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<document id="F82C05E823F81583055B3900E5D2ABCB" ID-DOI="10.3897/zookeys.1208.116083" ID-ZooBank="0215503D-E8B4-4179-89F6-FB2CCC59F37E" ID-publisher-id="116083" URI-arpha="CF5F342A-04A4-57D6-860A-D48A80ACC24A" URI-zoobank="http://zoobank.org/0215503D-E8B4-4179-89F6-FB2CCC59F37E" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1722361871078" checkinUser="pensoft" docAuthor="Man, Nem Sian, Ablett, Jonathan D., Lwin, Ngwe, Sutcharit, Chirasak &amp; Panha, Somsak" docDate="2024" docId="0C07C9B93D915570BDBC44A60DC4B9AA" docLanguage="en" docName="ZooKeys 1208: 173-239" docOrigin="ZooKeys 1208" docSource="https://zookeys.pensoft.net/article/116083/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Glessula von Martens 1860" docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="239" masterDocId="CF5F342A04A457D6860AD48A80ACC24A" masterDocTitle="Contributions on a small collection of the former Subulinidae Fischer &amp; Crosse, 1877 (Eupulmonata, Achatinoidea) with catalogue of the Glessula and Rishetia species recorded from Myanmar" masterLastPageNumber="239" masterPageNumber="173" pageNumber="173" updateTime="1722370176574" updateUser="admin">
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<mods:title id="F56200590729A003466FE51636E9049E">Contributions on a small collection of the former Subulinidae Fischer &amp; Crosse, 1877 (Eupulmonata, Achatinoidea) with catalogue of the Glessula and Rishetia species recorded from Myanmar</mods:title>
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<mods:namePart id="BD76869CE2BBF886E07B52019270B0B2">Man, Nem Sian</mods:namePart>
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<mods:affiliation id="9568457D3FBC7580CE359B9326C70C70">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand &amp; Department of Zoology, University of Yangon, University Avenue Road, Kamayut Township 11041, Yangon, Myanmar</mods:affiliation>
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<mods:namePart id="EFA001877D311952B014111BD2B9AC71">Ablett, Jonathan D.</mods:namePart>
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<mods:affiliation id="3C21AC8AF0058E9313B30670A10B46B7">Department of Life Sciences, The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
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<mods:affiliation id="3647F94533F1658159CE617D192A77D7">Fauna and Flora International, No. 35, 3 rd Floor, Shan Gone Condo, Myay Ni Gone Market Street, Sanchaung Township, Yangon, Myanmar</mods:affiliation>
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<mods:affiliation id="E7C4FA87239B5FDC4611CC6E0E532CFF">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand</mods:affiliation>
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<mods:affiliation id="27335E2361EDE73AF589CC8E2E678E3D">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand &amp; Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
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<taxonomicName id="8F79950B0BB9163C48B42DFD85334801" authority="von Martens, 1860" authorityName="von Martens" authorityYear="1860" class="Gastropoda" family="Subulinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="A5A8963BE0EFABFEC180C1AB87B4F64D" italics="true">Glessula</emphasis>
<bibRefCitation id="01CE199A1703623EE9070935E88834F1" author="von Martens E" journalOrPublisher="Proceedings of the Zoological Society of London" pagination="6-18" refId="B114" refString="von Martens E (1860) On the Mollusca of Siam. Proceedings of the Zoological Society of London 28: 6 - 18. https: // www. biodiversitylibrary. org / page / 12866504" title="On the Mollusca of Siam." url="https://www.biodiversitylibrary.org/page/12866504" volume="28" year="1860">von Martens, 1860</bibRefCitation>
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<taxonomicName id="F03B8EDBB57772F4047581D0362145E1" authority="Albers, 1850: 194" authorityName="Albers" authorityPageNumber="194" authorityYear="1850" class="Gastropoda" family="Subulinidae" genus="Electra" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="033AF1E044DF12289E89C2D574149581" italics="true">Electra</emphasis>
<bibRefCitation id="99DCA04CC86AFBD3ED7B6B8C49B5E49C" DOI="10.5962/bhl.title.11507" author="Albers JC" journalOrPublisher="Enslin, Berlin" refId="B2" refString="Albers JC (1850) Die Heliceen nach naturlicher Verwandtschaft systematisch geordnet. Enslin, Berlin, 262 pp. https: // doi. org / 10.5962 / bhl. title. 11507" title="Die Heliceen nach natürlicher Verwandtschaft systematisch geordnet." year="1850">Albers, 1850: 194</bibRefCitation>
</taxonomicName>
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. [non
<bibRefCitation id="9EC1793CB04255521361AA6202A32365" author="Lamouroux" firstAuthor="Lamouroux" journalOrPublisher="Poisson, Caen" refId="B56" refString="Lamouroux JVF (1816) Histoire des Polypiers Coralligènes Flexibles, Vulgairement Nommés Zoophytes. F. Poisson, Caen, 558 pp. https://www.biodiversitylibrary.org/page/11606092" title="Histoire des Polypiers Coralligènes Flexibles, Vulgairement Nommés Zoophytes. F." url="https://www.biodiversitylibrary.org/page/11606092" year="1816">Lamouroux 1816</bibRefCitation>
(Bryozoa)]. Type species:
<taxonomicName id="220EF53FF16133B8CB3E5676EC07903D" ID-CoL="969Y" authority="Pfeiffer, 1845" authorityName="Pfeiffer" authorityYear="1845" class="Gastropoda" family="Achatinidae" genus="Achatina" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="ceylanica">
<emphasis id="A09D62B44833075789E9D847BE3D7677" italics="true">Achatina ceylanica</emphasis>
<bibRefCitation id="5946B9A063A9050C6A668EE517813151" author="Pfeiffer L" journalOrPublisher="Zeitschrift für Malakozoologie" pagination="152-158" refId="B73" refString="Pfeiffer L (1845) Diagnosen einiger neuer Heliceen. Zeitschrift fur Malakozoologie 2: 152 - 158. https: // biodiversitylibrary. org / page / 16291740" title="Diagnosen einiger neuer Heliceen." url="https://biodiversitylibrary.org/page/16291740" volume="2" year="1845">Pfeiffer, 1845</bibRefCitation>
</taxonomicName>
.
<bibRefCitation id="93624B02FFCB0C8B9C87A4FFA503E9A6" author="Adams H &amp; Adams A" firstAuthor="Adams" journalOrPublisher="van Voorst, London" page="105" refId="B1" refString="Adams H, Adams A (1855) [1854–1858] The Genera of Recent Mollusca; Arranged According to their Organization. Volume 2. van Voorst, London, 661 pp. [pp 93–284 (1855)]" title="[1854-1858] The Genera of Recent Mollusca; Arranged According to their Organization. Volume 2." year="1855">Adams and Adams 1855: 105</bibRefCitation>
.
<bibRefCitation id="DA28F43AEA5AAC4F92DDAD5421632A96" author="Pfeiffer L" firstAuthor="Pfeiffer" journalOrPublisher="Malakozoologische Blätter" page="168" pagination="112-185" refId="B77" refString="Pfeiffer L (1856) Versuch einer Anordnung der Heliceen nach natürlichen Gruppen. Malakozoologische Blätter 2: 112–185. https://www.biodiversitylibrary.org/item/53876" title="Versuch einer Anordnung der Heliceen nach natürlichen Gruppen." url="https://www.biodiversitylibrary.org/item/53876" volume="2" year="1856">Pfeiffer 1856: 168</bibRefCitation>
.
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<taxonomicName id="5FEC9F6E9F70CED00657421C7354E3B4" authorityName="E. von Martens" authorityYear="1860" class="Gastropoda" family="Achatinidae" genus="Cionella" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="subGenus" subGenus="Glessula">
<emphasis id="3A7ADB00340CB554CD188E04697BEF13" italics="true">Cionella</emphasis>
(
<emphasis id="BDC1E6D019604E6CA7DCA740B29D00D9" italics="true">Glessula</emphasis>
)
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<comment id="95A54E8C1BEDC00037FEDEFA34905CF2">
— von Martens in
<treatmentCitation id="A1565F35370F1F55A0FE80D69F531AD9" author="Albers JC" page="254" year="1860">
<bibRefCitation id="AD37717A9AB9A25F5893F8AB74AE561D" DOI="10.5962/bhl.title.11218" author="Albers JC" firstAuthor="Albers" journalOrPublisher="Wilhelm Englemann, Leipzig" page="254" refId="B3" refString="Albers JC (1860) Die Heliceen nach Natürlicher Verwandtschaft Systematisch Geordnet, 2 nd edn. (manuscript edited by E. von Martens). Wilhelm Englemann, Leipzig, 359 pp. https://doi.org/10.5962/bhl.title.11218" title="Die Heliceen nach Natürlicher Verwandtschaft Systematisch Geordnet, 2 nd edn. (manuscript edited by E. von Martens)." year="1860">Albers 1860: 254</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="7326B0F6497502B0667F8F09DA31BC99" author="Pfeiffer L &amp; Clessin S" page="329" year="1881">
<bibRefCitation id="1BCA8F3FE52F5F956E553D157608DBAB" author="Pfeiffer L &amp; Clessin S" firstAuthor="Pfeiffer" journalOrPublisher="Kassel, Fischer" page="329" refId="B82" refString="Pfeiffer L, Clessin S (1881) Nomenclator Heliceorum Viventium: quo continetur nomina omnium hujus familiae generum et specierum hodie cognitarum, disposita ex affinitate naturali. Kassel, Fischer, 617 pp. https://www.biodiversitylibrary.org/item/47266" title="Nomenclator Heliceorum Viventium: quo continetur nomina omnium hujus familiae generum et specierum hodie cognitarum, disposita ex affinitate naturali." url="https://www.biodiversitylibrary.org/item/47266" year="1881">Pfeiffer and Clessin 1881: 329</bibRefCitation>
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.
