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<document id="D27A5CE2B6B89281C8A72D3230D32B29" ID-ISSN="0253-116X" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722341515648" checkinUser="felipe" docAuthor="Assing, V." docDate="2006" docId="DE7E8782C331761D3003FCE1FC4A77BF" docLanguage="en" docName="LinzerbiolBeitr.2006.38.2.1129-1137.pdf" docOrigin="Linzer biologische Beiträge 38 (2)" docStyle="DocumentStyle:6260912E6386EE56293601EB6FBA9D25.2:LinzerbiolBeitr.2006-.journal_article.type2" docStyleId="6260912E6386EE56293601EB6FBA9D25" docStyleName="LinzerbiolBeitr.2006-.journal_article.type2" docTitle="Atheta tenebrarum Assing, 2006, nov.sp." docType="treatment" docVersion="1" lastPageNumber="1137" masterDocId="2247FFFAC33776153157FFDAFF89740F" masterDocTitle="Three new species of Staphylinidae from Spain, with a new synonymy (Insecta: Coleoptera)" masterLastPageNumber="1137" masterPageNumber="1129" pageNumber="1129" updateTime="1722350314231" updateUser="felipe" zenodo-license-document="CC-BY-4.0">
<mods:mods id="62639779E94BEE4722765A097ECB755E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="90DF4DDFD9818F6E1F56751DEAA27579">
<mods:title id="EAF11D1B8F7A58C0FFB6B412C46CC71D">Three new species of Staphylinidae from Spain, with a new synonymy (Insecta: Coleoptera)</mods:title>
</mods:titleInfo>
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<mods:namePart id="E8816E85FABD09071B4D24B7C789B964">Assing, V.</mods:namePart>
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<mods:typeOfResource id="F75EC6088CEB45B7AEF52A94FA9E81C2">text</mods:typeOfResource>
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<mods:title id="E5CBCEF36E1D4DF1B19280677F2AC0BD">Linzer biologische Beiträge</mods:title>
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<mods:part id="6E0C45965D90A092E2152E89388326C1">
<mods:date id="0B74C1328026305C0B98FB0BAA10C6AF">2006</mods:date>
<mods:detail id="11D37E82655A666920C1F67AE79F13DA" type="pubDate">
<mods:number id="8FBBD60DD41A48193836D11D917321CF">2006-12-29</mods:number>
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<mods:detail id="D1FF6E2883880A60D48B8AE3436645EA" type="volume">
<mods:number id="8768763C727517FAC463AB0A65384882">38</mods:number>
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<mods:number id="B0F5067B11DC120C05477D45DF0BFA52">2</mods:number>
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<mods:classification id="D56E9F12D2172E8159E7C186DA2618E6">journal article</mods:classification>
<mods:identifier id="B82E2DEF698F00523E46FA748ABC8532" type="ISSN">0253-116X</mods:identifier>
</mods:mods>
<treatment id="DE7E8782C331761D3003FCE1FC4A77BF" LSID="urn:lsid:plazi:treatment:DE7E8782C331761D3003FCE1FC4A77BF" httpUri="http://treatment.plazi.org/id/DE7E8782C331761D3003FCE1FC4A77BF" lastPageId="8" pageId="6">
<subSubSection id="1ECD651FC33176133003FCE1FCCF775E" box="[340,838,827,849]" pageId="6" type="nomenclature">
<paragraph id="56683694C33176133003FCE1FCCF775E" blockId="6.[340,838,827,849]" box="[340,838,827,849]" pageId="6">
<heading id="0D2081F8C33176133003FCE1FCCF775E" box="[340,838,827,849]" fontSize="8" level="2" pageId="6" reason="2">
<emphasis id="64A3EA86C33176133003FCE1FCCF775E" bold="true" box="[340,838,827,849]" pageId="6">
<taxonomicName id="91D74D17C33176133003FCE1FD90775E" ID-CoL="688QK" authorityName="Assing" authorityYear="2006" box="[340,537,827,849]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="tenebrarum" status="sp. nov.">
<emphasis id="64A3EA86C33176133003FCE1FD90775E" bold="true" box="[340,537,827,849]" italics="true" pageId="6">Atheta tenebrarum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7F9057FDC33176133377FCE1FDE5775E" box="[544,620,827,849]" pageId="6" rank="species">nov.sp.</taxonomicNameLabel>
(
<figureCitation id="CEEC2A11C3317613332EFCE1FD65775E" box="[633,748,827,849]" captionStart="Figs 22-33" captionStartId="7.[340,380,1613,1633]" captionTargetBox="[340,1240,425,1579]" captionTargetId="figure-86@7.[340,1246,653,1129]" captionTargetPageId="7" captionText="Figs 22-33: Atheta tenebrarum nov.sp.: (22) habitus; (23) forebody; (24) head in lateral view; (25) antenna; (26) 3 tergite VIII; (27) posterior margin of 3 sternite VIII; (28-30) median lobe of aedeagus in lateral and in ventral view; (31) ♀ tergite VIII; (32) ♀ sternite VIII; (33) spermatheca. Scales: 22-23: 1.0 mm; 24-27, 31-32: 0.5 mm; 28-30, 33: 0.2 mm." pageId="6">Figs 22-33</figureCitation>
;
<figureCitation id="CEEC2A11C331761333AFFCE1FCB7775E" box="[760,830,827,849]" captionStart="Map 1" captionStartId="3.[340,385,1351,1371]" captionTargetBox="[340,1247,707,1340]" captionTargetId="figure-370@3.[340,1247,707,1340]" captionTargetPageId="3" captionText="Map 1: Distributions of Phloeocharis baenai nov.sp. (open circle), Geostiba jaenica nov.sp. (filled circles), and Atheta tenebrarum nov.sp. (square) in southern Spain." pageId="6">Map 1</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1ECD651FC331761D3003FCBBFC4A77BF" lastPageId="8" pageId="6" type="materials_examined">
<paragraph id="56683694C33176133003FCBBFE1F77AD" blockId="6.[340,1247,864,930]" pageId="6">
<materialsCitation id="E6BF3CC9C33176133003FCBBFBD8777A" box="[340,1105,864,885]" collectingDate="2006-03-18" collectorName="La Iruela" location="Cueva Secreta del Sagreo" pageId="6" specimenCount="1" typeStatus="holotype">
<typeStatus id="896C8836C33176133003FCBBFE20777A" box="[340,425,865,885]" pageId="6" type="holotype">Holotype</typeStatus>
3: E - Jaén,
<collectorName id="FB225342C3317613337DFCBBFD09777A" box="[554,640,865,885]" pageId="6">La Iruela</collectorName>
,
<location id="5308604FC331761333DCFCBBFCF2777A" LSID="urn:lsid:plazi:treatment:DE7E8782C331761D3003FCE1FC4A77BF:5308604FC331761333DCFCBBFCF2777A" box="[651,891,865,885]" name="Cueva Secreta del Sagreo" pageId="6">Cueva Secreta del Sagreo</location>
,
<date id="22691054C331761332D1FCBBFC67777A" box="[902,1006,865,885]" pageId="6" value="2006-03-18">
<collectingDate id="322DE9BCC331761332D1FCBBFC67777A" box="[902,1006,865,885]" pageId="6" value="2006-03-18">18.III.2006</collectingDate>
</date>
, leg. GEV
</materialsCitation>
/
<typeStatus id="896C8836C33176133531FCBBFB4C777A" box="[1126,1221,865,885]" pageId="6" type="holotype">Holotypus</typeStatus>
3
<taxonomicName id="91D74D17C33176133003FCA2FD887783" authorityName="Assing" authorityYear="2006" box="[340,513,888,908]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="tenebrarum" status="sp. nov.">
<emphasis id="64A3EA86C33176133003FCA2FD887783" box="[340,513,888,908]" italics="true" pageId="6">Atheta tenebrarum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7F9057FDC3317613335FFCA2FDBA7783" box="[520,563,888,908]" pageId="6" rank="species">sp.n.</taxonomicNameLabel>
det.
<collectionCode id="30C6AE51C33176133335FCA2FDFC7783" box="[610,629,888,908]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" type="Museum">V</collectionCode>
. Assing 2006 (cAss).
<typeStatus id="896C8836C33176133214FCA2FC167783" box="[835,927,888,908]" pageId="6" type="paratype">Paratypes</typeStatus>
: 233,
<specimenCount id="40D1FD1DC331761332B0FCA2FB8C779E" box="[999,1029,888,913]" count="1" pageId="6" type="female">1♀</specimenCount>
: same data as
<typeStatus id="896C8836C331761335C7FCA2FB567783" box="[1168,1247,888,908]" pageId="6" type="holotype">holotype</typeStatus>
(cAss).
</paragraph>
<paragraph id="56683694C33176133003FC68FBD57010" blockId="6.[340,1248,945,1055]" pageId="6">
D e s c r i p t i o n: Species of moderately large size,
<quantity id="912F9B71C331761332C6FC6BFB8277C8" box="[913,1035,945,967]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.2" metricValueMin="3.8" pageId="6" unit="mm" value="4.0" valueMax="4.2" valueMin="3.8">3.8-4.2 mm</quantity>
; facies as in
<figureCitation id="CEEC2A11C331761335C7FC6BFB5577C8" box="[1168,1244,945,967]" captionStart="Figs 22-33" captionStartId="7.[340,380,1613,1633]" captionTargetBox="[340,1240,425,1579]" captionTargetId="figure-86@7.[340,1246,653,1129]" captionTargetPageId="7" captionText="Figs 22-33: Atheta tenebrarum nov.sp.: (22) habitus; (23) forebody; (24) head in lateral view; (25) antenna; (26) 3 tergite VIII; (27) posterior margin of 3 sternite VIII; (28-30) median lobe of aedeagus in lateral and in ventral view; (31) ♀ tergite VIII; (32) ♀ sternite VIII; (33) spermatheca. Scales: 22-23: 1.0 mm; 24-27, 31-32: 0.5 mm; 28-30, 33: 0.2 mm." pageId="6">Fig. 22</figureCitation>
. Coloration: head dark brown; pronotum reddish brown; elytra yellowish to yellowish brown; abdomen reddish brown, with the preapical segments somewhat infuscate; legs yellowish; antennae dark brown, with the basal 3-4 antennomeres yellowish.
