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<document id="B8BA92E73ADF1EE7757DE7E33644EE57" ID-DOI="10.3897/caucasiana.3.e130238" ID-ZooBank="223368FA-DEC1-4E64-A79D-2C5EA7DBEC96" ID-publisher-id="130238" URI-arpha="676804BE-8338-559D-AA5E-F8E7F7AF0BFB" URI-zoobank="http://zoobank.org/223368FA-DEC1-4E64-A79D-2C5EA7DBEC96" article-type="research-article" checkinTime="1725892901502" checkinUser="pensoft" docAuthor="Kiria, Eleonora, Mumladze, Levan, Barjadze, Shalva &amp; Tuf, Ivan H." docDate="2024" docId="78AD1CD98B6D5205BFEF4C1F8429E1FD" docLanguage="en" docName="Caucasiana 3: 151-162" docOrigin="Caucasiana 3" docSource="https://caucasiana.pensoft.net/article/130238/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Henia hirsuta Verhoeff 1928" docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="162" masterDocId="676804BE8338559DAA5EF8E7F7AF0BFB" masterDocTitle="New records of centipedes (Myriapoda: Chilopoda) in the fauna of Georgia, South Caucasus" masterLastPageNumber="162" masterPageNumber="151" pageNumber="151" updateTime="1725893338483" updateUser="ExternalLinkService">
<mods:mods id="311C97AC5F3716A1B9EAB9C18B00C36C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D6332769D159FDF8D1F4E81C775694A9">
<mods:title id="4E1409CDC923E43517D1E7F3AAD905EF">New records of centipedes (Myriapoda: Chilopoda) in the fauna of Georgia, South Caucasus</mods:title>
</mods:titleInfo>
<mods:name id="F18D67BF93426028DDF8D7162C16EEAE" type="personal">
<mods:role id="1E260BCC562C99AB008052B8AD70933B">
<mods:roleTerm id="54053F8684C66818AF4AA02C552957DC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3F23AE9CB1C4BE09E47E22346CE72076">Kiria, Eleonora</mods:namePart>
<mods:nameIdentifier id="955F559BEB5399DB291C4666E5474304" type="ORCID">https://orcid.org/0009-0007-1711-2163</mods:nameIdentifier>
<mods:affiliation id="8570879882AD59D3FFA82F94DA54CE42">Ilia State University, Tbilisi, Georgia</mods:affiliation>
</mods:name>
<mods:name id="4CB783372795A1D98B2663454DEDE932" type="personal">
<mods:role id="6C6DE4CD8BDFF826CEFAA99D1D713B72">
<mods:roleTerm id="E0BE80995785DD66325D2054D047A178">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AEBED54911F16C0FC8783A8C3D08EB33">Mumladze, Levan</mods:namePart>
<mods:nameIdentifier id="18545447C9707A113C55089F2B6906AE" type="ORCID">0000-0002-2172-6973</mods:nameIdentifier>
<mods:affiliation id="C77C3289AE06CF0A2641AC510930C3A2">Ilia State University, Tbilisi, Georgia</mods:affiliation>
</mods:name>
<mods:name id="4C8B51DF890940A2FE414EE980C75027" type="personal">
<mods:role id="5D4917F5F7416DC5D5C148184045A9E0">
<mods:roleTerm id="5D0DB5838F7960D3809A65C887F2825E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D8427571ABB69808DC0C6E2727A78094">Barjadze, Shalva</mods:namePart>
<mods:nameIdentifier id="C7E228D15F7CD2AF7066068C6A945C90" type="ORCID">0000-0001-8992-4987</mods:nameIdentifier>
<mods:affiliation id="A3E2AA0541E888D9C0562312113C0DE1">Ilia State University, Tbilisi, Georgia</mods:affiliation>
</mods:name>
<mods:name id="859FBE74167822B8372C03FBE1BC284E" type="personal">
<mods:role id="DDBA53970BC6DBD4CD958F4186B68DD3">
<mods:roleTerm id="8AF7314B3B9EBDB32A8185E26D61D002">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7217C0B5542844B43C48ADCC13A1F63E">Tuf, Ivan H.</mods:namePart>
