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<mods:title id="0BB81BC7307A1D09137DDC5C3B2694EF">Description of a new species of the earthworm genus Moniligaster Perrier, 1872 (Clitellata, Moniligastridae) from the Nilgiri Biosphere Reserve of Tamil Nadu State, India</mods:title>
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<mods:namePart id="6957899244C2B03122A25D4A285DAFE3">Narayanan, S. Prasanth</mods:namePart>
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<mods:affiliation id="A31E286796B7935D5CEA799420C1552D">Advanced Centre of Environmental Studies and Sustainable Development, Mahatma Gandhi University, Priyadarsini Hills, Kottayam - 686560, Kerala, India</mods:affiliation>
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<mods:namePart id="EB6569300A0EBF6BCEBA93E1B7ADF5CB">Ahmed, Shakoor</mods:namePart>
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<mods:affiliation id="75E55C43000FA28238332325C79165A5">Zoological Survey of India (Ministry of Environment, Forest and Climate Change, Government of India), FPS Building, Indian Museum Complex, Kolkata - 700016, West Bengal, India</mods:affiliation>
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<mods:namePart id="BD677071F7A09DC0CA40A97C8AAB837B">Thomas, A. P.</mods:namePart>
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<mods:affiliation id="7B390484F24D1D74D9AF8DCEF0E3101B">Advanced Centre of Environmental Studies and Sustainable Development, Mahatma Gandhi University, Priyadarsini Hills, Kottayam - 686560, Kerala, India</mods:affiliation>
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<mods:date id="9AD355A2A269BAEA300A8EC70FA370D5">2025</mods:date>
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Key to the
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<taxonomicName id="78F1DCCAFFF3FF9FD721FC56FD87FC1E" ID-CoL="84QJY" authority="Perrier, 1872" authorityName="Perrier" authorityYear="1872" box="[291,618,906,933]" class="Clitellata" family="Moniligastridae" genus="Moniligaster" kingdom="Animalia" order="Moniligastrida" pageId="9" pageNumber="151" phylum="Annelida" rank="genus">
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<emphasis id="8D857B5BFFF3FF9FD721FC56FE53FC1F" bold="true" box="[291,446,906,933]" italics="true" pageId="9" pageNumber="151">Moniligaster</emphasis>
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<bibRefCitation id="DB60DAB8FFF3FF9FD7C5FC56FD87FC1E" author="Perrier, E." box="[455,618,906,932]" pageId="9" pageNumber="151" pagination="5 - 198" refId="ref10162" refString="Perrier, E. (1872) Recherches pour servir a l'histoire des lombriciens terrestres. Nouveaux Archives du Museum National d ' Histoire Naturelle, Paris, 8, 5-198. [in French] https://doi.org/10.5962/bhl.title.12201" type="journal article" year="1872">Perrier, 1872</bibRefCitation>
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</taxonomicName>
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species (modified and updated based on
|
||||
<bibRefCitation id="DB60DAB8FFF3FF9FD271FC56FAECFC1E" author="Gates, G. E." box="[1139,1281,906,932]" pageId="9" pageNumber="151" pagination="487 - 518" refId="ref8041" refString="Gates, G. E. (1940) Indian earthworms. XIII. The genus Moniligaster. Records of the Indian Museum, 42 (3), 487-518. https://doi.org/10.26515/rzsi/v42/i3/1940/162432" type="journal article" year="1940">Gates 1940</bibRefCitation>
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;
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<bibRefCitation id="DB60DAB8FFF3FF9FD312FC57FEC8FC72" author="Narayanan, S. P. & Sathrumithra, S. & Anuja, R. & Christopher, G. & Thomas, A. P. & Julka, J. M." pageId="9" pageNumber="151" pagination="381 - 397" refId="ref9276" refString="Narayanan, S. P., Sathrumithra, S., Anuja, R., Christopher, G., Thomas, A. P. & Julka, J. M. (2021 a) Three new species and four new species records of earthworms of the genus Moniligaster Perrier, 1872 (Clitellata: Moniligastridae) from Kerala region of the Western Ghats Biodiversity Hotspot, India. Zootaxa, 4949 (2), 381-397. https://doi.org/10.11646/zootaxa.4949.2.11" type="journal article" year="2021">
|
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Narayanan
|
||||
<emphasis id="8D857B5BFFF3FF9FD695FC73FF3FFC73" bold="true" box="[151,210,942,969]" italics="true" pageId="9" pageNumber="151">et al.</emphasis>
