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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.46.9069" ID-PMC="PMC4391959" ID-Pensoft-Pub="1314-2003-46-73" ID-Pensoft-UUID="4A26DD5DFFADE677FFA2A049FFFFFFC2" ID-PubMed="25878551" ID-Zenodo-Dep="576272" ModsDocID="1314-2003-46-73" checkinTime="1451251575242" checkinUser="pensoft" docAuthor="Deanna, Rocio, Gonzalez, Segundo Leiva &amp; Barboza, Gloria Estela" docDate="2015" docId="CC4B35A1EC085BE8AA5A65C51B7A0A4D" docLanguage="en" docName="PhytoKeys 46: 73-87" docOrigin="PhytoKeys 46" docPubDate="2015-02-27" docSource="http://dx.doi.org/10.3897/phytokeys.46.9069" docTitle="Deprea pomacochaensis Barboza 2015, comb. nov." docType="treatment" docVersion="3" id="4A26DD5DFFADE677FFA2A049FFFFFFC2" lastPageNumber="79" masterDocId="4A26DD5DFFADE677FFA2A049FFFFFFC2" masterDocTitle="Changes in the circumscription of Deprea (Physalideae, Solanaceae): thirty two new combinations" masterLastPageNumber="87" masterPageNumber="73" pageNumber="79" updateTime="1668141316567" updateUser="ExternalLinkService">
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<mods:title>Changes in the circumscription of Deprea (Physalideae, Solanaceae): thirty two new combinations</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Deanna, Rocio</mods:namePart>
<mods:affiliation>Instituto Multidisciplinario de Biologia Vegetal (IMBIV), CONICET and Universidad Nacional de Cordoba, CC 495, CP 5000, Cordoba, Argentina &amp; Facultad de Ciencias Quimicas, Universidad Nacional de Cordoba, Haya de la Torre y Medina Allende, Cordoba, Argentina</mods:affiliation>
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<mods:namePart>Gonzalez, Segundo Leiva</mods:namePart>
<mods:affiliation>Museo de Historia Natural, Universidad Privada Antenor Orrego de Trujillo, CC 1075, Trujillo, Peru</mods:affiliation>
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<mods:namePart>Barboza, Gloria Estela</mods:namePart>
<mods:affiliation>Instituto Multidisciplinario de Biologia Vegetal (IMBIV), CONICET and Universidad Nacional de Cordoba, CC 495, CP 5000, Cordoba, Argentina &amp; Facultad de Ciencias Quimicas, Universidad Nacional de Cordoba, Haya de la Torre y Medina Allende, Cordoba, Argentina</mods:affiliation>
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<taxonomicName LSID="CC4B35A1-EC08-5BE8-AA5A-65C51B7A0A4D" authority="(S. Leiva) Barboza" authorityName="Barboza" authorityYear="2015" baseAuthorityName="Deanna &amp; González &amp; Barboza" baseAuthorityYear="2015" class="Magnoliopsida" family="Solanaceae" genus="Deprea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Deprea pomacochaensis" order="Solanales" pageId="6" pageNumber="79" phylum="Tracheophyta" rank="species" species="pomacochaensis" status="comb. nov.">Deprea pomacochaensis (S.Leiva) Barboza</taxonomicName>
<taxonomicNameLabel pageId="6" pageNumber="79">comb. nov.</taxonomicNameLabel>
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<taxonomicName authorityName="Deanna &amp; González &amp; Barboza" authorityYear="2015" class="Magnoliopsida" family="Solanaceae" genus="Larnax" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Larnax pomacochaensis" order="Solanales" pageId="6" pageNumber="79" phylum="Tracheophyta" rank="species" species="pomacochaensis" status="comb. nov.">Larnax pomacochaensis</taxonomicName>
S.Leiva, Arnaldoa 13 (2): 299. 2006. Basionym
</paragraph>
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<paragraph pageId="6" pageNumber="79">Type.</paragraph>
<paragraph pageId="6" pageNumber="79">
PERU: Amazonas:
<normalizedToken originalValue="Bongará">Bongara</normalizedToken>
, km 328-329 carretera
<normalizedToken originalValue="Bongará-Nueva">Bongara-Nueva</normalizedToken>
Cajamarca, arriba de laguna Pomacochas, 2400 m, 5 Feb 2006 (fl, fr),
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(lectotype, designated in
<bibRefCitation author="Deanna, R" journalOrPublisher="Phytotaxa" pageId="11" pageNumber="84" pagination="1 - 34" publicationUrl="10.11646/phytotaxa.167.1.1" refId="B7" refString="Deanna, R, Leiva Gonzalez, S, Barboza, GE, 2014a. Four new species and eighteen lectotypifications of Larnax from Ecuador and Peru and a new synonym of Deprea orinocensis (Solanaceae: Solanoideae, Physalideae). Phytotaxa 167 (1): 1 - 34, DOI: 10.11646/phytotaxa.167.1.1" title="Four new species and eighteen lectotypifications of Larnax from Ecuador and Peru and a new synonym of Deprea orinocensis (Solanaceae: Solanoideae, Physalideae)." url="10.11646/phytotaxa.167.1.1" volume="167" year="2014 a">Deanna et al. 2014a</bibRefCitation>
, pg. 28: Leiva
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&amp;
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<normalizedToken originalValue="Rodríguez">Rodriguez</normalizedToken>
, Arnaldoa 13(2): 300. Fig. 5. 2006).
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<taxonomicName id="FACDC2B15AA796CC38263766F147A321" ID-CoL="3WXLH" authority="Schellenberg, 1950" authorityName="Schellenberg" authorityYear="1950" class="Malacostraca" family="Talitridae" genus="Macarorchestia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macarorchestia remyi" order="Amphipoda" pageId="1" pageNumber="134" phylum="Arthropoda" rank="species" species="remyi">Macarorchestia remyi (Schellenberg, 1950)</taxonomicName>
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<taxonomicName id="6B6FB6F9D1B22588EB0BB68C034F3DFC" class="Malacostraca" family="Talitridae" genus="Macarorchestia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macarorchestia remyi" order="Amphipoda" pageId="1" pageNumber="134" phylum="Arthropoda" rank="species" species="remyi">Macarorchestia remyi</taxonomicName>
:
<bibRefCitation id="3836858A0B96E9D7AF0939CCEF9D9D68" author="Schellenberg, A" journalOrPublisher="Archiv fur Hydrobiologie" pageId="8" pageNumber="141" pagination="325 - 330" title="Subterrane Amphipoden korsikanischer Biotope." volume="44" year="1950">Schellenberg 1950</bibRefCitation>
: 325;
<bibRefCitation id="052756244CDFFB8DFDAC714A714C676B" author="Ruffo, S" editor="Bellan-Santini, D" journalOrPublisher="Monaco" pageId="8" pageNumber="141" pagination="365 - 814" title="Memoires de l'Institut oceanographique." year="1993">Ruffo 1993</bibRefCitation>
: 739.
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<paragraph id="1A76C6DC2EC93DDB8111F30FFBDFA6F8" pageId="1" pageNumber="134">Material examined.</paragraph>
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Two males, 4 females, 1 juvenile (Crust. 25468) ZMB Museum
<normalizedToken id="E1C662394F16FE83C843F38E13BD9E00" originalValue="für">fuer</normalizedToken>
Naturkunde, Berlin. Collected by A. Schellenberg from a seashore cave at St. Barthelemy, Corisca. A total of 47 individuals (combined sample) of all life history stages, including females with ova, collected by L. Pavesi on 20 January and 15 April 2007 at Principina a Mare, Tyrrhenian Sea, Italy. Sample location co-ordinates:
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,
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. A further sample collected at Principina a Mare by L. Pavesi on 24 April 2011 consisted of 28 individuals of all life history stages, including females with ova. One individual from
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Island (Ionian Sea, Greece) collected by L. Pavesi in 2007 (Destructively used in DNA analysis).
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<paragraph id="5EC485DE6428F37F900DEE70B9AD4B49" pageId="1" pageNumber="134">Distribution.</paragraph>
<paragraph id="656E1B988264195E00AAE00FC87ED155" pageId="1" pageNumber="134">
Besides the locations found by L. Pavesi for this species, published records include that of
<bibRefCitation id="C33CB4DDCF6D5570C257B4D7F81903A7" pageId="1" pageNumber="134">Ruffo (1960)</bibRefCitation>
in Sardinia and
<bibRefCitation id="712BA4767683E33C250C243EDC0889AE" author="Ruffo, S" editor="Bellan-Santini, D" journalOrPublisher="Monaco" pageId="8" pageNumber="141" pagination="365 - 814" title="Memoires de l'Institut oceanographique." year="1993">Ruffo (1993)</bibRefCitation>
in France, Greece and Italy. Possibly some of these records refer to the species described below.
</paragraph>
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<paragraph id="224B4FD4EFB6D9FEF1D96BF1E39B1EEC" pageId="1" pageNumber="134">Epidermal pigment patterns.</paragraph>
<paragraph id="52A218091003D39B183208F5842BCFC0" pageId="1" pageNumber="134">Absent.</paragraph>
<paragraph id="2F00F7B3C451A60F1744FC01F3556E43" pageId="1" pageNumber="134">
Remarks This is the second largest
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. In recent times the type locality has been destroyed by cleaning beaches for the benefit of tourists (L. Pavesi, pers. comm.).
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<treatment id="CC4B87983246FF90FF1CFD44FCA5E3D0" ID-DOI="http://doi.org/10.5281/zenodo.5531639" ID-Zenodo-Dep="5531639" LSID="urn:lsid:plazi:treatment:CC4B87983246FF90FF1CFD44FCA5E3D0" httpUri="http://treatment.plazi.org/id/CC4B87983246FF90FF1CFD44FCA5E3D0" lastPageId="16" lastPageNumber="498" pageId="14" pageNumber="496">
<subSubSection id="0CF865053246FF8EFF1CFD44FDCDE034" box="[151,537,1835,1862]" pageId="14" pageNumber="496" type="nomenclature">
<paragraph id="445D368E3246FF8EFF1CFD44FDCDE034" blockId="14.[151,537,1835,1897]" box="[151,537,1835,1862]" pageId="14" pageNumber="496">
<heading id="1F1581E23246FF8EFF1CFD44FDCDE034" bold="true" box="[151,537,1835,1862]" fontSize="11" level="1" pageId="14" pageNumber="496" reason="1">
<emphasis id="7696EA9C3246FF8EFF1CFD44FDCDE034" bold="true" box="[151,537,1835,1862]" pageId="14" pageNumber="496">
<taxonomicName id="83E24D0D3246FF8EFF1CFD44FE60E037" ID-CoL="8SP7D" authority="Kasparek, 2021" authorityName="Kasparek" authorityYear="2021" box="[151,436,1835,1862]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="496" phylum="Arthropoda" rank="species" species="flavissimum" status="sp. nov.">
<emphasis id="7696EA9C3246FF8EFF1CFD44FE60E037" bold="true" box="[151,436,1835,1862]" italics="true" pageId="14" pageNumber="496">Anthidium flavissimum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6DA557E73246FF8EFE30FD44FDCDE034" box="[443,537,1835,1861]" pageId="14" pageNumber="496" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0CF865053246FF8EFF1CFD20FEF3E018" box="[151,295,1871,1897]" pageId="14" pageNumber="496" type="description">
<paragraph id="445D368E3246FF8EFF1CFD20FEF3E018" blockId="14.[151,537,1835,1897]" box="[151,295,1871,1897]" pageId="14" pageNumber="496">
(
<figureCitation id="DCD92A0B3246FF8EFF14FD20FF22E018" box="[159,246,1871,1897]" captionStart="FIGURE 10" captionStartId="13.[151,250,1274,1298]" captionTargetBox="[284,1304,188,1241]" captionTargetId="figure-39@13.[277,1311,181,1250]" captionTargetPageId="13" captionText="FIGURE 10. Schematic view of the head in the species of Anthidium, subgenus Gulanthidium. A. Lateral view; B. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5531655" httpUri="https://zenodo.org/record/5531655/files/figure.png" pageId="14" pageNumber="496">Figs 10</figureCitation>
,
<figureCitation id="DCD92A0B3246FF8EFE89FD20FECAE018" box="[258,286,1871,1897]" captionStart="FIGURE 13" captionStartId="15.[151,250,1958,1982]" captionTargetBox="[170,1417,272,1935]" captionTargetId="figure-88@15.[170,1417,272,1935]" captionTargetPageId="15" captionText="FIGURE 13. Anthidium flavissimum sp. nov. Female holotype. A. Habitus dorsal; B. Vertex; C. Habitus lateral; D. Face; E. Head, fore leg and mesepisternum with omaular carina; F. Scutellum and axillae; G. Abdominal terga." figureDoi="http://doi.org/10.5281/zenodo.5531661" httpUri="https://zenodo.org/record/5531661/files/figure.png" pageId="14" pageNumber="496">13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="0CF865053246FF8EFF1CFDF8FDEFE08B" pageId="14" pageNumber="496" type="materials_examined">
<paragraph id="445D368E3246FF8EFF1CFDF8FDEFE08B" blockId="14.[151,1437,1943,2042]" pageId="14" pageNumber="496">
<emphasis id="7696EA9C3246FF8EFF1CFDF8FE90E0C0" bold="true" box="[151,324,1943,1969]" pageId="14" pageNumber="496">Type material:</emphasis>
<materialsCitation id="F48A3CD33246FF8EFEC3FDF8FDEFE08B" collectingDate="1959-05-19" collectionCode="CMK" collectorName="W. Firestone" country="Afghanistan" determinerName="T. Griswold &amp; M. Schwarz" latitude="32.2" location="Dashi-Bakwa" longLatPrecision="1212" longitude="62.916668" pageId="14" pageNumber="496" specimenCount="1" typeStatus="holotype">
<typeStatus id="9B59882C3246FF8EFEC3FDF8FE34E0C0" box="[328,480,1943,1969]" pageId="14" pageNumber="496" type="holotype">HOLOTYPE</typeStatus>
.Female.
<collectingCountry id="3CF5761E3246FF8EFDC8FDF8FD1BE0C0" box="[579,719,1943,1969]" name="Afghanistan" pageId="14" pageNumber="496">Afghanistan</collectingCountry>
,
<location id="413D60553246FF8EFD5CFDF8FCBBE0C0" LSID="urn:lsid:plazi:treatment:CC4B87983246FF90FF1CFD44FCA5E3D0:413D60553246FF8EFD5CFDF8FCBBE0C0" box="[727,879,1943,1969]" country="Afghanistan" latitude="32.2" longLatPrecision="1212" longitude="62.916668" name="Dashi-Bakwa" pageId="14" pageNumber="496">Dashi-Bakwa</location>
[Dasht-e Bakwa, c.
<geoCoordinate id="21D650493246FF8EFBC2FDF8FB78E0C0" box="[1097,1196,1943,1969]" degrees="32" direction="north" minutes="12" orientation="latitude" pageId="14" pageNumber="496" precision="925" value="32.2">32°12’N</geoCoordinate>
,
<geoCoordinate id="21D650493246FF8EFB3EFDF8FAC3E0C3" box="[1205,1303,1943,1970]" degrees="62" direction="east" minutes="55" orientation="longitude" pageId="14" pageNumber="496" precision="925" value="62.916668">62°55’E</geoCoordinate>
],
<date id="305C104E3246FF8EFAACFDF8FA4CE0C3" box="[1319,1432,1943,1970]" pageId="14" pageNumber="496" value="1959-05-19">
<collectingDate id="2018E9A63246FF8EFAACFDF8FA4CE0C3" box="[1319,1432,1943,1970]" pageId="14" pageNumber="496" value="1959-05-19">19.v.1959</collectingDate>
</date>
,
<collectorName id="E91753583246FF8EFF1CFDD3FEF0E0A4" box="[151,292,1979,2006]" pageId="14" pageNumber="496">W. Firestone</collectorName>
leg., “
<emphasis id="7696EA9C3246FF8EFEE5FDD4FE26E0A7" box="[366,498,1979,2006]" italics="true" pageId="14" pageNumber="496">
<taxonomicName id="83E24D0D3246FF8EFEE5FDD4FE37E0A4" authorityName="Fabricius" authorityYear="1804" box="[366,483,1979,2005]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="496" phylum="Arthropoda" rank="genus">Anthidium</taxonomicName>
?
</emphasis>
<taxonomicName id="83E24D0D3246FF8EFE78FDD3FD6DE0A4" authority="Alfken" authorityName="Alfken" authorityYear="1938" box="[499,697,1979,2006]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="496" phylum="Arthropoda" rank="species" species="eremicum">
<emphasis id="7696EA9C3246FF8EFE78FDD3FDB0E0A7" box="[499,612,1980,2006]" italics="true" pageId="14" pageNumber="496">eremicum</emphasis>
Alfken
</taxonomicName>
, det.
<determinerName id="B181DA4A3246FF8EFD79FDD3FCA3E0A4" box="[754,887,1979,2006]" pageId="14" pageNumber="496">T. Griswold</determinerName>
1984”, “
<taxonomicName id="83E24D0D3246FF8EFC50FDD4FACDE0A4" authority="Alfken" authorityName="Alfken" authorityYear="1938" box="[987,1305,1979,2005]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="496" phylum="Arthropoda" rank="species" species="eremicum">
<emphasis id="7696EA9C3246FF8EFC50FDD4FB17E0A4" box="[987,1219,1979,2005]" italics="true" pageId="14" pageNumber="496">Anthidium eremicum</emphasis>
Alfken
</taxonomicName>
, female,
<determinerName id="B181DA4A3246FF8EFAF7FDD3FF23E088" pageId="14" pageNumber="496">M. Schwarz</determinerName>
det. 1986” (
<collectionCode id="22F3AE4B3246FF8EFE09FDB0FE12E088" box="[386,454,2015,2041]" pageId="14" pageNumber="496">CMK</collectionCode>
, ms2726).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="0CF865053247FF8FFF4CFAF9FD54E789" pageId="15" pageNumber="497" type="etymology">
<paragraph id="445D368E3247FF8FFF4CFAF9FD54E789" blockId="15.[151,1437,150,249]" pageId="15" pageNumber="497">
<emphasis id="7696EA9C3247FF8FFF4CFAF9FE72E7C1" bold="true" box="[199,422,150,176]" pageId="15" pageNumber="497">Derivatio nominis.</emphasis>
The epithet of the name refers to the striking yellow colouration of the species: flavus = yellow (adjective) in Latin. The suffix -
<emphasis id="7696EA9C3247FF8FFDC4FAD4FD43E7A5" box="[591,663,187,212]" italics="true" pageId="15" pageNumber="497">issimu</emphasis>
s (resp. -
<emphasis id="7696EA9C3247FF8FFD7CFAD4FC87E7A5" box="[759,851,187,212]" italics="true" pageId="15" pageNumber="497">issimum</emphasis>
as neuter) is added to form its superlative grade as it is the most yellow species of this subgenus.
</paragraph>
</subSubSection>
<caption id="109D66063247FF8FFF1CFDC9FB71E092" ID-DOI="http://doi.org/10.5281/zenodo.5531661" ID-Zenodo-Dep="5531661" httpUri="https://zenodo.org/record/5531661/files/figure.png" pageId="15" pageNumber="497" startId="15.[151,250,1958,1982]" targetBox="[170,1417,272,1935]" targetPageId="15">
<paragraph id="445D368E3247FF8FFF1CFDC9FB71E092" blockId="15.[151,1436,1958,2019]" pageId="15" pageNumber="497">
<emphasis id="7696EA9C3247FF8FFF1CFDC9FEF5E0CE" bold="true" box="[151,289,1958,1983]" pageId="15" pageNumber="497">FIGURE 13.</emphasis>
<taxonomicName id="83E24D0D3247FF8FFEA2FDC8FDC7E0CE" authority="Kasparek, 2021" authorityName="Kasparek" authorityYear="2021" box="[297,531,1959,1983]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="497" phylum="Arthropoda" rank="species" species="flavissimum" status="sp. nov.">
<emphasis id="7696EA9C3247FF8FFEA2FDC8FDC7E0CE" box="[297,531,1959,1983]" italics="true" pageId="15" pageNumber="497">Anthidium flavissimum</emphasis>
</taxonomicName>
<emphasis id="7696EA9C3247FF8FFD90FDC8FDB9E0CE" bold="true" box="[539,621,1959,1983]" pageId="15" pageNumber="497">
<taxonomicNameLabel id="6DA557E73247FF8FFD90FDC8FDB9E0CE" box="[539,621,1959,1983]" pageId="15" pageNumber="497" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Female holotype.
<emphasis id="7696EA9C3247FF8FFCA6FDC9FC92E0CF" bold="true" box="[813,838,1958,1982]" pageId="15" pageNumber="497">A.</emphasis>
Habitus dorsal;
<emphasis id="7696EA9C3247FF8FFC7AFDC8FBDCE0CE" bold="true" box="[1009,1032,1959,1983]" pageId="15" pageNumber="497">B.</emphasis>
Vertex;
<emphasis id="7696EA9C3247FF8FFBEAFDC9FBADE0CF" bold="true" box="[1121,1145,1958,1982]" pageId="15" pageNumber="497">C.</emphasis>
Habitus lateral;
<emphasis id="7696EA9C3247FF8FFAADFDC8FAEBE0CE" bold="true" box="[1318,1343,1959,1983]" pageId="15" pageNumber="497">D.</emphasis>
Face;
<emphasis id="7696EA9C3247FF8FFA0EFDC8FA48E0CE" bold="true" box="[1413,1436,1959,1983]" pageId="15" pageNumber="497">E.</emphasis>
Head, fore leg and mesepisternum with omaular carina;
<emphasis id="7696EA9C3247FF8FFD5BFDA4FD37E092" bold="true" box="[720,739,1995,2019]" pageId="15" pageNumber="497">F.</emphasis>
Scutellum and axillae;
<emphasis id="7696EA9C3247FF8FFC58FDA5FC39E093" bold="true" box="[979,1005,1994,2018]" pageId="15" pageNumber="497">G.</emphasis>
Abdominal terga.
</paragraph>
</caption>
<subSubSection id="0CF865053258FF90FF4CFAF9FACAE6DF" pageId="16" pageNumber="498" type="diagnosis">
<paragraph id="445D368E3258FF90FF4CFAF9FACAE6DF" blockId="16.[151,1437,150,1185]" pageId="16" pageNumber="498">
<emphasis id="7696EA9C3258FF90FF4CFAF9FE9CE7C1" bold="true" box="[199,328,150,176]" pageId="16" pageNumber="498">Diagnosis.</emphasis>
The female can easily be distinguished from all other species of the subgenus by its striking yellow (“canary yellow”) colouration of the entire body with the exception of some black remnants on scutum and in the upper face between antennal sockets and posterior ocelli. Distinguished from
<taxonomicName id="83E24D0D3258FF90FB3FFAB0FABEE788" authorityName="Morawitz" authorityYear="1888" box="[1204,1386,223,249]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="498" phylum="Arthropoda" rank="species" species="anguliventre">
<emphasis id="7696EA9C3258FF90FB3FFAB0FABEE788" box="[1204,1386,223,249]" italics="true" pageId="16" pageNumber="498">A. anguliventre</emphasis>
</taxonomicName>
and
<taxonomicName id="83E24D0D3258FF90FF1CFB6CFEFEE66C" authorityName="Warncke" authorityYear="1980" box="[151,298,259,285]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="498" phylum="Arthropoda" rank="species" species="rotundum">
<emphasis id="7696EA9C3258FF90FF1CFB6CFEFEE66C" box="[151,298,259,285]" italics="true" pageId="16" pageNumber="498">A. rotundum</emphasis>
</taxonomicName>
by a narrow (not wide) impunctate middle line on the clypeus, from
<taxonomicName id="83E24D0D3258FF90FBE8FB6CFB22E66D" authorityName="Alfken" authorityYear="1938" box="[1123,1270,259,285]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="498" phylum="Arthropoda" rank="species" species="eremicum">
<emphasis id="7696EA9C3258FF90FBE8FB6CFB22E66D" box="[1123,1270,259,285]" italics="true" pageId="16" pageNumber="498">A. eremicum</emphasis>
</taxonomicName>
by the lamellate upper omaulus (rounded in
<taxonomicName id="83E24D0D3258FF90FD9AFB48FD70E631" authorityName="Alfken" authorityYear="1938" box="[529,676,295,321]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="498" phylum="Arthropoda" rank="species" species="eremicum">
<emphasis id="7696EA9C3258FF90FD9AFB48FD70E631" box="[529,676,295,321]" italics="true" pageId="16" pageNumber="498">A. eremicum</emphasis>
</taxonomicName>
) and the laterally toothed T3–T5 (bulged in
<taxonomicName id="83E24D0D3258FF90FB3DFB48FA9DE631" authorityName="Alfken" authorityYear="1938" box="[1206,1353,295,321]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="498" phylum="Arthropoda" rank="species" species="eremicum">
<emphasis id="7696EA9C3258FF90FB3DFB48FA9DE631" box="[1206,1353,295,321]" italics="true" pageId="16" pageNumber="498">A. eremicum</emphasis>
</taxonomicName>
), from
<taxonomicName id="83E24D0D3258FF90FF1CFB24FE94E614" authorityName="Kasparek" authorityYear="2021" box="[151,320,331,357]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="498" phylum="Arthropoda" rank="species" species="occidentale">
<emphasis id="7696EA9C3258FF90FF1CFB24FE94E614" box="[151,320,331,357]" italics="true" pageId="16" pageNumber="498">A. occidentale</emphasis>
</taxonomicName>
by the lamellate upper omaulus (angulate in
<taxonomicName id="83E24D0D3258FF90FCD2FB24FBD6E614" authorityName="Kasparek" authorityYear="2021" box="[857,1026,331,357]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="498" phylum="Arthropoda" rank="species" species="occidentale">
<emphasis id="7696EA9C3258FF90FCD2FB24FBD6E614" box="[857,1026,331,357]" italics="true" pageId="16" pageNumber="498">A. occidentale</emphasis>
</taxonomicName>
), and from
<taxonomicName id="83E24D0D3258FF90FB04FB24FA9FE614" authorityName="Kasparek" authorityYear="2021" box="[1167,1355,331,357]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="498" phylum="Arthropoda" rank="species" species="preoccipitale">
<emphasis id="7696EA9C3258FF90FB04FB24FA9FE614" box="[1167,1355,331,357]" italics="true" pageId="16" pageNumber="498">A. preoccipitale</emphasis>
</taxonomicName>
by the rounded hind tibia (rudimentary carina in
<taxonomicName id="83E24D0D3258FF90FD00FB00FC9CE6F8" authorityName="Kasparek" authorityYear="2021" box="[651,840,367,393]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="498" phylum="Arthropoda" rank="species" species="preoccipitale">
<emphasis id="7696EA9C3258FF90FD00FB00FC9CE6F8" box="[651,840,367,393]" italics="true" pageId="16" pageNumber="498">A. preoccipitale</emphasis>
</taxonomicName>
) (
<tableCitation id="096003353258FF90FCE9FB00FC6FE6F8" box="[866,955,367,393]" captionStart="TABLE 1" captionStartId="7.[151,244,834,860]" captionText="TABLE 1. Comparison of some structural traits in the females of five species Anthidium, subgenus Gulanthidium. The female of A. preoccipitale is not known. Highly relevant distinguishing characters are given in bold." pageId="16" pageNumber="498">Table 1</tableCitation>
). Vertex longer than in its consubgeners (3.83 as long as hind ocellus diameter, whereas it varies between 2.4 and
<quantity id="831A9B6B3258FF90FB8AFBFCFB90E6DC" box="[1025,1092,403,429]" metricMagnitude="-2" metricUnit="m" metricValue="7.874" pageId="16" pageNumber="498" unit="in" value="3.1">3.1 in</quantity>
all other species).
</paragraph>
</subSubSection>
<subSubSection id="0CF865053258FF90FF4CFBD9FE43E344" pageId="16" pageNumber="498" type="description">
<paragraph id="445D368E3258FF90FF4CFBD9FBC7E360" blockId="16.[151,1437,150,1185]" pageId="16" pageNumber="498">
<emphasis id="7696EA9C3258FF90FF4CFBD9FE1BE6A0" bold="true" box="[199,463,438,465]" pageId="16" pageNumber="498">Description. Female.</emphasis>
<quantity id="831A9B6B3258FF90FE51FBD8FDF1E6A0" box="[474,549,439,465]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="16" pageNumber="498" unit="mm" value="9.0">9 mm</quantity>
.
<emphasis id="7696EA9C3258FF90FDBFFBD8FDAFE6A0" box="[564,635,439,465]" italics="true" pageId="16" pageNumber="498">Head.</emphasis>
Clypeus sub-trapezoid; anterior margin almost straight, posterior margin with transparent furrows, which are obliquely directed to the centre; clypeus punctate with impunctate middle line; punctures at base more scattered than at apex; mandibles yellow with 5 brown teeth (one tooth broken in the
<typeStatus id="9B59882C3258FF90FE09F84CFE3BE54F" box="[386,495,547,574]" pageId="16" pageNumber="498" type="holotype">holotype</typeStatus>
); teeth except of the lower outer tooth of almost equal size; subantennal suture straight; eyes greenish yellow; inner orbit modestly convergent; punctation on head finer than on clypeus; impunctate area in the supraclypeal area; supraclypeal area dirty-brown to black; upper face between antennal sockets and posterior ocelli black with brownish margins and a yellow spot below the anterior ocellus; area around antennal sockets and below ocelli with depressed white hairs; vertex in dorsal view moderately concave (
<figureCitation id="DCD92A0B3258FF90FE8CF8B8FEA6E580" box="[263,370,727,753]" captionStart="FIGURE 10" captionStartId="13.[151,250,1274,1298]" captionTargetBox="[284,1304,188,1241]" captionTargetId="figure-39@13.[277,1311,181,1250]" captionTargetPageId="13" captionText="FIGURE 10. Schematic view of the head in the species of Anthidium, subgenus Gulanthidium. A. Lateral view; B. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5531655" httpUri="https://zenodo.org/record/5531655/files/figure.png" pageId="16" pageNumber="498">Fig. 10A</figureCitation>
).—
<taxonomicName id="83E24D0D3258FF90FE14F8B8FDC8E581" box="[415,540,727,752]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="498" phylum="Arthropoda" rank="genus">
<emphasis id="7696EA9C3258FF90FE14F8B8FDC8E581" box="[415,540,727,752]" italics="true" pageId="16" pageNumber="498">Mesosoma</emphasis>
</taxonomicName>
. Scutum yellow with some brownish and black maculation, apparently remnants of the scutal pattern typical for the subgenus; scutellum widely rounded posteriorly in dorsal view with median emargination; axilla slightly protruding posteriorly; scutellum somewhat overhanging the propodeum; propodeal triangle rugulose or shagreened, punctate posteriorly; upper half of omaulus with low but distinctive lamella.—
<emphasis id="7696EA9C3258FF90FEFEF908FE3BE4F1" box="[373,495,871,896]" italics="true" pageId="16" pageNumber="498">Metasoma</emphasis>
. Terga yellow, with depressions semi-transparent brownish-yellow; punctation on depression finer than on discs; inconspicuous impunctate marginal zone; T6 concavely depressed on both sides, lateral lamellate protrusion in dorsal view; apex with narrow sting-wide incision; T3–T5 with lateral, partly transparent, acute projections, increasing in size towards T5; scopa white.—
<emphasis id="7696EA9C3258FF90FBF1F9BCFB68E49D" box="[1146,1212,979,1004]" italics="true" pageId="16" pageNumber="498">Legs:</emphasis>
Yellow; hind tibia rounded, hind basitarsus with transparent longitudinal carina on outer face.
</paragraph>
<paragraph id="445D368E3258FF90FF4CFE74FE43E344" blockId="16.[151,1437,150,1185]" box="[199,407,1051,1077]" pageId="16" pageNumber="498">
<emphasis id="7696EA9C3258FF90FF4CFE74FEDFE344" bold="true" box="[199,267,1051,1077]" pageId="16" pageNumber="498">Male.</emphasis>
Not known.
</paragraph>
</subSubSection>
<subSubSection id="0CF865053258FF90FF4CFE51FD0FE30C" pageId="16" pageNumber="498" type="distribution">
<paragraph id="445D368E3258FF90FF4CFE51FD0FE30C" blockId="16.[151,1437,150,1185]" pageId="16" pageNumber="498">
<emphasis id="7696EA9C3258FF90FF4CFE51FEB5E329" bold="true" box="[199,353,1086,1112]" pageId="16" pageNumber="498">Distribution.</emphasis>
Only known from Dasht-e Bakwa, an extensive alluvial plain in the southwest of
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, situated about
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).
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<emphasis id="7696EA9C3258FF90FF4CFEE9FEF3E3D1" bold="true" box="[199,295,1158,1184]" pageId="16" pageNumber="498">Biology.</emphasis>
Collected in May. No further information available.
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</treatment>
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<mods:title id="9A17B01B27FB42DD2149D1EA1548D665">Revision and description of three new species of the Palaearctic subgenus Gulanthidium of the wool carder bee genus Anthidium (Apoidea: Megachilidae: Anthidiini)</mods:title>
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<emphasis id="7696EA9C3258FF90FF1CFE9DFE7EE27C" bold="true" box="[151,426,1266,1293]" italics="true" pageId="16" pageNumber="498">Anthidium occidentale</emphasis>
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(
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,
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,
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<emphasis id="7696EA9C3258FF90FF1CFF30FE93E209" bold="true" box="[151,327,1374,1401]" pageId="16" pageNumber="498">Type material.</emphasis>
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,
<collectorName id="E91753583258FF90FDE3FFA4FD39E294" box="[616,749,1483,1509]" pageId="16" pageNumber="498">Ma. Halada</collectorName>
leg. (
<collectionCode id="22F3AE4B3258FF90FCB8FFA4FCAFE294" box="[819,891,1483,1509]" pageId="16" pageNumber="498">CMK</collectionCode>
).—
<specimenCount id="52E4FD073258FF90FC23FFA4FC11E294" box="[936,965,1483,1509]" pageId="16" pageNumber="498" type="male">1♂</specimenCount>
,
<collectorName id="E91753583258FF90FC59FFA4FBDBE294" box="[978,1039,1483,1509]" pageId="16" pageNumber="498">Agdz</collectorName>
env. [
<geoCoordinate id="21D650493258FF90FBDCFFA4FB68E294" box="[1111,1212,1483,1509]" degrees="30" direction="north" minutes="42" orientation="latitude" pageId="16" pageNumber="498" precision="925" value="30.7">30°42’N</geoCoordinate>
,
<geoCoordinate id="21D650493258FF90FB40FFA5FAFCE294" box="[1227,1320,1482,1509]" degrees="6" direction="west" minutes="27" orientation="longitude" pageId="16" pageNumber="498" precision="925" value="-6.45">6°27’W</geoCoordinate>
],
<quantity id="831A9B6B3258FF90FAB5FFA4FA43E294" box="[1342,1431,1483,1509]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="498" unit="m" value="1000.0">
<elevation id="CFCFD1BD3258FF90FAB5FFA4FA43E294" box="[1342,1431,1483,1509]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="498" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
<date id="305C104E3258FF90FF1CFF81FECDE178" box="[151,281,1518,1545]" pageId="16" pageNumber="498" value="1975-07-14">
<collectingDate id="2018E9A63258FF90FF1CFF81FECDE178" box="[151,281,1518,1545]" pageId="16" pageNumber="498" value="1975-07-14">14.vii.1975</collectingDate>
</date>
,
<collectorName id="E91753583258FF90FEA8FF80FE62E178" box="[291,438,1519,1545]" pageId="16" pageNumber="498">A. W. Ebmer</collectorName>
leg. (coll.
<collectorName id="E91753583258FF90FDA7FF80FD10E17B" box="[556,708,1519,1546]" pageId="16" pageNumber="498">A. W. Ebmer</collectorName>
)
</materialsCitation>
.
