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<document id="ABCE0E15E535EAF499FB44411C1A2DC2" ID-CLB-Dataset="310323" ID-DOI="10.1163/18759866-BJA10060" ID-GBIF-Dataset="814a05d4-fc17-4270-a6e5-e613b534360b" ID-ISSN="1875-9866" ID-Zenodo-Dep="13153846" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1720621781999" checkinUser="felipe" docAuthor="Arntzen, Jan W." docDate="2024" docId="03B4DA36EA09B45BFD44FE84E029FA7F" docLanguage="en" docName="ContribZool.93.3.252-267.pdf" docOrigin="Contributions to Zoology 93" docSource="http://dx.doi.org/10.1163/18759866-bja10060" docStyle="DocumentStyle:E1C3C5FD6CDDA08D62BFE928526826AF.3:ContribZool.2019-.journal_article" docStyleId="E1C3C5FD6CDDA08D62BFE928526826AF" docStyleName="ContribZool.2019-.journal_article" docStyleVersion="3" docTitle="Triturus marmoratus subsp. harmannis Arntzen, 2024, ssp. nov." docType="treatment" docVersion="2" lastPageNumber="267" masterDocId="FF8DA24EEA05B454FFC1FF95E36AFFD1" masterDocTitle="A subspecies of marbled newt (Triturus marmoratus) in the Iberian Peninsula newly resolved from congruent nuclear and mitochondrial DNA data" masterLastPageNumber="267" masterPageNumber="252" pageNumber="264" updateTime="1749481290728" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="3C97DA8DC4DC24093EA8FE18ECDF2DC0">A subspecies of marbled newt (Triturus marmoratus) in the Iberian Peninsula newly resolved from congruent nuclear and mitochondrial DNA data</mods:title>
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<mods:namePart id="4984B2A24D63878E486AEAF8264EE2D8">Arntzen, Jan W.</mods:namePart>
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<mods:affiliation id="127E3B75999F888ACECBF2A9D6F26410">Institute of Biology, Leiden University, 2333 BE Leiden, The Netherlands Naturalis Biodiversity Center, 2333 CR Leiden, The Netherlands</mods:affiliation>
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<mods:date id="A21FAFA3E2E81E1871D141679985965D">2024</mods:date>
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<emphasis id="B969B732EA09B458FD44FE84E046FEFA" bold="true" box="[645,812,273,299]" pageId="12" pageNumber="264">Description of</emphasis>
<taxonomicName id="4C1D10A3EA09B458FCF3FE87E1CFFE9E" authorityName="Arntzen" authorityYear="2024" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="12" pageNumber="264" phylum="Chordata" rank="subSpecies" species="marmoratus" status="subsp. nov." subSpecies="harmannis">
<emphasis id="B969B732EA09B458FCF3FE87E1CFFE9E" italics="true" pageId="12" pageNumber="264">Triturus marmoratus harmannis</emphasis>
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<emphasis id="B969B732EA09B458FD6AFEA3E060FE81" bold="true" box="[683,778,310,336]" pageId="12" pageNumber="264">
<taxonomicNameLabel id="A25A0A49EA09B458FD6AFEA3E060FE81" box="[683,778,310,336]" pageId="12" pageNumber="264" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
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<paragraph id="8BA26B20EA09B458FD44FE16E06FFDA5" blockId="12.[645,1157,387,1669]" pageId="12" pageNumber="264">
Descriptions were made from preserved material, except for the
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character state that was determined in the field. Dorsal colouration invariably dark, yet with the green colouration pattern discernible. Morphological data retrieved for a series of live animals are in
<tableCitation id="C69F5E9BEA09B458FD60FDCFE06AFDA5" box="[673,768,602,628]" captionStart="TABLE 1" captionStartId="5.[130,190,124,145]" captionTargetBox="[128,1187,306,1109]" captionTargetPageId="5" captionText="TABLE 1 Summary of morphometric data on males and females in two subspecies of Triturus marmoratus. Values to the left are averages of untransformed data in mm and to the right are the relative values obtained from division by snout-vent length (SVl1).The panel to the right gives the results of statistical testing on SVl1 and relative sizes, with in parentheses the identity of the group with significantly higher values." httpUri="http://table.plazi.org/id/DF623BA8EA00B451FF43FFE9E213FED6" pageId="12" pageNumber="264" tableUuid="DF623BA8EA00B451FF43FFE9E213FED6">Table A1</tableCitation>
.
</paragraph>
<paragraph id="8BA26B20EA09B458FD65FDEBE036FA27" blockId="12.[645,1157,387,1669]" pageId="12" pageNumber="264">
Description of type specimen preserved on ethanol in excellent storage condition at the Museo Nacional de Ciencias Naturales,
<collectingRegion id="49D9A5C2EA09B458FD44FD7FE1B0FCD5" box="[645,730,746,772]" country="Spain" name="Madrid" pageId="12" pageNumber="264">Madrid</collectingRegion>
,
<collectingCountry id="F30A2BB0EA09B458FD23FD7FE04BFCD5" box="[738,801,746,772]" name="Spain" pageId="12" pageNumber="264">Spain</collectingCountry>
under catalogue number MNCN 51792. Adult male in breeding condition with a well-developed crest and a large back cloaca. Thirteen crème-white bands over the head and body part of the crest. SVl1
<quantity id="4CE5C6C5EA09B458FBDEFCECE7EAFC45" box="[1055,1152,889,916]" metricMagnitude="-2" metricUnit="m" metricValue="6.45" pageId="12" pageNumber="264" unit="mm" value="64.5">64.5 mm</quantity>
, SVl2
<quantity id="4CE5C6C5EA09B458FD7EFC08E074FC69" box="[703,798,925,952]" metricMagnitude="-2" metricUnit="m" metricValue="7.35" pageId="12" pageNumber="264" unit="mm" value="73.5">73.5 mm</quantity>
, ILd
<quantity id="4CE5C6C5EA09B458FC9BFC08E0D3FC69" box="[858,953,925,952]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" pageId="12" pageNumber="264" unit="mm" value="32.5">32.5 mm</quantity>
, FLl
<quantity id="4CE5C6C5EA09B458FC32FC0BE739FC69" box="[1011,1107,926,952]" metricMagnitude="-2" metricUnit="m" metricValue="2.83" pageId="12" pageNumber="264" unit="mm" value="28.3">28.3 mm</quantity>
, TFl
<quantity id="4CE5C6C5EA09B458FD44FC54E1B3FC0D" box="[645,729,961,988]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="12" pageNumber="264" unit="mm" value="8.5">8.5 mm</quantity>
, HLl
<quantity id="4CE5C6C5EA09B458FCD1FC54E007FC0D" box="[784,877,961,988]" metricMagnitude="-2" metricUnit="m" metricValue="2.81" pageId="12" pageNumber="264" unit="mm" value="28.1">28.1 mm</quantity>
, FTl
<quantity id="4CE5C6C5EA09B458FC61FC54E093FC0D" box="[928,1017,961,988]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="12" pageNumber="264" unit="mm" value="11.6">11.6 mm</quantity>
, Hw
<quantity id="4CE5C6C5EA09B458FBEFFC54E7EEFC0D" box="[1070,1156,961,988]" metricMagnitude="-2" metricUnit="m" metricValue="1.41" pageId="12" pageNumber="264" unit="mm" value="14.1">14.1 mm</quantity>
and Hl
<quantity id="4CE5C6C5EA09B458FD10FC70E044FBD1" box="[721,814,997,1024]" metricMagnitude="-2" metricUnit="m" metricValue="1.97" pageId="12" pageNumber="264" unit="mm" value="19.7">19.7 mm</quantity>
.Total length
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.Relative finger lengths 1&lt;4&lt;2&lt;3. Relative toe lengths 1&lt;5&lt;2&lt;4&lt;3.
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left 0, right 0, total 0. Solid dark grey ventral colouration with many more or less evenly distributed white spots, denser in the middle part than towards the gular and cloacal regions. Throat region light grey with many small white dots. Underside of the legs and cloacal region with light and dark regions. The
<typeStatus id="54A6D582EA09B458FD7BFABDE074FA93" box="[698,798,1320,1346]" pageId="12" pageNumber="264" type="holotype">holotype</typeStatus>
is shown in fig. A1. A three-dimensional model of the entire skeleton of the type specimen obtained with CT-scanning is available for inspection at https://doi.org/10.6084/m9.figshare.25358890, courtesy of Dr. Tijana Vučić.
</paragraph>
<paragraph id="8BA26B20EA09B458FD65F995E040F954" blockId="12.[645,1157,387,1669]" pageId="12" pageNumber="264">
Description of
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Preserved on ethanol in excellent storage condition at the Museo Nacional de Ciencias Naturales,
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,
<collectingCountry id="F30A2BB0EA09B458FD25F9FEE04CF954" box="[740,806,1643,1669]" name="Spain" pageId="12" pageNumber="264">Spain</collectingCountry>
.
</paragraph>
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TABLE A1 Morphometric data for live adult
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<emphasis id="B969B732EA08B459FDEAFF33E03AFF6B" box="[555,848,165,187]" italics="true" pageId="13" pageNumber="265">Triturus marmoratus harmannis</emphasis>
</taxonomicName>
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<taxonomicNameLabel id="A25A0A49EA08B459FC94FF33E0C8FF6A" box="[853,930,166,187]" pageId="13" pageNumber="265" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
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from the type locality. Characters measured are described in the text.
</paragraph>
</caption>
<paragraph id="8BA26B20EA08B459FF40FE91E24DFAEC" pageId="13" pageNumber="265">
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<th id="76FC001EEA084BABFFBFFE91E7F4FECF" box="[126,1182,260,286]" colspan="12" colspanRight="11" gridcol="0" gridrow="0" pageId="13" pageNumber="265">
<emphasis id="B969B732EA08B459FF40FE91E3AFFECF" bold="true" box="[129,197,260,286]" pageId="13" pageNumber="265">Males</emphasis>
</th>
</tr>
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<th id="76FC001EEA084BABFFBFFED1E38BFEB6" box="[126,225,324,359]" gridcol="0" gridrow="1" pageId="13" pageNumber="265">
<emphasis id="B969B732EA08B459FF40FED0E38BFE8E" bold="true" box="[129,225,325,351]" pageId="13" pageNumber="265">Number</emphasis>
</th>
<th id="76FC001EEA084BABFE83FED1E218FEB6" box="[322,370,324,359]" gridcol="1" gridrow="1" pageId="13" pageNumber="265">
<emphasis id="B969B732EA08B459FE82FED1E218FE8F" bold="true" box="[323,370,324,350]" pageId="13" pageNumber="265">SVl1</emphasis>
</th>
<th id="76FC001EEA084BABFE4BFED1E2D5FEB6" box="[394,447,324,359]" gridcol="2" gridrow="1" pageId="13" pageNumber="265">
<emphasis id="B969B732EA08B459FE4DFED1E2D5FE8F" bold="true" box="[396,447,324,350]" pageId="13" pageNumber="265">SVl2</emphasis>
</th>
<th id="76FC001EEA084BABFE16FED1E16BFEB6" box="[471,513,324,359]" gridcol="3" gridrow="1" pageId="13" pageNumber="265">
<emphasis id="B969B732EA08B459FE18FED0E16BFE8E" bold="true" box="[473,513,325,351]" pageId="13" pageNumber="265">ILd</emphasis>
</th>
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<emphasis id="B969B732EA08B459FDDAFED0E12BFE8E" bold="true" box="[539,577,325,351]" pageId="13" pageNumber="265">FLl</emphasis>
</th>
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<emphasis id="B969B732EA08B459FDA5FED0E1E0FE8E" bold="true" box="[612,650,325,351]" pageId="13" pageNumber="265">TFl</emphasis>
</th>
<th id="76FC001EEA084BABFD65FED1E1BEFEB6" box="[676,724,324,359]" gridcol="6" gridrow="1" pageId="13" pageNumber="265">
<emphasis id="B969B732EA08B459FD64FED0E1BAFE8E" bold="true" box="[677,720,325,351]" pageId="13" pageNumber="265">HLl</emphasis>
</th>
<th id="76FC001EEA084BABFD2CFED1E07FFEB6" box="[749,789,324,359]" gridcol="7" gridrow="1" pageId="13" pageNumber="265">
<emphasis id="B969B732EA08B459FD2EFED0E07FFE8E" bold="true" box="[751,789,325,351]" pageId="13" pageNumber="265">FTl</emphasis>
</th>
<th id="76FC001EEA084BABFCEFFED1E030FEB6" box="[814,858,324,359]" gridcol="8" gridrow="1" pageId="13" pageNumber="265">
<emphasis id="B969B732EA08B459FCF1FED0E032FE8E" bold="true" box="[816,856,325,351]" pageId="13" pageNumber="265">Hw</emphasis>
</th>
<th id="76FC001EEA084BABFCB3FED1E0C9FEB6" box="[882,931,324,359]" gridcol="9" gridrow="1" pageId="13" pageNumber="265">
<emphasis id="B969B732EA08B459FCB5FED0E0FAFE8E" bold="true" box="[884,912,325,351]" pageId="13" pageNumber="265">Hl</emphasis>
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<th id="76FC001EEA084BABFC7FFED1E723FEB6" box="[958,1097,324,359]" gridcol="10" gridrow="1" pageId="13" pageNumber="265">
<emphasis id="B969B732EA08B459FC7FFED0E723FE8E" bold="true" box="[958,1097,325,351]" pageId="13" pageNumber="265">Total length</emphasis>
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<th id="76FC001EEA084BABFBA0FED1E7F4FEB6" box="[1121,1182,324,359]" gridcol="11" gridrow="1" pageId="13" pageNumber="265">
<emphasis id="B969B732EA08B459FBA2FED0E7F4FEB6" bold="true" box="[1123,1182,325,359]" pageId="13" pageNumber="265">Nlinks</emphasis>
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</tr>
<tr id="352D6962EA084BABFFBFFE13E7F4FE70" box="[126,1182,390,417]" gridrow="2" pageId="13" pageNumber="265">
<th id="76FC001EEA084BABFFBFFE13E38BFE70" box="[126,225,390,417]" gridcol="0" gridrow="2" pageId="13" pageNumber="265">1</th>
<td id="76FC001EEA084BABFE83FE13E218FE70" box="[322,370,390,417]" gridcol="1" gridrow="2" pageId="13" pageNumber="265">73</td>
<td id="76FC001EEA084BABFE4BFE13E2D5FE70" box="[394,447,390,417]" gridcol="2" gridrow="2" pageId="13" pageNumber="265">83</td>
<td id="76FC001EEA084BABFE16FE13E16BFE70" box="[471,513,390,417]" gridcol="3" gridrow="2" pageId="13" pageNumber="265">38</td>
<td id="76FC001EEA084BABFDDBFE13E123FE70" box="[538,585,390,417]" gridcol="4" gridrow="2" pageId="13" pageNumber="265">30.3</td>
<td id="76FC001EEA084BABFDA3FE13E1E0FE70" box="[610,650,390,417]" gridcol="5" gridrow="2" pageId="13" pageNumber="265">8.9</td>
<td id="76FC001EEA084BABFD65FE13E1BEFE70" box="[676,724,390,417]" gridcol="6" gridrow="2" pageId="13" pageNumber="265">32.6</td>
<td id="76FC001EEA084BABFD2CFE13E07FFE70" box="[749,789,390,417]" gridcol="7" gridrow="2" pageId="13" pageNumber="265">9.3</td>
<td id="76FC001EEA084BABFCEFFE13E030FE70" box="[814,858,390,417]" gridcol="8" gridrow="2" pageId="13" pageNumber="265">15.1</td>
<td id="76FC001EEA084BABFCB3FE13E0C9FE70" box="[882,931,390,417]" gridcol="9" gridrow="2" pageId="13" pageNumber="265">20.1</td>
<td id="76FC001EEA084BABFC7FFE13E723FE70" box="[958,1097,390,417]" gridcol="10" gridrow="2" pageId="13" pageNumber="265">151</td>
<td id="76FC001EEA084BABFBA0FE13E7F4FE70" box="[1121,1182,390,417]" gridcol="11" gridrow="2" pageId="13" pageNumber="265">1</td>
</tr>
<tr id="352D6962EA084BABFFBFFE3CE7F4FE15" box="[126,1182,425,452]" gridrow="3" pageId="13" pageNumber="265">
<th id="76FC001EEA084BABFFBFFE3CE38BFE15" box="[126,225,425,452]" gridcol="0" gridrow="3" pageId="13" pageNumber="265">2</th>
<td id="76FC001EEA084BABFE83FE3CE218FE15" box="[322,370,425,452]" gridcol="1" gridrow="3" pageId="13" pageNumber="265">58</td>
<td id="76FC001EEA084BABFE4BFE3CE2D5FE15" box="[394,447,425,452]" gridcol="2" gridrow="3" pageId="13" pageNumber="265">67</td>
<td id="76FC001EEA084BABFE16FE3CE16BFE15" box="[471,513,425,452]" gridcol="3" gridrow="3" pageId="13" pageNumber="265">31</td>
<td id="76FC001EEA084BABFDDBFE3CE123FE15" box="[538,585,425,452]" gridcol="4" gridrow="3" pageId="13" pageNumber="265">25.3</td>
<td id="76FC001EEA084BABFDA3FE3CE1E0FE15" box="[610,650,425,452]" gridcol="5" gridrow="3" pageId="13" pageNumber="265">7.8</td>
<td id="76FC001EEA084BABFD65FE3CE1BEFE15" box="[676,724,425,452]" gridcol="6" gridrow="3" pageId="13" pageNumber="265">25.1</td>
<td id="76FC001EEA084BABFD2CFE3CE07FFE15" box="[749,789,425,452]" gridcol="7" gridrow="3" pageId="13" pageNumber="265">7.3</td>
<td id="76FC001EEA084BABFCEFFE3CE030FE15" box="[814,858,425,452]" gridcol="8" gridrow="3" pageId="13" pageNumber="265">12.5</td>
<td id="76FC001EEA084BABFCB3FE3CE0C9FE15" box="[882,931,425,452]" gridcol="9" gridrow="3" pageId="13" pageNumber="265">17.9</td>
<td id="76FC001EEA084BABFC7FFE3CE723FE15" box="[958,1097,425,452]" gridcol="10" gridrow="3" pageId="13" pageNumber="265">123</td>
<td id="76FC001EEA084BABFBA0FE3CE7F4FE15" box="[1121,1182,425,452]" gridcol="11" gridrow="3" pageId="13" pageNumber="265">0</td>
</tr>
<tr id="352D6962EA084BABFFBFFE58E7F4FE39" box="[126,1182,461,488]" gridrow="4" pageId="13" pageNumber="265">
<th id="76FC001EEA084BABFFBFFE58E38BFE39" box="[126,225,461,488]" gridcol="0" gridrow="4" pageId="13" pageNumber="265">3</th>
<td id="76FC001EEA084BABFE83FE58E218FE39" box="[322,370,461,488]" gridcol="1" gridrow="4" pageId="13" pageNumber="265">65</td>
<td id="76FC001EEA084BABFE4BFE58E2D5FE39" box="[394,447,461,488]" gridcol="2" gridrow="4" pageId="13" pageNumber="265">75</td>
<td id="76FC001EEA084BABFE16FE58E16BFE39" box="[471,513,461,488]" gridcol="3" gridrow="4" pageId="13" pageNumber="265">32</td>
<td id="76FC001EEA084BABFDDBFE58E123FE39" box="[538,585,461,488]" gridcol="4" gridrow="4" pageId="13" pageNumber="265">28.8</td>
<td id="76FC001EEA084BABFDA3FE58E1E0FE39" box="[610,650,461,488]" gridcol="5" gridrow="4" pageId="13" pageNumber="265">7.4</td>
<td id="76FC001EEA084BABFD65FE58E1BEFE39" box="[676,724,461,488]" gridcol="6" gridrow="4" pageId="13" pageNumber="265">27.0</td>
<td id="76FC001EEA084BABFD2CFE58E07FFE39" box="[749,789,461,488]" gridcol="7" gridrow="4" pageId="13" pageNumber="265">8.3</td>
<td id="76FC001EEA084BABFCEFFE58E030FE39" box="[814,858,461,488]" gridcol="8" gridrow="4" pageId="13" pageNumber="265">13.3</td>
<td id="76FC001EEA084BABFCB3FE58E0C9FE39" box="[882,931,461,488]" gridcol="9" gridrow="4" pageId="13" pageNumber="265">19.0</td>
<td id="76FC001EEA084BABFC7FFE58E723FE39" box="[958,1097,461,488]" gridcol="10" gridrow="4" pageId="13" pageNumber="265">137</td>
<td id="76FC001EEA084BABFBA0FE58E7F4FE39" box="[1121,1182,461,488]" gridcol="11" gridrow="4" pageId="13" pageNumber="265">0</td>
</tr>
<tr id="352D6962EA084BABFFBFFE64E7F4FDDD" box="[126,1182,497,524]" gridrow="5" pageId="13" pageNumber="265">
<th id="76FC001EEA084BABFFBFFE64E38BFDDD" box="[126,225,497,524]" gridcol="0" gridrow="5" pageId="13" pageNumber="265">4</th>
<td id="76FC001EEA084BABFE83FE64E218FDDD" box="[322,370,497,524]" gridcol="1" gridrow="5" pageId="13" pageNumber="265">70</td>
<td id="76FC001EEA084BABFE4BFE64E2D5FDDD" box="[394,447,497,524]" gridcol="2" gridrow="5" pageId="13" pageNumber="265">79</td>
<td id="76FC001EEA084BABFE16FE64E16BFDDD" box="[471,513,497,524]" gridcol="3" gridrow="5" pageId="13" pageNumber="265">39</td>
<td id="76FC001EEA084BABFDDBFE64E123FDDD" box="[538,585,497,524]" gridcol="4" gridrow="5" pageId="13" pageNumber="265">27.4</td>
<td id="76FC001EEA084BABFDA3FE64E1E0FDDD" box="[610,650,497,524]" gridcol="5" gridrow="5" pageId="13" pageNumber="265">7.8</td>
<td id="76FC001EEA084BABFD65FE64E1BEFDDD" box="[676,724,497,524]" gridcol="6" gridrow="5" pageId="13" pageNumber="265">28.7</td>
<td id="76FC001EEA084BABFD2CFE64E07FFDDD" box="[749,789,497,524]" gridcol="7" gridrow="5" pageId="13" pageNumber="265">7.9</td>
<td id="76FC001EEA084BABFCEFFE64E030FDDD" box="[814,858,497,524]" gridcol="8" gridrow="5" pageId="13" pageNumber="265">14.3</td>
<td id="76FC001EEA084BABFCB3FE64E0C9FDDD" box="[882,931,497,524]" gridcol="9" gridrow="5" pageId="13" pageNumber="265">20.8</td>
<td id="76FC001EEA084BABFC7FFE64E723FDDD" box="[958,1097,497,524]" gridcol="10" gridrow="5" pageId="13" pageNumber="265">145</td>
<td id="76FC001EEA084BABFBA0FE64E7F4FDDD" box="[1121,1182,497,524]" gridcol="11" gridrow="5" pageId="13" pageNumber="265">0</td>
</tr>
<tr id="352D6962EA084BABFFBFFD80E7F4FDE1" box="[126,1182,533,560]" gridrow="6" pageId="13" pageNumber="265">
<th id="76FC001EEA084BABFFBFFD80E38BFDE1" box="[126,225,533,560]" gridcol="0" gridrow="6" pageId="13" pageNumber="265">5</th>
<td id="76FC001EEA084BABFE83FD80E218FDE1" box="[322,370,533,560]" gridcol="1" gridrow="6" pageId="13" pageNumber="265">70</td>
<td id="76FC001EEA084BABFE4BFD80E2D5FDE1" box="[394,447,533,560]" gridcol="2" gridrow="6" pageId="13" pageNumber="265">80</td>
<td id="76FC001EEA084BABFE16FD80E16BFDE1" box="[471,513,533,560]" gridcol="3" gridrow="6" pageId="13" pageNumber="265">38</td>
<td id="76FC001EEA084BABFDDBFD80E123FDE1" box="[538,585,533,560]" gridcol="4" gridrow="6" pageId="13" pageNumber="265">29.4</td>
<td id="76FC001EEA084BABFDA3FD80E1E0FDE1" box="[610,650,533,560]" gridcol="5" gridrow="6" pageId="13" pageNumber="265">8.4</td>
<td id="76FC001EEA084BABFD65FD80E1BEFDE1" box="[676,724,533,560]" gridcol="6" gridrow="6" pageId="13" pageNumber="265">28.7</td>
<td id="76FC001EEA084BABFD2CFD80E07FFDE1" box="[749,789,533,560]" gridcol="7" gridrow="6" pageId="13" pageNumber="265">8.8</td>
<td id="76FC001EEA084BABFCEFFD80E030FDE1" box="[814,858,533,560]" gridcol="8" gridrow="6" pageId="13" pageNumber="265">14.6</td>
<td id="76FC001EEA084BABFCB3FD80E0C9FDE1" box="[882,931,533,560]" gridcol="9" gridrow="6" pageId="13" pageNumber="265">18.0</td>
<td id="76FC001EEA084BABFC7FFD80E723FDE1" box="[958,1097,533,560]" gridcol="10" gridrow="6" pageId="13" pageNumber="265">147</td>
<td id="76FC001EEA084BABFBA0FD80E7F4FDE1" box="[1121,1182,533,560]" gridcol="11" gridrow="6" pageId="13" pageNumber="265">0</td>
</tr>
<tr id="352D6962EA084BABFFBFFDADE7F4FD82" box="[126,1182,568,595]" gridrow="7" pageId="13" pageNumber="265">
<th id="76FC001EEA084BABFFBFFDADE38BFD82" box="[126,225,568,595]" gridcol="0" gridrow="7" pageId="13" pageNumber="265">6</th>
<td id="76FC001EEA084BABFE83FDADE218FD82" box="[322,370,568,595]" gridcol="1" gridrow="7" pageId="13" pageNumber="265">66</td>
<td id="76FC001EEA084BABFE4BFDADE2D5FD82" box="[394,447,568,595]" gridcol="2" gridrow="7" pageId="13" pageNumber="265">75</td>
<td id="76FC001EEA084BABFE16FDADE16BFD82" box="[471,513,568,595]" gridcol="3" gridrow="7" pageId="13" pageNumber="265">37</td>
<td id="76FC001EEA084BABFDDBFDADE123FD82" box="[538,585,568,595]" gridcol="4" gridrow="7" pageId="13" pageNumber="265">27.9</td>
<td id="76FC001EEA084BABFDA3FDADE1E0FD82" box="[610,650,568,595]" gridcol="5" gridrow="7" pageId="13" pageNumber="265">7.6</td>
<td id="76FC001EEA084BABFD65FDADE1BEFD82" box="[676,724,568,595]" gridcol="6" gridrow="7" pageId="13" pageNumber="265">30.0</td>
<td id="76FC001EEA084BABFD2CFDADE07FFD82" box="[749,789,568,595]" gridcol="7" gridrow="7" pageId="13" pageNumber="265">8.5</td>
<td id="76FC001EEA084BABFCEFFDADE030FD82" box="[814,858,568,595]" gridcol="8" gridrow="7" pageId="13" pageNumber="265">13.8</td>
<td id="76FC001EEA084BABFCB3FDADE0C9FD82" box="[882,931,568,595]" gridcol="9" gridrow="7" pageId="13" pageNumber="265">18.8</td>
<td id="76FC001EEA084BABFC7FFDADE723FD82" box="[958,1097,568,595]" gridcol="10" gridrow="7" pageId="13" pageNumber="265">139</td>
<td id="76FC001EEA084BABFBA0FDADE7F4FD82" box="[1121,1182,568,595]" gridcol="11" gridrow="7" pageId="13" pageNumber="265">1</td>
</tr>
<tr id="352D6962EA084BABFFBFFDC9E7F4FDA6" box="[126,1182,604,631]" gridrow="8" pageId="13" pageNumber="265">
<th id="76FC001EEA084BABFFBFFDC9E38BFDA6" box="[126,225,604,631]" gridcol="0" gridrow="8" pageId="13" pageNumber="265">7</th>
<td id="76FC001EEA084BABFE83FDC9E218FDA6" box="[322,370,604,631]" gridcol="1" gridrow="8" pageId="13" pageNumber="265">59</td>
<td id="76FC001EEA084BABFE4BFDC9E2D5FDA6" box="[394,447,604,631]" gridcol="2" gridrow="8" pageId="13" pageNumber="265">63</td>
<td id="76FC001EEA084BABFE16FDC9E16BFDA6" box="[471,513,604,631]" gridcol="3" gridrow="8" pageId="13" pageNumber="265">30</td>
<td id="76FC001EEA084BABFDDBFDC9E123FDA6" box="[538,585,604,631]" gridcol="4" gridrow="8" pageId="13" pageNumber="265">25.8</td>
<td id="76FC001EEA084BABFDA3FDC9E1E0FDA6" box="[610,650,604,631]" gridcol="5" gridrow="8" pageId="13" pageNumber="265">6.7</td>
<td id="76FC001EEA084BABFD65FDC9E1BEFDA6" box="[676,724,604,631]" gridcol="6" gridrow="8" pageId="13" pageNumber="265">25.3</td>
<td id="76FC001EEA084BABFD2CFDC9E07FFDA6" box="[749,789,604,631]" gridcol="7" gridrow="8" pageId="13" pageNumber="265">7.2</td>
<td id="76FC001EEA084BABFCEFFDC9E030FDA6" box="[814,858,604,631]" gridcol="8" gridrow="8" pageId="13" pageNumber="265">13.2</td>
<td id="76FC001EEA084BABFCB3FDC9E0C9FDA6" box="[882,931,604,631]" gridcol="9" gridrow="8" pageId="13" pageNumber="265">17.5</td>
<td id="76FC001EEA084BABFC7FFDC9E723FDA6" box="[958,1097,604,631]" gridcol="10" gridrow="8" pageId="13" pageNumber="265">126</td>
<td id="76FC001EEA084BABFBA0FDC9E7F4FDA6" box="[1121,1182,604,631]" gridcol="11" gridrow="8" pageId="13" pageNumber="265">0</td>
</tr>
<tr id="352D6962EA084BABFFBFFD15E7F4FD4A" box="[126,1182,640,667]" gridrow="9" pageId="13" pageNumber="265">
<th id="76FC001EEA084BABFFBFFD15E38BFD4A" box="[126,225,640,667]" gridcol="0" gridrow="9" pageId="13" pageNumber="265">8</th>
<td id="76FC001EEA084BABFE83FD15E218FD4A" box="[322,370,640,667]" gridcol="1" gridrow="9" pageId="13" pageNumber="265">65</td>
<td id="76FC001EEA084BABFE4BFD15E2D5FD4A" box="[394,447,640,667]" gridcol="2" gridrow="9" pageId="13" pageNumber="265">75</td>
<td id="76FC001EEA084BABFE16FD15E16BFD4A" box="[471,513,640,667]" gridcol="3" gridrow="9" pageId="13" pageNumber="265">37</td>
<td id="76FC001EEA084BABFDDBFD15E123FD4A" box="[538,585,640,667]" gridcol="4" gridrow="9" pageId="13" pageNumber="265">27.3</td>
<td id="76FC001EEA084BABFDA3FD15E1E0FD4A" box="[610,650,640,667]" gridcol="5" gridrow="9" pageId="13" pageNumber="265">7.2</td>
<td id="76FC001EEA084BABFD65FD15E1BEFD4A" box="[676,724,640,667]" gridcol="6" gridrow="9" pageId="13" pageNumber="265">27.9</td>
<td id="76FC001EEA084BABFD2CFD15E07FFD4A" box="[749,789,640,667]" gridcol="7" gridrow="9" pageId="13" pageNumber="265">7.8</td>
<td id="76FC001EEA084BABFCEFFD15E030FD4A" box="[814,858,640,667]" gridcol="8" gridrow="9" pageId="13" pageNumber="265">13.8</td>
<td id="76FC001EEA084BABFCB3FD15E0C9FD4A" box="[882,931,640,667]" gridcol="9" gridrow="9" pageId="13" pageNumber="265">18.0</td>
<td id="76FC001EEA084BABFC7FFD15E723FD4A" box="[958,1097,640,667]" gridcol="10" gridrow="9" pageId="13" pageNumber="265">136</td>
<td id="76FC001EEA084BABFBA0FD15E7F4FD4A" box="[1121,1182,640,667]" gridcol="11" gridrow="9" pageId="13" pageNumber="265">0</td>
</tr>
<tr id="352D6962EA084BABFFBFFD31E7F4FD6E" box="[126,1182,676,703]" gridrow="10" pageId="13" pageNumber="265">
<th id="76FC001EEA084BABFFBFFD31E38BFD6E" box="[126,225,676,703]" gridcol="0" gridrow="10" pageId="13" pageNumber="265">9</th>
<td id="76FC001EEA084BABFE83FD31E218FD6E" box="[322,370,676,703]" gridcol="1" gridrow="10" pageId="13" pageNumber="265">74</td>
<td id="76FC001EEA084BABFE4BFD31E2D5FD6E" box="[394,447,676,703]" gridcol="2" gridrow="10" pageId="13" pageNumber="265">83</td>
<td id="76FC001EEA084BABFE16FD31E16BFD6E" box="[471,513,676,703]" gridcol="3" gridrow="10" pageId="13" pageNumber="265">42</td>
<td id="76FC001EEA084BABFDDBFD31E123FD6E" box="[538,585,676,703]" gridcol="4" gridrow="10" pageId="13" pageNumber="265">31.7</td>
<td id="76FC001EEA084BABFDA3FD31E1E0FD6E" box="[610,650,676,703]" gridcol="5" gridrow="10" pageId="13" pageNumber="265">9.2</td>
<td id="76FC001EEA084BABFD65FD31E1BEFD6E" box="[676,724,676,703]" gridcol="6" gridrow="10" pageId="13" pageNumber="265">33.0</td>
<td id="76FC001EEA084BABFD2CFD31E07FFD6E" box="[749,789,676,703]" gridcol="7" gridrow="10" pageId="13" pageNumber="265">9.6</td>
<td id="76FC001EEA084BABFCEFFD31E030FD6E" box="[814,858,676,703]" gridcol="8" gridrow="10" pageId="13" pageNumber="265">15.5</td>
<td id="76FC001EEA084BABFCB3FD31E0C9FD6E" box="[882,931,676,703]" gridcol="9" gridrow="10" pageId="13" pageNumber="265">21.1</td>
<td id="76FC001EEA084BABFC7FFD31E723FD6E" box="[958,1097,676,703]" gridcol="10" gridrow="10" pageId="13" pageNumber="265">139 #</td>
<td id="76FC001EEA084BABFBA0FD31E7F4FD6E" box="[1121,1182,676,703]" gridcol="11" gridrow="10" pageId="13" pageNumber="265">0</td>
</tr>
<tr id="352D6962EA084BABFFBFFD52E7F4FD33" box="[126,1182,711,738]" gridrow="11" pageId="13" pageNumber="265">
<th id="76FC001EEA084BABFFBFFD52E38BFD33" box="[126,225,711,738]" gridcol="0" gridrow="11" pageId="13" pageNumber="265">10</th>
<td id="76FC001EEA084BABFE83FD52E218FD33" box="[322,370,711,738]" gridcol="1" gridrow="11" pageId="13" pageNumber="265">67</td>
<td id="76FC001EEA084BABFE4BFD52E2D5FD33" box="[394,447,711,738]" gridcol="2" gridrow="11" pageId="13" pageNumber="265">75</td>
<td id="76FC001EEA084BABFE16FD52E16BFD33" box="[471,513,711,738]" gridcol="3" gridrow="11" pageId="13" pageNumber="265">33</td>
<td id="76FC001EEA084BABFDDBFD52E123FD33" box="[538,585,711,738]" gridcol="4" gridrow="11" pageId="13" pageNumber="265">30.5</td>
<td id="76FC001EEA084BABFDA3FD52E1E0FD33" box="[610,650,711,738]" gridcol="5" gridrow="11" pageId="13" pageNumber="265">8.6</td>
<td id="76FC001EEA084BABFD65FD52E1BEFD33" box="[676,724,711,738]" gridcol="6" gridrow="11" pageId="13" pageNumber="265">30.2</td>
<td id="76FC001EEA084BABFD2CFD52E07FFD33" box="[749,789,711,738]" gridcol="7" gridrow="11" pageId="13" pageNumber="265">9.1</td>
<td id="76FC001EEA084BABFCEFFD52E030FD33" box="[814,858,711,738]" gridcol="8" gridrow="11" pageId="13" pageNumber="265">14.2</td>
<td id="76FC001EEA084BABFCB3FD52E0C9FD33" box="[882,931,711,738]" gridcol="9" gridrow="11" pageId="13" pageNumber="265">16.8</td>
<td id="76FC001EEA084BABFC7FFD52E723FD33" box="[958,1097,711,738]" gridcol="10" gridrow="11" pageId="13" pageNumber="265">140</td>
<td id="76FC001EEA084BABFBA0FD52E7F4FD33" box="[1121,1182,711,738]" gridcol="11" gridrow="11" pageId="13" pageNumber="265">0</td>
</tr>
<tr id="352D6962EA084BABFFBFFD7EE7F4FCD7" box="[126,1182,747,774]" gridrow="12" pageId="13" pageNumber="265">
<th id="76FC001EEA084BABFFBFFD7EE38BFCD7" box="[126,225,747,774]" gridcol="0" gridrow="12" pageId="13" pageNumber="265">11</th>
<td id="76FC001EEA084BABFE83FD7EE218FCD7" box="[322,370,747,774]" gridcol="1" gridrow="12" pageId="13" pageNumber="265">66</td>
<td id="76FC001EEA084BABFE4BFD7EE2D5FCD7" box="[394,447,747,774]" gridcol="2" gridrow="12" pageId="13" pageNumber="265">75</td>
<td id="76FC001EEA084BABFE16FD7EE16BFCD7" box="[471,513,747,774]" gridcol="3" gridrow="12" pageId="13" pageNumber="265">34</td>
<td id="76FC001EEA084BABFDDBFD7EE123FCD7" box="[538,585,747,774]" gridcol="4" gridrow="12" pageId="13" pageNumber="265">30.0</td>
<td id="76FC001EEA084BABFDA3FD7EE1E0FCD7" box="[610,650,747,774]" gridcol="5" gridrow="12" pageId="13" pageNumber="265">8.8</td>
<td id="76FC001EEA084BABFD65FD7EE1BEFCD7" box="[676,724,747,774]" gridcol="6" gridrow="12" pageId="13" pageNumber="265">28.5</td>
<td id="76FC001EEA084BABFD2CFD7EE07FFCD7" box="[749,789,747,774]" gridcol="7" gridrow="12" pageId="13" pageNumber="265">8.3</td>
<td id="76FC001EEA084BABFCEFFD7EE030FCD7" box="[814,858,747,774]" gridcol="8" gridrow="12" pageId="13" pageNumber="265">14.5</td>
<td id="76FC001EEA084BABFCB3FD7EE0C9FCD7" box="[882,931,747,774]" gridcol="9" gridrow="12" pageId="13" pageNumber="265">18.5</td>
<td id="76FC001EEA084BABFC7FFD7EE723FCD7" box="[958,1097,747,774]" gridcol="10" gridrow="12" pageId="13" pageNumber="265">137</td>
<td id="76FC001EEA084BABFBA0FD7EE7F4FCD7" box="[1121,1182,747,774]" gridcol="11" gridrow="12" pageId="13" pageNumber="265">0</td>
</tr>
<tr id="352D6962EA084BABFFBFFC9AE7F4FCF8" box="[126,1182,783,809]" gridrow="13" pageId="13" pageNumber="265">
<th id="76FC001EEA084BABFFBFFC9AE38BFCF8" box="[126,225,783,809]" gridcol="0" gridrow="13" pageId="13" pageNumber="265">12</th>
<td id="76FC001EEA084BABFE83FC9AE218FCF8" box="[322,370,783,809]" gridcol="1" gridrow="13" pageId="13" pageNumber="265">62</td>
<td id="76FC001EEA084BABFE4BFC9AE2D5FCF8" box="[394,447,783,809]" gridcol="2" gridrow="13" pageId="13" pageNumber="265">70</td>
<td id="76FC001EEA084BABFE16FC9AE16BFCF8" box="[471,513,783,809]" gridcol="3" gridrow="13" pageId="13" pageNumber="265">36</td>
<td id="76FC001EEA084BABFDDBFC9AE123FCF8" box="[538,585,783,809]" gridcol="4" gridrow="13" pageId="13" pageNumber="265">28.3</td>
<td id="76FC001EEA084BABFDA3FC9AE1E0FCF8" box="[610,650,783,809]" gridcol="5" gridrow="13" pageId="13" pageNumber="265">8.8</td>
<td id="76FC001EEA084BABFD65FC9AE1BEFCF8" box="[676,724,783,809]" gridcol="6" gridrow="13" pageId="13" pageNumber="265">28.8</td>
<td id="76FC001EEA084BABFD2CFC9AE07FFCF8" box="[749,789,783,809]" gridcol="7" gridrow="13" pageId="13" pageNumber="265">9.3</td>
<td id="76FC001EEA084BABFCEFFC9AE030FCF8" box="[814,858,783,809]" gridcol="8" gridrow="13" pageId="13" pageNumber="265">13.6</td>
<td id="76FC001EEA084BABFCB3FC9AE0C9FCF8" box="[882,931,783,809]" gridcol="9" gridrow="13" pageId="13" pageNumber="265">16.1</td>
<td id="76FC001EEA084BABFC7FFC9AE723FCF8" box="[958,1097,783,809]" gridcol="10" gridrow="13" pageId="13" pageNumber="265">132</td>
<td id="76FC001EEA084BABFBA0FC9AE7F4FCF8" box="[1121,1182,783,809]" gridcol="11" gridrow="13" pageId="13" pageNumber="265">0</td>
</tr>
<tr id="352D6962EA084BABFFBFFCA8E7F4FC86" box="[126,1182,829,855]" gridrow="14" pageId="13" pageNumber="265">
<th id="76FC001EEA084BABFFBFFCA8E7F4FC86" box="[126,1182,829,855]" colspan="12" colspanRight="11" gridcol="0" gridrow="14" pageId="13" pageNumber="265">Females</th>
</tr>
<tr id="352D6962EA084BABFFBFFCFEE7F4FC57" box="[126,1182,875,902]" gridrow="15" pageId="13" pageNumber="265">
<th id="76FC001EEA084BABFFBFFCFEE38BFC57" box="[126,225,875,902]" gridcol="0" gridrow="15" pageId="13" pageNumber="265">13</th>
<td id="76FC001EEA084BABFE83FCFEE218FC57" box="[322,370,875,902]" gridcol="1" gridrow="15" pageId="13" pageNumber="265">73</td>
<td id="76FC001EEA084BABFE4BFCFEE2D5FC57" box="[394,447,875,902]" gridcol="2" gridrow="15" pageId="13" pageNumber="265">81</td>
<td id="76FC001EEA084BABFE16FCFEE16BFC57" box="[471,513,875,902]" gridcol="3" gridrow="15" pageId="13" pageNumber="265">39</td>
<td id="76FC001EEA084BABFDDBFCFEE123FC57" box="[538,585,875,902]" gridcol="4" gridrow="15" pageId="13" pageNumber="265">27.2</td>
<td id="76FC001EEA084BABFDA3FCFEE1E0FC57" box="[610,650,875,902]" gridcol="5" gridrow="15" pageId="13" pageNumber="265">7.8</td>
<td id="76FC001EEA084BABFD65FCFEE1BEFC57" box="[676,724,875,902]" gridcol="6" gridrow="15" pageId="13" pageNumber="265">27.5</td>
<td id="76FC001EEA084BABFD2CFCFEE07FFC57" box="[749,789,875,902]" gridcol="7" gridrow="15" pageId="13" pageNumber="265">6.3</td>
<td id="76FC001EEA084BABFCEFFCFEE030FC57" box="[814,858,875,902]" gridcol="8" gridrow="15" pageId="13" pageNumber="265">15.6</td>
<td id="76FC001EEA084BABFCB3FCFEE0C9FC57" box="[882,931,875,902]" gridcol="9" gridrow="15" pageId="13" pageNumber="265">19.4</td>
<td id="76FC001EEA084BABFC7FFCFEE723FC57" box="[958,1097,875,902]" gridcol="10" gridrow="15" pageId="13" pageNumber="265">151</td>
<td id="76FC001EEA084BABFBA0FCFEE7F4FC57" box="[1121,1182,875,902]" gridcol="11" gridrow="15" pageId="13" pageNumber="265">0</td>
</tr>
<tr id="352D6962EA084BABFFBFFC1AE7F4FC7B" box="[126,1182,911,938]" gridrow="16" pageId="13" pageNumber="265">
<th id="76FC001EEA084BABFFBFFC1AE38BFC7B" box="[126,225,911,938]" gridcol="0" gridrow="16" pageId="13" pageNumber="265">14</th>
<td id="76FC001EEA084BABFE83FC1AE218FC7B" box="[322,370,911,938]" gridcol="1" gridrow="16" pageId="13" pageNumber="265">73</td>
<td id="76FC001EEA084BABFE4BFC1AE2D5FC7B" box="[394,447,911,938]" gridcol="2" gridrow="16" pageId="13" pageNumber="265">81</td>
<td id="76FC001EEA084BABFE16FC1AE16BFC7B" box="[471,513,911,938]" gridcol="3" gridrow="16" pageId="13" pageNumber="265">40</td>
<td id="76FC001EEA084BABFDDBFC1AE123FC7B" box="[538,585,911,938]" gridcol="4" gridrow="16" pageId="13" pageNumber="265">27.7</td>
<td id="76FC001EEA084BABFDA3FC1AE1E0FC7B" box="[610,650,911,938]" gridcol="5" gridrow="16" pageId="13" pageNumber="265">6.7</td>
<td id="76FC001EEA084BABFD65FC1AE1BEFC7B" box="[676,724,911,938]" gridcol="6" gridrow="16" pageId="13" pageNumber="265">28.2</td>
<td id="76FC001EEA084BABFD2CFC1AE07FFC7B" box="[749,789,911,938]" gridcol="7" gridrow="16" pageId="13" pageNumber="265">7.1</td>
<td id="76FC001EEA084BABFCEFFC1AE030FC7B" box="[814,858,911,938]" gridcol="8" gridrow="16" pageId="13" pageNumber="265">14.2</td>
<td id="76FC001EEA084BABFCB3FC1AE0C9FC7B" box="[882,931,911,938]" gridcol="9" gridrow="16" pageId="13" pageNumber="265">20.5</td>
<td id="76FC001EEA084BABFC7FFC1AE723FC7B" box="[958,1097,911,938]" gridcol="10" gridrow="16" pageId="13" pageNumber="265">152</td>
<td id="76FC001EEA084BABFBA0FC1AE7F4FC7B" box="[1121,1182,911,938]" gridcol="11" gridrow="16" pageId="13" pageNumber="265">0</td>
</tr>
<tr id="352D6962EA084BABFFBFFC26E7F4FC1F" box="[126,1182,947,974]" gridrow="17" pageId="13" pageNumber="265">
<th id="76FC001EEA084BABFFBFFC26E38BFC1F" box="[126,225,947,974]" gridcol="0" gridrow="17" pageId="13" pageNumber="265">15</th>
<td id="76FC001EEA084BABFE83FC26E218FC1F" box="[322,370,947,974]" gridcol="1" gridrow="17" pageId="13" pageNumber="265">75</td>
<td id="76FC001EEA084BABFE4BFC26E2D5FC1F" box="[394,447,947,974]" gridcol="2" gridrow="17" pageId="13" pageNumber="265">82</td>
<td id="76FC001EEA084BABFE16FC26E16BFC1F" box="[471,513,947,974]" gridcol="3" gridrow="17" pageId="13" pageNumber="265">41</td>
<td id="76FC001EEA084BABFDDBFC26E123FC1F" box="[538,585,947,974]" gridcol="4" gridrow="17" pageId="13" pageNumber="265">19.3</td>
<td id="76FC001EEA084BABFDA3FC26E1E0FC1F" box="[610,650,947,974]" gridcol="5" gridrow="17" pageId="13" pageNumber="265">8.0</td>
<td id="76FC001EEA084BABFD65FC26E1BEFC1F" box="[676,724,947,974]" gridcol="6" gridrow="17" pageId="13" pageNumber="265">28.9</td>
<td id="76FC001EEA084BABFD2CFC26E07FFC1F" box="[749,789,947,974]" gridcol="7" gridrow="17" pageId="13" pageNumber="265">7.7</td>
<td id="76FC001EEA084BABFCEFFC26E030FC1F" box="[814,858,947,974]" gridcol="8" gridrow="17" pageId="13" pageNumber="265">14.7</td>
<td id="76FC001EEA084BABFCB3FC26E0C9FC1F" box="[882,931,947,974]" gridcol="9" gridrow="17" pageId="13" pageNumber="265">19.6</td>
<td id="76FC001EEA084BABFC7FFC26E723FC1F" box="[958,1097,947,974]" gridcol="10" gridrow="17" pageId="13" pageNumber="265">156</td>
<td id="76FC001EEA084BABFBA0FC26E7F4FC1F" box="[1121,1182,947,974]" gridcol="11" gridrow="17" pageId="13" pageNumber="265">0</td>
</tr>
<tr id="352D6962EA084BABFFBFFC42E7F4FC23" box="[126,1182,983,1010]" gridrow="18" pageId="13" pageNumber="265">
<th id="76FC001EEA084BABFFBFFC42E38BFC23" box="[126,225,983,1010]" gridcol="0" gridrow="18" pageId="13" pageNumber="265">16</th>
<td id="76FC001EEA084BABFE83FC42E218FC23" box="[322,370,983,1010]" gridcol="1" gridrow="18" pageId="13" pageNumber="265">76</td>
<td id="76FC001EEA084BABFE4BFC42E2D5FC23" box="[394,447,983,1010]" gridcol="2" gridrow="18" pageId="13" pageNumber="265">84</td>
<td id="76FC001EEA084BABFE16FC42E16BFC23" box="[471,513,983,1010]" gridcol="3" gridrow="18" pageId="13" pageNumber="265">43</td>
<td id="76FC001EEA084BABFDDBFC42E123FC23" box="[538,585,983,1010]" gridcol="4" gridrow="18" pageId="13" pageNumber="265">28.4</td>
<td id="76FC001EEA084BABFDA3FC42E1E0FC23" box="[610,650,983,1010]" gridcol="5" gridrow="18" pageId="13" pageNumber="265">7.4</td>
<td id="76FC001EEA084BABFD65FC42E1BEFC23" box="[676,724,983,1010]" gridcol="6" gridrow="18" pageId="13" pageNumber="265">28.0</td>
<td id="76FC001EEA084BABFD2CFC42E07FFC23" box="[749,789,983,1010]" gridcol="7" gridrow="18" pageId="13" pageNumber="265">7.1</td>
<td id="76FC001EEA084BABFCEFFC42E030FC23" box="[814,858,983,1010]" gridcol="8" gridrow="18" pageId="13" pageNumber="265">15.6</td>
<td id="76FC001EEA084BABFCB3FC42E0C9FC23" box="[882,931,983,1010]" gridcol="9" gridrow="18" pageId="13" pageNumber="265">21.3</td>
<td id="76FC001EEA084BABFC7FFC42E723FC23" box="[958,1097,983,1010]" gridcol="10" gridrow="18" pageId="13" pageNumber="265">155</td>
<td id="76FC001EEA084BABFBA0FC42E7F4FC23" box="[1121,1182,983,1010]" gridcol="11" gridrow="18" pageId="13" pageNumber="265">0</td>
</tr>
<tr id="352D6962EA084BABFFBFFC6FE7F4FBC4" box="[126,1182,1018,1045]" gridrow="19" pageId="13" pageNumber="265">
<th id="76FC001EEA084BABFFBFFC6FE38BFBC4" box="[126,225,1018,1045]" gridcol="0" gridrow="19" pageId="13" pageNumber="265">17</th>
<td id="76FC001EEA084BABFE83FC6FE218FBC4" box="[322,370,1018,1045]" gridcol="1" gridrow="19" pageId="13" pageNumber="265">76</td>
<td id="76FC001EEA084BABFE4BFC6FE2D5FBC4" box="[394,447,1018,1045]" gridcol="2" gridrow="19" pageId="13" pageNumber="265">83</td>
<td id="76FC001EEA084BABFE16FC6FE16BFBC4" box="[471,513,1018,1045]" gridcol="3" gridrow="19" pageId="13" pageNumber="265">42</td>
<td id="76FC001EEA084BABFDDBFC6FE123FBC4" box="[538,585,1018,1045]" gridcol="4" gridrow="19" pageId="13" pageNumber="265">28.9</td>
<td id="76FC001EEA084BABFDA3FC6FE1E0FBC4" box="[610,650,1018,1045]" gridcol="5" gridrow="19" pageId="13" pageNumber="265">8.4</td>
<td id="76FC001EEA084BABFD65FC6FE1BEFBC4" box="[676,724,1018,1045]" gridcol="6" gridrow="19" pageId="13" pageNumber="265">28.5</td>
<td id="76FC001EEA084BABFD2CFC6FE07FFBC4" box="[749,789,1018,1045]" gridcol="7" gridrow="19" pageId="13" pageNumber="265">7.8</td>
<td id="76FC001EEA084BABFCEFFC6FE030FBC4" box="[814,858,1018,1045]" gridcol="8" gridrow="19" pageId="13" pageNumber="265">15.1</td>
<td id="76FC001EEA084BABFCB3FC6FE0C9FBC4" box="[882,931,1018,1045]" gridcol="9" gridrow="19" pageId="13" pageNumber="265">18.9</td>
<td id="76FC001EEA084BABFC7FFC6FE723FBC4" box="[958,1097,1018,1045]" gridcol="10" gridrow="19" pageId="13" pageNumber="265">155</td>
<td id="76FC001EEA084BABFBA0FC6FE7F4FBC4" box="[1121,1182,1018,1045]" gridcol="11" gridrow="19" pageId="13" pageNumber="265">0</td>
</tr>
<tr id="352D6962EA084BABFFBFFB8BE7F4FBE8" box="[126,1182,1054,1081]" gridrow="20" pageId="13" pageNumber="265">
<th id="76FC001EEA084BABFFBFFB8BE38BFBE8" box="[126,225,1054,1081]" gridcol="0" gridrow="20" pageId="13" pageNumber="265">18</th>
<td id="76FC001EEA084BABFE83FB8BE218FBE8" box="[322,370,1054,1081]" gridcol="1" gridrow="20" pageId="13" pageNumber="265">71</td>
<td id="76FC001EEA084BABFE4BFB8BE2D5FBE8" box="[394,447,1054,1081]" gridcol="2" gridrow="20" pageId="13" pageNumber="265">78</td>
<td id="76FC001EEA084BABFE16FB8BE16BFBE8" box="[471,513,1054,1081]" gridcol="3" gridrow="20" pageId="13" pageNumber="265">40</td>
<td id="76FC001EEA084BABFDDBFB8BE123FBE8" box="[538,585,1054,1081]" gridcol="4" gridrow="20" pageId="13" pageNumber="265">26.5</td>
<td id="76FC001EEA084BABFDA3FB8BE1E0FBE8" box="[610,650,1054,1081]" gridcol="5" gridrow="20" pageId="13" pageNumber="265">7.4</td>
<td id="76FC001EEA084BABFD65FB8BE1BEFBE8" box="[676,724,1054,1081]" gridcol="6" gridrow="20" pageId="13" pageNumber="265">26.7</td>
<td id="76FC001EEA084BABFD2CFB8BE07FFBE8" box="[749,789,1054,1081]" gridcol="7" gridrow="20" pageId="13" pageNumber="265">6.6</td>
<td id="76FC001EEA084BABFCEFFB8BE030FBE8" box="[814,858,1054,1081]" gridcol="8" gridrow="20" pageId="13" pageNumber="265">15.5</td>
<td id="76FC001EEA084BABFCB3FB8BE0C9FBE8" box="[882,931,1054,1081]" gridcol="9" gridrow="20" pageId="13" pageNumber="265">20.6</td>
<td id="76FC001EEA084BABFC7FFB8BE723FBE8" box="[958,1097,1054,1081]" gridcol="10" gridrow="20" pageId="13" pageNumber="265">148</td>
<td id="76FC001EEA084BABFBA0FB8BE7F4FBE8" box="[1121,1182,1054,1081]" gridcol="11" gridrow="20" pageId="13" pageNumber="265">3</td>
</tr>
<tr id="352D6962EA084BABFFBFFBD7E7F4FB8C" box="[126,1182,1090,1117]" gridrow="21" pageId="13" pageNumber="265">
<th id="76FC001EEA084BABFFBFFBD7E38BFB8C" box="[126,225,1090,1117]" gridcol="0" gridrow="21" pageId="13" pageNumber="265">19</th>
<td id="76FC001EEA084BABFE83FBD7E218FB8C" box="[322,370,1090,1117]" gridcol="1" gridrow="21" pageId="13" pageNumber="265">75</td>
<td id="76FC001EEA084BABFE4BFBD7E2D5FB8C" box="[394,447,1090,1117]" gridcol="2" gridrow="21" pageId="13" pageNumber="265">84</td>
<td id="76FC001EEA084BABFE16FBD7E16BFB8C" box="[471,513,1090,1117]" gridcol="3" gridrow="21" pageId="13" pageNumber="265">40</td>
<td id="76FC001EEA084BABFDDBFBD7E123FB8C" box="[538,585,1090,1117]" gridcol="4" gridrow="21" pageId="13" pageNumber="265">28.1</td>
<td id="76FC001EEA084BABFDA3FBD7E1E0FB8C" box="[610,650,1090,1117]" gridcol="5" gridrow="21" pageId="13" pageNumber="265">6.4</td>
<td id="76FC001EEA084BABFD65FBD7E1BEFB8C" box="[676,724,1090,1117]" gridcol="6" gridrow="21" pageId="13" pageNumber="265">26.5</td>
<td id="76FC001EEA084BABFD2CFBD7E07FFB8C" box="[749,789,1090,1117]" gridcol="7" gridrow="21" pageId="13" pageNumber="265">6.2</td>
<td id="76FC001EEA084BABFCEFFBD7E030FB8C" box="[814,858,1090,1117]" gridcol="8" gridrow="21" pageId="13" pageNumber="265">15.4</td>
<td id="76FC001EEA084BABFCB3FBD7E0C9FB8C" box="[882,931,1090,1117]" gridcol="9" gridrow="21" pageId="13" pageNumber="265">21.5</td>
<td id="76FC001EEA084BABFC7FFBD7E723FB8C" box="[958,1097,1090,1117]" gridcol="10" gridrow="21" pageId="13" pageNumber="265">155</td>
<td id="76FC001EEA084BABFBA0FBD7E7F4FB8C" box="[1121,1182,1090,1117]" gridcol="11" gridrow="21" pageId="13" pageNumber="265">0</td>
</tr>
<tr id="352D6962EA084BABFFBFFBF3E7F4FB50" box="[126,1182,1126,1153]" gridrow="22" pageId="13" pageNumber="265">
<th id="76FC001EEA084BABFFBFFBF3E38BFB50" box="[126,225,1126,1153]" gridcol="0" gridrow="22" pageId="13" pageNumber="265">20</th>
<td id="76FC001EEA084BABFE83FBF3E218FB50" box="[322,370,1126,1153]" gridcol="1" gridrow="22" pageId="13" pageNumber="265">75</td>
<td id="76FC001EEA084BABFE4BFBF3E2D5FB50" box="[394,447,1126,1153]" gridcol="2" gridrow="22" pageId="13" pageNumber="265">83</td>
<td id="76FC001EEA084BABFE16FBF3E16BFB50" box="[471,513,1126,1153]" gridcol="3" gridrow="22" pageId="13" pageNumber="265">44</td>
<td id="76FC001EEA084BABFDDBFBF3E123FB50" box="[538,585,1126,1153]" gridcol="4" gridrow="22" pageId="13" pageNumber="265">28.5</td>
<td id="76FC001EEA084BABFDA3FBF3E1E0FB50" box="[610,650,1126,1153]" gridcol="5" gridrow="22" pageId="13" pageNumber="265">6.7</td>
<td id="76FC001EEA084BABFD65FBF3E1BEFB50" box="[676,724,1126,1153]" gridcol="6" gridrow="22" pageId="13" pageNumber="265">27.4</td>
<td id="76FC001EEA084BABFD2CFBF3E07FFB50" box="[749,789,1126,1153]" gridcol="7" gridrow="22" pageId="13" pageNumber="265">6.6</td>
<td id="76FC001EEA084BABFCEFFBF3E030FB50" box="[814,858,1126,1153]" gridcol="8" gridrow="22" pageId="13" pageNumber="265">15.6</td>
<td id="76FC001EEA084BABFCB3FBF3E0C9FB50" box="[882,931,1126,1153]" gridcol="9" gridrow="22" pageId="13" pageNumber="265">19.3</td>
<td id="76FC001EEA084BABFC7FFBF3E723FB50" box="[958,1097,1126,1153]" gridcol="10" gridrow="22" pageId="13" pageNumber="265">155</td>
<td id="76FC001EEA084BABFBA0FBF3E7F4FB50" box="[1121,1182,1126,1153]" gridcol="11" gridrow="22" pageId="13" pageNumber="265">4</td>
</tr>
<tr id="352D6962EA084BABFFBFFB1CE7F4FB75" box="[126,1182,1161,1188]" gridrow="23" pageId="13" pageNumber="265">
<th id="76FC001EEA084BABFFBFFB1CE38BFB75" box="[126,225,1161,1188]" gridcol="0" gridrow="23" pageId="13" pageNumber="265">21</th>
<td id="76FC001EEA084BABFE83FB1CE218FB75" box="[322,370,1161,1188]" gridcol="1" gridrow="23" pageId="13" pageNumber="265">72</td>
<td id="76FC001EEA084BABFE4BFB1CE2D5FB75" box="[394,447,1161,1188]" gridcol="2" gridrow="23" pageId="13" pageNumber="265">80</td>
<td id="76FC001EEA084BABFE16FB1CE16BFB75" box="[471,513,1161,1188]" gridcol="3" gridrow="23" pageId="13" pageNumber="265">40</td>
<td id="76FC001EEA084BABFDDBFB1CE123FB75" box="[538,585,1161,1188]" gridcol="4" gridrow="23" pageId="13" pageNumber="265">29.0</td>
<td id="76FC001EEA084BABFDA3FB1CE1E0FB75" box="[610,650,1161,1188]" gridcol="5" gridrow="23" pageId="13" pageNumber="265">7.5</td>
<td id="76FC001EEA084BABFD65FB1CE1BEFB75" box="[676,724,1161,1188]" gridcol="6" gridrow="23" pageId="13" pageNumber="265">28.8</td>
<td id="76FC001EEA084BABFD2CFB1CE07FFB75" box="[749,789,1161,1188]" gridcol="7" gridrow="23" pageId="13" pageNumber="265">6.6</td>
<td id="76FC001EEA084BABFCEFFB1CE030FB75" box="[814,858,1161,1188]" gridcol="8" gridrow="23" pageId="13" pageNumber="265">15.5</td>
<td id="76FC001EEA084BABFCB3FB1CE0C9FB75" box="[882,931,1161,1188]" gridcol="9" gridrow="23" pageId="13" pageNumber="265">19.5</td>
<td id="76FC001EEA084BABFC7FFB1CE723FB75" box="[958,1097,1161,1188]" gridcol="10" gridrow="23" pageId="13" pageNumber="265">150</td>
<td id="76FC001EEA084BABFBA0FB1CE7F4FB75" box="[1121,1182,1161,1188]" gridcol="11" gridrow="23" pageId="13" pageNumber="265">0</td>
</tr>
<tr id="352D6962EA084BABFFBFFB38E7F4FB19" box="[126,1182,1197,1224]" gridrow="24" pageId="13" pageNumber="265">
<th id="76FC001EEA084BABFFBFFB38E38BFB19" box="[126,225,1197,1224]" gridcol="0" gridrow="24" pageId="13" pageNumber="265">22</th>
<td id="76FC001EEA084BABFE83FB38E218FB19" box="[322,370,1197,1224]" gridcol="1" gridrow="24" pageId="13" pageNumber="265">76</td>
<td id="76FC001EEA084BABFE4BFB38E2D5FB19" box="[394,447,1197,1224]" gridcol="2" gridrow="24" pageId="13" pageNumber="265">83</td>
<td id="76FC001EEA084BABFE16FB38E16BFB19" box="[471,513,1197,1224]" gridcol="3" gridrow="24" pageId="13" pageNumber="265">40</td>
<td id="76FC001EEA084BABFDDBFB38E123FB19" box="[538,585,1197,1224]" gridcol="4" gridrow="24" pageId="13" pageNumber="265">28.6</td>
<td id="76FC001EEA084BABFDA3FB38E1E0FB19" box="[610,650,1197,1224]" gridcol="5" gridrow="24" pageId="13" pageNumber="265">6.5</td>
<td id="76FC001EEA084BABFD65FB38E1BEFB19" box="[676,724,1197,1224]" gridcol="6" gridrow="24" pageId="13" pageNumber="265">28.8</td>
<td id="76FC001EEA084BABFD2CFB38E07FFB19" box="[749,789,1197,1224]" gridcol="7" gridrow="24" pageId="13" pageNumber="265">7.7</td>
<td id="76FC001EEA084BABFCEFFB38E030FB19" box="[814,858,1197,1224]" gridcol="8" gridrow="24" pageId="13" pageNumber="265">14.4</td>
<td id="76FC001EEA084BABFCB3FB38E0C9FB19" box="[882,931,1197,1224]" gridcol="9" gridrow="24" pageId="13" pageNumber="265">22.2</td>
<td id="76FC001EEA084BABFC7FFB38E723FB19" box="[958,1097,1197,1224]" gridcol="10" gridrow="24" pageId="13" pageNumber="265">155</td>
<td id="76FC001EEA084BABFBA0FB38E7F4FB19" box="[1121,1182,1197,1224]" gridcol="11" gridrow="24" pageId="13" pageNumber="265">3</td>
</tr>
<tr id="352D6962EA084BABFFBFFB44E7F4FB3D" box="[126,1182,1233,1260]" gridrow="25" pageId="13" pageNumber="265">
<th id="76FC001EEA084BABFFBFFB44E38BFB3D" box="[126,225,1233,1260]" gridcol="0" gridrow="25" pageId="13" pageNumber="265">23</th>
<td id="76FC001EEA084BABFE83FB44E218FB3D" box="[322,370,1233,1260]" gridcol="1" gridrow="25" pageId="13" pageNumber="265">74</td>
<td id="76FC001EEA084BABFE4BFB44E2D5FB3D" box="[394,447,1233,1260]" gridcol="2" gridrow="25" pageId="13" pageNumber="265">80</td>
<td id="76FC001EEA084BABFE16FB44E16BFB3D" box="[471,513,1233,1260]" gridcol="3" gridrow="25" pageId="13" pageNumber="265">42</td>
<td id="76FC001EEA084BABFDDBFB44E123FB3D" box="[538,585,1233,1260]" gridcol="4" gridrow="25" pageId="13" pageNumber="265">28.2</td>
<td id="76FC001EEA084BABFDA3FB44E1E0FB3D" box="[610,650,1233,1260]" gridcol="5" gridrow="25" pageId="13" pageNumber="265">6.8</td>
<td id="76FC001EEA084BABFD65FB44E1BEFB3D" box="[676,724,1233,1260]" gridcol="6" gridrow="25" pageId="13" pageNumber="265">26.3</td>
<td id="76FC001EEA084BABFD2CFB44E07FFB3D" box="[749,789,1233,1260]" gridcol="7" gridrow="25" pageId="13" pageNumber="265">6.3</td>
<td id="76FC001EEA084BABFCEFFB44E030FB3D" box="[814,858,1233,1260]" gridcol="8" gridrow="25" pageId="13" pageNumber="265">14.6</td>
<td id="76FC001EEA084BABFCB3FB44E0C9FB3D" box="[882,931,1233,1260]" gridcol="9" gridrow="25" pageId="13" pageNumber="265">19.1</td>
<td id="76FC001EEA084BABFC7FFB44E723FB3D" box="[958,1097,1233,1260]" gridcol="10" gridrow="25" pageId="13" pageNumber="265">152</td>
<td id="76FC001EEA084BABFBA0FB44E7F4FB3D" box="[1121,1182,1233,1260]" gridcol="11" gridrow="25" pageId="13" pageNumber="265">3</td>
</tr>
<tr id="352D6962EA084BABFFBFFB61E7F4FADE" box="[126,1182,1268,1295]" gridrow="26" pageId="13" pageNumber="265">
<th id="76FC001EEA084BABFFBFFB61E38BFADE" box="[126,225,1268,1295]" gridcol="0" gridrow="26" pageId="13" pageNumber="265">24</th>
<td id="76FC001EEA084BABFE83FB61E218FADE" box="[322,370,1268,1295]" gridcol="1" gridrow="26" pageId="13" pageNumber="265">71</td>
<td id="76FC001EEA084BABFE4BFB61E2D5FADE" box="[394,447,1268,1295]" gridcol="2" gridrow="26" pageId="13" pageNumber="265">78</td>
<td id="76FC001EEA084BABFE16FB61E16BFADE" box="[471,513,1268,1295]" gridcol="3" gridrow="26" pageId="13" pageNumber="265">38</td>
<td id="76FC001EEA084BABFDDBFB61E123FADE" box="[538,585,1268,1295]" gridcol="4" gridrow="26" pageId="13" pageNumber="265">25.3</td>
<td id="76FC001EEA084BABFDA3FB61E1E0FADE" box="[610,650,1268,1295]" gridcol="5" gridrow="26" pageId="13" pageNumber="265">7.1</td>
<td id="76FC001EEA084BABFD65FB61E1BEFADE" box="[676,724,1268,1295]" gridcol="6" gridrow="26" pageId="13" pageNumber="265">25.5</td>
<td id="76FC001EEA084BABFD2CFB61E07FFADE" box="[749,789,1268,1295]" gridcol="7" gridrow="26" pageId="13" pageNumber="265">6.4</td>
<td id="76FC001EEA084BABFCEFFB61E030FADE" box="[814,858,1268,1295]" gridcol="8" gridrow="26" pageId="13" pageNumber="265">14.3</td>
<td id="76FC001EEA084BABFCB3FB61E0C9FADE" box="[882,931,1268,1295]" gridcol="9" gridrow="26" pageId="13" pageNumber="265">19.5</td>
<td id="76FC001EEA084BABFC7FFB61E723FADE" box="[958,1097,1268,1295]" gridcol="10" gridrow="26" pageId="13" pageNumber="265">147</td>
<td id="76FC001EEA084BABFBA0FB61E7F4FADE" box="[1121,1182,1268,1295]" gridcol="11" gridrow="26" pageId="13" pageNumber="265">0</td>
</tr>
<tr id="352D6962EA084BABFFBFFAB6E7F4FAEF" box="[126,1182,1315,1342]" gridrow="27" pageId="13" pageNumber="265">
<th id="76FC001EEA084BABFFBFFAB6E7F4FAEF" box="[126,1182,1315,1342]" colspan="12" colspanRight="11" gridcol="0" gridrow="27" pageId="13" pageNumber="265"># Tail damaged</th>
</tr>
</table>
</paragraph>
<paragraph id="8BA26B20EA08B459FF5EFA01E7C9FA03" blockId="13.[129,640,1428,1742]" lastBlockId="13.[676,1219,1428,1746]" pageId="13" pageNumber="265">
First
<typeStatus id="54A6D582EA08B459FF1BFA00E257FA7E" box="[218,317,1429,1455]" pageId="13" pageNumber="265" type="paratype">paratype</typeStatus>
MNCN 51793. Adult male in breeding condition with a well-developed crest and a large back cloaca. Seventeen crème-white bands over the head and body part of the crest. SVl1 65.0 mm, SVl2
<quantity id="4CE5C6C5EA08B459FDDCF9B1E111F9EF" box="[541,635,1572,1599]" metricMagnitude="-2" metricUnit="m" metricValue="7.35" pageId="13" pageNumber="265" unit="mm" value="73.5">73.5 mm</quantity>
, ILd 34.0 mm, FLl
<quantity id="4CE5C6C5EA08B459FE92F9DDE2DEF9B3" box="[339,436,1608,1634]" metricMagnitude="-2" metricUnit="m" metricValue="2.62" pageId="13" pageNumber="265" unit="mm" value="26.2">26.2 mm</quantity>
, TFl
<quantity id="4CE5C6C5EA08B459FE32F9DDE12DF9B3" box="[499,583,1608,1634]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="13" pageNumber="265" unit="mm" value="9.3">9.3 mm</quantity>
, HLl 27.0 mm, FTl
<quantity id="4CE5C6C5EA08B459FEE7F9FEE214F957" box="[294,382,1643,1670]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="13" pageNumber="265" unit="mm" value="11.5">11.5 mm</quantity>
, Hw
<quantity id="4CE5C6C5EA08B459FE05F9FEE175F957" box="[452,543,1643,1670]" metricMagnitude="-2" metricUnit="m" metricValue="1.3599999999999999" pageId="13" pageNumber="265" unit="mm" value="13.6">13.6 mm</quantity>
and Hl
<quantity id="4CE5C6C5EA08B459FF40F905E3B5F97B" box="[129,223,1680,1706]" metricMagnitude="-2" metricUnit="m" metricValue="1.9899999999999998" pageId="13" pageNumber="265" unit="mm" value="19.9">19.9 mm</quantity>
. Total length
<quantity id="4CE5C6C5EA08B459FE4BF91AE28BF97B" box="[394,481,1679,1706]" metricMagnitude="-1" metricUnit="m" metricValue="1.35" pageId="13" pageNumber="265" unit="mm" value="135.0">135 mm</quantity>
. Relative finger lengths 1=4&lt;2&lt;3. Relative toe lengths 1&lt;5&lt;2&lt;4&lt;3.
