: 110</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. 7, figs. 33ac;
<treatmentCitation id="0AC86F2FFFB53A1897BDFE37FC1534A1" author="Pearson, J." box="[734,919,400,424]" page="82" pageId="2" pageNumber="229" year="1913">
<bibRefCitation id="EFF834CFFFB53A1897BDFE37FC1534A1" author="Pearson, J." box="[734,919,400,424]" pageId="2" pageNumber="229" pagination="49 - 101" refId="ref8553" refString="Pearson, J. (1913) Notes on the Holothurioidea of the Indian Ocean. Spolia Zeylandica, 9, 49 - 101. https: // doi. org / 10.5962 / bhl. part. 7317" type="journal article" year="1913">Pearson, 1913: 82</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD6493EFFB53A1895F4FE09FD3434CF" blockId="2.[151,1437,399,577]" box="[151,694,430,454]" pageId="2" pageNumber="229">
<taxonomicName id="4C6932BDFFB53A1895F4FE09FEDF34CF" ID-CoL="3MB5W" authorityName="Sluiter" authorityYear="1901" box="[151,349,430,454]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Holothuriida" pageId="2" pageNumber="229" phylum="Echinodermata" rank="species" species="triremis">
<emphasis id="B91D952CFFB53A1895F4FE09FEDF34CF" box="[151,349,430,454]" italics="true" pageId="2" pageNumber="229">Holothuria triremis</emphasis>
</taxonomicName>
; Sluiter, 1901: 19, Taf. VI,
<figureCitation id="135255BBFFB53A189716FE09FD3034CF" box="[629,690,430,454]" captionStart="FIGURE 3" captionStartId="3.[151,250,1564,1588]" captionTargetBox="[257,1329,181,1540]" captionTargetId="figure-17@3.[257,1329,181,1540]" captionTargetPageId="3" captionText="FIGURE 3. Labidodemas rugosum (Ludwig, 1875). A, C, D, and E, MZB Hol.427; B, MZB Hol.390. Dorsal body wall, A: tables; B: buttons. Ventral body wall, C: tables; D: buttons. E: tentacle rods." pageId="2" pageNumber="229">Fig. 3</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3731AB5FFB53A1895F4FE6AFD7934EC" box="[151,763,461,485]" pageId="2" pageNumber="229" type="description">
<paragraph id="8BD6493EFFB53A1895F4FE6AFD7934EC" blockId="2.[151,1437,399,577]" box="[151,763,461,485]" pageId="2" pageNumber="229">
<taxonomicName id="4C6932BDFFB53A1895F4FE6AFE5E34EC" authorityName="Ludwig" authorityYear="1875" box="[151,476,461,485]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Holothuriida" pageId="2" pageNumber="229" phylum="Echinodermata" rank="species" species="rugosa" subGenus="Holothuria">
<emphasis id="B91D952CFFB53A1895F4FE6AFE8A34EC" box="[151,264,461,485]" italics="true" pageId="2" pageNumber="229">Holothuria</emphasis>
(
<emphasis id="B91D952CFFB53A189474FE6AFE0A34EC" box="[279,392,461,485]" italics="true" pageId="2" pageNumber="229">Holothuria</emphasis>
)
<emphasis id="B91D952CFFB53A1894F5FE69FE5E34EC" box="[406,476,462,485]" italics="true" pageId="2" pageNumber="229">rugosa</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFF834CFFFB53A189489FE6AFDFE34EC" author="Panning, A." box="[490,636,461,485]" pageId="2" pageNumber="229" pagination="65 - 84" refId="ref8520" refString="Panning, A. (1934 b) Die Gattung Holothuria. 3. Teil. Mitteilungen aus dem Zoologichen Staatsinstitur und Zoologischen Museum in Hamburg, 1934, 65 - 84." type="journal article" year="1934">Panning, 1934</bibRefCitation>
(teil. 3): 75.
</paragraph>
</subSubSection>
<subSubSection id="C3731AB5FFB53A1895F4FE4BFE823748" pageId="2" pageNumber="229" type="reference_group">
<paragraph id="8BD6493EFFB53A1895F4FE4BFD03370A" blockId="2.[151,1437,399,577]" box="[151,641,491,516]" pageId="2" pageNumber="229">
<treatmentCitationGroup id="AB796E10FFB53A1895F4FE4BFD03370A" box="[151,641,491,516]" pageId="2" pageNumber="229">
<taxonomicName id="4C6932BDFFB53A1895F4FE4BFEEE370A" ID-CoL="3MB4X" baseAuthorityName="Ludwig" baseAuthorityYear="1875" box="[151,364,492,516]" class="Holothuroidea" family="Holothuriidae" genus="Labidodemas" kingdom="Animalia" order="Holothuriida" pageId="2" pageNumber="229" phylum="Echinodermata" rank="species" species="rugosa">
<emphasis id="B91D952CFFB53A1895F4FE4BFEEE370A" box="[151,364,492,516]" italics="true" pageId="2" pageNumber="229">Labidodemas rugosa</emphasis>
</taxonomicName>
;
<treatmentCitation id="0AC86F2FFFB53A189419FE4BFDFF370A" author="Jangoux, M. &amp; De Ridder, C. &amp; Massin, C. &amp; Darsono, P." box="[378,637,491,516]" page="163" pageId="2" pageNumber="229" year="1989">
<bibRefCitation id="EFF834CFFFB53A189419FE4BFDFF370A" author="Jangoux, M. &amp; De Ridder, C. &amp; Massin, C. &amp; Darsono, P." box="[378,637,491,516]" pageId="2" pageNumber="229" pagination="161 - 170" refId="ref8131" refString="Jangoux, M., De Ridder, C., Massin, C. &amp; Darsono, P. (1989) The holothuroids, echinoids and asteroids (echinodermata) collected by the Snellius-II expedition. Netherlands Journal of Sea Research, 23, 161 - 170. https: // doi. org / 10.1016 / 0077 - 7579 (89) 90010 - 0" type="journal article" year="1989">
Jangoux
<emphasis id="B91D952CFFB53A1894B6FE4BFD8A370D" box="[469,520,492,516]" italics="true" pageId="2" pageNumber="229">et al.</emphasis>
, 1989: 163
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD6493EFFB53A1895F4FDADFE823748" blockId="2.[151,1437,399,577]" pageId="2" pageNumber="229">
<treatmentCitationGroup id="AB796E10FFB53A1895F4FDADFE823748" pageId="2" pageNumber="229">
<taxonomicName id="4C6932BDFFB53A1895F4FDADFEFD372B" ID-CoL="3RJJK" authorityName="Kudingareng-Sulawesi" baseAuthorityName="Ludwig" baseAuthorityYear="1875" box="[151,383,522,546]" class="Holothuroidea" family="Holothuriidae" genus="Labidodemas" kingdom="Animalia" order="Holothuriida" pageId="2" pageNumber="229" phylum="Echinodermata" rank="species" species="rugosum">
<emphasis id="B91D952CFFB53A1895F4FDADFEFD372B" box="[151,383,522,546]" italics="true" pageId="2" pageNumber="229">Labidodemas rugosum</emphasis>
</taxonomicName>
;
<treatmentCitation id="0AC86F2FFFB53A1894EFFDADFD0D372B" author="Massin, C. &amp; Samyn, Y. &amp; Thandar, A. S." box="[396,655,522,546]" page="1830" pageId="2" pageNumber="229" year="2004">
<bibRefCitation id="EFF834CFFFB53A1894EFFDADFD0D372B" author="Massin, C. &amp; Samyn, Y. &amp; Thandar, A. S." box="[396,655,522,546]" pageId="2" pageNumber="229" pagination="1811 - 1847" refId="ref8451" refString="Massin, C., Samyn, Y. &amp; Thandar, A. S. (2004) The genus Labidodemas (Holothuroidea: Aspidochirotida) revisited, with description of three new species and with re-positioning of Holothuria (Irenothuria) maccullochi Deichmann, 1958. Journal of Natural History, 38, 1811 - 1847. https: // doi. org / 10.1080 / 0022293031000156268" type="journal article" year="2004">
Massin
<emphasis id="B91D952CFFB53A1894B8FDACFD8C372B" box="[475,526,522,546]" italics="true" pageId="2" pageNumber="229">et al.</emphasis>
, 2004: 1830
</bibRefCitation>
</treatmentCitation>
, figs. 8AL, map 5 (records before 2004);
<treatmentCitation id="0AC86F2FFFB53A189123FDADFAE7372B" author="Purwati, P. &amp; Wirawati, I." box="[1088,1381,522,546]" page="7" pageId="2" pageNumber="229" year="2011">
<bibRefCitation id="EFF834CFFFB53A189123FDADFAE7372B" author="Purwati, P. &amp; Wirawati, I." box="[1088,1381,522,546]" pageId="2" pageNumber="229" pagination="1 - 10" refId="ref8592" refString="Purwati, P. &amp; Wirawati, I. (2011) Holothuriidae (Echinodermata, Holothuroidea, Aspidochirotida) Perairan Dangkal Lombok Barat Bagian II. Genus Actinopyga, Bohadschia, Pearsonothuria, Labidodemas. Jurnal Oseanologi, 3 (1 / 2), 1 - 10." type="journal article" year="2011">Purwati &amp; Wirawati, 2011: 7</bibRefCitation>
</treatmentCitation>
, figs. 6ab.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3731AB5FFB53A1F95F4FDC8FC4434C6" lastPageId="5" lastPageNumber="232" pageId="2" pageNumber="229" type="materials_examined">
<paragraph id="8BD6493EFFB53A1895F4FDC8FDDA3783" blockId="2.[151,1437,623,865]" box="[151,600,623,650]" pageId="2" pageNumber="229">
<emphasis id="B91D952CFFB53A1895F4FDC8FEBC3780" bold="true" box="[151,318,623,649]" pageId="2" pageNumber="229">Type material</emphasis>
.
<materialsCitation id="3B014363FFB53A18942FFDC8FDDA3783" box="[332,600,623,650]" collectionCode="ZMH" pageId="2" pageNumber="229" specimenCount="1" typeStatus="holotype">
<typeStatus id="54D2F79CFFB53A18942FFDC8FE363780" box="[332,436,623,649]" pageId="2" pageNumber="229" type="holotype">Holotype</typeStatus>
<collectionCode id="ED78D1FBFFB53A1894D8FDD7FE7B3783" box="[443,505,624,650]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="2" pageNumber="229" type="Museum">ZMH</collectionCode>
E.2625.
</materialsCitation>
</paragraph>
<paragraph id="8BD6493EFFB53A1895A4FD34FDFA37A7" blockId="2.[151,1437,623,865]" box="[199,632,659,686]" pageId="2" pageNumber="229">
<materialsCitation id="3B014363FFB53A1895A4FD34FDF637A7" box="[199,628,659,686]" country="Samoa" location="Samoa" pageId="2" pageNumber="229" specimenCount="1" typeStatus="holotype">
<emphasis id="B91D952CFFB53A1895A4FD34FEE237A4" bold="true" box="[199,352,659,685]" pageId="2" pageNumber="229">
<typeStatus id="54D2F79CFFB53A1895A4FD34FE8037A4" box="[199,258,659,685]" pageId="2" pageNumber="229">Type</typeStatus>
locality
</emphasis>
.
<collectingCountry id="F37E09AEFFB53A18940DFD34FE3837A4" box="[366,442,659,685]" name="Samoa" pageId="2" pageNumber="229">Samoa</collectingCountry>
(
<bibRefCitation id="EFF834CFFFB53A1894AAFD34FDEE37A7" author="Ludwig, H." box="[457,620,659,686]" pageId="2" pageNumber="229" pagination="77 - 118" refId="ref8343" refString="Ludwig, H. (1875) Beitrage zur Kenntniss der Holothurien. Arbeiten aus dem Zoologisch-Zootomischen Institut in Wurzburg, 2, 77 - 118." type="journal article" year="1875">Ludwig, 1875</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BD6493EFFB53A1895A4FD10FE9B3668" blockId="2.[151,1437,623,865]" pageId="2" pageNumber="229">
<emphasis id="B91D952CFFB53A1895A4FD10FE2F37D8" bold="true" box="[199,429,695,721]" pageId="2" pageNumber="229">Material examined</emphasis>
.
<materialsCitation id="3B014363FFB53A1894DDFD10FD4C37DB" box="[446,718,695,722]" collectionCode="MZB" pageId="2" pageNumber="229" specimenCount="3">
<specimenCount id="9D6F82B7FFB53A1894DDFD10FD0037D8" box="[446,642,695,721]" count="3" pageId="2" pageNumber="229" type="generic">Three specimens</specimenCount>
.
<collectionCode id="ED78D1FBFFB53A1897F2FD1FFD4C37DB" box="[657,718,696,722]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="2" pageNumber="229" type="Museum">MZB</collectionCode>
</materialsCitation>
<materialsCitation id="3B014363FFB53A1897BBFD10FB2437DB" box="[728,1190,695,722]" collectionCode="MZB" location="Hol." pageId="2" pageNumber="229" specimenCount="1">
<location id="8EB61FE5FFB53A1897BBFD10FC8A37D8" LSID="urn:lsid:plazi:treatment:03C0F828FFB53A1F95F4FE80FD6C3192:8EB61FE5FFB53A1897BBFD10FC8A37D8" box="[728,776,695,721]" name="Hol." pageId="2" pageNumber="229">Hol.</location>
390 (110x
<quantity id="4C91E4DBFFB53A18961DFD10FC5337DB" box="[894,977,695,722]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="2" pageNumber="229" unit="mm" value="55.0">55 mm</quantity>
in ethanol),
<collectionCode id="ED78D1FBFFB53A18910AFD1FFB2437DB" box="[1129,1190,696,722]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="2" pageNumber="229" type="Museum">MZB</collectionCode>
</materialsCitation>
<materialsCitation id="3B014363FFB53A1891D2FD10FCE33613" collectingDate="2018-07-01" collectionCode="MZB" collectorName="Tim Teripang P" county="Hol." elevation="4" location="Baluran National Park" municipality="Bilik Sajile" pageId="2" pageNumber="229" specimenCode="LIPI 2018" specimenCount="1" stateProvince="Jawa Timur">
<collectingCounty id="62B731B2FFB53A1891D2FD10FB6337D8" box="[1201,1249,695,721]" pageId="2" pageNumber="229">Hol.</collectingCounty>
427 (70x
<quantity id="4C91E4DBFFB53A18902FFD10FA1F37DB" box="[1356,1437,695,722]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="229" unit="mm" value="15.0">15 mm</quantity>
in ethanol); both collected in
<collectingMunicipality id="6BB2D344FFB53A189488FD7CFDED37FC" box="[491,623,731,757]" pageId="2" pageNumber="229">Bilik Sajile</collectingMunicipality>
,
<location id="8EB61FE5FFB53A18971FFD7CFCFC37FC" LSID="urn:lsid:plazi:treatment:03C0F828FFB53A1F95F4FE80FD6C3192:8EB61FE5FFB53A18971FFD7CFCFC37FC" box="[636,894,731,757]" county="Hol." municipality="Bilik Sajile" name="Baluran National Park" pageId="2" pageNumber="229" stateProvince="Jawa Timur">Baluran National Park</location>
,
<collectingRegion id="49AD87DCFFB53A1896E8FD7BFC7B37FF" box="[907,1017,732,758]" country="Indonesia" name="Jawa Timur" pageId="2" pageNumber="229">East Java</collectingRegion>
, by snorkeling, under a rock,
<elevation id="0044AE0DFFB53A189030FD7CFF5A3610" metricMagnitude="0" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="2" pageNumber="229" unit="m" value="3.5" valueMax="4.0" valueMin="3.0">
<quantity id="4C91E4DBFFB53A189030FD7CFA1E37FF" box="[1363,1436,731,758]" metricMagnitude="0" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="2" pageNumber="229" unit="m" value="3.5" valueMax="4.0" valueMin="3.0">34 m</quantity>
depth
</elevation>
, coll. by
<collectorName id="269C2CE8FFB53A18945EFD58FE693610" box="[317,491,767,793]" pageId="2" pageNumber="229">Tim Teripang P</collectorName>
2O
<specimenCode id="DBCFE145FFB53A189777FCA7FD0B3610" box="[532,649,767,794]" collectionCode="LIPI" pageId="2" pageNumber="229">LIPI 2018</specimenCode>
,
<date id="FFD76FFEFFB53A1897F7FD58FC993610" box="[660,795,767,793]" pageId="2" pageNumber="229" value="2018-07-01">
<collectingDate id="EF939616FFB53A1897F7FD58FC993610" box="[660,795,767,793]" pageId="2" pageNumber="229" value="2018-07-01">1 July 2018</collectingDate>
</date>
.
<collectionCode id="ED78D1FBFFB53A189647FCA7FCE33613" box="[804,865,768,794]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="2" pageNumber="229" type="Museum">MZB</collectionCode>
</materialsCitation>
<materialsCitation id="3B014363FFB53A18960BFD58FE973668" collectingDate="2016-04" collectionCode="LIPI" collectorName="Bakaheuni &amp; Tim Teripang P" elevation="1" location="South" municipality="Hol." pageId="2" pageNumber="229" specimenCode="LIPI 2016" specimenCount="1" stateProvince="Lampung">
<collectingMunicipality id="6BB2D344FFB53A18960BFD58FC1A3613" box="[872,920,767,794]" pageId="2" pageNumber="229">Hol.</collectingMunicipality>
425 (110x
<quantity id="4C91E4DBFFB53A18916AFD58FBDB3613" box="[1033,1113,767,794]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="229" unit="mm" value="40.0">40 mm</quantity>
in ethanol), coll.
<collectorName id="269C2CE8FFB53A18907EFD58FA1A3610" box="[1309,1432,767,793]" pageId="2" pageNumber="229">Bakaheuni</collectorName>
,
<collectingRegion id="49AD87DCFFB53A1895F4FC83FE873637" box="[151,261,804,830]" country="Indonesia" name="Lampung" pageId="2" pageNumber="229">Lampung</collectingRegion>
,
<location id="8EB61FE5FFB53A189472FC84FED03634" LSID="urn:lsid:plazi:treatment:03C0F828FFB53A1F95F4FE80FD6C3192:8EB61FE5FFB53A189472FC84FED03634" box="[273,338,803,829]" municipality="Hol." name="South" pageId="2" pageNumber="229" stateProvince="Lampung">South</location>
<collectingRegion id="49AD87DCFFB53A189439FC84FE453634" box="[346,455,803,829]" country="Indonesia" name="Sumatera Utara" pageId="2" pageNumber="229">Sumatera</collectingRegion>
; during low tide, under a rock,
<elevation id="0044AE0DFFB53A18964AFC84FC613634" box="[809,995,803,830]" metricMagnitude="0" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="2" pageNumber="229" unit="m" value="1.0" valueMax="1.5" valueMin="0.5">
<quantity id="4C91E4DBFFB53A18964AFC84FC183637" box="[809,922,803,830]" metricMagnitude="0" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="2" pageNumber="229" unit="m" value="1.0" valueMax="1.5" valueMin="0.5">0.51.5 m</quantity>
depth
</elevation>
, coll. by
<collectorName id="269C2CE8FFB53A189129FC84FB7B3634" box="[1098,1273,803,829]" pageId="2" pageNumber="229">Tim Teripang P</collectorName>
2O
<specimenCode id="DBCFE145FFB53A189041FC83FA1A3634" box="[1314,1432,803,830]" collectionCode="LIPI" pageId="2" pageNumber="229">LIPI 2016</specimenCode>
,
<date id="FFD76FFEFFB53A1895F4FCE0FE973668" box="[151,277,839,865]" pageId="2" pageNumber="229" value="2016-04">
<collectingDate id="EF939616FFB53A1895F4FCE0FE973668" box="[151,277,839,865]" pageId="2" pageNumber="229" value="2016-04">April 2016</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<caption id="DF1619B6FFB53A1895F4F97FFD21321D" pageId="2" pageNumber="229" startId="2.[151,250,1752,1776]" targetBox="[257,1329,889,1728]" targetPageId="2" targetType="figure">
<paragraph id="8BD6493EFFB53A1895F4F97FFD21321D" blockId="2.[151,1437,1752,1812]" pageId="2" pageNumber="229">
<emphasis id="B91D952CFFB53A1895F4F97FFE9433F9" bold="true" box="[151,278,1752,1776]" pageId="2" pageNumber="229">FIGURE 2.</emphasis>
<taxonomicName id="4C6932BDFFB53A18947CF97FFD3A33F9" authority="(Ludwig, 1875)" authorityName="Kudingareng-Sulawesi" baseAuthorityName="Ludwig" baseAuthorityYear="1875" box="[287,696,1752,1776]" class="Holothuroidea" family="Holothuriidae" genus="Labidodemas" kingdom="Animalia" order="Holothuriida" pageId="2" pageNumber="229" phylum="Echinodermata" rank="species" species="rugosum">
<emphasis id="B91D952CFFB53A18947CF97FFD8833F9" box="[287,522,1752,1776]" italics="true" pageId="2" pageNumber="229">Labidodemas rugosum</emphasis>
(
<bibRefCitation id="EFF834CFFFB53A189779F97FFD3233F9" author="Ludwig, H." box="[538,688,1752,1776]" pageId="2" pageNumber="229" pagination="77 - 118" refId="ref8343" refString="Ludwig, H. (1875) Beitrage zur Kenntniss der Holothurien. Arbeiten aus dem Zoologisch-Zootomischen Institut in Wurzburg, 2, 77 - 118." type="journal article" year="1875">Ludwig, 1875</bibRefCitation>
)
</taxonomicName>
. Live specimen photograph, MZB Hol.427 (70x15 mm in ethanol). A: dorsal; B. ventral; C: illustration of calcareous ring.
</paragraph>
</caption>
<paragraph id="8BD6493EFFB53A1895A4F8C1FDFD32E4" blockId="2.[151,1437,1894,2029]" pageId="2" pageNumber="229">
<emphasis id="B91D952CFFB53A1895A4F8C1FE393289" bold="true" box="[199,443,1894,1920]" pageId="2" pageNumber="229">External description</emphasis>
. Body cylindrical; specimen colour in ethanol white uniformly (
<figureCitation id="135255BBFFB53A1891E1F8C1FA873289" box="[1154,1285,1894,1920]" captionStart="FIGURE 2" captionStartId="2.[151,250,1752,1776]" captionTargetBox="[257,1329,889,1728]" captionTargetId="figure-190@2.[257,1329,889,1728]" captionTargetPageId="2" captionText="FIGURE 2. Labidodemas rugosum (Ludwig, 1875). Live specimen photograph, MZB Hol.427 (70x15 mm in ethanol). A: dorsal; B. ventral; C: illustration of calcareous ring." pageId="2" pageNumber="229">Figs. 2A, B</figureCitation>
). Body rather gritty,
<quantity id="4C91E4DBFFB53A189580F82DFEC632AC" box="[227,324,1930,1957]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="2" pageNumber="229" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
thick. Mouth and anus terminal; mouth white in colour, surrounded by 1820 yellow tentacles. Dorsal papillae small, scattered over the surface. Ventral tube feet longer than dorsal papillae (
<quantity id="4C91E4DBFFB53A1891DDF809FAC932C1" box="[1214,1355,1966,1993]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="2" pageNumber="229" unit="mm" value="2.0" valueMax="2.5" valueMin="1.5">1.52.5 mm</quantity>
), white to yellowish, 36 rows in ambulacral areas.
</paragraph>
<caption id="DF1619B6FFB43A1995F4F9BBFC1A3351" pageId="3" pageNumber="230" startId="3.[151,250,1564,1588]" targetBox="[257,1329,181,1540]" targetPageId="3" targetType="figure">
<paragraph id="8BD6493EFFB43A1995F4F9BBFC1A3351" blockId="3.[151,1437,1564,1625]" pageId="3" pageNumber="230">
<emphasis id="B91D952CFFB43A1995F4F9BBFE96333D" bold="true" box="[151,276,1564,1588]" pageId="3" pageNumber="230">FIGURE 3.</emphasis>
<taxonomicName id="4C6932BDFFB43A19947EF9BBFD30333D" authority="(Ludwig, 1875)" authorityName="Kudingareng-Sulawesi" baseAuthorityName="Ludwig" baseAuthorityYear="1875" box="[285,690,1564,1588]" class="Holothuroidea" family="Holothuriidae" genus="Labidodemas" kingdom="Animalia" order="Holothuriida" pageId="3" pageNumber="230" phylum="Echinodermata" rank="species" species="rugosum">
<emphasis id="B91D952CFFB43A19947EF9BBFD84333D" box="[285,518,1564,1588]" italics="true" pageId="3" pageNumber="230">Labidodemas rugosum</emphasis>
(
<bibRefCitation id="EFF834CFFFB43A199776F9BBFD2B333D" author="Ludwig, H." box="[533,681,1564,1588]" pageId="3" pageNumber="230" pagination="77 - 118" refId="ref8343" refString="Ludwig, H. (1875) Beitrage zur Kenntniss der Holothurien. Arbeiten aus dem Zoologisch-Zootomischen Institut in Wurzburg, 2, 77 - 118." type="journal article" year="1875">Ludwig, 1875</bibRefCitation>
)
</taxonomicName>
. A, C, D, and E, MZB Hol.427; B, MZB Hol.390. Dorsal body wall, A: tables; B: buttons. Ventral body wall, C: tables; D: buttons. E: tentacle rods.
</paragraph>
</caption>
<paragraph id="8BD6493EFFB43A1995A4F90CFB883294" blockId="3.[151,1437,1706,1949]" pageId="3" pageNumber="230">
<emphasis id="B91D952CFFB43A1995A4F90CFE3533CD" bold="true" box="[199,439,1706,1733]" pageId="3" pageNumber="230">Internal description</emphasis>
. Calcareous ring with radial pieces twice higher than interradial ones (radial pieces
<quantity id="4C91E4DBFFB43A199011F90CFF4133E0" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="3" pageNumber="230" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
wide and
<quantity id="4C91E4DBFFB43A199451F969FE2133E0" box="[306,419,1742,1769]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="3" pageNumber="230" unit="mm" value="2.25" valueMax="3.0" valueMin="1.5">1.53 mm</quantity>
high) (
<figureCitation id="135255BBFFB43A199492F968FDCB33E0" box="[497,585,1743,1769]" captionStart="FIGURE 2" captionStartId="2.[151,250,1752,1776]" captionTargetBox="[257,1329,889,1728]" captionTargetId="figure-190@2.[257,1329,889,1728]" captionTargetPageId="2" captionText="FIGURE 2. Labidodemas rugosum (Ludwig, 1875). Live specimen photograph, MZB Hol.427 (70x15 mm in ethanol). A: dorsal; B. ventral; C: illustration of calcareous ring." pageId="3" pageNumber="230">Fig. 2C</figureCitation>
). Cuvierian tubules present only in Hol.390, adhesive, long and thin, whitish, branched, distal part of the tube smooth, more than 10 attachment sites. Tentacle ampullae present,
<quantity id="4C91E4DBFFB43A19919FF955FAD83204" box="[1276,1370,1778,1805]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="6.0" metricValueMin="2.0" pageId="3" pageNumber="230" unit="mm" value="4.0" valueMax="6.0" valueMin="2.0">26 mm</quantity>
(1/35 to 1/27 body length). Single Polian vesicle, sac-like. Stone canal straight, present in Hol.425 (
<quantity id="4C91E4DBFFB43A1991DEF8B0FA833238" box="[1213,1281,1815,1841]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="230" unit="mm" value="4.0">4 mm</quantity>
) and Hol.427 (
<quantity id="4C91E4DBFFB43A1995C3F89CFF6A325C" box="[160,232,1851,1877]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="230" unit="mm" value="2.0">2 mm</quantity>
). Gonad only observed in Hol.425, one tuft, whitish, branched. Respiratory tree arises from single stalk. Longitudinal muscle flat, medially attached and lateral margins free. Inner body wall whitish with numerous black dots under the muscles. Intestine filled with small gravel and rough coral sand.
</paragraph>
<caption id="DF1619B6FFB33A1E95F4F907FCFD33D5" pageId="4" pageNumber="231" startId="4.[151,250,1696,1720]" targetBox="[257,1329,181,1672]" targetPageId="4" targetType="figure">
<paragraph id="8BD6493EFFB33A1E95F4F907FCFD33D5" blockId="4.[151,1437,1696,1756]" pageId="4" pageNumber="231">
<emphasis id="B91D952CFFB33A1E95F4F907FE9033B1" bold="true" box="[151,274,1696,1720]" pageId="4" pageNumber="231">FIGURE 4.</emphasis>
<taxonomicName id="4C6932BDFFB33A1E947BF907FD2433B1" authority="(Ludwig, 1875)" authorityName="Kudingareng-Sulawesi" baseAuthorityName="Ludwig" baseAuthorityYear="1875" box="[280,678,1696,1720]" class="Holothuroidea" family="Holothuriidae" genus="Labidodemas" kingdom="Animalia" order="Holothuriida" pageId="4" pageNumber="231" phylum="Echinodermata" rank="species" species="rugosum">
<emphasis id="B91D952CFFB33A1E947BF907FE7D33B1" box="[280,511,1696,1720]" italics="true" pageId="4" pageNumber="231">Labidodemas rugosum</emphasis>
(
<bibRefCitation id="EFF834CFFFB33A1E9768F907FD1F33B1" author="Ludwig, H." box="[523,669,1696,1720]" pageId="4" pageNumber="231" pagination="77 - 118" refId="ref8343" refString="Ludwig, H. (1875) Beitrage zur Kenntniss der Holothurien. Arbeiten aus dem Zoologisch-Zootomischen Institut in Wurzburg, 2, 77 - 118." type="journal article" year="1875">Ludwig, 1875</bibRefCitation>
)
</taxonomicName>
. MZB Hol.427. Dorsal papillae, A: rod-like buttons; B: rods; C: buttons; D: tables. Ventral tube feet, E: tables; F: buttons; G: buttons-plate; H: plates.
</paragraph>
</caption>
<paragraph id="8BD6493EFFB33A1E95A4F880FAD032FF" blockId="4.[151,1437,1831,2039]" pageId="4" pageNumber="231">
<emphasis id="B91D952CFFB33A1E95A4F880FEA43248" bold="true" box="[199,294,1831,1857]" pageId="4" pageNumber="231">Ossicles</emphasis>
. Dorsal body wall with tables and buttons; table discs 70110 µm across, disc rims very spinous; margin of the disc slightly turned upward; perforated by four central holes; spire moderate to high, with four undulating pillars united by one cross beam, terminating in narrow crown consisting of a cluster of stout spines (
<figureCitation id="135255BBFFB33A1E900CF8D7FF3F32A7" captionStart="FIGURE 3" captionStartId="3.[151,250,1564,1588]" captionTargetBox="[257,1329,181,1540]" captionTargetId="figure-17@3.[257,1329,181,1540]" captionTargetPageId="3" captionText="FIGURE 3. Labidodemas rugosum (Ludwig, 1875). A, C, D, and E, MZB Hol.427; B, MZB Hol.390. Dorsal body wall, A: tables; B: buttons. Ventral body wall, C: tables; D: buttons. E: tentacle rods." pageId="4" pageNumber="231">Fig. 3A</figureCitation>
). Buttons, 3040 µm long, with three to four pairs of irregular holes, margin smooth but undulating (
<figureCitation id="135255BBFFB33A1E9055F833FA0C32A7" box="[1334,1422,1940,1966]" captionStart="FIGURE 3" captionStartId="3.[151,250,1564,1588]" captionTargetBox="[257,1329,181,1540]" captionTargetId="figure-17@3.[257,1329,181,1540]" captionTargetPageId="3" captionText="FIGURE 3. Labidodemas rugosum (Ludwig, 1875). A, C, D, and E, MZB Hol.427; B, MZB Hol.390. Dorsal body wall, A: tables; B: buttons. Ventral body wall, C: tables; D: buttons. E: tentacle rods." pageId="4" pageNumber="231">Fig. 3B</figureCitation>
). Ventral body wall with tables and buttons; their shape similar to those of the dorsal body wall but bigger, and buttons more numerous than in dorsal body wall. Table discs 90120 µm, and buttons 5070 µm long (
<figureCitation id="135255BBFFB33A1E91A3F87BFAC432FF" box="[1216,1350,2012,2038]" captionStart="FIGURE 3" captionStartId="3.[151,250,1564,1588]" captionTargetBox="[257,1329,181,1540]" captionTargetId="figure-17@3.[257,1329,181,1540]" captionTargetPageId="3" captionText="FIGURE 3. Labidodemas rugosum (Ludwig, 1875). A, C, D, and E, MZB Hol.427; B, MZB Hol.390. Dorsal body wall, A: tables; B: buttons. Ventral body wall, C: tables; D: buttons. E: tentacle rods." pageId="4" pageNumber="231">Figs. 3C, D</figureCitation>
).
</paragraph>
<paragraph id="8BD6493EFFB23A1F95A4FF30FC4434C6" blockId="5.[151,1437,151,1179]" pageId="5" pageNumber="232">
Dorsal papillae with tables, buttons and rods; tables similar in shape to those of dorsal body wall, but generally bigger, table disc 80130 µm across (
<figureCitation id="135255BBFFB23A1F975DFF1CFD1B35DC" box="[574,665,187,213]" captionStart="FIGURE 4" captionStartId="4.[151,250,1696,1720]" captionTargetBox="[257,1329,181,1672]" captionTargetId="figure-25@4.[257,1329,181,1672]" captionTargetPageId="4" captionText="FIGURE 4. Labidodemas rugosum (Ludwig, 1875). MZB Hol.427. Dorsal papillae, A: rod-like buttons; B: rods; C: buttons; D: tables. Ventral tube feet, E: tables; F: buttons; G: buttons-plate; H: plates." pageId="5" pageNumber="232">Fig. 4D</figureCitation>
). Buttons in several shapes; rods-like buttons, 80110 µm long (
<figureCitation id="135255BBFFB23A1F900CFF1CFF3F35F0" captionStart="FIGURE 4" captionStartId="4.[151,250,1696,1720]" captionTargetBox="[257,1329,181,1672]" captionTargetId="figure-25@4.[257,1329,181,1672]" captionTargetPageId="4" captionText="FIGURE 4. Labidodemas rugosum (Ludwig, 1875). MZB Hol.427. Dorsal papillae, A: rod-like buttons; B: rods; C: buttons; D: tables. Ventral tube feet, E: tables; F: buttons; G: buttons-plate; H: plates." pageId="5" pageNumber="232">Fig. 4A</figureCitation>
); buttons with four to five pairs of irregular holes, smooth rim but undulating, 5080 µm long; buttons with rim rather spinous but blunt spines, only a few, 6070 µm long (
<figureCitation id="135255BBFFB23A1F9619FEA4FC573414" box="[890,981,259,285]" captionStart="FIGURE 4" captionStartId="4.[151,250,1696,1720]" captionTargetBox="[257,1329,181,1672]" captionTargetId="figure-25@4.[257,1329,181,1672]" captionTargetPageId="4" captionText="FIGURE 4. Labidodemas rugosum (Ludwig, 1875). MZB Hol.427. Dorsal papillae, A: rod-like buttons; B: rods; C: buttons; D: tables. Ventral tube feet, E: tables; F: buttons; G: buttons-plate; H: plates." pageId="5" pageNumber="232">Fig. 4C</figureCitation>
). Rods scarce, with several blunt spines and the extremities with or without perforation, 80110 µm long (
<figureCitation id="135255BBFFB23A1F96E5FE81FC623449" box="[902,992,294,320]" captionStart="FIGURE 4" captionStartId="4.[151,250,1696,1720]" captionTargetBox="[257,1329,181,1672]" captionTargetId="figure-25@4.[257,1329,181,1672]" captionTargetPageId="4" captionText="FIGURE 4. Labidodemas rugosum (Ludwig, 1875). MZB Hol.427. Dorsal papillae, A: rod-like buttons; B: rods; C: buttons; D: tables. Ventral tube feet, E: tables; F: buttons; G: buttons-plate; H: plates." pageId="5" pageNumber="232">Fig. 4B</figureCitation>
). Ventral tube feet with tables, buttons, and perforated plates and endplates. Tables and buttons shape similar to those in the ventral body wall, table discs 6090 µm across and button 6090 µm long (
<figureCitation id="135255BBFFB23A1F97FDFEC9FCA23481" box="[670,800,366,392]" captionStart="FIGURE 4" captionStartId="4.[151,250,1696,1720]" captionTargetBox="[257,1329,181,1672]" captionTargetId="figure-25@4.[257,1329,181,1672]" captionTargetPageId="4" captionText="FIGURE 4. Labidodemas rugosum (Ludwig, 1875). MZB Hol.427. Dorsal papillae, A: rod-like buttons; B: rods; C: buttons; D: tables. Ventral tube feet, E: tables; F: buttons; G: buttons-plate; H: plates." pageId="5" pageNumber="232">Figs. 4E, F</figureCitation>
). Perforated plates many, 60100 µm across, undulating or spinous edges (
<figureCitation id="135255BBFFB23A1F9408FE36FE4B34A5" box="[363,457,401,428]" captionStart="FIGURE 4" captionStartId="4.[151,250,1696,1720]" captionTargetBox="[257,1329,181,1672]" captionTargetId="figure-25@4.[257,1329,181,1672]" captionTargetPageId="4" captionText="FIGURE 4. Labidodemas rugosum (Ludwig, 1875). MZB Hol.427. Dorsal papillae, A: rod-like buttons; B: rods; C: buttons; D: tables. Ventral tube feet, E: tables; F: buttons; G: buttons-plate; H: plates." pageId="5" pageNumber="232">Fig. 4G</figureCitation>
). Endplates, 240300 µm across (
<figureCitation id="135255BBFFB23A1F9630FE36FC3334A5" box="[851,945,401,428]" captionStart="FIGURE 4" captionStartId="4.[151,250,1696,1720]" captionTargetBox="[257,1329,181,1672]" captionTargetId="figure-25@4.[257,1329,181,1672]" captionTargetPageId="4" captionText="FIGURE 4. Labidodemas rugosum (Ludwig, 1875). MZB Hol.427. Dorsal papillae, A: rod-like buttons; B: rods; C: buttons; D: tables. Ventral tube feet, E: tables; F: buttons; G: buttons-plate; H: plates." pageId="5" pageNumber="232">Fig. 4H</figureCitation>
). Tentacles with rods, mostly with smooth surfaces, straight or curved shapes, rarely with coarser surfaces (
<figureCitation id="135255BBFFB23A1F9609FE12FC3C34C6" box="[874,958,437,463]" captionStart="FIGURE 3" captionStartId="3.[151,250,1564,1588]" captionTargetBox="[257,1329,181,1540]" captionTargetId="figure-17@3.[257,1329,181,1540]" captionTargetPageId="3" captionText="FIGURE 3. Labidodemas rugosum (Ludwig, 1875). A, C, D, and E, MZB Hol.427; B, MZB Hol.390. Dorsal body wall, A: tables; B: buttons. Ventral body wall, C: tables; D: buttons. E: tentacle rods." pageId="5" pageNumber="232">Fig. 3E</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3731AB5FFB23A1F95A4FE7EFD7B36A9" pageId="5" pageNumber="232" type="discussion">
<paragraph id="8BD6493EFFB23A1F95A4FE7EFE55378B" blockId="5.[151,1437,151,1179]" pageId="5" pageNumber="232">
<emphasis id="B91D952CFFB23A1F95A4FE7EFEB634FA" bold="true" box="[199,308,473,499]" pageId="5" pageNumber="232">Remarks</emphasis>
. The body colour of the current materials is uniformly white, as described by
<bibRefCitation id="EFF834CFFFB23A1F91D3FE7EFA1A34FA" author="Massin, C. &amp; Samyn, Y. &amp; Thandar, A. S." box="[1200,1432,473,500]" pageId="5" pageNumber="232" pagination="1811 - 1847" refId="ref8451" refString="Massin, C., Samyn, Y. &amp; Thandar, A. S. (2004) The genus Labidodemas (Holothuroidea: Aspidochirotida) revisited, with description of three new species and with re-positioning of Holothuria (Irenothuria) maccullochi Deichmann, 1958. Journal of Natural History, 38, 1811 - 1847. https: // doi. org / 10.1080 / 0022293031000156268" type="journal article" year="2004">
Massin
<emphasis id="B91D952CFFB23A1F906AFE7DFAB934FA" box="[1289,1339,473,499]" italics="true" pageId="5" pageNumber="232">et al</emphasis>
. (2004)
</bibRefCitation>
,
<bibRefCitation id="EFF834CFFFB23A1F95F4FE5AFEBE371E" author="Massin, C." box="[151,316,509,535]" pageId="5" pageNumber="232" refId="ref8410" refString="Massin, C. (1999) Reef-dwelling Holothuroidea (Echinodermata) of the Spermonde Archipelago (South-West Sulawesi, Indonesia). Zoologische verhandelingen. Vol. 329. National Museum of Natural History, Leiden, 144 pp." type="book" year="1999">Massin (1999)</bibRefCitation>
, and
<bibRefCitation id="EFF834CFFFB23A1F9414FE5AFDA7371E" author="Pearson, J." box="[375,549,509,535]" pageId="5" pageNumber="232" pagination="49 - 101" refId="ref8553" refString="Pearson, J. (1913) Notes on the Holothurioidea of the Indian Ocean. Spolia Zeylandica, 9, 49 - 101. https: // doi. org / 10.5962 / bhl. part. 7317" type="journal article" year="1913">Pearson (1913)</bibRefCitation>
; not translucent pink, as described by
<bibRefCitation id="EFF834CFFFB23A1F96ACFE5AFBEA371E" author="James, D. B." box="[975,1128,509,535]" pageId="5" pageNumber="232" pagination="82 - 85" refId="ref8077" refString="James, D. B. (1981) Studies on Indian Echinoderms 7. On a new Family Labidodematidae (Holothurioidea: Aspidochirotida) with a detailed description of Labidodemas rugosum (Ludwig) from the Andamans. Journal of the Marine Biological Association of India, 23, 82 - 85." type="journal article" year="1981">James (1981)</bibRefCitation>
. Specimens from this study found 1820 tentacles, contrary to the previously observed 20 (
<bibRefCitation id="EFF834CFFFB23A1F9613FD87FBD13732" author="Massin, C. &amp; Samyn, Y. &amp; Thandar, A. S." box="[880,1107,544,571]" pageId="5" pageNumber="232" pagination="1811 - 1847" refId="ref8451" refString="Massin, C., Samyn, Y. &amp; Thandar, A. S. (2004) The genus Labidodemas (Holothuroidea: Aspidochirotida) revisited, with description of three new species and with re-positioning of Holothuria (Irenothuria) maccullochi Deichmann, 1958. Journal of Natural History, 38, 1811 - 1847. https: // doi. org / 10.1080 / 0022293031000156268" type="journal article" year="2004">
Massin
<emphasis id="B91D952CFFB23A1F96AFFD86FB823733" box="[972,1024,544,570]" italics="true" pageId="5" pageNumber="232">et al</emphasis>
., 2004
</bibRefCitation>
;
<bibRefCitation id="EFF834CFFFB23A1F9103FD87FB7F3732" author="Massin, C." box="[1120,1277,544,571]" pageId="5" pageNumber="232" refId="ref8410" refString="Massin, C. (1999) Reef-dwelling Holothuroidea (Echinodermata) of the Spermonde Archipelago (South-West Sulawesi, Indonesia). Zoologische verhandelingen. Vol. 329. National Museum of Natural History, Leiden, 144 pp." type="book" year="1999">Massin, 1999</bibRefCitation>
;
<bibRefCitation id="EFF834CFFFB23A1F9068FD86FA153733" author="Rowe, F. W. E." box="[1291,1431,544,571]" pageId="5" pageNumber="232" pagination="117 - 170" refId="ref8722" refString="Rowe, F. W. E. (1969) A Review of The Family Holothuriidae. (Holothurioidea: Aspidochirotida). Bulletin of the British Museum (Natural History), Zoology, 18, 117 - 170. https: // doi. org / 10.5962 / bhl. part. 18419" type="journal article" year="1969">Rowe, 1969</bibRefCitation>
;
<bibRefCitation id="EFF834CFFFB23A1F95F4FDE3FEEE3757" author="Cherbonnier, G." box="[151,364,580,606]" pageId="5" pageNumber="232" pagination="1 - 292" refId="ref7768" refString="Cherbonnier, G. (1988) Echinodermes: Holothurides. Faune de Madagascar, 70, 1 - 292." type="journal article" year="1988">Cherbonnier, 1988</bibRefCitation>
;
<bibRefCitation id="EFF834CFFFB23A1F941AFDE3FD993756" author="Ludwig, H." box="[377,539,580,607]" pageId="5" pageNumber="232" pagination="77 - 118" refId="ref8343" refString="Ludwig, H. (1875) Beitrage zur Kenntniss der Holothurien. Arbeiten aus dem Zoologisch-Zootomischen Institut in Wurzburg, 2, 77 - 118." type="journal article" year="1875">Ludwig, 1875</bibRefCitation>
). Cuvierian tubules are absent according to
<bibRefCitation id="EFF834CFFFB23A1F916DFDE3FB773757" author="Massin, C. &amp; Samyn, Y. &amp; Thandar, A. S." box="[1038,1269,580,607]" pageId="5" pageNumber="232" pagination="1811 - 1847" refId="ref8451" refString="Massin, C., Samyn, Y. &amp; Thandar, A. S. (2004) The genus Labidodemas (Holothuroidea: Aspidochirotida) revisited, with description of three new species and with re-positioning of Holothuria (Irenothuria) maccullochi Deichmann, 1958. Journal of Natural History, 38, 1811 - 1847. https: // doi. org / 10.1080 / 0022293031000156268" type="journal article" year="2004">
Massin
<emphasis id="B91D952CFFB23A1F9104FDE2FB1B3757" box="[1127,1177,580,606]" italics="true" pageId="5" pageNumber="232">et al</emphasis>
. (2004)
</bibRefCitation>
, but present in the specimen MZB Hol.390.
</paragraph>
<paragraph id="8BD6493EFFB23A1F95A4FD2BFD7B36A9" blockId="5.[151,1437,151,1179]" pageId="5" pageNumber="232">
The current material ossicles differ from previous author descriptions in some ways. Tentacle rods are short (4075 µm); many large table discs up to 100 µm, with four central holes and one ring of 812 peripheral holes, but occasionally with two rings of peripheral holes up to 14 holes in total, with the outer rings holes being smaller than the inner. The rim of the table disc is spinous, and the spines are longer and much more pointed than in
<bibRefCitation id="EFF834CFFFB23A1F9028FD50FEA1363C" author="Massin, C. &amp; Samyn, Y. &amp; Thandar, A. S." pageId="5" pageNumber="232" pagination="1811 - 1847" refId="ref8451" refString="Massin, C., Samyn, Y. &amp; Thandar, A. S. (2004) The genus Labidodemas (Holothuroidea: Aspidochirotida) revisited, with description of three new species and with re-positioning of Holothuria (Irenothuria) maccullochi Deichmann, 1958. Journal of Natural History, 38, 1811 - 1847. https: // doi. org / 10.1080 / 0022293031000156268" type="journal article" year="2004">
Massin
<emphasis id="B91D952CFFB23A1F95F4FCBCFF4A363C" box="[151,200,795,821]" italics="true" pageId="5" pageNumber="232">et al</emphasis>
. (2004)
</bibRefCitation>
,
<bibRefCitation id="EFF834CFFFB23A1F944DFCBCFE50363C" author="Massin, C." box="[302,466,795,821]" pageId="5" pageNumber="232" refId="ref8410" refString="Massin, C. (1999) Reef-dwelling Holothuroidea (Echinodermata) of the Spermonde Archipelago (South-West Sulawesi, Indonesia). Zoologische verhandelingen. Vol. 329. National Museum of Natural History, Leiden, 144 pp." type="book" year="1999">Massin (1999)</bibRefCitation>
, and
<bibRefCitation id="EFF834CFFFB23A1F976EFCBCFD24363C" author="James, D. B." box="[525,678,795,821]" pageId="5" pageNumber="232" pagination="82 - 85" refId="ref8077" refString="James, D. B. (1981) Studies on Indian Echinoderms 7. On a new Family Labidodematidae (Holothurioidea: Aspidochirotida) with a detailed description of Labidodemas rugosum (Ludwig) from the Andamans. Journal of the Marine Biological Association of India, 23, 82 - 85." type="journal article" year="1981">James (1981)</bibRefCitation>
, but it is similar to
<bibRefCitation id="EFF834CFFFB23A1F961FFCBCFBA8363C" author="Pearson, J." box="[892,1066,795,821]" pageId="5" pageNumber="232" pagination="49 - 101" refId="ref8553" refString="Pearson, J. (1913) Notes on the Holothurioidea of the Indian Ocean. Spolia Zeylandica, 9, 49 - 101. https: // doi. org / 10.5962 / bhl. part. 7317" type="journal article" year="1913">Pearson (1913)</bibRefCitation>
. Table crowns never have central perforation, whereas
<bibRefCitation id="EFF834CFFFB23A1F94E5FC99FDEE3651" author="Massin, C. &amp; Samyn, Y. &amp; Thandar, A. S." box="[390,620,830,857]" pageId="5" pageNumber="232" pagination="1811 - 1847" refId="ref8451" refString="Massin, C., Samyn, Y. &amp; Thandar, A. S. (2004) The genus Labidodemas (Holothuroidea: Aspidochirotida) revisited, with description of three new species and with re-positioning of Holothuria (Irenothuria) maccullochi Deichmann, 1958. Journal of Natural History, 38, 1811 - 1847. https: // doi. org / 10.1080 / 0022293031000156268" type="journal article" year="2004">
Massin
<emphasis id="B91D952CFFB23A1F94BCFC98FD903651" box="[479,530,830,856]" italics="true" pageId="5" pageNumber="232">et al</emphasis>
. (2004)
</bibRefCitation>
usually have a small central perforation. MZB Hol.427 lacks body wall buttons but is present in MZB Hol.390 and MZB Hol.425 with undulating margins. Rods in dorsal papillae of present materials have never been documented before.
</paragraph>
</subSubSection>
<subSubSection id="C3731AB5FFB23A1F95A4FC0DFD6C3192" pageId="5" pageNumber="232" type="biology_ecology">
<paragraph id="8BD6493EFFB23A1F95A4FC0DFCEB3102" blockId="5.[151,1437,151,1179]" pageId="5" pageNumber="232">
<emphasis id="B91D952CFFB23A1F95A4FC0DFEA436CD" bold="true" box="[199,294,938,964]" pageId="5" pageNumber="232">Ecology</emphasis>
. Found in shallow waters, always less than
<quantity id="4C91E4DBFFB23A1F966CFC0DFCCA36CD" box="[783,840,938,964]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="232" unit="m" value="20.0">20 m</quantity>
deep (
<bibRefCitation id="EFF834CFFFB23A1F96F1FC0DFBCF36CD" author="Lane, D. J. W. &amp; Marsh, L. M. &amp; VandenSpiegel, D. &amp; Rowe, F. W. E." box="[914,1101,937,964]" pageId="5" pageNumber="232" pagination="459 - 493" refId="ref8229" refString="Lane, D. J. W., Marsh, L. M., VandenSpiegel, D. &amp; Rowe, F. W. E. (2000) Echinoderm Fauna of the South China Sea: an Inventory and Analysis of Distribution Patterns. Raffles Bulletin of Zoology, 48, 459 - 493." type="journal article" year="2000">
Lane
<emphasis id="B91D952CFFB23A1F96B1FC0DFB8136CA" box="[978,1027,937,963]" italics="true" pageId="5" pageNumber="232">et al</emphasis>
. 2000
</bibRefCitation>
), but specimens from
<collectingCountry id="F37E09AEFFB23A1F9022FC0EFA1A36CA" box="[1345,1432,937,963]" name="Indonesia" pageId="5" pageNumber="232">Ambon</collectingCountry>
,
<collectingRegion id="49AD87DCFFB23A1F95F4FC6AFF6C36EE" box="[151,238,973,999]" country="Indonesia" name="Maluku Utara" pageId="5" pageNumber="232">Maluku</collectingRegion>
were found at
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depth (
<bibRefCitation id="EFF834CFFFB23A1F9757FC6AFC9F36EE" author="Jangoux, M. &amp; De Ridder, C. &amp; Massin, C. &amp; Darsono, P." box="[564,797,973,1000]" pageId="5" pageNumber="232" pagination="161 - 170" refId="ref8131" refString="Jangoux, M., De Ridder, C., Massin, C. &amp; Darsono, P. (1989) The holothuroids, echinoids and asteroids (echinodermata) collected by the Snellius-II expedition. Netherlands Journal of Sea Research, 23, 161 - 170. https: // doi. org / 10.1016 / 0077 - 7579 (89) 90010 - 0" type="journal article" year="1989">
Jangoux
<emphasis id="B91D952CFFB23A1F97FEFC69FD5336EE" box="[669,721,973,999]" italics="true" pageId="5" pageNumber="232">et al</emphasis>
. 1989
</bibRefCitation>
), detritus/deposit feeder, under coral rocks. Intestine all specimens of this study filled with coarse sand and small gravel.
</paragraph>
<paragraph id="8BD6493EFFB23A1F95A4FBB2FE443126" blockId="5.[151,1437,151,1179]" box="[199,454,1045,1071]" pageId="5" pageNumber="232">
<emphasis id="B91D952CFFB23A1F95A4FBB2FED93126" bold="true" box="[199,347,1045,1071]" pageId="5" pageNumber="232">Depth range</emphasis>
.
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.
</paragraph>
<paragraph id="8BD6493EFFB23A1F95A4FB9FFD6C3192" blockId="5.[151,1437,151,1179]" pageId="5" pageNumber="232">
<emphasis id="B91D952CFFB23A1F95A4FB9FFE68315B" bold="true" box="[199,490,1080,1106]" pageId="5" pageNumber="232">Geographic distribution</emphasis>
. The following new localities are added here since
<bibRefCitation id="EFF834CFFFB23A1F9154FB9EFADC315A" author="Massin, C. &amp; Samyn, Y. &amp; Thandar, A. S." box="[1079,1374,1081,1107]" pageId="5" pageNumber="232" pagination="1811 - 1847" refId="ref8451" refString="Massin, C., Samyn, Y. &amp; Thandar, A. S. (2004) The genus Labidodemas (Holothuroidea: Aspidochirotida) revisited, with description of three new species and with re-positioning of Holothuria (Irenothuria) maccullochi Deichmann, 1958. Journal of Natural History, 38, 1811 - 1847. https: // doi. org / 10.1080 / 0022293031000156268" type="journal article" year="2004">
Massin
<emphasis id="B91D952CFFB23A1F91F2FB9DFB46315A" box="[1169,1220,1081,1107]" italics="true" pageId="5" pageNumber="232">et al</emphasis>
. (2004: 1831
</bibRefCitation>
, map 5) publication: West Lombok Island,
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(
<bibRefCitation id="EFF834CFFFB23A1F97DFFBFAFC64317E" author="Purwati, P. &amp; Wirawati, I." box="[700,998,1117,1143]" pageId="5" pageNumber="232" pagination="1 - 10" refId="ref8592" refString="Purwati, P. &amp; Wirawati, I. (2011) Holothuriidae (Echinodermata, Holothuroidea, Aspidochirotida) Perairan Dangkal Lombok Barat Bagian II. Genus Actinopyga, Bohadschia, Pearsonothuria, Labidodemas. Jurnal Oseanologi, 3 (1 / 2), 1 - 10." type="journal article" year="2011">Purwati &amp; Wirawati 2011</bibRefCitation>
), Baluran National Park-East
<collectingRegion id="49AD87DCFFB23A1F9059FBFAFAEE317E" box="[1338,1388,1117,1143]" country="Indonesia" name="Jawa Timur" pageId="5" pageNumber="232">Java</collectingRegion>
and Lampung-South
<collectingRegion id="49AD87DCFFB23A1F9437FB26FE433192" box="[340,449,1153,1179]" country="Indonesia" name="Sumatera Utara" pageId="5" pageNumber="232">Sumatera</collectingRegion>
,
<collectingCountry id="F37E09AEFFB23A1F94AFFB26FDBB3192" box="[460,569,1153,1179]" name="Indonesia" pageId="5" pageNumber="232">Indonesia</collectingCountry>
(present study).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,817 @@
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<paragraph id="8BD6493EFFBF3A1295F4F9D0FEBF3398" blockId="8.[151,532,1618,1681]" box="[151,317,1655,1681]" pageId="8" pageNumber="235">
(
<figureCitation id="135255BBFFBF3A1295FCF9D0FEB73398" box="[159,309,1655,1681]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStart-2="FIGURE 11" captionStart-3="FIGURE 12" captionStart-4="FIGURE 13" captionStartId-0="9.[151,250,1679,1703]" captionStartId-1="10.[151,250,1681,1705]" captionStartId-2="11.[151,250,1483,1507]" captionStartId-3="12.[151,250,1143,1167]" captionStartId-4="14.[151,250,1716,1740]" captionTargetBox-0="[257,1329,181,1656]" captionTargetBox-1="[257,1329,181,1657]" captionTargetBox-2="[257,1329,273,1460]" captionTargetBox-3="[257,1329,181,1119]" captionTargetBox-4="[257,1329,181,1693]" captionTargetId-0="figure-17@9.[257,1329,181,1656]" captionTargetId-1="figure-25@10.[257,1329,181,1657]" captionTargetId-2="figure-67@11.[257,1329,273,1460]" captionTargetId-3="figure-25@12.[257,1329,181,1119]" captionTargetId-4="figure-25@14.[257,1329,181,1693]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="12" captionTargetPageId-4="14" captionText-0="FIGURE 9. Labidodemas javaensis sp. nov. Holotype, MZB Hol.413 (85x30 mm in ethanol). A: mouth; B: dorsal; C: anal papillae; D: ventral; E: tentacles; F: calcareous ring." captionText-1="FIGURE 10. Labidodemas javaensis sp. nov. Holotype, MZB Hol.413. Ossicles of dorsal body wall. A: tables; B: tri-radiate rods; C: regular rods; D: rods with perforations; E: pseudo-buttons. F: tables of ventral body wall." captionText-2="FIGURE 11. Labidodemas javaensis sp. nov. Holotype. MZB Hol.413. Ossicles of dorsal papillae.A: tables; B: smaller tables with spinous disc and crown; C: pseudo-buttons; D: rods with rugose surface; E: rods with smooth surface." captionText-3="FIGURE 12. Labidodemas javaensis sp. nov. Holotype, MZB Hol.413. Ossicles of ventral tube feet. A: tables; B: tables with reduce pillar and crowns; C: pseudo-buttons; D: rod; E: large perforation endplate." captionText-4="FIGURE 13. Labidodemas javaensis sp. nov. Holotype. MZB Hol.413. A: tentacle rods. Ossicles of anal papillae, B: tables; C: tri-radiate rods; D: rods with smooth surface; E: rods with rugose surface; F: rods rugose surface with perforations." pageId="8" pageNumber="235">Figures 913</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3731AB5FFBF3A1195F4F918FF7432B1" lastPageId="11" lastPageNumber="238" pageId="8" pageNumber="235" type="materials_examined">
<paragraph id="8BD6493EFFBF3A1295F4F918FEE133F4" blockId="8.[151,1437,1727,2041]" pageId="8" pageNumber="235">
<emphasis id="B91D952CFFBF3A1295F4F918FEBF33D0" bold="true" box="[151,317,1727,1753]" pageId="8" pageNumber="235">Type material</emphasis>
.
<materialsCitation id="3B014363FFBF3A129429F918FEE133F4" collectingDate="2018-07-01" collectionCode="MZB, LIPI" collectorName="Hol. &amp; Tim Teripang P" elevation="3" pageId="8" pageNumber="235" specimenCount="1" typeStatus="holotype">
<typeStatus id="54D2F79CFFBF3A129429F918FE3033D0" box="[330,434,1727,1753]" pageId="8" pageNumber="235" type="holotype">Holotype</typeStatus>
<collectionCode id="ED78D1FBFFBF3A1294DAF918FE7433D0" box="[441,502,1727,1753]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="8" pageNumber="235" type="Museum">MZB</collectionCode>
<collectorName id="269C2CE8FFBF3A12949FF918FDAC33D0" box="[508,558,1727,1753]" pageId="8" pageNumber="235">Hol.</collectorName>
413; 85x
<quantity id="4C91E4DBFFBF3A1297EDF918FD5F33D0" box="[654,733,1727,1754]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="235" unit="mm" value="30.0">30 mm</quantity>
in ethanol; under a rock,
<elevation id="0044AE0DFFBF3A12969BF918FBE833D0" box="[1016,1130,1727,1753]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="235" unit="m" value="3.0">
<quantity id="4C91E4DBFFBF3A12969BF918FBA033D0" box="[1016,1058,1727,1753]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="235" unit="m" value="3.0">3 m</quantity>
depth
</elevation>
; coll. by
<collectorName id="269C2CE8FFBF3A1291AEF918FAF833D0" box="[1229,1402,1727,1753]" pageId="8" pageNumber="235">Tim Teripang P</collectorName>
2O
<collectionCode id="ED78D1FBFFBF3A1295F4F944FF4F33F4" box="[151,205,1763,1789]" pageId="8" pageNumber="235">LIPI</collectionCode>
,
<date id="FFD76FFEFFBF3A1295BAF944FEDD33F4" box="[217,351,1763,1789]" pageId="8" pageNumber="235" value="2018-07-01">
<collectingDate id="EF939616FFBF3A1295BAF944FEDD33F4" box="[217,351,1763,1789]" pageId="8" pageNumber="235" value="2018-07-01">1 July 2018</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BD6493EFFBF3A1295A4F8A0FC6F3228" blockId="8.[151,1437,1727,2041]" box="[199,1005,1799,1825]" pageId="8" pageNumber="235">
<materialsCitation id="3B014363FFBF3A1295A4F8A0FC683228" box="[199,1002,1799,1825]" country="Indonesia" location="Bilik Sajile" municipality="Baluran National Park" pageId="8" pageNumber="235" specimenCount="1" stateProvince="Jawa Timur" typeStatus="holotype">
<emphasis id="B91D952CFFBF3A1295A4F8A0FEE43228" bold="true" box="[199,358,1799,1825]" pageId="8" pageNumber="235">
<typeStatus id="54D2F79CFFBF3A1295A4F8A0FE803228" box="[199,258,1799,1825]" pageId="8" pageNumber="235">Type</typeStatus>
locality.
</emphasis>
<location id="8EB61FE5FFBF3A12940FF8A0FE6C3228" LSID="urn:lsid:plazi:treatment:03C0F828FFBF3A1795F4F9F5FC7132FF:8EB61FE5FFBF3A12940FF8A0FE6C3228" box="[364,494,1799,1825]" country="Indonesia" municipality="Baluran National Park" name="Bilik Sajile" pageId="8" pageNumber="235" stateProvince="Jawa Timur">Bilik Sajile</location>
,
<collectingMunicipality id="6BB2D344FFBF3A12949AF8A0FD743228" box="[505,758,1799,1825]" pageId="8" pageNumber="235">Baluran National Park</collectingMunicipality>
,
<collectingRegion id="49AD87DCFFBF3A129661F8A0FCEF3228" box="[770,877,1799,1825]" country="Indonesia" name="Jawa Timur" pageId="8" pageNumber="235">East Java</collectingRegion>
,
<collectingCountry id="F37E09AEFFBF3A12961AF8A0FC683228" box="[889,1002,1799,1825]" name="Indonesia" pageId="8" pageNumber="235">Indonesia</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8BD6493EFFBF3A1295A4F88CFDD332F0" blockId="8.[151,1437,1727,2041]" pageId="8" pageNumber="235">
<emphasis id="B91D952CFFBF3A1295A4F88CFDEB324C" bold="true" box="[199,617,1835,1861]" pageId="8" pageNumber="235">
External morphology of
<typeStatus id="54D2F79CFFBF3A129498F88CFDE6324C" box="[507,612,1835,1861]" pageId="8" pageNumber="235" type="holotype">holotype</typeStatus>
.
</emphasis>
Medium-sized specimen (85 x
<quantity id="4C91E4DBFFBF3A129680F88CFBB8324C" box="[995,1082,1835,1861]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="235" unit="mm" value="30.0">30 mm</quantity>
), body cylindrical, worm-like. Colour in living animals and ethanol is similar, uniformly white, with dark brown colour at the anterior and posterior parts (
<figureCitation id="135255BBFFBF3A1295BFF8D4FEE03284" box="[220,354,1907,1933]" captionStart="FIGURE 9" captionStartId="9.[151,250,1679,1703]" captionTargetBox="[257,1329,181,1656]" captionTargetId="figure-17@9.[257,1329,181,1656]" captionTargetPageId="9" captionText="FIGURE 9. Labidodemas javaensis sp. nov. Holotype, MZB Hol.413 (85x30 mm in ethanol). A: mouth; B: dorsal; C: anal papillae; D: ventral; E: tentacles; F: calcareous ring." pageId="8" pageNumber="235">Figs. 9AD</figureCitation>
). Body wall
<quantity id="4C91E4DBFFBF3A12948EF8D4FDCB3284" box="[493,585,1907,1933]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="8" pageNumber="235" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">24 mm</quantity>
thick, firm, smooth to the touch. Mouth terminal, surrounded by 20 dendritic tentacles, yellowish (
<figureCitation id="135255BBFFBF3A1294EBF830FE6032B8" box="[392,482,1943,1969]" captionStart="FIGURE 9" captionStartId="9.[151,250,1679,1703]" captionTargetBox="[257,1329,181,1656]" captionTargetId="figure-17@9.[257,1329,181,1656]" captionTargetPageId="9" captionText="FIGURE 9. Labidodemas javaensis sp. nov. Holotype, MZB Hol.413 (85x30 mm in ethanol). A: mouth; B: dorsal; C: anal papillae; D: ventral; E: tentacles; F: calcareous ring." pageId="8" pageNumber="235">Fig. 9E</figureCitation>
). Anus terminal, encircled by 810 anal papillae, arranged solitary. Dorsal papillae small, yellowish, spread over the surface. Ventral tube feet numerous, yellowish, longer than dorsal papillae,
<quantity id="4C91E4DBFFBF3A129011F81CFF4732F0" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="8" pageNumber="235" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">34 mm</quantity>
, spread in three rows (ambulacral).
</paragraph>
<caption id="DF1619B6FFBE3A1395F4F928FD2A33C5" pageId="9" pageNumber="236" startId="9.[151,250,1679,1703]" targetBox="[257,1329,181,1656]" targetPageId="9" targetType="figure">
<paragraph id="8BD6493EFFBE3A1395F4F928FD2A33C5" blockId="9.[151,1436,1679,1740]" pageId="9" pageNumber="236">
<emphasis id="B91D952CFFBE3A1395F4F928FE9733AE" bold="true" box="[151,277,1679,1703]" pageId="9" pageNumber="236">FIGURE 9.</emphasis>
<taxonomicName id="4C6932BDFFBE3A13947EF937FD8E33AE" authorityName="Setyastuti &amp; Solis-Marin &amp; Lüter" authorityYear="2024" box="[285,524,1680,1704]" class="Holothuroidea" family="Holothuriidae" genus="Labidodemas" kingdom="Animalia" order="Holothuriida" pageId="9" pageNumber="236" phylum="Echinodermata" rank="species" species="javaensis" status="sp. nov.">
<emphasis id="B91D952CFFBE3A13947EF937FD8E33AE" box="[285,524,1680,1704]" italics="true" pageId="9" pageNumber="236">Labidodemas javaensis</emphasis>
</taxonomicName>
<emphasis id="B91D952CFFBE3A139776F937FDE533A1" bold="true" box="[533,615,1680,1704]" pageId="9" pageNumber="236">
<taxonomicNameLabel id="A22E2857FFBE3A139776F937FDE533A1" box="[533,615,1680,1704]" pageId="9" pageNumber="236" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype, MZB Hol.413 (85x30 mm in ethanol). A: mouth; B: dorsal; C: anal papillae; D: ventral; E: tentacles; F: calcareous ring.
</paragraph>
</caption>
<paragraph id="8BD6493EFFBE3A1395A4F8B9FEDC32E5" blockId="9.[151,1437,1821,2028]" pageId="9" pageNumber="236">
<emphasis id="B91D952CFFBE3A1395A4F8B9FDD43231" bold="true" box="[199,598,1821,1848]" pageId="9" pageNumber="236">
Internal morphology of
<typeStatus id="54D2F79CFFBE3A13948BF8B9FDD33231" box="[488,593,1822,1848]" pageId="9" pageNumber="236" type="holotype">holotype</typeStatus>
.
</emphasis>
Calcareous ring composed of huge quadrangular radial pieces and slender interradial pieces, posterior margin of radial pieces pronounced concave, anterior side with deep median notch (
<figureCitation id="135255BBFFBE3A1395FCF8C1FF743289" box="[159,246,1894,1920]" captionStart="FIGURE 9" captionStartId="9.[151,250,1679,1703]" captionTargetBox="[257,1329,181,1656]" captionTargetId="figure-17@9.[257,1329,181,1656]" captionTargetPageId="9" captionText="FIGURE 9. Labidodemas javaensis sp. nov. Holotype, MZB Hol.413 (85x30 mm in ethanol). A: mouth; B: dorsal; C: anal papillae; D: ventral; E: tentacles; F: calcareous ring." pageId="9" pageNumber="236">Fig. 9F</figureCitation>
). Radial pieces three times wider and higher than interradial pieces (radial pieces
<quantity id="4C91E4DBFFBE3A1391FCF8C1FB603289" box="[1183,1250,1894,1920]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="236" unit="mm" value="3.0">3 mm</quantity>
wide and high). Cuvierian tubules absent. Tentacle ampullae present,
<quantity id="4C91E4DBFFBE3A139797F82DFCD032AD" box="[756,850,1930,1957]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="9" pageNumber="236" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">45 mm</quantity>
long. Single Polian vesicle, tube-like,
<quantity id="4C91E4DBFFBE3A139068F82DFAD932AD" box="[1291,1371,1930,1956]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="236" unit="mm" value="10.0">10 mm</quantity>
long. Gonad one tuft, branched. Respiratory trees arise from a single stalk. Longitudinal muscles medially attachment, margins free, flat.
</paragraph>
<caption id="DF1619B6FFBD3A1095F4F936FBF533C4" pageId="10" pageNumber="237" startId="10.[151,250,1681,1705]" targetBox="[257,1329,181,1657]" targetPageId="10" targetType="figure">
<paragraph id="8BD6493EFFBD3A1095F4F936FBF533C4" blockId="10.[151,1436,1681,1741]" pageId="10" pageNumber="237">
<emphasis id="B91D952CFFBD3A1095F4F936FEA333A0" bold="true" box="[151,289,1681,1705]" pageId="10" pageNumber="237">FIGURE 10.</emphasis>
<taxonomicName id="4C6932BDFFBD3A10944BF936FD9433A0" authorityName="Setyastuti &amp; Solis-Marin &amp; Lüter" authorityYear="2024" box="[296,534,1681,1705]" class="Holothuroidea" family="Holothuriidae" genus="Labidodemas" kingdom="Animalia" order="Holothuriida" pageId="10" pageNumber="237" phylum="Echinodermata" rank="species" species="javaensis" status="sp. nov.">
<emphasis id="B91D952CFFBD3A10944BF936FD9433A0" box="[296,534,1681,1705]" italics="true" pageId="10" pageNumber="237">Labidodemas javaensis</emphasis>
</taxonomicName>
<emphasis id="B91D952CFFBD3A10977EF936FDED33A0" bold="true" box="[541,623,1681,1705]" pageId="10" pageNumber="237">
<taxonomicNameLabel id="A22E2857FFBD3A10977EF936FDED33A0" box="[541,623,1681,1705]" pageId="10" pageNumber="237" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype, MZB Hol.413. Ossicles of dorsal body wall. A: tables; B: tri-radiate rods; C: regular rods; D: rods with perforations; E: pseudo-buttons. F: tables of ventral body wall.
</paragraph>
</caption>
<paragraph id="8BD6493EFFBD3A1195A4F8B8FAA135F0" blockId="10.[151,1437,1823,2030]" lastBlockId="11.[151,1436,151,249]" lastPageId="11" lastPageNumber="238" pageId="10" pageNumber="237">
<emphasis id="B91D952CFFBD3A1095A4F8B8FE3B3230" bold="true" box="[199,441,1823,1849]" pageId="10" pageNumber="237">
Ossicles of
<typeStatus id="54D2F79CFFBD3A109428F8B8FE363230" box="[331,436,1823,1849]" pageId="10" pageNumber="237" type="holotype">holotype</typeStatus>
.
</emphasis>
Dorsal body wall and dorsal papillae hold tables, rods and pseudo-buttons (
<figureCitation id="135255BBFFBD3A10906BF8B8FAE73230" box="[1288,1381,1823,1849]" captionStart="FIGURE 10" captionStartId="10.[151,250,1681,1705]" captionTargetBox="[257,1329,181,1657]" captionTargetId="figure-25@10.[257,1329,181,1657]" captionTargetPageId="10" captionText="FIGURE 10. Labidodemas javaensis sp. nov. Holotype, MZB Hol.413. Ossicles of dorsal body wall. A: tables; B: tri-radiate rods; C: regular rods; D: rods with perforations; E: pseudo-buttons. F: tables of ventral body wall." pageId="10" pageNumber="237">Figs. 10</figureCitation>
,
<figureCitation id="135255BBFFBD3A109012F8B8FA0C3230" box="[1393,1422,1823,1849]" captionStart="FIGURE 11" captionStartId="11.[151,250,1483,1507]" captionTargetBox="[257,1329,273,1460]" captionTargetId="figure-67@11.[257,1329,273,1460]" captionTargetPageId="11" captionText="FIGURE 11. Labidodemas javaensis sp. nov. Holotype. MZB Hol.413. Ossicles of dorsal papillae.A: tables; B: smaller tables with spinous disc and crown; C: pseudo-buttons; D: rods with rugose surface; E: rods with smooth surface." pageId="10" pageNumber="237">11</figureCitation>
). Table discs rounded with spiny edges, usually perforated by four central holes, occasionally up to six central holes; rarely with peripheral holes, but if present, usually 24 holes; four short pillars united by a single cross beam, ending in a regular or irregular crown with 45 huge spines, usually with distal bifurcation, the crown of spines always wider than disc; disc 4070 µm across (
<figureCitation id="135255BBFFBD3A10972EF808FD3F32C0" box="[589,701,1967,1993]" captionStart="FIGURE 10" captionStartId="10.[151,250,1681,1705]" captionTargetBox="[257,1329,181,1657]" captionTargetId="figure-25@10.[257,1329,181,1657]" captionTargetPageId="10" captionText="FIGURE 10. Labidodemas javaensis sp. nov. Holotype, MZB Hol.413. Ossicles of dorsal body wall. A: tables; B: tri-radiate rods; C: regular rods; D: rods with perforations; E: pseudo-buttons. F: tables of ventral body wall." pageId="10" pageNumber="237">Figs. 10A</figureCitation>
,
<figureCitation id="135255BBFFBD3A1097ABF808FD7932C0" box="[712,763,1967,1993]" captionStart="FIGURE 11" captionStartId="11.[151,250,1483,1507]" captionTargetBox="[257,1329,273,1460]" captionTargetId="figure-67@11.[257,1329,273,1460]" captionTargetPageId="11" captionText="FIGURE 11. Labidodemas javaensis sp. nov. Holotype. MZB Hol.413. Ossicles of dorsal papillae.A: tables; B: smaller tables with spinous disc and crown; C: pseudo-buttons; D: rods with rugose surface; E: rods with smooth surface." pageId="10" pageNumber="237">11A</figureCitation>
). In dorsal papillae also present tables with very spinous disc rims, mostly with reduced pillars and ending in reduced crowns with numerous small spines, discs 4055 µm across (
<figureCitation id="135255BBFFBC3A1195FCFF30FE8435B8" box="[159,262,151,177]" captionStart="FIGURE 11" captionStartId="11.[151,250,1483,1507]" captionTargetBox="[257,1329,273,1460]" captionTargetId="figure-67@11.[257,1329,273,1460]" captionTargetPageId="11" captionText="FIGURE 11. Labidodemas javaensis sp. nov. Holotype. MZB Hol.413. Ossicles of dorsal papillae.A: tables; B: smaller tables with spinous disc and crown; C: pseudo-buttons; D: rods with rugose surface; E: rods with smooth surface." pageId="11" pageNumber="238">Fig. 11B</figureCitation>
). Rods observed in several shapes, smooth tri-radiate rods, 60100 µm long (
<figureCitation id="135255BBFFBC3A119117FF30FB5935B8" box="[1140,1243,151,177]" captionStart="FIGURE 10" captionStartId="10.[151,250,1681,1705]" captionTargetBox="[257,1329,181,1657]" captionTargetId="figure-25@10.[257,1329,181,1657]" captionTargetPageId="10" captionText="FIGURE 10. Labidodemas javaensis sp. nov. Holotype, MZB Hol.413. Ossicles of dorsal body wall. A: tables; B: tri-radiate rods; C: regular rods; D: rods with perforations; E: pseudo-buttons. F: tables of ventral body wall." pageId="11" pageNumber="238">Fig. 10B</figureCitation>
); smooth with or without perforations, 40100 µm long (
<figureCitation id="135255BBFFBC3A119734FF1CFD6935DC" box="[599,747,187,213]" captionStart="FIGURE 10" captionStartId="10.[151,250,1681,1705]" captionTargetBox="[257,1329,181,1657]" captionTargetId="figure-25@10.[257,1329,181,1657]" captionTargetPageId="10" captionText="FIGURE 10. Labidodemas javaensis sp. nov. Holotype, MZB Hol.413. Ossicles of dorsal body wall. A: tables; B: tri-radiate rods; C: regular rods; D: rods with perforations; E: pseudo-buttons. F: tables of ventral body wall." pageId="11" pageNumber="238">Figs. 10CD</figureCitation>
,
<figureCitation id="135255BBFFBC3A119794FF1CFCA535DC" box="[759,807,187,213]" captionStart="FIGURE 11" captionStartId="11.[151,250,1483,1507]" captionTargetBox="[257,1329,273,1460]" captionTargetId="figure-67@11.[257,1329,273,1460]" captionTargetPageId="11" captionText="FIGURE 11. Labidodemas javaensis sp. nov. Holotype. MZB Hol.413. Ossicles of dorsal papillae.A: tables; B: smaller tables with spinous disc and crown; C: pseudo-buttons; D: rods with rugose surface; E: rods with smooth surface." pageId="11" pageNumber="238">11E</figureCitation>
); rugose surface rods only found in papillae, straight or tri-radiate, 5070 µm long (
<figureCitation id="135255BBFFBC3A1194ACFF78FDB735F0" box="[463,565,223,249]" captionStart="FIGURE 11" captionStartId="11.[151,250,1483,1507]" captionTargetBox="[257,1329,273,1460]" captionTargetId="figure-67@11.[257,1329,273,1460]" captionTargetPageId="11" captionText="FIGURE 11. Labidodemas javaensis sp. nov. Holotype. MZB Hol.413. Ossicles of dorsal papillae.A: tables; B: smaller tables with spinous disc and crown; C: pseudo-buttons; D: rods with rugose surface; E: rods with smooth surface." pageId="11" pageNumber="238">Fig. 11D</figureCitation>
). Pseudo-buttons rarely found, 100110 µm long (
<figureCitation id="135255BBFFBC3A119108FF78FB5B35F0" box="[1131,1241,223,249]" captionStart="FIGURE 10" captionStartId="10.[151,250,1681,1705]" captionTargetBox="[257,1329,181,1657]" captionTargetId="figure-25@10.[257,1329,181,1657]" captionTargetPageId="10" captionText="FIGURE 10. Labidodemas javaensis sp. nov. Holotype, MZB Hol.413. Ossicles of dorsal body wall. A: tables; B: tri-radiate rods; C: regular rods; D: rods with perforations; E: pseudo-buttons. F: tables of ventral body wall." pageId="11" pageNumber="238">Figs. 10E</figureCitation>
,
<figureCitation id="135255BBFFBC3A119186FF78FA9435F0" box="[1253,1302,223,249]" captionStart="FIGURE 11" captionStartId="11.[151,250,1483,1507]" captionTargetBox="[257,1329,273,1460]" captionTargetId="figure-67@11.[257,1329,273,1460]" captionTargetPageId="11" captionText="FIGURE 11. Labidodemas javaensis sp. nov. Holotype. MZB Hol.413. Ossicles of dorsal papillae.A: tables; B: smaller tables with spinous disc and crown; C: pseudo-buttons; D: rods with rugose surface; E: rods with smooth surface." pageId="11" pageNumber="238">11C</figureCitation>
).
</paragraph>
<caption id="DF1619B6FFBC3A1195F4FA6CFB573301" pageId="11" pageNumber="238" startId="11.[151,250,1483,1507]" targetBox="[257,1329,273,1460]" targetPageId="11" targetType="figure">
<paragraph id="8BD6493EFFBC3A1195F4FA6CFB573301" blockId="11.[151,1436,1483,1544]" pageId="11" pageNumber="238">
<emphasis id="B91D952CFFBC3A1195F4FA6CFE9C30EA" bold="true" box="[151,286,1483,1507]" pageId="11" pageNumber="238">FIGURE 11.</emphasis>
<taxonomicName id="4C6932BDFFBC3A119447FA6BFD9330EA" authorityName="Setyastuti &amp; Solis-Marin &amp; Lüter" authorityYear="2024" box="[292,529,1484,1508]" class="Holothuroidea" family="Holothuriidae" genus="Labidodemas" kingdom="Animalia" order="Holothuriida" pageId="11" pageNumber="238" phylum="Echinodermata" rank="species" species="javaensis" status="sp. nov.">
<emphasis id="B91D952CFFBC3A119447FA6BFD9330EA" box="[292,529,1484,1508]" italics="true" pageId="11" pageNumber="238">Labidodemas javaensis</emphasis>
</taxonomicName>
<emphasis id="B91D952CFFBC3A119774FA6BFDEA30ED" bold="true" box="[535,616,1484,1508]" pageId="11" pageNumber="238">
<taxonomicNameLabel id="A22E2857FFBC3A119774FA6BFDEA30ED" box="[535,616,1484,1508]" pageId="11" pageNumber="238" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype. MZB Hol.413. Ossicles of dorsal papillae. A: tables; B: smaller tables with spinous disc and crown; C: pseudo-buttons; D: rods with rugose surface; E: rods with smooth surface.
</paragraph>
</caption>
<paragraph id="8BD6493EFFBC3A1195A4F9FDFB7333E9" blockId="11.[151,1437,1626,1976]" pageId="11" pageNumber="238">
Ventral body wall only present regular tables, discs 6580 µm across (
<figureCitation id="135255BBFFBC3A119684F9FDFBCE337D" box="[999,1100,1626,1652]" captionStart="FIGURE 10" captionStartId="10.[151,250,1681,1705]" captionTargetBox="[257,1329,181,1657]" captionTargetId="figure-25@10.[257,1329,181,1657]" captionTargetPageId="10" captionText="FIGURE 10. Labidodemas javaensis sp. nov. Holotype, MZB Hol.413. Ossicles of dorsal body wall. A: tables; B: tri-radiate rods; C: regular rods; D: rods with perforations; E: pseudo-buttons. F: tables of ventral body wall." pageId="11" pageNumber="238">Fig. 10F</figureCitation>
). Ventral tube feet with a few regular tables, discs 50100 µm across (
<figureCitation id="135255BBFFBC3A119705F9D9FD533391" box="[614,721,1662,1688]" captionStart="FIGURE 12" captionStartId="12.[151,250,1143,1167]" captionTargetBox="[257,1329,181,1119]" captionTargetId="figure-25@12.[257,1329,181,1119]" captionTargetPageId="12" captionText="FIGURE 12. Labidodemas javaensis sp. nov. Holotype, MZB Hol.413. Ossicles of ventral tube feet. A: tables; B: tables with reduce pillar and crowns; C: pseudo-buttons; D: rod; E: large perforation endplate." pageId="11" pageNumber="238">Fig. 12A</figureCitation>
), and very numerous tables similar to those in dorsal papillae, 4060 µm across (
<figureCitation id="135255BBFFBC3A119408F905FE5133B5" box="[363,467,1698,1724]" captionStart="FIGURE 12" captionStartId="12.[151,250,1143,1167]" captionTargetBox="[257,1329,181,1119]" captionTargetId="figure-25@12.[257,1329,181,1119]" captionTargetPageId="12" captionText="FIGURE 12. Labidodemas javaensis sp. nov. Holotype, MZB Hol.413. Ossicles of ventral tube feet. A: tables; B: tables with reduce pillar and crowns; C: pseudo-buttons; D: rod; E: large perforation endplate." pageId="11" pageNumber="238">Fig. 12B</figureCitation>
); large rods with smooth surface very rare, 180 µm long (
<figureCitation id="135255BBFFBC3A11910AF905FB5633B5" box="[1129,1236,1698,1724]" captionStart="FIGURE 12" captionStartId="12.[151,250,1143,1167]" captionTargetBox="[257,1329,181,1119]" captionTargetId="figure-25@12.[257,1329,181,1119]" captionTargetPageId="12" captionText="FIGURE 12. Labidodemas javaensis sp. nov. Holotype, MZB Hol.413. Ossicles of ventral tube feet. A: tables; B: tables with reduce pillar and crowns; C: pseudo-buttons; D: rod; E: large perforation endplate." pageId="11" pageNumber="238">Fig. 12D</figureCitation>
); large perforated plates many, 400450 µm across (
<figureCitation id="135255BBFFBC3A119776F961FDF933E9" box="[533,635,1734,1760]" captionStart="FIGURE 12" captionStartId="12.[151,250,1143,1167]" captionTargetBox="[257,1329,181,1119]" captionTargetId="figure-25@12.[257,1329,181,1119]" captionTargetPageId="12" captionText="FIGURE 12. Labidodemas javaensis sp. nov. Holotype, MZB Hol.413. Ossicles of ventral tube feet. A: tables; B: tables with reduce pillar and crowns; C: pseudo-buttons; D: rod; E: large perforation endplate." pageId="11" pageNumber="238">Fig. 12E</figureCitation>
). pseudo-buttons only a few, 6065 µm long (
<figureCitation id="135255BBFFBC3A11911EF961FB6633E9" box="[1149,1252,1734,1760]" captionStart="FIGURE 12" captionStartId="12.[151,250,1143,1167]" captionTargetBox="[257,1329,181,1119]" captionTargetId="figure-25@12.[257,1329,181,1119]" captionTargetPageId="12" captionText="FIGURE 12. Labidodemas javaensis sp. nov. Holotype, MZB Hol.413. Ossicles of ventral tube feet. A: tables; B: tables with reduce pillar and crowns; C: pseudo-buttons; D: rod; E: large perforation endplate." pageId="11" pageNumber="238">Fig. 12C</figureCitation>
).
</paragraph>
<paragraph id="8BD6493EFFBC3A1195A4F94DFF7432B1" blockId="11.[151,1437,1626,1976]" pageId="11" pageNumber="238">
Tentacles only present rods, straight or slightly curved, smooth or rugose surface with blunt spines, 3550 µm long (
<figureCitation id="135255BBFFBC3A11946BF8A9FEF13221" box="[264,371,1806,1832]" captionStart="FIGURE 13" captionStartId="14.[151,250,1716,1740]" captionTargetBox="[257,1329,181,1693]" captionTargetId="figure-25@14.[257,1329,181,1693]" captionTargetPageId="14" captionText="FIGURE 13. Labidodemas javaensis sp. nov. Holotype. MZB Hol.413. A: tentacle rods. Ossicles of anal papillae, B: tables; C: tri-radiate rods; D: rods with smooth surface; E: rods with rugose surface; F: rods rugose surface with perforations." pageId="11" pageNumber="238">Fig. 13A</figureCitation>
). Anal papillae always with tables disc with reduced pillars and crowns, spiny rims and more squarish, 2545 µm across (
<figureCitation id="135255BBFFBC3A1194B2F895FDB43245" box="[465,566,1842,1868]" captionStart="FIGURE 13" captionStartId="14.[151,250,1716,1740]" captionTargetBox="[257,1329,181,1693]" captionTargetId="figure-25@14.[257,1329,181,1693]" captionTargetPageId="14" captionText="FIGURE 13. Labidodemas javaensis sp. nov. Holotype. MZB Hol.413. A: tentacle rods. Ossicles of anal papillae, B: tables; C: tri-radiate rods; D: rods with smooth surface; E: rods with rugose surface; F: rods rugose surface with perforations." pageId="11" pageNumber="238">Fig. 13B</figureCitation>
); tri-radiate rods with spiny surfaces, 4060 µm long (
<figureCitation id="135255BBFFBC3A1191F4F895FB7F3245" box="[1175,1277,1842,1868]" captionStart="FIGURE 13" captionStartId="14.[151,250,1716,1740]" captionTargetBox="[257,1329,181,1693]" captionTargetId="figure-25@14.[257,1329,181,1693]" captionTargetPageId="14" captionText="FIGURE 13. Labidodemas javaensis sp. nov. Holotype. MZB Hol.413. A: tentacle rods. Ossicles of anal papillae, B: tables; C: tri-radiate rods; D: rods with smooth surface; E: rods with rugose surface; F: rods rugose surface with perforations." pageId="11" pageNumber="238">Fig. 13C</figureCitation>
); straight rods with smooth surface, 4560 µm long (
<figureCitation id="135255BBFFBC3A119728F8F1FD333279" box="[587,689,1878,1904]" captionStart="FIGURE 13" captionStartId="14.[151,250,1716,1740]" captionTargetBox="[257,1329,181,1693]" captionTargetId="figure-25@14.[257,1329,181,1693]" captionTargetPageId="14" captionText="FIGURE 13. Labidodemas javaensis sp. nov. Holotype. MZB Hol.413. A: tentacle rods. Ossicles of anal papillae, B: tables; C: tri-radiate rods; D: rods with smooth surface; E: rods with rugose surface; F: rods rugose surface with perforations." pageId="11" pageNumber="238">Fig. 13E</figureCitation>
); slightly curved and straight rods with spiny surfaces, 4055 µm long (
<figureCitation id="135255BBFFBC3A1195B8F8DDFEC4329D" box="[219,326,1914,1940]" captionStart="FIGURE 13" captionStartId="14.[151,250,1716,1740]" captionTargetBox="[257,1329,181,1693]" captionTargetId="figure-25@14.[257,1329,181,1693]" captionTargetPageId="14" captionText="FIGURE 13. Labidodemas javaensis sp. nov. Holotype. MZB Hol.413. A: tentacle rods. Ossicles of anal papillae, B: tables; C: tri-radiate rods; D: rods with smooth surface; E: rods with rugose surface; F: rods rugose surface with perforations." pageId="11" pageNumber="238">Fig. 13D</figureCitation>
); spiny surface rods with perforations, 4555 µm long (
<figureCitation id="135255BBFFBC3A1196B0F8DDFBB8329D" box="[979,1082,1914,1940]" captionStart="FIGURE 13" captionStartId="14.[151,250,1716,1740]" captionTargetBox="[257,1329,181,1693]" captionTargetId="figure-25@14.[257,1329,181,1693]" captionTargetPageId="14" captionText="FIGURE 13. Labidodemas javaensis sp. nov. Holotype. MZB Hol.413. A: tentacle rods. Ossicles of anal papillae, B: tables; C: tri-radiate rods; D: rods with smooth surface; E: rods with rugose surface; F: rods rugose surface with perforations." pageId="11" pageNumber="238">Fig. 13F</figureCitation>
). Longitudinal muscles devoid ossicles.
</paragraph>
</subSubSection>
<caption id="DF1619B6FFBB3A1695F4FBD0FC5F31BA" pageId="12" pageNumber="239" startId="12.[151,250,1143,1167]" targetBox="[257,1329,181,1119]" targetPageId="12" targetType="figure">
<paragraph id="8BD6493EFFBB3A1695F4FBD0FC5F31BA" blockId="12.[151,1436,1143,1203]" pageId="12" pageNumber="239">
<emphasis id="B91D952CFFBB3A1695F4FBD0FEA23186" bold="true" box="[151,288,1143,1167]" pageId="12" pageNumber="239">FIGURE 12.</emphasis>
<taxonomicName id="4C6932BDFFBB3A169444FBD0FD963186" authorityName="Setyastuti &amp; Solis-Marin &amp; Lüter" authorityYear="2024" box="[295,532,1143,1167]" class="Holothuroidea" family="Holothuriidae" genus="Labidodemas" kingdom="Animalia" order="Holothuriida" pageId="12" pageNumber="239" phylum="Echinodermata" rank="species" species="javaensis" status="sp. nov.">
<emphasis id="B91D952CFFBB3A169444FBD0FD963186" box="[295,532,1143,1167]" italics="true" pageId="12" pageNumber="239">Labidodemas javaensis</emphasis>
</taxonomicName>
<emphasis id="B91D952CFFBB3A16977FFBD0FDEF3186" bold="true" box="[540,621,1143,1167]" pageId="12" pageNumber="239">
<taxonomicNameLabel id="A22E2857FFBB3A16977FFBD0FDEF3186" box="[540,621,1143,1167]" pageId="12" pageNumber="239" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype, MZB Hol.413. Ossicles of ventral tube feet. A: tables; B: tables with reduce pillar and crowns; C: pseudo-buttons; D: rod; E: large perforation endplate.
</paragraph>
</caption>
<caption id="DF1619B6FFBB3A1695F4FAA0FB35304A" pageId="12" pageNumber="239" startId="12.[151,239,1287,1311]" targetType="table">
<paragraph id="8BD6493EFFBB3A1695F4FAA0FB35304A" blockId="12.[151,1436,1287,1347]" pageId="12" pageNumber="239">
<emphasis id="B91D952CFFBB3A1695F4FAA0FE893016" bold="true" box="[151,267,1287,1311]" pageId="12" pageNumber="239">TABLE 1.</emphasis>
Characters differentiate
<taxonomicName id="4C6932BDFFBB3A169775FAA0FC933016" authorityName="Setyastuti &amp; Solis-Marin &amp; Lüter" authorityYear="2024" box="[534,785,1287,1311]" class="Holothuroidea" family="Holothuriidae" genus="Labidodemas" kingdom="Animalia" order="Holothuriida" pageId="12" pageNumber="239" phylum="Echinodermata" rank="species" species="javaensis" status="sp. nov.">
<emphasis id="B91D952CFFBB3A169775FAA0FC933016" box="[534,785,1287,1311]" italics="true" pageId="12" pageNumber="239">Labidodemas javaensis</emphasis>
</taxonomicName>
<emphasis id="B91D952CFFBB3A16967AFAA0FCED3016" bold="true" box="[793,879,1287,1311]" pageId="12" pageNumber="239">
<taxonomicNameLabel id="A22E2857FFBB3A16967AFAA0FCED3016" box="[793,879,1287,1311]" pageId="12" pageNumber="239" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from
<taxonomicName id="4C6932BDFFBB3A1696D1FAAFFF4D304A" authority="Massin, Samyn &amp; Thandar 2004" authorityName="Massin, Samyn &amp; Thandar" authorityYear="2004" class="Holothuroidea" family="Holothuriidae" genus="Labidodemas" kingdom="Animalia" order="Holothuriida" pageId="12" pageNumber="239" phylum="Echinodermata" rank="species" species="quadripartitum">
<emphasis id="B91D952CFFBB3A1696D1FAAFFBF03016" box="[946,1138,1287,1311]" italics="true" pageId="12" pageNumber="239">L. quadripartitum</emphasis>
<bibRefCitation id="EFF834CFFFBB3A169119FAA0FF4D304A" author="Massin, C. &amp; Samyn, Y. &amp; Thandar, A. S." pageId="12" pageNumber="239" pagination="1811 - 1847" refId="ref8451" refString="Massin, C., Samyn, Y. &amp; Thandar, A. S. (2004) The genus Labidodemas (Holothuroidea: Aspidochirotida) revisited, with description of three new species and with re-positioning of Holothuria (Irenothuria) maccullochi Deichmann, 1958. Journal of Natural History, 38, 1811 - 1847. https: // doi. org / 10.1080 / 0022293031000156268" type="journal article" year="2004">Massin, Samyn &amp; Thandar 2004</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C6932BDFFBB3A1695B9FA8BFD9D304A" authority="Selenka, 1867" authorityName="Selenka" authorityYear="1867" box="[218,543,1323,1347]" class="Holothuroidea" family="Holothuriidae" genus="Labidodemas" kingdom="Animalia" order="Holothuriida" pageId="12" pageNumber="239" phylum="Echinodermata" rank="species" species="semperianum">
<emphasis id="B91D952CFFBB3A1695B9FA8BFEFD304A" box="[218,383,1324,1347]" italics="true" pageId="12" pageNumber="239">L semperianum</emphasis>
<bibRefCitation id="EFF834CFFFBB3A1694E6FA8CFD9D304A" author="Selenka, E." box="[389,543,1323,1347]" pageId="12" pageNumber="239" pagination="291 - 374" refId="ref8895" refString="Selenka, E. (1867) Beitrage zur anatomie und systematik der holothurien. Zeitschrift fur wissenschaftliche Zoologie, 17 (2), 291 - 374." type="journal article" year="1867">Selenka, 1867</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C6932BDFFBB3A169734FA8BFB31304A" authority="Massin, Samyn &amp; Thandar 2004" authorityName="Massin, Samyn &amp; Thandar" authorityYear="2004" box="[599,1203,1323,1347]" class="Holothuroidea" family="Holothuriidae" genus="Labidodemas" kingdom="Animalia" order="Holothuriida" pageId="12" pageNumber="239" phylum="Echinodermata" rank="species" species="pseudosemperianum">
<emphasis id="B91D952CFFBB3A169734FA8BFCCC304A" box="[599,846,1323,1347]" italics="true" pageId="12" pageNumber="239">L. pseudosemperianum</emphasis>
<bibRefCitation id="EFF834CFFFBB3A169636FA8CFB31304A" author="Massin, C. &amp; Samyn, Y. &amp; Thandar, A. S." box="[853,1203,1323,1347]" pageId="12" pageNumber="239" pagination="1811 - 1847" refId="ref8451" refString="Massin, C., Samyn, Y. &amp; Thandar, A. S. (2004) The genus Labidodemas (Holothuroidea: Aspidochirotida) revisited, with description of three new species and with re-positioning of Holothuria (Irenothuria) maccullochi Deichmann, 1958. Journal of Natural History, 38, 1811 - 1847. https: // doi. org / 10.1080 / 0022293031000156268" type="journal article" year="2004">Massin, Samyn &amp; Thandar 2004</bibRefCitation>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BD6493EFFBB3A1695C0FAF3FB5D3257" pageId="12" pageNumber="239">
<table id="F969BB9EFFBBC5E595C0FAF3FA053257" box="[163,1415,1364,1886]" colsContinueIn="13.[163,1419,202,1467]" gridcols="5" gridrows="17" pageId="12" pageNumber="239">
<tr id="35594B7CFFBBC5E595C0FAF3FA053063" box="[163,1415,1364,1386]" gridrow="0" pageId="12" pageNumber="239">
<th id="76882200FFBBC5E595C0FAF3FEC73063" box="[163,325,1364,1386]" gridcol="0" gridrow="0" pageId="12" pageNumber="239">Character</th>
<th id="76882200FFBBC5E59400FAF3FDC73063" box="[355,581,1364,1386]" gridcol="1" gridrow="0" pageId="12" pageNumber="239">
<taxonomicName id="4C6932BDFFBB3A169400FAF2FE573063" authorityName="Setyastuti &amp; Solis-Marin &amp; Lüter" authorityYear="2024" box="[355,469,1365,1386]" class="Holothuroidea" family="Holothuriidae" genus="Labidodemas" kingdom="Animalia" order="Holothuriida" pageId="12" pageNumber="239" phylum="Echinodermata" rank="species" species="javaensis" status="sp. nov.">
<emphasis id="B91D952CFFBB3A169400FAF2FE573063" box="[355,469,1365,1386]" italics="true" pageId="12" pageNumber="239">L. javaensis</emphasis>
</taxonomicName>
<emphasis id="B91D952CFFBB3A1694BFFAF2FDAA3063" bold="true" box="[476,552,1365,1386]" pageId="12" pageNumber="239">
<taxonomicNameLabel id="A22E2857FFBB3A1694BFFAF2FDAA3063" box="[476,552,1365,1386]" pageId="12" pageNumber="239" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<th id="76882200FFBBC5E5970EFAF3FCE83063" box="[621,874,1364,1386]" gridcol="2" gridrow="0" pageId="12" pageNumber="239">
<taxonomicName id="4C6932BDFFBB3A16970EFAF2FCE43063" authority="Massin" authorityName="Massin, Samyn &amp; Thandar" authorityYear="2004" box="[621,870,1364,1386]" class="Holothuroidea" family="Holothuriidae" genus="Labidodemas" kingdom="Animalia" order="Holothuriida" pageId="12" pageNumber="239" phylum="Echinodermata" rank="species" species="quadripartitum">
<emphasis id="B91D952CFFBB3A16970EFAF2FC9B3060" box="[621,793,1364,1386]" italics="true" pageId="12" pageNumber="239">L. quadripartitum</emphasis>
Massin
</taxonomicName>
,
</th>
<th id="76882200FFBBC5E596E3FAF3FB033063" box="[896,1153,1364,1386]" gridcol="3" gridrow="0" pageId="12" pageNumber="239">
<taxonomicName id="4C6932BDFFBB3A1696E3FAF2FBED3063" authority="Selenka" authorityName="Selenka" authorityYear="1867" box="[896,1135,1364,1386]" class="Holothuroidea" family="Holothuriidae" genus="Labidodemas" kingdom="Animalia" order="Holothuriida" pageId="12" pageNumber="239" phylum="Echinodermata" rank="species" species="semperianum">
<emphasis id="B91D952CFFBB3A1696E3FAF2FB993063" box="[896,1051,1365,1386]" italics="true" pageId="12" pageNumber="239">L. semperianum</emphasis>
Selenka
</taxonomicName>
,
</th>
<th id="76882200FFBBC5E591C1FAF3FA053063" box="[1186,1415,1364,1386]" gridcol="4" gridrow="0" pageId="12" pageNumber="239">
<taxonomicName id="4C6932BDFFBB3A1691C1FAF2FA033060" authority="Samyn &amp; Thandar, 2004" authorityName="Massin, Samyn &amp; Thandar" authorityYear="2004" box="[1186,1409,1364,1386]" class="Holothuroidea" family="Holothuriidae" genus="Labidodemas" kingdom="Animalia" order="Holothuriida" pageId="12" pageNumber="239" phylum="Echinodermata" rank="species" species="pseudosemperianum">
<emphasis id="B91D952CFFBB3A1691C1FAF2FA033060" box="[1186,1409,1364,1386]" italics="true" pageId="12" pageNumber="239">L. pseudosemperianum</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="35594B7CFFBBC5E595C0FAD5FA053081" box="[163,1415,1394,1416]" gridrow="1" pageId="12" pageNumber="239" rowspan-0="1" rowspan-1="1">
<td id="76882200FFBBC5E5970EFAD5FCE83081" box="[621,874,1394,1416]" gridcol="2" gridrow="1" pageId="12" pageNumber="239">Samyn &amp; Thandar, 2004</td>
<td id="76882200FFBBC5E596E3FAD5FB033081" box="[896,1153,1394,1416]" gridcol="3" gridrow="1" pageId="12" pageNumber="239">
1867 (
<bibRefCitation id="EFF834CFFFBB3A1696DEFAD5FBF1308E" author="Massin, C. &amp; Samyn, Y. &amp; Thandar, A. S." box="[957,1139,1394,1416]" pageId="12" pageNumber="239" pagination="1811 - 1847" refId="ref8451" refString="Massin, C., Samyn, Y. &amp; Thandar, A. S. (2004) The genus Labidodemas (Holothuroidea: Aspidochirotida) revisited, with description of three new species and with re-positioning of Holothuria (Irenothuria) maccullochi Deichmann, 1958. Journal of Natural History, 38, 1811 - 1847. https: // doi. org / 10.1080 / 0022293031000156268" type="journal article" year="2004">
Massin
<emphasis id="B91D952CFFBB3A169169FAD5FBB6308E" box="[1034,1076,1394,1415]" italics="true" pageId="12" pageNumber="239">et al</emphasis>
. 2004
</bibRefCitation>
;
</td>
<td id="76882200FFBBC5E591C1FAD5FA053081" box="[1186,1415,1394,1416]" gridcol="4" gridrow="1" pageId="12" pageNumber="239">
(
<bibRefCitation id="EFF834CFFFBB3A1691CAFAD5FADD308E" author="Massin, C. &amp; Samyn, Y. &amp; Thandar, A. S." box="[1193,1375,1394,1416]" pageId="12" pageNumber="239" pagination="1811 - 1847" refId="ref8451" refString="Massin, C., Samyn, Y. &amp; Thandar, A. S. (2004) The genus Labidodemas (Holothuroidea: Aspidochirotida) revisited, with description of three new species and with re-positioning of Holothuria (Irenothuria) maccullochi Deichmann, 1958. Journal of Natural History, 38, 1811 - 1847. https: // doi. org / 10.1080 / 0022293031000156268" type="journal article" year="2004">
Massin
<emphasis id="B91D952CFFBB3A169195FAD5FAA2308E" box="[1270,1312,1394,1415]" italics="true" pageId="12" pageNumber="239">et al</emphasis>
. 2004
</bibRefCitation>
;
</td>
</tr>
<tr id="35594B7CFFBBC5E595C0FA28FA0530AC" box="[163,1415,1423,1445]" gridrow="2" pageId="12" pageNumber="239" rowspan-0="1" rowspan-1="1">
<td id="76882200FFBBC5E5970EFA28FCE830AC" box="[621,874,1423,1445]" gridcol="2" gridrow="2" pageId="12" pageNumber="239">
(
<bibRefCitation id="EFF834CFFFBB3A169710FA28FCAB30AD" author="Massin, C. &amp; Samyn, Y. &amp; Thandar, A. S." box="[627,809,1423,1445]" pageId="12" pageNumber="239" pagination="1811 - 1847" refId="ref8451" refString="Massin, C., Samyn, Y. &amp; Thandar, A. S. (2004) The genus Labidodemas (Holothuroidea: Aspidochirotida) revisited, with description of three new species and with re-positioning of Holothuria (Irenothuria) maccullochi Deichmann, 1958. Journal of Natural History, 38, 1811 - 1847. https: // doi. org / 10.1080 / 0022293031000156268" type="journal article" year="2004">
Massin
<emphasis id="B91D952CFFBB3A1697A3FA37FD6930AD" box="[704,747,1423,1445]" italics="true" pageId="12" pageNumber="239">et al</emphasis>
. 2004
</bibRefCitation>
;
</td>
<td id="76882200FFBBC5E596E3FA28FB0330AC" box="[896,1153,1423,1445]" gridcol="3" gridrow="2" pageId="12" pageNumber="239">
<bibRefCitation id="EFF834CFFFBB3A1696E3FA28FBB230AC" author="Samyn, Y. &amp; Appeltans, W. &amp; Kerr, A. M." box="[896,1072,1423,1445]" pageId="12" pageNumber="239" pagination="103 - 120" refId="ref8831" refString="Samyn, Y., Appeltans, W. &amp; Kerr, A. M. (2005) Phylogeny of Labidodemas and the Holothuriidae (Holothuroidea: Aspidochirotida) as inferred from morphology. Zoological Journal of the Linnean Society, 144, 103 - 120. https: // doi. org / 10.1111 / j. 1096 - 3642.2005.00158. x" type="journal article" year="2005">
Samyn
<emphasis id="B91D952CFFBB3A1696AAFA37FC7130AD" box="[969,1011,1423,1445]" italics="true" pageId="12" pageNumber="239">et al</emphasis>
. 2005
</bibRefCitation>
)
</td>
<td id="76882200FFBBC5E591C1FA28FA0530AC" box="[1186,1415,1423,1445]" gridcol="4" gridrow="2" pageId="12" pageNumber="239">
<bibRefCitation id="EFF834CFFFBB3A1691C1FA28FAD030AC" author="Samyn, Y. &amp; Appeltans, W. &amp; Kerr, A. M." box="[1186,1362,1423,1445]" pageId="12" pageNumber="239" pagination="103 - 120" refId="ref8831" refString="Samyn, Y., Appeltans, W. &amp; Kerr, A. M. (2005) Phylogeny of Labidodemas and the Holothuriidae (Holothuroidea: Aspidochirotida) as inferred from morphology. Zoological Journal of the Linnean Society, 144, 103 - 120. https: // doi. org / 10.1111 / j. 1096 - 3642.2005.00158. x" type="journal article" year="2005">
Samyn
<emphasis id="B91D952CFFBB3A169188FA37FA9730AD" box="[1259,1301,1423,1445]" italics="true" pageId="12" pageNumber="239">et al</emphasis>
. 2005
</bibRefCitation>
)
</td>
</tr>
<tr id="35594B7CFFBBC5E595C0FA0BFA0530CB" box="[163,1415,1452,1474]" gridrow="3" pageId="12" pageNumber="239" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-4="1">
<td id="76882200FFBBC5E5970EFA0BFCE830CB" box="[621,874,1452,1474]" gridcol="2" gridrow="3" pageId="12" pageNumber="239">
<bibRefCitation id="EFF834CFFFBB3A16970EFA0AFC9F30CB" author="Samyn, Y. &amp; Appeltans, W. &amp; Kerr, A. M." box="[621,797,1452,1474]" pageId="12" pageNumber="239" pagination="103 - 120" refId="ref8831" refString="Samyn, Y., Appeltans, W. &amp; Kerr, A. M. (2005) Phylogeny of Labidodemas and the Holothuriidae (Holothuroidea: Aspidochirotida) as inferred from morphology. Zoological Journal of the Linnean Society, 144, 103 - 120. https: // doi. org / 10.1111 / j. 1096 - 3642.2005.00158. x" type="journal article" year="2005">
Samyn
<emphasis id="B91D952CFFBB3A1697D5FA0AFD6230C8" box="[694,736,1452,1474]" italics="true" pageId="12" pageNumber="239">et al</emphasis>
. 2005
</bibRefCitation>
)
</td>
</tr>
<tr id="35594B7CFFBBC5E595C0FA73FA0530E3" box="[163,1415,1492,1514]" gridrow="4" pageId="12" pageNumber="239">
<th id="76882200FFBBC5E595C0FA73FEC730E3" box="[163,325,1492,1514]" gridcol="0" gridrow="4" pageId="12" pageNumber="239">Body colour in</th>
<td id="76882200FFBBC5E59400FA73FDC730E3" box="[355,581,1492,1514]" gridcol="1" gridrow="4" pageId="12" pageNumber="239">White uniformly,</td>
<td id="76882200FFBBC5E5970EFA73FCE830E3" box="[621,874,1492,1514]" gridcol="2" gridrow="4" pageId="12" pageNumber="239">Dorsally whitish,</td>
<td id="76882200FFBBC5E596E3FA73FB0330E3" box="[896,1153,1492,1514]" gridcol="3" gridrow="4" pageId="12" pageNumber="239">According to the original</td>
<td id="76882200FFBBC5E591C1FA73FA0530E3" box="[1186,1415,1492,1514]" gridcol="4" gridrow="4" pageId="12" pageNumber="239">White-pink with yellow</td>
</tr>
<tr id="35594B7CFFBBC5E595C0FA56FA05330E" box="[163,1415,1521,1543]" gridrow="5" pageId="12" pageNumber="239">
<th id="76882200FFBBC5E595C0FA56FEC7330E" box="[163,325,1521,1543]" gridcol="0" gridrow="5" pageId="12" pageNumber="239">the living animal</th>
<td id="76882200FFBBC5E59400FA56FDC7330E" box="[355,581,1521,1543]" gridcol="1" gridrow="5" pageId="12" pageNumber="239">anterior and posterior</td>
<td id="76882200FFBBC5E5970EFA56FCE8330E" box="[621,874,1521,1543]" gridcol="2" gridrow="5" pageId="12" pageNumber="239">ventrally reddish</td>
<td id="76882200FFBBC5E596E3FA56FB03330E" box="[896,1153,1521,1543]" gridcol="3" gridrow="5" pageId="12" pageNumber="239">description, animal</td>
<td id="76882200FFBBC5E591C1FA56FA05330E" box="[1186,1415,1521,1543]" gridcol="4" gridrow="5" pageId="12" pageNumber="239">tube feet; around the</td>
</tr>
<tr id="35594B7CFFBBC5E595C0F9A9FA05332D" box="[163,1415,1550,1572]" gridrow="6" pageId="12" pageNumber="239" rowspan-0="1" rowspan-2="1">
<td id="76882200FFBBC5E59400F9A9FDC7332D" box="[355,581,1550,1572]" gridcol="1" gridrow="6" pageId="12" pageNumber="239">parts dark brown</td>
<td id="76882200FFBBC5E596E3F9A9FB03332D" box="[896,1153,1550,1572]" gridcol="3" gridrow="6" pageId="12" pageNumber="239">colourless, and brownish</td>
<td id="76882200FFBBC5E591C1F9A9FA05332D" box="[1186,1415,1550,1572]" gridcol="4" gridrow="6" pageId="12" pageNumber="239">mouth colour deep pink</td>
</tr>
<tr id="35594B7CFFBBC5E595C0F98BFA05334B" box="[163,1415,1580,1602]" gridrow="7" pageId="12" pageNumber="239" rowspan-0="1" rowspan-1="1" rowspan-2="1">
<td id="76882200FFBBC5E596E3F98BFB03334B" box="[896,1153,1580,1602]" gridcol="3" gridrow="7" pageId="12" pageNumber="239">at the ends of the body</td>
<td id="76882200FFBBC5E591C1F98BFA05334B" box="[1186,1415,1580,1602]" gridcol="4" gridrow="7" pageId="12" pageNumber="239">to brown</td>
</tr>
<tr id="35594B7CFFBBC5E595C0F9EEFA053356" box="[163,1415,1609,1631]" gridrow="8" pageId="12" pageNumber="239" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-4="1">
<td id="76882200FFBBC5E596E3F9EEFB033356" box="[896,1153,1609,1631]" gridcol="3" gridrow="8" pageId="12" pageNumber="239">(“Thier farbios, an den</td>
</tr>
<tr id="35594B7CFFBBC5E595C0F9C1FA053375" box="[163,1415,1638,1660]" gridrow="9" pageId="12" pageNumber="239" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-4="1">
<td id="76882200FFBBC5E596E3F9C1FB033375" box="[896,1153,1638,1660]" gridcol="3" gridrow="9" pageId="12" pageNumber="239">Körperenden bräunlich”).</td>
</tr>
<tr id="35594B7CFFBBC5E595C0F923FA053393" box="[163,1415,1668,1690]" gridrow="10" pageId="12" pageNumber="239" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-4="1">
<td id="76882200FFBBC5E596E3F923FB033393" box="[896,1153,1668,1690]" gridcol="3" gridrow="10" pageId="12" pageNumber="239">Not available for lectotype</td>
</tr>
<tr id="35594B7CFFBBC5E595C0F906FA0533BE" box="[163,1415,1697,1719]" gridrow="11" pageId="12" pageNumber="239" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-4="1">
<td id="76882200FFBBC5E596E3F906FB0333BE" box="[896,1153,1697,1719]" gridcol="3" gridrow="11" pageId="12" pageNumber="239">and paralectotype</td>
</tr>
<tr id="35594B7CFFBBC5E595C0F96FFA0533D7" box="[163,1415,1736,1758]" gridrow="12" pageId="12" pageNumber="239">
<th id="76882200FFBBC5E595C0F96FFEC733D7" box="[163,325,1736,1758]" gridcol="0" gridrow="12" pageId="12" pageNumber="239">Tentacle</th>
<td id="76882200FFBBC5E59400F96FFDC733D7" box="[355,581,1736,1758]" gridcol="1" gridrow="12" pageId="12" pageNumber="239">20, dendritic, yellowish</td>
<td id="76882200FFBBC5E5970EF96FFCE833D7" box="[621,874,1736,1758]" gridcol="2" gridrow="12" pageId="12" pageNumber="239">Tentacles retracted;</td>
<td id="76882200FFBBC5E596E3F96FFB0333D7" box="[896,1153,1736,1758]" gridcol="3" gridrow="12" pageId="12" pageNumber="239">12 in lectotype (but</td>
<td id="76882200FFBBC5E591C1F96FFA0533D7" box="[1186,1415,1736,1758]" gridcol="4" gridrow="12" pageId="12" pageNumber="239">20</td>
</tr>
<tr id="35594B7CFFBBC5E595C0F941FA0533F5" box="[163,1415,1766,1788]" gridrow="13" pageId="12" pageNumber="239" rowspan-0="1" rowspan-1="1" rowspan-4="1">
<td id="76882200FFBBC5E5970EF941FCE833F5" box="[621,874,1766,1788]" gridcol="2" gridrow="13" pageId="12" pageNumber="239">number could not be</td>
<td id="76882200FFBBC5E596E3F941FB0333F5" box="[896,1153,1766,1788]" gridcol="3" gridrow="13" pageId="12" pageNumber="239">probably the number</td>
</tr>
<tr id="35594B7CFFBBC5E595C0F8A4FA053210" box="[163,1415,1795,1817]" gridrow="14" pageId="12" pageNumber="239" rowspan-0="1" rowspan-1="1" rowspan-4="1">
<td id="76882200FFBBC5E5970EF8A4FCE83210" box="[621,874,1795,1817]" gridcol="2" gridrow="14" pageId="12" pageNumber="239">determined</td>
<td id="76882200FFBBC5E596E3F8A4FB033210" box="[896,1153,1795,1817]" gridcol="3" gridrow="14" pageId="12" pageNumber="239">is 20 as in all the other</td>
</tr>
<tr id="35594B7CFFBBC5E595C0F887FA05323F" box="[163,1415,1824,1846]" gridrow="15" pageId="12" pageNumber="239" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-4="1">
<td id="76882200FFBBC5E596E3F887FB03323F" box="[896,1153,1824,1846]" gridcol="3" gridrow="15" pageId="12" pageNumber="239">
<taxonomicName id="4C6932BDFFBB3A1696E3F887FBCD323F" box="[896,1103,1824,1846]" pageId="12" pageNumber="239">
<emphasis id="B91D952CFFBB3A1696E3F887FB83323C" box="[896,1025,1824,1845]" italics="true" pageId="12" pageNumber="239">Labidodemas</emphasis>
species
</taxonomicName>
)
</td>
</tr>
<tr id="35594B7CFFBBC5E595C0F8EFFA053257" box="[163,1415,1864,1886]" gridrow="16" pageId="12" pageNumber="239">
<th id="76882200FFBBC5E595C0F8EFFEC73257" box="[163,325,1864,1886]" gridcol="0" gridrow="16" pageId="12" pageNumber="239">Anal papillae</th>
<td id="76882200FFBBC5E59400F8EFFDC73257" box="[355,581,1864,1886]" gridcol="1" gridrow="16" pageId="12" pageNumber="239">present</td>
<td id="76882200FFBBC5E5970EF8EFFCE83257" box="[621,874,1864,1886]" gridcol="2" gridrow="16" pageId="12" pageNumber="239">absent</td>
<td id="76882200FFBBC5E596E3F8EFFB033257" box="[896,1153,1864,1886]" gridcol="3" gridrow="16" pageId="12" pageNumber="239">present</td>
<td id="76882200FFBBC5E591C1F8EFFA053257" box="[1186,1415,1864,1886]" gridcol="4" gridrow="16" pageId="12" pageNumber="239">absent</td>
</tr>
</table>
</paragraph>
<paragraph id="8BD6493EFFBB3A169109F8D7FA13328C" blockId="12.[1130,1425,1904,1925]" box="[1130,1425,1904,1925]" pageId="12" pageNumber="239">
<tableNote id="768F48B0FFBB3A169109F8D7FA13328C" box="[1130,1425,1904,1925]" pageId="12" pageNumber="239" targetBox="[163,1415,1364,1886]" targetPageId="12">......continued on the next page</tableNote>
</paragraph>
<caption id="DF1619B6FFBA3A1795C0FF04FE0635B0" box="[163,388,163,185]" isContinuationCaption="true" pageId="13" pageNumber="240" startId="13.[163,243,163,184]" targetType="table">
<paragraph id="8BD6493EFFBA3A1795C0FF04FE0635B0" blockId="13.[163,388,163,185]" box="[163,388,163,185]" pageId="13" pageNumber="240">
<emphasis id="B91D952CFFBA3A1795C0FF04FE8935B1" bold="true" box="[163,267,163,184]" pageId="13" pageNumber="240">TABLE 1.</emphasis>
(Continued)
</paragraph>
</caption>
<paragraph id="8BD6493EFFBA3A1795C0FF6DFEC730B2" pageId="13" pageNumber="240">
<table id="F969BB9EFFBAC5E595C0FF6DFA0930B2" box="[163,1419,202,1467]" colsContinueFrom="12.[163,1415,1364,1886]" gridcols="5" gridrows="41" pageId="13" pageNumber="240">
<tr id="35594B7CFFBAC5E595C0FF6DFA0935E9" box="[163,1419,202,224]" gridrow="0" pageId="13" pageNumber="240">
<th id="76882200FFBAC5E595C0FF6DFEC735E9" box="[163,325,202,224]" gridcol="0" gridrow="0" pageId="13" pageNumber="240">Character</th>
<th id="76882200FFBAC5E59400FF6DFDD535E9" box="[355,599,202,224]" gridcol="1" gridrow="0" pageId="13" pageNumber="240">
<taxonomicName id="4C6932BDFFBA3A179400FF6CFE5735E9" authorityName="Setyastuti &amp; Solis-Marin &amp; Lüter" authorityYear="2024" box="[355,469,203,224]" class="Holothuroidea" family="Holothuriidae" genus="Labidodemas" kingdom="Animalia" order="Holothuriida" pageId="13" pageNumber="240" phylum="Echinodermata" rank="species" species="javaensis" status="sp. nov.">
<emphasis id="B91D952CFFBA3A179400FF6CFE5735E9" box="[355,469,203,224]" italics="true" pageId="13" pageNumber="240">L. javaensis</emphasis>
</taxonomicName>
<emphasis id="B91D952CFFBA3A1794BFFF6DFDAA35D6" bold="true" box="[476,552,202,223]" pageId="13" pageNumber="240">
<taxonomicNameLabel id="A22E2857FFBA3A1794BFFF6DFDAA35D6" box="[476,552,202,223]" pageId="13" pageNumber="240" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<th id="76882200FFBAC5E5970EFF6DFCE835E9" box="[621,874,202,224]" gridcol="2" gridrow="0" pageId="13" pageNumber="240">
<taxonomicName id="4C6932BDFFBA3A17970EFF6CFCE435E9" authority="Massin" authorityName="Massin, Samyn &amp; Thandar" authorityYear="2004" box="[621,870,202,224]" class="Holothuroidea" family="Holothuriidae" genus="Labidodemas" kingdom="Animalia" order="Holothuriida" pageId="13" pageNumber="240" phylum="Echinodermata" rank="species" species="quadripartitum">
<emphasis id="B91D952CFFBA3A17970EFF6CFC9B35D6" box="[621,793,202,224]" italics="true" pageId="13" pageNumber="240">L. quadripartitum</emphasis>
Massin
</taxonomicName>
,
</th>
<th id="76882200FFBAC5E596E3FF6DFB0A35E9" box="[896,1160,202,224]" gridcol="3" gridrow="0" pageId="13" pageNumber="240">
<taxonomicName id="4C6932BDFFBA3A1796E3FF6CFBED35E9" authority="Selenka" authorityName="Selenka" authorityYear="1867" box="[896,1135,202,224]" class="Holothuroidea" family="Holothuriidae" genus="Labidodemas" kingdom="Animalia" order="Holothuriida" pageId="13" pageNumber="240" phylum="Echinodermata" rank="species" species="semperianum">
<emphasis id="B91D952CFFBA3A1796E3FF6CFB9935E9" box="[896,1051,203,224]" italics="true" pageId="13" pageNumber="240">L. semperianum</emphasis>
Selenka
</taxonomicName>
,
</th>
<th id="76882200FFBAC5E591C1FF6DFA0935E9" box="[1186,1419,202,224]" gridcol="4" gridrow="0" pageId="13" pageNumber="240">
<taxonomicName id="4C6932BDFFBA3A1791C1FF6CFA0335D6" authority="Samyn &amp; Thandar, 2004" authorityName="Massin, Samyn &amp; Thandar" authorityYear="2004" box="[1186,1409,202,224]" class="Holothuroidea" family="Holothuriidae" genus="Labidodemas" kingdom="Animalia" order="Holothuriida" pageId="13" pageNumber="240" phylum="Echinodermata" rank="species" species="pseudosemperianum">
<emphasis id="B91D952CFFBA3A1791C1FF6CFA0335D6" box="[1186,1409,202,224]" italics="true" pageId="13" pageNumber="240">L. pseudosemperianum</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="35594B7CFFBAC5E595C0FF40FA0935F4" box="[163,1419,231,253]" gridrow="1" pageId="13" pageNumber="240" rowspan-0="1" rowspan-1="1">
<td id="76882200FFBAC5E5970EFF40FCE835F4" box="[621,874,231,253]" gridcol="2" gridrow="1" pageId="13" pageNumber="240">Samyn &amp; Thandar, 2004</td>
<td id="76882200FFBAC5E596E3FF40FB0A35F4" box="[896,1160,231,253]" gridcol="3" gridrow="1" pageId="13" pageNumber="240">
1867 (
<bibRefCitation id="EFF834CFFFBA3A1796DEFF40FBF135F4" author="Massin, C. &amp; Samyn, Y. &amp; Thandar, A. S." box="[957,1139,231,253]" pageId="13" pageNumber="240" pagination="1811 - 1847" refId="ref8451" refString="Massin, C., Samyn, Y. &amp; Thandar, A. S. (2004) The genus Labidodemas (Holothuroidea: Aspidochirotida) revisited, with description of three new species and with re-positioning of Holothuria (Irenothuria) maccullochi Deichmann, 1958. Journal of Natural History, 38, 1811 - 1847. https: // doi. org / 10.1080 / 0022293031000156268" type="journal article" year="2004">
Massin
<emphasis id="B91D952CFFBA3A179169FF4FFBB635F5" box="[1034,1076,231,253]" italics="true" pageId="13" pageNumber="240">et al</emphasis>
. 2004
</bibRefCitation>
;
</td>
<td id="76882200FFBAC5E591C1FF40FA0935F4" box="[1186,1419,231,253]" gridcol="4" gridrow="1" pageId="13" pageNumber="240">
(
<bibRefCitation id="EFF834CFFFBA3A1791CAFF40FADD35F4" author="Massin, C. &amp; Samyn, Y. &amp; Thandar, A. S." box="[1193,1375,231,253]" pageId="13" pageNumber="240" pagination="1811 - 1847" refId="ref8451" refString="Massin, C., Samyn, Y. &amp; Thandar, A. S. (2004) The genus Labidodemas (Holothuroidea: Aspidochirotida) revisited, with description of three new species and with re-positioning of Holothuria (Irenothuria) maccullochi Deichmann, 1958. Journal of Natural History, 38, 1811 - 1847. https: // doi. org / 10.1080 / 0022293031000156268" type="journal article" year="2004">
Massin
<emphasis id="B91D952CFFBA3A179195FF4FFAA235F5" box="[1270,1312,231,253]" italics="true" pageId="13" pageNumber="240">et al</emphasis>
. 2004
</bibRefCitation>
;
</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FEA2FA093412" box="[163,1419,261,283]" gridrow="2" pageId="13" pageNumber="240" rowspan-0="1" rowspan-1="1">
<td id="76882200FFBAC5E5970EFEA2FCE83412" box="[621,874,261,283]" gridcol="2" gridrow="2" pageId="13" pageNumber="240">
(
<bibRefCitation id="EFF834CFFFBA3A179710FEA2FCAB3413" author="Massin, C. &amp; Samyn, Y. &amp; Thandar, A. S." box="[627,809,261,283]" pageId="13" pageNumber="240" pagination="1811 - 1847" refId="ref8451" refString="Massin, C., Samyn, Y. &amp; Thandar, A. S. (2004) The genus Labidodemas (Holothuroidea: Aspidochirotida) revisited, with description of three new species and with re-positioning of Holothuria (Irenothuria) maccullochi Deichmann, 1958. Journal of Natural History, 38, 1811 - 1847. https: // doi. org / 10.1080 / 0022293031000156268" type="journal article" year="2004">
Massin
<emphasis id="B91D952CFFBA3A1797A3FEA1FD693413" box="[704,747,261,283]" italics="true" pageId="13" pageNumber="240">et al</emphasis>
. 2004
</bibRefCitation>
;
</td>
<td id="76882200FFBAC5E596E3FEA2FB0A3412" box="[896,1160,261,283]" gridcol="3" gridrow="2" pageId="13" pageNumber="240">
<bibRefCitation id="EFF834CFFFBA3A1796E3FEA2FBB23412" author="Samyn, Y. &amp; Appeltans, W. &amp; Kerr, A. M." box="[896,1072,261,283]" pageId="13" pageNumber="240" pagination="103 - 120" refId="ref8831" refString="Samyn, Y., Appeltans, W. &amp; Kerr, A. M. (2005) Phylogeny of Labidodemas and the Holothuriidae (Holothuroidea: Aspidochirotida) as inferred from morphology. Zoological Journal of the Linnean Society, 144, 103 - 120. https: // doi. org / 10.1111 / j. 1096 - 3642.2005.00158. x" type="journal article" year="2005">
Samyn
<emphasis id="B91D952CFFBA3A1796AAFEA1FC713413" box="[969,1011,261,283]" italics="true" pageId="13" pageNumber="240">et al</emphasis>
. 2005
</bibRefCitation>
)
</td>
<td id="76882200FFBAC5E591C1FEA2FA093412" box="[1186,1419,261,283]" gridcol="4" gridrow="2" pageId="13" pageNumber="240">
<bibRefCitation id="EFF834CFFFBA3A1791C1FEA2FAD03412" author="Samyn, Y. &amp; Appeltans, W. &amp; Kerr, A. M." box="[1186,1362,261,283]" pageId="13" pageNumber="240" pagination="103 - 120" refId="ref8831" refString="Samyn, Y., Appeltans, W. &amp; Kerr, A. M. (2005) Phylogeny of Labidodemas and the Holothuriidae (Holothuroidea: Aspidochirotida) as inferred from morphology. Zoological Journal of the Linnean Society, 144, 103 - 120. https: // doi. org / 10.1111 / j. 1096 - 3642.2005.00158. x" type="journal article" year="2005">
Samyn
<emphasis id="B91D952CFFBA3A179188FEA1FA973413" box="[1259,1301,261,283]" italics="true" pageId="13" pageNumber="240">et al</emphasis>
. 2005
</bibRefCitation>
)
</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FE85FA093431" box="[163,1419,290,312]" gridrow="3" pageId="13" pageNumber="240" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-4="1">
<td id="76882200FFBAC5E5970EFE85FCE83431" box="[621,874,290,312]" gridcol="2" gridrow="3" pageId="13" pageNumber="240">
<bibRefCitation id="EFF834CFFFBA3A17970EFE85FC9F3431" author="Samyn, Y. &amp; Appeltans, W. &amp; Kerr, A. M." box="[621,797,290,312]" pageId="13" pageNumber="240" pagination="103 - 120" refId="ref8831" refString="Samyn, Y., Appeltans, W. &amp; Kerr, A. M. (2005) Phylogeny of Labidodemas and the Holothuriidae (Holothuroidea: Aspidochirotida) as inferred from morphology. Zoological Journal of the Linnean Society, 144, 103 - 120. https: // doi. org / 10.1111 / j. 1096 - 3642.2005.00158. x" type="journal article" year="2005">
Samyn
<emphasis id="B91D952CFFBA3A1797D5FE84FD62343E" box="[694,736,290,312]" italics="true" pageId="13" pageNumber="240">et al</emphasis>
. 2005
</bibRefCitation>
)
</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FEEDFA093469" box="[163,1419,330,352]" gridrow="4" pageId="13" pageNumber="240">
<th id="76882200FFBAC5E595C0FEEDFEC73469" box="[163,325,330,352]" gridcol="0" gridrow="4" pageId="13" pageNumber="240">Tables</th>
<td id="76882200FFBAC5E59400FEEDFDD53469" box="[355,599,330,352]" gridcol="1" gridrow="4" pageId="13" pageNumber="240">-Disc with 45 central</td>
<td id="76882200FFBAC5E5970EFEEDFCE83469" box="[621,874,330,352]" gridcol="2" gridrow="4" pageId="13" pageNumber="240">-Disc with 4 central holes</td>
<td id="76882200FFBAC5E596E3FEEDFB0A3469" box="[896,1160,330,352]" gridcol="3" gridrow="4" pageId="13" pageNumber="240">-Disc with 46 central</td>
<td id="76882200FFBAC5E591C1FEEDFA093469" box="[1186,1419,330,352]" gridcol="4" gridrow="4" pageId="13" pageNumber="240">-Disc with 58 central</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FEC0FA093474" box="[163,1419,359,381]" gridrow="5" pageId="13" pageNumber="240" rowspan-0="1">
<td id="76882200FFBAC5E59400FEC0FDD53474" box="[355,599,359,381]" gridcol="1" gridrow="5" pageId="13" pageNumber="240">holes</td>
<td id="76882200FFBAC5E5970EFEC0FCE83474" box="[621,874,359,381]" gridcol="2" gridrow="5" pageId="13" pageNumber="240">-Disc with 02</td>
<td id="76882200FFBAC5E596E3FEC0FB0A3474" box="[896,1160,359,381]" gridcol="3" gridrow="5" pageId="13" pageNumber="240">holes</td>
<td id="76882200FFBAC5E591C1FEC0FA093474" box="[1186,1419,359,381]" gridcol="4" gridrow="5" pageId="13" pageNumber="240">holes</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FE22FA093492" box="[163,1419,389,411]" gridrow="6" pageId="13" pageNumber="240" rowspan-0="1">
<td id="76882200FFBAC5E59400FE22FDD53492" box="[355,599,389,411]" gridcol="1" gridrow="6" pageId="13" pageNumber="240">-Mainly disc devoid of</td>
<td id="76882200FFBAC5E5970EFE22FCE83492" box="[621,874,389,411]" gridcol="2" gridrow="6" pageId="13" pageNumber="240">(occasionally six)</td>
<td id="76882200FFBAC5E596E3FE22FB0A3492" box="[896,1160,389,411]" gridcol="3" gridrow="6" pageId="13" pageNumber="240">-Disc with 510 peripheral</td>
<td id="76882200FFBAC5E591C1FE22FA093492" box="[1186,1419,389,411]" gridcol="4" gridrow="6" pageId="13" pageNumber="240">-Disc with 515</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FE05FA0934B1" box="[163,1419,418,440]" gridrow="7" pageId="13" pageNumber="240" rowspan-0="1">
<td id="76882200FFBAC5E59400FE05FDD534B1" box="[355,599,418,440]" gridcol="1" gridrow="7" pageId="13" pageNumber="240">peripheral holes, rarely</td>
<td id="76882200FFBAC5E5970EFE05FCE834B1" box="[621,874,418,440]" gridcol="2" gridrow="7" pageId="13" pageNumber="240">peripheral holes</td>
<td id="76882200FFBAC5E596E3FE05FB0A34B1" box="[896,1160,418,440]" gridcol="3" gridrow="7" pageId="13" pageNumber="240">holes</td>
<td id="76882200FFBAC5E591C1FE05FA0934B1" box="[1186,1419,418,440]" gridcol="4" gridrow="7" pageId="13" pageNumber="240">peripheral holes</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FE18FA0934DC" box="[163,1419,447,469]" gridrow="8" pageId="13" pageNumber="240" rowspan-0="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<td id="76882200FFBAC5E59400FE18FDD534DC" box="[355,599,447,469]" gridcol="1" gridrow="8" pageId="13" pageNumber="240">with 24 peripheral</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FE7AFA0934FA" box="[163,1419,477,499]" gridrow="9" pageId="13" pageNumber="240" rowspan-0="1" rowspan-3="1" rowspan-4="1">
<td id="76882200FFBAC5E59400FE7AFDD534FA" box="[355,599,477,499]" gridcol="1" gridrow="9" pageId="13" pageNumber="240">holes</td>
<td id="76882200FFBAC5E5970EFE7AFCE834FA" box="[621,874,477,499]" gridcol="2" gridrow="9" pageId="13" pageNumber="240">-Disc 3060 µm across</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FE5DFA093719" box="[163,1419,506,528]" gridrow="10" pageId="13" pageNumber="240" rowspan-0="1">
<td id="76882200FFBAC5E59400FE5DFDD53719" box="[355,599,506,528]" gridcol="1" gridrow="10" pageId="13" pageNumber="240">-Disc 40100 µm across</td>
<td id="76882200FFBAC5E5970EFE5DFCE83719" box="[621,874,506,528]" gridcol="2" gridrow="10" pageId="13" pageNumber="240">-Rim of the disc spiny</td>
<td id="76882200FFBAC5E596E3FE5DFB0A3719" box="[896,1160,506,528]" gridcol="3" gridrow="10" pageId="13" pageNumber="240">-Disc 5590 µm across</td>
<td id="76882200FFBAC5E591C1FE5DFA093719" box="[1186,1419,506,528]" gridcol="4" gridrow="10" pageId="13" pageNumber="240">-Disc 7095 µm across</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FDB0FA093724" box="[163,1419,535,557]" gridrow="11" pageId="13" pageNumber="240" rowspan-0="1">
<td id="76882200FFBAC5E59400FDB0FDD53724" box="[355,599,535,557]" gridcol="1" gridrow="11" pageId="13" pageNumber="240">-Rim of the disc very</td>
<td id="76882200FFBAC5E5970EFDB0FCE83724" box="[621,874,535,557]" gridcol="2" gridrow="11" pageId="13" pageNumber="240">-Crown with 4 huge</td>
<td id="76882200FFBAC5E596E3FDB0FB0A3724" box="[896,1160,535,557]" gridcol="3" gridrow="11" pageId="13" pageNumber="240">-Rim of the disc spiny</td>
<td id="76882200FFBAC5E591C1FDB0FA093724" box="[1186,1419,535,557]" gridcol="4" gridrow="11" pageId="13" pageNumber="240">-Rim of the disc smooth</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FD92FA093742" box="[163,1419,565,587]" gridrow="12" pageId="13" pageNumber="240" rowspan-0="1">
<td id="76882200FFBAC5E59400FD92FDD53742" box="[355,599,565,587]" gridcol="1" gridrow="12" pageId="13" pageNumber="240">spiny</td>
<td id="76882200FFBAC5E5970EFD92FCE83742" box="[621,874,565,587]" gridcol="2" gridrow="12" pageId="13" pageNumber="240">spines that occasionally</td>
<td id="76882200FFBAC5E596E3FD92FB0A3742" box="[896,1160,565,587]" gridcol="3" gridrow="12" pageId="13" pageNumber="240">-Crown with 45 huge</td>
<td id="76882200FFBAC5E591C1FD92FA093742" box="[1186,1419,565,587]" gridcol="4" gridrow="12" pageId="13" pageNumber="240">-Crown with 58 huge</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FDF5FA093761" box="[163,1419,594,616]" gridrow="13" pageId="13" pageNumber="240" rowspan-0="1">
<td id="76882200FFBAC5E59400FDF5FDD53761" box="[355,599,594,616]" gridcol="1" gridrow="13" pageId="13" pageNumber="240">-Crown with 45 huge</td>
<td id="76882200FFBAC5E5970EFDF5FCE83761" box="[621,874,594,616]" gridcol="2" gridrow="13" pageId="13" pageNumber="240">bifurcate distally</td>
<td id="76882200FFBAC5E596E3FDF5FB0A3761" box="[896,1160,594,616]" gridcol="3" gridrow="13" pageId="13" pageNumber="240">spines that bifurcate</td>
<td id="76882200FFBAC5E591C1FDF5FA093761" box="[1186,1419,594,616]" gridcol="4" gridrow="13" pageId="13" pageNumber="240">spines that bifurcate</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FDC8FA09378C" box="[163,1419,623,645]" gridrow="14" pageId="13" pageNumber="240" rowspan-0="1" rowspan-2="1">
<td id="76882200FFBAC5E59400FDC8FDD5378C" box="[355,599,623,645]" gridcol="1" gridrow="14" pageId="13" pageNumber="240">spines, arranged in</td>
<td id="76882200FFBAC5E596E3FDC8FB0A378C" box="[896,1160,623,645]" gridcol="3" gridrow="14" pageId="13" pageNumber="240">slightly distally</td>
<td id="76882200FFBAC5E591C1FDC8FA09378C" box="[1186,1419,623,645]" gridcol="4" gridrow="14" pageId="13" pageNumber="240">deeply distally</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FD2AFA0937AA" box="[163,1419,653,675]" gridrow="15" pageId="13" pageNumber="240" rowspan-0="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<td id="76882200FFBAC5E59400FD2AFDD537AA" box="[355,599,653,675]" gridcol="1" gridrow="15" pageId="13" pageNumber="240">regular or irregular/</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FD0DFA0937C9" box="[163,1419,682,704]" gridrow="16" pageId="13" pageNumber="240" rowspan-0="1" rowspan-3="1" rowspan-4="1">
<td id="76882200FFBAC5E59400FD0DFDD537C9" box="[355,599,682,704]" gridcol="1" gridrow="16" pageId="13" pageNumber="240">overlapping, always</td>
<td id="76882200FFBAC5E5970EFD0DFCE837C9" box="[621,874,682,704]" gridcol="2" gridrow="16" pageId="13" pageNumber="240">-Table crown&gt; table disc</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FD60FA0937D4" box="[163,1419,711,733]" gridrow="17" pageId="13" pageNumber="240" rowspan-0="1" rowspan-2="1">
<td id="76882200FFBAC5E59400FD60FDD537D4" box="[355,599,711,733]" gridcol="1" gridrow="17" pageId="13" pageNumber="240">bifurcate distally</td>
<td id="76882200FFBAC5E596E3FD60FB0A37D4" box="[896,1160,711,733]" gridcol="3" gridrow="17" pageId="13" pageNumber="240">-Table crown ≥ table disc</td>
<td id="76882200FFBAC5E591C1FD60FA0937D4" box="[1186,1419,711,733]" gridcol="4" gridrow="17" pageId="13" pageNumber="240">-Table crown &gt;&gt; table</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FD42FA0937F2" box="[163,1419,741,763]" gridrow="18" pageId="13" pageNumber="240" rowspan-0="1" rowspan-3="1">
<td id="76882200FFBAC5E59400FD42FDD537F2" box="[355,599,741,763]" gridcol="1" gridrow="18" pageId="13" pageNumber="240">-Table crown&gt; table disc</td>
<td id="76882200FFBAC5E5970EFD42FCE837F2" box="[621,874,741,763]" gridcol="2" gridrow="18" pageId="13" pageNumber="240">-Crown of spines with</td>
<td id="76882200FFBAC5E591C1FD42FA0937F2" box="[1186,1419,741,763]" gridcol="4" gridrow="18" pageId="13" pageNumber="240">disc (crown up to twice</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FCA5FA093611" box="[163,1419,770,792]" gridrow="19" pageId="13" pageNumber="240" rowspan-0="1" rowspan-1="1">
<td id="76882200FFBAC5E5970EFCA5FCE83611" box="[621,874,770,792]" gridcol="2" gridrow="19" pageId="13" pageNumber="240">one central hole</td>
<td id="76882200FFBAC5E596E3FCA5FB0A3611" box="[896,1160,770,792]" gridcol="3" gridrow="19" pageId="13" pageNumber="240">-Crown of spines with one</td>
<td id="76882200FFBAC5E591C1FCA5FA093611" box="[1186,1419,770,792]" gridcol="4" gridrow="19" pageId="13" pageNumber="240">the disc)</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FCB8FA09363C" box="[163,1419,799,821]" gridrow="20" pageId="13" pageNumber="240" rowspan-0="1" rowspan-2="1">
<td id="76882200FFBAC5E59400FCB8FDD5363C" box="[355,599,799,821]" gridcol="1" gridrow="20" pageId="13" pageNumber="240">-Crown of spines with</td>
<td id="76882200FFBAC5E596E3FCB8FB0A363C" box="[896,1160,799,821]" gridcol="3" gridrow="20" pageId="13" pageNumber="240">(occasionally two) central</td>
<td id="76882200FFBAC5E591C1FCB8FA09363C" box="[1186,1419,799,821]" gridcol="4" gridrow="20" pageId="13" pageNumber="240">-Crown of spines with</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FC9AFA09365A" box="[163,1419,829,851]" gridrow="21" pageId="13" pageNumber="240" rowspan-0="1">
<td id="76882200FFBAC5E59400FC9AFDD5365A" box="[355,599,829,851]" gridcol="1" gridrow="21" pageId="13" pageNumber="240">one central hole</td>
<td id="76882200FFBAC5E5970EFC9AFCE8365A" box="[621,874,829,851]" gridcol="2" gridrow="21" pageId="13" pageNumber="240">-Dorsal table few</td>
<td id="76882200FFBAC5E596E3FC9AFB0A365A" box="[896,1160,829,851]" gridcol="3" gridrow="21" pageId="13" pageNumber="240">hole</td>
<td id="76882200FFBAC5E591C1FC9AFA09365A" box="[1186,1419,829,851]" gridcol="4" gridrow="21" pageId="13" pageNumber="240">210 central holes</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FCD0FA093684" box="[163,1419,887,909]" gridrow="22" pageId="13" pageNumber="240" rowspan-0="1">
<td id="76882200FFBAC5E59400FCD0FDD53684" box="[355,599,887,909]" gridcol="1" gridrow="22" pageId="13" pageNumber="240">-Dorsal table numerous</td>
<td id="76882200FFBAC5E5970EFCD0FCE83684" box="[621,874,887,909]" gridcol="2" gridrow="22" pageId="13" pageNumber="240">-Ventral tube feet table</td>
<td id="76882200FFBAC5E596E3FCD0FB0A3684" box="[896,1160,887,909]" gridcol="3" gridrow="22" pageId="13" pageNumber="240">-Dorsal table numerous</td>
<td id="76882200FFBAC5E591C1FCD0FA093684" box="[1186,1419,887,909]" gridcol="4" gridrow="22" pageId="13" pageNumber="240">-Dorsal table numerous</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FC32FA0936A2" box="[163,1419,917,939]" gridrow="23" pageId="13" pageNumber="240" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-4="1">
<td id="76882200FFBAC5E5970EFC32FCE836A2" box="[621,874,917,939]" gridcol="2" gridrow="23" pageId="13" pageNumber="240">mostly reduced to</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FC15FA0936C1" box="[163,1419,946,968]" gridrow="24" pageId="13" pageNumber="240" rowspan-0="1">
<td id="76882200FFBAC5E59400FC15FDD536C1" box="[355,599,946,968]" gridcol="1" gridrow="24" pageId="13" pageNumber="240">-Ventral tube feet table</td>
<td id="76882200FFBAC5E5970EFC15FCE836C1" box="[621,874,946,968]" gridcol="2" gridrow="24" pageId="13" pageNumber="240">disc, 3342 µm across,</td>
<td id="76882200FFBAC5E596E3FC15FB0A36C1" box="[896,1160,946,968]" gridcol="3" gridrow="24" pageId="13" pageNumber="240">-Ventral tube feet tables are</td>
<td id="76882200FFBAC5E591C1FC15FA0936C1" box="[1186,1419,946,968]" gridcol="4" gridrow="24" pageId="13" pageNumber="240">-Ventral tube feet</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FC68FA0936EC" box="[163,1419,975,997]" gridrow="25" pageId="13" pageNumber="240" rowspan-0="1">
<td id="76882200FFBAC5E59400FC68FDD536EC" box="[355,599,975,997]" gridcol="1" gridrow="25" pageId="13" pageNumber="240">mostly reduced to</td>
<td id="76882200FFBAC5E5970EFC68FCE836EC" box="[621,874,975,997]" gridcol="2" gridrow="25" pageId="13" pageNumber="240">perforated by 34 central</td>
<td id="76882200FFBAC5E596E3FC68FB0A36EC" box="[896,1160,975,997]" gridcol="3" gridrow="25" pageId="13" pageNumber="240">reduced to the spiny-edge</td>
<td id="76882200FFBAC5E591C1FC68FA0936EC" box="[1186,1419,975,997]" gridcol="4" gridrow="25" pageId="13" pageNumber="240">table mostly reduced</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FC4AFA09310A" box="[163,1419,1005,1027]" gridrow="26" pageId="13" pageNumber="240" rowspan-0="1">
<td id="76882200FFBAC5E59400FC4AFDD5310A" box="[355,599,1005,1027]" gridcol="1" gridrow="26" pageId="13" pageNumber="240">disc, 4060 µm across,</td>
<td id="76882200FFBAC5E5970EFC4AFCE8310A" box="[621,874,1005,1027]" gridcol="2" gridrow="26" pageId="13" pageNumber="240">holes and occasionally a</td>
<td id="76882200FFBAC5E596E3FC4AFB0A310A" box="[896,1160,1005,1027]" gridcol="3" gridrow="26" pageId="13" pageNumber="240">disc, 4050 µm across,</td>
<td id="76882200FFBAC5E591C1FC4AFA09310A" box="[1186,1419,1005,1027]" gridcol="4" gridrow="26" pageId="13" pageNumber="240">to disc, 4550 µm</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FBADFA093129" box="[163,1419,1034,1056]" gridrow="27" pageId="13" pageNumber="240" rowspan-0="1">
<td id="76882200FFBAC5E59400FBADFDD53129" box="[355,599,1034,1056]" gridcol="1" gridrow="27" pageId="13" pageNumber="240">perforated by 45</td>
<td id="76882200FFBAC5E5970EFBADFCE83129" box="[621,874,1034,1056]" gridcol="2" gridrow="27" pageId="13" pageNumber="240">single peripheral hole, rim</td>
<td id="76882200FFBAC5E596E3FBADFB0A3129" box="[896,1160,1034,1056]" gridcol="3" gridrow="27" pageId="13" pageNumber="240">perforated by 34 central</td>
<td id="76882200FFBAC5E591C1FBADFA093129" box="[1186,1419,1034,1056]" gridcol="4" gridrow="27" pageId="13" pageNumber="240">across, perforated by</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FB80FA093134" box="[163,1419,1063,1085]" gridrow="28" pageId="13" pageNumber="240" rowspan-0="1">
<td id="76882200FFBAC5E59400FB80FDD53134" box="[355,599,1063,1085]" gridcol="1" gridrow="28" pageId="13" pageNumber="240">central holes, never with</td>
<td id="76882200FFBAC5E5970EFB80FCE83134" box="[621,874,1063,1085]" gridcol="2" gridrow="28" pageId="13" pageNumber="240">of disc slightly spinous</td>
<td id="76882200FFBAC5E596E3FB80FB0A3134" box="[896,1160,1063,1085]" gridcol="3" gridrow="28" pageId="13" pageNumber="240">holes and occasionally 12</td>
<td id="76882200FFBAC5E591C1FB80FA093134" box="[1186,1419,1063,1085]" gridcol="4" gridrow="28" pageId="13" pageNumber="240">46 central holes and</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FBE2FA093152" box="[163,1419,1093,1115]" gridrow="29" pageId="13" pageNumber="240" rowspan-0="1" rowspan-2="1">
<td id="76882200FFBAC5E59400FBE2FDD53152" box="[355,599,1093,1115]" gridcol="1" gridrow="29" pageId="13" pageNumber="240">peripheral holes, rim of</td>
<td id="76882200FFBAC5E596E3FBE2FB0A3152" box="[896,1160,1093,1115]" gridcol="3" gridrow="29" pageId="13" pageNumber="240">peripheral holes</td>
<td id="76882200FFBAC5E591C1FBE2FA093152" box="[1186,1419,1093,1115]" gridcol="4" gridrow="29" pageId="13" pageNumber="240">occasionally a single</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FBC5FA093171" box="[163,1419,1122,1144]" gridrow="30" pageId="13" pageNumber="240" rowspan-0="1" rowspan-2="1" rowspan-3="1">
<td id="76882200FFBAC5E59400FBC5FDD53171" box="[355,599,1122,1144]" gridcol="1" gridrow="30" pageId="13" pageNumber="240">disc very spinous with</td>
<td id="76882200FFBAC5E591C1FBC5FA093171" box="[1186,1419,1122,1144]" gridcol="4" gridrow="30" pageId="13" pageNumber="240">peripheral hole, rim of</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FBD8FA09319C" box="[163,1419,1151,1173]" gridrow="31" pageId="13" pageNumber="240" rowspan-0="1" rowspan-2="1" rowspan-3="1">
<td id="76882200FFBAC5E59400FBD8FDD5319C" box="[355,599,1151,1173]" gridcol="1" gridrow="31" pageId="13" pageNumber="240">several elongated spines</td>
<td id="76882200FFBAC5E591C1FBD8FA09319C" box="[1186,1419,1151,1173]" gridcol="4" gridrow="31" pageId="13" pageNumber="240">the disc slightly spinous</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FB3AFA0931BA" box="[163,1419,1181,1203]" gridrow="32" pageId="13" pageNumber="240" rowspan-0="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<td id="76882200FFBAC5E59400FB3AFDD531BA" box="[355,599,1181,1203]" gridcol="1" gridrow="32" pageId="13" pageNumber="240">that bifurcate distally</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FB63FA0931D3" box="[163,1419,1220,1242]" gridrow="33" pageId="13" pageNumber="240">
<th id="76882200FFBAC5E595C0FB63FEC731D3" box="[163,325,1220,1242]" gridcol="0" gridrow="33" pageId="13" pageNumber="240">Buttons</th>
<td id="76882200FFBAC5E59400FB63FDD531D3" box="[355,599,1220,1242]" gridcol="1" gridrow="33" pageId="13" pageNumber="240">Absent</td>
<td id="76882200FFBAC5E5970EFB63FCE831D3" box="[621,874,1220,1242]" gridcol="2" gridrow="33" pageId="13" pageNumber="240">Absent</td>
<td id="76882200FFBAC5E596E3FB63FB0A31D3" box="[896,1160,1220,1242]" gridcol="3" gridrow="33" pageId="13" pageNumber="240">Present in dorsal body wall</td>
<td id="76882200FFBAC5E591C1FB63FA0931D3" box="[1186,1419,1220,1242]" gridcol="4" gridrow="33" pageId="13" pageNumber="240">Present in dorsal and</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FB46FA0931FE" box="[163,1419,1249,1271]" gridrow="34" pageId="13" pageNumber="240" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<td id="76882200FFBAC5E591C1FB46FA0931FE" box="[1186,1419,1249,1271]" gridcol="4" gridrow="34" pageId="13" pageNumber="240">ventral body wall</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FAAEFA093016" box="[163,1419,1289,1311]" gridrow="35" pageId="13" pageNumber="240">
<th id="76882200FFBAC5E595C0FAAEFEC73016" box="[163,325,1289,1311]" gridcol="0" gridrow="35" pageId="13" pageNumber="240">Pseudo-buttons</th>
<td id="76882200FFBAC5E59400FAAEFDD53016" box="[355,599,1289,1311]" gridcol="1" gridrow="35" pageId="13" pageNumber="240">Present in dorsal body</td>
<td id="76882200FFBAC5E5970EFAAEFCE83016" box="[621,874,1289,1311]" gridcol="2" gridrow="35" pageId="13" pageNumber="240">Absent</td>
<td id="76882200FFBAC5E596E3FAAEFB0A3016" box="[896,1160,1289,1311]" gridcol="3" gridrow="35" pageId="13" pageNumber="240">Present in dorsal body wall</td>
<td id="76882200FFBAC5E591C1FAAEFA093016" box="[1186,1419,1289,1311]" gridcol="4" gridrow="35" pageId="13" pageNumber="240">Present (a few) in</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FA81FA093035" box="[163,1419,1318,1340]" gridrow="36" pageId="13" pageNumber="240" rowspan-0="1" rowspan-2="1">
<td id="76882200FFBAC5E59400FA81FDD53035" box="[355,599,1318,1340]" gridcol="1" gridrow="36" pageId="13" pageNumber="240">wall, dorsal papillae, and</td>
<td id="76882200FFBAC5E596E3FA81FB0A3035" box="[896,1160,1318,1340]" gridcol="3" gridrow="36" pageId="13" pageNumber="240">and dorsal papillae</td>
<td id="76882200FFBAC5E591C1FA81FA093035" box="[1186,1419,1318,1340]" gridcol="4" gridrow="36" pageId="13" pageNumber="240">ventral tube feet</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FAE4FA093050" box="[163,1419,1347,1369]" gridrow="37" pageId="13" pageNumber="240" rowspan-0="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<td id="76882200FFBAC5E59400FAE4FDD53050" box="[355,599,1347,1369]" gridcol="1" gridrow="37" pageId="13" pageNumber="240">ventral tube feet</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FACCFA093088" box="[163,1419,1387,1409]" gridrow="38" pageId="13" pageNumber="240">
<th id="76882200FFBAC5E595C0FACCFEC73088" box="[163,325,1387,1409]" gridcol="0" gridrow="38" pageId="13" pageNumber="240">Tri-radiate rods</th>
<td id="76882200FFBAC5E59400FACCFDD53088" box="[355,599,1387,1409]" gridcol="1" gridrow="38" pageId="13" pageNumber="240">Present</td>
<td id="76882200FFBAC5E5970EFACCFCE83088" box="[621,874,1387,1409]" gridcol="2" gridrow="38" pageId="13" pageNumber="240">Absent</td>
<td id="76882200FFBAC5E596E3FACCFB0A3088" box="[896,1160,1387,1409]" gridcol="3" gridrow="38" pageId="13" pageNumber="240">Absent</td>
<td id="76882200FFBAC5E591C1FACCFA093088" box="[1186,1419,1387,1409]" gridcol="4" gridrow="38" pageId="13" pageNumber="240">Absent</td>
</tr>
<tr id="35594B7CFFBAC5E595C0FA2FFA093097" box="[163,1419,1416,1438]" gridrow="39" pageId="13" pageNumber="240" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="76882200FFBAC5E595C0FA2FFEC73097" box="[163,325,1416,1438]" gridcol="0" gridrow="39" pageId="13" pageNumber="240">dorsal body wall</th>
</tr>
<tr id="35594B7CFFBAC5E595C0FA02FA0930B2" box="[163,1419,1445,1467]" gridrow="40" pageId="13" pageNumber="240" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="76882200FFBAC5E595C0FA02FEC730B2" box="[163,325,1445,1467]" gridcol="0" gridrow="40" pageId="13" pageNumber="240">and anal papillae</th>
</tr>
</table>
</paragraph>
<subSubSection id="C3731AB5FFBA3A1795A4F9AEFC85326E" pageId="13" pageNumber="240" type="discussion">
<paragraph id="8BD6493EFFBA3A1795A4F9AEFBDF334F" blockId="13.[151,1437,1544,2039]" pageId="13" pageNumber="240">
<emphasis id="B91D952CFFBA3A1795A4F9AEFEB6332A" bold="true" box="[199,308,1545,1571]" pageId="13" pageNumber="240">Remarks</emphasis>
. The new species is characterised by having a ribbon-like calcareous ring, three rows of ventral tube feet, and ossicle assemblage that undoubtedly belongs to
<taxonomicName id="4C6932BDFFBA3A17967AF98BFBDA334F" authority="Selenka, 1867" authorityName="Selenka" authorityYear="1867" box="[793,1112,1580,1606]" class="Holothuroidea" family="Holothuriidae" genus="Labidodemas" kingdom="Animalia" order="Holothuriida" pageId="13" pageNumber="240" phylum="Echinodermata" rank="genus">
<emphasis id="B91D952CFFBA3A17967AF98BFC32334F" box="[793,944,1580,1606]" italics="true" pageId="13" pageNumber="240">Labidodemas</emphasis>
<bibRefCitation id="EFF834CFFFBA3A1796D4F98BFBDA334F" author="Selenka, E." box="[951,1112,1580,1606]" pageId="13" pageNumber="240" pagination="291 - 374" refId="ref8895" refString="Selenka, E. (1867) Beitrage zur anatomie und systematik der holothurien. Zeitschrift fur wissenschaftliche Zoologie, 17 (2), 291 - 374." type="journal article" year="1867">Selenka, 1867</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD6493EFFBA3A1795A4F9F7FC1633DF" blockId="13.[151,1437,1544,2039]" pageId="13" pageNumber="240">
Three main characters specific to
<taxonomicName id="4C6932BDFFBA3A179720F9F6FD4B3363" authorityName="Setyastuti &amp; Solis-Marin &amp; Lüter" authorityYear="2024" box="[579,713,1617,1642]" class="Holothuroidea" family="Holothuriidae" genus="Labidodemas" kingdom="Animalia" order="Holothuriida" pageId="13" pageNumber="240" phylum="Echinodermata" rank="species" species="javaensis" status="sp. nov.">
<emphasis id="B91D952CFFBA3A179720F9F6FD4B3363" box="[579,713,1617,1642]" italics="true" pageId="13" pageNumber="240">L. javaensis</emphasis>
</taxonomicName>
<emphasis id="B91D952CFFBA3A1797B3F9F6FCA83362" bold="true" box="[720,810,1617,1643]" pageId="13" pageNumber="240">
<taxonomicNameLabel id="A22E2857FFBA3A1797B3F9F6FCA83362" box="[720,810,1617,1643]" pageId="13" pageNumber="240" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are (1) most table discs devoid of peripheral holes with rim of the discs very spiny, (2) dorsal papillae and ventral tube feet always with
<specimenCount id="9D6F82B7FFBA3A179169F9D2FBF73386" box="[1034,1141,1653,1679]" count="2" pageId="13" pageNumber="240" type="generic" typeStatus="types">two types</specimenCount>
of tables (i.e. regular table with crown spines always bifurcate distally and smaller table size with very spiny disc and crown), (3) the present of tri-radiate shape of rods in the dorsal body wall and anal papillae.
</paragraph>
<paragraph id="8BD6493EFFBA3A1795A4F947FC85326E" blockId="13.[151,1437,1544,2039]" pageId="13" pageNumber="240">
The ossicle assemblages show that
<taxonomicName id="4C6932BDFFBA3A179706F947FCED33F3" authorityName="Setyastuti &amp; Solis-Marin &amp; Lüter" authorityYear="2024" box="[613,879,1760,1786]" class="Holothuroidea" family="Holothuriidae" genus="Labidodemas" kingdom="Animalia" order="Holothuriida" pageId="13" pageNumber="240" phylum="Echinodermata" rank="species" species="javaensis" status="sp. nov.">
<emphasis id="B91D952CFFBA3A179706F947FCED33F3" box="[613,879,1760,1786]" italics="true" pageId="13" pageNumber="240">Labidodemas javaensis</emphasis>
</taxonomicName>
<emphasis id="B91D952CFFBA3A179619F946FC5533F2" bold="true" box="[890,983,1761,1787]" pageId="13" pageNumber="240">
<taxonomicNameLabel id="A22E2857FFBA3A179619F946FC5533F2" box="[890,983,1761,1787]" pageId="13" pageNumber="240" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is closely related to
<taxonomicName id="4C6932BDFFBA3A1791B3F946FD8D3216" authority="Massin, Samyn &amp; Thandar, 2004" authorityName="Massin, Samyn &amp; Thandar" authorityYear="2004" class="Holothuroidea" family="Holothuriidae" genus="Labidodemas" kingdom="Animalia" order="Holothuriida" pageId="13" pageNumber="240" phylum="Echinodermata" rank="species" species="quadripartitum">
<emphasis id="B91D952CFFBA3A1791B3F946FA1E33F3" box="[1232,1436,1760,1786]" italics="true" pageId="13" pageNumber="240">L. quadripartitum</emphasis>
<bibRefCitation id="EFF834CFFFBA3A1795F4F8A3FD8D3216" author="Massin, C. &amp; Samyn, Y. &amp; Thandar, A. S." box="[151,527,1796,1823]" pageId="13" pageNumber="240" pagination="1811 - 1847" refId="ref8451" refString="Massin, C., Samyn, Y. &amp; Thandar, A. S. (2004) The genus Labidodemas (Holothuroidea: Aspidochirotida) revisited, with description of three new species and with re-positioning of Holothuria (Irenothuria) maccullochi Deichmann, 1958. Journal of Natural History, 38, 1811 - 1847. https: // doi. org / 10.1080 / 0022293031000156268" type="journal article" year="2004">Massin, Samyn &amp; Thandar, 2004</bibRefCitation>
</taxonomicName>
, but several differences warrant its new species status.
<tableCitation id="C6EB7C85FFBA3A17911FF8A3FB4D3216" box="[1148,1231,1796,1823]" captionStart="TABLE 1" captionStartId="12.[151,239,1287,1311]" captionText="TABLE 1. Characters differentiate Labidodemas javaensis sp. nov. from L. quadripartitum Massin, Samyn &amp; Thandar 2004, L semperianum Selenka, 1867, and L. pseudosemperianum Massin, Samyn &amp; Thandar 2004." pageId="13" pageNumber="240">Table 1</tableCitation>
lists the character differences between both species and the other two closer-related species (
<taxonomicName id="4C6932BDFFBA3A17968FF88EFAC9324B" authority="Selenka, 1867" authorityName="Selenka" authorityYear="1867" box="[1004,1355,1832,1858]" class="Holothuroidea" family="Holothuriidae" genus="Labidodemas" kingdom="Animalia" order="Holothuriida" pageId="13" pageNumber="240" phylum="Echinodermata" rank="species" species="semperianum">
<emphasis id="B91D952CFFBA3A17968FF88EFB20324B" box="[1004,1186,1833,1858]" italics="true" pageId="13" pageNumber="240">L. semperianum</emphasis>
<bibRefCitation id="EFF834CFFFBA3A1791C8F88FFAC9324B" author="Selenka, E." box="[1195,1355,1832,1858]" pageId="13" pageNumber="240" pagination="291 - 374" refId="ref8895" refString="Selenka, E. (1867) Beitrage zur anatomie und systematik der holothurien. Zeitschrift fur wissenschaftliche Zoologie, 17 (2), 291 - 374." type="journal article" year="1867">Selenka, 1867</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C6932BDFFBA3A1790E5F88EFD79326E" authority="Massin, Samyn &amp; Thandar, 2004" authorityName="Massin, Samyn &amp; Thandar" authorityYear="2004" class="Holothuroidea" family="Holothuriidae" genus="Labidodemas" kingdom="Animalia" order="Holothuriida" pageId="13" pageNumber="240" phylum="Echinodermata" rank="species" species="pseudosemperianum">
<emphasis id="B91D952CFFBA3A1790E5F88EFEFF326F" italics="true" pageId="13" pageNumber="240">L. pseudosemperianum</emphasis>
<bibRefCitation id="EFF834CFFFBA3A1794E7F8EBFD79326E" author="Massin, C. &amp; Samyn, Y. &amp; Thandar, A. S." box="[388,763,1868,1895]" pageId="13" pageNumber="240" pagination="1811 - 1847" refId="ref8451" refString="Massin, C., Samyn, Y. &amp; Thandar, A. S. (2004) The genus Labidodemas (Holothuroidea: Aspidochirotida) revisited, with description of three new species and with re-positioning of Holothuria (Irenothuria) maccullochi Deichmann, 1958. Journal of Natural History, 38, 1811 - 1847. https: // doi. org / 10.1080 / 0022293031000156268" type="journal article" year="2004">Massin, Samyn &amp; Thandar, 2004</bibRefCitation>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3731AB5FFBA3A1795A4F8D6FBE83283" box="[199,1130,1904,1931]" pageId="13" pageNumber="240" type="etymology">
<paragraph id="8BD6493EFFBA3A1795A4F8D6FBE83283" blockId="13.[151,1437,1544,2039]" box="[199,1130,1904,1931]" pageId="13" pageNumber="240">
<emphasis id="B91D952CFFBA3A1795A4F8D6FECC3282" bold="true" box="[199,334,1905,1931]" pageId="13" pageNumber="240">Etymology.</emphasis>
Named after its
<typeStatus id="54D2F79CFFBA3A17976BF8D6FDBA3282" box="[520,568,1905,1931]" pageId="13" pageNumber="240">type</typeStatus>
locality and regional occurrence,
<collectingRegion id="49AD87DCFFBA3A1796D5F8D6FC683282" box="[950,1002,1905,1931]" country="Indonesia" name="Jawa Timur" pageId="13" pageNumber="240">Java</collectingRegion>
,
<collectingCountry id="F37E09AEFFBA3A179696F8D7FBE43283" box="[1013,1126,1904,1930]" name="Indonesia" pageId="13" pageNumber="240">Indonesia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3731AB5FFBA3A1795A4F832FC7132FF" pageId="13" pageNumber="240" type="biology_ecology">
<paragraph id="8BD6493EFFBA3A1795A4F832FC4932A7" blockId="13.[151,1437,1544,2039]" box="[199,971,1940,1967]" pageId="13" pageNumber="240">
<emphasis id="B91D952CFFBA3A1795A4F832FEA432A6" bold="true" box="[199,294,1941,1967]" pageId="13" pageNumber="240">Ecology</emphasis>
. A nocturnal animal, it hides under the rocks during the day.
</paragraph>
<paragraph id="8BD6493EFFBA3A1795A4F81EFE7F32DB" blockId="13.[151,1437,1544,2039]" box="[199,509,1976,2003]" pageId="13" pageNumber="240">
<emphasis id="B91D952CFFBA3A1795A4F81EFED932DA" bold="true" box="[199,347,1977,2003]" pageId="13" pageNumber="240">Depth range</emphasis>
.
<quantity id="4C91E4DBFFBA3A179409F81EFE3332DA" box="[362,433,1977,2003]" metricMagnitude="0" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="13" pageNumber="240" unit="m" value="5.0" valueMax="7.0" valueMin="3.0">37 m</quantity>
depth.
</paragraph>
<paragraph id="8BD6493EFFBA3A1795A4F87BFC7132FF" blockId="13.[151,1437,1544,2039]" box="[199,1011,2012,2039]" pageId="13" pageNumber="240">
<emphasis id="B91D952CFFBA3A1795A4F87BFE6A32FF" bold="true" box="[199,488,2012,2038]" pageId="13" pageNumber="240">Geographic distribution</emphasis>
. At present only known from the
<typeStatus id="54D2F79CFFBA3A179601F87AFC1032FE" box="[866,914,2013,2039]" pageId="13" pageNumber="240">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,285 @@
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<mods:title id="5DDD3B210FC2A716BAD27CAEB8B88D93">First record of Hornylia nalanda from India (Hemiptera: Heteroptera: Reduviidae: Emesinae)</mods:title>
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<mods:namePart id="DCA228CA8DDEF15B2D6BAB9BB17CB433">Boyane, Swapnil S.</mods:namePart>
<mods:affiliation id="77E2838C0AFF80B26B0994948FBBCE53">Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur Post, Bangalore 560064, India &amp; Current address: Department of Biological Sciences, Texas Tech University, Lubbock 79409, Texas, USA</mods:affiliation>
</mods:name>
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<mods:role id="5AE32C664B4FBCAFA2AC90D1D68D21C9">
<mods:roleTerm id="509CEDE71689572E400F90E580716BEF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A50FEA9ED248306EE126C9B7970A12D4">Ranjith, A. P.</mods:namePart>
<mods:nameIdentifier id="46A11B2F3FF0BE97D21E94DA3383A3C4" type="ORCID">0000-0001-7061-9659</mods:nameIdentifier>
<mods:affiliation id="9F0792BBD5AAF57B550163EC5B9ADC52">Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur Post, Bangalore 560064, India</mods:affiliation>
<mods:nameIdentifier id="E4341B8F8759809A63409EC5F8D85D0B" type="email">ridhuranjith@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="AD8D57EB32EC9B2DCA2881CE587C63A7" type="personal">
<mods:role id="1C41092EDEB053F9BD4D483F5BDB2D29">
<mods:roleTerm id="1FEF6CD95C9C3A039816CF6FE9488392">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A2EB541F91D44660AE10A9905DDF0614">Ghate, Hemant V.</mods:namePart>
<mods:affiliation id="B3DBB80C95BB56B5B7A44764B8B9EDF1">Post-Graduate Research Centre, Department of Zoology, Modern College of Arts, Science &amp; Commerce (Autonomous), Shivajinagar, Pune 411005, India</mods:affiliation>
</mods:name>
<mods:typeOfResource id="14741169185EB32E9F4D79D1F8666D6C">text</mods:typeOfResource>
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<mods:title id="B3867A2062E721F156577BE7E8AF7062">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="544310E594A89E315726416113169D2B">
<mods:date id="A44C6F015DCEB26F716DC125941EF50F">2024</mods:date>
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<mods:number id="4E08D800DEB6C8BDE729A9841B6E519B">2024-09-05</mods:number>
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<mods:number id="222EEC10D608F83CF003736827156769">5506</mods:number>
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<mods:start id="BD710163A2322F2A340443EADBF1DD8C">104</mods:start>
<mods:end id="9122C9781105E22C9722CC131911CE4D">112</mods:end>
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<mods:url id="6E4FDC9540C41DC7200375129B47BDD4">http://dx.doi.org/10.11646/zootaxa.5506.1.6</mods:url>
</mods:location>
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<mods:identifier id="50DBB4D1308F36A6E74554C9A8D28A3A" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="1802D5FABFA418A7FBF8AC034A8FC566" type="ZooBank">6AC60E48-93CF-4C0C-AC23-7873E005799B</mods:identifier>
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<treatment id="03C19E3EFF1E2A694CC3F773FE3564C8" ID-DOI="http://doi.org/10.5281/zenodo.13746401" ID-Zenodo-Dep="13746401" LSID="urn:lsid:plazi:treatment:03C19E3EFF1E2A694CC3F773FE3564C8" httpUri="http://treatment.plazi.org/id/03C19E3EFF1E2A694CC3F773FE3564C8" lastPageId="4" lastPageNumber="108" pageId="1" pageNumber="105">
<subSubSection id="C3727CA3FF1E2A6C4CC3F773FDC1642A" box="[151,614,1884,1911]" pageId="1" pageNumber="105" type="nomenclature">
<paragraph id="8BD72F28FF1E2A6C4CC3F773FDC1642A" blockId="1.[151,614,1884,1947]" box="[151,614,1884,1911]" pageId="1" pageNumber="105">
<heading id="D09F9844FF1E2A6C4CC3F773FDC1642A" bold="true" box="[151,614,1884,1911]" fontSize="11" level="1" pageId="1" pageNumber="105" reason="1">
<taxonomicName id="4C6854ABFF1E2A6C4CC3F773FDC1642A" ID-CoL="3MQ3Y" authority="Wygodzinsky, 1966" authorityName="Wygodzinsky" authorityYear="1966" box="[151,614,1884,1911]" class="Insecta" family="Reduviidae" genus="Hornylia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="105" phylum="Arthropoda" rank="species" species="nalanda">
<emphasis id="B91CF33AFF1E2A6C4CC3F773FDC1642A" bold="true" box="[151,614,1884,1911]" pageId="1" pageNumber="105">
<emphasis id="B91CF33AFF1E2A6C4CC3F773FEC8642A" bold="true" box="[151,367,1884,1911]" italics="true" pageId="1" pageNumber="105">Hornylia nalanda</emphasis>
<bibRefCitation id="EFF952D9FF1E2A6C4D22F772FDC1642A" author="Wygodzinsky, P. W." box="[374,614,1885,1911]" pageId="1" pageNumber="105" pagination="1 - 614" refId="ref3676" refString="Wygodzinsky, P. W. (1966) A monograph of the Emesinae (Reduviidae, Hemiptera). Bulletin of the American Museum of Natural History, 133, 1 - 614." type="journal article" year="1966">Wygodzinsky, 1966</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3727CA3FF1E2A6C4CC3F7AEFE8764C6" box="[151,288,1921,1947]" pageId="1" pageNumber="105" type="description">
<paragraph id="8BD72F28FF1E2A6C4CC3F7AEFE8764C6" blockId="1.[151,614,1884,1947]" box="[151,288,1921,1947]" pageId="1" pageNumber="105">
(
<figureCitation id="135333ADFF1E2A6C4CCBF7AEFEBF64C6" box="[159,280,1921,1947]" captionStart-0="FIGURES 25" captionStart-1="FIGURES 611" captionStart-2="FIGURES 1217" captionStart-3="FIGURES 18" captionStartId-0="2.[151,264,1943,1967]" captionStartId-1="3.[151,264,1422,1446]" captionStartId-2="5.[151,264,1908,1932]" captionStartId-3="6.[151,264,1454,1478]" captionTargetBox-0="[217,1370,181,1919]" captionTargetBox-1="[151,1436,181,1398]" captionTargetBox-2="[184,1404,181,1885]" captionTargetBox-3="[151,1436,181,1431]" captionTargetId-0="figure-17@2.[217,1370,181,1919]" captionTargetId-1="figure-32@3.[151,1436,181,1398]" captionTargetId-2="figure-32@5.[184,1404,181,1885]" captionTargetId-3="figure-17@6.[151,1436,181,1431]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURES 25. Hornylia nalanda Wygodzinsky, female. 2, habitus, lateral; 3, anterior part of body, lateral; 4, close up of head and pronotum, lateral; 5, habitus, dorsal." captionText-1="FIGURES 611. Hornylia nalanda Wygodzinsky, female. 6, abdomen, dorsal; 7, abdomen, ventral, inset showing 7th sternite with emargination and a blunt spine like projection; 811, abdomen apical area in dorsal, ventral, lateral, and posterior views, respectively." captionText-2="FIGURES 1217. Hornylia nalanda Wygodzinsky. female terminalia. 12, ventral view of tip of abdomen (KOH treated); 13, tergites 8 &amp; 9 in dorsal view slide mounted; 14. gonocoxa 8 with part of bursa copulatrix (ventral view); 15. gonocoxa 8 with attached gonapophysis; 16. bursa copulatrix (ventral view); 17. gonapophysis 9 (red arrow) and gonoplac (black arrow). Abbreviations: gap—gonapophysis, gcx—gonocoxae, gpl—gonoplac, T8 and T9 tergites eight and nine." captionText-3="FIGURES 18­­­­­­­ 19. Hornylia nalanda Wygodzinsky, holotype male. 18, dorsal view; 19, lateral view." pageId="1" pageNumber="105">Figs. 219</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3727CA3FF1E2A6C4CC3F790FE4764A8" pageId="1" pageNumber="105" type="reference_group">
<paragraph id="8BD72F28FF1E2A6C4CC3F790FE4764A8" blockId="1.[151,1437,1983,2037]" pageId="1" pageNumber="105">
<treatmentCitationGroup id="AB780806FF1E2A6C4CC3F790FE4764A8" pageId="1" pageNumber="105">
<taxonomicName id="4C6854ABFF1E2A6C4CC3F790FD84648A" ID-CoL="3MQ3Y" authority="Wygodzinsky, 1966" authorityName="Wygodzinsky" authorityYear="1966" box="[151,547,1983,2007]" class="Insecta" family="Reduviidae" genus="Hornylia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="105" phylum="Arthropoda" rank="species" species="nalanda">
<emphasis id="B91CF33AFF1E2A6C4CC3F790FEE8648A" box="[151,335,1983,2007]" italics="true" pageId="1" pageNumber="105">Hornylia nalanda</emphasis>
<bibRefCitation id="EFF952D9FF1E2A6C4D03F790FD84648A" author="Wygodzinsky, P. W." box="[343,547,1983,2007]" pageId="1" pageNumber="105" pagination="1 - 614" refId="ref3676" refString="Wygodzinsky, P. W. (1966) A monograph of the Emesinae (Reduviidae, Hemiptera). Bulletin of the American Museum of Natural History, 133, 1 - 614." type="journal article" year="1966">Wygodzinsky, 1966</bibRefCitation>
</taxonomicName>
(494: original description),
<treatmentCitation id="0AC90939FF1E2A6C4F1EF790FB31648A" author="Maldonado Capriles, J." box="[842,1174,1983,2007]" page="131" pageId="1" pageNumber="105" year="1990">
<bibRefCitation id="EFF952D9FF1E2A6C4F1EF790FB31648A" author="Maldonado Capriles, J." box="[842,1174,1983,2007]" pageId="1" pageNumber="105" refId="ref3486" refString="Maldonado Capriles, J. (1990) Systematic Catalogue of the Reduviidae of the World (Insecta: Heteroptera). A special edition of Caribbean Journal of Science, Mayaguez, x + 694 pp." type="book" year="1990">Maldonado Capriles (1990: 131</bibRefCitation>
</treatmentCitation>
, catalogued),
<treatmentCitation id="0AC90939FF1E2A6C497DF790FE9F64A8" author="Chen, Z. &amp; Li, H. &amp; Cai, W." page="107" pageId="1" pageNumber="105" year="2020">
<bibRefCitation id="EFF952D9FF1E2A6C497DF790FE9F64A8" author="Chen, Z. &amp; Li, H. &amp; Cai, W." pageId="1" pageNumber="105" pagination="105 - 115" refId="ref3369" refString="Chen, Z., Li, H. &amp; Cai, W. (2020) A new species of Hornylia Wygodzinsky (Hemiptera, Heteroptera, Reduviidae, Emesinae) from Thailand. ZooKeys, 917, 105 - 115. https: // doi. org / 10.3897 / zookeys. 917.46887" type="journal article" year="2020">
Chen
<emphasis id="B91CF33AFF1E2A6C4933F790FA3A648A" box="[1383,1437,1983,2007]" italics="true" pageId="1" pageNumber="105">et al.</emphasis>
(2020: 107
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</treatmentCitation>
, key to species).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="DF177FA0FF1D2A6F4CC3F7B8FD95648E" pageId="2" pageNumber="106" startId="2.[151,264,1943,1967]" targetBox="[217,1370,181,1919]" targetPageId="2" targetType="figure">
<paragraph id="8BD72F28FF1D2A6F4CC3F7B8FD95648E" blockId="2.[151,1436,1943,2003]" pageId="2" pageNumber="106">
<emphasis id="B91CF33AFF1D2A6F4CC3F7B8FD4664F2" bold="true" box="[151,737,1943,1967]" pageId="2" pageNumber="106">
FIGURES 25.
<taxonomicName id="4C6854ABFF1D2A6F4D6BF7B8FD3764F2" authority="Wygodzinsky" authorityName="Wygodzinsky" box="[319,656,1943,1967]" class="Insecta" family="Reduviidae" genus="Hornylia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="106" phylum="Arthropoda" rank="species" species="nalanda">
<emphasis id="B91CF33AFF1D2A6F4D6BF7B8FE5F64F2" bold="true" box="[319,504,1943,1967]" italics="true" pageId="2" pageNumber="106">Hornylia nalanda</emphasis>
Wygodzinsky
</taxonomicName>
, female
</emphasis>
. 2, habitus, lateral; 3, anterior part of body, lateral; 4, close up of head and pronotum, lateral; 5, habitus, dorsal.
</paragraph>
</caption>
<caption id="DF177FA0FF1C2A6E4CC3F5A1FEB066B3" pageId="3" pageNumber="107" startId="3.[151,264,1422,1446]" targetBox="[151,1436,181,1398]" targetPageId="3" targetType="figure">
<paragraph id="8BD72F28FF1C2A6E4CC3F5A1FEB066B3" blockId="3.[151,1437,1422,1518]" pageId="3" pageNumber="107">
<emphasis id="B91CF33AFF1C2A6E4CC3F5A1FD4966FB" bold="true" box="[151,750,1422,1446]" pageId="3" pageNumber="107">
FIGURES 611.
<taxonomicName id="4C6854ABFF1C2A6E4D1FF5A1FD3B66FB" authority="Wygodzinsky" authorityName="Wygodzinsky" box="[331,668,1422,1446]" class="Insecta" family="Reduviidae" genus="Hornylia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="nalanda">
<emphasis id="B91CF33AFF1C2A6E4D1FF5A1FDA266FB" bold="true" box="[331,517,1422,1446]" italics="true" pageId="3" pageNumber="107">Hornylia nalanda</emphasis>
Wygodzinsky
</taxonomicName>
, female
</emphasis>
. 6, abdomen, dorsal; 7, abdomen, ventral, inset showing 7th sternite with emargination and a blunt spine like projection; 811, abdomen apical area in dorsal, ventral, lateral, and posterior views, respectively.
</paragraph>
</caption>
<subSubSection id="C3727CA3FF1C2A6E4CC3F66EFB2065B6" pageId="3" pageNumber="107" type="materials_examined">
<paragraph id="8BD72F28FF1C2A6E4CC3F66EFD7165FE" blockId="3.[151,1436,1600,2023]" pageId="3" pageNumber="107">
<emphasis id="B91CF33AFF1C2A6E4CC3F66EFD8F6507" bold="true" box="[151,552,1600,1627]" pageId="3" pageNumber="107">Type specimen images examined.</emphasis>
<materialsCitation id="3B002575FF1C2A6E4E65F66FFD7165FE" collectionCode="AMNH" country="United States of America" location="American Museum of Natural History" pageId="3" pageNumber="107" specimenCode="IZC 00321017" specimenCount="1" stateProvince="New York" typeStatus="holotype">
<typeStatus id="54D3918AFF1C2A6E4E65F66FFD3E6507" box="[561,665,1600,1626]" pageId="3" pageNumber="107" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C6854ABFF1C2A6E4E96F66FFBD06507" authority="Wygodzinsky, 1966" authorityName="Wygodzinsky" authorityYear="1966" box="[706,1143,1600,1626]" class="Insecta" family="Reduviidae" genus="Hornylia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="nalanda">
<emphasis id="B91CF33AFF1C2A6E4E96F66FFC2B6507" box="[706,908,1600,1626]" italics="true" pageId="3" pageNumber="107">Hornylia nalanda</emphasis>
<bibRefCitation id="EFF952D9FF1C2A6E4FC0F66FFBD06507" author="Wygodzinsky, P. W." box="[916,1143,1600,1626]" pageId="3" pageNumber="107" pagination="1 - 614" refId="ref3676" refString="Wygodzinsky, P. W. (1966) A monograph of the Emesinae (Reduviidae, Hemiptera). Bulletin of the American Museum of Natural History, 133, 1 - 614." type="journal article" year="1966">Wygodzinsky, 1966</bibRefCitation>
</taxonomicName>
, apterous male,
<taxonomicName id="4C6854ABFF1C2A6E4967F66FFA3B6507" box="[1331,1436,1600,1626]" class="Insecta" family="Reduviidae" genus="Hornylia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="nalanda">“Nalanda</taxonomicName>
Ceylon Horn” (only examined images of this specimen, see
<figureCitation id="135333ADFF1C2A6E4F0AF64BFBA16522" box="[862,1030,1636,1663]" captionStart="FIGURES 18" captionStartId="6.[151,264,1454,1478]" captionTargetBox="[151,1436,181,1431]" captionTargetId="figure-17@6.[151,1436,181,1431]" captionTargetPageId="6" captionText="FIGURES 18­­­­­­­ 19. Hornylia nalanda Wygodzinsky, holotype male. 18, dorsal view; 19, lateral view." pageId="3" pageNumber="107">Figs. 18 &amp; 19</figureCitation>
), in
<location id="8EB779F3FF1C2A6E486BF64BFF4B65FF" LSID="urn:lsid:plazi:treatment:03C19E3EFF1E2A694CC3F773FE3564C8:8EB779F3FF1C2A6E486BF64BFF4B65FF" country="United States of America" name="American Museum of Natural History" pageId="3" pageNumber="107" stateProvince="New York">American Museum of Natural History</location>
,
<collectingRegion id="49ACE1CAFF1C2A6E4CA3F6A6FECB65FF" box="[247,364,1672,1699]" country="United States of America" name="New York" pageId="3" pageNumber="107">New York</collectingRegion>
,
<collectingCountry id="F37F6FB8FF1C2A6E4D23F6A6FE0A65FE" box="[375,429,1673,1699]" name="United States of America" pageId="3" pageNumber="107">USA</collectingCountry>
(
<collectionCode id="ED79B7EDFF1C2A6E4DE8F6A6FDB465FE" box="[444,531,1673,1699]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="3" pageNumber="107">AMNH</collectionCode>
_
<specimenCode id="DBCE8753FF1C2A6E4E75F6A6FD6D65FE" box="[545,714,1673,1699]" collectionCode="IZC" pageId="3" pageNumber="107">IZC 00321017</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BD72F28FF1C2A6E4C93F683FB2065B6" blockId="3.[151,1436,1600,2023]" pageId="3" pageNumber="107">
<emphasis id="B91CF33AFF1C2A6E4C93F683FDA5659B" bold="true" box="[199,514,1708,1734]" pageId="3" pageNumber="107">Other specimen examined</emphasis>
.
<materialsCitation id="3B002575FF1C2A6E4E46F682FB2465B6" collectingDate="2020-08-05" collectorName="A. P. Ranjith &amp; Pune" country="India" county="Malappuram" location="Zoological Survey of India" municipality="Calicut University Botanical Garden" pageId="3" pageNumber="107" specimenCount="1" specimenCount-female="1" stateProvince="Kerala">
<specimenCount id="9D6EE4A1FF1C2A6E4E46F682FD98659A" box="[530,575,1709,1735]" count="1" pageId="3" pageNumber="107" type="female">1 ♀</specimenCount>
.
<collectingCountry id="F37F6FB8FF1C2A6E4E1BF682FD04659A" box="[591,675,1709,1735]" name="India" pageId="3" pageNumber="107">INDIA</collectingCountry>
:
<collectingRegion id="49ACE1CAFF1C2A6E4EFBF683FD5B659B" box="[687,764,1708,1734]" country="India" name="Kerala" pageId="3" pageNumber="107">Kerala</collectingRegion>
,
<collectingCounty id="62B657A4FF1C2A6E4F5EF683FC3B659B" box="[778,924,1708,1734]" pageId="3" pageNumber="107">Malappuram</collectingCounty>
,
<collectingMunicipality id="6BB3B552FF1C2A6E4FFDF683FAEB659B" box="[937,1356,1708,1734]" pageId="3" pageNumber="107">Calicut University Botanical Garden</collectingMunicipality>
,
<collectingMethod id="5229573FFF1C2A6E490DF683FF7765B6" pageId="3" pageNumber="107">pitfall traps</collectingMethod>
,
<date id="FFD609E8FF1C2A6E4C88F6FFFEC165B6" box="[220,358,1744,1771]" pageId="3" pageNumber="107" value="2020-08-05">
<collectingDate id="EF92F000FF1C2A6E4C88F6FFFEC165B6" box="[220,358,1744,1771]" pageId="3" pageNumber="107" value="2020-08-05">05.viii.2020</collectingDate>
</date>
, coll.
