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(
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136
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Molecular phylogenetic placement of
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While the head of
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is suggestive of
|
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<taxonomicName id="4C524D13FFE3FFCA4BB8F901B160F8FA" box="[673,800,1769,1793]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="tribe" tribe="Trogastrini">Trogastrini</taxonomicName>
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, other characters typical of this tribe are absent. There is no dense genal beard of setae, the tarsal claws are both well-developed and equally-sized, rather than being unequal in size or consisting of one well-developed and one setiferous claw. A strongly-impressed median longitudinal sulcus on the pronotal disc is also lacking; although the sulcus is present, it is barely evident and only weaklydefined. Furthermore, the obcordate form of the pronotum, sharply constricted at the base, is far more similar to members of Jubini, another euplectite tribe.
|
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</paragraph>
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<emphasis id="B926EA82FFE2FFCB49D0FF70B306FF4B" box="[201,326,152,176]" italics="true" pageId="7" pageNumber="376">Jubogaster</emphasis>
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</taxonomicName>
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’s curious suite of features confounds a definitive assignment of the new taxon to tribe using morphology alone. To reliably place
|
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<emphasis id="B926EA82FFE2FFCB4B5FFF54B083FF2F" box="[582,707,188,212]" italics="true" pageId="7" pageNumber="376">Jubogaster</emphasis>
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phylogenetically, we employed a non-destructive procedure to extract DNA from the
|
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specimen, without damaging or altering its external morphology (
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Gilbert
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<emphasis id="B926EA82FFE2FFCB4C0BFF09B70CFF03" box="[1298,1356,224,248]" italics="true" pageId="7" pageNumber="376">et al.</emphasis>
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2007
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||||
</bibRefCitation>
|
||||
). Using this approach, we successfully amplified a fragment of 28s rDNA from
|
||||
<taxonomicName id="4C524D13FFE2FFCB4D16FEEDB628FEE7" box="[1039,1128,261,284]" class="Insecta" family="Silphidae" genus="Jubogaster" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="376" phylum="Arthropoda" rank="species" species="towai">
|
||||
<emphasis id="B926EA82FFE2FFCB4D16FEEDB628FEE7" box="[1039,1128,261,284]" italics="true" pageId="7" pageNumber="376">J. towai</emphasis>
|
||||
</taxonomicName>
|
||||
. This region is particularly informative for higher-level pselaphine systematics, resolving many relationships at the supertribal, tribal and subtribal levels (J. Parker and A. Vogler, unpublished data). We also sequenced this gene region from a range of other
|
||||
<taxonomicName id="4C524D13FFE2FFCB49C2FE87B321FE7C" box="[219,353,367,391]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="376" phylum="Arthropoda" rank="subFamily" subFamily="Pselaphinae">Pselaphinae</taxonomicName>
|
||||
including 5 genera of
|
||||
<taxonomicName id="4C524D13FFE2FFCB4B77FE87B0ACFE7C" box="[622,748,367,391]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="376" phylum="Arthropoda" rank="tribe" tribe="Trogastrini">Trogastrini</taxonomicName>
|
||||
, 3 Jubini genera, and representatives of all other euplectite tribes except Metopiasini, Dimerini and Mayetiini. A variety of non-euplectite pselaphines from different supertribes were included, totalling 24 ingroup taxa. Three other staphylinids (from the subfamilies
|
||||
<taxonomicName id="4C524D13FFE2FFCB4C0CFE5FB7D7FE34" box="[1301,1431,439,463]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="376" phylum="Arthropoda" rank="subFamily" subFamily="Proteininae">Proteininae</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C524D13FFE2FFCB498EFE34B373FE0F" box="[151,307,476,500]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="376" phylum="Arthropoda" rank="subFamily" subFamily="Euaesthetinae">Euaesthetinae</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C524D13FFE2FFCB4870FE34B04EFE0F" box="[361,526,476,500]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="376" phylum="Arthropoda" rank="subFamily" subFamily="Scydmaeninae">Scydmaeninae</taxonomicName>
|
||||
) and the silphid
|
||||
<taxonomicName id="4C524D13FFE2FFCB4BD1FE34B130FE0F" box="[712,880,476,500]" class="Insecta" family="Curculionidae" genus="Silpha" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="376" phylum="Arthropoda" rank="species" species="obscura">
|
||||
<emphasis id="B926EA82FFE2FFCB4BD1FE34B130FE0F" box="[712,880,476,500]" italics="true" pageId="7" pageNumber="376">Silpha obscura</emphasis>
|
||||
</taxonomicName>
|
||||
were used as outgroups.
|
||||
</paragraph>
|
||||
<paragraph id="8BED3690FFE2FFCB49DEFE17B35AFD14" blockId="7.[151,1436,151,751]" pageId="7" pageNumber="376">
|
||||
Both MP (
|
||||
<figureCitation id="13692A15FFE2FFCB4858FE17B3CDFDEC" box="[321,397,511,535]" captionStart="FIGURES 11 – 12" captionStartId="7.[151,264,1335,1358]" captionTargetBox="[159,1430,815,1302]" captionTargetId="figure@7.[151,1436,805,1314]" captionTargetPageId="7" captionText="FIGURES 11 – 12. Phylogenetic position of Jubogaster towai. Trees were rooted using Silpha obscura (Silphidae) as the designated outgroup. Grey branches denote outgroup taxa, black branches non-euplectite Pselaphinae, green branches Euplectitae, and purple branches Trogastrini. 11) Single most parsimonious tree of 2236 steps (CI = 0.50, RI = 0.56) found from an MP search on the default MUSCLE alignment. Values above branches are bootstrap proportions from 1000 replicates. ClustalX alignments made with a range of gap opening and extension costs produced highly similar topologies that differ only outside the Jubini + Trogastrini clade. 12) Bayesian consensus tree made from an unpartitioned analysis with the GTR + I + G model. Posterior probabilities for each node are shown above branches." httpUri="https://zenodo.org/record/222182/files/figure.png" pageId="7" pageNumber="376">Fig 11</figureCitation>
|
||||
) and Bayesian (
|
||||
<figureCitation id="13692A15FFE2FFCB4B53FE17B0D8FDEC" box="[586,664,511,535]" captionStart="FIGURES 11 – 12" captionStartId="7.[151,264,1335,1358]" captionTargetBox="[159,1430,815,1302]" captionTargetId="figure@7.[151,1436,805,1314]" captionTargetPageId="7" captionText="FIGURES 11 – 12. Phylogenetic position of Jubogaster towai. Trees were rooted using Silpha obscura (Silphidae) as the designated outgroup. Grey branches denote outgroup taxa, black branches non-euplectite Pselaphinae, green branches Euplectitae, and purple branches Trogastrini. 11) Single most parsimonious tree of 2236 steps (CI = 0.50, RI = 0.56) found from an MP search on the default MUSCLE alignment. Values above branches are bootstrap proportions from 1000 replicates. ClustalX alignments made with a range of gap opening and extension costs produced highly similar topologies that differ only outside the Jubini + Trogastrini clade. 12) Bayesian consensus tree made from an unpartitioned analysis with the GTR + I + G model. Posterior probabilities for each node are shown above branches." httpUri="https://zenodo.org/record/222182/files/figure.png" pageId="7" pageNumber="376">Fig 12</figureCitation>
|
||||
) analyses placed
|
||||
<taxonomicName id="4C524D13FFE2FFCB4A7FFDE8B1A3FDE3" box="[870,995,512,536]" class="Insecta" family="Silphidae" genus="Jubogaster" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="371" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B926EA82FFE2FFCB4A7FFDE8B1A3FDE3" box="[870,995,512,536]" italics="true" pageId="7" pageNumber="376">Jubogaster</emphasis>
|
||||
</taxonomicName>
|
||||
within the
|
||||
<taxonomicName id="4C524D13FFE2FFCB4D73FE17B6A5FDEC" box="[1130,1253,511,535]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="376" phylum="Arthropoda" rank="tribe" tribe="Trogastrini">Trogastrini</taxonomicName>
|
||||
with maximum support, forming a clade with
|
||||
<taxonomicName id="4C524D13FFE2FFCB4B10FDCCB0CBFDC7" box="[521,651,548,572]" class="Insecta" family="Staphylinidae" genus="Platomesus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="376" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B926EA82FFE2FFCB4B10FDCCB0CBFDC7" box="[521,651,548,572]" italics="true" pageId="7" pageNumber="376">Platomesus</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C524D13FFE2FFCB4BD4FDCCB166FDC7" box="[717,806,548,572]" class="Insecta" family="Staphylinidae" genus="Rhexius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="376" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B926EA82FFE2FFCB4BD4FDCCB166FDC7" box="[717,806,548,572]" italics="true" pageId="7" pageNumber="376">Rhexius</emphasis>
|
||||
</taxonomicName>
|
