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<document id="04D8C6231125B07364ECDA8BC7F2FE7D" ID-DOI="10.11646/zootaxa.5537.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="14239861" ID-ZooBank="6872E03D-49FF-44A0-82E9-AF7AB4407DDC" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1732801203152" checkinUser="plazi" docAuthor="Trýzna, Miloš &amp; Rakotonirina, Jean Claude" docDate="2024" docId="03FA87B21B00DE3A479E598BD672FBB8" docLanguage="en" docName="zootaxa.5537.3.2.pdf" docOrigin="Zootaxa 5537 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5537.3.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Apatenia Pascoe 1859" docType="key" docVersion="3" lastPageNumber="340" masterDocId="FFC3FFCA1B01DE3847095E15D413FF92" masterDocTitle="A new species of Apatenia Pascoe (Coleoptera: Anthribidae) from central Madagascar, with a key to species, additions to the distribution of rare species of the genus, and general notes about threats to anthribid diversity" masterLastPageNumber="347" masterPageNumber="325" pageNumber="326" updateTime="1733316570899" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:title id="A32D0237F686DE95D2B77499CA9CE65C">A new species of Apatenia Pascoe (Coleoptera: Anthribidae) from central Madagascar, with a key to species, additions to the distribution of rare species of the genus, and general notes about threats to anthribid diversity</mods:title>
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<mods:namePart id="696009CA02783479CBA4DE1BDAA91DA1">Trýzna, Miloš</mods:namePart>
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<mods:affiliation id="321B887DDDF03A8C0FCE15E7A76512BB">Mendel University in Brno, Faculty of Forestry and Wood Technology, Department of Forest Ecology, Zemědělská 3, CZ- 613 00 Brno, Czech Republic. &amp; Municipal Museum of Ústí nad Labem, Natural History Department, Masarykova 1000 / 3, CZ- 400 01 Ústí nad Labem, Czech Republic.</mods:affiliation>
<mods:nameIdentifier id="A40072315C04FDAAF5A18031F6D133BA" type="email">anthribidae@gmail.com</mods:nameIdentifier>
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<mods:namePart id="C33CD3952B7E2AB78CACC595D4DD9162">Rakotonirina, Jean Claude</mods:namePart>
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<mods:affiliation id="6B56F7CE926284FC7669D54428466123">Madagascar Biodiversity Center, BP 6257, Parc Botanique et Zoologique de Tsimbazaza, Antananarivo, Madagascar. &amp; Département d’Entomologie-Cultures, Elevage et Santé; Faculté des Sciences, BP 906, Université d’Antananarivo, Antananarivo, Madagascar.</mods:affiliation>
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Genus
<taxonomicName id="4C534D271B00DE3947E7598AD612F82A" ID-CoL="8HCZN" authority="Pascoe, 1859" authorityName="Pascoe" authorityYear="1859" box="[238,513,1950,1977]" class="Insecta" family="Anthribidae" genus="Apatenia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="326" phylum="Arthropoda" rank="genus">
<emphasis id="B927EAB61B00DE3947E7598AD548F82B" bold="true" box="[238,347,1951,1977]" italics="true" pageId="1" pageNumber="326">Apatenia</emphasis>
<bibRefCitation id="EFC24B551B00DE39466A598AD612F82A" author="Pascoe, F. P." box="[355,513,1950,1977]" pageId="1" pageNumber="326" pagination="431 - 439" refId="ref11869" refString="Pascoe, F. P. (1859) On some new Anthribidae. Annals and Magazine of Natural History, Series 3, 4, 431 - 439." type="journal article" year="1859">Pascoe, 1859</bibRefCitation>
</taxonomicName>
</emphasis>
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<paragraph id="8BEC36A41B03DE3A479E5E8CD04DFF23" blockId="2.[151,1436,152,238]" box="[151,1118,152,177]" pageId="2" pageNumber="327">
<typeStatus id="54E888061B03DE3A479E5E8CD4DAFF23" box="[151,201,153,177]" pageId="2" pageNumber="327">Type</typeStatus>
species:
<taxonomicName id="4C534D271B03DE3A462F5E8CD671FF22" authority="Pascoe, 1859" authorityName="Pascoe" authorityYear="1859" box="[294,610,152,176]" class="Insecta" family="Anthribidae" genus="Apatenia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="327" phylum="Arthropoda" rank="species" species="viduata">
<emphasis id="B927EAB61B03DE3A462F5E8CD5C7FF22" box="[294,468,152,176]" italics="true" pageId="2" pageNumber="327">Apatenia viduata</emphasis>
<bibRefCitation id="EFC24B551B03DE3A46D35E8CD671FF22" author="Pascoe, F. P." box="[474,610,152,176]" pageId="2" pageNumber="327" pagination="431 - 439" refId="ref11869" refString="Pascoe, F. P. (1859) On some new Anthribidae. Annals and Magazine of Natural History, Series 3, 4, 431 - 439." type="journal article" year="1859">Pascoe, 1859</bibRefCitation>
</taxonomicName>
, by original designation (described from Borneo).