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2021a
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, 2022)
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</emphasis>
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</heading>
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<paragraph id="BF4EA749FFF3FF9FD695FC28FA71FBB3" blockId="9.[151,1436,1012,1826]" box="[151,1436,1012,1034]" pageId="9" pageNumber="151">
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<keyLead id="0400047CFFF3FF9FD695FC28FA71FBB3" box="[151,1436,1012,1034]" pageId="9" pageNumber="151">1. Vas deferens penetrates longitudinal muscle layer prior to junction with prostate................................... 2</keyLead>
|
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</paragraph>
|
||||
<paragraph id="BF4EA749FFF3FF9FD695FBCDFA71FB9C" blockId="9.[151,1436,1012,1826]" box="[151,1436,1041,1063]" pageId="9" pageNumber="151">
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<keyLead id="0400047CFFF3FF9FD695FBCDFA71FB9C" box="[151,1436,1041,1063]" pageId="9" pageNumber="151">- Vas deferens enters prostate directly, without penetrating longitudinal muscle layer................................. 4</keyLead>
|
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</paragraph>
|
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</keyStep>
|
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<keyStep id="0405BFECFFF3FF9FD695FBF2FA71FBDB" pageId="9" pageNumber="151">
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<paragraph id="BF4EA749FFF3FF9FD695FBF2FA71FBF9" blockId="9.[151,1436,1012,1826]" box="[151,1436,1070,1092]" pageId="9" pageNumber="151">
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<keyLead id="0400047CFFF3FF9FD695FBF2FA71FBF9" box="[151,1436,1070,1092]" pageId="9" pageNumber="151">
|
||||
2. Prostatic capsule ovoidal to anvil-like.....................................................
|
||||
<taxonomicName id="78F1DCCAFFF3FF9FD2CEFBF3FA71FBF9" authority="Gates, 1940" authorityName="Gates" authorityYear="1940" box="[1228,1436,1070,1092]" class="Clitellata" family="Moniligastridae" genus="Moniligaster" kingdom="Animalia" order="Moniligastrida" pageId="9" pageNumber="151" phylum="Annelida" rank="species" species="horsti">
|
||||
<emphasis id="8D857B5BFFF3FF9FD2CEFBF3FAC9FBF9" box="[1228,1316,1070,1092]" italics="true" pageId="9" pageNumber="151">M. horsti</emphasis>
|
||||
<bibRefCitation id="DB60DAB8FFF3FF9FD328FBF2FA71FBF9" author="Gates, G. E." box="[1322,1436,1070,1091]" pageId="9" pageNumber="151" pagination="487 - 518" refId="ref8041" refString="Gates, G. E. (1940) Indian earthworms. XIII. The genus Moniligaster. Records of the Indian Museum, 42 (3), 487-518. https://doi.org/10.26515/rzsi/v42/i3/1940/162432" type="journal article" year="1940">Gates, 1940</bibRefCitation>
|
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</taxonomicName>
|
||||
</keyLead>
|
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</paragraph>
|
||||
<paragraph id="BF4EA749FFF3FF9FD695FB90FA71FBDB" blockId="9.[151,1436,1012,1826]" box="[151,1436,1100,1122]" pageId="9" pageNumber="151">
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<keyLead id="0400047CFFF3FF9FD695FB90FA71FBDB" box="[151,1436,1100,1122]" pageId="9" pageNumber="151">- Prostatic capsule different.............................................................................. 3</keyLead>
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</paragraph>
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<keyStep id="0405BFECFFF3FF9FD695FBB5FA71FB21" pageId="9" pageNumber="151">
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<paragraph id="BF4EA749FFF3FF9FD695FBB5FA71FBC4" blockId="9.[151,1436,1012,1826]" box="[151,1436,1129,1151]" pageId="9" pageNumber="151">
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<keyLead id="0400047CFFF3FF9FD695FBB5FA71FBC4" box="[151,1436,1129,1151]" pageId="9" pageNumber="151">
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3. Body length>
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<quantity id="78090AACFFF3FF9FD773FBB5FE2EFBC4" box="[369,451,1129,1150]" metricMagnitude="-1" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="151" unit="mm" value="400.0">400 mm</quantity>
|
||||
; prostatic capsule spheroidal.........................................