||||
. This placement was recovered across all alignment parameter combinations tested with consistently high support. Although the taxonomic coverage is limited,
|
||||
<taxonomicName id="4C524D13FFE2FFCB498EFD84B354FD7F" box="[151,276,620,644]" class="Insecta" family="Silphidae" genus="Jubogaster" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="371" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B926EA82FFE2FFCB498EFD84B354FD7F" box="[151,276,620,644]" italics="true" pageId="7" pageNumber="376">Jubogaster</emphasis>
|
||||
</taxonomicName>
|
||||
emerges in the same position in a far more comprehensive analysis of>150 ingroup taxa, with extensive taxon sampling from almost all pselaphine tribes and subtribes (data not shown; J. Parker and A. Vogler, unpublished data). Hence, we conclude with a high degree of certainty that
|
||||
<taxonomicName id="4C524D13FFE2FFCB4D17FD5CB6CAFD37" box="[1038,1162,692,716]" class="Insecta" family="Silphidae" genus="Jubogaster" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="371" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B926EA82FFE2FFCB4D17FD5CB6CAFD37" box="[1038,1162,692,716]" italics="true" pageId="7" pageNumber="376">Jubogaster</emphasis>
|
||||
</taxonomicName>
|
||||
is indeed a member of
|
||||
<taxonomicName id="4C524D13FFE2FFCB498EFD3FB356FD14" box="[151,278,727,751]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="376" phylum="Arthropoda" rank="tribe" tribe="Trogastrini">Trogastrini</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
347
data/03/FB/87/03FB8786FFE6FFCA498EFA2EB0FDF9B5.xml
Normal file
347
data/03/FB/87/03FB8786FFE6FFCA498EFA2EB0FDF9B5.xml
Normal file
|
|
@ -0,0 +1,347 @@
|
|||
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|
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<mods:mods id="F71BDBD859AA806A8F372DB0AF5CE560" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="2A4221EC4D86F30E64B549DFF6C3B404">
|
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<mods:title id="A8D626C735DEAD83B42C8B4D1A6E8EE4">Jubogaster towai, a new Neotropical genus and species of Trogastrini (Coleoptera: Staphylinidae: Pselaphinae) exhibiting myrmecophily and extreme body enlargement</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:name id="41261D38254B027428A32EA9B450DDF7" type="personal">
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<mods:role id="1277AA994809D9BC66B087D379504E2B">
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<mods:roleTerm id="B94F71E8D575489285AD78F8206D7C59">Author</mods:roleTerm>
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||||
</mods:role>
|
||||
<mods:namePart id="219E3E8F9A2EEBB0ABD49C71ACDCB819">Parker, Joseph</mods:namePart>
|
||||
</mods:name>
|
||||
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<mods:roleTerm id="B2715105788F119C8E6C00D1D8B154BF">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="317686E754F53B39D10821B8C4B27C8A">Maruyama, Munetoshi</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:typeOfResource id="A7F516DC2A3D2C8E1CCBF1B6D8326CA6">text</mods:typeOfResource>
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<mods:title id="B14D38580A43C238E00C33EFA6D9F6A8">Zootaxa</mods:title>
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</mods:titleInfo>
|
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<mods:part id="EFECE8F571C688A0680C5C5991A181EE">
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<mods:date id="1C75772094D3E85ADF71C18C12F14A29">2013</mods:date>
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<mods:detail id="1EA5A81B566095523B0A73A997713C7F" type="pubDate">
|
||||
<mods:number id="1260FCEA9433076630E19FEFBB2133A3">2013-03-20</mods:number>
|
||||
</mods:detail>
|
||||
<mods:detail id="47A71835D4BA9CD8E509CF7196237D7B" type="volume">
|
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<mods:number id="986B3DA622FB4F753015F1603BB38A87">3630</mods:number>
|
||||
</mods:detail>
|
||||
<mods:detail id="D09321D96660D7F6131C898FBFC7B1D4" type="issue">
|
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<mods:number id="582FFEE3DFEB671D9552D74BC239D7A0">2</mods:number>
|
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|
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|
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|
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|
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|
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<mods:classification id="AA7EDBF12834441BDA3CEBC3CB778241">journal article</mods:classification>
|
||||
<mods:identifier id="E357F3E5E07C3D5F18827FCC5AE0D5FF" type="DOI">10.11646/zootaxa.3630.2.11</mods:identifier>
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||||
<mods:identifier id="8EF40AEA0B6D5E3657D78B853207510E" type="ISSN">1175-5326</mods:identifier>
|
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<mods:identifier id="346BDEF985C7EE4B222DC30EBC8FB6C2" type="Zenodo-Dep">222179</mods:identifier>
|
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<mods:identifier id="A69DB409C7627712BD250B6CD8C00F94" type="ZooBank">70C11335-A5F6-4BCB-98E6-4D14EB1A4015</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment id="03FB8786FFE6FFCA498EFA2EB0FDF9B5" ID-DOI="http://doi.org/10.5281/zenodo.14040091" ID-Zenodo-Dep="14040091" LSID="urn:lsid:zoobank.org:act:F60FBF55-4ECB-4092-8A8E-5FF5D1C55E0C" httpUri="http://treatment.plazi.org/id/03FB8786FFE6FFCA498EFA2EB0FDF9B5" lastPageId="6" lastPageNumber="375" pageId="3" pageNumber="372">
|
||||
<subSubSection id="C348651BFFE6FFCF498EFA2EB3FCFA24" box="[151,444,1478,1503]" pageId="3" pageNumber="372" type="nomenclature">
|
||||
<paragraph id="8BED3690FFE6FFCF498EFA2EB3FCFA24" blockId="3.[151,444,1478,1503]" box="[151,444,1478,1503]" pageId="3" pageNumber="372">
|
||||
<emphasis id="B926EA82FFE6FFCF498EFA2EB3FCFA24" bold="true" box="[151,444,1478,1503]" pageId="3" pageNumber="372">
|
||||
<taxonomicName id="4C524D13FFE6FFCF498EFA2EB31BFA24" authority="Parker & Maruyama, 2013" authorityName="Parker & Maruyama" authorityYear="2013" box="[151,347,1478,1503]" class="Insecta" family="Silphidae" genus="Jubogaster" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="species" species="towai" status="sp. nov.">
|
||||
<emphasis id="B926EA82FFE6FFCF498EFA2EB31BFA24" bold="true" box="[151,347,1478,1503]" italics="true" pageId="3" pageNumber="372">Jubogaster towai</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A21557F9FFE6FFCF487AFA2FB3FCFA24" box="[355,444,1479,1503]" pageId="3" pageNumber="372" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C348651BFFE6FFCF498EF9E7B7DDF9B1" pageId="3" pageNumber="372" type="materials_examined">
|
||||
<paragraph id="8BED3690FFE6FFCF498EF9E7B7DDF9B1" blockId="3.[151,1437,1550,2006]" pageId="3" pageNumber="372">
|
||||
<emphasis id="B926EA82FFE6FFCF498EF9E7B37EF9DC" bold="true" box="[151,318,1550,1575]" pageId="3" pageNumber="372">
|
||||
<typeStatus id="54E98832FFE6FFCF498EF9E7B28BF9DC" box="[151,203,1551,1575]" pageId="3" pageNumber="372">type</typeStatus>
|
||||
material.
|
||||
</emphasis>
|
||||
<materialsCitation id="3B3A3CCDFFE6FFCF485DF9E6B7DDF9B1" collectingDate="2012-01-25" country="Peru" location="Santa Ana" municipality="Satipo" pageId="3" pageNumber="372" specimenCode="FMNH-INS 0 0 0 0 0 93 389" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||||
<typeStatus id="54E98832FFE6FFCF485DF9E6B3EFF9DD" box="[324,431,1550,1574]" pageId="3" pageNumber="372" type="holotype">Holotype</typeStatus>
|
||||
,
|
||||
<specimenCount id="9D54FD19FFE6FFCF48A2F9E6B3B1F9DD" box="[443,497,1550,1574]" pageId="3" pageNumber="372" type="male">male</specimenCount>
|
||||
,
|
||||
<collectingCountry id="F3457600FFE6FFCF48E6F9E7B075F9DD" box="[511,565,1551,1574]" name="Peru" pageId="3" pageNumber="372">Peru</collectingCountry>
|
||||
,
|
||||
<collectingMunicipality id="6B89ACEAFFE6FFCF4B58F9E6B0CBF9DD" box="[577,651,1550,1574]" pageId="3" pageNumber="372">Satipo</collectingMunicipality>
|
||||
,
|
||||
<location id="8E8D604BFFE6FFCF4B8EF9E6B14EF9DC" LSID="urn:lsid:plazi:treatment:03FB8786FFE6FFCA498EFA2EB0FDF9B5:8E8D604BFFE6FFCF4B8EF9E6B14EF9DC" box="[663,782,1550,1575]" country="Peru" municipality="Satipo" name="Santa Ana" pageId="3" pageNumber="372">Santa Ana</location>
|
||||
,
|
||||
<date id="FFEC1050FFE6FFCF4A00F9E6B1C7F9DD" box="[793,903,1550,1574]" pageId="3" pageNumber="372" value="2012-01-25">
|
||||
<collectingDate id="EFA8E9B8FFE6FFCF4A00F9E6B1C7F9DD" box="[793,903,1550,1574]" pageId="3" pageNumber="372" value="2012-01-25">25 I 2012</collectingDate>
|
||||
</date>
|
||||
, with
|
||||
<taxonomicName id="4C524D13FFE6FFCF4AD5F9E6B7DBF9DD" authority="Wilson. Specimen" authorityName="Wilson. Specimen" box="[972,1435,1550,1574]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="species" species="xanthogaster">
|
||||
<emphasis id="B926EA82FFE6FFCF4AD5F9E6B688F9DD" box="[972,1224,1550,1574]" italics="true" pageId="3" pageNumber="372">Pheidole xanthogaster</emphasis>
|
||||
Wilson. Specimen
|
||||
</taxonomicName>
|
||||
deposited in Field Museum of Natural History, Chicago, IL, USA, with catalog number:
|
||||
<specimenCode id="DBF49EEBFFE6FFCF4D76F9DAB7D8F9B1" box="[1135,1432,1586,1610]" pageId="3" pageNumber="372">FMNH-INS 0000 093 389</specimenCode>
|
||||
.
