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<document id="171F06B1649C688A213E5A5D8ABC4FDC" ID-DOI="10.5252/zoosystema2024v46a27" ID-ISSN="1638-9387" ID-Zenodo-Dep="13954689" ID-ZooBank="A46D3764-47EA-4E57-8AD8-14FEAB25AEA8" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1729334500832" checkinUser="plazi" docAuthor="Phuong, Lien Thi, Anh, Phuong Nguyen, Nguyen, Anh D., Dang, Hoa T., Michael, Seiki Yamane &amp; Engel, Michael S." docDate="2024" docId="03F88791FFA3FFCF044E077F80E8FBC5" docLanguage="en" docName="zoosystema.2024.46.27.pdf" docOrigin="Zoosystema 46 (27)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a27.pdf" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.9:Zoosystema.2020-.journal_article.type1" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article.type1" docStyleVersion="9" docTitle="Antepipona concava Nguyen, Dang &amp; A. D. Nguyen 2024, n. sp." docType="treatment" docVersion="2" lastPageNumber="711" masterDocId="FFC1FFE9FFAEFFDF004F010D8319FFEB" masterDocTitle="Taxonomic review of the potter wasp genus Antepipona de Saussure, 1855 (Hymenoptera, Vespidae, Eumeninae) from Vietnam, with descriptions of two new species and keys to the Oriental fauna" masterLastPageNumber="722" masterPageNumber="697" pageNumber="708" updateTime="1729334953978" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="D20A6C954C8A065F1F69A902CB3C8A01">Taxonomic review of the potter wasp genus Antepipona de Saussure, 1855 (Hymenoptera, Vespidae, Eumeninae) from Vietnam, with descriptions of two new species and keys to the Oriental fauna</mods:title>
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<mods:namePart id="1BEF807C4F0E2B46C5D33278774F29F3">Phuong, Lien Thi</mods:namePart>
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<mods:namePart id="A22E0C420BED8247DB4490047CA5EFCD">Anh, Phuong Nguyen</mods:namePart>
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<mods:namePart id="2F71E1EEBC83A9D5067E45E261681309">Nguyen, Anh D.</mods:namePart>
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<mods:namePart id="A97BFD190E726042F28F0B78E5CAD75B">Dang, Hoa T.</mods:namePart>
<mods:affiliation id="3BD676ECCA9B05EAF8955B024ED480D1">Insect Ecology Department, Institute of Ecology &amp; Biological Resources (IEBR), Vietnam Academy of Science &amp; Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi (Vietnam) and Graduate University of Science and Technology, Vietnam Academy of Science &amp; Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi (Vietnam) phuonglientit @ gmail. com (corresponding author: Lien Thi Phuong Nguyen)</mods:affiliation>
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<mods:namePart id="52B90DD401BE0E2DBC7512B2EB3DEFC8">Engel, Michael S.</mods:namePart>
<mods:affiliation id="CCAAC527DAE683CEDC86904054407CB5">Division of Invertebrate ZoÖlogy, American Museum of Natural History, 200 Central Park West, New York, New York 10024 - 5192 (United States)</mods:affiliation>
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<mods:date id="1FDF61B127222A5ACD0EF71412B23E57">2024</mods:date>
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<taxonomicName id="4C514D04FFA3FFD2044E077F87E4F946" authority="Nguyen, Dang &amp; A. D. Nguyen" authorityName="Nguyen, Dang &amp; A. D. Nguyen" authorityYear="2024" class="Insecta" family="Eumenidae" genus="Antepipona" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="708" phylum="Arthropoda" rank="species" species="concava" status="sp. nov.">
<emphasis id="B925EA95FFA3FFD2044E077F87C2F966" bold="true" box="[1025,1243,1650,1677]" italics="true" pageId="13" pageNumber="708">Antepipona concava</emphasis>
Nguyen, Dang &amp; A. D. Nguyen
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,
<taxonomicNameLabel id="A21657EEFFA3FFD20548079E865BF946" box="[1287,1346,1683,1709]" pageId="13" pageNumber="708" rank="species">n. sp.</taxonomicNameLabel>
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<subSubSection id="C34B650CFFA3FFD2046607BF868AF8EB" pageId="13" pageNumber="708" type="description">
<paragraph id="8BEE3687FFA3FFD2046607BF87ABF927" blockId="13.[922,1346,1650,1740]" box="[1065,1202,1714,1740]" pageId="13" pageNumber="708">
(
<figureCitation id="136A2A02FFA3FFD2047F07BF879BF927" box="[1072,1154,1714,1740]" captionStart="FIG" captionStartId="14.[134,145,1986,2004]" captionTargetBox="[188,1393,216,1919]" captionTargetId="figure-67@14.[589,791,1017,1218]" captionTargetPageId="14" captionText="FIG. 11. — Antepipona concava Nguyen, Dang &amp; A. D. Nguyen, n. sp., male holotype: A, head, facial view; B, antenna; C, mesosoma, dorsal view; D, head and mesosoma, lateral view; E, mesoscutellum, metanotum, and propodeum, posterior view; F, habitus, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13954721" httpUri="https://zenodo.org/record/13954721/files/figure.png" pageId="13" pageNumber="708">Figs 11</figureCitation>
;
<figureCitation id="136A2A02FFA3FFD204C107BF87B2F927" box="[1166,1195,1714,1740]" captionStart="FIG" captionStartId="15.[135,146,1386,1404]" captionTargetBox="[421,1177,215,1338]" captionTargetId="figure-313@15.[421,1177,215,1338]" captionTargetPageId="15" captionText="FIG. 12. — Antepipona concava Nguyen, Dang &amp; A. D. Nguyen, n. sp., genitalia: A, inner aspect of paramere with volsella and digitus; B, digitus; C, aedeagus, ventral view; D, aedeagus, lateral view. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13954723" httpUri="https://zenodo.org/record/13954723/files/figure.png" pageId="13" pageNumber="708">12</figureCitation>
)
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urn:lsid:zoobank.org:act:
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<subSubSection id="C34B650CFFA3FFD20362062F8661F863" pageId="13" pageNumber="708" type="materials_examined">
<paragraph id="8BEE3687FFA3FFD20362062F80FCF885" blockId="13.[811,1455,1825,2030]" pageId="13" pageNumber="708">
<materialsCitation id="3B393CDAFFA3FFD20362062F80FCF885" collectingDate="2018-06-22" collectionCode="IEBR" collectorName="Nguyen Quang Cuong" country="Vietnam" location="Cam Xuyen" municipality="Ke Go National Researve" pageId="13" pageNumber="708" specimenCount="1" specimenCount-male="1" stateProvince="Ha Tinh" typeStatus="holotype">
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. —
<emphasis id="B925EA95FFA3FFD2046D062C8798F8D2" bold="true" box="[1058,1153,1825,1849]" pageId="13" pageNumber="708">
<typeStatus id="54EA8825FFA3FFD2046D062C8798F8D2" box="[1058,1153,1825,1849]" pageId="13" pageNumber="708" type="holotype">Holotype</typeStatus>
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.
