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<document id="6BD1D623CDFBC177A9F9D292DA0C8000" ID-DOI="10.3897/zookeys.1208.116083" ID-ZooBank="0215503D-E8B4-4179-89F6-FB2CCC59F37E" ID-publisher-id="116083" URI-arpha="CF5F342A-04A4-57D6-860A-D48A80ACC24A" URI-zoobank="http://zoobank.org/0215503D-E8B4-4179-89F6-FB2CCC59F37E" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1722361871078" checkinUser="pensoft" docAuthor="Man, Nem Sian, Ablett, Jonathan D., Lwin, Ngwe, Sutcharit, Chirasak &amp; Panha, Somsak" docDate="2024" docId="04837403AA6A538D92CF1AD16EE01E7F" docLanguage="en" docName="ZooKeys 1208: 173-239" docOrigin="ZooKeys 1208" docSource="https://zookeys.pensoft.net/article/116083/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Rishetia burrailensis subsp. maxwelli" docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="239" masterDocId="CF5F342A04A457D6860AD48A80ACC24A" masterDocTitle="Contributions on a small collection of the former Subulinidae Fischer &amp; Crosse, 1877 (Eupulmonata, Achatinoidea) with catalogue of the Glessula and Rishetia species recorded from Myanmar" masterLastPageNumber="239" masterPageNumber="173" pageNumber="173" updateTime="1722370176574" updateUser="admin">
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<mods:title id="AA0DF8F17A01FF649E7DB73B1F797C6D">Contributions on a small collection of the former Subulinidae Fischer &amp; Crosse, 1877 (Eupulmonata, Achatinoidea) with catalogue of the Glessula and Rishetia species recorded from Myanmar</mods:title>
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<mods:namePart id="0B23C01F83816F407721DBE7612AFC19">Man, Nem Sian</mods:namePart>
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<mods:affiliation id="5429E617DE5E3F7D1F8E9BCFA76048E2">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand &amp; Department of Zoology, University of Yangon, University Avenue Road, Kamayut Township 11041, Yangon, Myanmar</mods:affiliation>
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<mods:namePart id="AF8F3575C8422101A1D412D01CF70331">Ablett, Jonathan D.</mods:namePart>
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<mods:affiliation id="977B46A1EF542388644EB685EF60B9EA">Department of Life Sciences, The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
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<mods:namePart id="B5A95C0042C27A406E116F709477308A">Lwin, Ngwe</mods:namePart>
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<mods:affiliation id="A646BE468DEDA5BCAA94FD63CCFA4E4A">Fauna and Flora International, No. 35, 3 rd Floor, Shan Gone Condo, Myay Ni Gone Market Street, Sanchaung Township, Yangon, Myanmar</mods:affiliation>
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<mods:namePart id="EC925A86E509B65EA5AD8447FB76C499">Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation id="C68F2129E6CFBA78C73CD3385B2426B4">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand</mods:affiliation>
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<mods:namePart id="D1609EC32AF875865FC686CF40D0AD21">Panha, Somsak</mods:namePart>
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<mods:affiliation id="21EF7AB7EA8C6EF16276C0D0BF0C0376">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand &amp; Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
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<mods:date id="348D2EC973AE1C60217619B732D8A9DD">2024</mods:date>
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<treatment id="04837403AA6A538D92CF1AD16EE01E7F" ID-DOI="http://doi.org/10.5281/zenodo.13137160" ID-Zenodo-Dep="13137160" ID-arpha="04837403-AA6A-538D-92CF-1AD16EE01E7F" LSID="urn:lsid:plazi:treatment:04837403AA6A538D92CF1AD16EE01E7F" httpUri="http://treatment.plazi.org/id/04837403AA6A538D92CF1AD16EE01E7F">
<subSubSection id="E74F3AB790EF06E7E0D0FF149FB2A8F8" type="nomenclature">
<paragraph id="30C0A765911DC06B0CF682F76952C01A">
<heading id="3AE76BF6B43FE8B984FAE7CE59048275" reason="label">30</heading>
<taxonomicName id="C1C127B12A9387ACAE03FDC8DC1F6EE7" authority="(Godwin-Austen, 1920)" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1920" class="Gastropoda" family="Subulinidae" genus="Rishetia" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="subSpecies" species="burrailensis" subSpecies="maxwelli">
<emphasis id="E55C90B5473DCB7F1732729473BF6C23" italics="true">Rishetia burrailensis maxwelli</emphasis>
(
<bibRefCitation id="D6BC3B6D1BDABD366BB2F81E05D726EF" author="Godwin-Austen HH" refId="B39" refString="Godwin-Austen HH (1920) Land and freshwater Mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other islands of the Indian Ocean. Supplementary to Messrs. Theobald and Hanley's Conchologia Indica. London, Taylor &amp; Francis, Volume 3, 1 - 65. [pls 159 - 165] https: // www. biodiversitylibrary. org / item / 57585" url="https://www.biodiversitylibrary.org/item/57585" year="1920">Godwin-Austen, 1920</bibRefCitation>
)
</taxonomicName>
comb. nov.