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<taxonomicName id="40A8AF71B8D86CD3C861B128DFC4B54C" authorityName="E. von Martens" authorityYear="1860" class="Gastropoda" family="Achatinidae" genus="Stenogyra" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="subGenus" subGenus="Glessula">
<emphasis id="7BEAAE0095F8893B77BD2D46B13BB80C" italics="true">Stenogyra</emphasis>
(
<emphasis id="D860C141E4293977FE5049D7E48D0E5C" italics="true">Glessula</emphasis>
)
</taxonomicName>
<comment id="2079CF7FCBACC3FDADB18422CB923624">
—
<treatmentCitation id="6B12D93F56402C05E0B644C1F964802F" author="Nevill G" page="166" year="1878">
<bibRefCitation id="9CF5D98BD1BB6533742D672D7D6A14B8" DOI="10.5962/bhl.title.11957" author="Nevill G" firstAuthor="Nevill" journalOrPublisher="Indian Museum, Calcutta" page="166" refId="B69" refString="Nevill G (1878) Hand list of Mollusca in the Indian Museum, Calcutta. Part I. Gastropoda. Indian Museum, Calcutta, 338 pp. https://doi.org/10.5962/bhl.title.11957" title="Hand list of Mollusca in the Indian Museum, Calcutta. Part I. Gastropoda." year="1878">Nevill 1878: 166</bibRefCitation>
</treatmentCitation>
.
</comment>
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</paragraph>
<paragraph id="255C518E3DA9C75D9DAFA87BA44B2BDF">
<treatmentCitationGroup id="F3047F6F11F0FB55907FB132EA283A99">
<taxonomicName id="48254FE09F150362D3510CFC468C9946" ID-CoL="7P4CF" authorityName="von Martens" authorityYear="1860" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="41B20E4E7BBB63EB5D6861C05AD2C928" italics="true">Glessula</emphasis>
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<comment id="743A9476D2617C6FE5DD7051B69CE697">
—
<treatmentCitation id="1BD49C0DB9CA98E52B0E4D5543F6CDFB" author="Tenison-Woods JE" page="1053" year="1888">
<bibRefCitation id="4DA512AD043C03029245472FC726B49F" DOI="10.5962/bhl.title.13026" author="Tenison-Woods JE" firstAuthor="Tenison-Woods" journalOrPublisher="Proceedings of the Linnean Society of New South Wales" page="1053" pagination="1003-1095" refId="B104" refString="Tenison-Woods JE (1888) Malaysian land and freshwater Mollusca. Proceedings of the Linnean Society of New South Wales 3: 1003–1095. https://doi.org/10.5962/bhl.title.13026" title="Malaysian land and freshwater Mollusca." volume="3" year="1888">Tenison-Woods 1888: 1053</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="AF4EEE776AD7A7C93E6D1C964B648DBF" author="Beddome CR" page="160" year="1906">
<bibRefCitation id="1372047B91A816861D34866623E72B1F" DOI="10.1093/oxfordjournals.mollus.a066160" author="Beddome CR" firstAuthor="Beddome" page="160" refId="B8" refString="Beddome CR (1906) Notes on Indian and Ceylonese species of Glessula. The Journal of Molluscan Studies 6 (3): 160–172. https://doi.org/10.1093/oxfordjournals.mollus.a066160" year="1906">Beddome 1906: 160</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="E1C60FAF2C3A463327B4CB25584A0E56" author="Pilsbry HA" page="50" year="1908">
<bibRefCitation id="7AB48FFD64BC00E4DECFC74A05A7368C" author="Pilsbry HA" firstAuthor="Pilsbry" page="50" refId="B87" refString="Pilsbry HA (1908) [1908–1910] Manual of Conchology, Structural and Systematic, with Illustrations of the Species, Series 2, Pulmonata. Volume 20: Caecilioides, Glessula and Partulidae. Conchological Section, Academy of Natural Sciences of Philadelphia, Philadelphia, 1–336. [pls 1–43] [pp 1–64, pls 1–10 (1908); pp 65–314, pls 22–36 (1909); pp 315–336, pls 37–43 (1910)] https://www.biodiversitylibrary.org/item/16293" url="https://www.biodiversitylibrary.org/item/16293" year="1908">Pilsbry 1908: 50</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="2E10BBCA5B8F7C65402026A6336DD443" author="Gude GK" page="377" year="1914">
<bibRefCitation id="1F01471E859E5C992B3D4C3682E8399B" DOI="10.5962/bhl.title.12891" author="Gude GK" firstAuthor="Gude" journalOrPublisher="Taylor &amp; Francis, London" page="377" refId="B42" refString="Gude GK (1914) Mollusca - II. Trochomorphidae - Janellidae. In: Shipley AE, Marshall GAK (Eds) The Fauna of British India, including Ceylon and Burma. Taylor &amp; Francis, London, 520 pp. https://doi.org/10.5962/bhl.title.12891" title="Mollusca - II. Trochomorphidae - Janellidae. In: Shipley AE, Marshall GAK (Eds) The Fauna of British India, including Ceylon and Burma." year="1914">Gude 1914: 377</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="1DDADA77549DE968913810E770F7EB81" author="Zilch A" page="343" year="1959">
<bibRefCitation id="97B9D8ED17618D4DB8D20219F82012CB" author="Zilch A" editor="Schindewolf OH" firstAuthor="Zilch" journalOrPublisher="Gebrüder Borntraeger, Berlin" page="343" pagination="201-400" refId="B124" refString="Zilch A (1959) Gastropoda, Euthyneura. In: Schindewolf OH (Ed.) Handbuch der Paläozoologie, Band 6, Gastropoda. Gebrüder Borntraeger, Berlin, 201–400." title="Gastropoda, Euthyneura." volumeTitle="Handbuch der Paläozoologie, Band 6, Gastropoda." year="1959">Zilch 1959: 343</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="58B0A75A67F9BAC3CE3ECE36A684D6EC" author="van Benthem Jutting WSS" page="373" year="1952">
<bibRefCitation id="0E555F4C19562FAA04791897F484F4AC" author="van Benthem Jutting WSS" firstAuthor="van Benthem Jutting" journalOrPublisher="Treubia" page="373" pagination="291-435" refId="B110" refString="van Benthem Jutting WSS (1952) Systematic studies on the non-marine Mollusca of the Ind-Australian archipelago. Treubia 21: 291–435." title="Systematic studies on the non-marine Mollusca of the Ind-Australian archipelago." volume="21" year="1952">van Benthem Jutting 1952: 373</bibRefCitation>
</treatmentCitation>
, 374.
<treatmentCitation id="E85B3C7B17B8D18DD5D61A0A9952C7D9" author="Schileyko AA" page="11" year="2011">
<bibRefCitation id="7F7E307F08D8DA02700A0DB6CBEA029F" author="Schileyko AA" firstAuthor="Schileyko" issue="1" journalOrPublisher="Ruthenica" page="11" pagination="1-68" refId="B96" refString="Schileyko AA (2011) Checklist of land pulmonate molluscs of Vietnam (Gastropoda: Stylommatophora). Ruthenica 21 (1): 1–68." title="Checklist of land pulmonate molluscs of Vietnam (Gastropoda: Stylommatophora)." volume="21" year="2011">Schileyko 2011: 11</bibRefCitation>
</treatmentCitation>
.