</paragraph>
<paragraph id="445D368E3258FF90FF4CFC7DFB9DE104" blockId="16.[151,1437,1374,2013]" pageId="16" pageNumber="498">
<emphasis id="7696EA9C3258FF90FF4CFC7DFDE5E15D" bold="true" box="[199,561,1554,1580]" pageId="16" pageNumber="498">Additional material examined.</emphasis>
<materialsCitation id="F48A3CD33258FF90FDBCFC7CFC73E120" collectingDate="1995-05-21" collectionCode="CMK" collectorName="Beni Bassia &amp; Ma." country="Morocco" latitude="32.516666" location="Bouarfa" longLatPrecision="1210" longitude="-1.9666667" pageId="16" pageNumber="498" specimenCount="1" specimenCount-male="1">
<specimenCount id="52E4FD073258FF90FDBCFC7CFD8FE15C" box="[567,603,1555,1581]" pageId="16" pageNumber="498" type="male">1♂</specimenCount>
,
<collectorName id="E91753583258FF90FDEDFC7CFD3EE15C" box="[614,746,1555,1581]" pageId="16" pageNumber="498">Beni Bassia</collectorName>
<locationDeviation id="55EF2FAC3258FF90FD7BFC7CFC12E15C" box="[752,966,1555,1581]" location="Boudnib" pageId="16" pageNumber="498">
<quantity id="831A9B6B3258FF90FD7BFC7CFCE1E15C" box="[752,821,1555,1581]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="16" pageNumber="498" unit="km" value="60.0">60 km</quantity>
NE Boudnib
</locationDeviation>
[
<geoCoordinate id="21D650493258FF90FC58FC7CFBECE15C" box="[979,1080,1555,1581]" degrees="32" direction="north" minutes="14" orientation="latitude" pageId="16" pageNumber="498" precision="925" value="32.233334">32°14’N</geoCoordinate>
,
<geoCoordinate id="21D650493258FF90FBC9FC7DFB4BE15C" box="[1090,1183,1554,1581]" degrees="3" direction="west" minutes="09" orientation="longitude" pageId="16" pageNumber="498" precision="925" value="-3.15">3°09’W</geoCoordinate>
],
<date id="305C104E3258FF90FB3AFC7DFAF6E15C" box="[1201,1314,1554,1581]" pageId="16" pageNumber="498" value="1995-05-21">
<collectingDate id="2018E9A63258FF90FB3AFC7DFAF6E15C" box="[1201,1314,1554,1581]" pageId="16" pageNumber="498" value="1995-05-21">21.v.1995</collectingDate>
</date>
,
<collectorName id="E91753583258FF90FAA7FC7CFA8DE15C" box="[1324,1369,1555,1581]" pageId="16" pageNumber="498">Ma.</collectorName>
Hala- da leg. (
<collectionCode id="22F3AE4B3258FF90FF78FC58FEE1E120" box="[243,309,1591,1617]" pageId="16" pageNumber="498">CMK</collectionCode>
).—
<collectingCountry id="3CF5761E3258FF90FED5FC58FE27E120" box="[350,499,1591,1617]" name="Morocco" pageId="16" pageNumber="498">MOROCCO</collectingCountry>
:
<quantity id="831A9B6B3258FF90FE74FC58FD93E120" box="[511,583,1591,1617]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="498" unit="km" value="10.0">10 km</quantity>
S
<location id="413D60553258FF90FDECFC58FD15E120" LSID="urn:lsid:plazi:treatment:CC4B87983258FF91FF1CFE9DFC8FE2B0:413D60553258FF90FDECFC58FD15E120" box="[615,705,1591,1617]" country="Morocco" latitude="32.516666" longLatPrecision="1210" longitude="-1.9666667" name="Bouarfa" pageId="16" pageNumber="498">Bouarfa</location>
[
<geoCoordinate id="21D650493258FF90FD5AFC58FCE2E120" box="[721,822,1591,1617]" degrees="32" direction="north" minutes="31" orientation="latitude" pageId="16" pageNumber="498" precision="925" value="32.516666">32°31’N</geoCoordinate>
,
<geoCoordinate id="21D650493258FF90FCC8FC59FC74E120" box="[835,928,1590,1617]" degrees="1" direction="west" minutes="58" orientation="longitude" pageId="16" pageNumber="498" precision="925" value="-1.9666667">1°58’W</geoCoordinate>
]
</materialsCitation>
,
<materialsCitation id="F48A3CD33258FF90FC3FFC59FD9BE104" collectingDate="1995-05-20" collectionCode="OLL" collectorName="Ma. Halada" country="Morocco" elevation="1000" latitude="30.7" longLatPrecision="1220" longitude="-6.45" pageId="16" pageNumber="498" specimenCount="95" specimenCount-female="95">
<specimenCount id="52E4FD073258FF90FC3FFC59FC33E120" box="[948,999,1590,1617]" pageId="16" pageNumber="498" type="female">95♀</specimenCount>
,
<date id="305C104E3258FF90FC7EFC59FBB2E120" box="[1013,1126,1590,1617]" pageId="16" pageNumber="498" value="1995-05-20">
<collectingDate id="2018E9A63258FF90FC7EFC59FBB2E120" box="[1013,1126,1590,1617]" pageId="16" pageNumber="498" value="1995-05-20">20.v.1995</collectingDate>
</date>
,
<collectorName id="E91753583258FF90FBF9FC58FB22E120" box="[1138,1270,1591,1617]" pageId="16" pageNumber="498">Ma. Halada</collectorName>
leg. (
<collectionCode id="22F3AE4B3258FF90FAB3FC58FAA6E120" box="[1336,1394,1591,1617]" pageId="16" pageNumber="498">OLL</collectionCode>
).— Agdz env. [
<geoCoordinate id="21D650493258FF90FE9CFC34FEAFE104" box="[279,379,1627,1653]" degrees="30" direction="north" minutes="42" orientation="latitude" pageId="16" pageNumber="498" precision="925" value="30.7">30°42’N</geoCoordinate>
,
<geoCoordinate id="21D650493258FF90FE03FC35FE31E104" box="[392,485,1626,1653]" degrees="6" direction="west" minutes="27" orientation="longitude" pageId="16" pageNumber="498" precision="925" value="-6.45">6°27’W</geoCoordinate>
],
<quantity id="831A9B6B3258FF90FE73FC34FD9BE104" box="[504,591,1627,1653]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="498" unit="m" value="1000.0">
<elevation id="CFCFD1BD3258FF90FE73FC34FD9BE104" box="[504,591,1627,1653]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="498" unit="m" value="1000.0">1000 m</elevation>
</quantity>
</materialsCitation>
,
<materialsCitation id="F48A3CD33258FF90FDD0FC34FB91E104" box="[603,1093,1626,1653]" collectingDate="1975-07-14" collectionCode="OLL" collectorName="J. Gusenleitner" country="Morocco" pageId="16" pageNumber="498" specimenCount="1" specimenCount-male="1">
<specimenCount id="52E4FD073258FF90FDD0FC34FDABE104" box="[603,639,1627,1653]" pageId="16" pageNumber="498" type="male">1♂</specimenCount>
,
<date id="305C104E3258FF90FD07FC35FCDAE104" box="[652,782,1626,1653]" pageId="16" pageNumber="498" value="1975-07-14">
<collectingDate id="2018E9A63258FF90FD07FC35FCDAE104" box="[652,782,1626,1653]" pageId="16" pageNumber="498" value="1975-07-14">14.vii.1975</collectingDate>
</date>
,
<collectorName id="E91753583258FF90FC92FC34FC15E104" box="[793,961,1627,1653]" pageId="16" pageNumber="498">J. Gusenleitner</collectorName>
leg. (
<collectionCode id="22F3AE4B3258FF90FB8BFC34FBE8E104" box="[1024,1084,1627,1653]" pageId="16" pageNumber="498">OLL</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0CF865053258FF90FF4CFC11FB85E1CC" pageId="16" pageNumber="498" type="etymology">
<paragraph id="445D368E3258FF90FF4CFC11FB85E1CC" blockId="16.[151,1437,1374,2013]" pageId="16" pageNumber="498">
<emphasis id="7696EA9C3258FF90FF4CFC11FE77E1E9" bold="true" box="[199,419,1662,1688]" pageId="16" pageNumber="498">Derivatio nominis.</emphasis>
The epithet is derived from Latin “occidentâlis” (occidentâle as neuter) which means “western” and indicates that this is the westernmost species of the subgenus
<taxonomicName id="83E24D0D3258FF90FC24FCCCFB9FE1CC" authorityName="Pasteels" authorityYear="1969" box="[943,1099,1699,1725]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="498" phylum="Arthropoda" rank="subGenus" subGenus="Gulanthidium">
<emphasis id="7696EA9C3258FF90FC24FCCCFB9FE1CC" box="[943,1099,1699,1725]" italics="true" pageId="16" pageNumber="498">Gulanthidium</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="0CF865053258FF91FF4CFCA9FC81E7A4" lastPageId="17" lastPageNumber="499" pageId="16" pageNumber="498" type="diagnosis">
<paragraph id="445D368E3258FF90FF4CFCA9FC2BE0C8" blockId="16.[151,1437,1374,2013]" pageId="16" pageNumber="498">
<emphasis id="7696EA9C3258FF90FF4CFCA9FE90E191" bold="true" box="[199,324,1734,1760]" pageId="16" pageNumber="498">Diagnosis:</emphasis>
The female is characterised by a narrow impunctate middle line on the clypeus, often with a somewhat scarred surface, which it shares with
<taxonomicName id="83E24D0D3258FF90FDF2FC84FCD2E075" authorityName="Alfken" authorityYear="1938" box="[633,774,1771,1797]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="498" phylum="Arthropoda" rank="species" species="eremicum">
<emphasis id="7696EA9C3258FF90FDF2FC84FCD2E075" box="[633,774,1771,1797]" italics="true" pageId="16" pageNumber="498">A. eremicum</emphasis>
</taxonomicName>
and
<taxonomicName id="83E24D0D3258FF90FCB5FC84FC30E074" authorityName="Kasparek" authorityYear="2021" box="[830,996,1771,1797]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="498" phylum="Arthropoda" rank="species" species="flavissimum">
<emphasis id="7696EA9C3258FF90FCB5FC84FC30E074" box="[830,996,1771,1797]" italics="true" pageId="16" pageNumber="498">A. flavissimum</emphasis>
</taxonomicName>
(
<tableCitation id="096003353258FF90FC78FC84FB9CE074" box="[1011,1096,1771,1797]" captionStart="TABLE 1" captionStartId="7.[151,244,834,860]" captionText="TABLE 1. Comparison of some structural traits in the females of five species Anthidium, subgenus Gulanthidium. The female of A. preoccipitale is not known. Highly relevant distinguishing characters are given in bold." pageId="16" pageNumber="498">Table 1</tableCitation>
). Among these species,
<taxonomicName id="83E24D0D3258FF90FAD7FC84FF3AE059" authorityName="Alfken" authorityYear="1938" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="498" phylum="Arthropoda" rank="species" species="eremicum">
<emphasis id="7696EA9C3258FF90FAD7FC84FF3AE059" italics="true" pageId="16" pageNumber="498">A. eremicum</emphasis>
</taxonomicName>
has a rounded omaulus and bulged lateral T4–T5 (omaulus angulate and T4–T5 toothed in
<taxonomicName id="83E24D0D3258FF90FB67FD60FA58E058" authorityName="Kasparek" authorityYear="2021" box="[1260,1420,1807,1833]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="498" phylum="Arthropoda" rank="species" species="occidentale">
<emphasis id="7696EA9C3258FF90FB67FD60FA58E058" box="[1260,1420,1807,1833]" italics="true" pageId="16" pageNumber="498">A. occidentale</emphasis>
</taxonomicName>
), and
<taxonomicName id="83E24D0D3258FF90FF43FD5CFEBAE03C" authorityName="Kasparek" authorityYear="2021" box="[200,366,1843,1869]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="498" phylum="Arthropoda" rank="species" species="flavissimum">
<emphasis id="7696EA9C3258FF90FF43FD5CFEBAE03C" box="[200,366,1843,1869]" italics="true" pageId="16" pageNumber="498">A. flavissimum</emphasis>
</taxonomicName>
a conspicuously lamellate upper omaulus (angulate in
<taxonomicName id="83E24D0D3258FF90FC6BFD5CFB55E03C" authorityName="Kasparek" authorityYear="2021" box="[992,1153,1843,1869]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="498" phylum="Arthropoda" rank="species" species="occidentale">
<emphasis id="7696EA9C3258FF90FC6BFD5CFB55E03C" box="[992,1153,1843,1869]" italics="true" pageId="16" pageNumber="498">A. occidentale</emphasis>
</taxonomicName>
).
<taxonomicName id="83E24D0D3258FF90FB11FD5CFA48E03C" authorityName="Kasparek" authorityYear="2021" box="[1178,1436,1843,1869]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="498" phylum="Arthropoda" rank="species" species="flavissimum">
<emphasis id="7696EA9C3258FF90FB11FD5CFA48E03C" box="[1178,1436,1843,1869]" italics="true" pageId="16" pageNumber="498">Anthidium flavissimum</emphasis>
</taxonomicName>
is also distinguished by its bright yellow colouration with only a few black remnants in the face and on the scutum (light yellow to reddish-brown with rich black maculation in
<taxonomicName id="83E24D0D3258FF90FCC4FD14FC24E0E4" authorityName="Kasparek" authorityYear="2021" box="[847,1008,1915,1941]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="498" phylum="Arthropoda" rank="species" species="occidentale">
<emphasis id="7696EA9C3258FF90FCC4FD14FC24E0E4" box="[847,1008,1915,1941]" italics="true" pageId="16" pageNumber="498">A. occidentale</emphasis>
</taxonomicName>
) and a broader clypeus (ratio clypeus length/maximal clypeus width&gt;1.4 as compared with &lt;
<quantity id="831A9B6B3258FF90FC83FDF0FC93E0C8" box="[776,839,1951,1977]" metricMagnitude="-2" metricUnit="m" metricValue="3.5559999999999996" pageId="16" pageNumber="498" unit="in" value="1.4">1.4 in</quantity>
<taxonomicName id="83E24D0D3258FF90FCC5FDF0FC3AE0C8" authorityName="Kasparek" authorityYear="2021" box="[846,1006,1951,1977]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="498" phylum="Arthropoda" rank="species" species="occidentale">
<emphasis id="7696EA9C3258FF90FCC5FDF0FC3AE0C8" box="[846,1006,1951,1977]" italics="true" pageId="16" pageNumber="498">A. occidentale</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="445D368E3258FF91FF4CFDACFC81E7A4" blockId="16.[151,1437,1374,2013]" lastBlockId="17.[151,1437,151,1473]" lastPageId="17" lastPageNumber="499" pageId="16" pageNumber="498">
The male is most similar to
<taxonomicName id="83E24D0D3258FF90FE75FDACFD5EE0AD" authorityName="Alfken" authorityYear="1938" box="[510,650,1987,2013]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="498" phylum="Arthropoda" rank="species" species="eremicum">
<emphasis id="7696EA9C3258FF90FE75FDACFD5EE0AD" box="[510,650,1987,2013]" italics="true" pageId="16" pageNumber="498">A. eremicum</emphasis>
</taxonomicName>
and is distinguished by the triangular to drop-shaped form of the apex of the penis valves (in dorsolateral view), which is elongate in
<taxonomicName id="83E24D0D3259FF91FCCCFAF8FC06E7C1" authorityName="Alfken" authorityYear="1938" box="[839,978,151,177]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="499" phylum="Arthropoda" rank="species" species="eremicum">
<emphasis id="7696EA9C3259FF91FCCCFAF8FC06E7C1" box="[839,978,151,177]" italics="true" pageId="17" pageNumber="499">A. eremicum</emphasis>
</taxonomicName>
.
<taxonomicName id="83E24D0D3259FF91FC54FAF8FB07E7C0" authorityName="Kasparek" authorityYear="2021" box="[991,1235,151,177]" class="Magnoliopsida" family="Asteraceae" genus="Anthidum" kingdom="Plantae" order="Asterales" pageId="17" pageNumber="499" phylum="Tracheophyta" rank="species" species="occidentale">
<emphasis id="7696EA9C3259FF91FC54FAF8FB07E7C0" box="[991,1235,151,177]" italics="true" pageId="17" pageNumber="499">Anthidum occidentale</emphasis>
</taxonomicName>
is the only known species of the subgenus
<taxonomicName id="83E24D0D3259FF91FE2DFAD4FD96E7A4" authorityName="Pasteels" authorityYear="1969" box="[422,578,187,213]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="499" phylum="Arthropoda" rank="subGenus" subGenus="Gulanthidium">
<emphasis id="7696EA9C3259FF91FE2DFAD4FD96E7A4" box="[422,578,187,213]" italics="true" pageId="17" pageNumber="499">Gulanthidium</emphasis>
</taxonomicName>
in north-western Africa.
</paragraph>
</subSubSection>
<subSubSection id="0CF865053259FF91FF4CFAB1FE87E27C" pageId="17" pageNumber="499" type="description">
<paragraph id="445D368E3259FF91FF4CFAB1FD18E5D8" blockId="17.[151,1437,151,1473]" pageId="17" pageNumber="499">
<emphasis id="7696EA9C3259FF91FF4CFAB1FE6AE788" bold="true" box="[199,446,222,249]" pageId="17" pageNumber="499">Description. Female.</emphasis>
<quantity id="831A9B6B3259FF91FE4EFAB0FDF1E788" box="[453,549,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="17" pageNumber="499" unit="mm" value="8.5" valueMax="9.0" valueMin="8.0">8–9 mm</quantity>
.
<emphasis id="7696EA9C3259FF91FDA4FAB0FDA1E788" box="[559,629,223,249]" italics="true" pageId="17" pageNumber="499">Head:</emphasis>
Clypeus (
<figureCitation id="DCD92A0B3259FF91FD6CFAB0FC94E788" box="[743,832,223,249]" captionStart="FIGURE 7" captionStartId="10.[151,250,1395,1419]" captionTargetBox="[152,1435,181,1371]" captionTargetId="figure-15@10.[152,1435,181,1371]" captionTargetPageId="10" captionText="FIGURE 7. Faces of the females of Anthidium anguliventre (A), A. eremicum (B), A. occidentale (C), and A. rotundum (D)." figureDoi="http://doi.org/10.5281/zenodo.5531649" httpUri="https://zenodo.org/record/5531649/files/figure.png" pageId="17" pageNumber="499">Fig. 7D</figureCitation>
) light yellow with straight anterior and posterior margins; light brown transparent apex; impunctate middle line approximately half an antennal diameter wide, often somewhat scarred; mandible yellow with five teeth and an additional minute tooth distal to the upper tooth; supraclypeal area yellow and reddish-brown; paraocular area below the antennal socket yellow, tapering above the antennal socket and becoming increasingly reddish-brown; vertex reddish-brown, gena yellow; preoccipital ridge rounded and moderately concave as seen from dorsal (
<figureCitation id="DCD92A0B3259FF91FC9AFBFCFCA9E6DC" box="[785,893,403,429]" captionStart="FIGURE 10" captionStartId="13.[151,250,1274,1298]" captionTargetBox="[284,1304,188,1241]" captionTargetId="figure-39@13.[277,1311,181,1250]" captionTargetPageId="13" captionText="FIGURE 10. Schematic view of the head in the species of Anthidium, subgenus Gulanthidium. A. Lateral view; B. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5531655" httpUri="https://zenodo.org/record/5531655/files/figure.png" pageId="17" pageNumber="499">Fig. 10A</figureCitation>
); antenna reddish-brown.—
<taxonomicName id="83E24D0D3259FF91FB37FBFCFA49E6DC" authority=": Scutum" authorityName="Scutum" box="[1212,1437,403,429]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="17" pageNumber="499" phylum="Arthropoda" rank="genus">
<emphasis id="7696EA9C3259FF91FB37FBFCFAE8E6DD" box="[1212,1340,403,428]" italics="true" pageId="17" pageNumber="499">Mesosoma:</emphasis>
Scutum
</taxonomicName>
black with yellow L-shaped anterolateral band and a yellow longitudinal crescent-shaped stripe next to the middle; scutellum yellow with median posterior impression and narrow transparent posterior margin; base reddish-brown; axilla slightly protruding, yellow and with some reddish-brown on the anterior side; mesepisternum yellow (smooth anterior face reddish-brown and black), omaulus angular.—
<emphasis id="7696EA9C3259FF91FCB6F84CFC66E54D" box="[829,946,547,572]" italics="true" pageId="17" pageNumber="499">Metasoma</emphasis>
: Terga reddish-brown to black, semi-transparent depressions; bands of T1–T6 yellow with reddish-yellow margins; T1–T2 rounded laterally; T3–T5 laterally toothed; T6 depressed in lateral view, light yellow with two dark spots next to the middle.—
<emphasis id="7696EA9C3259FF91FB6FF804FAF7E5F5" box="[1252,1315,619,644]" italics="true" pageId="17" pageNumber="499">Legs:</emphasis>
Hind tibia rounded, hind metatarsus with longitudinal carina.
</paragraph>
<paragraph id="445D368E3259FF91FF4CF8DCFE87E27C" blockId="17.[151,1437,151,1473]" pageId="17" pageNumber="499">
<emphasis id="7696EA9C3259FF91FF4CF8DCFEDFE5BC" bold="true" box="[199,267,691,717]" pageId="17" pageNumber="499">Male.</emphasis>
<quantity id="831A9B6B3259FF91FE9FF8DCFEA2E5BC" box="[276,374,691,717]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="17" pageNumber="499" unit="mm" value="8.5" valueMax="9.0" valueMin="8.0">8–9 mm</quantity>
.
<emphasis id="7696EA9C3259FF91FE09F8DCFE6BE5BC" box="[386,447,691,717]" italics="true" pageId="17" pageNumber="499">Head</emphasis>
: Clypeus bell-shaped with straight anterior margin; lateral margin slightly curved outwards posteriorly; posterior margin semi-transparent, deep yellow and straight; punctation scattered at base and sometimes along middle line but otherwise dense; mandible deep yellow with three strong black teeth; supraclypeal and paraocular area yellow; yellow upper paraocular area confluent with reddish-brown vertex; vertex reddishbrown, sometimes yellow at preoccipital ridge; preoccipital ridge angulate; gena yellow; upper face from posterior ocelli to antennal sockets black (or rarely [one case] reddish-brown), with black maculation rarely extending onto vertex; antenna reddish-brown, lighter on ventral than dorsal surface; ventral surface of scape yellow; entire face including clypeus with long adpressed white hairs.—
<taxonomicName id="83E24D0D3259FF91FD6CF9C0FC10E4B8" authority=": Scutum" authorityName="Scutum" box="[743,964,943,969]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="17" pageNumber="499" phylum="Arthropoda" rank="genus">
<emphasis id="7696EA9C3259FF91FD6CF9C0FC8BE4B9" box="[743,863,943,968]" italics="true" pageId="17" pageNumber="499">Mesosoma</emphasis>
: Scutum
</taxonomicName>
black with broad yellow L-shaped anterolateral band; semi-crescentic band next to the middle, with some variation to length and width (usually merged with the yellow lateral band at notaulus); scutellum widely rounded in dorsal view with median emargination; overhanging propodeum; scutellum and axilla yellow, with base of scutellum reddish-brown; propodeal triangle roughened, punctate posteriorly; mesepisternum yellow with dense white pubescence; pronotal lobe lamellate.—
<emphasis id="7696EA9C3259FF91FAABFE50FA41E329" box="[1312,1429,1087,1112]" italics="true" pageId="17" pageNumber="499">Metasoma</emphasis>
: Terga with yellow bands; depressions semi-transparent, light or dark brown; T3–T6 with acute lateral projection, increasing in size from T3 towards T6; disc of T6 bulging apically next to the centre; T7 semi-circular with some slight flattening at apex (
<figureCitation id="DCD92A0B3259FF91FE32FEC4FDCCE3B4" box="[441,536,1195,1221]" captionStart="FIGURE 8" captionStartId="11.[151,250,945,969]" captionTargetBox="[158,1429,181,920]" captionTargetId="figure-39@11.[158,1429,181,921]" captionTargetPageId="11" captionText="FIGURE 8. Apical terga of the male of five species of Anthidium, subgenus Gulanthidium: A. Anthidium anguliventre; B. A. eremicum; C. A. occidentale sp. nov.; D. A. preoccipitale sp. nov.; E. A. rotundum." figureDoi="http://doi.org/10.5281/zenodo.5531651" httpUri="https://zenodo.org/record/5531651/files/figure.png" pageId="17" pageNumber="499">Figs 8C</figureCitation>
,
<figureCitation id="DCD92A0B3259FF91FDACFEC4FD9EE3B4" box="[551,586,1195,1221]" captionStart="FIGURE 9" captionStartId="11.[151,250,1689,1713]" captionTargetBox="[168,1418,1036,1652]" captionTargetId="figure-81@11.[162,1425,1029,1665]" captionTargetPageId="11" captionText="FIGURE 9. Male of Anthidium, subgenus Gulanthidium. Apical terga (upper row) in dorsal and sternum 6 (S6) in ventral view. A + D. Anthidium anguliventre; B + E. A. eremicum and A. occidentale; C + F. A. preoccipitale and A. rotundum." figureDoi="http://doi.org/10.5281/zenodo.5531653" httpUri="https://zenodo.org/record/5531653/files/figure.png" pageId="17" pageNumber="499">9B</figureCitation>
); sterna light reddish-brown with white pubescence at apical margins; S5 widely emarginate; S6 semicircular, shining, with lateral depression (
<figureCitation id="DCD92A0B3259FF91FC2EFEA0FC2EE398" box="[933,1018,1231,1257]" captionStart="FIGURE 9" captionStartId="11.[151,250,1689,1713]" captionTargetBox="[168,1418,1036,1652]" captionTargetId="figure-81@11.[162,1425,1029,1665]" captionTargetPageId="11" captionText="FIGURE 9. Male of Anthidium, subgenus Gulanthidium. Apical terga (upper row) in dorsal and sternum 6 (S6) in ventral view. A + D. Anthidium anguliventre; B + E. A. eremicum and A. occidentale; C + F. A. preoccipitale and A. rotundum." figureDoi="http://doi.org/10.5281/zenodo.5531653" httpUri="https://zenodo.org/record/5531653/files/figure.png" pageId="17" pageNumber="499">Fig. 9E</figureCitation>
).—
<emphasis id="7696EA9C3259FF91FBAEFEA0FAB8E398" box="[1061,1388,1231,1257]" italics="true" pageId="17" pageNumber="499">Hidden sterna and genitalia:</emphasis>
See separate chapter.
</paragraph>
</subSubSection>
<subSubSection id="0CF865053259FF91FF4CFF79FA5FE227" pageId="17" pageNumber="499" type="distribution">
<paragraph id="445D368E3259FF91FF4CFF79FA5FE227" blockId="17.[151,1437,151,1473]" pageId="17" pageNumber="499">
<emphasis id="7696EA9C3259FF91FF4CFF79FEB5E241" bold="true" box="[199,353,1302,1328]" pageId="17" pageNumber="499">Distribution.</emphasis>
Endemic to
<collectingCountry id="3CF5761E3259FF91FE66FF78FD86E240" box="[493,594,1303,1329]" name="Morocco" pageId="17" pageNumber="499">Morocco</collectingCountry>
(
<figureCitation id="DCD92A0B3259FF91FDEBFF78FD67E240" box="[608,691,1303,1329]" captionStart="FIGURE 12" captionStartId="14.[151,250,1739,1763]" captionTargetBox="[151,1435,1073,1715]" captionTargetId="figure-52@14.[151,1435,1073,1715]" captionTargetPageId="14" captionText="FIGURE 12. Distribution of A. eremicum, A. flavissimum sp. nov., A. occidentale sp. nov., and A. preoccipitale sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5531659" httpUri="https://zenodo.org/record/5531659/files/figure.png" pageId="17" pageNumber="499">Fig. 12</figureCitation>
). Found in the southwestern and eastern High Atlas Mountains and the southern Saharan Atlas Mountains. Elevations varied between
<quantity id="831A9B6B3259FF91FCF5FF54FC11E224" box="[894,965,1339,1365]" metricMagnitude="2" metricUnit="m" metricValue="9.7" pageId="17" pageNumber="499" unit="m" value="970.0">970 m</quantity>
(Beni Bassia) and
<quantity id="831A9B6B3259FF91FB11FF54FB3BE224" box="[1178,1263,1339,1365]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="17" pageNumber="499" unit="m" value="1900.0">1900 m</quantity>
(Tizi-n-Test).
</paragraph>
</subSubSection>
<subSubSection id="0CF865053259FF91FF4CFF30FB93E2EC" pageId="17" pageNumber="499" type="discussion">
<paragraph id="445D368E3259FF91FF4CFF30FB93E2EC" blockId="17.[151,1437,151,1473]" pageId="17" pageNumber="499">
<emphasis id="7696EA9C3259FF91FF4CFF30FEEFE208" bold="true" box="[199,315,1375,1401]" pageId="17" pageNumber="499">Remarks.</emphasis>
It is likely that records listed by van der
<bibRefCitation id="20734B7F3259FF91FD75FF30FC77E20B" author="Zanden, G. van der" box="[766,931,1375,1402]" pageId="17" pageNumber="499" pagination="387 - 390" refId="ref18555" refString="Zanden, G. van der (1996) Neue Verbreitungsangaben zu einigen wenig bekannten palaarktischen Bienen-Arten (Insecta, Hymenoptera, Apoidea). Linzer biologische Beitrage, 28, 387 - 390." type="journal article" year="1996">Zanden (1996)</bibRefCitation>
under
<taxonomicName id="83E24D0D3259FF91FC64FF30FB4EE208" authorityName="Morawitz" authorityYear="1888" box="[1007,1178,1375,1401]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="499" phylum="Arthropoda" rank="species" species="anguliventre">
<emphasis id="7696EA9C3259FF91FC64FF30FB4EE208" box="[1007,1178,1375,1401]" italics="true" pageId="17" pageNumber="499">A. anguliventre</emphasis>
</taxonomicName>
and by Ascher &amp; Pickering (2021) under
<taxonomicName id="83E24D0D3259FF91FEFBFFECFE28E2ED" authorityName="Alfken" authorityYear="1938" box="[368,508,1411,1437]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="499" phylum="Arthropoda" rank="species" species="eremicum">
<emphasis id="7696EA9C3259FF91FEFBFFECFE28E2ED" box="[368,508,1411,1437]" italics="true" pageId="17" pageNumber="499">A. eremicum</emphasis>
</taxonomicName>
(see also
<bibRefCitation id="20734B7F3259FF91FDE7FFECFC8CE2EC" author="Lhomme, P. &amp; Michez, D. &amp; Christmann, S. &amp; Scheuchl, E. &amp; El Abdouni, I. &amp; Hamroud, L. &amp; Ihsane, O. &amp; Sentil, A. &amp; Smaili, M. C. &amp; Schwarz, M. &amp; Dathe, H. &amp; Straka, J. &amp; Pauly, A. &amp; Schmid-Egger, Ch. &amp; Patiny, S. &amp; Muller, A. &amp; Praz, Ch. &amp; Risch, S. &amp; Kasparek, M. &amp; Kuhlmann, M. &amp; Wood, Th. J. &amp; Bogusch, P. &amp; Ascher, J. &amp; Rasmont, R." box="[620,856,1411,1437]" pageId="17" pageNumber="499" pagination="1 - 159" refId="ref17546" refString="Lhomme, P., Michez, D., Christmann, S., Scheuchl, E., El Abdouni, I., Hamroud, L., Ihsane, O., Sentil, A., Smaili, M. C., Schwarz, M., Dathe, H., Straka, J., Pauly, A., Schmid-Egger, Ch. Patiny, S., Muller, A., Praz, Ch., Risch, S., Kasparek, M., Kuhlmann, M., Wood, Th. J., Bogusch, P., Ascher, J. &amp; Rasmont, R. (2020) The wild bees (Hymenoptera: Apoidea) of Morocco. Zootaxa, 4892 (1), 1 - 159. https: // doi. org / 10.11646 / zootaxa. 4892.1.1" type="journal article" year="2020">
Lhomme
<emphasis id="7696EA9C3259FF91FD53FFECFCC4E2EC" box="[728,784,1411,1437]" italics="true" pageId="17" pageNumber="499">et al.</emphasis>
, 2020
</bibRefCitation>
) refer to this species.
</paragraph>
</subSubSection>
<subSubSection id="0CF865053259FF91FF4CFFC9FC8FE2B0" box="[199,859,1446,1473]" pageId="17" pageNumber="499" type="biology_ecology">
<paragraph id="445D368E3259FF91FF4CFFC9FC8FE2B0" blockId="17.[151,1437,151,1473]" box="[199,859,1446,1473]" pageId="17" pageNumber="499">
<emphasis id="7696EA9C3259FF91FF4CFFC9FEF3E2B1" bold="true" box="[199,295,1446,1472]" pageId="17" pageNumber="499">Biology.</emphasis>
All material was collected between May and July.
</paragraph>
</subSubSection>
</treatment>
</document>

253
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<mods:title id="BA9A8733AB69BF8EF3979BBFB5E389AC">Revision and description of three new species of the Palaearctic subgenus Gulanthidium of the wool carder bee genus Anthidium (Apoidea: Megachilidae: Anthidiini)</mods:title>
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<heading id="1F1581E23259FF91FF1CFC7DFDF5E15C" bold="true" box="[151,545,1554,1581]" fontSize="11" level="1" pageId="17" pageNumber="499" reason="1">
<emphasis id="7696EA9C3259FF91FF1CFC7DFDF5E15C" bold="true" box="[151,545,1554,1581]" pageId="17" pageNumber="499">
<taxonomicName id="83E24D0D3259FF91FF1CFC7DFE68E15C" ID-CoL="8SPJQ" authority="Kasparek, 2021" authorityName="Kasparek" authorityYear="2021" box="[151,444,1554,1581]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="499" phylum="Arthropoda" rank="species" species="preoccipitale" status="sp. nov.">
<emphasis id="7696EA9C3259FF91FF1CFC7DFE68E15C" bold="true" box="[151,444,1554,1581]" italics="true" pageId="17" pageNumber="499">Anthidium preoccipitale</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6DA557E73259FF91FE48FC7CFDF5E15C" box="[451,545,1555,1581]" pageId="17" pageNumber="499" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</subSubSection>
<subSubSection id="0CF865053259FF91FF1CFC58FE78E120" box="[151,428,1591,1617]" pageId="17" pageNumber="499" type="description">
<paragraph id="445D368E3259FF91FF1CFC58FE78E120" blockId="17.[151,545,1554,1617]" box="[151,428,1591,1617]" pageId="17" pageNumber="499">
(
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,
<figureCitation id="DCD92A0B3259FF91FE8FFC58FE85E120" box="[260,337,1591,1617]" captionStart="FIGURE 9" captionStartId="11.[151,250,1689,1713]" captionTargetBox="[168,1418,1036,1652]" captionTargetId="figure-81@11.[162,1425,1029,1665]" captionTargetPageId="11" captionText="FIGURE 9. Male of Anthidium, subgenus Gulanthidium. Apical terga (upper row) in dorsal and sternum 6 (S6) in ventral view. A + D. Anthidium anguliventre; B + E. A. eremicum and A. occidentale; C + F. A. preoccipitale and A. rotundum." figureDoi="http://doi.org/10.5281/zenodo.5531653" httpUri="https://zenodo.org/record/5531653/files/figure.png" pageId="17" pageNumber="499">9C, 9F</figureCitation>
,
<figureCitation id="DCD92A0B3259FF91FED7FC58FEAEE120" box="[348,378,1591,1617]" captionStart="FIGURE 10" captionStartId="13.[151,250,1274,1298]" captionTargetBox="[284,1304,188,1241]" captionTargetId="figure-39@13.[277,1311,181,1250]" captionTargetPageId="13" captionText="FIGURE 10. Schematic view of the head in the species of Anthidium, subgenus Gulanthidium. A. Lateral view; B. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5531655" httpUri="https://zenodo.org/record/5531655/files/figure.png" pageId="17" pageNumber="499">10</figureCitation>
,
<figureCitation id="DCD92A0B3259FF91FE0DFC58FE77E120" box="[390,419,1591,1617]" captionStart="FIGURE 14" captionStartId="18.[151,250,1257,1281]" captionTargetBox="[189,1398,632,1234]" captionTargetId="figure-230@18.[189,1398,632,1234]" captionTargetPageId="18" captionText="FIGURE 14. Anthidium preoccipitale sp. nov. Head in frontal and lateral view. Note the lamella on the high preoccipital ridge (A) and the upwards projecting vertex with a sharply angular preoccipital ridge (B)." figureDoi="http://doi.org/10.5281/zenodo.5531663" httpUri="https://zenodo.org/record/5531663/files/figure.png" pageId="17" pageNumber="499">14</figureCitation>
)
</paragraph>
</subSubSection>
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<paragraph id="445D368E3259FF91FF1CFC10FCD6E058" blockId="17.[151,1437,1662,2013]" pageId="17" pageNumber="499">
<emphasis id="7696EA9C3259FF91FF1CFC10FE92E1E9" bold="true" box="[151,326,1662,1689]" pageId="17" pageNumber="499">Type material.</emphasis>
<typeStatus id="9B59882C3259FF91FEC5FC10FE62E1E8" box="[334,438,1663,1689]" pageId="17" pageNumber="499" type="holotype">Holotype</typeStatus>
(male). “Surabad / Typus / Turkestan 189 /
<taxonomicName id="83E24D0D3259FF91FC27FC10FAC5E1E8" authority="F. Mor." authorityName="F. Mor." box="[940,1297,1663,1689]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="499" phylum="Arthropoda" rank="species" species="anguliventre">
<emphasis id="7696EA9C3259FF91FC27FC10FB61E1E8" box="[940,1205,1663,1689]" italics="true" pageId="17" pageNumber="499">Anthidium anguliventre</emphasis>
F. Mor.
</taxonomicName>
/ http://coll. mfn-berlin.de/u” (
<collectionCode id="22F3AE4B3259FF91FEEEFCCCFE73E1CC" box="[357,423,1699,1725]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="17" pageNumber="499" type="Museum">ZMB</collectionCode>
). Note: Surabad is not located in
<collectingCountry id="3CF5761E3259FF91FCB8FCCCFC1AE1CC" box="[819,974,1699,1725]" name="Turkmenistan" pageId="17" pageNumber="499">Turkmenistan</collectingCountry>
as indicated on the label and by
<bibRefCitation id="20734B7F3259FF91FAD3FCCCFF30E190" author="Friese, H." pageId="17" pageNumber="499" refId="ref17078" refString="Friese, H. (1898) Die Bienen Europa ' s (Apidae europaeae) nach ihren Gattungen, Arten und Varietaten auf vergleichend morphologisch-biologischer Grundlage. Theil IV. Solitare Apiden: Genus Eriades. Genus Trachusa. Genus Anthidium. R. Friedla ¨ nder, Berlin, 303 pp." type="book" year="1898">Friese (1898)</bibRefCitation>
, but in
<collectingCountry id="3CF5761E3259FF91FEBDFCA8FEB7E190" box="[310,355,1735,1761]" name="Iran" pageId="17" pageNumber="499">Iran</collectingCountry>
as found out by
<bibRefCitation id="20734B7F3259FF91FD96FCA8FD65E190" author="Baker, D. B." box="[541,689,1735,1761]" pageId="17" pageNumber="499" pagination="823 - 860" refId="ref16669" refString="Baker, D. B. (1998) Taxonomic and phylogenetic problems in Old World eucerine bees, with special reference to the genus Tarsalia Morawitz, 1895 (Hymenoptera: Apoidea: Anthophoridae). Journal of Natural History, 32, 823 - 860. https: // doi. org / 10.1080 / 00222939800770441" type="journal article" year="1998">Baker (1998)</bibRefCitation>
based on the travel itinerary of the collector, D. K. Glasonov [Glazunov]. Coordinates: 35°
<geoCoordinate id="21D650493259FF91FE24FC85FE39E074" box="[431,493,1770,1797]" degrees="43" direction="north" orientation="latitude" pageId="17" pageNumber="499" precision="55555" value="43.0">43°N</geoCoordinate>
,
<geoCoordinate id="21D650493259FF91FE73FC85FD8CE074" box="[504,600,1770,1797]" degrees="61" direction="east" minutes="05" orientation="longitude" pageId="17" pageNumber="499" precision="925" value="61.083332">61°05’E</geoCoordinate>
. Genitalia are dissected (by F. Friese?) and available, but their preservation status does not allow examination of the relevant traits.