</paragraph>
<paragraph id="56683694C33176133003FBF4FB3270D9" blockId="6.[340,1248,1070,1238]" pageId="6">
Head slightly wider than long; eyes of normal size (
<figureCitation id="CEEC2A11C3317613322FFBF4FC4F704B" box="[888,966,1070,1092]" captionStart="Figs 22-33" captionStartId="7.[340,380,1613,1633]" captionTargetBox="[340,1240,425,1579]" captionTargetId="figure-86@7.[340,1246,653,1129]" captionTargetPageId="7" captionText="Figs 22-33: Atheta tenebrarum nov.sp.: (22) habitus; (23) forebody; (24) head in lateral view; (25) antenna; (26) 3 tergite VIII; (27) posterior margin of 3 sternite VIII; (28-30) median lobe of aedeagus in lateral and in ventral view; (31) ♀ tergite VIII; (32) ♀ sternite VIII; (33) spermatheca. Scales: 22-23: 1.0 mm; 24-27, 31-32: 0.5 mm; 28-30, 33: 0.2 mm." pageId="6">Fig. 24</figureCitation>
), approximately as long as postocular region in dorsal view, or slightly longer; postgenae finely carinate ventrally; integument with distinct microreticulation; puncturation rather dense and fine, barely visible in the microsculpture (
<figureCitation id="CEEC2A11C331761333D2FB5CFD587093" box="[645,721,1158,1180]" captionStart="Figs 22-33" captionStartId="7.[340,380,1613,1633]" captionTargetBox="[340,1240,425,1579]" captionTargetId="figure-86@7.[340,1246,653,1129]" captionTargetPageId="7" captionText="Figs 22-33: Atheta tenebrarum nov.sp.: (22) habitus; (23) forebody; (24) head in lateral view; (25) antenna; (26) 3 tergite VIII; (27) posterior margin of 3 sternite VIII; (28-30) median lobe of aedeagus in lateral and in ventral view; (31) ♀ tergite VIII; (32) ♀ sternite VIII; (33) spermatheca. Scales: 22-23: 1.0 mm; 24-27, 31-32: 0.5 mm; 28-30, 33: 0.2 mm." pageId="6">Fig. 23</figureCitation>
). Antennae slender; antennomere III slightly longer than II, IV weakly oblong, V-IX approximately as long as wide, X weakly transverse, and XI approximately as long as the combined length of IX-X or slightly longer (
<figureCitation id="CEEC2A11C33176133532FB1AFB3970D9" box="[1125,1200,1216,1238]" captionStart="Figs 22-33" captionStartId="7.[340,380,1613,1633]" captionTargetBox="[340,1240,425,1579]" captionTargetId="figure-86@7.[340,1246,653,1129]" captionTargetPageId="7" captionText="Figs 22-33: Atheta tenebrarum nov.sp.: (22) habitus; (23) forebody; (24) head in lateral view; (25) antenna; (26) 3 tergite VIII; (27) posterior margin of 3 sternite VIII; (28-30) median lobe of aedeagus in lateral and in ventral view; (31) ♀ tergite VIII; (32) ♀ sternite VIII; (33) spermatheca. Scales: 22-23: 1.0 mm; 24-27, 31-32: 0.5 mm; 28-30, 33: 0.2 mm." pageId="6">Fig. 25</figureCitation>
).
</paragraph>
<paragraph id="56683694C33176133003FB3CFDD07181" blockId="6.[340,1248,1254,1422]" pageId="6">
Pronotum approximately 1.3 times as wide as head and 1.35 times as wide as long; maximal width a short distance anterior to middle; posterior angles obtuse, weakly marked; puncturation and microreticulation similar to those of head (
<figureCitation id="CEEC2A11C33176133562FAFAFB0C7139" box="[1077,1157,1312,1334]" captionStart="Figs 22-33" captionStartId="7.[340,380,1613,1633]" captionTargetBox="[340,1240,425,1579]" captionTargetId="figure-86@7.[340,1246,653,1129]" captionTargetPageId="7" captionText="Figs 22-33: Atheta tenebrarum nov.sp.: (22) habitus; (23) forebody; (24) head in lateral view; (25) antenna; (26) 3 tergite VIII; (27) posterior margin of 3 sternite VIII; (28-30) median lobe of aedeagus in lateral and in ventral view; (31) ♀ tergite VIII; (32) ♀ sternite VIII; (33) spermatheca. Scales: 22-23: 1.0 mm; 24-27, 31-32: 0.5 mm; 28-30, 33: 0.2 mm." pageId="6">Fig. 23</figureCitation>
); pubescence directed cephalad in anterior 1/4 of midline, caudad in posterior 3/4, and diagonally latero-caudad in lateral area; marginal setae short, slightly shorter than maximal width of antennomere III.
</paragraph>
<paragraph id="56683694C33176133003FA47FBDE71E1" blockId="6.[340,1248,1437,1518]" pageId="6">
Elytra about 1.25 times as wide and at suture almost as long as pronotum; microsculpture and puncturation similar to those of head and pronotum; pubescence short and depressed (
<figureCitation id="CEEC2A11C3317613300BFA02FE2E71E1" box="[348,423,1496,1518]" captionStart="Figs 22-33" captionStartId="7.[340,380,1613,1633]" captionTargetBox="[340,1240,425,1579]" captionTargetId="figure-86@7.[340,1246,653,1129]" captionTargetPageId="7" captionText="Figs 22-33: Atheta tenebrarum nov.sp.: (22) habitus; (23) forebody; (24) head in lateral view; (25) antenna; (26) 3 tergite VIII; (27) posterior margin of 3 sternite VIII; (28-30) median lobe of aedeagus in lateral and in ventral view; (31) ♀ tergite VIII; (32) ♀ sternite VIII; (33) spermatheca. Scales: 22-23: 1.0 mm; 24-27, 31-32: 0.5 mm; 28-30, 33: 0.2 mm." pageId="6">Fig. 23</figureCitation>
). Hind wings fully developed. Tibiae without long prominent setae.
</paragraph>
<paragraph id="56683694C33176133003FA27FC327241" blockId="6.[340,1248,1533,1614]" pageId="6">
Abdomen slightly narrower than elytra, widest at segments IV/
<collectionCode id="30C6AE51C331761332BAFA27FC76721C" box="[1005,1023,1533,1555]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" type="Museum">V</collectionCode>
(
<figureCitation id="CEEC2A11C33176133547FA27FBE9721C" box="[1040,1120,1533,1555]" captionStart="Figs 22-33" captionStartId="7.[340,380,1613,1633]" captionTargetBox="[340,1240,425,1579]" captionTargetId="figure-86@7.[340,1246,653,1129]" captionTargetPageId="7" captionText="Figs 22-33: Atheta tenebrarum nov.sp.: (22) habitus; (23) forebody; (24) head in lateral view; (25) antenna; (26) 3 tergite VIII; (27) posterior margin of 3 sternite VIII; (28-30) median lobe of aedeagus in lateral and in ventral view; (31) ♀ tergite VIII; (32) ♀ sternite VIII; (33) spermatheca. Scales: 22-23: 1.0 mm; 24-27, 31-32: 0.5 mm; 28-30, 33: 0.2 mm." pageId="6">Fig. 22</figureCitation>
); tergal surfaces with fine transverse microscupture everywhere; puncturation relatively sparse and very fine, barely noticeable; pubescence dark and depressed.
</paragraph>
<paragraph id="56683694C33176133003F987FE1672A2" blockId="6.[340,1248,1629,1709]" pageId="6">
3: posterior margin of tergite VIII weakly concave and weakly serrate (
<figureCitation id="CEEC2A11C3317613356BF987FB00727C" box="[1084,1161,1629,1651]" captionStart="Figs 22-33" captionStartId="7.[340,380,1613,1633]" captionTargetBox="[340,1240,425,1579]" captionTargetId="figure-86@7.[340,1246,653,1129]" captionTargetPageId="7" captionText="Figs 22-33: Atheta tenebrarum nov.sp.: (22) habitus; (23) forebody; (24) head in lateral view; (25) antenna; (26) 3 tergite VIII; (27) posterior margin of 3 sternite VIII; (28-30) median lobe of aedeagus in lateral and in ventral view; (31) ♀ tergite VIII; (32) ♀ sternite VIII; (33) spermatheca. Scales: 22-23: 1.0 mm; 24-27, 31-32: 0.5 mm; 28-30, 33: 0.2 mm." pageId="6">Fig. 26</figureCitation>
), that of sternite VIII convex (
<figureCitation id="CEEC2A11C33176133366F9A0FDF6729F" box="[561,639,1658,1680]" captionStart="Figs 22-33" captionStartId="7.[340,380,1613,1633]" captionTargetBox="[340,1240,425,1579]" captionTargetId="figure-86@7.[340,1246,653,1129]" captionTargetPageId="7" captionText="Figs 22-33: Atheta tenebrarum nov.sp.: (22) habitus; (23) forebody; (24) head in lateral view; (25) antenna; (26) 3 tergite VIII; (27) posterior margin of 3 sternite VIII; (28-30) median lobe of aedeagus in lateral and in ventral view; (31) ♀ tergite VIII; (32) ♀ sternite VIII; (33) spermatheca. Scales: 22-23: 1.0 mm; 24-27, 31-32: 0.5 mm; 28-30, 33: 0.2 mm." pageId="6">Fig. 27</figureCitation>
); median lobe of aedeagus of highly distinctive shape (
<figureCitation id="CEEC2A11C331761335E1F9A0FE1A72A2" captionStart="Figs 22-33" captionStartId="7.[340,380,1613,1633]" captionTargetBox="[340,1240,425,1579]" captionTargetId="figure-86@7.[340,1246,653,1129]" captionTargetPageId="7" captionText="Figs 22-33: Atheta tenebrarum nov.sp.: (22) habitus; (23) forebody; (24) head in lateral view; (25) antenna; (26) 3 tergite VIII; (27) posterior margin of 3 sternite VIII; (28-30) median lobe of aedeagus in lateral and in ventral view; (31) ♀ tergite VIII; (32) ♀ sternite VIII; (33) spermatheca. Scales: 22-23: 1.0 mm; 24-27, 31-32: 0.5 mm; 28-30, 33: 0.2 mm." pageId="6">Fig. 28-30</figureCitation>
).
</paragraph>
<paragraph id="56683694C33176133003F919FE2A7302" blockId="6.[340,1248,1725,1805]" pageId="6">
<specimenCount id="40D1FD1DC33176133003F919FEE072D7" box="[340,361,1731,1752]" count="1" pageId="6" type="female">♀</specimenCount>
: posterior margin of tergite VIII in the middle weakly concave (
<figureCitation id="CEEC2A11C33176133551F967FBDB72DC" box="[1030,1106,1725,1747]" captionStart="Figs 22-33" captionStartId="7.[340,380,1613,1633]" captionTargetBox="[340,1240,425,1579]" captionTargetId="figure-86@7.[340,1246,653,1129]" captionTargetPageId="7" captionText="Figs 22-33: Atheta tenebrarum nov.sp.: (22) habitus; (23) forebody; (24) head in lateral view; (25) antenna; (26) 3 tergite VIII; (27) posterior margin of 3 sternite VIII; (28-30) median lobe of aedeagus in lateral and in ventral view; (31) ♀ tergite VIII; (32) ♀ sternite VIII; (33) spermatheca. Scales: 22-23: 1.0 mm; 24-27, 31-32: 0.5 mm; 28-30, 33: 0.2 mm." pageId="6">Fig. 31</figureCitation>
), that of sternite VIII broadly convex and with long thin marginal setae (
<figureCitation id="CEEC2A11C33176133291F900FB9C72FF" box="[966,1045,1754,1776]" captionStart="Figs 22-33" captionStartId="7.[340,380,1613,1633]" captionTargetBox="[340,1240,425,1579]" captionTargetId="figure-86@7.[340,1246,653,1129]" captionTargetPageId="7" captionText="Figs 22-33: Atheta tenebrarum nov.sp.: (22) habitus; (23) forebody; (24) head in lateral view; (25) antenna; (26) 3 tergite VIII; (27) posterior margin of 3 sternite VIII; (28-30) median lobe of aedeagus in lateral and in ventral view; (31) ♀ tergite VIII; (32) ♀ sternite VIII; (33) spermatheca. Scales: 22-23: 1.0 mm; 24-27, 31-32: 0.5 mm; 28-30, 33: 0.2 mm." pageId="6">Fig. 32</figureCitation>
); spermatheca as in
<figureCitation id="CEEC2A11C33176133003F92DFE167302" box="[340,415,1783,1805]" captionStart="Figs 22-33" captionStartId="7.[340,380,1613,1633]" captionTargetBox="[340,1240,425,1579]" captionTargetId="figure-86@7.[340,1246,653,1129]" captionTargetPageId="7" captionText="Figs 22-33: Atheta tenebrarum nov.sp.: (22) habitus; (23) forebody; (24) head in lateral view; (25) antenna; (26) 3 tergite VIII; (27) posterior margin of 3 sternite VIII; (28-30) median lobe of aedeagus in lateral and in ventral view; (31) ♀ tergite VIII; (32) ♀ sternite VIII; (33) spermatheca. Scales: 22-23: 1.0 mm; 24-27, 31-32: 0.5 mm; 28-30, 33: 0.2 mm." pageId="6">Fig. 33</figureCitation>
.