</comment>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EFFBI" type="type species">
<paragraph id="96BFB40E2A60726A2BAD1A6EB2176FCD">
<heading id="A964B48A5DA6B85215ECDD5363403C4E" reason="title">Type species.</heading>
</paragraph>
<paragraph id="D74E6808668768E29DA999D534F5F648">
<taxonomicName id="137A01C257D14DE3E2577B3F6F0C7146" authority="Reeve, 1850" authorityName="Reeve" authorityYear="1850" class="Gastropoda" family="Achatinidae" genus="Achatina" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="gemma">
<emphasis id="197440FFC067AB2C538FBFD232EA82A8" italics="true">Achatina gemma</emphasis>
Reeve, 1850
</taxonomicName>
by original designation.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWFBI" type="diagnosis">
<paragraph id="1CEBC18D0F59388295E4AE8FE9E5D7BB">
<heading id="77428BA232B350504174B7D7C34FA4E0" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="4421794B655F41591327F2646DB8D0EC">Shell ovate-conical in shape; spire low or high conical, and regularly attenuated; embryonic whorls smooth or with striations; subsequent whorls with equally or irregularly spaced radial striations or ribs. Aperture oblique, narrow to wide, and ovate; columella concave and truncated, and columellar margin not expanded. Penis large, thick, and moderately long; epiphallus short, stout, terminally pointed, and curved; flagellum present (rarely absent) with comb-like or hand-like structures; epiphallic caecum absent; vagina muscularly and nearly equal to ~ 1 / 5–2 / 5 of penis length.</paragraph>
</subSubSection>
<subSubSection id="SECID0E2FBI" type="remarks">
<paragraph id="F1EED80C58447BC9EB60DF7935D80F26">
<heading id="3A92F9C2C062677CA5AA829D3A7227CD" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="F5C6FF06F79E08A599D1AA3437651DC7">
Superficially,
<taxonomicName id="C51FF18366F41999EC65D6ACAC399DDE" authorityName="von Martens" authorityYear="1860" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="32C37CBAD5F8168BA99860F8CC04C19E" italics="true">Glessula</emphasis>
</taxonomicName>
and
<taxonomicName id="B752C2BF0FE72CAEFAB583C30507E515" authorityName="Godwin-Austen" authorityYear="1920" class="Gastropoda" family="Achatinidae" genus="Rishetia" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="A3ED46689F8BF5F1F046577A9AC00A8C" italics="true">Rishetia</emphasis>
</taxonomicName>
are similar in having a concave and truncated columella, but they differ in their shell shape and genital structure (
<bibRefCitation id="0D948D9003789339961C84BED469D532" author="Godwin-Austen" firstAuthor="Godwin-Austen" refId="B39" refString="Godwin-Austen HH (1920) Land and freshwater Mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other islands of the Indian Ocean. Supplementary to Messrs. Theobald and Hanley’s Conchologia Indica. London, Taylor &amp; Francis, Volume 3, 1–65. [pls 159–165] https://www.biodiversitylibrary.org/item/57585" url="https://www.biodiversitylibrary.org/item/57585" year="1920">Godwin-Austen 1920</bibRefCitation>
;
<bibRefCitation id="A2C7DF49C7BA4B1AFE6FC0AD49548E79" author="Raheem" etAl="et al." firstAuthor="Raheem" refId="B90" refString="Raheem DC, Taylor H, Ablett J, Preece RC, Aravind NA, Naggs F (2014) A systematic revision of the land snails of the Western Ghats of India. Tropical Natural History (Supplement 4): 1–294." year="2014">Raheem et al. 2014</bibRefCitation>
;
<bibRefCitation id="5DE0B56DC87C8EFE7F762D223BDCFAA7" DOI="10.3897/zookeys.675.13252" author="Budha" etAl="et al." firstAuthor="Budha" journalOrPublisher="ZooKeys" pagination="129-156" refId="B24" refString="Budha PB, Naggs F, Backeljau T (2017) Conchological differentiation and genital anatomy of Nepalese Glessulinae (Gastropoda, Stylommatophora, Subulinidae), with descriptions of six new species. ZooKeys 675: 129–156. https://doi.org/10.3897/zookeys.675.13252" title="Conchological differentiation and genital anatomy of Nepalese Glessulinae (Gastropoda, Stylommatophora, Subulinidae), with descriptions of six new species." volume="675" year="2017">Budha et al. 2017</bibRefCitation>
). In terms of shell morphology,
<taxonomicName id="1E949DACEB8E12EC6339181500DC2AFB" authorityName="von Martens" authorityYear="1860" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="D188AD1DC0C889A6E820F6A67AC0D0D6" italics="true">Glessula</emphasis>
</taxonomicName>
can be differentiated by its ovate-conical shape, short spire, large and blunt embryonic whorls, while
<taxonomicName id="7B154EC6AFE9DACB3DB7973605348312" authorityName="Godwin-Austen" authorityYear="1920" class="Gastropoda" family="Achatinidae" genus="Rishetia" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="7E729C7DEFB6D08B58557A8323B303F0" italics="true">Rishetia</emphasis>
</taxonomicName>
possesses a slender, conical-shaped elongate spire which narrow and rather acute embryonic whorls (Table
<tableCitation id="4821BB1A12A68F86C6D1CCA8D1825ABB" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Comparison of shell morphology among subulinid genera recorded in Myanmar. The superscript numbers are references; 1 = Pilsbry (1906), 2 = Gude (1914), 3 = Godwin-Austen (1920), 4 = Naggs (1994), 5 = Schileyko (1999), and 6 = Budha et al. (2017)." httpUri="http://table.plazi.org/id/7320BC8BE0D4AF3B31AE4617F5EE0127" tableUuid="7320BC8BE0D4AF3B31AE4617F5EE0127">2</tableCitation>
). Regarding the genital characters,
<taxonomicName id="746F8A6622E5C6F8DE53DFCF371E044E" authorityName="Godwin-Austen" authorityYear="1920" class="Gastropoda" family="Achatinidae" genus="Rishetia" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="3B02957B35E88A7D37E143B63E3E467E" italics="true">Rishetia</emphasis>
</taxonomicName>
exhibits a simple, tubular-shaped flagellum with the epiphallic caecum present (Table
<tableCitation id="AC9062A8EF8CF63A54174DB3CA36EC6C" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Comparison of terminal part of male genitalia between Glessula and Rishetia. References: 1 = Semper (1874); 2 = Godwin-Austen (1918); 3 = Godwin-Austen (1920); 4 = Schileyko and Kuznetsov (1996); 5 = Schileyko (1999); 6 = Budha et al. (2017). n / a = data not available (not mentioned in the description or could not differentiated from the original figure)." httpUri="http://table.plazi.org/id/6D80CD8EB26F35653BCD7153072A09C9" tableUuid="6D80CD8EB26F35653BCD7153072A09C9">3</tableCitation>
), while
<taxonomicName id="4D4DA9CE59035C1A5C33982C615677B4" authorityName="von Martens" authorityYear="1860" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="2770451D92B3F4F14AD29A28FD6A2DA7" italics="true">Glessula</emphasis>
</taxonomicName>
displays a well-developed comb-like flagellum (appendage) and the epiphallic caecum absent (
<bibRefCitation id="CE430CAC453AA280B55373FB39B52414" DOI="10.3897/zookeys.675.13252" author="Budha" etAl="et al." firstAuthor="Budha" journalOrPublisher="ZooKeys" pagination="129-156" refId="B24" refString="Budha PB, Naggs F, Backeljau T (2017) Conchological differentiation and genital anatomy of Nepalese Glessulinae (Gastropoda, Stylommatophora, Subulinidae), with descriptions of six new species. ZooKeys 675: 129–156. https://doi.org/10.3897/zookeys.675.13252" title="Conchological differentiation and genital anatomy of Nepalese Glessulinae (Gastropoda, Stylommatophora, Subulinidae), with descriptions of six new species." volume="675" year="2017">Budha et al. 2017</bibRefCitation>
); except
<taxonomicName id="F12FBF62ECC6B899FB05C33E5182D1E6" authorityName="Man &amp; Ablett &amp; Lwin &amp; Sutcharit &amp; Panha" authorityYear="2024" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="mandalayensis" status="sp. nov.">
<emphasis id="FAFF342480CA8E60E9DEFF7122BFD447" italics="true">G. mandalayensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F76DEBF54C94202DE6F5B119BE763A4D" rank="species">sp. nov.</taxonomicNameLabel>
which has no flagellum. Likewise,
<taxonomicName id="350F9C43ADAC5A180B473E659A19380C" authorityName="von Martens" authorityYear="1860" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="504AA41DD5CB210578B191B638F91FBD" italics="true">Glessula</emphasis>
</taxonomicName>
is also distinct from
<taxonomicName id="DBF8447E858C9128688314D856245AAE" authorityName="Chaper" authorityYear="1885" class="Gastropoda" family="Achatinidae" genus="Curvella" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="8CB1370B4B3C99E0F1F76A9D3481BCCC" italics="true">Curvella</emphasis>
</taxonomicName>
by its concave and truncated columella, not expanded columellar margin, and obliquely narrow and ovate aperture, while
<taxonomicName id="D11BB42B2DA8AD5EFE097B34CC243F27" authorityName="Chaper" authorityYear="1885" class="Gastropoda" family="Achatinidae" genus="Curvella" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="91EAAB0DBE8AA182DABCFA3982901E1D" italics="true">Curvella</emphasis>
</taxonomicName>
possesses a straight columella, expanded columellar margin, and more vertical, higher, and broader oblong aperture.