</paragraph>
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<treatmentCitationGroup id="AB43118A1B03DE3A479E5EA2D7F5FF7C" pageId="2" pageNumber="327">
<treatmentCitation id="0AF210B51B03DE3A479E5EA2D602FF5D" author="Pascoe, F. P." box="[151,529,183,207]" page="434" pageId="2" pageNumber="327" year="1859">
<taxonomicName id="4C534D271B03DE3A479E5EA2D602FF5D" ID-CoL="F8Q3" authority="Pascoe, 1859: 434" authorityName="Pascoe" authorityPageNumber="434" authorityYear="1859" box="[151,529,183,207]" class="Insecta" family="Anthribidae" genus="Apatenia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="327" phylum="Arthropoda" rank="species" species="viduata">
<emphasis id="B927EAB61B03DE3A479E5EA2D554FF5D" box="[151,327,183,207]" italics="true" pageId="2" pageNumber="327">Apatenia viduata</emphasis>
<bibRefCitation id="EFC24B551B03DE3A46595EADD602FF5D" author="Pascoe, F. P." box="[336,529,183,207]" pageId="2" pageNumber="327" pagination="431 - 439" refId="ref11869" refString="Pascoe, F. P. (1859) On some new Anthribidae. Annals and Magazine of Natural History, Series 3, 4, 431 - 439." type="journal article" year="1859">Pascoe, 1859: 434</bibRefCitation>
</taxonomicName>
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. Gemminger &amp; Harold 1872: 2735 (catalogue); Bovie 1906: 257 (catalogue);
<treatmentCitation id="0AF210B51B03DE3A42485EA2D533FF7C" author="Wolfrum, P." page="41" pageId="2" pageNumber="327" year="1929">
<bibRefCitation id="EFC24B551B03DE3A42485EA2D533FF7C" author="Wolfrum, P." pageId="2" pageNumber="327" refId="ref12591" refString="Wolfrum, P. (1929) Anthribidae. In: Schenkling, S. (Ed.), Coleopterorum Catalogus auspiciis et auxilio. Vol. 102. W. Junk, Berlin, pp. 1 - 145." type="book" year="1929">Wolfrum 1929: 41</bibRefCitation>
</treatmentCitation>
(catalogue), 1953: 18 (catalogue),
<treatmentCitation id="0AF210B51B03DE3A458D5EC3D775FF7C" author="Rheinheimer, J." box="[644,870,214,238]" page="44" pageId="2" pageNumber="327" year="2004">
<bibRefCitation id="EFC24B551B03DE3A458D5EC3D775FF7C" author="Rheinheimer, J." box="[644,870,214,238]" pageId="2" pageNumber="327" pagination="1 - 243" refId="ref11978" refString="Rheinheimer, J. (2004) Illustrierter Katalog und Bibliographie der Anthribidae der Welt (Insecta: Coleoptera). Mitteilungen des Entomologischen Vereins Stuttgart, 39 (1 / 2), 1 - 243." type="journal article" year="2004">Rheinheimer 2004: 44</bibRefCitation>
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(catalogue).
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Pictures/drawings of Malagasy
<taxonomicName id="4C534D271B03DE3A46FD5F09D64AFEA4" ID-CoL="8HCZN" box="[500,601,284,310]" class="Insecta" family="Anthribidae" genus="Apatenia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="327" phylum="Arthropoda" rank="genus">
<emphasis id="B927EAB61B03DE3A46FD5F09D64AFEA4" box="[500,601,284,310]" italics="true" pageId="2" pageNumber="327">Apatenia</emphasis>
</taxonomicName>
:
<bibRefCitation id="EFC24B551B03DE3A456E5F09D708FEA5" author="Frieser, R." box="[615,795,284,311]" pageId="2" pageNumber="327" pagination="35 - 51" refId="ref10720" refString="Frieser, R. (2000) Einige neue Anthribiden von Madagaskar und der Ile de la Reunion (Coleoptera: Anthribidae). Acta Coleopterologica, 16 (1), 35 - 51." type="journal article" year="2000">Frieser 2000: 51</bibRefCitation>
(fig. 4–dorsal habitus of
<taxonomicName id="4C534D271B03DE3A43275F09D18BFEA4" ID-CoL="67KHL" authority="Frieser, 2000" authorityName="Frieser" authorityYear="2000" box="[1070,1432,284,310]" class="Insecta" family="Anthribidae" genus="Apatenia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="327" phylum="Arthropoda" rank="species" species="oculifera">
<emphasis id="B927EAB61B03DE3A43275F09D0EDFEA4" box="[1070,1278,284,310]" italics="true" pageId="2" pageNumber="327">Apatenia oculifera</emphasis>
<bibRefCitation id="EFC24B551B03DE3A420A5F09D18BFEA4" author="Frieser, R." box="[1283,1432,284,310]" pageId="2" pageNumber="327" pagination="35 - 51" refId="ref10720" refString="Frieser, R. (2000) Einige neue Anthribiden von Madagaskar und der Ile de la Reunion (Coleoptera: Anthribidae). Acta Coleopterologica, 16 (1), 35 - 51." type="journal article" year="2000">Frieser, 2000</bibRefCitation>
</taxonomicName>
; fig. 5–dorsal habitus of
<taxonomicName id="4C534D271B03DE3A46AD5F55D731FEC8" ID-CoL="F8PW" authority="Frieser, 2000" authorityName="Frieser" authorityYear="2000" box="[420,802,320,346]" class="Insecta" family="Anthribidae" genus="Apatenia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="327" phylum="Arthropoda" rank="species" species="sulcicollis">
<emphasis id="B927EAB61B03DE3A46AD5F55D690FEC8" box="[420,643,320,346]" italics="true" pageId="2" pageNumber="327">Apatenia sulcicollis</emphasis>
<bibRefCitation id="EFC24B551B03DE3A45835F55D731FEC8" author="Frieser, R." box="[650,802,320,346]" pageId="2" pageNumber="327" pagination="35 - 51" refId="ref10720" refString="Frieser, R. (2000) Einige neue Anthribiden von Madagaskar und der Ile de la Reunion (Coleoptera: Anthribidae). Acta Coleopterologica, 16 (1), 35 - 51." type="journal article" year="2000">Frieser, 2000</bibRefCitation>
</taxonomicName>
);
<bibRefCitation id="EFC24B551B03DE3A443F5F55D022FEC8" author="Rheinheimer, J." box="[822,1073,320,346]" pageId="2" pageNumber="327" pagination="1 - 243" refId="ref11978" refString="Rheinheimer, J. (2004) Illustrierter Katalog und Bibliographie der Anthribidae der Welt (Insecta: Coleoptera). Mitteilungen des Entomologischen Vereins Stuttgart, 39 (1 / 2), 1 - 243." type="journal article" year="2004">Rheinheimer 2004: 44</bibRefCitation>
(catalogue), 221 (fig. 93–dorsal and lateral habitus of
<taxonomicName id="4C534D271B03DE3A46825F71D71BFEEC" ID-CoL="F8PW" authority="Frieser, 2000" authorityName="Frieser" authorityYear="2000" box="[395,776,356,382]" class="Insecta" family="Anthribidae" genus="Apatenia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="327" phylum="Arthropoda" rank="species" species="sulcicollis">
<emphasis id="B927EAB61B03DE3A46825F71D67AFEEC" box="[395,617,356,382]" italics="true" pageId="2" pageNumber="327">Apatenia sulcicollis</emphasis>