</paragraph>
</subSubSection>
<subSubSection id="2450291558CBF2841BC93013E24520CA" type="description">
<paragraph id="89026183D5B1149E219887476D3D1163">
<figureCitation id="4C66356CF05E4454ECF0EF64C4D42C33" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. A C Rishetia akouktoungensis, syntypes NHMUK 1906.1. 1.2207 from Akouktoung, Pegu D Rishetia baculina, syntypes NHMUK 1909.3. 15.9 from Kursiong, Darjiling E, F Rishetia basseinensis, syntypes NHMUK 1909.3. 15.19 from Bassein G I Rishetia burrailensis maxwelli G holotype NHMUK 1903.7. 1.1717 / 1 from Naga Hills, East of Kohima H specimen NHMUK 1988147 from Somra Tracts, Somra Khulen Post, west of Kyengdwen River, Burma and I paratypes NHMUK 1903.7. 1.1717 / 24 from Naga Hills, East of Kohima." figureDoi="10.3897/zookeys.1208.116083.figure13" httpUri="https://binary.pensoft.net/fig/1104511">Figs 13 G I</figureCitation>
,
<figureCitation id="38F0C0DBC3BE41482D606C6803406B60" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. A C Rishetia burrailensis maxwelli A, B specimen NHMUK 1903.7. 1.3742 from Somra Tracts, Somra Khulen Post, Burma and C specimen NHMUK 1903.7. 1.3744 from Somra Tracts, Somra Khulen Post, Burma D Rishetia hastula, neotype NHMUK 1906.1. 1.880 from Darjiling E, F Rishetia kentungensis, syntypes NHMUK 1903.7. 1.3650 from Mong Sing, Siam Boundary G Rishetia limborgi, holotype NHMUK 1903.7. 1.3200 from Tenasserim H, I Rishetia nathiana, syntypes NHMUK 1906.1. 1.2206 from Bassein District." figureDoi="10.3897/zookeys.1208.116083.figure14" httpUri="https://binary.pensoft.net/fig/1104512">14 A C</figureCitation>
</paragraph>
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<subSubSection id="F68642E0514A6EB82EC04D2A4CBDA5FC" type="reference_group">
<paragraph id="F67D843CD0A473EFC4EFD4F6989EB81D">
<treatmentCitationGroup id="B2969421E3EE38A5737F8B8D485575A4">
<treatmentCitation id="D3138760E1B5E5CAD8B89FA0245E01B9" author="Godwin-Austen HH" page="45" year="1920">
<taxonomicName id="F90123922B434511ADBCEC61FC7B5139" authority="Godwin-Austen, 1920: 45" authorityName="Godwin-Austen" authorityPageNumber="45" authorityYear="1920" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="variety" species="burrailensis" subGenus="Rishetia" variety="maxwelli">
<emphasis id="110AFC23D6983E5B9E818762B39C359B" italics="true">Glessula</emphasis>
(
<emphasis id="672812273BC33908D4F966E421CD1C9A" italics="true">Rishetia</emphasis>
)
<emphasis id="103814CE9F3010E023119D01E44127CE" italics="true">burrailensis</emphasis>
<comment id="5D76996F6AC0475E33CBFAB42B21C05F">
, var.
<emphasis id="BBFAA96AFE6B084400D79CA8B54F74F5" italics="true">maxwelli</emphasis>
<bibRefCitation id="986CE7E8F0967F49AD7CA157EDB642F3" author="Godwin-Austen HH" refId="B39" refString="Godwin-Austen HH (1920) Land and freshwater Mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other islands of the Indian Ocean. Supplementary to Messrs. Theobald and Hanley's Conchologia Indica. London, Taylor &amp; Francis, Volume 3, 1 - 65. [pls 159 - 165] https: // www. biodiversitylibrary. org / item / 57585" url="https://www.biodiversitylibrary.org/item/57585" year="1920">Godwin-Austen, 1920: 45</bibRefCitation>
, 46, pl. 160, figs 5, 6. Type locality: Naga Hills, east of Kohima and Somra, Khulen Post. West of Kyendwin or Chindwin River, Upper
<collectingCountry id="F98F805287BF118E052B92234FC9BB09" name="Myanmar">Burma</collectingCountry>
.
</comment>
</taxonomicName>
</treatmentCitation>
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</paragraph>
<paragraph id="461CD8133A28208D010B03BD7E9E90A9">
<treatmentCitationGroup id="8E7C6A9776EB99ED791CB43007BA8F35">
<taxonomicName id="51FF73E7498273E3E722A1879CD57EE7" authorityName="Godwin-Austen" authorityYear="1920" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="variety" species="burrailensis" variety="maxwelli">
<emphasis id="DBF869A9AE7F1AA4451BEC432C245117" italics="true">Glessula burrailensis</emphasis>
<comment id="2045087CCE6515B67F5226586E2C0F82">
, var.