<collectorName id="269D4AFEFF1C2A6E4DF3F6FEFD9565B7" box="[423,562,1744,1771]" pageId="3" pageNumber="107">A.P. Ranjith</collectorName>
, to be deposited in
<location id="8EB779F3FF1C2A6E4F5FF6FFFB9B65B7" LSID="urn:lsid:plazi:treatment:03C19E3EFF1E2A694CC3F773FE3564C8:8EB779F3FF1C2A6E4F5FF6FFFB9B65B7" box="[779,1084,1744,1771]" country="India" county="Malappuram" municipality="Calicut University Botanical Garden" name="Zoological Survey of India" pageId="3" pageNumber="107" stateProvince="Kerala">Zoological Survey of India</location>
,
<collectorName id="269D4AFEFF1C2A6E4813F6FEFB2465B6" box="[1095,1155,1745,1771]" pageId="3" pageNumber="107">Pune</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3727CA3FF1C2A694C93F6DBFE3564C8" lastPageId="4" lastPageNumber="108" pageId="3" pageNumber="107" type="description">
<paragraph id="8BD72F28FF1C2A6E4C93F6DBFEFE6453" blockId="3.[151,1436,1600,2023]" box="[199,345,1780,1806]" pageId="3" pageNumber="107">
<emphasis id="B91CF33AFF1C2A6E4C93F6DBFEFE6453" bold="true" box="[199,345,1780,1806]" pageId="3" pageNumber="107">Description.</emphasis>
</paragraph>
<paragraph id="8BD72F28FF1C2A6E4C93F737FD8D646E" blockId="3.[151,1436,1600,2023]" box="[199,554,1816,1843]" pageId="3" pageNumber="107">
<emphasis id="B91CF33AFF1C2A6E4C93F737FED1646E" bold="true" box="[199,374,1816,1843]" pageId="3" pageNumber="107">Apterous male</emphasis>
(
<figureCitation id="135333ADFF1C2A6E4DD1F737FD86646E" box="[389,545,1816,1843]" captionStart="FIGURES 18" captionStartId="6.[151,264,1454,1478]" captionTargetBox="[151,1436,181,1431]" captionTargetId="figure-17@6.[151,1436,181,1431]" captionTargetPageId="6" captionText="FIGURES 18­­­­­­­ 19. Hornylia nalanda Wygodzinsky, holotype male. 18, dorsal view; 19, lateral view." pageId="3" pageNumber="107">Figs. 18 &amp; 19</figureCitation>
)
</paragraph>
<paragraph id="8BD72F28FF1C2A6E4C93F712FAA2640A" blockId="3.[151,1436,1600,2023]" box="[199,1285,1852,1879]" pageId="3" pageNumber="107">
Described by
<bibRefCitation id="EFF952D9FF1C2A6E4D36F712FDE4640B" author="Wygodzinsky, P. W." box="[354,579,1852,1879]" pageId="3" pageNumber="107" pagination="1 - 614" refId="ref3676" refString="Wygodzinsky, P. W. (1966) A monograph of the Emesinae (Reduviidae, Hemiptera). Bulletin of the American Museum of Natural History, 133, 1 - 614." type="journal article" year="1966">Wygodzinsky, 1966</bibRefCitation>
, based on one apterous male (
<typeStatus id="54D3918AFF1C2A6E4FC6F712FC50640A" box="[914,1015,1853,1879]" pageId="3" pageNumber="107" type="holotype">holotype</typeStatus>
) specimen listed above.
</paragraph>
<paragraph id="8BD72F28FF1C2A6E4C93F74FFDBC6426" blockId="3.[151,1436,1600,2023]" box="[199,539,1888,1915]" pageId="3" pageNumber="107">
<emphasis id="B91CF33AFF1C2A6E4C93F74FFE2C6427" bold="true" box="[199,395,1888,1914]" pageId="3" pageNumber="107">Apterous female</emphasis>
(
<figureCitation id="135333ADFF1C2A6E4DCFF74FFDB46426" box="[411,531,1888,1915]" captionStart-0="FIGURES 25" captionStart-1="FIGURES 611" captionStart-2="FIGURES 1217" captionStartId-0="2.[151,264,1943,1967]" captionStartId-1="3.[151,264,1422,1446]" captionStartId-2="5.[151,264,1908,1932]" captionTargetBox-0="[217,1370,181,1919]" captionTargetBox-1="[151,1436,181,1398]" captionTargetBox-2="[184,1404,181,1885]" captionTargetId-0="figure-17@2.[217,1370,181,1919]" captionTargetId-1="figure-32@3.[151,1436,181,1398]" captionTargetId-2="figure-32@5.[184,1404,181,1885]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionText-0="FIGURES 25. Hornylia nalanda Wygodzinsky, female. 2, habitus, lateral; 3, anterior part of body, lateral; 4, close up of head and pronotum, lateral; 5, habitus, dorsal." captionText-1="FIGURES 611. Hornylia nalanda Wygodzinsky, female. 6, abdomen, dorsal; 7, abdomen, ventral, inset showing 7th sternite with emargination and a blunt spine like projection; 811, abdomen apical area in dorsal, ventral, lateral, and posterior views, respectively." captionText-2="FIGURES 1217. Hornylia nalanda Wygodzinsky. female terminalia. 12, ventral view of tip of abdomen (KOH treated); 13, tergites 8 &amp; 9 in dorsal view slide mounted; 14. gonocoxa 8 with part of bursa copulatrix (ventral view); 15. gonocoxa 8 with attached gonapophysis; 16. bursa copulatrix (ventral view); 17. gonapophysis 9 (red arrow) and gonoplac (black arrow). Abbreviations: gap—gonapophysis, gcx—gonocoxae, gpl—gonoplac, T8 and T9 tergites eight and nine." pageId="3" pageNumber="107">Figs. 217</figureCitation>
)
</paragraph>
<paragraph id="8BD72F28FF1C2A694C93F7ABFC7B6240" blockId="3.[151,1436,1600,2023]" lastBlockId="4.[151,1437,151,1941]" lastPageId="4" lastPageNumber="108" pageId="3" pageNumber="107">
<emphasis id="B91CF33AFF1C2A6E4C93F7ABFEF764C3" box="[199,336,1924,1950]" italics="true" pageId="3" pageNumber="107">Colouration</emphasis>
(
<figureCitation id="135333ADFF1C2A6E4D0AF7ABFE6D64C2" box="[350,458,1924,1951]" captionStart="FIGURES 25" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[217,1370,181,1919]" captionTargetId="figure-17@2.[217,1370,181,1919]" captionTargetPageId="2" captionText="FIGURES 25. Hornylia nalanda Wygodzinsky, female. 2, habitus, lateral; 3, anterior part of body, lateral; 4, close up of head and pronotum, lateral; 5, habitus, dorsal." pageId="3" pageNumber="107">Figs. 25</figureCitation>
): Head brownish but laterally dark brown; labium pale brown but visible second segment dark. Antennae with scape and pedicel dark, remaining two antennomeres very pale. Thorax brownish, laterally and ventrally dark brown. Abdomen brown with patches of dark brown on tergites, connexivum dark with anterior onefourth of each segment pale, underside very dark brown to piceous. All legs pale brown with some indistinct fuscous areas. Fore femora with very faint brownish areas that do not appear as well-defined annulations; fore tibia with basal and apical dark areas; fore tarsus darker in basal half. Mid- and hind legs pale brown, their femora with apical fuscous area preceded by indistinct pale area; tibia with dark apex (
<figureCitation id="135333ADFF1B2A694FDDF12CFC686240" box="[905,975,259,285]" captionStart="FIGURES 25" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[217,1370,181,1919]" captionTargetId="figure-17@2.[217,1370,181,1919]" captionTargetPageId="2" captionText="FIGURES 25. Hornylia nalanda Wygodzinsky, female. 2, habitus, lateral; 3, anterior part of body, lateral; 4, close up of head and pronotum, lateral; 5, habitus, dorsal." pageId="4" pageNumber="108">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8BD72F28FF1B2A694C93F108FBD162F0" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="108">
<emphasis id="B91CF33AFF1B2A694C93F108FE89621C" box="[199,302,295,321]" italics="true" pageId="4" pageNumber="108">Structure</emphasis>
(
<figureCitation id="135333ADFF1B2A694D69F108FE1E621C" box="[317,441,295,321]" captionStart-0="FIGURES 25" captionStart-1="FIGURES 611" captionStart-2="FIGURES 1217" captionStartId-0="2.[151,264,1943,1967]" captionStartId-1="3.[151,264,1422,1446]" captionStartId-2="5.[151,264,1908,1932]" captionTargetBox-0="[217,1370,181,1919]" captionTargetBox-1="[151,1436,181,1398]" captionTargetBox-2="[184,1404,181,1885]" captionTargetId-0="figure-17@2.[217,1370,181,1919]" captionTargetId-1="figure-32@3.[151,1436,181,1398]" captionTargetId-2="figure-32@5.[184,1404,181,1885]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionText-0="FIGURES 25. Hornylia nalanda Wygodzinsky, female. 2, habitus, lateral; 3, anterior part of body, lateral; 4, close up of head and pronotum, lateral; 5, habitus, dorsal." captionText-1="FIGURES 611. Hornylia nalanda Wygodzinsky, female. 6, abdomen, dorsal; 7, abdomen, ventral, inset showing 7th sternite with emargination and a blunt spine like projection; 811, abdomen apical area in dorsal, ventral, lateral, and posterior views, respectively." captionText-2="FIGURES 1217. Hornylia nalanda Wygodzinsky. female terminalia. 12, ventral view of tip of abdomen (KOH treated); 13, tergites 8 &amp; 9 in dorsal view slide mounted; 14. gonocoxa 8 with part of bursa copulatrix (ventral view); 15. gonocoxa 8 with attached gonapophysis; 16. bursa copulatrix (ventral view); 17. gonapophysis 9 (red arrow) and gonoplac (black arrow). Abbreviations: gap—gonapophysis, gcx—gonocoxae, gpl—gonoplac, T8 and T9 tergites eight and nine." pageId="4" pageNumber="108">Figs. 217</figureCitation>
):
<emphasis id="B91CF33AFF1B2A694D98F108FDA1621C" box="[460,518,295,321]" italics="true" pageId="4" pageNumber="108">Body</emphasis>
slender, surface dull, covered with granules. Head distinctly granular dorsally and laterally. Pronotum with scattered granules dorsally and laterally. Mesonotum and metanotum with sparse granules only laterally. Abdomen with sparse, scattered granules on dorsal aspect. Fine, short, erect setae present on fore femur and fore tibia ventrally. Very small, adpressed, spine-like setae present on all legs.
</paragraph>
<paragraph id="8BD72F28FF1B2A694C93F198FC8761D8" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="108">
<emphasis id="B91CF33AFF1B2A694C93F198FEAA628C" box="[199,269,439,465]" italics="true" pageId="4" pageNumber="108">Head:</emphasis>
Head longer than broad. Eyes very small, not reaching dorsal or ventral margin of head; transverse interocular sulcus distinct and situated just behind middle of eye; anteocular region twice as long as postocular in lateral view. Antennae 4-segmented, antenniferous tubercles situated about middle of anteocular area; scape longest. Labium straight, four segmented, visible first segment more than twice as long as second visible segment but not reaching anterior margin of eye in lateral view; first two visible segments thicker, visible third long and slender (
<figureCitation id="135333ADFF1B2A69493BF268FF0F61D8" captionStart="FIGURES 25" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[217,1370,181,1919]" captionTargetId="figure-17@2.[217,1370,181,1919]" captionTargetPageId="2" captionText="FIGURES 25. Hornylia nalanda Wygodzinsky, female. 2, habitus, lateral; 3, anterior part of body, lateral; 4, close up of head and pronotum, lateral; 5, habitus, dorsal." pageId="4" pageNumber="108">Fig. 4</figureCitation>
). A short, flap-like forward extension of labrum present.
</paragraph>
<paragraph id="8BD72F28FF1B2A694C93F2A0FBE56048" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="108">
<emphasis id="B91CF33AFF1B2A694C93F2A0FEBF61F4" box="[199,280,655,681]" italics="true" pageId="4" pageNumber="108">Thorax</emphasis>
: Pronotum longer than mesonotum, mesonotum longer than metanotum (
<figureCitation id="135333ADFF1B2A694832F2A0FB0A61F4" box="[1126,1197,654,681]" captionStart="FIGURES 25" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[217,1370,181,1919]" captionTargetId="figure-17@2.[217,1370,181,1919]" captionTargetPageId="2" captionText="FIGURES 25. Hornylia nalanda Wygodzinsky, female. 2, habitus, lateral; 3, anterior part of body, lateral; 4, close up of head and pronotum, lateral; 5, habitus, dorsal." pageId="4" pageNumber="108">Fig. 5</figureCitation>
). Pronotum broadest at anterior angles, gradually narrowed posteriorly; anterior lobe and posterior lobes not distinct from each other, latter very small and separated by very shallow transverse sulcus (
<figureCitation id="135333ADFF1B2A694F21F2F8FC1D61AC" box="[885,954,727,753]" captionStart="FIGURES 25" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[217,1370,181,1919]" captionTargetId="figure-17@2.[217,1370,181,1919]" captionTargetPageId="2" captionText="FIGURES 25. Hornylia nalanda Wygodzinsky, female. 2, habitus, lateral; 3, anterior part of body, lateral; 4, close up of head and pronotum, lateral; 5, habitus, dorsal." pageId="4" pageNumber="108">Fig. 4</figureCitation>
). Mesonotum and metanotum with delicate longitudinal median carina, lateral margins of meso- and metanotum raised (
<figureCitation id="135333ADFF1B2A694FBBF2D4FB936048" box="[1007,1076,762,789]" captionStart="FIGURES 25" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[217,1370,181,1919]" captionTargetId="figure-17@2.[217,1370,181,1919]" captionTargetPageId="2" captionText="FIGURES 25. Hornylia nalanda Wygodzinsky, female. 2, habitus, lateral; 3, anterior part of body, lateral; 4, close up of head and pronotum, lateral; 5, habitus, dorsal." pageId="4" pageNumber="108">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="8BD72F28FF1B2A694C93F330FC9B6720" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="108">
<emphasis id="B91CF33AFF1B2A694C93F330FEA16065" box="[199,262,799,824]" italics="true" pageId="4" pageNumber="108">Legs:</emphasis>
Fore coxae slender and long, parallel sided, almost as long as pronotum, without any spines or spiniferous processes ventrally; trochanters smooth, without spiniferous process. Fore femora ventrally with three series of spiniferous processes: anteroventral, posteroventral and an accessory series; there are in all six prominent posteroventral spiniferous processes—first process situated close to base longer than rest and longer than diameter of femur, but without penicillate process at tip; remaining five processes are shorter than diameter of femur; there are also many very small processes in between these long and short processes. Anteroventral series is made up of few medium sized processes and many much smaller processes, its first two basal processes are widely separated. Fore tibia less than half as long as femur, slightly dilated apically, with few denticles ventrally; fore tarsus unsegmented, subequal to tibia, gradually tapering, sclerotised, its ventral surface with adpressed setae (
<figureCitation id="135333ADFF1B2A694820F410FB106704" box="[1140,1207,1087,1113]" captionStart="FIGURES 25" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[217,1370,181,1919]" captionTargetId="figure-17@2.[217,1370,181,1919]" captionTargetPageId="2" captionText="FIGURES 25. Hornylia nalanda Wygodzinsky, female. 2, habitus, lateral; 3, anterior part of body, lateral; 4, close up of head and pronotum, lateral; 5, habitus, dorsal." pageId="4" pageNumber="108">Fig. 3</figureCitation>
). Mid- and hind legs very slender (
<figureCitation id="135333ADFF1B2A694D64F44CFED36720" box="[304,372,1123,1149]" captionStart="FIGURES 25" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[217,1370,181,1919]" captionTargetId="figure-17@2.[217,1370,181,1919]" captionTargetPageId="2" captionText="FIGURES 25. Hornylia nalanda Wygodzinsky, female. 2, habitus, lateral; 3, anterior part of body, lateral; 4, close up of head and pronotum, lateral; 5, habitus, dorsal." pageId="4" pageNumber="108">Fig. 2</figureCitation>
), hind femora passing apex of abdomen.
</paragraph>
<paragraph id="8BD72F28FF1B2A694C93F4A8FDD965E0" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="108">
<emphasis id="B91CF33AFF1B2A694C93F4A8FE9667FC" box="[199,305,1159,1185]" italics="true" pageId="4" pageNumber="108">Abdomen</emphasis>
: Narrow at base, gradually dilated posteriorly, broadest beyond middle and again slightly narrowed towards tip; connexivum narrow (
<figureCitation id="135333ADFF1B2A694E49F484FDC86798" box="[541,623,1194,1221]" captionStart="FIGURES 25" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[217,1370,181,1919]" captionTargetId="figure-17@2.[217,1370,181,1919]" captionTargetPageId="2" captionText="FIGURES 25. Hornylia nalanda Wygodzinsky, female. 2, habitus, lateral; 3, anterior part of body, lateral; 4, close up of head and pronotum, lateral; 5, habitus, dorsal." pageId="4" pageNumber="108">Figs. 5</figureCitation>
,
<figureCitation id="135333ADFF1B2A694E29F484FD2A6798" box="[637,653,1195,1221]" captionStart="FIGURES 611" captionStartId="3.[151,264,1422,1446]" captionTargetBox="[151,1436,181,1398]" captionTargetId="figure-32@3.[151,1436,181,1398]" captionTargetPageId="3" captionText="FIGURES 611. Hornylia nalanda Wygodzinsky, female. 6, abdomen, dorsal; 7, abdomen, ventral, inset showing 7th sternite with emargination and a blunt spine like projection; 811, abdomen apical area in dorsal, ventral, lateral, and posterior views, respectively." pageId="4" pageNumber="108">6</figureCitation>
). Basal (or second) abdominal sternite with small membranous area at base, distinct median keel visible from second to sixth segments ventrally, seventh sternite depressed ventrodorsally, with emargination around blunt median spine (
<figureCitation id="135333ADFF1B2A694F44F4DCFCF26650" box="[784,853,1267,1293]" captionStart="FIGURES 611" captionStartId="3.[151,264,1422,1446]" captionTargetBox="[151,1436,181,1398]" captionTargetId="figure-32@3.[151,1436,181,1398]" captionTargetPageId="3" captionText="FIGURES 611. Hornylia nalanda Wygodzinsky, female. 6, abdomen, dorsal; 7, abdomen, ventral, inset showing 7th sternite with emargination and a blunt spine like projection; 811, abdomen apical area in dorsal, ventral, lateral, and posterior views, respectively." pageId="4" pageNumber="108">Fig. 7</figureCitation>
).
<emphasis id="B91CF33AFF1B2A694F38F4DCFB8C6650" box="[876,1067,1267,1293]" italics="true" pageId="4" pageNumber="108">Female genitalia</emphasis>
(
<figureCitation id="135333ADFF1B2A69486FF4DCFB106650" box="[1083,1207,1267,1293]" captionStart-0="FIGURES 611" captionStart-1="FIGURES 1217" captionStartId-0="3.[151,264,1422,1446]" captionStartId-1="5.[151,264,1908,1932]" captionTargetBox-0="[151,1436,181,1398]" captionTargetBox-1="[184,1404,181,1885]" captionTargetId-0="figure-32@3.[151,1436,181,1398]" captionTargetId-1="figure-32@5.[184,1404,181,1885]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionText-0="FIGURES 611. Hornylia nalanda Wygodzinsky, female. 6, abdomen, dorsal; 7, abdomen, ventral, inset showing 7th sternite with emargination and a blunt spine like projection; 811, abdomen apical area in dorsal, ventral, lateral, and posterior views, respectively." captionText-1="FIGURES 1217. Hornylia nalanda Wygodzinsky. female terminalia. 12, ventral view of tip of abdomen (KOH treated); 13, tergites 8 &amp; 9 in dorsal view slide mounted; 14. gonocoxa 8 with part of bursa copulatrix (ventral view); 15. gonocoxa 8 with attached gonapophysis; 16. bursa copulatrix (ventral view); 17. gonapophysis 9 (red arrow) and gonoplac (black arrow). Abbreviations: gap—gonapophysis, gcx—gonocoxae, gpl—gonoplac, T8 and T9 tergites eight and nine." pageId="4" pageNumber="108">Figs. 817</figureCitation>
): terminalia (in situ, not dissected) in dorsal, ventral, lateral and posterior or caudal view are shown in
<figureCitation id="135333ADFF1B2A69484AF538FB32666C" box="[1054,1173,1303,1329]" captionStart="FIGURES 611" captionStartId="3.[151,264,1422,1446]" captionTargetBox="[151,1436,181,1398]" captionTargetId="figure-32@3.[151,1436,181,1398]" captionTargetPageId="3" captionText="FIGURES 611. Hornylia nalanda Wygodzinsky, female. 6, abdomen, dorsal; 7, abdomen, ventral, inset showing 7th sternite with emargination and a blunt spine like projection; 811, abdomen apical area in dorsal, ventral, lateral, and posterior views, respectively." pageId="4" pageNumber="108">Figs. 811</figureCitation>
. Eighth tergite slanting, rectangular with outer posterior angles bluntly prominent; ninth tergite almost vertical, its base partly covered under eighth, deeply emarginate at posterior border; setose posterior border of syngonapophysis (gonoplac) visible (
<figureCitation id="135333ADFF1B2A694913F570FA306624" box="[1351,1431,1375,1401]" captionStart="FIGURES 611" captionStartId="3.[151,264,1422,1446]" captionTargetBox="[151,1436,181,1398]" captionTargetId="figure-32@3.[151,1436,181,1398]" captionTargetPageId="3" captionText="FIGURES 611. Hornylia nalanda Wygodzinsky, female. 6, abdomen, dorsal; 7, abdomen, ventral, inset showing 7th sternite with emargination and a blunt spine like projection; 811, abdomen apical area in dorsal, ventral, lateral, and posterior views, respectively." pageId="4" pageNumber="108">Fig. 11</figureCitation>
; large gonocoxae of eighth segment seen laterally in caudal view of tip of abdomen (
<figureCitation id="135333ADFF1B2A694815F5ACFB3566C0" box="[1089,1170,1411,1437]" captionStart="FIGURES 611" captionStartId="3.[151,264,1422,1446]" captionTargetBox="[151,1436,181,1398]" captionTargetId="figure-32@3.[151,1436,181,1398]" captionTargetPageId="3" captionText="FIGURES 611. Hornylia nalanda Wygodzinsky, female. 6, abdomen, dorsal; 7, abdomen, ventral, inset showing 7th sternite with emargination and a blunt spine like projection; 811, abdomen apical area in dorsal, ventral, lateral, and posterior views, respectively." pageId="4" pageNumber="108">Fig. 11</figureCitation>
). At low magnification, KOH cleared tip of abdomen shows large, sclerotised gonocoxae 8 (gcx8) and associated gonapophyses 8 (gap8) as well as part of gonoplac (gpl) and posterior border of tergite 9 (T9) (
<figureCitation id="135333ADFF1B2A694FDFF5E4FC7966B8" box="[907,990,1483,1509]" captionStart="FIGURES 1217" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[184,1404,181,1885]" captionTargetId="figure-32@5.[184,1404,181,1885]" captionTargetPageId="5" captionText="FIGURES 1217. Hornylia nalanda Wygodzinsky. female terminalia. 12, ventral view of tip of abdomen (KOH treated); 13, tergites 8 &amp; 9 in dorsal view slide mounted; 14. gonocoxa 8 with part of bursa copulatrix (ventral view); 15. gonocoxa 8 with attached gonapophysis; 16. bursa copulatrix (ventral view); 17. gonapophysis 9 (red arrow) and gonoplac (black arrow). Abbreviations: gap—gonapophysis, gcx—gonocoxae, gpl—gonoplac, T8 and T9 tergites eight and nine." pageId="4" pageNumber="108">Fig. 12</figureCitation>
); at higher magnification slide-mounted tergites 8 and 9 show sparse punctures and setae as well as two tubercular areas and deep emargination of posterior margin of ninth tergite (
<figureCitation id="135333ADFF1B2A694DFBF63CFDA26570" box="[431,517,1555,1581]" captionStart="FIGURES 1217" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[184,1404,181,1885]" captionTargetId="figure-32@5.[184,1404,181,1885]" captionTargetPageId="5" captionText="FIGURES 1217. Hornylia nalanda Wygodzinsky. female terminalia. 12, ventral view of tip of abdomen (KOH treated); 13, tergites 8 &amp; 9 in dorsal view slide mounted; 14. gonocoxa 8 with part of bursa copulatrix (ventral view); 15. gonocoxa 8 with attached gonapophysis; 16. bursa copulatrix (ventral view); 17. gonapophysis 9 (red arrow) and gonoplac (black arrow). Abbreviations: gap—gonapophysis, gcx—gonocoxae, gpl—gonoplac, T8 and T9 tergites eight and nine." pageId="4" pageNumber="108">Fig. 13</figureCitation>
). Gonocoxa 8 along with part of bursa copulatrix is shown in
<figureCitation id="135333ADFF1B2A6948B4F63CFA926570" box="[1248,1333,1555,1581]" captionStart="FIGURES 1217" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[184,1404,181,1885]" captionTargetId="figure-32@5.[184,1404,181,1885]" captionTargetPageId="5" captionText="FIGURES 1217. Hornylia nalanda Wygodzinsky. female terminalia. 12, ventral view of tip of abdomen (KOH treated); 13, tergites 8 &amp; 9 in dorsal view slide mounted; 14. gonocoxa 8 with part of bursa copulatrix (ventral view); 15. gonocoxa 8 with attached gonapophysis; 16. bursa copulatrix (ventral view); 17. gonapophysis 9 (red arrow) and gonoplac (black arrow). Abbreviations: gap—gonapophysis, gcx—gonocoxae, gpl—gonoplac, T8 and T9 tergites eight and nine." pageId="4" pageNumber="108">Fig. 14</figureCitation>
. Isolated gonocoxa 8 appears pentagonal and more sclerotised (
<figureCitation id="135333ADFF1B2A694EA1F618FCE0650C" box="[757,839,1590,1617]" captionStart="FIGURES 1217" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[184,1404,181,1885]" captionTargetId="figure-32@5.[184,1404,181,1885]" captionTargetPageId="5" captionText="FIGURES 1217. Hornylia nalanda Wygodzinsky. female terminalia. 12, ventral view of tip of abdomen (KOH treated); 13, tergites 8 &amp; 9 in dorsal view slide mounted; 14. gonocoxa 8 with part of bursa copulatrix (ventral view); 15. gonocoxa 8 with attached gonapophysis; 16. bursa copulatrix (ventral view); 17. gonapophysis 9 (red arrow) and gonoplac (black arrow). Abbreviations: gap—gonapophysis, gcx—gonocoxae, gpl—gonoplac, T8 and T9 tergites eight and nine." pageId="4" pageNumber="108">Fig. 15</figureCitation>
) while gonapophysis 8 is triangular with sparse setae distally (posteriorly). Bursa copulatrix appears like a squarish sac with some sclerotised areas / sclerites proximally (
<figureCitation id="135333ADFF1B2A694CCBF650FF5465C4" box="[159,243,1663,1689]" captionStart="FIGURES 1217" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[184,1404,181,1885]" captionTargetId="figure-32@5.[184,1404,181,1885]" captionTargetPageId="5" captionText="FIGURES 1217. Hornylia nalanda Wygodzinsky. female terminalia. 12, ventral view of tip of abdomen (KOH treated); 13, tergites 8 &amp; 9 in dorsal view slide mounted; 14. gonocoxa 8 with part of bursa copulatrix (ventral view); 15. gonocoxa 8 with attached gonapophysis; 16. bursa copulatrix (ventral view); 17. gonapophysis 9 (red arrow) and gonoplac (black arrow). Abbreviations: gap—gonapophysis, gcx—gonocoxae, gpl—gonoplac, T8 and T9 tergites eight and nine." pageId="4" pageNumber="108">Fig. 16</figureCitation>
). Gonapophysis 9 with partly sclerotised median margin (arrow) and gonoplac with long setae at posterior margin are shown in ventral view (
<figureCitation id="135333ADFF1B2A694E4AF68CFDD665E0" box="[542,625,1699,1725]" captionStart="FIGURES 1217" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[184,1404,181,1885]" captionTargetId="figure-32@5.[184,1404,181,1885]" captionTargetPageId="5" captionText="FIGURES 1217. Hornylia nalanda Wygodzinsky. female terminalia. 12, ventral view of tip of abdomen (KOH treated); 13, tergites 8 &amp; 9 in dorsal view slide mounted; 14. gonocoxa 8 with part of bursa copulatrix (ventral view); 15. gonocoxa 8 with attached gonapophysis; 16. bursa copulatrix (ventral view); 17. gonapophysis 9 (red arrow) and gonoplac (black arrow). Abbreviations: gap—gonapophysis, gcx—gonocoxae, gpl—gonoplac, T8 and T9 tergites eight and nine." pageId="4" pageNumber="108">Fig. 17</figureCitation>
).
</paragraph>
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<emphasis id="B91CF33AFF1B2A694C93F6E8FECE65BD" box="[199,361,1735,1760]" italics="true" pageId="4" pageNumber="108">Measurements</emphasis>
(in mm): Total length 8.00. Head length 1.08; anteocular length 0.63; postocular length 0.25; eye diameter laterally 0.23. Labium: I visible segment 0.50, II visible segment 0.20, III visible segment 0.30. Antenna: I segment 5.00, II segment 3.25, segments III and IV missing. Thorax: pronotum length 1.35, mesonotum 0.75, metanotum 0.55; pronotal width at anterolateral angles 0.63, width at humeral angles 0.40; foreleg: coxa 1.55, femur 2.25, tibia 1.00, tarsus 0.80; mid leg: coxa 0.50, femur 3.8, tibia 5.00; hind leg: coxa 0.50, femur 5.50, tibia 7.50. Abdomen length 4.25.
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(
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<emphasis id="B918462EFFCC6F5EFF199083C3D2FB7A" box="[151,415,1231,1257]" italics="true" pageId="1" pageNumber="296">Varicopsella luzonensis</emphasis>
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(
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Distribution.
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Distribution.
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Distribution.
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Distribution.
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<emphasis id="B918462EFFCC6F5EFF19969BC3EEFD62" bold="true" box="[151,419,727,754]" italics="true" pageId="1" pageNumber="296">Varicopsella cornifera</emphasis>
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Distribution.
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<emphasis id="B918462EFFCC6F5EFF19970FC3FAFCCD" bold="true" box="[151,439,835,862]" italics="true" pageId="1" pageNumber="296">Varicopsella davaoensis</emphasis>
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<emphasis id="B918462EFFCC6F5EFF19972BC3EBFC12" box="[151,422,871,897]" italics="true" pageId="1" pageNumber="296">Varicopsella davaoensis</emphasis>
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<emphasis id="B918462EFFCC6F5EFF1997C7C3EBFC36" box="[151,422,907,933]" italics="true" pageId="1" pageNumber="296">Varicopsella davaoensis</emphasis>
(
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<bibRefCitation id="EFFDE7CDFFCC6F5EFDE997C7C11EFC36" author="Hamilton, K. G. A." box="[615,851,907,933]" pageId="1" pageNumber="296" pagination="875 - 932" refId="ref3094" refString="Hamilton, K. G. A. (1980) Contributions to the study of the world Macropsini (Rhynchota: Homoptera: Cicadellidae). The Canadian Entomologist, 112 (9), 875 - 932. https: // doi. org / 10.4039 / Ent 112875 - 9" type="journal article" year="1980">Hamilton, 1980: 900</bibRefCitation>
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Distribution.
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<paragraph id="8BD39A3CFFCC6F5EFF199057C0DDFBA6" blockId="1.[151,1003,1015,1149]" box="[151,656,1051,1077]" pageId="1" pageNumber="296">
<treatmentCitationGroup id="AB7CBD12FFCC6F5EFF199057C0DDFBA6" box="[151,656,1051,1077]" pageId="1" pageNumber="296">
<treatmentCitation id="0ACDBC2DFFCC6F5EFF199057C0C6FBA6" author="Viraktamath, C. A." box="[151,651,1051,1077]" page="306" pageId="1" pageNumber="296" year="1981">
<taxonomicName id="4C6CE1BFFFCC6F5EFF199057C0C6FBA6" ID-CoL="7FHXH" authority="Viraktamath 1981: 306" authorityName="Viraktamath" authorityPageNumber="306" authorityYear="1981" box="[151,651,1051,1077]" class="Insecta" family="Cicadellidae" genus="Varicopsella" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="296" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="B918462EFFCC6F5EFF199057C3CDFBA6" box="[151,384,1051,1077]" italics="true" pageId="1" pageNumber="296">Varicopsella elegans</emphasis>
<bibRefCitation id="EFFDE7CDFFCC6F5EFE089057C0C6FBA6" author="Viraktamath, C. A." box="[390,651,1051,1077]" pageId="1" pageNumber="296" pagination="295 - 310" refId="ref3494" refString="Viraktamath, C. A. (1981) Indian Macropsinae (Homoptera: Cicadellidae). II. Species described by W. L. Distant and descriptions of new species from the Indian subcontinent. Entomologica Scandinavica, 12, 295 - 310. https: // doi. org / 10.1163 / 187631281794709791" type="journal article" year="1981">Viraktamath 1981: 306</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD39A3CFFCC6F5EFF199073C1A6FBCA" blockId="1.[151,1003,1015,1149]" box="[151,1003,1087,1114]" pageId="1" pageNumber="296">
<treatmentCitationGroup id="AB7CBD12FFCC6F5EFF199073C1A6FBCA" box="[151,1003,1087,1114]" pageId="1" pageNumber="296">
<taxonomicName id="4C6CE1BFFFCC6F5EFF199073C01FFBCA" ID-CoL="7FHXH" authority="Viraktamath 1981" authorityName="Viraktamath" authorityYear="1981" box="[151,594,1087,1113]" class="Insecta" family="Cicadellidae" genus="Varicopsella" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="296" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="B918462EFFCC6F5EFF199073C3CDFBCA" box="[151,384,1087,1113]" italics="true" pageId="1" pageNumber="296">Varicopsella elegans</emphasis>
<bibRefCitation id="EFFDE7CDFFCC6F5EFE089073C01FFBCA" author="Viraktamath, C. A." box="[390,594,1087,1113]" pageId="1" pageNumber="296" pagination="295 - 310" refId="ref3494" refString="Viraktamath, C. A. (1981) Indian Macropsinae (Homoptera: Cicadellidae). II. Species described by W. L. Distant and descriptions of new species from the Indian subcontinent. Entomologica Scandinavica, 12, 295 - 310. https: // doi. org / 10.1163 / 187631281794709791" type="journal article" year="1981">Viraktamath 1981</bibRefCitation>
</taxonomicName>
;
<treatmentCitation id="0ACDBC2DFFCC6F5EFDD3900CC1AAFBCA" author="Yang, L. Y. &amp; Dietrich, C. H. &amp; Zhang, Y. L." box="[605,999,1087,1114]" page="193" pageId="1" pageNumber="296" year="2016">
<bibRefCitation id="EFFDE7CDFFCC6F5EFDD3900CC1AAFBCA" author="Yang, L. Y. &amp; Dietrich, C. H. &amp; Zhang, Y. L." box="[605,999,1087,1114]" pageId="1" pageNumber="296" pagination="187 - 194" refId="ref3611" refString="Yang, L. Y., Dietrich, C. H. &amp; Zhang, Y. L. (2016) Macropsini (Hemiptera: Cicadellidae) of Thailand, with description of two new species and three new country records. Zootaxa, 4168 (1), 187 - 194. https: // doi. org / 10.11646 / zootaxa. 4168.1.11" type="journal article" year="2016">Yang, Dietrich &amp; Zhang 2016: 193</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C376C9B7FFCC6F5EFF19902FC392FBEE" box="[151,479,1123,1149]" pageId="1" pageNumber="296" type="distribution">
<paragraph id="8BD39A3CFFCC6F5EFF19902FC392FBEE" blockId="1.[151,1003,1015,1149]" box="[151,479,1123,1149]" pageId="1" pageNumber="296">
Distribution.
<collectingCountry id="F37BDAACFFCC6F5EFEA3902FC324FBEE" box="[301,361,1123,1149]" name="India" pageId="1" pageNumber="296">India</collectingCountry>
,
<collectingCountry id="F37BDAACFFCC6F5EFEFA902FC396FBEE" box="[372,475,1123,1149]" name="Thailand" pageId="1" pageNumber="296">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,205 @@
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<mods:title id="BB461F375DC6A4C1F67A0C7254F09054">Taxonomy of the leafhopper genus Varicopsella (Hemiptera: Cicadellidae: Eurymelinae: Macropsini) with description of a new species from Thailand</mods:title>
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<mods:namePart id="C40F984C1583443B8C306F08A7013FCC">Zhang, Yalin</mods:namePart>
<mods:affiliation id="72506369BD79DD451DB59620C9FF51F5">Key Laboratory of Plant Protection Resources and Pest Management of the Ministry of Education, Entomological Museum, College of Plant Protection, Northwest A &amp; F University, Yangling, Shaanxi 712100, China</mods:affiliation>
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<subSubSection id="C376C9B7FFCF6F5DFF19908EC045FB4E" box="[151,520,1218,1245]" pageId="2" pageNumber="297" type="nomenclature">
<paragraph id="8BD39A3CFFCF6F5DFF19908EC045FB4E" blockId="2.[151,520,1218,1281]" box="[151,520,1218,1245]" pageId="2" pageNumber="297">
<heading id="D09B2D50FFCF6F5DFF19908EC045FB4E" bold="true" box="[151,520,1218,1245]" fontSize="11" level="1" pageId="2" pageNumber="297" reason="1">
<emphasis id="B918462EFFCF6F5DFF19908EC045FB4E" bold="true" box="[151,520,1218,1245]" pageId="2" pageNumber="297">
<taxonomicName id="4C6CE1BFFFCF6F5DFF19908EC3EEFB4F" authorityName="Wang &amp; Zhang" authorityYear="2024" box="[151,419,1218,1245]" class="Insecta" family="Cicadellidae" genus="Varicopsella" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="297" phylum="Arthropoda" rank="species" species="cornifera" status="sp. nov.">
<emphasis id="B918462EFFCF6F5DFF19908EC3EEFB4F" bold="true" box="[151,419,1218,1245]" italics="true" pageId="2" pageNumber="297">Varicopsella cornifera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22BFB55FFCF6F5DFE25908FC045FB4E" box="[427,520,1219,1245]" pageId="2" pageNumber="297" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C376C9B7FFCF6F5DFF1990AAC6CBF946" pageId="2" pageNumber="297" type="description">
<paragraph id="8BD39A3CFFCF6F5DFF1990AAC3C0FA92" blockId="2.[151,520,1218,1281]" box="[151,397,1254,1281]" pageId="2" pageNumber="297">
(
<figureCitation id="135786B9FFCF6F5DFF1190AAC37EFA92" box="[159,307,1254,1281]" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[185,1407,191,1909]" captionTargetId="figure-17@3.[175,1413,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. Varicopsella cornifera sp. nov.. A, E. habitus, dorsal view; B, F. habitus, lateral view; C, G. head and pronotum, dorsal view; D, H. face; AD, male; EH, female." pageId="2" pageNumber="297">Figs. 1. AH</figureCitation>
,
<figureCitation id="135786B9FFCF6F5DFEB090ABC3C8FA92" box="[318,389,1255,1281]" captionStart="FIGURE 2" captionStartId="4.[151,250,904,928]" captionTargetBox="[265,1314,181,877]" captionTargetId="figure-24@4.[265,1322,181,880]" captionTargetPageId="4" captionText="FIGURE 2. Varicopsella cornifera sp. nov. A, male genital capsule, lateral view; BC, aedeagus, ventral and lateral view; D, pygofer processes on ventral margin; EF, connective, lateral and ventral view; GH, dorsal connective, caudal and lateral view; I. style, lateral view." pageId="2" pageNumber="297">2. AI</figureCitation>
)
</paragraph>
<paragraph id="8BD39A3CFFCF6F5DFF199163C09CFADA" blockId="2.[151,1437,1326,2036]" box="[151,721,1326,1353]" pageId="2" pageNumber="297">
<emphasis id="B918462EFFCF6F5DFF199163C30EFADA" bold="true" box="[151,323,1327,1353]" pageId="2" pageNumber="297">Measurement.</emphasis>
Length.
<specimenCount id="9D6A51B5FFCF6F5DFE299163C3F1FADA" box="[423,444,1327,1353]" count="1" pageId="2" pageNumber="297" type="male"></specimenCount>
4.0
<quantity id="4C9437D9FFCF6F5DFE7B9163C001FADA" box="[501,588,1327,1353]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="2" pageNumber="297" unit="mm" value="4.1">4.1 mm</quantity>
,
<specimenCount id="9D6A51B5FFCF6F5DFDD99163C021FADA" box="[599,620,1327,1353]" count="1" pageId="2" pageNumber="297" type="female"></specimenCount>
<quantity id="4C9437D9FFCF6F5DFDFD9163C081FADA" box="[627,716,1327,1353]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="2" pageNumber="297" unit="mm" value="4.1">4.1 mm</quantity>
.
</paragraph>
<paragraph id="8BD39A3CFFCF6F5DFF49911EC1EAFA26" blockId="2.[151,1437,1326,2036]" pageId="2" pageNumber="297">
<emphasis id="B918462EFFCF6F5DFF49911EC305FAFF" bold="true" box="[199,328,1362,1388]" pageId="2" pageNumber="297">Coloration</emphasis>
. Crown, pronotum and scutellum yellowish, marked with brown spots and lines (
<figureCitation id="135786B9FFCF6F5DFB5E911EC718FAFE" box="[1232,1365,1362,1389]" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[185,1407,191,1909]" captionTargetId="figure-17@3.[175,1413,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. Varicopsella cornifera sp. nov.. A, E. habitus, dorsal view; B, F. habitus, lateral view; C, G. head and pronotum, dorsal view; D, H. face; AD, male; EH, female." pageId="2" pageNumber="297">Figs. 1C, G</figureCitation>
). Face pale yellowish; eyes grey; ocelli white (
<figureCitation id="135786B9FFCF6F5DFDD6913AC0ADFA02" box="[600,736,1398,1425]" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[185,1407,191,1909]" captionTargetId="figure-17@3.[175,1413,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. Varicopsella cornifera sp. nov.. A, E. habitus, dorsal view; B, F. habitus, lateral view; C, G. head and pronotum, dorsal view; D, H. face; AD, male; EH, female." pageId="2" pageNumber="297">Figs. 1D, H</figureCitation>
). Forewing subhyaline, with dense small irregular dark brown spots (
<figureCitation id="135786B9FFCF6F5DFF6E91D6C3EFFA26" box="[224,418,1434,1461]" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[185,1407,191,1909]" captionTargetId="figure-17@3.[175,1413,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. Varicopsella cornifera sp. nov.. A, E. habitus, dorsal view; B, F. habitus, lateral view; C, G. head and pronotum, dorsal view; D, H. face; AD, male; EH, female." pageId="2" pageNumber="297">Figs. 1A, B, E, F</figureCitation>
). Female colour similar to male (
<figureCitation id="135786B9FFCF6F5DFC9B91D6C1D7FA26" box="[789,922,1434,1461]" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[185,1407,191,1909]" captionTargetId="figure-17@3.[175,1413,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. Varicopsella cornifera sp. nov.. A, E. habitus, dorsal view; B, F. habitus, lateral view; C, G. head and pronotum, dorsal view; D, H. face; AD, male; EH, female." pageId="2" pageNumber="297">Figs. 1EH</figureCitation>
).
</paragraph>
<paragraph id="8BD39A3CFFCF6F5DFF4991F3C12DFA4B" blockId="2.[151,1437,1326,2036]" box="[199,864,1470,1497]" pageId="2" pageNumber="297">
<emphasis id="B918462EFFCF6F5DFF4991F3C316FA4A" bold="true" box="[199,347,1471,1497]" pageId="2" pageNumber="297">Morphology</emphasis>
. External characters as in generic description.
</paragraph>
<paragraph id="8BD39A3CFFCF6F5DFF4991AEC6CBF946" blockId="2.[151,1437,1326,2036]" pageId="2" pageNumber="297">
Male genitalia. Pygofer broad, with 2 spinelike distal processes on inner surface of lobe. Subgenital plate slender, with scattered setae, dorsal setae longest (
<figureCitation id="135786B9FFCF6F5DFDE0924AC085F9B2" box="[622,712,1542,1569]" captionStart="FIGURE 2" captionStartId="4.[151,250,904,928]" captionTargetBox="[265,1314,181,877]" captionTargetId="figure-24@4.[265,1322,181,880]" captionTargetPageId="4" captionText="FIGURE 2. Varicopsella cornifera sp. nov. A, male genital capsule, lateral view; BC, aedeagus, ventral and lateral view; D, pygofer processes on ventral margin; EF, connective, lateral and ventral view; GH, dorsal connective, caudal and lateral view; I. style, lateral view." pageId="2" pageNumber="297">Fig. 2A</figureCitation>
). Dorsal connectives divided into two sections, upper part broad basally and caudally directed ventrally, apex of ventral part acute in lateral view (
<figureCitation id="135786B9FFCF6F5DFBBD9266C6F0F9D6" box="[1075,1213,1578,1605]" captionStart="FIGURE 2" captionStartId="4.[151,250,904,928]" captionTargetBox="[265,1314,181,877]" captionTargetId="figure-24@4.[265,1322,181,880]" captionTargetPageId="4" captionText="FIGURE 2. Varicopsella cornifera sp. nov. A, male genital capsule, lateral view; BC, aedeagus, ventral and lateral view; D, pygofer processes on ventral margin; EF, connective, lateral and ventral view; GH, dorsal connective, caudal and lateral view; I. style, lateral view." pageId="2" pageNumber="297">Figs. 2G, H</figureCitation>
). Connective small, with fingerlike protrusion in middle between dorsally bent anterior arms (
<figureCitation id="135786B9FFCF6F5DFC6F9202C629F9FA" box="[993,1124,1614,1641]" captionStart="FIGURE 2" captionStartId="4.[151,250,904,928]" captionTargetBox="[265,1314,181,877]" captionTargetId="figure-24@4.[265,1322,181,880]" captionTargetPageId="4" captionText="FIGURE 2. Varicopsella cornifera sp. nov. A, male genital capsule, lateral view; BC, aedeagus, ventral and lateral view; D, pygofer processes on ventral margin; EF, connective, lateral and ventral view; GH, dorsal connective, caudal and lateral view; I. style, lateral view." pageId="2" pageNumber="297">Figs. 2E, F</figureCitation>
). Style slender, with sparse row of fine setae dorsolaterally, apex abruptly tapered and upturned (
<figureCitation id="135786B9FFCF6F5DFC10923EC1A0F91E" box="[926,1005,1650,1677]" captionStart="FIGURE 2" captionStartId="4.[151,250,904,928]" captionTargetBox="[265,1314,181,877]" captionTargetId="figure-24@4.[265,1322,181,880]" captionTargetPageId="4" captionText="FIGURE 2. Varicopsella cornifera sp. nov. A, male genital capsule, lateral view; BC, aedeagus, ventral and lateral view; D, pygofer processes on ventral margin; EF, connective, lateral and ventral view; GH, dorsal connective, caudal and lateral view; I. style, lateral view." pageId="2" pageNumber="297">Fig. 2I</figureCitation>
). Aedeagal shaft tubular, bent dorsally and evenly tapered to acute apex, with atrium broadened near base and shaft slightly tapered toward rounded apex in posterior view, dorsal apodeme and preatrium developed, gonopore apical (
<figureCitation id="135786B9FFCF6F5DFB8E92F6C634F946" box="[1024,1145,1722,1749]" captionStart="FIGURE 2" captionStartId="4.[151,250,904,928]" captionTargetBox="[265,1314,181,877]" captionTargetId="figure-24@4.[265,1322,181,880]" captionTargetPageId="4" captionText="FIGURE 2. Varicopsella cornifera sp. nov. A, male genital capsule, lateral view; BC, aedeagus, ventral and lateral view; D, pygofer processes on ventral margin; EF, connective, lateral and ventral view; GH, dorsal connective, caudal and lateral view; I. style, lateral view." pageId="2" pageNumber="297">Fig. 2B, C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C376C9B7FFCF6F5DFF499292C385F8F7" pageId="2" pageNumber="297" type="materials_examined">
<paragraph id="8BD39A3CFFCF6F5DFF499292C385F8F7" blockId="2.[151,1437,1326,2036]" pageId="2" pageNumber="297">
<emphasis id="B918462EFFCF6F5DFF499292C3E3F96B" bold="true" box="[199,430,1758,1784]" pageId="2" pageNumber="297">Material examined</emphasis>
.
<materialsCitation id="3B049061FFCF6F5DFE4F9292C618F88E" collectingDate="2008-07-08" collectingDateMax="2008-07-15" collectingDateMin="2008-07-08" collectionCode="QSBG" collectorName="Phu Toei NP &amp; Saunbua L." country="Thailand" elevation="763" latitude="14.973333" location="Suphanburi" longLatPrecision="92" longitude="99.43362" pageId="2" pageNumber="297" specimenCount="1" specimenCount-male="1" stateProvince="Suphan Buri" typeStatus="holotype">
<typeStatus id="54D7249EFFCF6F5DFE4F9292C064F96B" box="[449,553,1758,1784]" pageId="2" pageNumber="297" type="holotype">Holotype</typeStatus>
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(
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),
<collectingCountry id="F37BDAACFFCF6F5DFD489293C117F96A" box="[710,858,1759,1785]" name="Thailand" pageId="2" pageNumber="297">THAILAND</collectingCountry>
,
<collectingRegion id="49A854DEFFCF6F5DFCE49292C1A3F96B" box="[874,1006,1758,1784]" country="Thailand" name="Suphan Buri" pageId="2" pageNumber="297">Suphanburi</collectingRegion>
,
<collectorName id="2699FFEAFFCF6F5DFC709292C6D8F96A" box="[1022,1173,1758,1785]" pageId="2" pageNumber="297">Phu Toei NP</collectorName>
<taxonomicName id="4C6CE1BFFFCF6F5DFB2F9293C2ADF88E" authority="Jungh Forest" authorityName="Jungh Forest" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="2" pageNumber="297" phylum="Tracheophyta" rank="species" species="merkusii">
<emphasis id="B918462EFFCF6F5DFB2F9293C700F96B" box="[1185,1357,1758,1784]" italics="true" pageId="2" pageNumber="297">Pinus merkusii</emphasis>
Jungh Forest
</taxonomicName>
,
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,
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,
<quantity id="4C9437D9FFCF6F5DFD8B934EC00AF88F" box="[517,583,1794,1820]" metricMagnitude="2" metricUnit="m" metricValue="7.63" pageId="2" pageNumber="297" unit="m" value="763.0">
<elevation id="00417D0FFFCF6F5DFD8B934EC00AF88F" box="[517,583,1794,1820]" metricMagnitude="2" metricUnit="m" metricValue="7.63" pageId="2" pageNumber="297" unit="m" value="763.0">763m</elevation>
</quantity>
,
<collectingMethod id="522DE22BFFCF6F5DFDDD934EC0A4F88E" box="[595,745,1794,1821]" pageId="2" pageNumber="297">Malaise Trap</collectingMethod>
,
<date id="FFD2BCFCFFCF6F5DFD7A934EC1DFF88E" box="[756,914,1794,1821]" pageId="2" pageNumber="297" value="2008-07-08" valueMax="2008-07-15" valueMin="2008-07-08">
<collectingDate id="EF964514FFCF6F5DFD7A934EC1DFF88E" box="[756,914,1794,1821]" pageId="2" pageNumber="297" value="2008-07-08" valueMax="2008-07-15" valueMin="2008-07-08">815 vii 2008</collectingDate>
</date>
, coll.
<collectorName id="2699FFEAFFCF6F5DFC5B934EC618F88E" box="[981,1109,1794,1821]" pageId="2" pageNumber="297">Saunbua L.</collectorName>
</materialsCitation>
<materialsCitation id="3B049061FFCF6F5DFBD5934FC31EF8D3" collectingDate="2008-07-08" collectingDateMax="2008-07-15" collectingDateMin="2008-07-08" collectionCode="INHS" collectorName="Phu Toei NP &amp; Saunbua L." country="Thailand" elevation="763" latitude="14.973333" location="Suphanburi" longLatPrecision="92" longitude="99.43362" pageId="2" pageNumber="297" specimenCount="1" specimenCount-male="1" stateProvince="Suphan Buri" typeStatus="paratype">
<typeStatus id="54D7249EFFCF6F5DFBD5934FC681F88E" box="[1115,1228,1795,1821]" pageId="2" pageNumber="297" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D6A51B5FFCF6F5DFB59934FC6B7F88E" box="[1239,1274,1795,1821]" count="1" pageId="2" pageNumber="297" type="male">1♂</specimenCount>
(
<collectionCode id="ED7D02F9FFCF6F5DFA87934FC702F88E" box="[1289,1359,1795,1821]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34797" lsid="urn:lsid:biocol.org:col:34797" name="Illinois Natural History Survey" pageId="2" pageNumber="297" type="Other type of research institution/biorepository">INHS</collectionCode>
), same data as holotype
</materialsCitation>
;
<materialsCitation id="3B049061FFCF6F5DFED0936BC385F8F7" collectingDate="2007-04-12" collectingDateMax="2007-04-19" collectingDateMin="2007-04-12" collectionCode="INHS" collectorName="Tat Tone NP &amp; Tawit Jaruphan." country="Thailand" elevation="271" latitude="15.954" location="Chaiyaphum" longLatPrecision="13" longitude="101.90134" pageId="2" pageNumber="297" specimenCount="1" specimenCount-female="1" stateProvince="Chaiyaphum" typeStatus="paratype">
<specimenCount id="9D6A51B5FFCF6F5DFED0936BC3CCF8D2" box="[350,385,1831,1857]" count="1" pageId="2" pageNumber="297" type="female">1♀</specimenCount>
(
<collectionCode id="ED7D02F9FFCF6F5DFE1F936BC39AF8D2" box="[401,471,1831,1857]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34797" lsid="urn:lsid:biocol.org:col:34797" name="Illinois Natural History Survey" pageId="2" pageNumber="297" type="Other type of research institution/biorepository">INHS</collectionCode>
),
<collectingCountry id="F37BDAACFFCF6F5DFE65936BC032F8D2" box="[491,639,1831,1857]" name="Thailand" pageId="2" pageNumber="297">THAILAND</collectingCountry>
,
<collectingRegion id="49A854DEFFCF6F5DFD04936AC151F8D3" box="[650,796,1830,1856]" country="Thailand" name="Chaiyaphum" pageId="2" pageNumber="297">Chaiyaphum</collectingRegion>
,
<collectorName id="2699FFEAFFCF6F5DFCA6936BC1F8F8D2" box="[808,949,1831,1857]" pageId="2" pageNumber="297">Tat Tone NP</collectorName>
,
<geoCoordinate id="EE58FCFBFFCF6F5DFC4F9364C609F8D0" box="[961,1092,1832,1859]" degrees="15" direction="north" minutes="57.24" orientation="latitude" pageId="2" pageNumber="297" precision="9" value="15.954">15°57.24N</geoCoordinate>
,
<geoCoordinate id="EE58FCFBFFCF6F5DFBDE9364C693F8D0" box="[1104,1246,1832,1859]" degrees="101" direction="east" minutes="54.08" orientation="longitude" pageId="2" pageNumber="297" precision="9" value="101.90134">101°54.08E</geoCoordinate>
,
<quantity id="4C9437D9FFCF6F5DFB64936BC760F8D2" box="[1258,1325,1831,1857]" metricMagnitude="2" metricUnit="m" metricValue="2.71" pageId="2" pageNumber="297" unit="m" value="271.0">
<elevation id="00417D0FFFCF6F5DFB64936BC760F8D2" box="[1258,1325,1831,1857]" metricMagnitude="2" metricUnit="m" metricValue="2.71" pageId="2" pageNumber="297" unit="m" value="271.0">271m</elevation>
</quantity>
,
<date id="FFD2BCFCFFCF6F5DFAB7936BC29FF8F6" pageId="2" pageNumber="297" value="2007-04-12" valueMax="2007-04-19" valueMin="2007-04-12">
<collectingDate id="EF964514FFCF6F5DFAB7936BC29FF8F6" pageId="2" pageNumber="297" value="2007-04-12" valueMax="2007-04-19" valueMin="2007-04-12">1219 iv 2007</collectingDate>
</date>
, coll.
<collectorName id="2699FFEAFFCF6F5DFE9B9306C385F8F7" box="[277,456,1866,1892]" pageId="2" pageNumber="297">Tawit Jaruphan.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C376C9B7FFCF6F5DFF499322C6F4F83E" pageId="2" pageNumber="297" type="diagnosis">
<paragraph id="8BD39A3CFFCF6F5DFF499322C6F4F83E" blockId="2.[151,1437,1326,2036]" pageId="2" pageNumber="297">
<emphasis id="B918462EFFCF6F5DFF499322C377F81B" bold="true" box="[199,314,1902,1928]" pageId="2" pageNumber="297">Diagnosis</emphasis>
. This new species resembles
<taxonomicName id="4C6CE1BFFFCF6F5DFD179323C1EEF81B" authority="Viraktamath" authorityName="Viraktamath" authorityYear="1981" box="[665,931,1902,1928]" class="Insecta" family="Cicadellidae" genus="Varicopsella" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="297" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="B918462EFFCF6F5DFD179323C142F81B" box="[665,783,1902,1928]" italics="true" pageId="2" pageNumber="297">V. elegans</emphasis>
Viraktamath
</taxonomicName>
in the structure of the pygofer, but can be distinguished from the latter by the dorsal connective upper part with dorsal preapical process.
</paragraph>
</subSubSection>
<subSubSection id="C376C9B7FFCF6F5DFF4993FBC355F867" pageId="2" pageNumber="297" type="etymology">
<paragraph id="8BD39A3CFFCF6F5DFF4993FBC355F867" blockId="2.[151,1437,1326,2036]" pageId="2" pageNumber="297">
<emphasis id="B918462EFFCF6F5DFF4993FBC305F842" bold="true" box="[199,328,1975,2001]" pageId="2" pageNumber="297">Etymology</emphasis>
. This specific epithet is from the Latin word “
<emphasis id="B918462EFFCF6F5DFCDC93FBC1E6F843" box="[850,939,1975,2000]" italics="true" pageId="2" pageNumber="297">corneus</emphasis>
” referring to the hornlike apex of the dorsal connective.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,109 @@
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<mods:title id="D922444536E8DAF42F59217B8D4BDE0E">Taxonomy of the leafhopper genus Varicopsella (Hemiptera: Cicadellidae: Eurymelinae: Macropsini) with description of a new species from Thailand</mods:title>
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<paragraph id="8BD39A3CFFCF6F5DFF1996FDC3E7FD58" blockId="2.[151,426,689,716]" box="[151,426,689,716]" pageId="2" pageNumber="297">
<heading id="D09B2D50FFCF6F5DFF1996FDC3E7FD58" bold="true" box="[151,426,689,716]" fontSize="11" level="1" pageId="2" pageNumber="297" reason="1">
<taxonomicName id="4C6CE1BFFFCF6F5DFF1996FDC3E7FD58" ID-CoL="8HSP3" authority="Hamilton" authorityName="Hamilton" authorityYear="1980" box="[151,426,689,716]" class="Insecta" family="Cicadellidae" genus="Varicopsella" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="297" phylum="Arthropoda" rank="genus">
<emphasis id="B918462EFFCF6F5DFF1996FDC3E7FD58" bold="true" box="[151,426,689,716]" pageId="2" pageNumber="297">
<emphasis id="B918462EFFCF6F5DFF1996FDC366FD5F" bold="true" box="[151,299,689,716]" italics="true" pageId="2" pageNumber="297">Varicopsella</emphasis>
Hamilton
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C376C9B7FFCF6F5DFF1996BAC3BBFC9D" box="[151,502,758,782]" pageId="2" pageNumber="297" type="reference_group">
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<treatmentCitationGroup id="AB7CBD12FFCF6F5DFF1996BAC3BBFC9D" box="[151,502,758,782]" pageId="2" pageNumber="297">
<treatmentCitation id="0ACDBC2DFFCF6F5DFF1996BAC3BFFC9D" author="Hamilton, K. G. A." box="[151,498,758,782]" page="900" pageId="2" pageNumber="297" year="1980">
<taxonomicName id="4C6CE1BFFFCF6F5DFF1996BAC3BFFC9D" ID-CoL="8HSP3" authority="Hamilton, 1980: 900" authorityName="Hamilton" authorityPageNumber="900" authorityYear="1980" box="[151,498,758,782]" class="Insecta" family="Cicadellidae" genus="Varicopsella" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="297" phylum="Arthropoda" rank="genus">
<emphasis id="B918462EFFCF6F5DFF1996BAC35BFC9D" box="[151,278,758,782]" italics="true" pageId="2" pageNumber="297">Varicopsella</emphasis>
<bibRefCitation id="EFFDE7CDFFCF6F5DFE9296BAC3BFFC9D" author="Hamilton, K. G. A." box="[284,498,758,782]" pageId="2" pageNumber="297" pagination="875 - 932" refId="ref3094" refString="Hamilton, K. G. A. (1980) Contributions to the study of the world Macropsini (Rhynchota: Homoptera: Cicadellidae). The Canadian Entomologist, 112 (9), 875 - 932. https: // doi. org / 10.4039 / Ent 112875 - 9" type="journal article" year="1980">Hamilton, 1980: 900</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
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</paragraph>
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<paragraph id="8BD39A3CFFCF6F5DFF199759C033FCBF" blockId="2.[151,638,758,813]" box="[151,638,788,813]" pageId="2" pageNumber="297">
<typeStatus id="54D7249EFFCF6F5DFF199759C284FCBE" box="[151,201,789,813]" pageId="2" pageNumber="297">Type</typeStatus>
species:
<taxonomicName id="4C6CE1BFFFCF6F5DFEA89759C033FCBF" authority="Merino, 1936" authorityName="Merino" authorityYear="1936" box="[294,638,788,812]" class="Insecta" family="Cicadellidae" genus="Macropsis" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="297" phylum="Arthropoda" rank="species" species="breakeyi">
<emphasis id="B918462EFFCF6F5DFEA89759C3A0FCBF" box="[294,493,788,812]" italics="true" pageId="2" pageNumber="297">Macropsis breakeyi</emphasis>
<bibRefCitation id="EFFDE7CDFFCF6F5DFE7D9758C033FCBF" author="Merino, G." box="[499,638,788,812]" pageId="2" pageNumber="297" pagination="318 - 325" refId="ref3300" refString="Merino, G. (1936) Philippine Cicadellidae. The Philippine Journal of Science, 61 (3), 318 - 325." type="journal article" year="1936">Merino, 1936</bibRefCitation>
</taxonomicName>
</paragraph>
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<subSubSection id="C376C9B7FFCF6F5DFF199716C3EDFBDF" pageId="2" pageNumber="297" type="diagnosis">
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<emphasis id="B918462EFFCF6F5DFF199716C35FFCE7" bold="true" box="[151,274,858,884]" pageId="2" pageNumber="297">Diagnosis.</emphasis>
Body obviously depressed. Crown prominent, V-shaped in dorsal view. Face with lora fused to frontoclypeus, lacking clear sutures between them. Ocelli on face, closer to adjacent eyes than to each other. Pronotum slightly declivous, striations on surface dense and oblique. Scutellum triangular. Forewings with 3 anteapical cells, veins protruding. Male pygofer broad, with some short spinelike distal processes on inner surface of lobe. Subgenital plate slender, with scattered setae. Dorsal connective divided into two articulated portions. Connective small, posterior margin nearly truncate. Style slender, with sparse row of fine setae dorsolaterally. Aedeagus shaft tubular.
</paragraph>
</subSubSection>
<subSubSection id="C376C9B7FFCF6F5DFF49901AC1B9FBE3" box="[199,1012,1110,1137]" pageId="2" pageNumber="297" type="distribution">
<paragraph id="8BD39A3CFFCF6F5DFF49901AC1B9FBE3" blockId="2.[151,1437,858,1137]" box="[199,1012,1110,1137]" pageId="2" pageNumber="297">
<emphasis id="B918462EFFCF6F5DFF49901AC32CFBE3" bold="true" box="[199,353,1110,1136]" pageId="2" pageNumber="297">Distribution.</emphasis>
<collectingCountry id="F37BDAACFFCF6F5DFEE9901AC39CFBE3" box="[359,465,1110,1136]" name="Australia" pageId="2" pageNumber="297">Australia</collectingCountry>
, Borneo,
<collectingCountry id="F37BDAACFFCF6F5DFDB3901AC0CEFBE3" box="[573,643,1110,1136]" name="China" pageId="2" pageNumber="297">China</collectingCountry>
,
<collectingCountry id="F37BDAACFFCF6F5DFD03901AC084FBE3" box="[653,713,1110,1136]" name="India" pageId="2" pageNumber="297">India</collectingCountry>
,
<collectingCountry id="F37BDAACFFCF6F5DFD5B901AC11EFBE3" box="[725,851,1110,1136]" name="Philippines" pageId="2" pageNumber="297">Philippines</collectingCountry>
and
<collectingCountry id="F37BDAACFFCF6F5DFC07901AC1BDFBE3" box="[905,1008,1110,1136]" name="Thailand" pageId="2" pageNumber="297">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,312 @@
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<paragraph id="8BCA36ECFFB1FF84FF17FF16D318FF7C" blockId="1.[151,965,223,322]" box="[151,780,223,250]" pageId="1" pageNumber="146">
<heading id="D0828180FFB1FF84FF17FF16D318FF7C" bold="true" box="[151,780,223,250]" fontSize="11" level="1" pageId="1" pageNumber="146" reason="1">
<emphasis id="B901EAFEFFB1FF84FF17FF16D318FF7C" bold="true" box="[151,780,223,250]" pageId="1" pageNumber="146">
<taxonomicName id="4C754D6FFFB1FF84FF17FF16D2D6FF7C" authority="Borovec and Anderson" authorityName="Borovec and Anderson" authorityYear="2024" box="[151,706,223,250]" class="Insecta" family="Curculionidae" genus="Laohajekia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="146" phylum="Arthropoda" rank="species" species="uniformis" status="sp. nov.">
<emphasis id="B901EAFEFFB1FF84FF17FF16D18AFF7C" bold="true" box="[151,414,223,250]" italics="true" pageId="1" pageNumber="146">Laohajekia uniformis</emphasis>
Borovec and Anderson
</taxonomicName>
<taxonomicNameLabel id="A2325785FFB1FF84FD4AFF16D318FF7C" box="[714,780,223,249]" pageId="1" pageNumber="146" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="8BCA36ECFFB1FF84FF17FEEED157FEC7" blockId="1.[151,965,223,322]" box="[151,323,295,322]" pageId="1" pageNumber="146">
(
<figureCitation id="134E2A69FFB1FF84FF1FFEEED110FEC4" box="[159,260,295,321]" captionStart="FIGURES 14" captionStartId="1.[152,270,1594,1618]" captionTargetBox="[154,1436,771,1548]" captionTargetId="figure-218@1.[151,1436,771,1568]" captionTargetPageId="1" captionText="FIGURES 14. Dorsal and lateral habitus. 12. Laohajekia uniformis. 34. Trachyphloeosoma newthayorum." figureDoi="http://doi.org/10.5281/zenodo.13746095" httpUri="https://zenodo.org/record/13746095/files/figure.png" pageId="1" pageNumber="146">Figs 1, 2</figureCitation>
,
<figureCitation id="134E2A69FFB1FF84FE90FEEED12FFEC7" box="[272,315,295,322]" captionStart="FIGURES 513" captionStartId="2.[152,270,1652,1676]" captionTargetBox="[176,1410,392,1615]" captionTargetId="figure-108@2.[164,1423,381,1628]" captionTargetPageId="2" captionText="FIGURES 513. Structural details. 58. Laohajekia uniformis. 5. Head and rostrum, dorsal view. 6. Head and rostrum, lateral view. 7. Penis, dorsal and lateral view. 8. Spermatheca. 913. Trachyphloeosoma newthayorum. 9. Head and rostrum, dorsal view. 10. Head and rostrum, lateral view. 11. Penis, dorsal and lateral view. 12. Spermatheca. 13. Sternite VIII in female. Scale bars: 0.50 mm (5, 6, 9, 10, 13), 0.25 mm (7, 11) and 0.125 mm (8, 12)." figureDoi="http://doi.org/10.5281/zenodo.13746097" httpUri="https://zenodo.org/record/13746097/files/figure.png" pageId="1" pageNumber="146">58</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36F6567FFB1FF86FF17FEA2D504FD00" lastPageId="3" lastPageNumber="148" pageId="1" pageNumber="146" type="materials_examined">
<paragraph id="8BCA36ECFFB1FF84FF17FEA2D59DFE48" blockId="1.[151,1437,363,750]" pageId="1" pageNumber="146">
<emphasis id="B901EAFEFFB1FF84FF17FEA2D151FE00" bold="true" box="[151,325,363,389]" pageId="1" pageNumber="146">Type material.</emphasis>
<materialsCitation id="3B1D3CB1FFB1FF84FECCFEA2D37FFE48" collectingDate="2008-05-28" collectionCode="FMNH" collectorName="A. Newton &amp; M. Thayer" country="Laos" elevation="820" latitude="18.352467" location="Phou Khao Khouay" longLatPrecision="1" longitude="102.81115" pageId="1" pageNumber="146" specimenCount="1" specimenCount-male="1" stateProvince="Viangchan" typeStatus="holotype">
<typeStatus id="54CE884EFFB1FF84FECCFEA2D1ADFE00" box="[332,441,363,389]" pageId="1" pageNumber="146" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D73FD65FFB1FF84FE43FEA5D1CEFE03" box="[451,474,364,390]" count="1" pageId="1" pageNumber="146" type="male"></specimenCount>
,
<collectingCountry id="F362767CFFB1FF84FE66FEA5D20BFE03" box="[486,543,364,390]" name="Laos" pageId="1" pageNumber="146">Laos</collectingCountry>
,
<collectingRegion id="49B1F80EFFB1FF84FDAAFEA2D316FE00" box="[554,770,363,389]" country="Laos" name="Viangchan" pageId="1" pageNumber="146">Vientiane Province</collectingRegion>
,
<location id="8EAA6037FFB1FF84FC8DFEA2D3FEFE00" LSID="urn:lsid:plazi:treatment:03DC87FAFFB1FF86FF17FF16D488F9D4:8EAA6037FFB1FF84FC8DFEA2D3FEFE00" box="[781,1002,363,389]" country="Laos" latitude="18.352467" longLatPrecision="1" longitude="102.81115" name="Phou Khao Khouay" pageId="1" pageNumber="146" stateProvince="Viangchan">Phou Khao Khouay</location>
NBCA,
<locationDeviation id="9A782FCEFFB1FF84FBCDFEA5D53CFE00" box="[1101,1320,363,390]" location="Ban Vangheua" pageId="1" pageNumber="146">N of Ban Vangheua</locationDeviation>
(= Khua),
<quantity id="4C8D9B09FFB1FF84FF17FE46D0F4FE2F" box="[151,224,399,426]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="1" pageNumber="146" unit="m" value="820.0">
<elevation id="0058D1DFFFB1FF84FF17FE46D0F4FE2F" box="[151,224,399,426]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="1" pageNumber="146" unit="m" value="820.0">820 m</elevation>
</quantity>
,
<geoCoordinate id="EE41502BFFB1FF84FF6CFE46D16AFE2F" box="[236,382,399,426]" degrees="18" direction="north" minutes="21.148" orientation="latitude" pageId="1" pageNumber="146" precision="1" value="18.352467">18°21.148 N</geoCoordinate>
,
<geoCoordinate id="EE41502BFFB1FF84FE09FE46D231FE2F" box="[393,549,399,426]" degrees="102" direction="east" minutes="48.669" orientation="longitude" pageId="1" pageNumber="146" precision="1" value="102.81115">102°48.669 E</geoCoordinate>
,
<date id="FFCB102CFFB1FF84FDB1FE46D2B6FE2C" box="[561,674,399,425]" pageId="1" pageNumber="146" value="2008-05-28">
<collectingDate id="EF8FE9C4FFB1FF84FDB1FE46D2B6FE2C" box="[561,674,399,425]" pageId="1" pageNumber="146" value="2008-05-28">28.v.2008</collectingDate>
</date>
, old-growth tropical hardwood forest, FMHD2008-020; berlese, log &amp; leaf litter,
<collectorName id="2680533AFFB1FF84FEA5FE7AD1B4FE4B" box="[293,416,435,462]" pageId="1" pageNumber="146">A. Newton</collectorName>
&amp;
<collectorName id="2680533AFFB1FF84FE44FE7AD221FE48" box="[452,565,435,461]" pageId="1" pageNumber="146">M.Thayer</collectorName>
, ANMT site 1222 (
<collectionCode id="ED64AE29FFB1FF84FC8CFE7AD377FE48" box="[780,867,435,461]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="1" pageNumber="146" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B1D3CB1FFB1FF84FCF6FE7DD591FE48" box="[886,1413,435,462]" collectingDate="2008-05-28" collectionCode="CMNC" collectorName="A. Newton &amp; M. Thayer" country="Laos" elevation="820" latitude="18.352467" location="Phou Khao Khouay" longLatPrecision="1" longitude="102.81115" pageId="1" pageNumber="146" specimenCount="1" specimenCount-female="1" stateProvince="Viangchan" typeStatus="paratype">
<typeStatus id="54CE884EFFB1FF84FCF6FE7DD3C8FE4B" box="[886,988,436,462]" pageId="1" pageNumber="146" type="paratype">Paratype</typeStatus>
:
<specimenCount id="9D73FD65FFB1FF84FC66FE7AD404FE4B" box="[998,1040,435,462]" count="1" pageId="1" pageNumber="146" type="female">1 ♀</specimenCount>
, same data as holotype (
<collectionCode id="ED64AE29FFB1FF84FAA4FE7AD569FE48" box="[1316,1405,435,461]" pageId="1" pageNumber="146">CMNC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BCA36ECFFB1FF84FF17FE1ED343FD6B" blockId="1.[151,1437,363,750]" pageId="1" pageNumber="146">
Body length:
<typeStatus id="54CE884EFFB1FF84FEB0FE1ED186FE74" box="[304,402,471,497]" pageId="1" pageNumber="146" type="holotype">holotype</typeStatus>
<quantity id="4C8D9B09FFB1FF84FE19FE1ED1EBFE77" box="[409,511,471,498]" metricMagnitude="-3" metricUnit="m" metricValue="2.06" pageId="1" pageNumber="146" unit="mm" value="2.06">2.06 mm</quantity>
,
<typeStatus id="54CE884EFFB1FF84FD8AFE11D27EFE77" box="[522,618,472,498]" pageId="1" pageNumber="146" type="paratype">paratype</typeStatus>
<quantity id="4C8D9B09FFB1FF84FDF1FE1ED2CCFE77" box="[625,728,471,498]" metricMagnitude="-3" metricUnit="m" metricValue="2.13" pageId="1" pageNumber="146" unit="mm" value="2.13">2.13 mm</quantity>
. Body (
<figureCitation id="134E2A69FFB1FF84FCAEFE1ED386FE74" box="[814,914,471,497]" captionStart="FIGURES 14" captionStartId="1.[152,270,1594,1618]" captionTargetBox="[154,1436,771,1548]" captionTargetId="figure-218@1.[151,1436,771,1568]" captionTargetPageId="1" captionText="FIGURES 14. Dorsal and lateral habitus. 12. Laohajekia uniformis. 34. Trachyphloeosoma newthayorum." figureDoi="http://doi.org/10.5281/zenodo.13746095" httpUri="https://zenodo.org/record/13746095/files/figure.png" pageId="1" pageNumber="146">Figs 1, 2</figureCitation>
) dark brownish, basal parts of scapes, funicles with clubs and tarsi slightly paler, reddish brown.Appressed scales on dorsal part of body star-shaped, not completely covering integument, 23 across width of one interstria. Elytra with regular row of erect, narrow, long, sparse setae, slightly longer at posterior declivity than on the disc; setae apically rounded, widest at middle, at posterior declivity almost as long as the width of each interstria, distance between two setae 23 longer than length of one seta.Anterior margin of pronotum, head with rostrum with similar erect setae, slightly shorter than those on elytra. Antennal scapes, femora and tibiae with same appressed scales and erect setae as body, funicles with tarsi with semi-erect piliform setae, clubs densely setose with short appressed setae.
</paragraph>
<caption id="DF0A6664FFB1FF84FF18F9F3D4EEF9D7" ID-DOI="http://doi.org/10.5281/zenodo.13746095" ID-Zenodo-Dep="13746095" box="[152,1274,1594,1618]" httpUri="https://zenodo.org/record/13746095/files/figure.png" pageId="1" pageNumber="146" startId="1.[152,270,1594,1618]" targetBox="[154,1436,771,1548]" targetPageId="1" targetType="figure">
<paragraph id="8BCA36ECFFB1FF84FF18F9F3D4EEF9D7" blockId="1.[152,1274,1594,1618]" box="[152,1274,1594,1618]" pageId="1" pageNumber="146">
<emphasis id="B901EAFEFFB1FF84FF18F9F3D155F9D7" bold="true" box="[152,321,1594,1618]" pageId="1" pageNumber="146">FIGURES 14.</emphasis>
Dorsal and lateral habitus. 12.
<taxonomicName id="4C754D6FFFB1FF84FD0AF9F3D377F9D7" authorityName="Borovec and Anderson" authorityYear="2024" box="[650,867,1594,1618]" class="Insecta" family="Curculionidae" genus="Laohajekia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="146" phylum="Arthropoda" rank="species" species="uniformis">
<emphasis id="B901EAFEFFB1FF84FD0AF9F3D377F9D7" box="[650,867,1594,1618]" italics="true" pageId="1" pageNumber="146">Laohajekia uniformis</emphasis>
</taxonomicName>
. 34.
<taxonomicName id="4C754D6FFFB1FF84FC23F9F3D4E0F9D7" authorityName="Borovec and Anderson" authorityYear="2024" box="[931,1268,1594,1618]" class="Insecta" family="Curculionidae" genus="Trachyphloeosoma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="146" phylum="Arthropoda" rank="species" species="newthayorum">
<emphasis id="B901EAFEFFB1FF84FC23F9F3D4E0F9D7" box="[931,1268,1594,1618]" italics="true" pageId="1" pageNumber="146">Trachyphloeosoma newthayorum</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BCA36ECFFB1FF84FF47F954D529F80B" blockId="1.[151,1437,1692,2043]" pageId="1" pageNumber="146">
Rostrum (
<figureCitation id="134E2A69FFB1FF84FEB6F955D183F933" box="[310,407,1692,1719]" captionStart="FIGURES 513" captionStartId="2.[152,270,1652,1676]" captionTargetBox="[176,1410,392,1615]" captionTargetId="figure-108@2.[164,1423,381,1628]" captionTargetPageId="2" captionText="FIGURES 513. Structural details. 58. Laohajekia uniformis. 5. Head and rostrum, dorsal view. 6. Head and rostrum, lateral view. 7. Penis, dorsal and lateral view. 8. Spermatheca. 913. Trachyphloeosoma newthayorum. 9. Head and rostrum, dorsal view. 10. Head and rostrum, lateral view. 11. Penis, dorsal and lateral view. 12. Spermatheca. 13. Sternite VIII in female. Scale bars: 0.50 mm (5, 6, 9, 10, 13), 0.25 mm (7, 11) and 0.125 mm (8, 12)." figureDoi="http://doi.org/10.5281/zenodo.13746097" httpUri="https://zenodo.org/record/13746097/files/figure.png" pageId="1" pageNumber="146">Figs 5, 6</figureCitation>
) short and wide, 1.171.19× as wide as long, widest shortly before base, then slightly evenly tapering apicad with straight sides, apically broadly rounded; in lateral view regularly vaulted. Epifrons wide and flat, widest at base, distinctly evenly tapering apicad with straight sides, at base 1.361.41× as wide as at apex. Frons densely squamosa. Epistome small, U-shaped, glabrous, posteriorly faintly carinated. Antennal scrobes dorsally visible in anterior part of rostrum as narrowly reniform; in lateral view narrow, furrow-shaped, short, slightly curved, directed towards eye, separated from it by wide squamosa stripe. Eyes small, in dorsal view slightly prominent from outline of head; in lateral view distant from dorsal margin of head equal to eye diameter. Vertex flat, wide.
</paragraph>
<paragraph id="8BCA36ECFFB1FF87FF47F850D1ECFF50" blockId="1.[151,1437,1692,2043]" lastBlockId="2.[151,1437,150,357]" lastPageId="2" lastPageNumber="147" pageId="1" pageNumber="146">Antennae somewhat slender; scapes straight, evenly slightly enlarging apicad, 4.674.73× as long as at apex wide, at apex 0.8× as wide as club. Funicle of 7 antennomeres. Antennomeres 1 and 2 subequal in length, conical; antennomere 1 1.51.6× as long as wide; antennomere 2 1.71.8× as long as wide; antennomere 3 1.1× as wide as long; antennomere 4 isodiametric; antennomeres 5 and 6 1.1× as wide as long; antennomere 7 1.2× as wide as long; clubs 1.71.8× as long as wide.</paragraph>
<paragraph id="8BCA36ECFFB2FF87FF47FF16D487FEE0" blockId="2.[151,1437,150,357]" pageId="2" pageNumber="147">
Pronotum (
<figureCitation id="134E2A69FFB2FF87FEC5FF16D1BDFF7C" box="[325,425,223,249]" captionStart="FIGURES 14" captionStartId="1.[152,270,1594,1618]" captionTargetBox="[154,1436,771,1548]" captionTargetId="figure-218@1.[151,1436,771,1568]" captionTargetPageId="1" captionText="FIGURES 14. Dorsal and lateral habitus. 12. Laohajekia uniformis. 34. Trachyphloeosoma newthayorum." figureDoi="http://doi.org/10.5281/zenodo.13746095" httpUri="https://zenodo.org/record/13746095/files/figure.png" pageId="2" pageNumber="147">Figs 1, 2</figureCitation>
) slender, 1.421.47× as wide as long, widest at midlength with slightly regularly rounded sides, equally narrowing anteriad and posteriad with anterior margin indistinctly narrower than posterior one. Disc weakly regularly domed, in lateral view regularly convex. Integument under appressed scales roughly densely punctured, punctures identical to those in elytral rows, distance between them very narrow.
</paragraph>
<caption id="DF0A6664FFB2FF87FF18F9BDD370F97D" ID-DOI="http://doi.org/10.5281/zenodo.13746097" ID-Zenodo-Dep="13746097" httpUri="https://zenodo.org/record/13746097/files/figure.png" pageId="2" pageNumber="147" startId="2.[152,270,1652,1676]" targetBox="[176,1410,392,1615]" targetPageId="2" targetType="figure">
<paragraph id="8BCA36ECFFB2FF87FF18F9BDD370F97D" blockId="2.[151,1436,1652,1784]" pageId="2" pageNumber="147">
<emphasis id="B901EAFEFFB2FF87FF18F9BDD15AF909" bold="true" box="[152,334,1652,1676]" pageId="2" pageNumber="147">FIGURES 513.</emphasis>
Structural details. 58.
<taxonomicName id="4C754D6FFFB2FF87FDBFF9BDD30DF909" authorityName="Borovec and Anderson" authorityYear="2024" box="[575,793,1652,1676]" class="Insecta" family="Curculionidae" genus="Laohajekia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="147" phylum="Arthropoda" rank="species" species="uniformis">
<emphasis id="B901EAFEFFB2FF87FDBFF9BDD30DF909" box="[575,793,1652,1676]" italics="true" pageId="2" pageNumber="147">Laohajekia uniformis</emphasis>
</taxonomicName>
. 5. Head and rostrum, dorsal view. 6. Head and rostrum, lateral view. 7. Penis, dorsal and lateral view. 8. Spermatheca. 913.
<taxonomicName id="4C754D6FFFB2FF87FC9BF951D47AF935" authorityName="Borovec and Anderson" authorityYear="2024" box="[795,1134,1688,1712]" class="Insecta" family="Curculionidae" genus="Trachyphloeosoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="147" phylum="Arthropoda" rank="species" species="newthayorum">
<emphasis id="B901EAFEFFB2FF87FC9BF951D47AF935" box="[795,1134,1688,1712]" italics="true" pageId="2" pageNumber="147">Trachyphloeosoma newthayorum</emphasis>
</taxonomicName>
. 9. Head and rostrum, dorsal view. 10. Head and rostrum, lateral view. 11. Penis, dorsal and lateral view. 12. Spermatheca. 13. Sternite VIII in female. Scale bars: 0.50 mm (5, 6, 9, 10, 13), 0.25 mm (7, 11) and 0.125 mm (8, 12).
</paragraph>
</caption>
<paragraph id="8BCA36ECFFB2FF87FF47F883D1A9F8E1" blockId="2.[151,1436,1866,2036]" box="[199,445,1866,1893]" pageId="2" pageNumber="147">Scutellum not visible.</paragraph>
<paragraph id="8BCA36ECFFB2FF87FF47F8A7D11BF871" blockId="2.[151,1436,1866,2036]" pageId="2" pageNumber="147">
Elytra (
<figureCitation id="134E2A69FFB2FF87FE9BF8A7D194F80D" box="[283,384,1902,1928]" captionStart="FIGURES 14" captionStartId="1.[152,270,1594,1618]" captionTargetBox="[154,1436,771,1548]" captionTargetId="figure-218@1.[151,1436,771,1568]" captionTargetPageId="1" captionText="FIGURES 14. Dorsal and lateral habitus. 12. Laohajekia uniformis. 34. Trachyphloeosoma newthayorum." figureDoi="http://doi.org/10.5281/zenodo.13746095" httpUri="https://zenodo.org/record/13746095/files/figure.png" pageId="2" pageNumber="147">Figs 1, 2</figureCitation>
) in male subglobular, 1.21× as long as wide, in female oval, 1.28× as long as wide, widest at midlength, with distinctly regularly rounded sides, apically broadly rounded; in lateral view distinctly vaulted. Base equally wide as base of pronotum. Striae distinctly roughly punctured, about twice as wide as narrow, equally flat interstriae.
</paragraph>
<paragraph id="8BCA36ECFFB3FF86FF47FF5ED4F3FE98" blockId="3.[151,1437,151,1617]" pageId="3" pageNumber="148">All femora edentate. Tibiae robust, mucronate, not denticulate. Protibiae 4.274.31× as long as at midlength wide, with lateral margin straight and mesal margin bisinuate, apically obliquely subtruncated, fringed with short yellowish setae. Tarsi short; tarsomere 2 1.11.2× as wide as long; tarsomere 3 1.31.4× as wide as long and 1.3 1.4× as wide as tarsomere 2; onychium 0.91.1× as long as tarsomere 3. Claws fused in basal part.</paragraph>
<paragraph id="8BCA36ECFFB3FF86FF47FEEED273FE28" blockId="3.[151,1437,151,1617]" pageId="3" pageNumber="148">
Abdominal ventrites 1.031.07× as long as wide, densely squamose with appressed rounded scales, with metasternal process obtuse, wider than transverse diameter of metacoxa. Ventrite 1 large, in middle twice as long as ventrite 2; ventrite 2 slightly longer than ventrite 3 or 4; ventrite
<quantity id="4C8D9B09FFB3FF86FCE2FEA7D398FE0C" box="[866,908,366,393]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="3" pageNumber="148" unit="in" value="5.0">5 in</quantity>
male subtrapezoidal, apically obtuse, in female subtriangular, apically narrowly tapering.
</paragraph>
<paragraph id="8BCA36ECFFB3FF86FF47FE7ED57AFDB8" blockId="3.[151,1437,151,1617]" pageId="3" pageNumber="148">
Penis (
<figureCitation id="134E2A69FFB3FF86FE92FE7ED140FE54" box="[274,340,439,465]" captionStart="FIGURES 513" captionStartId="2.[152,270,1652,1676]" captionTargetBox="[176,1410,392,1615]" captionTargetId="figure-108@2.[164,1423,381,1628]" captionTargetPageId="2" captionText="FIGURES 513. Structural details. 58. Laohajekia uniformis. 5. Head and rostrum, dorsal view. 6. Head and rostrum, lateral view. 7. Penis, dorsal and lateral view. 8. Spermatheca. 913. Trachyphloeosoma newthayorum. 9. Head and rostrum, dorsal view. 10. Head and rostrum, lateral view. 11. Penis, dorsal and lateral view. 12. Spermatheca. 13. Sternite VIII in female. Scale bars: 0.50 mm (5, 6, 9, 10, 13), 0.25 mm (7, 11) and 0.125 mm (8, 12)." figureDoi="http://doi.org/10.5281/zenodo.13746097" httpUri="https://zenodo.org/record/13746097/files/figure.png" pageId="3" pageNumber="148">Fig. 7</figureCitation>
) in dorsal view widest at apical third, from base slightly evenly enlarging to this point, then slightly tapering apicad, apically broadly rounded; in lateral view distinctly regularly curved, evenly tapering apicad, tip narrow, elongate, slightly curved outside; endophallus with one short, straight sclerite; temones long, about twice as long as body of penis. Tegmen lacking parameres, its apodeme short, only faintly longer than diameter of ring.
</paragraph>
<paragraph id="8BCA36ECFFB3FF86FF47FD8ED504FD00" blockId="3.[151,1437,151,1617]" pageId="3" pageNumber="148">
Spermatheca (
<figureCitation id="134E2A69FFB3FF86FEEAFD8ED1BBFDE4" box="[362,431,583,609]" captionStart="FIGURES 513" captionStartId="2.[152,270,1652,1676]" captionTargetBox="[176,1410,392,1615]" captionTargetId="figure-108@2.[164,1423,381,1628]" captionTargetPageId="2" captionText="FIGURES 513. Structural details. 58. Laohajekia uniformis. 5. Head and rostrum, dorsal view. 6. Head and rostrum, lateral view. 7. Penis, dorsal and lateral view. 8. Spermatheca. 913. Trachyphloeosoma newthayorum. 9. Head and rostrum, dorsal view. 10. Head and rostrum, lateral view. 11. Penis, dorsal and lateral view. 12. Spermatheca. 13. Sternite VIII in female. Scale bars: 0.50 mm (5, 6, 9, 10, 13), 0.25 mm (7, 11) and 0.125 mm (8, 12)." figureDoi="http://doi.org/10.5281/zenodo.13746097" httpUri="https://zenodo.org/record/13746097/files/figure.png" pageId="3" pageNumber="148">Fig. 8</figureCitation>
) with slender and regularly curved cornu; ramus isodiametric; collum wider than long, equally long as ramus but conspicuously wider, regularly rounded. Female sternite VIII not examined.
</paragraph>
</subSubSection>
<subSubSection id="C36F6567FFB3FF86FF47FD46D53EFD2C" box="[199,1322,655,681]" pageId="3" pageNumber="148" type="etymology">
<paragraph id="8BCA36ECFFB3FF86FF47FD46D53EFD2C" blockId="3.[151,1437,151,1617]" box="[199,1322,655,681]" pageId="3" pageNumber="148">
<emphasis id="B901EAFEFFB3FF86FF47FD46D15AFD2C" bold="true" box="[199,334,655,681]" pageId="3" pageNumber="148">Etymology.</emphasis>
This species is named uniformis in recognition of the uniformly flat elytral interstriae.
</paragraph>
</subSubSection>
<subSubSection id="C36F6567FFB3FF86FF47FD7BD4B9FC90" pageId="3" pageNumber="148" type="biology_ecology">
<paragraph id="8BCA36ECFFB3FF86FF47FD7BD4B9FC90" blockId="3.[151,1437,151,1617]" pageId="3" pageNumber="148">
<emphasis id="B901EAFEFFB3FF86FF47FD7BD133FD49" bold="true" box="[199,295,690,716]" pageId="3" pageNumber="148">Biology.</emphasis>
The two known specimens were sifted from forest floor litter in a middle elevation fairly large (ca.
<quantity id="4C8D9B09FFB3FF86FA0EFD7AD0AFFD74" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="148" unit="km" value="1.0">1 km</quantity>
<superScript id="7C009BA4FFB3FF86FF3BFD1FD0D7FD61" attach="left" box="[187,195,726,740]" fontSize="6" pageId="3" pageNumber="148">2</superScript>
) patch of minimally disturbed old-growth tropical hardwood forest on level or gently rolling ground. This patch was surrounded by cultivated fields or scrubby cut-over forest, with rice paddies to the south.
</paragraph>
</subSubSection>
<subSubSection id="C36F6567FFB3FF86FF47FCD7D50BFB24" pageId="3" pageNumber="148" type="diagnosis">
<paragraph id="8BCA36ECFFB3FF86FF47FCD7D23DFC04" blockId="3.[151,1437,151,1617]" pageId="3" pageNumber="148">
<emphasis id="B901EAFEFFB3FF86FF47FCD7D1DBFCBD" bold="true" box="[199,463,798,824]" pageId="3" pageNumber="148">Differential diagnosis.</emphasis>
<taxonomicName id="4C754D6FFFB3FF86FE57FCD6D2DCFCBC" authorityName="Borovec and Anderson" authorityYear="2024" box="[471,712,799,825]" class="Insecta" family="Curculionidae" genus="Laohajekia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="species" species="uniformis">
<emphasis id="B901EAFEFFB3FF86FE57FCD6D2DCFCBC" box="[471,712,799,825]" italics="true" pageId="3" pageNumber="148">Laohajekia uniformis</emphasis>
</taxonomicName>
is very similar to the only other known species of the genus,
<taxonomicName id="4C754D6FFFB3FF86FA06FCD6D1C0FCD8" authority="Borovec" authorityName="Borovec" authorityYear="2014" class="Insecta" family="Curculionidae" genus="Laohajekia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="species" species="trachyphloeiformis">
<emphasis id="B901EAFEFFB3FF86FA06FCD6D178FCD8" italics="true" pageId="3" pageNumber="148">L. trachyphloeiformis</emphasis>
Borovec
</taxonomicName>
described from
<collectingRegion id="49B1F80EFFB3FF86FD11FC8AD395FCD8" box="[657,897,835,861]" country="Laos" name="Champasak" pageId="3" pageNumber="148">Champasak province</collectingRegion>
, also in
<collectingCountry id="F362767CFFB3FF86FC63FC8AD409FCD8" box="[995,1053,835,861]" name="Laos" pageId="3" pageNumber="148">Laos</collectingCountry>
. It is possible to separate the two species by the following characters:
</paragraph>
<paragraph id="8BCA36ECFFB3FF86FF47FC42D22DFB94" blockId="3.[151,1437,151,1617]" pageId="3" pageNumber="148">
<taxonomicName id="4C754D6FFFB3FF86FF47FC42D1ACFC20" authorityName="Borovec and Anderson" authorityYear="2024" box="[199,440,907,933]" class="Insecta" family="Curculionidae" genus="Laohajekia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="species" species="uniformis">
<emphasis id="B901EAFEFFB3FF86FF47FC42D1ACFC20" box="[199,440,907,933]" italics="true" pageId="3" pageNumber="148">Laohajekia uniformis</emphasis>
</taxonomicName>
: Antennal scrobes clearly visible in anterior part of rostrum; eyes in dorsal view slightly prominent from outline of head; erect elytral setae almost as long as width of interstria at posterior declivity; all elytral interstriae flat; rostrum 1.171.19× as wide as long; pronotum slender, 1.421.47× as wide as long; scapes slender, 4.7× as long as at apex wide.
</paragraph>
<paragraph id="8BCA36ECFFB3FF86FF47FBD2D50BFB24" blockId="3.[151,1437,151,1617]" pageId="3" pageNumber="148">
<taxonomicName id="4C754D6FFFB3FF86FF47FBD2D235FBB0" authorityName="Borovec" authorityYear="2014" box="[199,545,1051,1077]" class="Insecta" family="Curculionidae" genus="Laohajekia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="species" species="trachyphloeiformis">
<emphasis id="B901EAFEFFB3FF86FF47FBD2D235FBB0" box="[199,545,1051,1077]" italics="true" pageId="3" pageNumber="148">Laohajekia trachyphloeiformis</emphasis>
</taxonomicName>
: Antennal scrobes hardly visible as short and narrow furrows in anterior part of rostrum; eyes small, in dorsal view inconspicuous; erect elytral setae about as long as half the width of interstria at posterior declivity; interstriae 2, 4 and 6 slightly elevated, all interstriae with low, scarce bumps; rostrum 1.40× as wide as long; pronotum wider, 1.51× as wide as long; scapes more robust, 3.6× as long as at apex wide.
</paragraph>
</subSubSection>
<subSubSection id="C36F6567FFB3FF86FF47FB62D488F9D4" pageId="3" pageNumber="148" type="discussion">
<paragraph id="8BCA36ECFFB3FF86FF47FB62D488F9D4" blockId="3.[151,1437,151,1617]" pageId="3" pageNumber="148">
<emphasis id="B901EAFEFFB3FF86FF47FB62D12FFB40" bold="true" box="[199,315,1195,1221]" pageId="3" pageNumber="148">Remarks.</emphasis>
The monotypic genus
<taxonomicName id="4C754D6FFFB3FF86FDBBFB62D317FB40" authority="Morimoto" authorityName="Morimoto" authorityYear="2015" box="[571,771,1195,1221]" class="Insecta" family="Curculionidae" genus="Tokara" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="genus">
<emphasis id="B901EAFEFFB3FF86FDBBFB62D29DFB40" box="[571,649,1195,1221]" italics="true" pageId="3" pageNumber="148">Tokara</emphasis>
Morimoto
</taxonomicName>
described and known from
<collectingCountry id="F362767CFFB3FF86FBBCFB62D468FB40" box="[1084,1148,1195,1221]" name="Japan" pageId="3" pageNumber="148">Japan</collectingCountry>
following the description of
<taxonomicName id="4C754D6FFFB3FF86FF34FB06D182FB6C" authority="Borovec" authorityName="Borovec" authorityYear="2014" box="[180,406,1231,1257]" class="Insecta" family="Curculionidae" genus="Laohajekia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="genus">
<emphasis id="B901EAFEFFB3FF86FF34FB06D126FB6C" box="[180,306,1231,1257]" italics="true" pageId="3" pageNumber="148">Laohajekia</emphasis>
Borovec
</taxonomicName>
is very similar to
<taxonomicName id="4C754D6FFFB3FF86FDDAFB06D2CCFB6C" authorityName="Borovec" authorityYear="2014" box="[602,728,1231,1257]" class="Insecta" family="Curculionidae" genus="Laohajekia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="genus">
<emphasis id="B901EAFEFFB3FF86FDDAFB06D2CCFB6C" box="[602,728,1231,1257]" italics="true" pageId="3" pageNumber="148">Laohajekia</emphasis>
</taxonomicName>
. Based on its original description (
<bibRefCitation id="EFE44B1DFFB3FF86FBD6FB06D51DFB6C" author="Morimoto, K." box="[1110,1289,1230,1257]" pageId="3" pageNumber="148" refId="ref4157" refString="Morimoto, K. (2015) [new taxa] In: Morimoto, K., Nakamura, T., Kanno, K. Curculionidae: Entiminae (Part 2) (Coleoptera). The Insects of Japan. Vol. 4. Touka Shobo, Fukuoka, pp. 1 - 758." type="book" year="2015">Morimoto 2015</bibRefCitation>
) it is difficult to find any distinguishing characters except the funicle of 6 antennomeres and flagellate penis in
<taxonomicName id="4C754D6FFFB3FF86FB53FB3AD535FA88" authorityName="Morimoto" authorityYear="2015" box="[1235,1313,1267,1293]" class="Insecta" family="Curculionidae" genus="Tokara" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="genus">
<emphasis id="B901EAFEFFB3FF86FB53FB3AD535FA88" box="[1235,1313,1267,1293]" italics="true" pageId="3" pageNumber="148">Tokara</emphasis>
</taxonomicName>
. While the number of funicle antennomeres clearly distinguishes
<taxonomicName id="4C754D6FFFB3FF86FD6BFADED445FAB4" authority="Morimoto" authorityName="Morimoto" authorityYear="2015" box="[747,1105,1303,1329]" class="Insecta" family="Curculionidae" genus="Tokara" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="species" species="rotundipennis">
<emphasis id="B901EAFEFFB3FF86FD6BFADED3CEFAB4" box="[747,986,1303,1329]" italics="true" pageId="3" pageNumber="148">Tokara rotundipennis</emphasis>
Morimoto
</taxonomicName>
from both
<taxonomicName id="4C754D6FFFB3FF86FB48FADED588FAB4" box="[1224,1436,1303,1329]" pageId="3" pageNumber="148">
<emphasis id="B901EAFEFFB3FF86FB48FADED552FAB4" box="[1224,1350,1303,1329]" italics="true" pageId="3" pageNumber="148">Laohajekia</emphasis>
species
</taxonomicName>
from
<collectingCountry id="F362767CFFB3FF86FF57FAF2D104FAD0" box="[215,272,1339,1365]" name="Laos" pageId="3" pageNumber="148">Laos</collectingCountry>
, it is difficult to use this as a basis for distinguishing the genus. The number of funicle antennomeres often varies in Palaearctic
<taxonomicName id="4C754D6FFFB3FF86FE5AFA96D29CFAFC" baseAuthorityName="Borovec and Anderson" baseAuthorityYear="2022" box="[474,648,1375,1401]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="tribe" tribe="Trachyphloeini">Trachyphloeini</taxonomicName>
. For example, the nominal genus
<taxonomicName id="4C754D6FFFB3FF86FBA9FA96D526FAFC" authority="Germar" authorityName="Germar" authorityYear="1817" box="[1065,1330,1375,1401]" class="Insecta" family="Curculionidae" genus="Trachyphloeus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="genus">
<emphasis id="B901EAFEFFB3FF86FBA9FA96D4DAFAFC" box="[1065,1230,1375,1401]" italics="true" pageId="3" pageNumber="148">Trachyphloeus</emphasis>
Germar
</taxonomicName>
contains species with funicles having 5, 6 or 7 antennomeres. Also, the existence of a flagellum varies within species of a genus. For example, species of
<taxonomicName id="4C754D6FFFB3FF86FD83FA6ED305FA44" authority="Schoenherr" authorityName="Schoenherr" authorityYear="1845" box="[515,785,1447,1473]" class="Insecta" family="Curculionidae" genus="Porophorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="genus">
<emphasis id="B901EAFEFFB3FF86FD83FA6ED29CFA44" box="[515,648,1447,1473]" italics="true" pageId="3" pageNumber="148">Porophorus</emphasis>
Schoenherr
</taxonomicName>
(Tanyrhynchini Schoenherr) or
<taxonomicName id="4C754D6FFFB3FF86FB01FA6ED589FA44" authority="Schoenherr" authorityName="Schoenherr" authorityYear="1847" box="[1153,1437,1447,1473]" class="Insecta" family="Curculionidae" genus="Glyptosomus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="genus">
<emphasis id="B901EAFEFFB3FF86FB01FA6ED507FA44" box="[1153,1299,1447,1473]" italics="true" pageId="3" pageNumber="148">Glyptosomus</emphasis>
Schoenherr
</taxonomicName>
(Embrithini Marshall) either contain a long and conspicuous flagellum or the flagellum is completely absent. In all other characters, including male and female genitalia, which are well-illustrated in the description of
<taxonomicName id="4C754D6FFFB3FF86FA9CFA26D57EF98C" authorityName="Morimoto" authorityYear="2015" box="[1308,1386,1519,1545]" class="Insecta" family="Curculionidae" genus="Tokara" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="genus">
<emphasis id="B901EAFEFFB3FF86FA9CFA26D57EF98C" box="[1308,1386,1519,1545]" italics="true" pageId="3" pageNumber="148">Tokara</emphasis>
</taxonomicName>
, the characters of the two genera are very similar. It is apparent both genera are very close, but without the possibility of examining material of
<taxonomicName id="4C754D6FFFB3FF86FE15F9FED1F7F9D4" authorityName="Morimoto" authorityYear="2015" box="[405,483,1591,1617]" class="Insecta" family="Curculionidae" genus="Tokara" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="genus">
<emphasis id="B901EAFEFFB3FF86FE15F9FED1F7F9D4" box="[405,483,1591,1617]" italics="true" pageId="3" pageNumber="148">Tokara</emphasis>
</taxonomicName>
, it is not possible to synonymize (distinguish) the two genera.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,280 @@
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<emphasis id="B901EAFEFFB3FF86FF17F953D23FF930" bold="true" box="[151,555,1690,1717]" italics="true" pageId="3" pageNumber="148">Trachyphloeosoma newthayorum</emphasis>
Borovec and Anderson
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(
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,
<figureCitation id="134E2A69FFB3FF86FE90F92AD15DF978" box="[272,329,1763,1789]" captionStart="FIGURES 513" captionStartId="2.[152,270,1652,1676]" captionTargetBox="[176,1410,392,1615]" captionTargetId="figure-108@2.[164,1423,381,1628]" captionTargetPageId="2" captionText="FIGURES 513. Structural details. 58. Laohajekia uniformis. 5. Head and rostrum, dorsal view. 6. Head and rostrum, lateral view. 7. Penis, dorsal and lateral view. 8. Spermatheca. 913. Trachyphloeosoma newthayorum. 9. Head and rostrum, dorsal view. 10. Head and rostrum, lateral view. 11. Penis, dorsal and lateral view. 12. Spermatheca. 13. Sternite VIII in female. Scale bars: 0.50 mm (5, 6, 9, 10, 13), 0.25 mm (7, 11) and 0.125 mm (8, 12)." figureDoi="http://doi.org/10.5281/zenodo.13746097" httpUri="https://zenodo.org/record/13746097/files/figure.png" pageId="3" pageNumber="148">913</figureCitation>
)
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<paragraph id="8BCA36ECFFB3FF86FF17F8E2D4F3F834" blockId="3.[151,1437,1834,2041]" pageId="3" pageNumber="148">
<emphasis id="B901EAFEFFB3FF86FF17F8E2D156F8C1" bold="true" box="[151,322,1834,1861]" pageId="3" pageNumber="148">Type material.</emphasis>
<typeStatus id="54CE884EFFB3FF86FEC6F8E2D1A7F8C0" box="[326,435,1835,1861]" pageId="3" pageNumber="148" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D73FD65FFB3FF86FE3BF8E2D1C6F8C0" box="[443,466,1835,1861]" count="1" pageId="3" pageNumber="148" type="male"></specimenCount>
,
<collectingCountry id="F362767CFFB3FF86FE5BF8E2D200F8C0" box="[475,532,1835,1861]" name="Laos" pageId="3" pageNumber="148">Laos</collectingCountry>
,
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, Nakai-Nam Theun NBCA (N end), ca
<quantity id="4C8D9B09FFB3FF86FB40F8E2D4E2F8C0" box="[1216,1270,1835,1861]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="148" unit="km" value="2.0">2 km</quantity>
SW Kaew Nua Pass,
<quantity id="4C8D9B09FFB3FF86FF58F886D141F8EC" box="[216,341,1871,1897]" metricMagnitude="2" metricUnit="m" metricValue="7.0" metricValueMax="7.1" metricValueMin="6.9" pageId="3" pageNumber="148" unit="m" value="700.0" valueMax="710.0" valueMin="690.0">690-710 m</quantity>
,
<geoCoordinate id="EE41502BFFB3FF86FEE2F886D1E1F8EC" box="[354,501,1871,1897]" degrees="18" direction="north" minutes="22.292" orientation="latitude" pageId="3" pageNumber="148" precision="1" value="18.371532">18°22.292 N</geoCoordinate>
,
<geoCoordinate id="EE41502BFFB3FF86FD81F887D28AF8EC" box="[513,670,1870,1897]" degrees="105" direction="east" minutes="09.152" orientation="longitude" pageId="3" pageNumber="148" precision="1" value="105.152534">105°09.152 E</geoCoordinate>
,
<date id="FFCB102CFFB3FF86FD2BF887D303F8EC" box="[683,791,1870,1897]" pageId="3" pageNumber="148" value="2008-06-05">5.vi.2008</date>
, hardwood and bamboo forest w/tree ferns, FMHD2008- 029, berlese, litter incl. bamboo and under flowers of
<taxonomicName id="4C754D6FFFB3FF86FD74F8BAD39DF808" box="[756,905,1907,1933]" class="Magnoliopsida" family="Calophyllaceae" genus="Mesua" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="148" phylum="Tracheophyta" rank="species" species="ferrea">
<emphasis id="B901EAFEFFB3FF86FD74F8BAD39DF808" box="[756,905,1907,1933]" italics="true" pageId="3" pageNumber="148">Mesua ferrea</emphasis>
</taxonomicName>
(
<taxonomicName id="4C754D6FFFB3FF86FC18F8BAD403F808" box="[920,1047,1907,1933]" class="Magnoliopsida" family="Clusiaceae" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="148" phylum="Tracheophyta" rank="family">Clusiaceae</taxonomicName>
), A. Newton &amp; M. Thayer, ANMT site 1227 (
<collectionCode id="ED64AE29FFB3FF86FE8CF85ED177F834" box="[268,355,1943,1969]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="3" pageNumber="148" type="Museum">FMNH</collectionCode>
).
<typeStatus id="54CE884EFFB3FF86FEF6F85ED1F3F834" box="[374,487,1943,1969]" pageId="3" pageNumber="148" type="paratype">Paratypes</typeStatus>
:
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, same data as
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(2
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, 3
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, 1
<collectionCode id="ED64AE29FFB3FF86FB0EF85ED4CEF834" box="[1166,1242,1943,1969]" pageId="3" pageNumber="148">RBSC</collectionCode>
).
</paragraph>
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Body length:
<typeStatus id="54CE884EFFB3FF86FEE2F872D1D0F850" box="[354,452,1979,2005]" pageId="3" pageNumber="148" type="holotype">holotype</typeStatus>
<quantity id="4C8D9B09FFB3FF86FE4CF872D220F850" box="[460,564,1979,2005]" metricMagnitude="-3" metricUnit="m" metricValue="2.09" pageId="3" pageNumber="148" unit="mm" value="2.09">2.09 mm</quantity>
,
<typeStatus id="54CE884EFFB3FF86FDC1F872D2B8F850" box="[577,684,1979,2005]" pageId="3" pageNumber="148" type="paratype">paratypes</typeStatus>
<quantity id="4C8D9B09FFB3FF86FD34F872D34FF850" box="[692,859,1979,2005]" metricMagnitude="-3" metricUnit="m" metricValue="2.3499999999999996" metricValueMax="2.61" metricValueMin="2.09" pageId="3" pageNumber="148" unit="mm" value="2.3499999999999996" valueMax="2.61" valueMin="2.09">2.092.61 mm</quantity>
. Body (
<figureCitation id="134E2A69FFB3FF86FC34F872D40EF850" box="[948,1050,1979,2005]" captionStart="FIGURES 14" captionStartId="1.[152,270,1594,1618]" captionTargetBox="[154,1436,771,1548]" captionTargetId="figure-218@1.[151,1436,771,1568]" captionTargetPageId="1" captionText="FIGURES 14. Dorsal and lateral habitus. 12. Laohajekia uniformis. 34. Trachyphloeosoma newthayorum." figureDoi="http://doi.org/10.5281/zenodo.13746095" httpUri="https://zenodo.org/record/13746095/files/figure.png" pageId="3" pageNumber="148">Figs 3, 4</figureCitation>
) dark brownish, antennal funicles with clubs and tarsi slightly paler. Entire body except for antennae and tarsi covered with encrustation not allowing examination of appressed scales. Each elytral interstria with one dense and conspicuous row of erect piliform setae, slightly shorter than width of interstria; distance between two setae almost twice as long as length of one seta. Pronotum with similar setae, only slightly shorter than elytral ones, densely scattered. Head with rostrum with similar setae, slightly shorter than those on pronotum. Antennal scapes, femora and tibiae with short, erect piliform setae.
</paragraph>
<paragraph id="8BCA36ECFFB4FF81FF47FE82D3DDFD2C" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="149">
Rostrum (
<figureCitation id="134E2A69FFB4FF81FEBAFE82D1A7FEE0" box="[314,435,331,357]" captionStart="FIGURES 513" captionStartId="2.[152,270,1652,1676]" captionTargetBox="[176,1410,392,1615]" captionTargetId="figure-108@2.[164,1423,381,1628]" captionTargetPageId="2" captionText="FIGURES 513. Structural details. 58. Laohajekia uniformis. 5. Head and rostrum, dorsal view. 6. Head and rostrum, lateral view. 7. Penis, dorsal and lateral view. 8. Spermatheca. 913. Trachyphloeosoma newthayorum. 9. Head and rostrum, dorsal view. 10. Head and rostrum, lateral view. 11. Penis, dorsal and lateral view. 12. Spermatheca. 13. Sternite VIII in female. Scale bars: 0.50 mm (5, 6, 9, 10, 13), 0.25 mm (7, 11) and 0.125 mm (8, 12)." figureDoi="http://doi.org/10.5281/zenodo.13746097" httpUri="https://zenodo.org/record/13746097/files/figure.png" pageId="4" pageNumber="149">Figs 9, 10</figureCitation>
) short and wide, 1.061.11× as wide as long, widest at base, slightly evenly tapering apicad with straight sides, at base 1.091.14× as wide as at apex, apically broadly rounded; in lateral view regularly vaulted, in apical portion declined. Epifrons widest at base, distinctly tapering anteriad with straight sides, dorsally with indistinct median longitudinal furrow. Epistome not developed; fronto-epistomal area short and wide, glabrous, smooth, posteriorly contrastive separated from squamosa epifrons.Antennal scrobes in dorsal view visible as narrow furrows throughout entire length, in apical portion narrowly reniform; in lateral view conspicuously enlarged posteriad, with dorsal margin directed towards dorsal margin of eye, then curved downwards and reaching ventral margin of eye; ventral margin directed downwards and almost reaching ventral margin of rostrum. Eyes small, in dorsal view faintly prominent outline of head; in lateral view oval, placed above middle of head hight, its diameter smaller than width of scape at apex. Head coarsely punctate behind eyes.
</paragraph>
<paragraph id="8BCA36ECFFB4FF81FF47FD7AD3DEFCD8" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="149">Antennal scapes slender, faintly exceeding anterior margin of pronotum in repose, 4.5 × as long as wide at apex, weakly evenly enlarging apicad, at apex 0.60.7× as wide as club, curved at midlength. Funicle of 7 antennomeres; antennomere 1 1.92.1× as long as wide and 1.71.8× as long as antennomere 2, this is 1.51.6× as long as wide; antennomeres 3 and 4 isodiametric; antennomere 5 1.1× as wide as long; antennomere 6 1.21.3× as wide as long; antennomere 7 1.41.5× as wide as long; clubs 1.51.6× as long as wide.</paragraph>
<paragraph id="8BCA36ECFFB4FF81FF47FCAED34BFC68" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="149">
Pronotum (
<figureCitation id="134E2A69FFB4FF81FEC7FCAED1A5FC04" box="[327,433,871,897]" captionStart="FIGURES 14" captionStartId="1.[152,270,1594,1618]" captionTargetBox="[154,1436,771,1548]" captionTargetId="figure-218@1.[151,1436,771,1568]" captionTargetPageId="1" captionText="FIGURES 14. Dorsal and lateral habitus. 12. Laohajekia uniformis. 34. Trachyphloeosoma newthayorum." figureDoi="http://doi.org/10.5281/zenodo.13746095" httpUri="https://zenodo.org/record/13746095/files/figure.png" pageId="4" pageNumber="149">Figs 3, 4</figureCitation>
) as wide as long to 1.07× as wide as long, widest at anterior third, constricted behind anterior margin; anterior margin narrower than posterior one. Disc roughly, densely irregularly punctured with space between punctures conspicuously shorter than their diameter. Pronotum in lateral view regularly domed. Anterior margin bent strongly back beneath and towards coxae.
</paragraph>
<paragraph id="8BCA36ECFFB4FF81FF47FC3ED1A9FB94" blockId="4.[151,1437,151,2013]" box="[199,445,1015,1041]" pageId="4" pageNumber="149">Scutellum not visible.</paragraph>
<paragraph id="8BCA36ECFFB4FF81FF47FBD2D38CFBF8" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="149">
Elytra (
<figureCitation id="134E2A69FFB4FF81FE9CFBD2D196FBB0" box="[284,386,1051,1077]" captionStart="FIGURES 14" captionStartId="1.[152,270,1594,1618]" captionTargetBox="[154,1436,771,1548]" captionTargetId="figure-218@1.[151,1436,771,1568]" captionTargetPageId="1" captionText="FIGURES 14. Dorsal and lateral habitus. 12. Laohajekia uniformis. 34. Trachyphloeosoma newthayorum." figureDoi="http://doi.org/10.5281/zenodo.13746095" httpUri="https://zenodo.org/record/13746095/files/figure.png" pageId="4" pageNumber="149">Figs 3, 4</figureCitation>
) oval, 1.321.38× as long as wide, widest slightly before midlength, with regularly rounded sides, apically narrowly rounded. Striae densely coarsely punctured, interstriae flat, slightly wider than puncture diameter, somewhat smooth. Elytra in lateral view regularly vaulted.
</paragraph>
<paragraph id="8BCA36ECFFB4FF81FF47FB4ED2DBFAB4" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="149">Femora edentate. Protibiae somewhat slender, 5.155.34× as long as at middle wide, in apical quarter distinctly curved inside and here asquamose and densely covered with short appressed setae, apically subtruncated with dense fringe of short yellowish setae; all tibiae with long conspicuous mucro. Tarsi short; tarsomere 2 1.21.3× as wide as long; tarsomere 3 1.31.4× as wide as long and 1.31.4× as wide as tarsomere 2; onychium 0.80.9× as long as tarsomere 3; claws long, free, distinctly divaricate.</paragraph>
<paragraph id="8BCA36ECFFB4FF81FF47FAF2D383FA44" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="149">
Abdominal ventrites 1.091.15× as long as wide, roughly punctured, space between punctures slightly longer than their diameter; ventrite 1 at middle 1.5× as long as ventrite 2; ventrite 2 slightly longer than ventrites 3 and 4 together; ventrite
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males apically concave. Suture between ventrites 1 and 2 sinuose, the other straight. Metaventral process narrower than transverse diameter of metacoxa.
</paragraph>
<paragraph id="8BCA36ECFFB4FF81FF47FA02D356F9D4" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="149">
Penis (
<figureCitation id="134E2A69FFB4FF81FE93FA02D176FA60" box="[275,354,1483,1509]" captionStart="FIGURES 513" captionStartId="2.[152,270,1652,1676]" captionTargetBox="[176,1410,392,1615]" captionTargetId="figure-108@2.[164,1423,381,1628]" captionTargetPageId="2" captionText="FIGURES 513. Structural details. 58. Laohajekia uniformis. 5. Head and rostrum, dorsal view. 6. Head and rostrum, lateral view. 7. Penis, dorsal and lateral view. 8. Spermatheca. 913. Trachyphloeosoma newthayorum. 9. Head and rostrum, dorsal view. 10. Head and rostrum, lateral view. 11. Penis, dorsal and lateral view. 12. Spermatheca. 13. Sternite VIII in female. Scale bars: 0.50 mm (5, 6, 9, 10, 13), 0.25 mm (7, 11) and 0.125 mm (8, 12)." figureDoi="http://doi.org/10.5281/zenodo.13746097" httpUri="https://zenodo.org/record/13746097/files/figure.png" pageId="4" pageNumber="149">Fig. 11</figureCitation>
) 2.3× as long as wide, in dorsal view subparallel-sided, apical portion subtriangular with slightly concave sides, tip narrow, apically shortly obtuse; in lateral view narrow, in middle part almost straight, tip curved inside, evenly pointed. Temones twice as long as body of penis. Sternite IX with spiculum gastrale anteriorly curved, narrowed, posteriorly with basal arms fused, slender.
</paragraph>
<paragraph id="8BCA36ECFFB4FF81FF47F992D3BDF964" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="149">
Spermatheca (
<figureCitation id="134E2A69FFB4FF81FEE7F992D1ADF9F0" box="[359,441,1627,1653]" captionStart="FIGURES 513" captionStartId="2.[152,270,1652,1676]" captionTargetBox="[176,1410,392,1615]" captionTargetId="figure-108@2.[164,1423,381,1628]" captionTargetPageId="2" captionText="FIGURES 513. Structural details. 58. Laohajekia uniformis. 5. Head and rostrum, dorsal view. 6. Head and rostrum, lateral view. 7. Penis, dorsal and lateral view. 8. Spermatheca. 913. Trachyphloeosoma newthayorum. 9. Head and rostrum, dorsal view. 10. Head and rostrum, lateral view. 11. Penis, dorsal and lateral view. 12. Spermatheca. 13. Sternite VIII in female. Scale bars: 0.50 mm (5, 6, 9, 10, 13), 0.25 mm (7, 11) and 0.125 mm (8, 12)." figureDoi="http://doi.org/10.5281/zenodo.13746097" httpUri="https://zenodo.org/record/13746097/files/figure.png" pageId="4" pageNumber="149">Fig. 12</figureCitation>
) with slender and regularly curved cornu; corpus large, elongated; ramus not developed; collum very small, hump-shaped. Sternite VIII with long and slender apodeme, Y-shaped and terminating in basal part of plate; plate small, subtriangular with slender fenestra reaching middle of plate. Gonocoxites slender and long, in apical portion tube-shaped, with laterally directed apical styli.
</paragraph>
</subSubSection>
<subSubSection id="C36F6567FFB4FF81FF47F922D0F8F8C8" pageId="4" pageNumber="149" type="etymology">
<paragraph id="8BCA36ECFFB4FF81FF47F922D0F8F8C8" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="149">
<emphasis id="B901EAFEFFB4FF81FF47F922D15AF880" bold="true" box="[199,334,1771,1797]" pageId="4" pageNumber="149">Etymology.</emphasis>
This species is named after the collectors of the
<typeStatus id="54CE884EFFB4FF81FC11F922D3D5F880" box="[913,961,1771,1797]" pageId="4" pageNumber="149">type</typeStatus>
series, Al Newton and Margaret Thayer, Field Museum of Natural History (retired) by combining the first syllable of each of their surnames with the suffix orum.
</paragraph>
</subSubSection>
<subSubSection id="C36F6567FFB4FF81FF47F89FD2BEF858" pageId="4" pageNumber="149" type="biology_ecology">
<paragraph id="8BCA36ECFFB4FF81FF47F89FD2BEF858" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="149">
<emphasis id="B901EAFEFFB4FF81FF47F89FD133F8F5" bold="true" box="[199,295,1878,1904]" pageId="4" pageNumber="149">Biology.</emphasis>
All known specimens were sifted from forest floor litter in a middle elevation mountainous area along the Laos-Vietnam border, with extensive, moderately disturbed hardwood (tropical montane) forest on moderate slopes, with bamboo and tree ferns common in the understory. This forest area is very extensive and runs along the border for tens of kilometers in both directions.
</paragraph>
</subSubSection>
<subSubSection id="C36F6567FFB5FF80FF47FF5FD119FCBD" pageId="5" pageNumber="150" type="diagnosis">
<paragraph id="8BCA36ECFFB5FF80FF47FF5FD119FCBD" blockId="5.[151,1437,150,824]" pageId="5" pageNumber="150">
<emphasis id="B901EAFEFFB5FF80FF47FF5FD1C2FF35" bold="true" box="[199,470,150,176]" pageId="5" pageNumber="150">Differential diagnosis.</emphasis>
<taxonomicName id="4C754D6FFFB5FF80FE64FF5FD374FF35" authorityName="Borovec and Anderson" authorityYear="2024" box="[484,864,150,176]" class="Insecta" family="Curculionidae" genus="Trachyphloeosoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="species" species="newthayorum">
<emphasis id="B901EAFEFFB5FF80FE64FF5FD374FF35" box="[484,864,150,176]" italics="true" pageId="5" pageNumber="150">Trachyphloeosoma newthayorum</emphasis>
</taxonomicName>
has a funicle with 7 antennomeres, all elytral interstriae with a conspicuous row of long erect setae only slightly longer than setae on pronotum, rostrum long, slender and pronotum widest at anterior third. In these characters it is similar to
<taxonomicName id="4C754D6FFFB5FF80FBB5FF16D589FF7D" authority="Ren, Borovec &amp; Zhang" authorityName="Ren, Borovec &amp; Zhang" box="[1077,1437,222,249]" class="Insecta" family="Curculionidae" genus="Trachyphloeosoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="species" species="jirka">
<emphasis id="B901EAFEFFB5FF80FBB5FF16D49CFF7D" box="[1077,1160,222,248]" italics="true" pageId="5" pageNumber="150">T. jirka</emphasis>
Ren, Borovec &amp; Zhang
</taxonomicName>
known from
<collectingCountry id="F362767CFFB5FF80FEADFECBD164FE99" box="[301,368,258,284]" name="China" pageId="5" pageNumber="150">China</collectingCountry>
and
<collectingCountry id="F362767CFFB5FF80FE2AFECBD21DFE99" box="[426,521,258,284]" name="Vietnam" pageId="5" pageNumber="150">Vietnam</collectingCountry>
and
<taxonomicName id="4C754D6FFFB5FF80FDC4FECAD30CFE98" authority="Borovec" authorityName="Borovec" authorityYear="2021" box="[580,792,258,285]" class="Insecta" family="Curculionidae" genus="Trachyphloeosoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="species" species="lirenae">
<emphasis id="B901EAFEFFB5FF80FDC4FECAD2A4FE99" box="[580,688,258,284]" italics="true" pageId="5" pageNumber="150">T. lirenae</emphasis>
Borovec
</taxonomicName>
known from
<collectingCountry id="F362767CFFB5FF80FC36FECBD40DFE99" box="[950,1049,258,284]" name="Vietnam" pageId="5" pageNumber="150">Vietnam</collectingCountry>
.
<taxonomicName id="4C754D6FFFB5FF80FBA5FECBD588FE99" authorityName="Borovec and Anderson" authorityYear="2024" box="[1061,1436,258,284]" class="Insecta" family="Curculionidae" genus="Trachyphloeosoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="species" species="newthayorum">
<emphasis id="B901EAFEFFB5FF80FBA5FECBD588FE99" box="[1061,1436,258,284]" italics="true" pageId="5" pageNumber="150">Trachyphloeosoma newthayorum</emphasis>
</taxonomicName>
can be distinguished from both species by small eyes, in lateral view with diameter smaller than width of scape at apex (eyes mid-sized with diameter slightly longer than width of scape at apex in both other species) and ventrite
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males apically concave (ventrite
<quantity id="4C8D9B09FFB5FF80FDBFFEA7D27EFE0D" box="[575,618,366,393]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="5" pageNumber="150" unit="in" value="5.0">5 in</quantity>
males apically obtuse in both other species). It also differs from
<taxonomicName id="4C754D6FFFB5FF80FACBFEA6D588FE0D" box="[1355,1436,366,392]" class="Insecta" family="Curculionidae" genus="Trachyphloeosoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="species" species="jirka">
<emphasis id="B901EAFEFFB5FF80FACBFEA6D588FE0D" box="[1355,1436,366,392]" italics="true" pageId="5" pageNumber="150">T. jirka</emphasis>
</taxonomicName>
by a long funicular antennomere 2 (1.11.2× as long as wide in
<taxonomicName id="4C754D6FFFB5FF80FCF4FE5AD3D2FE29" box="[884,966,402,428]" class="Insecta" family="Curculionidae" genus="Trachyphloeosoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="species" species="jirka">
<emphasis id="B901EAFEFFB5FF80FCF4FE5AD3D2FE29" box="[884,966,402,428]" italics="true" pageId="5" pageNumber="150">T. jirka</emphasis>
</taxonomicName>
), penis with tip more elongate and wider in dorsal view, slender at middle and straight in lateral view (tip shorter and very narrow in dorsal view and wide and regularly curved in lateral view in
<taxonomicName id="4C754D6FFFB5FF80FDD3FE12D2B1FE71" box="[595,677,474,500]" class="Insecta" family="Curculionidae" genus="Trachyphloeosoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="species" species="jirka">
<emphasis id="B901EAFEFFB5FF80FDD3FE12D2B1FE71" box="[595,677,474,500]" italics="true" pageId="5" pageNumber="150">T. jirka</emphasis>
</taxonomicName>
) and female sternite VIII with plate short with fenestra (plate long and slender lacking fenestra in
<taxonomicName id="4C754D6FFFB5FF80FE7DFE36D25BFD9D" box="[509,591,510,536]" class="Insecta" family="Curculionidae" genus="Trachyphloeosoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="species" species="jirka">
<emphasis id="B901EAFEFFB5FF80FE7DFE36D25BFD9D" box="[509,591,510,536]" italics="true" pageId="5" pageNumber="150">T. jirka</emphasis>
</taxonomicName>
). It also differs from
<taxonomicName id="4C754D6FFFB5FF80FCC8FE36D3A7FD9D" authorityName="Borovec" authorityYear="2021" box="[840,947,510,536]" class="Insecta" family="Curculionidae" genus="Trachyphloeosoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="species" species="lirenae">
<emphasis id="B901EAFEFFB5FF80FCC8FE36D3A7FD9D" box="[840,947,510,536]" italics="true" pageId="5" pageNumber="150">T. lirenae</emphasis>
</taxonomicName>
by the head behind eyes punctured (finely longitudinally striate in
<taxonomicName id="4C754D6FFFB5FF80FE27FDEAD206FDB9" authorityName="Borovec" authorityYear="2021" box="[423,530,546,572]" class="Insecta" family="Curculionidae" genus="Trachyphloeosoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="species" species="lirenae">
<emphasis id="B901EAFEFFB5FF80FE27FDEAD206FDB9" box="[423,530,546,572]" italics="true" pageId="5" pageNumber="150">T. lirenae</emphasis>
</taxonomicName>
) and penis with elongate tip (not so in
<taxonomicName id="4C754D6FFFB5FF80FC53FDEAD42AFDB9" authorityName="Borovec" authorityYear="2021" box="[979,1086,546,572]" class="Insecta" family="Curculionidae" genus="Trachyphloeosoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="species" species="lirenae">
<emphasis id="B901EAFEFFB5FF80FC53FDEAD42AFDB9" box="[979,1086,546,572]" italics="true" pageId="5" pageNumber="150">T. lirenae</emphasis>
</taxonomicName>
). In its elytra with dense, long erect setae
<taxonomicName id="4C754D6FFFB5FF80FE97FD8FD299FDE5" authorityName="Borovec and Anderson" authorityYear="2024" box="[279,653,582,608]" class="Insecta" family="Curculionidae" genus="Trachyphloeosoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="species" species="newthayorum">
<emphasis id="B901EAFEFFB5FF80FE97FD8FD299FDE5" box="[279,653,582,608]" italics="true" pageId="5" pageNumber="150">Trachyphloeosoma newthayorum</emphasis>
</taxonomicName>
is similar also to
<taxonomicName id="4C754D6FFFB5FF80FCDDFD8ED4FDFDE5" authority="Borovec &amp; Anderson, 2022" authorityName="Borovec &amp; Anderson" authorityYear="2022" box="[861,1257,582,609]" class="Insecta" family="Curculionidae" genus="Trachyphloeosoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="species" species="ales">
<emphasis id="B901EAFEFFB5FF80FCDDFD8ED3B3FDE5" box="[861,935,582,608]" italics="true" pageId="5" pageNumber="150">T. ales</emphasis>
<bibRefCitation id="EFE44B1DFFB5FF80FC30FD8ED4FDFDE5" author="Borovec, R. &amp; Anderson, R. S." box="[944,1257,582,609]" pageId="5" pageNumber="150" pagination="93 - 100" refId="ref4092" refString="Borovec, R. &amp; Anderson, R. S. (2022) Three new species of Trachyphloeosoma Wollaston (Coleoptera: Curculionidae: Entiminae) from southeast Asia with the first record of the tribe Trachyphloeini from the Philippines. Zootaxa, 5182 (1), 93 - 100. https: // doi. org / 10.11646 / zootaxa. 5182.1.7" type="journal article" year="2022">Borovec &amp; Anderson, 2022</bibRefCitation>
</taxonomicName>
described from
<collectingCountry id="F362767CFFB5FF80FF17FDA3D0F8FD01" box="[151,236,618,644]" name="Taiwan" pageId="5" pageNumber="150">Taiwan</collectingCountry>
, but it is easily distinguished by a long series of characters: pronotum widest at anterior third and regularly domed on disc (widest at midlength, with wide longitudinal median furrow in
<taxonomicName id="4C754D6FFFB5FF80FB90FD46D44EFD2D" authorityName="Borovec &amp; Anderson" authorityYear="2022" box="[1040,1114,654,680]" class="Insecta" family="Curculionidae" genus="Trachyphloeosoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="species" species="ales">
<emphasis id="B901EAFEFFB5FF80FB90FD46D44EFD2D" box="[1040,1114,654,680]" italics="true" pageId="5" pageNumber="150">T. ales</emphasis>
</taxonomicName>
), head behind eyes punctate (finely striate in
<taxonomicName id="4C754D6FFFB5FF80FED6FD7AD1B4FD49" authorityName="Borovec &amp; Anderson" authorityYear="2022" box="[342,416,690,716]" class="Insecta" family="Curculionidae" genus="Trachyphloeosoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="species" species="ales">
<emphasis id="B901EAFEFFB5FF80FED6FD7AD1B4FD49" box="[342,416,690,716]" italics="true" pageId="5" pageNumber="150">T. ales</emphasis>
</taxonomicName>
), abdominal ventrites sparsely punctate (densely punctate in
<taxonomicName id="4C754D6FFFB5FF80FBDCFD7AD4B2FD49" authorityName="Borovec &amp; Anderson" authorityYear="2022" box="[1116,1190,690,716]" class="Insecta" family="Curculionidae" genus="Trachyphloeosoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="species" species="ales">
<emphasis id="B901EAFEFFB5FF80FBDCFD7AD4B2FD49" box="[1116,1190,690,716]" italics="true" pageId="5" pageNumber="150">T. ales</emphasis>
</taxonomicName>
), penis with elongate tip (apically broadly rounded in
<taxonomicName id="4C754D6FFFB5FF80FD90FD1ED24FFD75" authorityName="Borovec &amp; Anderson" authorityYear="2022" box="[528,603,726,752]" class="Insecta" family="Curculionidae" genus="Trachyphloeosoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="species" species="ales">
<emphasis id="B901EAFEFFB5FF80FD90FD1ED24FFD75" box="[528,603,726,752]" italics="true" pageId="5" pageNumber="150">T. ales</emphasis>
</taxonomicName>
), spermatheca with small, hump-shaped collum (corpus lacking collum, apically constricted and then regularly rounded in
<taxonomicName id="4C754D6FFFB5FF80FD53FD32D308FC91" authorityName="Borovec &amp; Anderson" authorityYear="2022" box="[723,796,762,788]" class="Insecta" family="Curculionidae" genus="Trachyphloeosoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="species" species="ales">
<emphasis id="B901EAFEFFB5FF80FD53FD32D308FC91" box="[723,796,762,788]" italics="true" pageId="5" pageNumber="150">T. ales</emphasis>
</taxonomicName>
) and female sternite VIII with fenestra (lacking fenestra in
<taxonomicName id="4C754D6FFFB5FF80FF34FCD6D0E8FCBD" authorityName="Borovec &amp; Anderson" authorityYear="2022" box="[180,252,798,824]" class="Insecta" family="Curculionidae" genus="Trachyphloeosoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="species" species="ales">
<emphasis id="B901EAFEFFB5FF80FF34FCD6D0E8FCBD" box="[180,252,798,824]" italics="true" pageId="5" pageNumber="150">T. ales</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,120 @@
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<mods:title id="2DC302ACC388CB93AD29CC6F50F055AA">Two new Trachyphloeini (Coleoptera; Curculionidae; Entiminae) from Laos</mods:title>
</mods:titleInfo>
<mods:name id="07A03AD80787A7F4D57A468C55C932E9" type="personal">
<mods:role id="F19EA868450C0CF043894E531FB61555">
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<mods:namePart id="FFAF23BFDEE99B24960DB61E76CA395E">Borovec, Roman</mods:namePart>
<mods:affiliation id="2B174B920296C3FFE2201A89DAC31ED4">Sloupno 64, 503 53 Smidary, Czech Republic.</mods:affiliation>
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<taxonomicName id="4C754D6FFFB5FF80FF17FC43D253FC21" ID-CoL="57S6B" authority="(Sharp)" baseAuthorityName="Sharp" box="[151,583,906,933]" class="Insecta" family="Curculionidae" genus="Trachyphloeosoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="species" species="roelofsi">
<emphasis id="B901EAFEFFB5FF80FF17FC43D253FC21" bold="true" box="[151,583,906,933]" pageId="5" pageNumber="150">
<emphasis id="B901EAFEFFB5FF80FF17FC43D1F4FC20" bold="true" box="[151,480,906,933]" italics="true" pageId="5" pageNumber="150">Trachyphloeosoma roelofsi</emphasis>
(Sharp)
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<emphasis id="B901EAFEFFB5FF80FF17FC1BD16AFC69" bold="true" box="[151,382,978,1004]" pageId="5" pageNumber="150">Material examined.</emphasis>
<materialsCitation id="3B1D3CB1FFB5FF80FE03FC1BD59FFBB1" collectingDate="2008-06-12" collectionCode="CMNC, FMNH" collectorName="A. Newton &amp; M. Thayer" country="Laos" elevation="1170" latitude="15.241567" location="Ban Thongvay" longLatPrecision="1" longitude="106.53011" municipality="Bolaven Plateau" pageId="5" pageNumber="150" specimenCount="3" specimenCount-female="3" stateProvince="Champasak">
<specimenCount id="9D73FD65FFB5FF80FE03FC1BD1D5FC68" box="[387,449,978,1005]" count="3" pageId="5" pageNumber="150" type="female">3 ♀♀</specimenCount>
,
<collectingCountry id="F362767CFFB5FF80FE4CFC1AD211FC68" box="[460,517,979,1005]" name="Laos" pageId="5" pageNumber="150">Laos</collectingCountry>
,
<collectingRegion id="49B1F80EFFB5FF80FD8EFC1BD2EFFC69" box="[526,763,978,1004]" country="Laos" name="Champasak" pageId="5" pageNumber="150">Champasak Province</collectingRegion>
,
<collectingMunicipality id="6BAEAC96FFB5FF80FC85FC1BD3A8FC69" box="[773,956,978,1004]" pageId="5" pageNumber="150">Bolaven Plateau</collectingMunicipality>
,
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(Xékatam) vic., old logging road N of village,
<quantity id="4C8D9B09FFB5FF80FEEFFC3FD1DCFB94" box="[367,456,1014,1041]" metricMagnitude="3" metricUnit="m" metricValue="1.17" pageId="5" pageNumber="150" unit="m" value="1170.0">
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,
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,
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,
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<collectingDate id="EF8FE9C4FFB5FF80FCA8FC3FD3B6FB95" box="[808,930,1014,1040]" pageId="5" pageNumber="150" value="2008-06-12">12.vi.2008</collectingDate>
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, selectively logged forest, FMHD2008-046, winkler extract, log &amp; leaf litter, much on boulders,
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&amp;
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, ANMT site 1232 (
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,
<collectionCode id="ED64AE29FFB5FF80FAACFBD3D596FBB1" box="[1324,1410,1050,1076]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="5" pageNumber="150" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BCA36ECFFB5FF80FF17FBF7D0C1FBF8" blockId="5.[151,1437,978,1149]" pageId="5" pageNumber="150">
This species is known from
<collectingCountry id="F362767CFFB5FF80FE63FBF6D237FBDC" box="[483,547,1087,1113]" name="Japan" pageId="5" pageNumber="150">Japan</collectingCountry>
and
<collectingCountry id="F362767CFFB5FF80FDE0FBF7D2A6FBDD" box="[608,690,1086,1112]" name="Taiwan" pageId="5" pageNumber="150">Taiwan</collectingCountry>
(
<bibRefCitation id="EFE44B1DFFB5FF80FD43FBF7D407FBDD" author="Alonso-Zarazaga, M. A. &amp; Barrios, H. &amp; Borovec, R. &amp; Bouchard, P. &amp; Caldara, R. &amp; Colonnelli, E. &amp; Gultekin, L. &amp; Hlavac, P. &amp; Korotyaev, B. &amp; Lyal, C. H. C. &amp; Machado, A. &amp; Meregalli, M. &amp; Pierotti, H. &amp; Ren, L. &amp; Sanchez-Ruiz, M. &amp; Sforzi, A. &amp; Silfverberg, H. &amp; Skuhrovec, J. &amp; Tryzna, M. &amp; Velazquez de Castro, A. J. &amp; Yunakov, N. N." box="[707,1043,1086,1112]" pageId="5" pageNumber="150" pagination="1 - 780" refId="ref3918" refString="Alonso-Zarazaga, M. A., Barrios, H., Borovec, R., Bouchard, P., Caldara, R., Colonnelli, E., Gultekin, L., Hlavac, P., Korotyaev, B., Lyal, C. H. C., Machado, A., Meregalli, M., Pierotti, H., Ren, L., Sanchez-Ruiz, M., Sforzi, A., Silfverberg, H., Skuhrovec, J., Tryzna, M., Velazquez de Castro, A. J. &amp; Yunakov, N. N. (2023) Cooperative Catalogue of Palaearctic Coleoptera Curculionoidea. 2 nd Edition. Monografias electronicas SEA, 14, 1 - 780." type="journal article" year="2023">
Alonso-Zarazaga
<emphasis id="B901EAFEFFB5FF80FC11FBF6D3D8FBDD" box="[913,972,1086,1112]" italics="true" pageId="5" pageNumber="150">et al.</emphasis>
2023
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); this is a new country record for
<collectingCountry id="F362767CFFB5FF80FF17FBAAD0C5FBF8" box="[151,209,1123,1149]" name="Laos" pageId="5" pageNumber="150">Laos</collectingCountry>
.
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View file

@ -0,0 +1,289 @@
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<typeStatus id="54F5F0E4FFA25052FF23FD40FF0C46D7" box="[151,201,652,676]" pageId="82" pageNumber="83">Type</typeStatus>
species:
<taxonomicName id="4C4E35C5FFA25052FE92FD40FD6C46D0" authority="Franz, 1986 b: 244" authorityName="Franz" authorityPageNumber="244" authorityYear="1986" box="[294,681,651,675]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="nitidulus">
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<bibRefCitation id="EFDF33B7FFA25052FE44FD40FD6C46D0" author="Franz, H." box="[496,681,651,675]" pageId="82" pageNumber="83" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 244</bibRefCitation>
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(here designated).
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<emphasis id="B93A9254FFA25052FF23FD1DFED74698" bold="true" box="[151,274,721,747]" pageId="82" pageNumber="83">Diagnosis.</emphasis>
Body (
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) with distinct constriction between head and pronotum and between pronotum and elytra; thick bristles present on genae and outer regions of hypomera (
<figureCitation id="137552C3FFA25052FCF9FD39FC6A4763" box="[845,943,757,784]" captionStart="FIGURES 312320" captionStartId="84.[151,264,1812,1836]" captionTargetBox="[151,1430,365,1776]" captionTargetId="figure-59@84.[151,1435,365,1789]" captionTargetPageId="84" captionText="FIGURES 312320. Clepsydraphes nitidulus (Franz), holotype. Habitus in dorsal view (312); head in anterodorsal (313) and dorsal (314) views; head and prothorax in lateral view (315); antenna (316); aedeagus in ventral (317, 319) and lateral (318, 320) views." figureDoi="http://doi.org/10.5281/zenodo.13746043" httpUri="https://zenodo.org/record/13746043/files/figure.png" pageId="82" pageNumber="83">Fig. 315</figureCitation>
); head (
<figureCitation id="137552C3FFA25052FBB0FD39FB644763" box="[1028,1185,757,784]" captionStart="FIGURES 312320" captionStartId="84.[151,264,1812,1836]" captionTargetBox="[151,1430,365,1776]" captionTargetId="figure-59@84.[151,1435,365,1789]" captionTargetPageId="84" captionText="FIGURES 312320. Clepsydraphes nitidulus (Franz), holotype. Habitus in dorsal view (312); head in anterodorsal (313) and dorsal (314) views; head and prothorax in lateral view (315); antenna (316); aedeagus in ventral (317, 319) and lateral (318, 320) views." figureDoi="http://doi.org/10.5281/zenodo.13746043" httpUri="https://zenodo.org/record/13746043/files/figure.png" pageId="82" pageNumber="83">Figs 313315</figureCitation>
,
<figureCitation id="137552C3FFA25052FB1FFD39FACB477C" box="[1195,1294,757,783]" captionStart="FIGURES 321323" captionStartId="85.[151,264,1018,1042]" captionTargetBox="[151,1435,181,995]" captionTargetId="figure-16@85.[151,1435,181,995]" captionTargetPageId="85" captionText="FIGURES 321323. Clepsydraphes nitidulus (Franz), holotype. Head, prothorax and elytral base in dorsal view (321); head and prothorax in ventral view (322); pterothorax in ventral view (323). Abbreviations: gp, gular plate; hr, hypomeral ridge; hs, hypostomal suture; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746047" httpUri="https://zenodo.org/record/13746047/files/figure.png" pageId="82" pageNumber="83">321322</figureCitation>
) round, with eyes situated closer to mandibular bases and antennal fossae than to occipital constriction; antennal fossae separated at middle by distance subequal to width of scape (
<figureCitation id="137552C3FFA25052FD58FCF1FC974724" box="[748,850,829,855]" captionStart="FIGURES 312320" captionStartId="84.[151,264,1812,1836]" captionTargetBox="[151,1430,365,1776]" captionTargetId="figure-59@84.[151,1435,365,1789]" captionTargetPageId="84" captionText="FIGURES 312320. Clepsydraphes nitidulus (Franz), holotype. Habitus in dorsal view (312); head in anterodorsal (313) and dorsal (314) views; head and prothorax in lateral view (315); antenna (316); aedeagus in ventral (317, 319) and lateral (318, 320) views." figureDoi="http://doi.org/10.5281/zenodo.13746043" httpUri="https://zenodo.org/record/13746043/files/figure.png" pageId="82" pageNumber="83">Fig. 313</figureCitation>
); antenna (
<figureCitation id="137552C3FFA25052FC61FCF1FBFC4724" box="[981,1081,829,855]" captionStart="FIGURES 312320" captionStartId="84.[151,264,1812,1836]" captionTargetBox="[151,1430,365,1776]" captionTargetId="figure-59@84.[151,1435,365,1789]" captionTargetPageId="84" captionText="FIGURES 312320. Clepsydraphes nitidulus (Franz), holotype. Habitus in dorsal view (312); head in anterodorsal (313) and dorsal (314) views; head and prothorax in lateral view (315); antenna (316); aedeagus in ventral (317, 319) and lateral (318, 320) views." figureDoi="http://doi.org/10.5281/zenodo.13746043" httpUri="https://zenodo.org/record/13746043/files/figure.png" pageId="82" pageNumber="83">Fig. 316</figureCitation>
) gradually thickening distally; frontoclypeal groove lacking; posterior margin of vertex (
<figureCitation id="137552C3FFA25052FCF5FCADFC604708" box="[833,933,865,891]" captionStart="FIGURES 312320" captionStartId="84.[151,264,1812,1836]" captionTargetBox="[151,1430,365,1776]" captionTargetId="figure-59@84.[151,1435,365,1789]" captionTargetPageId="84" captionText="FIGURES 312320. Clepsydraphes nitidulus (Franz), holotype. Habitus in dorsal view (312); head in anterodorsal (313) and dorsal (314) views; head and prothorax in lateral view (315); antenna (316); aedeagus in ventral (317, 319) and lateral (318, 320) views." figureDoi="http://doi.org/10.5281/zenodo.13746043" httpUri="https://zenodo.org/record/13746043/files/figure.png" pageId="82" pageNumber="83">Fig. 313</figureCitation>
) rounded and not bulging posterodorsally; submental region (
<figureCitation id="137552C3FFA25052FED8FC49FE1547EC" box="[364,464,901,927]" captionStart="FIGURES 321323" captionStartId="85.[151,264,1018,1042]" captionTargetBox="[151,1435,181,995]" captionTargetId="figure-16@85.[151,1435,181,995]" captionTargetPageId="85" captionText="FIGURES 321323. Clepsydraphes nitidulus (Franz), holotype. Head, prothorax and elytral base in dorsal view (321); head and prothorax in ventral view (322); pterothorax in ventral view (323). Abbreviations: gp, gular plate; hr, hypomeral ridge; hs, hypostomal suture; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746047" httpUri="https://zenodo.org/record/13746047/files/figure.png" pageId="82" pageNumber="83">Fig. 322</figureCitation>
;
<emphasis id="B93A9254FFA25052FE6AFC4AFDCE47EC" box="[478,523,902,927]" italics="true" pageId="82" pageNumber="83">smn</emphasis>
) demarcated laterally by hypostomal sutures (
<figureCitation id="137552C3FFA25052FBA9FC49FB4447EC" box="[1053,1153,901,927]" captionStart="FIGURES 321323" captionStartId="85.[151,264,1018,1042]" captionTargetBox="[151,1435,181,995]" captionTargetId="figure-16@85.[151,1435,181,995]" captionTargetPageId="85" captionText="FIGURES 321323. Clepsydraphes nitidulus (Franz), holotype. Head, prothorax and elytral base in dorsal view (321); head and prothorax in ventral view (322); pterothorax in ventral view (323). Abbreviations: gp, gular plate; hr, hypomeral ridge; hs, hypostomal suture; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746047" httpUri="https://zenodo.org/record/13746047/files/figure.png" pageId="82" pageNumber="83">Fig. 322</figureCitation>
;
<emphasis id="B93A9254FFA25052FB3BFC49FB6D47EC" box="[1167,1192,901,927]" italics="true" pageId="82" pageNumber="83">hs</emphasis>
) strongly converging posteriorly, area between sutures subtrapezoidal; hypostomal ridges (
<figureCitation id="137552C3FFA25052FC01FC65FBDF47B0" box="[949,1050,937,963]" captionStart="FIGURES 321323" captionStartId="85.[151,264,1018,1042]" captionTargetBox="[151,1435,181,995]" captionTargetId="figure-16@85.[151,1435,181,995]" captionTargetPageId="85" captionText="FIGURES 321323. Clepsydraphes nitidulus (Franz), holotype. Head, prothorax and elytral base in dorsal view (321); head and prothorax in ventral view (322); pterothorax in ventral view (323). Abbreviations: gp, gular plate; hr, hypomeral ridge; hs, hypostomal suture; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746047" httpUri="https://zenodo.org/record/13746047/files/figure.png" pageId="82" pageNumber="83">Fig. 322</figureCitation>
;
<emphasis id="B93A9254FFA25052FB9EFC65FB8647B0" box="[1066,1091,937,963]" italics="true" pageId="82" pageNumber="83">hr</emphasis>
) extending posteromesad and connecting to hypostomal sutures at level of posterior tentorial pits; pronotum (
<figureCitation id="137552C3FFA25052FBA1FC01FBB04794" box="[1045,1141,973,999]" captionStart="FIGURES 321323" captionStartId="85.[151,264,1018,1042]" captionTargetBox="[151,1435,181,995]" captionTargetId="figure-16@85.[151,1435,181,995]" captionTargetPageId="85" captionText="FIGURES 321323. Clepsydraphes nitidulus (Franz), holotype. Head, prothorax and elytral base in dorsal view (321); head and prothorax in ventral view (322); pterothorax in ventral view (323). Abbreviations: gp, gular plate; hr, hypomeral ridge; hs, hypostomal suture; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746047" httpUri="https://zenodo.org/record/13746047/files/figure.png" pageId="82" pageNumber="83">Fig. 321</figureCitation>
) oval, broadest in front of middle, lacking antebasal pits, grooves, impressions, and without lateral and sublateral carinae; prosternum (
<figureCitation id="137552C3FFA25052FADBFC3DFF06405C" captionStart="FIGURES 321323" captionStartId="85.[151,264,1018,1042]" captionTargetBox="[151,1435,181,995]" captionTargetId="figure-16@85.[151,1435,181,995]" captionTargetPageId="85" captionText="FIGURES 321323. Clepsydraphes nitidulus (Franz), holotype. Head, prothorax and elytral base in dorsal view (321); head and prothorax in ventral view (322); pterothorax in ventral view (323). Abbreviations: gp, gular plate; hr, hypomeral ridge; hs, hypostomal suture; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746047" httpUri="https://zenodo.org/record/13746047/files/figure.png" pageId="82" pageNumber="83">Fig. 322</figureCitation>
) with precoxal (= basisternal) region vestigial, much shorter than procoxal rests, prosternal process lacking; notosternal sutures (
<figureCitation id="137552C3FFA25052FECBFBF5FE264020" box="[383,483,1081,1107]" captionStart="FIGURES 321323" captionStartId="85.[151,264,1018,1042]" captionTargetBox="[151,1435,181,995]" captionTargetId="figure-16@85.[151,1435,181,995]" captionTargetPageId="85" captionText="FIGURES 321323. Clepsydraphes nitidulus (Franz), holotype. Head, prothorax and elytral base in dorsal view (321); head and prothorax in ventral view (322); pterothorax in ventral view (323). Abbreviations: gp, gular plate; hr, hypomeral ridge; hs, hypostomal suture; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746047" httpUri="https://zenodo.org/record/13746047/files/figure.png" pageId="82" pageNumber="83">Fig. 322</figureCitation>
;
<emphasis id="B93A9254FFA25052FE45FBF6FDD04020" box="[497,533,1082,1107]" italics="true" pageId="82" pageNumber="83">nss</emphasis>
) complete; procoxal cavity closed by postcoxal prosternal lobe overlapped by (but not fused with) postcoxal mesal lobe of hypomeron; hypomeral ridges lacking; mesoventral intercoxal process (
<figureCitation id="137552C3FFA25052FF2BFB4DFEC440E8" box="[159,257,1153,1179]" captionStart="FIGURES 321323" captionStartId="85.[151,264,1018,1042]" captionTargetBox="[151,1435,181,995]" captionTargetId="figure-16@85.[151,1435,181,995]" captionTargetPageId="85" captionText="FIGURES 321323. Clepsydraphes nitidulus (Franz), holotype. Head, prothorax and elytral base in dorsal view (321); head and prothorax in ventral view (322); pterothorax in ventral view (323). Abbreviations: gp, gular plate; hr, hypomeral ridge; hs, hypostomal suture; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746047" httpUri="https://zenodo.org/record/13746047/files/figure.png" pageId="82" pageNumber="83">Fig. 323</figureCitation>
<emphasis id="B93A9254FFA25052FEBFFB4EFE8040E8" box="[267,325,1154,1179]" italics="true" pageId="82" pageNumber="83">msvp</emphasis>
) elongate subtriangular, weakly elevated and posteriorly reducing in height, reaching middle of mesocoxal rests and not separating mesocoxae; metaventrite lacking anterior metaventral process; metaventral intermetacoxal process (
<figureCitation id="137552C3FFA25052FE1DFB05FDCF4090" box="[425,522,1225,1251]" captionStart="FIGURES 321323" captionStartId="85.[151,264,1018,1042]" captionTargetBox="[151,1435,181,995]" captionTargetId="figure-16@85.[151,1435,181,995]" captionTargetPageId="85" captionText="FIGURES 321323. Clepsydraphes nitidulus (Franz), holotype. Head, prothorax and elytral base in dorsal view (321); head and prothorax in ventral view (322); pterothorax in ventral view (323). Abbreviations: gp, gular plate; hr, hypomeral ridge; hs, hypostomal suture; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746047" httpUri="https://zenodo.org/record/13746047/files/figure.png" pageId="82" pageNumber="83">Fig. 323</figureCitation>
;
<emphasis id="B93A9254FFA25052FDA2FB06FD894090" box="[534,588,1226,1251]" italics="true" pageId="82" pageNumber="83">mtvp</emphasis>
) short and subtriangular, divided into lateral halves by deep median fissure and not separating metacoxae; each elytron (
<figureCitation id="137552C3FFA25052FD3BFB21FD2A4174" box="[655,751,1261,1287]" captionStart="FIGURES 321323" captionStartId="85.[151,264,1018,1042]" captionTargetBox="[151,1435,181,995]" captionTargetId="figure-16@85.[151,1435,181,995]" captionTargetPageId="85" captionText="FIGURES 321323. Clepsydraphes nitidulus (Franz), holotype. Head, prothorax and elytral base in dorsal view (321); head and prothorax in ventral view (322); pterothorax in ventral view (323). Abbreviations: gp, gular plate; hr, hypomeral ridge; hs, hypostomal suture; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746047" httpUri="https://zenodo.org/record/13746047/files/figure.png" pageId="82" pageNumber="83">Fig. 321</figureCitation>
) lacking any traces of basal foveae; mesoscutellar shield not exposed between elytral bases (
<figureCitation id="137552C3FFA25052FDB3FADDFDA84158" box="[519,621,1297,1323]" captionStart="FIGURES 321323" captionStartId="85.[151,264,1018,1042]" captionTargetBox="[151,1435,181,995]" captionTargetId="figure-16@85.[151,1435,181,995]" captionTargetPageId="85" captionText="FIGURES 321323. Clepsydraphes nitidulus (Franz), holotype. Head, prothorax and elytral base in dorsal view (321); head and prothorax in ventral view (322); pterothorax in ventral view (323). Abbreviations: gp, gular plate; hr, hypomeral ridge; hs, hypostomal suture; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746047" httpUri="https://zenodo.org/record/13746047/files/figure.png" pageId="82" pageNumber="83">Fig. 321</figureCitation>
); aedeagus (
<figureCitation id="137552C3FFA25052FD49FADDFC584158" box="[765,925,1297,1323]" captionStart="FIGURES 312320" captionStartId="84.[151,264,1812,1836]" captionTargetBox="[151,1430,365,1776]" captionTargetId="figure-59@84.[151,1435,365,1789]" captionTargetPageId="84" captionText="FIGURES 312320. Clepsydraphes nitidulus (Franz), holotype. Habitus in dorsal view (312); head in anterodorsal (313) and dorsal (314) views; head and prothorax in lateral view (315); antenna (316); aedeagus in ventral (317, 319) and lateral (318, 320) views." figureDoi="http://doi.org/10.5281/zenodo.13746043" httpUri="https://zenodo.org/record/13746043/files/figure.png" pageId="82" pageNumber="83">Figs 317320</figureCitation>
) with symmetrical median lobe and weakly sclerotized endophallus, with sclerotized ventral apical wall delimited from proximal area of median lobe by membrane, aedeagus with free, slender parameres.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFA25053FF73FAB1FAB4417E" lastPageId="83" lastPageNumber="84" pageId="82" pageNumber="83" type="description">
<paragraph id="8BF14E46FFA25052FF73FAB1FD3041C8" blockId="82.[151,1437,721,2008]" pageId="82" pageNumber="83">
<emphasis id="B93A9254FFA25052FF73FAB1FE9C41E4" bold="true" box="[199,345,1405,1431]" pageId="82" pageNumber="83">Description.</emphasis>
Body form (
<figureCitation id="137552C3FFA25052FE5FFAB1FD8841E4" box="[491,589,1405,1431]" captionStart="FIGURES 312320" captionStartId="84.[151,264,1812,1836]" captionTargetBox="[151,1430,365,1776]" captionTargetId="figure-59@84.[151,1435,365,1789]" captionTargetPageId="84" captionText="FIGURES 312320. Clepsydraphes nitidulus (Franz), holotype. Habitus in dorsal view (312); head in anterodorsal (313) and dorsal (314) views; head and prothorax in lateral view (315); antenna (316); aedeagus in ventral (317, 319) and lateral (318, 320) views." figureDoi="http://doi.org/10.5281/zenodo.13746043" httpUri="https://zenodo.org/record/13746043/files/figure.png" pageId="82" pageNumber="83">Fig. 312</figureCitation>
) elongate and slender, strongly convex, with distinct constrictions between head and pronotum and between pronotum and elytra.
</paragraph>
<paragraph id="8BF14E46FFA25052FF73FA09FCAA428C" blockId="82.[151,1437,721,2008]" pageId="82" pageNumber="83">
Head (
<figureCitation id="137552C3FFA25052FEA4FA09FE694193" box="[272,428,1477,1504]" captionStart="FIGURES 312320" captionStartId="84.[151,264,1812,1836]" captionTargetBox="[151,1430,365,1776]" captionTargetId="figure-59@84.[151,1435,365,1789]" captionTargetPageId="84" captionText="FIGURES 312320. Clepsydraphes nitidulus (Franz), holotype. Habitus in dorsal view (312); head in anterodorsal (313) and dorsal (314) views; head and prothorax in lateral view (315); antenna (316); aedeagus in ventral (317, 319) and lateral (318, 320) views." figureDoi="http://doi.org/10.5281/zenodo.13746043" httpUri="https://zenodo.org/record/13746043/files/figure.png" pageId="82" pageNumber="83">Figs 313315</figureCitation>
,
<figureCitation id="137552C3FFA25052FE02FA09FDDC41AC" box="[438,537,1477,1503]" captionStart="FIGURES 321323" captionStartId="85.[151,264,1018,1042]" captionTargetBox="[151,1435,181,995]" captionTargetId="figure-16@85.[151,1435,181,995]" captionTargetPageId="85" captionText="FIGURES 321323. Clepsydraphes nitidulus (Franz), holotype. Head, prothorax and elytral base in dorsal view (321); head and prothorax in ventral view (322); pterothorax in ventral view (323). Abbreviations: gp, gular plate; hr, hypomeral ridge; hs, hypostomal suture; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746047" httpUri="https://zenodo.org/record/13746047/files/figure.png" pageId="82" pageNumber="83">321322</figureCitation>
) rounded; compound eyes situated much closer to mandibular bases and antennal fossae than to occipital constriction, each with posterior emargination (
<figureCitation id="137552C3FFA25052FC73FA25FBEE4277" box="[967,1067,1513,1540]" captionStart="FIGURES 312320" captionStartId="84.[151,264,1812,1836]" captionTargetBox="[151,1430,365,1776]" captionTargetId="figure-59@84.[151,1435,365,1789]" captionTargetPageId="84" captionText="FIGURES 312320. Clepsydraphes nitidulus (Franz), holotype. Habitus in dorsal view (312); head in anterodorsal (313) and dorsal (314) views; head and prothorax in lateral view (315); antenna (316); aedeagus in ventral (317, 319) and lateral (318, 320) views." figureDoi="http://doi.org/10.5281/zenodo.13746043" httpUri="https://zenodo.org/record/13746043/files/figure.png" pageId="82" pageNumber="83">Fig. 315</figureCitation>
); tempora rounded; vertex (
<figureCitation id="137552C3FFA25052FADEFA25FF3E425B" captionStart="FIGURES 312320" captionStartId="84.[151,264,1812,1836]" captionTargetBox="[151,1430,365,1776]" captionTargetId="figure-59@84.[151,1435,365,1789]" captionTargetPageId="84" captionText="FIGURES 312320. Clepsydraphes nitidulus (Franz), holotype. Habitus in dorsal view (312); head in anterodorsal (313) and dorsal (314) views; head and prothorax in lateral view (315); antenna (316); aedeagus in ventral (317, 319) and lateral (318, 320) views." figureDoi="http://doi.org/10.5281/zenodo.13746043" httpUri="https://zenodo.org/record/13746043/files/figure.png" pageId="82" pageNumber="83">Figs 314315</figureCitation>
) not bulging posterodorsally, confluent with frons, its posterior margin rounded. Bristles present on genae (
<figureCitation id="137552C3FFA25052FF2BF9FDFECC423F" box="[159,265,1585,1612]" captionStart="FIGURES 312320" captionStartId="84.[151,264,1812,1836]" captionTargetBox="[151,1430,365,1776]" captionTargetId="figure-59@84.[151,1435,365,1789]" captionTargetPageId="84" captionText="FIGURES 312320. Clepsydraphes nitidulus (Franz), holotype. Habitus in dorsal view (312); head in anterodorsal (313) and dorsal (314) views; head and prothorax in lateral view (315); antenna (316); aedeagus in ventral (317, 319) and lateral (318, 320) views." figureDoi="http://doi.org/10.5281/zenodo.13746043" httpUri="https://zenodo.org/record/13746043/files/figure.png" pageId="82" pageNumber="83">Figs 315</figureCitation>
,
<figureCitation id="137552C3FFA25052FEA3F9FDFE804238" box="[279,325,1585,1611]" captionStart="FIGURES 321323" captionStartId="85.[151,264,1018,1042]" captionTargetBox="[151,1435,181,995]" captionTargetId="figure-16@85.[151,1435,181,995]" captionTargetPageId="85" captionText="FIGURES 321323. Clepsydraphes nitidulus (Franz), holotype. Head, prothorax and elytral base in dorsal view (321); head and prothorax in ventral view (322); pterothorax in ventral view (323). Abbreviations: gp, gular plate; hr, hypomeral ridge; hs, hypostomal suture; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746047" httpUri="https://zenodo.org/record/13746047/files/figure.png" pageId="82" pageNumber="83">322</figureCitation>
). Occipital constriction (
<figureCitation id="137552C3FFA25052FDD1F9FDFD0C4238" box="[613,713,1585,1611]" captionStart="FIGURES 321323" captionStartId="85.[151,264,1018,1042]" captionTargetBox="[151,1435,181,995]" captionTargetId="figure-16@85.[151,1435,181,995]" captionTargetPageId="85" captionText="FIGURES 321323. Clepsydraphes nitidulus (Franz), holotype. Head, prothorax and elytral base in dorsal view (321); head and prothorax in ventral view (322); pterothorax in ventral view (323). Abbreviations: gp, gular plate; hr, hypomeral ridge; hs, hypostomal suture; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746047" httpUri="https://zenodo.org/record/13746047/files/figure.png" pageId="82" pageNumber="83">Fig. 322</figureCitation>
) about as broad as half width of head; neck region strongly broadening posterad. Antennal fossae large and situated anterolaterally, separated at middle by median longitudinal frontal area about as wide as width of scape (
<figureCitation id="137552C3FFA25052FD00F9B5FCDF42E0" box="[692,794,1657,1683]" captionStart="FIGURES 312320" captionStartId="84.[151,264,1812,1836]" captionTargetBox="[151,1430,365,1776]" captionTargetId="figure-59@84.[151,1435,365,1789]" captionTargetPageId="84" captionText="FIGURES 312320. Clepsydraphes nitidulus (Franz), holotype. Habitus in dorsal view (312); head in anterodorsal (313) and dorsal (314) views; head and prothorax in lateral view (315); antenna (316); aedeagus in ventral (317, 319) and lateral (318, 320) views." figureDoi="http://doi.org/10.5281/zenodo.13746043" httpUri="https://zenodo.org/record/13746043/files/figure.png" pageId="82" pageNumber="83">Fig. 313</figureCitation>
). Frontoclypeal groove lacking; clypeus short, rounded anteriorly, with lateral margins converging towards labrum. Gular plate (
<figureCitation id="137552C3FFA25052FC76F951FBE642C4" box="[962,1059,1693,1719]" captionStart="FIGURES 321323" captionStartId="85.[151,264,1018,1042]" captionTargetBox="[151,1435,181,995]" captionTargetId="figure-16@85.[151,1435,181,995]" captionTargetPageId="85" captionText="FIGURES 321323. Clepsydraphes nitidulus (Franz), holotype. Head, prothorax and elytral base in dorsal view (321); head and prothorax in ventral view (322); pterothorax in ventral view (323). Abbreviations: gp, gular plate; hr, hypomeral ridge; hs, hypostomal suture; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746047" httpUri="https://zenodo.org/record/13746047/files/figure.png" pageId="82" pageNumber="83">Fig. 322</figureCitation>
;
<emphasis id="B93A9254FFA25052FB9AF952FB8F42C4" box="[1070,1098,1694,1719]" italics="true" pageId="82" pageNumber="83">gp</emphasis>
) subtrapezoidal, covered with transverse microsculpture, with indistinct gular sutures; posterior tentorial pits (
<figureCitation id="137552C3FFA25052FBA4F90DFBB442A8" box="[1040,1137,1729,1755]" captionStart="FIGURES 321323" captionStartId="85.[151,264,1018,1042]" captionTargetBox="[151,1435,181,995]" captionTargetId="figure-16@85.[151,1435,181,995]" captionTargetPageId="85" captionText="FIGURES 321323. Clepsydraphes nitidulus (Franz), holotype. Head, prothorax and elytral base in dorsal view (321); head and prothorax in ventral view (322); pterothorax in ventral view (323). Abbreviations: gp, gular plate; hr, hypomeral ridge; hs, hypostomal suture; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746047" httpUri="https://zenodo.org/record/13746047/files/figure.png" pageId="82" pageNumber="83">Fig. 322</figureCitation>
;
<emphasis id="B93A9254FFA25052FBC9F90EFB6442A8" box="[1149,1185,1730,1755]" italics="true" pageId="82" pageNumber="83">ptp</emphasis>
) situated in transverse impression demarcating ventrally 'neck' region, each punctiform.
</paragraph>
<paragraph id="8BF14E46FFA25053FF73F8C5FE974532" blockId="82.[151,1437,721,2008]" lastBlockId="83.[151,1437,151,1869]" lastPageId="83" lastPageNumber="84" pageId="82" pageNumber="83">
Submentum (
<figureCitation id="137552C3FFA25052FEE8F8C5FE7E4350" box="[348,443,1801,1827]" captionStart="FIGURES 321323" captionStartId="85.[151,264,1018,1042]" captionTargetBox="[151,1435,181,995]" captionTargetId="figure-16@85.[151,1435,181,995]" captionTargetPageId="85" captionText="FIGURES 321323. Clepsydraphes nitidulus (Franz), holotype. Head, prothorax and elytral base in dorsal view (321); head and prothorax in ventral view (322); pterothorax in ventral view (323). Abbreviations: gp, gular plate; hr, hypomeral ridge; hs, hypostomal suture; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746047" httpUri="https://zenodo.org/record/13746047/files/figure.png" pageId="82" pageNumber="83">Fig. 322</figureCitation>
;
<emphasis id="B93A9254FFA25052FE71F8C6FE374350" box="[453,498,1802,1827]" italics="true" pageId="82" pageNumber="83">smn</emphasis>
) subtrapezoidal, laterally demarcated by complete hypostomal sutures (
<figureCitation id="137552C3FFA25052FABDF8C5FAAD4350" box="[1289,1384,1801,1827]" captionStart="FIGURES 321323" captionStartId="85.[151,264,1018,1042]" captionTargetBox="[151,1435,181,995]" captionTargetId="figure-16@85.[151,1435,181,995]" captionTargetPageId="85" captionText="FIGURES 321323. Clepsydraphes nitidulus (Franz), holotype. Head, prothorax and elytral base in dorsal view (321); head and prothorax in ventral view (322); pterothorax in ventral view (323). Abbreviations: gp, gular plate; hr, hypomeral ridge; hs, hypostomal suture; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746047" httpUri="https://zenodo.org/record/13746047/files/figure.png" pageId="82" pageNumber="83">Fig. 322</figureCitation>
;
<emphasis id="B93A9254FFA25052FAC6F8C5FA4E4350" box="[1394,1419,1801,1827]" italics="true" pageId="82" pageNumber="83">hs</emphasis>
); mentum subtrapezoidal, broadest near base, with bisinuate anterior margin; prelabium short with moderately broadly separated insertions of labial palps; labial palpomere 1 annulate, palpomere 2 strongly elongate, subcylindrical, nearly 4 times as long as broad, palpomere 3 narrow, elongate but distinctly shorter than 2, tapering distally and straight. Maxilla generalized as in most Stenichnini, with large semicircular cardo, subtriangular basistipes, elongate mediostipes, elongate lacinia and galea, each with thick and not very numerous setae along distomesal margin, palpifer elongate, maxillary palp with distinctly elongate and subcylindrical palpomere 1, long, pipe-shaped and weakly curved palpomere 2, strongly elongate and slender fusiform palpomere 3 about twice as wide as palpomere 2, and minute, strongly elongate subconical and pointed palpomere 4. Mandibles poorly visible in studied specimen. Labrum strongly transverse, with rounded anterior margin, and with sparse, symmetrically distributed dorsal setae directed anterad.
</paragraph>
<paragraph id="8BF14E46FFA35053FF73FE87FB5D45FA" blockId="83.[151,1437,151,1869]" pageId="83" pageNumber="84">
Antenna (
<figureCitation id="137552C3FFA35053FE80FE87FE564516" box="[308,403,331,357]" captionStart="FIGURES 312320" captionStartId="84.[151,264,1812,1836]" captionTargetBox="[151,1430,365,1776]" captionTargetId="figure-59@84.[151,1435,365,1789]" captionTargetPageId="84" captionText="FIGURES 312320. Clepsydraphes nitidulus (Franz), holotype. Habitus in dorsal view (312); head in anterodorsal (313) and dorsal (314) views; head and prothorax in lateral view (315); antenna (316); aedeagus in ventral (317, 319) and lateral (318, 320) views." figureDoi="http://doi.org/10.5281/zenodo.13746043" httpUri="https://zenodo.org/record/13746043/files/figure.png" pageId="83" pageNumber="84">Fig. 316</figureCitation>
) short in relation to body length, slender, with scape and pedicel each elongate; flagellomeres loosely assembled; antenna gradually thickening distally. All antennomeres sparsely setose.
</paragraph>
<paragraph id="8BF14E46FFA35053FF73FE5FFE7D46DA" blockId="83.[151,1437,151,1869]" pageId="83" pageNumber="84">
Prothorax in dorsal view (
<figureCitation id="137552C3FFA35053FE47FE5FFD9145DE" box="[499,596,403,429]" captionStart="FIGURES 321323" captionStartId="85.[151,264,1018,1042]" captionTargetBox="[151,1435,181,995]" captionTargetId="figure-16@85.[151,1435,181,995]" captionTargetPageId="85" captionText="FIGURES 321323. Clepsydraphes nitidulus (Franz), holotype. Head, prothorax and elytral base in dorsal view (321); head and prothorax in ventral view (322); pterothorax in ventral view (323). Abbreviations: gp, gular plate; hr, hypomeral ridge; hs, hypostomal suture; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746047" httpUri="https://zenodo.org/record/13746047/files/figure.png" pageId="83" pageNumber="84">Fig. 321</figureCitation>
) oval, broadest distinctly in front of middle. Anterior and lateral margins confluent, without anterior angles; lateral margins strongly rounded; posterior corners not marked; posterior margin convex posteriorly. Pronotum lacking antebasal pits, impressions, sublateral and lateral carinae. Bristles present only on posterior regions of outer hypomera (
<figureCitation id="137552C3FFA35053FD12FE33FCCF466A" box="[678,778,511,537]" captionStart="FIGURES 321323" captionStartId="85.[151,264,1018,1042]" captionTargetBox="[151,1435,181,995]" captionTargetId="figure-16@85.[151,1435,181,995]" captionTargetPageId="85" captionText="FIGURES 321323. Clepsydraphes nitidulus (Franz), holotype. Head, prothorax and elytral base in dorsal view (321); head and prothorax in ventral view (322); pterothorax in ventral view (323). Abbreviations: gp, gular plate; hr, hypomeral ridge; hs, hypostomal suture; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746047" httpUri="https://zenodo.org/record/13746047/files/figure.png" pageId="83" pageNumber="84">Fig. 322</figureCitation>
). Hypomeral ridges lacking. Prosternum (
<figureCitation id="137552C3FFA35053FB5FFE33FA88466A" box="[1259,1357,511,537]" captionStart="FIGURES 321323" captionStartId="85.[151,264,1018,1042]" captionTargetBox="[151,1435,181,995]" captionTargetId="figure-16@85.[151,1435,181,995]" captionTargetPageId="85" captionText="FIGURES 321323. Clepsydraphes nitidulus (Franz), holotype. Head, prothorax and elytral base in dorsal view (321); head and prothorax in ventral view (322); pterothorax in ventral view (323). Abbreviations: gp, gular plate; hr, hypomeral ridge; hs, hypostomal suture; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746047" httpUri="https://zenodo.org/record/13746047/files/figure.png" pageId="83" pageNumber="84">Fig. 322</figureCitation>
) much shorter than half length of prothorax, with precoxal region vestigial, much shorter than coxal rests; notosternal sutures (
<figureCitation id="137552C3FFA35053FF42FD8BFE9D4612" box="[246,344,583,609]" captionStart="FIGURES 321323" captionStartId="85.[151,264,1018,1042]" captionTargetBox="[151,1435,181,995]" captionTargetId="figure-16@85.[151,1435,181,995]" captionTargetPageId="85" captionText="FIGURES 321323. Clepsydraphes nitidulus (Franz), holotype. Head, prothorax and elytral base in dorsal view (321); head and prothorax in ventral view (322); pterothorax in ventral view (323). Abbreviations: gp, gular plate; hr, hypomeral ridge; hs, hypostomal suture; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746047" httpUri="https://zenodo.org/record/13746047/files/figure.png" pageId="83" pageNumber="84">Fig. 322</figureCitation>
;
<emphasis id="B93A9254FFA35053FED0FD8BFE4D4613" box="[356,392,583,608]" italics="true" pageId="83" pageNumber="84">nss</emphasis>
) complete. Prosternal process not developed; posterior margin of furcasternum with shallow arcuate median emargination. Procoxal cavities closed; postcoxal hypomeral lobe overlapping (but not fused to) postcoxal prosternal lobe.
</paragraph>
<paragraph id="8BF14E46FFA35053FF73FD7FFB0146BE" blockId="83.[151,1437,151,1869]" box="[199,1220,691,717]" pageId="83" pageNumber="84">Meso- and metanotum not studied; mesoscutellar shield not exposed between elytral bases.</paragraph>
<paragraph id="8BF14E46FFA35053FF73FD1BFCA3472E" blockId="83.[151,1437,151,1869]" pageId="83" pageNumber="84">
Mesoventrite (
<figureCitation id="137552C3FFA35053FEDEFD1BFE0F4682" box="[362,458,727,753]" captionStart="FIGURES 321323" captionStartId="85.[151,264,1018,1042]" captionTargetBox="[151,1435,181,995]" captionTargetId="figure-16@85.[151,1435,181,995]" captionTargetPageId="85" captionText="FIGURES 321323. Clepsydraphes nitidulus (Franz), holotype. Head, prothorax and elytral base in dorsal view (321); head and prothorax in ventral view (322); pterothorax in ventral view (323). Abbreviations: gp, gular plate; hr, hypomeral ridge; hs, hypostomal suture; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746047" httpUri="https://zenodo.org/record/13746047/files/figure.png" pageId="83" pageNumber="84">Fig. 323</figureCitation>
) transverse, with massive prepectus, subquadrate and setose anteromedian impressions functioning as procoxal rests (
<figureCitation id="137552C3FFA35053FE5CFD37FD8D4766" box="[488,584,763,789]" captionStart="FIGURES 321323" captionStartId="85.[151,264,1018,1042]" captionTargetBox="[151,1435,181,995]" captionTargetId="figure-16@85.[151,1435,181,995]" captionTargetPageId="85" captionText="FIGURES 321323. Clepsydraphes nitidulus (Franz), holotype. Head, prothorax and elytral base in dorsal view (321); head and prothorax in ventral view (322); pterothorax in ventral view (323). Abbreviations: gp, gular plate; hr, hypomeral ridge; hs, hypostomal suture; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746047" httpUri="https://zenodo.org/record/13746047/files/figure.png" pageId="83" pageNumber="84">Fig. 323</figureCitation>
;
<emphasis id="B93A9254FFA35053FDE0FD37FDBC4767" box="[596,633,763,788]" italics="true" pageId="83" pageNumber="84">pcr</emphasis>
) separated at middle by longitudinal carina that posteriorly extends into narrow, subtriangular and weakly elevated mesoventral intercoxal process (
<figureCitation id="137552C3FFA35053FC4DFCD3FB98474A" box="[1017,1117,799,825]" captionStart="FIGURES 321323" captionStartId="85.[151,264,1018,1042]" captionTargetBox="[151,1435,181,995]" captionTargetId="figure-16@85.[151,1435,181,995]" captionTargetPageId="85" captionText="FIGURES 321323. Clepsydraphes nitidulus (Franz), holotype. Head, prothorax and elytral base in dorsal view (321); head and prothorax in ventral view (322); pterothorax in ventral view (323). Abbreviations: gp, gular plate; hr, hypomeral ridge; hs, hypostomal suture; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746047" httpUri="https://zenodo.org/record/13746047/files/figure.png" pageId="83" pageNumber="84">Fig. 323</figureCitation>
;
<emphasis id="B93A9254FFA35053FBD8FCD3FB63474B" box="[1132,1190,799,824]" italics="true" pageId="83" pageNumber="84">msvp</emphasis>
) which is posteriorly reducing in height and reaching only middle of mesocoxal rests.
</paragraph>
<paragraph id="8BF14E46FFA35053FF73FCABFE6D479E" blockId="83.[151,1437,151,1869]" pageId="83" pageNumber="84">
Metaventrite (
<figureCitation id="137552C3FFA35053FED0FCABFE0447F2" box="[356,449,871,897]" captionStart="FIGURES 321323" captionStartId="85.[151,264,1018,1042]" captionTargetBox="[151,1435,181,995]" captionTargetId="figure-16@85.[151,1435,181,995]" captionTargetPageId="85" captionText="FIGURES 321323. Clepsydraphes nitidulus (Franz), holotype. Head, prothorax and elytral base in dorsal view (321); head and prothorax in ventral view (322); pterothorax in ventral view (323). Abbreviations: gp, gular plate; hr, hypomeral ridge; hs, hypostomal suture; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746047" httpUri="https://zenodo.org/record/13746047/files/figure.png" pageId="83" pageNumber="84">Fig. 323</figureCitation>
) subtrapezoidal, anteriorly fused with mesoventrite, lacking anterior metaventral process; metaventral intermetacoxal process (
<figureCitation id="137552C3FFA35053FD82FC47FD5C47D6" box="[566,665,907,933]" captionStart="FIGURES 321323" captionStartId="85.[151,264,1018,1042]" captionTargetBox="[151,1435,181,995]" captionTargetId="figure-16@85.[151,1435,181,995]" captionTargetPageId="85" captionText="FIGURES 321323. Clepsydraphes nitidulus (Franz), holotype. Head, prothorax and elytral base in dorsal view (321); head and prothorax in ventral view (322); pterothorax in ventral view (323). Abbreviations: gp, gular plate; hr, hypomeral ridge; hs, hypostomal suture; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746047" httpUri="https://zenodo.org/record/13746047/files/figure.png" pageId="83" pageNumber="84">Fig. 323</figureCitation>
;
<emphasis id="B93A9254FFA35053FD12FC47FD1947D7" box="[678,732,907,932]" italics="true" pageId="83" pageNumber="84">mtvp</emphasis>
) short and broad, triangular and divided into lateral halves by deep median fissure, not separating metacoxae. Metanepisterna and metepimera fully demarcated and narrow, partly exposed in ventral view.
</paragraph>
<paragraph id="8BF14E46FFA35053FF73FC3BFDA34062" blockId="83.[151,1437,151,1869]" box="[199,614,1015,1041]" pageId="83" pageNumber="84">Legs short, lacking peculiar features.</paragraph>
<paragraph id="8BF14E46FFA35053FF73FBD7FF2C402A" blockId="83.[151,1437,151,1869]" pageId="83" pageNumber="84">
Elytra (
<figureCitation id="137552C3FFA35053FEAAFBD7FE424046" box="[286,391,1051,1077]" captionStart="FIGURES 312320" captionStartId="84.[151,264,1812,1836]" captionTargetBox="[151,1430,365,1776]" captionTargetId="figure-59@84.[151,1435,365,1789]" captionTargetPageId="84" captionText="FIGURES 312320. Clepsydraphes nitidulus (Franz), holotype. Habitus in dorsal view (312); head in anterodorsal (313) and dorsal (314) views; head and prothorax in lateral view (315); antenna (316); aedeagus in ventral (317, 319) and lateral (318, 320) views." figureDoi="http://doi.org/10.5281/zenodo.13746043" httpUri="https://zenodo.org/record/13746043/files/figure.png" pageId="83" pageNumber="84">Figs 312</figureCitation>
,
<figureCitation id="137552C3FFA35053FE22FBD7FE044046" box="[406,449,1051,1077]" captionStart="FIGURES 321323" captionStartId="85.[151,264,1018,1042]" captionTargetBox="[151,1435,181,995]" captionTargetId="figure-16@85.[151,1435,181,995]" captionTargetPageId="85" captionText="FIGURES 321323. Clepsydraphes nitidulus (Franz), holotype. Head, prothorax and elytral base in dorsal view (321); head and prothorax in ventral view (322); pterothorax in ventral view (323). Abbreviations: gp, gular plate; hr, hypomeral ridge; hs, hypostomal suture; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746047" httpUri="https://zenodo.org/record/13746047/files/figure.png" pageId="83" pageNumber="84">321</figureCitation>
) together oval, lacking basal impressions, humeral calli, humeral denticles and basal foveae.
</paragraph>
<paragraph id="8BF14E46FFA35053FF73FBAFFCF2400E" blockId="83.[151,1437,151,1869]" box="[199,823,1123,1149]" pageId="83" pageNumber="84">Hind wings in the only known species entirely reduced.</paragraph>
<paragraph id="8BF14E46FFA35053FF73FB4BFB4640B6" blockId="83.[151,1437,151,1869]" pageId="83" pageNumber="84">Abdomen subtriangular, slightly shorter than metaventrite, with six exposed sternites (sternites IIIVIII), division between VII and VIII slightly less distinct than between the remaining segments.</paragraph>
<paragraph id="8BF14E46FFA35053FF73FB03FAB4417E" blockId="83.[151,1437,151,1869]" pageId="83" pageNumber="84">
Aedeagus (
<figureCitation id="137552C3FFA35053FEFCFB03FE2C409A" box="[328,489,1231,1257]" captionStart="FIGURES 312320" captionStartId="84.[151,264,1812,1836]" captionTargetBox="[151,1430,365,1776]" captionTargetId="figure-59@84.[151,1435,365,1789]" captionTargetPageId="84" captionText="FIGURES 312320. Clepsydraphes nitidulus (Franz), holotype. Habitus in dorsal view (312); head in anterodorsal (313) and dorsal (314) views; head and prothorax in lateral view (315); antenna (316); aedeagus in ventral (317, 319) and lateral (318, 320) views." figureDoi="http://doi.org/10.5281/zenodo.13746043" httpUri="https://zenodo.org/record/13746043/files/figure.png" pageId="83" pageNumber="84">Figs 317320</figureCitation>
) with symmetrical median lobe and weakly sclerotized endophallus, with ventral apical wall delimited from proximal area of median lobe by membrane, aedeagus with free, slender parameres.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFA35053FF73FADBFD9941EE" pageId="83" pageNumber="84" type="etymology">
<paragraph id="8BF14E46FFA35053FF73FADBFC0D4126" blockId="83.[151,1437,151,1869]" pageId="83" pageNumber="84">
<emphasis id="B93A9254FFA35053FF73FADBFE8B4142" bold="true" box="[199,334,1303,1329]" pageId="83" pageNumber="84">Etymology.</emphasis>
<taxonomicName id="4C4E35C5FFA35053FEE2FADBFE394142" authorityName="Jałoszyński" authorityYear="2024" box="[342,508,1303,1329]" class="Insecta" family="Staphylinidae" genus="Clepsydraphes" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B93A9254FFA35053FEE2FADBFE394142" box="[342,508,1303,1329]" italics="true" pageId="83" pageNumber="84">Clepsydraphes</emphasis>
</taxonomicName>
combines
<emphasis id="B93A9254FFA35053FDCDFADBFD234142" box="[633,742,1303,1329]" italics="true" pageId="83" pageNumber="84">clepsydra</emphasis>
(water clock) alluding to the body shape in dorsal view, and
<emphasis id="B93A9254FFA35053FF23FAF7FF294126" box="[151,236,1339,1365]" italics="true" pageId="83" pageNumber="84">-raphes</emphasis>
used in generic names of several Stenichnini. Gender masculine.
</paragraph>
<paragraph id="8BF14E46FFA35053FF73FA92FD9941EE" blockId="83.[151,1437,151,1869]" pageId="83" pageNumber="84">
<emphasis id="B93A9254FFA35053FF73FA92FDFE410B" bold="true" box="[199,571,1374,1401]" pageId="83" pageNumber="84">Composition and distribution.</emphasis>
<taxonomicName id="4C4E35C5FFA35053FDFCFA93FD2B410A" authorityName="Jałoszyński" authorityYear="2024" box="[584,750,1375,1401]" class="Insecta" family="Staphylinidae" genus="Clepsydraphes" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B93A9254FFA35053FDFCFA93FD2B410A" box="[584,750,1375,1401]" italics="true" pageId="83" pageNumber="84">Clepsydraphes</emphasis>
</taxonomicName>
includes one nominal species known to occur in Central
<collectingCountry id="F3590ED6FFA35053FF23FA4FFEDB41EE" box="[151,286,1411,1437]" name="Madagascar" pageId="83" pageNumber="84">Madagascar</collectingCountry>
(Alaotra-Mangoro Region).