<bibRefCitation id="EFC24B551B03DE3A45795F71D71BFEEC" author="Frieser, R." box="[624,776,356,382]" pageId="2" pageNumber="327" pagination="35 - 51" refId="ref10720" refString="Frieser, R. (2000) Einige neue Anthribiden von Madagaskar und der Ile de la Reunion (Coleoptera: Anthribidae). Acta Coleopterologica, 16 (1), 35 - 51." type="journal article" year="2000">Frieser, 2000</bibRefCitation>
</taxonomicName>
);
<bibRefCitation id="EFC24B551B03DE3A44125F71D7C7FEEC" author="Frieser, R." box="[795,980,356,382]" pageId="2" pageNumber="327" pagination="3 - 22" refId="ref10788" refString="Frieser, R. (2010) Teilergebnisse der entomologischen Expedition von Milos Tryzna auf Madagaskar in 2007 mit Genehmigung ANGAP (Coleoptera: Anthribidae). Acta Coleopterologica, 26 (1), 3 - 22." type="journal article" year="2010">Frieser 2010: 20</bibRefCitation>
(fig. 1–dorsal habitus of
<taxonomicName id="4C534D271B03DE3A43F95F71D53EFE30" ID-CoL="8K5H9" authority="Frieser, 2010" authorityName="Frieser" authorityYear="2010" class="Insecta" family="Anthribidae" genus="Apatenia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="327" phylum="Arthropoda" rank="species" species="fallax">
<emphasis id="B927EAB61B03DE3A43F95F71D18FFEEC" box="[1264,1436,356,382]" italics="true" pageId="2" pageNumber="327">Apatenia fallax</emphasis>
<bibRefCitation id="EFC24B551B03DE3A479E5F9DD53EFE30" author="Frieser, R." box="[151,301,392,418]" pageId="2" pageNumber="327" pagination="3 - 22" refId="ref10788" refString="Frieser, R. (2010) Teilergebnisse der entomologischen Expedition von Milos Tryzna auf Madagaskar in 2007 mit Genehmigung ANGAP (Coleoptera: Anthribidae). Acta Coleopterologica, 26 (1), 3 - 22." type="journal article" year="2010">Frieser, 2010</bibRefCitation>
</taxonomicName>
);
<bibRefCitation id="EFC24B551B03DE3A46375F9DD674FE31" author="Tryzna, M. &amp; Banar, P." box="[318,615,392,419]" pageId="2" pageNumber="327" pagination="504 - 512" refId="ref12329" refString="Tryzna, M. &amp; Banar, P. (2013) A new species of the genus Apatenia (Coleoptera: Anthribidae) from Madagascar with notes on female genitalia, redescription of the female of Apatenia quadristigma Frieser and list of Madagascan species. Zootaxa, 3609 (5), 504 - 512. https: // doi. org / 10.11646 / zootaxa. 3609.5.6" type="journal article" year="2013">Trýzna &amp; Baňař 2013: 506</bibRefCitation>
,
<taxonomicName id="4C534D271B03DE3A45785F9DD018FE30" ID-CoL="F8PV" authority="Tryzna &amp; Banar, 2013" authorityName="Tryzna &amp; Banar" authorityYear="2013" box="[625,1035,392,418]" class="Insecta" family="Anthribidae" genus="Apatenia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="327" phylum="Arthropoda" rank="species" species="stysi">
<emphasis id="B927EAB61B03DE3A45785F9DD71EFE30" box="[625,781,392,418]" italics="true" pageId="2" pageNumber="327">Apatenia stysi</emphasis>
<bibRefCitation id="EFC24B551B03DE3A441B5F9DD018FE30" author="Tryzna, M. &amp; Banar, P." box="[786,1035,392,418]" pageId="2" pageNumber="327" pagination="504 - 512" refId="ref12329" refString="Tryzna, M. &amp; Banar, P. (2013) A new species of the genus Apatenia (Coleoptera: Anthribidae) from Madagascar with notes on female genitalia, redescription of the female of Apatenia quadristigma Frieser and list of Madagascan species. Zootaxa, 3609 (5), 504 - 512. https: // doi. org / 10.11646 / zootaxa. 3609.5.6" type="journal article" year="2013">Trýzna &amp; Baňař, 2013</bibRefCitation>
</taxonomicName>
, figs 1–4, female
<typeStatus id="54E888061B03DE3A43C25F9DD13EFE30" box="[1227,1325,392,418]" pageId="2" pageNumber="327" type="holotype">holotype</typeStatus>
(1–dorsal habitus, 2–head and pronotum, 3–head, 4–lateral habitus), 508,
<taxonomicName id="4C534D271B03DE3A44775FB9D134FE54" ID-CoL="8K5HK" authority="Frieser, 1981" authorityName="Frieser" authorityYear="1981" box="[894,1319,428,454]" class="Insecta" family="Anthribidae" genus="Apatenia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="327" phylum="Arthropoda" rank="species" species="quadristigma">
<emphasis id="B927EAB61B03DE3A44775FB9D090FE54" box="[894,1155,428,454]" italics="true" pageId="2" pageNumber="327">Apatenia quadristigma</emphasis>
<bibRefCitation id="EFC24B551B03DE3A43875FB9D134FE54" author="Frieser, R." box="[1166,1319,428,454]" pageId="2" pageNumber="327" pagination="249 - 258" refId="ref10687" refString="Frieser, R. (1981) Beitrag zur Kenntnis der Anthribiden (Coleoptera, Anthribidae). Entomologische Arbeiten aus dem Museum G. Frey, 29, 249 - 258." type="journal article" year="1981">Frieser, 1981</bibRefCitation>
</taxonomicName>
, figs 5–6, female
<typeStatus id="54E888061B03DE3A47E35FC5D557FE78" box="[234,324,464,490]" pageId="2" pageNumber="327" type="allotype">allotype</typeStatus>
(5–dorsal habitus, 6–head), 509,
<taxonomicName id="4C534D271B03DE3A45B25FC5D77CFE78" ID-CoL="8K5HK" authorityName="Frieser" authorityYear="1981" box="[699,879,464,490]" class="Insecta" family="Anthribidae" genus="Apatenia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="327" phylum="Arthropoda" rank="species" species="quadristigma">
<emphasis id="B927EAB61B03DE3A45B25FC5D77CFE78" box="[699,879,464,490]" italics="true" pageId="2" pageNumber="327">A. quadristigma</emphasis>
</taxonomicName>
, figs 7, 9, 12 (7–head, 9–right antenna, 12–dorsal pronotal carina),
<taxonomicName id="4C534D271B03DE3A465E5FE1D5BBFD9C" ID-CoL="F8PV" authorityName="Tryzna &amp; Banar" authorityYear="2013" box="[343,424,500,526]" class="Insecta" family="Anthribidae" genus="Apatenia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="327" phylum="Arthropoda" rank="species" species="stysi">
<emphasis id="B927EAB61B03DE3A465E5FE1D5BBFD9C" box="[343,424,500,526]" italics="true" pageId="2" pageNumber="327">A. stysi</emphasis>
</taxonomicName>
, figs 8, 10–11, 13–14 (8– head, 10–right antenna, 11–abdomen, 13–dorsal pronotal carina, 14–pygidium), 510,
<taxonomicName id="4C534D271B03DE3A46735C0DD5D9FDA0" ID-CoL="F8PV" authorityName="Tryzna &amp; Banar" authorityYear="2013" box="[378,458,536,562]" class="Insecta" family="Anthribidae" genus="Apatenia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="327" phylum="Arthropoda" rank="species" species="stysi">
<emphasis id="B927EAB61B03DE3A46735C0DD5D9FDA0" box="[378,458,536,562]" italics="true" pageId="2" pageNumber="327">A. stysi</emphasis>
</taxonomicName>
, figs 15–18 (15–hemisternites, 16–segment VIII, 17–toothed plate of right hemisternite, 18–bursa copulatrix and spermatheca).