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C348651BFFE6FFCF49D0F9BEB76DF921" pageId="3" pageNumber="372" type="diagnosis">
|
||||
<paragraph id="8BED3690FFE6FFCF49D0F9BEB76DF921" blockId="3.[151,1437,1550,2006]" pageId="3" pageNumber="372">
|
||||
<emphasis id="B926EA82FFE6FFCF49D0F9BEB303F994" bold="true" box="[201,323,1622,1647]" pageId="3" pageNumber="372">Diagnosis.</emphasis>
|
||||
The sole known species of
|
||||
<taxonomicName id="4C524D13FFE6FFCF4B92F9BEB148F995" box="[651,776,1622,1646]" class="Insecta" family="Silphidae" genus="Jubogaster" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="371" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B926EA82FFE6FFCF4B92F9BEB148F995" box="[651,776,1622,1646]" italics="true" pageId="3" pageNumber="372">Jubogaster</emphasis>
|
||||
</taxonomicName>
|
||||
is highly distinctive among
|
||||
<taxonomicName id="4C524D13FFE6FFCF4D4AF9BEB68FF995" box="[1107,1231,1622,1646]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="tribe" tribe="Trogastrini">Trogastrini</taxonomicName>
|
||||
due to its greatly enlarged and robust body covered in thick, blunt golden setae, the unique concave head that lacks vertexal fovea, the large, circular excavations on the frons that receive the long, geniculate antennae, the absence of a genal beard, an afoveate pronotum and elytra, the two equally-sized tarsal claws, and the bifid anterior protarsal claw.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C348651BFFE6FFCF49D0F90FB392F8D9" pageId="3" pageNumber="372" type="etymology">
|
||||
<paragraph id="8BED3690FFE6FFCF49D0F90FB392F8D9" blockId="3.[151,1437,1550,2006]" pageId="3" pageNumber="372">
|
||||
<emphasis id="B926EA82FFE6FFCF49D0F90FB310F904" bold="true" box="[201,336,1767,1791]" pageId="3" pageNumber="372">Etymology.</emphasis>
|
||||
The new species is named in honour of Towa Shimada, son of Taku Shimada, a remarkable collector of myrmecophiles.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C348651BFFE6FFC949D0F8C6B290F8C4" lastPageId="5" lastPageNumber="374" pageId="3" pageNumber="372" type="description">
|
||||
<paragraph id="8BED3690FFE6FFCF49D0F8C6B070F82D" blockId="3.[151,1437,1550,2006]" pageId="3" pageNumber="372">
|
||||
<emphasis id="B926EA82FFE6FFCF49D0F8C6B31AF8BC" bold="true" box="[201,346,1838,1863]" pageId="3" pageNumber="372">Description.</emphasis>
|
||||
Body length
|
||||
<quantity id="4CAA9B75FFE6FFCF48E3F8C6B009F8BD" box="[506,585,1838,1862]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="3" pageNumber="372" unit="mm" value="5.1">5.1mm</quantity>
|
||||
(
|
||||
<figureCitation id="13692A15FFE6FFCF4B43F8C6B0D8F8BD" box="[602,664,1838,1862]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="3" pageNumber="372">Fig 1</figureCitation>
|
||||
). Colour dark reddish brown, antennae and legs somewhat lighter. Integument shiny. Most dorsal regions covered by conspicuous thick, erect, blunt golden setae, arranged at regularly spaced intervals and orientated posteriorly (depicted most clearly in figures 4
|
||||
<emphasis id="B926EA82FFE6FFCF4D75F89FB63AF874" bold="true" box="[1132,1146,1911,1935]" pageId="3" pageNumber="372">–</emphasis>
|
||||
6). These are interspersed with shorter pubescence of the same colour, also orientated posteriorly. Ventral surfaces covered with fine, mostly aciculate golden pubescence (
|
||||
<figureCitation id="13692A15FFE6FFCF48FDF856B063F82D" box="[484,547,1982,2006]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="3" pageNumber="372">Fig 2</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="DF2D6618FFE1FFC8498EF89CB7DBF833" httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="4" pageNumber="373" targetBox="[151,1435,211,1884]" targetPageId="4">
|
||||
<paragraph id="8BED3690FFE1FFC8498EF89CB7DBF833" blockId="4.[151,1437,1908,1992]" pageId="4" pageNumber="373">
|
||||
<emphasis id="B926EA82FFE1FFC8498EF89CB37AF871" bold="true" box="[151,314,1908,1931]" pageId="4" pageNumber="373">FIGURES 1–7.</emphasis>
|
||||
<taxonomicName id="4C524D13FFE1FFC84858F89DB09FF871" authority="Parker and Maruyama." authorityName="Parker and Maruyama." box="[321,735,1908,1931]" class="Insecta" family="Silphidae" genus="Jubogaster" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="species" species="towai">
|
||||
<emphasis id="B926EA82FFE1FFC84858F89DB3AFF870" box="[321,495,1909,1931]" italics="true" pageId="4" pageNumber="373">Jubogaster towai</emphasis>
|
||||
Parker and Maruyama.
|
||||
</taxonomicName>
|
||||
<emphasis id="B926EA82FFE1FFC84BFDF89CB0BAF870" bold="true" box="[740,762,1908,1931]" pageId="4" pageNumber="373">1)</emphasis>
|
||||
dorsal habitus
|
||||
<emphasis id="B926EA82FFE1FFC84A8DF89CB1E8F870" bold="true" box="[916,936,1908,1931]" pageId="4" pageNumber="373">2)</emphasis>
|
||||
ventral habitus
|
||||
<emphasis id="B926EA82FFE1FFC84D55F89CB621F870" bold="true" box="[1100,1121,1908,1931]" pageId="4" pageNumber="373">3)</emphasis>
|
||||
left antenna, exterior face is to the left
|
||||
<emphasis id="B926EA82FFE1FFC849FEF87BB2BDF851" bold="true" box="[231,253,1939,1962]" pageId="4" pageNumber="373">4)</emphasis>
|
||||
vertex of head and pronotum
|
||||
<emphasis id="B926EA82FFE1FFC84B21F87BB00DF851" bold="true" box="[568,589,1939,1962]" pageId="4" pageNumber="373">5)</emphasis>
|
||||
frontal of view of head, showing excavations for antennae and maxillary palpi
|
||||
<emphasis id="B926EA82FFE1FFC84C9EF87BB7DCF851" bold="true" box="[1415,1436,1939,1962]" pageId="4" pageNumber="373">6)</emphasis>
|
||||
right elytron apical angle showing excision
|
||||
<emphasis id="B926EA82FFE1FFC84B4BF859B027F833" bold="true" box="[594,615,1969,1992]" pageId="4" pageNumber="373">7)</emphasis>
|
||||
left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BED3690FFE0FFC949D0FF7FB06FFD10" blockId="5.[151,1437,151,2034]" pageId="5" pageNumber="374">
|
||||
<emphasis id="B926EA82FFE0FFC949D0FF7FB350FF4B" bold="true" box="[201,272,151,176]" italics="true" pageId="5" pageNumber="374">Head:</emphasis>
|
||||
Eyes moderate in size. Transverse, triangular head (
|
||||
<figureCitation id="13692A15FFE0FFC94A7BFF7FB1E0FF54" box="[866,928,151,175]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="5" pageNumber="374">Fig 4</figureCitation>
|
||||
) widest just behind eyes. Vertex and occiput uniformly setiferous with thick, blunt, erect golden setae, and shorter, sharper pubescence. Margins of large lateral cavities for antennal insertion and maxillary palp retraction carinate; interiors of cavities glabrous (
|
||||
<figureCitation id="13692A15FFE0FFC94C10FF37B709FF0C" box="[1289,1353,223,247]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="5" pageNumber="374">Fig 5</figureCitation>
|
||||
). Neck approximately 0.4x head width between eyes, shining and with sparse, very short dark setae on dorsal side; glabrous and with finger-print-like (“strigate”) microsculpturation on ventral side. Fourth segments of maxillary palpi (
|
||||
<figureCitation id="13692A15FFE0FFC949FAFEA2B364FE99" box="[227,292,330,354]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="5" pageNumber="374">Fig 5</figureCitation>
|
||||
) densely covered in short golden setae. Antennae (
|
||||
<figureCitation id="13692A15FFE0FFC94A92FEA2B18CFE99" box="[907,972,330,354]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="5" pageNumber="374">Fig 3</figureCitation>
|
||||
) pigmented similarly to tibia and very elongate, approximately
|
||||
<date id="FFEC1050FFE0FFC948A8FE86B38CFE7D" box="[433,460,366,390]" pageId="5" pageNumber="374">2x</date>
|
||||
pronotum length. Antennomere I conical, widest in basal half and equal in length to III+IV combined. Laterodorsal margin with short arcuate carina extending to one-quarter antennomere length. Lateral face glabrous in basal half. With circular apical notch to permit geniculation with II; II small and cylindrical, approximately 0.6x the length of III; III cylindrical-obconical, equal in length to IV; IV cylindrical, twice as long as wide; V slightly shorter and narrower than IV; V
|
||||
<emphasis id="B926EA82FFE0FFC94A63FE15B1C8FDEE" bold="true" box="[890,904,509,533]" pageId="5" pageNumber="374">–</emphasis>
|
||||
VIII cylindrical-obconical, becoming gradually shorter apically, so that segment V is almost
|
||||
<date id="FFEC1050FFE0FFC94BB9FDC9B0FCFDC2" box="[672,700,545,569]" pageId="5" pageNumber="374">2x</date>
|
||||
longer than wide and VIII is as only slightly longer than wide; IX
|
||||
<emphasis id="B926EA82FFE0FFC949ADFDADB282FDA6" bold="true" box="[180,194,581,605]" pageId="5" pageNumber="374">–</emphasis>
|
||||
X much larger, IX almost as long as VI
|
||||
<emphasis id="B926EA82FFE0FFC94B91FDADB0D6FDA6" bold="true" box="[648,662,581,605]" pageId="5" pageNumber="374">–</emphasis>
|
||||
VIII combined; X slightly longer than VII+VIII combined. Both IX and X cylindrical-obconical, apical margins somewhat sloping interiorly due to slight lengthening of exterior antennomere face relative to interior face. Antennomere XI longest, twice the length of X; exterior face shallowly and evenly convex, interior face weakly concave before bending obtusely to form conical apex. Several long sensory setae on apical third (
|
||||
<figureCitation id="13692A15FFE0FFC948FAFD3BB061FD10" box="[483,545,723,747]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="5" pageNumber="374">Fig 3</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BED3690FFE0FFC949D0FD1FB687FBAA" blockId="5.[151,1437,151,2034]" pageId="5" pageNumber="374">
|
||||
<emphasis id="B926EA82FFE0FFC949D0FD1FB368FCEB" bold="true" box="[201,296,759,784]" italics="true" pageId="5" pageNumber="374">Thorax:</emphasis>
|
||||
Pronotum (
|
||||
<figureCitation id="13692A15FFE0FFC948B2FD1FB3A8FCF4" box="[427,488,759,783]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="5" pageNumber="374">Fig 4</figureCitation>
|
||||
) obcordate, margins smooth, lacking crenulations or other protuberances. Uniformly setiferous and pubescent. Prosternum with lateral longitudinal carinae abruptly bending dorsally on reaching base (
|
||||
<figureCitation id="13692A15FFE0FFC94986FCD7B29FFCAC" box="[159,223,831,855]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="5" pageNumber="374">Fig 2</figureCitation>
|
||||
). Region between carinae with reduced setation and pubescence relative to lateral regions, and with small transverse projection situated medially, anterior to procoxae. Region between carinae projecting sharply in basolateral portions to receive procoxae. Mesosternum with setiferous median mesosternal foveae (MMSF) and lateral mesocoxal fovea (LMCF). Region anterior to mesocoxae formed into a shield-shaped plate with carinate margins. Anterior margin of metasternum carinate around mesocoxae. Convex metasternum with median declivity adjacent to metacoxal projections. Metasternum uniformly covered with yellow pubescence of intermediate length. Median declivity somewhat less pubescent. Lateral metasternal fovea small and setiferous. Posterior margin of metasternum bow shaped (
|
||||
<figureCitation id="13692A15FFE0FFC948DBFBD1B040FBAA" box="[450,512,1081,1105]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="5" pageNumber="374">Fig 2</figureCitation>
|
||||
); projection between metacoxae with small medial indentation.