<emphasis id="E5B98F8582C9006A43D9B09ED7F5E31A" italics="true">maxwelli</emphasis>
<treatmentCitation id="DFFD58F66886B597A7BA6C34D267626F" author="Godwin-Austen HH" page="61" year="1920">
<bibRefCitation id="BC870E5AF1B0935CE6ECA8B9D2DD63F1" author="Godwin-Austen HH" firstAuthor="Godwin-Austen" page="61" refId="B39" refString="Godwin-Austen HH (1920) Land and freshwater Mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other islands of the Indian Ocean. Supplementary to Messrs. Theobald and Hanleys Conchologia Indica. London, Taylor &amp; Francis, Volume 3, 165. [pls 159165] https://www.biodiversitylibrary.org/item/57585" url="https://www.biodiversitylibrary.org/item/57585" year="1920">Godwin-Austen 1920: 61</bibRefCitation>
</treatmentCitation>
. Type locality: Somra Tracts, Somra Khulen Post, Upper
<collectingCountry id="6869DCA8B391E3863A6E601408E10725" name="Myanmar">Burma</collectingCountry>
.
</comment>
</taxonomicName>
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</paragraph>
</subSubSection>
<subSubSection id="SECID0EOGEK" type="Type specimens">
<paragraph id="0AE639C4B71CB19CB0C284EC50E1812C">
<heading id="8FA0E210575BDD13162B4D65A8B8D2EC" reason="title">Type specimens.</heading>
</paragraph>
<paragraph id="7D1C88EF044384ED0DA5EB98E7F1BEF3">
<materialsCitation id="B318366EFB10045330E9D7BA8756819E" collectionCode="NHMUK" collectorName="Godwin-Austen" county="Godwin-Austen" location="Naga Hills" municipality="Kohima" specimenCode="NHMUK 1903.7. 1.1717" specimenCount="1" stateProvince="Naga" typeStatus="holotype">
<emphasis id="E37055FC0AACCBDBFEFC71DD0D546B38" bold="true" italics="true">
<typeStatus id="C2E4C0A54DA5CE1D2DA6F6F7AD20A8FF" type="holotype">Holotype</typeStatus>
</emphasis>
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<named-content id="61474E64571F06715467E05CE57DFB2F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">NHMUK</named-content>
1903.7. 1.1717
</specimenCode>
/ 1 (Fig.
<figureCitation id="14C59386FC209E202C5B79BDCB18227D" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. A C Rishetia akouktoungensis, syntypes NHMUK 1906.1. 1.2207 from Akouktoung, Pegu D Rishetia baculina, syntypes NHMUK 1909.3. 15.9 from Kursiong, Darjiling E, F Rishetia basseinensis, syntypes NHMUK 1909.3. 15.19 from Bassein G I Rishetia burrailensis maxwelli G holotype NHMUK 1903.7. 1.1717 / 1 from Naga Hills, East of Kohima H specimen NHMUK 1988147 from Somra Tracts, Somra Khulen Post, west of Kyengdwen River, Burma and I paratypes NHMUK 1903.7. 1.1717 / 24 from Naga Hills, East of Kohima." figureDoi="10.3897/zookeys.1208.116083.figure13" httpUri="https://binary.pensoft.net/fig/1104511">13 G</figureCitation>
) ex.
<collectorName id="05318875C63214C4D39032277E3F0565">Godwin-Austen</collectorName>
collection from
<location id="8401E73D96C284E85601BBA5D33841D4" LSID="urn:lsid:plazi:treatment:04837403AA6A538D92CF1AD16EE01E7F:8401E73D96C284E85601BBA5D33841D4" county="Godwin-Austen" municipality="Kohima" name="Naga Hills" stateProvince="Naga">Naga Hills</location>
, East of
<collectingMunicipality id="2F013177AA5C3EE800069D5E57093DB1">Kohima</collectingMunicipality>
</materialsCitation>
.
<materialsCitation id="AA82F4BB2A27B83D4CA34AB04AA1715F" collectionCode="NHMUK" collectorName="Godwin-Austen" county="Godwin-Austen" location="Naga Hills" municipality="Kohima" specimenCode="1903.7. 1.1717" specimenCount="1" stateProvince="Naga" typeStatus="paratype">
<emphasis id="4B05709D2E19F095EF8919FB20EABA4E" bold="true" italics="true">
<typeStatus id="012DAE1F51251056B890266CBF204C24" type="paratype">Paratypes</typeStatus>
</emphasis>
<named-content id="0EA3F768E28D9F1D0B48BD2E4833E04C" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
<collectionCode id="6FE5EA9B68A2A99FA1ACB466660D6C42" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
<specimenCode id="4DD6741ECD11CBA3799D8B4A1128AAB6">1903.7. 1.1717</specimenCode>
/ 24 (3 shells; Fig.