</paragraph>
</subSubSection>
<subSubSection id="0CF865053259FF91FF4CFD5DFE3CE0C8" pageId="17" pageNumber="499" type="diagnosis">
<paragraph id="445D368E3259FF91FF4CFD5DFE3CE0C8" blockId="17.[151,1437,1662,2013]" pageId="17" pageNumber="499">
<emphasis id="7696EA9C3259FF91FF4CFD5DFE96E03D" bold="true" box="[199,322,1842,1868]" pageId="17" pageNumber="499">Diagnosis.</emphasis>
The most similar species is
<taxonomicName id="83E24D0D3259FF91FDF6FD5CFCDCE03C" authorityName="Warncke" authorityYear="1980" box="[637,776,1843,1869]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="499" phylum="Arthropoda" rank="species" species="rotundum">
<emphasis id="7696EA9C3259FF91FDF6FD5CFCDCE03C" box="[637,776,1843,1869]" italics="true" pageId="17" pageNumber="499">A. rotundum</emphasis>
</taxonomicName>
with which the male shares particularly the median furrow on S6. However, it is distinguished from that species and all other members of the subgenus by a strong lamella on the preoccipital ridge. Punctation on tergal discs more scattered than in
<taxonomicName id="83E24D0D3259FF91FC4FFD14FB84E0E4" authorityName="Warncke" authorityYear="1980" box="[964,1104,1915,1941]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="499" phylum="Arthropoda" rank="species" species="rotundum">
<emphasis id="7696EA9C3259FF91FC4FFD14FB84E0E4" box="[964,1104,1915,1941]" italics="true" pageId="17" pageNumber="499">A. rotundum</emphasis>
</taxonomicName>
. T2 laterally rounded (lateral tooth present in
<taxonomicName id="83E24D0D3259FF91FEC7FDF0FE03E0C8" authorityName="Warncke" authorityYear="1980" box="[332,471,1951,1977]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="499" phylum="Arthropoda" rank="species" species="rotundum">
<emphasis id="7696EA9C3259FF91FEC7FDF0FE03E0C8" box="[332,471,1951,1977]" italics="true" pageId="17" pageNumber="499">A. rotundum</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="0CF865053259FF92FF4CFDADFDCEE137" lastPageId="18" lastPageNumber="500" pageId="17" pageNumber="499" type="description">
<paragraph id="445D368E3259FF92FF4CFDADFBD5E511" blockId="17.[151,1437,1662,2013]" lastBlockId="18.[151,1437,150,609]" lastPageId="18" lastPageNumber="500" pageId="17" pageNumber="499">
<emphasis id="7696EA9C3259FF91FF4CFDADFE70E0AC" bold="true" box="[199,420,1986,2013]" pageId="17" pageNumber="499">Description. Male.</emphasis>
<quantity id="831A9B6B3259FF91FE20FDACFE29E0AC" box="[427,509,1987,2013]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="499" unit="mm" value="10.0">10 mm</quantity>
.
<emphasis id="7696EA9C3259FF91FD83FDACFD91E0AC" box="[520,581,1987,2013]" italics="true" pageId="17" pageNumber="499">Head</emphasis>
: Clypeus flat, anterior margin straight, posterior margin shallowly emarginate; apical margin brown, smooth; median third of clypeus almost impunctate; mandible yellow with three strong teeth; supraclypeal and lower paraocular area yellow, upper paraocular area with yellow band along inner orbit of eye and confluent with yellow maculation on vertex; vertex with broad ochreous to yellow transverse band, confluent with yellow gena; strong lamella (approx.
<quantity id="831A9B6B325AFF92FD62FB6DFC9AE66C" box="[745,846,258,285]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="18" pageNumber="500" unit="mm" value="0.13">0.13 mm</quantity>
high) extending to height of middle of eye; antenna ochreous; underside of scape yellow.—
<taxonomicName id="83E24D0D325AFF92FDDAFB48FCFAE631" authority=": Scutum" authorityName="Scutum" box="[593,814,294,321]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="18" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="7696EA9C325AFF92FDDAFB48FD1DE631" box="[593,713,295,320]" italics="true" pageId="18" pageNumber="500">Mesosoma</emphasis>
: Scutum
</taxonomicName>
black, densely punctated; broad L-shaped anterolateral yellow band; remnants of a thin yellow longitudinal line next to the middle; scutellum and axillae yellow, scutellum with some black colouration anteriorly and with median emargination; axilla slightly protruding laterally; propodeal triangle rugulose, coarser at base than apically; upper half of mesepisternum yellow, lower half black; pronotal lobe lamellate.—
<emphasis id="7696EA9C325AFF92FED2FBD8FE1AE6A1" box="[345,462,439,464]" italics="true" pageId="18" pageNumber="500">Metasoma</emphasis>
: Tergal discs punctate, with distance between punctures up to 1–2 puncture diameter; depressions T1–T6 chestnut-brown; T3–T6 each with a lateral tooth, small in T3 and becoming larger towards T6; teeth flat with transparent lamella-like margin; disc of T6 bulging on each side of the middle; T7 semicircular with transparent margin (
<figureCitation id="DCD92A0B325AFF92FEFCF84DFE07E54D" box="[375,467,546,572]" captionStart="FIGURE 8" captionStartId="11.[151,250,945,969]" captionTargetBox="[158,1429,181,920]" captionTargetId="figure-39@11.[158,1429,181,921]" captionTargetPageId="11" captionText="FIGURE 8. Apical terga of the male of five species of Anthidium, subgenus Gulanthidium: A. Anthidium anguliventre; B. A. eremicum; C. A. occidentale sp. nov.; D. A. preoccipitale sp. nov.; E. A. rotundum." figureDoi="http://doi.org/10.5281/zenodo.5531651" httpUri="https://zenodo.org/record/5531651/files/figure.png" pageId="18" pageNumber="500">Figs 8D</figureCitation>
,
<figureCitation id="DCD92A0B325AFF92FE55F84DFDD6E54D" box="[478,514,546,572]" captionStart="FIGURE 9" captionStartId="11.[151,250,1689,1713]" captionTargetBox="[168,1418,1036,1652]" captionTargetId="figure-81@11.[162,1425,1029,1665]" captionTargetPageId="11" captionText="FIGURE 9. Male of Anthidium, subgenus Gulanthidium. Apical terga (upper row) in dorsal and sternum 6 (S6) in ventral view. A + D. Anthidium anguliventre; B + E. A. eremicum and A. occidentale; C + F. A. preoccipitale and A. rotundum." figureDoi="http://doi.org/10.5281/zenodo.5531653" httpUri="https://zenodo.org/record/5531653/files/figure.png" pageId="18" pageNumber="500">9C</figureCitation>
); S6 semicircular with median furrow and lateral depression (
<figureCitation id="DCD92A0B325AFF92FB25F84DFAD4E54D" box="[1198,1280,546,572]" captionStart="FIGURE 9" captionStartId="11.[151,250,1689,1713]" captionTargetBox="[168,1418,1036,1652]" captionTargetId="figure-81@11.[162,1425,1029,1665]" captionTargetPageId="11" captionText="FIGURE 9. Male of Anthidium, subgenus Gulanthidium. Apical terga (upper row) in dorsal and sternum 6 (S6) in ventral view. A + D. Anthidium anguliventre; B + E. A. eremicum and A. occidentale; C + F. A. preoccipitale and A. rotundum." figureDoi="http://doi.org/10.5281/zenodo.5531653" httpUri="https://zenodo.org/record/5531653/files/figure.png" pageId="18" pageNumber="500">Fig. 9F</figureCitation>
).—
<emphasis id="7696EA9C325AFF92FAA0F84CFAB4E54D" box="[1323,1376,547,572]" italics="true" pageId="18" pageNumber="500">Legs</emphasis>
: Yellow, inner faces black or dark brown; hind basitarsus with longitudinal carina.
</paragraph>
<caption id="109D6606325AFF92FF1CFE86FC3CE254" ID-DOI="http://doi.org/10.5281/zenodo.5531663" ID-Zenodo-Dep="5531663" httpUri="https://zenodo.org/record/5531663/files/figure.png" pageId="18" pageNumber="500" startId="18.[151,250,1257,1281]" targetBox="[189,1398,632,1234]" targetPageId="18">
<paragraph id="445D368E325AFF92FF1CFE86FC3CE254" blockId="18.[151,1436,1257,1318]" pageId="18" pageNumber="500">
<emphasis id="7696EA9C325AFF92FF1CFE86FEF4E270" bold="true" box="[151,288,1257,1281]" pageId="18" pageNumber="500">FIGURE 14.</emphasis>
<taxonomicName id="83E24D0D325AFF92FEADFE85FDCFE273" authority="Kasparek, 2021" authorityName="Kasparek" authorityYear="2021" box="[294,539,1258,1282]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="500" phylum="Arthropoda" rank="species" species="preoccipitale" status="sp. nov.">
<emphasis id="7696EA9C325AFF92FEADFE85FDCFE273" box="[294,539,1258,1282]" italics="true" pageId="18" pageNumber="500">Anthidium preoccipitale</emphasis>
</taxonomicName>
<emphasis id="7696EA9C325AFF92FDAAFE85FDA5E273" bold="true" box="[545,625,1258,1282]" pageId="18" pageNumber="500">
<taxonomicNameLabel id="6DA557E7325AFF92FDAAFE85FDA5E273" box="[545,625,1258,1282]" pageId="18" pageNumber="500" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Head in frontal and lateral view. Note the lamella on the high preoccipital ridge
<emphasis id="7696EA9C325AFF92FF1CFF62FF6FE254" bold="true" box="[151,187,1293,1317]" pageId="18" pageNumber="500">(A)</emphasis>
and the upwards projecting vertex with a sharply angular preoccipital ridge
<emphasis id="7696EA9C325AFF92FC4BFF62FC36E254" bold="true" box="[960,994,1293,1317]" pageId="18" pageNumber="500">(B)</emphasis>
.
</paragraph>
</caption>
<paragraph id="445D368E325AFF92FF4CFF3BFDCEE137" blockId="18.[151,1437,1364,2038]" pageId="18" pageNumber="500">
<emphasis id="7696EA9C325AFF92FF4CFF3BFEA5E21F" bold="true" box="[199,369,1364,1390]" pageId="18" pageNumber="500">Morphometry</emphasis>
. Some measurements of the male
<typeStatus id="9B59882C325AFF92FC8FFF3BFCB2E21F" box="[772,870,1364,1390]" pageId="18" pageNumber="500" type="holotype">holotype</typeStatus>
of
<taxonomicName id="83E24D0D325AFF92FC19FF3BFB93E21F" authorityName="Kasparek" authorityYear="2021" box="[914,1095,1364,1390]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="500" phylum="Arthropoda" rank="species" species="preoccipitale">
<emphasis id="7696EA9C325AFF92FC19FF3BFB93E21F" box="[914,1095,1364,1390]" italics="true" pageId="18" pageNumber="500">A. preoccipitale</emphasis>
</taxonomicName>
were compared with a series (N=14) of males
<taxonomicName id="83E24D0D325AFF92FEDDFF17FE35E2E3" authorityName="Warncke" authorityYear="1980" box="[342,481,1400,1426]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="500" phylum="Arthropoda" rank="species" species="rotundum">
<emphasis id="7696EA9C325AFF92FEDDFF17FE35E2E3" box="[342,481,1400,1426]" italics="true" pageId="18" pageNumber="500">A. rotundum</emphasis>
</taxonomicName>
(see under “material examined” there): the ocellocular distance, the hind ocellus-vertex distance, the distance between the posterior ocelli, and the distance between anterior and posterior ocelli. None of these measurements alone was strong enough to distinguish
<taxonomicName id="83E24D0D325AFF92FCE0FFAFFBCBE2AB" authorityName="Kasparek" authorityYear="2021" box="[875,1055,1472,1498]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="500" phylum="Arthropoda" rank="species" species="preoccipitale">
<emphasis id="7696EA9C325AFF92FCE0FFAFFBCBE2AB" box="[875,1055,1472,1498]" italics="true" pageId="18" pageNumber="500">A. preoccipitale</emphasis>
</taxonomicName>
from
<taxonomicName id="83E24D0D325AFF92FBECFFAFFB21E2AB" authorityName="Warncke" authorityYear="1980" box="[1127,1269,1472,1498]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="500" phylum="Arthropoda" rank="species" species="rotundum">
<emphasis id="7696EA9C325AFF92FBECFFAFFB21E2AB" box="[1127,1269,1472,1498]" italics="true" pageId="18" pageNumber="500">A. rotundum</emphasis>
</taxonomicName>
. However, the combination of these five parameters showed in a Discriminant Function Analysis (DFA) that the discriminant score of
<taxonomicName id="83E24D0D325AFF92FF3DFC67FEBCE153" authorityName="Kasparek" authorityYear="2021" box="[182,360,1544,1570]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="500" phylum="Arthropoda" rank="species" species="preoccipitale">
<emphasis id="7696EA9C325AFF92FF3DFC67FEBCE153" box="[182,360,1544,1570]" italics="true" pageId="18" pageNumber="500">A. preoccipitale</emphasis>
</taxonomicName>
is higher than in
<taxonomicName id="83E24D0D325AFF92FDA0FC67FD63E153" authorityName="Warncke" authorityYear="1980" box="[555,695,1544,1570]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="500" phylum="Arthropoda" rank="species" species="rotundum">
<emphasis id="7696EA9C325AFF92FDA0FC67FD63E153" box="[555,695,1544,1570]" italics="true" pageId="18" pageNumber="500">A. rotundum</emphasis>
</taxonomicName>
(
<figureCitation id="DCD92A0B325AFF92FD4DFC67FCCCE152" box="[710,792,1544,1571]" captionStart="FIGURE 15" captionStartId="19.[151,250,853,877]" captionTargetBox="[160,1428,188,822]" captionTargetId="figure-39@19.[151,1436,181,829]" captionTargetPageId="19" captionText="FIGURE 15. Comparison of some morphometric parameters regarding the distance of the ocelli from the eye and the preoccipital ridge (four parameters) between male Anthidium rotundum (grey) (N=14) and A. preoccipitale sp. nov. (scarlet) (N=1), based on a Discriminant Function Analysis." figureDoi="http://doi.org/10.5281/zenodo.5531665" httpUri="https://zenodo.org/record/5531665/files/figure.png" pageId="18" pageNumber="500">Fig. 15</figureCitation>
), and this may be understood as further indication for the distinctiveness of
<taxonomicName id="83E24D0D325AFF92FEEAFC43FDC7E137" authorityName="Kasparek" authorityYear="2021" box="[353,531,1580,1606]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="500" phylum="Arthropoda" rank="species" species="preoccipitale">
<emphasis id="7696EA9C325AFF92FEEAFC43FDC7E137" box="[353,531,1580,1606]" italics="true" pageId="18" pageNumber="500">A. preoccipitale</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="0CF86505325AFF92FF4CFC20FB1AE11B" box="[199,1230,1615,1642]" pageId="18" pageNumber="500" type="etymology">
<paragraph id="445D368E325AFF92FF4CFC20FB1AE11B" blockId="18.[151,1437,1364,2038]" box="[199,1230,1615,1642]" pageId="18" pageNumber="500">
<emphasis id="7696EA9C325AFF92FF4CFC20FE70E118" bold="true" box="[199,420,1615,1641]" pageId="18" pageNumber="500">Derivatio nominis.</emphasis>
The name refers to the preoccipital ridge which bears a distinct lamella.
</paragraph>
</subSubSection>
<subSubSection id="0CF86505325AFF92FF4CFC1CFE70E1FF" box="[199,420,1651,1678]" pageId="18" pageNumber="500" type="biology_ecology">
<paragraph id="445D368E325AFF92FF4CFC1CFE70E1FF" blockId="18.[151,1437,1364,2038]" box="[199,420,1651,1678]" pageId="18" pageNumber="500">
<emphasis id="7696EA9C325AFF92FF4CFC1CFEF3E1FC" bold="true" box="[199,295,1651,1677]" pageId="18" pageNumber="500">Biology.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="0CF86505325AFF92FF4CFCF8FB7EE1C3" box="[199,1194,1687,1714]" pageId="18" pageNumber="500" type="distribution">
<paragraph id="445D368E325AFF92FF4CFCF8FB7EE1C3" blockId="18.[151,1437,1364,2038]" box="[199,1194,1687,1714]" pageId="18" pageNumber="500">
<emphasis id="7696EA9C325AFF92FF4CFCF8FEB5E1C0" bold="true" box="[199,353,1687,1713]" pageId="18" pageNumber="500">Distribution.</emphasis>
Only known from the
<typeStatus id="9B59882C325AFF92FDEBFCF7FD44E1C3" box="[608,656,1688,1714]" pageId="18" pageNumber="500">type</typeStatus>
locality, situated in north-eastern
<collectingCountry id="3CF5761E325AFF92FB86FCF7FBEEE1C3" box="[1037,1082,1688,1714]" name="Iran" pageId="18" pageNumber="500">Iran</collectingCountry>
(
<figureCitation id="DCD92A0B325AFF92FBC1FCF7FB49E1C3" box="[1098,1181,1688,1714]" captionStart="FIGURE 12" captionStartId="14.[151,250,1739,1763]" captionTargetBox="[151,1435,1073,1715]" captionTargetId="figure-52@14.[151,1435,1073,1715]" captionTargetPageId="14" captionText="FIGURE 12. Distribution of A. eremicum, A. flavissimum sp. nov., A. occidentale sp. nov., and A. preoccipitale sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5531659" httpUri="https://zenodo.org/record/5531659/files/figure.png" pageId="18" pageNumber="500">Fig. 12</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0CF86505325AFF92FF4CFCD3FE2CE087" pageId="18" pageNumber="500" type="discussion">
<paragraph id="445D368E325AFF92FF4CFCD3FE2CE087" blockId="18.[151,1437,1364,2038]" pageId="18" pageNumber="500">
<emphasis id="7696EA9C325AFF92FF4CFCD3FEEFE1A7" bold="true" box="[199,315,1724,1750]" pageId="18" pageNumber="500">Remarks.</emphasis>
The
<typeStatus id="9B59882C325AFF92FEFDFCD3FE0CE1A7" box="[374,472,1724,1750]" pageId="18" pageNumber="500" type="holotype">holotype</typeStatus>
of
<taxonomicName id="83E24D0D325AFF92FE75FCD3FD65E1A7" authorityName="Kasparek" authorityYear="2021" box="[510,689,1724,1750]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="500" phylum="Arthropoda" rank="species" species="preoccipitale">
<emphasis id="7696EA9C325AFF92FE75FCD3FD65E1A7" box="[510,689,1724,1750]" italics="true" pageId="18" pageNumber="500">A. preoccipitale</emphasis>
</taxonomicName>
male was provided to H. Friese together with a female
<taxonomicName id="83E24D0D325AFF92FAA3FCD3FA49E1A7" authorityName="Fabricius" authorityYear="1804" box="[1320,1437,1724,1750]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="7696EA9C325AFF92FAA3FCD3FA49E1A7" box="[1320,1437,1724,1750]" italics="true" pageId="18" pageNumber="500">Anthidium</emphasis>
</taxonomicName>
from the same location, and both specimens were regarded by him as
<typeStatus id="9B59882C325AFF92FC39FC8FFBC9E18B" box="[946,1053,1760,1786]" pageId="18" pageNumber="500" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="83E24D0D325AFF92FBCFFC8FFB25E18B" box="[1092,1265,1760,1786]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="500" phylum="Arthropoda" rank="species" species="anguliventre">
<emphasis id="7696EA9C325AFF92FBCFFC8FFB25E18B" box="[1092,1265,1760,1786]" italics="true" pageId="18" pageNumber="500">A. anguliventre</emphasis>
</taxonomicName>
. As the collection date and other details are not available, it cannot be said whether these
<specimenCount id="52E4FD07325AFF92FC67FD6BFB46E06F" box="[1004,1170,1796,1822]" pageId="18" pageNumber="500" type="generic">two specimens</specimenCount>
were actually collected together. The identity of the female as
<taxonomicName id="83E24D0D325AFF92FDC5FD47FD2EE033" box="[590,762,1832,1858]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="500" phylum="Arthropoda" rank="species" species="anguliventre">
<emphasis id="7696EA9C325AFF92FDC5FD47FD2EE033" box="[590,762,1832,1858]" italics="true" pageId="18" pageNumber="500">A. anguliventre</emphasis>
</taxonomicName>
was confirmed here by a flat clypeus with a medially wide impunctate middle area, the punctation of T5 reaching the posterior margin, the laterally rounded T1–T2, and the presence of blunt teeth on T3–T5. In a Discriminant Function Analysis of six morphometric parameters of the clypeus, the female specimen also clearly clustered with
<taxonomicName id="83E24D0D325AFF92FD67FDFBFC4CE0DF" box="[748,920,1940,1966]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="500" phylum="Arthropoda" rank="species" species="anguliventre">
<emphasis id="7696EA9C325AFF92FD67FDFBFC4CE0DF" box="[748,920,1940,1966]" italics="true" pageId="18" pageNumber="500">A. anguliventre</emphasis>
</taxonomicName>
. Nevertheless, it cannot be ruled out that this female is the unknown female of
<taxonomicName id="83E24D0D325AFF92FD9EFDD7FD1CE0A3" authorityName="Kasparek" authorityYear="2021" box="[533,712,1976,2002]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="500" phylum="Arthropoda" rank="species" species="preoccipitale">
<emphasis id="7696EA9C325AFF92FD9EFDD7FD1CE0A3" box="[533,712,1976,2002]" italics="true" pageId="18" pageNumber="500">A. preoccipitale</emphasis>
</taxonomicName>
which would then be indistinguishable from
<taxonomicName id="83E24D0D325AFF92FB40FDD7FAACE0A3" box="[1227,1400,1976,2002]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="500" phylum="Arthropoda" rank="species" species="anguliventre">
<emphasis id="7696EA9C325AFF92FB40FDD7FAACE0A3" box="[1227,1400,1976,2002]" italics="true" pageId="18" pageNumber="500">A. anguliventre</emphasis>
</taxonomicName>
on the basis of external characters.
</paragraph>
</subSubSection>
</treatment>
</document>

894
data/CC/4B/87/CC4B8798325BFF95FF1CFE69FE3FE614.xml Normal file
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@ -0,0 +1,894 @@
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<mods:title id="2EFA7A9A1AF8053EC26CAD3D806BEED8">Revision and description of three new species of the Palaearctic subgenus Gulanthidium of the wool carder bee genus Anthidium (Apoidea: Megachilidae: Anthidiini)</mods:title>
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<paragraph id="445D368E325BFF93FF1CFE69FD88E350" blockId="19.[151,608,1030,1094]" box="[151,604,1030,1057]" pageId="19" pageNumber="501">
<heading id="1F1581E2325BFF93FF1CFE69FD88E350" bold="true" box="[151,604,1030,1057]" fontSize="11" level="1" pageId="19" pageNumber="501" reason="1">
<taxonomicName id="83E24D0D325BFF93FF1CFE69FD88E350" ID-CoL="5VCJD" authority="Warncke, 1980" authorityName="Warncke" authorityYear="1980" box="[151,604,1030,1057]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="501" phylum="Arthropoda" rank="species" species="rotundum">
<emphasis id="7696EA9C325BFF93FF1CFE69FD88E350" bold="true" box="[151,604,1030,1057]" pageId="19" pageNumber="501">
<emphasis id="7696EA9C325BFF93FF1CFE69FE4EE350" bold="true" box="[151,410,1030,1057]" italics="true" pageId="19" pageNumber="501">Anthidium rotundum</emphasis>
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</emphasis>
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</heading>
</paragraph>
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<subSubSection id="0CF86505325BFF93FF1CFE44FDB4E334" box="[151,608,1067,1094]" pageId="19" pageNumber="501" type="description">
<paragraph id="445D368E325BFF93FF1CFE44FDB4E334" blockId="19.[151,608,1030,1094]" box="[151,608,1067,1094]" pageId="19" pageNumber="501">
(
<figureCitation id="DCD92A0B325BFF93FF14FE44FF32E334" box="[159,230,1067,1093]" captionStart="FIGURE 1" captionStartId="2.[151,250,1927,1951]" captionTargetBox="[162,1427,568,1891]" captionTargetId="figure-200@2.[151,1436,560,1903]" captionTargetPageId="2" captionText="FIGURE 1. Comparison of the hidden sterna S7 (A) and S8 (B) and the apices of the penis valves in dorsolateral view (C) in four species of Anthidium, subgenus Gulanthidium." figureDoi="http://doi.org/10.5281/zenodo.5531633" httpUri="https://zenodo.org/record/5531633/files/figure.png" pageId="19" pageNumber="501">Figs 1</figureCitation>
,
<figureCitation id="DCD92A0B325BFF93FF7FFE44FECCE334" box="[244,280,1067,1093]" captionStart="FIGURE 2" captionStartId="3.[151,250,1432,1456]" captionTargetBox="[166,1421,181,1409]" captionTargetId="figure-39@3.[166,1421,181,1409]" captionTargetPageId="3" captionText="FIGURE 2. Male genitalia of four species of Anthidium (Gulanthidium) in dorsal view: A. A. anguliventre; B. A. eremicum; C. A. occidentale sp. nov.; D. A. rotundum." figureDoi="http://doi.org/10.5281/zenodo.5531635" httpUri="https://zenodo.org/record/5531635/files/figure.png" pageId="19" pageNumber="501">2D</figureCitation>
,
<figureCitation id="DCD92A0B325BFF93FEAFFE44FEA2E337" box="[292,374,1067,1094]" captionStart="FIGURE 6" captionStartId="9.[151,250,1360,1384]" captionTargetBox="[155,1432,181,1337]" captionTargetId="figure-39@9.[155,1432,181,1337]" captionTargetPageId="9" captionText="FIGURE 6. Habitus dorsal of four species of Anthidium, subgenus Gulanthidium. Upper row females, lower row males. A + E. Anthidium anguliventre; B + F. A. eremicum; C + G. A. occidentale sp. nov.; D + H. A. rotundum." figureDoi="http://doi.org/10.5281/zenodo.5531647" httpUri="https://zenodo.org/record/5531647/files/figure.png" pageId="19" pageNumber="501">6D, 6H</figureCitation>
,
<figureCitation id="DCD92A0B325BFF93FE0FFE43FE7DE337" box="[388,425,1068,1094]" captionStart="FIGURE 7" captionStartId="10.[151,250,1395,1419]" captionTargetBox="[152,1435,181,1371]" captionTargetId="figure-15@10.[152,1435,181,1371]" captionTargetPageId="10" captionText="FIGURE 7. Faces of the females of Anthidium anguliventre (A), A. eremicum (B), A. occidentale (C), and A. rotundum (D)." figureDoi="http://doi.org/10.5281/zenodo.5531649" httpUri="https://zenodo.org/record/5531649/files/figure.png" pageId="19" pageNumber="501">7D</figureCitation>
,
<figureCitation id="DCD92A0B325BFF93FE3EFE44FE02E334" box="[437,470,1067,1093]" captionStart="FIGURE 8" captionStartId="11.[151,250,945,969]" captionTargetBox="[158,1429,181,920]" captionTargetId="figure-39@11.[158,1429,181,921]" captionTargetPageId="11" captionText="FIGURE 8. Apical terga of the male of five species of Anthidium, subgenus Gulanthidium: A. Anthidium anguliventre; B. A. eremicum; C. A. occidentale sp. nov.; D. A. preoccipitale sp. nov.; E. A. rotundum." figureDoi="http://doi.org/10.5281/zenodo.5531651" httpUri="https://zenodo.org/record/5531651/files/figure.png" pageId="19" pageNumber="501">8E</figureCitation>
,
<figureCitation id="DCD92A0B325BFF93FE69FE44FDFBE334" box="[482,559,1067,1093]" captionStart="FIGURE 9" captionStartId="11.[151,250,1689,1713]" captionTargetBox="[168,1418,1036,1652]" captionTargetId="figure-81@11.[162,1425,1029,1665]" captionTargetPageId="11" captionText="FIGURE 9. Male of Anthidium, subgenus Gulanthidium. Apical terga (upper row) in dorsal and sternum 6 (S6) in ventral view. A + D. Anthidium anguliventre; B + E. A. eremicum and A. occidentale; C + F. A. preoccipitale and A. rotundum." figureDoi="http://doi.org/10.5281/zenodo.5531653" httpUri="https://zenodo.org/record/5531653/files/figure.png" pageId="19" pageNumber="501">9C, 9F</figureCitation>
,
<figureCitation id="DCD92A0B325BFF93FDB1FE44FD83E334" box="[570,599,1067,1093]" captionStart="FIGURE 10" captionStartId="13.[151,250,1274,1298]" captionTargetBox="[284,1304,188,1241]" captionTargetId="figure-39@13.[277,1311,181,1250]" captionTargetPageId="13" captionText="FIGURE 10. Schematic view of the head in the species of Anthidium, subgenus Gulanthidium. A. Lateral view; B. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5531655" httpUri="https://zenodo.org/record/5531655/files/figure.png" pageId="19" pageNumber="501">10</figureCitation>
)
</paragraph>
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<treatmentCitationGroup id="64F211A0325BFF93FF1CFE00FD38E3F6" box="[151,748,1135,1160]" pageId="19" pageNumber="501">
<treatmentCitation id="C543109F325BFF93FF1CFE00FDB9E3F6" author="Warncke, K." box="[151,621,1135,1160]" page="187" pageId="19" pageNumber="501" year="1980">
<taxonomicName id="83E24D0D325BFF93FF1CFE00FDB9E3F6" ID-CoL="5VCJD" authority="Warncke, 1980: 186 - 187" authorityName="Warncke" authorityPageNumber="186 - 187" authorityYear="1980" box="[151,621,1135,1160]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="501" phylum="Arthropoda" rank="species" species="rotundum">
<emphasis id="7696EA9C325BFF93FF1CFE00FEBDE3F6" box="[151,361,1135,1159]" italics="true" pageId="19" pageNumber="501">Anthidium rotundum</emphasis>
<bibRefCitation id="20734B7F325BFF93FEE4FE00FDB9E3F6" author="Warncke, K." box="[367,621,1135,1160]" pageId="19" pageNumber="501" pagination="119 - 210" refId="ref18439" refString="Warncke, K. (1980) Die Bienengattung Anthidium Fabricius, 1804 in der Westpalaarktis und im turkestanischen Becken. Entomofauna, 1, 119 - 210." type="journal article" year="1980">Warncke, 1980: 186–187</bibRefCitation>
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(
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; male)
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</paragraph>
</subSubSection>
<subSubSection id="0CF86505325BFF94FF1CFED9FACCE510" lastPageId="20" lastPageNumber="502" pageId="19" pageNumber="501" type="materials_examined">
<paragraph id="445D368E325BFF93FF1CFED9FE85E211" blockId="19.[151,1437,1205,2024]" pageId="19" pageNumber="501">
<emphasis id="7696EA9C325BFF93FF1CFED9FE14E3BE" bold="true" box="[151,448,1205,1232]" pageId="19" pageNumber="501">Type material examined.</emphasis>
<materialsCitation id="F48A3CD3325BFF93FE4CFED9FDD7E385" collectingDate="1965-07-09" collectingDateMax="1965-07-10" collectingDateMin="1965-07-09" collectionCode="OLL" county="Elburz" location="Tehran prov." municipality="Ab Ali" pageId="19" pageNumber="501" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="9B59882C325BFF93FE4CFED9FD8BE3A1" box="[455,607,1206,1232]" pageId="19" pageNumber="501" type="holotype">HOLOTYPE</typeStatus>
:
<location id="413D6055325BFF93FDE2FED9FD2FE3A1" LSID="urn:lsid:plazi:treatment:CC4B8798325BFF95FF1CFE69FE3FE614:413D6055325BFF93FDE2FED9FD2FE3A1" box="[617,763,1206,1232]" county="Elburz" municipality="Ab Ali" name="Tehran prov." pageId="19" pageNumber="501">Tehran prov.</location>
:
<collectingMunicipality id="A439ACF4325BFF93FC8FFED9FC84E3A1" box="[772,848,1206,1232]" pageId="19" pageNumber="501">Ab Ali</collectingMunicipality>
/
<collectingCounty id="AD3C4E02325BFF93FCD2FED9FC71E3A1" box="[857,933,1206,1232]" pageId="19" pageNumber="501">Elburz</collectingCounty>
,
<specimenCount id="52E4FD07325BFF93FC3AFED9FC01E3A1" box="[945,981,1206,1232]" pageId="19" pageNumber="501" type="male">1♂</specimenCount>
,
<date id="305C104E325BFF93FC69FEDAFB53E3A1" box="[994,1159,1205,1232]" pageId="19" pageNumber="501" value="1965-07-09" valueMax="1965-07-10" valueMin="1965-07-09">
<collectingDate id="2018E9A6325BFF93FC69FEDAFB53E3A1" box="[994,1159,1205,1232]" pageId="19" pageNumber="501" value="1965-07-09" valueMax="1965-07-10" valueMin="1965-07-09">9.–10.vii.1965</collectingDate>
</date>
, Soika &amp; Mavromousta- kis leg. (
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;
<bibRefCitation id="20734B7F325BFF93FEC4FEB5FE29E385" author="Warncke, K." box="[335,509,1242,1268]" pageId="19" pageNumber="501" pagination="119 - 210" refId="ref18439" refString="Warncke, K. (1980) Die Bienengattung Anthidium Fabricius, 1804 in der Westpalaarktis und im turkestanischen Becken. Entomofauna, 1, 119 - 210." type="journal article" year="1980">Warncke, 1980</bibRefCitation>
)
</materialsCitation>
.—
<materialsCitation id="F48A3CD3325BFF93FDAFFEB5FCEBE269" collectingDate="1960-06-27" collectingDateMax="1960-07-03" collectingDateMin="1960-06-27" collectionCode="OLL" country="Lebanon" location="Becharre" pageId="19" pageNumber="501" specimenCount="8" specimenCount-female="6" specimenCount-male="2" typeStatus="paratype">
<typeStatus id="9B59882C325BFF93FDAFFEB5FD1DE385" box="[548,713,1242,1268]" pageId="19" pageNumber="501" type="paratype">PARATYPES</typeStatus>
: N.