|
||||
</paragraph>
|
||||
<paragraph id="8BED3690FFE0FFC949D0FBB4B093FA69" blockId="5.[151,1437,151,2034]" pageId="5" pageNumber="374">
|
||||
<emphasis id="B926EA82FFE0FFC949D0FBB4B300FB8E" bold="true" box="[201,320,1116,1141]" pageId="5" pageNumber="374">
|
||||
<emphasis id="B926EA82FFE0FFC949D0FBB4B377FB8E" bold="true" box="[201,311,1116,1141]" italics="true" pageId="5" pageNumber="374">Abdomen</emphasis>
|
||||
:
|
||||
</emphasis>
|
||||
Tergites strongly convex with visible tergite 1 (IV) sloping to abdomen base. Visible tergite 2 (V) broadest, slightly broader than 1 (IV), with segments 3
|
||||
<emphasis id="B926EA82FFE0FFC94A0FFB69B164FB62" bold="true" box="[790,804,1153,1177]" pageId="5" pageNumber="374">–</emphasis>
|
||||
5 (VI
|
||||
<emphasis id="B926EA82FFE0FFC94A7DFB69B132FB62" bold="true" box="[868,882,1153,1177]" pageId="5" pageNumber="374">–</emphasis>
|
||||
VIII) evenly narrowing. Tergite 4 (VII) longest, approximately 1.5x tergite 3 (VI). Broad paratergites on segments 1
|
||||
<emphasis id="B926EA82FFE0FFC94A81FB4DB1E6FB46" bold="true" box="[920,934,1189,1213]" pageId="5" pageNumber="374">–</emphasis>
|
||||
3 (IV
|
||||
<emphasis id="B926EA82FFE0FFC94AFAFB4DB1B1FB46" bold="true" box="[995,1009,1189,1213]" pageId="5" pageNumber="374">–</emphasis>
|
||||
VI); paratergite margins of segment 4 converging apically to create a triangular paratergite. Anterior edges of abdominal tergites with narrow band of black pigment. Remainder of tergite reddish brown; intertergital membrane yellow with latticed microsculpture. Base of visible tergite 1 (IV) with short, blunt tubercles, situated halfway between midline and lateral margin. First visible sternite (IV) medially acuminate in anterior half, projecting ventrally between metacoxae. Last visible segment (6; VIII) with median declivity. Sternite 7 (XI) internalised. Pubescence on sternites 2
|
||||
<emphasis id="B926EA82FFE0FFC94C1CFABFB753FA94" bold="true" box="[1285,1299,1367,1391]" pageId="5" pageNumber="374">–</emphasis>
|
||||
6 (IV
|
||||
<emphasis id="B926EA82FFE0FFC94C4DFABFB722FA94" bold="true" box="[1364,1378,1367,1391]" pageId="5" pageNumber="374">–</emphasis>
|
||||
VIII) becoming finer and sparser toward midline (
|
||||
<figureCitation id="13692A15FFE0FFC94B9EFA92B086FA69" box="[647,710,1402,1426]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="5" pageNumber="374">Fig 2</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BED3690FFE0FFC949D0FA76B0AAFA21" blockId="5.[151,1437,151,2034]" pageId="5" pageNumber="374">
|
||||
<emphasis id="B926EA82FFE0FFC949D0FA76B359FA4D" bold="true" box="[201,281,1438,1463]" pageId="5" pageNumber="374">
|
||||
<emphasis id="B926EA82FFE0FFC949D0FA76B351FA4C" bold="true" box="[201,273,1438,1463]" italics="true" pageId="5" pageNumber="374">Elytra</emphasis>
|
||||
:
|
||||
</emphasis>
|
||||
Blunt setae on elytra somewhat longer and slightly thicker than on other parts of body. Apicolateral angles of elytra with margins weakly excised (
|
||||
<figureCitation id="13692A15FFE0FFC94B87FA2AB09CFA21" box="[670,732,1474,1498]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="5" pageNumber="374">Fig 6</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BED3690FFE0FFC949D0FA0EB66BF92F" blockId="5.[151,1437,151,2034]" pageId="5" pageNumber="374">
|
||||
<emphasis id="B926EA82FFE0FFC949D0FA0EB348FA05" bold="true" box="[201,264,1510,1534]" pageId="5" pageNumber="374">
|
||||
<emphasis id="B926EA82FFE0FFC949D0FA0EB2BFFA06" bold="true" box="[201,255,1510,1533]" italics="true" pageId="5" pageNumber="374">Legs</emphasis>
|
||||
:
|
||||
</emphasis>
|
||||
Coxae contiguous. Mesocoxae with proximodistal carina along posterior face. Metacoxae glabrous except for pubescence on internal face and around projection to trochanter. Apices of metacoxae produced into flat, blunt spines that extend posteriorly to half trochanter length (
|
||||
<figureCitation id="13692A15FFE0FFC94A50F9C5B1C7F9BE" box="[841,903,1581,1605]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="5" pageNumber="374">Fig 7</figureCitation>
|
||||
). Metatrochanters with small rhomboid-shaped tubercles on exterior face (
|
||||
<figureCitation id="13692A15FFE0FFC948D0F9B9B049F992" box="[457,521,1617,1641]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="5" pageNumber="374">Fig 7</figureCitation>
|
||||
). Femora brownish red, covered with thick, blunt golden setae. Tibia somewhat flattened in proximal half, carinate along anterior edge to half-tibia length. Tibia lighter in colour than femora, yellowish brown, with similar but shorter setae. Tarsi narrow and longer than half tibial length; also yellowish brown, with fine, sharp, aciculate setae. Protarsus with bifid anterior claw (
|
||||
<figureCitation id="13692A15FFE0FFC94AC6F954B65DF92F" box="[991,1053,1724,1748]" captionStart="FIGURES 8 – 10. 8" captionStartId="6.[151,264,1096,1119]" captionTargetBox="[163,1421,201,1057]" captionTargetId="figure@6.[157,1429,193,1075]" captionTargetPageId="6" captionText="FIGURES 8 – 10. 8) apical protarsus showing bifid anterior protarsal claw 9) aedeagus, ventral view 10) aedeagus, dorsal view." httpUri="https://zenodo.org/record/222181/files/figure.png" pageId="5" pageNumber="374">Fig 8</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BED3690FFE0FFC949D0F937B290F8C4" blockId="5.[151,1437,151,2034]" pageId="5" pageNumber="374">
|
||||
<emphasis id="B926EA82FFE0FFC949D0F937B302F903" bold="true" box="[201,322,1759,1784]" pageId="5" pageNumber="374">
|
||||
<emphasis id="B926EA82FFE0FFC949D0F937B379F903" bold="true" box="[201,313,1759,1784]" italics="true" pageId="5" pageNumber="374">Aedeagus</emphasis>
|
||||
:
|
||||
</emphasis>
|
||||
Asymmetric, broad, lacking parameres (
|
||||
<figureCitation id="13692A15FFE0FFC94A09F908B1E5F903" box="[784,933,1760,1784]" captionStart="FIGURES 8 – 10. 8" captionStartId="6.[151,264,1096,1119]" captionTargetBox="[163,1421,201,1057]" captionTargetId="figure@6.[157,1429,193,1075]" captionTargetPageId="6" captionText="FIGURES 8 – 10. 8) apical protarsus showing bifid anterior protarsal claw 9) aedeagus, ventral view 10) aedeagus, dorsal view." httpUri="https://zenodo.org/record/222181/files/figure.png" pageId="5" pageNumber="374">Fig 9 and 10</figureCitation>
|
||||
). Wide, globose basal bulb with asymmetric left and right apical projections. An asymmetric, apically-truncate sclerotised stylus emerges from within the basal bulb.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C348651BFFE0FFC949D0F8A2B295F809" pageId="5" pageNumber="374" type="biology_ecology">
|
||||
<paragraph id="8BED3690FFE0FFC949D0F8A2B295F809" blockId="5.[151,1437,151,2034]" pageId="5" pageNumber="374">
|
||||
<emphasis id="B926EA82FFE0FFC949D0F8A2B0C9F898" bold="true" box="[201,649,1866,1891]" pageId="5" pageNumber="374">Biological and collection information.</emphasis>
|
||||
The single specimen of
|
||||
<taxonomicName id="4C524D13FFE0FFC94A83F8A4B1B2F898" box="[922,1010,1868,1891]" class="Insecta" family="Silphidae" genus="Jubogaster" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="towai">
|
||||
<emphasis id="B926EA82FFE0FFC94A83F8A4B1B2F898" box="[922,1010,1868,1891]" italics="true" pageId="5" pageNumber="374">J. towai</emphasis>
|
||||
</taxonomicName>
|
||||
was collected in a colony of
|
||||
<taxonomicName id="4C524D13FFE0FFC94C21F8A3B3C4F87C" authority="Wilson" authorityName="Wilson" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="xanthogaster">
|
||||
<emphasis id="B926EA82FFE0FFC94C21F8A3B369F87C" italics="true" pageId="5" pageNumber="374">Pheidole xanthogaster</emphasis>
|
||||
Wilson
|
||||
</taxonomicName>
|
||||
under a log. The ants were apparently unaware of the beetle, which was was found deep inside the nest. This suggests that
|
||||
<taxonomicName id="4C524D13FFE0FFC94B0DF87CB02DF850" box="[532,621,1940,1963]" class="Insecta" family="Silphidae" genus="Jubogaster" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="towai">