<emphasis id="B925EA95FFA3FFD204C3062E87FBF8D0" bold="true" box="[1164,1250,1827,1851]" pageId="13" pageNumber="708">
<collectingCountry id="F3467617FFA3FFD204C3062E87FBF8D0" box="[1164,1250,1827,1851]" name="Vietnam" pageId="13" pageNumber="708">Vietnam</collectingCountry>
</emphasis>
<specimenCount id="9D57FD0EFFA3FFD204B9062B8613F8D6" box="[1270,1290,1830,1853]" count="1" pageId="13" pageNumber="708" type="male"></specimenCount>
;
<collectingMunicipality id="6B8AACFDFFA3FFD2055A062F809FF8B8" pageId="13" pageNumber="708">Ke Go National Researve</collectingMunicipality>
,
<location id="8E8E605CFFA3FFD203DC06318708F8BF" LSID="urn:lsid:plazi:treatment:03F88791FFA3FFCF044E077F80E8FBC5:8E8E605CFFA3FFD203DC06318708F8BF" box="[915,1041,1852,1876]" country="Vietnam" municipality="Ke Go National Researve" name="Cam Xuyen" pageId="13" pageNumber="708" stateProvince="Ha Tinh">Cam Xuyen</location>
,
<collectingRegion id="4995F865FFA3FFD2045106318760F8BF" box="[1054,1145,1852,1876]" country="Vietnam" name="Ha Tinh" pageId="13" pageNumber="708">Ha Tinh</collectingRegion>
;
<date id="FFEF1047FFA3FFD204C906318619F8BF" box="[1158,1280,1852,1876]" pageId="13" pageNumber="708" value="2018-06-22">
<collectingDate id="EFABE9AFFFA3FFD204C906318619F8BF" box="[1158,1280,1852,1876]" pageId="13" pageNumber="708" value="2018-06-22">22.VI.2018</collectingDate>
</date>
;
<collectorName id="26A45351FFA3FFD205420631806BF885" pageId="13" pageNumber="708">Nguyen Quang Cuong</collectorName>
leg.;
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.
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<materialsCitation id="3B393CDAFFA3FFD20362067C866DF863" box="[813,1396,1905,1932]" collectingDate="2018-06-22" collectionCode="IEBR" collectorName="Nguyen Quang Cuong" country="Vietnam" location="Cam Xuyen" municipality="Ke Go National Researve" pageId="13" pageNumber="708" specimenCount="4" specimenCount-male="4" stateProvince="Ha Tinh" typeStatus="paratype">
<emphasis id="B925EA95FFA3FFD20362067C80F6F861" bold="true" box="[813,1007,1905,1930]" pageId="13" pageNumber="708">
<typeStatus id="54EA8825FFA3FFD20362067C8094F862" box="[813,909,1905,1929]" pageId="13" pageNumber="708" type="paratype">Paratypes</typeStatus>
.
<collectingCountry id="F3467617FFA3FFD203D6067F80F6F861" box="[921,1007,1906,1930]" name="Vietnam" pageId="13" pageNumber="708">Vietnam</collectingCountry>
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<specimenCount id="9D57FD0EFFA3FFD20449067C8733F867" box="[1030,1066,1905,1932]" count="4" pageId="13" pageNumber="708" type="male">4 ♂</specimenCount>
; same data as for holotype;
<collectionCode id="ED40AE42FFA3FFD20575067C866DF863" box="[1338,1396,1905,1928]" pageId="13" pageNumber="708">IEBR</collectionCode>
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.
</paragraph>
</subSubSection>
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. — This species can be distinguished from congeners by the following character combination: Inner compound eye margins strongly convergent ventrally, in anterior view
<emphasis id="B925EA95FFA3FFD204A306DB87E2F806" box="[1260,1275,2006,2029]" italics="true" pageId="13" pageNumber="708">c.</emphasis>
1.7× further apart from each other at vertex than at clypeus; clypeus with apical margin widely and deeply emarginate medially, forming rather blunt tooth on each side, distance between teeth nearly half width of clypeus between inner compound eye margins (
<emphasis id="B925EA95FFA1FFD00242072C8104F9D3" box="[525,541,1569,1592]" italics="true" pageId="15" pageNumber="710">c.</emphasis>
0.45× width of clypeus between inner compound eye margins); propodeum with deep concavities, the concavities shiny and with dense strong punctures, corner between dorsal and posterior surfaces angled, posterior surface without carinae; SII not bulging at base.
</paragraph>
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. 11. —
<taxonomicName id="4C514D04FFA0FFD100AE06CF8196F83F" authority="Nguyen, Dang &amp; A. D. Nguyen" authorityName="Nguyen, Dang &amp; A. D. Nguyen" authorityYear="2024" box="[225,655,1986,2004]" class="Insecta" family="Eumenidae" genus="Antepipona" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="709" phylum="Arthropoda" rank="species" species="concava" status="sp. nov.">
<emphasis id="B925EA95FFA0FFD100AE06CF8292F83F" box="[225,395,1986,2004]" italics="true" pageId="14" pageNumber="709">Antepipona concava</emphasis>
Nguyen, Dang &amp; A. D. Nguyen
</taxonomicName>
,
<taxonomicNameLabel id="A21657EEFFA0FFD102D806CF81D0F83F" box="[663,713,1986,2004]" pageId="14" pageNumber="709" rank="species">n. sp.</taxonomicNameLabel>
, male holotype:
<emphasis id="B925EA95FFA0FFD1031F06CF8044F83F" bold="true" box="[848,861,1986,2004]" pageId="14" pageNumber="709">A</emphasis>
, head, facial view;
<emphasis id="B925EA95FFA0FFD103B206CF8713F83F" bold="true" box="[1021,1034,1986,2004]" pageId="14" pageNumber="709">B</emphasis>
, antenna;
<emphasis id="B925EA95FFA0FFD1042D06CC8769F838" bold="true" box="[1122,1136,1985,2003]" pageId="14" pageNumber="709">C</emphasis>
, mesosoma, dorsal view;
<emphasis id="B925EA95FFA0FFD1050506CF8641F83F" bold="true" box="[1354,1368,1986,2004]" pageId="14" pageNumber="709">D</emphasis>
, head and mesosoma, lateral view;
<emphasis id="B925EA95FFA0FFD1011B06D58279F801" bold="true" box="[340,352,2008,2026]" pageId="14" pageNumber="709">E</emphasis>
, mesoscutellum, metanotum, and propodeum, posterior view;
<emphasis id="B925EA95FFA0FFD1032306D5806EF801" bold="true" box="[876,887,2008,2026]" pageId="14" pageNumber="709">F</emphasis>
, habitus, lateral view. Scale bars: 1 mm.
</paragraph>
</caption>
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<smallCapsWord id="8D08A05BFFA1FFD000C804678386FA90" baselines="1399,1400" box="[135,159,1386,1404]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="15" pageNumber="710">FIG</smallCapsWord>
. 12. —
<taxonomicName id="4C514D04FFA1FFD000AD0467818BFA97" authority="Nguyen, Dang &amp; A. D. Nguyen" authorityName="Nguyen, Dang &amp; A. D. Nguyen" authorityYear="2024" box="[226,658,1386,1404]" class="Insecta" family="Eumenidae" genus="Antepipona" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="710" phylum="Arthropoda" rank="species" species="concava" status="sp. nov.">
<emphasis id="B925EA95FFA1FFD000AD04678295FA97" box="[226,396,1386,1404]" italics="true" pageId="15" pageNumber="710">Antepipona concava</emphasis>
Nguyen, Dang &amp; A. D. Nguyen
</taxonomicName>
,
<taxonomicNameLabel id="A21657EEFFA1FFD002D5046781D4FA97" box="[666,717,1386,1404]" pageId="15" pageNumber="710" rank="species">n. sp.</taxonomicNameLabel>
, genitalia:
<emphasis id="B925EA95FFA1FFD0036B04678028FA97" bold="true" box="[804,817,1386,1404]" pageId="15" pageNumber="710">A</emphasis>
, inner aspect of paramere with volsella and digitus;
<emphasis id="B925EA95FFA1FFD004AA046787EBFA97" bold="true" box="[1253,1266,1386,1404]" pageId="15" pageNumber="710">B</emphasis>
, digitus;
<emphasis id="B925EA95FFA1FFD0050F04678657FA97" bold="true" box="[1344,1358,1386,1404]" pageId="15" pageNumber="710">C</emphasis>
, aedeagus, ventral view;
<emphasis id="B925EA95FFA1FFD000BB048C821BFA78" bold="true" box="[244,258,1409,1427]" pageId="15" pageNumber="710">D</emphasis>
, aedeagus, lateral view. Scale bar: 1 mm.