<collectingCountry id="3CF5761E325BFF93FC89FEB5FCBCE385" box="[770,872,1242,1268]" name="Lebanon" pageId="19" pageNumber="501">Lebanon</collectingCountry>
:
<location id="413D6055325BFF93FCF3FEB5FC34E385" LSID="urn:lsid:plazi:treatment:CC4B8798325BFF95FF1CFE69FE3FE614:413D6055325BFF93FCF3FEB5FC34E385" box="[888,992,1242,1268]" country="Lebanon" name="Becharre" pageId="19" pageNumber="501">Becharre</location>
,
<specimenCount id="52E4FD07325BFF93FC7AFEB5FBC0E385" box="[1009,1044,1242,1268]" pageId="19" pageNumber="501" type="female">1♀</specimenCount>
,
<specimenCount id="52E4FD07325BFF93FBA3FEB5FB99E385" box="[1064,1101,1242,1268]" pageId="19" pageNumber="501" type="male">1♂</specimenCount>
,
<date id="305C104E325BFF93FBD4FEB5FB00E385" box="[1119,1236,1242,1268]" pageId="19" pageNumber="501" value="1960-07-01">
<collectingDate id="2018E9A6325BFF93FBD4FEB5FB00E385" box="[1119,1236,1242,1268]" pageId="19" pageNumber="501" value="1960-07-01">1.vii.1960</collectingDate>
</date>
;
<specimenCount id="52E4FD07325BFF93FB6DFEB5FADDE385" box="[1254,1289,1242,1268]" pageId="19" pageNumber="501" type="female">1♀</specimenCount>
,
<date id="305C104E325BFF93FA96FEB5FA4CE385" box="[1309,1432,1242,1268]" pageId="19" pageNumber="501" value="1960-06-27">
<collectingDate id="2018E9A6325BFF93FA96FEB5FA4CE385" box="[1309,1432,1242,1268]" pageId="19" pageNumber="501" value="1960-06-27">27.vi.1960</collectingDate>
</date>
;
<specimenCount id="52E4FD07325BFF93FF1CFE91FF68E269" box="[151,188,1278,1304]" pageId="19" pageNumber="501" type="female">4♀</specimenCount>
, 2,
<date id="305C104E325BFF93FF55FE91FE94E269" box="[222,320,1278,1304]" pageId="19" pageNumber="501" value="1960-07">
<collectingDate id="2018E9A6325BFF93FF55FE91FE94E269" box="[222,320,1278,1304]" pageId="19" pageNumber="501" value="1960-07">vii.1960</collectingDate>
</date>
;
<specimenCount id="52E4FD07325BFF93FEC5FE91FEA7E269" box="[334,371,1278,1304]" pageId="19" pageNumber="501" type="male">1♂</specimenCount>
,
<date id="305C104E325BFF93FE09FE91FE20E269" box="[386,500,1278,1304]" pageId="19" pageNumber="501" value="1960-07-03">
<collectingDate id="2018E9A6325BFF93FE09FE91FE20E269" box="[386,500,1278,1304]" pageId="19" pageNumber="501" value="1960-07-03">3.vii.1960</collectingDate>
</date>
(
<collectionCode id="22F3AE4B325BFF93FD8DFE91FD95E269" box="[518,577,1278,1304]" pageId="19" pageNumber="501">OLL</collectionCode>
;
<bibRefCitation id="20734B7F325BFF93FDC4FE91FD2AE269" author="Warncke, K." box="[591,766,1278,1304]" pageId="19" pageNumber="501" pagination="119 - 210" refId="ref18439" refString="Warncke, K. (1980) Die Bienengattung Anthidium Fabricius, 1804 in der Westpalaarktis und im turkestanischen Becken. Entomofauna, 1, 119 - 210." type="journal article" year="1980">Warncke, 1980</bibRefCitation>
).—N
</materialsCitation>
.
<materialsCitation id="F48A3CD3325BFF93FCC7FE91FE98E211" collectingDate="21. vi. 1960, 27. vi. 1960, 22. vi. 1960" country="Lebanon" location="Kadisha" pageId="19" pageNumber="501" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="paratype">
<collectingCountry id="3CF5761E325BFF93FCC7FE91FC66E269" box="[844,946,1278,1304]" name="Lebanon" pageId="19" pageNumber="501">Lebanon</collectingCountry>
:
<location id="413D6055325BFF93FC4BFE91FBCBE269" LSID="urn:lsid:plazi:treatment:CC4B8798325BFF95FF1CFE69FE3FE614:413D6055325BFF93FC4BFE91FBCBE269" box="[960,1055,1278,1304]" country="Lebanon" name="Kadisha" pageId="19" pageNumber="501">Kadisha</location>
,
<specimenCount id="52E4FD07325BFF93FBA6FE91FB85E269" box="[1069,1105,1278,1304]" pageId="19" pageNumber="501" type="female">2♀</specimenCount>
,
<specimenCount id="52E4FD07325BFF93FBEAFE91FB51E269" box="[1121,1157,1278,1304]" pageId="19" pageNumber="501" type="male">1♂</specimenCount>
,
<collectingDate id="2018E9A6325BFF93FB1EFE91FEDBE24D" isEnumeration="true" pageId="19" pageNumber="501">
<date id="305C104E325BFF93FB1EFE91FADBE269" box="[1173,1295,1278,1304]" pageId="19" pageNumber="501" value="1960-06-21">21.vi.1960</date>
,
<date id="305C104E325BFF93FA96FE91FA43E269" box="[1309,1431,1278,1304]" pageId="19" pageNumber="501" value="1960-06-27">27.vi.1960</date>
,
<date id="305C104E325BFF93FF1CFF4DFEDBE24D" box="[151,271,1314,1340]" pageId="19" pageNumber="501" value="1960-06-22">22.vi.1960</date>
</collectingDate>
(listed by Mavromoustakis, 1962 as
<taxonomicName id="83E24D0D325BFF93FD4BFF4DFCBAE24D" box="[704,878,1314,1340]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="501" phylum="Arthropoda" rank="species" species="anguliventre">
<emphasis id="7696EA9C325BFF93FD4BFF4DFCBAE24D" box="[704,878,1314,1340]" italics="true" pageId="19" pageNumber="501">A. anguliventre</emphasis>
</taxonomicName>
, but transferred to
<taxonomicName id="83E24D0D325BFF93FBDAFF4DFB0BE24D" authorityName="Warncke" authorityYear="1980" box="[1105,1247,1314,1340]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="501" phylum="Arthropoda" rank="species" species="rotundum">
<emphasis id="7696EA9C325BFF93FBDAFF4DFB0BE24D" box="[1105,1247,1314,1340]" italics="true" pageId="19" pageNumber="501">A. rotundum</emphasis>
</taxonomicName>
as
<typeStatus id="9B59882C325BFF93FA80FF4DFAA2E24D" box="[1291,1398,1314,1340]" pageId="19" pageNumber="501" type="paratype">paratypes</typeStatus>
by
<bibRefCitation id="20734B7F325BFF93FF1CFF29FE90E211" author="Warncke, K." box="[151,324,1350,1376]" pageId="19" pageNumber="501" pagination="119 - 210" refId="ref18439" refString="Warncke, K. (1980) Die Bienengattung Anthidium Fabricius, 1804 in der Westpalaarktis und im turkestanischen Becken. Entomofauna, 1, 119 - 210." type="journal article" year="1980">Warncke, 1980</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="445D368E325BFF93FF4CFF06FCFBE029" blockId="19.[151,1437,1205,2024]" pageId="19" pageNumber="501">
<emphasis id="7696EA9C325BFF93FF4CFF06FE21E2F2" bold="true" box="[199,501,1385,1411]" pageId="19" pageNumber="501">Other material examined.</emphasis>
<materialsCitation id="F48A3CD3325BFF93FE72FF06FD29E12D" collectingDate="1960-06-27" collectingDateMax="2019-07-06" collectingDateMin="1960-06-27" collectionCode="CMK" collectorName="N. Lebanon &amp; G. Mavromoustakis &amp; N. Liban &amp; Barouk &amp; X. van Achter &amp; M. Boustani &amp; Mbou &amp; Laqlouq &amp; Matoube" country="Lebanon" county="Bcharre" elevation="1692" latitude="34.116665" location="Shouf Biosphere Reserve" longLatPrecision="1201" longitude="35.85" municipality="Shouf Biosphere Reserve" pageId="19" pageNumber="501" specimenCount="8" specimenCount-female="5" specimenCount-male="3">
<emphasis id="7696EA9C325BFF93FE72FF06FD57E2F2" bold="true" box="[505,643,1385,1411]" pageId="19" pageNumber="501">
<collectingCountry id="3CF5761E325BFF93FE72FF06FD57E2F2" box="[505,643,1385,1411]" name="Lebanon" pageId="19" pageNumber="501">LEBANON</collectingCountry>
</emphasis>
(see also
<bibRefCitation id="20734B7F325BFF93FD61FF05FC1AE2F5" author="Boustani, M. &amp; Dathe, H. &amp; Ghisbain, G. &amp; Kasparek, M. &amp; Michez, D. &amp; Nemer, N. &amp; Pauly, A. &amp; Risch, S. &amp; Terzo, M. &amp; Van Achter, X. &amp; Wood, T. &amp; Rasmont, P." box="[746,974,1386,1412]" pageId="19" pageNumber="501" pagination="1 - 146" refId="ref16727" refString="Boustani, M., Dathe, H., Ghisbain, G., Kasparek, M., Michez, D., Nemer, N., Pauly, A., Risch, S., Terzo, M., Van Achter, X., Wood, T. &amp; Rasmont, P. (2021) The bees of Lebanon (Hymenoptera: Apoidea: Anthophila). Zootaxa, 4976, 1 - 146. https: // doi. org / 10.11646 / zootaxa. 4976.1.1" type="journal article" year="2021">
Boustani
<emphasis id="7696EA9C325BFF93FCDAFF05FC53E2F5" box="[849,903,1386,1412]" italics="true" pageId="19" pageNumber="501">et al.</emphasis>
, 2021
</bibRefCitation>
):
<collectorName id="E9175358325BFF93FC55FF05FBB7E2F5" box="[990,1123,1386,1412]" pageId="19" pageNumber="501">N.Lebanon</collectorName>
:nr.
<collectingCounty id="AD3C4E02325BFF93FB07FF05FB3CE2F5" box="[1164,1256,1386,1412]" pageId="19" pageNumber="501">Bcharré</collectingCounty>
,
<specimenCount id="52E4FD07325BFF93FB64FF05FAC0E2F5" box="[1263,1300,1386,1412]" pageId="19" pageNumber="501" type="male">1♂</specimenCount>
,
<date id="305C104E325BFF93FA96FF05FA43E2F5" box="[1309,1431,1386,1412]" pageId="19" pageNumber="501" value="1960-06-27">
<collectingDate id="2018E9A6325BFF93FA96FF05FA43E2F5" box="[1309,1431,1386,1412]" pageId="19" pageNumber="501" value="1960-06-27">27.vi.1960</collectingDate>
</date>
,
<collectorName id="E9175358325BFF93FF1CFFE1FEA1E2D9" box="[151,373,1422,1448]" pageId="19" pageNumber="501">G. Mavromoustakis</collectorName>
leg. (
<collectionCode id="22F3AE4B325BFF93FE38FFE1FE23E2D9" box="[435,503,1422,1448]" pageId="19" pageNumber="501">CMK</collectionCode>
, ms3907).—
<collectorName id="E9175358325BFF93FD02FFE1FD3AE2D9" box="[649,750,1422,1448]" pageId="19" pageNumber="501">N. Liban</collectorName>
:
<collectorName id="E9175358325BFF93FD7CFFE1FC98E2D9" box="[759,844,1422,1448]" pageId="19" pageNumber="501">Barouk</collectorName>
,
<collectingMunicipality id="A439ACF4325BFF93FCDCFFE1FBA6E2D9" box="[855,1138,1422,1448]" pageId="19" pageNumber="501">Shouf Biosphere Reserve</collectingMunicipality>
(
<geoCoordinate id="21D65049325BFF93FB0AFFE1FB31E2D9" box="[1153,1253,1422,1448]" degrees="33" direction="north" minutes="40" orientation="latitude" pageId="19" pageNumber="501" precision="925" value="33.666668">33°40’N</geoCoordinate>
,
<geoCoordinate id="21D65049325BFF93FB7BFFE2FA84E2D9" box="[1264,1360,1421,1448]" degrees="35" direction="east" minutes="41" orientation="longitude" pageId="19" pageNumber="501" precision="925" value="35.683334">35°41’E</geoCoordinate>
),
<quantity id="831A9B6B325BFF93FAEFFFE1FF7BE2BD" metricMagnitude="3" metricUnit="m" metricValue="1.6919999999999997" pageId="19" pageNumber="501" unit="m" value="1692.0">
<elevation id="CFCFD1BD325BFF93FAEFFFE1FF7BE2BD" metricMagnitude="3" metricUnit="m" metricValue="1.6919999999999997" pageId="19" pageNumber="501" unit="m" value="1692.0">1692 m</elevation>
</quantity>
,
<specimenCount id="52E4FD07325BFF93FF31FFDDFF0AE2BD" box="[186,222,1458,1484]" pageId="19" pageNumber="501" type="male">1♂</specimenCount>
,
<date id="305C104E325BFF93FF62FFDDFEBFE2BD" box="[233,363,1458,1484]" pageId="19" pageNumber="501" value="2019-07-02">
<collectingDate id="2018E9A6325BFF93FF62FFDDFEBFE2BD" box="[233,363,1458,1484]" pageId="19" pageNumber="501" value="2019-07-02">02.vii.2019</collectingDate>
</date>
,
<collectorName id="E9175358325BFF93FEFDFFDDFDDBE2BD" box="[374,527,1458,1484]" pageId="19" pageNumber="501">X. van Achter</collectorName>
leg. (coll.
<collectorName id="E9175358325BFF93FD0FFFDDFCC4E2BD" box="[644,784,1458,1484]" pageId="19" pageNumber="501">M. Boustani</collectorName>
,
<collectorName id="E9175358325BFF93FC91FFDDFCB6E2BC" box="[794,866,1458,1485]" pageId="19" pageNumber="501">Mbou</collectorName>
124).—
<collectorName id="E9175358325BFF93FC3DFFDDFBCEE2BD" box="[950,1050,1458,1485]" pageId="19" pageNumber="501">N. Liban</collectorName>
:
<collectorName id="E9175358325BFF93FBA8FFDDFBACE2BD" box="[1059,1144,1458,1484]" pageId="19" pageNumber="501">Barouk</collectorName>
,
<location id="413D6055325BFF93FB09FFDDFA48E2BD" LSID="urn:lsid:plazi:treatment:CC4B8798325BFF95FF1CFE69FE3FE614:413D6055325BFF93FB09FFDDFA48E2BD" box="[1154,1436,1458,1484]" country="Lebanon" county="Bcharre" latitude="34.116665" longLatPrecision="1201" longitude="35.85" municipality="Shouf Biosphere Reserve" name="Shouf Biosphere Reserve" pageId="19" pageNumber="501">Shouf Biosphere Reserve</location>
(
<geoCoordinate id="21D65049325BFF93FF2BFFB9FED0E281" box="[160,260,1494,1520]" degrees="33" direction="north" minutes="39" orientation="latitude" pageId="19" pageNumber="501" precision="925" value="33.65">33°39’N</geoCoordinate>
,
<geoCoordinate id="21D65049325BFF93FE98FFBAFEA0E281" box="[275,372,1493,1520]" degrees="35" direction="east" minutes="40" orientation="longitude" pageId="19" pageNumber="501" precision="925" value="35.666668">35°40’E</geoCoordinate>
),
<quantity id="831A9B6B325BFF93FE00FFB9FE32E281" box="[395,486,1494,1520]" metricMagnitude="3" metricUnit="m" metricValue="1.382" pageId="19" pageNumber="501" unit="m" value="1382.0">
<elevation id="CFCFD1BD325BFF93FE00FFB9FE32E281" box="[395,486,1494,1520]" metricMagnitude="3" metricUnit="m" metricValue="1.382" pageId="19" pageNumber="501" unit="m" value="1382.0">1382 m</elevation>
</quantity>
,
<specimenCount id="52E4FD07325BFF93FE7EFFB9FDCCE281" box="[501,536,1494,1520]" pageId="19" pageNumber="501" type="female">1♀</specimenCount>
,
<specimenCount id="52E4FD07325BFF93FDA2FFB9FD9AE281" box="[553,590,1494,1520]" pageId="19" pageNumber="501" type="male">1♂</specimenCount>
,
<date id="305C104E325BFF93FDD5FFB9FD34E281" box="[606,736,1494,1520]" pageId="19" pageNumber="501" value="2019-07-03">
<collectingDate id="2018E9A6325BFF93FDD5FFB9FD34E281" box="[606,736,1494,1520]" pageId="19" pageNumber="501" value="2019-07-03">03.vii.2019</collectingDate>
</date>
,
<collectorName id="E9175358325BFF93FD64FFB9FC46E281" box="[751,914,1494,1520]" pageId="19" pageNumber="501">X. van Achter</collectorName>
leg. (coll.
<collectorName id="E9175358325BFF93FB9EFFB9FB71E281" box="[1045,1189,1494,1520]" pageId="19" pageNumber="501">M. Boustani</collectorName>
,
<collectorName id="E9175358325BFF93FB3FFFBAFB29E281" box="[1204,1277,1493,1520]" pageId="19" pageNumber="501">Mbou</collectorName>
125-126).—
<collectorName id="E9175358325BFF93FA08FFBAFF09E165" pageId="19" pageNumber="501">N. Liban</collectorName>
:
<collectorName id="E9175358325BFF93FF62FF95FEEAE165" box="[233,318,1530,1556]" pageId="19" pageNumber="501">Barouk</collectorName>
,
<location id="413D6055325BFF93FEC1FF95FDBEE165" LSID="urn:lsid:plazi:treatment:CC4B8798325BFF95FF1CFE69FE3FE614:413D6055325BFF93FEC1FF95FDBEE165" box="[330,618,1530,1556]" country="Lebanon" county="Bcharre" latitude="34.116665" longLatPrecision="1201" longitude="35.85" municipality="Shouf Biosphere Reserve" name="Shouf Biosphere Reserve" pageId="19" pageNumber="501">Shouf Biosphere Reserve</location>
(
<geoCoordinate id="21D65049325BFF93FDF0FF95FD0AE165" box="[635,734,1530,1556]" degrees="33" direction="north" minutes="40" orientation="latitude" pageId="19" pageNumber="501" precision="925" value="33.666668">33°40’N</geoCoordinate>
,
<geoCoordinate id="21D65049325BFF93FD67FF96FC99E165" box="[748,845,1529,1556]" degrees="35" direction="east" minutes="42" orientation="longitude" pageId="19" pageNumber="501" precision="925" value="35.7">35°42’E</geoCoordinate>
),
<quantity id="831A9B6B325BFF93FCE9FF95FC6EE165" box="[866,954,1530,1556]" metricMagnitude="3" metricUnit="m" metricValue="1.716" pageId="19" pageNumber="501" unit="m" value="1716.0">
<elevation id="CFCFD1BD325BFF93FCE9FF95FC6EE165" box="[866,954,1530,1556]" metricMagnitude="3" metricUnit="m" metricValue="1.716" pageId="19" pageNumber="501" unit="m" value="1716.0">1716 m</elevation>
</quantity>
,
<specimenCount id="52E4FD07325BFF93FC4CFF95FC38E165" box="[967,1004,1530,1556]" pageId="19" pageNumber="501" type="female">2♀</specimenCount>
,
<date id="305C104E325BFF93FC71FF95FBA8E165" box="[1018,1148,1530,1556]" pageId="19" pageNumber="501" value="2019-07-04">
<collectingDate id="2018E9A6325BFF93FC71FF95FBA8E165" box="[1018,1148,1530,1556]" pageId="19" pageNumber="501" value="2019-07-04">04.vii.2019</collectingDate>
</date>
,
<collectorName id="E9175358325BFF93FB02FF95FAFCE165" box="[1161,1320,1530,1556]" pageId="19" pageNumber="501">X. van Achter</collectorName>
leg. (coll.
<collectorName id="E9175358325BFF93FF1CFC71FEF1E149" box="[151,293,1566,1592]" pageId="19" pageNumber="501">M. Boustani</collectorName>
, MBou127-128).—
<collectorName id="E9175358325BFF93FD8EFC71FDB8E149" box="[517,620,1566,1592]" pageId="19" pageNumber="501">N. Liban</collectorName>
:
<collectorName id="E9175358325BFF93FDF3FC71FD0CE149" box="[632,728,1566,1592]" pageId="19" pageNumber="501">Laqlouq</collectorName>
,
<collectorName id="E9175358325BFF93FD6FFC71FC9CE149" box="[740,840,1566,1592]" pageId="19" pageNumber="501">Matoube</collectorName>
(
<geoCoordinate id="21D65049325BFF93FCD3FC71FC68E149" box="[856,956,1566,1592]" degrees="34" direction="north" minutes="07" orientation="latitude" pageId="19" pageNumber="501" precision="925" value="34.116665">34°07’N</geoCoordinate>
,
<geoCoordinate id="21D65049325BFF93FC42FC72FBFDE149" box="[969,1065,1565,1592]" degrees="35" direction="east" minutes="51" orientation="longitude" pageId="19" pageNumber="501" precision="925" value="35.85">35°51’E</geoCoordinate>
), 1660/
<quantity id="831A9B6B325BFF93FBF4FC71FB03E149" box="[1151,1239,1566,1592]" metricMagnitude="3" metricUnit="m" metricValue="1.699" pageId="19" pageNumber="501" unit="m" value="1699.0">
<elevation id="CFCFD1BD325BFF93FBF4FC71FB03E149" box="[1151,1239,1566,1592]" metricMagnitude="3" metricUnit="m" metricValue="1.699" pageId="19" pageNumber="501" unit="m" value="1699.0">1699 m</elevation>
</quantity>
,
<specimenCount id="52E4FD07325BFF93FB6FFC71FADCE149" box="[1252,1288,1566,1592]" pageId="19" pageNumber="501" type="female">2♀</specimenCount>
,
<date id="305C104E325BFF93FA9DFC71FA43E149" box="[1302,1431,1566,1592]" pageId="19" pageNumber="501" value="2019-07-06">
<collectingDate id="2018E9A6325BFF93FA9DFC71FA43E149" box="[1302,1431,1566,1592]" pageId="19" pageNumber="501" value="2019-07-06">06.vii.2019</collectingDate>
</date>
,
<collectorName id="E9175358325BFF93FF1CFC2DFEE2E12D" box="[151,310,1602,1628]" pageId="19" pageNumber="501">X. van Achter</collectorName>
leg. (coll.
<collectorName id="E9175358325BFF93FE38FC2DFD95E12D" box="[435,577,1602,1628]" pageId="19" pageNumber="501">M. Boustani</collectorName>
, MBou129-130)
</materialsCitation>
.—
<materialsCitation id="F48A3CD3325BFF93FC94FC2EFE80E1F1" collectingDate="2000-06-21" collectionCode="CMK" collectorName="M. Halada" country="Syria" location="Kafr Suwayda" pageId="19" pageNumber="501" specimenCount="1" specimenCount-male="1">
<emphasis id="7696EA9C325BFF93FC94FC2EFCABE12A" bold="true" box="[799,895,1601,1627]" pageId="19" pageNumber="501">
<collectingCountry id="3CF5761E325BFF93FC94FC2EFCAEE12A" box="[799,890,1601,1627]" name="Syria" pageId="19" pageNumber="501">SYRIA</collectingCountry>
:
</emphasis>
<location id="413D6055325BFF93FC03FC2DFBF9E12D" LSID="urn:lsid:plazi:treatment:CC4B8798325BFF95FF1CFE69FE3FE614:413D6055325BFF93FC03FC2DFBF9E12D" box="[904,1069,1602,1628]" country="Syria" name="Kafr Suwayda" pageId="19" pageNumber="501">Kafr Suwayda</location>
,
<specimenCount id="52E4FD07325BFF93FBB1FC2DFB8AE12D" box="[1082,1118,1602,1628]" pageId="19" pageNumber="501" type="male">1♂</specimenCount>
,
<date id="305C104E325BFF93FBE7FC2DFB32E12D" box="[1132,1254,1602,1628]" pageId="19" pageNumber="501" value="2000-06-21">
<collectingDate id="2018E9A6325BFF93FBE7FC2DFB32E12D" box="[1132,1254,1602,1628]" pageId="19" pageNumber="501" value="2000-06-21">21.vi.2000</collectingDate>
</date>
,
<collectorName id="E9175358325BFF93FB78FC2DFABEE12D" box="[1267,1386,1602,1628]" pageId="19" pageNumber="501">M. Halada</collectorName>
leg. (
<collectionCode id="22F3AE4B325BFF93FF14FC09FF37E1F1" box="[159,227,1638,1664]" pageId="19" pageNumber="501">CMK</collectionCode>
, ms2900)
</materialsCitation>
.—
<materialsCitation id="F48A3CD3325BFF93FEFDFC09FF0CE1D4" collectingDate="1985-06-21" collectionCode="CMK" collectorName="M. Schwarz" country="Turkey" location="Golbasi" pageId="19" pageNumber="501" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Adiyaman">
<emphasis id="7696EA9C325BFF93FEFDFC09FE20E1F1" bold="true" box="[374,500,1638,1664]" pageId="19" pageNumber="501">
<collectingCountry id="3CF5761E325BFF93FEFDFC09FE24E1F1" box="[374,496,1638,1664]" name="Turkey" pageId="19" pageNumber="501">TURKEY</collectingCountry>
:
</emphasis>
<collectingRegion id="8626F86C325BFF93FE75FC09FD6DE1F1" box="[510,697,1638,1664]" country="Turkey" name="Adiyaman" pageId="19" pageNumber="501">Adıyaman prov.</collectingRegion>
:
<location id="413D6055325BFF93FD4EFC09FCCBE1F1" LSID="urn:lsid:plazi:treatment:CC4B8798325BFF95FF1CFE69FE3FE614:413D6055325BFF93FD4EFC09FCCBE1F1" box="[709,799,1638,1664]" country="Turkey" name="Golbasi" pageId="19" pageNumber="501" stateProvince="Adiyaman">Gölbaşı</location>
,
<specimenCount id="52E4FD07325BFF93FCA6FC09FC85E1F1" box="[813,849,1638,1664]" pageId="19" pageNumber="501" type="female">2♀</specimenCount>
,
<specimenCount id="52E4FD07325BFF93FCEBFC09FC51E1F1" box="[864,901,1638,1664]" pageId="19" pageNumber="501" type="male">3♂</specimenCount>
,
<date id="305C104E325BFF93FC18FC0AFBD9E1F1" box="[915,1037,1637,1664]" pageId="19" pageNumber="501" value="1985-06-21">
<collectingDate id="2018E9A6325BFF93FC18FC0AFBD9E1F1" box="[915,1037,1637,1664]" pageId="19" pageNumber="501" value="1985-06-21">21.vi.1985</collectingDate>
</date>
,
<collectorName id="E9175358325BFF93FB90FC09FB70E1F1" box="[1051,1188,1638,1664]" pageId="19" pageNumber="501">M. Schwarz</collectorName>
leg. (
<collectionCode id="22F3AE4B325BFF93FB62FC09FAF9E1F1" box="[1257,1325,1638,1664]" pageId="19" pageNumber="501">CMK</collectionCode>
, ms2301- 2302)
</materialsCitation>
.—
<materialsCitation id="F48A3CD3325BFF93FF73FCE5FAE8E1D4" box="[248,1340,1674,1701]" collectingDate="1993-07-02" collectionCode="CMK" collectorName="Jirousek" country="Turkey" location="Karadut" municipality="Nemrut Dagi" pageId="19" pageNumber="501" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Adiyaman">
<collectingRegion id="8626F86C325BFF93FF73FCE5FE62E1D5" box="[248,438,1674,1701]" country="Turkey" name="Adiyaman" pageId="19" pageNumber="501">Adıyaman prov.</collectingRegion>
:
<collectingMunicipality id="A439ACF4325BFF93FE49FCE5FD8DE1D5" box="[450,601,1674,1700]" pageId="19" pageNumber="501">Nemrut Dağı</collectingMunicipality>
,
<location id="413D6055325BFF93FDEDFCE5FD17E1D5" LSID="urn:lsid:plazi:treatment:CC4B8798325BFF95FF1CFE69FE3FE614:413D6055325BFF93FDEDFCE5FD17E1D5" box="[614,707,1674,1700]" country="Turkey" municipality="Nemrut Dagi" name="Karadut" pageId="19" pageNumber="501" stateProvince="Adiyaman">Karadut</location>
,
<specimenCount id="52E4FD07325BFF93FD5BFCE5FD27E1D5" box="[720,755,1674,1700]" pageId="19" pageNumber="501" type="female">1♀</specimenCount>
,
<specimenCount id="52E4FD07325BFF93FC88FCE5FCFCE1D5" box="[771,808,1674,1700]" pageId="19" pageNumber="501" type="male">2♂</specimenCount>
,
<date id="305C104E325BFF93FCBEFCE5FC63E1D5" box="[821,951,1674,1700]" pageId="19" pageNumber="501" value="1993-07-02">
<collectingDate id="2018E9A6325BFF93FCBEFCE5FC63E1D5" box="[821,951,1674,1700]" pageId="19" pageNumber="501" value="1993-07-02">02.vii.1993</collectingDate>
</date>
,
<collectorName id="E9175358325BFF93FC4EFCE5FBF6E1D5" box="[965,1058,1674,1700]" pageId="19" pageNumber="501">Jirousek</collectorName>
leg. (
<collectionCode id="22F3AE4B325BFF93FBEFFCE5FB7CE1D5" box="[1124,1192,1674,1700]" pageId="19" pageNumber="501">CMK</collectionCode>
, ms3924-26)
</materialsCitation>
.—
<materialsCitation id="F48A3CD3325BFF93FAD6FCE5FC10E1B9" collectionCode="ZMB" country="Turkey" location="Izmir" pageId="19" pageNumber="501" specimenCode="ZMB145" specimenCount="1" specimenCount-male="1" stateProvince="Izmir">
<collectingRegion id="8626F86C325BFF93FAD6FCE5FA49E1D4" box="[1373,1437,1674,1701]" country="Turkey" name="Izmir" pageId="19" pageNumber="501">Izmir</collectingRegion>
(„
<collectingRegion id="8626F86C325BFF93FF21FCC1FEDEE1B9" box="[170,266,1710,1736]" country="Turkey" name="Izmir" pageId="19" pageNumber="501">Smyrna</collectingRegion>
“),
<specimenCount id="52E4FD07325BFF93FEA0FCC1FE84E1B9" box="[299,336,1710,1736]" pageId="19" pageNumber="501" type="male">1♂</specimenCount>
, http://coll.mfn-berlin.de/u/31ef29 zmb145 (
<specimenCode id="14449EF5325BFF93FCDBFCC2FC68E1B9" box="[848,956,1709,1736]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="19" pageNumber="501" type="Museum">ZMB145</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="F48A3CD3325BFF93FC59FCC1FD91E19C" collectingDate="1983-08-10" collectionCode="OLL" collectorName="S Beyetussebap &amp; K. Warncke" country="Turkey" elevation="1250" location="Hakkari prov." pageId="19" pageNumber="501" specimenCount="1" stateProvince="Hakkari">
<collectingRegion id="8626F86C325BFF93FC59FCC1FBA0E1B9" box="[978,1140,1710,1736]" country="Turkey" name="Hakkari" pageId="19" pageNumber="501">Hakkari prov.</collectingRegion>
:
<collectorName id="E9175358325BFF93FB09FCC1FAEDE1B9" box="[1154,1337,1710,1736]" pageId="19" pageNumber="501">S Beyetüşşebap</collectorName>
,
<quantity id="831A9B6B325BFF93FACCFCC2FA4CE1B9" box="[1351,1432,1709,1736]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="19" pageNumber="501" unit="m" value="1250.0">
<elevation id="CFCFD1BD325BFF93FACCFCC2FA4CE1B9" box="[1351,1432,1709,1736]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="19" pageNumber="501" unit="m" value="1250.0">1250m</elevation>
</quantity>
,
<date id="305C104E325BFF93FF1CFCBDFEF5E19D" box="[151,289,1746,1772]" pageId="19" pageNumber="501" value="1983-08-10">
<collectingDate id="2018E9A6325BFF93FF1CFCBDFEF5E19D" box="[151,289,1746,1772]" pageId="19" pageNumber="501" value="1983-08-10">10.viii.1983</collectingDate>
</date>
,
<collectorName id="E9175358325BFF93FEB9FCBDFE6FE19D" box="[306,443,1746,1772]" pageId="19" pageNumber="501">K. Warncke</collectorName>
leg. (
<collectionCode id="22F3AE4B325BFF93FD8FFCBDFDE9E19C" box="[516,573,1746,1773]" pageId="19" pageNumber="501">OLL</collectionCode>
)
</materialsCitation>
.—
<materialsCitation id="F48A3CD3325BFF93FDEDFCBDFD20E061" collectingDate="2006-07-02" collectionCode="CMK" collectorName="J. Halada" country="Turkey" elevation="170" latitude="38.433334" location="15 km SEE Salihli" longLatPrecision="1175" longitude="28.316668" pageId="19" pageNumber="501" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Manisa">
<collectingRegion id="8626F86C325BFF93FDEDFCBDFCD0E19D" box="[614,772,1746,1773]" country="Turkey" name="Manisa" pageId="19" pageNumber="501">Manisa prov.</collectingRegion>
:
<location id="413D6055325BFF93FC98FCBEFC32E19D" LSID="urn:lsid:plazi:treatment:CC4B8798325BFF95FF1CFE69FE3FE614:413D6055325BFF93FC98FCBEFC32E19D" box="[787,998,1745,1772]" country="Turkey" latitude="38.433334" longLatPrecision="1175" longitude="28.316668" name="15 km SEE Salihli" pageId="19" pageNumber="501" stateProvince="Manisa">
<quantity id="831A9B6B325BFF93FC98FCBEFC87E19D" box="[787,851,1745,1772]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="19" pageNumber="501" unit="km" value="15.0">15km</quantity>
SEE Salihli
</location>
(
<geoCoordinate id="21D65049325BFF93FC70FCBDFB8AE19D" box="[1019,1118,1746,1772]" degrees="38" direction="north" minutes="26" orientation="latitude" pageId="19" pageNumber="501" precision="925" value="38.433334">38°26’N</geoCoordinate>
,
<geoCoordinate id="21D65049325BFF93FBE4FCBDFB1BE19D" box="[1135,1231,1746,1772]" degrees="28" direction="east" minutes="19" orientation="longitude" pageId="19" pageNumber="501" precision="925" value="28.316668">28°19’E</geoCoordinate>
),
<quantity id="831A9B6B325BFF93FB62FCBDFAF8E19D" box="[1257,1324,1746,1772]" metricMagnitude="2" metricUnit="m" metricValue="1.7" pageId="19" pageNumber="501" unit="m" value="170.0">
<elevation id="CFCFD1BD325BFF93FB62FCBDFAF8E19D" box="[1257,1324,1746,1772]" metricMagnitude="2" metricUnit="m" metricValue="1.7" pageId="19" pageNumber="501" unit="m" value="170.0">170m</elevation>
</quantity>
,
<specimenCount id="52E4FD07325BFF93FAB7FCBDFA8BE19D" box="[1340,1375,1746,1772]" pageId="19" pageNumber="501" type="female">1♀</specimenCount>
,
<specimenCount id="52E4FD07325BFF93FAF9FCBDFA43E19D" box="[1394,1431,1746,1772]" pageId="19" pageNumber="501" type="male">3♂</specimenCount>
,
<date id="305C104E325BFF93FF1CFC99FECDE061" box="[151,281,1782,1808]" pageId="19" pageNumber="501" value="2006-07-02">
<collectingDate id="2018E9A6325BFF93FF1CFC99FECDE061" box="[151,281,1782,1808]" pageId="19" pageNumber="501" value="2006-07-02">02.vii.2006</collectingDate>
</date>
,
<collectorName id="E9175358325BFF93FEA8FC99FE5EE061" box="[291,394,1782,1808]" pageId="19" pageNumber="501">J. Halada</collectorName>
leg. (
<collectionCode id="22F3AE4B325BFF93FE4CFC99FDDFE061" box="[455,523,1782,1808]" pageId="19" pageNumber="501">CMK</collectionCode>
, ms798-800, ms943)
</materialsCitation>
.—
<materialsCitation id="F48A3CD3325BFF93FC9DFC99FE12E044" collectingDate="2010-07-11" collectorName="M. Kasparek" country="Turkey" location="Kuyucak" municipality="Akyaka" pageId="19" pageNumber="501" specimenCount="1" specimenCount-male="1" stateProvince="Mugla">
<collectingRegion id="8626F86C325BFF93FC9DFC99FC76E061" box="[790,930,1782,1808]" country="Turkey" name="Mugla" pageId="19" pageNumber="501">Muğla prov.</collectingRegion>
: betw.