</paragraph>
<paragraph id="56683694C33076123255FEBFFCBB7575" blockId="7.[770,818,357,378]" box="[770,818,357,378]" pageId="7">1136</paragraph>
<caption id="02A8661CC33076123003F997FC2672AA" pageId="7" startId="7.[340,380,1613,1633]" targetBox="[340,1240,425,1579]" targetPageId="7" targetType="figure">
<paragraph id="56683694C33076123003F997FC2672AA" blockId="7.[340,1249,1613,1701]" pageId="7">
<emphasis id="64A3EA86C33076123003F997FE31726E" bold="true" box="[340,440,1613,1633]" pageId="7">Figs 22-33</emphasis>
:
<taxonomicName id="91D74D17C33076123092F997FDF8726E" authorityName="Assing" authorityYear="2006" box="[453,625,1613,1633]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="tenebrarum" status="sp. nov.">
<emphasis id="64A3EA86C33076123092F997FDF8726E" box="[453,625,1613,1633]" italics="true" pageId="7">Atheta tenebrarum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7F9057FDC33076123320F997FD34726E" box="[631,701,1613,1633]" pageId="7" rank="species">nov.sp.</taxonomicNameLabel>
: (
<emphasis id="64A3EA86C3307612339AF997FD6D726E" bold="true" box="[717,740,1613,1633]" pageId="7">22</emphasis>
) habitus; (
<emphasis id="64A3EA86C3307612321FF997FCD6726E" bold="true" box="[840,863,1613,1633]" pageId="7">23</emphasis>
) forebody; (
<emphasis id="64A3EA86C33076123284F997FC63726E" bold="true" box="[979,1002,1613,1633]" pageId="7">24</emphasis>
) head in lateral view; (
<emphasis id="64A3EA86C33076123596F997FB51726E" bold="true" box="[1217,1240,1613,1633]" pageId="7">25</emphasis>
) antenna; (
<emphasis id="64A3EA86C330761230E5F9BEFE407277" bold="true" box="[434,457,1636,1656]" pageId="7">26</emphasis>
) 3 tergite VIII; (
<emphasis id="64A3EA86C3307612332EF9BEFD197277" bold="true" box="[633,656,1636,1656]" pageId="7">27</emphasis>
) posterior margin of 3 sternite VIII; (
<emphasis id="64A3EA86C3307612355EF9BEFBB77277" bold="true" box="[1033,1086,1636,1656]" pageId="7">28-30</emphasis>
) median lobe of aedeagus in lateral and in ventral view; (
<emphasis id="64A3EA86C3307612339FF9A0FD567281" bold="true" box="[712,735,1658,1678]" pageId="7">31</emphasis>
) ♀ tergite VIII; (
<emphasis id="64A3EA86C330761232D5F9A0FC107281" bold="true" box="[898,921,1658,1678]" pageId="7">32</emphasis>
) ♀ sternite VIII; (
<emphasis id="64A3EA86C33076123513F9A0FBD27281" bold="true" box="[1092,1115,1658,1678]" pageId="7">33</emphasis>
) spermatheca. Scales: 22-23: 1.0 mm; 24-27, 31-32: 0.5 mm; 28-30, 33: 0.2 mm.
</paragraph>
</caption>
<paragraph id="56683694C33F761D3255FEBFFCBB7575" blockId="8.[770,818,357,378]" box="[770,818,357,378]" pageId="8">1137</paragraph>
<paragraph id="56683694C33F761D3003FE75FD7475EF" blockId="8.[340,1247,430,480]" pageId="8">E t y m o l o g y: The name (Lat.: noun, genitive: of the darkness) alludes to the fact that this species was discovered in a cave.</paragraph>
<paragraph id="56683694C33F761D3003FE34FDFF7713" blockId="8.[340,1248,494,796]" pageId="8">
C o m p a r a t i v e n o t e s:
<taxonomicName id="91D74D17C33F761D33E5FE34FCFB760B" authorityName="Assing" authorityYear="2006" box="[690,882,494,516]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="tenebrarum">
<emphasis id="64A3EA86C33F761D33E5FE34FCFB760B" box="[690,882,494,516]" italics="true" pageId="8">Atheta tenebrarum</emphasis>
</taxonomicName>
is distinguished from all its congeners especially by the highly distinctive morphology of the median lobe of the aedeagus and also by the shape of the spermatheca. Based on external morphology (pronotal pubescence pattern, chaetotaxy) and on the primary and secondary sexual characters, the new species is closely related to
<taxonomicName id="91D74D17C33F761D33E3FD85FC14767B" authority="(KRAATZ)" baseAuthorityName="KRAATZ" box="[692,925,606,628]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="trinotata">
<emphasis id="64A3EA86C33F761D33E3FD85FCA5767B" box="[692,812,607,628]" italics="true" pageId="8">A. trinotata</emphasis>
(KRAATZ)
</taxonomicName>
,
<taxonomicName id="91D74D17C33F761D32FBFD85FBE1767B" authority="SHARP" authorityName="SHARP" box="[940,1128,606,628]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="hybrida">
<emphasis id="64A3EA86C33F761D32FBFD85FB92767B" box="[940,1051,606,628]" italics="true" pageId="8">A. hybrida</emphasis>
SHARP
</taxonomicName>
, and allied species, its closest relative being
<taxonomicName id="91D74D17C33F761D33E6FDA1FC97769F" box="[689,798,634,656]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="hybrida">
<emphasis id="64A3EA86C33F761D33E6FDA1FC97769F" box="[689,798,634,656]" italics="true" pageId="8">A. hybrida</emphasis>
</taxonomicName>
, as is suggested particularly by the similar internal structures and shape of the median lobe of the aedeagus. From both
<taxonomicName id="91D74D17C33F761D353EFD4DFB6976A3" box="[1129,1248,663,684]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="trinotata">
<emphasis id="64A3EA86C33F761D353EFD4DFB6976A3" box="[1129,1248,663,684]" italics="true" pageId="8">A. trinotata</emphasis>
</taxonomicName>
and
<taxonomicName id="91D74D17C33F761D30D7FD69FE6576C7" box="[384,492,690,712]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="hybrida">
<emphasis id="64A3EA86C33F761D30D7FD69FE6576C7" box="[384,492,690,712]" italics="true" pageId="8">A. hybrida</emphasis>
</taxonomicName>
, it is separated also by the longer antennae, the relatively larger eyes, the paler and broader pronotum, the uniformly yellowish elytra, the different coloration of the abdomen (in
<taxonomicName id="91D74D17C33F761D3359FD31FD01770F" box="[526,648,747,768]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="trinotata">
<emphasis id="64A3EA86C33F761D3359FD31FD01770F" box="[526,648,747,768]" italics="true" pageId="8">A. trinotata</emphasis>
</taxonomicName>
and
<taxonomicName id="91D74D17C33F761D3394FD31FCBA770F" box="[707,819,746,768]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="hybrida">
<emphasis id="64A3EA86C33F761D3394FD31FCBA770F" box="[707,819,746,768]" italics="true" pageId="8">A. hybrida</emphasis>
</taxonomicName>
almost uniformly blackish), and by the secondary sexual characters.
</paragraph>
<paragraph id="56683694C33F761D3003FCF1FC4A77BF" blockId="8.[340,1248,810,944]" pageId="8">
D i s t r i b u t i o n a n d b i o n o m i c s: The species is currently known only from one cave near La Iruela in the Sierra de Cazorla, eastern Jaén province (altitude: slightly less than
<quantity id="912F9B71C33F761D3352FCB8FDDC7777" box="[517,597,866,888]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="8" unit="m" value="1200.0">1200 m</quantity>
). The
<taxonomicName id="91D74D17C33F761D33C2FCB8FCA97777" box="[661,800,866,888]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
from this cave were collected using various methods, so that the particular circumstances, under which the
<specimenCount id="40D1FD1DC33F761D328FFCA4FBBF779B" box="[984,1078,894,916]" count="4" pageId="8" type="generic" typeStatus="type">four type</specimenCount>
specimens of
<taxonomicName id="91D74D17C33F761D359EFCA5FE4377BF" authorityName="Assing" authorityYear="2006" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="tenebrarum">
<emphasis id="64A3EA86C33F761D359EFCA5FE4377BF" italics="true" pageId="8">A. tenebrarum</emphasis>
</taxonomicName>
were found, are unknown (
<collectionCode id="30C6AE51C33F761D33B6FC40FCA377BF" box="[737,810,922,944]" pageId="8">BAENA</collectionCode>
, pers. comm.).
</paragraph>
</subSubSection>
</treatment>
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<emphasis id="64A3EA86C33476163003FA41FC3071BE" bold="true" box="[340,953,1435,1457]" pageId="3">
<taxonomicName id="91D74D17C33476163003FA41FD1071BE" ID-CoL="3FSQW" authorityName="Assing" authorityYear="2006" box="[340,665,1435,1457]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="jaenica" status="sp. nov." subGenus="Sipalotricha">
<emphasis id="64A3EA86C33476163003FA41FE2671BE" bold="true" box="[340,431,1435,1457]" italics="true" pageId="3">Geostiba</emphasis>
(
<emphasis id="64A3EA86C334761630E9FA41FDB771BE" bold="true" box="[446,574,1435,1457]" italics="true" pageId="3">Sipalotricha</emphasis>
)
<emphasis id="64A3EA86C3347616331AFA41FD1071BE" bold="true" box="[589,665,1435,1457]" italics="true" pageId="3">jaenica</emphasis>
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(
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;
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<typeStatus id="896C8836C33476163003FA18FE2071D9" box="[340,425,1474,1494]" pageId="3" type="holotype">Holotype</typeStatus>
3: E - No. 7,
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,
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Mancha Real, Sierra Almadén,
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,
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,
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. Assing
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/
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3
<taxonomicName id="91D74D17C33476163246FA03FC2571E2" authorityName="Assing" authorityYear="2006" box="[785,940,1497,1517]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="jaenica" status="sp. nov.">
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det.
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. Assing 2006 (cAss).
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<typeStatus id="896C8836C33476163003FA35FE39720C" box="[340,432,1519,1539]" pageId="3" type="paratype">Paratypes</typeStatus>
: 733,
<specimenCount id="40D1FD1DC33476163354FA35FDC97207" box="[515,576,1519,1544]" count="13" pageId="3" type="female">13♀♀</specimenCount>
: same data as holotype (cAss,
<collectionCode id="30C6AE51C334761632D3FA35FC4C720C" box="[900,965,1519,1539]" pageId="3">OÖLL</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="E6BF3CC9C3347616328BFA35FE1B723E" collectingDate="2003-12-26" collectionCode="V" collectorName="E Jaen &amp; Assing" elevation="1650" latitude="37.75" location="Sierra Almaden" longLatPrecision="1179" longitude="-3.55" municipality="No." pageId="3" specimenCount="6" specimenCount-female="6" stateProvince="Andalucia" typeStatus="paratype">
1033,
<specimenCount id="40D1FD1DC3347616357CFA35FBD47207" box="[1067,1117,1519,1544]" count="6" pageId="3" type="female">6♀♀</specimenCount>
: E -
<collectingMunicipality id="B60CACEEC334761635F7FA35FB4B720C" box="[1184,1218,1519,1539]" pageId="3">No.</collectingMunicipality>
8,
<collectingRegion id="9413F876C33476163003F9DCFE3D7215" box="[340,436,1542,1562]" country="Spain" name="Andalucia" pageId="3">Andalucía</collectingRegion>
,
<collectorName id="FB225342C33476163095F9DCFD8F7215" box="[450,518,1542,1562]" pageId="3">E Jaén</collectorName>
,
<location id="5308604FC33476163343F9DCFD257215" LSID="urn:lsid:plazi:treatment:DE7E8782C33476133003FA41FCBB7575:5308604FC33476163343F9DCFD257215" box="[532,684,1542,1562]" latitude="37.75" longLatPrecision="1179" longitude="-3.55" municipality="No." name="Sierra Almaden" pageId="3" stateProvince="Andalucia">Sierra Almadén</location>
,
<quantity id="912F9B71C334761633EDF9DCFCB27215" box="[698,827,1542,1562]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.85" metricValueMin="1.45" pageId="3" unit="m" value="1650.0" valueMax="1850.0" valueMin="1450.0">
<elevation id="DDFAD1A7C334761633EDF9DCFCB27215" box="[698,827,1542,1562]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.85" metricValueMin="1.45" pageId="3" unit="m" value="1650.0" valueMax="1850.0" valueMin="1450.0">1450-1850 m</elevation>
</quantity>
,
<geoCoordinate id="33E35053C3347616321DF9DCFC1A7215" box="[842,915,1542,1562]" degrees="37" direction="north" minutes="45" orientation="latitude" pageId="3" precision="925" value="37.75">37°45N</geoCoordinate>
,
<geoCoordinate id="33E35053C334761632F5F9DCFC797215" box="[930,1008,1542,1562]" degrees="03" direction="west" minutes="33" orientation="longitude" pageId="3" precision="925" value="-3.55">03°33W</geoCoordinate>
,
<date id="22691054C334761632A8F9DCFBF97215" box="[1023,1136,1542,1562]" pageId="3" value="2003-12-26">
<collectingDate id="322DE9BCC334761632A8F9DCFBF97215" box="[1023,1136,1542,1562]" pageId="3" value="2003-12-26">26.XII.2003</collectingDate>
</date>
,
<collectionCode id="30C6AE51C33476163528F9DCFB1B7215" box="[1151,1170,1542,1562]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" type="Museum">V</collectionCode>
.