|
||||
<emphasis id="B926EA82FFE0FFC94B0DF87CB02DF850" box="[532,621,1940,1963]" italics="true" pageId="5" pageNumber="374">J. towai</emphasis>
|
||||
</taxonomicName>
|
||||
is a true myrmecophile, but information about its biology and behaviour are unavailable at this time. It has no clear myrmecophilous adaptions, so may belong to the “obligate synoekete” class.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="DF2D6618FFE3FFCA498EFBA0B7DDFBA5" box="[151,1437,1096,1119]" httpUri="https://zenodo.org/record/222181/files/figure.png" pageId="6" pageNumber="375" targetBox="[163,1421,201,1057]" targetPageId="6">
|
||||
<paragraph id="8BED3690FFE3FFCA498EFBA0B7DDFBA5" blockId="6.[151,1437,1096,1119]" box="[151,1437,1096,1119]" pageId="6" pageNumber="375">
|
||||
<emphasis id="B926EA82FFE3FFCA498EFBA0B322FBA4" bold="true" box="[151,354,1096,1119]" pageId="6" pageNumber="375">FIGURES 8–10. 8)</emphasis>
|
||||
apical protarsus showing bifid anterior protarsal claw
|
||||
<emphasis id="B926EA82FFE3FFCA4A90FBA0B1DEFBA4" bold="true" box="[905,926,1096,1119]" pageId="6" pageNumber="375">9)</emphasis>
|
||||
aedeagus, ventral view
|
||||
<emphasis id="B926EA82FFE3FFCA4D88FBA0B6F2FBA4" bold="true" box="[1169,1202,1096,1119]" pageId="6" pageNumber="375">10)</emphasis>
|
||||
aedeagus, dorsal view.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<subSubSection id="C348651BFFE3FFCA49D0FB6DB0FDF9B5" pageId="6" pageNumber="375" type="discussion">
|
||||
<paragraph id="8BED3690FFE3FFCA49D0FB6DB0FDF9B5" blockId="6.[151,1436,1157,1615]" pageId="6" pageNumber="375">
|
||||
<emphasis id="B926EA82FFE3FFCA49D0FB6DB311FB65" bold="true" box="[201,337,1157,1182]" pageId="6" pageNumber="375">Comments.</emphasis>
|
||||
<taxonomicName id="4C524D13FFE3FFCA4841FB6FB3F2FB65" box="[344,434,1159,1182]" class="Insecta" family="Silphidae" genus="Jubogaster" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="species" species="towai">
|
||||
<emphasis id="B926EA82FFE3FFCA4841FB6FB3F2FB65" box="[344,434,1159,1182]" italics="true" pageId="6" pageNumber="375">J. towai</emphasis>
|
||||
</taxonomicName>
|
||||
has distinctly internalised terminal tergites and sternites. The plesiomorphic condition in
|
||||
<taxonomicName id="4C524D13FFE3FFCA498EFB41B35DFB3A" box="[151,285,1193,1217]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="subFamily" subFamily="Pselaphinae">Pselaphinae</taxonomicName>
|
||||
is for visible sternite 7 (IX) to form a penial plate, flanked by left and right sclerites formed from visible tergite 6 (which is also segment IX). However, in
|
||||
<taxonomicName id="4C524D13FFE3FFCA4A26FB27B1DBFB1D" box="[831,923,1231,1254]" class="Insecta" family="Silphidae" genus="Jubogaster" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="species" species="towai">
|
||||
<emphasis id="B926EA82FFE3FFCA4A26FB27B1DBFB1D" box="[831,923,1231,1254]" italics="true" pageId="6" pageNumber="375">J. towai</emphasis>
|
||||
</taxonomicName>
|
||||
, segment IX is internalized; in figure 2 the internal sclerites can be seen enclosed within the genital aperture. This arrangement makes segment VIII (visible sternite 6 and visible tergite 5) the posterior-most external segment. Don Chandler (University of New Hampshire) has kindly surveyed the character state of segment 7 (XI) and found internalisation to be relatively common in
|
||||
<taxonomicName id="4C524D13FFE3FFCA498EFAB6B357FA8D" box="[151,279,1374,1398]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="tribe" tribe="Trogastrini">Trogastrini</taxonomicName>
|
||||
: in many genera the segment is not externally visibile (
|
||||
<taxonomicName id="4C524D13FFE3FFCA4A94FAB6B60FFA8D" authority="Raffray" authorityName="Raffray" box="[909,1103,1374,1398]" class="Insecta" family="Staphylinidae" genus="Rhexinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B926EA82FFE3FFCA4A94FAB6B1B1FA8D" box="[909,1009,1374,1398]" italics="true" pageId="6" pageNumber="375">Rhexinia</emphasis>
|
||||
Raffray
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C524D13FFE3FFCA4D45FAB6B750FA8D" authority="Raffray" authorityName="Raffray" box="[1116,1296,1374,1398]" class="Insecta" family="Staphylinidae" genus="Xerhius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B926EA82FFE3FFCA4D45FAB6B6F3FA8D" box="[1116,1203,1374,1398]" italics="true" pageId="6" pageNumber="375">Xerhius</emphasis>
|
||||
Raffray
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C524D13FFE3FFCA4C04FAB6B2ADFA62" authority="Raffray" authorityName="Raffray" class="Insecta" family="Staphylinidae" genus="Aporhexius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B926EA82FFE3FFCA4C04FAB6B7DCFA8D" box="[1309,1436,1374,1398]" italics="true" pageId="6" pageNumber="375">Aporhexius</emphasis>
|
||||
Raffray
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C524D13FFE3FFCA49E2FA6AB381FA62" authority="LeConte" authorityName="LeConte" box="[251,449,1409,1434]" class="Insecta" family="Staphylinidae" genus="Rhexius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B926EA82FFE3FFCA49E2FA6AB314FA61" box="[251,340,1410,1434]" italics="true" pageId="6" pageNumber="375">Rhexius</emphasis>
|
||||
LeConte
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C524D13FFE3FFCA48D7FA6AB0CDFA62" authority="Sharp" authorityName="Sharp" box="[462,653,1409,1434]" class="Insecta" family="Staphylinidae" genus="Eurhexius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B926EA82FFE3FFCA48D7FA6AB07FFA61" box="[462,575,1410,1434]" italics="true" pageId="6" pageNumber="375">Eurhexius</emphasis>
|
||||
Sharp
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C524D13FFE3FFCA4B82FA6AB1CDFA62" authority="Chandler" authorityName="Chandler" box="[667,909,1409,1434]" class="Insecta" family="Staphylinidae" genus="Platomesus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B926EA82FFE3FFCA4B82FA6AB15CFA61" box="[667,796,1410,1434]" italics="true" pageId="6" pageNumber="375">Platomesus</emphasis>
|
||||
Chandler
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C524D13FFE3FFCA4A85FA6AB6EAFA62" authority="Chandler" authorityName="Chandler" box="[924,1194,1409,1434]" class="Insecta" family="Staphylinidae" genus="Austrorhexius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B926EA82FFE3FFCA4A85FA6AB678FA61" box="[924,1080,1410,1434]" italics="true" pageId="6" pageNumber="375">Austrorhexius</emphasis>
|
||||
Chandler
|
||||
</taxonomicName>
|
||||
) while in others it is clearly present (
|
||||
<taxonomicName id="4C524D13FFE3FFCA484DFA4EB17DFA45" authority="Brendel, Fletcherhexius Park" authorityName="Brendel, Fletcherhexius Park" box="[340,829,1446,1470]" class="Insecta" family="Staphylinidae" genus="Conoplectus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B926EA82FFE3FFCA484DFA4EB3A0FA45" box="[340,480,1446,1470]" italics="true" pageId="6" pageNumber="375">Conoplectus</emphasis>
|
||||
Brendel,
|
||||
<emphasis id="B926EA82FFE3FFCA4B4CFA4EB0BBFA45" box="[597,763,1446,1470]" italics="true" pageId="6" pageNumber="375">Fletcherhexius</emphasis>
|
||||
Park
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C524D13FFE3FFCA4A55FA4EB66EFA45" authority="Raffray" authorityName="Raffray" box="[844,1070,1446,1470]" class="Insecta" family="Staphylinidae" genus="Adrogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B926EA82FFE3FFCA4A55FA4EB189FA45" box="[844,969,1446,1470]" italics="true" pageId="6" pageNumber="375">Adrogaster</emphasis>
|
||||
Raffray
|
||||
</taxonomicName>
|
||||
). In still others, the segment is visible but thin (
|
||||
<taxonomicName id="4C524D13FFE3FFCA484EFA22B07FFA1A" authority="Raffray" authorityName="Raffray" box="[343,575,1481,1506]" class="Insecta" family="Staphylinidae" genus="Anarmodius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B926EA82FFE3FFCA484EFA22B3A0FA19" box="[343,480,1482,1506]" italics="true" pageId="6" pageNumber="375">Anarmodius</emphasis>
|
||||
Raffray
