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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1147.93824" ID-Pensoft-Pub="1313-2970-1147-1" ID-Pensoft-UUID="FF2B6B317B505F9EA0E1000BDCD16CE7" ID-ZooBank="9BB9881B0076473D8E53155D37CA1F50" ModsDocID="1313-2970-1147-1" checkinTime="1676378625502" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Hunyadi, Andras, Vermeulen, Jaap J., Grego, Jozef, Sutcharit, Chirasak, Reischuetz, Alexander, Dumrongrojwattana, Pongrat, Botta-Dukat, Zoltan, Oerstan, Aydin, Fekete, Judit &amp; Jochum, Adrienne" docDate="2023" docId="CCD4366580D95139BA3F3FC55EFC40D5" docLanguage="en" docName="ZooKeys 1147: 1-177" docOrigin="ZooKeys 1147" docPubDate="2023-02-13" docSource="http://dx.doi.org/10.3897/zookeys.1147.93824" docTitle="Angustopila tetradon Pall-Gergely &amp; Hunyadi 2023, sp. nov." docType="treatment" docUuid="F84D6F13-0CC2-4D02-955A-1C4549C121B7" docUuidSource="ZooBank" docVersion="1" id="FF2B6B317B505F9EA0E1000BDCD16CE7" lastPageNumber="1" masterDocId="FF2B6B317B505F9EA0E1000BDCD16CE7" masterDocTitle="Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)" masterLastPageNumber="177" masterPageNumber="1" pageNumber="1" updateTime="1676378625502" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pall-Gergely, Barna</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
<mods:affiliation>Centre for Agricultural Research, Plant Protection Institute, Eoetvoes Lorand Research Network, Herman Otto ut 15, H- 1022 Budapest, Hungary</mods:affiliation>
<mods:nameIdentifier type="email">pallgergely2@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hunyadi, Andras</mods:namePart>
<mods:affiliation>Adria setany 10 G 2 / 5., H- 1148 Budapest, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vermeulen, Jaap J.</mods:namePart>
<mods:affiliation>JK Art and Science, Lauwerbes 8, 2318 AT Leiden, Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Grego, Jozef</mods:namePart>
<mods:affiliation>Horna Micina 219, SK- 97401 Banska Bystrica, Slovakia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sutcharit, Chirasak</mods:namePart>
<mods:affiliation>Animal Systematic Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Reischuetz, Alexander</mods:namePart>
<mods:affiliation>Puechhaimgasse 52, A- 3580 Horn, Austria</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dumrongrojwattana, Pongrat</mods:namePart>
<mods:affiliation>Department of Biology, Faculty of Science, Burapha University, 169 Longhardbangsaen Road, Muang District, Chonburi, 20131, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Botta-Dukat, Zoltan</mods:namePart>
<mods:affiliation>Centre for Ecological Research, Institute of Ecology and Botany, Alkotmany 2 - 4, H- 2600, Vacratot, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Oerstan, Aydin</mods:namePart>
<mods:affiliation>12501 Milestone Manor Lane, Germantown, Maryland, 20876, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fekete, Judit</mods:namePart>
<mods:affiliation>University of Pannonia, Centre of Natural Science, Research Group of Limnology, Egyetem u. 10, H- 8200 Veszprem, Hungary &amp; Centre for Ecological Research, Institute of Aquatic Ecology, Department of Tisza Research, 18 / c Bem square, H- 4026 Debrecen, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jochum, Adrienne</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6624-6412</mods:nameIdentifier>
<mods:affiliation>Naturhistorisches Museum der Burgergemeinde Bern, CH- 3005 Bern, Switzerland &amp; Institute of Ecology and Evolution, University of Bern, CH- 3012 Bern, Switzerland &amp; Senckenberg Forschungsinstitut und Naturmuseum, 60325 Frankfurt am Main, Germany</mods:affiliation>
</mods:name>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-02-13</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1147</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>177</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1147.93824</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1147.93824</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1147-1</mods:identifier>
<mods:identifier type="ZooBank">9BB9881B0076473D8E53155D37CA1F50</mods:identifier>
<mods:identifier type="Pensoft-UUID">FF2B6B317B505F9EA0E1000BDCD16CE7</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:zoobank.org:act:F84D6F13-0CC2-4D02-955A-1C4549C121B7" httpUri="http://treatment.plazi.org/id/CCD4366580D95139BA3F3FC55EFC40D5" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="https://zoobank.org/F84D6F13-0CC2-4D02-955A-1C4549C121B7" authority="Páll-Gergely &amp; Hunyadi" authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Angustopila tetradon" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tetradon" status="sp. nov.">
Angustopila tetradon