</paragraph>
</caption>
<subSubSection id="C34B650CFFA1FFD000CB07B682F3F939" box="[132,490,1722,1746]" pageId="15" pageNumber="710" type="distribution">
<paragraph id="8BEE3687FFA1FFD000CB07B682F3F939" blockId="15.[132,490,1722,1746]" box="[132,490,1722,1746]" pageId="15" pageNumber="710">
<smallCapsWord id="8D08A05BFFA1FFD000CB07B6820BF93A" baselines="1740,1741" box="[132,274,1723,1746]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Distribution" pageId="15" pageNumber="710">DISTRIBUTION</smallCapsWord>
. — Central
<collectingCountry id="F3467617FFA1FFD001C107B782FFF939" box="[398,486,1722,1746]" name="Vietnam" pageId="15" pageNumber="710">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C34B650CFFA1FFD000CB07E7810BF8DC" pageId="15" pageNumber="710" type="etymology">
<paragraph id="8BEE3687FFA1FFD000CB07E7810BF8DC" blockId="15.[131,775,1770,1847]" pageId="15" pageNumber="710">
<smallCapsWord id="8D08A05BFFA1FFD000CB07E783E7F8EB" baselines="1787,1788" box="[132,254,1770,1793]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Etymology" pageId="15" pageNumber="710">ETYMOLOGY</smallCapsWord>
. — The specific epithet is the Latin adjective
<emphasis id="B925EA95FFA1FFD002E007E68019F8E9" box="[687,768,1771,1794]" italics="true" pageId="15" pageNumber="710">concavus</emphasis>
, meaning, “concave”, and refers to the deep concavity of the posterior surface of the propodeum in this species.
</paragraph>
</subSubSection>
<subSubSection id="C34B650CFFA1FFCF00CB067F80E8FBC5" lastPageId="16" lastPageNumber="711" pageId="15" pageNumber="710" type="description">
<paragraph id="8BEE3687FFA1FFD000CB067F820CF861" blockId="15.[130,775,1906,2028]" box="[132,277,1906,1932]" pageId="15" pageNumber="710">
<smallCapsWord id="8D08A05BFFA1FFD000CB067F820CF861" baselines="1925,1925" box="[132,277,1906,1932]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="15" pageNumber="710">DESCRIPTION</smallCapsWord>
</paragraph>
<paragraph id="8BEE3687FFA1FFD000CB069C8234F840" blockId="15.[130,775,1906,2028]" box="[132,301,1937,1964]" pageId="15" pageNumber="710">
<emphasis id="B925EA95FFA1FFD000CB069C8234F840" box="[132,301,1937,1964]" italics="true" pageId="15" pageNumber="710">
Male (
<figureCitation id="136A2A02FFA1FFD00086069F823CF840" box="[201,293,1937,1964]" captionStart="FIG" captionStartId="14.[134,145,1986,2004]" captionTargetBox="[188,1393,216,1919]" captionTargetId="figure-67@14.[589,791,1017,1218]" captionTargetPageId="14" captionText="FIG. 11. — Antepipona concava Nguyen, Dang &amp; A. D. Nguyen, n. sp., male holotype: A, head, facial view; B, antenna; C, mesosoma, dorsal view; D, head and mesosoma, lateral view; E, mesoscutellum, metanotum, and propodeum, posterior view; F, habitus, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13954721" httpUri="https://zenodo.org/record/13954721/files/figure.png" pageId="15" pageNumber="710">Fig. 11F</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8BEE3687FFA1FFD000CB06BC8180F800" blockId="15.[130,775,1906,2028]" pageId="15" pageNumber="710">
<emphasis id="B925EA95FFA1FFD000CB06BC8231F820" bold="true" box="[132,296,1969,1995]" pageId="15" pageNumber="710">Measurements.</emphasis>
Body length 5.5-6.0 mm (
<typeStatus id="54EA8825FFA1FFD0027406BC8184F820" box="[571,669,1969,1995]" pageId="15" pageNumber="710" type="holotype">holotype</typeStatus>
:
<quantity id="4CA99B62FFA1FFD002EA06BC81E2F820" box="[677,763,1969,1996]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="15" pageNumber="710" unit="mm" value="5.7">5.7 mm</quantity>
); forewing length
<quantity id="4CA99B62FFA1FFD0017C06DC82ACF807" box="[307,437,2001,2028]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="5.8" metricValueMin="5.2" pageId="15" pageNumber="710" unit="mm" value="5.5" valueMax="5.8" valueMin="5.2">5.2-5.8 mm</quantity>
(
<typeStatus id="54EA8825FFA1FFD0018B06DC8131F800" box="[452,552,2001,2027]" pageId="15" pageNumber="710" type="holotype">holotype</typeStatus>
:
<quantity id="4CA99B62FFA1FFD0027D06DC8195F800" box="[562,652,2001,2028]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="15" pageNumber="710" unit="mm" value="5.5">5.5 mm</quantity>
).
</paragraph>
<paragraph id="8BEE3687FFA1FFCF036204DF8287FE5A" blockId="15.[813,1457,1490,2028]" lastBlockId="16.[130,777,215,433]" lastPageId="16" lastPageNumber="711" pageId="15" pageNumber="710">
<emphasis id="B925EA95FFA1FFD0036204DF8085FA06" bold="true" box="[813,924,1490,1517]" pageId="15" pageNumber="710">Structure.</emphasis>
Head in facial view transverse, 1.2× as wide as high (
<figureCitation id="136A2A02FFA1FFD0033D04FE80C3F9E6" box="[882,986,1523,1549]" captionStart="FIG" captionStartId="14.[134,145,1986,2004]" captionTargetBox="[188,1393,216,1919]" captionTargetId="figure-67@14.[589,791,1017,1218]" captionTargetPageId="14" captionText="FIG. 11. — Antepipona concava Nguyen, Dang &amp; A. D. Nguyen, n. sp., male holotype: A, head, facial view; B, antenna; C, mesosoma, dorsal view; D, head and mesosoma, lateral view; E, mesoscutellum, metanotum, and propodeum, posterior view; F, habitus, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13954721" httpUri="https://zenodo.org/record/13954721/files/figure.png" pageId="15" pageNumber="710">Fig. 11A</figureCitation>
). Vertex without cephalic fovea. Distance from posterior ocellus to posterior margin of vertex 1.4× distance from posterior ocellus to inner compound eye margin (
<figureCitation id="136A2A02FFA1FFD0037B075F8083F987" box="[820,922,1618,1644]" captionStart="FIG" captionStartId="14.[134,145,1986,2004]" captionTargetBox="[188,1393,216,1919]" captionTargetId="figure-67@14.[589,791,1017,1218]" captionTargetPageId="14" captionText="FIG. 11. — Antepipona concava Nguyen, Dang &amp; A. D. Nguyen, n. sp., male holotype: A, head, facial view; B, antenna; C, mesosoma, dorsal view; D, head and mesosoma, lateral view; E, mesoscutellum, metanotum, and propodeum, posterior view; F, habitus, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13954721" httpUri="https://zenodo.org/record/13954721/files/figure.png" pageId="15" pageNumber="710">Fig. 11C</figureCitation>
). Gena in lateral view narrower than compound eye, about 0.8× as wide as compound eye; occipital carina not complete, present only along length of gena, obsolete dorsally. Inner compound eye margins strongly convergent ventrally, in anterior view
<emphasis id="B925EA95FFA1FFD0041B07DE877CF907" box="[1108,1125,1747,1772]" italics="true" pageId="15" pageNumber="710">c.</emphasis>
1.7× further apart from each other at vertex than at clypeus (
<figureCitation id="136A2A02FFA1FFD0043007FF87F8F8E7" box="[1151,1249,1778,1804]" captionStart="FIG" captionStartId="14.[134,145,1986,2004]" captionTargetBox="[188,1393,216,1919]" captionTargetId="figure-67@14.[589,791,1017,1218]" captionTargetPageId="14" captionText="FIG. 11. — Antepipona concava Nguyen, Dang &amp; A. D. Nguyen, n. sp., male holotype: A, head, facial view; B, antenna; C, mesosoma, dorsal view; D, head and mesosoma, lateral view; E, mesoscutellum, metanotum, and propodeum, posterior view; F, habitus, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13954721" httpUri="https://zenodo.org/record/13954721/files/figure.png" pageId="15" pageNumber="710">Fig. 11A</figureCitation>
). Clypeus in lateral view gradually convex from base to apical margin; in frontal view slightly wider than high (