<subScript id="17996965EA08B459FCFDFA36E008FA67" attach="left" box="[828,866,1443,1462]" fontSize="8" pageId="13" pageNumber="265">Nlinks</subScript>
left 0, right 1, total 1. Grey ventral colouration with many dark dots.
</paragraph>
<paragraph id="8BA26B20EA08B459FD65FA48E196F957" blockId="13.[676,1219,1428,1746]" pageId="13" pageNumber="265">Many white spots mostly positioned on the undotted sections. Colour of the throat continuous with that of the belly. Underside of the legs and cloacal region with light and dark regions.</paragraph>
<paragraph id="8BA26B20EA08B45AFD03F91AE108FACF" blockId="13.[676,1219,1428,1746]" lastBlockId="14.[98,610,854,1705]" lastPageId="14" lastPageNumber="266" pageId="13" pageNumber="265">
Second
<typeStatus id="54A6D582EA08B459FCDCF905E0EAF97B" box="[797,896,1680,1706]" pageId="13" pageNumber="265" type="paratype">paratype</typeStatus>
MNCN 51794. Adult male in breeding condition with a low crest and a large back cloaca. Eleven crème-white bands over the head and body part of the low crest. SVl1
<quantity id="4CE5C6C5EA0BB45AFF5FFC08E395FC69" box="[158,255,925,952]" metricMagnitude="-2" metricUnit="m" metricValue="5.65" pageId="14" pageNumber="266" unit="mm" value="56.5">56.5 mm</quantity>
, SVl2 64.0 mm, ILd
<quantity id="4CE5C6C5EA0BB45AFE3CFC08E137FC69" box="[509,605,925,952]" metricMagnitude="-2" metricUnit="m" metricValue="2.85" pageId="14" pageNumber="266" unit="mm" value="28.5">28.5 mm</quantity>
, FLl
<quantity id="4CE5C6C5EA0BB45AFF53FC54E398FC0D" box="[146,242,961,988]" metricMagnitude="-2" metricUnit="m" metricValue="2.34" pageId="14" pageNumber="266" unit="mm" value="23.4">23.4 mm</quantity>
, TFl
<quantity id="4CE5C6C5EA0BB45AFEF2FC57E2ECFC0D" box="[307,390,962,988]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="14" pageNumber="266" unit="mm" value="7.7">7.7 mm</quantity>
, HLl
<quantity id="4CE5C6C5EA0BB45AFE0DFC54E146FC0D" box="[460,556,961,988]" metricMagnitude="-2" metricUnit="m" metricValue="2.36" pageId="14" pageNumber="266" unit="mm" value="23.6">23.6 mm</quantity>
, FTl
<quantity id="4CE5C6C5EA0BB45AFFA3FC70E3DDFBD1" box="[98,183,997,1024]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="14" pageNumber="266" unit="mm" value="9.6">9.6 mm</quantity>
, Hw
<quantity id="4CE5C6C5EA0BB45AFF3BFC70E23FFBD1" box="[250,341,997,1024]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="14" pageNumber="266" unit="mm" value="12.6">12.6 mm</quantity>
and Hl
<quantity id="4CE5C6C5EA0BB45AFE7FFC70E171FBD1" box="[446,539,997,1024]" metricMagnitude="-2" metricUnit="m" metricValue="1.77" pageId="14" pageNumber="266" unit="mm" value="17.7">17.7 mm</quantity>
. Total length
<quantity id="4CE5C6C5EA0BB45AFF72FB9CE26CFBF5" box="[179,262,1033,1060]" metricMagnitude="-1" metricUnit="m" metricValue="1.18" pageId="14" pageNumber="266" unit="mm" value="118.0">118 mm</quantity>
. Tail tip regenerating. Relative finger lengths 1&lt;4&lt;2&lt;3. Relative toe lengths 1&lt;5&lt;2&lt;4&lt;3.
<subScript id="17996965EA0BB45AFF3BFBF5E24AFBA2" attach="left" box="[250,288,1120,1139]" fontSize="8" pageId="14" pageNumber="266">Nlinks</subScript>
left 1, right 2, total 3. Solid black ventral colouration with many, evenly distributed small white spots. Throat colouration continuous with belly with equally dense but larger white spots. Underside of the legs and cloacal region light with few dark regions.
</paragraph>
<footnote id="E806772EEA08B459FD3EF956E7A9F8C3" pageId="13" pageNumber="265">
<paragraph id="8BA26B20EA08B459FD3EF956E7A9F8C3" blockId="13.[676,1219,1428,1746]" lastBlockId="13.[498,1220,1752,1810]" pageId="13" pageNumber="265">Downloaded from Brill.com 07/10/2024 02:02:47PM via Open Access. This is an open access article distributed under the terms of the CC BY 4.0 license. https://creativecommons.org/licenses/by/4.0/</paragraph>
</footnote>
<caption id="DF623BA8EA0BB45AFFA3FD22E74AFD3B" ID-DOI="http://doi.org/10.5281/zenodo.13153858" ID-Zenodo-Dep="13153858" httpUri="https://zenodo.org/record/13153858/files/figure.png" pageId="14" pageNumber="266" startId="14.[98,165,695,716]" targetBox="[243,1011,164,667]" targetPageId="14" targetType="figure">
<paragraph id="8BA26B20EA0BB45AFFA3FD22E74AFD3B" blockId="14.[98,1148,695,747]" pageId="14" pageNumber="266">
FIGURE A1 Holotype of
<taxonomicName id="4C1D10A3EA0BB45AFE8CFD2DE118FD1D" authorityName="Arntzen" authorityYear="2024" box="[333,626,695,717]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="14" pageNumber="266" phylum="Chordata" rank="subSpecies" species="marmoratus" status="subsp. nov." subSpecies="harmannis">
<emphasis id="B969B732EA0BB45AFE8CFD2DE118FD1D" box="[333,626,695,717]" italics="true" pageId="14" pageNumber="266">Triturus marmoratus harmannis</emphasis>
</taxonomicName>
<emphasis id="B969B732EA0BB45AFDB6FD22E1AEFD1D" bold="true" box="[631,708,695,716]" pageId="14" pageNumber="266">
<taxonomicNameLabel id="A25A0A49EA0BB45AFDB6FD22E1AEFD1D" box="[631,708,695,716]" pageId="14" pageNumber="266" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
</emphasis>
at right and ventral view. Size bar is 1 cm. Stored at the Museo Nacional de Ciencias Naturales, Madrid, Spain under catalogue number 51792.
</paragraph>
</caption>
<paragraph id="8BA26B20EA0BB45AFF40FABDE198FC45" blockId="14.[98,610,854,1705]" lastBlockId="14.[645,1157,854,1706]" pageId="14" pageNumber="266">
Third
<typeStatus id="54A6D582EA0BB45AFF06FABCE240FA92" box="[199,298,1321,1347]" pageId="14" pageNumber="266" type="paratype">paratype</typeStatus>
MNCN 51795. Adult female in breeding condition with ovaries shining through. SVl1 64.0 mm, SVl2 70.0 mm, ILd 33.0 mm, FLl
<quantity id="4CE5C6C5EA0BB45AFEF6FA01E2F9FA7E" box="[311,403,1428,1455]" metricMagnitude="-2" metricUnit="m" metricValue="2.51" pageId="14" pageNumber="266" unit="mm" value="25.1">25.1 mm</quantity>
, TFl
<quantity id="4CE5C6C5EA0BB45AFE15FA01E142FA7E" box="[468,552,1428,1455]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="14" pageNumber="266" unit="mm" value="8.3">8.3 mm</quantity>
, HLl
<quantity id="4CE5C6C5EA0BB45AFFA3FA2DE3ABFA02" box="[98,193,1464,1491]" metricMagnitude="-2" metricUnit="m" metricValue="2.37" pageId="14" pageNumber="266" unit="mm" value="23.7">23.7 mm</quantity>
, FTl
<quantity id="4CE5C6C5EA0BB45AFEC9FA2DE236FA02" box="[264,348,1464,1491]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="14" pageNumber="266" unit="mm" value="7.6">7.6 mm</quantity>
, Hw
<quantity id="4CE5C6C5EA0BB45AFE65FA2DE295FA02" box="[420,511,1464,1491]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="14" pageNumber="266" unit="mm" value="14.5">14.5 mm</quantity>
and Hl
<quantity id="4CE5C6C5EA0BB45AFFA3FA49E3ABFA27" box="[98,193,1500,1526]" metricMagnitude="-2" metricUnit="m" metricValue="1.96" pageId="14" pageNumber="266" unit="mm" value="19.6">19.6 mm</quantity>
. Total length
<quantity id="4CE5C6C5EA0BB45AFE42FA49E285FA27" box="[387,495,1500,1527]" metricMagnitude="-1" metricUnit="m" metricValue="1.405" pageId="14" pageNumber="266" unit="mm" value="140.5">140.5 mm</quantity>
. Relative finger lengths 1&lt;4&lt;2&lt;3. Relative toe lengths 1=5&lt;2&lt;4&lt;3.
<subScript id="17996965EA0BB45AFF38F9A6E275F997" attach="left" box="[249,287,1587,1606]" fontSize="8" pageId="14" pageNumber="266">Nlinks</subScript>
left 2, right 2, total 4. Solid, medium-grey ventral colouration with few, evenly distributed small white spots. Throat coloration continuous with that of the belly. Underside of the legs and cloacal region solid light grey.
</paragraph>
<paragraph id="8BA26B20EA0BB45AFD65FC08E1B4FAB7" blockId="14.[645,1157,854,1706]" pageId="14" pageNumber="266">
Fourth
<typeStatus id="54A6D582EA0BB45AFD34FC0BE032FC69" box="[757,856,926,952]" pageId="14" pageNumber="266" type="paratype">paratype</typeStatus>
MNCN 51796. Adult female in breeding condition with ovaries shining through. SVl1 73.0 mm, SVl2
<quantity id="4CE5C6C5EA0BB45AFBE0FC70E7EAFBD1" box="[1057,1152,997,1024]" metricMagnitude="-2" metricUnit="m" metricValue="7.95" pageId="14" pageNumber="266" unit="mm" value="79.5">79.5 mm</quantity>
, ILd
<quantity id="4CE5C6C5EA0BB45AFD79FB9CE072FBF5" box="[696,792,1033,1060]" metricMagnitude="-2" metricUnit="m" metricValue="3.95" pageId="14" pageNumber="266" unit="mm" value="39.5">39.5 mm</quantity>
, FLl
<quantity id="4CE5C6C5EA0BB45AFC98FB9CE0DCFBF5" box="[857,950,1033,1060]" metricMagnitude="-2" metricUnit="m" metricValue="2.81" pageId="14" pageNumber="266" unit="mm" value="28.1">28.1 mm</quantity>
, TFl
<quantity id="4CE5C6C5EA0BB45AFC36FB9CE721FBF5" box="[1015,1099,1033,1060]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="14" pageNumber="266" unit="mm" value="9.3">9.3 mm</quantity>
, HLl
<quantity id="4CE5C6C5EA0BB45AFD44FBB8E18DFB99" box="[645,743,1069,1096]" metricMagnitude="-2" metricUnit="m" metricValue="2.6399999999999997" pageId="14" pageNumber="266" unit="mm" value="26.4">26.4 mm</quantity>
, FTl
<quantity id="4CE5C6C5EA0BB45AFCEDFBB8E0EBFB99" box="[812,897,1069,1096]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="14" pageNumber="266" unit="mm" value="8.8">8.8 mm</quantity>
, Hw
<quantity id="4CE5C6C5EA0BB45AFC09FBB8E749FB99" box="[968,1059,1069,1096]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="14" pageNumber="266" unit="mm" value="16.5">16.5 mm</quantity>
and Hl
<quantity id="4CE5C6C5EA0BB45AFD44FBC4E188FBBD" box="[645,738,1105,1132]" metricMagnitude="-2" metricUnit="m" metricValue="2.17" pageId="14" pageNumber="266" unit="mm" value="21.7">21.7 mm</quantity>
. Total length
<quantity id="4CE5C6C5EA0BB45AFC4CFBC4E08CFBBD" box="[909,998,1105,1132]" metricMagnitude="-1" metricUnit="m" metricValue="1.49" pageId="14" pageNumber="266" unit="mm" value="149.0">149 mm</quantity>
. Relative finger lengths 1&lt;4&lt;2&lt;3. Relative toe lengths 1&lt;5&lt;2&lt;4&lt;3.
<subScript id="17996965EA0BB45AFCCBFB0CE028FB6A" attach="left" box="[778,834,1177,1211]" fontSize="8" pageId="14" pageNumber="266">Nlinks</subScript>
left 0, right 0, total 0. Dark grey ventral colouration with few small dark dots and few white spots. Colouration of throat continuous with belly. Underside of the legs and cloacal region light with few dark regions.
</paragraph>
<paragraph id="8BA26B20EA0BB45BFD65FAE5E212FEE0" blockId="14.[645,1157,854,1706]" lastBlockId="15.[129,642,171,1455]" lastPageId="15" lastPageNumber="267" pageId="14" pageNumber="266">
Fifth
<typeStatus id="54A6D582EA0BB45AFD25FAE4E02DFA5A" box="[740,839,1393,1419]" pageId="14" pageNumber="266" type="paratype">paratype</typeStatus>
MNCN 51797. Adult female in breeding condition with ovaries shining through. SVl1 71.0 mm, SVl2
<quantity id="4CE5C6C5EA0BB45AFBE0FA2DE7EAFA02" box="[1057,1152,1464,1491]" metricMagnitude="-2" metricUnit="m" metricValue="7.75" pageId="14" pageNumber="266" unit="mm" value="77.5">77.5 mm</quantity>
, ILd
<quantity id="4CE5C6C5EA0BB45AFD79FA49E07DFA26" box="[696,791,1500,1527]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" pageId="14" pageNumber="266" unit="mm" value="37.5">37.5 mm</quantity>
, FLl
<quantity id="4CE5C6C5EA0BB45AFC98FA49E0D3FA26" box="[857,953,1500,1527]" metricMagnitude="-2" metricUnit="m" metricValue="2.7399999999999998" pageId="14" pageNumber="266" unit="mm" value="27.4">27.4 mm</quantity>
, TFl
<quantity id="4CE5C6C5EA0BB45AFC3AFA49E721FA26" box="[1019,1099,1500,1527]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="14" pageNumber="266" unit="mm" value="9.1">9.1 mm</quantity>
, HLl 26.0 mm, FTl
<quantity id="4CE5C6C5EA0BB45AFCEFF995E0EBF9CB" box="[814,897,1536,1562]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="14" pageNumber="266" unit="mm" value="7.2">7.2 mm</quantity>
, Hw
<quantity id="4CE5C6C5EA0BB45AFC09F995E748F9CB" box="[968,1058,1536,1563]" metricMagnitude="-2" metricUnit="m" metricValue="1.54" pageId="14" pageNumber="266" unit="mm" value="15.4">15.4 mm</quantity>
and Hl
<quantity id="4CE5C6C5EA0BB45AFD44F9B1E18CF9EF" box="[645,742,1572,1598]" metricMagnitude="-2" metricUnit="m" metricValue="2.2199999999999998" pageId="14" pageNumber="266" unit="mm" value="22.2">22.2 mm</quantity>
. Total length
<quantity id="4CE5C6C5EA0BB45AFC66F9B1E778F9EF" box="[935,1042,1572,1599]" metricMagnitude="-1" metricUnit="m" metricValue="1.505" pageId="14" pageNumber="266" unit="mm" value="150.5">150.5 mm</quantity>
. Relative finger lengths 1=4&lt;2&lt;3. Relative toe lengths 1=5&lt;2&lt;4&lt;3.
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left 1, right 2, total 3. Light grey ventral colouration with large dark dots and few white spots towards the flanks. Colouration of throat continuous with belly. Underside of the legs and cloacal region light with few dark regions.
</paragraph>
<paragraph id="8BA26B20EA0AB45BFF5EFEAEE212FCD5" blockId="15.[129,642,171,1455]" pageId="15" pageNumber="267">
Sixth
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MNCN 51798. Adult female in breeding condition with ovaries shining through. SVl1 72.0 mm, SVl2
<quantity id="4CE5C6C5EA0AB45BFDDDFE16E111FE4C" box="[540,635,387,414]" metricMagnitude="-2" metricUnit="m" metricValue="7.85" pageId="15" pageNumber="267" unit="mm" value="78.5">78.5 mm</quantity>
, ILd
<quantity id="4CE5C6C5EA0AB45BFF73FE32E27BFE10" box="[178,273,423,450]" metricMagnitude="-2" metricUnit="m" metricValue="3.85" pageId="15" pageNumber="267" unit="mm" value="38.5">38.5 mm</quantity>
, FLl 28.0 mm, TFl 9.0 mm, HLl
<quantity id="4CE5C6C5EA0AB45BFF40FE5EE38AFE34" box="[129,224,459,485]" metricMagnitude="-2" metricUnit="m" metricValue="2.7199999999999998" pageId="15" pageNumber="267" unit="mm" value="27.2">27.2 mm</quantity>
, FTl
<quantity id="4CE5C6C5EA0AB45BFEE6FE5EE211FE34" box="[295,379,459,485]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="15" pageNumber="267" unit="mm" value="8.7">8.7 mm</quantity>
, Hw
<quantity id="4CE5C6C5EA0AB45BFE04FE5EE176FE34" box="[453,540,459,485]" metricMagnitude="-2" metricUnit="m" metricValue="1.61" pageId="15" pageNumber="267" unit="mm" value="16.1">16.1 mm</quantity>
and Hl
<quantity id="4CE5C6C5EA0AB45BFF40FE7AE3B3FDD8" box="[129,217,495,521]" metricMagnitude="-2" metricUnit="m" metricValue="2.11" pageId="15" pageNumber="267" unit="mm" value="21.1">21.1 mm</quantity>
. Total length
<quantity id="4CE5C6C5EA0AB45BFE46FE7BE2B5FDD8" box="[391,479,494,521]" metricMagnitude="-1" metricUnit="m" metricValue="1.46" pageId="15" pageNumber="267" unit="mm" value="146.0">146 mm</quantity>
. Relative finger lengths 1=4&lt;2&lt;3. Relative toe lengths 1&lt;5&lt;2&lt;3=4.
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left 2, right 2, total 4. Light grey ventral colouration with medium number of large dark dots and few white spots. Colouration of throat continuous with belly. Underside of the legs and cloacal region light with few dark regions.
</paragraph>
</subSubSection>
<subSubSection id="C30738ABEA0AB45BFF5EFC9BE246FC0A" pageId="15" pageNumber="267" type="materials_examined">
<paragraph id="8BA26B20EA0AB45BFF5EFC9BE246FC0A" blockId="15.[129,642,171,1455]" pageId="15" pageNumber="267">
<materialsCitation id="3B75617DEA0AB45BFF5EFC9BE246FC0A" collectingDate="2013-03-20" collectionCode="MNCN" collectorName="J. W. Arntzen." country="Portugal" county="Elevation" elevation="396" location="Arrochela" municipality="Arrochela" pageId="15" pageNumber="267" specimenCount="1" stateProvince="Madeira">
Locality and date of collecting
<collectingMunicipality id="6BC6F15AEA0AB45BFDCAFC9BE116FCF9" box="[523,636,782,808]" pageId="15" pageNumber="267">Arrochela</collectingMunicipality>
, near
<collectingRegion id="49D9A5C2EA0AB45BFF76FCA7E27CFC9D" box="[183,278,818,844]" country="Portugal" name="Madeira" pageId="15" pageNumber="267">Madeirã</collectingRegion>
,
<collectingCountry id="F30A2BB0EA0AB45BFEDFFCA7E216FC9D" box="[286,380,818,844]" name="Portugal" pageId="15" pageNumber="267">Portugal</collectingCountry>
at 39.9386northern latitude and 8.1025 western longitude. Elevation
<elevation id="00308C13EA0AB45BFF40FCECE394FC42" box="[129,254,889,916]" metricMagnitude="2" metricUnit="m" metricValue="3.96" pageId="15" pageNumber="267" unit="m" value="396.0">
<quantity id="4CE5C6C5EA0AB45BFF40FCECE3ADFC45" box="[129,199,889,916]" metricMagnitude="2" metricUnit="m" metricValue="3.96" pageId="15" pageNumber="267" unit="m" value="396.0">396 m</quantity>
a.s.l.
</elevation>
Date of collecting
<date id="FFA34DE0EA0AB45BFE16FCEFE116FC45" box="[471,636,889,916]" pageId="15" pageNumber="267" value="2013-03-20">
<collectingDate id="EFE7B408EA0AB45BFE16FCEFE116FC45" box="[471,636,889,916]" pageId="15" pageNumber="267" value="2013-03-20">20 March 2013</collectingDate>
</date>
, leg.
<collectorName id="26E80EF6EA0AB45BFF6DFC0BE229FC69" box="[172,323,926,952]" pageId="15" pageNumber="267">J. W. Arntzen.</collectorName>
Date of deposition at
<collectionCode id="ED0CF3E5EA0AB45BFDF9FC0BE1EAFC69" box="[568,640,926,952]" pageId="15" pageNumber="267">MNCN</collectionCode>
<date id="FFA34DE0EA0AB45BFF40FC54E24DFC0A" box="[129,295,961,988]" pageId="15" pageNumber="267" value="2023-08-16">16 August 2023</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30738ABEA0AB45BFF5EFC70E149FAB7" pageId="15" pageNumber="267" type="diagnosis">
<paragraph id="8BA26B20EA0AB45BFF5EFC70E149FAB7" blockId="15.[129,642,171,1455]" pageId="15" pageNumber="267">
Diagnostic features the newly recognized subspecies is closely related to and morphometrically similar to
<emphasis id="B969B732EA0AB45BFEACFBBBE159FB96" box="[365,563,1070,1095]" italics="true" pageId="15" pageNumber="267">
T. m.
<taxonomicName id="4C1D10A3EA0AB45BFE68FBBBE159FB96" baseAuthorityName="Latreille" baseAuthorityYear="1800" box="[425,563,1070,1095]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="15" pageNumber="267" phylum="Chordata" rank="species" species="marmoratus">marmoratus</taxonomicName>
</emphasis>
, yet at the population level characterized by a lower number of green coloured dorso-lateral transversal bands (links). In comparison with
<emphasis id="B969B732EA0AB45BFDADFB0FE244FB07" italics="true" pageId="15" pageNumber="267">
T. m.
<taxonomicName id="4C1D10A3EA0AB45BFF65FB28E244FB07" baseAuthorityName="Latreille" baseAuthorityYear="1800" box="[164,302,1213,1238]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="15" pageNumber="267" phylum="Chordata" rank="species" species="marmoratus">marmoratus</taxonomicName>
</emphasis>
, it has a small body size, significantly shorter extremities in females and a relatively big head in both sexes. The distinctiveness of the subspecies is supported by nuclear and mitochondrial DNA data.
</paragraph>
</subSubSection>
<subSubSection id="C30738ABEA0AB45BFF5EFAE4E192FD31" pageId="15" pageNumber="267" type="etymology">
<paragraph id="8BA26B20EA0AB45BFF5EFAE4E192FD31" blockId="15.[129,642,171,1455]" lastBlockId="15.[676,1188,171,1454]" pageId="15" pageNumber="267">
<emphasis id="B969B732EA0AB45BFF5EFAE4E20CFA5B" box="[159,358,1393,1418]" italics="true" pageId="15" pageNumber="267">Derivatio nominis</emphasis>
the subspecies name is chosen to commemorate the Dutch couple Harm and Annie (or Ann) Walen, who lived from 19262005 (Mr. H. C. Walen) and 1928 2018 (Mrs. A. A. van Silfhout). After Harms retirement from his taxidermist job at the Zoological Museum in Amsterdam, the twosome eventually landed in Nisa,
<collectingCountry id="F30A2BB0EA0AB45BFB82FECAE7C9FEA8" box="[1091,1187,351,377]" name="Portugal" pageId="15" pageNumber="267">Portugal</collectingCountry>
where they constructed their own quinta and felt enormously at place. Without harmanns hospitality and moral support, my extensive fieldwork in
<collectingCountry id="F30A2BB0EA0AB45BFC66FE7BE76DFDD9" box="[935,1031,494,520]" name="Portugal" pageId="15" pageNumber="267">Portugal</collectingCountry>
and adjacent
<collectingCountry id="F30A2BB0EA0AB45BFD65FD87E18EFDFD" box="[676,740,530,556]" name="Spain" pageId="15" pageNumber="267">Spain</collectingCountry>
late last century would hardly have been sustainable. The new subspecies name refers to a lifelong couple and only indirectly to individual people so that, matrimony being gender neutral, the third declension is used.
</paragraph>
</subSubSection>
<subSubSection id="C30738ABEA0AB45BFD03FD7FE0DCFBBA" pageId="15" pageNumber="267" type="vernacular_names">
<paragraph id="8BA26B20EA0AB45BFD03FD7FE0DCFBBA" blockId="15.[676,1188,171,1454]" pageId="15" pageNumber="267">
<location id="8EC23DFBEA0AB45BFD03FD7FE058FCD5" LSID="urn:lsid:plazi:treatment:03B4DA36EA09B45BFD44FE84E029FA7F:8EC23DFBEA0AB45BFD03FD7FE058FCD5" box="[706,818,746,772]" country="Georgia" county="The" latitude="41.5" municipality="Distribution" name="Suggested" pageId="15" pageNumber="267">Suggested</location>
vernacular name
<location id="8EC23DFBEA0AB45BFBE8FD7FE7C9FCD5" LSID="urn:lsid:plazi:treatment:03B4DA36EA09B45BFD44FE84E029FA7F:8EC23DFBEA0AB45BFBE8FD7FE7C9FCD5" box="[1065,1187,746,772]" country="Georgia" county="The" latitude="41.5" municipality="Distribution" name="Harmann's" pageId="15" pageNumber="267">Harmanns</location>
marbled newt, or
<location id="8EC23DFBEA0AB45BFC47FC9BE75FFCF9" LSID="urn:lsid:plazi:treatment:03B4DA36EA09B45BFD44FE84E029FA7F:8EC23DFBEA0AB45BFC47FC9BE75FFCF9" box="[902,1077,782,808]" country="Georgia" county="The" latitude="41.5" municipality="Distribution" name="central Iberian" pageId="15" pageNumber="267">central Iberian</location>
marbled newt
<collectingMunicipality id="6BC6F15AEA0AB45BFD2BFCA7E01EFC9D" box="[746,884,818,844]" pageId="15" pageNumber="267">Distribution</collectingMunicipality>
central-western
<collectingCountry id="F30A2BB0EA0AB45BFB9AFCA7E7CAFC9D" box="[1115,1184,818,844]" name="Georgia" pageId="15" pageNumber="267">Iberia</collectingCountry>
.
<collectingCounty id="62C313ACEA0AB45BFD65FCC0E1A5FCBE" box="[676,719,853,879]" pageId="15" pageNumber="267">The</collectingCounty>
southern range border is determined by a sharp, parapatric, yet mosaic range border with two pygmy newt species, namely
<taxonomicName id="4C1D10A3EA0AB45BFD65FC57E062FC0A" authorityName="Arntzen" authorityYear="2024" box="[676,776,961,987]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="15" pageNumber="267" phylum="Chordata" rank="species" species="rudolfi">
<emphasis id="B969B732EA0AB45BFD65FC57E062FC0A" box="[676,776,961,987]" italics="true" pageId="15" pageNumber="267">T. rudolfi</emphasis>
</taxonomicName>
in the west and
<taxonomicName id="4C1D10A3EA0AB45BFC0DFC57E73CFC0A" baseAuthorityName="Wolterstorff" baseAuthorityYear="1905" box="[972,1110,962,987]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="15" pageNumber="267" phylum="Chordata" rank="species" species="pygmaeus">
<emphasis id="B969B732EA0AB45BFC0DFC57E73CFC0A" box="[972,1110,962,987]" italics="true" pageId="15" pageNumber="267">T. pygmaeus</emphasis>
</taxonomicName>
in the centre of the Iberian Peninsula. The northern edge of the range is positioned at ca.
<geoCoordinate id="EE290DE7EA0AB45BFB9FFB9CE7F5FBF5" box="[1118,1183,1033,1060]" degrees="41.5" direction="north" orientation="latitude" pageId="15" pageNumber="267" precision="5555" value="41.5">41.5 N</geoCoordinate>
. The transition area with the nominotypical subspecies may be wide.
</paragraph>
</subSubSection>
<subSubSection id="C30738ABEA0AB45BFD03FBE0E029FA7F" pageId="15" pageNumber="267" type="description">
<paragraph id="8BA26B20EA0AB45BFD03FBE0E029FA7F" blockId="15.[676,1188,171,1454]" pageId="15" pageNumber="267">
Nomenclatorial act the electronic on-line early version of this article is considered a published work according to the International Code of Zoological Nomenclature. The new name has also been registered in ZooBank (http://zoobank.org/) where it can be accessed under http://zoobank.org/
<uri id="FF8C6722EA0AB45BFC29FAD9E055FA7F" pageId="15" pageNumber="267">
urn:lsid:zoobank.org:pub:
<uuid id="FFBB51F5EA0AB45BFCC9FAE5E055FA7F" pageId="15" pageNumber="267">6DE32B0B-FB92-4F45-80BB-15 EBC49DD23E</uuid>
</uri>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,295 @@
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<treatment id="03B59F7DFF9C7138F8F06CECFAB7FA52" LSID="urn:lsid:plazi:treatment:03B59F7DFF9C7138F8F06CECFAB7FA52" httpUri="http://treatment.plazi.org/id/03B59F7DFF9C7138F8F06CECFAB7FA52" lastPageNumber="238" pageId="1" pageNumber="238">
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<taxonomicName id="4C1C55E8FF9C7138F8F06CECFD6DFE62" ID-CoL="7427H" authority="(Reissek) Biral (2017: 688)" authorityName="Biral" authorityPageNumber="688" authorityYear="2017" baseAuthorityName="Reissek" box="[136,727,468,495]" class="Magnoliopsida" family="Celastraceae" genus="Monteverdia" kingdom="Plantae" order="Celastrales" pageId="1" pageNumber="238" phylum="Tracheophyta" rank="species" species="communis">
<emphasis id="B968F279FF9C7138F8F06CECFE23FE62" bold="true" box="[136,409,468,495]" italics="true" pageId="1" pageNumber="238">Monteverdia communis</emphasis>
(Reissek)
<bibRefCitation id="EF8D539AFF9C7138FA696CECFD6DFE62" author="Biral, L. &amp; Simmons, M. P. &amp; Smidt, E. &amp; Tembrock, L. R. &amp; Bolson, M. &amp; Archer, R. H. &amp; Lombardi, J. A." box="[529,727,468,494]" pageId="1" pageNumber="238" pagination="680 - 693" refId="ref5712" refString="Biral, L., Simmons, M. P., Smidt, E., Tembrock, L. R. Bolson, M. Archer, R. H. &amp; Lombardi, J. A. (2017) Systematics of the New World Maytenus (Celastraceae) and a new delimitation of the genus. Systematic Botany 42 (4): 680 - 693. https: // doi. org / 10.1600 / 036364417 X 696456" type="journal article" year="2017">Biral (2017: 688)</bibRefCitation>
</taxonomicName>
.
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, t. IV, f. 2, t. VIII).
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s.n.
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[
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).
</paragraph>
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<paragraph id="8BA32E6BFF9C7138F8F06E24FED3FCF5" blockId="1.[136,1452,796,890]" pageId="1" pageNumber="238">
=
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<emphasis id="B968F279FF9C7138F8E26E24FED1FCBD" box="[154,363,796,817]" italics="true" pageId="1" pageNumber="238">Maytenus grandiflora</emphasis>
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, t. III, f. 3).
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: in prope urbem principale, s.d. [1817-1821],
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s.n.
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(
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W [0059534]! image, designated here;
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F [870327, barcode V0361497F]!, W [0059532 &amp; 0059533]! images).
<emphasis id="B968F279FF9C7138F96D6E5CFED3FCF5" box="[277,361,868,889]" italics="true" pageId="1" pageNumber="238">syn. nov.</emphasis>
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<emphasis id="B968F279FF9C7138F8F06E91FE3AFC4E" box="[136,384,936,962]" italics="true" pageId="1" pageNumber="238">Maytenus grandiflora</emphasis>
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was described by
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based on three different collections gathered from the city of
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by Johann Christian Mikan, Heinrich Wilhelm Schott and Guilherme Schüch de Capanema. After analysis of protologue and examination of specimens mentioned, no relevant difference between the circumscription of
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<emphasis id="B968F279FF9C7138F8D0692DFEEBFBA2" box="[168,337,1044,1070]" italics="true" pageId="1" pageNumber="238">M. grandiflora</emphasis>
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and
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<emphasis id="B968F279FF9C7138F9F4692CFD2FFBA2" box="[396,661,1044,1070]" italics="true" pageId="1" pageNumber="238">Monteverdia communis</emphasis>
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could be observed. It is a typical species from the Brazilian Atlantic Forest, characterized by lanceolate to elliptic leaves, drying brown on the abaxial side, and multiflowered cymes with a conspicuous peduncle, always longer than pedicels, and flowers distributed along the axis. These two species were published in the same volume of
<taxonomicName id="4C1C55E8FF9C7138FA3C69B8FC72FB16" authority="(Reissek 1861)" baseAuthorityName="Reissek" baseAuthorityYear="1861" box="[580,968,1152,1178]" class="Magnoliopsida" family="Celastraceae" genus="Flora" kingdom="Plantae" order="Celastrales" pageId="1" pageNumber="238" phylum="Tracheophyta" rank="species" species="brasiliensis">
<emphasis id="B968F279FF9C7138FA3C69B8FCABFB16" box="[580,785,1152,1178]" italics="true" pageId="1" pageNumber="238">Flora Brasiliensis</emphasis>
(
<bibRefCitation id="EF8D539AFF9C7138FB5969B8FC05FB16" author="Reissek, S." box="[801,959,1152,1178]" pageId="1" pageNumber="238" pagination="1 - 34" refId="ref6671" refString="Reissek, S. (1861) Celastrinea. In: Martius, C. F. P. &amp; Eichler, A. W. (Eds.) Flora Brasiliensis, v. 11, pars 1. Frid. Fleischer, Lipsiae [Leipzig], pp. 1 - 34, pl. 1 - 10." type="book chapter" year="1861">Reissek 1861</bibRefCitation>
)
</taxonomicName>
; in this case the name
<taxonomicName id="4C1C55E8FF9C7138FCB569B9FADCFB16" authorityName="Biral" authorityYear="2017" baseAuthorityName="Reissek" box="[1229,1382,1153,1178]" class="Magnoliopsida" family="Celastraceae" genus="Monteverdia" kingdom="Plantae" order="Celastrales" pageId="1" pageNumber="238" phylum="Tracheophyta" rank="species" species="communis">
<emphasis id="B968F279FF9C7138FCB569B9FADCFB16" box="[1229,1382,1153,1178]" italics="true" pageId="1" pageNumber="238">M. communis</emphasis>
</taxonomicName>
is the preferable to be used as its basionym is most commonly found in literature and herbarium identifications (see article 11.5. of the Shenzen Code,
<bibRefCitation id="EF8D539AFF9C7138F9C469F0FD2DFB6E" author="Turland, N. J. &amp; Wiersema, J. H. &amp; Barrie, F. R. &amp; Greuter, W. &amp; Hawksworth, D. L. &amp; Herendeen, P. S. &amp; Knapp, S. &amp; Kusber, W. - H. &amp; Li, D. - Z. &amp; Marhold, K. &amp; May, T. W. &amp; McNeill, J. &amp; Monro, A. M. &amp; Prado, J. &amp; Price, M. J. &amp; Smith, G. F." box="[444,663,1224,1250]" pageId="1" pageNumber="238" refId="ref7074" refString="Turland, N. J., Wiersema, J. H., Barrie, F. R., Greuter, W., Hawksworth, D. L., Herendeen, P. S., Knapp, S., Kusber, W. - H., Li, D. - Z., Marhold, K., May, T. W., McNeill, J., Monro, A. M., Prado, J., Price, M. J. &amp; Smith, G. F. (Eds.) (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzen Code): Adopted by the Nineteenth International Botanical Congress Shenzen, China, July 2017. Regnum Vegetabile 159. Koeltz Scientific Books, Glashutten, 254 pp. https: // doi. org / 10.12705 / Code. 2018" type="book" year="2018">
Turland
<emphasis id="B968F279FF9C7138FA6369F1FDE9FB6E" box="[539,595,1224,1250]" italics="true" pageId="1" pageNumber="238">et al.</emphasis>
2018
</bibRefCitation>
).
</paragraph>
<paragraph id="8BA32E6BFF9C7138F8C569D4FAB7FA52" blockId="1.[136,1452,936,1503]" pageId="1" pageNumber="238">
The specimen W0059534 is selected as
<typeStatus id="54A790C9FF9C7138FAE869D4FD4DFA8A" box="[656,759,1260,1286]" pageId="1" pageNumber="238" type="lectotype">lectotype</typeStatus>
for
<emphasis id="B968F279FF9C7138FB5469D5FB93FA8A" box="[812,1065,1260,1286]" italics="true" pageId="1" pageNumber="238">
<taxonomicName id="4C1C55E8FF9C7138FB5469D5FB9FFA8A" box="[812,1061,1260,1286]" class="Magnoliopsida" family="Celastraceae" genus="Maytenus" kingdom="Plantae" order="Celastrales" pageId="1" pageNumber="238" phylum="Tracheophyta" rank="species" species="grandifolia">Maytenus grandifolia</taxonomicName>
.
</emphasis>
This specimen is the unique that bears a label that includes a handwritten description by Reissek, very similar to the one he provided in the protologue, which constitutes clear evidence by the author in applying this name (Recommendation 9a.3 from the Shenzen Code,
<bibRefCitation id="EF8D539AFF9C7138F8F06860FEDDFAFE" author="Turland, N. J. &amp; Wiersema, J. H. &amp; Barrie, F. R. &amp; Greuter, W. &amp; Hawksworth, D. L. &amp; Herendeen, P. S. &amp; Knapp, S. &amp; Kusber, W. - H. &amp; Li, D. - Z. &amp; Marhold, K. &amp; May, T. W. &amp; McNeill, J. &amp; Monro, A. M. &amp; Prado, J. &amp; Price, M. J. &amp; Smith, G. F." box="[136,359,1368,1394]" pageId="1" pageNumber="238" refId="ref7074" refString="Turland, N. J., Wiersema, J. H., Barrie, F. R., Greuter, W., Hawksworth, D. L., Herendeen, P. S., Knapp, S., Kusber, W. - H., Li, D. - Z., Marhold, K., May, T. W., McNeill, J., Monro, A. M., Prado, J., Price, M. J. &amp; Smith, G. F. (Eds.) (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzen Code): Adopted by the Nineteenth International Botanical Congress Shenzen, China, July 2017. Regnum Vegetabile 159. Koeltz Scientific Books, Glashutten, 254 pp. https: // doi. org / 10.12705 / Code. 2018" type="book" year="2018">
Turland
<emphasis id="B968F279FF9C7138F8906861FE98FAFE" box="[232,290,1368,1394]" italics="true" pageId="1" pageNumber="238">et al.</emphasis>
2018
</bibRefCitation>
). No date is provided on the label; however, according to
<bibRefCitation id="EF8D539AFF9C7138FB866860FB20FAFE" author="Urban, I." box="[1022,1178,1368,1394]" pageId="1" pageNumber="238" pagination="1 - 268" refId="ref7239" refString="Urban, I. (1906) Vitae itineraque collectorum botanicorum, notae collaboratum biographicae, Flora Brasiliensis ratio edendi chronologica, systema, index familiarum. In: Martius, C. F. P., Eichler, A. W. &amp; Urban, I. (Eds.) Flora Brasiliensis, v. 1, pars 1. R. Oldenbourg, Munchen et Leipzig, pp. 1 - 268." type="book chapter" year="1906">Urban (1906)</bibRefCitation>
, Schott collected in and near the city of
<collectingRegion id="49D8E089FF9C7138F94E6844FE6CFA1A" box="[310,470,1404,1430]" country="Brazil" name="Rio de Janeiro" pageId="1" pageNumber="238">Rio de Janeiro</collectingRegion>
between 1817 and 1820. The specimen represented by negative number 19567 from the Field Museums Type Photograph Collection (previously deposited in B, and presumably destroyed) is not type of
<taxonomicName id="4C1C55E8FF9C7138FDF56899FEBCFA52" class="Magnoliopsida" family="Celastraceae" genus="Maytenus" kingdom="Plantae" order="Celastrales" pageId="1" pageNumber="238" phylum="Tracheophyta" rank="species" species="grandifolia">
<emphasis id="B968F279FF9C7138FDF56899FEBCFA52" italics="true" pageId="1" pageNumber="238">M. grandifolia</emphasis>
</taxonomicName>
because it is of a specimen gathered by Riedel and, thus, is not part of the original material.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,202 @@
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<emphasis id="B968F279FF9C7138F8F06B34FE25F9AB" bold="true" box="[136,415,1548,1575]" italics="true" pageId="1" pageNumber="238">Monteverdia floribunda</emphasis>
(Reissek)
<bibRefCitation id="EF8D539AFF9C7138FA636B34FD5CF9AA" author="Biral, L. &amp; Simmons, M. P. &amp; Smidt, E. &amp; Tembrock, L. R. &amp; Bolson, M. &amp; Archer, R. H. &amp; Lombardi, J. A." box="[539,742,1548,1574]" pageId="1" pageNumber="238" pagination="680 - 693" refId="ref5712" refString="Biral, L., Simmons, M. P., Smidt, E., Tembrock, L. R. Bolson, M. Archer, R. H. &amp; Lombardi, J. A. (2017) Systematics of the New World Maytenus (Celastraceae) and a new delimitation of the genus. Systematic Botany 42 (4): 680 - 693. https: // doi. org / 10.1600 / 036364417 X 696456" type="journal article" year="2017">Biral (2017: 688)</bibRefCitation>
</taxonomicName>
.
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<taxonomicName id="4C1C55E8FF9C7138FB746B35FB69F9AA" authority="Reissek (1861: 16" authorityName="Reissek" authorityYear="1861" box="[780,1235,1548,1574]" class="Magnoliopsida" family="Celastraceae" genus="Maytenus" kingdom="Plantae" order="Celastrales" pageId="1" pageNumber="238" phylum="Tracheophyta" rank="species" species="floribunda">
<emphasis id="B968F279FF9C7138FB746B35FC42F9AA" box="[780,1016,1548,1574]" italics="true" pageId="1" pageNumber="238">Maytenus floribunda</emphasis>
<bibRefCitation id="EF8D539AFF9C7138FC7A6B34FB69F9AA" author="Reissek, S." box="[1026,1235,1548,1574]" pageId="1" pageNumber="238" pagination="1 - 34" refId="ref6671" refString="Reissek, S. (1861) Celastrinea. In: Martius, C. F. P. &amp; Eichler, A. W. (Eds.) Flora Brasiliensis, v. 11, pars 1. Frid. Fleischer, Lipsiae [Leipzig], pp. 1 - 34, pl. 1 - 10." type="book chapter" year="1861">Reissek (1861: 16</bibRefCitation>
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, t. IV, f. 3, t. VII).
</paragraph>
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<collectingMunicipality id="6BC7B411FF9C7138F8F06B08FF79F9C6" box="[136,195,1584,1610]" pageId="1" pageNumber="238">Type</collectingMunicipality>
:—
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<collectingCountry id="F30B6EFBFF9C7138F89C6B08FEF7F9C6" box="[228,333,1584,1610]" name="Brazil" pageId="1" pageNumber="238">BRAZIL</collectingCountry>
.
<collectingRegion id="49D8E089FF9C7138F9206B08FE24F9C6" box="[344,414,1584,1610]" country="Brazil" name="Goias" pageId="1" pageNumber="238">Goiás</collectingRegion>
:
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,
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<collectingDate id="EFE6F143FF9C7138FA5E6B09FD27F9C6" box="[550,669,1584,1611]" pageId="1" pageNumber="238" value="1819-05">May 1819</collectingDate>
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,
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<collectorName id="26E94BBDFF9C7138FAD26B08FD65F9C6" box="[682,735,1584,1610]" pageId="1" pageNumber="238">Pohl</collectorName>
s.n.
</emphasis>
(
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,
<collectionCode id="ED0DB6AEFF9C7138FBEE6B09FC0AF9C7" box="[918,944,1585,1611]" pageId="1" pageNumber="238">W</collectionCode>
[
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] [U neg 1177]! image, designated here
</materialsCitation>
;
<materialsCitation id="3B742436FF9C7138F8B06B6CFDD6F9E3" box="[200,620,1620,1647]" collectingDate="1819-05" collectionCode="BR" collectorName="Pohl" country="Brazil" location="ad Crixas" pageId="1" pageNumber="238" specimenCode="0000005737673" specimenCount="1" stateProvince="Goias" typeStatus="isolectotype">
<typeStatus id="54A790C9FF9C7138F8B06B6CFEE7F9E2" box="[200,349,1620,1646]" pageId="1" pageNumber="238" type="isolectotype">isolectotypes</typeStatus>
,
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[
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]!
</materialsCitation>
,
<materialsCitation id="3B742436FF9C7138FA0F6B6DFE34F91E" collectingDate="1819-05" collectionCode="LE" collectorName="Pohl" country="Brazil" location="ad Crixas" pageId="1" pageNumber="238" specimenCount="1" stateProvince="Goias" typeStatus="isolectotype">
<collectionCode id="ED0DB6AEFF9C7138FA0F6B6DFD23F9E3" box="[631,665,1621,1647]" pageId="1" pageNumber="238">LE</collectionCode>
[n.v., indicated by
<bibRefCitation id="EF8D539AFF9C7138FB0C6B6CFB21F9E2" author="Carvalho-Okano, R. M." box="[884,1179,1620,1646]" pageId="1" pageNumber="238" refId="ref5826" refString="Carvalho-Okano, R. M. (1992) Estudos taxonomicos do genero Maytenus (Celastraceae) no Brasil extra-amazonico. Thesis, Universidade Estadual de Campinas, Campinas, x + 253 pp." type="book" year="1992">Carvalho-Okano 1992: 62</bibRefCitation>
and
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],
</materialsCitation>
<materialsCitation id="3B742436FF9C7138F9ED6B41FD3EF91E" box="[405,644,1656,1683]" collectingDate="1819-05" collectionCode="W" collectorName="Pohl" country="Brazil" location="ad Crixas" pageId="1" pageNumber="238" specimenCode="0047063" specimenCount="1" stateProvince="Goias" typeStatus="isolectotype">
<collectionCode id="ED0DB6AEFF9C7138F9ED6B41FE15F91F" box="[405,431,1657,1683]" pageId="1" pageNumber="238">W</collectionCode>
[
<specimenCode id="DBBA8610FF9C7138F9C46B40FD93F91E" box="[444,553,1656,1682]" pageId="1" pageNumber="238">0047063</specimenCode>
]! image
</materialsCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3067DE0FF9C7138F8F06BFCFA96F971" pageId="1" pageNumber="238" type="reference_group">
<paragraph id="8BA32E6BFF9C7138F8F06BFCFA96F971" blockId="1.[136,1451,1732,1790]" pageId="1" pageNumber="238">
=
<taxonomicName id="4C1C55E8FF9C7138F8E46BFCFEDEF955" authorityName="Reissek" authorityYear="1861" box="[156,356,1732,1753]" class="Magnoliopsida" family="Celastraceae" genus="Maytenus" kingdom="Plantae" order="Celastrales" pageId="1" pageNumber="238" phylum="Tracheophyta" rank="species" species="floribunda">
<emphasis id="B968F279FF9C7138F8E46BFCFEDEF955" box="[156,356,1732,1753]" italics="true" pageId="1" pageNumber="238">Maytenus floribunda</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF8D539AFF9C7138F9096BFCFDA0F956" author="Reissek, S." box="[369,538,1732,1754]" pageId="1" pageNumber="238" pagination="1 - 34" refId="ref6671" refString="Reissek, S. (1861) Celastrinea. In: Martius, C. F. P. &amp; Eichler, A. W. (Eds.) Flora Brasiliensis, v. 11, pars 1. Frid. Fleischer, Lipsiae [Leipzig], pp. 1 - 34, pl. 1 - 10." type="book chapter" year="1861">Reissek 1861: 16</bibRefCitation>
, t. IV, f. 3, t. VII)
<taxonomicName id="4C1C55E8FF9C7138FAB06BFCFB92F956" authority="Warming (1880: 366)" authorityName="Warming" authorityPageNumber="366" authorityYear="1880" box="[712,1064,1732,1754]" class="Magnoliopsida" family="Celastraceae" genus="Maytenus" kingdom="Plantae" order="Celastrales" pageId="1" pageNumber="238" phylum="Tracheophyta" rank="variety" species="floribunda" variety="parvifolia">
var.
<emphasis id="B968F279FF9C7138FA896BFCFCEAF955" box="[753,848,1732,1753]" italics="true" pageId="1" pageNumber="238">parvifolia</emphasis>
<bibRefCitation id="EF8D539AFF9C7138FB2D6BFCFB92F956" author="Warming, E." box="[853,1064,1732,1754]" pageId="1" pageNumber="238" pagination="363 - 366" refId="ref7316" refString="Warming, E. (1880) Symbolae ad floram Brasiliae centralis cognoscendam. Particula XXVI. Fam. Celastraceae. Videnskabelige Meddelelser fraden naturhistoriske Forening i Kjoebenhavn 412: 363 - 366." type="journal article" year="1880">Warming (1880: 366)</bibRefCitation>
</taxonomicName>
. Type:—
<collectingCountry id="F30B6EFBFF9C7138FC076BFCFB63F955" box="[1151,1241,1732,1753]" name="Brazil" pageId="1" pageNumber="238">BRAZIL</collectingCountry>
.
<collectingRegion id="49D8E089FF9C7138FC9A6BFCFADCF956" box="[1250,1382,1732,1754]" country="Brazil" name="Minas Gerais" pageId="1" pageNumber="238">Minas Gerais</collectingRegion>
: Lagoa Santa [Lapa Vermelha], in silvis, s.d. [
<date id="FFA208ABFF9C7138FA536BD0FD53F972" box="[555,745,1768,1790]" pageId="1" pageNumber="238" value="1864-11-11">11 November 1864</date>
],
<emphasis id="B968F279FF9C7138FA816BD0FCC2F971" box="[761,888,1768,1789]" italics="true" pageId="1" pageNumber="238">Warming s.n.</emphasis>
(
<typeStatus id="54A790C9FF9C7138FBFC6BD0FC66F972" box="[900,988,1768,1790]" pageId="1" pageNumber="238" type="holotype">holotype</typeStatus>
, C [C10023017]! image).
<emphasis id="B968F279FF9C7138FCA06BD0FA96F971" box="[1240,1324,1768,1789]" italics="true" pageId="1" pageNumber="238">syn. nov.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3067DE0FF9C713BF8F06A15FEB6FEE0" lastPageId="2" lastPageNumber="239" pageId="1" pageNumber="238" type="discussion">
<paragraph id="8BA32E6BFF9C713BF8F06A15FDA3FE8C" blockId="1.[136,1452,1836,2079]" lastBlockId="2.[136,1452,158,364]" lastPageId="2" lastPageNumber="239" pageId="1" pageNumber="238">
<taxonomicName id="4C1C55E8FF9C7138F8F06A15FDABF8CA" authorityName="Warming" authorityYear="1880" box="[136,529,1836,1863]" class="Magnoliopsida" family="Celastraceae" genus="Maytenus" kingdom="Plantae" order="Celastrales" pageId="1" pageNumber="238" phylum="Tracheophyta" rank="variety" species="floribunda" variety="parvifolia">
<emphasis id="B968F279FF9C7138F8F06A15FECAF8CA" box="[136,368,1836,1862]" italics="true" pageId="1" pageNumber="238">Maytenus floribunda</emphasis>
var.
<emphasis id="B968F279FF9C7138F9DB6A14FDABF8CA" box="[419,529,1836,1862]" italics="true" pageId="1" pageNumber="238">parvifolia</emphasis>
</taxonomicName>
was described by Eugenius Warming based on one of his collections from the Minas Gerais state in Brazils Southeastern region, where he lived from 1863 to 1866 (
<bibRefCitation id="EF8D539AFF9C7138FB8D6A68FBC2F8E6" author="Klein, A. L." box="[1013,1144,1872,1898]" pageId="1" pageNumber="238" refId="ref6105" refString="Klein, A. L. (org.) (2002) Eugen Warming e o cerrado brasileiro: um seculo depois. Editora Unesp, Imprensa Oficial do Estado, Sao Paulo, 156 pp." type="book" year="2002">Klein 2002</bibRefCitation>
).
<taxonomicName id="4C1C55E8FF9C7138FCF16A68FA29F8E6" authorityName="Biral" authorityYear="2017" baseAuthorityName="Reissek" box="[1161,1427,1872,1898]" class="Magnoliopsida" family="Celastraceae" genus="Monteverdia" kingdom="Plantae" order="Celastrales" pageId="1" pageNumber="238" phylum="Tracheophyta" rank="species" species="floribunda">
<emphasis id="B968F279FF9C7138FCF16A68FA29F8E6" box="[1161,1427,1872,1898]" italics="true" pageId="1" pageNumber="238">Monteverdia floribunda</emphasis>
</taxonomicName>
is a widely distributed species in South America. It is most prevalent in the Brazilian areas containing Cerrado vegetation but has also been collected in
<collectingCountry id="F30B6EFBFF9C7138F9AF6AA0FD96F83E" box="[471,556,1944,1970]" name="Bolivia" pageId="1" pageNumber="238">Bolivia</collectingCountry>
,
<collectingCountry id="F30B6EFBFF9C7138FA406AA0FD1CF83E" box="[568,678,1944,1970]" name="Colombia" pageId="1" pageNumber="238">Colombia</collectingCountry>
and Guyanas (
<bibRefCitation id="EF8D539AFF9C7138FB356AA0FBB0F83F" author="Biral, L. &amp; Smidt, E. C. &amp; Bolson, M. &amp; Lombardi, J. A." box="[845,1034,1944,1971]" pageId="1" pageNumber="238" pagination="53 - 62" refId="ref5596" refString="Biral, L., Smidt, E. C., Bolson, M. &amp; Lombardi, J. A. (2015) A new species of Maytenus (Celastraceae) from the Brazilian Atlantic Forest, with evidence of molecular phylogeny, and two new synonyms for Maytenus floridunda. Phytotaxa 231 (1): 53 - 62. https: // doi. org / 10.11646 / phytotaxa. 231.1.5" type="journal article" year="2015">
Biral
<emphasis id="B968F279FF9C7138FBF66AA1FC7CF83E" box="[910,966,1944,1970]" italics="true" pageId="1" pageNumber="238">et al.</emphasis>
2015
</bibRefCitation>
).
<taxonomicName id="4C1C55E8FF9C7138FC646AA1FA11F83E" authorityName="Warming" authorityYear="1880" box="[1052,1451,1944,1971]" class="Magnoliopsida" family="Celastraceae" genus="Maytenus" kingdom="Plantae" order="Celastrales" pageId="1" pageNumber="238" phylum="Tracheophyta" rank="variety" species="floribunda" variety="parvifolia">
<emphasis id="B968F279FF9C7138FC646AA1FABCF83E" box="[1052,1286,1944,1970]" italics="true" pageId="1" pageNumber="238">Maytenus floribunda</emphasis>
var.
<emphasis id="B968F279FF9C7138FD456AA0FA11F83E" box="[1341,1451,1944,1970]" italics="true" pageId="1" pageNumber="238">parvifolia</emphasis>
</taxonomicName>
was ignored in
<bibRefCitation id="EF8D539AFF9C7138F9416A84FDE3F85A" author="Carvalho-Okano, R. M." box="[313,601,1980,2006]" pageId="1" pageNumber="238" refId="ref5826" refString="Carvalho-Okano, R. M. (1992) Estudos taxonomicos do genero Maytenus (Celastraceae) no Brasil extra-amazonico. Thesis, Universidade Estadual de Campinas, Campinas, x + 253 pp." type="book" year="1992">Carvalho-Okanos (1992)</bibRefCitation>
<taxonomicName id="4C1C55E8FF9C7138FA1A6A85FD75F85A" box="[610,719,1981,2006]" class="Magnoliopsida" family="Celastraceae" genus="Maytenus" kingdom="Plantae" order="Celastrales" pageId="1" pageNumber="238" phylum="Tracheophyta" rank="genus">
<emphasis id="B968F279FF9C7138FA1A6A85FD75F85A" box="[610,719,1981,2006]" italics="true" pageId="1" pageNumber="238">Maytenus</emphasis>
</taxonomicName>
revision and it did not appear in other references I traced. After contacting the herbarium C, I was able to inspect a high-resolution image of the
<typeStatus id="54A790C9FF9C7138FC516AD8FB31F876" box="[1065,1163,2016,2042]" pageId="1" pageNumber="238" type="holotype">holotype</typeStatus>
and to determine that its continued designation as a variety of
<taxonomicName id="4C1C55E8FF9C7138FA4B653CFCFAF792" authorityName="Biral" authorityYear="2017" baseAuthorityName="Reissek" box="[563,832,2052,2078]" class="Magnoliopsida" family="Celastraceae" genus="Monteverdia" kingdom="Plantae" order="Celastrales" pageId="1" pageNumber="238" phylum="Tracheophyta" rank="species" species="floribunda">
<emphasis id="B968F279FF9C7138FA4B653CFCFAF792" box="[563,832,2052,2078]" italics="true" pageId="1" pageNumber="238">Monteverdia floribunda</emphasis>
</taxonomicName>
, based solely on leaf characters, is not supported. The leaves of
<taxonomicName id="4C1C55E8FF9F713BF88B6DA7FE35FF34" authorityName="Biral" authorityYear="2017" baseAuthorityName="Reissek" box="[243,399,158,184]" class="Magnoliopsida" family="Celastraceae" genus="Monteverdia" kingdom="Plantae" order="Celastrales" pageId="2" pageNumber="239" phylum="Tracheophyta" rank="species" species="floribunda">
<emphasis id="B968F279FF9F713BF88B6DA7FE35FF34" box="[243,399,158,184]" italics="true" pageId="2" pageNumber="239">M. floribunda</emphasis>
</taxonomicName>
are quite variable in size but relatively constant in other features, such as having an elliptic to obovate shape, a margin that is crenate only close to the apex, 11-14 pairs of ascending secondary veins, and multi-flowered fasciculate inflorescences.
</paragraph>
<paragraph id="8BA32E6BFF9F713BF8C56C32FEB6FEE0" blockId="2.[136,1452,158,364]" pageId="2" pageNumber="239">
The specimen W0047062 is elected as
<typeStatus id="54A790C9FF9F713BFA1F6C32FD74FEA8" box="[615,718,266,292]" pageId="2" pageNumber="239" type="lectotype">lectotype</typeStatus>
for
<taxonomicName id="4C1C55E8FF9F713BFA816C32FBB8FEA8" authorityName="Biral" authorityYear="2017" baseAuthorityName="Reissek" box="[761,1026,266,292]" class="Magnoliopsida" family="Celastraceae" genus="Monteverdia" kingdom="Plantae" order="Celastrales" pageId="2" pageNumber="239" phylum="Tracheophyta" rank="species" species="floribunda">
<emphasis id="B968F279FF9F713BFA816C32FBB8FEA8" box="[761,1026,266,292]" italics="true" pageId="2" pageNumber="239">Monteverdia floribunda</emphasis>
</taxonomicName>
because it is constituted by a complete branch with flowers and contains two labels, one with the name authors identification and other bearing a handwritten description.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,225 @@
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<taxonomicName id="4C1C55E8FF9E713AF8F06C16FD28FEC4" ID-CoL="6RQ9B" authority="(Reissek) Biral (2017: 689)" authorityName="Biral" authorityPageNumber="689" authorityYear="2017" baseAuthorityName="Reissek" box="[136,658,302,329]" class="Magnoliopsida" family="Celastraceae" genus="Monteverdia" kingdom="Plantae" order="Celastrales" pageId="3" pageNumber="240" phylum="Tracheophyta" rank="species" species="laevis">
<emphasis id="B968F279FF9E713AF8F06C16FEE5FEC5" bold="true" box="[136,351,302,329]" italics="true" pageId="3" pageNumber="240">Monteverdia laevis</emphasis>
(Reissek)
<bibRefCitation id="EF8D539AFF9E713AF9AA6C16FD28FEC4" author="Biral, L. &amp; Simmons, M. P. &amp; Smidt, E. &amp; Tembrock, L. R. &amp; Bolson, M. &amp; Archer, R. H. &amp; Lombardi, J. A." box="[466,658,302,328]" pageId="3" pageNumber="240" pagination="680 - 693" refId="ref5712" refString="Biral, L., Simmons, M. P., Smidt, E., Tembrock, L. R. Bolson, M. Archer, R. H. &amp; Lombardi, J. A. (2017) Systematics of the New World Maytenus (Celastraceae) and a new delimitation of the genus. Systematic Botany 42 (4): 680 - 693. https: // doi. org / 10.1600 / 036364417 X 696456" type="journal article" year="2017">Biral (2017:689)</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA32E6BFF9E713AFAE26C17FB25FEC4" blockId="3.[136,1452,302,508]" box="[666,1183,302,329]" pageId="3" pageNumber="240">
<taxonomicName id="4C1C55E8FF9E713AFAD66C17FB91FEC4" authority="Reissek (1861: 27" authorityName="Reissek" authorityYear="1861" box="[686,1067,302,328]" class="Magnoliopsida" family="Celastraceae" genus="Maytenus" kingdom="Plantae" order="Celastrales" pageId="3" pageNumber="240" phylum="Tracheophyta" rank="species" species="laevis">
<emphasis id="B968F279FF9E713AFAD66C17FCDAFEC4" box="[686,864,302,328]" italics="true" pageId="3" pageNumber="240">Maytenus laevis</emphasis>
<bibRefCitation id="EF8D539AFF9E713AFB1C6C16FB91FEC4" author="Reissek, S." box="[868,1067,302,328]" pageId="3" pageNumber="240" pagination="1 - 34" refId="ref6671" refString="Reissek, S. (1861) Celastrinea. In: Martius, C. F. P. &amp; Eichler, A. W. (Eds.) Flora Brasiliensis, v. 11, pars 1. Frid. Fleischer, Lipsiae [Leipzig], pp. 1 - 34, pl. 1 - 10." type="book chapter" year="1861">Reissek (1861: 27</bibRefCitation>
</taxonomicName>
, t. IV, f. 6).
</paragraph>
<paragraph id="8BA32E6BFF9E713AFCDB6C16FF52FE70" blockId="3.[136,1452,302,508]" pageId="3" pageNumber="240">
<collectingCounty id="62C256E7FF9E713AFCDB6C16FB67FEC4" box="[1187,1245,302,328]" pageId="3" pageNumber="240">Type</collectingCounty>
:—
<materialsCitation id="3B742436FF9E713AFC856C16FC31FE1C" pageId="3" pageNumber="240">
<collectingCountry id="F30B6EFBFF9E713AFC856C16FA13FEC4" box="[1277,1449,302,328]" name="Venezuela" pageId="3" pageNumber="240">VENEZUELA</collectingCountry>
.
<collectingRegion id="49D8E089FF9E713AF8F06C6AFEB9FEE0" box="[136,259,338,364]" country="Venezuela" name="Amazonas" pageId="3" pageNumber="240">Amazonas</collectingRegion>
: habitat ad flum. [Guainia v.]
<collectingMunicipality id="6BC7B411FF9E713AFAFA6C6AFCBAFEE1" box="[642,768,338,365]" pageId="3" pageNumber="240">Rio Negro</collectingMunicipality>
, supra ostium flum.
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,
<date id="FFA208ABFF9E713AFCF76C6BFAB3FEE1" box="[1167,1289,338,365]" pageId="3" pageNumber="240" value="1854-06">
<collectingDate id="EFE6F143FF9E713AFCF76C6BFAB3FEE1" box="[1167,1289,338,365]" pageId="3" pageNumber="240" value="1854-06">June 1854</collectingDate>
</date>
,
<emphasis id="B968F279FF9E713AFD616C6BFA11FEE0" box="[1305,1451,338,365]" italics="true" pageId="3" pageNumber="240">Spruce 2284</emphasis>
(
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, BR [BR0000005737055]!, designated here;
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</materialsCitation>
,
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,
<materialsCitation id="3B742436FF9E713AFC906C4FFF75FE38" pageId="3" pageNumber="240">E [E000775445]! image</materialsCitation>
,
<materialsCitation id="3B742436FF9E713AF8A36CA3FD0EFE38" box="[219,692,410,437]" pageId="3" pageNumber="240">F [871648, barcode VF0053757F]! image</materialsCitation>
,
<materialsCitation id="3B742436FF9E713AFAB86CA2FD65FE38" box="[704,735,410,436]" pageId="3" pageNumber="240">G!</materialsCitation>
,
<materialsCitation id="3B742436FF9E713AFA946CA2FBA5FE38" box="[748,1055,410,437]" pageId="3" pageNumber="240">GH [GH00049865]! image</materialsCitation>
,
<materialsCitation id="3B742436FF9E713AFC526CA3FF60FE54" pageId="3" pageNumber="240">K [K000494623 &amp; K000494624]! images</materialsCitation>
,
<materialsCitation id="3B742436FF9E713AF89F6C86FD19FE54" box="[231,675,446,472]" pageId="3" pageNumber="240">MO [1624264, barcode MO-2196878]</materialsCitation>
!,
<materialsCitation id="3B742436FF9E713AFACA6C87FC3CFE54" box="[690,902,446,473]" pageId="3" pageNumber="240">NY [NY00380604</materialsCitation>
,
<materialsCitation id="3B742436FF9E713AFBEB6C86FBFCFE55" box="[915,1094,446,473]" pageId="3" pageNumber="240">MO neg 5035]!</materialsCitation>
,
<materialsCitation id="3B742436FF9E713AFC2B6C87FF66FE70" pageId="3" pageNumber="240">P [P02274006 &amp; P05585758]! images</materialsCitation>
).
</paragraph>
<paragraph id="8BA32E6BFF9E713AF8F06F17FCF5FDE4" blockId="3.[136,1452,558,616]" pageId="3" pageNumber="240">
<emphasis id="B968F279FF9E713AF8F06F17FF22FDC8" box="[136,152,559,580]" italics="true" pageId="3" pageNumber="240">=</emphasis>
<taxonomicName id="4C1C55E8FF9E713AF8E66F16FD1EFDC8" authority="(Reissek) Biral (2017: 690)" authorityName="Biral" authorityPageNumber="690" authorityYear="2017" baseAuthorityName="Reissek" box="[158,676,558,580]" class="Magnoliopsida" family="Celastraceae" genus="Monteverdia" kingdom="Plantae" order="Celastrales" pageId="3" pageNumber="240" phylum="Tracheophyta" rank="species" species="sapotiformis">
<emphasis id="B968F279FF9E713AF8E66F16FE2FFDCF" box="[158,405,558,579]" italics="true" pageId="3" pageNumber="240">Monteverdia sapotiformis</emphasis>
<emphasis id="B968F279FF9E713AF9E36F16FE19FDC8" bold="true" box="[411,419,558,580]" pageId="3" pageNumber="240">(</emphasis>
Reissek)
<bibRefCitation id="EF8D539AFF9E713AF9836F16FD1EFDC8" author="Biral, L. &amp; Simmons, M. P. &amp; Smidt, E. &amp; Tembrock, L. R. &amp; Bolson, M. &amp; Archer, R. H. &amp; Lombardi, J. A." box="[507,676,558,580]" pageId="3" pageNumber="240" pagination="680 - 693" refId="ref5712" refString="Biral, L., Simmons, M. P., Smidt, E., Tembrock, L. R. Bolson, M. Archer, R. H. &amp; Lombardi, J. A. (2017) Systematics of the New World Maytenus (Celastraceae) and a new delimitation of the genus. Systematic Botany 42 (4): 680 - 693. https: // doi. org / 10.1600 / 036364417 X 696456" type="journal article" year="2017">Biral (2017: 690)</bibRefCitation>
</taxonomicName>
. ≡
<taxonomicName id="4C1C55E8FF9E713AFAB86F17FBEAFDCF" authority="Reissek (1861: 27" authorityName="Reissek" authorityYear="1861" box="[704,1104,558,580]" class="Magnoliopsida" family="Celastraceae" genus="Maytenus" kingdom="Plantae" order="Celastrales" pageId="3" pageNumber="240" phylum="Tracheophyta" rank="species" species="sapotiformis">
<emphasis id="B968F279FF9E713AFAB86F17FC21FDCF" box="[704,923,558,580]" italics="true" pageId="3" pageNumber="240">Maytenus sapotiformis</emphasis>
<bibRefCitation id="EF8D539AFF9E713AFBD96F16FBEAFDCF" author="Reissek, S." box="[929,1104,558,580]" pageId="3" pageNumber="240" pagination="1 - 34" refId="ref6671" refString="Reissek, S. (1861) Celastrinea. In: Martius, C. F. P. &amp; Eichler, A. W. (Eds.) Flora Brasiliensis, v. 11, pars 1. Frid. Fleischer, Lipsiae [Leipzig], pp. 1 - 34, pl. 1 - 10." type="book chapter" year="1861">Reissek (1861: 27</bibRefCitation>
</taxonomicName>
, t. IV, f. 5). Type:—
<collectingCountry id="F30B6EFBFF9E713AFD766F16FAC2FDCF" box="[1294,1400,558,579]" name="Guyana" pageId="3" pageNumber="240">GUYANA</collectingCountry>
. S.d.,
<emphasis id="B968F279FF9E713AF8C56F6AFEBFFDEB" box="[189,261,594,615]" italics="true" pageId="3" pageNumber="240">Lindley</emphasis>
[
<emphasis id="B968F279FF9E713AF96B6F6AFEECFDEB" box="[275,342,594,615]" italics="true" pageId="3" pageNumber="240">Martin</emphasis>
]
<emphasis id="B968F279FF9E713AF91C6F6BFE3CFDE4" box="[356,390,595,616]" italics="true" pageId="3" pageNumber="240">s.n.</emphasis>
(
<typeStatus id="54A790C9FF9E713AF9EA6F6AFE52FDE4" box="[402,488,594,616]" pageId="3" pageNumber="240" type="holotype">holotype</typeStatus>
BR [BR0000005798797]!).