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFA35053FF73FA6BFDBA433E" pageId="83" pageNumber="84" type="discussion">
<paragraph id="8BF14E46FFA35053FF73FA6BFDBA433E" blockId="83.[151,1437,151,1869]" pageId="83" pageNumber="84">
<emphasis id="B93A9254FFA35053FF73FA6BFEFE41B2" bold="true" box="[199,315,1447,1473]" pageId="83" pageNumber="84">Remarks.</emphasis>
<taxonomicName id="4C4E35C5FFA35053FEF6FA6BFE2D41B2" authorityName="Jałoszyński" authorityYear="2024" box="[322,488,1447,1473]" class="Insecta" family="Staphylinidae" genus="Clepsydraphes" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B93A9254FFA35053FEF6FA6BFE2D41B2" box="[322,488,1447,1473]" italics="true" pageId="83" pageNumber="84">Clepsydraphes</emphasis>
</taxonomicName>
was included in the key to world genera of Stenichnini (
<bibRefCitation id="EFDF33B7FFA35053FBD5FA6BFAE041B2" author="Jaloszynski, P." box="[1121,1317,1447,1473]" pageId="83" pageNumber="84" pagination="549 - 570" refId="ref55749" refString="Jaloszynski, P. (2024) Palposcydmus gen. n. from India, with a preliminary key to world genera of Stenichnini (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 5428 (4), 549 - 570. https: // doi. org / 10.11646 / zootaxa. 5428.4.5" type="journal article" year="2024">Jałoszyński 2024</bibRefCitation>
) under the code name Mdg genus 1. A combination of gradually thickened antennae, anteriorly situated eyes, narrow occipital constriction, hypostomal sutures demarcating elongate subtrapezoidal submentum, oval pronotum broadest in front of middle, complete notosternal sutures, lack of hypomeral ridges, subtriangular mesoventral process posteriorly reaching middle of mesocoxal rests, and contiguous metacoxae distinguish
<taxonomicName id="4C4E35C5FFA35053FBB0F9FBFB6F4222" authorityName="Jałoszyński" authorityYear="2024" box="[1028,1194,1591,1617]" class="Insecta" family="Staphylinidae" genus="Clepsydraphes" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B93A9254FFA35053FBB0F9FBFB6F4222" box="[1028,1194,1591,1617]" italics="true" pageId="83" pageNumber="84">Clepsydraphes</emphasis>
</taxonomicName>
from all stenichnine genera. In dorsal view,
<taxonomicName id="4C4E35C5FFA35053FE2BF997FD804206" authorityName="Jałoszyński" authorityYear="2024" box="[415,581,1627,1653]" class="Insecta" family="Staphylinidae" genus="Clepsydraphes" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B93A9254FFA35053FE2BF997FD804206" box="[415,581,1627,1653]" italics="true" pageId="83" pageNumber="84">Clepsydraphes</emphasis>
</taxonomicName>
resembles mostly the New Guinean
<taxonomicName id="4C4E35C5FFA35053FC51F997FA9E4206" authority="Jaloszynski, 2020 c" authorityName="Jaloszynski" authorityYear="2020" box="[997,1371,1627,1653]" class="Insecta" family="Staphylinidae" genus="Aenigmaphes" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B93A9254FFA35053FC51F997FBB94206" box="[997,1148,1627,1653]" italics="true" pageId="83" pageNumber="84">Aenigmaphes</emphasis>
<bibRefCitation id="EFDF33B7FFA35053FB37F997FA9E4206" author="Jaloszynski, P." box="[1155,1371,1627,1653]" pageId="83" pageNumber="84" pagination="4439 - 446" refId="ref55453" refString="Jaloszynski, P. (2020 c) Aenigmaphes gen. n., a new glandulariine genus of the Australian region (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 4731 (3), 4439 - 446. https: // doi. org / 10.11646 / zootaxa. 4731.3.12" type="journal article" year="2020">Jałoszyński, 2020c</bibRefCitation>
</taxonomicName>
. Both genera have adults showing a similar body shape, with round pronota setose only in posterolateral regions and elytra broadest far behind middle, strongly narrowing anteriorly. However,
<taxonomicName id="4C4E35C5FFA35053FC01F96FFB8942CE" authorityName="Jaloszynski" authorityYear="2020" box="[949,1100,1699,1725]" class="Insecta" family="Staphylinidae" genus="Aenigmaphes" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B93A9254FFA35053FC01F96FFB8942CE" box="[949,1100,1699,1725]" italics="true" pageId="83" pageNumber="84">Aenigmaphes</emphasis>
</taxonomicName>
has dimerous antennal clubs (antennae gradually thickened in
<taxonomicName id="4C4E35C5FFA35053FDACF90BFD7B4292" authorityName="Jałoszyński" authorityYear="2024" box="[536,702,1735,1761]" class="Insecta" family="Staphylinidae" genus="Clepsydraphes" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B93A9254FFA35053FDACF90BFD7B4292" box="[536,702,1735,1761]" italics="true" pageId="83" pageNumber="84">Clepsydraphes</emphasis>
</taxonomicName>
), one median antebasal pronotal pit anteriorly prolonged into a median longitudinal groove (pronotum lacking any pits or grooves in
<taxonomicName id="4C4E35C5FFA35053FC2BF927FB804376" authorityName="Jałoszyński" authorityYear="2024" box="[927,1093,1771,1797]" class="Insecta" family="Staphylinidae" genus="Clepsydraphes" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B93A9254FFA35053FC2BF927FB804376" box="[927,1093,1771,1797]" italics="true" pageId="83" pageNumber="84">Clepsydraphes</emphasis>
</taxonomicName>
), the head lacking hypostomal sutures (present in
<taxonomicName id="4C4E35C5FFA35053FEDFF8C3FDD4435A" authorityName="Jałoszyński" authorityYear="2024" box="[363,529,1807,1833]" class="Insecta" family="Staphylinidae" genus="Clepsydraphes" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B93A9254FFA35053FEDFF8C3FDD4435A" box="[363,529,1807,1833]" italics="true" pageId="83" pageNumber="84">Clepsydraphes</emphasis>
</taxonomicName>
), and a long keel-like mesoventral intermesocoxal process posteriorly fused with the metaventrite (absent in
<taxonomicName id="4C4E35C5FFA35053FE7CF8FFFDAB433E" authorityName="Jałoszyński" authorityYear="2024" box="[456,622,1843,1869]" class="Insecta" family="Staphylinidae" genus="Clepsydraphes" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B93A9254FFA35053FE7CF8FFFDAB433E" box="[456,622,1843,1869]" italics="true" pageId="83" pageNumber="84">Clepsydraphes</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,282 @@
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<mods:title id="07974A17206D87A2E7541EBC84601106">The curse of Horaeomorphus (almost) lifted. Revision of misplaced species from Madagascar, Comoros and Mascarenes (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<subSubSection id="C3541DCDFFA65056FF23FD7FFE6946BE" box="[151,428,691,717]" pageId="86" pageNumber="87" type="nomenclature">
<paragraph id="8BF14E46FFA65056FF23FD7FFE6946BE" blockId="86.[151,428,691,717]" box="[151,428,691,717]" pageId="86" pageNumber="87">
<heading id="D0B9F92AFFA65056FF23FD7FFE6946BE" bold="true" box="[151,428,691,717]" fontSize="11" level="2" pageId="86" pageNumber="87" reason="4">
<emphasis id="B93A9254FFA65056FF23FD7FFE6946BE" bold="true" box="[151,428,691,717]" pageId="86" pageNumber="87">
<taxonomicName id="4C4E35C5FFA65056FF23FD7FFE9746BE" authorityName="Jałoszyński" authorityYear="2024" box="[151,338,691,717]" class="Insecta" family="Staphylinidae" genus="Contreuconnus" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B93A9254FFA65056FF23FD7FFE9746BE" bold="true" box="[151,338,691,717]" italics="true" pageId="86" pageNumber="87">Contreuconnus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2092F2FFFA65056FEEEFD7FFE6946BE" box="[346,428,691,717]" pageId="86" pageNumber="87" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFA65056FF23FD34FC15477C" box="[151,976,759,784]" pageId="86" pageNumber="87" type="materials_examined">
<paragraph id="8BF14E46FFA65056FF23FD34FC15477C" blockId="86.[151,976,759,784]" box="[151,976,759,784]" pageId="86" pageNumber="87">
<typeStatus id="54F5F0E4FFA65056FF23FD34FF0C4763" box="[151,201,760,784]" pageId="86" pageNumber="87">Type</typeStatus>
species:
<taxonomicName id="4C4E35C5FFA65056FE92FD34FCD5477C" authority="Franz, 1986 b: 237" authorityName="Franz" authorityPageNumber="237" authorityYear="1986" box="[294,784,759,783]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="ambatolampyensis">
<emphasis id="B93A9254FFA65056FE92FD34FD8B477C" box="[294,590,759,783]" italics="true" pageId="86" pageNumber="87">Euconnus ambatolampyensis</emphasis>
<bibRefCitation id="EFDF33B7FFA65056FDE7FD34FCD5477C" author="Franz, H." box="[595,784,759,783]" pageId="86" pageNumber="87" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 237</bibRefCitation>
</taxonomicName>
. (here designated).
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFA65056FF23FCF1FB984193" pageId="86" pageNumber="87" type="diagnosis">
<paragraph id="8BF14E46FFA65056FF23FCF1FB984193" blockId="86.[151,1437,829,2008]" pageId="86" pageNumber="87">
<emphasis id="B93A9254FFA65056FF23FCF1FED74724" bold="true" box="[151,274,829,855]" pageId="86" pageNumber="87">Diagnosis.</emphasis>
Body (
<figureCitation id="137552C3FFA65056FED7FCF1FE0C4724" box="[355,457,829,855]" captionStart="FIGURES 324331" captionStartId="88.[151,264,1672,1696]" captionTargetBox="[151,1419,322,1642]" captionTargetId="figure-77@88.[151,1435,322,1649]" captionTargetPageId="88" captionText="FIGURES 324331. Contreuconnus ambatolampyensis (Franz), holotype. Habitus in dorsal view (324); head in anterodorsal view (325); head and prothorax in lateral view (326); antenna (327); aedeagus in ventral (328, 330) and lateral (329, 331) views." figureDoi="http://doi.org/10.5281/zenodo.13746049" httpUri="https://zenodo.org/record/13746049/files/figure.png" pageId="86" pageNumber="87">Figs 324</figureCitation>
,
<figureCitation id="137552C3FFA65056FE60FCF1FE3A472B" box="[468,511,829,856]" captionStart="FIGURES 335342" captionStartId="90.[151,264,1484,1508]" captionTargetBox="[151,1425,181,1446]" captionTargetId="figure-28@90.[151,1435,181,1461]" captionTargetPageId="90" captionText="FIGURES 335342. Contreuconnus ambatolampyensis (Franz), holotype of Euconnus (Anthicimorphus) angustus Franz. Habitus in dorsal view (335); head in anterodorsal view (336); head and prothorax in lateral view (337); antenna (338); aedeagus in ventral (339, 341) and lateral (340, 342) views." figureDoi="http://doi.org/10.5281/zenodo.13746057" httpUri="https://zenodo.org/record/13746057/files/figure.png" pageId="86" pageNumber="87">335</figureCitation>
) with distinct constriction between head and pronotum and between pronotum and elytra; thick bristles present on genae and outer regions of hypomera (
<figureCitation id="137552C3FFA65056FC77FCADFBED4708" box="[963,1064,865,891]" captionStart="FIGURES 324331" captionStartId="88.[151,264,1672,1696]" captionTargetBox="[151,1419,322,1642]" captionTargetId="figure-77@88.[151,1435,322,1649]" captionTargetPageId="88" captionText="FIGURES 324331. Contreuconnus ambatolampyensis (Franz), holotype. Habitus in dorsal view (324); head in anterodorsal view (325); head and prothorax in lateral view (326); antenna (327); aedeagus in ventral (328, 330) and lateral (329, 331) views." figureDoi="http://doi.org/10.5281/zenodo.13746049" httpUri="https://zenodo.org/record/13746049/files/figure.png" pageId="86" pageNumber="87">Fig. 326</figureCitation>
); head (
<figureCitation id="137552C3FFA65056FB31FCADFAE0470F" box="[1157,1317,865,892]" captionStart="FIGURES 324331" captionStartId="88.[151,264,1672,1696]" captionTargetBox="[151,1419,322,1642]" captionTargetId="figure-77@88.[151,1435,322,1649]" captionTargetPageId="88" captionText="FIGURES 324331. Contreuconnus ambatolampyensis (Franz), holotype. Habitus in dorsal view (324); head in anterodorsal view (325); head and prothorax in lateral view (326); antenna (327); aedeagus in ventral (328, 330) and lateral (329, 331) views." figureDoi="http://doi.org/10.5281/zenodo.13746049" httpUri="https://zenodo.org/record/13746049/files/figure.png" pageId="86" pageNumber="87">Figs 325326</figureCitation>
,
<figureCitation id="137552C3FFA65056FA87FCADFA5D4708" box="[1331,1432,865,891]" captionStart="FIGURES 332334" captionStartId="89.[151,264,1174,1198]" captionTargetBox="[151,1435,273,1151]" captionTargetId="figure-61@89.[151,1435,273,1151]" captionTargetPageId="89" captionText="FIGURES 332334. Contreuconnus ambatolampyensis (Franz), holotype. Head, prothorax and elytral base in dorsal view (332); head and prothorax in ventral view (333); pterothorax in ventral view (334). Abbreviations: abp, antebasal pit; gp, gular plate; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746055" httpUri="https://zenodo.org/record/13746055/files/figure.png" pageId="86" pageNumber="87">332333</figureCitation>
,
<figureCitation id="137552C3FFA65056FF23FC49FF3E47EC" box="[151,251,901,927]" captionStart="FIGURES 335342" captionStartId="90.[151,264,1484,1508]" captionTargetBox="[151,1425,181,1446]" captionTargetId="figure-28@90.[151,1435,181,1461]" captionTargetPageId="90" captionText="FIGURES 335342. Contreuconnus ambatolampyensis (Franz), holotype of Euconnus (Anthicimorphus) angustus Franz. Habitus in dorsal view (335); head in anterodorsal view (336); head and prothorax in lateral view (337); antenna (338); aedeagus in ventral (339, 341) and lateral (340, 342) views." figureDoi="http://doi.org/10.5281/zenodo.13746057" httpUri="https://zenodo.org/record/13746057/files/figure.png" pageId="86" pageNumber="87">336337</figureCitation>
) round, with eyes situated closer to mandibular bases and antennal fossae than to occipital constriction; antennal fossae separated at middle by distance wider than width of scape (
<figureCitation id="137552C3FFA65056FBBEFC65FBB547B7" box="[1034,1136,937,964]" captionStart="FIGURES 324331" captionStartId="88.[151,264,1672,1696]" captionTargetBox="[151,1419,322,1642]" captionTargetId="figure-77@88.[151,1435,322,1649]" captionTargetPageId="88" captionText="FIGURES 324331. Contreuconnus ambatolampyensis (Franz), holotype. Habitus in dorsal view (324); head in anterodorsal view (325); head and prothorax in lateral view (326); antenna (327); aedeagus in ventral (328, 330) and lateral (329, 331) views." figureDoi="http://doi.org/10.5281/zenodo.13746049" httpUri="https://zenodo.org/record/13746049/files/figure.png" pageId="86" pageNumber="87">Fig. 325</figureCitation>
); antenna (
<figureCitation id="137552C3FFA65056FB45FC65FA9F47B0" box="[1265,1370,937,963]" captionStart="FIGURES 324331" captionStartId="88.[151,264,1672,1696]" captionTargetBox="[151,1419,322,1642]" captionTargetId="figure-77@88.[151,1435,322,1649]" captionTargetPageId="88" captionText="FIGURES 324331. Contreuconnus ambatolampyensis (Franz), holotype. Habitus in dorsal view (324); head in anterodorsal view (325); head and prothorax in lateral view (326); antenna (327); aedeagus in ventral (328, 330) and lateral (329, 331) views." figureDoi="http://doi.org/10.5281/zenodo.13746049" httpUri="https://zenodo.org/record/13746049/files/figure.png" pageId="86" pageNumber="87">Figs 327</figureCitation>
,
<figureCitation id="137552C3FFA65056FADDFC65FA5147B0" box="[1385,1428,937,963]" captionStart="FIGURES 335342" captionStartId="90.[151,264,1484,1508]" captionTargetBox="[151,1425,181,1446]" captionTargetId="figure-28@90.[151,1435,181,1461]" captionTargetPageId="90" captionText="FIGURES 335342. Contreuconnus ambatolampyensis (Franz), holotype of Euconnus (Anthicimorphus) angustus Franz. Habitus in dorsal view (335); head in anterodorsal view (336); head and prothorax in lateral view (337); antenna (338); aedeagus in ventral (339, 341) and lateral (340, 342) views." figureDoi="http://doi.org/10.5281/zenodo.13746057" httpUri="https://zenodo.org/record/13746057/files/figure.png" pageId="86" pageNumber="87">338</figureCitation>
) gradually thickening distally; frontoclypeal groove lacking; posterior margin of vertex (
<figureCitation id="137552C3FFA65056FB38FC01FAEA4794" box="[1164,1327,973,1000]" captionStart="FIGURES 335342" captionStartId="90.[151,264,1484,1508]" captionTargetBox="[151,1425,181,1446]" captionTargetId="figure-28@90.[151,1435,181,1461]" captionTargetPageId="90" captionText="FIGURES 335342. Contreuconnus ambatolampyensis (Franz), holotype of Euconnus (Anthicimorphus) angustus Franz. Habitus in dorsal view (335); head in anterodorsal view (336); head and prothorax in lateral view (337); antenna (338); aedeagus in ventral (339, 341) and lateral (340, 342) views." figureDoi="http://doi.org/10.5281/zenodo.13746057" httpUri="https://zenodo.org/record/13746057/files/figure.png" pageId="86" pageNumber="87">Figs 335, 336</figureCitation>
) rounded or nearly straight and not bulging posterodorsally; submental region (
<figureCitation id="137552C3FFA65056FC1BFC3DFBD74078" box="[943,1042,1009,1035]" captionStart="FIGURES 332334" captionStartId="89.[151,264,1174,1198]" captionTargetBox="[151,1435,273,1151]" captionTargetId="figure-61@89.[151,1435,273,1151]" captionTargetPageId="89" captionText="FIGURES 332334. Contreuconnus ambatolampyensis (Franz), holotype. Head, prothorax and elytral base in dorsal view (332); head and prothorax in ventral view (333); pterothorax in ventral view (334). Abbreviations: abp, antebasal pit; gp, gular plate; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746055" httpUri="https://zenodo.org/record/13746055/files/figure.png" pageId="86" pageNumber="87">Fig. 333</figureCitation>
;
<emphasis id="B93A9254FFA65056FB94FC3EFB884078" box="[1056,1101,1010,1035]" italics="true" pageId="86" pageNumber="87">smn</emphasis>
) lacking hypostomal sutures; hypostomal ridges (
<figureCitation id="137552C3FFA65056FECFFBD9FE25405C" box="[379,480,1045,1071]" captionStart="FIGURES 332334" captionStartId="89.[151,264,1174,1198]" captionTargetBox="[151,1435,273,1151]" captionTargetId="figure-61@89.[151,1435,273,1151]" captionTargetPageId="89" captionText="FIGURES 332334. Contreuconnus ambatolampyensis (Franz), holotype. Head, prothorax and elytral base in dorsal view (332); head and prothorax in ventral view (333); pterothorax in ventral view (334). Abbreviations: abp, antebasal pit; gp, gular plate; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746055" httpUri="https://zenodo.org/record/13746055/files/figure.png" pageId="86" pageNumber="87">Fig. 333</figureCitation>
;
<emphasis id="B93A9254FFA65056FE5BFBD9FDCD405C" box="[495,520,1045,1071]" italics="true" pageId="86" pageNumber="87">hr</emphasis>
) extending posteromesad and not reaching posterior tentorial pits, but laterally demarcating slightly elevated smooth platform; pronotum (
<figureCitation id="137552C3FFA65056FCE0FBF5FC724020" box="[852,951,1081,1107]" captionStart="FIGURES 332334" captionStartId="89.[151,264,1174,1198]" captionTargetBox="[151,1435,273,1151]" captionTargetId="figure-61@89.[151,1435,273,1151]" captionTargetPageId="89" captionText="FIGURES 332334. Contreuconnus ambatolampyensis (Franz), holotype. Head, prothorax and elytral base in dorsal view (332); head and prothorax in ventral view (333); pterothorax in ventral view (334). Abbreviations: abp, antebasal pit; gp, gular plate; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746055" httpUri="https://zenodo.org/record/13746055/files/figure.png" pageId="86" pageNumber="87">Fig. 332</figureCitation>
) bell-shaped, broadest in front of middle, with two pairs of antebasal pits, inner pair connected by shallow transverse impression, pronotum lacking lateral and sublateral carinae; prosternum (
<figureCitation id="137552C3FFA65056FD8EFB4DFD5840E8" box="[570,669,1153,1179]" captionStart="FIGURES 332334" captionStartId="89.[151,264,1174,1198]" captionTargetBox="[151,1435,273,1151]" captionTargetId="figure-61@89.[151,1435,273,1151]" captionTargetPageId="89" captionText="FIGURES 332334. Contreuconnus ambatolampyensis (Franz), holotype. Head, prothorax and elytral base in dorsal view (332); head and prothorax in ventral view (333); pterothorax in ventral view (334). Abbreviations: abp, antebasal pit; gp, gular plate; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746055" httpUri="https://zenodo.org/record/13746055/files/figure.png" pageId="86" pageNumber="87">Fig. 333</figureCitation>
) with precoxal (= basisternal) region much shorter than procoxal rests, prosternal process lacking; notosternal sutures (
<figureCitation id="137552C3FFA65056FD45FB69FC9740CC" box="[753,850,1189,1215]" captionStart="FIGURES 332334" captionStartId="89.[151,264,1174,1198]" captionTargetBox="[151,1435,273,1151]" captionTargetId="figure-61@89.[151,1435,273,1151]" captionTargetPageId="89" captionText="FIGURES 332334. Contreuconnus ambatolampyensis (Franz), holotype. Head, prothorax and elytral base in dorsal view (332); head and prothorax in ventral view (333); pterothorax in ventral view (334). Abbreviations: abp, antebasal pit; gp, gular plate; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746055" httpUri="https://zenodo.org/record/13746055/files/figure.png" pageId="86" pageNumber="87">Fig. 333</figureCitation>
;
<emphasis id="B93A9254FFA65056FCEBFB6AFC4640CC" box="[863,899,1190,1215]" italics="true" pageId="86" pageNumber="87">nss</emphasis>
) complete; procoxal cavity closed by postcoxal prosternal lobe overlapped by (but not fused with) postcoxal mesal lobe of hypomeron; hypomeral ridges nearly complete (slightly rounded off and less distinct in anterior fourth), each delimiting narrow subtriangular inner (adcoxal) unsculptured region of hypomeron; mesoventral intercoxal process (
<figureCitation id="137552C3FFA65056FC41FADDFB974158" box="[1013,1106,1297,1323]" captionStart="FIGURES 332334" captionStartId="89.[151,264,1174,1198]" captionTargetBox="[151,1435,273,1151]" captionTargetId="figure-61@89.[151,1435,273,1151]" captionTargetPageId="89" captionText="FIGURES 332334. Contreuconnus ambatolampyensis (Franz), holotype. Head, prothorax and elytral base in dorsal view (332); head and prothorax in ventral view (333); pterothorax in ventral view (334). Abbreviations: abp, antebasal pit; gp, gular plate; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746055" httpUri="https://zenodo.org/record/13746055/files/figure.png" pageId="86" pageNumber="87">Fig. 333</figureCitation>
<emphasis id="B93A9254FFA65056FBE3FADEFB544158" box="[1111,1169,1298,1323]" italics="true" pageId="86" pageNumber="87">msvp</emphasis>
) elongate subtriangular, barely marked, reaching middle of mesocoxal rests and not separating mesocoxae; metaventrite lacking anterior metaventral process; metaventral intermetacoxal process (
<figureCitation id="137552C3FFA65056FC92FA95FC4D4100" box="[806,904,1369,1395]" captionStart="FIGURES 332334" captionStartId="89.[151,264,1174,1198]" captionTargetBox="[151,1435,273,1151]" captionTargetId="figure-61@89.[151,1435,273,1151]" captionTargetPageId="89" captionText="FIGURES 332334. Contreuconnus ambatolampyensis (Franz), holotype. Head, prothorax and elytral base in dorsal view (332); head and prothorax in ventral view (333); pterothorax in ventral view (334). Abbreviations: abp, antebasal pit; gp, gular plate; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746055" httpUri="https://zenodo.org/record/13746055/files/figure.png" pageId="86" pageNumber="87">Fig. 333</figureCitation>
;
<emphasis id="B93A9254FFA65056FC21FA96FC0E4100" box="[917,971,1370,1395]" italics="true" pageId="86" pageNumber="87">mtvp</emphasis>
) short and broad, with concave posterior margin, widely separating metacoxae; elytra (
<figureCitation id="137552C3FFA65056FD1EFAB1FCCB41E4" box="[682,782,1405,1431]" captionStart="FIGURES 332334" captionStartId="89.[151,264,1174,1198]" captionTargetBox="[151,1435,273,1151]" captionTargetId="figure-61@89.[151,1435,273,1151]" captionTargetPageId="89" captionText="FIGURES 332334. Contreuconnus ambatolampyensis (Franz), holotype. Head, prothorax and elytral base in dorsal view (332); head and prothorax in ventral view (333); pterothorax in ventral view (334). Abbreviations: abp, antebasal pit; gp, gular plate; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746055" httpUri="https://zenodo.org/record/13746055/files/figure.png" pageId="86" pageNumber="87">Fig. 332</figureCitation>
) lacking basal foveae; mesoscutellar shield not exposed between elytral bases (
<figureCitation id="137552C3FFA65056FE2FFA6DFDC541C8" box="[411,512,1441,1467]" captionStart="FIGURES 332334" captionStartId="89.[151,264,1174,1198]" captionTargetBox="[151,1435,273,1151]" captionTargetId="figure-61@89.[151,1435,273,1151]" captionTargetPageId="89" captionText="FIGURES 332334. Contreuconnus ambatolampyensis (Franz), holotype. Head, prothorax and elytral base in dorsal view (332); head and prothorax in ventral view (333); pterothorax in ventral view (334). Abbreviations: abp, antebasal pit; gp, gular plate; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746055" httpUri="https://zenodo.org/record/13746055/files/figure.png" pageId="86" pageNumber="87">Fig. 332</figureCitation>
); aedeagus (
<figureCitation id="137552C3FFA65056FD39FA6DFCE941C8" box="[653,812,1441,1467]" captionStart="FIGURES 324331" captionStartId="88.[151,264,1672,1696]" captionTargetBox="[151,1419,322,1642]" captionTargetId="figure-77@88.[151,1435,322,1649]" captionTargetPageId="88" captionText="FIGURES 324331. Contreuconnus ambatolampyensis (Franz), holotype. Habitus in dorsal view (324); head in anterodorsal view (325); head and prothorax in lateral view (326); antenna (327); aedeagus in ventral (328, 330) and lateral (329, 331) views." figureDoi="http://doi.org/10.5281/zenodo.13746049" httpUri="https://zenodo.org/record/13746049/files/figure.png" pageId="86" pageNumber="87">Figs 328331</figureCitation>
,
<figureCitation id="137552C3FFA65056FC8DFA6DFC5841C8" box="[825,925,1441,1467]" captionStart="FIGURES 335342" captionStartId="90.[151,264,1484,1508]" captionTargetBox="[151,1425,181,1446]" captionTargetId="figure-28@90.[151,1435,181,1461]" captionTargetPageId="90" captionText="FIGURES 335342. Contreuconnus ambatolampyensis (Franz), holotype of Euconnus (Anthicimorphus) angustus Franz. Habitus in dorsal view (335); head in anterodorsal view (336); head and prothorax in lateral view (337); antenna (338); aedeagus in ventral (339, 341) and lateral (340, 342) views." figureDoi="http://doi.org/10.5281/zenodo.13746057" httpUri="https://zenodo.org/record/13746057/files/figure.png" pageId="86" pageNumber="87">339342</figureCitation>
) with symmetrical median lobe and strongly sclerotized, asymmetrical and complex endophallus, and with free, slender parameres.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFA65057FF73FA25FCCD427A" lastPageId="87" lastPageNumber="88" pageId="86" pageNumber="87" type="description">
<paragraph id="8BF14E46FFA65056FF73FA25FC9C4254" blockId="86.[151,1437,829,2008]" pageId="86" pageNumber="87">
<emphasis id="B93A9254FFA65056FF73FA25FE9C4270" bold="true" box="[199,345,1513,1539]" pageId="86" pageNumber="87">Description.</emphasis>
Body form (
<figureCitation id="137552C3FFA65056FE40FA25FD9B4270" box="[500,606,1513,1539]" captionStart="FIGURES 324331" captionStartId="88.[151,264,1672,1696]" captionTargetBox="[151,1419,322,1642]" captionTargetId="figure-77@88.[151,1435,322,1649]" captionTargetPageId="88" captionText="FIGURES 324331. Contreuconnus ambatolampyensis (Franz), holotype. Habitus in dorsal view (324); head in anterodorsal view (325); head and prothorax in lateral view (326); antenna (327); aedeagus in ventral (328, 330) and lateral (329, 331) views." figureDoi="http://doi.org/10.5281/zenodo.13746049" httpUri="https://zenodo.org/record/13746049/files/figure.png" pageId="86" pageNumber="87">Figs 324</figureCitation>
,
<figureCitation id="137552C3FFA65056FDD9FA25FD5D4277" box="[621,664,1513,1540]" captionStart="FIGURES 335342" captionStartId="90.[151,264,1484,1508]" captionTargetBox="[151,1425,181,1446]" captionTargetId="figure-28@90.[151,1435,181,1461]" captionTargetPageId="90" captionText="FIGURES 335342. Contreuconnus ambatolampyensis (Franz), holotype of Euconnus (Anthicimorphus) angustus Franz. Habitus in dorsal view (335); head in anterodorsal view (336); head and prothorax in lateral view (337); antenna (338); aedeagus in ventral (339, 341) and lateral (340, 342) views." figureDoi="http://doi.org/10.5281/zenodo.13746057" httpUri="https://zenodo.org/record/13746057/files/figure.png" pageId="86" pageNumber="87">335</figureCitation>
) elongate and slender, strongly convex, with distinct constrictions between head and pronotum and between pronotum and elytra.
</paragraph>
<paragraph id="8BF14E46FFA65056FF73F9FDFAC24318" blockId="86.[151,1437,829,2008]" pageId="86" pageNumber="87">
Head (
<figureCitation id="137552C3FFA65056FEBBF9FDFE6F423F" box="[271,426,1585,1612]" captionStart="FIGURES 324331" captionStartId="88.[151,264,1672,1696]" captionTargetBox="[151,1419,322,1642]" captionTargetId="figure-77@88.[151,1435,322,1649]" captionTargetPageId="88" captionText="FIGURES 324331. Contreuconnus ambatolampyensis (Franz), holotype. Habitus in dorsal view (324); head in anterodorsal view (325); head and prothorax in lateral view (326); antenna (327); aedeagus in ventral (328, 330) and lateral (329, 331) views." figureDoi="http://doi.org/10.5281/zenodo.13746049" httpUri="https://zenodo.org/record/13746049/files/figure.png" pageId="86" pageNumber="87">Figs 325326</figureCitation>
,
<figureCitation id="137552C3FFA65056FE00F9FDFDDD4238" box="[436,536,1585,1611]" captionStart="FIGURES 332334" captionStartId="89.[151,264,1174,1198]" captionTargetBox="[151,1435,273,1151]" captionTargetId="figure-61@89.[151,1435,273,1151]" captionTargetPageId="89" captionText="FIGURES 332334. Contreuconnus ambatolampyensis (Franz), holotype. Head, prothorax and elytral base in dorsal view (332); head and prothorax in ventral view (333); pterothorax in ventral view (334). Abbreviations: abp, antebasal pit; gp, gular plate; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746055" httpUri="https://zenodo.org/record/13746055/files/figure.png" pageId="86" pageNumber="87">332333</figureCitation>
,
<figureCitation id="137552C3FFA65056FD96F9FDFD404238" box="[546,645,1585,1611]" captionStart="FIGURES 335342" captionStartId="90.[151,264,1484,1508]" captionTargetBox="[151,1425,181,1446]" captionTargetId="figure-28@90.[151,1435,181,1461]" captionTargetPageId="90" captionText="FIGURES 335342. Contreuconnus ambatolampyensis (Franz), holotype of Euconnus (Anthicimorphus) angustus Franz. Habitus in dorsal view (335); head in anterodorsal view (336); head and prothorax in lateral view (337); antenna (338); aedeagus in ventral (339, 341) and lateral (340, 342) views." figureDoi="http://doi.org/10.5281/zenodo.13746057" httpUri="https://zenodo.org/record/13746057/files/figure.png" pageId="86" pageNumber="87">336337</figureCitation>
) rounded; compound eyes situated much closer to mandibular bases and antennal fossae than to occipital constriction, each with posterior emargination (
<figureCitation id="137552C3FFA65056FB94F999FB43421C" box="[1056,1158,1621,1647]" captionStart="FIGURES 324331" captionStartId="88.[151,264,1672,1696]" captionTargetBox="[151,1419,322,1642]" captionTargetId="figure-77@88.[151,1435,322,1649]" captionTargetPageId="88" captionText="FIGURES 324331. Contreuconnus ambatolampyensis (Franz), holotype. Habitus in dorsal view (324); head in anterodorsal view (325); head and prothorax in lateral view (326); antenna (327); aedeagus in ventral (328, 330) and lateral (329, 331) views." figureDoi="http://doi.org/10.5281/zenodo.13746049" httpUri="https://zenodo.org/record/13746049/files/figure.png" pageId="86" pageNumber="87">Figs 326</figureCitation>
,
<figureCitation id="137552C3FFA65056FB26F999FB05421C" box="[1170,1216,1621,1647]" captionStart="FIGURES 335342" captionStartId="90.[151,264,1484,1508]" captionTargetBox="[151,1425,181,1446]" captionTargetId="figure-28@90.[151,1435,181,1461]" captionTargetPageId="90" captionText="FIGURES 335342. Contreuconnus ambatolampyensis (Franz), holotype of Euconnus (Anthicimorphus) angustus Franz. Habitus in dorsal view (335); head in anterodorsal view (336); head and prothorax in lateral view (337); antenna (338); aedeagus in ventral (339, 341) and lateral (340, 342) views." figureDoi="http://doi.org/10.5281/zenodo.13746057" httpUri="https://zenodo.org/record/13746057/files/figure.png" pageId="86" pageNumber="87">337</figureCitation>
); tempora rounded; vertex (
<figureCitation id="137552C3FFA65056FF44F9B5FE9C42E7" box="[240,345,1657,1684]" captionStart="FIGURES 324331" captionStartId="88.[151,264,1672,1696]" captionTargetBox="[151,1419,322,1642]" captionTargetId="figure-77@88.[151,1435,322,1649]" captionTargetPageId="88" captionText="FIGURES 324331. Contreuconnus ambatolampyensis (Franz), holotype. Habitus in dorsal view (324); head in anterodorsal view (325); head and prothorax in lateral view (326); antenna (327); aedeagus in ventral (328, 330) and lateral (329, 331) views." figureDoi="http://doi.org/10.5281/zenodo.13746049" httpUri="https://zenodo.org/record/13746049/files/figure.png" pageId="86" pageNumber="87">Figs 325</figureCitation>
,
<figureCitation id="137552C3FFA65056FEDCF9B5FE5142E0" box="[360,404,1657,1683]" captionStart="FIGURES 335342" captionStartId="90.[151,264,1484,1508]" captionTargetBox="[151,1425,181,1446]" captionTargetId="figure-28@90.[151,1435,181,1461]" captionTargetPageId="90" captionText="FIGURES 335342. Contreuconnus ambatolampyensis (Franz), holotype of Euconnus (Anthicimorphus) angustus Franz. Habitus in dorsal view (335); head in anterodorsal view (336); head and prothorax in lateral view (337); antenna (338); aedeagus in ventral (339, 341) and lateral (340, 342) views." figureDoi="http://doi.org/10.5281/zenodo.13746057" httpUri="https://zenodo.org/record/13746057/files/figure.png" pageId="86" pageNumber="87">336</figureCitation>
) not bulging posterodorsally, confluent with frons, its posterior margin rounded. Bristles present on genae (
<figureCitation id="137552C3FFA65056FED5F951FE0342C4" box="[353,454,1693,1719]" captionStart="FIGURES 324331" captionStartId="88.[151,264,1672,1696]" captionTargetBox="[151,1419,322,1642]" captionTargetId="figure-77@88.[151,1435,322,1649]" captionTargetPageId="88" captionText="FIGURES 324331. Contreuconnus ambatolampyensis (Franz), holotype. Habitus in dorsal view (324); head in anterodorsal view (325); head and prothorax in lateral view (326); antenna (327); aedeagus in ventral (328, 330) and lateral (329, 331) views." figureDoi="http://doi.org/10.5281/zenodo.13746049" httpUri="https://zenodo.org/record/13746049/files/figure.png" pageId="86" pageNumber="87">Figs 326</figureCitation>
,
<figureCitation id="137552C3FFA65056FE65F951FE3B42C4" box="[465,510,1693,1719]" captionStart="FIGURES 332334" captionStartId="89.[151,264,1174,1198]" captionTargetBox="[151,1435,273,1151]" captionTargetId="figure-61@89.[151,1435,273,1151]" captionTargetPageId="89" captionText="FIGURES 332334. Contreuconnus ambatolampyensis (Franz), holotype. Head, prothorax and elytral base in dorsal view (332); head and prothorax in ventral view (333); pterothorax in ventral view (334). Abbreviations: abp, antebasal pit; gp, gular plate; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746055" httpUri="https://zenodo.org/record/13746055/files/figure.png" pageId="86" pageNumber="87">333</figureCitation>
,
<figureCitation id="137552C3FFA65056FDBCF951FDF342C4" box="[520,566,1693,1719]" captionStart="FIGURES 335342" captionStartId="90.[151,264,1484,1508]" captionTargetBox="[151,1425,181,1446]" captionTargetId="figure-28@90.[151,1435,181,1461]" captionTargetPageId="90" captionText="FIGURES 335342. Contreuconnus ambatolampyensis (Franz), holotype of Euconnus (Anthicimorphus) angustus Franz. Habitus in dorsal view (335); head in anterodorsal view (336); head and prothorax in lateral view (337); antenna (338); aedeagus in ventral (339, 341) and lateral (340, 342) views." figureDoi="http://doi.org/10.5281/zenodo.13746057" httpUri="https://zenodo.org/record/13746057/files/figure.png" pageId="86" pageNumber="87">337</figureCitation>
). Occipital constriction (
<figureCitation id="137552C3FFA65056FCFDF951FC6C42C4" box="[841,937,1693,1719]" captionStart="FIGURES 332334" captionStartId="89.[151,264,1174,1198]" captionTargetBox="[151,1435,273,1151]" captionTargetId="figure-61@89.[151,1435,273,1151]" captionTargetPageId="89" captionText="FIGURES 332334. Contreuconnus ambatolampyensis (Franz), holotype. Head, prothorax and elytral base in dorsal view (332); head and prothorax in ventral view (333); pterothorax in ventral view (334). Abbreviations: abp, antebasal pit; gp, gular plate; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746055" httpUri="https://zenodo.org/record/13746055/files/figure.png" pageId="86" pageNumber="87">Fig. 332</figureCitation>
) about as broad as half width of head; neck region strongly broadening posterad. Antennal fossae large and situated anterolaterally, separated at middle by median longitudinal frontal area wider than scape (
<figureCitation id="137552C3FFA65056FD65F929FCF64373" box="[721,819,1765,1792]" captionStart="FIGURES 324331" captionStartId="88.[151,264,1672,1696]" captionTargetBox="[151,1419,322,1642]" captionTargetId="figure-77@88.[151,1435,322,1649]" captionTargetPageId="88" captionText="FIGURES 324331. Contreuconnus ambatolampyensis (Franz), holotype. Habitus in dorsal view (324); head in anterodorsal view (325); head and prothorax in lateral view (326); antenna (327); aedeagus in ventral (328, 330) and lateral (329, 331) views." figureDoi="http://doi.org/10.5281/zenodo.13746049" httpUri="https://zenodo.org/record/13746049/files/figure.png" pageId="86" pageNumber="87">Fig. 325</figureCitation>
). Frontoclypeal groove lacking; clypeus short, rounded anteriorly, with lateral margins converging towards labrum. Gular plate (
<figureCitation id="137552C3FFA65056FC76F8C5FBE64350" box="[962,1059,1801,1827]" captionStart="FIGURES 332334" captionStartId="89.[151,264,1174,1198]" captionTargetBox="[151,1435,273,1151]" captionTargetId="figure-61@89.[151,1435,273,1151]" captionTargetPageId="89" captionText="FIGURES 332334. Contreuconnus ambatolampyensis (Franz), holotype. Head, prothorax and elytral base in dorsal view (332); head and prothorax in ventral view (333); pterothorax in ventral view (334). Abbreviations: abp, antebasal pit; gp, gular plate; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746055" httpUri="https://zenodo.org/record/13746055/files/figure.png" pageId="86" pageNumber="87">Fig. 333</figureCitation>
;
<emphasis id="B93A9254FFA65056FB9AF8C6FB8F4350" box="[1070,1098,1802,1827]" italics="true" pageId="86" pageNumber="87">gp</emphasis>
) subtrapezoidal, covered with transverse microsculpture, lacking gular sutures; posterior tentorial pits (
<figureCitation id="137552C3FFA65056FC7BF8E1FBF74334" box="[975,1074,1837,1863]" captionStart="FIGURES 332334" captionStartId="89.[151,264,1174,1198]" captionTargetBox="[151,1435,273,1151]" captionTargetId="figure-61@89.[151,1435,273,1151]" captionTargetPageId="89" captionText="FIGURES 332334. Contreuconnus ambatolampyensis (Franz), holotype. Head, prothorax and elytral base in dorsal view (332); head and prothorax in ventral view (333); pterothorax in ventral view (334). Abbreviations: abp, antebasal pit; gp, gular plate; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746055" httpUri="https://zenodo.org/record/13746055/files/figure.png" pageId="86" pageNumber="87">Fig. 333</figureCitation>
;
<emphasis id="B93A9254FFA65056FB8BF8E2FBA64334" box="[1087,1123,1838,1863]" italics="true" pageId="86" pageNumber="87">ptp</emphasis>
) situated clearly in front of transverse impression demarcating ventrally 'neck' region, each strongly elongate and weakly curved.
</paragraph>
<paragraph id="8BF14E46FFA65057FF73F8B9FE1845A2" blockId="86.[151,1437,829,2008]" lastBlockId="87.[151,1437,151,1761]" lastPageId="87" lastPageNumber="88" pageId="86" pageNumber="87">
Submentum (
<figureCitation id="137552C3FFA65056FEEAF8B9FE7A43FC" box="[350,447,1909,1935]" captionStart="FIGURES 332334" captionStartId="89.[151,264,1174,1198]" captionTargetBox="[151,1435,273,1151]" captionTargetId="figure-61@89.[151,1435,273,1151]" captionTargetPageId="89" captionText="FIGURES 332334. Contreuconnus ambatolampyensis (Franz), holotype. Head, prothorax and elytral base in dorsal view (332); head and prothorax in ventral view (333); pterothorax in ventral view (334). Abbreviations: abp, antebasal pit; gp, gular plate; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746055" httpUri="https://zenodo.org/record/13746055/files/figure.png" pageId="86" pageNumber="87">Fig. 333</figureCitation>
;
<emphasis id="B93A9254FFA65056FE7FF8BAFE3D43FC" box="[459,504,1910,1935]" italics="true" pageId="86" pageNumber="87">smn</emphasis>
) laterally not demarcated by hypostomal sutures; mentum subtrapezoidal, broadest near base, with anterior margin rounded on sides and indistinctly concave at middle; prelabium short with narrowly separated insertions of labial palps and with narrow and short ligula bearing two long setae; labial palpomere 1 annulate, palpomere 2 strongly elongate, subcylindrical, nearly 4 times as long as broad, palpomere 3 narrow, elongate but distinctly shorter than 2, tapering distally and weakly curved. Maxilla generalized as in most Stenichnini, with large semicircular cardo, subtriangular basistipes, elongate mediostipes, elongate lacinia and galea, each with thick and not very numerous setae along distomesal margin, palpifer elongate, maxillary palp with distinctly elongate and distally slightly broadening palpomere 1, long, pipe-shaped and weakly curved palpomere 2, strongly elongate and slender fusiform palpomere 3 about twice as wide as palpomere 2, and minute, strongly elongate subconical and pointed palpomere 4. Mandibles broadly subtriangular, each with one preapical mesal tooth and short setose prostheca. Labrum strongly transverse, with rounded anterior margin, and with sparse, symmetrically distributed dorsal setae directed anterad.
</paragraph>
<paragraph id="8BF14E46FFA75057FF73FE17FF26464E" blockId="87.[151,1437,151,1761]" pageId="87" pageNumber="88">
Antenna (
<figureCitation id="137552C3FFA75057FE8EFE17FE604586" box="[314,421,475,501]" captionStart="FIGURES 324331" captionStartId="88.[151,264,1672,1696]" captionTargetBox="[151,1419,322,1642]" captionTargetId="figure-77@88.[151,1435,322,1649]" captionTargetPageId="88" captionText="FIGURES 324331. Contreuconnus ambatolampyensis (Franz), holotype. Habitus in dorsal view (324); head in anterodorsal view (325); head and prothorax in lateral view (326); antenna (327); aedeagus in ventral (328, 330) and lateral (329, 331) views." figureDoi="http://doi.org/10.5281/zenodo.13746049" httpUri="https://zenodo.org/record/13746049/files/figure.png" pageId="87" pageNumber="88">Figs 327</figureCitation>
,
<figureCitation id="137552C3FFA75057FE02FE17FE244586" box="[438,481,475,501]" captionStart="FIGURES 335342" captionStartId="90.[151,264,1484,1508]" captionTargetBox="[151,1425,181,1446]" captionTargetId="figure-28@90.[151,1435,181,1461]" captionTargetPageId="90" captionText="FIGURES 335342. Contreuconnus ambatolampyensis (Franz), holotype of Euconnus (Anthicimorphus) angustus Franz. Habitus in dorsal view (335); head in anterodorsal view (336); head and prothorax in lateral view (337); antenna (338); aedeagus in ventral (339, 341) and lateral (340, 342) views." figureDoi="http://doi.org/10.5281/zenodo.13746057" httpUri="https://zenodo.org/record/13746057/files/figure.png" pageId="87" pageNumber="88">338</figureCitation>
) long in relation to body length, slender, with scape and pedicel each elongate; flagellomeres loosely assembled; antenna gradually and weakly thickening distally. All antennomeres sparsely setose.
</paragraph>
<paragraph id="8BF14E46FFA75057FF73FD8BFDE747D6" blockId="87.[151,1437,151,1761]" pageId="87" pageNumber="88">
Prothorax in dorsal view (
<figureCitation id="137552C3FFA75057FE4CFD8BFD9E4612" box="[504,603,583,609]" captionStart="FIGURES 332334" captionStartId="89.[151,264,1174,1198]" captionTargetBox="[151,1435,273,1151]" captionTargetId="figure-61@89.[151,1435,273,1151]" captionTargetPageId="89" captionText="FIGURES 332334. Contreuconnus ambatolampyensis (Franz), holotype. Head, prothorax and elytral base in dorsal view (332); head and prothorax in ventral view (333); pterothorax in ventral view (334). Abbreviations: abp, antebasal pit; gp, gular plate; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746055" httpUri="https://zenodo.org/record/13746055/files/figure.png" pageId="87" pageNumber="88">Fig. 332</figureCitation>
) bell-shaped, broadest distinctly in front of middle. Anterior and lateral margins confluent, without anterior angles; lateral margins strongly rounded in anterior half and sinuate posteriorly; posterior corners distinct but blunt; posterior margin weakly convex posteriorly. Pronotal base with two pairs of antebasal pits, inner pair connected by shallow transverse impression; sublateral and lateral carinae lacking. Bristles present only on anterior regions of outer hypomera (
<figureCitation id="137552C3FFA75057FD5BFD1BFC554682" box="[751,912,727,753]" captionStart="FIGURES 332334" captionStartId="89.[151,264,1174,1198]" captionTargetBox="[151,1435,273,1151]" captionTargetId="figure-61@89.[151,1435,273,1151]" captionTargetPageId="89" captionText="FIGURES 332334. Contreuconnus ambatolampyensis (Franz), holotype. Head, prothorax and elytral base in dorsal view (332); head and prothorax in ventral view (333); pterothorax in ventral view (334). Abbreviations: abp, antebasal pit; gp, gular plate; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746055" httpUri="https://zenodo.org/record/13746055/files/figure.png" pageId="87" pageNumber="88">Figs 332333</figureCitation>
), slightly expanding onto dorsum. Hypomeral ridges complete (slightly rounded off anteriorly), demarcating narrow and unsculptured inner regions of hypomera. Prosternum (
<figureCitation id="137552C3FFA75057FE9CFCD3FE4D474A" box="[296,392,799,825]" captionStart="FIGURES 332334" captionStartId="89.[151,264,1174,1198]" captionTargetBox="[151,1435,273,1151]" captionTargetId="figure-61@89.[151,1435,273,1151]" captionTargetPageId="89" captionText="FIGURES 332334. Contreuconnus ambatolampyensis (Franz), holotype. Head, prothorax and elytral base in dorsal view (332); head and prothorax in ventral view (333); pterothorax in ventral view (334). Abbreviations: abp, antebasal pit; gp, gular plate; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746055" httpUri="https://zenodo.org/record/13746055/files/figure.png" pageId="87" pageNumber="88">Fig. 333</figureCitation>
) slightly shorter than half length of prothorax, with precoxal region much shorter than coxal rests; notosternal sutures (
<figureCitation id="137552C3FFA75057FE07FC8FFDD4472E" box="[435,529,835,861]" captionStart="FIGURES 332334" captionStartId="89.[151,264,1174,1198]" captionTargetBox="[151,1435,273,1151]" captionTargetId="figure-61@89.[151,1435,273,1151]" captionTargetPageId="89" captionText="FIGURES 332334. Contreuconnus ambatolampyensis (Franz), holotype. Head, prothorax and elytral base in dorsal view (332); head and prothorax in ventral view (333); pterothorax in ventral view (334). Abbreviations: abp, antebasal pit; gp, gular plate; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746055" httpUri="https://zenodo.org/record/13746055/files/figure.png" pageId="87" pageNumber="88">Fig.333</figureCitation>
;
<emphasis id="B93A9254FFA75057FDAEFC8FFDFB472F" box="[538,574,835,860]" italics="true" pageId="87" pageNumber="88">nss</emphasis>
) complete.Prosternal process not developed; posterior margin of furcasternum with shallow arcuate median emargination. Procoxal cavities closed; postcoxal hypomeral lobe overlapping (but not fused to) postcoxal prosternal lobe.
</paragraph>
<paragraph id="8BF14E46FFA75057FF73FC63FB0147BA" blockId="87.[151,1437,151,1761]" box="[199,1220,943,969]" pageId="87" pageNumber="88">Meso- and metanotum not studied; mesoscutellar shield not exposed between elytral bases.</paragraph>
<paragraph id="8BF14E46FFA75057FF73FC1FFCA8402A" blockId="87.[151,1437,151,1761]" pageId="87" pageNumber="88">
Mesoventrite (
<figureCitation id="137552C3FFA75057FEDEFC1FFE0E479E" box="[362,459,979,1005]" captionStart="FIGURES 332334" captionStartId="89.[151,264,1174,1198]" captionTargetBox="[151,1435,273,1151]" captionTargetId="figure-61@89.[151,1435,273,1151]" captionTargetPageId="89" captionText="FIGURES 332334. Contreuconnus ambatolampyensis (Franz), holotype. Head, prothorax and elytral base in dorsal view (332); head and prothorax in ventral view (333); pterothorax in ventral view (334). Abbreviations: abp, antebasal pit; gp, gular plate; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746055" httpUri="https://zenodo.org/record/13746055/files/figure.png" pageId="87" pageNumber="88">Fig. 334</figureCitation>
) transverse, with massive prepectus, subquadrate and setose anteromedian impressions functioning as procoxal rests (
<figureCitation id="137552C3FFA75057FE4AFC3BFDA04062" box="[510,613,1015,1041]" captionStart="FIGURES 332334" captionStartId="89.[151,264,1174,1198]" captionTargetBox="[151,1435,273,1151]" captionTargetId="figure-61@89.[151,1435,273,1151]" captionTargetPageId="89" captionText="FIGURES 332334. Contreuconnus ambatolampyensis (Franz), holotype. Head, prothorax and elytral base in dorsal view (332); head and prothorax in ventral view (333); pterothorax in ventral view (334). Abbreviations: abp, antebasal pit; gp, gular plate; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746055" httpUri="https://zenodo.org/record/13746055/files/figure.png" pageId="87" pageNumber="88">Fig. 334</figureCitation>
;
<emphasis id="B93A9254FFA75057FDC2FC3BFD5E4063" box="[630,667,1015,1040]" italics="true" pageId="87" pageNumber="88">pcr</emphasis>
) only anteriorly separated at middle by short longitudinal carina posteriorly not connected to broadly subtriangular and barely marked mesoventral intercoxal process (
<figureCitation id="137552C3FFA75057FA87FBD7FA524046" box="[1331,1431,1051,1077]" captionStart="FIGURES 332334" captionStartId="89.[151,264,1174,1198]" captionTargetBox="[151,1435,273,1151]" captionTargetId="figure-61@89.[151,1435,273,1151]" captionTargetPageId="89" captionText="FIGURES 332334. Contreuconnus ambatolampyensis (Franz), holotype. Head, prothorax and elytral base in dorsal view (332); head and prothorax in ventral view (333); pterothorax in ventral view (334). Abbreviations: abp, antebasal pit; gp, gular plate; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746055" httpUri="https://zenodo.org/record/13746055/files/figure.png" pageId="87" pageNumber="88">Fig. 334</figureCitation>
;
<emphasis id="B93A9254FFA75057FF23FBF3FF14402B" box="[151,209,1087,1112]" italics="true" pageId="87" pageNumber="88">msvp</emphasis>
) which posteriorly reaches only middle of mesocoxal rests.
</paragraph>
<paragraph id="8BF14E46FFA75057FF73FBAFFEED409A" blockId="87.[151,1437,151,1761]" pageId="87" pageNumber="88">
Metaventrite (
<figureCitation id="137552C3FFA75057FED9FBAFFE10400E" box="[365,469,1123,1149]" captionStart="FIGURES 332334" captionStartId="89.[151,264,1174,1198]" captionTargetBox="[151,1435,273,1151]" captionTargetId="figure-61@89.[151,1435,273,1151]" captionTargetPageId="89" captionText="FIGURES 332334. Contreuconnus ambatolampyensis (Franz), holotype. Head, prothorax and elytral base in dorsal view (332); head and prothorax in ventral view (333); pterothorax in ventral view (334). Abbreviations: abp, antebasal pit; gp, gular plate; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746055" httpUri="https://zenodo.org/record/13746055/files/figure.png" pageId="87" pageNumber="88">Fig. 334</figureCitation>
) subrectangular, anteriorly fused with mesoventrite, lacking anterior metaventral process; metaventral intermetacoxal process (
<figureCitation id="137552C3FFA75057FD1CFB4BFCCB40D2" box="[680,782,1159,1185]" captionStart="FIGURES 332334" captionStartId="89.[151,264,1174,1198]" captionTargetBox="[151,1435,273,1151]" captionTargetId="figure-61@89.[151,1435,273,1151]" captionTargetPageId="89" captionText="FIGURES 332334. Contreuconnus ambatolampyensis (Franz), holotype. Head, prothorax and elytral base in dorsal view (332); head and prothorax in ventral view (333); pterothorax in ventral view (334). Abbreviations: abp, antebasal pit; gp, gular plate; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746055" httpUri="https://zenodo.org/record/13746055/files/figure.png" pageId="87" pageNumber="88">Fig. 334</figureCitation>
;
<emphasis id="B93A9254FFA75057FCAAFB4BFC9140D3" box="[798,852,1159,1184]" italics="true" pageId="87" pageNumber="88">mtvp</emphasis>
) short and broad, with concave posterior margin, widely separating metacoxae. Metanepisterna and metepimera fully demarcated and narrow, partly exposed in ventral view.
</paragraph>
<paragraph id="8BF14E46FFA75057FF73FB3FFDA3417E" blockId="87.[151,1437,151,1761]" box="[199,614,1267,1293]" pageId="87" pageNumber="88">Legs short, lacking peculiar features.</paragraph>
<paragraph id="8BF14E46FFA75057FF73FADBFEEF4126" blockId="87.[151,1437,151,1761]" pageId="87" pageNumber="88">
Elytra (
<figureCitation id="137552C3FFA75057FEAEFADBFE454142" box="[282,384,1303,1329]" captionStart="FIGURES 324331" captionStartId="88.[151,264,1672,1696]" captionTargetBox="[151,1419,322,1642]" captionTargetId="figure-77@88.[151,1435,322,1649]" captionTargetPageId="88" captionText="FIGURES 324331. Contreuconnus ambatolampyensis (Franz), holotype. Habitus in dorsal view (324); head in anterodorsal view (325); head and prothorax in lateral view (326); antenna (327); aedeagus in ventral (328, 330) and lateral (329, 331) views." figureDoi="http://doi.org/10.5281/zenodo.13746049" httpUri="https://zenodo.org/record/13746049/files/figure.png" pageId="87" pageNumber="88">Figs 324</figureCitation>
,
<figureCitation id="137552C3FFA75057FE3FFADBFE7D4142" box="[395,440,1303,1329]" captionStart="FIGURES 332334" captionStartId="89.[151,264,1174,1198]" captionTargetBox="[151,1435,273,1151]" captionTargetId="figure-61@89.[151,1435,273,1151]" captionTargetPageId="89" captionText="FIGURES 332334. Contreuconnus ambatolampyensis (Franz), holotype. Head, prothorax and elytral base in dorsal view (332); head and prothorax in ventral view (333); pterothorax in ventral view (334). Abbreviations: abp, antebasal pit; gp, gular plate; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13746055" httpUri="https://zenodo.org/record/13746055/files/figure.png" pageId="87" pageNumber="88">332</figureCitation>
,
<figureCitation id="137552C3FFA75057FE77FADAFE2B4142" box="[451,494,1302,1329]" captionStart="FIGURES 335342" captionStartId="90.[151,264,1484,1508]" captionTargetBox="[151,1425,181,1446]" captionTargetId="figure-28@90.[151,1435,181,1461]" captionTargetPageId="90" captionText="FIGURES 335342. Contreuconnus ambatolampyensis (Franz), holotype of Euconnus (Anthicimorphus) angustus Franz. Habitus in dorsal view (335); head in anterodorsal view (336); head and prothorax in lateral view (337); antenna (338); aedeagus in ventral (339, 341) and lateral (340, 342) views." figureDoi="http://doi.org/10.5281/zenodo.13746057" httpUri="https://zenodo.org/record/13746057/files/figure.png" pageId="87" pageNumber="88">335</figureCitation>
) together oval, with humeral calli, lacking basal impressions, humeral denticles and basal foveae.
</paragraph>
<paragraph id="8BF14E46FFA75057FF73FA93FCF4410A" blockId="87.[151,1437,151,1761]" box="[199,817,1375,1401]" pageId="87" pageNumber="88">Hind wings in the only known species fully developed.</paragraph>
<paragraph id="8BF14E46FFA75057FF73FA4FFB4641B2" blockId="87.[151,1437,151,1761]" pageId="87" pageNumber="88">Abdomen subtriangular, slightly shorter than metaventrite, with six exposed sternites (sternites IIIVIII), division between VII and VIII slightly less distinct than between the remaining segments.</paragraph>
<paragraph id="8BF14E46FFA75057FF73FA07FCCD427A" blockId="87.[151,1437,151,1761]" pageId="87" pageNumber="88">
Aedeagus (
<figureCitation id="137552C3FFA75057FEF1FA07FE274196" box="[325,482,1483,1509]" captionStart="FIGURES 324331" captionStartId="88.[151,264,1672,1696]" captionTargetBox="[151,1419,322,1642]" captionTargetId="figure-77@88.[151,1435,322,1649]" captionTargetPageId="88" captionText="FIGURES 324331. Contreuconnus ambatolampyensis (Franz), holotype. Habitus in dorsal view (324); head in anterodorsal view (325); head and prothorax in lateral view (326); antenna (327); aedeagus in ventral (328, 330) and lateral (329, 331) views." figureDoi="http://doi.org/10.5281/zenodo.13746049" httpUri="https://zenodo.org/record/13746049/files/figure.png" pageId="87" pageNumber="88">Figs 328331</figureCitation>
,
<figureCitation id="137552C3FFA75057FE5AFA07FD974196" box="[494,594,1483,1509]" captionStart="FIGURES 335342" captionStartId="90.[151,264,1484,1508]" captionTargetBox="[151,1425,181,1446]" captionTargetId="figure-28@90.[151,1435,181,1461]" captionTargetPageId="90" captionText="FIGURES 335342. Contreuconnus ambatolampyensis (Franz), holotype of Euconnus (Anthicimorphus) angustus Franz. Habitus in dorsal view (335); head in anterodorsal view (336); head and prothorax in lateral view (337); antenna (338); aedeagus in ventral (339, 341) and lateral (340, 342) views." figureDoi="http://doi.org/10.5281/zenodo.13746057" httpUri="https://zenodo.org/record/13746057/files/figure.png" pageId="87" pageNumber="88">339342</figureCitation>
) with symmetrical median lobe and strongly sclerotized, asymmetrical and complex endophallus, and with free, slender parameres.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFA75057FF73F9DFFC834206" pageId="87" pageNumber="88" type="etymology">
<paragraph id="8BF14E46FFA75057FF73F9DFFAD1425E" blockId="87.[151,1437,151,1761]" box="[199,1300,1555,1581]" pageId="87" pageNumber="88">
<emphasis id="B93A9254FFA75057FF73F9DFFE8B425E" bold="true" box="[199,334,1555,1581]" pageId="87" pageNumber="88">Etymology.</emphasis>
<taxonomicName id="4C4E35C5FFA75057FEE1F9DFFE3A425E" authorityName="Jałoszyński" authorityYear="2024" box="[341,511,1555,1581]" class="Insecta" family="Staphylinidae" genus="Contreuconnus" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="genus">
<emphasis id="B93A9254FFA75057FEE1F9DFFE3A425E" box="[341,511,1555,1581]" italics="true" pageId="87" pageNumber="88">Contreuconnus</emphasis>
</taxonomicName>
combines
<emphasis id="B93A9254FFA75057FDCCF9DFFD04425F" box="[632,705,1555,1580]" italics="true" pageId="87" pageNumber="88">contra</emphasis>
(Latin “against”) with
<taxonomicName id="4C4E35C5FFA75057FC71F9DFFBF6425F" box="[965,1075,1555,1580]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="genus">
<emphasis id="B93A9254FFA75057FC71F9DFFBF6425F" box="[965,1075,1555,1580]" italics="true" pageId="87" pageNumber="88">Euconnus</emphasis>
</taxonomicName>
. Gender masculine.
</paragraph>
<paragraph id="8BF14E46FFA75057FF73F9FAFC834206" blockId="87.[151,1437,151,1761]" pageId="87" pageNumber="88">
<emphasis id="B93A9254FFA75057FF73F9FAFDFE4223" bold="true" box="[199,571,1590,1617]" pageId="87" pageNumber="88">Composition and distribution.</emphasis>
<taxonomicName id="4C4E35C5FFA75057FDF3F9FBFD344222" authorityName="Jałoszyński" authorityYear="2024" box="[583,753,1591,1617]" class="Insecta" family="Staphylinidae" genus="Contreuconnus" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="genus">
<emphasis id="B93A9254FFA75057FDF3F9FBFD344222" box="[583,753,1591,1617]" italics="true" pageId="87" pageNumber="88">Contreuconnus</emphasis>
</taxonomicName>
includes one nominal species known to occur in Central
<collectingCountry id="F3590ED6FFA75057FF23F997FEDB4206" box="[151,286,1627,1653]" name="Madagascar" pageId="87" pageNumber="88">Madagascar</collectingCountry>
(Alaotra-Mangoro and Vakinankaratra Regions).
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFA75057FF73F9B3FC7F4292" pageId="87" pageNumber="88" type="discussion">
<paragraph id="8BF14E46FFA75057FF73F9B3FC7F4292" blockId="87.[151,1437,151,1761]" pageId="87" pageNumber="88">
<emphasis id="B93A9254FFA75057FF73F9B3FEFE42EA" bold="true" box="[199,315,1663,1689]" pageId="87" pageNumber="88">Remarks.</emphasis>
<taxonomicName id="4C4E35C5FFA75057FEF1F9B3FE2A42EA" authorityName="Jałoszyński" authorityYear="2024" box="[325,495,1663,1689]" class="Insecta" family="Staphylinidae" genus="Contreuconnus" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="genus">
<emphasis id="B93A9254FFA75057FEF1F9B3FE2A42EA" box="[325,495,1663,1689]" italics="true" pageId="87" pageNumber="88">Contreuconnus</emphasis>
</taxonomicName>
was included in the key to world genera of Stenichnini (
<bibRefCitation id="EFDF33B7FFA75057FB31F9B3FA8E42EA" author="Jaloszynski, P." box="[1157,1355,1663,1689]" pageId="87" pageNumber="88" pagination="549 - 570" refId="ref55749" refString="Jaloszynski, P. (2024) Palposcydmus gen. n. from India, with a preliminary key to world genera of Stenichnini (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 5428 (4), 549 - 570. https: // doi. org / 10.11646 / zootaxa. 5428.4.5" type="journal article" year="2024">Jałoszyński 2024</bibRefCitation>
) under the code name Mdg genus 2. It is most similar to
<taxonomicName id="4C4E35C5FFA75057FD6FF96FFC8C42CF" box="[731,841,1699,1724]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="genus">
<emphasis id="B93A9254FFA75057FD6FF96FFC8C42CF" box="[731,841,1699,1724]" italics="true" pageId="87" pageNumber="88">Euconnus</emphasis>
</taxonomicName>
, but does not have the highly elevated and complete mesoventral process that in
<taxonomicName id="4C4E35C5FFA75057FE64F90BFDFB4293" box="[464,574,1735,1760]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="genus">
<emphasis id="B93A9254FFA75057FE64F90BFDFB4293" box="[464,574,1735,1760]" italics="true" pageId="87" pageNumber="88">Euconnus</emphasis>
</taxonomicName>
completely separates mesocoxae.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,317 @@
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<subSubSection id="C3541DCDFFBF504FFF23FE5FFCFE45DE" box="[151,827,402,429]" pageId="79" pageNumber="80" type="nomenclature">
<paragraph id="8BF14E46FFBF504FFF23FE5FFCFE45DE" blockId="79.[151,827,402,465]" box="[151,827,402,429]" pageId="79" pageNumber="80">
<heading id="D0B9F92AFFBF504FFF23FE5FFCFE45DE" bold="true" box="[151,827,402,429]" fontSize="11" level="2" pageId="79" pageNumber="80" reason="4">
<emphasis id="B93A9254FFBF504FFF23FE5FFCFE45DE" bold="true" box="[151,827,402,429]" pageId="79" pageNumber="80">
<taxonomicName id="4C4E35C5FFBF504FFF23FE5FFD0645DF" authority="(Lhoste)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[151,707,402,429]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="mauritiensis" status="comb. nov." subGenus="Tetramelus">
<emphasis id="B93A9254FFBF504FFF23FE5FFED645DE" bold="true" box="[151,275,403,429]" italics="true" pageId="79" pageNumber="80">Euconnus</emphasis>
(
<emphasis id="B93A9254FFBF504FFE90FE5EFE6F45DE" bold="true" box="[292,426,402,429]" italics="true" pageId="79" pageNumber="80">Tetramelus</emphasis>
)
<emphasis id="B93A9254FFBF504FFE0FFE5FFD9445DE" bold="true" box="[443,593,403,429]" italics="true" pageId="79" pageNumber="80">mauritiensis</emphasis>
(Lhoste)
</taxonomicName>
,
<taxonomicNameLabel id="A2092F2FFFBF504FFD64FE5FFCFE45DE" box="[720,827,403,429]" pageId="79" pageNumber="80" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFBF504FFF23FE7BFE8645A2" box="[151,323,438,465]" pageId="79" pageNumber="80" type="description">
<paragraph id="8BF14E46FFBF504FFF23FE7BFE8645A2" blockId="79.[151,827,402,465]" box="[151,323,438,465]" pageId="79" pageNumber="80">
(
<figureCitation id="137552C3FFBF504FFF2BFE7BFEF945A2" box="[159,316,438,465]" captionStart="FIGURES 304311" captionStartId="80.[151,264,1602,1626]" captionTargetBox="[164,1435,181,1568]" captionTargetId="figure-28@80.[151,1435,181,1578]" captionTargetPageId="80" captionText="FIGURES 304311. Euconnus (Tetramelus) mauritiensis (Lhoste), reproduced from Lhoste (1956) (304305), and Horaeomorphus sp. mislabeled with original labels for syntypes of Scydmaenus mauritiensis Lhoste (307311). Habitus in dorsal view (304); antenna (305); aedeagus in dorsal view (306); dorsal habitus of female from NHMW (307); aedeagus of a specimen from NHMUK in ventral (308, 310) and lateral (309, 311) views." figureDoi="http://doi.org/10.5281/zenodo.13746039" httpUri="https://zenodo.org/record/13746039/files/figure.png" pageId="79" pageNumber="80">Figs 304305</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFBF504FFF23FE37FD0B4641" pageId="79" pageNumber="80" type="reference_group">
<paragraph id="8BF14E46FFBF504FFF23FE37FD234660" blockId="79.[151,742,507,562]" box="[151,742,507,531]" pageId="79" pageNumber="80">
<treatmentCitationGroup id="AB5E6968FFBF504FFF23FE37FD234660" box="[151,742,507,531]" pageId="79" pageNumber="80">
<treatmentCitation id="0AEF6857FFBF504FFF23FE37FD274660" author="Lhoste, J." box="[151,738,507,531]" page="284" pageId="79" pageNumber="80" year="1956">
<taxonomicName id="4C4E35C5FFBF504FFF23FE37FD274660" ID-CoL="6XZVK" authority="Lhoste, 1956: 284" authorityName="Lhoste" authorityPageNumber="284" authorityYear="1956" box="[151,738,507,531]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="mauritiensis" subGenus="Vinsoniana">
<emphasis id="B93A9254FFBF504FFF23FE37FED14660" box="[151,276,507,531]" italics="true" pageId="79" pageNumber="80">Scydmaenus</emphasis>
(
<emphasis id="B93A9254FFBF504FFE97FE30FE504660" box="[291,405,508,531]" italics="true" pageId="79" pageNumber="80">Vinsoniana</emphasis>
)
<emphasis id="B93A9254FFBF504FFE17FE30FDE54660" box="[419,544,508,531]" italics="true" pageId="79" pageNumber="80">mauritiensis</emphasis>
<bibRefCitation id="EFDF33B7FFBF504FFD93FE37FD274660" author="Lhoste, J." box="[551,738,507,531]" pageId="79" pageNumber="80" pagination="283 - 286" refId="ref56286" refString="Lhoste, J. (1956) Sur un sous-genre nouveau de Scydmaenus Latreille, originaire de l'Ile Maurice (Col. Scydmaenidae). Mauritius Institute Bulletin, 3, 283 - 286." type="journal article" year="1956">Lhoste, 1956: 284</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF14E46FFBF504FFF23FDD6FD0B4641" blockId="79.[151,742,507,562]" box="[151,718,538,562]" pageId="79" pageNumber="80">
<treatmentCitationGroup id="AB5E6968FFBF504FFF23FDD6FD0B4641" box="[151,718,538,562]" pageId="79" pageNumber="80">
<taxonomicName id="4C4E35C5FFBF504FFF23FDD6FDE54641" ID-CoL="3MNKP" authority="(Lhoste)" baseAuthorityName="Lhoste" baseAuthorityYear="1956" box="[151,544,538,562]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="mauritiensis">
<emphasis id="B93A9254FFBF504FFF23FDD6FE044642" box="[151,449,538,562]" italics="true" pageId="79" pageNumber="80">Horaeomorphus mauritiensis</emphasis>
(Lhoste)
</taxonomicName>
;
<treatmentCitation id="0AEF6857FFBF504FFD9EFDD6FD0F4641" author="Franz, H." box="[554,714,538,562]" page="6" pageId="79" pageNumber="80" year="1972">
<bibRefCitation id="EFDF33B7FFBF504FFD9EFDD6FD774641" author="Franz, H." box="[554,690,538,562]" pageId="79" pageNumber="80" pagination="3 - 14" refId="ref53178" refString="Franz, H. (1972) Die Scydmaenidenfauna der Insel Mauritius. Koleopterologische Rundschau, 50, 3 - 14." type="journal article" year="1972">Franz (1972)</bibRefCitation>
: 6
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFBF5051FF23FDACFCD5448B" lastPageId="81" lastPageNumber="82" pageId="79" pageNumber="80" type="discussion">
<paragraph id="8BF14E46FFBF504FFF23FDACFE024790" blockId="79.[151,1437,608,2002]" pageId="79" pageNumber="80">
<emphasis id="B93A9254FFBF504FFF23FDACFECE4609" bold="true" box="[151,267,608,634]" pageId="79" pageNumber="80">Remarks.</emphasis>
The history of misidentification of this species is complicated and difficult to understand, and perfectly exemplifies the
<taxonomicName id="4C4E35C5FFBF504FFEE1FD48FDC846ED" box="[341,525,644,670]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="genus">
<emphasis id="B93A9254FFBF504FFEE1FD48FDC846ED" box="[341,525,644,670]" italics="true" pageId="79" pageNumber="80">Horaeomorphus</emphasis>
</taxonomicName>
curse.
<bibRefCitation id="EFDF33B7FFBF504FFDDCFD48FCC346EC" author="Lhoste, J." box="[616,774,644,671]" pageId="79" pageNumber="80" pagination="283 - 286" refId="ref56286" refString="Lhoste, J. (1956) Sur un sous-genre nouveau de Scydmaenus Latreille, originaire de l'Ile Maurice (Col. Scydmaenidae). Mauritius Institute Bulletin, 3, 283 - 286." type="journal article" year="1956">Lhoste (1956)</bibRefCitation>
described
<taxonomicName id="4C4E35C5FFBF504FFC36FD48FB5B46ED" authorityName="Lhoste" authorityYear="1956" box="[898,1182,644,670]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="mauritiensis">
<emphasis id="B93A9254FFBF504FFC36FD48FB5B46ED" box="[898,1182,644,670]" italics="true" pageId="79" pageNumber="80">Scydmaenus mauritiensis</emphasis>
</taxonomicName>
based on
<specimenCount id="9D4885CFFFBF504FFAA6FD48FA5946EC" box="[1298,1436,644,671]" count="6" pageId="79" pageNumber="80" type="generic" typeStatus="syntype">six syntypes</specimenCount>
collected on
<date id="FFF06886FFBF504FFE98FD64FE6F46B0" box="[300,426,680,707]" pageId="79" pageNumber="80" value="1954-10-31">31.10.1954</date>
by J. Vinson on Mt. Cocotte,
<collectingCountry id="F3590ED6FFBF504FFCBBFD64FC4546B1" box="[783,896,680,706]" name="Mauritius" pageId="79" pageNumber="80">Mauritius</collectingCountry>
. His illustrations (figs 13, reproduced here in
<figureCitation id="137552C3FFBF504FFF23FD00FEF64695" box="[151,307,716,742]" captionStart="FIGURES 304311" captionStartId="80.[151,264,1602,1626]" captionTargetBox="[164,1435,181,1568]" captionTargetId="figure-28@80.[151,1435,181,1578]" captionTargetPageId="80" captionText="FIGURES 304311. Euconnus (Tetramelus) mauritiensis (Lhoste), reproduced from Lhoste (1956) (304305), and Horaeomorphus sp. mislabeled with original labels for syntypes of Scydmaenus mauritiensis Lhoste (307311). Habitus in dorsal view (304); antenna (305); aedeagus in dorsal view (306); dorsal habitus of female from NHMW (307); aedeagus of a specimen from NHMUK in ventral (308, 310) and lateral (309, 311) views." figureDoi="http://doi.org/10.5281/zenodo.13746039" httpUri="https://zenodo.org/record/13746039/files/figure.png" pageId="79" pageNumber="80">Figs 304306</figureCitation>
) clearly show a stenichnine beetle (not Scydmaenini!) with an elongate pronotum broadest in front of base and bearing two pairs of antebasal pits, antennae with elongate proximal flagellomeres and distinctly delimited tetramerous clubs, and the aedeagus, which was not completely dissected from surrounding structures, because the illustrated dorsal wall with a transverse row of setae is most likely the tergite X, showed as it was an integral part of the median lobe. All hitherto known species of
<taxonomicName id="4C4E35C5FFBF504FFCFFFC90FBC64705" box="[843,1027,860,886]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="genus">
<emphasis id="B93A9254FFBF504FFCFFFC90FBC64705" box="[843,1027,860,886]" italics="true" pageId="79" pageNumber="80">Horaeomorphus</emphasis>
</taxonomicName>
have an uneven number of antebasal pronotal pits (three or five) and gradually broadening antennae, and for these reasons this species cannot belong in
<taxonomicName id="4C4E35C5FFBF504FFF02FC68FEAB47CD" box="[182,366,932,958]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="genus">
<emphasis id="B93A9254FFBF504FFF02FC68FEAB47CD" box="[182,366,932,958]" italics="true" pageId="79" pageNumber="80">Horaeomorphus</emphasis>
</taxonomicName>
. The illustrated aedeagus is also clearly different from the form of the male copulatory organ typical of
<taxonomicName id="4C4E35C5FFBF504FFEBDFC04FE044791" box="[265,449,968,994]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="genus">
<emphasis id="B93A9254FFBF504FFEBDFC04FE044791" box="[265,449,968,994]" italics="true" pageId="79" pageNumber="80">Horaeomorphus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF14E46FFBF504FFF73FC20FA4141C5" blockId="79.[151,1437,608,2002]" pageId="79" pageNumber="80">
Despite these features,
<bibRefCitation id="EFDF33B7FFBF504FFE68FC21FDB64075" author="Franz, H." box="[476,627,1004,1031]" pageId="79" pageNumber="80" pagination="3 - 14" refId="ref53178" refString="Franz, H. (1972) Die Scydmaenidenfauna der Insel Mauritius. Koleopterologische Rundschau, 50, 3 - 14." type="journal article" year="1972">Franz (1972)</bibRefCitation>
transferred
<taxonomicName id="4C4E35C5FFBF504FFCB2FC20FB454075" authority="Lhoste" authorityName="Lhoste" authorityYear="1956" box="[774,1152,1004,1030]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="mauritiensis">
<emphasis id="B93A9254FFBF504FFCB2FC20FBE24075" box="[774,1063,1004,1030]" italics="true" pageId="79" pageNumber="80">Scydmaenus mauritiensis</emphasis>
Lhoste
</taxonomicName>
to
<taxonomicName id="4C4E35C5FFBF504FFB04FC20FAAD4075" box="[1200,1384,1004,1030]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="genus">
<emphasis id="B93A9254FFBF504FFB04FC20FAAD4075" box="[1200,1384,1004,1030]" italics="true" pageId="79" pageNumber="80">Horaeomorphus</emphasis>
</taxonomicName>
and redescribed it. He mentions “4 leider durchwegs sehr stark beschädigte Exemplare (
<specimenCount id="9D4885CFFFBF504FFBD5FBDCFB424059" box="[1121,1159,1040,1066]" count="1" pageId="79" pageNumber="80" type="female">♀♀</specimenCount>
)” (
<emphasis id="B93A9254FFBF504FFB05FBDDFB114059" box="[1201,1236,1041,1066]" italics="true" pageId="79" pageNumber="80">i.e.</emphasis>
, “4 unfortunately very badly damaged specimens (
<specimenCount id="9D4885CFFFBF504FFDBFFBF8FDF4403D" box="[523,561,1076,1102]" count="1" pageId="79" pageNumber="80" type="female">♀♀</specimenCount>
)”) examined for his redescription, collected by J. Vinson on
<date id="FFF06886FFBF504FFB4EFBF8FABD403C" box="[1274,1400,1076,1103]" pageId="79" pageNumber="80" value="1954-10-31">31.10.1954</date>
on Mt. Cocotte. Franz did not disclose a source or depository of these specimens. His redescription fully agrees with the original description, and the aedeagus is not illustrated, but described in reference to the Lhostes figure. Later,
<bibRefCitation id="EFDF33B7FFBF504FFF23FB6DFEFD40C9" author="Franz, H." box="[151,312,1184,1211]" pageId="79" pageNumber="80" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986b)</bibRefCitation>
again treated this species, but this time he stated that the type specimens were not found neither in Muséum National d'Histoire Naturelle, Paris nor in Natural History Museum, London. Interestingly, he mentions that he had received
<specimenCount id="9D4885CFFFBF504FFE33FB25FDE64171" box="[391,547,1256,1283]" count="1" pageId="79" pageNumber="80" type="generic">one specimen</specimenCount>
from L. Gomy, which belonged in the original type series. This specimen, of an unspecified sex, according to
<bibRefCitation id="EFDF33B7FFBF504FFDBEFAC1FD6F4155" author="Franz, H." box="[522,682,1292,1319]" pageId="79" pageNumber="80" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986b)</bibRefCitation>
bears a label “
<collectingCountry id="F3590ED6FFBF504FFCE5FAC0FC044155" box="[849,961,1292,1318]" name="Mauritius" pageId="79" pageNumber="80">Mauritius</collectingCountry>
Mt. Cocotte
<date id="FFF06886FFBF504FFBE2FAC0FB084154" box="[1110,1229,1292,1319]" pageId="79" pageNumber="80" value="1954-10-31">31.X.1954</date>
J. Vinson”, which indeed agrees with the
<bibRefCitation id="EFDF33B7FFBF504FFE28FAFCFD8B4138" author="Lhoste, J." box="[412,590,1328,1355]" pageId="79" pageNumber="80" pagination="283 - 286" refId="ref56286" refString="Lhoste, J. (1956) Sur un sous-genre nouveau de Scydmaenus Latreille, originaire de l'Ile Maurice (Col. Scydmaenidae). Mauritius Institute Bulletin, 3, 283 - 286." type="journal article" year="1956">Lhostes (1956)</bibRefCitation>
description. Moreover,
<bibRefCitation id="EFDF33B7FFBF504FFCE9FAFDFC384139" author="Franz, H." box="[861,1021,1328,1355]" pageId="79" pageNumber="80" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986b)</bibRefCitation>
designated this single specimen as a
<typeStatus id="54F5F0E4FFBF504FFF23FA98FF3B411D" box="[151,254,1364,1390]" pageId="79" pageNumber="80" type="lectotype">lectotype</typeStatus>
and deposited it in his private collection. However, he again redescribes this species, as
<taxonomicName id="4C4E35C5FFBF504FFB51FA98FEE441E1" baseAuthorityName="Lhoste" baseAuthorityYear="1956" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="mauritiensis">
<emphasis id="B93A9254FFBF504FFB51FA98FEE441E1" italics="true" pageId="79" pageNumber="80">Horaeomorphus mauritiensis</emphasis>
</taxonomicName>
, but not using the
<typeStatus id="54F5F0E4FFBF504FFDB7FAB4FDAF41E1" box="[515,618,1400,1426]" pageId="79" pageNumber="80" type="lectotype">lectotype</typeStatus>
for this purpose. Instead, he simply cites verbatim the entire Lhostes description in French and reproduces the Lhostes illustrations (
<bibRefCitation id="EFDF33B7FFBF504FFCEBFA50FC3741C5" author="Franz, H." box="[863,1010,1436,1462]" pageId="79" pageNumber="80" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz 1986b</bibRefCitation>
), as if no specimens were available.
</paragraph>
<paragraph id="8BF14E46FFBF504FFF73FA0CFD3C436D" blockId="79.[151,1437,608,2002]" pageId="79" pageNumber="80">
The lectotype in Franz Coll., currently at NHMW, indeed bears the possibly original label cited by
<bibRefCitation id="EFDF33B7FFBF504FFAE8FA0DFF37418D" author="Franz, H." pageId="79" pageNumber="80" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986b)</bibRefCitation>
, and a red handwritten “
<typeStatus id="54F5F0E4FFBF504FFDB8FA28FD4E418D" box="[524,651,1508,1534]" pageId="79" pageNumber="80" type="lectotype">Lectotypus</typeStatus>
” label. However, this specimen, illustrated here in
<figureCitation id="137552C3FFBF504FFB66FA28FAF1418D" box="[1234,1332,1508,1534]" captionStart="FIGURES 304311" captionStartId="80.[151,264,1602,1626]" captionTargetBox="[164,1435,181,1568]" captionTargetId="figure-28@80.[151,1435,181,1578]" captionTargetPageId="80" captionText="FIGURES 304311. Euconnus (Tetramelus) mauritiensis (Lhoste), reproduced from Lhoste (1956) (304305), and Horaeomorphus sp. mislabeled with original labels for syntypes of Scydmaenus mauritiensis Lhoste (307311). Habitus in dorsal view (304); antenna (305); aedeagus in dorsal view (306); dorsal habitus of female from NHMW (307); aedeagus of a specimen from NHMUK in ventral (308, 310) and lateral (309, 311) views." figureDoi="http://doi.org/10.5281/zenodo.13746039" httpUri="https://zenodo.org/record/13746039/files/figure.png" pageId="79" pageNumber="80">Fig. 307</figureCitation>
, is a true
<taxonomicName id="4C4E35C5FFBF504FFF23F9C4FE8A4251" box="[151,335,1544,1570]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="genus">
<emphasis id="B93A9254FFBF504FFF23F9C4FE8A4251" box="[151,335,1544,1570]" italics="true" pageId="79" pageNumber="80">Horaeomorphus</emphasis>
</taxonomicName>
(female) with three antebasal pronotal pits. Without doubt, this specimen is different from what
<bibRefCitation id="EFDF33B7FFBF504FFF23F9E0FEFF4234" author="Lhoste, J." box="[151,314,1580,1607]" pageId="79" pageNumber="80" pagination="283 - 286" refId="ref56286" refString="Lhoste, J. (1956) Sur un sous-genre nouveau de Scydmaenus Latreille, originaire de l'Ile Maurice (Col. Scydmaenidae). Mauritius Institute Bulletin, 3, 283 - 286." type="journal article" year="1956">Lhoste (1956)</bibRefCitation>
illustrated, and what
<bibRefCitation id="EFDF33B7FFBF504FFD8BF9E1FD0B4235" author="Franz, H." box="[575,718,1580,1607]" pageId="79" pageNumber="80" pagination="3 - 14" refId="ref53178" refString="Franz, H. (1972) Die Scydmaenidenfauna der Insel Mauritius. Koleopterologische Rundschau, 50, 3 - 14." type="journal article" year="1972">Franz (1972</bibRefCitation>
,
<bibRefCitation id="EFDF33B7FFBF504FFD6AF9E0FCE04235" author="Franz, H." box="[734,805,1580,1606]" pageId="79" pageNumber="80" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">1986b</bibRefCitation>
) twice redescribed. Furthermore, in Natural History Museum, London, there are two “unfortunately very badly damaged specimens” bearing the same original Vinsons labels, both with only partly preserved remains of very likely the same true
<taxonomicName id="4C4E35C5FFBF504FFC56F9B8FB5F42FD" box="[994,1178,1652,1678]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="genus">
<emphasis id="B93A9254FFBF504FFC56F9B8FB5F42FD" box="[994,1178,1652,1678]" italics="true" pageId="79" pageNumber="80">Horaeomorphus</emphasis>
</taxonomicName>
as the lectotype. One of them is a female lacking the head, the other one is a pterothorax with only one hind leg, elytra with the scutellar shield and abdomen of a male. I dissected the abdomen, and the aedeagus (
<figureCitation id="137552C3FFBF504FFC45F970FB4842A5" box="[1009,1165,1724,1750]" captionStart="FIGURES 304311" captionStartId="80.[151,264,1602,1626]" captionTargetBox="[164,1435,181,1568]" captionTargetId="figure-28@80.[151,1435,181,1578]" captionTargetPageId="80" captionText="FIGURES 304311. Euconnus (Tetramelus) mauritiensis (Lhoste), reproduced from Lhoste (1956) (304305), and Horaeomorphus sp. mislabeled with original labels for syntypes of Scydmaenus mauritiensis Lhoste (307311). Habitus in dorsal view (304); antenna (305); aedeagus in dorsal view (306); dorsal habitus of female from NHMW (307); aedeagus of a specimen from NHMUK in ventral (308, 310) and lateral (309, 311) views." figureDoi="http://doi.org/10.5281/zenodo.13746039" httpUri="https://zenodo.org/record/13746039/files/figure.png" pageId="79" pageNumber="80">Figs 308311</figureCitation>
) can be unambiguously recognized as representing a morphological form typical of
<taxonomicName id="4C4E35C5FFBF504FFCE1F92CFBC84289" box="[853,1037,1760,1786]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="genus">
<emphasis id="B93A9254FFBF504FFCE1F92CFBC84289" box="[853,1037,1760,1786]" italics="true" pageId="79" pageNumber="80">Horaeomorphus</emphasis>
</taxonomicName>
, and it belongs to an undescribed species, very likely of a SE Asian or Australian origin.
</paragraph>
<paragraph id="8BF14E46FFBF504FFF73F8E5FED243A1" blockId="79.[151,1437,608,2002]" pageId="79" pageNumber="80">
<materialsCitation id="3B26441BFFBF504FFF73F8E5FED643A1" collectionCode="NHMW" pageId="79" pageNumber="80" specimenCount="1" typeStatus="lectotype">
To sum up, none of the specimens with the possibly original Vinsons labels deposited at
<collectionCode id="ED5FD683FFBF504FFB29F8E5FB3D4330" box="[1181,1272,1833,1859]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="79" pageNumber="80" type="Museum">NHMW</collectionCode>
and in Natural History Museum, London is congeneric with
<taxonomicName id="4C4E35C5FFBF504FFD3DF880FC3D4315" authority="Lhoste" authorityName="Lhoste" authorityYear="1956" box="[649,1016,1868,1894]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="mauritiensis">
<emphasis id="B93A9254FFBF504FFD3DF880FC664315" box="[649,931,1868,1894]" italics="true" pageId="79" pageNumber="80">Scydmaenus mauritiensis</emphasis>
Lhoste
</taxonomicName>
, and the
<typeStatus id="54F5F0E4FFBF504FFBE2F880FB784315" box="[1110,1213,1868,1894]" pageId="79" pageNumber="80" type="lectotype">lectotype</typeStatus>
designation was not valid, because it was based on a misidentified specimen. It seems clear that these specimens have been mislabeled, but I was unable to figure out what had happened with the original Vinsons specimens used by Lhoste to describe his species
</materialsCitation>
.
</paragraph>
<caption id="DF311ECEFFA05050FF23F98EFC5642B5" ID-DOI="http://doi.org/10.5281/zenodo.13746039" ID-Zenodo-Dep="13746039" httpUri="https://zenodo.org/record/13746039/files/figure.png" pageId="80" pageNumber="81" startId="80.[151,264,1602,1626]" targetBox="[164,1435,181,1568]" targetPageId="80" targetType="figure">
<paragraph id="8BF14E46FFA05050FF23F98EFC5642B5" blockId="80.[151,1437,1602,1734]" pageId="80" pageNumber="81">
<emphasis id="B93A9254FFA05050FF23F98EFEB34229" bold="true" box="[151,374,1602,1626]" pageId="80" pageNumber="81">FIGURES 304311.</emphasis>
<taxonomicName id="4C4E35C5FFA05050FE33F98FFCB64229" authority="(Lhoste)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[391,883,1602,1626]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="mauritiensis" subGenus="Tetramelus">
<emphasis id="B93A9254FFA05050FE33F98FFE2E4229" box="[391,491,1603,1626]" italics="true" pageId="80" pageNumber="81">Euconnus</emphasis>
(
<emphasis id="B93A9254FFA05050FDB0F98EFDB14229" box="[516,628,1602,1626]" italics="true" pageId="80" pageNumber="81">Tetramelus</emphasis>
)
<emphasis id="B93A9254FFA05050FD39F98FFCCF4229" box="[653,778,1603,1626]" italics="true" pageId="80" pageNumber="81">mauritiensis</emphasis>
(Lhoste)
</taxonomicName>
, reproduced from
<bibRefCitation id="EFDF33B7FFA05050FBF9F98EFB234229" author="Lhoste, J." box="[1101,1254,1602,1626]" pageId="80" pageNumber="81" pagination="283 - 286" refId="ref56286" refString="Lhoste, J. (1956) Sur un sous-genre nouveau de Scydmaenus Latreille, originaire de l'Ile Maurice (Col. Scydmaenidae). Mauritius Institute Bulletin, 3, 283 - 286." type="journal article" year="1956">Lhoste (1956)</bibRefCitation>
(304305), and
<taxonomicName id="4C4E35C5FFA05050FF23F9AAFEA1420D" box="[151,356,1638,1662]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B93A9254FFA05050FF23F9AAFEF8420D" box="[151,317,1638,1662]" italics="true" pageId="80" pageNumber="81">Horaeomorphus</emphasis>
sp.
</taxonomicName>
mislabeled with original labels for syntypes of
<taxonomicName id="4C4E35C5FFA05050FCE9F9AAFB6A420D" authority="Lhoste" authorityName="Lhoste" authorityYear="1956" box="[861,1199,1638,1662]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="mauritiensis">
<emphasis id="B93A9254FFA05050FCE9F9AAFBA4420D" box="[861,1121,1638,1662]" italics="true" pageId="80" pageNumber="81">Scydmaenus mauritiensis</emphasis>
Lhoste
</taxonomicName>
(307311). Habitus in dorsal view (304); antenna (305); aedeagus in dorsal view (306); dorsal habitus of female from NHMW (307); aedeagus of a specimen from NHMUK in ventral (308, 310) and lateral (309, 311) views.
</paragraph>
</caption>
<paragraph id="8BF14E46FFA05050FF73F8D4FE6643B0" blockId="80.[151,1436,1816,1987]" pageId="80" pageNumber="81">
The aedeagus of
<taxonomicName id="4C4E35C5FFA05050FE23F8D4FD7D4341" authorityName="Lhoste" authorityYear="1956" box="[407,696,1816,1842]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="mauritiensis">
<emphasis id="B93A9254FFA05050FE23F8D4FD7D4341" box="[407,696,1816,1842]" italics="true" pageId="80" pageNumber="81">Scydmaenus mauritiensis</emphasis>
</taxonomicName>
(
<figureCitation id="137552C3FFA05050FD79F8D4FCF64341" box="[717,819,1816,1842]" captionStart="FIGURES 304311" captionStartId="80.[151,264,1602,1626]" captionTargetBox="[164,1435,181,1568]" captionTargetId="figure-28@80.[151,1435,181,1578]" captionTargetPageId="80" captionText="FIGURES 304311. Euconnus (Tetramelus) mauritiensis (Lhoste), reproduced from Lhoste (1956) (304305), and Horaeomorphus sp. mislabeled with original labels for syntypes of Scydmaenus mauritiensis Lhoste (307311). Habitus in dorsal view (304); antenna (305); aedeagus in dorsal view (306); dorsal habitus of female from NHMW (307); aedeagus of a specimen from NHMUK in ventral (308, 310) and lateral (309, 311) views." figureDoi="http://doi.org/10.5281/zenodo.13746039" httpUri="https://zenodo.org/record/13746039/files/figure.png" pageId="80" pageNumber="81">Fig. 306</figureCitation>
) is highly similar to that of
<taxonomicName id="4C4E35C5FFA05050FB27F8D5FA504341" authorityName="Franz" authorityYear="1986" box="[1171,1429,1816,1842]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="andasyensis">
<emphasis id="B93A9254FFA05050FB27F8D5FA504341" box="[1171,1429,1816,1842]" italics="true" pageId="80" pageNumber="81">Euconnus andasyensis</emphasis>
</taxonomicName>
, and although Lhoste illustrated the endophallus as symmetrical, while all Malagasy species of
<taxonomicName id="4C4E35C5FFA05050FB56F8F0FA984325" box="[1250,1373,1852,1878]" pageId="80" pageNumber="52" rank="subGenus" subGenus="Tetramelus">
<emphasis id="B93A9254FFA05050FB56F8F0FA984325" box="[1250,1373,1852,1878]" italics="true" pageId="80" pageNumber="81">Tetramelus</emphasis>
</taxonomicName>
have asymmetrical endophallic sclerites, it is possible that the dark structures in his figure represent the combined most darkly sclerotized median proximal and distolateral endophallic structures, similar to those in
<taxonomicName id="4C4E35C5FFA05050FB1EF849FA5943ED" authorityName="Franz" authorityYear="1986" box="[1194,1436,1924,1950]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="matsaboryi">
<emphasis id="B93A9254FFA05050FB1EF849FA5943ED" box="[1194,1436,1924,1950]" italics="true" pageId="80" pageNumber="81">Euconnus matsaboryi</emphasis>
</taxonomicName>
(
<figureCitation id="137552C3FFA05050FF2BF864FF3A43B1" box="[159,255,1960,1986]" captionStart="FIGURES 264271" captionStartId="70.[151,264,1875,1899]" captionTargetBox="[151,1411,509,1836]" captionTargetId="figure-104@70.[151,1435,509,1852]" captionTargetPageId="70" captionText="FIGURES 264271. Euconnus (Tetramelus) matsaboryi Franz, holotype. Habitus in dorsal view (264); head in anterodorsal view (265); head and prothorax in lateral view (only bristles on head are shown) (266); antenna (267); aedeagus in ventral (268, 270) and lateral (269, 271) views." figureDoi="http://doi.org/10.5281/zenodo.13746023" httpUri="https://zenodo.org/record/13746023/files/figure.png" pageId="80" pageNumber="81">Fig. 268</figureCitation>
) or
<taxonomicName id="4C4E35C5FFA05050FE99F865FE5843B1" authorityName="Franz" authorityYear="1986" box="[301,413,1960,1986]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="pellifer">
<emphasis id="B93A9254FFA05050FE99F865FE5843B1" box="[301,413,1960,1986]" italics="true" pageId="80" pageNumber="81">E. pellifer</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF14E46FFA15051FF73FF5AFCD5448B" blockId="81.[151,1437,150,249]" pageId="81" pageNumber="82">
<emphasis id="B93A9254FFA15051FF73FF5AFCD5448B" bold="true" pageId="81" pageNumber="82">
The transfer of
<taxonomicName id="4C4E35C5FFA15051FE33FF5AFC6C44C3" authority="Lhoste" authorityName="Lhoste" authorityYear="1956" box="[391,937,150,177]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="mauritiensis" subGenus="Vinsoniana">
<emphasis id="B93A9254FFA15051FE33FF5AFDDD44C2" bold="true" box="[391,536,150,177]" italics="true" pageId="81" pageNumber="82">Scydmaenus</emphasis>
(
<emphasis id="B93A9254FFA15051FD9EFF5AFD6B44C3" bold="true" box="[554,686,150,176]" italics="true" pageId="81" pageNumber="82">Vinsoniana</emphasis>
)
<emphasis id="B93A9254FFA15051FD74FF5AFC8A44C3" bold="true" box="[704,847,150,176]" italics="true" pageId="81" pageNumber="82">mauritiensis</emphasis>
Lhoste
</taxonomicName>
(
<taxonomicName id="4C4E35C5FFA15051FC70FF5AFADC44C3" baseAuthorityName="Lhoste" baseAuthorityYear="1956" box="[964,1305,150,177]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="mauritiensis">
<emphasis id="B93A9254FFA15051FC70FF5AFADC44C3" bold="true" box="[964,1305,150,177]" italics="true" pageId="81" pageNumber="82">Horaeomorphus mauritiensis</emphasis>
</taxonomicName>
of Franz) to
<taxonomicName id="4C4E35C5FFA15051FF0FFF77FE1544A7" authorityName="Motschulsky" authorityYear="1869" box="[187,464,186,213]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="subGenus" subGenus="Tetramelus">
<emphasis id="B93A9254FFA15051FF0FFF77FEF444A6" bold="true" box="[187,305,187,213]" italics="true" pageId="81" pageNumber="82">Euconnus</emphasis>
(
<emphasis id="B93A9254FFA15051FEF3FF76FE0244A6" bold="true" box="[327,455,186,213]" italics="true" pageId="81" pageNumber="82">Tetramelus</emphasis>
)
</taxonomicName>
affects the status of the name
<taxonomicName id="4C4E35C5FFA15051FCEAFF76FC2744A7" box="[862,994,186,212]" pageId="81" pageNumber="4" rank="subGenus" subGenus="Vinsoniana">
<emphasis id="B93A9254FFA15051FCEAFF76FC2744A7" bold="true" box="[862,994,186,212]" italics="true" pageId="81" pageNumber="82">Vinsoniana</emphasis>
</taxonomicName>
, which is not a junior synonym of
<taxonomicName id="4C4E35C5FFA15051FF23FF12FE90448A" box="[151,341,222,249]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="genus">
<emphasis id="B93A9254FFA15051FF23FF12FE90448A" bold="true" box="[151,341,222,249]" italics="true" pageId="81" pageNumber="82">Horaeomorphus</emphasis>
</taxonomicName>
, but a junior synonym of
<taxonomicName id="4C4E35C5FFA15051FD3DFF12FCCC448A" box="[649,777,222,249]" pageId="81" pageNumber="52" rank="subGenus" subGenus="Tetramelus">
<emphasis id="B93A9254FFA15051FD3DFF12FCCC448A" bold="true" box="[649,777,222,249]" italics="true" pageId="81" pageNumber="82">Tetramelus</emphasis>
</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,203 @@
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<subSubSection id="C3541DCDFFC05030FF23FF5BFD6744C2" box="[151,674,151,178]" pageId="48" pageNumber="49" type="nomenclature">
<paragraph id="8BF14E46FFC05030FF23FF5BFD6744C2" blockId="48.[151,674,151,213]" box="[151,674,151,178]" pageId="48" pageNumber="49">
<heading id="D0B9F92AFFC05030FF23FF5BFD6744C2" bold="true" box="[151,674,151,178]" fontSize="11" level="2" pageId="48" pageNumber="49" reason="4">
<emphasis id="B93A9254FFC05030FF23FF5BFD6744C2" bold="true" box="[151,674,151,178]" pageId="48" pageNumber="49">
<taxonomicName id="4C4E35C5FFC05030FF23FF5BFDEC44C2" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[151,553,151,178]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="vellyi" status="comb. nov.">
<emphasis id="B93A9254FFC05030FF23FF5BFE0644C1" bold="true" box="[151,451,151,178]" italics="true" pageId="48" pageNumber="49">Anhoraeomorphus vellyi</emphasis>
(Franz)
</taxonomicName>
,
<taxonomicNameLabel id="A2092F2FFFC05030FD83FF5BFD6744C2" box="[567,674,151,177]" pageId="48" pageNumber="49" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFC05030FF23FF77FE8644A6" box="[151,323,187,213]" pageId="48" pageNumber="49" type="description">
<paragraph id="8BF14E46FFC05030FF23FF77FE8644A6" blockId="48.[151,674,151,213]" box="[151,323,187,213]" pageId="48" pageNumber="49">
(
<figureCitation id="137552C3FFC05030FF2BFF77FEFE44A6" box="[159,315,187,213]" captionStart="FIGURES 179186" captionStartId="49.[151,264,1641,1665]" captionTargetBox="[151,1415,181,1602]" captionTargetId="figure-16@49.[151,1435,181,1617]" captionTargetPageId="49" captionText="FIGURES 179186. Anhoraeomorphus vellyi (Franz), holotype. Habitus in dorsal view (179); head in anterodorsal view (180); head and prothorax in lateral view (only bristles on head are shown) (181); antenna (182); aedeagus in ventral (183, 185) and lateral (184, 186) views." figureDoi="http://doi.org/10.5281/zenodo.13745983" httpUri="https://zenodo.org/record/13745983/files/figure.png" pageId="48" pageNumber="49">Figs 179186</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFC05030FF23FF33FD844564" box="[151,577,255,279]" pageId="48" pageNumber="49" type="reference_group">
<paragraph id="8BF14E46FFC05030FF23FF33FD844564" blockId="48.[151,577,255,279]" box="[151,577,255,279]" pageId="48" pageNumber="49">
<treatmentCitationGroup id="AB5E6968FFC05030FF23FF33FD844564" box="[151,577,255,279]" pageId="48" pageNumber="49">
<treatmentCitation id="0AEF6857FFC05030FF23FF33FDF84564" author="Franz, H." box="[151,573,255,279]" page="158" pageId="48" pageNumber="49" year="1986">
<taxonomicName id="4C4E35C5FFC05030FF23FF33FDF84564" ID-CoL="3MNLZ" authority="Franz, 1986 b: 158" authorityName="Franz" authorityPageNumber="158" authorityYear="1986" box="[151,573,255,279]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="vellyi">
<emphasis id="B93A9254FFC05030FF23FF33FEBE4564" box="[151,379,255,279]" italics="true" pageId="48" pageNumber="49">Horaeomorphus vellyi</emphasis>
<bibRefCitation id="EFDF33B7FFC05030FE35FECCFDF84564" author="Franz, H." box="[385,573,255,279]" pageId="48" pageNumber="49" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 158</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFC05030FF23FE8AFE0446BB" pageId="48" pageNumber="49" type="materials_examined">
<paragraph id="8BF14E46FFC05030FF23FE8AFD1D45DB" blockId="48.[151,1437,325,1864]" pageId="48" pageNumber="49">
<emphasis id="B93A9254FFC05030FF23FE8AFE5A452C" bold="true" box="[151,415,325,352]" pageId="48" pageNumber="49">Type material studied.</emphasis>
<materialsCitation id="3B26441BFFC05030FE10FE8AFD1D45DB" collectionCode="NHMW" collectorName="H. Franz" country="Madagascar" determinerName="H. Franz" location="Madagascar" pageId="48" pageNumber="49" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
<emphasis id="B93A9254FFC05030FE10FE8AFD78452C" bold="true" box="[420,701,325,352]" pageId="48" pageNumber="49">
<typeStatus id="54F5F0E4FFC05030FE10FE8AFDD44513" box="[420,529,326,352]" pageId="48" pageNumber="49" type="holotype">Holotype</typeStatus>
(
<collectingCountry id="F3590ED6FFC05030FDA9FE89FD71452C" box="[541,692,325,351]" name="Madagascar" pageId="48" pageNumber="49">Madagascar</collectingCountry>
)
</emphasis>
:
<specimenCount id="9D4885CFFFC05030FD7DFE8AFD254513" box="[713,736,326,352]" count="1" pageId="48" pageNumber="49" type="male"></specimenCount>
, four labels: “Madagaskar,Plateau / du Tampoketsa / lg.
<collectorName id="26BB2B90FFC05030FA8AFE8AFA594513" box="[1342,1436,326,352]" pageId="48" pageNumber="49">H.Franz</collectorName>
1969” [white, printed], “
<specimenCount id="9D4885CFFFC05030FE05FEA6FE0245F7" box="[433,455,362,388]" count="1" pageId="48" pageNumber="49" type="male"></specimenCount>
” [white, printed], “
<taxonomicName id="4C4E35C5FFC05030FD12FEA6FC9845F7" box="[678,861,362,388]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/ vellyi m. / det.
<determinerName id="7E2DA282FFC05030FBABFEA6FBB845F7" box="[1055,1149,362,388]" pageId="48" pageNumber="49">H.Franz</determinerName>
” [yellowish, handwritten and printed], “Typus” [red, handwritten] (
<collectionCode id="ED5FD683FFC05030FDDEFE42FD0845DB" box="[618,717,398,424]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="48" pageNumber="49" type="Museum">NHMW</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BF14E46FFC05030FF73FE7DFE0446BB" blockId="48.[151,1437,325,1864]" pageId="48" pageNumber="49">
<emphasis id="B93A9254FFC05030FF73FE7DFE6345B8" bold="true" box="[199,422,433,459]" pageId="48" pageNumber="49">Revised diagnosis.</emphasis>
Body (
<figureCitation id="137552C3FFC05030FDB0FE7EFDAC45BF" box="[516,617,434,460]" captionStart="FIGURES 179186" captionStartId="49.[151,264,1641,1665]" captionTargetBox="[151,1415,181,1602]" captionTargetId="figure-16@49.[151,1435,181,1617]" captionTargetPageId="49" captionText="FIGURES 179186. Anhoraeomorphus vellyi (Franz), holotype. Habitus in dorsal view (179); head in anterodorsal view (180); head and prothorax in lateral view (only bristles on head are shown) (181); antenna (182); aedeagus in ventral (183, 185) and lateral (184, 186) views." figureDoi="http://doi.org/10.5281/zenodo.13745983" httpUri="https://zenodo.org/record/13745983/files/figure.png" pageId="48" pageNumber="49">Fig. 179</figureCitation>
) slender; head (
<figureCitation id="137552C3FFC05030FC9FFE7EFC0945BF" box="[811,972,434,460]" captionStart="FIGURES 179186" captionStartId="49.[151,264,1641,1665]" captionTargetBox="[151,1415,181,1602]" captionTargetId="figure-16@49.[151,1435,181,1617]" captionTargetPageId="49" captionText="FIGURES 179186. Anhoraeomorphus vellyi (Franz), holotype. Habitus in dorsal view (179); head in anterodorsal view (180); head and prothorax in lateral view (only bristles on head are shown) (181); antenna (182); aedeagus in ventral (183, 185) and lateral (184, 186) views." figureDoi="http://doi.org/10.5281/zenodo.13745983" httpUri="https://zenodo.org/record/13745983/files/figure.png" pageId="48" pageNumber="49">Figs 180181</figureCitation>
) round, with vertex posteriorly weakly rounded; pronotum (
<figureCitation id="137552C3FFC05030FECAFE1AFE1B4583" box="[382,478,470,496]" captionStart="FIGURES 179186" captionStartId="49.[151,264,1641,1665]" captionTargetBox="[151,1415,181,1602]" captionTargetId="figure-16@49.[151,1435,181,1617]" captionTargetPageId="49" captionText="FIGURES 179186. Anhoraeomorphus vellyi (Franz), holotype. Habitus in dorsal view (179); head in anterodorsal view (180); head and prothorax in lateral view (only bristles on head are shown) (181); antenna (182); aedeagus in ventral (183, 185) and lateral (184, 186) views." figureDoi="http://doi.org/10.5281/zenodo.13745983" httpUri="https://zenodo.org/record/13745983/files/figure.png" pageId="48" pageNumber="49">Fig. 179</figureCitation>
) bell-shaped, broadest near anterior third, with one pair of large, transverse antebasal pits; antennomeres 34 (
<figureCitation id="137552C3FFC05030FE07FE36FDD04667" box="[435,533,506,532]" captionStart="FIGURES 179186" captionStartId="49.[151,264,1641,1665]" captionTargetBox="[151,1415,181,1602]" captionTargetId="figure-16@49.[151,1435,181,1617]" captionTargetPageId="49" captionText="FIGURES 179186. Anhoraeomorphus vellyi (Franz), holotype. Habitus in dorsal view (179); head in anterodorsal view (180); head and prothorax in lateral view (only bristles on head are shown) (181); antenna (182); aedeagus in ventral (183, 185) and lateral (184, 186) views." figureDoi="http://doi.org/10.5281/zenodo.13745983" httpUri="https://zenodo.org/record/13745983/files/figure.png" pageId="48" pageNumber="49">Fig. 182</figureCitation>
) each about as long as broad, 5 slightly elongate, 610 each about as long as broad; all femora similarly slender; metatibiae in male modified, slightly recurved and broadened in distal region, broadened area with elongate outer impression bordered by sharp outer edge; aedeagus (
<figureCitation id="137552C3FFC05030FB37FD8EFAE5462F" box="[1155,1312,578,604]" captionStart="FIGURES 179186" captionStartId="49.[151,264,1641,1665]" captionTargetBox="[151,1415,181,1602]" captionTargetId="figure-16@49.[151,1435,181,1617]" captionTargetPageId="49" captionText="FIGURES 179186. Anhoraeomorphus vellyi (Franz), holotype. Habitus in dorsal view (179); head in anterodorsal view (180); head and prothorax in lateral view (only bristles on head are shown) (181); antenna (182); aedeagus in ventral (183, 185) and lateral (184, 186) views." figureDoi="http://doi.org/10.5281/zenodo.13745983" httpUri="https://zenodo.org/record/13745983/files/figure.png" pageId="48" pageNumber="49">Figs 183186</figureCitation>
) in ventral view broadest near middle, apical region of dorsal wall in lateral view weakly curved dorsally, distolateral endophallic sclerites narrowly separated, each elongate and with blunt subtriangular apex, each paramere with group of six short subapical and apical setae.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFC05030FF73FD1DFE5B4203" pageId="48" pageNumber="49" type="description">
<paragraph id="8BF14E46FFC05030FF73FD1DFD914763" blockId="48.[151,1437,325,1864]" pageId="48" pageNumber="49">
<emphasis id="B93A9254FFC05030FF73FD1DFEB04698" bold="true" box="[199,373,721,747]" pageId="48" pageNumber="49">Redescription.</emphasis>
Body of male (
<figureCitation id="137552C3FFC05030FD87FD1EFD52469F" box="[563,663,722,748]" captionStart="FIGURES 179186" captionStartId="49.[151,264,1641,1665]" captionTargetBox="[151,1415,181,1602]" captionTargetId="figure-16@49.[151,1435,181,1617]" captionTargetPageId="49" captionText="FIGURES 179186. Anhoraeomorphus vellyi (Franz), holotype. Habitus in dorsal view (179); head in anterodorsal view (180); head and prothorax in lateral view (only bristles on head are shown) (181); antenna (182); aedeagus in ventral (183, 185) and lateral (184, 186) views." figureDoi="http://doi.org/10.5281/zenodo.13745983" httpUri="https://zenodo.org/record/13745983/files/figure.png" pageId="48" pageNumber="49">Fig. 179</figureCitation>
) strongly convex, moderately dark brown, tarsi and palps slightly lighter; setae light brown; BL
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.
</paragraph>
<paragraph id="8BF14E46FFC05030FF73FCD6FF1A4043" blockId="48.[151,1437,325,1864]" pageId="48" pageNumber="49">
Head (
<figureCitation id="137552C3FFC05030FEA0FCD6FEB04747" box="[276,373,794,820]" captionStart="FIGURES 179186" captionStartId="49.[151,264,1641,1665]" captionTargetBox="[151,1415,181,1602]" captionTargetId="figure-16@49.[151,1435,181,1617]" captionTargetPageId="49" captionText="FIGURES 179186. Anhoraeomorphus vellyi (Franz), holotype. Habitus in dorsal view (179); head in anterodorsal view (180); head and prothorax in lateral view (only bristles on head are shown) (181); antenna (182); aedeagus in ventral (183, 185) and lateral (184, 186) views." figureDoi="http://doi.org/10.5281/zenodo.13745983" httpUri="https://zenodo.org/record/13745983/files/figure.png" pageId="48" pageNumber="49">Fig. 179</figureCitation>
) in anterodorsal view round, broadest at eyes, HL
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, HW
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; temple in lateral view (
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) subequal to eye; vertex and frons confluent, together weakly convex and slightly transverse; vertex posteriorly weakly rounded; each eye large and weakly convex, oval, not emarginate posteriorly, weakly projecting from head silhouette. Punctures on vertex fine, inconspicuous; setae short, dense and suberect; frons and vertex densely covered with thick bristles directed posteriorly. Antenna (
<figureCitation id="137552C3FFC05030FCCAFC66FC1B47B7" box="[894,990,938,964]" captionStart="FIGURES 179186" captionStartId="49.[151,264,1641,1665]" captionTargetBox="[151,1415,181,1602]" captionTargetId="figure-16@49.[151,1435,181,1617]" captionTargetPageId="49" captionText="FIGURES 179186. Anhoraeomorphus vellyi (Franz), holotype. Habitus in dorsal view (179); head in anterodorsal view (180); head and prothorax in lateral view (only bristles on head are shown) (181); antenna (182); aedeagus in ventral (183, 185) and lateral (184, 186) views." figureDoi="http://doi.org/10.5281/zenodo.13745983" httpUri="https://zenodo.org/record/13745983/files/figure.png" pageId="48" pageNumber="49">Fig. 182</figureCitation>
) slender but short, slightly shorter than half BL, AnL
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, antennomeres 12 each elongate, 34 each about as long as broad, 5 slightly elongate, 610 each about as long as broad, 11 slightly shorter than 9 and 10 combined, as wide as 10, about 1.7 times as long as broad.
</paragraph>
<paragraph id="8BF14E46FFC05030FF73FBF6FEEA4097" blockId="48.[151,1437,325,1864]" pageId="48" pageNumber="49">
Pronotum bell-shaped, broadest near anterior third; PL
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, PW
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. Anterior margin strongly rounded, lateral margins rounded in anterior half, weakly sinuate in posterior half; posterior corners blunt; posterior margin nearly straight. Base with one pair of large and strongly transverse pits, sublateral carinae short but distinct. Disc with fine, inconspicuous punctures; setae dense, short and suberect, lateral surface of pronotum with dense thick bristles.
</paragraph>
<paragraph id="8BF14E46FFC05030FF73FB22FACC4123" blockId="48.[151,1437,325,1864]" pageId="48" pageNumber="49">
Elytra together oval, broadest distinctly in front of middle; EL
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, EW
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, EI 1.45; humeral calli weakly elevated, basal elytral foveae barely discernible. Punctures small and shallow but distinct, those on anterior half separated by spaces slightly shorter than diameters of punctures; setae sparse, short and suberect.
</paragraph>
<paragraph id="8BF14E46FFC05030FF73FA96FAE941EB" blockId="48.[151,1437,325,1864]" pageId="48" pageNumber="49">Legs conspicuously long and slender, unmodified except hind tibiae, which are slightly recurved and slightly broadened in distal regions, with the broadened area longitudinally impressed and with outer sharp edge.</paragraph>
<paragraph id="8BF14E46FFC05030FF73FA6EFAF2423F" blockId="48.[151,1437,325,1864]" pageId="48" pageNumber="49">
Aedeagus (
<figureCitation id="137552C3FFC05030FEFDFA6EFE2D41CF" box="[329,488,1442,1468]" captionStart="FIGURES 179186" captionStartId="49.[151,264,1641,1665]" captionTargetBox="[151,1415,181,1602]" captionTargetId="figure-16@49.[151,1435,181,1617]" captionTargetPageId="49" captionText="FIGURES 179186. Anhoraeomorphus vellyi (Franz), holotype. Habitus in dorsal view (179); head in anterodorsal view (180); head and prothorax in lateral view (only bristles on head are shown) (181); antenna (182); aedeagus in ventral (183, 185) and lateral (184, 186) views." figureDoi="http://doi.org/10.5281/zenodo.13745983" httpUri="https://zenodo.org/record/13745983/files/figure.png" pageId="48" pageNumber="49">Figs 183186</figureCitation>
) stout, AeL
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; in ventral view nearly drop-shaped, broadest near middle, distally gradually narrowing and in subapical region rapidly narrowed to form subtriangular apex; in lateral view apical region of dorsal wall weakly curved dorsally; endophallus with all sclerotized structures asymmetrical, distolateral sclerites narrowly separated, each with subtriangular and blunt apical region; parameres broad and their apices not projecting beyond aedeagal apex, each paramere with subapical/apical group of six short setae.
</paragraph>
<paragraph id="8BF14E46FFC05030FF73F99AFE5B4203" blockId="48.[151,1437,325,1864]" box="[199,414,1622,1648]" pageId="48" pageNumber="49">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFC05030FF73F9B5FC8A42E7" box="[199,847,1657,1684]" pageId="48" pageNumber="49" type="distribution">
<paragraph id="8BF14E46FFC05030FF73F9B5FC8A42E7" blockId="48.[151,1437,325,1864]" box="[199,847,1657,1684]" pageId="48" pageNumber="49">
<emphasis id="B93A9254FFC05030FF73F9B5FEA442E0" bold="true" box="[199,353,1657,1683]" pageId="48" pageNumber="49">Distribution.</emphasis>
Central
<collectingCountry id="F3590ED6FFC05030FE75F9B6FD8D42E7" box="[449,584,1658,1684]" name="Madagascar" pageId="48" pageNumber="49">Madagascar</collectingCountry>
(Analamanga Region).
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFC05030FF73F952FD40433B" pageId="48" pageNumber="49" type="discussion">
<paragraph id="8BF14E46FFC05030FF73F952FC2942AF" blockId="48.[151,1437,325,1864]" pageId="48" pageNumber="49">
<emphasis id="B93A9254FFC05030FF73F952FEFE42CB" bold="true" box="[199,315,1694,1720]" pageId="48" pageNumber="49">Remarks.</emphasis>
<taxonomicName id="4C4E35C5FFC05030FEF7F952FD9C42CB" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[323,601,1694,1720]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="vellyi">
<emphasis id="B93A9254FFC05030FEF7F952FD9C42CB" box="[323,601,1694,1720]" italics="true" pageId="48" pageNumber="49">Anhoraeomorphus vellyi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4E35C5FFC05030FD25F952FCB542CB" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[657,880,1694,1720]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="lambomakandroi">
<emphasis id="B93A9254FFC05030FD25F952FCB542CB" box="[657,880,1694,1720]" italics="true" pageId="48" pageNumber="49">A. lambomakandroi</emphasis>
</taxonomicName>
may be conspecific, but the available material is not sufficient to clarify this suspicion. See Remarks for
<taxonomicName id="4C4E35C5FFC05030FCBCF90EFC2042AF" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[776,997,1730,1756]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="lambomakandroi">
<emphasis id="B93A9254FFC05030FCBCF90EFC2042AF" box="[776,997,1730,1756]" italics="true" pageId="48" pageNumber="49">A. lambomakandroi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF14E46FFC05030FF73F92AFD40433B" blockId="48.[151,1437,325,1864]" pageId="48" pageNumber="49">
Similarly as for
<taxonomicName id="4C4E35C5FFC05030FECEF92AFE384373" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[378,509,1766,1792]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="barbatus">
<emphasis id="B93A9254FFC05030FECEF92AFE384373" box="[378,509,1766,1792]" italics="true" pageId="48" pageNumber="49">A. barbatus</emphasis>
</taxonomicName>
, “Plateau du Tampoketsa” is the only locality given in label data, but in the original description
<bibRefCitation id="EFDF33B7FFC05030FEA8F8C6FE784357" author="Franz, H." box="[284,445,1802,1828]" pageId="48" pageNumber="49" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986b)</bibRefCitation>
adds more detailed information: “Massif du Tampokeza {sic!} nördlich Tananarive bei der Forststation Manonkazo am
<date id="FFF06886FFC05030FDB6F8E2FDB2433B" box="[514,631,1838,1864]" pageId="48" pageNumber="49" value="1969-11-04">11.4.1969</date>
”.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,102 @@
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<emphasis id="B93A9254FFC15031FF23F8D7FD2D4345" bold="true" box="[151,744,1819,1846]" pageId="49" pageNumber="50">
<taxonomicName id="4C4E35C5FFC15031FF23F8D7FDAA4346" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[151,623,1819,1846]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="ankaratrae" status="comb. nov.">
<emphasis id="B93A9254FFC15031FF23F8D7FDCC4345" bold="true" box="[151,521,1819,1846]" italics="true" pageId="49" pageNumber="50">Anhoraeomorphus ankaratrae</emphasis>
(Franz)
</taxonomicName>
,
<taxonomicNameLabel id="A2092F2FFFC15031FDC9F8D0FD2D4345" box="[637,744,1820,1846]" pageId="49" pageNumber="50" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFC15031FF23F8ACFDBE430B" box="[151,635,1888,1912]" pageId="49" pageNumber="50" type="reference_group">
<paragraph id="8BF14E46FFC15031FF23F8ACFDBE430B" blockId="49.[151,635,1888,1912]" box="[151,635,1888,1912]" pageId="49" pageNumber="50">
<treatmentCitationGroup id="AB5E6968FFC15031FF23F8ACFDBE430B" box="[151,635,1888,1912]" pageId="49" pageNumber="50">
<treatmentCitation id="0AEF6857FFC15031FF23F8ACFDB2430B" author="Franz, H." box="[151,631,1888,1912]" page="160" pageId="49" pageNumber="50" year="1986">
<taxonomicName id="4C4E35C5FFC15031FF23F8ACFDB2430B" ID-CoL="3MNJ6" authority="Franz, 1986 b: 160" authorityName="Franz" authorityPageNumber="160" authorityYear="1986" box="[151,631,1888,1912]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="ankaratrae">
<emphasis id="B93A9254FFC15031FF23F8ACFE70430B" box="[151,437,1888,1912]" italics="true" pageId="49" pageNumber="50">Horaeomorphus ankaratrae</emphasis>
<bibRefCitation id="EFDF33B7FFC15031FE0FF8ACFDB2430B" author="Franz, H." box="[443,631,1888,1912]" pageId="49" pageNumber="50" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 160</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFC15032FF23F86AFEE3448A" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50" type="discussion">
<paragraph id="8BF14E46FFC15032FF23F86AFEE3448A" blockId="49.[151,1436,1958,2021]" lastBlockId="50.[151,1437,151,285]" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50">
<emphasis id="B93A9254FFC15031FF23F86AFECE43B3" bold="true" box="[151,267,1958,1984]" pageId="49" pageNumber="50">Remarks.</emphasis>
The
<typeStatus id="54F5F0E4FFC15031FEFCF86AFE6F43B3" box="[328,426,1958,1984]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
(the only known specimen) is deposited in Muséum National dHistoire Naturelle, Paris and was not available for my study. Characters described by Franz, especially the illustrated aedeagus with a pair of elongate distolateral endophallic sclerites, and the male metatibiae sinuate and distally broadened leave no doubts that this species belongs in
<taxonomicName id="4C4E35C5FFC25032FE6CFF77FD6D44A6" box="[472,680,187,213]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B93A9254FFC25032FE6CFF77FD6D44A6" box="[472,680,187,213]" italics="true" pageId="50" pageNumber="51">Anhoraeomorphus</emphasis>
</taxonomicName>
. According to
<bibRefCitation id="EFDF33B7FFC25032FCE2FF77FC3E44A6" author="Franz, H." box="[854,1019,187,213]" pageId="50" pageNumber="51" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986b)</bibRefCitation>
,
<taxonomicName id="4C4E35C5FFC25032FBBDFF77FB6244A6" authorityName="Franz" authorityYear="1986" box="[1033,1191,187,213]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="ankaratrae">
<emphasis id="B93A9254FFC25032FBBDFF77FB6244A6" box="[1033,1191,187,213]" italics="true" pageId="50" pageNumber="51">A. ankaratrae</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C4E35C5FFC25032FA30FF77FEDB448A" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="adamsoniae">
<emphasis id="B93A9254FFC25032FA30FF77FEDB448A" italics="true" pageId="50" pageNumber="51">A. adamsoniae</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFC25032FF73FECEFCAE456E" box="[199,875,258,285]" pageId="50" pageNumber="51" type="distribution">
<paragraph id="8BF14E46FFC25032FF73FECEFCAE456E" blockId="50.[151,1437,151,285]" box="[199,875,258,285]" pageId="50" pageNumber="51">
<emphasis id="B93A9254FFC25032FF73FECEFEA4456F" bold="true" box="[199,353,258,284]" pageId="50" pageNumber="51">Distribution.</emphasis>
Central
<collectingCountry id="F3590ED6FFC25032FE75FECFFD8D456E" box="[449,584,259,285]" name="Madagascar" pageId="50" pageNumber="51">Madagascar</collectingCountry>
(Vakinankaratra Region).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,118 @@
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<subSubSection id="C3541DCDFFC25032FF23FA79FD2541BC" box="[151,736,1461,1488]" pageId="50" pageNumber="51" type="nomenclature">
<paragraph id="8BF14E46FFC25032FF23FA79FD2541BC" blockId="50.[151,736,1461,1488]" box="[151,736,1461,1488]" pageId="50" pageNumber="51">
<heading id="D0B9F92AFFC25032FF23FA79FD2541BC" bold="true" box="[151,736,1461,1488]" fontSize="11" level="2" pageId="50" pageNumber="51" reason="4">
<emphasis id="B93A9254FFC25032FF23FA79FD2541BC" bold="true" box="[151,736,1461,1488]" pageId="50" pageNumber="51">
<taxonomicName id="4C4E35C5FFC25032FF23FA79FDA241BC" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[151,615,1461,1488]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="longicollis" status="comb. nov.">
<emphasis id="B93A9254FFC25032FF23FA79FDC441A3" bold="true" box="[151,513,1461,1488]" italics="true" pageId="50" pageNumber="51">Anhoraeomorphus longicollis</emphasis>
(Franz)
</taxonomicName>
,
<taxonomicNameLabel id="A2092F2FFFC25032FDC1FA79FD2541BC" box="[629,736,1461,1487]" pageId="50" pageNumber="51" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFC25032FF23FA36FDB04261" box="[151,629,1530,1554]" pageId="50" pageNumber="51" type="reference_group">
<paragraph id="8BF14E46FFC25032FF23FA36FDB04261" blockId="50.[151,629,1530,1554]" box="[151,629,1530,1554]" pageId="50" pageNumber="51">
<treatmentCitationGroup id="AB5E6968FFC25032FF23FA36FDB04261" box="[151,629,1530,1554]" pageId="50" pageNumber="51">
<treatmentCitation id="0AEF6857FFC25032FF23FA36FDB44261" author="Franz, H." box="[151,625,1530,1554]" page="157" pageId="50" pageNumber="51" year="1986">
<taxonomicName id="4C4E35C5FFC25032FF23FA36FDB44261" ID-CoL="3MNKJ" authority="Franz, 1986 b: 157" authorityName="Franz" authorityPageNumber="157" authorityYear="1986" box="[151,625,1530,1554]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="longicollis">
<emphasis id="B93A9254FFC25032FF23FA36FE6A4261" box="[151,431,1530,1554]" italics="true" pageId="50" pageNumber="51">Horaeomorphus longicollis</emphasis>
<bibRefCitation id="EFDF33B7FFC25032FE01FA36FDB44261" author="Franz, H." box="[437,625,1530,1554]" pageId="50" pageNumber="51" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 157</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFC25032FF23F98CFCF64325" pageId="50" pageNumber="51" type="discussion">
<paragraph id="8BF14E46FFC25032FF23F98CFCF64325" blockId="50.[151,1437,1600,1914]" pageId="50" pageNumber="51">
<emphasis id="B93A9254FFC25032FF23F98CFECE4229" bold="true" box="[151,267,1600,1626]" pageId="50" pageNumber="51">Remarks.</emphasis>
The
<typeStatus id="54F5F0E4FFC25032FEFCF98CFE6F4229" box="[328,426,1600,1626]" pageId="50" pageNumber="51" type="holotype">holotype</typeStatus>
(the only known specimen) is deposited in Muséum National dHistoire Naturelle, Paris and was not available for my study. Characters described and illustrated by Franz include the aedeagus with a pair of elongate distolateral endophallic sclerites accompanied by median proximal assemblage of asymmetrical structures, and male metatibiae broadened in distal third, which are typical of
<taxonomicName id="4C4E35C5FFC25032FC34F960FB9542B5" box="[896,1104,1708,1734]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B93A9254FFC25032FC34F960FB9542B5" box="[896,1104,1708,1734]" italics="true" pageId="50" pageNumber="51">Anhoraeomorphus</emphasis>
</taxonomicName>
. According to
<bibRefCitation id="EFDF33B7FFC25032FB42F961FA5242B5" author="Franz, H." box="[1270,1431,1708,1735]" pageId="50" pageNumber="51" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986b)</bibRefCitation>
,
<taxonomicName id="4C4E35C5FFC25032FF23F91CFEE94299" authorityName="Franz" authorityYear="1986" box="[151,300,1744,1770]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="longicollis">
<emphasis id="B93A9254FFC25032FF23F91CFEE94299" box="[151,300,1744,1770]" italics="true" pageId="50" pageNumber="51">A. longicollis</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C4E35C5FFC25032FE49F91CFDA64299" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[509,611,1744,1770]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="diegoi">
<emphasis id="B93A9254FFC25032FE49F91CFDA64299" box="[509,611,1744,1770]" italics="true" pageId="50" pageNumber="51">A. diegoi</emphasis>
</taxonomicName>
. Judging from the aedeagus of
<taxonomicName id="4C4E35C5FFC25032FC70F91CFB9C4299" authorityName="Franz" authorityYear="1986" box="[964,1113,1744,1770]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="longicollis">
<emphasis id="B93A9254FFC25032FC70F91CFB9C4299" box="[964,1113,1744,1770]" italics="true" pageId="50" pageNumber="51">A. longicollis</emphasis>
</taxonomicName>
illustrated in
<bibRefCitation id="EFDF33B7FFC25032FB41F91DFA5D4299" author="Franz, H." box="[1269,1432,1744,1771]" pageId="50" pageNumber="51" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986b)</bibRefCitation>
: fig. 141), and that of
<taxonomicName id="4C4E35C5FFC25032FE33F938FE28437D" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[391,493,1780,1806]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="diegoi">
<emphasis id="B93A9254FFC25032FE33F938FE28437D" box="[391,493,1780,1806]" italics="true" pageId="50" pageNumber="51">A. diegoi</emphasis>
</taxonomicName>
studied here, these two names may be synonyms. Re-mounting and examination of the aedeagus of the
<typeStatus id="54F5F0E4FFC25032FECEF8D4FE194341" box="[378,476,1816,1842]" pageId="50" pageNumber="51" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C4E35C5FFC25032FDB7F8D4FD5C4341" authorityName="Franz" authorityYear="1986" box="[515,665,1816,1842]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="longicollis">
<emphasis id="B93A9254FFC25032FDB7F8D4FD5C4341" box="[515,665,1816,1842]" italics="true" pageId="50" pageNumber="51">A. longicollis</emphasis>
</taxonomicName>
is necessary to verify this view, but currently the Muséum National d'Histoire Naturelle does not issue loans of type specimens.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFC25032FF73F8ACFCA64309" box="[199,867,1888,1914]" pageId="50" pageNumber="51" type="distribution">
<paragraph id="8BF14E46FFC25032FF73F8ACFCA64309" blockId="50.[151,1437,1600,1914]" box="[199,867,1888,1914]" pageId="50" pageNumber="51">
<emphasis id="B93A9254FFC25032FF73F8ACFEA44309" bold="true" box="[199,353,1888,1914]" pageId="50" pageNumber="51">Distribution.</emphasis>
Northern
<collectingCountry id="F3590ED6FFC25032FE60F8ACFD9E4309" box="[468,603,1888,1914]" name="Madagascar" pageId="50" pageNumber="51">Madagascar</collectingCountry>
(Analanjirofo Region).
</paragraph>
</subSubSection>
</treatment>
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<mods:title id="933CEB4B6EFA64CE03B797F965210613">The curse of Horaeomorphus (almost) lifted. Revision of misplaced species from Madagascar, Comoros and Mascarenes (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<emphasis id="B93A9254FFC25032FF23FBC6FE064056" bold="true" box="[151,451,1034,1061]" italics="true" pageId="50" pageNumber="51">Anhoraeomorphus gigas</emphasis>
(Franz)
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,
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<emphasis id="B93A9254FFC25032FF23FB83FEBE4014" box="[151,379,1103,1127]" italics="true" pageId="50" pageNumber="51">Horaeomorphus gigas</emphasis>
<bibRefCitation id="EFDF33B7FFC25032FE35FB9CFDF84014" author="Franz, H." box="[385,573,1103,1127]" pageId="50" pageNumber="51" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 171</bibRefCitation>
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.
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<emphasis id="B93A9254FFC25032FF23FB59FECE40DC" bold="true" box="[151,267,1173,1199]" pageId="50" pageNumber="51">Remarks.</emphasis>
According to
<bibRefCitation id="EFDF33B7FFC25032FE1FFB5AFD8940DC" author="Franz, H." box="[427,588,1173,1200]" pageId="50" pageNumber="51" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986b)</bibRefCitation>
, the
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female (the only known specimen) of this species is deposited in Tervuren. This specimen was not found in Musée Royal de lAfrique Centrale, Tervuren,
<collectingCountry id="F3590ED6FFC25032FB19FB75FAC840A0" box="[1197,1293,1209,1235]" name="Belgium" pageId="50" pageNumber="51">Belgium</collectingCountry>
(email from Stéphane Hanot dated
<date id="FFF06886FFC25032FE20FB11FDD24084" box="[404,535,1245,1271]" pageId="50" pageNumber="51" value="2024-02-04">02.04.2024</date>
), and it is not present in Franz Coll. at NHMW (pers. obs.).
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<bibRefCitation id="EFDF33B7FFC25032FF73FACEFEA04168" author="Franz, H." box="[199,357,1281,1308]" pageId="50" pageNumber="51" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986b)</bibRefCitation>
described this species as most similar to
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<emphasis id="B93A9254FFC25032FC92FACEFC6D4168" box="[806,936,1282,1308]" italics="true" pageId="50" pageNumber="51">A. maximus</emphasis>
</taxonomicName>
, and there is little doubt that
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<emphasis id="B93A9254FFC25032FB51FACDFF11414C" italics="true" pageId="50" pageNumber="51">Horaeomorphus gigas</emphasis>
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is also a member of
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<emphasis id="B93A9254FFC25032FE09FAE9FD48414C" box="[445,653,1317,1343]" italics="true" pageId="50" pageNumber="51">Anhoraeomorphus</emphasis>
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.
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<emphasis id="B93A9254FFC25032FF73FA85FEA44110" bold="true" box="[199,353,1353,1379]" pageId="50" pageNumber="51">Distribution.</emphasis>
Northern
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(Analanjirofo Region).
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(
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)
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<taxonomicName id="4C4E35C5FFD05020FF23FCF3FDA54724" ID-CoL="3MNK2" authority="Franz, 1986 b: 169" authorityName="Franz" authorityPageNumber="169" authorityYear="1986" box="[151,608,831,855]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="globosus">
<emphasis id="B93A9254FFD05020FF23FCF3FE5B4724" box="[151,414,831,855]" italics="true" pageId="32" pageNumber="33">Horaeomorphus globosus</emphasis>
<bibRefCitation id="EFDF33B7FFD05020FE10FC8CFDA54724" author="Franz, H." box="[420,608,831,855]" pageId="32" pageNumber="33" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 169</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD05020FF23FC49FD90415F" pageId="32" pageNumber="33" type="materials_examined">
<paragraph id="8BF14E46FFD05020FF23FC49FE1B405C" blockId="32.[151,1437,901,1935]" pageId="32" pageNumber="33">
<emphasis id="B93A9254FFD05020FF23FC49FE7C47EC" bold="true" box="[151,441,901,927]" pageId="32" pageNumber="33">Type material studied.</emphasis>
<materialsCitation id="3B26441BFFD05020FE7EFC49FE1B405C" collectionCode="NHMW" country="Madagascar" determinerName="H. Franz" elevation="2030" location="MADAGASCAR" pageId="32" pageNumber="33" specimenCount="59" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B93A9254FFD05020FE7EFC49FD3547EC" bold="true" box="[458,752,901,927]" pageId="32" pageNumber="33">
<typeStatus id="54F5F0E4FFD05020FE7EFC49FDF247EC" box="[458,567,901,927]" pageId="32" pageNumber="33" type="holotype">Holotype</typeStatus>
(Madagascar)
</emphasis>
:
<specimenCount id="9D4885CFFFD05020FCBDFC4AFCE547D3" box="[777,800,902,928]" count="1" pageId="32" pageNumber="33" type="male"></specimenCount>
, five labels: “Madagascar-Centre / Forét Imaitso / Anjavidilava
<quantity id="4CB6E3A3FFD05020FE9AFC65FEB947B0" box="[302,380,937,963]" metricMagnitude="3" metricUnit="m" metricValue="2.03" pageId="32" pageNumber="33" unit="m" value="2030.0">
<elevation id="0063A975FFD05020FE9AFC65FEB947B0" box="[302,380,937,963]" metricMagnitude="3" metricUnit="m" metricValue="2.03" pageId="32" pageNumber="33" unit="m" value="2030.0">2030m</elevation>
</quantity>
/ Andingitra-Ambalavao / 19.1.
<specimenCount id="9D4885CFFFD05020FD6FFC65FCD647B7" box="[731,787,937,964]" count="58" pageId="32" pageNumber="33" type="generic">58 P.</specimenCount>
Grivaud” [white, printed], “INSTITUT / SCIENTIFIQUE /
<collectingCountry id="F3590ED6FFD05020FF1FFC01FEB74794" box="[171,370,973,999]" name="Madagascar" pageId="32" pageNumber="33">MADAGASCAR</collectingCountry>
” [bluish, printed], “Hinterhüften / schmal getrennt {
<emphasis id="B93A9254FFD05020FC52FC02FBCD4794" box="[998,1032,974,999]" italics="true" pageId="32" pageNumber="33">i.e.</emphasis>
, hind coxae narrowly separated}” [yellow, handwritten], “
<taxonomicName id="4C4E35C5FFD05020FE2AFC3DFD904078" box="[414,597,1009,1035]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/ globosus m. / det.
<determinerName id="7E2DA282FFD05020FC99FC3DFC4E4078" box="[813,907,1009,1035]" pageId="32" pageNumber="33">H.Franz</determinerName>
” [yellowish, handwritten and printed], “Typus” [red, handwritten] (
<collectionCode id="ED5FD683FFD05020FEC4FBD9FE16405C" box="[368,467,1045,1071]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="32" pageNumber="33" type="Museum">NHMW</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BF14E46FFD05020FF73FBF5FD90415F" blockId="32.[151,1437,901,1935]" pageId="32" pageNumber="33">
<emphasis id="B93A9254FFD05020FF73FBF5FE6D4020" bold="true" box="[199,424,1081,1107]" pageId="32" pageNumber="33">Revised diagnosis.</emphasis>
Body (
<figureCitation id="137552C3FFD05020FDB3FBF5FDA94020" box="[519,620,1081,1107]" captionStart="FIGURES 119126" captionStartId="33.[151,264,1567,1591]" captionTargetBox="[151,1420,181,1531]" captionTargetId="figure-16@33.[151,1435,181,1543]" captionTargetPageId="33" captionText="FIGURES 119126. Anhoraeomorphus globosus (Franz), holotype. Habitus in dorsal view (119); head in anterodorsal view (120); head and prothorax in lateral view (only bristles on head are shown) (121); antenna (122); aedeagus in ventral (123, 125) and lateral (124, 126) views." figureDoi="http://doi.org/10.5281/zenodo.13745963" httpUri="https://zenodo.org/record/13745963/files/figure.png" pageId="32" pageNumber="33">Fig. 119</figureCitation>
) slender; head (
<figureCitation id="137552C3FFD05020FC85FBF5FC164020" box="[817,979,1081,1107]" captionStart="FIGURES 119126" captionStartId="33.[151,264,1567,1591]" captionTargetBox="[151,1420,181,1531]" captionTargetId="figure-16@33.[151,1435,181,1543]" captionTargetPageId="33" captionText="FIGURES 119126. Anhoraeomorphus globosus (Franz), holotype. Habitus in dorsal view (119); head in anterodorsal view (120); head and prothorax in lateral view (only bristles on head are shown) (121); antenna (122); aedeagus in ventral (123, 125) and lateral (124, 126) views." figureDoi="http://doi.org/10.5281/zenodo.13745963" httpUri="https://zenodo.org/record/13745963/files/figure.png" pageId="32" pageNumber="33">Figs 120121</figureCitation>
) oval, with vertex posteriorly broadly rounded; pronotum (
<figureCitation id="137552C3FFD05020FE35FB91FE244004" box="[385,481,1117,1143]" captionStart="FIGURES 119126" captionStartId="33.[151,264,1567,1591]" captionTargetBox="[151,1420,181,1531]" captionTargetId="figure-16@33.[151,1435,181,1543]" captionTargetPageId="33" captionText="FIGURES 119126. Anhoraeomorphus globosus (Franz), holotype. Habitus in dorsal view (119); head in anterodorsal view (120); head and prothorax in lateral view (only bristles on head are shown) (121); antenna (122); aedeagus in ventral (123, 125) and lateral (124, 126) views." figureDoi="http://doi.org/10.5281/zenodo.13745963" httpUri="https://zenodo.org/record/13745963/files/figure.png" pageId="32" pageNumber="33">Fig. 119</figureCitation>
) bell-shaped, broadest near anterior third, with one pair of large and deep antebasal pits; antennomeres 36 (
<figureCitation id="137552C3FFD05020FE1CFB4DFDC240E8" box="[424,519,1153,1179]" captionStart="FIGURES 119126" captionStartId="33.[151,264,1567,1591]" captionTargetBox="[151,1420,181,1531]" captionTargetId="figure-16@33.[151,1435,181,1543]" captionTargetPageId="33" captionText="FIGURES 119126. Anhoraeomorphus globosus (Franz), holotype. Habitus in dorsal view (119); head in anterodorsal view (120); head and prothorax in lateral view (only bristles on head are shown) (121); antenna (122); aedeagus in ventral (123, 125) and lateral (124, 126) views." figureDoi="http://doi.org/10.5281/zenodo.13745963" httpUri="https://zenodo.org/record/13745963/files/figure.png" pageId="32" pageNumber="33">Fig. 122</figureCitation>
) each elongate, 7 about as long as broad, 810 each slightly transverse; profemora slightly thicker than meso- and metafemora; hind tibiae in male unmodified; aedeagus (
<figureCitation id="137552C3FFD05020FB35FB69FADA40CC" box="[1153,1311,1189,1215]" captionStart="FIGURES 119126" captionStartId="33.[151,264,1567,1591]" captionTargetBox="[151,1420,181,1531]" captionTargetId="figure-16@33.[151,1435,181,1543]" captionTargetPageId="33" captionText="FIGURES 119126. Anhoraeomorphus globosus (Franz), holotype. Habitus in dorsal view (119); head in anterodorsal view (120); head and prothorax in lateral view (only bristles on head are shown) (121); antenna (122); aedeagus in ventral (123, 125) and lateral (124, 126) views." figureDoi="http://doi.org/10.5281/zenodo.13745963" httpUri="https://zenodo.org/record/13745963/files/figure.png" pageId="32" pageNumber="33">Figs 123126</figureCitation>
) in ventral view broadest in sub-basal third, apical region of dorsal wall in lateral view abruptly and strongly bent dorsally, distolateral endophallic sclerites broadly separated, each elongate and with blunt subtriangular apex, each paramere with group of several short apical setae.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD05022FF73FAF9FE5B44C2" lastPageId="34" lastPageNumber="35" pageId="32" pageNumber="33" type="description">
<paragraph id="8BF14E46FFD05020FF73FAF9FE474107" blockId="32.[151,1437,901,1935]" pageId="32" pageNumber="33">
<emphasis id="B93A9254FFD05020FF73FAF9FEB0413C" bold="true" box="[199,373,1333,1359]" pageId="32" pageNumber="33">Redescription.</emphasis>
Body of male (
<figureCitation id="137552C3FFD05020FDABFAF9FDB9413C" box="[543,636,1333,1359]" captionStart="FIGURES 119126" captionStartId="33.[151,264,1567,1591]" captionTargetBox="[151,1420,181,1531]" captionTargetId="figure-16@33.[151,1435,181,1543]" captionTargetPageId="33" captionText="FIGURES 119126. Anhoraeomorphus globosus (Franz), holotype. Habitus in dorsal view (119); head in anterodorsal view (120); head and prothorax in lateral view (only bristles on head are shown) (121); antenna (122); aedeagus in ventral (123, 125) and lateral (124, 126) views." figureDoi="http://doi.org/10.5281/zenodo.13745963" httpUri="https://zenodo.org/record/13745963/files/figure.png" pageId="32" pageNumber="33">Fig. 119</figureCitation>
) strongly convex, dark brown, tarsi and palps slightly lighter; setae light brown; BL
<quantity id="4CB6E3A3FFD05020FEA3FA95FEBB4107" box="[279,382,1369,1396]" metricMagnitude="-3" metricUnit="m" metricValue="2.53" pageId="32" pageNumber="33" unit="mm" value="2.53">2.53 mm</quantity>
.
</paragraph>
<paragraph id="8BF14E46FFD05020FF73FAB1FAC0421C" blockId="32.[151,1437,901,1935]" pageId="32" pageNumber="33">
Head (
<figureCitation id="137552C3FFD05020FEA5FAB1FEB541E4" box="[273,368,1405,1431]" captionStart="FIGURES 119126" captionStartId="33.[151,264,1567,1591]" captionTargetBox="[151,1420,181,1531]" captionTargetId="figure-16@33.[151,1435,181,1543]" captionTargetPageId="33" captionText="FIGURES 119126. Anhoraeomorphus globosus (Franz), holotype. Habitus in dorsal view (119); head in anterodorsal view (120); head and prothorax in lateral view (only bristles on head are shown) (121); antenna (122); aedeagus in ventral (123, 125) and lateral (124, 126) views." figureDoi="http://doi.org/10.5281/zenodo.13745963" httpUri="https://zenodo.org/record/13745963/files/figure.png" pageId="32" pageNumber="33">Fig. 120</figureCitation>
) in anterodorsal view oval, broadest at eyes, HL
<quantity id="4CB6E3A3FFD05020FC27FAB1FC3D41EB" box="[915,1016,1405,1432]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="32" pageNumber="33" unit="mm" value="0.5">0.50 mm</quantity>
, HW
<quantity id="4CB6E3A3FFD05020FB8CFAB1FB5A41EB" box="[1080,1183,1405,1432]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="32" pageNumber="33" unit="mm" value="0.45">0.45 mm</quantity>
; temple in lateral view (
<figureCitation id="137552C3FFD05020FF2BFA6DFF3A41C8" box="[159,255,1441,1467]" captionStart="FIGURES 119126" captionStartId="33.[151,264,1567,1591]" captionTargetBox="[151,1420,181,1531]" captionTargetId="figure-16@33.[151,1435,181,1543]" captionTargetPageId="33" captionText="FIGURES 119126. Anhoraeomorphus globosus (Franz), holotype. Habitus in dorsal view (119); head in anterodorsal view (120); head and prothorax in lateral view (only bristles on head are shown) (121); antenna (122); aedeagus in ventral (123, 125) and lateral (124, 126) views." figureDoi="http://doi.org/10.5281/zenodo.13745963" httpUri="https://zenodo.org/record/13745963/files/figure.png" pageId="32" pageNumber="33">Fig. 121</figureCitation>
) slightly more than twice as long as eye; vertex and frons confluent, together weakly convex and slightly elongate; vertex posteriorly broadly rounded; each eye conspicuously small and weakly convex, nearly circular, weakly projecting from head silhouette. Punctures on vertex fine, inconspicuous; setae short, dense and suberect; frons and vertex densely covered with thick bristles directed posteriorly. Antenna (
<figureCitation id="137552C3FFD05020FBE9F9C1FB054254" box="[1117,1216,1549,1575]" captionStart="FIGURES 119126" captionStartId="33.[151,264,1567,1591]" captionTargetBox="[151,1420,181,1531]" captionTargetId="figure-16@33.[151,1435,181,1543]" captionTargetPageId="33" captionText="FIGURES 119126. Anhoraeomorphus globosus (Franz), holotype. Habitus in dorsal view (119); head in anterodorsal view (120); head and prothorax in lateral view (only bristles on head are shown) (121); antenna (122); aedeagus in ventral (123, 125) and lateral (124, 126) views." figureDoi="http://doi.org/10.5281/zenodo.13745963" httpUri="https://zenodo.org/record/13745963/files/figure.png" pageId="32" pageNumber="33">Fig. 122</figureCitation>
) slender but short, as long as half BL, AnL
<quantity id="4CB6E3A3FFD05020FE05F9FDFDDC423F" box="[433,537,1585,1612]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="32" pageNumber="33" unit="mm" value="1.25">1.25 mm</quantity>
, antennomeres 16 each elongate, 7 about as long as broad, 810 each slightly transverse, 11 as long as 9 and 10 combined, slightly wider than 10, about 1.8 times as long as broad.
</paragraph>
<paragraph id="8BF14E46FFD05020FF73F9B6FEEA4350" blockId="32.[151,1437,901,1935]" pageId="32" pageNumber="33">
Pronotum bell-shaped, broadest near anterior third; PL
<quantity id="4CB6E3A3FFD05020FCE6F9B5FC7842E7" box="[850,957,1657,1684]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="32" pageNumber="33" unit="mm" value="0.7">0.70 mm</quantity>
, PW
<quantity id="4CB6E3A3FFD05020FBB6F9B5FBAB42E7" box="[1026,1134,1657,1684]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="32" pageNumber="33" unit="mm" value="0.58">0.58 mm</quantity>
. Anterior margin strongly rounded, lateral margins rounded in anterior half, weakly sinuate in posterior half; posterior corners blunt; posterior margin nearly straight. Base with one pair of conspicuously large and deep pits, sublateral carinae short but distinct. Disc with fine, inconspicuous punctures; setae dense, short and suberect, lateral surface of pronotum with dense thick bristles.
</paragraph>
<paragraph id="8BF14E46FFD05020FF73F8E1FE1443FC" blockId="32.[151,1437,901,1935]" pageId="32" pageNumber="33">
Elytra together oval, broadest slightly in front of middle; EL
<quantity id="4CB6E3A3FFD05020FCD9F8E1FC17433B" box="[877,978,1837,1864]" metricMagnitude="-3" metricUnit="m" metricValue="1.33" pageId="32" pageNumber="33" unit="mm" value="1.33">1.33 mm</quantity>
, EW
<quantity id="4CB6E3A3FFD05020FBBBF8E1FBB1433B" box="[1039,1140,1837,1864]" metricMagnitude="-3" metricUnit="m" metricValue="1.03" pageId="32" pageNumber="33" unit="mm" value="1.03">1.03 mm</quantity>
, EI 1.29; humeral calli not marked, basal elytral foveae indiscernible. Punctures dense but superficial and diffuse, inconspicuous; setae sparse, short and nearly recumbent.
</paragraph>
<caption id="DF311ECEFFD15021FF23F9D3FE7F420C" ID-DOI="http://doi.org/10.5281/zenodo.13745963" ID-Zenodo-Dep="13745963" httpUri="https://zenodo.org/record/13745963/files/figure.png" pageId="33" pageNumber="34" startId="33.[151,264,1567,1591]" targetBox="[151,1420,181,1531]" targetPageId="33" targetType="figure">
<paragraph id="8BF14E46FFD15021FF23F9D3FE7F420C" blockId="33.[151,1436,1567,1663]" pageId="33" pageNumber="34">
<emphasis id="B93A9254FFD15021FF23F9D3FEAB4244" bold="true" box="[151,366,1567,1591]" pageId="33" pageNumber="34">FIGURES 119126.</emphasis>
<taxonomicName id="4C4E35C5FFD15021FEC1F9D3FD2C4244" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[373,745,1567,1591]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="globosus">
<emphasis id="B93A9254FFD15021FEC1F9D3FD514244" box="[373,660,1567,1591]" italics="true" pageId="33" pageNumber="34">Anhoraeomorphus globosus</emphasis>
(Franz)
</taxonomicName>
, holotype. Habitus in dorsal view (
<emphasis id="B93A9254FFD15021FBE2F9D3FBBE4244" bold="true" box="[1110,1147,1567,1591]" pageId="33" pageNumber="34">119</emphasis>
); head in anterodorsal view (
<emphasis id="B93A9254FFD15021FF14F98FFF034228" bold="true" box="[160,198,1603,1627]" pageId="33" pageNumber="34">120</emphasis>
); head and prothorax in lateral view (only bristles on head are shown) (
<emphasis id="B93A9254FFD15021FC23F98FFC784228" bold="true" box="[919,957,1603,1627]" pageId="33" pageNumber="34">121</emphasis>
); antenna (
<emphasis id="B93A9254FFD15021FB9BF98FFB904228" bold="true" box="[1071,1109,1603,1627]" pageId="33" pageNumber="34">122</emphasis>
); aedeagus in ventral (
<emphasis id="B93A9254FFD15021FA88F98FFA514228" bold="true" box="[1340,1428,1603,1627]" pageId="33" pageNumber="34">123, 125</emphasis>
) and lateral (
<emphasis id="B93A9254FFD15021FEA4F9ABFEAC420C" bold="true" box="[272,361,1639,1663]" pageId="33" pageNumber="34">124, 126</emphasis>
) views.
</paragraph>
</caption>
<paragraph id="8BF14E46FFD15021FF73F908FE434357" blockId="33.[151,1437,1731,2042]" pageId="33" pageNumber="34">Legs moderately long, slender, unmodified except tibiae: protibiae with distal region curved mesally and with large distomesal concave patch of dense adhesive setae; mesotibiae with distal region curved mesally; and metatibiae indistinctly recurved.</paragraph>
<paragraph id="8BF14E46FFD15021FF73F8E2FE324389" blockId="33.[151,1437,1731,2042]" pageId="33" pageNumber="34">
Aedeagus (
<figureCitation id="137552C3FFD15021FEF0F8E2FE1B433B" box="[324,478,1838,1864]" captionStart="FIGURES 119126" captionStartId="33.[151,264,1567,1591]" captionTargetBox="[151,1420,181,1531]" captionTargetId="figure-16@33.[151,1435,181,1543]" captionTargetPageId="33" captionText="FIGURES 119126. Anhoraeomorphus globosus (Franz), holotype. Habitus in dorsal view (119); head in anterodorsal view (120); head and prothorax in lateral view (only bristles on head are shown) (121); antenna (122); aedeagus in ventral (123, 125) and lateral (124, 126) views." figureDoi="http://doi.org/10.5281/zenodo.13745963" httpUri="https://zenodo.org/record/13745963/files/figure.png" pageId="33" pageNumber="34">Figs 123126</figureCitation>
) stout, AeL
<quantity id="4CB6E3A3FFD15021FDD2F8E1FD09433B" box="[614,716,1837,1864]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="33" pageNumber="34" unit="mm" value="0.45">0.45 mm</quantity>
; in ventral view pear-shaped, broadest in sub-basal third, distally gradually narrowing to truncate apex with slightly arcuate distal margin; in lateral view apical region of dorsal wall strongly bent dorsally; endophallus with asymmetrical and large distolateral sclerites and with strongly asymmetrical assemblage of median sclerotized structures, distolateral sclerites broadly separated, each with subtriangular and blunt apical region; parameres broad and their apices not projecting beyond aedeagal apex, each paramere with apical group of 45 short setae.
</paragraph>
<paragraph id="8BF14E46FFD25022FF73FF5BFE5B44C2" blockId="34.[151,1436,151,357]" box="[199,414,151,177]" pageId="34" pageNumber="35">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD25022FF73FF76FC6044A6" box="[199,933,186,213]" pageId="34" pageNumber="35" type="distribution">
<paragraph id="8BF14E46FFD25022FF73FF76FC6044A6" blockId="34.[151,1436,151,357]" box="[199,933,186,213]" pageId="34" pageNumber="35">
<emphasis id="B93A9254FFD25022FF73FF76FEA444A7" bold="true" box="[199,353,186,212]" pageId="34" pageNumber="35">Distribution.</emphasis>
Southern
<collectingCountry id="F3590ED6FFD25022FE60FF77FD9E44A6" box="[468,603,187,213]" name="Madagascar" pageId="34" pageNumber="35">Madagascar</collectingCountry>
(Matsiatra Ambony Region).
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD25022FF73FF13FEC04516" pageId="34" pageNumber="35" type="discussion">
<paragraph id="8BF14E46FFD25022FF73FF13FC1C448A" blockId="34.[151,1436,151,357]" box="[199,985,223,249]" pageId="34" pageNumber="35">
<emphasis id="B93A9254FFD25022FF73FF13FEFE448A" bold="true" box="[199,315,223,249]" pageId="34" pageNumber="35">Remarks.</emphasis>
<bibRefCitation id="EFDF33B7FFD25022FEF6FF13FE24448A" author="Franz, H." box="[322,481,223,249]" pageId="34" pageNumber="35" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986b)</bibRefCitation>
misidentified the
<typeStatus id="54F5F0E4FFD25022FD18FF13FCCB448A" box="[684,782,223,249]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
male as a female.
</paragraph>
<paragraph id="8BF14E46FFD25022FF73FECFFEC04516" blockId="34.[151,1436,151,357]" pageId="34" pageNumber="35">
<taxonomicName id="4C4E35C5FFD25022FF73FECFFDC2456E" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[199,519,259,285]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="globosus">
<emphasis id="B93A9254FFD25022FF73FECFFDC2456E" box="[199,519,259,285]" italics="true" pageId="34" pageNumber="35">Anhoraeomorphus globosus</emphasis>
</taxonomicName>
can be easily distinguished from all remaining congeners by the body shape, tempora nearly twice as long as small eyes, slender legs with unmodified metatibiae, and unique structures of the aedeagus.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="DB2A495A410CF0A20FB025C8630E1948">The curse of Horaeomorphus (almost) lifted. Revision of misplaced species from Madagascar, Comoros and Mascarenes (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<subSubSection id="C3541DCDFFD25022FF23FE7AFD7845A2" box="[151,701,438,465]" pageId="34" pageNumber="35" type="nomenclature">
<paragraph id="8BF14E46FFD25022FF23FE7AFD7845A2" blockId="34.[151,701,438,501]" box="[151,701,438,465]" pageId="34" pageNumber="35">
<heading id="D0B9F92AFFD25022FF23FE7AFD7845A2" bold="true" box="[151,701,438,465]" fontSize="11" level="2" pageId="34" pageNumber="35" reason="4">
<emphasis id="B93A9254FFD25022FF23FE7AFD7845A2" bold="true" box="[151,701,438,465]" pageId="34" pageNumber="35">
<taxonomicName id="4C4E35C5FFD25022FF23FE7AFD8045A3" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[151,581,438,465]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="inermis" status="comb. nov.">
<emphasis id="B93A9254FFD25022FF23FE7AFE1B45A2" bold="true" box="[151,478,438,465]" italics="true" pageId="34" pageNumber="35">Anhoraeomorphus inermis</emphasis>
(Franz)
</taxonomicName>
,
<taxonomicNameLabel id="A2092F2FFFD25022FDE7FE7BFD7845A2" box="[595,701,439,465]" pageId="34" pageNumber="35" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3541DCDFFD25022FF23FE17FE864586" box="[151,323,475,501]" pageId="34" pageNumber="35" type="description">
<paragraph id="8BF14E46FFD25022FF23FE17FE864586" blockId="34.[151,701,438,501]" box="[151,323,475,501]" pageId="34" pageNumber="35">
(
<figureCitation id="137552C3FFD25022FF2BFE17FEFE4586" box="[159,315,475,501]" captionStart="FIGURES 127130" captionStartId="35.[151,264,1033,1057]" captionTargetBox="[151,1416,237,1009]" captionTargetId="figure-37@35.[151,1435,237,1009]" captionTargetPageId="35" captionText="FIGURES 127130. Anhoraeomorphus inermis (Franz), paratype. Habitus in dorsal view (127); head in anterodorsal view (128); head and prothorax in lateral view (only bristles on head are shown) (129); antenna (130)." figureDoi="http://doi.org/10.5281/zenodo.13745965" httpUri="https://zenodo.org/record/13745965/files/figure.png" pageId="34" pageNumber="35">Figs 127130</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD25022FF23FDD3FD934644" box="[151,598,543,567]" pageId="34" pageNumber="35" type="reference_group">
<paragraph id="8BF14E46FFD25022FF23FDD3FD934644" blockId="34.[151,598,543,567]" box="[151,598,543,567]" pageId="34" pageNumber="35">
<treatmentCitationGroup id="AB5E6968FFD25022FF23FDD3FD934644" box="[151,598,543,567]" pageId="34" pageNumber="35">
<treatmentCitation id="0AEF6857FFD25022FF23FDD3FD974644" author="Franz, H." box="[151,594,543,567]" page="163" pageId="34" pageNumber="35" year="1986">
<taxonomicName id="4C4E35C5FFD25022FF23FDD3FD974644" ID-CoL="3MNK9" authority="Franz, 1986 b: 163" authorityName="Franz" authorityPageNumber="163" authorityYear="1986" box="[151,594,543,567]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="inermis">
<emphasis id="B93A9254FFD25022FF23FDD3FE554644" box="[151,400,543,567]" italics="true" pageId="34" pageNumber="35">Horaeomorphus inermis</emphasis>
<bibRefCitation id="EFDF33B7FFD25022FE22FDECFD974644" author="Franz, H." box="[406,594,543,567]" pageId="34" pageNumber="35" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 163</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD25022FF23FDA9FE6447B0" pageId="34" pageNumber="35" type="materials_examined">
<paragraph id="8BF14E46FFD25022FF23FDA9FE1B46B4" blockId="34.[151,1437,613,2008]" pageId="34" pageNumber="35">
<emphasis id="B93A9254FFD25022FF23FDA9FE60460C" bold="true" box="[151,421,613,639]" pageId="34" pageNumber="35">Type material studied.</emphasis>
<materialsCitation id="3B26441BFFD25022FE19FDA9FE1E46B4" collectionCode="NHMW" collectorName="Ch. Alluaud" country="Madagascar" determinerName="H. Franz" location="Madagascar" pageId="34" pageNumber="35" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<emphasis id="B93A9254FFD25022FE19FDA9FD11460C" bold="true" box="[429,724,613,639]" pageId="34" pageNumber="35">
<typeStatus id="54F5F0E4FFD25022FE19FDA9FDE0460C" box="[429,549,613,639]" pageId="34" pageNumber="35" type="paratype">Paratypes</typeStatus>
(
<collectingCountry id="F3590ED6FFD25022FD81FDA9FD09460C" box="[565,716,613,639]" name="Madagascar" pageId="34" pageNumber="35">Madagascar</collectingCountry>
)
</emphasis>
:
<specimenCount id="9D4885CFFFD25022FD50FDA9FCE046F3" box="[740,805,613,640]" count="2" pageId="34" pageNumber="35" type="female">2 ♀♀</specimenCount>
, two labels: “Madagascar / Diego-Suarez /
<collectorName id="26BB2B90FFD25022FAA0FDA9FA59460C" box="[1300,1436,613,639]" pageId="34" pageNumber="35">Ch. Alluaud</collectorName>
1893” [white, printed], “
<taxonomicName id="4C4E35C5FFD25022FE1CFD45FD9A46D0" box="[424,607,649,675]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/ inermis m. / det.
<determinerName id="7E2DA282FFD25022FC9EFD45FC4D46D0" box="[810,904,649,675]" pageId="34" pageNumber="35">H.Franz</determinerName>
” [yellowish, handwritten and printed], “Typus” [red, handwritten] (
<collectionCode id="ED5FD683FFD25022FEC4FD61FE1646B4" box="[368,467,685,711]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="34" pageNumber="35" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF14E46FFD25022FF73FD1DFE6447B0" blockId="34.[151,1437,613,2008]" pageId="34" pageNumber="35">
<emphasis id="B93A9254FFD25022FF73FD1DFE5B4698" bold="true" box="[199,414,721,747]" pageId="34" pageNumber="35">Revised diagnosis.</emphasis>
Body (
<figureCitation id="137552C3FFD25022FE5FFD1DFD824698" box="[491,583,721,747]" captionStart="FIGURES 127130" captionStartId="35.[151,264,1033,1057]" captionTargetBox="[151,1416,237,1009]" captionTargetId="figure-37@35.[151,1435,237,1009]" captionTargetPageId="35" captionText="FIGURES 127130. Anhoraeomorphus inermis (Franz), paratype. Habitus in dorsal view (127); head in anterodorsal view (128); head and prothorax in lateral view (only bristles on head are shown) (129); antenna (130)." figureDoi="http://doi.org/10.5281/zenodo.13745965" httpUri="https://zenodo.org/record/13745965/files/figure.png" pageId="34" pageNumber="35">Fig.127</figureCitation>
) slender; head (
<figureCitation id="137552C3FFD25022FD45FD1DFC4F4698" box="[753,906,721,747]" captionStart="FIGURES 127130" captionStartId="35.[151,264,1033,1057]" captionTargetBox="[151,1416,237,1009]" captionTargetId="figure-37@35.[151,1435,237,1009]" captionTargetPageId="35" captionText="FIGURES 127130. Anhoraeomorphus inermis (Franz), paratype. Habitus in dorsal view (127); head in anterodorsal view (128); head and prothorax in lateral view (only bristles on head are shown) (129); antenna (130)." figureDoi="http://doi.org/10.5281/zenodo.13745965" httpUri="https://zenodo.org/record/13745965/files/figure.png" pageId="34" pageNumber="35">Figs 128129</figureCitation>
) round, with vertex posteriorly broadly rounded; pronotum (
<figureCitation id="137552C3FFD25022FEA2FD39FEB2477C" box="[278,375,757,783]" captionStart="FIGURES 127130" captionStartId="35.[151,264,1033,1057]" captionTargetBox="[151,1416,237,1009]" captionTargetId="figure-37@35.[151,1435,237,1009]" captionTargetPageId="35" captionText="FIGURES 127130. Anhoraeomorphus inermis (Franz), paratype. Habitus in dorsal view (127); head in anterodorsal view (128); head and prothorax in lateral view (only bristles on head are shown) (129); antenna (130)." figureDoi="http://doi.org/10.5281/zenodo.13745965" httpUri="https://zenodo.org/record/13745965/files/figure.png" pageId="34" pageNumber="35">Fig. 127</figureCitation>
) bell-shaped, broadest near anterior third, with one pair of conspicuously large, transversely elongate antebasal pits; antennomeres 37 (
<figureCitation id="137552C3FFD25022FDCAFCD5FD184740" box="[638,733,793,819]" captionStart="FIGURES 127130" captionStartId="35.[151,264,1033,1057]" captionTargetBox="[151,1416,237,1009]" captionTargetId="figure-37@35.[151,1435,237,1009]" captionTargetPageId="35" captionText="FIGURES 127130. Anhoraeomorphus inermis (Franz), paratype. Habitus in dorsal view (127); head in anterodorsal view (128); head and prothorax in lateral view (only bristles on head are shown) (129); antenna (130)." figureDoi="http://doi.org/10.5281/zenodo.13745965" httpUri="https://zenodo.org/record/13745965/files/figure.png" pageId="34" pageNumber="35">Fig. 130</figureCitation>
) each about as long as broad, 810 each slightly transverse; all femora similarly slender; metatibiae in male unmodified; aedeagus (
<bibRefCitation id="EFDF33B7FFD25022FC27FCF1FBE34724" author="Franz, H." box="[915,1062,829,855]" pageId="34" pageNumber="35" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz 1986b</bibRefCitation>
: fig. 147) in dorsal view suboval, with sub-parallel lateral margins and broadly rounded apex, distolateral sclerites asymmetrical, strongly elongate, broadly separated and each with subtriangular apex, (parameres were illustrated without setae, but it may be an artefact of preparation).
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD25022FF73FC01FE6F4357" pageId="34" pageNumber="35" type="description">
<paragraph id="8BF14E46FFD25022FF73FC01FDFE407F" blockId="34.[151,1437,613,2008]" pageId="34" pageNumber="35">
<emphasis id="B93A9254FFD25022FF73FC01FEB04794" bold="true" box="[199,373,973,999]" pageId="34" pageNumber="35">Redescription.</emphasis>
Body of female (
<figureCitation id="137552C3FFD25022FDF5FC01FD664794" box="[577,675,973,999]" captionStart="FIGURES 127130" captionStartId="35.[151,264,1033,1057]" captionTargetBox="[151,1416,237,1009]" captionTargetId="figure-37@35.[151,1435,237,1009]" captionTargetPageId="35" captionText="FIGURES 127130. Anhoraeomorphus inermis (Franz), paratype. Habitus in dorsal view (127); head in anterodorsal view (128); head and prothorax in lateral view (only bristles on head are shown) (129); antenna (130)." figureDoi="http://doi.org/10.5281/zenodo.13745965" httpUri="https://zenodo.org/record/13745965/files/figure.png" pageId="34" pageNumber="35">Fig. 127</figureCitation>
) flattened, moderately dark brown, tarsi and palps slightly lighter; setae light brown; BL
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.
</paragraph>
<paragraph id="8BF14E46FFD25022FF73FBD9FDC24158" blockId="34.[151,1437,613,2008]" pageId="34" pageNumber="35">
Head (
<figureCitation id="137552C3FFD25022FEA7FBD9FEB6405C" box="[275,371,1045,1071]" captionStart="FIGURES 127130" captionStartId="35.[151,264,1033,1057]" captionTargetBox="[151,1416,237,1009]" captionTargetId="figure-37@35.[151,1435,237,1009]" captionTargetPageId="35" captionText="FIGURES 127130. Anhoraeomorphus inermis (Franz), paratype. Habitus in dorsal view (127); head in anterodorsal view (128); head and prothorax in lateral view (only bristles on head are shown) (129); antenna (130)." figureDoi="http://doi.org/10.5281/zenodo.13745965" httpUri="https://zenodo.org/record/13745965/files/figure.png" pageId="34" pageNumber="35">Fig. 128</figureCitation>
) in anterodorsal view round, broadest at eyes, HL
<quantity id="4CB6E3A3FFD25022FC07FBD9FB9F4043" box="[947,1114,1045,1072]" metricMagnitude="-4" metricUnit="m" metricValue="3.3999999999999995" metricValueMax="3.5" metricValueMin="3.3" pageId="34" pageNumber="35" unit="mm" value="0.33999999999999997" valueMax="0.35" valueMin="0.33">0.330.35 mm</quantity>
, HW
<quantity id="4CB6E3A3FFD25022FB28FBD9FA814043" box="[1180,1348,1045,1072]" metricMagnitude="-4" metricUnit="m" metricValue="3.65" metricValueMax="3.8" metricValueMin="3.5" pageId="34" pageNumber="35" unit="mm" value="0.365" valueMax="0.38" valueMin="0.35">0.350.38 mm</quantity>
; temple in lateral view (
<figureCitation id="137552C3FFD25022FEF3FBF5FE634020" box="[327,422,1081,1107]" captionStart="FIGURES 127130" captionStartId="35.[151,264,1033,1057]" captionTargetBox="[151,1416,237,1009]" captionTargetId="figure-37@35.[151,1435,237,1009]" captionTargetPageId="35" captionText="FIGURES 127130. Anhoraeomorphus inermis (Franz), paratype. Habitus in dorsal view (127); head in anterodorsal view (128); head and prothorax in lateral view (only bristles on head are shown) (129); antenna (130)." figureDoi="http://doi.org/10.5281/zenodo.13745965" httpUri="https://zenodo.org/record/13745965/files/figure.png" pageId="34" pageNumber="35">Fig. 129</figureCitation>
) slightly shorter than eye; vertex and frons confluent, together weakly convex and slightly transverse; vertex posteriorly broadly rounded; each eye large and weakly convex, oval with slightly emarginate posterior margin, weakly projecting from head silhouette. Punctures on vertex fine, inconspicuous; setae short, dense and suberect; frons and vertex densely covered with thick bristles directed posteriorly. Antenna (
<figureCitation id="137552C3FFD25022FA86FB69FA5140CC" box="[1330,1428,1189,1215]" captionStart="FIGURES 119126" captionStartId="33.[151,264,1567,1591]" captionTargetBox="[151,1420,181,1531]" captionTargetId="figure-16@33.[151,1435,181,1543]" captionTargetPageId="33" captionText="FIGURES 119126. Anhoraeomorphus globosus (Franz), holotype. Habitus in dorsal view (119); head in anterodorsal view (120); head and prothorax in lateral view (only bristles on head are shown) (121); antenna (122); aedeagus in ventral (123, 125) and lateral (124, 126) views." figureDoi="http://doi.org/10.5281/zenodo.13745963" httpUri="https://zenodo.org/record/13745963/files/figure.png" pageId="34" pageNumber="35">Fig. 122</figureCitation>
) slender but short, slightly shorter than half BL, AnL 0.850,88 mm, antennomeres 12 each elongate, 37 each about as long as broad, 810 each slightly transverse, 11 slightly shorter than 9 and 10 combined, as wide as 10, about 1.7 times as long as broad.
</paragraph>
<paragraph id="8BF14E46FFD25022FF73FAFAFE6B41AC" blockId="34.[151,1437,613,2008]" pageId="34" pageNumber="35">
Pronotum bell-shaped, broadest near anterior third; PL
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, PW
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. Anterior margin strongly rounded, lateral margins rounded in anterior half, distinctly sinuate in posterior half; posterior corners blunt; posterior margin nearly straight. Base with one pair of conspicuously large, strongly transverse pits, sublateral carinae short but distinct. Disc with fine, inconspicuous punctures; setae dense, short and suberect, lateral surface of pronotum with dense thick bristles.
</paragraph>
<paragraph id="8BF14E46FFD25022FF73FA25FAE24238" blockId="34.[151,1437,613,2008]" pageId="34" pageNumber="35">
Elytra together oval, broadest indistinctly in front of middle; EL
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, EW
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, EI 1.41 1.43; humeral calli weakly elevated, basal elytral foveae vestigial, barely discernible. Punctures dense but superficial and diffuse, so that elytral surface is slightly less glossy than pronotum; setae sparse, short and suberect.
</paragraph>
<paragraph id="8BF14E46FFD25022FF73F99AFD75421C" blockId="34.[151,1437,613,2008]" box="[199,688,1621,1648]" pageId="34" pageNumber="35">Legs moderately long, slender, unmodified.</paragraph>
<paragraph id="8BF14E46FFD25022FF73F9B5FE6F4357" blockId="34.[151,1437,613,2008]" pageId="34" pageNumber="35">
Male. Unstudied (See Remarks). According to
<bibRefCitation id="EFDF33B7FFD25022FD62F9B6FCBD42E0" author="Franz, H." box="[726,888,1657,1684]" pageId="34" pageNumber="35" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986b)</bibRefCitation>
, metatibiae unmodified; aedeagus in dorsal view suboval in shape, with sub-parallel lateral margins and broadly rounded apex, distolateral sclerites asymmetrical, strongly elongate, broadly separated and each with subtriangular apex, median sclerotized endophallic structures strongly asymmetrical, parameres slender, their apices not projecting beyond apex of median lobe, their apices were illustrated without setae.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD25022FF73F8E1FC884334" box="[199,845,1837,1863]" pageId="34" pageNumber="35" type="distribution">
<paragraph id="8BF14E46FFD25022FF73F8E1FC884334" blockId="34.[151,1437,613,2008]" box="[199,845,1837,1863]" pageId="34" pageNumber="35">
<emphasis id="B93A9254FFD25022FF73F8E1FEA44334" bold="true" box="[199,353,1837,1863]" pageId="34" pageNumber="35">Distribution.</emphasis>
Northernmost
<collectingCountry id="F3590ED6FFD25022FDBEF8E1FD544334" box="[522,657,1837,1863]" name="Madagascar" pageId="34" pageNumber="35">Madagascar</collectingCountry>
(Diana Region).
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD25023FF73F89DFED444A6" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35" type="discussion">
<paragraph id="8BF14E46FFD25023FF73F89DFED444A6" blockId="34.[151,1437,613,2008]" lastBlockId="35.[151,1436,151,213]" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
<emphasis id="B93A9254FFD25022FF73F89DFEFE4318" bold="true" box="[199,315,1873,1899]" pageId="34" pageNumber="35">Remarks.</emphasis>
<bibRefCitation id="EFDF33B7FFD25022FEF4F89EFE184318" author="Franz, H." box="[320,477,1873,1900]" pageId="34" pageNumber="35" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986b)</bibRefCitation>
studied
<specimenCount id="9D4885CFFFD25022FD83F89DFD294318" box="[567,748,1873,1899]" count="8" pageId="34" pageNumber="35" type="generic">eight specimens</specimenCount>
, of which six, including the
<typeStatus id="54F5F0E4FFD25022FBAFF89DFB454318" box="[1051,1152,1873,1899]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
, are deposited in Muséum National d'Histoire Naturelle, Paris and were not available for detailed examination. The
<specimenCount id="9D4885CFFFD25022FB12F8BAFAE343FC" box="[1190,1318,1909,1936]" count="2" pageId="34" pageNumber="35" type="female">two female</specimenCount>
<typeStatus id="54F5F0E4FFD25022FA85F8BAFA5943E3" box="[1329,1436,1910,1936]" pageId="34" pageNumber="35" type="paratype">paratypes</typeStatus>
examined during the present study are small-bodied, but otherwise unremarkable and difficult to distinguish from females of other small
<taxonomicName id="4C4E35C5FFD25022FE23F871FD7A43A4" box="[407,703,1981,2007]" pageId="34" pageNumber="35">
<emphasis id="B93A9254FFD25022FE23F871FDA243A4" box="[407,615,1981,2007]" italics="true" pageId="34" pageNumber="35">Anhoraeomorphus</emphasis>
species
</taxonomicName>
with the pronotum broadest in front of middle. The illustration of aedeagus given by
<bibRefCitation id="EFDF33B7FFD35023FEDBFF5BFDCF44C2" author="Franz, H." box="[367,522,151,177]" pageId="35" pageNumber="36" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986b</bibRefCitation>
: fig. 147) must be compared with genital organs of studied specimens to identify
<taxonomicName id="4C4E35C5FFD35023FF23FF77FECF44A7" authorityName="Franz" authorityYear="1986" box="[151,266,187,213]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="inermis">
<emphasis id="B93A9254FFD35023FF23FF77FECF44A7" box="[151,266,187,213]" italics="true" pageId="35" pageNumber="36">A. inermis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,264 @@
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<paragraph id="8BF14E46FFD35023FF23FB5BFCF440C1" blockId="35.[151,817,1175,1238]" box="[151,817,1175,1202]" pageId="35" pageNumber="36">
<heading id="D0B9F92AFFD35023FF23FB5BFCF440C1" bold="true" box="[151,817,1175,1202]" fontSize="11" level="2" pageId="35" pageNumber="36" reason="4">
<emphasis id="B93A9254FFD35023FF23FB5BFCF440C1" bold="true" box="[151,817,1175,1202]" pageId="35" pageNumber="36">
<taxonomicName id="4C4E35C5FFD35023FF23FB5BFD7C40C2" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[151,697,1175,1202]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="lambomakandroi" status="comb. nov.">
<emphasis id="B93A9254FFD35023FF23FB5BFD9740C1" bold="true" box="[151,594,1175,1202]" italics="true" pageId="35" pageNumber="36">Anhoraeomorphus lambomakandroi</emphasis>
(Franz)
</taxonomicName>
,
<taxonomicNameLabel id="A2092F2FFFD35023FD72FB54FCF440C1" box="[710,817,1176,1202]" pageId="35" pageNumber="36" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD35023FF23FB77FE8640A5" box="[151,323,1211,1238]" pageId="35" pageNumber="36" type="description">
<paragraph id="8BF14E46FFD35023FF23FB77FE8640A5" blockId="35.[151,817,1175,1238]" box="[151,323,1211,1238]" pageId="35" pageNumber="36">
(
<figureCitation id="137552C3FFD35023FF2BFB77FEFF40A5" box="[159,314,1211,1238]" captionStart="FIGURES 131138" captionStartId="36.[151,264,1837,1861]" captionTargetBox="[151,1423,237,1794]" captionTargetId="figure-67@36.[151,1435,237,1814]" captionTargetPageId="36" captionText="FIGURES 131138. Anhoraeomorphus lambomakandroi (Franz), holotype. Habitus in dorsal view (131); head in anterodorsal view (132); head and prothorax in lateral view (only bristles on head are shown) (133); antenna (134); aedeagus in ventral (135, 137) and lateral (136, 138) views." figureDoi="http://doi.org/10.5281/zenodo.13745967" httpUri="https://zenodo.org/record/13745967/files/figure.png" pageId="35" pageNumber="36">Figs 131138</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD35023FF23FACCFD73416B" box="[151,694,1280,1304]" pageId="35" pageNumber="36" type="reference_group">
<paragraph id="8BF14E46FFD35023FF23FACCFD73416B" blockId="35.[151,694,1280,1304]" box="[151,694,1280,1304]" pageId="35" pageNumber="36">
<treatmentCitationGroup id="AB5E6968FFD35023FF23FACCFD73416B" box="[151,694,1280,1304]" pageId="35" pageNumber="36">
<treatmentCitation id="0AEF6857FFD35023FF23FACCFD77416B" author="Franz, H." box="[151,690,1280,1304]" page="154" pageId="35" pageNumber="36" year="1986">
<taxonomicName id="4C4E35C5FFD35023FF23FACCFD77416B" ID-CoL="3MNKG" authority="Franz, 1986 b: 154" authorityName="Franz" authorityPageNumber="154" authorityYear="1986" box="[151,690,1280,1304]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="lambomakandroi">
<emphasis id="B93A9254FFD35023FF23FACCFE35416B" box="[151,496,1280,1304]" italics="true" pageId="35" pageNumber="36">Horaeomorphus lambomakandroi</emphasis>
<bibRefCitation id="EFDF33B7FFD35023FE42FACCFD77416B" author="Franz, H." box="[502,690,1280,1304]" pageId="35" pageNumber="36" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 154</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD35023FF23FA8AFE67429E" pageId="35" pageNumber="36" type="materials_examined">
<paragraph id="8BF14E46FFD35023FF23FA8AFE6641BF" blockId="35.[151,1437,1350,2025]" pageId="35" pageNumber="36">
<emphasis id="B93A9254FFD35023FF23FA8AFE654113" bold="true" box="[151,416,1350,1376]" pageId="35" pageNumber="36">Type material studied.</emphasis>
<materialsCitation id="3B26441BFFD35023FE12FA8AFE6641BF" collectionCode="VII-, NHMW" country="Madagascar" determinerName="H. Franz" elevation="500" location="MADAGASCAR" pageId="35" pageNumber="36" specimenCount="58" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B93A9254FFD35023FE12FA8AFD7A4113" bold="true" box="[422,703,1350,1376]" pageId="35" pageNumber="36">
<typeStatus id="54F5F0E4FFD35023FE12FA8AFDD64113" box="[422,531,1350,1376]" pageId="35" pageNumber="36" type="holotype">Holotype</typeStatus>
(Madagascar)
</emphasis>
:
<specimenCount id="9D4885CFFFD35023FD78FA8BFD264112" box="[716,739,1351,1377]" count="1" pageId="35" pageNumber="36" type="male"></specimenCount>
, four labels: “Madagascar Sud-Ouest / LAMBOMAKANDRO
<quantity id="4CB6E3A3FFD35023FF23FAA6FF1241F6" box="[151,215,1386,1413]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="35" pageNumber="36" unit="m" value="500.0">
<elevation id="0063A975FFD35023FF23FAA6FF1241F6" box="[151,215,1386,1413]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="35" pageNumber="36" unit="m" value="500.0">500m</elevation>
</quantity>
/ Tuléar / 18-
<collectionCode id="ED5FD683FFD35023FEC0FAA6FE6241F6" box="[372,423,1386,1413]" pageId="35" pageNumber="36">VII-</collectionCode>
<specimenCount id="9D4885CFFFD35023FE13FAA6FE2441F7" box="[423,481,1386,1413]" count="57" pageId="35" pageNumber="36" type="generic">57 P.</specimenCount>
Griv.” [white, printed], “INSTITUT / SCIENTIFIQUE /
<collectingCountry id="F3590ED6FFD35023FBDFFAA6FAF041F7" box="[1131,1333,1386,1412]" name="Madagascar" pageId="35" pageNumber="36">MADAGASCAR</collectingCountry>
” [bluish, printed], “
<taxonomicName id="4C4E35C5FFD35023FEB3FA42FE7B41DB" box="[263,446,1422,1448]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/ lambomakandroi m. / det.
<determinerName id="7E2DA282FFD35023FD45FA42FC8A41DB" box="[753,847,1422,1448]" pageId="35" pageNumber="36">H.Franz</determinerName>
” [yellowish, handwritten and printed], “Typus” [red, handwritten] (
<collectionCode id="ED5FD683FFD35023FE81FA7EFE5D41BF" box="[309,408,1458,1484]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="35" pageNumber="36" type="Museum">NHMW</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BF14E46FFD35023FF73FA1AFE67429E" blockId="35.[151,1437,1350,2025]" pageId="35" pageNumber="36">
<emphasis id="B93A9254FFD35023FF73FA1AFE634183" bold="true" box="[199,422,1494,1520]" pageId="35" pageNumber="36">Revised diagnosis.</emphasis>
Body (
<figureCitation id="137552C3FFD35023FDB0FA1AFDAC4183" box="[516,617,1494,1520]" captionStart="FIGURES 131138" captionStartId="36.[151,264,1837,1861]" captionTargetBox="[151,1423,237,1794]" captionTargetId="figure-67@36.[151,1435,237,1814]" captionTargetPageId="36" captionText="FIGURES 131138. Anhoraeomorphus lambomakandroi (Franz), holotype. Habitus in dorsal view (131); head in anterodorsal view (132); head and prothorax in lateral view (only bristles on head are shown) (133); antenna (134); aedeagus in ventral (135, 137) and lateral (136, 138) views." figureDoi="http://doi.org/10.5281/zenodo.13745967" httpUri="https://zenodo.org/record/13745967/files/figure.png" pageId="35" pageNumber="36">Fig. 131</figureCitation>
) slender; head (
<figureCitation id="137552C3FFD35023FC9FFA1AFC094183" box="[811,972,1494,1520]" captionStart="FIGURES 131138" captionStartId="36.[151,264,1837,1861]" captionTargetBox="[151,1423,237,1794]" captionTargetId="figure-67@36.[151,1435,237,1814]" captionTargetPageId="36" captionText="FIGURES 131138. Anhoraeomorphus lambomakandroi (Franz), holotype. Habitus in dorsal view (131); head in anterodorsal view (132); head and prothorax in lateral view (only bristles on head are shown) (133); antenna (134); aedeagus in ventral (135, 137) and lateral (136, 138) views." figureDoi="http://doi.org/10.5281/zenodo.13745967" httpUri="https://zenodo.org/record/13745967/files/figure.png" pageId="35" pageNumber="36">Figs 132133</figureCitation>
) round, with vertex posteriorly weakly rounded; pronotum (
<figureCitation id="137552C3FFD35023FECAFA36FE1B4267" box="[382,478,1530,1556]" captionStart="FIGURES 131138" captionStartId="36.[151,264,1837,1861]" captionTargetBox="[151,1423,237,1794]" captionTargetId="figure-67@36.[151,1435,237,1814]" captionTargetPageId="36" captionText="FIGURES 131138. Anhoraeomorphus lambomakandroi (Franz), holotype. Habitus in dorsal view (131); head in anterodorsal view (132); head and prothorax in lateral view (only bristles on head are shown) (133); antenna (134); aedeagus in ventral (135, 137) and lateral (136, 138) views." figureDoi="http://doi.org/10.5281/zenodo.13745967" httpUri="https://zenodo.org/record/13745967/files/figure.png" pageId="35" pageNumber="36">Fig. 131</figureCitation>
) bell-shaped, broadest near anterior third, with one pair of large, transverse antebasal pits; antennomere 3 (
<figureCitation id="137552C3FFD35023FE3EF9D2FE2E424B" box="[394,491,1566,1592]" captionStart="FIGURES 131138" captionStartId="36.[151,264,1837,1861]" captionTargetBox="[151,1423,237,1794]" captionTargetId="figure-67@36.[151,1435,237,1814]" captionTargetPageId="36" captionText="FIGURES 131138. Anhoraeomorphus lambomakandroi (Franz), holotype. Habitus in dorsal view (131); head in anterodorsal view (132); head and prothorax in lateral view (only bristles on head are shown) (133); antenna (134); aedeagus in ventral (135, 137) and lateral (136, 138) views." figureDoi="http://doi.org/10.5281/zenodo.13745967" httpUri="https://zenodo.org/record/13745967/files/figure.png" pageId="35" pageNumber="36">Fig. 134</figureCitation>
) about as long as broad, 47 each elongate, 810 each about as long as broad; all femora similarly slender; metatibiae in male modified, slightly recurved and broadened in distal region, broadened area with elongate outer impression bordered by sharp outer edge; aedeagus (
<figureCitation id="137552C3FFD35023FC5EF9AAFB4042F2" box="[1002,1157,1638,1665]" captionStart="FIGURES 131138" captionStartId="36.[151,264,1837,1861]" captionTargetBox="[151,1423,237,1794]" captionTargetId="figure-67@36.[151,1435,237,1814]" captionTargetPageId="36" captionText="FIGURES 131138. Anhoraeomorphus lambomakandroi (Franz), holotype. Habitus in dorsal view (131); head in anterodorsal view (132); head and prothorax in lateral view (only bristles on head are shown) (133); antenna (134); aedeagus in ventral (135, 137) and lateral (136, 138) views." figureDoi="http://doi.org/10.5281/zenodo.13745967" httpUri="https://zenodo.org/record/13745967/files/figure.png" pageId="35" pageNumber="36">Figs 135138</figureCitation>
) in ventral view broadest near middle, apical region of dorsal wall in lateral view weakly curved dorsally, distolateral endophallic sclerites broadly separated, each elongate and with blunt subtriangular apex, each paramere with group of five short subapical setae distant from apex.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD35025FF73F93AFE5B46BE" lastPageId="37" lastPageNumber="38" pageId="35" pageNumber="36" type="description">
<paragraph id="8BF14E46FFD35023FF73F93AFD914346" blockId="35.[151,1437,1350,2025]" pageId="35" pageNumber="36">
<emphasis id="B93A9254FFD35023FF73F93AFEB04363" bold="true" box="[199,373,1782,1808]" pageId="35" pageNumber="36">Redescription.</emphasis>
Body of male (
<figureCitation id="137552C3FFD35023FD87F93AFD524363" box="[563,663,1782,1808]" captionStart="FIGURES 131138" captionStartId="36.[151,264,1837,1861]" captionTargetBox="[151,1423,237,1794]" captionTargetId="figure-67@36.[151,1435,237,1814]" captionTargetPageId="36" captionText="FIGURES 131138. Anhoraeomorphus lambomakandroi (Franz), holotype. Habitus in dorsal view (131); head in anterodorsal view (132); head and prothorax in lateral view (only bristles on head are shown) (133); antenna (134); aedeagus in ventral (135, 137) and lateral (136, 138) views." figureDoi="http://doi.org/10.5281/zenodo.13745967" httpUri="https://zenodo.org/record/13745967/files/figure.png" pageId="35" pageNumber="36">Fig. 131</figureCitation>
) strongly convex, moderately dark brown, tarsi and palps slightly lighter; setae light brown; BL
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.