<collectingMunicipality id="A439ACF4325BFF93FC60FC99FB96E061" box="[1003,1090,1782,1808]" pageId="19" pageNumber="501">Akyaka</collectingMunicipality>
and
<location id="413D6055325BFF93FBFEFC99FB09E061" LSID="urn:lsid:plazi:treatment:CC4B8798325BFF95FF1CFE69FE3FE614:413D6055325BFF93FBFEFC99FB09E061" box="[1141,1245,1782,1808]" country="Turkey" municipality="Akyaka" name="Kuyucak" pageId="19" pageNumber="501" stateProvince="Mugla">Kuyucak</location>
,
<specimenCount id="52E4FD07325BFF93FB6CFC99FADFE061" box="[1255,1291,1782,1808]" pageId="19" pageNumber="501" type="male">1♂</specimenCount>
,
<date id="305C104E325BFF93FA9CFC99FA4CE061" box="[1303,1432,1782,1808]" pageId="19" pageNumber="501" value="2010-07-11">
<collectingDate id="2018E9A6325BFF93FA9CFC99FA4CE061" box="[1303,1432,1782,1808]" pageId="19" pageNumber="501" value="2010-07-11">11.vii.2010</collectingDate>
</date>
,
<collectorName id="E9175358325BFF93FF1CFD75FEFCE045" box="[151,296,1818,1844]" pageId="19" pageNumber="501">M. Kasparek</collectorName>
leg. (MK433)
</materialsCitation>
.—
<materialsCitation id="F48A3CD3325BFF93FE6DFD75FB65E044" box="[486,1201,1818,1845]" collectingDate="1977-06-14" collectingDateMax="1977-06-17" collectingDateMin="1977-06-14" collectionCode="SMF" collectorName="J. Heinrich" country="Turkey" location="Urfa prov." pageId="19" pageNumber="501" specimenCode="SMF238" specimenCount="1" specimenCount-male="1" stateProvince="Sanliurfa">
<collectingRegion id="8626F86C325BFF93FE6DFD75FDB5E045" box="[486,609,1818,1845]" country="Turkey" name="Sanliurfa" pageId="19" pageNumber="501">Urfa prov.</collectingRegion>
:
<collectingRegion id="8626F86C325BFF93FDE0FD75FD75E045" box="[619,673,1818,1844]" country="Turkey" name="Sanliurfa" pageId="19" pageNumber="501">Urfa</collectingRegion>
,
<specimenCount id="52E4FD07325BFF93FD20FD75FD04E045" box="[683,720,1818,1844]" pageId="19" pageNumber="501" type="male">1♂</specimenCount>
,
<date id="305C104E325BFF93FD57FD75FC56E045" box="[732,898,1818,1844]" pageId="19" pageNumber="501" value="1977-06-14" valueMax="1977-06-17" valueMin="1977-06-14">
<collectingDate id="2018E9A6325BFF93FD57FD75FC56E045" box="[732,898,1818,1844]" pageId="19" pageNumber="501" value="1977-06-14" valueMax="1977-06-17" valueMin="1977-06-14">14.-17.vi.1977</collectingDate>
</date>
,
<collectorName id="E9175358325BFF93FC06FD75FBDCE045" box="[909,1032,1818,1844]" pageId="19" pageNumber="501">J. Heinrich</collectorName>
leg. (
<specimenCode id="14449EF5325BFF93FBCEFD75FB7DE044" box="[1093,1193,1818,1845]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="19" pageNumber="501">SMF238</specimenCode>
)
</materialsCitation>
.—
<materialsCitation id="F48A3CD3325BFF93FB59FD75FCFFE029" collectingDate="1981-06-18" collectionCode="OLL" collectorName="K. Warncke" country="Turkey" elevation="500" location="20 km N Ceylanpinar" pageId="19" pageNumber="501" specimenCount="1" stateProvince="Sanliurfa">
<collectingRegion id="8626F86C325BFF93FB59FD75FA98E045" box="[1234,1356,1818,1845]" country="Turkey" name="Sanliurfa" pageId="19" pageNumber="501">Urfa prov.</collectingRegion>
:
<location id="413D6055325BFF93FADDFD75FEE9E029" LSID="urn:lsid:plazi:treatment:CC4B8798325BFF95FF1CFE69FE3FE614:413D6055325BFF93FADDFD75FEE9E029" country="Turkey" name="20 km N Ceylanpinar" pageId="19" pageNumber="501" stateProvince="Sanliurfa">
<quantity id="831A9B6B325BFF93FADDFD75FA48E045" box="[1366,1436,1818,1844]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="501" unit="km" value="20.0">20 km</quantity>
N Ceylanpınar
</location>
,
<quantity id="831A9B6B325BFF93FEC3FD52FE45E029" box="[328,401,1853,1880]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="19" pageNumber="501" unit="m" value="500.0">
<elevation id="CFCFD1BD325BFF93FEC3FD52FE45E029" box="[328,401,1853,1880]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="19" pageNumber="501" unit="m" value="500.0">500 m</elevation>
</quantity>
,
<date id="305C104E325BFF93FE16FD51FDC3E029" box="[413,535,1854,1880]" pageId="19" pageNumber="501" value="1981-06-18">
<collectingDate id="2018E9A6325BFF93FE16FD51FDC3E029" box="[413,535,1854,1880]" pageId="19" pageNumber="501" value="1981-06-18">18.vi.1981</collectingDate>
</date>
,
<collectorName id="E9175358325BFF93FDA8FD51FD73E029" box="[547,679,1854,1880]" pageId="19" pageNumber="501">K. Warncke</collectorName>
leg. (
<collectionCode id="22F3AE4B325BFF93FD6CFD51FCF7E029" box="[743,803,1854,1880]" pageId="19" pageNumber="501">OLL</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="445D368E325BFF94FF4CFD0EFB8AE684" blockId="19.[151,1437,1205,2024]" lastBlockId="20.[151,1437,150,2014]" lastPageId="20" lastPageNumber="502" pageId="19" pageNumber="501">
<emphasis id="7696EA9C325BFF93FF4CFD0EFD76E00A" bold="true" box="[199,674,1889,1916]" pageId="19" pageNumber="501">
Material not examined.
<collectingCountry id="3CF5761E325BFF93FE6EFD0EFD49E00A" box="[485,669,1889,1915]" name="Azerbaijan" pageId="19" pageNumber="501">AZERBAIJAN</collectingCountry>
.
</emphasis>
<collectingRegion id="8626F86C325BFF93FD22FD0DFCE4E00D" box="[681,816,1890,1916]" country="Azerbaijan" name="Naxcivan" pageId="19" pageNumber="501">Nakhchivan</collectingRegion>
Autonomous Republic:
<collectingRegion id="8626F86C325BFF93FBC9FD0DFB67E00D" box="[1090,1203,1890,1916]" country="Azerbaijan" name="Sahbuz" pageId="19" pageNumber="501">Shakhbuz</collectingRegion>
, Zarnatun (
<geoCoordinate id="21D65049325BFF93FABFFD0DFA43E00D" box="[1332,1431,1890,1916]" degrees="39" direction="north" minutes="31" orientation="latitude" pageId="19" pageNumber="501" precision="925" value="39.516666">39°31‘N</geoCoordinate>
,
<geoCoordinate id="21D65049325BFF93FF1CFDEAFF2CE0D1" box="[151,248,1925,1952]" degrees="45" direction="east" minutes="46" orientation="longitude" pageId="19" pageNumber="501" precision="925" value="45.766666">45°46‘E</geoCoordinate>
),
<quantity id="831A9B6B325BFF93FE84FDEAFEBDE0D1" box="[271,361,1925,1952]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="19" pageNumber="501" unit="m" value="1550.0">1550 m</quantity>
,
<specimenCount id="52E4FD07325BFF93FEF3FDE9FE49E0D1" box="[376,413,1926,1952]" pageId="19" pageNumber="501" type="female">4♀</specimenCount>
,
<specimenCount id="52E4FD07325BFF93FE27FDE9FE06E0D1" box="[428,466,1926,1952]" pageId="19" pageNumber="501" type="male">7♂</specimenCount>
,
<date id="305C104E325BFF93FE6AFDEAFD51E0D1" box="[481,645,1925,1952]" pageId="19" pageNumber="501" value="2018-07-24" valueMax="2018-07-25" valueMin="2018-07-24">24-25.vii.2018</date>
(
<bibRefCitation id="20734B7F325BFF93FD1CFDE9FC58E0D0" author="Fateryga, A. V. &amp; Proshchalykin, M. Yu. &amp; Maharramov, M. M." box="[663,908,1926,1953]" pageId="19" pageNumber="501" pagination="323 - 336" refId="ref17018" refString="Fateryga, A. V., Proshchalykin, M. Yu. &amp; Maharramov, M. M. (2020) Bees of the tribe Anthidiini (Hymenoptera, Megachilidae) of Nakhchivan Autonomous Republic of Azerbaijan. Entomological Review, 100, 323 - 336. https: // doi. org / 10.1134 / S 0013873820030069" type="journal article" year="2020">
Fateryga
<emphasis id="7696EA9C325BFF93FC8EFDE9FC94E0D1" box="[773,832,1926,1952]" italics="true" pageId="19" pageNumber="501">et al.</emphasis>
, 2020
</bibRefCitation>
).—
<collectingRegion id="8626F86C325BFF93FC3EFDE9FC3BE0D0" box="[949,1007,1926,1953]" country="Azerbaijan" name="Culfa" pageId="19" pageNumber="501">Julfa</collectingRegion>
, Gazanchi (
<geoCoordinate id="21D65049325BFF93FBF1FDE9FB09E0D1" box="[1146,1245,1926,1952]" degrees="39" direction="north" minutes="13" orientation="latitude" pageId="19" pageNumber="501" precision="925" value="39.216667">39°13’N</geoCoordinate>
,
<geoCoordinate id="21D65049325BFF93FB67FDEAFA99E0D1" box="[1260,1357,1925,1952]" degrees="45" direction="east" minutes="41" orientation="longitude" pageId="19" pageNumber="501" precision="925" value="45.683334">45°41’E</geoCoordinate>
),
<quantity id="831A9B6B325BFF93FAEFFDE9FF7BE0B5" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="19" pageNumber="501" unit="m" value="1300.0">1300 m</quantity>
,
<specimenCount id="52E4FD07325BFF93FF34FDC5FF36E0B5" box="[191,226,1962,1988]" pageId="19" pageNumber="501" type="female">1♀</specimenCount>
,
<specimenCount id="52E4FD07325BFF93FF7FFDC5FECDE0B5" box="[244,281,1962,1988]" pageId="19" pageNumber="501" type="male">2♂</specimenCount>
,
<date id="305C104E325BFF93FEA2FDC5FE19E0B5" box="[297,461,1962,1988]" pageId="19" pageNumber="501" value="2018-07-26" valueMax="2018-07-27" valueMin="2018-07-26">26-27.vii.2018</date>
(
<bibRefCitation id="20734B7F325BFF93FE6BFDC5FD02E0B4" author="Fateryga, A. V. &amp; Proshchalykin, M. Yu. &amp; Maharramov, M. M." box="[480,726,1962,1989]" pageId="19" pageNumber="501" pagination="323 - 336" refId="ref17018" refString="Fateryga, A. V., Proshchalykin, M. Yu. &amp; Maharramov, M. M. (2020) Bees of the tribe Anthidiini (Hymenoptera, Megachilidae) of Nakhchivan Autonomous Republic of Azerbaijan. Entomological Review, 100, 323 - 336. https: // doi. org / 10.1134 / S 0013873820030069" type="journal article" year="2020">
Fateryga
<emphasis id="7696EA9C325BFF93FDC5FDC5FD5FE0B5" box="[590,651,1962,1988]" italics="true" pageId="19" pageNumber="501">et al.</emphasis>
, 2020
</bibRefCitation>
).—
<collectingRegion id="8626F86C325BFF93FD74FDC5FCB2E0B4" box="[767,870,1962,1989]" country="Azerbaijan" name="Ordubad" pageId="19" pageNumber="501">Ordubad</collectingRegion>
, Aghdara (
<geoCoordinate id="21D65049325BFF93FC6CFDC5FB9EE0B5" box="[999,1098,1962,1988]" degrees="39" direction="north" minutes="06" orientation="latitude" pageId="19" pageNumber="501" precision="925" value="39.1">39°06’N</geoCoordinate>
,
<geoCoordinate id="21D65049325BFF93FBD1FDC6FB6FE0B5" box="[1114,1211,1961,1988]" degrees="45" direction="east" minutes="54" orientation="longitude" pageId="19" pageNumber="501" precision="925" value="45.9">45°54’E</geoCoordinate>
),
<quantity id="831A9B6B325BFF93FB58FDC5FAFAE0B5" box="[1235,1326,1962,1988]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="501" unit="m" value="2000.0">2000 m</quantity>
,
<specimenCount id="52E4FD07325BFF93FAB6FDC5FAB4E0B5" box="[1341,1376,1962,1988]" pageId="19" pageNumber="501" type="female">1♀</specimenCount>
,
<specimenCount id="52E4FD07325BFF93FAF9FDC5FA43E0B5" box="[1394,1431,1962,1988]" pageId="19" pageNumber="501" type="male">1♂</specimenCount>
,
<date id="305C104E325BFF93FF1CFDA1FEC2E099" box="[151,278,1998,2024]" pageId="19" pageNumber="501" value="2018-07-28">28.vii.2018</date>
(
<bibRefCitation id="20734B7F325BFF93FEACFDA1FDC2E099" author="Fateryga, A. V. &amp; Proshchalykin, M. Yu. &amp; Maharramov, M. M." box="[295,534,1998,2024]" pageId="19" pageNumber="501" pagination="323 - 336" refId="ref17018" refString="Fateryga, A. V., Proshchalykin, M. Yu. &amp; Maharramov, M. M. (2020) Bees of the tribe Anthidiini (Hymenoptera, Megachilidae) of Nakhchivan Autonomous Republic of Azerbaijan. Entomological Review, 100, 323 - 336. https: // doi. org / 10.1134 / S 0013873820030069" type="journal article" year="2020">
Fateryga
<emphasis id="7696EA9C325BFF93FE18FDA1FE18E099" box="[403,460,1998,2024]" italics="true" pageId="19" pageNumber="501">et al.</emphasis>
, 2020
</bibRefCitation>
).—
<emphasis id="7696EA9C325BFF93FDCBFDA2FD73E096" bold="true" box="[576,679,1997,2023]" pageId="19" pageNumber="501">
<collectingCountry id="3CF5761E325BFF93FDCBFDA2FD77E096" box="[576,675,1997,2023]" name="Egypt" pageId="19" pageNumber="501">EGYPT</collectingCountry>
:
</emphasis>
Rd. Maudi-Sokma [= Maadi, today part of
<collectingRegion id="8626F86C325BFF93FB10FDA1FB08E099" box="[1179,1244,1998,2024]" country="Egypt" name="Al Qahirah" pageId="19" pageNumber="501">Cairo</collectingRegion>
, and Ain Suchna (Ain Sokhna) in the Gulf of
<collectingRegion id="8626F86C325CFF94FE55FAF9FDCDE7C0" box="[478,537,150,177]" country="Egypt" name="As Suways" pageId="20" pageNumber="502">Suez</collectingRegion>
],
<specimenCount id="52E4FD07325CFF94FDA5FAF8FD86E7C0" box="[558,594,151,177]" pageId="20" pageNumber="502" type="male">1♂</specimenCount>
,
<date id="305C104E325CFF94FDEBFAF8FD18E7C0" box="[608,716,151,177]" pageId="20" pageNumber="502" value="1991-06-01">1.vi.1991</date>
, A. Mochi leg. (van der
<bibRefCitation id="20734B7F325CFF94FC6DFAF8FB54E7C3" author="Zanden, G. van der" box="[998,1152,151,178]" pageId="20" pageNumber="502" pagination="387 - 390" refId="ref18555" refString="Zanden, G. van der (1996) Neue Verbreitungsangaben zu einigen wenig bekannten palaarktischen Bienen-Arten (Insecta, Hymenoptera, Apoidea). Linzer biologische Beitrage, 28, 387 - 390." type="journal article" year="1996">Zanden, 1996</bibRefCitation>
).—Ikingi-Alexandria [c.
<geoCoordinate id="21D65049325CFF94FF1CFAD5FF2EE7A4" box="[151,250,186,213]" degrees="30" direction="north" minutes="59" orientation="latitude" pageId="20" pageNumber="502" precision="925" value="30.983334">30°59’N</geoCoordinate>
,
<geoCoordinate id="21D65049325CFF94FE8DFAD5FEBCE7A4" box="[262,360,186,213]" degrees="29" direction="east" minutes="45" orientation="longitude" pageId="20" pageNumber="502" precision="925" value="29.75">29°45’E</geoCoordinate>
],
<specimenCount id="52E4FD07325CFF94FEF0FAD4FE4BE7A4" box="[379,415,187,213]" pageId="20" pageNumber="502" type="male">1♂</specimenCount>
,
<date id="305C104E325CFF94FE27FAD4FDF9E7A4" box="[428,557,187,213]" pageId="20" pageNumber="502" value="1980-07-12">12.vii.1980</date>
, A. Mochi leg. (van der
<bibRefCitation id="20734B7F325CFF94FCB5FAD4FC08E7A7" author="Zanden, G. van der" box="[830,988,187,214]" pageId="20" pageNumber="502" pagination="387 - 390" refId="ref18555" refString="Zanden, G. van der (1996) Neue Verbreitungsangaben zu einigen wenig bekannten palaarktischen Bienen-Arten (Insecta, Hymenoptera, Apoidea). Linzer biologische Beitrage, 28, 387 - 390." type="journal article" year="1996">Zanden, 1996</bibRefCitation>
). Confirmation required.—
<emphasis id="7696EA9C325CFF94FA86FAD5FA8AE7A5" bold="true" box="[1293,1374,186,212]" pageId="20" pageNumber="502">
<collectingCountry id="3CF5761E325CFF94FA86FAD5FA8DE7A5" box="[1293,1369,186,212]" name="Iran" pageId="20" pageNumber="502">IRAN</collectingCountry>
:
</emphasis>
<collectingRegion id="8626F86C325CFF94FAEFFAD4FF00E788" country="Iran" name="Yazd" pageId="20" pageNumber="502">Yazd prov.</collectingRegion>
: Mehriz, Damgahan,
<specimenCount id="52E4FD07325CFF94FE49FAB0FE33E788" box="[450,487,223,249]" pageId="20" pageNumber="502" type="male">6♂</specimenCount>
,
<date id="305C104E325CFF94FE79FAB0FDA5E788" box="[498,625,223,249]" pageId="20" pageNumber="502" value="2013-07-03">03.vii.2013</date>
(
<bibRefCitation id="20734B7F325CFF94FDF4FAB0FC19E78B" author="Dehghan Dehnavi, L. &amp; Talebi, A. A. &amp; Goldasteh, Sh. &amp; Vafaei Shooshtari, R. &amp; Nadimi, A." box="[639,973,223,250]" pageId="20" pageNumber="502" pagination="1 - 21" refId="ref16822" refString="Dehghan Dehnavi, L., Talebi, A. A., Goldasteh, Sh., Vafaei Shooshtari, R. &amp; Nadimi, A. (2014) Contribution to the knowledge of Megachilidae (Hymenoptera: Apoidea) in the Yazd province, Iran. Journal of Entomological Research, 7, 1 - 21." type="journal article" year="2014">
Dehghan Dehnavi
<emphasis id="7696EA9C325CFF94FCDBFAB0FC5CE788" box="[848,904,223,249]" italics="true" pageId="20" pageNumber="502">et al.</emphasis>
, 2014
</bibRefCitation>
).—
<collectingRegion id="8626F86C325CFF94FC7EFAB0FBA0E788" box="[1013,1140,223,250]" country="Iran" name="Yazd" pageId="20" pageNumber="502">Yazd prov.</collectingRegion>
: Taft, Nir,
<specimenCount id="52E4FD07325CFF94FB6CFAB0FACDE788" box="[1255,1305,223,249]" pageId="20" pageNumber="502" type="male">11♂</specimenCount>
,
<date id="305C104E325CFF94FAAFFAB0FA49E788" box="[1316,1437,223,249]" pageId="20" pageNumber="502" value="2013-06-21">21.vi.2013</date>
(
<bibRefCitation id="20734B7F325CFF94FF14FB6CFE38E66F" author="Dehghan Dehnavi, L. &amp; Talebi, A. A. &amp; Goldasteh, Sh. &amp; Vafaei Shooshtari, R. &amp; Nadimi, A." box="[159,492,259,286]" pageId="20" pageNumber="502" pagination="1 - 21" refId="ref16822" refString="Dehghan Dehnavi, L., Talebi, A. A., Goldasteh, Sh., Vafaei Shooshtari, R. &amp; Nadimi, A. (2014) Contribution to the knowledge of Megachilidae (Hymenoptera: Apoidea) in the Yazd province, Iran. Journal of Entomological Research, 7, 1 - 21." type="journal article" year="2014">
Dehghan Dehnavi
<emphasis id="7696EA9C325CFF94FEFBFB6CFE73E66C" box="[368,423,259,285]" italics="true" pageId="20" pageNumber="502">et al.</emphasis>
, 2014
</bibRefCitation>
).—
<collectingRegion id="8626F86C325CFF94FD9FFB6CFD47E66C" box="[532,659,259,286]" country="Iran" name="Yazd" pageId="20" pageNumber="502">Yazd prov.</collectingRegion>
: Taft, Saleh abad,
<quantity id="831A9B6B325CFF94FCD1FB6DFC40E66C" box="[858,916,258,285]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="20" pageNumber="502" unit="m" value="15.0">15 m</quantity>
,
<date id="305C104E325CFF94FC15FB6CFBC2E66C" box="[926,1046,259,285]" pageId="20" pageNumber="502" value="2013-06-28">28.vi.2013</date>
(
<bibRefCitation id="20734B7F325CFF94FBA8FB6CFAA7E66C" author="Dehghan Dehnavi, L. &amp; Talebi, A. A. &amp; Goldasteh, Sh. &amp; Vafaei Shooshtari, R. &amp; Nadimi, A." box="[1059,1395,259,285]" pageId="20" pageNumber="502" pagination="1 - 21" refId="ref16822" refString="Dehghan Dehnavi, L., Talebi, A. A., Goldasteh, Sh., Vafaei Shooshtari, R. &amp; Nadimi, A. (2014) Contribution to the knowledge of Megachilidae (Hymenoptera: Apoidea) in the Yazd province, Iran. Journal of Entomological Research, 7, 1 - 21." type="journal article" year="2014">
Dehghan Dehnavi
<emphasis id="7696EA9C325CFF94FB7FFB6CFAFFE66C" box="[1268,1323,259,285]" italics="true" pageId="20" pageNumber="502">et al.</emphasis>
, 2014
</bibRefCitation>
).—
<collectingRegion id="8626F86C325CFF94FF1CFB48FEC7E630" box="[151,275,295,321]" country="Iran" name="Yazd" pageId="20" pageNumber="502">Yazd prov.</collectingRegion>
: Taft, Rahat abad,
<specimenCount id="52E4FD07325CFF94FE6BFB49FDD1E630" box="[480,517,294,321]" pageId="20" pageNumber="502" type="male">5♂</specimenCount>
,
<date id="305C104E325CFF94FD9AFB48FD45E630" box="[529,657,295,321]" pageId="20" pageNumber="502" value="2013-07-14">14.vii.2013</date>
(
<bibRefCitation id="20734B7F325CFF94FD15FB48FC27E630" author="Dehghan Dehnavi, L. &amp; Talebi, A. A. &amp; Goldasteh, Sh. &amp; Vafaei Shooshtari, R. &amp; Nadimi, A." box="[670,1011,295,321]" pageId="20" pageNumber="502" pagination="1 - 21" refId="ref16822" refString="Dehghan Dehnavi, L., Talebi, A. A., Goldasteh, Sh., Vafaei Shooshtari, R. &amp; Nadimi, A. (2014) Contribution to the knowledge of Megachilidae (Hymenoptera: Apoidea) in the Yazd province, Iran. Journal of Entomological Research, 7, 1 - 21." type="journal article" year="2014">
Dehghan Dehnavi
<emphasis id="7696EA9C325CFF94FCFAFB48FC7DE630" box="[881,937,295,321]" italics="true" pageId="20" pageNumber="502">et al.</emphasis>
, 2014
</bibRefCitation>
). –
<collectingRegion id="8626F86C325CFF94FB98FB48FB70E630" box="[1043,1188,295,321]" country="Iran" name="Tehran" pageId="20" pageNumber="502">Tehran prov.</collectingRegion>
: Monti steppici
<quantity id="831A9B6B325CFF94FADDFB48FA48E630" box="[1366,1436,295,321]" metricMagnitude="4" metricUnit="m" metricValue="2.4" pageId="20" pageNumber="502" unit="km" value="24.0">24 km</quantity>
S Ab Ali,
<quantity id="831A9B6B325CFF94FE8EFB24FE88E614" box="[261,348,331,357]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="20" pageNumber="502" unit="m" value="1300.0">1300 m</quantity>
,
<specimenCount id="52E4FD07325CFF94FEECFB24FE58E614" box="[359,396,331,357]" pageId="20" pageNumber="502" type="male">1♂</specimenCount>
,
<date id="305C104E325CFF94FE13FB25FDCCE614" box="[408,536,330,357]" pageId="20" pageNumber="502" value="1965-07-10">10.vii.1965</date>
(
<bibRefCitation id="20734B7F325CFF94FDADFB24FD04E617" author="Warncke, K." box="[550,720,331,358]" pageId="20" pageNumber="502" pagination="119 - 210" refId="ref18439" refString="Warncke, K. (1980) Die Bienengattung Anthidium Fabricius, 1804 in der Westpalaarktis und im turkestanischen Becken. Entomofauna, 1, 119 - 210." type="journal article" year="1980">Warncke, 1980</bibRefCitation>
).—
<collectingRegion id="8626F86C325CFF94FD72FB24FC5AE614" box="[761,910,331,358]" country="Iran" name="Alborz" pageId="20" pageNumber="502">Alborz prov.</collectingRegion>
: Dam
<quantity id="831A9B6B325CFF94FC5EFB24FBC8E614" box="[981,1052,331,357]" metricMagnitude="4" metricUnit="m" metricValue="2.3" pageId="20" pageNumber="502" unit="km" value="23.0">23 km</quantity>
N Karaj,
<quantity id="831A9B6B325CFF94FB01FB24FB34E614" box="[1162,1248,331,357]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="20" pageNumber="502" unit="m" value="1700.0">1700 m</quantity>
,
<specimenCount id="52E4FD07325CFF94FB67FB24FAC4E614" box="[1260,1296,331,357]" pageId="20" pageNumber="502" type="male">1♂</specimenCount>
,
<date id="305C104E325CFF94FA96FB25FA48E614" box="[1309,1436,330,357]" pageId="20" pageNumber="502" value="1965-07-08">08.vii.1965</date>
(
<bibRefCitation id="20734B7F325CFF94FF14FB00FE98E6F8" author="Warncke, K." box="[159,332,367,393]" pageId="20" pageNumber="502" pagination="119 - 210" refId="ref18439" refString="Warncke, K. (1980) Die Bienengattung Anthidium Fabricius, 1804 in der Westpalaarktis und im turkestanischen Becken. Entomofauna, 1, 119 - 210." type="journal article" year="1980">Warncke, 1980</bibRefCitation>
).—
<emphasis id="7696EA9C325CFF94FEFCFB01FDCDE6F9" bold="true" box="[375,537,366,392]" pageId="20" pageNumber="502">
<collectingCountry id="3CF5761E325CFF94FEFCFB01FDC0E6F9" box="[375,532,366,392]" name="Morocco" pageId="20" pageNumber="502">MOROCCO</collectingCountry>
:
</emphasis>
Tarondona [location not identified],
<specimenCount id="52E4FD07325CFF94FC37FB00FC34E6F8" box="[956,992,367,393]" pageId="20" pageNumber="502" type="male">1♂</specimenCount>
,
<date id="305C104E325CFF94FC65FB00FBBCE6F8" box="[1006,1128,367,393]" pageId="20" pageNumber="502" value="1968-06-21">21.vi.1968</date>
, Verhoeff leg., Leiden Mu- seum (van der
<bibRefCitation id="20734B7F325CFF94FEB6FBFCFE0FE6DF" author="Zanden, G. van der" box="[317,475,403,430]" pageId="20" pageNumber="502" pagination="387 - 390" refId="ref18555" refString="Zanden, G. van der (1996) Neue Verbreitungsangaben zu einigen wenig bekannten palaarktischen Bienen-Arten (Insecta, Hymenoptera, Apoidea). Linzer biologische Beitrage, 28, 387 - 390." type="journal article" year="1996">Zanden, 1996</bibRefCitation>
). Record not regarded as reliable (
<taxonomicName id="83E24D0D325CFF94FCD6FBFCFC29E6DC" authorityName="Kasparek" authorityYear="2021" box="[861,1021,403,429]" class="Magnoliopsida" family="Asteraceae" genus="Anthidum" kingdom="Plantae" order="Asterales" pageId="20" pageNumber="502" phylum="Tracheophyta" rank="species" species="occidentale">
<emphasis id="7696EA9C325CFF94FCD6FBFCFC29E6DC" box="[861,1021,403,429]" italics="true" pageId="20" pageNumber="502">A. occidentale</emphasis>
</taxonomicName>
?).—
<emphasis id="7696EA9C325CFF94FBBDFBFCFB60E6DC" bold="true" box="[1078,1204,403,429]" pageId="20" pageNumber="502">
<collectingCountry id="3CF5761E325CFF94FBBDFBFCFB64E6DC" box="[1078,1200,403,429]" name="Turkey" pageId="20" pageNumber="502">TURKEY</collectingCountry>
:
</emphasis>
<collectingRegion id="8626F86C325CFF94FB30FBFCFA8EE6DC" box="[1211,1370,403,429]" country="Turkey" name="Hakkari" pageId="20" pageNumber="502">Hakkari prov.</collectingRegion>
: Yüksekova,
<quantity id="831A9B6B325CFF94FF79FBD8FE9CE6A0" box="[242,328,439,465]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="502" unit="m" value="2000.0">2000 m</quantity>
,
<specimenCount id="52E4FD07325CFF94FED8FBD8FEA2E6A0" box="[339,374,439,465]" pageId="20" pageNumber="502" type="female">1♀</specimenCount>
,
<date id="305C104E325CFF94FE0FFBD8FDDEE6A0" box="[388,522,439,465]" pageId="20" pageNumber="502" value="1973-08-11">11.viii.1973</date>
(
<bibRefCitation id="20734B7F325CFF94FD93FBD8FD12E6A0" author="Warncke, K." box="[536,710,439,465]" pageId="20" pageNumber="502" pagination="119 - 210" refId="ref18439" refString="Warncke, K. (1980) Die Bienengattung Anthidium Fabricius, 1804 in der Westpalaarktis und im turkestanischen Becken. Entomofauna, 1, 119 - 210." type="journal article" year="1980">Warncke, 1980</bibRefCitation>
).
<collectingRegion id="8626F86C325CFF94FD5CFBD8FCCBE6A0" box="[727,799,439,465]" country="Turkey" name="Afyonkarahisar" pageId="20" pageNumber="502">Afyon</collectingRegion>
Karahisar prov.: Bolvadin,
<quantity id="831A9B6B325CFF94FBD9FBD9FB40E6A0" box="[1106,1172,438,465]" metricMagnitude="2" metricUnit="m" metricValue="9.85" pageId="20" pageNumber="502" unit="m" value="985.0">985m</quantity>
,
<date id="305C104E325CFF94FB14FBD8FACEE6A0" box="[1183,1306,439,465]" pageId="20" pageNumber="502" value="2006-06-23">23.vi.2006</date>
,
<specimenCount id="52E4FD07325CFF94FAAFFBD8FA93E6A0" box="[1316,1351,439,465]" pageId="20" pageNumber="502" type="female">1♀</specimenCount>
(
<bibRefCitation id="20734B7F325CFF94FADEFBD8FF1AE687" author="Guler, Y." pageId="20" pageNumber="502" pagination="731 - 746" refId="ref17159" refString="Guler, Y. (2011) The Wild Bee Fauna of Afyonkarahisar Province: Andrenidae, Anthophoridae and Megachilidae (Hymenoptera: Apoidea). Linzer biologische Beitrage, 43, 731 - 746." type="journal article" year="2011">Güler, 2011</bibRefCitation>
).—
<collectingRegion id="8626F86C325CFF94FF7CFBB4FE96E687" box="[247,322,475,502]" country="Turkey" name="Afyonkarahisar" pageId="20" pageNumber="502">Afyon</collectingRegion>
Karahisar prov.: Sandıklı Daylık,
<quantity id="831A9B6B325CFF94FD48FBB5FCD1E684" box="[707,773,474,501]" metricMagnitude="2" metricUnit="m" metricValue="9.85" pageId="20" pageNumber="502" unit="m" value="985.0">985m</quantity>
,
<date id="305C104E325CFF94FC9BFBB4FC5FE684" box="[784,907,475,501]" pageId="20" pageNumber="502" value="2007-06-26">26.vi.2007</date>
,
<specimenCount id="52E4FD07325CFF94FC1DFBB4FC6DE684" box="[918,953,475,501]" pageId="20" pageNumber="502" type="female">1♀</specimenCount>
(
<bibRefCitation id="20734B7F325CFF94FC43FBB4FB86E684" author="Guler, Y." box="[968,1106,475,501]" pageId="20" pageNumber="502" pagination="731 - 746" refId="ref17159" refString="Guler, Y. (2011) The Wild Bee Fauna of Afyonkarahisar Province: Andrenidae, Anthophoridae and Megachilidae (Hymenoptera: Apoidea). Linzer biologische Beitrage, 43, 731 - 746." type="journal article" year="2011">Güler, 2011</bibRefCitation>
).
</paragraph>
<paragraph id="445D368E325CFF94FF4CFB90FACCE510" blockId="20.[151,1437,150,2014]" pageId="20" pageNumber="502">
<materialsCitation id="F48A3CD3325CFF94FF4CFB90FAC7E510" country="Iran" latitude="35.25" location="The" longLatPrecision="1194" longitude="51.733334" pageId="20" pageNumber="502" specimenCount="1">
<bibRefCitation id="20734B7F325CFF94FF4CFB90FE64E568" author="Nadimi, A. &amp; Talebi, A. A. &amp; Zhu, C. - D. &amp; Fathipour, Y." box="[199,432,511,537]" pageId="20" pageNumber="502" pagination="413 - 424" refId="ref18189" refString="Nadimi, A., Talebi, A. A., Zhu, C. - D. &amp; Fathipour, Y. (2014) Study of the tribe Anthidiini (Hymenoptera: Megachilidae) in northern Iran, with the description of a new species. North-Western Journal of Zoology, 10, 413 - 424." type="journal article" year="2014">
Nadimi
<emphasis id="7696EA9C325CFF94FEAFFB90FE89E568" box="[292,349,511,537]" italics="true" pageId="20" pageNumber="502">et al.</emphasis>
(2014)
</bibRefCitation>
also mention Peykanshahr (
<geoCoordinate id="21D65049325CFF94FD78FB91FC82E568" box="[755,854,510,537]" degrees="35" direction="north" minutes="44" orientation="latitude" pageId="20" pageNumber="502" precision="925" value="35.733334">35°44’N</geoCoordinate>
,
<geoCoordinate id="21D65049325CFF94FCE8FB91FC16E568" box="[867,962,510,537]" degrees="51" direction="east" minutes="09" orientation="longitude" pageId="20" pageNumber="502" precision="925" value="51.15">51°09’E</geoCoordinate>
) and Pishva-Mohammad Abad (
<geoCoordinate id="21D65049325CFF94FABFFB91FA43E568" box="[1332,1431,510,537]" degrees="35" direction="north" minutes="15" orientation="latitude" pageId="20" pageNumber="502" precision="925" value="35.25">35°15’N</geoCoordinate>
,
<geoCoordinate id="21D65049325CFF94FF1CF84DFF21E54C" box="[151,245,546,573]" degrees="51" direction="east" minutes="44" orientation="longitude" pageId="20" pageNumber="502" precision="925" value="51.733334">51°44’E</geoCoordinate>
) in northern
<collectingCountry id="3CF5761E325CFF94FE00F84CFE6CE54C" box="[395,440,547,573]" name="Iran" pageId="20" pageNumber="502">Iran</collectingCountry>
as “
<taxonomicName id="83E24D0D325CFF94FE67F84CFD4AE54C" authorityName="Warncke" authorityYear="1980" box="[492,670,547,573]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="502" phylum="Arthropoda" rank="species" species="rotundum">
<emphasis id="7696EA9C325CFF94FE67F84CFDD1E54C" box="[492,517,547,573]" italics="true" pageId="20" pageNumber="502">A.</emphasis>
cf.
<emphasis id="7696EA9C325CFF94FDB9F84CFD4AE54C" box="[562,670,547,573]" italics="true" pageId="20" pageNumber="502">rotundum</emphasis>
</taxonomicName>
”. A male in OLL is labelled as “
<taxonomicName id="83E24D0D325CFF94FB9AF84CFB59E54C" authorityName="Fabricius" authorityYear="1804" box="[1041,1165,547,573]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="502" phylum="Arthropoda" rank="genus">Anthidium</taxonomicName>
flavitarse. Serbien: Rndow”, without date or collector.