<figureCitation id="136A2A02FFA1FFD00438063C87C0F8A7" box="[1143,1241,1841,1868]" captionStart="FIG" captionStartId="14.[134,145,1986,2004]" captionTargetBox="[188,1393,216,1919]" captionTargetId="figure-67@14.[589,791,1017,1218]" captionTargetPageId="14" captionText="FIG. 11. — Antepipona concava Nguyen, Dang &amp; A. D. Nguyen, n. sp., male holotype: A, head, facial view; B, antenna; C, mesosoma, dorsal view; D, head and mesosoma, lateral view; E, mesoscutellum, metanotum, and propodeum, posterior view; F, habitus, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13954721" httpUri="https://zenodo.org/record/13954721/files/figure.png" pageId="15" pageNumber="710">Fig. 11A</figureCitation>
), with basal margin slightly concave medially and distinctly separated from antennal toruli; apical margin widely and deeply emarginate medially, forming rather blunt tooth on each side, distance between teeth nearly half width of clypeus between inner compound eye margins (
<emphasis id="B925EA95FFA1FFD0047106DF8756F800" box="[1086,1103,2002,2027]" italics="true" pageId="15" pageNumber="710">c.</emphasis>
0.45× width of clypeus between inner compound eye margins). Mandible with five prominent teeth. Antennal scape
<emphasis id="B925EA95FFBEFFCF013901F5829EFEFA" box="[374,391,248,273]" italics="true" pageId="16" pageNumber="711">c.</emphasis>
4× as long as its maximum width, slightly curved; FI
<emphasis id="B925EA95FFBEFFCF010200158247FEDA" box="[333,350,280,305]" italics="true" pageId="16" pageNumber="711">c.</emphasis>
2× longer than wide, FII slightly longer than wide, FIII-IV as wide as long, FV-FVIII wider than long, FX small, terminal flagellomere slightly curved, long and gradually narrow from base to apex,
<emphasis id="B925EA95FFBEFFCF0205007A8142FE7B" box="[586,603,375,400]" italics="true" pageId="16" pageNumber="711">c.</emphasis>
2.2× as long as its basal width (
<figureCitation id="136A2A02FFBEFFCF0160009B828BFE5A" box="[303,402,406,433]" captionStart="FIG" captionStartId="14.[134,145,1986,2004]" captionTargetBox="[188,1393,216,1919]" captionTargetId="figure-67@14.[589,791,1017,1218]" captionTargetPageId="14" captionText="FIG. 11. — Antepipona concava Nguyen, Dang &amp; A. D. Nguyen, n. sp., male holotype: A, head, facial view; B, antenna; C, mesosoma, dorsal view; D, head and mesosoma, lateral view; E, mesoscutellum, metanotum, and propodeum, posterior view; F, habitus, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13954721" httpUri="https://zenodo.org/record/13954721/files/figure.png" pageId="16" pageNumber="711">Fig. 11B</figureCitation>
).
</paragraph>
<paragraph id="8BEE3687FFBEFFCF00CB00DB823BFBE5" blockId="16.[130,777,470,1198]" pageId="16" pageNumber="711">
<emphasis id="B925EA95FFBEFFCF00CB00DB821CFE1B" bold="true" box="[132,261,470,496]" pageId="16" pageNumber="711">Mesosoma.</emphasis>
Longer than wide in dorsal view (
<figureCitation id="136A2A02FFBEFFCF02DE00DB81E3FE1B" box="[657,762,470,496]" captionStart="FIG" captionStartId="14.[134,145,1986,2004]" captionTargetBox="[188,1393,216,1919]" captionTargetId="figure-67@14.[589,791,1017,1218]" captionTargetPageId="14" captionText="FIG. 11. — Antepipona concava Nguyen, Dang &amp; A. D. Nguyen, n. sp., male holotype: A, head, facial view; B, antenna; C, mesosoma, dorsal view; D, head and mesosoma, lateral view; E, mesoscutellum, metanotum, and propodeum, posterior view; F, habitus, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13954721" httpUri="https://zenodo.org/record/13954721/files/figure.png" pageId="16" pageNumber="711">Fig. 11C</figureCitation>
). Pronotal carina fully developed, reaching ventral corner of pronotum, somewhat weak dorsally; anterior surface of pronotum with two close, deeply impressed pits medially, with dense strong punctures mediolaterally. Mesoscutum weakly convex, slightly longer than wide between tegulae, without depressed and oblique furrows apically (
<figureCitation id="136A2A02FFBEFFCF0273039B81BBFD5B" box="[572,674,662,688]" captionStart="FIG" captionStartId="14.[134,145,1986,2004]" captionTargetBox="[188,1393,216,1919]" captionTargetId="figure-67@14.[589,791,1017,1218]" captionTargetPageId="14" captionText="FIG. 11. — Antepipona concava Nguyen, Dang &amp; A. D. Nguyen, n. sp., male holotype: A, head, facial view; B, antenna; C, mesosoma, dorsal view; D, head and mesosoma, lateral view; E, mesoscutellum, metanotum, and propodeum, posterior view; F, habitus, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13954721" httpUri="https://zenodo.org/record/13954721/files/figure.png" pageId="16" pageNumber="711">Fig. 11C</figureCitation>
). Disc of mesoscutellum convex, in lateral view at same level of mesoscutum (
<figureCitation id="136A2A02FFBEFFCF00BE03D8824FFD1B" box="[241,342,725,752]" captionStart="FIG" captionStartId="14.[134,145,1986,2004]" captionTargetBox="[188,1393,216,1919]" captionTargetId="figure-67@14.[589,791,1017,1218]" captionTargetPageId="14" captionText="FIG. 11. — Antepipona concava Nguyen, Dang &amp; A. D. Nguyen, n. sp., male holotype: A, head, facial view; B, antenna; C, mesosoma, dorsal view; D, head and mesosoma, lateral view; E, mesoscutellum, metanotum, and propodeum, posterior view; F, habitus, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13954721" httpUri="https://zenodo.org/record/13954721/files/figure.png" pageId="16" pageNumber="711">Fig. 11D</figureCitation>
), narrowly depressed basally, with a row of short longitudinal carinae basally and apically. Metanotum weakly convex, with two large and sharp tubercles, distance between tubercles slightly longer than distant from tubercle to lateral margin of metanotum (
<figureCitation id="136A2A02FFBEFFCF01980258812AFC84" box="[471,563,853,879]" captionStart="FIG" captionStartId="14.[134,145,1986,2004]" captionTargetBox="[188,1393,216,1919]" captionTargetId="figure-67@14.[589,791,1017,1218]" captionTargetPageId="14" captionText="FIG. 11. — Antepipona concava Nguyen, Dang &amp; A. D. Nguyen, n. sp., male holotype: A, head, facial view; B, antenna; C, mesosoma, dorsal view; D, head and mesosoma, lateral view; E, mesoscutellum, metanotum, and propodeum, posterior view; F, habitus, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13954721" httpUri="https://zenodo.org/record/13954721/files/figure.png" pageId="16" pageNumber="711">Fig. 11E</figureCitation>
). Propodeal dorsum below level with metanotum; declivity of propodeum with strong carina medially; concavity deep and wide, without short carinae (
<figureCitation id="136A2A02FFBEFFCF016C02B8829CFC24" box="[291,389,949,975]" captionStart="FIG" captionStartId="14.[134,145,1986,2004]" captionTargetBox="[188,1393,216,1919]" captionTargetId="figure-67@14.[589,791,1017,1218]" captionTargetPageId="14" captionText="FIG. 11. — Antepipona concava Nguyen, Dang &amp; A. D. Nguyen, n. sp., male holotype: A, head, facial view; B, antenna; C, mesosoma, dorsal view; D, head and mesosoma, lateral view; E, mesoscutellum, metanotum, and propodeum, posterior view; F, habitus, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13954721" httpUri="https://zenodo.org/record/13954721/files/figure.png" pageId="16" pageNumber="711">Fig. 11E</figureCitation>
), corner between lateral and dorsal surfaces angulate, corner between dorsal and posterior surfaces angulate.