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Hunyadi
</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 93" captionStartId="F93" captionText="Figure 93. Angustopila tetradon Pall-Gergely &amp; Hunyadi, sp. nov. Paratype (A, G) and holotype (HNHM 105320) (B-F, H-J). Apertural (A, B), lateral (C), apical (D) and ventral (E) sides of the shell; sculpture on the protoconch (F); aperture (G, H); ventral (I) and frontal (J) surface of the body whorl." figureDoi="10.3897/zookeys.1147.93824.figure93" httpUri="https://binary.pensoft.net/fig/808902" pageId="0" pageNumber="1">Figs 93</figureCitation>
<figureCitation captionStart="Figure 94" captionStartId="F94" captionText="Figure 94. Angustopila tetradon Pall-Gergely &amp; Hunyadi, sp. nov., sample 2020 / 40. Specimen 1 (A, C) and Specimen 2 (B, D)." figureDoi="10.3897/zookeys.1147.93824.figure94" httpUri="https://binary.pensoft.net/fig/808903" pageId="0" pageNumber="1">, 94</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
: Vietnam • 1 empty shell (H: 0.82 mm, D: 0.79); Thanh
<normalizedToken originalValue="Hóa">Hoa</normalizedToken>
Province,
<normalizedToken originalValue="Như">Như</normalizedToken>
Thanh District, Hải
<normalizedToken originalValue="Vân">Van</normalizedToken>
, Hang
<normalizedToken originalValue="Lò">Lo</normalizedToken>
Cao
<normalizedToken originalValue="Kháng">Khang</normalizedToken>
Chiến, vicinity of the cave (locality code: 2020/41);
<geoCoordinate degrees="19" direction="north" minutes="37.08" orientation="latitude" precision="9" value="19.618">19°37.08'N</geoCoordinate>
,
<geoCoordinate degrees="105" direction="east" minutes="34.63" orientation="longitude" precision="9" value="105.57716">105°34.63'E</geoCoordinate>
; 20 m a.s.l.; 14 Feb. 2020; A. Hunyadi leg.; HNHM 105320.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
: Vietnam • 76 shells; same data as for holotype; coll. HA • 3 shells; same data as for holotype; coll. JJV • 48 shells; Thanh
<normalizedToken originalValue="Hóa">Hoa</normalizedToken>
Province,
<normalizedToken originalValue="Như">Như</normalizedToken>
Thanh District, 600 m south from
<normalizedToken originalValue="Xuân">Xuan</normalizedToken>
Khang along road no. 45, around the cave temple (locality code: 2020/40);
<geoCoordinate degrees="19" direction="north" minutes="40.35" orientation="latitude" precision="9" value="19.6725">19°40.35'N</geoCoordinate>
,
<geoCoordinate degrees="105" direction="east" minutes="31.07" orientation="longitude" precision="9" value="105.51783">105°31.07'E</geoCoordinate>
; 80 m a.s.l.; 14 Feb. 2020; A. Hunyadi leg.; coll. HA.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Additional material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Vietnam" location="Thanh District" specimenCount="1" stateProvince="Thanh Hoa Province">
<collectingCountry name="Vietnam">Vietnam</collectingCountry>
• 22 j/b shells; same data as for holotype; coll. HA • 1 j/b shell;
<collectingRegion country="Vietnam" name="Thanh Hoa">
Thanh
<normalizedToken originalValue="Hóa">Hoa</normalizedToken>
Province
</collectingRegion>
,
<normalizedToken originalValue="Như">Như</normalizedToken>
<location LSID="urn:lsid:plazi:treatment:CCD4366580D95139BA3F3FC55EFC40D5:FD581B4106F68853706B04FAA470D49B" country="Vietnam" name="Thanh District" stateProvince="Thanh Hoa Province">Thanh District</location>
</materialsCitation>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">600 m</quantity>
south from
<normalizedToken originalValue="Xuân">Xuan</normalizedToken>
Khang along road no. 45, around the cave temple, (locality code: 2020/40);
<geoCoordinate degrees="19" direction="north" minutes="40.35" orientation="latitude" precision="9" value="19.6725">19°40.35'N</geoCoordinate>
,
<geoCoordinate degrees="105" direction="east" minutes="31.07" orientation="longitude" precision="9" value="105.51783">105°31.07'E</geoCoordinate>
;
<quantity metricMagnitude="1" metricUnit="m" metricValue="8.0" unit="m" value="80.0">80 m</quantity>
a.s.l.;
<date value="2020-02-14">14 Feb. 2020</date>
; A. Hunyadi leg.; coll. HA.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
A small, conical
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila</emphasis>
</taxonomicName>