<collectorName id="FB225342C334761635F7F9DCFB567215" box="[1184,1247,1542,1562]" pageId="3">Assing</collectorName>
(cAss)
</materialsCitation>
;
<materialsCitation id="E6BF3CC9C334761630CAF9C7FDF97247" collectingDate="2003-12-26" collectionCode="V" collectorName="E Jaen &amp; Assing" elevation="1300" latitude="37.75" location="Sierra Almaden" longLatPrecision="1179" longitude="-3.5833333" municipality="No." pageId="3" specimenCount="2" specimenCount-female="2" stateProvince="Andalucia" typeStatus="paratype">
13,
<specimenCount id="40D1FD1DC3347616309EF9C7FE737239" box="[457,506,1565,1590]" count="2" pageId="3" type="female">2♀♀</specimenCount>
: E -
<collectingMunicipality id="B60CACEEC3347616337AF9C7FDD9723E" box="[557,592,1565,1585]" pageId="3">No.</collectingMunicipality>
9,
<collectingRegion id="9413F876C33476163338F9C7FD46723E" box="[623,719,1565,1585]" country="Spain" name="Andalucia" pageId="3">Andalucía</collectingRegion>
,
<collectorName id="FB225342C3347616338DF9C7FC93723E" box="[730,794,1565,1585]" pageId="3">E Jaén</collectorName>
,
<location id="5308604FC33476163272F9C7FC30723E" LSID="urn:lsid:plazi:treatment:DE7E8782C33476133003FA41FCBB7575:5308604FC33476163272F9C7FC30723E" box="[805,953,1565,1585]" latitude="37.75" longLatPrecision="1179" longitude="-3.5833333" municipality="No." name="Sierra Almaden" pageId="3" stateProvince="Andalucia">Sierra Almadén</location>
,
<quantity id="912F9B71C33476163293F9C7FB84723E" box="[964,1037,1565,1585]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="3" unit="m" value="1300.0">
<elevation id="DDFAD1A7C33476163293F9C7FB84723E" box="[964,1037,1565,1585]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="3" unit="m" value="1300.0">1300 m</elevation>
</quantity>
, ca.
<geoCoordinate id="33E35053C3347616356EF9C7FB0B723E" box="[1081,1154,1565,1585]" degrees="37" direction="north" minutes="45" orientation="latitude" pageId="3" precision="925" value="37.75">37°45N</geoCoordinate>
,
<geoCoordinate id="33E35053C334761635D9F9C7FB55723E" box="[1166,1244,1565,1585]" degrees="03" direction="west" minutes="35" orientation="longitude" pageId="3" precision="925" value="-3.5833333">03°35W</geoCoordinate>
,
<date id="22691054C33476163003F9EEFE4F7247" box="[340,454,1588,1608]" pageId="3" value="2003-12-26">
<collectingDate id="322DE9BCC33476163003F9EEFE4F7247" box="[340,454,1588,1608]" pageId="3" value="2003-12-26">26.XII.2003</collectingDate>
</date>
,
<collectionCode id="30C6AE51C33476163098F9EEFE6B7247" box="[463,482,1588,1608]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" type="Museum">V</collectionCode>
.
<collectorName id="FB225342C334761630BBF9EEFDA27247" box="[492,555,1588,1608]" pageId="3">Assing</collectorName>
(cAss)
</materialsCitation>
;
<materialsCitation id="E6BF3CC9C3347616332DF9EEFC6B7251" collectingDate="2003-12-29" collectionCode="V" collectorName="SE Jaen &amp; Sierra de la Pandera &amp; Assing" elevation="1500" latitude="37.666668" location="Puerto Alto" longLatPrecision="1180" longitude="-3.6666667" municipality="No." pageId="3" specimenCount="2" specimenCount-female="2" stateProvince="Andalucia" typeStatus="paratype">
233,
<specimenCount id="40D1FD1DC334761633EFF9EEFD607242" box="[696,745,1588,1613]" count="2" pageId="3" type="female">2♀♀</specimenCount>
: E -
<collectingMunicipality id="B60CACEEC3347616324EF9EEFCB27247" box="[793,827,1588,1608]" pageId="3">No.</collectingMunicipality>
16,
<collectingRegion id="9413F876C33476163233F9EEFC4D7247" box="[868,964,1588,1608]" country="Spain" name="Andalucia" pageId="3">Andalucía</collectingRegion>
,
<collectorName id="FB225342C33476163299F9EEFB937247" box="[974,1050,1588,1608]" pageId="3">SE Jaén</collectorName>
,
<quantity id="912F9B71C33476163573F9EEFBE57247" box="[1060,1132,1588,1608]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="3" unit="m" value="1500.0">
<elevation id="DDFAD1A7C33476163573F9EEFBE57247" box="[1060,1132,1588,1608]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="3" unit="m" value="1500.0">1500 m</elevation>
</quantity>
,
<collectorName id="FB225342C33476163521F9EEFE297251" pageId="3">Sierra de la Pandera</collectorName>
,
<location id="5308604FC334761630FEF990FD9C7251" LSID="urn:lsid:plazi:treatment:DE7E8782C33476133003FA41FCBB7575:5308604FC334761630FEF990FD9C7251" box="[425,533,1610,1630]" latitude="37.666668" longLatPrecision="1180" longitude="-3.6666667" municipality="No." name="Puerto Alto" pageId="3" stateProvince="Andalucia">Puerto Alto</location>
,
<geoCoordinate id="33E35053C33476163349F990FDE17251" box="[542,616,1610,1630]" degrees="37" direction="north" minutes="40" orientation="latitude" pageId="3" precision="925" value="37.666668">37°40N</geoCoordinate>
,
<geoCoordinate id="33E35053C33476163326F990FD377251" box="[625,702,1610,1630]" degrees="03" direction="west" minutes="40" orientation="longitude" pageId="3" precision="925" value="-3.6666667">03°40W</geoCoordinate>
,
<date id="22691054C3347616339FF990FCB37251" box="[712,826,1610,1630]" pageId="3" value="2003-12-29">
<collectingDate id="322DE9BCC3347616339FF990FCB37251" box="[712,826,1610,1630]" pageId="3" value="2003-12-29">29.XII.2003</collectingDate>
</date>
,
<collectionCode id="30C6AE51C33476163214F990FCDF7251" box="[835,854,1610,1630]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" type="Museum">V</collectionCode>
.
<collectorName id="FB225342C33476163208F990FC177251" box="[863,926,1610,1630]" pageId="3">Assing</collectorName>
(cAss)
</materialsCitation>
;
<materialsCitation id="E6BF3CC9C334761632BBF990FCC4727A" collectingDate="2005-04-16" collectorName="M. Baena" county="Jaen" location="Pico Almaden" municipality="Mancha Real" pageId="3" specimenCount="1" specimenCount-female="1" stateProvince="Andalucia" typeStatus="paratype">
13,
<specimenCount id="40D1FD1DC33476163541F990FBBD726C" box="[1046,1076,1610,1635]" count="1" pageId="3" type="female">1♀</specimenCount>
: E -
<collectingCounty id="BF094E18C33476163535F990FB047251" box="[1122,1165,1610,1630]" pageId="3">Jaén</collectingCounty>
,
<collectingMunicipality id="B60CACEEC334761635C1F990FE09727A" pageId="3">Mancha Real</collectingMunicipality>
,
<location id="5308604FC334761630DEF9BBFD87727A" LSID="urn:lsid:plazi:treatment:DE7E8782C33476133003FA41FCBB7575:5308604FC334761630DEF9BBFD87727A" box="[393,526,1633,1653]" county="Jaen" municipality="Mancha Real" name="Pico Almaden" pageId="3" stateProvince="Andalucia">Pico Almadén</location>
,
<date id="22691054C33476163341F9BBFDF6727A" box="[534,639,1633,1653]" pageId="3" value="2005-04-16">
<collectingDate id="322DE9BCC33476163341F9BBFDF6727A" box="[534,639,1633,1653]" pageId="3" value="2005-04-16">16.IV.2005</collectingDate>
</date>
, leg.
<collectorName id="FB225342C334761633E7F9BBFC80727A" box="[688,777,1633,1653]" pageId="3">M. Baena</collectorName>
(cAss)
</materialsCitation>
.
</paragraph>
<paragraph id="56683694C33476163003F959FC3772DF" blockId="3.[340,1248,1666,1744]" pageId="3">
D e s c r i p t i o n: Small species,
<quantity id="912F9B71C3347616338AF958FCD37297" box="[733,858,1666,1688]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" metricValueMax="2.5" metricValueMin="1.9" pageId="3" unit="mm" value="2.2" valueMax="2.5" valueMin="1.9">1.9-2.5 mm</quantity>
; facies as in
<figureCitation id="CEEC2A11C334761632B9F958FBB97297" box="[1006,1072,1666,1688]" captionStart="Figs 8-21" captionStartId="5.[340,380,1392,1412]" captionTargetBox="[343,1247,434,1370]" captionTargetId="figure-194@5.[340,1248,434,910]" captionTargetPageId="5" captionText="Figs 8-21: Geostiba jaenica nov.sp.: (8) habitus; (9) forebody; (10) head in lateral view; (11) antenna; (12) 3 tergite VIII; (13) posterior margin of 3 tergite VIII; (14) 3 sternite VIII; (15-17) median lobe of aedeagus in lateral and in ventral view; (18) apical lobe of paramere; (19) ♀ sternite VIII; (20) posterior margin of ♀ sternite VIII; (21) spermatheca. Scales: 8-9: 0.5 mm; 10-21: 0.1 mm." pageId="3">Fig. 8</figureCitation>
. Coloration uniformly yellowish to reddish yellow, usually with the middle of abdominal tergite
<collectionCode id="30C6AE51C334761635CEF945FB3D72BB" box="[1177,1204,1695,1716]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="3">VI</collectionCode>
and sometimes also the middle of tergite
<collectionCode id="30C6AE51C3347616339EF961FD5272DF" box="[713,731,1723,1744]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" type="Museum">V</collectionCode>
indistinctly darkened.