<figureCitation id="2FC8A83DE2FB8A93E667EE56C1E2B51D" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. A C Rishetia akouktoungensis, syntypes NHMUK 1906.1. 1.2207 from Akouktoung, Pegu D Rishetia baculina, syntypes NHMUK 1909.3. 15.9 from Kursiong, Darjiling E, F Rishetia basseinensis, syntypes NHMUK 1909.3. 15.19 from Bassein G I Rishetia burrailensis maxwelli G holotype NHMUK 1903.7. 1.1717 / 1 from Naga Hills, East of Kohima H specimen NHMUK 1988147 from Somra Tracts, Somra Khulen Post, west of Kyengdwen River, Burma and I paratypes NHMUK 1903.7. 1.1717 / 24 from Naga Hills, East of Kohima." figureDoi="10.3897/zookeys.1208.116083.figure13" httpUri="https://binary.pensoft.net/fig/1104511">13 I</figureCitation>
) ex. Godwin-Austen collection from the location same as the holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EOHEK" type="material">
<paragraph id="B1E7CF27ECD6D75F8CCE55829E5A7C21">
<heading id="FD562F0EAC712853B28E7158E90C3F23" reason="title">Other material.</heading>
</paragraph>
<paragraph id="563290340032E3BA2EE18CE2BE5E1035">
<named-content id="BCFD670FD8B5079C73FFF8B664923660" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
<collectionCode id="09E74D31EF8E8D6377CEA27C8877EE24" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
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1903.7. 1.3742 (7 shells + 1 broken shell; Fig.
<figureCitation id="5B55957A2346D082D27208979692BA81" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. A C Rishetia burrailensis maxwelli A, B specimen NHMUK 1903.7. 1.3742 from Somra Tracts, Somra Khulen Post, Burma and C specimen NHMUK 1903.7. 1.3744 from Somra Tracts, Somra Khulen Post, Burma D Rishetia hastula, neotype NHMUK 1906.1. 1.880 from Darjiling E, F Rishetia kentungensis, syntypes NHMUK 1903.7. 1.3650 from Mong Sing, Siam Boundary G Rishetia limborgi, holotype NHMUK 1903.7. 1.3200 from Tenasserim H, I Rishetia nathiana, syntypes NHMUK 1906.1. 1.2206 from Bassein District." figureDoi="10.3897/zookeys.1208.116083.figure14" httpUri="https://binary.pensoft.net/fig/1104512">14 A, B</figureCitation>
) collected by F. Ede ex. Godwin-Austen collection from Somra Tracts, Somra Khulen Post,
<collectingCountry id="198E75DE73522C770BE21A67567F474B" name="Myanmar">Burma</collectingCountry>
(labelled as
<taxonomicName id="18054BB616C6788AADA19681D6831CFA" rank="variety" variety="somraensis">
var.
<emphasis id="523C793E2B78221B877AA57327604D92" italics="true">somraensis</emphasis>
</taxonomicName>
).
<named-content id="F8CACAA9394C8D21AAE6A35D073EE562" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
<collectionCode id="04C59A332646F1B17F567270BFC93A2F" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
1903.7. 1.3744 (2 shells; Fig.
<figureCitation id="D81A9A3C6DDA7666AA3FD0C400788D86" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. A C Rishetia burrailensis maxwelli A, B specimen NHMUK 1903.7. 1.3742 from Somra Tracts, Somra Khulen Post, Burma and C specimen NHMUK 1903.7. 1.3744 from Somra Tracts, Somra Khulen Post, Burma D Rishetia hastula, neotype NHMUK 1906.1. 1.880 from Darjiling E, F Rishetia kentungensis, syntypes NHMUK 1903.7. 1.3650 from Mong Sing, Siam Boundary G Rishetia limborgi, holotype NHMUK 1903.7. 1.3200 from Tenasserim H, I Rishetia nathiana, syntypes NHMUK 1906.1. 1.2206 from Bassein District." figureDoi="10.3897/zookeys.1208.116083.figure14" httpUri="https://binary.pensoft.net/fig/1104512">14 C</figureCitation>
) collected by F. Ede ex. Godwin-Austen collection from Somra Tracts, Somra Khulen Post,
<collectingCountry id="9B24E47DADD75AD80D54CA9E775BA531" name="Myanmar">Burma</collectingCountry>
(labelled as
<taxonomicName id="A36AEB830C01A66D9E5C4E646BE80C35" rank="variety" variety="somraensis">
var.
<emphasis id="06EBFED0A2DDFC56608ED7239B23652C" italics="true">somraensis</emphasis>
</taxonomicName>
).
<named-content id="B6353512ECBD107740C8F4DB8F6BFF1F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
<collectionCode id="8552001D5820D1CAB0CA2F4DFEE01DE9" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
1988147 (4 shells; Fig.