</paragraph>
<caption id="6D80CD8EB26F35653BCD7153072A09C9" ID-Table-UUID="6D80CD8EB26F35653BCD7153072A09C9" httpUri="http://table.plazi.org/id/6D80CD8EB26F35653BCD7153072A09C9" startId="T3">
<paragraph id="273051DB0C3286A75EB9847366338813">
<label id="24A3CD1F1045FD75836B3495C842EBA6">Table 3.</label>
</paragraph>
<paragraph id="4B1F1B84849E602B756256C9D4E655F1">
Comparison of terminal part of male genitalia between
<taxonomicName id="A7FC095C76B9C2BDD43C2B89511AACF5" authorityName="von Martens" authorityYear="1860" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="FC7868CD8DBF6A9C69038109312F6DF6" italics="true">Glessula</emphasis>
</taxonomicName>
and
<taxonomicName id="52FB11BDAE0B40608F76924754DD1630" authorityName="Godwin-Austen" authorityYear="1920" class="Gastropoda" family="Achatinidae" genus="Rishetia" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="563543E717179AAF99B80D0432142348" italics="true">Rishetia</emphasis>
</taxonomicName>
. References: 1 =
<bibRefCitation id="84E150282118005860FF7B8CA7E8FCA9" author="Semper C" firstAuthor="Semper" journalOrPublisher="Band 3, Landmollusken. Wiesbaden: Kreidel" refId="B98" refString="Semper C (1874) [1870–1885] Reisen im Archipel der Philippinen, Theil 2. Wissenschaftliche Resultate. Band 3, Landmollusken. Wiesbaden: Kreidel, 327 pp. [pp 129–168 (1874)]" title="[1870-1885] Reisen im Archipel der Philippinen, Theil 2. Wissenschaftliche Resultate." year="1874">Semper (1874)</bibRefCitation>
; 2 =
<bibRefCitation id="45DC372725684825A9EF93708748CEED" DOI="10.26515/rzsi/v8/i12/1918/163139" author="Godwin-Austen HH" firstAuthor="Godwin-Austen" issue="12" journalOrPublisher="Records of the Indian Museum" pagination="601-621" refId="B38" refString="Godwin-Austen HH (1918) Zoological results of the Abor Expedition, 1911–1912. XLIX. Mollusca, IX. Records of the Indian Museum 8 (12): 601–621. https://doi.org/10.26515/rzsi/v8/i12/1918/163139" title="Zoological results of the Abor Expedition, 1911-1912. XLIX. Mollusca, IX." volume="8" year="1918">Godwin-Austen (1918)</bibRefCitation>
; 3 =
<bibRefCitation id="49338DDA64B51E729FFDB7FEC1B7E2A2" author="Godwin-Austen HH" firstAuthor="Godwin-Austen" refId="B39" refString="Godwin-Austen HH (1920) Land and freshwater Mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other islands of the Indian Ocean. Supplementary to Messrs. Theobald and Hanley’s Conchologia Indica. London, Taylor &amp; Francis, Volume 3, 1–65. [pls 159–165] https://www.biodiversitylibrary.org/item/57585" url="https://www.biodiversitylibrary.org/item/57585" year="1920">Godwin-Austen (1920)</bibRefCitation>
; 4 =
<bibRefCitation id="7EC0FFA384A9C2B36F46E97639B9EB3E" author="Schileyko AA &amp; Kuznetsov AG" firstAuthor="Schileyko" issue="2" journalOrPublisher="Ruthenica" pagination="158-160" refId="B97" refString="Schileyko AA, Kuznetsov AG (1996) A new genus of the Subulinidae (Pulmonata) from Nepal. Ruthenica 5 (2): 158–160." title="A new genus of the Subulinidae (Pulmonata) from Nepal." volume="5" year="1996">Schileyko and Kuznetsov (1996)</bibRefCitation>
; 5 =
<bibRefCitation id="25B1521C0CF7F299E0F20D65FD02D3B7" author="Schileyko AA" firstAuthor="Schileyko" refId="B95" refString="Schileyko AA (1999) Treatise on recent terrestrial pulmonate molluscs Part 4. Draparnaudiidae, Caryodidae, Macrocyclidae, Acavidae, Clavatoridae, Dorcasiidae, Sculptariidae, Corillidae, Plectopylidae, Megalobulimidae, Strophocheilidae, Cerionidae, Achatinidae, Subulinidae, Glessulidae, Micractaeonidae, Ferrussaciidae. Ruthenica (Supplement 2): 437–564." year="1999">Schileyko (1999)</bibRefCitation>
; 6 =
<bibRefCitation id="92B896460BD6C447CB9D6711B492E7CF" DOI="10.3897/zookeys.675.13252" author="Budha PB &amp; Naggs F &amp; Backeljau T" etAl="et al." firstAuthor="Budha" journalOrPublisher="ZooKeys" pagination="129-156" refId="B24" refString="Budha PB, Naggs F, Backeljau T (2017) Conchological differentiation and genital anatomy of Nepalese Glessulinae (Gastropoda, Stylommatophora, Subulinidae), with descriptions of six new species. ZooKeys 675: 129–156. https://doi.org/10.3897/zookeys.675.13252" title="Conchological differentiation and genital anatomy of Nepalese Glessulinae (Gastropoda, Stylommatophora, Subulinidae), with descriptions of six new species." volume="675" year="2017">Budha et al. (2017)</bibRefCitation>
. n / a = data not available (not mentioned in the description or could not differentiated from the original figure).
</paragraph>
</caption>
<paragraph id="88E990AF0AA3F7A6162DBF3E06CA92E8">
<table id="TID0E5WDK" rules="all">
<tbody id="773D99E476B32664A5BFBB94784CCD5C">
<tr id="5D9258524A2F80F112582A93DDA7E9C4">
<th id="7CD04F5632A076A92D4F3238D945A70E" colspan="1" rowspan="1">Species</th>
<th id="4CE6D65553EB7B1185EC15894076037E" colspan="1" rowspan="1">Epiphallic caecum</th>
<th id="A230FD83D4960415C694035114172072" colspan="1" rowspan="1">Flagellum</th>
<th id="BF621D966A03B7074895056C3ECFA13B" colspan="1" rowspan="1">References</th>
</tr>
<tr id="34AA3438075C0AF26B5E57651C74E8EC">
<td id="F015643E893CFD480DF752DC6A84F572" colspan="4" rowspan="1">
<taxonomicName id="D9C780175CCB71684782EAFC5D6054DC" authority="von Martens, 1860" authorityName="von Martens" authorityYear="1860" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="9B6DB0DA5CE73CFD9368D15188D7855B" bold="true">
<emphasis id="FB430A16DC054AED3000243082606ACD" bold="true" italics="true">Glessula</emphasis>
<bibRefCitation id="23D00CBE6BB09C05BF4E064CC921E3F7" author="von Martens E" journalOrPublisher="Proceedings of the Zoological Society of London" pagination="6-18" refId="B114" refString="von Martens E (1860) On the Mollusca of Siam. Proceedings of the Zoological Society of London 28: 6 - 18. https: // www. biodiversitylibrary. org / page / 12866504" title="On the Mollusca of Siam." url="https://www.biodiversitylibrary.org/page/12866504" volume="28" year="1860">von Martens, 1860</bibRefCitation>
</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="C0EB25DFDB73EA38243DFFF48524629D">
<td id="CF2E5A9189C7B032ABA5CE806652F5E2" colspan="1" rowspan="1">
<taxonomicName id="77A8E56EE666AB5B642692BFC6625A40" authority="Godwin-Austen, 1920" authorityName="Godwin-Austen" authorityYear="1920" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="hebetata">
<emphasis id="D2AB830254AA2F7B004EA29CB96360A1" italics="true">G. hebetata</emphasis>
<bibRefCitation id="657C9F6252AAC7BAC82251DD23D75400" author="Godwin-Austen HH" refId="B39" refString="Godwin-Austen HH (1920) Land and freshwater Mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other islands of the Indian Ocean. Supplementary to Messrs. Theobald and Hanley's Conchologia Indica. London, Taylor &amp; Francis, Volume 3, 1 - 65. [pls 159 - 165] https: // www. biodiversitylibrary. org / item / 57585" url="https://www.biodiversitylibrary.org/item/57585" year="1920">Godwin-Austen, 1920</bibRefCitation>
</taxonomicName>
</td>
<td id="1F42A32CCBCA755B9D15274E9C9040D4" colspan="1" rowspan="1">absent</td>
<td id="C2F792E920B1F072BA4323A0F01BCB9D" colspan="1" rowspan="1">comb-like with numerous notches</td>
<td id="B450DA0C73413D4D54E80C416E2355CE" colspan="1" rowspan="1">6</td>
</tr>
<tr id="2AD79E0E4C2A24C01FB560C2E9876397">
<td id="4F90F137CC0AE989B640CB1792B94AEA" colspan="1" rowspan="1">
<taxonomicName id="80DAADC45AA87835B418BAA43BEBFB94" authority="(Pfeiffer, 1853)" baseAuthorityName="Pfeiffer" baseAuthorityYear="1853" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="inornata">
<emphasis id="85614E1E40A319D12300FFCD3FD8452A" italics="true">G. inornata</emphasis>
(
<bibRefCitation id="DD8E3FCA01DD4B34C26F5C84EA7A4822" author="Pfeiffer L" refId="B76" refString="Pfeiffer L (1853) Monographia Heliceorum Viventium: sistens descriptiones systematicas et criticas omnium huius familiae generum et specierum hodie cognitarum. Volumen Tertium. Lipsiae: F. A. Brockhaus, 711 pp. https: // www. biodiversitylibrary. org / item / 47349" url="https://www.biodiversitylibrary.org/item/47349" year="1853">Pfeiffer, 1853</bibRefCitation>
)
</taxonomicName>
</td>
<td id="171B940AE265ECC6FAA32671A528A7F8" colspan="1" rowspan="1">absent</td>
<td id="35243952D59C442F6BB1EE618B36CBEB" colspan="1" rowspan="1">comb-like with numerous notches</td>
<td id="58BF8B691721A330A9EE8910794C8A81" colspan="1" rowspan="1">3</td>
</tr>
<tr id="F12E88C8B7B3D973CAB559EDC9BEE785">