<location id="413D6055325CFF94FD8BF828FDF8E510" LSID="urn:lsid:plazi:treatment:CC4B8798325BFF95FF1CFE69FE3FE614:413D6055325CFF94FD8BF828FDF8E510" box="[512,556,583,609]" country="Iran" latitude="35.25" longLatPrecision="1194" longitude="51.733334" name="The" pageId="20" pageNumber="502">The</location>
locality could not be identified and the record needs confirmation
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0CF86505325CFF94FF4CF805FAC7E4D4" pageId="20" pageNumber="502" type="diagnosis">
<paragraph id="445D368E325CFF94FF4CF805FB12E464" blockId="20.[151,1437,150,2014]" pageId="20" pageNumber="502">
<emphasis id="7696EA9C325CFF94FF4CF805FE96E5F5" bold="true" box="[199,322,618,644]" pageId="20" pageNumber="502">Diagnosis.</emphasis>
The female is characterised by a deeply concave vertex (moderately concave in the other species) which is slightly curled up in lateral view (lamellate in
<taxonomicName id="83E24D0D325CFF94FC9BF8E0FC17E5D8" authorityName="Kasparek" authorityYear="2021" box="[784,963,655,681]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="502" phylum="Arthropoda" rank="species" species="preoccipitale">
<emphasis id="7696EA9C325CFF94FC9BF8E0FC17E5D8" box="[784,963,655,681]" italics="true" pageId="20" pageNumber="502">A. preoccipitale</emphasis>
</taxonomicName>
, rounded to angular in the other species). Clypeus is flat with a broad (approx. one antenna diameter wide) impunctate middle line (shared with
<taxonomicName id="83E24D0D325CFF94FA8DF8DCFF61E581" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="502" phylum="Arthropoda" rank="species" species="anguliventre">
<emphasis id="7696EA9C325CFF94FA8DF8DCFF61E581" italics="true" pageId="20" pageNumber="502">A. anguliventre</emphasis>
</taxonomicName>
; narrower middle line in the other species). Terga with a narrow (1–2 puncture diameter wide) smooth marginal zone (punctation reaching apical margin in all other species). See
<tableCitation id="09600335325CFF94FCF3F894FC1FE464" box="[888,971,763,789]" captionStart="TABLE 1" captionStartId="7.[151,244,834,860]" captionText="TABLE 1. Comparison of some structural traits in the females of five species Anthidium, subgenus Gulanthidium. The female of A. preoccipitale is not known. Highly relevant distinguishing characters are given in bold." pageId="20" pageNumber="502">Table 1</tableCitation>
for further characters.
</paragraph>
<paragraph id="445D368E325CFF94FF4CF970FAC7E4D4" blockId="20.[151,1437,150,2014]" pageId="20" pageNumber="502">
The male of
<taxonomicName id="83E24D0D325CFF94FEDDF970FE35E448" authorityName="Warncke" authorityYear="1980" box="[342,481,799,825]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="502" phylum="Arthropoda" rank="species" species="rotundum">
<emphasis id="7696EA9C325CFF94FEDDF970FE35E448" box="[342,481,799,825]" italics="true" pageId="20" pageNumber="502">A. rotundum</emphasis>
</taxonomicName>
is well characterised by the semicircular T6, not flattened or emarginated at the apex as in the other species of the subgenus, and an angulate vertex (lamellate or rounded in the other species). Yellow band on T1 mostly broadly interrupted (uninterrupted in the other species).
<taxonomicName id="83E24D0D325CFF94FC57F908FB17E4F0" authorityName="Warncke" authorityYear="1980" box="[988,1219,871,897]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="502" phylum="Arthropoda" rank="species" species="rotundum">
<emphasis id="7696EA9C325CFF94FC57F908FB17E4F0" box="[988,1219,871,897]" italics="true" pageId="20" pageNumber="502">Anthidium rotundum</emphasis>
</taxonomicName>
has a characteristic S6 with a median longitudinal furrow.
<taxonomicName id="83E24D0D325CFF94FDC3F9E4FCE4E4D4" authorityName="Warncke" authorityYear="1980" box="[584,816,907,933]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="502" phylum="Arthropoda" rank="species" species="rotundum">
<emphasis id="7696EA9C325CFF94FDC3F9E4FCE4E4D4" box="[584,816,907,933]" italics="true" pageId="20" pageNumber="502">Anthidium rotundum</emphasis>
</taxonomicName>
is on average larger than the other species.
</paragraph>
</subSubSection>
<subSubSection id="0CF86505325CFF94FF4CF9C1FA5EE0C8" pageId="20" pageNumber="502" type="description">
<paragraph id="445D368E325CFF94FF4CF9C1FE0AE208" blockId="20.[151,1437,150,2014]" pageId="20" pageNumber="502">
<emphasis id="7696EA9C325CFF94FF4CF9C1FE69E4B8" bold="true" box="[199,445,942,969]" pageId="20" pageNumber="502">Description. Female.</emphasis>
<quantity id="831A9B6B325CFF94FE48F9C0FDE5E4B8" box="[451,561,943,969]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.2" metricValueMin="0.9" pageId="20" pageNumber="502" unit="mm" value="10.5" valueMax="12.0" valueMin="9.0">9–12 mm</quantity>
.
<emphasis id="7696EA9C325CFF94FDB1F9C0FD54E4B8" box="[570,640,943,969]" italics="true" pageId="20" pageNumber="502">Head:</emphasis>
Clypeus (
<figureCitation id="DCD92A0B325CFF94FD7AF9C0FC9DE4B8" box="[753,841,943,969]" captionStart="FIGURE 7" captionStartId="10.[151,250,1395,1419]" captionTargetBox="[152,1435,181,1371]" captionTargetId="figure-15@10.[152,1435,181,1371]" captionTargetPageId="10" captionText="FIGURE 7. Faces of the females of Anthidium anguliventre (A), A. eremicum (B), A. occidentale (C), and A. rotundum (D)." figureDoi="http://doi.org/10.5281/zenodo.5531649" httpUri="https://zenodo.org/record/5531649/files/figure.png" pageId="20" pageNumber="502">Fig. 7D</figureCitation>
) flat with a broad almost entirely impunctate, smooth middle band; anterior and posterior margins straight, posterior margin smooth and bulging, and with a narrow, semitransparent light brown apex; supraclypeal area yellow, with scattered punctation; paraocular area yellow, with a broad band almost reaching top of eye; mandible yellow with five dark brown teeth, and sometimes an additional minute tooth distal to the uppermost; vertex slightly curled up, preoccipital ridge angular; vertex in dorsal view deeply concave (
<figureCitation id="DCD92A0B325CFF94FEDDFE0CFE6AE30C" box="[342,446,1123,1149]" captionStart="FIGURE 10" captionStartId="13.[151,250,1274,1298]" captionTargetBox="[284,1304,188,1241]" captionTargetId="figure-39@13.[277,1311,181,1250]" captionTargetPageId="13" captionText="FIGURE 10. Schematic view of the head in the species of Anthidium, subgenus Gulanthidium. A. Lateral view; B. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5531655" httpUri="https://zenodo.org/record/5531655/files/figure.png" pageId="20" pageNumber="502">Fig. 10B</figureCitation>
); yellow preoccipital band; gena yellow except for some black on the inner side.—
<taxonomicName id="83E24D0D325CFF94FAE4FE0CFEEAE3D0" authority=": Black" authorityName="Black" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="20" pageNumber="502" phylum="Arthropoda" rank="genus">
<emphasis id="7696EA9C325CFF94FAE4FE0CFF20E3D1" italics="true" pageId="20" pageNumber="502">Mesosoma:</emphasis>
Black
</taxonomicName>
with broad yellow, reverse U-shaped band; mesepisternum yellow with very low omaular lamella; pronotal lobe yellow with high lamella; scutellum widely rounded in dorsal view with posterior semi-transparent margin laterally; posterior margin of axilla set back against posterior margin of scutellum; scutellum and axilla yellow, with black base of scutellum.—
<emphasis id="7696EA9C325CFF94FDD6FE9CFD0FE27D" box="[605,731,1267,1292]" italics="true" pageId="20" pageNumber="502">Metasoma:</emphasis>
Discs of T1–T5 yellow, depressions black; punctation on depressions finer than on discs, punctures leave an impunctate posterior margin; T6 yellow (except of some black laterally), depressed as seen in profile; small sting-wide incision at apex.—
<emphasis id="7696EA9C325CFF94FC62FF54FBCAE225" box="[1001,1054,1339,1364]" italics="true" pageId="20" pageNumber="502">Legs</emphasis>
: Yellow; hind tibia rounded; hind basitarsus with strong carina.
</paragraph>
<paragraph id="445D368E325CFF94FF4CFFECFA5EE0C8" blockId="20.[151,1437,150,2014]" pageId="20" pageNumber="502">
<emphasis id="7696EA9C325CFF94FF4CFFECFEDFE2EC" bold="true" box="[199,267,1411,1437]" pageId="20" pageNumber="502">Male.</emphasis>
<quantity id="831A9B6B325CFF94FE99FFECFE54E2EC" box="[274,384,1411,1437]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.4" metricValueMin="0.9" pageId="20" pageNumber="502" unit="mm" value="11.5" valueMax="14.0" valueMin="9.0">9–14 mm</quantity>
.
<emphasis id="7696EA9C325CFF94FE00FFECFE05E2EC" box="[395,465,1411,1437]" italics="true" pageId="20" pageNumber="502">Head:</emphasis>
Clypeus flat with straight basal and lateral margins; broadly impunctate medially, apical margin with a thin semi-transparent end line; very shallowly emarginate; dense punctation laterally; mandible yellow with 3 black teeth; supraclypeal area yellow with scattered punctation; paraocular area yellow, with yellow maculation reaching upper end of eye; vertex strongly concave in dorsal view with angulate margin (
<figureCitation id="DCD92A0B325CFF94FAADFF80FA5AE178" box="[1318,1422,1519,1545]" captionStart="FIGURE 10" captionStartId="13.[151,250,1274,1298]" captionTargetBox="[284,1304,188,1241]" captionTargetId="figure-39@13.[277,1311,181,1250]" captionTargetPageId="13" captionText="FIGURE 10. Schematic view of the head in the species of Anthidium, subgenus Gulanthidium. A. Lateral view; B. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5531655" httpUri="https://zenodo.org/record/5531655/files/figure.png" pageId="20" pageNumber="502">Fig. 10B</figureCitation>
); yellow preoccipital band reaching mandibles; preoccipital band sometimes interrupted medially with some yellow remnants in the middle; gena broad, broader than eye in lateral view; scape dark yellow; proximal segments of antenna ochreous.—
<taxonomicName id="83E24D0D325CFF94FEEFFC34FD97E104" authority=": Scutum" authorityName="Scutum" box="[356,579,1627,1653]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="20" pageNumber="502" phylum="Arthropoda" rank="genus">
<emphasis id="7696EA9C325CFF94FEEFFC34FE31E105" box="[356,485,1627,1652]" italics="true" pageId="20" pageNumber="502">Mesosoma:</emphasis>
Scutum
</taxonomicName>
black with yellow, inverted U-shaped yellow band; inner arm much thinner than lateral arm; scutellum widely rounded in dorsal view, somewhat overhanging propodeum; apical margin emarginate in the centre; scutellum and axillae yellow except at the base of scutellum; pronotal lobe yellow with high lamella; mesepisternum with yellow maculation; upper part of omaulus slightly carinate, but hardly visible because of dense white pubescence.—
<emphasis id="7696EA9C325CFF94FE61FC84FDBCE075" box="[490,616,1771,1796]" italics="true" pageId="20" pageNumber="502">Metasoma:</emphasis>
T1–T6 with yellow bands, mostly interrupted on T1, otherwise uninterrupted; T2 bulged laterally, T3 with rounded lateral projection, T4–T6 with acute projections; apical margin of T6 bulged; T7 yellow, semicircular, with fine longitudinal carina medially and transparent apical margin (
<figureCitation id="DCD92A0B325CFF94FA8DFD5CFA8BE03C" box="[1286,1375,1843,1869]" captionStart="FIGURE 8" captionStartId="11.[151,250,945,969]" captionTargetBox="[158,1429,181,920]" captionTargetId="figure-39@11.[158,1429,181,921]" captionTargetPageId="11" captionText="FIGURE 8. Apical terga of the male of five species of Anthidium, subgenus Gulanthidium: A. Anthidium anguliventre; B. A. eremicum; C. A. occidentale sp. nov.; D. A. preoccipitale sp. nov.; E. A. rotundum." figureDoi="http://doi.org/10.5281/zenodo.5531651" httpUri="https://zenodo.org/record/5531651/files/figure.png" pageId="20" pageNumber="502">Figs 8E</figureCitation>
,
<figureCitation id="DCD92A0B325CFF94FAE1FD5CFA5AE03C" box="[1386,1422,1843,1869]" captionStart="FIGURE 9" captionStartId="11.[151,250,1689,1713]" captionTargetBox="[168,1418,1036,1652]" captionTargetId="figure-81@11.[162,1425,1029,1665]" captionTargetPageId="11" captionText="FIGURE 9. Male of Anthidium, subgenus Gulanthidium. Apical terga (upper row) in dorsal and sternum 6 (S6) in ventral view. A + D. Anthidium anguliventre; B + E. A. eremicum and A. occidentale; C + F. A. preoccipitale and A. rotundum." figureDoi="http://doi.org/10.5281/zenodo.5531653" httpUri="https://zenodo.org/record/5531653/files/figure.png" pageId="20" pageNumber="502">9C</figureCitation>
); S6 semicircular with median furrow, broadening towards the apex, and a lateral depression (
<figureCitation id="DCD92A0B325CFF94FB19FD38FB30E000" box="[1170,1252,1879,1905]" captionStart="FIGURE 9" captionStartId="11.[151,250,1689,1713]" captionTargetBox="[168,1418,1036,1652]" captionTargetId="figure-81@11.[162,1425,1029,1665]" captionTargetPageId="11" captionText="FIGURE 9. Male of Anthidium, subgenus Gulanthidium. Apical terga (upper row) in dorsal and sternum 6 (S6) in ventral view. A + D. Anthidium anguliventre; B + E. A. eremicum and A. occidentale; C + F. A. preoccipitale and A. rotundum." figureDoi="http://doi.org/10.5281/zenodo.5531653" httpUri="https://zenodo.org/record/5531653/files/figure.png" pageId="20" pageNumber="502">Fig. 9F</figureCitation>
).—
<emphasis id="7696EA9C325CFF94FA84FD38FA9AE001" box="[1295,1358,1879,1904]" italics="true" pageId="20" pageNumber="502">Legs:</emphasis>
Femora with much black maculation; tibiae yellow with black inner face; tarsi yellow.—
<emphasis id="7696EA9C325CFF94FBBEFD14FAACE0E4" box="[1077,1400,1915,1941]" italics="true" pageId="20" pageNumber="502">Hidden sterna and genitalia:</emphasis>
S7 elongate with hairy apex; S8 with triangular apex; apex of penis valves spearhead-like. See also separate chapter.
</paragraph>
</subSubSection>
<subSubSection id="0CF86505325CFF95FF4CFDADFBEEE7A5" lastPageId="21" lastPageNumber="503" pageId="20" pageNumber="502" type="biology_ecology">
<paragraph id="445D368E325CFF95FF4CFDADFBEEE7A5" blockId="20.[151,1437,150,2014]" lastBlockId="21.[151,1436,150,357]" lastPageId="21" lastPageNumber="503" pageId="20" pageNumber="502">
<emphasis id="7696EA9C325CFF94FF4CFDADFEF3E0AD" bold="true" box="[199,295,1986,2012]" pageId="20" pageNumber="502">Biology.</emphasis>
All records are from June and July. According to
<bibRefCitation id="20734B7F325CFF94FCE2FDACFBDFE0AF" author="Muller, A." box="[873,1035,1987,2014]" pageId="20" pageNumber="502" pagination="235 - 257" refId="ref18149" refString="Muller, A. (1996) Host-Plant Specialization in Western Palearctic Anthidine Bees (Hymenoptera: Apoidea: Megachilidae. Ecological Monographs, 66, 235 - 257. https: // doi. org / 10.2307 / 2963476" type="journal article" year="1996">Müller (1996)</bibRefCitation>
,
<taxonomicName id="83E24D0D325CFF94FB92FDACFACDE0AC" authority="" box="[1049,1305,1987,2013]" class="Magnoliopsida" family="Asteraceae" genus="Cardueae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Asterales" pageId="20" pageNumber="502" phylum="Tracheophyta" rank="subGenus" subGenus="Asteraceae">Cardueae (Asteraceae)</taxonomicName>
are the ex- clusive pollen sources of
<taxonomicName id="83E24D0D325DFF95FE36FAF8FD9FE7C1" authorityName="Warncke" authorityYear="1980" box="[445,587,150,177]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="503" phylum="Arthropoda" rank="species" species="rotundum">
<emphasis id="7696EA9C325DFF95FE36FAF8FD9FE7C1" box="[445,587,150,177]" italics="true" pageId="21" pageNumber="503">A. rotundum</emphasis>
</taxonomicName>
. The species was collected in
<collectingCountry id="3CF5761E325DFF95FC27FAF9FBDAE7C1" box="[940,1038,150,176]" name="Lebanon" pageId="21" pageNumber="503">Lebanon</collectingCountry>
from
<taxonomicName id="83E24D0D325DFF95FBDCFAF8FB22E7C0" box="[1111,1270,151,177]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="21" pageNumber="503" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="7696EA9C325DFF95FBDCFAF8FB18E7C0" box="[1111,1228,151,177]" italics="true" pageId="21" pageNumber="503">Centaurea</emphasis>
sp.
</taxonomicName>
(M. Boustani, unpubl.), and in
<collectingCountry id="3CF5761E325DFF95FEDBFAD4FEA9E7A4" box="[336,381,187,213]" name="Iran" pageId="21" pageNumber="503">Iran</collectingCountry>
from
<taxonomicName id="83E24D0D325DFF95FE4AFAD4FBE2E7A5" authority="(Dehghan Dehnavi et al., 2014)" baseAuthorityName="Dehghan Dehnavi" baseAuthorityYear="2014" box="[449,1078,186,213]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="21" pageNumber="503" phylum="Tracheophyta" rank="species" species="bruguierana">
<emphasis id="7696EA9C325DFF95FE4AFAD4FD1DE7A5" box="[449,713,186,213]" italics="true" pageId="21" pageNumber="503">Centaurea bruguierana</emphasis>
(
<bibRefCitation id="20734B7F325DFF95FD53FAD5FBFAE7A5" author="Dehghan Dehnavi, L. &amp; Talebi, A. A. &amp; Goldasteh, Sh. &amp; Vafaei Shooshtari, R. &amp; Nadimi, A." box="[728,1070,186,213]" pageId="21" pageNumber="503" pagination="1 - 21" refId="ref16822" refString="Dehghan Dehnavi, L., Talebi, A. A., Goldasteh, Sh., Vafaei Shooshtari, R. &amp; Nadimi, A. (2014) Contribution to the knowledge of Megachilidae (Hymenoptera: Apoidea) in the Yazd province, Iran. Journal of Entomological Research, 7, 1 - 21." type="journal article" year="2014">
Dehghan Dehnavi
<emphasis id="7696EA9C325DFF95FC20FAD4FC30E7A5" box="[939,996,186,212]" italics="true" pageId="21" pageNumber="503">et al.</emphasis>
, 2014
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="0CF86505325DFF95FF4CFAB1FE3FE614" pageId="21" pageNumber="503" type="distribution">
<paragraph id="445D368E325DFF95FF4CFAB1FE3FE614" blockId="21.[151,1436,150,357]" pageId="21" pageNumber="503">
<emphasis id="7696EA9C325DFF95FF4CFAB1FEB5E789" bold="true" box="[199,353,222,248]" pageId="21" pageNumber="503">Distribution.</emphasis>
Known from
<collectingCountry id="3CF5761E325DFF95FD82FAB1FD8FE789" box="[521,603,222,248]" name="Greece" pageId="21" pageNumber="503">Greece</collectingCountry>
,
<collectingCountry id="3CF5761E325DFF95FDE1FAB0FD4EE788" box="[618,666,223,249]" name="Iran" pageId="21" pageNumber="503">Iran</collectingCountry>
,
<collectingCountry id="3CF5761E325DFF95FD22FAB1FCDAE789" box="[681,782,222,248]" name="Lebanon" pageId="21" pageNumber="503">Lebanon</collectingCountry>
,
<collectingCountry id="3CF5761E325DFF95FC96FAB1FC8FE789" box="[797,859,222,248]" name="Syria" pageId="21" pageNumber="503">Syria</collectingCountry>
,
<collectingCountry id="3CF5761E325DFF95FCE2FAB1FC6EE789" box="[873,954,222,248]" name="Turkey" pageId="21" pageNumber="503">Turkey</collectingCountry>
, and
<collectingCountry id="3CF5761E325DFF95FC70FAB1FBA2E789" box="[1019,1142,222,248]" name="Azerbaijan" pageId="21" pageNumber="503">Azerbaijan</collectingCountry>
(
<figureCitation id="DCD92A0B325DFF95FB02FAB1FB34E788" box="[1161,1248,222,249]" captionStart="FIGURE 16" captionStartId="21.[151,250,1125,1149]" captionTargetBox="[163,1425,380,1100]" captionTargetId="figure-125@21.[163,1425,380,1100]" captionTargetPageId="21" captionText="FIGURE 16. Distribution of Anthidium rotundum. Records in Egypt need confirmation." figureDoi="http://doi.org/10.5281/zenodo.5531667" httpUri="https://zenodo.org/record/5531667/files/figure.png" pageId="21" pageNumber="503">Fig. 16</figureCitation>
). A record from
<collectingCountry id="3CF5761E325DFF95FF1CFB6CFF28E66C" box="[151,252,259,285]" name="Morocco" pageId="21" pageNumber="503">Morocco</collectingCountry>
(van den
<bibRefCitation id="20734B7F325DFF95FEFFFB6DFDC5E66C" author="Zanden, G. van der" box="[372,529,258,285]" pageId="21" pageNumber="503" pagination="387 - 390" refId="ref18555" refString="Zanden, G. van der (1996) Neue Verbreitungsangaben zu einigen wenig bekannten palaarktischen Bienen-Arten (Insecta, Hymenoptera, Apoidea). Linzer biologische Beitrage, 28, 387 - 390." type="journal article" year="1996">Zanden, 1996</bibRefCitation>
) seems to be unreliable and records from
<collectingCountry id="3CF5761E325DFF95FC76FB6CFB94E66C" box="[1021,1088,259,285]" name="Egypt" pageId="21" pageNumber="503">Egypt</collectingCountry>
(van den
<bibRefCitation id="20734B7F325DFF95FB3CFB6DFA81E66C" author="Zanden, G. van der" box="[1207,1365,258,285]" pageId="21" pageNumber="503" pagination="387 - 390" refId="ref18555" refString="Zanden, G. van der (1996) Neue Verbreitungsangaben zu einigen wenig bekannten palaarktischen Bienen-Arten (Insecta, Hymenoptera, Apoidea). Linzer biologische Beitrage, 28, 387 - 390." type="journal article" year="1996">Zanden, 1996</bibRefCitation>
) need confirmation. The distribution in the west extends to Lesbos (northern Aegean;
<bibRefCitation id="20734B7F325DFF95FB9EFB49FB75E631" author="Grace, A." box="[1045,1185,294,320]" pageId="21" pageNumber="503" refId="ref17132" refString="Grace, A. (2010) Introductory biogeography to bees of the Eastern Mediterranean and Near East. Bexhill Museum, Sussex, 284 pp." type="book" year="2010">Grace, 2010</bibRefCitation>
) and western
<collectingCountry id="3CF5761E325DFF95FAB7FB49FA48E631" box="[1340,1436,294,320]" name="Turkey" pageId="21" pageNumber="503">Anatolia</collectingCountry>
(
<collectingRegion id="8626F86C325DFF95FF14FB25FF09E615" box="[159,221,330,356]" country="Turkey" name="Izmir" pageId="21" pageNumber="503">Izmir</collectingRegion>
and
<collectingRegion id="8626F86C325DFF95FE9FFB25FE8BE614" box="[276,351,330,357]" country="Turkey" name="Mugla" pageId="21" pageNumber="503">Muğla</collectingRegion>
) provinces).
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="8F22CBA223FF3F76839FA6C31492E1E9" ID-CLB-Dataset="39150" ID-DOI="10.11646/zootaxa.4107.3.8" ID-GBIF-Dataset="a9ae982c-e6f3-4876-9600-dbb53ab224aa" ID-ISSN="1175-5326" ID-Zenodo-Dep="255951" ID-ZooBank="FF41E034-0EDA-410D-940E-F67D949D4A2C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1462257912970" checkinUser="plazi" docAuthor="Huang, Hao" docDate="2016" docId="CC4BB639EB2800666E80C3DD4F7B8E91" docLanguage="en" docName="zootaxa.4107.3.8.pdf.imf" docOrigin="Zootaxa 4107 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Shaanxiana langi Huang, 2016, sp. n." docType="treatment" docVersion="9" lastPageNumber="411" masterDocId="3072CE41EB29006E6E17C34A4E418C16" masterDocTitle="Discovery of a fourth species of Shaanxiana (Lepidoptera: Lycaenidae: Theclinae) from China" masterLastPageNumber="412" masterPageNumber="403" pageNumber="404" updateTime="1698662204794" updateUser="ExternalLinkService">
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<mods:title id="DB73C91A63BD0AA1912887E2F6B5CAB8">Discovery of a fourth species of Shaanxiana (Lepidoptera: Lycaenidae: Theclinae) from China</mods:title>
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<mods:namePart id="5D7418B531EDCA370AF5212DA450C591">Huang, Hao</mods:namePart>
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<mods:date id="35638E2F4B17E723611AFC6DDFA27DE9">2016</mods:date>
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<treatment id="CC4BB639EB2800666E80C3DD4F7B8E91" ID-DOI="http://doi.org/10.5281/zenodo.6086665" ID-GBIF-Taxon="120620776" ID-Zenodo-Dep="6086665" LSID="urn:lsid:plazi:treatment:CC4BB639EB2800666E80C3DD4F7B8E91" httpUri="http://treatment.plazi.org/id/CC4BB639EB2800666E80C3DD4F7B8E91" lastPageId="8" lastPageNumber="411" pageId="1" pageNumber="404">
<subSubSection id="0CF854A4EB28006F6E80C3DD4C2C8CC5" pageId="1" pageNumber="404" type="nomenclature">
<paragraph id="445D072FEB28006F6E80C3DD4FF98CA4" blockId="1.[151,621,151,211]" box="[151,440,151,178]" pageId="1" pageNumber="404">
<heading id="1F15B043EB28006F6E80C3DD4FF98CA4" bold="true" box="[151,440,151,178]" fontSize="11" level="1" pageId="1" pageNumber="404" reason="1">
<emphasis id="7696DB3DEB28006F6E80C3DD4FF98CA4" bold="true" box="[151,440,151,178]" pageId="1" pageNumber="404">
<taxonomicName id="83E27CACEB28006F6E80C3DD4F2E8CA7" box="[151,367,151,177]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="404" phylum="Arthropoda" rank="species" species="langi" status="sp. nov.">
<emphasis id="7696DB3DEB28006F6E80C3DD4F2E8CA7" bold="true" box="[151,367,151,177]" italics="true" pageId="1" pageNumber="404">Shaanxiana langi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6DA56646EB28006F6F61C3D24FF98CA4" box="[374,440,152,178]" pageId="1" pageNumber="404" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="445D072FEB28006F6E80C3F04C2C8CC5" blockId="1.[151,621,151,211]" box="[151,621,186,211]" pageId="1" pageNumber="404">
(
<figureCitation id="DCD91BAAEB28006F6E88C3F04F4A8CC5" box="[159,267,186,211]" captionStart="FIGURES 1 – 8" captionStartId="1.[151,264,1903,1926]" captionTargetBox="[151,1436,391,1880]" captionTargetId="figure@1.[151,1436,383,1881]" captionTargetPageId="1" captionText="FIGURES 1 – 8. Relative size of Shaanxiana species at the same scale (scale bar 1 cm). 1 – 2 — Shaanxiana australis, male holotype (reproduced from Hsu, 2015 with permission by chief editor of Zootaxa); 3 – 4 — Shaanxiana langi, male holotype; 5 – 6 — Shaanxiana langi, female paratype; 7 – 8 — Shaanxiana takashimai; 7 — male from Zhouzhi, Shaanxi; 8 — female from Fengxian, Shaanxi. 1, 3, 5 — upperside; 2, 4, 6 — underside; 7 – 8 — upperside (left half) and underside (right half)." httpUri="https://zenodo.org/record/255952/files/figure.png" pageId="1" pageNumber="404">Figs. 3–6</figureCitation>
,
<figureCitation id="DCD91BAAEB28006F6F01C3F14F648CC5" box="[278,293,187,211]" captionStart="FIGURE 9" captionStartId="2.[151,250,1643,1665]" captionTargetBox="[151,1436,193,1603]" captionTargetId="figure@2.[151,1436,193,1604]" captionTargetPageId="2" captionText="FIGURE 9. Morphological differences between S. langi and S. australis, the numbers of arrows correspond to those of the characters under the Diagnosis heading in the text (a, b at same scale; c, d at same scale; e – i at same scale); a, b—wing-pattern of S. langi, male; c—dorsum of S. australis in flattened shape (reproduced from Hsu, 2015 with permission by chief editor of Zootaxa); d—dorsum of S. langi in dorsal view of natural shape; e—male genitalia of S. langi in lateral view of natural shape; f—left valva of S. australis in flattened shape (reproduced from Hsu, 2015 with permission by chief editor of Zootaxa); g—left valva of S. langi, holotype in flattened shape; h—right valva of S. takashimai in flattened shape (reproduced from Hsu, 2015 with permission by chief editor of Zootaxa); i—right valva of S. takashimai in flattened shape, taken from specimen shown in Fig. 7." httpUri="https://zenodo.org/record/255953/files/figure.png" pageId="1" pageNumber="404">9</figureCitation>
a, 9b, 9d, 9e,
<quantity id="831AAACAEB28006F6FACC3F14F988CC5" box="[443,473,187,211]" metricMagnitude="-3" metricUnit="kg" metricValue="9.0" pageId="1" pageNumber="404" unit="g" value="9.0">9g</quantity>
,, 10, 12, 14)
</paragraph>
</subSubSection>
<subSubSection id="0CF854A4EB28006F6E80C2484D588D75" pageId="1" pageNumber="404" type="materials_examined">
<paragraph id="445D072FEB28006F6E80C2484D588D75" blockId="1.[151,1436,258,355]" pageId="1" pageNumber="404">
<emphasis id="7696DB3DEB28006F6E80C2484FF98D0D" bold="true" box="[151,440,258,283]" pageId="1" pageNumber="404">
<typeStatus id="9B59B98DEB28006F6E80C2484E938D0D" box="[151,210,258,283]" pageId="1" pageNumber="404">Type</typeStatus>
material.
<typeStatus id="9B59B98DEB28006F6F5CC2484FF98D0D" box="[331,440,258,283]" pageId="1" pageNumber="404" type="holotype">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="DCD91BAAEB28006F6FD1C2484C728D0D" box="[454,563,258,283]" captionStart="FIGURES 1 – 8" captionStartId="1.[151,264,1903,1926]" captionTargetBox="[151,1436,391,1880]" captionTargetId="figure@1.[151,1436,383,1881]" captionTargetPageId="1" captionText="FIGURES 1 – 8. Relative size of Shaanxiana species at the same scale (scale bar 1 cm). 1 – 2 — Shaanxiana australis, male holotype (reproduced from Hsu, 2015 with permission by chief editor of Zootaxa); 3 – 4 — Shaanxiana langi, male holotype; 5 – 6 — Shaanxiana langi, female paratype; 7 – 8 — Shaanxiana takashimai; 7 — male from Zhouzhi, Shaanxi; 8 — female from Fengxian, Shaanxi. 1, 3, 5 — upperside; 2, 4, 6 — underside; 7 – 8 — upperside (left half) and underside (right half)." httpUri="https://zenodo.org/record/255952/files/figure.png" pageId="1" pageNumber="404">Figs. 3–4</figureCitation>
):
<emphasis id="7696DB3DEB28006F6C51C2484CE28D0D" bold="true" box="[582,675,258,283]" pageId="1" pageNumber="404">
<collectingCountry id="3CF547BFEB28006F6C51C2484CE28D0D" box="[582,675,258,283]" name="China" pageId="1" pageNumber="404">CHINA</collectingCountry>
</emphasis>
: ♂, Guangxi province, Laibin City, Jinxiu County, Dayaoshan Mts.,
<quantity id="831AAACAEB28006F6E80C26D4EA68D29" box="[151,231,295,319]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="404" unit="m" value="1000.0">1000m</quantity>
,
<date id="305C21EFEB28006F6EE4C26C4F288D29" box="[243,361,294,319]" pageId="1" pageNumber="404" value="2015-05-21">21.V.2015</date>
, Song-Yun Lang leg., to be deposited in
<collectionCode id="22F39FEAEB28006F6D22C26D4DC58D29" box="[821,900,295,319]" pageId="1" pageNumber="404">SNUC</collectionCode>
.
<emphasis id="7696DB3DEB28006F6D99C26C4DBA8D29" bold="true" box="[910,1019,294,319]" pageId="1" pageNumber="404">
<typeStatus id="9B59B98DEB28006F6D99C26C4DBA8D29" box="[910,1019,294,319]" pageId="1" pageNumber="404" type="paratype">Paratype</typeStatus>
</emphasis>
(
<figureCitation id="DCD91BAAEB28006F6A1CC26C4A388D29" box="[1035,1145,294,319]" captionStart="FIGURES 1 – 8" captionStartId="1.[151,264,1903,1926]" captionTargetBox="[151,1436,391,1880]" captionTargetId="figure@1.[151,1436,383,1881]" captionTargetPageId="1" captionText="FIGURES 1 – 8. Relative size of Shaanxiana species at the same scale (scale bar 1 cm). 1 – 2 — Shaanxiana australis, male holotype (reproduced from Hsu, 2015 with permission by chief editor of Zootaxa); 3 – 4 — Shaanxiana langi, male holotype; 5 – 6 — Shaanxiana langi, female paratype; 7 – 8 — Shaanxiana takashimai; 7 — male from Zhouzhi, Shaanxi; 8 — female from Fengxian, Shaanxi. 1, 3, 5 — upperside; 2, 4, 6 — underside; 7 – 8 — upperside (left half) and underside (right half)." httpUri="https://zenodo.org/record/255952/files/figure.png" pageId="1" pageNumber="404">Figs. 5–6</figureCitation>
):
<emphasis id="7696DB3DEB28006F6A99C26C4AAA8D29" bold="true" box="[1166,1259,294,319]" pageId="1" pageNumber="404">
<collectingCountry id="3CF547BFEB28006F6A99C26C4AAA8D29" box="[1166,1259,294,319]" name="China" pageId="1" pageNumber="404">CHINA</collectingCountry>
</emphasis>
: 1♀, same data as
<typeStatus id="9B59B98DEB28006F6EA2C2004F5B8D75" box="[181,282,330,355]" pageId="1" pageNumber="404" type="holotype">holotype</typeStatus>
, deposited in Hao Huang’s private collection.
</paragraph>
</subSubSection>
<caption id="109D57A7EB28006F6E80C4254B498BF7" httpUri="https://zenodo.org/record/255952/files/figure.png" pageId="1" pageNumber="404" targetBox="[151,1436,391,1880]" targetPageId="1">
<paragraph id="445D072FEB28006F6E80C4254B498BF7" blockId="1.[151,1436,1903,2017]" pageId="1" pageNumber="404">
<emphasis id="7696DB3DEB28006F6E80C4254F7C8B92" bold="true" box="[151,317,1903,1926]" pageId="1" pageNumber="404">FIGURES 1–8.</emphasis>
Relative size of
<taxonomicName id="83E27CACEB28006F6FE0C4254C2E8B93" box="[503,623,1903,1925]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="404" phylum="Arthropoda" rank="genus">
<emphasis id="7696DB3DEB28006F6FE0C4254C2E8B93" box="[503,623,1903,1925]" italics="true" pageId="1" pageNumber="404">Shaanxiana</emphasis>
</taxonomicName>
species at the same scale (scale bar 1cm). 1–2—
<taxonomicName id="83E27CACEB28006F6A97C4254B1B8B93" box="[1152,1370,1903,1925]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="404" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="7696DB3DEB28006F6A97C4254B1B8B93" box="[1152,1370,1903,1925]" italics="true" pageId="1" pageNumber="404">Shaanxiana australis</emphasis>
</taxonomicName>
, male holotype (reproduced from Hsu, 2015 with permission by chief editor of Zootaxa); 3–4—
<emphasis id="7696DB3DEB28006F6A28C4C44ABB8BB2" box="[1087,1274,1934,1956]" italics="true" pageId="1" pageNumber="404">
<taxonomicName id="83E27CACEB28006F6A28C4C44AB48BB2" box="[1087,1269,1934,1956]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="404" phylum="Arthropoda" rank="species" species="langi">Shaanxiana langi</taxonomicName>
,
</emphasis>
male holotype; 5–6—
<emphasis id="7696DB3DEB28006F6EC1C4E64FD38BD4" box="[214,402,1964,1986]" italics="true" pageId="1" pageNumber="404">
<taxonomicName id="83E27CACEB28006F6EC1C4E64FCF8BD4" box="[214,398,1964,1986]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="404" phylum="Arthropoda" rank="species" species="langi">Shaanxiana langi</taxonomicName>
,
</emphasis>
female paratype; 7–8—
<taxonomicName id="83E27CACEB28006F6C85C4E64DC38BD4" box="[658,898,1964,1986]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="404" phylum="Arthropoda" rank="species" species="takashimai">
<emphasis id="7696DB3DEB28006F6C85C4E64DC38BD4" box="[658,898,1964,1986]" italics="true" pageId="1" pageNumber="404">Shaanxiana takashimai</emphasis>
</taxonomicName>
; 7—male from Zhouzhi, Shaanxi; 8—female from Fengxian, Shaanxi. 1, 3, 5—upperside; 2, 4, 6—underside; 7–8—upperside (left half) and underside (right half).