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||||
<taxonomicName id="78F1DCCAFFF3FF9FD2CEFBB6FA71FBC4" authority="Gates, 1940" authorityName="Gates" authorityYear="1940" box="[1228,1436,1129,1151]" class="Clitellata" family="Moniligastridae" genus="Moniligaster" kingdom="Animalia" order="Moniligastrida" pageId="9" pageNumber="151" phylum="Annelida" rank="species" species="aiyeri">
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<emphasis id="8D857B5BFFF3FF9FD2CEFBB6FAC9FBC5" box="[1228,1316,1130,1151]" italics="true" pageId="9" pageNumber="151">M. aiyeri</emphasis>
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<bibRefCitation id="DB60DAB8FFF3FF9FD328FBB5FA71FBC4" author="Gates, G. E." box="[1322,1436,1129,1150]" pageId="9" pageNumber="151" pagination="487 - 518" refId="ref8041" refString="Gates, G. E. (1940) Indian earthworms. XIII. The genus Moniligaster. Records of the Indian Museum, 42 (3), 487-518. https://doi.org/10.26515/rzsi/v42/i3/1940/162432" type="journal article" year="1940">Gates, 1940</bibRefCitation>
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</taxonomicName>
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<paragraph id="BF4EA749FFF3FF9FD695FB5BFA71FB21" blockId="9.[151,1436,1012,1826]" box="[151,1436,1158,1180]" pageId="9" pageNumber="151">
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<keyLead id="0400047CFFF3FF9FD695FB5BFA71FB21" box="[151,1436,1158,1180]" pageId="9" pageNumber="151">
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- Body length <
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<quantity id="78090AACFFF3FF9FD773FB5AFE2EFB26" box="[369,451,1158,1180]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="151" unit="mm" value="200.0">200 mm</quantity>
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; prostatic capsule disc-like and latero-mesially flattened...............
|
||||
<taxonomicName id="78F1DCCAFFF3FF9FD290FB5BFA71FB21" authority="Gates, 1940" authorityName="Gates" authorityYear="1940" box="[1170,1436,1158,1180]" class="Clitellata" family="Moniligastridae" genus="Moniligaster" kingdom="Animalia" order="Moniligastrida" pageId="9" pageNumber="151" phylum="Annelida" rank="species" species="stephensoni">
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<emphasis id="8D857B5BFFF3FF9FD290FB5BFACEFB21" box="[1170,1315,1158,1180]" italics="true" pageId="9" pageNumber="151">M. stephensoni</emphasis>
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<bibRefCitation id="DB60DAB8FFF3FF9FD328FB5AFA71FB21" author="Gates, G. E." box="[1322,1436,1158,1179]" pageId="9" pageNumber="151" pagination="487 - 518" refId="ref8041" refString="Gates, G. E. (1940) Indian earthworms. XIII. The genus Moniligaster. Records of the Indian Museum, 42 (3), 487-518. https://doi.org/10.26515/rzsi/v42/i3/1940/162432" type="journal article" year="1940">Gates, 1940</bibRefCitation>
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</taxonomicName>
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</keyLead>
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<keyStep id="0405BFECFFF3FF9FD695FB78FA71FB6C" pageId="9" pageNumber="151">
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<paragraph id="BF4EA749FFF3FF9FD695FB78FA71FB03" blockId="9.[151,1436,1012,1826]" box="[151,1436,1187,1210]" pageId="9" pageNumber="151">
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<keyLead id="0400047CFFF3FF9FD695FB78FA71FB03" box="[151,1436,1187,1210]" pageId="9" pageNumber="151">4. Spermathecal atrial glands confined to segment 7............................................................ 5</keyLead>
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</paragraph>
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<paragraph id="BF4EA749FFF3FF9FD695FB1DFA71FB6C" blockId="9.[151,1436,1012,1826]" box="[151,1436,1217,1239]" pageId="9" pageNumber="151">
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<keyLead id="0400047CFFF3FF9FD695FB1DFA71FB6C" box="[151,1436,1217,1239]" pageId="9" pageNumber="151">- Spermathecal atrial glands in segments 7 and 8............................................................. 13</keyLead>
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</paragraph>
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<keyStep id="0405BFECFFF3FF9FD695FB02FA71FAAB" pageId="9" pageNumber="151">
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<paragraph id="BF4EA749FFF3FF9FD695FB02FA71FB4E" blockId="9.[151,1436,1012,1826]" box="[151,1436,1246,1268]" pageId="9" pageNumber="151">
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<keyLead id="0400047CFFF3FF9FD695FB02FA71FB4E" box="[151,1436,1246,1268]" pageId="9" pageNumber="151">5. Leaflet-like glands present on vas deferens................................................................. 6</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="BF4EA749FFF3FF9FD695FB20FA71FAAB" blockId="9.[151,1436,1012,1826]" box="[151,1436,1276,1298]" pageId="9" pageNumber="151">
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<keyLead id="0400047CFFF3FF9FD695FB20FA71FAAB" box="[151,1436,1276,1298]" pageId="9" pageNumber="151">- Leaflet-like glands absent on vas deferens.................................................................. 7</keyLead>
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</paragraph>
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</keyStep>
|
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<keyStep id="0405BFECFFF3FF9FD695FAC5FA71FAF1" pageId="9" pageNumber="151">
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<paragraph id="BF4EA749FFF3FF9FD695FAC5FA71FA94" blockId="9.[151,1436,1012,1826]" box="[151,1436,1305,1327]" pageId="9" pageNumber="151">
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<keyLead id="0400047CFFF3FF9FD695FAC5FA71FA94" box="[151,1436,1305,1327]" pageId="9" pageNumber="151">
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6. Prostate long, capsule with crisscrossed ridges, prostatic duct slender........................