</paragraph>
<paragraph id="56683694C33476163003F904FB69731F" blockId="3.[340,1248,1758,1808]" pageId="3">
Head approximately as wide as long to weakly oblong; eyes very small, composed of only approximately 5 ommatidia (
<figureCitation id="CEEC2A11C334761633E5F920FD76731F" box="[690,767,1786,1808]" captionStart="Figs 8-21" captionStartId="5.[340,380,1392,1412]" captionTargetBox="[343,1247,434,1370]" captionTargetId="figure-194@5.[340,1248,434,910]" captionTargetPageId="5" captionText="Figs 8-21: Geostiba jaenica nov.sp.: (8) habitus; (9) forebody; (10) head in lateral view; (11) antenna; (12) 3 tergite VIII; (13) posterior margin of 3 tergite VIII; (14) 3 sternite VIII; (15-17) median lobe of aedeagus in lateral and in ventral view; (18) apical lobe of paramere; (19) ♀ sternite VIII; (20) posterior margin of ♀ sternite VIII; (21) spermatheca. Scales: 8-9: 0.5 mm; 10-21: 0.1 mm." pageId="3">Fig. 10</figureCitation>
), in dorsal view 1/6-1/5 the length of postocu-
</paragraph>
<paragraph id="56683694C33376113255FEBFFCBB7575" blockId="4.[770,818,357,378]" box="[770,818,357,378]" pageId="4">1133</paragraph>
<paragraph id="56683694C33376113003FE74FD447617" blockId="4.[340,1248,430,536]" pageId="4">
lar region, not projecting from lateral outline of head (
<figureCitation id="CEEC2A11C333761132F4FE74FC6F75CB" box="[931,998,430,452]" captionStart="Figs 8-21" captionStartId="5.[340,380,1392,1412]" captionTargetBox="[343,1247,434,1370]" captionTargetId="figure-194@5.[340,1248,434,910]" captionTargetPageId="5" captionText="Figs 8-21: Geostiba jaenica nov.sp.: (8) habitus; (9) forebody; (10) head in lateral view; (11) antenna; (12) 3 tergite VIII; (13) posterior margin of 3 tergite VIII; (14) 3 sternite VIII; (15-17) median lobe of aedeagus in lateral and in ventral view; (18) apical lobe of paramere; (19) ♀ sternite VIII; (20) posterior margin of ♀ sternite VIII; (21) spermatheca. Scales: 8-9: 0.5 mm; 10-21: 0.1 mm." pageId="4">Fig. 9</figureCitation>
); integument with very shallow microreticulation and some shine; puncturation extremely fine and sparse, barely noticeable. Antenna with antennomere IV strongly transverse, IV-X of increasing width and increasingly transverse (
<figureCitation id="CEEC2A11C33376113321FDD8FD497617" box="[630,704,514,536]" captionStart="Figs 8-21" captionStartId="5.[340,380,1392,1412]" captionTargetBox="[343,1247,434,1370]" captionTargetId="figure-194@5.[340,1248,434,910]" captionTargetPageId="5" captionText="Figs 8-21: Geostiba jaenica nov.sp.: (8) habitus; (9) forebody; (10) head in lateral view; (11) antenna; (12) 3 tergite VIII; (13) posterior margin of 3 tergite VIII; (14) 3 sternite VIII; (15-17) median lobe of aedeagus in lateral and in ventral view; (18) apical lobe of paramere; (19) ♀ sternite VIII; (20) posterior margin of ♀ sternite VIII; (21) spermatheca. Scales: 8-9: 0.5 mm; 10-21: 0.1 mm." pageId="4">Fig. 11</figureCitation>
).
</paragraph>
<paragraph id="56683694C33376113003FDFDFB337657" blockId="4.[340,1247,550,600]" pageId="4">
Pronotum 1.05-1.10 times as wide as head and 1.05-1.10 times as wide as long; maximal width in anterior half; puncturation and microreticulation similar to those of head (
<figureCitation id="CEEC2A11C33376113526FD98FB247657" box="[1137,1197,578,600]" captionStart="Figs 8-21" captionStartId="5.[340,380,1392,1412]" captionTargetBox="[343,1247,434,1370]" captionTargetId="figure-194@5.[340,1248,434,910]" captionTargetPageId="5" captionText="Figs 8-21: Geostiba jaenica nov.sp.: (8) habitus; (9) forebody; (10) head in lateral view; (11) antenna; (12) 3 tergite VIII; (13) posterior margin of 3 tergite VIII; (14) 3 sternite VIII; (15-17) median lobe of aedeagus in lateral and in ventral view; (18) apical lobe of paramere; (19) ♀ sternite VIII; (20) posterior margin of ♀ sternite VIII; (21) spermatheca. Scales: 8-9: 0.5 mm; 10-21: 0.1 mm." pageId="4">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="56683694C33376113003FDBCFC4576BB" blockId="4.[340,1248,614,692]" pageId="4">
Elytra without sexual dimorphism, at suture 0.55-0.60 times as long as pronotum; microsculpture more distinct than that of pronotum; puncturation extremely fine, barely noticeable in the microsculpture (
<figureCitation id="CEEC2A11C333761133F0FD44FD6F76BB" box="[679,742,670,692]" captionStart="Figs 8-21" captionStartId="5.[340,380,1392,1412]" captionTargetBox="[343,1247,434,1370]" captionTargetId="figure-194@5.[340,1248,434,910]" captionTargetPageId="5" captionText="Figs 8-21: Geostiba jaenica nov.sp.: (8) habitus; (9) forebody; (10) head in lateral view; (11) antenna; (12) 3 tergite VIII; (13) posterior margin of 3 tergite VIII; (14) 3 sternite VIII; (15-17) median lobe of aedeagus in lateral and in ventral view; (18) apical lobe of paramere; (19) ♀ sternite VIII; (20) posterior margin of ♀ sternite VIII; (21) spermatheca. Scales: 8-9: 0.5 mm; 10-21: 0.1 mm." pageId="4">Fig. 9</figureCitation>
). Hind wings reduced.
</paragraph>
<paragraph id="56683694C33376113003FD18FE507747" blockId="4.[340,1248,706,840]" pageId="4">
Abdomen slightly (approximately 1.1 ✕) wider than elytra, widest at segments
<collectionCode id="30C6AE51C333761135F2FD18FB3176D7" box="[1189,1208,706,728]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" type="Museum">V</collectionCode>
/
<collectionCode id="30C6AE51C33376113597FD18FB5576D7" box="[1216,1244,706,728]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="4">VI</collectionCode>
; puncturation sparse and very fine, barely noticeable; microsculpture shallow; posterior margin of tergite VII without palisade fringe; tergite VII and VIII without sexual dimorphism; posterior margin of tergite VIII in the middle more or less distinctly concave (
<figureCitation id="CEEC2A11C3337611300CFCE8FE457747" box="[347,460,818,840]" captionStart="Figs 8-21" captionStartId="5.[340,380,1392,1412]" captionTargetBox="[343,1247,434,1370]" captionTargetId="figure-194@5.[340,1248,434,910]" captionTargetPageId="5" captionText="Figs 8-21: Geostiba jaenica nov.sp.: (8) habitus; (9) forebody; (10) head in lateral view; (11) antenna; (12) 3 tergite VIII; (13) posterior margin of 3 tergite VIII; (14) 3 sternite VIII; (15-17) median lobe of aedeagus in lateral and in ventral view; (18) apical lobe of paramere; (19) ♀ sternite VIII; (20) posterior margin of ♀ sternite VIII; (21) spermatheca. Scales: 8-9: 0.5 mm; 10-21: 0.1 mm." pageId="4">Figs 12-13</figureCitation>
).
</paragraph>
<paragraph id="56683694C33376113003FC82FD5A7784" blockId="4.[340,1247,856,907]" pageId="4">
3: posterior margin of sternite VIII convex (
<figureCitation id="CEEC2A11C33376113273FC82FCF97761" box="[804,880,856,878]" captionStart="Figs 8-21" captionStartId="5.[340,380,1392,1412]" captionTargetBox="[343,1247,434,1370]" captionTargetId="figure-194@5.[340,1248,434,910]" captionTargetPageId="5" captionText="Figs 8-21: Geostiba jaenica nov.sp.: (8) habitus; (9) forebody; (10) head in lateral view; (11) antenna; (12) 3 tergite VIII; (13) posterior margin of 3 tergite VIII; (14) 3 sternite VIII; (15-17) median lobe of aedeagus in lateral and in ventral view; (18) apical lobe of paramere; (19) ♀ sternite VIII; (20) posterior margin of ♀ sternite VIII; (21) spermatheca. Scales: 8-9: 0.5 mm; 10-21: 0.1 mm." pageId="4">Fig. 14</figureCitation>
); median lobe of aedeagus and apical lobe of paramere as in
<figureCitation id="CEEC2A11C33376113308FCAFFD467784" box="[607,719,885,907]" captionStart="Figs 8-21" captionStartId="5.[340,380,1392,1412]" captionTargetBox="[343,1247,434,1370]" captionTargetId="figure-194@5.[340,1248,434,910]" captionTargetPageId="5" captionText="Figs 8-21: Geostiba jaenica nov.sp.: (8) habitus; (9) forebody; (10) head in lateral view; (11) antenna; (12) 3 tergite VIII; (13) posterior margin of 3 tergite VIII; (14) 3 sternite VIII; (15-17) median lobe of aedeagus in lateral and in ventral view; (18) apical lobe of paramere; (19) ♀ sternite VIII; (20) posterior margin of ♀ sternite VIII; (21) spermatheca. Scales: 8-9: 0.5 mm; 10-21: 0.1 mm." pageId="4">Figs 15-18</figureCitation>
.