</paragraph>
</subSubSection>
<subSubSection id="C349652F1B03DE3A47CE5C75D50CFCC0" pageId="2" pageNumber="327" type="diagnosis">
<paragraph id="8BEC36A41B03DE3A47CE5C75D50CFCC0" blockId="2.[151,1437,284,1066]" pageId="2" pageNumber="327">
<emphasis id="B927EAB61B03DE3A47CE5C75D5C5FDE8" bold="true" box="[199,470,608,634]" pageId="2" pageNumber="327">
Diagnosis of
<taxonomicName id="4C534D271B03DE3A466E5C75D5DCFDE8" box="[359,463,608,634]" class="Insecta" family="Anthribidae" genus="Apatenia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="327" phylum="Arthropoda" rank="genus">
<emphasis id="B927EAB61B03DE3A466E5C75D5DCFDE8" bold="true" box="[359,463,608,634]" italics="true" pageId="2" pageNumber="327">Apatenia</emphasis>
</taxonomicName>
.
</emphasis>
Head long, narrow, rostrum lengthened, flat and broad, mandibles robust, strongly toothed. Eyes laterally situated.Antennae inserted at about midway between anterior margin of eyes and mandibles. Antennae longer than head, antennomeres I and II thick, III–VI more or less cylindrical, VII–VIII subtriangular, IX–XI forming loose club with antennomeres IX and X triangular, terminal antennomere rounded. Pronotum slightly transverse, narrow in front. Dorsal transverse carina of pronotum antebasal or sub-basal, its lateral margins directed anteriorly to about halfway along the side of prothorax. Elytra slightly depressed. Legs moderate, tarsi rather short.
</paragraph>
</subSubSection>
<subSubSection id="C349652F1B03DE3A47CE5D49D672FBB8" pageId="2" pageNumber="327" type="discussion">
<paragraph id="8BEC36A41B03DE3A47CE5D49D672FBB8" blockId="2.[151,1437,284,1066]" pageId="2" pageNumber="327">
<emphasis id="B927EAB61B03DE3A47CE5D49D519FCE4" bold="true" box="[199,266,860,886]" pageId="2" pageNumber="327">Notes</emphasis>
. The classification of the genera
<taxonomicName id="4C534D271B03DE3A45795D49D6C6FCE4" box="[624,725,860,886]" class="Insecta" family="Anthribidae" genus="Apatenia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="327" phylum="Arthropoda" rank="genus">
<emphasis id="B927EAB61B03DE3A45795D49D6C6FCE4" box="[624,725,860,886]" italics="true" pageId="2" pageNumber="327">Apatenia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C534D271B03DE3A440E5D49D76FFCE4" authorityName="Pascoe" authorityYear="1859" box="[775,892,860,886]" class="Insecta" family="Anthribidae" genus="Phaulimia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="327" phylum="Arthropoda" rank="genus">
<emphasis id="B927EAB61B03DE3A440E5D49D76FFCE4" box="[775,892,860,886]" italics="true" pageId="2" pageNumber="327">Phaulimia</emphasis>
</taxonomicName>
(both placed in the tribe
<taxonomicName id="4C534D271B03DE3A438E5D49D18BFCE4" authority="Bedel, 1882" authorityName="Bedel" authorityYear="1882" box="[1159,1432,860,886]" class="Insecta" family="Anthribidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="327" phylum="Arthropoda" rank="tribe" tribe="Platyrhinini">
Platyrhinini
<bibRefCitation id="EFC24B551B03DE3A42195D49D18BFCE4" author="Bedel, L. E. M." box="[1296,1432,860,886]" pageId="2" pageNumber="327" pagination="1 - 16" refId="ref10467" refString="Bedel, L. E. M. (1882) Faune des coleopteres du Bassin de la Seine. VI. Rhynchophora. Annales de la Societe Entomologique de France, Series 6, 2, 1 - 16. https: // doi. org / 10.5962 / bhl. title. 8757" type="journal article" year="1882">Bedel, 1882</bibRefCitation>
</taxonomicName>
, according
<bibRefCitation id="EFC24B551B03DE3A46045D95D63FFC08" author="Tryzna, M. &amp; Valentine, B. D." box="[269,556,896,922]" pageId="2" pageNumber="327" pagination="90 - 107" refId="ref12453" refString="Tryzna, M. &amp; Valentine, B. D. (2011) Anthribidae. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 7. Apollo Books, Stenstrup, pp. 64 - 66 + 90 - 107." type="book chapter" year="2011">Trýzna &amp; Valentine 2011</bibRefCitation>
,
<bibRefCitation id="EFC24B551B03DE3A453E5D95D6DBFC08" author="Tryzna, M. &amp; Alonso-Zarazaga, M. A. &amp; Barrios, H. &amp; Borovec, R. &amp; Bouchard, P. &amp; Caldara, R. &amp; Colonnelli, E. &amp; Gultekin, L. &amp; Hlavac, P. &amp; Korotyaev, B. &amp; Lyal, C. H. C. &amp; Machado, A. &amp; Meregalli, M. &amp; Pierotti, H. &amp; Ren, L. &amp; Sanchez-Ruiz, M. &amp; Sforzi, A. &amp; Silfverberg, H. &amp; Skuhrovec, J. &amp; Tryzna, M. &amp; Velazquez de Castro, A. J. &amp; Yunakov, N. N." box="[567,712,896,922]" pageId="2" pageNumber="327" pagination="18 - 33" refId="ref12016" refString="Tryzna, M. (2017) Anthribidae. In: Alonso-Zarazaga, M. A., Barrios, H., Borovec, R., Bouchard, P., Caldara, R., Colonnelli, E., Gultekin, L., Hlavac, P., Korotyaev, B., Lyal, C. H. C., Machado, A., Meregalli, M., Pierotti, H., Ren, L., Sanchez-Ruiz, M., Sforzi, A., Silfverberg, H., Skuhrovec, J., Tryzna, M., Velazquez de Castro, A. J. &amp; Yunakov, N. N. (2017) Cooperative Catalogue of Palaearctic Coleoptera. Curculionoidea. Monografias electronicas SEA 8. Sociedad Entomologica Aragonesa (S. E. A.), Zaragoza, pp. 18 - 33." type="journal article" year="2017">Trýzna 2017</bibRefCitation>