<mods:nameIdentifier id="10B0741BD60F112C04408A7ABF2FE16B" type="ORCID">0000-0003-0250-0482</mods:nameIdentifier>
<mods:affiliation id="E789B17C1FDD7E0F9558A7A8E72B3110">Faculty of Science, Palacký University, Olomouc, Czech Republic</mods:affiliation>
</mods:name>
<mods:typeOfResource id="02FCDEDF930115AE1463CAA2B19C6106">text</mods:typeOfResource>
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<mods:titleInfo id="393486A704EC17B1DD835DEA64369371">
<mods:title id="68A6AD087CE271CD65D3B7E30A550D4F">Caucasiana</mods:title>
</mods:titleInfo>
<mods:part id="56FE8D0A44BBD43C0CF8BB73564121C4">
<mods:date id="10F6CECB67EF113A2F5C2AC09B8EF265">2024</mods:date>
<mods:detail id="77CE74327DFC3E32FFA0F4A898913DD4" type="pubDate">
<mods:number id="C2F39D83946E7B216D4BEE9530953309">2024-09-09</mods:number>
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<mods:number id="2492FBFF094593AF7C39A6E63BBEF871">3</mods:number>
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<mods:start id="4BE7988EEF3F7DA8428E431548F18D60">151</mods:start>
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<mods:identifier id="458671A155C4A17A14168B597E61528F" type="DOI">10.3897/caucasiana.3.e130238</mods:identifier>
<mods:identifier id="4A2AFC2388A9D812DAB3650751A42C42" type="ZooBank">223368FA-DEC1-4E64-A79D-2C5EA7DBEC96</mods:identifier>
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<treatment id="78AD1CD98B6D5205BFEF4C1F8429E1FD" ID-DOI="http://doi.org/10.5281/zenodo.13737081" ID-Zenodo-Dep="13737081" ID-arpha="78AD1CD9-8B6D-5205-BFEF-4C1F8429E1FD" LSID="urn:lsid:plazi:treatment:78AD1CD98B6D5205BFEF4C1F8429E1FD" httpUri="http://treatment.plazi.org/id/78AD1CD98B6D5205BFEF4C1F8429E1FD">
<subSubSection id="C6E82B6C11A517E3B3539D46E452A848" type="nomenclature">
<paragraph id="73198A2E82864E95220D02DBB0B85444">
<heading id="76E34852DC511707E1F5B88266834AFF" reason="label">2.</heading>
<taxonomicName id="EF95F399F5337248719F390A930C8FD2" ID-CoL="6LPV8" authority="Verhoeff, 1928" authorityName="Verhoeff" authorityYear="1928" class="Chilopoda" family="Dignathodontidae" genus="Henia" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="41C005DB7F4C711B1648D5F896583453" italics="true">Henia hirsuta</emphasis>
<bibRefCitation id="7422F84DCAC95F5BF6F914B3F7518FC8" author="Verhoeff KW" refId="B27" refString="Verhoeff KW (1928) Geophilomorphen-Beitrage und eine Lithobius - Form. Mitteilungen aus dem Zoologischen Museum in Berlin 14 (2): 227 - 286. (in German)" year="1928">Verhoeff, 1928</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="585950365C9292D6F49394142CE9486F" type="description">
<paragraph id="E326109116C5F0BAF562A49E634C07AE">
<figureCitation id="F2110CB3A2D2A690855111DF97067139" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Henia hirsuta Verhoeff, 1928, male, (a) head ventral view; (b) head and first two anterior sternites, ventral view; (c) posterior segments, ventral view; (d) segments showing sternal pores, ventral view. Scale bars: 0.2 mm (a, d); 0.5 mm (b, c)." figureDoi="10.3897/caucasiana.3.e130238.figure3" httpUri="https://binary.pensoft.net/fig/1130296">Fig. 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EMMAC" type="type locality">
<paragraph id="6D3A6D28DD6EAA1B987919DADD5D96ED">
<heading id="5D69F90A84FF8A42E1F5E51ED15B55A9" reason="title">
<typeStatus id="D2C44A96F3DFA9E3F41EBD521695214D">Type</typeStatus>
locality.