<emphasis id="B968F279FF9E713AFA836F6BFCF5FDE4" box="[763,847,595,616]" italics="true" pageId="3" pageNumber="240">syn. nov.</emphasis>
</paragraph>
<paragraph id="8BA32E6BFF9E713AF8F06FAEFC92FC5C" blockId="3.[136,1452,662,1192]" pageId="3" pageNumber="240">
Revisiting the
<taxonomicName id="4C1C55E8FF9E713AF95E6FAEFE09FD3C" box="[294,435,662,688]" class="Magnoliopsida" family="Celastraceae" kingdom="Plantae" order="Celastrales" pageId="3" pageNumber="240" phylum="Tracheophyta" rank="family">Celastraceae</taxonomicName>
names published by
<bibRefCitation id="EF8D539AFF9E713AFAEE6FAEFC86FD3C" author="Reissek, S." box="[662,828,662,688]" pageId="3" pageNumber="240" pagination="1 - 34" refId="ref6671" refString="Reissek, S. (1861) Celastrinea. In: Martius, C. F. P. &amp; Eichler, A. W. (Eds.) Flora Brasiliensis, v. 11, pars 1. Frid. Fleischer, Lipsiae [Leipzig], pp. 1 - 34, pl. 1 - 10." type="book chapter" year="1861">Reissek (1861)</bibRefCitation>
in Martius
<taxonomicName id="4C1C55E8FF9E713AFBC76FAEFB3DFD3C" baseAuthorityName="Reissek" baseAuthorityYear="1861" box="[959,1159,662,688]" class="Magnoliopsida" family="Celastraceae" genus="Flora" kingdom="Plantae" order="Celastrales" pageId="3" pageNumber="240" phylum="Tracheophyta" rank="species" species="brasiliensis">
<emphasis id="B968F279FF9E713AFBC76FAEFB3DFD3C" box="[959,1159,662,688]" italics="true" pageId="3" pageNumber="240">Flora Brasiliensis</emphasis>
</taxonomicName>
, no substantial differences could be found between
<typeStatus id="54A790C9FF9E713AF9DC6F83FE65FD59" box="[420,479,699,725]" pageId="3" pageNumber="240">types</typeStatus>
and protologues of
<taxonomicName id="4C1C55E8FF9E713AFAB06F83FCC5FD58" authorityName="Reissek" authorityYear="1861" box="[712,895,698,724]" class="Magnoliopsida" family="Celastraceae" genus="Maytenus" kingdom="Plantae" order="Celastrales" pageId="3" pageNumber="240" phylum="Tracheophyta" rank="species" species="laevis">
<emphasis id="B968F279FF9E713AFAB06F83FCC5FD58" box="[712,895,698,724]" italics="true" pageId="3" pageNumber="240">Maytenus laevis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1C55E8FF9E713AFBC36F83FBD4FD58" authorityName="Reissek" authorityYear="1861" box="[955,1134,698,724]" class="Magnoliopsida" family="Celastraceae" genus="Maytenus" kingdom="Plantae" order="Celastrales" pageId="3" pageNumber="240" phylum="Tracheophyta" rank="species" species="sapotiformis">
<emphasis id="B968F279FF9E713AFBC36F83FBD4FD58" box="[955,1134,698,724]" italics="true" pageId="3" pageNumber="240">M. sapotiformis</emphasis>
</taxonomicName>
. They are indistinguishable morphologically, being characterized by carenated young twigs, coriaceous leaves with obscure venation on both sides, pruinose, usually blackening as they dry, and fasciculate inflorescences. Both names were described on the same page of the
<taxonomicName id="4C1C55E8FF9E713AF9336E1EFE62FCCC" box="[331,472,806,832]" class="Magnoliopsida" family="Celastraceae" kingdom="Plantae" order="Celastrales" pageId="3" pageNumber="240" phylum="Tracheophyta" rank="family">Celastraceae</taxonomicName>
treatment in
<taxonomicName id="4C1C55E8FF9E713AFA146E1EFB95FCCC" authority="(Reissek 1861, p. 27)" baseAuthorityName="Reissek" baseAuthorityPageNumber="27" baseAuthorityYear="1861" box="[620,1071,806,833]" class="Magnoliopsida" family="Celastraceae" genus="Flora" kingdom="Plantae" order="Celastrales" pageId="3" pageNumber="240" phylum="Tracheophyta" rank="species" species="brasiliensis">
<emphasis id="B968F279FF9E713AFA146E1EFC8DFCCC" box="[620,823,806,832]" italics="true" pageId="3" pageNumber="240">Flora Brasiliensis</emphasis>
(
<bibRefCitation id="EF8D539AFF9E713AFB3D6E1EFC65FCCC" author="Reissek, S." box="[837,991,806,832]" pageId="3" pageNumber="240" pagination="1 - 34" refId="ref6671" refString="Reissek, S. (1861) Celastrinea. In: Martius, C. F. P. &amp; Eichler, A. W. (Eds.) Flora Brasiliensis, v. 11, pars 1. Frid. Fleischer, Lipsiae [Leipzig], pp. 1 - 34, pl. 1 - 10." type="book chapter" year="1861">Reissek 1861</bibRefCitation>
, p. 27)
</taxonomicName>
, so there is not an older name that must have priority. However, the combination based on
<taxonomicName id="4C1C55E8FF9E713AFA806E73FC16FCE8" authorityName="Reissek" authorityYear="1861" box="[760,940,842,868]" class="Magnoliopsida" family="Celastraceae" genus="Maytenus" kingdom="Plantae" order="Celastrales" pageId="3" pageNumber="240" phylum="Tracheophyta" rank="species" species="laevis">
<emphasis id="B968F279FF9E713AFA806E73FC16FCE8" box="[760,940,842,868]" italics="true" pageId="3" pageNumber="240">Maytenus laevis</emphasis>
</taxonomicName>
is preferable to be used as it is more common in literature and herbarium determinations (see article 11.5. of the Shenzen Code,
<bibRefCitation id="EF8D539AFF9E713AFC4D6E56FAAFFC04" author="Turland, N. J. &amp; Wiersema, J. H. &amp; Barrie, F. R. &amp; Greuter, W. &amp; Hawksworth, D. L. &amp; Herendeen, P. S. &amp; Knapp, S. &amp; Kusber, W. - H. &amp; Li, D. - Z. &amp; Marhold, K. &amp; May, T. W. &amp; McNeill, J. &amp; Monro, A. M. &amp; Prado, J. &amp; Price, M. J. &amp; Smith, G. F." box="[1077,1301,878,904]" pageId="3" pageNumber="240" refId="ref7074" refString="Turland, N. J., Wiersema, J. H., Barrie, F. R., Greuter, W., Hawksworth, D. L., Herendeen, P. S., Knapp, S., Kusber, W. - H., Li, D. - Z., Marhold, K., May, T. W., McNeill, J., Monro, A. M., Prado, J., Price, M. J. &amp; Smith, G. F. (Eds.) (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzen Code): Adopted by the Nineteenth International Botanical Congress Shenzen, China, July 2017. Regnum Vegetabile 159. Koeltz Scientific Books, Glashutten, 254 pp. https: // doi. org / 10.12705 / Code. 2018" type="book" year="2018">
Turland
<emphasis id="B968F279FF9E713AFCED6E57FB75FC04" box="[1173,1231,878,904]" italics="true" pageId="3" pageNumber="240">et al.</emphasis>
2018
</bibRefCitation>
). I have seen herbarium specimens of
<taxonomicName id="4C1C55E8FF9E713AF9DE6EAAFDC5FC20" authorityName="Biral" authorityYear="2017" baseAuthorityName="Reissek" box="[422,639,914,940]" class="Magnoliopsida" family="Celastraceae" genus="Monteverdia" kingdom="Plantae" order="Celastrales" pageId="3" pageNumber="240" phylum="Tracheophyta" rank="species" species="laevis">
<emphasis id="B968F279FF9E713AF9DE6EAAFDC5FC20" box="[422,639,914,940]" italics="true" pageId="3" pageNumber="240">Monteverdia laevis</emphasis>
</taxonomicName>
from
<collectingCountry id="F30B6EFBFF9E713AFAB36EAAFC83FC20" box="[715,825,914,940]" name="Colombia" pageId="3" pageNumber="240">Colombia</collectingCountry>
(
<collectingRegion id="49D8E089FF9E713AFB336EAAFC1BFC20" box="[843,929,914,940]" country="Colombia" name="Vaupes" pageId="3" pageNumber="240">Vaupés</collectingRegion>
, but also cited for
<collectingRegion id="49D8E089FF9E713AFCF86EAAFB63FC20" box="[1152,1241,914,940]" country="Colombia" name="Guainia" pageId="3" pageNumber="240">Guainía</collectingRegion>
by
<bibRefCitation id="EF8D539AFF9E713AFD716EABFA25FC20" author="Kearns, D." box="[1289,1439,914,941]" pageId="3" pageNumber="240" pagination="190 - 197" refId="ref6037" refString="Kearns, D. (1998) Celastraceae. In: Steyermark, J. A., Berry, P. E., Holst, B. K. &amp; Yatskievych, K. (eds) Flora of the Venezuelan Guayana, vol. 4. Caesalpiniaceae - Ericaceae. Missouri Botanical Garden Press, St. Louis, pp. 190 - 197." type="book chapter" year="1998">Kearns 1998</bibRefCitation>
),
<collectingCountry id="F30B6EFBFF9E713AF8F06E8EFF41FC5C" box="[136,251,950,976]" name="Venezuela" pageId="3" pageNumber="240">Venezuela</collectingCountry>
(
<collectingRegion id="49D8E089FF9E713AF9726E8EFE38FC5C" box="[266,386,950,976]" country="Venezuela" name="Amazonas" pageId="3" pageNumber="240">Amazonas</collectingRegion>
and
<collectingRegion id="49D8E089FF9E713AF9C06E8EFDB4FC5C" box="[440,526,950,976]" country="Venezuela" name="Bolivar" pageId="3" pageNumber="240">Bolívar</collectingRegion>
) and
<collectingCountry id="F30B6EFBFF9E713AFA346E8EFD2AFC5C" box="[588,656,950,976]" name="Brazil" pageId="3" pageNumber="240">Brazil</collectingCountry>
(Amazonas).
</paragraph>
<paragraph id="8BA32E6BFF9E713AF8C56EE3FD3EFB24" blockId="3.[136,1452,662,1192]" pageId="3" pageNumber="240">
In the protologue,
<bibRefCitation id="EF8D539AFF9E713AF9E96EE2FD81FC78" author="Reissek, S." box="[401,571,986,1012]" pageId="3" pageNumber="240" pagination="1 - 34" refId="ref6671" refString="Reissek, S. (1861) Celastrinea. In: Martius, C. F. P. &amp; Eichler, A. W. (Eds.) Flora Brasiliensis, v. 11, pars 1. Frid. Fleischer, Lipsiae [Leipzig], pp. 1 - 34, pl. 1 - 10." type="book chapter" year="1861">Reissek (1861)</bibRefCitation>
cited the type of
<taxonomicName id="4C1C55E8FF9E713AFB7E6EE3FC02FC78" authorityName="Reissek" authorityYear="1861" box="[774,952,986,1012]" class="Magnoliopsida" family="Celastraceae" genus="Maytenus" kingdom="Plantae" order="Celastrales" pageId="3" pageNumber="240" phylum="Tracheophyta" rank="species" species="sapotiformis">
<emphasis id="B968F279FF9E713AFB7E6EE3FC02FC78" box="[774,952,986,1012]" italics="true" pageId="3" pageNumber="240">M. sapotiformis</emphasis>
</taxonomicName>
as deposited in the
<emphasis id="B968F279FF9E713AFCE66EE2FAD1FC78" box="[1182,1387,986,1012]" italics="true" pageId="3" pageNumber="240">Herbarium Martii</emphasis>
(“hb. Mart.”), currently housed at BR (
<bibRefCitation id="EF8D539AFF9E713AF9846EC6FD2EFB94" author="Forther, H." box="[508,660,1022,1048]" pageId="3" pageNumber="240" pagination="5 - 24" refId="ref5974" refString="Forther, H. (1994) Die Geschichte des Martius-Herbariums: seine Brasilienkollektion und Empfehlungen zur Typswahl. Sendtnera 2: 5 - 24" type="journal article" year="1994">Förther 1994</bibRefCitation>
). During a visit to BR, I located a unique specimen of
<taxonomicName id="4C1C55E8FF9E713AFC826EC7FA11FB94" authorityName="Reissek" authorityYear="1861" box="[1274,1451,1022,1048]" class="Magnoliopsida" family="Celastraceae" genus="Maytenus" kingdom="Plantae" order="Celastrales" pageId="3" pageNumber="240" phylum="Tracheophyta" rank="species" species="sapotiformis">
<emphasis id="B968F279FF9E713AFC826EC7FA11FB94" box="[1274,1451,1022,1048]" italics="true" pageId="3" pageNumber="240">M. sapotiformis</emphasis>
</taxonomicName>
in its
<taxonomicName id="4C1C55E8FF9E713AF8BE691AFEE9FBB0" box="[198,339,1058,1084]" class="Magnoliopsida" family="Celastraceae" kingdom="Plantae" order="Celastrales" pageId="3" pageNumber="240" phylum="Tracheophyta" rank="family">Celastraceae</taxonomicName>
collection; the specimen is annotated with Reisseks determination and his signature indicating that it is clearly the
<typeStatus id="54A790C9FF9E713AF94B697EFE2FFBEC" box="[307,405,1094,1120]" pageId="3" pageNumber="240" type="holotype">holotype</typeStatus>
(following
<bibRefCitation id="EF8D539AFF9E713AFA60697EFD03FBEC" author="McNeill, J." box="[536,697,1094,1120]" pageId="3" pageNumber="240" pagination="112 - 113" refId="ref6494" refString="McNeill, J. (2014) Holotype specimens and type citations: General issues. Taxon 63 (5): 112 - 113. https: // doi. org / 10.12705 / 635.7" type="journal article" year="2014">McNeill 2014</bibRefCitation>
). The specimen BR0000005737055 is selected here as
<typeStatus id="54A790C9FF9E713AFD66697EFA3FFBEC" box="[1310,1413,1094,1120]" pageId="3" pageNumber="240" type="lectotype">lectotype</typeStatus>
for
<taxonomicName id="4C1C55E8FF9E713AF8F06953FF4AFB08" authorityName="Biral" authorityYear="2017" baseAuthorityName="Reissek" box="[136,240,1130,1156]" class="Magnoliopsida" family="Celastraceae" genus="Monteverdia" kingdom="Plantae" order="Celastrales" pageId="3" pageNumber="240" phylum="Tracheophyta" rank="species" species="laevis">
<emphasis id="B968F279FF9E713AF8F06953FF4AFB08" box="[136,240,1130,1156]" italics="true" pageId="3" pageNumber="240">M. laevis</emphasis>
</taxonomicName>
because it is the unique mentioned specimen that brings a label signed by Reissek (Recommendation 9a.3 from the Shenzen Code,
<bibRefCitation id="EF8D539AFF9E713AF9E569B6FDC2FB24" author="Turland, N. J. &amp; Wiersema, J. H. &amp; Barrie, F. R. &amp; Greuter, W. &amp; Hawksworth, D. L. &amp; Herendeen, P. S. &amp; Knapp, S. &amp; Kusber, W. - H. &amp; Li, D. - Z. &amp; Marhold, K. &amp; May, T. W. &amp; McNeill, J. &amp; Monro, A. M. &amp; Prado, J. &amp; Price, M. J. &amp; Smith, G. F." box="[413,632,1166,1192]" pageId="3" pageNumber="240" refId="ref7074" refString="Turland, N. J., Wiersema, J. H., Barrie, F. R., Greuter, W., Hawksworth, D. L., Herendeen, P. S., Knapp, S., Kusber, W. - H., Li, D. - Z., Marhold, K., May, T. W., McNeill, J., Monro, A. M., Prado, J., Price, M. J. &amp; Smith, G. F. (Eds.) (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzen Code): Adopted by the Nineteenth International Botanical Congress Shenzen, China, July 2017. Regnum Vegetabile 159. Koeltz Scientific Books, Glashutten, 254 pp. https: // doi. org / 10.12705 / Code. 2018" type="book" year="2018">
Turland
<emphasis id="B968F279FF9E713AF98469B7FD8EFB24" box="[508,564,1166,1192]" italics="true" pageId="3" pageNumber="240">et al.</emphasis>
2018
</bibRefCitation>
).
</paragraph>
</treatment>
</document>

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@ -0,0 +1,369 @@
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<mods:identifier id="B5CBC40BB202EB73D66FD069273332B5" type="Zenodo-Dep">13878069</mods:identifier>
</mods:mods>
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<subSubSection id="C3067DE0FF9F713BF8F06CA2FC8EFE38" box="[136,820,410,437]" pageId="2" pageNumber="239" type="nomenclature">
<paragraph id="8BA32E6BFF9F713BF8F06CA2FC8EFE38" blockId="2.[136,1451,410,509]" box="[136,820,410,437]" pageId="2" pageNumber="239">
<taxonomicName id="4C1C55E8FF9F713BF8F06CA2FC8AFE38" ID-CoL="73Q7B" authority="(Mart. ex Reissek) Biral (2017: 689)" authorityName="Biral" authorityPageNumber="689" authorityYear="2017" baseAuthorityName="Mart. ex Reissek" box="[136,816,410,437]" class="Magnoliopsida" family="Celastraceae" genus="Monteverdia" kingdom="Plantae" order="Celastrales" pageId="2" pageNumber="239" phylum="Tracheophyta" rank="species" species="ilicifolia">
<emphasis id="B968F279FF9F713BF8F06CA2FE39FE39" bold="true" box="[136,387,410,437]" italics="true" pageId="2" pageNumber="239">Monteverdia ilicifolia</emphasis>
(Mart. ex Reissek)
<bibRefCitation id="EF8D539AFF9F713BFA1F6CA2FC8AFE38" author="Biral, L. &amp; Simmons, M. P. &amp; Smidt, E. &amp; Tembrock, L. R. &amp; Bolson, M. &amp; Archer, R. H. &amp; Lombardi, J. A." box="[615,816,410,436]" pageId="2" pageNumber="239" pagination="680 - 693" refId="ref5712" refString="Biral, L., Simmons, M. P., Smidt, E., Tembrock, L. R. Bolson, M. Archer, R. H. &amp; Lombardi, J. A. (2017) Systematics of the New World Maytenus (Celastraceae) and a new delimitation of the genus. Systematic Botany 42 (4): 680 - 693. https: // doi. org / 10.1600 / 036364417 X 696456" type="journal article" year="2017">Biral (2017: 689)</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3067DE0FF9F713BFB456CA3FEFDFE54" pageId="2" pageNumber="239" type="reference_group">
<paragraph id="8BA32E6BFF9F713BFB456CA3FEFDFE54" blockId="2.[136,1451,410,509]" pageId="2" pageNumber="239">
<taxonomicName id="4C1C55E8FF9F713BFB2D6CA3FACFFE38" authority="Martius ex Reissek (1861: 8" authorityName="Reissek" authorityYear="1861" box="[853,1397,410,437]" class="Magnoliopsida" family="Celastraceae" genus="Maytenus" kingdom="Plantae" order="Celastrales" pageId="2" pageNumber="239" phylum="Tracheophyta" rank="species" species="ilicifolia">
<emphasis id="B968F279FF9F713BFB2D6CA3FB93FE38" box="[853,1065,410,436]" italics="true" pageId="2" pageNumber="239">Maytenus ilicifolia</emphasis>
Martius ex
<bibRefCitation id="EF8D539AFF9F713BFCCD6CA2FACFFE38" author="Reissek, S." box="[1205,1397,410,436]" pageId="2" pageNumber="239" pagination="1 - 34" refId="ref6671" refString="Reissek, S. (1861) Celastrinea. In: Martius, C. F. P. &amp; Eichler, A. W. (Eds.) Flora Brasiliensis, v. 11, pars 1. Frid. Fleischer, Lipsiae [Leipzig], pp. 1 - 34, pl. 1 - 10." type="book chapter" year="1861">Reissek (1861: 8</bibRefCitation>
</taxonomicName>
, t. I, f. 11 a, b, t. VI).
</paragraph>
</subSubSection>
<subSubSection id="C3067DE0FF9F713BF9286C86FC08FE70" pageId="2" pageNumber="239" type="materials_examined">
<paragraph id="8BA32E6BFF9F713BF9286C86FC08FE70" blockId="2.[136,1451,410,509]" pageId="2" pageNumber="239">
Type:—
<materialsCitation id="3B742436FF9F713BF9D36C86FC14FE70" collectionCode="M" collectorName="Sellow" country="Uruguay" location="In Brasilia meridionali extratropica et ad Montevideo" pageId="2" pageNumber="239" specimenCount="1" typeStatus="lectotype">
BRAZIL,
<collectingCountry id="F30B6EFBFF9F713BFA596C86FD11FE54" box="[545,683,446,472]" name="Uruguay" pageId="2" pageNumber="239">URUGUAY</collectingCountry>
.
<location id="8EC378B0FF9F713BFACF6C87FAA7FE54" LSID="urn:lsid:plazi:treatment:03B59F7DFF9F713AF8F06CA2FF72FE8C:8EC378B0FF9F713BFACF6C87FAA7FE54" box="[695,1309,446,473]" country="Uruguay" name="In Brasilia meridionali extratropica et ad Montevideo" pageId="2" pageNumber="239">In Brasilia meridionali extratropica et ad Montevideo</location>
, 1821-1823,
<emphasis id="B968F279FF9F713BF8F06CDAFEA8FE70" box="[136,274,482,508]" italics="true" pageId="2" pageNumber="239">
<collectorName id="26E94BBDFF9F713BF8F06CDAFF69FE70" box="[136,211,482,508]" pageId="2" pageNumber="239">Sellow</collectorName>
d768
</emphasis>
(
<typeStatus id="54A790C9FF9F713BF9586CDAFE31FE70" box="[288,395,482,508]" pageId="2" pageNumber="239" type="lectotype">lectotype</typeStatus>
,
<collectionCode id="ED0DB6AEFF9F713BF9EF6CDBFE08FE71" box="[407,434,483,509]" pageId="2" pageNumber="239">M</collectionCode>
, designated by
<bibRefCitation id="EF8D539AFF9F713BFA276CDAFC1FFE70" author="Biral, L. &amp; Lombardi, J. A." box="[607,933,482,508]" pageId="2" pageNumber="239" pagination="468 - 469" refId="ref5532" refString="Biral, L. &amp; Lombardi, J. A. (2012) (2057) Proposal to conserve the name Maytenus ilicifolia Mart. ex Reissek against M. ilicifolia (Schrad.) Planch. (Celastraceae). Taxon 61 (2): 468 - 469. https: // doi. org / 10.1002 / tax. 612022" type="journal article" year="2012">Biral &amp; Lombardi 2012: 468</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3067DE0FF9F713BF8F06F16FD68FCCC" pageId="2" pageNumber="239" type="reference_group">
<paragraph id="8BA32E6BFF9F713BF8F06F16FC5EFDE4" blockId="2.[136,1452,558,832]" pageId="2" pageNumber="239">
=
<taxonomicName id="4C1C55E8FF9F713BF8E36F17FD94FDC8" authority="Reissek (1861: 6" authorityName="Reissek" authorityYear="1861" box="[155,558,558,580]" class="Magnoliopsida" family="Celastraceae" genus="Maytenus" kingdom="Plantae" order="Celastrales" pageId="2" pageNumber="239" phylum="Tracheophyta" rank="species" species="acanthophylla">
<emphasis id="B968F279FF9F713BF8E36F17FE3CFDCF" box="[155,390,558,580]" italics="true" pageId="2" pageNumber="239">Maytenus acanthophylla</emphasis>
<bibRefCitation id="EF8D539AFF9F713BF9F46F16FD94FDC8" author="Reissek, S." box="[396,558,558,580]" pageId="2" pageNumber="239" pagination="1 - 34" refId="ref6671" refString="Reissek, S. (1861) Celastrinea. In: Martius, C. F. P. &amp; Eichler, A. W. (Eds.) Flora Brasiliensis, v. 11, pars 1. Frid. Fleischer, Lipsiae [Leipzig], pp. 1 - 34, pl. 1 - 10." type="book chapter" year="1861">Reissek (1861: 6</bibRefCitation>
</taxonomicName>
, t. I, f. 5)
<taxonomicName id="4C1C55E8FF9F713BFAF16F16FCE5FDCF" authority="Loesener" authorityName="Loesener" authorityYear="2020" box="[649,863,558,580]" class="Magnoliopsida" family="Celastraceae" genus="Maytenus" kingdom="Plantae" order="Celastrales" pageId="2" pageNumber="239" phylum="Tracheophyta" rank="variety" species="acanthophylla" variety="eurostos">
var.
<emphasis id="B968F279FF9F713BFACA6F17FCB8FDC8" box="[690,770,559,580]" italics="true" pageId="2" pageNumber="239">eurostos</emphasis>
Loesener
</taxonomicName>
in
<bibRefCitation id="EF8D539AFF9F713BFB056F16FB94FDCF" author="Kuntze, C. E. O." box="[893,1070,558,579]" pageId="2" pageNumber="239" refId="ref6143" refString="Kuntze, C. E. O. (1898) Revisio Generum Plantarum vascularium omnium atque cellularium multarum secundum Leges Nomenclaturae Internationales cum enumeratione Plantarum Exoticarum in itineribus mundi collectarum. Pars III. Part 2. Commissionen. Arthur Feliz, Leipzig; Dulau &amp; Co., London; U. Hoepli, Milano; Gust. E. Stechert, New York; Charles Klincksieck, Paris, 201 pp." type="book" year="1898">Kuntze (1898: 37)</bibRefCitation>
. Type:—
<collectingCountry id="F30B6EFBFF9F713BFCFC6F16FB64FDCF" box="[1156,1246,558,579]" name="Brazil" pageId="2" pageNumber="239">BRAZIL</collectingCountry>
.
<collectingRegion id="49D8E089FF9F713BFC9E6F16FA13FDC8" box="[1254,1449,558,580]" country="Brazil" name="Mato Grosso do Sul" pageId="2" pageNumber="239">Mato Grosso do Sul</collectingRegion>
: Corumbá, [late]
<date id="FFA208ABFF9F713BF9226F6AFE6DFDEB" box="[346,471,594,615]" pageId="2" pageNumber="239" value="1892-08">August 1892</date>
,
<emphasis id="B968F279FF9F713BF9996F6BFDF6FDE4" box="[481,588,595,616]" italics="true" pageId="2" pageNumber="239">Kuntze s.n.</emphasis>
(
<typeStatus id="54A790C9FF9F713BFA206F6AFD0AFDE4" box="[600,688,594,616]" pageId="2" pageNumber="239" type="holotype">holotype</typeStatus>
, NY [NY01031056]!).
<emphasis id="B968F279FF9F713BFBE86F6BFC5EFDE4" box="[912,996,595,616]" italics="true" pageId="2" pageNumber="239">syn. nov.</emphasis>
</paragraph>
<paragraph id="8BA32E6BFF9F713BF8F06F4EFAA6FD58" blockId="2.[136,1452,558,832]" pageId="2" pageNumber="239">
=
<taxonomicName id="4C1C55E8FF9F713BF8E26F4FFD95FD07" authority="Reissek (1861: 7" authorityName="Reissek" authorityYear="1861" box="[154,559,630,652]" class="Magnoliopsida" family="Celastraceae" genus="Maytenus" kingdom="Plantae" order="Celastrales" pageId="2" pageNumber="239" phylum="Tracheophyta" rank="species" species="castaneiformis">
<emphasis id="B968F279FF9F713BF8E26F4FFE33FD07" box="[154,393,630,652]" italics="true" pageId="2" pageNumber="239">Maytenus castaneiformis</emphasis>
<bibRefCitation id="EF8D539AFF9F713BF9F66F4EFD95FD07" author="Reissek, S." box="[398,559,630,652]" pageId="2" pageNumber="239" pagination="1 - 34" refId="ref6671" refString="Reissek, S. (1861) Celastrinea. In: Martius, C. F. P. &amp; Eichler, A. W. (Eds.) Flora Brasiliensis, v. 11, pars 1. Frid. Fleischer, Lipsiae [Leipzig], pp. 1 - 34, pl. 1 - 10." type="book chapter" year="1861">Reissek (1861: 7</bibRefCitation>
</taxonomicName>
, t. I, f. 7, as “
<emphasis id="B968F279FF9F713BFAD16F4EFCFCFD07" box="[681,838,630,651]" italics="true" pageId="2" pageNumber="239">castaneaeformis</emphasis>
”, see the article 60.10 of the Shenzen Code,
<bibRefCitation id="EF8D539AFF9F713BFC926F4EFA1BFD07" author="Turland, N. J. &amp; Wiersema, J. H. &amp; Barrie, F. R. &amp; Greuter, W. &amp; Hawksworth, D. L. &amp; Herendeen, P. S. &amp; Knapp, S. &amp; Kusber, W. - H. &amp; Li, D. - Z. &amp; Marhold, K. &amp; May, T. W. &amp; McNeill, J. &amp; Monro, A. M. &amp; Prado, J. &amp; Price, M. J. &amp; Smith, G. F." box="[1258,1441,630,652]" pageId="2" pageNumber="239" refId="ref7074" refString="Turland, N. J., Wiersema, J. H., Barrie, F. R., Greuter, W., Hawksworth, D. L., Herendeen, P. S., Knapp, S., Kusber, W. - H., Li, D. - Z., Marhold, K., May, T. W., McNeill, J., Monro, A. M., Prado, J., Price, M. J. &amp; Smith, G. F. (Eds.) (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzen Code): Adopted by the Nineteenth International Botanical Congress Shenzen, China, July 2017. Regnum Vegetabile 159. Koeltz Scientific Books, Glashutten, 254 pp. https: // doi. org / 10.12705 / Code. 2018" type="book" year="2018">
Turland
<emphasis id="B968F279FF9F713BFD426F4FFAD3FD07" box="[1338,1385,630,652]" italics="true" pageId="2" pageNumber="239">et al.</emphasis>
2018
</bibRefCitation>
).
<collectingMunicipality id="6BC7B411FF9F713BF8C56FA2FF55FD23" box="[189,239,666,687]" pageId="2" pageNumber="239">Type</collectingMunicipality>
:—
<collectingCountry id="F30B6EFBFF9F713BF9746FA2FEDCFD23" box="[268,358,666,687]" name="Brazil" pageId="2" pageNumber="239">BRAZIL</collectingCountry>
.
<collectingRegion id="49D8E089FF9F713BF9176FA2FE11FD3C" box="[367,427,666,688]" country="Brazil" name="Goias" pageId="2" pageNumber="239">Goiás</collectingRegion>
: [Buenolândia] ad
<location id="8EC378B0FF9F713BFA1A6FA2FD75FD23" LSID="urn:lsid:plazi:treatment:03B59F7DFF9F713AF8F06CA2FF72FE8C:8EC378B0FF9F713BFA1A6FA2FD75FD23" box="[610,719,666,688]" country="Brazil" municipality="Type" name="Rio Bugres" pageId="2" pageNumber="239" stateProvince="Goias">Rio Bugres</location>
[prope Barra], s.d. [
<date id="FFA208ABFF9F713BFBEB6FA2FBB9FD23" box="[915,1027,666,688]" pageId="2" pageNumber="239" value="1819-04">
<collectingDate id="EFE6F143FF9F713BFBEB6FA2FBB9FD23" box="[915,1027,666,688]" pageId="2" pageNumber="239" value="1819-04">April 1819</collectingDate>
</date>
],
<emphasis id="B968F279FF9F713BFC6B6FA2FBD3FD3C" box="[1043,1129,666,688]" italics="true" pageId="2" pageNumber="239">Pohl s.n.</emphasis>
[
<emphasis id="B968F279FF9F713BFC0F6FA2FB1DFD23" box="[1143,1191,666,687]" italics="true" pageId="2" pageNumber="239">1710</emphasis>
] (
<typeStatus id="54A790C9FF9F713BFCC56FA2FAA3FD3C" box="[1213,1305,666,688]" pageId="2" pageNumber="239" type="lectotype">lectotype</typeStatus>
, W [0059514]! image, designated here;
<typeStatus id="54A790C9FF9F713BF9DF6F86FD9DFD58" box="[423,551,702,724]" pageId="2" pageNumber="239" type="isolectotype">isolectotypes</typeStatus>
, BR [BR0000005739677]!, W [0059515]! image, B† [F neg 13276]).
<emphasis id="B968F279FF9F713BFCB06F87FAA6FD58" box="[1224,1308,703,724]" italics="true" pageId="2" pageNumber="239">syn. nov.</emphasis>
</paragraph>
<paragraph id="8BA32E6BFF9F713BF8F06FDAFD68FCCC" blockId="2.[136,1452,558,832]" pageId="2" pageNumber="239">
=
<taxonomicName id="4C1C55E8FF9F713BF8E76FDBFDFAFD74" authority="Reissek (1861: 6 - 7" authorityName="Reissek" authorityYear="1861" box="[159,576,738,760]" class="Magnoliopsida" family="Celastraceae" genus="Maytenus" kingdom="Plantae" order="Celastrales" pageId="2" pageNumber="239" phylum="Tracheophyta" rank="species" species="macrodonta">
<emphasis id="B968F279FF9F713BF8E76FDBFEC3FD7B" box="[159,377,738,760]" italics="true" pageId="2" pageNumber="239">Maytenus macrodonta</emphasis>
<bibRefCitation id="EF8D539AFF9F713BF9FA6FDAFDFAFD74" author="Reissek, S." box="[386,576,738,760]" pageId="2" pageNumber="239" pagination="1 - 34" refId="ref6671" refString="Reissek, S. (1861) Celastrinea. In: Martius, C. F. P. &amp; Eichler, A. W. (Eds.) Flora Brasiliensis, v. 11, pars 1. Frid. Fleischer, Lipsiae [Leipzig], pp. 1 - 34, pl. 1 - 10." type="book chapter" year="1861">Reissek (1861: 6-7</bibRefCitation>
</taxonomicName>
, t. I, f. 6).
<collectingMunicipality id="6BC7B411FF9F713BFACE6FDAFD52FD7B" box="[694,744,738,759]" pageId="2" pageNumber="239">Type</collectingMunicipality>
:—
<collectingCountry id="F30B6EFBFF9F713BFB7D6FDAFCE5FD7B" box="[773,863,738,759]" name="Brazil" pageId="2" pageNumber="239">BRAZIL</collectingCountry>
. [
<collectingRegion id="49D8E089FF9F713BFB086FDAFC17FD74" box="[880,941,738,760]" country="Brazil" name="Goias" pageId="2" pageNumber="239">Goiás</collectingRegion>
]
<location id="8EC378B0FF9F713BFBC46FDAFBC7FD74" LSID="urn:lsid:plazi:treatment:03B59F7DFF9F713AF8F06CA2FF72FE8C:8EC378B0FF9F713BFBC46FDAFBC7FD74" box="[956,1149,738,760]" country="Brazil" municipality="Type" name="Habitat in Brasiliae" pageId="2" pageNumber="239" stateProvince="Goias">Habitat in Brasiliae</location>
meridionalis interioribus [Rio Pillano], s.d. [late
<date id="FFA208ABFF9F713BF9176E3EFE58FC90" box="[367,482,774,796]" pageId="2" pageNumber="239" value="1919-03">
<collectingDate id="EFE6F143FF9F713BF9176E3EFE58FC90" box="[367,482,774,796]" pageId="2" pageNumber="239" value="1919-03">March 1919</collectingDate>
</date>
-
<date id="FFA208ABFF9F713BF9906E3EFDC7FC97" box="[488,637,774,796]" pageId="2" pageNumber="239" value="1820-02">
<collectingDate id="EFE6F143FF9F713BF9906E3EFDC7FC97" box="[488,637,774,796]" pageId="2" pageNumber="239" value="1820-02">February 1820</collectingDate>
</date>
],
<emphasis id="B968F279FF9F713BFAF56E3EFD59FC90" box="[653,739,774,796]" italics="true" pageId="2" pageNumber="239">Pohl s.n.</emphasis>
[
<emphasis id="B968F279FF9F713BFA896E3EFC9BFC97" box="[753,801,774,795]" italics="true" pageId="2" pageNumber="239">2444</emphasis>
] (
<typeStatus id="54A790C9FF9F713BFB4D6E3EFC2BFC90" box="[821,913,774,796]" pageId="2" pageNumber="239" type="lectotype">lectotype</typeStatus>
, W [0059540]! image, designated here;
<typeStatus id="54A790C9FF9F713BFD6C6E3EFA2EFC90" box="[1300,1428,774,796]" pageId="2" pageNumber="239" type="isolectotype">isolectotypes</typeStatus>
, F [frag., 870042, barcode V0044275F]! image).
<emphasis id="B968F279FF9F713BFA036E12FD68FCCC" box="[635,722,810,832]" italics="true" pageId="2" pageNumber="239">Syn. nov.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3067DE0FF9F713AF8F06E57FF72FE8C" lastPageId="3" lastPageNumber="240" pageId="2" pageNumber="239" type="discussion">
<paragraph id="8BA32E6BFF9F713BF8F06E57FCEDFAB5" blockId="2.[136,1452,878,2093]" pageId="2" pageNumber="239">
<taxonomicName id="4C1C55E8FF9F713BF8F06E57FD81FC04" authorityName="Loesener" authorityYear="2020" box="[136,571,878,905]" class="Magnoliopsida" family="Celastraceae" genus="Maytenus" kingdom="Plantae" order="Celastrales" pageId="2" pageNumber="239" phylum="Tracheophyta" rank="variety" species="acanthophylla" variety="eurostos">
<emphasis id="B968F279FF9F713BF8F06E57FE25FC04" box="[136,415,878,904]" italics="true" pageId="2" pageNumber="239">Maytenus acanthophylla</emphasis>
var.
<emphasis id="B968F279FF9F713BF9A56E57FD81FC04" box="[477,571,879,904]" italics="true" pageId="2" pageNumber="239">eurostos</emphasis>
</taxonomicName>
is a forgotten name, absent in all
<taxonomicName id="4C1C55E8FF9F713BFBB56E56FBE0FC04" box="[973,1114,878,904]" class="Magnoliopsida" family="Celastraceae" kingdom="Plantae" order="Celastrales" pageId="2" pageNumber="239" phylum="Tracheophyta" rank="family">Celastraceae</taxonomicName>
literature I accessed after its publication. I came cross the
<typeStatus id="54A790C9FF9F713BF99E6EAAFDF2FC20" box="[486,584,914,940]" pageId="2" pageNumber="239" type="holotype">holotype</typeStatus>
(not annotated as such) when examined New World
<taxonomicName id="4C1C55E8FF9F713BFCC76EABFA96FC20" box="[1215,1324,915,940]" class="Magnoliopsida" family="Celastraceae" genus="Maytenus" kingdom="Plantae" order="Celastrales" pageId="2" pageNumber="239" phylum="Tracheophyta" rank="genus">
<emphasis id="B968F279FF9F713BFCC76EABFA96FC20" box="[1215,1324,915,940]" italics="true" pageId="2" pageNumber="239">Maytenus</emphasis>
</taxonomicName>
specimens provided on loan from the NY herbarium. The specimen is derived from the Herbarium of Otto Kuntze (purchased by Andrew Carnegie for the New York Botanical Garden, see
<bibRefCitation id="EF8D539AFF9F713BFB6E6EE2FC1DFC78" author="Zanoni, T. A." box="[790,935,986,1012]" pageId="2" pageNumber="239" pagination="551 - 571" refId="ref7447" refString="Zanoni, T. A. (1980) Otto Kuntze, botanist. I. Biography, bibliography and travels. Brittonia 32 (4): 551 - 571. https: // doi. org / 10.2307 / 2806169" type="journal article" year="1980">Zanoni 1980</bibRefCitation>
), signed by Ludwig Eduard Theodor Loesener (the species author) and its label includes the following diagnosis: “
<emphasis id="B968F279FF9F713BFBE06EC7FB2CFBB0" italics="true" pageId="2" pageNumber="239">
<taxonomicName id="4C1C55E8FF9F713BFBE06EC7FF7EFBB0" authority="Loes." authorityName="Loesener" authorityYear="2020" class="Magnoliopsida" family="Celastraceae" form="eurostos" genus="Maytenus" kingdom="Plantae" order="Celastrales" pageId="2" pageNumber="239" phylum="Tracheophyta" rank="form" species="acanthophylla" status="f. nov.">Maytenus acanthophylla Reiss. forma eurostos Loes.</taxonomicName>
<taxonomicNameLabel id="A25B4F02FF9F713BF8B5691AFEF5FBB0" box="[205,335,1058,1084]" pageId="2" pageNumber="239" rank="form">forma nova</taxonomicNameLabel>
foliis paullo latioribus, basi magis truncatis spinis robustioribus diversa
</emphasis>
”. Later, when the name was validly published, Loesener (in
<bibRefCitation id="EF8D539AFF9F713BFA59697FFD09FBEC" author="Kuntze, C. E. O." box="[545,691,1094,1121]" pageId="2" pageNumber="239" refId="ref6143" refString="Kuntze, C. E. O. (1898) Revisio Generum Plantarum vascularium omnium atque cellularium multarum secundum Leges Nomenclaturae Internationales cum enumeratione Plantarum Exoticarum in itineribus mundi collectarum. Pars III. Part 2. Commissionen. Arthur Feliz, Leipzig; Dulau &amp; Co., London; U. Hoepli, Milano; Gust. E. Stechert, New York; Charles Klincksieck, Paris, 201 pp." type="book" year="1898">Kuntze 1898</bibRefCitation>
) did so as a variety of
<taxonomicName id="4C1C55E8FF9F713BFBCE697FFB71FBEC" authorityName="Reissek" authorityYear="1861" box="[950,1227,1094,1120]" class="Magnoliopsida" family="Celastraceae" genus="Maytenus" kingdom="Plantae" order="Celastrales" pageId="2" pageNumber="239" phylum="Tracheophyta" rank="species" species="acanthophylla">
<emphasis id="B968F279FF9F713BFBCE697FFB71FBEC" box="[950,1227,1094,1120]" italics="true" pageId="2" pageNumber="239">Maytenus acanthophylla</emphasis>
</taxonomicName>
and not as a forma. Some specimens of
<taxonomicName id="4C1C55E8FF9F713BF91D6952FDE2FB08" authorityName="Biral" authorityYear="2017" baseAuthorityName="Mart. ex Reissek" box="[357,600,1130,1156]" class="Magnoliopsida" family="Celastraceae" genus="Monteverdia" kingdom="Plantae" order="Celastrales" pageId="2" pageNumber="239" phylum="Tracheophyta" rank="species" species="ilicifolia">
<emphasis id="B968F279FF9F713BF91D6952FDE2FB08" box="[357,600,1130,1156]" italics="true" pageId="2" pageNumber="239">Monteverdia ilicifolia</emphasis>
</taxonomicName>
, especially those from
<collectingCountry id="F30B6EFBFF9F713BFB206952FC10FB08" box="[856,938,1130,1156]" name="Bolivia" pageId="2" pageNumber="239">Bolivia</collectingCountry>
and Brazilian states of Mato Grosso and Mato Grosso do Sul, are confused with
<taxonomicName id="4C1C55E8FF9F713BF98369B6FC94FB24" box="[507,814,1166,1192]" class="Magnoliopsida" family="Celastraceae" genus="Monteverdia" kingdom="Plantae" order="Celastrales" pageId="2" pageNumber="239" phylum="Tracheophyta" rank="species" species="acanthophylla">
<emphasis id="B968F279FF9F713BF98369B6FC94FB24" box="[507,814,1166,1192]" italics="true" pageId="2" pageNumber="239">Monteverdia acanthophylla</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF8D539AFF9F713BFB4469B6FC4AFB24" author="Reissek, S." box="[828,1008,1166,1192]" pageId="2" pageNumber="239" pagination="1 - 34" refId="ref6671" refString="Reissek, S. (1861) Celastrinea. In: Martius, C. F. P. &amp; Eichler, A. W. (Eds.) Flora Brasiliensis, v. 11, pars 1. Frid. Fleischer, Lipsiae [Leipzig], pp. 1 - 34, pl. 1 - 10." type="book chapter" year="1861">Reissek 1861: 6</bibRefCitation>
, t. I, f. 5)
<bibRefCitation id="EF8D539AFF9F713BFC2F69B6FAADFB24" author="Biral, L. &amp; Simmons, M. P. &amp; Smidt, E. &amp; Tembrock, L. R. &amp; Bolson, M. &amp; Archer, R. H. &amp; Lombardi, J. A." box="[1111,1303,1166,1192]" pageId="2" pageNumber="239" pagination="680 - 693" refId="ref5712" refString="Biral, L., Simmons, M. P., Smidt, E., Tembrock, L. R. Bolson, M. Archer, R. H. &amp; Lombardi, J. A. (2017) Systematics of the New World Maytenus (Celastraceae) and a new delimitation of the genus. Systematic Botany 42 (4): 680 - 693. https: // doi. org / 10.1600 / 036364417 X 696456" type="journal article" year="2017">Biral (2017: 688)</bibRefCitation>
because they have broad leaves with long spines on the margin but can be differentiated by their spherical fruits, as opposed to the tetragonal fruits of
<taxonomicName id="4C1C55E8FF9F713BF92669EEFD28FB7C" box="[350,658,1238,1264]" class="Magnoliopsida" family="Celastraceae" genus="Monteverdia" kingdom="Plantae" order="Celastrales" pageId="2" pageNumber="239" phylum="Tracheophyta" rank="species" species="acanthophylla">
<emphasis id="B968F279FF9F713BF92669EEFD28FB7C" box="[350,658,1238,1264]" italics="true" pageId="2" pageNumber="239">Monteverdia acanthophylla</emphasis>
</taxonomicName>
.
<taxonomicName id="4C1C55E8FF9F713BFAD869EEFC29FB7C" authorityName="Biral" authorityYear="2017" baseAuthorityName="Mart. ex Reissek" box="[672,915,1238,1264]" class="Magnoliopsida" family="Celastraceae" genus="Monteverdia" kingdom="Plantae" order="Celastrales" pageId="2" pageNumber="239" phylum="Tracheophyta" rank="species" species="ilicifolia">
<emphasis id="B968F279FF9F713BFAD869EEFC29FB7C" box="[672,915,1238,1264]" italics="true" pageId="2" pageNumber="239">Monteverdia ilicifolia</emphasis>
</taxonomicName>
is widely distributed in the forests of
<collectingCountry id="F30B6EFBFF9F713BFD4D69EEFA1DFB7C" box="[1333,1447,1238,1264]" name="Argentina" pageId="2" pageNumber="239">Argentina</collectingCountry>
,
<collectingCountry id="F30B6EFBFF9F713BF8F069C2FF67FA98" box="[136,221,1274,1300]" name="Bolivia" pageId="2" pageNumber="239">Bolivia</collectingCountry>
,
<collectingCountry id="F30B6EFBFF9F713BF89269C2FE94FA98" box="[234,302,1274,1300]" name="Brazil" pageId="2" pageNumber="239">Brazil</collectingCountry>
(chiefly, in south region),
<collectingCountry id="F30B6EFBFF9F713BFA2769C3FD7DFA99" box="[607,711,1275,1301]" name="Paraguay" pageId="2" pageNumber="239">Paraguay</collectingCountry>
and
<collectingCountry id="F30B6EFBFF9F713BFB7869C3FCD9FA99" box="[768,867,1275,1301]" name="Uruguay" pageId="2" pageNumber="239">Uruguay</collectingCountry>
, whereas
<taxonomicName id="4C1C55E8FF9F713BFBAC69C2FAB1FA98" box="[980,1291,1274,1300]" class="Magnoliopsida" family="Celastraceae" genus="Monteverdia" kingdom="Plantae" order="Celastrales" pageId="2" pageNumber="239" phylum="Tracheophyta" rank="species" species="acanthophylla">
<emphasis id="B968F279FF9F713BFBAC69C2FAB1FA98" box="[980,1291,1274,1300]" italics="true" pageId="2" pageNumber="239">Monteverdia acanthophylla</emphasis>
</taxonomicName>
is endemic to
<collectingCountry id="F30B6EFBFF9F713BF8F06826FF75FAB4" box="[136,207,1310,1336]" name="Brazil" pageId="2" pageNumber="239">Brazil</collectingCountry>
, especially in areas with caatinga vegetation (
<bibRefCitation id="EF8D539AFF9F713BFAB76826FCF1FAB5" author="BFG &amp; The Brazil Flora Group" box="[719,843,1310,1337]" pageId="2" pageNumber="239" pagination="1085 - 1113" refId="ref5489" refString="BFG - The Brazil Flora Group (2015) Growing knowledge: an overview of Seed Plant diversity in Brazil. Rodriguesia 66 (4): 1085 - 1113. https: // doi. org / 10.1590 / 2175 - 7860201566409" type="journal article" year="2015">BFG 2015</bibRefCitation>
).
</paragraph>
<paragraph id="8BA32E6BFF9F713BF8C5687AFD3DF92C" blockId="2.[136,1452,878,2093]" pageId="2" pageNumber="239">
<bibRefCitation id="EF8D539AFF9F713BF8C5687AFDBBFAD0" author="Carvalho-Okano, R. M." box="[189,513,1346,1372]" pageId="2" pageNumber="239" refId="ref5826" refString="Carvalho-Okano, R. M. (1992) Estudos taxonomicos do genero Maytenus (Celastraceae) no Brasil extra-amazonico. Thesis, Universidade Estadual de Campinas, Campinas, x + 253 pp." type="book" year="1992">Carvalho-Okano (1992: 228)</bibRefCitation>
considered
<taxonomicName id="4C1C55E8FF9F713BFAFF687BFC25FAD0" authorityName="Reissek" authorityYear="1861" box="[647,927,1346,1372]" class="Magnoliopsida" family="Celastraceae" genus="Maytenus" kingdom="Plantae" order="Celastrales" pageId="2" pageNumber="239" phylum="Tracheophyta" rank="species" species="castaneiformis">
<emphasis id="B968F279FF9F713BFAFF687BFC25FAD0" box="[647,927,1346,1372]" italics="true" pageId="2" pageNumber="239">Maytenus castaneiformis</emphasis>
</taxonomicName>
in the “Untreated species” section of her thesis because she did not have access to any original material from this species. In addition to this mention, I found no citation of this name in literature. The morphological characters available from the type specimens and mentioned in the protologue overlap those found for
<taxonomicName id="4C1C55E8FF9F713BFA406896FC91FA44" authorityName="Biral" authorityYear="2017" baseAuthorityName="Mart. ex Reissek" box="[568,811,1454,1480]" class="Magnoliopsida" family="Celastraceae" genus="Monteverdia" kingdom="Plantae" order="Celastrales" pageId="2" pageNumber="239" phylum="Tracheophyta" rank="species" species="ilicifolia">
<emphasis id="B968F279FF9F713BFA406896FC91FA44" box="[568,811,1454,1480]" italics="true" pageId="2" pageNumber="239">Monteverdia ilicifolia</emphasis>
</taxonomicName>
. The leaves with two or three pairs of spines concentrated only on the apex presented in the type specimens are quite characteristics of
<taxonomicName id="4C1C55E8FF9F713BFBB268EBFBF7FA60" authorityName="Biral" authorityYear="2017" baseAuthorityName="Mart. ex Reissek" box="[970,1101,1490,1517]" class="Magnoliopsida" family="Celastraceae" genus="Monteverdia" kingdom="Plantae" order="Celastrales" pageId="2" pageNumber="239" phylum="Tracheophyta" rank="species" species="ilicifolia">
<emphasis id="B968F279FF9F713BFBB268EBFC5BFA60" box="[970,993,1491,1516]" italics="true" pageId="2" pageNumber="239">M</emphasis>
.
<emphasis id="B968F279FF9F713BFB9668EAFBF7FA60" box="[1006,1101,1490,1516]" italics="true" pageId="2" pageNumber="239">ilicifolia</emphasis>
</taxonomicName>
; thus,
<taxonomicName id="4C1C55E8FF9F713BFCEC68EBFA11FA60" authorityName="Reissek" authorityYear="1861" box="[1172,1451,1490,1516]" class="Magnoliopsida" family="Celastraceae" genus="Maytenus" kingdom="Plantae" order="Celastrales" pageId="2" pageNumber="239" phylum="Tracheophyta" rank="species" species="castaneiformis">
<emphasis id="B968F279FF9F713BFCEC68EBFA11FA60" box="[1172,1451,1490,1516]" italics="true" pageId="2" pageNumber="239">Maytenus castaneiformis</emphasis>
</taxonomicName>
is considered here as its synonym. The specimen
<emphasis id="B968F279FF9F713BFAB868CEFC8CF99C" box="[704,822,1526,1552]" italics="true" pageId="2" pageNumber="239">
Pohl
<quantity id="4CE4838EFF9F713BFA8668CEFC8CF99C" box="[766,822,1526,1552]" metricMagnitude="1" metricUnit="m" metricValue="4.3434" pageId="2" pageNumber="239" unit="in" value="1710.0">1710</quantity>
</emphasis>
in W [0059514] is designated here as
<typeStatus id="54A790C9FF9F713BFC8C68CEFAE1F99C" box="[1268,1371,1526,1552]" pageId="2" pageNumber="239" type="lectotype">lectotype</typeStatus>
for
<taxonomicName id="4C1C55E8FF9F713BFDF568CFFE97F9B8" authorityName="Reissek" authorityYear="1861" class="Magnoliopsida" family="Celastraceae" genus="Maytenus" kingdom="Plantae" order="Celastrales" pageId="2" pageNumber="239" phylum="Tracheophyta" rank="species" species="castaneiformis">
<emphasis id="B968F279FF9F713BFDF568CFFA1EF99C" box="[1421,1444,1527,1552]" italics="true" pageId="2" pageNumber="239">M</emphasis>
.
<emphasis id="B968F279FF9F713BF8F06B22FE97F9B8" box="[136,301,1562,1588]" italics="true" pageId="2" pageNumber="239">castaneiformis</emphasis>
</taxonomicName>
because it is constituted by a complete branch with an attached label bearing a handwritten morphological description by
<location id="8EC378B0FF9F713BF94B6B06FE30F9D4" LSID="urn:lsid:plazi:treatment:03B59F7DFF9F713AF8F06CA2FF72FE8C:8EC378B0FF9F713BF94B6B06FE30F9D4" box="[307,394,1598,1624]" country="Brazil" county="Pohl's" municipality="The" name="Reissek" pageId="2" pageNumber="239">Reissek</location>
that is identical to that provided by him in the protologue (see Recommendation 9a.3. from the Shenzen Code,
<bibRefCitation id="EF8D539AFF9F713BF9276B5AFD83F9F0" author="Turland, N. J. &amp; Wiersema, J. H. &amp; Barrie, F. R. &amp; Greuter, W. &amp; Hawksworth, D. L. &amp; Herendeen, P. S. &amp; Knapp, S. &amp; Kusber, W. - H. &amp; Li, D. - Z. &amp; Marhold, K. &amp; May, T. W. &amp; McNeill, J. &amp; Monro, A. M. &amp; Prado, J. &amp; Price, M. J. &amp; Smith, G. F." box="[351,569,1634,1660]" pageId="2" pageNumber="239" refId="ref7074" refString="Turland, N. J., Wiersema, J. H., Barrie, F. R., Greuter, W., Hawksworth, D. L., Herendeen, P. S., Knapp, S., Kusber, W. - H., Li, D. - Z., Marhold, K., May, T. W., McNeill, J., Monro, A. M., Prado, J., Price, M. J. &amp; Smith, G. F. (Eds.) (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzen Code): Adopted by the Nineteenth International Botanical Congress Shenzen, China, July 2017. Regnum Vegetabile 159. Koeltz Scientific Books, Glashutten, 254 pp. https: // doi. org / 10.12705 / Code. 2018" type="book" year="2018">
Turland
<emphasis id="B968F279FF9F713BF9C56B5BFE4CF9F0" box="[445,502,1634,1660]" italics="true" pageId="2" pageNumber="239">et al.</emphasis>
2018
</bibRefCitation>
).
<collectingMunicipality id="6BC7B411FF9F713BFA336B5AFDCDF9F0" box="[587,631,1634,1660]" pageId="2" pageNumber="239">The</collectingMunicipality>
collection date is not provided on the label but was taken from the
<collectingCounty id="62C256E7FF9F713BFD1D6B5AFA11F9F0" box="[1381,1451,1634,1660]" pageId="2" pageNumber="239">Pohls</collectingCounty>
itinerary in
<collectingCountry id="F30B6EFBFF9F713BF9736BBEFEF5F92C" box="[267,335,1670,1696]" name="Brazil" pageId="2" pageNumber="239">Brazil</collectingCountry>
according to
<bibRefCitation id="EF8D539AFF9F713BF9906BBEFD39F92C" author="Urban, I." box="[488,643,1670,1696]" pageId="2" pageNumber="239" pagination="1 - 268" refId="ref7239" refString="Urban, I. (1906) Vitae itineraque collectorum botanicorum, notae collaboratum biographicae, Flora Brasiliensis ratio edendi chronologica, systema, index familiarum. In: Martius, C. F. P., Eichler, A. W. &amp; Urban, I. (Eds.) Flora Brasiliensis, v. 1, pars 1. R. Oldenbourg, Munchen et Leipzig, pp. 1 - 268." type="book chapter" year="1906">Urban (1906)</bibRefCitation>
.
</paragraph>
<paragraph id="8BA32E6BFF9F713AF8C56B93FF72FE8C" blockId="2.[136,1452,878,2093]" lastBlockId="3.[136,1451,158,256]" lastPageId="3" lastPageNumber="240" pageId="2" pageNumber="239">
<taxonomicName id="4C1C55E8FF9F713BF8C56B93FE0DF948" authorityName="Reissek" authorityYear="1861" box="[189,439,1706,1732]" class="Magnoliopsida" family="Celastraceae" genus="Maytenus" kingdom="Plantae" order="Celastrales" pageId="2" pageNumber="239" phylum="Tracheophyta" rank="species" species="macrodonta">
<emphasis id="B968F279FF9F713BF8C56B93FE0DF948" box="[189,439,1706,1732]" italics="true" pageId="2" pageNumber="239">Maytenus macrodonta</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="EF8D539AFF9F713BFAFC6B92FC96F948" author="Reissek, S." box="[644,812,1706,1732]" pageId="2" pageNumber="239" pagination="1 - 34" refId="ref6671" refString="Reissek, S. (1861) Celastrinea. In: Martius, C. F. P. &amp; Eichler, A. W. (Eds.) Flora Brasiliensis, v. 11, pars 1. Frid. Fleischer, Lipsiae [Leipzig], pp. 1 - 34, pl. 1 - 10." type="book chapter" year="1861">Reissek (1861)</bibRefCitation>
in Martius
<taxonomicName id="4C1C55E8FF9F713BFBCC6B92FBC4F948" baseAuthorityName="Reissek" baseAuthorityYear="1861" box="[948,1150,1706,1732]" class="Magnoliopsida" family="Celastraceae" genus="Flora" kingdom="Plantae" order="Celastrales" pageId="2" pageNumber="239" phylum="Tracheophyta" rank="species" species="brasiliensis">
<emphasis id="B968F279FF9F713BFBCC6B92FBC4F948" box="[948,1150,1706,1732]" italics="true" pageId="2" pageNumber="239">Flora Brasiliensis</emphasis>
</taxonomicName>
. The description presented in the protologue is short and essentially based on vegetative characters, such as branches and leaves (shape, measures, number of spines and secondary veins). No flower or fruit features are mentioned and the unique information cited about the inflorescence [“
<emphasis id="B968F279FF9F713BF9CF6A2EFD50F8BC" box="[439,746,1814,1840]" italics="true" pageId="2" pageNumber="239">
<collectingCountry id="F30B6EFBFF9F713BF9CF6A2EFE45F8BC" box="[439,511,1814,1840]" name="Indonesia" pageId="2" pageNumber="239">Flores</collectingCountry>
axillares, aggregati
</emphasis>
”,
<bibRefCitation id="EF8D539AFF9F713BFB716A2EFC60F8BC" author="Reissek, S." box="[777,986,1814,1840]" pageId="2" pageNumber="239" pagination="1 - 34" refId="ref6671" refString="Reissek, S. (1861) Celastrinea. In: Martius, C. F. P. &amp; Eichler, A. W. (Eds.) Flora Brasiliensis, v. 11, pars 1. Frid. Fleischer, Lipsiae [Leipzig], pp. 1 - 34, pl. 1 - 10." type="book chapter" year="1861">Reissek (1861: 7)</bibRefCitation>
] is not informative since all
<taxonomicName id="4C1C55E8FF9F713BFD4F6A2FFA1EF8BC" box="[1335,1444,1815,1840]" class="Magnoliopsida" family="Celastraceae" genus="Maytenus" kingdom="Plantae" order="Celastrales" pageId="2" pageNumber="239" phylum="Tracheophyta" rank="genus">
<emphasis id="B968F279FF9F713BFD4F6A2FFA1EF8BC" box="[1335,1444,1815,1840]" italics="true" pageId="2" pageNumber="239">Maytenus</emphasis>
</taxonomicName>
/
<taxonomicName id="4C1C55E8FF9F713BF8F06A02FED4F8D8" box="[136,366,1850,1876]" pageId="2" pageNumber="239">
<emphasis id="B968F279FF9F713BF8F06A02FEACF8D8" box="[136,278,1850,1876]" italics="true" pageId="2" pageNumber="239">Monteverdia</emphasis>
species
</taxonomicName>
with spinose leaves share this type of inflorescence. Because the description lacks details and no original material was accessible,
<bibRefCitation id="EF8D539AFF9F713BFA7C6A66FCA8F8F4" author="Carvalho-Okano, R. M." box="[516,786,1886,1912]" pageId="2" pageNumber="239" refId="ref5826" refString="Carvalho-Okano, R. M. (1992) Estudos taxonomicos do genero Maytenus (Celastraceae) no Brasil extra-amazonico. Thesis, Universidade Estadual de Campinas, Campinas, x + 253 pp." type="book" year="1992">Carvalho-Okano (1992)</bibRefCitation>
excluded the name from her thesis. Now, considering the protologue, the type collections and the
<taxonomicName id="4C1C55E8FF9F713BFA3D6ABAFD68F810" box="[581,722,1922,1948]" class="Magnoliopsida" family="Celastraceae" kingdom="Plantae" order="Celastrales" pageId="2" pageNumber="239" phylum="Tracheophyta" rank="family">Celastraceae</taxonomicName>
collections from Goiás state, it is determined that
<taxonomicName id="4C1C55E8FF9F713BFD786ABBFA11F810" authorityName="Reissek" authorityYear="1861" box="[1280,1451,1922,1948]" class="Magnoliopsida" family="Celastraceae" genus="Maytenus" kingdom="Plantae" order="Celastrales" pageId="2" pageNumber="239" phylum="Tracheophyta" rank="species" species="macrodonta">
<emphasis id="B968F279FF9F713BFD786ABBFA11F810" box="[1280,1451,1922,1948]" italics="true" pageId="2" pageNumber="239">M. macrodonta</emphasis>
</taxonomicName>
is conspecific with
<taxonomicName id="4C1C55E8FF9F713BF91F6A9EFDE7F84C" authorityName="Biral" authorityYear="2017" baseAuthorityName="Mart. ex Reissek" box="[359,605,1958,1984]" class="Magnoliopsida" family="Celastraceae" genus="Monteverdia" kingdom="Plantae" order="Celastrales" pageId="2" pageNumber="239" phylum="Tracheophyta" rank="species" species="ilicifolia">
<emphasis id="B968F279FF9F713BF91F6A9EFDE7F84C" box="[359,605,1958,1984]" italics="true" pageId="2" pageNumber="239">Monteverdia ilicifolia</emphasis>
</taxonomicName>
.
<taxonomicName id="4C1C55E8FF9F713BFA156A9EFCD9F84C" authorityName="Biral" authorityYear="2017" baseAuthorityName="Mart. ex Reissek" box="[621,867,1958,1984]" class="Magnoliopsida" family="Celastraceae" genus="Monteverdia" kingdom="Plantae" order="Celastrales" pageId="2" pageNumber="239" phylum="Tracheophyta" rank="species" species="ilicifolia">
<emphasis id="B968F279FF9F713BFA156A9EFCD9F84C" box="[621,867,1958,1984]" italics="true" pageId="2" pageNumber="239">Monteverdia ilicifolia</emphasis>
</taxonomicName>
is easily characterized by young twigs flattened or carinated, coriaceous leaves spirally arranged, leaf with one to seven pairs of spines regularly arranged in the margin or concentrated in the apex, and multi-flowered fasciculate inflorescence. The protologue of
<taxonomicName id="4C1C55E8FF9F713BFCE26AD7FAFFF784" authorityName="Reissek" authorityYear="1861" box="[1178,1349,2030,2056]" class="Magnoliopsida" family="Celastraceae" genus="Maytenus" kingdom="Plantae" order="Celastrales" pageId="2" pageNumber="239" phylum="Tracheophyta" rank="species" species="macrodonta">
<emphasis id="B968F279FF9F713BFCE26AD7FAFFF784" box="[1178,1349,2030,2056]" italics="true" pageId="2" pageNumber="239">M. macrodonta</emphasis>
</taxonomicName>
does not mention locality, collector number, or date of collection. The collector number [
<emphasis id="B968F279FF9F713BFC6E652AFBF4F7A0" box="[1046,1102,2066,2092]" italics="true" pageId="2" pageNumber="239">2444</emphasis>
] and the locality [Rio Pillano] included in this paper are taken from the label of the designated
<typeStatus id="54A790C9FF9E713AFB256DA6FC7DFF34" box="[861,967,158,184]" pageId="3" pageNumber="240" type="lectotype">lectotype</typeStatus>
. According to Pohls itinerary, provided by
<bibRefCitation id="EF8D539AFF9E713AF8F06DFAFE98FF50" author="Urban, I." box="[136,290,194,220]" pageId="3" pageNumber="240" pagination="1 - 268" refId="ref7239" refString="Urban, I. (1906) Vitae itineraque collectorum botanicorum, notae collaboratum biographicae, Flora Brasiliensis ratio edendi chronologica, systema, index familiarum. In: Martius, C. F. P., Eichler, A. W. &amp; Urban, I. (Eds.) Flora Brasiliensis, v. 1, pars 1. R. Oldenbourg, Munchen et Leipzig, pp. 1 - 268." type="book chapter" year="1906">Urban (1906)</bibRefCitation>
, he visited the Pilões river in the Brazilian state of Goiás twice between late
<date id="FFA208ABFF9E713AFCFE6DFAFAB5FF50" box="[1158,1295,194,220]" pageId="3" pageNumber="240" value="1919-03">March 1919</date>
and
<date id="FFA208ABFF9E713AFD3E6DFAFF79FE8C" pageId="3" pageNumber="240" value="1820-02">February 1820</date>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,355 @@
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<mods:title id="34DD4E2DE9AB685646CC03C7DF681D2E">The Dacini fruit flies of Borneo: an annotated checklist with 89 species including three new to science (Tephritidae, Dacinae)</mods:title>
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<mods:namePart id="55E6A79869B7EF4B541976F889733E2A">Doorenweerd, Camiel</mods:namePart>
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<mods:affiliation id="79E3CCDE1FAB10326F43D8087D698CC7">Department of Plant and Environmental Protection Sciences, Entomology Section, College of Tropical Agriculture and Human Resources, University of Hawaiʻi at Mānoa, 3050 Maile Way, Honolulu, Hawaii, 96822 - 2231, USA</mods:affiliation>
</mods:name>
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<mods:namePart id="0015B0DE84463342883DF4FBA2B0A5F9">Chung, Arthur Y. C.</mods:namePart>
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<mods:affiliation id="DF7A6079D10AC1512C5BAE7106698D71">Sabah Forestry Department, Forest Research Centre, P. O. Box 1407, 90715 Sandakan, Sabah, Malaysia</mods:affiliation>
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<mods:namePart id="B27C2CF4AA9807D6778526F50C2A86A3">Mustapeng, Andi Maryani A.</mods:namePart>
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<mods:affiliation id="EF60500B3F480057565B42273CE7BC5C">Sabah Forestry Department, Forest Research Centre, P. O. Box 1407, 90715 Sandakan, Sabah, Malaysia</mods:affiliation>
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<mods:namePart id="6176548150903B036269D5B8965FE268">Rubinoff, Daniel</mods:namePart>
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<mods:affiliation id="CF1E15D08560D2DC92CAFD2F2D7D2428">Department of Plant and Environmental Protection Sciences, Entomology Section, College of Tropical Agriculture and Human Resources, University of Hawaiʻi at Mānoa, 3050 Maile Way, Honolulu, Hawaii, 96822 - 2231, USA</mods:affiliation>
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<subSubSection id="100CAB54640DB876EF7D6E38BDEF7FB1" type="nomenclature">
<paragraph id="49E6494DB2FD27F45D60EE3F4DAB650B">
<taxonomicName id="BEF39FBA5BF9B078963B5EACB313C146" authority="Doorenweerd" authorityName="Doorenweerd &amp; Chung &amp; Mustapeng &amp; Rubinoff" authorityYear="2025" class="Insecta" family="Tephritidae" genus="Zeugodacus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="cataracta" status="sp. nov." subGenus="Zeugodacus">
<emphasis id="FFAD5308AE1C2EB8EB4283D0B0DA0D02" italics="true">Zeugodacus</emphasis>
(
<emphasis id="F2DD38051DC2C46644594ADBF7EA6F35" italics="true">Zeugodacus</emphasis>
)
<emphasis id="8AD78114A1AF69AA44F506953664E7EA" italics="true">cataracta</emphasis>
Doorenweerd
</taxonomicName>
<taxonomicNameLabel id="EC9E5E036E5693BFA19EA2F14B117401" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="73532FCDE188A2999BEDDB56A990E9B4" type="description">
<paragraph id="DD7544208D722D2FF68584F486E2C86C">
<figureCitation id="B9DA438F8B99E1D4013EA664EEF0AEF4" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Holotype male of Zeugodacus cataracta sp. nov., UHIM. ms 08929 A dorsal view of head and thorax B dorsal view of abdomen C anterior view of the head D lateral view E dorso-anterior view of the abdomen showing the ceromata F detail image of the genitalia showing the surstylus length G left wing, dissected and slide mounted H dorsal view of the abdomen of the paratype (ms 08929) showing variation in the abdomen markings." figureDoi="10.3897/zookeys.1240.148768.figure5" httpUri="https://binary.pensoft.net/fig/1348697">Fig. 5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5YBI" type="type material">
<paragraph id="4110248F2F65286FDC99333774EA53FB">
<heading id="5F4B4B38CDC2CE900B6684869D6CD6D9" reason="title">Type material.</heading>
</paragraph>
<paragraph id="F04572894A1BA6BE71A735AFED6DE689">
<materialsCitation id="FA5917A71BFE7C2BDA6E7C112E45141A" collectingDate="2018-12-03" collectingDateMax="2018-12-05" collectingDateMin="2018-12-03" collectionCode="UHIM" collectorName="Leg. D. Rubinoff &amp; C. Doorenweerd." country="Malaysia" latitude="4.9506" location="University of Hawaii Insect Museum" longLatPrecision="7" longitude="117.8061" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<emphasis id="0C546107B27B748FA8240EFD3505223D" bold="true">
<emphasis id="8941416C56900723F620096BA6C314CC" bold="true" italics="true">
<typeStatus id="57A96B1A289CC1584F336E6CBB5900E9" type="holotype">Holotype</typeStatus>
</emphasis>
.
</emphasis>
Male. “
<collectingCountry id="C8F2CE0AC05D47D83FB84E5ABD35A9FB" name="Malaysia">Malaysia</collectingCountry>
:
<collectingRegion id="3C63CA9B89B8B1DD9A007E59CF74B070" country="Malaysia" name="Sabah">Sabah</collectingRegion>
: Danum Valley Tembaling waterfall trail. WGS 84
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EMZBI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[117.806100,4.950600]}">
<geoCoordinate id="A96F12A84A58878B09FE834D2F9F76D2" degrees="4.9506" direction="north" orientation="latitude" precision="5" value="4.9506">4.9506</geoCoordinate>
,
<geoCoordinate id="EF00E88E653C68A0C3885ACD2ED13DEE" degrees="117.8061" direction="east" orientation="longitude" precision="5" value="117.8061">117.8061</geoCoordinate>
</named-content>
<date id="54315D4C03CADBFC7CB3D2446B0FCE16" value="2018-12-03" valueMax="2018-12-05" valueMin="2018-12-03">
<collectingDate id="F69E04C527D72771A16779DA395F711D" value="2018-12-03" valueMax="2018-12-05" valueMin="2018-12-03">35. xii. 2018</collectingDate>
</date>
<collectingMethod id="D072342423E31DAF2FAA8B977C24D0BB">Zingerone trap</collectingMethod>
.