) need to be revised. Both genera contain numerous species, and occur in a large area (Oriental, Afrotropical and Australian distribution). Their generic descriptions were based upon the description of the first included species and have not been suitably adjusted to accommodate all subsequently described species. It is likely that the two genera will be maintained but perhaps with different species compositions. This will be the subject of future studies.
</paragraph>
</subSubSection>
<subSubSection id="C349652F1B0EDE37479E5E82D628FF23" box="[151,571,151,177]" pageId="15" pageNumber="340" type="nomenclature">
<paragraph id="8BEC36A41B0EDE37479E5E82D628FF23" blockId="15.[151,571,151,177]" box="[151,571,151,177]" pageId="15" pageNumber="340">
<heading id="D0A481C81B0EDE37479E5E82D628FF23" bold="true" box="[151,571,151,177]" fontSize="11" level="1" pageId="15" pageNumber="340" reason="1">
<emphasis id="B927EAB61B0EDE37479E5E82D628FF23" bold="true" box="[151,571,151,177]" pageId="15" pageNumber="340">
Key to Malagasy
<taxonomicName id="4C534D271B0EDE3746785E82D5CDFF23" ID-CoL="8HCZN" authority="Pascoe, 1859" authorityName="Pascoe" authorityYear="1859" box="[369,478,151,177]" class="Insecta" family="Anthribidae" genus="Apatenia" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="340" phylum="Arthropoda" rank="genus">
<emphasis id="B927EAB61B0EDE3746785E82D5CDFF23" bold="true" box="[369,478,151,177]" italics="true" pageId="15" pageNumber="340">Apatenia</emphasis>
</taxonomicName>
species
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C349652F1B0EDE37479E5EC9D18FFA15" pageId="15" pageNumber="340" type="key">
<keyStep id="30A72E011B0EDE37479E5EC9D18FFEBE" pageId="15" pageNumber="340">
<paragraph id="8BEC36A41B0EDE37479E5EC9D18FFE9D" blockId="15.[151,1437,220,1415]" pageId="15" pageNumber="340">
<keyLead id="30A295911B0EDE37479E5EC9D18FFE9D" pageId="15" pageNumber="340">
1 Dorsal transverse carina of pronotum distinctly antebasal (
<figureCitation id="13682A211B0EDE37443F5EC9D7BCFF60" box="[822,943,220,242]" captionStart="FIGURES 23–26" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[182,1408,185,1918]" captionTargetId="figure-18@10.[178,1408,181,1918]" captionTargetPageId="10" captionText="FIGURES 23–26. Malagasy Apatenia species, habitus in dorsal view. 23, A. quadristigma Frieser, 1981 (female); 24, A. oculifera Frieser 2000 (male); 25, A. stysi Trýzna &amp; Baňař, 2013 (paratype, female); 26, A. sulcicollis Frieser, 2000 (male)." figureDoi="http://doi.org/10.5281/zenodo.14239881" httpUri="https://zenodo.org/record/14239881/files/figure.png" pageId="15" pageNumber="340">Figs 24, 25</figureCitation>
), i.e. markedly distant from posterior margin of pronotum........................................................................................... 2
</keyLead>
</paragraph>
<paragraph id="8BEC36A41B0EDE37479E5F02D18FFEBE" blockId="15.[151,1437,220,1415]" box="[151,1436,279,301]" pageId="15" pageNumber="340">
<keyLead id="30A295911B0EDE37479E5F02D18FFEBE" box="[151,1436,279,301]" pageId="15" pageNumber="340">
- Dorsal transverse carina of pronotum sub-basal (
<figureCitation id="13682A211B0EDE3745BD5F02D730FEBF" box="[692,803,279,301]" captionStart="FIGURES 23–26" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[182,1408,185,1918]" captionTargetId="figure-18@10.[178,1408,181,1918]" captionTargetPageId="10" captionText="FIGURES 23–26. Malagasy Apatenia species, habitus in dorsal view. 23, A. quadristigma Frieser, 1981 (female); 24, A. oculifera Frieser 2000 (male); 25, A. stysi Trýzna &amp; Baňař, 2013 (paratype, female); 26, A. sulcicollis Frieser, 2000 (male)." figureDoi="http://doi.org/10.5281/zenodo.14239881" httpUri="https://zenodo.org/record/14239881/files/figure.png" pageId="15" pageNumber="340">Figs 23, 26</figureCitation>
,
<figureCitation id="13682A211B0EDE3744245F02D77FFEBE" box="[813,876,279,300]" captionStart="FIGURES 27–31" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[209,1378,184,1919]" captionTargetId="figure-17@11.[197,1390,181,1919]" captionTargetPageId="11" captionText="FIGURES 27–31. Malagasy Apatenia species, habitus in dorsal view. 27, A. longiclava Wolfrum, 1955 (male); 28, A. fallax Frieser, 2010 (female); 29–31, A. mesostigma Wolfrum, 1961 (female): 29, habitus in dorsal view; 30 and 31, colour of elytral spot in different viewing angles." figureDoi="http://doi.org/10.5281/zenodo.14239883" httpUri="https://zenodo.org/record/14239883/files/figure.png" pageId="15" pageNumber="340">27–29</figureCitation>
), i.e. approaching posterior margin of pronotum....... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A72E011B0EDE37479E5F21D18FFD85" pageId="15" pageNumber="340">
<paragraph id="8BEC36A41B0EDE37479E5F21D18FFE30" blockId="15.[151,1437,220,1415]" pageId="15" pageNumber="340">
<keyLead id="30A295911B0EDE37479E5F21D18FFE30" pageId="15" pageNumber="340">
2 Rostrum almost as long as wide. Eyes small, minimal distance between eyes about twice as wide as length of eye (only females known). Dorsal transverse carina of pronotum only weakly bisinuate, not convex in middle. Disc of pronotum moderately convex, without weak transverse depression. Elytra black, without basal elevation, with only two pairs of transverse brilliant white stripes in basal and sub-basal area.........................................