<td id="FA0F16B0CA09E48EDD53DFB6DCBA02C4" colspan="1" rowspan="1">
<taxonomicName id="0A8495C4BB2D730DBF7E57DABFA12886" authority="Godwin-Austen, 1918" authorityName="Godwin-Austen" authorityYear="1918" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="oakesi">
<emphasis id="F41BEDDD620D3C87D0675C43862470ED" italics="true">G. oakesi</emphasis>
<bibRefCitation id="C3800E278C68A56F3B5C173267954D47" DOI="10.26515/rzsi/v8/i12/1918/163139" author="Godwin-Austen HH" issue="12" journalOrPublisher="Records of the Indian Museum" pagination="601-621" refId="B38" refString="Godwin-Austen HH (1918) Zoological results of the Abor Expedition, 1911 - 1912. XLIX. Mollusca, IX. Records of the Indian Museum 8 (12): 601 - 621. https: // doi. org / 10.26515 / rzsi / v 8 / i 12 / 1918 / 163139" title="Zoological results of the Abor Expedition, 1911-1912. XLIX. Mollusca, IX." volume="8" year="1918">Godwin-Austen, 1918</bibRefCitation>
</taxonomicName>
</td>
<td id="BCA97F30AFB9BD3BED70A9FC003E147C" colspan="1" rowspan="1">absent</td>
<td id="61F84B4313ED48C1281BAEAAE3CBC5F4" colspan="1" rowspan="1">comb-like with three notches</td>
<td id="AABE2B5E1F1C1847C26DD3F210321B95" colspan="1" rowspan="1">2, 3</td>
</tr>
<tr id="2DC8A1A8CF51F65057846D093A2205C3">
<td id="E32BDA6F131AAC70142F90E2BFB0FB50" colspan="1" rowspan="1">
<taxonomicName id="2F067003D75A5F89782F34B987404417" authority="Godwin-Austen, 1918" authorityName="Godwin-Austen" authorityYear="1918" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="ochracea">
<emphasis id="78E6ECE6D18814BBF72D19AECBAEE6BA" italics="true">G. ochracea</emphasis>
<bibRefCitation id="4574F06DE31AE674CB7637968E4E8147" DOI="10.26515/rzsi/v8/i12/1918/163139" author="Godwin-Austen HH" issue="12" journalOrPublisher="Records of the Indian Museum" pagination="601-621" refId="B38" refString="Godwin-Austen HH (1918) Zoological results of the Abor Expedition, 1911 - 1912. XLIX. Mollusca, IX. Records of the Indian Museum 8 (12): 601 - 621. https: // doi. org / 10.26515 / rzsi / v 8 / i 12 / 1918 / 163139" title="Zoological results of the Abor Expedition, 1911-1912. XLIX. Mollusca, IX." volume="8" year="1918">Godwin-Austen, 1918</bibRefCitation>
</taxonomicName>
</td>
<td id="114C0DB9C99E3B04B21F680E4F2DA8DB" colspan="1" rowspan="1">absent</td>
<td id="B0BD1EF8F3D9027719FE776F1963119A" colspan="1" rowspan="1">comb-like with numerous notches</td>
<td id="4D17BC66F73614787DF8335DCDE8E251" colspan="1" rowspan="1">2, 3</td>
</tr>
<tr id="2AAB49877948999DE5FA1E72051462A1">
<td id="081640E054446A13669642AA8AFE21B3" colspan="1" rowspan="1">
<taxonomicName id="A3EC74F5A21C13DBA3A930E69F619DA7" authority="(Benson, 1860)" baseAuthorityName="Benson" baseAuthorityYear="1860" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="orobia">
<emphasis id="F6AF0E6E58714485BA059B89A4F76924" italics="true">G. orobia</emphasis>
(
<bibRefCitation id="79E8F55B8497827547CA1BF01B3BE19E" DOI="10.1080/00222936008697255" author="Benson WH" refId="B12" refString="Benson WH (1860) On new land-shells from Darjiling, with a series of new Indian species of Achatina. Annals &amp; Magazine of Natural History Series 3 5 (30): 460 - 466. https: // doi. org / 10.1080 / 00222936008697255" year="1860">Benson, 1860</bibRefCitation>
)
</taxonomicName>
</td>
<td id="AF80F62F38E45A4AB986A83BB41B292F" colspan="1" rowspan="1">absent</td>
<td id="598E6543ABFDFA0E74850A4E94FD9641" colspan="1" rowspan="1">comb-like with five notches</td>
<td id="6025B4BCB2F10253B7397DC565D824C1" colspan="1" rowspan="1">3, 6</td>
</tr>
<tr id="858DCA9C5C2EE3870F185276D42D030C">
<td id="E547EFB13E8822D492051F211EB51886" colspan="1" rowspan="1">
<taxonomicName id="E677C5AD1C42575D4BB76EF50231D71D" authority="(Reeve, 1849)" baseAuthorityName="Reeve" baseAuthorityYear="1849" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="orophila">
<emphasis id="291A919B58EC57910A9581BA0958A368" italics="true">G. orophila</emphasis>
(Reeve, 1849)
</taxonomicName>
</td>
<td id="A497067F37070BE15C8D6E87930FB316" colspan="1" rowspan="1">absent</td>
<td id="B700E4B1BADED66CB9B4617858E41C10" colspan="1" rowspan="1">comb-like with numerous notches</td>
<td id="CE53E388CED1CCA6937CA12128FB1ACC" colspan="1" rowspan="1">1</td>
</tr>
<tr id="6AE2A1241AF722A97AB88469AB786B90">
<td id="FF71BD17FCE0E398AB68FD22A724842D" colspan="1" rowspan="1">
<taxonomicName id="CD9AB73408E36D87D69114017701E3B8" authority="(Benson, 1860)" baseAuthorityName="Benson" baseAuthorityYear="1860" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="serena">
<emphasis id="6EE3A0DB0D3D916DFF311E74C07ABF45" italics="true">G. serena</emphasis>
(
<bibRefCitation id="CFC7BBC3DFD19992460BD470F3929873" DOI="10.1080/00222936008697255" author="Benson WH" refId="B12" refString="Benson WH (1860) On new land-shells from Darjiling, with a series of new Indian species of Achatina. Annals &amp; Magazine of Natural History Series 3 5 (30): 460 - 466. https: // doi. org / 10.1080 / 00222936008697255" year="1860">Benson, 1860</bibRefCitation>
)
</taxonomicName>
</td>
<td id="C55C4CF1A9068C1A30A66E95735F7282" colspan="1" rowspan="1">absent</td>
<td id="F2B524696F022100A8488C8E57B9C3AA" colspan="1" rowspan="1">comb-like with numerous notches</td>
<td id="6065F95B18E29DE2D1B330E4CF79620C" colspan="1" rowspan="1">5</td>
</tr>
<tr id="8FA512CE10C78E914A494E0AF31FB165">
<td id="DF38B34025CEC7234E7C9F5726C38C46" colspan="1" rowspan="1">
<taxonomicName id="EA017039C562FC97C1048C864EA0D1CE" authority="Budha &amp; Backeljau, 2017" authorityName="Budha &amp; Backeljau" authorityYear="2017" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="tamakoshi">
<emphasis id="DE630B8E9D33F04E9E6F1385A7A1E9BF" italics="true">G. tamakoshi</emphasis>
<bibRefCitation id="A605E37CA2D4E0232BA9EF8421ABE19D" DOI="10.3897/zookeys.675.13252" author="Budha PB &amp; Naggs F &amp; Backeljau T" journalOrPublisher="ZooKeys" pagination="129-156" refId="B24" refString="Budha PB, Naggs F, Backeljau T (2017) Conchological differentiation and genital anatomy of Nepalese Glessulinae (Gastropoda, Stylommatophora, Subulinidae), with descriptions of six new species. ZooKeys 675: 129 - 156. https: // doi. org / 10.3897 / zookeys. 675.13252" title="Conchological differentiation and genital anatomy of Nepalese Glessulinae (Gastropoda, Stylommatophora, Subulinidae), with descriptions of six new species." volume="675" year="2017">Budha &amp; Backeljau, 2017</bibRefCitation>
</taxonomicName>
</td>
<td id="ABD641486B9737F3E1496C9B4882C5AD" colspan="1" rowspan="1">absent</td>
<td id="50FF0576FF37DE1B3D7C0BB69F5D5EC1" colspan="1" rowspan="1">comb-like with numerous notches</td>
<td id="A23DE2E0394B28C4975A10839934F236" colspan="1" rowspan="1">6</td>
</tr>
<tr id="AD6AD1A109DFAEB478F728A7C075BF5A">
<td id="8212A2780AA435DA9952EAA5409B18A0" colspan="1" rowspan="1">
<taxonomicName id="9C4834D768385116439EA9AC81554C8C" authorityName="Man &amp; Ablett &amp; Lwin &amp; Sutcharit &amp; Panha" authorityYear="2024" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="mandalayensis" status="sp. nov.">
<emphasis id="1F2ACD3988C5593BB6806211D42EEA4A" italics="true">G. mandalayensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3AF2EA0AE0315ECEFED7E7408D8F108B" rank="species">sp. nov.</taxonomicNameLabel>
</td>
<td id="BB922DC05EED7647D1F2C3CED1A29D38" colspan="1" rowspan="1">absent</td>
<td id="195D1281ABB75CFA3AF08AD781506C41" colspan="1" rowspan="1">absent</td>
<td id="FE4BDAD6769669699B43201D4C720C10" colspan="1" rowspan="1">This study</td>
</tr>
<tr id="E44E0A7B3C333C9DA37B30C1CD073261">
<td id="CAF67B4BDD764B924C2B4B8813F8F093" colspan="4" rowspan="1">
<taxonomicName id="8A408F4EFAE2FF6FB7337A4CE93E3ECF" authority="Godwin-Austen, 1920" authorityName="Godwin-Austen" authorityYear="1920" class="Gastropoda" family="Achatinidae" genus="Rishetia" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="D142A59162B655D6EF56FB45E02FE7D3" bold="true">
<emphasis id="C24CE6E76DA5424F3FAD891347C24503" bold="true" italics="true">Rishetia</emphasis>
<bibRefCitation id="D5325EAF8250C9B8C53C68BB614D7F18" author="Godwin-Austen HH" refId="B39" refString="Godwin-Austen HH (1920) Land and freshwater Mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other islands of the Indian Ocean. Supplementary to Messrs. Theobald and Hanley's Conchologia Indica. London, Taylor &amp; Francis, Volume 3, 1 - 65. [pls 159 - 165] https: // www. biodiversitylibrary. org / item / 57585" url="https://www.biodiversitylibrary.org/item/57585" year="1920">Godwin-Austen, 1920</bibRefCitation>