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C524D13FFE3FFCA4B54FA22B14EFA1A" authority="Casey" authorityName="Casey" box="[589,782,1481,1506]" class="Insecta" family="Staphylinidae" genus="Rhexidius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B926EA82FFE3FFCA4B54FA22B0FBFA19" box="[589,699,1482,1506]" italics="true" pageId="6" pageNumber="375">Rhexidius</emphasis>
|
||||
Casey
|
||||
</taxonomicName>
|
||||
). Two genera,
|
||||
<taxonomicName id="4C524D13FFE3FFCA4AA3FA22B64DFA1A" box="[954,1037,1482,1505]" class="Insecta" family="Staphylinidae" genus="Oropus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B926EA82FFE3FFCA4AA3FA22B64DFA1A" box="[954,1037,1482,1505]" italics="true" pageId="6" pageNumber="375">Oropus</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C524D13FFE3FFCA4D51FA22B689FA19" box="[1096,1225,1482,1506]" class="Insecta" family="Staphylinidae" genus="Mesoplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B926EA82FFE3FFCA4D51FA22B689FA19" box="[1096,1225,1482,1506]" italics="true" pageId="6" pageNumber="375">Mesoplatus</emphasis>
|
||||
</taxonomicName>
|
||||
, are polymorphic, with species falling into two or all of the three classes. Furthermore, in most Jubini and some Goniaceritae, segment 7 is not externally apparent (D. S. Chandler,
|
||||
<emphasis id="B926EA82FFE3FFCA4BF4F9FBB127F9D1" box="[749,871,1555,1578]" italics="true" pageId="6" pageNumber="375">pers comm</emphasis>
|
||||
.). Hence, a recurring trend is for internalization of the terminal segment, as exemplified by
|
||||
<taxonomicName id="4C524D13FFE3FFCA4B47F9DFB0F6F9B5" box="[606,694,1591,1614]" class="Insecta" family="Silphidae" genus="Jubogaster" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="species" species="towai">
|
||||
<emphasis id="B926EA82FFE3FFCA4B47F9DFB0F6F9B5" box="[606,694,1591,1614]" italics="true" pageId="6" pageNumber="375">J. towai</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
212
data/03/FB/87/03FB8786FFE7FFCF498EFAC6B0DBFA83.xml
Normal file
212
data/03/FB/87/03FB8786FFE7FFCF498EFAC6B0DBFA83.xml
Normal file
|
|
@ -0,0 +1,212 @@
|
|||
<document id="7C25F8BAC4FF4BF575917C5A780CEA06" ID-DOI="10.11646/zootaxa.3630.2.11" ID-ISSN="1175-5326" ID-Zenodo-Dep="222179" ID-ZooBank="70C11335-A5F6-4BCB-98E6-4D14EB1A4015" IM.bibliography_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1460584307806" checkinUser="plazi" docAuthor="Parker, Joseph & Maruyama, Munetoshi" docDate="2013" docId="03FB8786FFE7FFCF498EFAC6B0DBFA83" docLanguage="en" docName="zt03630p378.pdf" docOrigin="Zootaxa 3630 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Jubogaster Parker & Maruyama 2013, gen. nov." docType="treatment" docUuid="C4617491-F55F-4757-9189-A09F56E27AE2" docUuidSource="ZooBank" docVersion="2" lastPageNumber="372" masterDocId="FFC2FFFEFFE5FFCC4919FFE8B240FFFB" masterDocTitle="Jubogaster towai, a new Neotropical genus and species of Trogastrini (Coleoptera: Staphylinidae: Pselaphinae) exhibiting myrmecophily and extreme body enlargement" masterLastPageNumber="378" masterPageNumber="369" pageNumber="371" updateTime="1730809485467" updateUser="ExternalLinkService">
|
||||
<mods:mods id="766E5D489533B18820D68CAAB346E789" xmlns:mods="http://www.loc.gov/mods/v3">
|
||||
<mods:titleInfo id="7372C62D664B19287054F1F96E59779B">
|
||||
<mods:title id="680FA9A8B06D22FDAEA5BBD120DD44D3">Jubogaster towai, a new Neotropical genus and species of Trogastrini (Coleoptera: Staphylinidae: Pselaphinae) exhibiting myrmecophily and extreme body enlargement</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:name id="8F21DDD9A473EF4A58701A5D17736821" type="personal">
|
||||
<mods:role id="31024DFE86775633296CF74C227F0B07">
|
||||
<mods:roleTerm id="0FBF976F511936D04928DA4E786CE0DD">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="64B214466E52A94CD3DB5BAB37058EBF">Parker, Joseph</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name id="D3B5833A8C2FA45DDE448AB355DC059C" type="personal">
|
||||
<mods:role id="37862A7445129EB446A3FFD460F10E11">
|
||||
<mods:roleTerm id="52238953EC4575290058AC814022CA0C">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="3B236B6E510A66B50AAEDD4CA50FBC5C">Maruyama, Munetoshi</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:typeOfResource id="DD5D754FB376479BEEDB4CD37673FAA9">text</mods:typeOfResource>
|
||||
<mods:relatedItem id="F46411A677BC253F744D9860B52E8A8A" type="host">
|
||||
<mods:titleInfo id="C19FFC065DAF749135AEE167E4768E9E">
|
||||
<mods:title id="200A845623AF8D03E45AE935F3E7D4E4">Zootaxa</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:part id="B4EF0C898543A27D53167E8307798BBF">
|
||||
<mods:date id="BB173061770AA8C5D44DA6BC3E7C5F5F">2013</mods:date>
|
||||
<mods:detail id="4D3E7D2E5AD58B3E132B8733C8390575" type="pubDate">
|
||||
<mods:number id="23E59DDEA809EC025246B48729CA4973">2013-03-20</mods:number>
|
||||
</mods:detail>
|
||||
<mods:detail id="71AF65D12D9A2DBDE5BE38FA9641FFBB" type="volume">
|
||||
<mods:number id="E8EF333DA449E3C6D20654859F4EE278">3630</mods:number>
|
||||
</mods:detail>
|
||||
<mods:detail id="FC1FB826C0EAE452B3ADB3D951D53F15" type="issue">
|
||||
<mods:number id="1A4E571957E220D977E79DA3073F877C">2</mods:number>
|
||||
</mods:detail>
|
||||
<mods:extent id="FFD8E7456C2618EAF8B15FC3D1764E93" unit="page">
|
||||
<mods:start id="32E5E1A5355A342242BE184B6CBEF3AB">369</mods:start>
|
||||
<mods:end id="6A7EAF4241F48FF0269BD0710FFE44D2">378</mods:end>
|
||||
</mods:extent>
|
||||
</mods:part>
|
||||
</mods:relatedItem>
|
||||
<mods:classification id="EE490EC8DB68D658B9B1C1CF4914AC34">journal article</mods:classification>
|
||||
<mods:identifier id="F42A4DFEBBCD82FB970B889B50481107" type="DOI">10.11646/zootaxa.3630.2.11</mods:identifier>
|
||||
<mods:identifier id="A05A61D4F58B93EE030DF7B2E30970AE" type="ISSN">1175-5326</mods:identifier>
|
||||
<mods:identifier id="93C0BE10A6E3044590913D97C3735905" type="Zenodo-Dep">222179</mods:identifier>
|
||||
<mods:identifier id="E38D2BE2B47E964C88FD0BD66F52D90B" type="ZooBank">70C11335-A5F6-4BCB-98E6-4D14EB1A4015</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment id="03FB8786FFE7FFCF498EFAC6B0DBFA83" ID-DOI="http://doi.org/10.5281/zenodo.14040089" ID-Zenodo-Dep="14040089" LSID="urn:lsid:zoobank.org:act:C4617491-F55F-4757-9189-A09F56E27AE2" httpUri="http://treatment.plazi.org/id/03FB8786FFE7FFCF498EFAC6B0DBFA83" lastPageId="3" lastPageNumber="372" pageId="2" pageNumber="371">
|
||||
<subSubSection id="C348651BFFE7FFCE498EFAC6B3C6FABC" box="[151,390,1326,1351]" pageId="2" pageNumber="371" type="nomenclature">
|
||||
<paragraph id="8BED3690FFE7FFCE498EFAC6B3C6FABC" blockId="2.[151,390,1326,1351]" box="[151,390,1326,1351]" pageId="2" pageNumber="371">
|
||||
<emphasis id="B926EA82FFE7FFCE498EFAC6B3C6FABC" bold="true" box="[151,390,1326,1351]" pageId="2" pageNumber="371">
|
||||
<taxonomicName id="4C524D13FFE7FFCE498EFAC6B357FABC" authority="Parker & Maruyama, 2013" authorityName="Parker & Maruyama" authorityYear="2013" box="[151,279,1326,1351]" class="Insecta" family="Silphidae" genus="Jubogaster" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B926EA82FFE7FFCE498EFAC6B357FABC" bold="true" box="[151,279,1326,1351]" italics="true" pageId="2" pageNumber="371">Jubogaster</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A21557F9FFE7FFCE4804FAC7B3C6FABC" box="[285,390,1327,1351]" pageId="2" pageNumber="371" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C348651BFFE7FFCE498EFA98B155FA7C" box="[151,789,1391,1416]" pageId="2" pageNumber="371" type="type_taxon">
|
||||
<paragraph id="8BED3690FFE7FFCE498EFA98B155FA7C" blockId="2.[151,1436,1391,2028]" box="[151,789,1391,1416]" pageId="2" pageNumber="371">
|
||||
<emphasis id="B926EA82FFE7FFCE498EFA98B372FA73" bold="true" box="[151,306,1391,1416]" pageId="2" pageNumber="371">
|
||||
<typeStatus id="54E98832FFE7FFCE498EFA98B292FA73" box="[151,210,1392,1416]" pageId="2" pageNumber="371">Type</typeStatus>
|
||||
species.