</paragraph>
<paragraph id="8BEE3687FFBEFFCF00CB051982B7FB45" blockId="16.[130,777,470,1198]" pageId="16" pageNumber="711">
Metasomal segment I narrower than segment II (
<figureCitation id="136A2A02FFBEFFCF0230051981C3FBC5" box="[639,730,1044,1070]" captionStart="FIG" captionStartId="14.[134,145,1986,2004]" captionTargetBox="[188,1393,216,1919]" captionTargetId="figure-67@14.[589,791,1017,1218]" captionTargetPageId="14" captionText="FIG. 11. — Antepipona concava Nguyen, Dang &amp; A. D. Nguyen, n. sp., male holotype: A, head, facial view; B, antenna; C, mesosoma, dorsal view; D, head and mesosoma, lateral view; E, mesoscutellum, metanotum, and propodeum, posterior view; F, habitus, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13954721" httpUri="https://zenodo.org/record/13954721/files/figure.png" pageId="16" pageNumber="711">Fig. 11F</figureCitation>
); TI conical in shape, in dorsal view almost half as long as wide; TII wider than long, about 1.1× as wide as long in dorsal view; SII depressed at base, in lateral view slightly convex from base to apical margin.
</paragraph>
<paragraph id="8BEE3687FFBEFFCF00CB05DE865BFE1A" blockId="16.[130,777,1235,2027]" lastBlockId="16.[811,1458,215,497]" pageId="16" pageNumber="711">
<emphasis id="B925EA95FFBEFFCF00CB05DE8208FB05" bold="true" box="[132,273,1235,1262]" pageId="16" pageNumber="711">Sculpturing.</emphasis>
Clypeus with dense strong punctures, interspaces between punctures smooth, larger than puncture diameter at lateral parts, each puncture bearing a short bristle. Frons densely covered with coarse flat-bottomed punctures, interspaces between punctures raised to form reticulations. Vertex with punctures similar with those on frons, but larger; gena with large strong punctures, punctures weaker and smaller near base; occipital carina weakly widened laterally (
<figureCitation id="136A2A02FFBEFFCF009E04DE8225FA06" box="[209,316,1491,1517]" captionStart="FIG" captionStartId="14.[134,145,1986,2004]" captionTargetBox="[188,1393,216,1919]" captionTargetId="figure-67@14.[589,791,1017,1218]" captionTargetPageId="14" captionText="FIG. 11. — Antepipona concava Nguyen, Dang &amp; A. D. Nguyen, n. sp., male holotype: A, head, facial view; B, antenna; C, mesosoma, dorsal view; D, head and mesosoma, lateral view; E, mesoscutellum, metanotum, and propodeum, posterior view; F, habitus, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13954721" httpUri="https://zenodo.org/record/13954721/files/figure.png" pageId="16" pageNumber="711">Fig. 11D</figureCitation>
). Pronotum with punctures coarser than those on vertex, interspaces between punctures thicker and raised to form reticulation. Mesoscutum covered with coarse punctures, punctures equal in size, smaller than those on pronotum, interspaces between punctures smooth, raised to form reticulations; mesoscutellum with punctures similar to those on pronotum, punctures on metanotum smaller than those on mesoscutellum. Mesepisternum with flat-bottomed punctures similar to those on pronotum posterodorsally, smooth anteroventrally; border between posterodorsal and anteroventral parts distinct, without epicnemial carina. Dorsal part of metapleuron largely smooth and with several short weak striae, ventral part with sparse shallow punctures. Propodeum with coarse punctures dorsally, interspaces between punctures strongly raised to form reticulation, punctures much shallower and weaker laterally, dorso-lateral margin of propodeum somewhat angled; posterior surface shiny, with dense strong punctures.Tegula with minute punctures. Metasomal TI covered with dense strong punctures, punctures on lateral and apical margins sparser and weaker; punctures on TII smaller and shallower than those on TI, punctures on apical margin of TIII-IV larger than those on apical margin of TII, punctures on apical margin of TV-VI smaller than those on apical margin of TIII-TIV, punctures on TVII and SVII small and sparse, punctures on SII clear, deeper than those on TII, interspaces with minute punctures.
</paragraph>
<paragraph id="8BEE3687FFBEFFCF0362031B80E8FBC5" blockId="16.[811,1458,534,1070]" pageId="16" pageNumber="711">
<emphasis id="B925EA95FFBEFFCF0362031B806BFDDB" bold="true" box="[813,882,534,560]" pageId="16" pageNumber="711">Color.</emphasis>
Body black, with following parts yellow: clypeus except apical margin, a large spot on frons, a large mark along inner orbit to clypeus, mandible except base and teeth, antennal scape except a longitudinal black strip dorsally, two spots on vertex below compound eye margins, two large spots on pronotum, two large spots on mesoscutellum and two smaller spots on metanotum, a large spot on mesopleuron, two very large spots on propodeum laterally, tegulae except large brown-black spots centrally and parategulae, two large spots on TI laterally and apical margin, two large spots laterally and a thick band apically on TII, bands on TIII and TIV laterally, two small spots at lower corner laterally on SIII and SIV, apical margin of TV-TVI, coxae of all legs except dorsal part black, all femora except dorsal part brown on profemur and black on mesofemur and apex of metafemur. Valvulae brown. Wing membrane transparent, veins dark brown (
<figureCitation id="136A2A02FFBEFFCF03CB051980FDFBC5" box="[900,996,1044,1070]" captionStart="FIG" captionStartId="14.[134,145,1986,2004]" captionTargetBox="[188,1393,216,1919]" captionTargetId="figure-67@14.[589,791,1017,1218]" captionTargetPageId="14" captionText="FIG. 11. — Antepipona concava Nguyen, Dang &amp; A. D. Nguyen, n. sp., male holotype: A, head, facial view; B, antenna; C, mesosoma, dorsal view; D, head and mesosoma, lateral view; E, mesoscutellum, metanotum, and propodeum, posterior view; F, habitus, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13954721" httpUri="https://zenodo.org/record/13954721/files/figure.png" pageId="16" pageNumber="711">Fig. 11F</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="C2ACAB56587AE2278A926F5FC49FABAA" ID-DOI="10.5252/zoosystema2024v46a27" ID-ISSN="1638-9387" ID-Zenodo-Dep="13954689" ID-ZooBank="A46D3764-47EA-4E57-8AD8-14FEAB25AEA8" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1729334500832" checkinUser="plazi" docAuthor="Phuong, Lien Thi, Anh, Phuong Nguyen, Nguyen, Anh D., Dang, Hoa T., Michael, Seiki Yamane &amp; Engel, Michael S." docDate="2024" docId="03F88791FFBFFFCC011401DA8252F8C7" docLanguage="en" docName="zoosystema.2024.46.27.pdf" docOrigin="Zoosystema 46 (27)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a27.pdf" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.9:Zoosystema.2020-.journal_article.type1" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article.type1" docStyleVersion="9" docTitle="Antepipona crenula Nguyen, Yamane &amp; Engel 2024, n. sp." docType="treatment" docVersion="2" lastPageNumber="712" masterDocId="FFC1FFE9FFAEFFDF004F010D8319FFEB" masterDocTitle="Taxonomic review of the potter wasp genus Antepipona de Saussure, 1855 (Hymenoptera, Vespidae, Eumeninae) from Vietnam, with descriptions of two new species and keys to the Oriental fauna" masterLastPageNumber="722" masterPageNumber="697" pageNumber="712" updateTime="1729334953978" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="08CC0179AC0B3B571A453906673EAF62">Taxonomic review of the potter wasp genus Antepipona de Saussure, 1855 (Hymenoptera, Vespidae, Eumeninae) from Vietnam, with descriptions of two new species and keys to the Oriental fauna</mods:title>