|
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<taxonomicName id="78F1DCCAFFF3FF9FD29FFAC6FA71FA94" ID-CoL="853JQ" authority="Perrier, 1872" authorityName="Perrier" authorityYear="1872" box="[1181,1436,1305,1327]" class="Clitellata" family="Moniligastridae" genus="Moniligaster" kingdom="Animalia" order="Moniligastrida" pageId="9" pageNumber="151" phylum="Annelida" rank="species" species="deshayesi">
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<emphasis id="8D857B5BFFF3FF9FD29FFAC6FAF7FA94" box="[1181,1306,1305,1327]" italics="true" pageId="9" pageNumber="151">M. deshayesi</emphasis>
|
||||
<bibRefCitation id="DB60DAB8FFF3FF9FD322FAC5FA71FA94" author="Perrier, E." box="[1312,1436,1305,1327]" pageId="9" pageNumber="151" pagination="5 - 198" refId="ref10162" refString="Perrier, E. (1872) Recherches pour servir a l'histoire des lombriciens terrestres. Nouveaux Archives du Museum National d ' Histoire Naturelle, Paris, 8, 5-198. [in French] https://doi.org/10.5962/bhl.title.12201" type="journal article" year="1872">Perrier, 1872</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="BF4EA749FFF3FF9FD695FAEBFA71FAF1" blockId="9.[151,1436,1012,1826]" box="[151,1436,1334,1356]" pageId="9" pageNumber="151">
|
||||
<keyLead id="0400047CFFF3FF9FD695FAEBFA71FAF1" box="[151,1436,1334,1356]" pageId="9" pageNumber="151">
|
||||
- Prostate short, capsule smooth, prostatic duct bulged....................................
|
||||
<taxonomicName id="78F1DCCAFFF3FF9FD290FAEBFA71FAF1" authority="Michaelsen, 1913" authorityName="Michaelsen" authorityYear="1913" box="[1170,1436,1334,1356]" class="Clitellata" family="Moniligastridae" genus="Moniligaster" kingdom="Animalia" order="Moniligastrida" pageId="9" pageNumber="151" phylum="Annelida" rank="species" species="minor">
|
||||
<emphasis id="8D857B5BFFF3FF9FD290FAEBFB06FAF6" box="[1170,1259,1335,1356]" italics="true" pageId="9" pageNumber="151">M. minor</emphasis>
|
||||
<bibRefCitation id="DB60DAB8FFF3FF9FD2F0FAEAFA71FAF1" author="Michaelsen, W." box="[1266,1436,1334,1356]" pageId="9" pageNumber="151" pagination="73 - 92" refId="ref8620" refString="Michaelsen, W. (1913) Oligochaten von Travancore und Borneo. Jahrbuch der hamburgischen wissenschaftlichen Anstalten, 30, 73-92. [in German]" type="journal article" year="1913">Michaelsen, 1913</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="0405BFECFFF3FF9FD695FA88FA71FA3C" pageId="9" pageNumber="151">
|
||||
<paragraph id="BF4EA749FFF3FF9FD695FA88FA71FAD3" blockId="9.[151,1436,1012,1826]" box="[151,1436,1364,1386]" pageId="9" pageNumber="151">
|
||||
<keyLead id="0400047CFFF3FF9FD695FA88FA71FAD3" box="[151,1436,1364,1386]" pageId="9" pageNumber="151">7. Vas deferens joins prostate at or close to ental end........................................................... 8</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="BF4EA749FFF3FF9FD695FAADFA71FA3C" blockId="9.[151,1436,1012,1826]" box="[151,1436,1393,1415]" pageId="9" pageNumber="151">
|
||||
<keyLead id="0400047CFFF3FF9FD695FAADFA71FA3C" box="[151,1436,1393,1415]" pageId="9" pageNumber="151">- Vas deferens joins prostate at or close to ectal end.......................................................... 11</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="0405BFECFFF3FF9FD695FA52FA71FA7B" pageId="9" pageNumber="151">
|
||||
<paragraph id="BF4EA749FFF3FF9FD695FA52FA71FA1E" blockId="9.[151,1436,1012,1826]" box="[151,1436,1422,1444]" pageId="9" pageNumber="151">
|
||||
<keyLead id="0400047CFFF3FF9FD695FA52FA71FA1E" box="[151,1436,1422,1444]" pageId="9" pageNumber="151">