<taxonomicName id="4C534D271B0EDE3743535F99D151FE33" ID-CoL="F8PV" authority="Tryzna &amp; Banar" authorityName="Tryzna &amp; Banar" authorityYear="2013" box="[1114,1346,396,418]" class="Insecta" family="Anthribidae" genus="Apatenia" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="340" phylum="Arthropoda" rank="species" species="stysi">
<emphasis id="B927EAB61B0EDE3743535F99D0B2FE33" bold="true" box="[1114,1185,396,417]" italics="true" pageId="15" pageNumber="340">A. stysi</emphasis>
Trýzna &amp; Baňař
</taxonomicName>
(
<figureCitation id="13682A211B0EDE3742465F99D186FE30" box="[1359,1429,396,418]" captionStart="FIGURES 23–26" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[182,1408,185,1918]" captionTargetId="figure-18@10.[178,1408,181,1918]" captionTargetPageId="10" captionText="FIGURES 23–26. Malagasy Apatenia species, habitus in dorsal view. 23, A. quadristigma Frieser, 1981 (female); 24, A. oculifera Frieser 2000 (male); 25, A. stysi Trýzna &amp; Baňař, 2013 (paratype, female); 26, A. sulcicollis Frieser, 2000 (male)." figureDoi="http://doi.org/10.5281/zenodo.14239881" httpUri="https://zenodo.org/record/14239881/files/figure.png" pageId="15" pageNumber="340">Fig. 25</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="8BEC36A41B0EDE37479E5FBFD18FFD85" blockId="15.[151,1437,220,1415]" pageId="15" pageNumber="340">
<keyLead id="30A295911B0EDE37479E5FBFD18FFD85" pageId="15" pageNumber="340">
- Rostrum noticeably wider than longer. Eyes large, length of eye about twice longer than minimal distance between eyes (in both sexes). Dorsal transverse carina of pronotum bisinuate, convex in middle. Disc of pronotum with weak transverse depression. Elytra grey, with basal elevation from elytral interval I to III, with two distinct black spots on basal elevation surrounded by rusty or light brown margin, and transverse light brown stripe at apex.....................
<taxonomicName id="4C534D271B0EDE37438D5C14D150FD85" ID-CoL="67KHL" authority="Frieser" authorityName="Frieser" authorityYear="2000" box="[1156,1347,513,535]" class="Insecta" family="Anthribidae" genus="Apatenia" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="340" phylum="Arthropoda" rank="species" species="oculifera">
<emphasis id="B927EAB61B0EDE37438D5C14D0EAFD85" bold="true" box="[1156,1273,513,535]" italics="true" pageId="15" pageNumber="340">A. oculifera</emphasis>
Frieser
</taxonomicName>
(
<figureCitation id="13682A211B0EDE3742465C14D186FD85" box="[1359,1429,513,535]" captionStart="FIGURES 23–26" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[182,1408,185,1918]" captionTargetId="figure-18@10.[178,1408,181,1918]" captionTargetPageId="10" captionText="FIGURES 23–26. Malagasy Apatenia species, habitus in dorsal view. 23, A. quadristigma Frieser, 1981 (female); 24, A. oculifera Frieser 2000 (male); 25, A. stysi Trýzna &amp; Baňař, 2013 (paratype, female); 26, A. sulcicollis Frieser, 2000 (male)." figureDoi="http://doi.org/10.5281/zenodo.14239881" httpUri="https://zenodo.org/record/14239881/files/figure.png" pageId="15" pageNumber="340">Fig. 24</figureCitation>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A72E011B0EDE37479E5C0AD18FFD3B" pageId="15" pageNumber="340">
<paragraph id="8BEC36A41B0EDE37479E5C0AD18FFDFD" blockId="15.[151,1437,220,1415]" pageId="15" pageNumber="340">
<keyLead id="30A295911B0EDE37479E5C0AD18FFDFD" pageId="15" pageNumber="340">
3 Disc of pronotum with one pair of distinct tubercles, tops of tubercles widely rounded and blunt. Elytra with two pairs of distinct tubercles on elytral interval II in sub-basal and median area, tops of tubercles more or less pointed...........................................................................................