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<mods:namePart id="B7189A4542004AC5B21AC652089F7EAB">Phuong, Lien Thi</mods:namePart>
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<mods:namePart id="6842984D662869A3CC9DF191C3F8B541">Dang, Hoa T.</mods:namePart>
<mods:affiliation id="70F9D674265A737399D5AA1827DA03C5">Insect Ecology Department, Institute of Ecology &amp; Biological Resources (IEBR), Vietnam Academy of Science &amp; Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi (Vietnam) and Graduate University of Science and Technology, Vietnam Academy of Science &amp; Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi (Vietnam) phuonglientit @ gmail. com (corresponding author: Lien Thi Phuong Nguyen)</mods:affiliation>
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<mods:affiliation id="2EEC745CD6246EF9F67D5B7CA9BF01BA">Division of Invertebrate ZoÖlogy, American Museum of Natural History, 200 Central Park West, New York, New York 10024 - 5192 (United States)</mods:affiliation>
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<taxonomicName id="4C514D04FFBFFFCE011401DA8128FEFA" authority="Nguyen, Yamane &amp; Engel" authorityName="Nguyen, Yamane &amp; Engel" authorityYear="2024" class="Insecta" family="Eumenidae" genus="Antepipona" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="712" phylum="Arthropoda" rank="species" species="crenula" status="sp. nov.">
<emphasis id="B925EA95FFBFFFCE011401DA8129FF19" bold="true" box="[347,560,215,242]" italics="true" pageId="17" pageNumber="712">Antepipona crenula</emphasis>
Nguyen, Yamane &amp; Engel
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,
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(
<figureCitation id="136A2A02FFBFFFCE01C7001A82C0FEDA" box="[392,473,279,305]" captionStart="FIG" captionStartId="18.[132,143,1985,2003]" captionTargetBox="[189,1361,216,1873]" captionTargetId="figure-15@18.[189,1361,216,1874]" captionTargetPageId="18" captionText="FIG. 13. — Antepipona crenula Nguyen, Yamane &amp; Engel, n. sp., male holotype: A, head, facial view; B, head, dorsal view; C, antenna; D, pronotum, frontal view; E, mesosoma, dorsal view; F, metanotum and propodeum, posterior view; G, habitus, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13954727" httpUri="https://zenodo.org/record/13954727/files/figure.png" pageId="17" pageNumber="712">Figs 13</figureCitation>
;
<figureCitation id="136A2A02FFBFFFCE01AA001A8118FEDA" box="[485,513,279,305]" captionStart="FIG" captionStartId="19.[132,143,1397,1415]" captionTargetBox="[438,1152,215,1344]" captionTargetId="figure-17@19.[438,1152,215,1344]" captionTargetPageId="19" captionText="FIG. 14. — Antepipona crenula Nguyen, Yamane &amp; Engel, n. sp., genitalia: A, inner aspect of paramere with volsella and digitus; B, digitus; C, aedeagus, ventral view; D, aedeagus, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13954729" httpUri="https://zenodo.org/record/13954729/files/figure.png" pageId="17" pageNumber="712">14</figureCitation>
)
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<materialsCitation id="3B393CDAFFBFFFCE00CB008583E8FE3F" collectingDate="2009-09-11" collectionCode="IEBR" collectorName="Nguyen Huu Thao" country="Vietnam" location="Xuan Son" municipality="Xuan Son National Park" pageId="17" pageNumber="712" specimenCount="1" specimenCount-male="1" stateProvince="Phu Tho" typeStatus="holotype">
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. —
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<typeStatus id="54EA8825FFBFFFCE01CF008582C6FE4B" box="[384,479,392,416]" pageId="17" pageNumber="712" type="holotype">Holotype</typeStatus>
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.
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;
<collectingMunicipality id="6B8AACFDFFBFFFCE0235008583E8FE51" pageId="17" pageNumber="712">Xuan Son National Park</collectingMunicipality>
,
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,
<collectingRegion id="4995F865FFBFFFCE012500AF82DAFE51" box="[362,451,418,442]" country="Vietnam" name="Phu Tho" pageId="17" pageNumber="712">Phu Tho</collectingRegion>
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<collectingDate id="EFABE9AFFFBFFFCE018200AF8158FE51" box="[461,577,418,442]" pageId="17" pageNumber="712" value="2009-09-11">11.IX.2009</collectingDate>
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<collectorName id="26A45351FFBFFFCE020300AE801EFE51" box="[588,775,418,443]" pageId="17" pageNumber="712">Nguyen Huu Thao</collectorName>
leg.;
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.
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<paragraph id="8BEE3687FFBFFFCE00CB00E38293FCB6" blockId="17.[130,777,494,862]" pageId="17" pageNumber="712">
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. — This species can be distinguished from congeners by the following character combination: Clypeus in frontal view slightly wider than high; inner compound eye margins strongly convergent ventrally, in anterior view slightly more than 1.5× further apart from each other at vertex than at clypeus; mesoscutum shorter than wide between tegulae (
<emphasis id="B925EA95FFBFFFCE0181037E82C4FD61" box="[462,477,627,650]" italics="true" pageId="17" pageNumber="712">c.</emphasis>
0.9× shorter than wide between tegulae), with faint short longitudinal carina laterally; metanotum weakly convex, with two large sharp tubercles; propodeum with slightly oblique short carinae, with corner between lateral and dorsal surfaces rounded, corner between dorsal and posterior surfaces angled; SII depressed at base, in lateral view convex from base to apical margin. Penis valve about 1.7× as long as basal apodeme, in profile apical part strongly produced into a large rounded lobe, with apical margin serrate.
</paragraph>
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<emphasis id="B925EA95FFBFFFCE00CB02B88228FC24" box="[132,305,949,975]" italics="true" pageId="17" pageNumber="712">
Male (
<figureCitation id="136A2A02FFBFFFCE008602B88230FC24" box="[201,297,949,975]" captionStart="FIG" captionStartId="18.[132,143,1985,2003]" captionTargetBox="[189,1361,216,1873]" captionTargetId="figure-15@18.[189,1361,216,1874]" captionTargetPageId="18" captionText="FIG. 13. — Antepipona crenula Nguyen, Yamane &amp; Engel, n. sp., male holotype: A, head, facial view; B, head, dorsal view; C, antenna; D, pronotum, frontal view; E, mesosoma, dorsal view; F, metanotum and propodeum, posterior view; G, habitus, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13954727" httpUri="https://zenodo.org/record/13954727/files/figure.png" pageId="17" pageNumber="712">Fig. 13G</figureCitation>
)
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<emphasis id="B925EA95FFBFFFCE00CB02D88233FC04" bold="true" box="[132,298,981,1007]" pageId="17" pageNumber="712">Measurements.</emphasis>
Body length
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; forewing length
<quantity id="4CA99B62FFBFFFCE02E302D9801AFC04" box="[684,771,980,1007]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="17" pageNumber="712" unit="mm" value="6.5">6.5 mm</quantity>
.