|
||||
8. Atrial gland single on each side...................................................
|
||||
<taxonomicName id="78F1DCCAFFF3FF9FD27FFA53FA71FA1E" ID-CoL="BXFT6" authority="Stephenson, 1915" authorityName="Stephenson" authorityYear="1915" box="[1149,1436,1422,1444]" class="Clitellata" family="Moniligastridae" genus="Moniligaster" kingdom="Animalia" order="Moniligastrida" pageId="9" pageNumber="151" phylum="Annelida" rank="species" species="gravelyi">
|
||||
<emphasis id="8D857B5BFFF3FF9FD27FFA53FB01FA19" box="[1149,1260,1422,1444]" italics="true" pageId="9" pageNumber="151">M. gravelyi</emphasis>
|
||||
<bibRefCitation id="DB60DAB8FFF3FF9FD2F0FA52FA71FA1E" author="Stephenson, J." box="[1266,1436,1422,1444]" pageId="9" pageNumber="151" pagination="35 - 108" refId="ref10437" refString="Stephenson, J. (1915) On some Indian Oligochaeta, mainly from Southern India and Ceylon. Memoirs of the Indian Museum, 6, 35-108." type="journal article" year="1915">Stephenson, 1915</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="BF4EA749FFF3FF9FD695FA70FA71FA7B" blockId="9.[151,1436,1012,1826]" box="[151,1436,1452,1474]" pageId="9" pageNumber="151">
|
||||
<keyLead id="0400047CFFF3FF9FD695FA70FA71FA7B" box="[151,1436,1452,1474]" pageId="9" pageNumber="151">- Atrial glands paired on each side......................................................................... 9</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="0405BFECFFF3FF9FD695FA15FA71FA41" pageId="9" pageNumber="151">
|
||||
<paragraph id="BF4EA749FFF3FF9FD695FA15FA71FA64" blockId="9.[151,1436,1012,1826]" box="[151,1436,1481,1503]" pageId="9" pageNumber="151">
|
||||
<keyLead id="0400047CFFF3FF9FD695FA15FA71FA64" box="[151,1436,1481,1503]" pageId="9" pageNumber="151">
|
||||
9. Prostate bluntly club-shaped, nodulations absent on capsule.....................
|
||||
<taxonomicName id="78F1DCCAFFF3FF9FD214FA16FA71FA64" authority="Narayanan & Julka, 2021" authorityName="Narayanan & Julka" authorityYear="2021" box="[1046,1436,1481,1503]" class="Clitellata" family="Moniligastridae" genus="Moniligaster" kingdom="Animalia" order="Moniligastrida" pageId="9" pageNumber="151" phylum="Annelida" rank="species" species="blakemorei">
|
||||
<emphasis id="8D857B5BFFF3FF9FD214FA16FB4CFA64" box="[1046,1185,1481,1503]" italics="true" pageId="9" pageNumber="151">M. blakemorei</emphasis>
|
||||
Narayanan & Julka, 2021
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="BF4EA749FFF3FF9FD695FA3BFA71FA41" blockId="9.[151,1436,1012,1826]" box="[151,1436,1510,1532]" pageId="9" pageNumber="151">
|
||||
<keyLead id="0400047CFFF3FF9FD695FA3BFA71FA41" box="[151,1436,1510,1532]" pageId="9" pageNumber="151">- Prostate strap-shaped, nodulations present on capsule........................................................ 10</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="0405BFECFFF3FF9FD695F9D8FA71F98C" pageId="9" pageNumber="151">
|
||||
<paragraph id="BF4EA749FFF3FF9FD695F9D8FA71F9A3" blockId="9.[151,1436,1012,1826]" box="[151,1436,1540,1562]" pageId="9" pageNumber="151">
|
||||
<keyLead id="0400047CFFF3FF9FD695F9D8FA71F9A3" box="[151,1436,1540,1562]" pageId="9" pageNumber="151">