<collectorName id="0A93129705FDC78CF90CF89892FF02E7">Leg. D. Rubinoff</collectorName>
&amp;
<collectorName id="E373CB1F066665F5A30835864BE71E15">C. Doorenweerd.</collectorName>
DNA sample ms 08933. ” Deposited at the
<location id="91A111DA9451348860B561249D6A4BB4" LSID="urn:lsid:plazi:treatment:217BC2A4A4225A07BD31FB265F842C3D:91A111DA9451348860B561249D6A4BB4" country="Malaysia" latitude="4.9506" longLatPrecision="7" longitude="117.8061" name="University of Hawaii Insect Museum" stateProvince="Sabah">University of Hawaii Insect Museum</location>
reg. no.
<abbrev content-type="institution" id="ABBRID0ERZBI" xlink_title="University of Hawaiʻi at Mānoa">
<collectionCode id="9C1A492BB314C52FB7C41B12C6CEF8F4">UHIM</collectionCode>
</abbrev>
. ms 08933
</materialsCitation>
.
<emphasis id="3A7D9FDDE2AF6316FB79D25675AEE5DB" bold="true" italics="true">
<typeStatus id="69CDF81F0B7B57D6C4D4EC34D533A2C0" type="paratype">Paratype</typeStatus>
</emphasis>
.
<specimenCount id="39707E49D94771A2A06834DA5E5DEFE7" count="1" type="male">One male</specimenCount>
. “
<collectingCountry id="9537F8B7C6C24AB7EDB49199F2C52F1E" name="Malaysia">Malaysia</collectingCountry>
,
<collectingRegion id="484BB6D4AD3AF10BC085D864CC4D0F59" country="Malaysia" name="Sabah">Sabah</collectingRegion>
, Danum Valley access road. WGS 84
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E3ZBI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[117.812600,4.968900]}">
<geoCoordinate id="97A3213DF79DD94852AFA420E9B6829C" degrees="4.9689" direction="north" orientation="latitude" precision="5" value="4.9689">4.9689</geoCoordinate>
,
<geoCoordinate id="35E6B5D85BE098A2B68425620570906B" degrees="117.8126" direction="east" orientation="longitude" precision="5" value="117.8126">117.8126</geoCoordinate>
</named-content>
<date id="A384B738E923E454CD7EE1D9D925AB8E" value="2018-12-03" valueMax="2018-12-05" valueMin="2018-12-03">35. xii. 2018</date>
Zingerone trap. Leg. D. Rubinoff &amp; C. Doorenweerd. DNA sample ms 08929. ” Deposited at the
<collectingRegion id="F7CCF65788BEB23C263D8E12591884DC" country="Malaysia" name="Sabah">Sabah</collectingRegion>
Forestry Department Insect Collection reg. no.
<abbrev content-type="institution" id="ABBRID0EB1BI" xlink_title="University of Hawaiʻi at Mānoa">
<collectionCode id="C52ED519DF4C07AD9B447F5C6E5EAA10">UHIM</collectionCode>
</abbrev>
. ms 08929.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EG1BI" type="Differential diagnosis">
<paragraph id="5F128703C1B577D568559E9574BDF905">
<heading id="B0E85223771051375B504274F5211D0A" reason="title">Differential diagnosis.</heading>
</paragraph>
<paragraph id="6E4D7BD8C52D95139E78095704854399">
The wing pattern of
<taxonomicName id="C04F1A75BC493FC86DCAB2B95C9AB9F4" authorityName="Doorenweerd &amp; Chung &amp; Mustapeng &amp; Rubinoff" authorityYear="2025" class="Insecta" family="Tephritidae" genus="Zeugodacus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="cataracta">
<emphasis id="9928E79D0754FC952B412DCECA0C21E6" italics="true">Zeugodacus cataracta</emphasis>
</taxonomicName>
is strikingly different from any other species of
<taxonomicName id="413E8ABF0C861D860C457A7B39465D46" class="Insecta" family="Tephritidae" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="tribe" tribe="Dacini">Dacini</taxonomicName>
, with a medial dark band that extends down to midway of cell
<emphasis id="9452400B17AAAA135B3FF79816C709D0" italics="true">dm</emphasis>
, dark marking throughout cell
<emphasis id="A1880ED158B2925D1EF1321396F89AEF" italics="true">br</emphasis>
, and three dark smudges in the distal part of the wing. The species is further unique in having vein CuA
<subScript id="4AD1EAF84A67C08361679F1DE877E5F9">2</subScript>
and A
<subScript id="19A601FC260F058E4AA0DE5B6C5A0724">1</subScript>
merge at ~ 1 / 2 the length of A
<subScript id="A03792E7CF3D3967B71B3E267BE671A3">1</subScript>
, leaving a short cell
<emphasis id="B0FC280793D760B7562631958C5437D6" italics="true">cup</emphasis>
, and all antennal segments are remarkably short, combined less than half the height of the head.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EI2BI" type="Molecular diagnostics">
<paragraph id="3886A8502090895EDE125FC5526D0865">
<heading id="DF44949CD3F3D98061C95179C8D5E8F0" reason="title">Molecular diagnostics.</heading>
</paragraph>
<paragraph id="8011FBD5218256B2AB4760F4FBCE8978">
The
<abbrev id="ABBRID0EO2BI" xlink_title="Cytochrome C Oxidase I">COI</abbrev>
- 5 P 3 P sequences of ms 08929 and ms 08933 are most similar to published sequences of
<taxonomicName id="109CFCD282D554C0D92D1401DBD1B6A6" authority="(Bezzi, 1913)" baseAuthorityName="Bezzi" baseAuthorityYear="1913" class="Insecta" family="Tephritidae" genus="Zeugodacus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="scutellaris">
<emphasis id="497E4107C169F9CACA21F98BFF33AD11" italics="true">Zeugodacus scutellaris</emphasis>
(Bezzi, 1913)
</taxonomicName>
, but at 8.9 % minimum pairwise difference (
<bibRefCitation id="EADDBB0FBFC83F17D85A55CAA7365090" DOI="10.1111/1755-0998.13987" author="Doorenweerd C &amp; San Jose M &amp; Leblanc L &amp; Barr N &amp; Geib SM &amp; Chung AYC &amp; Dupuis JR &amp; Ekayanti A &amp; Fiegalan E &amp; Hemachandra KS &amp; Hossain MA &amp; Huang C &amp; Hsu Y &amp; Morris KY &amp; Maryani A &amp; Mustapeng A &amp; Niogret J &amp; Pham TH &amp; Thi Nguyen N &amp; Sirisena UGAI &amp; Todd T &amp; Rubinoff D" refId="B15" refString="Doorenweerd C, San Jose M, Leblanc L, Barr N, Geib SM, Chung AYC, Dupuis JR, Ekayanti A, Fiegalan E, Hemachandra KS, Hossain MA, Huang C, Hsu Y, Morris KY, Maryani A, Mustapeng A, Niogret J, Pham TH, Thi Nguyen N, Sirisena UGAI, Todd T, Rubinoff D (2024 b) Towards a better future for DNA barcoding: Evaluating monophyly- and distance-based species identification using COI gene fragments of Dacini fruit flies. Molecular Ecology Resources 13987. https://doi.org/10.1111/1755-0998.13987">Doorenweerd et al. 2024 b</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EB3BI" type="description">
<paragraph id="AD24D5E10685B59A7712AC59F40136F0">
<heading id="D40E767573E21C50D6421C9295D30CFF" reason="title">Description.</heading>
</paragraph>
<paragraph id="D2D66B48EE4001EC32DEAAA32D1C10A2">
<emphasis id="81DC81AC6124CB5502FBF93AED7A7B08" bold="true">
Male.
<emphasis id="2637D3F1B2CA5AB12A0DAAEAA6922634" bold="true" italics="true">Head</emphasis>
</emphasis>
(Fig.
<figureCitation id="C6562A728480C60FBBF70D2B1DC3DD71" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Holotype male of Zeugodacus cataracta sp. nov., UHIM. ms 08929 A dorsal view of head and thorax B dorsal view of abdomen C anterior view of the head D lateral view E dorso-anterior view of the abdomen showing the ceromata F detail image of the genitalia showing the surstylus length G left wing, dissected and slide mounted H dorsal view of the abdomen of the paratype (ms 08929) showing variation in the abdomen markings." figureDoi="10.3897/zookeys.1240.148768.figure5" httpUri="https://binary.pensoft.net/fig/1348697">5 C</figureCitation>
). Fulvous; face dark with indistinct oval black spots in the antennal furrows. Antennal segments dark fulvous and short, combined length less than half the height of the head.
<emphasis id="C1DAF0841A790880C373BA321BF8A384" bold="true" italics="true">Thorax</emphasis>
(Fig.
<figureCitation id="F8C1A091F60E146FCBCEB7935DCF654F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Holotype male of Zeugodacus cataracta sp. nov., UHIM. ms 08929 A dorsal view of head and thorax B dorsal view of abdomen C anterior view of the head D lateral view E dorso-anterior view of the abdomen showing the ceromata F detail image of the genitalia showing the surstylus length G left wing, dissected and slide mounted H dorsal view of the abdomen of the paratype (ms 08929) showing variation in the abdomen markings." figureDoi="10.3897/zookeys.1240.148768.figure5" httpUri="https://binary.pensoft.net/fig/1348697">5 A, D</figureCitation>
). Scutum and pleural areas black with narrow red-brown areas surrounding parts of the yellow markings. Yellow markings: postpronotal lobes; notopleura; presutural area adjacent to notopleura; postsutural yellow lateral vittae that narrow sharply posteriorly and do not reach intra-alar seta; medial elongate postsutural vitta; mesopleural stripe dorsally wider than notopleuron but does not reach postpronotal lobe; anatergite; katatergite. Scutellum yellow with a narrow black basal band. Setae: two pair scutellar; one pair prescutellar; one pair intra-alar; one pair posterior supra-alar; one pair anterior supra-alar; two pair notopleural and four scapular.
<emphasis id="C3E9399174C235B1941E6417ABE0577C" bold="true" italics="true">Abdomen</emphasis>
(Fig.
<figureCitation id="AF7BD7032BEFE555ECF32A7ACA4C5A4C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Holotype male of Zeugodacus cataracta sp. nov., UHIM. ms 08929 A dorsal view of head and thorax B dorsal view of abdomen C anterior view of the head D lateral view E dorso-anterior view of the abdomen showing the ceromata F detail image of the genitalia showing the surstylus length G left wing, dissected and slide mounted H dorsal view of the abdomen of the paratype (ms 08929) showing variation in the abdomen markings." figureDoi="10.3897/zookeys.1240.148768.figure5" httpUri="https://binary.pensoft.net/fig/1348697">5 B, E, F, H</figureCitation>
). Oval to diamond-shaped; terga free; pecten of setae on tergum III absent. Posterior lobe of the male surstylus long (Fig.
<figureCitation id="F81852A7A294CE341D464365C145E19C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Holotype male of Zeugodacus cataracta sp. nov., UHIM. ms 08929 A dorsal view of head and thorax B dorsal view of abdomen C anterior view of the head D lateral view E dorso-anterior view of the abdomen showing the ceromata F detail image of the genitalia showing the surstylus length G left wing, dissected and slide mounted H dorsal view of the abdomen of the paratype (ms 08929) showing variation in the abdomen markings." figureDoi="10.3897/zookeys.1240.148768.figure5" httpUri="https://binary.pensoft.net/fig/1348697">5 F</figureCitation>
). Abdomen overall fulvous with interrupted black T on segments III
<collectionCode id="EE73A322766B4D497D56084A4058F3A6" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
. Anterolateral corners of segment
<collectionCode id="D7B3F496B03C8714FE20D0C533DB32D9">IV</collectionCode>
black.
<typeStatus id="34E146887FE46B473B133315D29FFBAA" type="paratype">Paratype</typeStatus>
with more extensive black lateral markings on tergum
<collectionCode id="D5454747D4DBE0909AEB5B1CE0FC764E">IV</collectionCode>
and
<collectionCode id="252B0EEC466ABE3EAC3A025A5C1AF424" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
(Fig.
<figureCitation id="6AC83C5B35C4889BFEFA0E8C6168B60A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Holotype male of Zeugodacus cataracta sp. nov., UHIM. ms 08929 A dorsal view of head and thorax B dorsal view of abdomen C anterior view of the head D lateral view E dorso-anterior view of the abdomen showing the ceromata F detail image of the genitalia showing the surstylus length G left wing, dissected and slide mounted H dorsal view of the abdomen of the paratype (ms 08929) showing variation in the abdomen markings." figureDoi="10.3897/zookeys.1240.148768.figure5" httpUri="https://binary.pensoft.net/fig/1348697">5 H</figureCitation>
). Ceromata yellow and indistinct.
<emphasis id="55E2C8AD542D9471F8ED8A3FA39C2B30" bold="true" italics="true">Legs</emphasis>
(Fig.
<figureCitation id="E60F057300FD08BA9F85E52F5CE03E96" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Holotype male of Zeugodacus cataracta sp. nov., UHIM. ms 08929 A dorsal view of head and thorax B dorsal view of abdomen C anterior view of the head D lateral view E dorso-anterior view of the abdomen showing the ceromata F detail image of the genitalia showing the surstylus length G left wing, dissected and slide mounted H dorsal view of the abdomen of the paratype (ms 08929) showing variation in the abdomen markings." figureDoi="10.3897/zookeys.1240.148768.figure5" httpUri="https://binary.pensoft.net/fig/1348697">5 D</figureCitation>
). Legs overall fulvous. Outer surface of the fore tibia black, distal third of hind tibia black.
<emphasis id="66D5E287EE39913A98A3F50E468FE9AF" bold="true" italics="true">Wings</emphasis>
(Fig.
<figureCitation id="6550D6E91D010219DB7378FCC83D862C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Holotype male of Zeugodacus cataracta sp. nov., UHIM. ms 08929 A dorsal view of head and thorax B dorsal view of abdomen C anterior view of the head D lateral view E dorso-anterior view of the abdomen showing the ceromata F detail image of the genitalia showing the surstylus length G left wing, dissected and slide mounted H dorsal view of the abdomen of the paratype (ms 08929) showing variation in the abdomen markings." figureDoi="10.3897/zookeys.1240.148768.figure5" httpUri="https://binary.pensoft.net/fig/1348697">5 G</figureCitation>
). Wing length
<quantity id="27A48DC4EE26B392462CAB85FBB56E65" metricMagnitude="-3" metricUnit="m" metricValue="5.8" unit="mm" value="5.8">5.8 mm</quantity>
. Cells
<emphasis id="D740F643831EE233161B40EEED4EACF4" italics="true">bc</emphasis>
and
<emphasis id="913C85026930F1C2EEF3E690522E96D8" italics="true">c</emphasis>
clear, faint tint in cell
<emphasis id="B0099018588197B8C1681DAADB25C7FD" italics="true">sc.</emphasis>
Costal band confluent with vein
<collectionCode id="947A302C5C0965A83868534C062DC869" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
<subScript id="FA1E74C07F5DFD49B51CD0B36D4A8938">2 + 3</subScript>
and continues at more or less the same width until just past where vein
<collectionCode id="72C47DCD9C0446016C22F2B9D4D39258" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
<subScript id="D625D7FEA8BA4817D70C81C06D3CB80B">4 + 5</subScript>
reaches the costa. Anal streak absent, vein CuA
<subScript id="4338A555FBFE5FB1CA9F06C3B2E6236B">2</subScript>
and A
<subScript id="F308EDDA86FAA4074476C7C4A03B8225">1</subScript>
merge at # 1 / 2 the length of A
<subScript id="FC9C7F95EC96FAE71605FA2A8A18573F">1</subScript>
. A medial dark band extends from the costal band down to midway of cell
<emphasis id="49E323B36517732B151BCEA71CEDAF6E" italics="true">dm</emphasis>
and dark marking throughout cell
<emphasis id="0E25A7637A3D005AFCEFD21121781B77" italics="true">br.</emphasis>
The distal part of the wing has three dark brown smudges below vein
<collectionCode id="DBE4FC370C1842B93039EAB8DD59148F" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
<subScript id="7949AA0074D366DAF2136B59B55726F8">4 + 5</subScript>
.
<emphasis id="B84E570271645E4E3BD6A1D53F3B4892" bold="true">Female.</emphasis>
Unknown.
</paragraph>
<caption id="ECB8EBD6C8E7E9F2B67885DA1005581E" ID-DOI="10.3897/zookeys.1240.148768.figure5" ID-arpha="8EBF0663-DC9D-5471-90A7-630387C90E9F" httpUri="https://binary.pensoft.net/fig/1348697" startId="F5">
<paragraph id="2E86EE8D2F3A94FEDFD29DDD904160B0">
<label id="DE69099FD0D8CA0937880AA544CFAF8F">Figure 5.</label>
</paragraph>
<paragraph id="DA5BEA3DA880EA9C9C46AD91FE5A2CFE">
Holotype male of
<taxonomicName id="CF11AF5AA23CBE6CFCEFCFBE0448F1C9" authorityName="Doorenweerd &amp; Chung &amp; Mustapeng &amp; Rubinoff" authorityYear="2025" class="Insecta" family="Tephritidae" genus="Zeugodacus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="cataracta" status="sp. nov.">
<emphasis id="49EA59377A696EC641ECF8D433F07D41" italics="true">Zeugodacus cataracta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="80C3312276B9C127ADA30A2443E7B322" rank="species">sp. nov.</taxonomicNameLabel>
,
<abbrev content-type="institution" id="ABBRID0E55BI" xlink_title="University of Hawaiʻi at Mānoa">UHIM</abbrev>
. ms 08929
<emphasis id="4885702F025989F31FEEF66B051099D4" bold="true">A</emphasis>
dorsal view of head and thorax
<emphasis id="BEEED6709E315E4048D0EC1FD642AC19" bold="true">B</emphasis>
dorsal view of abdomen
<emphasis id="7562245B81FA9BE052FF4F6D1C7B4905" bold="true">C</emphasis>
anterior view of the head
<emphasis id="498CAD49849290D04463E0E7A4845E8E" bold="true">D</emphasis>
lateral view
<emphasis id="92D607358C0810DF9A5B3E462EADD329" bold="true">E</emphasis>
dorso-anterior view of the abdomen showing the ceromata
<emphasis id="05164E7FDDF0C7E3DA64DA43287CD6A7" bold="true">F</emphasis>
detail image of the genitalia showing the surstylus length
<emphasis id="FAC45D84F8B349A6C5F0EC6B3D33043D" bold="true">G</emphasis>
left wing, dissected and slide mounted
<emphasis id="92CB13D0881C647F5401E6C0FF255D76" bold="true">H</emphasis>
dorsal view of the abdomen of the paratype (ms 08929) showing variation in the abdomen markings.
</paragraph>
</caption>
<paragraph id="AE0A50C7A8E936648BEBCFED9AC6A851">
<emphasis id="82CC47FC96627789DF2D70D1C08FFAE5" bold="true">Male lure.</emphasis>
Zingerone.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E36BI" type="host">
<paragraph id="901E4F6CCBC2023DBD11F9DB0B0584FB">
<heading id="78FB5D40E007603E244984B138B7DA75" reason="title">Host plant.</heading>
</paragraph>
<paragraph id="B255D97FBE69BFA46871660830194457">Unknown.</paragraph>
</subSubSection>
<subSubSection id="SECID0EBACI" type="etymology">
<paragraph id="2ED0C1F6CE2DB148F29188521D6773FB">
<heading id="1D0B4B2DCEC3BAFB9B6BEEAFD7ED4D63" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="F9C5DAE7927710231777AF927FD4EDE1">
The species epithet
<emphasis id="1BF28A11FA8B37095B3C641182968FEB" italics="true">cataracta</emphasis>
, Latin for waterfall, is a noun in apposition that refers to both the peculiar wing pattern that resembles a waterfall going over the edge and the
<typeStatus id="B92D3D2418B9A0E79D8667D61C995C03">type</typeStatus>
locality; the Tembaling waterfall trail in the Danum Valley Conservation Area.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERACI" type="comments">
<paragraph id="E9DCC73A0124032312AD47329A43407F">
<heading id="6753CBBAD25228ADF2451A29445DC18E" reason="title">Comments.</heading>
</paragraph>
<paragraph id="FD10CF0BC36AA39A00AB11E2EA2468BD">
This species was included as
<taxonomicName id="FF61B92B8B83B3E9B59F2FD4065D5985" class="Insecta" family="Tephritidae" genus="Zeugodacus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">
<emphasis id="43806402FE8EE21116F136BA1330E38A" italics="true">Zeugodacus</emphasis>
</taxonomicName>
spnMalaysia 02 in the DNA barcoding study of
<bibRefCitation id="E783DF945C6FD1E1408663ECACC59D35" DOI="10.1111/1755-0998.13987" author="Doorenweerd C &amp; San Jose M &amp; Leblanc L &amp; Barr N &amp; Geib SM &amp; Chung AYC &amp; Dupuis JR &amp; Ekayanti A &amp; Fiegalan E &amp; Hemachandra KS &amp; Hossain MA &amp; Huang C &amp; Hsu Y &amp; Morris KY &amp; Maryani A &amp; Mustapeng A &amp; Niogret J &amp; Pham TH &amp; Thi Nguyen N &amp; Sirisena UGAI &amp; Todd T &amp; Rubinoff D" refId="B15" refString="Doorenweerd C, San Jose M, Leblanc L, Barr N, Geib SM, Chung AYC, Dupuis JR, Ekayanti A, Fiegalan E, Hemachandra KS, Hossain MA, Huang C, Hsu Y, Morris KY, Maryani A, Mustapeng A, Niogret J, Pham TH, Thi Nguyen N, Sirisena UGAI, Todd T, Rubinoff D (2024 b) Towards a better future for DNA barcoding: Evaluating monophyly- and distance-based species identification using COI gene fragments of Dacini fruit flies. Molecular Ecology Resources 13987. https://doi.org/10.1111/1755-0998.13987">Doorenweerd et al. (2024 b</bibRefCitation>
). The combination of a short cell
<emphasis id="2C974794A5C06113C5536E0E52DDCEA7" italics="true">cup</emphasis>
and antennae shorter than the height of the head makes accurate systematic placement of this species challenging. Within
<taxonomicName id="E93F80B7088668021A2784827069F38F" class="Insecta" family="Tephritidae" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="tribe" tribe="Dacini">Dacini</taxonomicName>
, this combination is only known in the monotypic
<emphasis id="91B2CF7C7BC08D7E32F86FB69BF6E528" italics="true">
<taxonomicName id="8A76D2C2A256252732B09A8BAD1D62AD" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">Bactrocera</taxonomicName>
subgenus Queenslandacus
</emphasis>
;
<taxonomicName id="2C983E8E73ECC969C309B5D716929468" baseAuthorityName="May" baseAuthorityYear="1958" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="exigua">
<emphasis id="D38F36EDC289F5D0D30A4FB2A12D6D51" italics="true">B. exigua</emphasis>
</taxonomicName>
(May), known from
<collectingCountry id="5A7E59790F5BB641F600AB12700F9800" name="Australia">Australia</collectingCountry>
. However, the yellow presutural markings and long posterior lobes of the male surstylus clearly place this species in
<taxonomicName id="3364D147DF8F524C159E4182709712E8" class="Insecta" family="Tephritidae" genus="Zeugodacus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">
<emphasis id="96905972F7C0DE7F8DEC71A138186CFA" italics="true">Zeugodacus</emphasis>
</taxonomicName>
, and we find placement in subgenus
<taxonomicName id="7BB3081EB10C4E631E401ABCD2098E7E" rank="subGenus" subGenus="Zeugodacus">
<emphasis id="731C216F93EE15397EDEDEC78B7DB334" italics="true">Zeugodacus</emphasis>
</taxonomicName>
best fitting and consider the similarity with
<taxonomicName id="0691411E920455D4E6B4029B3C5FFA93" baseAuthorityName="May" baseAuthorityYear="1958" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="exigua">
<emphasis id="4FBF6B4A7135BC6BB81617AFA6C2618B" italics="true">B. exigua</emphasis>
</taxonomicName>
paralogous.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,296 @@
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<mods:title id="160A34E9788F34F82196505CC110A38D">The Dacini fruit flies of Borneo: an annotated checklist with 89 species including three new to science (Tephritidae, Dacinae)</mods:title>
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<mods:affiliation id="4D19025650143CA272AA1CC5288CF0D2">Sabah Forestry Department, Forest Research Centre, P. O. Box 1407, 90715 Sandakan, Sabah, Malaysia</mods:affiliation>
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<subSubSection id="C2481AEC54B4B948279680DADE4C8E67" type="nomenclature">
<paragraph id="23DF4FB892E7913C81EBACF319A3CF26">
<taxonomicName id="F622F743299B95ADD87A058B8F83A84C" authority="Doorenweerd" authorityName="Doorenweerd &amp; Chung &amp; Mustapeng &amp; Rubinoff" authorityYear="2025" class="Insecta" family="Tephritidae" genus="Dacus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="danumensis" status="sp. nov." subGenus="Mellesis">
<emphasis id="E9475A2E57D7721710B20DDA1251A6D4" italics="true">Dacus</emphasis>
(
<emphasis id="3FDCD5F116F471F5A6FAE77E61C74DA5" italics="true">Mellesis</emphasis>
)
<emphasis id="0468C54184FC28F675844E904F6975DB" italics="true">danumensis</emphasis>
Doorenweerd
</taxonomicName>
<taxonomicNameLabel id="8DD1A4FBA56F3446AF676A37C04EB9CF" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="B8329D5542B50F32B95316ED5024A1B0" type="description">
<paragraph id="D528CE7FCE34430E7C326E3D833541CC">
<figureCitation id="63C4BAE165A98DC04E7FF7A8F9B4D52B" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Holotype male of Dacus danumensis sp. n., UHIM. ms 08931 A dorsal view B anterior view of the head C lateral view D dorso-anterior view of the abdomen showing the ceromata E right wing, dissected and slide mounted F detail image of the genitalia showing the surstylus length." figureDoi="10.3897/zookeys.1240.148768.figure4" httpUri="https://binary.pensoft.net/fig/1348696">Fig. 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EKPBI" type="type material">
<paragraph id="151C5242D2A33D1B42E7CFB580246751">
<heading id="D4211F3D564B5E445237A870A5B2829B" reason="title">Type material.</heading>
</paragraph>
<paragraph id="3A0AC8D9D01D4E46A17B220FADC1E15B">
<materialsCitation id="97898D0E4D13027B686593C2C2DE3D2A" collectingDate="2018-12-03" collectingDateMax="2018-12-05" collectingDateMin="2018-12-03" collectionCode="UHIM" collectorName="Leg. D. Rubinoff &amp; C. Doorenweerd." country="Malaysia" latitude="4.9534" location="University of Hawaii Insect Museum" longLatPrecision="7" longitude="117.8062" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<emphasis id="DF4FC51916B267A67E7133BAEFAA8D62" bold="true">
<emphasis id="D1EEE32A1A17E4B659021A668A55E780" bold="true" italics="true">
<typeStatus id="AA3B90120EBA183057B810D1C3DB7C89" type="holotype">Holotype</typeStatus>
</emphasis>
.
</emphasis>
Male. “
<collectingCountry id="923581703CF32A29000DD3C8550AFC4E" name="Malaysia">Malaysia</collectingCountry>
:
<collectingRegion id="FB40D50719BC805AD3CDA8FF7F2A9FE0" country="Malaysia" name="Sabah">Sabah</collectingRegion>
: Danum valley Tembaling waterfall trail. WGS 84
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EYPBI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[117.806200,4.953400]}">
<geoCoordinate id="30A45CAA9E225FA604BFDBE6D395514D" degrees="4.9534" direction="north" orientation="latitude" precision="5" value="4.9534">4.9534</geoCoordinate>
,
<geoCoordinate id="13BDA1B290495FA11086A4108701645C" degrees="117.8062" direction="east" orientation="longitude" precision="5" value="117.8062">117.8062</geoCoordinate>
</named-content>
<date id="9AE19BFDC064BFFC7079495B4E3E13D7" value="2018-12-03" valueMax="2018-12-05" valueMin="2018-12-03">
<collectingDate id="BE92DCD69B995082D71BD2A38270E570" value="2018-12-03" valueMax="2018-12-05" valueMin="2018-12-03">35. xii. 2018</collectingDate>
</date>
<collectingMethod id="B55BD5B2C83296C491201ADDC0F65A8E">Zingerone trap</collectingMethod>
.
<collectorName id="DA3CD05958B1640C50CE46C0DA2E9FA3">Leg. D. Rubinoff</collectorName>
&amp;
<collectorName id="9208A4509E62A160446D3D16B848F5D3">C. Doorenweerd.</collectorName>
DNA sample ms 08931. ” Deposited at the
<location id="1C7D92358036E5F7A23A0407202633B3" LSID="urn:lsid:plazi:treatment:3A334A530B295861BFAEBFA058163E70:1C7D92358036E5F7A23A0407202633B3" country="Malaysia" latitude="4.9534" longLatPrecision="7" longitude="117.8062" name="University of Hawaii Insect Museum" stateProvince="Sabah">University of Hawaii Insect Museum</location>
reg. no.
<abbrev content-type="institution" id="ABBRID0E4PBI" xlink_title="University of Hawaiʻi at Mānoa">
<collectionCode id="336591434BDDB242208FFA4F32C3067C">UHIM</collectionCode>
</abbrev>
. ms 08931.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECQBI" type="Differential diagnosis">
<paragraph id="DDAAF1FE1B9F8E83406B8468A78CB88B">
<heading id="CF5AB2B672C6554D3A0F957EFFCCB3D9" reason="title">Differential diagnosis.</heading>
</paragraph>
<paragraph id="CDA0A0703310F4D765F04D4BF1A8A510">
The combination of an elongate and petiolate ( wasp-like) abdomen, a uniquely shaped postsutural medial yellow vitta that is almost equally wide as long, the wing with a costal band that reaches up to vein
<collectionCode id="6FD686BF86C0D9D61D782D993414B075" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
<subScript id="19A052AF6682B22E17004E40DF174B6D">4 + 5</subScript>
and does not expand apically, and the absence of an anal streak, is unique within
<taxonomicName id="A7DEA1B74526C5993248ACBD13105F75" class="Insecta" family="Tephritidae" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="tribe" tribe="Dacini">Dacini</taxonomicName>
. The most closely related species based on
<abbrev id="ABBRID0EPQBI" xlink_title="Cytochrome C Oxidase I">COI</abbrev>
sequence data is
<taxonomicName id="656362F9C1372B12F3B057A8A1DDC39D" authority="Wang" authorityName="Wang" class="Insecta" family="Tephritidae" genus="Dacus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="EB05D2E24A93EE955EA4CB96AE63AF7D" italics="true">Dacus sinensis</emphasis>
Wang
</taxonomicName>
, from which
<taxonomicName id="BAE3E0E6324E72EE09033BD3FEEFF68A" authorityName="Doorenweerd &amp; Chung &amp; Mustapeng &amp; Rubinoff" authorityYear="2025" class="Insecta" family="Tephritidae" genus="Dacus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="danumensis">
<emphasis id="6F8EF741D465E2CC4375C0D00565FA14" italics="true">D. danumensis</emphasis>
</taxonomicName>
can be morphologically distinguished by having a medial postsutural yellow vitta, which is absent in
<taxonomicName id="B8E40D3257ABF049FD43D599CDB72B50" class="Insecta" family="Tephritidae" genus="Dacus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="0EFB699E58BECE6BA99826763FE2DB91" italics="true">D. sinensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EURBI" type="Molecular diagnostics">
<paragraph id="8B6F28E7AD58E7C1148D223B3640C1EB">
<heading id="DB4C215565C3E3ED37C56BFCA5D84F4F" reason="title">Molecular diagnostics.</heading>
</paragraph>
<paragraph id="AB72F1A810CD9388CEB3A4F396D34C80">
<taxonomicName id="A58A0C1A38603997F46C97E3BFEB5FAD" authority="COI" authorityName="COI" class="Insecta" family="Tephritidae" genus="Dacus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="danumensis">
<emphasis id="12F9BB693F25BAB346FD0F17DE1DC795" italics="true">Dacus danumensis</emphasis>
<abbrev id="ABBRID0EFSBI" xlink_title="Cytochrome C Oxidase I">COI</abbrev>
</taxonomicName>
- 5 P 3 P sequences are most similar to sequences of
<taxonomicName id="FE86A7F29E877A3C5734115F6E4C56AA" class="Insecta" family="Tephritidae" genus="Dacus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="EEEE78F3FDEE30B7769740AEDE61A42F" italics="true">D. sinensis</emphasis>
</taxonomicName>
at 8.6 % minimum pairwise difference (
<bibRefCitation id="B867137882F76DBCC3179D549789F17D" DOI="10.1111/1755-0998.13987" author="Doorenweerd C &amp; San Jose M &amp; Leblanc L &amp; Barr N &amp; Geib SM &amp; Chung AYC &amp; Dupuis JR &amp; Ekayanti A &amp; Fiegalan E &amp; Hemachandra KS &amp; Hossain MA &amp; Huang C &amp; Hsu Y &amp; Morris KY &amp; Maryani A &amp; Mustapeng A &amp; Niogret J &amp; Pham TH &amp; Thi Nguyen N &amp; Sirisena UGAI &amp; Todd T &amp; Rubinoff D" refId="B15" refString="Doorenweerd C, San Jose M, Leblanc L, Barr N, Geib SM, Chung AYC, Dupuis JR, Ekayanti A, Fiegalan E, Hemachandra KS, Hossain MA, Huang C, Hsu Y, Morris KY, Maryani A, Mustapeng A, Niogret J, Pham TH, Thi Nguyen N, Sirisena UGAI, Todd T, Rubinoff D (2024 b) Towards a better future for DNA barcoding: Evaluating monophyly- and distance-based species identification using COI gene fragments of Dacini fruit flies. Molecular Ecology Resources 13987. https://doi.org/10.1111/1755-0998.13987">Doorenweerd et al. 2024 b</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EYSBI" type="description">
<paragraph id="FA1AB05630309F3801298537D5CB7A81">
<heading id="C30E1A2A882F9D796DFEC1991CCC1715" reason="title">Description.</heading>
</paragraph>
<paragraph id="E53E9DCA0452E508534E535DD54E630A">
<emphasis id="A511CAE040A2339BF05A5441F6E487A8" bold="true">
Male.
<emphasis id="982799E7DB6DA75287CEEB18D2629F55" bold="true" italics="true">Head</emphasis>
</emphasis>
(Fig.
<figureCitation id="706B7FC936A7AFC0F240B2451F43363D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Holotype male of Dacus danumensis sp. n., UHIM. ms 08931 A dorsal view B anterior view of the head C lateral view D dorso-anterior view of the abdomen showing the ceromata E right wing, dissected and slide mounted F detail image of the genitalia showing the surstylus length." figureDoi="10.3897/zookeys.1240.148768.figure4" httpUri="https://binary.pensoft.net/fig/1348696">4 B</figureCitation>
). Fulvous with two waterdrop-shaped black facial spots in the antennal furrows and a black band connecting both compound eyes across the occiput. Antennae dark fulvous. Combined length of antennal segments greater than the height of the head
<emphasis id="89B98A6BCC34DCCAAF5F7C6085EC16FB" bold="true" italics="true">Thorax</emphasis>
(Fig.
<figureCitation id="F0DFD81708D3BD50DD9683512671D67C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Holotype male of Dacus danumensis sp. n., UHIM. ms 08931 A dorsal view B anterior view of the head C lateral view D dorso-anterior view of the abdomen showing the ceromata E right wing, dissected and slide mounted F detail image of the genitalia showing the surstylus length." figureDoi="10.3897/zookeys.1240.148768.figure4" httpUri="https://binary.pensoft.net/fig/1348696">4 A, C</figureCitation>
). Scutum and pleural areas completely black. Yellow markings: postpronotal lobes; notopleura and adjacent presutural area; medial postsutural roughly waterdrop-shaped vitta; mesopleural stripe approx. the width of the notopleuron, anterior margin slightly convex; anatergite, katatergite; scutellum. Setae: one pair scutellar; prescutellar setae absent; one pair intra-alar; one pair posterior supra-alar; one pair anterior supra-alar; two pair notopleural; four scapular.
<emphasis id="B9F2E45474ABDFAC526914D3EFFF048C" bold="true" italics="true">Abdomen</emphasis>
(Fig.
<figureCitation id="202946FB034C1671F5FD56208A7D1E29" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Holotype male of Dacus danumensis sp. n., UHIM. ms 08931 A dorsal view B anterior view of the head C lateral view D dorso-anterior view of the abdomen showing the ceromata E right wing, dissected and slide mounted F detail image of the genitalia showing the surstylus length." figureDoi="10.3897/zookeys.1240.148768.figure4" httpUri="https://binary.pensoft.net/fig/1348696">4 A, C, D, F</figureCitation>
). Abdomen shape petiolate with narrow and elongated segment I; terga fused; pecten present on tergum III; posterior lobe of the male surstylus short (Fig.
<figureCitation id="681A6F019A37F2337F0F12A148402B02" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Holotype male of Dacus danumensis sp. n., UHIM. ms 08931 A dorsal view B anterior view of the head C lateral view D dorso-anterior view of the abdomen showing the ceromata E right wing, dissected and slide mounted F detail image of the genitalia showing the surstylus length." figureDoi="10.3897/zookeys.1240.148768.figure4" httpUri="https://binary.pensoft.net/fig/1348696">4 F</figureCitation>
); abdominal sternum
<collectionCode id="FFA5D602D48896F600470EF253C8EEFC" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
with a narrow concavity on the posterior margin. Tergum I black with yellow posterior band; tergum II yellow with a black T-shape; tergum III with a second black T-shape that covers most of the segment and a medial black line that continuous from tergum II down to the posterior of tergum
<collectionCode id="CCD2EA35C5059EA0E23FEC0860C817C2" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
; lateral black bands on tergum
<collectionCode id="7241FA994ED9112D06E48DC1967C75A3">IV</collectionCode>
and
<collectionCode id="6A9B233DD03F75BE3FC5CE0727E43975" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
. Ceromata yellow and indistinct.
<emphasis id="B7B0E6429865EA1B1C5B1A1ADF040705" bold="true" italics="true">Legs</emphasis>
(Fig.
<figureCitation id="0C35606648C1FB71347CA1207D46EF3E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Holotype male of Dacus danumensis sp. n., UHIM. ms 08931 A dorsal view B anterior view of the head C lateral view D dorso-anterior view of the abdomen showing the ceromata E right wing, dissected and slide mounted F detail image of the genitalia showing the surstylus length." figureDoi="10.3897/zookeys.1240.148768.figure4" httpUri="https://binary.pensoft.net/fig/1348696">4 C</figureCitation>
). Fore tibia with dark mark on the outer surface that covers most of the tibial length; mid tibia with dark marks on the outer and inner surface that extend from one-third the distal end; hind tibia with distal quarter black. All other leg segments yellow to fulvous.
<emphasis id="3CC8FF3810CD6276EBA01D21D5A2AC37" bold="true" italics="true">Wings</emphasis>
(Fig.
<figureCitation id="780D93E405F81855D535ADC6D407E99E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Holotype male of Dacus danumensis sp. n., UHIM. ms 08931 A dorsal view B anterior view of the head C lateral view D dorso-anterior view of the abdomen showing the ceromata E right wing, dissected and slide mounted F detail image of the genitalia showing the surstylus length." figureDoi="10.3897/zookeys.1240.148768.figure4" httpUri="https://binary.pensoft.net/fig/1348696">4 E</figureCitation>
). Wing length
<quantity id="66E2A8C05460301E3882F27C80BE0157" metricMagnitude="-3" metricUnit="m" metricValue="4.9" unit="mm" value="4.9">4.9 mm</quantity>
. Cells
<emphasis id="863F6987D2F47B1F9F87AAD619F72D8A" italics="true">bc</emphasis>
and
<emphasis id="CF06CF74298E5A356A7A9299721216EB" italics="true">c</emphasis>
clear, faint tint in cell
<emphasis id="7CBDC6EEBE78A9D7AA8F4E8CCBE33F51" italics="true">sc.</emphasis>
Costal band reaches up to vein
<collectionCode id="2DFCAAAD51A08A892C84F5907FA71A19" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
<subScript id="1296E9DCB9716FCEBB13A8D94B1CE520">4 + 5</subScript>
, slightly crosses
<collectionCode id="C12850B308685A2BE01B1BE658EBBC11" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
<subScript id="E8F493C0DAAF89E9E8430254845EBAB8">4 + 5</subScript>
apically. Anal streak absent, supernumerary lobe not pronounced.
<emphasis id="538CD12023D18DCAC1782F6FE3DB513A" bold="true">Female.</emphasis>
Unknown.
</paragraph>
<caption id="CB121375F67F085C46BC92FCC14B238E" ID-DOI="10.3897/zookeys.1240.148768.figure4" ID-arpha="2E077206-16F6-5244-8C1B-69B4CE27F9F7" httpUri="https://binary.pensoft.net/fig/1348696" startId="F4">
<paragraph id="84BFC941AE7F9E43A2F43FA959FCF839">
<label id="4081C63C00B02125EE6BA23E54AA552F">Figure 4.</label>
</paragraph>
<paragraph id="FFFA0FC61BBA2A5F17B6C176A3B1E549">
Holotype male of
<taxonomicName id="4E13CAF3B558601559224ABD7A68A54F" authorityName="Doorenweerd &amp; Chung &amp; Mustapeng &amp; Rubinoff" authorityYear="2025" class="Insecta" family="Tephritidae" genus="Dacus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="danumensis" status="sp. nov.">
<emphasis id="0726CBECEBF46069A270837C92752162" italics="true">Dacus danumensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="502C838FA55981A87C79AE0345910A5E" rank="species">sp. n.</taxonomicNameLabel>
,
<abbrev content-type="institution" id="ABBRID0EFVBI" xlink_title="University of Hawaiʻi at Mānoa">UHIM</abbrev>
. ms 08931
<emphasis id="480A482C3B1C3080CFA721E8835D0936" bold="true">A</emphasis>
dorsal view
<emphasis id="18F8D5386B581A6CD118143A08928A49" bold="true">B</emphasis>
anterior view of the head
<emphasis id="BB5F9F0F2924B040F0C889F9C20A6F50" bold="true">C</emphasis>
lateral view
<emphasis id="EEEE1BB6B724B573CB474C7096CE6568" bold="true">D</emphasis>
dorso-anterior view of the abdomen showing the ceromata
<emphasis id="DD229C594B8D6A58F440D5D80210F27D" bold="true">E</emphasis>
right wing, dissected and slide mounted
<emphasis id="57DDB7D241910E2CEE634AB6C27F66F1" bold="true">F</emphasis>
detail image of the genitalia showing the surstylus length.
</paragraph>
</caption>
<paragraph id="1C6EFC1964D9DEDD66B5D2FA607A3EEA">
<emphasis id="AFC7489A3D08D6EE421AC24AD88A3686" bold="true">Male lure.</emphasis>
Zingerone.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E6VBI" type="host">
<paragraph id="6E8DD9435D42BEAECFFF2D60AAAC7E46">
<heading id="C28F79AE982FEA7C21DB0A70C366AB58" reason="title">Host plant.</heading>
</paragraph>
<paragraph id="ECACDFEDB8FF23BDA23EFBFCE8A8D754">Unknown.</paragraph>
</subSubSection>
<subSubSection id="SECID0EEWBI" type="etymology">
<paragraph id="4EB91046A0DAE4A75363291ADB4A67E1">
<heading id="0822656EC360A66FACBB577B3291D9B1" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="9523FAE20A21E4EEE65550A9BF38C307">
The species epithet
<emphasis id="F690489E4E2AFFE66597ECA813DEAF34" italics="true">danumensis</emphasis>
is a noun in apposition that refers to the
<typeStatus id="F1445073EBFBA9D9ACC35A02D0CC9384">type</typeStatus>
locality; the Danum Valley Conservation Area in
<collectingRegion id="F6A043B27FCFEA66D62415AF969F5183" country="Malaysia" name="Sabah">Sabah</collectingRegion>
,
<collectingCountry id="28EF5812ABD5CDDFCEB691D4315FDCD1" name="Malaysia">Malaysia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUWBI" type="comments">
<paragraph id="174FA47FD52878CB284B0040437E4A3A">
<heading id="2A74EA88384573D166445E7BAC4B9D61" reason="title">Comments.</heading>
</paragraph>
<paragraph id="1E11536F0757E3A75F8B9EA8AD5679DA">
This species was included as
<taxonomicName id="C4E5753579D1A0C10BF66485B0FA4797" class="Insecta" family="Tephritidae" genus="Dacus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">
<emphasis id="50A558C14CDC9E519C863FC57B0E66A1" italics="true">Dacus</emphasis>
</taxonomicName>
spnMalaysia 01 in the DNA barcoding study of
<bibRefCitation id="70A9A5856C70E3FEDAD5C7D641D5E6A1" DOI="10.1111/1755-0998.13987" author="Doorenweerd C &amp; San Jose M &amp; Leblanc L &amp; Barr N &amp; Geib SM &amp; Chung AYC &amp; Dupuis JR &amp; Ekayanti A &amp; Fiegalan E &amp; Hemachandra KS &amp; Hossain MA &amp; Huang C &amp; Hsu Y &amp; Morris KY &amp; Maryani A &amp; Mustapeng A &amp; Niogret J &amp; Pham TH &amp; Thi Nguyen N &amp; Sirisena UGAI &amp; Todd T &amp; Rubinoff D" refId="B15" refString="Doorenweerd C, San Jose M, Leblanc L, Barr N, Geib SM, Chung AYC, Dupuis JR, Ekayanti A, Fiegalan E, Hemachandra KS, Hossain MA, Huang C, Hsu Y, Morris KY, Maryani A, Mustapeng A, Niogret J, Pham TH, Thi Nguyen N, Sirisena UGAI, Todd T, Rubinoff D (2024 b) Towards a better future for DNA barcoding: Evaluating monophyly- and distance-based species identification using COI gene fragments of Dacini fruit flies. Molecular Ecology Resources 13987. https://doi.org/10.1111/1755-0998.13987">Doorenweerd et al. (2024 b</bibRefCitation>
). Placement in subgenus
<taxonomicName id="2C63210924A33CA165BB4E5F10FEA9B1" rank="subGenus" subGenus="Mellesis">
<emphasis id="21591F8C9DA835FCF26AD79A4FCA0F59" italics="true">Mellesis</emphasis>
</taxonomicName>
is based on lacking prescutellar setae, a slight concavity on the posterior margin of sternum
<collectionCode id="A869396CF987102707DE5E01AE15324F" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
and its elongate and petiolate abdomen shape.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="922FAB0021F8C41CD430C356B4E742AC" ID-DOI="10.3897/zookeys.1240.148768" ID-ZooBank="DC035B97-5EF7-4FBB-8589-36C59208E745" ID-publisher-id="148768" URI-arpha="483F65DB-D8A4-5E3A-BE2A-8EAAF6DB1564" URI-zoobank="http://zoobank.org/DC035B97-5EF7-4FBB-8589-36C59208E745" article-type="research-article" checkinTime="1749481029109" checkinUser="pensoft" docAuthor="Doorenweerd, Camiel, Chung, Arthur Y. C., Mustapeng, Andi Maryani A. &amp; Rubinoff, Daniel" docDate="2025" docId="47665AA36DEA54A9A9D09D2210832D13" docLanguage="en" docName="ZooKeys 1240: 305-325" docOrigin="ZooKeys 1240" docSource="https://zookeys.pensoft.net/article/148768/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Bactrocera (Bactrocera) melanobivittata Doorenweerd &amp; Chung &amp; Mustapeng &amp; Rubinoff 2025, sp. nov." docType="treatment" docVersion="1" dtd-version="3.0" lastPageNumber="325" masterDocId="483F65DBD8A45E3ABE2A8EAAF6DB1564" masterDocTitle="The Dacini fruit flies of Borneo: an annotated checklist with 89 species including three new to science (Tephritidae, Dacinae)" masterLastPageNumber="325" masterPageNumber="305" pageNumber="305" updateTime="1749481098078" updateUser="pensoft">
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<mods:title id="19A61730CFD64D637A3BCE3866F5C5E8">The Dacini fruit flies of Borneo: an annotated checklist with 89 species including three new to science (Tephritidae, Dacinae)</mods:title>
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<mods:namePart id="A827BF305EBF05FF4511E3E08239ED8B">Doorenweerd, Camiel</mods:namePart>
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<mods:affiliation id="F59808445B713463C17F7B882196BC97">Department of Plant and Environmental Protection Sciences, Entomology Section, College of Tropical Agriculture and Human Resources, University of Hawaiʻi at Mānoa, 3050 Maile Way, Honolulu, Hawaii, 96822 - 2231, USA</mods:affiliation>
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<mods:namePart id="C1BAC6A903661545C2910B88E6E82221">Chung, Arthur Y. C.</mods:namePart>
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<mods:affiliation id="F8AC484E43DF5129272B00D1157B10BB">Sabah Forestry Department, Forest Research Centre, P. O. Box 1407, 90715 Sandakan, Sabah, Malaysia</mods:affiliation>
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<mods:namePart id="1243C6C3A788BAAE52049F93A1DA1FF0">Mustapeng, Andi Maryani A.</mods:namePart>
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<mods:affiliation id="EE247A4F9FFCFD950B2472D1CDD6C1C7">Sabah Forestry Department, Forest Research Centre, P. O. Box 1407, 90715 Sandakan, Sabah, Malaysia</mods:affiliation>
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<mods:namePart id="6DA2EBBF6EC4190AEA64FCB671B70796">Rubinoff, Daniel</mods:namePart>
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<mods:affiliation id="7072753D4F75DA710C5671AC938411AD">Department of Plant and Environmental Protection Sciences, Entomology Section, College of Tropical Agriculture and Human Resources, University of Hawaiʻi at Mānoa, 3050 Maile Way, Honolulu, Hawaii, 96822 - 2231, USA</mods:affiliation>
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<subSubSection id="45DFF275DBAD434C6CB3A8C55618194A" type="nomenclature">
<paragraph id="EE87B06053CAD42041B2B7D398E5D507">
<taxonomicName id="4AEE0A70D8DB5E987FA8B01E962ABC49" authority="Doorenweerd" authorityName="Doorenweerd &amp; Chung &amp; Mustapeng &amp; Rubinoff" authorityYear="2025" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="melanobivittata" status="sp. nov." subGenus="Bactrocera">
<emphasis id="C376D8AE29BEE61278974E12CC1A4739" italics="true">Bactrocera</emphasis>
(
<emphasis id="4DEB2E60475A2AC6CF7CF4AD641CE8E2" italics="true">Bactrocera</emphasis>
)
<emphasis id="82F2370EE7652C0E9F73160B87E4E223" italics="true">melanobivittata</emphasis>
Doorenweerd
</taxonomicName>
<taxonomicNameLabel id="B8214601C38F0A640AE7D7643B41BC06" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="94591AF7A8F2C365509F90EFF86C2C26" type="description">
<paragraph id="4DE3C1063E379EA4DD74ED940CD030C2">
<figureCitation id="2515AB219AF75986CB838624AC540EA9" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Holotype male of Bactrocera melanobivittata sp. nov., UHIM. ms 12171 A dorsal view of head and thorax B dorsal view of abdomen C anterior view of the head D lateral view of habitus E posterior view of abdomen showing the ceromata F left wing, dissected and slide mounted G detail image of the genitalia showing the surstylus length." figureDoi="10.3897/zookeys.1240.148768.figure2" httpUri="https://binary.pensoft.net/fig/1348694">Fig. 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E46AI" type="type material">
<paragraph id="17B0F2C1F857DB4C9A923E772CB7D66E">
<heading id="F96F3B9FF403D83B64B84780E0E9D500" reason="title">Type material.</heading>
</paragraph>
<paragraph id="99D42F71D147CE0710576C31442B924B">
<materialsCitation id="0D0FCDFA15D5BC3A4A8725583E2D9A0C" collectingDate="2018-12-03" collectingDateMax="2018-12-05" collectingDateMin="2018-12-03" collectionCode="UHIM" collectorName="Leg. D. Rubinoff &amp; C. Doorenweerd." country="Malaysia" location="University of Hawaii Insect Museum" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<emphasis id="C59A93E399109A0ECCFBF214E5C4F755" bold="true">
<emphasis id="C947C5A8ADF8D9A943C2A557166B2B14" bold="true" italics="true">
<typeStatus id="D43134E0F2602E9E323516C45C1299C7" type="holotype">Holotype</typeStatus>
</emphasis>
.
</emphasis>
Male. “
<collectingCountry id="5E61DFBC2FB34865677CAD1AF5C7CB24" name="Malaysia">Malaysia</collectingCountry>
:
<collectingRegion id="7EC8C9A74EF8FC98F1006EB2EB1EDB81" country="Malaysia" name="Sabah">Sabah</collectingRegion>
.
<date id="A773AA419F787A370767CC50601182CF" value="2018-12-03" valueMax="2018-12-05" valueMin="2018-12-03">
<collectingDate id="82DC674F88A54F8A3BCC56E44D3B94CA" value="2018-12-03" valueMax="2018-12-05" valueMin="2018-12-03">35. xii. 2018</collectingDate>
</date>
. Danum Valley access rd.
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ELABI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[117.817300,4.968300]}">4.9683 &quot; N, 117.8173 &quot; E</named-content>
.
<collectingMethod id="89997DC4E54C2BC14CCDEAEB82031AC6">Methyl eugenol trap</collectingMethod>
FF 18 Ma 065.
<collectorName id="F918739553A2DEFF53DAF57D6393341B">Leg. D. Rubinoff</collectorName>
&amp;
<collectorName id="67A12A8963DF5C5517326075C5BAD980">C. Doorenweerd.</collectorName>
DNA sample ms 12171 ”. Deposited at the
<location id="CB71E689BB02294C418EDFEBE766BE42" LSID="urn:lsid:plazi:treatment:47665AA36DEA54A9A9D09D2210832D13:CB71E689BB02294C418EDFEBE766BE42" country="Malaysia" name="University of Hawaii Insect Museum" stateProvince="Sabah">University of Hawaii Insect Museum</location>
reg. no.
<abbrev content-type="institution" id="ABBRID0EQABI" xlink_title="University of Hawaiʻi at Mānoa">
<collectionCode id="08D2DA3807DF93C077D1AD69A4EC7812">UHIM</collectionCode>
</abbrev>
. ms 12171
</materialsCitation>
.
<emphasis id="8C33E173D72F2702B928CBCA19B1F293" bold="true" italics="true">
<typeStatus id="3A6370549E17641FE328B015FB772D00" type="paratype">Paratype</typeStatus>
</emphasis>
.
<specimenCount id="00B7FAB249EBC057B77E6AB8616E5E0B" count="1" type="male">One male</specimenCount>
. “
<collectingCountry id="CD9223910DE76CE76711916D68EA828F" name="Malaysia">Malaysia</collectingCountry>
,
<collectingRegion id="2F685EB6DDB85BC03422B0A6716A2FD1" country="Malaysia" name="Sabah">Sabah</collectingRegion>
.
<date id="7DB7EF62C445E8F29038C99B3BD931CC" value="2019-08-02" valueMax="2019-08-04" valueMin="2019-08-02">24. viii. 2019</date>
. Maliau Basin: camp Agathis trail.
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E2ABI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[116.894700,4.707900]}">4.7079 &quot; N, 116.8947 &quot; E</named-content>
. Methyl eugenol trap FF 19 Ma 036. Leg. D. Rubinoff &amp; C. Doorenweerd. DNA sample ms 08978. ” Deposited at the
<collectingRegion id="62DAD7F34B3E58A104F7464719C293C8" country="Malaysia" name="Sabah">Sabah</collectingRegion>
Forestry Department Insect Collection, reg. no.
<abbrev content-type="institution" id="ABBRID0EABBI" xlink_title="University of Hawaiʻi at Mānoa">
<collectionCode id="7A7EF81901789A0D9B5AFBC222947BEB">UHIM</collectionCode>
</abbrev>
. ms 08978.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EFBBI" type="Differential diagnosis">
<paragraph id="033150CE75A6729E6F1ED2ECE93E65B1">
<heading id="0DAE2AB261DC6DE2BE1305913EDF7558" reason="title">Differential diagnosis.</heading>
</paragraph>
<paragraph id="8CA336BA7F1CB53C9D273231A0B215A3">
The two lateral longitudinal red stripes on the black scutum distinguish
<taxonomicName id="ABC681E86538FE0CC1F5BACCAC0A72BD" authorityName="Doorenweerd &amp; Chung &amp; Mustapeng &amp; Rubinoff" authorityYear="2025" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="melanobivittata">
<emphasis id="0DC25FBF83D813D19338EB9869C6DEC8" italics="true">B. melanobivittata</emphasis>
</taxonomicName>
from most other
<taxonomicName id="5B3D94376ABE83641AD4ACE6C0908678" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">
<emphasis id="B038339528C635FC8E13EFF75769F8BC" italics="true">Bactrocera</emphasis>
</taxonomicName>
. The most similar species is
<taxonomicName id="6A034D29C77C264AA09B699D33CB5621" authority="Lin &amp; Wang" authorityName="Lin &amp; Wang" authorityYear="2005" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="bivittata">
<emphasis id="3C2B185B3FA7EC546E908D13CD125926" italics="true">B. bivittata</emphasis>
Lin &amp; Wang
</taxonomicName>
, but
<taxonomicName id="E87A1CB6E7BB9E0BB5E1FA2B0A5EA5E5" authorityName="Doorenweerd &amp; Chung &amp; Mustapeng &amp; Rubinoff" authorityYear="2025" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="melanobivittata">
<emphasis id="7B358158B9122E8BB0A976EB5869EBE5" italics="true">B. melanobivittata</emphasis>
</taxonomicName>
can be distinguished by having a scutum that is almost completely black except for two longitudinal red stripes, and the yellow postsutural vittae. There is intraspecific variation in the amount of black on the scutum in
<taxonomicName id="BA3231B5C222A43486864800B2BDD98D" authorityName="Lin &amp; Wang" authorityYear="2005" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="bivittata">
<emphasis id="3C0A97D3CB95E86880C2584FD33F2743" italics="true">B. bivittata</emphasis>
</taxonomicName>
(Fig.
<figureCitation id="D09362921DB2E9C5466DC4D60CB29DA6" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Variation in dorsal scutum coloration of Bactrocera bivittata, sister species to B. melanobivittata sp. nov. Specimens from China and Laos, photographed in ethanol. University of Hawaii Insect Museum specimen identifiers, from left to right, top to bottom: ms 03606, ms 01304, ms 03607, ms 01305, ms 03604, ms 03609, ms 03608, and ms 01790." figureDoi="10.3897/zookeys.1240.148768.figure3" httpUri="https://binary.pensoft.net/fig/1348695">3</figureCitation>
) but the red stripes are obscured in darker colored specimens of
<taxonomicName id="E01294BCB9B3AF0CF52BAAC9F7D5AA68" authorityName="Lin &amp; Wang" authorityYear="2005" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="bivittata">
<emphasis id="B84E3F8EEE6643B36E65F1B5706102EE" italics="true">B. bivittata</emphasis>
</taxonomicName>
whereas they are of roughly equal width and clearly contrasting in
<taxonomicName id="D519BF804F400E77585862DDA0826B6F" authorityName="Doorenweerd &amp; Chung &amp; Mustapeng &amp; Rubinoff" authorityYear="2025" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="melanobivittata">
<emphasis id="28E2F0EC466781C270597928F665E7C3" italics="true">B. melanobivittata</emphasis>
</taxonomicName>
.
<taxonomicName id="E66C745813564D0DB69DEA048B6E9DA0" authorityName="Doorenweerd &amp; Chung &amp; Mustapeng &amp; Rubinoff" authorityYear="2025" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="melanobivittata">
<emphasis id="C802E2C7FCEB092116743936A3DE9339" italics="true">Bactrocera melanobivittata</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="2CF4C1F716EAA0C0890B9DE6BEEE524E" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="ellenriederae">
<emphasis id="997631A4A2F448480172E3CFCC192D1D" italics="true">B. ellenriederae</emphasis>
</taxonomicName>
, which also has two red stripes on the scutum, by the costal band on the wing that ends between vein
<collectionCode id="FECFC3D34176590D8E8F9F30CE4DF853" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
<subScript id="65B50894164C95E95417C524B0D543C4">4 + 5</subScript>
and M, while the costal band of
<taxonomicName id="570C81B090CCAAE29899D3DE9F771644" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="ellenriederae">
<emphasis id="206D100474A3BC870137A644F9460DDF" italics="true">B. ellenriederae</emphasis>
</taxonomicName>
reaches vein M, and
<taxonomicName id="693E07ED28A89E114DF1358AE8B425A4" authorityName="Doorenweerd &amp; Chung &amp; Mustapeng &amp; Rubinoff" authorityYear="2025" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="melanobivittata">
<emphasis id="DED156D79D3A0F745B45DA8D7747B760" italics="true">B. melanobivittata</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="0F65885B5F5E93DB995DD4F15A05527C" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="youngi">
<emphasis id="4E0A80103E6E141E67C385420349247B" italics="true">B. youngi</emphasis>
</taxonomicName>
by having only the anterior lateral corners of tergite
<collectionCode id="46B8B90D5100D707DAE839445F44AFDD">IV</collectionCode>
black, whereas
<taxonomicName id="2C5D2AEED21AE28D298212D277DC710D" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="youngi">
<emphasis id="CA95A9681E41ED151CCC023CD75E1CD3" italics="true">B. youngi</emphasis>
</taxonomicName>
has the lateral quarters of tergite
<collectionCode id="98CDAE3016A1C014494DC6E839EFC485">IV</collectionCode>
black. The most similar species in Borneo is
<taxonomicName id="5A0884ECCA6AF323C680BBD9FD700162" authorityName="Drew &amp; Hancock" authorityYear="1994" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="lateritaenia">
<emphasis id="E3BF375B7071517091CE4B8318FF15A0" italics="true">B. lateritaenia</emphasis>
</taxonomicName>
, which can be distinguished by having tapering yellow postsutural vittae, which are parallel in
<taxonomicName id="6D669BE606BFC5D67998390C03E503E8" authorityName="Doorenweerd &amp; Chung &amp; Mustapeng &amp; Rubinoff" authorityYear="2025" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="melanobivittata">
<emphasis id="EA0C3A37847C426FD88D67557AE80AE1" italics="true">B. melanobivittata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ESGBI" type="Molecular diagnostics">
<paragraph id="FF2AC32AFE3BE7F1706F1DC6BF425D53">
<heading id="77D3EA44B2BD7A81BC86D331D0EA0712" reason="title">Molecular diagnostics.</heading>
</paragraph>
<paragraph id="1A5CDDBF1995817987CB936A1DFA5D0B">
The
<abbrev id="ABBRID0EYGBI" xlink_title="Cytochrome C Oxidase I">COI</abbrev>
- 5 P 3 P sequences of ms 12171 and ms 08978 are most similar to sequences of
<taxonomicName id="D57810F9849E1738C90123983B57DCF5" authorityName="Lin &amp; Wang" authorityYear="2005" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="bivittata">
<emphasis id="7EBA25A749946910386334DD35ECE075" italics="true">Bactrocera bivittata</emphasis>
</taxonomicName>
, but at 6.5 % minimum pairwise difference (
<bibRefCitation id="CE2C64EB17CE1DD9E167A1960A58EF0E" DOI="10.1111/1755-0998.13987" author="Doorenweerd C &amp; San Jose M &amp; Leblanc L &amp; Barr N &amp; Geib SM &amp; Chung AYC &amp; Dupuis JR &amp; Ekayanti A &amp; Fiegalan E &amp; Hemachandra KS &amp; Hossain MA &amp; Huang C &amp; Hsu Y &amp; Morris KY &amp; Maryani A &amp; Mustapeng A &amp; Niogret J &amp; Pham TH &amp; Thi Nguyen N &amp; Sirisena UGAI &amp; Todd T &amp; Rubinoff D" refId="B15" refString="Doorenweerd C, San Jose M, Leblanc L, Barr N, Geib SM, Chung AYC, Dupuis JR, Ekayanti A, Fiegalan E, Hemachandra KS, Hossain MA, Huang C, Hsu Y, Morris KY, Maryani A, Mustapeng A, Niogret J, Pham TH, Thi Nguyen N, Sirisena UGAI, Todd T, Rubinoff D (2024 b) Towards a better future for DNA barcoding: Evaluating monophyly- and distance-based species identification using COI gene fragments of Dacini fruit flies. Molecular Ecology Resources 13987. https://doi.org/10.1111/1755-0998.13987">Doorenweerd et al. 2024 b</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELHBI" type="description">
<paragraph id="67EAD8FDE240C3A67FD508C94B3F8DBE">
<heading id="2153F1994037E57C9764866BAA12339C" reason="title">Description.</heading>
</paragraph>
<paragraph id="591B6DD79ADA497A3A38411A8F2A7512">
<emphasis id="5590EFC3BB9188CA5D1FABA8E8F20CB7" bold="true">
Male.
<emphasis id="4EE0BB20A3D50421DDF323F6E6479679" bold="true" italics="true">Head</emphasis>
</emphasis>
(Fig.
<figureCitation id="93933E8460799D6F40C795484195AE3B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Holotype male of Bactrocera melanobivittata sp. nov., UHIM. ms 12171 A dorsal view of head and thorax B dorsal view of abdomen C anterior view of the head D lateral view of habitus E posterior view of abdomen showing the ceromata F left wing, dissected and slide mounted G detail image of the genitalia showing the surstylus length." figureDoi="10.3897/zookeys.1240.148768.figure2" httpUri="https://binary.pensoft.net/fig/1348694">2 C</figureCitation>
). Fulvous with oval black spots in the antennal furrows. Antennal segments fulvous to dark fulvous distally, combined length less than the height of the head.
<emphasis id="A7AD95EEB65A730890C1877250094242" bold="true" italics="true">Thorax</emphasis>
(Fig.
<figureCitation id="C256B23C86B55E332D08C2603F958ECF" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Holotype male of Bactrocera melanobivittata sp. nov., UHIM. ms 12171 A dorsal view of head and thorax B dorsal view of abdomen C anterior view of the head D lateral view of habitus E posterior view of abdomen showing the ceromata F left wing, dissected and slide mounted G detail image of the genitalia showing the surstylus length." figureDoi="10.3897/zookeys.1240.148768.figure2" httpUri="https://binary.pensoft.net/fig/1348694">2 A, B, D, E</figureCitation>
). Scutum and pleural areas all black with narrow red-brown areas surrounding the yellow markings. Two lateral longitudinal red stripes reach from the anterior of the scutum to half-way of the postsutural yellow vittae. Yellow markings: postpronotal lobes; notopleura; presutural area adjacent to notopleura; postsutural yellow lateral vittae of equal width throughout and reach just past intra-alar seta; mesopleural stripe dorsally wider than notopleuron but does not reach postpronotal lobe; anatergite and katatergite. Scutellum yellow with a narrow black basal band. Setae: one pair scutellar; one pair prescutellar; one pair intra-alar; one pair posterior supra-alar; one pair anterior supra-alar; two pair notopleural and four scapular.
<emphasis id="91729C72DB774CAB5B5932A97AC5663D" bold="true" italics="true">Abdomen</emphasis>
(Fig.
<figureCitation id="CBCEDEE671C1AC07DC7D5E17EA60875E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Holotype male of Bactrocera melanobivittata sp. nov., UHIM. ms 12171 A dorsal view of head and thorax B dorsal view of abdomen C anterior view of the head D lateral view of habitus E posterior view of abdomen showing the ceromata F left wing, dissected and slide mounted G detail image of the genitalia showing the surstylus length." figureDoi="10.3897/zookeys.1240.148768.figure2" httpUri="https://binary.pensoft.net/fig/1348694">2 B, D, E, G</figureCitation>
). Diamond-shaped; terga free; pecten of setae on tergum III present. Dorsal side of abdomen fulvous with a black T on segments III
<collectionCode id="1FAAC40E8A609A67035F168CE560755B" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
, the medial line narrows posteriorly to a point on segment
<collectionCode id="41CCBC17C094441B1A37C17D8CAFA40C" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
. Anterolateral corners of segment
<collectionCode id="5121ABC0C7FB120C558F295DFA09A952">IV</collectionCode>
black. Ceromata brown. Posterior lobe of the male surstylus short (Fig.
<figureCitation id="FE10DA230E0631228F144E482D7798B5" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Holotype male of Bactrocera melanobivittata sp. nov., UHIM. ms 12171 A dorsal view of head and thorax B dorsal view of abdomen C anterior view of the head D lateral view of habitus E posterior view of abdomen showing the ceromata F left wing, dissected and slide mounted G detail image of the genitalia showing the surstylus length." figureDoi="10.3897/zookeys.1240.148768.figure2" httpUri="https://binary.pensoft.net/fig/1348694">2 G</figureCitation>
). Sternum
<collectionCode id="692A35025B88CFA19EF8E13867E1F325" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
with a narrow concavity on the posterior margin.
<emphasis id="2965F77E7265DCB428FCFF93D126476F" bold="true" italics="true">Legs</emphasis>
(Fig.
<figureCitation id="04D2ED1F854DDD487E48B3AC9C355D04" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Holotype male of Bactrocera melanobivittata sp. nov., UHIM. ms 12171 A dorsal view of head and thorax B dorsal view of abdomen C anterior view of the head D lateral view of habitus E posterior view of abdomen showing the ceromata F left wing, dissected and slide mounted G detail image of the genitalia showing the surstylus length." figureDoi="10.3897/zookeys.1240.148768.figure2" httpUri="https://binary.pensoft.net/fig/1348694">2 D</figureCitation>
). All femora and tibia fulvous.
<emphasis id="D46A44160029618F3259494CB6D4A6E2" bold="true" italics="true">Wings</emphasis>
(Fig.
<figureCitation id="4B2395F8A8131B45E0B2625500EF752C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Holotype male of Bactrocera melanobivittata sp. nov., UHIM. ms 12171 A dorsal view of head and thorax B dorsal view of abdomen C anterior view of the head D lateral view of habitus E posterior view of abdomen showing the ceromata F left wing, dissected and slide mounted G detail image of the genitalia showing the surstylus length." figureDoi="10.3897/zookeys.1240.148768.figure2" httpUri="https://binary.pensoft.net/fig/1348694">2 F</figureCitation>
). Wing length
<quantity id="95897B7230E5ED4D1FC574A4DB42184C" metricMagnitude="-3" metricUnit="m" metricValue="6.5" unit="mm" value="6.5">6.5 mm</quantity>
. Cells
<emphasis id="670292DC6CA752F123C672D4455360F7" italics="true">bc</emphasis>
and
<emphasis id="CB2B76AEAE8AF085BF13C98163617874" italics="true">c</emphasis>
clear, faint tint in cell
<emphasis id="C887D8F7782DAF40BAE7F73112D6FE40" italics="true">sc.</emphasis>
Costal band confluent with vein
<collectionCode id="854559468663A72A206E8CE7C1D05250" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
<subScript id="1C44710F92285A01948F595885C0A378">2 + 3</subScript>
and continues at more or less the same width until just past where vein
<collectionCode id="BFBF62530293EBCC8DACC13315BF5E9B" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
<subScript id="1C88EC4871300E3A44E90570E7EEB07B">4 + 5</subScript>
reaches the costa. Anal streak absent, vein CuA
<subScript id="5CBA9263A274CC4F36C3EC1225C5CEA8">2</subScript>
and A
<subScript id="9069FB41853A07A3D37E82FCBE3ACC46">1</subScript>
merge at ~ 0.8 length of A
<subScript id="C9F3D5A1BEA9BF09CF60AA7EEDA44AC8">1</subScript>
. Supernumerary lobe not pronounced, inconspicuous.