</paragraph>
<paragraph id="8BF14E46FFD35024FF73F8F2FB0644A6" blockId="35.[151,1437,1350,2025]" lastBlockId="36.[151,1436,151,213]" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
Head (
<figureCitation id="137552C3FFD35023FEA4F8F2FEAB432B" box="[272,366,1854,1880]" captionStart="FIGURES 131138" captionStartId="36.[151,264,1837,1861]" captionTargetBox="[151,1423,237,1794]" captionTargetId="figure-67@36.[151,1435,237,1814]" captionTargetPageId="36" captionText="FIGURES 131138. Anhoraeomorphus lambomakandroi (Franz), holotype. Habitus in dorsal view (131); head in anterodorsal view (132); head and prothorax in lateral view (only bristles on head are shown) (133); antenna (134); aedeagus in ventral (135, 137) and lateral (136, 138) views." figureDoi="http://doi.org/10.5281/zenodo.13745967" httpUri="https://zenodo.org/record/13745967/files/figure.png" pageId="35" pageNumber="36">Fig. 132</figureCitation>
) in anterodorsal view round, broadest at eyes, HL
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, HW
<quantity id="4CB6E3A3FFD35023FB89F8F2FB67432A" box="[1085,1186,1854,1881]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="35" pageNumber="36" unit="mm" value="0.48">0.48 mm</quantity>
; temple in lateral view (
<figureCitation id="137552C3FFD35023FF2BF8AEFEC4430F" box="[159,257,1890,1916]" captionStart="FIGURES 131138" captionStartId="36.[151,264,1837,1861]" captionTargetBox="[151,1423,237,1794]" captionTargetId="figure-67@36.[151,1435,237,1814]" captionTargetPageId="36" captionText="FIGURES 131138. Anhoraeomorphus lambomakandroi (Franz), holotype. Habitus in dorsal view (131); head in anterodorsal view (132); head and prothorax in lateral view (only bristles on head are shown) (133); antenna (134); aedeagus in ventral (135, 137) and lateral (136, 138) views." figureDoi="http://doi.org/10.5281/zenodo.13745967" httpUri="https://zenodo.org/record/13745967/files/figure.png" pageId="35" pageNumber="36">Fig. 133</figureCitation>
) slightly shorter than eye; vertex and frons confluent, together weakly convex and slightly transverse; vertex posteriorly weakly rounded; each eye large and weakly convex, nearly circular, weakly projecting from head silhouette. Punctures on vertex fine, inconspicuous; setae short, dense and suberect; frons and vertex densely covered with thick bristles directed posteriorly. Antenna (
<figureCitation id="137552C3FFD35023FCA9F802FCB8439B" box="[797,893,1998,2024]" captionStart="FIGURES 131138" captionStartId="36.[151,264,1837,1861]" captionTargetBox="[151,1423,237,1794]" captionTargetId="figure-67@36.[151,1435,237,1814]" captionTargetPageId="36" captionText="FIGURES 131138. Anhoraeomorphus lambomakandroi (Franz), holotype. Habitus in dorsal view (131); head in anterodorsal view (132); head and prothorax in lateral view (only bristles on head are shown) (133); antenna (134); aedeagus in ventral (135, 137) and lateral (136, 138) views." figureDoi="http://doi.org/10.5281/zenodo.13745967" httpUri="https://zenodo.org/record/13745967/files/figure.png" pageId="35" pageNumber="36">Fig. 134</figureCitation>
) slender but short, as long as half BL, AnL
<quantity id="4CB6E3A3FFD35024FADFF802FF0044C2" lastPageId="36" lastPageNumber="37" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="35" pageNumber="36" unit="mm" value="1.25">1.25 mm</quantity>
, antennomeres 12 each elongate, 3 about as long as broad, 47 each elongate, 810 each about as long as broad, 11 slightly shorter than 9 and 10 combined, as wide as 10, nearly twice as long as broad.
</paragraph>
<caption id="DF311ECEFFD45024FF23F8E1FE2A43FE" ID-DOI="http://doi.org/10.5281/zenodo.13745967" ID-Zenodo-Dep="13745967" httpUri="https://zenodo.org/record/13745967/files/figure.png" pageId="36" pageNumber="37" startId="36.[151,264,1837,1861]" targetBox="[151,1423,237,1794]" targetPageId="36" targetType="figure">
<paragraph id="8BF14E46FFD45024FF23F8E1FE2A43FE" blockId="36.[151,1436,1837,1933]" pageId="36" pageNumber="37">
<emphasis id="B93A9254FFD45024FF23F8E1FEA84336" bold="true" box="[151,365,1837,1861]" pageId="36" pageNumber="37">FIGURES 131138.</emphasis>
<taxonomicName id="4C4E35C5FFD45024FEC7F8E1FCF14336" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[371,820,1837,1861]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="lambomakandroi">
<emphasis id="B93A9254FFD45024FEC7F8E1FD244336" box="[371,737,1837,1861]" italics="true" pageId="36" pageNumber="37">Anhoraeomorphus lambomakandroi</emphasis>
(Franz)
</taxonomicName>
, holotype. Habitus in dorsal view (
<emphasis id="B93A9254FFD45024FB20F8E1FB7F4336" bold="true" box="[1172,1210,1837,1861]" pageId="36" pageNumber="37">131</emphasis>
); head in anterodorsal view (
<emphasis id="B93A9254FFD45024FF63F89DFF38431A" bold="true" box="[215,253,1873,1897]" pageId="36" pageNumber="37">132</emphasis>
); head and prothorax in lateral view (only bristles on head are shown) (
<emphasis id="B93A9254FFD45024FC78F89DFC37431A" bold="true" box="[972,1010,1873,1897]" pageId="36" pageNumber="37">133</emphasis>
); antenna (
<emphasis id="B93A9254FFD45024FBD0F89DFB4F431A" bold="true" box="[1124,1162,1873,1897]" pageId="36" pageNumber="37">134</emphasis>
); aedeagus in ventral (
<emphasis id="B93A9254FFD45024FAC4F89DFF7843FE" bold="true" pageId="36" pageNumber="37">135, 137</emphasis>
) and lateral (
<emphasis id="B93A9254FFD45024FEF1F8B9FE5B43FE" bold="true" box="[325,414,1909,1933]" pageId="36" pageNumber="37">136, 138</emphasis>
) views.
</paragraph>
</caption>
<paragraph id="8BF14E46FFD55025FF73FF5BFEEA4532" blockId="37.[151,1437,150,1257]" pageId="37" pageNumber="38">
Pronotum bell-shaped, broadest near anterior third; PL
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, PW
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. Anterior margin strongly rounded, lateral margins rounded in anterior half, weakly sinuate in posterior half; posterior corners blunt; posterior margin nearly straight. Base with one pair of large and strongly transverse pits, sublateral carinae short but distinct. Disc with fine, inconspicuous punctures; setae dense, short and suberect, lateral surface of pronotum with dense thick bristles.
</paragraph>
<paragraph id="8BF14E46FFD55025FF73FE87FACC45DE" blockId="37.[151,1437,150,1257]" pageId="37" pageNumber="38">
Elytra together oval, broadest distinctly in front of middle; EL
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, EW
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, EI 1.50; humeral calli weakly elevated, basal elytral foveae barely discernible. Punctures small and shallow but distinct, those on anterior half separated by spaces slightly shorter than diameters of punctures; setae sparse, short and suberect.
</paragraph>
<paragraph id="8BF14E46FFD55025FF73FE7BFB164586" blockId="37.[151,1437,150,1257]" pageId="37" pageNumber="38">Legs conspicuously long and slender, unmodified except hind tibiae, which are recurved and slightly broadened in distal regions, with dorsal broadened area longitudinally impressed and with outer sharp edge.</paragraph>
<paragraph id="8BF14E46FFD55025FF73FE33FB7046DA" blockId="37.[151,1437,150,1257]" pageId="37" pageNumber="38">
Aedeagus (
<figureCitation id="137552C3FFD55025FEFDFE33FE2E466A" box="[329,491,510,537]" captionStart="FIGURES 131138" captionStartId="36.[151,264,1837,1861]" captionTargetBox="[151,1423,237,1794]" captionTargetId="figure-67@36.[151,1435,237,1814]" captionTargetPageId="36" captionText="FIGURES 131138. Anhoraeomorphus lambomakandroi (Franz), holotype. Habitus in dorsal view (131); head in anterodorsal view (132); head and prothorax in lateral view (only bristles on head are shown) (133); antenna (134); aedeagus in ventral (135, 137) and lateral (136, 138) views." figureDoi="http://doi.org/10.5281/zenodo.13745967" httpUri="https://zenodo.org/record/13745967/files/figure.png" pageId="37" pageNumber="38">Figs 135138</figureCitation>
) slender, AeL
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; in ventral view suboval, broadest near middle, distally gradually narrowing and in subapical region rapidly narrowed to subtriangular apex; in lateral view apical region of dorsal wall weakly curved dorsally; endophallus with all sclerotized structures asymmetrical, distolateral sclerites broadly separated, each with subtriangular and blunt apical region; parameres broad and their apices not projecting beyond aedeagal apex, each paramere with subapical group of 5 short setae distant from apex.
</paragraph>
<paragraph id="8BF14E46FFD55025FF73FD7FFE5B46BE" blockId="37.[151,1437,150,1257]" box="[199,414,691,717]" pageId="37" pageNumber="38">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD55025FF73FD1AFC634682" box="[199,934,726,753]" pageId="37" pageNumber="38" type="distribution">
<paragraph id="8BF14E46FFD55025FF73FD1AFC634682" blockId="37.[151,1437,150,1257]" box="[199,934,726,753]" pageId="37" pageNumber="38">
<emphasis id="B93A9254FFD55025FF73FD1AFEA44683" bold="true" box="[199,353,726,752]" pageId="37" pageNumber="38">Distribution.</emphasis>
Southern
<collectingCountry id="F3590ED6FFD55025FE60FD1BFD9E4682" box="[468,603,727,753]" name="Madagascar" pageId="37" pageNumber="38">Madagascar</collectingCountry>
(Atsimo-Andrefana Region).
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD55025FF73FD37FE5C409A" pageId="37" pageNumber="38" type="discussion">
<paragraph id="8BF14E46FFD55025FF73FD37FE5C409A" blockId="37.[151,1437,150,1257]" pageId="37" pageNumber="38">
<emphasis id="B93A9254FFD55025FF73FD37FEFE4766" bold="true" box="[199,315,763,789]" pageId="37" pageNumber="38">Remarks.</emphasis>
Males of
<taxonomicName id="4C4E35C5FFD55025FE75FD37FC9B4766" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[449,862,763,789]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="lambomakandroi">
<emphasis id="B93A9254FFD55025FE75FD37FC9B4766" box="[449,862,763,789]" italics="true" pageId="37" pageNumber="38">Anhoraeomorphus lambomakandroi</emphasis>
</taxonomicName>
can be easily distinguished from all remaining congeners, except for one, by the body shape, slender legs with metatibiae recurved and slightly broadened in distal regions, with the broadened area longitudinally impressed and laterally delimited by an outer sharp edge, and by unique structures of the aedeagus. The exception is
<taxonomicName id="4C4E35C5FFD55025FD64FCABFCEF47F2" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[720,810,871,897]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="vellyi">
<emphasis id="B93A9254FFD55025FD64FCABFCEF47F2" box="[720,810,871,897]" italics="true" pageId="37" pageNumber="38">A. vellyi</emphasis>
</taxonomicName>
known to occur in Central
<collectingCountry id="F3590ED6FFD55025FBE0FCABFB1847F2" box="[1108,1245,871,897]" name="Madagascar" pageId="37" pageNumber="38">Madagascar</collectingCountry>
, and these names may turn out to be synonyms. Externally, the
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of both species (each known from
<specimenCount id="9D4885CFFFD55025FBCBFC47FADC47D6" box="[1151,1305,907,933]" count="1" pageId="37" pageNumber="38" type="generic">one specimen</specimenCount>
) differ only in body length (
<taxonomicName id="4C4E35C5FFD55025FEFDFC63FDE347BA" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[329,550,943,969]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="lambomakandroi">
<emphasis id="B93A9254FFD55025FEFDFC63FDE347BA" box="[329,550,943,969]" italics="true" pageId="37" pageNumber="38">A. lambomakandroi</emphasis>
</taxonomicName>
<quantity id="4CB6E3A3FFD55025FD99FC63FD5647BA" box="[557,659,943,969]" metricMagnitude="-3" metricUnit="m" metricValue="2.43" pageId="37" pageNumber="38" unit="mm" value="2.43">2.43 mm</quantity>
,
<taxonomicName id="4C4E35C5FFD55025FD2AFC63FD3F47BA" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[670,762,943,969]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="vellyi">
<emphasis id="B93A9254FFD55025FD2AFC63FD3F47BA" box="[670,762,943,969]" italics="true" pageId="37" pageNumber="38">A. vellyi</emphasis>
</taxonomicName>
<quantity id="4CB6E3A3FFD55025FCB4FC62FCAD47BA" box="[768,872,942,969]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" pageId="37" pageNumber="38" unit="mm" value="1.95">1.95 mm</quantity>
), relative lengths of antennae (
<taxonomicName id="4C4E35C5FFD55025FB0BFC63FA5947BA" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[1215,1436,943,969]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="lambomakandroi">
<emphasis id="B93A9254FFD55025FB0BFC63FA5947BA" box="[1215,1436,943,969]" italics="true" pageId="37" pageNumber="38">A. lambomakandroi</emphasis>
</taxonomicName>
AnL/Bl 0.52,
<taxonomicName id="4C4E35C5FFD55025FE8FFC1FFE5E479E" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[315,411,979,1005]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="vellyi">
<emphasis id="B93A9254FFD55025FE8FFC1FFE5E479E" box="[315,411,979,1005]" italics="true" pageId="37" pageNumber="38">A. vellyi</emphasis>
</taxonomicName>
0.42), and in slightly (but rather indistinctively) different proportions of antennomeres. Unfortunately, the aedeagus of the
<typeStatus id="54F5F0E4FFD55025FDA9FC3BFDBA4062" box="[541,639,1015,1041]" pageId="37" pageNumber="38" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C4E35C5FFD55025FD10FC3BFC444062" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[676,897,1015,1041]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="lambomakandroi">
<emphasis id="B93A9254FFD55025FD10FC3BFC444062" box="[676,897,1015,1041]" italics="true" pageId="37" pageNumber="38">A. lambomakandroi</emphasis>
</taxonomicName>
is partly erected, so the shapes and arrangement of twisted and displaced endophallic structures are difficult to compare with those in the
<typeStatus id="54F5F0E4FFD55025FBDFFBD7FB084046" box="[1131,1229,1051,1077]" pageId="37" pageNumber="38" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C4E35C5FFD55025FB44FBD7FA8E4046" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[1264,1355,1051,1077]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="vellyi">
<emphasis id="B93A9254FFD55025FB44FBD7FA8E4046" box="[1264,1355,1051,1077]" italics="true" pageId="37" pageNumber="38">A. vellyi</emphasis>
</taxonomicName>
, which are in repose (
<figureCitation id="137552C3FFD55025FE8EFBF3FE1E402A" box="[314,475,1087,1113]" captionStart="FIGURES 179186" captionStartId="49.[151,264,1641,1665]" captionTargetBox="[151,1415,181,1602]" captionTargetId="figure-16@49.[151,1435,181,1617]" captionTargetPageId="49" captionText="FIGURES 179186. Anhoraeomorphus vellyi (Franz), holotype. Habitus in dorsal view (179); head in anterodorsal view (180); head and prothorax in lateral view (only bristles on head are shown) (181); antenna (182); aedeagus in ventral (183, 185) and lateral (184, 186) views." figureDoi="http://doi.org/10.5281/zenodo.13745983" httpUri="https://zenodo.org/record/13745983/files/figure.png" pageId="37" pageNumber="38">Figs 183186</figureCitation>
). Some of the endophallic structures seem identical, but comparison is inconclusive. The dorsal apical plate in the
<typeStatus id="54F5F0E4FFD55025FE45FBAFFD96400E" box="[497,595,1123,1149]" pageId="37" pageNumber="38" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C4E35C5FFD55025FDC9FBAFFC99400E" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[637,860,1123,1149]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="lambomakandroi">
<emphasis id="B93A9254FFD55025FDC9FBAFFC99400E" box="[637,860,1123,1149]" italics="true" pageId="37" pageNumber="38">A. lambomakandroi</emphasis>
</taxonomicName>
is clearly more elongate, narrower than that in
<taxonomicName id="4C4E35C5FFD55025FA30FBAFFF1140D2" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="vellyi">
<emphasis id="B93A9254FFD55025FA30FBAFFF1140D2" italics="true" pageId="37" pageNumber="38">A. vellyi</emphasis>
</taxonomicName>
, but the distortion of median lobe does not allow for comparing other external aedeagal structures. With only
<specimenCount id="9D4885CFFFD55025FF23FB67FEF840B6" box="[151,317,1195,1221]" count="2" pageId="37" pageNumber="38" type="generic">two specimens</specimenCount>
at hand, I refrain from synonymizing these species names; more male specimens must be examined to clarify this problem.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,212 @@
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<mods:title id="8B984B0E71E797466FCCA79CFF4C4E44">The curse of Horaeomorphus (almost) lifted. Revision of misplaced species from Madagascar, Comoros and Mascarenes (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<subSubSection id="C3541DCDFFD55025FF23FAF6FCF54126" box="[151,816,1338,1365]" pageId="37" pageNumber="38" type="nomenclature">
<paragraph id="8BF14E46FFD55025FF23FAF6FCF54126" blockId="37.[151,816,1338,1401]" box="[151,816,1338,1365]" pageId="37" pageNumber="38">
<heading id="D0B9F92AFFD55025FF23FAF6FCF54126" bold="true" box="[151,816,1338,1365]" fontSize="11" level="2" pageId="37" pageNumber="38" reason="4">
<emphasis id="B93A9254FFD55025FF23FAF6FCF54126" bold="true" box="[151,816,1338,1365]" pageId="37" pageNumber="38">
<taxonomicName id="4C4E35C5FFD55025FF23FAF6FD724127" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[151,695,1338,1365]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="manangotryanus" status="comb. nov.">
<emphasis id="B93A9254FFD55025FF23FAF6FD954126" bold="true" box="[151,592,1338,1365]" italics="true" pageId="37" pageNumber="38">Anhoraeomorphus manangotryanus</emphasis>
(Franz)
</taxonomicName>
,
<taxonomicNameLabel id="A2092F2FFFD55025FD71FAF7FCF54126" box="[709,816,1339,1365]" pageId="37" pageNumber="38" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD55025FF23FA93FE86410A" box="[151,323,1375,1401]" pageId="37" pageNumber="38" type="description">
<paragraph id="8BF14E46FFD55025FF23FA93FE86410A" blockId="37.[151,816,1338,1401]" box="[151,323,1375,1401]" pageId="37" pageNumber="38">
(
<figureCitation id="137552C3FFD55025FF2BFA93FEFE410A" box="[159,315,1375,1401]" captionStart="FIGURES 139146" captionStartId="38.[151,264,1606,1630]" captionTargetBox="[151,1413,181,1565]" captionTargetId="figure-28@38.[151,1435,181,1582]" captionTargetPageId="38" captionText="FIGURES 139146. Anhoraeomorphus manangotryanus (Franz), holotype. Habitus in dorsal view (139); head in anterodorsal view (140); head and prothorax in lateral view (only bristles on head are shown) (141); antenna (142); aedeagus in ventral (143, 145) and lateral (144, 146) views." figureDoi="http://doi.org/10.5281/zenodo.13745969" httpUri="https://zenodo.org/record/13745969/files/figure.png" pageId="37" pageNumber="38">Figs 139146</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD55025FF23FA6FFD7141C8" box="[151,692,1443,1467]" pageId="37" pageNumber="38" type="reference_group">
<paragraph id="8BF14E46FFD55025FF23FA6FFD7141C8" blockId="37.[151,692,1443,1467]" box="[151,692,1443,1467]" pageId="37" pageNumber="38">
<treatmentCitationGroup id="AB5E6968FFD55025FF23FA6FFD7141C8" box="[151,692,1443,1467]" pageId="37" pageNumber="38">
<treatmentCitation id="0AEF6857FFD55025FF23FA6FFD7541C8" author="Franz, H." box="[151,688,1443,1467]" page="147" pageId="37" pageNumber="38" year="1986">
<taxonomicName id="4C4E35C5FFD55025FF23FA6FFD7541C8" ID-CoL="3MNKL" authority="Franz, 1986 b: 147" authorityName="Franz" authorityPageNumber="147" authorityYear="1986" box="[151,688,1443,1467]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="manangotryanus">
<emphasis id="B93A9254FFD55025FF23FA6FFE2B41C8" box="[151,494,1443,1467]" italics="true" pageId="37" pageNumber="38">Horaeomorphus manangotryanus</emphasis>
<bibRefCitation id="EFDF33B7FFD55025FE40FA68FD7541C8" author="Franz, H." box="[500,688,1443,1467]" pageId="37" pageNumber="38" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 147</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD55025FF23FA25FCE1431F" pageId="37" pageNumber="38" type="materials_examined">
<paragraph id="8BF14E46FFD55025FF23FA25FC5F4238" blockId="37.[151,1436,1513,1972]" pageId="37" pageNumber="38">
<emphasis id="B93A9254FFD55025FF23FA25FE594270" bold="true" box="[151,412,1513,1539]" pageId="37" pageNumber="38">Type material studied.</emphasis>
<materialsCitation id="3B26441BFFD55025FE2BFA25FC5F4238" collectionCode="NHMW" collectorName="Ft. Dauphin" country="Madagascar" determinerName="H. Franz" location="Madagascar" pageId="37" pageNumber="38" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
<emphasis id="B93A9254FFD55025FE2BFA25FD7A4270" bold="true" box="[415,703,1513,1539]" pageId="37" pageNumber="38">
<typeStatus id="54F5F0E4FFD55025FE2BFA25FDC94270" box="[415,524,1513,1539]" pageId="37" pageNumber="38" type="holotype">Holotype</typeStatus>
(
<collectingCountry id="F3590ED6FFD55025FDA3FA25FD774270" box="[535,690,1513,1539]" name="Madagascar" pageId="37" pageNumber="38">Madagascar</collectingCountry>
):
</emphasis>
<specimenCount id="9D4885CFFFD55025FD77FA26FD1F4277" box="[707,730,1514,1540]" count="1" pageId="37" pageNumber="38" type="male"></specimenCount>
,four labels:“S-Madagaskar,Col / de Manangotry b./
<collectorName id="26BB2B90FFD55025FAA9FA25FA594270" box="[1309,1436,1513,1539]" pageId="37" pageNumber="38">Ft.Dauphin</collectorName>
/ lg.Franz” [white, printed], “
<taxonomicName id="4C4E35C5FFD55025FE58F9C1FD664254" box="[492,675,1549,1575]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/ manangotryanus m. / det.
<determinerName id="7E2DA282FFD55025FC5CF9C1FB834254" box="[1000,1094,1549,1575]" pageId="37" pageNumber="38">H.Franz</determinerName>
” [yellowish, handwritten and printed], “Typus” [red, handwritten], “
<specimenCount id="9D4885CFFFD55025FDF2F9FDFD994238" box="[582,604,1585,1611]" count="1" pageId="37" pageNumber="38" type="male"></specimenCount>
” [white, printed] (
<collectionCode id="ED5FD683FFD55025FC98F9FDFC4A4238" box="[812,911,1585,1611]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="37" pageNumber="38" type="Museum">NHMW</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BF14E46FFD55025FF73F999FCE1431F" blockId="37.[151,1436,1513,1972]" pageId="37" pageNumber="38">
<emphasis id="B93A9254FFD55025FF73F999FE63421C" bold="true" box="[199,422,1621,1647]" pageId="37" pageNumber="38">Revised diagnosis.</emphasis>
Body (
<figureCitation id="137552C3FFD55025FDB5F999FDA0421C" box="[513,613,1621,1647]" captionStart="FIGURES 139146" captionStartId="38.[151,264,1606,1630]" captionTargetBox="[151,1413,181,1565]" captionTargetId="figure-28@38.[151,1435,181,1582]" captionTargetPageId="38" captionText="FIGURES 139146. Anhoraeomorphus manangotryanus (Franz), holotype. Habitus in dorsal view (139); head in anterodorsal view (140); head and prothorax in lateral view (only bristles on head are shown) (141); antenna (142); aedeagus in ventral (143, 145) and lateral (144, 146) views." figureDoi="http://doi.org/10.5281/zenodo.13745969" httpUri="https://zenodo.org/record/13745969/files/figure.png" pageId="37" pageNumber="38">Fig. 139</figureCitation>
) slender; head (
<figureCitation id="137552C3FFD55025FC91F999FC00421C" box="[805,965,1621,1647]" captionStart="FIGURES 139146" captionStartId="38.[151,264,1606,1630]" captionTargetBox="[151,1413,181,1565]" captionTargetId="figure-28@38.[151,1435,181,1582]" captionTargetPageId="38" captionText="FIGURES 139146. Anhoraeomorphus manangotryanus (Franz), holotype. Habitus in dorsal view (139); head in anterodorsal view (140); head and prothorax in lateral view (only bristles on head are shown) (141); antenna (142); aedeagus in ventral (143, 145) and lateral (144, 146) views." figureDoi="http://doi.org/10.5281/zenodo.13745969" httpUri="https://zenodo.org/record/13745969/files/figure.png" pageId="37" pageNumber="38">Figs 140141</figureCitation>
) round, with vertex posteriorly strongly rounded; pronotum (
<figureCitation id="137552C3FFD55025FE36F9B5FE2142E0" box="[386,484,1657,1683]" captionStart="FIGURES 139146" captionStartId="38.[151,264,1606,1630]" captionTargetBox="[151,1413,181,1565]" captionTargetId="figure-28@38.[151,1435,181,1582]" captionTargetPageId="38" captionText="FIGURES 139146. Anhoraeomorphus manangotryanus (Franz), holotype. Habitus in dorsal view (139); head in anterodorsal view (140); head and prothorax in lateral view (only bristles on head are shown) (141); antenna (142); aedeagus in ventral (143, 145) and lateral (144, 146) views." figureDoi="http://doi.org/10.5281/zenodo.13745969" httpUri="https://zenodo.org/record/13745969/files/figure.png" pageId="37" pageNumber="38">Fig. 139</figureCitation>
) subtrapezoidal, broadest near middle and anteriorly narrowing more strongly than posteriorly, with one pair of large, transverse antebasal pits; antennomeres 35 (
<figureCitation id="137552C3FFD55025FB99F951FB4A42C4" box="[1069,1167,1693,1719]" captionStart="FIGURES 139146" captionStartId="38.[151,264,1606,1630]" captionTargetBox="[151,1413,181,1565]" captionTargetId="figure-28@38.[151,1435,181,1582]" captionTargetPageId="38" captionText="FIGURES 139146. Anhoraeomorphus manangotryanus (Franz), holotype. Habitus in dorsal view (139); head in anterodorsal view (140); head and prothorax in lateral view (only bristles on head are shown) (141); antenna (142); aedeagus in ventral (143, 145) and lateral (144, 146) views." figureDoi="http://doi.org/10.5281/zenodo.13745969" httpUri="https://zenodo.org/record/13745969/files/figure.png" pageId="37" pageNumber="38">Fig. 142</figureCitation>
) each weakly elongate, 610 each about as long as broad; all femora similarly slender; hind tibiae in male unmodified; aedeagus (
<figureCitation id="137552C3FFD55025FA98F90DFF07428C" captionStart="FIGURES 139146" captionStartId="38.[151,264,1606,1630]" captionTargetBox="[151,1413,181,1565]" captionTargetId="figure-28@38.[151,1435,181,1582]" captionTargetPageId="38" captionText="FIGURES 139146. Anhoraeomorphus manangotryanus (Franz), holotype. Habitus in dorsal view (139); head in anterodorsal view (140); head and prothorax in lateral view (only bristles on head are shown) (141); antenna (142); aedeagus in ventral (143, 145) and lateral (144, 146) views." figureDoi="http://doi.org/10.5281/zenodo.13745969" httpUri="https://zenodo.org/record/13745969/files/figure.png" pageId="37" pageNumber="38">Figs 143 146</figureCitation>
) in ventral view broadest near basal third, apical region of dorsal wall in lateral view weakly curved dorsally, distolateral endophallic sclerites narrowly separated, each elongate and asymmetrical, right sclerite in ventral view with apical region bent outwards at right angle and with blunt subtriangular apex, each paramere with numerous short subapical setae distributed on distal 3/4 of paramere.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD55027FF73F8B9FE5B4682" lastPageId="39" lastPageNumber="40" pageId="37" pageNumber="38" type="description">
<paragraph id="8BF14E46FFD55025FF73F8B9FE4743C7" blockId="37.[151,1436,1513,1972]" pageId="37" pageNumber="38">
<emphasis id="B93A9254FFD55025FF73F8B9FEB043FC" bold="true" box="[199,373,1909,1935]" pageId="37" pageNumber="38">Redescription.</emphasis>
Body of male (
<figureCitation id="137552C3FFD55025FDAAF8B9FDB843FC" box="[542,637,1909,1935]" captionStart="FIGURES 139146" captionStartId="38.[151,264,1606,1630]" captionTargetBox="[151,1413,181,1565]" captionTargetId="figure-28@38.[151,1435,181,1582]" captionTargetPageId="38" captionText="FIGURES 139146. Anhoraeomorphus manangotryanus (Franz), holotype. Habitus in dorsal view (139); head in anterodorsal view (140); head and prothorax in lateral view (only bristles on head are shown) (141); antenna (142); aedeagus in ventral (143, 145) and lateral (144, 146) views." figureDoi="http://doi.org/10.5281/zenodo.13745969" httpUri="https://zenodo.org/record/13745969/files/figure.png" pageId="37" pageNumber="38">Fig. 139</figureCitation>
) strongly convex, dark brown, tarsi and palps slightly lighter; setae light brown; BL
<quantity id="4CB6E3A3FFD55025FEA3F855FEBB43C7" box="[279,382,1945,1972]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" pageId="37" pageNumber="38" unit="mm" value="3.05">3.05 mm</quantity>
.
</paragraph>
<caption id="DF311ECEFFD65026FF23F98AFE2A42D5" ID-DOI="http://doi.org/10.5281/zenodo.13745969" ID-Zenodo-Dep="13745969" httpUri="https://zenodo.org/record/13745969/files/figure.png" pageId="38" pageNumber="39" startId="38.[151,264,1606,1630]" targetBox="[151,1413,181,1565]" targetPageId="38" targetType="figure">
<paragraph id="8BF14E46FFD65026FF23F98AFE2A42D5" blockId="38.[151,1436,1606,1702]" pageId="38" pageNumber="39">
<emphasis id="B93A9254FFD65026FF23F98AFEA8422D" bold="true" box="[151,365,1606,1630]" pageId="38" pageNumber="39">FIGURES 139146.</emphasis>
<taxonomicName id="4C4E35C5FFD65026FEC7F98AFCF6422D" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[371,819,1606,1630]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="manangotryanus">
<emphasis id="B93A9254FFD65026FEC7F98AFD1A422D" box="[371,735,1606,1630]" italics="true" pageId="38" pageNumber="39">Anhoraeomorphus manangotryanus</emphasis>
(Franz)
</taxonomicName>
, holotype. Habitus in dorsal view (
<emphasis id="B93A9254FFD65026FB20F98AFB7F422D" bold="true" box="[1172,1210,1606,1630]" pageId="38" pageNumber="39">139</emphasis>
); head in anterodorsal view (
<emphasis id="B93A9254FFD65026FF63F9A6FF3842F1" bold="true" box="[215,253,1642,1666]" pageId="38" pageNumber="39">140</emphasis>
); head and prothorax in lateral view (only bristles on head are shown) (
<emphasis id="B93A9254FFD65026FC78F9A6FC3742F1" bold="true" box="[972,1010,1642,1666]" pageId="38" pageNumber="39">141</emphasis>
); antenna (
<emphasis id="B93A9254FFD65026FBD0F9A6FB4F42F1" bold="true" box="[1124,1162,1642,1666]" pageId="38" pageNumber="39">142</emphasis>
); aedeagus in ventral (
<emphasis id="B93A9254FFD65026FAC4F9A6FF7842D5" bold="true" pageId="38" pageNumber="39">143, 145</emphasis>
) and lateral (
<emphasis id="B93A9254FFD65026FEF1F942FE5B42D5" bold="true" box="[325,414,1678,1702]" pageId="38" pageNumber="39">144, 146</emphasis>
) views.
</paragraph>
</caption>
<paragraph id="8BF14E46FFD65026FF73F934FCC14399" blockId="38.[151,1437,1784,2027]" pageId="38" pageNumber="39">
Head (
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) in anterodorsal view round, broadest at eyes, HL
<quantity id="4CB6E3A3FFD65026FC18F934FBD74360" box="[940,1042,1784,1811]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="38" pageNumber="39" unit="mm" value="0.53">0.53 mm</quantity>
, HW
<quantity id="4CB6E3A3FFD65026FBE0F934FB7E4360" box="[1108,1211,1784,1811]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="38" pageNumber="39" unit="mm" value="0.53">0.53 mm</quantity>
; length of temple in lateral view (
<figureCitation id="137552C3FFD65026FE99F8D0FE4B4344" box="[301,398,1820,1847]" captionStart="FIGURES 139146" captionStartId="38.[151,264,1606,1630]" captionTargetBox="[151,1413,181,1565]" captionTargetId="figure-28@38.[151,1435,181,1582]" captionTargetPageId="38" captionText="FIGURES 139146. Anhoraeomorphus manangotryanus (Franz), holotype. Habitus in dorsal view (139); head in anterodorsal view (140); head and prothorax in lateral view (only bristles on head are shown) (141); antenna (142); aedeagus in ventral (143, 145) and lateral (144, 146) views." figureDoi="http://doi.org/10.5281/zenodo.13745969" httpUri="https://zenodo.org/record/13745969/files/figure.png" pageId="38" pageNumber="39">Fig. 141</figureCitation>
) subequal to eye; vertex and frons confluent, together weakly convex and about as long as broad; vertex posteriorly strongly rounded; each eye large and weakly convex, nearly circular, weakly projecting from head silhouette. Punctures on vertex fine, inconspicuous; setae short, dense and suberect; frons and vertex densely covered with thick bristles directed posteriorly. Antenna (
<figureCitation id="137552C3FFD65026FC37F844FC2143D0" box="[899,996,1928,1955]" captionStart="FIGURES 139146" captionStartId="38.[151,264,1606,1630]" captionTargetBox="[151,1413,181,1565]" captionTargetId="figure-28@38.[151,1435,181,1582]" captionTargetPageId="38" captionText="FIGURES 139146. Anhoraeomorphus manangotryanus (Franz), holotype. Habitus in dorsal view (139); head in anterodorsal view (140); head and prothorax in lateral view (only bristles on head are shown) (141); antenna (142); aedeagus in ventral (143, 145) and lateral (144, 146) views." figureDoi="http://doi.org/10.5281/zenodo.13745969" httpUri="https://zenodo.org/record/13745969/files/figure.png" pageId="38" pageNumber="39">Fig. 142</figureCitation>
) slender but short, as long as half BL, AnL
<quantity id="4CB6E3A3FFD65026FF65F860FEF243B4" box="[209,311,1964,1991]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="38" pageNumber="39" unit="mm" value="1.6">1.60 mm</quantity>
, antennomeres 15 each weakly elongate, 610 each about as long as broad, 11 much shorter than 9 and 10 combined, as wide as 10, only slightly elongate.
</paragraph>
<paragraph id="8BF14E46FFD75027FF73FF5BFC944532" blockId="39.[151,1437,150,969]" pageId="39" pageNumber="40">
Pronotum subtrapezoidal, broadest near middle, strongly narrowing anteriorly and weakly posteriorly; PL
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, PW
<quantity id="4CB6E3A3FFD75027FEB5FF77FEAD44A6" box="[257,360,187,213]" metricMagnitude="-4" metricUnit="m" metricValue="6.3" pageId="39" pageNumber="40" unit="mm" value="0.63">0.63 mm</quantity>
. Anterior margin weakly rounded, lateral margins weakly rounded in anterior half, slightly more so in posterior half; posterior corners blunt; posterior margin nearly straight. Base with one pair of large and strongly transverse pits, sublateral carinae short but distinct. Disc with fine, inconspicuous punctures; setae dense, short and suberect, lateral surface of pronotum with dense thick bristles.
</paragraph>
<paragraph id="8BF14E46FFD75027FF73FE87FEAE45DE" blockId="39.[151,1437,150,969]" pageId="39" pageNumber="40">
Elytra together oval, broadest distinctly in front of middle; EL
<quantity id="4CB6E3A3FFD75027FC39FE87FC314516" box="[909,1012,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.78" pageId="39" pageNumber="40" unit="mm" value="1.78">1.78 mm</quantity>
, EW
<quantity id="4CB6E3A3FFD75027FB87FE87FB5F4516" box="[1075,1178,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="39" pageNumber="40" unit="mm" value="1.2">1.20 mm</quantity>
, EI 1.48; humeral calli weakly elevated, basal elytral foveae barely discernible. Punctures small and shallow, inconspicuous; setae sparse, short and suberect.
</paragraph>
<paragraph id="8BF14E46FFD75027FF73FE7BFBA045A2" blockId="39.[151,1437,150,969]" box="[199,1125,439,465]" pageId="39" pageNumber="40">Legs long and slender, unmodified except slightly recurved fore and middle tibiae.</paragraph>
<paragraph id="8BF14E46FFD75027FF73FE17FEC346BE" blockId="39.[151,1437,150,969]" pageId="39" pageNumber="40">
Aedeagus (
<figureCitation id="137552C3FFD75027FEF3FE17FE214586" box="[327,484,475,501]" captionStart="FIGURES 139146" captionStartId="38.[151,264,1606,1630]" captionTargetBox="[151,1413,181,1565]" captionTargetId="figure-28@38.[151,1435,181,1582]" captionTargetPageId="38" captionText="FIGURES 139146. Anhoraeomorphus manangotryanus (Franz), holotype. Habitus in dorsal view (139); head in anterodorsal view (140); head and prothorax in lateral view (only bristles on head are shown) (141); antenna (142); aedeagus in ventral (143, 145) and lateral (144, 146) views." figureDoi="http://doi.org/10.5281/zenodo.13745969" httpUri="https://zenodo.org/record/13745969/files/figure.png" pageId="39" pageNumber="40">Figs 143146</figureCitation>
) slender, AeL
<quantity id="4CB6E3A3FFD75027FD3AFE17FD324586" box="[654,759,475,501]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="39" pageNumber="40" unit="mm" value="0.6">0.60 mm</quantity>
; in ventral view pear-shaped, broadest near proximal third, distally gradually narrowing and in subapical region rapidly narrowed to form subtriangular apex with broadly rounded distal margin; in lateral view apical region of dorsal wall weakly curved dorsally; endophallus with all sclerotized structures asymmetrical, distolateral sclerites narrowly separated, right sclerite in ventral view with apical region bent outwards at straight angle and with blunt subtriangular apex; parameres slender and their twisted apices slightly projecting beyond aedeagal apex, each paramere with numerous short subapical setae distributed on distal 3/4.
</paragraph>
<paragraph id="8BF14E46FFD75027FF73FD1BFE5B4682" blockId="39.[151,1437,150,969]" box="[199,414,727,753]" pageId="39" pageNumber="40">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD75027FF73FD36FCD84766" box="[199,797,762,789]" pageId="39" pageNumber="40" type="distribution">
<paragraph id="8BF14E46FFD75027FF73FD36FCD84766" blockId="39.[151,1437,150,969]" box="[199,797,762,789]" pageId="39" pageNumber="40">
<emphasis id="B93A9254FFD75027FF73FD36FEA44767" bold="true" box="[199,353,762,788]" pageId="39" pageNumber="40">Distribution.</emphasis>
Southern
<collectingCountry id="F3590ED6FFD75027FE60FD37FD9E4766" box="[468,603,763,789]" name="Madagascar" pageId="39" pageNumber="40">Madagascar</collectingCountry>
(Anosy Region).
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD75027FF73FCD3FEC047BA" pageId="39" pageNumber="40" type="discussion">
<paragraph id="8BF14E46FFD75027FF73FCD3FEC047BA" blockId="39.[151,1437,150,969]" pageId="39" pageNumber="40">
<emphasis id="B93A9254FFD75027FF73FCD3FEFE474A" bold="true" box="[199,315,799,825]" pageId="39" pageNumber="40">Remarks.</emphasis>
Males of
<taxonomicName id="4C4E35C5FFD75027FE79FCD3FCA9474B" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[461,876,799,825]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="manangotryanus">
<emphasis id="B93A9254FFD75027FE79FCD3FCA9474B" box="[461,876,799,825]" italics="true" pageId="39" pageNumber="40">Anhoraeomorphus manangotryanus</emphasis>
</taxonomicName>
belong to the largest among all species of
<taxonomicName id="4C4E35C5FFD75027FF23FC8FFEA2472E" box="[151,359,835,861]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B93A9254FFD75027FF23FC8FFEA2472E" box="[151,359,835,861]" italics="true" pageId="39" pageNumber="40">Anhoraeomorphus</emphasis>
</taxonomicName>
and exceed
<quantity id="4CB6E3A3FFD75027FE5CFC8FFDE0472E" box="[488,549,835,861]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="39" pageNumber="40" unit="mm" value="3.0">3mm</quantity>
in BL,but otherwise are relatively unremarkable,showing no tibial modifications. They can be identified by their unique aedeagal structures, including strongly asymmetrical distolateral endophallic sclerites, of which the right one in ventral view is bent outwards at a right angle in relation to the long axis of aedeagus.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,231 @@
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<subSubSection id="C3541DCDFFD75027FF23FBD6FD144046" box="[151,721,1050,1077]" pageId="39" pageNumber="40" type="nomenclature">
<paragraph id="8BF14E46FFD75027FF23FBD6FD144046" blockId="39.[151,721,1050,1113]" box="[151,721,1050,1077]" pageId="39" pageNumber="40">
<heading id="D0B9F92AFFD75027FF23FBD6FD144046" bold="true" box="[151,721,1050,1077]" fontSize="11" level="2" pageId="39" pageNumber="40" reason="4">
<emphasis id="B93A9254FFD75027FF23FBD6FD144046" bold="true" box="[151,721,1050,1077]" pageId="39" pageNumber="40">
<taxonomicName id="4C4E35C5FFD75027FF23FBD6FD9C4047" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[151,601,1050,1077]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="maximus" status="comb. nov.">
<emphasis id="B93A9254FFD75027FF23FBD6FE374046" bold="true" box="[151,498,1050,1077]" italics="true" pageId="39" pageNumber="40">Anhoraeomorphus maximus</emphasis>
(Franz)
</taxonomicName>
,
<taxonomicNameLabel id="A2092F2FFFD75027FDD2FBD7FD144046" box="[614,721,1051,1077]" pageId="39" pageNumber="40" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD75027FF23FBF3FE86402A" box="[151,323,1086,1113]" pageId="39" pageNumber="40" type="description">
<paragraph id="8BF14E46FFD75027FF23FBF3FE86402A" blockId="39.[151,721,1050,1113]" box="[151,323,1086,1113]" pageId="39" pageNumber="40">
(
<figureCitation id="137552C3FFD75027FF2BFBF3FEFE402A" box="[159,315,1086,1113]" captionStart="FIGURES 147150" captionStartId="40.[151,264,933,957]" captionTargetBox="[151,1425,181,909]" captionTargetId="figure-28@40.[151,1435,181,909]" captionTargetPageId="40" captionText="FIGURES 147150. Anhoraeomorphus maximus (Franz), holotype. Habitus in dorsal view (147); head in anterodorsal view (148); head and prothorax in lateral view (only bristles on head are shown) (149); antenna (150)." figureDoi="http://doi.org/10.5281/zenodo.13745973" httpUri="https://zenodo.org/record/13745973/files/figure.png" pageId="39" pageNumber="40">Figs 147150</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD75027FF23FB4FFDA140E8" box="[151,612,1155,1179]" pageId="39" pageNumber="40" type="reference_group">
<paragraph id="8BF14E46FFD75027FF23FB4FFDA140E8" blockId="39.[151,612,1155,1179]" box="[151,612,1155,1179]" pageId="39" pageNumber="40">
<treatmentCitationGroup id="AB5E6968FFD75027FF23FB4FFDA140E8" box="[151,612,1155,1179]" pageId="39" pageNumber="40">
<treatmentCitation id="0AEF6857FFD75027FF23FB4FFDA540E8" author="Franz, H." box="[151,608,1155,1179]" page="170" pageId="39" pageNumber="40" year="1986">
<taxonomicName id="4C4E35C5FFD75027FF23FB4FFDA540E8" ID-CoL="3MNKQ" authority="Franz, 1986 b: 170" authorityName="Franz" authorityPageNumber="170" authorityYear="1986" box="[151,608,1155,1179]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="maximus">
<emphasis id="B93A9254FFD75027FF23FB4FFE5B40E8" box="[151,414,1155,1179]" italics="true" pageId="39" pageNumber="40">Horaeomorphus maximus</emphasis>
<bibRefCitation id="EFDF33B7FFD75027FE10FB48FDA540E8" author="Franz, H." box="[420,608,1155,1179]" pageId="39" pageNumber="40" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 170</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD75027FF23FB05FD2041AC" pageId="39" pageNumber="40" type="materials_examined">
<paragraph id="8BF14E46FFD75027FF23FB05FD51413C" blockId="39.[151,1437,1225,2007]" pageId="39" pageNumber="40">
<emphasis id="B93A9254FFD75027FF23FB05FE774090" bold="true" box="[151,434,1225,1251]" pageId="39" pageNumber="40">Type material studied.</emphasis>
<materialsCitation id="3B26441BFFD75027FE74FB05FEE9413C" collectionCode="NHMW" collectorName="H. Franz" country="Madagascar" determinerName="H. Franz" pageId="39" pageNumber="40" specimenCount="3" specimenCount-female="3" typeStatus="holotype">
<emphasis id="B93A9254FFD75027FE74FB05FD2E4090" bold="true" box="[448,747,1225,1251]" pageId="39" pageNumber="40">
<typeStatus id="54F5F0E4FFD75027FE74FB05FDE84090" box="[448,557,1225,1251]" pageId="39" pageNumber="40" type="holotype">Holotype</typeStatus>
(
<collectingCountry id="F3590ED6FFD75027FDF7FB05FD1B4090" box="[579,734,1225,1251]" name="Madagascar" pageId="39" pageNumber="40">Madagascar</collectingCountry>
):
</emphasis>
<specimenCount id="9D4885CFFFD75027FD4DFB06FCCB4097" box="[761,782,1226,1252]" count="1" pageId="39" pageNumber="40" type="female"></specimenCount>
, four labels: “Massif de lAn- / karatra,Madagaskar / lg.
<collectorName id="26BB2B90FFD75027FF05FB21FECA4174" box="[177,271,1261,1287]" pageId="39" pageNumber="40">H.Franz</collectorName>
1969” [white, printed], “
<taxonomicName id="4C4E35C5FFD75027FD8DFB21FD354174" box="[569,752,1261,1287]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/ maximus m. / det.
<determinerName id="7E2DA282FFD75027FC52FB21FB814174" box="[998,1092,1261,1287]" pageId="39" pageNumber="40">H.Franz</determinerName>
” [yellowish, handwritten and printed], “Typus” [red, handwritten], “
<specimenCount id="9D4885CFFFD75027FDFFFADDFD984158" box="[587,605,1297,1323]" count="1" pageId="39" pageNumber="40" type="female"></specimenCount>
” [white, printed] (
<collectionCode id="ED5FD683FFD75027FC85FADDFC514158" box="[817,916,1297,1323]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="39" pageNumber="40" type="Museum">NHMW</collectionCode>
).
<emphasis id="B93A9254FFD75027FC13FADDFBD84158" bold="true" box="[935,1053,1297,1323]" pageId="39" pageNumber="40">
<typeStatus id="54F5F0E4FFD75027FC13FADDFBDD4158" box="[935,1048,1297,1323]" pageId="39" pageNumber="40" type="paratype">Paratype</typeStatus>
:
</emphasis>
<specimenCount id="9D4885CFFFD75027FB91FADEFBFF415F" box="[1061,1082,1298,1324]" count="1" pageId="39" pageNumber="40" type="female"></specimenCount>
, same data as for holotype, but with yellow “
</materialsCitation>
<materialsCitation id="3B26441BFFD75027FE99FAF9FD55413C" box="[301,656,1333,1359]" collectionCode="NHMW" country="Madagascar" pageId="39" pageNumber="40" specimenCount="1" typeStatus="paratype">
<typeStatus id="54F5F0E4FFD75027FE99FAF9FE14413C" box="[301,465,1333,1359]" pageId="39" pageNumber="40" type="paratype">PARATYPUS</typeStatus>
” label (
<collectionCode id="ED5FD683FFD75027FD92FAF9FD4C413C" box="[550,649,1333,1359]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="39" pageNumber="40" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF14E46FFD75027FF73FA95FD2041AC" blockId="39.[151,1437,1225,2007]" pageId="39" pageNumber="40">
<emphasis id="B93A9254FFD75027FF73FA95FE5A4100" bold="true" box="[199,415,1369,1395]" pageId="39" pageNumber="40">Revised diagnosis.</emphasis>
Body (
<figureCitation id="137552C3FFD75027FE59FA95FD8F4100" box="[493,586,1369,1395]" captionStart="FIGURES 147150" captionStartId="40.[151,264,933,957]" captionTargetBox="[151,1425,181,909]" captionTargetId="figure-28@40.[151,1435,181,909]" captionTargetPageId="40" captionText="FIGURES 147150. Anhoraeomorphus maximus (Franz), holotype. Habitus in dorsal view (147); head in anterodorsal view (148); head and prothorax in lateral view (only bristles on head are shown) (149); antenna (150)." figureDoi="http://doi.org/10.5281/zenodo.13745973" httpUri="https://zenodo.org/record/13745973/files/figure.png" pageId="39" pageNumber="40">Fig. 147</figureCitation>
) slender; head (
<figureCitation id="137552C3FFD75027FD42FA95FC554107" box="[758,912,1369,1396]" captionStart="FIGURES 147150" captionStartId="40.[151,264,933,957]" captionTargetBox="[151,1425,181,909]" captionTargetId="figure-28@40.[151,1435,181,909]" captionTargetPageId="40" captionText="FIGURES 147150. Anhoraeomorphus maximus (Franz), holotype. Habitus in dorsal view (147); head in anterodorsal view (148); head and prothorax in lateral view (only bristles on head are shown) (149); antenna (150)." figureDoi="http://doi.org/10.5281/zenodo.13745973" httpUri="https://zenodo.org/record/13745973/files/figure.png" pageId="39" pageNumber="40">Figs 148150</figureCitation>
) oval, with vertex posteriorly strongly rounded; pronotum (
<figureCitation id="137552C3FFD75027FEA2FAB1FEBD41E4" box="[278,376,1405,1431]" captionStart="FIGURES 147150" captionStartId="40.[151,264,933,957]" captionTargetBox="[151,1425,181,909]" captionTargetId="figure-28@40.[151,1435,181,909]" captionTargetPageId="40" captionText="FIGURES 147150. Anhoraeomorphus maximus (Franz), holotype. Habitus in dorsal view (147); head in anterodorsal view (148); head and prothorax in lateral view (only bristles on head are shown) (149); antenna (150)." figureDoi="http://doi.org/10.5281/zenodo.13745973" httpUri="https://zenodo.org/record/13745973/files/figure.png" pageId="39" pageNumber="40">Fig. 147</figureCitation>
) bell-shaped, broadest near anterior third, with two pairs of large subcircular antebasal pits; antennomeres 16 (
<figureCitation id="137552C3FFD75027FEC7FA6DFE1741CF" box="[371,466,1441,1468]" captionStart="FIGURES 147150" captionStartId="40.[151,264,933,957]" captionTargetBox="[151,1425,181,909]" captionTargetId="figure-28@40.[151,1435,181,909]" captionTargetPageId="40" captionText="FIGURES 147150. Anhoraeomorphus maximus (Franz), holotype. Habitus in dorsal view (147); head in anterodorsal view (148); head and prothorax in lateral view (only bristles on head are shown) (149); antenna (150)." figureDoi="http://doi.org/10.5281/zenodo.13745973" httpUri="https://zenodo.org/record/13745973/files/figure.png" pageId="39" pageNumber="40">Fig. 150</figureCitation>
) each elongate, 78 each about as long as broad, 9 and 10 each indistinctly transverse; profemora much thicker than meso-and metafemora.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD75028FF73FA25FE4040AD" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40" type="description">
<paragraph id="8BF14E46FFD75027FF73FA25FD2B425B" blockId="39.[151,1437,1225,2007]" pageId="39" pageNumber="40">
<emphasis id="B93A9254FFD75027FF73FA25FEB04270" bold="true" box="[199,373,1513,1539]" pageId="39" pageNumber="40">Redescription.</emphasis>
Body of female (
<figureCitation id="137552C3FFD75027FD82FA25FD504270" box="[566,661,1513,1539]" captionStart="FIGURES 147150" captionStartId="40.[151,264,933,957]" captionTargetBox="[151,1425,181,909]" captionTargetId="figure-28@40.[151,1435,181,909]" captionTargetPageId="40" captionText="FIGURES 147150. Anhoraeomorphus maximus (Franz), holotype. Habitus in dorsal view (147); head in anterodorsal view (148); head and prothorax in lateral view (only bristles on head are shown) (149); antenna (150)." figureDoi="http://doi.org/10.5281/zenodo.13745973" httpUri="https://zenodo.org/record/13745973/files/figure.png" pageId="39" pageNumber="40">Fig. 147</figureCitation>
) strongly convex, dark brown, elytra with reddish hue, tarsi and palps slightly lighter; setae light brown; BL
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.
</paragraph>
<paragraph id="8BF14E46FFD75027FF73F9FDFA544350" blockId="39.[151,1437,1225,2007]" pageId="39" pageNumber="40">
Head (
<figureCitation id="137552C3FFD75027FEA5F9FDFEB54238" box="[273,368,1585,1611]" captionStart="FIGURES 147150" captionStartId="40.[151,264,933,957]" captionTargetBox="[151,1425,181,909]" captionTargetId="figure-28@40.[151,1435,181,909]" captionTargetPageId="40" captionText="FIGURES 147150. Anhoraeomorphus maximus (Franz), holotype. Habitus in dorsal view (147); head in anterodorsal view (148); head and prothorax in lateral view (only bristles on head are shown) (149); antenna (150)." figureDoi="http://doi.org/10.5281/zenodo.13745973" httpUri="https://zenodo.org/record/13745973/files/figure.png" pageId="39" pageNumber="40">Fig. 148</figureCitation>
) in anterodorsal view oval, broadest at eyes, HL
<quantity id="4CB6E3A3FFD75027FC25F9FDFBF3423F" box="[913,1078,1585,1612]" metricMagnitude="-4" metricUnit="m" metricValue="6.15" metricValueMax="6.3" metricValueMin="6.0" pageId="39" pageNumber="40" unit="mm" value="0.615" valueMax="0.63" valueMin="0.6">0.600.63 mm</quantity>
, HW
<quantity id="4CB6E3A3FFD75027FBC2F9FDFB19423F" box="[1142,1244,1585,1612]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="39" pageNumber="40" unit="mm" value="0.55">0.55 mm</quantity>
; temple in lateral view (
<figureCitation id="137552C3FFD75027FF6AF999FEFB421C" box="[222,318,1621,1647]" captionStart="FIGURES 147150" captionStartId="40.[151,264,933,957]" captionTargetBox="[151,1425,181,909]" captionTargetId="figure-28@40.[151,1435,181,909]" captionTargetPageId="40" captionText="FIGURES 147150. Anhoraeomorphus maximus (Franz), holotype. Habitus in dorsal view (147); head in anterodorsal view (148); head and prothorax in lateral view (only bristles on head are shown) (149); antenna (150)." figureDoi="http://doi.org/10.5281/zenodo.13745973" httpUri="https://zenodo.org/record/13745973/files/figure.png" pageId="39" pageNumber="40">Fig. 149</figureCitation>
) slightly longer than eye; vertex and frons confluent, together weakly convex and about as long as broad; vertex posteriorly strongly rounded; each eye small and weakly convex, nearly circular, weakly projecting from head silhouette. Punctures on vertex fine, inconspicuous; setae short, dense and suberect; frons and vertex densely covered with thick bristles directed posteriorly. Antenna (
<figureCitation id="137552C3FFD75027FC32F90DFC2D42AF" box="[902,1000,1729,1756]" captionStart="FIGURES 147150" captionStartId="40.[151,264,933,957]" captionTargetBox="[151,1425,181,909]" captionTargetId="figure-28@40.[151,1435,181,909]" captionTargetPageId="40" captionText="FIGURES 147150. Anhoraeomorphus maximus (Franz), holotype. Habitus in dorsal view (147); head in anterodorsal view (148); head and prothorax in lateral view (only bristles on head are shown) (149); antenna (150)." figureDoi="http://doi.org/10.5281/zenodo.13745973" httpUri="https://zenodo.org/record/13745973/files/figure.png" pageId="39" pageNumber="40">Fig. 150</figureCitation>
) slender but short, indistinctly shorter than half BL, AnL
<quantity id="4CB6E3A3FFD75027FEDDF929FDC84373" box="[361,525,1765,1792]" metricMagnitude="-3" metricUnit="m" metricValue="1.39" metricValueMax="1.4" metricValueMin="1.38" pageId="39" pageNumber="40" unit="mm" value="1.39" valueMax="1.4" valueMin="1.38">1.381.40 mm</quantity>
, antennomeres 16 each elongate, 78 each about as long as broad, 9 and 10 each indistinctly transverse, 11 as long as 9 and 10 combined, indistinctly wider than 10, nearly twice as long as broad.
</paragraph>
<paragraph id="8BF14E46FFD75027FF73F8E2FEEA43A4" blockId="39.[151,1437,1225,2007]" pageId="39" pageNumber="40">
Pronotum bell-shaped, broadest near anterior third; PL
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, PW
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. Anterior margin strongly rounded, lateral margins strongly rounded in anterior half, sinuate in posterior half; posterior corners blunt; posterior margin nearly straight. Base with two pairs of large subcircular pits, sublateral carinae short but distinct. Disc with fine, inconspicuous punctures; setae dense, short and suberect, lateral surface of pronotum with dense thick bristles.
</paragraph>
<caption id="DF311ECEFFD85028FF23FC69FBAF4791" ID-DOI="http://doi.org/10.5281/zenodo.13745973" ID-Zenodo-Dep="13745973" httpUri="https://zenodo.org/record/13745973/files/figure.png" pageId="40" pageNumber="41" startId="40.[151,264,933,957]" targetBox="[151,1425,181,909]" targetPageId="40" targetType="figure">
<paragraph id="8BF14E46FFD85028FF23FC69FBAF4791" blockId="40.[151,1436,933,994]" pageId="40" pageNumber="41">
<emphasis id="B93A9254FFD85028FF23FC69FEAA47CE" bold="true" box="[151,367,933,957]" pageId="40" pageNumber="41">FIGURES 147150.</emphasis>
<taxonomicName id="4C4E35C5FFD85028FEC2FC6AFD2C47CD" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[374,745,934,958]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="maximus">
<emphasis id="B93A9254FFD85028FEC2FC6AFD5047CE" box="[374,661,934,958]" italics="true" pageId="40" pageNumber="41">Anhoraeomorphus maximus</emphasis>
(Franz)
</taxonomicName>
, holotype. Habitus in dorsal view (
<emphasis id="B93A9254FFD85028FBE2FC69FBB947CE" bold="true" box="[1110,1148,933,957]" pageId="40" pageNumber="41">147</emphasis>
); head in anterodorsal view (
<emphasis id="B93A9254FFD85028FF14FC05FF034792" bold="true" box="[160,198,969,993]" pageId="40" pageNumber="41">148</emphasis>
); head and prothorax in lateral view (only bristles on head are shown) (
<emphasis id="B93A9254FFD85028FC28FC05FC074792" bold="true" box="[924,962,969,993]" pageId="40" pageNumber="41">149</emphasis>
); antenna (
<emphasis id="B93A9254FFD85028FB82FC05FB994792" bold="true" box="[1078,1116,969,993]" pageId="40" pageNumber="41">150</emphasis>
).
</paragraph>
</caption>
<paragraph id="8BF14E46FFD85028FF73FBF8FC1540E5" blockId="40.[151,1436,1076,1606]" pageId="40" pageNumber="41">
Elytra together oval, conspicuously convex, broadest slightly in front of middle; EL
<quantity id="4CB6E3A3FFD85028FB31FBF8FAEE403D" box="[1157,1323,1076,1103]" metricMagnitude="-3" metricUnit="m" metricValue="1.64" metricValueMax="1.65" metricValueMin="1.63" pageId="40" pageNumber="41" unit="mm" value="1.64" valueMax="1.65" valueMin="1.63">1.631.65 mm</quantity>
, EW
<quantity id="4CB6E3A3FFD85028FADFFBF8FF004001" metricMagnitude="-3" metricUnit="m" metricValue="1.13" pageId="40" pageNumber="41" unit="mm" value="1.13">1.13 mm</quantity>
, EI 1.441.47; humeral calli weakly elevated, basal elytral foveae barely discernible. Punctures small and shallow, inconspicuous; setae sparse, conspicuously short and recumbent.
</paragraph>
<paragraph id="8BF14E46FFD85028FF73FB6CFC3340C9" blockId="40.[151,1436,1076,1606]" box="[199,1014,1184,1210]" pageId="40" pageNumber="41">Legs long and slender, unmodified except strongly thickened profemora.</paragraph>
<paragraph id="8BF14E46FFD85028FF73FB08FE4040AD" blockId="40.[151,1436,1076,1606]" box="[199,389,1220,1246]" pageId="40" pageNumber="41">Male. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD85028FF73FB24FCAE4171" box="[199,875,1256,1282]" pageId="40" pageNumber="41" type="distribution">
<paragraph id="8BF14E46FFD85028FF73FB24FCAE4171" blockId="40.[151,1436,1076,1606]" box="[199,875,1256,1282]" pageId="40" pageNumber="41">
<emphasis id="B93A9254FFD85028FF73FB24FEA44171" bold="true" box="[199,353,1256,1282]" pageId="40" pageNumber="41">Distribution.</emphasis>
Central
<collectingCountry id="F3590ED6FFD85028FE75FB24FD8D4171" box="[449,584,1256,1282]" name="Madagascar" pageId="40" pageNumber="41">Madagascar</collectingCountry>
(Vakinankaratra Region).
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD85028FF73FAC0FE464235" pageId="40" pageNumber="41" type="discussion">
<paragraph id="8BF14E46FFD85028FF73FAC0FE2341E1" blockId="40.[151,1436,1076,1606]" pageId="40" pageNumber="41">
<emphasis id="B93A9254FFD85028FF73FAC0FEFE4155" bold="true" box="[199,315,1292,1318]" pageId="40" pageNumber="41">Remarks.</emphasis>
The label data for the
<typeStatus id="54F5F0E4FFD85028FDF8FAC0FD6B4155" box="[588,686,1292,1318]" pageId="40" pageNumber="41" type="holotype">holotype</typeStatus>
and
<typeStatus id="54F5F0E4FFD85028FD5FFAC0FC8E4155" box="[747,843,1292,1318]" pageId="40" pageNumber="41" type="paratype">paratype</typeStatus>
are the same, but
<bibRefCitation id="EFDF33B7FFD85028FB9CFAC0FB0F4155" author="Franz, H." box="[1064,1226,1292,1318]" pageId="40" pageNumber="41" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986b)</bibRefCitation>
mentions that the
<typeStatus id="54F5F0E4FFD85028FF23FAFCFF3C4139" box="[151,249,1328,1354]" pageId="40" pageNumber="41" type="holotype">holotype</typeStatus>
was collected on
<date id="FFF06886FFD85028FE09FAFCFDFE4139" box="[445,571,1328,1354]" pageId="40" pageNumber="41" value="1969-12-04">12.04.1969</date>
above a forestry station (“Station forestière”), and the
<typeStatus id="54F5F0E4FFD85028FB22FAFCFB334139" box="[1174,1270,1328,1354]" pageId="40" pageNumber="41" type="paratype">paratype</typeStatus>
on
<date id="FFF06886FFD85028FAAAFAFCFA594139" box="[1310,1436,1328,1354]" pageId="40" pageNumber="41" value="1969-04-14">14.04.1969</date>
at the upper edge of the forest at Col de Manontongana, altitude nearly
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, both localities in a “Reserve forestière” of Manjakotompo.
</paragraph>
<paragraph id="8BF14E46FFD85028FF73FA50FE464235" blockId="40.[151,1436,1076,1606]" pageId="40" pageNumber="41">
Females of
<taxonomicName id="4C4E35C5FFD85028FEFAFA50FE1641C5" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[334,467,1436,1462]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="maximus">
<emphasis id="B93A9254FFD85028FEFAFA50FE1641C5" box="[334,467,1436,1462]" italics="true" pageId="40" pageNumber="41">A. maximus</emphasis>
</taxonomicName>
are the largest among all species of
<taxonomicName id="4C4E35C5FFD85028FCCDFA50FB8C41C5" box="[889,1097,1436,1462]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B93A9254FFD85028FCCDFA50FB8C41C5" box="[889,1097,1436,1462]" italics="true" pageId="40" pageNumber="41">Anhoraeomorphus</emphasis>
</taxonomicName>
, reaching ≥
<quantity id="4CB6E3A3FFD85028FB6FFA50FA8141C5" box="[1243,1348,1436,1462]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="40" pageNumber="41" unit="mm" value="3.1">3.10 mm</quantity>
. Beside the exceptional body length, they can be distinguished from all remaining species by profemora much thicker than meso- and metafemora, slender antennae with antennomere 11 as long as 9 and 10 combined, bell-shaped and elongate pronotum with two pairs of antebasal pits, and strongly convex elytra covered with conspicuously short and recumbent setae.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,262 @@
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<treatment id="03E7FF50FFD8502AFF23F946FE04409A" ID-DOI="http://doi.org/10.5281/zenodo.13746071" ID-Zenodo-Dep="13746071" LSID="urn:lsid:plazi:treatment:03E7FF50FFD8502AFF23F946FE04409A" httpUri="http://treatment.plazi.org/id/03E7FF50FFD8502AFF23F946FE04409A" lastPageId="42" lastPageNumber="43" pageId="40" pageNumber="41">
<subSubSection id="C3541DCDFFD85028FF23F946FD3742D6" box="[151,754,1674,1701]" pageId="40" pageNumber="41" type="nomenclature">
<paragraph id="8BF14E46FFD85028FF23F946FD3742D6" blockId="40.[151,754,1674,1737]" box="[151,754,1674,1701]" pageId="40" pageNumber="41">
<heading id="D0B9F92AFFD85028FF23F946FD3742D6" bold="true" box="[151,754,1674,1701]" fontSize="11" level="2" pageId="40" pageNumber="41" reason="4">
<emphasis id="B93A9254FFD85028FF23F946FD3742D6" bold="true" box="[151,754,1674,1701]" pageId="40" pageNumber="41">
<taxonomicName id="4C4E35C5FFD85028FF23F946FDBC42D7" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[151,633,1674,1701]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="moheliensis" status="comb. nov.">
<emphasis id="B93A9254FFD85028FF23F946FDD742D6" bold="true" box="[151,530,1674,1701]" italics="true" pageId="40" pageNumber="41">Anhoraeomorphus moheliensis</emphasis>
(Franz)
</taxonomicName>
,
<taxonomicNameLabel id="A2092F2FFFD85028FD33F947FD3742D6" box="[647,754,1675,1701]" pageId="40" pageNumber="41" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD85028FF23F962FE8642BA" box="[151,323,1710,1737]" pageId="40" pageNumber="41" type="description">
<paragraph id="8BF14E46FFD85028FF23F962FE8642BA" blockId="40.[151,754,1674,1737]" box="[151,323,1710,1737]" pageId="40" pageNumber="41">
(
<figureCitation id="137552C3FFD85028FF2BF962FEFF42BA" box="[159,314,1710,1737]" captionStart="FIGURES 151158" captionStartId="41.[151,264,1514,1538]" captionTargetBox="[151,1416,181,1476]" captionTargetId="figure-16@41.[151,1435,181,1489]" captionTargetPageId="41" captionText="FIGURES 151158. Anhoraeomorphus moheliensis (Franz), holotype. Habitus in dorsal view (151); head in anterodorsal view (152); head and prothorax in lateral view (only bristles on head are shown) (153); antenna (154); aedeagus in ventral (155, 157) and lateral (156, 158) views." figureDoi="http://doi.org/10.5281/zenodo.13745975" httpUri="https://zenodo.org/record/13745975/files/figure.png" pageId="40" pageNumber="41">Figs 151158</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD85028FF23F920FD474377" box="[151,642,1772,1796]" pageId="40" pageNumber="41" type="reference_group">
<paragraph id="8BF14E46FFD85028FF23F920FD474377" blockId="40.[151,642,1772,1796]" box="[151,642,1772,1796]" pageId="40" pageNumber="41">
<treatmentCitationGroup id="AB5E6968FFD85028FF23F920FD474377" box="[151,642,1772,1796]" pageId="40" pageNumber="41">
<treatmentCitation id="0AEF6857FFD85028FF23F920FDBB4377" author="Franz, H." box="[151,638,1772,1796]" page="159" pageId="40" pageNumber="41" year="1986">
<taxonomicName id="4C4E35C5FFD85028FF23F920FDBB4377" ID-CoL="3MNKT" authority="Franz, 1986 b: 159" authorityName="Franz" authorityPageNumber="159" authorityYear="1986" box="[151,638,1772,1796]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="moheliensis">
<emphasis id="B93A9254FFD85028FF23F920FE794377" box="[151,444,1772,1796]" italics="true" pageId="40" pageNumber="41">Horaeomorphus moheliensis</emphasis>
<bibRefCitation id="EFDF33B7FFD85028FE76F921FDBB4377" author="Franz, H." box="[450,638,1772,1796]" pageId="40" pageNumber="41" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 159</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD85029FF23F8E0FDFC43C0" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41" type="materials_examined">
<paragraph id="8BF14E46FFD85028FF23F8E0FE664389" blockId="40.[151,1436,1835,2042]" pageId="40" pageNumber="41">
<emphasis id="B93A9254FFD85028FF23F8E0FE614335" bold="true" box="[151,420,1836,1862]" pageId="40" pageNumber="41">Type material studied.</emphasis>
<materialsCitation id="3B26441BFFD85028FE1FF8E0FE664389" collectionCode="NHMW" collectorName="G. Gomy" country="Comoros" county="Insel" determinerName="H. Franz" elevation="420" location="Typus" municipality="Moihani" pageId="40" pageNumber="41" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Moheli" typeStatus="holotype">
<emphasis id="B93A9254FFD85028FE1FF8E0FC9A4336" bold="true" box="[427,863,1835,1862]" pageId="40" pageNumber="41">
<typeStatus id="54F5F0E4FFD85028FE1FF8E0FDDD4335" box="[427,536,1836,1862]" pageId="40" pageNumber="41" type="holotype">Holotype</typeStatus>
(
<collectingCountry id="F3590ED6FFD85028FD93F8E7FD5C4336" box="[551,665,1835,1861]" name="Comoros" pageId="40" pageNumber="41">Comoros</collectingCountry>
:
<collectingRegion id="498A80A4FFD85028FD12F8E7FC974336" box="[678,850,1835,1862]" country="Comoros" name="Moheli" pageId="40" pageNumber="41">Mohéli Island</collectingRegion>
):
</emphasis>
<specimenCount id="9D4885CFFFD85028FCD2F8E0FCB84335" box="[870,893,1836,1862]" count="1" pageId="40" pageNumber="41" type="male"></specimenCount>
, four labels: “(
<collectingCounty id="629036CAFFD85028FB97F8E0FB984335" box="[1059,1117,1836,1862]" pageId="40" pageNumber="41">Insel</collectingCounty>
<collectingRegion id="498A80A4FFD85028FBD0F8E0FB704335" box="[1124,1205,1836,1862]" country="Comoros" name="Moheli" pageId="40" pageNumber="41">Moheli</collectingRegion>
/
<collectingMunicipality id="6B95D43CFFD85028FB78F8E0FAEE4335" box="[1228,1323,1836,1862]" pageId="40" pageNumber="41">Moihani</collectingMunicipality>
/ 20.8.69,
<collectorName id="26BB2B90FFD85028FF23F89CFEC04319" box="[151,261,1872,1898]" pageId="40" pageNumber="41">G. Gomy</collectorName>
” [white, handwritten], “
<taxonomicName id="4C4E35C5FFD85028FDABF89CFD134319" box="[543,726,1872,1898]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/ moheliensis m. / det.
<determinerName id="7E2DA282FFD85028FC5CF89CFB834319" box="[1000,1094,1872,1898]" pageId="40" pageNumber="41">H.Franz</determinerName>
” [yellowish, handwritten and printed], “
<location id="8E91189DFFD85028FEBFF8B8FE9143FD" LSID="urn:lsid:plazi:treatment:03E7FF50FFD8502AFF23F946FE04409A:8E91189DFFD85028FEBFF8B8FE9143FD" box="[267,340,1908,1934]" country="Comoros" county="Insel" municipality="Moihani" name="Typus" pageId="40" pageNumber="41" stateProvince="Moheli">Typus</location>
” [red, handwritten], “
<specimenCount id="9D4885CFFFD85028FDE5F8B8FDA243FD" box="[593,615,1908,1934]" count="1" pageId="40" pageNumber="41" type="male"></specimenCount>
” [white, printed] (
<collectionCode id="ED5FD683FFD85028FC8BF8B8FC6743FD" box="[831,930,1908,1934]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="40" pageNumber="41" type="Museum">NHMW</collectionCode>
).
<emphasis id="B93A9254FFD85028FC02F8B8FBE843FD" bold="true" box="[950,1069,1908,1934]" pageId="40" pageNumber="41">
<typeStatus id="54F5F0E4FFD85028FC02F8B8FBE243FD" box="[950,1063,1908,1934]" pageId="40" pageNumber="41" type="paratype">Paratype</typeStatus>
:
</emphasis>
<specimenCount id="9D4885CFFFD85028FB82F8B8FB8E43FD" box="[1078,1099,1908,1934]" count="1" pageId="40" pageNumber="41" type="female"></specimenCount>
, “
<collectingRegion id="498A80A4FFD85028FBD2F8B8FB7C43FD" box="[1126,1209,1908,1934]" country="Comoros" name="Moheli" pageId="40" pageNumber="41">Moheli</collectingRegion>
/
<location id="8E91189DFFD85028FB61F8B8FA9643FD" LSID="urn:lsid:plazi:treatment:03E7FF50FFD8502AFF23F946FE04409A:8E91189DFFD85028FB61F8B8FA9643FD" box="[1237,1363,1908,1934]" country="Comoros" county="Insel" municipality="Moihani" name="Bandamale" pageId="40" pageNumber="41" stateProvince="Moheli">Bandamalé</location>
<quantity id="4CB6E3A3FFD85028FAE8F8B8FA5943FD" box="[1372,1436,1908,1934]" metricMagnitude="2" metricUnit="m" metricValue="4.2" pageId="40" pageNumber="41" unit="m" value="420.0">
<elevation id="0063A975FFD85028FAE8F8B8FA5943FD" box="[1372,1436,1908,1934]" metricMagnitude="2" metricUnit="m" metricValue="4.2" pageId="40" pageNumber="41" unit="m" value="420.0">420m</elevation>
</quantity>
/ IX-58 Raharizonina” [white, printed], “INSTITUT / SCIENTIFIQUE / MADAGASCAR” [bluish, printed], “
<specimenCount id="9D4885CFFFD85028FF11F870FF7243A5" box="[165,183,1980,2006]" count="1" pageId="40" pageNumber="41" type="female"></specimenCount>
”, “
<taxonomicName id="4C4E35C5FFD85028FF55F870FE5D43A5" box="[225,408,1980,2006]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/ moheliensis m. / det.
<determinerName id="7E2DA282FFD85028FD1FF870FCCD43A5" box="[683,776,1980,2006]" pageId="40" pageNumber="41">H.Franz</determinerName>
” [yellowish, handwritten and printed], “
<typeStatus id="54F5F0E4FFD85028FB54F870FA9643A5" box="[1248,1363,1980,2006]" pageId="40" pageNumber="41" type="allotype">Allotypus</typeStatus>
” [red, handwritten] (
<collectionCode id="ED5FD683FFD85028FE81F82CFE5D4389" box="[309,408,2016,2042]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="40" pageNumber="41" type="Museum">NHMW</collectionCode>
).
</materialsCitation>
</paragraph>
<caption id="DF311ECEFFD95029FF23FA26FE7F4239" ID-DOI="http://doi.org/10.5281/zenodo.13745975" ID-Zenodo-Dep="13745975" httpUri="https://zenodo.org/record/13745975/files/figure.png" pageId="41" pageNumber="42" startId="41.[151,264,1514,1538]" targetBox="[151,1416,181,1476]" targetPageId="41" targetType="figure">
<paragraph id="8BF14E46FFD95029FF23FA26FE7F4239" blockId="41.[151,1436,1514,1610]" pageId="41" pageNumber="42">
<emphasis id="B93A9254FFD95029FF23FA26FEA84271" bold="true" box="[151,365,1514,1538]" pageId="41" pageNumber="42">FIGURES 151158.</emphasis>
<taxonomicName id="4C4E35C5FFD95029FEC6FA26FD3A4271" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[370,767,1514,1538]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="moheliensis">
<emphasis id="B93A9254FFD95029FEC6FA26FD694271" box="[370,684,1514,1538]" italics="true" pageId="41" pageNumber="42">Anhoraeomorphus moheliensis</emphasis>
(Franz)
</taxonomicName>
, holotype. Habitus in dorsal view (
<emphasis id="B93A9254FFD95029FBEAFA26FB414271" bold="true" box="[1118,1156,1514,1538]" pageId="41" pageNumber="42">151</emphasis>
); head in anterodorsal view (
<emphasis id="B93A9254FFD95029FF14F9C2FF034255" bold="true" box="[160,198,1550,1574]" pageId="41" pageNumber="42">152</emphasis>
); head and prothorax in lateral view (only bristles on head are shown) (
<emphasis id="B93A9254FFD95029FC23F9C2FC784255" bold="true" box="[919,957,1550,1574]" pageId="41" pageNumber="42">153</emphasis>
); antenna (
<emphasis id="B93A9254FFD95029FB9BF9C2FB904255" bold="true" box="[1071,1109,1550,1574]" pageId="41" pageNumber="42">154</emphasis>
); aedeagus in ventral (
<emphasis id="B93A9254FFD95029FA88F9C2FA514255" bold="true" box="[1340,1428,1550,1574]" pageId="41" pageNumber="42">155, 157</emphasis>
) and lateral (
<emphasis id="B93A9254FFD95029FEA4F9FEFEAC4239" bold="true" box="[272,361,1586,1610]" pageId="41" pageNumber="42">156, 158</emphasis>
) views.
</paragraph>
</caption>
<paragraph id="8BF14E46FFD95029FF73F950FDFC43C0" blockId="41.[151,1437,1692,2043]" pageId="41" pageNumber="42">
<emphasis id="B93A9254FFD95029FF73F950FE6042C5" bold="true" box="[199,421,1692,1718]" pageId="41" pageNumber="42">Revised diagnosis.</emphasis>
Body (
<figureCitation id="137552C3FFD95029FDB5F950FDA142C4" box="[513,612,1692,1719]" captionStart="FIGURES 151158" captionStartId="41.[151,264,1514,1538]" captionTargetBox="[151,1416,181,1476]" captionTargetId="figure-16@41.[151,1435,181,1489]" captionTargetPageId="41" captionText="FIGURES 151158. Anhoraeomorphus moheliensis (Franz), holotype. Habitus in dorsal view (151); head in anterodorsal view (152); head and prothorax in lateral view (only bristles on head are shown) (153); antenna (154); aedeagus in ventral (155, 157) and lateral (156, 158) views." figureDoi="http://doi.org/10.5281/zenodo.13745975" httpUri="https://zenodo.org/record/13745975/files/figure.png" pageId="41" pageNumber="42">Fig. 151</figureCitation>
) slender; head (
<figureCitation id="137552C3FFD95029FC90F950FC0142C4" box="[804,964,1692,1719]" captionStart="FIGURES 151158" captionStartId="41.[151,264,1514,1538]" captionTargetBox="[151,1416,181,1476]" captionTargetId="figure-16@41.[151,1435,181,1489]" captionTargetPageId="41" captionText="FIGURES 151158. Anhoraeomorphus moheliensis (Franz), holotype. Habitus in dorsal view (151); head in anterodorsal view (152); head and prothorax in lateral view (only bristles on head are shown) (153); antenna (154); aedeagus in ventral (155, 157) and lateral (156, 158) views." figureDoi="http://doi.org/10.5281/zenodo.13745975" httpUri="https://zenodo.org/record/13745975/files/figure.png" pageId="41" pageNumber="42">Figs 152153</figureCitation>
) sub-pentagonal, with vertex posteriorly weakly rounded; pronotum (
<figureCitation id="137552C3FFD95029FE55F90CFD8142A8" box="[481,580,1728,1755]" captionStart="FIGURES 151158" captionStartId="41.[151,264,1514,1538]" captionTargetBox="[151,1416,181,1476]" captionTargetId="figure-16@41.[151,1435,181,1489]" captionTargetPageId="41" captionText="FIGURES 151158. Anhoraeomorphus moheliensis (Franz), holotype. Habitus in dorsal view (151); head in anterodorsal view (152); head and prothorax in lateral view (only bristles on head are shown) (153); antenna (154); aedeagus in ventral (155, 157) and lateral (156, 158) views." figureDoi="http://doi.org/10.5281/zenodo.13745975" httpUri="https://zenodo.org/record/13745975/files/figure.png" pageId="41" pageNumber="42">Fig. 151</figureCitation>
) bell-shaped, broadest near anterior third, with one pair of small circular antebasal pits; antennomeres 35 (
<figureCitation id="137552C3FFD95029FDADF928FDBC428C" box="[537,633,1764,1791]" captionStart="FIGURES 151158" captionStartId="41.[151,264,1514,1538]" captionTargetBox="[151,1416,181,1476]" captionTargetId="figure-16@41.[151,1435,181,1489]" captionTargetPageId="41" captionText="FIGURES 151158. Anhoraeomorphus moheliensis (Franz), holotype. Habitus in dorsal view (151); head in anterodorsal view (152); head and prothorax in lateral view (only bristles on head are shown) (153); antenna (154); aedeagus in ventral (155, 157) and lateral (156, 158) views." figureDoi="http://doi.org/10.5281/zenodo.13745975" httpUri="https://zenodo.org/record/13745975/files/figure.png" pageId="41" pageNumber="42">Fig. 154</figureCitation>
) each weakly elongate, 69 each about as long as broad, 10 indistinctly transverse; profemora slightly thicker than meso- and metafemora; in both sexes pro- and mesotibiae strongly and evenly curved, in males metatibiae curved in proximal third, in female metatibiae evenly curved; aedeagus (
<figureCitation id="137552C3FFD95029FADEF8E0FF3E4318" captionStart="FIGURES 151158" captionStartId="41.[151,264,1514,1538]" captionTargetBox="[151,1416,181,1476]" captionTargetId="figure-16@41.[151,1435,181,1489]" captionTargetPageId="41" captionText="FIGURES 151158. Anhoraeomorphus moheliensis (Franz), holotype. Habitus in dorsal view (151); head in anterodorsal view (152); head and prothorax in lateral view (only bristles on head are shown) (153); antenna (154); aedeagus in ventral (155, 157) and lateral (156, 158) views." figureDoi="http://doi.org/10.5281/zenodo.13745975" httpUri="https://zenodo.org/record/13745975/files/figure.png" pageId="41" pageNumber="42">Figs 155158</figureCitation>
) in ventral view broadest near middle, apical region of dorsal wall in lateral view weakly curved dorsally, distolateral endophallic sclerites broadly separated, each elongate and asymmetrical, with truncate apices, each paramere with two short apical setae.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD9502AFF73F870FA9B4046" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42" type="description">
<paragraph id="8BF14E46FFD95029FF73F870FE784388" blockId="41.[151,1437,1692,2043]" pageId="41" pageNumber="42">
<emphasis id="B93A9254FFD95029FF73F870FEB043A5" bold="true" box="[199,373,1980,2006]" pageId="41" pageNumber="42">Redescription.</emphasis>
Body of male (
<figureCitation id="137552C3FFD95029FD94F870FDBB43A5" box="[544,638,1980,2006]" captionStart="FIGURES 139146" captionStartId="38.[151,264,1606,1630]" captionTargetBox="[151,1413,181,1565]" captionTargetId="figure-28@38.[151,1435,181,1582]" captionTargetPageId="38" captionText="FIGURES 139146. Anhoraeomorphus manangotryanus (Franz), holotype. Habitus in dorsal view (139); head in anterodorsal view (140); head and prothorax in lateral view (only bristles on head are shown) (141); antenna (142); aedeagus in ventral (143, 145) and lateral (144, 146) views." figureDoi="http://doi.org/10.5281/zenodo.13745969" httpUri="https://zenodo.org/record/13745969/files/figure.png" pageId="41" pageNumber="42">Fig. 139</figureCitation>
) flattened, moderately dark brown, tarsi and palps slightly lighter; setae light brown; BL
<quantity id="4CB6E3A3FFD95029FEE6F82CFE7C4388" box="[338,441,2016,2043]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" pageId="41" pageNumber="42" unit="mm" value="2.45">2.45 mm</quantity>
.
</paragraph>
<paragraph id="8BF14E46FFDA502AFF73FF5BFD4845DE" blockId="42.[151,1437,150,1257]" pageId="42" pageNumber="43">
Head (
<figureCitation id="137552C3FFDA502AFEA0FF5BFEB344C2" box="[276,374,150,177]" captionStart="FIGURES 151158" captionStartId="41.[151,264,1514,1538]" captionTargetBox="[151,1416,181,1476]" captionTargetId="figure-16@41.[151,1435,181,1489]" captionTargetPageId="41" captionText="FIGURES 151158. Anhoraeomorphus moheliensis (Franz), holotype. Habitus in dorsal view (151); head in anterodorsal view (152); head and prothorax in lateral view (only bristles on head are shown) (153); antenna (154); aedeagus in ventral (155, 157) and lateral (156, 158) views." figureDoi="http://doi.org/10.5281/zenodo.13745975" httpUri="https://zenodo.org/record/13745975/files/figure.png" pageId="42" pageNumber="43">Fig. 152</figureCitation>
) in anterodorsal view sub-pentagonal, broadest at eyes, HL
<quantity id="4CB6E3A3FFDA502AFB9FFF5AFB5144C2" box="[1067,1172,150,177]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="42" pageNumber="43" unit="mm" value="0.53">0.53 mm</quantity>
, HW
<quantity id="4CB6E3A3FFDA502AFB6DFF5AFA8744C2" box="[1241,1346,150,177]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="42" pageNumber="43" unit="mm" value="0.45">0.45 mm</quantity>
; temple in lateral view (
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) nearly twice as long as eye; vertex and frons confluent, together weakly convex and indistinctly transverse; vertex posteriorly weakly rounded; each eye small and weakly convex, oval, not emarginate posteriorly, weakly projecting from head silhouette. Punctures on vertex fine, inconspicuous; setae short, dense and suberect; frons and vertex densely covered with thick bristles directed posteriorly. Antenna (
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) slender but short, indistinctly shorter than half BL, AnL
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, antennomeres 12 each distinctly elongate, 35 each weakly elongate, 69 each about as long as broad, 10 indistinctly transverse, 11 slightly shorter than 9 and 10 combined, as wide as 10, about 1.6 times as long as broad.
</paragraph>
<paragraph id="8BF14E46FFDA502AFF73FE7BFEEA4612" blockId="42.[151,1437,150,1257]" pageId="42" pageNumber="43">
Pronotum bell-shaped, broadest near anterior third; PL
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, PW
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. Anterior margin strongly rounded, lateral margins strongly rounded in anterior half and sinuate in posterior half; posterior corners blunt; posterior margin nearly straight. Base with one pair of small subcircular pits, sublateral carinae short but distinct. Disc with fine, inconspicuous punctures; setae dense, short and suberect, lateral surface of pronotum with dense thick bristles.
</paragraph>
<paragraph id="8BF14E46FFDA502AFF73FDA7FEAE46BE" blockId="42.[151,1437,150,1257]" pageId="42" pageNumber="43">
Elytra together oval, broadest distinctly in front of middle; EL
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, EW
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, EI 1.53; humeral calli weakly elevated, basal elytral foveae barely discernible. Punctures small and shallow, inconspicuous; setae sparse, short and suberect.
</paragraph>
<paragraph id="8BF14E46FFDA502AFF73FD1BFBC04766" blockId="42.[151,1437,150,1257]" pageId="42" pageNumber="43">Legs moderately long, slender, fore femora slightly thicker than meso- and metafemora, pro- and mesotibiae strongly and evenly curved, metatibiae strongly curved only in proximal third.</paragraph>
<paragraph id="8BF14E46FFDA502AFF73FCD3FCBD479E" blockId="42.[151,1437,150,1257]" pageId="42" pageNumber="43">
Aedeagus (
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) stout, AeL
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; in ventral view pear-shaped, broadest near middle, distally rapidly but weakly narrowing towards subtriangular apex with broadly rounded distal margin hidden behind broad and distally broadly rounded ventral apical plate; in lateral view apical region of dorsal wall weakly curved dorsally; endophallus with all sclerotized structures asymmetrical, distolateral sclerites broadly separated, strongly elongate and each with truncate apex; parameres in lateral view conspicuously broadened and their apices not projecting beyond aedeagal apex, each paramere with two short apical setae.
</paragraph>
<paragraph id="8BF14E46FFDA502AFF73FC3BFA9B4046" blockId="42.[151,1437,150,1257]" pageId="42" pageNumber="43">
Female. Externally similar to male, but with all tibiae, including hind ones, evenly curved. BL
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; HL
<quantity id="4CB6E3A3FFDA502AFF23FBD6FF384046" box="[151,253,1050,1077]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="42" pageNumber="43" unit="mm" value="0.5">0.50 mm</quantity>
, HW
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, AnL
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; PL
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, PW
<quantity id="4CB6E3A3FFDA502AFC96FBD6FC4C4046" box="[802,905,1050,1077]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="42" pageNumber="43" unit="mm" value="0.58">0.58 mm</quantity>
; EL
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, EW
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, EI
<quantity id="4CB6E3A3FFDA502AFB47FBD6FA9F4046" box="[1267,1370,1050,1077]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="42" pageNumber="43" unit="mm" value="1.5">1.50 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFDA502AFF73FBF2FD45402A" box="[199,640,1086,1113]" pageId="42" pageNumber="43" type="distribution">
<paragraph id="8BF14E46FFDA502AFF73FBF2FD45402A" blockId="42.[151,1437,150,1257]" box="[199,640,1086,1113]" pageId="42" pageNumber="43">
<emphasis id="B93A9254FFDA502AFF73FBF2FEA4402B" bold="true" box="[199,353,1086,1112]" pageId="42" pageNumber="43">Distribution.</emphasis>
<collectingRegion id="498A80A4FFDA502AFED3FBF3FDC3402A" box="[359,518,1087,1113]" country="Comoros" name="Moheli" pageId="42" pageNumber="43">Mohéli Island</collectingRegion>
,
<collectingCountry id="F3590ED6FFDA502AFDA6FBF3FDB9402A" box="[530,636,1087,1113]" name="Comoros" pageId="42" pageNumber="43">Comoros</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFDA502AFF73FBAFFE04409A" pageId="42" pageNumber="43" type="discussion">
<paragraph id="8BF14E46FFDA502AFF73FBAFFE04409A" blockId="42.[151,1437,150,1257]" pageId="42" pageNumber="43">
<emphasis id="B93A9254FFDA502AFF73FBAFFEFE400E" bold="true" box="[199,315,1123,1149]" pageId="42" pageNumber="43">Remarks.</emphasis>
This small-bodied and slender species is unique in having all tibiae in both sexes strongly curved, with metatibiae in males curved only in proximal third, and in female evenly curved along entire length. Besides the
<typeStatus id="54F5F0E4FFDA502AFF23FB67FF3C40B6" box="[151,249,1195,1221]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
and
<typeStatus id="54F5F0E4FFDA502AFE84FB67FE5540B6" box="[304,400,1195,1221]" pageId="42" pageNumber="43" type="paratype">paratype</typeStatus>
deposited at NHMW, according to
<bibRefCitation id="EFDF33B7FFDA502AFC94FB67FC7A40B6" author="Franz, H." box="[800,959,1195,1221]" pageId="42" pageNumber="43" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986b)</bibRefCitation>
<specimenCount id="9D4885CFFFDA502AFC73FB67FB5140B6" box="[967,1172,1195,1221]" count="1" pageId="42" pageNumber="43" type="male" typeStatus="paratype">one paratype male</specimenCount>
is in Muséum National d'Histoire Naturelle, Paris.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,198 @@
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<paragraph id="8BF14E46FFDA502AFF23FAF6FD2C4126" blockId="42.[151,745,1338,1401]" box="[151,745,1338,1365]" pageId="42" pageNumber="43">
<heading id="D0B9F92AFFDA502AFF23FAF6FD2C4126" bold="true" box="[151,745,1338,1365]" fontSize="11" level="2" pageId="42" pageNumber="43" reason="4">
<emphasis id="B93A9254FFDA502AFF23FAF6FD2C4126" bold="true" box="[151,745,1338,1365]" pageId="42" pageNumber="43">
<taxonomicName id="4C4E35C5FFDA502AFF23FAF6FDB44127" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[151,625,1338,1365]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="nudipennis" status="comb. nov.">
<emphasis id="B93A9254FFDA502AFF23FAF6FDCF4126" bold="true" box="[151,522,1338,1365]" italics="true" pageId="42" pageNumber="43">Anhoraeomorphus nudipennis</emphasis>
(Franz)
</taxonomicName>
,
<taxonomicNameLabel id="A2092F2FFFDA502AFDCBFAF7FD2C4126" box="[639,745,1339,1365]" pageId="42" pageNumber="43" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C3541DCDFFDA502AFF23FA93FE86410A" box="[151,323,1374,1401]" pageId="42" pageNumber="43" type="description">
<paragraph id="8BF14E46FFDA502AFF23FA93FE86410A" blockId="42.[151,745,1338,1401]" box="[151,323,1374,1401]" pageId="42" pageNumber="43">
(
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)
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFDA502AFF23FA6FFDBC41C8" box="[151,633,1443,1467]" pageId="42" pageNumber="43" type="reference_group">
<paragraph id="8BF14E46FFDA502AFF23FA6FFDBC41C8" blockId="42.[151,633,1443,1467]" box="[151,633,1443,1467]" pageId="42" pageNumber="43">
<treatmentCitationGroup id="AB5E6968FFDA502AFF23FA6FFDBC41C8" box="[151,633,1443,1467]" pageId="42" pageNumber="43">
<treatmentCitation id="0AEF6857FFDA502AFF23FA6FFDB041C8" author="Franz, H." box="[151,629,1443,1467]" page="148" pageId="42" pageNumber="43" year="1986">
<taxonomicName id="4C4E35C5FFDA502AFF23FA6FFDB041C8" ID-CoL="3MNKZ" authority="Franz, 1986 b: 148" authorityName="Franz" authorityPageNumber="148" authorityYear="1986" box="[151,629,1443,1467]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="nudipennis">
<emphasis id="B93A9254FFDA502AFF23FA6FFE7641C8" box="[151,435,1443,1467]" italics="true" pageId="42" pageNumber="43">Horaeomorphus nudipennis</emphasis>
<bibRefCitation id="EFDF33B7FFDA502AFE0DFA68FDB041C8" author="Franz, H." box="[441,629,1443,1467]" pageId="42" pageNumber="43" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 148</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFDA502AFF23FA25FDC0428C" pageId="42" pageNumber="43" type="materials_examined">
<paragraph id="8BF14E46FFDA502AFF23FA25FE1B4238" blockId="42.[151,1436,1513,2008]" pageId="42" pageNumber="43">
<emphasis id="B93A9254FFDA502AFF23FA25FE614270" bold="true" box="[151,420,1513,1539]" pageId="42" pageNumber="43">Type material studied.</emphasis>
<materialsCitation id="3B26441BFFDA502AFE1FFA25FE1B4238" collectionCode="NHMW" collectorName="H. Franz" country="Madagascar" determinerName="H. Franz" location="Madagascar" pageId="42" pageNumber="43" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="B93A9254FFDA502AFE1FFA25FD0A4270" bold="true" box="[427,719,1513,1539]" pageId="42" pageNumber="43">
<typeStatus id="54F5F0E4FFDA502AFE1FFA25FDDD4270" box="[427,536,1513,1539]" pageId="42" pageNumber="43" type="holotype">Holotype</typeStatus>
(
<collectingCountry id="F3590ED6FFDA502AFD93FA25FD074270" box="[551,706,1513,1539]" name="Madagascar" pageId="42" pageNumber="43">Madagascar</collectingCountry>
):
</emphasis>
<specimenCount id="9D4885CFFFDA502AFD62FA26FD2E4277" box="[726,747,1514,1540]" count="1" pageId="42" pageNumber="43" type="female"></specimenCount>
, three labels: “Madagaskar / Umg.Périnet / lg.
<collectorName id="26BB2B90FFDA502AFB46FA25FA954270" box="[1266,1360,1513,1539]" pageId="42" pageNumber="43">H.Franz</collectorName>
1969” [white, printed], “
<taxonomicName id="4C4E35C5FFDA502AFED7F9C1FDDF4254" box="[355,538,1549,1575]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/ nudipennis m. / det.
<determinerName id="7E2DA282FFDA502AFCAFF9C1FCBC4254" box="[795,889,1549,1575]" pageId="42" pageNumber="43">H.Franz</determinerName>
” [yellowish, handwritten and printed], “Typus” [red, handwritten] (
<collectionCode id="ED5FD683FFDA502AFEC4F9FDFE164238" box="[368,467,1585,1611]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="42" pageNumber="43" type="Museum">NHMW</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BF14E46FFDA502AFF73F999FDC0428C" blockId="42.[151,1436,1513,2008]" pageId="42" pageNumber="43">
<emphasis id="B93A9254FFDA502AFF73F999FE62421C" bold="true" box="[199,423,1621,1647]" pageId="42" pageNumber="43">Revised diagnosis.</emphasis>
Body (
<figureCitation id="137552C3FFDA502AFDB2F999FDA94203" box="[518,620,1621,1648]" captionStart="FIGURES 159162" captionStartId="43.[151,264,1115,1139]" captionTargetBox="[151,1412,309,1092]" captionTargetId="figure-76@43.[151,1435,309,1092]" captionTargetPageId="43" captionText="FIGURES 159162. Anhoraeomorphus nudipennis (Franz), holotype. Habitus in dorsal view (159); head in anterodorsal view (160); head and prothorax in lateral view (only bristles on head are shown) (161); antenna (162)." figureDoi="http://doi.org/10.5281/zenodo.13745977" httpUri="https://zenodo.org/record/13745977/files/figure.png" pageId="42" pageNumber="43">Fig. 159</figureCitation>
) moderately stout; head (
<figureCitation id="137552C3FFDA502AFC15F999FB86421C" box="[929,1091,1621,1647]" captionStart="FIGURES 159162" captionStartId="43.[151,264,1115,1139]" captionTargetBox="[151,1412,309,1092]" captionTargetId="figure-76@43.[151,1435,309,1092]" captionTargetPageId="43" captionText="FIGURES 159162. Anhoraeomorphus nudipennis (Franz), holotype. Habitus in dorsal view (159); head in anterodorsal view (160); head and prothorax in lateral view (only bristles on head are shown) (161); antenna (162)." figureDoi="http://doi.org/10.5281/zenodo.13745977" httpUri="https://zenodo.org/record/13745977/files/figure.png" pageId="42" pageNumber="43">Figs 160161</figureCitation>
) sub-pentagonal, with vertex posteriorly weakly rounded; pronotum (
<figureCitation id="137552C3FFDA502AFDEDF9B5FD7F42E7" box="[601,698,1657,1684]" captionStart="FIGURES 159162" captionStartId="43.[151,264,1115,1139]" captionTargetBox="[151,1412,309,1092]" captionTargetId="figure-76@43.[151,1435,309,1092]" captionTargetPageId="43" captionText="FIGURES 159162. Anhoraeomorphus nudipennis (Franz), holotype. Habitus in dorsal view (159); head in anterodorsal view (160); head and prothorax in lateral view (only bristles on head are shown) (161); antenna (162)." figureDoi="http://doi.org/10.5281/zenodo.13745977" httpUri="https://zenodo.org/record/13745977/files/figure.png" pageId="42" pageNumber="43">Fig. 159</figureCitation>
) bell-shaped, broadest slightly in front of anterior third, with one pair of large subcircular antebasal pits; antennomeres 16 (
<figureCitation id="137552C3FFDA502AFCF5F951FC6642C4" box="[833,931,1693,1719]" captionStart="FIGURES 159162" captionStartId="43.[151,264,1115,1139]" captionTargetBox="[151,1412,309,1092]" captionTargetId="figure-76@43.[151,1435,309,1092]" captionTargetPageId="43" captionText="FIGURES 159162. Anhoraeomorphus nudipennis (Franz), holotype. Habitus in dorsal view (159); head in anterodorsal view (160); head and prothorax in lateral view (only bristles on head are shown) (161); antenna (162)." figureDoi="http://doi.org/10.5281/zenodo.13745977" httpUri="https://zenodo.org/record/13745977/files/figure.png" pageId="42" pageNumber="43">Fig. 162</figureCitation>
) each elongate, 710 each about as long as broad; all femora similarly slender and densely setose in lateroventral distal region; metatibiae in female with flattened and matt dorsal surface
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFDA502BFF73F8C5FE4042E3" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43" type="description">
<paragraph id="8BF14E46FFDA502AFF73F8C5FF0F433B" blockId="42.[151,1436,1513,2008]" pageId="42" pageNumber="43">
<emphasis id="B93A9254FFDA502AFF73F8C5FEB04350" bold="true" box="[199,373,1801,1827]" pageId="42" pageNumber="43">Redescription.</emphasis>
Body of female (
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) moderately convex, yellowish brown; setae yellowish; BL
<quantity id="4CB6E3A3FFDA502AFADFF8C5FF03433B" metricMagnitude="-3" metricUnit="m" metricValue="2.55" pageId="42" pageNumber="43" unit="mm" value="2.55">2.55 mm</quantity>
.
</paragraph>
<paragraph id="8BF14E46FFDA502BFF73F89DFF1A456E" blockId="42.[151,1436,1513,2008]" lastBlockId="43.[151,1436,151,285]" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
Head (
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) in anterodorsal view sub-pentagonal, broadest at eyes, HL
<quantity id="4CB6E3A3FFDA502AFBA3F89DFBB8431F" box="[1047,1149,1873,1900]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="42" pageNumber="43" unit="mm" value="0.43">0.43 mm</quantity>
, HW
<quantity id="4CB6E3A3FFDA502AFB0BF89DFAE2431F" box="[1215,1319,1873,1900]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="42" pageNumber="43" unit="mm" value="0.45">0.45 mm</quantity>
; temple in lateral view (
<figureCitation id="137552C3FFDA502AFE9AF8B9FE4A43FC" box="[302,399,1909,1935]" captionStart="FIGURES 159162" captionStartId="43.[151,264,1115,1139]" captionTargetBox="[151,1412,309,1092]" captionTargetId="figure-76@43.[151,1435,309,1092]" captionTargetPageId="43" captionText="FIGURES 159162. Anhoraeomorphus nudipennis (Franz), holotype. Habitus in dorsal view (159); head in anterodorsal view (160); head and prothorax in lateral view (only bristles on head are shown) (161); antenna (162)." figureDoi="http://doi.org/10.5281/zenodo.13745977" httpUri="https://zenodo.org/record/13745977/files/figure.png" pageId="42" pageNumber="43">Fig. 161</figureCitation>
) slightly shorter than eye; vertex and frons confluent, together weakly convex and weakly transverse; vertex posteriorly weakly rounded; each eye large and weakly convex, oval and posteriorly barely noticeably emarginate, weakly projecting from head silhouette. Punctures on vertex fine, inconspicuous; setae short, dense and suberect; frons and vertex densely covered with thick bristles directed posteriorly. Antenna (
<figureCitation id="137552C3FFDB502BFA86FF5BFA5144C2" box="[1330,1428,151,177]" captionStart="FIGURES 159162" captionStartId="43.[151,264,1115,1139]" captionTargetBox="[151,1412,309,1092]" captionTargetId="figure-76@43.[151,1435,309,1092]" captionTargetPageId="43" captionText="FIGURES 159162. Anhoraeomorphus nudipennis (Franz), holotype. Habitus in dorsal view (159); head in anterodorsal view (160); head and prothorax in lateral view (only bristles on head are shown) (161); antenna (162)." figureDoi="http://doi.org/10.5281/zenodo.13745977" httpUri="https://zenodo.org/record/13745977/files/figure.png" pageId="43" pageNumber="44">Fig. 162</figureCitation>
) slender but short, indistinctly shorter than half BL, AnL
<quantity id="4CB6E3A3FFDB502BFCB6FF77FCA244A6" box="[770,871,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="1.23" pageId="43" pageNumber="44" unit="mm" value="1.23">1.23 mm</quantity>
, antennomeres 16 each elongate, 710 each about as long as broad, 11 much shorter than 9 and 10 combined, slightly narrower than 10, about 1.7 times as long as broad.
</paragraph>
<caption id="DF311ECEFFDB502BFF23FB97FBAF40EB" ID-DOI="http://doi.org/10.5281/zenodo.13745977" ID-Zenodo-Dep="13745977" httpUri="https://zenodo.org/record/13745977/files/figure.png" pageId="43" pageNumber="44" startId="43.[151,264,1115,1139]" targetBox="[151,1412,309,1092]" targetPageId="43" targetType="figure">
<paragraph id="8BF14E46FFDB502BFF23FB97FBAF40EB" blockId="43.[151,1436,1115,1176]" pageId="43" pageNumber="44">
<emphasis id="B93A9254FFDB502BFF23FB97FEAB4007" bold="true" box="[151,366,1115,1140]" pageId="43" pageNumber="44">FIGURES 159162.</emphasis>
<taxonomicName id="4C4E35C5FFDB502BFEC7FB90FD3C4007" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[371,761,1116,1140]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="nudipennis">
<emphasis id="B93A9254FFDB502BFEC7FB90FD604007" box="[371,677,1116,1140]" italics="true" pageId="43" pageNumber="44">Anhoraeomorphus nudipennis</emphasis>
(Franz)
</taxonomicName>
, holotype. Habitus in dorsal view (
<emphasis id="B93A9254FFDB502BFBE8FB90FB474007" bold="true" box="[1116,1154,1116,1140]" pageId="43" pageNumber="44">159</emphasis>
); head in anterodorsal view (
<emphasis id="B93A9254FFDB502BFF14FB4CFF0340EB" bold="true" box="[160,198,1152,1176]" pageId="43" pageNumber="44">160</emphasis>
); head and prothorax in lateral view (only bristles on head are shown) (
<emphasis id="B93A9254FFDB502BFC28FB4CFC0740EB" bold="true" box="[924,962,1152,1176]" pageId="43" pageNumber="44">161</emphasis>
); antenna (
<emphasis id="B93A9254FFDB502BFB82FB4CFB9940EB" bold="true" box="[1078,1116,1152,1176]" pageId="43" pageNumber="44">162</emphasis>
).
</paragraph>
</caption>
<paragraph id="8BF14E46FFDB502BFF73FB27FDE741E7" blockId="43.[151,1437,1258,2005]" pageId="43" pageNumber="44">
Pronotum bell-shaped, broadest slightly in front of anterior third; PL
<quantity id="4CB6E3A3FFDB502BFC66FB26FBFC4176" box="[978,1081,1258,1285]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="43" pageNumber="44" unit="mm" value="0.68">0.68 mm</quantity>
, PW
<quantity id="4CB6E3A3FFDB502BFBC2FB26FB1B4176" box="[1142,1246,1258,1285]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="43" pageNumber="44" unit="mm" value="0.6">0.60 mm</quantity>
. Anterior margin strongly rounded, lateral margins strongly rounded in anterior half, sinuate in posterior half; posterior corners blunt; posterior margin nearly straight. Base with one pair of large subcircular pits, sublateral carinae conspicuously long and sharply marked. Disc with fine, inconspicuous punctures; setae dense, short and suberect, lateral surface of pronotum with dense thick bristles.
</paragraph>
<paragraph id="8BF14E46FFDB502BFF73FA52FF354256" blockId="43.[151,1437,1258,2005]" pageId="43" pageNumber="44">
Elytra together oval, moderately convex, dorsally distinctly impressed along suture in anterior third, broadest slightly in front of middle; EL
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, EW
<quantity id="4CB6E3A3FFDB502BFD08FA0EFCE341AE" box="[700,806,1474,1501]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="43" pageNumber="44" unit="mm" value="1.05">1.05 mm</quantity>
, EI 1.38; humeral calli weakly elevated, basal elytral foveae barely discernible. Punctures small and shallow, inconspicuous; setae extremely short, so that elytra appear asetose.
</paragraph>
<paragraph id="8BF14E46FFDB502BFF73F9E3FCF5421E" blockId="43.[151,1437,1258,2005]" pageId="43" pageNumber="44">Legs long and slender, all femora densely setose in lateroventral distal region; all tibiae slightly recurved, metatibiae with dorsal surface distinctly flattened and matt.</paragraph>
<paragraph id="8BF14E46FFDB502BFF73F9BAFE4042E3" blockId="43.[151,1437,1258,2005]" box="[199,389,1654,1680]" pageId="43" pageNumber="44">Male. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFDB502BFF73F956FC4042C7" box="[199,901,1690,1716]" pageId="43" pageNumber="44" type="distribution">
<paragraph id="8BF14E46FFDB502BFF73F956FC4042C7" blockId="43.[151,1437,1258,2005]" box="[199,901,1690,1716]" pageId="43" pageNumber="44">
<emphasis id="B93A9254FFDB502BFF73F956FEA442C7" bold="true" box="[199,353,1690,1716]" pageId="43" pageNumber="44">Distribution.</emphasis>
Central
<collectingCountry id="F3590ED6FFDB502BFE75F956FD8D42C7" box="[449,584,1690,1716]" name="Madagascar" pageId="43" pageNumber="44">Madagascar</collectingCountry>
(Alaotra-Mangoro Region).
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFDB502BFF73F972FC2A43A7" pageId="43" pageNumber="44" type="discussion">
<paragraph id="8BF14E46FFDB502BFF73F972FEF44353" blockId="43.[151,1437,1258,2005]" pageId="43" pageNumber="44">
<emphasis id="B93A9254FFDB502BFF73F972FEFE42AB" bold="true" box="[199,315,1726,1752]" pageId="43" pageNumber="44">Remarks.</emphasis>
Périnet (= Analamazaotra Special Reserve, often referred to as Andasibe) was a part of the large Mantadia National Park, but currently is a separated component of Andasibe-Mantadia NP divided by deforestation.
</paragraph>
<paragraph id="8BF14E46FFDB502BFF73F8E6FC2A43A7" blockId="43.[151,1437,1258,2005]" pageId="43" pageNumber="44">
The only known female of
<taxonomicName id="4C4E35C5FFDB502BFE40F8E6FD484337" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[500,653,1834,1860]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="nudipennis">
<emphasis id="B93A9254FFDB502BFE40F8E6FD484337" box="[500,653,1834,1860]" italics="true" pageId="43" pageNumber="44">A. nudipennis</emphasis>
</taxonomicName>
has the elytra so finely setose that at magnification 80 times it appears virtually asetose. All femora have a dense group of setae in the lateroventral distal region (this feature is not as distinct as in illustration in
<bibRefCitation id="EFDF33B7FFDB502BFE67F8BFFDBD43FF" author="Franz, H." box="[467,632,1906,1933]" pageId="43" pageNumber="44" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986b)</bibRefCitation>
: fig. 130), and the metatibiae have distinctly flattened and matt dorsal surface, suggesting that in males they may be more conspicuously modified. The conspicuously short antennomere
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relation to 9 and 10 combined is another unique feature of this species.
</paragraph>
</subSubSection>
</treatment>
</document>

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<emphasis id="B93A9254FFDC502CFF23FF5BFD3844C2" bold="true" box="[151,765,151,178]" pageId="44" pageNumber="45">
<taxonomicName id="4C4E35C5FFDC502CFF23FF5BFD4044C2" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[151,645,151,178]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="roussettensis" status="comb. nov.">
<emphasis id="B93A9254FFDC502CFF23FF5BFDDB44C2" bold="true" box="[151,542,151,178]" italics="true" pageId="44" pageNumber="45">Anhoraeomorphus roussettensis</emphasis>
(Franz)
</taxonomicName>
,
<taxonomicNameLabel id="A2092F2FFFDC502CFD26FF5BFD3844C2" box="[658,765,151,177]" pageId="44" pageNumber="45" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFDC502CFF23FF77FE8644A6" box="[151,323,187,213]" pageId="44" pageNumber="45" type="description">
<paragraph id="8BF14E46FFDC502CFF23FF77FE8644A6" blockId="44.[151,765,151,213]" box="[151,323,187,213]" pageId="44" pageNumber="45">
(
<figureCitation id="137552C3FFDC502CFF2BFF77FEF944A6" box="[159,316,187,213]" captionStart="FIGURES 163170" captionStartId="45.[151,264,1727,1751]" captionTargetBox="[151,1412,181,1692]" captionTargetId="figure-16@45.[151,1435,181,1704]" captionTargetPageId="45" captionText="FIGURES 163170. Anhoraeomorphus roussettensis (Franz), holotype. Habitus in dorsal view (163); head in anterodorsal view (164); head and prothorax in lateral view (only bristles on head are shown) (165); antenna (166); aedeagus in ventral (167, 169) and lateral (168, 170) views." figureDoi="http://doi.org/10.5281/zenodo.13745979" httpUri="https://zenodo.org/record/13745979/files/figure.png" pageId="44" pageNumber="45">Figs 163170</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFDC502CFF23FF33FD494564" box="[151,652,255,279]" pageId="44" pageNumber="45" type="reference_group">
<paragraph id="8BF14E46FFDC502CFF23FF33FD494564" blockId="44.[151,652,255,279]" box="[151,652,255,279]" pageId="44" pageNumber="45">
<treatmentCitationGroup id="AB5E6968FFDC502CFF23FF33FD494564" box="[151,652,255,279]" pageId="44" pageNumber="45">
<treatmentCitation id="0AEF6857FFDC502CFF23FF33FD4D4564" author="Franz, H." box="[151,648,255,279]" page="150" pageId="44" pageNumber="45" year="1986">
<taxonomicName id="4C4E35C5FFDC502CFF23FF33FD4D4564" ID-CoL="3MNLD" authority="Franz, 1986 b: 150" authorityName="Franz" authorityPageNumber="150" authorityYear="1986" box="[151,648,255,279]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="roussettensis">
<emphasis id="B93A9254FFDC502CFF23FF33FE034564" box="[151,454,255,279]" italics="true" pageId="44" pageNumber="45">Horaeomorphus roussettensis</emphasis>
<bibRefCitation id="EFDF33B7FFDC502CFE78FECCFD4D4564" author="Franz, H." box="[460,648,255,279]" pageId="44" pageNumber="45" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 150</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFDC502CFF23FE8AFB5646BB" pageId="44" pageNumber="45" type="materials_examined">
<paragraph id="8BF14E46FFDC502CFF23FE8AFA4C45BF" blockId="44.[151,1438,325,1936]" pageId="44" pageNumber="45">
<emphasis id="B93A9254FFDC502CFF23FE8AFE62452C" bold="true" box="[151,423,325,352]" pageId="44" pageNumber="45">Type material studied.</emphasis>
<materialsCitation id="3B26441BFFDC502CFE04FE8AFB9745DB" collectingDate="1954-11" collectingDateMax="1954-12-31" collectingDateMin="1954-11" collectionCode="NHMW" collectorName="A. Robinson" country="Madagascar" determinerName="H. Franz" location="Montagne d'Ambre" municipality="Les Roussettes" pageId="44" pageNumber="45" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
<emphasis id="B93A9254FFDC502CFE04FE8AFD08452C" bold="true" box="[432,717,325,352]" pageId="44" pageNumber="45">
<typeStatus id="54F5F0E4FFDC502CFE04FE8AFDD84513" box="[432,541,326,352]" pageId="44" pageNumber="45" type="holotype">Holotype</typeStatus>
(
<collectingCountry id="F3590ED6FFDC502CFD99FE89FD01452C" box="[557,708,325,351]" name="Madagascar" pageId="44" pageNumber="45">Madagascar</collectingCountry>
)
</emphasis>
:
<specimenCount id="9D4885CFFFDC502CFD69FE8AFD314513" box="[733,756,326,352]" count="1" pageId="44" pageNumber="45" type="male"></specimenCount>
, four labels: “N-Madagaskar /
<location id="8E91189DFFDC502CFBEDFE8AFAFF4513" LSID="urn:lsid:plazi:treatment:03E7FF50FFDC502CFF23FF5BFC0743E3:8E91189DFFDC502CFBEDFE8AFAFF4513" box="[1113,1338,326,352]" country="Madagascar" municipality="Les Roussettes" name="Montagne d'Ambre" pageId="44" pageNumber="45">Montagne dAmbre</location>
” [white, printed], “
<collectingMunicipality id="6B95D43CFFDC502CFEBFFEA6FE7645F7" box="[267,435,362,388]" pageId="44" pageNumber="45">Les Roussettes</collectingMunicipality>
/
<date id="FFF06886FFDC502CFE7EFEA6FDA045F6" box="[458,613,362,389]" pageId="44" pageNumber="45" value="1954-11" valueMax="1954-12-31" valueMin="1954-11-00">
<collectingDate id="EFB4916EFFDC502CFE7EFEA6FDA045F6" box="[458,613,362,389]" pageId="44" pageNumber="45" value="1954-11" valueMax="1954-12-31" valueMin="1954-11-00">XI.-XII. 1954</collectingDate>
</date>
/ lg.
<collectorName id="26BB2B90FFDC502CFD2BFEA6FCF545F7" box="[671,816,362,388]" pageId="44" pageNumber="45">A. Robinson</collectorName>
” [white, handwritten], “
<taxonomicName id="4C4E35C5FFDC502CFB8BFEA6FB3345F7" box="[1087,1270,362,388]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/ roussettensis m. / det.
<determinerName id="7E2DA282FFDC502CFF45FE42FE8A45DB" box="[241,335,398,424]" pageId="44" pageNumber="45">H.Franz</determinerName>
” [yellowish, handwritten and printed], “
<specimenCount id="9D4885CFFFDC502CFCBBFE42FCE345DB" box="[783,806,398,424]" count="1" pageId="44" pageNumber="45" type="male"></specimenCount>
”, white, printed (
<collectionCode id="ED5FD683FFDC502CFC53FE42FB8F45DB" box="[999,1098,398,424]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="44" pageNumber="45" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B26441BFFDC502CFBEFFE42FA4045BF" collectingDate="1954-11" collectingDateMax="1954-12-31" collectingDateMin="1954-11" collectionCode="NHMW" collectorName="A. Robinson" country="Madagascar" determinerName="H. Franz" location="Montagne d'Ambre" municipality="Les Roussettes" pageId="44" pageNumber="45" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<emphasis id="B93A9254FFDC502CFBEFFE42FB1945DB" bold="true" box="[1115,1244,398,424]" pageId="44" pageNumber="45">
<typeStatus id="54F5F0E4FFDC502CFBEFFE42FB1245DB" box="[1115,1239,398,424]" pageId="44" pageNumber="45" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D4885CFFFDC502CFB56FE42FAE745DB" box="[1250,1314,398,424]" count="2" pageId="44" pageNumber="45" type="female">2 ♀♀</specimenCount>
, same data as for holotype, but one with red handwritten “
<typeStatus id="54F5F0E4FFDC502CFD16FE7EFCD245BF" box="[674,791,434,460]" pageId="44" pageNumber="45" type="allotype">Allotypus</typeStatus>
”, and one with yellow “
<typeStatus id="54F5F0E4FFDC502CFB95FE7EFB0145BF" box="[1057,1220,434,460]" pageId="44" pageNumber="45" type="paratype">PARATYPUS</typeStatus>
” label (
<collectionCode id="ED5FD683FFDC502CFAAFFE7EFABB45BF" box="[1307,1406,434,460]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="44" pageNumber="45" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF14E46FFDC502CFF73FE19FB5646BB" blockId="44.[151,1438,325,1936]" pageId="44" pageNumber="45">
<emphasis id="B93A9254FFDC502CFF73FE19FE63459C" bold="true" box="[199,422,469,495]" pageId="44" pageNumber="45">Revised diagnosis.</emphasis>
Body (
<figureCitation id="137552C3FFDC502CFDB7FE1AFDA24583" box="[515,615,470,496]" captionStart="FIGURES 163170" captionStartId="45.[151,264,1727,1751]" captionTargetBox="[151,1412,181,1692]" captionTargetId="figure-16@45.[151,1435,181,1704]" captionTargetPageId="45" captionText="FIGURES 163170. Anhoraeomorphus roussettensis (Franz), holotype. Habitus in dorsal view (163); head in anterodorsal view (164); head and prothorax in lateral view (only bristles on head are shown) (165); antenna (166); aedeagus in ventral (167, 169) and lateral (168, 170) views." figureDoi="http://doi.org/10.5281/zenodo.13745979" httpUri="https://zenodo.org/record/13745979/files/figure.png" pageId="44" pageNumber="45">Fig. 163</figureCitation>
) slender; head (
<figureCitation id="137552C3FFDC502CFC9DFE1AFC0C4582" box="[809,969,470,497]" captionStart="FIGURES 163170" captionStartId="45.[151,264,1727,1751]" captionTargetBox="[151,1412,181,1692]" captionTargetId="figure-16@45.[151,1435,181,1704]" captionTargetPageId="45" captionText="FIGURES 163170. Anhoraeomorphus roussettensis (Franz), holotype. Habitus in dorsal view (163); head in anterodorsal view (164); head and prothorax in lateral view (only bristles on head are shown) (165); antenna (166); aedeagus in ventral (167, 169) and lateral (168, 170) views." figureDoi="http://doi.org/10.5281/zenodo.13745979" httpUri="https://zenodo.org/record/13745979/files/figure.png" pageId="44" pageNumber="45">Figs 164165</figureCitation>
) round, with vertex posteriorly broadly rounded; pronotum (
<figureCitation id="137552C3FFDC502CFEC8FE36FE1F4667" box="[380,474,506,532]" captionStart="FIGURES 163170" captionStartId="45.[151,264,1727,1751]" captionTargetBox="[151,1412,181,1692]" captionTargetId="figure-16@45.[151,1435,181,1704]" captionTargetPageId="45" captionText="FIGURES 163170. Anhoraeomorphus roussettensis (Franz), holotype. Habitus in dorsal view (163); head in anterodorsal view (164); head and prothorax in lateral view (only bristles on head are shown) (165); antenna (166); aedeagus in ventral (167, 169) and lateral (168, 170) views." figureDoi="http://doi.org/10.5281/zenodo.13745979" httpUri="https://zenodo.org/record/13745979/files/figure.png" pageId="44" pageNumber="45">Fig. 163</figureCitation>
) bell-shaped, broadest in anterior third, with one pair of large transverse antebasal pits; antennomeres 37 (
<figureCitation id="137552C3FFDC502CFEC3FDD2FE1D464B" box="[375,472,542,568]" captionStart="FIGURES 163170" captionStartId="45.[151,264,1727,1751]" captionTargetBox="[151,1412,181,1692]" captionTargetId="figure-16@45.[151,1435,181,1704]" captionTargetPageId="45" captionText="FIGURES 163170. Anhoraeomorphus roussettensis (Franz), holotype. Habitus in dorsal view (163); head in anterodorsal view (164); head and prothorax in lateral view (only bristles on head are shown) (165); antenna (166); aedeagus in ventral (167, 169) and lateral (168, 170) views." figureDoi="http://doi.org/10.5281/zenodo.13745979" httpUri="https://zenodo.org/record/13745979/files/figure.png" pageId="44" pageNumber="45">Fig. 166</figureCitation>
) each about as long as broad, 810 each indistinctly transverse; all femora similarly slender; metatibiae in male modified, strongly broadening to distal third, then narrowing to apex, with tuft of setae on outer surface at widest point; aedeagus (
<figureCitation id="137552C3FFDC502CFD34FDAAFCD846F3" box="[640,797,614,640]" captionStart="FIGURES 163170" captionStartId="45.[151,264,1727,1751]" captionTargetBox="[151,1412,181,1692]" captionTargetId="figure-16@45.[151,1435,181,1704]" captionTargetPageId="45" captionText="FIGURES 163170. Anhoraeomorphus roussettensis (Franz), holotype. Habitus in dorsal view (163); head in anterodorsal view (164); head and prothorax in lateral view (only bristles on head are shown) (165); antenna (166); aedeagus in ventral (167, 169) and lateral (168, 170) views." figureDoi="http://doi.org/10.5281/zenodo.13745979" httpUri="https://zenodo.org/record/13745979/files/figure.png" pageId="44" pageNumber="45">Figs 167170</figureCitation>
) in ventral view with sub-parallel lateral margins, apical region of dorsal wall in lateral view weakly curved dorsally, distolateral endophallic sclerites widely separated, each strongly elongate and with subtriangular apex, parameres in the only studied male missing.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFDC502CFF73FD1DFA4442AE" pageId="44" pageNumber="45" type="description">
<paragraph id="8BF14E46FFDC502CFF73FD1DFEC74763" blockId="44.[151,1438,325,1936]" pageId="44" pageNumber="45">
<emphasis id="B93A9254FFDC502CFF73FD1DFEB04698" bold="true" box="[199,373,721,747]" pageId="44" pageNumber="45">Redescription.</emphasis>
Body of male (
<figureCitation id="137552C3FFDC502CFD9EFD1EFD4E469F" box="[554,651,722,748]" captionStart="FIGURES 163170" captionStartId="45.[151,264,1727,1751]" captionTargetBox="[151,1412,181,1692]" captionTargetId="figure-16@45.[151,1435,181,1704]" captionTargetPageId="45" captionText="FIGURES 163170. Anhoraeomorphus roussettensis (Franz), holotype. Habitus in dorsal view (163); head in anterodorsal view (164); head and prothorax in lateral view (only bristles on head are shown) (165); antenna (166); aedeagus in ventral (167, 169) and lateral (168, 170) views." figureDoi="http://doi.org/10.5281/zenodo.13745979" httpUri="https://zenodo.org/record/13745979/files/figure.png" pageId="44" pageNumber="45">Fig. 163</figureCitation>
) dark brown, tarsi and palps distinctly lighter, setae light brown; BL
<quantity id="4CB6E3A3FFDC502CFF23FD3AFF3B4763" box="[151,254,758,785]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" pageId="44" pageNumber="45" unit="mm" value="2.35">2.35 mm</quantity>
.
</paragraph>
<paragraph id="8BF14E46FFDC502CFF73FCD6FAC7407F" blockId="44.[151,1438,325,1936]" pageId="44" pageNumber="45">
Head (
<figureCitation id="137552C3FFDC502CFEA0FCD6FEB04747" box="[276,373,794,820]" captionStart="FIGURES 163170" captionStartId="45.[151,264,1727,1751]" captionTargetBox="[151,1412,181,1692]" captionTargetId="figure-16@45.[151,1435,181,1704]" captionTargetPageId="45" captionText="FIGURES 163170. Anhoraeomorphus roussettensis (Franz), holotype. Habitus in dorsal view (163); head in anterodorsal view (164); head and prothorax in lateral view (only bristles on head are shown) (165); antenna (166); aedeagus in ventral (167, 169) and lateral (168, 170) views." figureDoi="http://doi.org/10.5281/zenodo.13745979" httpUri="https://zenodo.org/record/13745979/files/figure.png" pageId="44" pageNumber="45">Fig. 164</figureCitation>
) in anterodorsal view round, broadest at eyes, HL
<quantity id="4CB6E3A3FFDC502CFC0AFCD6FBE24747" box="[958,1063,794,820]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="44" pageNumber="45" unit="mm" value="0.4">0.40 mm</quantity>
, HW
<quantity id="4CB6E3A3FFDC502CFBDFFCD6FB104747" box="[1131,1237,794,820]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="44" pageNumber="45" unit="mm" value="0.43">0.43 mm</quantity>
; temple in lateral view (
<figureCitation id="137552C3FFDC502CFF69FCF2FEF9472A" box="[221,316,830,857]" captionStart="FIGURES 163170" captionStartId="45.[151,264,1727,1751]" captionTargetBox="[151,1412,181,1692]" captionTargetId="figure-16@45.[151,1435,181,1704]" captionTargetPageId="45" captionText="FIGURES 163170. Anhoraeomorphus roussettensis (Franz), holotype. Habitus in dorsal view (163); head in anterodorsal view (164); head and prothorax in lateral view (only bristles on head are shown) (165); antenna (166); aedeagus in ventral (167, 169) and lateral (168, 170) views." figureDoi="http://doi.org/10.5281/zenodo.13745979" httpUri="https://zenodo.org/record/13745979/files/figure.png" pageId="44" pageNumber="45">Fig. 165</figureCitation>
) subequal to eye; vertex and frons confluent, together weakly convex and weakly transverse; vertex posteriorly broadly rounded; each eye large and weakly convex, deeply emarginate posteriorly, weakly projecting from head silhouette. Punctures on vertex fine, inconspicuous; setae short, dense and suberect; frons and vertex densely covered with thick bristles directed posteriorly. Antenna (
<figureCitation id="137552C3FFDC502CFCCAFC66FC1B47B7" box="[894,990,938,964]" captionStart="FIGURES 163170" captionStartId="45.[151,264,1727,1751]" captionTargetBox="[151,1412,181,1692]" captionTargetId="figure-16@45.[151,1435,181,1704]" captionTargetPageId="45" captionText="FIGURES 163170. Anhoraeomorphus roussettensis (Franz), holotype. Habitus in dorsal view (163); head in anterodorsal view (164); head and prothorax in lateral view (only bristles on head are shown) (165); antenna (166); aedeagus in ventral (167, 169) and lateral (168, 170) views." figureDoi="http://doi.org/10.5281/zenodo.13745979" httpUri="https://zenodo.org/record/13745979/files/figure.png" pageId="44" pageNumber="45">Fig. 166</figureCitation>
) slender but short, slightly shorter than half BL, AnL
<quantity id="4CB6E3A3FFDC502CFE82FC02FE58479B" box="[310,413,974,1001]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="44" pageNumber="45" unit="mm" value="1.05">1.05 mm</quantity>
, antennomeres 12 each elongate, 37 each about as long as broad, 810 each indistinctly transverse, 11 much shorter than 9 and 10 combined, as wide as 10, about 1.7 times as long as broad.
</paragraph>
<paragraph id="8BF14E46FFDC502CFF73FBDAFF3540B3" blockId="44.[151,1438,325,1936]" pageId="44" pageNumber="45">
Pronotum bell-shaped, broadest in anterior third; PL
<quantity id="4CB6E3A3FFDC502CFCA4FBDAFCB04043" box="[784,885,1046,1072]" metricMagnitude="-4" metricUnit="m" metricValue="6.3" pageId="44" pageNumber="45" unit="mm" value="0.63">0.63 mm</quantity>
, PW
<quantity id="4CB6E3A3FFDC502CFC04FBDAFBD34043" box="[944,1046,1046,1073]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="44" pageNumber="45" unit="mm" value="0.53">0.53 mm</quantity>
. Anterior margin strongly rounded, lateral margins rounded in anterior half, sinuate in posterior half; posterior corners blunt; posterior margin nearly straight. Base with one pair of large transversely oval pits, and with short but distinct sublateral carinae. Disc with fine, inconspicuous punctures; setae dense, short and suberect, lateral surface of pronotum with dense thick bristles.
</paragraph>
<paragraph id="8BF14E46FFDC502CFF73FB06FEAE4123" blockId="44.[151,1438,325,1936]" pageId="44" pageNumber="45">
Elytra together oval, broadest distinctly in front of middle; EL
<quantity id="4CB6E3A3FFDC502CFC39FB06FC314097" box="[909,1012,1226,1252]" metricMagnitude="-3" metricUnit="m" metricValue="1.33" pageId="44" pageNumber="45" unit="mm" value="1.33">1.33 mm</quantity>
, EW
<quantity id="4CB6E3A3FFDC502CFB87FB06FB5F4097" box="[1075,1178,1226,1252]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" pageId="44" pageNumber="45" unit="mm" value="0.88">0.88 mm</quantity>
, EI 1.51; humeral calli weakly elevated, basal elytral foveae vestigial, barely discernible. Punctures small and shallow, but more distinct than those on pronotum, those in anterior half separated by spaces subequal to diameters of punctures; setae sparse, short and suberect.
</paragraph>
<paragraph id="8BF14E46FFDC502CFF73FA96FF3E4193" blockId="44.[151,1438,325,1936]" pageId="44" pageNumber="45">Legs moderately long, slender, unmodified except large and concave patch of adhesive setae on apical mesal region of protibia and slightly recurved and strongly broadening metatibiae, which are broadest near distal third, where outer margin bears tuft of long setae, and from this point tibia narrowing distally and its outer margin concave.</paragraph>
<paragraph id="8BF14E46FFDC502CFF73FA26FBBE42E7" blockId="44.[151,1438,325,1936]" pageId="44" pageNumber="45">
Aedeagus (
<figureCitation id="137552C3FFDC502CFEFCFA26FE234277" box="[328,486,1514,1540]" captionStart="FIGURES 163170" captionStartId="45.[151,264,1727,1751]" captionTargetBox="[151,1412,181,1692]" captionTargetId="figure-16@45.[151,1435,181,1704]" captionTargetPageId="45" captionText="FIGURES 163170. Anhoraeomorphus roussettensis (Franz), holotype. Habitus in dorsal view (163); head in anterodorsal view (164); head and prothorax in lateral view (only bristles on head are shown) (165); antenna (166); aedeagus in ventral (167, 169) and lateral (168, 170) views." figureDoi="http://doi.org/10.5281/zenodo.13745979" httpUri="https://zenodo.org/record/13745979/files/figure.png" pageId="44" pageNumber="45">Figs 167170</figureCitation>
) slender, AeL
<quantity id="4CB6E3A3FFDC502CFD27FA26FD384277" box="[659,765,1514,1540]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="44" pageNumber="45" unit="mm" value="0.43">0.43 mm</quantity>
; in ventral view drop-shaped, with lateral margins largely subparallel, distally rapidly narrowing towards broadly subtriangular apex with rounded tip; in lateral view apical region of dorsal wall weakly curved dorsally; endophallus with asymmetrical and conspicuously long distolateral sclerites and with strongly asymmetrical assemblage of median proximal sclerotized structures, distolateral sclerites broadly separated, each with subtriangular apex; parameres in studied specimen missing.
</paragraph>
<paragraph id="8BF14E46FFDC502CFF73F952FA4442AE" blockId="44.[151,1438,325,1936]" pageId="44" pageNumber="45">
Female. Externally similar to male, but with unmodified metatibiae. BL
<quantity id="4CB6E3A3FFDC502CFC57F952FB8D42CB" box="[995,1096,1694,1720]" metricMagnitude="-3" metricUnit="m" metricValue="2.33" pageId="44" pageNumber="45" unit="mm" value="2.33">2.33 mm</quantity>
; HL
<quantity id="4CB6E3A3FFDC502CFBC8F952FADA42CB" box="[1148,1311,1694,1720]" metricMagnitude="-4" metricUnit="m" metricValue="4.15" metricValueMax="4.3" metricValueMin="4.0" pageId="44" pageNumber="45" unit="mm" value="0.415" valueMax="0.43" valueMin="0.4">0.400.43 mm</quantity>
, HW
<quantity id="4CB6E3A3FFDC502CFAE9F952FF3842AF" metricMagnitude="-4" metricUnit="m" metricValue="4.4" metricValueMax="4.5" metricValueMin="4.3" pageId="44" pageNumber="45" unit="mm" value="0.44" valueMax="0.45" valueMin="0.43">0.43 0.45 mm</quantity>
, AnL 0.931.00 mm; PL
<quantity id="4CB6E3A3FFDC502CFDACF90EFD7842AF" box="[536,701,1730,1756]" metricMagnitude="-4" metricUnit="m" metricValue="6.15" metricValueMax="6.3" metricValueMin="6.0" pageId="44" pageNumber="45" unit="mm" value="0.615" valueMax="0.63" valueMin="0.6">0.600.63 mm</quantity>
, PW
<quantity id="4CB6E3A3FFDC502CFD4EF90EFCA442AF" box="[762,865,1730,1757]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="44" pageNumber="45" unit="mm" value="0.55">0.55 mm</quantity>
; EL
<quantity id="4CB6E3A3FFDC502CFC20F90EFC3F42AF" box="[916,1018,1730,1756]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="44" pageNumber="45" unit="mm" value="1.3">1.30 mm</quantity>
, EW
<quantity id="4CB6E3A3FFDC502CFB8CF90EFB1842AF" box="[1080,1245,1730,1757]" metricMagnitude="-4" metricUnit="m" metricValue="8.65" metricValueMax="8.8" metricValueMin="8.5" pageId="44" pageNumber="45" unit="mm" value="0.865" valueMax="0.88" valueMin="0.85">0.850.88 mm</quantity>
, EI 1.491.53.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFDC502CFF73F929FC884373" box="[199,845,1765,1792]" pageId="44" pageNumber="45" type="distribution">
<paragraph id="8BF14E46FFDC502CFF73F929FC884373" blockId="44.[151,1438,325,1936]" box="[199,845,1765,1792]" pageId="44" pageNumber="45">
<emphasis id="B93A9254FFDC502CFF73F929FEA4428C" bold="true" box="[199,353,1765,1791]" pageId="44" pageNumber="45">Distribution.</emphasis>
Northernmost
<collectingCountry id="F3590ED6FFDC502CFDBEF92AFD544373" box="[522,657,1766,1792]" name="Madagascar" pageId="44" pageNumber="45">Madagascar</collectingCountry>
(Diana Region).
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFDC502CFF73F8C6FC0743E3" pageId="44" pageNumber="45" type="discussion">
<paragraph id="8BF14E46FFDC502CFF73F8C6FF35433B" blockId="44.[151,1438,325,1936]" pageId="44" pageNumber="45">
<emphasis id="B93A9254FFDC502CFF73F8C6FEFE4357" bold="true" box="[199,315,1802,1828]" pageId="44" pageNumber="45">Remarks.</emphasis>
<taxonomicName id="4C4E35C5FFDC502CFEFCF8C6FD704357" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[328,693,1802,1828]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="roussettensis">
<emphasis id="B93A9254FFDC502CFEFCF8C6FD704357" box="[328,693,1802,1828]" italics="true" pageId="44" pageNumber="45">Anhoraeomorphus roussettensis</emphasis>
</taxonomicName>
is highly similar to
<taxonomicName id="4C4E35C5FFDC502CFC07F8C6FB944357" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[947,1105,1802,1828]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="anosyensis">
<emphasis id="B93A9254FFDC502CFC07F8C6FB944357" box="[947,1105,1802,1828]" italics="true" pageId="44" pageNumber="45">A. anosyensis</emphasis>
</taxonomicName>
, see Remarks for the latter species.
</paragraph>
<paragraph id="8BF14E46FFDC502CFF73F89EFC0743E3" blockId="44.[151,1438,325,1936]" pageId="44" pageNumber="45">
<materialsCitation id="3B26441BFFDC502CFF73F89EFC7B43E3" collectionCode="NHMW" pageId="44" pageNumber="45" specimenCount="5" specimenCount-female="1" specimenCount-male="2" typeStatus="holotype">
According to
<bibRefCitation id="EFDF33B7FFDC502CFED5F89EFDC6431F" author="Franz, H." box="[353,515,1874,1900]" pageId="44" pageNumber="45" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986b)</bibRefCitation>
, in addition to the
<typeStatus id="54F5F0E4FFDC502CFD61F89EFCF2431F" box="[725,823,1874,1900]" pageId="44" pageNumber="45" type="holotype">holotype</typeStatus>
and
<specimenCount id="9D4885CFFFDC502CFCD9F89EFBCC431F" box="[877,1033,1874,1900]" count="2" pageId="44" pageNumber="45" type="generic" typeStatus="paratype">two paratypes</specimenCount>
deposited at
<collectionCode id="ED5FD683FFDC502CFB2AF89EFB3F431F" box="[1182,1274,1874,1900]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="44" pageNumber="45" type="Museum">NHMW</collectionCode>
,
<specimenCount id="9D4885CFFFDC502CFAB1F89EFAA9431F" box="[1285,1388,1874,1900]" count="2" pageId="44" pageNumber="45" type="male">two male</specimenCount>
and
<specimenCount id="9D4885CFFFDC502CFF23F8BAFED643E3" box="[151,275,1910,1936]" count="1" pageId="44" pageNumber="45" type="female">one female</specimenCount>
<typeStatus id="54F5F0E4FFDC502CFEAEF8BAFEBF43E3" box="[282,378,1910,1936]" pageId="44" pageNumber="45" type="paratype">paratype</typeStatus>
are in Muséum National d'Histoire Naturelle, Paris
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,256 @@
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<subSubSection id="C3541DCDFFDE502EFF23FF5BFD7044C2" box="[151,693,151,178]" pageId="46" pageNumber="47" type="nomenclature">
<paragraph id="8BF14E46FFDE502EFF23FF5BFD7044C2" blockId="46.[151,693,151,213]" box="[151,693,151,178]" pageId="46" pageNumber="47">
<heading id="D0B9F92AFFDE502EFF23FF5BFD7044C2" bold="true" box="[151,693,151,178]" fontSize="11" level="2" pageId="46" pageNumber="47" reason="4">
<emphasis id="B93A9254FFDE502EFF23FF5BFD7044C2" bold="true" box="[151,693,151,178]" pageId="46" pageNumber="47">
<taxonomicName id="4C4E35C5FFDE502EFF23FF5BFDF844C2" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[151,573,151,178]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="sikorai" status="comb. nov.">
<emphasis id="B93A9254FFDE502EFF23FF5BFE1344C1" bold="true" box="[151,470,151,178]" italics="true" pageId="46" pageNumber="47">Anhoraeomorphus sikorai</emphasis>
(Franz)
</taxonomicName>
,
<taxonomicNameLabel id="A2092F2FFFDE502EFDFEFF5BFD7044C2" box="[586,693,151,177]" pageId="46" pageNumber="47" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFDE502EFF23FF77FE8644A6" box="[151,323,187,213]" pageId="46" pageNumber="47" type="description">
<paragraph id="8BF14E46FFDE502EFF23FF77FE8644A6" blockId="46.[151,693,151,213]" box="[151,323,187,213]" pageId="46" pageNumber="47">
(
<figureCitation id="137552C3FFDE502EFF2BFF77FEFF44A6" box="[159,314,187,213]" captionStart="FIGURES 171178" captionStartId="47.[151,264,1540,1564]" captionTargetBox="[151,1407,181,1508]" captionTargetId="figure-16@47.[151,1435,181,1516]" captionTargetPageId="47" captionText="FIGURES 171178. Anhoraeomorphus sikorai (Franz), holotype. Habitus in dorsal view (171); head in anterodorsal view (172); head and prothorax in lateral view (only bristles on head are shown) (173); antenna (174); aedeagus in ventral (175, 177) and lateral (176, 178) views." figureDoi="http://doi.org/10.5281/zenodo.13745981" httpUri="https://zenodo.org/record/13745981/files/figure.png" pageId="46" pageNumber="47">Figs 171178</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFDE502EFF23FECCFD954564" box="[151,592,255,280]" pageId="46" pageNumber="47" type="reference_group">
<paragraph id="8BF14E46FFDE502EFF23FECCFD954564" blockId="46.[151,592,255,280]" box="[151,592,255,280]" pageId="46" pageNumber="47">
<treatmentCitationGroup id="AB5E6968FFDE502EFF23FECCFD954564" box="[151,592,255,280]" pageId="46" pageNumber="47">
<treatmentCitation id="0AEF6857FFDE502EFF23FECCFD894564" author="Franz, H." box="[151,588,255,280]" page="162" pageId="46" pageNumber="47" year="1986">
<taxonomicName id="4C4E35C5FFDE502EFF23FECCFD894564" ID-CoL="3MNLK" authority="Franz, 1986 b: 162" authorityName="Franz" authorityPageNumber="162" authorityYear="1986" box="[151,588,255,280]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="sikorai">
<emphasis id="B93A9254FFDE502EFF23FECCFE4F456B" box="[151,394,256,280]" italics="true" pageId="46" pageNumber="47">Horaeomorphus sikorai</emphasis>
<bibRefCitation id="EFDF33B7FFDE502EFE24FECCFD894564" author="Franz, H." box="[400,588,255,279]" pageId="46" pageNumber="47" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 162</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFDE502EFF23FE8AFD05469F" pageId="46" pageNumber="47" type="materials_examined">
<paragraph id="8BF14E46FFDE502EFF23FE8AFA8245DB" blockId="46.[151,1437,325,1936]" pageId="46" pageNumber="47">
<emphasis id="B93A9254FFDE502EFF23FE8AFE77452C" bold="true" box="[151,434,325,352]" pageId="46" pageNumber="47">Type material studied.</emphasis>
<materialsCitation id="3B26441BFFDE502EFE75FE8AFECC45DB" collectionCode="NHMW" country="Madagascar" determinerName="H. Franz" location="Madagascar" pageId="46" pageNumber="47" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B93A9254FFDE502EFE75FE8AFD26452C" bold="true" box="[449,739,325,352]" pageId="46" pageNumber="47">
<typeStatus id="54F5F0E4FFDE502EFE75FE8AFDEB4513" box="[449,558,326,352]" pageId="46" pageNumber="47" type="holotype">Holotype</typeStatus>
(
<collectingCountry id="F3590ED6FFDE502EFDF0FE89FD1E452C" box="[580,731,325,351]" name="Madagascar" pageId="46" pageNumber="47">Madagascar</collectingCountry>
)
</emphasis>
:
<specimenCount id="9D4885CFFFDE502EFD4DFE8AFCD54513" box="[761,784,326,352]" count="1" pageId="46" pageNumber="47" type="male"></specimenCount>
, three labels: “Madagaskar / Sikora” [white, printed], “
<taxonomicName id="4C4E35C5FFDE502EFF15FEA6FE9D45F7" box="[161,344,362,388]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/ sikorai m. / det.
<determinerName id="7E2DA282FFDE502EFD81FEA6FD5645F7" box="[565,659,362,388]" pageId="46" pageNumber="47">H.Franz</determinerName>
” [yellowish, handwritten and printed], “Typus” [red, handwritten] (
<collectionCode id="ED5FD683FFDE502EFF2BFE42FEC745DB" box="[159,258,398,424]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="46" pageNumber="47" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B26441BFFDE502EFEA0FE42FA8145DB" box="[276,1348,398,424]" collectionCode="NHMW" country="Madagascar" determinerName="H. Franz" location="Madagascar" pageId="46" pageNumber="47" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<emphasis id="B93A9254FFDE502EFEA0FE42FE5045DB" bold="true" box="[276,405,398,424]" pageId="46" pageNumber="47">
<typeStatus id="54F5F0E4FFDE502EFEA0FE42FE5545DB" box="[276,400,398,424]" pageId="46" pageNumber="47" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D4885CFFFDE502EFE28FE42FE1945DB" box="[412,476,398,424]" count="2" pageId="46" pageNumber="47" type="female">2 ♀♀</specimenCount>
, same data as for holotype, but with yellow “
<typeStatus id="54F5F0E4FFDE502EFC61FE42FBBD45DB" box="[981,1144,398,424]" pageId="46" pageNumber="47" type="paratype">PARATYPUS</typeStatus>
” labels (
<collectionCode id="ED5FD683FFDE502EFB6DFE42FAF945DB" box="[1241,1340,398,424]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="46" pageNumber="47" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF14E46FFDE502EFF73FE7DFD05469F" blockId="46.[151,1437,325,1936]" pageId="46" pageNumber="47">
<emphasis id="B93A9254FFDE502EFF73FE7DFE6345B8" bold="true" box="[199,422,433,459]" pageId="46" pageNumber="47">Revised diagnosis.</emphasis>
Body (
<figureCitation id="137552C3FFDE502EFDB7FE7EFDA245BF" box="[515,615,434,460]" captionStart="FIGURES 171178" captionStartId="47.[151,264,1540,1564]" captionTargetBox="[151,1407,181,1508]" captionTargetId="figure-16@47.[151,1435,181,1516]" captionTargetPageId="47" captionText="FIGURES 171178. Anhoraeomorphus sikorai (Franz), holotype. Habitus in dorsal view (171); head in anterodorsal view (172); head and prothorax in lateral view (only bristles on head are shown) (173); antenna (174); aedeagus in ventral (175, 177) and lateral (176, 178) views." figureDoi="http://doi.org/10.5281/zenodo.13745981" httpUri="https://zenodo.org/record/13745981/files/figure.png" pageId="46" pageNumber="47">Fig. 171</figureCitation>
) slender; head (
<figureCitation id="137552C3FFDE502EFC9DFE7EFC0C45BF" box="[809,969,434,460]" captionStart="FIGURES 171178" captionStartId="47.[151,264,1540,1564]" captionTargetBox="[151,1407,181,1508]" captionTargetId="figure-16@47.[151,1435,181,1516]" captionTargetPageId="47" captionText="FIGURES 171178. Anhoraeomorphus sikorai (Franz), holotype. Habitus in dorsal view (171); head in anterodorsal view (172); head and prothorax in lateral view (only bristles on head are shown) (173); antenna (174); aedeagus in ventral (175, 177) and lateral (176, 178) views." figureDoi="http://doi.org/10.5281/zenodo.13745981" httpUri="https://zenodo.org/record/13745981/files/figure.png" pageId="46" pageNumber="47">Figs 172173</figureCitation>
) round, with vertex posteriorly broadly rounded; pronotum (
<figureCitation id="137552C3FFDE502EFE37FE1AFE204583" box="[387,485,470,496]" captionStart="FIGURES 171178" captionStartId="47.[151,264,1540,1564]" captionTargetBox="[151,1407,181,1508]" captionTargetId="figure-16@47.[151,1435,181,1516]" captionTargetPageId="47" captionText="FIGURES 171178. Anhoraeomorphus sikorai (Franz), holotype. Habitus in dorsal view (171); head in anterodorsal view (172); head and prothorax in lateral view (only bristles on head are shown) (173); antenna (174); aedeagus in ventral (175, 177) and lateral (176, 178) views." figureDoi="http://doi.org/10.5281/zenodo.13745981" httpUri="https://zenodo.org/record/13745981/files/figure.png" pageId="46" pageNumber="47">Fig. 171</figureCitation>
) bell-shaped, broadest in anterior third, with one pair of large transverse antebasal pits; antennomere 3 (
<figureCitation id="137552C3FFDE502EFECBFE36FE184667" box="[383,477,506,532]" captionStart="FIGURES 171178" captionStartId="47.[151,264,1540,1564]" captionTargetBox="[151,1407,181,1508]" captionTargetId="figure-16@47.[151,1435,181,1516]" captionTargetPageId="47" captionText="FIGURES 171178. Anhoraeomorphus sikorai (Franz), holotype. Habitus in dorsal view (171); head in anterodorsal view (172); head and prothorax in lateral view (only bristles on head are shown) (173); antenna (174); aedeagus in ventral (175, 177) and lateral (176, 178) views." figureDoi="http://doi.org/10.5281/zenodo.13745981" httpUri="https://zenodo.org/record/13745981/files/figure.png" pageId="46" pageNumber="47">Fig. 174</figureCitation>
) as long as broad, 45 each slightly elongate, 6 as long as broad, 710 each indistinctly transverse; profemora similarly slender as meso- and metafemora; metatibiae in male modified, but only slightly strongly broadening to distal third and then narrowing to apex; aedeagus (
<figureCitation id="137552C3FFDE502EFC6AFD8EFBBE462E" box="[990,1147,578,605]" captionStart="FIGURES 171178" captionStartId="47.[151,264,1540,1564]" captionTargetBox="[151,1407,181,1508]" captionTargetId="figure-16@47.[151,1435,181,1516]" captionTargetPageId="47" captionText="FIGURES 171178. Anhoraeomorphus sikorai (Franz), holotype. Habitus in dorsal view (171); head in anterodorsal view (172); head and prothorax in lateral view (only bristles on head are shown) (173); antenna (174); aedeagus in ventral (175, 177) and lateral (176, 178) views." figureDoi="http://doi.org/10.5281/zenodo.13745981" httpUri="https://zenodo.org/record/13745981/files/figure.png" pageId="46" pageNumber="47">Figs 175178</figureCitation>
) in ventral view broadest near basal third, apical region of dorsal wall in lateral view weakly curved dorsally, distolateral endophallic sclerites separated except their apices touching at middle, each strongly elongate and with distal margin concave with outer and inner subtriangular apex, parameres with several subapical setae distributed along more than half of length and increasing in length from proximal to distal ones.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFDE502EFF73FD39FAB24373" pageId="46" pageNumber="47" type="description">
<paragraph id="8BF14E46FFDE502EFF73FD39FCAF4747" blockId="46.[151,1437,325,1936]" pageId="46" pageNumber="47">
<emphasis id="B93A9254FFDE502EFF73FD39FEB0477C" bold="true" box="[199,373,757,783]" pageId="46" pageNumber="47">Redescription.</emphasis>
Body of male (
<figureCitation id="137552C3FFDE502EFDABFD3AFDB84763" box="[543,637,758,784]" captionStart="FIGURES 171178" captionStartId="47.[151,264,1540,1564]" captionTargetBox="[151,1407,181,1508]" captionTargetId="figure-16@47.[151,1435,181,1516]" captionTargetPageId="47" captionText="FIGURES 171178. Anhoraeomorphus sikorai (Franz), holotype. Habitus in dorsal view (171); head in anterodorsal view (172); head and prothorax in lateral view (only bristles on head are shown) (173); antenna (174); aedeagus in ventral (175, 177) and lateral (176, 178) views." figureDoi="http://doi.org/10.5281/zenodo.13745981" httpUri="https://zenodo.org/record/13745981/files/figure.png" pageId="46" pageNumber="47">Fig. 171</figureCitation>
) moderately dark brown, elytra slightly lighter than head and pronotum, tarsi and palps distinctly lighter, setae light brown; BL
<quantity id="4CB6E3A3FFDE502EFD4AFCD6FCA04747" box="[766,869,794,821]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" pageId="46" pageNumber="47" unit="mm" value="2.15">2.15 mm</quantity>
.
</paragraph>
<paragraph id="8BF14E46FFDE502EFF73FCF2FDF44027" blockId="46.[151,1437,325,1936]" pageId="46" pageNumber="47">
Head (
<figureCitation id="137552C3FFDE502EFEA0FCF2FEB0472B" box="[276,373,830,856]" captionStart="FIGURES 171178" captionStartId="47.[151,264,1540,1564]" captionTargetBox="[151,1407,181,1508]" captionTargetId="figure-16@47.[151,1435,181,1516]" captionTargetPageId="47" captionText="FIGURES 171178. Anhoraeomorphus sikorai (Franz), holotype. Habitus in dorsal view (171); head in anterodorsal view (172); head and prothorax in lateral view (only bristles on head are shown) (173); antenna (174); aedeagus in ventral (175, 177) and lateral (176, 178) views." figureDoi="http://doi.org/10.5281/zenodo.13745981" httpUri="https://zenodo.org/record/13745981/files/figure.png" pageId="46" pageNumber="47">Fig. 172</figureCitation>
) in anterodorsal view round, broadest at eyes, HL
<quantity id="4CB6E3A3FFDE502EFC0BFCF2FBE2472B" box="[959,1063,830,856]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="46" pageNumber="47" unit="mm" value="0.38">0.38 mm</quantity>
, HW
<quantity id="4CB6E3A3FFDE502EFBD8FCF2FB10472B" box="[1132,1237,830,856]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="46" pageNumber="47" unit="mm" value="0.38">0.38 mm</quantity>
; temple in lateral view (
<figureCitation id="137552C3FFDE502EFF69FCAEFEF9470F" box="[221,316,866,892]" captionStart="FIGURES 171178" captionStartId="47.[151,264,1540,1564]" captionTargetBox="[151,1407,181,1508]" captionTargetId="figure-16@47.[151,1435,181,1516]" captionTargetPageId="47" captionText="FIGURES 171178. Anhoraeomorphus sikorai (Franz), holotype. Habitus in dorsal view (171); head in anterodorsal view (172); head and prothorax in lateral view (only bristles on head are shown) (173); antenna (174); aedeagus in ventral (175, 177) and lateral (176, 178) views." figureDoi="http://doi.org/10.5281/zenodo.13745981" httpUri="https://zenodo.org/record/13745981/files/figure.png" pageId="46" pageNumber="47">Fig. 174</figureCitation>
) subequal to eye; vertex and frons confluent, together weakly convex and weakly transverse; vertex posteriorly broadly rounded; each eye large and weakly convex, shallowly emarginate posteriorly, weakly projecting from head silhouette. Punctures on vertex fine, inconspicuous; setae short, dense and suberect; frons and vertex densely covered with thick bristles directed posteriorly. Antenna (
<figureCitation id="137552C3FFDE502EFCCAFC02FC1B479B" box="[894,990,974,1000]" captionStart="FIGURES 171178" captionStartId="47.[151,264,1540,1564]" captionTargetBox="[151,1407,181,1508]" captionTargetId="figure-16@47.[151,1435,181,1516]" captionTargetPageId="47" captionText="FIGURES 171178. Anhoraeomorphus sikorai (Franz), holotype. Habitus in dorsal view (171); head in anterodorsal view (172); head and prothorax in lateral view (only bristles on head are shown) (173); antenna (174); aedeagus in ventral (175, 177) and lateral (176, 178) views." figureDoi="http://doi.org/10.5281/zenodo.13745981" httpUri="https://zenodo.org/record/13745981/files/figure.png" pageId="46" pageNumber="47">Fig. 174</figureCitation>
) slender but short, slightly shorter than half BL, AnL
<quantity id="4CB6E3A3FFDE502EFE81FC3EFE59407F" box="[309,412,1010,1036]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="46" pageNumber="47" unit="mm" value="0.9">0.90 mm</quantity>
, antennomeres 12 each elongate, 3 as long as broad, 45 each slightly elongate, 6 as long as broad, 710 each indistinctly transverse, 11 distinctly shorter than 9 and 10 combined, indistinctly narrower than 10, about 1.7 times as long as broad.
</paragraph>
<paragraph id="8BF14E46FFDE502EFF73FB92FF35417B" blockId="46.[151,1437,325,1936]" pageId="46" pageNumber="47">
Pronotum bell-shaped, broadest in anterior third; PL
<quantity id="4CB6E3A3FFDE502EFCA4FB92FCB0400B" box="[784,885,1118,1145]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="46" pageNumber="47" unit="mm" value="0.5">0.50 mm</quantity>
, PW
<quantity id="4CB6E3A3FFDE502EFC04FB92FBD3400B" box="[944,1046,1118,1145]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="46" pageNumber="47" unit="mm" value="0.5">0.50 mm</quantity>
. Anterior margin strongly rounded, lateral margins rounded in anterior half, sinuate in posterior half; posterior corners blunt; posterior margin nearly straight. Base with one pair of large transversely oval pits, and with short but distinct sublateral carinae. Disc with fine, inconspicuous punctures; setae dense, short and suberect, lateral surface of pronotum with dense thick bristles.
</paragraph>
<paragraph id="8BF14E46FFDE502EFF73FADEFEAE4107" blockId="46.[151,1437,325,1936]" pageId="46" pageNumber="47">
Elytra together oval, broadest distinctly in front of middle; EL
<quantity id="4CB6E3A3FFDE502EFC39FADEFC31415F" box="[909,1012,1298,1324]" metricMagnitude="-3" metricUnit="m" metricValue="1.28" pageId="46" pageNumber="47" unit="mm" value="1.28">1.28 mm</quantity>
, EW
<quantity id="4CB6E3A3FFDE502EFB87FADEFB5F415F" box="[1075,1178,1298,1324]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="46" pageNumber="47" unit="mm" value="0.8">0.80 mm</quantity>
, EI 1.59; humeral calli weakly elevated, basal elytral foveae vestigial, barely discernible. Punctures fine and inconspicuous; setae sparse, short and suberect.
</paragraph>
<paragraph id="8BF14E46FFDE502EFF73FAB2FE8C41CF" blockId="46.[151,1437,325,1936]" pageId="46" pageNumber="47">Legs moderately long, slender, mesotibiae distinctly curved, metatibiae slightly broadening to distal third and then narrowing.</paragraph>
<paragraph id="8BF14E46FFDE502EFF73FA0AFAFE42CB" blockId="46.[151,1437,325,1936]" pageId="46" pageNumber="47">
Aedeagus (
<figureCitation id="137552C3FFDE502EFEF0FA0AFE1A4192" box="[324,479,1478,1505]" captionStart="FIGURES 171178" captionStartId="47.[151,264,1540,1564]" captionTargetBox="[151,1407,181,1508]" captionTargetId="figure-16@47.[151,1435,181,1516]" captionTargetPageId="47" captionText="FIGURES 171178. Anhoraeomorphus sikorai (Franz), holotype. Habitus in dorsal view (171); head in anterodorsal view (172); head and prothorax in lateral view (only bristles on head are shown) (173); antenna (174); aedeagus in ventral (175, 177) and lateral (176, 178) views." figureDoi="http://doi.org/10.5281/zenodo.13745981" httpUri="https://zenodo.org/record/13745981/files/figure.png" pageId="46" pageNumber="47">Figs 175178</figureCitation>
) stout, AeL
<quantity id="4CB6E3A3FFDE502EFDD2FA0AFD084193" box="[614,717,1478,1505]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="46" pageNumber="47" unit="mm" value="0.35">0.35 mm</quantity>
; in ventral view drop-shaped, distally rapidly narrowing towards broadly subtrapezoidal apex with rounded distal margin; in lateral view apical region of dorsal wall weakly curved dorsally; endophallus with asymmetrical and long distolateral sclerites and with strongly asymmetrical assemblage of median proximal sclerotized structures, distolateral sclerites separated except for their tips touching at middle, each with concave distal margin and outer and inner subtriangular apices; parameres strongly broadening distally, their apices slightly projecting beyond apex of median lobe, each paramere with subapical group of setae distributed along more than distal half of outer lateral margin, setae increasing in length from proximal to distal ones.
</paragraph>
<paragraph id="8BF14E46FFDE502EFF73F90EFAB24373" blockId="46.[151,1437,325,1936]" pageId="46" pageNumber="47">
Female. Externally similar to male, but with slenderer hind tibiae. BL
<quantity id="4CB6E3A3FFDE502EFC6CF90EFBB842AF" box="[984,1149,1730,1757]" metricMagnitude="-3" metricUnit="m" metricValue="2.19" metricValueMax="2.23" metricValueMin="2.15" pageId="46" pageNumber="47" unit="mm" value="2.19" valueMax="2.23" valueMin="2.15">2.152.23 mm</quantity>
; HL 0.400.43
<quantity id="4CB6E3A3FFDE502EFA87F90EFA5D42AF" box="[1331,1432,1730,1756]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="46" pageNumber="47" unit="mm" value="0.43">0.43 mm</quantity>
, HW
<quantity id="4CB6E3A3FFDE502EFF78F92AFEF74373" box="[204,306,1766,1792]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="46" pageNumber="47" unit="mm" value="0.43">0.43 mm</quantity>
, AnL
<quantity id="4CB6E3A3FFDE502EFEC2F92AFDD94373" box="[374,540,1766,1792]" metricMagnitude="-4" metricUnit="m" metricValue="9.55" metricValueMax="10.3" metricValueMin="8.8" pageId="46" pageNumber="47" unit="mm" value="0.955" valueMax="1.03" valueMin="0.88">0.881.03 mm</quantity>
; PL
<quantity id="4CB6E3A3FFDE502EFDFAF92AFD364373" box="[590,755,1766,1793]" metricMagnitude="-4" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="46" pageNumber="47" unit="mm" value="0.525" valueMax="0.55" valueMin="0.5">0.500.55 mm</quantity>
, PW
<quantity id="4CB6E3A3FFDE502EFC9BF92AFC104373" box="[815,981,1766,1793]" metricMagnitude="-4" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="46" pageNumber="47" unit="mm" value="0.525" valueMax="0.55" valueMin="0.5">0.500.55 mm</quantity>
; EL
<quantity id="4CB6E3A3FFDE502EFBBCF92AFBAB4373" box="[1032,1134,1766,1793]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="46" pageNumber="47" unit="mm" value="1.25">1.25 mm</quantity>
, EW
<quantity id="4CB6E3A3FFDE502EFB18F92AFAD74373" box="[1196,1298,1766,1793]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="46" pageNumber="47" unit="mm" value="0.85">0.85 mm</quantity>
, EI 1.47.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFDE502EFF73F8C5FD5B4357" box="[199,670,1801,1828]" pageId="46" pageNumber="47" type="distribution">
<paragraph id="8BF14E46FFDE502EFF73F8C5FD5B4357" blockId="46.[151,1437,325,1936]" box="[199,670,1801,1828]" pageId="46" pageNumber="47">
<emphasis id="B93A9254FFDE502EFF73F8C5FEA44350" bold="true" box="[199,353,1801,1827]" pageId="46" pageNumber="47">Distribution.</emphasis>
<collectingCountry id="F3590ED6FFDE502EFED3F8C6FE2B4357" box="[359,494,1802,1828]" name="Madagascar" pageId="46" pageNumber="47">Madagascar</collectingCountry>
(see Remarks).
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFDE502FFF73F8E2FE16434E" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47" type="discussion">
<paragraph id="8BF14E46FFDE502EFF73F8E2FB2043E3" blockId="46.[151,1437,325,1936]" pageId="46" pageNumber="47">
<emphasis id="B93A9254FFDE502EFF73F8E2FEFE433B" bold="true" box="[199,315,1838,1864]" pageId="46" pageNumber="47">Remarks.</emphasis>
<taxonomicName id="4C4E35C5FFDE502EFEF2F8E2FDB5433B" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[326,624,1838,1864]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="sikorai">
<emphasis id="B93A9254FFDE502EFEF2F8E2FDB5433B" box="[326,624,1838,1864]" italics="true" pageId="46" pageNumber="47">Anhoraeomorphus sikorai</emphasis>
</taxonomicName>
is unremarkable and the aedeagus must be examined to identify this species. The large asymmetrical endophallic distolateral sclerites touching distally at middle, each with concave distal margin with inner and outer subtriangular process are unique among
<taxonomicName id="4C4E35C5FFDE502EFC6BF8BAFB2043E3" box="[991,1253,1910,1936]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B93A9254FFDE502EFC6BF8BAFB6A43E3" box="[991,1199,1910,1936]" italics="true" pageId="46" pageNumber="47">Anhoraeomorphus</emphasis>
spp.
</taxonomicName>
</paragraph>
<caption id="DF311ECEFFDF502FFF23F9C8FE7F4217" ID-DOI="http://doi.org/10.5281/zenodo.13745981" ID-Zenodo-Dep="13745981" httpUri="https://zenodo.org/record/13745981/files/figure.png" pageId="47" pageNumber="48" startId="47.[151,264,1540,1564]" targetBox="[151,1407,181,1508]" targetPageId="47" targetType="figure">
<paragraph id="8BF14E46FFDF502FFF23F9C8FE7F4217" blockId="47.[151,1436,1540,1637]" pageId="47" pageNumber="48">
<emphasis id="B93A9254FFDF502FFF23F9C8FEB5426F" bold="true" box="[151,368,1540,1564]" pageId="47" pageNumber="48">FIGURES 171178.</emphasis>
<taxonomicName id="4C4E35C5FFDF502FFECDF9C8FD1E426F" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[377,731,1540,1564]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="sikorai">
<emphasis id="B93A9254FFDF502FFECDF9C8FD40426F" box="[377,645,1540,1564]" italics="true" pageId="47" pageNumber="48">Anhoraeomorphus sikorai</emphasis>
(Franz)
</taxonomicName>
, holotype. Habitus in dorsal view (
<emphasis id="B93A9254FFDF502FFBE4F9C8FBB3426F" bold="true" box="[1104,1142,1540,1564]" pageId="47" pageNumber="48">171</emphasis>
); head in anterodorsal view (
<emphasis id="B93A9254FFDF502FFF14F9E4FF034233" bold="true" box="[160,198,1576,1600]" pageId="47" pageNumber="48">172</emphasis>
); head and prothorax in lateral view (only bristles on head are shown) (
<emphasis id="B93A9254FFDF502FFC23F9E4FC784233" bold="true" box="[919,957,1576,1600]" pageId="47" pageNumber="48">173</emphasis>
); antenna (
<emphasis id="B93A9254FFDF502FFB9BF9E4FB904233" bold="true" box="[1071,1109,1576,1600]" pageId="47" pageNumber="48">174</emphasis>
); aedeagus in ventral (
<emphasis id="B93A9254FFDF502FFA88F9E4FA514233" bold="true" box="[1340,1428,1576,1600]" pageId="47" pageNumber="48">175, 177</emphasis>
) and lateral (
<emphasis id="B93A9254FFDF502FFEA4F980FEAC4217" bold="true" box="[272,361,1612,1636]" pageId="47" pageNumber="48">176, 178</emphasis>
) views.
</paragraph>
</caption>
<paragraph id="8BF14E46FFDF502FFF73F97AFE16434E" blockId="47.[151,1437,1718,1853]" pageId="47" pageNumber="48">
The specimens used by Franz to describe
<taxonomicName id="4C4E35C5FFDF502FFD1DF97BFCDD42A3" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[681,792,1718,1745]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="sikorai">
<emphasis id="B93A9254FFDF502FFD1DF97BFCDD42A3" box="[681,792,1718,1745]" italics="true" pageId="47" pageNumber="48">A. sikorai</emphasis>
</taxonomicName>
were deposited at NHMW before he worked in Vienna. The exact collecting site remains unknown. These specimens were collected by Franz Sikora (18631902), an Austrian traveler and collector of various biological specimens active in
<collectingCountry id="F3590ED6FFDF502FFC68F932FBA6436B" box="[988,1123,1790,1816]" name="Madagascar" pageId="47" pageNumber="48">Madagascar</collectingCountry>
and
<collectingCountry id="F3590ED6FFDF502FFB2BF932FB3B436B" box="[1183,1278,1790,1816]" name="Réunion" pageId="47" pageNumber="48">Réunion</collectingCountry>
during 1890s (
<emphasis id="B93A9254FFDF502FFF15F8EFFF0C434F" box="[161,201,1827,1852]" italics="true" pageId="47" pageNumber="48">e.g.</emphasis>
,
<bibRefCitation id="EFDF33B7FFDF502FFF63F8EEFE0A434E" author="Bolotov, I. N. &amp; Eliseeva, T. A. &amp; Tsiplenkina, I. G. &amp; Gofarov, M. Y. &amp; Kondakov, A. V." box="[215,463,1826,1853]" pageId="47" pageNumber="48" pagination="56 - 67" refId="ref52681" refString="Bolotov, I. N., Eliseeva, T. A., Tsiplenkina, I. G., Gofarov, M. Y. &amp; Kondakov, A. V. (2023) Long forgotten record of a freshwater leech on Madagascar indicates a host-associated dispersal event from continental Africa. Ecologica Montenegrina, 61, 56 - 67. https: // doi. org / 10.37828 / em. 2023.61.7" type="journal article" year="2023">
Bolotov
<emphasis id="B93A9254FFDF502FFE8CF8EFFEB5434F" box="[312,368,1826,1852]" italics="true" pageId="47" pageNumber="48">et al.</emphasis>
(2023))
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,216 @@
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<mods:title id="E4CC5769CA7461936E8687E940A40646">The curse of Horaeomorphus (almost) lifted. Revision of misplaced species from Madagascar, Comoros and Mascarenes (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<paragraph id="8BF14E46FFE05010FF23FBAEFD3A400E" blockId="16.[151,767,1122,1185]" box="[151,767,1122,1149]" pageId="16" pageNumber="17">
<heading id="D0B9F92AFFE05010FF23FBAEFD3A400E" bold="true" box="[151,767,1122,1149]" fontSize="11" level="2" pageId="16" pageNumber="17" reason="4">
<emphasis id="B93A9254FFE05010FF23FBAEFD3A400E" bold="true" box="[151,767,1122,1149]" pageId="16" pageNumber="17">
<taxonomicName id="4C4E35C5FFE05010FF23FBAEFD43400F" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[151,646,1122,1149]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="sakarahanus" status="comb. nov.">
<emphasis id="B93A9254FFE05010FF23FBAEFDDA400E" bold="true" box="[151,543,1122,1149]" italics="true" pageId="16" pageNumber="17">Anhoraeomorphus sakarahanus</emphasis>
(Franz)
</taxonomicName>
,
<taxonomicNameLabel id="A2092F2FFFE05010FD20FBAFFD3A400E" box="[660,767,1123,1149]" pageId="16" pageNumber="17" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C3541DCDFFE05010FF23FB4BFEE240D2" box="[151,295,1158,1185]" pageId="16" pageNumber="17" type="description">
<paragraph id="8BF14E46FFE05010FF23FB4BFEE240D2" blockId="16.[151,767,1122,1185]" box="[151,295,1158,1185]" pageId="16" pageNumber="17">
(
<figureCitation id="137552C3FFE05010FF2BFB4BFEDA40D2" box="[159,287,1158,1185]" captionStart="FIGURES 4451" captionStartId="17.[151,264,1523,1547]" captionTargetBox="[151,1422,181,1488]" captionTargetId="figure-16@17.[151,1435,181,1500]" captionTargetPageId="17" captionText="FIGURES 4451. Anhoraeomorphus sakarahanus (Franz), holotype. Habitus in dorsal view (44); head in anterodorsal view (45); head and prothorax in lateral view (only bristles on head are shown) (46); antenna (47); aedeagus in ventral (48, 50) and lateral (49, 51) views." figureDoi="http://doi.org/10.5281/zenodo.13745937" httpUri="https://zenodo.org/record/13745937/files/figure.png" pageId="16" pageNumber="17">Figs 4451</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFE05010FF23FB07FD4E4090" box="[151,651,1227,1251]" pageId="16" pageNumber="17" type="reference_group">
<paragraph id="8BF14E46FFE05010FF23FB07FD4E4090" blockId="16.[151,651,1227,1251]" box="[151,651,1227,1251]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="AB5E6968FFE05010FF23FB07FD4E4090" box="[151,651,1227,1251]" pageId="16" pageNumber="17">
<treatmentCitation id="0AEF6857FFE05010FF23FB07FD424090" author="Franz, H." box="[151,647,1227,1251]" page="165" pageId="16" pageNumber="17" year="1986">
<taxonomicName id="4C4E35C5FFE05010FF23FB07FD424090" ID-CoL="3MNLF" authority="Franz, 1986 b: 165" authorityName="Franz" authorityPageNumber="165" authorityYear="1986" box="[151,647,1227,1251]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="sakarahanus">
<emphasis id="B93A9254FFE05010FF23FB07FE004090" box="[151,453,1227,1251]" italics="true" pageId="16" pageNumber="17">Horaeomorphus sakarahanus</emphasis>
<bibRefCitation id="EFDF33B7FFE05010FE7FFB00FD424090" author="Franz, H." box="[459,647,1227,1251]" pageId="16" pageNumber="17" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 165</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFE05010FF23FADDFEE24238" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8BF14E46FFE05010FF23FADDFC5F4100" blockId="16.[151,1437,1297,1972]" pageId="16" pageNumber="17">
<emphasis id="B93A9254FFE05010FF23FADDFE6E4158" bold="true" box="[151,427,1297,1323]" pageId="16" pageNumber="17">Type material studied.</emphasis>
<materialsCitation id="3B26441BFFE05010FE02FADDFC5F4100" collectionCode="NHMW" collectorName="H. Franz" country="Madagascar" determinerName="H. Franz" location="Foret de Sakahara" pageId="16" pageNumber="17" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
<emphasis id="B93A9254FFE05010FE02FADDFD184158" bold="true" box="[438,733,1297,1323]" pageId="16" pageNumber="17">
<typeStatus id="54F5F0E4FFE05010FE02FADDFDE64158" box="[438,547,1297,1323]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
(
<collectingCountry id="F3590ED6FFE05010FD81FADDFD154158" box="[565,720,1297,1323]" name="Madagascar" pageId="16" pageNumber="17">Madagascar</collectingCountry>
):
</emphasis>
<specimenCount id="9D4885CFFFE05010FD5CFADEFD3A415F" box="[744,767,1298,1324]" count="1" pageId="16" pageNumber="17" type="male"></specimenCount>
, four labels: “S-Madagaskar /
<location id="8E91189DFFE05010FBDDFADEFAF94158" LSID="urn:lsid:plazi:treatment:03E7FF50FFE05012FF23FBAEFA844586:8E91189DFFE05010FBDDFADEFAF94158" box="[1129,1340,1297,1324]" country="Madagascar" name="Foret de Sakahara" pageId="16" pageNumber="17">Foret de Sakahara</location>
{sic!} / lg.
<collectorName id="26BB2B90FFE05010FF05FAF9FECA413C" box="[177,271,1333,1359]" pageId="16" pageNumber="17">H.Franz</collectorName>
1969” [white, printed], “
<taxonomicName id="4C4E35C5FFE05010FD9AFAF9FD20413C" box="[558,741,1333,1359]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/ sakarahanus m. / det.
<determinerName id="7E2DA282FFE05010FC5BFAF9FB89413C" box="[1007,1100,1333,1359]" pageId="16" pageNumber="17">H.Franz</determinerName>
” [yellowish, handwritten and printed], “Typus” [red, handwritten], “
<specimenCount id="9D4885CFFFE05010FDF2FA95FD994100" box="[582,604,1369,1395]" count="1" pageId="16" pageNumber="17" type="male"></specimenCount>
” [white, printed] (
<collectionCode id="ED5FD683FFE05010FC98FA95FC4A4100" box="[812,911,1369,1395]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="16" pageNumber="17" type="Museum">NHMW</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BF14E46FFE05010FF73FAB1FEE24238" blockId="16.[151,1437,1297,1972]" pageId="16" pageNumber="17">
<emphasis id="B93A9254FFE05010FF73FAB1FE6441E4" bold="true" box="[199,417,1405,1431]" pageId="16" pageNumber="17">Revised diagnosis.</emphasis>
Body (
<figureCitation id="137552C3FFE05010FE40FAB1FD8341E4" box="[500,582,1405,1431]" captionStart="FIGURES 4451" captionStartId="17.[151,264,1523,1547]" captionTargetBox="[151,1422,181,1488]" captionTargetId="figure-16@17.[151,1435,181,1500]" captionTargetPageId="17" captionText="FIGURES 4451. Anhoraeomorphus sakarahanus (Franz), holotype. Habitus in dorsal view (44); head in anterodorsal view (45); head and prothorax in lateral view (only bristles on head are shown) (46); antenna (47); aedeagus in ventral (48, 50) and lateral (49, 51) views." figureDoi="http://doi.org/10.5281/zenodo.13745937" httpUri="https://zenodo.org/record/13745937/files/figure.png" pageId="16" pageNumber="17">Fig. 44</figureCitation>
) stout; head (
<figureCitation id="137552C3FFE05010FD6BFAB1FCF441EB" box="[735,817,1405,1432]" captionStart="FIGURES 4451" captionStartId="17.[151,264,1523,1547]" captionTargetBox="[151,1422,181,1488]" captionTargetId="figure-16@17.[151,1435,181,1500]" captionTargetPageId="17" captionText="FIGURES 4451. Anhoraeomorphus sakarahanus (Franz), holotype. Habitus in dorsal view (44); head in anterodorsal view (45); head and prothorax in lateral view (only bristles on head are shown) (46); antenna (47); aedeagus in ventral (48, 50) and lateral (49, 51) views." figureDoi="http://doi.org/10.5281/zenodo.13745937" httpUri="https://zenodo.org/record/13745937/files/figure.png" pageId="16" pageNumber="17">Fig. 45</figureCitation>
) about as long as wide, with vertex posteriorly broadly rounded; pronotum (
<figureCitation id="137552C3FFE05010FECEFA6DFE0C41C8" box="[378,457,1441,1467]" captionStart="FIGURES 4451" captionStartId="17.[151,264,1523,1547]" captionTargetBox="[151,1422,181,1488]" captionTargetId="figure-16@17.[151,1435,181,1500]" captionTargetPageId="17" captionText="FIGURES 4451. Anhoraeomorphus sakarahanus (Franz), holotype. Habitus in dorsal view (44); head in anterodorsal view (45); head and prothorax in lateral view (only bristles on head are shown) (46); antenna (47); aedeagus in ventral (48, 50) and lateral (49, 51) views." figureDoi="http://doi.org/10.5281/zenodo.13745937" httpUri="https://zenodo.org/record/13745937/files/figure.png" pageId="16" pageNumber="17">Fig. 44</figureCitation>
) bell-shaped, broadest near middle, with one pair of distinct antebasal pits; antennomeres 310 not elongate; all femora similarly slender; metatibiae in males unmodified; aedeagus (
<figureCitation id="137552C3FFE05010FB15FA09FAE74193" box="[1185,1314,1477,1504]" captionStart="FIGURES 4451" captionStartId="17.[151,264,1523,1547]" captionTargetBox="[151,1422,181,1488]" captionTargetId="figure-16@17.[151,1435,181,1500]" captionTargetPageId="17" captionText="FIGURES 4451. Anhoraeomorphus sakarahanus (Franz), holotype. Habitus in dorsal view (44); head in anterodorsal view (45); head and prothorax in lateral view (only bristles on head are shown) (46); antenna (47); aedeagus in ventral (48, 50) and lateral (49, 51) views." figureDoi="http://doi.org/10.5281/zenodo.13745937" httpUri="https://zenodo.org/record/13745937/files/figure.png" pageId="16" pageNumber="17">Figs 4851</figureCitation>
) in ventral view broadest near middle, with apical region of dorsal wall abruptly bent dorsally, distolateral endophallic sclerites widely separated and with rounded apices, each paramere with a row of subapical setae distributed on less than distal fourth.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFE05012FF73F999FE5B45FA" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17" type="description">
<paragraph id="8BF14E46FFE05010FF73F999FF0F42E7" blockId="16.[151,1437,1297,1972]" pageId="16" pageNumber="17">
<emphasis id="B93A9254FFE05010FF73F999FEB0421C" bold="true" box="[199,373,1621,1647]" pageId="16" pageNumber="17">Redescription.</emphasis>
Body of male (
<figureCitation id="137552C3FFE05010FD93F999FDBF421C" box="[551,634,1621,1647]" captionStart="FIGURES 4451" captionStartId="17.[151,264,1523,1547]" captionTargetBox="[151,1422,181,1488]" captionTargetId="figure-16@17.[151,1435,181,1500]" captionTargetPageId="17" captionText="FIGURES 4451. Anhoraeomorphus sakarahanus (Franz), holotype. Habitus in dorsal view (44); head in anterodorsal view (45); head and prothorax in lateral view (only bristles on head are shown) (46); antenna (47); aedeagus in ventral (48, 50) and lateral (49, 51) views." figureDoi="http://doi.org/10.5281/zenodo.13745937" httpUri="https://zenodo.org/record/13745937/files/figure.png" pageId="16" pageNumber="17">Fig. 44</figureCitation>
) light brown, appendages indistinctly lighter, setae yellowish; BL
<quantity id="4CB6E3A3FFE05010FADFF999FF0342E7" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="16" pageNumber="17" unit="mm" value="1.4">1.40 mm</quantity>
.
</paragraph>
<paragraph id="8BF14E46FFE05010FF73F951FC8F43C0" blockId="16.[151,1437,1297,1972]" pageId="16" pageNumber="17">
Head (
<figureCitation id="137552C3FFE05010FEA4F951FE4A42CB" box="[272,399,1693,1720]" captionStart="FIGURES 4451" captionStartId="17.[151,264,1523,1547]" captionTargetBox="[151,1422,181,1488]" captionTargetId="figure-16@17.[151,1435,181,1500]" captionTargetPageId="17" captionText="FIGURES 4451. Anhoraeomorphus sakarahanus (Franz), holotype. Habitus in dorsal view (44); head in anterodorsal view (45); head and prothorax in lateral view (only bristles on head are shown) (46); antenna (47); aedeagus in ventral (48, 50) and lateral (49, 51) views." figureDoi="http://doi.org/10.5281/zenodo.13745937" httpUri="https://zenodo.org/record/13745937/files/figure.png" pageId="16" pageNumber="17">Figs 4546</figureCitation>
) in anterodorsal view rounded, broadest at eyes, HL
<quantity id="4CB6E3A3FFE05010FC61F951FBFF42CB" box="[981,1082,1693,1720]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="16" pageNumber="17" unit="mm" value="0.28">0.28 mm</quantity>
, HW
<quantity id="4CB6E3A3FFE05010FBCDF951FB1B42CB" box="[1145,1246,1693,1720]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="16" pageNumber="17" unit="mm" value="0.28">0.28 mm</quantity>
; temple in lateral view (
<figureCitation id="137552C3FFE05010FF6AF90DFEF542A8" box="[222,304,1729,1755]" captionStart="FIGURES 4451" captionStartId="17.[151,264,1523,1547]" captionTargetBox="[151,1422,181,1488]" captionTargetId="figure-16@17.[151,1435,181,1500]" captionTargetPageId="17" captionText="FIGURES 4451. Anhoraeomorphus sakarahanus (Franz), holotype. Habitus in dorsal view (44); head in anterodorsal view (45); head and prothorax in lateral view (only bristles on head are shown) (46); antenna (47); aedeagus in ventral (48, 50) and lateral (49, 51) views." figureDoi="http://doi.org/10.5281/zenodo.13745937" httpUri="https://zenodo.org/record/13745937/files/figure.png" pageId="16" pageNumber="17">Fig. 46</figureCitation>
) subequal to length of eye; vertex and frons confluent, together weakly convex and about as long as broad; vertex posteriorly strongly rounded; each eye moderately large and weakly convex, with deep posteromedian emargination, weakly projecting from head silhouette. Punctures on vertex fine, inconspicuous; setae short, sparse and suberect; posterior region of frons and entire vertex densely covered with thick bristles directed posteriorly. Antenna (
<figureCitation id="137552C3FFE05010FEB1F89DFE904318" box="[261,341,1873,1899]" captionStart="FIGURES 4451" captionStartId="17.[151,264,1523,1547]" captionTargetBox="[151,1422,181,1488]" captionTargetId="figure-16@17.[151,1435,181,1500]" captionTargetPageId="17" captionText="FIGURES 4451. Anhoraeomorphus sakarahanus (Franz), holotype. Habitus in dorsal view (44); head in anterodorsal view (45); head and prothorax in lateral view (only bristles on head are shown) (46); antenna (47); aedeagus in ventral (48, 50) and lateral (49, 51) views." figureDoi="http://doi.org/10.5281/zenodo.13745937" httpUri="https://zenodo.org/record/13745937/files/figure.png" pageId="16" pageNumber="17">Fig. 47</figureCitation>
) slender but short, slightly shorter than half BL, AnL
<quantity id="4CB6E3A3FFE05010FC1DF89DFBCB431F" box="[937,1038,1873,1900]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="16" pageNumber="17" unit="mm" value="0.55">0.55 mm</quantity>
, antennomeres 1 and 2 each weakly elongate, 3 slightly transverse, 47 each about as long as broad, 810 each transverse; 11 slightly shorter than 9 and 10 combined, as wide as 10, about 1.8 times as long as broad.