</heading>
</paragraph>
<paragraph id="19E4A99812198BB620641246A06671AE">
<collectingCountry id="284B1F376BA5481B2B1FB3FEEF205A5A" name="Greece">Greece</collectingCountry>
: Mesolongi.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERMAC" type="material">
<paragraph id="E1E92C871999A347CCBBA1EE01C07F90">
<heading id="AF7EE369D581F4C80F7A822F35F06A64" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="57A77B5DD936B90E324B86475D5C44E7">
<materialsCitation id="8D4543D7BBADB13118BA3F455CA237C3" collectingDate="2013-06-03" collectorName="L. Mumladze" country="Georgia" county="Lagodekhi National Park" elevation="2347" latitude="41.87408" location="eastern Georgia" longLatPrecision="1" longitude="46.37877" municipality="Lagodekhi" specimenCount="3" specimenCount-male="3">
<collectingCountry id="A7D6E17DDCF48C48C670D05A84480E43" name="Georgia">GEORGIA</collectingCountry>
<specimenCount id="5ED173649EA4E3A1C0CDA5030C731655" count="3" type="male">3 ♂</specimenCount>
;
<location id="568C2F1D27617C180FE594659264AC30" LSID="urn:lsid:plazi:treatment:78AD1CD98B6D5205BFEF4C1F8429E1FD:568C2F1D27617C180FE594659264AC30" country="Georgia" county="Lagodekhi National Park" latitude="41.87408" longLatPrecision="1" longitude="46.37877" municipality="Lagodekhi" name="eastern Georgia">eastern Georgia</location>
,
<collectingMunicipality id="B3B43A29193FAD8DCE2385D808CC7B15">Lagodekhi</collectingMunicipality>
municipality,
<collectingCounty id="60651B91689007C447B535ED1202B47C">Lagodekhi National Park</collectingCounty>
, subalpine meadow;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E1MAC" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[46.378770,41.874080]}">
<geoCoordinate id="CD7FE4E0819981F6A8529BFEA257B380" degrees="41.87408" direction="north" orientation="latitude" precision="1" value="41.87408">N 41.87408 °</geoCoordinate>
,
<geoCoordinate id="0447E114CE4D1F9546855181206424A2" degrees="46.37877" direction="east" orientation="longitude" precision="1" value="46.37877">E 46.37877 °</geoCoordinate>
</named-content>
;
<elevation id="B770B35CBBFFDB11E1927BBA52646894" metricMagnitude="3" metricUnit="m" metricValue="2.347" unit="m" value="2347.0">
<quantity id="DAADD34F9452324CCA247360C6144495" metricMagnitude="3" metricUnit="m" metricValue="2.347" unit="m" value="2347.0">2347 m</quantity>
a. s. l.
</elevation>
;
<date id="8979E6CCFB99DCEB03418E680E3D9232" value="2013-06-03">
<collectingDate id="6CC92F3F3C5CB6960097CD8B1650A214" value="2013-06-03">3 Jun. 2013</collectingDate>
</date>
; leg.
<collectorName id="2640C3F19AF242F9876B9AAD93E99E48">L. Mumladze</collectorName>
</materialsCitation>
; •
<materialsCitation id="42CC438789CB69CE24E69C16E0EC4D5C" collectingDate="2023-05-26" collectorName="E. Kiria &amp; Kiria, M &amp; Gogshelidze, L &amp; Shavadze, A" country="Georgia" elevation="652" latitude="41.85232" location="Lagodekhi National Park" longLatPrecision="1" longitude="46.28744" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<specimenCount id="BC4F0A99D7EBB9AE89E7F28BEA3CFBC3" count="2" type="male">2 ♂</specimenCount>
,
<specimenCount id="34172CB83C296EC09F5A33795E3568F5" count="1" type="female">1 ♀</specimenCount>
;
<location id="2685A181C41C9466983DB0AB03C83BA4" LSID="urn:lsid:plazi:treatment:78AD1CD98B6D5205BFEF4C1F8429E1FD:2685A181C41C9466983DB0AB03C83BA4" country="Georgia" latitude="41.85232" longLatPrecision="1" longitude="46.28744" name="Lagodekhi National Park">Lagodekhi National Park</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ECNAC" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[46.287440,41.852320]}">
<geoCoordinate id="FDB271B0EA89607CE1346A1A470F6213" degrees="41.85232" direction="north" orientation="latitude" precision="1" value="41.85232">N 41.85232 °</geoCoordinate>
,
<geoCoordinate id="AE4461BA05BC17B14ECA4FE98B1DDE06" degrees="46.28744" direction="east" orientation="longitude" precision="1" value="46.28744">E 46.28744 °</geoCoordinate>
</named-content>
;
<elevation id="F4ADA78B5E8840B498BF4C0552177193" metricMagnitude="2" metricUnit="m" metricValue="6.52" unit="m" value="652.0">
<quantity id="8E221EDD414EDEFB50471547BA0BCD17" metricMagnitude="2" metricUnit="m" metricValue="6.52" unit="m" value="652.0">652 m</quantity>
a. s. l.