<emphasis id="4BE09CD8C93DE6B4D3DB0E2D42C579DA" bold="true">Female.</emphasis>
Unknown.
</paragraph>
<caption id="1EBB2EE2754BE22E90D2AF31516F1BEC" ID-DOI="10.3897/zookeys.1240.148768.figure2" ID-arpha="D2322B26-9B03-515A-AB39-3E51F216AAAA" httpUri="https://binary.pensoft.net/fig/1348694" startId="F2">
<paragraph id="3EAE4638C22E1DD904DE3E348CEB165C">
<label id="75B48B5635EB2442A969E45D59432BBD">Figure 2.</label>
</paragraph>
<paragraph id="DD7F820062F02B1D2A343FCC662102CE">
Holotype male of
<taxonomicName id="98525266F2D756B06E4D78A50034BDDD" authorityName="Doorenweerd &amp; Chung &amp; Mustapeng &amp; Rubinoff" authorityYear="2025" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="melanobivittata" status="sp. nov.">
<emphasis id="99B97FD478BD98CA05D619199592AAED" italics="true">Bactrocera melanobivittata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="24F1759F7DD00106535C48899C5337EF" rank="species">sp. nov.</taxonomicNameLabel>
,
<abbrev content-type="institution" id="ABBRID0E5JBI" xlink_title="University of Hawaiʻi at Mānoa">UHIM</abbrev>
. ms 12171
<emphasis id="E61ED616ADEE011D383334279C71FA01" bold="true">A</emphasis>
dorsal view of head and thorax
<emphasis id="2D78D0D3E1BA9CD837FE823816F46C35" bold="true">B</emphasis>
dorsal view of abdomen
<emphasis id="CD96F22426F3D547A0F1B573045E8EB8" bold="true">C</emphasis>
anterior view of the head
<emphasis id="E9A0DCDB3A31BF49F747AB6BC1FE717E" bold="true">D</emphasis>
lateral view of habitus
<emphasis id="D8C5002218CA774163B2F1E0D884280E" bold="true">E</emphasis>
posterior view of abdomen showing the ceromata
<emphasis id="EF658669B0CA298BEDD8CD2CBEA0A5BA" bold="true">F</emphasis>
left wing, dissected and slide mounted
<emphasis id="A240BF1D87003A284CCF5360F3397D7B" bold="true">G</emphasis>
detail image of the genitalia showing the surstylus length.
</paragraph>
</caption>
<caption id="837C9B14F6A2DBDCE28363BA34DCC559" ID-DOI="10.3897/zookeys.1240.148768.figure3" ID-arpha="A79AB5A3-5B2E-594B-BA7C-E8851D1FB254" httpUri="https://binary.pensoft.net/fig/1348695" startId="F3">
<paragraph id="929354FA76AE136E05A45FD1634CDEA6">
<label id="5B77F583BC46AE72070515AFFA102786">Figure 3.</label>
</paragraph>
<paragraph id="33B37CEBB99044CB86215CFD06CC4CE4">
Variation in dorsal scutum coloration of
<taxonomicName id="46CB3E5C97B9C67B595383FE7408BBAE" authorityName="Lin &amp; Wang" authorityYear="2005" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="bivittata">
<emphasis id="32FEECF39D7BFBF0A9A9983D798C0EB0" italics="true">Bactrocera bivittata</emphasis>
</taxonomicName>
, sister species to
<taxonomicName id="359B2469724ECFFB5334EF9B194E2505" authorityName="Doorenweerd &amp; Chung &amp; Mustapeng &amp; Rubinoff" authorityYear="2025" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="melanobivittata" status="sp. nov.">
<emphasis id="F4C31720D02D6C5FFCA5E2DDE866A9C3" italics="true">B. melanobivittata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E514EAD6646005390B635097CC648D9A" rank="species">sp. nov.</taxonomicNameLabel>
Specimens from China and Laos, photographed in ethanol. University of Hawaii Insect Museum specimen identifiers, from left to right, top to bottom: ms 03606, ms 01304, ms 03607, ms 01305, ms 03604, ms 03609, ms 03608, and ms 01790.
</paragraph>
</caption>
<paragraph id="003AD55AC017B1B1255376A1AB65D7D3">
<emphasis id="F9CA0A43C1413BD6B629D8BC5D9EFF5C" bold="true">Male lure.</emphasis>
Methyl eugenol.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4LBI" type="host">
<paragraph id="105D04F4B24CDF6AD77D2BF783B22158">
<heading id="AAF1A1620541A52E67A898F580C2D938" reason="title">Host plant.</heading>
</paragraph>
<paragraph id="F3404B9EC3A4C08324B987D71AD29EE2">Unknown.</paragraph>
</subSubSection>
<subSubSection id="SECID0ECMBI" type="etymology">
<paragraph id="CC19EF7277D9212C0BBC796080A30AFD">
<heading id="51183AFEF1EB7999730B59BFFB356DA9" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="A6B16E20CA5A0347BB8DA6C680521B59">
The species epithet is a compound adjective formed from the Latin
<emphasis id="F5E137865E99F58C225A66724596BCF5" italics="true">melano</emphasis>
, meaning dark, and its nearest sister species
<emphasis id="67BE9B4E04F29D9E96B9999731129093" italics="true">B. bivittata</emphasis>
, because it has an overall darker appearance of the latter.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EVMBI" type="comments">
<paragraph id="7A785A1FFA584AF24E1C331CAF16EB49">
<heading id="D7AE3888B98EA9D590B81487499A1A49" reason="title">Comments.</heading>
</paragraph>
<paragraph id="13C14D077C3E80E5F7269055B55044BD">
<taxonomicName id="593C217D1A6A609A0842DFFD4B59155C" authorityName="Doorenweerd &amp; Chung &amp; Mustapeng &amp; Rubinoff" authorityYear="2025" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="melanobivittata">
<emphasis id="92EC076C8CB3D6402B8B497F78A8D922" italics="true">Bactrocera melanobivittata</emphasis>
</taxonomicName>
was included as
<emphasis id="50D66F9813E808C2BC107228D1352DC2" italics="true">B.</emphasis>
spMalaysia 05 in the DNA barcoding study of
<bibRefCitation id="7FF51D5DEABE5F83D1BB9C043E5DDD51" DOI="10.1111/1755-0998.13987" author="Doorenweerd C &amp; San Jose M &amp; Leblanc L &amp; Barr N &amp; Geib SM &amp; Chung AYC &amp; Dupuis JR &amp; Ekayanti A &amp; Fiegalan E &amp; Hemachandra KS &amp; Hossain MA &amp; Huang C &amp; Hsu Y &amp; Morris KY &amp; Maryani A &amp; Mustapeng A &amp; Niogret J &amp; Pham TH &amp; Thi Nguyen N &amp; Sirisena UGAI &amp; Todd T &amp; Rubinoff D" refId="B15" refString="Doorenweerd C, San Jose M, Leblanc L, Barr N, Geib SM, Chung AYC, Dupuis JR, Ekayanti A, Fiegalan E, Hemachandra KS, Hossain MA, Huang C, Hsu Y, Morris KY, Maryani A, Mustapeng A, Niogret J, Pham TH, Thi Nguyen N, Sirisena UGAI, Todd T, Rubinoff D (2024 b) Towards a better future for DNA barcoding: Evaluating monophyly- and distance-based species identification using COI gene fragments of Dacini fruit flies. Molecular Ecology Resources 13987. https://doi.org/10.1111/1755-0998.13987">Doorenweerd et al. (2024 b</bibRefCitation>
). It is placed in subgenus
<taxonomicName id="33D95228EA1CAB0F1792C40C449640E7" rank="subGenus" subGenus="Bactrocera">
<emphasis id="6EBF7E6B5028E74C8F09F011059F13D3" italics="true">Bactrocera</emphasis>
</taxonomicName>
based on having a short posterior lobe of the male surstylus, sternum
<collectionCode id="F555241BAD8BEBB8E307E27B4A09474B" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
has a narrow concavity on the posterior margin, lateral postsutural yellow vitta present, medial vitta is absent, one pair of prescutellar setae and the presence of a pair of anterior supra-alar seta.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,584 @@
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<treatment id="51A5EFBD0EDD54D4979AF4650B2C3157" ID-DOI="http://doi.org/10.5281/zenodo.15625178" ID-Zenodo-Dep="15625178" ID-arpha="51A5EFBD-0EDD-54D4-979A-F4650B2C3157" LSID="urn:lsid:plazi:treatment:51A5EFBD0EDD54D4979AF4650B2C3157" httpUri="http://treatment.plazi.org/id/51A5EFBD0EDD54D4979AF4650B2C3157">
<subSubSection id="E9093247F9678D107E10182800ABD726" type="nomenclature">
<paragraph id="5E49704ABBE82226E15B51C779414D2A">
<taxonomicName id="D04EAE1CC582C715062E4516F9465709" ID-CoL="C82XC" authority="Jang, Lee, Yoo &amp; Kim, 2024" authorityName="Jang, Lee, Yoo &amp; Kim" authorityYear="2024" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="paiki">
<emphasis id="4E65BA0236CD8421632614062328F4D9" italics="true">Philodromus paiki</emphasis>
<bibRefCitation id="2446C0250FCE66EE41BE0206C4629E84" DOI="10.11626/KJEB.2023.41.4.497" author="Jang CM &amp; Lee SY &amp; Yoo JS &amp; Kim ST" refId="B24" refString="Jang CM, Lee SY, Yoo JS, Kim ST (2024 a) Description of Philodromus paiki sp. nov. and Philodromus spinitarsis Simon, 1895 (Araneae: Philodromidae) from Korea. Korean Journal of Environmental Biology 41 (4) [2023]: 497 - 504. [publ. in January 2024] https://doi.org/10.11626/KJEB.2023.41.4.497" year="2024">Jang, Lee, Yoo &amp; Kim, 2024</bibRefCitation>
</taxonomicName>
</paragraph>
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<subSubSection id="62C98071BDD4A6FC2A3CC1688D37D7D8" type="description">
<paragraph id="0DFEE74163074D2F67F88B2DD7CF42B9">
<figureCitation id="977224058434BCA50A5CBED8D2B34D16" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Sample collection locations for this project. Hubei Province 1 Jiugongshan National Nature Reserve 2 Hubei University of Science and Technology 3 Jiuling Mountain 4 Qizimei Mountain National Nature Reserve; Guizhou Province 5 Fanjingshan National Nature Reserve 6 Mayanghe National Nature Reserve 7 Xishui National Nature Reserve 8 Guiyang City, Kaiyang County 9 Guiyang City, Wudang District 10 Guiyang City, Yunyan District 11 Guiyang City, Huaxi District." figureDoi="10.3897/zookeys.1240.149456.figure1" httpUri="https://binary.pensoft.net/fig/1348759">Figs 1</figureCitation>
,
<figureCitation id="395ADF9A3F0242B31AD02BAFD7FCD7EE" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Living specimens of some Philodromidae species treated in this paper A, B Philodromus paiki (♂ ♀) C, D Philodromus rufus (♂ ♀). Photographs by Q. Lu (Shenzhen)." figureDoi="10.3897/zookeys.1240.149456.figure3" httpUri="https://binary.pensoft.net/fig/1348761">3 A, B</figureCitation>
,
<figureCitation id="01EB67084DAFA9CE9ECB6CA8C4E73F7E" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Philodromus paiki, male, habitus (A C) and left palp (D F) A dorsal view B ventral view C lateral view D prolateral view E ventral view F retrolateral view (teal green dashed lines represent the contours of DTA, RTA and VTA). Abbreviations: aSDL = ascending part of sperm duct loop; Con = conductor; Cy = cymbium; dSDL = descending part of sperm duct loop; DTA = dorsal tibial apophysis; Em = embolus; EmB = embolic base; EmT = embolic tip; IR = intertegular retinaculum; RTA = retrolateral tibial apophysis; SD = sperm duct; Te = tegulum; VTA = ventral tibial apophysis. Scale bars: 1 mm (equal for A C); 0.5 mm (equal for D F)." figureDoi="10.3897/zookeys.1240.149456.figure12" httpUri="https://binary.pensoft.net/fig/1348770">12</figureCitation>
,
<figureCitation id="47F35DE427DB03A392542247542E234C" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Philodromus paiki, female, habitus (A C) and epigyne (D, E) A dorsal view B ventral view C lateral view D ventral view E dorsal view. Abbreviations: CD = copulatory duct; EG = epigynal groove; FD = fertilisation duct; MS = median septum; R = receptaculum; rMS = rim of median septum (teal green line); rSEF = rim of sclerotised epigynal fold (blue line); SEF = sclerotised epigynal fold. Scale bars: 1 mm (equal for A C); 0.2 mm (equal for D, E)." figureDoi="10.3897/zookeys.1240.149456.figure13" httpUri="https://binary.pensoft.net/fig/1348771">13</figureCitation>
,
<figureCitation id="7B201C1C61D72DFAD7E7572ACB33501C" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Distribution records of the philodromid species in Guizhou Province (A) and Hubei Province (B)." figureDoi="10.3897/zookeys.1240.149456.figure23" httpUri="https://binary.pensoft.net/fig/1348781">23 A, B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="D366BFA43E7ED3EDEC307EB0DC7B1677" type="reference_group">
<paragraph id="5A47F4A47B724C83B5E480532478D125">
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<treatmentCitation id="736495D2534189895FA6DF3578E41942">
<taxonomicName id="45F48C11CB7AC2258F5948B59A1B1738" ID-CoL="4G8S5" baseAuthorityName="De Geer" baseAuthorityYear="1778" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="fuscomarginatus">
<emphasis id="D5F02C80063EE4AE33AD527B87B56C84" italics="true">Philodromus fuscomarginatus</emphasis>
</taxonomicName>
<comment id="C4828DE877FA8174281A4A87582BC9FC">
:
<treatmentCitation id="31660BB69922F1ECDDEE49327A468A2E" author="Nakatsudi K" page="14" year="1942">
<bibRefCitation id="AA0C4E45BED93758A42FF3911BFBABEB" DOI="10.2476/asjaa.7.7" author="Nakatsudi K" firstAuthor="Nakatsudi" issue="1" journalOrPublisher="Acta Arachnologica" page="14" pagination="7-18" refId="B37" refString="Nakatsudi K (1942) Spiders from Heiho Prefecture, North Manchuria, China. Acta Arachnologica 7 (1): 718. https://doi.org/10.2476/asjaa.7.7" title="Spiders from Heiho Prefecture, North Manchuria, China." volume="7" year="1942">Nakatsudi 1942: 14</bibRefCitation>
</treatmentCitation>
, fig. 5 A, B (
<specimenCount id="49C873DEB32728B9EC66CE9A72B96401" count="1" type="male"></specimenCount>
; misidentified per
<treatmentCitation id="3FDC9A1BEFA175F786C968C5D0BD0729" author="Jang CM &amp; Lee SY &amp; Yoo JS &amp; Kim ST" page="498" year="">
<treatmentCitation id="4D98C01DBFF76ED2405BFCE7F5C47867">
<bibRefCitation id="217563273E5E14ABEDCF0B8E5BF52025" DOI="10.11626/KJEB.2023.41.4.497" author="Jang CM &amp; Lee SY &amp; Yoo JS &amp; Kim ST" refId="B24" refString="Jang CM, Lee SY, Yoo JS, Kim ST (2024 a) Description of Philodromus paiki sp. nov. and Philodromus spinitarsis Simon, 1895 (Araneae: Philodromidae) from Korea. Korean Journal of Environmental Biology 41 (4) [2023]: 497504. [publ. in January 2024] https://doi.org/10.11626/KJEB.2023.41.4.497">Jang et al. 2024 a</bibRefCitation>
</treatmentCitation>
: 498
</treatmentCitation>
);
<treatmentCitation id="B5D62FBCF3DBC5009910B72C50FD95C4" author="Paik KY" page="430" year="1979">
<bibRefCitation id="1875B2936F10AF0844520038E31FD4E0" author="Paik KY" firstAuthor="Paik" journalOrPublisher="Research Review of Kyungpook National University" page="430" pagination="421-452" refId="B43" refString="Paik KY (1979) Korean spiders of the genus Philodromus (Araneae: Thomisidae). Research Review of Kyungpook National University 28: 421452." title="Korean spiders of the genus Philodromus (Araneae: Thomisidae)." volume="28" year="1979">Paik 1979: 430</bibRefCitation>
</treatmentCitation>
, figs 4060 (
<specimenCount id="274B53CD0855D6DD15095F121EE2369E" count="1" type="male"></specimenCount>
<specimenCount id="959922396F890F09982F8D24372B41F1" count="1" type="female"></specimenCount>
; misidentified per
<treatmentCitation id="190AF0A37004E1E1563D3D3FAF29CE60" author="Jang CM &amp; Lee SY &amp; Yoo JS &amp; Kim ST" page="498" year="">
<treatmentCitation id="410D080972B7A89A48051372B26682A4">
<bibRefCitation id="9E1FC139B83742E7999EF5914691AE85" DOI="10.11626/KJEB.2023.41.4.497" author="Jang CM &amp; Lee SY &amp; Yoo JS &amp; Kim ST" refId="B24" refString="Jang CM, Lee SY, Yoo JS, Kim ST (2024 a) Description of Philodromus paiki sp. nov. and Philodromus spinitarsis Simon, 1895 (Araneae: Philodromidae) from Korea. Korean Journal of Environmental Biology 41 (4) [2023]: 497504. [publ. in January 2024] https://doi.org/10.11626/KJEB.2023.41.4.497">Jang et al. 2024 a</bibRefCitation>
</treatmentCitation>
: 498
</treatmentCitation>
).
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="512B0214843D11305AA649481599030E">
<treatmentCitationGroup id="0B9BCC2BA72FBF1D83014F4FA7464C53">
<treatmentCitation id="8882EA4E7A8F6AE9E9B676D0FBC0A470">
<taxonomicName id="E8AA301D46FFFFA212B8B3780ACC618C" ID-CoL="4G8VX" authorityName="Simon" authorityYear="1895" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="spinitarsis">
<emphasis id="E799D118B03088F2A5B62ED26041BC7D" italics="true">Philodromus spinitarsis</emphasis>
</taxonomicName>
<comment id="E4C1A912FE99E6D03B0B8497E2A1872D">
:
<treatmentCitation id="523DD576E303B03B777137B68A618981" author="Kim JP &amp; Jung JY" page="201" year="2001">
<bibRefCitation id="5255D97AC082CC472F98DC1052F364F7" author="Kim JP &amp; Jung JY" firstAuthor="Kim" journalOrPublisher="Korean Arachnology" page="201" pagination="185-222" refId="B27" refString="Kim JP, Jung JY (2001) A revisional study of the spider family Philodromidae O. P. - Cambridge, 1871 (Arachnida: Araneae) from Korea. Korean Arachnology 17: 185222." title="A revisional study of the spider family Philodromidae O. P. - Cambridge, 1871 (Arachnida: Araneae) from Korea." volume="17" year="2001">Kim and Jung 2001: 201</bibRefCitation>
</treatmentCitation>
, figs 6668 (
<specimenCount id="B08F878D48C2E8B963E58B91E830C99B" count="1" type="male"></specimenCount>
<specimenCount id="36B92BA3E03FC06A02E05A06CF97B256" count="1" type="female"></specimenCount>
; misidentified per
<treatmentCitation id="906ED0896DFAEDD610331A4C6DA8C05D" author="Jang CM &amp; Lee SY &amp; Yoo JS &amp; Kim ST" page="498" year="">
<treatmentCitation id="22395807D84D5DA16A340BAA3D0A439C">
<bibRefCitation id="C8AF7E0101BE6544952C0025361B3B90" DOI="10.11626/KJEB.2023.41.4.497" author="Jang CM &amp; Lee SY &amp; Yoo JS &amp; Kim ST" refId="B24" refString="Jang CM, Lee SY, Yoo JS, Kim ST (2024 a) Description of Philodromus paiki sp. nov. and Philodromus spinitarsis Simon, 1895 (Araneae: Philodromidae) from Korea. Korean Journal of Environmental Biology 41 (4) [2023]: 497504. [publ. in January 2024] https://doi.org/10.11626/KJEB.2023.41.4.497">Jang et al. 2024 a</bibRefCitation>
</treatmentCitation>
: 498
</treatmentCitation>
);
<treatmentCitation id="7AC39BC11570E855C7FE89E845BD3532" author="Namkung J" page="510" year="2002">
<bibRefCitation id="8889AD1FAF1E32CAD4056DA1A72740BA" author="Namkung J" firstAuthor="Namkung" journalOrPublisher="Kyo-Hak Publishing Co, Seoul" page="510" refId="B38" refString="Namkung J (2002) The spiders of Korea. Kyo-Hak Publishing Co, Seoul, 648 pp." title="The spiders of Korea." year="2002">Namkung 2002: 510</bibRefCitation>
</treatmentCitation>
, fig. 41.8 a, b (
<specimenCount id="3A07B75C8F47150338F83869B4A3DD03" count="1" type="male"></specimenCount>
<specimenCount id="CA36AC8035322D140CE91A54F926A4DC" count="1" type="female"></specimenCount>
; misidentified per
<treatmentCitation id="49312402C69DB29F11A869AD4602A609" author="Jang CM &amp; Lee SY &amp; Yoo JS &amp; Kim ST" page="498" year="">
<treatmentCitation id="3A9EF4638F270F7FDED516D711DFD8D1">
<bibRefCitation id="245EC7B3EFD379FA9EC25012E8825385" DOI="10.11626/KJEB.2023.41.4.497" author="Jang CM &amp; Lee SY &amp; Yoo JS &amp; Kim ST" refId="B24" refString="Jang CM, Lee SY, Yoo JS, Kim ST (2024 a) Description of Philodromus paiki sp. nov. and Philodromus spinitarsis Simon, 1895 (Araneae: Philodromidae) from Korea. Korean Journal of Environmental Biology 41 (4) [2023]: 497504. [publ. in January 2024] https://doi.org/10.11626/KJEB.2023.41.4.497">Jang et al. 2024 a</bibRefCitation>
</treatmentCitation>
: 498
</treatmentCitation>
);
<treatmentCitation id="D986D01884EDA1035C2399882ED3BB65" author="Namkung J" page="513" year="2003">
<bibRefCitation id="4806F4702CF7CF60E3FD366B64AB1881" author="Namkung J" firstAuthor="Namkung" journalOrPublisher="Kyo-Hak Publishing Co, Seoul" page="513" refId="B39" refString="Namkung J (2003) The spiders of Korea, 2 nd edn. Kyo-Hak Publishing Co, Seoul, 648 pp." title="The spiders of Korea, 2 nd edn." year="2003">Namkung 2003: 513</bibRefCitation>
</treatmentCitation>
, fig. 41.8 a, b (
<specimenCount id="A66B32C2111F3CF136769FD0C867A657" count="1" type="male"></specimenCount>
<specimenCount id="916DE74448C0A7A71E7892878A9C6D29" count="1" type="female"></specimenCount>
; misidentified per
<treatmentCitation id="3AA77EFB3C6BC1D17CD58C8F8CB7519B" author="Jang CM &amp; Lee SY &amp; Yoo JS &amp; Kim ST" page="498" year="">
<treatmentCitation id="171AED89C42949640795FEF00080670C">
<bibRefCitation id="02CA9DA874831B8704D9B88279EF1512" DOI="10.11626/KJEB.2023.41.4.497" author="Jang CM &amp; Lee SY &amp; Yoo JS &amp; Kim ST" refId="B24" refString="Jang CM, Lee SY, Yoo JS, Kim ST (2024 a) Description of Philodromus paiki sp. nov. and Philodromus spinitarsis Simon, 1895 (Araneae: Philodromidae) from Korea. Korean Journal of Environmental Biology 41 (4) [2023]: 497504. [publ. in January 2024] https://doi.org/10.11626/KJEB.2023.41.4.497">Jang et al. 2024 a</bibRefCitation>
</treatmentCitation>
: 498
</treatmentCitation>
).
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="84F0847E2317FE5CD5DBD2EC25D7446E">
<treatmentCitationGroup id="AC757FA5408CB250B85E520737483E26">
<treatmentCitation id="3BA8967762182D55EF8138BAA1902EEA">
<taxonomicName id="A81F71CA5D8EA5F9DE89C1BFAF9E2C0B" ID-CoL="779RM" baseAuthorityName="Thorell" baseAuthorityYear="1872" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="poecilus">
<emphasis id="ABE792952C82D6DA62AB72A0044133ED" italics="true">Philodromus poecilus</emphasis>
</taxonomicName>
<comment id="46A9EE9420E9191C2DEA0C598ACD8BA9">
:
<treatmentCitation id="EC02763AA52C3E6E758CAD453B390678" author="Kim ST &amp; Lee SY" page="74" year="2017">
<bibRefCitation id="F49ADA50AFF187AD31B7A1AD9FED2392" author="Kim ST &amp; Lee SY" firstAuthor="Kim" issue="42" journalOrPublisher="Invertebrate Fauna of Korea" page="74" pagination="1-122" refId="B28" refString="Kim ST, Lee SY (2017) Arthropoda: Arachnida: Aaraneae [sic]: Oecobiidae, Oxyopidae, Cybaeidae, Dictynidae, Sparassidae, Philodromidae. Spiders II. Invertebrate Fauna of Korea 21 (42): 1122." title="Arthropoda: Arachnida: Aaraneae [sic]: Oecobiidae, Oxyopidae, Cybaeidae, Dictynidae, Sparassidae, Philodromidae. Spiders II." volume="21" year="2017">Kim and Lee 2017: 74</bibRefCitation>
</treatmentCitation>
, fig. 42 A D (
<specimenCount id="5DF2C9E5458FDE061C70E54800D55040" count="1" type="male"></specimenCount>
<specimenCount id="A9B70F2D24D4F38DFC46F315C75433E5" count="1" type="female"></specimenCount>
; misidentified per
<treatmentCitation id="1C68E9AA2A0EF069790CB29AC5D9D092" author="Jang CM &amp; Lee SY &amp; Yoo JS &amp; Kim ST" page="498" year="">
<treatmentCitation id="3F928B99005E883B2302B2054C1A23D1">
<bibRefCitation id="43B87A3A0A9FDB3E882C9CBC0421109D" DOI="10.11626/KJEB.2023.41.4.497" author="Jang CM &amp; Lee SY &amp; Yoo JS &amp; Kim ST" refId="B24" refString="Jang CM, Lee SY, Yoo JS, Kim ST (2024 a) Description of Philodromus paiki sp. nov. and Philodromus spinitarsis Simon, 1895 (Araneae: Philodromidae) from Korea. Korean Journal of Environmental Biology 41 (4) [2023]: 497504. [publ. in January 2024] https://doi.org/10.11626/KJEB.2023.41.4.497">Jang et al. 2024 a</bibRefCitation>
</treatmentCitation>
: 498
</treatmentCitation>
).
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="41143E2D4006DD953828E246AA1BAA34">
<treatmentCitationGroup id="494A7C1CC599E5E4B0D205F4488757B0">
<treatmentCitation id="3505BF2D7B5448F8188435B9CAE1AAB7" author="Jang CM &amp; Lee SY &amp; Yoo JS &amp; Kim ST" page="498" year="">
<taxonomicName id="964BEFE3FA780EEF057164A856016293" ID-CoL="C82XC" authority="Jang et al. 2024 a: 498" authorityName="Jang" authorityPageNumber="498" authorityYear="2024" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="paiki">
<emphasis id="CDF3021F8C101687AE06E406E91F7DF7" italics="true">Philodromus paiki</emphasis>
<comment id="CA731A372360B494EA667A91CAF8EF63">
<bibRefCitation id="D5EDA6671AA02DE6BFE0A074743B6262" DOI="10.11626/KJEB.2023.41.4.497" author="Jang CM &amp; Lee SY &amp; Yoo JS &amp; Kim ST" refId="B24" refString="Jang CM, Lee SY, Yoo JS, Kim ST (2024 a) Description of Philodromus paiki sp. nov. and Philodromus spinitarsis Simon, 1895 (Araneae: Philodromidae) from Korea. Korean Journal of Environmental Biology 41 (4) [2023]: 497504. [publ. in January 2024] https://doi.org/10.11626/KJEB.2023.41.4.497">Jang et al. 2024 a</bibRefCitation>
: 498, fig. 1 A K (
<specimenCount id="A8C56D604E01E6D4D5F13ED9124DFE1F" count="1" type="male"></specimenCount>
<specimenCount id="E4C6AF564135E32360F86D0C8C3C4015" count="1" type="female"></specimenCount>
).
</comment>
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</paragraph>
</subSubSection>
<subSubSection id="SECID0E3TAI" type="material">
<paragraph id="04AA51A2D2B214E1CDD24A4E389A101B">
<heading id="D062C973F8FB6CE9C8ABCFD65ADF85EC" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="258526711BA7A21C38A91B9404990D0E">
<materialsCitation id="7B27D75878C47C7EC667E095F6F89743" country="China" location="Guizhou Province" specimenCount="1" stateProvince="Guizhou">
<emphasis id="D48CE304BAE9F2B9C2D367AA8BB53CAE" bold="true">
<collectingCountry id="29E2B367BB7ED1F49B57D6A58CD7D7AA" name="China">China</collectingCountry>
</emphasis>
:
<collectingRegion id="4A70ED5A6DEE6DAF5DEAB0D3D1769A41" country="China" name="Guizhou">Guizhou Province</collectingRegion>
</materialsCitation>
: •
<materialsCitation id="1CAD106F209A897124F156281A8982EA" collectingDate="2015-05-25" collectorName="H. Yu" country="China" county="Guiyang City" elevation="1071" latitude="26.64" location="Dongfeng Town" longLatPrecision="744" longitude="106.8" municipality="Wudang District" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Guizhou">
<specimenCount id="09F21D9CB82862766BDB5993B4B2F8D5" count="1" type="male">1 ♂</specimenCount>
,
<specimenCount id="AE38549763999D758D89809DCD92D8A0" count="1" type="female">1 ♀</specimenCount>
,
<collectingCounty id="F940AEA4AB5EBCB592228A446B98A2A6">Guiyang City</collectingCounty>
,
<collectingMunicipality id="F0401A07E8D122E4CCF0BB73E260B4D7">Wudang District</collectingMunicipality>
,
<location id="904446059C0D4D11F2FA2DE49ECB62AA" LSID="urn:lsid:plazi:treatment:51A5EFBD0EDD54D4979AF4650B2C3157:904446059C0D4D11F2FA2DE49ECB62AA" country="China" county="Guiyang City" latitude="26.64" longLatPrecision="744" longitude="106.8" municipality="Wudang District" name="Dongfeng Town" stateProvince="Guizhou">Dongfeng Town</location>
,
<collectingRegion id="B49EDA2EB1E0C4BBD04A357B7B63FF8F" country="China" name="Guizhou">Guizhou</collectingRegion>
<location id="E0D654EEF6B1B976CD631281AB315521" LSID="urn:lsid:plazi:treatment:51A5EFBD0EDD54D4979AF4650B2C3157:E0D654EEF6B1B976CD631281AB315521" country="China" county="Guiyang City" latitude="26.64" longLatPrecision="744" longitude="106.8" municipality="Wudang District" name="Education University" stateProvince="Guizhou">Education University</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EHUAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[106.800000,26.640000]}">
<geoCoordinate id="28CB379D241896749FDCE619B57C8FB6" degrees="26.64" direction="north" orientation="latitude" precision="555" value="26.64">26.64 ° N</geoCoordinate>
,
<geoCoordinate id="977138CFC20C5699168381AF5CBB431D" degrees="106.80" direction="east" orientation="longitude" precision="555" value="106.8">106.80 ° E</geoCoordinate>
</named-content>
;
<elevation id="5F1B29ABB2FCBC6186024768C27D6227" metricMagnitude="3" metricUnit="m" metricValue="1.071" unit="m" value="1071.0">
<quantity id="7CD30E5E95DCC82F9ABEA406A88EBF66" metricMagnitude="3" metricUnit="m" metricValue="1.071" unit="m" value="1071.0">1071 m</quantity>
a. s. l.
</elevation>
;
<date id="620D3F7037D674C7D56892BBBE290FA9" value="2015-05-25">
<collectingDate id="B08C27C9A5B80EC4753538BB28D36F2D" value="2015-05-25">25 May 2015</collectingDate>
</date>
;
<collectorName id="6D4E5875F5289E8648DFED62607C0C0F">H. Yu</collectorName>
et al. leg
</materialsCitation>
. •
<materialsCitation id="C7DFA503A936EDBCF23EE0537DE4C459" collectingDate="2022-06-10" collectionCode="YHGY" collectorName="H. Yu &amp; Q. Lu" country="China" county="Kaiyang County" elevation="812" latitude="26.93" location="Table" longLatPrecision="744" longitude="107.07" municipality="GenBank" specimenCode="YHGY 325" specimenCount="1" specimenCount-female="1" stateProvince="Hubei">
<specimenCount id="F6B058A37E7C4604BA17D485BC40084D" count="1" type="female">1 ♀</specimenCount>
(
<specimenCode id="7D1165B05986572B74D14D9B5FC80274" collectionCode="YHGY">YHGY 325</specimenCode>
used for sequencing,
<collectingMunicipality id="0D68877DA339DF34BEB8F06284E24E73">GenBank</collectingMunicipality>
accession numbers in
<location id="1CB0FDCB2A00E9D2B6610B773FDACFE3" LSID="urn:lsid:plazi:treatment:51A5EFBD0EDD54D4979AF4650B2C3157:1CB0FDCB2A00E9D2B6610B773FDACFE3" country="China" county="Kaiyang County" latitude="26.93" longLatPrecision="744" longitude="107.07" municipality="GenBank" name="Table" stateProvince="Hubei">Table</location>
<tableCitation id="75F1B7DAB6DE39725B79F9084B21C311" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Voucher specimen information." httpUri="http://table.plazi.org/id/12F54BE0BDE6512C62E0465BE733F517" tableUuid="12F54BE0BDE6512C62E0465BE733F517">1</tableCitation>
),
<location id="902CBDB15C4ADAE5F97D99B5EFFF57B9" LSID="urn:lsid:plazi:treatment:51A5EFBD0EDD54D4979AF4650B2C3157:902CBDB15C4ADAE5F97D99B5EFFF57B9" country="China" county="Kaiyang County" latitude="26.93" longLatPrecision="744" longitude="107.07" municipality="GenBank" name="Guiyang City" stateProvince="Hubei">Guiyang City</location>
,
<collectingCounty id="43BB3CA187A385ECE09AD24381616FF7">Kaiyang County</collectingCounty>
,
<location id="33EE7F50CE4D711E29A4CE1061743A23" LSID="urn:lsid:plazi:treatment:51A5EFBD0EDD54D4979AF4650B2C3157:33EE7F50CE4D711E29A4CE1061743A23" country="China" county="Kaiyang County" latitude="26.93" longLatPrecision="744" longitude="107.07" municipality="GenBank" name="Longgang Town" stateProvince="Hubei">Longgang Town</location>
,
<location id="ADE73152B1789D517437ED60E1F6CA0D" LSID="urn:lsid:plazi:treatment:51A5EFBD0EDD54D4979AF4650B2C3157:ADE73152B1789D517437ED60E1F6CA0D" country="China" county="Kaiyang County" latitude="26.93" longLatPrecision="744" longitude="107.07" municipality="GenBank" name="Pingshan Village" stateProvince="Hubei">Pingshan Village</location>
,
<location id="0648FEE76A885051F2360C926BBDD83F" LSID="urn:lsid:plazi:treatment:51A5EFBD0EDD54D4979AF4650B2C3157:0648FEE76A885051F2360C926BBDD83F" country="China" county="Kaiyang County" latitude="26.93" longLatPrecision="744" longitude="107.07" municipality="GenBank" name="Zijiang Rift Scenic Area" stateProvince="Hubei">Zijiang Rift Scenic Area</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ETUAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[107.070000,26.930000]}">
<geoCoordinate id="07BE2C2DDED8D43F6BA73074D23C18E8" degrees="26.93" direction="north" orientation="latitude" precision="555" value="26.93">26.93 ° N</geoCoordinate>
,
<geoCoordinate id="71786C816F1C9B0774255956CF357A2D" degrees="107.07" direction="east" orientation="longitude" precision="555" value="107.07">107.07 ° E</geoCoordinate>
</named-content>
;
<elevation id="373B9199FEF3D9B78EC71677C756DB1E" metricMagnitude="2" metricUnit="m" metricValue="8.12" unit="m" value="812.0">
<quantity id="0BD06D7A65482A569A276B5549C4513F" metricMagnitude="2" metricUnit="m" metricValue="8.12" unit="m" value="812.0">812 m</quantity>
a. s. l.
</elevation>
;
<date id="D5F9229562D4A79C926F93857F85A1F0" value="2022-06-10">
<collectingDate id="4F1B07DB4771593D886BC7AE4BBA11DB" value="2022-06-10">10 June 2022</collectingDate>
</date>
;
<collectorName id="20C3E7329B3A4E903E395CC245C71BD6">H. Yu</collectorName>
&amp;
<collectorName id="DD4D0568F951555AB0BC4488B41FFA8E">Q. Lu</collectorName>
leg.;
<collectingRegion id="130B515013988D2B59EB1D7CB71CFFA4" country="China" name="Hubei">Hubei Province</collectingRegion>
</materialsCitation>
: •
<materialsCitation id="1FE5BF353A342169C05088238652C6D6" collectingDate="2020-07-03" collectionCode="YHPHI" collectorName="Q. Lu" country="China" county="GenBank" elevation="480" latitude="29.39" location="Xianning City" longLatPrecision="736" longitude="114.65" municipality="Table" specimenCode="YHPHI 005" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<specimenCount id="2BEBB4617716320A956E46D5BC7C853F" count="1" type="male">1 ♂</specimenCount>
,
<specimenCount id="F20BB6F967D3BE990F408AE4EE3D64B5" count="1" type="female">1 ♀</specimenCount>
(
<specimenCount id="8847582A315BAABA32DE60DE5D231535" count="1" type="male">1 ♂</specimenCount>
,
<specimenCode id="15E48401053C0D0DFE728DB7C950C415" collectionCode="YHPHI">YHPHI 005</specimenCode>
used for sequencing,
<collectingCounty id="56742E967801790077BBD399EE18A753">GenBank</collectingCounty>
accession numbers in
<collectingMunicipality id="4C3812780EF151BF66F3C6E6E5051FBF">Table</collectingMunicipality>
<tableCitation id="CD664B66BDC762232046C59B73D33CD7" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Voucher specimen information." httpUri="http://table.plazi.org/id/12F54BE0BDE6512C62E0465BE733F517" tableUuid="12F54BE0BDE6512C62E0465BE733F517">1</tableCitation>
),
<location id="DD0A87CDF891652FF5FA1EB7CF9DA01B" LSID="urn:lsid:plazi:treatment:51A5EFBD0EDD54D4979AF4650B2C3157:DD0A87CDF891652FF5FA1EB7CF9DA01B" country="China" county="GenBank" latitude="29.39" longLatPrecision="736" longitude="114.65" municipality="Table" name="Xianning City">Xianning City</location>
, Jiugongshan National Nature Reserve,
<location id="154BA0A4A7D5EEAE4BAFC1506AC4B480" LSID="urn:lsid:plazi:treatment:51A5EFBD0EDD54D4979AF4650B2C3157:154BA0A4A7D5EEAE4BAFC1506AC4B480" country="China" county="GenBank" latitude="29.39" longLatPrecision="736" longitude="114.65" municipality="Table" name="Yunzhonghu Scenic Area">Yunzhonghu Scenic Area</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E6UAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[114.650000,29.390000]}">
<geoCoordinate id="F0015778AF22B663B123D8623D6F8B5A" degrees="29.39" direction="north" orientation="latitude" precision="555" value="29.39">29.39 ° N</geoCoordinate>
,
<geoCoordinate id="B73EC596696C4A9B38326CB85F050D5D" degrees="114.65" direction="east" orientation="longitude" precision="555" value="114.65">114.65 ° E</geoCoordinate>
</named-content>
;
<elevation id="F1573BE698BCEAF4E30A046225FB28EB" metricMagnitude="2" metricUnit="m" metricValue="4.8" unit="m" value="480.0">
<quantity id="E7A352A40DB430EEDEF0772CA9C9E074" metricMagnitude="2" metricUnit="m" metricValue="4.8" unit="m" value="480.0">480 m</quantity>
a. s. l.
</elevation>
;
<date id="1CBB3CAB1F87CCF99C8E0A5D952CF0DB" value="2020-07-03">
<collectingDate id="640751A1D7E63329B8AC0A338EBA1A18" value="2020-07-03">3 July 2020</collectingDate>
</date>
;
<collectorName id="24284688077E042360C637CB450D9114">Q. Lu</collectorName>
et al. leg
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEVAI" type="diagnosis">
<paragraph id="0C0115E3E2EA2CD314EECBC7C0201F83">
<heading id="1230F9978473B54D3DC52F4FEFBB7214" reason="title">Diagnosis and description.</heading>
</paragraph>
<paragraph id="6460AE863819C40956FCA313778D98AF">
See
<bibRefCitation id="9BA160380E8A9F87BD2872B341B9346A" DOI="10.11626/KJEB.2023.41.4.497" author="Jang CM &amp; Lee SY &amp; Yoo JS &amp; Kim ST" refId="B24" refString="Jang CM, Lee SY, Yoo JS, Kim ST (2024 a) Description of Philodromus paiki sp. nov. and Philodromus spinitarsis Simon, 1895 (Araneae: Philodromidae) from Korea. Korean Journal of Environmental Biology 41 (4) [2023]: 497504. [publ. in January 2024] https://doi.org/10.11626/KJEB.2023.41.4.497">Jang et al. (2024 a</bibRefCitation>
). Habitus as in Figs
<figureCitation id="EE8599BAD8E09C1E7CF3C5E66F29B55A" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Philodromus paiki, male, habitus (A C) and left palp (D F) A dorsal view B ventral view C lateral view D prolateral view E ventral view F retrolateral view (teal green dashed lines represent the contours of DTA, RTA and VTA). Abbreviations: aSDL = ascending part of sperm duct loop; Con = conductor; Cy = cymbium; dSDL = descending part of sperm duct loop; DTA = dorsal tibial apophysis; Em = embolus; EmB = embolic base; EmT = embolic tip; IR = intertegular retinaculum; RTA = retrolateral tibial apophysis; SD = sperm duct; Te = tegulum; VTA = ventral tibial apophysis. Scale bars: 1 mm (equal for A C); 0.5 mm (equal for D F)." figureDoi="10.3897/zookeys.1240.149456.figure12" httpUri="https://binary.pensoft.net/fig/1348770">12 A C</figureCitation>
,
<figureCitation id="CCB91544C7A7B5BF5D48CF4619E522E4" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Philodromus paiki, female, habitus (A C) and epigyne (D, E) A dorsal view B ventral view C lateral view D ventral view E dorsal view. Abbreviations: CD = copulatory duct; EG = epigynal groove; FD = fertilisation duct; MS = median septum; R = receptaculum; rMS = rim of median septum (teal green line); rSEF = rim of sclerotised epigynal fold (blue line); SEF = sclerotised epigynal fold. Scale bars: 1 mm (equal for A C); 0.2 mm (equal for D, E)." figureDoi="10.3897/zookeys.1240.149456.figure13" httpUri="https://binary.pensoft.net/fig/1348771">13 A C</figureCitation>
. Male palp as in Fig.
<figureCitation id="F8C0D8D3D8D9B83FD6ACE7ACB3761E6D" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Philodromus paiki, male, habitus (A C) and left palp (D F) A dorsal view B ventral view C lateral view D prolateral view E ventral view F retrolateral view (teal green dashed lines represent the contours of DTA, RTA and VTA). Abbreviations: aSDL = ascending part of sperm duct loop; Con = conductor; Cy = cymbium; dSDL = descending part of sperm duct loop; DTA = dorsal tibial apophysis; Em = embolus; EmB = embolic base; EmT = embolic tip; IR = intertegular retinaculum; RTA = retrolateral tibial apophysis; SD = sperm duct; Te = tegulum; VTA = ventral tibial apophysis. Scale bars: 1 mm (equal for A C); 0.5 mm (equal for D F)." figureDoi="10.3897/zookeys.1240.149456.figure12" httpUri="https://binary.pensoft.net/fig/1348770">12 D F</figureCitation>
, epigyne as in Fig.
<figureCitation id="0D8C0ABA18688F12AC5D089B9EEF324F" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Philodromus paiki, female, habitus (A C) and epigyne (D, E) A dorsal view B ventral view C lateral view D ventral view E dorsal view. Abbreviations: CD = copulatory duct; EG = epigynal groove; FD = fertilisation duct; MS = median septum; R = receptaculum; rMS = rim of median septum (teal green line); rSEF = rim of sclerotised epigynal fold (blue line); SEF = sclerotised epigynal fold. Scale bars: 1 mm (equal for A C); 0.2 mm (equal for D, E)." figureDoi="10.3897/zookeys.1240.149456.figure13" httpUri="https://binary.pensoft.net/fig/1348771">13 D, E</figureCitation>
.
</paragraph>
<caption id="0D7034BBCDBBA43770CB919BFFF4E92E" ID-DOI="10.3897/zookeys.1240.149456.figure12" ID-arpha="7BBC9021-FFC3-5C32-8634-3FD5B5696CE5" httpUri="https://binary.pensoft.net/fig/1348770" startId="F12">
<paragraph id="0576DE3C29466337AE787C88C839D9B2">
<label id="7C6A905278423ECA95B3499A0C31308E">Figure 12.</label>
</paragraph>
<paragraph id="EDC2CC7E815A8F358B301C5834C1268E">
<taxonomicName id="2D014B9E78046C6AE2ABF3A86B8E58B6" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="paiki">
<emphasis id="51824CDC89390689C4E558FC00A220A3" italics="true">Philodromus paiki</emphasis>
</taxonomicName>
, male, habitus (
<emphasis id="CD2B88EABA88C2594EBA79AC347CA89F" bold="true">A C</emphasis>
) and left palp (
<emphasis id="4BD4C3D563ECB29FE3DEB9B3BEBD532A" bold="true">D F</emphasis>
)
<emphasis id="C9FA660A4E4E88A90FCF21986444FC1D" bold="true">A</emphasis>
dorsal view
<emphasis id="4644F1F5434A2E8618046C757EE527C4" bold="true">B</emphasis>
ventral view
<emphasis id="D6EFFBF01529684E7C59B29E40DDB24E" bold="true">C</emphasis>
lateral view
<emphasis id="AFCDA5CECB3AED60C057CFF3CB16F338" bold="true">D</emphasis>
prolateral view
<emphasis id="24511187F9F459ED0FCECC83B9202BA0" bold="true">E</emphasis>
ventral view
<emphasis id="7E3AAE7821B43CE49CA4E93A7FD5AFF5" bold="true">F</emphasis>
retrolateral view (teal green dashed lines represent the contours of
<abbrev id="ABBRID0EBXAI" xlink_title="dorsal tibial apophysis">DTA</abbrev>
,
<abbrev id="ABBRID0EFXAI" xlink_title="retrolateral tibial apophysis">RTA</abbrev>
and
<abbrev id="ABBRID0EJXAI" xlink_title="ventral tibial apophysis">VTA</abbrev>
). Abbreviations:
<abbrev id="ABBRID0ENXAI" xlink_title="ascending part of sperm duct loop">aSDL</abbrev>
= ascending part of sperm duct loop;
<abbrev id="ABBRID0ERXAI" xlink_title="conductor">Con</abbrev>
= conductor;
<abbrev id="ABBRID0EVXAI" xlink_title="cymbium">Cy</abbrev>
= cymbium;
<abbrev id="ABBRID0EZXAI" xlink_title="descending part of sperm duct loop">dSDL</abbrev>
= descending part of sperm duct loop;
<abbrev id="ABBRID0E4XAI" xlink_title="dorsal tibial apophysis">DTA</abbrev>
= dorsal tibial apophysis;
<abbrev id="ABBRID0EBYAI" xlink_title="embolus">Em</abbrev>
= embolus;
<abbrev id="ABBRID0EFYAI" xlink_title="embolic base">EmB</abbrev>
= embolic base;
<abbrev id="ABBRID0EJYAI" xlink_title="embolic tip">EmT</abbrev>
= embolic tip;
<abbrev id="ABBRID0ENYAI" xlink_title="intertegular retinaculum">IR</abbrev>
= intertegular retinaculum;
<abbrev id="ABBRID0ERYAI" xlink_title="retrolateral tibial apophysis">RTA</abbrev>
= retrolateral tibial apophysis;
<abbrev id="ABBRID0EVYAI" xlink_title="sperm duct">SD</abbrev>
= sperm duct;
<abbrev id="ABBRID0EZYAI" xlink_title="tegulum">Te</abbrev>
= tegulum;
<abbrev id="ABBRID0E4YAI" xlink_title="ventral tibial apophysis">VTA</abbrev>
= ventral tibial apophysis. Scale bars: 1 mm (equal for
<emphasis id="1A648B4F672BEBA547345F7D19575759" bold="true">A C</emphasis>
); 0.5 mm (equal for
<emphasis id="130C8018234E2555BF0860AE492003E7" bold="true">D F</emphasis>
).
</paragraph>
</caption>
<caption id="1D4F096456727D64CEA52A20BE808D07" ID-DOI="10.3897/zookeys.1240.149456.figure13" ID-arpha="8580CDDF-B85E-5D8B-9C97-8CB634ABBB37" httpUri="https://binary.pensoft.net/fig/1348771" startId="F13">
<paragraph id="82B97DDBB6A4A7127A894183D62658C6">
<label id="CAE2C2071AE5E9B606B7C7D9EBE09E57">Figure 13.</label>
</paragraph>
<paragraph id="C3FD4E011A9174B005A51601C3A36D98">
<taxonomicName id="4616EDCCF7D952D14262665E1D6C224C" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="paiki">
<emphasis id="A574A76383217D833318A0BFDC9DE925" italics="true">Philodromus paiki</emphasis>
</taxonomicName>
, female, habitus (
<emphasis id="48457BF30F125F7E07E345B1D807943D" bold="true">A C</emphasis>
) and epigyne (
<emphasis id="C0BB4DAC9BD7D436B76E173792F87766" bold="true">D, E</emphasis>
)
<emphasis id="E938E95E46562E348E8F1021643A1B5E" bold="true">A</emphasis>
dorsal view
<emphasis id="463E34549C690DA5FA571B76EA869F61" bold="true">B</emphasis>
ventral view
<emphasis id="26C246680F680B7E6B0B4DBAA0C7314B" bold="true">C</emphasis>
lateral view
<emphasis id="46A8648A9B42B64E67F262AF5AD78F78" bold="true">D</emphasis>
ventral view
<emphasis id="812349C776D1CB0F7DB16A3386C3384C" bold="true">E</emphasis>
dorsal view. Abbreviations:
<abbrev id="ABBRID0EL1AI" xlink_title="copulatory duct">CD</abbrev>
= copulatory duct;
<abbrev id="ABBRID0EP1AI" xlink_title="epigynal groove">EG</abbrev>
= epigynal groove;
<abbrev id="ABBRID0ET1AI" xlink_title="fertilisation duct">FD</abbrev>
= fertilisation duct;
<abbrev id="ABBRID0EX1AI" xlink_title="median septum">MS</abbrev>
= median septum; R = receptaculum;
<abbrev id="ABBRID0E21AI" xlink_title="rim of median septum">rMS</abbrev>
= rim of median septum (teal green line);
<abbrev id="ABBRID0E61AI" xlink_title="rim of sclerotised epigynal fold">rSEF</abbrev>
= rim of sclerotised epigynal fold (blue line);
<abbrev id="ABBRID0ED2AI" xlink_title="sclerotised epigynal fold">SEF</abbrev>
= sclerotised epigynal fold. Scale bars: 1 mm (equal for
<emphasis id="0E169F1A666499601AC3C0CE59449C23" bold="true">A C</emphasis>
); 0.2 mm (equal for
<emphasis id="F243F1BEDB0433CFC338676300C9654F" bold="true">D, E</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EQ2AI" type="distribution">
<paragraph id="4881A98D641F79E53890A99C2C403D59">
<heading id="0A0F54B5EA5406E46DBAEB4C334ED0C3" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="A8F24612B624C08D1D34FD552E08792C">
<collectingCountry id="043ECCAE6A96C8D6FF3F2D4A800BE561" name="China">China</collectingCountry>
(
<collectingRegion id="C36E482C83F5E043FB87FBF243498D6D" country="China" name="Guizhou">Guizhou</collectingRegion>
,
<collectingRegion id="DA62CE5E094C8777860CCD34A75B1882" country="China" name="Hubei">Hubei</collectingRegion>
; Fig.
<figureCitation id="3D66C3A60C004D042741CC631E9BDC04" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Distribution records of the philodromid species in Guizhou Province (A) and Hubei Province (B)." figureDoi="10.3897/zookeys.1240.149456.figure23" httpUri="https://binary.pensoft.net/fig/1348781">23</figureCitation>
),
<collectingCountry id="8245024A17C5B69AFBB6A53AF15B6A4B" name="South Korea">Korea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E12AI" type="comments">
<paragraph id="01D9CC6AE346463B172A6C5011375A96">
<heading id="856511756F867D38A27E1BAB73E7F5ED" reason="title">Comments.</heading>
</paragraph>
<paragraph id="BE2A7252DEB0B02651E6932F911F4F3D">
This species is easily confused with
<taxonomicName id="5F356DCB9ACB05735F235D446691EAEE" authorityName="Simon" authorityYear="1895" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="spinitarsis">
<emphasis id="5BA1099264570533873835EAE77E2D53" italics="true">P. spinitarsis</emphasis>
</taxonomicName>
due to its similar genital morphology.
<bibRefCitation id="AB9C4395ED7704BD8A2CB152A3656544" DOI="10.11626/KJEB.2023.41.4.497" author="Jang CM &amp; Lee SY &amp; Yoo JS &amp; Kim ST" refId="B24" refString="Jang CM, Lee SY, Yoo JS, Kim ST (2024 a) Description of Philodromus paiki sp. nov. and Philodromus spinitarsis Simon, 1895 (Araneae: Philodromidae) from Korea. Korean Journal of Environmental Biology 41 (4) [2023]: 497504. [publ. in January 2024] https://doi.org/10.11626/KJEB.2023.41.4.497">Jang et al. (2024 a</bibRefCitation>
) described it as a new species based on specimens from
<collectingCountry id="D7E96CC9A146E1C614FB503EA6F0037F" name="South Korea">Korea</collectingCountry>
and pointed out that previous records of
<taxonomicName id="28C043F17BB0B1CB6C077EFBA662F02E" authorityName="Simon" authorityYear="1895" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="spinitarsis">
<emphasis id="530DC98AF0B817A62AAFE39024A21D6F" italics="true">P. spinitarsis</emphasis>
</taxonomicName>
in
<collectingCountry id="ECB78B53E955FC919FE495D8996258F5" name="South Korea">Korea</collectingCountry>
in multiple studies were likely misidentifications, most of which should be attributed to
<taxonomicName id="30C4F0F702A69FEBE75782759A7BFBF4" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="paiki">
<emphasis id="8B2A279DF2F3D32639CEA37A4151124F" italics="true">P. paiki</emphasis>
</taxonomicName>
. It appears that many records of
<taxonomicName id="0B7E81B45893958209BE32F5F3D446DC" authorityName="Simon" authorityYear="1895" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="spinitarsis">
<emphasis id="31C425034923360595E29B448D734D5E" italics="true">P. spinitarsis</emphasis>
</taxonomicName>
in Chinese literature may also be misidentifications. However, since we have not examined the original specimens, our study is currently based on materials from Guizhou and Hubei provinces.
</paragraph>
<paragraph id="39C5CDF1B03EB6D927420BDDE73AA2FE">
<bibRefCitation id="6393A0AF6240A005A924A1BC2776E812" DOI="10.3969/j.issn.1005-9628.2022.02.009" author="Long FY &amp; Deng LJ &amp; Mao JL &amp; Yu H" etAl="et al." firstAuthor="Long" issue="2" journalOrPublisher="Acta Arachnologica Sinica" pagination="113-120" refId="B33" refString="Long FY, Deng LJ, Mao JL, Yu H (2022) A survey of genus Philodromus (Araneae: Philodromidae) from Guiyang, Guizhou, China, with description of a new species. Acta Arachnologica Sinica 31 (2): 113120. https://doi.org/10.3969/j.issn.1005-9628.2022.02.009" title="A survey of genus Philodromus (Araneae: Philodromidae) from Guiyang, Guizhou, China, with description of a new species." volume="31" year="2022">Long et al. (2022)</bibRefCitation>
reported
<taxonomicName id="06AEF9CDB3B6D5306E0BCC44153F11AA" authorityName="Simon" authorityYear="1895" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="spinitarsis">
<emphasis id="61D793B8DB8776B3AE4901A85455079C" italics="true">P. spinitarsis</emphasis>
</taxonomicName>
as a new record for
<collectingRegion id="87F96792181DCCBAB38533B1B607D15E" country="China" name="Guizhou">Guizhou Province</collectingRegion>
based on specimens from Guiyang City but did not provide diagnostic illustrations. Upon re-examining the specimens from
<bibRefCitation id="87C08E4D8F630A9235155104DBFBBAB4" DOI="10.3969/j.issn.1005-9628.2022.02.009" author="Long FY &amp; Deng LJ &amp; Mao JL &amp; Yu H" etAl="et al." firstAuthor="Long" issue="2" journalOrPublisher="Acta Arachnologica Sinica" pagination="113-120" refId="B33" refString="Long FY, Deng LJ, Mao JL, Yu H (2022) A survey of genus Philodromus (Araneae: Philodromidae) from Guiyang, Guizhou, China, with description of a new species. Acta Arachnologica Sinica 31 (2): 113120. https://doi.org/10.3969/j.issn.1005-9628.2022.02.009" title="A survey of genus Philodromus (Araneae: Philodromidae) from Guiyang, Guizhou, China, with description of a new species." volume="31" year="2022">Long et al. (2022)</bibRefCitation>
, we determined that they should all belong to
<taxonomicName id="F3CBBA43686F07CD6CC46709B70960B5" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="paiki">
<emphasis id="80B47988EE9678FC41CD392DF8BE20FE" italics="true">P. paiki</emphasis>
</taxonomicName>
. Therefore, there is currently no confirmed record of
<taxonomicName id="27F7BC30CA6361327EB50AE8C17F5AD1" authorityName="Simon" authorityYear="1895" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="spinitarsis">
<emphasis id="D2430CB5E7BD5AF4A665772A61788A8C" italics="true">P. spinitarsis</emphasis>
</taxonomicName>
in
<collectingRegion id="953758358704BD5FDA2841B15BB5A036" country="China" name="Guizhou">Guizhou</collectingRegion>
. The newly available specimens indicate that this species is also distributed in Fanjingshan National Nature Reserve,
<collectingRegion id="77BC305537A6A0F207657FA3C6C39480" country="China" name="Guizhou">Guizhou Province</collectingRegion>
(Fig.
<figureCitation id="C797246091B11ECCB179672C1186EE28" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Distribution records of the philodromid species in Guizhou Province (A) and Hubei Province (B)." figureDoi="10.3897/zookeys.1240.149456.figure23" httpUri="https://binary.pensoft.net/fig/1348781">23 A</figureCitation>
).
</paragraph>
<paragraph id="EF665223A435BF9ACA3E917C19AEB206">
Several studies have recorded the presence of
<taxonomicName id="4424F677D1F3CC0CC1356B3738A45669" authorityName="Simon" authorityYear="1895" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="spinitarsis">
<emphasis id="E6FD0A1C0A372F60A8001F63D1BED3A8" italics="true">P. spinitarsis</emphasis>
</taxonomicName>
in
<collectingRegion id="37582054E417C2923A0C5747BD0471FE" country="China" name="Hubei">Hubei</collectingRegion>
, such as
<bibRefCitation id="E2FD93B38A296E6B9C435D9C5D4C22F3" author="Zhao JZ" firstAuthor="Zhao" journalOrPublisher="Wuhan Publishing House, Wuhan, China" refId="B65" refString="Zhao JZ (1993) Spiders in the cotton fields in China. Wuhan Publishing House, Wuhan, China, 552 pp." title="Spiders in the cotton fields in China." year="1993">Zhao (1993)</bibRefCitation>
,
<bibRefCitation id="602ED7589FAEEF77066F76C5701F91DA" author="Song DX &amp; Zhu MS" firstAuthor="Song" journalOrPublisher="Science Press, Beijing" refId="B48" refString="Song DX, Zhu MS (1997) Fauna Sinica: Arachnida: Araneae: Thomisidae, Philodromidae. Science Press, Beijing, 259 pp." title="Fauna Sinica: Arachnida: Araneae: Thomisidae, Philodromidae." year="1997">Song and Zhu (1997)</bibRefCitation>
,
<bibRefCitation id="40E1D7085A731147A58F33A5A6638EBA" author="Song DX &amp; Zhu MS &amp; Chen J" etAl="et al." firstAuthor="Song" journalOrPublisher="Hebei Science and Technology Publishing House, Shijiazhuang" refId="B51" refString="Song DX, Zhu MS, Chen J (1999) The spiders of China. Hebei Science and Technology Publishing House, Shijiazhuang, 640 pp." title="The spiders of China." year="1999">Song et al. (1999)</bibRefCitation>
,
<bibRefCitation id="1497D1971096D5D3447151B8C9F12C2A" author="Zhu MS &amp; Zhang BS" firstAuthor="Zhu" journalOrPublisher="Science Press, Beijing" refId="B67" refString="Zhu MS, Zhang BS (2011) Spider Fauna of Henan: Arachnida: Araneae. Science Press, Beijing, 558 pp." title="Spider Fauna of Henan: Arachnida: Araneae." year="2011">Zhu and Zhang (2011)</bibRefCitation>
,
<bibRefCitation id="7BDC2FEF977B0CA574726456C12249CD" author="Yin CM &amp; Peng XJ &amp; Yan HM &amp; Bao YH &amp; Xu X &amp; Tang G &amp; Zhou QS &amp; Liu P" etAl="et al." firstAuthor="Yin" journalOrPublisher="Hunan Science and Technology Press, Changsha" refId="B62" refString="Yin CM, Peng XJ, Yan HM, Bao YH, Xu X, Tang G, Zhou QS, Liu P (2012) Fauna Hunan: Araneae in Hunan, China. Hunan Science and Technology Press, Changsha, 1590 pp." title="Fauna Hunan: Araneae in Hunan, China." year="2012">Yin et al. (2012)</bibRefCitation>
,
<bibRefCitation id="4A0BF7EB36AFB90A6F2D40FD0C1DA513" author="Zhang F &amp; Peng JY &amp; Zhang BS" etAl="et al." firstAuthor="Zhang" journalOrPublisher="Science Press, Beijing" refId="B64" refString="Zhang F, Peng JY, Zhang BS (2022) Spiders of Mt. Xiaowutai. Science Press, Beijing, 387 pp." title="Spiders of Mt. Xiaowutai." year="2022">Zhang et al. (2022)</bibRefCitation>
, but did not provide specific distribution points or coordinates. Additionally, the diagnostic illustrations provided are relatively crude and were not based on specimens from
<collectingRegion id="3CF1298F137A22E1BCD0DBF0518D9BC1" country="China" name="Hubei">Hubei</collectingRegion>
. Therefore, the reported distribution of
<taxonomicName id="641264FC41E76C8D5A4349F22CFCE970" authorityName="Simon" authorityYear="1895" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="spinitarsis">
<emphasis id="68ACE8EB5AC9A53187F67BD6155ABD65" italics="true">P. spinitarsis</emphasis>
</taxonomicName>
in
<collectingRegion id="DD94ECC86C751EB2873D9A2CD7C9CC6D" country="China" name="Hubei">Hubei</collectingRegion>
remains questionable. Our various field trips in
<collectingRegion id="39A11054A025AB569C8A42E30CF36563" country="China" name="Hubei">Hubei</collectingRegion>
have not yielded any
<taxonomicName id="BA6BAD4AC7F0E8BC6C5389009EF273FB" authorityName="Simon" authorityYear="1895" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="spinitarsis">
<emphasis id="9914682CFF72BF44411160826C1C2768" italics="true">P. spinitarsis</emphasis>
</taxonomicName>
specimens. In contrast, paired specimens of
<taxonomicName id="BD0F2D9EDB57D50D3B36B5E9A0B9E3E3" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="paiki">
<emphasis id="C041324EFB6D6E284FAC375428A65BB0" italics="true">P. paiki</emphasis>
</taxonomicName>
were obtained from Jiugongshan National Nature Reserve (Fig.
<figureCitation id="483B6F903980EBF9FE2FE08D143EB3F2" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Distribution records of the philodromid species in Guizhou Province (A) and Hubei Province (B)." figureDoi="10.3897/zookeys.1240.149456.figure23" httpUri="https://binary.pensoft.net/fig/1348781">23 B</figureCitation>
). This paper represents the first formal report of the distribution of
<taxonomicName id="ACA244C8867498B87419578F4610BD13" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="paiki">
<emphasis id="1675D3AFFD225F90EB7C4125534CFC51" italics="true">P. paiki</emphasis>
</taxonomicName>
in
<collectingRegion id="54BD2B5CAA7D2C098CCAA8C4C9E2D786" country="China" name="Hubei">Hubei Province</collectingRegion>
.
</paragraph>
</subSubSection>
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<taxonomicName id="E279284538171F36D3A8179332F30DBA" ID-CoL="C82X9" authority="Long &amp; Yu, 2022" authorityName="Long &amp; Yu" authorityYear="2022" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="guiyang">
<emphasis id="F9ECAE9E7B549E88714D61D09FEB4CA1" italics="true">Philodromus guiyang</emphasis>
<bibRefCitation id="F672FA02D6DD2C79E0846C681FF306C9" DOI="10.3969/j.issn.1005-9628.2022.02.009" author="Long FY &amp; Deng LJ &amp; Mao JL &amp; Yu H" issue="2" journalOrPublisher="Acta Arachnologica Sinica" pagination="113-120" refId="B33" refString="Long FY, Deng LJ, Mao JL, Yu H (2022) A survey of genus Philodromus (Araneae: Philodromidae) from Guiyang, Guizhou, China, with description of a new species. Acta Arachnologica Sinica 31 (2): 113-120. https://doi.org/10.3969/j.issn.1005-9628.2022.02.009" title="A survey of genus Philodromus (Araneae: Philodromidae) from Guiyang, Guizhou, China, with description of a new species." volume="31" year="2022">Long &amp; Yu, 2022</bibRefCitation>
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<paragraph id="FD049133638C9ADFCE35C9012468698A">
<figureCitation id="76E4ADB97610ABC7314268E59DA84AA2" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Sample collection locations for this project. Hubei Province 1 Jiugongshan National Nature Reserve 2 Hubei University of Science and Technology 3 Jiuling Mountain 4 Qizimei Mountain National Nature Reserve; Guizhou Province 5 Fanjingshan National Nature Reserve 6 Mayanghe National Nature Reserve 7 Xishui National Nature Reserve 8 Guiyang City, Kaiyang County 9 Guiyang City, Wudang District 10 Guiyang City, Yunyan District 11 Guiyang City, Huaxi District." figureDoi="10.3897/zookeys.1240.149456.figure1" httpUri="https://binary.pensoft.net/fig/1348759">Figs 1</figureCitation>
,
<figureCitation id="A2DF9864F2B9D5810D775A414B6E58F5" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Living specimens of some Philodromidae species treated in this paper A, B Philodromus auricomus (♂ ♀) C, D Philodromus guiyang (♂) E, F Philodromus subaureolus (♂ ♀). Photographs by Q. Lu (Shenzhen)." figureDoi="10.3897/zookeys.1240.149456.figure2" httpUri="https://binary.pensoft.net/fig/1348760">2 C, D</figureCitation>
,
<figureCitation id="560A274C8119F1E523FE6AB2FEC7403F" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Philodromus guiyang, female (A C) and male (D F), habitus A, D dorsal view B, E ventral view C, F lateral view. Scale bars: 1 mm (equal for A C, equal for D F)." figureDoi="10.3897/zookeys.1240.149456.figure7" httpUri="https://binary.pensoft.net/fig/1348765">7</figureCitation>
,
<figureCitation id="CED8D4430D3BF85AFAF34ADE1D576162" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Philodromus guiyang, female (A, C G) and male (B), frontal view of carapace (A, B) and epigyne (C G) A female B male C intact, ventral view D macerated, ventral view E macerated, dorsal view F macerated and embedded in arabic gum, ventral view G macerated and embedded in Arabic gum, dorsal view. Abbreviations: A = atrium; CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; GM = glandular mound; MS = median septum; R = receptaculum; rMS = rim of median septum (teal green line); rSEF = rim of sclerotised epigynal fold (blue line); SEF = sclerotised epigynal fold. Scale bars: 0.5 mm (A, B); 0.2 mm (equal for C G)." figureDoi="10.3897/zookeys.1240.149456.figure8" httpUri="https://binary.pensoft.net/fig/1348766">8</figureCitation>
,
<figureCitation id="7BB8EE3FAF4E0E2381DD6614F99AF776" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Philodromus guiyang, male, left palp A ventral view B dorsal view C prolateral view D retrolateral view. Abbreviations: aSDL = ascending part of sperm duct loop; Con = conductor; CP = cymbial process; Cy = cymbium; dRTA = dorsal branch of RTA; dSDL = descending part of sperm duct loop; Em = embolus; EmB = embolic base; EmT = embolic tip; IR = intertegular retinaculum; pCon = prolateral part of conductor; RTA = retrolateral tibial apophysis; SD = sperm duct; tCon = tip of conductor; Te = tegulum; vRTA = ventral branch of RTA; VTA = ventral tibial apophysis. Scale bar: 0.2 mm (equal for A D)." figureDoi="10.3897/zookeys.1240.149456.figure9" httpUri="https://binary.pensoft.net/fig/1348767">9</figureCitation>
,
<figureCitation id="7A888DE04379DF17482B276DAEA806EF" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Distribution records of the philodromid species in Guizhou Province (A) and Hubei Province (B)." figureDoi="10.3897/zookeys.1240.149456.figure23" httpUri="https://binary.pensoft.net/fig/1348781">23 A</figureCitation>
</paragraph>
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<paragraph id="F58794F45CAFD790851C6184F4F36EE7">
<treatmentCitationGroup id="7566A79AAC60B86D62ADA2CA225DFA4B">
<treatmentCitation id="2F5028B381CD16F4BE23A6BE74371327">
<taxonomicName id="1F6CD2900C7053CD31950EEBD162F603" ID-CoL="C82X9" authority="Long &amp; Yu" authorityName="Long &amp; Yu" authorityYear="2022" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="guiyang">
<emphasis id="2F53BA1700F36ED009641E233B574695" italics="true">Philodromus guiyang</emphasis>
Long &amp; Yu
</taxonomicName>
, in
<bibRefCitation id="90927FDFB2534E1A3FFC6CDE8BB3F178" DOI="10.3969/j.issn.1005-9628.2022.02.009" author="Long FY &amp; Deng LJ &amp; Mao JL &amp; Yu H" etAl="et al." firstAuthor="Long" issue="2" journalOrPublisher="Acta Arachnologica Sinica" page="118" pagination="113-120" refId="B33" refString="Long FY, Deng LJ, Mao JL, Yu H (2022) A survey of genus Philodromus (Araneae: Philodromidae) from Guiyang, Guizhou, China, with description of a new species. Acta Arachnologica Sinica 31 (2): 113120. https://doi.org/10.3969/j.issn.1005-9628.2022.02.009" title="A survey of genus Philodromus (Araneae: Philodromidae) from Guiyang, Guizhou, China, with description of a new species." volume="31" year="2022">Long et al. 2022: 118</bibRefCitation>
</treatmentCitation>
, figs 2 A D, 3 A D (
<specimenCount id="7FE62074F5A344C56DD52C0D0D72D110" count="1" type="male"></specimenCount>
);
<treatmentCitation id="B4A84CA93B47756734902C98ED0F3793">
<bibRefCitation id="E1B6884C62CCF863AFCD7423544A92D6" DOI="10.3897/zookeys.1221.137930" author="Wang ZJ &amp; Yao YB &amp; Tang ZY &amp; Li WH &amp; Liu KK &amp; Xu X" etAl="et al." firstAuthor="Wang" journalOrPublisher="ZooKeys" page="281" pagination="279-296" refId="B55" refString="Wang ZJ, Yao YB, Tang ZY, Li WH, Liu KK, Xu X (2024) A new genus, Sinodromus gen. nov., with two new species and the first description of the female of Philodromus guiyang Long &amp; Yu, 2022 (Arachnida, Araneae, Philodromidae) from China. ZooKeys 296: 279296. https://doi.org/10.3897/zookeys.1221.137930" title="A new genus, Sinodromus gen. nov., with two new species and the first description of the female of Philodromus guiyang Long &amp; Yu, 2022 (Arachnida, Araneae, Philodromidae) from China." volume="296" year="2024">Wang et al. 2024: 281</bibRefCitation>
</treatmentCitation>
, figs 1 A D, 2 A E, 8 A, 9 A, B (
<specimenCount id="8B1B04DBC9F5E71E728394B645F83AD9" count="1" type="male"></specimenCount>
<specimenCount id="D3D0FC75330B3F3560A8FBA44BECB49E" count="1" type="female"></specimenCount>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDCBG" type="type material">
<paragraph id="BD6ED8A42579AA592FB4763F3D017F66">
<heading id="B746A8C6179C53709435458596416901" reason="title">
<typeStatus id="0739D70C54D4ED7C67778647C769C6C2">Type</typeStatus>
material examined.