</paragraph>
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<paragraph id="8BF14E46FFE15011FF23FA3FFEB34227" blockId="17.[151,1437,1523,1620]" pageId="17" pageNumber="18">
<emphasis id="B93A9254FFE15011FF23FA3FFE93427F" bold="true" box="[151,342,1523,1548]" pageId="17" pageNumber="18">FIGURES 4451.</emphasis>
<taxonomicName id="4C4E35C5FFE15011FEE9FA38FD32427F" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[349,759,1524,1548]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="sakarahanus">
<emphasis id="B93A9254FFE15011FEE9FA38FD67427F" box="[349,674,1524,1548]" italics="true" pageId="17" pageNumber="18">Anhoraeomorphus sakarahanus</emphasis>
(Franz)
</taxonomicName>
, holotype. Habitus in dorsal view (
<emphasis id="B93A9254FFE15011FBD7FA38FBB9427F" bold="true" box="[1123,1148,1524,1548]" pageId="17" pageNumber="18">44</emphasis>
); head in anterodorsal view (
<emphasis id="B93A9254FFE15011FF14F9D4FF7C4243" bold="true" box="[160,185,1560,1584]" pageId="17" pageNumber="18">45</emphasis>
); head and prothorax in lateral view (only bristles on head are shown) (
<emphasis id="B93A9254FFE15011FC23F9D4FC754243" bold="true" box="[919,944,1560,1584]" pageId="17" pageNumber="18">46</emphasis>
); antenna (
<emphasis id="B93A9254FFE15011FB91F9D4FBFB4243" bold="true" box="[1061,1086,1560,1584]" pageId="17" pageNumber="18">47</emphasis>
); aedeagus in ventral (
<emphasis id="B93A9254FFE15011FA9CF9D4FAAD4243" bold="true" box="[1320,1384,1560,1584]" pageId="17" pageNumber="18">48, 50</emphasis>
) and lateral (
<emphasis id="B93A9254FFE15011FF51F9F0FEE14227" bold="true" box="[229,292,1596,1620]" pageId="17" pageNumber="18">49, 51</emphasis>
) views.
</paragraph>
</caption>
<paragraph id="8BF14E46FFE15011FF73F96AFCE84323" blockId="17.[151,1437,1702,2016]" pageId="17" pageNumber="18">
Pronotum (
<figureCitation id="137552C3FFE15011FEFCF96AFE5942B3" box="[328,412,1702,1728]" captionStart="FIGURES 4451" captionStartId="17.[151,264,1523,1547]" captionTargetBox="[151,1422,181,1488]" captionTargetId="figure-16@17.[151,1435,181,1500]" captionTargetPageId="17" captionText="FIGURES 4451. Anhoraeomorphus sakarahanus (Franz), holotype. Habitus in dorsal view (44); head in anterodorsal view (45); head and prothorax in lateral view (only bristles on head are shown) (46); antenna (47); aedeagus in ventral (48, 50) and lateral (49, 51) views." figureDoi="http://doi.org/10.5281/zenodo.13745937" httpUri="https://zenodo.org/record/13745937/files/figure.png" pageId="17" pageNumber="18">Fig. 44</figureCitation>
) bell-shaped, broadest near middle; PL
<quantity id="4CB6E3A3FFE15011FCDFF96AFC1642B3" box="[875,979,1702,1728]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="17" pageNumber="18" unit="mm" value="0.38">0.38 mm</quantity>
, PW
<quantity id="4CB6E3A3FFE15011FBA1F96AFBBA42B3" box="[1045,1151,1702,1729]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="17" pageNumber="18" unit="mm" value="0.35">0.35 mm</quantity>
. Anterior margin weakly arcuate, lateral margins rounded, anteriorly converging stronger than posteriorly; posterior corners blunt; posterior margin indistinctly convex. Base with one pair of round and deep pits, and with barely discernible, extremely short vestiges of sublateral carinae. Disc with fine, inconspicuous punctures; setae sparse, short and suberect, lateral and lateroventral surface of pronotum with dense thick bristles.
</paragraph>
<paragraph id="8BF14E46FFE15011FF73F896FE3B43CF" blockId="17.[151,1437,1702,2016]" pageId="17" pageNumber="18">
Elytra (
<figureCitation id="137552C3FFE15011FEAAF896FEB64307" box="[286,371,1882,1908]" captionStart="FIGURES 4451" captionStartId="17.[151,264,1523,1547]" captionTargetBox="[151,1422,181,1488]" captionTargetId="figure-16@17.[151,1435,181,1500]" captionTargetPageId="17" captionText="FIGURES 4451. Anhoraeomorphus sakarahanus (Franz), holotype. Habitus in dorsal view (44); head in anterodorsal view (45); head and prothorax in lateral view (only bristles on head are shown) (46); antenna (47); aedeagus in ventral (48, 50) and lateral (49, 51) views." figureDoi="http://doi.org/10.5281/zenodo.13745937" httpUri="https://zenodo.org/record/13745937/files/figure.png" pageId="17" pageNumber="18">Fig. 44</figureCitation>
) together oval, broadest distinctly in front of middle; EL
<quantity id="4CB6E3A3FFE15011FBACF896FB474307" box="[1048,1154,1882,1909]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="17" pageNumber="18" unit="mm" value="0.75">0.75 mm</quantity>
, EW
<quantity id="4CB6E3A3FFE15011FB72F896FAF54307" box="[1222,1328,1882,1909]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="17" pageNumber="18" unit="mm" value="0.58">0.58 mm</quantity>
, EI 1.30; humeral calli weakly elevated, basal elytral foveae vestigial, barely discernible. Punctures fine and inconspicuous; setae sparse, short and suberect.
</paragraph>
<paragraph id="8BF14E46FFE15011FF73F80AFBCC4393" blockId="17.[151,1437,1702,2016]" box="[199,1033,1990,2016]" pageId="17" pageNumber="18">Legs moderately long, slender, unmodified except slightly recurved tibiae.</paragraph>
<paragraph id="8BF14E46FFE25012FF73FF5BFE2F4516" blockId="18.[151,1437,150,501]" pageId="18" pageNumber="19">
Aedeagus (
<figureCitation id="137552C3FFE25012FEF3FF5BFE0F44C2" box="[327,458,150,177]" captionStart="FIGURES 4451" captionStartId="17.[151,264,1523,1547]" captionTargetBox="[151,1422,181,1488]" captionTargetId="figure-16@17.[151,1435,181,1500]" captionTargetPageId="17" captionText="FIGURES 4451. Anhoraeomorphus sakarahanus (Franz), holotype. Habitus in dorsal view (44); head in anterodorsal view (45); head and prothorax in lateral view (only bristles on head are shown) (46); antenna (47); aedeagus in ventral (48, 50) and lateral (49, 51) views." figureDoi="http://doi.org/10.5281/zenodo.13745937" httpUri="https://zenodo.org/record/13745937/files/figure.png" pageId="18" pageNumber="19">Figs 4851</figureCitation>
) stout, AeL
<quantity id="4CB6E3A3FFE25012FDE8FF5BFD0344C2" box="[604,710,151,177]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="18" pageNumber="19" unit="mm" value="0.23">0.23 mm</quantity>
; in ventral view broadest near middle and distinctly narrowing towards truncate apex; in lateral view apical region of dorsal wall strongly and abruptly bent dorsally; endophallus with symmetrical and elongate distolateral sclerites and basilateral sclerites, and with asymmetrical weakly sclerotized median structures, distolateral sclerites broadly separated and each with rounded apex; parameres slender and their apices only slightly projecting beyond aedeagal apex, each paramere with a row of sparse and short setae occupying slightly less than distal fourth.
</paragraph>
<paragraph id="8BF14E46FFE25012FF73FEA3FE5B45FA" blockId="18.[151,1437,150,501]" box="[199,414,367,393]" pageId="18" pageNumber="19">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFE25012FF73FE5EFC6345DE" box="[199,934,402,429]" pageId="18" pageNumber="19" type="distribution">
<paragraph id="8BF14E46FFE25012FF73FE5EFC6345DE" blockId="18.[151,1437,150,501]" box="[199,934,402,429]" pageId="18" pageNumber="19">
<emphasis id="B93A9254FFE25012FF73FE5EFEA445DF" bold="true" box="[199,353,402,428]" pageId="18" pageNumber="19">Distribution.</emphasis>
Southern
<collectingCountry id="F3590ED6FFE25012FE60FE5FFD9E45DE" box="[468,603,403,429]" name="Madagascar" pageId="18" pageNumber="19">Madagascar</collectingCountry>
(Atsimo-Andrefana Region).
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFE25012FF73FE7BFA844586" pageId="18" pageNumber="19" type="discussion">
<paragraph id="8BF14E46FFE25012FF73FE7BFC3C45A2" blockId="18.[151,1437,150,501]" box="[199,1017,439,465]" pageId="18" pageNumber="19">
<emphasis id="B93A9254FFE25012FF73FE7BFEFE45A2" bold="true" box="[199,315,439,465]" pageId="18" pageNumber="19">Remarks.</emphasis>
Franz misspelled Sakaraha in the locality label as “Sakahara”.
</paragraph>
<paragraph id="8BF14E46FFE25012FF73FE17FA844586" blockId="18.[151,1437,150,501]" box="[199,1345,475,501]" pageId="18" pageNumber="19">
<taxonomicName id="4C4E35C5FFE25012FF73FE17FDEB4586" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[199,558,475,501]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="sakarahanus">
<emphasis id="B93A9254FFE25012FF73FE17FDEB4586" box="[199,558,475,501]" italics="true" pageId="18" pageNumber="19">Anhoraeomorphus sakarahanus</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C4E35C5FFE25012FD49FE17FC714586" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[765,948,475,501]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="ankaratranus">
<emphasis id="B93A9254FFE25012FD49FE17FC714586" box="[765,948,475,501]" italics="true" pageId="18" pageNumber="19">A. ankaratranus</emphasis>
</taxonomicName>
; see Remarks for the latter species.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,241 @@
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<subSubSection id="C3541DCDFFE75017FF23FACAFCDF4152" box="[151,794,1286,1313]" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="8BF14E46FFE75017FF23FACAFCDF4152" blockId="23.[151,794,1286,1349]" box="[151,794,1286,1313]" pageId="23" pageNumber="24">
<heading id="D0B9F92AFFE75017FF23FACAFCDF4152" bold="true" box="[151,794,1286,1313]" fontSize="11" level="2" pageId="23" pageNumber="24" reason="4">
<emphasis id="B93A9254FFE75017FF23FACAFCDF4152" bold="true" box="[151,794,1286,1313]" pageId="23" pageNumber="24">
<taxonomicName id="4C4E35C5FFE75017FF23FACAFD674153" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[151,674,1286,1313]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="ambodivoangyi" status="comb. nov.">
<emphasis id="B93A9254FFE75017FF23FACAFDFE4152" bold="true" box="[151,571,1286,1313]" italics="true" pageId="23" pageNumber="24">Anhoraeomorphus ambodivoangyi</emphasis>
(Franz)
</taxonomicName>
,
<taxonomicNameLabel id="A2092F2FFFE75017FD1BFACBFCDF4152" box="[687,794,1287,1313]" pageId="23" pageNumber="24" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFE75017FF23FAE6FEE24136" box="[151,295,1322,1349]" pageId="23" pageNumber="24" type="description">
<paragraph id="8BF14E46FFE75017FF23FAE6FEE24136" blockId="23.[151,794,1286,1349]" box="[151,295,1322,1349]" pageId="23" pageNumber="24">
(
<figureCitation id="137552C3FFE75017FF2BFAE6FEDA4136" box="[159,287,1322,1349]" captionStart="FIGURES 8390" captionStartId="24.[151,264,1732,1756]" captionTargetBox="[151,1422,381,1694]" captionTargetId="figure-114@24.[151,1435,381,1709]" captionTargetPageId="24" captionText="FIGURES 8390. Anhoraeomorphus ambodivoangyi (Franz), paratype female (83, 86) and paratype male (8485, 8790). Habitus in dorsal view (83); head in anterodorsal view (84); head and prothorax in lateral view (only bristles on head are shown) (85); antenna (86); aedeagus in ventral (87, 89) and lateral (88, 90) views." figureDoi="http://doi.org/10.5281/zenodo.13745953" httpUri="https://zenodo.org/record/13745953/files/figure.png" pageId="23" pageNumber="24">Figs 8390</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFE75017FF23FAA3FD6341F4" box="[151,678,1391,1415]" pageId="23" pageNumber="24" type="reference_group">
<paragraph id="8BF14E46FFE75017FF23FAA3FD6341F4" blockId="23.[151,678,1391,1415]" box="[151,678,1391,1415]" pageId="23" pageNumber="24">
<treatmentCitationGroup id="AB5E6968FFE75017FF23FAA3FD6341F4" box="[151,678,1391,1415]" pageId="23" pageNumber="24">
<treatmentCitation id="0AEF6857FFE75017FF23FAA3FD6741F4" author="Franz, H." box="[151,674,1391,1415]" page="149" pageId="23" pageNumber="24" year="1986">
<taxonomicName id="4C4E35C5FFE75017FF23FAA3FD6741F4" ID-CoL="3MNHX" authority="Franz, 1986 b: 149" authorityName="Franz" authorityPageNumber="149" authorityYear="1986" box="[151,674,1391,1415]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="ambodivoangyi">
<emphasis id="B93A9254FFE75017FF23FAA3FE2541F4" box="[151,480,1391,1415]" italics="true" pageId="23" pageNumber="24">Horaeomorphus ambodivoangyi</emphasis>
<bibRefCitation id="EFDF33B7FFE75017FE52FAA3FD6741F4" author="Franz, H." box="[486,674,1391,1415]" pageId="23" pageNumber="24" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 149</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFE75017FF23FA79FBB24367" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="8BF14E46FFE75017FF23FA79FBD84264" blockId="23.[151,1437,1461,2028]" pageId="23" pageNumber="24">
<emphasis id="B93A9254FFE75017FF23FA79FE6B41BC" bold="true" box="[151,430,1461,1487]" pageId="23" pageNumber="24">Type material studied.</emphasis>
<materialsCitation id="3B26441BFFE75017FE0FFA79FBDF4264" collectionCode="NHMW" collectorName="J. Vadon" country="Madagascar" determinerName="H. Franz" location="MADAGASCAR" pageId="23" pageNumber="24" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B93A9254FFE75017FE0FFA79FD2341BC" bold="true" box="[443,742,1461,1487]" pageId="23" pageNumber="24">
<typeStatus id="54F5F0E4FFE75017FE0FFA79FDF641BC" box="[443,563,1461,1487]" pageId="23" pageNumber="24" type="paratype">Paratypes</typeStatus>
(Madagascar)
</emphasis>
:
<specimenCount id="9D4885CFFFE75017FD4EFA79FCEE41A3" box="[762,811,1461,1488]" count="1" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D4885CFFFE75017FC89FA79FCA941A3" box="[829,876,1461,1488]" count="1" pageId="23" pageNumber="24" type="female">1 ♀</specimenCount>
, three labels: “Maroantsetra / Ambodivoangy /
<collectorName id="26BB2B90FFE75017FF23FA15FF364180" box="[151,243,1497,1523]" pageId="23" pageNumber="24">J.Vadon</collectorName>
” [white, printed], “
<taxonomicName id="4C4E35C5FFE75017FE77FA15FDBF4180" box="[451,634,1497,1523]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/ ambodivoangyi m. / det.
<determinerName id="7E2DA282FFE75017FC22FA15FC314180" box="[918,1012,1497,1523]" pageId="23" pageNumber="24">H.Franz</determinerName>
” [yellowish, handwritten and printed], “INSTITUT / SCIENTIFIQUE /
<collectingCountry id="F3590ED6FFE75017FDBEFA31FD144264" box="[522,721,1533,1559]" name="Madagascar" pageId="23" pageNumber="24">MADAGASCAR</collectingCountry>
” [bluish, printed], (
<collectionCode id="ED5FD683FFE75017FC1BFA31FBD74264" box="[943,1042,1533,1559]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="23" pageNumber="24" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF14E46FFE75017FF73F9EDFBB24367" blockId="23.[151,1437,1461,2028]" pageId="23" pageNumber="24">
<emphasis id="B93A9254FFE75017FF73F9EDFE604248" bold="true" box="[199,421,1569,1595]" pageId="23" pageNumber="24">Revised diagnosis.</emphasis>
Body (
<figureCitation id="137552C3FFE75017FE4BF9EDFD904248" box="[511,597,1569,1595]" captionStart="FIGURES 8390" captionStartId="24.[151,264,1732,1756]" captionTargetBox="[151,1422,381,1694]" captionTargetId="figure-114@24.[151,1435,381,1709]" captionTargetPageId="24" captionText="FIGURES 8390. Anhoraeomorphus ambodivoangyi (Franz), paratype female (83, 86) and paratype male (8485, 8790). Habitus in dorsal view (83); head in anterodorsal view (84); head and prothorax in lateral view (only bristles on head are shown) (85); antenna (86); aedeagus in ventral (87, 89) and lateral (88, 90) views." figureDoi="http://doi.org/10.5281/zenodo.13745953" httpUri="https://zenodo.org/record/13745953/files/figure.png" pageId="23" pageNumber="24">Fig. 83</figureCitation>
) stout; head (
<figureCitation id="137552C3FFE75017FD4DF9EDFC8B4248" box="[761,846,1569,1595]" captionStart="FIGURES 8390" captionStartId="24.[151,264,1732,1756]" captionTargetBox="[151,1422,381,1694]" captionTargetId="figure-114@24.[151,1435,381,1709]" captionTargetPageId="24" captionText="FIGURES 8390. Anhoraeomorphus ambodivoangyi (Franz), paratype female (83, 86) and paratype male (8485, 8790). Habitus in dorsal view (83); head in anterodorsal view (84); head and prothorax in lateral view (only bristles on head are shown) (85); antenna (86); aedeagus in ventral (87, 89) and lateral (88, 90) views." figureDoi="http://doi.org/10.5281/zenodo.13745953" httpUri="https://zenodo.org/record/13745953/files/figure.png" pageId="23" pageNumber="24">Fig. 84</figureCitation>
) clearly elongate, with vertex posteriorly truncate; pronotum (
<figureCitation id="137552C3FFE75017FEA2F989FEAF422C" box="[278,362,1605,1631]" captionStart="FIGURES 8390" captionStartId="24.[151,264,1732,1756]" captionTargetBox="[151,1422,381,1694]" captionTargetId="figure-114@24.[151,1435,381,1709]" captionTargetPageId="24" captionText="FIGURES 8390. Anhoraeomorphus ambodivoangyi (Franz), paratype female (83, 86) and paratype male (8485, 8790). Habitus in dorsal view (83); head in anterodorsal view (84); head and prothorax in lateral view (only bristles on head are shown) (85); antenna (86); aedeagus in ventral (87, 89) and lateral (88, 90) views." figureDoi="http://doi.org/10.5281/zenodo.13745953" httpUri="https://zenodo.org/record/13745953/files/figure.png" pageId="23" pageNumber="24">Fig. 83</figureCitation>
) rounded, broadest in anterior third, but weakly narrowing posteriorly, with one pair of large antebasal pits; antennomeres 36 (
<figureCitation id="137552C3FFE75017FDAAF9A5FDB542F0" box="[542,624,1641,1667]" captionStart="FIGURES 8390" captionStartId="24.[151,264,1732,1756]" captionTargetBox="[151,1422,381,1694]" captionTargetId="figure-114@24.[151,1435,381,1709]" captionTargetPageId="24" captionText="FIGURES 8390. Anhoraeomorphus ambodivoangyi (Franz), paratype female (83, 86) and paratype male (8485, 8790). Habitus in dorsal view (83); head in anterodorsal view (84); head and prothorax in lateral view (only bristles on head are shown) (85); antenna (86); aedeagus in ventral (87, 89) and lateral (88, 90) views." figureDoi="http://doi.org/10.5281/zenodo.13745953" httpUri="https://zenodo.org/record/13745953/files/figure.png" pageId="23" pageNumber="24">Fig. 86</figureCitation>
) elongate; all femora similarly slender; hind tibiae in male unmodified; aedeagus (
<figureCitation id="137552C3FFE75017FEBAF941FE5542D4" box="[270,400,1677,1703]" captionStart="FIGURES 8390" captionStartId="24.[151,264,1732,1756]" captionTargetBox="[151,1422,381,1694]" captionTargetId="figure-114@24.[151,1435,381,1709]" captionTargetPageId="24" captionText="FIGURES 8390. Anhoraeomorphus ambodivoangyi (Franz), paratype female (83, 86) and paratype male (8485, 8790). Habitus in dorsal view (83); head in anterodorsal view (84); head and prothorax in lateral view (only bristles on head are shown) (85); antenna (86); aedeagus in ventral (87, 89) and lateral (88, 90) views." figureDoi="http://doi.org/10.5281/zenodo.13745953" httpUri="https://zenodo.org/record/13745953/files/figure.png" pageId="23" pageNumber="24">Figs 8790</figureCitation>
) in ventral view broadest near base, lateral margins sinuate, apical region of dorsal wall in lateral view weakly curved dorsally but with dorsal surface convex and ventral concave, distolateral endophallic sclerites widely separated, each only about as long as wide, with subtriangular apex, each paramere with a short subapical row of several long setae and proximal to them with two short subapical setae.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFE75019FF73F8D1FC5B464E" lastPageId="25" lastPageNumber="26" pageId="23" pageNumber="24" type="description">
<paragraph id="8BF14E46FFE75017FF73F8D1FEC7432F" blockId="23.[151,1437,1461,2028]" pageId="23" pageNumber="24">
<emphasis id="B93A9254FFE75017FF73F8D1FEB04344" bold="true" box="[199,373,1821,1847]" pageId="23" pageNumber="24">Redescription.</emphasis>
Body of male moderately dark brown, tarsi and palps distinctly lighter, setae yellowish; BL
<quantity id="4CB6E3A3FFE75017FF23F88DFF3B432F" box="[151,254,1857,1884]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="23" pageNumber="24" unit="mm" value="2.2">2.20 mm</quantity>
.
</paragraph>
<paragraph id="8BF14E46FFE75018FF73F8A9FE0C44A6" blockId="23.[151,1437,1461,2028]" lastBlockId="24.[151,1436,151,357]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
Head (
<figureCitation id="137552C3FFE75017FEA6F8A9FE5143F3" box="[274,404,1893,1920]" captionStart="FIGURES 8390" captionStartId="24.[151,264,1732,1756]" captionTargetBox="[151,1422,381,1694]" captionTargetId="figure-114@24.[151,1435,381,1709]" captionTargetPageId="24" captionText="FIGURES 8390. Anhoraeomorphus ambodivoangyi (Franz), paratype female (83, 86) and paratype male (8485, 8790). Habitus in dorsal view (83); head in anterodorsal view (84); head and prothorax in lateral view (only bristles on head are shown) (85); antenna (86); aedeagus in ventral (87, 89) and lateral (88, 90) views." figureDoi="http://doi.org/10.5281/zenodo.13745953" httpUri="https://zenodo.org/record/13745953/files/figure.png" pageId="23" pageNumber="24">Figs 8485</figureCitation>
) in anterodorsal view distinctly elongate, conspicuously narrower than prothorax, broadest at eyes, HL
<quantity id="4CB6E3A3FFE75017FE92F845FE4B43D7" box="[294,398,1929,1956]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="23" pageNumber="24" unit="mm" value="0.48">0.48 mm</quantity>
, HW
<quantity id="4CB6E3A3FFE75017FE60F845FDFB43D7" box="[468,574,1929,1956]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="23" pageNumber="24" unit="mm" value="0.38">0.38 mm</quantity>
; temple in lateral view (
<figureCitation id="137552C3FFE75017FCEEF845FC6B43D7" box="[858,942,1929,1956]" captionStart="FIGURES 8390" captionStartId="24.[151,264,1732,1756]" captionTargetBox="[151,1422,381,1694]" captionTargetId="figure-114@24.[151,1435,381,1709]" captionTargetPageId="24" captionText="FIGURES 8390. Anhoraeomorphus ambodivoangyi (Franz), paratype female (83, 86) and paratype male (8485, 8790). Habitus in dorsal view (83); head in anterodorsal view (84); head and prothorax in lateral view (only bristles on head are shown) (85); antenna (86); aedeagus in ventral (87, 89) and lateral (88, 90) views." figureDoi="http://doi.org/10.5281/zenodo.13745953" httpUri="https://zenodo.org/record/13745953/files/figure.png" pageId="23" pageNumber="24">Fig. 85</figureCitation>
) slightly longer than eye; vertex and frons confluent, together weakly convex and elongate; vertex posteriorly truncate; each eye moderately large and weakly convex, oval, weakly projecting from head silhouette. Punctures on vertex fine, inconspicuous; setae short, dense and suberect; frons and vertex densely covered with thick bristles directed posteriorly. Antennae in the only studied male specimen are lacking.
</paragraph>
<paragraph id="8BF14E46FFE85018FF73FF13FB024516" blockId="24.[151,1436,151,357]" pageId="24" pageNumber="25">
Pronotum round, broadest in anterior third; PL
<quantity id="4CB6E3A3FFE85018FD5AFF12FC9D448A" box="[750,856,222,249]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="24" pageNumber="25" unit="mm" value="0.53">0.53 mm</quantity>
, PW
<quantity id="4CB6E3A3FFE85018FC2EFF12FBC0448A" box="[922,1029,222,249]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="24" pageNumber="25" unit="mm" value="0.5">0.50 mm</quantity>
. Anterior margin strongly rounded, lateral margins rounded in anterior half, nearly straight in posterior half; posterior corners blunt; posterior margin straight. Base with one pair of large round pits, and with short sublateral carinae. Disc with fine, inconspicuous punctures; setae dense, short and suberect, lateral surface of pronotum with dense thick bristles.
</paragraph>
<caption id="DF311ECEFFE85018FF23F908FC404357" ID-DOI="http://doi.org/10.5281/zenodo.13745953" ID-Zenodo-Dep="13745953" httpUri="https://zenodo.org/record/13745953/files/figure.png" pageId="24" pageNumber="25" startId="24.[151,264,1732,1756]" targetBox="[151,1422,381,1694]" targetPageId="24" targetType="figure">
<paragraph id="8BF14E46FFE85018FF23F908FC404357" blockId="24.[151,1436,1732,1829]" pageId="24" pageNumber="25">
<emphasis id="B93A9254FFE85018FF23F908FE9242AF" bold="true" box="[151,343,1732,1756]" pageId="24" pageNumber="25">FIGURES 8390.</emphasis>
<taxonomicName id="4C4E35C5FFE85018FED4F908FCD242AE" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[352,791,1732,1757]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="ambodivoangyi">
<emphasis id="B93A9254FFE85018FED4F908FD0442AF" box="[352,705,1732,1756]" italics="true" pageId="24" pageNumber="25">Anhoraeomorphus ambodivoangyi</emphasis>
(Franz)
</taxonomicName>
, paratype female (
<emphasis id="B93A9254FFE85018FC6FF908FBD842AF" bold="true" box="[987,1053,1732,1756]" pageId="24" pageNumber="25">83, 86</emphasis>
) and paratype male (
<emphasis id="B93A9254FFE85018FB4BF908FA4842AF" bold="true" box="[1279,1421,1732,1756]" pageId="24" pageNumber="25">8485, 8790</emphasis>
). Habitus in dorsal view (
<emphasis id="B93A9254FFE85018FE33F924FE654373" bold="true" box="[391,416,1768,1792]" pageId="24" pageNumber="25">83</emphasis>
); head in anterodorsal view (
<emphasis id="B93A9254FFE85018FD72F924FD1A4373" bold="true" box="[710,735,1768,1792]" pageId="24" pageNumber="25">84</emphasis>
); head and prothorax in lateral view (only bristles on head are shown) (
<emphasis id="B93A9254FFE85018FF14F8C0FF7C4357" bold="true" box="[160,185,1804,1828]" pageId="24" pageNumber="25">85</emphasis>
); antenna (
<emphasis id="B93A9254FFE85018FE98F8C0FE804357" bold="true" box="[300,325,1804,1828]" pageId="24" pageNumber="25">86</emphasis>
); aedeagus in ventral (
<emphasis id="B93A9254FFE85018FD9AF8C0FDA84357" bold="true" box="[558,621,1804,1828]" pageId="24" pageNumber="25">87, 89</emphasis>
) and lateral (
<emphasis id="B93A9254FFE85018FD41F8C0FCF14357" bold="true" box="[757,820,1804,1828]" pageId="24" pageNumber="25">88, 90</emphasis>
) views.
</paragraph>
</caption>
<paragraph id="8BF14E46FFE85018FF73F8BCFCD343A1" blockId="24.[151,1436,1904,2039]" pageId="24" pageNumber="25">
Elytra together oval, broadest distinctly in front of middle; EL
<quantity id="4CB6E3A3FFE85018FC39F8BCFC3143F8" box="[909,1012,1904,1931]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="24" pageNumber="25" unit="mm" value="1.2">1.20 mm</quantity>
, EW
<quantity id="4CB6E3A3FFE85018FB87F8BCFB5F43F8" box="[1075,1178,1904,1931]" metricMagnitude="-4" metricUnit="m" metricValue="9.3" pageId="24" pageNumber="25" unit="mm" value="0.93">0.93 mm</quantity>
, EI 1.30; humeral calli weakly elevated, basal elytral foveae vestigial, barely discernible. Punctures small and shallow, but more distinct than those on pronotum; setae sparse, short and suberect.
</paragraph>
<paragraph id="8BF14E46FFE85018FF73F811FB384385" blockId="24.[151,1436,1904,2039]" box="[199,1277,2012,2039]" pageId="24" pageNumber="25">Legs moderately long, slender, unmodified except slightly recurved tibiae (especially protibiae).</paragraph>
<paragraph id="8BF14E46FFE95019FF73FF5BFB8345FA" blockId="25.[151,1437,150,897]" pageId="25" pageNumber="26">
Aedeagus (
<figureCitation id="137552C3FFE95019FEF0FF5BFE0644C2" box="[324,451,151,177]" captionStart="FIGURES 8390" captionStartId="24.[151,264,1732,1756]" captionTargetBox="[151,1422,381,1694]" captionTargetId="figure-114@24.[151,1435,381,1709]" captionTargetPageId="24" captionText="FIGURES 8390. Anhoraeomorphus ambodivoangyi (Franz), paratype female (83, 86) and paratype male (8485, 8790). Habitus in dorsal view (83); head in anterodorsal view (84); head and prothorax in lateral view (only bristles on head are shown) (85); antenna (86); aedeagus in ventral (87, 89) and lateral (88, 90) views." figureDoi="http://doi.org/10.5281/zenodo.13745953" httpUri="https://zenodo.org/record/13745953/files/figure.png" pageId="25" pageNumber="26">Figs 8790</figureCitation>
) slender, AeL
<quantity id="4CB6E3A3FFE95019FDD0FF5AFD0F44C2" box="[612,714,150,177]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="25" pageNumber="26" unit="mm" value="0.5">0.50 mm</quantity>
; in ventral view broadest near base, distinctly narrowing towards abruptly delimited and broadly subtriangular apex, lateral margins of median lobe sinuate; in lateral view apical region of dorsal wall weakly curved dorsally, with dorsal wall convex and ventral wall concave; endophallus with symmetrical and short distolateral sclerites and with strongly asymmetrical assemblage of median sclerotized structures, distolateral sclerites broadly separated, each about as long as broad, subquadrate with distomesal subtriangular projection; parameres broad and their apices not projecting beyond aedeagal apex, each paramere with a longitudinal row of five long and proximal to them two short subapical setae.
</paragraph>
<paragraph id="8BF14E46FFE95019FF73FE5FFC5B464E" blockId="25.[151,1437,150,897]" pageId="25" pageNumber="26">
Female (
<figureCitation id="137552C3FFE95019FE9EFE5FFEBB45DE" box="[298,382,403,429]" captionStart="FIGURES 8390" captionStartId="24.[151,264,1732,1756]" captionTargetBox="[151,1422,381,1694]" captionTargetId="figure-114@24.[151,1435,381,1709]" captionTargetPageId="24" captionText="FIGURES 8390. Anhoraeomorphus ambodivoangyi (Franz), paratype female (83, 86) and paratype male (8485, 8790). Habitus in dorsal view (83); head in anterodorsal view (84); head and prothorax in lateral view (only bristles on head are shown) (85); antenna (86); aedeagus in ventral (87, 89) and lateral (88, 90) views." figureDoi="http://doi.org/10.5281/zenodo.13745953" httpUri="https://zenodo.org/record/13745953/files/figure.png" pageId="25" pageNumber="26">Fig. 83</figureCitation>
). Externally similar to male, but protibiae only weakly curved mesally, meso- and metatibiae nearly straight. BL
<quantity id="4CB6E3A3FFE95019FECCFE7BFE2745A2" box="[376,482,439,465]" metricMagnitude="-3" metricUnit="m" metricValue="2.18" pageId="25" pageNumber="26" unit="mm" value="2.18">2.18 mm</quantity>
; HL
<quantity id="4CB6E3A3FFE95019FDAAFE7BFD4345A2" box="[542,646,439,465]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="25" pageNumber="26" unit="mm" value="0.48">0.48 mm</quantity>
, HW
<quantity id="4CB6E3A3FFE95019FD78FE7BFCF045A2" box="[716,821,439,465]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="25" pageNumber="26" unit="mm" value="0.38">0.38 mm</quantity>
, AnL
<quantity id="4CB6E3A3FFE95019FCCAFE7BFC2245A2" box="[894,999,439,465]" metricMagnitude="-3" metricUnit="m" metricValue="1.03" pageId="25" pageNumber="26" unit="mm" value="1.03">1.03 mm</quantity>
; PL
<quantity id="4CB6E3A3FFE95019FBABFE7AFB4245A2" box="[1055,1159,438,465]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="25" pageNumber="26" unit="mm" value="0.55">0.55 mm</quantity>
, PW
<quantity id="4CB6E3A3FFE95019FB7DFE7AFAF745A2" box="[1225,1330,438,465]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="25" pageNumber="26" unit="mm" value="0.53">0.53 mm</quantity>
; EL
<quantity id="4CB6E3A3FFE95019FADFFE7AFF004586" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="25" pageNumber="26" unit="mm" value="1.15">1.15 mm</quantity>
, EW
<quantity id="4CB6E3A3FFE95019FEBFFE17FEB34586" box="[267,374,475,501]" metricMagnitude="-4" metricUnit="m" metricValue="9.3" pageId="25" pageNumber="26" unit="mm" value="0.93">0.93 mm</quantity>
, EI 1.24. Antenna (
<figureCitation id="137552C3FFE95019FDD4FE17FD704586" box="[608,693,475,501]" captionStart="FIGURES 8390" captionStartId="24.[151,264,1732,1756]" captionTargetBox="[151,1422,381,1694]" captionTargetId="figure-114@24.[151,1435,381,1709]" captionTargetPageId="24" captionText="FIGURES 8390. Anhoraeomorphus ambodivoangyi (Franz), paratype female (83, 86) and paratype male (8485, 8790). Habitus in dorsal view (83); head in anterodorsal view (84); head and prothorax in lateral view (only bristles on head are shown) (85); antenna (86); aedeagus in ventral (87, 89) and lateral (88, 90) views." figureDoi="http://doi.org/10.5281/zenodo.13745953" httpUri="https://zenodo.org/record/13745953/files/figure.png" pageId="25" pageNumber="26">Fig. 86</figureCitation>
) indistinctly shorter than half BL, strongly broadening distally, antennomeres 16 each elongate (5 longest), 7 about as long as broad, 810 each transverse, 11 much shorter than 9 and 10 combined, slightly narrower than 10, only slightly elongate.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFE95019FF73FD8AFCA64612" box="[199,867,582,609]" pageId="25" pageNumber="26" type="distribution">
<paragraph id="8BF14E46FFE95019FF73FD8AFCA64612" blockId="25.[151,1437,150,897]" box="[199,867,582,609]" pageId="25" pageNumber="26">
<emphasis id="B93A9254FFE95019FF73FD8AFEA44613" bold="true" box="[199,353,582,608]" pageId="25" pageNumber="26">Distribution.</emphasis>
Northern
<collectingCountry id="F3590ED6FFE95019FE60FD8BFD9E4612" box="[468,603,583,609]" name="Madagascar" pageId="25" pageNumber="26">Madagascar</collectingCountry>
(Analanjirofo Region).
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFE95019FF73FDA7FD0947F2" pageId="25" pageNumber="26" type="discussion">
<paragraph id="8BF14E46FFE95019FF73FDA7FF124766" blockId="25.[151,1437,150,897]" pageId="25" pageNumber="26">
<emphasis id="B93A9254FFE95019FF73FDA7FEFE46F6" bold="true" box="[199,315,619,645]" pageId="25" pageNumber="26">Remarks.</emphasis>
<taxonomicName id="4C4E35C5FFE95019FEF6FDA7FD0246F6" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[322,711,619,645]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="ambodivoangyi">
<emphasis id="B93A9254FFE95019FEF6FDA7FD0246F6" box="[322,711,619,645]" italics="true" pageId="25" pageNumber="26">Anhoraeomorphus ambodivoangyi</emphasis>
</taxonomicName>
is unique in having a distinctly elongate head, slender antennae, pronotum about as long as broad, with one pair of pits, and a slender aedeagus with abruptly delimited broadly subtriangular apex, endophallic distolateral sclerites conspicuously short, about as long as broad, and the setal pattern of parameral apices, each bearing a longitudinal row of five long and proximal to them two short subapical setae.
</paragraph>
<paragraph id="8BF14E46FFE95019FF73FCD3FD0947F2" blockId="25.[151,1437,150,897]" pageId="25" pageNumber="26">
<materialsCitation id="3B26441BFFE95019FF73FCD3FD0D47F2" collectionCode="NHMW" pageId="25" pageNumber="26" specimenCount="2" typeStatus="holotype">
The
<typeStatus id="54F5F0E4FFE95019FF4EFCD3FE99474A" box="[250,348,799,825]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
of this species is deposited in Muséum National d'Histoire Naturelle, Paris and was not available for my study.
<specimenCount id="9D4885CFFFE95019FE9AFC8FFE16472E" box="[302,467,835,861]" count="2" pageId="25" pageNumber="26" type="generic" typeStatus="paratype">Two paratypes</specimenCount>
, male and female, are in Franz Coll. (
<collectionCode id="ED5FD683FFE95019FCD8FC8FFC0B472E" box="[876,974,835,861]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="25" pageNumber="26" type="Museum">NHMW</collectionCode>
), and the aedeagus of a dissected
<typeStatus id="54F5F0E4FFE95019FA88FC8FFA59472E" box="[1340,1436,835,861]" pageId="25" pageNumber="26" type="paratype">paratype</typeStatus>
matches the illustration in the original description
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="9E2FFDE5B9B3FC7E64D1CDC1CB32A814" ID-DOI="10.11646/zootaxa.5505.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="13745912" ID-ZooBank="93F5E9F6-9EDC-4ED2-8F2B-1E6D0C18F893" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726052234810" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2024" docId="03E7FF50FFE9501BFF23FC1EFCFB47BA" docLanguage="en" docName="zootaxa.5505.1.1.pdf" docOrigin="Zootaxa 5505 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5505.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Anhoraeomorphus anosyensis Jałoszyński 2024, comb. n." docType="treatment" docVersion="2" lastPageNumber="28" masterDocId="FFDE8728FFF05000FFB4FFCCFFC54473" masterDocTitle="The curse of Horaeomorphus (almost) lifted. Revision of misplaced species from Madagascar, Comoros and Mascarenes (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="96" masterPageNumber="1" pageNumber="26" updateTime="1726053143081" updateUser="ExternalLinkService">
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<mods:title id="D443C93101AB76BE1FEE5DB6BE88D6DA">The curse of Horaeomorphus (almost) lifted. Revision of misplaced species from Madagascar, Comoros and Mascarenes (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:date id="6B74F04C1ABE106CAA8CBE454591D50F">2024</mods:date>
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<subSubSection id="C3541DCDFFE95019FF23FC1EFD20479E" box="[151,741,978,1005]" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="8BF14E46FFE95019FF23FC1EFD20479E" blockId="25.[151,741,978,1041]" box="[151,741,978,1005]" pageId="25" pageNumber="26">
<heading id="D0B9F92AFFE95019FF23FC1EFD20479E" bold="true" box="[151,741,978,1005]" fontSize="11" level="2" pageId="25" pageNumber="26" reason="4">
<emphasis id="B93A9254FFE95019FF23FC1EFD20479E" bold="true" box="[151,741,978,1005]" pageId="25" pageNumber="26">
<taxonomicName id="4C4E35C5FFE95019FF23FC1EFDA9479F" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[151,620,978,1005]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="anosyensis" status="comb. nov.">
<emphasis id="B93A9254FFE95019FF23FC1EFDC0479E" bold="true" box="[151,517,978,1005]" italics="true" pageId="25" pageNumber="26">Anhoraeomorphus anosyensis</emphasis>
(Franz)
</taxonomicName>
,
<taxonomicNameLabel id="A2092F2FFFE95019FDCEFC1FFD20479E" box="[634,741,979,1005]" pageId="25" pageNumber="26" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3541DCDFFE95019FF23FC3BFEE24062" box="[151,295,1015,1041]" pageId="25" pageNumber="26" type="description">
<paragraph id="8BF14E46FFE95019FF23FC3BFEE24062" blockId="25.[151,741,978,1041]" box="[151,295,1015,1041]" pageId="25" pageNumber="26">
(
<figureCitation id="137552C3FFE95019FF2BFC3BFEDA4062" box="[159,287,1015,1041]" captionStart="FIGURES 9198" captionStartId="26.[151,264,1681,1705]" captionTargetBox="[151,1418,181,1636]" captionTargetId="figure-28@26.[151,1435,181,1657]" captionTargetPageId="26" captionText="FIGURES 9198. Anhoraeomorphus anosyensis (Franz), paratype. Habitus in dorsal view (91); head in anterodorsal view (92); head and prothorax in lateral view (only bristles on head are shown) (93); antenna (94); aedeagus in ventral (95, 97) and lateral (96, 98) views." figureDoi="http://doi.org/10.5281/zenodo.13745955" httpUri="https://zenodo.org/record/13745955/files/figure.png" pageId="25" pageNumber="26">Figs 9198</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFE95019FF23FBF7FDBD4020" box="[151,632,1083,1107]" pageId="25" pageNumber="26" type="reference_group">
<paragraph id="8BF14E46FFE95019FF23FBF7FDBD4020" blockId="25.[151,632,1083,1107]" box="[151,632,1083,1107]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="AB5E6968FFE95019FF23FBF7FDBD4020" box="[151,632,1083,1107]" pageId="25" pageNumber="26">
<treatmentCitation id="0AEF6857FFE95019FF23FBF7FDB14020" author="Franz, H." box="[151,628,1083,1107]" page="151" pageId="25" pageNumber="26" year="1986">
<taxonomicName id="4C4E35C5FFE95019FF23FBF7FDB14020" ID-CoL="3MNJ8" authority="Franz, 1986 b: 151" authorityName="Franz" authorityPageNumber="151" authorityYear="1986" box="[151,628,1083,1107]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="anosyensis">
<emphasis id="B93A9254FFE95019FF23FBF7FE774020" box="[151,434,1083,1107]" italics="true" pageId="25" pageNumber="26">Horaeomorphus anosyensis</emphasis>
<bibRefCitation id="EFDF33B7FFE95019FE0CFBF0FDB14020" author="Franz, H." box="[440,628,1083,1107]" pageId="25" pageNumber="26" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 151</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFE95019FF23FB4DFDBF4193" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="8BF14E46FFE95019FF23FB4DFEA84090" blockId="25.[151,1437,1153,1972]" pageId="25" pageNumber="26">
<emphasis id="B93A9254FFE95019FF23FB4DFE6240E8" bold="true" box="[151,423,1153,1179]" pageId="25" pageNumber="26">Type material studied.</emphasis>
<materialsCitation id="3B26441BFFE95019FE04FB4DFEAC4090" collectionCode="NHMW" collectorName="Foret N Ft. Dauphin" country="Madagascar" determinerName="H. Franz" location="Madagascar" pageId="25" pageNumber="26" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B93A9254FFE95019FE04FB4DFD0840E8" bold="true" box="[432,717,1153,1179]" pageId="25" pageNumber="26">
<typeStatus id="54F5F0E4FFE95019FE04FB4DFDD840E8" box="[432,541,1153,1179]" pageId="25" pageNumber="26" type="paratype">Paratype</typeStatus>
(
<collectingCountry id="F3590ED6FFE95019FD9AFB4DFD0040E8" box="[558,709,1153,1179]" name="Madagascar" pageId="25" pageNumber="26">Madagascar</collectingCountry>
)
</emphasis>
:
<specimenCount id="9D4885CFFFE95019FD69FB4EFD3140EF" box="[733,756,1154,1180]" count="1" pageId="25" pageNumber="26" type="male"></specimenCount>
, two labels: “Madagascar (Süd) /
<collectorName id="26BB2B90FFE95019FBC8FB4DFAA040E8" box="[1148,1381,1153,1180]" pageId="25" pageNumber="26">Forét N Ft. Dauphin</collectorName>
/ lg. Alluaud 1900” [white, printed], “
<taxonomicName id="4C4E35C5FFE95019FDBDFB69FD0540CC" box="[521,704,1189,1215]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/ anosiensis {sic!} m. / det.
<determinerName id="7E2DA282FFE95019FC40FB69FB9740CC" box="[1012,1106,1189,1215]" pageId="25" pageNumber="26">H.Franz</determinerName>
” [yellowish, handwritten and printed] (
<collectionCode id="ED5FD683FFE95019FF4BFB05FEA74090" box="[255,354,1225,1251]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="25" pageNumber="26" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF14E46FFE95019FF73FB21FDBF4193" blockId="25.[151,1437,1153,1972]" pageId="25" pageNumber="26">
<emphasis id="B93A9254FFE95019FF73FB21FE644174" bold="true" box="[199,417,1261,1287]" pageId="25" pageNumber="26">Revised diagnosis.</emphasis>
Body (
<figureCitation id="137552C3FFE95019FE40FB21FD804174" box="[500,581,1261,1287]" captionStart="FIGURES 9198" captionStartId="26.[151,264,1681,1705]" captionTargetBox="[151,1418,181,1636]" captionTargetId="figure-28@26.[151,1435,181,1657]" captionTargetPageId="26" captionText="FIGURES 9198. Anhoraeomorphus anosyensis (Franz), paratype. Habitus in dorsal view (91); head in anterodorsal view (92); head and prothorax in lateral view (only bristles on head are shown) (93); antenna (94); aedeagus in ventral (95, 97) and lateral (96, 98) views." figureDoi="http://doi.org/10.5281/zenodo.13745955" httpUri="https://zenodo.org/record/13745955/files/figure.png" pageId="25" pageNumber="26">Fig. 91</figureCitation>
) slender; head (
<figureCitation id="137552C3FFE95019FD4CFB21FCBD4174" box="[760,888,1261,1287]" captionStart="FIGURES 9198" captionStartId="26.[151,264,1681,1705]" captionTargetBox="[151,1418,181,1636]" captionTargetId="figure-28@26.[151,1435,181,1657]" captionTargetPageId="26" captionText="FIGURES 9198. Anhoraeomorphus anosyensis (Franz), paratype. Habitus in dorsal view (91); head in anterodorsal view (92); head and prothorax in lateral view (only bristles on head are shown) (93); antenna (94); aedeagus in ventral (95, 97) and lateral (96, 98) views." figureDoi="http://doi.org/10.5281/zenodo.13745955" httpUri="https://zenodo.org/record/13745955/files/figure.png" pageId="25" pageNumber="26">Figs 9293</figureCitation>
) round, with vertex posteriorly broadly rounded; pronotum (
<figureCitation id="137552C3FFE95019FEA0FADDFEA34158" box="[276,358,1297,1323]" captionStart="FIGURES 9198" captionStartId="26.[151,264,1681,1705]" captionTargetBox="[151,1418,181,1636]" captionTargetId="figure-28@26.[151,1435,181,1657]" captionTargetPageId="26" captionText="FIGURES 9198. Anhoraeomorphus anosyensis (Franz), paratype. Habitus in dorsal view (91); head in anterodorsal view (92); head and prothorax in lateral view (only bristles on head are shown) (93); antenna (94); aedeagus in ventral (95, 97) and lateral (96, 98) views." figureDoi="http://doi.org/10.5281/zenodo.13745955" httpUri="https://zenodo.org/record/13745955/files/figure.png" pageId="25" pageNumber="26">Fig. 91</figureCitation>
) bell-shaped, broadest in anterior third, with one pair of large antebasal pits; antennomeres 36 (
<figureCitation id="137552C3FFE95019FF2BFAF9FF36413C" box="[159,243,1333,1359]" captionStart="FIGURES 9198" captionStartId="26.[151,264,1681,1705]" captionTargetBox="[151,1418,181,1636]" captionTargetId="figure-28@26.[151,1435,181,1657]" captionTargetPageId="26" captionText="FIGURES 9198. Anhoraeomorphus anosyensis (Franz), paratype. Habitus in dorsal view (91); head in anterodorsal view (92); head and prothorax in lateral view (only bristles on head are shown) (93); antenna (94); aedeagus in ventral (95, 97) and lateral (96, 98) views." figureDoi="http://doi.org/10.5281/zenodo.13745955" httpUri="https://zenodo.org/record/13745955/files/figure.png" pageId="25" pageNumber="26">Fig. 94</figureCitation>
) each elongate; all femora similarly slender; metatibiae in male modified, strongly broadening to distal third, then narrowing to apex, with tuft of setae on outer surface at widest point; aedeagus (
<figureCitation id="137552C3FFE95019FB14FA95FAE44107" box="[1184,1313,1369,1396]" captionStart="FIGURES 9198" captionStartId="26.[151,264,1681,1705]" captionTargetBox="[151,1418,181,1636]" captionTargetId="figure-28@26.[151,1435,181,1657]" captionTargetPageId="26" captionText="FIGURES 9198. Anhoraeomorphus anosyensis (Franz), paratype. Habitus in dorsal view (91); head in anterodorsal view (92); head and prothorax in lateral view (only bristles on head are shown) (93); antenna (94); aedeagus in ventral (95, 97) and lateral (96, 98) views." figureDoi="http://doi.org/10.5281/zenodo.13745955" httpUri="https://zenodo.org/record/13745955/files/figure.png" pageId="25" pageNumber="26">Figs 9598</figureCitation>
) in ventral view broadest near base, apical region of dorsal wall in lateral view weakly curved dorsally, distolateral endophallic sclerites narrowly separated, each strongly elongate and with subtriangular apex, each paramere with subapical group of five short setae distant from apex.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFE9501BFF73FA25FD9C464E" lastPageId="27" lastPageNumber="28" pageId="25" pageNumber="26" type="description">
<paragraph id="8BF14E46FFE95019FF73FA25FEE9425B" blockId="25.[151,1437,1153,1972]" pageId="25" pageNumber="26">
<emphasis id="B93A9254FFE95019FF73FA25FEB04270" bold="true" box="[199,373,1513,1539]" pageId="25" pageNumber="26">Redescription.</emphasis>
Body of male (
<figureCitation id="137552C3FFE95019FDA8FA25FDAE4270" box="[540,619,1513,1539]" captionStart="FIGURES 9198" captionStartId="26.[151,264,1681,1705]" captionTargetBox="[151,1418,181,1636]" captionTargetId="figure-28@26.[151,1435,181,1657]" captionTargetPageId="26" captionText="FIGURES 9198. Anhoraeomorphus anosyensis (Franz), paratype. Habitus in dorsal view (91); head in anterodorsal view (92); head and prothorax in lateral view (only bristles on head are shown) (93); antenna (94); aedeagus in ventral (95, 97) and lateral (96, 98) views." figureDoi="http://doi.org/10.5281/zenodo.13745955" httpUri="https://zenodo.org/record/13745955/files/figure.png" pageId="25" pageNumber="26">Fig. 91</figureCitation>
) moderately dark brown, tarsi and palps distinctly lighter, setae yellowish; BL
<quantity id="4CB6E3A3FFE95019FF75F9C1FEED425B" box="[193,296,1549,1576]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" pageId="25" pageNumber="26" unit="mm" value="2.15">2.15 mm</quantity>
.
</paragraph>
<paragraph id="8BF14E46FFE95019FF73F9FDFCE24350" blockId="25.[151,1437,1153,1972]" pageId="25" pageNumber="26">
Head (
<figureCitation id="137552C3FFE95019FEA0F9FDFEAC4238" box="[276,361,1585,1611]" captionStart="FIGURES 9198" captionStartId="26.[151,264,1681,1705]" captionTargetBox="[151,1418,181,1636]" captionTargetId="figure-28@26.[151,1435,181,1657]" captionTargetPageId="26" captionText="FIGURES 9198. Anhoraeomorphus anosyensis (Franz), paratype. Habitus in dorsal view (91); head in anterodorsal view (92); head and prothorax in lateral view (only bristles on head are shown) (93); antenna (94); aedeagus in ventral (95, 97) and lateral (96, 98) views." figureDoi="http://doi.org/10.5281/zenodo.13745955" httpUri="https://zenodo.org/record/13745955/files/figure.png" pageId="25" pageNumber="26">Fig. 92</figureCitation>
) in anterodorsal view round, broadest at eyes, HL
<quantity id="4CB6E3A3FFE95019FC0DF9FDFBE7423F" box="[953,1058,1585,1612]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="25" pageNumber="26" unit="mm" value="0.33">0.33 mm</quantity>
, HW
<quantity id="4CB6E3A3FFE95019FBDCF9FDFB17423F" box="[1128,1234,1585,1612]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="25" pageNumber="26" unit="mm" value="0.4">0.40 mm</quantity>
; temple in lateral view (
<figureCitation id="137552C3FFE95019FF6AF999FEF5421C" box="[222,304,1621,1647]" captionStart="FIGURES 9198" captionStartId="26.[151,264,1681,1705]" captionTargetBox="[151,1418,181,1636]" captionTargetId="figure-28@26.[151,1435,181,1657]" captionTargetPageId="26" captionText="FIGURES 9198. Anhoraeomorphus anosyensis (Franz), paratype. Habitus in dorsal view (91); head in anterodorsal view (92); head and prothorax in lateral view (only bristles on head are shown) (93); antenna (94); aedeagus in ventral (95, 97) and lateral (96, 98) views." figureDoi="http://doi.org/10.5281/zenodo.13745955" httpUri="https://zenodo.org/record/13745955/files/figure.png" pageId="25" pageNumber="26">Fig. 93</figureCitation>
) subequal to eye; vertex and frons confluent, together weakly convex and weakly transverse; vertex posteriorly broadly rounded; each eye moderately large and weakly convex, oval, weakly projecting from head silhouette. Punctures on vertex fine, inconspicuous; setae short, dense and suberect; frons and vertex densely covered with thick bristles directed posteriorly. Antenna (
<figureCitation id="137552C3FFE95019FC97F90DFCB042A8" box="[803,885,1729,1755]" captionStart="FIGURES 9198" captionStartId="26.[151,264,1681,1705]" captionTargetBox="[151,1418,181,1636]" captionTargetId="figure-28@26.[151,1435,181,1657]" captionTargetPageId="26" captionText="FIGURES 9198. Anhoraeomorphus anosyensis (Franz), paratype. Habitus in dorsal view (91); head in anterodorsal view (92); head and prothorax in lateral view (only bristles on head are shown) (93); antenna (94); aedeagus in ventral (95, 97) and lateral (96, 98) views." figureDoi="http://doi.org/10.5281/zenodo.13745955" httpUri="https://zenodo.org/record/13745955/files/figure.png" pageId="25" pageNumber="26">Fig. 94</figureCitation>
) slender but short, as long as half BL, AnL
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, antennomeres 16 each weakly elongate, 710 each about as long as broad, 11 much shorter than 9 and 10 combined, as wide as 10, about 1.8 times as long as broad.
</paragraph>
<paragraph id="8BF14E46FFE95019FF73F8E2FAAA43C0" blockId="25.[151,1437,1153,1972]" pageId="25" pageNumber="26">
Pronotum bell-shaped, broadest in anterior third; PL
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, PW
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. Anterior margin strongly rounded, lateral margins rounded in anterior half, weakly sinuate in posterior half; posterior corners blunt; posterior margin nearly straight. Base with one pair of large round pits, and with short but distinct sublateral carinae. Disc with fine, inconspicuous punctures; setae dense, short and suberect, lateral surface of pronotum with dense thick bristles.
</paragraph>
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<paragraph id="8BF14E46FFEA501AFF23F95DFEF54282" blockId="26.[151,1436,1681,1777]" pageId="26" pageNumber="27">
<emphasis id="B93A9254FFEA501AFF23F95DFE9642DA" bold="true" box="[151,339,1681,1705]" pageId="26" pageNumber="27">FIGURES 9198.</emphasis>
<taxonomicName id="4C4E35C5FFEA501AFEECF95DFD1F42DA" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[344,730,1681,1705]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="anosyensis">
<emphasis id="B93A9254FFEA501AFEECF95DFD4D42DA" box="[344,648,1681,1705]" italics="true" pageId="26" pageNumber="27">Anhoraeomorphus anosyensis</emphasis>
(Franz)
</taxonomicName>
, paratype. Habitus in dorsal view (
<emphasis id="B93A9254FFEA501AFB82F95DFB8A42DA" bold="true" box="[1078,1103,1681,1705]" pageId="26" pageNumber="27">91</emphasis>
); head in anterodorsal view (
<emphasis id="B93A9254FFEA501AFAC7F95DFA4942DA" bold="true" box="[1395,1420,1681,1705]" pageId="26" pageNumber="27">92</emphasis>
); head and prothorax in lateral view (only bristles on head are shown) (
<emphasis id="B93A9254FFEA501AFCE3F979FCB542BE" bold="true" box="[855,880,1717,1741]" pageId="26" pageNumber="27">93</emphasis>
); antenna (
<emphasis id="B93A9254FFEA501AFC57F979FC3942BE" bold="true" box="[995,1020,1717,1741]" pageId="26" pageNumber="27">94</emphasis>
); aedeagus in ventral (
<emphasis id="B93A9254FFEA501AFB50F979FAE642BE" bold="true" box="[1252,1315,1717,1741]" pageId="26" pageNumber="27">95, 97</emphasis>
) and lateral (
<emphasis id="B93A9254FFEA501AFF14F915FF1A4282" bold="true" box="[160,223,1753,1777]" pageId="26" pageNumber="27">96, 98</emphasis>
) views.
</paragraph>
</caption>
<paragraph id="8BF14E46FFEA501AFF73F88FFEAE43BA" blockId="26.[151,1436,1859,1993]" pageId="26" pageNumber="27">
Elytra together oval, broadest distinctly in front of middle; EL
<quantity id="4CB6E3A3FFEA501AFC39F88FFC31432D" box="[909,1012,1859,1886]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="26" pageNumber="27" unit="mm" value="1.25">1.25 mm</quantity>
, EW
<quantity id="4CB6E3A3FFEA501AFB87F88FFB5F432D" box="[1075,1178,1859,1886]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="26" pageNumber="27" unit="mm" value="0.85">0.85 mm</quantity>
, EI 1.47; humeral calli weakly elevated, basal elytral foveae vestigial, barely discernible. Punctures small and shallow, but more distinct than those on pronotum, those in anterior half separated by spaces subequal to diameters of punctures; setae sparse, short and suberect.
</paragraph>
<paragraph id="8BF14E46FFEB501BFF73FF5BFF3E456E" blockId="27.[151,1437,151,969]" pageId="27" pageNumber="28">Legs moderately long, slender, unmodified except large and concave patch of adhesive setae on apical mesal region of protibia and slightly recurved and strongly broadening metatibiae, which are broadest near distal third, where outer margin bears tuft of long setae, and from this point tibia is narrowing distally and its outer margin is concave.</paragraph>
<paragraph id="8BF14E46FFEB501BFF73FEEBFE67466A" blockId="27.[151,1437,151,969]" pageId="27" pageNumber="28">
Aedeagus (
<figureCitation id="137552C3FFEB501BFEF6FEEBFE054532" box="[322,448,294,321]" captionStart="FIGURES 9198" captionStartId="26.[151,264,1681,1705]" captionTargetBox="[151,1418,181,1636]" captionTargetId="figure-28@26.[151,1435,181,1657]" captionTargetPageId="26" captionText="FIGURES 9198. Anhoraeomorphus anosyensis (Franz), paratype. Habitus in dorsal view (91); head in anterodorsal view (92); head and prothorax in lateral view (only bristles on head are shown) (93); antenna (94); aedeagus in ventral (95, 97) and lateral (96, 98) views." figureDoi="http://doi.org/10.5281/zenodo.13745955" httpUri="https://zenodo.org/record/13745955/files/figure.png" pageId="27" pageNumber="28">Figs 9598</figureCitation>
) slender,AeL
<quantity id="4CB6E3A3FFEB501BFDEFFEEBFD054532" box="[603,704,295,321]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="27" pageNumber="28" unit="mm" value="0.4">0.40 mm</quantity>
; in ventral view broadest near base, drop-shaped, distally gradually narrowing towards broadly subtriangular apex with rounded tip; in lateral view apical region of dorsal wall weakly curved dorsally; endophallus with asymmetrical and conspicuously long distolateral sclerites and with strongly asymmetrical assemblage of median proximal sclerotized structures, distolateral sclerites narrowly separated, each with subtriangular apex; parameres broad and their apices slightly projecting beyond aedeagal apex, each paramere with twisted short distal region appearing in ventral view as an apical loop, and with subapical group of five short setae distant from apex.
</paragraph>
<paragraph id="8BF14E46FFEB501BFF73FDEFFD9C464E" blockId="27.[151,1437,151,969]" box="[199,601,547,573]" pageId="27" pageNumber="28">Female. Not studied (see Remarks).</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFEB501BFF73FD8AFCD84612" box="[199,797,582,609]" pageId="27" pageNumber="28" type="distribution">
<paragraph id="8BF14E46FFEB501BFF73FD8AFCD84612" blockId="27.[151,1437,151,969]" box="[199,797,582,609]" pageId="27" pageNumber="28">
<emphasis id="B93A9254FFEB501BFF73FD8AFEA44613" bold="true" box="[199,353,582,608]" pageId="27" pageNumber="28">Distribution.</emphasis>
Southern
<collectingCountry id="F3590ED6FFEB501BFE60FD8BFD9E4612" box="[468,603,583,609]" name="Madagascar" pageId="27" pageNumber="28">Madagascar</collectingCountry>
(Anosy Region).
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFEB501BFF73FDA7FCFB47BA" pageId="27" pageNumber="28" type="discussion">
<paragraph id="8BF14E46FFEB501BFF73FDA7FAED474A" blockId="27.[151,1437,151,969]" pageId="27" pageNumber="28">
<emphasis id="B93A9254FFEB501BFF73FDA7FEFE46F6" bold="true" box="[199,315,619,645]" pageId="27" pageNumber="28">Remarks.</emphasis>
<taxonomicName id="4C4E35C5FFEB501BFEF7FDA7FD5146F7" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[323,660,619,645]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="anosyensis">
<emphasis id="B93A9254FFEB501BFEF7FDA7FD5146F7" box="[323,660,619,645]" italics="true" pageId="27" pageNumber="28">Anhoraeomorphus anosyensis</emphasis>
</taxonomicName>
is highly similar to
<taxonomicName id="4C4E35C5FFEB501BFCCEFDA7FBEE46F7" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[890,1067,619,645]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="roussettensis">
<emphasis id="B93A9254FFEB501BFCCEFDA7FBEE46F7" box="[890,1067,619,645]" italics="true" pageId="27" pageNumber="28">A. roussettensis</emphasis>
</taxonomicName>
. These are the only species with males having metatibiae strongly broadening distally, but in subapical regions rapidly narrowing, so that the outer subapical margin is concave, and there is a tuft of setae on the outer angulate broadening near distal third. These two species share also some structures of the aedeagus,
<emphasis id="B93A9254FFEB501BFCA1FD1BFCF84683" box="[789,829,727,752]" italics="true" pageId="27" pageNumber="28">e.g.</emphasis>
, a pair of very long and dark distolateral endophallic sclerites. However,
<taxonomicName id="4C4E35C5FFEB501BFEC1FD37FDCB4767" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[373,526,763,789]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="anosyensis">
<emphasis id="B93A9254FFEB501BFEC1FD37FDCB4767" box="[373,526,763,789]" italics="true" pageId="27" pageNumber="28">A. anosyensis</emphasis>
</taxonomicName>
has distinctly slenderer antennae with antennomeres 36 each elongate, whereas in
<taxonomicName id="4C4E35C5FFEB501BFF00FCD3FEA6474B" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[180,355,799,825]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="roussettensis">
<emphasis id="B93A9254FFEB501BFF00FCD3FEA6474B" box="[180,355,799,825]" italics="true" pageId="27" pageNumber="28">A. roussettensis</emphasis>
</taxonomicName>
antennomeres 36 are as long as broad, and the antennae are distinctly more massive.
</paragraph>
<paragraph id="8BF14E46FFEB501BFF73FC8FFCFB47BA" blockId="27.[151,1437,151,969]" pageId="27" pageNumber="28">
<materialsCitation id="3B26441BFFEB501BFF73FC8FFCFF47BA" collectionCode="NHMW" pageId="27" pageNumber="28" specimenCount="3" typeStatus="holotype">
The
<typeStatus id="54F5F0E4FFEB501BFF4DFC8FFE9E472E" box="[249,347,835,861]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
male and a
<typeStatus id="54F5F0E4FFEB501BFE69FC8FFDF8472E" box="[477,573,835,861]" pageId="27" pageNumber="28" type="paratype">paratype</typeStatus>
female of this species are deposited in Muséum National d'Histoire Naturelle, Paris and were not available for my study. The examined
<typeStatus id="54F5F0E4FFEB501BFC92FCABFC4347F2" box="[806,902,871,897]" pageId="27" pageNumber="28" type="paratype">paratype</typeStatus>
male (
<collectionCode id="ED5FD683FFEB501BFC60FCABFBF647F2" box="[980,1075,871,897]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="27" pageNumber="28" type="Museum">NHMW</collectionCode>
) has the aedeagus matching the illustration in the original description (including the uniquely shaped parameral apices). Fort-Dauphin indicated in label data is a French name for the Malagasy city Tolagnaro
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,268 @@
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<subSubSection id="C3541DCDFFEB501BFF23FBD6FD094046" box="[151,716,1050,1077]" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="8BF14E46FFEB501BFF23FBD6FD094046" blockId="27.[151,716,1050,1113]" box="[151,716,1050,1077]" pageId="27" pageNumber="28">
<heading id="D0B9F92AFFEB501BFF23FBD6FD094046" bold="true" box="[151,716,1050,1077]" fontSize="11" level="2" pageId="27" pageNumber="28" reason="4">
<emphasis id="B93A9254FFEB501BFF23FBD6FD094046" bold="true" box="[151,716,1050,1077]" pageId="27" pageNumber="28">
<taxonomicName id="4C4E35C5FFEB501BFF23FBD6FD914047" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[151,596,1050,1077]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="barbatus" status="comb. nov.">
<emphasis id="B93A9254FFEB501BFF23FBD6FE284046" bold="true" box="[151,493,1050,1077]" italics="true" pageId="27" pageNumber="28">Anhoraeomorphus barbatus</emphasis>
(Franz)
</taxonomicName>
,
<taxonomicNameLabel id="A2092F2FFFEB501BFDD5FBD7FD094046" box="[609,716,1051,1077]" pageId="27" pageNumber="28" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFEB501BFF23FBF3FEF0402A" box="[151,309,1087,1113]" pageId="27" pageNumber="28" type="description">
<paragraph id="8BF14E46FFEB501BFF23FBF3FEF0402A" blockId="27.[151,716,1050,1113]" box="[151,309,1087,1113]" pageId="27" pageNumber="28">
(
<figureCitation id="137552C3FFEB501BFF2BFBF3FEEB402A" box="[159,302,1087,1113]" captionStart="FIGURES 99106" captionStartId="28.[151,264,1481,1505]" captionTargetBox="[151,1428,181,1441]" captionTargetId="figure-28@28.[151,1435,181,1457]" captionTargetPageId="28" captionText="FIGURES 99106. Anhoraeomorphus barbatus (Franz), holotype. Habitus in dorsal view (99); head in anterodorsal view (100); head and prothorax in lateral view (only bristles on head are shown) (101); antenna (102); aedeagus in ventral (103, 105) and lateral (104, 106) views." figureDoi="http://doi.org/10.5281/zenodo.13745957" httpUri="https://zenodo.org/record/13745957/files/figure.png" pageId="27" pageNumber="28">Figs 99106</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFEB501BFF23FB4FFDA140E8" box="[151,612,1155,1179]" pageId="27" pageNumber="28" type="reference_group">
<paragraph id="8BF14E46FFEB501BFF23FB4FFDA140E8" blockId="27.[151,612,1155,1179]" box="[151,612,1155,1179]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="AB5E6968FFEB501BFF23FB4FFDA140E8" box="[151,612,1155,1179]" pageId="27" pageNumber="28">
<treatmentCitation id="0AEF6857FFEB501BFF23FB4FFDA540E8" author="Franz, H." box="[151,608,1155,1179]" page="167" pageId="27" pageNumber="28" year="1986">
<taxonomicName id="4C4E35C5FFEB501BFF23FB4FFDA540E8" ID-CoL="3MNJH" authority="Franz, 1986 b: 167" authorityName="Franz" authorityPageNumber="167" authorityYear="1986" box="[151,608,1155,1179]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="barbatus">
<emphasis id="B93A9254FFEB501BFF23FB4FFE5B40E8" box="[151,414,1155,1179]" italics="true" pageId="27" pageNumber="28">Horaeomorphus barbatus</emphasis>
<bibRefCitation id="EFDF33B7FFEB501BFE10FB48FDA540E8" author="Franz, H." box="[420,608,1155,1179]" pageId="27" pageNumber="28" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 167</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFEB501BFF23FB05FB004254" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="8BF14E46FFEB501BFF23FB05FD3A413C" blockId="27.[151,1437,1225,1936]" pageId="27" pageNumber="28">
<emphasis id="B93A9254FFEB501BFF23FB05FE5A4090" bold="true" box="[151,415,1225,1251]" pageId="27" pageNumber="28">Type material studied.</emphasis>
<materialsCitation id="3B26441BFFEB501BFE10FB05FD164158" collectionCode="NHMW" collectorName="H. Franz" country="Madagascar" determinerName="H. Franz" location="Madagascar" pageId="27" pageNumber="28" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
<emphasis id="B93A9254FFEB501BFE10FB05FD784090" bold="true" box="[420,701,1225,1251]" pageId="27" pageNumber="28">
<typeStatus id="54F5F0E4FFEB501BFE10FB05FDD44090" box="[420,529,1225,1251]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
(
<collectingCountry id="F3590ED6FFEB501BFDA9FB05FD714090" box="[541,692,1225,1251]" name="Madagascar" pageId="27" pageNumber="28">Madagascar</collectingCountry>
)
</emphasis>
:
<specimenCount id="9D4885CFFFEB501BFD7DFB06FD254097" box="[713,736,1226,1252]" count="1" pageId="27" pageNumber="28" type="male"></specimenCount>
, four labels: “Madagaskar,Plateau / du Tampoketsa / lg.
<collectorName id="26BB2B90FFEB501BFA8AFB05FA594090" box="[1342,1436,1225,1251]" pageId="27" pageNumber="28">H.Franz</collectorName>
1969” [white, printed], “
<specimenCount id="9D4885CFFFEB501BFE1EFB21FE054174" box="[426,448,1261,1287]" count="1" pageId="27" pageNumber="28" type="male"></specimenCount>
” [white, printed], “
<taxonomicName id="4C4E35C5FFEB501BFD23FB21FC8B4174" box="[663,846,1261,1287]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/ barbatus m. / det.
<determinerName id="7E2DA282FFEB501BFB90FB21FB474174" box="[1060,1154,1261,1287]" pageId="27" pageNumber="28">H.Franz</determinerName>
” [yellowish, handwritten and printed], “Typus” [red, handwritten] (
<collectionCode id="ED5FD683FFEB501BFDDCFADDFD0E4158" box="[616,715,1297,1323]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="27" pageNumber="28" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B26441BFFEB501BFD69FADDFD3E413C" collectionCode="NHMW" collectorName="H. Franz" country="Madagascar" determinerName="H. Franz" location="Madagascar" pageId="27" pageNumber="28" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B93A9254FFEB501BFD69FADDFC904158" bold="true" box="[733,853,1297,1323]" pageId="27" pageNumber="28">
<typeStatus id="54F5F0E4FFEB501BFD69FADDFC904158" box="[733,853,1297,1323]" pageId="27" pageNumber="28" type="paratype">Paratypes</typeStatus>
</emphasis>
(see Remarks):
<specimenCount id="9D4885CFFFEB501BFBBEFADDFBF3415F" box="[1034,1078,1297,1324]" count="1" pageId="27" pageNumber="28" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D4885CFFFEB501BFBF5FADDFBAE415F" box="[1089,1131,1297,1324]" count="1" pageId="27" pageNumber="28" type="female">1 ♀</specimenCount>
, same data as for holotype, except for yellowish “
<typeStatus id="54F5F0E4FFEB501BFE38FAF9FDF5413C" box="[396,560,1333,1359]" pageId="27" pageNumber="28" type="paratype">PARATYPUS</typeStatus>
” labels (
<collectionCode id="ED5FD683FFEB501BFD24FAF9FD36413C" box="[656,755,1333,1359]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="27" pageNumber="28" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF14E46FFEB501BFF73FA95FB004254" blockId="27.[151,1437,1225,1936]" pageId="27" pageNumber="28">
<emphasis id="B93A9254FFEB501BFF73FA95FE5A4100" bold="true" box="[199,415,1369,1395]" pageId="27" pageNumber="28">Revised diagnosis.</emphasis>
Body (
<figureCitation id="137552C3FFEB501BFE5AFA95FDF84100" box="[494,573,1369,1395]" captionStart="FIGURES 99106" captionStartId="28.[151,264,1481,1505]" captionTargetBox="[151,1428,181,1441]" captionTargetId="figure-28@28.[151,1435,181,1457]" captionTargetPageId="28" captionText="FIGURES 99106. Anhoraeomorphus barbatus (Franz), holotype. Habitus in dorsal view (99); head in anterodorsal view (100); head and prothorax in lateral view (only bristles on head are shown) (101); antenna (102); aedeagus in ventral (103, 105) and lateral (104, 106) views." figureDoi="http://doi.org/10.5281/zenodo.13745957" httpUri="https://zenodo.org/record/13745957/files/figure.png" pageId="27" pageNumber="28">Fig. 99</figureCitation>
) slender; head (
<figureCitation id="137552C3FFEB501BFD5EFA95FC464100" box="[746,899,1369,1395]" captionStart="FIGURES 99106" captionStartId="28.[151,264,1481,1505]" captionTargetBox="[151,1428,181,1441]" captionTargetId="figure-28@28.[151,1435,181,1457]" captionTargetPageId="28" captionText="FIGURES 99106. Anhoraeomorphus barbatus (Franz), holotype. Habitus in dorsal view (99); head in anterodorsal view (100); head and prothorax in lateral view (only bristles on head are shown) (101); antenna (102); aedeagus in ventral (103, 105) and lateral (104, 106) views." figureDoi="http://doi.org/10.5281/zenodo.13745957" httpUri="https://zenodo.org/record/13745957/files/figure.png" pageId="27" pageNumber="28">Figs 100101</figureCitation>
) round, with vertex posteriorly broadly rounded; pronotum (
<figureCitation id="137552C3FFEB501BFEA2FAB1FEAC41E4" box="[278,361,1405,1431]" captionStart="FIGURES 99106" captionStartId="28.[151,264,1481,1505]" captionTargetBox="[151,1428,181,1441]" captionTargetId="figure-28@28.[151,1435,181,1457]" captionTargetPageId="28" captionText="FIGURES 99106. Anhoraeomorphus barbatus (Franz), holotype. Habitus in dorsal view (99); head in anterodorsal view (100); head and prothorax in lateral view (only bristles on head are shown) (101); antenna (102); aedeagus in ventral (103, 105) and lateral (104, 106) views." figureDoi="http://doi.org/10.5281/zenodo.13745957" httpUri="https://zenodo.org/record/13745957/files/figure.png" pageId="27" pageNumber="28">Fig. 99</figureCitation>
) bell-shaped, broadest near anterior third, with one pair of large antebasal pits; antennomeres 37 (
<figureCitation id="137552C3FFEB501BFF66FA6DFEE041C8" box="[210,293,1441,1467]" captionStart="FIGURES 9198" captionStartId="26.[151,264,1681,1705]" captionTargetBox="[151,1418,181,1636]" captionTargetId="figure-28@26.[151,1435,181,1657]" captionTargetPageId="26" captionText="FIGURES 9198. Anhoraeomorphus anosyensis (Franz), paratype. Habitus in dorsal view (91); head in anterodorsal view (92); head and prothorax in lateral view (only bristles on head are shown) (93); antenna (94); aedeagus in ventral (95, 97) and lateral (96, 98) views." figureDoi="http://doi.org/10.5281/zenodo.13745955" httpUri="https://zenodo.org/record/13745955/files/figure.png" pageId="27" pageNumber="28">Fig. 94</figureCitation>
) each slightly elongate; all femora similarly slender; hind tibiae in male unmodified; aedeagus (
<figureCitation id="137552C3FFEB501BFADEFA6DFF3F41AC" captionStart="FIGURES 99106" captionStartId="28.[151,264,1481,1505]" captionTargetBox="[151,1428,181,1441]" captionTargetId="figure-28@28.[151,1435,181,1457]" captionTargetPageId="28" captionText="FIGURES 99106. Anhoraeomorphus barbatus (Franz), holotype. Habitus in dorsal view (99); head in anterodorsal view (100); head and prothorax in lateral view (only bristles on head are shown) (101); antenna (102); aedeagus in ventral (103, 105) and lateral (104, 106) views." figureDoi="http://doi.org/10.5281/zenodo.13745957" httpUri="https://zenodo.org/record/13745957/files/figure.png" pageId="27" pageNumber="28">Figs 103106</figureCitation>
) in ventral view broadest in sub-basal third, apical region of dorsal wall in lateral view strongly curved dorsally, distolateral endophallic sclerites narrowly separated, each elongate and with subtriangular apex, each paramere with group of numerous subapical and apical setae occupying about half of its length.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFEB501DFF73F9FDFDAB45A2" lastPageId="29" lastPageNumber="30" pageId="27" pageNumber="28" type="description">
<paragraph id="8BF14E46FFEB501BFF73F9FDFEAE4203" blockId="27.[151,1437,1225,1936]" pageId="27" pageNumber="28">
<emphasis id="B93A9254FFEB501BFF73F9FDFEB04238" bold="true" box="[199,373,1585,1611]" pageId="27" pageNumber="28">Redescription.</emphasis>
Body of male (
<figureCitation id="137552C3FFEB501BFDA8F9FDFDAE4238" box="[540,619,1585,1611]" captionStart="FIGURES 99106" captionStartId="28.[151,264,1481,1505]" captionTargetBox="[151,1428,181,1441]" captionTargetId="figure-28@28.[151,1435,181,1457]" captionTargetPageId="28" captionText="FIGURES 99106. Anhoraeomorphus barbatus (Franz), holotype. Habitus in dorsal view (99); head in anterodorsal view (100); head and prothorax in lateral view (only bristles on head are shown) (101); antenna (102); aedeagus in ventral (103, 105) and lateral (104, 106) views." figureDoi="http://doi.org/10.5281/zenodo.13745957" httpUri="https://zenodo.org/record/13745957/files/figure.png" pageId="27" pageNumber="28">Fig. 99</figureCitation>
) moderately dark brown, tarsi and palps distinctly lighter, setae yellowish; BL
<quantity id="4CB6E3A3FFEB501BFF75F999FEA24203" box="[193,359,1621,1648]" metricMagnitude="-3" metricUnit="m" metricValue="2.3049999999999997" metricValueMax="2.31" metricValueMin="2.3" pageId="27" pageNumber="28" unit="mm" value="2.3049999999999997" valueMax="2.31" valueMin="2.3">2.302.31 mm</quantity>
.
</paragraph>
<paragraph id="8BF14E46FFEB501BFF73F9B5FDC243FC" blockId="27.[151,1437,1225,1936]" pageId="27" pageNumber="28">
Head (
<figureCitation id="137552C3FFEB501BFEA7F9B5FEB642E0" box="[275,371,1657,1683]" captionStart="FIGURES 99106" captionStartId="28.[151,264,1481,1505]" captionTargetBox="[151,1428,181,1441]" captionTargetId="figure-28@28.[151,1435,181,1457]" captionTargetPageId="28" captionText="FIGURES 99106. Anhoraeomorphus barbatus (Franz), holotype. Habitus in dorsal view (99); head in anterodorsal view (100); head and prothorax in lateral view (only bristles on head are shown) (101); antenna (102); aedeagus in ventral (103, 105) and lateral (104, 106) views." figureDoi="http://doi.org/10.5281/zenodo.13745957" httpUri="https://zenodo.org/record/13745957/files/figure.png" pageId="27" pageNumber="28">Fig. 100</figureCitation>
) in anterodorsal view round, broadest at eyes, HL
<quantity id="4CB6E3A3FFEB501BFC07F9B5FB9F42E7" box="[947,1114,1657,1684]" metricMagnitude="-4" metricUnit="m" metricValue="4.15" metricValueMax="4.3" metricValueMin="4.0" pageId="27" pageNumber="28" unit="mm" value="0.415" valueMax="0.43" valueMin="0.4">0.400.43 mm</quantity>
, HW
<quantity id="4CB6E3A3FFEB501BFB28F9B5FA8142E7" box="[1180,1348,1657,1684]" metricMagnitude="-4" metricUnit="m" metricValue="4.15" metricValueMax="4.3" metricValueMin="4.0" pageId="27" pageNumber="28" unit="mm" value="0.415" valueMax="0.43" valueMin="0.4">0.400.43 mm</quantity>
; temple in lateral view (
<figureCitation id="137552C3FFEB501BFEF8F951FE6842C4" box="[332,429,1693,1719]" captionStart="FIGURES 99106" captionStartId="28.[151,264,1481,1505]" captionTargetBox="[151,1428,181,1441]" captionTargetId="figure-28@28.[151,1435,181,1457]" captionTargetPageId="28" captionText="FIGURES 99106. Anhoraeomorphus barbatus (Franz), holotype. Habitus in dorsal view (99); head in anterodorsal view (100); head and prothorax in lateral view (only bristles on head are shown) (101); antenna (102); aedeagus in ventral (103, 105) and lateral (104, 106) views." figureDoi="http://doi.org/10.5281/zenodo.13745957" httpUri="https://zenodo.org/record/13745957/files/figure.png" pageId="27" pageNumber="28">Fig. 101</figureCitation>
) subequal to eye; vertex and frons confluent, together weakly convex and about as long as broad; vertex posteriorly broadly rounded; each eye moderately large and weakly convex, oval with distinct posteromedian emargination, weakly projecting from head silhouette. Punctures on vertex fine, inconspicuous; setae short, dense and suberect; frons and vertex densely covered with thick bristles directed posteriorly. Antenna (
<figureCitation id="137552C3FFEB501BFF2BF8E1FF3A4334" box="[159,255,1837,1863]" captionStart="FIGURES 99106" captionStartId="28.[151,264,1481,1505]" captionTargetBox="[151,1428,181,1441]" captionTargetId="figure-28@28.[151,1435,181,1457]" captionTargetPageId="28" captionText="FIGURES 99106. Anhoraeomorphus barbatus (Franz), holotype. Habitus in dorsal view (99); head in anterodorsal view (100); head and prothorax in lateral view (only bristles on head are shown) (101); antenna (102); aedeagus in ventral (103, 105) and lateral (104, 106) views." figureDoi="http://doi.org/10.5281/zenodo.13745957" httpUri="https://zenodo.org/record/13745957/files/figure.png" pageId="27" pageNumber="28">Fig. 102</figureCitation>
) slender but short, indistinctly shorter than half BL, AnL
<quantity id="4CB6E3A3FFEB501BFC3EF8E1FBF5433B" box="[906,1072,1837,1864]" metricMagnitude="-3" metricUnit="m" metricValue="1.09" metricValueMax="1.1" metricValueMin="1.08" pageId="27" pageNumber="28" unit="mm" value="1.09" valueMax="1.1" valueMin="1.08">1.081.10 mm</quantity>
, antennomeres 17 each weakly elongate, 89 each about as long as broad, 10 slightly transverse, 11 shorter than 9 and 10 combined, as wide as 10, about 1.8 times as long as broad.
</paragraph>
<caption id="DF311ECEFFEC501CFF23FA05FE7F425A" ID-DOI="http://doi.org/10.5281/zenodo.13745957" ID-Zenodo-Dep="13745957" httpUri="https://zenodo.org/record/13745957/files/figure.png" pageId="28" pageNumber="29" startId="28.[151,264,1481,1505]" targetBox="[151,1428,181,1441]" targetPageId="28" targetType="figure">
<paragraph id="8BF14E46FFEC501CFF23FA05FE7F425A" blockId="28.[151,1436,1481,1578]" pageId="28" pageNumber="29">
<emphasis id="B93A9254FFEC501CFF23FA05FEA14192" bold="true" box="[151,356,1481,1505]" pageId="28" pageNumber="29">FIGURES 99106.</emphasis>
<taxonomicName id="4C4E35C5FFEC501CFED9FA05FD214191" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[365,740,1481,1506]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="barbatus">
<emphasis id="B93A9254FFEC501CFED9FA05FD4B4192" box="[365,654,1481,1505]" italics="true" pageId="28" pageNumber="29">Anhoraeomorphus barbatus</emphasis>
(Franz)
</taxonomicName>
, holotype. Habitus in dorsal view (
<emphasis id="B93A9254FFEC501CFBEFFA05FBB14192" bold="true" box="[1115,1140,1481,1505]" pageId="28" pageNumber="29">99</emphasis>
); head in anterodorsal view (
<emphasis id="B93A9254FFEC501CFF14FA21FF034276" bold="true" box="[160,198,1517,1541]" pageId="28" pageNumber="29">100</emphasis>
); head and prothorax in lateral view (only bristles on head are shown) (
<emphasis id="B93A9254FFEC501CFC23FA21FC784276" bold="true" box="[919,957,1517,1541]" pageId="28" pageNumber="29">101</emphasis>
); antenna (
<emphasis id="B93A9254FFEC501CFB9BFA21FB904276" bold="true" box="[1071,1109,1517,1541]" pageId="28" pageNumber="29">102</emphasis>
); aedeagus in ventral (
<emphasis id="B93A9254FFEC501CFA88FA21FA514276" bold="true" box="[1340,1428,1517,1541]" pageId="28" pageNumber="29">103, 105</emphasis>
) and lateral (
<emphasis id="B93A9254FFEC501CFEA4F9DDFEAC425A" bold="true" box="[272,361,1553,1577]" pageId="28" pageNumber="29">104, 106</emphasis>
) views.
</paragraph>
</caption>
<paragraph id="8BF14E46FFEC501CFF73F9B0FEEA4355" blockId="28.[151,1437,1659,1974]" pageId="28" pageNumber="29">
Pronotum bell-shaped, broadest near anterior third; PL
<quantity id="4CB6E3A3FFEC501CFCF2F9B0FC2B42E5" box="[838,1006,1660,1686]" metricMagnitude="-4" metricUnit="m" metricValue="6.2" metricValueMax="6.3" metricValueMin="6.1" pageId="28" pageNumber="29" unit="mm" value="0.62" valueMax="0.63" valueMin="0.61">0.610.63 mm</quantity>
, PW
<quantity id="4CB6E3A3FFEC501CFB84F9B7FB1C42E5" box="[1072,1241,1659,1686]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" metricValueMax="5.4" metricValueMin="5.0" pageId="28" pageNumber="29" unit="mm" value="0.52" valueMax="0.54" valueMin="0.5">0.500.54 mm</quantity>
. Anterior margin strongly rounded, lateral margins rounded in anterior half, weakly sinuate in posterior half; posterior corners blunt; posterior margin nearly straight. Base with one pair of large round pits, and with short but distinct sublateral carinae. Disc with fine, inconspicuous punctures; setae dense, short and suberect, lateral surface of pronotum with dense thick bristles.
</paragraph>
<paragraph id="8BF14E46FFEC501CFF73F8FCFA5E43E1" blockId="28.[151,1437,1659,1974]" pageId="28" pageNumber="29">
Elytra together oval, broadest slightly in front of middle; EL
<quantity id="4CB6E3A3FFEC501CFC2CF8FCFB844339" box="[920,1089,1840,1867]" metricMagnitude="-3" metricUnit="m" metricValue="1.275" metricValueMax="1.3" metricValueMin="1.25" pageId="28" pageNumber="29" unit="mm" value="1.275" valueMax="1.3" valueMin="1.25">1.251.30 mm</quantity>
, EW
<quantity id="4CB6E3A3FFEC501CFB32F8FCFB354339" box="[1158,1264,1840,1866]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" pageId="28" pageNumber="29" unit="mm" value="0.88">0.88 mm</quantity>
, EI 1.431.49; humeral calli weakly elevated, basal elytral foveae vestigial, barely discernible. Punctures very dense but extremely shallow and diffuse, so that surface of elytra is slightly less glossy than pronotum; setae sparse, short and suberect.
</paragraph>
<paragraph id="8BF14E46FFEC501CFF73F850FB3943C5" blockId="28.[151,1437,1659,1974]" box="[199,1276,1948,1974]" pageId="28" pageNumber="29">Legs moderately long, slender, unmodified except weakly curved apices of pro- and mesotibiae.</paragraph>
<paragraph id="8BF14E46FFED501DFF73FF5BFB2045FA" blockId="29.[151,1437,151,753]" pageId="29" pageNumber="30">
Aedeagus (
<figureCitation id="137552C3FFED501DFEF0FF5BFE2544C2" box="[324,480,151,177]" captionStart="FIGURES 99106" captionStartId="28.[151,264,1481,1505]" captionTargetBox="[151,1428,181,1441]" captionTargetId="figure-28@28.[151,1435,181,1457]" captionTargetPageId="28" captionText="FIGURES 99106. Anhoraeomorphus barbatus (Franz), holotype. Habitus in dorsal view (99); head in anterodorsal view (100); head and prothorax in lateral view (only bristles on head are shown) (101); antenna (102); aedeagus in ventral (103, 105) and lateral (104, 106) views." figureDoi="http://doi.org/10.5281/zenodo.13745957" httpUri="https://zenodo.org/record/13745957/files/figure.png" pageId="29" pageNumber="30">Figs 103106</figureCitation>
) slender, AeL
<quantity id="4CB6E3A3FFED501DFD37FF5BFD2F44C2" box="[643,746,151,177]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="29" pageNumber="30" unit="mm" value="0.38">0.38 mm</quantity>
; in ventral view broadest in sub-basal third, distally gradually narrowing and in subapical region rapidly narrowing towards broadly subtriangular apex with rounded tip; in lateral view apical region of dorsal wall strongly curved dorsally; endophallus with nearly symmetrical and conspicuously broad distolateral sclerites and with strongly asymmetrical assemblage of median sclerotized structures, distolateral sclerites narrowly separated, each with subtriangular apex; parameres broad and their apices slightly projecting beyond aedeagal apex, each paramere with twisted short distal region appearing in ventral view as an apical loop, and with apical-subapical group of 12 short setae proximally reaching middle of parameral length.
</paragraph>
<paragraph id="8BF14E46FFED501DFF73FE5FFDAB45A2" blockId="29.[151,1437,151,753]" pageId="29" pageNumber="30">
Female. Externally similar to male. BL
<quantity id="4CB6E3A3FFED501DFD34FE5FFD2245DE" box="[640,743,403,429]" metricMagnitude="-3" metricUnit="m" metricValue="2.33" pageId="29" pageNumber="30" unit="mm" value="2.33">2.33 mm</quantity>
; HL
<quantity id="4CB6E3A3FFED501DFCA9FE5FFC4745DE" box="[797,898,403,429]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="29" pageNumber="30" unit="mm" value="0.43">0.43 mm</quantity>
, HW
<quantity id="4CB6E3A3FFED501DFC76FE5EFBED45DE" box="[962,1064,402,429]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="29" pageNumber="30" unit="mm" value="0.45">0.45 mm</quantity>
, AnL
<quantity id="4CB6E3A3FFED501DFBDEFE5FFB1445DE" box="[1130,1233,403,429]" metricMagnitude="-3" metricUnit="m" metricValue="1.03" pageId="29" pageNumber="30" unit="mm" value="1.03">1.03 mm</quantity>
; PL
<quantity id="4CB6E3A3FFED501DFAB6FE5FFAAD45DE" box="[1282,1384,403,429]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="29" pageNumber="30" unit="mm" value="0.6">0.60 mm</quantity>
, PW
<quantity id="4CB6E3A3FFED501DFF23FE7AFF3B45A2" box="[151,254,438,465]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="29" pageNumber="30" unit="mm" value="0.53">0.53 mm</quantity>
; EL 1.30, EW
<quantity id="4CB6E3A3FFED501DFE17FE7BFDCC45A2" box="[419,521,439,465]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" pageId="29" pageNumber="30" unit="mm" value="0.88">0.88 mm</quantity>
, EI 1.49.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFED501DFF73FE16FC8A4586" box="[199,847,474,501]" pageId="29" pageNumber="30" type="distribution">
<paragraph id="8BF14E46FFED501DFF73FE16FC8A4586" blockId="29.[151,1437,151,753]" box="[199,847,474,501]" pageId="29" pageNumber="30">
<emphasis id="B93A9254FFED501DFF73FE16FEA44587" bold="true" box="[199,353,474,500]" pageId="29" pageNumber="30">Distribution.</emphasis>
Central
<collectingCountry id="F3590ED6FFED501DFE75FE17FD8D4586" box="[449,584,475,501]" name="Madagascar" pageId="29" pageNumber="30">Madagascar</collectingCountry>
(Analamanga Region).
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFED501DFF73FE33FB664682" pageId="29" pageNumber="30" type="discussion">
<paragraph id="8BF14E46FFED501DFF73FE33FEDA464E" blockId="29.[151,1437,151,753]" pageId="29" pageNumber="30">
<emphasis id="B93A9254FFED501DFF73FE33FEFE466A" bold="true" box="[199,315,511,537]" pageId="29" pageNumber="30">Remarks.</emphasis>
<taxonomicName id="4C4E35C5FFED501DFEF7FE33FD45466A" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[323,640,511,537]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="barbatus">
<emphasis id="B93A9254FFED501DFEF7FE33FD45466A" box="[323,640,511,537]" italics="true" pageId="29" pageNumber="30">Anhoraeomorphus barbatus</emphasis>
</taxonomicName>
is unremarkable and examination of aedeagus is necessary to identify this species.
</paragraph>
<paragraph id="8BF14E46FFED501DFF73FD8BFB664682" blockId="29.[151,1437,151,753]" pageId="29" pageNumber="30">
<materialsCitation id="3B26441BFFED501DFF73FD8BFB5A4682" collectionCode="NHMW" collectorName="All &amp; Franz" elevation="1500" location="Seehohe" pageId="29" pageNumber="30" specimenCount="6" specimenCount-female="1" specimenCount-male="1" typeStatus="holotype">
<bibRefCitation id="EFDF33B7FFED501DFF73FD8BFEA34612" author="Franz, H." box="[199,358,583,609]" pageId="29" pageNumber="30" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986b)</bibRefCitation>
mentions
<specimenCount id="9D4885CFFFED501DFE6EFD8BFD414612" box="[474,644,583,609]" count="4" pageId="29" pageNumber="30" type="generic">four specimens</specimenCount>
included in the type series, of which
<specimenCount id="9D4885CFFFED501DFB90FD8BFB4C4612" box="[1060,1161,583,609]" count="1" pageId="29" pageNumber="30" type="male">one male</specimenCount>
was fixed as a
<typeStatus id="54F5F0E4FFED501DFA87FD8BFA5C4612" box="[1331,1433,583,609]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
.
<collectorName id="26BB2B90FFED501DFF23FDA7FF7E46F6" box="[151,187,619,645]" pageId="29" pageNumber="30">All</collectorName>
specimens are preserved at
<collectionCode id="ED5FD683FFED501DFDB1FDA7FDA446F6" box="[517,609,619,645]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="29" pageNumber="30" type="Museum">NHMW</collectionCode>
, but
<specimenCount id="9D4885CFFFED501DFD28FDA7FC4046F6" box="[668,901,619,645]" count="1" pageId="29" pageNumber="30" type="female" typeStatus="paratype">one paratype female</specimenCount>
is clearly different from remaining specimens and does not belong in
<taxonomicName id="4C4E35C5FFED501DFE19FD43FDF146DA" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[429,564,655,681]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="barbatus">
<emphasis id="B93A9254FFED501DFE19FD43FDF146DA" box="[429,564,655,681]" italics="true" pageId="29" pageNumber="30">A. barbatus</emphasis>
</taxonomicName>
; it must be excluded from the type series. “Plateau du Tampoketsa” is the only locality given in label data, but in the original description
<collectorName id="26BB2B90FFED501DFCD6FD7FFC6746BE" box="[866,930,691,717]" pageId="29" pageNumber="30">Franz</collectorName>
adds more detailed information: “Plateau du Tampokeza {sic!} dAnkazobe nächst der Forststation Manonkazo im ca.
<quantity id="4CB6E3A3FFED501DFC60FD1AFBEC4682" box="[980,1065,726,753]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="29" pageNumber="30" unit="m" value="1500.0">
<elevation id="0063A975FFED501DFC60FD1AFBEC4682" box="[980,1065,726,753]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="29" pageNumber="30" unit="m" value="1500.0">1500 m</elevation>
</quantity>
<location id="8E91189DFFED501DFB84FD1BFB504682" LSID="urn:lsid:plazi:treatment:03E7FF50FFEB501DFF23FBD6FB664682:8E91189DFFED501DFB84FD1BFB504682" box="[1072,1173,727,753]" name="Seehohe" pageId="29" pageNumber="30">Seehöhe</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,208 @@
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<treatment id="03E7FF50FFED501EFF23FC8EFAD44016" ID-DOI="http://doi.org/10.5281/zenodo.13746041" ID-Zenodo-Dep="13746041" LSID="urn:lsid:plazi:treatment:03E7FF50FFED501EFF23FC8EFAD44016" httpUri="http://treatment.plazi.org/id/03E7FF50FFED501EFF23FC8EFAD44016" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
<subSubSection id="C3541DCDFFED501DFF23FC8EFD26472E" box="[151,739,834,861]" pageId="29" pageNumber="30" type="nomenclature">
<paragraph id="8BF14E46FFED501DFF23FC8EFD26472E" blockId="29.[151,739,834,897]" box="[151,739,834,861]" pageId="29" pageNumber="30">
<heading id="D0B9F92AFFED501DFF23FC8EFD26472E" bold="true" box="[151,739,834,861]" fontSize="11" level="2" pageId="29" pageNumber="30" reason="4">
<emphasis id="B93A9254FFED501DFF23FC8EFD26472E" bold="true" box="[151,739,834,861]" pageId="29" pageNumber="30">
<taxonomicName id="4C4E35C5FFED501DFF23FC8EFDAE472F" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[151,619,834,861]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="compactus" status="comb. nov.">
<emphasis id="B93A9254FFED501DFF23FC8EFDC1472E" bold="true" box="[151,516,834,861]" italics="true" pageId="29" pageNumber="30">Anhoraeomorphus compactus</emphasis>
(Franz)
</taxonomicName>
,
<taxonomicNameLabel id="A2092F2FFFED501DFDCCFC8FFD26472E" box="[632,739,835,861]" pageId="29" pageNumber="30" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFED501DFF23FCABFE8747F2" box="[151,322,871,897]" pageId="29" pageNumber="30" type="description">
<paragraph id="8BF14E46FFED501DFF23FCABFE8747F2" blockId="29.[151,739,834,897]" box="[151,322,871,897]" pageId="29" pageNumber="30">
(
<figureCitation id="137552C3FFED501DFF2BFCABFEFF47F2" box="[159,314,871,897]" captionStart="FIGURES 107110" captionStartId="30.[151,264,777,801]" captionTargetBox="[151,1425,181,753]" captionTargetId="figure-28@30.[151,1435,181,753]" captionTargetPageId="30" captionText="FIGURES 107110. Anhoraeomorphus compactus (Franz), holotype. Habitus in dorsal view (107); head in anterodorsal view (108); head and prothorax in lateral view (only bristles on head are shown) (109); antenna (110)." figureDoi="http://doi.org/10.5281/zenodo.13745959" httpUri="https://zenodo.org/record/13745959/files/figure.png" pageId="29" pageNumber="30">Figs 107110</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFED501DFF23FC67FDB347B0" box="[151,630,939,963]" pageId="29" pageNumber="30" type="reference_group">
<paragraph id="8BF14E46FFED501DFF23FC67FDB347B0" blockId="29.[151,630,939,963]" box="[151,630,939,963]" pageId="29" pageNumber="30">
<treatmentCitationGroup id="AB5E6968FFED501DFF23FC67FDB347B0" box="[151,630,939,963]" pageId="29" pageNumber="30">
<treatmentCitation id="0AEF6857FFED501DFF23FC67FDB747B0" author="Franz, H." box="[151,626,939,963]" page="168" pageId="29" pageNumber="30" year="1986">
<taxonomicName id="4C4E35C5FFED501DFF23FC67FDB747B0" ID-CoL="3MNJR" authority="Franz, 1986 b: 168" authorityName="Franz" authorityPageNumber="168" authorityYear="1986" box="[151,626,939,963]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="compactus">
<emphasis id="B93A9254FFED501DFF23FC67FE7547B0" box="[151,432,939,963]" italics="true" pageId="29" pageNumber="30">Horaeomorphus compactus</emphasis>
<bibRefCitation id="EFDF33B7FFED501DFE02FC60FDB747B0" author="Franz, H." box="[438,626,939,963]" pageId="29" pageNumber="30" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 168</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFED501DFF23FC3DFA9F4090" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="8BF14E46FFED501DFF23FC3DFC334020" blockId="29.[151,1437,1009,2007]" pageId="29" pageNumber="30">
<emphasis id="B93A9254FFED501DFF23FC3DFE634078" bold="true" box="[151,422,1009,1035]" pageId="29" pageNumber="30">Type material studied.</emphasis>
<materialsCitation id="3B26441BFFED501DFE1BFC3DFC334020" collectionCode="NHMW" collectorName="Umg. Ft. Dauphin &amp; H. Franz" country="Madagascar" determinerName="H. Franz" location="Madagascar" pageId="29" pageNumber="30" specimenCount="2" specimenCount-female="2" typeStatus="holotype">
<emphasis id="B93A9254FFED501DFE1BFC3DFD094078" bold="true" box="[431,716,1009,1035]" pageId="29" pageNumber="30">
<typeStatus id="54F5F0E4FFED501DFE1BFC3DFDD94078" box="[431,540,1009,1035]" pageId="29" pageNumber="30" type="holotype">Holotype</typeStatus>
(
<collectingCountry id="F3590ED6FFED501DFD98FC3DFD064078" box="[556,707,1009,1035]" name="Madagascar" pageId="29" pageNumber="30">Madagascar</collectingCountry>
)
</emphasis>
:
<specimenCount id="9D4885CFFFED501DFD68FC3EFD34407F" box="[732,753,1010,1036]" count="1" pageId="29" pageNumber="30" type="female"></specimenCount>
, five labels: “S-Madagaskar /
<collectorName id="26BB2B90FFED501DFBFAFC3DFAC94078" box="[1102,1292,1009,1035]" pageId="29" pageNumber="30">Umg.Ft.Dauphin</collectorName>
/ lg.
<collectorName id="26BB2B90FFED501DFA8AFC3DFA594078" box="[1342,1436,1009,1035]" pageId="29" pageNumber="30">H.Franz</collectorName>
1969” [white, printed], “
<specimenCount id="9D4885CFFFED501DFE07FBD9FE00405C" box="[435,453,1045,1071]" count="1" pageId="29" pageNumber="30" type="female"></specimenCount>
” [white, printed], “Scierie de / Bimanguidy” [white, handwritten], “
<taxonomicName id="4C4E35C5FFED501DFB60FBD9FA4E405C" box="[1236,1419,1045,1071]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/ compactus m. / det.
<determinerName id="7E2DA282FFED501DFEC5FBF5FE0A4020" box="[369,463,1081,1107]" pageId="29" pageNumber="30">H.Franz</determinerName>
” [yellowish, handwritten and printed] (
<collectionCode id="ED5FD683FFED501DFC3CFBF5FC2E4020" box="[904,1003,1081,1107]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="29" pageNumber="30" type="Museum">NHMW</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BF14E46FFED501DFF73FB91FA9F4090" blockId="29.[151,1437,1009,2007]" pageId="29" pageNumber="30">
<emphasis id="B93A9254FFED501DFF73FB91FE674004" bold="true" box="[199,418,1117,1143]" pageId="29" pageNumber="30">Revised diagnosis.</emphasis>
Body (
<figureCitation id="137552C3FFED501DFE42FB91FD934004" box="[502,598,1117,1143]" captionStart="FIGURES 107110" captionStartId="30.[151,264,777,801]" captionTargetBox="[151,1425,181,753]" captionTargetId="figure-28@30.[151,1435,181,753]" captionTargetPageId="30" captionText="FIGURES 107110. Anhoraeomorphus compactus (Franz), holotype. Habitus in dorsal view (107); head in anterodorsal view (108); head and prothorax in lateral view (only bristles on head are shown) (109); antenna (110)." figureDoi="http://doi.org/10.5281/zenodo.13745959" httpUri="https://zenodo.org/record/13745959/files/figure.png" pageId="29" pageNumber="30">Fig. 107</figureCitation>
) stout and conspicuously strongly convex; head (
<figureCitation id="137552C3FFED501DFB30FB91FAE44004" box="[1156,1313,1117,1143]" captionStart="FIGURES 107110" captionStartId="30.[151,264,777,801]" captionTargetBox="[151,1425,181,753]" captionTargetId="figure-28@30.[151,1435,181,753]" captionTargetPageId="30" captionText="FIGURES 107110. Anhoraeomorphus compactus (Franz), holotype. Habitus in dorsal view (107); head in anterodorsal view (108); head and prothorax in lateral view (only bristles on head are shown) (109); antenna (110)." figureDoi="http://doi.org/10.5281/zenodo.13745959" httpUri="https://zenodo.org/record/13745959/files/figure.png" pageId="29" pageNumber="30">Figs 108109</figureCitation>
) strikingly small compared to prothorax and distinctly elongate, with vertex posteriorly broadly rounded; pronotum (
<figureCitation id="137552C3FFED501DFADBFB4DFF0740CC" captionStart="FIGURES 107110" captionStartId="30.[151,264,777,801]" captionTargetBox="[151,1425,181,753]" captionTargetId="figure-28@30.[151,1435,181,753]" captionTargetPageId="30" captionText="FIGURES 107110. Anhoraeomorphus compactus (Franz), holotype. Habitus in dorsal view (107); head in anterodorsal view (108); head and prothorax in lateral view (only bristles on head are shown) (109); antenna (110)." figureDoi="http://doi.org/10.5281/zenodo.13745959" httpUri="https://zenodo.org/record/13745959/files/figure.png" pageId="29" pageNumber="30">Fig. 107</figureCitation>
) round, broadest between middle and anterior third, lacking antebasal pits; antennomeres 35 (
<figureCitation id="137552C3FFED501DFB4DFB69FA9C40CC" box="[1273,1369,1189,1215]" captionStart="FIGURES 107110" captionStartId="30.[151,264,777,801]" captionTargetBox="[151,1425,181,753]" captionTargetId="figure-28@30.[151,1435,181,753]" captionTargetPageId="30" captionText="FIGURES 107110. Anhoraeomorphus compactus (Franz), holotype. Habitus in dorsal view (107); head in anterodorsal view (108); head and prothorax in lateral view (only bristles on head are shown) (109); antenna (110)." figureDoi="http://doi.org/10.5281/zenodo.13745959" httpUri="https://zenodo.org/record/13745959/files/figure.png" pageId="29" pageNumber="30">Fig. 110</figureCitation>
) each elongate, 6 as long as broad, and 710 each transverse; profemora much thicker than meso- and metafemora.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFED501DFF73FB21FE4043C0" pageId="29" pageNumber="30" type="description">
<paragraph id="8BF14E46FFED501DFF73FB21FE6D415F" blockId="29.[151,1437,1009,2007]" pageId="29" pageNumber="30">
<emphasis id="B93A9254FFED501DFF73FB21FEB04174" bold="true" box="[199,373,1261,1287]" pageId="29" pageNumber="30">Redescription.</emphasis>
Body of female (
<figureCitation id="137552C3FFED501DFD8DFB21FD5D4174" box="[569,664,1261,1287]" captionStart="FIGURES 107110" captionStartId="30.[151,264,777,801]" captionTargetBox="[151,1425,181,753]" captionTargetId="figure-28@30.[151,1435,181,753]" captionTargetPageId="30" captionText="FIGURES 107110. Anhoraeomorphus compactus (Franz), holotype. Habitus in dorsal view (107); head in anterodorsal view (108); head and prothorax in lateral view (only bristles on head are shown) (109); antenna (110)." figureDoi="http://doi.org/10.5281/zenodo.13745959" httpUri="https://zenodo.org/record/13745959/files/figure.png" pageId="29" pageNumber="30">Fig. 107</figureCitation>
) dark brown with reddish hue, tarsi and palps distinctly lighter, setae yellowish; BL
<quantity id="4CB6E3A3FFED501DFE89FADDFE61415F" box="[317,420,1297,1324]" metricMagnitude="-3" metricUnit="m" metricValue="2.78" pageId="29" pageNumber="30" unit="mm" value="2.78">2.78 mm</quantity>
.
</paragraph>
<paragraph id="8BF14E46FFED501DFF73FAF9FADF4254" blockId="29.[151,1437,1009,2007]" pageId="29" pageNumber="30">
Head (
<figureCitation id="137552C3FFED501DFEA3FAF9FEBE413C" box="[279,379,1333,1359]" captionStart="FIGURES 107110" captionStartId="30.[151,264,777,801]" captionTargetBox="[151,1425,181,753]" captionTargetId="figure-28@30.[151,1435,181,753]" captionTargetPageId="30" captionText="FIGURES 107110. Anhoraeomorphus compactus (Franz), holotype. Habitus in dorsal view (107); head in anterodorsal view (108); head and prothorax in lateral view (only bristles on head are shown) (109); antenna (110)." figureDoi="http://doi.org/10.5281/zenodo.13745959" httpUri="https://zenodo.org/record/13745959/files/figure.png" pageId="29" pageNumber="30">Fig. 108</figureCitation>
) in anterodorsal view elongate, broadest at eyes, HL
<quantity id="4CB6E3A3FFED501DFC48FAF9FBA24123" box="[1020,1127,1333,1360]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="29" pageNumber="30" unit="mm" value="0.58">0.58 mm</quantity>
, HW
<quantity id="4CB6E3A3FFED501DFB06FAF9FADB4123" box="[1202,1310,1333,1360]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="29" pageNumber="30" unit="mm" value="0.48">0.48 mm</quantity>
; temple in lateral view (
<figureCitation id="137552C3FFED501DFE9BFA95FE544100" box="[303,401,1369,1395]" captionStart="FIGURES 107110" captionStartId="30.[151,264,777,801]" captionTargetBox="[151,1425,181,753]" captionTargetId="figure-28@30.[151,1435,181,753]" captionTargetPageId="30" captionText="FIGURES 107110. Anhoraeomorphus compactus (Franz), holotype. Habitus in dorsal view (107); head in anterodorsal view (108); head and prothorax in lateral view (only bristles on head are shown) (109); antenna (110)." figureDoi="http://doi.org/10.5281/zenodo.13745959" httpUri="https://zenodo.org/record/13745959/files/figure.png" pageId="29" pageNumber="30">Fig. 109</figureCitation>
) slightly longer than eye; vertex and frons confluent, together weakly convex and clearly elongate; vertex posteriorly broadly rounded; each eye small and weakly convex, oval with shallow posteromedian emargination, weakly projecting from head silhouette. Punctures on vertex fine, inconspicuous; setae short, dense and suberect; frons and vertex densely covered with thick bristles directed posteriorly. Antenna (
<figureCitation id="137552C3FFED501DFB60FA09FAF741AC" box="[1236,1330,1477,1503]" captionStart="FIGURES 107110" captionStartId="30.[151,264,777,801]" captionTargetBox="[151,1425,181,753]" captionTargetId="figure-28@30.[151,1435,181,753]" captionTargetPageId="30" captionText="FIGURES 107110. Anhoraeomorphus compactus (Franz), holotype. Habitus in dorsal view (107); head in anterodorsal view (108); head and prothorax in lateral view (only bristles on head are shown) (109); antenna (110)." figureDoi="http://doi.org/10.5281/zenodo.13745959" httpUri="https://zenodo.org/record/13745959/files/figure.png" pageId="29" pageNumber="30">Fig. 110</figureCitation>
) massive and short, as long as half BL, AnL
<quantity id="4CB6E3A3FFED501DFD92FA25FD494277" box="[550,652,1513,1540]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" pageId="29" pageNumber="30" unit="mm" value="1.35">1.35 mm</quantity>
, antennomeres 15 each distinctly elongate, 6 as long as broad, 710 each transverse, 11 clearly longer than 9 and 10 combined, as wide as 10, about twice as long as broad.
</paragraph>
<paragraph id="8BF14E46FFED501DFF73F9FEFAAA42C4" blockId="29.[151,1437,1009,2007]" pageId="29" pageNumber="30">
Pronotum round, broadest between middle and anterior third; PL
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, PW
<quantity id="4CB6E3A3FFED501DFBD9F9FDFB1D423F" box="[1133,1240,1585,1612]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="29" pageNumber="30" unit="mm" value="0.7">0.70 mm</quantity>
. Anterior margin strongly rounded, lateral margins rounded in anterior half, nearly straight in posterior half; posterior corners blunt; posterior margin nearly straight. Base lacking pits, but with short but distinct sublateral carinae. Disc with fine, inconspicuous punctures; setae dense, short and suberect, lateral surface of pronotum with dense thick bristles.
</paragraph>
<paragraph id="8BF14E46FFED501DFF73F90DFED04334" blockId="29.[151,1437,1009,2007]" pageId="29" pageNumber="30">
Elytra together oval and conspicuously strongly convex, broadest slightly in front of middle; EL
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, EW
<quantity id="4CB6E3A3FFED501DFF23F929FF384373" box="[151,253,1765,1792]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="29" pageNumber="30" unit="mm" value="1.2">1.20 mm</quantity>
, EI 1.23; humeral calli not marked, basal elytral foveae indiscernible. Punctures very dense but extremely shallow and diffuse, so that surface of elytra is slightly less glossy than pronotum; setae sparse, short and nearly recumbent.
</paragraph>
<paragraph id="8BF14E46FFED501DFF73F89EFE1443FC" blockId="29.[151,1437,1009,2007]" pageId="29" pageNumber="30">Legs long, slender, unmodified except thickened protibiae with large distal patch of adhesive setae and weakly curved pro- and mesotibiae.</paragraph>
<paragraph id="8BF14E46FFED501DFF73F855FE4043C0" blockId="29.[151,1437,1009,2007]" box="[199,389,1945,1971]" pageId="29" pageNumber="30">Male. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFED501DFF73F871FCD843A4" box="[199,797,1981,2007]" pageId="29" pageNumber="30" type="distribution">
<paragraph id="8BF14E46FFED501DFF73F871FCD843A4" blockId="29.[151,1437,1009,2007]" box="[199,797,1981,2007]" pageId="29" pageNumber="30">
<emphasis id="B93A9254FFED501DFF73F871FEA443A4" bold="true" box="[199,353,1981,2007]" pageId="29" pageNumber="30">Distribution.</emphasis>
Southern
<collectingCountry id="F3590ED6FFED501DFE60F871FD9E43A4" box="[468,603,1981,2007]" name="Madagascar" pageId="29" pageNumber="30">Madagascar</collectingCountry>
(Anosy Region).
</paragraph>
</subSubSection>
<caption id="DF311ECEFFEE501EFF23FCC5FBAC4736" ID-DOI="http://doi.org/10.5281/zenodo.13745959" ID-Zenodo-Dep="13745959" httpUri="https://zenodo.org/record/13745959/files/figure.png" pageId="30" pageNumber="31" startId="30.[151,264,777,801]" targetBox="[151,1425,181,753]" targetPageId="30" targetType="figure">
<paragraph id="8BF14E46FFEE501EFF23FCC5FBAC4736" blockId="30.[151,1436,777,837]" pageId="30" pageNumber="31">
<emphasis id="B93A9254FFEE501EFF23FCC5FEA94752" bold="true" box="[151,364,777,801]" pageId="30" pageNumber="31">FIGURES 107110.</emphasis>
<taxonomicName id="4C4E35C5FFEE501EFEC7FCC5FD334752" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[371,758,777,801]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="compactus">
<emphasis id="B93A9254FFEE501EFEC7FCC5FD674752" box="[371,674,777,801]" italics="true" pageId="30" pageNumber="31">Anhoraeomorphus compactus</emphasis>
(Franz)
</taxonomicName>
, holotype. Habitus in dorsal view (
<emphasis id="B93A9254FFEE501EFBEEFCC5FB454752" bold="true" box="[1114,1152,777,801]" pageId="30" pageNumber="31">107</emphasis>
); head in anterodorsal view (
<emphasis id="B93A9254FFEE501EFF14FCE1FF034736" bold="true" box="[160,198,813,837]" pageId="30" pageNumber="31">108</emphasis>
); head and prothorax in lateral view (only bristles on head are shown) (
<emphasis id="B93A9254FFEE501EFC28FCE1FC074736" bold="true" box="[924,962,813,837]" pageId="30" pageNumber="31">109</emphasis>
); antenna (
<emphasis id="B93A9254FFEE501EFB82FCE1FB9E4736" bold="true" box="[1078,1115,813,837]" pageId="30" pageNumber="31">110</emphasis>
).
</paragraph>
</caption>
<subSubSection id="C3541DCDFFEE501EFF73FC54FAD44016" pageId="30" pageNumber="31" type="discussion">
<paragraph id="8BF14E46FFEE501EFF73FC54FA8D4789" blockId="30.[151,1437,919,1126]" pageId="30" pageNumber="31">
<emphasis id="B93A9254FFEE501EFF73FC54FEFE47C1" bold="true" box="[199,315,920,946]" pageId="30" pageNumber="31">Remarks.</emphasis>
<taxonomicName id="4C4E35C5FFEE501EFEF7FC5BFD5647C2" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[323,659,919,945]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="compactus">
<emphasis id="B93A9254FFEE501EFEF7FC5BFD5647C2" box="[323,659,919,945]" italics="true" pageId="30" pageNumber="31">Anhoraeomorphus compactus</emphasis>
</taxonomicName>
is known from only a single
<typeStatus id="54F5F0E4FFEE501EFC54FC5BFB8747C2" box="[992,1090,919,945]" pageId="30" pageNumber="31" type="holotype">holotype</typeStatus>
female, which differs from all congeners in strongly convex and stout elytra (EI merely 1.23), round pronotum lacking antebasal pits, small and elongate head, and massive antennae strongly broadening and with strongly transverse antennomeres 710.
</paragraph>
<paragraph id="8BF14E46FFEE501EFF73FBCFFAD44016" blockId="30.[151,1437,919,1126]" pageId="30" pageNumber="31">
The label information “Umg.Ft.Dauphin” means in fact ~
<quantity id="4CB6E3A3FFEE501EFCEDFBC8FC74406E" box="[857,945,1027,1054]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="30" pageNumber="31" unit="km" value="100.0">100 km</quantity>
north of Fort-Dauphin, according to
<bibRefCitation id="EFDF33B7FFEE501EFAE8FBC8FF364032" author="Franz, H." pageId="30" pageNumber="31" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986b)</bibRefCitation>
. An additional label written in a hand different than Franzs reads “Scierie de / Bimanguidy” (
<emphasis id="B93A9254FFEE501EFAA7FBE4FAF34032" box="[1299,1334,1064,1089]" italics="true" pageId="30" pageNumber="31">i.e.</emphasis>
, sawmill Bimanguidy), whereas
<bibRefCitation id="EFDF33B7FFEE501EFE28FB80FDFE4016" author="Franz, H." box="[412,571,1099,1126]" pageId="30" pageNumber="31" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986b)</bibRefCitation>
translates this into German with an error, as “Säge Bemangidy”.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,255 @@
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<subSubSection id="C3541DCDFFEE501EFF23FB7BFD6E40A1" box="[151,683,1207,1234]" pageId="30" pageNumber="31" type="nomenclature">
<paragraph id="8BF14E46FFEE501EFF23FB7BFD6E40A1" blockId="30.[151,683,1207,1270]" box="[151,683,1207,1234]" pageId="30" pageNumber="31">
<heading id="D0B9F92AFFEE501EFF23FB7BFD6E40A1" bold="true" box="[151,683,1207,1234]" fontSize="11" level="2" pageId="30" pageNumber="31" reason="4">
<emphasis id="B93A9254FFEE501EFF23FB7BFD6E40A1" bold="true" box="[151,683,1207,1234]" pageId="30" pageNumber="31">
<taxonomicName id="4C4E35C5FFEE501EFF23FB7BFDF640A2" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[151,563,1207,1234]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="diegoi" status="comb. nov.">
<emphasis id="B93A9254FFEE501EFF23FB7BFE0940A1" bold="true" box="[151,460,1207,1234]" italics="true" pageId="30" pageNumber="31">Anhoraeomorphus diegoi</emphasis>
(Franz)
</taxonomicName>
,
<taxonomicNameLabel id="A2092F2FFFEE501EFDF5FB74FD6E40A1" box="[577,683,1208,1234]" pageId="30" pageNumber="31" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFEE501EFF23FB17FE854085" box="[151,320,1243,1270]" pageId="30" pageNumber="31" type="description">
<paragraph id="8BF14E46FFEE501EFF23FB17FE854085" blockId="30.[151,683,1207,1270]" box="[151,320,1243,1270]" pageId="30" pageNumber="31">
(
<figureCitation id="137552C3FFEE501EFF2BFB17FEFD4085" box="[159,312,1243,1270]" captionStart="FIGURES 111118" captionStartId="31.[151,264,1391,1415]" captionTargetBox="[151,1422,181,1357]" captionTargetId="figure-16@31.[151,1435,181,1367]" captionTargetPageId="31" captionText="FIGURES 111118. Anhoraeomorphus diegoi (Franz), paratype. Habitus in dorsal view (111); head in anterodorsal view (112); head and prothorax in lateral view (only bristles on head are shown) (113); antenna (114); aedeagus in ventral (115, 117) and lateral (116, 118) views." figureDoi="http://doi.org/10.5281/zenodo.13745961" httpUri="https://zenodo.org/record/13745961/files/figure.png" pageId="30" pageNumber="31">Figs 111118</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFEE501EFF23FAD5FD8C4142" box="[151,585,1305,1329]" pageId="30" pageNumber="31" type="reference_group">
<paragraph id="8BF14E46FFEE501EFF23FAD5FD8C4142" blockId="30.[151,585,1305,1329]" box="[151,585,1305,1329]" pageId="30" pageNumber="31">
<treatmentCitationGroup id="AB5E6968FFEE501EFF23FAD5FD8C4142" box="[151,585,1305,1329]" pageId="30" pageNumber="31">
<treatmentCitation id="0AEF6857FFEE501EFF23FAD5FD804142" author="Franz, H." box="[151,581,1305,1329]" page="156" pageId="30" pageNumber="31" year="1986">
<taxonomicName id="4C4E35C5FFEE501EFF23FAD5FD804142" ID-CoL="6MDVK" authority="Franz, 1986 b: 156" authorityName="Franz" authorityPageNumber="156" authorityYear="1986" box="[151,581,1305,1329]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="diegoi">
<emphasis id="B93A9254FFEE501EFF23FAD5FE464142" box="[151,387,1305,1329]" italics="true" pageId="30" pageNumber="31">Horaeomorphus diegoi</emphasis>
<bibRefCitation id="EFDF33B7FFEE501EFE3DFAD6FD804142" author="Franz, H." box="[393,581,1305,1329]" pageId="30" pageNumber="31" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 156</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFEE501EFF23FA95FB0742C4" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="8BF14E46FFEE501EFF23FA95FD6841C8" blockId="30.[151,1437,1368,2043]" pageId="30" pageNumber="31">
<emphasis id="B93A9254FFEE501EFF23FA95FE754101" bold="true" box="[151,432,1368,1395]" pageId="30" pageNumber="31">Type material studied.</emphasis>
<materialsCitation id="3B26441BFFEE501EFE0AFA95FD6F41C8" collectionCode="MUSEUM, PARIS, BATHIE, NHMW" country="Madagascar" determinerName="H. Franz" location="MADAGASCAR" pageId="30" pageNumber="31" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B93A9254FFEE501EFE0AFA95FD254101" bold="true" box="[446,736,1368,1395]" pageId="30" pageNumber="31">
<typeStatus id="54F5F0E4FFEE501EFE0AFA95FDEE4100" box="[446,555,1369,1395]" pageId="30" pageNumber="31" type="paratype">Paratype</typeStatus>
(Madagascar)
</emphasis>
:
<specimenCount id="9D4885CFFFEE501EFD40FA95FCCE4100" box="[756,779,1369,1395]" count="1" pageId="30" pageNumber="31" type="male"></specimenCount>
, three labels: “
<collectionCode id="ED5FD683FFEE501EFC70FA95FB584100" box="[964,1181,1369,1395]" pageId="30" pageNumber="31">MUSEUM PARIS</collectionCode>
/
<collectingCountry id="F3590ED6FFEE501EFB74FA95FA424100" box="[1216,1415,1369,1395]" name="Madagascar" pageId="30" pageNumber="31">MADAGASCAR</collectingCountry>
/ (PERRIER DE LA
<collectionCode id="ED5FD683FFEE501EFE30FAB1FE2941E4" box="[388,492,1405,1431]" pageId="30" pageNumber="31">BATHIE</collectionCode>
)” [bluish, printed], “
<taxonomicName id="4C4E35C5FFEE501EFD56FAB1FC5C41E4" box="[738,921,1405,1431]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/ diegoi m. / det.
<determinerName id="7E2DA282FFEE501EFBC0FAB1FB1441E4" box="[1140,1233,1405,1431]" pageId="30" pageNumber="31">H.Franz</determinerName>
/
<typeStatus id="54F5F0E4FFEE501EFB46FAB1FA5641E4" box="[1266,1427,1405,1431]" pageId="30" pageNumber="31" type="paratype">PARATYPUS</typeStatus>
” [yellowish, handwritten and printed] (
<collectionCode id="ED5FD683FFEE501EFD8BFA6DFD6741C8" box="[575,674,1441,1467]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="30" pageNumber="31" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF14E46FFEE501EFF73FA08FB0742C4" blockId="30.[151,1437,1368,2043]" pageId="30" pageNumber="31">
<emphasis id="B93A9254FFEE501EFF73FA08FE5B41AD" bold="true" box="[199,414,1476,1502]" pageId="30" pageNumber="31">Revised diagnosis.</emphasis>
Body (
<figureCitation id="137552C3FFEE501EFE58FA09FD8341AC" box="[492,582,1477,1503]" captionStart="FIGURES 111118" captionStartId="31.[151,264,1391,1415]" captionTargetBox="[151,1422,181,1357]" captionTargetId="figure-16@31.[151,1435,181,1367]" captionTargetPageId="31" captionText="FIGURES 111118. Anhoraeomorphus diegoi (Franz), paratype. Habitus in dorsal view (111); head in anterodorsal view (112); head and prothorax in lateral view (only bristles on head are shown) (113); antenna (114); aedeagus in ventral (115, 117) and lateral (116, 118) views." figureDoi="http://doi.org/10.5281/zenodo.13745961" httpUri="https://zenodo.org/record/13745961/files/figure.png" pageId="30" pageNumber="31">Fig.111</figureCitation>
) slender; head (
<figureCitation id="137552C3FFEE501EFD45FA09FC4D41AC" box="[753,904,1477,1503]" captionStart="FIGURES 111118" captionStartId="31.[151,264,1391,1415]" captionTargetBox="[151,1422,181,1357]" captionTargetId="figure-16@31.[151,1435,181,1367]" captionTargetPageId="31" captionText="FIGURES 111118. Anhoraeomorphus diegoi (Franz), paratype. Habitus in dorsal view (111); head in anterodorsal view (112); head and prothorax in lateral view (only bristles on head are shown) (113); antenna (114); aedeagus in ventral (115, 117) and lateral (116, 118) views." figureDoi="http://doi.org/10.5281/zenodo.13745961" httpUri="https://zenodo.org/record/13745961/files/figure.png" pageId="30" pageNumber="31">Figs 112113</figureCitation>
) round, with vertex posteriorly broadly rounded; pronotum (
<figureCitation id="137552C3FFEE501EFEACFA25FEBC4270" box="[280,377,1513,1539]" captionStart="FIGURES 111118" captionStartId="31.[151,264,1391,1415]" captionTargetBox="[151,1422,181,1357]" captionTargetId="figure-16@31.[151,1435,181,1367]" captionTargetPageId="31" captionText="FIGURES 111118. Anhoraeomorphus diegoi (Franz), paratype. Habitus in dorsal view (111); head in anterodorsal view (112); head and prothorax in lateral view (only bristles on head are shown) (113); antenna (114); aedeagus in ventral (115, 117) and lateral (116, 118) views." figureDoi="http://doi.org/10.5281/zenodo.13745961" httpUri="https://zenodo.org/record/13745961/files/figure.png" pageId="30" pageNumber="31">Fig. 111</figureCitation>
) bell-shaped, broadest between middle and anterior third, with one pair of large and deep antebasal pits; antennomeres 36 (
<figureCitation id="137552C3FFEE501EFDA8F9C1FDBE4254" box="[540,635,1549,1575]" captionStart="FIGURES 111118" captionStartId="31.[151,264,1391,1415]" captionTargetBox="[151,1422,181,1357]" captionTargetId="figure-16@31.[151,1435,181,1367]" captionTargetPageId="31" captionText="FIGURES 111118. Anhoraeomorphus diegoi (Franz), paratype. Habitus in dorsal view (111); head in anterodorsal view (112); head and prothorax in lateral view (only bristles on head are shown) (113); antenna (114); aedeagus in ventral (115, 117) and lateral (116, 118) views." figureDoi="http://doi.org/10.5281/zenodo.13745961" httpUri="https://zenodo.org/record/13745961/files/figure.png" pageId="30" pageNumber="31">Fig. 114</figureCitation>
) each about as long as broad, 710 each slightly transverse; profemora slightly thicker than meso- and metafemora; hind tibiae in male modified, strongly thickening to distal third and then narrowing towards apex; aedeagus (
<figureCitation id="137552C3FFEE501EFDD4F999FD3E4203" box="[608,763,1621,1648]" captionStart="FIGURES 111118" captionStartId="31.[151,264,1391,1415]" captionTargetBox="[151,1422,181,1357]" captionTargetId="figure-16@31.[151,1435,181,1367]" captionTargetPageId="31" captionText="FIGURES 111118. Anhoraeomorphus diegoi (Franz), paratype. Habitus in dorsal view (111); head in anterodorsal view (112); head and prothorax in lateral view (only bristles on head are shown) (113); antenna (114); aedeagus in ventral (115, 117) and lateral (116, 118) views." figureDoi="http://doi.org/10.5281/zenodo.13745961" httpUri="https://zenodo.org/record/13745961/files/figure.png" pageId="30" pageNumber="31">Figs 115116</figureCitation>
) in ventral view broadest in sub-basal third, apical region of dorsal wall in lateral view weakly curved dorsally, distolateral endophallic sclerites broadly separated, each elongate and with curved subtriangular apex, each paramere with minute 23 apical and subapical setae.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFEE5020FF73F90CFE5B4532" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31" type="description">
<paragraph id="8BF14E46FFEE501EFF73F90CFF0F428C" blockId="30.[151,1437,1368,2043]" pageId="30" pageNumber="31">
<emphasis id="B93A9254FFEE501EFF73F90CFEB042A9" bold="true" box="[199,373,1728,1754]" pageId="30" pageNumber="31">Redescription.</emphasis>
Body of male (
<figureCitation id="137552C3FFEE501EFD96F90DFDBA42A8" box="[546,639,1729,1755]" captionStart="FIGURES 111118" captionStartId="31.[151,264,1391,1415]" captionTargetBox="[151,1422,181,1357]" captionTargetId="figure-16@31.[151,1435,181,1367]" captionTargetPageId="31" captionText="FIGURES 111118. Anhoraeomorphus diegoi (Franz), paratype. Habitus in dorsal view (111); head in anterodorsal view (112); head and prothorax in lateral view (only bristles on head are shown) (113); antenna (114); aedeagus in ventral (115, 117) and lateral (116, 118) views." figureDoi="http://doi.org/10.5281/zenodo.13745961" httpUri="https://zenodo.org/record/13745961/files/figure.png" pageId="30" pageNumber="31">Fig. 111</figureCitation>
) flattened, moderately uniformly light brown, setae yellowish; BL
<quantity id="4CB6E3A3FFEE501EFADFF90DFF03428C" metricMagnitude="-3" metricUnit="m" metricValue="1.79" pageId="30" pageNumber="31" unit="mm" value="1.79">1.79 mm</quantity>
.
</paragraph>
<paragraph id="8BF14E46FFEE501EFF73F8C5FA844388" blockId="30.[151,1437,1368,2043]" pageId="30" pageNumber="31">
Head (
<figureCitation id="137552C3FFEE501EFEA0F8C5FEB14350" box="[276,372,1801,1827]" captionStart="FIGURES 111118" captionStartId="31.[151,264,1391,1415]" captionTargetBox="[151,1422,181,1357]" captionTargetId="figure-16@31.[151,1435,181,1367]" captionTargetPageId="31" captionText="FIGURES 111118. Anhoraeomorphus diegoi (Franz), paratype. Habitus in dorsal view (111); head in anterodorsal view (112); head and prothorax in lateral view (only bristles on head are shown) (113); antenna (114); aedeagus in ventral (115, 117) and lateral (116, 118) views." figureDoi="http://doi.org/10.5281/zenodo.13745961" httpUri="https://zenodo.org/record/13745961/files/figure.png" pageId="30" pageNumber="31">Fig. 112</figureCitation>
) in anterodorsal view round, broadest at eyes, HL
<quantity id="4CB6E3A3FFEE501EFC0AF8C5FBE34350" box="[958,1062,1801,1827]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="30" pageNumber="31" unit="mm" value="0.3">0.30 mm</quantity>
, HW
<quantity id="4CB6E3A3FFEE501EFBDFF8C5FB114350" box="[1131,1236,1801,1827]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="30" pageNumber="31" unit="mm" value="0.3">0.30 mm</quantity>
; temple in lateral view (
<figureCitation id="137552C3FFEE501EFF6AF8E1FEFB4334" box="[222,318,1837,1863]" captionStart="FIGURES 111118" captionStartId="31.[151,264,1391,1415]" captionTargetBox="[151,1422,181,1357]" captionTargetId="figure-16@31.[151,1435,181,1367]" captionTargetPageId="31" captionText="FIGURES 111118. Anhoraeomorphus diegoi (Franz), paratype. Habitus in dorsal view (111); head in anterodorsal view (112); head and prothorax in lateral view (only bristles on head are shown) (113); antenna (114); aedeagus in ventral (115, 117) and lateral (116, 118) views." figureDoi="http://doi.org/10.5281/zenodo.13745961" httpUri="https://zenodo.org/record/13745961/files/figure.png" pageId="30" pageNumber="31">Fig. 113</figureCitation>
) subequal to eye; vertex and frons confluent, together weakly convex and about as long as broad; vertex posteriorly broadly rounded; each eye moderately large and weakly convex, oval, weakly projecting from head silhouette. Punctures on vertex fine, inconspicuous; setae short, dense and suberect; frons and vertex densely covered with thick bristles directed posteriorly. Antenna (
<figureCitation id="137552C3FFEE501EFC95F855FC4543C0" box="[801,896,1945,1971]" captionStart="FIGURES 111118" captionStartId="31.[151,264,1391,1415]" captionTargetBox="[151,1422,181,1357]" captionTargetId="figure-16@31.[151,1435,181,1367]" captionTargetPageId="31" captionText="FIGURES 111118. Anhoraeomorphus diegoi (Franz), paratype. Habitus in dorsal view (111); head in anterodorsal view (112); head and prothorax in lateral view (only bristles on head are shown) (113); antenna (114); aedeagus in ventral (115, 117) and lateral (116, 118) views." figureDoi="http://doi.org/10.5281/zenodo.13745961" httpUri="https://zenodo.org/record/13745961/files/figure.png" pageId="30" pageNumber="31">Fig. 114</figureCitation>
) slender but short, indistinctly shorter than half BL, AnL
<quantity id="4CB6E3A3FFEE501EFEB0F871FEAE43A4" box="[260,363,1981,2007]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="30" pageNumber="31" unit="mm" value="0.8">0.80 mm</quantity>
, antennomeres 12 each weakly elongate, 36 each about as long as broad, 710 each weakly transverse, 11 indistinctly shorter than 9 and 10 combined, as wide as 10, about 1.8 times as long as broad.
</paragraph>
<caption id="DF311ECEFFEF501FFF23FAA3FE4941BC" ID-DOI="http://doi.org/10.5281/zenodo.13745961" ID-Zenodo-Dep="13745961" httpUri="https://zenodo.org/record/13745961/files/figure.png" pageId="31" pageNumber="32" startId="31.[151,264,1391,1415]" targetBox="[151,1422,181,1357]" targetPageId="31" targetType="figure">
<paragraph id="8BF14E46FFEF501FFF23FAA3FE4941BC" blockId="31.[151,1437,1391,1487]" pageId="31" pageNumber="32">
<emphasis id="B93A9254FFEF501FFF23FAA3FEAD41F4" bold="true" box="[151,360,1391,1415]" pageId="31" pageNumber="32">FIGURES 111118.</emphasis>
<taxonomicName id="4C4E35C5FFEF501FFEDAFAA3FD0741F4" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[366,706,1391,1415]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="diegoi">
<emphasis id="B93A9254FFEF501FFEDAFAA3FDAA41F4" box="[366,623,1391,1415]" italics="true" pageId="31" pageNumber="32">Anhoraeomorphus diegoi</emphasis>
(Franz)
</taxonomicName>
, paratype. Habitus in dorsal view (
<emphasis id="B93A9254FFEF501FFBABFAA3FB8741F4" bold="true" box="[1055,1090,1391,1415]" pageId="31" pageNumber="32">111</emphasis>
); head in anterodorsal view (
<emphasis id="B93A9254FFEF501FFADCFAA3FA4841F4" bold="true" box="[1384,1421,1391,1415]" pageId="31" pageNumber="32">112</emphasis>
); head and prothorax in lateral view (only bristles on head are shown) (
<emphasis id="B93A9254FFEF501FFCD1FA5FFC4F41D8" bold="true" box="[869,906,1427,1451]" pageId="31" pageNumber="32">113</emphasis>
); antenna (
<emphasis id="B93A9254FFEF501FFBB4FA5FFBE041D8" bold="true" box="[1024,1061,1427,1451]" pageId="31" pageNumber="32">114</emphasis>
); aedeagus in ventral (
<emphasis id="B93A9254FFEF501FFAA5FA5FFAAD41D8" bold="true" box="[1297,1384,1427,1451]" pageId="31" pageNumber="32">115, 117</emphasis>
) and lateral (
<emphasis id="B93A9254FFEF501FFF51FA7BFEF941BC" bold="true" box="[229,316,1463,1487]" pageId="31" pageNumber="32">116, 118</emphasis>
) views.
</paragraph>
</caption>
<paragraph id="8BF14E46FFEF501FFF73F9EEFF3542B8" blockId="31.[151,1436,1569,2028]" pageId="31" pageNumber="32">
Pronotum bell-shaped, broadest between middle and anterior third; PL
<quantity id="4CB6E3A3FFEF501FFC6FF9EEFB85424F" box="[987,1088,1570,1596]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="31" pageNumber="32" unit="mm" value="0.48">0.48 mm</quantity>
, PW
<quantity id="4CB6E3A3FFEF501FFBCEF9EEFB25424F" box="[1146,1248,1570,1596]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="31" pageNumber="32" unit="mm" value="0.4">0.40 mm</quantity>
. Anterior margin strongly rounded, lateral margins rounded in anterior half, strongly sinuate in posterior half; posterior corners blunt; posterior margin nearly straight. Base with one pair of large and deep pits, sublateral carinae indiscernible. Disc with fine, inconspicuous punctures; setae dense, short and suberect, lateral surface of pronotum with dense thick bristles.
</paragraph>
<paragraph id="8BF14E46FFEF501FFF73F919FA944344" blockId="31.[151,1436,1569,2028]" pageId="31" pageNumber="32">
Elytra together oval, broadest distinctly in front of middle; EL
<quantity id="4CB6E3A3FFEF501FFC39F91AFC314283" box="[909,1012,1750,1776]" metricMagnitude="-4" metricUnit="m" metricValue="9.3" pageId="31" pageNumber="32" unit="mm" value="0.93">0.93 mm</quantity>
, EW
<quantity id="4CB6E3A3FFEF501FFB87F919FB5F4283" box="[1075,1178,1749,1776]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="31" pageNumber="32" unit="mm" value="0.6">0.60 mm</quantity>
, EI 1.54; humeral calli weakly elevated, basal elytral foveae vestigial, barely discernible. Punctures very dense but extremely shallow and diffuse, so that surface of elytra appears slightly less glossy than pronotum; setae sparse, short and suberect.
</paragraph>
<paragraph id="8BF14E46FFEF501FFF73F88EFED743D0" blockId="31.[151,1436,1569,2028]" pageId="31" pageNumber="32">Legs moderately long, slender, unmodified except all tibiae, which are thick and slightly recurved, fore tibiae with large distomesal concave patch of dense adhesive setae, metatibiae strongly broadening to distal third and then narrowing.</paragraph>
<paragraph id="8BF14E46FFEF5020FF73F861FA40456E" blockId="31.[151,1436,1569,2028]" lastBlockId="32.[151,1437,151,645]" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
Aedeagus (
<figureCitation id="137552C3FFEF501FFEFCF861FE2343BB" box="[328,486,1965,1992]" captionStart="FIGURES 111118" captionStartId="31.[151,264,1391,1415]" captionTargetBox="[151,1422,181,1357]" captionTargetId="figure-16@31.[151,1435,181,1367]" captionTargetPageId="31" captionText="FIGURES 111118. Anhoraeomorphus diegoi (Franz), paratype. Habitus in dorsal view (111); head in anterodorsal view (112); head and prothorax in lateral view (only bristles on head are shown) (113); antenna (114); aedeagus in ventral (115, 117) and lateral (116, 118) views." figureDoi="http://doi.org/10.5281/zenodo.13745961" httpUri="https://zenodo.org/record/13745961/files/figure.png" pageId="31" pageNumber="32">Figs 115118</figureCitation>
) slender, AeL
<quantity id="4CB6E3A3FFEF501FFD27F862FD3B43BB" box="[659,766,1966,1992]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="31" pageNumber="32" unit="mm" value="0.28">0.28 mm</quantity>
; in ventral view broadest in sub-basal third, drop-shaped, distally gradually narrowing to broadly subtriangular apex with weakly marked obtusely angulate tip; in lateral view apical region of dorsal wall weakly curved dorsally; endophallus with nearly symmetrical and conspicuously large distolateral sclerites and with strongly asymmetrical assemblage of median sclerotized structures, distolateral sclerites broadly separated, each with long subtriangular and curved apical region; parameres broad and their apices not projecting beyond aedeagal apex, each paramere with apical-subapical group of 23 barely discernible setae.
</paragraph>
<paragraph id="8BF14E46FFD05020FF73FEEBFE5B4532" blockId="32.[151,1437,151,645]" box="[199,414,295,321]" pageId="32" pageNumber="33">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD05020FF73FE86FC884516" box="[199,845,330,357]" pageId="32" pageNumber="33" type="distribution">
<paragraph id="8BF14E46FFD05020FF73FE86FC884516" blockId="32.[151,1437,151,645]" box="[199,845,330,357]" pageId="32" pageNumber="33">
<emphasis id="B93A9254FFD05020FF73FE86FEA44517" bold="true" box="[199,353,330,356]" pageId="32" pageNumber="33">Distribution.</emphasis>
Northernmost
<collectingCountry id="F3590ED6FFD05020FDBEFE87FD544516" box="[522,657,331,357]" name="Madagascar" pageId="32" pageNumber="33">Madagascar</collectingCountry>
(Diana Region).
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD05020FF73FEA3FC8346F6" pageId="32" pageNumber="33" type="discussion">
<paragraph id="8BF14E46FFD05020FF73FEA3FC7345A2" blockId="32.[151,1437,151,645]" pageId="32" pageNumber="33">
<emphasis id="B93A9254FFD05020FF73FEA3FEFE45FA" bold="true" box="[199,315,367,393]" pageId="32" pageNumber="33">Remarks.</emphasis>
Males of
<taxonomicName id="4C4E35C5FFD05020FE1FFEA3FD0D45FA" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[427,712,367,393]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="diegoi">
<emphasis id="B93A9254FFD05020FE1FFEA3FD0D45FA" box="[427,712,367,393]" italics="true" pageId="32" pageNumber="33">Anhoraeomorphus diegoi</emphasis>
</taxonomicName>
have the hind tibiae similarly modified as those in
<taxonomicName id="4C4E35C5FFD05020FAB0FEA3FA5945FB" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[1284,1436,367,393]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="anosyensis">
<emphasis id="B93A9254FFD05020FAB0FEA3FA5945FB" box="[1284,1436,367,393]" italics="true" pageId="32" pageNumber="33">A. anosyensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4E35C5FFD05020FF73FE5FFEB245DF" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[199,375,403,429]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="roussettensis">
<emphasis id="B93A9254FFD05020FF73FE5FFEB245DF" box="[199,375,403,429]" italics="true" pageId="32" pageNumber="33">A. roussettensis</emphasis>
</taxonomicName>
. However, the body of
<taxonomicName id="4C4E35C5FFD05020FD35FE5FFD2245DE" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[641,743,403,429]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="diegoi">
<emphasis id="B93A9254FFD05020FD35FE5FFD2245DE" box="[641,743,403,429]" italics="true" pageId="32" pageNumber="33">A. diegoi</emphasis>
</taxonomicName>
is slenderer and the aedeagus has the distolateral endophallic sclerites much broader than those in
<taxonomicName id="4C4E35C5FFD05020FD86FE7BFD0F45A3" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[562,714,439,465]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="anosyensis">
<emphasis id="B93A9254FFD05020FD86FE7BFD0F45A3" box="[562,714,439,465]" italics="true" pageId="32" pageNumber="33">A. anosyensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4E35C5FFD05020FCB5FE7BFC7545A3" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[769,944,439,465]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="roussettensis">
<emphasis id="B93A9254FFD05020FCB5FE7BFC7545A3" box="[769,944,439,465]" italics="true" pageId="32" pageNumber="33">A. roussettensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF14E46FFD05020FF73FE17FC8346F6" blockId="32.[151,1437,151,645]" pageId="32" pageNumber="33">
<materialsCitation id="3B26441BFFD05020FF73FE17FC8746F6" collectionCode="NHMW" collectorName="Charles Alluaud" location="Antsiranana" pageId="32" pageNumber="33" specimenCount="6" stateProvince="Antsiranana" typeStatus="holotype">
<bibRefCitation id="EFDF33B7FFD05020FF73FE17FEA04586" author="Franz, H." box="[199,357,475,501]" pageId="32" pageNumber="33" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986b)</bibRefCitation>
mentions
<specimenCount id="9D4885CFFFD05020FE62FE17FD494586" box="[470,652,475,501]" count="3" pageId="32" pageNumber="33" type="generic">three specimens</specimenCount>
, all males, included in the type series. The
<typeStatus id="54F5F0E4FFD05020FBEAFE17FB054586" box="[1118,1216,475,501]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
comes from Diego-Suarez (currently
<collectingRegion id="498A80A4FFD05020FED0FE33FE28466A" box="[356,493,511,537]" country="Madagascar" name="Antsiranana" pageId="32" pageNumber="33">Antsiranana</collectingRegion>
) and was collected by
<collectorName id="26BB2B90FFD05020FD4FFE33FC73466A" box="[763,950,511,537]" pageId="32" pageNumber="33">Charles Alluaud</collectorName>
,
<specimenCount id="9D4885CFFFD05020FC70FE33FBA6466A" box="[964,1123,511,537]" count="2" pageId="32" pageNumber="33" type="generic" typeStatus="paratype">two paratypes</specimenCount>
are from Léon Fairmaires collection and their collecting site is unknown. The
<typeStatus id="54F5F0E4FFD05020FD57FDEFFC80464E" box="[739,837,547,573]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
and
<specimenCount id="9D4885CFFFD05020FCCAFDEFFBCB464E" box="[894,1038,547,573]" count="1" pageId="32" pageNumber="33" type="generic" typeStatus="paratype">one paratype</specimenCount>
are deposited in Muséum National d'Histoire Naturelle, Paris and were not available for my study. The examined
<typeStatus id="54F5F0E4FFD05020FB9CFD8BFB4D4612" box="[1064,1160,583,609]" pageId="32" pageNumber="33" type="paratype">paratype</typeStatus>
male (
<collectionCode id="ED5FD683FFD05020FB6EFD8BFAFC4612" box="[1242,1337,583,609]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="32" pageNumber="33" type="Museum">NHMW</collectionCode>
) has the aedeagus matching the illustration in the original description
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,417 @@
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<mods:title id="7F902C672156B2422FF3F0EF622E036B">The curse of Horaeomorphus (almost) lifted. Revision of misplaced species from Madagascar, Comoros and Mascarenes (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<subSubSection id="C3541DCDFFF85008FF23FDEEFCC2464E" box="[151,775,546,573]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="8BF14E46FFF85008FF23FDEEFCC2464E" blockId="8.[151,775,546,609]" box="[151,775,546,573]" pageId="8" pageNumber="9">
<heading id="D0B9F92AFFF85008FF23FDEEFCC2464E" bold="true" box="[151,775,546,573]" fontSize="11" level="2" pageId="8" pageNumber="9" reason="4">
<emphasis id="B93A9254FFF85008FF23FDEEFCC2464E" bold="true" box="[151,775,546,573]" pageId="8" pageNumber="9">
<taxonomicName id="4C4E35C5FFF85008FF23FDEEFD4B464F" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[151,654,546,573]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="ankaratranus" status="comb. nov.">
<emphasis id="B93A9254FFF85008FF23FDEEFDED464E" bold="true" box="[151,552,546,573]" italics="true" pageId="8" pageNumber="9">Anhoraeomorphus ankaratranus</emphasis>
(Franz)
</taxonomicName>
,
<taxonomicNameLabel id="A2092F2FFFF85008FD28FDEFFCC2464E" box="[668,775,547,573]" pageId="8" pageNumber="9" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFF85008FF23FD8BFEDC4612" box="[151,281,583,609]" pageId="8" pageNumber="9" type="description">
<paragraph id="8BF14E46FFF85008FF23FD8BFEDC4612" blockId="8.[151,775,546,609]" box="[151,281,583,609]" pageId="8" pageNumber="9">
(
<figureCitation id="137552C3FFF85008FF2BFD8BFED54612" box="[159,272,583,609]" captionStart-0="FIGURES 18" captionStart-1="FIGURES 911" captionStart-2="FIGURES 1219" captionStartId-0="9.[151,264,1405,1429]" captionStartId-1="10.[151,264,705,729]" captionStartId-2="10.[151,264,1945,1969]" captionTargetBox-0="[151,1429,181,1369]" captionTargetBox-1="[183,1404,181,681]" captionTargetBox-2="[199,1390,902,1907]" captionTargetId-0="figure-16@9.[151,1435,181,1381]" captionTargetId-1="figure-28@10.[183,1404,181,681]" captionTargetId-2="figure-107@10.[189,1397,898,1922]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="10" captionText-0="FIGURES 18. Anhoraeomorphus ankaratranus (Franz), holotype. Habitus in dorsal view (1); head in anterodorsal view (2); head and prothorax in lateral view (only bristles on head are shown) (3); antenna (4); aedeagus in ventral (5, 7) and lateral (6, 8) views." captionText-1="FIGURES 911. Anhoraeomorphus ankaratranus (Franz), holotype. Head, prothorax and elytral base in dorsal view (9); head, prothorax and anterior region of mesothorax in ventral view (10); pterothorax in ventral view (11). Abbreviations: abp, antebasal pit; bef, basal elytral fovea; gp, gular plate; gs, gular suture; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; pcr, procoxal rest; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum." captionText-2="FIGURES 1219. Anhoraeomorphus ankaratranus (Franz), holotype of Horaeomorphus manjakotompoi Franz. Habitus in dorsal view (12); head in anterodorsal view (13); head and prothorax in lateral view (only bristles on head are shown) (14); antenna (15); aedeagus in ventral (16, 18) and lateral (17, 19) views." figureDoi-0="http://doi.org/10.5281/zenodo.13745916" figureDoi-1="http://doi.org/10.5281/zenodo.13745921" figureDoi-2="http://doi.org/10.5281/zenodo.13745923" httpUri-0="https://zenodo.org/record/13745916/files/figure.png" httpUri-1="https://zenodo.org/record/13745921/files/figure.png" httpUri-2="https://zenodo.org/record/13745923/files/figure.png" pageId="8" pageNumber="9">Figs 119</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFF85008FF23FD47FD3246B1" pageId="8" pageNumber="9" type="reference_group">
<paragraph id="8BF14E46FFF85008FF23FD47FD5646D0" blockId="8.[151,759,651,706]" box="[151,659,651,675]" pageId="8" pageNumber="9">
<treatmentCitationGroup id="AB5E6968FFF85008FF23FD47FD5646D0" box="[151,659,651,675]" pageId="8" pageNumber="9">
<treatmentCitation id="0AEF6857FFF85008FF23FD47FD4A46D0" author="Franz, H." box="[151,655,651,675]" page="164" pageId="8" pageNumber="9" year="1986">
<taxonomicName id="4C4E35C5FFF85008FF23FD47FD4A46D0" ID-CoL="3MNJ7" authority="Franz, 1986 b: 164" authorityName="Franz" authorityPageNumber="164" authorityYear="1986" box="[151,655,651,675]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="ankaratranus">
<emphasis id="B93A9254FFF85008FF23FD47FE0846D0" box="[151,461,651,675]" italics="true" pageId="8" pageNumber="9">Horaeomorphus ankaratranus</emphasis>
<bibRefCitation id="EFDF33B7FFF85008FE66FD40FD4A46D0" author="Franz, H." box="[466,655,651,675]" pageId="8" pageNumber="9" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 164</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF14E46FFF85008FF23FD66FD3246B1" blockId="8.[151,759,651,706]" box="[151,759,681,706]" pageId="8" pageNumber="9">
<treatmentCitationGroup id="AB5E6968FFF85008FF23FD66FD3246B1" box="[151,759,681,706]" pageId="8" pageNumber="9">
<treatmentCitation id="0AEF6857FFF85008FF23FD66FD6146B1" author="Franz, H." box="[151,676,682,706]" page="173" pageId="8" pageNumber="9" year="1986">
<taxonomicName id="4C4E35C5FFF85008FF23FD66FD6146B1" ID-CoL="3MNKM" authority="Franz, 1986 b: 173" authorityName="Franz" authorityPageNumber="173" authorityYear="1986" box="[151,676,682,706]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="manjakotompoi">
<emphasis id="B93A9254FFF85008FF23FD66FE2746B1" box="[151,482,682,706]" italics="true" pageId="8" pageNumber="9">Horaeomorphus manjakotompoi</emphasis>
<bibRefCitation id="EFDF33B7FFF85008FE5CFD66FD6146B1" author="Franz, H." box="[488,676,682,706]" pageId="8" pageNumber="9" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 173</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<emphasis id="B93A9254FFF85008FD19FD65FD3246B1" bold="true" box="[685,759,681,706]" pageId="8" pageNumber="9">Syn. n.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFF85008FF23FD3CFD2440C9" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="8BF14E46FFF85008FF23FD3CFC154791" blockId="8.[151,1437,751,1931]" pageId="8" pageNumber="9">
<emphasis id="B93A9254FFF85008FF23FD3CFE744779" bold="true" box="[151,433,752,778]" pageId="8" pageNumber="9">Type material studied.</emphasis>
<materialsCitation id="3B26441BFFF85008FE0BFD3CFC364721" collectionCode="NHMW" collectorName="H. Franz" country="Madagascar" determinerName="H. Franz" location="Madagascar" pageId="8" pageNumber="9" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B93A9254FFF85008FE0BFD3CFC204779" bold="true" box="[447,997,751,778]" pageId="8" pageNumber="9">
<typeStatus id="54F5F0E4FFF85008FE0BFD3CFDE94779" box="[447,556,752,778]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C4E35C5FFF85008FDEBFD3CFCE24779" authorityName="Franz" authorityYear="1986" box="[607,807,751,778]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="ankaratranus">
<emphasis id="B93A9254FFF85008FDEBFD3CFCE24779" bold="true" box="[607,807,751,778]" italics="true" pageId="8" pageNumber="9">H. ankaratranus</emphasis>
</taxonomicName>
(
<collectingCountry id="F3590ED6FFF85008FC89FD3CFC1D4779" box="[829,984,752,778]" name="Madagascar" pageId="8" pageNumber="9">Madagascar</collectingCountry>
):
</emphasis>
<specimenCount id="9D4885CFFFF85008FC47FD3DFBCF4778" box="[1011,1034,753,779]" count="1" pageId="8" pageNumber="9" type="male"></specimenCount>
, three labels: “Massif de lAn- / karatra,Madagaskar / lg.
<collectorName id="26BB2B90FFF85008FE01FCD8FDD7475D" box="[437,530,788,814]" pageId="8" pageNumber="9">H.Franz</collectorName>
1969” [white, printed], “
<taxonomicName id="4C4E35C5FFF85008FCF8FCD8FBC6475D" box="[844,1027,788,814]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/ ankaratranus m. / det.
<determinerName id="7E2DA282FFF85008FA80FCD8FA57475D" box="[1332,1426,788,814]" pageId="8" pageNumber="9">H.Franz</determinerName>
” [yellowish, handwritten and printed], “Typus” [red, handwritten] (
<collectionCode id="ED5FD683FFF85008FC3CFCF4FC2E4721" box="[904,1003,824,850]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="9" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B26441BFFF85008FC4BFCF4FC914705" collectionCode="NHMW" collectorName="H. Franz" country="Madagascar" determinerName="H. Franz" location="Madagascar" pageId="8" pageNumber="9" specimenCount="2" typeStatus="paralectotype">
<emphasis id="B93A9254FFF85008FC4BFCF4FB6A4721" bold="true" box="[1023,1199,824,850]" pageId="8" pageNumber="9">
<typeStatus id="54F5F0E4FFF85008FC4BFCF4FB6A4721" box="[1023,1199,824,850]" pageId="8" pageNumber="9" type="paralectotype">Paralectotypes</typeStatus>
</emphasis>
: 26 exx., data as for holotype, except each with yellow “
<typeStatus id="54F5F0E4FFF85008FD96FC90FD504705" box="[546,661,860,886]" pageId="8" pageNumber="9" type="paratype">Paratypus</typeStatus>
” label (
<collectionCode id="ED5FD683FFF85008FD5EFC90FC884705" box="[746,845,860,886]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="9" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B26441BFFF85008FCE9FC90FC094791" collectionCode="R, VII, NHMW" country="Madagascar" determinerName="H. Franz" location="Manjakotompo" pageId="8" pageNumber="9" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
<emphasis id="B93A9254FFF85008FCE9FC90FABF4705" bold="true" box="[861,1402,859,886]" pageId="8" pageNumber="9">
<typeStatus id="54F5F0E4FFF85008FCE9FC90FC0F4705" box="[861,970,860,886]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C4E35C5FFF85008FC59FC90FB014705" box="[1005,1220,859,886]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="manjakotompoi">
<emphasis id="B93A9254FFF85008FC59FC90FB014705" bold="true" box="[1005,1220,859,886]" italics="true" pageId="8" pageNumber="9">H. manjakotompoi</emphasis>
</taxonomicName>
(
<collectingCountry id="F3590ED6FFF85008FB66FC90FAA84705" box="[1234,1389,860,886]" name="Madagascar" pageId="8" pageNumber="9">Madagascar</collectingCountry>
):
</emphasis>
<specimenCount id="9D4885CFFFF85008FA34FC91FA524704" box="[1408,1431,861,887]" count="1" pageId="8" pageNumber="9" type="male"></specimenCount>
, five labels: “
<location id="8E91189DFFF85008FE94FC4CFE0B47E9" LSID="urn:lsid:plazi:treatment:03E7FF50FFF8500BFF23FDEEFCA846DA:8E91189DFFF85008FE94FC4CFE0B47E9" box="[288,462,896,922]" country="Madagascar" name="Manjakotompo" pageId="8" pageNumber="9">Manjakotompo</location>
/ 6.vii.56 (E.
<collectionCode id="ED5FD683FFF85008FDEEFC4CFDAA47E9" box="[602,623,896,922]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="8" pageNumber="9">R</collectionCode>
{or K})” [brownish, handwritten], “
<specimenCount id="9D4885CFFFF85008FBB5FC4CFBDD47E9" box="[1025,1048,896,922]" count="1" pageId="8" pageNumber="9" type="male"></specimenCount>
,” [white, printed],
<taxonomicName id="4C4E35C5FFF85008FB5CFC4CFA5947E9" box="[1256,1436,896,922]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/ manjakotompoi m. / det.
<determinerName id="7E2DA282FFF85008FE61FC68FDF747CD" box="[469,562,932,958]" pageId="8" pageNumber="9">H.Franz</determinerName>
” [yellowish, handwritten and printed], “Manjakotompo /
<collectionCode id="ED5FD683FFF85008FB5EFC68FAD647CC" box="[1258,1299,932,959]" pageId="8" pageNumber="9">VII</collectionCode>
.56 (E.
<collectionCode id="ED5FD683FFF85008FAD7FC69FAB347CC" box="[1379,1398,933,959]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="8" pageNumber="9">R</collectionCode>
.) /
<collectingCountry id="F3590ED6FFF85008FF23FC04FEE44791" box="[151,289,968,994]" name="Madagascar" pageId="8" pageNumber="9">Madagascar</collectingCountry>
” [white, handwritten], “Typus” [red, handwritten] (
<collectionCode id="ED5FD683FFF85008FCD5FC04FC014791" box="[865,964,968,994]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="9" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF14E46FFF85008FF73FC20FD2440C9" blockId="8.[151,1437,751,1931]" pageId="8" pageNumber="9">
<emphasis id="B93A9254FFF85008FF73FC20FE634075" bold="true" box="[199,422,1004,1030]" pageId="8" pageNumber="9">Revised diagnosis.</emphasis>
Body (
<figureCitation id="137552C3FFF85008FDB0FC20FD8C4075" box="[516,585,1004,1030]" captionStart="FIGURES 18" captionStartId="9.[151,264,1405,1429]" captionTargetBox="[151,1429,181,1369]" captionTargetId="figure-16@9.[151,1435,181,1381]" captionTargetPageId="9" captionText="FIGURES 18. Anhoraeomorphus ankaratranus (Franz), holotype. Habitus in dorsal view (1); head in anterodorsal view (2); head and prothorax in lateral view (only bristles on head are shown) (3); antenna (4); aedeagus in ventral (5, 7) and lateral (6, 8) views." figureDoi="http://doi.org/10.5281/zenodo.13745916" httpUri="https://zenodo.org/record/13745916/files/figure.png" pageId="8" pageNumber="9">Fig. 1</figureCitation>
) stout; head (
<figureCitation id="137552C3FFF85008FD41FC20FCF84075" box="[757,829,1004,1030]" captionStart="FIGURES 18" captionStartId="9.[151,264,1405,1429]" captionTargetBox="[151,1429,181,1369]" captionTargetId="figure-16@9.[151,1435,181,1381]" captionTargetPageId="9" captionText="FIGURES 18. Anhoraeomorphus ankaratranus (Franz), holotype. Habitus in dorsal view (1); head in anterodorsal view (2); head and prothorax in lateral view (only bristles on head are shown) (3); antenna (4); aedeagus in ventral (5, 7) and lateral (6, 8) views." figureDoi="http://doi.org/10.5281/zenodo.13745916" httpUri="https://zenodo.org/record/13745916/files/figure.png" pageId="8" pageNumber="9">Fig. 2</figureCitation>
) about as long as wide, with vertex subconical, in anterodorsal view its posterior margin subtriangular; pronotum (
<figureCitation id="137552C3FFF85008FCDAFBDCFC774059" box="[878,946,1040,1066]" captionStart="FIGURES 911" captionStartId="10.[151,264,705,729]" captionTargetBox="[183,1404,181,681]" captionTargetId="figure-28@10.[183,1404,181,681]" captionTargetPageId="10" captionText="FIGURES 911. Anhoraeomorphus ankaratranus (Franz), holotype. Head, prothorax and elytral base in dorsal view (9); head, prothorax and anterior region of mesothorax in ventral view (10); pterothorax in ventral view (11). Abbreviations: abp, antebasal pit; bef, basal elytral fovea; gp, gular plate; gs, gular suture; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; pcr, procoxal rest; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13745921" httpUri="https://zenodo.org/record/13745921/files/figure.png" pageId="8" pageNumber="9">Fig. 9</figureCitation>
) subtrapezoidal, broadest at base, with two pairs of distinct antebasal pits; antennomeres 310 not elongate; all femora similarly slender; metatibiae in males unmodified; aedeagus (
<figureCitation id="137552C3FFF85008FE28FB94FE3A4000" box="[412,511,1112,1139]" captionStart="FIGURES 18" captionStartId="9.[151,264,1405,1429]" captionTargetBox="[151,1429,181,1369]" captionTargetId="figure-16@9.[151,1435,181,1381]" captionTargetPageId="9" captionText="FIGURES 18. Anhoraeomorphus ankaratranus (Franz), holotype. Habitus in dorsal view (1); head in anterodorsal view (2); head and prothorax in lateral view (only bristles on head are shown) (3); antenna (4); aedeagus in ventral (5, 7) and lateral (6, 8) views." figureDoi="http://doi.org/10.5281/zenodo.13745916" httpUri="https://zenodo.org/record/13745916/files/figure.png" pageId="8" pageNumber="9">Figs 58</figureCitation>
) in ventral view broadest near base, with apical region of dorsal wall abruptly bent dorsad, distolateral endophallic sclerites widely separated and with rounded apices, each paramere with a row of subapical setae distributed on more than distal third.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFF85009FF73FB08FD894319" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="description">
<paragraph id="8BF14E46FFF85008FF73FB08FB434170" blockId="8.[151,1437,751,1931]" pageId="8" pageNumber="9">
<emphasis id="B93A9254FFF85008FF73FB08FEB040AD" bold="true" box="[199,373,1220,1246]" pageId="8" pageNumber="9">Redescription.</emphasis>
Body of male (
<figureCitation id="137552C3FFF85008FDA9FB08FDA740AD" box="[541,610,1220,1246]" captionStart="FIGURES 18" captionStartId="9.[151,264,1405,1429]" captionTargetBox="[151,1429,181,1369]" captionTargetId="figure-16@9.[151,1435,181,1381]" captionTargetPageId="9" captionText="FIGURES 18. Anhoraeomorphus ankaratranus (Franz), holotype. Habitus in dorsal view (1); head in anterodorsal view (2); head and prothorax in lateral view (only bristles on head are shown) (3); antenna (4); aedeagus in ventral (5, 7) and lateral (6, 8) views." figureDoi="http://doi.org/10.5281/zenodo.13745916" httpUri="https://zenodo.org/record/13745916/files/figure.png" pageId="8" pageNumber="9">Figs 1</figureCitation>
,
<figureCitation id="137552C3FFF85008FDDBFB08FD4940AD" box="[623,652,1220,1246]" captionStart="FIGURES 1219" captionStartId="10.[151,264,1945,1969]" captionTargetBox="[199,1390,902,1907]" captionTargetId="figure-107@10.[189,1397,898,1922]" captionTargetPageId="10" captionText="FIGURES 1219. Anhoraeomorphus ankaratranus (Franz), holotype of Horaeomorphus manjakotompoi Franz. Habitus in dorsal view (12); head in anterodorsal view (13); head and prothorax in lateral view (only bristles on head are shown) (14); antenna (15); aedeagus in ventral (16, 18) and lateral (17, 19) views." figureDoi="http://doi.org/10.5281/zenodo.13745923" httpUri="https://zenodo.org/record/13745923/files/figure.png" pageId="8" pageNumber="9">12</figureCitation>
) moderately dark to strongly dark brown with reddish hue, appendages indistinctly lighter, setae lighter than cuticle, yellowish to light brown; BL
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.
</paragraph>
<paragraph id="8BF14E46FFF85008FF73FAC0FB784251" blockId="8.[151,1437,751,1931]" pageId="8" pageNumber="9">
Head (
<figureCitation id="137552C3FFF85008FEA5FAC0FEB34155" box="[273,374,1292,1318]" captionStart="FIGURES 18" captionStartId="9.[151,264,1405,1429]" captionTargetBox="[151,1429,181,1369]" captionTargetId="figure-16@9.[151,1435,181,1381]" captionTargetPageId="9" captionText="FIGURES 18. Anhoraeomorphus ankaratranus (Franz), holotype. Habitus in dorsal view (1); head in anterodorsal view (2); head and prothorax in lateral view (only bristles on head are shown) (3); antenna (4); aedeagus in ventral (5, 7) and lateral (6, 8) views." figureDoi="http://doi.org/10.5281/zenodo.13745916" httpUri="https://zenodo.org/record/13745916/files/figure.png" pageId="8" pageNumber="9">Figs 23</figureCitation>
,
<figureCitation id="137552C3FFF85008FE35FAC0FE7E4155" box="[385,443,1292,1318]" captionStart="FIGURES 911" captionStartId="10.[151,264,705,729]" captionTargetBox="[183,1404,181,681]" captionTargetId="figure-28@10.[183,1404,181,681]" captionTargetPageId="10" captionText="FIGURES 911. Anhoraeomorphus ankaratranus (Franz), holotype. Head, prothorax and elytral base in dorsal view (9); head, prothorax and anterior region of mesothorax in ventral view (10); pterothorax in ventral view (11). Abbreviations: abp, antebasal pit; bef, basal elytral fovea; gp, gular plate; gs, gular suture; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; pcr, procoxal rest; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13745921" httpUri="https://zenodo.org/record/13745921/files/figure.png" pageId="8" pageNumber="9">910</figureCitation>
,
<figureCitation id="137552C3FFF85008FE73FAC0FDCB4155" box="[455,526,1292,1318]" captionStart="FIGURES 1219" captionStartId="10.[151,264,1945,1969]" captionTargetBox="[199,1390,902,1907]" captionTargetId="figure-107@10.[189,1397,898,1922]" captionTargetPageId="10" captionText="FIGURES 1219. Anhoraeomorphus ankaratranus (Franz), holotype of Horaeomorphus manjakotompoi Franz. Habitus in dorsal view (12); head in anterodorsal view (13); head and prothorax in lateral view (only bristles on head are shown) (14); antenna (15); aedeagus in ventral (16, 18) and lateral (17, 19) views." figureDoi="http://doi.org/10.5281/zenodo.13745923" httpUri="https://zenodo.org/record/13745923/files/figure.png" pageId="8" pageNumber="9">1314</figureCitation>
) in anterodorsal view almost triangular, broadest at eyes, HL
<quantity id="4CB6E3A3FFF85008FB0AFAC0FAA64154" box="[1214,1379,1292,1319]" metricMagnitude="-4" metricUnit="m" metricValue="3.15" metricValueMax="3.3" metricValueMin="3.0" pageId="8" pageNumber="9" unit="mm" value="0.315" valueMax="0.33" valueMin="0.3">0.300.33 mm</quantity>
, HW
<quantity id="4CB6E3A3FFF85008FF23FAFCFEF94138" box="[151,316,1328,1355]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" metricValueMax="3.0" metricValueMin="2.8" pageId="8" pageNumber="9" unit="mm" value="0.29" valueMax="0.3" valueMin="0.28">0.280.30 mm</quantity>
; temple in lateral view (
<figureCitation id="137552C3FFF85008FDF1FAFCFD434139" box="[581,646,1328,1354]" captionStart="FIGURES 18" captionStartId="9.[151,264,1405,1429]" captionTargetBox="[151,1429,181,1369]" captionTargetId="figure-16@9.[151,1435,181,1381]" captionTargetPageId="9" captionText="FIGURES 18. Anhoraeomorphus ankaratranus (Franz), holotype. Habitus in dorsal view (1); head in anterodorsal view (2); head and prothorax in lateral view (only bristles on head are shown) (3); antenna (4); aedeagus in ventral (5, 7) and lateral (6, 8) views." figureDoi="http://doi.org/10.5281/zenodo.13745916" httpUri="https://zenodo.org/record/13745916/files/figure.png" pageId="8" pageNumber="9">Fig. 3</figureCitation>
) subequal to length of eye; vertex and frons confluent, together weakly convex and slightly elongate; vertex subconical, with its posterior margin nearly triangular; each eye moderately large and weakly convex, with small posteromedian emargination, weakly projecting from head silhouette. Punctures on vertex fine, inconspicuous; setae short, sparse and suberect; posterior region of frons and entire vertex densely covered with thick bristles directed posteriorly. Antenna (
<figureCitation id="137552C3FFF85008FCA1FA0CFC9841A9" box="[789,861,1472,1498]" captionStart="FIGURES 18" captionStartId="9.[151,264,1405,1429]" captionTargetBox="[151,1429,181,1369]" captionTargetId="figure-16@9.[151,1435,181,1381]" captionTargetPageId="9" captionText="FIGURES 18. Anhoraeomorphus ankaratranus (Franz), holotype. Habitus in dorsal view (1); head in anterodorsal view (2); head and prothorax in lateral view (only bristles on head are shown) (3); antenna (4); aedeagus in ventral (5, 7) and lateral (6, 8) views." figureDoi="http://doi.org/10.5281/zenodo.13745916" httpUri="https://zenodo.org/record/13745916/files/figure.png" pageId="8" pageNumber="9">Figs 4</figureCitation>
,
<figureCitation id="137552C3FFF85008FCDCFA0CFC4041A8" box="[872,901,1472,1499]" captionStart="FIGURES 1219" captionStartId="10.[151,264,1945,1969]" captionTargetBox="[199,1390,902,1907]" captionTargetId="figure-107@10.[189,1397,898,1922]" captionTargetPageId="10" captionText="FIGURES 1219. Anhoraeomorphus ankaratranus (Franz), holotype of Horaeomorphus manjakotompoi Franz. Habitus in dorsal view (12); head in anterodorsal view (13); head and prothorax in lateral view (only bristles on head are shown) (14); antenna (15); aedeagus in ventral (16, 18) and lateral (17, 19) views." figureDoi="http://doi.org/10.5281/zenodo.13745923" httpUri="https://zenodo.org/record/13745923/files/figure.png" pageId="8" pageNumber="9">15</figureCitation>
) slender but short, slightly shorter than half BL, AnL
<quantity id="4CB6E3A3FFF85008FF65FA28FEB3418C" box="[209,374,1508,1535]" metricMagnitude="-4" metricUnit="m" metricValue="6.65" metricValueMax="6.8" metricValueMin="6.5" pageId="8" pageNumber="9" unit="mm" value="0.665" valueMax="0.68" valueMin="0.65">0.650.68 mm</quantity>
, antennomeres 1 and 2 each weakly elongate, 310 each about as long as broad or indistinctly transverse; 11 shorter than 9 and 10 combined, as wide as 10, about 1.8 times as long as broad.
</paragraph>
<paragraph id="8BF14E46FFF85008FF73F9E1FCE842A5" blockId="8.[151,1437,751,1931]" pageId="8" pageNumber="9">
Pronotum (
<figureCitation id="137552C3FFF85008FEFCF9E0FE4B4235" box="[328,398,1580,1606]" captionStart="FIGURES 911" captionStartId="10.[151,264,705,729]" captionTargetBox="[183,1404,181,681]" captionTargetId="figure-28@10.[183,1404,181,681]" captionTargetPageId="10" captionText="FIGURES 911. Anhoraeomorphus ankaratranus (Franz), holotype. Head, prothorax and elytral base in dorsal view (9); head, prothorax and anterior region of mesothorax in ventral view (10); pterothorax in ventral view (11). Abbreviations: abp, antebasal pit; bef, basal elytral fovea; gp, gular plate; gs, gular suture; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; pcr, procoxal rest; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13745921" httpUri="https://zenodo.org/record/13745921/files/figure.png" pageId="8" pageNumber="9">Fig. 9</figureCitation>
) subtrapezoidal, broadest at base; PL
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, PW
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. Anterior margin weakly arcuate, lateral margins weakly convex and strongly diverging posteriorly; posterior corners blunt; posterior margin indistinctly convex. Base with two pairs of round and deep pits, and with barely discernible, extremely short vestiges of sublateral carinae. Disc with fine, inconspicuous punctures; setae sparse, short and suberect, lateral and lateroventral surface of pronotum with dense thick bristles.
</paragraph>
<paragraph id="8BF14E46FFF85008FF73F92CFC414331" blockId="8.[151,1437,751,1931]" pageId="8" pageNumber="9">
Elytra (
<figureCitation id="137552C3FFF85008FEA9F92CFE9A4289" box="[285,351,1760,1786]" captionStart="FIGURES 18" captionStartId="9.[151,264,1405,1429]" captionTargetBox="[151,1429,181,1369]" captionTargetId="figure-16@9.[151,1435,181,1381]" captionTargetPageId="9" captionText="FIGURES 18. Anhoraeomorphus ankaratranus (Franz), holotype. Habitus in dorsal view (1); head in anterodorsal view (2); head and prothorax in lateral view (only bristles on head are shown) (3); antenna (4); aedeagus in ventral (5, 7) and lateral (6, 8) views." figureDoi="http://doi.org/10.5281/zenodo.13745916" httpUri="https://zenodo.org/record/13745916/files/figure.png" pageId="8" pageNumber="9">Fig. 1</figureCitation>
) together oval, broadest slightly in front of middle; EL
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, EW
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, EI 1.251.32; humeral calli weakly elevated, basal elytral foveae (
<figureCitation id="137552C3FFF85008FCE7F8C8FC50436D" box="[851,917,1796,1822]" captionStart="FIGURES 911" captionStartId="10.[151,264,705,729]" captionTargetBox="[183,1404,181,681]" captionTargetId="figure-28@10.[183,1404,181,681]" captionTargetPageId="10" captionText="FIGURES 911. Anhoraeomorphus ankaratranus (Franz), holotype. Head, prothorax and elytral base in dorsal view (9); head, prothorax and anterior region of mesothorax in ventral view (10); pterothorax in ventral view (11). Abbreviations: abp, antebasal pit; bef, basal elytral fovea; gp, gular plate; gs, gular suture; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; pcr, procoxal rest; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13745921" httpUri="https://zenodo.org/record/13745921/files/figure.png" pageId="8" pageNumber="9">Fig. 9</figureCitation>
) vestigial, barely discernible even under SEM. Punctures fine and inconspicuous; setae sparse, short and suberect.
</paragraph>
<paragraph id="8BF14E46FFF85008FF73F880FB934315" blockId="8.[151,1437,751,1931]" box="[199,1110,1868,1895]" pageId="8" pageNumber="9">
Metaventrite (
<figureCitation id="137552C3FFF85008FED2F880FE704315" box="[358,437,1868,1894]" captionStart="FIGURES 911" captionStartId="10.[151,264,705,729]" captionTargetBox="[183,1404,181,681]" captionTargetId="figure-28@10.[183,1404,181,681]" captionTargetPageId="10" captionText="FIGURES 911. Anhoraeomorphus ankaratranus (Franz), holotype. Head, prothorax and elytral base in dorsal view (9); head, prothorax and anterior region of mesothorax in ventral view (10); pterothorax in ventral view (11). Abbreviations: abp, antebasal pit; bef, basal elytral fovea; gp, gular plate; gs, gular suture; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; pcr, procoxal rest; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13745921" httpUri="https://zenodo.org/record/13745921/files/figure.png" pageId="8" pageNumber="9">Fig. 11</figureCitation>
) with setae similar to those on elytra but nearly impunctate.
</paragraph>
<paragraph id="8BF14E46FFF85008FF73F8BDFAAD43F9" blockId="8.[151,1437,751,1931]" box="[199,1384,1904,1931]" pageId="8" pageNumber="9">Legs moderately long, slender, unmodified except slightly recurved tibiae (mesotibiae most distinctly so).</paragraph>
<caption id="DF311ECEFFF95009FF23FAB1FF3041AE" ID-DOI="http://doi.org/10.5281/zenodo.13745916" ID-Zenodo-Dep="13745916" httpUri="https://zenodo.org/record/13745916/files/figure.png" pageId="9" pageNumber="10" startId="9.[151,264,1405,1429]" targetBox="[151,1429,181,1369]" targetPageId="9" targetType="figure">
<paragraph id="8BF14E46FFF95009FF23FAB1FF3041AE" blockId="9.[151,1437,1405,1502]" pageId="9" pageNumber="10">
<emphasis id="B93A9254FFF95009FF23FAB1FEFE41E6" bold="true" box="[151,315,1405,1429]" pageId="9" pageNumber="10">FIGURES 18.</emphasis>
<taxonomicName id="4C4E35C5FFF95009FEF6FAB1FD2441E6" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[322,737,1405,1429]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ankaratranus">
<emphasis id="B93A9254FFF95009FEF6FAB1FD4B41E6" box="[322,654,1405,1429]" italics="true" pageId="9" pageNumber="10">Anhoraeomorphus ankaratranus</emphasis>
(Franz)
</taxonomicName>
, holotype. Habitus in dorsal view (
<emphasis id="B93A9254FFF95009FBF3FAB1FB9141E6" bold="true" box="[1095,1108,1405,1429]" pageId="9" pageNumber="10">1</emphasis>
); head in anterodorsal view (
<emphasis id="B93A9254FFF95009FA34FAB1FA4841E6" bold="true" box="[1408,1421,1405,1429]" pageId="9" pageNumber="10">2</emphasis>
); head and prothorax in lateral view (only bristles on head are shown) (
<emphasis id="B93A9254FFF95009FCD4FA6DFCA841CA" bold="true" box="[864,877,1441,1465]" pageId="9" pageNumber="10">3</emphasis>
); antenna (
<emphasis id="B93A9254FFF95009FC55FA6DFC2B41CA" bold="true" box="[993,1006,1441,1465]" pageId="9" pageNumber="10">4</emphasis>
); aedeagus in ventral (
<emphasis id="B93A9254FFF95009FB6DFA6DFAC541CA" bold="true" box="[1241,1280,1441,1465]" pageId="9" pageNumber="10">5, 7</emphasis>
) and lateral (
<emphasis id="B93A9254FFF95009FA3DFA6DFF6141AE" bold="true" pageId="9" pageNumber="10">6, 8</emphasis>
) views.
</paragraph>
</caption>
<paragraph id="8BF14E46FFF95009FF73F9E3FE2F428E" blockId="9.[151,1437,1583,1933]" pageId="9" pageNumber="10">
Aedeagus (
<figureCitation id="137552C3FFF95009FEF2F9E3FE684239" box="[326,429,1583,1610]" captionStart="FIGURES 18" captionStartId="9.[151,264,1405,1429]" captionTargetBox="[151,1429,181,1369]" captionTargetId="figure-16@9.[151,1435,181,1381]" captionTargetPageId="9" captionText="FIGURES 18. Anhoraeomorphus ankaratranus (Franz), holotype. Habitus in dorsal view (1); head in anterodorsal view (2); head and prothorax in lateral view (only bristles on head are shown) (3); antenna (4); aedeagus in ventral (5, 7) and lateral (6, 8) views." figureDoi="http://doi.org/10.5281/zenodo.13745916" httpUri="https://zenodo.org/record/13745916/files/figure.png" pageId="9" pageNumber="10">Figs 58</figureCitation>
,
<figureCitation id="137552C3FFF95009FE0FF9E3FDC4423A" box="[443,513,1583,1609]" captionStart="FIGURES 1219" captionStartId="10.[151,264,1945,1969]" captionTargetBox="[199,1390,902,1907]" captionTargetId="figure-107@10.[189,1397,898,1922]" captionTargetPageId="10" captionText="FIGURES 1219. Anhoraeomorphus ankaratranus (Franz), holotype of Horaeomorphus manjakotompoi Franz. Habitus in dorsal view (12); head in anterodorsal view (13); head and prothorax in lateral view (only bristles on head are shown) (14); antenna (15); aedeagus in ventral (16, 18) and lateral (17, 19) views." figureDoi="http://doi.org/10.5281/zenodo.13745923" httpUri="https://zenodo.org/record/13745923/files/figure.png" pageId="9" pageNumber="10">1619</figureCitation>
) stout, AeL
<quantity id="4CB6E3A3FFF95009FD25F9E3FD3F4239" box="[657,762,1583,1610]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="9" pageNumber="10" unit="mm" value="0.25">0.25 mm</quantity>
; in ventral view broadest near base and narrowing towards truncate apex; in lateral view apical region of dorsal wall strongly and abruptly bent dorsally; endophallus with symmetrical and elongate distolateral sclerites and basilateral sclerites and with asymmetrical weakly sclerotized median structures, distolateral sclerites broadly separated and each with rounded apex; parameres slender and their apices only slightly projecting beyond aedeagal apex, each paramere with a row of sparse and short setae occupying slightly more than distal third.
</paragraph>
<paragraph id="8BF14E46FFF95009FF73F8CBFD894319" blockId="9.[151,1437,1583,1933]" pageId="9" pageNumber="10">
Female. Externally similar to male, with indistinctly shorter antennae in relation to body length. BL
<quantity id="4CB6E3A3FFF95009FA98F8CBFF034335" metricMagnitude="-3" metricUnit="m" metricValue="1.64" metricValueMax="1.7" metricValueMin="1.58" pageId="9" pageNumber="10" unit="mm" value="1.64" valueMax="1.7" valueMin="1.58">1.581.70 mm</quantity>
; HL
<quantity id="4CB6E3A3FFF95009FF4AF8E0FE614335" box="[254,420,1836,1862]" metricMagnitude="-4" metricUnit="m" metricValue="3.15" metricValueMax="3.3" metricValueMin="3.0" pageId="9" pageNumber="10" unit="mm" value="0.315" valueMax="0.33" valueMin="0.3">0.300.33 mm</quantity>
, HW
<quantity id="4CB6E3A3FFF95009FE53F8E0FD884335" box="[487,589,1836,1862]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="10" unit="mm" value="0.3">0.30 mm</quantity>
, AnL
<quantity id="4CB6E3A3FFF95009FD26F8E7FCFC4335" box="[658,825,1835,1862]" metricMagnitude="-4" metricUnit="m" metricValue="5.9" metricValueMax="6.3" metricValueMin="5.5" pageId="9" pageNumber="10" unit="mm" value="0.59" valueMax="0.63" valueMin="0.55">0.550.63 mm</quantity>
; PL
<quantity id="4CB6E3A3FFF95009FCD9F8E7FBD64335" box="[877,1043,1835,1862]" metricMagnitude="-4" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="9" pageNumber="10" unit="mm" value="0.425" valueMax="0.45" valueMin="0.4">0.400.45 mm</quantity>
, PW
<quantity id="4CB6E3A3FFF95009FBE4F8E0FB324335" box="[1104,1271,1836,1862]" metricMagnitude="-4" metricUnit="m" metricValue="4.05" metricValueMax="4.3" metricValueMin="3.8" pageId="9" pageNumber="10" unit="mm" value="0.405" valueMax="0.43" valueMin="0.38">0.380.43 mm</quantity>
; EL
<quantity id="4CB6E3A3FFF95009FA98F8E7FF004319" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="9.5" metricValueMin="8.5" pageId="9" pageNumber="10" unit="mm" value="0.8999999999999999" valueMax="0.95" valueMin="0.85">0.850.95 mm</quantity>
, EW
<quantity id="4CB6E3A3FFF95009FEB7F883FE6D4319" box="[259,424,1871,1898]" metricMagnitude="-4" metricUnit="m" metricValue="7.05" metricValueMax="7.3" metricValueMin="6.8" pageId="9" pageNumber="10" unit="mm" value="0.705" valueMax="0.73" valueMin="0.68">0.680.73 mm</quantity>
, EI 1.251.31.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFF95009FF73F8BFFCAE43FE" box="[199,875,1907,1933]" pageId="9" pageNumber="10" type="distribution">
<paragraph id="8BF14E46FFF95009FF73F8BFFCAE43FE" blockId="9.[151,1437,1583,1933]" box="[199,875,1907,1933]" pageId="9" pageNumber="10">
<emphasis id="B93A9254FFF95009FF73F8BFFEA443FE" bold="true" box="[199,353,1907,1933]" pageId="9" pageNumber="10">Distribution.</emphasis>
Central
<collectingCountry id="F3590ED6FFF95009FE75F8BFFD8D43FE" box="[449,584,1907,1933]" name="Madagascar" pageId="9" pageNumber="10">Madagascar</collectingCountry>
(Vakinankaratra Region).