</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="2283B93CBFC33FC99618253C524D4539">
<td id="CD105ADCABBFA6287B7C1BA7613D9CA4" colspan="1" rowspan="1">
<taxonomicName id="7557ED2049C5FEA413449231663987A8" authority="(Godwin-Austen, 1920)" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1920" class="Gastropoda" family="Achatinidae" genus="Rishetia" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="garoense">
<emphasis id="D206087DCA982FE4448F6728E84299F7" italics="true">R. garoense</emphasis>
(
<bibRefCitation id="8B463B3E58C45F47A48C18C41C93E625" author="Godwin-Austen HH" refId="B39" refString="Godwin-Austen HH (1920) Land and freshwater Mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other islands of the Indian Ocean. Supplementary to Messrs. Theobald and Hanley's Conchologia Indica. London, Taylor &amp; Francis, Volume 3, 1 - 65. [pls 159 - 165] https: // www. biodiversitylibrary. org / item / 57585" url="https://www.biodiversitylibrary.org/item/57585" year="1920">Godwin-Austen, 1920</bibRefCitation>
)
</taxonomicName>
</td>
<td id="D5AF6DC3CB73F0BFDF80196939688ED1" colspan="1" rowspan="1">n / a</td>
<td id="3E36B316FD6CDDADF6716471A31E2974" colspan="1" rowspan="1">long cylindrical</td>
<td id="28674AD66EE1797CFA0D147F75EE4FEA" colspan="1" rowspan="1">3</td>
</tr>
<tr id="8DFE8B19E58B02B61EDD28BD47E85D5F">
<td id="47EB681C6B533DD485A92CFF1DE1F43A" colspan="1" rowspan="1">
<taxonomicName id="57693C2C51D8422F5F472DD865BC15B3" authority="(Benson, 1860)" baseAuthorityName="Benson" baseAuthorityYear="1860" class="Gastropoda" family="Achatinidae" genus="Rishetia" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="hastula">
<emphasis id="FEF06EF278AF63F4D8A5CE2512206827" italics="true">R. hastula</emphasis>
(
<bibRefCitation id="2097401B1D7C7675EB8401E79A71194B" DOI="10.1080/00222936008697255" author="Benson WH" refId="B12" refString="Benson WH (1860) On new land-shells from Darjiling, with a series of new Indian species of Achatina. Annals &amp; Magazine of Natural History Series 3 5 (30): 460 - 466. https: // doi. org / 10.1080 / 00222936008697255" year="1860">Benson, 1860</bibRefCitation>
)
</taxonomicName>
</td>
<td id="530A348B479BAFB5E95C96AC77E8274D" colspan="1" rowspan="1">short knob-like</td>
<td id="A6145D6D4984DBA279D6A4C86DF849C4" colspan="1" rowspan="1">short tubular</td>
<td id="D13FE3781FB8822969081A222A371C93" colspan="1" rowspan="1">6</td>
</tr>
<tr id="73B8D061B4DA27C2861F4D8D15B9FCC1">
<td id="80514208D3C6266F3FD38D9B6A2A895C" colspan="1" rowspan="1">
<taxonomicName id="55C3CC192CC27EFD6E4731FB1E82E025" authority="Budha &amp; Backeljau, 2017" authorityName="Budha &amp; Backeljau" authorityYear="2017" class="Gastropoda" family="Achatinidae" genus="Rishetia" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="kathmandica">
<emphasis id="D9AE9DEB8B2F146AAEB0231D17094C9C" italics="true">R. kathmandica</emphasis>
<bibRefCitation id="CC45331F0E2E72E5AB205F3D35619DD1" DOI="10.3897/zookeys.675.13252" author="Budha PB &amp; Naggs F &amp; Backeljau T" journalOrPublisher="ZooKeys" pagination="129-156" refId="B24" refString="Budha PB, Naggs F, Backeljau T (2017) Conchological differentiation and genital anatomy of Nepalese Glessulinae (Gastropoda, Stylommatophora, Subulinidae), with descriptions of six new species. ZooKeys 675: 129 - 156. https: // doi. org / 10.3897 / zookeys. 675.13252" title="Conchological differentiation and genital anatomy of Nepalese Glessulinae (Gastropoda, Stylommatophora, Subulinidae), with descriptions of six new species." volume="675" year="2017">Budha &amp; Backeljau, 2017</bibRefCitation>
</taxonomicName>
</td>
<td id="7349A88B79AAA1E0B953779AA015C67D" colspan="1" rowspan="1">short cylindrical</td>
<td id="7F4C53BE521FCC66B8017AD476C8467D" colspan="1" rowspan="1">long cylindrical</td>
<td id="94A9BA4BDA53F0C320C0866FB04A7DF4" colspan="1" rowspan="1">6</td>
</tr>
<tr id="B8874F1987C1E357B1D1DF5CA421FE00">
<td id="748553E20C9100E0024DC9E4EC12C3AA" colspan="1" rowspan="1">
<taxonomicName id="0F0E453C28676166335002EC38529ACA" authority="(Godwin-Austen, 1920)" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1920" class="Gastropoda" family="Achatinidae" genus="Rishetia" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="longispira">
<emphasis id="08655074F2F683BEAB20407509014C4D" italics="true">R. longispira</emphasis>
(
<bibRefCitation id="AA82CEBBF2F3A49F9DFBC28B471E497E" author="Godwin-Austen HH" refId="B39" refString="Godwin-Austen HH (1920) Land and freshwater Mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other islands of the Indian Ocean. Supplementary to Messrs. Theobald and Hanley's Conchologia Indica. London, Taylor &amp; Francis, Volume 3, 1 - 65. [pls 159 - 165] https: // www. biodiversitylibrary. org / item / 57585" url="https://www.biodiversitylibrary.org/item/57585" year="1920">Godwin-Austen, 1920</bibRefCitation>
)
</taxonomicName>
</td>
<td id="4598C41BA639B866B5E23F8B8FD7A2CE" colspan="1" rowspan="1">n / a</td>
<td id="12B44B91BF1BA976CEC3DB8CE21E700F" colspan="1" rowspan="1">long cylindrical</td>
<td id="21FDF08001F0F7DE0A7054210E54B175" colspan="1" rowspan="1">3, 6</td>
</tr>
<tr id="959E4EACD7FD68DAC88F380069204FE2">
<td id="41C3D348EA7A5FDCC4A14C88169BC869" colspan="1" rowspan="1">
<taxonomicName id="22561ABC03487B532BEB6EFB0DAC27CF" authority="(Godwin-Austen, 1920)" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1920" class="Gastropoda" family="Achatinidae" genus="Rishetia" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="mastersi">
<emphasis id="9874C74598B90CFFFBC6203A63924B32" italics="true">R. mastersi</emphasis>
(
<bibRefCitation id="13790C54926A8FF16DFB3737ED4258C2" author="Godwin-Austen HH" refId="B39" refString="Godwin-Austen HH (1920) Land and freshwater Mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other islands of the Indian Ocean. Supplementary to Messrs. Theobald and Hanley's Conchologia Indica. London, Taylor &amp; Francis, Volume 3, 1 - 65. [pls 159 - 165] https: // www. biodiversitylibrary. org / item / 57585" url="https://www.biodiversitylibrary.org/item/57585" year="1920">Godwin-Austen, 1920</bibRefCitation>
)
</taxonomicName>
</td>
<td id="0DA250A3351C79D83D8D97C9F38AB18F" colspan="1" rowspan="1">short cylindrical</td>
<td id="F5CF758B00F98A8DD5C3FCDB10B1F712" colspan="1" rowspan="1">long cylindrical</td>
<td id="AFB0598975B5E2210403D4EBAD84FB8C" colspan="1" rowspan="1">6</td>
</tr>
<tr id="2CAEEB9CE54AA794558570C4640F57DA">
<td id="3769737BA3DF3B4348D6ACDAD4A24157" colspan="1" rowspan="1">
<taxonomicName id="342EAD93FF80D31AA9ACFE1932DE6534" authority="Budha &amp; Naggs, 2017" authorityName="Budha &amp; Naggs" authorityYear="2017" class="Gastropoda" family="Achatinidae" genus="Rishetia" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="nagarjunensis">
<emphasis id="CDDD3379488903F82CC944FF1D934CEA" italics="true">R. nagarjunensis</emphasis>
<bibRefCitation id="65F669E6AE99E96F4E09CD427571AEC8" DOI="10.3897/zookeys.675.13252" author="Budha PB &amp; Naggs F &amp; Backeljau T" journalOrPublisher="ZooKeys" pagination="129-156" refId="B24" refString="Budha PB, Naggs F, Backeljau T (2017) Conchological differentiation and genital anatomy of Nepalese Glessulinae (Gastropoda, Stylommatophora, Subulinidae), with descriptions of six new species. ZooKeys 675: 129 - 156. https: // doi. org / 10.3897 / zookeys. 675.13252" title="Conchological differentiation and genital anatomy of Nepalese Glessulinae (Gastropoda, Stylommatophora, Subulinidae), with descriptions of six new species." volume="675" year="2017">Budha &amp; Naggs, 2017</bibRefCitation>
</taxonomicName>
</td>
<td id="D2C39EF015ECC457272B295B4ADD113E" colspan="1" rowspan="1">short knob</td>
<td id="D4F676667F8DB5B598A0371E274A283B" colspan="1" rowspan="1">short knob</td>
<td id="2C7DF34B74FC7198CC962BF087F0D0CC" colspan="1" rowspan="1">6</td>
</tr>
<tr id="68EF97B734D80F5E7B07532A0A1EB12F">
<td id="4256014601DC0DC17024D60570D50DB6" colspan="1" rowspan="1">
<taxonomicName id="02004F9DEC5E8F4A57A6B6D5179791A5" authority="Budha &amp; Naggs, 2017" authorityName="Budha &amp; Naggs" authorityYear="2017" class="Gastropoda" family="Achatinidae" genus="Rishetia" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="rishikeshi">