</paragraph>
</caption>
<caption id="109D57A7EB2B006C6E80C5214E998B4E" httpUri="https://zenodo.org/record/255953/files/figure.png" pageId="2" pageNumber="405" targetBox="[151,1436,193,1603]" targetPageId="2">
<paragraph id="445D072FEB2B006C6E80C5214E998B4E" blockId="2.[151,1436,1643,1880]" pageId="2" pageNumber="405">
<emphasis id="7696DB3DEB2B006C6E80C5214F548A96" bold="true" box="[151,277,1643,1665]" pageId="2" pageNumber="405">FIGURE 9.</emphasis>
Morphological differences between
<taxonomicName id="83E27CACEB2B006C6C85C5264CA08A97" box="[658,737,1643,1665]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="7696DB3DEB2B006C6C85C5264CA08A97" box="[658,737,1643,1665]" italics="true" pageId="2" pageNumber="405">S. langi</emphasis>
</taxonomicName>
and
<taxonomicName id="83E27CACEB2B006C6D00C5264DCA8A97" box="[791,907,1643,1665]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="7696DB3DEB2B006C6D00C5264DCA8A97" box="[791,907,1643,1665]" italics="true" pageId="2" pageNumber="405">S. australis</emphasis>
</taxonomicName>
, the numbers of arrows correspond to those of the characters under the
<emphasis id="7696DB3DEB2B006C6F7EC5C04F908AB6" bold="true" box="[361,465,1674,1696]" pageId="2" pageNumber="405">Diagnosis</emphasis>
heading in the text (a, b at same scale; c, d at same scale; e–i at same scale); a, b—wing-pattern of
<taxonomicName id="83E27CACEB2B006C6EA4C5E34F408AA8" box="[179,257,1704,1726]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="7696DB3DEB2B006C6EA4C5E34F408AA8" box="[179,257,1704,1726]" italics="true" pageId="2" pageNumber="405">S. langi</emphasis>
</taxonomicName>
, male; c—dorsum of
<taxonomicName id="83E27CACEB2B006C6FF5C5E34C148AA8" box="[482,597,1704,1726]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="7696DB3DEB2B006C6FF5C5E34C148AA8" box="[482,597,1704,1726]" italics="true" pageId="2" pageNumber="405">S. australis</emphasis>
</taxonomicName>
in flattened shape (reproduced from Hsu, 2015 with permission by chief editor of Zootaxa); d—dorsum of
<taxonomicName id="83E27CACEB2B006C6F83C5824FA38ACB" box="[404,482,1735,1757]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="7696DB3DEB2B006C6F83C5824FA38ACB" box="[404,482,1735,1757]" italics="true" pageId="2" pageNumber="405">S. langi</emphasis>
</taxonomicName>
in dorsal view of natural shape; e—male genitalia of
<taxonomicName id="83E27CACEB2B006C6A11C5824A128ACB" box="[1030,1107,1735,1757]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="7696DB3DEB2B006C6A11C5824A128ACB" box="[1030,1107,1735,1757]" italics="true" pageId="2" pageNumber="405">S. langi</emphasis>
</taxonomicName>
in lateral view of natural shape; f—left valva of
<taxonomicName id="83E27CACEB2B006C6F2EC5AC4FEA8AED" box="[313,427,1765,1787]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="7696DB3DEB2B006C6F2EC5AC4FEA8AED" box="[313,427,1765,1787]" italics="true" pageId="2" pageNumber="405">S. australis</emphasis>
</taxonomicName>
in flattened shape (reproduced from Hsu, 2015 with permission by chief editor of Zootaxa); g—left valva of
<taxonomicName id="83E27CACEB2B006C6EE5C44F4F018B0C" box="[242,320,1796,1818]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="7696DB3DEB2B006C6EE5C44F4F018B0C" box="[242,320,1796,1818]" italics="true" pageId="2" pageNumber="405">S. langi</emphasis>
</taxonomicName>
, holotype in flattened shape; h—right valva of
<taxonomicName id="83E27CACEB2B006C6D3DC44F4DF58B0C" box="[810,948,1796,1818]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="takashimai">
<emphasis id="7696DB3DEB2B006C6D3DC44F4DF58B0C" box="[810,948,1796,1818]" italics="true" pageId="2" pageNumber="405">S. takashimai</emphasis>
</taxonomicName>
in flattened shape (reproduced from Hsu, 2015 with permission by chief editor of Zootaxa); i—right valva of
<taxonomicName id="83E27CACEB2B006C6D05C46E4DDA8B2F" box="[786,923,1827,1849]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="takashimai">
<emphasis id="7696DB3DEB2B006C6D05C46E4DDA8B2F" box="[786,923,1827,1849]" italics="true" pageId="2" pageNumber="405">S. takashimai</emphasis>
</taxonomicName>
in flattened shape, taken from specimen shown in Fig. 7.
</paragraph>
</caption>
<subSubSection id="0CF854A4EB2B006B6ED0C4354CCC8DA4" lastPageId="5" lastPageNumber="408" pageId="2" pageNumber="405" type="description">
<paragraph id="445D072FEB2B006C6ED0C4354B408BF6" blockId="2.[151,1436,1919,2016]" pageId="2" pageNumber="405">
<emphasis id="7696DB3DEB2B006C6ED0C4354C458B8E" bold="true" box="[199,516,1919,1944]" pageId="2" pageNumber="405">
Male
<typeStatus id="9B59B98DEB2B006C6F1DC4354F308B8E" box="[266,369,1919,1944]" pageId="2" pageNumber="405" type="holotype">holotype</typeStatus>
description.
</emphasis>
Length of forewing:
<quantity id="831AAACAEB2B006C6CE3C4354D1A8B81" box="[756,859,1919,1943]" metricMagnitude="-2" metricUnit="m" metricValue="1.78" pageId="2" pageNumber="405" unit="mm" value="17.8">17.8 mm</quantity>
. Eye smooth. Antenna similar to that of
<taxonomicName id="83E27CACEB2B006C6B09C4CA4F4D8BAA" authority="Hsu, 2015" authorityName="Hsu" authorityYear="2015" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="7696DB3DEB2B006C6B09C4CA4BDD8B8E" box="[1310,1436,1919,1944]" italics="true" pageId="2" pageNumber="405">S. australis</emphasis>
<bibRefCitation id="20737ADEEB2B006C6E80C4EE4F4D8BAA" author="Hsu" box="[151,268,1955,1980]" pageId="2" pageNumber="405" refString="Hsu, Y. - F. (2015) A new species of Shaanxiana (Lepidoptera: Lycaenidae) from southern China. Zootaxa, 4027 (1), 130 - 134. http: // dx. doi. org / 10.11646 / zootaxa. 4027.1.6" type="journal article" year="2015">Hsu, 2015</bibRefCitation>
</taxonomicName>
, with black club markedly longer than that of
<taxonomicName id="83E27CACEB2B006C6D1CC4EE4DE08BAA" box="[779,929,1955,1980]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="takashimai">
<emphasis id="7696DB3DEB2B006C6D1CC4EE4DE08BAA" box="[779,929,1955,1980]" italics="true" pageId="2" pageNumber="405">S. takashimai</emphasis>
</taxonomicName>
and
<taxonomicName id="83E27CACEB2B006C6DC1C4EE4B418BAA" authority="Sugiyama, 1994" authorityName="Sugiyama" authorityYear="1994" box="[982,1280,1955,1980]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="pauper">
<emphasis id="7696DB3DEB2B006C6DC1C4EE4A018BAD" box="[982,1088,1956,1979]" italics="true" pageId="2" pageNumber="405">S. pauper</emphasis>
<bibRefCitation id="20737ADEEB2B006C6A50C4E94B418BAA" author="Sugiyama" box="[1095,1280,1955,1980]" pageId="2" pageNumber="405" refString="Sugiyama, H. (1994) New butterflies from western China. Pallarge, 3, 1 - 12." type="journal article" year="1994">Sugiyama, 1994</bibRefCitation>
</taxonomicName>
. Foretarsus 5- segmented. Hindwing with a long black tail tipped with white, and with anal lobe weakly developed.
</paragraph>
<caption id="109D57A7EB2A006D6E80C7BB4D028975" httpUri="https://zenodo.org/record/255954/files/figure.png" pageId="3" pageNumber="406" targetBox="[151,1436,193,1243]" targetPageId="3">
<paragraph id="445D072FEB2A006D6E80C7BB4D028975" blockId="3.[151,1437,1265,1379]" pageId="3" pageNumber="406">
<emphasis id="7696DB3DEB2A006D6E80C7BB4F138910" bold="true" box="[151,338,1265,1288]" pageId="3" pageNumber="406">FIGURES 10–11.</emphasis>
Male genitalia at same scale. 10—
<taxonomicName id="83E27CACEB2A006D6CA4C7B84D418911" box="[691,768,1265,1287]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="406" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="7696DB3DEB2A006D6CA4C7B84D418911" box="[691,768,1265,1287]" italics="true" pageId="3" pageNumber="406">S. langi</emphasis>
</taxonomicName>
, holotype; 11—
<taxonomicName id="83E27CACEB2A006D6DB3C7B84A6D8911" box="[932,1068,1265,1287]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="406" phylum="Arthropoda" rank="species" species="takashimai">
<emphasis id="7696DB3DEB2A006D6DB3C7B84A6D8911" box="[932,1068,1265,1287]" italics="true" pageId="3" pageNumber="406">S. takashimai</emphasis>
</taxonomicName>
, taken from specimen shown in Fig. 7. Gl = genitalia in lateral view; Ff = falces in full face view; Dd = dorsum in dorsal view; Vv = valvae in ventral view; Gp = genitalia in posterior view; Vi = left valva in inner lateral view; Vo = left valva in outer lateral view; Vd = left valva in dorsal view; Pd = phallus in dorsal view; Pl = phallus in right lateral view.
</paragraph>
</caption>
<paragraph id="445D072FEB2A006D6ED0C6DA4BCF8ADE" blockId="3.[151,1437,1424,2025]" pageId="3" pageNumber="406">Wing-pattern. Upperside. Forewing uniformly dark brown and unmarked; ciliae mostly lost, those remaining dark brown mixed with white. Hindwing dark brown; an ill-defined black marginal spot in space 2; a well-marked orange submarginal patch in space 1c, extending a little into spaces 1b and 2 on both sides; anal lobe black; ciliae lost. Underside. Forewing mostly pale yellow and unmarked except for postdiscal, submarginal and marginal areas; a postdiscal series of silvery-white, crescent-shaped spots continuous from space 1b to space 6, basal to a black submarginal band and a whitish marginal band, the latter with two short black dashes in each space; ciliae mostly lost, those remaining in space 1c white, but those in spaces 3 and 5 brown mixed with white. Hindwing similarly marked as forewing except for marginal and submarginal markings in spaces 1b–2; a distinct black marginal spot in space 2 as on upperside; and an orange submarginal patch present in spaces 1b–1c, extending a little into space 2.</paragraph>
<paragraph id="445D072FEB2A006A6ED0C59E4F2F8CEE" blockId="3.[151,1437,1424,2025]" lastBlockId="4.[151,1436,151,248]" lastPageId="4" lastPageNumber="407" pageId="3" pageNumber="406">
Male genitalia (
<figureCitation id="DCD91BAAEB2A006D6F6FC59E4F878AFB" box="[376,454,1748,1773]" captionStart="FIGURE 9" captionStartId="2.[151,250,1643,1665]" captionTargetBox="[151,1436,193,1603]" captionTargetId="figure@2.[151,1436,193,1604]" captionTargetPageId="2" captionText="FIGURE 9. Morphological differences between S. langi and S. australis, the numbers of arrows correspond to those of the characters under the Diagnosis heading in the text (a, b at same scale; c, d at same scale; e – i at same scale); a, b—wing-pattern of S. langi, male; c—dorsum of S. australis in flattened shape (reproduced from Hsu, 2015 with permission by chief editor of Zootaxa); d—dorsum of S. langi in dorsal view of natural shape; e—male genitalia of S. langi in lateral view of natural shape; f—left valva of S. australis in flattened shape (reproduced from Hsu, 2015 with permission by chief editor of Zootaxa); g—left valva of S. langi, holotype in flattened shape; h—right valva of S. takashimai in flattened shape (reproduced from Hsu, 2015 with permission by chief editor of Zootaxa); i—right valva of S. takashimai in flattened shape, taken from specimen shown in Fig. 7." httpUri="https://zenodo.org/record/255953/files/figure.png" pageId="3" pageNumber="406">Figs. 9</figureCitation>
d, 9e,
<quantity id="831AAACAEB2A006D6C1DC59F4C698AFB" box="[522,552,1749,1773]" metricMagnitude="-3" metricUnit="kg" metricValue="9.0" pageId="3" pageNumber="406" unit="g" value="9.0">9g</quantity>
, 10, 12). Tegumen in lateral view as in
<taxonomicName id="83E27CACEB2A006D6DF3C59F4A228AFB" box="[996,1123,1748,1773]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="406" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="7696DB3DEB2A006D6DF3C59F4A228AFB" box="[996,1123,1748,1773]" italics="true" pageId="3" pageNumber="406">S. australis</emphasis>
</taxonomicName>
, more robust than that of
<taxonomicName id="83E27CACEB2A006D6B90C59F4F538B07" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="406" phylum="Arthropoda" rank="species" species="takashimai">
<emphasis id="7696DB3DEB2A006D6B90C59F4F538B07" italics="true" pageId="3" pageNumber="406">S. takashimai</emphasis>
</taxonomicName>
and
<taxonomicName id="83E27CACEB2A006D6F46C5B34F818B06" box="[337,448,1785,1808]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="406" phylum="Arthropoda" rank="species" species="pauper">
<emphasis id="7696DB3DEB2A006D6F46C5B34F818B06" box="[337,448,1785,1808]" italics="true" pageId="3" pageNumber="406">S. pauper</emphasis>
</taxonomicName>
. Uncus in dorsal view protruding medially (differing from
<taxonomicName id="83E27CACEB2A006D6A96C5B34B428B07" box="[1153,1283,1784,1809]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="406" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="7696DB3DEB2A006D6A96C5B34B428B07" box="[1153,1283,1784,1809]" italics="true" pageId="3" pageNumber="406">S. australis</emphasis>
</taxonomicName>
), but largely concealed by very well developed socii. Socii more developed posteriorly than those of
<taxonomicName id="83E27CACEB2A006D6ABBC4574B068B23" box="[1196,1351,1820,1845]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="406" phylum="Arthropoda" rank="species" species="takashimai">
<emphasis id="7696DB3DEB2A006D6ABBC4574B068B23" box="[1196,1351,1820,1845]" italics="true" pageId="3" pageNumber="406">S. takashimai</emphasis>
</taxonomicName>
and
<taxonomicName id="83E27CACEB2A006D6B90C4574EA48B4E" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="406" phylum="Arthropoda" rank="species" species="pauper">
<emphasis id="7696DB3DEB2A006D6B90C4574EA48B4E" italics="true" pageId="3" pageNumber="406">S. pauper</emphasis>
</taxonomicName>
, with posterior margin convex in lateral view. Saccus as long as that of
<taxonomicName id="83E27CACEB2A006D6A32C40B4AFF8B4F" box="[1061,1214,1856,1881]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="406" phylum="Arthropoda" rank="species" species="takashimai">
<emphasis id="7696DB3DEB2A006D6A32C40B4AFF8B4F" box="[1061,1214,1856,1881]" italics="true" pageId="3" pageNumber="406">S. takashimai</emphasis>
</taxonomicName>
, approximately 1/6 times as long as height of ring (tegumen + vinculum). Falces concealed by socius and lateral process of tegumen in lateral view, gradually hooked near apex as in
<taxonomicName id="83E27CACEB2A006D6CA6C4C34D708BB7" box="[689,817,1928,1953]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="406" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="7696DB3DEB2A006D6CA6C4C34D708BB7" box="[689,817,1928,1953]" italics="true" pageId="3" pageNumber="406">S. australis</emphasis>
</taxonomicName>
, not bent medially as in
<taxonomicName id="83E27CACEB2A006D6A44C4C34AAD8BB7" box="[1107,1260,1928,1953]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="406" phylum="Arthropoda" rank="species" species="takashimai">
<emphasis id="7696DB3DEB2A006D6A44C4C34AAD8BB7" box="[1107,1260,1928,1953]" italics="true" pageId="3" pageNumber="406">S. takashimai</emphasis>
</taxonomicName>
. Valva with no harpe, in lateral view with no distal process. Juxta U-shaped in posterior view. Phallus similar to that of
<taxonomicName id="83E27CACEB2A006D6B00C4E74BD48BD3" box="[1303,1429,1964,1989]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="406" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="7696DB3DEB2A006D6B00C4E74BD48BD3" box="[1303,1429,1964,1989]" italics="true" pageId="3" pageNumber="406">S. australis</emphasis>
</taxonomicName>
, with two carinae penis strongly bent upwards beyond dorsal wall of aedeagus; dorsal wall of aedeagus colorless; entrance of vesica (internal sac) oblique, postero-ventrally downward-facing and very close to the zone on ventral surface of aedeagus; vesica when fully everted extending ventroposteriorly, non-bifurcate, bearing two cornuti, one dorsal, one ventral.
</paragraph>
<caption id="109D57A7EB2D006A6E80C50B4B428A60" httpUri="https://zenodo.org/record/255955/files/figure.png" pageId="4" pageNumber="407" targetBox="[151,1436,301,1579]" targetPageId="4">
<paragraph id="445D072FEB2D006A6E80C50B4B428A60" blockId="4.[151,1436,1601,1654]" pageId="4" pageNumber="407">
<emphasis id="7696DB3DEB2D006A6E80C50B4F148A41" bold="true" box="[151,341,1601,1624]" pageId="4" pageNumber="407">FIGURES 12–13.</emphasis>
Phallus at same scale. 12—
<taxonomicName id="83E27CACEB2D006A6C6CC5084C8B8A4E" box="[635,714,1602,1624]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="407" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="7696DB3DEB2D006A6C6CC5084C8B8A4E" box="[635,714,1602,1624]" italics="true" pageId="4" pageNumber="407">S. langi</emphasis>
</taxonomicName>
. 13—
<taxonomicName id="83E27CACEB2D006A6D1CC5084DD48A4E" box="[779,917,1602,1624]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="407" phylum="Arthropoda" rank="species" species="takashimai">
<emphasis id="7696DB3DEB2D006A6D1CC5084DD48A4E" box="[779,917,1602,1624]" italics="true" pageId="4" pageNumber="407">S. takashimai</emphasis>
</taxonomicName>
. Pll = phallus in left lateral view; Pd = phallus in dorsal view; Prl = phallus in right lateral view; Pv = phallus in ventral view; Pvl = phallus in ventrolateral view.
</paragraph>
</caption>
<paragraph id="445D072FEB2D006A6ED0C5D74FFA8BC0" blockId="4.[151,1437,1693,2006]" pageId="4" pageNumber="407">
<emphasis id="7696DB3DEB2D006A6ED0C5D74C058AA0" bold="true" box="[199,580,1693,1718]" pageId="4" pageNumber="407">
Description of female
<typeStatus id="9B59B98DEB2D006A6FCEC5D74C058AA0" box="[473,580,1693,1718]" pageId="4" pageNumber="407" type="paratype">paratype</typeStatus>
</emphasis>
. Length of forewing:
<quantity id="831AAACAEB2D006A6D5CC5D74DF78AA0" box="[843,950,1693,1718]" metricMagnitude="-2" metricUnit="m" metricValue="2.26" pageId="4" pageNumber="407" unit="mm" value="22.6">22.6 mm</quantity>
. Sexual dimorphism well marked in size, wing-shape, ground color of forewing upperside and submarginal markings on both wings underside. Wings broader than the male, the dorsum of the forewing and the costa of the hindwing each relatively longer with respect to the respective termen. Forewing upperside with an extensive, non-iridescent pale blue area from wing-base to postdiscal area, leaving only costal and submarginal areas black. Hindwing ground color brownish-grey, markedly paler than in male, extensively dusted with blue scales in the postdiscal and submarginal areas. Both wings underside with black submarginal band broader than in male, centrally dusted with some obscure, dull-orange scales. Hindwing underside with orange patch in spaces 1b–2 more pronounced than in male. Tails markedly longer than those of male.
</paragraph>
<paragraph id="445D072FEB2C006B6ED0C3D04CCC8DA4" blockId="5.[151,1437,151,434]" pageId="5" pageNumber="408">
Female genitalia (
<figureCitation id="DCD91BAAEB2C006B6F81C3D04FAD8CA5" box="[406,492,154,179]" captionStart="FIGURES 14 – 15" captionStartId="5.[151,264,1795,1818]" captionTargetBox="[151,1436,486,1771]" captionTargetId="figure@5.[151,1436,486,1771]" captionTargetPageId="5" captionText="FIGURES 14 – 15. Female genitalia at same scale. 14 — S. langi; 15 — S. takashimai. A = female genitalia in lateral view; B = 8 th tergum, lamella and antrum in lateral view; C = female genitalia in lateral view, with 7 th tergum and 7 th sternum removed; D = lamella and antrum in ventral view; E = 8 th tergum and papilla analis in dorsal view; F = 8 th tergum and papilla analis in ventral view, removed from genitalia; G, H = ostium bursae and lamella in posterior view; I, J = lamella and ostium bursae in dorsolateral view, removed from 8 th tergum; K, L = lamella, ostium bursae and corpus bursae in dorsal view, removed from 8 th tergum." httpUri="https://zenodo.org/record/255956/files/figure.png" pageId="5" pageNumber="408">Fig. 14</figureCitation>
). Lodix longer than that of
<taxonomicName id="83E27CACEB2C006B6D26C3D14D8A8CA5" box="[817,971,154,179]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="408" phylum="Arthropoda" rank="species" species="takashimai">
<emphasis id="7696DB3DEB2C006B6D26C3D14D8A8CA5" box="[817,971,154,179]" italics="true" pageId="5" pageNumber="408">S. takashimai</emphasis>
</taxonomicName>
. 8th tergum in lateral view with anterior margin sinuate. Apophysis anterioris absent. Apophysis posterioris nearly as long as 8th tergum and abruptly enlarged at apex. Papilla analis elongate, depressed dorsoventrally. Lamella developed as a ring around entrance of ostium, with lamella postvaginalis interrupted medially. Antrum abnormal, with a dorsal pouch which is pigmented on its inner wall. Ductus bursae absent, replaced by a cervix bursae immediately connected to antrum. Attachment point of ductus seminalis located on dorsal surface of the beginning of cervix bursae. Cephalic end of cervix bursae sclerotized and pigmented. Corpus bursae global. Signa weakly marked as two groups of sclerotized and pigmented teeth on surface of corpus bursae.
</paragraph>
</subSubSection>
<caption id="109D57A7EB2C006B6E80C4494EA48BAC" httpUri="https://zenodo.org/record/255956/files/figure.png" pageId="5" pageNumber="408" targetBox="[151,1436,486,1771]" targetPageId="5">
<paragraph id="445D072FEB2C006B6E80C4494EA48BAC" blockId="5.[151,1436,1793,1978]" pageId="5" pageNumber="408">
<emphasis id="7696DB3DEB2C006B6E80C4494F128B0F" bold="true" box="[151,339,1795,1818]" pageId="5" pageNumber="408">FIGURES 14–15.</emphasis>
Female genitalia at same scale. 14—
<taxonomicName id="83E27CACEB2C006B6CDEC44E4D578B0C" box="[713,790,1796,1818]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="408" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="7696DB3DEB2C006B6CDEC44E4D578B0C" box="[713,790,1796,1818]" italics="true" pageId="5" pageNumber="408">S. langi</emphasis>
</taxonomicName>
; 15—
<taxonomicName id="83E27CACEB2C006B6D42C44E4D9C8B0C" box="[853,989,1796,1818]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="408" phylum="Arthropoda" rank="species" species="takashimai">
<emphasis id="7696DB3DEB2C006B6D42C44E4D9C8B0C" box="[853,989,1796,1818]" italics="true" pageId="5" pageNumber="408">S. takashimai</emphasis>
</taxonomicName>
. A = female genitalia in lateral view; B = 8th tergum, lamella and antrum in lateral view; C = female genitalia in lateral view, with 7th tergum and 7th sternum removed; D = lamella and antrum in ventral view; E = 8th tergum and papilla analis in dorsal view; F = 8th tergum and papilla analis in ventral view, removed from genitalia; G, H = ostium bursae and lamella in posterior view; I, J = lamella and ostium bursae in dorsolateral view, removed from 8th tergum; K, L = lamella, ostium bursae and corpus bursae in dorsal view, removed from 8th tergum.
</paragraph>
</caption>
<subSubSection id="0CF854A4EB2F00686ED0C3DD4C5E8CC3" pageId="6" pageNumber="409" type="etymology">
<paragraph id="445D072FEB2F00686ED0C3DD4C5E8CC3" blockId="6.[151,1436,151,285]" pageId="6" pageNumber="409">
<emphasis id="7696DB3DEB2F00686ED0C3DD4F0F8CA6" bold="true" box="[199,334,151,176]" pageId="6" pageNumber="409">Etymology.</emphasis>
This new species is named in honor of Dr. Song-Yun Lang, from Chongqing, who collected the
<typeStatus id="9B59B98DEB2F00686E80C3F74E868CC3" box="[151,199,189,213]" pageId="6" pageNumber="409">type</typeStatus>
specimens of the new species.
</paragraph>
</subSubSection>
<subSubSection id="0CF854A4EB2F00686ED0C3954A4B8F2E" pageId="6" pageNumber="409" type="diagnosis">
<paragraph id="445D072FEB2F00686ED0C3954A1E8D0B" blockId="6.[151,1436,151,285]" pageId="6" pageNumber="409">
<emphasis id="7696DB3DEB2F00686ED0C3954FD88CEE" bold="true" box="[199,409,223,248]" pageId="6" pageNumber="409">
Diagnosis (
<figureCitation id="DCD91BAAEB2F00686F5DC3954FD08CEE" box="[330,401,223,248]" captionStart="FIGURE 9" captionStartId="2.[151,250,1643,1665]" captionTargetBox="[151,1436,193,1603]" captionTargetId="figure@2.[151,1436,193,1604]" captionTargetPageId="2" captionText="FIGURE 9. Morphological differences between S. langi and S. australis, the numbers of arrows correspond to those of the characters under the Diagnosis heading in the text (a, b at same scale; c, d at same scale; e – i at same scale); a, b—wing-pattern of S. langi, male; c—dorsum of S. australis in flattened shape (reproduced from Hsu, 2015 with permission by chief editor of Zootaxa); d—dorsum of S. langi in dorsal view of natural shape; e—male genitalia of S. langi in lateral view of natural shape; f—left valva of S. australis in flattened shape (reproduced from Hsu, 2015 with permission by chief editor of Zootaxa); g—left valva of S. langi, holotype in flattened shape; h—right valva of S. takashimai in flattened shape (reproduced from Hsu, 2015 with permission by chief editor of Zootaxa); i—right valva of S. takashimai in flattened shape, taken from specimen shown in Fig. 7." httpUri="https://zenodo.org/record/255953/files/figure.png" pageId="6" pageNumber="409">Fig. 9</figureCitation>
)
</emphasis>
. This new species is similar to
<taxonomicName id="83E27CACEB2F00686D16C3AA4DC18CEE" box="[769,896,223,248]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="409" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="7696DB3DEB2F00686D16C3AA4DC18CEE" box="[769,896,223,248]" italics="true" pageId="6" pageNumber="409">S. australis</emphasis>
</taxonomicName>
, but can be distinguished from the latter by the following combination of male characters (the female of
<taxonomicName id="83E27CACEB2F00686D01C24F4DD58D0B" box="[790,916,260,285]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="409" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="7696DB3DEB2F00686D01C24F4DD58D0B" box="[790,916,260,285]" italics="true" pageId="6" pageNumber="409">S. australis</emphasis>
</taxonomicName>
is still unknown):
</paragraph>
<paragraph id="445D072FEB2F00686E80C2064A8C8D72" blockId="6.[151,1436,332,824]" box="[151,1229,332,357]" pageId="6" pageNumber="409">
1 Size is markedly smaller, with a forewing-length of
<quantity id="831AAACAEB2F00686D19C2064D2A8D72" box="[782,875,332,356]" metricMagnitude="-2" metricUnit="m" metricValue="1.78" pageId="6" pageNumber="409" unit="mm" value="17.8">17.8mm</quantity>
against
<quantity id="831AAACAEB2F00686DDFC2064A658D72" box="[968,1060,332,356]" metricMagnitude="-2" metricUnit="m" metricValue="2.2199999999999998" pageId="6" pageNumber="409" unit="mm" value="22.2">22.2mm</quantity>
in
<taxonomicName id="83E27CACEB2F00686A5FC2074A878D73" box="[1096,1222,332,357]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="409" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="7696DB3DEB2F00686A5FC2074A878D73" box="[1096,1222,332,357]" italics="true" pageId="6" pageNumber="409">S. australis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="445D072FEB2F00686E80C23A4B368D9E" blockId="6.[151,1436,332,824]" box="[151,1399,367,392]" pageId="6" pageNumber="409">2 Hindwing upperside without a clearly defined orange patch just basal to the black marginal spot in space 2.</paragraph>
<paragraph id="445D072FEB2F00686E80C2DE4D878DD9" blockId="6.[151,1436,332,824]" pageId="6" pageNumber="409">
3 Black submarginal band on both wings underside less marked by orange scales, although the condition of the specimen examined is fresher in the new species than in
<taxonomicName id="83E27CACEB2F00686D56C2F24DFE8DC6" box="[833,959,439,464]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="409" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="7696DB3DEB2F00686D56C2F24DFE8DC6" box="[833,959,439,464]" italics="true" pageId="6" pageNumber="409">S. australis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="445D072FEB2F00686E80C2964ACF8DE2" blockId="6.[151,1436,332,824]" box="[151,1166,476,501]" pageId="6" pageNumber="409">4 Hindwing underside without a clearly defined orange patch basal to the marginal spot.</paragraph>
<paragraph id="445D072FEB2F00686E80C2B54B6E8E2B" blockId="6.[151,1436,332,824]" pageId="6" pageNumber="409">
5 Uncus in male genitalia well developed and protruding medially in dorsal view, whereas that of
<taxonomicName id="83E27CACEB2F00686B14C14A4BC08E0E" box="[1283,1409,511,536]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="409" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="7696DB3DEB2F00686B14C14A4BC08E0E" box="[1283,1409,511,536]" italics="true" pageId="6" pageNumber="409">S. australis</emphasis>
</taxonomicName>
is weakly developed as “a barely recognizable extension with shallow medial depression” (
<bibRefCitation id="20737ADEEB2F00686ABDC16E4B628E2B" author="Hsu" box="[1194,1315,548,573]" pageId="6" pageNumber="409" refString="Hsu, Y. - F. (2015) A new species of Shaanxiana (Lepidoptera: Lycaenidae) from southern China. Zootaxa, 4027 (1), 130 - 134. http: // dx. doi. org / 10.11646 / zootaxa. 4027.1.6" type="journal article" year="2015">Hsu, 2015</bibRefCitation>
).
</paragraph>
<paragraph id="445D072FEB2F00686E80C10D4B238E49" blockId="6.[151,1436,332,824]" box="[151,1378,583,608]" pageId="6" pageNumber="409">
6 Saccus in male genitalia markedly shorter than in
<taxonomicName id="83E27CACEB2F00686CE0C1024D348E76" box="[759,885,583,608]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="409" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="7696DB3DEB2F00686CE0C1024D348E76" box="[759,885,583,608]" italics="true" pageId="6" pageNumber="409">S. australis</emphasis>
</taxonomicName>
, and nearly as long as that of
<taxonomicName id="83E27CACEB2F00686AD3C1024B1A8E76" box="[1220,1371,583,608]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="409" phylum="Arthropoda" rank="species" species="takashimai">
<emphasis id="7696DB3DEB2F00686AD3C1024B1A8E76" box="[1220,1371,583,608]" italics="true" pageId="6" pageNumber="409">S. takashimai</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="445D072FEB2F00686E80C1274A4B8F2E" blockId="6.[151,1436,332,824]" pageId="6" pageNumber="409">
7 Valva of male genitalia, when flattened in full-face view, not strongly arched at upper margin as that of
<taxonomicName id="83E27CACEB2F00686B90C1274F688EBE" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="409" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="7696DB3DEB2F00686B90C1274F688EBE" italics="true" pageId="6" pageNumber="409">S. australis</emphasis>
</taxonomicName>
. (As shown in
<figureCitation id="DCD91BAAEB2F00686FC1C1C54C5A8EBE" box="[470,539,655,680]" captionStart="FIGURE 9" captionStartId="2.[151,250,1643,1665]" captionTargetBox="[151,1436,193,1603]" captionTargetId="figure@2.[151,1436,193,1604]" captionTargetPageId="2" captionText="FIGURE 9. Morphological differences between S. langi and S. australis, the numbers of arrows correspond to those of the characters under the Diagnosis heading in the text (a, b at same scale; c, d at same scale; e – i at same scale); a, b—wing-pattern of S. langi, male; c—dorsum of S. australis in flattened shape (reproduced from Hsu, 2015 with permission by chief editor of Zootaxa); d—dorsum of S. langi in dorsal view of natural shape; e—male genitalia of S. langi in lateral view of natural shape; f—left valva of S. australis in flattened shape (reproduced from Hsu, 2015 with permission by chief editor of Zootaxa); g—left valva of S. langi, holotype in flattened shape; h—right valva of S. takashimai in flattened shape (reproduced from Hsu, 2015 with permission by chief editor of Zootaxa); i—right valva of S. takashimai in flattened shape, taken from specimen shown in Fig. 7." httpUri="https://zenodo.org/record/255953/files/figure.png" pageId="6" pageNumber="409">Fig. 9</figureCitation>
, the valvae of both
<taxonomicName id="83E27CACEB2F00686CEBC1DA4D138EBE" box="[764,850,655,680]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="409" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="7696DB3DEB2F00686CEBC1DA4D138EBE" box="[764,850,655,680]" italics="true" pageId="6" pageNumber="409">S. langi</emphasis>
</taxonomicName>
and
<taxonomicName id="83E27CACEB2F00686D9DC1DA4A488EBE" box="[906,1033,655,680]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="409" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="7696DB3DEB2F00686D9DC1DA4A488EBE" box="[906,1033,655,680]" italics="true" pageId="6" pageNumber="409">S. australis</emphasis>
</taxonomicName>
are spread without any part folded, thus a comparison between them is convincing. To give a better understanding for the figures published by
<bibRefCitation id="20737ADEEB2F00686B78C1FF4F558EE6" author="Hsu" pageId="6" pageNumber="409" refString="Hsu, Y. - F. (2015) A new species of Shaanxiana (Lepidoptera: Lycaenidae) from southern China. Zootaxa, 4027 (1), 130 - 134. http: // dx. doi. org / 10.11646 / zootaxa. 4027.1.6" type="journal article" year="2015">Hsu (2015)</bibRefCitation>
, the valva of a male of
<taxonomicName id="83E27CACEB2F00686C31C1924CFE8EE6" box="[550,703,727,752]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="409" phylum="Arthropoda" rank="species" species="takashimai">
<emphasis id="7696DB3DEB2F00686C31C1924CFE8EE6" box="[550,703,727,752]" italics="true" pageId="6" pageNumber="409">S. takashimai</emphasis>
</taxonomicName>
was flattened in the same condition and compared with the one given by
<bibRefCitation id="20737ADEEB2F00686F27C1B74FF38F03" author="Hsu" box="[304,434,764,789]" pageId="6" pageNumber="409" refString="Hsu, Y. - F. (2015) A new species of Shaanxiana (Lepidoptera: Lycaenidae) from southern China. Zootaxa, 4027 (1), 130 - 134. http: // dx. doi. org / 10.11646 / zootaxa. 4027.1.6" type="journal article" year="2015">Hsu (2015)</bibRefCitation>
; the difference between
<figureCitation id="DCD91BAAEB2F00686CD1C1B64D478F03" box="[710,774,764,789]" captionStart="FIGURE 9" captionStartId="2.[151,250,1643,1665]" captionTargetBox="[151,1436,193,1603]" captionTargetId="figure@2.[151,1436,193,1604]" captionTargetPageId="2" captionText="FIGURE 9. Morphological differences between S. langi and S. australis, the numbers of arrows correspond to those of the characters under the Diagnosis heading in the text (a, b at same scale; c, d at same scale; e – i at same scale); a, b—wing-pattern of S. langi, male; c—dorsum of S. australis in flattened shape (reproduced from Hsu, 2015 with permission by chief editor of Zootaxa); d—dorsum of S. langi in dorsal view of natural shape; e—male genitalia of S. langi in lateral view of natural shape; f—left valva of S. australis in flattened shape (reproduced from Hsu, 2015 with permission by chief editor of Zootaxa); g—left valva of S. langi, holotype in flattened shape; h—right valva of S. takashimai in flattened shape (reproduced from Hsu, 2015 with permission by chief editor of Zootaxa); i—right valva of S. takashimai in flattened shape, taken from specimen shown in Fig. 7." httpUri="https://zenodo.org/record/255953/files/figure.png" pageId="6" pageNumber="409">Fig. 9</figureCitation>
h and
<figureCitation id="DCD91BAAEB2F00686D5AC1B64DCE8F03" box="[845,911,764,789]" captionStart="FIGURE 9" captionStartId="2.[151,250,1643,1665]" captionTargetBox="[151,1436,193,1603]" captionTargetId="figure@2.[151,1436,193,1604]" captionTargetPageId="2" captionText="FIGURE 9. Morphological differences between S. langi and S. australis, the numbers of arrows correspond to those of the characters under the Diagnosis heading in the text (a, b at same scale; c, d at same scale; e – i at same scale); a, b—wing-pattern of S. langi, male; c—dorsum of S. australis in flattened shape (reproduced from Hsu, 2015 with permission by chief editor of Zootaxa); d—dorsum of S. langi in dorsal view of natural shape; e—male genitalia of S. langi in lateral view of natural shape; f—left valva of S. australis in flattened shape (reproduced from Hsu, 2015 with permission by chief editor of Zootaxa); g—left valva of S. langi, holotype in flattened shape; h—right valva of S. takashimai in flattened shape (reproduced from Hsu, 2015 with permission by chief editor of Zootaxa); i—right valva of S. takashimai in flattened shape, taken from specimen shown in Fig. 7." httpUri="https://zenodo.org/record/255953/files/figure.png" pageId="6" pageNumber="409">
Fig.