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<emphasis id="B925EA95FFBFFFCE00CD051983F6FBC4" bold="true" box="[130,239,1044,1071]" pageId="17" pageNumber="712">Structure.</emphasis>
Head in facial view transverse, 1.2× as wide as high (
<figureCitation id="136A2A02FFBFFFCE00C5053983F0FBA5" box="[138,233,1076,1102]" captionStart="FIG" captionStartId="18.[132,143,1985,2003]" captionTargetBox="[189,1361,216,1873]" captionTargetId="figure-15@18.[189,1361,216,1874]" captionTargetPageId="18" captionText="FIG. 13. — Antepipona crenula Nguyen, Yamane &amp; Engel, n. sp., male holotype: A, head, facial view; B, head, dorsal view; C, antenna; D, pronotum, frontal view; E, mesosoma, dorsal view; F, metanotum and propodeum, posterior view; G, habitus, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13954727" httpUri="https://zenodo.org/record/13954727/files/figure.png" pageId="17" pageNumber="712">Fig. 13A</figureCitation>
). Vertex without cephalic fovea. Distance from posterior ocellus to posterior margin of vertex 1.2× distance from posterior ocellus to inner compound eye margin (
<figureCitation id="136A2A02FFBFFFCE02D8057981E1FB65" box="[663,760,1140,1166]" captionStart="FIG" captionStartId="18.[132,143,1985,2003]" captionTargetBox="[189,1361,216,1873]" captionTargetId="figure-15@18.[189,1361,216,1874]" captionTargetPageId="18" captionText="FIG. 13. — Antepipona crenula Nguyen, Yamane &amp; Engel, n. sp., male holotype: A, head, facial view; B, head, dorsal view; C, antenna; D, pronotum, frontal view; E, mesosoma, dorsal view; F, metanotum and propodeum, posterior view; G, habitus, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13954727" httpUri="https://zenodo.org/record/13954727/files/figure.png" pageId="17" pageNumber="712">Fig. 13B</figureCitation>
). Gena in lateral view narrower than compound eye, about 0.8× as wide as compound eye; occipital carina not complete, present along length of gena laterally, obsolete dorsally. Inner compound eye margins strongly convergent ventrally, in anterior view slightly more than 1.5× further apart from each other at vertex than at clypeus (
<figureCitation id="136A2A02FFBFFFCE019B043E812EFAA6" box="[468,567,1331,1357]" captionStart="FIG" captionStartId="18.[132,143,1985,2003]" captionTargetBox="[189,1361,216,1873]" captionTargetId="figure-15@18.[189,1361,216,1874]" captionTargetPageId="18" captionText="FIG. 13. — Antepipona crenula Nguyen, Yamane &amp; Engel, n. sp., male holotype: A, head, facial view; B, head, dorsal view; C, antenna; D, pronotum, frontal view; E, mesosoma, dorsal view; F, metanotum and propodeum, posterior view; G, habitus, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13954727" httpUri="https://zenodo.org/record/13954727/files/figure.png" pageId="17" pageNumber="712">Fig. 13A</figureCitation>
). Clypeus in lateral view gradually convex from base to middle, then straight to apical margin; in frontal view slightly wider than high (
<figureCitation id="136A2A02FFBFFFCE00C1049E83E8FA46" box="[142,241,1427,1453]" captionStart="FIG" captionStartId="18.[132,143,1985,2003]" captionTargetBox="[189,1361,216,1873]" captionTargetId="figure-15@18.[189,1361,216,1874]" captionTargetPageId="18" captionText="FIG. 13. — Antepipona crenula Nguyen, Yamane &amp; Engel, n. sp., male holotype: A, head, facial view; B, head, dorsal view; C, antenna; D, pronotum, frontal view; E, mesosoma, dorsal view; F, metanotum and propodeum, posterior view; G, habitus, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13954727" httpUri="https://zenodo.org/record/13954727/files/figure.png" pageId="17" pageNumber="712">Fig. 13A</figureCitation>
), with basal margin slightly convex medially and distinctly separated from antennal toruli; apical margin narrowly and deeply emarginate medially, forming slightly sharp tooth on each side, distance between teeth nearly 1/3 width of clypeus between inner compound eye margins (
<emphasis id="B925EA95FFBFFFCE02E0071E81D9F9C7" box="[687,704,1555,1580]" italics="true" pageId="17" pageNumber="712">c.</emphasis>
0.30× width of clypeus between inner compound eye margins). Mandible with five prominent teeth. Antennal scape 3.7× as long as its maximum width, slightly curved; FI 1.5× longer than wide, FII slightly longer than wide, FIII-IV as wide as long, FV-FVIII wider than long, FX small, terminal flagellomere slightly curved, long and gradually narrowed from base to apex, 2× as long as its basal width.
</paragraph>
<paragraph id="8BEE3687FFBFFFCE00CB063F87AAFD7B" blockId="17.[130,777,1841,2028]" lastBlockId="17.[811,1457,215,784]" pageId="17" pageNumber="712">
<emphasis id="B925EA95FFBFFFCE00CB063F821FF8A7" bold="true" box="[132,262,1842,1868]" pageId="17" pageNumber="712">Mesosoma.</emphasis>
Longer than wide in dorsal view (
<figureCitation id="136A2A02FFBFFFCE02DC063C81E0F8A7" box="[659,761,1841,1868]" captionStart="FIG" captionStartId="18.[132,143,1985,2003]" captionTargetBox="[189,1361,216,1873]" captionTargetId="figure-15@18.[189,1361,216,1874]" captionTargetPageId="18" captionText="FIG. 13. — Antepipona crenula Nguyen, Yamane &amp; Engel, n. sp., male holotype: A, head, facial view; B, head, dorsal view; C, antenna; D, pronotum, frontal view; E, mesosoma, dorsal view; F, metanotum and propodeum, posterior view; G, habitus, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13954727" httpUri="https://zenodo.org/record/13954727/files/figure.png" pageId="17" pageNumber="712">Fig. 13E</figureCitation>
). Pronotal carina developed at lateral sides, reaching ventral corner of pronotum, obsoleted dorsally, in dorsal view slightly produced at lateral corners; anterior surface of pronotum with two closely, deeply impressed pits medially, with dense strong punctures mediolaterally (
<figureCitation id="136A2A02FFBFFFCE01B706DC817AF800" box="[504,611,2001,2027]" captionStart="FIG" captionStartId="18.[132,143,1985,2003]" captionTargetBox="[189,1361,216,1873]" captionTargetId="figure-15@18.[189,1361,216,1874]" captionTargetPageId="18" captionText="FIG. 13. — Antepipona crenula Nguyen, Yamane &amp; Engel, n. sp., male holotype: A, head, facial view; B, head, dorsal view; C, antenna; D, pronotum, frontal view; E, mesosoma, dorsal view; F, metanotum and propodeum, posterior view; G, habitus, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13954727" httpUri="https://zenodo.org/record/13954727/files/figure.png" pageId="17" pageNumber="712">Fig. 13D</figureCitation>
). Mesoscutum convex, shorter than wide between tegulae (
<emphasis id="B925EA95FFBFFFCE055D01D5863AFF1A" box="[1298,1315,216,241]" italics="true" pageId="17" pageNumber="712">c.</emphasis>
0.9× shorter than wide between tegulae), with faint short longitudinal carina laterally (
<figureCitation id="136A2A02FFBFFFCE03AF001A875DFEDA" box="[992,1092,279,305]" captionStart="FIG" captionStartId="18.[132,143,1985,2003]" captionTargetBox="[189,1361,216,1873]" captionTargetId="figure-15@18.[189,1361,216,1874]" captionTargetPageId="18" captionText="FIG. 13. — Antepipona crenula Nguyen, Yamane &amp; Engel, n. sp., male holotype: A, head, facial view; B, head, dorsal view; C, antenna; D, pronotum, frontal view; E, mesosoma, dorsal view; F, metanotum and propodeum, posterior view; G, habitus, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13954727" httpUri="https://zenodo.org/record/13954727/files/figure.png" pageId="17" pageNumber="712">Fig. 13E</figureCitation>