<emphasis id="99FA9F05B791A4DB4B4C07E02327BA36" italics="true">R. rishikeshi</emphasis>
<bibRefCitation id="AD136230738DC0F8E11600A13A72C3E6" DOI="10.3897/zookeys.675.13252" author="Budha PB &amp; Naggs F &amp; Backeljau T" journalOrPublisher="ZooKeys" pagination="129-156" refId="B24" refString="Budha PB, Naggs F, Backeljau T (2017) Conchological differentiation and genital anatomy of Nepalese Glessulinae (Gastropoda, Stylommatophora, Subulinidae), with descriptions of six new species. ZooKeys 675: 129 - 156. https: // doi. org / 10.3897 / zookeys. 675.13252" title="Conchological differentiation and genital anatomy of Nepalese Glessulinae (Gastropoda, Stylommatophora, Subulinidae), with descriptions of six new species." volume="675" year="2017">Budha &amp; Naggs, 2017</bibRefCitation>
</taxonomicName>
</td>
<td id="145BEE895522C9D7393EA4C70A34D374" colspan="1" rowspan="1">short cylindrical</td>
<td id="859E71CB83D40BB632E585C597C9D6DC" colspan="1" rowspan="1">long cylindrical</td>
<td id="4804A0021BE490CAE7173D8B637D21A7" colspan="1" rowspan="1">6</td>
</tr>
<tr id="3EE0F3C3B31FBA1120920F6EA827185D">
<td id="750104A1F1AA92CA604C9D3000647591" colspan="1" rowspan="1">
<taxonomicName id="16DB0A135E0F8811BF60696D945C2EB5" authority="Budha &amp; Naggs, 2017" authorityName="Budha &amp; Naggs" authorityYear="2017" class="Gastropoda" family="Achatinidae" genus="Rishetia" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="subulata">
<emphasis id="D126FA134EC4D52CDE4F153884DD22C7" italics="true">R. subulata</emphasis>
<bibRefCitation id="86A0ECC6CCE8F36F7F19561F48838D76" DOI="10.3897/zookeys.675.13252" author="Budha PB &amp; Naggs F &amp; Backeljau T" journalOrPublisher="ZooKeys" pagination="129-156" refId="B24" refString="Budha PB, Naggs F, Backeljau T (2017) Conchological differentiation and genital anatomy of Nepalese Glessulinae (Gastropoda, Stylommatophora, Subulinidae), with descriptions of six new species. ZooKeys 675: 129 - 156. https: // doi. org / 10.3897 / zookeys. 675.13252" title="Conchological differentiation and genital anatomy of Nepalese Glessulinae (Gastropoda, Stylommatophora, Subulinidae), with descriptions of six new species." volume="675" year="2017">Budha &amp; Naggs, 2017</bibRefCitation>
</taxonomicName>
</td>
<td id="1469E1053C2572E83A55F6600D87BA3B" colspan="1" rowspan="1">short cylindrical</td>
<td id="12377ED1B6699C49246C4FF4D8255464" colspan="1" rowspan="1">long cylindrical</td>
<td id="85C8C6F1765DBDADE28FFA9AB3C89651" colspan="1" rowspan="1">6</td>
</tr>
<tr id="51C123441B719A9D87E7FEA64043BEF4">
<td id="8779C56251A2DD53DEDB0F7DEC46D57C" colspan="1" rowspan="1">
<taxonomicName id="072504615935069CF72C1B01F8CB8891" authority="Budha, 2017" authorityName="Budha" authorityYear="2017" class="Gastropoda" family="Achatinidae" genus="Rishetia" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="tribhuvana">
<emphasis id="B719C712CC2F255A8AA4877DEABC23B2" italics="true">R. tribhuvana</emphasis>
Budha, 2017
</taxonomicName>
</td>
<td id="4C5F797805D9A41BBD9A88C73D75A85F" colspan="1" rowspan="1">short cylindrical</td>
<td id="F22448BA1DE2B5D1E3CE7E381A146F18" colspan="1" rowspan="1">long cylindrical</td>
<td id="0B59FFC10CE245229BDD9CDF292C47A2" colspan="1" rowspan="1">6</td>
</tr>
<tr id="09052399C270C4B1C85C887E98B8BCB8">
<td id="0F7FCF17968EB75463536DD23027557F" colspan="1" rowspan="1">
<taxonomicName id="315EC042D34CE982A9FF43F8B5A05F7C" authority="(Benson, 1936)" baseAuthorityName="Benson" baseAuthorityYear="1936" class="Gastropoda" family="Achatinidae" genus="Achatina" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="tenuispira">
<emphasis id="0BFFDD2A8E6263DB5A3C61E163C21623" italics="true">R. tenuispira</emphasis>
(Benson, 1936)
</taxonomicName>
</td>
<td id="35A63649A894EFC16552996F7B84D317" colspan="1" rowspan="1">short cylindrical</td>
<td id="4E7636FD34556058B00385B13588F34B" colspan="1" rowspan="1">long cylindrical</td>
<td id="F1E3F514BE00F424A9127C8023630FDB" colspan="1" rowspan="1">4, 5</td>
</tr>
</tbody>
</table>
</paragraph>
<paragraph id="99422C6C205A672A00D257741C007647">
<taxonomicName id="76C835728ACFEACF8932E65C6CE1EB5B" authorityName="von Martens" authorityYear="1860" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="36B95ADAD57608082C5A5147FE006EC2" italics="true">Glessula</emphasis>
</taxonomicName>
is widely distributed throughout the Indian subcontinent, and nearly 100 species have been described (
<bibRefCitation id="C79AB8F4DF2A44D769118512B68499A1" author="Godwin-Austen" firstAuthor="Godwin-Austen" refId="B39" refString="Godwin-Austen HH (1920) Land and freshwater Mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other islands of the Indian Ocean. Supplementary to Messrs. Theobald and Hanley’s Conchologia Indica. London, Taylor &amp; Francis, Volume 3, 1–65. [pls 159–165] https://www.biodiversitylibrary.org/item/57585" url="https://www.biodiversitylibrary.org/item/57585" year="1920">Godwin-Austen 1920</bibRefCitation>
;
<bibRefCitation id="39BCB33A3D2579CE97CE196DAEAC64A4" author="Schileyko" firstAuthor="Schileyko" refId="B95" refString="Schileyko AA (1999) Treatise on recent terrestrial pulmonate molluscs Part 4. Draparnaudiidae, Caryodidae, Macrocyclidae, Acavidae, Clavatoridae, Dorcasiidae, Sculptariidae, Corillidae, Plectopylidae, Megalobulimidae, Strophocheilidae, Cerionidae, Achatinidae, Subulinidae, Glessulidae, Micractaeonidae, Ferrussaciidae. Ruthenica (Supplement 2): 437–564." year="1999">Schileyko 1999</bibRefCitation>
;
<bibRefCitation id="CDBD7649711F9FFAB5BCE78AD78111FC" author="Ramakrishna" etAl="et al." firstAuthor="Ramakrishna" refId="B91" refString="Ramakrishna, Mitra SC, Dey A (2010) Annotated checklist of Indian land molluscs. Occasional paper no. 306. Zoological Survey of India, 359 pp." year="2010">Ramakrishna et al. 2010</bibRefCitation>
;
<bibRefCitation id="F02F7DE81389CFF5DA7E7FF69CD364FF" DOI="10.14284/448" author="MolluscaBase" firstAuthor="MolluscaBase" refId="B61" refString="MolluscaBase eds (2023) MolluscaBase. https://www.molluscabase.org [in 2023–07 – 17] https://doi.org/10.14284/448" url="https://www.molluscabase.org" year="2023">MolluscaBase 2023</bibRefCitation>
). Among the sporadic species recorded in Southeast Asia,
<collectingCountry id="D043817482311D8621C1EB0A01F409AD" name="Myanmar">Myanmar</collectingCountry>
has 12 species (
<bibRefCitation id="638CFC25E5A4486E304D7D3490B03404" DOI="10.5962/bhl.title.12891" author="Gude" firstAuthor="Gude" journalOrPublisher="Taylor &amp; Francis, London" refId="B42" refString="Gude GK (1914) Mollusca - II. Trochomorphidae - Janellidae. In: Shipley AE, Marshall GAK (Eds) The Fauna of British India, including Ceylon and Burma. Taylor &amp; Francis, London, 520 pp. https://doi.org/10.5962/bhl.title.12891" title="Mollusca - II. Trochomorphidae - Janellidae. In: Shipley AE, Marshall GAK (Eds) The Fauna of British India, including Ceylon and Burma." year="1914">Gude 1914</bibRefCitation>
;
<bibRefCitation id="A7EE946DF7372CFA9C5F1D8425E48E8D" author="Godwin-Austen" firstAuthor="Godwin-Austen" refId="B39" refString="Godwin-Austen HH (1920) Land and freshwater Mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other islands of the Indian Ocean. Supplementary to Messrs. Theobald and Hanley’s Conchologia Indica. London, Taylor &amp; Francis, Volume 3, 1–65. [pls 159–165] https://www.biodiversitylibrary.org/item/57585" url="https://www.biodiversitylibrary.org/item/57585" year="1920">Godwin-Austen 1920</bibRefCitation>
), while two species,
<taxonomicName id="D4B572AA4352F97F4A4082E5EAE1C350" authority="von Mollendorff, 1899" authorityName="von Mollendorff" authorityYear="1899" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="latestriata">
<emphasis id="158F54163AE139E98DFAE2D464AD4408" italics="true">G. latestriata</emphasis>
<bibRefCitation id="EFD2588E20312AE5C56329BA8AB3E3AC" author="von Möllendorff OF" refId="B119" refString="von Mollendorff OF (1899) Neue arten aus Hinterindien. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 31: 165, 166. https: // biodiversitylibrary. org / page / 28228787" url="https://biodiversitylibrary.org/page/28228787" year="1899">von Möllendorff, 1899</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="E86F81A258631BA52C056CCB9595952B" authority="Morlet 1893" authorityName="Morlet" authorityYear="1893" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="paviei">