<figureCitation id="7EE0FB95DFBADB15139007FD630ECA43" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. A C Rishetia akouktoungensis, syntypes NHMUK 1906.1. 1.2207 from Akouktoung, Pegu D Rishetia baculina, syntypes NHMUK 1909.3. 15.9 from Kursiong, Darjiling E, F Rishetia basseinensis, syntypes NHMUK 1909.3. 15.19 from Bassein G I Rishetia burrailensis maxwelli G holotype NHMUK 1903.7. 1.1717 / 1 from Naga Hills, East of Kohima H specimen NHMUK 1988147 from Somra Tracts, Somra Khulen Post, west of Kyengdwen River, Burma and I paratypes NHMUK 1903.7. 1.1717 / 24 from Naga Hills, East of Kohima." figureDoi="10.3897/zookeys.1208.116083.figure13" httpUri="https://binary.pensoft.net/fig/1104511">13 H</figureCitation>
) collected by L. R. Mawson from Somra Tracts, Somra Khulen Post, west of Kyengdwen River,
<collectingCountry id="F9A07A329EDC1CBCB006173568BF7CDF" name="Myanmar">Burma</collectingCountry>
.
</paragraph>
<caption id="4F306B49B86B6EF06FD04868EB681E17" ID-DOI="10.3897/zookeys.1208.116083.figure14" ID-arpha="BDF5BDC7-85B5-5D09-AA93-EE64C882A3B0" httpUri="https://binary.pensoft.net/fig/1104512" startId="F14">
<paragraph id="BA7486E208BE4B2D1EB5378F0E5560FF">
<label id="0060295E3CEC4DC3641C522185A25BA5">Figure 14.</label>
</paragraph>
<paragraph id="9684500AB26F1F1FF462E9E4B092E9B9">
<emphasis id="4C87E0F3242FCE4535188123F4AEAAC7" bold="true">A C</emphasis>
<taxonomicName id="41FD237C6DC49C848D672C6AC3DB2BD7" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1920" class="Gastropoda" family="Achatinidae" genus="Rishetia" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="subSpecies" species="burrailensis" subSpecies="maxwelli">
<emphasis id="D18B707CC0A966F14D8D8D044D16E030" italics="true">Rishetia burrailensis maxwelli</emphasis>
</taxonomicName>
<emphasis id="684390457346B3F6211CA0A68EB39C9A" bold="true">A, B</emphasis>
specimen
<named-content id="1B0312B10606E5BA67B8EC08EB94F1E3" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">NHMUK</named-content>
1903.7. 1.3742 from Somra Tracts, Somra Khulen Post, Burma and
<emphasis id="FB5077743F4DFC7A944EE61C9305D349" bold="true">C</emphasis>
specimen
<named-content id="D884F895950C8FFD7C618E91ADB55F61" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">NHMUK</named-content>
1903.7. 1.3744 from Somra Tracts, Somra Khulen Post, Burma
<emphasis id="07942914B3B0FD90CE21A79B6838882A" bold="true">D</emphasis>
<taxonomicName id="885924250BFADA342379CC93E9EF9853" baseAuthorityName="Benson" baseAuthorityYear="1860" class="Gastropoda" family="Achatinidae" genus="Rishetia" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="hastula">
<emphasis id="4EFF0B263D424D3F1638AC14F891809A" italics="true">Rishetia hastula</emphasis>
</taxonomicName>
, neotype
<named-content id="2ACBDD3124B03D872318AEAEBE4EABDE" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">NHMUK</named-content>
1906.1. 1.880 from Darjiling
<emphasis id="2E348CCDDCC178D8A3FFBF93B0D6FE1E" bold="true">E, F</emphasis>
<taxonomicName id="DC0782735F7C38F41B3D1812C77FBFF0" authorityName="Man &amp; Ablett &amp; Lwin &amp; Sutcharit &amp; Panha" authorityYear="2024" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1920" class="Gastropoda" family="Achatinidae" genus="Rishetia" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="kentungensis">
<emphasis id="802415FEA5E11A9B0DBC1B39A369C54B" italics="true">Rishetia kentungensis</emphasis>
</taxonomicName>
, syntypes
<named-content id="48F85599A92A9C9D0DAA22B31F3063AE" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">NHMUK</named-content>
1903.7. 1.3650 from Mong Sing, Siam Boundary
<emphasis id="159E1E469B734C2FE817A4AB8A353B7D" bold="true">G</emphasis>
<taxonomicName id="B42EE9B8D9C98ADB1E1A98174B3468B0" authorityName="Man &amp; Ablett &amp; Lwin &amp; Sutcharit &amp; Panha" authorityYear="2024" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1920" class="Gastropoda" family="Achatinidae" genus="Rishetia" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="limborgi">