</paragraph>
</subSubSection>
<caption id="DF311ECEFFFA500AFF23FD0DFED04719" ID-DOI="http://doi.org/10.5281/zenodo.13745921" ID-Zenodo-Dep="13745921" httpUri="https://zenodo.org/record/13745921/files/figure.png" pageId="10" pageNumber="11" startId="10.[151,264,705,729]" targetBox="[183,1404,181,681]" targetPageId="10" targetType="figure">
<paragraph id="8BF14E46FFFA500AFF23FD0DFED04719" blockId="10.[151,1436,705,874]" pageId="10" pageNumber="11">
<emphasis id="B93A9254FFFA500AFF23FD0DFE8346AA" bold="true" box="[151,326,705,729]" pageId="10" pageNumber="11">FIGURES 911.</emphasis>
<taxonomicName id="4C4E35C5FFFA500AFEFFFD0EFD2C46A9" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[331,745,706,730]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ankaratranus">
<emphasis id="B93A9254FFFA500AFEFFFD0EFD5246A9" box="[331,663,706,730]" italics="true" pageId="10" pageNumber="11">Anhoraeomorphus ankaratranus</emphasis>
(Franz)
</taxonomicName>
, holotype. Head, prothorax and elytral base in dorsal view (
<emphasis id="B93A9254FFFA500AFAF0FD0DFA9446AA" bold="true" box="[1348,1361,705,729]" pageId="10" pageNumber="11">9</emphasis>
); head, prothorax and anterior region of mesothorax in ventral view (
<emphasis id="B93A9254FFFA500AFD48FD29FCD0468E" bold="true" box="[764,789,741,765]" pageId="10" pageNumber="11">10</emphasis>
); pterothorax in ventral view (
<emphasis id="B93A9254FFFA500AFBF3FD29FB9A468E" bold="true" box="[1095,1119,741,765]" pageId="10" pageNumber="11">11</emphasis>
). Abbreviations: abp, antebasal pit; bef, basal elytral fovea; gp, gular plate; gs, gular suture; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; pcr, procoxal rest; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum.
</paragraph>
</caption>
<caption id="DF311ECEFFFA500AFF23F855FC8B438A" ID-DOI="http://doi.org/10.5281/zenodo.13745923" ID-Zenodo-Dep="13745923" httpUri="https://zenodo.org/record/13745923/files/figure.png" pageId="10" pageNumber="11" startId="10.[151,264,1945,1969]" targetBox="[199,1390,902,1907]" targetPageId="10" targetType="figure">
<paragraph id="8BF14E46FFFA500AFF23F855FC8B438A" blockId="10.[151,1436,1945,2042]" pageId="10" pageNumber="11">
<emphasis id="B93A9254FFFA500AFF23F855FE9243C2" bold="true" box="[151,343,1945,1969]" pageId="10" pageNumber="11">FIGURES 1219.</emphasis>
<taxonomicName id="4C4E35C5FFFA500AFED4F855FCC143C1" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[352,772,1945,1970]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ankaratranus">
<emphasis id="B93A9254FFFA500AFED4F855FD6A43C2" box="[352,687,1945,1969]" italics="true" pageId="10" pageNumber="11">Anhoraeomorphus ankaratranus</emphasis>
(Franz)
</taxonomicName>
, holotype of
<taxonomicName id="4C4E35C5FFFA500AFC25F855FA4543C2" authority="Franz. Habitus" authorityName="Franz. Habitus" box="[913,1408,1945,1969]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="manjakotompoi">
<emphasis id="B93A9254FFFA500AFC25F855FB1A43C2" box="[913,1247,1945,1969]" italics="true" pageId="10" pageNumber="11">Horaeomorphus manjakotompoi</emphasis>
Franz. Habitus
</taxonomicName>
in dorsal view (
<emphasis id="B93A9254FFFA500AFEABF871FEFD43A6" bold="true" box="[287,312,1981,2005]" pageId="10" pageNumber="11">12</emphasis>
); head in anterodorsal view (
<emphasis id="B93A9254FFFA500AFDDFF871FD4143A6" bold="true" box="[619,644,1981,2005]" pageId="10" pageNumber="11">13</emphasis>
); head and prothorax in lateral view (only bristles on head are shown) (
<emphasis id="B93A9254FFFA500AFAC7F871FA4943A6" bold="true" box="[1395,1420,1981,2005]" pageId="10" pageNumber="11">14</emphasis>
); antenna (
<emphasis id="B93A9254FFFA500AFF41F82DFECB438A" bold="true" box="[245,270,2017,2041]" pageId="10" pageNumber="11">15</emphasis>
); aedeagus in ventral (
<emphasis id="B93A9254FFFA500AFE42F82DFDF0438A" bold="true" box="[502,565,2017,2041]" pageId="10" pageNumber="11">16, 18</emphasis>
) and lateral (
<emphasis id="B93A9254FFFA500AFD09F82DFD39438A" bold="true" box="[701,764,2017,2041]" pageId="10" pageNumber="11">17, 19</emphasis>
) views.
</paragraph>
</caption>
<subSubSection id="C3541DCDFFFB500BFF73FF5BFCA846DA" pageId="11" pageNumber="12" type="discussion">
<paragraph id="8BF14E46FFFB500BFF73FF5BFC2C4532" blockId="11.[151,1437,151,681]" pageId="11" pageNumber="12">
<emphasis id="B93A9254FFFB500BFF73FF5BFEFE44C2" bold="true" box="[199,315,151,177]" pageId="11" pageNumber="12">Remarks.</emphasis>
The
<typeStatus id="54F5F0E4FFFB500BFEC7FF5BFE6644C2" box="[371,419,151,177]" pageId="11" pageNumber="12">type</typeStatus>
specimens of
<taxonomicName id="4C4E35C5FFFB500BFDF6FF5BFD3E44C2" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[578,763,151,177]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ankaratranus">
<emphasis id="B93A9254FFFB500BFDF6FF5BFD3E44C2" box="[578,763,151,177]" italics="true" pageId="11" pageNumber="12">H. ankaratranus</emphasis>
</taxonomicName>
(
<figureCitation id="137552C3FFFB500BFCBDFF5BFCBC44C2" box="[777,889,151,177]" captionStart-0="FIGURES 18" captionStart-1="FIGURES 911" captionStartId-0="9.[151,264,1405,1429]" captionStartId-1="10.[151,264,705,729]" captionTargetBox-0="[151,1429,181,1369]" captionTargetBox-1="[183,1404,181,681]" captionTargetId-0="figure-16@9.[151,1435,181,1381]" captionTargetId-1="figure-28@10.[183,1404,181,681]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURES 18. Anhoraeomorphus ankaratranus (Franz), holotype. Habitus in dorsal view (1); head in anterodorsal view (2); head and prothorax in lateral view (only bristles on head are shown) (3); antenna (4); aedeagus in ventral (5, 7) and lateral (6, 8) views." captionText-1="FIGURES 911. Anhoraeomorphus ankaratranus (Franz), holotype. Head, prothorax and elytral base in dorsal view (9); head, prothorax and anterior region of mesothorax in ventral view (10); pterothorax in ventral view (11). Abbreviations: abp, antebasal pit; bef, basal elytral fovea; gp, gular plate; gs, gular suture; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; pcr, procoxal rest; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum." figureDoi-0="http://doi.org/10.5281/zenodo.13745916" figureDoi-1="http://doi.org/10.5281/zenodo.13745921" httpUri-0="https://zenodo.org/record/13745916/files/figure.png" httpUri-1="https://zenodo.org/record/13745921/files/figure.png" pageId="11" pageNumber="12">Figs 111</figureCitation>
) and
<taxonomicName id="4C4E35C5FFFB500BFC03FF5BFB4344C2" box="[951,1158,151,177]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="manjakotompoi">
<emphasis id="B93A9254FFFB500BFC03FF5BFB4344C2" box="[951,1158,151,177]" italics="true" pageId="11" pageNumber="12">H. manjakotompoi</emphasis>
</taxonomicName>
(
<figureCitation id="137552C3FFFB500BFB21FF5BFAD144C2" box="[1173,1300,151,177]" captionStart="FIGURES 1219" captionStartId="10.[151,264,1945,1969]" captionTargetBox="[199,1390,902,1907]" captionTargetId="figure-107@10.[189,1397,898,1922]" captionTargetPageId="10" captionText="FIGURES 1219. Anhoraeomorphus ankaratranus (Franz), holotype of Horaeomorphus manjakotompoi Franz. Habitus in dorsal view (12); head in anterodorsal view (13); head and prothorax in lateral view (only bristles on head are shown) (14); antenna (15); aedeagus in ventral (16, 18) and lateral (17, 19) views." figureDoi="http://doi.org/10.5281/zenodo.13745923" httpUri="https://zenodo.org/record/13745923/files/figure.png" pageId="11" pageNumber="12">Figs 1219</figureCitation>
) come from the same area of the Ankaratra volcano range (including specimens of
<taxonomicName id="4C4E35C5FFFB500BFC29FF77FB9044A6" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[925,1109,187,213]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ankaratranus">
<emphasis id="B93A9254FFFB500BFC29FF77FB9044A6" box="[925,1109,187,213]" italics="true" pageId="11" pageNumber="12">H. ankaratranus</emphasis>
</taxonomicName>
collected in Manjakotompo). They differ only in the pigmentation (
<taxonomicName id="4C4E35C5FFFB500BFDF8FF13FCCD448A" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[588,776,223,249]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ankaratranus">
<emphasis id="B93A9254FFFB500BFDF8FF13FCCD448A" box="[588,776,223,249]" italics="true" pageId="11" pageNumber="12">H. ankaratranus</emphasis>
</taxonomicName>
slightly darker) and the preservation of setae and bristles on the head (in
<taxonomicName id="4C4E35C5FFFB500BFEF9FECFFDDA456E" box="[333,543,259,285]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="manjakotompoi">
<emphasis id="B93A9254FFFB500BFEF9FECFFDDA456E" box="[333,543,259,285]" italics="true" pageId="11" pageNumber="12">H. manjakotompoi</emphasis>
</taxonomicName>
most of them are broken off). The aedeagi have the same shapes and internal structures, and there is no doubt that these specimens belong to one species.
</paragraph>
<paragraph id="8BF14E46FFFB500BFF73FE87FDC945DE" blockId="11.[151,1437,151,681]" pageId="11" pageNumber="12">
<taxonomicName id="4C4E35C5FFFB500BFF73FE87FDFC4516" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[199,569,331,357]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ankaratranus">
<emphasis id="B93A9254FFFB500BFF73FE87FDFC4516" box="[199,569,331,357]" italics="true" pageId="11" pageNumber="12">Anhoraeomorphus ankaratranus</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C4E35C5FFFB500BFCADFE87FC0F4516" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[793,970,331,357]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="sakarahanus">
<emphasis id="B93A9254FFFB500BFCADFE87FC0F4516" box="[793,970,331,357]" italics="true" pageId="11" pageNumber="12">A. sakarahanus</emphasis>
</taxonomicName>
, including external morphology and the aedeagus. These species clearly differ in the shape of the pronotum, in
<taxonomicName id="4C4E35C5FFFB500BFC63FEA3FB5545FA" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[983,1168,367,393]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ankaratranus">
<emphasis id="B93A9254FFFB500BFC63FEA3FB5545FA" box="[983,1168,367,393]" italics="true" pageId="11" pageNumber="12">A. ankaratranus</emphasis>
</taxonomicName>
broadest at base, in
<taxonomicName id="4C4E35C5FFFB500BFA30FEA3FEE345DE" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="sakarahanus">
<emphasis id="B93A9254FFFB500BFA30FEA3FEE345DE" italics="true" pageId="11" pageNumber="12">A. sakarahanus</emphasis>
</taxonomicName>
near posterior third.
</paragraph>
<paragraph id="8BF14E46FFFB500BFF73FE7BFCA846DA" blockId="11.[151,1437,151,681]" pageId="11" pageNumber="12">
<specimenCount id="9D4885CFFFFB500BFF73FE7BFEA345A2" box="[199,358,439,465]" count="1" pageId="11" pageNumber="12" type="generic">One specimen</specimenCount>
of
<taxonomicName id="4C4E35C5FFFB500BFE3DFE7BFDFB45A2" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[393,574,439,465]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ankaratranus">
<emphasis id="B93A9254FFFB500BFE3DFE7BFDFB45A2" box="[393,574,439,465]" italics="true" pageId="11" pageNumber="12">A. ankaratranus</emphasis>
</taxonomicName>
in the type series is labeled as “Typus”, while all others as “
<typeStatus id="54F5F0E4FFFB500BFB66FE7BFA8D45A2" box="[1234,1352,439,465]" pageId="11" pageNumber="12" type="paratype">Paratypus</typeStatus>
”.
<bibRefCitation id="EFDF33B7FFFB500BFAE8FE7BFF354586" author="Franz, H." pageId="11" pageNumber="12" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986b)</bibRefCitation>
uses the terms “
<typeStatus id="54F5F0E4FFFB500BFE17FE17FDD94586" box="[419,540,475,501]" pageId="11" pageNumber="12" type="holotype">Holotypus</typeStatus>
” and “
<typeStatus id="54F5F0E4FFFB500BFDD1FE17FD124586" box="[613,727,475,501]" pageId="11" pageNumber="12" type="paratype">Paratypes</typeStatus>
” in his description, so the “Typus” is a correctly fixed
<typeStatus id="54F5F0E4FFFB500BFA87FE17FA5C4586" box="[1331,1433,475,501]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
. All 27 studied specimens (from Franz Coll. in NHMW) were collected in Ankaratra volcanic range situated ~
<quantity id="4CB6E3A3FFFB500BFACBFE32FF7E464E" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="12" unit="km" value="50.0">50 km</quantity>
SW of
<collectingRegion id="498A80A4FFFB500BFEA4FDEFFE69464E" box="[272,428,547,573]" country="Madagascar" name="Antananarivo" pageId="11" pageNumber="12">Antananarivo</collectingRegion>
.
<bibRefCitation id="EFDF33B7FFFB500BFE02FDEFFD93464E" author="Franz, H." box="[438,598,547,573]" pageId="11" pageNumber="12" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986b)</bibRefCitation>
states that he examined
<specimenCount id="9D4885CFFFFB500BFCDDFDEFFBC4464E" box="[873,1025,547,573]" count="39" pageId="11" pageNumber="12" type="generic">39 specimens</specimenCount>
coming from Ankaratra, some taken in Manjakotompo, and some in Col de Manontongana. Only “Massif de lAnkaratra” is given on all labels, without exact localities. According to
<bibRefCitation id="EFDF33B7FFFB500BFE51FDA7FD4246F6" author="Franz, H." box="[485,647,619,645]" pageId="11" pageNumber="12" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986b)</bibRefCitation>
, only a part of the
<typeStatus id="54F5F0E4FFFB500BFCEEFDA7FC0046F6" box="[858,965,619,645]" pageId="11" pageNumber="12" type="paratype">paratypes</typeStatus>
is deposited in his collection (currently at NHMW); the depository of the remaining
<typeStatus id="54F5F0E4FFFB500BFDC6FD43FD1846DA" box="[626,733,655,681]" pageId="11" pageNumber="12" type="paratype">paratypes</typeStatus>
is unknown.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="5613CEE758CDD27C907E99BD89C8CF45">The curse of Horaeomorphus (almost) lifted. Revision of misplaced species from Madagascar, Comoros and Mascarenes (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:date id="AA43A132913EFD42CF03AAAE944B2FC3">2024</mods:date>
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<subSubSection id="C3541DCDFFFB500BFF23FD36FD2C4766" box="[151,745,762,789]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="8BF14E46FFFB500BFF23FD36FD2C4766" blockId="11.[151,745,762,825]" box="[151,745,762,789]" pageId="11" pageNumber="12">
<heading id="D0B9F92AFFFB500BFF23FD36FD2C4766" bold="true" box="[151,745,762,789]" fontSize="11" level="2" pageId="11" pageNumber="12" reason="4">
<emphasis id="B93A9254FFFB500BFF23FD36FD2C4766" bold="true" box="[151,745,762,789]" pageId="11" pageNumber="12">
<taxonomicName id="4C4E35C5FFFB500BFF23FD36FDB44767" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[151,625,762,789]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="andapensis" status="comb. nov.">
<emphasis id="B93A9254FFFB500BFF23FD36FDCF4766" bold="true" box="[151,522,762,789]" italics="true" pageId="11" pageNumber="12">Anhoraeomorphus andapensis</emphasis>
(Franz)
</taxonomicName>
,
<taxonomicNameLabel id="A2092F2FFFFB500BFDCBFD37FD2C4766" box="[639,745,763,789]" pageId="11" pageNumber="12" rank="species">comb. n.</taxonomicNameLabel>
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<subSubSection id="C3541DCDFFFB500BFF23FCD3FEE2474A" box="[151,295,799,825]" pageId="11" pageNumber="12" type="description">
<paragraph id="8BF14E46FFFB500BFF23FCD3FEE2474A" blockId="11.[151,745,762,825]" box="[151,295,799,825]" pageId="11" pageNumber="12">
(
<figureCitation id="137552C3FFFB500BFF2BFCD3FEE5474A" box="[159,288,799,825]" captionStart="FIGURES 2027" captionStartId="12.[151,264,1502,1526]" captionTargetBox="[151,1409,181,1461]" captionTargetId="figure-28@12.[151,1435,181,1478]" captionTargetPageId="12" captionText="FIGURES 2027. Anhoraeomorphus andapensis (Franz), holotype. Habitus in dorsal view (20); head in anterodorsal view (21); head and prothorax in lateral view (22); antenna (23); aedeagus in ventral (24, 26) and lateral (25, 27) views." figureDoi="http://doi.org/10.5281/zenodo.13745929" httpUri="https://zenodo.org/record/13745929/files/figure.png" pageId="11" pageNumber="12">Figs 2027</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFFB500BFF23FCAFFDB94708" box="[151,636,867,891]" pageId="11" pageNumber="12" type="reference_group">
<paragraph id="8BF14E46FFFB500BFF23FCAFFDB94708" blockId="11.[151,636,867,891]" box="[151,636,867,891]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="AB5E6968FFFB500BFF23FCAFFDB94708" box="[151,636,867,891]" pageId="11" pageNumber="12">
<treatmentCitation id="0AEF6857FFFB500BFF23FCAFFDBD4708" author="Franz, H." box="[151,632,867,891]" page="172" pageId="11" pageNumber="12" year="1986">
<taxonomicName id="4C4E35C5FFFB500BFF23FCAFFDBD4708" ID-CoL="3MNHY" authority="Franz, 1986 b: 172" authorityName="Franz" authorityPageNumber="172" authorityYear="1986" box="[151,632,867,891]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="andapensis">
<emphasis id="B93A9254FFFB500BFF23FCAFFE734708" box="[151,438,867,891]" italics="true" pageId="11" pageNumber="12">Horaeomorphus andapensis</emphasis>
<bibRefCitation id="EFDF33B7FFFB500BFE08FCA8FDBD4708" author="Franz, H." box="[444,632,867,891]" pageId="11" pageNumber="12" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 172</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFFB500BFF23FC65FE574097" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="8BF14E46FFFB500BFF23FC65FC5F4078" blockId="11.[151,1437,937,1936]" pageId="11" pageNumber="12">
<emphasis id="B93A9254FFFB500BFF23FC65FE6647B0" bold="true" box="[151,419,937,963]" pageId="11" pageNumber="12">Type material studied.</emphasis>
<materialsCitation id="3B26441BFFFB500BFE1DFC65FC5F4078" collectionCode="NHMW" collectorName="H. Franz" country="Madagascar" determinerName="H. Franz" location="Madagascar" pageId="11" pageNumber="12" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
<emphasis id="B93A9254FFFB500BFE1DFC65FD0947B0" bold="true" box="[425,716,937,963]" pageId="11" pageNumber="12">
<typeStatus id="54F5F0E4FFFB500BFE1DFC65FDD347B0" box="[425,534,937,963]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
(
<collectingCountry id="F3590ED6FFFB500BFD90FC65FD7A47B0" box="[548,703,937,963]" name="Madagascar" pageId="11" pageNumber="12">Madagascar</collectingCountry>
):
</emphasis>
<specimenCount id="9D4885CFFFFB500BFD67FC66FD2F47B7" box="[723,746,938,964]" count="1" pageId="11" pageNumber="12" type="male"></specimenCount>
, four labels: “Madagaskar,Montagne / de Marojezy b.Andape / lg.
<collectorName id="26BB2B90FFFB500BFF75FC01FEDA4794" box="[193,287,973,999]" pageId="11" pageNumber="12">H.Franz</collectorName>
1969” [white, printed], “
<taxonomicName id="4C4E35C5FFFB500BFD8AFC01FD304794" box="[574,757,973,999]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/ andapensis m. / det.
<determinerName id="7E2DA282FFFB500BFC5BFC01FB884794" box="[1007,1101,973,999]" pageId="11" pageNumber="12">H.Franz</determinerName>
” [yellowish, handwritten and printed], “Typus” [red, handwritten], “
<specimenCount id="9D4885CFFFFB500BFDF2FC3DFD994078" box="[582,604,1009,1035]" count="1" pageId="11" pageNumber="12" type="male"></specimenCount>
” [white, printed] (
<collectionCode id="ED5FD683FFFB500BFC98FC3DFC4A4078" box="[812,911,1009,1035]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="12" type="Museum">NHMW</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BF14E46FFFB500BFF73FBD9FE574097" blockId="11.[151,1437,937,1936]" pageId="11" pageNumber="12">
<emphasis id="B93A9254FFFB500BFF73FBD9FE66405C" bold="true" box="[199,419,1045,1071]" pageId="11" pageNumber="12">Revised diagnosis.</emphasis>
Body (
<figureCitation id="137552C3FFFB500BFE4EFBD9FD8B405C" box="[506,590,1045,1071]" captionStart="FIGURES 2027" captionStartId="12.[151,264,1502,1526]" captionTargetBox="[151,1409,181,1461]" captionTargetId="figure-28@12.[151,1435,181,1478]" captionTargetPageId="12" captionText="FIGURES 2027. Anhoraeomorphus andapensis (Franz), holotype. Habitus in dorsal view (20); head in anterodorsal view (21); head and prothorax in lateral view (22); antenna (23); aedeagus in ventral (24, 26) and lateral (25, 27) views." figureDoi="http://doi.org/10.5281/zenodo.13745929" httpUri="https://zenodo.org/record/13745929/files/figure.png" pageId="11" pageNumber="12">Fig. 20</figureCitation>
) slender; head (
<figureCitation id="137552C3FFFB500BFCB3FBD9FC9F405C" box="[775,858,1045,1071]" captionStart="FIGURES 2027" captionStartId="12.[151,264,1502,1526]" captionTargetBox="[151,1409,181,1461]" captionTargetId="figure-28@12.[151,1435,181,1478]" captionTargetPageId="12" captionText="FIGURES 2027. Anhoraeomorphus andapensis (Franz), holotype. Habitus in dorsal view (20); head in anterodorsal view (21); head and prothorax in lateral view (22); antenna (23); aedeagus in ventral (24, 26) and lateral (25, 27) views." figureDoi="http://doi.org/10.5281/zenodo.13745929" httpUri="https://zenodo.org/record/13745929/files/figure.png" pageId="11" pageNumber="12">Fig. 21</figureCitation>
) about as long as wide, with vertex subconical, in anterodorsal view its posterior margin subtriangular; pronotum (
<figureCitation id="137552C3FFFB500BFCD3FBF5FC7C4020" box="[871,953,1081,1107]" captionStart="FIGURES 2027" captionStartId="12.[151,264,1502,1526]" captionTargetBox="[151,1409,181,1461]" captionTargetId="figure-28@12.[151,1435,181,1478]" captionTargetPageId="12" captionText="FIGURES 2027. Anhoraeomorphus andapensis (Franz), holotype. Habitus in dorsal view (20); head in anterodorsal view (21); head and prothorax in lateral view (22); antenna (23); aedeagus in ventral (24, 26) and lateral (25, 27) views." figureDoi="http://doi.org/10.5281/zenodo.13745929" httpUri="https://zenodo.org/record/13745929/files/figure.png" pageId="11" pageNumber="12">Fig. 20</figureCitation>
) subtrapezoidal, broadest at base, with one pair of shallow antebasal pits; antennomeres 310 not elongate; all femora similarly slender; metatibiae in males unmodified; aedeagus (
<figureCitation id="137552C3FFFB500BFE15FB4DFDE640E8" box="[417,547,1153,1179]" captionStart="FIGURES 2027" captionStartId="12.[151,264,1502,1526]" captionTargetBox="[151,1409,181,1461]" captionTargetId="figure-28@12.[151,1435,181,1478]" captionTargetPageId="12" captionText="FIGURES 2027. Anhoraeomorphus andapensis (Franz), holotype. Habitus in dorsal view (20); head in anterodorsal view (21); head and prothorax in lateral view (22); antenna (23); aedeagus in ventral (24, 26) and lateral (25, 27) views." figureDoi="http://doi.org/10.5281/zenodo.13745929" httpUri="https://zenodo.org/record/13745929/files/figure.png" pageId="11" pageNumber="12">Figs 2427</figureCitation>
) in ventral view broadest near base, with apical region of dorsal wall weakly curved distoventrally, distolateral endophallic sclerites widely separated and with emarginate apices, each paramere with only apical setae.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFFB500CFF73FB21FE5B432C" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="description">
<paragraph id="8BF14E46FFFB500BFF73FB21FD5E415F" blockId="11.[151,1437,937,1936]" pageId="11" pageNumber="12">
<emphasis id="B93A9254FFFB500BFF73FB21FEB04174" bold="true" box="[199,373,1261,1287]" pageId="11" pageNumber="12">Redescription.</emphasis>
Body of male (
<figureCitation id="137552C3FFFB500BFD94FB21FDB54174" box="[544,624,1261,1287]" captionStart="FIGURES 2027" captionStartId="12.[151,264,1502,1526]" captionTargetBox="[151,1409,181,1461]" captionTargetId="figure-28@12.[151,1435,181,1478]" captionTargetPageId="12" captionText="FIGURES 2027. Anhoraeomorphus andapensis (Franz), holotype. Habitus in dorsal view (20); head in anterodorsal view (21); head and prothorax in lateral view (22); antenna (23); aedeagus in ventral (24, 26) and lateral (25, 27) views." figureDoi="http://doi.org/10.5281/zenodo.13745929" httpUri="https://zenodo.org/record/13745929/files/figure.png" pageId="11" pageNumber="12">Fig. 20</figureCitation>
) dark brown, nearly black, tarsi and palps distinctly lighter, setae slightly lighter than cuticle, dark brown; BL
<quantity id="4CB6E3A3FFFB500BFD84FADDFD52415F" box="[560,663,1297,1324]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="12" unit="mm" value="1.5">1.50 mm</quantity>
.
</paragraph>
<paragraph id="8BF14E46FFFB500BFF73FAF9FB294238" blockId="11.[151,1437,937,1936]" pageId="11" pageNumber="12">
Head (
<figureCitation id="137552C3FFFB500BFEA4FAF9FE55413C" box="[272,400,1333,1359]" captionStart="FIGURES 2027" captionStartId="12.[151,264,1502,1526]" captionTargetBox="[151,1409,181,1461]" captionTargetId="figure-28@12.[151,1435,181,1478]" captionTargetPageId="12" captionText="FIGURES 2027. Anhoraeomorphus andapensis (Franz), holotype. Habitus in dorsal view (20); head in anterodorsal view (21); head and prothorax in lateral view (22); antenna (23); aedeagus in ventral (24, 26) and lateral (25, 27) views." figureDoi="http://doi.org/10.5281/zenodo.13745929" httpUri="https://zenodo.org/record/13745929/files/figure.png" pageId="11" pageNumber="12">Figs 2122</figureCitation>
) in anterodorsal view almost triangular, broadest at eyes, HL
<quantity id="4CB6E3A3FFFB500BFB8FFAF9FB654123" box="[1083,1184,1333,1360]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="12" unit="mm" value="0.3">0.30 mm</quantity>
, HW
<quantity id="4CB6E3A3FFFB500BFB6BFAF9FA804123" box="[1247,1349,1333,1360]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="12" unit="mm" value="0.3">0.30 mm</quantity>
; temple in lateral view (
<figureCitation id="137552C3FFFB500BFEF3FA95FE5D4100" box="[327,408,1369,1395]" captionStart="FIGURES 2027" captionStartId="12.[151,264,1502,1526]" captionTargetBox="[151,1409,181,1461]" captionTargetId="figure-28@12.[151,1435,181,1478]" captionTargetPageId="12" captionText="FIGURES 2027. Anhoraeomorphus andapensis (Franz), holotype. Habitus in dorsal view (20); head in anterodorsal view (21); head and prothorax in lateral view (22); antenna (23); aedeagus in ventral (24, 26) and lateral (25, 27) views." figureDoi="http://doi.org/10.5281/zenodo.13745929" httpUri="https://zenodo.org/record/13745929/files/figure.png" pageId="11" pageNumber="12">Fig. 22</figureCitation>
) subequal to length of eye; vertex and frons confluent, together weakly convex and slightly elongate; vertex subconical, with its posterior margin nearly triangular; each eye moderately large and weakly convex, with barely marked posteromedian emargination, weakly projecting from head silhouette. Punctures on vertex fine, inconspicuous; setae short, sparse and suberect; posterior region of frons and entire vertex densely covered with thick bristles directed posteriorly. Antenna (
<figureCitation id="137552C3FFFB500BFC93FA25FCBF4270" box="[807,890,1513,1539]" captionStart="FIGURES 2027" captionStartId="12.[151,264,1502,1526]" captionTargetBox="[151,1409,181,1461]" captionTargetId="figure-28@12.[151,1435,181,1478]" captionTargetPageId="12" captionText="FIGURES 2027. Anhoraeomorphus andapensis (Franz), holotype. Habitus in dorsal view (20); head in anterodorsal view (21); head and prothorax in lateral view (22); antenna (23); aedeagus in ventral (24, 26) and lateral (25, 27) views." figureDoi="http://doi.org/10.5281/zenodo.13745929" httpUri="https://zenodo.org/record/13745929/files/figure.png" pageId="11" pageNumber="12">Fig. 23</figureCitation>
) slender but short, indistinctly shorter than half BL, AnL
<quantity id="4CB6E3A3FFFB500BFEB7F9C1FEAF425B" box="[259,362,1549,1576]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="11" pageNumber="12" unit="mm" value="0.68">0.68 mm</quantity>
, antennomeres 1 and 2 each weakly elongate, 310 each about as long as broad or indistinctly transverse; 11 shorter than 9 and 10 combined, almost as wide as 10, nearly twice as long as broad.
</paragraph>
<paragraph id="8BF14E46FFFB500BFF73F99AFD5D428C" blockId="11.[151,1437,937,1936]" pageId="11" pageNumber="12">
Pronotum (
<figureCitation id="137552C3FFFB500BFEF7F999FE56421C" box="[323,403,1621,1647]" captionStart="FIGURES 2027" captionStartId="12.[151,264,1502,1526]" captionTargetBox="[151,1409,181,1461]" captionTargetId="figure-28@12.[151,1435,181,1478]" captionTargetPageId="12" captionText="FIGURES 2027. Anhoraeomorphus andapensis (Franz), holotype. Habitus in dorsal view (20); head in anterodorsal view (21); head and prothorax in lateral view (22); antenna (23); aedeagus in ventral (24, 26) and lateral (25, 27) views." figureDoi="http://doi.org/10.5281/zenodo.13745929" httpUri="https://zenodo.org/record/13745929/files/figure.png" pageId="11" pageNumber="12">Fig. 20</figureCitation>
) subtrapezoidal, broadest at base; PL
<quantity id="4CB6E3A3FFFB500BFC87F999FC524203" box="[819,919,1621,1648]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="11" pageNumber="12" unit="mm" value="0.38">0.38 mm</quantity>
, PW
<quantity id="4CB6E3A3FFFB500BFC64F999FBF34203" box="[976,1078,1621,1648]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="11" pageNumber="12" unit="mm" value="0.35">0.35 mm</quantity>
.Anterior margin straight, lateral margins almost straight and moderately diverging posteriorly; posterior corners blunt; posterior margin indistinctly convex. Base with one pair of round and shallow pits, and with barely discernible, extremely short vestiges of sublateral carinae. Disc with fine, inconspicuous punctures; setae sparse, short and suberect, lateral and lateroventral surface of pronotum with dense thick bristles.
</paragraph>
<paragraph id="8BF14E46FFFB500BFF73F8C5FE7A4318" blockId="11.[151,1437,937,1936]" pageId="11" pageNumber="12">
Elytra (
<figureCitation id="137552C3FFFB500BFEAEF8C5FEAE4350" box="[282,363,1801,1827]" captionStart="FIGURES 2027" captionStartId="12.[151,264,1502,1526]" captionTargetBox="[151,1409,181,1461]" captionTargetId="figure-28@12.[151,1435,181,1478]" captionTargetPageId="12" captionText="FIGURES 2027. Anhoraeomorphus andapensis (Franz), holotype. Habitus in dorsal view (20); head in anterodorsal view (21); head and prothorax in lateral view (22); antenna (23); aedeagus in ventral (24, 26) and lateral (25, 27) views." figureDoi="http://doi.org/10.5281/zenodo.13745929" httpUri="https://zenodo.org/record/13745929/files/figure.png" pageId="11" pageNumber="12">Fig. 20</figureCitation>
) together oval, broadest slightly in front of middle; EL
<quantity id="4CB6E3A3FFFB500BFC64F8C5FBF04357" box="[976,1077,1801,1828]" metricMagnitude="-4" metricUnit="m" metricValue="8.299999999999999" pageId="11" pageNumber="12" unit="mm" value="0.83">0.83 mm</quantity>
, EW
<quantity id="4CB6E3A3FFFB500BFBC5F8C5FB134357" box="[1137,1238,1801,1828]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="11" pageNumber="12" unit="mm" value="0.68">0.68 mm</quantity>
, EI 1.22; humeral calli weakly elevated, basal elytral foveae vestigial, barely discernible. Punctures fine and inconspicuous; setae sparse, short and suberect.
</paragraph>
<paragraph id="8BF14E46FFFB500BFF73F8BAFC3643FC" blockId="11.[151,1437,937,1936]" box="[199,1011,1909,1936]" pageId="11" pageNumber="12">Legs moderately long, slender, unmodified except slightly curved tibiae.</paragraph>
<caption id="DF311ECEFFFC500CFF23FA12FADB4268" ID-DOI="http://doi.org/10.5281/zenodo.13745929" ID-Zenodo-Dep="13745929" httpUri="https://zenodo.org/record/13745929/files/figure.png" pageId="12" pageNumber="13" startId="12.[151,264,1502,1526]" targetBox="[151,1409,181,1461]" targetPageId="12" targetType="figure">
<paragraph id="8BF14E46FFFC500CFF23FA12FADB4268" blockId="12.[151,1436,1502,1563]" pageId="12" pageNumber="13">
<emphasis id="B93A9254FFFC500CFF23FA12FE924185" bold="true" box="[151,343,1502,1526]" pageId="12" pageNumber="13">FIGURES 2027.</emphasis>
<taxonomicName id="4C4E35C5FFFC500CFEEBFA13FD294184" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[351,748,1503,1527]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="andapensis">
<emphasis id="B93A9254FFFC500CFEEBFA13FD534184" box="[351,662,1503,1527]" italics="true" pageId="12" pageNumber="13">Anhoraeomorphus andapensis</emphasis>
(Franz)
</taxonomicName>
, holotype. Habitus in dorsal view (
<emphasis id="B93A9254FFFC500CFBEAFA12FBB24185" bold="true" box="[1118,1143,1502,1526]" pageId="12" pageNumber="13">20</emphasis>
); head in anterodorsal view (
<emphasis id="B93A9254FFFC500CFF14F9CEFF7C4269" bold="true" box="[160,185,1538,1562]" pageId="12" pageNumber="13">21</emphasis>
); head and prothorax in lateral view (
<emphasis id="B93A9254FFFC500CFD8CF9CEFD944269" bold="true" box="[568,593,1538,1562]" pageId="12" pageNumber="13">22</emphasis>
); antenna (
<emphasis id="B93A9254FFFC500CFD71F9CEFD1B4269" bold="true" box="[709,734,1538,1562]" pageId="12" pageNumber="13">23</emphasis>
); aedeagus in ventral (
<emphasis id="B93A9254FFFC500CFC72F9CEFBC04269" bold="true" box="[966,1029,1538,1562]" pageId="12" pageNumber="13">24, 26</emphasis>
) and lateral (
<emphasis id="B93A9254FFFC500CFB3AF9CEFB084269" bold="true" box="[1166,1229,1538,1562]" pageId="12" pageNumber="13">25, 27</emphasis>
) views.
</paragraph>
</caption>
<paragraph id="8BF14E46FFFC500CFF73F9A1FBAD4348" blockId="12.[151,1437,1645,2031]" pageId="12" pageNumber="13">
Aedeagus (
<figureCitation id="137552C3FFFC500CFEF0F9A1FE0142FB" box="[324,452,1645,1672]" captionStart="FIGURES 2027" captionStartId="12.[151,264,1502,1526]" captionTargetBox="[151,1409,181,1461]" captionTargetId="figure-28@12.[151,1435,181,1478]" captionTargetPageId="12" captionText="FIGURES 2027. Anhoraeomorphus andapensis (Franz), holotype. Habitus in dorsal view (20); head in anterodorsal view (21); head and prothorax in lateral view (22); antenna (23); aedeagus in ventral (24, 26) and lateral (25, 27) views." figureDoi="http://doi.org/10.5281/zenodo.13745929" httpUri="https://zenodo.org/record/13745929/files/figure.png" pageId="12" pageNumber="13">Figs 2425</figureCitation>
) stout, AeL
<quantity id="4CB6E3A3FFFC500CFDF8F9A1FD7742F4" box="[588,690,1645,1671]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="12" pageNumber="13" unit="mm" value="0.28">0.28 mm</quantity>
; in ventral view broadest near base and narrowing towards broadly rounded apex; in lateral view apical region of dorsal plate weakly curved distoventrally; endophallus with slightly asymmetrical and elongate distolateral sclerites, symmetrical basilateral sclerites and with asymmetrical weakly sclerotized median structures containing a group of elongate sclerites situated in basal region; distolateral sclerites broadly separated and each with distinctly emarginate apex; parameres slender with broadened apical regions, their apices not projecting beyond aedeagal apex, each paramere with two short apical setae.
</paragraph>
<paragraph id="8BF14E46FFFC500CFF73F889FE5B432C" blockId="12.[151,1437,1645,2031]" box="[199,414,1861,1887]" pageId="12" pageNumber="13">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFFC500CFF73F8A5FCCF43F0" box="[199,778,1897,1923]" pageId="12" pageNumber="13" type="distribution">
<paragraph id="8BF14E46FFFC500CFF73F8A5FCCF43F0" blockId="12.[151,1437,1645,2031]" box="[199,778,1897,1923]" pageId="12" pageNumber="13">
<emphasis id="B93A9254FFFC500CFF73F8A5FEA443F0" bold="true" box="[199,353,1897,1923]" pageId="12" pageNumber="13">Distribution.</emphasis>
Northern
<collectingCountry id="F3590ED6FFFC500CFE60F8A5FD9E43F0" box="[468,603,1897,1923]" name="Madagascar" pageId="12" pageNumber="13">Madagascar</collectingCountry>
(Sava Region).
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFFC500DFF73F841FDFC44A6" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="discussion">
<paragraph id="8BF14E46FFFC500CFF73F841FEE143B8" blockId="12.[151,1437,1645,2031]" pageId="12" pageNumber="13">
<emphasis id="B93A9254FFFC500CFF73F841FEFE43D4" bold="true" box="[199,315,1933,1959]" pageId="12" pageNumber="13">Remarks.</emphasis>
The names “Marojezy” and “Andape” from the locality label of the
<typeStatus id="54F5F0E4FFFC500CFB9DF841FB4E43D4" box="[1065,1163,1933,1959]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
are misspelled Marojejy and Andapa.
</paragraph>
<paragraph id="8BF14E46FFFC500DFF73F819FDFC44A6" blockId="12.[151,1437,1645,2031]" lastBlockId="13.[151,1436,151,213]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<taxonomicName id="4C4E35C5FFFC500CFF73F819FDD8439C" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[199,541,2005,2031]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="andapensis">
<emphasis id="B93A9254FFFC500CFF73F819FDD8439C" box="[199,541,2005,2031]" italics="true" pageId="12" pageNumber="13">Anhoraeomorphus andapensis</emphasis>
</taxonomicName>
is externally similar to
<taxonomicName id="4C4E35C5FFFC500CFC92F819FC18439C" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[806,989,2005,2031]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="ankaratranus">
<emphasis id="B93A9254FFFC500CFC92F819FC18439C" box="[806,989,2005,2031]" italics="true" pageId="12" pageNumber="13">A. ankaratranus</emphasis>
</taxonomicName>
, both species have the pronota broadest at base. However, in
<taxonomicName id="4C4E35C5FFFD500DFECBFF5BFDDE44C2" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[383,539,151,177]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="andapensis">
<emphasis id="B93A9254FFFD500DFECBFF5BFDDE44C2" box="[383,539,151,177]" italics="true" pageId="13" pageNumber="14">A. andapensis</emphasis>
</taxonomicName>
the pronotum has nearly straight lateral margins, whereas in
<taxonomicName id="4C4E35C5FFFD500DFB0BFF5BFAB144C2" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[1215,1396,151,177]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ankaratranus">
<emphasis id="B93A9254FFFD500DFB0BFF5BFAB144C2" box="[1215,1396,151,177]" italics="true" pageId="13" pageNumber="14">A. ankaratranus</emphasis>
</taxonomicName>
the pronotum has rounded, convex sides.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,230 @@
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<subSubSection id="C3541DCDFFFD500DFF23FED5FD234540" box="[151,742,281,308]" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="8BF14E46FFFD500DFF23FED5FD234540" blockId="13.[151,742,281,344]" box="[151,742,281,308]" pageId="13" pageNumber="14">
<heading id="D0B9F92AFFFD500DFF23FED5FD234540" bold="true" box="[151,742,281,308]" fontSize="11" level="2" pageId="13" pageNumber="14" reason="4">
<emphasis id="B93A9254FFFD500DFF23FED5FD234540" bold="true" box="[151,742,281,308]" pageId="13" pageNumber="14">
<taxonomicName id="4C4E35C5FFFD500DFF23FED5FDAB4540" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[151,622,281,308]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ivelonanus" status="comb. nov.">
<emphasis id="B93A9254FFFD500DFF23FED5FDC24547" bold="true" box="[151,519,281,308]" italics="true" pageId="13" pageNumber="14">Anhoraeomorphus ivelonanus</emphasis>
(Franz)
</taxonomicName>
,
<taxonomicNameLabel id="A2092F2FFFFD500DFDCFFED5FD234540" box="[635,742,281,307]" pageId="13" pageNumber="14" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFFD500DFF23FEF1FEE2452B" box="[151,295,317,344]" pageId="13" pageNumber="14" type="description">
<paragraph id="8BF14E46FFFD500DFF23FEF1FEE2452B" blockId="13.[151,742,281,344]" box="[151,295,317,344]" pageId="13" pageNumber="14">
(
<figureCitation id="137552C3FFFD500DFF2BFEF1FEDA452B" box="[159,287,317,344]" captionStart="FIGURES 2835" captionStartId="13.[151,264,1909,1933]" captionTargetBox="[151,1434,614,1874]" captionTargetId="figure-99@13.[151,1435,614,1886]" captionTargetPageId="13" captionText="FIGURES 2835. Anhoraeomorphus ivelonanus (Franz), holotype. Habitus in dorsal view (28); head in anterodorsal view (29); head and prothorax in lateral view (only bristles on head are shown) (30); antenna (31); aedeagus in ventral (32, 34) and lateral (33, 35) views." figureDoi="http://doi.org/10.5281/zenodo.13745931" httpUri="https://zenodo.org/record/13745931/files/figure.png" pageId="13" pageNumber="14">Figs 2835</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFFD500DFF23FE4EFDBD45E9" box="[151,632,386,410]" pageId="13" pageNumber="14" type="reference_group">
<paragraph id="8BF14E46FFFD500DFF23FE4EFDBD45E9" blockId="13.[151,632,386,410]" box="[151,632,386,410]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="AB5E6968FFFD500DFF23FE4EFDBD45E9" box="[151,632,386,410]" pageId="13" pageNumber="14">
<treatmentCitation id="0AEF6857FFFD500DFF23FE4EFDB145E9" author="Franz, H." box="[151,628,386,410]" page="171" pageId="13" pageNumber="14" year="1986">
<taxonomicName id="4C4E35C5FFFD500DFF23FE4EFDB145E9" ID-CoL="3MNKB" authority="Franz, 1986 b: 171" authorityName="Franz" authorityPageNumber="171" authorityYear="1986" box="[151,628,386,410]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ivelonanus">
<emphasis id="B93A9254FFFD500DFF23FE4EFE7745E9" box="[151,434,386,410]" italics="true" pageId="13" pageNumber="14">Horaeomorphus ivelonanus</emphasis>
<bibRefCitation id="EFDF33B7FFFD500DFE0CFE4EFDB145E9" author="Franz, H." box="[440,628,386,410]" pageId="13" pageNumber="14" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 171</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFFD500EFF23FE04FB524516" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="8BF14E46FFFD500DFF23FE04FEC8463D" blockId="13.[151,1436,456,590]" pageId="13" pageNumber="14">
<emphasis id="B93A9254FFFD500DFF23FE04FE774591" bold="true" box="[151,434,456,482]" pageId="13" pageNumber="14">Type material studied.</emphasis>
<materialsCitation id="3B26441BFFFD500DFE74FE04FEC8463D" collectionCode="NHMW" country="Madagascar" determinerName="H. Franz" location="MADAGASCAR" pageId="13" pageNumber="14" specimenCount="58" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B93A9254FFFD500DFE74FE04FD2E4591" bold="true" box="[448,747,456,482]" pageId="13" pageNumber="14">
<typeStatus id="54F5F0E4FFFD500DFE74FE04FDE84591" box="[448,557,456,482]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
(Madagascar):
</emphasis>
<specimenCount id="9D4885CFFFFD500DFD4EFE05FCD44590" box="[762,785,457,483]" count="1" pageId="13" pageNumber="14" type="male"></specimenCount>
, four labels: “Réserenat. III / Andranomalaza / Valeé dIvelona / X.
<specimenCount id="9D4885CFFFFD500DFE82FE20FEB44674" box="[310,369,492,519]" count="57" pageId="13" pageNumber="14" type="generic">57 P.</specimenCount>
Soga” [white, printed], “INSTITUT / SCIENTIFIQUE /
<collectingCountry id="F3590ED6FFFD500DFBB6FE20FB0C4675" box="[1026,1225,492,518]" name="Madagascar" pageId="13" pageNumber="14">MADAGASCAR</collectingCountry>
” [bluish, printed], “
<taxonomicName id="4C4E35C5FFFD500DFF15FDDCFE9D4659" box="[161,344,528,554]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/ ivelonanus m. / det.
<determinerName id="7E2DA282FFFD500DFDE4FDDCFD684659" box="[592,685,528,554]" pageId="13" pageNumber="14">H.Franz</determinerName>
” [yellowish, handwritten and printed], “Typus” [red, handwritten] (
<collectionCode id="ED5FD683FFFD500DFF2BFDF8FEC7463D" box="[159,258,564,590]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="13" pageNumber="14" type="Museum">NHMW</collectionCode>
).
</materialsCitation>
</paragraph>
<caption id="DF311ECEFFFD500DFF23F8B9FEF543A5" ID-DOI="http://doi.org/10.5281/zenodo.13745931" ID-Zenodo-Dep="13745931" httpUri="https://zenodo.org/record/13745931/files/figure.png" pageId="13" pageNumber="14" startId="13.[151,264,1909,1933]" targetBox="[151,1434,614,1874]" targetPageId="13" targetType="figure">
<paragraph id="8BF14E46FFFD500DFF23F8B9FEF543A5" blockId="13.[151,1436,1909,2006]" pageId="13" pageNumber="14">
<emphasis id="B93A9254FFFD500DFF23F8B9FE9643FD" bold="true" box="[151,339,1909,1934]" pageId="13" pageNumber="14">FIGURES 2835.</emphasis>
<taxonomicName id="4C4E35C5FFFD500DFEECF8BAFD1C43FD" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[344,729,1910,1934]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ivelonanus">
<emphasis id="B93A9254FFFD500DFEECF8BAFD4D43FD" box="[344,648,1910,1934]" italics="true" pageId="13" pageNumber="14">Anhoraeomorphus ivelonanus</emphasis>
(Franz)
</taxonomicName>
, holotype. Habitus in dorsal view (
<emphasis id="B93A9254FFFD500DFB82F8BAFB8A43FD" bold="true" box="[1078,1103,1910,1934]" pageId="13" pageNumber="14">28</emphasis>
); head in anterodorsal view (
<emphasis id="B93A9254FFFD500DFAC7F8BAFA4943FD" bold="true" box="[1395,1420,1910,1934]" pageId="13" pageNumber="14">29</emphasis>
); head and prothorax in lateral view (only bristles on head are shown) (
<emphasis id="B93A9254FFFD500DFCE3F856FCB543C1" bold="true" box="[855,880,1946,1970]" pageId="13" pageNumber="14">30</emphasis>
); antenna (
<emphasis id="B93A9254FFFD500DFC57F856FC3943C1" bold="true" box="[995,1020,1946,1970]" pageId="13" pageNumber="14">31</emphasis>
); aedeagus in ventral (
<emphasis id="B93A9254FFFD500DFB50F856FAE643C1" bold="true" box="[1252,1315,1946,1970]" pageId="13" pageNumber="14">32, 34</emphasis>
) and lateral (
<emphasis id="B93A9254FFFD500DFF14F872FF1A43A5" bold="true" box="[160,223,1982,2006]" pageId="13" pageNumber="14">33, 35</emphasis>
) views.
</paragraph>
</caption>
<paragraph id="8BF14E46FFFE500EFF73FF5AFB524516" blockId="14.[151,1437,150,1473]" pageId="14" pageNumber="15">
<emphasis id="B93A9254FFFE500EFF73FF5AFE6344C3" bold="true" box="[199,422,150,176]" pageId="14" pageNumber="15">Revised diagnosis.</emphasis>
Body (
<figureCitation id="137552C3FFFE500EFDB0FF5BFD9F44C2" box="[516,602,151,177]" captionStart="FIGURES 2835" captionStartId="13.[151,264,1909,1933]" captionTargetBox="[151,1434,614,1874]" captionTargetId="figure-99@13.[151,1435,614,1886]" captionTargetPageId="13" captionText="FIGURES 2835. Anhoraeomorphus ivelonanus (Franz), holotype. Habitus in dorsal view (28); head in anterodorsal view (29); head and prothorax in lateral view (only bristles on head are shown) (30); antenna (31); aedeagus in ventral (32, 34) and lateral (33, 35) views." figureDoi="http://doi.org/10.5281/zenodo.13745931" httpUri="https://zenodo.org/record/13745931/files/figure.png" pageId="14" pageNumber="15">Fig. 28</figureCitation>
) slender; head (
<figureCitation id="137552C3FFFE500EFCA9FF5BFCB644C2" box="[797,883,151,177]" captionStart="FIGURES 2835" captionStartId="13.[151,264,1909,1933]" captionTargetBox="[151,1434,614,1874]" captionTargetId="figure-99@13.[151,1435,614,1886]" captionTargetPageId="13" captionText="FIGURES 2835. Anhoraeomorphus ivelonanus (Franz), holotype. Habitus in dorsal view (28); head in anterodorsal view (29); head and prothorax in lateral view (only bristles on head are shown) (30); antenna (31); aedeagus in ventral (32, 34) and lateral (33, 35) views." figureDoi="http://doi.org/10.5281/zenodo.13745931" httpUri="https://zenodo.org/record/13745931/files/figure.png" pageId="14" pageNumber="15">Fig. 29</figureCitation>
) slightly elongate, with vertex semicircular, in anterodorsal view its posterior margin evenly rounded; pronotum (
<figureCitation id="137552C3FFFE500EFC3AFF77FC2444A6" box="[910,993,187,213]" captionStart="FIGURES 2835" captionStartId="13.[151,264,1909,1933]" captionTargetBox="[151,1434,614,1874]" captionTargetId="figure-99@13.[151,1435,614,1886]" captionTargetPageId="13" captionText="FIGURES 2835. Anhoraeomorphus ivelonanus (Franz), holotype. Habitus in dorsal view (28); head in anterodorsal view (29); head and prothorax in lateral view (only bristles on head are shown) (30); antenna (31); aedeagus in ventral (32, 34) and lateral (33, 35) views." figureDoi="http://doi.org/10.5281/zenodo.13745931" httpUri="https://zenodo.org/record/13745931/files/figure.png" pageId="14" pageNumber="15">Fig. 28</figureCitation>
) subtrapezoidal, broadest at base, with two pairs of deep antebasal pits; antennomeres 17 distinctly elongate, 810 each about as long as broad; all femora similarly slender; metatibiae in males unmodified; aedeagus (
<figureCitation id="137552C3FFFE500EFCEEFECFFC19456E" box="[858,988,258,285]" captionStart="FIGURES 2835" captionStartId="13.[151,264,1909,1933]" captionTargetBox="[151,1434,614,1874]" captionTargetId="figure-99@13.[151,1435,614,1886]" captionTargetPageId="13" captionText="FIGURES 2835. Anhoraeomorphus ivelonanus (Franz), holotype. Habitus in dorsal view (28); head in anterodorsal view (29); head and prothorax in lateral view (only bristles on head are shown) (30); antenna (31); aedeagus in ventral (32, 34) and lateral (33, 35) views." figureDoi="http://doi.org/10.5281/zenodo.13745931" httpUri="https://zenodo.org/record/13745931/files/figure.png" pageId="14" pageNumber="15">Figs 3235</figureCitation>
) in ventral view broadest near middle, with apical region of dorsal wall weakly curved distoventrally, distolateral endophallic sclerites curved mesad and overlapping at middle, each with rounded apex, each paramere with three long apical setae.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFFE500EFF73FEA2FE5B409A" pageId="14" pageNumber="15" type="description">
<paragraph id="8BF14E46FFFE500EFF73FEA2FD5845DE" blockId="14.[151,1437,150,1473]" pageId="14" pageNumber="15">
<emphasis id="B93A9254FFFE500EFF73FEA2FEB045FB" bold="true" box="[199,373,366,392]" pageId="14" pageNumber="15">Redescription.</emphasis>
Body of male (
<figureCitation id="137552C3FFFE500EFDABFEA3FDAA45FA" box="[543,623,367,393]" captionStart="FIGURES 2835" captionStartId="13.[151,264,1909,1933]" captionTargetBox="[151,1434,614,1874]" captionTargetId="figure-99@13.[151,1435,614,1886]" captionTargetPageId="13" captionText="FIGURES 2835. Anhoraeomorphus ivelonanus (Franz), holotype. Habitus in dorsal view (28); head in anterodorsal view (29); head and prothorax in lateral view (only bristles on head are shown) (30); antenna (31); aedeagus in ventral (32, 34) and lateral (33, 35) views." figureDoi="http://doi.org/10.5281/zenodo.13745931" httpUri="https://zenodo.org/record/13745931/files/figure.png" pageId="14" pageNumber="15">Fig. 28</figureCitation>
) light brown, tarsi and palps distinctly lighter, antennae slightly infuscate in distal halves, setae yellowish; BL
<quantity id="4CB6E3A3FFFE500EFD86FE5FFD5C45DE" box="[562,665,403,429]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="14" pageNumber="15" unit="mm" value="1.6">1.60 mm</quantity>
.
</paragraph>
<paragraph id="8BF14E46FFFE500EFF73FE7BFBB846DA" blockId="14.[151,1437,150,1473]" pageId="14" pageNumber="15">
Head (
<figureCitation id="137552C3FFFE500EFEA6FE7BFE5645A2" box="[274,403,439,465]" captionStart="FIGURES 2835" captionStartId="13.[151,264,1909,1933]" captionTargetBox="[151,1434,614,1874]" captionTargetId="figure-99@13.[151,1435,614,1886]" captionTargetPageId="13" captionText="FIGURES 2835. Anhoraeomorphus ivelonanus (Franz), holotype. Habitus in dorsal view (28); head in anterodorsal view (29); head and prothorax in lateral view (only bristles on head are shown) (30); antenna (31); aedeagus in ventral (32, 34) and lateral (33, 35) views." figureDoi="http://doi.org/10.5281/zenodo.13745931" httpUri="https://zenodo.org/record/13745931/files/figure.png" pageId="14" pageNumber="15">Figs 2930</figureCitation>
) in anterodorsal view subtrapezoidal, broadest at eyes, HL
<quantity id="4CB6E3A3FFFE500EFB87FE7BFB5F45A2" box="[1075,1178,439,465]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="15" unit="mm" value="0.3">0.30 mm</quantity>
, HW
<quantity id="4CB6E3A3FFFE500EFB68FE7BFA8145A2" box="[1244,1348,439,465]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="14" pageNumber="15" unit="mm" value="0.28">0.28 mm</quantity>
; temple in lateral view (
<figureCitation id="137552C3FFFE500EFEFDFE17FE5E4586" box="[329,411,475,501]" captionStart="FIGURES 2835" captionStartId="13.[151,264,1909,1933]" captionTargetBox="[151,1434,614,1874]" captionTargetId="figure-99@13.[151,1435,614,1886]" captionTargetPageId="13" captionText="FIGURES 2835. Anhoraeomorphus ivelonanus (Franz), holotype. Habitus in dorsal view (28); head in anterodorsal view (29); head and prothorax in lateral view (only bristles on head are shown) (30); antenna (31); aedeagus in ventral (32, 34) and lateral (33, 35) views." figureDoi="http://doi.org/10.5281/zenodo.13745931" httpUri="https://zenodo.org/record/13745931/files/figure.png" pageId="14" pageNumber="15">Fig. 30</figureCitation>
) slightly shorter than eye; vertex and frons confluent, together weakly convex and slightly elongate; posterior margin of vertex weakly and evenly rounded; each eye large and strongly convex, with small but distinct posteromedian emargination. Punctures on vertex fine, inconspicuous; setae short, sparse and suberect; frons and vertex densely covered with thick bristles directed posteriorly.Antenna (
<figureCitation id="137552C3FFFE500EFB91FD8BFBB04612" box="[1061,1141,583,609]" captionStart="FIGURES 2835" captionStartId="13.[151,264,1909,1933]" captionTargetBox="[151,1434,614,1874]" captionTargetId="figure-99@13.[151,1435,614,1886]" captionTargetPageId="13" captionText="FIGURES 2835. Anhoraeomorphus ivelonanus (Franz), holotype. Habitus in dorsal view (28); head in anterodorsal view (29); head and prothorax in lateral view (only bristles on head are shown) (30); antenna (31); aedeagus in ventral (32, 34) and lateral (33, 35) views." figureDoi="http://doi.org/10.5281/zenodo.13745931" httpUri="https://zenodo.org/record/13745931/files/figure.png" pageId="14" pageNumber="15">Fig. 31</figureCitation>
) slender but short, slightly shorter than half BL, AnL
<quantity id="4CB6E3A3FFFE500EFE77FDA7FDEF46F6" box="[451,554,619,645]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" pageId="14" pageNumber="15" unit="mm" value="0.78">0.78 mm</quantity>
, antennomeres 17 each elongate, 810 each about as long as broad; 11 much shorter than 9 and 10 combined, slightly narrower than 10, nearly twice as long as broad.
</paragraph>
<paragraph id="8BF14E46FFFE500EFF73FD7FFA57472E" blockId="14.[151,1437,150,1473]" pageId="14" pageNumber="15">
Pronotum (
<figureCitation id="137552C3FFFE500EFEF3FD7FFE5F46BE" box="[327,410,691,717]" captionStart="FIGURES 2835" captionStartId="13.[151,264,1909,1933]" captionTargetBox="[151,1434,614,1874]" captionTargetId="figure-99@13.[151,1435,614,1886]" captionTargetPageId="13" captionText="FIGURES 2835. Anhoraeomorphus ivelonanus (Franz), holotype. Habitus in dorsal view (28); head in anterodorsal view (29); head and prothorax in lateral view (only bristles on head are shown) (30); antenna (31); aedeagus in ventral (32, 34) and lateral (33, 35) views." figureDoi="http://doi.org/10.5281/zenodo.13745931" httpUri="https://zenodo.org/record/13745931/files/figure.png" pageId="14" pageNumber="15">Fig. 28</figureCitation>
) subtrapezoidal, equally broad from base to middle; PL
<quantity id="4CB6E3A3FFFE500EFB96FD7FFB4F46BE" box="[1058,1162,691,717]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="15" unit="mm" value="0.4">0.40 mm</quantity>
, PW
<quantity id="4CB6E3A3FFFE500EFB7EFD7FFAF746BE" box="[1226,1330,691,717]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="14" pageNumber="15" unit="mm" value="0.33">0.33 mm</quantity>
. Anterior margin weakly arcuate, lateral margins slightly rounded in anterior half and strongly diverging posteriorly, in posterior half parallel; posterior corners blunt; posterior margin indistinctly convex. Base with two pairs of round and deep pits, and with barely discernible, extremely short vestiges of sublateral carinae. Disc with fine, inconspicuous punctures; setae sparse, short and suberect, lateral and lateroventral surface of pronotum with dense thick bristles.
</paragraph>
<paragraph id="8BF14E46FFFE500EFF73FCABFF3947BA" blockId="14.[151,1437,150,1473]" pageId="14" pageNumber="15">
Elytra (
<figureCitation id="137552C3FFFE500EFEAFFCABFEA947F2" box="[283,364,871,897]" captionStart="FIGURES 2835" captionStartId="13.[151,264,1909,1933]" captionTargetBox="[151,1434,614,1874]" captionTargetId="figure-99@13.[151,1435,614,1886]" captionTargetPageId="13" captionText="FIGURES 2835. Anhoraeomorphus ivelonanus (Franz), holotype. Habitus in dorsal view (28); head in anterodorsal view (29); head and prothorax in lateral view (only bristles on head are shown) (30); antenna (31); aedeagus in ventral (32, 34) and lateral (33, 35) views." figureDoi="http://doi.org/10.5281/zenodo.13745931" httpUri="https://zenodo.org/record/13745931/files/figure.png" pageId="14" pageNumber="15">Fig. 28</figureCitation>
) together oval, broadest near middle; EL
<quantity id="4CB6E3A3FFFE500EFC89FCABFC6647F2" box="[829,931,871,897]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="14" pageNumber="15" unit="mm" value="0.9">0.90 mm</quantity>
, EW
<quantity id="4CB6E3A3FFFE500EFC54FCAAFB8347F2" box="[992,1094,870,897]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="14" pageNumber="15" unit="mm" value="0.65">0.65 mm</quantity>
, EI 1.38; humeral calli weakly elevated, basal elytral foveae vestigial, barely discernible. Punctures fine and inconspicuous; setae sparse, short and suberect.
</paragraph>
<paragraph id="8BF14E46FFFE500EFF73FC1FFBCC479E" blockId="14.[151,1437,150,1473]" box="[199,1033,979,1005]" pageId="14" pageNumber="15">Legs moderately long, slender, unmodified except slightly recurved tibiae.</paragraph>
<paragraph id="8BF14E46FFFE500EFF73FC3BFA8F40B6" blockId="14.[151,1437,150,1473]" pageId="14" pageNumber="15">
Aedeagus (
<figureCitation id="137552C3FFFE500EFEFDFC3BFE084062" box="[329,461,1014,1041]" captionStart="FIGURES 2835" captionStartId="13.[151,264,1909,1933]" captionTargetBox="[151,1434,614,1874]" captionTargetId="figure-99@13.[151,1435,614,1886]" captionTargetPageId="13" captionText="FIGURES 2835. Anhoraeomorphus ivelonanus (Franz), holotype. Habitus in dorsal view (28); head in anterodorsal view (29); head and prothorax in lateral view (only bristles on head are shown) (30); antenna (31); aedeagus in ventral (32, 34) and lateral (33, 35) views." figureDoi="http://doi.org/10.5281/zenodo.13745931" httpUri="https://zenodo.org/record/13745931/files/figure.png" pageId="14" pageNumber="15">Figs 3235</figureCitation>
) stout, AeL
<quantity id="4CB6E3A3FFFE500EFDD0FC3BFD0A4062" box="[612,719,1015,1041]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="14" pageNumber="15" unit="mm" value="0.23">0.23 mm</quantity>
; in ventral view broadest near middle and weakly narrowing towards step-wise delimited apical region, which is subrectangular, transverse and shallowly emarginate; in lateral view apical region of dorsal plate weakly curved distoventrally; endophallus with slightly asymmetrical and elongate distolateral sclerites, symmetrical lateral groups of long and dense basal sclerites and asymmetrical weakly sclerotized median structures, distolateral sclerites strongly curved mesad and overlapping at middle, each with rounded apex; parameres thick, much shorter than median lobe, each paramere with three long apical setae.
</paragraph>
<paragraph id="8BF14E46FFFE500EFF73FB03FE5B409A" blockId="14.[151,1437,150,1473]" box="[199,414,1231,1257]" pageId="14" pageNumber="15">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFFE500EFF73FB3EFC8A417E" box="[199,847,1266,1293]" pageId="14" pageNumber="15" type="distribution">
<paragraph id="8BF14E46FFFE500EFF73FB3EFC8A417E" blockId="14.[151,1437,150,1473]" box="[199,847,1266,1293]" pageId="14" pageNumber="15">
<emphasis id="B93A9254FFFE500EFF73FB3EFEA4417F" bold="true" box="[199,353,1266,1292]" pageId="14" pageNumber="15">Distribution.</emphasis>
Central
<collectingCountry id="F3590ED6FFFE500EFE75FB3FFD8D417E" box="[449,584,1267,1293]" name="Madagascar" pageId="14" pageNumber="15">Madagascar</collectingCountry>
(Analamanga Region).
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFFE500EFF73FADBFC3841B2" pageId="14" pageNumber="15" type="discussion">
<paragraph id="8BF14E46FFFE500EFF73FADBFA444142" blockId="14.[151,1437,150,1473]" box="[199,1409,1303,1329]" pageId="14" pageNumber="15">
<emphasis id="B93A9254FFFE500EFF73FADBFEFE4142" bold="true" box="[199,315,1303,1329]" pageId="14" pageNumber="15">Remarks.</emphasis>
According to
<bibRefCitation id="EFDF33B7FFFE500EFE6EFADBFDA04142" author="Franz, H." box="[474,613,1303,1329]" pageId="14" pageNumber="15" pagination="1 - 93" refId="ref52970" refString="Franz, H. (1963) Scydmaeniden (Coleoptera) aus Ostafrika, dem Kongo und der Tschadrepublik (vorwiegend aus den im Sommer 1962 vom Verfasser selbst gesammelten Ausbeuten). Stuttgarter Beitrage zur Naturkunde, 120, 1 - 93." type="journal article" year="1963">Franz (1963</bibRefCitation>
b), the word “Réserenat.” on the label is a misspelling of Réserve nat.
</paragraph>
<paragraph id="8BF14E46FFFE500EFF73FAF7FC3841B2" blockId="14.[151,1437,150,1473]" pageId="14" pageNumber="15">
<taxonomicName id="4C4E35C5FFFE500EFF73FAF7FDDF4126" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[199,538,1339,1365]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="ivelonanus">
<emphasis id="B93A9254FFFE500EFF73FAF7FDDF4126" box="[199,538,1339,1365]" italics="true" pageId="14" pageNumber="15">Anhoraeomorphus ivelonanus</emphasis>
</taxonomicName>
is externally slightly similar to
<taxonomicName id="4C4E35C5FFFE500EFC39FAF7FBCE4126" box="[909,1035,1339,1365]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="grucheti">
<emphasis id="B93A9254FFFE500EFC39FAF7FBCE4126" box="[909,1035,1339,1365]" italics="true" pageId="14" pageNumber="15">A. grucheti</emphasis>
</taxonomicName>
, but it can be distinguished by the stronger elongate pronotum and distinctly elongate antennomeres 46 (about as long as broad in
<taxonomicName id="4C4E35C5FFFE500EFABFFA93FA49410A" box="[1291,1420,1375,1401]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="grucheti">
<emphasis id="B93A9254FFFE500EFABFFA93FA49410A" box="[1291,1420,1375,1401]" italics="true" pageId="14" pageNumber="15">A. grucheti</emphasis>
</taxonomicName>
). Another externally similar species is
<taxonomicName id="4C4E35C5FFFE500EFD86FA4FFD7741EF" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[562,690,1411,1437]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="procerus">
<emphasis id="B93A9254FFFE500EFD86FA4FFD7741EF" box="[562,690,1411,1437]" italics="true" pageId="14" pageNumber="15">A. procerus</emphasis>
</taxonomicName>
. However, in
<taxonomicName id="4C4E35C5FFFE500EFCF9FA4FFC0841EF" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[845,973,1411,1437]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="procerus">
<emphasis id="B93A9254FFFE500EFCF9FA4FFC0841EF" box="[845,973,1411,1437]" italics="true" pageId="14" pageNumber="15">A. procerus</emphasis>
</taxonomicName>
, the posterior margin of vertex is truncate and straight, whereas that in
<taxonomicName id="4C4E35C5FFFE500EFE6DFA6BFDB441B2" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[473,625,1447,1473]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="ivelonanus">
<emphasis id="B93A9254FFFE500EFE6DFA6BFDB441B2" box="[473,625,1447,1473]" italics="true" pageId="14" pageNumber="15">A. ivelonanus</emphasis>
</taxonomicName>
is rounded and posteriorly convex.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="F86C905E8549551F0324C9E4A140E940" ID-DOI="10.11646/zootaxa.5505.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="13745912" ID-ZooBank="93F5E9F6-9EDC-4ED2-8F2B-1E6D0C18F893" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726052234810" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2024" docId="03E7FF50FFFE5010FF23F9DEFAF54062" docLanguage="en" docName="zootaxa.5505.1.1.pdf" docOrigin="Zootaxa 5505 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5505.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Anhoraeomorphus procerus Jałoszyński 2024, comb. n." docType="treatment" docVersion="2" lastPageNumber="17" masterDocId="FFDE8728FFF05000FFB4FFCCFFC54473" masterDocTitle="The curse of Horaeomorphus (almost) lifted. Revision of misplaced species from Madagascar, Comoros and Mascarenes (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="96" masterPageNumber="1" pageNumber="15" updateTime="1726053143081" updateUser="ExternalLinkService">
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<mods:title id="45EE4D21E878C36B1ED6F6A0EEDE8A4D">The curse of Horaeomorphus (almost) lifted. Revision of misplaced species from Madagascar, Comoros and Mascarenes (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:date id="6DB8A176E30EC151A9FB71C901CC5DAA">2024</mods:date>
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<paragraph id="8BF14E46FFFE500EFF23F9DEFD09425E" blockId="14.[151,716,1554,1617]" box="[151,716,1554,1581]" pageId="14" pageNumber="15">
<heading id="D0B9F92AFFFE500EFF23F9DEFD09425E" bold="true" box="[151,716,1554,1581]" fontSize="11" level="2" pageId="14" pageNumber="15" reason="4">
<emphasis id="B93A9254FFFE500EFF23F9DEFD09425E" bold="true" box="[151,716,1554,1581]" pageId="14" pageNumber="15">
<taxonomicName id="4C4E35C5FFFE500EFF23F9DEFD91425F" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[151,596,1554,1581]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="procerus" status="comb. nov.">
<emphasis id="B93A9254FFFE500EFF23F9DEFE28425E" bold="true" box="[151,493,1554,1581]" italics="true" pageId="14" pageNumber="15">Anhoraeomorphus procerus</emphasis>
(Franz)
</taxonomicName>
,
<taxonomicNameLabel id="A2092F2FFFFE500EFDD5F9DFFD09425E" box="[609,716,1555,1581]" pageId="14" pageNumber="15" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFFE500EFF23F9FBFEE24222" box="[151,295,1591,1617]" pageId="14" pageNumber="15" type="description">
<paragraph id="8BF14E46FFFE500EFF23F9FBFEE24222" blockId="14.[151,716,1554,1617]" box="[151,295,1591,1617]" pageId="14" pageNumber="15">
(
<figureCitation id="137552C3FFFE500EFF2BF9FBFEDA4222" box="[159,287,1591,1617]" captionStart="FIGURES 3643" captionStartId="15.[151,264,1679,1703]" captionTargetBox="[151,1418,273,1635]" captionTargetId="figure-64@15.[151,1435,273,1655]" captionTargetPageId="15" captionText="FIGURES 3643. Anhoraeomorphus procerus (Franz), holotype. Habitus in dorsal view (36); head in anterodorsal view (37); head and prothorax in lateral view (only bristles on head are shown) (38); antenna (39); aedeagus in ventral (40, 42) and lateral (41, 43) views." figureDoi="http://doi.org/10.5281/zenodo.13745935" httpUri="https://zenodo.org/record/13745935/files/figure.png" pageId="14" pageNumber="15">Figs 3643</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFFE500EFF23F9B7FDA642E0" box="[151,611,1659,1683]" pageId="14" pageNumber="15" type="reference_group">
<paragraph id="8BF14E46FFFE500EFF23F9B7FDA642E0" blockId="14.[151,611,1659,1683]" box="[151,611,1659,1683]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="AB5E6968FFFE500EFF23F9B7FDA642E0" box="[151,611,1659,1683]" pageId="14" pageNumber="15">
<treatmentCitation id="0AEF6857FFFE500EFF23F9B7FD9A42E0" author="Franz, H." box="[151,607,1659,1683]" page="166" pageId="14" pageNumber="15" year="1986">
<taxonomicName id="4C4E35C5FFFE500EFF23F9B7FD9A42E0" ID-CoL="3MNL6" authority="Franz, 1986 b: 166" authorityName="Franz" authorityPageNumber="166" authorityYear="1986" box="[151,607,1659,1683]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="procerus">
<emphasis id="B93A9254FFFE500EFF23F9B7FE5842E0" box="[151,413,1659,1683]" italics="true" pageId="14" pageNumber="15">Horaeomorphus procerus</emphasis>
<bibRefCitation id="EFDF33B7FFFE500EFE17F9B0FD9A42E0" author="Franz, H." box="[419,607,1659,1683]" pageId="14" pageNumber="15" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 166</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFFE500FFF23F90DFADB448A" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="8BF14E46FFFE500EFF23F90DFEC84334" blockId="14.[151,1437,1729,2008]" pageId="14" pageNumber="15">
<emphasis id="B93A9254FFFE500EFF23F90DFE7042A8" bold="true" box="[151,437,1729,1755]" pageId="14" pageNumber="15">Type material studied.</emphasis>
<materialsCitation id="3B26441BFFFE500EFE71F90DFEC84334" collectionCode="NHMW" country="Madagascar" determinerName="H. Franz" location="MADAGASCAR" pageId="14" pageNumber="15" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B93A9254FFFE500EFE71F90DFD3742A8" bold="true" box="[453,754,1729,1755]" pageId="14" pageNumber="15">
<typeStatus id="54F5F0E4FFFE500EFE71F90DFDF742A8" box="[453,562,1729,1755]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
(Madagascar):
</emphasis>
<specimenCount id="9D4885CFFFFE500EFCB6F90EFCDC42AF" box="[770,793,1730,1756]" count="1" pageId="14" pageNumber="15" type="male"></specimenCount>
, three labels: “Madagascar Est / district Sambava / Marojejy / Ambinanitelo” [white, printed], “INSTITUT / SCIENTIFIQUE /
<collectingCountry id="F3590ED6FFFE500EFBB6F929FB0C428C" box="[1026,1225,1765,1791]" name="Madagascar" pageId="14" pageNumber="15">MADAGASCAR</collectingCountry>
” [bluish, printed], “
<taxonomicName id="4C4E35C5FFFE500EFF15F8C5FE9D4350" box="[161,344,1801,1827]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/ procerus m. / det.
<determinerName id="7E2DA282FFFE500EFDF5F8C5FD5A4350" box="[577,671,1801,1827]" pageId="14" pageNumber="15">H.Franz</determinerName>
” [yellowish, handwritten and printed], “Typus” [red, handwritten] (
<collectionCode id="ED5FD683FFFE500EFF2BF8E1FEC74334" box="[159,258,1837,1863]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="14" pageNumber="15" type="Museum">NHMW</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BF14E46FFFE500FFF73F89DFADB448A" blockId="14.[151,1437,1729,2008]" lastBlockId="15.[151,1436,151,249]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<emphasis id="B93A9254FFFE500EFF73F89DFE664318" bold="true" box="[199,419,1873,1899]" pageId="14" pageNumber="15">Revised diagnosis.</emphasis>
Body (
<figureCitation id="137552C3FFFE500EFE4EF89DFD884318" box="[506,589,1873,1899]" captionStart="FIGURES 3643" captionStartId="15.[151,264,1679,1703]" captionTargetBox="[151,1418,273,1635]" captionTargetId="figure-64@15.[151,1435,273,1655]" captionTargetPageId="15" captionText="FIGURES 3643. Anhoraeomorphus procerus (Franz), holotype. Habitus in dorsal view (36); head in anterodorsal view (37); head and prothorax in lateral view (only bristles on head are shown) (38); antenna (39); aedeagus in ventral (40, 42) and lateral (41, 43) views." figureDoi="http://doi.org/10.5281/zenodo.13745935" httpUri="https://zenodo.org/record/13745935/files/figure.png" pageId="14" pageNumber="15">Fig. 36</figureCitation>
) slender; head (
<figureCitation id="137552C3FFFE500EFCB1F89DFC9C4318" box="[773,857,1873,1899]" captionStart="FIGURES 3643" captionStartId="15.[151,264,1679,1703]" captionTargetBox="[151,1418,273,1635]" captionTargetId="figure-64@15.[151,1435,273,1655]" captionTargetPageId="15" captionText="FIGURES 3643. Anhoraeomorphus procerus (Franz), holotype. Habitus in dorsal view (36); head in anterodorsal view (37); head and prothorax in lateral view (only bristles on head are shown) (38); antenna (39); aedeagus in ventral (40, 42) and lateral (41, 43) views." figureDoi="http://doi.org/10.5281/zenodo.13745935" httpUri="https://zenodo.org/record/13745935/files/figure.png" pageId="14" pageNumber="15">Fig. 37</figureCitation>
) about as long as broad, with vertex transverse, in anterodorsal view its posterior margin truncate at middle; pronotum (
<figureCitation id="137552C3FFFE500EFC04F8B9FBC143FC" box="[944,1028,1909,1935]" captionStart="FIGURES 3643" captionStartId="15.[151,264,1679,1703]" captionTargetBox="[151,1418,273,1635]" captionTargetId="figure-64@15.[151,1435,273,1655]" captionTargetPageId="15" captionText="FIGURES 3643. Anhoraeomorphus procerus (Franz), holotype. Habitus in dorsal view (36); head in anterodorsal view (37); head and prothorax in lateral view (only bristles on head are shown) (38); antenna (39); aedeagus in ventral (40, 42) and lateral (41, 43) views." figureDoi="http://doi.org/10.5281/zenodo.13745935" httpUri="https://zenodo.org/record/13745935/files/figure.png" pageId="14" pageNumber="15">Fig. 36</figureCitation>
) bell-shaped, broadest near middle, with two pairs of deep antebasal pits: large and deep inner pits and barely discernible, small and shallow outer pits; antennomeres 12 elongate, 3 about as long as broad, 47 each elongate, 810 each about as long as broad; all femora similarly slender; metatibiae in males unmodified; aedeagus (
<figureCitation id="137552C3FFFF500FFC71FF5BFB8344C2" box="[965,1094,151,177]" captionStart="FIGURES 3643" captionStartId="15.[151,264,1679,1703]" captionTargetBox="[151,1418,273,1635]" captionTargetId="figure-64@15.[151,1435,273,1655]" captionTargetPageId="15" captionText="FIGURES 3643. Anhoraeomorphus procerus (Franz), holotype. Habitus in dorsal view (36); head in anterodorsal view (37); head and prothorax in lateral view (only bristles on head are shown) (38); antenna (39); aedeagus in ventral (40, 42) and lateral (41, 43) views." figureDoi="http://doi.org/10.5281/zenodo.13745935" httpUri="https://zenodo.org/record/13745935/files/figure.png" pageId="15" pageNumber="16">Figs 4043</figureCitation>
) in ventral view broadest near base, with apical region of dorsal wall nearly straight, distolateral endophallic sclerites broadly separated, each with hook-like apex curved mesad, each paramere with one long and four subapical setae of various lengths.
</paragraph>
</subSubSection>
<caption id="DF311ECEFFFF500FFF23F943FEF5429C" ID-DOI="http://doi.org/10.5281/zenodo.13745935" ID-Zenodo-Dep="13745935" httpUri="https://zenodo.org/record/13745935/files/figure.png" pageId="15" pageNumber="16" startId="15.[151,264,1679,1703]" targetBox="[151,1418,273,1635]" targetPageId="15" targetType="figure">
<paragraph id="8BF14E46FFFF500FFF23F943FEF5429C" blockId="15.[151,1436,1679,1775]" pageId="15" pageNumber="16">
<emphasis id="B93A9254FFFF500FFF23F943FE9042D4" bold="true" box="[151,341,1679,1703]" pageId="15" pageNumber="16">FIGURES 3643.</emphasis>
<taxonomicName id="4C4E35C5FFFF500FFEEFF943FD0F42D4" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[347,714,1679,1703]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="procerus">
<emphasis id="B93A9254FFFF500FFEEFF943FDB242D4" box="[347,631,1679,1703]" italics="true" pageId="15" pageNumber="16">Anhoraeomorphus procerus</emphasis>
(Franz)
</taxonomicName>
, holotype. Habitus in dorsal view (
<emphasis id="B93A9254FFFF500FFB9BF943FB8D42D4" bold="true" box="[1071,1096,1679,1703]" pageId="15" pageNumber="16">36</emphasis>
); head in anterodorsal view (
<emphasis id="B93A9254FFFF500FFAC7F943FA4942D4" bold="true" box="[1395,1420,1679,1703]" pageId="15" pageNumber="16">37</emphasis>
); head and prothorax in lateral view (only bristles on head are shown) (
<emphasis id="B93A9254FFFF500FFCE3F97FFCB542B8" bold="true" box="[855,880,1715,1739]" pageId="15" pageNumber="16">38</emphasis>
); antenna (
<emphasis id="B93A9254FFFF500FFC57F97FFC3942B8" bold="true" box="[995,1020,1715,1739]" pageId="15" pageNumber="16">39</emphasis>
); aedeagus in ventral (
<emphasis id="B93A9254FFFF500FFB50F97FFAE642B8" bold="true" box="[1252,1315,1715,1739]" pageId="15" pageNumber="16">40, 42</emphasis>
) and lateral (
<emphasis id="B93A9254FFFF500FFF14F91BFF1A429C" bold="true" box="[160,223,1751,1775]" pageId="15" pageNumber="16">41, 43</emphasis>
) views.
</paragraph>
</caption>
<subSubSection id="C3541DCDFFFF5010FF73F88DFE5B472E" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="description">
<paragraph id="8BF14E46FFFF500FFF73F88DFF0F43F3" blockId="15.[151,1436,1857,2028]" pageId="15" pageNumber="16">
<emphasis id="B93A9254FFFF500FFF73F88DFEB04328" bold="true" box="[199,373,1857,1883]" pageId="15" pageNumber="16">Redescription.</emphasis>
Body of male (
<figureCitation id="137552C3FFFF500FFD91F88DFDB34328" box="[549,630,1857,1883]" captionStart="FIGURES 3643" captionStartId="15.[151,264,1679,1703]" captionTargetBox="[151,1418,273,1635]" captionTargetId="figure-64@15.[151,1435,273,1655]" captionTargetPageId="15" captionText="FIGURES 3643. Anhoraeomorphus procerus (Franz), holotype. Habitus in dorsal view (36); head in anterodorsal view (37); head and prothorax in lateral view (only bristles on head are shown) (38); antenna (39); aedeagus in ventral (40, 42) and lateral (41, 43) views." figureDoi="http://doi.org/10.5281/zenodo.13745935" httpUri="https://zenodo.org/record/13745935/files/figure.png" pageId="15" pageNumber="16">Fig. 36</figureCitation>
) light brown, tarsi and palps distinctly lighter, setae yellowish; BL
<quantity id="4CB6E3A3FFFF500FFADFF88DFF0343F3" metricMagnitude="-3" metricUnit="m" metricValue="1.53" pageId="15" pageNumber="16" unit="mm" value="1.53">1.53 mm</quantity>
.
</paragraph>
<paragraph id="8BF14E46FFFF5010FF73F845FB1B456E" blockId="15.[151,1436,1857,2028]" lastBlockId="16.[151,1437,151,1041]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
Head (
<figureCitation id="137552C3FFFF500FFEA0F845FE5243D0" box="[276,407,1929,1955]" captionStart="FIGURES 3643" captionStartId="15.[151,264,1679,1703]" captionTargetBox="[151,1418,273,1635]" captionTargetId="figure-64@15.[151,1435,273,1655]" captionTargetPageId="15" captionText="FIGURES 3643. Anhoraeomorphus procerus (Franz), holotype. Habitus in dorsal view (36); head in anterodorsal view (37); head and prothorax in lateral view (only bristles on head are shown) (38); antenna (39); aedeagus in ventral (40, 42) and lateral (41, 43) views." figureDoi="http://doi.org/10.5281/zenodo.13745935" httpUri="https://zenodo.org/record/13745935/files/figure.png" pageId="15" pageNumber="16">Figs 3738</figureCitation>
) in anterodorsal view subquadrate, broadest at eyes, HL
<quantity id="4CB6E3A3FFFF500FFB9EF845FB5743D7" box="[1066,1170,1929,1956]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="15" pageNumber="16" unit="mm" value="0.25">0.25 mm</quantity>
, HW
<quantity id="4CB6E3A3FFFF500FFB6CF845FA8743D7" box="[1240,1346,1929,1956]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="15" pageNumber="16" unit="mm" value="0.28">0.28 mm</quantity>
; temple in lateral view (
<figureCitation id="137552C3FFFF500FFEFAF861FE6743B4" box="[334,418,1965,1991]" captionStart="FIGURES 3643" captionStartId="15.[151,264,1679,1703]" captionTargetBox="[151,1418,273,1635]" captionTargetId="figure-64@15.[151,1435,273,1655]" captionTargetPageId="15" captionText="FIGURES 3643. Anhoraeomorphus procerus (Franz), holotype. Habitus in dorsal view (36); head in anterodorsal view (37); head and prothorax in lateral view (only bristles on head are shown) (38); antenna (39); aedeagus in ventral (40, 42) and lateral (41, 43) views." figureDoi="http://doi.org/10.5281/zenodo.13745935" httpUri="https://zenodo.org/record/13745935/files/figure.png" pageId="15" pageNumber="16">Fig. 38</figureCitation>
) slightly longer than eye; vertex and frons confluent, together weakly convex and about as long as broad; posterior margin of vertex truncate at middle; each eye large and strongly convex, with straight posteromedian margin. Punctures on vertex fine, inconspicuous; setae short, sparse and suberect; frons and vertex densely covered with thick bristles directed posteriorly. Antenna (
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) slender but short, slightly shorter than half BL, AnL
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, antennomeres 12 elongate, 3 about as long as broad, 47 each elongate, 810 each about as long as broad; 11 shorter than 9 and 10 combined, as wide as 10, nearly twice as long as broad.
</paragraph>
<paragraph id="8BF14E46FFE05010FF73FEEBFE6B4586" blockId="16.[151,1437,151,1041]" pageId="16" pageNumber="17">
Pronotum (
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) bell-shaped, broadest near middle; PL
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, PW
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. Anterior margin nearly straight, lateral margins rounded, more strongly converging anteriorly than posteriorly; posterior corners blunt; posterior margin indistinctly convex. Base with two pairs of pits: inner pits conspicuously large, deep, outer pits barely discernible, small and shallow, and with barely discernible, extremely short vestiges of sublateral carinae. Disc with fine, inconspicuous punctures; setae sparse, short and suberect, lateral and lateroventral surface of pronotum with dense thick bristles.
</paragraph>
<paragraph id="8BF14E46FFE05010FF73FE33FE7A4612" blockId="16.[151,1437,151,1041]" pageId="16" pageNumber="17">
Elytra (
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) together oval, broadest slightly anterior to middle; EL
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, EW
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, EI 1.48; humeral calli weakly elevated, basal elytral foveae vestigial, barely discernible. Punctures fine and inconspicuous; setae sparse, short and suberect.
</paragraph>
<paragraph id="8BF14E46FFE05010FF73FDA7FBCC46F6" blockId="16.[151,1437,151,1041]" box="[199,1033,619,645]" pageId="16" pageNumber="17">Legs moderately long, slender, unmodified except slightly recurved tibiae.</paragraph>
<paragraph id="8BF14E46FFE05010FF73FD43FABF474A" blockId="16.[151,1437,151,1041]" pageId="16" pageNumber="17">
Aedeagus (
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) slender, AeL
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; in ventral view broadest near base and weakly narrowing towards step-wise delimited apical region, which is subtriangular with truncate apex; in lateral view apical region of dorsal plate nearly straight; endophallus with slightly asymmetrical and elongate distolateral sclerites and asymmetrical median structures, distolateral sclerites broadly separated, each with hook-like apex curved mesally; parameres thick, shorter than median lobe, each paramere with one long apical and four subapical setae of various lengths.
</paragraph>
<paragraph id="8BF14E46FFE05010FF73FC8FFE5B472E" blockId="16.[151,1437,151,1041]" box="[199,414,835,861]" pageId="16" pageNumber="17">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFE05010FF73FCAAFCCF47F2" box="[199,778,870,897]" pageId="16" pageNumber="17" type="distribution">
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<emphasis id="B93A9254FFE05010FF73FCAAFEA447F3" bold="true" box="[199,353,870,896]" pageId="16" pageNumber="17">Distribution.</emphasis>
Northern
<collectingCountry id="F3590ED6FFE05010FE60FCABFD9E47F2" box="[468,603,871,897]" name="Madagascar" pageId="16" pageNumber="17">Madagascar</collectingCountry>
(Sava Region).
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFE05010FF73FC47FAF54062" pageId="16" pageNumber="17" type="discussion">
<paragraph id="8BF14E46FFE05010FF73FC47FB47479E" blockId="16.[151,1437,151,1041]" pageId="16" pageNumber="17">
<emphasis id="B93A9254FFE05010FF73FC47FEFE47D6" bold="true" box="[199,315,907,933]" pageId="16" pageNumber="17">Remarks.</emphasis>
<taxonomicName id="4C4E35C5FFE05010FEF1FC47FD4747D7" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[325,642,907,933]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="procerus">
<emphasis id="B93A9254FFE05010FEF1FC47FD4747D7" box="[325,642,907,933]" italics="true" pageId="16" pageNumber="17">Anhoraeomorphus procerus</emphasis>
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was described based on
<specimenCount id="9D4885CFFFE05010FC10FC47FB9547D6" box="[932,1104,907,933]" count="2" pageId="16" pageNumber="17" type="generic">two specimens</specimenCount>
; only the
<typeStatus id="54F5F0E4FFE05010FB71FC47FAE247D6" box="[1221,1319,907,933]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
male was studied. The
<typeStatus id="54F5F0E4FFE05010FE9FFC63FE4E47BA" box="[299,395,943,969]" pageId="16" pageNumber="17" type="paratype">paratype</typeStatus>
male is deposited in Muséum National d'Histoire Naturelle, Paris and was not available for my study. The village Ambinanitelo was misspelled by
<bibRefCitation id="EFDF33B7FFE05010FCB0FC1FFC66479E" author="Franz, H." box="[772,931,979,1005]" pageId="16" pageNumber="17" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986b)</bibRefCitation>
as “Ambinaniteta”.
</paragraph>
<paragraph id="8BF14E46FFE05010FF73FC3BFAF54062" blockId="16.[151,1437,151,1041]" box="[199,1328,1015,1041]" pageId="16" pageNumber="17">
<taxonomicName id="4C4E35C5FFE05010FF73FC3BFDC74063" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[199,514,1015,1041]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="procerus">
<emphasis id="B93A9254FFE05010FF73FC3BFDC74063" box="[199,514,1015,1041]" italics="true" pageId="16" pageNumber="17">Anhoraeomorphus procerus</emphasis>
</taxonomicName>
is externally similar to
<taxonomicName id="4C4E35C5FFE05010FCBEFC3BFC674062" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[778,930,1015,1041]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ivelonanus">
<emphasis id="B93A9254FFE05010FCBEFC3BFC674062" box="[778,930,1015,1041]" italics="true" pageId="16" pageNumber="17">A. ivelonanus</emphasis>
</taxonomicName>
, see Remarks for the latter species.
</paragraph>
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</treatment>
</document>

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@ -0,0 +1,115 @@
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<emphasis id="B5A9EA9CFF807715FF79F069FE08042D" bold="true" box="[151,408,1570,1597]" italics="true" pageId="16" pageNumber="51">Dysomma robinsorum</emphasis>
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Mok
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<elevation id="0CF0D1BDFF9F770AFD87F6BEFD24031E" box="[617,692,244,271]" metricMagnitude="2" metricUnit="m" metricValue="1.15" pageId="15" pageNumber="50" unit="m" value="115.0">115 m</elevation>
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<emphasis id="B5A9EA9CFF9F770AFF50F764FE300357" box="[190,416,302,328]" italics="true" pageId="15" pageNumber="50">Sinomyrus angustus</emphasis>
<materialsCitation id="37B53CD3FF9F770AFE47F767FDAF0357" box="[425,575,301,327]" collectionCode="FESC, PRFRI" country="China" location="Hoitow" pageId="15" pageNumber="50" specimenCount="1" stateProvince="Hainan" typeStatus="holotype">
<bibRefCitation id="E34C4B7FFF9F770AFE47F767FDAF0357" author="Lin, S. - Y." box="[425,575,301,327]" pageId="15" pageNumber="50" pagination="93 - 96" refId="ref18255" refString="Lin, S. - Y. (1933) A new genus and three new species of marine fish from Hainan Island. Lingnan Science Journal, Canton, 12 (1), 93 - 96." type="journal article" year="1933">Lin 1933: 94</bibRefCitation>
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<figureCitation id="1FE62A0BFF9F770AFDA2F767FD000357" box="[588,656,301,327]" captionStart="FIGURE 1" captionStartId="4.[151,250,973,997]" captionTargetBox="[170,1418,181,949]" captionTargetId="figure-34@4.[170,1418,181,949]" captionTargetPageId="4" captionText="FIGURE 1. Ilyophis arx. Swarm of eels attracted to a baited camera near summit of seamount APEI 7 (4° 52' 57.36&quot; N, 141° 46' 45.48&quot; W, 3203 m) (photo grab from https://oceanexplorer.noaa.gov/explorations/18ccz/logs/photolog/photolog.html#cbpi=../ may25/media/DC03.html" figureDoi="http://doi.org/10.5281/zenodo.13746101" httpUri="https://zenodo.org/record/13746101/files/figure.png" pageId="15" pageNumber="50">Fig. 1</figureCitation>
.
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: FESC [now PRFRI] 405. Type locality:
<location id="82026055FF9F770AFB0DF767FAA80357" LSID="urn:lsid:plazi:treatment:0F748798FF9F770AFF79F6DDFCB60094:82026055FF9F770AFB0DF767FAA80357" box="[1251,1336,301,327]" country="China" name="Hoitow" pageId="15" pageNumber="50" stateProvince="Hainan">Hoitow</location>
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<collectingCountry id="FFCA761EFF9F770AFF50F71BFE95037B" box="[190,261,337,363]" name="China" pageId="15" pageNumber="50">China</collectingCountry>
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<treatmentCitation id="067C109FFF9F770AFF50F7C1FD3C03B5" author="Ginsburg, I." box="[190,684,394,421]" page="452" pageId="15" pageNumber="50" year="1951">
<taxonomicName id="40DD4D0DFF9F770AFF50F7C1FD3C03B5" ID-CoL="389ZC" authority="Ginsburg 1951: 452" authorityName="Ginsburg" authorityPageNumber="452" authorityYear="1951" box="[190,684,394,421]" family="Synaphobranchidae" genus="Dysomma" kingdom="Animalia" order="Anguilliformes" pageId="15" pageNumber="50" phylum="Chordata" rank="species" species="aphododera">
<emphasis id="B5A9EA9CFF9F770AFF50F7C1FE2D03B5" box="[190,445,395,421]" italics="true" pageId="15" pageNumber="50">Dysomma aphododera</emphasis>
<materialsCitation id="37B53CD3FF9F770AFE28F7C1FD3C03B5" box="[454,684,394,421]" collectionCode="USNM" elevation="92" latitude="26.5" location="Gulf" longLatPrecision="1241" longitude="-96.433334" pageId="15" pageNumber="50" specimenCode="USNM 154992" specimenCount="1" stateProvince="Mexico" typeStatus="holotype">
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.
<typeStatus id="5866882CFF9F770AFCE5F7C1FCE803B5" box="[779,888,395,421]" pageId="15" pageNumber="50" type="holotype">Holotype</typeStatus>
:
<specimenCode id="D77B9EF5FF9F770AFC6DF7C1FBA503B5" box="[899,1077,394,421]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="15" pageNumber="50" type="Museum">USNM 154992</specimenCode>
. Type locality:
<location id="82026055FF9F770AFB0AF7C1FA8703B5" LSID="urn:lsid:plazi:treatment:0F748798FF9F770AFF79F6DDFCB60094:82026055FF9F770AFB0AF7C1FA8703B5" box="[1252,1303,395,421]" latitude="26.5" longLatPrecision="1241" longitude="-96.433334" name="Gulf" pageId="15" pageNumber="50" stateProvince="Mexico">Gulf</location>
of
<collectingRegion id="4519F86CFF9F770AFAAEF7C1FA0803B5" box="[1344,1432,395,421]" country="Mexico" name="Mexico" pageId="15" pageNumber="50">Mexico</collectingRegion>
,
<geoCoordinate id="E2E95049FF9F770AFF50F7E5FEBD03D9" box="[190,301,431,457]" degrees="26" direction="north" minutes="30" orientation="latitude" pageId="15" pageNumber="50" precision="925" value="26.5">26° 30 N</geoCoordinate>
,
<geoCoordinate id="E2E95049FF9F770AFED6F7E5FE3B03D9" box="[312,427,431,457]" degrees="96" direction="west" minutes="26" orientation="longitude" pageId="15" pageNumber="50" precision="925" value="-96.433334">96° 26 W</geoCoordinate>
, 50 fm (
<quantity id="40259B6BFF9F770AFDE9F7E5FDD503D9" box="[519,581,431,457]" metricMagnitude="1" metricUnit="m" metricValue="9.2" pageId="15" pageNumber="50" unit="m" value="92.0">
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<taxonomicName id="40DD4D0DFF9F770AFF50F7A3FD050012" ID-CoL="389ZJ" authority="Matsubara 1936: 961" authorityName="Matsubara" authorityPageNumber="961" authorityYear="1936" box="[190,661,488,514]" family="Synaphobranchidae" genus="Dysomma" kingdom="Animalia" order="Anguilliformes" pageId="15" pageNumber="50" phylum="Chordata" rank="species" species="japonicus">
<emphasis id="B5A9EA9CFF9F770AFF50F7A3FE310012" box="[190,417,489,514]" italics="true" pageId="15" pageNumber="50">Dysomma japonicus</emphasis>
<materialsCitation id="37B53CD3FF9F770AFE49F7A2FD050012" box="[423,661,488,514]" collectionCode="FAKU" country="Japan" elevation="275" location="Nada" pageId="15" pageNumber="50" specimenCode="FAKU 1941" specimenCount="1" typeStatus="holotype">
<bibRefCitation id="E34C4B7FFF9F770AFE49F7A2FD050012" author="Matsubara, K." box="[423,661,488,514]" pageId="15" pageNumber="50" pagination="960 - 962" refId="ref18294" refString="Matsubara, K. (1936) Studies on the deep-sea fishes of Japan. I. On a new apodal fish, Dysomma japonicus, with an emendation of the genus Dysomma. Zoological Magazine Tokyo, 48 (11), 960 - 962." type="journal article" year="1936">Matsubara 1936: 961</bibRefCitation>
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, Fig. unnumb.
<typeStatus id="5866882CFF9F770AFCD5F7A2FC380012" box="[827,936,488,514]" pageId="15" pageNumber="50" type="holotype">Holotype</typeStatus>
:
<specimenCode id="D77B9EF5FF9F770AFC5FF7A2FBAD0012" box="[945,1085,488,514]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="15" pageNumber="50">FAKU 1941</specimenCode>
. Type locality:
<collectingCountry id="FFCA761EFF9F770AFB08F7A3FAB90013" box="[1254,1321,489,515]" name="Japan" pageId="15" pageNumber="50">Japan</collectingCountry>
, Kumano-
<location id="82026055FF9F770AFF50F446FF6C0036" LSID="urn:lsid:plazi:treatment:0F748798FF9F770AFF79F6DDFCB60094:82026055FF9F770AFF50F446FF6C0036" box="[190,252,524,550]" country="Japan" name="Nada" pageId="15" pageNumber="50">Nada</location>
,
<locationDeviation id="96D02FACFF9F770AFEE9F446FE770036" box="[263,487,524,550]" location="Kii Peninsula" pageId="15" pageNumber="50">SE of Kii Peninsula</locationDeviation>
, 150 fm (
<quantity id="40259B6BFF9F770AFDBFF446FD0E0036" box="[593,670,524,551]" metricMagnitude="2" metricUnit="m" metricValue="2.75" pageId="15" pageNumber="50" unit="m" value="275.0">
<elevation id="0CF0D1BDFF9F770AFDBFF446FD0E0036" box="[593,670,524,551]" metricMagnitude="2" metricUnit="m" metricValue="2.75" pageId="15" pageNumber="50" unit="m" value="275.0">275 m</elevation>
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<treatmentCitationGroup id="A7CD11A0FF9F770AFF50F40CFCB60094" pageId="15" pageNumber="50">
<treatmentCitation id="067C109FFF9F770AFF50F40CFD38004F" author="Norman, J. R." box="[190,680,581,608]" page="44" pageId="15" pageNumber="50" year="1939">
<taxonomicName id="40DD4D0DFF9F770AFF50F40CFD38004F" ID-CoL="389ZT" authority="Norman 1939: 44" authorityName="Norman" authorityPageNumber="44" authorityYear="1939" box="[190,680,581,608]" family="Synaphobranchidae" genus="Dysomma" kingdom="Animalia" order="Anguilliformes" pageId="15" pageNumber="50" phylum="Chordata" rank="species" species="zanzibarensis">
<emphasis id="B5A9EA9CFF9F770AFF50F40CFE5F004F" box="[190,463,581,607]" italics="true" pageId="15" pageNumber="50">Dysomma zanzibarensis</emphasis>
<materialsCitation id="37B53CD3FF9F770AFE37F40CFD38004F" box="[473,680,581,608]" collectionCode="BMNH" elevation="189" latitude="-5.648333" location="western Indian Ocean" longLatPrecision="21" longitude="39.261665" pageId="15" pageNumber="50" specimenCode="BMNH 1939.524" specimenCount="1" typeStatus="holotype">
<bibRefCitation id="E34C4B7FFF9F770AFE37F40CFD38004F" author="Norman, J. R." box="[473,680,581,608]" pageId="15" pageNumber="50" pagination="1 - 116" refId="ref18464" refString="Norman, J. R. (1939) Fishes. The John Murray Expedition 1933 - 34. Scientific Reports, John Murray Expedition, 7 (1), 1 - 116." type="journal article" year="1939">Norman 1939: 44</bibRefCitation>
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, Fig. 17.
<typeStatus id="5866882CFF9F770AFCF4F40FFC17004F" box="[794,903,581,607]" pageId="15" pageNumber="50" type="holotype">Holotype</typeStatus>
:
<specimenCode id="D77B9EF5FF9F770AFC7AF40CFBF30070" box="[916,1123,581,608]" collectionCode="BMNH" pageId="15" pageNumber="50">BMNH 1939.524</specimenCode>
.651. Type locality:
<location id="82026055FF9F770AFAABF40CFEC60093" LSID="urn:lsid:plazi:treatment:0F748798FF9F770AFF79F6DDFCB60094:82026055FF9F770AFAABF40CFEC60093" latitude="-5.648333" longLatPrecision="21" longitude="39.261665" name="western Indian Ocean" pageId="15" pageNumber="50">western Indian Ocean</location>
,
<geoCoordinate id="E2E95049FF9F770AFE8CF423FE7C0093" box="[354,492,617,644]" degrees="5" direction="south" minutes="38" orientation="latitude" pageId="15" pageNumber="50" precision="15" seconds="54" value="-5.648333">5° 38 54” S</geoCoordinate>
,
<geoCoordinate id="E2E95049FF9F770AFE17F423FD040094" box="[505,660,617,644]" degrees="39" direction="east" minutes="15" orientation="longitude" pageId="15" pageNumber="50" precision="15" seconds="42" value="39.261665">39° 15 42” E</geoCoordinate>
,
<quantity id="40259B6BFF9F770AFD4EF423FCB20094" box="[672,802,617,644]" metricMagnitude="2" metricUnit="m" metricValue="1.885" metricValueMax="1.94" metricValueMin="1.83" pageId="15" pageNumber="50" unit="m" value="188.5" valueMax="194.0" valueMin="183.0">
<elevation id="0CF0D1BDFF9F770AFD4EF423FCB20094" box="[672,802,617,644]" metricMagnitude="2" metricUnit="m" metricValue="1.885" metricValueMax="1.94" metricValueMin="1.83" pageId="15" pageNumber="50" unit="m" value="188.5" valueMax="194.0" valueMin="183.0">183194 m</elevation>
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View file

@ -0,0 +1,74 @@
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Genus:
<taxonomicName id="68914D73FFB38F67B6DFFE48D55AFDE0" ID-CoL="635DK" authority="Vavra, 1891" authorityName="Vavra" authorityYear="1891" box="[248,515,510,537]" class="Ostracoda" family="Cyprididae" genus="Eucypris" kingdom="Animalia" order="Podocopida" pageId="2" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="9DE5EAE2FFB38F67B6DFFE48D63DFDE0" bold="true" box="[248,356,511,537]" italics="true" pageId="2" pageNumber="60">Eucypris</emphasis>
<bibRefCitation id="CB004B01FFB38F67B74CFE49D55AFDE0" author="Vavra, W." box="[363,515,510,537]" pageId="2" pageNumber="60" pagination="1 - 116" refId="ref11727" refString="Vavra, W. (1891) Monographie der Ostracoden Bohmens. Archiv fur die naturwissenschaftliche Landesdurchforschung von Bohmen, 8, 1 - 116." type="journal article" year="1891">Vávra, 1891</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E78B657BFFB38F67B6B0FDF0D4F6FCC0" pageId="2" pageNumber="60" type="diagnosis">
<paragraph id="AF2E36F0FFB38F67B6B0FDF0D4F6FCC0" blockId="2.[151,1437,583,826]" pageId="2" pageNumber="60">
Diagnosis (modified from
<bibRefCitation id="CB004B01FFB38F67B7E2FDF0D502FD98" author="Meisch, C." box="[453,603,583,609]" pageId="2" pageNumber="60" pagination="1 - 522" refId="ref10628" refString="Meisch, C. (2000) Freshwater Ostracoda of Western and Central Europe. In: Schwoerbel, J. &amp; Zwick, P. (Eds.), Susswasserfauna von Mitteleuropa 8 / 3. Spektrum Akademischer Verlag, Berlin, pp. 1 - 522." type="book chapter" year="2000">Meisch 2000</bibRefCitation>
;
<bibRefCitation id="CB004B01FFB38F67B440FDF0D478FD98" author="Karanovic, I." box="[615,801,583,609]" pageId="2" pageNumber="60" refId="ref10112" refString="Karanovic, I. (2012) Recent freshwater ostracods of the world: Crustacea, Ostracoda, Podocopida. Springer, New York, 608 pp. https: // doi. org / 10.1007 / 978 - 3 - 642 - 21810 - 1" type="book" year="2012">Karanovic 2012</bibRefCitation>
). Carapace elliptical in lateral view. RV smooth in both, anterior and posterior margins. Hinge adont. Both valves with a broad inner lamella and a wide vestibulum. Selvage usually peripheral when present. Wart-like elevations (
<emphasis id="9DE5EAE2FFB38F67B4DAFD27D4D7FD50" box="[765,910,656,681]" italics="true" pageId="2" pageNumber="60">porenwarzen</emphasis>
) on anterior valve surface can be seen in some species. Anteroventral margin concave with a rounded expansion LV overlapping RV. A2 with swimming setae. Second segment of the Mxl palp cylindrical in shape and longer than broad. T1 with c-seta. T2 (walking leg) five-segmented, d1 seta (if present) longer than d2. T3 (cleaning leg) three-segmented and pincer organ present. UR well-developed with Triebel loop and basal triangular structure absent.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,301 @@
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<paragraph id="AF2E36F0FFB38F67B6B0FC3CD6E0FC5C" blockId="2.[151,441,907,969]" box="[151,441,907,934]" pageId="2" pageNumber="60">
<heading id="F466819CFFB38F67B6B0FC3CD6E0FC5C" bold="true" box="[151,441,907,934]" fontSize="11" level="1" pageId="2" pageNumber="60" reason="1">
<emphasis id="9DE5EAE2FFB38F67B6B0FC3CD6E0FC5C" bold="true" box="[151,441,907,934]" pageId="2" pageNumber="60">
<taxonomicName id="68914D73FFB38F67B6B0FC3CD636FC5F" authorityName="Külköylüoğlu &amp; Özdilek" authorityYear="2024" box="[151,367,907,934]" class="Ostracoda" family="Cyprididae" genus="Eucypris" kingdom="Animalia" order="Podocopida" pageId="2" pageNumber="60" phylum="Arthropoda" rank="species" species="denktasi" status="sp. nov.">
<emphasis id="9DE5EAE2FFB38F67B6B0FC3CD636FC5F" bold="true" box="[151,367,907,934]" italics="true" pageId="2" pageNumber="60">Eucypris denktasi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="86D65799FFB38F67B751FC3CD6E0FC5C" box="[374,441,907,933]" pageId="2" pageNumber="60" rank="species">n. sp.</taxonomicNameLabel>
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<subSubSection id="E78B657BFFB38F6DB6B0FC18D49DFE9C" lastPageId="8" lastPageNumber="66" pageId="2" pageNumber="60" type="description">
<paragraph id="AF2E36F0FFB38F67B6B0FC18D64BFC30" blockId="2.[151,441,907,969]" box="[151,274,943,969]" pageId="2" pageNumber="60">
(
<figureCitation id="37AA2A75FFB38F67B6B8FC18D653FC30" box="[159,266,943,969]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStartId-0="3.[151,250,1388,1412]" captionStartId-1="4.[151,250,1943,1967]" captionStartId-2="5.[151,250,1943,1967]" captionStartId-3="6.[151,250,1820,1844]" captionStartId-4="7.[151,250,1505,1529]" captionTargetBox-0="[183,1404,181,1365]" captionTargetBox-1="[285,1301,181,1919]" captionTargetBox-2="[201,1386,181,1919]" captionTargetBox-3="[151,1432,189,1796]" captionTargetBox-4="[152,1436,245,1482]" captionTargetId-0="figure-27@3.[183,1404,181,1365]" captionTargetId-1="figure-17@4.[285,1301,181,1919]" captionTargetId-2="figure-27@5.[201,1386,181,1919]" captionTargetId-3="figure-17@6.[151,1436,181,1796]" captionTargetId-4="figure-57@7.[151,1436,237,1482]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionText-0="FIGURE 2. Eucypris denktasi sp. nov. Female. (A) RV, external view. (B) LV, external view. (C) carapace in dorsal view. (D) muscle scars. Scale bar: 133 µm for A and C. 145 µm for B. 94 µm for D." captionText-1="FIGURE 3. Eucypris denktasi sp. nov. Female. (A) RV internal view. (B) RV internal view with enlarged anterior margin. (C) RV internal view with enlarged posterior margin. (D) LV internal view. (E) LV internal view with posterior margin extended. (F) LV internal view with anterior margin extended. Scale bars: 140 µm for A and D. 88 µm for B and C. 100 µm for E and F." captionText-2="FIGURE 4. Eucypris denktasi sp. nov. (A) A1. (B) A2. (C) Md. Scale bar: 75 µm for A. 100 µm for B. 90 µm for C." captionText-3="FIGURE 5. Eucypris denktasi sp. nov. (A) Mx1. (B) Mx1 vibratory plate. (C) T1. (D) T2. Scale bar: 100 µm for A. 70 µm for B. 460 µm for C. 40 µm for D." captionText-4="FIGURE 6. Eucypris denktasi sp. nov. (A) T3. (B) Terminal part of T3. (C) Hypostome. (D) Genital organ. (E) Rake-like organ. (F) Uropod with attachment. Scale bar: 70 µm for A and B. 150 µm for C. 200 µm for D. 155 µm for E.80 µm for F." pageId="2" pageNumber="60">Figs. 26</figureCitation>
)
</paragraph>
<paragraph id="AF2E36F0FFB38F67B6B0FC40D385FBE8" blockId="2.[151,1437,1015,2014]" box="[151,1244,1015,1041]" pageId="2" pageNumber="60">
Etymology. The species is named after Rauf Raif DenktaŞ, the first president of
<collectingCountry id="D7867660FFB38F67B230FC40D381FBE8" box="[1047,1240,1015,1041]" name="Northern Cyprus" pageId="2" pageNumber="60">Northern Cyprus</collectingCountry>
.
</paragraph>
<paragraph id="AF2E36F0FFB38F67B6E0FBABD6F2FBA0" blockId="2.[151,1437,1015,2014]" pageId="2" pageNumber="60">
<materialsCitation id="1FF93CADFFB38F67B6E0FBABD622FBA0" country="Northern Cyprus" county="Elevation" elevation="136" location="Temporary creek" municipality="LefkoSa" pageId="2" pageNumber="60" specimenCount="136">
Type locality:
<location id="AA4E602BFFB38F67B743FBABD57AFBCC" LSID="urn:lsid:plazi:treatment:273887E6FFB38F6DB6B0FC3CD6FCFD08:AA4E602BFFB38F67B743FBABD57AFBCC" box="[356,547,1051,1078]" country="Northern Cyprus" county="Elevation" municipality="LefkoSa" name="Temporary creek" pageId="2" pageNumber="60">Temporary creek</location>
,
<location id="AA4E602BFFB38F67B40DFBACD5A4FBCC" LSID="urn:lsid:plazi:treatment:273887E6FFB38F6DB6B0FC3CD6FCFD08:AA4E602BFFB38F67B40DFBACD5A4FBCC" box="[554,765,1051,1077]" country="Northern Cyprus" county="Elevation" municipality="LefkoSa" name="Yilmazkoy district" pageId="2" pageNumber="60">Yılmazköy district</location>
,
<collectingMunicipality id="4F4AAC8AFFB38F67B522FBACD43BFBCC" box="[773,866,1051,1077]" pageId="2" pageNumber="60">LefkoŞa</collectingMunicipality>
,
<collectingCountry id="D7867660FFB38F67B54CFBACD37FFBCC" box="[875,1062,1051,1077]" name="Northern Cyprus" pageId="2" pageNumber="60">Northern Cyprus</collectingCountry>
(35˚1293.2N, 033˚1076.7E).
<collectingCounty id="464F4E7CFFB38F67B6B0FB88D65BFBA0" box="[151,258,1087,1113]" pageId="2" pageNumber="60">Elevation</collectingCounty>
ca
<specimenCount id="B997FD79FFB38F67B70EFB88D622FBA0" box="[297,379,1087,1113]" count="136" pageId="2" pageNumber="60" type="generic">
<quantity id="68699B15FFB38F67B70EFB88D62EFBA0" box="[297,375,1087,1113]" metricMagnitude="2" metricUnit="m" metricValue="1.3599999999999999" pageId="2" pageNumber="60" unit="m" value="136.0">
<elevation id="24BCD1C3FFB38F67B70EFB88D62EFBA0" box="[297,375,1087,1113]" metricMagnitude="2" metricUnit="m" metricValue="1.3599999999999999" pageId="2" pageNumber="60" unit="m" value="136.0">136 m</elevation>
</quantity>
.
</specimenCount>
</materialsCitation>
a.s.l.
</paragraph>
<paragraph id="AF2E36F0FFB38F67B6E0FBD4D7A1FB15" blockId="2.[151,1437,1015,2014]" pageId="2" pageNumber="60">
Material examined.
<typeStatus id="702A8852FFB38F67B78CFBD4D541FB84" box="[427,536,1123,1149]" pageId="2" pageNumber="60" type="holotype">Holotype</typeStatus>
:
<specimenCount id="B997FD79FFB38F67B403FBD3D5F8FB84" box="[548,673,1123,1150]" count="1" pageId="2" pageNumber="60" type="female">one female</specimenCount>
dissected in lactophenol solution with soft body parts on one slide mounted and sealed with translucent nail polish (no: OK-CYP1); valves in micropalaeontological cavity slides (no: OK-CYP-V1). Collected from type locality on
<date id="DB2F1030FFB38F67B4C4FB1CD423FB3C" box="[739,890,1195,1222]" pageId="2" pageNumber="60" value="2022-05-21">21 May 2022</date>
by Okan Külköylüoğlu, Ahmet Özdilek and Ali Özdilek.
</paragraph>
<paragraph id="AF2E36F0FFB38F67B6E0FB44D6E9FACD" blockId="2.[151,1437,1015,2014]" pageId="2" pageNumber="60">
Dissected
<typeStatus id="702A8852FFB38F67B71EFB43D6F0FAF7" box="[313,425,1268,1294]" pageId="2" pageNumber="60" type="paratype">paratypes</typeStatus>
:
<specimenCount id="B997FD79FFB38F67B795FB44D576FAF4" box="[434,559,1267,1293]" count="6" pageId="2" pageNumber="60" type="female">six females</specimenCount>
from type locality (no: OK-CYP2-7). Collected by Okan Külköylüoğlu, Ahmet Özdilek and Ali Özdilek.
</paragraph>
<paragraph id="AF2E36F0FFB38F67B6E0FA8CD5F2FA85" blockId="2.[151,1437,1015,2014]" pageId="2" pageNumber="60">
Non-dissected
<typeStatus id="702A8852FFB38F67B753FA8BD6BDFAAF" box="[372,484,1340,1366]" pageId="2" pageNumber="60" type="paratype">paratypes</typeStatus>
: forty-eight females kept in 70% ethanol. Collected from type locality by Okan Külköylüoğlu, Ahmet Özdilek and Ali Özdilek.
</paragraph>
<paragraph id="AF2E36F0FFB38F67B6E0FA34D5C3FA38" blockId="2.[151,1437,1015,2014]" pageId="2" pageNumber="60">
<materialsCitation id="1FF93CADFFB38F67B6E0FA34D5C3FA38" country="Turkey" county="Abant Izzet Baysal University" location="Limnology Laboratory" municipality="Biology Department" pageId="2" pageNumber="60" specimenCount="2" stateProvince="Bolu" typeStatus="holotype">
Repository.
<typeStatus id="702A8852FFB38F67B777FA34D6E1FA64" box="[336,440,1411,1437]" pageId="2" pageNumber="60" type="holotype">Holotype</typeStatus>
and all
<typeStatus id="702A8852FFB38F67B432FA33D5D9FA67" box="[533,640,1412,1438]" pageId="2" pageNumber="60" type="paratype">paratypes</typeStatus>
stored at the
<location id="AA4E602BFFB38F67B53BFA34D344FA64" LSID="urn:lsid:plazi:treatment:273887E6FFB38F6DB6B0FC3CD6FCFD08:AA4E602BFFB38F67B53BFA34D344FA64" box="[796,1053,1411,1437]" country="Turkey" county="Abant Izzet Baysal University" municipality="Biology Department" name="Limnology Laboratory" pageId="2" pageNumber="60" stateProvince="Bolu">Limnology Laboratory</location>
of the
<collectingMunicipality id="4F4AAC8AFFB38F67B257FA34D200FA67" box="[1136,1369,1411,1438]" pageId="2" pageNumber="60">Biology Department</collectingMunicipality>
,
<collectingRegion id="6D55F812FFB38F67B341FA34D2C5FA64" box="[1382,1436,1411,1437]" country="Turkey" name="Bolu" pageId="2" pageNumber="60">Bolu</collectingRegion>
<collectingCounty id="464F4E7CFFB38F67B6B0FA10D6B3FA38" box="[151,490,1447,1476]" pageId="2" pageNumber="60">Abant İzzet Baysal University</collectingCounty>
,
<collectingRegion id="6D55F812FFB38F67B7D1FA10D576FA38" box="[502,559,1447,1473]" country="Turkey" name="Bolu" pageId="2" pageNumber="60">Bolu</collectingRegion>
,
<collectingCountry id="D7867660FFB38F67B41DFA10D5CFFA38" box="[570,662,1447,1473]" name="Turkey" pageId="2" pageNumber="60">Türkiye</collectingCountry>
.
</materialsCitation>
</paragraph>
<paragraph id="AF2E36F0FFB38F67B6E0FA7BD6F4F9F3" blockId="2.[151,1437,1015,2014]" pageId="2" pageNumber="60">Measurements (mm). Female (n=7): Length: 1.121.32 (mean= 1.22), Height: 0.610.78 (mean= 0.71), Width: 0.530.63 (mean= 0.58).</paragraph>
<paragraph id="AF2E36F0FFB38F67B6E0F9A4D470F963" blockId="2.[151,1437,1015,2014]" pageId="2" pageNumber="60">
Description: Carapace (
<figureCitation id="37AA2A75FFB38F67B7F1F9A4D51EF9D4" box="[470,583,1555,1581]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="3.[151,250,1388,1412]" captionStartId-1="4.[151,250,1943,1967]" captionTargetBox-0="[183,1404,181,1365]" captionTargetBox-1="[285,1301,181,1919]" captionTargetId-0="figure-27@3.[183,1404,181,1365]" captionTargetId-1="figure-17@4.[285,1301,181,1919]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 2. Eucypris denktasi sp. nov. Female. (A) RV, external view. (B) LV, external view. (C) carapace in dorsal view. (D) muscle scars. Scale bar: 133 µm for A and C. 145 µm for B. 94 µm for D." captionText-1="FIGURE 3. Eucypris denktasi sp. nov. Female. (A) RV internal view. (B) RV internal view with enlarged anterior margin. (C) RV internal view with enlarged posterior margin. (D) LV internal view. (E) LV internal view with posterior margin extended. (F) LV internal view with anterior margin extended. Scale bars: 140 µm for A and D. 88 µm for B and C. 100 µm for E and F." pageId="2" pageNumber="60">Figs. 23</figureCitation>
): In lateral view carapace ovoid, in dorsal view carapace anteriorly beak-shaped, LV&gt; RV, carapace surface smooth without wart-like elevations, selvage present in RV, LV with inner list close to the outer margin, hinge adont, greatest height in the middle of carapace, rounded margin present anteriorly and posteriorly, valve surface with normal pores and setae.
</paragraph>
<paragraph id="AF2E36F0FFB38F67B6E0F914D7A5F827" blockId="2.[151,1437,1015,2014]" pageId="2" pageNumber="60">
A1 (
<figureCitation id="37AA2A75FFB38F67B6DCF914D601F944" box="[251,344,1699,1725]" captionStart="FIGURE 4" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[201,1386,181,1919]" captionTargetId="figure-27@5.[201,1386,181,1919]" captionTargetPageId="5" captionText="FIGURE 4. Eucypris denktasi sp. nov. (A) A1. (B) A2. (C) Md. Scale bar: 75 µm for A. 100 µm for B. 90 µm for C." pageId="2" pageNumber="60">Fig. 4A</figureCitation>
): Seven-segmented. Wouters organ and Rome organ absent. Four setae on first segment: a thin dorsal smooth seta, a long and smooth seta ventrally, extending to the end of the second segment, one of the two unequally long and smooth ventro-apical setae reaching to the tips of sixth segment, short seta slightly extending beyond the third segment. The second segment partially quadrangular with a medium-sized smooth dorsal seta. The third segment with one dorsal and one ventral apical setae. The fourth segment with two very long dorso-apical setae, twice the total length of the last three segments and with two equally long smooth apical setae. The fifth segment with two very long dorso-apical setae and with two ventro-apical setae. Penultimate (sixth) segment with rather long four setae. Terminal segment with three long setae, aesthetasc ya smooth, twice the length of the terminal segment.
</paragraph>
<caption id="FBEE6678FFB28F66B6B0FADBD4DBFA51" pageId="3" pageNumber="61" startId="3.[151,250,1388,1412]" targetBox="[183,1404,181,1365]" targetPageId="3" targetType="figure">
<paragraph id="AF2E36F0FFB28F66B6B0FADBD4DBFA51" blockId="3.[151,1437,1388,1449]" pageId="3" pageNumber="61">
<emphasis id="9DE5EAE2FFB28F66B6B0FADBD64AFA7C" bold="true" box="[151,275,1388,1413]" pageId="3" pageNumber="61">FIGURE 2.</emphasis>
<taxonomicName id="68914D73FFB28F66B73DFADAD696FA7C" authorityName="Külköylüoğlu &amp; Özdilek" authorityYear="2024" box="[282,463,1389,1413]" class="Ostracoda" family="Cyprididae" genus="Eucypris" kingdom="Animalia" order="Podocopida" pageId="3" pageNumber="61" phylum="Arthropoda" rank="species" species="denktasi" status="sp. nov.">
<emphasis id="9DE5EAE2FFB28F66B73DFADAD696FA7C" box="[282,463,1389,1413]" italics="true" pageId="3" pageNumber="61">Eucypris denktasi</emphasis>
</taxonomicName>
<emphasis id="9DE5EAE2FFB28F66B7F1FADAD57EFA7C" bold="true" box="[470,551,1389,1413]" pageId="3" pageNumber="61">
<taxonomicNameLabel id="86D65799FFB28F66B7F1FADAD57EFA7C" box="[470,551,1389,1413]" pageId="3" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Female. (A) RV, external view. (B) LV, external view. (C) carapace in dorsal view. (D) muscle scars. Scale bar: 133 µm for A and C. 145 µm for B. 94 µm for D.
</paragraph>
</caption>
<paragraph id="AF2E36F0FFB28F66B6E0FA4CD41EF8CC" blockId="3.[151,1437,1531,2025]" pageId="3" pageNumber="61">
A2 (
<figureCitation id="37AA2A75FFB28F66B6DFFA4CD609F9EC" box="[248,336,1531,1557]" captionStart="FIGURE 4" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[201,1386,181,1919]" captionTargetId="figure-27@5.[201,1386,181,1919]" captionTargetPageId="5" captionText="FIGURE 4. Eucypris denktasi sp. nov. (A) A1. (B) A2. (C) Md. Scale bar: 75 µm for A. 100 µm for B. 90 µm for C." pageId="3" pageNumber="61">Fig. 4B</figureCitation>
): Five-segmented. First protopodal segment with two short postero-distal setae. Second protopodal segment with one long basal seta on the ventro-apical part. First endopodal segment with one long and two short exopodal setae (long seta reaching to the end of second segment) postero-apical seta plumose about the length of second segment, two-segmented aesthetasc Y smooth about one fourth of the segment in length, first five natatory setae long, extending the tips of the terminal claws, the sixth natatory seta short, about the length of second segment. Penultimate segment undivided with four unequally long t-setae (t4&gt;t3&gt;t2&gt;t1, t2 plumose) present, z-setae (z3&gt;z2&gt;z1) smooth, and two equally long smooth setae. Aesthetasc y2 absent. Claws (G1&gt;G3&gt;G2) well developed and serrated. Terminal (third endopodal segment) segment with one long
<collectingRegion id="6D55F812FFB28F66B5E6F940D4B7F8E8" box="[961,1006,1783,1809]" country="Iran" name="Qom" pageId="3" pageNumber="61">GM</collectingRegion>
and one Gm (shorter than
<collectingRegion id="6D55F812FFB28F66B334F940D21DF8E8" box="[1299,1348,1783,1809]" country="Iran" name="Qom" pageId="3" pageNumber="61">GM</collectingRegion>
), length of the y3 aesthetasc about the length of the terminal segment.
</paragraph>
<paragraph id="AF2E36F0FFB28F66B6E0F888D3E5F810" blockId="3.[151,1437,1531,2025]" pageId="3" pageNumber="61">
Md (
<figureCitation id="37AA2A75FFB28F66B6D9F888D600F8A0" box="[254,345,1855,1881]" captionStart="FIGURE 4" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[201,1386,181,1919]" captionTargetId="figure-27@5.[201,1386,181,1919]" captionTargetPageId="5" captionText="FIGURE 4. Eucypris denktasi sp. nov. (A) A1. (B) A2. (C) Md. Scale bar: 75 µm for A. 100 µm for B. 90 µm for C." pageId="3" pageNumber="61">Fig. 4C</figureCitation>
): Md-coxa with seven strong teeth and a thin inner seta and long, thin dorsal seta. Md-palp with four segments. First segment with two unequally long S1 and S2 setae plumose and a thin α seta, vibratory plate with six unequally long smooth setae; second segment with four plumose setae, a thick plumose β seta, and three long external setae; penultimate (third) segment with four dorsal setae, one thick cone-shaped plumose γ seta, three apical setae smooth and two ventral setae; terminal segment with three claw like smooth setae.
</paragraph>
<caption id="FBEE6678FFB58F61B6B0F820D235F80E" pageId="4" pageNumber="62" startId="4.[151,250,1943,1967]" targetBox="[285,1301,181,1919]" targetPageId="4" targetType="figure">
<paragraph id="AF2E36F0FFB58F61B6B0F820D235F80E" blockId="4.[151,1437,1943,2040]" pageId="4" pageNumber="62">
<emphasis id="9DE5EAE2FFB58F61B6B0F820D64AF856" bold="true" box="[151,275,1943,1967]" pageId="4" pageNumber="62">FIGURE 3.</emphasis>
<taxonomicName id="68914D73FFB58F61B73DF82FD697F856" authorityName="Külköylüoğlu &amp; Özdilek" authorityYear="2024" box="[282,462,1943,1967]" class="Ostracoda" family="Cyprididae" genus="Eucypris" kingdom="Animalia" order="Podocopida" pageId="4" pageNumber="62" phylum="Arthropoda" rank="species" species="denktasi" status="sp. nov.">
<emphasis id="9DE5EAE2FFB58F61B73DF82FD697F856" box="[282,462,1943,1967]" italics="true" pageId="4" pageNumber="62">Eucypris denktasi</emphasis>
</taxonomicName>
<emphasis id="9DE5EAE2FFB58F61B7F2F820D57FF856" bold="true" box="[469,550,1943,1967]" pageId="4" pageNumber="62">
<taxonomicNameLabel id="86D65799FFB58F61B7F2F820D57FF856" box="[469,550,1943,1967]" pageId="4" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Female. (A) RV internal view. (B) RV internal view with enlarged anterior margin. (C) RV internal view with enlarged posterior margin. (D) LV internal view. (E) LV internal view with posterior margin extended. (F) LV internal view with anterior margin extended. Scale bars: 140 µm for A and D. 88 µm for B and C. 100 µm for E and F.
</paragraph>
</caption>
<caption id="FBEE6678FFB48F60B6B0F820D260F856" box="[151,1337,1943,1968]" pageId="5" pageNumber="63" startId="5.[151,250,1943,1967]" targetBox="[201,1386,181,1919]" targetPageId="5" targetType="figure">
<paragraph id="AF2E36F0FFB48F60B6B0F820D260F856" blockId="5.[151,1337,1943,1968]" box="[151,1337,1943,1968]" pageId="5" pageNumber="63">
<emphasis id="9DE5EAE2FFB48F60B6B0F820D64AF856" bold="true" box="[151,275,1943,1967]" pageId="5" pageNumber="63">FIGURE 4.</emphasis>
<taxonomicName id="68914D73FFB48F60B73EF82FD697F856" authorityName="Külköylüoğlu &amp; Özdilek" authorityYear="2024" box="[281,462,1943,1967]" class="Ostracoda" family="Cyprididae" genus="Eucypris" kingdom="Animalia" order="Podocopida" pageId="5" pageNumber="63" phylum="Arthropoda" rank="species" species="denktasi" status="sp. nov.">
<emphasis id="9DE5EAE2FFB48F60B73EF82FD697F856" box="[281,462,1943,1967]" italics="true" pageId="5" pageNumber="63">Eucypris denktasi</emphasis>
</taxonomicName>
<emphasis id="9DE5EAE2FFB48F60B7F2F820D57CF856" bold="true" box="[469,549,1943,1967]" pageId="5" pageNumber="63">
<taxonomicNameLabel id="86D65799FFB48F60B7F2F820D57CF856" box="[469,549,1943,1967]" pageId="5" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A) A1. (B) A2. (C) Md. Scale bar: 75 µm for A. 100 µm for B. 90 µm for C.
</paragraph>
</caption>
<caption id="FBEE6678FFB78F63B6B0F8ABD689F8A1" pageId="6" pageNumber="64" startId="6.[151,250,1820,1844]" targetBox="[151,1432,189,1796]" targetPageId="6" targetType="figure">
<paragraph id="AF2E36F0FFB78F63B6B0F8ABD689F8A1" blockId="6.[151,1437,1820,1881]" pageId="6" pageNumber="64">
<emphasis id="9DE5EAE2FFB78F63B6B0F8ABD64AF8CC" bold="true" box="[151,275,1820,1845]" pageId="6" pageNumber="64">FIGURE 5.</emphasis>
<taxonomicName id="68914D73FFB78F63B73DF8AAD697F8CD" authorityName="Külköylüoğlu &amp; Özdilek" authorityYear="2024" box="[282,462,1820,1844]" class="Ostracoda" family="Cyprididae" genus="Eucypris" kingdom="Animalia" order="Podocopida" pageId="6" pageNumber="64" phylum="Arthropoda" rank="species" species="denktasi" status="sp. nov.">
<emphasis id="9DE5EAE2FFB78F63B73DF8AAD697F8CD" box="[282,462,1820,1844]" italics="true" pageId="6" pageNumber="64">Eucypris denktasi</emphasis>
</taxonomicName>
<emphasis id="9DE5EAE2FFB78F63B7F2F8AAD57CF8CC" bold="true" box="[469,549,1821,1845]" pageId="6" pageNumber="64">
<taxonomicNameLabel id="86D65799FFB78F63B7F2F8AAD57CF8CC" box="[469,549,1821,1845]" pageId="6" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A) Mx1. (B) Mx1 vibratory plate. (C) T1. (D) T2. Scale bar: 100 µm for A. 70 µm for B. 460 µm for C. 40 µm for D.
</paragraph>
</caption>
<paragraph id="AF2E36F0FFB78F62B6E0F81CD346FF2C" blockId="6.[151,1437,1963,2025]" lastBlockId="7.[151,1436,151,213]" lastPageId="7" lastPageNumber="65" pageId="6" pageNumber="64">
Mx1 (
<figureCitation id="37AA2A75FFB78F63B72CF81CD6D6F83C" box="[267,399,1963,1990]" captionStart="FIGURE 5" captionStartId="6.[151,250,1820,1844]" captionTargetBox="[151,1432,189,1796]" captionTargetId="figure-17@6.[151,1436,181,1796]" captionTargetPageId="6" captionText="FIGURE 5. Eucypris denktasi sp. nov. (A) Mx1. (B) Mx1 vibratory plate. (C) T1. (D) T2. Scale bar: 100 µm for A. 70 µm for B. 460 µm for C. 40 µm for D." pageId="6" pageNumber="64">Figs. 5A, B</figureCitation>
): with three endites and two-segmented palp; vibratory plate with 20 setose setae; first endite with 12 plumose and two smooth setae; second endite with nine setae; third endite with six setae and two serrated teeth bristles (Zahnborsten). First segment of Mxl palp with seven setae (one seta slightly plumose); second segment of Mxl-palp rectangular in shape with six (two claw-like and four smooth) setae.
</paragraph>
<caption id="FBEE6678FFB68F62B6B0FA56D268F9E4" pageId="7" pageNumber="65" startId="7.[151,250,1505,1529]" targetBox="[152,1436,245,1482]" targetPageId="7" targetType="figure">
<paragraph id="AF2E36F0FFB68F62B6B0FA56D268F9E4" blockId="7.[151,1437,1505,1566]" pageId="7" pageNumber="65">
<emphasis id="9DE5EAE2FFB68F62B6B0FA56D64BFA00" bold="true" box="[151,274,1505,1529]" pageId="7" pageNumber="65">FIGURE 6.</emphasis>
<taxonomicName id="68914D73FFB68F62B730FA55D692FA00" authorityName="Külköylüoğlu &amp; Özdilek" authorityYear="2024" box="[279,459,1505,1529]" class="Ostracoda" family="Cyprididae" genus="Eucypris" kingdom="Animalia" order="Podocopida" pageId="7" pageNumber="65" phylum="Arthropoda" rank="species" species="denktasi" status="sp. nov.">
<emphasis id="9DE5EAE2FFB68F62B730FA55D692FA00" box="[279,459,1505,1529]" italics="true" pageId="7" pageNumber="65">Eucypris denktasi</emphasis>
</taxonomicName>
<emphasis id="9DE5EAE2FFB68F62B7F7FA55D579FA03" bold="true" box="[464,544,1506,1530]" pageId="7" pageNumber="65">
<taxonomicNameLabel id="86D65799FFB68F62B7F7FA55D579FA03" box="[464,544,1506,1530]" pageId="7" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A) T3. (B) Terminal part of T3. (C) Hypostome. (D) Genital organ. (E) Rake-like organ. (F) Uropod with attachment. Scale bar: 70 µm for A and B. 150 µm for C. 200 µm for D. 155 µm for E.80 µm for F.
</paragraph>
</caption>
<paragraph id="AF2E36F0FFB68F62B6E0F9C7D55DF957" blockId="7.[151,1437,1647,2034]" pageId="7" pageNumber="65">
T1 (
<figureCitation id="37AA2A75FFB68F62B6D3F9C7D615F973" box="[244,332,1647,1674]" captionStart="FIGURE 5" captionStartId="6.[151,250,1820,1844]" captionTargetBox="[151,1432,189,1796]" captionTargetId="figure-17@6.[151,1436,181,1796]" captionTargetPageId="6" captionText="FIGURE 5. Eucypris denktasi sp. nov. (A) Mx1. (B) Mx1 vibratory plate. (C) T1. (D) T2. Scale bar: 100 µm for A. 70 µm for B. 460 µm for C. 40 µm for D." pageId="7" pageNumber="65">Fig. 5C</figureCitation>
): with two unequally long a setae, setae b, c, and d present; length of setae h: h2&gt;h1&gt; h3. Vibratory plate with five long setose setae.
</paragraph>
<paragraph id="AF2E36F0FFB68F62B6E0F90FD64FF8C7" blockId="7.[151,1437,1647,2034]" pageId="7" pageNumber="65">
T2 (
<figureCitation id="37AA2A75FFB68F62B6D1F90FD608F92B" box="[246,337,1719,1746]" captionStart="FIGURE 5" captionStartId="6.[151,250,1820,1844]" captionTargetBox="[151,1432,189,1796]" captionTargetId="figure-17@6.[151,1436,181,1796]" captionTargetPageId="6" captionText="FIGURE 5. Eucypris denktasi sp. nov. (A) Mx1. (B) Mx1 vibratory plate. (C) T1. (D) T2. Scale bar: 100 µm for A. 70 µm for B. 460 µm for C. 40 µm for D." pageId="7" pageNumber="65">Fig. 5D</figureCitation>
): Five-segmented. First segment with a short d1 seta; d2 and dp setae absent. e seta smooth, f and g setae slightly plumose (length ratio: e&gt; f&gt; g); seta h1 short, slightly extending the length of terminal segment, seta h2 a well-developed serrated claw, longer than the terminal three segments together. Seta h3 ½ the length of the h1 seta.
</paragraph>
<paragraph id="AF2E36F0FFB68F62B6E0F8FFD6EAF853" blockId="7.[151,1437,1647,2034]" pageId="7" pageNumber="65">
T3 (
<figureCitation id="37AA2A75FFB68F62B6D1F8FFD62BF89B" box="[246,370,1864,1890]" captionStart="FIGURE 6" captionStartId="7.[151,250,1505,1529]" captionTargetBox="[152,1436,245,1482]" captionTargetId="figure-57@7.[151,1436,237,1482]" captionTargetPageId="7" captionText="FIGURE 6. Eucypris denktasi sp. nov. (A) T3. (B) Terminal part of T3. (C) Hypostome. (D) Genital organ. (E) Rake-like organ. (F) Uropod with attachment. Scale bar: 70 µm for A and B. 150 µm for C. 200 µm for D. 155 µm for E.80 µm for F." pageId="7" pageNumber="65">Fig. 6A, B</figureCitation>
): Three-segmented. d1 and d2 setae smooth, dp seta plumose. Length ratio of d-setae: dp&gt; d2&gt; d1. Setae e and f plumose (length of seta f about half of seta e). Pincer organ present. Length ratio of h-setae: h3&gt; h2&gt; h1 (hook-like h2).
</paragraph>
<paragraph id="AF2E36F0FFB68F62B6E0F803D4A4F837" blockId="7.[151,1437,1647,2034]" box="[199,1021,1972,1998]" pageId="7" pageNumber="65">
Hypostome (
<figureCitation id="37AA2A75FFB68F62B77FF803D6E9F837" box="[344,432,1972,1998]" captionStart="FIGURE 6" captionStartId="7.[151,250,1505,1529]" captionTargetBox="[152,1436,245,1482]" captionTargetId="figure-57@7.[151,1436,237,1482]" captionTargetPageId="7" captionText="FIGURE 6. Eucypris denktasi sp. nov. (A) T3. (B) Terminal part of T3. (C) Hypostome. (D) Genital organ. (E) Rake-like organ. (F) Uropod with attachment. Scale bar: 70 µm for A and B. 150 µm for C. 200 µm for D. 155 µm for E.80 µm for F." pageId="7" pageNumber="65">Fig. 6C</figureCitation>
): with a wide base covered with thin pseudochaetae.
</paragraph>
<paragraph id="AF2E36F0FFB68F62B6E0F86FD308F80B" blockId="7.[151,1437,1647,2034]" box="[199,1105,2008,2034]" pageId="7" pageNumber="65">
Genital organ (
<figureCitation id="37AA2A75FFB68F62B748F86FD693F80B" box="[367,458,2008,2034]" captionStart="FIGURE 6" captionStartId="7.[151,250,1505,1529]" captionTargetBox="[152,1436,245,1482]" captionTargetId="figure-57@7.[151,1436,237,1482]" captionTargetPageId="7" captionText="FIGURE 6. Eucypris denktasi sp. nov. (A) T3. (B) Terminal part of T3. (C) Hypostome. (D) Genital organ. (E) Rake-like organ. (F) Uropod with attachment. Scale bar: 70 µm for A and B. 150 µm for C. 200 µm for D. 155 µm for E.80 µm for F." pageId="7" pageNumber="65">Fig. 6D</figureCitation>
): smooth and rounded with a hook-shape structure inside.
</paragraph>
<paragraph id="AF2E36F0FFB98F6DB6E0FF20D40EFF48" blockId="8.[151,1437,151,753]" box="[199,855,151,177]" pageId="8" pageNumber="66">
Rake-like organ (
<figureCitation id="37AA2A75FFB98F6DB7ADFF20D6B8FF48" box="[394,481,151,177]" captionStart="FIGURE 6" captionStartId="7.[151,250,1505,1529]" captionTargetBox="[152,1436,245,1482]" captionTargetId="figure-57@7.[151,1436,237,1482]" captionTargetPageId="7" captionText="FIGURE 6. Eucypris denktasi sp. nov. (A) T3. (B) Terminal part of T3. (C) Hypostome. (D) Genital organ. (E) Rake-like organ. (F) Uropod with attachment. Scale bar: 70 µm for A and B. 150 µm for C. 200 µm for D. 155 µm for E.80 µm for F." pageId="8" pageNumber="66">Fig. 6E</figureCitation>
): with well-developed nine teeth.
</paragraph>
<paragraph id="AF2E36F0FFB98F6DB6E0FF0CD619FEE4" blockId="8.[151,1437,151,753]" pageId="8" pageNumber="66">
Uropod (
<figureCitation id="37AA2A75FFB98F6DB717FF0CD6D2FF2C" box="[304,395,187,213]" captionStart="FIGURE 6" captionStartId="7.[151,250,1505,1529]" captionTargetBox="[152,1436,245,1482]" captionTargetId="figure-57@7.[151,1436,237,1482]" captionTargetPageId="7" captionText="FIGURE 6. Eucypris denktasi sp. nov. (A) T3. (B) Terminal part of T3. (C) Hypostome. (D) Genital organ. (E) Rake-like organ. (F) Uropod with attachment. Scale bar: 70 µm for A and B. 150 µm for C. 200 µm for D. 155 µm for E.80 µm for F." pageId="8" pageNumber="66">Fig. 6F</figureCitation>
): slightly curved, ventral margin smooth; Gp longer than half of Ga (both claws slightly serrated); posterior seta (sp) plumose and ca. sa = sp. Uropodal attachment smooth, thin and curved with a short ventral branch.
</paragraph>
<paragraph id="AF2E36F0FFB98F6DB6E0FE90D626FEB8" blockId="8.[151,1437,151,753]" box="[199,383,295,321]" pageId="8" pageNumber="66">Male: unknown.</paragraph>
<paragraph id="AF2E36F0FFB98F6DB6E0FEFCD49DFE9C" blockId="8.[151,1437,151,753]" box="[199,964,331,357]" pageId="8" pageNumber="66">
Accompanying ostracod species:
<taxonomicName id="68914D73FFB98F6DB418FEFCD4E6FE9C" authority="(Ramdohr, 1808)" baseAuthorityName="Ramdohr" baseAuthorityYear="1808" box="[575,959,331,357]" class="Ostracoda" family="Cyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="8" pageNumber="66" phylum="Arthropoda" rank="species" species="gibba">
<emphasis id="9DE5EAE2FFB98F6DB418FEFCD5AFFE9C" box="[575,758,331,357]" italics="true" pageId="8" pageNumber="66">Ilyocypris gibba</emphasis>
(
<bibRefCitation id="CB004B01FFB98F6DB523FEFCD4EEFE9C" author="Ramdohr, A. F." box="[772,951,331,357]" pageId="8" pageNumber="66" pagination="85 - 93" refId="ref11410" refString="Ramdohr, A. F. (1808) Uber die Gattung Cypris Muller und drei zu derselben gehorige neue Arten. Magazin der Gesellschaft Naturforschender Freunde zu Berlin fur die Neuesten Entdeckungen in der Gesammten Naturkunde, 2, 85 - 93." type="journal article" year="1808">Ramdohr, 1808</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="E78B657BFFB98F6DB6E0FED9D464FD98" pageId="8" pageNumber="66" type="diagnosis">
<paragraph id="AF2E36F0FFB98F6DB6E0FED9D464FD98" blockId="8.[151,1437,151,753]" pageId="8" pageNumber="66">
<emphasis id="9DE5EAE2FFB98F6DB6E0FED9D61BFE71" bold="true" box="[199,322,366,392]" pageId="8" pageNumber="66">Diagnosis.</emphasis>
In lateral view, carapace smooth, ovoid, and anterior margin beak shaped. A1 without Wouters and Rome organs. A2 length of t-setae: t4&gt;t3&gt;t2&gt;t1, t-2 plumose, Aesthetasc Y and y3 present, y2 absent, natatory setae extending slightly beyond terminal claws. Md vibratory plate with six smooth setae. Terminal segment of Mx-palp rectangular with two well-developed claw-like and four smooth setae, vibratory plate with 20 plumose setae. Rake-like organ with nine teeth. T1 with five setae on vibratory plate (all plumose), length of h-setae: h2&gt;h1&gt;h3. T2 five-segmented, d1 seta short, d2 absent, length of e-g setae: e&gt;f&gt;g, f and g plumose. T3 three-segmented. Uropod well-developed, Gp more than 1/2 of the Ga, sp equals to sa.
</paragraph>
</subSubSection>
<subSubSection id="E78B657BFFB98F6DB6E0FDDCD6FCFD08" pageId="8" pageNumber="66" type="biology_ecology">
<paragraph id="AF2E36F0FFB98F6DB6E0FDDCD6FCFD08" blockId="8.[151,1437,151,753]" pageId="8" pageNumber="66">
<emphasis id="9DE5EAE2FFB98F6DB6E0FDDCD67FFD7C" bold="true" box="[199,294,619,645]" pageId="8" pageNumber="66">Ecology</emphasis>
:
<taxonomicName id="68914D73FFB98F6DB711FDDCD6A7FD7C" authorityName="Külköylüoğlu &amp; Özdilek" authorityYear="2024" box="[310,510,619,645]" class="Ostracoda" family="Cyprididae" genus="Eucypris" kingdom="Animalia" order="Podocopida" pageId="8" pageNumber="66" phylum="Arthropoda" rank="species" species="denktasi" status="sp. nov.">
<emphasis id="9DE5EAE2FFB98F6DB711FDDCD6A7FD7C" box="[310,510,619,645]" italics="true" pageId="8" pageNumber="66">Eucypris denktasi</emphasis>
</taxonomicName>
<emphasis id="9DE5EAE2FFB98F6DB420FDDCD538FD7C" bold="true" box="[519,609,619,645]" pageId="8" pageNumber="66">
<taxonomicNameLabel id="86D65799FFB98F6DB420FDDCD538FD7C" box="[519,609,619,645]" pageId="8" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was collected from a temporary creek water puddle with about
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of depth. Physical variables of the
<typeStatus id="702A8852FFB98F6DB7DDFD38D573FD50" box="[506,554,655,681]" pageId="8" pageNumber="66">type</typeStatus>
locality: elevation
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<quantity id="68699B15FFB98F6DB4D8FD38D415FD50" box="[767,844,655,681]" metricMagnitude="2" metricUnit="m" metricValue="1.3599999999999999" pageId="8" pageNumber="66" unit="m" value="136.0">136 m</quantity>
.
</specimenCount>
a.s.l., water temperature 18.16 °C, pH 7.98, electrical conductivity 26.30 mS/cm, oxidation reduction potential 99.8 mV, total dissolved solids 13.16 ppt, atmospheric pressure 14.526 mmHg.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,400 @@
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<mods:title id="6230B225CD4B134F41E7830237C062F9">Two new species of the genus Eucypris (Crustacea: Ostracoda) from Northern Cyprus</mods:title>
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<treatment id="273887E6FFB98F6BB6B0FCF4D7B4FE28" LSID="urn:lsid:plazi:treatment:273887E6FFB98F6BB6B0FCF4D7B4FE28" httpUri="http://treatment.plazi.org/id/273887E6FFB98F6BB6B0FCF4D7B4FE28" lastPageId="14" lastPageNumber="72" pageId="8" pageNumber="66">
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<emphasis id="9DE5EAE2FFB98F6DB6B0FCF4D6BEFCA4" bold="true" box="[151,487,834,861]" pageId="8" pageNumber="66">
<taxonomicName id="68914D73FFB98F6DB6B0FCF4D6C4FCA4" authorityName="Külköylüoğlu &amp; Özdilek" authorityYear="2024" box="[151,413,834,861]" class="Ostracoda" family="Cyprididae" genus="Eucypris" kingdom="Animalia" order="Podocopida" pageId="8" pageNumber="66" phylum="Arthropoda" rank="species" species="lefkosaensis" status="sp. nov.">
<emphasis id="9DE5EAE2FFB98F6DB6B0FCF4D6C4FCA4" bold="true" box="[151,413,834,861]" italics="true" pageId="8" pageNumber="66">Eucypris lefkosaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="86D65799FFB98F6DB783FCF4D6BEFCA4" box="[420,487,835,861]" pageId="8" pageNumber="66" rank="species">n. sp.</taxonomicNameLabel>
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<subSubSection id="E78B657BFFB98F68B6B0FCD0D6D4F996" lastPageId="13" lastPageNumber="71" pageId="8" pageNumber="66" type="description">
<paragraph id="AF2E36F0FFB98F6DB6B0FCD0D646FC78" blockId="8.[151,487,834,897]" box="[151,287,871,897]" pageId="8" pageNumber="66">
(
<figureCitation id="37AA2A75FFB98F6DB6B8FCD0D64EFC78" box="[159,279,871,897]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStart-2="FIGURE 9" captionStart-3="FIGURE 10" captionStart-4="FIGURE 11" captionStartId-0="9.[151,250,1374,1398]" captionStartId-1="10.[151,249,1943,1967]" captionStartId-2="11.[151,250,1373,1397]" captionStartId-3="12.[151,250,1825,1849]" captionStartId-4="13.[151,250,1373,1397]" captionTargetBox-0="[151,1436,181,1350]" captionTargetBox-1="[242,1345,181,1919]" captionTargetBox-2="[151,1436,181,1349]" captionTargetBox-3="[152,1436,181,1801]" captionTargetBox-4="[207,1380,185,1350]" captionTargetId-0="figure-27@9.[151,1436,181,1350]" captionTargetId-1="figure-17@10.[242,1345,181,1919]" captionTargetId-2="figure-27@11.[151,1436,181,1349]" captionTargetId-3="figure-17@12.[151,1436,181,1801]" captionTargetId-4="figure-27@13.[207,1380,181,1350]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="12" captionTargetPageId-4="13" captionText-0="FIGURE 7. Eucypris lefkosaensis sp. nov. Female. (A) RV, external view. (B) LV, external view. (C) carapace in dorsal view (leaning slightly to the right). (D) muscle scars. Scale bar: 200 µm for A and B. 160 µm for C. 53 µm for D." captionText-1="FIGURE 8. Eucypris lefkosaensis sp. nov. Female. (A) RV internal view. (B) RV internal view with enlarged anterior margin. (C) RV internal view with enlarged posterior margin. (D) LV internal view. (E) LV internal view with posterior margin extended. (F) LV internal view with anterior margin extended. Scale bars: 200 µm for A and D and C. 100 µm for B. 133 µm for E and F." captionText-2="FIGURE 9. Eucypris lefkosaensis sp. nov. (A) A1. (B) A2. (C) Md. Scale bar: 100 µm for A. 95 µm for B. 140 µm for C." captionText-3="FIGURE 10. Eucypris lefkosaensis sp. nov. (A) Mx1. (B) Mx1 vibratory plate. (C) T1. (D) T2. Scale bar: 110 µm for A. 70 µm for B. 90 µm for C. 80 µm for D." captionText-4="FIGURE 11. Eucypris lefkosaensis sp. nov. (A) Terminal part of T3. (B) T3. (C) Rake-like organ. (D) Genital organ. (E) Hypostome. (F) Uropod with attachment. Scale bar: 59 µm for A. 60 µm for B. 186 µm for C. 240 µm for D. 110 µm for E. 93 µm for F." pageId="8" pageNumber="66">Figs. 711</figureCitation>
)
</paragraph>
<paragraph id="AF2E36F0FFB98F6DB6B0FC18D2C8FC30" blockId="8.[151,1437,943,1977]" box="[151,1425,943,969]" pageId="8" pageNumber="66">
Etymology. The species is named after LefkoŞa, the capital city of
<collectingCountry id="D7867660FFB98F6DB5A5FC18D366FC30" box="[898,1087,943,969]" name="Northern Cyprus" pageId="8" pageNumber="66">Northern Cyprus</collectingCountry>
where the new species found.
</paragraph>
<paragraph id="AF2E36F0FFB98F6DB6E0FC64D455FBE8" blockId="8.[151,1437,943,1977]" pageId="8" pageNumber="66">
<materialsCitation id="1FF93CADFFB98F6DB6E0FC64D585FBE8" country="Northern Cyprus" county="LefkoSa" elevation="142" latitude="35.223305" location="Dar Dere" longLatPrecision="1" longitude="33.17689" municipality="Yilmazkoy district" pageId="8" pageNumber="66" specimenCount="142">
Type locality:
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, shallow puddle of a temporary creek,
<collectingMunicipality id="4F4AAC8AFFB98F6DB5B5FC64D331FC14" box="[914,1128,979,1005]" pageId="8" pageNumber="66">Yılmazköy district</collectingMunicipality>
,
<collectingCounty id="464F4E7CFFB98F6DB253FC64D388FC14" box="[1140,1233,979,1005]" pageId="8" pageNumber="66">LefkoŞa</collectingCounty>
,
<collectingCountry id="D7867660FFB98F6DB2F9FC64D2C5FC14" box="[1246,1436,979,1005]" name="Northern Cyprus" pageId="8" pageNumber="66">Northern Cyprus</collectingCountry>
(
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<geoCoordinate id="CAA55037FFB98F6DB761FC40D6BCFBE8" box="[326,485,1015,1041]" degrees="33" direction="east" minutes="10" orientation="longitude" pageId="8" pageNumber="66" precision="1" seconds="36.8" value="33.17689">33°1036.8”E</geoCoordinate>
). Elevation ca
<specimenCount id="B997FD79FFB98F6DB4ADFC40D585FBE8" box="[650,732,1015,1041]" count="142" pageId="8" pageNumber="66" type="generic">
<quantity id="68699B15FFB98F6DB4ADFC40D581FBE8" box="[650,728,1015,1041]" metricMagnitude="2" metricUnit="m" metricValue="1.42" pageId="8" pageNumber="66" unit="m" value="142.0">
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.
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a.s.l.
</paragraph>
<paragraph id="AF2E36F0FFB98F6DB6E0FBACD215FB86" blockId="8.[151,1437,943,1977]" pageId="8" pageNumber="66">
Material examined.
<typeStatus id="702A8852FFB98F6DB780FBACD54DFBCC" box="[423,532,1051,1077]" pageId="8" pageNumber="66" type="holotype">Holotype</typeStatus>
: Female dissected in lactophenol solution with soft body parts in a slide mounted and sealed with translucent nail polish (no: OK-CY1); valves in micropalaeontological cavity slides (no: OK-CY- V1). Collected from type locality on
<date id="DB2F1030FFB98F6DB411FBD4D592FB84" box="[566,715,1123,1149]" pageId="8" pageNumber="66" value="2022-05-21">21 May 2022</date>
by Okan Külköylüoğlu, Ahmet Özdilek and Ali Özdilek.
</paragraph>
<paragraph id="AF2E36F0FFB98F6DB6E0FB30D4CEFB58" blockId="8.[151,1437,943,1977]" box="[199,919,1159,1185]" pageId="8" pageNumber="66">
Dissected
<typeStatus id="702A8852FFB98F6DB71CFB30D6F2FB58" box="[315,427,1159,1185]" pageId="8" pageNumber="66" type="paratype">paratypes</typeStatus>
: a female from type locality (no: OK-CY2).
</paragraph>
<paragraph id="AF2E36F0FFB98F6DB6E0FB1CD55AFB12" blockId="8.[151,1437,943,1977]" pageId="8" pageNumber="66">
Non-dissected
<typeStatus id="702A8852FFB98F6DB74CFB1CD682FB3C" box="[363,475,1195,1221]" pageId="8" pageNumber="66" type="paratype">paratypes</typeStatus>
:
<specimenCount id="B997FD79FFB98F6DB7C5FB1CD5EBFB3C" box="[482,690,1195,1221]" count="34" pageId="8" pageNumber="66" type="female">thirty-four females</specimenCount>
in70%ethanol.Collected from type locality by Okan Külköylüoğlu, Ahmet Özdilek and Ali Özdilek.
</paragraph>
<paragraph id="AF2E36F0FFB98F6DB6E0FB44D33BFA80" blockId="8.[151,1437,943,1977]" pageId="8" pageNumber="66">
Other materials:
<materialsCitation id="1FF93CADFFB98F6DB7A9FB44D22AFAC8" collectingDate="2022-05-19" latitude="35.16003" location="Glapsides" longLatPrecision="1" longitude="33.91303" pageId="8" pageNumber="66" specimenCount="2" specimenCount-female="2" stateProvince="Famagusta">
1)
<collectingRegion id="6D55F812FFB98F6DB796FB44D5E9FAF4" box="[433,688,1267,1293]" country="Cyprus" name="Famagusta" pageId="8" pageNumber="66">Gazimağusa province</collectingRegion>
: one and
<specimenCount id="B997FD79FFB98F6DB501FB44D4EDFAF4" box="[806,948,1267,1293]" count="2" pageId="8" pageNumber="66" type="female">two females</specimenCount>
collected from coastal zone at
<location id="AA4E602BFFB98F6DB30AFB44D2C4FAF4" LSID="urn:lsid:plazi:treatment:273887E6FFB98F6BB6B0FCF4D7B4FE28:AA4E602BFFB98F6DB30AFB44D2C4FAF4" box="[1325,1437,1267,1293]" latitude="35.16003" longLatPrecision="1" longitude="33.91303" name="Glapsides" pageId="8" pageNumber="66" stateProvince="Famagusta">Glapsides</location>
wetlands on
<date id="DB2F1030FFB98F6DB713FAA0D681FAC8" box="[308,472,1303,1329]" pageId="8" pageNumber="66" value="2022-05-19">
<collectingDate id="CB6BE9D8FFB98F6DB713FAA0D681FAC8" box="[308,472,1303,1329]" pageId="8" pageNumber="66" value="2022-05-19">19 May 2022</collectingDate>
</date>
(
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<geoCoordinate id="CAA55037FFB98F6DB4BAFAA1D463FAC8" box="[669,826,1302,1329]" degrees="33" direction="east" minutes="54" orientation="longitude" pageId="8" pageNumber="66" precision="1" seconds="43.7" value="33.91214">33°5443.7”E</geoCoordinate>
and
<geoCoordinate id="CAA55037FFB98F6DB559FAA1D347FAC8" box="[894,1054,1302,1329]" degrees="35" direction="north" minutes="09" orientation="latitude" pageId="8" pageNumber="66" precision="1" seconds="36.1" value="35.16003">35°0936.1”N</geoCoordinate>
<geoCoordinate id="CAA55037FFB98F6DB20BFAA1D392FAC8" box="[1068,1227,1302,1329]" degrees="33" direction="east" minutes="54" orientation="longitude" pageId="8" pageNumber="66" precision="1" seconds="46.9" value="33.91303">33°5446.9”E</geoCoordinate>
, respectively)
</materialsCitation>
,
<materialsCitation id="1FF93CADFFB98F6DB3A2FAA0D304FA80" collectingDate="2022-05-21" latitude="35.27336" location="Yilmazkoy district" longLatPrecision="1" longitude="33.16942" municipality="Sirinevler district" pageId="8" pageNumber="66" specimenCount="31" specimenCount-female="31" stateProvince="Gazimagusa province">
2)
<collectingMunicipality id="4F4AAC8AFFB98F6DB6B0FA8CD63BFAAC" box="[151,354,1339,1365]" pageId="8" pageNumber="66">Şirinevler district</collectingMunicipality>
:
<specimenCount id="B997FD79FFB98F6DB749FA8CD6B2FAAC" box="[366,491,1339,1365]" count="1" pageId="8" pageNumber="66" type="female">one female</specimenCount>
from an unnamed creek (
<geoCoordinate id="CAA55037FFB98F6DB535FA8DD4EAFAAC" box="[786,947,1338,1365]" degrees="35" direction="north" minutes="16" orientation="latitude" pageId="8" pageNumber="66" precision="1" seconds="24.1" value="35.27336">35°1624.1”N</geoCoordinate>
<geoCoordinate id="CAA55037FFB98F6DB59CFA8CD302FAAC" box="[955,1115,1339,1365]" degrees="33" direction="east" minutes="10" orientation="longitude" pageId="8" pageNumber="66" precision="1" seconds="09.9" value="33.16942">33°1009.9”E</geoCoordinate>
), and 3)
<location id="AA4E602BFFB98F6DB2E7FA8CD2C0FAAC" LSID="urn:lsid:plazi:treatment:273887E6FFB98F6BB6B0FCF4D7B4FE28:AA4E602BFFB98F6DB2E7FA8CD2C0FAAC" box="[1216,1433,1339,1365]" latitude="35.27336" longLatPrecision="1" longitude="33.16942" municipality="Sirinevler district" name="Yilmazkoy district" pageId="8" pageNumber="66" stateProvince="Gazimagusa province">Yılmazköy district</location>
:
<specimenCount id="B997FD79FFB98F6DB6B0FAE8D66DFA80" box="[151,308,1375,1401]" count="30" pageId="8" pageNumber="66" type="female">thirty females</specimenCount>
from an unnamed creek (35˚1240N, 033˚1012E) on
<date id="DB2F1030FFB98F6DB5E4FAE8D304FA80" box="[963,1117,1375,1401]" pageId="8" pageNumber="66" value="2022-05-21">
<collectingDate id="CB6BE9D8FFB98F6DB5E4FAE8D304FA80" box="[963,1117,1375,1401]" pageId="8" pageNumber="66" value="2022-05-21">21 May 2022</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="AF2E36F0FFB98F6DB6E0FA34D5C3FA38" blockId="8.[151,1437,943,1977]" pageId="8" pageNumber="66">
<materialsCitation id="1FF93CADFFB98F6DB6E0FA34D5C3FA38" country="Turkey" county="Abant Izzet Baysal University" location="Limnology Laboratory" municipality="Biology Department" pageId="8" pageNumber="66" specimenCount="2" stateProvince="Bolu" typeStatus="holotype">
Repository.
<typeStatus id="702A8852FFB98F6DB777FA34D6E1FA64" box="[336,440,1411,1437]" pageId="8" pageNumber="66" type="holotype">Holotype</typeStatus>
and all
<typeStatus id="702A8852FFB98F6DB432FA34D5D9FA64" box="[533,640,1411,1437]" pageId="8" pageNumber="66" type="paratype">paratypes</typeStatus>
stored at the
<location id="AA4E602BFFB98F6DB53BFA34D344FA64" LSID="urn:lsid:plazi:treatment:273887E6FFB98F6BB6B0FCF4D7B4FE28:AA4E602BFFB98F6DB53BFA34D344FA64" box="[796,1053,1411,1437]" country="Turkey" county="Abant Izzet Baysal University" municipality="Biology Department" name="Limnology Laboratory" pageId="8" pageNumber="66" stateProvince="Bolu">Limnology Laboratory</location>
of the
<collectingMunicipality id="4F4AAC8AFFB98F6DB257FA34D200FA64" box="[1136,1369,1411,1437]" pageId="8" pageNumber="66">Biology Department</collectingMunicipality>
,
<collectingRegion id="6D55F812FFB98F6DB341FA34D2C5FA64" box="[1382,1436,1411,1437]" country="Turkey" name="Bolu" pageId="8" pageNumber="66">Bolu</collectingRegion>
<collectingCounty id="464F4E7CFFB98F6DB6B0FA10D6B3FA38" box="[151,490,1447,1475]" pageId="8" pageNumber="66">Abant İzzet Baysal University</collectingCounty>
,
<collectingRegion id="6D55F812FFB98F6DB7D1FA10D576FA38" box="[502,559,1447,1473]" country="Turkey" name="Bolu" pageId="8" pageNumber="66">Bolu</collectingRegion>
,
<collectingCountry id="D7867660FFB98F6DB41DFA10D5CFFA38" box="[570,662,1447,1473]" name="Turkey" pageId="8" pageNumber="66">Türkiye</collectingCountry>
.
</materialsCitation>
</paragraph>
<paragraph id="AF2E36F0FFB98F6DB6E0FA7CD6F4F9F0" blockId="8.[151,1437,943,1977]" pageId="8" pageNumber="66">Measurements (mm). Female (n=2): Length: 1.521.79 (mean= 1.66), Height: 0.871.06 (mean= 0.97), Width: 0.710.91 (mean= 0.81).</paragraph>
<paragraph id="AF2E36F0FFB98F6DB6E0F9A4D392F9D4" blockId="8.[151,1437,943,1977]" box="[199,1227,1555,1581]" pageId="8" pageNumber="66">
Description: Carapace (
<figureCitation id="37AA2A75FFB98F6DB7F7F9A4D56EF9D4" box="[464,567,1555,1581]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStartId-0="9.[151,250,1374,1398]" captionStartId-1="10.[151,249,1943,1967]" captionTargetBox-0="[151,1436,181,1350]" captionTargetBox-1="[242,1345,181,1919]" captionTargetId-0="figure-27@9.[151,1436,181,1350]" captionTargetId-1="figure-17@10.[242,1345,181,1919]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURE 7. Eucypris lefkosaensis sp. nov. Female. (A) RV, external view. (B) LV, external view. (C) carapace in dorsal view (leaning slightly to the right). (D) muscle scars. Scale bar: 200 µm for A and B. 160 µm for C. 53 µm for D." captionText-1="FIGURE 8. Eucypris lefkosaensis sp. nov. Female. (A) RV internal view. (B) RV internal view with enlarged anterior margin. (C) RV internal view with enlarged posterior margin. (D) LV internal view. (E) LV internal view with posterior margin extended. (F) LV internal view with anterior margin extended. Scale bars: 200 µm for A and D and C. 100 µm for B. 133 µm for E and F." pageId="8" pageNumber="66">Figs.78</figureCitation>
): Opaque whitish. In lateral view ovoid in shape, LV&gt; RV,
</paragraph>
<paragraph id="AF2E36F0FFB98F6DB6E0F980D4D6F960" blockId="8.[151,1437,943,1977]" pageId="8" pageNumber="66">LV with wider inner lamella than RV, inner list closer to the outer margin in LV, selvage present in RV, LV without selvage, hinge adont, greatest height found in the middle of the carapace, anterior and posterior margins rounded, carapace surface smooth with normal pores and thin setae.</paragraph>
<paragraph id="AF2E36F0FFB98F6DB6E0F914D4AEF840" blockId="8.[151,1437,943,1977]" pageId="8" pageNumber="66">
A1 (
<figureCitation id="37AA2A75FFB98F6DB6D0F914D612F944" box="[247,331,1699,1725]" captionStart="FIGURE 9" captionStartId="11.[151,250,1373,1397]" captionTargetBox="[151,1436,181,1349]" captionTargetId="figure-27@11.[151,1436,181,1349]" captionTargetPageId="11" captionText="FIGURE 9. Eucypris lefkosaensis sp. nov. (A) A1. (B) A2. (C) Md. Scale bar: 100 µm for A. 95 µm for B. 140 µm for C." pageId="8" pageNumber="66">Fig.9A</figureCitation>
): Seven-segmented. First segment without Wouters organ, and with two unequally long ventral setae and one smooth dorsal seta reaching about middle of the second segment. Second segment partially quadrangular with one smooth dorso-medial seta and without Rome organ; third segment with one long dorsal and two small ventro-apical setae; fourth segment with two unequally long ventro-apical setae and with two long dorsal setae; fifth segment with five unequally long setae; one long dorso-apical seta, reaching half way of terminal setae, one medium sized seta, and one very thin seta, two ventral setae long, extending ¾ of terminal setae; sixth (penultimate) segment with two equally long dorso-apical setae and three ventral setae; terminal (seventh) segment with three long setae and aesthetasc ya, three times longer than the length of the terminal segment.
</paragraph>
<caption id="FBEE6678FFB88F6CB6B0FAE9D385FA63" pageId="9" pageNumber="67" startId="9.[151,250,1374,1398]" targetBox="[151,1436,181,1350]" targetPageId="9" targetType="figure">
<paragraph id="AF2E36F0FFB88F6CB6B0FAE9D385FA63" blockId="9.[151,1436,1374,1435]" pageId="9" pageNumber="67">
<emphasis id="9DE5EAE2FFB88F6CB6B0FAE9D64DFA8E" bold="true" box="[151,276,1374,1399]" pageId="9" pageNumber="67">FIGURE 7.</emphasis>
<taxonomicName id="68914D73FFB88F6CB73CFAE8D6AFFA8F" authorityName="Külköylüoğlu &amp; Özdilek" authorityYear="2024" box="[283,502,1374,1398]" class="Ostracoda" family="Cyprididae" genus="Eucypris" kingdom="Animalia" order="Podocopida" pageId="9" pageNumber="67" phylum="Arthropoda" rank="species" species="lefkosaensis" status="sp. nov.">
<emphasis id="9DE5EAE2FFB88F6CB73CFAE8D6AFFA8F" box="[283,502,1374,1398]" italics="true" pageId="9" pageNumber="67">Eucypris lefkosaensis</emphasis>
</taxonomicName>
<emphasis id="9DE5EAE2FFB88F6CB7D9FAE8D516FA8E" bold="true" box="[510,591,1375,1399]" pageId="9" pageNumber="67">
<taxonomicNameLabel id="86D65799FFB88F6CB7D9FAE8D516FA8E" box="[510,591,1375,1399]" pageId="9" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Female. (A) RV, external view. (B) LV, external view. (C) carapace in dorsal view (leaning slightly to the right). (D) muscle scars. Scale bar: 200 µm for A and B. 160 µm for C. 53 µm for D.
</paragraph>
</caption>
<paragraph id="AF2E36F0FFB88F6CB6E0FA5AD506F926" blockId="9.[151,1437,1517,2011]" pageId="9" pageNumber="67">
A2 (
<figureCitation id="37AA2A75FFB88F6CB6DDFA5AD60CF9FE" box="[250,341,1517,1543]" captionStart="FIGURE 9" captionStartId="11.[151,250,1373,1397]" captionTargetBox="[151,1436,181,1349]" captionTargetId="figure-27@11.[151,1436,181,1349]" captionTargetPageId="11" captionText="FIGURE 9. Eucypris lefkosaensis sp. nov. (A) A1. (B) A2. (C) Md. Scale bar: 100 µm for A. 95 µm for B. 140 µm for C." pageId="9" pageNumber="67">Fig. 9B</figureCitation>
): Four-segmented. First segment with one smooth ventro-apical seta and three short basal setae. Second (first endopodal) segment with one long and two short exopodal setae, one short (sixth seta) and five long natatory setae extending tips of the terminal claws. Aesthetasc Y two-segmented. Penultimate (third) segment undivided with two long dorsal setae, four t-setae (t3&gt;t4&gt;t2&gt;t1, t1 and t2 plumose), aesthetasc y2 longer than terminal segment. z-setae (z3&gt;z2&gt;z1) unequally long. Claws (G1&gt;G3&gt;G2) well developed and serrated. Terminal segment with well-developed
<collectingRegion id="6D55F812FFB88F6CB7C3F916D548F942" box="[484,529,1697,1723]" country="Iran" name="Qom" pageId="9" pageNumber="67">GM</collectingRegion>
and a short Gm claw (short, about 2/4 of
<collectingRegion id="6D55F812FFB88F6CB5FFF916D350F942" box="[984,1033,1697,1723]" country="Iran" name="Qom" pageId="9" pageNumber="67">GM</collectingRegion>
), both serrated; aesthetasc y3 almost the same length as the terminal segment.
</paragraph>
<paragraph id="AF2E36F0FFB88F6CB6E0F95ED518F822" blockId="9.[151,1437,1517,2011]" pageId="9" pageNumber="67">
Md (
<figureCitation id="37AA2A75FFB88F6CB726F95ED604F8FA" box="[257,349,1769,1795]" captionStart="FIGURE 9" captionStartId="11.[151,250,1373,1397]" captionTargetBox="[151,1436,181,1349]" captionTargetId="figure-27@11.[151,1436,181,1349]" captionTargetPageId="11" captionText="FIGURE 9. Eucypris lefkosaensis sp. nov. (A) A1. (B) A2. (C) Md. Scale bar: 100 µm for A. 95 µm for B. 140 µm for C." pageId="9" pageNumber="67">Fig. 9C</figureCitation>
): Md-coxa with seven strong teeth, thin inner setae, and long, thin dorsal seta. Md-palp four-segmented; first segment with a long smooth seta, unequally long, plumose S1 and S2 setae, α setae thin and lunate. Vibratory plate with five plumose setae in different lengths. Second segment with four long plumose setae, β seta very thin, plumose, about the length of the second segment, and three external setae of almost equal length. Penultimate (third) segment with five long smooth external setae, γ seta robust with setules distally (reaching beyond the terminal segment), three smooth unequally long internal setae and two small setae. Terminal segment with two claws and two smooth setae.
</paragraph>
<caption id="FBEE6678FFBB8F6FB6B0F820D235F80E" pageId="10" pageNumber="68" startId="10.[151,249,1943,1967]" targetBox="[242,1345,181,1919]" targetPageId="10" targetType="figure">
<paragraph id="AF2E36F0FFBB8F6FB6B0F820D235F80E" blockId="10.[151,1437,1943,2040]" pageId="10" pageNumber="68">
<emphasis id="9DE5EAE2FFBB8F6FB6B0F820D649F856" bold="true" box="[151,272,1943,1967]" pageId="10" pageNumber="68">FIGURE 8.</emphasis>
<taxonomicName id="68914D73FFBB8F6FB731F82FD6B4F856" authorityName="Külköylüoğlu &amp; Özdilek" authorityYear="2024" box="[278,493,1943,1967]" class="Ostracoda" family="Cyprididae" genus="Eucypris" kingdom="Animalia" order="Podocopida" pageId="10" pageNumber="68" phylum="Arthropoda" rank="species" species="lefkosaensis" status="sp. nov.">
<emphasis id="9DE5EAE2FFBB8F6FB731F82FD6B4F856" box="[278,493,1943,1967]" italics="true" pageId="10" pageNumber="68">Eucypris lefkosaensis</emphasis>
</taxonomicName>
<emphasis id="9DE5EAE2FFBB8F6FB7D5F820D518F856" bold="true" box="[498,577,1943,1967]" pageId="10" pageNumber="68">
<taxonomicNameLabel id="86D65799FFBB8F6FB7D5F820D518F856" box="[498,577,1943,1967]" pageId="10" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Female. (A) RV internal view. (B) RV internal view with enlarged anterior margin. (C) RV internal view with enlarged posterior margin. (D) LV internal view. (E) LV internal view with posterior margin extended. (F) LV internal view with anterior margin extended. Scale bars: 200 µm for A and D and C. 100 µm for B. 133 µm for E and F.
</paragraph>
</caption>
<caption id="FBEE6678FFBA8F6EB6B0FAEAD232FA8C" box="[151,1387,1373,1397]" pageId="11" pageNumber="69" startId="11.[151,250,1373,1397]" targetBox="[151,1436,181,1349]" targetPageId="11" targetType="figure">
<paragraph id="AF2E36F0FFBA8F6EB6B0FAEAD232FA8C" blockId="11.[151,1387,1373,1397]" box="[151,1387,1373,1397]" pageId="11" pageNumber="69">
<emphasis id="9DE5EAE2FFBA8F6EB6B0FAEAD64AFA8C" bold="true" box="[151,275,1373,1397]" pageId="11" pageNumber="69">FIGURE 9.</emphasis>
<taxonomicName id="68914D73FFBA8F6EB73EFAEAD6ADFA8C" authorityName="Külköylüoğlu &amp; Özdilek" authorityYear="2024" box="[281,500,1373,1397]" class="Ostracoda" family="Cyprididae" genus="Eucypris" kingdom="Animalia" order="Podocopida" pageId="11" pageNumber="69" phylum="Arthropoda" rank="species" species="lefkosaensis" status="sp. nov.">
<emphasis id="9DE5EAE2FFBA8F6EB73EFAEAD6ADFA8C" box="[281,500,1373,1397]" italics="true" pageId="11" pageNumber="69">Eucypris lefkosaensis</emphasis>
</taxonomicName>
<emphasis id="9DE5EAE2FFBA8F6EB7DCFAEAD512FA8C" bold="true" box="[507,587,1373,1397]" pageId="11" pageNumber="69">
<taxonomicNameLabel id="86D65799FFBA8F6EB7DCFAEAD512FA8C" box="[507,587,1373,1397]" pageId="11" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A) A1. (B) A2. (C) Md. Scale bar: 100 µm for A. 95 µm for B. 140 µm for C.
</paragraph>
</caption>
<paragraph id="AF2E36F0FFBA8F6EB6E0FA70D3CBF9B7" blockId="11.[151,1437,1479,2010]" pageId="11" pageNumber="69">
Mx1 (
<figureCitation id="37AA2A75FFBA8F6EB729FA70D6C5FA18" box="[270,412,1479,1505]" captionStart="FIGURE 10" captionStartId="12.[151,250,1825,1849]" captionTargetBox="[152,1436,181,1801]" captionTargetId="figure-17@12.[151,1436,181,1801]" captionTargetPageId="12" captionText="FIGURE 10. Eucypris lefkosaensis sp. nov. (A) Mx1. (B) Mx1 vibratory plate. (C) T1. (D) T2. Scale bar: 110 µm for A. 70 µm for B. 90 µm for C. 80 µm for D." pageId="11" pageNumber="69">Figs.10AB</figureCitation>
): with three endites and two-segmented palps. First endite with 12 setae distally and two smooth setae at the base. Second and third endites with nine and six setae, respectively, both tooth bristles on third endite slightly serrated. First segment of palp with seven setae. Length of terminal palp segment slightly longer than broad and with four well-developed claw like setae. Vibratory plate with 18 plumose setae.
</paragraph>
<paragraph id="AF2E36F0FFBA8F6EB6E0F9E0D4A9F940" blockId="11.[151,1437,1479,2010]" pageId="11" pageNumber="69">
T1 (
<figureCitation id="37AA2A75FFBA8F6EB6D1F9E0D607F988" box="[246,350,1623,1649]" captionStart="FIGURE 10" captionStartId="12.[151,250,1825,1849]" captionTargetBox="[152,1436,181,1801]" captionTargetId="figure-17@12.[151,1436,181,1801]" captionTargetPageId="12" captionText="FIGURE 10. Eucypris lefkosaensis sp. nov. (A) Mx1. (B) Mx1 vibratory plate. (C) T1. (D) T2. Scale bar: 110 µm for A. 70 µm for B. 90 µm for C. 80 µm for D." pageId="11" pageNumber="69">Fig. 10C</figureCitation>
): Two small a setae equally long and plumose, b seta smooth, d seta smooth and long, extending tips of terminal setae, c seta 2/3 of d seta and plumose. Length ratio of h setae: h2&gt;h1&gt;h3, both h1-3 plumose. Vibratory plate with four plumose and one smooth setae of different lengths.
</paragraph>
<paragraph id="AF2E36F0FFBA8F6EB6E0F974D224F8F8" blockId="11.[151,1437,1479,2010]" pageId="11" pageNumber="69">
T2 (
<figureCitation id="37AA2A75FFBA8F6EB6D1F974D639F924" box="[246,352,1731,1757]" captionStart="FIGURE 10" captionStartId="12.[151,250,1825,1849]" captionTargetBox="[152,1436,181,1801]" captionTargetId="figure-17@12.[151,1436,181,1801]" captionTargetPageId="12" captionText="FIGURE 10. Eucypris lefkosaensis sp. nov. (A) Mx1. (B) Mx1 vibratory plate. (C) T1. (D) T2. Scale bar: 110 µm for A. 70 µm for B. 90 µm for C. 80 µm for D." pageId="11" pageNumber="69">Fig. 10D</figureCitation>
): Five-segmented. d1and d2 setae absent. Seta e plumose, f and g setae smooth (e=f&gt;g); h1 seta short, h2 seta well developed, serrated and claw-like, seta h3 present and as long as seta h1 (but see discussion).
</paragraph>
<paragraph id="AF2E36F0FFBA8F6EB6E0F8BCD528F894" blockId="11.[151,1437,1479,2010]" pageId="11" pageNumber="69">
T3 (
<figureCitation id="37AA2A75FFBA8F6EB6D3F8BCD6DDF8DC" box="[244,388,1803,1829]" captionStart="FIGURE 11" captionStartId="13.[151,250,1373,1397]" captionTargetBox="[207,1380,185,1350]" captionTargetId="figure-27@13.[207,1380,181,1350]" captionTargetPageId="13" captionText="FIGURE 11. Eucypris lefkosaensis sp. nov. (A) Terminal part of T3. (B) T3. (C) Rake-like organ. (D) Genital organ. (E) Hypostome. (F) Uropod with attachment. Scale bar: 59 µm for A. 60 µm for B. 186 µm for C. 240 µm for D. 110 µm for E. 93 µm for F." pageId="11" pageNumber="69">Figs. 11A, B</figureCitation>
): Three-segmented with smooth d1, d2 and dp setae. Length ratio of d-setae: dp&gt;d2&gt;d1. Seta e slightly plumose anteriorly, and seta f plumose (ca. length of f seta about half of seta e). Pincer organ present. Length ratio of h-setae: h3&gt;h2&gt;h1 (h1 hook-like).
</paragraph>
<paragraph id="AF2E36F0FFBA8F6EB6E0F8C0D5E3F868" blockId="11.[151,1437,1479,2010]" box="[199,698,1911,1938]" pageId="11" pageNumber="69">
Rake-like organ (
<figureCitation id="37AA2A75FFBA8F6EB7ADF8C0D6A9F868" box="[394,496,1911,1937]" captionStart="FIGURE 11" captionStartId="13.[151,250,1373,1397]" captionTargetBox="[207,1380,185,1350]" captionTargetId="figure-27@13.[207,1380,181,1350]" captionTargetPageId="13" captionText="FIGURE 11. Eucypris lefkosaensis sp. nov. (A) Terminal part of T3. (B) T3. (C) Rake-like organ. (D) Genital organ. (E) Hypostome. (F) Uropod with attachment. Scale bar: 59 µm for A. 60 µm for B. 186 µm for C. 240 µm for D. 110 µm for E. 93 µm for F." pageId="11" pageNumber="69">Fig. 11C</figureCitation>
): with eight teeth.
</paragraph>
<paragraph id="AF2E36F0FFBA8F6EB6E0F82CD58EF84C" blockId="11.[151,1437,1479,2010]" box="[199,727,1947,1974]" pageId="11" pageNumber="69">
Genital organ (
<figureCitation id="37AA2A75FFBA8F6EB748F82CD68EF84C" box="[367,471,1947,1973]" captionStart="FIGURE 11" captionStartId="13.[151,250,1373,1397]" captionTargetBox="[207,1380,185,1350]" captionTargetId="figure-27@13.[207,1380,181,1350]" captionTargetPageId="13" captionText="FIGURE 11. Eucypris lefkosaensis sp. nov. (A) Terminal part of T3. (B) T3. (C) Rake-like organ. (D) Genital organ. (E) Hypostome. (F) Uropod with attachment. Scale bar: 59 µm for A. 60 µm for B. 186 µm for C. 240 µm for D. 110 µm for E. 93 µm for F." pageId="11" pageNumber="69">Fig. 11D</figureCitation>
): smooth and rounded.
</paragraph>
<paragraph id="AF2E36F0FFBA8F6EB6E0F877D350F820" blockId="11.[151,1437,1479,2010]" box="[199,1033,1983,2010]" pageId="11" pageNumber="69">
Hypostome (
<figureCitation id="37AA2A75FFBA8F6EB77FF808D6E4F820" box="[344,445,1983,2009]" captionStart="FIGURE 11" captionStartId="13.[151,250,1373,1397]" captionTargetBox="[207,1380,185,1350]" captionTargetId="figure-27@13.[207,1380,181,1350]" captionTargetPageId="13" captionText="FIGURE 11. Eucypris lefkosaensis sp. nov. (A) Terminal part of T3. (B) T3. (C) Rake-like organ. (D) Genital organ. (E) Hypostome. (F) Uropod with attachment. Scale bar: 59 µm for A. 60 µm for B. 186 µm for C. 240 µm for D. 110 µm for E. 93 µm for F." pageId="11" pageNumber="69">Fig. 11E</figureCitation>
): with a wide base covered with thin pseudochaetae.
</paragraph>
<caption id="FBEE6678FFBD8F69B6B0F896D6B1F8A4" pageId="12" pageNumber="70" startId="12.[151,250,1825,1849]" targetBox="[152,1436,181,1801]" targetPageId="12" targetType="figure">
<paragraph id="AF2E36F0FFBD8F69B6B0F896D6B1F8A4" blockId="12.[151,1436,1825,1886]" pageId="12" pageNumber="70">
<emphasis id="9DE5EAE2FFBD8F69B6B0F896D646F8C0" bold="true" box="[151,287,1825,1849]" pageId="12" pageNumber="70">FIGURE 10.</emphasis>
<taxonomicName id="68914D73FFBD8F69B702F895D6A6F8C0" authorityName="Külköylüoğlu &amp; Özdilek" authorityYear="2024" box="[293,511,1825,1849]" class="Ostracoda" family="Cyprididae" genus="Eucypris" kingdom="Animalia" order="Podocopida" pageId="12" pageNumber="70" phylum="Arthropoda" rank="species" species="lefkosaensis" status="sp. nov.">
<emphasis id="9DE5EAE2FFBD8F69B702F895D6A6F8C0" box="[293,511,1825,1849]" italics="true" pageId="12" pageNumber="70">Eucypris lefkosaensis</emphasis>
</taxonomicName>
<emphasis id="9DE5EAE2FFBD8F69B422F895D50CF8C3" bold="true" box="[517,597,1826,1850]" pageId="12" pageNumber="70">
<taxonomicNameLabel id="86D65799FFBD8F69B422F895D50CF8C3" box="[517,597,1826,1850]" pageId="12" pageNumber="70" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A) Mx1. (B) Mx1 vibratory plate. (C) T1. (D) T2. Scale bar: 110 µm for A. 70 µm for B. 90 µm for C. 80 µm for D.
</paragraph>
</caption>
<paragraph id="AF2E36F0FFBD8F69B6E0F807D4A5F817" blockId="12.[151,1436,1968,2030]" pageId="12" pageNumber="70">
Uropod (
<figureCitation id="37AA2A75FFBD8F69B70AF807D6C9F833" box="[301,400,1968,1994]" captionStart="FIGURE 11" captionStartId="13.[151,250,1373,1397]" captionTargetBox="[207,1380,185,1350]" captionTargetId="figure-27@13.[207,1380,181,1350]" captionTargetPageId="13" captionText="FIGURE 11. Eucypris lefkosaensis sp. nov. (A) Terminal part of T3. (B) T3. (C) Rake-like organ. (D) Genital organ. (E) Hypostome. (F) Uropod with attachment. Scale bar: 59 µm for A. 60 µm for B. 186 µm for C. 240 µm for D. 110 µm for E. 93 µm for F." pageId="12" pageNumber="70">Fig. 11F</figureCitation>
): Claw Gp long, ca. 3/4 of claw Ga. Both claws serrated. Seta sp plumose; seta sa claw-like and smooth. Uropodal attachment slightly curved with a short ventral branch.