<emphasis id="4CB6AAC00AD3EC1667B8DB266169B700" italics="true">G. paviei</emphasis>
<bibRefCitation id="9637A87EB32F945DEAA19605F8D72D33" author="Morlet" firstAuthor="Morlet" issue="1892" journalOrPublisher="Journal de Conchyliologie" pagination="315-329" refId="B63" refString="Morlet L (1893) Descriptions ďespèces nouvelles provenant de ľIndochine. Journal de Conchyliologie 40 [1892]: 315–329." title="Descriptions ďespèces nouvelles provenant de ľIndochine." volume="40" year="1893">Morlet 1893</bibRefCitation>
</taxonomicName>
, have been recorded from
<collectingCountry id="E4B2A7DC8E445A8581AC4DF2223ECEB4" name="Vietnam">Vietnam</collectingCountry>
,
<collectingCountry id="1445CC35F000FC821B312E1480A5D760" name="Laos">Laos</collectingCountry>
, and
<collectingCountry id="6F9F5615CC9ADDC3A2970B3EF7156E2E" name="Thailand">Thailand</collectingCountry>
, and another two species,
<taxonomicName id="3F6403F10C42336FBCA9956970D1734B" authority="(von Martens, 1864)" baseAuthorityName="von Martens" baseAuthorityYear="1864" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="sumatrana">
<emphasis id="7D4F52C4825C44CFB5F531995D1BCA72" italics="true">G. sumatrana</emphasis>
(
<bibRefCitation id="C5F1C88E7A568B6C4DBB53759D21C4C8" author="von Martens E" journalOrPublisher="Monatsberichte der Königlichen Preussischen Akademien der Wissenschaften zu Berlin" pagination="523-529" refId="B115" refString="von Martens E (1864) Diagnosen neuer Heliceen aus dem ostasiatischen Archipel. Monatsberichte der Koniglichen Preussischen Akademien der Wissenschaften zu Berlin 1864: 523 - 529. https: // www. biodiversitylibrary. org / page / 35180469 # page / 541 /" title="Diagnosen neuer Heliceen aus dem ostasiatischen Archipel." url="https://www.biodiversitylibrary.org/page/35180469#page/541/" volume="1864" year="1864">von Martens, 1864</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="103C5771CF5EB813D63B3410A508FBB3" authority="(Pfeiffer, 1856)" baseAuthorityName="Pfeiffer" baseAuthorityYear="1856" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="wallacei">
<emphasis id="E461835DCCCAF426DA7A2CFFE3FBF252" italics="true">G. wallacei</emphasis>
(
<bibRefCitation id="63EAF18A6BFB4F4EBF1DB84CAFD6162F" author="Pfeiffer L" journalOrPublisher="Malakozoologische Blätter" pagination="112-185" refId="B77" refString="Pfeiffer L (1856) Versuch einer Anordnung der Heliceen nach naturlichen Gruppen. Malakozoologische Blatter 2: 112 - 185. https: // www. biodiversitylibrary. org / item / 53876" title="Versuch einer Anordnung der Heliceen nach natürlichen Gruppen." url="https://www.biodiversitylibrary.org/item/53876" volume="2" year="1856">Pfeiffer, 1856</bibRefCitation>
)
</taxonomicName>
from
<collectingCountry id="13967BE3B861BAAB9A9D98791C8DBD91" name="Indonesia">Indonesia</collectingCountry>
(
<bibRefCitation id="147390F0607F9AB814EC4DFFBAAF02DF" DOI="10.5962/bhl.title.13026" author="Tenison-Woods" firstAuthor="Tenison-Woods" journalOrPublisher="Proceedings of the Linnean Society of New South Wales" pagination="1003-1095" refId="B104" refString="Tenison-Woods JE (1888) Malaysian land and freshwater Mollusca. Proceedings of the Linnean Society of New South Wales 3: 1003–1095. https://doi.org/10.5962/bhl.title.13026" title="Malaysian land and freshwater Mollusca." volume="3" year="1888">Tenison-Woods 1888</bibRefCitation>
;
<bibRefCitation id="23D87819A2C8B42965B2D94A9D898513" author="van Benthem Jutting" firstAuthor="van Benthem Jutting" journalOrPublisher="Treubia" pagination="291-435" refId="B110" refString="van Benthem Jutting WSS (1952) Systematic studies on the non-marine Mollusca of the Ind-Australian archipelago. Treubia 21: 291–435." title="Systematic studies on the non-marine Mollusca of the Ind-Australian archipelago." volume="21" year="1952">van Benthem Jutting 1952</bibRefCitation>
;
<bibRefCitation id="26F414ECF3EE388DA2DFA7A095269A0E" author="Solem" firstAuthor="Solem" journalOrPublisher="Spolia Zoologia Musei Hauniensis" pagination="1-110" refId="B99" refString="Solem A (1966) Some non-marine mollusks from Thailand. Spolia Zoologia Musei Hauniensis 24: 1–110." title="Some non-marine mollusks from Thailand." volume="24" year="1966">Solem 1966</bibRefCitation>
;
<bibRefCitation id="62C13564D5977EFE4440ECC5F077B732" author="Schileyko" firstAuthor="Schileyko" issue="1" journalOrPublisher="Ruthenica" pagination="1-68" refId="B96" refString="Schileyko AA (2011) Checklist of land pulmonate molluscs of Vietnam (Gastropoda: Stylommatophora). Ruthenica 21 (1): 1–68." title="Checklist of land pulmonate molluscs of Vietnam (Gastropoda: Stylommatophora)." volume="21" year="2011">Schileyko 2011</bibRefCitation>
;
<bibRefCitation id="8A7069BF1B338D86E740EC4F6B303A64" DOI="10.3897/zookeys.834.28800" author="Inkhavilay" etAl="et al." firstAuthor="Inkhavilay" journalOrPublisher="ZooKeys" pagination="1-166" refId="B47" refString="Inkhavilay K, Sutcharit C, Bantaowong U, Chanabun R, Siriwut W, Srisonchai R, Pholyotha A, Jirapatrasilp P, Panha S (2019) Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda). ZooKeys 834: 1–166. https://doi.org/10.3897/zookeys.834.28800" title="Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda)." volume="834" year="2019">Inkhavilay et al. 2019</bibRefCitation>
).
</paragraph>
<paragraph id="95B42AECBFD2E704226F8463FF4DF3CC">
In this study, only five species including a new species,
<taxonomicName id="3DAEE4BE365D307020DD70DF52F5C620" authority="Nevill, 1877" authorityName="Nevill" authorityYear="1877" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="blanfordiana">
<emphasis id="CA986C68A0B18455E51E21092D406CF5" italics="true">G. blanfordiana</emphasis>
<bibRefCitation id="41E30B97C645B48D933021B92BC2A23A" author="Nevill G" issue="2" journalOrPublisher="Journal of the Asiatic Society of Bengal" pagination="14-41" refId="B68" refString="Nevill G (1877) List of the Mollusca brought back by Dr. J. Anderson from Yunnan and Upper Burma, with descriptions of new species. Journal of the Asiatic Society of Bengal 46 (2): 14 - 41. https: // www. biodiversitylibrary. org / item / 110105" title="List of the Mollusca brought back by Dr. J. Anderson from Yunnan and Upper Burma, with descriptions of new species." url="https://www.biodiversitylibrary.org/item/110105" volume="46" year="1877">Nevill, 1877</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="21F14EDEF3E943C858102BC8478D6451" authority="Godwin-Austen, 1920" authorityName="Godwin-Austen" authorityYear="1920" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="feddeni">
<emphasis id="A242A451B6FFF3A315E9F463D5C4B94E" italics="true">G. feddeni</emphasis>
<bibRefCitation id="7AFB29D501233E9C56D9D64D43028B9C" author="Godwin-Austen HH" refId="B39" refString="Godwin-Austen HH (1920) Land and freshwater Mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other islands of the Indian Ocean. Supplementary to Messrs. Theobald and Hanley's Conchologia Indica. London, Taylor &amp; Francis, Volume 3, 1 - 65. [pls 159 - 165] https: // www. biodiversitylibrary. org / item / 57585" url="https://www.biodiversitylibrary.org/item/57585" year="1920">Godwin-Austen, 1920</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="ACA53D212384AE8A79E7E5B558DE9783" authority="(Reeve, 1850)" baseAuthorityName="Reeve" baseAuthorityYear="1850" class="Gastropoda" family="Achatinidae" genus="Achatina" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="gemma">
<emphasis id="C48DC48F32331E284E16CF23EF7CFE89" italics="true">G. gemma</emphasis>
(Reeve, 1850)
</taxonomicName>
,
<taxonomicName id="79A045069743FB7F12A9077D29FE8316" authorityName="von Mollendorff" authorityYear="1899" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="latestriata">
<emphasis id="8B45509414165F994AF96E790DEE4CFE" italics="true">G. latestriata</emphasis>
</taxonomicName>
, and
<taxonomicName id="219268ED78C41181284A1FF8C3C4F909" authorityName="Man &amp; Ablett &amp; Lwin &amp; Sutcharit &amp; Panha" authorityYear="2024" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="mandalayensis" status="sp. nov.">
<emphasis id="991E73E1E55125FAD422C8342676946E" italics="true">G. mandalayensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FE659E92E63C5F5E4D1D83E02F95365A" rank="species">sp. nov.</taxonomicNameLabel>
have been collected and re-described.
</paragraph>
</subSubSection>
</treatment>
</document>