<emphasis id="FEE0F71293DCE426B95FD8B06AF3AC93" italics="true">Rishetia limborgi</emphasis>
</taxonomicName>
, holotype
<named-content id="69B1D4D8C19647196BCF1B4D2F077C47" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">NHMUK</named-content>
1903.7. 1.3200 from Tenasserim
<emphasis id="7A7898CACA7975D4B1E55EFF19E8EDB4" bold="true">H, I</emphasis>
<taxonomicName id="6202F0D0DE6145FBE9D9C17B9C23A6A3" authorityName="Man &amp; Ablett &amp; Lwin &amp; Sutcharit &amp; Panha" authorityYear="2024" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1920" class="Gastropoda" family="Achatinidae" genus="Rishetia" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="nathiana">
<emphasis id="84F927D2F2A4EBB65575992E178A09B4" italics="true">Rishetia nathiana</emphasis>
</taxonomicName>
, syntypes
<named-content id="042A30166B7383572F33B993CA7E3FAF" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">NHMUK</named-content>
1906.1. 1.2206 from Bassein District.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EBNEK" type="diagnosis">
<paragraph id="426FDD9511BDE2957EE0D6DEA074B5E8">
<heading id="ACA130A94A38D93E4239C868F8D4E01C" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="E106F5FA8B94DAACBF758867AFABA6B9">Shell elongate, turreted, and regularly attenuated; apex rounded and blunt; subsequent whorls nearly smooth with fine irregularly spaced radial ridges throughout, and coarser ridges appearing near suture. Suture impressed and whorls flattened. Aperture ovate; columella concave and truncated.</paragraph>
</subSubSection>
<subSubSection id="SECID0EGNEK" type="distribution">
<paragraph id="2E79CB59A4E6C00BC2C441B2D8A983CE">
<heading id="1E5D80BE097E874A4F00FD904B23137E" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="8E3B23BE63EF955A12CBB951FEBD14CB">
This subspecies was originally described from the Naga Hills. The locality reported in
<collectingCountry id="EEB6CBF698B1E628902BE1871F3A0AF4" name="Myanmar">Myanmar</collectingCountry>
is Somra Town (~
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EPNEK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[94.687333,25.362111]}">
<geoCoordinate id="ADE1D8242488E46813075DD44E8A4942" degrees="25" direction="north" minutes="21" orientation="latitude" precision="1" seconds="43.6" value="25.362112">25 ° 21 ' 43.6 &quot; N</geoCoordinate>
,
<geoCoordinate id="B8C4CB7D608A32D803F0244143FDEE6C" degrees="94" direction="east" minutes="41" orientation="longitude" precision="1" seconds="14.4" value="94.68733">94 ° 41 ' 14.4 &quot; E</geoCoordinate>
</named-content>
), a mountainous area in Leshi or Layshi Township,
<collectingRegion id="837A84C0296F30AA4FF5658FA1251D01" country="Myanmar" name="Sagaing">Sagaing Region</collectingRegion>
in northwestern
<collectingCountry id="4BF9C2C58BF2C4651400C38D380181E8" name="Myanmar">Myanmar</collectingCountry>
. It is neighbouring Nagaland to the west and
<collectingRegion id="25C8CD6F02F33C46DFD7EA146F151440" country="Myanmar" name="Kachin">Kachin State</collectingRegion>
to the east and along the Chindwin River.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUNEK" type="remarks">
<paragraph id="72C000FD282DE56A1EF22A2D7882F264">
<heading id="9FFCEEBF1B97D7773B34E4C8E154CAC3" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="7FB2E96EDF6E7024374D4F81D86881DD">
The name
<emphasis id="B43F2C5AAEEEF9777F72E0ED690766CF" italics="true">maxwelli</emphasis>
was stated twice in the same publication, on pages 45 and 61. The first mention is
<taxonomicName id="1553FF51AEC702419911EEA29AA4CF1C" authorityName="Godwin-Austen" authorityYear="1920" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="variety" species="burrailensis" subGenus="Rishetia" variety="maxwelli">
<emphasis id="4A529266CABAAE34ABA8F030A201C251" italics="true">Glessula</emphasis>
(
<emphasis id="FF70BADE0B1A8ED99FED2F3365AB713D" italics="true">Rishetia</emphasis>
)
<emphasis id="863760AA694C9F843C3E94B8664065C7" italics="true">burrailensis</emphasis>
, var.