|
||||
10. Prostatic capsule tubular, nodulation confined to ental end..........................
|
||||
<taxonomicName id="78F1DCCAFFF3FF9FD252F9D8FA71F9A3" authority="Narayanan & Julka, 2021" authorityName="Narayanan & Julka" authorityYear="2021" box="[1104,1436,1540,1562]" class="Clitellata" family="Moniligastridae" genus="Moniligaster" kingdom="Animalia" order="Moniligastrida" pageId="9" pageNumber="151" phylum="Annelida" rank="species" species="bahli">
|
||||
<emphasis id="8D857B5BFFF3FF9FD252F9D8FB4CF9A3" box="[1104,1185,1540,1561]" italics="true" pageId="9" pageNumber="151">M. bahli</emphasis>
|
||||
Narayanan & Julka, 2021
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="BF4EA749FFF3FF9FD695F9FDFA71F98C" blockId="9.[151,1436,1012,1826]" box="[151,1436,1569,1591]" pageId="9" pageNumber="151">
|
||||
<keyLead id="0400047CFFF3FF9FD695F9FDFA71F98C" box="[151,1436,1569,1591]" pageId="9" pageNumber="151">
|
||||
- Prostatic capsule tubular, full surface with low nodulations.....................
|
||||
<taxonomicName id="78F1DCCAFFF3FF9FD215F9FEFAABF98C" authority="Ahmed & Narayanan" authorityName="Ahmed & Narayanan" authorityYear="2025" box="[1047,1350,1569,1591]" class="Clitellata" family="Moniligastridae" genus="Moniligaster" kingdom="Animalia" order="Moniligastrida" pageId="9" pageNumber="151" phylum="Annelida" rank="species" species="gatesi" status="sp. nov.">
|
||||
<emphasis id="8D857B5BFFF3FF9FD215F9FEFB9DF98D" box="[1047,1136,1570,1591]" italics="true" pageId="9" pageNumber="151">M. gatesi</emphasis>
|
||||
Ahmed & Narayanan
|
||||
</taxonomicName>
|
||||
,
|
||||
<emphasis id="8D857B5BFFF3FF9FD352F9FDFA71F98C" bold="true" box="[1360,1436,1569,1590]" pageId="9" pageNumber="151">
|
||||
<taxonomicNameLabel id="96B6C620FFF3FF9FD352F9FDFA71F98C" box="[1360,1436,1569,1590]" pageId="9" pageNumber="151" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="0405BFECFFF3FF9FD695F9E2FA71F9CB" pageId="9" pageNumber="151">
|
||||
<paragraph id="BF4EA749FFF3FF9FD695F9E2FA71F9E9" blockId="9.[151,1436,1012,1826]" box="[151,1436,1597,1620]" pageId="9" pageNumber="151">
|
||||
<keyLead id="0400047CFFF3FF9FD695F9E2FA71F9E9" box="[151,1436,1597,1620]" pageId="9" pageNumber="151">