</paragraph>
<paragraph id="56683694C33376113003FC7AFC0877C2" blockId="4.[340,1247,922,973]" pageId="4">
<specimenCount id="40D1FD1DC33376113003FC7AFEE077BA" box="[340,361,928,949]" count="1" pageId="4" type="female">♀</specimenCount>
: posterior margin of sternite VIII in the middle distinctly concave and with long stout marginal setae (
<figureCitation id="CEEC2A11C333761130A3FC6DFDEF77C2" box="[500,614,951,973]" captionStart="Figs 8-21" captionStartId="5.[340,380,1392,1412]" captionTargetBox="[343,1247,434,1370]" captionTargetId="figure-194@5.[340,1248,434,910]" captionTargetPageId="5" captionText="Figs 8-21: Geostiba jaenica nov.sp.: (8) habitus; (9) forebody; (10) head in lateral view; (11) antenna; (12) 3 tergite VIII; (13) posterior margin of 3 tergite VIII; (14) 3 sternite VIII; (15-17) median lobe of aedeagus in lateral and in ventral view; (18) apical lobe of paramere; (19) ♀ sternite VIII; (20) posterior margin of ♀ sternite VIII; (21) spermatheca. Scales: 8-9: 0.5 mm; 10-21: 0.1 mm." pageId="4">Figs 19-20</figureCitation>
); spermatheca as in
<figureCitation id="CEEC2A11C33376113265FC6DFC0877C2" box="[818,897,951,973]" captionStart="Figs 8-21" captionStartId="5.[340,380,1392,1412]" captionTargetBox="[343,1247,434,1370]" captionTargetId="figure-194@5.[340,1248,434,910]" captionTargetPageId="5" captionText="Figs 8-21: Geostiba jaenica nov.sp.: (8) habitus; (9) forebody; (10) head in lateral view; (11) antenna; (12) 3 tergite VIII; (13) posterior margin of 3 tergite VIII; (14) 3 sternite VIII; (15-17) median lobe of aedeagus in lateral and in ventral view; (18) apical lobe of paramere; (19) ♀ sternite VIII; (20) posterior margin of ♀ sternite VIII; (21) spermatheca. Scales: 8-9: 0.5 mm; 10-21: 0.1 mm." pageId="4">Fig. 21.</figureCitation>
</paragraph>
<paragraph id="56683694C33376113003FC07FE7E701F" blockId="4.[340,1248,989,1040]" pageId="4">E t y m o l o g y: The name (adj.) is derived from Jaén, the province where the species was discovered.</paragraph>
<paragraph id="56683694C33376113003FBC4FB5470AB" blockId="4.[340,1248,1054,1188]" pageId="4">
C o m p a r a t i v e n o t e s a n d s y s t e m a t i c s: Basedontheabsenceofa sexual dimorphism of the pronotum, elytra, and of the abdominal tergites VII and VIII, as well as on the morphology of the primary sexual characters,
<taxonomicName id="91D74D17C3337611328CFB8DFBCD7063" authorityName="Assing" authorityYear="2006" box="[987,1092,1111,1132]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="jaenica">
<emphasis id="64A3EA86C3337611328CFB8DFBCD7063" box="[987,1092,1111,1132]" italics="true" pageId="4">G. jaenica</emphasis>
</taxonomicName>
is attributed to the subgenus
<taxonomicName id="91D74D17C333761130B5FBA8FCBE7087" authority="SCHEERPELTZ. From" authorityName="SCHEERPELTZ. From" box="[482,823,1138,1160]" pageId="4" rank="subGenus" subGenus="Sipalotricha">
<emphasis id="64A3EA86C333761130B5FBA8FDD67087" box="[482,607,1138,1160]" italics="true" pageId="4">Sipalotricha</emphasis>
SCHEERPELTZ. From
</taxonomicName>
other species of this subgenus occurring in southern
<collectingCountry id="2EC07604C3337611309BFB54FD8170AB" box="[460,520,1166,1188]" name="Spain" pageId="4">Spain</collectingCountry>
, it is distinguished by the sexual characters and additionally as follows:
</paragraph>
<paragraph id="56683694C33376113003FB68FB2370EB" blockId="4.[340,1248,1202,1252]" pageId="4">
from the widespread
<taxonomicName id="91D74D17C33376113366FB69FC9570C7" authority="(FAUVEL)" baseAuthorityName="FAUVEL" box="[561,796,1202,1224]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="plicatella">
<emphasis id="64A3EA86C33376113366FB69FD3B70C7" box="[561,690,1202,1224]" italics="true" pageId="4">G. plicatella</emphasis>
(FAUVEL)
</taxonomicName>
by paler coloration, smaller size, relatively shorter elytra, much smaller eyes, and more strongly transverse antennomeres IV-X;
</paragraph>
<paragraph id="56683694C33376113003FB28FBA8712B" blockId="4.[340,1247,1266,1316]" pageId="4">
from the similarly small and microphthalmous
<taxonomicName id="91D74D17C3337611326EFB29FBAE7107" authority="(FAGEL)" baseAuthorityName="FAGEL" box="[825,1063,1266,1288]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="cazorlensis">
<emphasis id="64A3EA86C3337611326EFB29FC457107" box="[825,972,1266,1288]" italics="true" pageId="4">G. cazorlensis</emphasis>
(FAGEL)
</taxonomicName>
by the infuscated preapical abdominal segments and by the indistinctly punctured elytra;
</paragraph>
<paragraph id="56683694C33376113003FAE8FE1C716B" blockId="4.[340,1247,1330,1380]" pageId="4">
from
<taxonomicName id="91D74D17C333761130D9FAE9FDE47147" authority="(FAGEL)" baseAuthorityName="FAGEL" box="[398,621,1330,1352]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="besucheti">
<emphasis id="64A3EA86C333761130D9FAE9FD867147" box="[398,527,1330,1352]" italics="true" pageId="4">G. besucheti</emphasis>
(FAGEL)
</taxonomicName>
by much paler coloration and by the indistinctly punctured elytra;
</paragraph>
</subSubSection>
<subSubSection id="1ECD651FC33376133003FAA8FCBB7575" lastPageId="6" pageId="4" type="description">
<paragraph id="56683694C33376113003FAA8FD6971AB" blockId="4.[340,1248,1394,1444]" pageId="4">
from
<taxonomicName id="91D74D17C333761130C7FAA9FDD87187" authority="PACE" authorityName="PACE" box="[400,593,1394,1416]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="tarifensis">
<emphasis id="64A3EA86C333761130C7FAA9FD987187" box="[400,529,1394,1416]" italics="true" pageId="4">G. tarifensis</emphasis>
PACE
</taxonomicName>
by smaller size, paler coloration, smaller eyes, and the more strongly transverse antennomere IV-X;
</paragraph>
<paragraph id="56683694C33376113003FA68FBB871EB" blockId="4.[340,1248,1458,1508]" pageId="4">
from
<taxonomicName id="91D74D17C333761130DBFA69FD0F71C7" authority="PACE" authorityName="PACE" box="[396,646,1459,1480]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="jarasteparensis">
<emphasis id="64A3EA86C333761130DBFA69FDC171C7" box="[396,584,1459,1480]" italics="true" pageId="4">G. jarasteparensis</emphasis>
PACE
</taxonomicName>
by the infuscated preapical abdominal segments, the more transverse antennomeres IV-X, and the less distinct elytral puncturation;
</paragraph>
<paragraph id="56683694C33376113003FA28FCCC722B" blockId="4.[340,1248,1522,1572]" pageId="4">
from
<taxonomicName id="91D74D17C333761130C6FA29FDEB7207" authority="(FAGEL)" baseAuthorityName="FAGEL" box="[401,610,1522,1544]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="fretoria">
<emphasis id="64A3EA86C333761130C6FA29FD8B7207" box="[401,514,1522,1544]" italics="true" pageId="4">G. fretoria</emphasis>
(FAGEL)
</taxonomicName>
by the infuscated preapical abdominal segments, the more transverse antennomeres IV-X, and smaller eyes;
</paragraph>
<paragraph id="56683694C33376113003F9E9FC297269" blockId="4.[340,1247,1587,1638]" pageId="4">
and from
<taxonomicName id="91D74D17C333761130ECF9EEFD1D7246" authority="PACE" authorityName="PACE" box="[443,660,1588,1609]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="araneaensis">
<emphasis id="64A3EA86C333761130ECF9EEFDDF7246" box="[443,598,1588,1609]" italics="true" pageId="4">G. araneaensis</emphasis>
PACE
</taxonomicName>
by less pronounced microsculpture on the forebody and by less distinctly infuscate preapical abdominal segments.
</paragraph>
<paragraph id="56683694C33276103255FEBFFCBB7575" blockId="5.[770,818,357,378]" box="[770,818,357,378]" pageId="5">1134</paragraph>
<caption id="02A8661CC33276103003FAAAFEF771D0" pageId="5" startId="5.[340,380,1392,1412]" targetBox="[343,1247,434,1370]" targetPageId="5" targetType="figure">
<paragraph id="56683694C33276103003FAAAFEF771D0" blockId="5.[340,1248,1392,1503]" pageId="5">
<emphasis id="64A3EA86C33276103003FAAAFE39718B" bold="true" box="[340,432,1392,1412]" pageId="5">Figs 8-21</emphasis>
:
<taxonomicName id="91D74D17C33276103097FAAAFDD6718B" authorityName="Assing" authorityYear="2006" box="[448,607,1392,1412]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="jaenica" status="sp. nov.">
<emphasis id="64A3EA86C33276103097FAAAFDD6718B" box="[448,607,1392,1412]" italics="true" pageId="5">Geostiba jaenica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7F9057FDC3327610333EFAAAFD26718B" box="[617,687,1392,1412]" pageId="5" rank="species">nov.sp.</taxonomicNameLabel>
: (
<emphasis id="64A3EA86C33276103394FAAAFD47718B" bold="true" box="[707,718,1392,1412]" pageId="5">8</emphasis>
) habitus; (
<emphasis id="64A3EA86C3327610326DFAAAFCCC718B" bold="true" box="[826,837,1392,1412]" pageId="5">9</emphasis>
) forebody; (
<emphasis id="64A3EA86C33276103296FAAAFC51718B" bold="true" box="[961,984,1392,1412]" pageId="5">10</emphasis>
) head in lateral view; (
<emphasis id="64A3EA86C33276103596FAAAFB51718B" bold="true" box="[1217,1240,1392,1412]" pageId="5">11</emphasis>
) antenna; (
<emphasis id="64A3EA86C332761030F8FA5DFE4F7194" bold="true" box="[431,454,1415,1435]" pageId="5">12</emphasis>
) 3 tergite VIII; (
<emphasis id="64A3EA86C3327610333CFA5DFD0B7194" bold="true" box="[619,642,1415,1435]" pageId="5">13</emphasis>
) posterior margin of 3 tergite VIII; (
<emphasis id="64A3EA86C33276103289FA5DFC7C7194" bold="true" box="[990,1013,1415,1435]" pageId="5">14</emphasis>
) 3 sternite VIII; (
<emphasis id="64A3EA86C332761035F4FA5DFB5E7194" bold="true" box="[1187,1239,1415,1435]" pageId="5">15-17</emphasis>
) median lobe of aedeagus in lateral and in ventral view; (
<emphasis id="64A3EA86C33276103204FA44FCE371BD" bold="true" box="[851,874,1438,1458]" pageId="5">18</emphasis>
) apical lobe of paramere; (
<emphasis id="64A3EA86C33276103537FA44FBFE71BD" bold="true" box="[1120,1143,1438,1458]" pageId="5">19</emphasis>
) ♀ sternite VIII; (
<emphasis id="64A3EA86C332761030C7FA6FFE2E71C6" bold="true" box="[400,423,1461,1481]" pageId="5">20</emphasis>
) posterior margin of ♀ sternite VIII; (
<emphasis id="64A3EA86C33276103245FA6FFCA071C6" bold="true" box="[786,809,1461,1481]" pageId="5">21</emphasis>
) spermatheca. Scales: 8-9: 0.5 mm; 10-21: 0.1 mm.
</paragraph>
</caption>
<paragraph id="56683694C33276103003F9CEFD3472B4" blockId="5.[340,1248,1555,1723]" pageId="5">
D i s t r i b u t i o n a n d b i o n o m i c s: The absence of records from other localities, as well as the adaptive reductions of the eyes, pigmentation, and wings suggest that
<taxonomicName id="91D74D17C332761030D4F997FDBB726C" authorityName="Assing" authorityYear="2006" box="[387,562,1613,1635]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="jaenica">
<emphasis id="64A3EA86C332761030D4F997FDBB726C" box="[387,562,1613,1635]" italics="true" pageId="5">Geostiba jaenica</emphasis>
</taxonomicName>
has a restricted distribution; it seems very likely that it is an endemic of the Sierra Almadén and the Sierra de la Pandera near Jaén (
<figureCitation id="CEEC2A11C33276103570F9B1FBE5728E" box="[1063,1132,1643,1665]" captionStart="Map 1" captionStartId="3.[340,385,1351,1371]" captionTargetBox="[340,1247,707,1340]" captionTargetId="figure-370@3.[340,1247,707,1340]" captionTargetPageId="3" captionText="Map 1: Distributions of Phloeocharis baenai nov.sp. (open circle), Geostiba jaenica nov.sp. (filled circles), and Atheta tenebrarum nov.sp. (square) in southern Spain." pageId="5">Map 1</figureCitation>
). The
<typeStatus id="896C8836C332761035E3F9B6FB69728E" box="[1204,1248,1644,1665]" pageId="5">type</typeStatus>
specimens were collected by sifting grass roots, moss, and leaf litter in
<emphasis id="64A3EA86C3327610350CF953FB567291" box="[1115,1247,1672,1694]" italics="true" pageId="5">Quercus ilex</emphasis>
forests at altitudes of
<quantity id="912F9B71C33276103379F97FFD3172B4" box="[558,696,1701,1723]" metricMagnitude="3" metricUnit="m" metricValue="1.575" metricValueMax="1.85" metricValueMin="1.3" pageId="5" unit="m" value="1575.0" valueMax="1850.0" valueMin="1300.0">1300-1850 m</quantity>
.