<emphasis id="26CBB0AC9D9A2ECDD95B88ED0D231353" italics="true">maxwelli</emphasis>
</taxonomicName>
; when proposing this name,
<bibRefCitation id="E844401E7992B1B966D4D08066073461" author="Godwin-Austen HH" firstAuthor="Godwin-Austen" page="45" refId="B39" refString="Godwin-Austen HH (1920) Land and freshwater Mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other islands of the Indian Ocean. Supplementary to Messrs. Theobald and Hanleys Conchologia Indica. London, Taylor &amp; Francis, Volume 3, 165. [pls 159165] https://www.biodiversitylibrary.org/item/57585" url="https://www.biodiversitylibrary.org/item/57585" year="1920">Godwin-Austen (1920: 45)</bibRefCitation>
recognised
<specimenCount id="4E7140A7EF3CEB46EA5F7F53012FF199" count="2" type="generic">two specimen</specimenCount>
lots from Naga Hills … (collected by Col. H. St. P. Maxwell) and from Somra … Upper
<collectingCountry id="062CA2857C47BB2A6290F09CAB4C54BF" name="Myanmar">Burma</collectingCountry>
(collected by F. Ede). Although the species description is brief, Godwin-Austen provides the catalogue number No. 1717
<abbrev content-type="institution" id="ABBRID0EFPEK" xlink_title="used in old labels for the British Museum">
<collectionCode id="E37A4836D305C60B28CF610E6BE2484D">B</collectionCode>
. M
</abbrev>
. , two sets of shell dimensions, and illustrations of two shells from Naga Hill. We have examined this specimen lot,
<named-content id="49DE7A8A08FF4D1DF361DE0391497DD6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
<collectionCode id="CB9FBFD40A7A2A136A8D246AA8A1F48F" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
1903.7. 1.1717, from Naga Hills, which consists of four shells. In the original description,
<bibRefCitation id="23B8467857DBEED7837D8519F3EAD047" author="Godwin-Austen HH" firstAuthor="Godwin-Austen" page="46" refId="B39" refString="Godwin-Austen HH (1920) Land and freshwater Mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other islands of the Indian Ocean. Supplementary to Messrs. Theobald and Hanleys Conchologia Indica. London, Taylor &amp; Francis, Volume 3, 165. [pls 159165] https://www.biodiversitylibrary.org/item/57585" url="https://www.biodiversitylibrary.org/item/57585" year="1920">Godwin-Austen (1920: 46)</bibRefCitation>
further states, Size: Fig.
<figureCitation id="B8EA3123809C24407AD016310BE5BA46" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A D Glessula blanfordiana A, B syntypes NHMUK 1988150 from Ponsee, Yunnan, W. China and C, D specimen CUMZ 13067 from Taunggyi, Shan State, Myanmar E, F Glessula crassilabris E syntype UMZC I. 102430 from Bengal (after Preece et al. 2022: fig. 51 e) and F specimen NHMUK from Cherrapoonji (M 10604 / 2)." figureDoi="10.3897/zookeys.1208.116083.figure5" httpUri="https://binary.pensoft.net/fig/1104503">5</figureCitation>
. Type and in the figure caption as Fig.
<figureCitation id="744CFE8A1395F8233DFA5ABC84699D83" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A D Glessula blanfordiana A, B syntypes NHMUK 1988150 from Ponsee, Yunnan, W. China and C, D specimen CUMZ 13067 from Taunggyi, Shan State, Myanmar E, F Glessula crassilabris E syntype UMZC I. 102430 from Bengal (after Preece et al. 2022: fig. 51 e) and F specimen NHMUK from Cherrapoonji (M 10604 / 2)." figureDoi="10.3897/zookeys.1208.116083.figure5" httpUri="https://binary.pensoft.net/fig/1104503">5</figureCitation>
. (1
<superScript id="66CEEDBADADE1F28040B4EB6CCD4743D">st</superScript>
Type) from Naga Hills and Fig.
<figureCitation id="08B58B0B5E8352C61B7581DB2B53217B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A F Glessula feddeni A, B syntypes NHMUK 1906.2. 2.261 from Shan Hills, E. of Ava C specimen NHMUK 1906.3. 3.41 from Shan Hills and D F specimen CUMZ 13068 from Pyinyaung Village, Mandalay Region, Myanmar." figureDoi="10.3897/zookeys.1208.116083.figure6" httpUri="https://binary.pensoft.net/fig/1104504">6</figureCitation>
. (2
<superScript id="E2774B3CC16EA0C9D2DB281FC6CDEDFB">nd</superScript>
Type) from Naga Hills. It seemed that Godwin-Austen intended to designate the specimen figured in plate 160, fig. 5, as the unique name-bearing type, which we consider as the
<typeStatus id="6AA46A609358B9366989DB11ED30C006" type="holotype">holotype</typeStatus>
. The other
<specimenCount id="0E46B7128A4AF4FF3028847EC76BE870" count="3" type="generic">three specimens</specimenCount>
from the same lot are therefore
<typeStatus id="9A79CB724D0E1424A0693AFF7FBBA44F" type="paratype">paratypes</typeStatus>
.
</paragraph>
<paragraph id="8813EE52E92EC4014B6069C954681E5F">
The second mention is
<taxonomicName id="E5D15A91B0044466756597033B3B947E" authorityName="Godwin-Austen" authorityYear="1920" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="variety" species="burrailensis" variety="maxwelli">
<emphasis id="442340C3DDE47957236CA0C84F068AB5" italics="true">Glessula burrailensis</emphasis>
, var.