</paragraph>
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<paragraph id="AF2E36F0FFBC8F68B6B0FAEAD7AEFA44" blockId="13.[151,1437,1373,1470]" pageId="13" pageNumber="71">
<emphasis id="9DE5EAE2FFBC8F68B6B0FAEAD67BFA8C" bold="true" box="[151,290,1373,1397]" pageId="13" pageNumber="71">FIGURE 11.</emphasis>
<taxonomicName id="68914D73FFBC8F68B70CFAE9D551FA8F" authorityName="Külköylüoğlu &amp; Özdilek" authorityYear="2024" box="[299,520,1374,1398]" class="Ostracoda" family="Cyprididae" genus="Eucypris" kingdom="Animalia" order="Podocopida" pageId="13" pageNumber="71" phylum="Arthropoda" rank="species" species="lefkosaensis" status="sp. nov.">
<emphasis id="9DE5EAE2FFBC8F68B70CFAE9D551FA8F" box="[299,520,1374,1398]" italics="true" pageId="13" pageNumber="71">Eucypris lefkosaensis</emphasis>
</taxonomicName>
<emphasis id="9DE5EAE2FFBC8F68B435FAE9D53FFA8F" bold="true" box="[530,614,1374,1398]" pageId="13" pageNumber="71">
<taxonomicNameLabel id="86D65799FFBC8F68B435FAE9D53FFA8F" box="[530,614,1374,1398]" pageId="13" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A) Terminal part of T3. (B) T3. (C) Rake-like organ. (D) Genital organ. (E) Hypostome. (F) Uropod with attachment. Scale bar: 59 µm for A. 60 µm for B. 186 µm for C. 240 µm for D. 110 µm for E. 93 µm for F.
</paragraph>
</caption>
<paragraph id="AF2E36F0FFBC8F68B6E0FA5FD6D3F9FB" blockId="13.[151,1437,1512,2043]" box="[199,394,1512,1538]" pageId="13" pageNumber="71">Males: unknown.</paragraph>
<paragraph id="AF2E36F0FFBC8F68B6E0F9BBD6D4F996" blockId="13.[151,1437,1512,2043]" pageId="13" pageNumber="71">
Accompanying ostracod species:
<taxonomicName id="68914D73FFBC8F68B468F9BBD4E3F9DE" authority="(Jones, 1850)" baseAuthorityName="Jones" baseAuthorityYear="1850" box="[591,954,1548,1575]" class="Ostracoda" family="Cytherideidae" genus="Cyprideis" kingdom="Animalia" order="Podocopida" pageId="13" pageNumber="71" phylum="Arthropoda" rank="species" species="torosa">
<emphasis id="9DE5EAE2FFBC8F68B468F9BBD456F9DF" box="[591,783,1548,1574]" italics="true" pageId="13" pageNumber="71">Cyprideis torosa</emphasis>
(
<bibRefCitation id="CB004B01FFBC8F68B504F9BBD4EBF9DE" author="Jones, T. R." box="[803,946,1548,1575]" pageId="13" pageNumber="71" pagination="25 - 28" refId="ref10028" refString="Jones, T. R. (1850) Description of the Entomostraca of the Pleistocene beds of Newbury, Copford, Clacton, and Grays. Annals and Magazine of Natural History, Series 2, 6, 25 - 28." type="journal article" year="1850">Jones, 1850</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="68914D73FFBC8F68B5ECF9BBD3B3F9DF" baseAuthorityName="Costa" baseAuthorityYear="1847" box="[971,1258,1548,1574]" class="Ostracoda" family="Cyprididae" genus="Sarscypridopsis" kingdom="Animalia" order="Podocopida" pageId="13" pageNumber="71" phylum="Arthropoda" rank="species" species="aculeata">
<emphasis id="9DE5EAE2FFBC8F68B5ECF9BBD3B3F9DF" box="[971,1258,1548,1574]" italics="true" pageId="13" pageNumber="71">Sarscypridopsis aculeata</emphasis>
</taxonomicName>
(
<bibRefCitation id="CB004B01FFBC8F68B2D8F9BBD2C9F9DF" author="Costa, O. G." box="[1279,1424,1548,1574]" pageId="13" pageNumber="71" pagination="7 - 12" refId="ref9081" refString="Costa, O. G. (1847) Entomostraci, Ostracodi. Fauna del Regno di Napoli, ossia enumerazione di tutti gli animali che abitano la diverse regioni di questo Regno a le acque che le bagnano. Animali Articolati, Crostacei, 1, 7 - 12." type="journal article" year="1847">
<collectingRegion id="6D55F812FFBC8F68B2D8F9BBD21AF9DF" box="[1279,1347,1548,1574]" country="New Zealand" name="West Coast" pageId="13" pageNumber="71">Costa</collectingRegion>
, 1847
</bibRefCitation>
),
<taxonomicName id="68914D73FFBC8F68B6B0F986D5D7F9B3" authority="(Ramdohr, 1808)" baseAuthorityName="Ramdohr" baseAuthorityYear="1808" box="[151,654,1584,1610]" class="Ostracoda" family="Cyprididae" genus="Heterocypris" kingdom="Animalia" order="Podocopida" pageId="13" pageNumber="71" phylum="Arthropoda" rank="species" species="incongruens">
<emphasis id="9DE5EAE2FFBC8F68B6B0F986D6E4F9B3" box="[151,445,1585,1610]" italics="true" pageId="13" pageNumber="71">Heterocypris incongruens</emphasis>
(
<bibRefCitation id="CB004B01FFBC8F68B7F7F987D5DEF9B3" author="Ramdohr, A. F." box="[464,647,1584,1610]" pageId="13" pageNumber="71" pagination="85 - 93" refId="ref11410" refString="Ramdohr, A. F. (1808) Uber die Gattung Cypris Muller und drei zu derselben gehorige neue Arten. Magazin der Gesellschaft Naturforschender Freunde zu Berlin fur die Neuesten Entdeckungen in der Gesammten Naturkunde, 2, 85 - 93." type="journal article" year="1808">Ramdohr, 1808</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="68914D73FFBC8F68B4B9F986D4ECF9B3" authority="(Brady, 1868)" baseAuthorityName="Brady" baseAuthorityYear="1868" box="[670,949,1584,1610]" class="Ostracoda" family="Cyprididae" genus="Heterocypris" kingdom="Animalia" order="Podocopida" pageId="13" pageNumber="71" phylum="Arthropoda" rank="species" species="salina">
<emphasis id="9DE5EAE2FFBC8F68B4B9F986D451F9B3" box="[670,776,1584,1610]" italics="true" pageId="13" pageNumber="71">H. salina</emphasis>
(
<bibRefCitation id="CB004B01FFBC8F68B53CF987D4F4F9B3" author="Brady, G. S." box="[795,941,1584,1610]" pageId="13" pageNumber="71" pagination="353 - 495" refId="ref8942" refString="Brady, G. S. (1868) A Monograph of the Recent British Ostracoda. Transactions of the Linnean Society of London, 26 (2), 353 - 495. https: // doi. org / 10.1111 / j. 1096 - 3642.1968. tb 00199. x" type="journal article" year="1868">Brady, 1868</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="68914D73FFBC8F68B5E2F987D217F9B3" authority="(Ramdohr, 1808)" baseAuthorityName="Ramdohr" baseAuthorityYear="1808" box="[965,1358,1584,1610]" class="Ostracoda" family="Cyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="13" pageNumber="71" phylum="Arthropoda" rank="species" species="gibba">
<emphasis id="9DE5EAE2FFBC8F68B5E2F987D326F9B3" box="[965,1151,1584,1610]" italics="true" pageId="13" pageNumber="71">Ilyocypris gibba</emphasis>
(
<bibRefCitation id="CB004B01FFBC8F68B2B5F987D21CF9B3" author="Ramdohr, A. F." box="[1170,1349,1584,1610]" pageId="13" pageNumber="71" pagination="85 - 93" refId="ref11410" refString="Ramdohr, A. F. (1808) Uber die Gattung Cypris Muller und drei zu derselben gehorige neue Arten. Magazin der Gesellschaft Naturforschender Freunde zu Berlin fur die Neuesten Entdeckungen in der Gesammten Naturkunde, 2, 85 - 93." type="journal article" year="1808">Ramdohr, 1808</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="68914D73FFBC8F68B3ABF986D6D0F996" authority="Masi, 1905" authorityName="Masi" authorityYear="1905" class="Ostracoda" family="Cyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="13" pageNumber="71" phylum="Arthropoda" rank="species" species="decipiens">
<emphasis id="9DE5EAE2FFBC8F68B3ABF986D658F997" italics="true" pageId="13" pageNumber="71">I. decipiens</emphasis>
<bibRefCitation id="CB004B01FFBC8F68B72FF9E3D6D0F996" author="Masi, L." box="[264,393,1620,1647]" pageId="13" pageNumber="71" pagination="115 - 128" refId="ref10591" refString="Masi, L. (1905) Nota sugli Ostracodi viventi nei dintorni di Roma ed osservazioni sulla classificazione delle Cypridae. Bollettino della Societa Zoologica Italiana, Series 2, 6, 115 - 128." type="journal article" year="1905">Masi, 1905</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="E78B657BFFBC8F68B6E0F9CFD203F893" pageId="13" pageNumber="71" type="diagnosis">
<paragraph id="AF2E36F0FFBC8F68B6E0F9CFD203F893" blockId="13.[151,1437,1512,2043]" pageId="13" pageNumber="71">
<emphasis id="9DE5EAE2FFBC8F68B6E0F9CFD663F96B" bold="true" box="[199,314,1656,1682]" pageId="13" pageNumber="71">Diagnosis</emphasis>
. In lateral view, carapace smooth, ovoid, and anteriorly beak shaped. A1 without Wouters and Rome organs. A2 length of t-setae: t3&gt;t4&gt;t2&gt;t1, plumose t-1 and t-2, Aesthetasc Y, y2 and y3 present, natatory setae extending tips of the terminal claws. Md vibratory plate with five smooth setae. Terminal segment of Mxl-palp slightly longer than broad with four well-developed claw like setae, vibratory plate with 18 plumose setae. Rake-like organ with eight teeth. Vibratory plate on T1 with four plumose and one smooth setae. Length of setae-h: h2&gt;h1&gt;h3. T2 five-segmented, d1 and d2 setae absent, length of e and g setae: e=f&gt;g, e seta plumose. T3 three-segmented. Uropod well-developed, Ga claw more than 1/2 of the Gp claw, sp more than 1/2 of the Gp claw.
</paragraph>
</subSubSection>
<subSubSection id="E78B657BFFBC8F6BB6E0F8C3D7B4FE28" lastPageId="14" lastPageNumber="72" pageId="13" pageNumber="71" type="biology_ecology">
<paragraph id="AF2E36F0FFBC8F68B6E0F8C3D56CF803" blockId="13.[151,1437,1512,2043]" pageId="13" pageNumber="71">
<emphasis id="9DE5EAE2FFBC8F68B6E0F8C3D67FF877" bold="true" box="[199,294,1908,1934]" pageId="13" pageNumber="71">Ecology</emphasis>
:
<taxonomicName id="68914D73FFBC8F68B712F8C2D57EF877" authorityName="Külköylüoğlu &amp; Özdilek" authorityYear="2024" box="[309,551,1908,1934]" class="Ostracoda" family="Cyprididae" genus="Eucypris" kingdom="Animalia" order="Podocopida" pageId="13" pageNumber="71" phylum="Arthropoda" rank="species" species="lefkosaensis" status="sp. nov.">
<emphasis id="9DE5EAE2FFBC8F68B712F8C2D57EF877" box="[309,551,1908,1934]" italics="true" pageId="13" pageNumber="71">Eucypris lefkosaensis</emphasis>
</taxonomicName>
<emphasis id="9DE5EAE2FFBC8F68B408F8C3D5D0F877" bold="true" box="[559,649,1908,1934]" pageId="13" pageNumber="71">
<taxonomicNameLabel id="86D65799FFBC8F68B408F8C3D5D0F877" box="[559,649,1908,1934]" pageId="13" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was collected from a shallow puddle of a temporary creek with about
<quantity id="68699B15FFBC8F68B6B0F82FD786F84B" box="[151,223,1944,1970]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="71" unit="cm" value="10.0">10 cm</quantity>
of depth. Physical variables of the
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locality: elevation
<specimenCount id="B997FD79FFBC8F68B5B4F82FD4B1F84B" box="[915,1000,1944,1970]" count="142" pageId="13" pageNumber="71" type="generic">
<quantity id="68699B15FFBC8F68B5B4F82FD4BDF84B" box="[915,996,1944,1970]" metricMagnitude="2" metricUnit="m" metricValue="1.42" pageId="13" pageNumber="71" unit="m" value="142.0">142 m</quantity>
.
</specimenCount>
a.s.l., water temperature 20.83 °C, pH 8.06, electrical conductivity 32.99 mS/cm, oxidation reduction potential 99.8 mV, total dissolved solids 16,46 ppt, atmospheric pressure 14.532 mmHg.
</paragraph>
<paragraph id="AF2E36F0FFBF8F6BB6E0FF20D7B4FE28" blockId="14.[151,1436,151,465]" pageId="14" pageNumber="72">
Other locations: Glapsides wetland where one and
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were found from two sites: depth 20 and
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, respectively; elevation
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.
</specimenCount>
a.s.l., water temperature 25.24 and 28.27 °C, pH 9.35 and 9.14, electrical conductivity 37.17 and 39.99 mS/cm, oxidation reduction potential 63.5 and 20.3 mV, total dissolved solids 18.79 and 20.01 ppt, atmospheric pressure 14.734 and 14.739 mmHg, respectively. Unnamed creek in the Şirinevler district: depth
<quantity id="68699B15FFBF8F6BB3A7FEB4D7E5FEB8" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="72" unit="cm" value="10.0">10 cm</quantity>
, elevation
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, water temperature 17.21°C, pH 7.43, electrical conductivity 4.524 mS/cm, oxidation reduction potential 92 mV, total dissolved solids 2.262 ppt, atmospheric pressure 14.417 mmHg. Unnamed creek in the Yılmazköy district: depth
<quantity id="68699B15FFBF8F6BB79AFED8D55FFE70" box="[445,518,367,393]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" pageId="14" pageNumber="72" unit="cm" value="13.0">13 cm</quantity>
, elevation
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.
</specimenCount>
a.s.l., water temperature 18.48 °C, pH 8, electrical conductivity 27.78 mS/cm, oxidation reduction potential 101 mV, total dissolved solids 13.89 ppt, atmospheric pressure 14.545 mmHg.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,239 @@
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<paragraph id="057E36F2FFD5B435FF14FB6BD13DFACE" blockId="6.[151,1138,1274,1337]" box="[151,1138,1274,1301]" pageId="6" pageNumber="99">
<heading id="5E36819EFFD5B435FF14FB6BD13DFACE" bold="true" box="[151,1138,1274,1301]" fontSize="11" level="1" pageId="6" pageNumber="99" reason="1">
<emphasis id="37B5EAE0FFD5B435FF14FB6BD13DFACE" bold="true" box="[151,1138,1274,1301]" pageId="6" pageNumber="99">
Larva of
<taxonomicName id="C2C14D71FFD5B435FE88FB6AD688FACF" ID-CoL="9X2ML" authority="Bota-Sierra &amp; Novelo-Gutierrez" authorityName="Bota-Sierra &amp; Novelo-Gutierrez" authorityYear="2020" box="[267,967,1274,1301]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="99" phylum="Arthropoda" rank="species" species="brillantina">
<emphasis id="37B5EAE0FFD5B435FE88FB6AD763FACF" bold="true" box="[267,556,1274,1301]" italics="true" pageId="6" pageNumber="99">Epigomphus brillantina</emphasis>
Bota-Sierra &amp; Novelo-Gutiérrez
</taxonomicName>
(supposition)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="4DDB6579FFD5B435FF14FA8FD45DFAE3" box="[151,274,1311,1337]" pageId="6" pageNumber="99" type="description">
<paragraph id="057E36F2FFD5B435FF14FA8FD45DFAE3" blockId="6.[151,1138,1274,1337]" box="[151,274,1311,1337]" pageId="6" pageNumber="99">
(
<figureCitation id="9DFA2A77FFD5B435FF1CFA8FD445FAE3" box="[159,266,1311,1337]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStartId-0="7.[151,250,1268,1292]" captionStartId-1="8.[151,250,1943,1967]" captionStartId-2="9.[151,250,1168,1192]" captionTargetBox-0="[151,1436,181,1244]" captionTargetBox-1="[205,1383,181,1919]" captionTargetBox-2="[151,1435,181,1145]" captionTargetId-0="figure-17@7.[151,1436,181,1244]" captionTargetId-1="figure-23@8.[205,1383,181,1919]" captionTargetId-2="figure-17@9.[151,1435,181,1145]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionText-0="FIGURE 6. Epigomphus brillantina: a) F-1 female larva, dorsal view; b) Ídem, ventral view." captionText-1="FIGURE 7. Epigomphus brillantina: a) antennae, dorsal view; b) ventral view of mouth parts showing the hypopharynx; c) detail of the ligula and palps, ventral view (right movable hook broken); d) left hind leg; e) sterna 510; f) sterna 89 showing the female gonapophyses; g) S10 and caudal appendages, dorsal." captionText-2="FIGURE 8. Epigomphus brillantina larval habitat at Finca La Brillantina." pageId="6" pageNumber="99">Figs. 68</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="4DDB6579FFD5B435FF14FAF6D056FA13" pageId="6" pageNumber="99" type="materials_examined">
<paragraph id="057E36F2FFD5B435FF14FAF6D056FA13" blockId="6.[151,1437,1382,1877]" pageId="6" pageNumber="99">
<emphasis id="37B5EAE0FFD5B435FF14FAF6D449FA5A" bold="true" box="[151,262,1382,1408]" pageId="6" pageNumber="99">Material.</emphasis>
<materialsCitation id="B5A93CAFFFD5B435FE93FAF7D4ABFA5B" box="[272,484,1383,1409]" location="One" pageId="6" pageNumber="99" specimenCount="2" specimenCount-female="1" specimenCount-larva="1">
<location id="001E6029FFD5B435FE93FAF7D470FA5B" LSID="urn:lsid:plazi:treatment:8D6887E4FFD5B434FF14FB6BD026F897:001E6029FFD5B435FE93FAF7D470FA5B" box="[272,319,1383,1409]" name="One" pageId="6" pageNumber="99">One</location>
F-
<specimenCount id="13C7FD7BFFD5B435FEE5FAF7D4FEFA5B" box="[358,433,1383,1409]" count="1" pageId="6" pageNumber="99" type="larva">1 larva</specimenCount>
(
<specimenCount id="13C7FD7BFFD5B435FE45FAF7D496FA5B" box="[454,473,1383,1409]" count="1" pageId="6" pageNumber="99" type="female"></specimenCount>
)
</materialsCitation>
.
<materialsCitation id="B5A93CAFFFD5B435FE70FAF7D1BFFA7F" collectingDate="2019-02-21" country="Colombia" county="Remedios Municipality" elevation="593" latitude="6.90593" location="Finca La Brillantina" longLatPrecision="1" longitude="-74.57738" municipality="La Cruz Township" pageId="6" pageNumber="99" specimenCount="2" specimenCount-larva="2" stateProvince="Antioquia">
<collectingCountry id="7DD67662FFD5B435FE70FAF7D7C4FA5B" box="[499,651,1383,1409]" name="Colombia" pageId="6" pageNumber="99">COLOMBIA</collectingCountry>
,
<collectingRegion id="C705F810FFD5B435FD1BFAF7D6D6FA5B" box="[664,921,1383,1409]" country="Colombia" name="Antioquia" pageId="6" pageNumber="99">Antioquia Department</collectingRegion>
,
<collectingCounty id="EC1F4E7EFFD5B435FC24FAF7D1FCFA5B" box="[935,1203,1383,1409]" pageId="6" pageNumber="99">Remedios Municipality</collectingCounty>
,
<collectingMunicipality id="E51AAC88FFD5B435FB43FAF7D0D7FA5B" box="[1216,1432,1383,1409]" pageId="6" pageNumber="99">La Cruz Township</collectingMunicipality>
,
<location id="001E6029FFD5B435FF14FA1BD4CFFA7F" LSID="urn:lsid:plazi:treatment:8D6887E4FFD5B434FF14FB6BD026F897:001E6029FFD5B435FF14FA1BD4CFFA7F" box="[151,384,1419,1445]" country="Colombia" county="Remedios Municipality" latitude="6.90593" longLatPrecision="1" longitude="-74.57738" municipality="La Cruz Township" name="Finca La Brillantina" pageId="6" pageNumber="99" stateProvince="Antioquia">Finca La Brillantina</location>
, stream elev.
<quantity id="C2399B17FFD5B435FD9DFA1BD728FA7F" box="[542,615,1419,1446]" metricMagnitude="2" metricUnit="m" metricValue="5.93" pageId="6" pageNumber="99" unit="m" value="593.0">
<elevation id="8EECD1C1FFD5B435FD9DFA1BD728FA7F" box="[542,615,1419,1446]" metricMagnitude="2" metricUnit="m" metricValue="5.93" pageId="6" pageNumber="99" unit="m" value="593.0">593 m</elevation>
</quantity>
(
<geoCoordinate id="60F55035FFD5B435FDFBFA1BD7B9FA7C" box="[632,758,1419,1446]" degrees="6.90593" direction="north" orientation="latitude" pageId="6" pageNumber="99" precision="1" value="6.90593">6.90593°N</geoCoordinate>
, -
<geoCoordinate id="60F55035FFD5B435FC8FFA1BD6EFFA7C" box="[780,928,1419,1446]" degrees="74.57738" direction="west" orientation="longitude" pageId="6" pageNumber="99" precision="1" value="-74.57738">74.57738°W</geoCoordinate>
),
<date id="717F1032FFD5B435FC35FA1BD1CBFA7F" box="[950,1156,1419,1445]" pageId="6" pageNumber="99" value="2019-02-21">
<collectingDate id="613BE9DAFFD5B435FC35FA1BD1CBFA7F" box="[950,1156,1419,1445]" pageId="6" pageNumber="99" value="2019-02-21">21 February 2019</collectingDate>
</date>
,
<specimenCount id="13C7FD7BFFD5B435FB12FA1BD1BFFA7F" box="[1169,1264,1419,1445]" count="2" pageId="6" pageNumber="99" type="larva">2 larvae</specimenCount>
</materialsCitation>
;
<materialsCitation id="B5A93CAFFFD5B435FB7DFA1BD6ABFA13" collectingDate="2019-02-23" country="Colombia" county="Remedios Municipality" elevation="530" latitude="6.89664" location="Finca La Brillantina" longLatPrecision="1" longitude="-74.57579" municipality="La Cruz Township" pageId="6" pageNumber="99" specimenCount="8" specimenCount-larva="8" stateProvince="Antioquia">
same data but stream elev.
<quantity id="C2399B17FFD5B435FEA7FA3FD424FA13" box="[292,363,1455,1482]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="6" pageNumber="99" unit="m" value="530.0">
<elevation id="8EECD1C1FFD5B435FEA7FA3FD424FA13" box="[292,363,1455,1482]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="6" pageNumber="99" unit="m" value="530.0">530 m</elevation>
</quantity>
(
<geoCoordinate id="60F55035FFD5B435FEFAFA3FD4B7FA13" box="[377,504,1455,1481]" degrees="6.89664" direction="north" orientation="latitude" pageId="6" pageNumber="99" precision="1" value="6.89664">6.89664°N</geoCoordinate>
, -
<geoCoordinate id="60F55035FFD5B435FD88FA3FD7D0FA10" box="[523,671,1455,1482]" degrees="74.57579" direction="west" orientation="longitude" pageId="6" pageNumber="99" precision="1" value="-74.57579">74.57579°W</geoCoordinate>
),
<date id="717F1032FFD5B435FD31FA3FD632FA13" box="[690,893,1455,1481]" pageId="6" pageNumber="99" value="2019-02-23">
<collectingDate id="613BE9DAFFD5B435FD31FA3FD632FA13" box="[690,893,1455,1481]" pageId="6" pageNumber="99" value="2019-02-23">23 February 2019</collectingDate>
</date>
,
<specimenCount id="13C7FD7BFFD5B435FC0BFA3FD6ABFA13" box="[904,996,1455,1481]" count="8" pageId="6" pageNumber="99" type="larva">8 larvae</specimenCount>
</materialsCitation>
,
<materialsCitation id="B5A93CAFFFD5B435FC6CFA3FD056FA13" box="[1007,1305,1455,1481]" collectionCode="R" collectorName="Bota &amp; Novelo" country="Colombia" pageId="6" pageNumber="99" specimenCount="1" stateProvince="Antioquia">
all C.
<collectorName id="A8345324FFD5B435FBB0FA3FD124FA13" box="[1075,1131,1455,1481]" pageId="6" pageNumber="99">Bota</collectorName>
,
<collectionCode id="63D0AE37FFD5B435FBF6FA3FD1C4FA13" box="[1141,1163,1455,1481]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="99">R</collectionCode>
.
<collectorName id="A8345324FFD5B435FB15FA3FD1A7FA13" box="[1174,1256,1455,1481]" pageId="6" pageNumber="99">Novelo</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="4DDB6579FFD5B434FF44FA42D026F897" lastPageId="7" lastPageNumber="100" pageId="6" pageNumber="99" type="description">
<paragraph id="057E36F2FFD5B435FF44FA42D591F9EF" blockId="6.[151,1437,1382,1877]" pageId="6" pageNumber="99">
<emphasis id="37B5EAE0FFD5B435FF44FA42D416FA36" bold="true" box="[199,345,1490,1516]" pageId="6" pageNumber="99">Description.</emphasis>
Larva F-1
<quantity id="C2399B17FFD5B435FE5FFA43D70CFA37" box="[476,579,1491,1518]" metricMagnitude="-2" metricUnit="m" metricValue="2.55" pageId="6" pageNumber="99" unit="mm" value="25.5">25.5 mm</quantity>
total length, body robust, antennae, legs and sides of thorax setose, gently tapering caudad, body mostly reddish-brown, dark brown on S910 with a complex color pattern on abdomen (
<figureCitation id="9DFA2A77FFD5B435FAE7FA67D59DF9EF" captionStart="FIGURE 6" captionStartId="7.[151,250,1268,1292]" captionTargetBox="[151,1436,181,1244]" captionTargetId="figure-17@7.[151,1436,181,1244]" captionTargetPageId="7" captionText="FIGURE 6. Epigomphus brillantina: a) F-1 female larva, dorsal view; b) Ídem, ventral view." pageId="6" pageNumber="99">Figs. 6ab</figureCitation>
).
</paragraph>
<paragraph id="057E36F2FFD5B435FF44F9AFD043F933" blockId="6.[151,1437,1382,1877]" pageId="6" pageNumber="99">
<emphasis id="37B5EAE0FFD5B435FF44F9AFD44BF983" box="[199,260,1599,1625]" italics="true" pageId="6" pageNumber="99">Head</emphasis>
: As described for
<taxonomicName id="C2C14D71FFD5B435FE4CF9D0D768F983" authorityName="Bota-Sierra &amp; Novelo-Gutierrez" authorityYear="2020" box="[463,551,1599,1625]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="99" phylum="Arthropoda" rank="species" species="rufus">
<emphasis id="37B5EAE0FFD5B435FE4CF9D0D768F983" box="[463,551,1599,1625]" italics="true" pageId="6" pageNumber="99">E. rufus</emphasis>
</taxonomicName>
except: 3
<superScript id="F2B49BBAFFD5B435FD10F9AED7EEF996" attach="left" box="[659,673,1598,1612]" fontSize="6" pageId="6" pageNumber="99">rd</superScript>
antennomere 2x as long as widest part, 4
<superScript id="F2B49BBAFFD5B435FBE9F9AED138F996" attach="left" box="[1130,1143,1598,1612]" fontSize="6" pageId="6" pageNumber="99">th</superScript>
antennomere more or less subspherical (
<figureCitation id="9DFA2A77FFD5B435FEB6F9F3D4C7F9A7" box="[309,392,1635,1661]" captionStart="FIGURE 7" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[205,1383,181,1919]" captionTargetId="figure-23@8.[205,1383,181,1919]" captionTargetPageId="8" captionText="FIGURE 7. Epigomphus brillantina: a) antennae, dorsal view; b) ventral view of mouth parts showing the hypopharynx; c) detail of the ligula and palps, ventral view (right movable hook broken); d) left hind leg; e) sterna 510; f) sterna 89 showing the female gonapophyses; g) S10 and caudal appendages, dorsal." pageId="6" pageNumber="99">Fig. 7a</figureCitation>
), antennomeres proportion: 0.32, 0.14, 1.0, 0.06. Mandibles formula: L 1234 0 a(m
<superScript id="F2B49BBAFFD5B435FAC4F9F3D03BF9AB" attach="left" box="[1351,1396,1635,1649]" fontSize="6" pageId="6" pageNumber="99">1,2,3,4</superScript>
)b / R 1234 y a(m
<superScript id="F2B49BBAFFD5B435FEB0F916D423F94E" attach="right" box="[307,364,1670,1684]" fontSize="6" pageId="6" pageNumber="99">1,2,3,4,5</superScript>
)b, tooth a&gt;b in both mandibles. Labium: prementum slightly wider than long; ligula W/L ratio 1:0.16, with a ventral row of 910 dark reddish-brown, short, truncate teeth on middle (
<figureCitation id="9DFA2A77FFD5B435FBEEF93BD1F0F91F" box="[1133,1215,1707,1733]" captionStart="FIGURE 7" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[205,1383,181,1919]" captionTargetId="figure-23@8.[205,1383,181,1919]" captionTargetPageId="8" captionText="FIGURE 7. Epigomphus brillantina: a) antennae, dorsal view; b) ventral view of mouth parts showing the hypopharynx; c) detail of the ligula and palps, ventral view (right movable hook broken); d) left hind leg; e) sterna 510; f) sterna 89 showing the female gonapophyses; g) S10 and caudal appendages, dorsal." pageId="6" pageNumber="99">Fig. 7c</figureCitation>
). Labial palp apical lobe with tip finely serrate, internal margin slightly concave with 1012 small truncate teeth (
<figureCitation id="9DFA2A77FFD5B435FB2EF95FD1B0F933" box="[1197,1279,1743,1769]" captionStart="FIGURE 7" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[205,1383,181,1919]" captionTargetId="figure-23@8.[205,1383,181,1919]" captionTargetPageId="8" captionText="FIGURE 7. Epigomphus brillantina: a) antennae, dorsal view; b) ventral view of mouth parts showing the hypopharynx; c) detail of the ligula and palps, ventral view (right movable hook broken); d) left hind leg; e) sterna 510; f) sterna 89 showing the female gonapophyses; g) S10 and caudal appendages, dorsal." pageId="6" pageNumber="99">Fig. 7c</figureCitation>
).
</paragraph>
<paragraph id="057E36F2FFD5B435FF44F963D5B6F88F" blockId="6.[151,1437,1382,1877]" pageId="6" pageNumber="99">
<emphasis id="37B5EAE0FFD5B435FF44F963D457F8D7" box="[199,280,1779,1805]" italics="true" pageId="6" pageNumber="99">Thorax</emphasis>
: As described for
<taxonomicName id="C2C14D71FFD5B435FE64F964D70EF8D7" authorityName="Bota-Sierra &amp; Novelo-Gutierrez" authorityYear="2020" box="[487,577,1779,1805]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="99" phylum="Arthropoda" rank="species" species="rufus">
<emphasis id="37B5EAE0FFD5B435FE64F964D70EF8D7" box="[487,577,1779,1805]" italics="true" pageId="6" pageNumber="99">E. rufus</emphasis>
</taxonomicName>
but: Femora (
<figureCitation id="9DFA2A77FFD5B435FD63F963D67DF8D7" box="[736,818,1779,1805]" captionStart="FIGURE 7" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[205,1383,181,1919]" captionTargetId="figure-23@8.[205,1383,181,1919]" captionTargetPageId="8" captionText="FIGURE 7. Epigomphus brillantina: a) antennae, dorsal view; b) ventral view of mouth parts showing the hypopharynx; c) detail of the ligula and palps, ventral view (right movable hook broken); d) left hind leg; e) sterna 510; f) sterna 89 showing the female gonapophyses; g) S10 and caudal appendages, dorsal." pageId="6" pageNumber="99">Fig. 7d</figureCitation>
) reddish-brown on basal 0.80, a yellow band on apical 0.20, tibiae reddish-yellow, tarsal claws yellow (
<figureCitation id="9DFA2A77FFD5B435FD38F887D640F8EB" box="[699,783,1815,1841]" captionStart="FIGURE 7" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[205,1383,181,1919]" captionTargetId="figure-23@8.[205,1383,181,1919]" captionTargetPageId="8" captionText="FIGURE 7. Epigomphus brillantina: a) antennae, dorsal view; b) ventral view of mouth parts showing the hypopharynx; c) detail of the ligula and palps, ventral view (right movable hook broken); d) left hind leg; e) sterna 510; f) sterna 89 showing the female gonapophyses; g) S10 and caudal appendages, dorsal." pageId="6" pageNumber="99">Fig. 7d</figureCitation>
). Wing sheaths light yellow, with abundant, long setae on margins.
</paragraph>
<caption id="51BE667AFFD4B434FF14FB64D108FAD6" box="[151,1095,1268,1295]" pageId="7" pageNumber="100" startId="7.[151,250,1268,1292]" targetBox="[151,1436,181,1244]" targetPageId="7" targetType="figure">
<paragraph id="057E36F2FFD4B434FF14FB64D108FAD6" blockId="7.[151,1095,1268,1295]" box="[151,1095,1268,1295]" pageId="7" pageNumber="100">
<emphasis id="37B5EAE0FFD4B434FF14FB64D45CFAD6" bold="true" box="[151,275,1268,1292]" pageId="7" pageNumber="100">FIGURE 6.</emphasis>
<taxonomicName id="C2C14D71FFD4B434FE9AFB64D745FAD6" authorityName="Bota-Sierra &amp; Novelo-Gutierrez" authorityYear="2020" box="[281,522,1268,1292]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="100" phylum="Arthropoda" rank="species" species="brillantina">
<emphasis id="37B5EAE0FFD4B434FE9AFB64D745FAD6" box="[281,522,1268,1292]" italics="true" pageId="7" pageNumber="100">Epigomphus brillantina</emphasis>
</taxonomicName>
: a) F-1 female larva, dorsal view; b)
<emphasis id="37B5EAE0FFD4B434FC06FB66D6F7FAD5" box="[901,952,1270,1295]" italics="true" pageId="7" pageNumber="100">Ídem</emphasis>
, ventral view.
</paragraph>
</caption>
<paragraph id="057E36F2FFD4B434FF44FACFD69DF93B" blockId="7.[151,1437,1375,1869]" pageId="7" pageNumber="100">
<emphasis id="37B5EAE0FFD4B434FF44FACFD47EFAA3" box="[199,305,1375,1401]" italics="true" pageId="7" pageNumber="100">Abdomen</emphasis>
: As described for
<taxonomicName id="C2C14D71FFD4B434FD81FACFD713FAA3" authorityName="Bota-Sierra &amp; Novelo-Gutierrez" authorityYear="2020" box="[514,604,1375,1401]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="100" phylum="Arthropoda" rank="species" species="rufus">
<emphasis id="37B5EAE0FFD4B434FD81FACFD713FAA3" box="[514,604,1375,1401]" italics="true" pageId="7" pageNumber="100">E. rufus</emphasis>
</taxonomicName>
except: Mostly brown with a complex pattern of yellow spots on tergum, S910 dark reddish-brown (
<figureCitation id="9DFA2A77FFD4B434FE4DFA13D750FA46" box="[462,543,1410,1437]" captionStart="FIGURE 6" captionStartId="7.[151,250,1268,1292]" captionTargetBox="[151,1436,181,1244]" captionTargetId="figure-17@7.[151,1436,181,1244]" captionTargetPageId="7" captionText="FIGURE 6. Epigomphus brillantina: a) F-1 female larva, dorsal view; b) Ídem, ventral view." pageId="7" pageNumber="100">Fig. 6a</figureCitation>
); lateral margins of S79 serrate, serrations increasing in number and robustness caudad (
<figureCitation id="9DFA2A77FFD4B434FF7AFA37D401FA1B" box="[249,334,1447,1473]" captionStart="FIGURE 7" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[205,1383,181,1919]" captionTargetId="figure-23@8.[205,1383,181,1919]" captionTargetPageId="8" captionText="FIGURE 7. Epigomphus brillantina: a) antennae, dorsal view; b) ventral view of mouth parts showing the hypopharynx; c) detail of the ligula and palps, ventral view (right movable hook broken); d) left hind leg; e) sterna 510; f) sterna 89 showing the female gonapophyses; g) S10 and caudal appendages, dorsal." pageId="7" pageNumber="100">Fig. 7e</figureCitation>
); tergites 23 with long, bristle-like setae, tergites 49 mostly bare, with some scattered, long, delicate setae; posterior margins of S29 with a dense row of minute, whitish, scalelike setae, S10 glabrous. Sterna 17 mostly yellowish-brown (
<figureCitation id="9DFA2A77FFD4B434FE6BFA7FD774F9D2" box="[488,571,1518,1545]" captionStart="FIGURE 6" captionStartId="7.[151,250,1268,1292]" captionTargetBox="[151,1436,181,1244]" captionTargetId="figure-17@7.[151,1436,181,1244]" captionTargetPageId="7" captionText="FIGURE 6. Epigomphus brillantina: a) F-1 female larva, dorsal view; b) Ídem, ventral view." pageId="7" pageNumber="100">Fig. 6b</figureCitation>
), with posterior margin of middle sternites dark reddish-brown (
<figureCitation id="9DFA2A77FFD4B434FA8AFA7FD029F9D2" box="[1289,1382,1518,1545]" captionStart="FIGURE 6" captionStartId="7.[151,250,1268,1292]" captionTargetBox="[151,1436,181,1244]" captionTargetId="figure-17@7.[151,1436,181,1244]" captionTargetPageId="7" captionText="FIGURE 6. Epigomphus brillantina: a) F-1 female larva, dorsal view; b) Ídem, ventral view." pageId="7" pageNumber="100">Figs. 6b</figureCitation>
,
<figureCitation id="9DFA2A77FFD4B434FAF1FA7FD0C0F9D3" box="[1394,1423,1519,1545]" captionStart="FIGURE 7" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[205,1383,181,1919]" captionTargetId="figure-23@8.[205,1383,181,1919]" captionTargetPageId="8" captionText="FIGURE 7. Epigomphus brillantina: a) antennae, dorsal view; b) ventral view of mouth parts showing the hypopharynx; c) detail of the ligula and palps, ventral view (right movable hook broken); d) left hind leg; e) sterna 510; f) sterna 89 showing the female gonapophyses; g) S10 and caudal appendages, dorsal." pageId="7" pageNumber="100">7e</figureCitation>
), sternum 8 light reddish-yellow, sternum 9 reddish-brown, sternum 10 dark reddish-brown (
<figureCitation id="9DFA2A77FFD4B434FB3BF983D043F9F7" box="[1208,1292,1555,1581]" captionStart="FIGURE 7" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[205,1383,181,1919]" captionTargetId="figure-23@8.[205,1383,181,1919]" captionTargetPageId="8" captionText="FIGURE 7. Epigomphus brillantina: a) antennae, dorsal view; b) ventral view of mouth parts showing the hypopharynx; c) detail of the ligula and palps, ventral view (right movable hook broken); d) left hind leg; e) sterna 510; f) sterna 89 showing the female gonapophyses; g) S10 and caudal appendages, dorsal." pageId="7" pageNumber="100">Fig. 7e</figureCitation>
); sterna 28 with minute spiniform setae on middle sternites only visible at great magnification, posterior margins bare; middle sternite on sternum 9 with small, sparse, delicate setae, posterior margin of sternum 9 with a row of abundant, minute, white scale-like setae. Female gonapophyses vestigial (
<figureCitation id="9DFA2A77FFD4B434FCE3F9EFD6E0F943" box="[864,943,1663,1689]" captionStart="FIGURE 7" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[205,1383,181,1919]" captionTargetId="figure-23@8.[205,1383,181,1919]" captionTargetPageId="8" captionText="FIGURE 7. Epigomphus brillantina: a) antennae, dorsal view; b) ventral view of mouth parts showing the hypopharynx; c) detail of the ligula and palps, ventral view (right movable hook broken); d) left hind leg; e) sterna 510; f) sterna 89 showing the female gonapophyses; g) S10 and caudal appendages, dorsal." pageId="7" pageNumber="100">Fig. 7f</figureCitation>
), roundly-pointed. Caudal appendages (
<figureCitation id="9DFA2A77FFD4B434FAECF9EFD5F8F967" captionStart="FIGURE 7" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[205,1383,181,1919]" captionTargetId="figure-23@8.[205,1383,181,1919]" captionTargetPageId="8" captionText="FIGURE 7. Epigomphus brillantina: a) antennae, dorsal view; b) ventral view of mouth parts showing the hypopharynx; c) detail of the ligula and palps, ventral view (right movable hook broken); d) left hind leg; e) sterna 510; f) sterna 89 showing the female gonapophyses; g) S10 and caudal appendages, dorsal." pageId="7" pageNumber="100">
Fig.
<quantity id="C2399B17FFD4B434FF14F933D5F8F967" box="[151,183,1699,1725]" metricMagnitude="-3" metricUnit="kg" metricValue="7.0" pageId="7" pageNumber="100" unit="g" value="7.0">7g</quantity>
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): in dorsal view, tips of cerci divergent, tips of paraprocts more or less straight, in lateral view tips of epiproct and cerci straight; cerci and paraprocts subequal, epiproct slightly shorter.
</paragraph>
<paragraph id="057E36F2FFD4B434FF44F97BD1C2F8F3" blockId="7.[151,1437,1375,1869]" pageId="7" pageNumber="100">
<emphasis id="37B5EAE0FFD4B434FF44F97BD4A4F8DF" bold="true" box="[199,491,1770,1797]" pageId="7" pageNumber="100">Measurements [in mm]:</emphasis>
F-
<specimenCount id="13C7FD7BFFD4B434FD91F97BD72EF8DF" box="[530,609,1771,1797]" count="1" pageId="7" pageNumber="100" type="larva">1 larva</specimenCount>
: TL 25.5; AL (ventral, excl. caudal app.) 14; MWh 5.6; HfL (lateral) 5; MWa (ventral) 8; Ep 1.2, Ce 1.3, Pp 1.3; spine on
<geoCoordinate id="60F55035FFD4B434FD49F89FD65CF8F3" box="[714,787,1807,1833]" degrees="7" direction="south" minutes="0. 4" orientation="latitude" pageId="7" pageNumber="100" precision="92" value="-7.0066667">S7 0.4</geoCoordinate>
, spine on
<geoCoordinate id="60F55035FFD4B434FC07F89FD681F8F2" box="[900,974,1806,1833]" degrees="8" direction="south" minutes="0. 6" orientation="latitude" pageId="7" pageNumber="100" precision="92" value="-8.01">S8 0.6</geoCoordinate>
, spine on
<geoCoordinate id="60F55035FFD4B434FBBCF89FD1C6F8F3" box="[1087,1161,1807,1833]" degrees="9" direction="south" minutes="0. 8" orientation="latitude" pageId="7" pageNumber="100" precision="92" value="-9.013333">S9 0.8</geoCoordinate>
.
</paragraph>
<paragraph id="057E36F2FFD4B434FF44F8A2D026F897" blockId="7.[151,1437,1375,1869]" box="[199,1385,1842,1869]" pageId="7" pageNumber="100">
<emphasis id="37B5EAE0FFD4B434FF44F8A2D465F896" bold="true" box="[199,298,1842,1868]" pageId="7" pageNumber="100">Habitat.</emphasis>
Larvae were found in a small, still, shallow stream with abundant leaf litter in the forest (
<figureCitation id="9DFA2A77FFD4B434FA95F8A3D014F897" box="[1302,1371,1843,1869]" captionStart="FIGURE 8" captionStartId="9.[151,250,1168,1192]" captionTargetBox="[151,1435,181,1145]" captionTargetId="figure-17@9.[151,1435,181,1145]" captionTargetPageId="9" captionText="FIGURE 8. Epigomphus brillantina larval habitat at Finca La Brillantina." pageId="7" pageNumber="100">Fig. 8</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,128 @@
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<paragraph id="182436E2FF83FFD2C5AAF971FDE913A2" blockId="10.[151,854,1667,2018]" box="[151,553,1739,1766]" pageId="10" pageNumber="11">
<heading id="436C818EFF83FFD2C5AAF971FDE913A2" bold="true" box="[151,553,1739,1766]" fontSize="11" level="2" pageId="10" pageNumber="11" reason="4">
<taxonomicName id="DF9B4D61FF83FFD2C5AAF971FDE913A2" ID-CoL="6SGTN" authority="Emery, 1892" authorityName="Emery" authorityYear="1892" box="[151,553,1739,1766]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="bauri">
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<emphasis id="2AEFEAF0FF83FFD2C5AAF971FE4C13A2" bold="true" box="[151,396,1739,1766]" italics="true" pageId="10" pageNumber="11">Odontomachus bauri</emphasis>
<bibRefCitation id="7C0A4B13FF83FFD2C4AEF976FDE913A2" author="Emery, C." box="[403,553,1740,1766]" pageId="10" pageNumber="11" pagination="553 - 574" refId="ref93113" refString="Emery, C. (1892) Voyage de M. Ch. Alluaud dans le territoire d'Assinie (Afrique occidentale) en juillet et aout 1886. Formicides. Annales de La Societe Entomologique de France, 60, 553 - 574." type="journal article" year="1892">Emery, 1892</bibRefCitation>
</emphasis>
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=
<taxonomicName id="DF9B4D61FF83FFD2C5E3F94AFD52124E" authority="Roger, 1861" authorityName="Roger" authorityYear="1861" box="[222,658,1776,1803]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="pubescens">
<emphasis id="2AEFEAF0FF83FFD2C5E3F94AFDC2124E" box="[222,514,1776,1802]" italics="true" pageId="10" pageNumber="11">Odontomachus pubescens</emphasis>
<bibRefCitation id="7C0A4B13FF83FFD2C734F94BFD52124E" author="Roger, J." box="[521,658,1776,1803]" pageId="10" pageNumber="11" pagination="278 - 312" refId="ref97876" refString="Roger, J. (1861) die Ponera-artigen Ameisen. Berliner Entomologische Zeitschrift, 4, 278 - 312." type="journal article" year="1861">Roger, 1861</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="182436E2FF83FFD2C5FAF8AFFC96126A" blockId="10.[151,854,1667,2018]" box="[199,854,1812,1839]" pageId="10" pageNumber="11">
=
<taxonomicName id="DF9B4D61FF83FFD2C5E3F8AEFC96126A" authority="Emery, 1890" authorityName="Emery" authorityYear="1890" box="[222,854,1812,1839]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="subSpecies" species="haematodus" subSpecies="microcephalus">
<emphasis id="2AEFEAF0FF83FFD2C5E3F8AEFD01126A" box="[222,705,1812,1838]" italics="true" pageId="10" pageNumber="11">Odontomachus haematodus microcephalus</emphasis>
<bibRefCitation id="7C0A4B13FF83FFD2C7F5F8AFFC96126A" author="Emery, C." box="[712,854,1812,1839]" pageId="10" pageNumber="11" pagination="38 - 80" refId="ref93088" refString="Emery, C. (1890) Studi sulle formiche della fauna neotropical. Bullettino della Societa Entomologica Italiana, 22, 38 - 80" type="journal article" year="1890">Emery, 1890</bibRefCitation>
</taxonomicName>
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<paragraph id="182436E2FF83FFD2C5FAF883FCE41216" blockId="10.[151,854,1667,2018]" box="[199,804,1848,1875]" pageId="10" pageNumber="11">
=
<taxonomicName id="DF9B4D61FF83FFD2C5E3F882FCE41216" authority="Emery, 1893" authorityName="Emery" authorityYear="1893" box="[222,804,1848,1875]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="subSpecies" species="haematodus" subSpecies="paucidens">
<emphasis id="2AEFEAF0FF83FFD2C5E3F882FD4F1216" box="[222,655,1848,1874]" italics="true" pageId="10" pageNumber="11">Odontomachus haematodus paucidens</emphasis>
<bibRefCitation id="7C0A4B13FF83FFD2C7A8F883FCE41216" author="Emery, C." box="[661,804,1848,1875]" pageId="10" pageNumber="11" pagination="89 - 92" refId="ref93157" refString="Emery, C. (1893) Notice sur quelques fourmis des iles Galapagos. Annales de La Societe Entomologique de France, 62, 89 - 92." type="journal article" year="1893">Emery, 1893</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="182436E2FF83FFD2C5AAF8E7FD461233" blockId="10.[151,854,1667,2018]" box="[151,646,1884,1911]" pageId="10" pageNumber="11">
=
<taxonomicName id="DF9B4D61FF83FFD2C593F8E6FD461233" authority="Provancher, 1895" authorityName="Provancher" authorityYear="1895" box="[174,646,1884,1911]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="sericeus">
<emphasis id="2AEFEAF0FF83FFD2C593F8E6FE7B1232" box="[174,443,1884,1910]" italics="true" pageId="10" pageNumber="11">Odontomachus sericeus</emphasis>
<bibRefCitation id="7C0A4B13FF83FFD2C4FCF8E6FD461233" author="Provancher, L." box="[449,646,1884,1911]" pageId="10" pageNumber="11" pagination="95 - 97" refId="ref97004" refString="Provancher, L. (1895) Les dernieres descriptions de l'Abbe Provancher. Naturaliste Canadien, Quebec, 22, 95 - 97." type="journal article" year="1895">Provancher, 1895</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="182436E2FF83FFD2C5FAF83BFCEB12DE" blockId="10.[151,854,1667,2018]" box="[199,811,1920,1947]" pageId="10" pageNumber="11">
=
<taxonomicName id="DF9B4D61FF83FFD2C5E3F83AFCEB12DE" authority="Forel, 1908" authorityName="Forel" authorityYear="1908" box="[222,811,1920,1946]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="subSpecies" species="haematodus" subSpecies="rugisquama">
<emphasis id="2AEFEAF0FF83FFD2C5E3F83AFD6312DE" box="[222,675,1920,1946]" italics="true" pageId="10" pageNumber="11">Odontomachus haematodus rugisquama</emphasis>
<bibRefCitation id="7C0A4B13FF83FFD2C797F83AFCEB12DE" author="Forel, A." box="[682,811,1920,1946]" pageId="10" pageNumber="11" pagination="35 - 72" refId="ref93901" refString="Forel, A. (1908) Fourmis de Costa-Rica recoltees par M. Paul Biolley. Bulletin de la Societe Vaudoise des Sciences Naturelles, 44, 35 - 72." type="journal article" year="1908">Forel, 1908</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="182436E2FF83FFD2C5FAF81FFD3E12FA" blockId="10.[151,854,1667,2018]" box="[199,766,1956,1983]" pageId="10" pageNumber="11">
=
<taxonomicName id="DF9B4D61FF83FFD2C5E3F81EFD3E12FA" authority="Mann, 1920" authorityName="Mann" authorityYear="1920" box="[222,766,1956,1983]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="subSpecies" species="haematodus" subSpecies="notatus">
<emphasis id="2AEFEAF0FF83FFD2C5E3F81EFDAF12FA" box="[222,623,1956,1982]" italics="true" pageId="10" pageNumber="11">Odontomachus haematodus notatus</emphasis>
<bibRefCitation id="7C0A4B13FF83FFD2C74BF81FFD3E12FA" author="Mann, W. M." box="[630,766,1956,1983]" pageId="10" pageNumber="11" pagination="403 - 439" refId="ref96023" refString="Mann, W. M. (1920) Additions to the ant fauna of the West Indies and Central America. Bulletin of the American Museum of Natural History, 42, 403 - 439." type="journal article" year="1920">Mann, 1920</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="182436E2FF83FFD2C5AAF87DFDEE12A6" blockId="10.[151,854,1667,2018]" box="[151,558,1991,2018]" pageId="10" pageNumber="11">
<taxonomicName id="DF9B4D61FF83FFD2C5AAF87DFDEE12A6" ID-CoL="48QBB" authority="Forel, 1908" authorityName="Forel" authorityYear="1908" box="[151,558,1991,2018]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="biolleyi">
<emphasis id="2AEFEAF0FF83FFD2C5AAF87DFDEE12A6" bold="true" box="[151,558,1991,2018]" pageId="10" pageNumber="11">
<emphasis id="2AEFEAF0FF83FFD2C5AAF87DFE6212A6" bold="true" box="[151,418,1991,2018]" italics="true" pageId="10" pageNumber="11">Odontomachus biolleyi</emphasis>
<bibRefCitation id="7C0A4B13FF83FFD2C494F872FDEE12A6" author="Forel, A." box="[425,558,1992,2018]" pageId="10" pageNumber="11" pagination="35 - 72" refId="ref93901" refString="Forel, A. (1908) Fourmis de Costa-Rica recoltees par M. Paul Biolley. Bulletin de la Societe Vaudoise des Sciences Naturelles, 44, 35 - 72." type="journal article" year="1908">Forel, 1908</bibRefCitation>
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View file

@ -0,0 +1,202 @@
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<mods:title id="1D55F6923E1C3E22AFAE0F822E6C583D">New species of little-known crickets from the subfamily Pteroplistinae (Orthoptera, Grylloidea) from Sabah and contribution to their bioacoustics</mods:title>
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<mods:affiliation id="C6D2ED33C72C49D23320338FB4B7C99F">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore &amp; Institut de Systématique, Evolution, Biodiversité (ISYEB), Muséum national dHistoire naturelle, CNRS, SU, EPHE-PSL, UA, 57 rue Cuvier, CP 50, 75231 Paris Cedex 05, France &amp; Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore</mods:affiliation>
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<emphasis id="6B874948FFCD3E110C95F964FE7D009F" bold="true" box="[151,397,1690,1717]" italics="true" pageId="18" pageNumber="223">Pteroplistes lagrecai</emphasis>
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(
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,
<figureCitation id="C1C889DFFFCD3E110D04F940FED900F2" box="[262,297,1726,1752]" captionStart="FIGURE 3" captionStartId="6.[151,250,1766,1790]" captionTargetBox="[162,1425,320,1732]" captionTargetId="figure-71@6.[151,1436,309,1743]" captionTargetPageId="6" captionText="FIGURE 3. Oscillograms of the calling songs of the pteroplistine cricket species: three echemes with different lengths of Tembelingiola biaculeata sp. nov. (A), two echeme-sequencesof Tembelingiola kabili sp. nov. (B), continuous trill of Pteroplistes silam sp. nov. (C), an echeme-sequence of Pteroplistes lagrecai Gorochov, 2004 (D) and three echemes of Singapuriola separata Gorochov &amp; Tan, 2012 (E). Scale bar: 1 s." pageId="18" pageNumber="223">3B</figureCitation>
,
<figureCitation id="C1C889DFFFCD3E110D37F940FEA800F2" box="[309,344,1726,1752]" captionStart="FIGURE 4" captionStartId="7.[151,250,1500,1524]" captionTargetBox="[162,1425,192,1466]" captionTargetId="figure-17@7.[151,1436,181,1477]" captionTargetPageId="7" captionText="FIGURE 4. Oscillograms of the syllables of the pteroplistine cricket species: Tembelingiola biaculeata sp. nov. (A), Tembelingiola kabili sp. nov. (B), Pteroplistes silam sp. nov. (C), Pteroplistes lagrecai Gorochov, 2004 (D) and Singapuriola separata Gorochov &amp; Tan, 2012 (E). Scale bar: 0.05 s." pageId="18" pageNumber="223">4B</figureCitation>
,
<figureCitation id="C1C889DFFFCD3E110D66F940FE6300F3" box="[356,403,1726,1753]" captionStart="FIGURE 15" captionStartId="19.[151,250,1771,1795]" captionTargetBox="[151,1436,181,1747]" captionTargetId="figure-17@19.[151,1436,181,1747]" captionTargetPageId="19" captionText="FIGURE 15. Pteroplistes silam sp. nov. calling on the tree trunk in Mount Silam (A), and Pteroplistes lagrecai Gorochov, 2004 calling on the tree in Sepilok (B)." pageId="18" pageNumber="223">15B</figureCitation>
)
</paragraph>
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<emphasis id="6B874948FFCD3E110C95F8FDFE990131" box="[151,361,1795,1819]" italics="true" pageId="18" pageNumber="223">Pteroplistes lagrecai</emphasis>
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<bibRefCitation id="3D62E8ABFFCD3E110E5EF8FDFCC30131" author="Gorochov, A. V." box="[604,819,1795,1819]" pageId="18" pageNumber="223" pagination="445 - 456" refId="ref7996" refString="Gorochov, A. V. (2022) New taxa of the cricket subfamily Pteroplistinae (Orthoptera: Gryllidae) from Indo Malayan Region. Amurian Zoological Journal, 14 (3), 445 - 456. https: // doi. org / 10.33910 / 2686 - 9519 - 2022 - 14 - 3 - 445 - 456" type="journal article" year="2022">Gorochov, 2022: 447</bibRefCitation>
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<emphasis id="6B874948FFCD3E110C95F8B7FE750149" bold="true" box="[151,389,1865,1891]" pageId="18" pageNumber="223">Material examined.</emphasis>
<materialsCitation id="E99B9F07FFCD3E110D93F8B7FC510181" collectingDate="2022-05-18" collectorName="T. Robillard &amp; M. K. Tan" country="Malaysia" county="Sandakan District" elevation="54" latitude="5.87395" location="Rainforest Discovery Centre" longLatPrecision="1" longitude="117.93871" municipality="Sepilok" pageId="18" pageNumber="223" specimenCount="1" specimenCount-male="1" stateProvince="Sabah">
EAST
<collectingCountry id="21E4D5CAFFCD3E110DE0F8B7FD830149" box="[482,627,1865,1891]" name="Malaysia" pageId="18" pageNumber="223">MALAYSIA</collectingCountry>
: •
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;
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,
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,
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,
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, dipterocarp forest;
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,
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,
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;
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<collectingDate id="3D094A72FFCD3E110F3BF893FC5901AD" box="[825,937,1901,1927]" pageId="18" pageNumber="223" value="2022-05-18">18.v.2022</collectingDate>
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, night (19h); on tree trunk;
<collectorName id="F406F08CFFCD3E1108F0F890FA8C01AD" box="[1266,1404,1901,1928]" pageId="18" pageNumber="223">T. Robillard</collectorName>
&amp;
<collectorName id="F406F08CFFCD3E110C95F86CFEF30186" box="[151,259,1938,1964]" pageId="18" pageNumber="223">M.K. Tan</collectorName>
leg.; TR22-27, call recording video 0062; MNHN-EO1288
</materialsCitation>
.
</paragraph>
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<subSubSection id="11E9C6D1FFCD3E110CC5F84BFAAB01E5" box="[199,1371,1973,2000]" pageId="18" pageNumber="223" type="biology_ecology">
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<emphasis id="6B874948FFCD3E110CC5F84BFEDC01E5" bold="true" box="[199,300,1973,1999]" pageId="18" pageNumber="223">Ecology.</emphasis>
This species was found calling on a tree trunk at night, where
<specimenCount id="4FF55ED3FFCD3E110FEBF848FB8101E5" box="[1001,1137,1973,2000]" count="2" pageId="18" pageNumber="223" type="female">two females</specimenCount>
were located near it.
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFCD3E170CC5F827FB31074F" lastPageId="20" lastPageNumber="225" pageId="18" pageNumber="223" type="distribution">
<paragraph id="594C955AFFCD3E110CC5F827FCF401D9" blockId="18.[151,1436,1865,2035]" box="[199,772,2009,2035]" pageId="18" pageNumber="223">
<emphasis id="6B874948FFCD3E110CC5F827FE9101D9" bold="true" box="[199,353,2009,2035]" pageId="18" pageNumber="223">Distribution.</emphasis>
EAST
<collectingCountry id="21E4D5CAFFCD3E110DB6F827FDBA01D9" box="[436,586,2009,2035]" name="Malaysia" pageId="18" pageNumber="223">MALAYSIA</collectingCountry>
:
<collectingRegion id="9B375BB8FFCD3E110E56F827FD6D01D9" box="[596,669,2009,2035]" country="Malaysia" name="Sabah" pageId="18" pageNumber="223">Sabah</collectingRegion>
: Sepilok.
</paragraph>
<caption id="0D8CC5D2FFCC3E100C95F915FE1A010D" pageId="19" pageNumber="224" startId="19.[151,250,1771,1795]" targetBox="[151,1436,181,1747]" targetPageId="19" targetType="figure">
<paragraph id="594C955AFFCC3E100C95F915FE1A010D" blockId="19.[151,1437,1771,1831]" pageId="19" pageNumber="224">
<emphasis id="6B874948FFCC3E100C95F915FEEF0129" bold="true" box="[151,287,1771,1795]" pageId="19" pageNumber="224">FIGURE 15.</emphasis>
<taxonomicName id="9EF3EED9FFCC3E100D26F915FE290129" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" box="[292,473,1771,1795]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="19" pageNumber="224" phylum="Arthropoda" rank="species" species="silam" status="sp. nov.">
<emphasis id="6B874948FFCC3E100D26F915FE290129" box="[292,473,1771,1795]" italics="true" pageId="19" pageNumber="224">Pteroplistes silam</emphasis>
</taxonomicName>
<emphasis id="6B874948FFCC3E100DDCF915FDDE0129" bold="true" box="[478,558,1771,1795]" pageId="19" pageNumber="224">
<taxonomicNameLabel id="70B4F433FFCC3E100DDCF915FDDE0129" box="[478,558,1771,1795]" pageId="19" pageNumber="224" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
calling on the tree trunk in Mount Silam (A), and
<taxonomicName id="9EF3EED9FFCC3E100826F915FA6D0129" authority="Gorochov, 2004" authorityName="Gorochov" authorityYear="2004" box="[1060,1437,1771,1795]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="19" pageNumber="224" phylum="Arthropoda" rank="species" species="lagrecai">
<emphasis id="6B874948FFCC3E100826F915FB040129" box="[1060,1268,1771,1795]" italics="true" pageId="19" pageNumber="224">Pteroplistes lagrecai</emphasis>
<bibRefCitation id="3D62E8ABFFCC3E1008F8F915FA6D0129" author="Gorochov, A. V." box="[1274,1437,1771,1795]" pageId="19" pageNumber="224" pagination="379 - 396" refId="ref7847" refString="Gorochov, A. V. (2004) Review of the subfamily Pteroplistinae (Orthoptera Gryllidae). Memorie della Societa Entomologica Italiana, 82 (2), 379 - 396." type="journal article" year="2004">Gorochov, 2004</bibRefCitation>
</taxonomicName>
calling on the tree in Sepilok (B).
</paragraph>
</caption>
<paragraph id="594C955AFFCC3E170CC5F887FB31074F" blockId="19.[151,1437,1913,2012]" lastBlockId="20.[151,1437,151,357]" lastPageId="20" lastPageNumber="225" pageId="19" pageNumber="224">
<emphasis id="6B874948FFCC3E100CC5F887FDB101B9" bold="true" box="[199,577,1913,1939]" pageId="19" pageNumber="224">
Calling song (
<figureCitation id="C1C889DFFFCC3E100D6CF887FE3E01B9" box="[366,462,1913,1939]" captionStart="FIGURE 2" captionStartId="5.[151,250,1600,1624]" captionTargetBox="[161,1426,191,1567]" captionTargetId="figure-17@5.[151,1436,181,1577]" captionTargetPageId="5" captionText="FIGURE 2. Power spectra of the calling songs of the pteroplistine cricket species: Tembelingiola biaculeata sp. nov. (A), Tembelingiola kabili sp. nov. (B), Pteroplistes silam sp. nov. (C), Pteroplistes lagrecai Gorochov, 2004 (D) and Singapuriola separata Gorochov &amp; Tan, 2012 (E). The grey line represents the power [dB] (axis name and scale in grey) and the black solid peak represents the relative amplitude (axis name in black)." pageId="19" pageNumber="224">Figs 2D</figureCitation>
,
<figureCitation id="C1C889DFFFCC3E100DDFF887FDF001B9" box="[477,512,1913,1939]" captionStart="FIGURE 3" captionStartId="6.[151,250,1766,1790]" captionTargetBox="[162,1425,320,1732]" captionTargetId="figure-71@6.[151,1436,309,1743]" captionTargetPageId="6" captionText="FIGURE 3. Oscillograms of the calling songs of the pteroplistine cricket species: three echemes with different lengths of Tembelingiola biaculeata sp. nov. (A), two echeme-sequencesof Tembelingiola kabili sp. nov. (B), continuous trill of Pteroplistes silam sp. nov. (C), an echeme-sequence of Pteroplistes lagrecai Gorochov, 2004 (D) and three echemes of Singapuriola separata Gorochov &amp; Tan, 2012 (E). Scale bar: 1 s." pageId="19" pageNumber="224">3D</figureCitation>
,
<figureCitation id="C1C889DFFFCC3E100E0DF887FDC401B9" box="[527,564,1913,1939]" captionStart="FIGURE 4" captionStartId="7.[151,250,1500,1524]" captionTargetBox="[162,1425,192,1466]" captionTargetId="figure-17@7.[151,1436,181,1477]" captionTargetPageId="7" captionText="FIGURE 4. Oscillograms of the syllables of the pteroplistine cricket species: Tembelingiola biaculeata sp. nov. (A), Tembelingiola kabili sp. nov. (B), Pteroplistes silam sp. nov. (C), Pteroplistes lagrecai Gorochov, 2004 (D) and Singapuriola separata Gorochov &amp; Tan, 2012 (E). Scale bar: 0.05 s." pageId="19" pageNumber="224">4D</figureCitation>
).
</emphasis>
Consists of an echeme-sequence. At 26°C, each echeme-sequence consists two parts, the first being 6±2 (510) isolated syllables, followed by the second being 11±4 (516) echemes made up of two syllables (doublet) each. For the first part, the average syllable duration is 22.1±1.5 ms (19.625.2 ms) and the average interval between consecutive syllables is 0.14±
<geoCoordinate id="3CC7F39DFFCB3E170F38FF69FC8F069B" box="[826,895,151,177]" degrees="0.03" direction="south" orientation="latitude" pageId="20" pageNumber="225" precision="555" value="-0.03">0.03 s</geoCoordinate>
(0.12
<geoCoordinate id="3CC7F39DFFCB3E170FCDFF69FBE6069B" box="[975,1046,151,177]" degrees="0.20" direction="south" orientation="latitude" pageId="20" pageNumber="225" precision="555" value="-0.2">0.20 s</geoCoordinate>
). For the second part, the average echeme duration is 58.8±1.8 ms (57.564.1 ms) and the average silent interval between consecutive echemes is the same as the average silent interval between consecutive syllables in the first part of the song. The average syllable duration of the doublet is 22.4±0.5 ms (21.523.2 ms), barely longer than that in the first part; and the average silent interval between the two syllables is 13.1±0.7 ms (11.814.1 ms). The frequency spectrum is pure-tonal and forms an harmonic series, with the energy peaking at a dominant fundamental frequency of 7.39 kHz.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,308 @@
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<mods:title id="051B08BDE815DFFEF6D9D7E4D30AB0A5">New species of little-known crickets from the subfamily Pteroplistinae (Orthoptera, Grylloidea) from Sabah and contribution to their bioacoustics</mods:title>
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<mods:affiliation id="95CBE5EA41EF5B27AF2817D87BC15264">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore &amp; Institut de Systématique, Evolution, Biodiversité (ISYEB), Muséum national dHistoire naturelle, CNRS, SU, EPHE-PSL, UA, 57 rue Cuvier, CP 50, 75231 Paris Cedex 05, France &amp; Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore</mods:affiliation>
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<heading id="02042236FFD13E0D0C95FB06FCC80339" bold="true" box="[151,824,1272,1299]" fontSize="11" level="1" pageId="14" pageNumber="219" reason="1">
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<taxonomicName id="9EF3EED9FFD13E0D0C95FB06FD3E0339" authority="Tan, Gorochov &amp; Robillard" authorityName="Tan, Gorochov &amp; Robillard" authorityYear="2024" box="[151,718,1272,1299]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="219" phylum="Arthropoda" rank="species" species="silam" status="sp. nov.">
<emphasis id="6B874948FFD13E0D0C95FB06FE9C0339" bold="true" box="[151,364,1272,1299]" italics="true" pageId="14" pageNumber="219">Pteroplistes silam</emphasis>
Tan, Gorochov &amp; Robillard
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,
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<subSubSection id="11E9C6D1FFD13E0D0C95FAE3FE010312" box="[151,497,1309,1336]" pageId="14" pageNumber="219" type="description">
<paragraph id="594C955AFFD13E0D0C95FAE3FE010312" blockId="14.[151,824,1272,1336]" box="[151,497,1309,1336]" pageId="14" pageNumber="219">
(
<figureCitation id="C1C889DFFFD13E0D0C9DFAE3FF0B031D" box="[159,251,1309,1335]" captionStart="FIGURE 2" captionStartId="5.[151,250,1600,1624]" captionTargetBox="[161,1426,191,1567]" captionTargetId="figure-17@5.[151,1436,181,1577]" captionTargetPageId="5" captionText="FIGURE 2. Power spectra of the calling songs of the pteroplistine cricket species: Tembelingiola biaculeata sp. nov. (A), Tembelingiola kabili sp. nov. (B), Pteroplistes silam sp. nov. (C), Pteroplistes lagrecai Gorochov, 2004 (D) and Singapuriola separata Gorochov &amp; Tan, 2012 (E). The grey line represents the power [dB] (axis name and scale in grey) and the black solid peak represents the relative amplitude (axis name in black)." pageId="14" pageNumber="219">Figs 2C</figureCitation>
,
<figureCitation id="C1C889DFFFD13E0D0D04FAE3FED9031D" box="[262,297,1309,1335]" captionStart="FIGURE 3" captionStartId="6.[151,250,1766,1790]" captionTargetBox="[162,1425,320,1732]" captionTargetId="figure-71@6.[151,1436,309,1743]" captionTargetPageId="6" captionText="FIGURE 3. Oscillograms of the calling songs of the pteroplistine cricket species: three echemes with different lengths of Tembelingiola biaculeata sp. nov. (A), two echeme-sequencesof Tembelingiola kabili sp. nov. (B), continuous trill of Pteroplistes silam sp. nov. (C), an echeme-sequence of Pteroplistes lagrecai Gorochov, 2004 (D) and three echemes of Singapuriola separata Gorochov &amp; Tan, 2012 (E). Scale bar: 1 s." pageId="14" pageNumber="219">3C</figureCitation>
,
<figureCitation id="C1C889DFFFD13E0D0D37FAE3FEA8031D" box="[309,344,1309,1335]" captionStart="FIGURE 4" captionStartId="7.[151,250,1500,1524]" captionTargetBox="[162,1425,192,1466]" captionTargetId="figure-17@7.[151,1436,181,1477]" captionTargetPageId="7" captionText="FIGURE 4. Oscillograms of the syllables of the pteroplistine cricket species: Tembelingiola biaculeata sp. nov. (A), Tembelingiola kabili sp. nov. (B), Pteroplistes silam sp. nov. (C), Pteroplistes lagrecai Gorochov, 2004 (D) and Singapuriola separata Gorochov &amp; Tan, 2012 (E). Scale bar: 0.05 s." pageId="14" pageNumber="219">4C</figureCitation>
,
<figureCitation id="C1C889DFFFD13E0D0D66FAE3FE72031D" box="[356,386,1309,1335]" captionStart="FIGURE 13" captionStartId="17.[151,250,1744,1768]" captionTargetBox="[152,1436,181,1720]" captionTargetId="figure-17@17.[151,1436,181,1720]" captionTargetPageId="17" captionText="FIGURE 13. Pteroplistes silam sp. nov.: head and pronotum in dorsal (A) and lateral (B) views, face in anterior view (C), left (D) and right (E) FWs in dorsal view, supra-anal plate in dorsal view (F). Scale bars: 5 mm (A, D, E); 2 mm (rest)." pageId="14" pageNumber="219">13</figureCitation>
,
<figureCitation id="C1C889DFFFD13E0D0D8CFAE3FE5C031D" box="[398,428,1309,1335]" captionStart="FIGURE 14" captionStartId="18.[151,250,1548,1572]" captionTargetBox="[166,1421,194,1500]" captionTargetId="figure-20@18.[151,1436,181,1524]" captionTargetPageId="18" captionText="FIGURE 14. Male genitalia of Pteroplistes silam sp. nov. in posterodorsal (A), dorsal (B), ventral (C) and lateral (D) views, before (A) and after (BD) cleaning with KOH. Scale bar: 1 mm." pageId="14" pageNumber="219">14</figureCitation>
,
<figureCitation id="C1C889DFFFD13E0D0DBAFAE3FE190312" box="[440,489,1309,1336]" captionStart="FIGURE 15" captionStartId="19.[151,250,1771,1795]" captionTargetBox="[151,1436,181,1747]" captionTargetId="figure-17@19.[151,1436,181,1747]" captionTargetPageId="19" captionText="FIGURE 15. Pteroplistes silam sp. nov. calling on the tree trunk in Mount Silam (A), and Pteroplistes lagrecai Gorochov, 2004 calling on the tree in Sepilok (B)." pageId="14" pageNumber="219">15A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFD13E0D0C95FA9BFE9A03ED" pageId="14" pageNumber="219" type="materials_examined">
<paragraph id="594C955AFFD13E0D0C95FA9BFE9A03ED" blockId="14.[151,1437,1381,1911]" pageId="14" pageNumber="219">
<emphasis id="6B874948FFD13E0D0C95FA9BFE710355" bold="true" box="[151,385,1381,1407]" pageId="14" pageNumber="219">Material examined.</emphasis>
<materialsCitation id="E99B9F07FFD13E0D0D8AFA9BFE9A03ED" collectingDate="2022-05-12" collectionCode="FRC" collectorName="M. K. Tan &amp; T. Robillard &amp; R. Japir" country="Malaysia" elevation="5" latitude="4.96878" location="Lahad Datu" longLatPrecision="1" longitude="118.17189" municipality="Mount Silam" pageId="14" pageNumber="219" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="holotype">
<emphasis id="6B874948FFD13E0D0D8AFA9BFE0E0355" bold="true" box="[392,510,1381,1407]" pageId="14" pageNumber="219">
<typeStatus id="86482BF8FFD13E0D0D8AFA9BFE090355" box="[392,505,1381,1407]" pageId="14" pageNumber="219" type="holotype">Holotype</typeStatus>
:
</emphasis>
EAST
<collectingCountry id="21E4D5CAFFD13E0D0E50FA9BFD180355" box="[594,744,1381,1407]" name="Malaysia" pageId="14" pageNumber="219">MALAYSIA</collectingCountry>
: •
<specimenCount id="4FF55ED3FFD13E0D0F06FA9BFCEC0355" box="[772,796,1381,1407]" count="1" pageId="14" pageNumber="219" type="male"></specimenCount>
;
<collectingRegion id="9B375BB8FFD13E0D0F2AFA9BFC5F0355" box="[808,943,1381,1407]" country="Malaysia" name="Sabah" pageId="14" pageNumber="219">Sabah State</collectingRegion>
,
<collectingMunicipality id="B9280F20FFD13E0D0FB8FA9BFBA00355" box="[954,1104,1381,1407]" pageId="14" pageNumber="219">Mount Silam</collectingMunicipality>
, near
<location id="5C2CC381FFD13E0D0891FA9BFAEA0355" LSID="urn:lsid:plazi:treatment:D15A244CFFD13E130C95FB06FC590193:5C2CC381FFD13E0D0891FA9BFAEA0355" box="[1171,1306,1381,1407]" country="Malaysia" latitude="4.96878" longLatPrecision="1" longitude="118.17189" municipality="Mount Silam" name="Lahad Datu" pageId="14" pageNumber="219" stateProvince="Sabah">Lahad Datu</location>
;
<geoCoordinate id="3CC7F39DFFD13E0D0924FA9BFA680355" box="[1318,1432,1381,1407]" degrees="4.96878" direction="north" orientation="latitude" pageId="14" pageNumber="219" precision="1" value="4.96878">N4.96878</geoCoordinate>
,
<geoCoordinate id="3CC7F39DFFD13E0D0C95FA77FED10389" box="[151,289,1417,1443]" degrees="118.17189" direction="east" orientation="longitude" pageId="14" pageNumber="219" precision="1" value="118.17189">E118.17189</geoCoordinate>
, 736.1±
<quantity id="9E0B38BFFFD13E0D0D7FFA77FE360389" box="[381,454,1417,1444]" metricMagnitude="0" metricUnit="m" metricValue="5.4" pageId="14" pageNumber="219" unit="m" value="5.4">
<elevation id="D2DE7269FFD13E0D0D7FFA77FE360389" box="[381,454,1417,1444]" metricMagnitude="0" metricUnit="m" metricValue="5.4" pageId="14" pageNumber="219" unit="m" value="5.4">5.4 m</elevation>
</quantity>
.a.s.l.; calling on tree trunk;
<date id="2D4DB39AFFD13E0D0F07FA77FC860389" box="[773,886,1417,1443]" pageId="14" pageNumber="219" value="2022-05-12">
<collectingDate id="3D094A72FFD13E0D0F07FA77FC860389" box="[773,886,1417,1443]" pageId="14" pageNumber="219" value="2022-05-12">12.v.2022</collectingDate>
</date>
, 21h56;
<collectorName id="F406F08CFFD13E0D0FD8FA77FBBA0389" box="[986,1098,1417,1443]" pageId="14" pageNumber="219">M.K. Tan</collectorName>
,
<collectorName id="F406F08CFFD13E0D0855FA77FB110389" box="[1111,1249,1417,1443]" pageId="14" pageNumber="219">T. Robillard</collectorName>
&amp;
<collectorName id="F406F08CFFD13E0D090BFA77FA920389" box="[1289,1378,1417,1443]" pageId="14" pageNumber="219">R. Japir</collectorName>
leg.; SBH.22.42 (
<collectionCode id="3FE20D9FFFD13E0D0D21FA53FEAE03ED" box="[291,350,1453,1479]" country="USA" httpUri="http://grbio.org/cool/z9c1-g2b0" name="Fusarium Research Center" pageId="14" pageNumber="219">FRC</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFD13E0D0CC5FA2FFD7D0179" pageId="14" pageNumber="219" type="diagnosis">
<paragraph id="594C955AFFD13E0D0CC5FA2FFD7D0179" blockId="14.[151,1437,1381,1911]" pageId="14" pageNumber="219">
<emphasis id="6B874948FFD13E0D0CC5FA2FFEB203C1" bold="true" box="[199,322,1489,1515]" pageId="14" pageNumber="219">Diagnosis.</emphasis>
This species is most similar to
<taxonomicName id="9EF3EED9FFD13E0D0EB3FA2FFB9403C1" authority="Saussure, 1877" authorityName="Saussure" authorityYear="1877" box="[689,1124,1489,1515]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="219" phylum="Arthropoda" rank="species" species="acinaceus">
<emphasis id="6B874948FFD13E0D0EB3FA2FFC5F03C1" box="[689,943,1489,1515]" italics="true" pageId="14" pageNumber="219">Pteroplistes acinaceus</emphasis>
Saussure, 1877
</taxonomicName>
from Malay Peninsula and
<taxonomicName id="9EF3EED9FFD13E0D0C95FA0BFCE50025" authority="Tan, Gorochov &amp; Wahab, 2019" authorityName="Tan, Gorochov &amp; Wahab" authorityYear="2019" box="[151,789,1525,1551]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="219" phylum="Arthropoda" rank="species" species="bruneiensis">
<emphasis id="6B874948FFD13E0D0C95FA0BFE560025" box="[151,422,1525,1551]" italics="true" pageId="14" pageNumber="219">Pteroplistes bruneiensis</emphasis>
<bibRefCitation id="3D62E8ABFFD13E0D0DADFA0BFCE50025" author="Tan, M. K. &amp; Gorochov, A. V. &amp; Wahab, R. A. &amp; Japir, R. &amp; Chung, A. Y. C." box="[431,789,1525,1551]" pageId="14" pageNumber="219" pagination="101 - 117" refId="ref8234" refString="Tan, M. K., Gorochov, A. V., Wahab, R. A., Japir, R. &amp; Chung, A. Y. C. (2019) New taxa of crickets (Orthoptera: Grylloidea: Gryllinae, Phaloriinae and Pteroplistinae) from northern Borneo (Belait and Sandakan). Zootaxa, 4661 (1), 101 - 117. https: // doi. org / 10.11646 / zootaxa. 4661.1.4" type="journal article" year="2019">Tan, Gorochov &amp; Wahab, 2019</bibRefCitation>
</taxonomicName>
from Borneo in the ectophallic fold [rachis] of the male genitalia with a pair of rather large and sclerotized processes on its dorsal surface (dp) (before the apical part) but distinguished from them by these processes thickened, almost straight and with acute apices (these processes in
<taxonomicName id="9EF3EED9FFD13E0D0C95F99CFED50051" authorityName="Saussure" authorityYear="1877" box="[151,293,1634,1659]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="219" phylum="Arthropoda" rank="species" species="acinaceus">
<emphasis id="6B874948FFD13E0D0C95F99CFED50051" box="[151,293,1634,1659]" italics="true" pageId="14" pageNumber="219">P. acinaceus</emphasis>
</taxonomicName>
with characteristic hooks distally, and in
<taxonomicName id="9EF3EED9FFD13E0D0F0AF99CFC570051" authorityName="Tan, Gorochov &amp; Wahab" authorityYear="2019" box="[776,935,1633,1659]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="219" phylum="Arthropoda" rank="species" species="bruneiensis">
<emphasis id="6B874948FFD13E0D0F0AF99CFC570051" box="[776,935,1633,1659]" italics="true" pageId="14" pageNumber="219">P. bruneiensis</emphasis>
</taxonomicName>
, they are triangular). It is also similar to
<taxonomicName id="9EF3EED9FFD13E0D098AF99CFEF700B5" authorityName="Saussure" authorityYear="1877" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="219" phylum="Arthropoda" rank="species" species="acinaceus">
<emphasis id="6B874948FFD13E0D098AF99CFEF700B5" italics="true" pageId="14" pageNumber="219">P. acinaceus</emphasis>
</taxonomicName>
,
<taxonomicName id="9EF3EED9FFD13E0D0D15F978FE4500B5" authorityName="Tan, Gorochov &amp; Wahab" authorityYear="2019" box="[279,437,1669,1695]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="219" phylum="Arthropoda" rank="species" species="bruneiensis">
<emphasis id="6B874948FFD13E0D0D15F978FE4500B5" box="[279,437,1669,1695]" italics="true" pageId="14" pageNumber="219">P. bruneiensis</emphasis>
</taxonomicName>
and
<taxonomicName id="9EF3EED9FFD13E0D0DEDF97BFD2900B5" authorityName="Gorochov" authorityYear="2004" box="[495,729,1669,1695]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="219" phylum="Arthropoda" rank="species" species="lagrecai">
<emphasis id="6B874948FFD13E0D0DEDF97BFD2900B5" box="[495,729,1669,1695]" italics="true" pageId="14" pageNumber="219">Pteroplistes lagrecai</emphasis>
</taxonomicName>
by the shapes of the anal plate having a pair of small lobules (hooks), but differs from them by the lobules spaced widely apart (instead of nearly touching each other in
<taxonomicName id="9EF3EED9FFD13E0D098AF954FEE800CD" authorityName="Tan, Gorochov &amp; Wahab" authorityYear="2019" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="219" phylum="Arthropoda" rank="species" species="bruneiensis">
<emphasis id="6B874948FFD13E0D098AF954FEE800CD" italics="true" pageId="14" pageNumber="219">P. bruneiensis</emphasis>
</taxonomicName>
and
<taxonomicName id="9EF3EED9FFD13E0D0D52F933FDC800CD" authorityName="Gorochov" authorityYear="2004" box="[336,568,1741,1767]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="219" phylum="Arthropoda" rank="species" species="lagrecai">
<emphasis id="6B874948FFD13E0D0D52F933FDC800CD" box="[336,568,1741,1767]" italics="true" pageId="14" pageNumber="219">Pteroplistes lagrecai</emphasis>
</taxonomicName>
) and with the apices of the lobules slightly enlarged (instead of tapering into acute apices in
<taxonomicName id="9EF3EED9FFD13E0D0D48F90CFE270121" authorityName="Saussure" authorityYear="1877" box="[330,471,1778,1803]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="219" phylum="Arthropoda" rank="species" species="acinaceus">
<emphasis id="6B874948FFD13E0D0D48F90CFE270121" box="[330,471,1778,1803]" italics="true" pageId="14" pageNumber="219">P. acinaceus</emphasis>
</taxonomicName>
). The new species also differs from congeners from
<collectingCountry id="21E4D5CAFFD13E0D083BF90FFB830121" box="[1081,1139,1777,1803]" name="India" pageId="14" pageNumber="219">India</collectingCountry>
(
<taxonomicName id="9EF3EED9FFD13E0D0887F90FFA660121" authorityName="Jaiswara" authorityYear="2014" box="[1157,1430,1777,1803]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="219" phylum="Arthropoda" rank="species" species="masinagudi">
<emphasis id="6B874948FFD13E0D0887F90FFA660121" box="[1157,1430,1777,1803]" italics="true" pageId="14" pageNumber="219">Pteroplistes masinagudi</emphasis>
</taxonomicName>
,
<taxonomicName id="9EF3EED9FFD13E0D0C95F8EBFE7B0105" authorityName="Jaiswara" authorityYear="2014" box="[151,395,1813,1839]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="219" phylum="Arthropoda" rank="species" species="kervasae">
<emphasis id="6B874948FFD13E0D0C95F8EBFE7B0105" box="[151,395,1813,1839]" italics="true" pageId="14" pageNumber="219">Pteroplistes kervasae</emphasis>
</taxonomicName>
and
<taxonomicName id="9EF3EED9FFD13E0D0DCAF8EBFD330105" authorityName="Bolivar" authorityYear="1900" box="[456,707,1813,1839]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="219" phylum="Arthropoda" rank="species" species="platycleis">
<emphasis id="6B874948FFD13E0D0DCAF8EBFD330105" box="[456,707,1813,1839]" italics="true" pageId="14" pageNumber="219">Pteroplistes platycleis</emphasis>
</taxonomicName>
) by the head dorsum without distinct pale bands behind eyes, shapes of male anal plate and male genitalia.
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFD13E0D0CC5F8A3FC28015D" box="[199,984,1885,1911]" pageId="14" pageNumber="219" type="etymology">
<paragraph id="594C955AFFD13E0D0CC5F8A3FC28015D" blockId="14.[151,1437,1381,1911]" box="[199,984,1885,1911]" pageId="14" pageNumber="219">
<emphasis id="6B874948FFD13E0D0CC5F8A3FEBE015D" bold="true" box="[199,334,1885,1911]" pageId="14" pageNumber="219">Etymology.</emphasis>
The species is named after Mount Silam, its
<typeStatus id="86482BF8FFD13E0D0F45F8A3FC87015D" box="[839,887,1885,1911]" pageId="14" pageNumber="219">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<caption id="0D8CC5D2FFD03E0C0C95F8A6FF3001BF" pageId="15" pageNumber="220" startId="15.[151,250,1880,1904]" targetBox="[151,1436,181,1857]" targetPageId="15" targetType="figure">
<paragraph id="594C955AFFD03E0C0C95F8A6FF3001BF" blockId="15.[151,1436,1880,1941]" pageId="15" pageNumber="220">
<emphasis id="6B874948FFD03E0C0C95F8A6FED1015B" bold="true" box="[151,289,1880,1905]" pageId="15" pageNumber="220">FIGURE 12.</emphasis>
<taxonomicName id="9EF3EED9FFD03E0C0D2BF8A7FE0B015B" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" box="[297,507,1881,1905]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="220" phylum="Arthropoda" rank="species" species="kabili" status="sp. nov.">
<emphasis id="6B874948FFD03E0C0D2BF8A7FE0B015B" box="[297,507,1881,1905]" italics="true" pageId="15" pageNumber="220">Tembelingiola kabili</emphasis>
</taxonomicName>
<emphasis id="6B874948FFD03E0C0E01F8A7FDA5015B" bold="true" box="[515,597,1881,1905]" pageId="15" pageNumber="220">
<taxonomicNameLabel id="70B4F433FFD03E0C0E01F8A7FDA5015B" box="[515,597,1881,1905]" pageId="15" pageNumber="220" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: a male with two females on a tree trunk in Sepilok (A); the male started calling (B).
</paragraph>
</caption>
<subSubSection id="11E9C6D1FFCF3E130CC5FF68FED300CB" pageId="16" pageNumber="221" type="description">
<paragraph id="594C955AFFCF3E130CC5FF68FB3C044B" blockId="16.[151,1437,150,1977]" pageId="16" pageNumber="221">
<emphasis id="6B874948FFCF3E130CC5FF68FEA9069A" bold="true" box="[199,345,150,176]" pageId="16" pageNumber="221">Description.</emphasis>
Body distinctly dorsoventrally compressed, with head and pronotum finely pubescent. Head rostrum 1.3 times as wide as scape, with apex truncated (
<figureCitation id="C1C889DFFFCF3E130F34FF45FC5106FF" box="[822,929,187,213]" captionStart="FIGURE 13" captionStartId="17.[151,250,1744,1768]" captionTargetBox="[152,1436,181,1720]" captionTargetId="figure-17@17.[151,1436,181,1720]" captionTargetPageId="17" captionText="FIGURE 13. Pteroplistes silam sp. nov.: head and pronotum in dorsal (A) and lateral (B) views, face in anterior view (C), left (D) and right (E) FWs in dorsal view, supra-anal plate in dorsal view (F). Scale bars: 5 mm (A, D, E); 2 mm (rest)." pageId="16" pageNumber="221">Fig. 13A</figureCitation>
). Head wider than high in facial view (
<figureCitation id="C1C889DFFFCF3E13096DFF45FF3906D3" captionStart="FIGURE 13" captionStartId="17.[151,250,1744,1768]" captionTargetBox="[152,1436,181,1720]" captionTargetId="figure-17@17.[151,1436,181,1720]" captionTargetPageId="17" captionText="FIGURE 13. Pteroplistes silam sp. nov.: head and pronotum in dorsal (A) and lateral (B) views, face in anterior view (C), left (D) and right (E) FWs in dorsal view, supra-anal plate in dorsal view (F). Scale bars: 5 mm (A, D, E); 2 mm (rest)." pageId="16" pageNumber="221">Fig. 13C</figureCitation>
). Maxillary palps elongated, with apical segment widened apically and subequal to third segment in length, and with subapical segment longest and slightly broadened apically (
<figureCitation id="C1C889DFFFCF3E130FB7FEFDFBEE0737" box="[949,1054,259,285]" captionStart="FIGURE 13" captionStartId="17.[151,250,1744,1768]" captionTargetBox="[152,1436,181,1720]" captionTargetId="figure-17@17.[151,1436,181,1720]" captionTargetPageId="17" captionText="FIGURE 13. Pteroplistes silam sp. nov.: head and pronotum in dorsal (A) and lateral (B) views, face in anterior view (C), left (D) and right (E) FWs in dorsal view, supra-anal plate in dorsal view (F). Scale bars: 5 mm (A, D, E); 2 mm (rest)." pageId="16" pageNumber="221">Fig. 13B</figureCitation>
). Eyes elongated in profile view, slightly protruding anteriorly (
<figureCitation id="C1C889DFFFCF3E130DE5FED9FDBB076B" box="[487,587,295,321]" captionStart="FIGURE 13" captionStartId="17.[151,250,1744,1768]" captionTargetBox="[152,1436,181,1720]" captionTargetId="figure-17@17.[151,1436,181,1720]" captionTargetPageId="17" captionText="FIGURE 13. Pteroplistes silam sp. nov.: head and pronotum in dorsal (A) and lateral (B) views, face in anterior view (C), left (D) and right (E) FWs in dorsal view, supra-anal plate in dorsal view (F). Scale bars: 5 mm (A, D, E); 2 mm (rest)." pageId="16" pageNumber="221">Fig. 13B</figureCitation>
); median ocellus small and rounded; lateral ocelli larger and more elongated; fenestrae between scapes rounded, small like median ocellus (
<figureCitation id="C1C889DFFFCF3E130F58FEB5FC32074F" box="[858,962,331,357]" captionStart="FIGURE 13" captionStartId="17.[151,250,1744,1768]" captionTargetBox="[152,1436,181,1720]" captionTargetId="figure-17@17.[151,1436,181,1720]" captionTargetPageId="17" captionText="FIGURE 13. Pteroplistes silam sp. nov.: head and pronotum in dorsal (A) and lateral (B) views, face in anterior view (C), left (D) and right (E) FWs in dorsal view, supra-anal plate in dorsal view (F). Scale bars: 5 mm (A, D, E); 2 mm (rest)." pageId="16" pageNumber="221">Fig. 13C</figureCitation>
). Pronotal disc 1.3 times as wide as long, finely pubescent along anterior and posterior margins; anterior margin of disc concave; posterior margin of disc straight; lateral margins sub straight and parallel (
<figureCitation id="C1C889DFFFCF3E130ECDFE6DFCC90787" box="[719,825,403,429]" captionStart="FIGURE 13" captionStartId="17.[151,250,1744,1768]" captionTargetBox="[152,1436,181,1720]" captionTargetId="figure-17@17.[151,1436,181,1720]" captionTargetPageId="17" captionText="FIGURE 13. Pteroplistes silam sp. nov.: head and pronotum in dorsal (A) and lateral (B) views, face in anterior view (C), left (D) and right (E) FWs in dorsal view, supra-anal plate in dorsal view (F). Scale bars: 5 mm (A, D, E); 2 mm (rest)." pageId="16" pageNumber="221">Fig. 13A</figureCitation>
). Pronotal lateral lobe 2.1 times as long as high, with ventral margin rising posteriorly (
<figureCitation id="C1C889DFFFCF3E130E19FE49FD7407FB" box="[539,644,439,465]" captionStart="FIGURE 13" captionStartId="17.[151,250,1744,1768]" captionTargetBox="[152,1436,181,1720]" captionTargetId="figure-17@17.[151,1436,181,1720]" captionTargetPageId="17" captionText="FIGURE 13. Pteroplistes silam sp. nov.: head and pronotum in dorsal (A) and lateral (B) views, face in anterior view (C), left (D) and right (E) FWs in dorsal view, supra-anal plate in dorsal view (F). Scale bars: 5 mm (A, D, E); 2 mm (rest)." pageId="16" pageNumber="221">Fig. 13B</figureCitation>
). TI with minute oval tympana on both sides; legs I and II generally pubescent (especially along ventral margin) and with a few stout setae usually located along dorsal margin, and their tarsus with a row of stout setae on ventral surface; FIII pubescent and flattened, without ventral spines; TIII also pubescent, with about 18 small and stout spines on each dorsal side, and with 2 long ventral apical spurs on inner margin and 4 shorter other apical spurs; hind basitarsus with about 7 inner and 7 outer denticles.
</paragraph>
<paragraph id="594C955AFFCF3E130CC5FD95FE8A021F" blockId="16.[151,1437,150,1977]" pageId="16" pageNumber="221">
<emphasis id="6B874948FFCF3E130CC5FD95FEFB04AF" bold="true" box="[199,267,619,645]" pageId="16" pageNumber="221">Male.</emphasis>
Metanotal gland with two lateral areas cream coloured, broadly triangular and slightly raised (
<figureCitation id="C1C889DFFFCF3E13092AFD95FA6004AF" box="[1320,1424,619,645]" captionStart="FIGURE 13" captionStartId="17.[151,250,1744,1768]" captionTargetBox="[152,1436,181,1720]" captionTargetId="figure-17@17.[151,1436,181,1720]" captionTargetPageId="17" captionText="FIGURE 13. Pteroplistes silam sp. nov.: head and pronotum in dorsal (A) and lateral (B) views, face in anterior view (C), left (D) and right (E) FWs in dorsal view, supra-anal plate in dorsal view (F). Scale bars: 5 mm (A, D, E); 2 mm (rest)." pageId="16" pageNumber="221">Fig. 13A</figureCitation>
). FW extending beyond abdominal apex, with dorsal field slightly longer than lateral field (
<figureCitation id="C1C889DFFFCF3E130895FD71FAB20483" box="[1175,1346,655,681]" captionStart="FIGURE 13" captionStartId="17.[151,250,1744,1768]" captionTargetBox="[152,1436,181,1720]" captionTargetId="figure-17@17.[151,1436,181,1720]" captionTargetPageId="17" captionText="FIGURE 13. Pteroplistes silam sp. nov.: head and pronotum in dorsal (A) and lateral (B) views, face in anterior view (C), left (D) and right (E) FWs in dorsal view, supra-anal plate in dorsal view (F). Scale bars: 5 mm (A, D, E); 2 mm (rest)." pageId="16" pageNumber="221">Figs 13D, 13E</figureCitation>
); dorsal tegminal field (
<figureCitation id="C1C889DFFFCF3E130D44FD4DFE1F04E7" box="[326,495,691,717]" captionStart="FIGURE 13" captionStartId="17.[151,250,1744,1768]" captionTargetBox="[152,1436,181,1720]" captionTargetId="figure-17@17.[151,1436,181,1720]" captionTargetPageId="17" captionText="FIGURE 13. Pteroplistes silam sp. nov.: head and pronotum in dorsal (A) and lateral (B) views, face in anterior view (C), left (D) and right (E) FWs in dorsal view, supra-anal plate in dorsal view (F). Scale bars: 5 mm (A, D, E); 2 mm (rest)." pageId="16" pageNumber="221">Figs 13D, 13E</figureCitation>
) in harp area with 6 oblique veins, mostly almost straight and parallel (except for posterior one which slightly sinuous); mirror in this field large and rounded, as wide as long, with anterior margin roundly angular, with posterior margin somewhat more widely rounded, and with two parallel dividing veins (these dividing veins arcuate and located close to one another); apical field about as long as length of mirror, separated from mirror by distinctive transverse cell (
<figureCitation id="C1C889DFFFCF3E130E83FCBDFCDD0577" box="[641,813,835,861]" captionStart="FIGURE 13" captionStartId="17.[151,250,1744,1768]" captionTargetBox="[152,1436,181,1720]" captionTargetId="figure-17@17.[151,1436,181,1720]" captionTargetPageId="17" captionText="FIGURE 13. Pteroplistes silam sp. nov.: head and pronotum in dorsal (A) and lateral (B) views, face in anterior view (C), left (D) and right (E) FWs in dorsal view, supra-anal plate in dorsal view (F). Scale bars: 5 mm (A, D, E); 2 mm (rest)." pageId="16" pageNumber="221">Figs 13D, 13E</figureCitation>
); lateral field wide, with about 20 branches of Sc and about 10 cross-veins (some veins faint) between R and M, as well as with R and M mostly parallel and strongly converging towards the apex (
<figureCitation id="C1C889DFFFCF3E130DF1FC75FD50058F" box="[499,672,907,933]" captionStart="FIGURE 13" captionStartId="17.[151,250,1744,1768]" captionTargetBox="[152,1436,181,1720]" captionTargetId="figure-17@17.[151,1436,181,1720]" captionTargetPageId="17" captionText="FIGURE 13. Pteroplistes silam sp. nov.: head and pronotum in dorsal (A) and lateral (B) views, face in anterior view (C), left (D) and right (E) FWs in dorsal view, supra-anal plate in dorsal view (F). Scale bars: 5 mm (A, D, E); 2 mm (rest)." pageId="16" pageNumber="221">Figs 13D, 13E</figureCitation>
). Hind wings slightly surpassing FWs. Anal plate broadly tongueshaped and slightly bilobed apically, with a pair of small lobules (hooks) in central part of dorsal surface but not very near each other; each of these lobules pointing inwards and tapering into subacute apex (
<figureCitation id="C1C889DFFFCF3E1308ACFC2DFAE105C7" box="[1198,1297,979,1005]" captionStart="FIGURE 13" captionStartId="17.[151,250,1744,1768]" captionTargetBox="[152,1436,181,1720]" captionTargetId="figure-17@17.[151,1436,181,1720]" captionTargetPageId="17" captionText="FIGURE 13. Pteroplistes silam sp. nov.: head and pronotum in dorsal (A) and lateral (B) views, face in anterior view (C), left (D) and right (E) FWs in dorsal view, supra-anal plate in dorsal view (F). Scale bars: 5 mm (A, D, E); 2 mm (rest)." pageId="16" pageNumber="221">Fig. 13F</figureCitation>
). Subgenital plate trapezoidal, about as long as wide; anterior margin wide, tapers slightly posteriorly, and with posterior margin deeply emarginated.
</paragraph>
<paragraph id="594C955AFFCF3E130CC5FBC1FE6A00B3" blockId="16.[151,1437,150,1977]" pageId="16" pageNumber="221">
Male genitalia as shown in
<figureCitation id="C1C889DFFFCF3E130E1FFBC1FD870273" box="[541,631,1087,1113]" captionStart="FIGURE 14" captionStartId="18.[151,250,1548,1572]" captionTargetBox="[166,1421,194,1500]" captionTargetId="figure-20@18.[151,1436,181,1524]" captionTargetPageId="18" captionText="FIGURE 14. Male genitalia of Pteroplistes silam sp. nov. in posterodorsal (A), dorsal (B), ventral (C) and lateral (D) views, before (A) and after (BD) cleaning with KOH. Scale bar: 1 mm." pageId="16" pageNumber="221">Fig. 14</figureCitation>
. Pseudepiphallus [epiphallus] almost H-shaped, with transverse but moderately long median bridge between lateral parts, moderately short posterolateral lobes and long anterolateral parts; pseudepiphallic lophi [posterolateral epiphallic lobes] lamellate, with inner margin strongly sclerotized and concave, with lateral margin distinctly convex, and with apical part forming small hook directed dorso-externally; median bridge (mb) in shape of parallelogram, with anterior margin 1.9 times as wide as long; anterolateral parts of pseudepiphallus lamellate, sinuate and divergent, anteriorly fused with rami. Ectophallic fold [rachis (= guiding rod)] very strongly sclerotized and directed almost perpendicularly downwards in relation to the longitudinal axis of genitalia, with distal half having anterior margin slightly convex, with apical third tapering into acute apex, with basal part having upper process (up) almost perpendicular to ectophallic fold (this process distally bifurcated into two short triangular lobes with subacute apices); dorsum of upper process with pair of dark lateral sclerites (dp) projecting laterally and posteriorly, these projections connect with ectophallic apodemes. Ectophallic apodemes [endoparameres] strongly sclerotized, slender, with middle parts almost touching each other, and with anterior parts in shape of long arcuate ribbons curved downwards and aside (
<figureCitation id="C1C889DFFFCF3E130F58FA11FC300023" box="[858,960,1519,1545]" captionStart="FIGURE 14" captionStartId="18.[151,250,1548,1572]" captionTargetBox="[166,1421,194,1500]" captionTargetId="figure-20@18.[151,1436,181,1524]" captionTargetPageId="18" captionText="FIGURE 14. Male genitalia of Pteroplistes silam sp. nov. in posterodorsal (A), dorsal (B), ventral (C) and lateral (D) views, before (A) and after (BD) cleaning with KOH. Scale bar: 1 mm." pageId="16" pageNumber="221">Fig. 14B</figureCitation>
). Endophallic sclerite [formula (= mold of spermatophore attachment plate)] consists of three small but elongated plates (a pair of lateral oblique plates and unpaired transverse one) as well as a pair of larger and longer but oblique and isolated plates in anteroventral parts of genitalia. Rami rather short and narrow, strongly (arcuately) curved, lamellate and with anterior end tapering almost into acute apex.
</paragraph>
<paragraph id="594C955AFFCF3E130CC5F95DFED300CB" blockId="16.[151,1437,150,1977]" pageId="16" pageNumber="221">
<emphasis id="6B874948FFCF3E130CC5F95DFE8E0097" bold="true" box="[199,382,1699,1725]" pageId="16" pageNumber="221">Measurements.</emphasis>
<specimenCount id="4FF55ED3FFCF3E130D9BF95DFE5E0097" box="[409,430,1699,1725]" count="1" pageId="16" pageNumber="221" type="male"></specimenCount>
<typeStatus id="86482BF8FFCF3E130DB5F95DFDED0097" box="[439,541,1699,1725]" pageId="16" pageNumber="221" type="holotype">holotype</typeStatus>
: BL = 14.4; BWL = 18.8; HL = 1.9; PronL = 3.3; PronW = 4.5; FWL = 15.3; FWW = 6.6.
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFCF3E130CC5F915FBE4012F" box="[199,1044,1770,1797]" pageId="16" pageNumber="221" type="biology_ecology">
<paragraph id="594C955AFFCF3E130CC5F915FBE4012F" blockId="16.[151,1437,150,1977]" box="[199,1044,1770,1797]" pageId="16" pageNumber="221">
<emphasis id="6B874948FFCF3E130CC5F915FEDC012F" bold="true" box="[199,300,1771,1797]" pageId="16" pageNumber="221">Ecology.</emphasis>
This species was found calling on a tree trunk at night (
<figureCitation id="C1C889DFFFCF3E130F9DF915FBF8012F" box="[927,1032,1770,1797]" captionStart="FIGURE 15" captionStartId="19.[151,250,1771,1795]" captionTargetBox="[151,1436,181,1747]" captionTargetId="figure-17@19.[151,1436,181,1747]" captionTargetPageId="19" captionText="FIGURE 15. Pteroplistes silam sp. nov. calling on the tree trunk in Mount Silam (A), and Pteroplistes lagrecai Gorochov, 2004 calling on the tree in Sepilok (B)." pageId="16" pageNumber="221">Fig. 15A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFCF3E130CC5F8F0FC590193" pageId="16" pageNumber="221" type="distribution">
<paragraph id="594C955AFFCF3E130CC5F8F0FCCA0103" blockId="16.[151,1437,150,1977]" box="[199,826,1806,1833]" pageId="16" pageNumber="221">
<emphasis id="6B874948FFCF3E130CC5F8F0FE910102" bold="true" box="[199,353,1806,1832]" pageId="16" pageNumber="221">Distribution.</emphasis>
EAST
<collectingCountry id="21E4D5CAFFCF3E130DB6F8F1FDBA0103" box="[436,586,1807,1833]" name="Malaysia" pageId="16" pageNumber="221">MALAYSIA</collectingCountry>
:
<collectingRegion id="9B375BB8FFCF3E130E56F8F1FD6D0103" box="[596,669,1807,1833]" country="Malaysia" name="Sabah" pageId="16" pageNumber="221">Sabah</collectingRegion>
: Mount Silam
</paragraph>
<paragraph id="594C955AFFCF3E130CC5F8CCFC590193" blockId="16.[151,1437,150,1977]" pageId="16" pageNumber="221">
<emphasis id="6B874948FFCF3E130CC5F8CCFDC50166" bold="true" box="[199,565,1842,1869]" pageId="16" pageNumber="221">
Calling song (
<figureCitation id="C1C889DFFFCF3E130D6BF8CCFE360166" box="[361,454,1842,1868]" captionStart="FIGURE 2" captionStartId="5.[151,250,1600,1624]" captionTargetBox="[161,1426,191,1567]" captionTargetId="figure-17@5.[151,1436,181,1577]" captionTargetPageId="5" captionText="FIGURE 2. Power spectra of the calling songs of the pteroplistine cricket species: Tembelingiola biaculeata sp. nov. (A), Tembelingiola kabili sp. nov. (B), Pteroplistes silam sp. nov. (C), Pteroplistes lagrecai Gorochov, 2004 (D) and Singapuriola separata Gorochov &amp; Tan, 2012 (E). The grey line represents the power [dB] (axis name and scale in grey) and the black solid peak represents the relative amplitude (axis name in black)." pageId="16" pageNumber="221">Figs 2C</figureCitation>
,
<figureCitation id="C1C889DFFFCF3E130DD1F8CCFE060166" box="[467,502,1842,1868]" captionStart="FIGURE 3" captionStartId="6.[151,250,1766,1790]" captionTargetBox="[162,1425,320,1732]" captionTargetId="figure-71@6.[151,1436,309,1743]" captionTargetPageId="6" captionText="FIGURE 3. Oscillograms of the calling songs of the pteroplistine cricket species: three echemes with different lengths of Tembelingiola biaculeata sp. nov. (A), two echeme-sequencesof Tembelingiola kabili sp. nov. (B), continuous trill of Pteroplistes silam sp. nov. (C), an echeme-sequence of Pteroplistes lagrecai Gorochov, 2004 (D) and three echemes of Singapuriola separata Gorochov &amp; Tan, 2012 (E). Scale bar: 1 s." pageId="16" pageNumber="221">3C</figureCitation>
,
<figureCitation id="C1C889DFFFCF3E130E00F8CCFDD70166" box="[514,551,1842,1868]" captionStart="FIGURE 4" captionStartId="7.[151,250,1500,1524]" captionTargetBox="[162,1425,192,1466]" captionTargetId="figure-17@7.[151,1436,181,1477]" captionTargetPageId="7" captionText="FIGURE 4. Oscillograms of the syllables of the pteroplistine cricket species: Tembelingiola biaculeata sp. nov. (A), Tembelingiola kabili sp. nov. (B), Pteroplistes silam sp. nov. (C), Pteroplistes lagrecai Gorochov, 2004 (D) and Singapuriola separata Gorochov &amp; Tan, 2012 (E). Scale bar: 0.05 s." pageId="16" pageNumber="221">4C</figureCitation>
).
</emphasis>
Consists of a trill made up of long syllables irregular spaced-apart. At 23.6°C, average syllable duration is 26.8±1.1 ms (25.629.7 ms) and average silent interval between consecutive syllables is 115.8±40.1 ms (89.8249.2 ms). The frequency spectrum is pure-tonal and forms a clear harmonic series, with the energy peaking at a dominant fundamental frequency of 5.34 kHz.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,141 @@
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<mods:title id="D2CD88E6D96E7D878817449C4E405EB7">New species of little-known crickets from the subfamily Pteroplistinae (Orthoptera, Grylloidea) from Sabah and contribution to their bioacoustics</mods:title>
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Genus
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<emphasis id="6B874948FFD33E0F0CECFA4EFE8A03E1" bold="true" box="[238,378,1456,1483]" italics="true" pageId="12" pageNumber="217">Pteroplistes</emphasis>
<bibRefCitation id="3D62E8ABFFD33E0F0D80FA4EFD0A03E0" author="Brunner von Wattenwyl, C." box="[386,762,1456,1482]" pageId="12" pageNumber="217" pagination="163 - 170" refId="ref7541" refString="Brunner von Wattenwyl, C. (1873) Systeme des Gryllides. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 4 (4), 163 - 170." type="journal article" year="1873">Brunner von Wattenwyl, 1873</bibRefCitation>
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<bibRefCitation id="3D62E8ABFFD33E0F0F43FA0BFBFA0027" author="Chopard, L." box="[833,1034,1525,1549]" pageId="12" pageNumber="217" refId="ref7631" refString="Chopard, L. (1968) Fam. Gryllidae: Subfam. Mogoplistinae, Myrmecophilinae, Scleropterinae, Cachoplistinae, Pteroplistinae, Pentacentrinae, Phalangopsinae, Trigonidiinae, Eneopterinae; Fam. Oecanthidae, Gryllotalpidae. In: Beier, M. (Ed.), Orthopterorum Catalogus. Vol. 12. Junk, ' s-Gravenhage, The Hague, pp. 213 - 500." type="book" year="1968">Chopard 1968: 254</bibRefCitation>
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Tan
<emphasis id="6B874948FFD33E0F0EE0F9EAFCE50001" box="[738,789,1555,1579]" italics="true" pageId="12" pageNumber="217">et al.</emphasis>
, 2019
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<typeStatus id="86482BF8FFD33E0F0C95F9A4FF22005E" box="[151,210,1626,1652]" pageId="12" pageNumber="217">Type</typeStatus>
species:
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<taxonomicName id="9EF3EED9FFD33E0F0D3EF9A4FD54005E" ID-CoL="4Q2NV" authority="Saussure" authorityName="Saussure" authorityYear="1877" box="[316,676,1626,1652]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="217" phylum="Arthropoda" rank="species" species="acinaceus">
<emphasis id="6B874948FFD33E0F0D3EF9A4FDC70059" box="[316,567,1626,1652]" italics="true" pageId="12" pageNumber="217">Pteroplistes acinaceus</emphasis>
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<emphasis id="6B874948FFD73E0B0C95FAF0FE7E0303" bold="true" box="[151,398,1294,1321]" italics="true" pageId="8" pageNumber="213">Tembelingiola kabili</emphasis>
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(
<figureCitation id="C1C889DFFFD73E0B0C9DFACDFF0B0367" box="[159,251,1331,1357]" captionStart="FIGURE 2" captionStartId="5.[151,250,1600,1624]" captionTargetBox="[161,1426,191,1567]" captionTargetId="figure-17@5.[151,1436,181,1577]" captionTargetPageId="5" captionText="FIGURE 2. Power spectra of the calling songs of the pteroplistine cricket species: Tembelingiola biaculeata sp. nov. (A), Tembelingiola kabili sp. nov. (B), Pteroplistes silam sp. nov. (C), Pteroplistes lagrecai Gorochov, 2004 (D) and Singapuriola separata Gorochov &amp; Tan, 2012 (E). The grey line represents the power [dB] (axis name and scale in grey) and the black solid peak represents the relative amplitude (axis name in black)." pageId="8" pageNumber="213">Figs 2B</figureCitation>
,
<figureCitation id="C1C889DFFFD73E0B0D04FACDFED90367" box="[262,297,1331,1357]" captionStart="FIGURE 3" captionStartId="6.[151,250,1766,1790]" captionTargetBox="[162,1425,320,1732]" captionTargetId="figure-71@6.[151,1436,309,1743]" captionTargetPageId="6" captionText="FIGURE 3. Oscillograms of the calling songs of the pteroplistine cricket species: three echemes with different lengths of Tembelingiola biaculeata sp. nov. (A), two echeme-sequencesof Tembelingiola kabili sp. nov. (B), continuous trill of Pteroplistes silam sp. nov. (C), an echeme-sequence of Pteroplistes lagrecai Gorochov, 2004 (D) and three echemes of Singapuriola separata Gorochov &amp; Tan, 2012 (E). Scale bar: 1 s." pageId="8" pageNumber="213">3B</figureCitation>
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<figureCitation id="C1C889DFFFD73E0B0D37FACDFEA80367" box="[309,344,1331,1357]" captionStart="FIGURE 4" captionStartId="7.[151,250,1500,1524]" captionTargetBox="[162,1425,192,1466]" captionTargetId="figure-17@7.[151,1436,181,1477]" captionTargetPageId="7" captionText="FIGURE 4. Oscillograms of the syllables of the pteroplistine cricket species: Tembelingiola biaculeata sp. nov. (A), Tembelingiola kabili sp. nov. (B), Pteroplistes silam sp. nov. (C), Pteroplistes lagrecai Gorochov, 2004 (D) and Singapuriola separata Gorochov &amp; Tan, 2012 (E). Scale bar: 0.05 s." pageId="8" pageNumber="213">4B</figureCitation>
,
<figureCitation id="C1C889DFFFD73E0B0D66FACDFE6C0367" box="[356,412,1331,1357]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[154,1419,181,1900]" captionTargetId="figure-20@10.[153,1434,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Tembelingiola biaculeata sp. nov. (AD) and Tembelingiola kabili sp. nov. (E I): head and pronotum in dorsal (A, E) and lateral (B, F) views, face in anterior view (C, G), male FW in dorsal view (D, H), female FWs in dorsal view (I). Scale bars: 2 mm (D, H, I), 1 mm (rest)." pageId="8" pageNumber="213">7EI</figureCitation>
,
<figureCitation id="C1C889DFFFD73E0B0DAAFACDFE120367" box="[424,482,1331,1357]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStart-2="FIGURE 11" captionStart-3="FIGURE 12" captionStartId-0="12.[151,250,1350,1374]" captionStartId-1="13.[151,250,1588,1612]" captionStartId-2="14.[151,250,1166,1190]" captionStartId-3="15.[151,250,1880,1904]" captionTargetBox-0="[176,1425,193,1326]" captionTargetBox-1="[151,1436,345,1564]" captionTargetBox-2="[151,1436,181,1143]" captionTargetBox-3="[151,1436,181,1857]" captionTargetId-0="figure-20@12.[151,1436,181,1326]" captionTargetId-1="figure-44@13.[151,1436,345,1564]" captionTargetId-2="figure-20@14.[151,1436,181,1143]" captionTargetId-3="figure-17@15.[151,1436,181,1857]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="14" captionTargetPageId-3="15" captionText-0="FIGURE 9. Habitus of Tembelingiola kabili sp. nov., male in dorsal (A) and lateral (B) views. Scale bars: 2 mm." captionText-1="FIGURE 10. Male genitalia of Tembelingiola kabili sp. nov. in dorsal (A), ventral (B) and lateral (C) views. Scale bar: 0.5 mm." captionText-2="FIGURE 11. Habitus of Tembelingiola kabili sp. nov., female in dorsal (A) and lateral (B) views. Scale bars: 2 mm." captionText-3="FIGURE 12. Tembelingiola kabili sp. nov.: a male with two females on a tree trunk in Sepilok (A); the male started calling (B)." pageId="8" pageNumber="213">912</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFD73E0B0C95FA85FE090047" pageId="8" pageNumber="213" type="materials_examined">
<paragraph id="594C955AFFD73E0B0C95FA85FD7A03F7" blockId="8.[151,1437,1403,2041]" pageId="8" pageNumber="213">
<emphasis id="6B874948FFD73E0B0C95FA85FE7703BF" bold="true" box="[151,391,1403,1429]" pageId="8" pageNumber="213">Material examined.</emphasis>
<materialsCitation id="E99B9F07FFD73E0B0D96FA85FD7A03F7" collectingDate="2022-05-18" collectionCode="FRC" collectorName="T. Robillard &amp; M. K. Tan" country="Malaysia" county="Sandakan District" elevation="54" latitude="5.87395" location="Rainforest Discovery Centre" longLatPrecision="1" longitude="117.93871" municipality="Sepilok" pageId="8" pageNumber="213" specimenCount="1" specimenCount-female="1" stateProvince="Sabah" typeStatus="holotype">
<emphasis id="6B874948FFD73E0B0D96FA85FDFB03BF" bold="true" box="[404,523,1403,1429]" pageId="8" pageNumber="213">
<typeStatus id="86482BF8FFD73E0B0D96FA85FDF503BF" box="[404,517,1403,1429]" pageId="8" pageNumber="213" type="holotype">Holotype</typeStatus>
:
</emphasis>
EAST
<collectingCountry id="21E4D5CAFFD73E0B0E69FA85FCF103BF" box="[619,769,1403,1429]" name="Malaysia" pageId="8" pageNumber="213">MALAYSIA</collectingCountry>
:
<specimenCount id="4FF55ED3FFD73E0B0F10FA85FCD803BF" box="[786,808,1403,1429]" count="1" pageId="8" pageNumber="213" type="female"></specimenCount>
;
<collectingRegion id="9B375BB8FFD73E0B0F3EFA85FC3903BF" box="[828,969,1403,1429]" country="Malaysia" name="Sabah" pageId="8" pageNumber="213">Sabah State</collectingRegion>
,
<collectingCounty id="B02DEDD6FFD73E0B0FD9FA85FB5C03BF" box="[987,1196,1403,1429]" pageId="8" pageNumber="213">Sandakan District</collectingCounty>
,
<collectingMunicipality id="B9280F20FFD73E0B08BCFA85FAE603BF" box="[1214,1302,1403,1429]" pageId="8" pageNumber="213">Sepilok</collectingMunicipality>
,
<location id="5C2CC381FFD73E0B092AFA85FE920393" LSID="urn:lsid:plazi:treatment:D15A244CFFD73E080C95FAF0FBDF01F4:5C2CC381FFD73E0B092AFA85FE920393" country="Malaysia" county="Sandakan District" latitude="5.87395" longLatPrecision="1" longitude="117.93871" municipality="Sepilok" name="Rainforest Discovery Centre" pageId="8" pageNumber="213" stateProvince="Sabah">Rainforest Discovery Centre</location>
, dipterocarp forest;
<geoCoordinate id="3CC7F39DFFD73E0B0E4BFA61FD390390" box="[585,713,1439,1466]" degrees="05.87395" direction="north" orientation="latitude" pageId="8" pageNumber="213" precision="1" value="5.87395">N05.87395</geoCoordinate>
,
<geoCoordinate id="3CC7F39DFFD73E0B0ED4FA61FC900393" box="[726,864,1439,1465]" degrees="117.93871" direction="east" orientation="longitude" pageId="8" pageNumber="213" precision="1" value="117.93871">E117.93871</geoCoordinate>
,
<quantity id="9E0B38BFFFD73E0B0F6CFA61FC5C0393" box="[878,940,1439,1466]" metricMagnitude="1" metricUnit="m" metricValue="5.4" pageId="8" pageNumber="213" unit="m" value="54.0">
<elevation id="D2DE7269FFD73E0B0F6CFA61FC5C0393" box="[878,940,1439,1466]" metricMagnitude="1" metricUnit="m" metricValue="5.4" pageId="8" pageNumber="213" unit="m" value="54.0">54 m</elevation>
</quantity>
;
<date id="2D4DB39AFFD73E0B0FB8FA61FBDB0393" box="[954,1067,1439,1465]" pageId="8" pageNumber="213" value="2022-05-18">
<collectingDate id="3D094A72FFD73E0B0FB8FA61FBDB0393" box="[954,1067,1439,1465]" pageId="8" pageNumber="213" value="2022-05-18">18.v.2022</collectingDate>
</date>
, night; on tree trunk (h=
<quantity id="9E0B38BFFFD73E0B094EFA61FA800393" box="[1356,1392,1439,1465]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="213" unit="m" value="1.0">
<elevation id="D2DE7269FFD73E0B094EFA61FA800393" box="[1356,1392,1439,1465]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="213" unit="m" value="1.0">1m</elevation>
</quantity>
);
<collectorName id="F406F08CFFD73E0B0984FA61FEF103F7" pageId="8" pageNumber="213">T. Robillard</collectorName>
&amp;
<collectorName id="F406F08CFFD73E0B0D27FA3DFE6003F7" box="[293,400,1475,1501]" pageId="8" pageNumber="213">M.K. Tan</collectorName>
leg.; TR22-30 (
<collectionCode id="3FE20D9FFFD73E0B0E41FA3DFD8E03F7" box="[579,638,1475,1501]" country="USA" httpUri="http://grbio.org/cool/z9c1-g2b0" name="Fusarium Research Center" pageId="8" pageNumber="213">FRC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="594C955AFFD73E0B0CC5FA19FE090047" blockId="8.[151,1437,1403,2041]" pageId="8" pageNumber="213">
<materialsCitation id="E99B9F07FFD73E0B0CC5FA19FE050047" collectingDate="2022-05-18" collectorName="T. Robillard &amp; M. K. Tan &amp; Pap" country="Malaysia" county="Sandakan District" elevation="54" latitude="5.87395" location="Rainforest Discovery Centre" longLatPrecision="1" longitude="117.93871" municipality="Sepilok" pageId="8" pageNumber="213" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Sabah" typeStatus="paratype">
<emphasis id="6B874948FFD73E0B0CC5FA19FECF002B" bold="true" box="[199,319,1511,1537]" pageId="8" pageNumber="213">
<typeStatus id="86482BF8FFD73E0B0CC5FA19FECF002B" box="[199,319,1511,1537]" pageId="8" pageNumber="213" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="4FF55ED3FFD73E0B0D4DFA19FE84002B" box="[335,372,1511,1537]" count="1" pageId="8" pageNumber="213" type="male">1♂</specimenCount>
,
<specimenCount id="4FF55ED3FFD73E0B0D7DFA19FE50002B" box="[383,416,1511,1537]" count="1" pageId="8" pageNumber="213" type="female">1♀</specimenCount>
): EAST
<collectingCountry id="21E4D5CAFFD73E0B0E05FA19FD6D002B" box="[519,669,1511,1537]" name="Malaysia" pageId="8" pageNumber="213">MALAYSIA</collectingCountry>
: •
<specimenCount id="4FF55ED3FFD73E0B0EB4FA19FD2B002B" box="[694,731,1511,1537]" count="1" pageId="8" pageNumber="213" type="male">1♂</specimenCount>
;
<collectingRegion id="9B375BB8FFD73E0B0EE5FA19FC9D002B" box="[743,877,1511,1537]" country="Malaysia" name="Sabah" pageId="8" pageNumber="213">Sabah State</collectingRegion>
,
<collectingCounty id="B02DEDD6FFD73E0B0F75FA19FBB1002B" box="[887,1089,1511,1537]" pageId="8" pageNumber="213">Sandakan District</collectingCounty>
,
<collectingMunicipality id="B9280F20FFD73E0B0849FA19FB53002B" box="[1099,1187,1511,1537]" pageId="8" pageNumber="213">Sepilok</collectingMunicipality>
,
<location id="5C2CC381FFD73E0B08ACFA19FF15000F" LSID="urn:lsid:plazi:treatment:D15A244CFFD73E080C95FAF0FBDF01F4:5C2CC381FFD73E0B08ACFA19FF15000F" country="Malaysia" county="Sandakan District" latitude="5.87395" longLatPrecision="1" longitude="117.93871" municipality="Sepilok" name="Rainforest Discovery Centre" pageId="8" pageNumber="213" stateProvince="Sabah">Rainforest Discovery Centre</location>
, dipterocarp forest;
<geoCoordinate id="3CC7F39DFFD73E0B0DCAF9F5FDB8000C" box="[456,584,1547,1574]" degrees="05.87395" direction="north" orientation="latitude" pageId="8" pageNumber="213" precision="1" value="5.87395">N05.87395</geoCoordinate>
,
<geoCoordinate id="3CC7F39DFFD73E0B0E56F9F5FD2D000F" box="[596,733,1547,1573]" degrees="117.93871" direction="east" orientation="longitude" pageId="8" pageNumber="213" precision="1" value="117.93871">E117.93871</geoCoordinate>
,
<quantity id="9E0B38BFFFD73E0B0EE8F9F5FCD7000F" box="[746,807,1547,1574]" metricMagnitude="1" metricUnit="m" metricValue="5.4" pageId="8" pageNumber="213" unit="m" value="54.0">
<elevation id="D2DE7269FFD73E0B0EE8F9F5FCD7000F" box="[746,807,1547,1574]" metricMagnitude="1" metricUnit="m" metricValue="5.4" pageId="8" pageNumber="213" unit="m" value="54.0">54 m</elevation>
</quantity>
;
<date id="2D4DB39AFFD73E0B0F36F9F5FC54000F" box="[820,932,1547,1573]" pageId="8" pageNumber="213" value="2022-05-18">
<collectingDate id="3D094A72FFD73E0B0F36F9F5FC54000F" box="[820,932,1547,1573]" pageId="8" pageNumber="213" value="2022-05-18">18.v.2022</collectingDate>
</date>
, night; on tree trunk (h=
<quantity id="9E0B38BFFFD73E0B08BCF9F5FB11000F" box="[1214,1249,1547,1573]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="213" unit="m" value="1.0">
<elevation id="D2DE7269FFD73E0B08BCF9F5FB11000F" box="[1214,1249,1547,1573]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="213" unit="m" value="1.0">1m</elevation>
</quantity>
);
<collectorName id="F406F08CFFD73E0B08F5F9F5FA8F000F" box="[1271,1407,1547,1573]" pageId="8" pageNumber="213">T. Robillard</collectorName>
&amp;
<collectorName id="F406F08CFFD73E0B0C95F9D1FEF30063" box="[151,259,1583,1609]" pageId="8" pageNumber="213">M.K. Tan</collectorName>
leg.; call recording: video files 6566, sound files
<collectorName id="F406F08CFFD73E0B0F40F9D1FC9E0063" box="[834,878,1583,1609]" pageId="8" pageNumber="213">Pap</collectorName>
268271, TR22-30 (MNHN-EO1277) •
<specimenCount id="4FF55ED3FFD73E0B0932F9D1FAA50063" box="[1328,1365,1583,1609]" count="1" pageId="8" pageNumber="213" type="female">1♀</specimenCount>
; same information (MNHN-EO1278)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFD73E0B0CC5F989FD2B016F" pageId="8" pageNumber="213" type="diagnosis">
<paragraph id="594C955AFFD73E0B0CC5F989FD2B016F" blockId="8.[151,1437,1403,2041]" pageId="8" pageNumber="213">
<emphasis id="6B874948FFD73E0B0CC5F989FEB200BB" bold="true" box="[199,322,1655,1681]" pageId="8" pageNumber="213">Diagnosis.</emphasis>
This species differs from all congeners by the male genitalia with its pseudepiphallic lophi [posterolateral epiphallic lobes] straight (bent in
<taxonomicName id="9EF3EED9FFD73E0B0EA5F962FCF0009F" authorityName="Gorochov" authorityYear="2004" box="[679,768,1691,1717]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="213" phylum="Arthropoda" rank="species" species="plana">
<emphasis id="6B874948FFD73E0B0EA5F962FCF0009F" box="[679,768,1691,1717]" italics="true" pageId="8" pageNumber="213">T. plana</emphasis>
</taxonomicName>
and
<taxonomicName id="9EF3EED9FFD73E0B0F32F962FC30009F" authorityName="Tan, Gorochov &amp; Wahab" authorityYear="2019" box="[816,960,1691,1717]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="213" phylum="Arthropoda" rank="species" species="belaitensis">
<emphasis id="6B874948FFD73E0B0F32F962FC30009F" box="[816,960,1691,1717]" italics="true" pageId="8" pageNumber="213">T.belaitensis</emphasis>
</taxonomicName>
) or narrow (narrower than in
<taxonomicName id="9EF3EED9FFD73E0B08FEF962FA7B009F" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" box="[1276,1419,1691,1717]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="213" phylum="Arthropoda" rank="species" species="biaculeata">
<emphasis id="6B874948FFD73E0B08FEF962FA7B009F" box="[1276,1419,1691,1717]" italics="true" pageId="8" pageNumber="213">T.biaculeata</emphasis>
</taxonomicName>
); sclerotized bridge connecting the two lateral parts of pseudepiphallus distinctly longer; pseudepiphallic parameres [ectoparameres] membraneous; ectophallic fold [rachis] with two long and slender anterolateral ribbons; endophallic sclerite [formula] nearly divided into two compact lateral parts (each part somewhat U-shaped); additional plates rather long and narrow as well as obliquely located.
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFD73E0B0CC5F8B1FD4F01A7" pageId="8" pageNumber="213" type="etymology">
<paragraph id="594C955AFFD73E0B0CC5F8B1FD4F01A7" blockId="8.[151,1437,1403,2041]" pageId="8" pageNumber="213">
<emphasis id="6B874948FFD73E0B0CC5F8B1FEBE0143" bold="true" box="[199,334,1871,1897]" pageId="8" pageNumber="213">Etymology.</emphasis>
The species name refers to the Kabili Monster tree, an Obah Suluk tree (
<taxonomicName id="9EF3EED9FFD73E0B0875F8B1FAB10143" box="[1143,1345,1871,1897]" class="Magnoliopsida" family="Dipterocarpaceae" genus="Shorea" kingdom="Plantae" order="Malvales" pageId="8" pageNumber="213" phylum="Tracheophyta" rank="species" species="pauciflora">
<emphasis id="6B874948FFD73E0B0875F8B1FAB10143" box="[1143,1345,1871,1897]" italics="true" pageId="8" pageNumber="213">Shorea pauciflora</emphasis>
</taxonomicName>
) near to where the species was found; noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFD73E080CC5F869FADC008A" lastPageId="11" lastPageNumber="216" pageId="8" pageNumber="213" type="description">
<paragraph id="594C955AFFD73E0B0CC5F869FA8D01D3" blockId="8.[151,1437,1403,2041]" pageId="8" pageNumber="213">
<emphasis id="6B874948FFD73E0B0CC5F869FEA9019B" bold="true" box="[199,345,1943,1969]" pageId="8" pageNumber="213">Description.</emphasis>
General appearance, including tegminal structure, very similar to
<taxonomicName id="9EF3EED9FFD73E0B0851F869FA81019B" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" box="[1107,1393,1943,1969]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="213" phylum="Arthropoda" rank="species" species="biaculeata" status="sp. nov.">
<emphasis id="6B874948FFD73E0B0851F869FA81019B" box="[1107,1393,1943,1969]" italics="true" pageId="8" pageNumber="213">Tembelingiola biaculeata</emphasis>
</taxonomicName>
<emphasis id="6B874948FFD73E0B0979F869FF3801FF" bold="true" pageId="8" pageNumber="213">
<taxonomicNameLabel id="70B4F433FFD73E0B0979F869FF3801FF" pageId="8" pageNumber="213" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
but with following differences (
<figureCitation id="C1C889DFFFD73E0B0E31F845FD8D01FF" box="[563,637,1979,2005]" captionStart="FIGURE 9" captionStartId="12.[151,250,1350,1374]" captionTargetBox="[176,1425,193,1326]" captionTargetId="figure-20@12.[151,1436,181,1326]" captionTargetPageId="12" captionText="FIGURE 9. Habitus of Tembelingiola kabili sp. nov., male in dorsal (A) and lateral (B) views. Scale bars: 2 mm." pageId="8" pageNumber="213">
Figs
<geoCoordinate id="3CC7F39DFFD73E0B0E6FF845FD8D01FF" box="[621,637,1979,2005]" degrees="9, 7" direction="east" orientation="longitude" pageId="8" pageNumber="213" precision="5555" value="9.7">9</geoCoordinate>
</figureCitation>
,
<figureCitation id="C1C889DFFFD73E0B0E8BF845FD2001FF" box="[649,720,1979,2005]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[154,1419,181,1900]" captionTargetId="figure-20@10.[153,1434,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Tembelingiola biaculeata sp. nov. (AD) and Tembelingiola kabili sp. nov. (E I): head and pronotum in dorsal (A, E) and lateral (B, F) views, face in anterior view (C, G), male FW in dorsal view (D, H), female FWs in dorsal view (I). Scale bars: 2 mm (D, H, I), 1 mm (rest)." pageId="8" pageNumber="213">7EG</figureCitation>
): rostrum between antennal cavities wider, 1.3 times wider than scapes (
<figureCitation id="C1C889DFFFD73E0B0CF2F821FEB701D3" box="[240,327,2015,2041]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[154,1419,181,1900]" captionTargetId="figure-20@10.[153,1434,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Tembelingiola biaculeata sp. nov. (AD) and Tembelingiola kabili sp. nov. (E I): head and pronotum in dorsal (A, E) and lateral (B, F) views, face in anterior view (C, G), male FW in dorsal view (D, H), female FWs in dorsal view (I). Scale bars: 2 mm (D, H, I), 1 mm (rest)." pageId="8" pageNumber="213">Fig. 7E</figureCitation>
); maxillary palp with apical segment slightly more elongate; pronotum less pubescent (
<figureCitation id="C1C889DFFFD73E0B091BF821FA8001D3" box="[1305,1392,2015,2041]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[154,1419,181,1900]" captionTargetId="figure-20@10.[153,1434,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Tembelingiola biaculeata sp. nov. (AD) and Tembelingiola kabili sp. nov. (E I): head and pronotum in dorsal (A, E) and lateral (B, F) views, face in anterior view (C, G), male FW in dorsal view (D, H), female FWs in dorsal view (I). Scale bars: 2 mm (D, H, I), 1 mm (rest)." pageId="8" pageNumber="213">Fig. 7E</figureCitation>
).
</paragraph>
<caption id="0D8CC5D2FFD63E0A0C95FB6DFAB50280" box="[151,1349,1171,1195]" pageId="9" pageNumber="214" startId="9.[151,250,1171,1195]" targetBox="[151,1423,195,1147]" targetPageId="9" targetType="figure">
<paragraph id="594C955AFFD63E0A0C95FB6DFAB50280" blockId="9.[151,1349,1171,1195]" box="[151,1349,1171,1195]" pageId="9" pageNumber="214">
<emphasis id="6B874948FFD63E0A0C95FB6DFEE30281" bold="true" box="[151,275,1171,1195]" pageId="9" pageNumber="214">FIGURE 6.</emphasis>
Habitus of
<taxonomicName id="9EF3EED9FFD63E0A0D88FB6DFD7B0281" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" box="[394,651,1171,1195]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="214" phylum="Arthropoda" rank="species" species="biaculeata" status="sp. nov.">
<emphasis id="6B874948FFD63E0A0D88FB6DFD7B0281" box="[394,651,1171,1195]" italics="true" pageId="9" pageNumber="214">Tembelingiola biaculeata</emphasis>
</taxonomicName>
<emphasis id="6B874948FFD63E0A0E90FB6DFD120281" bold="true" box="[658,738,1171,1195]" pageId="9" pageNumber="214">
<taxonomicNameLabel id="70B4F433FFD63E0A0E90FB6DFD120281" box="[658,738,1171,1195]" pageId="9" pageNumber="214" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male in dorsal (A) and lateral (B) views. Scale bars: 2 mm.
</paragraph>
</caption>
<paragraph id="594C955AFFD63E0A0CC5FB11FE4500D0" blockId="9.[151,1437,1263,2035]" pageId="9" pageNumber="214">
<emphasis id="6B874948FFD63E0A0CC5FB11FEFB0323" bold="true" box="[199,267,1263,1289]" pageId="9" pageNumber="214">Male.</emphasis>
FW brown, reaching abdominal apex, 2.2 times as long as wide (
<figureCitation id="C1C889DFFFD63E0A0FE1FB11FBCD0323" box="[995,1085,1263,1289]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[154,1419,181,1900]" captionTargetId="figure-20@10.[153,1434,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Tembelingiola biaculeata sp. nov. (AD) and Tembelingiola kabili sp. nov. (E I): head and pronotum in dorsal (A, E) and lateral (B, F) views, face in anterior view (C, G), male FW in dorsal view (D, H), female FWs in dorsal view (I). Scale bars: 2 mm (D, H, I), 1 mm (rest)." pageId="9" pageNumber="214">Fig. 7H</figureCitation>
). Dorsal field more slender, 2.2 times as long as wide, with only 4 harp veins, and with mirror longer than wide; with a faint diagonal vein; harps with 4 oblique veins including 2 long, parallel sinusoidal posterior veins fused anteriorly, and 2 anterior shorter and fainter one; mirror approximately 1.1 times longer than wide, separated by two dividing veins, dividing veins mostly straight but slightly bent at anal end.Apical field moderately short (
<figureCitation id="C1C889DFFFD63E0A0F7AFA83FC2003BD" box="[888,976,1405,1431]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[154,1419,181,1900]" captionTargetId="figure-20@10.[153,1434,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Tembelingiola biaculeata sp. nov. (AD) and Tembelingiola kabili sp. nov. (E I): head and pronotum in dorsal (A, E) and lateral (B, F) views, face in anterior view (C, G), male FW in dorsal view (D, H), female FWs in dorsal view (I). Scale bars: 2 mm (D, H, I), 1 mm (rest)." pageId="9" pageNumber="214">Fig. 7H</figureCitation>
). Lateral field with about 7 branches of Sc and 9 cross veins between R and M. Hind wings not exposed (
<figureCitation id="C1C889DFFFD63E0A0F43FA5FFC740391" box="[833,900,1441,1467]" captionStart="FIGURE 9" captionStartId="12.[151,250,1350,1374]" captionTargetBox="[176,1425,193,1326]" captionTargetId="figure-20@12.[151,1436,181,1326]" captionTargetPageId="12" captionText="FIGURE 9. Habitus of Tembelingiola kabili sp. nov., male in dorsal (A) and lateral (B) views. Scale bars: 2 mm." pageId="9" pageNumber="214">Fig. 9</figureCitation>
). Genitalia as shown in
<figureCitation id="C1C889DFFFD63E0A088EFA5FFB2D0391" box="[1164,1245,1441,1467]" captionStart="FIGURE 10" captionStartId="13.[151,250,1588,1612]" captionTargetBox="[151,1436,345,1564]" captionTargetId="figure-44@13.[151,1436,345,1564]" captionTargetPageId="13" captionText="FIGURE 10. Male genitalia of Tembelingiola kabili sp. nov. in dorsal (A), ventral (B) and lateral (C) views. Scale bar: 0.5 mm." pageId="9" pageNumber="214">Fig. 10</figureCitation>
. Pseudepiphallus [epiphallus] consisting of two lateral parts connected with each other by broad bridge; pseudepiphallic lophi [posterolateral epiphallic lobes] dorso-ventrally compressed, straight, diverging slightly posteriorly, and with apices roundly truncated; between these lophi, pseudepiphallus posteriorly angularly concave. Pseudepiphallic parameres [ectoparameres] membraneous. Ectophallic fold [rachis] semi-tube-like, narrowing posteriorly into acute apex, slightly surpassing pseudepiphallic bridge, and with a pair of long and slender anterolateral sclerotized ribbons. Endophallic sclerite [formula] lamellate and somewhat transverse but compact, almost divided into two U-shaped lateral parts (each part with medial half more sclerotized and lateral half almost semi-sclerotized. Rami weakly sclerotized, fused with epiphallus. Additional lateral sclerotized plates (p) very long and slender, obliquely located, slightly curved in profile.
</paragraph>
<paragraph id="594C955AFFD63E0A0CC5F8FDFDD701D8" blockId="9.[151,1437,1263,2035]" pageId="9" pageNumber="214">
<emphasis id="6B874948FFD63E0A0CC5F8FDFE660137" bold="true" box="[199,406,1795,1821]" pageId="9" pageNumber="214">
Female (
<figureCitation id="C1C889DFFFD63E0A0D33F8FDFE770137" box="[305,391,1795,1821]" captionStart="FIGURE 11" captionStartId="14.[151,250,1166,1190]" captionTargetBox="[151,1436,181,1143]" captionTargetId="figure-20@14.[151,1436,181,1143]" captionTargetPageId="14" captionText="FIGURE 11. Habitus of Tembelingiola kabili sp. nov., female in dorsal (A) and lateral (B) views. Scale bars: 2 mm." pageId="9" pageNumber="214">Fig. 11</figureCitation>
).
</emphasis>
Colouration and structure of body similar to male. Tegminal dorsal field with 4 oblique longitudinal branches and with numerous and slightly irregular cross-veins (some cross-veins rather long and very oblique, i.e., situated almost longitudinally); most-anal branch after middle splits into two sub-branches (
<figureCitation id="C1C889DFFFD63E0A093DF8B4FA7F014E" box="[1343,1423,1866,1892]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[154,1419,181,1900]" captionTargetId="figure-20@10.[153,1434,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Tembelingiola biaculeata sp. nov. (AD) and Tembelingiola kabili sp. nov. (E I): head and pronotum in dorsal (A, E) and lateral (B, F) views, face in anterior view (C, G), male FW in dorsal view (D, H), female FWs in dorsal view (I). Scale bars: 2 mm (D, H, I), 1 mm (rest)." pageId="9" pageNumber="214">Fig. 7I</figureCitation>
). Lateral field with 3 branches of Sc and with cross-veins between R and M indistinct. Subgenital plate wide but gradually narrowing backwards, with almost widely truncate apex having shallow and rather wide postero-median notch. Ovipositor dark rufous, weakly arcuate in profile, with middle part rather high (wide) and distal third gradually narrowing to acute apex (
<figureCitation id="C1C889DFFFD63E0A0DB6F826FDEA01D8" box="[436,538,2008,2034]" captionStart="FIGURE 11" captionStartId="14.[151,250,1166,1190]" captionTargetBox="[151,1436,181,1143]" captionTargetId="figure-20@14.[151,1436,181,1143]" captionTargetPageId="14" captionText="FIGURE 11. Habitus of Tembelingiola kabili sp. nov., female in dorsal (A) and lateral (B) views. Scale bars: 2 mm." pageId="9" pageNumber="214">Fig. 11B</figureCitation>
).
</paragraph>
<caption id="0D8CC5D2FFD53E090C95F869FE0101D2" pageId="10" pageNumber="215" startId="10.[151,250,1943,1967]" targetBox="[154,1419,181,1900]" targetPageId="10" targetType="figure">
<paragraph id="594C955AFFD53E090C95F869FE0101D2" blockId="10.[151,1436,1943,2040]" pageId="10" pageNumber="215">
<emphasis id="6B874948FFD53E090C95F869FEE2019A" bold="true" box="[151,274,1943,1968]" pageId="10" pageNumber="215">FIGURE 7.</emphasis>
<taxonomicName id="9EF3EED9FFD53E090D15F869FDE70185" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" box="[279,535,1943,1967]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="215" phylum="Arthropoda" rank="species" species="biaculeata" status="sp. nov.">
<emphasis id="6B874948FFD53E090D15F869FDE70185" box="[279,535,1943,1967]" italics="true" pageId="10" pageNumber="215">Tembelingiola biaculeata</emphasis>
</taxonomicName>
<emphasis id="6B874948FFD53E090E1FF866FD9C019A" bold="true" box="[541,620,1944,1968]" pageId="10" pageNumber="215">
<taxonomicNameLabel id="70B4F433FFD53E090E1FF866FD9C019A" box="[541,620,1944,1968]" pageId="10" pageNumber="215" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(AD) and
<taxonomicName id="9EF3EED9FFD53E090EE1F869FC420185" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" box="[739,946,1943,1967]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="215" phylum="Arthropoda" rank="species" species="kabili" status="sp. nov.">
<emphasis id="6B874948FFD53E090EE1F869FC420185" box="[739,946,1943,1967]" italics="true" pageId="10" pageNumber="215">Tembelingiola kabili</emphasis>
</taxonomicName>
<emphasis id="6B874948FFD53E090FB5F866FBF7019A" bold="true" box="[951,1031,1944,1968]" pageId="10" pageNumber="215">
<taxonomicNameLabel id="70B4F433FFD53E090FB5F866FBF7019A" box="[951,1031,1944,1968]" pageId="10" pageNumber="215" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(E I): head and pronotum in dorsal (A, E) and lateral (B, F) views, face in anterior view (C, G), male FW in dorsal view (D, H), female FWs in dorsal view (I). Scale bars: 2 mm (D, H, I), 1 mm (rest).
</paragraph>
</caption>
<caption id="0D8CC5D2FFD43E080C95FA72FF3603E2" pageId="11" pageNumber="216" startId="11.[151,250,1420,1444]" targetBox="[174,1404,193,1378]" targetPageId="11" targetType="figure">
<paragraph id="594C955AFFD43E080C95FA72FF3603E2" blockId="11.[151,1437,1420,1480]" pageId="11" pageNumber="216">
<emphasis id="6B874948FFD43E080C95FA72FEE3038F" bold="true" box="[151,275,1420,1445]" pageId="11" pageNumber="216">FIGURE 8.</emphasis>
Male genitalia of
<taxonomicName id="9EF3EED9FFD43E080DCFFA73FD3F038F" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" box="[461,719,1421,1445]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="216" phylum="Arthropoda" rank="species" species="biaculeata" status="sp. nov.">
<emphasis id="6B874948FFD43E080DCFFA73FD3F038F" box="[461,719,1421,1445]" italics="true" pageId="11" pageNumber="216">Tembelingiola biaculeata</emphasis>
</taxonomicName>
<emphasis id="6B874948FFD43E080ED7FA73FCD6038F" bold="true" box="[725,806,1421,1445]" pageId="11" pageNumber="216">
<taxonomicNameLabel id="70B4F433FFD43E080ED7FA73FCD6038F" box="[725,806,1421,1445]" pageId="11" pageNumber="216" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in dorsal (A), ventral (B) and lateral (C) views. Scale bar: 0.5 mm.
</paragraph>
</caption>
<paragraph id="594C955AFFD43E080CC5F9E5FADC008A" blockId="11.[151,1437,1562,2015]" pageId="11" pageNumber="216">
<emphasis id="6B874948FFD43E080CC5F9E5FE8E001F" bold="true" box="[199,382,1563,1589]" pageId="11" pageNumber="216">Measurements.</emphasis>
<specimenCount id="4FF55ED3FFD43E080D96F9E5FE59001F" box="[404,425,1563,1589]" count="1" pageId="11" pageNumber="216" type="male"></specimenCount>
<typeStatus id="86482BF8FFD43E080DADF9E4FDE5001E" box="[431,533,1562,1588]" pageId="11" pageNumber="216" type="holotype">holotype</typeStatus>
: BL = 8.0; BWL = 9.3; HL = 1.2; PronL = 2.2; PronW = 2.5; FWL = 6.1; FWW = 2.8; FIIIL = 5.8; TIIIL = 3.6; TaIIIL = 2.8 •
<specimenCount id="4FF55ED3FFD43E080E78F9C0FD7F0072" box="[634,655,1598,1624]" count="1" pageId="11" pageNumber="216" type="female"></specimenCount>
<typeStatus id="86482BF8FFD43E080E94F9C0FD0B0072" box="[662,763,1598,1624]" pageId="11" pageNumber="216" type="paratype">paratype</typeStatus>
: BL = 7.9; BWL = 9.5; HL = 1.0; PronL = 2.4; PronW = 2.6; FWL = 6.4; FWW = 1.8; FIIIL = 6.9; TIIIL = 3.9; TaIIIL = 3.0; OL = 5.4 •
<specimenCount id="4FF55ED3FFD43E080FD8F99CFC1F0056" box="[986,1007,1634,1660]" count="1" pageId="11" pageNumber="216" type="female"></specimenCount>
<typeStatus id="86482BF8FFD43E080FF7F99CFBAA0056" box="[1013,1114,1634,1660]" pageId="11" pageNumber="216" type="paratype">paratype</typeStatus>
: BL = 7.3; BWL = 9.5; HL = 1.1; PronL = 2.4; PronW = 2.7; FWL = 6.4; FWW = 1.7; FIIIL = 6.2; TIIIL = 3.5; TaIIIL = 2.9; OL = 5.2
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFD43E080CC5F956FA6800E8" box="[199,1432,1704,1731]" pageId="11" pageNumber="216" type="biology_ecology">
<paragraph id="594C955AFFD43E080CC5F956FA6800E8" blockId="11.[151,1437,1562,2015]" box="[199,1432,1704,1731]" pageId="11" pageNumber="216">
<emphasis id="6B874948FFD43E080CC5F956FEDC00E8" bold="true" box="[199,300,1704,1730]" pageId="11" pageNumber="216">Ecology.</emphasis>
A male was observed producing short echemes behind
<specimenCount id="4FF55ED3FFD43E080F98F957FBD200E8" box="[922,1058,1704,1731]" count="2" pageId="11" pageNumber="216" type="female">two females</specimenCount>
on a tree trunk at night (
<figureCitation id="C1C889DFFFD43E080935F956FA7A00E8" box="[1335,1418,1704,1730]" captionStart="FIGURE 12" captionStartId="15.[151,250,1880,1904]" captionTargetBox="[151,1436,181,1857]" captionTargetId="figure-17@15.[151,1436,181,1857]" captionTargetPageId="15" captionText="FIGURE 12. Tembelingiola kabili sp. nov.: a male with two females on a tree trunk in Sepilok (A); the male started calling (B)." pageId="11" pageNumber="216">Fig. 12</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFD43E080CC5F935FBDF01F4" pageId="11" pageNumber="216" type="distribution">
<paragraph id="594C955AFFD43E080CC5F935FD0D00CC" blockId="11.[151,1437,1562,2015]" box="[199,765,1739,1766]" pageId="11" pageNumber="216">
<emphasis id="6B874948FFD43E080CC5F935FE9100CF" bold="true" box="[199,353,1739,1765]" pageId="11" pageNumber="216">Distribution.</emphasis>
EAST
<collectingCountry id="21E4D5CAFFD43E080DB6F932FDBA00CC" box="[436,586,1740,1766]" name="Malaysia" pageId="11" pageNumber="216">MALAYSIA</collectingCountry>
:
<collectingRegion id="9B375BB8FFD43E080E56F932FD6D00CC" box="[596,669,1740,1766]" country="Malaysia" name="Sabah" pageId="11" pageNumber="216">Sabah</collectingRegion>
: Sepilok
</paragraph>
<paragraph id="594C955AFFD43E080CC5F911FBDF01F4" blockId="11.[151,1437,1562,2015]" pageId="11" pageNumber="216">
<emphasis id="6B874948FFD43E080CC5F911FDC20123" bold="true" box="[199,562,1775,1801]" pageId="11" pageNumber="216">
Calling song (
<figureCitation id="C1C889DFFFD43E080D68F911FE350123" box="[362,453,1775,1801]" captionStart="FIGURE 2" captionStartId="5.[151,250,1600,1624]" captionTargetBox="[161,1426,191,1567]" captionTargetId="figure-17@5.[151,1436,181,1577]" captionTargetPageId="5" captionText="FIGURE 2. Power spectra of the calling songs of the pteroplistine cricket species: Tembelingiola biaculeata sp. nov. (A), Tembelingiola kabili sp. nov. (B), Pteroplistes silam sp. nov. (C), Pteroplistes lagrecai Gorochov, 2004 (D) and Singapuriola separata Gorochov &amp; Tan, 2012 (E). The grey line represents the power [dB] (axis name and scale in grey) and the black solid peak represents the relative amplitude (axis name in black)." pageId="11" pageNumber="216">Figs 2B</figureCitation>
,
<figureCitation id="C1C889DFFFD43E080DD0F911FE040123" box="[466,500,1775,1801]" captionStart="FIGURE 3" captionStartId="6.[151,250,1766,1790]" captionTargetBox="[162,1425,320,1732]" captionTargetId="figure-71@6.[151,1436,309,1743]" captionTargetPageId="6" captionText="FIGURE 3. Oscillograms of the calling songs of the pteroplistine cricket species: three echemes with different lengths of Tembelingiola biaculeata sp. nov. (A), two echeme-sequencesof Tembelingiola kabili sp. nov. (B), continuous trill of Pteroplistes silam sp. nov. (C), an echeme-sequence of Pteroplistes lagrecai Gorochov, 2004 (D) and three echemes of Singapuriola separata Gorochov &amp; Tan, 2012 (E). Scale bar: 1 s." pageId="11" pageNumber="216">3B</figureCitation>
,
<figureCitation id="C1C889DFFFD43E080E02F911FDD30123" box="[512,547,1775,1801]" captionStart="FIGURE 4" captionStartId="7.[151,250,1500,1524]" captionTargetBox="[162,1425,192,1466]" captionTargetId="figure-17@7.[151,1436,181,1477]" captionTargetPageId="7" captionText="FIGURE 4. Oscillograms of the syllables of the pteroplistine cricket species: Tembelingiola biaculeata sp. nov. (A), Tembelingiola kabili sp. nov. (B), Pteroplistes silam sp. nov. (C), Pteroplistes lagrecai Gorochov, 2004 (D) and Singapuriola separata Gorochov &amp; Tan, 2012 (E). Scale bar: 0.05 s." pageId="11" pageNumber="216">4B</figureCitation>
).
</emphasis>
Consists of echeme-sequence, each made up of 6 to 8 echemes. At 26°C, each echeme-sequence has an average duration of 0.84±
<geoCoordinate id="3CC7F39DFFD43E080EE3F8EDFCD70107" box="[737,807,1811,1837]" degrees="0.08" direction="south" orientation="latitude" pageId="11" pageNumber="216" precision="555" value="-0.08">0.08 s</geoCoordinate>
(0.67
<geoCoordinate id="3CC7F39DFFD43E080F7AF8EDFC4F0107" box="[888,959,1811,1837]" degrees="0.99" direction="south" orientation="latitude" pageId="11" pageNumber="216" precision="555" value="-0.99">0.99 s</geoCoordinate>
). The silent interval between consecutive echeme-sequences is 11.9±
<geoCoordinate id="3CC7F39DFFD43E080DC8F8C8FDF0017B" box="[458,512,1846,1873]" degrees="5.7" direction="south" orientation="latitude" pageId="11" pageNumber="216" precision="5555" value="-5.7">5.7 s</geoCoordinate>
(7.4
<geoCoordinate id="3CC7F39DFFD43E080E40F8C8FD78017A" box="[578,648,1846,1872]" degrees="29.7" direction="south" orientation="latitude" pageId="11" pageNumber="216" precision="5555" value="-29.7">29.7 s</geoCoordinate>
). Each echeme consists of 3 syllables and has an average duration of 56.2±1.4 ms (53.659 ms). The silent interval between consecutive echemes is 66.4±3.5 ms (60.572.1 ms). Within each echeme, the average syllable duration is 11.5±0.5 ms (10.512.2 ms) and the average silent interval between consecutive syllables is 9.5±0.5 ms (8.610.2 ms). The frequency spectrum is pure-tonal and forms a clear harmonic series, with the energy peaking at a dominant fundamental frequency of 5.91 kHz.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,432 @@
<document id="F318344294517A427654908B9C9DC8EA" ID-DOI="10.11646/zootaxa.5506.2.3" ID-ISSN="1175-5326" ID-ZooBank="8D2FAB83-6C2B-424E-BDA3-9EB430B9EDDC" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726053455536" checkinUser="plazi" docAuthor="Tan, Ming Kai, Gorochov, Andrei, Japir, Razy, Chung, Arthur Y. C. &amp; Robillard, Tony" docDate="2024" docId="D15A244CFFDB3E0B0C95FE17FD6A0737" docLanguage="en" docName="zootaxa.5506.2.3.pdf" docOrigin="Zootaxa 5506 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5506.2.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tembelingiola biaculeata Tan, Gorochov &amp; Robillard 2024, sp. nov." docType="treatment" docVersion="1" lastPageNumber="213" masterDocId="2D635C34FFDF3E030C02FFFEFFF0062A" masterDocTitle="New species of little-known crickets from the subfamily Pteroplistinae (Orthoptera, Grylloidea) from Sabah and contribution to their bioacoustics" masterLastPageNumber="226" masterPageNumber="205" pageNumber="209" updateTime="1726053593371" updateUser="GgImagineBatch" zenodo-license-document="CLOSED">
<mods:mods id="40E02AF1781125DAEE8684DA5A5AB113" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="51F9D643B8A1E7FAEA65DBAED17448ED">
<mods:title id="A41F73DB94E14341007BF6E7C25234ED">New species of little-known crickets from the subfamily Pteroplistinae (Orthoptera, Grylloidea) from Sabah and contribution to their bioacoustics</mods:title>
</mods:titleInfo>
<mods:name id="72C166E13123DADD7E9DEE9355A68F3B" type="personal">
<mods:role id="D1476BFB7A10F12DF069E993A2354724">
<mods:roleTerm id="1AE929AA3E19B6ABE5F9929D42869E51">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EEDC8745D43B3EEA7DC23D71CA4FEC23">Tan, Ming Kai</mods:namePart>
<mods:affiliation id="2A0F82130AF620A3D2E6315E53AAB2EE">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore &amp; Institut de Systématique, Evolution, Biodiversité (ISYEB), Muséum national dHistoire naturelle, CNRS, SU, EPHE-PSL, UA, 57 rue Cuvier, CP 50, 75231 Paris Cedex 05, France</mods:affiliation>
</mods:name>
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<mods:role id="2D04A625AAFFAAD4D0583BB3A3FC3219">
<mods:roleTerm id="9BB04DE06672650BDF1FEB0C3B2E5771">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="174081ADBCEE3427C381B6A0FDFF6319">Gorochov, Andrei</mods:namePart>
<mods:affiliation id="2795E6CA38B67655C595514CE1BBF677">Zoological Institute, Russian Academy of Sciences, 1 Universitetskaya Emb., St Petersburg 199034, Russia</mods:affiliation>
</mods:name>
<mods:name id="83BE170B7E99920E87250B0D1865DA64" type="personal">
<mods:role id="0722A2EC9DF821F0D1502877AB72F8CC">
<mods:roleTerm id="1E3AEA94599D58FA519F754F57DFBA7F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="904F84725281D586E02B3A91B5F1DA0C">Japir, Razy</mods:namePart>
<mods:affiliation id="15CE4580320FD873D67CB190EF3A2FD5">Forest Research Centre (Sepilok), Sabah Forestry Department, PO Box 1407, 90715 Sandakan, Sabah, Malaysia</mods:affiliation>
</mods:name>
<mods:name id="BD17C4516D7B3D607537E6FE45A796CC" type="personal">
<mods:role id="B9F75D7800319FF154CAC1FC76B9CC7C">
<mods:roleTerm id="DE1454F78786D17463B18C83962E1A43">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A305ADAF7E23216402F9EBCF53B5A8F9">Chung, Arthur Y. C.</mods:namePart>
<mods:affiliation id="60F5B512F1D202A6681EB095BF749B5B">Forest Research Centre (Sepilok), Sabah Forestry Department, PO Box 1407, 90715 Sandakan, Sabah, Malaysia</mods:affiliation>
</mods:name>
<mods:name id="06DF0E8FF2A8466F8970482E89846EEC" type="personal">
<mods:role id="6BE33F6926BD7D3F808DE102689EAE3E">
<mods:roleTerm id="1B3B590361DE0D597CB885DCCCA0B2B7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4EC264CBDCF2359A14F98CEE41B10F59">Robillard, Tony</mods:namePart>
<mods:affiliation id="A62EA24BAD0382B0085CB8BBF1134118">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore &amp; Institut de Systématique, Evolution, Biodiversité (ISYEB), Muséum national dHistoire naturelle, CNRS, SU, EPHE-PSL, UA, 57 rue Cuvier, CP 50, 75231 Paris Cedex 05, France &amp; Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore</mods:affiliation>
</mods:name>
<mods:typeOfResource id="0B557D0F661433FC40CF521E7D036CCA">text</mods:typeOfResource>
<mods:relatedItem id="1E295D666A42ACA43B5854270D2945B5" type="host">
<mods:titleInfo id="754C1C2A41E054ACCEA27B9AC170B34B">
<mods:title id="FA54EF167D763D11A73FAD23E9636F9F">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="AB33185B0EE27139B6F3AEAB4E4AA38D">
<mods:date id="14508173BB011F4042FAA15896B278C4">2024</mods:date>
<mods:detail id="E946AE6841A1A7F03327031D4A1AB505" type="pubDate">
<mods:number id="3C5B24101D9B365DC22F3FEB43F0B55D">2024-09-06</mods:number>
</mods:detail>
<mods:detail id="F72370ED4B58B68EB5A103E375198B30" type="volume">
<mods:number id="60D985B8582AE19BD2499125ABA49048">5506</mods:number>
</mods:detail>
<mods:detail id="B16BEFC4192D9D2257FE550AB877824E" type="issue">
<mods:number id="EA43D4C0CCA21B15EE28D9F2B305195B">2</mods:number>
</mods:detail>
<mods:extent id="7040448214F3FE239073FBFB4B611888" unit="page">
<mods:start id="2C7692E930DD465BB975F83013430AA7">205</mods:start>
<mods:end id="4BA3F0FE1064F0B93F52871D66C93892">226</mods:end>
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<mods:location id="0E6969D86A8703B91493B4D32611E9B5">
<mods:url id="7EC3BCF9C3EE3A3B65B491BDC0F5AABF">http://dx.doi.org/10.11646/zootaxa.5506.2.3</mods:url>
</mods:location>
<mods:classification id="49464D216AFD9D2E5A3C28DF2E7B29AE">journal article</mods:classification>
<mods:identifier id="6B217FCFC9FA23E71641615649D43187" type="DOI">10.11646/zootaxa.5506.2.3</mods:identifier>
<mods:identifier id="DFE776E81441E21777BF9237B6F57382" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="FCA203443FB761BF71249924E1151681" type="ZooBank">8D2FAB83-6C2B-424E-BDA3-9EB430B9EDDC</mods:identifier>
</mods:mods>
<treatment id="D15A244CFFDB3E0B0C95FE17FD6A0737" LSID="urn:lsid:plazi:treatment:D15A244CFFDB3E0B0C95FE17FD6A0737" httpUri="http://treatment.plazi.org/id/D15A244CFFDB3E0B0C95FE17FD6A0737" lastPageId="8" lastPageNumber="213" pageId="4" pageNumber="209">
<subSubSection id="11E9C6D1FFDB3E070C95FE17FC65042E" box="[151,917,489,516]" pageId="4" pageNumber="209" type="nomenclature">
<paragraph id="594C955AFFDB3E070C95FE17FC65042E" blockId="4.[151,917,489,553]" box="[151,917,489,516]" pageId="4" pageNumber="209">
<heading id="02042236FFDB3E070C95FE17FC65042E" bold="true" box="[151,917,489,516]" fontSize="11" level="1" pageId="4" pageNumber="209" reason="1">
<emphasis id="6B874948FFDB3E070C95FE17FC65042E" bold="true" box="[151,917,489,516]" pageId="4" pageNumber="209">
<taxonomicName id="9EF3EED9FFDB3E070C95FE17FCDA0429" authority="Tan, Gorochov &amp; Robillard" authorityName="Tan, Gorochov &amp; Robillard" authorityYear="2024" box="[151,810,489,516]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="species" species="biaculeata" status="sp. nov.">
<emphasis id="6B874948FFDB3E070C95FE17FE37042E" bold="true" box="[151,455,489,516]" italics="true" pageId="4" pageNumber="209">Tembelingiola biaculeata</emphasis>
Tan, Gorochov &amp; Robillard
</taxonomicName>
,
<taxonomicNameLabel id="70B4F433FFDB3E070F35FE14FC65042E" box="[823,917,490,516]" pageId="4" pageNumber="209" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFDB3E070C95FDF0FDFA0402" box="[151,522,526,553]" pageId="4" pageNumber="209" type="description">
<paragraph id="594C955AFFDB3E070C95FDF0FDFA0402" blockId="4.[151,917,489,553]" box="[151,522,526,553]" pageId="4" pageNumber="209">
(
<figureCitation id="C1C889DFFFDB3E070C9DFDF0FF0C0402" box="[159,252,526,552]" captionStart="FIGURE 2" captionStartId="5.[151,250,1600,1624]" captionTargetBox="[161,1426,191,1567]" captionTargetId="figure-17@5.[151,1436,181,1577]" captionTargetPageId="5" captionText="FIGURE 2. Power spectra of the calling songs of the pteroplistine cricket species: Tembelingiola biaculeata sp. nov. (A), Tembelingiola kabili sp. nov. (B), Pteroplistes silam sp. nov. (C), Pteroplistes lagrecai Gorochov, 2004 (D) and Singapuriola separata Gorochov &amp; Tan, 2012 (E). The grey line represents the power [dB] (axis name and scale in grey) and the black solid peak represents the relative amplitude (axis name in black)." pageId="4" pageNumber="209">Figs 2A</figureCitation>
,
<figureCitation id="C1C889DFFFDB3E070D0AFDF0FEDC0402" box="[264,300,526,552]" captionStart="FIGURE 3" captionStartId="6.[151,250,1766,1790]" captionTargetBox="[162,1425,320,1732]" captionTargetId="figure-71@6.[151,1436,309,1743]" captionTargetPageId="6" captionText="FIGURE 3. Oscillograms of the calling songs of the pteroplistine cricket species: three echemes with different lengths of Tembelingiola biaculeata sp. nov. (A), two echeme-sequencesof Tembelingiola kabili sp. nov. (B), continuous trill of Pteroplistes silam sp. nov. (C), an echeme-sequence of Pteroplistes lagrecai Gorochov, 2004 (D) and three echemes of Singapuriola separata Gorochov &amp; Tan, 2012 (E). Scale bar: 1 s." pageId="4" pageNumber="209">3A</figureCitation>
,
<figureCitation id="C1C889DFFFDB3E070D3AFDF0FEAD0402" box="[312,349,526,552]" captionStart="FIGURE 4" captionStartId="7.[151,250,1500,1524]" captionTargetBox="[162,1425,192,1466]" captionTargetId="figure-17@7.[151,1436,181,1477]" captionTargetPageId="7" captionText="FIGURE 4. Oscillograms of the syllables of the pteroplistine cricket species: Tembelingiola biaculeata sp. nov. (A), Tembelingiola kabili sp. nov. (B), Pteroplistes silam sp. nov. (C), Pteroplistes lagrecai Gorochov, 2004 (D) and Singapuriola separata Gorochov &amp; Tan, 2012 (E). Scale bar: 0.05 s." pageId="4" pageNumber="209">4A</figureCitation>
,
<figureCitation id="C1C889DFFFDB3E070D6AFDF0FE880403" box="[360,376,526,553]" captionStart="FIGURE 5" captionStartId="8.[151,250,1188,1212]" captionTargetBox="[151,1435,309,1165]" captionTargetId="figure-78@8.[151,1435,309,1165]" captionTargetPageId="8" captionText="FIGURE 5. Male Tembelingiola biaculeata sp. nov. in its natural habitat in Sepilok." pageId="4" pageNumber="209">5</figureCitation>
,
<figureCitation id="C1C889DFFFDB3E070D86FDF0FE640402" box="[388,404,526,552]" captionStart="FIGURE 6" captionStartId="9.[151,250,1171,1195]" captionTargetBox="[151,1423,195,1147]" captionTargetId="figure-17@9.[151,1436,178,1147]" captionTargetPageId="9" captionText="FIGURE 6. Habitus of Tembelingiola biaculeata sp. nov., male in dorsal (A) and lateral (B) views. Scale bars: 2 mm." pageId="4" pageNumber="209">6</figureCitation>
,
<figureCitation id="C1C889DFFFDB3E070DA2FDF0FE180402" box="[416,488,526,552]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[154,1419,181,1900]" captionTargetId="figure-20@10.[153,1434,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Tembelingiola biaculeata sp. nov. (AD) and Tembelingiola kabili sp. nov. (E I): head and pronotum in dorsal (A, E) and lateral (B, F) views, face in anterior view (C, G), male FW in dorsal view (D, H), female FWs in dorsal view (I). Scale bars: 2 mm (D, H, I), 1 mm (rest)." pageId="4" pageNumber="209">7AD</figureCitation>
,
<figureCitation id="C1C889DFFFDB3E070DF1FDF0FDF10402" box="[499,513,526,552]" captionStart="FIGURE 8" captionStartId="11.[151,250,1420,1444]" captionTargetBox="[174,1404,193,1378]" captionTargetId="figure-17@11.[151,1436,181,1397]" captionTargetPageId="11" captionText="FIGURE 8. Male genitalia of Tembelingiola biaculeata sp. nov. in dorsal (A), ventral (B) and lateral (C) views. Scale bar: 0.5 mm." pageId="4" pageNumber="209">8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFDB3E070C95FDABFD7C050E" pageId="4" pageNumber="209" type="materials_examined">
<paragraph id="594C955AFFDB3E070C95FDABFC560492" blockId="4.[151,1437,597,1920]" pageId="4" pageNumber="209">
<emphasis id="6B874948FFDB3E070C95FDABFE750445" bold="true" box="[151,389,597,623]" pageId="4" pageNumber="209">Material examined.</emphasis>
<materialsCitation id="E99B9F07FFDB3E070D93FDA8FC560492" collectingDate="2022-05-18" collectionCode="FRC" collectorName="M. K. Tan &amp; T. Robillard" country="Malaysia" county="Sandakan District" latitude="5.87552" location="Rainforest Discovery Centre" longLatPrecision="1" longitude="117.94131" municipality="Sepilok" pageId="4" pageNumber="209" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="holotype">
<emphasis id="6B874948FFDB3E070D93FDA8FDF7045A" bold="true" box="[401,519,598,624]" pageId="4" pageNumber="209">
<typeStatus id="86482BF8FFDB3E070D93FDA8FDF2045A" box="[401,514,598,624]" pageId="4" pageNumber="209" type="holotype">Holotype</typeStatus>
:
</emphasis>
EAST
<collectingCountry id="21E4D5CAFFDB3E070E66FDA8FD0A045A" box="[612,762,598,624]" name="Malaysia" pageId="4" pageNumber="209">MALAYSIA</collectingCountry>
: •
<specimenCount id="4FF55ED3FFDB3E070F1FFDA8FCC5045A" box="[797,821,598,624]" count="1" pageId="4" pageNumber="209" type="male"></specimenCount>
;
<collectingRegion id="9B375BB8FFDB3E070F44FDA8FC21045A" box="[838,977,598,624]" country="Malaysia" name="Sabah" pageId="4" pageNumber="209">Sabah State</collectingRegion>
,
<collectingCounty id="B02DEDD6FFDB3E070FE2FDA8FB40045A" box="[992,1200,598,624]" pageId="4" pageNumber="209">Sandakan District</collectingCounty>
,
<collectingMunicipality id="B9280F20FFDB3E0708BDFDA8FAE7045A" box="[1215,1303,598,624]" pageId="4" pageNumber="209">Sepilok</collectingMunicipality>
,
<location id="5C2CC381FFDB3E07092AFDA8FE9204BE" LSID="urn:lsid:plazi:treatment:D15A244CFFDB3E0B0C95FE17FD6A0737:5C2CC381FFDB3E07092AFDA8FE9204BE" country="Malaysia" county="Sandakan District" latitude="5.87552" longLatPrecision="1" longitude="117.94131" municipality="Sepilok" name="Rainforest Discovery Centre" pageId="4" pageNumber="209" stateProvince="Sabah">Rainforest Discovery Centre</location>
, dipterocarp forest;
<geoCoordinate id="3CC7F39DFFDB3E070E48FD84FD4C04BF" box="[586,700,634,661]" degrees="5.87552" direction="north" orientation="latitude" pageId="4" pageNumber="209" precision="1" value="5.87552">N5.87552</geoCoordinate>
,
<geoCoordinate id="3CC7F39DFFDB3E070EC8FD84FCA304BE" box="[714,851,634,660]" degrees="117.94131" direction="east" orientation="longitude" pageId="4" pageNumber="209" precision="1" value="117.94131">E117.94131</geoCoordinate>
, 62.6±6.0 m.a.s.l.;
<date id="2D4DB39AFFDB3E070833FD84FB5204BE" box="[1073,1186,634,660]" pageId="4" pageNumber="209" value="2022-05-18">
<collectingDate id="3D094A72FFDB3E070833FD84FB5204BE" box="[1073,1186,634,660]" pageId="4" pageNumber="209" value="2022-05-18">18.v.2022</collectingDate>
</date>
, 22h36; on foliage of shrub near ground;
<collectorName id="F406F08CFFDB3E070D70FD60FE2D0492" box="[370,477,670,696]" pageId="4" pageNumber="209">M.K. Tan</collectorName>
&amp;
<collectorName id="F406F08CFFDB3E070E02FD60FD770492" box="[512,647,670,696]" pageId="4" pageNumber="209">T. Robillard</collectorName>
leg.; SBH.22.159 (
<collectionCode id="3FE20D9FFFDB3E070F62FD60FC6B0492" box="[864,923,670,696]" country="USA" httpUri="http://grbio.org/cool/z9c1-g2b0" name="Fusarium Research Center" pageId="4" pageNumber="209">FRC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="594C955AFFDB3E070CC5FD3CFD7C050E" blockId="4.[151,1437,597,1920]" pageId="4" pageNumber="209">
<materialsCitation id="E99B9F07FFDB3E070CC5FD3CFD78050E" collectingDate="2022-05-19" collectionCode="ZRC" collectorName="M. K. Tan &amp; T. Robillard" country="Malaysia" county="Sandakan District" elevation="7" latitude="5.87536" location="Rainforest Discovery Centre" longLatPrecision="1" longitude="117.93977" municipality="Sepilok" pageId="4" pageNumber="209" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="paratype">
<emphasis id="6B874948FFDB3E070CC5FD3CFECD04F6" bold="true" box="[199,317,706,732]" pageId="4" pageNumber="209">
<typeStatus id="86482BF8FFDB3E070CC5FD3CFEC804F6" box="[199,312,706,732]" pageId="4" pageNumber="209" type="paratype">Paratype</typeStatus>
:
</emphasis>
EAST
<collectingCountry id="21E4D5CAFFDB3E070D95FD3CFDDD04F6" box="[407,557,706,732]" name="Malaysia" pageId="4" pageNumber="209">MALAYSIA</collectingCountry>
: •
<specimenCount id="4FF55ED3FFDB3E070E4CFD3CFD9604F6" box="[590,614,706,732]" count="1" pageId="4" pageNumber="209" type="male"></specimenCount>
;
<collectingRegion id="9B375BB8FFDB3E070E77FD3CFD0E04F6" box="[629,766,706,732]" country="Malaysia" name="Sabah" pageId="4" pageNumber="209">Sabah State</collectingRegion>
,
<collectingCounty id="B02DEDD6FFDB3E070F0EFD3CFC2A04F6" box="[780,986,706,732]" pageId="4" pageNumber="209">Sandakan District</collectingCounty>
,
<collectingMunicipality id="B9280F20FFDB3E070FEAFD3CFBB004F6" box="[1000,1088,706,732]" pageId="4" pageNumber="209">Sepilok</collectingMunicipality>
,
<location id="5C2CC381FFDB3E07084DFD3CFA6904F6" LSID="urn:lsid:plazi:treatment:D15A244CFFDB3E0B0C95FE17FD6A0737:5C2CC381FFDB3E07084DFD3CFA6904F6" box="[1103,1433,706,732]" country="Malaysia" county="Sandakan District" latitude="5.87536" longLatPrecision="1" longitude="117.93977" municipality="Sepilok" name="Rainforest Discovery Centre" pageId="4" pageNumber="209" stateProvince="Sabah">Rainforest Discovery Centre</location>
, dipterocarp forest;
<geoCoordinate id="3CC7F39DFFDB3E070D6DFD18FE11052B" box="[367,481,742,769]" degrees="5.87536" direction="north" orientation="latitude" pageId="4" pageNumber="209" precision="1" value="5.87536">N5.87536</geoCoordinate>
,
<geoCoordinate id="3CC7F39DFFDB3E070DEFFD18FD87052A" box="[493,631,742,768]" degrees="117.93977" direction="east" orientation="longitude" pageId="4" pageNumber="209" precision="1" value="117.93977">E117.93977</geoCoordinate>
, 57.6±
<quantity id="9E0B38BFFFDB3E070EC6FD18FCFC052A" box="[708,780,742,769]" metricMagnitude="0" metricUnit="m" metricValue="6.8" pageId="4" pageNumber="209" unit="m" value="6.8">
<elevation id="D2DE7269FFDB3E070EC6FD18FCFC052A" box="[708,780,742,769]" metricMagnitude="0" metricUnit="m" metricValue="6.8" pageId="4" pageNumber="209" unit="m" value="6.8">6.8 m</elevation>
</quantity>
.a.s.l.;
<date id="2D4DB39AFFDB3E070F52FD18FC31052A" box="[848,961,742,768]" pageId="4" pageNumber="209" value="2022-05-19">
<collectingDate id="3D094A72FFDB3E070F52FD18FC31052A" box="[848,961,742,768]" pageId="4" pageNumber="209" value="2022-05-19">19.v.2022</collectingDate>
</date>
, 21h06; among branches and twigs;
<collectorName id="F406F08CFFDB3E070963FD18FF31050E" pageId="4" pageNumber="209">M.K. Tan</collectorName>
&amp;
<collectorName id="F406F08CFFDB3E070CE6FCF4FE9B050E" box="[228,363,778,804]" pageId="4" pageNumber="209">T. Robillard</collectorName>
leg.; SBH.22.165 (
<collectionCode id="3FE20D9FFFDB3E070E41FCF4FD8F050E" box="[579,639,778,804]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="209">ZRC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFDB3E070CC5FCD3FBC802A6" pageId="4" pageNumber="209" type="diagnosis">
<paragraph id="594C955AFFDB3E070CC5FCD3FDEA059E" blockId="4.[151,1437,597,1920]" pageId="4" pageNumber="209">
<emphasis id="6B874948FFDB3E070CC5FCD3FEB2056D" bold="true" box="[199,322,813,839]" pageId="4" pageNumber="209">Diagnosis.</emphasis>
This species differs from
<taxonomicName id="9EF3EED9FFDB3E070E71FCD0FBEB0562" authority="Gorochov, 2004" authorityName="Gorochov" authorityYear="2004" box="[627,1051,814,840]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="species" species="plana">
<emphasis id="6B874948FFDB3E070E71FCD0FCAB0562" box="[627,859,814,840]" italics="true" pageId="4" pageNumber="209">Tembelingiola plana</emphasis>
<bibRefCitation id="3D62E8ABFFDB3E070F67FCD0FBEB0562" author="Gorochov, A. V." box="[869,1051,814,840]" pageId="4" pageNumber="209" pagination="379 - 396" refId="ref7847" refString="Gorochov, A. V. (2004) Review of the subfamily Pteroplistinae (Orthoptera Gryllidae). Memorie della Societa Entomologica Italiana, 82 (2), 379 - 396." type="journal article" year="2004">Gorochov, 2004</bibRefCitation>
</taxonomicName>
from Peninsular
<collectingCountry id="21E4D5CAFFDB3E0708E6FCD0FABB0562" box="[1252,1355,814,840]" name="Malaysia" pageId="4" pageNumber="209">Malaysia</collectingCountry>
by the pseudepiphallic paramere [ectoparamere] thicker (less slender), directed downwards (but not backwards), with acute and hooked apical part; and the ectophallic fold [rachis] narrowing posteriorly into acute apex and slightly surpassing pseudepiphallic bridge.
</paragraph>
<paragraph id="594C955AFFDB3E070CC5FC40FBC802A6" blockId="4.[151,1437,597,1920]" pageId="4" pageNumber="209">
It also differs from
<taxonomicName id="9EF3EED9FFDB3E070DB3FC40FBBD05F2" authority="Tan, Gorochov &amp; Wahab, 2019" authorityName="Tan, Gorochov &amp; Wahab" authorityYear="2019" box="[433,1101,958,984]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="species" species="belaitensis">
<emphasis id="6B874948FFDB3E070DB3FC40FD2405F2" box="[433,724,958,984]" italics="true" pageId="4" pageNumber="209">Tembelingiola belaitensis</emphasis>
<bibRefCitation id="3D62E8ABFFDB3E070EDCFC40FBBD05F2" author="Tan, M. K. &amp; Gorochov, A. V. &amp; Wahab, R. A. &amp; Japir, R. &amp; Chung, A. Y. C." box="[734,1101,958,984]" pageId="4" pageNumber="209" pagination="101 - 117" refId="ref8234" refString="Tan, M. K., Gorochov, A. V., Wahab, R. A., Japir, R. &amp; Chung, A. Y. C. (2019) New taxa of crickets (Orthoptera: Grylloidea: Gryllinae, Phaloriinae and Pteroplistinae) from northern Borneo (Belait and Sandakan). Zootaxa, 4661 (1), 101 - 117. https: // doi. org / 10.11646 / zootaxa. 4661.1.4" type="journal article" year="2019">Tan, Gorochov &amp; Wahab, 2019</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="21E4D5CAFFDB3E07089BFC40FA8505F2" box="[1177,1397,958,984]" name="Brunei" pageId="4" pageNumber="209">Brunei Darussalam</collectingCountry>
by the pseudepiphallic lophi [posterolateral epiphallic lobes] directed partly externally (but not curved internally in distal part); pseudepiphallic parameres [ectoparameres] completely sclerotized and somewhat longer but directed downwards (in
<taxonomicName id="9EF3EED9FFDB3E070D4DFBD5FE18026E" authorityName="Tan, Gorochov &amp; Wahab" authorityYear="2019" box="[335,488,1066,1092]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="species" species="belaitensis">
<emphasis id="6B874948FFDB3E070D4DFBD5FE18026E" box="[335,488,1066,1092]" italics="true" pageId="4" pageNumber="209">T. belaitensis</emphasis>
</taxonomicName>
, they are in the form of membranous lobules directed backwards and with thin sclerotized ribbon ventrally); ectophallic fold [rachis] much shorter and less strongly sclerotized; and endophallic sclerite [formula] clearly less wide and not divided into a pair of sclerotized plates.
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFDB3E070CC5FB68FB5102FE" pageId="4" pageNumber="209" type="etymology">
<paragraph id="594C955AFFDB3E070CC5FB68FB5102FE" blockId="4.[151,1437,597,1920]" pageId="4" pageNumber="209">
<emphasis id="6B874948FFDB3E070CC5FB68FEBE029A" bold="true" box="[199,334,1174,1200]" pageId="4" pageNumber="209">Etymology.</emphasis>
The species name refers to the acute apices at the posterior end of both the pseudepiphallic parameres [ectoparameres]; bi- referring to two (a pair), and aculeatus = with spike in Latin.
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFDB3E040CC5FB23FC7D013F" lastPageId="7" lastPageNumber="212" pageId="4" pageNumber="209" type="description">
<paragraph id="594C955AFFDB3E070CC5FB23FA800082" blockId="4.[151,1437,597,1920]" pageId="4" pageNumber="209">
<emphasis id="6B874948FFDB3E070CC5FB23FEA902DD" bold="true" box="[199,345,1245,1271]" pageId="4" pageNumber="209">Description.</emphasis>
Small-sized, body well pubescent, brown with head, pronotum and abdomen with rufous tinge (
<figureCitation id="C1C889DFFFDB3E070C9DFAFCFF180337" box="[159,232,1282,1309]" captionStart="FIGURE 5" captionStartId="8.[151,250,1188,1212]" captionTargetBox="[151,1435,309,1165]" captionTargetId="figure-78@8.[151,1435,309,1165]" captionTargetPageId="8" captionText="FIGURE 5. Male Tembelingiola biaculeata sp. nov. in its natural habitat in Sepilok." pageId="4" pageNumber="209">Figs 5</figureCitation>
,
<figureCitation id="C1C889DFFFDB3E070CF6FAFCFEF40336" box="[244,260,1282,1308]" captionStart="FIGURE 6" captionStartId="9.[151,250,1171,1195]" captionTargetBox="[151,1423,195,1147]" captionTargetId="figure-17@9.[151,1436,178,1147]" captionTargetPageId="9" captionText="FIGURE 6. Habitus of Tembelingiola biaculeata sp. nov., male in dorsal (A) and lateral (B) views. Scale bars: 2 mm." pageId="4" pageNumber="209">6</figureCitation>
). Rostrum between antennal cavities 1.1 times wider than scapes (
<figureCitation id="C1C889DFFFDB3E070FF2FAFCFBBA0336" box="[1008,1098,1282,1308]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[154,1419,181,1900]" captionTargetId="figure-20@10.[153,1434,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Tembelingiola biaculeata sp. nov. (AD) and Tembelingiola kabili sp. nov. (E I): head and pronotum in dorsal (A, E) and lateral (B, F) views, face in anterior view (C, G), male FW in dorsal view (D, H), female FWs in dorsal view (I). Scale bars: 2 mm (D, H, I), 1 mm (rest)." pageId="4" pageNumber="209">Fig. 7A</figureCitation>
), angularly-rounded in profile (
<figureCitation id="C1C889DFFFDB3E070C9DFAD8FF08036A" box="[159,248,1318,1344]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[154,1419,181,1900]" captionTargetId="figure-20@10.[153,1434,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Tembelingiola biaculeata sp. nov. (AD) and Tembelingiola kabili sp. nov. (E I): head and pronotum in dorsal (A, E) and lateral (B, F) views, face in anterior view (C, G), male FW in dorsal view (D, H), female FWs in dorsal view (I). Scale bars: 2 mm (D, H, I), 1 mm (rest)." pageId="4" pageNumber="209">Fig. 7B</figureCitation>
). Maxillary palps pale coloured, with apical, subapical and third segments of subequal lengths; with apical segment elongated-triangular, very broadly expanded apically (
<figureCitation id="C1C889DFFFDB3E070F5EFAB4FC45034E" box="[860,949,1354,1380]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[154,1419,181,1900]" captionTargetId="figure-20@10.[153,1434,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Tembelingiola biaculeata sp. nov. (AD) and Tembelingiola kabili sp. nov. (E I): head and pronotum in dorsal (A, E) and lateral (B, F) views, face in anterior view (C, G), male FW in dorsal view (D, H), female FWs in dorsal view (I). Scale bars: 2 mm (D, H, I), 1 mm (rest)." pageId="4" pageNumber="209">Fig. 7B</figureCitation>
). Head wider than high in facial view. Face dark brown (
<figureCitation id="C1C889DFFFDB3E070D24FA90FE8E03A2" box="[294,382,1390,1416]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[154,1419,181,1900]" captionTargetId="figure-20@10.[153,1434,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Tembelingiola biaculeata sp. nov. (AD) and Tembelingiola kabili sp. nov. (E I): head and pronotum in dorsal (A, E) and lateral (B, F) views, face in anterior view (C, G), male FW in dorsal view (D, H), female FWs in dorsal view (I). Scale bars: 2 mm (D, H, I), 1 mm (rest)." pageId="4" pageNumber="209">Fig. 7C</figureCitation>
); median ocellus present, round; fenestrae between the scapes, anterior of median ocellus, also round and similar in size as median ocellus. Pronotal dorsal disc densely pubescent, weakly transverse, 1.11.3 times as wide as long, faintly widening posteriorly; anterior and posterior margins fairly straight to faintly concave and convex, respectively and with numerous long and thick setae (
<figureCitation id="C1C889DFFFDB3E070F82FA24FC2A03DE" box="[896,986,1498,1524]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[154,1419,181,1900]" captionTargetId="figure-20@10.[153,1434,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Tembelingiola biaculeata sp. nov. (AD) and Tembelingiola kabili sp. nov. (E I): head and pronotum in dorsal (A, E) and lateral (B, F) views, face in anterior view (C, G), male FW in dorsal view (D, H), female FWs in dorsal view (I). Scale bars: 2 mm (D, H, I), 1 mm (rest)." pageId="4" pageNumber="209">Fig. 7A</figureCitation>
). Lateral lobe 1.7 times as long as wide; posterior margin oblique, with numerous long and thick setae (more so than other parts of lateral lobe); ventral margin raised posteriorly; vaguely darker than dorsal disc (
<figureCitation id="C1C889DFFFDB3E070F35F9DCFC610016" box="[823,913,1570,1596]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[154,1419,181,1900]" captionTargetId="figure-20@10.[153,1434,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Tembelingiola biaculeata sp. nov. (AD) and Tembelingiola kabili sp. nov. (E I): head and pronotum in dorsal (A, E) and lateral (B, F) views, face in anterior view (C, G), male FW in dorsal view (D, H), female FWs in dorsal view (I). Scale bars: 2 mm (D, H, I), 1 mm (rest)." pageId="4" pageNumber="209">Fig. 7B</figureCitation>
). Inner tympanum on TI large, elongated oval (opened); outer tympanum undeveloped. TIII with only numerous small dorsal denticles; with 3 pairs of apical spurs, inner ones typically longer than outer ones; hind basitarsus with 2 rows of dorsal spines (each row with 56 spines). Legs generally pale brown. FIII pale brown, but distal part and knee dark rufous, and TIII dark rufous.