</paragraph>
<paragraph id="56683694C33176133255FEBFFCBB7575" blockId="6.[770,818,357,378]" box="[770,818,357,378]" pageId="6">1135</paragraph>
</subSubSection>
</treatment>
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270
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<taxonomicName id="91D74D17C33676143003FCE8FDA37747" ID-CoL="4GG9V" authorityName="Assing" authorityYear="2006" box="[340,554,818,840]" class="Insecta" family="Staphylinidae" genus="Phloeocharis" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="baenai" status="sp. nov.">
<emphasis id="64A3EA86C33676143003FCE8FDA37747" bold="true" box="[340,554,818,840]" italics="true" pageId="1">Phloeocharis baenai</emphasis>
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(
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;
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)
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<typeStatus id="896C8836C33676143003FC83FE207762" box="[340,425,857,877]" pageId="1" type="holotype">Holotype</typeStatus>
3: E -
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,
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,
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, leg.
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</materialsCitation>
/
<typeStatus id="896C8836C33676143537FC83FB367762" box="[1120,1215,857,877]" pageId="1" type="holotype">Holotypus</typeStatus>
3
<taxonomicName id="91D74D17C33676143003FCAAFD98778B" authorityName="Assing" authorityYear="2006" box="[340,529,880,900]" class="Insecta" family="Staphylinidae" genus="Phloeocharis" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="baenai" status="sp. nov.">
<emphasis id="64A3EA86C33676143003FCAAFD98778B" box="[340,529,880,900]" italics="true" pageId="1">Phloeocharis baenai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7F9057FDC33676143341FCAAFDCB778B" box="[534,578,880,900]" pageId="1" rank="species">sp.n.</taxonomicNameLabel>
det.
<collectionCode id="30C6AE51C33676143339FCAAFD08778B" box="[622,641,880,900]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" type="Museum">V</collectionCode>
. Assing 2006 (cAss).
</paragraph>
<paragraph id="56683694C33676143003FC49FCC677ED" blockId="1.[340,1249,914,994]" pageId="1">
D e s c r i p t i o n: Measurements (in mm) and ratios: AL: 0.50;
<collectionCode id="30C6AE51C33676143552FC49FBBA77A7" box="[1029,1075,915,936]" pageId="1">HW</collectionCode>
: 0.32;
<collectionCode id="30C6AE51C3367614352FFC49FB2B77A7" box="[1144,1186,915,936]" country="Thailand" name="Paleontological Collections" pageId="1">PW</collectionCode>
: 0.41;
<collectionCode id="30C6AE51C33676143003FC6AFEFC77CA" box="[340,373,944,965]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="1" type="Museum">PL</collectionCode>
: 0.30;
<collectionCode id="30C6AE51C336761430E8FC6AFE6B77CA" box="[447,482,944,965]" pageId="1">EL</collectionCode>
: 0.34; EW: 0.41; AW: 0.40;
<collectionCode id="30C6AE51C33676143240FC6AFCC877CA" box="[791,833,944,965]" country="France" name="Musee de Lectoure" pageId="1">ML</collectionCode>
: 0.38; TL: 2.2;
<collectionCode id="30C6AE51C336761432BEFC75FB9877CA" box="[1001,1041,943,965]" country="Thailand" name="Paleontological Collections" pageId="1">PW</collectionCode>
/
<collectionCode id="30C6AE51C3367614354EFC75FBCC77CA" box="[1049,1093,943,965]" pageId="1">HW</collectionCode>
: 1.26;
<collectionCode id="30C6AE51C336761435D9FC75FB3F77CA" box="[1166,1206,943,965]" country="Thailand" name="Paleontological Collections" pageId="1">PW</collectionCode>
/
<collectionCode id="30C6AE51C336761435E9FC75FB5477CA" box="[1214,1245,943,965]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="1" type="Museum">PL</collectionCode>
: 1.35;
<collectionCode id="30C6AE51C336761430D9FC16FE2777ED" box="[398,430,972,994]" pageId="1">EL</collectionCode>
/
<collectionCode id="30C6AE51C336761430E1FC16FE5C77ED" box="[438,469,972,994]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="1" type="Museum">PL</collectionCode>
: 1.13; EW/
<collectionCode id="30C6AE51C3367614331DFC16FDFB77ED" box="[586,626,972,994]" country="Thailand" name="Paleontological Collections" pageId="1">PW</collectionCode>
: 1.00; AW/EW: 0.98.
</paragraph>
<paragraph id="56683694C33676143003FC28FD3A704D" blockId="1.[340,1247,1010,1090]" pageId="1">
Small species (see measurements); facies as in
<figureCitation id="CEEC2A11C3367614326BFC28FCF37007" box="[828,890,1010,1032]" captionStart="Figs 1-7" captionStartId="2.[340,380,1447,1467]" captionTargetBox="[343,1240,843,1436]" captionTargetId="figure-398@2.[340,1248,962,1436]" captionTargetPageId="2" captionText="Figs 1-7: Phloeocharis baenai nov.sp.: (1) habitus; (2) forebody; (3) head in lateral view; (4) antenna; (5-7) aedeagus in lateral and in ventral view. Scales: 1-2: 0.5 mm; 3-7: 0.1 mm." pageId="1">Fig. 1</figureCitation>
. Coloration: body brown, with the posterior margins of abdominal segments III-VII, and abdominal apex paler; legs light brown; antennae yellowish brown.
</paragraph>
<paragraph id="56683694C33676143003FB88FC1170F5" blockId="1.[340,1248,1106,1274]" pageId="1">
Head with distinct microreticulation and almost matt; puncturation extremely fine and sparse, barely noticeable; eyes not reduced, composed of numerous ommatidia (
<figureCitation id="CEEC2A11C336761435C2FBB5FEEB70AD" captionStart="Figs 1-7" captionStartId="2.[340,380,1447,1467]" captionTargetBox="[343,1240,843,1436]" captionTargetId="figure-398@2.[340,1248,962,1436]" captionTargetPageId="2" captionText="Figs 1-7: Phloeocharis baenai nov.sp.: (1) habitus; (2) forebody; (3) head in lateral view; (4) antenna; (5-7) aedeagus in lateral and in ventral view. Scales: 1-2: 0.5 mm; 3-7: 0.1 mm." pageId="1">Figs 2- 3</figureCitation>
). Antenna with antennomere IV approximately as wide as long,
<collectionCode id="30C6AE51C33676143551FB57FB9170AD" box="[1030,1048,1165,1186]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" type="Museum">V</collectionCode>
weakly transverse, V-IX increasingly transverse and of increasing width, IX twice as wide as long, X distinctly longer and less transverse than IX, and XI weakly oblong, somewhat longer than X, but shorter than the combined length of IX-X (
<figureCitation id="CEEC2A11C3367614321AFB3EFC0270F5" box="[845,907,1252,1274]" captionStart="Figs 1-7" captionStartId="2.[340,380,1447,1467]" captionTargetBox="[343,1240,843,1436]" captionTargetId="figure-398@2.[340,1248,962,1436]" captionTargetPageId="2" captionText="Figs 1-7: Phloeocharis baenai nov.sp.: (1) habitus; (2) forebody; (3) head in lateral view; (4) antenna; (5-7) aedeagus in lateral and in ventral view. Scales: 1-2: 0.5 mm; 3-7: 0.1 mm." pageId="1">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="56683694C33676143003FAD0FC117178" blockId="1.[340,1248,1289,1399]" pageId="1">
Pronotum relatively slender, but wider than head and transverse (see ratios
<collectionCode id="30C6AE51C336761435D6FAD3FB217110" box="[1153,1192,1289,1311]" country="Thailand" name="Paleontological Collections" pageId="1">PW</collectionCode>
/
<collectionCode id="30C6AE51C336761435E6FAD3FB557110" box="[1201,1244,1289,1311]" pageId="1">HW</collectionCode>
,
<collectionCode id="30C6AE51C33676143003FAFCFEF27133" box="[340,379,1318,1340]" country="Thailand" name="Paleontological Collections" pageId="1">PW</collectionCode>
/
<collectionCode id="30C6AE51C336761430D4FAFCFE287133" box="[387,417,1318,1340]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="1" type="Museum">PL</collectionCode>
, and
<figureCitation id="CEEC2A11C33676143080FAFCFD9C7133" box="[471,533,1318,1340]" captionStart="Figs 1-7" captionStartId="2.[340,380,1447,1467]" captionTargetBox="[343,1240,843,1436]" captionTargetId="figure-398@2.[340,1248,962,1436]" captionTargetPageId="2" captionText="Figs 1-7: Phloeocharis baenai nov.sp.: (1) habitus; (2) forebody; (3) head in lateral view; (4) antenna; (5-7) aedeagus in lateral and in ventral view. Scales: 1-2: 0.5 mm; 3-7: 0.1 mm." pageId="1">Fig. 2</figureCitation>
); maximum width about in the middle; lateral margins weakly convex in dorsal view; posterior angles obtuse, but well-marked; posterior margin straight; microsculpture and puncturation similar to those of head.
</paragraph>
<paragraph id="56683694C33676143003FA5CFB0C71FB" blockId="1.[340,1248,1414,1524]" pageId="1">
Elytra about as wide as and at suture longer than pronotum (see ratios
<collectionCode id="30C6AE51C33676143565FA5CFBD37193" box="[1074,1114,1414,1436]" pageId="1">EW</collectionCode>
/
<collectionCode id="30C6AE51C33676143535FA5CFB007193" box="[1122,1161,1414,1436]" country="Thailand" name="Paleontological Collections" pageId="1">PW</collectionCode>
,
<collectionCode id="30C6AE51C336761435C1FA5CFB3F7193" box="[1174,1206,1414,1436]" pageId="1">EL</collectionCode>
/
<collectionCode id="30C6AE51C336761435EAFA5CFB557193" box="[1213,1244,1414,1436]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="1" type="Museum">PL</collectionCode>
, and
<figureCitation id="CEEC2A11C336761430D7FA7EFE3771B5" box="[384,446,1444,1466]" captionStart="Figs 1-7" captionStartId="2.[340,380,1447,1467]" captionTargetBox="[343,1240,843,1436]" captionTargetId="figure-398@2.[340,1248,962,1436]" captionTargetPageId="2" captionText="Figs 1-7: Phloeocharis baenai nov.sp.: (1) habitus; (2) forebody; (3) head in lateral view; (4) antenna; (5-7) aedeagus in lateral and in ventral view. Scales: 1-2: 0.5 mm; 3-7: 0.1 mm." pageId="1">Fig. 2</figureCitation>
); posterior margin weakly convex; microsculpture shallower than that of head and pronotum, surface consequently with more shine; puncturation fine, but more distinct than that of head and pronotum. Hind wings not examined.. Legs not distinctive.
</paragraph>
<paragraph id="56683694C33676143003F9D9FB8E727E" blockId="1.[340,1248,1539,1649]" pageId="1">
Abdomen approximately as wide as combined width of elytra (see ratio
<collectionCode id="30C6AE51C3367614350FF9D9FB0B7216" box="[1112,1154,1539,1561]" pageId="1">AW</collectionCode>
/
<collectionCode id="30C6AE51C336761435DEF9D9FB397216" box="[1161,1200,1539,1561]" pageId="1">EW</collectionCode>
and
<figureCitation id="CEEC2A11C33676143003F9FBFE187238" box="[340,401,1569,1591]" captionStart="Figs 1-7" captionStartId="2.[340,380,1447,1467]" captionTargetBox="[343,1240,843,1436]" captionTargetId="figure-398@2.[340,1248,962,1436]" captionTargetPageId="2" captionText="Figs 1-7: Phloeocharis baenai nov.sp.: (1) habitus; (2) forebody; (3) head in lateral view; (4) antenna; (5-7) aedeagus in lateral and in ventral view. Scales: 1-2: 0.5 mm; 3-7: 0.1 mm." pageId="1">Fig. 1</figureCitation>
); individual segments less transverse than in
<taxonomicName id="91D74D17C3367614323AF9F8FB8D7238" box="[877,1028,1569,1591]" class="Insecta" family="Staphylinidae" genus="Phloeocharis" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="subtilissima">
<emphasis id="64A3EA86C3367614323AF9F8FB8D7238" box="[877,1028,1569,1591]" italics="true" pageId="1">P. subtilissima</emphasis>
</taxonomicName>
; with distinct microreticulation and subdued shine; puncturation extremely fine and relatively sparse; posterior margin of tergite VII with narrow rudiment of a palisade fringe.