<emphasis id="2EC9DA3402D3E5605EBC8B025A7E3AF5" italics="true">maxwelli</emphasis>
</taxonomicName>
, for which
<bibRefCitation id="7447B55555AA189640D92636A7F72512" author="Godwin-Austen HH" firstAuthor="Godwin-Austen" page="61" refId="B39" refString="Godwin-Austen HH (1920) Land and freshwater Mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other islands of the Indian Ocean. Supplementary to Messrs. Theobald and Hanleys Conchologia Indica. London, Taylor &amp; Francis, Volume 3, 165. [pls 159165] https://www.biodiversitylibrary.org/item/57585" url="https://www.biodiversitylibrary.org/item/57585" year="1920">Godwin-Austen (1920: 61)</bibRefCitation>
provided a very detailed description together with specimen lot number No. 3742
<abbrev content-type="institution" id="ABBRID0EDREK" xlink_title="used in old labels for the British Museum">
<collectionCode id="9351DFE4E6D8071E39AB8AC3C0C71032">B</collectionCode>
. M
</abbrev>
. , two sets of shell measurements, from Somra Tracts, Somra Khulen Post, Upper
<collectingCountry id="59A5EAFAAFEDFAA9627F7DD974278C5B" name="Myanmar">Burma</collectingCountry>
, but without illustration. The
<named-content id="5ABBD48D861E638F8ACD14242067F058" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
<collectionCode id="F9F3A4C7BF9EBAC52835492E69653439" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
collections contain specimen lot
<named-content id="C4A93DA6D042248EBD48F1C08EEFF723" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
<collectionCode id="EA451EF1C6F1D3DFB7048975CFDB6B9D" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
1903.7. 1.3742, consisting of
<specimenCount id="5DE4C32F7C6CB261740081E217F98942" count="8" type="generic">eight specimens</specimenCount>
with the original label stating
<taxonomicName id="91113F99938F91A116E62E6827B55F5C" authority=", GA." authorityName="GA." rank="variety" variety="somraensis">
var.
<emphasis id="28D66CAD388FD9AC6C471DA7B0D178F7" italics="true">somraensis</emphasis>
, GA.
</taxonomicName>
. This manuscript name has never properly been published and made available (
<collectionCode id="0FDD6BB075A0DA8DCE66F7B91DB3C5B4">ICZN</collectionCode>
1999: Art. 10), the specimen lot
<named-content id="67FC484703BDAD99213D5B17D77A621A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
<collectionCode id="1E9D5E619B2D0F0B561281B2E5295BB5" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
1903.7. 1.3742, with its original label stating
<taxonomicName id="90B8BFFFD9B7985FA4A8E4D502ED3574" rank="variety" variety="somraensis">
var.
<emphasis id="3FBA886BF6CC336B971851DA782CE2EB" italics="true">somraensis</emphasis>
</taxonomicName>
is considered as the
<typeStatus id="026B6B804784B5F96C5BD2535F28D8F0" type="syntype">syntype</typeStatus>
. Therefore, this second combination becomes an available name (
<collectionCode id="AF04E52DD7D7B85584C308D483BD7D01">ICZN</collectionCode>
1999: Arts 11, 12), and it is considered a junior (primary) homonym (
<collectionCode id="171BE5D3786B58150DD5206BF7996255">ICZN</collectionCode>
1999: Arts 24.2. 1, 24.2. 2, 53.3, 57.2). Additionally,
<typeStatus id="59D336D058C7E0B17F432E0627CE5916" type="syntype">syntypes</typeStatus>
also include,
<named-content id="C3682E21DD06F98EC776526AC713E231" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
<collectionCode id="348C124739213FF79FC1346A93716283" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
1988147 (4 shells), from Somra, Khulen Post, West of Kyendwin River, Upper Burmah, as
<bibRefCitation id="C61DA31B0EEFE1C0405BE1C7BC778984" author="Godwin-Austen HH" firstAuthor="Godwin-Austen" page="61" refId="B39" refString="Godwin-Austen HH (1920) Land and freshwater Mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other islands of the Indian Ocean. Supplementary to Messrs. Theobald and Hanleys Conchologia Indica. London, Taylor &amp; Francis, Volume 3, 165. [pls 159165] https://www.biodiversitylibrary.org/item/57585" url="https://www.biodiversitylibrary.org/item/57585" year="1920">Godwin-Austen (1920: 61)</bibRefCitation>
recognised in the original description.
</paragraph>
<paragraph id="7FD12CE88F1B17A7FDDFAFB671F97905">
After examining the type specimens carefully, we found no differences in the diagnostic characters between the senior homonym (
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1903.7. 1.1717 / 1 and
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1903.7. 1.1717 / 24) and junior homonym (
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1903.7. 1.3742,
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1903.7. 1.3744 and
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1988147) specimens, especially shell shape, shell sculpture, and protoconch sculpture. The only detected difference is the brownish or yellowish brown periostracum, which we consider as intraspecific variation. Therefore, it seems unnecessary to propose a new replacement name for this junior homonym.
</paragraph>
</subSubSection>
</treatment>
</document>