</elevation>
;
<date id="AB9FD1F3726E6EDD5762D0823EF4F462" value="2023-05-26">
<collectingDate id="843CCD40ADEB087C1294C9D89CD95B0E" value="2023-05-26">26 May 2023</collectingDate>
</date>
; leg.
<collectorName id="1BF12C31CB226556D5A7E2D11A99E274">
E.
<collectorName id="2EDB56B5B852C58D4ED29F7D9B1E3538">Kiria, M</collectorName>
</collectorName>
.
<collectorName id="EE44148336561E7318ACF795F816DE38">Gogshelidze, L</collectorName>
.
<collectorName id="47F67A81347650A60EB071D35F17AF80">Shavadze, A</collectorName>
. Margalitadze
</materialsCitation>
; •
<materialsCitation id="01642BE4BE54350B89E98DD1EBD282E5" collectingDate="2017-05-24" collectorName="L. Mumladze." country="Georgia" county="Bochorma" elevation="965" latitude="41.92391" location="eastern Georgia" longLatPrecision="1" longitude="45.10189" municipality="Tianeti" specimenCount="2" specimenCount-male="2">
<specimenCount id="E00C0B8800DA39FCBEFBC72EBFB77464" count="2" type="male">2 ♂</specimenCount>
;
<location id="D2DC541C5807C4B645312A56ABD08C95" LSID="urn:lsid:plazi:treatment:78AD1CD98B6D5205BFEF4C1F8429E1FD:D2DC541C5807C4B645312A56ABD08C95" country="Georgia" county="Bochorma" latitude="41.92391" longLatPrecision="1" longitude="45.10189" municipality="Tianeti" name="eastern Georgia">eastern Georgia</location>
,
<collectingMunicipality id="387E361E73701C1DC0AC1CEE1896F682">Tianeti</collectingMunicipality>
municipality, village
<collectingCounty id="7386ECD9ACE24A9DA140790CB47369FB">Bochorma</collectingCounty>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EKNAC" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[45.101890,41.923910]}">
<geoCoordinate id="C5D90E493D6869504BB7B0C5BF68612E" degrees="41.92391" direction="north" orientation="latitude" precision="1" value="41.92391">N 41.92391 °</geoCoordinate>
,
<geoCoordinate id="AE0A89033EE3D4F276194B554E5F9445" degrees="45.10189" direction="east" orientation="longitude" precision="1" value="45.10189">E 45.10189 °</geoCoordinate>
</named-content>
;
<elevation id="238A7F98CFFE5B1D47C6228E5DF9F95C" metricMagnitude="2" metricUnit="m" metricValue="9.65" unit="m" value="965.0">
<quantity id="8F9FB111DF971E6EC009E1E45FDBFEE6" metricMagnitude="2" metricUnit="m" metricValue="9.65" unit="m" value="965.0">965 m</quantity>
a. s. l.
</elevation>
;
<date id="4DFE7A46F5977B127F42FD435EFB6F31" value="2017-05-24">
<collectingDate id="A7F588F333189404D0649FD745D65344" value="2017-05-24">24 May 2017</collectingDate>
</date>
; leg.