</heading>
</paragraph>
<paragraph id="B07B645C86906DD9B46EB4806AE9DF94">
<materialsCitation id="031B6AE3FB1409CF59BA86B77ED7A381" collectingDate="2022-06-10" collectionCode="YHGY" collectorName="H. Yu &amp; Q. Lu" country="China" county="Kaiyang County" elevation="812" latitude="26.93" location="Longgang Town" longLatPrecision="744" longitude="107.07" municipality="Guiyang City" specimenCode="YHGY 213" specimenCount="1" specimenCount-male="1" stateProvince="Guizhou" typeStatus="holotype">
<emphasis id="D102207BF19B42FF37FAD1CCE97EE1CF" bold="true">
<collectingCountry id="F266415A4DE0DE1D66676A18C3214861" name="China">China</collectingCountry>
</emphasis>
:
<collectingRegion id="04D30CB25D293DF7BF4AE276F02A23B8" country="China" name="Guizhou">Guizhou Province</collectingRegion>
:
<emphasis id="5C2355F922F0822E4242265D18F009A9" bold="true" italics="true">
<typeStatus id="8093BC818A56B60D97E9F95F93452274" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="A4926ACDF33DF059EEE7BA2A3535FD94" count="1" type="male"></specimenCount>
(
<specimenCode id="20437001FC927E01EFED7D1CFD889692" collectionCode="YHGY">YHGY 213</specimenCode>
used for sequencing, GenBank accession numbers in Table
<tableCitation id="FD959EDC1F8BC4DEA4EE4B956E4A0200" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Voucher specimen information." httpUri="http://table.plazi.org/id/12F54BE0BDE6512C62E0465BE733F517" tableUuid="12F54BE0BDE6512C62E0465BE733F517">1</tableCitation>
),
<collectingMunicipality id="60F6B48798678348B99253E3F500CCC8">Guiyang City</collectingMunicipality>
,
<collectingCounty id="9DEABE6B242AB8AF66C957E6345C3127">Kaiyang County</collectingCounty>
,
<location id="D61087479D66BE25C4680308EF549CC5" LSID="urn:lsid:plazi:treatment:5C66494BBC5E5EEAA427896890BAF57C:D61087479D66BE25C4680308EF549CC5" country="China" county="Kaiyang County" latitude="26.93" longLatPrecision="744" longitude="107.07" municipality="Guiyang City" name="Longgang Town" stateProvince="Guizhou">Longgang Town</location>
,
<location id="353D06DA460C55368C7A6F13EB0BA860" LSID="urn:lsid:plazi:treatment:5C66494BBC5E5EEAA427896890BAF57C:353D06DA460C55368C7A6F13EB0BA860" country="China" county="Kaiyang County" latitude="26.93" longLatPrecision="744" longitude="107.07" municipality="Guiyang City" name="Pingshan Village" stateProvince="Guizhou">Pingshan Village</location>
,
<location id="91D0E0DDEA4651AAE0A1AAA7C9912D0C" LSID="urn:lsid:plazi:treatment:5C66494BBC5E5EEAA427896890BAF57C:91D0E0DDEA4651AAE0A1AAA7C9912D0C" country="China" county="Kaiyang County" latitude="26.93" longLatPrecision="744" longitude="107.07" municipality="Guiyang City" name="Zijiang Rift Scenic Area" stateProvince="Guizhou">Zijiang Rift Scenic Area</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EVCBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[107.070000,26.930000]}">
<geoCoordinate id="0BD27935C8DADDF8D9C9B17BE032FBA9" degrees="26.93" direction="north" orientation="latitude" precision="555" value="26.93">26.93 ° N</geoCoordinate>
,
<geoCoordinate id="611EAE5268C8EB2188C1F476247839B7" degrees="107.07" direction="east" orientation="longitude" precision="555" value="107.07">107.07 ° E</geoCoordinate>
</named-content>
;
<elevation id="BE77D1B9A0736716316796CDF3402617" metricMagnitude="2" metricUnit="m" metricValue="8.12" unit="m" value="812.0">
<quantity id="F04193C43E5789E90DE24E53DA656A33" metricMagnitude="2" metricUnit="m" metricValue="8.12" unit="m" value="812.0">812 m</quantity>
a. s. l.
</elevation>
;
<date id="760612F7196AA2BE55573DFDEA5A8BFB" value="2022-06-10">
<collectingDate id="5E23264DB7B08B090CA26B742E40761B" value="2022-06-10">10 June 2022</collectingDate>
</date>
;
<collectorName id="6E0F3638E44A7886CEE3F16A6241D479">H. Yu</collectorName>
&amp;
<collectorName id="B4545D9CAF9D5FA6948203986F0F7C82">Q. Lu</collectorName>
leg.
</materialsCitation>
<materialsCitation id="B8C50409580965659330FC4BB2C1E451" collectingDate="2022-06-10" collectionCode="YHGY" collectorName="H. Yu &amp; Q. Lu" country="China" county="Kaiyang County" elevation="812" latitude="26.93" location="Longgang Town" longLatPrecision="744" longitude="107.07" municipality="Guiyang City" specimenCount="1" specimenCount-male="1" stateProvince="Paratype" typeStatus="holotype">
<emphasis id="BBCDED51BF9D2A89B56D5E1BE3ADC4AE" bold="true" italics="true">
<typeStatus id="A8BD65F7CD9A3049BCF4B37B5029F093" type="paratype">
<collectingRegion id="1B6E7B7237A84FC22B5165FC808D2AB2">Paratype</collectingRegion>
</typeStatus>
</emphasis>
<specimenCount id="B8EA63C8DF85AC1A1FE9F357270A58E9" count="1" type="male">1 ♂</specimenCount>
, the same data as the holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4CBG" type="material">
<paragraph id="242E381BA1CD9AC622BDF0DF7F5FAD9F">
<heading id="6B3233C162BF1189FFE14215D13607D1" reason="title">Other material examined.</heading>
</paragraph>
<paragraph id="741F394A2F235B9C265C8AE6F5BE8CD8">
<materialsCitation id="496EDA2BBF2ED259EDF8498815DF6376" collectingDate="2023-08-08" collectionCode="MYHPHI" collectorName="Y. Zhou" country="China" county="Yanhe County" elevation="1194" latitude="28.69" location="Mayanghe National Nature Reserve" longLatPrecision="738" longitude="108.16" municipality="Tongren City" specimenCode="MYHPHI 001" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Guizhou">
<emphasis id="FFFF1FF08D67BBB98D6972AE3079D5BE" bold="true">
<collectingCountry id="CA1525A6745F1F4AF96F8FF3CFC17CEA" name="China">China</collectingCountry>
</emphasis>
:
<collectingRegion id="3CC237A6097D3F11F717151835768C65" country="China" name="Guizhou">Guizhou Province</collectingRegion>
: •
<specimenCount id="7EF85B315F6E9B3BEAE531B969B1D977" count="1" type="male">1 ♂</specimenCount>
,
<specimenCount id="3F1A680E126CDAA1BE36A0B09946D855" count="1" type="female">1 ♀</specimenCount>
(
<specimenCount id="0E070F4E7BDBC4C8FCFCDF1CB46888C3" count="1" type="female">1 ♀</specimenCount>
,
<specimenCode id="59FE4A134B72EBDE1BBD42DDFC7A6DCD" collectionCode="MYHPHI">MYHPHI 001</specimenCode>
used for sequencing, GenBank accession numbers in Table
<tableCitation id="FA7E116F89727EEDA11C3B36871D97AC" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Voucher specimen information." httpUri="http://table.plazi.org/id/12F54BE0BDE6512C62E0465BE733F517" tableUuid="12F54BE0BDE6512C62E0465BE733F517">1</tableCitation>
),
<collectingMunicipality id="8039C374E5C33D22CB496EF39A6F698D">Tongren City</collectingMunicipality>
,
<location id="ED23CA1901B0A5AB17ED17956477AB03" LSID="urn:lsid:plazi:treatment:5C66494BBC5E5EEAA427896890BAF57C:ED23CA1901B0A5AB17ED17956477AB03" country="China" county="Yanhe County" latitude="28.69" longLatPrecision="738" longitude="108.16" municipality="Tongren City" name="Mayanghe National Nature Reserve" stateProvince="Guizhou">Mayanghe National Nature Reserve</location>
,
<collectingCounty id="139C86F74956B80BB310535438C5E622">Yanhe County</collectingCounty>
,
<location id="8CB1C47DE802E492FE290BB0E65D25BB" LSID="urn:lsid:plazi:treatment:5C66494BBC5E5EEAA427896890BAF57C:8CB1C47DE802E492FE290BB0E65D25BB" country="China" county="Yanhe County" latitude="28.69" longLatPrecision="738" longitude="108.16" municipality="Tongren City" name="Huangtu Town" stateProvince="Guizhou">Huangtu Town</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EMDBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[108.160000,28.690000]}">
<geoCoordinate id="D90DE3B5996115AB6C9A1065F93DA391" degrees="28.69" direction="north" orientation="latitude" precision="555" value="28.69">28.69 ° N</geoCoordinate>
,
<geoCoordinate id="E6ACB267E69F68F5475493F451E3F7D5" degrees="108.16" direction="east" orientation="longitude" precision="555" value="108.16">108.16 ° E</geoCoordinate>
</named-content>
;
<elevation id="71C536EAE2A8CB18173189E4099ABD58" metricMagnitude="3" metricUnit="m" metricValue="1.194" unit="m" value="1194.0">
<quantity id="AC3B74C16911447C8398AEA7F6ADA791" metricMagnitude="3" metricUnit="m" metricValue="1.194" unit="m" value="1194.0">1194 m</quantity>
a. s. l.
</elevation>
;
<date id="8A39352D112749F9B337DF7C5997E700" value="2023-08-08">
<collectingDate id="2C29FBAFA965E2523EF23316F25CDA78" value="2023-08-08">8 August 2023</collectingDate>
</date>
;
<collectorName id="9E3D01A742950B16B35D2FF70A74F617">Y. Zhou</collectorName>
et al. leg
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERDBG" type="diagnosis">
<paragraph id="BF1F696A7D2A811C9C2863577C686146">
<heading id="1F79C6567394C7539F20FD5E5022E245" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="E42781E0B5376B92593264F751B20F0C">
Females resemble those of
<taxonomicName id="1A161045126F10BBBEE2C263BEA6444D" authorityName="Bosenberg &amp; Strand" authorityYear="1906" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="subaureolus">
<emphasis id="6C53EC44AD89539D7DA0A25FB91FA7B5" italics="true">P. subaureolus</emphasis>
</taxonomicName>
in having the similarly bell-shaped
<abbrev id="ABBRID0ECEBG" xlink_title="median septum">MS</abbrev>
which is not delimited to
<abbrev id="ABBRID0EGEBG" xlink_title="sclerotised epigynal fold">SEF</abbrev>
, but can be recognised by: (1) A comma-shaped (vs elongate-oval, nearly funnel-shaped) (cf. Fig.
<figureCitation id="03D1ABA58DD217EFC727AB6E6FE09BC1" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Philodromus guiyang, female (A, C G) and male (B), frontal view of carapace (A, B) and epigyne (C G) A female B male C intact, ventral view D macerated, ventral view E macerated, dorsal view F macerated and embedded in arabic gum, ventral view G macerated and embedded in Arabic gum, dorsal view. Abbreviations: A = atrium; CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; GM = glandular mound; MS = median septum; R = receptaculum; rMS = rim of median septum (teal green line); rSEF = rim of sclerotised epigynal fold (blue line); SEF = sclerotised epigynal fold. Scale bars: 0.5 mm (A, B); 0.2 mm (equal for C G)." figureDoi="10.3897/zookeys.1240.149456.figure8" httpUri="https://binary.pensoft.net/fig/1348766">8 C, D, F</figureCitation>
and Fig.
<figureCitation id="B8EBF475EB2C851783B1047AFFF70D2C" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Philodromus subaureolus, female, habitus (A C) and epigyne (D, E) A dorsal view B ventral view C lateral view D ventral view E dorsal view. Abbreviations: A = atrium; cCD = course of copulatory duct (red line); CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; GM = glandular mound; MS = median septum; R = receptaculum; rMS = rim of median septum (teal green line); rSEF = rim of sclerotised epigynal fold (blue line); SEF = sclerotised epigynal fold. Scale bars: 1 mm (equal for A C); 0.2 mm (equal for D, E)." figureDoi="10.3897/zookeys.1240.149456.figure11" httpUri="https://binary.pensoft.net/fig/1348769">11 D</figureCitation>
); (2) anterior keel of
<abbrev id="ABBRID0ESEBG" xlink_title="median septum">MS</abbrev>
relatively wider, ~ 1 / 51 / 4 epigyne width (vs distinctly narrower, ~ 1 / 101 / 8 epigyne width) (cf. Fig.
<figureCitation id="2907E78C8C65507B4C3E41F3BA8204A0" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Philodromus guiyang, female (A, C G) and male (B), frontal view of carapace (A, B) and epigyne (C G) A female B male C intact, ventral view D macerated, ventral view E macerated, dorsal view F macerated and embedded in arabic gum, ventral view G macerated and embedded in Arabic gum, dorsal view. Abbreviations: A = atrium; CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; GM = glandular mound; MS = median septum; R = receptaculum; rMS = rim of median septum (teal green line); rSEF = rim of sclerotised epigynal fold (blue line); SEF = sclerotised epigynal fold. Scale bars: 0.5 mm (A, B); 0.2 mm (equal for C G)." figureDoi="10.3897/zookeys.1240.149456.figure8" httpUri="https://binary.pensoft.net/fig/1348766">8 D</figureCitation>
and Fig.
<figureCitation id="5B3EA19BD2DCBC7201D34DB7262030FD" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Philodromus subaureolus, female, habitus (A C) and epigyne (D, E) A dorsal view B ventral view C lateral view D ventral view E dorsal view. Abbreviations: A = atrium; cCD = course of copulatory duct (red line); CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; GM = glandular mound; MS = median septum; R = receptaculum; rMS = rim of median septum (teal green line); rSEF = rim of sclerotised epigynal fold (blue line); SEF = sclerotised epigynal fold. Scale bars: 1 mm (equal for A C); 0.2 mm (equal for D, E)." figureDoi="10.3897/zookeys.1240.149456.figure11" httpUri="https://binary.pensoft.net/fig/1348769">11 D</figureCitation>
); (3)
<abbrev id="ABBRID0E5EBG" xlink_title="copulatory duct">
<collectionCode id="529CC3BC7BF2377E59CF87528C502B04">CD</collectionCode>
</abbrev>
heavily sclerotised, not looped, distinctly thick and short, almost as thick as
<collectionCode id="28D03ACF0384C6F4351B37BCBA836664" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
, length ~ 1 / 2 epigyne (vs weakly sclerotised, distinctly thinner and longer, with a long
<abbrev id="ABBRID0ECFBG" xlink_title="course of copulatory duct">cCD</abbrev>
forming two loops before entering
<collectionCode id="40CEDE175022D355687B87A58F29A609" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
, their thickness no more than 1 / 2 the diameter of
<collectionCode id="2008182564DC1F8A5ABFCB109A307429" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
, length longer than epigyne) (cf. Fig.
<figureCitation id="7009D3BC571FDA60DA9499248BD86FE8" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Philodromus guiyang, female (A, C G) and male (B), frontal view of carapace (A, B) and epigyne (C G) A female B male C intact, ventral view D macerated, ventral view E macerated, dorsal view F macerated and embedded in arabic gum, ventral view G macerated and embedded in Arabic gum, dorsal view. Abbreviations: A = atrium; CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; GM = glandular mound; MS = median septum; R = receptaculum; rMS = rim of median septum (teal green line); rSEF = rim of sclerotised epigynal fold (blue line); SEF = sclerotised epigynal fold. Scale bars: 0.5 mm (A, B); 0.2 mm (equal for C G)." figureDoi="10.3897/zookeys.1240.149456.figure8" httpUri="https://binary.pensoft.net/fig/1348766">8 E, G</figureCitation>
and Fig.
<figureCitation id="211A566D1D1BDEEE4B70BB5E985A6D18" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Philodromus subaureolus, female, habitus (A C) and epigyne (D, E) A dorsal view B ventral view C lateral view D ventral view E dorsal view. Abbreviations: A = atrium; cCD = course of copulatory duct (red line); CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; GM = glandular mound; MS = median septum; R = receptaculum; rMS = rim of median septum (teal green line); rSEF = rim of sclerotised epigynal fold (blue line); SEF = sclerotised epigynal fold. Scale bars: 1 mm (equal for A C); 0.2 mm (equal for D, E)." figureDoi="10.3897/zookeys.1240.149456.figure11" httpUri="https://binary.pensoft.net/fig/1348769">11 E</figureCitation>
); (4)
<collectionCode id="5C88D85DB331416C0E3A8DE890E0AA7E" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
elongate-oval, anteriorly separated by ~ 0.5 × diameters, posteriorly separated by ~ 1.3 × diameters (vs nearly spherical, separated by ~ 1 diameter) (cf. Fig.
<figureCitation id="B9845FCE28FF7A2CAF329117FAC0B312" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Philodromus guiyang, female (A, C G) and male (B), frontal view of carapace (A, B) and epigyne (C G) A female B male C intact, ventral view D macerated, ventral view E macerated, dorsal view F macerated and embedded in arabic gum, ventral view G macerated and embedded in Arabic gum, dorsal view. Abbreviations: A = atrium; CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; GM = glandular mound; MS = median septum; R = receptaculum; rMS = rim of median septum (teal green line); rSEF = rim of sclerotised epigynal fold (blue line); SEF = sclerotised epigynal fold. Scale bars: 0.5 mm (A, B); 0.2 mm (equal for C G)." figureDoi="10.3897/zookeys.1240.149456.figure8" httpUri="https://binary.pensoft.net/fig/1348766">8 E, G</figureCitation>
and Fig.
<figureCitation id="775F9F48102711F1F3208145D06DE828" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Philodromus subaureolus, female, habitus (A C) and epigyne (D, E) A dorsal view B ventral view C lateral view D ventral view E dorsal view. Abbreviations: A = atrium; cCD = course of copulatory duct (red line); CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; GM = glandular mound; MS = median septum; R = receptaculum; rMS = rim of median septum (teal green line); rSEF = rim of sclerotised epigynal fold (blue line); SEF = sclerotised epigynal fold. Scale bars: 1 mm (equal for A C); 0.2 mm (equal for D, E)." figureDoi="10.3897/zookeys.1240.149456.figure11" httpUri="https://binary.pensoft.net/fig/1348769">11 E</figureCitation>
). Males of
<taxonomicName id="7BA9808E8302FCF258188A66B4A5E2AA" authorityName="Long &amp; Yu" authorityYear="2022" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="guiyang">
<emphasis id="227BB8FA608D89584365F3D5F42B6E42" italics="true">P. guiyang</emphasis>
</taxonomicName>
are also similar to those of
<taxonomicName id="30348FD5941019B523B28E2DB8C37577" authorityName="Bosenberg &amp; Strand" authorityYear="1906" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="subaureolus">
<emphasis id="13C1DE4FE1F92721CE7288E9E32A2C78" italics="true">P. subaureolus</emphasis>
</taxonomicName>
by the similar, blade-shaped
<abbrev id="ABBRID0EMGBG" xlink_title="ventral tibial apophysis">VTA</abbrev>
, lamellar
<abbrev id="ABBRID0EQGBG" xlink_title="retrolateral tibial apophysis">
<collectionCode id="2EAE43FBB96E4A749CE272C52AF16B9D">RTA</collectionCode>
</abbrev>
with a bifurcated tip, spine-shaped
<abbrev id="ABBRID0EUGBG" xlink_title="intertegular retinaculum">IR</abbrev>
, and the more or less S-shaped
<abbrev id="ABBRID0EYGBG" xlink_title="sperm duct loop">SDL</abbrev>
, but can be distinguished from the latter by: (1)
<abbrev id="ABBRID0E3GBG" xlink_title="embolus">Em</abbrev>
claw-shaped, distinctly shorter, originating at the 10 oclock position, terminating at the ~ 1 oclock position (vs filiform, distinctly longer, originating at 8 oclock position, terminating at ~ 1 oclock position) (cf. Fig.
<figureCitation id="32252ADE79A378DAA1FD60FA9B27DAF9" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Philodromus guiyang, male, left palp A ventral view B dorsal view C prolateral view D retrolateral view. Abbreviations: aSDL = ascending part of sperm duct loop; Con = conductor; CP = cymbial process; Cy = cymbium; dRTA = dorsal branch of RTA; dSDL = descending part of sperm duct loop; Em = embolus; EmB = embolic base; EmT = embolic tip; IR = intertegular retinaculum; pCon = prolateral part of conductor; RTA = retrolateral tibial apophysis; SD = sperm duct; tCon = tip of conductor; Te = tegulum; vRTA = ventral branch of RTA; VTA = ventral tibial apophysis. Scale bar: 0.2 mm (equal for A D)." figureDoi="10.3897/zookeys.1240.149456.figure9" httpUri="https://binary.pensoft.net/fig/1348767">9 A, C, D</figureCitation>
and Fig.
<figureCitation id="B267CABCD341EC23F6021296574B2CB2" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Philodromus subaureolus, male, habitus (A C) and left palp (D F) A dorsal view B ventral view C lateral view D prolateral view E ventral view F retrolateral view. Abbreviations: aSDL = ascending part of sperm duct loop; Con = conductor; Cy = cymbium; dRTA = dorsal branch of RTA; dSDL = descending part of sperm duct loop; Em = embolus; EmB = embolic base; EmT = embolic tip; IR = intertegular retinaculum; pCon = prolateral part of conductor; RTA = retrolateral tibial apophysis; SD = sperm duct; tCon = tip of conductor; Te = tegulum; vRTA = ventral branch of RTA; VTA = ventral tibial apophysis. Scale bars: 1 mm (equal for A C); 0.2 mm (equal for D F)." figureDoi="10.3897/zookeys.1240.149456.figure10" httpUri="https://binary.pensoft.net/fig/1348768">10 D F</figureCitation>
); (2)
<abbrev id="ABBRID0EIHBG" xlink_title="conductor">Con</abbrev>
distinctly shorter, enveloping the second half of
<abbrev id="ABBRID0EMHBG" xlink_title="embolus">Em</abbrev>
, the coverage ranges from the 11 oclock to the 1 oclock position (vs distinctly longer, almost enveloping the entire
<abbrev id="ABBRID0EQHBG" xlink_title="embolus">Em</abbrev>
, the coverage ranges from the 9 oclock to the 1 oclock position) (cf. Fig.
<figureCitation id="03BC4F5677BBCB4AF850352571AD69DC" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Philodromus guiyang, male, left palp A ventral view B dorsal view C prolateral view D retrolateral view. Abbreviations: aSDL = ascending part of sperm duct loop; Con = conductor; CP = cymbial process; Cy = cymbium; dRTA = dorsal branch of RTA; dSDL = descending part of sperm duct loop; Em = embolus; EmB = embolic base; EmT = embolic tip; IR = intertegular retinaculum; pCon = prolateral part of conductor; RTA = retrolateral tibial apophysis; SD = sperm duct; tCon = tip of conductor; Te = tegulum; vRTA = ventral branch of RTA; VTA = ventral tibial apophysis. Scale bar: 0.2 mm (equal for A D)." figureDoi="10.3897/zookeys.1240.149456.figure9" httpUri="https://binary.pensoft.net/fig/1348767">9 A, C, D</figureCitation>
and Fig.
<figureCitation id="101B89D6D3233F1C3ED6DADD5746FC42" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Philodromus subaureolus, male, habitus (A C) and left palp (D F) A dorsal view B ventral view C lateral view D prolateral view E ventral view F retrolateral view. Abbreviations: aSDL = ascending part of sperm duct loop; Con = conductor; Cy = cymbium; dRTA = dorsal branch of RTA; dSDL = descending part of sperm duct loop; Em = embolus; EmB = embolic base; EmT = embolic tip; IR = intertegular retinaculum; pCon = prolateral part of conductor; RTA = retrolateral tibial apophysis; SD = sperm duct; tCon = tip of conductor; Te = tegulum; vRTA = ventral branch of RTA; VTA = ventral tibial apophysis. Scale bars: 1 mm (equal for A C); 0.2 mm (equal for D F)." figureDoi="10.3897/zookeys.1240.149456.figure10" httpUri="https://binary.pensoft.net/fig/1348768">10 D F</figureCitation>
); (3) in retrolateral view, the middle section of
<abbrev id="ABBRID0E3HBG" xlink_title="retrolateral tibial apophysis">
<collectionCode id="FECB07D4324B0C70352E10B0148496ED">RTA</collectionCode>
</abbrev>
distinctly narrower than its base and tip (vs not distinctly narrowed, the entire
<abbrev id="ABBRID0EAIBG" xlink_title="retrolateral tibial apophysis">
<collectionCode id="750FF99386266D6A883755673B88AD90">RTA</collectionCode>
</abbrev>
is almost uniform in width) (cf. Fig.
<figureCitation id="6AB5124348730E558D17E1A7719925FE" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Philodromus guiyang, male, left palp A ventral view B dorsal view C prolateral view D retrolateral view. Abbreviations: aSDL = ascending part of sperm duct loop; Con = conductor; CP = cymbial process; Cy = cymbium; dRTA = dorsal branch of RTA; dSDL = descending part of sperm duct loop; Em = embolus; EmB = embolic base; EmT = embolic tip; IR = intertegular retinaculum; pCon = prolateral part of conductor; RTA = retrolateral tibial apophysis; SD = sperm duct; tCon = tip of conductor; Te = tegulum; vRTA = ventral branch of RTA; VTA = ventral tibial apophysis. Scale bar: 0.2 mm (equal for A D)." figureDoi="10.3897/zookeys.1240.149456.figure9" httpUri="https://binary.pensoft.net/fig/1348767">9 D</figureCitation>
and Fig.
<figureCitation id="B05DC9121FD9BD1F91DAE2DA4128F375" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Philodromus subaureolus, male, habitus (A C) and left palp (D F) A dorsal view B ventral view C lateral view D prolateral view E ventral view F retrolateral view. Abbreviations: aSDL = ascending part of sperm duct loop; Con = conductor; Cy = cymbium; dRTA = dorsal branch of RTA; dSDL = descending part of sperm duct loop; Em = embolus; EmB = embolic base; EmT = embolic tip; IR = intertegular retinaculum; pCon = prolateral part of conductor; RTA = retrolateral tibial apophysis; SD = sperm duct; tCon = tip of conductor; Te = tegulum; vRTA = ventral branch of RTA; VTA = ventral tibial apophysis. Scale bars: 1 mm (equal for A C); 0.2 mm (equal for D F)." figureDoi="10.3897/zookeys.1240.149456.figure10" httpUri="https://binary.pensoft.net/fig/1348768">10 F</figureCitation>
); and (4)
<abbrev id="ABBRID0EMIBG" xlink_title="ventral branch of RTA">vRTA</abbrev>
noticeably prominent, distinctly longer and sharper than
<abbrev id="ABBRID0EQIBG" xlink_title="dorsal branch of RTA">dRTA</abbrev>
(vs
<abbrev id="ABBRID0EUIBG" xlink_title="ventral branch of RTA">vRTA</abbrev>
small, both branches of
<abbrev id="ABBRID0EYIBG" xlink_title="retrolateral tibial apophysis">
<collectionCode id="7850C114C2F222892223AECFEC20C8B0">RTA</collectionCode>
</abbrev>
are similar in size and shape) (cf. Fig.
<figureCitation id="E6D9DBE23D0662DF63678F5B28249E37" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Philodromus guiyang, male, left palp A ventral view B dorsal view C prolateral view D retrolateral view. Abbreviations: aSDL = ascending part of sperm duct loop; Con = conductor; CP = cymbial process; Cy = cymbium; dRTA = dorsal branch of RTA; dSDL = descending part of sperm duct loop; Em = embolus; EmB = embolic base; EmT = embolic tip; IR = intertegular retinaculum; pCon = prolateral part of conductor; RTA = retrolateral tibial apophysis; SD = sperm duct; tCon = tip of conductor; Te = tegulum; vRTA = ventral branch of RTA; VTA = ventral tibial apophysis. Scale bar: 0.2 mm (equal for A D)." figureDoi="10.3897/zookeys.1240.149456.figure9" httpUri="https://binary.pensoft.net/fig/1348767">9 D</figureCitation>
and Fig.
<figureCitation id="D329C3AFAC5F2F0B24EEB9089E52F594" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Philodromus subaureolus, male, habitus (A C) and left palp (D F) A dorsal view B ventral view C lateral view D prolateral view E ventral view F retrolateral view. Abbreviations: aSDL = ascending part of sperm duct loop; Con = conductor; Cy = cymbium; dRTA = dorsal branch of RTA; dSDL = descending part of sperm duct loop; Em = embolus; EmB = embolic base; EmT = embolic tip; IR = intertegular retinaculum; pCon = prolateral part of conductor; RTA = retrolateral tibial apophysis; SD = sperm duct; tCon = tip of conductor; Te = tegulum; vRTA = ventral branch of RTA; VTA = ventral tibial apophysis. Scale bars: 1 mm (equal for A C); 0.2 mm (equal for D F)." figureDoi="10.3897/zookeys.1240.149456.figure10" httpUri="https://binary.pensoft.net/fig/1348768">10 F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEJBG" type="description">
<paragraph id="691FCD32AA4EBFFD26C3CE7214BED57E">
<heading id="D76D32E7580355B93A033FDB98729273" reason="title">Description.</heading>
</paragraph>
<paragraph id="F0C103FB31F87B304C7053C1FC372D2D">
<emphasis id="3EBDFF226E5E864B604F4D0D340E86B5" bold="true">Female.</emphasis>
Total length 4.05. Carapace 1.62 long, 1.57 wide. Abdomen 2.65 long, 1.82 wide.
<emphasis id="6EEE5808A2B9371B02507F2255E5DF84" bold="true" italics="true">Eye sizes and interdistances</emphasis>
:
<abbrev id="ABBRID0EPJBG" xlink_title="anterior median eyes">AME</abbrev>
0.07,
<abbrev id="ABBRID0ETJBG" xlink_title="anterior lateral eyes">ALE</abbrev>
0.07,
<abbrev id="ABBRID0EXJBG" xlink_title="posterior median eyes">PME</abbrev>
0.06,
<abbrev id="ABBRID0E2JBG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.08,
<abbrev id="ABBRID0E6JBG" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0EDKBG" xlink_title="anterior median eyes">AME</abbrev>
0.19,
<abbrev id="ABBRID0EHKBG" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0ELKBG" xlink_title="anterior lateral eyes">ALE</abbrev>
0.10,
<abbrev id="ABBRID0EPKBG" xlink_title="posterior median eyes">PME</abbrev>
<abbrev id="ABBRID0ETKBG" xlink_title="posterior median eyes">PME</abbrev>
0.36,
<abbrev id="ABBRID0EXKBG" xlink_title="posterior median eyes">PME</abbrev>
<abbrev id="ABBRID0E2KBG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.21,
<abbrev id="ABBRID0E6KBG" xlink_title="MOQ length">
<collectionCode id="42D3446991BC3F0094DB9047211B2CB8">MOQL</collectionCode>
</abbrev>
0.31,
<abbrev id="ABBRID0EDLBG" xlink_title="MOQ anterior width">
<collectionCode id="8C28AC98D1721CD89D4ED29FE0B14EE8">MOQA</collectionCode>
</abbrev>
0.32,
<abbrev id="ABBRID0EHLBG" xlink_title="MOQ posterior width">
<collectionCode id="935FE129D0B125496B6D7FEFF7D01B19">MOQP</collectionCode>
</abbrev>
0.48,
<abbrev id="ABBRID0ELLBG" xlink_title="clypeal height">
<collectingCountry id="3F85F4F758751797573516F7A43F638F" name="Switzerland">CH</collectingCountry>
</abbrev>
0.22. Sternum 0.89 long, 0.80 wide.
<emphasis id="73C8AD0D83B7D4B62F639BFA290EE250" bold="true" italics="true">Measurements of legs</emphasis>
: I 6.50 (1.85, 2.38, 1.40, 0.87), II 7.88 (2.24, 2.92, 1.71, 1.01), III 5.42 (1.72, 1.88, 1.20, 0.62), IV 5.57 (1.77, 1.91, 1.25, 0.64). Leg formula: II-I-IV-III. Cheliceral furrow with one promarginal tooth.
</paragraph>
<paragraph id="00766E5A546239D912F4246E1F30D657">
<emphasis id="CD736FB2493893A0FD8705272B768D8E" bold="true" italics="true">Colouration in ethanol</emphasis>
(Figs
<figureCitation id="83D81A6F84B3289CB9E624FC814B2509" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Philodromus guiyang, female (A C) and male (D F), habitus A, D dorsal view B, E ventral view C, F lateral view. Scale bars: 1 mm (equal for A C, equal for D F)." figureDoi="10.3897/zookeys.1240.149456.figure7" httpUri="https://binary.pensoft.net/fig/1348765">7 A C</figureCitation>
,
<figureCitation id="BD80D598AFF8BE4B9C947D37C717B7D8" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Philodromus guiyang, female (A, C G) and male (B), frontal view of carapace (A, B) and epigyne (C G) A female B male C intact, ventral view D macerated, ventral view E macerated, dorsal view F macerated and embedded in arabic gum, ventral view G macerated and embedded in Arabic gum, dorsal view. Abbreviations: A = atrium; CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; GM = glandular mound; MS = median septum; R = receptaculum; rMS = rim of median septum (teal green line); rSEF = rim of sclerotised epigynal fold (blue line); SEF = sclerotised epigynal fold. Scale bars: 0.5 mm (A, B); 0.2 mm (equal for C G)." figureDoi="10.3897/zookeys.1240.149456.figure8" httpUri="https://binary.pensoft.net/fig/1348766">8 A</figureCitation>
). Carapace nearly pear-shaped, ocular region distinctly narrowed, tegument relatively smooth, with numerous hair bases (all hairs detached); lateral bands dark brown, ~ 1 / 6 of carapace width, respectively; median band wide, ~ 2 / 3 of carapace width, bright yellowish-brown and distinctly delimited to lateral bands, centrally with V-shaped white stripe starting from behind
<abbrev id="ABBRID0EBMBG" xlink_title="posterior lateral eyes">PLE</abbrev>
, almost reaching indistinct cervical groove; radial furrows and fovea indistinctly marked. Cheliceral base coloured slightly darker than median band, with pale brown fangs. Sternum uniformly yellowish-white. Endites and labium coloured as cheliceral base, both with dense scopulae on anterior margins. Legs yellowish-brown, without distinct markings, and covered by short spines. Abdomen elongate-oval, dorsum with a narrow, purplish median band starting from behind pedicel, reaching 1 / 2 of abdomen length; posteriorly with numerous purplish-black streaks interspersed with many pale brown spots, forming a reticulated pattern; ventral abdomen uniformly pale brownish.
</paragraph>
<caption id="92903EE06D6FDCF0E7897BC956878CD7" ID-DOI="10.3897/zookeys.1240.149456.figure7" ID-arpha="6F8045C4-CE1A-5AE3-8E97-4A1AF3A3B7A4" httpUri="https://binary.pensoft.net/fig/1348765" startId="F7">
<paragraph id="E6D3499C3B9B697543562A0E3DFE1AA9">
<label id="DF03D527102D79A3E44C1344DC8E8C87">Figure 7.</label>
</paragraph>
<paragraph id="3574A655A99200AC7F00F58772106AB0">
<taxonomicName id="C1D986A10CCEDBF988651D74AB1BA36D" authorityName="Long &amp; Yu" authorityYear="2022" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="guiyang">
<emphasis id="9EC8B8AFE5D6B55DACBFAD82D2ACA2E5" italics="true">Philodromus guiyang</emphasis>
</taxonomicName>
, female (
<emphasis id="D50D60FB472FD5AE25965AE7B5964D0E" bold="true">A C</emphasis>
) and male (
<emphasis id="8A73CD209193E02CD56482193ABA1893" bold="true">D F</emphasis>
), habitus
<emphasis id="1AD83A465E1FACD6EAE09BE825498889" bold="true">A, D</emphasis>
dorsal view
<emphasis id="C330FABE13E2CB7D5B705557255272AC" bold="true">B, E</emphasis>
ventral view
<emphasis id="23B05409639769C2097D274C60399622" bold="true">C, F</emphasis>
lateral view. Scale bars: 1 mm (equal for
<emphasis id="63645FF8511B3DEF39971B5126DB40BC" bold="true">A C</emphasis>
, equal for
<emphasis id="8270AA1E1B2872477D8A9BE03B6B473B" bold="true">D F</emphasis>
).
</paragraph>
</caption>
<caption id="E0F7B03B2239DE1E438D487633F73F91" ID-DOI="10.3897/zookeys.1240.149456.figure8" ID-arpha="3426E89D-5524-50A3-979F-D824AB58DD6B" httpUri="https://binary.pensoft.net/fig/1348766" startId="F8">
<paragraph id="33218808D883842C602CBA5E7B75E83B">
<label id="AA334184594BD02B9957848E95D25C65">Figure 8.</label>
</paragraph>
<paragraph id="3FEB209176C4485FBD7667E7C3C64977">
<taxonomicName id="C1FB0E456FA40BD87DEE2D26963F128A" authorityName="Long &amp; Yu" authorityYear="2022" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="guiyang">
<emphasis id="FB4DD613C2C74D320F923A829AACA690" italics="true">Philodromus guiyang</emphasis>
</taxonomicName>
, female (
<emphasis id="EF05100BC1A04057A44D132A6E04C103" bold="true">A, C G</emphasis>
) and male (
<emphasis id="97419ED3E8629B6FA8F60867F10E9DB6" bold="true">B</emphasis>
), frontal view of carapace (
<emphasis id="CB73BC006FE7CD5749709061FFDC0173" bold="true">A, B</emphasis>
) and epigyne (
<emphasis id="7B76EC99735B26C3520BA2D0EB7D0087" bold="true">C G</emphasis>
)
<emphasis id="CFB3C261064D6A8FED3467D828E25AFC" bold="true">A</emphasis>
female
<emphasis id="DE42E7E5E91A3E8E0B56ADCDA601C4F4" bold="true">B</emphasis>
male
<emphasis id="E6ADDD4DE075C008A71431FA3ACE79B8" bold="true">C</emphasis>
intact, ventral view
<emphasis id="89F194EDA0F3548F8D7BAC2E576FCCAF" bold="true">D</emphasis>
macerated, ventral view
<emphasis id="76AED2B143D20EF820872FA60169BE59" bold="true">E</emphasis>
macerated, dorsal view
<emphasis id="3929ED9BA909737D90359212C6ECDEE4" bold="true">F</emphasis>
macerated and embedded in arabic gum, ventral view
<emphasis id="F2D836F672AF31B89BB93E459E8A5023" bold="true">G</emphasis>
macerated and embedded in Arabic gum, dorsal view. Abbreviations: A = atrium;
<abbrev id="ABBRID0EUOBG" xlink_title="copulatory duct">CD</abbrev>
= copulatory duct;
<abbrev id="ABBRID0EYOBG" xlink_title="copulatory opening">CO</abbrev>
= copulatory opening;
<abbrev id="ABBRID0E3OBG" xlink_title="fertilisation duct">FD</abbrev>
= fertilisation duct;
<abbrev id="ABBRID0EAPBG" xlink_title="glandular mound">GM</abbrev>
= glandular mound;
<abbrev id="ABBRID0EEPBG" xlink_title="median septum">MS</abbrev>
= median septum; R = receptaculum;
<abbrev id="ABBRID0EIPBG" xlink_title="rim of median septum">rMS</abbrev>
= rim of median septum (teal green line);
<abbrev id="ABBRID0EMPBG" xlink_title="rim of sclerotised epigynal fold">rSEF</abbrev>
= rim of sclerotised epigynal fold (blue line);
<abbrev id="ABBRID0EQPBG" xlink_title="sclerotised epigynal fold">SEF</abbrev>
= sclerotised epigynal fold. Scale bars: 0.5 mm (
<emphasis id="E97BD2A48AAE1C06454F891B29DBD7F9" bold="true">A, B</emphasis>
); 0.2 mm (equal for
<emphasis id="B36469314A592715608631B145950488" bold="true">C G</emphasis>
).
</paragraph>
</caption>
<caption id="A3DEC855A9D331F8BE3C19F251D8F0A1" ID-DOI="10.3897/zookeys.1240.149456.figure9" ID-arpha="F7CF1281-F509-52C7-B990-41D594C7792D" httpUri="https://binary.pensoft.net/fig/1348767" startId="F9">
<paragraph id="148B5825EB160F586B0F3ADC00BBE6F6">
<label id="86C9BAB6678426F91BB1411E1B38E2D3">Figure 9.</label>
</paragraph>
<paragraph id="A77C158A3B35AAA44768E70F60C9A614">
<taxonomicName id="BD122E33A3397108FA6E98956F4A48AF" authorityName="Long &amp; Yu" authorityYear="2022" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="guiyang">
<emphasis id="4D4660878A012F4B8BA7B14891E92A80" italics="true">Philodromus guiyang</emphasis>
</taxonomicName>
, male, left palp
<emphasis id="5ECB92083ED728A26C25ABCAAFE3A3AE" bold="true">A</emphasis>
ventral view
<emphasis id="1081BA532E58E08DE8E6CA3126E84F40" bold="true">B</emphasis>
dorsal view
<emphasis id="70EDE638B7D11678929FBB4EB575ECEF" bold="true">C</emphasis>
prolateral view
<emphasis id="71FDB5FFDC2AA9DE5E64D0F287091653" bold="true">D</emphasis>
retrolateral view. Abbreviations:
<abbrev id="ABBRID0EYQBG" xlink_title="ascending part of sperm duct loop">aSDL</abbrev>
= ascending part of sperm duct loop;
<abbrev id="ABBRID0E3QBG" xlink_title="conductor">Con</abbrev>
= conductor;
<abbrev id="ABBRID0EARBG" xlink_title="cymbial process">CP</abbrev>
= cymbial process;
<abbrev id="ABBRID0EERBG" xlink_title="cymbium">Cy</abbrev>
= cymbium;
<abbrev id="ABBRID0EIRBG" xlink_title="dorsal branch of RTA">dRTA</abbrev>
= dorsal branch of
<abbrev id="ABBRID0EMRBG" xlink_title="retrolateral tibial apophysis">RTA</abbrev>
;
<abbrev id="ABBRID0EQRBG" xlink_title="descending part of sperm duct loop">dSDL</abbrev>
= descending part of sperm duct loop;
<abbrev id="ABBRID0EURBG" xlink_title="embolus">Em</abbrev>
= embolus;
<abbrev id="ABBRID0EYRBG" xlink_title="embolic base">EmB</abbrev>
= embolic base;
<abbrev id="ABBRID0E3RBG" xlink_title="embolic tip">EmT</abbrev>
= embolic tip;
<abbrev id="ABBRID0EASBG" xlink_title="intertegular retinaculum">IR</abbrev>
= intertegular retinaculum;
<abbrev id="ABBRID0EESBG" xlink_title="prolateral part of conductor">pCon</abbrev>
= prolateral part of conductor;
<abbrev id="ABBRID0EISBG" xlink_title="retrolateral tibial apophysis">RTA</abbrev>
= retrolateral tibial apophysis;
<abbrev id="ABBRID0EMSBG" xlink_title="sperm duct">SD</abbrev>
= sperm duct;
<abbrev id="ABBRID0EQSBG" xlink_title="tip of conductor">tCon</abbrev>
= tip of conductor;
<abbrev id="ABBRID0EUSBG" xlink_title="tegulum">Te</abbrev>
= tegulum;
<abbrev id="ABBRID0EYSBG" xlink_title="ventral branch of RTA">vRTA</abbrev>
= ventral branch of
<abbrev id="ABBRID0E3SBG" xlink_title="retrolateral tibial apophysis">RTA</abbrev>
;
<abbrev id="ABBRID0EATBG" xlink_title="ventral tibial apophysis">VTA</abbrev>
= ventral tibial apophysis. Scale bar: 0.2 mm (equal for
<emphasis id="6F1A8FD9DDA28E7EF279787C84E1E506" bold="true">A D</emphasis>
).
</paragraph>
</caption>
<caption id="C91212CA7F600D0D4A68EEA83EC38577" ID-DOI="10.3897/zookeys.1240.149456.figure10" ID-arpha="A1E48F13-0387-5F26-B889-12A8FDF3D5F5" httpUri="https://binary.pensoft.net/fig/1348768" startId="F10">
<paragraph id="A91DDE908A9A2EDD1325D5C4049CC52C">
<label id="C424494D5E9B7F98E637F1D9F36D3929">Figure 10.</label>
</paragraph>
<paragraph id="8B1FC0B3B0E99ED5EEE608382E1C192F">
<taxonomicName id="26193A285A5F27B72876025A911557A0" authorityName="Bosenberg &amp; Strand" authorityYear="1906" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="subaureolus">
<emphasis id="51E886E4921B61B5B9EBB793B5E69D2F" italics="true">Philodromus subaureolus</emphasis>
</taxonomicName>
, male, habitus (
<emphasis id="9799F8CB76C55F1D404807079FED0771" bold="true">A C</emphasis>
) and left palp (
<emphasis id="C6C4FED4EBA7F55C24C745FA71720164" bold="true">D F</emphasis>
)
<emphasis id="8A10A1C3F887CA2EDD2B0714FF308351" bold="true">A</emphasis>
dorsal view
<emphasis id="DBF550EB272475EECDB5AB9DD51683E6" bold="true">B</emphasis>
ventral view
<emphasis id="4A7ECB949D2FC65F8C63F0DD81899531" bold="true">C</emphasis>
lateral view
<emphasis id="067A6256B14F60F7FEB5A48E1B447D38" bold="true">D</emphasis>
prolateral view
<emphasis id="FF8DC4BABDB2D73908673BCBAAA32718" bold="true">E</emphasis>
ventral view
<emphasis id="564AE4C94841731E97E4B75AFCC4C8E2" bold="true">F</emphasis>
retrolateral view. Abbreviations:
<abbrev id="ABBRID0EOUBG" xlink_title="ascending part of sperm duct loop">aSDL</abbrev>
= ascending part of sperm duct loop;
<abbrev id="ABBRID0ESUBG" xlink_title="conductor">Con</abbrev>
= conductor;
<abbrev id="ABBRID0EWUBG" xlink_title="cymbium">Cy</abbrev>
= cymbium;
<abbrev id="ABBRID0E1UBG" xlink_title="dorsal branch of RTA">dRTA</abbrev>
= dorsal branch of
<abbrev id="ABBRID0E5UBG" xlink_title="retrolateral tibial apophysis">RTA</abbrev>
;
<abbrev id="ABBRID0ECVBG" xlink_title="descending part of sperm duct loop">dSDL</abbrev>
= descending part of sperm duct loop;
<abbrev id="ABBRID0EGVBG" xlink_title="embolus">Em</abbrev>
= embolus;
<abbrev id="ABBRID0EKVBG" xlink_title="embolic base">EmB</abbrev>
= embolic base;
<abbrev id="ABBRID0EOVBG" xlink_title="embolic tip">EmT</abbrev>
= embolic tip;
<abbrev id="ABBRID0ESVBG" xlink_title="intertegular retinaculum">IR</abbrev>
= intertegular retinaculum;
<abbrev id="ABBRID0EWVBG" xlink_title="prolateral part of conductor">pCon</abbrev>
= prolateral part of conductor;
<abbrev id="ABBRID0E1VBG" xlink_title="retrolateral tibial apophysis">RTA</abbrev>
= retrolateral tibial apophysis;
<abbrev id="ABBRID0E5VBG" xlink_title="sperm duct">SD</abbrev>
= sperm duct;
<abbrev id="ABBRID0ECWBG" xlink_title="tip of conductor">tCon</abbrev>
= tip of conductor;
<abbrev id="ABBRID0EGWBG" xlink_title="tegulum">Te</abbrev>
= tegulum;
<abbrev id="ABBRID0EKWBG" xlink_title="ventral branch of RTA">vRTA</abbrev>
= ventral branch of
<abbrev id="ABBRID0EOWBG" xlink_title="retrolateral tibial apophysis">RTA</abbrev>
;
<abbrev id="ABBRID0ESWBG" xlink_title="ventral tibial apophysis">VTA</abbrev>
= ventral tibial apophysis. Scale bars: 1 mm (equal for
<emphasis id="88ABEEFA22C751BE4A37018E13E3C6AA" bold="true">A C</emphasis>
); 0.2 mm (equal for
<emphasis id="95DAC5F51B2F562942492A3345C231EA" bold="true">D F</emphasis>
).
</paragraph>
</caption>
<caption id="92E059497A791636F5E3F6F2ADA32AD2" ID-DOI="10.3897/zookeys.1240.149456.figure11" ID-arpha="0CEC2C9B-63ED-5753-8068-6332E80FA49F" httpUri="https://binary.pensoft.net/fig/1348769" startId="F11">
<paragraph id="077D80F32D0002442BED680F44B3C667">
<label id="EDE57BC00D9B3FA4B7B191AE3B9EB2CA">Figure 11.</label>
</paragraph>
<paragraph id="8BBC0209F553F03644B31717E7226578">
<taxonomicName id="4A112D27DE6218EFE4A3CB16B2B11F3F" authorityName="Bosenberg &amp; Strand" authorityYear="1906" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="subaureolus">
<emphasis id="FFDA08938E55B1525D3B45C3FE0A2E82" italics="true">Philodromus subaureolus</emphasis>
</taxonomicName>
, female, habitus (
<emphasis id="95425D49B89D2DB9A1563C41143BFAD0" bold="true">A C</emphasis>
) and epigyne (
<emphasis id="788A429CF75187AC4A188FE44C18D017" bold="true">D, E</emphasis>
)
<emphasis id="834292CB8330D11B9426771449C350ED" bold="true">A</emphasis>
dorsal view
<emphasis id="C130C44C398C6CF78EF68E48FE74CAFF" bold="true">B</emphasis>
ventral view
<emphasis id="14D936331FDD7487B233A21687D0F61A" bold="true">C</emphasis>
lateral view
<emphasis id="255C4BD97943147BB03B5EAD85305D08" bold="true">D</emphasis>
ventral view
<emphasis id="B760F9A345D044480E7F9746975DBB0B" bold="true">E</emphasis>
dorsal view. Abbreviations: A = atrium;
<abbrev id="ABBRID0EAYBG" xlink_title="course of copulatory duct">cCD</abbrev>
= course of copulatory duct (red line);
<abbrev id="ABBRID0EEYBG" xlink_title="copulatory duct">CD</abbrev>
= copulatory duct;
<abbrev id="ABBRID0EIYBG" xlink_title="copulatory opening">CO</abbrev>
= copulatory opening;
<abbrev id="ABBRID0EMYBG" xlink_title="fertilisation duct">FD</abbrev>
= fertilisation duct;
<abbrev id="ABBRID0EQYBG" xlink_title="glandular mound">GM</abbrev>
= glandular mound;
<abbrev id="ABBRID0EUYBG" xlink_title="median septum">MS</abbrev>
= median septum; R = receptaculum;
<abbrev id="ABBRID0EYYBG" xlink_title="rim of median septum">rMS</abbrev>
= rim of median septum (teal green line);
<abbrev id="ABBRID0E3YBG" xlink_title="rim of sclerotised epigynal fold">rSEF</abbrev>
= rim of sclerotised epigynal fold (blue line);
<abbrev id="ABBRID0EAZBG" xlink_title="sclerotised epigynal fold">SEF</abbrev>
= sclerotised epigynal fold. Scale bars: 1 mm (equal for
<emphasis id="65B713922049FB8A4A8063204FB676BA" bold="true">A C</emphasis>
); 0.2 mm (equal for
<emphasis id="DCF9745948CF872B8E98F507A7E6F4C5" bold="true">D, E</emphasis>
).
</paragraph>
</caption>
<paragraph id="F02A8D096E2514CEEF19F2F9F6374FB6">
<emphasis id="2DDD171354E17D565E70A15A358FAC56" bold="true" italics="true">Epigyne</emphasis>
(Fig.
<figureCitation id="8479777D1984B54F9A71837D694BCD25" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Philodromus guiyang, female (A, C G) and male (B), frontal view of carapace (A, B) and epigyne (C G) A female B male C intact, ventral view D macerated, ventral view E macerated, dorsal view F macerated and embedded in arabic gum, ventral view G macerated and embedded in Arabic gum, dorsal view. Abbreviations: A = atrium; CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; GM = glandular mound; MS = median septum; R = receptaculum; rMS = rim of median septum (teal green line); rSEF = rim of sclerotised epigynal fold (blue line); SEF = sclerotised epigynal fold. Scale bars: 0.5 mm (A, B); 0.2 mm (equal for C G)." figureDoi="10.3897/zookeys.1240.149456.figure8" httpUri="https://binary.pensoft.net/fig/1348766">8 C G</figureCitation>
). Epigynal field slightly longer than wide; anterior and lateral margins not delimited, posterior margin rebordered; the arrangement of the various parts of the vulva (
<abbrev id="ABBRID0EYZBG" xlink_title="copulatory duct">
<collectionCode id="DA294C9C42D54B1AA32BF3C16330F87E">CD</collectionCode>
</abbrev>
and
<collectionCode id="5050AF858528BE64D92671971FD6C1C8" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
) are distinctly visible through integument. A small, located at anterior part of epigynal plate, divided by anterior keel of
<abbrev id="ABBRID0E3ZBG" xlink_title="median septum">MS</abbrev>
, represented by two comma-shaped cavities; the two cavities separated by ~ 1 diameter.
<abbrev id="ABBRID0EA1BG" xlink_title="median septum">MS</abbrev>
bell-shaped, broad; anteriorly narrowed abruptly, ~ 1 / 5 epigyne width, with distinct edges and distinctly delimited to A; posteriorly widen gradually, ~ 1 / 2 epigyne width, with indistinct lateral
<abbrev id="ABBRID0EE1BG" xlink_title="rim of median septum">rMS</abbrev>
alongside with
<abbrev id="ABBRID0EI1BG" xlink_title="rim of sclerotised epigynal fold">rSEF</abbrev>
.
<abbrev id="ABBRID0EM1BG" xlink_title="sclerotised epigynal fold">SEF</abbrev>
anteriorly and posteriorly narrowed, medially widened, ~ 1 / 3 epigyne width, not delimited to
<abbrev id="ABBRID0EQ1BG" xlink_title="median septum">MS</abbrev>
.
<abbrev id="ABBRID0EU1BG" xlink_title="copulatory opening">CO</abbrev>
indistinct, located at basolateral atrial borders, leading to
<abbrev id="ABBRID0EY1BG" xlink_title="copulatory duct">
<collectionCode id="ECE8E8CF802ED86E9DED945B6C74FE05">CD</collectionCode>
</abbrev>
which extend to connect with
<collectionCode id="EB63221D6CE701F1DBA1451B5031161B" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
.
<abbrev id="ABBRID0E31BG" xlink_title="copulatory duct">
<collectionCode id="E8C1C4386077D1C8DC02CE377A952B64">CD</collectionCode>
</abbrev>
distinctly thick and heavily sclerotised, almost as thick as
<collectionCode id="C3BFC24EA71157FB6DDBCD7DC8D625DF" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
, ~ 1 / 4 epigyne width; anteriorly convergent on the central axis, posteriorly descend obliquely, separated by ~ 1 diameter, finally connected to
<collectionCode id="925D2A114B12DF675AF02869803F4514" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
at midlength of epigyne.
<collectionCode id="B0FD8A028B582DAF8C8AC5FEF51B53F9" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
elongate-oval, ~ 1 / 2 epigyne length and 1 / 4 epigyne width, arranged obliquely; anteriorly separated by ~ 0.5 × diameter, posteriorly separated by ~ 1.3 × diameter.
<abbrev id="ABBRID0EA2BG" xlink_title="glandular mound">
<collectingRegion id="4670620A1BF702F5237534C33390C89B" country="Iran" name="Qom">GM</collectingRegion>
</abbrev>
distinctly small, represented by small humps that locate at the antero-lateral surfaces of
<collectionCode id="CE6F36EC60C2C456CD2641C39040AF91" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
.
<abbrev id="ABBRID0EE2BG" xlink_title="fertilisation duct">FD</abbrev>
membranous and acicular, large, ~ 2 / 3 of
<collectionCode id="670F96283EF4A8BF4F7606C32F678244" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
length, originating from the posterior surface of
<collectionCode id="7F2C36CB9280A0C5DF03080F75F7B705" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
, anterolaterally extending.
</paragraph>
<paragraph id="F43B73C4B806228B753F7E99868EF7C2">
<emphasis id="3380D6E12F7F8A44565A9B345CF042D1" bold="true">Male.</emphasis>
Total length 3.25. Carapace 1.49 long, 1.42 wide. Abdomen 1.76 long, 1.15 wide.
<emphasis id="E18F9EEAF3A10BEB03A63B6AC8B44BED" bold="true" italics="true">Eye sizes and interdistances</emphasis>
:
<abbrev id="ABBRID0EP2BG" xlink_title="anterior median eyes">AME</abbrev>
0.07,
<abbrev id="ABBRID0ET2BG" xlink_title="anterior lateral eyes">ALE</abbrev>
0.06,
<abbrev id="ABBRID0EX2BG" xlink_title="posterior median eyes">PME</abbrev>
0.06,
<abbrev id="ABBRID0E22BG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.08,
<abbrev id="ABBRID0E62BG" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0ED3BG" xlink_title="anterior median eyes">AME</abbrev>
0.16,
<abbrev id="ABBRID0EH3BG" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0EL3BG" xlink_title="anterior lateral eyes">ALE</abbrev>
0.08,
<abbrev id="ABBRID0EP3BG" xlink_title="posterior median eyes">PME</abbrev>
<abbrev id="ABBRID0ET3BG" xlink_title="posterior median eyes">PME</abbrev>
0.30,
<abbrev id="ABBRID0EX3BG" xlink_title="posterior median eyes">PME</abbrev>
<abbrev id="ABBRID0E23BG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.19,
<abbrev id="ABBRID0E63BG" xlink_title="MOQ length">
<collectionCode id="C31EA90EE9F7A203EE327401EC43B293">MOQL</collectionCode>
</abbrev>
0.30,
<abbrev id="ABBRID0ED4BG" xlink_title="MOQ anterior width">
<collectionCode id="FE7B6F64B447AF02285D4821D2816C48">MOQA</collectionCode>
</abbrev>
0.30,
<abbrev id="ABBRID0EH4BG" xlink_title="MOQ posterior width">
<collectionCode id="892B4CEF91663DD82844C841A4DDFCB2">MOQP</collectionCode>
</abbrev>
0.41,
<abbrev id="ABBRID0EL4BG" xlink_title="clypeal height">
<collectingCountry id="B11101E5FE1411995FCEE3F54F2C1FAE" name="Switzerland">CH</collectingCountry>
</abbrev>
0.19. Sternum 0.89 long, 0.81 wide.
<emphasis id="176A6EC3A4F3E55213D773C7349D05CE" bold="true" italics="true">Measurements of legs</emphasis>
: I 7.78 (1.99, 2.66, 1.92, 1.21), II 9.70 (2.59, 3.27, 2.37, 1.47), III 6.05 (1.87, 2.01, 1.41, 0.76), IV 5.23 (1.79, 1.88, 1.03, 0.53). Leg formula: II-I-III-IV. Cheliceral furrow with one promarginal teeth. Colouration in ethanol as in females, but body slightly paler (Figs
<figureCitation id="AEBB8D70BD38C36DA037786E71B32767" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Philodromus guiyang, female (A C) and male (D F), habitus A, D dorsal view B, E ventral view C, F lateral view. Scale bars: 1 mm (equal for A C, equal for D F)." figureDoi="10.3897/zookeys.1240.149456.figure7" httpUri="https://binary.pensoft.net/fig/1348765">7 D F</figureCitation>
,
<figureCitation id="D0574C9A5A4F10748E7CDBCCF1DD0480" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Philodromus guiyang, female (A, C G) and male (B), frontal view of carapace (A, B) and epigyne (C G) A female B male C intact, ventral view D macerated, ventral view E macerated, dorsal view F macerated and embedded in arabic gum, ventral view G macerated and embedded in Arabic gum, dorsal view. Abbreviations: A = atrium; CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; GM = glandular mound; MS = median septum; R = receptaculum; rMS = rim of median septum (teal green line); rSEF = rim of sclerotised epigynal fold (blue line); SEF = sclerotised epigynal fold. Scale bars: 0.5 mm (A, B); 0.2 mm (equal for C G)." figureDoi="10.3897/zookeys.1240.149456.figure8" httpUri="https://binary.pensoft.net/fig/1348766">8 B</figureCitation>
; see
<bibRefCitation id="67D604747B612CCF00806385EC21D90A" DOI="10.3969/j.issn.1005-9628.2022.02.009" author="Long FY &amp; Deng LJ &amp; Mao JL &amp; Yu H" etAl="et al." firstAuthor="Long" issue="2" journalOrPublisher="Acta Arachnologica Sinica" pagination="113-120" refId="B33" refString="Long FY, Deng LJ, Mao JL, Yu H (2022) A survey of genus Philodromus (Araneae: Philodromidae) from Guiyang, Guizhou, China, with description of a new species. Acta Arachnologica Sinica 31 (2): 113120. https://doi.org/10.3969/j.issn.1005-9628.2022.02.009" title="A survey of genus Philodromus (Araneae: Philodromidae) from Guiyang, Guizhou, China, with description of a new species." volume="31" year="2022">Long et al. (2022)</bibRefCitation>
for others described).
</paragraph>
<paragraph id="DCC2F3C8E4ECBDA6671A065B6CA1A770">
<emphasis id="9AC925D703EBC89655B725E82FECED7B" bold="true" italics="true">Palp</emphasis>
(Fig.
<figureCitation id="932A6AF47E71E81671CAB8CCF1190036" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Philodromus guiyang, male, left palp A ventral view B dorsal view C prolateral view D retrolateral view. Abbreviations: aSDL = ascending part of sperm duct loop; Con = conductor; CP = cymbial process; Cy = cymbium; dRTA = dorsal branch of RTA; dSDL = descending part of sperm duct loop; Em = embolus; EmB = embolic base; EmT = embolic tip; IR = intertegular retinaculum; pCon = prolateral part of conductor; RTA = retrolateral tibial apophysis; SD = sperm duct; tCon = tip of conductor; Te = tegulum; vRTA = ventral branch of RTA; VTA = ventral tibial apophysis. Scale bar: 0.2 mm (equal for A D)." figureDoi="10.3897/zookeys.1240.149456.figure9" httpUri="https://binary.pensoft.net/fig/1348767">9 A D</figureCitation>
). Tibia relatively long, ~ 2 / 3 of
<abbrev id="ABBRID0EJ5BG" xlink_title="cymbium">Cy</abbrev>
length, with two apophyses arising distally from tibia. Both tibial apophyses are lamellar and almost equal in length, nearly as long as palpal tibia length, including: a weakly sclerotised
<abbrev id="ABBRID0EN5BG" xlink_title="ventral tibial apophysis">VTA</abbrev>
, blade-shaped in ventral view and finger-shaped in lateral views; and a more sclerotised
<abbrev id="ABBRID0ER5BG" xlink_title="retrolateral tibial apophysis">
<collectionCode id="91448A9C6614F0F99BD2D398A68E1AE4">RTA</collectionCode>
</abbrev>
which with wide base, narrowed middle section, and more or less biforked tip; both branches of
<abbrev id="ABBRID0EV5BG" xlink_title="retrolateral tibial apophysis">
<collectionCode id="531B7C08061B4F4ACCD321E313FC416D">RTA</collectionCode>
</abbrev>
nearly triangular,
<abbrev id="ABBRID0EZ5BG" xlink_title="ventral branch of RTA">vRTA</abbrev>
heavily sclerotised, surface and edges smooth, apex angle is approximately 30 °,
<abbrev id="ABBRID0E45BG" xlink_title="dorsal branch of RTA">dRTA</abbrev>
hyaline, surface rough and with several scratch-like textures, distal edge jagged, apex angle is approximately 90 °.
<abbrev id="ABBRID0EB6BG" xlink_title="cymbium">Cy</abbrev>
distinctly longer than tibia, basoretrolaterally with an indistinct
<abbrev id="ABBRID0EF6BG" xlink_title="cymbial process">CP</abbrev>
.
<abbrev id="ABBRID0EJ6BG" xlink_title="tegulum">Te</abbrev>
oval, ~ 1.37 longer than wide, proximally slightly swollen, prolatero-apically slightly excavated to accommodate
<abbrev id="ABBRID0EN6BG" xlink_title="embolus">Em</abbrev>
and
<abbrev id="ABBRID0ER6BG" xlink_title="conductor">Con</abbrev>
.
<abbrev id="ABBRID0EV6BG" xlink_title="sperm duct">SD</abbrev>
sinuate, originating at distal portion of
<abbrev id="ABBRID0EZ6BG" xlink_title="tegulum">Te</abbrev>
, aligning clockwise along the tegular retrolateral margin, forming a S-shaped
<abbrev id="ABBRID0E46BG" xlink_title="sperm duct loop">SDL</abbrev>
in ventral view, finally terminating at the ~ 10 oclock position, and entering
<abbrev id="ABBRID0ECAAI" xlink_title="embolic base">EmB</abbrev>
.
<abbrev id="ABBRID0EGAAI" xlink_title="embolus">Em</abbrev>
claw-shaped,
<abbrev id="ABBRID0EKAAI" xlink_title="embolic base">EmB</abbrev>
thick, inserted prolatero-apically (approximately 10 oclock relative to
<abbrev id="ABBRID0EOAAI" xlink_title="tegulum">Te</abbrev>
), gradually tapering toward apex;
<abbrev id="ABBRID0ESAAI" xlink_title="embolic tip">EmT</abbrev>
sharp and retrolaterally pointed, terminating at ~ 1 oclock position.
<abbrev id="ABBRID0EWAAI" xlink_title="conductor">Con</abbrev>
membranous, axe-shaped, aligning transversely on apical part of the
<abbrev id="ABBRID0E1AAI" xlink_title="tegulum">Te</abbrev>
, enveloping the second half of
<abbrev id="ABBRID0E5AAI" xlink_title="embolus">Em</abbrev>
.
<abbrev id="ABBRID0ECBAI" xlink_title="intertegular retinaculum">IR</abbrev>
distinctly small, spine-shaped, located at the ~ 2 oclock position.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGBAI" type="distribution">
<paragraph id="885B23ECD28152AD64598B19CC2346E3">
<heading id="97720063A66C92D97F7DFDD11B5BFC3C" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="2A3F7658B3AC3C533B3DD7C290DD7B30">
<collectingCountry id="9F6E52E3A4D175D533CF23BE05FEB7C4" name="China">China</collectingCountry>
(
<collectingRegion id="D3B1836723ADFE675EDA739AE4ED0375" country="China" name="Fujian">Fujian</collectingRegion>
,
<collectingRegion id="D3CEFABE7CD3FC6CA432A0DC0E527D60" country="China" name="Guizhou">Guizhou</collectingRegion>
,
<collectingRegion id="32EFF8475E2FF9475033F1059EE5DFC3" country="China" name="Hunan">Hunan</collectingRegion>
,
<collectingRegion id="AE5FC7B2E58E7BCB1A755458D986CB81" country="China" name="Jiangxi">Jiangxi</collectingRegion>
; distribution records in
<collectingRegion id="7D1CC4C3930346659F5C389297D12A39" country="China" name="Guizhou">Guizhou</collectingRegion>
as in Fig.
<figureCitation id="07D11F990BF430570B51C23AC000E01B" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Distribution records of the philodromid species in Guizhou Province (A) and Hubei Province (B)." figureDoi="10.3897/zookeys.1240.149456.figure23" httpUri="https://binary.pensoft.net/fig/1348781">23 A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQBAI" type="comments">
<paragraph id="78F21A7B835438D75DCBEEBE84E2A9EE">
<heading id="B40EF42754005BB8E355FC825212DB58" reason="title">Comments.</heading>
</paragraph>
<paragraph id="15D78F18C7C851E52FF52E1C8B52D356">
<bibRefCitation id="A331474068255241EBEFCC6F6CE9F6CB" DOI="10.3969/j.issn.1005-9628.2022.02.009" author="Long FY &amp; Deng LJ &amp; Mao JL &amp; Yu H" etAl="et al." firstAuthor="Long" issue="2" journalOrPublisher="Acta Arachnologica Sinica" pagination="113-120" refId="B33" refString="Long FY, Deng LJ, Mao JL, Yu H (2022) A survey of genus Philodromus (Araneae: Philodromidae) from Guiyang, Guizhou, China, with description of a new species. Acta Arachnologica Sinica 31 (2): 113120. https://doi.org/10.3969/j.issn.1005-9628.2022.02.009" title="A survey of genus Philodromus (Araneae: Philodromidae) from Guiyang, Guizhou, China, with description of a new species." volume="31" year="2022">Long et al. (2022)</bibRefCitation>
described the
<typeStatus id="04B2656636400A5B7C9E27026BD9C122" type="holotype">holotype</typeStatus>
male of
<taxonomicName id="08B3807C7303A3A8D07E02D3E5B2CE2C" authorityName="Long &amp; Yu" authorityYear="2022" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="guiyang">
<emphasis id="5A2A9B5EBA8F3A0834340D1B55BA0C75" italics="true">P. guiyang</emphasis>
</taxonomicName>
in Chinese in the original paper.
<bibRefCitation id="A7B0EF8CD3DAB5B33AD12A5631DDBF2D" DOI="10.3897/zookeys.1221.137930" author="Wang ZJ &amp; Yao YB &amp; Tang ZY &amp; Li WH &amp; Liu KK &amp; Xu X" etAl="et al." firstAuthor="Wang" journalOrPublisher="ZooKeys" pagination="279-296" refId="B55" refString="Wang ZJ, Yao YB, Tang ZY, Li WH, Liu KK, Xu X (2024) A new genus, Sinodromus gen. nov., with two new species and the first description of the female of Philodromus guiyang Long &amp; Yu, 2022 (Arachnida, Araneae, Philodromidae) from China. ZooKeys 296: 279296. https://doi.org/10.3897/zookeys.1221.137930" title="A new genus, Sinodromus gen. nov., with two new species and the first description of the female of Philodromus guiyang Long &amp; Yu, 2022 (Arachnida, Araneae, Philodromidae) from China." volume="296" year="2024">Wang et al. (2024)</bibRefCitation>
later described the female of the species for the first time but did not provide a diagnosis and redescription for the male. Therefore, to date, the male of this species lacks an English description. Here we diagnose and describe the male in English for the first time. Newly available specimens indicate that the species is also distributed in Mayanghe National Nature Reserve in
<collectingRegion id="1CBDFB34D196222BE6DE3030E386B42A" country="China" name="Guizhou">Guizhou Province</collectingRegion>
(Fig.
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).