species with a strong parietal tooth, a strong subcolumellar tooth, and two strong palatal teeth situated relatively close to peristome edge.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">Shell small for the genus, slightly higher than wide or slightly wider than high; colourless, translucent, conical-globular with a high-domed spire; body whorl widest from standard apertural view; protoconch consists of ca. 1.5 whorls, with weak signs of spiral striation preceding the first teleoconch whorl; teleoconch with some fine, irregular radial growth lines and much stronger, equidistantly-arranged spiral striae (ca. 15-17 on body whorl from standard apertural view); whorls 3.75-4, slightly shouldered; aperture slightly curved and in line with shell axis in lateral view; umbilicus wide; aperture subquadrate, sinulus broad; peristome expanded, not reflected; parietal callus not protruding, not detached from penultimate whorl; parietal tooth prominent, high, perpendicular to parietal side or slightly bent, almost reaching edge of peristome; palatal wall with two rather low but prominent palatal teeth (upper and lower palatal teeth), situated in rather small distance from peristome; subcolumellar tooth low, blunt, nearly reaching peristome edge.</paragraph>
<caption doi="10.3897/zookeys.1147.93824.figure93" httpUri="https://binary.pensoft.net/fig/808902" pageId="0" pageNumber="1" start="Figure 93" startId="F93">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 93.</emphasis>
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila tetradon" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tetradon">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila tetradon</emphasis>
</taxonomicName>
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Hunyadi, sp. nov. Paratype (
<emphasis bold="true" pageId="0" pageNumber="1">A, G</emphasis>
) and holotype (HNHM 105320) (
<emphasis bold="true" pageId="0" pageNumber="1">B-F, H-J</emphasis>
). Apertural (
<emphasis bold="true" pageId="0" pageNumber="1">A, B</emphasis>
), lateral (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
), apical (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
) and ventral (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
) sides of the shell; sculpture on the protoconch (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
); aperture (
<emphasis bold="true" pageId="0" pageNumber="1">G, H</emphasis>
); ventral (
<emphasis bold="true" pageId="0" pageNumber="1">I</emphasis>
) and frontal (
<emphasis bold="true" pageId="0" pageNumber="1">J</emphasis>
) surface of the body whorl.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1147.93824.figure94" httpUri="https://binary.pensoft.net/fig/808903" pageId="0" pageNumber="1" start="Figure 94" startId="F94">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 94.</emphasis>
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila tetradon" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tetradon">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila tetradon</emphasis>
</taxonomicName>
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Hunyadi, sp. nov., sample 2020/40. Specimen 1 (
<emphasis bold="true" pageId="0" pageNumber="1">A, C</emphasis>
) and Specimen 2 (
<emphasis bold="true" pageId="0" pageNumber="1">B, D</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Measurements (in mm).</paragraph>
<paragraph pageId="0" pageNumber="1">
H = 0.73-0.87, D = 0.74-0.84, H/D*100 = 92.8-113 (
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
= 14), RUD = 26.3-30.8 (
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
= 4).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
The upper palatal tooth of
<taxonomicName lsidName="A. quadridens" pageId="0" pageNumber="1" rank="species" species="quadridens">
<emphasis italics="true" pageId="0" pageNumber="1">A. quadridens</emphasis>
</taxonomicName>
sp. nov. is situated deeper, its subcolumellar tooth is more elevated, and its sinulus is narrower resulting in a more elongated aperture shape. Moreover, the shells of
<taxonomicName lsidName="A. tetradon" pageId="0" pageNumber="1" rank="species" species="tetradon">
<emphasis italics="true" pageId="0" pageNumber="1">A. tetradon</emphasis>
</taxonomicName>
sp. nov. are more conical (instead of conical-globular) than those of
<taxonomicName lsidName="A. quadridens" pageId="0" pageNumber="1" rank="species" species="quadridens">
<emphasis italics="true" pageId="0" pageNumber="1">A. quadridens</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
The specific epithet refers to the four apertural barriers of this new species (
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila tetradon" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tetradon">Angustopila tetradon</taxonomicName>
:
<emphasis italics="true" pageId="0" pageNumber="1">
<normalizedToken originalValue="τετρα">τετρα</normalizedToken>
</emphasis>
= four, and
<emphasis italics="true" pageId="0" pageNumber="1">
<normalizedToken originalValue="ὀδούς">ὀδούς</normalizedToken>
</emphasis>
= tooth).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is known from two geographically adjacent sites in Thanh
<normalizedToken originalValue="Hóa">Hoa</normalizedToken>
Province, northern Vietnam (Fig.