|
||||
11. Prostates clavate; duct of each atrial gland short, about ¼ the length of common atrial duct.........
|
||||
<taxonomicName id="78F1DCCAFFF3FF9FD2B6F9E3FA71F9E9" authority="Jamieson, 1977" authorityName="Jamieson" authorityYear="1977" box="[1204,1436,1598,1620]" class="Clitellata" family="Moniligastridae" genus="Moniligaster" kingdom="Animalia" order="Moniligastrida" pageId="9" pageNumber="151" phylum="Annelida" rank="species" species="troyi">
|
||||
<emphasis id="8D857B5BFFF3FF9FD2B6F9E3FAEDF9EE" box="[1204,1280,1599,1620]" italics="true" pageId="9" pageNumber="151">M. troyi</emphasis>
|
||||
<bibRefCitation id="DB60DAB8FFF3FF9FD305F9E2FA71F9E9" author="Jamieson, B. G. M." box="[1287,1436,1598,1620]" pageId="9" pageNumber="151" pagination="95 - 114" refId="ref8219" refString="Jamieson, B. G. M. (1977) On the phylogeny of the Moniligastridae, with description of new species of Moniligaster (Oligochaeta, Annelida). Evolutionary Theory, 2, 95-114." type="journal article" year="1977">Jamieson, 1977</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="BF4EA749FFF3FF9FD695F980FA71F9CB" blockId="9.[151,1436,1012,1826]" box="[151,1436,1628,1650]" pageId="9" pageNumber="151">
|
||||
<keyLead id="0400047CFFF3FF9FD695F980FA71F9CB" box="[151,1436,1628,1650]" pageId="9" pageNumber="151">- Prostates tubular, duct of each atrial gland equal to or larger than common atrial duct.............................. 12</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="0405BFECFFF3FF9FD695F9A5FA71F911" pageId="9" pageNumber="151">
|
||||
<paragraph id="BF4EA749FFF3FF9FD695F9A5FA71F934" blockId="9.[151,1436,1012,1826]" box="[151,1436,1657,1679]" pageId="9" pageNumber="151">
|
||||
<keyLead id="0400047CFFF3FF9FD695F9A5FA71F934" box="[151,1436,1657,1679]" pageId="9" pageNumber="151">
|
||||
12. Duct of each atrial gland about as long as common atrial duct...................
|
||||
<taxonomicName id="78F1DCCAFFF3FF9FD222F9A6FA71F934" authority="Narayanan & Julka, 2021" authorityName="Narayanan & Julka" authorityYear="2021" box="[1056,1436,1657,1679]" class="Clitellata" family="Moniligastridae" genus="Moniligaster" kingdom="Animalia" order="Moniligastrida" pageId="9" pageNumber="151" phylum="Annelida" rank="species" species="keralensis">
|
||||
<emphasis id="8D857B5BFFF3FF9FD222F9A6FB4CF934" box="[1056,1185,1657,1679]" italics="true" pageId="9" pageNumber="151">M. keralensis</emphasis>
|
||||
Narayanan & Julka, 2021
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="BF4EA749FFF3FF9FD695F94BFA71F911" blockId="9.[151,1436,1012,1826]" box="[151,1436,1686,1708]" pageId="9" pageNumber="151">
|
||||
<keyLead id="0400047CFFF3FF9FD695F94BFA71F911" box="[151,1436,1686,1708]" pageId="9" pageNumber="151">
|
||||
- Duct of each atrial gland about five times the length of common atrial duct..........
|
||||
<taxonomicName id="78F1DCCAFFF3FF9FD237F94BFA71F911" authority="Narayanan & Paliwal, 2022" authorityName="Narayanan & Paliwal" authorityYear="2022" box="[1077,1436,1686,1708]" class="Clitellata" family="Moniligastridae" genus="Moniligaster" kingdom="Animalia" order="Moniligastrida" pageId="9" pageNumber="151" phylum="Annelida" rank="species" species="julkai">
|
||||
<emphasis id="8D857B5BFFF3FF9FD237F94BFB61F911" box="[1077,1164,1686,1708]" italics="true" pageId="9" pageNumber="151">M. julkai</emphasis>
|
||||
Narayanan & Paliwal, 2022
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="0405BFECFFF3FF9FD695F968FA71F95C" pageId="9" pageNumber="151">
|
||||
<paragraph id="BF4EA749FFF3FF9FD695F968FA71F973" blockId="9.[151,1436,1012,1826]" box="[151,1436,1716,1738]" pageId="9" pageNumber="151">
|
||||
<keyLead id="0400047CFFF3FF9FD695F968FA71F973" box="[151,1436,1716,1738]" pageId="9" pageNumber="151">