|
||||
</emphasis>
|
||||
<taxonomicName id="4C524D13FFE7FFCE4820FA98B3BAFA73" box="[313,506,1392,1416]" class="Insecta" family="Silphidae" genus="Jubogaster" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="towai" status="sp. nov.">
|
||||
<emphasis id="B926EA82FFE7FFCE4820FA98B3BAFA73" box="[313,506,1392,1416]" italics="true" pageId="2" pageNumber="371">Jubogaster towai</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A21557F9FFE7FFCE4B18FA98B018FA7C" box="[513,600,1392,1415]" pageId="2" pageNumber="371" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
here designated.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C348651BFFE7FFCE498EFA7BB2B6F90B" pageId="2" pageNumber="371" type="diagnosis">
|
||||
<paragraph id="8BED3690FFE7FFCE498EFA7BB2B6F90B" blockId="2.[151,1436,1391,2028]" pageId="2" pageNumber="371">
|
||||
<emphasis id="B926EA82FFE7FFCE498EFA7BB351FA57" bold="true" box="[151,273,1427,1452]" pageId="2" pageNumber="371">Diagnosis.</emphasis>
|
||||
Euplectite pselaphines with the following characters: transverse head articulating ventrally under pronotum when deflexed, typical of
|
||||
<taxonomicName id="4C524D13FFE7FFCE4B21FA5FB0F7FA34" box="[568,695,1463,1487]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="tribe" tribe="Trogastrini">Trogastrini</taxonomicName>
|
||||
. Head with vertex excavate, lacking vertexal fovea; ventral head without the typical dense “genal beard” of setae common to most
|
||||
<taxonomicName id="4C524D13FFE7FFCE4AADFA34B673FA0F" box="[948,1075,1500,1524]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="tribe" tribe="Trogastrini">Trogastrini</taxonomicName>
|
||||
. Eleven-segmented moderately geniculate antennae, received by large excavations anterior to eyes. Pronotum jubine-like: broad and obcordate, constricted at basal quarter, with weakly-evident medial longitudinal sulcus and distinct antebasal sulcus. Pronotum afoveate, with margins smooth, devoid of crenulations or small spines found in other
|
||||
<taxonomicName id="4C524D13FFE7FFCE4DD7F9AFB70DF9A4" box="[1230,1357,1607,1631]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="tribe" tribe="Trogastrini">Trogastrini</taxonomicName>
|
||||
. Elytra lacking any stereotypical fovea. Legs with contiguous, projecting coxae, broad femora, basally flattened tibia, and three tarsal segments; segment 1 very small, 2 and 3 elongate and subequal in length. Tarsi with two equally-sized claws, unlike other
|
||||
<taxonomicName id="4C524D13FFE7FFCE486DF95CB3AFF937" box="[372,495,1716,1740]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="tribe" tribe="Trogastrini">Trogastrini</taxonomicName>
|
||||
which have unequally-sized claws. Bifid anterior protarsal claw in the
|
||||
<typeStatus id="54E98832FFE7FFCE4C0AF95DB703F936" box="[1299,1347,1717,1741]" pageId="2" pageNumber="371">type</typeStatus>
|
||||
species
|
||||
<taxonomicName id="4C524D13FFE7FFCE498EF931B2AFF90B" box="[151,239,1753,1776]" class="Insecta" family="Silphidae" genus="Jubogaster" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="towai">
|
||||
<emphasis id="B926EA82FFE7FFCE498EF931B2AFF90B" box="[151,239,1753,1776]" italics="true" pageId="2" pageNumber="371">J. towai</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C348651BFFE7FFCE49D0F914B1CEF85F" pageId="2" pageNumber="371" type="etymology">
|
||||
<paragraph id="8BED3690FFE7FFCE49D0F914B1CEF85F" blockId="2.[151,1436,1391,2028]" pageId="2" pageNumber="371">
|
||||
<emphasis id="B926EA82FFE7FFCE49D0F914B30FF8EF" bold="true" box="[201,335,1788,1812]" pageId="2" pageNumber="371">Etymology.</emphasis>
|
||||
A combination of
|
||||
<taxonomicName id="4C524D13FFE7FFCE4B38F914B09AF8EF" authority="Schaufuss" authorityName="Schaufuss" box="[545,730,1788,1812]" class="Insecta" family="Staphylinidae" genus="Jubus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B926EA82FFE7FFCE4B38F914B022F8EF" box="[545,610,1788,1812]" italics="true" pageId="2" pageNumber="371">Jubus</emphasis>
|
||||
Schaufuss
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C524D13FFE7FFCE4A09F915B186F8EF" authority="Sharp" authorityName="Sharp" box="[784,966,1788,1812]" class="Insecta" family="Staphylinidae" genus="Trogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B926EA82FFE7FFCE4A09F915B13CF8EF" box="[784,892,1789,1812]" italics="true" pageId="2" pageNumber="371">Trogaster</emphasis>
|
||||
Sharp
|
||||
</taxonomicName>
|
||||
, the
|
||||
<typeStatus id="54E98832FFE7FFCE4AE3F915B66AF8EE" box="[1018,1066,1789,1813]" pageId="2" pageNumber="371">type</typeStatus>
|
||||
genera of Jubini and
|
||||
<taxonomicName id="4C524D13FFE7FFCE4C03F914B7D8F8EF" box="[1306,1432,1788,1812]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="tribe" tribe="Trogastrini">Trogastrini</taxonomicName>
|
||||
, respectively. The name is chosen on account of the membership of the new genus in
|
||||
<taxonomicName id="4C524D13FFE7FFCE4D43F8F7B698F8CC" box="[1114,1240,1823,1847]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="tribe" tribe="Trogastrini">Trogastrini</taxonomicName>
|
||||
, despite apparent Jubini-like characters such as the shape of the pronotum and the presence of two equally-sized tarsal claws (a character state found in Jubini, but not other Euplectitae). The bifid anterior protarsal claw of the
|
||||
<typeStatus id="54E98832FFE7FFCE4DEFF881B766F87A" box="[1270,1318,1897,1921]" pageId="2" pageNumber="371">type</typeStatus>
|
||||
species
|
||||
<taxonomicName id="4C524D13FFE7FFCE4C91F881B295F85F" class="Insecta" family="Silphidae" genus="Jubogaster" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="towai">
|
||||
<emphasis id="B926EA82FFE7FFCE4C91F881B295F85F" italics="true" pageId="2" pageNumber="371">J. towai</emphasis>
|
||||
</taxonomicName>
|
||||
is also shared with
|
||||
<taxonomicName id="4C524D13FFE7FFCE48AAF864B15EF85F" authority="Silvestri" authorityName="Silvestri" box="[435,798,1932,1956]" class="Insecta" family="Staphylinidae" genus="Phamisus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="myrmophilus">
|
||||
<emphasis id="B926EA82FFE7FFCE48AAF864B0F8F85F" box="[435,696,1932,1956]" italics="true" pageId="2" pageNumber="371">Phamisus myrmophilus</emphasis>
|
||||
Silvestri
|
||||
</taxonomicName>
|
||||
of Jubini.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C348651BFFE7FFCF49D0F847B0DBFA83" lastPageId="3" lastPageNumber="372" pageId="2" pageNumber="371" type="description">
|
||||
<paragraph id="8BED3690FFE7FFCE49D0F847B388F817" blockId="2.[151,1436,1391,2028]" pageId="2" pageNumber="371">
|
||||
<emphasis id="B926EA82FFE7FFCE49D0F847B31AF833" bold="true" box="[201,346,1967,1992]" pageId="2" pageNumber="371">Description.</emphasis>
|
||||
Body large (
|
||||
<quantity id="4CAA9B75FFE7FFCE48F4F858B07EF833" box="[493,574,1968,1992]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="2" pageNumber="371" unit="mm" value="5.1">5.1mm</quantity>
|
||||
for the
|
||||
<typeStatus id="54E98832FFE7FFCE4B8EF859B087F832" box="[663,711,1969,1993]" pageId="2" pageNumber="371">type</typeStatus>
|
||||
species
|
||||
<taxonomicName id="4C524D13FFE7FFCE4A31F859B1C1F833" box="[808,897,1969,1992]" class="Insecta" family="Silphidae" genus="Jubogaster" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="towai">
|
||||
<emphasis id="B926EA82FFE7FFCE4A31F859B1C1F833" box="[808,897,1969,1992]" italics="true" pageId="2" pageNumber="371">J. towai</emphasis>
|
||||
</taxonomicName>
|
||||
shown in
|
||||
<figureCitation id="13692A15FFE7FFCE4AEEF858B615F833" box="[1015,1109,1968,1992]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="2" pageNumber="371">Figure 1</figureCitation>
|
||||
). Body shape approximately ovoid and strongly convex.
|
||||
</paragraph>
|
||||
<paragraph id="8BED3690FFE6FFCF49D0FF7FB63FFE0F" blockId="3.[151,1437,151,1401]" pageId="3" pageNumber="372">
|
||||
<emphasis id="B926EA82FFE6FFCF49D0FF7FB350FF4B" bold="true" box="[201,272,151,176]" italics="true" pageId="3" pageNumber="372">Head:</emphasis>
|
||||
Approximately triangular, strongly transverse (
|
||||
<figureCitation id="13692A15FFE6FFCF4A30FF7FB129FF54" box="[809,873,151,175]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="3" pageNumber="372">Fig 4</figureCitation>
|
||||
). Vertex afoveate, concave, with moderately deep excavation extending from basal quarter to anterior margin of frons; excavation narrowing apically along midline as it passes between raised frontolateral margins. Frontolateral margins carinate, extending roughly arcuately from posterior of antennal insertion cavity to clypeus (
|
||||
<figureCitation id="13692A15FFE6FFCF4BD7FEECB14FFEE7" box="[718,783,260,284]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="3" pageNumber="372">Fig 5</figureCitation>
|
||||
). Frontolateral face accommodating large circular cavity anterior to eyes to receive antennae (
|
||||
<figureCitation id="13692A15FFE6FFCF4B50FECFB0CBFEC4" box="[585,651,295,319]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="3" pageNumber="372">Fig 5</figureCitation>
|
||||
). Eye flanked directly anteriorly by additional large excavation for retracted maxillary palpus (
|
||||
<figureCitation id="13692A15FFE6FFCF48CAFEA4B054FE9F" box="[467,532,332,356]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="3" pageNumber="372">Fig 5</figureCitation>
|
||||
). Ventral surface of head bisected by longitudinal gular carina situated within broader gular sulcus terminating in gular fovea. Gular sulcus widens basally, spanning the head-neck interface. Maxillary palpi apparently 4-segmented, with segment II long and curved at apex, III short and triangular, wider than II, IV oval, approximately
|
||||
<date id="FFEC1050FFE6FFCF4B18FE5FB05DFE34" box="[513,541,439,463]" pageId="3" pageNumber="372">2x</date>
|
||||
longer than wide (
|
||||
<figureCitation id="13692A15FFE6FFCF4BECFE5FB174FE34" box="[757,820,439,463]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="3" pageNumber="372">Fig 5</figureCitation>
|
||||
). Antennae (
|
||||
<figureCitation id="13692A15FFE6FFCF4ADFFE5FB643FE34" box="[966,1027,439,463]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="3" pageNumber="372">Fig 3</figureCitation>
|
||||
) with 11 antennomeres, moderately geniculate with enlarged segment I. Antennomeres IX
|
||||
<emphasis id="B926EA82FFE6FFCF4BEDFE34B142FE0F" bold="true" box="[756,770,476,500]" pageId="3" pageNumber="372">–</emphasis>
|
||||
XI strongly elongate (XI longest).