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<subSubSection id="7AD0C3C89A6B5A61C54143C34A3A9312" type="nomenclature">
<paragraph id="3F6739A038ED3DD1A8626E086EB1A5D7">
<heading id="2B6915695F7E0BEAA78837B18C979AEE" reason="label">Genus</heading>
<taxonomicName id="BA771556FACB50AA923882E564E5B5BA" ID-CoL="7WRZ" authority="Simon, 1875" authorityName="Simon" authorityYear="1875" class="Arachnida" family="Philodromidae" genus="Tibellus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="55170649797A6C40344B227EDBB07C01" italics="true">Tibellus</emphasis>
<bibRefCitation id="80F183FC92E3C04A4A0598E8052DA125" author="Simon E" refId="B45" refString="Simon E (1875) Les Arachnides de France. Tome seconde. Contenant les familles des Urocteidae, Agelenidae, Thomisidae et Sparassidae. Roret, Paris, 360 pp [pls 4 - 8]." year="1875">Simon, 1875</bibRefCitation>
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<subSubSection id="SECID0EDVCI" type="type species">
<paragraph id="FA539231509E51CEA6A07A9CBC8492D5">
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species.
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<paragraph id="F98A50B96D6A4FD3307B1D5A09991024">
<taxonomicName id="56ACB56B425475692D71EF0D8CA1E853" authority="Walckenaer, 1802" authorityName="Walckenaer" authorityYear="1802" class="Gastropoda" family="Muricidae" genus="Aranea" kingdom="Animalia" order="Neogastropoda" phylum="Mollusca" rank="species" species="oblonga">
<emphasis id="6593E8895BEDD7AE7B71703C1DA747F8" italics="true">Aranea oblonga</emphasis>
Walckenaer, 1802
</taxonomicName>
from North America, Europe, North Africa,
<collectingCountry id="F2338403E0CF9F830F402E976088680C" name="Turkey">Turkey</collectingCountry>
,
<collectingCountry id="AC292A4619BD79C4B5A820FB4DA4CF79" name="Israel">Israel</collectingCountry>
, Caucasus,
<collectingCountry id="F9789EBD3F04EC34E88E999DAF47B0DB" name="Russia">Russia</collectingCountry>
(Europe to Far East),
<collectingCountry id="955433FB7D0A9E6FD0296DD95A344842" name="Kazakhstan">Kazakhstan</collectingCountry>
,
<collectingCountry id="863D4C30900D75DE40DF3A0013FA9AC8" name="Iran">Iran</collectingCountry>
, Central Asia,
<collectingCountry id="CB88991C6C14DA3F8B2341C9ED5DD4B7" name="Mongolia">Mongolia</collectingCountry>
,
<collectingCountry id="914989B6166B7EBA3F96793960F69C56" name="China">China</collectingCountry>
,
<collectingCountry id="BBFF7AE087B81DA22ABD9CB9FF210BCD" name="South Korea">Korea</collectingCountry>
,
<collectingCountry id="F74CC2C0D9351F332CF9549FC2D1A906" name="Japan">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUVCI" type="diagnosis">
<paragraph id="F32F99010A9D9A69D41A6DC7D94CD2D0">
<heading id="B7B953A824F2F7EAAE8FE04D3440F996" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="8354649D1F2CEA1D498026FF38A95656">
See
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<collectingRegion id="FCEB3DFF04BCD1361484CA19794AFBE5" country="Turkey" name="Van">Van</collectingRegion>
den Berg and Dippenaar-Schoeman (1994)
</bibRefCitation>
and
<bibRefCitation id="9ECF381789C11AC0FCD63552C6F181BD" author="Efimik VE" firstAuthor="Efimik" journalOrPublisher="Arthropoda Selecta" pagination="103-124" refId="B18" refString="Efimik VE (1999) A review of the spider genus Tibellus Simon, 1875 of the East Palearctic (Aranei: Philodromidae). Arthropoda Selecta 8: 103124." title="A review of the spider genus Tibellus Simon, 1875 of the East Palearctic (Aranei: Philodromidae)." volume="8" year="1999">Efimik (1999)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECWCI" type="comments">
<paragraph id="47227C820C0DDAA528E39B01AD4B795E">
<heading id="CE4640CA30446B847513FDDCEA209266" reason="title">Comments.</heading>
</paragraph>
<paragraph id="3804562D527CD5A87D49E76879E058D9">
The genus has been widely considered as putatively monophyletic; it presents a distinct set of characters (
<bibRefCitation id="6071941F185EAF23E012DCA1FF4DD6DB" DOI="10.4102/koedoe.v37i1.327" author="Van den Berg and Dippenaar-Schoeman" firstAuthor="Van den Berg" issue="1" journalOrPublisher="Koedoe" pagination="67-114" refId="B52" refString="Van den Berg A, Dippenaar-Schoeman AS (1994) A revision of the Afrotropical species of the genus Tibellus Simon (Araneae: Philodromidae). Koedoe 37 (1): 67114. https://doi.org/10.4102/koedoe.v37i1.327" title="A revision of the Afrotropical species of the genus Tibellus Simon (Araneae: Philodromidae)." volume="37" year="1994">
<collectingRegion id="E41F0E0C7BBF2B2775F472079DE2CF0E" country="Turkey" name="Van">Van</collectingRegion>
den Berg and Dippenaar-Schoeman 1994
</bibRefCitation>
;
<bibRefCitation id="B9C52BFD1C0CB343893B68D3D4E44B5B" author="Efimik" firstAuthor="Efimik" journalOrPublisher="Arthropoda Selecta" pagination="103-124" refId="B18" refString="Efimik VE (1999) A review of the spider genus Tibellus Simon, 1875 of the East Palearctic (Aranei: Philodromidae). Arthropoda Selecta 8: 103124." title="A review of the spider genus Tibellus Simon, 1875 of the East Palearctic (Aranei: Philodromidae)." volume="8" year="1999">Efimik 1999</bibRefCitation>
;
<bibRefCitation id="E593AC9AA2286ABEDA564F6305D6BAC6" DOI="10.5635/ASED.2023.39.4.044" author="Jang" etAl="et al." firstAuthor="Jang" issue="4" journalOrPublisher="Animal Systematics, Evolution and Diversity" pagination="272-283" refId="B23" refString="Jang CM, Lee SY, Yoo JS, Bae YS, Kim ST (2023) Six new records of running crab spiders of the genus Tibellus with four new species (Araneae: Philodromidae) from Korea. Animal Systematics, Evolution and Diversity 39 (4): 272283. https://doi.org/10.5635/ASED.2023.39.4.044" title="Six new records of running crab spiders of the genus Tibellus with four new species (Araneae: Philodromidae) from Korea." volume="39" year="2023">Jang et al. 2023</bibRefCitation>
), and its species composition is relatively stable (
<bibRefCitation id="03F63C3C26662F443F622472DF5D39B3" refId="B57" refString="WSC (2025) World Spider Catalog. Version 25.5. Natural History Museum Bern. http://wsc.nmbe.ch [accessed on 12 January 2025]" url="http://wsc.nmbe.ch" year="2025">WSC 2025</bibRefCitation>
). However, some species have extraordinarily intraspecific morphological variation and low levels of interspecific variation, and together with the insufficiency of alpha taxonomic information (lacking high-quality illustrations, detailed descriptions, and molecular data), have hindered species recognition and has resulted in several wrong descriptions and misidentifications (
<bibRefCitation id="B93697F73B275E9975B22120135775B7" author="Efimik" firstAuthor="Efimik" journalOrPublisher="Arthropoda Selecta" pagination="103-124" refId="B18" refString="Efimik VE (1999) A review of the spider genus Tibellus Simon, 1875 of the East Palearctic (Aranei: Philodromidae). Arthropoda Selecta 8: 103124." title="A review of the spider genus Tibellus Simon, 1875 of the East Palearctic (Aranei: Philodromidae)." volume="8" year="1999">Efimik 1999</bibRefCitation>
).
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</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="6A0423D296E4E1B7168751739C2DB741">journal article</mods:classification>
<mods:identifier id="A7A0D5902F394FAABA9FC4DA81165324" type="DOI">10.3897/zookeys.1240.149456</mods:identifier>
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</mods:mods>
<treatment id="B7C1840AFBB0593484F10AE3D37D649E" ID-DOI="http://doi.org/10.5281/zenodo.15625172" ID-Zenodo-Dep="15625172" ID-arpha="B7C1840A-FBB0-5934-84F1-0AE3D37D649E" LSID="urn:lsid:plazi:treatment:B7C1840AFBB0593484F10AE3D37D649E" httpUri="http://treatment.plazi.org/id/B7C1840AFBB0593484F10AE3D37D649E">
<subSubSection id="8E8E58D0B94E5937710A540C7B61094C" type="nomenclature">
<paragraph id="01B1D88DE7BF37F92E2D689995660A32">
<heading id="5A7083AE6CD88CF5DE17DE81682C34B8" reason="label">Genus</heading>
<taxonomicName id="3F24FB7AA69D2525CB0EE0DD860D660B" ID-CoL="6NGM" authority="Walckenaer, 1826" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="CF4B75C2353CB1707A6595AFA47E0E5B" italics="true">Philodromus</emphasis>
<bibRefCitation id="C666835F63C0CA894719958B08C8C756" author="Walckenaer CA" refId="B53" refString="Walckenaer CA (1826) Araneides. In: Faune francaise ou histoire naturelle generale et particuliere des animaux qui se trouvent en France, constamment ou passagerement, a la surface du sol, dans les eaux qui le baignent et dans le littoral des mers qui le bornent par Viellot, Desmarrey, Ducrotoy, Audinet, Lepelletier et Walckenaer. Paris, livr. 11 - 12: 1 - 96." year="1826">Walckenaer, 1826</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E6MAG" type="type species">
<paragraph id="5BFD4849D6CC517547FB36D56FA37C57">
<heading id="5168D706063C8573AA961F0E34CC6714" reason="title">
<typeStatus id="B05B1600A123A30C7221FC15BBA82FE2">Type</typeStatus>
species.
</heading>
</paragraph>
<paragraph id="A612FAE8537906BACD04234C6B25FAA8">
<taxonomicName id="D28C3E5FA3EA2E677899CA86826537FF" authority="Clerck, 1757" authorityName="Clerck" authorityYear="1757" class="Arachnida" family="Araneidae" genus="Araneus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="aureolus">
<emphasis id="E8B986717AD63974638C622BCFEA5CB6" italics="true">Araneus aureolus</emphasis>
Clerck, 1757
</taxonomicName>
from Europe,
<collectingCountry id="2B3F59BCCD25641DFDE8CD4552778C67" name="Turkey">Turkey</collectingCountry>
, Caucasus,
<collectingCountry id="278F9F5E5DD0395D573942BAF35A1951" name="Russia">Russia</collectingCountry>
(Europe to Central Asia and Middle Siberia),
<collectingCountry id="5E923F34153F9FDCC184DC9D16E7C7D6" name="Kazakhstan">Kazakhstan</collectingCountry>
,
<collectingCountry id="E023CC2A01E04FAC972CFDEAA20175A2" name="Iran">Iran</collectingCountry>
, Central Asia,
<collectingCountry id="34ACF25C8492C9F0EBC2BD8A74FEB7C6" name="Mongolia">Mongolia</collectingCountry>
,
<collectingCountry id="EA530DF9272FAE04951297ED189D3C4A" name="China">China</collectingCountry>
,
<collectingCountry id="74B4F5BDB6D7D584BCCF579DABE2F541" name="South Korea">Korea</collectingCountry>
,
<collectingCountry id="AFAD237EF7ACB890090C19CC8AC0D6E6" name="Japan">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQNAG" type="diagnosis">
<paragraph id="FAAC7935FB12ED059AE49C696AD7999E">
<heading id="C30EA3DA50257CD3EAA90F2140568638" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="7A4593B595E773EE7DCBB4FF0707EA65">
See
<bibRefCitation id="6CC3EDFEFCD065056554EC5F8969C7AE" DOI="10.4039/Ent108127-2" author="Dondale CD &amp; Redner JH" firstAuthor="Dondale" issue="2" journalOrPublisher="Canadian Entomologist" pagination="127-157" refId="B16" refString="Dondale CD, Redner JH (1976) A review of the spider genus Philodromus in the Americas (Araneida: Philodromidae). Canadian Entomologist 108 (2): 127157. https://doi.org/10.4039/Ent108127-2" title="A review of the spider genus Philodromus in the Americas (Araneida: Philodromidae)." volume="108" year="1976">Dondale and Redner (1976)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E1NAG" type="comments">
<paragraph id="856526041F641C422DE8B43D22BFBA5E">
<heading id="3FAA31A176B7889413BCE497E79C5AFF" reason="title">Comments.</heading>
</paragraph>
<paragraph id="5CF78C485C9F4892C8573E07844676AE">
<taxonomicName id="B14E7262EAAF3125993B068CE732C6E6" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="F7787C03E1710B4D0740EB6996BF634F" italics="true">Philodromus</emphasis>
</taxonomicName>
is the
<typeStatus id="62481C750B81BCD2D0DAEDAE38EAB06F">type</typeStatus>
genus of
<taxonomicName id="23AC9CDE7C214A634B95642BF8AACC4E" authorityName="Thorell" authorityYear="1869" class="Arachnida" family="Philodromidae" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="family">Philodromidae</taxonomicName>
and currently includes 214 extant species that are found worldwide except for the Polar Regions (mainly distributed in the Old World and North America, except three from
<collectingCountry id="EF0562197E63A4265CB418F2952C1998" name="Australia">Australia</collectingCountry>
and only one recorded in South America respectively) (
<bibRefCitation id="8CFE949F77DF418A1C7BE440D42D63B8" refId="B57" refString="WSC (2025) World Spider Catalog. Version 25.5. Natural History Museum Bern. http://wsc.nmbe.ch [accessed on 12 January 2025]" url="http://wsc.nmbe.ch" year="2025">WSC 2025</bibRefCitation>
), 21 species of which have been recorded from
<collectingCountry id="D8CB7C1B44C8F72099F155F2045D7CC4" name="China">China</collectingCountry>
. This genus is one of the largest of
<taxonomicName id="36977E4C284ADA4EBE62AA6742048AC7" authorityName="Thorell" authorityYear="1869" class="Arachnida" family="Philodromidae" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="family">Philodromidae</taxonomicName>
and comprises 41 % of the total number of species of the family (
<bibRefCitation id="973CF2CD6146FE0B3AF729C35D7347E6" refId="B57" refString="WSC (2025) World Spider Catalog. Version 25.5. Natural History Museum Bern. http://wsc.nmbe.ch [accessed on 12 January 2025]" url="http://wsc.nmbe.ch" year="2025">WSC 2025</bibRefCitation>
).
</paragraph>
<paragraph id="C9383A73A959828C4FFF9364AE2B6B46">
Although
<taxonomicName id="8E59BC1DA5A3A948C505791489CC1792" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="3A91F0112AB6D7B66963E5E8420330E5" italics="true">Philodromus</emphasis>
</taxonomicName>
is rather well known for its high species diversity, the genus remains inadequately studied, and the species diversity is still insufficiently known (mostly related to its alpha taxonomy). The possible reasons include, but are not limited to the following: almost half of the species are described based on a single sex or juveniles (15 from males only, 79 from females only, eight from juveniles only) (
<bibRefCitation id="2614D192AFF8D9AFC3764E7AF691C5CD" refId="B57" refString="WSC (2025) World Spider Catalog. Version 25.5. Natural History Museum Bern. http://wsc.nmbe.ch [accessed on 12 January 2025]" url="http://wsc.nmbe.ch" year="2025">WSC 2025</bibRefCitation>
); for many species described in earlier studies, original descriptions are rather brief, and illustrations are absent or inadequate (
<bibRefCitation id="3D937586681A083ADA3A003BBA9D3F53" DOI="10.3969/j.issn.1005-9628.2022.02.009" author="Long" etAl="et al." firstAuthor="Long" issue="2" journalOrPublisher="Acta Arachnologica Sinica" pagination="113-120" refId="B33" refString="Long FY, Deng LJ, Mao JL, Yu H (2022) A survey of genus Philodromus (Araneae: Philodromidae) from Guiyang, Guizhou, China, with description of a new species. Acta Arachnologica Sinica 31 (2): 113120. https://doi.org/10.3969/j.issn.1005-9628.2022.02.009" title="A survey of genus Philodromus (Araneae: Philodromidae) from Guiyang, Guizhou, China, with description of a new species." volume="31" year="2022">Long et al. 2022</bibRefCitation>
;
<bibRefCitation id="9C1F8E8A7DF4E45C62D81948D56F2685" refId="B57" refString="WSC (2025) World Spider Catalog. Version 25.5. Natural History Museum Bern. http://wsc.nmbe.ch [accessed on 12 January 2025]" url="http://wsc.nmbe.ch" year="2025">WSC 2025</bibRefCitation>
); and the lack of available molecular data we can obtain
<abbrev id="ABBRID0EPPAG" xlink_title="cytochrome c oxidase subunit I">
<emphasis id="4A964F78DD3185AA1557EDCE39A888A1" italics="true">COI</emphasis>
</abbrev>
sequences for only 45 species through
<bibRefCitation id="B3E021BBCD991D68637592EA83FD82EA" refId="B40" refString="NCBI (2025) National Library of Medicine. The National Center for Biotechnology Information. https://www.ncbi.nlm.nih.gov/ [accessed on 19 January 2025]" url="https://www.ncbi.nlm.nih.gov/" year="2025">NCBI (2025</bibRefCitation>
).
</paragraph>
<paragraph id="B26FA6E012ECC2F2A6005B09373EADBA">
The insufficiency of fundamental information in alpha taxonomy poses a significant obstacle to the advancement of beta taxonomy (i. e., phylogenetic studies). Several major taxonomic studies on a regional scale have been conducted, e. g.,
<bibRefCitation id="D9926DD7C0FFB02471ACDA683514C7AC" DOI="10.4039/Ent93199-3" author="Dondale" firstAuthor="Dondale" issue="3" journalOrPublisher="Canadian Entomologist" pagination="199-222" refId="B10" refString="Dondale CD (1961) Revision of the aureolus group of the genus Philodromus (Araneae: Thomisidae) in North America. Canadian Entomologist 93 (3): 199222. https://doi.org/10.4039/Ent93199-3" title="Revision of the aureolus group of the genus Philodromus (Araneae: Thomisidae) in North America." volume="93" year="1961">Dondale (1961</bibRefCitation>
,
<bibRefCitation id="458C07220ABA05EF70BA354C656FF729" DOI="10.1155/1963/53435" author="Dondale" firstAuthor="Dondale" issue="1" journalOrPublisher="Psyche (Cambridge, Massachusetts)" pagination="34-43" refId="B11" refString="Dondale CD (1963) Florida spiders in the rufus group in the genus Philodromus (Araneae: Thomisidae). Psyche (Cambridge, Massachusetts) 70 (1): 3443. https://doi.org/10.1155/1963/53435" title="Florida spiders in the rufus group in the genus Philodromus (Araneae: Thomisidae)." volume="70" year="1963">1963</bibRefCitation>
), and
<bibRefCitation id="A09F55C972F1579217D4A9CC03ABE392" DOI="10.4039/entm10055fv" author="Dondale and Redner" firstAuthor="Dondale" refId="B13" refString="Dondale CD, Redner JH (1968) The imbecillus and rufus groups of the spider genus Philodromus in North America (Araneida: Thomisidae). Memoirs of the Entomological Society of Canada 55 (S 55): 178. https://doi.org/10.4039/entm10055fv" year="1968">Dondale and Redner (1968</bibRefCitation>
,
<bibRefCitation id="2B5F6F9E38671358AC94F87591BD0A48" DOI="10.4039/Ent101921-9" author="Dondale and Redner" firstAuthor="Dondale" issue="9" journalOrPublisher="Canadian Entomologist" pagination="921-954" refId="B14" refString="Dondale CD, Redner JH (1969) The infuscatus and dispar groups of the spider genus Philodromus in North and Central America and the West Indies (Araneida: Thomisidae). Canadian Entomologist 101 (9): 921954. https://doi.org/10.4039/Ent101921-9" title="The infuscatus and dispar groups of the spider genus Philodromus in North and Central America and the West Indies (Araneida: Thomisidae)." volume="101" year="1969">1969</bibRefCitation>
,
<bibRefCitation id="472CDDF5E3BC91DAEEA2684A4DD10A75" DOI="10.4039/Ent107369-4" author="Dondale and Redner" firstAuthor="Dondale" issue="4" journalOrPublisher="Canadian Entomologist" pagination="369-384" refId="B15" refString="Dondale CD, Redner JH (1975) The fuscomarginatus and histrio groups of the spider genus Philodromus in North America (Araneida: Thomisidae). Canadian Entomologist 107 (4): 369384. https://doi.org/10.4039/Ent107369-4" title="The fuscomarginatus and histrio groups of the spider genus Philodromus in North America (Araneida: Thomisidae)." volume="107" year="1975">1975</bibRefCitation>
,
<bibRefCitation id="21F860FD21B6EE4CE6616F6F3B5CC0E9" DOI="10.4039/Ent108127-2" author="Dondale and Redner" firstAuthor="Dondale" issue="2" journalOrPublisher="Canadian Entomologist" pagination="127-157" refId="B16" refString="Dondale CD, Redner JH (1976) A review of the spider genus Philodromus in the Americas (Araneida: Philodromidae). Canadian Entomologist 108 (2): 127157. https://doi.org/10.4039/Ent108127-2" title="A review of the spider genus Philodromus in the Americas (Araneida: Philodromidae)." volume="108" year="1976">1976</bibRefCitation>
,
<bibRefCitation id="B7E6D830B4F379B288A6BBC725D99FB0" author="Dondale and Redner" firstAuthor="Dondale" journalOrPublisher="Research Branch Agriculture Canada Publication" pagination="1-255" refId="B17" refString="Dondale CD, Redner JH (1978) The insects and arachnids of Canada, Part 5. The crab spiders of Canada and Alaska, Araneae: Philodromidae and Thomisidae. Research Branch Agriculture Canada Publication 1663: 1255." title="The insects and arachnids of Canada, Part 5. The crab spiders of Canada and Alaska, Araneae: Philodromidae and Thomisidae." volume="1663" year="1978">1978</bibRefCitation>
) for the North American species, Muster and his coauthors (
<bibRefCitation id="6F65B9DE04EE6B367C02B943E527123A" author="Muster and Thaler" editor="Thaler K" firstAuthor="Muster" journalOrPublisher="Denisia" pagination="305-326" refId="B35" refString="Muster C, Thaler K (2004) New species and records of Mediterranean Philodromidae (Arachnida, Araneae): I. Philodromus aureolus group. In: Thaler K (ed.) Diversität und Biologie von Webspinnen, Skorpionen und anderen Spinnentieren. Denisia 12: 305326." title="New species and records of Mediterranean Philodromidae (Arachnida, Araneae): I. Philodromus aureolus group." volume="12" volumeTitle="Diversität und Biologie von Webspinnen, Skorpionen und anderen Spinnentieren." year="2004">Muster and Thaler 2004</bibRefCitation>
;
<bibRefCitation id="B99BA4449E6C9A77BCAA0E2AE37308C4" DOI="10.1071/IS06014" author="Muster" etAl="et al." firstAuthor="Muster" issue="1" journalOrPublisher="Invertebrate Systematics" pagination="39-72" refId="B36" refString="Muster C, Bosmans R, Thaler K (2007) The Philodromus pulchellus - group in the Mediterranean: taxonomic revision, phylogenetic analysis and biogeography (Araneae: Philodromidae). Invertebrate Systematics 21 (1): 3972. https://doi.org/10.1071/IS06014" title="The Philodromus pulchellus - group in the Mediterranean: taxonomic revision, phylogenetic analysis and biogeography (Araneae: Philodromidae)." volume="21" year="2007">Muster et al. 2007</bibRefCitation>
;
<bibRefCitation id="06DD1D6BCFF1D854715A4BF19F23771A" DOI="10.1071/IS08044" author="Muster" firstAuthor="Muster" issue="2" journalOrPublisher="Invertebrate Systematics" pagination="135-169" refId="B34" refString="Muster C (2009) Phylogenetic relationships within Philodromidae, with a taxonomic revision of Philodromus subgenus Artanes in the western Palearctic (Arachnida: Araneae). Invertebrate Systematics 23 (2): 135169. https://doi.org/10.1071/IS08044" title="Phylogenetic relationships within Philodromidae, with a taxonomic revision of Philodromus subgenus Artanes in the western Palearctic (Arachnida: Araneae)." volume="23" year="2009">Muster 2009</bibRefCitation>
), and
<bibRefCitation id="CEAE5F50BE82E669512B957D232DB15C" author="Wunderlich J" firstAuthor="Wunderlich" journalOrPublisher="Beiträge zur Araneologie" pagination="1-246" refId="B58" refString="Wunderlich J (2012) Fifteen papers on extant and fossil spiders (Araneae). Beiträge zur Araneologie 7: 1246." title="Fifteen papers on extant and fossil spiders (Araneae)." volume="7" year="2012">Wunderlich (2012)</bibRefCitation>
for the European species; however, these revisionary studies often exclude species from Asia (potentially due to the aforementioned lack of foundational taxonomic information), and the debate on the groups limits and internal structure of this family remains open (
<bibRefCitation id="EA4931464A5116FFCB9DF1C4CF88565D" refId="B57" refString="WSC (2025) World Spider Catalog. Version 25.5. Natural History Museum Bern. http://wsc.nmbe.ch [accessed on 12 January 2025]" url="http://wsc.nmbe.ch" year="2025">WSC 2025</bibRefCitation>
). According to the quite diverse copulatory structures of both sexes, different camouflage behaviours and habitat preferences,
<taxonomicName id="A102B0E7F5FA261BAA02E8B3090276B2" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="06DB3F2DC99818442D2E6FB2989F5EC0" italics="true">Philodromus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EAD43871FD1486C9F620BECE60719E42" sensu="lato">sensu lato</taxonomicNameLabel>
has been regarded as paraphyletic and needs to be split (
<bibRefCitation id="674AC5B8918F372865C2EC230D529751" author="Wunderlich" firstAuthor="Wunderlich" journalOrPublisher="Beiträge zur Araneologie" pagination="1-246" refId="B58" refString="Wunderlich J (2012) Fifteen papers on extant and fossil spiders (Araneae). Beiträge zur Araneologie 7: 1246." title="Fifteen papers on extant and fossil spiders (Araneae)." volume="7" year="2012">Wunderlich 2012</bibRefCitation>
). However, we agree with
<bibRefCitation id="C0BF811704DB94DB332978BAF4712534" DOI="10.1071/IS08044" author="Muster C" firstAuthor="Muster" issue="2" journalOrPublisher="Invertebrate Systematics" pagination="135-169" refId="B34" refString="Muster C (2009) Phylogenetic relationships within Philodromidae, with a taxonomic revision of Philodromus subgenus Artanes in the western Palearctic (Arachnida: Araneae). Invertebrate Systematics 23 (2): 135169. https://doi.org/10.1071/IS08044" title="Phylogenetic relationships within Philodromidae, with a taxonomic revision of Philodromus subgenus Artanes in the western Palearctic (Arachnida: Araneae)." volume="23" year="2009">Muster (2009)</bibRefCitation>
that the elevation of an autapomorphic species group would render
<taxonomicName id="8363D64E043291C844C3331FE3E9ED5F" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="AE664FB27EF011BCD0A8964E135A685C" italics="true">Philodromus</emphasis>
</taxonomicName>
paraphyletic and in need of an extensive, large-scale review of the genus. Consequently, the present study follows the
<bibRefCitation id="B897F8097338695257D4EF636C4BD06B" refId="B57" refString="WSC (2025) World Spider Catalog. Version 25.5. Natural History Museum Bern. http://wsc.nmbe.ch [accessed on 12 January 2025]" url="http://wsc.nmbe.ch" year="2025">WSC (2025</bibRefCitation>
), and places all treated species in
<taxonomicName id="B81EA4A47BEA94191DC610DF0EC40151" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="AB992D200362D3A9138F990494AC547B" italics="true">Philodromus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9621937C20705160E940CAD29394B7AE" sensu="lato">sensu lato</taxonomicNameLabel>
. We provide only the fundamental information on these species (such as detailed descriptions, supplementary illustrations, and DNA barcodes) for species delimitation, matching of sexes and future use. A review of the genus is not within the scope of this work.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,95 @@
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<paragraph id="2DB7B2272DAA78B636DD0296C68C482A">
<heading id="8CD443274C0BC39CFA659735C98A986A" reason="label">Genus</heading>
<taxonomicName id="853AE33FC315A6EF6D18B08C7689721D" ID-CoL="CVWP6" authority="Yao &amp; Liu, 2024" authorityName="Yao &amp; Liu" authorityYear="2024" class="Arachnida" family="Philodromidae" genus="Sinodromus" higherTaxonomySource="CoL" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="229E523C091849F62EE852B411E50B9A" italics="true">Sinodromus</emphasis>
Yao &amp; Liu, 2024
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<typeStatus id="8AD34B1F6731117DBA6BB9357F6573B1">Type</typeStatus>
species.
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<paragraph id="D0326E4055DFCA260FE971CCF851FC15">
<taxonomicName id="ED0628FA26A3BF3BFD1EE197DFB137AA" authority="Yao &amp; Liu, 2024" authorityName="Yao &amp; Liu" authorityYear="2024" class="Arachnida" family="Philodromidae" genus="Sinodromus" higherTaxonomySource="CoL" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="fujianensis">
<emphasis id="92D1021C4BEDE5D04A2628DDF3A3B6A6" italics="true">Sinodromus fujianensis</emphasis>
Yao &amp; Liu, 2024
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from
<collectingRegion id="F1125DE8BE7DF857C913B41B60532D86" country="China" name="Fujian">Fujian</collectingRegion>
and
<collectingRegion id="C20BE8888F7C3E1C93319283EB37380C" country="China" name="Jiangxi">Jiangxi</collectingRegion>
provinces,
<collectingCountry id="65799B80C64A5CE36F88B3DEEA41A7DB" name="China">China</collectingCountry>
.
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<subSubSection id="SECID0EYVBI" type="diagnosis">
<paragraph id="A5FC0311DAF4AA8C8BA68963DE9879A1">
<heading id="D797261CE1ADA7C721634F62123FD4BC" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="82573BE1ACC6032E2390B1A0792936C4">
See
<bibRefCitation id="EB20F6C247A39011D88C0B43BB22A2C9" DOI="10.3897/zookeys.1221.137930" author="Wang ZJ &amp; Yao YB &amp; Tang ZY &amp; Li WH &amp; Liu KK &amp; Xu X" etAl="et al." firstAuthor="Wang" journalOrPublisher="ZooKeys" pagination="279-296" refId="B55" refString="Wang ZJ, Yao YB, Tang ZY, Li WH, Liu KK, Xu X (2024) A new genus, Sinodromus gen. nov., with two new species and the first description of the female of Philodromus guiyang Long &amp; Yu, 2022 (Arachnida, Araneae, Philodromidae) from China. ZooKeys 296: 279296. https://doi.org/10.3897/zookeys.1221.137930" title="A new genus, Sinodromus gen. nov., with two new species and the first description of the female of Philodromus guiyang Long &amp; Yu, 2022 (Arachnida, Araneae, Philodromidae) from China." volume="296" year="2024">Wang et al. (2024)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,732 @@
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<emphasis id="8881AD90C1EEBAD21C6C8C9DA853DFEF" italics="true">Sinodromus lanyue</emphasis>
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<taxonomicNameLabel id="3CF8C389EB7B98A90D358396110819E7" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="38AB25CB06973C4921325C42A9FF5144">
<figureCitation id="E5116A2CD7EFF5AA57E2907F20D7E1D5" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Sample collection locations for this project. Hubei Province 1 Jiugongshan National Nature Reserve 2 Hubei University of Science and Technology 3 Jiuling Mountain 4 Qizimei Mountain National Nature Reserve; Guizhou Province 5 Fanjingshan National Nature Reserve 6 Mayanghe National Nature Reserve 7 Xishui National Nature Reserve 8 Guiyang City, Kaiyang County 9 Guiyang City, Wudang District 10 Guiyang City, Yunyan District 11 Guiyang City, Huaxi District." figureDoi="10.3897/zookeys.1240.149456.figure1" httpUri="https://binary.pensoft.net/fig/1348759">Figs 1</figureCitation>
,
<figureCitation id="E7D5E83F3FBC572752A486EC48B84ABC" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Living specimens of some Philodromidae species treated in this paper A, B Sinodromus lanyue sp. nov. (♂ ♀) C, D Tibellus japonicus (♂ ♀). Photographs by Q. Lu (Shenzhen)." figureDoi="10.3897/zookeys.1240.149456.figure4" httpUri="https://binary.pensoft.net/fig/1348762">4 A, B</figureCitation>
,
<figureCitation id="F761AED9F4D1874434AA5985ADF16D2B" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Sinodromus lanyue sp. nov., holotype male, left palp A ventral view B dorsal view. Abbreviations: Con = conductor; CP = cymbial process; Cy = cymbium; Em = embolus; SDL = sperm duct loop; TA = tegular apophysis; Te = tegulum; VPTA = ventro-prolateral tibial apophysis; vRTA = ventral branch of RTA. Scale bar: 0.2 mm (equal for A, B)." figureDoi="10.3897/zookeys.1240.149456.figure16" httpUri="https://binary.pensoft.net/fig/1348774">16</figureCitation>
,
<figureCitation id="7062457A519B13C0D16CD875FEA938F3" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Sinodromus lanyue sp. nov., holotype male, left palp A prolateral view B retrolateral view. Abbreviations: Con = conductor; CP = cymbial process; Cy = cymbium; dRTA = dorsal branch of RTA; EmT = embolic tip; SD = sperm duct; SDL = sperm duct loop; TA = tegular apophysis; Te = tegulum; VPTA = ventro-prolateral tibial apophysis; vRTA = ventral branch of RTA. Scale bar: 0.2 mm (equal for A, B)." figureDoi="10.3897/zookeys.1240.149456.figure17" httpUri="https://binary.pensoft.net/fig/1348775">17</figureCitation>
,
<figureCitation id="7F555E10DDE6473CFB0DEA6F34CA7BF9" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Sinodromus lanyue sp. nov., holotype male, left palpal bulb A ventral view B dorsal view C prolateral view D retrolateral view. Abbreviations: Con = conductor; Em = embolus; SD = sperm duct; SDL = sperm duct loop; TA = tegular apophysis; Te = tegulum. Scale bar: 0.2 mm (equal for A D)." figureDoi="10.3897/zookeys.1240.149456.figure18" httpUri="https://binary.pensoft.net/fig/1348776">18</figureCitation>
,
<figureCitation id="A65E58A6D14BCBC3E94E9C2F1E6D0CDF" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Sinodromus lanyue sp. nov., paratype female (A E, G) and holotype male (F), epigyne (A E) and frontal view of carapace (F, G) A macerated, ventral view B macerated, dorsal view C macerated and embedded in arabic gum, ventral view D macerated and embedded in arabic gum, dorsal view E intact, ventral view F male G female. Abbreviations: A = atrium; cCD = course of copulatory duct (red line); CD = copulatory duct; CO = copulatory opening; ET = epigynal tooth; FD = fertilisation duct; GM = glandular mound; MS = median septum; R = receptaculum; rMS = rim of median septum (teal green line); rSEF = rim of sclerotised epigynal fold (blue line); SEF = sclerotised epigynal fold. Scale bars: 0.2 mm (A E); 0.5 mm (equal for F, G)." figureDoi="10.3897/zookeys.1240.149456.figure19" httpUri="https://binary.pensoft.net/fig/1348777">19</figureCitation>
,
<figureCitation id="FC8884DED6D9A3D238F459D2DEBBC173" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Sinodromus lanyue sp. nov., holotype male (A C) and paratype female (D F), habitus A, D dorsal view B, E ventral view C, F lateral view Scale bars: 1 mm (equal for A C, equal for D F)." figureDoi="10.3897/zookeys.1240.149456.figure20" httpUri="https://binary.pensoft.net/fig/1348778">20</figureCitation>
,
<figureCitation id="E5161C073D27CD38B7A457FC060BF2AB" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Distribution records of the philodromid species in Guizhou Province (A) and Hubei Province (B)." figureDoi="10.3897/zookeys.1240.149456.figure23" httpUri="https://binary.pensoft.net/fig/1348781">23 B</figureCitation>
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<heading id="568152462865F7FA23131213F3F2C51D" reason="title">
<typeStatus id="F8A92D24D93D3912C023900A293EECBD">Type</typeStatus>
material.
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<paragraph id="A92E2654E542630DD95954517BC41580">
<materialsCitation id="9F35873509FDE58FD0DE3C6F54C6149E" collectingDate="2023-03-22" collectionCode="MGNU" collectorName="Y. Zhong &amp; Q. Lu" country="China" county="Xianning City" latitude="29.85" location="University of Science" longLatPrecision="734" longitude="114.34" municipality="Xianan District" specimenCode="PHISL 001" specimenCount="1" specimenCount-male="1" stateProvince="Hubei" typeStatus="holotype">
<emphasis id="48AD763AF29B75CDC929901FBF35224C" bold="true">
<collectingCountry id="6E3BC5148D255EDB393DE4195D6C9714" name="China">China</collectingCountry>
</emphasis>
:
<collectingRegion id="1418858E13888066A8D0BBDE8B718999" country="China" name="Hubei">Hubei Province</collectingRegion>
:
<emphasis id="C86F822960A4A1FD3E05358B86AA8825" bold="true" italics="true">
<typeStatus id="C6D98A07C428E0D0E7E8F61D8F96571D" type="holotype">Holotype</typeStatus>
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<specimenCount id="616AAEA76CFEBAC64BDCA57F953941E3" count="1" type="male"></specimenCount>
:
<collectingCounty id="B9936C8F5AFD77E800CDEC1AD5FF06C7">Xianning City</collectingCounty>
,
<collectingMunicipality id="03D42B6853F2F1327CB8BA43BDD13E1D">Xianan District</collectingMunicipality>
,
<collectingRegion id="22218FD6ABEBF59CBC263B151C29B4B4" country="China" name="Hubei">Hubei</collectingRegion>
<location id="ED237725C9C229737E4216DE009420D1" LSID="urn:lsid:plazi:treatment:FAD04B0075A15956A2F39A3A4EBABB51:ED237725C9C229737E4216DE009420D1" country="China" county="Xianning City" latitude="29.85" longLatPrecision="734" longitude="114.34" municipality="Xianan District" name="University of Science" stateProvince="Hubei">University of Science</location>
and
<location id="27CF0F9BD66E134360F43C18BF5D6A0C" LSID="urn:lsid:plazi:treatment:FAD04B0075A15956A2F39A3A4EBABB51:27CF0F9BD66E134360F43C18BF5D6A0C" country="China" county="Xianning City" latitude="29.85" longLatPrecision="734" longitude="114.34" municipality="Xianan District" name="Technology" stateProvince="Hubei">Technology</location>
, the bamboo forest on the hill behind
<location id="058ADD65AF14EC1F9FEB6705737673E5" LSID="urn:lsid:plazi:treatment:FAD04B0075A15956A2F39A3A4EBABB51:058ADD65AF14EC1F9FEB6705737673E5" country="China" county="Xianning City" latitude="29.85" longLatPrecision="734" longitude="114.34" municipality="Xianan District" name="Lanyue Lake" stateProvince="Hubei">Lanyue Lake</location>
;
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<geoCoordinate id="A999423B59A4B3AB12E9D02BBDBE0910" degrees="29.85" direction="north" orientation="latitude" precision="555" value="29.85">29.85 ° N</geoCoordinate>
,
<geoCoordinate id="767EDEADE1D08B16B74297B504F6B581" degrees="114.34" direction="east" orientation="longitude" precision="555" value="114.34">114.34 ° E</geoCoordinate>
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;
<date id="9E8D65C5002245879DC8C811444D4174" value="2023-03-22">
<collectingDate id="5CB5B7F061387A3258FBB611D86F36DB" value="2023-03-22">22 March 2023</collectingDate>
</date>
;
<collectorName id="61AD54280B41AC6ABD7B9EF33548C1A5">Y. Zhong</collectorName>
&amp;
<collectorName id="A06DD3B20929789A96DF799EBED8E8D2">Q. Lu</collectorName>
leg. (Inventory number:
<named-content id="CD2D0E38CB2572AC6F4240F9048FC205" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/guizhou-normal-university" xlink_title="Museum of Guizhou Normal University">
<collectionCode id="D14D40D27258AA69715EADBD92B79E43">MGNU</collectionCode>
</named-content>
- 2025 -
<specimenCode id="CF77EA3791DC5D03BE7781F8C7CBAB3D" collectionCode="PHISL">PHISL 001</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="4DDD82C245D376931113C4605C5787CB" collectingDate="2023-03-22" collectionCode="MGNU" collectorName="Y. Zhong &amp; Q. Lu" country="China" county="Xianning City" latitude="29.85" location="University of Science" longLatPrecision="734" longitude="114.34" municipality="Xianan District" specimenCode="PHISL 002" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Paratypes" typeStatus="holotype">
<emphasis id="48BD3C40E1140A152BE4BF76F0937DF1" bold="true" italics="true">
<typeStatus id="891DF42251BA3EE53BA8FCD47E441539" type="paratype">
<collectingRegion id="84E72CB817E8FDA42937CAF2377829DB">Paratypes</collectingRegion>
</typeStatus>
</emphasis>
<specimenCount id="163CA8340977681B28E551AABEB61B78" count="1" type="male">1 ♂</specimenCount>
,
<specimenCount id="752B16709BC07AC3EF922ACCF3E2D848" count="1" type="female">1 ♀</specimenCount>
(Inventory number:
<named-content id="4310EF1FD27FF8A6768D852F6C5D8BF6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/guizhou-normal-university" xlink_title="Museum of Guizhou Normal University">
<collectionCode id="42F5EEE4C00DC6F1184B94F9C1061CFD">MGNU</collectionCode>
</named-content>
- 2025 -
<specimenCode id="21A434FD706C4991CE24506FB983CC7B" collectionCode="PHISL">PHISL 002</specimenCode>
~ 003), the same data as the holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJZBI" type="material">
<paragraph id="579068E2765FD92027E7D0B8F0F9AF81">
<heading id="FEB763B6C25643E7C237316775E0C298" reason="title">Other material examined.</heading>
</paragraph>
<paragraph id="12C187B9CAF188C2039BEF36C4F10674">
<specimenCount id="AC162A1D40ECE6F07190C0A9C9790CD5" count="1" type="male">1 ♂</specimenCount>
,
<specimenCount id="ECE3EFFAD1742BAB8483B5E5998DA962" count="1" type="female">1 ♀</specimenCount>
(
<collectionCode id="E29F2D78369CD42801B00FFD40FDC424">YHPHI</collectionCode>
008 and
<collectionCode id="A5E81F7925D31A229F9514219F353DD5">YHPHI</collectionCode>
009 used for sequencing, GenBank accession numbers in Table
<tableCitation id="BDFD87D35ED3A06AFFD4107BA30F85B2" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Voucher specimen information." httpUri="http://table.plazi.org/id/12F54BE0BDE6512C62E0465BE733F517" tableUuid="12F54BE0BDE6512C62E0465BE733F517">1</tableCitation>
), the same data as the
<typeStatus id="20AF8DCDDDB9F0AA657025689C8A5785" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ETZBI" type="etymology">
<paragraph id="41F1055190EA15345D2DAE7AC77FEE34">
<heading id="5F895DDBCF7375D2D07D3A4E85563EA2" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="3FE2EFC3729E8D2D352507A1D8E184A0">
The species name is derived from the
<typeStatus id="33FD45813AA2EDAE48B76A1CBBD54410">type</typeStatus>
locality; noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EYZBI" type="diagnosis">
<paragraph id="100D70FAE1EA258C136A34BEB1C2F4F0">
<heading id="071496F95B06F200D22B39C4470B0A3F" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="0E4A1CF2F453EE3EA0BDFE3C3EEE2DB3">
Males of the new species are easily distinguished from
<taxonomicName id="0F4219F7F053BF440D66CA6203D23374" authority="Yao &amp; Liu, 2024" authorityName="Yao &amp; Liu" authorityYear="2024" class="Arachnida" family="Philodromidae" genus="Sinodromus" higherTaxonomySource="CoL" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="fujianensis">
<emphasis id="1819BCA411AF1D242BF821A7E7D51834" italics="true">Sinodromus fujianensis</emphasis>
Yao &amp; Liu, 2024
</taxonomicName>
(only congener with described male) by the following combination of morphological characteristics: (1)
<abbrev id="ABBRID0EJ1BI" xlink_title="tegular apophysis">
<collectionCode id="2BB1CC894C1C4B31B0F5D97FB1D72A76" country="0" name="Timescale Adventures Research and Interpretive Center">TA</collectionCode>
</abbrev>
shaped like a cocks head, with a hump-like, not folded basal apophysis (vs horn-shaped, with a lamellar, folded basal apophysis) (cf. Figs
<figureCitation id="C07A04E3C9119566F2BE3200FA543AEB" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Sinodromus lanyue sp. nov., holotype male, left palp A ventral view B dorsal view. Abbreviations: Con = conductor; CP = cymbial process; Cy = cymbium; Em = embolus; SDL = sperm duct loop; TA = tegular apophysis; Te = tegulum; VPTA = ventro-prolateral tibial apophysis; vRTA = ventral branch of RTA. Scale bar: 0.2 mm (equal for A, B)." figureDoi="10.3897/zookeys.1240.149456.figure16" httpUri="https://binary.pensoft.net/fig/1348774">16 A</figureCitation>
,
<figureCitation id="94E4068657386885F8498566CE78B430" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Sinodromus lanyue sp. nov., holotype male, left palp A prolateral view B retrolateral view. Abbreviations: Con = conductor; CP = cymbial process; Cy = cymbium; dRTA = dorsal branch of RTA; EmT = embolic tip; SD = sperm duct; SDL = sperm duct loop; TA = tegular apophysis; Te = tegulum; VPTA = ventro-prolateral tibial apophysis; vRTA = ventral branch of RTA. Scale bar: 0.2 mm (equal for A, B)." figureDoi="10.3897/zookeys.1240.149456.figure17" httpUri="https://binary.pensoft.net/fig/1348775">17 A, B</figureCitation>
,
<figureCitation id="43209B06D89EBC45905068736E80838A" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Sinodromus lanyue sp. nov., holotype male, left palpal bulb A ventral view B dorsal view C prolateral view D retrolateral view. Abbreviations: Con = conductor; Em = embolus; SD = sperm duct; SDL = sperm duct loop; TA = tegular apophysis; Te = tegulum. Scale bar: 0.2 mm (equal for A D)." figureDoi="10.3897/zookeys.1240.149456.figure18" httpUri="https://binary.pensoft.net/fig/1348776">18 A, C, D</figureCitation>
and
<bibRefCitation id="57FDFB478D5E55C0886189B586D1FC05" DOI="10.3897/zookeys.1221.137930" author="Wang" etAl="et al." firstAuthor="Wang" journalOrPublisher="ZooKeys" pagination="279-296" refId="B55" refString="Wang ZJ, Yao YB, Tang ZY, Li WH, Liu KK, Xu X (2024) A new genus, Sinodromus gen. nov., with two new species and the first description of the female of Philodromus guiyang Long &amp; Yu, 2022 (Arachnida, Araneae, Philodromidae) from China. ZooKeys 296: 279296. https://doi.org/10.3897/zookeys.1221.137930" title="A new genus, Sinodromus gen. nov., with two new species and the first description of the female of Philodromus guiyang Long &amp; Yu, 2022 (Arachnida, Araneae, Philodromidae) from China." volume="296" year="2024">Wang et al. 2024</bibRefCitation>
: figs 4 B E, 5 C, G, I, K); and (2)
<abbrev id="ABBRID0E41BI" xlink_title="conductor">Con</abbrev>
wider than
<abbrev id="ABBRID0EB2BI" xlink_title="tegular apophysis">
<collectionCode id="1DAF36FBE5F078B7E509E03481095A65" country="0" name="Timescale Adventures Research and Interpretive Center">TA</collectionCode>
</abbrev>
, surface relatively smooth (vs narrower than
<abbrev id="ABBRID0EF2BI" xlink_title="tegular apophysis">
<collectionCode id="47BB42B1905DEC41A80D5940559C94C2" country="0" name="Timescale Adventures Research and Interpretive Center">TA</collectionCode>
</abbrev>
, with many scaly serrations) (cf. Figs
<figureCitation id="A2CFCD42505004D708C34D5D2D792845" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Sinodromus lanyue sp. nov., holotype male, left palp A ventral view B dorsal view. Abbreviations: Con = conductor; CP = cymbial process; Cy = cymbium; Em = embolus; SDL = sperm duct loop; TA = tegular apophysis; Te = tegulum; VPTA = ventro-prolateral tibial apophysis; vRTA = ventral branch of RTA. Scale bar: 0.2 mm (equal for A, B)." figureDoi="10.3897/zookeys.1240.149456.figure16" httpUri="https://binary.pensoft.net/fig/1348774">16 A</figureCitation>
,
<figureCitation id="8A664924A828586F9D5BDD4E79876C1B" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Sinodromus lanyue sp. nov., holotype male, left palp A prolateral view B retrolateral view. Abbreviations: Con = conductor; CP = cymbial process; Cy = cymbium; dRTA = dorsal branch of RTA; EmT = embolic tip; SD = sperm duct; SDL = sperm duct loop; TA = tegular apophysis; Te = tegulum; VPTA = ventro-prolateral tibial apophysis; vRTA = ventral branch of RTA. Scale bar: 0.2 mm (equal for A, B)." figureDoi="10.3897/zookeys.1240.149456.figure17" httpUri="https://binary.pensoft.net/fig/1348775">17 A, B</figureCitation>
,
<figureCitation id="4ECA2EC02CDE7F804C00F2728B117A9D" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Sinodromus lanyue sp. nov., holotype male, left palpal bulb A ventral view B dorsal view C prolateral view D retrolateral view. Abbreviations: Con = conductor; Em = embolus; SD = sperm duct; SDL = sperm duct loop; TA = tegular apophysis; Te = tegulum. Scale bar: 0.2 mm (equal for A D)." figureDoi="10.3897/zookeys.1240.149456.figure18" httpUri="https://binary.pensoft.net/fig/1348776">18 A D</figureCitation>
and
<bibRefCitation id="43990C7E496B9BB62EAC20D12877EBF5" DOI="10.3897/zookeys.1221.137930" author="Wang" etAl="et al." firstAuthor="Wang" journalOrPublisher="ZooKeys" pagination="279-296" refId="B55" refString="Wang ZJ, Yao YB, Tang ZY, Li WH, Liu KK, Xu X (2024) A new genus, Sinodromus gen. nov., with two new species and the first description of the female of Philodromus guiyang Long &amp; Yu, 2022 (Arachnida, Araneae, Philodromidae) from China. ZooKeys 296: 279296. https://doi.org/10.3897/zookeys.1221.137930" title="A new genus, Sinodromus gen. nov., with two new species and the first description of the female of Philodromus guiyang Long &amp; Yu, 2022 (Arachnida, Araneae, Philodromidae) from China." volume="296" year="2024">Wang et al. 2024</bibRefCitation>
: figs 4 B E, 5 C, G, I, K). Females of the new species resemble those of
<taxonomicName id="DFFA6977D1EFD3A34A48A064D450E921" authority="Yao &amp; Liu, 2024" authorityName="Yao &amp; Liu" authorityYear="2024" class="Arachnida" family="Philodromidae" genus="Sinodromus" higherTaxonomySource="CoL" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="perbrevis">
<emphasis id="7C715C73CA53460D09DC76E069ECE539" italics="true">S. perbrevis</emphasis>
Yao &amp; Liu,
<quantity id="FF9B6EE65755A667109FC5ED36481483" metricMagnitude="1" metricUnit="m" metricValue="5.14096" unit="in" value="2024.0">2024 in</quantity>
</taxonomicName>
having a similar
<abbrev id="ABBRID0EE3BI" xlink_title="median septum">MS</abbrev>
and endogyne but can be recognised by: (1)
<abbrev id="ABBRID0EI3BI" xlink_title="epigynal tooth">
<collectionCode id="E125C7263BF81DA429FB27C22CE1E514" country="0" name="East Texas State University">ET</collectionCode>
</abbrev>
axe-shaped, distinctly widened, wider than midsection of
<abbrev id="ABBRID0EM3BI" xlink_title="sclerotised epigynal fold">SEF</abbrev>
(vs ear-shaped, not widened, nearly as wide as midsection of
<abbrev id="ABBRID0EQ3BI" xlink_title="sclerotised epigynal fold">SEF</abbrev>
) (cf. Fig.
<figureCitation id="AEA3863FF9154E8E6258AFAFBF4EEA65" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Sinodromus lanyue sp. nov., paratype female (A E, G) and holotype male (F), epigyne (A E) and frontal view of carapace (F, G) A macerated, ventral view B macerated, dorsal view C macerated and embedded in arabic gum, ventral view D macerated and embedded in arabic gum, dorsal view E intact, ventral view F male G female. Abbreviations: A = atrium; cCD = course of copulatory duct (red line); CD = copulatory duct; CO = copulatory opening; ET = epigynal tooth; FD = fertilisation duct; GM = glandular mound; MS = median septum; R = receptaculum; rMS = rim of median septum (teal green line); rSEF = rim of sclerotised epigynal fold (blue line); SEF = sclerotised epigynal fold. Scale bars: 0.2 mm (A E); 0.5 mm (equal for F, G)." figureDoi="10.3897/zookeys.1240.149456.figure19" httpUri="https://binary.pensoft.net/fig/1348777">19 A, C, E</figureCitation>
and
<bibRefCitation id="F0E71E3A8EF4501DFFC3210866D632E0" DOI="10.3897/zookeys.1221.137930" author="Wang" etAl="et al." firstAuthor="Wang" journalOrPublisher="ZooKeys" pagination="279-296" refId="B55" refString="Wang ZJ, Yao YB, Tang ZY, Li WH, Liu KK, Xu X (2024) A new genus, Sinodromus gen. nov., with two new species and the first description of the female of Philodromus guiyang Long &amp; Yu, 2022 (Arachnida, Araneae, Philodromidae) from China. ZooKeys 296: 279296. https://doi.org/10.3897/zookeys.1221.137930" title="A new genus, Sinodromus gen. nov., with two new species and the first description of the female of Philodromus guiyang Long &amp; Yu, 2022 (Arachnida, Araneae, Philodromidae) from China." volume="296" year="2024">Wang et al. 2024</bibRefCitation>
: figs 7 C, 8 C); and (2)
<collectionCode id="B26B5793FAC67D9281BD52FFD181C56E" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
oval, close together (vs globular, widely separated) (cf. Fig.
<figureCitation id="5C3D78EC803D7D29AEA8F1CC158CBB0E" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Sinodromus lanyue sp. nov., paratype female (A E, G) and holotype male (F), epigyne (A E) and frontal view of carapace (F, G) A macerated, ventral view B macerated, dorsal view C macerated and embedded in arabic gum, ventral view D macerated and embedded in arabic gum, dorsal view E intact, ventral view F male G female. Abbreviations: A = atrium; cCD = course of copulatory duct (red line); CD = copulatory duct; CO = copulatory opening; ET = epigynal tooth; FD = fertilisation duct; GM = glandular mound; MS = median septum; R = receptaculum; rMS = rim of median septum (teal green line); rSEF = rim of sclerotised epigynal fold (blue line); SEF = sclerotised epigynal fold. Scale bars: 0.2 mm (A E); 0.5 mm (equal for F, G)." figureDoi="10.3897/zookeys.1240.149456.figure19" httpUri="https://binary.pensoft.net/fig/1348777">19 B, D</figureCitation>
and
<bibRefCitation id="19FAD714B3A34EC6559E43E35E65B46C" DOI="10.3897/zookeys.1221.137930" author="Wang" etAl="et al." firstAuthor="Wang" journalOrPublisher="ZooKeys" pagination="279-296" refId="B55" refString="Wang ZJ, Yao YB, Tang ZY, Li WH, Liu KK, Xu X (2024) A new genus, Sinodromus gen. nov., with two new species and the first description of the female of Philodromus guiyang Long &amp; Yu, 2022 (Arachnida, Araneae, Philodromidae) from China. ZooKeys 296: 279296. https://doi.org/10.3897/zookeys.1221.137930" title="A new genus, Sinodromus gen. nov., with two new species and the first description of the female of Philodromus guiyang Long &amp; Yu, 2022 (Arachnida, Araneae, Philodromidae) from China." volume="296" year="2024">Wang et al. 2024</bibRefCitation>
: fig. 7 D).
</paragraph>
<caption id="D879C24EF7BB6ED9945E3EB5EFF579FC" ID-DOI="10.3897/zookeys.1240.149456.figure16" ID-arpha="3320D463-07F6-507E-BA22-A3F15CAAF840" httpUri="https://binary.pensoft.net/fig/1348774" startId="F16">
<paragraph id="CD21B49275FC5B63777677E6E8F26C54">
<label id="D0BFD5D6AB97C77D46AE74A9451DD36D">Figure 16.</label>
</paragraph>
<paragraph id="6E112D90EBE91595BAFEFBC7B47AF018">
<taxonomicName id="8F0F83ADC3F44A3007F105D4F3C8ECE6" authorityName="Zhang &amp; Zhang &amp; Zhong" authorityYear="2025" class="Arachnida" family="Philodromidae" genus="Sinodromus" higherTaxonomySource="CoL" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="lanyue" status="sp. nov.">
<emphasis id="98BF21AC35AAD8494FA7CC0462ECFFA3" italics="true">Sinodromus lanyue</emphasis>
</taxonomicName>
<taxonomicNameLabel id="111E0906281B2ED3AC7C29F22F81DD99" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male, left palp
<emphasis id="4FE59CC878630C3D82F062B93EE5F317" bold="true">A</emphasis>
ventral view
<emphasis id="9314176EB5A4EF363B7785E38C0D7709" bold="true">B</emphasis>
dorsal view. Abbreviations:
<abbrev id="ABBRID0E24BI" xlink_title="conductor">Con</abbrev>
= conductor;
<abbrev id="ABBRID0E64BI" xlink_title="cymbial process">CP</abbrev>
= cymbial process;
<abbrev id="ABBRID0ED5BI" xlink_title="cymbium">Cy</abbrev>
= cymbium;
<abbrev id="ABBRID0EH5BI" xlink_title="embolus">Em</abbrev>
= embolus;
<abbrev id="ABBRID0EL5BI" xlink_title="sperm duct loop">SDL</abbrev>
= sperm duct loop;
<abbrev id="ABBRID0EP5BI" xlink_title="tegular apophysis">TA</abbrev>
= tegular apophysis;
<abbrev id="ABBRID0ET5BI" xlink_title="tegulum">Te</abbrev>
= tegulum;
<abbrev id="ABBRID0EX5BI" xlink_title="ventro-prolateral tibial apophysis">VPTA</abbrev>
= ventro-prolateral tibial apophysis;
<abbrev id="ABBRID0E25BI" xlink_title="ventral branch of RTA">vRTA</abbrev>
= ventral branch of
<abbrev id="ABBRID0E65BI" xlink_title="retrolateral tibial apophysis">RTA</abbrev>
. Scale bar: 0.2 mm (equal for
<emphasis id="703160A3BD5F50878298267BC371FC55" bold="true">A, B</emphasis>
).
</paragraph>
</caption>
<caption id="9E5FC9CA7A1F4AC52A44B2D4163C1570" ID-DOI="10.3897/zookeys.1240.149456.figure17" ID-arpha="3607FCE5-17D3-5024-9FA5-E6EE659A1E13" httpUri="https://binary.pensoft.net/fig/1348775" startId="F17">
<paragraph id="8EF0993C0DEC1F81B528749C636052B5">
<label id="1C59FC3FD57F90FE545B27FD7645DC00">Figure 17.</label>
</paragraph>
<paragraph id="47249C2CE2B058BDAE72207DC7926E4B">
<taxonomicName id="7BE0708BA731DB52BD9CB7DDF18EBB85" authorityName="Zhang &amp; Zhang &amp; Zhong" authorityYear="2025" class="Arachnida" family="Philodromidae" genus="Sinodromus" higherTaxonomySource="CoL" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="lanyue" status="sp. nov.">
<emphasis id="D8313E24F71EE3008290B8D7F9E27B6D" italics="true">Sinodromus lanyue</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C647FFA55C3392E0FB1A870839CCC86E" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male, left palp
<emphasis id="981EFFE48A80D9B2F135F49E78A8260A" bold="true">A</emphasis>
prolateral view
<emphasis id="6CC02AA36E8EF9D19C7984EC6DBD8993" bold="true">B</emphasis>
retrolateral view. Abbreviations:
<abbrev id="ABBRID0EBACI" xlink_title="conductor">Con</abbrev>
= conductor;
<abbrev id="ABBRID0EFACI" xlink_title="cymbial process">CP</abbrev>
= cymbial process;
<abbrev id="ABBRID0EJACI" xlink_title="cymbium">Cy</abbrev>
= cymbium;
<abbrev id="ABBRID0ENACI" xlink_title="dorsal branch of RTA">dRTA</abbrev>
= dorsal branch of
<abbrev id="ABBRID0ERACI" xlink_title="retrolateral tibial apophysis">RTA</abbrev>
;
<abbrev id="ABBRID0EVACI" xlink_title="embolic tip">EmT</abbrev>
= embolic tip;
<abbrev id="ABBRID0EZACI" xlink_title="sperm duct">SD</abbrev>
= sperm duct;
<abbrev id="ABBRID0E4ACI" xlink_title="sperm duct loop">SDL</abbrev>
= sperm duct loop;
<abbrev id="ABBRID0EBBCI" xlink_title="tegular apophysis">TA</abbrev>
= tegular apophysis;
<abbrev id="ABBRID0EFBCI" xlink_title="tegulum">Te</abbrev>
= tegulum;
<abbrev id="ABBRID0EJBCI" xlink_title="ventro-prolateral tibial apophysis">VPTA</abbrev>
= ventro-prolateral tibial apophysis;
<abbrev id="ABBRID0ENBCI" xlink_title="ventral branch of RTA">vRTA</abbrev>
= ventral branch of
<abbrev id="ABBRID0ERBCI" xlink_title="retrolateral tibial apophysis">RTA</abbrev>
. Scale bar: 0.2 mm (equal for
<emphasis id="DFEC7C9B81AF0B87DE1BEDA991565570" bold="true">A, B</emphasis>
).
</paragraph>
</caption>
<caption id="0A5DA0687083E5337C6E966D422C8E23" ID-DOI="10.3897/zookeys.1240.149456.figure18" ID-arpha="1F2F8683-F76F-5151-AC46-533C7E272B3D" httpUri="https://binary.pensoft.net/fig/1348776" startId="F18">
<paragraph id="BACA373D02EE77F2E7B88142A629E2B1">
<label id="26CF9F00E6A4C60EB3F1A9683ABD6B6A">Figure 18.</label>
</paragraph>
<paragraph id="1E74BD1FB5BAAC2787F169BB0539CE24">
<taxonomicName id="9F8836ACB1107AE71F6A46E1DFF03360" authorityName="Zhang &amp; Zhang &amp; Zhong" authorityYear="2025" class="Arachnida" family="Philodromidae" genus="Sinodromus" higherTaxonomySource="CoL" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="lanyue" status="sp. nov.">
<emphasis id="55CAA14F173B585E8914E55AE6E351E1" italics="true">Sinodromus lanyue</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7D7D8243700931D380D78AEA85C78B1" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male, left palpal bulb
<emphasis id="06736453CDE1A6B829330A2625BEB689" bold="true">A</emphasis>
ventral view
<emphasis id="F50EA034A26307D52975B165DBF12651" bold="true">B</emphasis>
dorsal view
<emphasis id="9DFE000383B7E85DD7525EF148CAB431" bold="true">C</emphasis>
prolateral view
<emphasis id="FD32A6F28A6F7682CF1CC2027F7D3CB5" bold="true">D</emphasis>
retrolateral view. Abbreviations:
<abbrev id="ABBRID0EXCCI" xlink_title="conductor">Con</abbrev>
= conductor;
<abbrev id="ABBRID0E2CCI" xlink_title="embolus">Em</abbrev>
= embolus;
<abbrev id="ABBRID0E6CCI" xlink_title="sperm duct">SD</abbrev>
= sperm duct;
<abbrev id="ABBRID0EDDCI" xlink_title="sperm duct loop">SDL</abbrev>
= sperm duct loop;
<abbrev id="ABBRID0EHDCI" xlink_title="tegular apophysis">TA</abbrev>
= tegular apophysis;
<abbrev id="ABBRID0ELDCI" xlink_title="tegulum">Te</abbrev>
= tegulum. Scale bar: 0.2 mm (equal for
<emphasis id="168B79FE041425CF491090C8443B644B" bold="true">A D</emphasis>
).
</paragraph>
</caption>
<caption id="DEC21F8173F713E5A97CE337AEC67F3B" ID-DOI="10.3897/zookeys.1240.149456.figure19" ID-arpha="0D512FBC-4183-5545-91A2-5F3DB30E2B6D" httpUri="https://binary.pensoft.net/fig/1348777" startId="F19">
<paragraph id="33E52BC5BBD591E67398BE0F5F5F0DE7">
<label id="1F4CE9A329EAC8BA03956F353DF139E4">Figure 19.</label>
</paragraph>
<paragraph id="0C90DF3A2A0415098D03F67B00201B7C">
<taxonomicName id="806832B3460E2F71DCF287DC7CB0E449" authorityName="Zhang &amp; Zhang &amp; Zhong" authorityYear="2025" class="Arachnida" family="Philodromidae" genus="Sinodromus" higherTaxonomySource="CoL" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="lanyue" status="sp. nov.">
<emphasis id="7E52287C92DDBE35E17C94A94A3E0F8C" italics="true">Sinodromus lanyue</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9B04E6C9E43D47B67A7A3E9F5A6D3911" rank="species">sp. nov.</taxonomicNameLabel>
, paratype female (
<emphasis id="6F273AF2F83C1DA2302D9AADEC162F89" bold="true">A E, G</emphasis>
) and holotype male (
<emphasis id="59CA2E30C93D35035BA062507446AF3C" bold="true">F</emphasis>
), epigyne (
<emphasis id="038A0CF379F85E95BCD348D74C3525B1" bold="true">A E</emphasis>
) and frontal view of carapace (
<emphasis id="3BAA2FB40CB5D3E1D3D1E776DA5568EB" bold="true">F, G</emphasis>
)
<emphasis id="F863C1399FAC194092265692F039800E" bold="true">A</emphasis>
macerated, ventral view
<emphasis id="6749F800916D7360ADCCA0E3EB06626B" bold="true">B</emphasis>
macerated, dorsal view
<emphasis id="BCFC48D9A63DC8B1B2A070143D7228CB" bold="true">C</emphasis>
macerated and embedded in arabic gum, ventral view
<emphasis id="7D9B20A5C08A66C1E7F69613F6D24414" bold="true">D</emphasis>
macerated and embedded in arabic gum, dorsal view
<emphasis id="EC5A86FB477D5CEE2AFC27A916CE9862" bold="true">E</emphasis>
intact, ventral view
<emphasis id="1140327088B89045B94432AD370EF573" bold="true">F</emphasis>
male
<emphasis id="479BD62988D922BCDDE066414BE0310F" bold="true">G</emphasis>
female. Abbreviations: A = atrium;
<abbrev id="ABBRID0E6ECI" xlink_title="course of copulatory duct">cCD</abbrev>
= course of copulatory duct (red line);
<abbrev id="ABBRID0EDFCI" xlink_title="copulatory duct">CD</abbrev>
= copulatory duct;
<abbrev id="ABBRID0EHFCI" xlink_title="copulatory opening">CO</abbrev>
= copulatory opening;
<abbrev id="ABBRID0ELFCI" xlink_title="epigynal tooth">ET</abbrev>
= epigynal tooth;
<abbrev id="ABBRID0EPFCI" xlink_title="fertilisation duct">FD</abbrev>
= fertilisation duct;
<abbrev id="ABBRID0ETFCI" xlink_title="glandular mound">GM</abbrev>
= glandular mound;
<abbrev id="ABBRID0EXFCI" xlink_title="median septum">MS</abbrev>
= median septum; R = receptaculum;
<abbrev id="ABBRID0E2FCI" xlink_title="rim of median septum">rMS</abbrev>
= rim of median septum (teal green line);
<abbrev id="ABBRID0E6FCI" xlink_title="rim of sclerotised epigynal fold">rSEF</abbrev>
= rim of sclerotised epigynal fold (blue line);
<abbrev id="ABBRID0EDGCI" xlink_title="sclerotised epigynal fold">SEF</abbrev>
= sclerotised epigynal fold. Scale bars: 0.2 mm (
<emphasis id="4C80BCB0F0A091F3ACD62373B000A8C8" bold="true">A E</emphasis>
); 0.5 mm (equal for
<emphasis id="1AC9858896D7A557F0B92C1717BA50FB" bold="true">F, G</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EQGCI" type="description">
<paragraph id="4837EEEF5DEC3009FF118D0B3F9AD039">
<heading id="C4344FEE1FA1879939EF8C81FBB08387" reason="title">Description.</heading>
</paragraph>
<paragraph id="08F7101BE6F52758D9806DEC09A0FC8D">
<emphasis id="18F5662FC05562DA43F3B882ACF8707D" bold="true">
Male (
<named-content id="2052E1A55049DB962406E7FB8656B523" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/guizhou-normal-university" xlink_title="Museum of Guizhou Normal University">
<collectionCode id="2C93A813C73E973ED1E7997E507A5D2F">MGNU</collectionCode>
</named-content>
- 2025 -
<collectionCode id="6F457B0119C6B66C4D1C0CC64C99DF0A">PHISL</collectionCode>
001).
</emphasis>
Total length 3.64. Carapace 1.46 long, 1.29 wide. Abdomen 2.29 long, 0.93 wide.
<emphasis id="681C749B5C33697799247242B34C400D" bold="true" italics="true">Eye sizes and interdistances</emphasis>
:
<abbrev id="ABBRID0EAHCI" xlink_title="anterior median eyes">AME</abbrev>
0.05,
<abbrev id="ABBRID0EEHCI" xlink_title="anterior lateral eyes">ALE</abbrev>
0.05,
<abbrev id="ABBRID0EIHCI" xlink_title="posterior median eyes">PME</abbrev>
0.03,
<abbrev id="ABBRID0EMHCI" xlink_title="posterior lateral eyes">PLE</abbrev>
0.06,
<abbrev id="ABBRID0EQHCI" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0EUHCI" xlink_title="anterior median eyes">AME</abbrev>
0.16,
<abbrev id="ABBRID0EYHCI" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0E3HCI" xlink_title="anterior lateral eyes">ALE</abbrev>
0.10,
<abbrev id="ABBRID0EAICI" xlink_title="posterior median eyes">PME</abbrev>
<abbrev id="ABBRID0EEICI" xlink_title="posterior median eyes">PME</abbrev>
0.23,
<abbrev id="ABBRID0EIICI" xlink_title="posterior median eyes">PME</abbrev>
<abbrev id="ABBRID0EMICI" xlink_title="posterior lateral eyes">PLE</abbrev>
0.24,
<abbrev id="ABBRID0EQICI" xlink_title="MOQ length">
<collectionCode id="36D23A5B66A2B5D06EDCC635D24FF875">MOQL</collectionCode>
</abbrev>
0.21,
<abbrev id="ABBRID0EUICI" xlink_title="MOQ anterior width">
<collectionCode id="2FAB6C639FA4FDA7E5B4974B31A1882D">MOQA</collectionCode>
</abbrev>
0.26,
<abbrev id="ABBRID0EYICI" xlink_title="MOQ posterior width">
<collectionCode id="B3363B235CF147396F9D17E06275C6B1">MOQP</collectionCode>
</abbrev>
0.29,
<abbrev id="ABBRID0E3ICI" xlink_title="clypeal height">
<collectingCountry id="2432CD72C900784B396A82488DF2BCB4" name="Switzerland">CH</collectingCountry>
</abbrev>
0.15. Sternum 0.87 long, 0.68 wide.
<emphasis id="0EDBA313E3D5CFCBADD52622D675D9E1" bold="true" italics="true">Measurements of legs</emphasis>
: I 6.03 (1.76, 2.23, 1.56, 1.38, 0.66), II 7.61 (2.18, 1.38, 2.03, 1.78, 0.89), III 4.94 (1.6, 1.67, 1.08, 0.59), IV 5.94 (1.91, 1.95, 1.43, 0.65). Leg formula: II-I-IV-III. Cheliceral furrow with one promarginal tooth.