<date id="305C21EFEB2F00686D96C1B64DCE8F03" box="[897,911,764,789]" pageId="6" pageNumber="409">9</date>
</figureCitation>
i indicates that in the figures published by
<bibRefCitation id="20737ADEEB2F00686B78C1B74F558F2E" author="Hsu" pageId="6" pageNumber="409" refString="Hsu, Y. - F. (2015) A new species of Shaanxiana (Lepidoptera: Lycaenidae) from southern China. Zootaxa, 4027 (1), 130 - 134. http: // dx. doi. org / 10.11646 / zootaxa. 4027.1.6" type="journal article" year="2015">Hsu (2015)</bibRefCitation>
, some part of the upper marginal area was folded, not fully spread.)
</paragraph>
</subSubSection>
<subSubSection id="0CF854A4EB2F00666ED0C02D4F7B8E91" lastPageId="8" lastPageNumber="411" pageId="6" pageNumber="409" type="discussion">
<paragraph id="445D072FEB2F00686ED0C02D4E9F8819" blockId="6.[151,1437,871,2013]" pageId="6" pageNumber="409">
<emphasis id="7696DB3DEB2F00686ED0C02D4F7A8F96" bold="true" box="[199,315,871,896]" pageId="6" pageNumber="409">Remarks.</emphasis>
Some of the above-mentioned morphological differences might be explained by individual variation within a single species, such as size, details of wing-pattern, and length of saccus and shape of valva in the male genitalia. However, individual variation affecting the form of the uncus is very seldom seen in the
<taxonomicName id="83E27CACEB2F00686B14C0E54B258FDE" box="[1283,1380,943,968]" class="Insecta" family="Lycaenidae" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="409" phylum="Arthropoda" rank="tribe" tribe="Theclini">Theclini</taxonomicName>
. The difference between the uncus of
<taxonomicName id="83E27CACEB2F00686C02C09F4C2E8FFB" box="[533,623,980,1005]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="409" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="7696DB3DEB2F00686C02C09F4C2E8FFB" box="[533,623,980,1005]" italics="true" pageId="6" pageNumber="409">S. langi</emphasis>
</taxonomicName>
and
<taxonomicName id="83E27CACEB2F00686CBBC09F4D6F8FFB" box="[684,814,980,1005]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="409" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="7696DB3DEB2F00686CBBC09F4D6F8FFB" box="[684,814,980,1005]" italics="true" pageId="6" pageNumber="409">S. australis</emphasis>
</taxonomicName>
is therefore considered indicative of separate species status.
</paragraph>
<paragraph id="445D072FEB2F00686ED0C7564ACF886B" blockId="6.[151,1437,871,2013]" pageId="6" pageNumber="409">
The female
<typeStatus id="9B59B98DEB2F00686F5BC7574FED8823" box="[332,428,1053,1077]" pageId="6" pageNumber="409" type="paratype">paratype</typeStatus>
of
<taxonomicName id="83E27CACEB2F00686FC7C7574C648823" box="[464,549,1052,1077]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="409" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="7696DB3DEB2F00686FC7C7574C648823" box="[464,549,1052,1077]" italics="true" pageId="6" pageNumber="409">S. langi</emphasis>
</taxonomicName>
has nearly the same forewing-length as the
<typeStatus id="9B59B98DEB2F00686A19C7564A318823" box="[1038,1136,1052,1077]" pageId="6" pageNumber="409" type="holotype">holotype</typeStatus>
of
<taxonomicName id="83E27CACEB2F00686A83C7574B538823" box="[1172,1298,1052,1077]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="409" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="7696DB3DEB2F00686A83C7574B538823" box="[1172,1298,1052,1077]" italics="true" pageId="6" pageNumber="409">S. australis</emphasis>
</taxonomicName>
, however, it has a relatively slender body and shorter antenna than the latter. It is possible that
<taxonomicName id="83E27CACEB2F00686A42C70A4A97884E" box="[1109,1238,1087,1112]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="409" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="7696DB3DEB2F00686A42C70A4A97884E" box="[1109,1238,1087,1112]" italics="true" pageId="6" pageNumber="409">S. australis</emphasis>
</taxonomicName>
is a more robust species than
<taxonomicName id="83E27CACEB2F00686F31C72F4F3A886B" box="[294,379,1124,1149]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="409" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="7696DB3DEB2F00686F31C72F4F3A886B" box="[294,379,1124,1149]" italics="true" pageId="6" pageNumber="409">S. langi</emphasis>
</taxonomicName>
, with a longer antenna and stouter body in proportion to its wing size.
</paragraph>
<paragraph id="445D072FEB2F00686ED0C7CD4BD788D3" blockId="6.[151,1437,871,2013]" pageId="6" pageNumber="409">
The orange scales of the forewing underside submarginal band of the female
<taxonomicName id="83E27CACEB2F00686A28C7C24AD688B6" box="[1087,1175,1159,1184]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="409" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="7696DB3DEB2F00686A28C7C24AD688B6" box="[1087,1175,1159,1184]" italics="true" pageId="6" pageNumber="409">S. langi</emphasis>
</taxonomicName>
<typeStatus id="9B59B98DEB2F00686AB7C7C34B4188B7" box="[1184,1280,1161,1185]" pageId="6" pageNumber="409" type="paratype">paratype</typeStatus>
, even though this specimen is in markedly fresher condition than the
<typeStatus id="9B59B98DEB2F00686D13C7E64D2788D3" box="[772,870,1196,1221]" pageId="6" pageNumber="409" type="holotype">holotype</typeStatus>
of
<taxonomicName id="83E27CACEB2F00686D9CC7E74A4888D3" box="[907,1033,1196,1221]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="409" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="7696DB3DEB2F00686D9CC7E74A4888D3" box="[907,1033,1196,1221]" italics="true" pageId="6" pageNumber="409">S. australis</emphasis>
</taxonomicName>
, are less apparent than in the latter.
</paragraph>
<paragraph id="445D072FEB2F00686ED0C7854DDB8A1E" blockId="6.[151,1437,871,2013]" pageId="6" pageNumber="409">
<emphasis id="7696DB3DEB2F00686ED0C7854F8A88FE" bold="true" box="[199,459,1231,1256]" pageId="6" pageNumber="409">Generic classification</emphasis>
. The discovery of
<taxonomicName id="83E27CACEB2F00686CB9C79A4D7188FE" box="[686,816,1231,1256]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="409" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="7696DB3DEB2F00686CB9C79A4D7188FE" box="[686,816,1231,1256]" italics="true" pageId="6" pageNumber="409">S. australis</emphasis>
</taxonomicName>
requires some adjustments to the characterization of
<taxonomicName id="83E27CACEB2F00686E80C7BE4F5D891B" box="[151,284,1268,1293]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="409" phylum="Arthropoda" rank="genus">
<emphasis id="7696DB3DEB2F00686E80C7BE4F5D891B" box="[151,284,1268,1293]" italics="true" pageId="6" pageNumber="409">Shaanxiana</emphasis>
</taxonomicName>
given by (
<bibRefCitation id="20737ADEEB2F00686F80C7BE4C53891B" author="Hsu" box="[407,530,1268,1293]" pageId="6" pageNumber="409" refString="Hsu, Y. - F. (2015) A new species of Shaanxiana (Lepidoptera: Lycaenidae) from southern China. Zootaxa, 4027 (1), 130 - 134. http: // dx. doi. org / 10.11646 / zootaxa. 4027.1.6" type="journal article" year="2015">Hsu, 2015</bibRefCitation>
): 1) the uncus can be obsolete or bifurcate; and 2) the harpe can be absent. The female of
<taxonomicName id="83E27CACEB2F00686F1DC6524F218926" box="[266,352,1303,1328]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="409" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="7696DB3DEB2F00686F1DC6524F218926" box="[266,352,1303,1328]" italics="true" pageId="6" pageNumber="409">S. langi</emphasis>
</taxonomicName>
demonstrates that marked sexual dimorphism can occur in the genus: the female differs sharply from the male in having extensive pale blue areas on upperside of both wings. A careful examination of both male and female genitalia reveals more specific variations as follows: falces can be concealed by socii and tegumen or entirely exposed; the everted vesica (internal sac) of phallus may be branched or not (
<figureCitation id="DCD91BAAEB2F00686A90C6CE4B55898A" box="[1159,1300,1412,1437]" captionStart="FIGURES 12 – 13" captionStartId="4.[151,264,1601,1624]" captionTargetBox="[151,1436,301,1579]" captionTargetId="figure@4.[151,1436,301,1580]" captionTargetPageId="4" captionText="FIGURES 12 – 13. Phallus at same scale. 12 — S. langi. 13 — S. takashimai. Pll = phallus in left lateral view; Pd = phallus in dorsal view; Prl = phallus in right lateral view; Pv = phallus in ventral view; Pvl = phallus in ventrolateral view." httpUri="https://zenodo.org/record/255955/files/figure.png" pageId="6" pageNumber="409">Figs. 12–13</figureCitation>
); antrum of female genitalia can be normal (
<figureCitation id="DCD91BAAEB2F00686C04C6ED4C1C89D6" box="[531,605,1447,1472]" captionStart="FIGURES 14 – 15" captionStartId="5.[151,264,1795,1818]" captionTargetBox="[151,1436,486,1771]" captionTargetId="figure@5.[151,1436,486,1771]" captionTargetPageId="5" captionText="FIGURES 14 – 15. Female genitalia at same scale. 14 — S. langi; 15 — S. takashimai. A = female genitalia in lateral view; B = 8 th tergum, lamella and antrum in lateral view; C = female genitalia in lateral view, with 7 th tergum and 7 th sternum removed; D = lamella and antrum in ventral view; E = 8 th tergum and papilla analis in dorsal view; F = 8 th tergum and papilla analis in ventral view, removed from genitalia; G, H = ostium bursae and lamella in posterior view; I, J = lamella and ostium bursae in dorsolateral view, removed from 8 th tergum; K, L = lamella, ostium bursae and corpus bursae in dorsal view, removed from 8 th tergum." httpUri="https://zenodo.org/record/255956/files/figure.png" pageId="6" pageNumber="409">Fig.15</figureCitation>
C, 15K) or specialized into a dorsal pouch (
<figureCitation id="DCD91BAAEB2F00686A74C6ED4AF889D6" box="[1123,1209,1447,1472]" captionStart="FIGURES 14 – 15" captionStartId="5.[151,264,1795,1818]" captionTargetBox="[151,1436,486,1771]" captionTargetId="figure@5.[151,1436,486,1771]" captionTargetPageId="5" captionText="FIGURES 14 – 15. Female genitalia at same scale. 14 — S. langi; 15 — S. takashimai. A = female genitalia in lateral view; B = 8 th tergum, lamella and antrum in lateral view; C = female genitalia in lateral view, with 7 th tergum and 7 th sternum removed; D = lamella and antrum in ventral view; E = 8 th tergum and papilla analis in dorsal view; F = 8 th tergum and papilla analis in ventral view, removed from genitalia; G, H = ostium bursae and lamella in posterior view; I, J = lamella and ostium bursae in dorsolateral view, removed from 8 th tergum; K, L = lamella, ostium bursae and corpus bursae in dorsal view, removed from 8 th tergum." httpUri="https://zenodo.org/record/255956/files/figure.png" pageId="6" pageNumber="409">
Fig.
<date id="305C21EFEB2F00686A8CC6E24AF889D6" box="[1179,1209,1448,1472]" pageId="6" pageNumber="409" value="2014-01">14</date>
</figureCitation>
I); ductus bursae of female genitalia can be normal (
<figureCitation id="DCD91BAAEB2F00686C16C6864C1089F3" box="[513,593,1484,1509]" captionStart="FIGURES 14 – 15" captionStartId="5.[151,264,1795,1818]" captionTargetBox="[151,1436,486,1771]" captionTargetId="figure@5.[151,1436,486,1771]" captionTargetPageId="5" captionText="FIGURES 14 – 15. Female genitalia at same scale. 14 — S. langi; 15 — S. takashimai. A = female genitalia in lateral view; B = 8 th tergum, lamella and antrum in lateral view; C = female genitalia in lateral view, with 7 th tergum and 7 th sternum removed; D = lamella and antrum in ventral view; E = 8 th tergum and papilla analis in dorsal view; F = 8 th tergum and papilla analis in ventral view, removed from genitalia; G, H = ostium bursae and lamella in posterior view; I, J = lamella and ostium bursae in dorsolateral view, removed from 8 th tergum; K, L = lamella, ostium bursae and corpus bursae in dorsal view, removed from 8 th tergum." httpUri="https://zenodo.org/record/255956/files/figure.png" pageId="6" pageNumber="409">Fig. 15</figureCitation>
C) or absent and replaced by a cervix bursae (
<figureCitation id="DCD91BAAEB2F00686A40C6864AE889F2" box="[1111,1193,1484,1509]" captionStart="FIGURES 14 – 15" captionStartId="5.[151,264,1795,1818]" captionTargetBox="[151,1436,486,1771]" captionTargetId="figure@5.[151,1436,486,1771]" captionTargetPageId="5" captionText="FIGURES 14 – 15. Female genitalia at same scale. 14 — S. langi; 15 — S. takashimai. A = female genitalia in lateral view; B = 8 th tergum, lamella and antrum in lateral view; C = female genitalia in lateral view, with 7 th tergum and 7 th sternum removed; D = lamella and antrum in ventral view; E = 8 th tergum and papilla analis in dorsal view; F = 8 th tergum and papilla analis in ventral view, removed from genitalia; G, H = ostium bursae and lamella in posterior view; I, J = lamella and ostium bursae in dorsolateral view, removed from 8 th tergum; K, L = lamella, ostium bursae and corpus bursae in dorsal view, removed from 8 th tergum." httpUri="https://zenodo.org/record/255956/files/figure.png" pageId="6" pageNumber="409">Fig. 14</figureCitation>
C); corpus bursae can be simply spherical (
<figureCitation id="DCD91BAAEB2F00686F97C6A54F918A1E" box="[384,464,1519,1544]" captionStart="FIGURES 14 – 15" captionStartId="5.[151,264,1795,1818]" captionTargetBox="[151,1436,486,1771]" captionTargetId="figure@5.[151,1436,486,1771]" captionTargetPageId="5" captionText="FIGURES 14 – 15. Female genitalia at same scale. 14 — S. langi; 15 — S. takashimai. A = female genitalia in lateral view; B = 8 th tergum, lamella and antrum in lateral view; C = female genitalia in lateral view, with 7 th tergum and 7 th sternum removed; D = lamella and antrum in ventral view; E = 8 th tergum and papilla analis in dorsal view; F = 8 th tergum and papilla analis in ventral view, removed from genitalia; G, H = ostium bursae and lamella in posterior view; I, J = lamella and ostium bursae in dorsolateral view, removed from 8 th tergum; K, L = lamella, ostium bursae and corpus bursae in dorsal view, removed from 8 th tergum." httpUri="https://zenodo.org/record/255956/files/figure.png" pageId="6" pageNumber="409">Fig. 14</figureCitation>
C) or bearing lateral pouches (
<figureCitation id="DCD91BAAEB2F00686D32C6A54D348A1E" box="[805,885,1519,1544]" captionStart="FIGURES 14 – 15" captionStartId="5.[151,264,1795,1818]" captionTargetBox="[151,1436,486,1771]" captionTargetId="figure@5.[151,1436,486,1771]" captionTargetPageId="5" captionText="FIGURES 14 – 15. Female genitalia at same scale. 14 — S. langi; 15 — S. takashimai. A = female genitalia in lateral view; B = 8 th tergum, lamella and antrum in lateral view; C = female genitalia in lateral view, with 7 th tergum and 7 th sternum removed; D = lamella and antrum in ventral view; E = 8 th tergum and papilla analis in dorsal view; F = 8 th tergum and papilla analis in ventral view, removed from genitalia; G, H = ostium bursae and lamella in posterior view; I, J = lamella and ostium bursae in dorsolateral view, removed from 8 th tergum; K, L = lamella, ostium bursae and corpus bursae in dorsal view, removed from 8 th tergum." httpUri="https://zenodo.org/record/255956/files/figure.png" pageId="6" pageNumber="409">Fig. 15</figureCitation>
K).
</paragraph>
<paragraph id="445D072FEB2F00686ED0C55E4DCA8B5B" blockId="6.[151,1437,871,2013]" pageId="6" pageNumber="409">
The four known species of
<taxonomicName id="83E27CACEB2F00686C10C55E4CCA8A3B" box="[519,651,1556,1581]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="409" phylum="Arthropoda" rank="genus">
<emphasis id="7696DB3DEB2F00686C10C55E4CCA8A3B" box="[519,651,1556,1581]" italics="true" pageId="6" pageNumber="409">Shaanxiana</emphasis>
</taxonomicName>
clearly divide into two groups:
<taxonomicName id="83E27CACEB2F00686A15C55F4ADD8A3B" box="[1026,1180,1556,1581]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="409" phylum="Arthropoda" rank="species" species="takashimai">
<emphasis id="7696DB3DEB2F00686A15C55F4ADD8A3B" box="[1026,1180,1556,1581]" italics="true" pageId="6" pageNumber="409">S. takashimai</emphasis>
</taxonomicName>
and
<taxonomicName id="83E27CACEB2F00686ACFC55F4B078A3A" box="[1240,1350,1557,1580]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="409" phylum="Arthropoda" rank="species" species="pauper">
<emphasis id="7696DB3DEB2F00686ACFC55F4B078A3A" box="[1240,1350,1557,1580]" italics="true" pageId="6" pageNumber="409">S. pauper</emphasis>
</taxonomicName>
have a bifurcate uncus, less developed socii, exposed falces, an elongate valva with a harpe, less developed carinae penis on the aedeagus, bifurcate everted vesica of the phallus, and in the female genitalia, probably a shorter lodix, a more elongate papilla analis, a normal antrum, a ductus bursae, no cervix bursae, lateral pouches on the corpus bursae, and lacking sexual dimorphism affecting the upperside wing-patterns. In contrast,
<taxonomicName id="83E27CACEB2F00686A98C5EF4B4F8AAB" box="[1167,1294,1700,1725]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="409" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="7696DB3DEB2F00686A98C5EF4B4F8AAB" box="[1167,1294,1700,1725]" italics="true" pageId="6" pageNumber="409">S. australis</emphasis>
</taxonomicName>
and
<taxonomicName id="83E27CACEB2F00686B52C5EF4BDD8AAB" box="[1349,1436,1700,1725]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="409" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="7696DB3DEB2F00686B52C5EF4BDD8AAB" box="[1349,1436,1700,1725]" italics="true" pageId="6" pageNumber="409">S. langi</emphasis>
</taxonomicName>
have a non-bifurcate uncus, more developed socii, concealed falces, a stouter valva without a harpe, better developed carinae penis of the aedeagus, non-bifurcate everted vesica of the phallus, and in the female, probably a longer lodix, stouter papilla analis, a specialized antrum, no ductus bursae, a cervix bursae, no lateral pouches on the ductus bursae, and sexually dimorphic upperside wing-patterns.
</paragraph>
<paragraph id="445D072FEB2F00696ED0C41D4C518D56" blockId="6.[151,1437,871,2013]" lastBlockId="7.[151,1436,151,320]" lastPageId="7" lastPageNumber="410" pageId="6" pageNumber="409">
Arguably
<taxonomicName id="83E27CACEB2F00686F2EC4124FF98B66" box="[313,440,1879,1904]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="409" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="7696DB3DEB2F00686F2EC4124FF98B66" box="[313,440,1879,1904]" italics="true" pageId="6" pageNumber="409">S. australis</emphasis>
</taxonomicName>
and
<taxonomicName id="83E27CACEB2F00686FE5C4124C088B66" box="[498,585,1879,1904]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="409" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="7696DB3DEB2F00686FE5C4124C088B66" box="[498,585,1879,1904]" italics="true" pageId="6" pageNumber="409">S. langi</emphasis>
</taxonomicName>
represent a genus separate from
<taxonomicName id="83E27CACEB2F00686DD5C41D4A078B66" box="[962,1094,1879,1904]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="409" phylum="Arthropoda" rank="genus">
<emphasis id="7696DB3DEB2F00686DD5C41D4A078B66" box="[962,1094,1879,1904]" italics="true" pageId="6" pageNumber="409">Shaanxiana</emphasis>
</taxonomicName>
, as the genital differences are comparable to those of other genera of
<taxonomicName id="83E27CACEB2F00686C45C4364CEE8B83" box="[594,687,1916,1941]" class="Insecta" family="Lycaenidae" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="409" phylum="Arthropoda" rank="tribe" tribe="Theclini">Theclini</taxonomicName>
recognised by
<bibRefCitation id="20737ADEEB2F00686D4BC4364A578B82" author="Koiwaya" box="[860,1046,1916,1941]" pageId="6" pageNumber="409" refString="Koiwaya, S. (2007) The Zephyrus hairstreaks of the World. Mushi-Sha, Tokyo, 300 pp. [in Japanese]" type="book" year="2007">Koiwaya (2007)</bibRefCitation>
. However, there seems little doubt that the two species groups of
<taxonomicName id="83E27CACEB2F00686FEFC4EA4C3C8BAF" box="[504,637,1952,1977]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="409" phylum="Arthropoda" rank="genus">
<emphasis id="7696DB3DEB2F00686FEFC4EA4C3C8BAF" box="[504,637,1952,1977]" italics="true" pageId="6" pageNumber="409">Shaanxiana</emphasis>
</taxonomicName>
together constitute a monophyletic group, and cannot be separated in any phylogenetic analysis using morphological characters, as they share the following genital characters besides their striking exophenotypic similarity: 1) carinae penis paired and well developed; 2) entrance of vesica of phallus large and oblique ventrally; 3) cornuti of phallus paired and well marked; 4) papilla analis elongate and depressed dorsoventrally; 5) lamella ring-shaped and interrupted at middle of lamella postvaginalis; 6) attachment point of ductus seminalis close to entrance of ostium; and 7) signum represented by a group of sclerotized and pigmented teeth on surface of corpus bursae.
</paragraph>
<caption id="109D57A7EB2E00696E80C5E74DB78AD5" box="[151,1014,1709,1731]" httpUri="https://zenodo.org/record/255957/files/figure.png" pageId="7" pageNumber="410" targetBox="[151,1436,373,1687]" targetPageId="7">
<paragraph id="445D072FEB2E00696E80C5E74DB78AD5" blockId="7.[151,1014,1709,1731]" box="[151,1014,1709,1731]" pageId="7" pageNumber="410">
<emphasis id="7696DB3DEB2E00696E80C5E74F5E8AD4" bold="true" box="[151,287,1709,1731]" pageId="7" pageNumber="410">FIGURE 16.</emphasis>
Habitat of
<taxonomicName id="83E27CACEB2E00696F85C5E44F9E8AD5" box="[402,479,1709,1731]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="410" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="7696DB3DEB2E00696F85C5E44F9E8AD5" box="[402,479,1709,1731]" italics="true" pageId="7" pageNumber="410">S. langi</emphasis>
</taxonomicName>
at Dayaoshan, Guangxi (photograph by S.-Y. Lang).
</paragraph>
</caption>
<paragraph id="445D072FEB2E00666ED0C5A04FCA8DC6" blockId="7.[151,1436,1770,2011]" lastBlockId="8.[151,1437,151,647]" lastPageId="8" lastPageNumber="411" pageId="7" pageNumber="410">
The author prefers to use the following indicators (‘rules of thumb’) to delimit genera when there are various choices regarding different clades, giving preference to: 1) the clade with a higher Bremer Support value and resampling support value (such as jackknifing or bootstrapping); 2) the clade with more synapomorphies; 3) the clade with more diagnostic characters, regardless of source (external, internal, larval, molecular etc.); 4) finally, historical usage is preferred if the first three indicators do not help. Koiwaya’s (2007) indicators for generic status were evidently based very largely on genital characters, neglecting many potentially important external features in cases such as
<taxonomicName id="83E27CACEB2E00696F24C4884C788BCD" authority="Fabricius, 1807" authorityName="Fabricius" authorityYear="1807" box="[307,569,1986,2011]" class="Insecta" family="Lycaenidae" genus="Thecla" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="410" phylum="Arthropoda" rank="genus">
<emphasis id="7696DB3DEB2E00696F24C4884FC18BCD" box="[307,384,1986,2011]" italics="true" pageId="7" pageNumber="410">Thecla</emphasis>
Fabricius, 1807
</taxonomicName>
, and
<emphasis id="7696DB3DEB2E00696C63C4884CAE8BCD" box="[628,751,1986,2011]" italics="true" pageId="7" pageNumber="410">Iozephyrus</emphasis>
Wang &amp; Fan, 2002, and of
<taxonomicName id="83E27CACEB2E00666A3CC4884E938CA6" authority="Shirozu &amp; Yamamoto, 1956" authorityName="Shirozu &amp; Yamamoto" authorityYear="1956" class="Insecta" family="Lycaenidae" genus="Gonerilia" kingdom="Animalia" lastPageId="8" lastPageNumber="411" order="Lepidoptera" pageId="7" pageNumber="410" phylum="Arthropoda" rank="genus">
<emphasis id="7696DB3DEB2E00696A3CC4884AD98BCD" box="[1067,1176,1986,2011]" italics="true" pageId="7" pageNumber="410">Gonerilia</emphasis>
Shirôzu &amp; Yamamoto, 1956
</taxonomicName>
,
<taxonomicName id="83E27CACEB2100666EC8C3DD4CCF8CA6" authority="Shirozu &amp; Yamamoto, 1956" authorityName="Shirozu &amp; Yamamoto" authorityYear="1956" box="[223,654,151,176]" class="Insecta" family="Lycaenidae" genus="Cordelia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="411" phylum="Arthropoda" rank="genus">
<emphasis id="7696DB3DEB2100666EC8C3DD4F038CA6" box="[223,322,151,176]" italics="true" pageId="8" pageNumber="411">Cordelia</emphasis>
Shirôzu &amp; Yamamoto, 1956
</taxonomicName>
,
<emphasis id="7696DB3DEB2100666C8CC3DD4D6E8CA6" box="[667,815,151,176]" italics="true" pageId="8" pageNumber="411">Neogonerilia</emphasis>
<bibRefCitation id="20737ADEEB2100666D20C3DD4DA98CA6" author="Koiwaya" box="[823,1000,151,176]" pageId="8" pageNumber="411" refString="Koiwaya, S. (2007) The Zephyrus hairstreaks of the World. Mushi-Sha, Tokyo, 300 pp. [in Japanese]" type="book" year="2007">Koiwaya, 2007</bibRefCitation>
, and
<emphasis id="7696DB3DEB2100666A31C3DD4A9E8CA6" box="[1062,1247,151,176]" italics="true" pageId="8" pageNumber="411">Pseudogonerilia</emphasis>
<bibRefCitation id="20737ADEEB2100666AFFC3DD4BD98CA6" author="Koiwaya" box="[1256,1432,151,176]" pageId="8" pageNumber="411" refString="Koiwaya, S. (2007) The Zephyrus hairstreaks of the World. Mushi-Sha, Tokyo, 300 pp. [in Japanese]" type="book" year="2007">Koiwaya, 2007</bibRefCitation>
, for which generic separations were rejected by
<bibRefCitation id="20737ADEEB2100666CD7C3F64A708CC2" author="Weidenhoffer" box="[704,1073,188,213]" pageId="8" pageNumber="411" refString="Weidenhoffer, Z. &amp; Bozano, G. C. (2007) Guide to the butterflies of the Palearctic Region - Lycaenidae part III. Omnes Artes, Milano, 97 pp." type="book" year="2007">Weidenhoffer &amp; Bozano (2007)</bibRefCitation>
. As there is no sound work on morphological phylogeny for
<emphasis id="7696DB3DEB2100666FF2C3954C318CEE" box="[485,624,223,248]" italics="true" pageId="8" pageNumber="411">
<taxonomicName id="83E27CACEB2100666FF2C3954C2D8CEE" box="[485,620,223,248]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="411" phylum="Arthropoda" rank="genus">Shaanxiana</taxonomicName>
,
</emphasis>
<taxonomicName id="83E27CACEB2100666C60C3AB4D248CEE" authority="Tutt, 1907" authorityName="Tutt" authorityYear="1907" box="[631,869,224,248]" class="Insecta" family="Lycaenidae" genus="Ussuriana" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="411" phylum="Arthropoda" rank="genus">
<emphasis id="7696DB3DEB2100666C60C3AB4CAD8CEE" box="[631,748,225,248]" italics="true" pageId="8" pageNumber="411">Ussuriana</emphasis>
Tutt, 1907
</taxonomicName>
,
<taxonomicName id="83E27CACEB2100666D64C3954DA18CEE" box="[883,992,223,248]" class="Insecta" family="Lycaenidae" genus="Gonerilia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="411" phylum="Arthropoda" rank="genus">
<emphasis id="7696DB3DEB2100666D64C3954DA18CEE" box="[883,992,223,248]" italics="true" pageId="8" pageNumber="411">Gonerilia</emphasis>
</taxonomicName>
(
<emphasis id="7696DB3DEB2100666DE7C3AB4A6F8CEE" box="[1008,1070,225,248]" italics="true" pageId="8" pageNumber="411">sensu</emphasis>
<bibRefCitation id="20737ADEEB2100666A23C3954BD18CEE" author="Weidenhoffer" box="[1076,1424,223,248]" pageId="8" pageNumber="411" refString="Weidenhoffer, Z. &amp; Bozano, G. C. (2007) Guide to the butterflies of the Palearctic Region - Lycaenidae part III. Omnes Artes, Milano, 97 pp." type="book" year="2007">Weidenhoffer &amp; Bozano, 2007</bibRefCitation>
),
<taxonomicName id="83E27CACEB2100666E80C24F4F348D0A" authority="Tutt, 1907" authorityName="Tutt" authorityYear="1907" box="[151,373,260,284]" class="Insecta" family="Lycaenidae" genus="Coreana" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="411" phylum="Arthropoda" rank="genus">
<emphasis id="7696DB3DEB2100666E80C24F4EB98D0A" box="[151,248,261,284]" italics="true" pageId="8" pageNumber="411">Coreana</emphasis>
Tutt, 1907
</taxonomicName>
,
<taxonomicName id="83E27CACEB2100666F68C24F4CD98D0B" authority="Rambur, 1858" authorityName="Rambur" authorityYear="1858" box="[383,664,260,285]" class="Insecta" family="Lycaenidae" genus="Laeosopis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="411" phylum="Arthropoda" rank="genus">
<emphasis id="7696DB3DEB2100666F68C24F4FB08D0A" box="[383,497,261,284]" italics="true" pageId="8" pageNumber="411">Laeosopis</emphasis>
Rambur, 1858
</taxonomicName>
, and
<taxonomicName id="83E27CACEB2100666CC4C24E4A468D0A" authority="Chapman, 1909" authorityName="Chapman" authorityYear="1909" box="[723,1031,260,285]" class="Insecta" family="Lycaenidae" genus="Artopoetes" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="411" phylum="Arthropoda" rank="genus">
<emphasis id="7696DB3DEB2100666CC4C24E4D0D8D0D" box="[723,844,260,283]" italics="true" pageId="8" pageNumber="411">Artopoetes</emphasis>
Chapman, 1909
</taxonomicName>
, which were supposed to be close to
<taxonomicName id="83E27CACEB2100666E80C26D4F5D8D56" box="[151,284,295,320]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="411" phylum="Arthropoda" rank="genus">
<emphasis id="7696DB3DEB2100666E80C26D4F5D8D56" box="[151,284,295,320]" italics="true" pageId="8" pageNumber="411">Shaanxiana</emphasis>
</taxonomicName>
in the phylogeny by
<bibRefCitation id="20737ADEEB2100666C18C26D4C888D56" author="Koiwaya" box="[527,713,295,320]" pageId="8" pageNumber="411" refString="Koiwaya, S. (2007) The Zephyrus hairstreaks of the World. Mushi-Sha, Tokyo, 300 pp. [in Japanese]" type="book" year="2007">Koiwaya (2007)</bibRefCitation>
, the above-mentioned Indicator 3 is preferred in this work. The genus
<taxonomicName id="83E27CACEB2100666EC8C2064F258D73" box="[223,356,332,357]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="411" phylum="Arthropoda" rank="genus">
<emphasis id="7696DB3DEB2100666EC8C2064F258D73" box="[223,356,332,357]" italics="true" pageId="8" pageNumber="411">Shaanxiana</emphasis>
</taxonomicName>
is separable from related genera not only by the seven genital characters listed above but also by various striking external characters, such as the wing-pattern. Since the number of diagnostic characters of the two species groups of
<taxonomicName id="83E27CACEB2100666F73C2DE4FA88DBB" box="[356,489,404,429]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="411" phylum="Arthropoda" rank="genus">
<emphasis id="7696DB3DEB2100666F73C2DE4FA88DBB" box="[356,489,404,429]" italics="true" pageId="8" pageNumber="411">Shaanxiana</emphasis>
</taxonomicName>
is less than that of the collective genus, it is at best to treat them as species groups within a single genus.
</paragraph>
<paragraph id="445D072FEB2100666ED0C2964F7B8E91" blockId="8.[151,1437,151,647]" pageId="8" pageNumber="411">
<emphasis id="7696DB3DEB2100666ED0C2964FE38DE3" bold="true" box="[199,418,476,501]" pageId="8" pageNumber="411">Field observations</emphasis>
. The
<typeStatus id="9B59B98DEB2100666FF4C2974C528DE3" box="[483,531,477,501]" pageId="8" pageNumber="411">type</typeStatus>
specimens were found perching and flying around a tree of
<taxonomicName id="83E27CACEB2100666AACC2964BDD8DE3" box="[1211,1436,476,501]" class="Actinopterygii" family="Osmeridae" genus="Mallotus" kingdom="Animalia" order="Osmeriformes" pageId="8" pageNumber="411" phylum="Chordata" rank="species" species="tenuifolius">
<emphasis id="7696DB3DEB2100666AACC2964BDD8DE3" box="[1211,1436,476,501]" italics="true" pageId="8" pageNumber="411">Mallotus tenuifolius</emphasis>
</taxonomicName>
<taxonomicName id="83E27CACEB2100666E80C1494FBE8E0D" box="[151,511,514,539]" pageId="8" pageNumber="411" rank="variety" variety="tenuifolius">
var.
<emphasis id="7696DB3DEB2100666EDFC1484F7F8E0D" box="[200,318,514,539]" italics="true" pageId="8" pageNumber="411">tenuifolius</emphasis>
(Euphorbiaceae)
</taxonomicName>
at 1:00–2:00 pm on
<date id="305C21EFEB2100666CF9C1494DD38E0D" box="[750,914,511,539]" pageId="8" pageNumber="411" value="2015-05-21">21st May 2015</date>
, at an elevation of
<quantity id="831AAACAEB2100666A7BC1494AFC8E0D" box="[1132,1213,515,539]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="411" unit="m" value="1000.0">1000m</quantity>
. It was very cloudy and the female eventually captured stayed on the tree when it was disturbed by a strike on nearby branches. The two
<taxonomicName id="83E27CACEB2100666EDFC1004F438E75" box="[200,258,586,611]" class="Insecta" family="Lycaenidae" genus="Shaanxiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="411" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="7696DB3DEB2100666EDFC1004F438E75" box="[200,258,586,611]" italics="true" pageId="8" pageNumber="411">langi</emphasis>
</taxonomicName>
were collected along with five adults of
<taxonomicName id="83E27CACEB2100666CDDC1064A368E75" authority="Oberthur, 1880" authorityName="Oberthur" authorityYear="1880" box="[714,1143,586,611]" class="Insecta" family="Lycaenidae" genus="Ussuriana" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="411" phylum="Arthropoda" rank="species" species="michaelis">
<emphasis id="7696DB3DEB2100666CDDC1064DF18E75" box="[714,944,586,611]" italics="true" pageId="8" pageNumber="411">Ussuriana michaelis</emphasis>
(Oberthür, 1880)
</taxonomicName>
, found on the same tree at the same time.
</paragraph>
</subSubSection>
</treatment>
</document>