|
||||
13. Prostate and duct bound to parietes in a C-shaped figure, capsule ovoidal....................
|
||||
<taxonomicName id="78F1DCCAFFF3FF9FD293F968FA71F973" authority="Gates, 1940" authorityName="Gates" authorityYear="1940" box="[1169,1436,1716,1737]" class="Clitellata" family="Moniligastridae" genus="Moniligaster" kingdom="Animalia" order="Moniligastrida" pageId="9" pageNumber="151" phylum="Annelida" rank="species" species="michaelseni">
|
||||
<emphasis id="8D857B5BFFF3FF9FD293F968FAC9F973" box="[1169,1316,1716,1737]" italics="true" pageId="9" pageNumber="151">M. michaelseni</emphasis>
|
||||
<bibRefCitation id="DB60DAB8FFF3FF9FD328F968FA71F973" author="Gates, G. E." box="[1322,1436,1716,1737]" pageId="9" pageNumber="151" pagination="487 - 518" refId="ref8041" refString="Gates, G. E. (1940) Indian earthworms. XIII. The genus Moniligaster. Records of the Indian Museum, 42 (3), 487-518. https://doi.org/10.26515/rzsi/v42/i3/1940/162432" type="journal article" year="1940">Gates, 1940</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="BF4EA749FFF3FF9FD695F90DFA71F95C" blockId="9.[151,1436,1012,1826]" box="[151,1436,1745,1767]" pageId="9" pageNumber="151">
|
||||
<keyLead id="0400047CFFF3FF9FD695F90DFA71F95C" box="[151,1436,1745,1767]" pageId="9" pageNumber="151">- Prostate and duct not so bound to parietes, capsule shape otherwise............................................. 14</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="0405BFECFFF3FF9FD695F932FA71F898" pageId="9" pageNumber="151">
|
||||
<paragraph id="BF4EA749FFF3FF9FD695F932FA71F8B9" blockId="9.[151,1436,1012,1826]" box="[151,1436,1774,1796]" pageId="9" pageNumber="151">
|
||||
<keyLead id="0400047CFFF3FF9FD695F932FA71F8B9" box="[151,1436,1774,1796]" pageId="9" pageNumber="151">
|
||||
14. Prostatic capsule reniform, without nodulations.......................................
|
||||
<taxonomicName id="78F1DCCAFFF3FF9FD283F933FA71F8B9" ID-CoL="CKM68" authority="Michaelsen, 1907" authorityName="Michaelsen" authorityYear="1907" box="[1153,1436,1774,1796]" class="Clitellata" family="Moniligastridae" genus="Moniligaster" kingdom="Animalia" order="Moniligastrida" pageId="9" pageNumber="151" phylum="Annelida" rank="species" species="perrieri">
|
||||
<emphasis id="8D857B5BFFF3FF9FD283F933FB01F8BE" box="[1153,1260,1775,1796]" italics="true" pageId="9" pageNumber="151">M. perrieri</emphasis>
|
||||
<bibRefCitation id="DB60DAB8FFF3FF9FD2F0F932FA71F8B9" author="Michaelsen, W." box="[1266,1436,1774,1796]" pageId="9" pageNumber="151" pagination="143 - 188" refId="ref8550" refString="Michaelsen, W. (1907) Neue Oligochaten von Vorder-Indien, Ceylon, Birma, und den Andaman-Inseln. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 24, 143-188. [in German]" type="journal article" year="1907">Michaelsen, 1907</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="BF4EA749FFF3FF9FD695F8D0FA71F898" blockId="9.[151,1436,1012,1826]" box="[151,1436,1804,1826]" pageId="9" pageNumber="151">
|
||||
<keyLead id="0400047CFFF3FF9FD695F8D0FA71F898" box="[151,1436,1804,1826]" pageId="9" pageNumber="151">
|
||||
- Prostatic capsule short, tubular and U-shaped, with irregularly placed nodulations.............
|
||||
<taxonomicName id="78F1DCCAFFF3FF9FD28CF8D0FA71F898" authority="Gates, 1962" authorityName="Gates" authorityYear="1962" box="[1166,1436,1804,1826]" class="Clitellata" family="Moniligastridae" genus="Moniligaster" kingdom="Animalia" order="Moniligastrida" pageId="9" pageNumber="151" phylum="Annelida" rank="species" species="cernosvitovi">
|
||||
<emphasis id="8D857B5BFFF3FF9FD28CF8D0FACEF89B" box="[1166,1315,1804,1825]" italics="true" pageId="9" pageNumber="151">M. cernosvitovi</emphasis>
|
||||
<bibRefCitation id="DB60DAB8FFF3FF9FD328F8D0FA71F898" author="Gates, G. E." box="[1322,1436,1804,1826]" pageId="9" pageNumber="151" pagination="297 - 373" refId="ref8123" refString="Gates, G. E. (1962) On some Burmese earthworms of the moniligastrid genus Drawida. Bulletin of the Museum of the Comparative Zoology at Harvard College, 127 (5), 297-373." type="journal article" year="1962">Gates, 1962</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
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