|
||||
</paragraph>
|
||||
<paragraph id="8BED3690FFE6FFCF49D0FE17B10CFCCC" blockId="3.[151,1437,151,1401]" pageId="3" pageNumber="372">
|
||||
<emphasis id="B926EA82FFE6FFCF49D0FE17B368FDE3" bold="true" box="[201,296,511,536]" italics="true" pageId="3" pageNumber="372">Thorax:</emphasis>
|
||||
Pronotum (
|
||||
<figureCitation id="13692A15FFE6FFCF48A8FE17B3B0FDEC" box="[433,496,511,535]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="3" pageNumber="372">Fig 4</figureCitation>
|
||||
) obcordate,
|
||||
<date id="FFEC1050FFE6FFCF4B98FE17B0DDFDEC" box="[641,669,511,535]" pageId="3" pageNumber="372">2x</date>
|
||||
wider than head, almost as wide as elytra. Afoveate, with smooth margins. Anterior margin broadly arcuate, largely obscuring the deflexed head when viewed from above, and constricted sharply in basal quarter, so that pronotum is laterally expanded into broad, blunt spurs with curved edges. Medial longitudinal sulcus present on pronotal disc, weakly impressed, thin, terminating at deeper, bowshaped antebasal sulcus that extends between bases of mediolateral spurs. Prosternum afoveate, with lateral longitudinal carinae extending from sides of neck to base. Mesosternum with median mesosternal foveae (MMSF), lateral mesosternal fovea (LMSF) and lateral mesocoxal fovea (LMCF) present. Other typical pselaphine mesosternal fovea listed in
|
||||
<bibRefCitation id="EFC34B61FFE6FFCF48CDFD14B0CFFCEF" author="Chandler, D. S." box="[468,655,764,788]" journalOrPublisher="Memoirs on Entomology International" pageId="3" pageNumber="372" pagination="1 - 560" part="15" refId="ref6070" refString="Chandler, D. S. (2001) Biology, morphology, and systematics of the ant-like litter beetles of Australia (Coleoptera: Staphylinidae: Pselaphinae). Memoirs on Entomology International, 15, 1 - 560." title="Biology, morphology, and systematics of the ant-like litter beetles of Australia (Coleoptera: Staphylinidae: Pselaphinae)" type="journal article" year="2001">Chandler (2001)</bibRefCitation>
|
||||
lacking. Metasternum strongly convex. Lateral metathoracic foveae (LMTF) present; median metathoracic fovea (MMTF) absent.
|
||||
</paragraph>
|
||||
<paragraph id="8BED3690FFE6FFCF49D0FCABB646FC17" blockId="3.[151,1437,151,1401]" pageId="3" pageNumber="372">
|
||||
<emphasis id="B926EA82FFE6FFCF49D0FCABB301FCA7" bold="true" box="[201,321,835,860]" italics="true" pageId="3" pageNumber="372">Abdomen:</emphasis>
|
||||
Abdomen with tergites strongly convex. Visible segments 1
|
||||
<emphasis id="B926EA82FFE6FFCF4AF3FCACB1B8FCA7" bold="true" box="[1002,1016,836,860]" pageId="3" pageNumber="372">–</emphasis>
|
||||
3 (IV
|
||||
<emphasis id="B926EA82FFE6FFCF4D2DFCACB602FCA7" bold="true" box="[1076,1090,836,860]" pageId="3" pageNumber="372">–</emphasis>
|
||||
VI) subequal in length (
|
||||
<figureCitation id="13692A15FFE6FFCF4C57FCACB7CAFCA7" box="[1358,1418,836,860]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="3" pageNumber="372">Fig 1</figureCitation>
|
||||
), segment 4 (VII) longest (not apparent in
|
||||
<figureCitation id="13692A15FFE6FFCF4B8AFC8FB095FC84" box="[659,725,871,895]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="3" pageNumber="372">Fig 1</figureCitation>
|
||||
due to curvature of abdomen). Tergites 1
|
||||
<emphasis id="B926EA82FFE6FFCF4DCFFC80B6A4FC7B" bold="true" box="[1238,1252,872,896]" pageId="3" pageNumber="372">–</emphasis>
|
||||
3 (IV
|
||||
<emphasis id="B926EA82FFE6FFCF4C3EFC80B775FC7B" bold="true" box="[1319,1333,872,896]" pageId="3" pageNumber="372">–</emphasis>
|
||||
VI) with pronounced paratergites, 4 (VII) with small tapering paratergites. Tergite 6 (IX) internalised. Six sternites visible (
|
||||
<figureCitation id="13692A15FFE6FFCF4986FC47B29EFC3C" box="[159,222,943,967]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="3" pageNumber="372">Fig 2</figureCitation>
|
||||
), sternite 7 (IX) internalised. First visible sternite (III) with median process projecting between metacoxae. Sternites 2 and 3 (IV and V) subequal in length, 4 (VI) slightly shorter (
|
||||
<figureCitation id="13692A15FFE6FFCF4AA0FC3CB1B8FC17" box="[953,1016,980,1004]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="3" pageNumber="372">Fig 2</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BED3690FFE6FFCF49D0FC1FB723FB5B" blockId="3.[151,1437,151,1401]" pageId="3" pageNumber="372">
|
||||
<emphasis id="B926EA82FFE6FFCF49D0FC1FB35AFBEB" bold="true" box="[201,282,1015,1040]" italics="true" pageId="3" pageNumber="372">Elytra:</emphasis>
|
||||
Strongly convex, slightly broader than pronotum (
|
||||
<figureCitation id="13692A15FFE6FFCF4A70FC1FB1E7FBF4" box="[873,935,1015,1039]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="3" pageNumber="372">Fig 1</figureCitation>
|
||||
), approximately 0.3x total body length. At their longest, equal in length to visible abdominal segments 2+3 (V+VI) combined. Elytral humeri broadly rounded to almost half elytral length. Sutural stria entire; lacking sutural fovea and other stereotypical antebasal fovea. Lacking subhumeral fovea and subhumeral stria. One weakly defined, somewhat discal depression is evident halfway between suture and humerus; its homology to canonical pselaphine antebasal elytral fovea is unclear.
|
||||
</paragraph>
|
||||
<paragraph id="8BED3690FFE6FFCF49D0FB44B0DBFA83" blockId="3.[151,1437,151,1401]" pageId="3" pageNumber="372">
|
||||
<emphasis id="B926EA82FFE6FFCF49D0FB44B349FB38" bold="true" box="[201,265,1196,1219]" italics="true" pageId="3" pageNumber="372">Legs:</emphasis>
|
||||
All coxae contiguous (
|
||||
<figureCitation id="13692A15FFE6FFCF4B03FB44B01BFB3F" box="[538,603,1196,1220]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="3" pageNumber="372">Fig 2</figureCitation>
|
||||
), procoxae conical, projecting ventrally, mesocoxae globose, metacoxae (
|
||||
<figureCitation id="13692A15FFE6FFCF4987FB38B29BFB13" box="[158,219,1232,1256]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="3" pageNumber="372">Fig 7</figureCitation>
|
||||
) transverse-conical, spanning from ventral midline to metasternal margin, and projecting posteriorly. Femora somewhat thickened, and excavated apically to receive retracted tibia. Tibia flattened basally. Femora and tibia lacking spines or processes. Tarsi elongate, 3-segmented, first segment extremely short, 0.2x the length of segment 2. Segment 2 and 3 subequal in length. All tarsi with 2 well-developed, equally-sized tarsal claws. Bifid anterior protarsal claw in
|
||||
<typeStatus id="54E98832FFE6FFCF4841FA89B3C8FA82" box="[344,392,1377,1401]" pageId="3" pageNumber="372">type</typeStatus>
|
||||
species
|
||||
<taxonomicName id="4C524D13FFE6FFCF48FEFA89B000FA83" box="[487,576,1377,1400]" class="Insecta" family="Silphidae" genus="Jubogaster" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="species" species="towai">
|
||||
<emphasis id="B926EA82FFE6FFCF48FEFA89B000FA83" box="[487,576,1377,1400]" italics="true" pageId="3" pageNumber="372">J. towai</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="13692A15FFE6FFCF4B56FA88B0CDFA83" box="[591,653,1376,1400]" captionStart="FIGURES 8 – 10. 8" captionStartId="6.[151,264,1096,1119]" captionTargetBox="[163,1421,201,1057]" captionTargetId="figure@6.[157,1429,193,1075]" captionTargetPageId="6" captionText="FIGURES 8 – 10. 8) apical protarsus showing bifid anterior protarsal claw 9) aedeagus, ventral view 10) aedeagus, dorsal view." httpUri="https://zenodo.org/record/222181/files/figure.png" pageId="3" pageNumber="372">Fig 8</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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Reference in a new issue