<taxonomicName id="4C534D271B0EDE3743725C4CD150FDFD" ID-CoL="F8PW" authority="Frieser" authorityName="Frieser" authorityYear="2000" box="[1147,1347,601,623]" class="Insecta" family="Anthribidae" genus="Apatenia" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="340" phylum="Arthropoda" rank="species" species="sulcicollis">
<emphasis id="B927EAB61B0EDE3743725C4CD0E9FDFD" bold="true" box="[1147,1274,601,623]" italics="true" pageId="15" pageNumber="340">A. sulcicollis</emphasis>
Frieser
</taxonomicName>
(
<figureCitation id="13682A211B0EDE3742465C4CD186FDFD" box="[1359,1429,601,623]" captionStart="FIGURES 23–26" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[182,1408,185,1918]" captionTargetId="figure-18@10.[178,1408,181,1918]" captionTargetPageId="10" captionText="FIGURES 23–26. Malagasy Apatenia species, habitus in dorsal view. 23, A. quadristigma Frieser, 1981 (female); 24, A. oculifera Frieser 2000 (male); 25, A. stysi Trýzna &amp; Baňař, 2013 (paratype, female); 26, A. sulcicollis Frieser, 2000 (male)." figureDoi="http://doi.org/10.5281/zenodo.14239881" httpUri="https://zenodo.org/record/14239881/files/figure.png" pageId="15" pageNumber="340">Fig. 26</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="8BEC36A41B0EDE37479E5C62D18FFD3B" blockId="15.[151,1437,220,1415]" pageId="15" pageNumber="340">
<keyLead id="30A295911B0EDE37479E5C62D18FFD3B" pageId="15" pageNumber="340">- Disc of pronotum convex, without any tubercles. Elytra at most with only indistinct small elevations on elytral interval II in sub-basal area....................................................................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A72E011B0EDE37479E5CA4D18FFC8D" pageId="15" pageNumber="340">
<paragraph id="8BEC36A41B0EDE37479E5CA4D18FFD77" blockId="15.[151,1437,220,1415]" pageId="15" pageNumber="340">
<keyLead id="30A295911B0EDE37479E5CA4D18FFD77" pageId="15" pageNumber="340">4 Elytra with one distinct black spot on disc, or with four distinct, sharply delimited transverese and oblique brilliant white stripes.............................................................................................. 5</keyLead>
</paragraph>
<paragraph id="8BEC36A41B0EDE37479E5CF9D18FFC8D" blockId="15.[151,1437,220,1415]" pageId="15" pageNumber="340">
<keyLead id="30A295911B0EDE37479E5CF9D18FFC8D" pageId="15" pageNumber="340">- Elytra without distinct spot or white transverse stripes, either dark brown, or yellowish to light brown with more or less alternating areas of light and dark colour on individual elytral intervals.......................................... 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A72E011B0EDE37479E5D32D18FFBB5" pageId="15" pageNumber="340">
<paragraph id="8BEC36A41B0EDE37479E5D32D18FFC06" blockId="15.[151,1437,220,1415]" pageId="15" pageNumber="340">
<keyLead id="30A295911B0EDE37479E5D32D18FFC06" pageId="15" pageNumber="340">
5 Elytra grey, disc of elytra with one oval black spot reaching laterally to elytral interval III, area of this spot smooth and without tubercles, remainder of elytra with distinct rows of coarse striae. Even-numbered interstriae II, IV, VI and VIII with number of small, blunt tubercles. Frons with coarse sculpture. Underside of body, femora and tibiae white speckled.........................................................................................
<taxonomicName id="4C534D271B0EDE37435F5D6AD151FC07" ID-CoL="5VKHL" authority="Wolfrum" authorityName="Wolfrum" authorityYear="1961" box="[1110,1346,895,917]" class="Insecta" family="Anthribidae" genus="Apatenia" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="340" phylum="Arthropoda" rank="species" species="mesostigma">
<emphasis id="B927EAB61B0EDE37435F5D6AD0F5FC06" bold="true" box="[1110,1254,895,916]" italics="true" pageId="15" pageNumber="340">A. mesostigma</emphasis>
Wolfrum
</taxonomicName>
(
<figureCitation id="13682A211B0EDE3742465D6AD186FC06" box="[1359,1429,895,917]" captionStart="FIGURES 27–31" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[209,1378,184,1919]" captionTargetId="figure-17@11.[197,1390,181,1919]" captionTargetPageId="11" captionText="FIGURES 27–31. Malagasy Apatenia species, habitus in dorsal view. 27, A. longiclava Wolfrum, 1955 (male); 28, A. fallax Frieser, 2010 (female); 29–31, A. mesostigma Wolfrum, 1961 (female): 29, habitus in dorsal view; 30 and 31, colour of elytral spot in different viewing angles." figureDoi="http://doi.org/10.5281/zenodo.14239883" httpUri="https://zenodo.org/record/14239883/files/figure.png" pageId="15" pageNumber="340">Fig. 29</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="8BEC36A41B0EDE37479E5D89D18FFBB5" blockId="15.[151,1437,220,1415]" pageId="15" pageNumber="340">
<keyLead id="30A295911B0EDE37479E5D89D18FFBB5" pageId="15" pageNumber="340">
- Elytra with rows of fine striae, without any tubercles, blue-grey, disc of elytra with horseshoe-shaped black stripe, open in basal part of elytra, stripe extending laterally on interstriae VII, with two pairs of brilliant white stripes situated inside, one transverse pair in basal area anteriorly of sub-basal tubercles, the second oblique pair posterior to tubercles. Frons with delicate sculpture. Underside of body covered with monochromatic grey pubescence, only posterior margin of metasternum and mesepimeron with white pubescence.......................................................
<taxonomicName id="4C534D271B0EDE3743525A04D150FBB5" ID-CoL="8K5HK" authority="Frieser" authorityName="Frieser" authorityYear="1981" box="[1115,1347,1041,1063]" class="Insecta" family="Anthribidae" genus="Apatenia" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="340" phylum="Arthropoda" rank="species" species="quadristigma">
<emphasis id="B927EAB61B0EDE3743525A04D0E9FBB5" bold="true" box="[1115,1274,1041,1063]" italics="true" pageId="15" pageNumber="340">A. quadristigma</emphasis>
Frieser
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(
<figureCitation id="13682A211B0EDE3742465A04D186FBB5" box="[1359,1429,1041,1063]" captionStart="FIGURES 23–26" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[182,1408,185,1918]" captionTargetId="figure-18@10.[178,1408,181,1918]" captionTargetPageId="10" captionText="FIGURES 23–26. Malagasy Apatenia species, habitus in dorsal view. 23, A. quadristigma Frieser, 1981 (female); 24, A. oculifera Frieser 2000 (male); 25, A. stysi Trýzna &amp; Baňař, 2013 (paratype, female); 26, A. sulcicollis Frieser, 2000 (male)." figureDoi="http://doi.org/10.5281/zenodo.14239881" httpUri="https://zenodo.org/record/14239881/files/figure.png" pageId="15" pageNumber="340">Fig. 23</figureCitation>
)
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<paragraph id="8BEC36A41B0EDE37479E5A3AD18FFBF0" blockId="15.[151,1437,220,1415]" pageId="15" pageNumber="340">
<keyLead id="30A295911B0EDE37479E5A3AD18FFBF0" pageId="15" pageNumber="340">
6 Elytra dark brown, odd- and even-numbered interstriae uniform in colour, with only one pair of very small light-coloured dots on interstria II in median area.....................................................