<collectorName id="E992F164062AFC712176659B43B1F5D0">L. Mumladze.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EPNAC" type="distribution">
<paragraph id="33EB59B7B43718E68F3609A977C2A7CD">
<heading id="9738CB35D802EE64EE339B4FD58B4DB8" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="896DE13973D3D7EC6970231D7C7E933E">
<collectingCountry id="03E22EF6DD785EA6C390B8B86AC66075" name="Greece">Greece</collectingCountry>
(
<bibRefCitation id="7453D9B9713D19CB6E19FD3F1943B5FD" author="Verhoeff" firstAuthor="Verhoeff" refId="B27" refString="Verhoeff KW (1928) Geophilomorphen-Beiträge und eine Lithobius - Form. Mitteilungen aus dem Zoologischen Museum in Berlin 14 (2): 227-286. (in German)" year="1928">Verhoeff 1928</bibRefCitation>
,
<bibRefCitation id="35E528699236E2386CEA87EF769AF955" author="Kanellis" firstAuthor="Kanellis" journalOrPublisher="Scientific Annals of the Faculty of Physics and Mathematics of the University of Thessaloniki" pagination="1-56" refId="B12" refString="Kanellis A (1959) Die Chilopodenfauna Griechenlands. Scientific Annals of the Faculty of Physics and Mathematics of the University of Thessaloniki 1: 156." title="Die Chilopodenfauna Griechenlands." volume="1" year="1959">Kanellis 1959</bibRefCitation>
,
<bibRefCitation id="70CE5D1C17F3796F13BAD3EDB711731B" DOI="10.5962/bhl.part.91440" author="Matic" firstAuthor="Matic" journalOrPublisher="Revue suisse de Zoologie" pagination="287-306" refId="B17" refString="Matic Z (1976) Sur quelques Myriapodes Chilopodes du Muséum dHistoire naturelle de Genève. Revue suisse de Zoologie 83: 287306. https://doi.org/10.5962/bhl.part.91440" title="Sur quelques Myriapodes Chilopodes du Muséum dHistoire naturelle de Genève." volume="83" year="1976">Matic 1976</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EBOAC" type="remarks">
<paragraph id="1A1B5016FF87FC020DBC995840EDAAFE">
<heading id="558FD9A0CB93DBB61C5AE95605E15738" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="5F19D78A847B2152327375BE98DAC71B">
This species was described from a specimen found near Mesolongi, central Greece (
<bibRefCitation id="0C776D2C401CC1D6DD25BAD5DB591A2F" author="Verhoeff" firstAuthor="Verhoeff" refId="B27" refString="Verhoeff KW (1928) Geophilomorphen-Beiträge und eine Lithobius - Form. Mitteilungen aus dem Zoologischen Museum in Berlin 14 (2): 227-286. (in German)" year="1928">Verhoeff 1928</bibRefCitation>
). This was the only information on its distribution until now. This is the first record of this species for
<collectingCountry id="87D4FFFE293B9B3DA1887BCA24425225" name="Georgia">Georgia</collectingCountry>
and also for the entire Caucasus.
</paragraph>
<caption id="53B2C1B0C371C7E03C47778F4876415D" ID-DOI="10.3897/caucasiana.3.e130238.figure3" ID-arpha="9A65BC1F-8AE6-5298-AC77-9C2A61867BD4" httpUri="https://binary.pensoft.net/fig/1130296" startId="F3">
<paragraph id="30BBAE4363D29A58FF76A1EC04BC376D">
<label id="3143F2EC6A376928CD672FC127AF2DBE">Figure 3.</label>
</paragraph>
<paragraph id="70BD7B02877A40DF5449750A6EF2E40E">
<taxonomicName id="6AD60F7CF16F51FF894046F79C425E0E" authority="Verhoeff, 1928" authorityName="Verhoeff" authorityYear="1928" class="Chilopoda" family="Dignathodontidae" genus="Henia" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="AF581DB19BA33056D43889EE6DE5E58F" italics="true">Henia hirsuta</emphasis>
<bibRefCitation id="88668D0750690C56104AC7E1157111A8" author="Verhoeff KW" refId="B27" refString="Verhoeff KW (1928) Geophilomorphen-Beitrage und eine Lithobius - Form. Mitteilungen aus dem Zoologischen Museum in Berlin 14 (2): 227 - 286. (in German)" year="1928">Verhoeff, 1928</bibRefCitation>
</taxonomicName>
, male, (a) head ventral view; (b) head and first two anterior sternites, ventral view; (c) posterior segments, ventral view; (d) segments showing sternal pores, ventral view. Scale bars: 0.2 mm (a, d); 0.5 mm (b, c).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>