). Disc of mesoscutellum convex, in lateral view not at same level of mesoscutum (
<figureCitation id="136A2A02FFBFFFCE0571003A86BAFEBA" box="[1342,1443,311,337]" captionStart="FIG" captionStartId="18.[132,143,1985,2003]" captionTargetBox="[189,1361,216,1873]" captionTargetId="figure-15@18.[189,1361,216,1874]" captionTargetPageId="18" captionText="FIG. 13. — Antepipona crenula Nguyen, Yamane &amp; Engel, n. sp., male holotype: A, head, facial view; B, head, dorsal view; C, antenna; D, pronotum, frontal view; E, mesosoma, dorsal view; F, metanotum and propodeum, posterior view; G, habitus, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13954727" httpUri="https://zenodo.org/record/13954727/files/figure.png" pageId="17" pageNumber="712">Fig. 13G</figureCitation>
), narrowly depressed basally, with a row of short longitudinal carinae basally and apically, with longitudinal carina laterally. Metanotum weakly convex, with two large sharp tubercles, distance between tubercles slightly longer than distance from tubercle to lateral margin of metanotum. Propodeal dorsum below level with metanotum; declivity of propodeum with narrow longitudinal carina medially, with slightly oblique short carinae (
<figureCitation id="136A2A02FFBFFFCE0381033B8729FDBB" box="[974,1072,566,592]" captionStart="FIG" captionStartId="18.[132,143,1985,2003]" captionTargetBox="[189,1361,216,1873]" captionTargetId="figure-15@18.[189,1361,216,1874]" captionTargetPageId="18" captionText="FIG. 13. — Antepipona crenula Nguyen, Yamane &amp; Engel, n. sp., male holotype: A, head, facial view; B, head, dorsal view; C, antenna; D, pronotum, frontal view; E, mesosoma, dorsal view; F, metanotum and propodeum, posterior view; G, habitus, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13954727" httpUri="https://zenodo.org/record/13954727/files/figure.png" pageId="17" pageNumber="712">Fig. 13F</figureCitation>
); concavity deep and wide, corner between lateral and dorsal surfaces rounded, corner between dorsal and posterior surfaces angled.
</paragraph>
<paragraph id="8BEE3687FFBFFFCE0362039B865AFCE4" blockId="17.[811,1457,215,784]" pageId="17" pageNumber="712">
Metasomal segment I narrower than segment II (
<figureCitation id="136A2A02FFBFFFCE0572039B86B8FD5B" box="[1341,1441,662,688]" captionStart="FIG" captionStartId="18.[132,143,1985,2003]" captionTargetBox="[189,1361,216,1873]" captionTargetId="figure-15@18.[189,1361,216,1874]" captionTargetPageId="18" captionText="FIG. 13. — Antepipona crenula Nguyen, Yamane &amp; Engel, n. sp., male holotype: A, head, facial view; B, head, dorsal view; C, antenna; D, pronotum, frontal view; E, mesosoma, dorsal view; F, metanotum and propodeum, posterior view; G, habitus, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13954727" httpUri="https://zenodo.org/record/13954727/files/figure.png" pageId="17" pageNumber="712">Fig. 13G</figureCitation>
); TI conical in shape, in dorsal view 1.6× as wide as long; TII wider than long, 1.2× as wide as long; SII depressed at base, in lateral view convex from base to apical margin.
</paragraph>
<paragraph id="8BEE3687FFBFFFCE036202388735F8C7" blockId="17.[811,1458,821,1836]" pageId="17" pageNumber="712">
<emphasis id="B925EA95FFBFFFCE0362023880A0FCA4" bold="true" box="[813,953,821,847]" pageId="17" pageNumber="712">Sculpturing.</emphasis>
Clypeus with dense strong punctures, interspaces between punctures smooth, each puncture bearing a short bristle. Frons densely covered with coarse flat-bottomed punctures, interspaces between punctures raised to form reticulation. Vertex with punctures similar with those on frons, but coarser; gena with strong and large punctures; occipital carina not widened laterally. Pronotum with punctures similar to those on vertex. Mesoscutum covered with coarse punctures larger than those on pronotum, interspaces between punctures smooth, raised to form reticulation; mesoscutellum with punctures similar to those on pronotum, punctures on metanotum smaller than those on mesoscutellum. Mesepisternum with flat-bottomed punctures, punctures similar to those on pronotum posterodorsally, with sparse strong punctures anteroventrally; border between posterodorsal and anteroventral parts distinct, looking like a carina, without epicnemial carina.Dorsal part of metapleuron largely smooth, with several short and weak striae, ventral part with dense strong punctures. Propodeum with coarse punctures dorsally, punctures much shallower and weaker to form weak transverse striae laterally, interspaces between punctures strongly raised to form reticulation, dorso-lateral margin of propodeum somewhat angled; posterior surface shiny, with dense strong punctures. Tegula with dense small punctures. Metasomal TI covered with dense strong punctures, punctures on lateral and apical margins sparser and weaker; punctures on TII smaller and shallower than those on TI, punctures on apical margin of TIII larger than those on apical margin of TII, punctures on apical margin of TIV-VI smaller than those on apical margin of TIII-TIV, punctures on TVII and SVII small and sparse, punctures on SII clear, deeper than those on TII, interspaces with minute punctures.
</paragraph>
<paragraph id="8BEE3687FFBFFFCC0362065C8252F8C7" blockId="17.[811,1458,1873,2028]" lastBlockId="19.[130,777,1491,1836]" lastPageId="19" lastPageNumber="714" pageId="17" pageNumber="712">
<emphasis id="B925EA95FFBFFFCE0362065C806BF887" bold="true" box="[813,882,1873,1900]" pageId="17" pageNumber="712">Color.</emphasis>
Body black, with following parts yellow: clypeus except small brown spot at central and black lateral margins, a large spot on frons, a large mark along inner orbit extended close to clypeus, mandible except base and teeth, antennal scape except a longitudinal black strip dorsally, flagellomeres VII-X beneath, last flagellomere entirely, two spots on vertex below compound eye margins, a thick transverse strip on pronotum, scutellum except black margins, metanotum except black margins, a large spot on mesopleuron, two large spots on propodeum laterally, parategulae, apical margin of TI, two large spots laterally near base and a thick band apically on TII, bands on TIII-TV, SIII-SV apically, tibiae and basitarsi of all legs, and femora of all legs except 1/3 basally. Tegulae dark brown with a black mark medially. Valvulae brown. Wing membrane with apical part dark infumate, veins dark brown (
<figureCitation id="136A2A02FFBDFFCC0096061F8227F8C7" box="[217,318,1810,1836]" captionStart="FIG" captionStartId="18.[132,143,1985,2003]" captionTargetBox="[189,1361,216,1873]" captionTargetId="figure-15@18.[189,1361,216,1874]" captionTargetPageId="18" captionText="FIG. 13. — Antepipona crenula Nguyen, Yamane &amp; Engel, n. sp., male holotype: A, head, facial view; B, head, dorsal view; C, antenna; D, pronotum, frontal view; E, mesosoma, dorsal view; F, metanotum and propodeum, posterior view; G, habitus, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13954727" httpUri="https://zenodo.org/record/13954727/files/figure.png" pageId="19" pageNumber="714">Fig. 13G</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>