</paragraph>
<paragraph id="594C955AFFDB3E070CC5F94CFBBD01AA" blockId="4.[151,1437,597,1920]" pageId="4" pageNumber="209">
<emphasis id="6B874948FFDB3E070CC5F94CFEFB00E6" bold="true" box="[199,267,1714,1740]" pageId="4" pageNumber="209">Male.</emphasis>
FW light brown; barely reaching abdominal apex, 1.81.9 times as long as wide (
<figureCitation id="C1C889DFFFDB3E0708AAF94CFAF300E6" box="[1192,1283,1714,1740]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[154,1419,181,1900]" captionTargetId="figure-20@10.[153,1434,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Tembelingiola biaculeata sp. nov. (AD) and Tembelingiola kabili sp. nov. (E I): head and pronotum in dorsal (A, E) and lateral (B, F) views, face in anterior view (C, G), male FW in dorsal view (D, H), female FWs in dorsal view (I). Scale bars: 2 mm (D, H, I), 1 mm (rest)." pageId="4" pageNumber="209">Fig. 7D</figureCitation>
). Dorsal field with diagonal vein long and slightly sinuous; 5 oblique veins (3 long, parallel sinusoidal posterior ones and 2 anterior shorter and fainter ones); mirror approximately 1.1 times as wide as long, separated by two dividing veins, dividing veins mostly straight but at anal end slightly bent; with moderately short apical area (
<figureCitation id="C1C889DFFFDB3E0708A5F8E0FAF10112" box="[1191,1281,1822,1848]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[154,1419,181,1900]" captionTargetId="figure-20@10.[153,1434,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Tembelingiola biaculeata sp. nov. (AD) and Tembelingiola kabili sp. nov. (E I): head and pronotum in dorsal (A, E) and lateral (B, F) views, face in anterior view (C, G), male FW in dorsal view (D, H), female FWs in dorsal view (I). Scale bars: 2 mm (D, H, I), 1 mm (rest)." pageId="4" pageNumber="209">Fig. 7D</figureCitation>
). Lateral field of FW with about 9 branches of Sc and 12 cross veins between R and M. Hind wings not exposed (
<figureCitation id="C1C889DFFFDB3E0708E3F8BCFAD40176" box="[1249,1316,1858,1884]" captionStart="FIGURE 6" captionStartId="9.[151,250,1171,1195]" captionTargetBox="[151,1423,195,1147]" captionTargetId="figure-17@9.[151,1436,178,1147]" captionTargetPageId="9" captionText="FIGURE 6. Habitus of Tembelingiola biaculeata sp. nov., male in dorsal (A) and lateral (B) views. Scale bars: 2 mm." pageId="4" pageNumber="209">Fig. 6</figureCitation>
). Thoracic sternites brown. Subgenital plate with apex having an angular notched in the middle.
</paragraph>
<caption id="0D8CC5D2FFDA3E060C95F9BEFD0300EF" pageId="5" pageNumber="210" startId="5.[151,250,1600,1624]" targetBox="[161,1426,191,1567]" targetPageId="5" targetType="figure">
<paragraph id="594C955AFFDA3E060C95F9BEFD0300EF" blockId="5.[151,1436,1600,1733]" pageId="5" pageNumber="210">
<emphasis id="6B874948FFDA3E060C95F9BEFEE60072" bold="true" box="[151,278,1600,1624]" pageId="5" pageNumber="210">FIGURE 2.</emphasis>
Power spectra of the calling songs of the pteroplistine cricket species:
<taxonomicName id="9EF3EED9FFDA3E06080AF9BFFAFC0073" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" box="[1032,1292,1601,1625]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="210" phylum="Arthropoda" rank="species" species="biaculeata" status="sp. nov.">
<emphasis id="6B874948FFDA3E06080AF9BFFAFC0073" box="[1032,1292,1601,1625]" italics="true" pageId="5" pageNumber="210">Tembelingiola biaculeata</emphasis>
</taxonomicName>
<emphasis id="6B874948FFDA3E060914F9BFFA990073" bold="true" box="[1302,1385,1601,1625]" pageId="5" pageNumber="210">
<taxonomicNameLabel id="70B4F433FFDA3E060914F9BFFA990073" box="[1302,1385,1601,1625]" pageId="5" pageNumber="210" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A),
<taxonomicName id="9EF3EED9FFDA3E060C95F99BFE980057" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" box="[151,360,1637,1661]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="210" phylum="Arthropoda" rank="species" species="kabili" status="sp. nov.">
<emphasis id="6B874948FFDA3E060C95F99BFE980057" box="[151,360,1637,1661]" italics="true" pageId="5" pageNumber="210">Tembelingiola kabili</emphasis>
</taxonomicName>
<emphasis id="6B874948FFDA3E060D6DF99BFE310057" bold="true" box="[367,449,1637,1661]" pageId="5" pageNumber="210">
<taxonomicNameLabel id="70B4F433FFDA3E060D6DF99BFE310057" box="[367,449,1637,1661]" pageId="5" pageNumber="210" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(B),
<taxonomicName id="9EF3EED9FFDA3E060DF5F99BFD5E0057" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" box="[503,686,1637,1661]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="210" phylum="Arthropoda" rank="species" species="silam" status="sp. nov.">
<emphasis id="6B874948FFDA3E060DF5F99BFD5E0057" box="[503,686,1637,1661]" italics="true" pageId="5" pageNumber="210">Pteroplistes silam</emphasis>
</taxonomicName>
<emphasis id="6B874948FFDA3E060EB7F99BFCF60057" bold="true" box="[693,774,1637,1661]" pageId="5" pageNumber="210">
<taxonomicNameLabel id="70B4F433FFDA3E060EB7F99BFCF60057" box="[693,774,1637,1661]" pageId="5" pageNumber="210" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(C),
<taxonomicName id="9EF3EED9FFDA3E060F3FF99BFB4B0057" authority="Gorochov, 2004" authorityName="Gorochov" authorityYear="2004" box="[829,1211,1637,1661]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="210" phylum="Arthropoda" rank="species" species="lagrecai">
<emphasis id="6B874948FFDA3E060F3FF99BFBFF0057" box="[829,1039,1637,1661]" italics="true" pageId="5" pageNumber="210">Pteroplistes lagrecai</emphasis>
<bibRefCitation id="3D62E8ABFFDA3E060815F99BFB4B0057" author="Gorochov, A. V." box="[1047,1211,1637,1661]" pageId="5" pageNumber="210" pagination="379 - 396" refId="ref7847" refString="Gorochov, A. V. (2004) Review of the subfamily Pteroplistinae (Orthoptera Gryllidae). Memorie della Societa Entomologica Italiana, 82 (2), 379 - 396." type="journal article" year="2004">Gorochov, 2004</bibRefCitation>
</taxonomicName>
(D) and
<taxonomicName id="9EF3EED9FFDA3E06091AF99BFE11008B" authority="Gorochov &amp; Tan, 2012" authorityName="Gorochov &amp; Tan" authorityYear="2012" class="Insecta" family="Pteroplistidae" genus="Singapuriola" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="210" phylum="Arthropoda" rank="species" species="separata">
<emphasis id="6B874948FFDA3E06091AF99BFF00008A" italics="true" pageId="5" pageNumber="210">Singapuriola separata</emphasis>
<bibRefCitation id="3D62E8ABFFDA3E060CF4F977FE11008B" author="Gorochov, A. V. &amp; Tan, M. K." box="[246,481,1673,1697]" pageId="5" pageNumber="210" pagination="18 - 34" refId="ref8057" refString="Gorochov, A. V. &amp; Tan, M. K. (2012) New crickets of the subfamilies Phaloriinae and Pteroplistinae (Orthoptera: Gryllidae) from Singapore. Zootaxa, 3525 (1), 18 - 34. https: // doi. org / 10.11646 / zootaxa. 3525.1.2" type="journal article" year="2012">Gorochov &amp; Tan, 2012</bibRefCitation>
</taxonomicName>
(E). The grey line represents the power [dB] (axis name and scale in grey) and the black solid peak represents the relative amplitude (axis name in black).
</paragraph>
</caption>
<paragraph id="594C955AFFDA3E050CC5F8E9FC9E0737" blockId="5.[151,1437,1815,2021]" lastBlockId="6.[151,1436,151,285]" lastPageId="6" lastPageNumber="211" pageId="5" pageNumber="210">
Genitalia as shown in
<figureCitation id="C1C889DFFFDA3E060DC2F8E9FDF4011B" box="[448,516,1815,1841]" captionStart="FIGURE 8" captionStartId="11.[151,250,1420,1444]" captionTargetBox="[174,1404,193,1378]" captionTargetId="figure-17@11.[151,1436,181,1397]" captionTargetPageId="11" captionText="FIGURE 8. Male genitalia of Tembelingiola biaculeata sp. nov. in dorsal (A), ventral (B) and lateral (C) views. Scale bar: 0.5 mm." pageId="5" pageNumber="210">Fig. 8</figureCitation>
. Pseudepiphallus [epiphallus] divided into two lateral parts. Pseudepiphallic lophi [posterolateral epiphallic lobes] dorso-ventrally compressed; basal third broad, after basal third taper and bent externally, with external margins concave; apex roundly truncated. Pseudepiphallic lophi [epiphallic lateral lobes] connected by narrow bridge; pseudepiphallus with anterior part transversely straight. Pseudepiphallic parameres [ectoparameres] strongly sclerotized, elongated, not directed downwards and with hooks apically; hook with apex acute, not reaching apex of pseudepiphallic lophi. Ectophallic fold [rachis] semi-tube-like, approximately triangular with basal part having two lateral plates; ectophallic fold narrowing posteriorly into acute apex and slightly surpassing pseudepiphallic bridge. Endophallic sclerite [formula] lamellate, transverse and strongly sclerotized with lateral tongues at anterior and posterior parts. Rami fused with pseudepiphallus. Additional lateral sclerotized plate (p) elongated-pyriform with posterior apex narrow and subacute.
</paragraph>
<caption id="0D8CC5D2FFD93E050C95F918FDB20140" pageId="6" pageNumber="211" startId="6.[151,250,1766,1790]" targetBox="[162,1425,320,1732]" targetPageId="6" targetType="figure">
<paragraph id="594C955AFFD93E050C95F918FDB20140" blockId="6.[151,1437,1766,1898]" pageId="6" pageNumber="211">
<emphasis id="6B874948FFD93E050C95F918FEE600D4" bold="true" box="[151,278,1766,1790]" pageId="6" pageNumber="211">FIGURE 3.</emphasis>
Oscillograms of the calling songs of the pteroplistine cricket species: three echemes with different lengths of
<taxonomicName id="9EF3EED9FFD93E050C95F8F4FE660108" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" box="[151,406,1802,1826]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="211" phylum="Arthropoda" rank="species" species="biaculeata" status="sp. nov.">
<emphasis id="6B874948FFD93E050C95F8F4FE660108" box="[151,406,1802,1826]" italics="true" pageId="6" pageNumber="211">Tembelingiola biaculeata</emphasis>
</taxonomicName>
<emphasis id="6B874948FFD93E050D98F8F4FE180108" bold="true" box="[410,488,1802,1826]" pageId="6" pageNumber="211">
<taxonomicNameLabel id="70B4F433FFD93E050D98F8F4FE180108" box="[410,488,1802,1826]" pageId="6" pageNumber="211" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A), two echeme-sequences of
<taxonomicName id="9EF3EED9FFD93E050F1CF8F4FC1C0108" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" box="[798,1004,1802,1826]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="211" phylum="Arthropoda" rank="species" species="kabili" status="sp. nov.">
<emphasis id="6B874948FFD93E050F1CF8F4FC1C0108" box="[798,1004,1802,1826]" italics="true" pageId="6" pageNumber="211">Tembelingiola kabili</emphasis>
</taxonomicName>
<emphasis id="6B874948FFD93E050FF2F8F4FBCE0108" bold="true" box="[1008,1086,1802,1826]" pageId="6" pageNumber="211">
<taxonomicNameLabel id="70B4F433FFD93E050FF2F8F4FBCE0108" box="[1008,1086,1802,1826]" pageId="6" pageNumber="211" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(B), continuous trill of
<taxonomicName id="9EF3EED9FFD93E050926F8F4FF3E016C" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="211" phylum="Arthropoda" rank="species" species="silam" status="sp. nov.">
<emphasis id="6B874948FFD93E050926F8F4FF3E016C" italics="true" pageId="6" pageNumber="211">Pteroplistes silam</emphasis>
</taxonomicName>
<emphasis id="6B874948FFD93E050CD1F8D0FED2016C" bold="true" box="[211,290,1838,1862]" pageId="6" pageNumber="211">
<taxonomicNameLabel id="70B4F433FFD93E050CD1F8D0FED2016C" box="[211,290,1838,1862]" pageId="6" pageNumber="211" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(C), an echeme-sequence of
<taxonomicName id="9EF3EED9FFD93E050E43F8D0FC48016C" authority="Gorochov, 2004" authorityName="Gorochov" authorityYear="2004" box="[577,952,1838,1862]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="211" phylum="Arthropoda" rank="species" species="lagrecai">
<emphasis id="6B874948FFD93E050E43F8D0FCE1016C" box="[577,785,1838,1862]" italics="true" pageId="6" pageNumber="211">Pteroplistes lagrecai</emphasis>
<bibRefCitation id="3D62E8ABFFD93E050F14F8D0FC48016C" author="Gorochov, A. V." box="[790,952,1838,1862]" pageId="6" pageNumber="211" pagination="379 - 396" refId="ref7847" refString="Gorochov, A. V. (2004) Review of the subfamily Pteroplistinae (Orthoptera Gryllidae). Memorie della Societa Entomologica Italiana, 82 (2), 379 - 396." type="journal article" year="2004">Gorochov, 2004</bibRefCitation>
</taxonomicName>
(D) and three echemes of
<taxonomicName id="9EF3EED9FFD93E0508B9F8D0FE720140" authority="Gorochov &amp; Tan, 2012" authorityName="Gorochov &amp; Tan" authorityYear="2012" class="Insecta" family="Pteroplistidae" genus="Singapuriola" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="211" phylum="Arthropoda" rank="species" species="separata">
<emphasis id="6B874948FFD93E0508B9F8D0FA6D016C" box="[1211,1437,1838,1862]" italics="true" pageId="6" pageNumber="211">Singapuriola separata</emphasis>
<bibRefCitation id="3D62E8ABFFD93E050C95F8ACFE720140" author="Gorochov, A. V. &amp; Tan, M. K." box="[151,386,1874,1898]" pageId="6" pageNumber="211" pagination="18 - 34" refId="ref8057" refString="Gorochov, A. V. &amp; Tan, M. K. (2012) New crickets of the subfamilies Phaloriinae and Pteroplistinae (Orthoptera: Gryllidae) from Singapore. Zootaxa, 3525 (1), 18 - 34. https: // doi. org / 10.11646 / zootaxa. 3525.1.2" type="journal article" year="2012">Gorochov &amp; Tan, 2012</bibRefCitation>
</taxonomicName>
(E). Scale bar: 1 s.
</paragraph>
</caption>
<caption id="0D8CC5D2FFD83E040C95FA22FD310016" pageId="7" pageNumber="212" startId="7.[151,250,1500,1524]" targetBox="[162,1425,192,1466]" targetPageId="7" targetType="figure">
<paragraph id="594C955AFFD83E040C95FA22FD310016" blockId="7.[151,1436,1500,1597]" pageId="7" pageNumber="212">
<emphasis id="6B874948FFD83E040C95FA22FEEA03DE" bold="true" box="[151,282,1500,1524]" pageId="7" pageNumber="212">FIGURE 4.</emphasis>
Oscillograms of the syllables of the pteroplistine cricket species:
<taxonomicName id="9EF3EED9FFD83E040FFBFA23FAF103DF" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" box="[1017,1281,1501,1525]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="212" phylum="Arthropoda" rank="species" species="biaculeata" status="sp. nov.">
<emphasis id="6B874948FFD83E040FFBFA23FAF103DF" box="[1017,1281,1501,1525]" italics="true" pageId="7" pageNumber="212">Tembelingiola biaculeata</emphasis>
</taxonomicName>
<emphasis id="6B874948FFD83E04090CFA23FA9503DF" bold="true" box="[1294,1381,1501,1525]" pageId="7" pageNumber="212">
<taxonomicNameLabel id="70B4F433FFD83E04090CFA23FA9503DF" box="[1294,1381,1501,1525]" pageId="7" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A),
<taxonomicName id="9EF3EED9FFD83E040C95F9FFFE980033" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" box="[151,360,1537,1561]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="212" phylum="Arthropoda" rank="species" species="kabili" status="sp. nov.">
<emphasis id="6B874948FFD83E040C95F9FFFE980033" box="[151,360,1537,1561]" italics="true" pageId="7" pageNumber="212">Tembelingiola kabili</emphasis>
</taxonomicName>
<emphasis id="6B874948FFD83E040D6DF9FFFE310033" bold="true" box="[367,449,1537,1561]" pageId="7" pageNumber="212">
<taxonomicNameLabel id="70B4F433FFD83E040D6DF9FFFE310033" box="[367,449,1537,1561]" pageId="7" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(B),
<taxonomicName id="9EF3EED9FFD83E040DF5F9FFFD5E0033" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" box="[503,686,1537,1561]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="212" phylum="Arthropoda" rank="species" species="silam" status="sp. nov.">
<emphasis id="6B874948FFD83E040DF5F9FFFD5E0033" box="[503,686,1537,1561]" italics="true" pageId="7" pageNumber="212">Pteroplistes silam</emphasis>
</taxonomicName>
<emphasis id="6B874948FFD83E040EB7F9FFFCF60033" bold="true" box="[693,774,1537,1561]" pageId="7" pageNumber="212">
<taxonomicNameLabel id="70B4F433FFD83E040EB7F9FFFCF60033" box="[693,774,1537,1561]" pageId="7" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(C),
<taxonomicName id="9EF3EED9FFD83E040F3FF9FFFB4B0033" authority="Gorochov, 2004" authorityName="Gorochov" authorityYear="2004" box="[829,1211,1537,1561]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="212" phylum="Arthropoda" rank="species" species="lagrecai">
<emphasis id="6B874948FFD83E040F3FF9FFFBFF0033" box="[829,1039,1537,1561]" italics="true" pageId="7" pageNumber="212">Pteroplistes lagrecai</emphasis>
<bibRefCitation id="3D62E8ABFFD83E040815F9FFFB4B0033" author="Gorochov, A. V." box="[1047,1211,1537,1561]" pageId="7" pageNumber="212" pagination="379 - 396" refId="ref7847" refString="Gorochov, A. V. (2004) Review of the subfamily Pteroplistinae (Orthoptera Gryllidae). Memorie della Societa Entomologica Italiana, 82 (2), 379 - 396." type="journal article" year="2004">Gorochov, 2004</bibRefCitation>
</taxonomicName>
(D) and
<taxonomicName id="9EF3EED9FFD83E04091AF9FFFE110017" authority="Gorochov &amp; Tan, 2012" authorityName="Gorochov &amp; Tan" authorityYear="2012" class="Insecta" family="Pteroplistidae" genus="Singapuriola" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="212" phylum="Arthropoda" rank="species" species="separata">
<emphasis id="6B874948FFD83E04091AF9FFFF000016" italics="true" pageId="7" pageNumber="212">Singapuriola separata</emphasis>
<bibRefCitation id="3D62E8ABFFD83E040CF4F9DBFE110017" author="Gorochov, A. V. &amp; Tan, M. K." box="[246,481,1573,1597]" pageId="7" pageNumber="212" pagination="18 - 34" refId="ref8057" refString="Gorochov, A. V. &amp; Tan, M. K. (2012) New crickets of the subfamilies Phaloriinae and Pteroplistinae (Orthoptera: Gryllidae) from Singapore. Zootaxa, 3525 (1), 18 - 34. https: // doi. org / 10.11646 / zootaxa. 3525.1.2" type="journal article" year="2012">Gorochov &amp; Tan, 2012</bibRefCitation>
</taxonomicName>
(E). Scale bar: 0.05 s.
</paragraph>
</caption>
<paragraph id="594C955AFFD83E040CC5F971FE6A0083" blockId="7.[151,1437,1679,2030]" box="[199,410,1679,1705]" pageId="7" pageNumber="212">
<emphasis id="6B874948FFD83E040CC5F971FED50083" bold="true" box="[199,293,1679,1705]" pageId="7" pageNumber="212">Female.</emphasis>
Unknown
</paragraph>
<paragraph id="594C955AFFD83E040CC5F94DFC7D013F" blockId="7.[151,1437,1679,2030]" pageId="7" pageNumber="212">
<emphasis id="6B874948FFD83E040CC5F94DFE8E00E7" bold="true" box="[199,382,1715,1741]" pageId="7" pageNumber="212">Measurements.</emphasis>
<specimenCount id="4FF55ED3FFD83E040D94F94DFE5B00E7" box="[406,427,1715,1741]" count="1" pageId="7" pageNumber="212" type="male"></specimenCount>
<typeStatus id="86482BF8FFD83E040DB0F94DFDE800E7" box="[434,536,1715,1741]" pageId="7" pageNumber="212" type="holotype">holotype</typeStatus>
: BL = 8.9; BWL = 9.6; HL = 1.9; PronL = 2.2; PronW = 2.8; FWL = 5.9; FWW = 3.3; FIIIL = 6.3; TIIIL = 4.0; TaIIIL = 3.1 •
<specimenCount id="4FF55ED3FFD83E040E9FF929FD4200DB" box="[669,690,1751,1777]" count="1" pageId="7" pageNumber="212" type="male"></specimenCount>
<typeStatus id="86482BF8FFD83E040EBBF929FCEE00DB" box="[697,798,1751,1777]" pageId="7" pageNumber="212" type="paratype">paratype</typeStatus>
: BL = 8.3; BWL = 9.4; HL = 1.3; PronL = 2.3; PronW = 2.6; FWL = 5.8; FWW = 3.1; FIIIL = 5.9; TIIIL = 3.6; TaIIIL = 3.0.
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFD83E040CC5F8E1FBB70174" pageId="7" pageNumber="212" type="biology_ecology">
<paragraph id="594C955AFFD83E040CC5F8E1FBB70174" blockId="7.[151,1437,1679,2030]" pageId="7" pageNumber="212">
<emphasis id="6B874948FFD83E040CC5F8E1FEDC0113" bold="true" box="[199,300,1823,1849]" pageId="7" pageNumber="212">Ecology.</emphasis>
This species occurs in sympatry with
<taxonomicName id="9EF3EED9FFD83E040ED1F8E1FC480113" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" box="[723,952,1823,1849]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="212" phylum="Arthropoda" rank="species" species="kabili" status="sp. nov.">
<emphasis id="6B874948FFD83E040ED1F8E1FC480113" box="[723,952,1823,1849]" italics="true" pageId="7" pageNumber="212">Tembelingiola kabili</emphasis>
</taxonomicName>
<emphasis id="6B874948FFD83E040FBCF8E1FBE70113" bold="true" box="[958,1047,1823,1849]" pageId="7" pageNumber="212">
<taxonomicNameLabel id="70B4F433FFD83E040FBCF8E1FBE70113" box="[958,1047,1823,1849]" pageId="7" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<specimenCount id="4FF55ED3FFD83E04081FF8E1FB780113" box="[1053,1160,1823,1849]" count="1" pageId="7" pageNumber="212" type="male">One male</specimenCount>
individual was found on a foliage of a low-shrub, whereas another male was found on a dead branch (
<figureCitation id="C1C889DFFFD83E040FF6F8BDFBC90174" box="[1012,1081,1859,1886]" captionStart="FIGURE 5" captionStartId="8.[151,250,1188,1212]" captionTargetBox="[151,1435,309,1165]" captionTargetId="figure-78@8.[151,1435,309,1165]" captionTargetPageId="8" captionText="FIGURE 5. Male Tembelingiola biaculeata sp. nov. in its natural habitat in Sepilok." pageId="7" pageNumber="212">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFD83E0B0CC5F899FD6A0737" lastPageId="8" lastPageNumber="213" pageId="7" pageNumber="212" type="distribution">
<paragraph id="594C955AFFD83E040CC5F899FD0D01AB" blockId="7.[151,1437,1679,2030]" box="[199,765,1895,1921]" pageId="7" pageNumber="212">
<emphasis id="6B874948FFD83E040CC5F899FE9101AB" bold="true" box="[199,353,1895,1921]" pageId="7" pageNumber="212">Distribution.</emphasis>
EAST
<collectingCountry id="21E4D5CAFFD83E040DB6F899FDBA01AB" box="[436,586,1895,1921]" name="Malaysia" pageId="7" pageNumber="212">MALAYSIA</collectingCountry>
:
<collectingRegion id="9B375BB8FFD83E040E56F899FD6D01AB" box="[596,669,1895,1921]" country="Malaysia" name="Sabah" pageId="7" pageNumber="212">Sabah</collectingRegion>
: Sepilok
</paragraph>
<paragraph id="594C955AFFD83E040CC5F875FA6D01C7" blockId="7.[151,1437,1679,2030]" pageId="7" pageNumber="212">
<emphasis id="6B874948FFD83E040CC5F875FDCB018E" bold="true" box="[199,571,1930,1957]" pageId="7" pageNumber="212">
Calling song (
<figureCitation id="C1C889DFFFD83E040D6EF874FE3A018F" box="[364,458,1930,1957]" captionStart="FIGURE 2" captionStartId="5.[151,250,1600,1624]" captionTargetBox="[161,1426,191,1567]" captionTargetId="figure-17@5.[151,1436,181,1577]" captionTargetPageId="5" captionText="FIGURE 2. Power spectra of the calling songs of the pteroplistine cricket species: Tembelingiola biaculeata sp. nov. (A), Tembelingiola kabili sp. nov. (B), Pteroplistes silam sp. nov. (C), Pteroplistes lagrecai Gorochov, 2004 (D) and Singapuriola separata Gorochov &amp; Tan, 2012 (E). The grey line represents the power [dB] (axis name and scale in grey) and the black solid peak represents the relative amplitude (axis name in black)." pageId="7" pageNumber="212">Figs 2A</figureCitation>
,
<figureCitation id="C1C889DFFFD83E040DDAF875FE0B018F" box="[472,507,1931,1957]" captionStart="FIGURE 3" captionStartId="6.[151,250,1766,1790]" captionTargetBox="[162,1425,320,1732]" captionTargetId="figure-71@6.[151,1436,309,1743]" captionTargetPageId="6" captionText="FIGURE 3. Oscillograms of the calling songs of the pteroplistine cricket species: three echemes with different lengths of Tembelingiola biaculeata sp. nov. (A), two echeme-sequencesof Tembelingiola kabili sp. nov. (B), continuous trill of Pteroplistes silam sp. nov. (C), an echeme-sequence of Pteroplistes lagrecai Gorochov, 2004 (D) and three echemes of Singapuriola separata Gorochov &amp; Tan, 2012 (E). Scale bar: 1 s." pageId="7" pageNumber="212">3A</figureCitation>
,
<figureCitation id="C1C889DFFFD83E040E0BF874FDDE018E" box="[521,558,1930,1956]" captionStart="FIGURE 4" captionStartId="7.[151,250,1500,1524]" captionTargetBox="[162,1425,192,1466]" captionTargetId="figure-17@7.[151,1436,181,1477]" captionTargetPageId="7" captionText="FIGURE 4. Oscillograms of the syllables of the pteroplistine cricket species: Tembelingiola biaculeata sp. nov. (A), Tembelingiola kabili sp. nov. (B), Pteroplistes silam sp. nov. (C), Pteroplistes lagrecai Gorochov, 2004 (D) and Singapuriola separata Gorochov &amp; Tan, 2012 (E). Scale bar: 0.05 s." pageId="7" pageNumber="212">4A</figureCitation>
).
</emphasis>
Consists of either short or long echemes. At 31.5031.75°C, a short echeme has a duration of 0.26±
<geoCoordinate id="3CC7F39DFFD83E040DA3F851FE1701E3" box="[417,487,1967,1993]" degrees="0.02" direction="south" orientation="latitude" pageId="7" pageNumber="212" precision="555" value="-0.02">0.02 s</geoCoordinate>
(0.23
<geoCoordinate id="3CC7F39DFFD83E040E35F851FD8D01E3" box="[567,637,1967,1993]" degrees="0.29" direction="south" orientation="latitude" pageId="7" pageNumber="212" precision="555" value="-0.29">0.29 s</geoCoordinate>
) and consists of an average of nine syllables (810 syllables). A long echeme has a duration of 3.02±
<geoCoordinate id="3CC7F39DFFD83E040DFAF82DFDCB01C7" box="[504,571,2003,2029]" degrees="0.48" direction="south" orientation="latitude" pageId="7" pageNumber="212" precision="555" value="-0.48">0.48 s</geoCoordinate>
(2.50
<geoCoordinate id="3CC7F39DFFD83E040E88F82DFD3E01C7" box="[650,718,2003,2030]" degrees="3.77" direction="south" orientation="latitude" pageId="7" pageNumber="212" precision="555" value="-3.77">3.77 s</geoCoordinate>
) and consists of an average of 101 syllables (82127 syllables).
</paragraph>
<paragraph id="594C955AFFD73E0B0C95FF69FD6A0737" blockId="8.[151,1437,151,285]" pageId="8" pageNumber="213">
The silent interval between consecutive echemes is 1.7±
<geoCoordinate id="3CC7F39DFFD73E0B0F09FF69FCB1069B" box="[779,833,151,177]" degrees="0.8" direction="south" orientation="latitude" pageId="8" pageNumber="213" precision="5555" value="-0.8">0.8 s</geoCoordinate>
(0.9
<geoCoordinate id="3CC7F39DFFD73E0B0F83FF69FC48069B" box="[897,952,151,177]" degrees="3.4" direction="south" orientation="latitude" pageId="8" pageNumber="213" precision="5555" value="-3.4">3.4 s</geoCoordinate>
). Within each echeme, the average syllable duration is 18.8±2.1 ms (14.021.9 ms) and the average silent interval between consecutive syllables is 11.1±2.9 ms (7.317.3 ms). The frequency spectrum is pure-tonal and forms a harmonics, with the energy peaking at a dominant frequency of 5.66±0.09 kHz (5.535.72 kHz).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,111 @@
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<mods:title id="8CAAC40EB059A9B70043ECED7592F729">New species of little-known crickets from the subfamily Pteroplistinae (Orthoptera, Grylloidea) from Sabah and contribution to their bioacoustics</mods:title>
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<mods:identifier id="5E2F6BA928F70D6B0404728D8395E734" type="ZooBank">7385E4AA-D7AD-4E79-A05C-684319E0843B</mods:identifier>
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<treatment id="FA1F073A4507FF82FF411E84FB1329F0" LSID="urn:lsid:zoobank.org:act:D8C20638-F2AA-4F8D-A762-31CB4CB6C241" httpUri="http://treatment.plazi.org/id/FA1F073A4507FF82FF411E84FB1329F0" lastPageId="62" lastPageNumber="63" pageId="50" pageNumber="51">
<subSubSection id="3AACE5A74507FF8EFF411E84FC5D2B3C" pageId="50" pageNumber="51" type="nomenclature">
@ -81,12 +82,12 @@ urn:lsid:zoobank.org:act:
<subSubSection id="3AACE5A74507FF8EFF411D5DFE242B18" box="[151,426,1555,1581]" pageId="50" pageNumber="51" type="description">
<paragraph id="7209B62C4507FF8EFF411D5DFE242B18" blockId="50.[151,979,1482,1581]" box="[151,426,1555,1581]" pageId="50" pageNumber="51">
(
<figureCitation id="EA8DAAA94507FF8EFF491D5DFECA2B18" box="[159,324,1555,1581]" captionStart-0="FIGURE 32" captionStart-1="FIGURE 33" captionStart-2="FIGURE 34" captionStart-3="FIGURE 35" captionStart-4="FIGURE 36" captionStart-5="FIGURE 37" captionStart-6="FIGURE 38" captionStart-7="FIGURE 39" captionStartId-0="51.[151,250,1943,1967]" captionStartId-1="52.[151,250,1520,1544]" captionStartId-2="54.[151,250,1520,1544]" captionStartId-3="55.[151,250,1943,1967]" captionStartId-4="56.[151,250,1943,1967]" captionStartId-5="57.[151,250,1981,2005]" captionStartId-6="58.[151,250,1943,1967]" captionStartId-7="59.[151,250,1960,1984]" captionTargetBox-0="[179,1408,181,1920]" captionTargetBox-1="[151,1436,181,1496]" captionTargetBox-2="[151,1436,181,1496]" captionTargetBox-3="[186,1401,185,1915]" captionTargetBox-4="[166,1418,185,1915]" captionTargetBox-5="[223,1388,222,1895]" captionTargetBox-6="[189,1403,188,1912]" captionTargetBox-7="[175,1409,202,1932]" captionTargetId-0="figure-17@51.[179,1408,181,1920]" captionTargetId-1="figure-31@52.[151,1436,181,1496]" captionTargetId-2="figure-31@54.[151,1436,181,1496]" captionTargetId-3="figure-17@55.[182,1405,181,1919]" captionTargetId-4="figure-31@56.[166,1422,181,1919]" captionTargetId-5="figure-17@57.[166,1422,181,1957]" captionTargetId-6="figure-31@58.[179,1408,181,1919]" captionTargetId-7="figure-17@59.[175,1413,198,1936]" captionTargetPageId-0="51" captionTargetPageId-1="52" captionTargetPageId-2="54" captionTargetPageId-3="55" captionTargetPageId-4="56" captionTargetPageId-5="57" captionTargetPageId-6="58" captionTargetPageId-7="59" captionText-0="FIGURE 32. Benincandona trapezoides gen. et spec. nov., valves (AC, GK ♂; DF ♀). AF, H. CpRl (AC, H. INV.159276, DF. INV.1592 INV.159279). G, I, J. INV.159279. CpV. K. INV.159277. CpD. Scale bars. 1 (B, E, G, K) = 500 µm; 2 (A, C, D, F, I) = 200 µm, 3 (H, J) = 40 µm." captionText-1="FIGURE 33. Benincandona trapezoides gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159263). B, E, F, H. RVi (INV.159263). G, H. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (GH) = 50 µm." captionText-2="FIGURE 34. Benincandona trapezoides gen. et spec. nov., valves (♀). A, C, D, G. LVi (INV.159264). B, E, F, H. RVi (INV.159264). G, H. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (GH) = 50 µm." captionText-3="FIGURE 35. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. A2 (INV.159280). B. A1 (INV.159265). C. MdCoxa (INV.159263). Scale bar = 50 µm." captionText-4="FIGURE 36. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Mx1, respiratory plate (INV.159263). B. Mx1 (Chaetotaxy incomplete) (INV.159263). C. MdPalp (INV.159263). Scale bar. 1 (A), 2 (C), 3 (B) = 50 µm." captionText-5="FIGURE 37. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. T2 (INV.159263). B. T3 (INV.159263). C. CR (INV.159280). D. CRAtt (INV.159280). Scale bar = 50 µm." captionText-6="FIGURE 38. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159263). BD. Hp (INV.159263). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." captionText-7="FIGURE 39. Benincandona trapezoides gen. et spec. nov., limbs (AB ♀ and CE ♂). A. T1, Lpp (INV.159280). B. T1, Rpp (INV.159280). C. A2 (INV.159264, INV.159269). D. T1 (INV.159268). E. CR (INV.159264). Scale bar = 50 µm." pageId="50" pageNumber="51">
<figureCitation id="EA8DAAA94507FF8EFF491D5DFECA2B18" box="[159,324,1555,1581]" captionStart-0="FIGURE 32" captionStart-1="FIGURE 33" captionStart-2="FIGURE 34" captionStart-3="FIGURE 35" captionStart-4="FIGURE 36" captionStart-5="FIGURE 37" captionStart-6="FIGURE 38" captionStart-7="FIGURE 39" captionStartId-0="51.[151,250,1943,1967]" captionStartId-1="52.[151,250,1520,1544]" captionStartId-2="54.[151,250,1520,1544]" captionStartId-3="55.[151,250,1943,1967]" captionStartId-4="56.[151,250,1943,1967]" captionStartId-5="57.[151,250,1981,2005]" captionStartId-6="58.[151,250,1943,1967]" captionStartId-7="59.[151,250,1960,1984]" captionTargetBox-0="[179,1408,181,1920]" captionTargetBox-1="[151,1436,181,1496]" captionTargetBox-2="[151,1436,181,1496]" captionTargetBox-3="[186,1401,185,1915]" captionTargetBox-4="[166,1418,185,1915]" captionTargetBox-5="[223,1388,222,1895]" captionTargetBox-6="[189,1403,188,1912]" captionTargetBox-7="[175,1409,202,1932]" captionTargetId-0="figure-17@51.[179,1408,181,1920]" captionTargetId-1="figure-31@52.[151,1436,181,1496]" captionTargetId-2="figure-31@54.[151,1436,181,1496]" captionTargetId-3="figure-17@55.[182,1405,181,1919]" captionTargetId-4="figure-31@56.[166,1422,181,1919]" captionTargetId-5="figure-17@57.[166,1422,181,1957]" captionTargetId-6="figure-31@58.[179,1408,181,1919]" captionTargetId-7="figure-17@59.[175,1413,198,1936]" captionTargetPageId-0="51" captionTargetPageId-1="52" captionTargetPageId-2="54" captionTargetPageId-3="55" captionTargetPageId-4="56" captionTargetPageId-5="57" captionTargetPageId-6="58" captionTargetPageId-7="59" captionText-0="FIGURE 32. Benincandona trapezoides gen. et spec. nov., valves (AC, GK ♂; DF ♀). AF, H. CpRl (AC, H. INV.159276, DF. INV.1592 INV.159279). G, I, J. INV.159279. CpV. K. INV.159277. CpD. Scale bars. 1 (B, E, G, K) = 500 µm; 2 (A, C, D, F, I) = 200 µm, 3 (H, J) = 40 µm." captionText-1="FIGURE 33. Benincandona trapezoides gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159263). B, E, F, H. RVi (INV.159263). G, H. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (GH) = 50 µm." captionText-2="FIGURE 34. Benincandona trapezoides gen. et spec. nov., valves (♀). A, C, D, G. LVi (INV.159264). B, E, F, H. RVi (INV.159264). G, H. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (GH) = 50 µm." captionText-3="FIGURE 35. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. A2 (INV.159280). B. A1 (INV.159265). C. MdCoxa (INV.159263). Scale bar = 50 µm." captionText-4="FIGURE 36. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Mx1, respiratory plate (INV.159263). B. Mx1 (Chaetotaxy incomplete) (INV.159263). C. MdPalp (INV.159263). Scale bar. 1 (A), 2 (C), 3 (B) = 50 µm." captionText-5="FIGURE 37. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. T2 (INV.159263). B. T3 (INV.159263). C. CR (INV.159280). D. CRAtt (INV.159280). Scale bar = 50 µm." captionText-6="FIGURE 38. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159263). BD. Hp (INV.159263). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." captionText-7="FIGURE 39. Benincandona trapezoides gen. et spec. nov., limbs (AB ♀ and CE ♂). A. T1, Lpp (INV.159280). B. T1, Rpp (INV.159280). C. A2 (INV.159264, INV.159269). D. T1 (INV.159268). E. CR (INV.159264). Scale bar = 50 µm." figureDoi-0="http://doi.org/10.5281/zenodo.13745788" figureDoi-1="http://doi.org/10.5281/zenodo.13745790" figureDoi-2="http://doi.org/10.5281/zenodo.13745792" figureDoi-3="http://doi.org/10.5281/zenodo.13745794" figureDoi-4="http://doi.org/10.5281/zenodo.13745796" figureDoi-5="http://doi.org/10.5281/zenodo.13745798" figureDoi-6="http://doi.org/10.5281/zenodo.13745800" figureDoi-7="http://doi.org/10.5281/zenodo.13745802" httpUri-0="https://zenodo.org/record/13745788/files/figure.png" httpUri-1="https://zenodo.org/record/13745790/files/figure.png" httpUri-2="https://zenodo.org/record/13745792/files/figure.png" httpUri-3="https://zenodo.org/record/13745794/files/figure.png" httpUri-4="https://zenodo.org/record/13745796/files/figure.png" httpUri-5="https://zenodo.org/record/13745798/files/figure.png" httpUri-6="https://zenodo.org/record/13745800/files/figure.png" httpUri-7="https://zenodo.org/record/13745802/files/figure.png" pageId="50" pageNumber="51">
Figures 32
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,
<figureCitation id="EA8DAAA94507FF8EFE861D5DFE2C2B18" box="[336,418,1555,1581]" captionStart="FIGURE 41" captionStartId="61.[151,250,1113,1137]" captionTargetBox="[151,1436,181,1089]" captionTargetId="figure-17@61.[151,1436,181,1089]" captionTargetPageId="61" captionText="FIGURE 41. Benincandona gen. et spec. nov., CpRl. (A, C, E, F, G, I, K, N, O, Q ♂ and B, D, H, J, L, M, P, R ♀). AD. B. martini. EF. B. ibikounlei. GJ. B. sakitii. KN. B. cebios. OR. B. trapezoides. Scale bar = 500 µm." pageId="50" pageNumber="51">41OR</figureCitation>
<figureCitation id="EA8DAAA94507FF8EFE861D5DFE2C2B18" box="[336,418,1555,1581]" captionStart="FIGURE 41" captionStartId="61.[151,250,1113,1137]" captionTargetBox="[151,1436,181,1089]" captionTargetId="figure-17@61.[151,1436,181,1089]" captionTargetPageId="61" captionText="FIGURE 41. Benincandona gen. et spec. nov., CpRl. (A, C, E, F, G, I, K, N, O, Q ♂ and B, D, H, J, L, M, P, R ♀). AD. B. martini. EF. B. ibikounlei. GJ. B. sakitii. KN. B. cebios. OR. B. trapezoides. Scale bar = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.13745806" httpUri="https://zenodo.org/record/13745806/files/figure.png" pageId="50" pageNumber="51">41OR</figureCitation>
)
</paragraph>
</subSubSection>
@ -108,7 +109,7 @@ one third of the entire length, with greatest height in the middle; LV dorsally
. 1.5x basal W. T3 with seta d2 absent. Hp with ls and ms with overlapping positions, lobe ls broad and rounded, extending beyond lobe ms with more than half of its length.
</paragraph>
</subSubSection>
<caption id="26C9E6A44506FF8FFF411CD9FD892ACD" pageId="51" pageNumber="52" startId="51.[151,250,1943,1967]" targetBox="[179,1408,181,1920]" targetPageId="51" targetType="figure">
<caption id="26C9E6A44506FF8FFF411CD9FD892ACD" ID-DOI="http://doi.org/10.5281/zenodo.13745788" ID-Zenodo-Dep="13745788" httpUri="https://zenodo.org/record/13745788/files/figure.png" pageId="51" pageNumber="52" startId="51.[151,250,1943,1967]" targetBox="[179,1408,181,1920]" targetPageId="51" targetType="figure">
<paragraph id="7209B62C4506FF8FFF411CD9FD892ACD" blockId="51.[151,1436,1943,2040]" pageId="51" pageNumber="52">
<emphasis id="40C26A3E4506FF8FFF411CD9FE912A9A" bold="true" box="[151,287,1943,1967]" pageId="51" pageNumber="52">FIGURE 32.</emphasis>
<taxonomicName id="B5B6CDAF4506FF8FFEF21CD6FDBF2A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[292,561,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="trapezoides" status="gen. et spec. nov.">
@ -124,7 +125,7 @@ one third of the entire length, with greatest height in the middle; LV dorsally
. INV.159277. CpD. Scale bars. 1 (B, E, G, K) = 500 µm; 2 (A, C, D, F, I) = 200 µm, 3 (H, J) = 40 µm.
</paragraph>
</caption>
<caption id="26C9E6A44501FF88FF411EBEFBFC2B18" pageId="52" pageNumber="53" startId="52.[151,250,1520,1544]" targetBox="[151,1436,181,1496]" targetPageId="52" targetType="figure">
<caption id="26C9E6A44501FF88FF411EBEFBFC2B18" ID-DOI="http://doi.org/10.5281/zenodo.13745790" ID-Zenodo-Dep="13745790" httpUri="https://zenodo.org/record/13745790/files/figure.png" pageId="52" pageNumber="53" startId="52.[151,250,1520,1544]" targetBox="[151,1436,181,1496]" targetPageId="52" targetType="figure">
<paragraph id="7209B62C4501FF88FF411EBEFBFC2B18" blockId="52.[151,1437,1520,1581]" pageId="52" pageNumber="53">
<emphasis id="40C26A3E4501FF88FF411EBEFEAA2B3D" bold="true" box="[151,292,1520,1544]" pageId="52" pageNumber="53">FIGURE 33.</emphasis>
<taxonomicName id="B5B6CDAF4501FF88FEF81EBEFDCE2B3D" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[302,576,1520,1544]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="trapezoides" status="gen. et spec. nov.">
@ -241,39 +242,39 @@ and
</paragraph>
<paragraph id="7209B62C4500FF89FF411AB1FCD82FF8" blockId="53.[151,1437,474,1905]" pageId="53" pageNumber="54">
CpRl (
<figureCitation id="EA8DAAA94500FF89FF361AB1FE0D2F2C" box="[224,387,511,537]" captionStart="FIGURE 32" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure-17@51.[179,1408,181,1920]" captionTargetPageId="51" captionText="FIGURE 32. Benincandona trapezoides gen. et spec. nov., valves (AC, GK ♂; DF ♀). AF, H. CpRl (AC, H. INV.159276, DF. INV.1592 INV.159279). G, I, J. INV.159279. CpV. K. INV.159277. CpD. Scale bars. 1 (B, E, G, K) = 500 µm; 2 (A, C, D, F, I) = 200 µm, 3 (H, J) = 40 µm." pageId="53" pageNumber="54">Figure 32AC</figureCitation>
<figureCitation id="EA8DAAA94500FF89FF361AB1FE0D2F2C" box="[224,387,511,537]" captionStart="FIGURE 32" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure-17@51.[179,1408,181,1920]" captionTargetPageId="51" captionText="FIGURE 32. Benincandona trapezoides gen. et spec. nov., valves (AC, GK ♂; DF ♀). AF, H. CpRl (AC, H. INV.159276, DF. INV.1592 INV.159279). G, I, J. INV.159279. CpV. K. INV.159277. CpD. Scale bars. 1 (B, E, G, K) = 500 µm; 2 (A, C, D, F, I) = 200 µm, 3 (H, J) = 40 µm." figureDoi="http://doi.org/10.5281/zenodo.13745788" httpUri="https://zenodo.org/record/13745788/files/figure.png" pageId="53" pageNumber="54">Figure 32AC</figureCitation>
) with trapezoidal shape (L/H ratio=2.0); dorsal margin straight in the middle 1/3, with greatest height in the middle, anterior margin broadly rounded, posterior margin running straight towards ventral margin, the latter almost straight; LV overlapping RV along anterior, ventral and posterior sides, dorsally strongly overlapping RV along the entire length of the straight part; valve surface set with ridges in the middle part of the valves (
<figureCitation id="EA8DAAA94500FF89FA871925FF6B2F9C" captionStart="FIGURE 32" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure-17@51.[179,1408,181,1920]" captionTargetPageId="51" captionText="FIGURE 32. Benincandona trapezoides gen. et spec. nov., valves (AC, GK ♂; DF ♀). AF, H. CpRl (AC, H. INV.159276, DF. INV.1592 INV.159279). G, I, J. INV.159279. CpV. K. INV.159277. CpD. Scale bars. 1 (B, E, G, K) = 500 µm; 2 (A, C, D, F, I) = 200 µm, 3 (H, J) = 40 µm." pageId="53" pageNumber="54">Figure 32HJ</figureCitation>
<figureCitation id="EA8DAAA94500FF89FA871925FF6B2F9C" captionStart="FIGURE 32" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure-17@51.[179,1408,181,1920]" captionTargetPageId="51" captionText="FIGURE 32. Benincandona trapezoides gen. et spec. nov., valves (AC, GK ♂; DF ♀). AF, H. CpRl (AC, H. INV.159276, DF. INV.1592 INV.159279). G, I, J. INV.159279. CpV. K. INV.159277. CpD. Scale bars. 1 (B, E, G, K) = 500 µm; 2 (A, C, D, F, I) = 200 µm, 3 (H, J) = 40 µm." figureDoi="http://doi.org/10.5281/zenodo.13745788" httpUri="https://zenodo.org/record/13745788/files/figure.png" pageId="53" pageNumber="54">Figure 32HJ</figureCitation>
). CpD and CpV (
<figureCitation id="EA8DAAA94500FF89FE7019C1FDC32F9C" box="[422,589,655,681]" captionStart="FIGURE 32" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure-17@51.[179,1408,181,1920]" captionTargetPageId="51" captionText="FIGURE 32. Benincandona trapezoides gen. et spec. nov., valves (AC, GK ♂; DF ♀). AF, H. CpRl (AC, H. INV.159276, DF. INV.1592 INV.159279). G, I, J. INV.159279. CpV. K. INV.159277. CpD. Scale bars. 1 (B, E, G, K) = 500 µm; 2 (A, C, D, F, I) = 200 µm, 3 (H, J) = 40 µm." pageId="53" pageNumber="54">Figure 32G, K</figureCitation>
<figureCitation id="EA8DAAA94500FF89FE7019C1FDC32F9C" box="[422,589,655,681]" captionStart="FIGURE 32" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure-17@51.[179,1408,181,1920]" captionTargetPageId="51" captionText="FIGURE 32. Benincandona trapezoides gen. et spec. nov., valves (AC, GK ♂; DF ♀). AF, H. CpRl (AC, H. INV.159276, DF. INV.1592 INV.159279). G, I, J. INV.159279. CpV. K. INV.159277. CpD. Scale bars. 1 (B, E, G, K) = 500 µm; 2 (A, C, D, F, I) = 200 µm, 3 (H, J) = 40 µm." figureDoi="http://doi.org/10.5281/zenodo.13745788" httpUri="https://zenodo.org/record/13745788/files/figure.png" pageId="53" pageNumber="54">Figure 32G, K</figureCitation>
) relatively wide, pointed sharply anteriorly and bluntly posteriorly, greatest width in the middle, lateral margins straight in the middle part.
</paragraph>
<paragraph id="7209B62C4500FF89FF111999FB412E0C" blockId="53.[151,1437,474,1905]" pageId="53" pageNumber="54">
LVi (
<figureCitation id="EA8DAAA94500FF89FF281999FE6F2FC4" box="[254,481,727,753]" captionStart="FIGURE 33" captionStartId="52.[151,250,1520,1544]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-31@52.[151,1436,181,1496]" captionTargetPageId="52" captionText="FIGURE 33. Benincandona trapezoides gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159263). B, E, F, H. RVi (INV.159263). G, H. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (GH) = 50 µm." pageId="53" pageNumber="54">Figure 33A, C, D, G</figureCitation>
<figureCitation id="EA8DAAA94500FF89FF281999FE6F2FC4" box="[254,481,727,753]" captionStart="FIGURE 33" captionStartId="52.[151,250,1520,1544]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-31@52.[151,1436,181,1496]" captionTargetPageId="52" captionText="FIGURE 33. Benincandona trapezoides gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159263). B, E, F, H. RVi (INV.159263). G, H. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (GH) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745790" httpUri="https://zenodo.org/record/13745790/files/figure.png" pageId="53" pageNumber="54">Figure 33A, C, D, G</figureCitation>
) elongated and sub-triangular, dorsal margin straight in the middle third, with greatest height in the middle, anterior margin broadly rounded; posterior margin sloping towards ventral margin, the latter almost straight; ventral il without antero-ventral tooth. CMS (
<figureCitation id="EA8DAAA94500FF89FC9F1851FC452E0C" box="[841,971,799,825]" captionStart="FIGURE 33" captionStartId="52.[151,250,1520,1544]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-31@52.[151,1436,181,1496]" captionTargetPageId="52" captionText="FIGURE 33. Benincandona trapezoides gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159263). B, E, F, H. RVi (INV.159263). G, H. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (GH) = 50 µm." pageId="53" pageNumber="54">Figure 33G</figureCitation>
<figureCitation id="EA8DAAA94500FF89FC9F1851FC452E0C" box="[841,971,799,825]" captionStart="FIGURE 33" captionStartId="52.[151,250,1520,1544]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-31@52.[151,1436,181,1496]" captionTargetPageId="52" captionText="FIGURE 33. Benincandona trapezoides gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159263). B, E, F, H. RVi (INV.159263). G, H. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (GH) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745790" httpUri="https://zenodo.org/record/13745790/files/figure.png" pageId="53" pageNumber="54">Figure 33G</figureCitation>
) as in the
<typeStatus id="AD0D088E4500FF89FB961851FBFE2E0C" box="[1088,1136,799,825]" pageId="53" pageNumber="54">type</typeStatus>
species.
</paragraph>
<paragraph id="7209B62C4500FF89FF11180DFBF02E90" blockId="53.[151,1437,474,1905]" pageId="53" pageNumber="54">
RVi (
<figureCitation id="EA8DAAA94500FF89FED4180DFE682E68" box="[258,486,835,861]" captionStart="FIGURE 33" captionStartId="52.[151,250,1520,1544]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-31@52.[151,1436,181,1496]" captionTargetPageId="52" captionText="FIGURE 33. Benincandona trapezoides gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159263). B, E, F, H. RVi (INV.159263). G, H. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (GH) = 50 µm." pageId="53" pageNumber="54">Figure 33B, E, F, H</figureCitation>
<figureCitation id="EA8DAAA94500FF89FED4180DFE682E68" box="[258,486,835,861]" captionStart="FIGURE 33" captionStartId="52.[151,250,1520,1544]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-31@52.[151,1436,181,1496]" captionTargetPageId="52" captionText="FIGURE 33. Benincandona trapezoides gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159263). B, E, F, H. RVi (INV.159263). G, H. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (GH) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745790" httpUri="https://zenodo.org/record/13745790/files/figure.png" pageId="53" pageNumber="54">Figure 33B, E, F, H</figureCitation>
) elongated, with dorsal margin straight in the middle third and with greatest height in the middle, anterior margin broadly rounded; posterior margin sloping towards ventral margin, the latter almost straight; ventral il without antero-ventral tooth. CMS (
<figureCitation id="EA8DAAA94500FF89FD2118C5FCF42E90" box="[759,890,907,933]" captionStart="FIGURE 33" captionStartId="52.[151,250,1520,1544]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-31@52.[151,1436,181,1496]" captionTargetPageId="52" captionText="FIGURE 33. Benincandona trapezoides gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159263). B, E, F, H. RVi (INV.159263). G, H. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (GH) = 50 µm." pageId="53" pageNumber="54">Figure 33H</figureCitation>
<figureCitation id="EA8DAAA94500FF89FD2118C5FCF42E90" box="[759,890,907,933]" captionStart="FIGURE 33" captionStartId="52.[151,250,1520,1544]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-31@52.[151,1436,181,1496]" captionTargetPageId="52" captionText="FIGURE 33. Benincandona trapezoides gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159263). B, E, F, H. RVi (INV.159263). G, H. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (GH) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745790" httpUri="https://zenodo.org/record/13745790/files/figure.png" pageId="53" pageNumber="54">Figure 33H</figureCitation>
) as in the
<typeStatus id="AD0D088E4500FF89FC3818C5FB902E90" box="[1006,1054,907,933]" pageId="53" pageNumber="54">type</typeStatus>
species.
</paragraph>
<paragraph id="7209B62C4500FF89FF1118E1FB0A2838" blockId="53.[151,1437,474,1905]" pageId="53" pageNumber="54">
A1 (
<figureCitation id="EA8DAAA94500FF89FF2D18E1FEF12EFC" box="[251,383,942,969]" captionStart="FIGURE 35" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[186,1401,185,1915]" captionTargetId="figure-17@55.[182,1405,181,1919]" captionTargetPageId="55" captionText="FIGURE 35. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. A2 (INV.159280). B. A1 (INV.159265). C. MdCoxa (INV.159263). Scale bar = 50 µm." pageId="53" pageNumber="54">Figure 35B</figureCitation>
<figureCitation id="EA8DAAA94500FF89FF2D18E1FEF12EFC" box="[251,383,942,969]" captionStart="FIGURE 35" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[186,1401,185,1915]" captionTargetId="figure-17@55.[182,1405,181,1919]" captionTargetPageId="55" captionText="FIGURE 35. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. A2 (INV.159280). B. A1 (INV.159265). C. MdCoxa (INV.159263). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745794" httpUri="https://zenodo.org/record/13745794/files/figure.png" pageId="53" pageNumber="54">Figure 35B</figureCitation>
) with seven segments, one basal and six distal. Basal segment with two sub-apical, ventral setae, one long and one shorter, the latter about half of the length of the longer one, and two shorter setae on the mid-dorsal side (one not illustrated here), situated some distance from each other. First distal segment about 2.5 times as long as the basal width, with one short dorso-subapical seta. Second distal segment with about the same length/width ratio as the previous segment, set with one short dorso-apical seta. Third distal segment also with about the same length/width ratio, carrying one short apical setae on each side. Fourth distal segment about three times as long as wide, and set with one short ventro-apical seta and two long dorso-apical. Fifth distal segment about five time as long as wide, with two long dorso-apical setae and one short ventro-apical seta clearly exceeding the next segment. Sixth distal segment narrow, at least five times as long as wide, with four apical structures: one aesthetasc Ya of intermediate length, two long and one short setae (about one third the length of Ya).
</paragraph>
<paragraph id="7209B62C4500FF89FF111E59FA9A2A1C" blockId="53.[151,1437,474,1905]" pageId="53" pageNumber="54">
A2 (
<figureCitation id="EA8DAAA94500FF89FF2D1E59FE0D2804" box="[251,387,1302,1329]" captionStart="FIGURE 35" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[186,1401,185,1915]" captionTargetId="figure-17@55.[182,1405,181,1919]" captionTargetPageId="55" captionText="FIGURE 35. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. A2 (INV.159280). B. A1 (INV.159265). C. MdCoxa (INV.159263). Scale bar = 50 µm." pageId="53" pageNumber="54">Figure 35A</figureCitation>
<figureCitation id="EA8DAAA94500FF89FF2D1E59FE0D2804" box="[251,387,1302,1329]" captionStart="FIGURE 35" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[186,1401,185,1915]" captionTargetId="figure-17@55.[182,1405,181,1919]" captionTargetPageId="55" captionText="FIGURE 35. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. A2 (INV.159280). B. A1 (INV.159265). C. MdCoxa (INV.159263). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745794" httpUri="https://zenodo.org/record/13745794/files/figure.png" pageId="53" pageNumber="54">Figure 35A</figureCitation>
) with basal segment (first protopodite) broad and rounded, mid-ventrally with one seta of medium length, subapically with two setae of unequal length, proximal one
<emphasis id="40C26A3E4500FF89FBD41E75FB9B2861" box="[1026,1045,1339,1364]" italics="true" pageId="53" pageNumber="54">c.</emphasis>
four times the length of the more apical one. Second segment (second protopodite) large and
@ -281,15 +282,15 @@ four times the length of the more apical one. Second segment (second protopodite
twice as long as basal width, set with one long ventro-subapical seta X. No remnant of exopodite visible. Endopodite consisting of four segments (penultimate segment divided). First endopodal segment large and stout, five times as long as basal width, carrying one long and narrow mid-ventral aesthetasc Y, and two ventro-apical setae, one short and one large and stout, hirsute in its distal 4/5. Second endopodal segment (segment 2A)
<emphasis id="40C26A3E4500FF89FDAD1EA1FD002B3D" box="[635,654,1519,1544]" italics="true" pageId="53" pageNumber="54">c.</emphasis>
twice as long as wide, carrying one stout dorso-apical plumose seta and three t-setae; seta t1 long, narrow and plumose, accompanied by a slender aesthetasc y1; t2 and t3 setae hyper-developed into male bristles, t2 ventro-apical, t3 centro-apical. Third endopodal segment (segment 2B) smaller than previous segment and about 1.5x as long as wide, set with a complex (and sexually dimorphic) apical chaetotaxy (
<figureCitation id="EA8DAAA94500FF89FF491D31FE9F2BAC" box="[159,273,1663,1689]" captionStart="FIGURE 40" captionStartId="60.[151,250,1516,1540]" captionTargetBox="[151,1435,181,1492]" captionTargetId="figure-31@60.[151,1435,181,1492]" captionTargetPageId="60" captionText="FIGURE 40. Diagrams of apical views of the A2 of Benincandona, showing position of (sub-) terminal claws, setae and aesthetascs of the terminal (small circle) and penultimate segment (large circle). For explanations of codes, see Material and Methods (Abbreviations used in text and figures)." pageId="53" pageNumber="54">Figure 40</figureCitation>
<figureCitation id="EA8DAAA94500FF89FF491D31FE9F2BAC" box="[159,273,1663,1689]" captionStart="FIGURE 40" captionStartId="60.[151,250,1516,1540]" captionTargetBox="[151,1435,181,1492]" captionTargetId="figure-31@60.[151,1435,181,1492]" captionTargetPageId="60" captionText="FIGURE 40. Diagrams of apical views of the A2 of Benincandona, showing position of (sub-) terminal claws, setae and aesthetascs of the terminal (small circle) and penultimate segment (large circle). For explanations of codes, see Material and Methods (Abbreviations used in text and figures)." figureDoi="http://doi.org/10.5281/zenodo.13745804" httpUri="https://zenodo.org/record/13745804/files/figure.png" pageId="53" pageNumber="54">Figure 40</figureCitation>
): ventro-apically with one slender aesthetasc y2 and claw G3 transformed into one short seta about equal length as y2; centrally with one short plumose seta z3; dorso-apically, z1 claw-like about 1.5x as long as penultimate segment. Claw G2 and G1 about 2.5 times as long as z1; seta z2 transformed into long claw, reaching tip of G1. Fourth, terminal, endopodal segment with one long claw (Gm), one short claw (GM), a long aesthetasc y3 with shorter accompanying seta, with fused part short, and a seta g (slightly exceeding tip of aesthetasc y3).
</paragraph>
<paragraph id="7209B62C4500FF89FF111C7DFAA52A44" blockId="53.[151,1437,474,1905]" pageId="53" pageNumber="54">
MdCoxa (
<figureCitation id="EA8DAAA94500FF89FEED1C7DFE4E2A78" box="[315,448,1842,1869]" captionStart="FIGURE 35" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[186,1401,185,1915]" captionTargetId="figure-17@55.[182,1405,181,1919]" captionTargetPageId="55" captionText="FIGURE 35. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. A2 (INV.159280). B. A1 (INV.159265). C. MdCoxa (INV.159263). Scale bar = 50 µm." pageId="53" pageNumber="54">Figure 35C</figureCitation>
<figureCitation id="EA8DAAA94500FF89FEED1C7DFE4E2A78" box="[315,448,1842,1869]" captionStart="FIGURE 35" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[186,1401,185,1915]" captionTargetId="figure-17@55.[182,1405,181,1919]" captionTargetPageId="55" captionText="FIGURE 35. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. A2 (INV.159280). B. A1 (INV.159265). C. MdCoxa (INV.159263). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745794" httpUri="https://zenodo.org/record/13745794/files/figure.png" pageId="53" pageNumber="54">Figure 35C</figureCitation>
) elongated, proximally sharply pointed, medially widened, distally with an obliquely widened end, set with a series (67) of strong teeth, interspaced with setae; subapically with a short seta.
</paragraph>
<caption id="26C9E6A44503FF8AFF411EBEFBFC2B18" pageId="54" pageNumber="55" startId="54.[151,250,1520,1544]" targetBox="[151,1436,181,1496]" targetPageId="54" targetType="figure">
<caption id="26C9E6A44503FF8AFF411EBEFBFC2B18" ID-DOI="http://doi.org/10.5281/zenodo.13745792" ID-Zenodo-Dep="13745792" httpUri="https://zenodo.org/record/13745792/files/figure.png" pageId="54" pageNumber="55" startId="54.[151,250,1520,1544]" targetBox="[151,1436,181,1496]" targetPageId="54" targetType="figure">
<paragraph id="7209B62C4503FF8AFF411EBEFBFC2B18" blockId="54.[151,1437,1520,1581]" pageId="54" pageNumber="55">
<emphasis id="40C26A3E4503FF8AFF411EBEFEAA2B3D" bold="true" box="[151,292,1520,1544]" pageId="54" pageNumber="55">FIGURE 34.</emphasis>
<taxonomicName id="B5B6CDAF4503FF8AFEF81EBEFDCE2B3D" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[302,576,1520,1544]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="trapezoides" status="gen. et spec. nov.">
@ -307,10 +308,10 @@ CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (GH) = 50 µm.
</caption>
<paragraph id="7209B62C4503FF8AFF111D30FEF82A8D" blockId="54.[151,1437,1662,1976]" pageId="54" pageNumber="55">
MdPalp (
<figureCitation id="EA8DAAA94503FF8AFEF91D30FE3C2BAD" box="[303,434,1662,1688]" captionStart="FIGURE 36" captionStartId="56.[151,250,1943,1967]" captionTargetBox="[166,1418,185,1915]" captionTargetId="figure-31@56.[166,1422,181,1919]" captionTargetPageId="56" captionText="FIGURE 36. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Mx1, respiratory plate (INV.159263). B. Mx1 (Chaetotaxy incomplete) (INV.159263). C. MdPalp (INV.159263). Scale bar. 1 (A), 2 (C), 3 (B) = 50 µm." pageId="54" pageNumber="55">Figure 36C</figureCitation>
<figureCitation id="EA8DAAA94503FF8AFEF91D30FE3C2BAD" box="[303,434,1662,1688]" captionStart="FIGURE 36" captionStartId="56.[151,250,1943,1967]" captionTargetBox="[166,1418,185,1915]" captionTargetId="figure-31@56.[166,1422,181,1919]" captionTargetPageId="56" captionText="FIGURE 36. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Mx1, respiratory plate (INV.159263). B. Mx1 (Chaetotaxy incomplete) (INV.159263). C. MdPalp (INV.159263). Scale bar. 1 (A), 2 (C), 3 (B) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745796" httpUri="https://zenodo.org/record/13745796/files/figure.png" pageId="54" pageNumber="55">Figure 36C</figureCitation>
) with four segments. Basal segment ventro-subapically carrying two hirsute setae (S1 &amp; S2), S1 long, reaching tip of terminal segment and S2 short and stout, length about one fourth of the length of S1; in between with the short flagellum-like alpha seta (thin and as short as S2), and one smooth and medium size (slightly shorter than S1) more proximal seta. Second segment of sub-triangular shape, dorso-apically with two setae of unequal length; ventro-apically with a group of four long setae, one of these being the beta-seta. Third segment with two unequal dorso-subapical setae, two mid-dorsal setae, one of which being the gamma seta, and ventro-apically with two setae of sub-equal length. Terminal segment sub-rectangular, apically set with one large claw, fused with the segment over its entire distal margin, accompanied by one smooth lateral seta on each side, tips slightly shorter than the fused claw.
</paragraph>
<caption id="26C9E6A44502FF8BFF411CD9FE7C2AE1" pageId="55" pageNumber="56" startId="55.[151,250,1943,1967]" targetBox="[186,1401,185,1915]" targetPageId="55" targetType="figure">
<caption id="26C9E6A44502FF8BFF411CD9FE7C2AE1" ID-DOI="http://doi.org/10.5281/zenodo.13745794" ID-Zenodo-Dep="13745794" httpUri="https://zenodo.org/record/13745794/files/figure.png" pageId="55" pageNumber="56" startId="55.[151,250,1943,1967]" targetBox="[186,1401,185,1915]" targetPageId="55" targetType="figure">
<paragraph id="7209B62C4502FF8BFF411CD9FE7C2AE1" blockId="55.[151,1437,1943,2004]" pageId="55" pageNumber="56">
<emphasis id="40C26A3E4502FF8BFF411CD9FEAE2A9A" bold="true" box="[151,288,1943,1967]" pageId="55" pageNumber="56">FIGURE 35.</emphasis>
<taxonomicName id="B5B6CDAF4502FF8BFEF11CD6FDBB2A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[295,565,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="trapezoides" status="gen. et spec. nov.">
@ -326,7 +327,7 @@ MdPalp (
MdCoxa (INV.159263). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A4450DFF84FF411CD9FB492AE1" pageId="56" pageNumber="57" startId="56.[151,250,1943,1967]" targetBox="[166,1418,185,1915]" targetPageId="56" targetType="figure">
<caption id="26C9E6A4450DFF84FF411CD9FB492AE1" ID-DOI="http://doi.org/10.5281/zenodo.13745796" ID-Zenodo-Dep="13745796" httpUri="https://zenodo.org/record/13745796/files/figure.png" pageId="56" pageNumber="57" startId="56.[151,250,1943,1967]" targetBox="[166,1418,185,1915]" targetPageId="56" targetType="figure">
<paragraph id="7209B62C450DFF84FF411CD9FB492AE1" blockId="56.[151,1436,1943,2004]" pageId="56" pageNumber="57">
<emphasis id="40C26A3E450DFF84FF411CD9FEAB2A9A" bold="true" box="[151,293,1943,1967]" pageId="56" pageNumber="57">FIGURE 36.</emphasis>
<taxonomicName id="B5B6CDAF450DFF84FEF91CD6FDCC2A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[303,578,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="trapezoides" status="gen. et spec. nov.">
@ -342,7 +343,7 @@ MdCoxa (INV.159263). Scale bar = 50 µm.
. MdPalp (INV.159263). Scale bar. 1 (A), 2 (C), 3 (B) = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A4450CFF85FF411CF3FD7D2ACF" pageId="57" pageNumber="58" startId="57.[151,250,1981,2005]" targetBox="[223,1388,222,1895]" targetPageId="57" targetType="figure">
<caption id="26C9E6A4450CFF85FF411CF3FD7D2ACF" ID-DOI="http://doi.org/10.5281/zenodo.13745798" ID-Zenodo-Dep="13745798" httpUri="https://zenodo.org/record/13745798/files/figure.png" pageId="57" pageNumber="58" startId="57.[151,250,1981,2005]" targetBox="[223,1388,222,1895]" targetPageId="57" targetType="figure">
<paragraph id="7209B62C450CFF85FF411CF3FD7D2ACF" blockId="57.[151,1436,1981,2042]" pageId="57" pageNumber="58">
<emphasis id="40C26A3E450CFF85FF411CF3FEAA2AE0" bold="true" box="[151,292,1981,2005]" pageId="57" pageNumber="58">FIGURE 37.</emphasis>
<taxonomicName id="B5B6CDAF450CFF85FEF81CF3FDCE2AE0" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[302,576,1981,2005]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="trapezoides" status="gen. et spec. nov.">
@ -360,7 +361,7 @@ MdCoxa (INV.159263). Scale bar = 50 µm.
. CRAtt (INV.159280). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A4450FFF86FF411CD9FDB32ACD" pageId="58" pageNumber="59" startId="58.[151,250,1943,1967]" targetBox="[189,1403,188,1912]" targetPageId="58" targetType="figure">
<caption id="26C9E6A4450FFF86FF411CD9FDB32ACD" ID-DOI="http://doi.org/10.5281/zenodo.13745800" ID-Zenodo-Dep="13745800" httpUri="https://zenodo.org/record/13745800/files/figure.png" pageId="58" pageNumber="59" startId="58.[151,250,1943,1967]" targetBox="[189,1403,188,1912]" targetPageId="58" targetType="figure">
<paragraph id="7209B62C450FFF86FF411CD9FDB32ACD" blockId="58.[151,1437,1943,2040]" pageId="58" pageNumber="59">
<emphasis id="40C26A3E450FFF86FF411CD9FEAE2A9A" bold="true" box="[151,288,1943,1967]" pageId="58" pageNumber="59">FIGURE 38.</emphasis>
<taxonomicName id="B5B6CDAF450FFF86FEF01CD9FDBD2A9A" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[294,563,1943,1967]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="trapezoides" status="gen. et spec. nov.">
@ -380,7 +381,7 @@ MdCoxa (INV.159263). Scale bar = 50 µm.
. Internal view (photograph with interference contrast). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A4450EFF87FF411CE6FA9F2AD0" pageId="59" pageNumber="60" startId="59.[151,250,1960,1984]" targetBox="[175,1409,202,1932]" targetPageId="59" targetType="figure">
<caption id="26C9E6A4450EFF87FF411CE6FA9F2AD0" ID-DOI="http://doi.org/10.5281/zenodo.13745802" ID-Zenodo-Dep="13745802" httpUri="https://zenodo.org/record/13745802/files/figure.png" pageId="59" pageNumber="60" startId="59.[151,250,1960,1984]" targetBox="[175,1409,202,1932]" targetPageId="59" targetType="figure">
<paragraph id="7209B62C450EFF87FF411CE6FA9F2AD0" blockId="59.[151,1436,1960,2021]" pageId="59" pageNumber="60">
<emphasis id="40C26A3E450EFF87FF411CE6FEAE2AF4" bold="true" box="[151,288,1960,1985]" pageId="59" pageNumber="60">FIGURE 39.</emphasis>
<taxonomicName id="B5B6CDAF450EFF87FEF11CE7FDBB2AF4" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[295,565,1961,1985]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="trapezoides" status="gen. et spec. nov.">
@ -400,7 +401,7 @@ MdCoxa (INV.159263). Scale bar = 50 µm.
. CR (INV.159264). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A44509FF80FF411EA2FD1F2B79" pageId="60" pageNumber="61" startId="60.[151,250,1516,1540]" targetBox="[151,1435,181,1492]" targetPageId="60" targetType="figure">
<caption id="26C9E6A44509FF80FF411EA2FD1F2B79" ID-DOI="http://doi.org/10.5281/zenodo.13745804" ID-Zenodo-Dep="13745804" httpUri="https://zenodo.org/record/13745804/files/figure.png" pageId="60" pageNumber="61" startId="60.[151,250,1516,1540]" targetBox="[151,1435,181,1492]" targetPageId="60" targetType="figure">
<paragraph id="7209B62C4509FF80FF411EA2FD1F2B79" blockId="60.[151,1437,1516,1612]" pageId="60" pageNumber="61">
<emphasis id="40C26A3E4509FF80FF411EA2FEAD2B31" bold="true" box="[151,291,1516,1540]" pageId="60" pageNumber="61">FIGURE 40.</emphasis>
Diagrams of apical views of the A2 of
@ -412,20 +413,20 @@ Diagrams of apical views of the A2 of
</caption>
<paragraph id="7209B62C4509FF80FF111DD0FCE42AA4" blockId="60.[151,1437,1694,1937]" pageId="60" pageNumber="61">
Mx1 (
<figureCitation id="EA8DAAA94509FF80FED91DD0FE322B8D" box="[271,444,1694,1720]" captionStart="FIGURE 36" captionStartId="56.[151,250,1943,1967]" captionTargetBox="[166,1418,185,1915]" captionTargetId="figure-31@56.[166,1422,181,1919]" captionTargetPageId="56" captionText="FIGURE 36. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Mx1, respiratory plate (INV.159263). B. Mx1 (Chaetotaxy incomplete) (INV.159263). C. MdPalp (INV.159263). Scale bar. 1 (A), 2 (C), 3 (B) = 50 µm." pageId="60" pageNumber="61">Figure 36A, B</figureCitation>
<figureCitation id="EA8DAAA94509FF80FED91DD0FE322B8D" box="[271,444,1694,1720]" captionStart="FIGURE 36" captionStartId="56.[151,250,1943,1967]" captionTargetBox="[166,1418,185,1915]" captionTargetId="figure-31@56.[166,1422,181,1919]" captionTargetPageId="56" captionText="FIGURE 36. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Mx1, respiratory plate (INV.159263). B. Mx1 (Chaetotaxy incomplete) (INV.159263). C. MdPalp (INV.159263). Scale bar. 1 (A), 2 (C), 3 (B) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745796" httpUri="https://zenodo.org/record/13745796/files/figure.png" pageId="60" pageNumber="61">Figure 36A, B</figureCitation>
) with a basal (protopodite) part carrying a large respiratory plate (exopodite), three endites and a two-segmented palp (endopodite). Respiratory plate (
<figureCitation id="EA8DAAA94509FF80FC5C1D8CFB802BE9" box="[906,1038,1730,1756]" captionStart="FIGURE 36" captionStartId="56.[151,250,1943,1967]" captionTargetBox="[166,1418,185,1915]" captionTargetId="figure-31@56.[166,1422,181,1919]" captionTargetPageId="56" captionText="FIGURE 36. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Mx1, respiratory plate (INV.159263). B. Mx1 (Chaetotaxy incomplete) (INV.159263). C. MdPalp (INV.159263). Scale bar. 1 (A), 2 (C), 3 (B) = 50 µm." pageId="60" pageNumber="61">Figure 36A</figureCitation>
<figureCitation id="EA8DAAA94509FF80FC5C1D8CFB802BE9" box="[906,1038,1730,1756]" captionStart="FIGURE 36" captionStartId="56.[151,250,1943,1967]" captionTargetBox="[166,1418,185,1915]" captionTargetId="figure-31@56.[166,1422,181,1919]" captionTargetPageId="56" captionText="FIGURE 36. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Mx1, respiratory plate (INV.159263). B. Mx1 (Chaetotaxy incomplete) (INV.159263). C. MdPalp (INV.159263). Scale bar. 1 (A), 2 (C), 3 (B) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745796" httpUri="https://zenodo.org/record/13745796/files/figure.png" pageId="60" pageNumber="61">Figure 36A</figureCitation>
) elongated, carrying
<emphasis id="40C26A3E4509FF80FB291D8DFA9C2BE9" box="[1279,1298,1731,1756]" italics="true" pageId="60" pageNumber="61">c.</emphasis>
18 plumose respiratory rays, two short distal setae and a group of shorter proximal setae (not illustrated). Palp (
<figureCitation id="EA8DAAA94509FF80FAC61DA8FA1A2A35" box="[1296,1428,1766,1792]" captionStart="FIGURE 36" captionStartId="56.[151,250,1943,1967]" captionTargetBox="[166,1418,185,1915]" captionTargetId="figure-31@56.[166,1422,181,1919]" captionTargetPageId="56" captionText="FIGURE 36. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Mx1, respiratory plate (INV.159263). B. Mx1 (Chaetotaxy incomplete) (INV.159263). C. MdPalp (INV.159263). Scale bar. 1 (A), 2 (C), 3 (B) = 50 µm." pageId="60" pageNumber="61">Figure 36B</figureCitation>
<figureCitation id="EA8DAAA94509FF80FAC61DA8FA1A2A35" box="[1296,1428,1766,1792]" captionStart="FIGURE 36" captionStartId="56.[151,250,1943,1967]" captionTargetBox="[166,1418,185,1915]" captionTargetId="figure-31@56.[166,1422,181,1919]" captionTargetPageId="56" captionText="FIGURE 36. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Mx1, respiratory plate (INV.159263). B. Mx1 (Chaetotaxy incomplete) (INV.159263). C. MdPalp (INV.159263). Scale bar. 1 (A), 2 (C), 3 (B) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745796" httpUri="https://zenodo.org/record/13745796/files/figure.png" pageId="60" pageNumber="61">Figure 36B</figureCitation>
) with first segment carrying four long dorso-apical setae; terminal segment short and rectangular, carrying four distal setae, some slightly claw-like; chaetotaxy of three endites incompletely illustrated: third endite with two strong teeth carrying lateral spines (
<emphasis id="40C26A3E4509FF80FE0F1C1CFDE92A59" box="[473,615,1874,1900]" italics="true" pageId="60" pageNumber="61">Zahnborsten</emphasis>
), basally with one lateral seta; sideways directed bristles near base of first endite, one long and one short,
<emphasis id="40C26A3E4509FF80FE211C39FD842AA5" box="[503,522,1911,1936]" italics="true" pageId="60" pageNumber="61">c.</emphasis>
half the length of the long one.
</paragraph>
<caption id="26C9E6A44508FF81FF411F17FB2529A3" pageId="61" pageNumber="62" startId="61.[151,250,1113,1137]" targetBox="[151,1436,181,1089]" targetPageId="61" targetType="figure">
<caption id="26C9E6A44508FF81FF411F17FB2529A3" ID-DOI="http://doi.org/10.5281/zenodo.13745806" ID-Zenodo-Dep="13745806" httpUri="https://zenodo.org/record/13745806/files/figure.png" pageId="61" pageNumber="62" startId="61.[151,250,1113,1137]" targetBox="[151,1436,181,1089]" targetPageId="61" targetType="figure">
<paragraph id="7209B62C4508FF81FF411F17FB2529A3" blockId="61.[151,1437,1113,1174]" pageId="61" pageNumber="62">
<emphasis id="40C26A3E4508FF81FF411F17FEAF2944" bold="true" box="[151,289,1113,1137]" pageId="61" pageNumber="62">FIGURE 41.</emphasis>
<taxonomicName id="B5B6CDAF4508FF81FEFE1F17FE342944" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[296,442,1113,1137]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="61" pageNumber="62" phylum="Arthropoda" rank="genus" status="gen. et spec. nov.">
@ -468,47 +469,47 @@ Scale bar = 500 µm.
</caption>
<paragraph id="7209B62C4508FF81FF111FA6FDA328A7" blockId="61.[151,1437,1256,2038]" pageId="61" pageNumber="62">
T1 (
<figureCitation id="EA8DAAA94508FF81FF201FA6FE152837" box="[246,411,1256,1282]" captionStart="FIGURE 39" captionStartId="59.[151,250,1960,1984]" captionTargetBox="[175,1409,202,1932]" captionTargetId="figure-17@59.[175,1413,198,1936]" captionTargetPageId="59" captionText="FIGURE 39. Benincandona trapezoides gen. et spec. nov., limbs (AB ♀ and CE ♂). A. T1, Lpp (INV.159280). B. T1, Rpp (INV.159280). C. A2 (INV.159264, INV.159269). D. T1 (INV.159268). E. CR (INV.159264). Scale bar = 50 µm." pageId="61" pageNumber="62">Figure 39A, B</figureCitation>
<figureCitation id="EA8DAAA94508FF81FF201FA6FE152837" box="[246,411,1256,1282]" captionStart="FIGURE 39" captionStartId="59.[151,250,1960,1984]" captionTargetBox="[175,1409,202,1932]" captionTargetId="figure-17@59.[175,1413,198,1936]" captionTargetPageId="59" captionText="FIGURE 39. Benincandona trapezoides gen. et spec. nov., limbs (AB ♀ and CE ♂). A. T1, Lpp (INV.159280). B. T1, Rpp (INV.159280). C. A2 (INV.159264, INV.159269). D. T1 (INV.159268). E. CR (INV.159264). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745802" httpUri="https://zenodo.org/record/13745802/files/figure.png" pageId="61" pageNumber="62">Figure 39A, B</figureCitation>
) consisting of basal part (protopodite), carrying respiratory plates (exopodites not shown). Protopodite (
<figureCitation id="EA8DAAA94508FF81FEFB1E42FE3C2813" box="[301,434,1292,1318]" captionStart="FIGURE 39" captionStartId="59.[151,250,1960,1984]" captionTargetBox="[175,1409,202,1932]" captionTargetId="figure-17@59.[175,1413,198,1936]" captionTargetPageId="59" captionText="FIGURE 39. Benincandona trapezoides gen. et spec. nov., limbs (AB ♀ and CE ♂). A. T1, Lpp (INV.159280). B. T1, Rpp (INV.159280). C. A2 (INV.159264, INV.159269). D. T1 (INV.159268). E. CR (INV.159264). Scale bar = 50 µm." pageId="61" pageNumber="62">Figure 39B</figureCitation>
<figureCitation id="EA8DAAA94508FF81FEFB1E42FE3C2813" box="[301,434,1292,1318]" captionStart="FIGURE 39" captionStartId="59.[151,250,1960,1984]" captionTargetBox="[175,1409,202,1932]" captionTargetId="figure-17@59.[175,1413,198,1936]" captionTargetPageId="59" captionText="FIGURE 39. Benincandona trapezoides gen. et spec. nov., limbs (AB ♀ and CE ♂). A. T1, Lpp (INV.159280). B. T1, Rpp (INV.159280). C. A2 (INV.159264, INV.159269). D. T1 (INV.159268). E. CR (INV.159264). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745802" httpUri="https://zenodo.org/record/13745802/files/figure.png" pageId="61" pageNumber="62">Figure 39B</figureCitation>
) distally set with almost 10 plumose setae of different morphology and length; more proximally set with d, d and b setae. Prehensile palps asymmetrical Rpp (
<figureCitation id="EA8DAAA94508FF81FBD11E7EFB03287F" box="[1031,1165,1328,1354]" captionStart="FIGURE 39" captionStartId="59.[151,250,1960,1984]" captionTargetBox="[175,1409,202,1932]" captionTargetId="figure-17@59.[175,1413,198,1936]" captionTargetPageId="59" captionText="FIGURE 39. Benincandona trapezoides gen. et spec. nov., limbs (AB ♀ and CE ♂). A. T1, Lpp (INV.159280). B. T1, Rpp (INV.159280). C. A2 (INV.159264, INV.159269). D. T1 (INV.159268). E. CR (INV.159264). Scale bar = 50 µm." pageId="61" pageNumber="62">Figure 39B</figureCitation>
<figureCitation id="EA8DAAA94508FF81FBD11E7EFB03287F" box="[1031,1165,1328,1354]" captionStart="FIGURE 39" captionStartId="59.[151,250,1960,1984]" captionTargetBox="[175,1409,202,1932]" captionTargetId="figure-17@59.[175,1413,198,1936]" captionTargetPageId="59" captionText="FIGURE 39. Benincandona trapezoides gen. et spec. nov., limbs (AB ♀ and CE ♂). A. T1, Lpp (INV.159280). B. T1, Rpp (INV.159280). C. A2 (INV.159264, INV.159269). D. T1 (INV.159268). E. CR (INV.159264). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745802" httpUri="https://zenodo.org/record/13745802/files/figure.png" pageId="61" pageNumber="62">Figure 39B</figureCitation>
) slightly more swollen than Lpp (
<figureCitation id="EA8DAAA94508FF81FEDB1E1AFE1B285B" box="[269,405,1364,1390]" captionStart="FIGURE 39" captionStartId="59.[151,250,1960,1984]" captionTargetBox="[175,1409,202,1932]" captionTargetId="figure-17@59.[175,1413,198,1936]" captionTargetPageId="59" captionText="FIGURE 39. Benincandona trapezoides gen. et spec. nov., limbs (AB ♀ and CE ♂). A. T1, Lpp (INV.159280). B. T1, Rpp (INV.159280). C. A2 (INV.159264, INV.159269). D. T1 (INV.159268). E. CR (INV.159264). Scale bar = 50 µm." pageId="61" pageNumber="62">Figure 39A</figureCitation>
<figureCitation id="EA8DAAA94508FF81FEDB1E1AFE1B285B" box="[269,405,1364,1390]" captionStart="FIGURE 39" captionStartId="59.[151,250,1960,1984]" captionTargetBox="[175,1409,202,1932]" captionTargetId="figure-17@59.[175,1413,198,1936]" captionTargetPageId="59" captionText="FIGURE 39. Benincandona trapezoides gen. et spec. nov., limbs (AB ♀ and CE ♂). A. T1, Lpp (INV.159280). B. T1, Rpp (INV.159280). C. A2 (INV.159264, INV.159269). D. T1 (INV.159268). E. CR (INV.159264). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745802" httpUri="https://zenodo.org/record/13745802/files/figure.png" pageId="61" pageNumber="62">Figure 39A</figureCitation>
); one-segmented, distally sigmoid hook-like, both palps set with a short sensory organ and proximally with two subequal setae.
</paragraph>
<paragraph id="7209B62C4508FF81FF111ED2FB4E2B5F" blockId="61.[151,1437,1256,2038]" pageId="61" pageNumber="62">
T2 (
<figureCitation id="EA8DAAA94508FF81FF231ED2FEF62883" box="[245,376,1436,1462]" captionStart="FIGURE 37" captionStartId="57.[151,250,1981,2005]" captionTargetBox="[223,1388,222,1895]" captionTargetId="figure-17@57.[166,1422,181,1957]" captionTargetPageId="57" captionText="FIGURE 37. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. T2 (INV.159263). B. T3 (INV.159263). C. CR (INV.159280). D. CRAtt (INV.159280). Scale bar = 50 µm." pageId="61" pageNumber="62">Figure 37A</figureCitation>
<figureCitation id="EA8DAAA94508FF81FF231ED2FEF62883" box="[245,376,1436,1462]" captionStart="FIGURE 37" captionStartId="57.[151,250,1981,2005]" captionTargetBox="[223,1388,222,1895]" captionTargetId="figure-17@57.[166,1422,181,1957]" captionTargetPageId="57" captionText="FIGURE 37. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. T2 (INV.159263). B. T3 (INV.159263). C. CR (INV.159280). D. CRAtt (INV.159280). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745798" httpUri="https://zenodo.org/record/13745798/files/figure.png" pageId="61" pageNumber="62">Figure 37A</figureCitation>
) with 4-segmented endopodite (penultimate segment divided), generally elongated. First basal segment without setae d1 and d2. First segment of endopodite elongated (length more than four times basal width), without seta. Second segment (segment 2A) with length
<emphasis id="40C26A3E4508FF81FCF81EAAFCCF28C8" box="[814,833,1508,1533]" italics="true" pageId="61" pageNumber="62">c.</emphasis>
4x width with one short ventro-apical seta f. Third segment (segment 2B) about the same length/width ratio as the previous segment, set with one ventro-subapical seta g. Terminal segment (segment 3) almost as long as width, apically with two short setae (h1 and h3), h1 (ventral) and twice the length of h3 (dorsal), and a long and slender claw (h2), distally set with short setulae.
</paragraph>
<paragraph id="7209B62C4508FF81FF111D3AFF642A77" blockId="61.[151,1437,1256,2038]" pageId="61" pageNumber="62">
T3 (
<figureCitation id="EA8DAAA94508FF81FF231D3AFEF82BBB" box="[245,374,1652,1678]" captionStart="FIGURE 37" captionStartId="57.[151,250,1981,2005]" captionTargetBox="[223,1388,222,1895]" captionTargetId="figure-17@57.[166,1422,181,1957]" captionTargetPageId="57" captionText="FIGURE 37. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. T2 (INV.159263). B. T3 (INV.159263). C. CR (INV.159280). D. CRAtt (INV.159280). Scale bar = 50 µm." pageId="61" pageNumber="62">Figure 37B</figureCitation>
<figureCitation id="EA8DAAA94508FF81FF231D3AFEF82BBB" box="[245,374,1652,1678]" captionStart="FIGURE 37" captionStartId="57.[151,250,1981,2005]" captionTargetBox="[223,1388,222,1895]" captionTargetId="figure-17@57.[166,1422,181,1957]" captionTargetPageId="57" captionText="FIGURE 37. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. T2 (INV.159263). B. T3 (INV.159263). C. CR (INV.159280). D. CRAtt (INV.159280). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745798" httpUri="https://zenodo.org/record/13745798/files/figure.png" pageId="61" pageNumber="62">Figure 37B</figureCitation>
) with first segment carrying only one subapical seta d1, of intermediate length. Second (knee-) segment without seta d2. Third segment about four times as long as wide, not carrying any seta. Fourth segment narrower and at least 8x basal width and carrying one short subapical seta g. Terminal segment well-separated from penultimate segment, about twice as long as basal width and carrying three setae: one long and reflexed (h3), almost reaching knee-segment and two ventrally directed and curved setae h1 and h2, the later slightly exceeding of the former.
</paragraph>
<paragraph id="7209B62C4508FF81FF111C02FC862A9B" blockId="61.[151,1437,1256,2038]" pageId="61" pageNumber="62">
CR (
<figureCitation id="EA8DAAA94508FF81FF2C1C02FEF52A53" box="[250,379,1868,1894]" captionStart="FIGURE 37" captionStartId="57.[151,250,1981,2005]" captionTargetBox="[223,1388,222,1895]" captionTargetId="figure-17@57.[166,1422,181,1957]" captionTargetPageId="57" captionText="FIGURE 37. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. T2 (INV.159263). B. T3 (INV.159263). C. CR (INV.159280). D. CRAtt (INV.159280). Scale bar = 50 µm." pageId="61" pageNumber="62">Figure 37C</figureCitation>
<figureCitation id="EA8DAAA94508FF81FF2C1C02FEF52A53" box="[250,379,1868,1894]" captionStart="FIGURE 37" captionStartId="57.[151,250,1981,2005]" captionTargetBox="[223,1388,222,1895]" captionTargetId="figure-17@57.[166,1422,181,1957]" captionTargetPageId="57" captionText="FIGURE 37. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. T2 (INV.159263). B. T3 (INV.159263). C. CR (INV.159280). D. CRAtt (INV.159280). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745798" httpUri="https://zenodo.org/record/13745798/files/figure.png" pageId="61" pageNumber="62">Figure 37C</figureCitation>
) with stout ramus; claw Ga well-developed, stout with length slightly exceeding half the length of ramus, setae Sp, Sa and claw Gp reduced to small setae; seta Sp situated about at one-fifth length of ramus from distal end, Sa dorso-subapical and Gp ventro-subapical.
</paragraph>
<paragraph id="7209B62C4508FF81FF111CF6FA802AE7" blockId="61.[151,1437,1256,2038]" box="[199,1294,1976,2002]" pageId="61" pageNumber="62">
CRAtt (
<figureCitation id="EA8DAAA94508FF81FEC91CF6FE2C2AE7" box="[287,418,1976,2002]" captionStart="FIGURE 37" captionStartId="57.[151,250,1981,2005]" captionTargetBox="[223,1388,222,1895]" captionTargetId="figure-17@57.[166,1422,181,1957]" captionTargetPageId="57" captionText="FIGURE 37. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. T2 (INV.159263). B. T3 (INV.159263). C. CR (INV.159280). D. CRAtt (INV.159280). Scale bar = 50 µm." pageId="61" pageNumber="62">Figure 37D</figureCitation>
<figureCitation id="EA8DAAA94508FF81FEC91CF6FE2C2AE7" box="[287,418,1976,2002]" captionStart="FIGURE 37" captionStartId="57.[151,250,1981,2005]" captionTargetBox="[223,1388,222,1895]" captionTargetId="figure-17@57.[166,1422,181,1957]" captionTargetPageId="57" captionText="FIGURE 37. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. T2 (INV.159263). B. T3 (INV.159263). C. CR (INV.159280). D. CRAtt (INV.159280). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745798" httpUri="https://zenodo.org/record/13745798/files/figure.png" pageId="61" pageNumber="62">Figure 37D</figureCitation>
) long and stout, distally bifurcated, subapically with additional lateral branch.
</paragraph>
<paragraph id="7209B62C4508FF82FF111C92FD4E2DE0" blockId="61.[151,1437,1256,2038]" lastBlockId="62.[151,1436,151,249]" lastPageId="62" lastPageNumber="63" pageId="61" pageNumber="62">
Hp (
<figureCitation id="EA8DAAA94508FF81FF211C92FE142AC3" box="[247,410,2012,2038]" captionStart="FIGURE 38" captionStartId="58.[151,250,1943,1967]" captionTargetBox="[189,1403,188,1912]" captionTargetId="figure-31@58.[179,1408,181,1919]" captionTargetPageId="58" captionText="FIGURE 38. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159263). BD. Hp (INV.159263). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." pageId="61" pageNumber="62">Figure 38BD</figureCitation>
<figureCitation id="EA8DAAA94508FF81FF211C92FE142AC3" box="[247,410,2012,2038]" captionStart="FIGURE 38" captionStartId="58.[151,250,1943,1967]" captionTargetBox="[189,1403,188,1912]" captionTargetId="figure-31@58.[179,1408,181,1919]" captionTargetPageId="58" captionText="FIGURE 38. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159263). BD. Hp (INV.159263). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745800" httpUri="https://zenodo.org/record/13745800/files/figure.png" pageId="61" pageNumber="62">Figure 38BD</figureCitation>
) with robust and rounded lobe ls, reaching well beyond the asymmetrically rounded lobe ms, both lobes overlapping, not diverging; lobe h shorter and sub-rectangular; internal anatomy (
<figureCitation id="EA8DAAA9450BFF82FB611BD9FAD52D84" box="[1207,1371,151,177]" captionStart="FIGURE 38" captionStartId="58.[151,250,1943,1967]" captionTargetBox="[189,1403,188,1912]" captionTargetId="figure-31@58.[179,1408,181,1919]" captionTargetPageId="58" captionText="FIGURE 38. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159263). BD. Hp (INV.159263). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." pageId="62" pageNumber="63">Figure 38C, D</figureCitation>
<figureCitation id="EA8DAAA9450BFF82FB611BD9FAD52D84" box="[1207,1371,151,177]" captionStart="FIGURE 38" captionStartId="58.[151,250,1943,1967]" captionTargetBox="[189,1403,188,1912]" captionTargetId="figure-31@58.[179,1408,181,1919]" captionTargetPageId="58" captionText="FIGURE 38. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159263). BD. Hp (INV.159263). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745800" httpUri="https://zenodo.org/record/13745800/files/figure.png" pageId="62" pageNumber="63">Figure 38C, D</figureCitation>
) with postlabyrinthal spermiduct with one circular coil.
</paragraph>
<paragraph id="7209B62C450BFF82FF111B91FC382DCC" blockId="62.[151,1436,151,249]" box="[199,950,223,249]" pageId="62" pageNumber="63">
Zenker organ (
<figureCitation id="EA8DAAA9450BFF82FEBD1B91FE602DCC" box="[363,494,223,249]" captionStart="FIGURE 38" captionStartId="58.[151,250,1943,1967]" captionTargetBox="[189,1403,188,1912]" captionTargetId="figure-31@58.[179,1408,181,1919]" captionTargetPageId="58" captionText="FIGURE 38. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159263). BD. Hp (INV.159263). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." pageId="62" pageNumber="63">Figure 38A</figureCitation>
<figureCitation id="EA8DAAA9450BFF82FEBD1B91FE602DCC" box="[363,494,223,249]" captionStart="FIGURE 38" captionStartId="58.[151,250,1943,1967]" captionTargetBox="[189,1403,188,1912]" captionTargetId="figure-31@58.[179,1408,181,1919]" captionTargetPageId="58" captionText="FIGURE 38. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159263). BD. Hp (INV.159263). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745800" httpUri="https://zenodo.org/record/13745800/files/figure.png" pageId="62" pageNumber="63">Figure 38A</figureCitation>
) elongated with
<emphasis id="40C26A3E450BFF82FD7C1B91FD332DCD" box="[682,701,223,248]" italics="true" pageId="62" pageNumber="63">c.</emphasis>
seven spinous whirls.
@ -518,26 +519,26 @@ seven spinous whirls.
</paragraph>
<paragraph id="7209B62C450BFF82FF411A05FB2E2C50" blockId="62.[151,1437,294,825]" box="[151,1184,331,357]" pageId="62" pageNumber="63">
Cp and valves (
<figureCitation id="EA8DAAA9450BFF82FE921A05FE7E2C50" box="[324,496,331,357]" captionStart="FIGURE 32" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure-17@51.[179,1408,181,1920]" captionTargetPageId="51" captionText="FIGURE 32. Benincandona trapezoides gen. et spec. nov., valves (AC, GK ♂; DF ♀). AF, H. CpRl (AC, H. INV.159276, DF. INV.1592 INV.159279). G, I, J. INV.159279. CpV. K. INV.159277. CpD. Scale bars. 1 (B, E, G, K) = 500 µm; 2 (A, C, D, F, I) = 200 µm, 3 (H, J) = 40 µm." pageId="62" pageNumber="63">Figures 32DF</figureCitation>
<figureCitation id="EA8DAAA9450BFF82FE921A05FE7E2C50" box="[324,496,331,357]" captionStart="FIGURE 32" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure-17@51.[179,1408,181,1920]" captionTargetPageId="51" captionText="FIGURE 32. Benincandona trapezoides gen. et spec. nov., valves (AC, GK ♂; DF ♀). AF, H. CpRl (AC, H. INV.159276, DF. INV.1592 INV.159279). G, I, J. INV.159279. CpV. K. INV.159277. CpD. Scale bars. 1 (B, E, G, K) = 500 µm; 2 (A, C, D, F, I) = 200 µm, 3 (H, J) = 40 µm." figureDoi="http://doi.org/10.5281/zenodo.13745788" httpUri="https://zenodo.org/record/13745788/files/figure.png" pageId="62" pageNumber="63">Figures 32DF</figureCitation>
,
<figureCitation id="EA8DAAA9450BFF82FE2D1A05FDC02C50" box="[507,590,331,357]" captionStart="FIGURE 34" captionStartId="54.[151,250,1520,1544]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-31@54.[151,1436,181,1496]" captionTargetPageId="54" captionText="FIGURE 34. Benincandona trapezoides gen. et spec. nov., valves (♀). A, C, D, G. LVi (INV.159264). B, E, F, H. RVi (INV.159264). G, H. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (GH) = 50 µm." pageId="62" pageNumber="63">34AH</figureCitation>
<figureCitation id="EA8DAAA9450BFF82FE2D1A05FDC02C50" box="[507,590,331,357]" captionStart="FIGURE 34" captionStartId="54.[151,250,1520,1544]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-31@54.[151,1436,181,1496]" captionTargetPageId="54" captionText="FIGURE 34. Benincandona trapezoides gen. et spec. nov., valves (♀). A, C, D, G. LVi (INV.159264). B, E, F, H. RVi (INV.159264). G, H. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (GH) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745792" httpUri="https://zenodo.org/record/13745792/files/figure.png" pageId="62" pageNumber="63">34AH</figureCitation>
) as in the male, without obvious sexual dimorphism.
</paragraph>
<paragraph id="7209B62C450BFF82FF111A21FB072CBC" blockId="62.[151,1437,294,825]" box="[199,1161,367,393]" pageId="62" pageNumber="63">
Limbs (
<figureCitation id="EA8DAAA9450BFF82FECB1A21FE312CBC" box="[285,447,367,393]" captionStart="FIGURE 39" captionStartId="59.[151,250,1960,1984]" captionTargetBox="[175,1409,202,1932]" captionTargetId="figure-17@59.[175,1413,198,1936]" captionTargetPageId="59" captionText="FIGURE 39. Benincandona trapezoides gen. et spec. nov., limbs (AB ♀ and CE ♂). A. T1, Lpp (INV.159280). B. T1, Rpp (INV.159280). C. A2 (INV.159264, INV.159269). D. T1 (INV.159268). E. CR (INV.159264). Scale bar = 50 µm." pageId="62" pageNumber="63">Figure 39CE</figureCitation>
<figureCitation id="EA8DAAA9450BFF82FECB1A21FE312CBC" box="[285,447,367,393]" captionStart="FIGURE 39" captionStartId="59.[151,250,1960,1984]" captionTargetBox="[175,1409,202,1932]" captionTargetId="figure-17@59.[175,1413,198,1936]" captionTargetPageId="59" captionText="FIGURE 39. Benincandona trapezoides gen. et spec. nov., limbs (AB ♀ and CE ♂). A. T1, Lpp (INV.159280). B. T1, Rpp (INV.159280). C. A2 (INV.159264, INV.159269). D. T1 (INV.159268). E. CR (INV.159264). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745802" httpUri="https://zenodo.org/record/13745802/files/figure.png" pageId="62" pageNumber="63">Figure 39CE</figureCitation>
) largely as in the male, but with sexually dimorphic A2 and T1.
</paragraph>
<paragraph id="7209B62C450BFF82FF111ADDFDCA2F08" blockId="62.[151,1437,294,825]" pageId="62" pageNumber="63">
A2 (
<figureCitation id="EA8DAAA9450BFF82FF2F1ADDFEF22C98" box="[249,380,403,429]" captionStart="FIGURE 39" captionStartId="59.[151,250,1960,1984]" captionTargetBox="[175,1409,202,1932]" captionTargetId="figure-17@59.[175,1413,198,1936]" captionTargetPageId="59" captionText="FIGURE 39. Benincandona trapezoides gen. et spec. nov., limbs (AB ♀ and CE ♂). A. T1, Lpp (INV.159280). B. T1, Rpp (INV.159280). C. A2 (INV.159264, INV.159269). D. T1 (INV.159268). E. CR (INV.159264). Scale bar = 50 µm." pageId="62" pageNumber="63">Figure 39C</figureCitation>
<figureCitation id="EA8DAAA9450BFF82FF2F1ADDFEF22C98" box="[249,380,403,429]" captionStart="FIGURE 39" captionStartId="59.[151,250,1960,1984]" captionTargetBox="[175,1409,202,1932]" captionTargetId="figure-17@59.[175,1413,198,1936]" captionTargetPageId="59" captionText="FIGURE 39. Benincandona trapezoides gen. et spec. nov., limbs (AB ♀ and CE ♂). A. T1, Lpp (INV.159280). B. T1, Rpp (INV.159280). C. A2 (INV.159264, INV.159269). D. T1 (INV.159268). E. CR (INV.159264). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745802" httpUri="https://zenodo.org/record/13745802/files/figure.png" pageId="62" pageNumber="63">Figure 39C</figureCitation>
) with second endopodal segment undivided, with t1-seta (long, narrow and plumose) and t2- seta (thin and smooth); apical chaetotaxy (
<figureCitation id="EA8DAAA9450BFF82FDA91AF9FD7C2CE4" box="[639,754,439,465]" captionStart="FIGURE 40" captionStartId="60.[151,250,1516,1540]" captionTargetBox="[151,1435,181,1492]" captionTargetId="figure-31@60.[151,1435,181,1492]" captionTargetPageId="60" captionText="FIGURE 40. Diagrams of apical views of the A2 of Benincandona, showing position of (sub-) terminal claws, setae and aesthetascs of the terminal (small circle) and penultimate segment (large circle). For explanations of codes, see Material and Methods (Abbreviations used in text and figures)." pageId="62" pageNumber="63">Figure 40</figureCitation>
<figureCitation id="EA8DAAA9450BFF82FDA91AF9FD7C2CE4" box="[639,754,439,465]" captionStart="FIGURE 40" captionStartId="60.[151,250,1516,1540]" captionTargetBox="[151,1435,181,1492]" captionTargetId="figure-31@60.[151,1435,181,1492]" captionTargetPageId="60" captionText="FIGURE 40. Diagrams of apical views of the A2 of Benincandona, showing position of (sub-) terminal claws, setae and aesthetascs of the terminal (small circle) and penultimate segment (large circle). For explanations of codes, see Material and Methods (Abbreviations used in text and figures)." figureDoi="http://doi.org/10.5281/zenodo.13745804" httpUri="https://zenodo.org/record/13745804/files/figure.png" pageId="62" pageNumber="63">Figure 40</figureCitation>
) with z1claw-like, z2 and z3 being both setae, z1 about 3x as long as z2, and z3 slightly shorter than z2; claws G1, G2, G3 and GM all reaching about the same point; seta g and aesthetasc y3 as in the male and the tip of Gm slightly longer than the tip of y3. Small claw Gm in female homologous to large claw in the male.
</paragraph>
<paragraph id="7209B62C450BFF82FF111909FB952F9C" blockId="62.[151,1437,294,825]" pageId="62" pageNumber="63">
T1 (
<figureCitation id="EA8DAAA9450BFF82FF221909FEF92F54" box="[244,375,583,609]" captionStart="FIGURE 39" captionStartId="59.[151,250,1960,1984]" captionTargetBox="[175,1409,202,1932]" captionTargetId="figure-17@59.[175,1413,198,1936]" captionTargetPageId="59" captionText="FIGURE 39. Benincandona trapezoides gen. et spec. nov., limbs (AB ♀ and CE ♂). A. T1, Lpp (INV.159280). B. T1, Rpp (INV.159280). C. A2 (INV.159264, INV.159269). D. T1 (INV.159268). E. CR (INV.159264). Scale bar = 50 µm." pageId="62" pageNumber="63">Figure 39D</figureCitation>
<figureCitation id="EA8DAAA9450BFF82FF221909FEF92F54" box="[244,375,583,609]" captionStart="FIGURE 39" captionStartId="59.[151,250,1960,1984]" captionTargetBox="[175,1409,202,1932]" captionTargetId="figure-17@59.[175,1413,198,1936]" captionTargetPageId="59" captionText="FIGURE 39. Benincandona trapezoides gen. et spec. nov., limbs (AB ♀ and CE ♂). A. T1, Lpp (INV.159280). B. T1, Rpp (INV.159280). C. A2 (INV.159264, INV.159269). D. T1 (INV.159268). E. CR (INV.159264). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745802" httpUri="https://zenodo.org/record/13745802/files/figure.png" pageId="62" pageNumber="63">Figure 39D</figureCitation>
) with basal part with
<emphasis id="40C26A3E450BFF82FDB01909FDF42F55" box="[614,634,583,608]" italics="true" pageId="62" pageNumber="63">c.</emphasis>
10 distal setae of unequal length and morphology, setae d and b present, with long additional seta (d) present close to seta d; respiratory plate represented by two long rays. Endopodite set with a short and narrow palp, with three subequal but short apical setae (h1h3).
@ -574,9 +575,9 @@ Other candonid species from the wells in
</paragraph>
<paragraph id="7209B62C450BFF82FF4118E1FE152EFC" blockId="62.[151,739,906,969]" box="[151,411,943,969]" pageId="62" pageNumber="63">
(
<figureCitation id="EA8DAAA9450BFF82FF4918E1FE952EFC" box="[159,283,943,969]" captionStart="FIGURE 42" captionStartId="64.[151,250,1943,1967]" captionTargetBox="[163,1424,181,1920]" captionTargetId="figure-31@64.[163,1424,181,1920]" captionTargetPageId="64" captionText="FIGURE 42. Scanning electron microscope illustration of fifteen of thirty further undescribed candonid species, (CpRl). Numbered: AO. Scale bar (for all individuals) = 500 µm." pageId="62" pageNumber="63">Figures 42</figureCitation>
<figureCitation id="EA8DAAA9450BFF82FF4918E1FE952EFC" box="[159,283,943,969]" captionStart="FIGURE 42" captionStartId="64.[151,250,1943,1967]" captionTargetBox="[163,1424,181,1920]" captionTargetId="figure-31@64.[163,1424,181,1920]" captionTargetPageId="64" captionText="FIGURE 42. Scanning electron microscope illustration of fifteen of thirty further undescribed candonid species, (CpRl). Numbered: AO. Scale bar (for all individuals) = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.13745808" httpUri="https://zenodo.org/record/13745808/files/figure.png" pageId="62" pageNumber="63">Figures 42</figureCitation>
,
<figureCitation id="EA8DAAA9450BFF82FEF018E1FED52EFC" box="[294,347,943,969]" captionStart="FIGURE 43" captionStartId="65.[151,250,1943,1967]" captionTargetBox="[160,1427,181,1920]" captionTargetId="figure-17@65.[160,1427,181,1920]" captionTargetPageId="65" captionText="FIGURE 43. Scanning electron microscope illustration of fifteen of thirty further undescribed candonid species, (CpRl). (Continued PAD). Scale bar (for all individuals) = 500 µm." pageId="62" pageNumber="63">43 A</figureCitation>
<figureCitation id="EA8DAAA9450BFF82FEF018E1FED52EFC" box="[294,347,943,969]" captionStart="FIGURE 43" captionStartId="65.[151,250,1943,1967]" captionTargetBox="[160,1427,181,1920]" captionTargetId="figure-17@65.[160,1427,181,1920]" captionTargetPageId="65" captionText="FIGURE 43. Scanning electron microscope illustration of fifteen of thirty further undescribed candonid species, (CpRl). (Continued PAD). Scale bar (for all individuals) = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.13745810" httpUri="https://zenodo.org/record/13745810/files/figure.png" pageId="62" pageNumber="63">43 A</figureCitation>
<collectingRegion id="B07278CE450BFF82FEBC18E1FE1D2EFC" box="[362,403,943,969]" country="Serbia" name="Ada" pageId="62" pageNumber="63">AD</collectingRegion>
)
@ -585,15 +586,15 @@ Other candonid species from the wells in
Thirty further candonid species, in addition to the five new species described above, have been found in the samples from the wells in
<collectingCountry id="0AA1F6BC450BFF82FE8F1F55FE2E2900" box="[345,416,1051,1077]" name="Benin" pageId="62" pageNumber="63">Benin</collectingCountry>
. Most of these are also (sub-) triangular in lateral view, with strong (
<figureCitation id="EA8DAAA9450BFF82FB441F55FA852900" box="[1170,1291,1051,1077]" captionStart="FIGURE 42" captionStartId="64.[151,250,1943,1967]" captionTargetBox="[163,1424,181,1920]" captionTargetId="figure-31@64.[163,1424,181,1920]" captionTargetPageId="64" captionText="FIGURE 42. Scanning electron microscope illustration of fifteen of thirty further undescribed candonid species, (CpRl). Numbered: AO. Scale bar (for all individuals) = 500 µm." pageId="62" pageNumber="63">Figures 42</figureCitation>
<figureCitation id="EA8DAAA9450BFF82FB441F55FA852900" box="[1170,1291,1051,1077]" captionStart="FIGURE 42" captionStartId="64.[151,250,1943,1967]" captionTargetBox="[163,1424,181,1920]" captionTargetId="figure-31@64.[163,1424,181,1920]" captionTargetPageId="64" captionText="FIGURE 42. Scanning electron microscope illustration of fifteen of thirty further undescribed candonid species, (CpRl). Numbered: AO. Scale bar (for all individuals) = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.13745808" httpUri="https://zenodo.org/record/13745808/files/figure.png" pageId="62" pageNumber="63">Figures 42</figureCitation>
/43E, N, AB) or weak (
<figureCitation id="EA8DAAA9450BFF82FF291F71FEF6296C" box="[255,376,1087,1113]" captionStart="FIGURE 42" captionStartId="64.[151,250,1943,1967]" captionTargetBox="[163,1424,181,1920]" captionTargetId="figure-31@64.[163,1424,181,1920]" captionTargetPageId="64" captionText="FIGURE 42. Scanning electron microscope illustration of fifteen of thirty further undescribed candonid species, (CpRl). Numbered: AO. Scale bar (for all individuals) = 500 µm." pageId="62" pageNumber="63">Figures 42</figureCitation>
<figureCitation id="EA8DAAA9450BFF82FF291F71FEF6296C" box="[255,376,1087,1113]" captionStart="FIGURE 42" captionStartId="64.[151,250,1943,1967]" captionTargetBox="[163,1424,181,1920]" captionTargetId="figure-31@64.[163,1424,181,1920]" captionTargetPageId="64" captionText="FIGURE 42. Scanning electron microscope illustration of fifteen of thirty further undescribed candonid species, (CpRl). Numbered: AO. Scale bar (for all individuals) = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.13745808" httpUri="https://zenodo.org/record/13745808/files/figure.png" pageId="62" pageNumber="63">Figures 42</figureCitation>
/43K, M, Z) external lateral ridges; others have different shapes (
<figureCitation id="EA8DAAA9450BFF82FB9B1F71FB4B296C" box="[1101,1221,1087,1113]" captionStart="FIGURE 42" captionStartId="64.[151,250,1943,1967]" captionTargetBox="[163,1424,181,1920]" captionTargetId="figure-31@64.[163,1424,181,1920]" captionTargetPageId="64" captionText="FIGURE 42. Scanning electron microscope illustration of fifteen of thirty further undescribed candonid species, (CpRl). Numbered: AO. Scale bar (for all individuals) = 500 µm." pageId="62" pageNumber="63">Figures 42</figureCitation>
<figureCitation id="EA8DAAA9450BFF82FB9B1F71FB4B296C" box="[1101,1221,1087,1113]" captionStart="FIGURE 42" captionStartId="64.[151,250,1943,1967]" captionTargetBox="[163,1424,181,1920]" captionTargetId="figure-31@64.[163,1424,181,1920]" captionTargetPageId="64" captionText="FIGURE 42. Scanning electron microscope illustration of fifteen of thirty further undescribed candonid species, (CpRl). Numbered: AO. Scale bar (for all individuals) = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.13745808" httpUri="https://zenodo.org/record/13745808/files/figure.png" pageId="62" pageNumber="63">Figures 42</figureCitation>
/43F, G, O, V, AC). Some species (
<figureCitation id="EA8DAAA9450BFF82FEE81F2DFD892948" box="[318,519,1123,1149]" captionStart="FIGURE 42" captionStartId="64.[151,250,1943,1967]" captionTargetBox="[163,1424,181,1920]" captionTargetId="figure-31@64.[163,1424,181,1920]" captionTargetPageId="64" captionText="FIGURE 42. Scanning electron microscope illustration of fifteen of thirty further undescribed candonid species, (CpRl). Numbered: AO. Scale bar (for all individuals) = 500 µm." pageId="62" pageNumber="63">Figure 42A, D, N</figureCitation>
<figureCitation id="EA8DAAA9450BFF82FEE81F2DFD892948" box="[318,519,1123,1149]" captionStart="FIGURE 42" captionStartId="64.[151,250,1943,1967]" captionTargetBox="[163,1424,181,1920]" captionTargetId="figure-31@64.[163,1424,181,1920]" captionTargetPageId="64" captionText="FIGURE 42. Scanning electron microscope illustration of fifteen of thirty further undescribed candonid species, (CpRl). Numbered: AO. Scale bar (for all individuals) = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.13745808" httpUri="https://zenodo.org/record/13745808/files/figure.png" pageId="62" pageNumber="63">Figure 42A, D, N</figureCitation>
) are much larger than most, and some are much smaller (
<figureCitation id="EA8DAAA9450BFF82FB5A1F2DFA882948" box="[1164,1286,1123,1149]" captionStart="FIGURE 42" captionStartId="64.[151,250,1943,1967]" captionTargetBox="[163,1424,181,1920]" captionTargetId="figure-31@64.[163,1424,181,1920]" captionTargetPageId="64" captionText="FIGURE 42. Scanning electron microscope illustration of fifteen of thirty further undescribed candonid species, (CpRl). Numbered: AO. Scale bar (for all individuals) = 500 µm." pageId="62" pageNumber="63">Figures 42</figureCitation>
<figureCitation id="EA8DAAA9450BFF82FB5A1F2DFA882948" box="[1164,1286,1123,1149]" captionStart="FIGURE 42" captionStartId="64.[151,250,1943,1967]" captionTargetBox="[163,1424,181,1920]" captionTargetId="figure-31@64.[163,1424,181,1920]" captionTargetPageId="64" captionText="FIGURE 42. Scanning electron microscope illustration of fifteen of thirty further undescribed candonid species, (CpRl). Numbered: AO. Scale bar (for all individuals) = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.13745808" httpUri="https://zenodo.org/record/13745808/files/figure.png" pageId="62" pageNumber="63">Figures 42</figureCitation>
/43O, X, AA, AD). Based on preliminary analyses of the soft parts of these species, it is at present most likely that all of these species belong to
<taxonomicName id="B5B6CDAF450BFF82FE891FE5FD6529F0" authority="Hotekpo &amp; Martens" authorityName="Hotekpo &amp; Martens" box="[351,747,1195,1221]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="62" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="40C26A3E450BFF82FE891FE5FD8F29F0" box="[351,513,1195,1221]" italics="true" pageId="62" pageNumber="63">Benincandona</emphasis>

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@ -1,64 +1,65 @@
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(
<figureCitation id="EA8DAAA9451DFF94FF491909FECA2F54" box="[159,324,583,609]" captionStart-0="FIGURE 24" captionStart-1="FIGURE 25" captionStart-2="FIGURE 26" captionStart-3="FIGURE 27" captionStart-4="FIGURE 28" captionStart-5="FIGURE 29" captionStart-6="FIGURE 30" captionStart-7="FIGURE 31" captionStartId-0="41.[151,250,1944,1968]" captionStartId-1="42.[151,250,1673,1697]" captionStartId-2="44.[151,250,1673,1697]" captionStartId-3="45.[151,250,1943,1967]" captionStartId-4="46.[151,250,1943,1967]" captionStartId-5="47.[151,250,1981,2005]" captionStartId-6="48.[151,250,1943,1967]" captionStartId-7="49.[151,250,1943,1967]" captionTargetBox-0="[170,1417,181,1921]" captionTargetBox-1="[151,1436,181,1649]" captionTargetBox-2="[151,1436,181,1649]" captionTargetBox-3="[173,1414,183,1915]" captionTargetBox-4="[188,1401,185,1915]" captionTargetBox-5="[197,1421,196,1942]" captionTargetBox-6="[194,1393,191,1910]" captionTargetBox-7="[184,1406,184,1915]" captionTargetId-0="figure-17@41.[170,1417,181,1921]" captionTargetId-1="figure-31@42.[151,1436,181,1649]" captionTargetId-2="figure-31@44.[151,1436,181,1649]" captionTargetId-3="figure-17@45.[171,1417,181,1919]" captionTargetId-4="figure-31@46.[184,1403,181,1919]" captionTargetId-5="figure-17@47.[166,1422,181,1957]" captionTargetId-6="figure-31@48.[184,1403,181,1920]" captionTargetId-7="figure-17@49.[180,1407,181,1919]" captionTargetPageId-0="41" captionTargetPageId-1="42" captionTargetPageId-2="44" captionTargetPageId-3="45" captionTargetPageId-4="46" captionTargetPageId-5="47" captionTargetPageId-6="48" captionTargetPageId-7="49" captionText-0="FIGURE 24. Benincandona cebios gen. et spec. nov., valves (AC, JK ♂; DI ♀). AI. CpRl (AC. INV.159255, DI. INV.159258). J. INV.159256. CpD. K. INV.159257. CpV. Scale bars. 1 (B, E, J, K) = 500 µm; 2 (A, C, D, F, H, I) = 200 µm." captionText-1="FIGURE 25. Benincandona cebios gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159244). B, E, F. RVi (INV.159244). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." captionText-2="FIGURE 26. Benincandona cebios gen. et spec. nov., valves (♀). A, C, D, G. LVi (INV.159253). B, E, F. RVi (INV.159253). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." captionText-3="FIGURE 27. Benincandona cebios gen. et spec. nov., limbs (♂). A. A2 (INV.159244). B. A1 (INV.159244). C. MdCoxa (INV.159248). Scale bar = 50 µm." captionText-4="FIGURE 28. Benincandona cebios gen. et spec. nov., limbs (♂). A. Mx1-palp (INV.159244). B. Mx1, respiratory plate (INV.159248). C. MdPalp (INV.159244). Scale bars. 1 (B), 2 (A), 3 (C) = 50 µm." captionText-5="FIGURE 29. Benincandona cebios gen. et spec. nov., limbs (♂). A. T3 (INV.159244). B. T2 (INV.159244). C. CRAtt (INV.159244). D. CR (INV.159244). Scale bar = 50 µm." captionText-6="FIGURE 30. Benincandona cebios gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159247). BD. Hp (INV.159244). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." captionText-7="FIGURE 31. Benincandona cebios gen. et spec. nov., limbs (AB ♂ and CE ♀). C. A2 (INV.159245). D. CR (INV.159245). E. T1 (INV.159245). A. T1, Lpp (INV.159244). B. T1, Rpp (INV.159244). Scale bar = 50 µm." pageId="40" pageNumber="41">Figures 2431</figureCitation>
<figureCitation id="EA8DAAA9451DFF94FF491909FECA2F54" box="[159,324,583,609]" captionStart-0="FIGURE 24" captionStart-1="FIGURE 25" captionStart-2="FIGURE 26" captionStart-3="FIGURE 27" captionStart-4="FIGURE 28" captionStart-5="FIGURE 29" captionStart-6="FIGURE 30" captionStart-7="FIGURE 31" captionStartId-0="41.[151,250,1944,1968]" captionStartId-1="42.[151,250,1673,1697]" captionStartId-2="44.[151,250,1673,1697]" captionStartId-3="45.[151,250,1943,1967]" captionStartId-4="46.[151,250,1943,1967]" captionStartId-5="47.[151,250,1981,2005]" captionStartId-6="48.[151,250,1943,1967]" captionStartId-7="49.[151,250,1943,1967]" captionTargetBox-0="[170,1417,181,1921]" captionTargetBox-1="[151,1436,181,1649]" captionTargetBox-2="[151,1436,181,1649]" captionTargetBox-3="[173,1414,183,1915]" captionTargetBox-4="[188,1401,185,1915]" captionTargetBox-5="[197,1421,196,1942]" captionTargetBox-6="[194,1393,191,1910]" captionTargetBox-7="[184,1406,184,1915]" captionTargetId-0="figure-17@41.[170,1417,181,1921]" captionTargetId-1="figure-31@42.[151,1436,181,1649]" captionTargetId-2="figure-31@44.[151,1436,181,1649]" captionTargetId-3="figure-17@45.[171,1417,181,1919]" captionTargetId-4="figure-31@46.[184,1403,181,1919]" captionTargetId-5="figure-17@47.[166,1422,181,1957]" captionTargetId-6="figure-31@48.[184,1403,181,1920]" captionTargetId-7="figure-17@49.[180,1407,181,1919]" captionTargetPageId-0="41" captionTargetPageId-1="42" captionTargetPageId-2="44" captionTargetPageId-3="45" captionTargetPageId-4="46" captionTargetPageId-5="47" captionTargetPageId-6="48" captionTargetPageId-7="49" captionText-0="FIGURE 24. Benincandona cebios gen. et spec. nov., valves (AC, JK ♂; DI ♀). AI. CpRl (AC. INV.159255, DI. INV.159258). J. INV.159256. CpD. K. INV.159257. CpV. Scale bars. 1 (B, E, J, K) = 500 µm; 2 (A, C, D, F, H, I) = 200 µm." captionText-1="FIGURE 25. Benincandona cebios gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159244). B, E, F. RVi (INV.159244). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." captionText-2="FIGURE 26. Benincandona cebios gen. et spec. nov., valves (♀). A, C, D, G. LVi (INV.159253). B, E, F. RVi (INV.159253). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." captionText-3="FIGURE 27. Benincandona cebios gen. et spec. nov., limbs (♂). A. A2 (INV.159244). B. A1 (INV.159244). C. MdCoxa (INV.159248). Scale bar = 50 µm." captionText-4="FIGURE 28. Benincandona cebios gen. et spec. nov., limbs (♂). A. Mx1-palp (INV.159244). B. Mx1, respiratory plate (INV.159248). C. MdPalp (INV.159244). Scale bars. 1 (B), 2 (A), 3 (C) = 50 µm." captionText-5="FIGURE 29. Benincandona cebios gen. et spec. nov., limbs (♂). A. T3 (INV.159244). B. T2 (INV.159244). C. CRAtt (INV.159244). D. CR (INV.159244). Scale bar = 50 µm." captionText-6="FIGURE 30. Benincandona cebios gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159247). BD. Hp (INV.159244). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." captionText-7="FIGURE 31. Benincandona cebios gen. et spec. nov., limbs (AB ♂ and CE ♀). C. A2 (INV.159245). D. CR (INV.159245). E. T1 (INV.159245). A. T1, Lpp (INV.159244). B. T1, Rpp (INV.159244). Scale bar = 50 µm." figureDoi-0="http://doi.org/10.5281/zenodo.13745770" figureDoi-1="http://doi.org/10.5281/zenodo.13745772" figureDoi-2="http://doi.org/10.5281/zenodo.13745774" figureDoi-3="http://doi.org/10.5281/zenodo.13745776" figureDoi-4="http://doi.org/10.5281/zenodo.13745778" figureDoi-5="http://doi.org/10.5281/zenodo.13745780" figureDoi-6="http://doi.org/10.5281/zenodo.13745782" figureDoi-7="http://doi.org/10.5281/zenodo.13745784" httpUri-0="https://zenodo.org/record/13745770/files/figure.png" httpUri-1="https://zenodo.org/record/13745772/files/figure.png" httpUri-2="https://zenodo.org/record/13745774/files/figure.png" httpUri-3="https://zenodo.org/record/13745776/files/figure.png" httpUri-4="https://zenodo.org/record/13745778/files/figure.png" httpUri-5="https://zenodo.org/record/13745780/files/figure.png" httpUri-6="https://zenodo.org/record/13745782/files/figure.png" httpUri-7="https://zenodo.org/record/13745784/files/figure.png" pageId="40" pageNumber="41">Figures 2431</figureCitation>
,
<figureCitation id="EA8DAAA9451DFF94FE861909FE2D2F54" box="[336,419,583,609]" captionStart="FIGURE 41" captionStartId="61.[151,250,1113,1137]" captionTargetBox="[151,1436,181,1089]" captionTargetId="figure-17@61.[151,1436,181,1089]" captionTargetPageId="61" captionText="FIGURE 41. Benincandona gen. et spec. nov., CpRl. (A, C, E, F, G, I, K, N, O, Q ♂ and B, D, H, J, L, M, P, R ♀). AD. B. martini. EF. B. ibikounlei. GJ. B. sakitii. KN. B. cebios. OR. B. trapezoides. Scale bar = 500 µm." pageId="40" pageNumber="41">41KN</figureCitation>
<figureCitation id="EA8DAAA9451DFF94FE861909FE2D2F54" box="[336,419,583,609]" captionStart="FIGURE 41" captionStartId="61.[151,250,1113,1137]" captionTargetBox="[151,1436,181,1089]" captionTargetId="figure-17@61.[151,1436,181,1089]" captionTargetPageId="61" captionText="FIGURE 41. Benincandona gen. et spec. nov., CpRl. (A, C, E, F, G, I, K, N, O, Q ♂ and B, D, H, J, L, M, P, R ♀). AD. B. martini. EF. B. ibikounlei. GJ. B. sakitii. KN. B. cebios. OR. B. trapezoides. Scale bar = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.13745806" httpUri="https://zenodo.org/record/13745806/files/figure.png" pageId="40" pageNumber="41">41KN</figureCitation>
)
</paragraph>
</subSubSection>
@ -213,7 +214,7 @@ See
for a list of localities where the present species was collected.
</paragraph>
</subSubSection>
<caption id="26C9E6A4451CFF95FF411CD6FA052AE0" pageId="41" pageNumber="42" startId="41.[151,250,1944,1968]" targetBox="[170,1417,181,1921]" targetPageId="41" targetType="figure">
<caption id="26C9E6A4451CFF95FF411CD6FA052AE0" ID-DOI="http://doi.org/10.5281/zenodo.13745770" ID-Zenodo-Dep="13745770" httpUri="https://zenodo.org/record/13745770/files/figure.png" pageId="41" pageNumber="42" startId="41.[151,250,1944,1968]" targetBox="[170,1417,181,1921]" targetPageId="41" targetType="figure">
<paragraph id="7209B62C451CFF95FF411CD6FA052AE0" blockId="41.[151,1437,1944,2005]" pageId="41" pageNumber="42">
<emphasis id="40C26A3E451CFF95FF411CD6FEAD2A85" bold="true" box="[151,291,1944,1968]" pageId="41" pageNumber="42">FIGURE 24.</emphasis>
<taxonomicName id="B5B6CDAF451CFF95FEFB1CD7FD842A84" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[301,522,1945,1969]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="cebios" status="gen. et spec. nov.">
@ -229,7 +230,7 @@ for a list of localities where the present species was collected.
. INV.159257. CpV. Scale bars. 1 (B, E, J, K) = 500 µm; 2 (A, C, D, F, H, I) = 200 µm.
</paragraph>
</caption>
<caption id="26C9E6A4451FFF96FF411DC7FC172BF3" pageId="42" pageNumber="43" startId="42.[151,250,1673,1697]" targetBox="[151,1436,181,1649]" targetPageId="42" targetType="figure">
<caption id="26C9E6A4451FFF96FF411DC7FC172BF3" ID-DOI="http://doi.org/10.5281/zenodo.13745772" ID-Zenodo-Dep="13745772" httpUri="https://zenodo.org/record/13745772/files/figure.png" pageId="42" pageNumber="43" startId="42.[151,250,1673,1697]" targetBox="[151,1436,181,1649]" targetPageId="42" targetType="figure">
<paragraph id="7209B62C451FFF96FF411DC7FC172BF3" blockId="42.[151,1437,1673,1734]" pageId="42" pageNumber="43">
<emphasis id="40C26A3E451FFF96FF411DC7FEAE2B94" bold="true" box="[151,288,1673,1697]" pageId="42" pageNumber="43">FIGURE 25.</emphasis>
<taxonomicName id="B5B6CDAF451FFF96FEF11DC7FD8F2B94" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[295,513,1673,1697]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="cebios" status="gen. et spec. nov.">
@ -251,33 +252,33 @@ CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm.
</paragraph>
<paragraph id="7209B62C451FFF97FF411C72FDA32DE0" blockId="42.[151,1436,1815,2022]" lastBlockId="43.[151,1437,151,2013]" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
CpRl (
<figureCitation id="EA8DAAA9451FFF96FF351C72FE042A63" box="[227,394,1852,1878]" captionStart="FIGURE 24" captionStartId="41.[151,250,1944,1968]" captionTargetBox="[170,1417,181,1921]" captionTargetId="figure-17@41.[170,1417,181,1921]" captionTargetPageId="41" captionText="FIGURE 24. Benincandona cebios gen. et spec. nov., valves (AC, JK ♂; DI ♀). AI. CpRl (AC. INV.159255, DI. INV.159258). J. INV.159256. CpD. K. INV.159257. CpV. Scale bars. 1 (B, E, J, K) = 500 µm; 2 (A, C, D, F, H, I) = 200 µm." pageId="42" pageNumber="43">Figure 24AC</figureCitation>
<figureCitation id="EA8DAAA9451FFF96FF351C72FE042A63" box="[227,394,1852,1878]" captionStart="FIGURE 24" captionStartId="41.[151,250,1944,1968]" captionTargetBox="[170,1417,181,1921]" captionTargetId="figure-17@41.[170,1417,181,1921]" captionTargetPageId="41" captionText="FIGURE 24. Benincandona cebios gen. et spec. nov., valves (AC, JK ♂; DI ♀). AI. CpRl (AC. INV.159255, DI. INV.159258). J. INV.159256. CpD. K. INV.159257. CpV. Scale bars. 1 (B, E, J, K) = 500 µm; 2 (A, C, D, F, H, I) = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.13745770" httpUri="https://zenodo.org/record/13745770/files/figure.png" pageId="42" pageNumber="43">Figure 24AC</figureCitation>
) elongated (L/H ratio=2.0) and sub-triangular, dorsal margin bluntly pointed, with greatest height slightly behind the middle, anterior part of dorsal margin sloping almost straight to anterior margin, passing into it with a blunt angle, anteroventral part of anterior margin straight, posterior part of dorsal margin sloping to ventral margin, the latter almost straight; LV overlapping RV along anterior, ventral and posterior sides, dorsally strongly overlapping RV; valve surface smooth in the anterior and posterior parts, with slight ridges in the middle section. CpD and CpV (
<figureCitation id="EA8DAAA9451EFF97FE7E1BD9FDC62D84" box="[424,584,151,177]" captionStart="FIGURE 24" captionStartId="41.[151,250,1944,1968]" captionTargetBox="[170,1417,181,1921]" captionTargetId="figure-17@41.[170,1417,181,1921]" captionTargetPageId="41" captionText="FIGURE 24. Benincandona cebios gen. et spec. nov., valves (AC, JK ♂; DI ♀). AI. CpRl (AC. INV.159255, DI. INV.159258). J. INV.159256. CpD. K. INV.159257. CpV. Scale bars. 1 (B, E, J, K) = 500 µm; 2 (A, C, D, F, H, I) = 200 µm." pageId="43" pageNumber="44">Figure 24J, K</figureCitation>
<figureCitation id="EA8DAAA9451EFF97FE7E1BD9FDC62D84" box="[424,584,151,177]" captionStart="FIGURE 24" captionStartId="41.[151,250,1944,1968]" captionTargetBox="[170,1417,181,1921]" captionTargetId="figure-17@41.[170,1417,181,1921]" captionTargetPageId="41" captionText="FIGURE 24. Benincandona cebios gen. et spec. nov., valves (AC, JK ♂; DI ♀). AI. CpRl (AC. INV.159255, DI. INV.159258). J. INV.159256. CpD. K. INV.159257. CpV. Scale bars. 1 (B, E, J, K) = 500 µm; 2 (A, C, D, F, H, I) = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.13745770" httpUri="https://zenodo.org/record/13745770/files/figure.png" pageId="43" pageNumber="44">Figure 24J, K</figureCitation>
) narrow and pointed on both anterior and posterior sides, greatest width in the middle, lateral margins rounded.
</paragraph>
<paragraph id="7209B62C451EFF97FF111B91FEA92CBC" blockId="43.[151,1437,151,2013]" pageId="43" pageNumber="44">
LVi (
<figureCitation id="EA8DAAA9451EFF97FED71B91FE7F2DCC" box="[257,497,222,249]" captionStart="FIGURE 25" captionStartId="42.[151,250,1673,1697]" captionTargetBox="[151,1436,181,1649]" captionTargetId="figure-31@42.[151,1436,181,1649]" captionTargetPageId="42" captionText="FIGURE 25. Benincandona cebios gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159244). B, E, F. RVi (INV.159244). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." pageId="43" pageNumber="44">Figure 25A, C, D, G</figureCitation>
<figureCitation id="EA8DAAA9451EFF97FED71B91FE7F2DCC" box="[257,497,222,249]" captionStart="FIGURE 25" captionStartId="42.[151,250,1673,1697]" captionTargetBox="[151,1436,181,1649]" captionTargetId="figure-31@42.[151,1436,181,1649]" captionTargetPageId="42" captionText="FIGURE 25. Benincandona cebios gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159244). B, E, F. RVi (INV.159244). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745772" httpUri="https://zenodo.org/record/13745772/files/figure.png" pageId="43" pageNumber="44">Figure 25A, C, D, G</figureCitation>
) elongated and sub-triangular, dorsal margin bluntly pointed, with greatest height slightly behind the middle, anterior part of dorsal margin sloping almost straight to anterior margin, passing into it with a blunt angle, anteroventral part of anterior margin straight; posterior part of dorsal margin sloping towards ventral margin, the latter almost straight; one ventral il without antero-ventral tooth. CMS (
<figureCitation id="EA8DAAA9451EFF97FB711A05FAA42C50" box="[1191,1322,330,357]" captionStart="FIGURE 25" captionStartId="42.[151,250,1673,1697]" captionTargetBox="[151,1436,181,1649]" captionTargetId="figure-31@42.[151,1436,181,1649]" captionTargetPageId="42" captionText="FIGURE 25. Benincandona cebios gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159244). B, E, F. RVi (INV.159244). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." pageId="43" pageNumber="44">Figure 25G</figureCitation>
<figureCitation id="EA8DAAA9451EFF97FB711A05FAA42C50" box="[1191,1322,330,357]" captionStart="FIGURE 25" captionStartId="42.[151,250,1673,1697]" captionTargetBox="[151,1436,181,1649]" captionTargetId="figure-31@42.[151,1436,181,1649]" captionTargetPageId="42" captionText="FIGURE 25. Benincandona cebios gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159244). B, E, F. RVi (INV.159244). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745772" httpUri="https://zenodo.org/record/13745772/files/figure.png" pageId="43" pageNumber="44">Figure 25G</figureCitation>
) as in the
<typeStatus id="AD0D088E451EFF97FF411A21FF492CBC" box="[151,199,367,393]" pageId="43" pageNumber="44">type</typeStatus>
species.
</paragraph>
<paragraph id="7209B62C451EFF97FF111ADDFC882F2C" blockId="43.[151,1437,151,2013]" pageId="43" pageNumber="44">
RVi (
<figureCitation id="EA8DAAA9451EFF97FED41ADDFE4A2C98" box="[258,452,402,429]" captionStart="FIGURE 25" captionStartId="42.[151,250,1673,1697]" captionTargetBox="[151,1436,181,1649]" captionTargetId="figure-31@42.[151,1436,181,1649]" captionTargetPageId="42" captionText="FIGURE 25. Benincandona cebios gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159244). B, E, F. RVi (INV.159244). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." pageId="43" pageNumber="44">Figure 25B, E, F</figureCitation>
<figureCitation id="EA8DAAA9451EFF97FED41ADDFE4A2C98" box="[258,452,402,429]" captionStart="FIGURE 25" captionStartId="42.[151,250,1673,1697]" captionTargetBox="[151,1436,181,1649]" captionTargetId="figure-31@42.[151,1436,181,1649]" captionTargetPageId="42" captionText="FIGURE 25. Benincandona cebios gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159244). B, E, F. RVi (INV.159244). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745772" httpUri="https://zenodo.org/record/13745772/files/figure.png" pageId="43" pageNumber="44">Figure 25B, E, F</figureCitation>
) elongated, with dorsal margin partly straight and with greatest height well behind the middle, anterior part of dorsal margin sloping almost straight to anterior margin, passing into it with a blunt angle and with anteroventral part straight, posterior part of dorsal margin sloping towards ventral margin, the latter almost straight; il running submarginally along ventral margin.
</paragraph>
<paragraph id="7209B62C451EFF97FF11196DFC8B2E90" blockId="43.[151,1437,151,2013]" pageId="43" pageNumber="44">
A1 (
<figureCitation id="EA8DAAA9451EFF97FF2C196DFEF02F08" box="[250,382,547,573]" captionStart="FIGURE 27" captionStartId="45.[151,250,1943,1967]" captionTargetBox="[173,1414,183,1915]" captionTargetId="figure-17@45.[171,1417,181,1919]" captionTargetPageId="45" captionText="FIGURE 27. Benincandona cebios gen. et spec. nov., limbs (♂). A. A2 (INV.159244). B. A1 (INV.159244). C. MdCoxa (INV.159248). Scale bar = 50 µm." pageId="43" pageNumber="44">Figure 27B</figureCitation>
<figureCitation id="EA8DAAA9451EFF97FF2C196DFEF02F08" box="[250,382,547,573]" captionStart="FIGURE 27" captionStartId="45.[151,250,1943,1967]" captionTargetBox="[173,1414,183,1915]" captionTargetId="figure-17@45.[171,1417,181,1919]" captionTargetPageId="45" captionText="FIGURE 27. Benincandona cebios gen. et spec. nov., limbs (♂). A. A2 (INV.159244). B. A1 (INV.159244). C. MdCoxa (INV.159248). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745776" httpUri="https://zenodo.org/record/13745776/files/figure.png" pageId="43" pageNumber="44">Figure 27B</figureCitation>
) consisting of seven articulated segments, one basal and six distal. Basal segment with two ventro-subapical setae, one long and one shorter, the latter about one third of the length of the longer seta, and two shorter setae on the mid-dorsal side, situated some distance from each other. First distal segment about three times as long as the basal width, with one short dorso-subapical seta. Second distal segment with about the same length/ width ratio as the previous segment, set with one short dorso-apical seta. Third distal segment also with about the same length/width ratio as the previous segment, carrying one short apical setae on each side. Fourth distal segment more than three times as long as wide, and set with two short setae as on the previous segment. Fifth distal segment more than five times as long as wide, set with one short ventro-apical seta and two long dorso-apical plumose setae. Sixth distal segment narrow, more than four times as long as wide, with four apical structures: one aesthetasc Ya of intermediate length, two long but unequal length plumose setae, about twice and 1.5x the length of Ya respectively, and one short plumose seta, about half the length of Ya.
</paragraph>
<paragraph id="7209B62C451EFF97FF1118E1FA9A28D0" blockId="43.[151,1437,151,2013]" pageId="43" pageNumber="44">
A2 (
<figureCitation id="EA8DAAA9451EFF97FF2D18E1FE0C2EFC" box="[251,386,943,969]" captionStart="FIGURE 27" captionStartId="45.[151,250,1943,1967]" captionTargetBox="[173,1414,183,1915]" captionTargetId="figure-17@45.[171,1417,181,1919]" captionTargetPageId="45" captionText="FIGURE 27. Benincandona cebios gen. et spec. nov., limbs (♂). A. A2 (INV.159244). B. A1 (INV.159244). C. MdCoxa (INV.159248). Scale bar = 50 µm." pageId="43" pageNumber="44">Figure 27A</figureCitation>
<figureCitation id="EA8DAAA9451EFF97FF2D18E1FE0C2EFC" box="[251,386,943,969]" captionStart="FIGURE 27" captionStartId="45.[151,250,1943,1967]" captionTargetBox="[173,1414,183,1915]" captionTargetId="figure-17@45.[171,1417,181,1919]" captionTargetPageId="45" captionText="FIGURE 27. Benincandona cebios gen. et spec. nov., limbs (♂). A. A2 (INV.159244). B. A1 (INV.159244). C. MdCoxa (INV.159248). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745776" httpUri="https://zenodo.org/record/13745776/files/figure.png" pageId="43" pageNumber="44">Figure 27A</figureCitation>
) with basal segment (first protopodite) broad and rounded, mid-ventrally with one seta of medium length, subapically with two setae of unequal length, proximal one
<emphasis id="40C26A3E451EFF97FBD2189DFB992ED9" box="[1028,1047,979,1004]" italics="true" pageId="43" pageNumber="44">c.</emphasis>
five times the length of the more apical one (plumose). Second segment (second protopodite) large and
@ -287,29 +288,29 @@ twice as long as basal width, set with one short and plumose ventro-subapical se
three times as long as basal width, carrying one long mid-ventral aesthetasc Y (tip clearly exceeding half of segment) and two ventro-apical setae, one short and one large and stout, hirsute in its distal 4/5. Second endopodal segment (segment 2A)
<emphasis id="40C26A3E451EFF97FA931FC9FAD62995" box="[1349,1368,1159,1184]" italics="true" pageId="43" pageNumber="44">c.</emphasis>
twice as long as wide, carrying one stout dorso-apical plumose seta and three t-setae; seta t1 long, narrow and plumose, accompanied by a long and slender, but well-developed aesthetasc y1; setae t2 and t3 respectively ventro and centro-apical, hyper-developed into male bristles. Third endopodal segment (segment 2B) smaller than previous segment and about 1.5x as long as wide, set with a complex (and sexually dimorphic) apical chaetotaxy (
<figureCitation id="EA8DAAA9451EFF97FB101E59FAB72804" box="[1222,1337,1303,1329]" captionStart="FIGURE 40" captionStartId="60.[151,250,1516,1540]" captionTargetBox="[151,1435,181,1492]" captionTargetId="figure-31@60.[151,1435,181,1492]" captionTargetPageId="60" captionText="FIGURE 40. Diagrams of apical views of the A2 of Benincandona, showing position of (sub-) terminal claws, setae and aesthetascs of the terminal (small circle) and penultimate segment (large circle). For explanations of codes, see Material and Methods (Abbreviations used in text and figures)." pageId="43" pageNumber="44">Figure 40</figureCitation>
<figureCitation id="EA8DAAA9451EFF97FB101E59FAB72804" box="[1222,1337,1303,1329]" captionStart="FIGURE 40" captionStartId="60.[151,250,1516,1540]" captionTargetBox="[151,1435,181,1492]" captionTargetId="figure-31@60.[151,1435,181,1492]" captionTargetPageId="60" captionText="FIGURE 40. Diagrams of apical views of the A2 of Benincandona, showing position of (sub-) terminal claws, setae and aesthetascs of the terminal (small circle) and penultimate segment (large circle). For explanations of codes, see Material and Methods (Abbreviations used in text and figures)." figureDoi="http://doi.org/10.5281/zenodo.13745804" httpUri="https://zenodo.org/record/13745804/files/figure.png" pageId="43" pageNumber="44">Figure 40</figureCitation>
): ventro-apically with one slender aesthetasc y2 and claw G3 transformed into short seta of almost equal length; centrally with one short plumose seta z3; dorso-apically, z1 claw-like, length about 1.7x as long as penultimate segment. Claw G2 about 2.5 times as long as z1 with G1 slightly shorter; z2 seta transformed into long claw, reaching tip of G1. Fourth, terminal segment (segment 3) with one long claw (Gm), one short claw (GM), a long aesthetasc y3 with shorter accompanying seta, with fused part short, and a seta g (slightly exceeding tip of aesthetasc y3).
</paragraph>
<paragraph id="7209B62C451EFF97FF111EA1FAA52B18" blockId="43.[151,1437,151,2013]" pageId="43" pageNumber="44">
MdCoxa (
<figureCitation id="EA8DAAA9451EFF97FEED1EA1FE4E2B3C" box="[315,448,1519,1545]" captionStart="FIGURE 27" captionStartId="45.[151,250,1943,1967]" captionTargetBox="[173,1414,183,1915]" captionTargetId="figure-17@45.[171,1417,181,1919]" captionTargetPageId="45" captionText="FIGURE 27. Benincandona cebios gen. et spec. nov., limbs (♂). A. A2 (INV.159244). B. A1 (INV.159244). C. MdCoxa (INV.159248). Scale bar = 50 µm." pageId="43" pageNumber="44">Figure 27C</figureCitation>
<figureCitation id="EA8DAAA9451EFF97FEED1EA1FE4E2B3C" box="[315,448,1519,1545]" captionStart="FIGURE 27" captionStartId="45.[151,250,1943,1967]" captionTargetBox="[173,1414,183,1915]" captionTargetId="figure-17@45.[171,1417,181,1919]" captionTargetPageId="45" captionText="FIGURE 27. Benincandona cebios gen. et spec. nov., limbs (♂). A. A2 (INV.159244). B. A1 (INV.159244). C. MdCoxa (INV.159248). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745776" httpUri="https://zenodo.org/record/13745776/files/figure.png" pageId="43" pageNumber="44">Figure 27C</figureCitation>
) elongated, proximally sharply pointed, medially widened, distally with an obliquely widened end, set with a series (67) of strong teeth, interspaced with setae; subapically with a short seta.
</paragraph>
<paragraph id="7209B62C451EFF97FF111D79FAEA2A78" blockId="43.[151,1437,151,2013]" pageId="43" pageNumber="44">
MdPalp (
<figureCitation id="EA8DAAA9451EFF97FEF91D79FE3F2B64" box="[303,433,1591,1617]" captionStart="FIGURE 28" captionStartId="46.[151,250,1943,1967]" captionTargetBox="[188,1401,185,1915]" captionTargetId="figure-31@46.[184,1403,181,1919]" captionTargetPageId="46" captionText="FIGURE 28. Benincandona cebios gen. et spec. nov., limbs (♂). A. Mx1-palp (INV.159244). B. Mx1, respiratory plate (INV.159248). C. MdPalp (INV.159244). Scale bars. 1 (B), 2 (A), 3 (C) = 50 µm." pageId="43" pageNumber="44">Figure 28C</figureCitation>
<figureCitation id="EA8DAAA9451EFF97FEF91D79FE3F2B64" box="[303,433,1591,1617]" captionStart="FIGURE 28" captionStartId="46.[151,250,1943,1967]" captionTargetBox="[188,1401,185,1915]" captionTargetId="figure-31@46.[184,1403,181,1919]" captionTargetPageId="46" captionText="FIGURE 28. Benincandona cebios gen. et spec. nov., limbs (♂). A. Mx1-palp (INV.159244). B. Mx1, respiratory plate (INV.159248). C. MdPalp (INV.159244). Scale bars. 1 (B), 2 (A), 3 (C) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745778" httpUri="https://zenodo.org/record/13745778/files/figure.png" pageId="43" pageNumber="44">Figure 28C</figureCitation>
) with four segments. Basal segment ventro-apically with two hirsute setae (S1 &amp; S2); S1 long, reaching tip of terminal segment and S2 short and stout, length about one fifth of the length of S1; in between with the short flagellum-like alpha seta (thin and about twice the length of S2), and one smooth and medium size, more proximal seta. Second segment of sub-triangular shape, dorso-apically with two setae of unequal length; ventro-apically with a group of four long setae, one of these being the beta-seta. Third segment with two unequal dorso-subapical setae, two mid-dorsal setae, one of which being the gamma seta, and ventro-apically with one seta of medium length. Terminal segment sub-rectangular, apically set with one large claw, fused with the segment over its entire distal margin, accompanied by one smooth lateral seta on each side, slightly shorter than fused claw.
</paragraph>
<paragraph id="7209B62C451EFF97FF111C19FF6B2AE8" blockId="43.[151,1437,151,2013]" pageId="43" pageNumber="44">
Mx1 (
<figureCitation id="EA8DAAA9451EFF97FEDC1C19FE232A44" box="[266,429,1879,1905]" captionStart="FIGURE 28" captionStartId="46.[151,250,1943,1967]" captionTargetBox="[188,1401,185,1915]" captionTargetId="figure-31@46.[184,1403,181,1919]" captionTargetPageId="46" captionText="FIGURE 28. Benincandona cebios gen. et spec. nov., limbs (♂). A. Mx1-palp (INV.159244). B. Mx1, respiratory plate (INV.159248). C. MdPalp (INV.159244). Scale bars. 1 (B), 2 (A), 3 (C) = 50 µm." pageId="43" pageNumber="44">Figure 28A, B</figureCitation>
<figureCitation id="EA8DAAA9451EFF97FEDC1C19FE232A44" box="[266,429,1879,1905]" captionStart="FIGURE 28" captionStartId="46.[151,250,1943,1967]" captionTargetBox="[188,1401,185,1915]" captionTargetId="figure-31@46.[184,1403,181,1919]" captionTargetPageId="46" captionText="FIGURE 28. Benincandona cebios gen. et spec. nov., limbs (♂). A. Mx1-palp (INV.159244). B. Mx1, respiratory plate (INV.159248). C. MdPalp (INV.159244). Scale bars. 1 (B), 2 (A), 3 (C) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745778" httpUri="https://zenodo.org/record/13745778/files/figure.png" pageId="43" pageNumber="44">Figure 28A, B</figureCitation>
) with first palp segment carrying four long dorso-apical setae; terminal segment short and rectangular, carrying four distal setae, some slightly claw-like. Respiratory plate (
<figureCitation id="EA8DAAA9451EFF97FBE61C35FB3D2AA0" box="[1072,1203,1915,1941]" captionStart="FIGURE 28" captionStartId="46.[151,250,1943,1967]" captionTargetBox="[188,1401,185,1915]" captionTargetId="figure-31@46.[184,1403,181,1919]" captionTargetPageId="46" captionText="FIGURE 28. Benincandona cebios gen. et spec. nov., limbs (♂). A. Mx1-palp (INV.159244). B. Mx1, respiratory plate (INV.159248). C. MdPalp (INV.159244). Scale bars. 1 (B), 2 (A), 3 (C) = 50 µm." pageId="43" pageNumber="44">Figure 28B</figureCitation>
<figureCitation id="EA8DAAA9451EFF97FBE61C35FB3D2AA0" box="[1072,1203,1915,1941]" captionStart="FIGURE 28" captionStartId="46.[151,250,1943,1967]" captionTargetBox="[188,1401,185,1915]" captionTargetId="figure-31@46.[184,1403,181,1919]" captionTargetPageId="46" captionText="FIGURE 28. Benincandona cebios gen. et spec. nov., limbs (♂). A. Mx1-palp (INV.159244). B. Mx1, respiratory plate (INV.159248). C. MdPalp (INV.159244). Scale bars. 1 (B), 2 (A), 3 (C) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745778" httpUri="https://zenodo.org/record/13745778/files/figure.png" pageId="43" pageNumber="44">Figure 28B</figureCitation>
) elongated, carrying
<emphasis id="40C26A3E451EFF97FF411CD1FF242A8D" box="[151,170,1951,1976]" italics="true" pageId="43" pageNumber="44">c.</emphasis>
16 plumose respiratory rays, two short distal setae and a group of about seven shorter proximal setae of unequal length.
</paragraph>
<caption id="26C9E6A44519FF90FF411DC7FC172BF3" pageId="44" pageNumber="45" startId="44.[151,250,1673,1697]" targetBox="[151,1436,181,1649]" targetPageId="44" targetType="figure">
<caption id="26C9E6A44519FF90FF411DC7FC172BF3" ID-DOI="http://doi.org/10.5281/zenodo.13745774" ID-Zenodo-Dep="13745774" httpUri="https://zenodo.org/record/13745774/files/figure.png" pageId="44" pageNumber="45" startId="44.[151,250,1673,1697]" targetBox="[151,1436,181,1649]" targetPageId="44" targetType="figure">
<paragraph id="7209B62C4519FF90FF411DC7FC172BF3" blockId="44.[151,1437,1673,1734]" pageId="44" pageNumber="45">
<emphasis id="40C26A3E4519FF90FF411DC7FEAE2B94" bold="true" box="[151,288,1673,1697]" pageId="44" pageNumber="45">FIGURE 26.</emphasis>
<taxonomicName id="B5B6CDAF4519FF90FEF11DC7FD8F2B94" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[295,513,1673,1697]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="cebios" status="gen. et spec. nov.">
@ -327,16 +328,16 @@ CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm.
</caption>
<paragraph id="7209B62C4519FF90FF111C56FB2B2AF7" blockId="44.[151,1436,1816,1986]" pageId="44" pageNumber="45">
T1 (
<figureCitation id="EA8DAAA94519FF90FF231C56FE142A07" box="[245,410,1816,1842]" captionStart="FIGURE 31" captionStartId="49.[151,250,1943,1967]" captionTargetBox="[184,1406,184,1915]" captionTargetId="figure-17@49.[180,1407,181,1919]" captionTargetPageId="49" captionText="FIGURE 31. Benincandona cebios gen. et spec. nov., limbs (AB ♂ and CE ♀). C. A2 (INV.159245). D. CR (INV.159245). E. T1 (INV.159245). A. T1, Lpp (INV.159244). B. T1, Rpp (INV.159244). Scale bar = 50 µm." pageId="44" pageNumber="45">Figure 31A, B</figureCitation>
<figureCitation id="EA8DAAA94519FF90FF231C56FE142A07" box="[245,410,1816,1842]" captionStart="FIGURE 31" captionStartId="49.[151,250,1943,1967]" captionTargetBox="[184,1406,184,1915]" captionTargetId="figure-17@49.[180,1407,181,1919]" captionTargetPageId="49" captionText="FIGURE 31. Benincandona cebios gen. et spec. nov., limbs (AB ♂ and CE ♀). C. A2 (INV.159245). D. CR (INV.159245). E. T1 (INV.159245). A. T1, Lpp (INV.159244). B. T1, Rpp (INV.159244). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745784" httpUri="https://zenodo.org/record/13745784/files/figure.png" pageId="44" pageNumber="45">Figure 31A, B</figureCitation>
) consisting of basal part (protopodite), carrying respiratory plates (exopodites) represented by three plumose setae, two short and one medium length. Protopodite (
<figureCitation id="EA8DAAA94519FF90FC011C72FBD42A63" box="[983,1114,1852,1878]" captionStart="FIGURE 31" captionStartId="49.[151,250,1943,1967]" captionTargetBox="[184,1406,184,1915]" captionTargetId="figure-17@49.[180,1407,181,1919]" captionTargetPageId="49" captionText="FIGURE 31. Benincandona cebios gen. et spec. nov., limbs (AB ♂ and CE ♀). C. A2 (INV.159245). D. CR (INV.159245). E. T1 (INV.159245). A. T1, Lpp (INV.159244). B. T1, Rpp (INV.159244). Scale bar = 50 µm." pageId="44" pageNumber="45">Figure 31B</figureCitation>
<figureCitation id="EA8DAAA94519FF90FC011C72FBD42A63" box="[983,1114,1852,1878]" captionStart="FIGURE 31" captionStartId="49.[151,250,1943,1967]" captionTargetBox="[184,1406,184,1915]" captionTargetId="figure-17@49.[180,1407,181,1919]" captionTargetPageId="49" captionText="FIGURE 31. Benincandona cebios gen. et spec. nov., limbs (AB ♂ and CE ♀). C. A2 (INV.159245). D. CR (INV.159245). E. T1 (INV.159245). A. T1, Lpp (INV.159244). B. T1, Rpp (INV.159244). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745784" httpUri="https://zenodo.org/record/13745784/files/figure.png" pageId="44" pageNumber="45">Figure 31B</figureCitation>
) distally set with almost 10 plumose setae of different morphology and length; more proximally set with d, d and b setae. Prehensile palps asymmetrical with Rpp (
<figureCitation id="EA8DAAA94519FF90FE711CCAFDA92AAB" box="[423,551,1924,1950]" captionStart="FIGURE 31" captionStartId="49.[151,250,1943,1967]" captionTargetBox="[184,1406,184,1915]" captionTargetId="figure-17@49.[180,1407,181,1919]" captionTargetPageId="49" captionText="FIGURE 31. Benincandona cebios gen. et spec. nov., limbs (AB ♂ and CE ♀). C. A2 (INV.159245). D. CR (INV.159245). E. T1 (INV.159245). A. T1, Lpp (INV.159244). B. T1, Rpp (INV.159244). Scale bar = 50 µm." pageId="44" pageNumber="45">Figure 31B</figureCitation>
<figureCitation id="EA8DAAA94519FF90FE711CCAFDA92AAB" box="[423,551,1924,1950]" captionStart="FIGURE 31" captionStartId="49.[151,250,1943,1967]" captionTargetBox="[184,1406,184,1915]" captionTargetId="figure-17@49.[180,1407,181,1919]" captionTargetPageId="49" captionText="FIGURE 31. Benincandona cebios gen. et spec. nov., limbs (AB ♂ and CE ♀). C. A2 (INV.159245). D. CR (INV.159245). E. T1 (INV.159245). A. T1, Lpp (INV.159244). B. T1, Rpp (INV.159244). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745784" httpUri="https://zenodo.org/record/13745784/files/figure.png" pageId="44" pageNumber="45">Figure 31B</figureCitation>
) slightly more swollen than Lpp (
<figureCitation id="EA8DAAA94519FF90FC4A1CCAFBAE2AAB" box="[924,1056,1924,1950]" captionStart="FIGURE 31" captionStartId="49.[151,250,1943,1967]" captionTargetBox="[184,1406,184,1915]" captionTargetId="figure-17@49.[180,1407,181,1919]" captionTargetPageId="49" captionText="FIGURE 31. Benincandona cebios gen. et spec. nov., limbs (AB ♂ and CE ♀). C. A2 (INV.159245). D. CR (INV.159245). E. T1 (INV.159245). A. T1, Lpp (INV.159244). B. T1, Rpp (INV.159244). Scale bar = 50 µm." pageId="44" pageNumber="45">Figure 31A</figureCitation>
<figureCitation id="EA8DAAA94519FF90FC4A1CCAFBAE2AAB" box="[924,1056,1924,1950]" captionStart="FIGURE 31" captionStartId="49.[151,250,1943,1967]" captionTargetBox="[184,1406,184,1915]" captionTargetId="figure-17@49.[180,1407,181,1919]" captionTargetPageId="49" captionText="FIGURE 31. Benincandona cebios gen. et spec. nov., limbs (AB ♂ and CE ♀). C. A2 (INV.159245). D. CR (INV.159245). E. T1 (INV.159245). A. T1, Lpp (INV.159244). B. T1, Rpp (INV.159244). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745784" httpUri="https://zenodo.org/record/13745784/files/figure.png" pageId="44" pageNumber="45">Figure 31A</figureCitation>
); one-segmented, distally sigmoid hook-like, both palps set with a short sensory organ and proximally with two subequal setae.
</paragraph>
<caption id="26C9E6A44518FF91FF411CD9FE7C2AE1" pageId="45" pageNumber="46" startId="45.[151,250,1943,1967]" targetBox="[173,1414,183,1915]" targetPageId="45" targetType="figure">
<caption id="26C9E6A44518FF91FF411CD9FE7C2AE1" ID-DOI="http://doi.org/10.5281/zenodo.13745776" ID-Zenodo-Dep="13745776" httpUri="https://zenodo.org/record/13745776/files/figure.png" pageId="45" pageNumber="46" startId="45.[151,250,1943,1967]" targetBox="[173,1414,183,1915]" targetPageId="45" targetType="figure">
<paragraph id="7209B62C4518FF91FF411CD9FE7C2AE1" blockId="45.[151,1437,1943,2004]" pageId="45" pageNumber="46">
<emphasis id="40C26A3E4518FF91FF411CD9FEAA2A9A" bold="true" box="[151,292,1943,1967]" pageId="45" pageNumber="46">FIGURE 27.</emphasis>
<taxonomicName id="B5B6CDAF4518FF91FEFB1CD6FD842A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[301,522,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="cebios" status="gen. et spec. nov.">
@ -352,7 +353,7 @@ T1 (
. MdCoxa (INV.159248). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A4451BFF92FF411CD9FC5F2AE1" pageId="46" pageNumber="47" startId="46.[151,250,1943,1967]" targetBox="[188,1401,185,1915]" targetPageId="46" targetType="figure">
<caption id="26C9E6A4451BFF92FF411CD9FC5F2AE1" ID-DOI="http://doi.org/10.5281/zenodo.13745778" ID-Zenodo-Dep="13745778" httpUri="https://zenodo.org/record/13745778/files/figure.png" pageId="46" pageNumber="47" startId="46.[151,250,1943,1967]" targetBox="[188,1401,185,1915]" targetPageId="46" targetType="figure">
<paragraph id="7209B62C451BFF92FF411CD9FC5F2AE1" blockId="46.[151,1436,1943,2004]" pageId="46" pageNumber="47">
<emphasis id="40C26A3E451BFF92FF411CD9FEAB2A9A" bold="true" box="[151,293,1943,1967]" pageId="46" pageNumber="47">FIGURE 28.</emphasis>
<taxonomicName id="B5B6CDAF451BFF92FEF91CD6FD802A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[303,526,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="cebios" status="gen. et spec. nov.">
@ -368,7 +369,7 @@ T1 (
. MdPalp (INV.159244). Scale bars. 1 (B), 2 (A), 3 (C) = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A4451AFF93FF411CF3FD5D2ACF" pageId="47" pageNumber="48" startId="47.[151,250,1981,2005]" targetBox="[197,1421,196,1942]" targetPageId="47" targetType="figure">
<caption id="26C9E6A4451AFF93FF411CF3FD5D2ACF" ID-DOI="http://doi.org/10.5281/zenodo.13745780" ID-Zenodo-Dep="13745780" httpUri="https://zenodo.org/record/13745780/files/figure.png" pageId="47" pageNumber="48" startId="47.[151,250,1981,2005]" targetBox="[197,1421,196,1942]" targetPageId="47" targetType="figure">
<paragraph id="7209B62C451AFF93FF411CF3FD5D2ACF" blockId="47.[151,1436,1981,2042]" pageId="47" pageNumber="48">
<emphasis id="40C26A3E451AFF93FF411CF3FEAB2AE0" bold="true" box="[151,293,1981,2005]" pageId="47" pageNumber="48">FIGURE 29.</emphasis>
<taxonomicName id="B5B6CDAF451AFF93FEE61CF3FD812AE0" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[304,527,1981,2005]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="cebios" status="gen. et spec. nov.">
@ -386,7 +387,7 @@ T1 (
. CR (INV.159244). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A44505FF8CFF411CD9FDB32ACD" pageId="48" pageNumber="49" startId="48.[151,250,1943,1967]" targetBox="[194,1393,191,1910]" targetPageId="48" targetType="figure">
<caption id="26C9E6A44505FF8CFF411CD9FDB32ACD" ID-DOI="http://doi.org/10.5281/zenodo.13745782" ID-Zenodo-Dep="13745782" httpUri="https://zenodo.org/record/13745782/files/figure.png" pageId="48" pageNumber="49" startId="48.[151,250,1943,1967]" targetBox="[194,1393,191,1910]" targetPageId="48" targetType="figure">
<paragraph id="7209B62C4505FF8CFF411CD9FDB32ACD" blockId="48.[151,1437,1943,2040]" pageId="48" pageNumber="49">
<emphasis id="40C26A3E4505FF8CFF411CD9FEAD2A9A" bold="true" box="[151,291,1943,1967]" pageId="48" pageNumber="49">FIGURE 30.</emphasis>
<taxonomicName id="B5B6CDAF4505FF8CFEFA1CD6FD872A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[300,521,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="cebios" status="gen. et spec. nov.">
@ -406,7 +407,7 @@ T1 (
. Internal view (photograph with interference contrast). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A44504FF8DFF411CD9FBDC2AE1" pageId="49" pageNumber="50" startId="49.[151,250,1943,1967]" targetBox="[184,1406,184,1915]" targetPageId="49" targetType="figure">
<caption id="26C9E6A44504FF8DFF411CD9FBDC2AE1" ID-DOI="http://doi.org/10.5281/zenodo.13745784" ID-Zenodo-Dep="13745784" httpUri="https://zenodo.org/record/13745784/files/figure.png" pageId="49" pageNumber="50" startId="49.[151,250,1943,1967]" targetBox="[184,1406,184,1915]" targetPageId="49" targetType="figure">
<paragraph id="7209B62C4504FF8DFF411CD9FBDC2AE1" blockId="49.[151,1437,1943,2004]" pageId="49" pageNumber="50">
<emphasis id="40C26A3E4504FF8DFF411CD9FEAE2A9A" bold="true" box="[151,288,1943,1967]" pageId="49" pageNumber="50">FIGURE 31.</emphasis>
<taxonomicName id="B5B6CDAF4504FF8DFEF11CD6FD8F2A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[295,513,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="cebios" status="gen. et spec. nov.">
@ -428,36 +429,36 @@ T1 (
</caption>
<paragraph id="7209B62C4507FF8EFF111BD9FC692C50" blockId="50.[151,1437,151,1401]" pageId="50" pageNumber="51">
T2 (
<figureCitation id="EA8DAAA94507FF8EFF231BD9FEF92D84" box="[245,375,151,177]" captionStart="FIGURE 29" captionStartId="47.[151,250,1981,2005]" captionTargetBox="[197,1421,196,1942]" captionTargetId="figure-17@47.[166,1422,181,1957]" captionTargetPageId="47" captionText="FIGURE 29. Benincandona cebios gen. et spec. nov., limbs (♂). A. T3 (INV.159244). B. T2 (INV.159244). C. CRAtt (INV.159244). D. CR (INV.159244). Scale bar = 50 µm." pageId="50" pageNumber="51">Figure 29B</figureCitation>
<figureCitation id="EA8DAAA94507FF8EFF231BD9FEF92D84" box="[245,375,151,177]" captionStart="FIGURE 29" captionStartId="47.[151,250,1981,2005]" captionTargetBox="[197,1421,196,1942]" captionTargetId="figure-17@47.[166,1422,181,1957]" captionTargetPageId="47" captionText="FIGURE 29. Benincandona cebios gen. et spec. nov., limbs (♂). A. T3 (INV.159244). B. T2 (INV.159244). C. CRAtt (INV.159244). D. CR (INV.159244). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745780" httpUri="https://zenodo.org/record/13745780/files/figure.png" pageId="50" pageNumber="51">Figure 29B</figureCitation>
) with 4-segmented endopodite (penultimate segment divided), generally elongated. First basal segment without setae d1 and d2. First segment of endopodite elongated (length almost 4x the basal width), without seta. Second endopodal segment (segment 2A), with length
<emphasis id="40C26A3E4507FF8EFCEC1B91FCC32DCD" box="[826,845,223,248]" italics="true" pageId="50" pageNumber="51">c.</emphasis>
3x basal width, set with one short ventro-apical seta f. Third endopodal segment (segment 2B) of about the same length/width ratio as the previous segment, ventro-subapically set with a g-seta. Terminal segment about 1.5 times as long as basal width, apically with two short setae (h1 and h3) and a long and slender claw (h2), distally set with short setulae.
</paragraph>
<paragraph id="7209B62C4507FF8EFF111A21FF642F08" blockId="50.[151,1437,151,1401]" pageId="50" pageNumber="51">
T3 (
<figureCitation id="EA8DAAA94507FF8EFF211A21FEF02CBC" box="[247,382,367,393]" captionStart="FIGURE 29" captionStartId="47.[151,250,1981,2005]" captionTargetBox="[197,1421,196,1942]" captionTargetId="figure-17@47.[166,1422,181,1957]" captionTargetPageId="47" captionText="FIGURE 29. Benincandona cebios gen. et spec. nov., limbs (♂). A. T3 (INV.159244). B. T2 (INV.159244). C. CRAtt (INV.159244). D. CR (INV.159244). Scale bar = 50 µm." pageId="50" pageNumber="51">Figure 29A</figureCitation>
<figureCitation id="EA8DAAA94507FF8EFF211A21FEF02CBC" box="[247,382,367,393]" captionStart="FIGURE 29" captionStartId="47.[151,250,1981,2005]" captionTargetBox="[197,1421,196,1942]" captionTargetId="figure-17@47.[166,1422,181,1957]" captionTargetPageId="47" captionText="FIGURE 29. Benincandona cebios gen. et spec. nov., limbs (♂). A. T3 (INV.159244). B. T2 (INV.159244). C. CRAtt (INV.159244). D. CR (INV.159244). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745780" httpUri="https://zenodo.org/record/13745780/files/figure.png" pageId="50" pageNumber="51">Figure 29A</figureCitation>
) with first segment carrying only one subapical seta d1, of medium length. Second (knee-) segment with short seta d2. Third segment about 5x as long as wide, not carrying any seta. Fourth segment narrower and at least 5x basal width, carrying one short subapical seta g. Terminal segment well-separated from penultimate segment, length about twice as long as basal width, carrying three setae: one long and reflexed (h3), almost reaching knee-segment and two short ventrally directed and curved setae h1 &amp; h2, the latter slightly exceeding tip of the former.
</paragraph>
<paragraph id="7209B62C4507FF8EFF111909FC862F9C" blockId="50.[151,1437,151,1401]" pageId="50" pageNumber="51">
CR (
<figureCitation id="EA8DAAA94507FF8EFF2C1909FEF32F54" box="[250,381,583,609]" captionStart="FIGURE 29" captionStartId="47.[151,250,1981,2005]" captionTargetBox="[197,1421,196,1942]" captionTargetId="figure-17@47.[166,1422,181,1957]" captionTargetPageId="47" captionText="FIGURE 29. Benincandona cebios gen. et spec. nov., limbs (♂). A. T3 (INV.159244). B. T2 (INV.159244). C. CRAtt (INV.159244). D. CR (INV.159244). Scale bar = 50 µm." pageId="50" pageNumber="51">Figure 29D</figureCitation>
<figureCitation id="EA8DAAA94507FF8EFF2C1909FEF32F54" box="[250,381,583,609]" captionStart="FIGURE 29" captionStartId="47.[151,250,1981,2005]" captionTargetBox="[197,1421,196,1942]" captionTargetId="figure-17@47.[166,1422,181,1957]" captionTargetPageId="47" captionText="FIGURE 29. Benincandona cebios gen. et spec. nov., limbs (♂). A. T3 (INV.159244). B. T2 (INV.159244). C. CRAtt (INV.159244). D. CR (INV.159244). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745780" httpUri="https://zenodo.org/record/13745780/files/figure.png" pageId="50" pageNumber="51">Figure 29D</figureCitation>
) with stout ramus; claw Ga well-developed, stout with length slightly exceeding half the length of ramus, setae Sp, Sa and claw Gp reduced to small setae; seta Sp situated about at one-fifth length of ramus from distal end, Sa dorso-subapical and Gp ventro-subapical.
</paragraph>
<paragraph id="7209B62C4507FF8EFF1119FDFA832FF8" blockId="50.[151,1437,151,1401]" box="[199,1293,691,717]" pageId="50" pageNumber="51">
CRAtt (
<figureCitation id="EA8DAAA94507FF8EFEC919FDFE2F2FF8" box="[287,417,691,717]" captionStart="FIGURE 29" captionStartId="47.[151,250,1981,2005]" captionTargetBox="[197,1421,196,1942]" captionTargetId="figure-17@47.[166,1422,181,1957]" captionTargetPageId="47" captionText="FIGURE 29. Benincandona cebios gen. et spec. nov., limbs (♂). A. T3 (INV.159244). B. T2 (INV.159244). C. CRAtt (INV.159244). D. CR (INV.159244). Scale bar = 50 µm." pageId="50" pageNumber="51">Figure 29C</figureCitation>
<figureCitation id="EA8DAAA94507FF8EFEC919FDFE2F2FF8" box="[287,417,691,717]" captionStart="FIGURE 29" captionStartId="47.[151,250,1981,2005]" captionTargetBox="[197,1421,196,1942]" captionTargetId="figure-17@47.[166,1422,181,1957]" captionTargetPageId="47" captionText="FIGURE 29. Benincandona cebios gen. et spec. nov., limbs (♂). A. T3 (INV.159244). B. T2 (INV.159244). C. CRAtt (INV.159244). D. CR (INV.159244). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745780" httpUri="https://zenodo.org/record/13745780/files/figure.png" pageId="50" pageNumber="51">Figure 29C</figureCitation>
) long and stout, distally bifurcated, subapically with additional lateral branch.
</paragraph>
<paragraph id="7209B62C4507FF8EFF111999FD772E0C" blockId="50.[151,1437,151,1401]" pageId="50" pageNumber="51">
Hp (
<figureCitation id="EA8DAAA94507FF8EFF2C1999FE2E2FC4" box="[250,416,727,753]" captionStart="FIGURE 30" captionStartId="48.[151,250,1943,1967]" captionTargetBox="[194,1393,191,1910]" captionTargetId="figure-31@48.[184,1403,181,1920]" captionTargetPageId="48" captionText="FIGURE 30. Benincandona cebios gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159247). BD. Hp (INV.159244). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." pageId="50" pageNumber="51">Figure 30BD</figureCitation>
<figureCitation id="EA8DAAA94507FF8EFF2C1999FE2E2FC4" box="[250,416,727,753]" captionStart="FIGURE 30" captionStartId="48.[151,250,1943,1967]" captionTargetBox="[194,1393,191,1910]" captionTargetId="figure-31@48.[184,1403,181,1920]" captionTargetPageId="48" captionText="FIGURE 30. Benincandona cebios gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159247). BD. Hp (INV.159244). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745782" httpUri="https://zenodo.org/record/13745782/files/figure.png" pageId="50" pageNumber="51">Figure 30BD</figureCitation>
) with robust lobe ls, apically rounded, reaching well beyond symmetrically rounded lobe ms, both lobes overlapping, not diverging; lobe h shorter and sub-rectangular; internal anatomy (
<figureCitation id="EA8DAAA94507FF8EFB3819B5FA1A2E20" box="[1262,1428,763,789]" captionStart="FIGURE 30" captionStartId="48.[151,250,1943,1967]" captionTargetBox="[194,1393,191,1910]" captionTargetId="figure-31@48.[184,1403,181,1920]" captionTargetPageId="48" captionText="FIGURE 30. Benincandona cebios gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159247). BD. Hp (INV.159244). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." pageId="50" pageNumber="51">Figure 30C, D</figureCitation>
<figureCitation id="EA8DAAA94507FF8EFB3819B5FA1A2E20" box="[1262,1428,763,789]" captionStart="FIGURE 30" captionStartId="48.[151,250,1943,1967]" captionTargetBox="[194,1393,191,1910]" captionTargetId="figure-31@48.[184,1403,181,1920]" captionTargetPageId="48" captionText="FIGURE 30. Benincandona cebios gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159247). BD. Hp (INV.159244). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745782" httpUri="https://zenodo.org/record/13745782/files/figure.png" pageId="50" pageNumber="51">Figure 30C, D</figureCitation>
) with postlabyrinthal spermiduct with one circular coil.
</paragraph>
<paragraph id="7209B62C4507FF8EFF11180DFBDD2E68" blockId="50.[151,1437,151,1401]" box="[199,1107,835,861]" pageId="50" pageNumber="51">
Zenker organ (
<figureCitation id="EA8DAAA94507FF8EFEBD180DFE612E68" box="[363,495,835,861]" captionStart="FIGURE 30" captionStartId="48.[151,250,1943,1967]" captionTargetBox="[194,1393,191,1910]" captionTargetId="figure-31@48.[184,1403,181,1920]" captionTargetPageId="48" captionText="FIGURE 30. Benincandona cebios gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159247). BD. Hp (INV.159244). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." pageId="50" pageNumber="51">Figure 30A</figureCitation>
<figureCitation id="EA8DAAA94507FF8EFEBD180DFE612E68" box="[363,495,835,861]" captionStart="FIGURE 30" captionStartId="48.[151,250,1943,1967]" captionTargetBox="[194,1393,191,1910]" captionTargetId="figure-31@48.[184,1403,181,1920]" captionTargetPageId="48" captionText="FIGURE 30. Benincandona cebios gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159247). BD. Hp (INV.159244). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745782" httpUri="https://zenodo.org/record/13745782/files/figure.png" pageId="50" pageNumber="51">Figure 30A</figureCitation>
) long and slightly broad, with
<emphasis id="40C26A3E4507FF8EFC91180DFCD52E69" box="[839,859,835,860]" italics="true" pageId="50" pageNumber="51">c.</emphasis>
seven spinous whirls.
@ -467,26 +468,26 @@ seven spinous whirls.
</paragraph>
<paragraph id="7209B62C4507FF8EFF1118C5FB422E90" blockId="50.[151,1437,151,1401]" box="[199,1228,907,933]" pageId="50" pageNumber="51">
Cp and valves (
<figureCitation id="EA8DAAA94507FF8EFEA218C5FD952E90" box="[372,539,907,933]" captionStart="FIGURE 24" captionStartId="41.[151,250,1944,1968]" captionTargetBox="[170,1417,181,1921]" captionTargetId="figure-17@41.[170,1417,181,1921]" captionTargetPageId="41" captionText="FIGURE 24. Benincandona cebios gen. et spec. nov., valves (AC, JK ♂; DI ♀). AI. CpRl (AC. INV.159255, DI. INV.159258). J. INV.159256. CpD. K. INV.159257. CpV. Scale bars. 1 (B, E, J, K) = 500 µm; 2 (A, C, D, F, H, I) = 200 µm." pageId="50" pageNumber="51">Figures 24DI</figureCitation>
<figureCitation id="EA8DAAA94507FF8EFEA218C5FD952E90" box="[372,539,907,933]" captionStart="FIGURE 24" captionStartId="41.[151,250,1944,1968]" captionTargetBox="[170,1417,181,1921]" captionTargetId="figure-17@41.[170,1417,181,1921]" captionTargetPageId="41" captionText="FIGURE 24. Benincandona cebios gen. et spec. nov., valves (AC, JK ♂; DI ♀). AI. CpRl (AC. INV.159255, DI. INV.159258). J. INV.159256. CpD. K. INV.159257. CpV. Scale bars. 1 (B, E, J, K) = 500 µm; 2 (A, C, D, F, H, I) = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.13745770" httpUri="https://zenodo.org/record/13745770/files/figure.png" pageId="50" pageNumber="51">Figures 24DI</figureCitation>
,
<figureCitation id="EA8DAAA94507FF8EFDF118C5FDF72E90" box="[551,633,907,933]" captionStart="FIGURE 26" captionStartId="44.[151,250,1673,1697]" captionTargetBox="[151,1436,181,1649]" captionTargetId="figure-31@44.[151,1436,181,1649]" captionTargetPageId="44" captionText="FIGURE 26. Benincandona cebios gen. et spec. nov., valves (♀). A, C, D, G. LVi (INV.159253). B, E, F. RVi (INV.159253). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." pageId="50" pageNumber="51">26AG</figureCitation>
<figureCitation id="EA8DAAA94507FF8EFDF118C5FDF72E90" box="[551,633,907,933]" captionStart="FIGURE 26" captionStartId="44.[151,250,1673,1697]" captionTargetBox="[151,1436,181,1649]" captionTargetId="figure-31@44.[151,1436,181,1649]" captionTargetPageId="44" captionText="FIGURE 26. Benincandona cebios gen. et spec. nov., valves (♀). A, C, D, G. LVi (INV.159253). B, E, F. RVi (INV.159253). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745774" httpUri="https://zenodo.org/record/13745774/files/figure.png" pageId="50" pageNumber="51">26AG</figureCitation>
) as in the male, without obvious sexual dimorphism.
</paragraph>
<paragraph id="7209B62C4507FF8EFF1118E1FB072EFC" blockId="50.[151,1437,151,1401]" box="[199,1161,943,969]" pageId="50" pageNumber="51">
Limbs (
<figureCitation id="EA8DAAA94507FF8EFECB18E1FE302EFC" box="[285,446,943,969]" captionStart="FIGURE 31" captionStartId="49.[151,250,1943,1967]" captionTargetBox="[184,1406,184,1915]" captionTargetId="figure-17@49.[180,1407,181,1919]" captionTargetPageId="49" captionText="FIGURE 31. Benincandona cebios gen. et spec. nov., limbs (AB ♂ and CE ♀). C. A2 (INV.159245). D. CR (INV.159245). E. T1 (INV.159245). A. T1, Lpp (INV.159244). B. T1, Rpp (INV.159244). Scale bar = 50 µm." pageId="50" pageNumber="51">Figure 31CE</figureCitation>
<figureCitation id="EA8DAAA94507FF8EFECB18E1FE302EFC" box="[285,446,943,969]" captionStart="FIGURE 31" captionStartId="49.[151,250,1943,1967]" captionTargetBox="[184,1406,184,1915]" captionTargetId="figure-17@49.[180,1407,181,1919]" captionTargetPageId="49" captionText="FIGURE 31. Benincandona cebios gen. et spec. nov., limbs (AB ♂ and CE ♀). C. A2 (INV.159245). D. CR (INV.159245). E. T1 (INV.159245). A. T1, Lpp (INV.159244). B. T1, Rpp (INV.159244). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745784" httpUri="https://zenodo.org/record/13745784/files/figure.png" pageId="50" pageNumber="51">Figure 31CE</figureCitation>
) largely as in the male, but with sexually dimorphic A2 and T1.
</paragraph>
<paragraph id="7209B62C4507FF8EFF11189DFB53296C" blockId="50.[151,1437,151,1401]" pageId="50" pageNumber="51">
A2 (
<figureCitation id="EA8DAAA94507FF8EFF2F189DFEF22ED8" box="[249,380,979,1005]" captionStart="FIGURE 31" captionStartId="49.[151,250,1943,1967]" captionTargetBox="[184,1406,184,1915]" captionTargetId="figure-17@49.[180,1407,181,1919]" captionTargetPageId="49" captionText="FIGURE 31. Benincandona cebios gen. et spec. nov., limbs (AB ♂ and CE ♀). C. A2 (INV.159245). D. CR (INV.159245). E. T1 (INV.159245). A. T1, Lpp (INV.159244). B. T1, Rpp (INV.159244). Scale bar = 50 µm." pageId="50" pageNumber="51">Figure 31C</figureCitation>
<figureCitation id="EA8DAAA94507FF8EFF2F189DFEF22ED8" box="[249,380,979,1005]" captionStart="FIGURE 31" captionStartId="49.[151,250,1943,1967]" captionTargetBox="[184,1406,184,1915]" captionTargetId="figure-17@49.[180,1407,181,1919]" captionTargetPageId="49" captionText="FIGURE 31. Benincandona cebios gen. et spec. nov., limbs (AB ♂ and CE ♀). C. A2 (INV.159245). D. CR (INV.159245). E. T1 (INV.159245). A. T1, Lpp (INV.159244). B. T1, Rpp (INV.159244). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745784" httpUri="https://zenodo.org/record/13745784/files/figure.png" pageId="50" pageNumber="51">Figure 31C</figureCitation>
) with second endopodal segment undivided, with t1-seta (long, narrow and plumose) and t2- seta (thin and smooth); apical chaetotaxy (
<figureCitation id="EA8DAAA94507FF8EFDA418B9FD6C2924" box="[626,738,1015,1041]" captionStart="FIGURE 40" captionStartId="60.[151,250,1516,1540]" captionTargetBox="[151,1435,181,1492]" captionTargetId="figure-31@60.[151,1435,181,1492]" captionTargetPageId="60" captionText="FIGURE 40. Diagrams of apical views of the A2 of Benincandona, showing position of (sub-) terminal claws, setae and aesthetascs of the terminal (small circle) and penultimate segment (large circle). For explanations of codes, see Material and Methods (Abbreviations used in text and figures)." pageId="50" pageNumber="51">Figure 40</figureCitation>
<figureCitation id="EA8DAAA94507FF8EFDA418B9FD6C2924" box="[626,738,1015,1041]" captionStart="FIGURE 40" captionStartId="60.[151,250,1516,1540]" captionTargetBox="[151,1435,181,1492]" captionTargetId="figure-31@60.[151,1435,181,1492]" captionTargetPageId="60" captionText="FIGURE 40. Diagrams of apical views of the A2 of Benincandona, showing position of (sub-) terminal claws, setae and aesthetascs of the terminal (small circle) and penultimate segment (large circle). For explanations of codes, see Material and Methods (Abbreviations used in text and figures)." figureDoi="http://doi.org/10.5281/zenodo.13745804" httpUri="https://zenodo.org/record/13745804/files/figure.png" pageId="50" pageNumber="51">Figure 40</figureCitation>
) with z1 claw-like, z2 and z3 being both setae, z1 about twice as long as z2, and z3 slightly shorter than z2; claws G1, G2, G3 and GM all reaching about the same point; seta g and aesthetasc y3 as in the male. Small claw Gm in female homologous to large claw in the male.
</paragraph>
<paragraph id="7209B62C4507FF8EFF111F2DFBDF29F0" blockId="50.[151,1437,151,1401]" pageId="50" pageNumber="51">
T1 (
<figureCitation id="EA8DAAA94507FF8EFF221F2DFEFA2948" box="[244,372,1123,1149]" captionStart="FIGURE 31" captionStartId="49.[151,250,1943,1967]" captionTargetBox="[184,1406,184,1915]" captionTargetId="figure-17@49.[180,1407,181,1919]" captionTargetPageId="49" captionText="FIGURE 31. Benincandona cebios gen. et spec. nov., limbs (AB ♂ and CE ♀). C. A2 (INV.159245). D. CR (INV.159245). E. T1 (INV.159245). A. T1, Lpp (INV.159244). B. T1, Rpp (INV.159244). Scale bar = 50 µm." pageId="50" pageNumber="51">Figure 31E</figureCitation>
<figureCitation id="EA8DAAA94507FF8EFF221F2DFEFA2948" box="[244,372,1123,1149]" captionStart="FIGURE 31" captionStartId="49.[151,250,1943,1967]" captionTargetBox="[184,1406,184,1915]" captionTargetId="figure-17@49.[180,1407,181,1919]" captionTargetPageId="49" captionText="FIGURE 31. Benincandona cebios gen. et spec. nov., limbs (AB ♂ and CE ♀). C. A2 (INV.159245). D. CR (INV.159245). E. T1 (INV.159245). A. T1, Lpp (INV.159244). B. T1, Rpp (INV.159244). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745784" httpUri="https://zenodo.org/record/13745784/files/figure.png" pageId="50" pageNumber="51">Figure 31E</figureCitation>
) with basal part with
<emphasis id="40C26A3E4507FF8EFDB21F2DFDF62949" box="[612,632,1123,1148]" italics="true" pageId="50" pageNumber="51">c.</emphasis>
10 distal setae of unequal length and morphology, setae d and b present, with long additional seta (d) present close to seta d; respiratory plate represented by two long and one short rays. Endopodite with a broad palp, carrying three subequal but short apical setae (h1h3).

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