</paragraph>
<paragraph id="7EFC19B8CDE55B9E17A6CC6FB46B1C31">
<emphasis id="B7E94B942789F58E15DC935DCB28A930" bold="true" italics="true">Colouration in ethanol</emphasis>
(Figs
<figureCitation id="2F4A330BC59520FDC818632E3833FDD2" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Sinodromus lanyue sp. nov., paratype female (A E, G) and holotype male (F), epigyne (A E) and frontal view of carapace (F, G) A macerated, ventral view B macerated, dorsal view C macerated and embedded in arabic gum, ventral view D macerated and embedded in arabic gum, dorsal view E intact, ventral view F male G female. Abbreviations: A = atrium; cCD = course of copulatory duct (red line); CD = copulatory duct; CO = copulatory opening; ET = epigynal tooth; FD = fertilisation duct; GM = glandular mound; MS = median septum; R = receptaculum; rMS = rim of median septum (teal green line); rSEF = rim of sclerotised epigynal fold (blue line); SEF = sclerotised epigynal fold. Scale bars: 0.2 mm (A E); 0.5 mm (equal for F, G)." figureDoi="10.3897/zookeys.1240.149456.figure19" httpUri="https://binary.pensoft.net/fig/1348777">19 F</figureCitation>
,
<figureCitation id="A50133748D393DEA741244693E8D0E8B" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Sinodromus lanyue sp. nov., holotype male (A C) and paratype female (D F), habitus A, D dorsal view B, E ventral view C, F lateral view Scale bars: 1 mm (equal for A C, equal for D F)." figureDoi="10.3897/zookeys.1240.149456.figure20" httpUri="https://binary.pensoft.net/fig/1348778">20 A C</figureCitation>
). Carapace basically yellow-brown, nearly pear-shaped, ocular region distinctly narrowed, tegument relatively smooth; with three pairs of indistinct, brown, longitudinal stripes, each one including dense black spots: the central pair starting from
<abbrev id="ABBRID0ESJCI" xlink_title="posterior lateral eyes">PLE</abbrev>
, extending obliquely at front and vertically at rear, forming a funnel shape, or shaped like capital letter Y; the second pair also starting from
<abbrev id="ABBRID0EWJCI" xlink_title="posterior lateral eyes">PLE</abbrev>
and extending almost vertically; the third pair running along the edge of the carapace, slightly curved, resembling a pair of parentheses. Chelicerae coloured slightly paler than carapace, cheliceral base with sparse black spots. Sternum uniformly yellowish-white, laterally with many black dots. Endites and labium coloured as cheliceral base, both with dense scopulae on anterior margins. All legs proximally yellowish-white (coxae, trochanters, and femora), distally brown (patellae, tibiae, metatarsi, and tarsi), with many small black dots on dorsal and lateral surfaces, covered by short spines. Abdomen elongate-oval, dorsum brown, clothed with dense hairs and covered by countless black spots, with two pairs of longitudinal, white lines, reaching entire abdominal length: the central pair anteriorly long and widely separated, posteriorly short and convergent, shaped like a tuning fork; the lateral pair running along the edge of the abdomen, almost ascending parallel; ventral abdomen basically yellowish-white, marked with small dense black dots.
</paragraph>
<caption id="680C749E1882BA626B279EE01757AEB8" ID-DOI="10.3897/zookeys.1240.149456.figure20" ID-arpha="87AA7FCE-89B5-5A1D-82B7-ED1D3638E4F3" httpUri="https://binary.pensoft.net/fig/1348778" startId="F20">
<paragraph id="4FE30C511C431DEC6F0FDF9C880649E5">
<label id="20760004792F364939AFA28379363598">Figure 20.</label>
</paragraph>
<paragraph id="1DB5DC3C57FC61A2C206852B0035230F">
<taxonomicName id="C247EFF6A22DB69F260138CFB5F765A7" authorityName="Zhang &amp; Zhang &amp; Zhong" authorityYear="2025" class="Arachnida" family="Philodromidae" genus="Sinodromus" higherTaxonomySource="CoL" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="lanyue" status="sp. nov.">
<emphasis id="82B9154202674EAFFAE7F97F885CFEFC" italics="true">Sinodromus lanyue</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58EE8E792E36B136FF0392D810593204" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male (
<emphasis id="52C1ADDCA36F7D6D9D7E308F4444AD9A" bold="true">A C</emphasis>
) and paratype female (
<emphasis id="64F6BC7FA78F5317E0313E4D50212EBB" bold="true">D F</emphasis>
), habitus
<emphasis id="DA06834C90E416B34FDFB125FD0B1291" bold="true">A, D</emphasis>
dorsal view
<emphasis id="CA9A2470E72E4EC019524A48B52638A9" bold="true">B, E</emphasis>
ventral view
<emphasis id="B8ED4864C2BABAAA4B37C3F0013A7411" bold="true">C, F</emphasis>
lateral view Scale bars: 1 mm (equal for
<emphasis id="F8063D07812FED843031CEACE881B90A" bold="true">A C</emphasis>
, equal for
<emphasis id="AEA703B91C2B4C17A110A87011CC1270" bold="true">D F</emphasis>
).
</paragraph>
</caption>
<paragraph id="CA930F6E9B6935103B9616CC4C7D4632">
<emphasis id="823EDD885DA8766A3E0F28B3F0D21BD5" bold="true" italics="true">Palp</emphasis>
(Figs
<figureCitation id="C3D65D86489EF90B7F317176A7EB82AE" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Sinodromus lanyue sp. nov., holotype male, left palp A ventral view B dorsal view. Abbreviations: Con = conductor; CP = cymbial process; Cy = cymbium; Em = embolus; SDL = sperm duct loop; TA = tegular apophysis; Te = tegulum; VPTA = ventro-prolateral tibial apophysis; vRTA = ventral branch of RTA. Scale bar: 0.2 mm (equal for A, B)." figureDoi="10.3897/zookeys.1240.149456.figure16" httpUri="https://binary.pensoft.net/fig/1348774">16 A, B</figureCitation>
,
<figureCitation id="B1EBAC3C92085CC6211BF8B857AF3CC1" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Sinodromus lanyue sp. nov., holotype male, left palp A prolateral view B retrolateral view. Abbreviations: Con = conductor; CP = cymbial process; Cy = cymbium; dRTA = dorsal branch of RTA; EmT = embolic tip; SD = sperm duct; SDL = sperm duct loop; TA = tegular apophysis; Te = tegulum; VPTA = ventro-prolateral tibial apophysis; vRTA = ventral branch of RTA. Scale bar: 0.2 mm (equal for A, B)." figureDoi="10.3897/zookeys.1240.149456.figure17" httpUri="https://binary.pensoft.net/fig/1348775">17 A, B</figureCitation>
,
<figureCitation id="E44AC6A5FDF178354199C56030830619" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Sinodromus lanyue sp. nov., holotype male, left palpal bulb A ventral view B dorsal view C prolateral view D retrolateral view. Abbreviations: Con = conductor; Em = embolus; SD = sperm duct; SDL = sperm duct loop; TA = tegular apophysis; Te = tegulum. Scale bar: 0.2 mm (equal for A D)." figureDoi="10.3897/zookeys.1240.149456.figure18" httpUri="https://binary.pensoft.net/fig/1348776">18 A D</figureCitation>
). Tibia relatively long, ~ 2 / 3 of
<abbrev id="ABBRID0ETLCI" xlink_title="cymbium">Cy</abbrev>
length, with two apophyses arising distally from tibia:
<abbrev id="ABBRID0EXLCI" xlink_title="ventro-prolateral tibial apophysis">
<collectionCode id="E7FBB3CFA742DC63A513807059F9A9CF">VPTA</collectionCode>
</abbrev>
relatively short, ~ 1 / 61 / 5 tibia length, subtriangular and nearly erect in ventral view, distinctly curved and dorsally toward posterior part of
<abbrev id="ABBRID0E2LCI" xlink_title="tegulum">Te</abbrev>
in prolateral view;
<abbrev id="ABBRID0E6LCI" xlink_title="retrolateral tibial apophysis">
<collectionCode id="4DA148D1078F80ACAE461CE30A36DC41">RTA</collectionCode>
</abbrev>
bifurcated, with a membranous, thumb-like
<abbrev id="ABBRID0EDMCI" xlink_title="dorsal branch of RTA">dRTA</abbrev>
and a relatively sclerotised, dagger-like
<abbrev id="ABBRID0EHMCI" xlink_title="ventral branch of RTA">vRTA</abbrev>
,
<abbrev id="ABBRID0ELMCI" xlink_title="ventral branch of RTA">vRTA</abbrev>
relatively long, ~ 1 / 3 tibia length, twice longer than
<abbrev id="ABBRID0EPMCI" xlink_title="dorsal branch of RTA">dRTA</abbrev>
.
<abbrev id="ABBRID0ETMCI" xlink_title="cymbium">Cy</abbrev>
~ 1.9 × longer than wide, basoretrolaterally with an indistinct
<abbrev id="ABBRID0EXMCI" xlink_title="cymbial process">CP</abbrev>
.
<abbrev id="ABBRID0E2MCI" xlink_title="tegulum">Te</abbrev>
egg-shaped, ~ 1.55 × longer than wide, proximally slightly swollen, prolatero-apically slightly excavated to accommodate
<abbrev id="ABBRID0E6MCI" xlink_title="embolus">Em</abbrev>
and
<abbrev id="ABBRID0EDNCI" xlink_title="conductor">Con</abbrev>
.
<abbrev id="ABBRID0EHNCI" xlink_title="sperm duct">SD</abbrev>
sinuate, originating at retrolatero-distal portion of
<abbrev id="ABBRID0ELNCI" xlink_title="tegulum">Te</abbrev>
, proximally aligning clockwise along the tegular retrolateral margin, medially forming a S-shaped
<abbrev id="ABBRID0EPNCI" xlink_title="sperm duct loop">SDL</abbrev>
in ventral view, with its distal end hidden behind
<abbrev id="ABBRID0ETNCI" xlink_title="tegulum">Te</abbrev>
and covered by
<abbrev id="ABBRID0EXNCI" xlink_title="conductor">Con</abbrev>
, ultimately entering
<abbrev id="ABBRID0E2NCI" xlink_title="embolic base">EmB</abbrev>
.
<abbrev id="ABBRID0E6NCI" xlink_title="embolus">Em</abbrev>
distinctly simplified and small, ~ 1 / 4
<abbrev id="ABBRID0EDOCI" xlink_title="tegulum">Te</abbrev>
length, slightly curved, spine-like;
<abbrev id="ABBRID0EHOCI" xlink_title="embolic tip">EmT</abbrev>
sharply pointed and directed retrolatero-distally, terminating at ~ 1 oclock position.
<abbrev id="ABBRID0ELOCI" xlink_title="conductor">Con</abbrev>
weakly sclerotised, with moderate size, ~ 1 / 3
<abbrev id="ABBRID0EPOCI" xlink_title="tegulum">Te</abbrev>
length, basally columnar and slightly torqued along its length, apex triangular and terminating at ~ 1 oclock position, covers
<abbrev id="ABBRID0ETOCI" xlink_title="embolus">Em</abbrev>
.
<abbrev id="ABBRID0EXOCI" xlink_title="tegular apophysis">
<collectionCode id="559C58E7D3C2823D1FF7DDBE8FC5AA81" country="0" name="Timescale Adventures Research and Interpretive Center">TA</collectionCode>
</abbrev>
heavily sclerotised, shaped like a cocks head, with a hump-like basal apophysis directed anteriorly and a sharp, beak-like apex pointing retrolaterally.
</paragraph>
<paragraph id="6E41A3A9258879675679132445B144A5">
<emphasis id="A4E349A295893AAB046F738697A381DF" bold="true">
Female (
<named-content id="F2AFBCAB293713E39EFEA448CB7E133C" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/guizhou-normal-university" xlink_title="Museum of Guizhou Normal University">
<collectionCode id="28E8C67632D092022A65E9AA5552806E">MGNU</collectionCode>
</named-content>
- 2025 -
<collectionCode id="6843A55710FA0F5438CCF54B974B8244">PHISL</collectionCode>
002).
</emphasis>
Total length 3.93. Carapace 1.45 long, 1.31 wide. Abdomen 2.66 long, 1.14 wide.
<emphasis id="EC9441E0F4B73EBC149D5BDC0444C222" bold="true" italics="true">Eye sizes and interdistances</emphasis>
:
<abbrev id="ABBRID0EHPCI" xlink_title="anterior median eyes">AME</abbrev>
0.05,
<abbrev id="ABBRID0ELPCI" xlink_title="anterior lateral eyes">ALE</abbrev>
0.05,
<abbrev id="ABBRID0EPPCI" xlink_title="posterior median eyes">PME</abbrev>
0.03,
<abbrev id="ABBRID0ETPCI" xlink_title="posterior lateral eyes">PLE</abbrev>
0.06,
<abbrev id="ABBRID0EXPCI" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0E2PCI" xlink_title="anterior median eyes">AME</abbrev>
0.19,
<abbrev id="ABBRID0E6PCI" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0EDQCI" xlink_title="anterior lateral eyes">ALE</abbrev>
0.11,
<abbrev id="ABBRID0EHQCI" xlink_title="posterior median eyes">PME</abbrev>
<abbrev id="ABBRID0ELQCI" xlink_title="posterior median eyes">PME</abbrev>
0.28,
<abbrev id="ABBRID0EPQCI" xlink_title="posterior median eyes">PME</abbrev>
<abbrev id="ABBRID0ETQCI" xlink_title="posterior lateral eyes">PLE</abbrev>
0.25,
<abbrev id="ABBRID0EXQCI" xlink_title="MOQ length">
<collectionCode id="6F8FEEDC8BFF1F863EB2588D9C2C19FD">MOQL</collectionCode>
</abbrev>
0.21,
<abbrev id="ABBRID0E2QCI" xlink_title="MOQ anterior width">
<collectionCode id="6E4374D9F836104280F81C40CE3FABE0">MOQA</collectionCode>
</abbrev>
0.28,
<abbrev id="ABBRID0E6QCI" xlink_title="MOQ posterior width">
<collectionCode id="60D6931487E9F461A4A511F7A7DBCDBF">MOQP</collectionCode>
</abbrev>
0.34,
<abbrev id="ABBRID0EDRCI" xlink_title="clypeal height">
<collectingCountry id="BDD98EFABA76D605138F73709EF0F213" name="Switzerland">CH</collectingCountry>
</abbrev>
0.17. Sternum 0.88 long, 0.64 wide.
<emphasis id="18500ACBCBA3A7EC7F61D392BE4E965F" bold="true" italics="true">Measurements of legs</emphasis>
: I 4.73 (1.36, 1.81, 1.16, 0.99, 0.57), II 5.37 (1.64, 1.93, 1.31, 1.14, 0.66), III 4.32 (1.4, 1.50, 0.92, 0.5), IV 5.21 (1.73, 1.76, 1.14, 0.58). Leg formula: II-IV-I-III. Cheliceral furrow with one promarginal tooth. Colouration in ethanol as in males, but body slightly paler (Figs
<figureCitation id="91C695664CFA54CD39CD834E7F8709F4" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Sinodromus lanyue sp. nov., paratype female (A E, G) and holotype male (F), epigyne (A E) and frontal view of carapace (F, G) A macerated, ventral view B macerated, dorsal view C macerated and embedded in arabic gum, ventral view D macerated and embedded in arabic gum, dorsal view E intact, ventral view F male G female. Abbreviations: A = atrium; cCD = course of copulatory duct (red line); CD = copulatory duct; CO = copulatory opening; ET = epigynal tooth; FD = fertilisation duct; GM = glandular mound; MS = median septum; R = receptaculum; rMS = rim of median septum (teal green line); rSEF = rim of sclerotised epigynal fold (blue line); SEF = sclerotised epigynal fold. Scale bars: 0.2 mm (A E); 0.5 mm (equal for F, G)." figureDoi="10.3897/zookeys.1240.149456.figure19" httpUri="https://binary.pensoft.net/fig/1348777">19 G</figureCitation>
,
<figureCitation id="121A840D65AEE466A9F607ABE4614B73" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Sinodromus lanyue sp. nov., holotype male (A C) and paratype female (D F), habitus A, D dorsal view B, E ventral view C, F lateral view Scale bars: 1 mm (equal for A C, equal for D F)." figureDoi="10.3897/zookeys.1240.149456.figure20" httpUri="https://binary.pensoft.net/fig/1348778">20 D F</figureCitation>
).
</paragraph>
<paragraph id="A7E640D7CC71CD68B67133A658FCB305">
<emphasis id="0F26E1D912DADCF4511662196181457A" bold="true" italics="true">Epigyne</emphasis>
(Fig.
<figureCitation id="F03FDC45469E4859432B39FDC0D1002E" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Sinodromus lanyue sp. nov., paratype female (A E, G) and holotype male (F), epigyne (A E) and frontal view of carapace (F, G) A macerated, ventral view B macerated, dorsal view C macerated and embedded in arabic gum, ventral view D macerated and embedded in arabic gum, dorsal view E intact, ventral view F male G female. Abbreviations: A = atrium; cCD = course of copulatory duct (red line); CD = copulatory duct; CO = copulatory opening; ET = epigynal tooth; FD = fertilisation duct; GM = glandular mound; MS = median septum; R = receptaculum; rMS = rim of median septum (teal green line); rSEF = rim of sclerotised epigynal fold (blue line); SEF = sclerotised epigynal fold. Scale bars: 0.2 mm (A E); 0.5 mm (equal for F, G)." figureDoi="10.3897/zookeys.1240.149456.figure19" httpUri="https://binary.pensoft.net/fig/1348777">19 A E</figureCitation>
). Epigynal field slightly wider than long; anterior and lateral margins not rebordered, posterior margin delimited;
<abbrev id="ABBRID0E4RCI" xlink_title="copulatory duct">
<collectionCode id="E78CDA74F861588809499F01CF90AB54">CD</collectionCode>
</abbrev>
and
<collectionCode id="68DA09F213EE8490ABF5F295D7458950" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
obscured through epigynal plate in ventral view. A small, located at antero-lateral part of epigynal plate, divided by anterior keel of
<abbrev id="ABBRID0EBSCI" xlink_title="median septum">MS</abbrev>
, represented by two C-shaped depressions; the two depressions separated by ~ five diameters.
<abbrev id="ABBRID0EFSCI" xlink_title="median septum">MS</abbrev>
more or less U-shaped, or vase-shaped, broad; anterior keel slightly narrowed, ~ 2 / 5 epigyne width, with distinct edges and delimited to A; medial stem slightly widened, ~ 1 / 2 epigyne width; posterior base nearly as wide as anterior keel; both middle stem and posterior base with indistinct lateral
<abbrev id="ABBRID0EJSCI" xlink_title="rim of median septum">rMS</abbrev>
alongside with
<abbrev id="ABBRID0ENSCI" xlink_title="rim of sclerotised epigynal fold">rSEF</abbrev>
.
<abbrev id="ABBRID0ERSCI" xlink_title="sclerotised epigynal fold">SEF</abbrev>
shaped like a pair of parentheses; anteriorly distinctly widened, forming axe-shaped
<abbrev id="ABBRID0EVSCI" xlink_title="epigynal tooth">
<collectionCode id="218C174B13C72AD32CC1135AD60C440A" country="0" name="East Texas State University">ET</collectionCode>
</abbrev>
; midsection narrowed, with distinct edges and delimited to A; posteriorly widened,
<abbrev id="ABBRID0EZSCI" xlink_title="rim of sclerotised epigynal fold">rSEF</abbrev>
not distinct and alongside with
<abbrev id="ABBRID0E4SCI" xlink_title="rim of median septum">rMS</abbrev>
.
<abbrev id="ABBRID0EBTCI" xlink_title="copulatory opening">CO</abbrev>
indistinct, located at antero-lateral borders of
<abbrev id="ABBRID0EFTCI" xlink_title="median septum">MS</abbrev>
, leading to
<abbrev id="ABBRID0EJTCI" xlink_title="copulatory duct">
<collectionCode id="A6599A6ACDA00A13FDC254AFEA6655A4">CD</collectionCode>
</abbrev>
which looped to connect with
<collectionCode id="C061A8D5375C926E366F9BEF93F61EB7" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
.
<abbrev id="ABBRID0ENTCI" xlink_title="copulatory duct">
<collectionCode id="C3EC1DC5463045AB06B5BB8E4B76F1AB">CD</collectionCode>
</abbrev>
relatively short, ~ 1 / 4 epigyne length, with a course forming one loop before entering
<collectionCode id="9953EC00AECB7A9B3AFDF4EF58FDA8A2" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
.
<collectionCode id="B952569B65FC1468242206900D8A84F5" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
close together, oval, ~ 1.2 × longer than wide, ~ 1 / 2 epigyne length and 1 / 3 epigyne width; receptacular surface hyaline and smooth, inside pigmented, sclerotised and granular.
<abbrev id="ABBRID0ERTCI" xlink_title="glandular mound">
<collectingRegion id="03A1C8ABB9BB09698E1FECC0331FE6F2" country="Iran" name="Qom">GM</collectingRegion>
</abbrev>
distinctly small, slightly protruding, papilliform, located at the antero-lateral surfaces of
<collectionCode id="CEB8912F7CDD5595074466494EB79A75" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
.
<abbrev id="ABBRID0EVTCI" xlink_title="fertilisation duct">FD</abbrev>
membranous and acicular, moderately long, ~ 2 / 5 of
<collectionCode id="65235AE0CFB0C8E7CD2B1A81A826A226" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
length, originating from the posterior surface of
<collectionCode id="ADFC0A1EDE71B93907168E8DE54FD5E7" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
, directing antero-laterally.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EZTCI" type="distribution">
<paragraph id="90675A0E8321A16BD6864FD2803D366C">
<heading id="29D1253AD4D48308CAF5EBF5BFF2F593" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="D1753C0D27F3F740D043A3ED299B3326">
Known from the
<typeStatus id="0515D7B1BCDB6432B93B490DCE19F8DA">type</typeStatus>
locality in
<collectingRegion id="2711C3AA8D81BB03C0E96E5992D3F451" country="China" name="Hubei">Hubei Province</collectingRegion>
,
<collectingCountry id="2B13F6CA738C40DC9CDA13BD2D2A1386" name="China">China</collectingCountry>
(Fig.
<figureCitation id="F8434F82FEFF85DCEB30CCD44E59E764" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Distribution records of the philodromid species in Guizhou Province (A) and Hubei Province (B)." figureDoi="10.3897/zookeys.1240.149456.figure23" httpUri="https://binary.pensoft.net/fig/1348781">23 B</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,518 @@
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<taxonomicName id="11367C63E0EC23DE58B1FF7E716797DB" ID-CoL="7BYTN" authority="Efimik, 1999" authorityName="Efimik" authorityYear="1999" class="Arachnida" family="Philodromidae" genus="Tibellus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="F6B28D0AD1E5C6D2457954FC4107FBDD" italics="true">Tibellus japonicus</emphasis>
<bibRefCitation id="0552CB1AD4225E636033680C121BA4F9" author="Efimik VE" journalOrPublisher="Arthropoda Selecta" pagination="103-124" refId="B18" refString="Efimik VE (1999) A review of the spider genus Tibellus Simon, 1875 of the East Palearctic (Aranei: Philodromidae). Arthropoda Selecta 8: 103-124." title="A review of the spider genus Tibellus Simon, 1875 of the East Palearctic (Aranei: Philodromidae)." volume="8" year="1999">Efimik, 1999</bibRefCitation>
</taxonomicName>
</paragraph>
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<subSubSection id="81098004E6283D46049B0775C8DAC046" type="description">
<paragraph id="7181DB374D41CB015CCC742FE10F26E1">
<figureCitation id="5C320A64F81C0EA3F79EB057AB913617" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Sample collection locations for this project. Hubei Province 1 Jiugongshan National Nature Reserve 2 Hubei University of Science and Technology 3 Jiuling Mountain 4 Qizimei Mountain National Nature Reserve; Guizhou Province 5 Fanjingshan National Nature Reserve 6 Mayanghe National Nature Reserve 7 Xishui National Nature Reserve 8 Guiyang City, Kaiyang County 9 Guiyang City, Wudang District 10 Guiyang City, Yunyan District 11 Guiyang City, Huaxi District." figureDoi="10.3897/zookeys.1240.149456.figure1" httpUri="https://binary.pensoft.net/fig/1348759">Figs 1</figureCitation>
,
<figureCitation id="B1203BD04CEDB693DE6A6BB122748582" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Living specimens of some Philodromidae species treated in this paper A, B Sinodromus lanyue sp. nov. (♂ ♀) C, D Tibellus japonicus (♂ ♀). Photographs by Q. Lu (Shenzhen)." figureDoi="10.3897/zookeys.1240.149456.figure4" httpUri="https://binary.pensoft.net/fig/1348762">4 C, D</figureCitation>
,
<figureCitation id="1AB68C3F60F471374345B1EB34A53818" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Tibellus japonicus, male, habitus (A C) and left palp (D F) A dorsal view B ventral view C lateral view D prolateral view E ventral view F retrolateral view. Abbreviations: aSDL = ascending part of sperm duct loop; Con = conductor; Cy = cymbium; dSDL = descending part of sperm duct loop; EmB = embolic base; EmT = embolic tip; RTA = retrolateral tibial apophysis; SD = sperm duct; Te = tegulum. Scale bars: 1 mm (equal for A C); 0.2 mm (equal for D F)." figureDoi="10.3897/zookeys.1240.149456.figure21" httpUri="https://binary.pensoft.net/fig/1348779">21</figureCitation>
,
<figureCitation id="ED9C3BBBF8C7DA7F3CFE872380451D8B" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Tibellus japonicus, female, habitus (A C) and epigyne (D, E) A dorsal view B ventral view C lateral view D ventral view E dorsal view. Abbreviations: CO = copulatory opening; FD = fertilisation duct; GM = glandular mound; MS = median septum; R = receptaculum. Scale bars: 2 mm (equal for A C); 0.2 mm (equal for D, E)." figureDoi="10.3897/zookeys.1240.149456.figure22" httpUri="https://binary.pensoft.net/fig/1348780">22</figureCitation>
,
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</paragraph>
</subSubSection>
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<taxonomicName id="40014512289A530C5C9719890A169A7B" ID-CoL="7BYGD" baseAuthorityName="L. Koch" baseAuthorityYear="1876" class="Arachnida" family="Philodromidae" genus="Tibellus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tenellus">
<emphasis id="31E149E364C1CE7055EB9458AF6A960F" italics="true">Tibellus tenellus</emphasis>
</taxonomicName>
<comment id="3736ABFBEBBB6ED6A4072CDCEBBF9780">
:
<treatmentCitation id="F7F0D23179943F16993ABC6F230692D4" author="Bösenberg W &amp; Strand E" page="271" year="1906">
<bibRefCitation id="20CFB5C8F2F549F8D1BF953386DBB1FC" author="Bösenberg W &amp; Strand E" firstAuthor="Bosenberg" journalOrPublisher="Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft" page="271" pagination="93-422" refId="B5" refString="Bösenberg W, Strand E (1906) Japanische Spinnen. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 30: 93422. https://www.biodiversitylibrary.org/part/49839" title="Japanische Spinnen." url="https://www.biodiversitylibrary.org/part/49839" volume="30" year="1906">Bösenberg and Strand 1906: 271</bibRefCitation>
</treatmentCitation>
, pl. 8, fig. 112, pl. 10, fig. 156 (
<specimenCount id="9A46AB10BC88A39879531AA9F9B26A2A" count="1" type="male"></specimenCount>
<specimenCount id="CFEA4DA15D5E59E3201D9F19A3650EE8" count="1" type="female"></specimenCount>
; misidentified per
<treatmentCitation id="B8F2BDE76EE10AD8E072EC3ED1083E59" author="Efimik VE" page="112" year="1999">
<bibRefCitation id="8A19F043EB0F701AD7A7784E0B6BD78B" author="Efimik VE" firstAuthor="Efimik" journalOrPublisher="Arthropoda Selecta" page="112" pagination="103-124" refId="B18" refString="Efimik VE (1999) A review of the spider genus Tibellus Simon, 1875 of the East Palearctic (Aranei: Philodromidae). Arthropoda Selecta 8: 103124." title="A review of the spider genus Tibellus Simon, 1875 of the East Palearctic (Aranei: Philodromidae)." volume="8" year="1999">Efimik 1999: 112</bibRefCitation>
</treatmentCitation>
);
<treatmentCitation id="D2FBF7E27E2ECBB385B31C198E539B89" author="Chikuni Y" page="133" year="1989">
<bibRefCitation id="F7B81382235B73CF3DD1712AC4DC373C" author="Chikuni Y" firstAuthor="Chikuni" journalOrPublisher="Kaisei-sha Publishing Co., Tokyo" page="133" refId="B9" refString="Chikuni Y (1989) Pictorial encyclopedia of spiders in Japan. Kaisei-sha Publishing Co., Tokyo, 310 pp." title="Pictorial encyclopedia of spiders in Japan." year="1989">Chikuni 1989: 133</bibRefCitation>
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, fig. 2 (
<specimenCount id="2817E173FF78C3D67EF0927D40ECC776" count="1" type="male"></specimenCount>
<specimenCount id="B5A1333972D0363B1EBA8A5A8E00B653" count="1" type="female"></specimenCount>
; misidentified per
<treatmentCitation id="2967CDD9FC6B9257F374393B89A45A91" author="Efimik VE" page="112" year="1999">
<bibRefCitation id="230C702A826F1ABA88C645EBCD6F871F" author="Efimik VE" firstAuthor="Efimik" journalOrPublisher="Arthropoda Selecta" page="112" pagination="103-124" refId="B18" refString="Efimik VE (1999) A review of the spider genus Tibellus Simon, 1875 of the East Palearctic (Aranei: Philodromidae). Arthropoda Selecta 8: 103124." title="A review of the spider genus Tibellus Simon, 1875 of the East Palearctic (Aranei: Philodromidae)." volume="8" year="1999">Efimik 1999: 112</bibRefCitation>
</treatmentCitation>
).
</comment>
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<treatmentCitation id="9E43165F22C2564A06AAF4CFC4C2F721" author="Efimik VE" page="112" year="1999">
<taxonomicName id="AECE86B977D199F002B8692A73CA7676" ID-CoL="7BYTN" authority="Efimik, 1999: 112" authorityName="Efimik" authorityPageNumber="112" authorityYear="1999" class="Arachnida" family="Philodromidae" genus="Tibellus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="3ECD89FA1E5BFA5247AA8A4CEF743019" italics="true">Tibellus japonicus</emphasis>
<comment id="D64F323756F626109C88AC450DE1147C">
<bibRefCitation id="EB3B98990E3274D0E41ABB55C55C91C7" author="Efimik VE" journalOrPublisher="Arthropoda Selecta" pagination="103-124" refId="B18" refString="Efimik VE (1999) A review of the spider genus Tibellus Simon, 1875 of the East Palearctic (Aranei: Philodromidae). Arthropoda Selecta 8: 103-124." title="A review of the spider genus Tibellus Simon, 1875 of the East Palearctic (Aranei: Philodromidae)." volume="8" year="1999">Efimik, 1999: 112</bibRefCitation>
, figs 35, 46, 52, 65 (
<specimenCount id="7D4DAADED986BBBB8DE785E5FF353F6A" count="1" type="female"></specimenCount>
);
<treatmentCitation id="D5F5F8108E7C43439520435E79EA04EE" author="Chen HM &amp; Zhang JX &amp; Song DX" page="91" year="2003">
<bibRefCitation id="92433CE2E05A194C3CE2E2242AD9B7C5" DOI="10.3969/j.issn.1005-9628.2003.02.006" author="Chen HM &amp; Zhang JX &amp; Song DX" etAl="et al." firstAuthor="Chen" journalOrPublisher="Acta Arachnologica Sinica" page="91" pagination="91-93" refId="B8" refString="Chen HM, Zhang JX, Song DX (2003) A newly recorded species of the family Philodromidae from China (Arachnida: Araneae). Acta Arachnologica Sinica 12: 9193. https://doi.org/10.3969/j.issn.1005-9628.2003.02.006" title="A newly recorded species of the family Philodromidae from China (Arachnida: Araneae)." volume="12" year="2003">Chen et al. 2003: 91</bibRefCitation>
</treatmentCitation>
, figs 15 (
<specimenCount id="FD5FF6142165BA0E8FEC8533CC0E9C3F" count="1" type="male"></specimenCount>
<specimenCount id="8B9BCE3662155BFACBCDBB6CAE00E185" count="1" type="female"></specimenCount>
);
<treatmentCitation id="F16677F94D7E09978349C57D595EE5F1" author="Ono H &amp; Ban M" page="478" year="2009">
<bibRefCitation id="7617FC3BA62B056146ABD54AE2EF1E79" author="Ono H &amp; Ban M" editor="Ono H" firstAuthor="Ono" journalOrPublisher="Tokai University Press, Kanagawa, 249-250" page="478" pagination="476-481" refId="B42" refString="Ono H, Ban M (2009) Oxyopidae, Philodromidae. In: Ono H (Ed.) The spiders of Japan with keys to the families and genera and illustrations of the species. Tokai University Press, Kanagawa, 249250, 476481." title="Oxyopidae, Philodromidae." volumeTitle="The spiders of Japan with keys to the families and genera and illustrations of the species." year="2009">Ono and Ban 2009: 478</bibRefCitation>
</treatmentCitation>
, figs 2427 (
<specimenCount id="9C5405FEC11330B0688FF1122AB3BB7E" count="1" type="male"></specimenCount>
<specimenCount id="CEA208785F9F721D8D6CBAED66996F62" count="1" type="female"></specimenCount>
);
<treatmentCitation id="28A387302C5E84439CCAC940169D02DE" author="Yin CM &amp; Peng XJ &amp; Yan HM &amp; Bao YH &amp; Xu X &amp; Tang G &amp; Zhou QS &amp; Liu P" page="1252" year="2012">
<bibRefCitation id="43EBBBA8474BCA637F959E323D158B84" author="Yin CM &amp; Peng XJ &amp; Yan HM &amp; Bao YH &amp; Xu X &amp; Tang G &amp; Zhou QS &amp; Liu P" etAl="et al." firstAuthor="Yin" journalOrPublisher="Hunan Science and Technology Press, Changsha" page="1252" refId="B62" refString="Yin CM, Peng XJ, Yan HM, Bao YH, Xu X, Tang G, Zhou QS, Liu P (2012) Fauna Hunan: Araneae in Hunan, China. Hunan Science and Technology Press, Changsha, 1590 pp." title="Fauna Hunan: Araneae in Hunan, China." year="2012">Yin et al. 2012: 1252</bibRefCitation>
</treatmentCitation>
, fig. 673 a d (
<specimenCount id="0BD8BA7F28B1DB90FCBAF0BBA11C4EAD" count="1" type="male"></specimenCount>
<specimenCount id="582F8F36AC8DEBA2C0AE6ABE05FABC4A" count="1" type="female"></specimenCount>
);
<treatmentCitation id="7AF6AB5344C62ABD836E70CA9E7DC4CB" author="Jang CM &amp; Lee SY &amp; Yoo JS &amp; Bae YS &amp; Kim ST" page="278" year="2023">
<bibRefCitation id="1BAF28E04CE1AA9832346B8129C4C944" DOI="10.5635/ASED.2023.39.4.044" author="Jang CM &amp; Lee SY &amp; Yoo JS &amp; Bae YS &amp; Kim ST" etAl="et al." firstAuthor="Jang" issue="4" journalOrPublisher="Animal Systematics, Evolution and Diversity" page="278" pagination="272-283" refId="B23" refString="Jang CM, Lee SY, Yoo JS, Bae YS, Kim ST (2023) Six new records of running crab spiders of the genus Tibellus with four new species (Araneae: Philodromidae) from Korea. Animal Systematics, Evolution and Diversity 39 (4): 272283. https://doi.org/10.5635/ASED.2023.39.4.044" title="Six new records of running crab spiders of the genus Tibellus with four new species (Araneae: Philodromidae) from Korea." volume="39" year="2023">Jang et al. 2023: 278</bibRefCitation>
</treatmentCitation>
, fig. 4 A H (
<specimenCount id="A97FEC65FCD116983EF436E8A664A8C4" count="1" type="male"></specimenCount>
).
<superScript id="026B324BCEBD92FFB9C6621CB38DC1BA">
<xref id="2ABC3028B8E76A3B61E0B15797A3AE52" ref-type="fn" rid="en4">4</xref>
</superScript>
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EU1CI" type="material">
<paragraph id="6D839F4216611F1FCF7667423457F707">
<heading id="03818AFD709217DA211B774005B2D483" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="D0345AD08437F56B314F49A65E71A1DF">
<materialsCitation id="E5FD0FAF0B4E3E86A12E39DF6A74B6E0" collectingDate="2022-06-04" collectionCode="YHGY" collectorName="H. Yu" country="China" county="GenBank" elevation="1310" latitude="26.63" location="Guiyang City" longLatPrecision="744" longitude="106.7" municipality="Table" specimenCode="YHGY 301, YHGY 303" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Guizhou">
<emphasis id="6A9A9E268EA3D05ED93D4A5255334126" bold="true">
<collectingCountry id="86C758AA7B506688AFE82DAA3889E856" name="China">China</collectingCountry>
</emphasis>
:
<collectingRegion id="3CAB1D64766B5FA06D313C348F632688" country="China" name="Guizhou">Guizhou Province</collectingRegion>
: •
<specimenCount id="EB919B1E6C97E8AF929FD7E810715EA1" count="1" type="male">1 ♂</specimenCount>
,
<specimenCount id="12D826A244369D2F92F38E90A2141E19" count="1" type="female">1 ♀</specimenCount>
(
<specimenCode id="9ED0C571825A58E194DA287C3AECA6A3" collectionCode="YHGY">YHGY 301</specimenCode>
and
<specimenCode id="1A859A23A8CC1986B47E03F081FBE9D6" collectionCode="YHGY">YHGY 303</specimenCode>
used for sequencing,
<collectingCounty id="A6496CA3DF5D1E81783FD85D5C6DB5D1">GenBank</collectingCounty>
accession numbers in
<collectingMunicipality id="463FD8E3A2E0842F6FC64696B3B2BCAA">Table</collectingMunicipality>
<tableCitation id="B42CC9A77F1B296DB8505A80AC12C94F" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Voucher specimen information." httpUri="http://table.plazi.org/id/12F54BE0BDE6512C62E0465BE733F517" tableUuid="12F54BE0BDE6512C62E0465BE733F517">1</tableCitation>
),
<location id="786F4A5D8199E399E307FB3D2ABF7DB0" LSID="urn:lsid:plazi:treatment:FFF24DD002B45D6DBEC999D53444FF12:786F4A5D8199E399E307FB3D2ABF7DB0" country="China" county="GenBank" latitude="26.63" longLatPrecision="744" longitude="106.7" municipality="Table" name="Guiyang City" stateProvince="Guizhou">Guiyang City</location>
,
<location id="9422E4458AAB3D788BF3D5C0754307C0" LSID="urn:lsid:plazi:treatment:FFF24DD002B45D6DBEC999D53444FF12:9422E4458AAB3D788BF3D5C0754307C0" country="China" county="GenBank" latitude="26.63" longLatPrecision="744" longitude="106.7" municipality="Table" name="Yunyan District" stateProvince="Guizhou">Yunyan District</location>
,
<location id="83A6FD99D6466867D576F01A0585B21B" LSID="urn:lsid:plazi:treatment:FFF24DD002B45D6DBEC999D53444FF12:83A6FD99D6466867D576F01A0585B21B" country="China" county="GenBank" latitude="26.63" longLatPrecision="744" longitude="106.7" municipality="Table" name="Luchongguan Forest Park" stateProvince="Guizhou">Luchongguan Forest Park</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ED2CI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[106.700000,26.630000]}">
<geoCoordinate id="DEDBE380E2978294C538EF6A67F07741" degrees="26.63" direction="north" orientation="latitude" precision="555" value="26.63">26.63 ° N</geoCoordinate>
,
<geoCoordinate id="BA9E83CF92917B45F586F365C9543FCA" degrees="106.70" direction="east" orientation="longitude" precision="555" value="106.7">106.70 ° E</geoCoordinate>
</named-content>
;
<elevation id="2E818076775B7A7E5994D8334A032B5B" metricMagnitude="3" metricUnit="m" metricValue="1.31" unit="m" value="1310.0">
<quantity id="B6FDABCF0A7D21944D96F2D7A0CD4C10" metricMagnitude="3" metricUnit="m" metricValue="1.31" unit="m" value="1310.0">1310 m</quantity>
a. s. l.
</elevation>
;
<date id="D67E1AB1567F5579BD20085BF413EEB7" value="2022-06-04">
<collectingDate id="E59F44782401E7A2EF56371B250C648E" value="2022-06-04">4 June 2022</collectingDate>
</date>
;
<collectorName id="0F81F407B9728A8B0B05B7944FF54445">H. Yu</collectorName>
et al. leg
</materialsCitation>
. •
<materialsCitation id="EC43BB7C0F748AB03F24AD11720E2651" collectingDate="2022-06-02" collectionCode="YHGY" collectorName="H. Yu" country="China" county="GenBank" elevation="1172" latitude="26.74" location="Guiyang City" longLatPrecision="744" longitude="106.88" municipality="Table" specimenCode="YHGY 014" specimenCount="1" specimenCount-female="1">
<specimenCount id="A3A3757013DCA52AE8F865163D89349B" count="1" type="female">1 ♀</specimenCount>
(
<specimenCode id="7B83C74E42D7D3990D32E51CD8A6DB2C" collectionCode="YHGY">YHGY 014</specimenCode>
used for sequencing,
<collectingCounty id="7A9D78BF916465A1273DE478E5DB61E8">GenBank</collectingCounty>
accession numbers in
<collectingMunicipality id="BBB295C5C25D39EABC65C115CBD9DA74">Table</collectingMunicipality>
<tableCitation id="25DFA4EFE3EA2A733EFB814AABF2B7E4" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Voucher specimen information." httpUri="http://table.plazi.org/id/12F54BE0BDE6512C62E0465BE733F517" tableUuid="12F54BE0BDE6512C62E0465BE733F517">1</tableCitation>
),
<location id="A4C17417534AADEFEE9DF09FC7377D3F" LSID="urn:lsid:plazi:treatment:FFF24DD002B45D6DBEC999D53444FF12:A4C17417534AADEFEE9DF09FC7377D3F" country="China" county="GenBank" latitude="26.74" longLatPrecision="744" longitude="106.88" municipality="Table" name="Guiyang City">Guiyang City</location>
,
<location id="F8346B4E02BB15D2075762DA2748B592" LSID="urn:lsid:plazi:treatment:FFF24DD002B45D6DBEC999D53444FF12:F8346B4E02BB15D2075762DA2748B592" country="China" county="GenBank" latitude="26.74" longLatPrecision="744" longitude="106.88" municipality="Table" name="Wudang District">Wudang District</location>
,
<location id="B852F58770B9C16B99734D443CE8EF86" LSID="urn:lsid:plazi:treatment:FFF24DD002B45D6DBEC999D53444FF12:B852F58770B9C16B99734D443CE8EF86" country="China" county="GenBank" latitude="26.74" longLatPrecision="744" longitude="106.88" municipality="Table" name="Panlongshan Forest Park">Panlongshan Forest Park</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EP2CI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[106.880000,26.740000]}">
<geoCoordinate id="68474DA14DADD078E0FEECC123F4CDD9" degrees="26.74" direction="north" orientation="latitude" precision="555" value="26.74">26.74 ° N</geoCoordinate>
,
<geoCoordinate id="BEB38E63199CEBB0B072982349BA090C" degrees="106.88" direction="east" orientation="longitude" precision="555" value="106.88">106.88 ° E</geoCoordinate>
</named-content>
;
<elevation id="AAA3AF8255AE1DCF7933CB43B79E6C26" metricMagnitude="3" metricUnit="m" metricValue="1.172" unit="m" value="1172.0">
<quantity id="36102AA00565740797DA188D7C60C8D2" metricMagnitude="3" metricUnit="m" metricValue="1.172" unit="m" value="1172.0">1172 m</quantity>
a. s. l.
</elevation>
;
<date id="32D3BE3F805A31CC76BC2F4D5A69C151" value="2022-06-02">
<collectingDate id="39B6E0FCABC8D01BE0837E42A1DC785C" value="2022-06-02">2 June 2022</collectingDate>
</date>
;
<collectorName id="63A06446D2B16427B863910668B80B42">H. Yu</collectorName>
et al. leg
</materialsCitation>
. •
<materialsCitation id="4D12492E1AFB6EB0C462751F64E20D82" collectingDate="2021-07-21" collectorName="H. Yu" country="China" county="Miao Autonomous County" elevation="1462" latitude="28.0" location="Fanjingshan National Nature Reserve" longLatPrecision="740" longitude="108.74" municipality="Tongren City" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="47A15E57150729E50F748D6BED939D21" count="1" type="male">1 ♂</specimenCount>
,
<specimenCount id="E500D3D9AE1F460D5EB4689E42EE85CE" count="1" type="female">1 ♀</specimenCount>
,
<collectingMunicipality id="54E503B866145C390B391994D4DBD55F">Tongren City</collectingMunicipality>
,
<location id="27FCDF85583FA4FFE6C76810F9F2E98A" LSID="urn:lsid:plazi:treatment:FFF24DD002B45D6DBEC999D53444FF12:27FCDF85583FA4FFE6C76810F9F2E98A" country="China" county="Miao Autonomous County" latitude="28.0" longLatPrecision="740" longitude="108.74" municipality="Tongren City" name="Fanjingshan National Nature Reserve">Fanjingshan National Nature Reserve</location>
,
<location id="0F774C8D2E352091BAB3C50A618EE0F2" LSID="urn:lsid:plazi:treatment:FFF24DD002B45D6DBEC999D53444FF12:0F774C8D2E352091BAB3C50A618EE0F2" country="China" county="Miao Autonomous County" latitude="28.0" longLatPrecision="740" longitude="108.74" municipality="Tongren City" name="Yinjiang Tujia">Yinjiang Tujia</location>
and
<collectingCounty id="05E254FDE062AF5FC1DC20B8F51CFCDD">Miao Autonomous County</collectingCounty>
,
<location id="4A879EBF93A2EBC2D621C35484FCCEC5" LSID="urn:lsid:plazi:treatment:FFF24DD002B45D6DBEC999D53444FF12:4A879EBF93A2EBC2D621C35484FCCEC5" country="China" county="Miao Autonomous County" latitude="28.0" longLatPrecision="740" longitude="108.74" municipality="Tongren City" name="Tianqing Temple">Tianqing Temple</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EX2CI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[108.740000,28.000000]}">
<geoCoordinate id="E8A5DAC161CD13F34536B41A13233314" degrees="28.00" direction="north" orientation="latitude" precision="555" value="28.0">28.00 ° N</geoCoordinate>
,
<geoCoordinate id="CC8F97CCD795052952E7DD7A47975C99" degrees="108.74" direction="east" orientation="longitude" precision="555" value="108.74">108.74 ° E</geoCoordinate>
</named-content>
;
<elevation id="AC47E952AB1285301E023383EC6DA7E3" metricMagnitude="3" metricUnit="m" metricValue="1.462" unit="m" value="1462.0">
<quantity id="1E996F424BA77CBB45D44316F9DDD9CF" metricMagnitude="3" metricUnit="m" metricValue="1.462" unit="m" value="1462.0">1462 m</quantity>
a. s. l.
</elevation>
;
<date id="3091A2C57AFFEFFFE4E6B20357676A71" value="2021-07-21">
<collectingDate id="826473523C55152A73A34D3DF5836FEA" value="2021-07-21">21 July 2021</collectingDate>
</date>
;
<collectorName id="0566485C4464474FFF04994BF848CF03">H. Yu</collectorName>
et al. leg
</materialsCitation>
. •
<materialsCitation id="89198B8EBF0D5F9D82012D495CE65AC3" collectingDate="2022-05-28" collectorName="H. Yu" country="China" county="Xishui County" elevation="981" latitude="28.4" location="Xishui National Nature Reserve" longLatPrecision="739" longitude="106.18" municipality="Zunyi City" specimenCount="1" specimenCount-female="1" stateProvince="Hubei">
<specimenCount id="D33C6E99103BE32C357E080E5C75C190" count="1" type="female">1 ♀</specimenCount>
,
<collectingMunicipality id="20CB593C8749CE5C40D210553D870E5D">Zunyi City</collectingMunicipality>
,
<location id="667CD08CCE8FB770CCF42EEE4C60D114" LSID="urn:lsid:plazi:treatment:FFF24DD002B45D6DBEC999D53444FF12:667CD08CCE8FB770CCF42EEE4C60D114" country="China" county="Xishui County" latitude="28.4" longLatPrecision="739" longitude="106.18" municipality="Zunyi City" name="Xishui National Nature Reserve" stateProvince="Hubei">Xishui National Nature Reserve</location>
,
<collectingCounty id="CDCBCB5F2D66A9A142CBE188168331C8">Xishui County</collectingCounty>
,
<location id="B16D9227B17C155CA27A34B9AEFAAE1D" LSID="urn:lsid:plazi:treatment:FFF24DD002B45D6DBEC999D53444FF12:B16D9227B17C155CA27A34B9AEFAAE1D" country="China" county="Xishui County" latitude="28.4" longLatPrecision="739" longitude="106.18" municipality="Zunyi City" name="Donghuang Town" stateProvince="Hubei">Donghuang Town</location>
,
<location id="FC1BFEC7415813FBB1FC7080971ABCE7" LSID="urn:lsid:plazi:treatment:FFF24DD002B45D6DBEC999D53444FF12:FC1BFEC7415813FBB1FC7080971ABCE7" country="China" county="Xishui County" latitude="28.4" longLatPrecision="739" longitude="106.18" municipality="Zunyi City" name="Changqiangou Village" stateProvince="Hubei">Changqiangou Village</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E62CI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[106.180000,28.400000]}">
<geoCoordinate id="822B81E0A3D003F0F8E159DA0A9ED307" degrees="28.40" direction="north" orientation="latitude" precision="555" value="28.4">28.40 ° N</geoCoordinate>
,
<geoCoordinate id="4B614E3A6CFDB6F2E14C02C2165E4237" degrees="106.18" direction="east" orientation="longitude" precision="555" value="106.18">106.18 ° E</geoCoordinate>
</named-content>
;
<elevation id="FA9268489BD4D1B860FC69810ED77E68" metricMagnitude="2" metricUnit="m" metricValue="9.809999999999999" unit="m" value="981.0">
<quantity id="C6A56525349054E524998965FEC2F629" metricMagnitude="2" metricUnit="m" metricValue="9.809999999999999" unit="m" value="981.0">981 m</quantity>
a. s. l.
</elevation>
;
<date id="417B8F67CE357564E60F2B4E07480DD2" value="2022-05-28">
<collectingDate id="2BF2C5ED18FCD6A64B9E496DDDB27222" value="2022-05-28">28 May 2022</collectingDate>
</date>
;
<collectorName id="99647BFD52CF00E987EB5437A9CEC7B2">H. Yu</collectorName>
et al. leg.;
<collectingRegion id="864BD1FDCDC032374E13B148859A72F5" country="China" name="Hubei">Hubei Province</collectingRegion>
</materialsCitation>
: •
<materialsCitation id="2A15CDBFB10CBAE94E351945978CB376" collectingDate="2020-07-03" collectorName="Q. Lu" country="China" county="Xianning City" elevation="480" latitude="29.39" location="Yunzhonghu Scenic Area" longLatPrecision="736" longitude="114.65" municipality="Jiugongshan National Nature Reserve" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="A062C9CBC71B38CC54ADACDAC9254919" count="1" type="male">1 ♂</specimenCount>
,
<specimenCount id="5F7871984B555DF067D6F512A917119E" count="1" type="female">1 ♀</specimenCount>
,
<collectingCounty id="8F907D998B1CAEE29E46BD0F9C86FD4C">Xianning City</collectingCounty>
,
<collectingMunicipality id="92FFC34646BFD15575F25BF083305E2D">Jiugongshan National Nature Reserve</collectingMunicipality>
,
<location id="306EB4E522C00A3DBADDD940C9F2F1D2" LSID="urn:lsid:plazi:treatment:FFF24DD002B45D6DBEC999D53444FF12:306EB4E522C00A3DBADDD940C9F2F1D2" country="China" county="Xianning City" latitude="29.39" longLatPrecision="736" longitude="114.65" municipality="Jiugongshan National Nature Reserve" name="Yunzhonghu Scenic Area">Yunzhonghu Scenic Area</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EH3CI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[114.650000,29.390000]}">
<geoCoordinate id="1C6BDAF732F8A339ED4B0F0F7DA29BB5" degrees="29.39" direction="north" orientation="latitude" precision="555" value="29.39">29.39 ° N</geoCoordinate>
,
<geoCoordinate id="15E35F383B1A331184F67D60D6EAC0A7" degrees="114.65" direction="east" orientation="longitude" precision="555" value="114.65">114.65 ° E</geoCoordinate>
</named-content>
;
<elevation id="909C924FA120C881DFE39C30F4C6ACDD" metricMagnitude="2" metricUnit="m" metricValue="4.8" unit="m" value="480.0">
<quantity id="2868A68F607A97029A8F28F3B843E9B0" metricMagnitude="2" metricUnit="m" metricValue="4.8" unit="m" value="480.0">480 m</quantity>
a. s. l.
</elevation>
;
<date id="4A33440FB962473BC55F6858F9F53696" value="2020-07-03">
<collectingDate id="01BBDA3E9F5510F03C826A2F1F1B0518" value="2020-07-03">3 July 2020</collectingDate>
</date>
;
<collectorName id="F7AE13CBDBB7688765FA83E3C6687280">Q. Lu</collectorName>
et al. leg
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EM3CI" type="diagnosis">
<paragraph id="C2DB306F7CC75BCD07C8D762D00665AA">
<heading id="F4DAD56B39E45EDAF01610305A861978" reason="title">Diagnosis and description.</heading>
</paragraph>
<paragraph id="BCE830F6378484F7E24C6FA21A031EEE">
See
<bibRefCitation id="3E4B5DF9F4CD381400A29B5DB4440107" author="Efimik VE" firstAuthor="Efimik" journalOrPublisher="Arthropoda Selecta" pagination="103-124" refId="B18" refString="Efimik VE (1999) A review of the spider genus Tibellus Simon, 1875 of the East Palearctic (Aranei: Philodromidae). Arthropoda Selecta 8: 103124." title="A review of the spider genus Tibellus Simon, 1875 of the East Palearctic (Aranei: Philodromidae)." volume="8" year="1999">Efimik (1999)</bibRefCitation>
,
<bibRefCitation id="7FF9064D51A5E5A21F583E293C996E00" DOI="10.3969/j.issn.1005-9628.2003.02.006" author="Chen HM &amp; Zhang JX &amp; Song DX" etAl="et al." firstAuthor="Chen" journalOrPublisher="Acta Arachnologica Sinica" pagination="91-93" refId="B8" refString="Chen HM, Zhang JX, Song DX (2003) A newly recorded species of the family Philodromidae from China (Arachnida: Araneae). Acta Arachnologica Sinica 12: 9193. https://doi.org/10.3969/j.issn.1005-9628.2003.02.006" title="A newly recorded species of the family Philodromidae from China (Arachnida: Araneae)." volume="12" year="2003">Chen et al. (2003)</bibRefCitation>
,
<bibRefCitation id="EB8A58963B9AA62503F3A8EDCD987EE9" author="Yin CM &amp; Peng XJ &amp; Yan HM &amp; Bao YH &amp; Xu X &amp; Tang G &amp; Zhou QS &amp; Liu P" etAl="et al." firstAuthor="Yin" journalOrPublisher="Hunan Science and Technology Press, Changsha" refId="B62" refString="Yin CM, Peng XJ, Yan HM, Bao YH, Xu X, Tang G, Zhou QS, Liu P (2012) Fauna Hunan: Araneae in Hunan, China. Hunan Science and Technology Press, Changsha, 1590 pp." title="Fauna Hunan: Araneae in Hunan, China." year="2012">Yin et al. (2012)</bibRefCitation>
, and
<bibRefCitation id="FC809188FC008B9A7FD64497D5DEFC7D" DOI="10.5635/ASED.2023.39.4.044" author="Jang CM &amp; Lee SY &amp; Yoo JS &amp; Bae YS &amp; Kim ST" etAl="et al." firstAuthor="Jang" issue="4" journalOrPublisher="Animal Systematics, Evolution and Diversity" pagination="272-283" refId="B23" refString="Jang CM, Lee SY, Yoo JS, Bae YS, Kim ST (2023) Six new records of running crab spiders of the genus Tibellus with four new species (Araneae: Philodromidae) from Korea. Animal Systematics, Evolution and Diversity 39 (4): 272283. https://doi.org/10.5635/ASED.2023.39.4.044" title="Six new records of running crab spiders of the genus Tibellus with four new species (Araneae: Philodromidae) from Korea." volume="39" year="2023">Jang et al. (2023)</bibRefCitation>
. Habitus as in Figs
<figureCitation id="BC0F65FE6159DD4B2F45B6577B808F89" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Tibellus japonicus, male, habitus (A C) and left palp (D F) A dorsal view B ventral view C lateral view D prolateral view E ventral view F retrolateral view. Abbreviations: aSDL = ascending part of sperm duct loop; Con = conductor; Cy = cymbium; dSDL = descending part of sperm duct loop; EmB = embolic base; EmT = embolic tip; RTA = retrolateral tibial apophysis; SD = sperm duct; Te = tegulum. Scale bars: 1 mm (equal for A C); 0.2 mm (equal for D F)." figureDoi="10.3897/zookeys.1240.149456.figure21" httpUri="https://binary.pensoft.net/fig/1348779">21 A C</figureCitation>
,
<figureCitation id="96F7E6BCF31EE6DFDD54D1FE8B946F05" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Tibellus japonicus, female, habitus (A C) and epigyne (D, E) A dorsal view B ventral view C lateral view D ventral view E dorsal view. Abbreviations: CO = copulatory opening; FD = fertilisation duct; GM = glandular mound; MS = median septum; R = receptaculum. Scale bars: 2 mm (equal for A C); 0.2 mm (equal for D, E)." figureDoi="10.3897/zookeys.1240.149456.figure22" httpUri="https://binary.pensoft.net/fig/1348780">22 A C</figureCitation>
. Male palp as in Fig.
<figureCitation id="7938DB1AB7A52A1C8905F5BA3F6F766A" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Tibellus japonicus, male, habitus (A C) and left palp (D F) A dorsal view B ventral view C lateral view D prolateral view E ventral view F retrolateral view. Abbreviations: aSDL = ascending part of sperm duct loop; Con = conductor; Cy = cymbium; dSDL = descending part of sperm duct loop; EmB = embolic base; EmT = embolic tip; RTA = retrolateral tibial apophysis; SD = sperm duct; Te = tegulum. Scale bars: 1 mm (equal for A C); 0.2 mm (equal for D F)." figureDoi="10.3897/zookeys.1240.149456.figure21" httpUri="https://binary.pensoft.net/fig/1348779">21 D F</figureCitation>
, epigyne as in Fig.
<figureCitation id="77220A96983B6068FBCC498EEAEC4944" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Tibellus japonicus, female, habitus (A C) and epigyne (D, E) A dorsal view B ventral view C lateral view D ventral view E dorsal view. Abbreviations: CO = copulatory opening; FD = fertilisation duct; GM = glandular mound; MS = median septum; R = receptaculum. Scale bars: 2 mm (equal for A C); 0.2 mm (equal for D, E)." figureDoi="10.3897/zookeys.1240.149456.figure22" httpUri="https://binary.pensoft.net/fig/1348780">22 D, E</figureCitation>
.
</paragraph>
<caption id="67DB2F7316FC31F9F9B8FEF06F9A210D" ID-DOI="10.3897/zookeys.1240.149456.figure21" ID-arpha="5D9471DE-D74F-56CC-9301-29BB85C3C67B" httpUri="https://binary.pensoft.net/fig/1348779" startId="F21">
<paragraph id="CEBA99ACD4F2B92E584663D266D9415E">
<label id="216CC9F6CB1CD00F49D635DB96413932">Figure 21.</label>
</paragraph>
<paragraph id="E19B4D73AE26477D12BE9BE32AC8A718">
<taxonomicName id="140AAF4A722664D07D860160F01498DF" authorityName="Efimik" authorityYear="1999" class="Arachnida" family="Philodromidae" genus="Tibellus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="88B35EA65078F3DCEE2673AEB3BEDF91" italics="true">Tibellus japonicus</emphasis>
</taxonomicName>
, male, habitus (
<emphasis id="23FA434D77A3F4C7DB16FE3133452E8F" bold="true">A C</emphasis>
) and left palp (
<emphasis id="AEA32BACCB96CAC2A39B397A95F36AA9" bold="true">D F</emphasis>
)
<emphasis id="880D58C3E38CBDB6A8464F9068DB80C6" bold="true">A</emphasis>
dorsal view
<emphasis id="F7522A5F2473E295D255D8A025A99008" bold="true">B</emphasis>
ventral view
<emphasis id="9B3AAF8C58D0D11BC71D86D18593C949" bold="true">C</emphasis>
lateral view
<emphasis id="3F0A63AACAAC727A6C5F18049A04C964" bold="true">D</emphasis>
prolateral view
<emphasis id="1A8A98C245158D2A2AE8C6A4EFC2BA0E" bold="true">E</emphasis>
ventral view
<emphasis id="E3AE66C7761AACC487A00C2E3486C53A" bold="true">F</emphasis>
retrolateral view. Abbreviations:
<abbrev id="ABBRID0EV5CI" xlink_title="ascending part of sperm duct loop">aSDL</abbrev>
= ascending part of sperm duct loop;
<abbrev id="ABBRID0EZ5CI" xlink_title="conductor">Con</abbrev>
= conductor;
<abbrev id="ABBRID0E45CI" xlink_title="cymbium">Cy</abbrev>
= cymbium;
<abbrev id="ABBRID0EB6CI" xlink_title="descending part of sperm duct loop">dSDL</abbrev>
= descending part of sperm duct loop;
<abbrev id="ABBRID0EF6CI" xlink_title="embolic base">EmB</abbrev>
= embolic base;
<abbrev id="ABBRID0EJ6CI" xlink_title="embolic tip">EmT</abbrev>
= embolic tip;
<abbrev id="ABBRID0EN6CI" xlink_title="retrolateral tibial apophysis">RTA</abbrev>
= retrolateral tibial apophysis;
<abbrev id="ABBRID0ER6CI" xlink_title="sperm duct">SD</abbrev>
= sperm duct;
<abbrev id="ABBRID0EV6CI" xlink_title="tegulum">Te</abbrev>
= tegulum. Scale bars: 1 mm (equal for
<emphasis id="1ED972BAB516DE2E84F1A7A3D2CEA03E" bold="true">A C</emphasis>
); 0.2 mm (equal for
<emphasis id="4FDA9CD99ABD3666E7FEC43F8159977A" bold="true">D F</emphasis>
).
</paragraph>
</caption>
<caption id="24592C5195B3E741779BE777EFEB8815" ID-DOI="10.3897/zookeys.1240.149456.figure22" ID-arpha="F9596938-E310-5DA9-AD36-E9B6F4892310" httpUri="https://binary.pensoft.net/fig/1348780" startId="F22">
<paragraph id="E7B5696DB27F4C6AE36C2BE65ECE124F">
<label id="74538A5504BBDB78BACA1F378E5A19E8">Figure 22.</label>
</paragraph>
<paragraph id="D53018896B455CFE99219C8A642F30DC">
<taxonomicName id="D87194C8A0BD4486AC803EED6CB2566C" authorityName="Efimik" authorityYear="1999" class="Arachnida" family="Philodromidae" genus="Tibellus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="A41CE4D027D5B134C3E32637FF1F4FFD" italics="true">Tibellus japonicus</emphasis>
</taxonomicName>
, female, habitus (
<emphasis id="8AF458076BD0A3AB63C13C976D73CF2B" bold="true">A C</emphasis>
) and epigyne (
<emphasis id="765AF70A19D1B2FF3999D75000B5DB78" bold="true">D, E</emphasis>
)
<emphasis id="AA27427A32188A77AE4C2E161E22E03D" bold="true">A</emphasis>
dorsal view
<emphasis id="F7A6E41294CDBFF7C7EEC4008593573A" bold="true">B</emphasis>
ventral view
<emphasis id="597668830CFD6006D04E511D1D7B56F7" bold="true">C</emphasis>
lateral view
<emphasis id="4F2B2B480030459F00F400221A2E29C9" bold="true">D</emphasis>
ventral view
<emphasis id="7FE8FB30013B30622245D2CD01A399CF" bold="true">E</emphasis>
dorsal view. Abbreviations:
<abbrev id="ABBRID0EDBDI" xlink_title="copulatory opening">CO</abbrev>
= copulatory opening;
<abbrev id="ABBRID0EHBDI" xlink_title="fertilisation duct">FD</abbrev>
= fertilisation duct;
<abbrev id="ABBRID0ELBDI" xlink_title="glandular mound">GM</abbrev>
= glandular mound;
<abbrev id="ABBRID0EPBDI" xlink_title="median septum">MS</abbrev>
= median septum; R = receptaculum. Scale bars: 2 mm (equal for
<emphasis id="1EEE15DBEA825609743CBD634A33CE2A" bold="true">A C</emphasis>
); 0.2 mm (equal for
<emphasis id="5AD604F3EEAABCD684EA537D536D4177" bold="true">D, E</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0E3BDI" type="distribution">
<paragraph id="7DECDCEE44975ADB3977BAF517E2A623">
<heading id="8CBE5848BBE921D029185E0597E24B4A" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="ADE9C9914149E80F4DE7982A2ABDBF4B">
<collectingCountry id="DD0DCE9FD3930C6B90968CE58A00BE13" name="Russia">Russia</collectingCountry>
(Southern
<collectingRegion id="865E952236A18C1641DD0B6161EA3471" country="Russia" name="Sakhalin">Sakhalin</collectingRegion>
),
<collectingCountry id="6F1137E1365F5BC030A9AF49F303B01C" name="China">China</collectingCountry>
(
<collectingRegion id="652FCB3E84F6A4297E1D4A7C35E361D1" country="China" name="Guizhou">Guizhou</collectingRegion>
,
<collectingRegion id="B8D7F35E5C2FC33CF15C6B0DCE76FADB" country="China" name="Hubei">Hubei</collectingRegion>
; Fig.
<figureCitation id="E07CD4C58943A1A2F48BE221F40D5D1C" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Distribution records of the philodromid species in Guizhou Province (A) and Hubei Province (B)." figureDoi="10.3897/zookeys.1240.149456.figure23" httpUri="https://binary.pensoft.net/fig/1348781">23</figureCitation>
),
<collectingCountry id="AEC61DCA3304F2DD1B6DBFDE19D98869" name="South Korea">Korea</collectingCountry>
,
<collectingCountry id="2E1ED911FFF8667E4AEDE1E44EBFA6B1" name="Japan">Japan</collectingCountry>
.
</paragraph>
<caption id="DFEF0254FA8DEE7886CE9A8ADCE7D686" ID-DOI="10.3897/zookeys.1240.149456.figure23" ID-arpha="C89D364B-7F7B-53DB-941F-8CC0633E8B65" httpUri="https://binary.pensoft.net/fig/1348781" startId="F23">
<paragraph id="8724D40DB0C6FF7202EAB4A83D047D8A">
<label id="A0E45BFCB9CF0B0A299DFD31BDD4EFC0">Figure 23.</label>
</paragraph>
<paragraph id="A4DA9F95491943A1B88D68B6C20475FD">
Distribution records of the philodromid species in Guizhou Province (
<emphasis id="0D55A78C009C3A23BC418D2879CCF020" bold="true">A</emphasis>
) and Hubei Province (
<emphasis id="61496F4210AED756EFD885BAE54406F6" bold="true">B</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EXCDI" type="comments">
<paragraph id="B2CF8C5B9ED0C6D17F5E1DADE0743858">
<heading id="F0A0805A0DBFB53C35F43996ED674BE6" reason="title">Comments.</heading>
</paragraph>
<paragraph id="EEFBA9A08E5217AAC0A626B95FFA02E1">
The species was reported as a new record for
<collectingCountry id="42BA4D86C211905450FF50F59426AC85" name="China">China</collectingCountry>
by
<bibRefCitation id="D0F92B8E761E9F47855240124EFC26A6" DOI="10.3969/j.issn.1005-9628.2003.02.006" author="Chen HM &amp; Zhang JX &amp; Song DX" etAl="et al." firstAuthor="Chen" journalOrPublisher="Acta Arachnologica Sinica" pagination="91-93" refId="B8" refString="Chen HM, Zhang JX, Song DX (2003) A newly recorded species of the family Philodromidae from China (Arachnida: Araneae). Acta Arachnologica Sinica 12: 9193. https://doi.org/10.3969/j.issn.1005-9628.2003.02.006" title="A newly recorded species of the family Philodromidae from China (Arachnida: Araneae)." volume="12" year="2003">Chen et al. (2003)</bibRefCitation>
based on materials from Xishui National Nature Reserve in
<collectingRegion id="1258F10876DE5B04E42B6D29B78544D1" country="China" name="Guizhou">Guizhou</collectingRegion>
. Subsequent literature has reported the distribution of this species in
<collectingRegion id="7AEFA9609F2D5EEC7DFDD3960B3B647C" country="China" name="Henan">Henan</collectingRegion>
and
<collectingRegion id="D67A2C17A06F86C73706CBA41B231AC5" country="China" name="Hunan">Hunan</collectingRegion>
provinces (
<bibRefCitation id="673E7FA62B3B769797B0E63481134450" author="Zhu and Zhang" firstAuthor="Zhu" journalOrPublisher="Science Press, Beijing" refId="B67" refString="Zhu MS, Zhang BS (2011) Spider Fauna of Henan: Arachnida: Araneae. Science Press, Beijing, 558 pp." title="Spider Fauna of Henan: Arachnida: Araneae." year="2011">Zhu and Zhang 2011</bibRefCitation>
;
<bibRefCitation id="75BAA9AA41A82FE02E53CDE4E9E77A7A" author="Yin" etAl="et al." firstAuthor="Yin" journalOrPublisher="Hunan Science and Technology Press, Changsha" refId="B62" refString="Yin CM, Peng XJ, Yan HM, Bao YH, Xu X, Tang G, Zhou QS, Liu P (2012) Fauna Hunan: Araneae in Hunan, China. Hunan Science and Technology Press, Changsha, 1590 pp." title="Fauna Hunan: Araneae in Hunan, China." year="2012">Yin et al. 2012</bibRefCitation>
). Newly available specimens indicate that the species is also distributed in Guiyang City and Fanjingshan National Nature Reserve in
<collectingRegion id="7A7AE59D9D605CD4C03F26B414BE6CB4" country="China" name="Guizhou">Guizhou Province</collectingRegion>
(Fig.
<figureCitation id="727E8766F2EC3EC7985899858CF49DD5" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Distribution records of the philodromid species in Guizhou Province (A) and Hubei Province (B)." figureDoi="10.3897/zookeys.1240.149456.figure23" httpUri="https://binary.pensoft.net/fig/1348781">23 A</figureCitation>
), as well as Jiugongshan National Nature Reserve in
<collectingRegion id="D3CED8644D31CBAA2F373DF4FCA14428" country="China" name="Hubei">Hubei Province</collectingRegion>
(Fig.
<figureCitation id="2533E8DF09D6D74B9F22165D1CE97E8C" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Distribution records of the philodromid species in Guizhou Province (A) and Hubei Province (B)." figureDoi="10.3897/zookeys.1240.149456.figure23" httpUri="https://binary.pensoft.net/fig/1348781">23 B</figureCitation>
). This paper provides the first formal report of the species distribution in
<collectingRegion id="8A32BF672AB6A4B275CD4CA654370E76" country="China" name="Hubei">Hubei Province</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>