</paragraph>
<paragraph id="56683694C33676143003F95BFEF272BB" blockId="1.[340,1247,1665,1716]" pageId="1">
3: sternites VII and VIII unmodified; aedeagus as in
<figureCitation id="CEEC2A11C3367614322DF95BFC597298" box="[890,976,1665,1687]" captionStart="Figs 1-7" captionStartId="2.[340,380,1447,1467]" captionTargetBox="[343,1240,843,1436]" captionTargetId="figure-398@2.[340,1248,962,1436]" captionTargetPageId="2" captionText="Figs 1-7: Phloeocharis baenai nov.sp.: (1) habitus; (2) forebody; (3) head in lateral view; (4) antenna; (5-7) aedeagus in lateral and in ventral view. Scales: 1-2: 0.5 mm; 3-7: 0.1 mm." pageId="1">Figs 5-7</figureCitation>
, without spines in internal sac.
</paragraph>
<paragraph id="56683694C33676143003F91EFE5072F9" blockId="1.[340,1247,1731,1782]" pageId="1">
E t y m o l o g y: The species is dedicated to Manuel Baena, Córdoba, who collected the
<typeStatus id="896C8836C3367614302EF93AFE5C72F9" box="[377,469,1760,1782]" pageId="1" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="56683694C33676143003F8DCFB697313" blockId="1.[340,1248,1798,1820]" box="[340,1248,1798,1820]" pageId="1">
C o m p a r a t i v e n o t e s: From the widespread
<taxonomicName id="91D74D17C33676143279F8DDFBD07314" authority="MANNERHEIM" authorityName="MANNERHEIM" box="[814,1113,1798,1820]" class="Insecta" family="Staphylinidae" genus="Phloeocharis" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="subtilissima">
<emphasis id="64A3EA86C33676143279F8DDFC4D7313" box="[814,964,1798,1820]" italics="true" pageId="1">P. subtilissima</emphasis>
MANNERHEIM
</taxonomicName>
,
<taxonomicName id="91D74D17C33676143531F8DDFB417313" authorityName="Assing" authorityYear="2006" box="[1126,1224,1798,1820]" class="Insecta" family="Staphylinidae" genus="Phloeocharis" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="baenai">
<emphasis id="64A3EA86C33676143531F8DDFB417313" box="[1126,1224,1798,1820]" italics="true" pageId="1">P. baenai</emphasis>
</taxonomicName>
is
</paragraph>
<paragraph id="56683694C33576173255FEBFFCBB7575" blockId="2.[770,818,357,378]" box="[770,818,357,378]" pageId="2">1131</paragraph>
<paragraph id="56683694C33576173003FE75FB577634" blockId="2.[340,1248,431,571]" pageId="2">
distinguished by its distinctly smaller size and the more slender body (less transverse pronotum, elytra, and abdominal segments), the darker coloration of the elytra, the different morphology of the antennae (in
<taxonomicName id="91D74D17C335761733B1FE31FCF4760F" box="[742,893,490,512]" class="Insecta" family="Staphylinidae" genus="Phloeocharis" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="subtilissima">
<emphasis id="64A3EA86C335761733B1FE31FCF4760F" box="[742,893,490,512]" italics="true" pageId="2">P. subtilissima</emphasis>
</taxonomicName>
, antennomere
<collectionCode id="30C6AE51C33576173541FE31FBA1760F" box="[1046,1064,491,512]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" type="Museum">V</collectionCode>
is not transverse, VI-IX are weakly transverse, and the club is formed by antennomeres
<collectionCode id="30C6AE51C33576173513FDDDFB0F7612" box="[1092,1158,519,541]" pageId="2">IX-XI</collectionCode>
), the reduced palisade fringe at the posterior margin of tergite VIII, and by the smaller aedeagus.
</paragraph>
<paragraph id="56683694C33576173003FD90FCA87745" blockId="2.[340,1248,586,842]" pageId="2">
Previously, only one endemic
<taxonomicName id="91D74D17C335761733C2FD90FCE7766F" box="[661,878,586,608]" pageId="2">
<emphasis id="64A3EA86C335761733C2FD90FC95766F" box="[661,796,586,608]" italics="true" pageId="2">Phloeocharis</emphasis>
species
</taxonomicName>
had become known from southern
<collectingCountry id="2EC07604C33576173003FDBDFE1B7672" box="[340,402,615,637]" name="Spain" pageId="2">Spain</collectingCountry>
:
<taxonomicName id="91D74D17C335761730C9FDB2FD307672" authority="ASSING. From" authorityName="ASSING. From" box="[414,697,615,637]" class="Insecta" family="Staphylinidae" genus="Phloeocharis" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="bermejae">
<emphasis id="64A3EA86C335761730C9FDB2FD957672" box="[414,540,615,637]" italics="true" pageId="2">P. bermejae</emphasis>
ASSING. From
</taxonomicName>
this species,
<taxonomicName id="91D74D17C3357617321EFDB2FC247672" authorityName="Assing" authorityYear="2006" box="[841,941,615,637]" class="Insecta" family="Staphylinidae" genus="Phloeocharis" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="baenai">
<emphasis id="64A3EA86C3357617321EFDB2FC247672" box="[841,941,615,637]" italics="true" pageId="2">P. baenai</emphasis>
</taxonomicName>
is separated by much darker coloration (body in
<taxonomicName id="91D74D17C3357617337FFD5CFD2F7695" box="[552,678,644,667]" class="Insecta" family="Staphylinidae" genus="Phloeocharis" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="bermejae">
<emphasis id="64A3EA86C3357617337FFD5CFD2F7695" box="[552,678,644,667]" italics="true" pageId="2">P. bermejae</emphasis>
</taxonomicName>
uniformly dark yellowish), less bulging eyes, shorter antennae with much more transverse antennomeres V-IX, a more slender and less transverse pronotum with less convex lateral margins (dorsal view), distinctly longer and more slender elytra, a more slender abdomen, the presence of a rudiment of a palisade fringe at posterior margin of tergite VII (in
<taxonomicName id="91D74D17C33576173240FD21FC1B771F" box="[791,914,762,784]" class="Insecta" family="Staphylinidae" genus="Phloeocharis" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="bermejae">
<emphasis id="64A3EA86C33576173240FD21FC1B771F" box="[791,914,762,784]" italics="true" pageId="2">P. bermejae</emphasis>
</taxonomicName>
absent), and a smaller aedeagus with a more slender ventral process (lateral view). For illustrations of external and sexual characters of
<taxonomicName id="91D74D17C3357617308BFCEFFDDE7745" box="[476,599,820,842]" class="Insecta" family="Staphylinidae" genus="Phloeocharis" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="bermejae">
<emphasis id="64A3EA86C3357617308BFCEFFDDE7745" box="[476,599,820,842]" italics="true" pageId="2">P. bermejae</emphasis>
</taxonomicName>
see
<collectionCode id="30C6AE51C335761733D3FCEEFD597745" box="[644,720,820,842]" pageId="2">ASSING</collectionCode>
(2003).
</paragraph>
<caption id="02A8661CC33576173003FA7DFBF271DD" pageId="2" startId="2.[340,380,1447,1467]" targetBox="[343,1240,843,1436]" targetPageId="2" targetType="figure">
<paragraph id="56683694C33576173003FA7DFBF271DD" blockId="2.[340,1248,1447,1490]" pageId="2">
<emphasis id="64A3EA86C33576173003FA7DFE2C71B4" bold="true" box="[340,421,1447,1467]" pageId="2">Figs 1-7</emphasis>
:
<taxonomicName id="91D74D17C335761730E2FA7DFDFF71B4" authorityName="Assing" authorityYear="2006" box="[437,630,1447,1467]" class="Insecta" family="Staphylinidae" genus="Phloeocharis" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="baenai" status="sp. nov.">
<emphasis id="64A3EA86C335761730E2FA7DFDFF71B4" box="[437,630,1447,1467]" italics="true" pageId="2">Phloeocharis baenai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7F9057FDC335761733D7FA7DFD4F71B4" box="[640,710,1447,1467]" pageId="2" rank="species">nov.sp.</taxonomicNameLabel>
: (
<emphasis id="64A3EA86C3357617338DFA7DFD6C71B4" bold="true" box="[730,741,1447,1467]" pageId="2">1</emphasis>
) habitus; (
<emphasis id="64A3EA86C33576173206FA7DFCD571B4" bold="true" box="[849,860,1447,1467]" pageId="2">2</emphasis>
) forebody; (
<emphasis id="64A3EA86C3357617328FFA7DFC6A71B4" bold="true" box="[984,995,1447,1467]" pageId="2">3</emphasis>
) head in lateral view; (
<emphasis id="64A3EA86C3357617359AFA7DFB5171B4" bold="true" box="[1229,1240,1447,1467]" pageId="2">4</emphasis>
) antenna; (
<emphasis id="64A3EA86C335761730F9FA64FE4571DD" bold="true" box="[430,460,1470,1490]" pageId="2">5-7</emphasis>
) aedeagus in lateral and in ventral view. Scales: 1-2: 0.5 mm; 3-7: 0.1 mm.
</paragraph>
</caption>
<paragraph id="56683694C33576173003FA3BFB0C7286" blockId="2.[340,1248,1504,1673]" pageId="2">
D i s t r i b u t i o n a n d b i o n o m i c s: The type locality is situated some
<quantity id="912F9B71C33576173591FA3AFEFD721C" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="2" unit="km" value="10.0">10 km</quantity>
to the southeast of Baena, Córdoba (
<figureCitation id="CEEC2A11C335761733A4FA27FCBE721C" box="[755,823,1533,1555]" captionStart="Map 1" captionStartId="3.[340,385,1351,1371]" captionTargetBox="[340,1247,707,1340]" captionTargetId="figure-370@3.[340,1247,707,1340]" captionTargetPageId="3" captionText="Map 1: Distributions of Phloeocharis baenai nov.sp. (open circle), Geostiba jaenica nov.sp. (filled circles), and Atheta tenebrarum nov.sp. (square) in southern Spain." pageId="2">Map 1</figureCitation>
). The reduced palisade fringe at the posterior margin of tergite VII is characteristic of species with reduced wings and wing muscles. This, as well as the absence of other records suggests that
<taxonomicName id="91D74D17C33576173554F9E3FBEC7241" authorityName="Assing" authorityYear="2006" box="[1027,1125,1592,1614]" class="Insecta" family="Staphylinidae" genus="Phloeocharis" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="baenai">
<emphasis id="64A3EA86C33576173554F9E3FBEC7241" box="[1027,1125,1592,1614]" italics="true" pageId="2">P. baenai</emphasis>
</taxonomicName>
may have a restricted distribution. The
<typeStatus id="896C8836C33576173338F98FFD417264" box="[623,712,1621,1643]" pageId="2" type="holotype">holotype</typeStatus>
was found under a stone at an altitude of approximately
<quantity id="912F9B71C335761730C8F9A9FE567287" box="[415,479,1651,1673]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="2" unit="m" value="800.0">800 m</quantity>
(
<collectionCode id="30C6AE51C335761730BAF9A9FDBF7287" box="[493,566,1651,1673]" pageId="2">BAENA</collectionCode>
pers. comm.). Additional bionomic data are not available.
</paragraph>
</subSubSection>
</treatment>
</document>