<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Distribution of Angustopila species." figureDoi="10.3897/zookeys.1147.93824.figure55" httpUri="https://binary.pensoft.net/fig/808864" pageId="0" pageNumber="1">55</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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data/CC/D4/CF/CCD4CF8C9D1A538F8702BE19E776D116.xml Normal file
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<mods:affiliation>Sofia University, Sofia, Bulgaria</mods:affiliation>
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<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-05-09</mods:number>
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<mods:number>10</mods:number>
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<taxonomicName LSID="CCD4CF8C-9D1A-538F-8702-BE19E776D116" authority="(Forel, 1911)" authorityName="Mayr" authorityYear="1861" baseAuthorityName="Forel" baseAuthorityYear="1911" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Fungi" lsidName="Temnothorax aeolius" order="Hymenoptera" pageId="0" pageNumber="83658" phylum="Ascomycota" rank="species" species="aeolius">Temnothorax aeolius (Forel, 1911)</taxonomicName>
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<paragraph pageId="0" pageNumber="83658">Distribution</paragraph>
<paragraph pageId="0" pageNumber="83658">
New records: South Pirin Mt., Kalimantsi vill., 06.04.-10.05.2002, pitfall traps, 1 w.; same place, 06.08-08.09.2002, pitfall traps, 1 w., leg. M. Langourov; Struma Valley, Kamenitsa vill., 23.06-08.08.2002, pitfall traps, 1 w., leg. D. Chobanov. Detailed occurrence data:
<bibRefCitation author="Lapeva-Gjonova, A." journalOrPublisher="Biodiversity Data Journal" pageId="0" pageNumber="83658" refId="B7741022" refString="Lapeva-Gjonova, A., Borowiec, L., 2022. New and little-known ant species (Hymenoptera, Formicidae) from Bulgaria. In: Biodiversity Data Journal. Occurrence dataset." title="New and little-known ant species (Hymenoptera, Formicidae) from Bulgaria" volumeTitle="Biodiversity Data Journal. Occurrence dataset." year="2022">Lapeva-Gjonova and Borowiec (2022)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="83658" type="notes">
<paragraph pageId="0" pageNumber="83658">Notes</paragraph>
<paragraph pageId="0" pageNumber="83658">
First record for Bulgaria. A rarely collected ant that is a member of the
<taxonomicName authority="Mayr, 1861" authorityName="Mayr" authorityYear="1861" baseAuthorityName="Forel" baseAuthorityYear="1911" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Fungi" lsidName="Temnothorax graecus" order="Hymenoptera" pageId="0" pageNumber="83658" phylum="Ascomycota" rank="species" species="graecus">
<emphasis italics="true" pageId="0" pageNumber="83658">Temnothorax graecus</emphasis>
</taxonomicName>
species group (
<bibRefCitation author="Salata, S." journalOrPublisher="ZooKeys" pageId="0" pageNumber="83658" pagination="81 - 131" refId="B7725952" refString="Salata, S., Borowiec, L., 2019. Preliminary division of not socially parasitic Greek Temnothorax Mayr, 1861 (Hymenoptera, Formicidae) with a description of three new species. ZooKeys 87: 81 - 131" title="Preliminary division of not socially parasitic Greek Temnothorax Mayr, 1861 (Hymenoptera, Formicidae) with a description of three new species" volume="87" year="2019">Salata and Borowiec 2019b</bibRefCitation>
). This species is known from Turkey, its type locality, as well as from Israel and the Greek islands (
<bibRefCitation author="Borowiec, L." journalOrPublisher="Genus" pageId="0" pageNumber="83658" pagination="461 - 563" refId="B7725651" refString="Borowiec, L., Salata, S., 2012. Ants of Greece - checklist, comments and new faunistic data (Hymenoptera: Formicidae). Genus 23 (4): 461 - 563" title="Ants of Greece - checklist, comments and new faunistic data (Hymenoptera: Formicidae)" volume="23" year="2012">Borowiec and Salata 2012</bibRefCitation>
). It was recently recorded in Greek Thrace (
<bibRefCitation author="Bracko, G." journalOrPublisher="Biodiversity Data Journal" pageId="0" pageNumber="83658" refId="B7725696" refString="Bracko, G., Kiran, K., Karaman, C., Salata, S., Borowiec, L., 2016. Survey of the ants (Hymenoptera: Formicidae) of the Greek Thrace. Biodiversity Data Journal 4 (7945)" title="Survey of the ants (Hymenoptera: Formicidae) of the Greek Thrace" volume="4" year="2016">
<normalizedToken originalValue="Bračko">Bracko</normalizedToken>
et al. 2016
</bibRefCitation>
).
</paragraph>
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</treatment>
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<emphasis id="BB69D23C40D55181AC99C9959EA8B05C" italics="true" pageNumber="924">Tachyoryctes ruandae</emphasis>
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<emphasis id="986A99282173D2EE5396E9AE76861646" inLineHeading="true" pageNumber="924">Type Locality:</emphasis>
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<collectingCountry id="78557FDCABB9373ECB4FBB5D5731F12A" name="Rwanda">Rwanda</collectingCountry>
, Mt Muhavura.
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<emphasis id="E1C789FB09B1779EC7744E1CC695A3F5" inLineHeading="true" pageNumber="924">Vernacular Names:</emphasis>
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<collectingCountry id="B4B29D9C92678B0F5DC0514154E56143" name="Rwanda">Rwanda</collectingCountry>
African Mole Rat
</vernacularName>
.
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<paragraph id="55EEB8D77110CE1FFDA40B53499B7A3B" pageNumber="924">
<emphasis id="EFEDD2D6360F33C17201A65F045E2129" inLineHeading="true" pageNumber="924">Distribution:</emphasis>
E Dem. Rep.
<collectingCountry id="3D1C91D8554AE1CA49EEF969B23CFB26" name="Democratic Republic of the Congo">Congo</collectingCountry>
(
<collectingRegion id="8672BADD88602282F7C1BCC71DCF9176" country="Democratic Republic of the Congo" name="Sud-Kivu">Kivu</collectingRegion>
),
<collectingCountry id="1BA2FC940635FD4A1AE96765A955CBC6" name="Rwanda">Rwanda</collectingCountry>
, and
<collectingCountry id="2C568BE2DD7CB26A1BE0E658F0E2770A" name="Burundi">Burundi</collectingCountry>
.
</paragraph>
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<emphasis id="EDAB7B880E2BD7D55530D821663BD34F" inLineHeading="true" pageNumber="924">Conservation:</emphasis>
<collectionCode id="70C2BDAA19E397882912D260140A8410">IUCN</collectionCode>
– Lower Risk (lc).
</paragraph>
</subSubSection>
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<paragraph id="43E9E9CFF0989552A2266C2A4D4012E1" pageNumber="924">
<emphasis id="AB4DC90A3D074E4E2C0F562DCEF43D3E" inLineHeading="true" pageNumber="924">Discussion:</emphasis>
See
<bibRefCitation id="C1C83A615905592DAC2331F8948E4DE1" author="Elbl, A. &amp; U. Rahm &amp; G. Mathys" refId="ref92480" refString="Elbl, A., U. Rahm, and G. Mathys. 1966. Les mammiferes et leurs tiques dans La Foret du Ruggege (Republique Rwandaise). Acta Tropica, 23: 223 - 263." year="1966">Elbl et al. (1966)</bibRefCitation>
and
<bibRefCitation id="8420E022812101977DA6A9324B42E149" author="Rahm, U." refId="ref296017" refString="Rahm, U. 1967. Les Murides des environs du Lac Kivu et des regions voisines (Afrique Centrale) et leur ecologie. Revue Suisse de Zoologie, 74: 439 - 519." year="1967">Rahm (1967)</bibRefCitation>
for distributional and ecological information. Stomach morphology described by
<bibRefCitation id="8C55150D37A47ED1696ACBDA95334116" author="Rahm, U." refId="ref296105" refString="Rahm, U. 1976. Zur Morphologie des Magens von Tachyoryctes splendens Ruppell, 1835 (Rodentia, Rhizomyidae). Saugetierkundliche Mitteilungen, 24: 148 - 150." year="1976">Rahm (1976</bibRefCitation>
, as
<taxonomicName id="FAD4A16F913CEAA5B055EFEE716FE037" authorityName="Rüppell" authorityYear="1835" class="Mammalia" family="Spalacidae" genus="Tachyoryctes" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="splendens">
<emphasis id="6F47106EBE58F4559D67FFCFED488BB7" italics="true" pageNumber="924">splendens</emphasis>
</taxonomicName>
) and chromosomal data (2n = 48, FN = 54) reported by
<bibRefCitation id="78D30FB01D5DB096C05EF0AB89BB003A" author="Matthey, R." refId="ref231476" refString="Matthey, R. 1967 a. Note sur la cytogenetique de quelques murides Africains. Mammalia, 31: 281 - 287." year="1967">
Matthey (1967
<emphasis id="98A0EDC822599049A04CA0D6E5D4864B" italics="true" pageNumber="924">a</emphasis>
)
</bibRefCitation>
.
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