<emphasis id="B927EAB61B0EDE37438C5A59D120FBF3" bold="true" box="[1157,1331,1100,1122]" pageId="15" pageNumber="340">
<taxonomicName id="4C534D271B0EDE37438C5A59D0F3FBF0" authorityName="Trýzna &amp; Rakotonirina" authorityYear="2024" box="[1157,1248,1100,1122]" class="Insecta" family="Anthribidae" genus="Apatenia" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="340" phylum="Arthropoda" rank="species" species="kuntei" status="sp. nov.">
<emphasis id="B927EAB61B0EDE37438C5A59D0F3FBF0" bold="true" box="[1157,1248,1100,1122]" italics="true" pageId="15" pageNumber="340">A. kuntei</emphasis>
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<taxonomicNameLabel id="A21457CD1B0EDE3743EF5A59D120FBF3" box="[1254,1331,1100,1121]" pageId="15" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13682A211B0EDE3742495A59D16FFBF3" box="[1344,1404,1100,1122]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[179,1408,183,1916]" captionTargetId="figure-17@3.[178,1408,179,1916]" captionTargetPageId="3" captionText="FIGURES 1–5. Apatenia kuntei, male holotype, 1, habitus in dorsal view; 2, head; 3, pronotum; 4, right antenna; 5, habitus in lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.14239863" httpUri="https://zenodo.org/record/14239863/files/figure.png" pageId="15" pageNumber="340">Figs 1</figureCitation>
,
<figureCitation id="13682A211B0EDE3742815A59D186FBF0" box="[1416,1429,1100,1122]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[179,1401,182,1918]" captionTargetId="figure-18@4.[178,1408,181,1918]" captionTargetPageId="4" captionText="FIGURES 6–10. Apatenia kuntei, female allotype, 6, habitus in dorsal view; 7, head; 8, pronotum; 9, left antenna; 10, habitus in lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.14239867" httpUri="https://zenodo.org/record/14239867/files/figure.png" pageId="15" pageNumber="340">6</figureCitation>
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<paragraph id="8BEC36A41B0EDE37479E5A7FD18FFB0E" blockId="15.[151,1437,220,1415]" pageId="15" pageNumber="340">
<keyLead id="30A295911B0EDE37479E5A7FD18FFB0E" pageId="15" pageNumber="340">- Elytra yellowish to light brown, odd-numbered interstriae monochromatic yellowish to brownish, even-numbered interstriae with alternating areas of yellow, light brown and blackish colour............................................... 7</keyLead>
</paragraph>
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<keyStep id="30A72E011B0EDE37479E5AB1D18FFA15" pageId="15" pageNumber="340">
<paragraph id="8BEC36A41B0EDE37479E5AB1D18FFA83" blockId="15.[151,1437,220,1415]" pageId="15" pageNumber="340">
<keyLead id="30A295911B0EDE37479E5AB1D18FFA83" pageId="15" pageNumber="340">
7 Dorsal transverse carina of pronotum bisinuate, distinctly approached to posterior margin of pronotum in middle, in contact with lateral carina forms double obtuse angle, lateral carinae of pronotum reaching to midlength of pronotum. Eyes laterally distinctly protruding and regularly convex, distance between eyes larger, ocular index from 1.00 (male) to 1.70 (female). Elytral declivity with pair of small tubercles..................................................
<taxonomicName id="4C534D271B0EDE3743AD5AE9D150FA80" ID-CoL="8K5H9" authority="Frieser" authorityName="Frieser" authorityYear="2010" box="[1188,1347,1276,1298]" class="Insecta" family="Anthribidae" genus="Apatenia" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="340" phylum="Arthropoda" rank="species" species="fallax">
<emphasis id="B927EAB61B0EDE3743AD5AE9D0EAFA80" bold="true" box="[1188,1273,1276,1298]" italics="true" pageId="15" pageNumber="340">A. fallax</emphasis>
Frieser
</taxonomicName>
(
<figureCitation id="13682A211B0EDE3742465AE9D186FA83" box="[1359,1429,1276,1298]" captionStart="FIGURES 27–31" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[209,1378,184,1919]" captionTargetId="figure-17@11.[197,1390,181,1919]" captionTargetPageId="11" captionText="FIGURES 27–31. Malagasy Apatenia species, habitus in dorsal view. 27, A. longiclava Wolfrum, 1955 (male); 28, A. fallax Frieser, 2010 (female); 29–31, A. mesostigma Wolfrum, 1961 (female): 29, habitus in dorsal view; 30 and 31, colour of elytral spot in different viewing angles." figureDoi="http://doi.org/10.5281/zenodo.14239883" httpUri="https://zenodo.org/record/14239883/files/figure.png" pageId="15" pageNumber="340">Fig. 28</figureCitation>
)
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</paragraph>
<paragraph id="8BEC36A41B0EDE37479E5B0FD18FFA15" blockId="15.[151,1437,220,1415]" pageId="15" pageNumber="340">
<keyLead id="30A295911B0EDE37479E5B0FD18FFA15" pageId="15" pageNumber="340">
- Dorsal transverse carina of pronotum sinuate, equidistant from posterior margin of pronotum along its entire length, in middle very shortly interrupted, lateral carinae of pronotum very short, not reaching to midlength of pronotum. Eyes more flat, rather convex in posterior part, distance between eyes smaller, ocular index from 0.70–0.80 (male) to 0.90–1.20 (female). Elytral declivity without pair of small tubercles..........................................
<taxonomicName id="4C534D271B0EDE37436D5B64D151FA15" ID-CoL="F8P7" authority="Wolfrum" authorityName="Wolfrum" authorityYear="1955" box="[1124,1346,1393,1415]" class="Insecta" family="Anthribidae" genus="Apatenia" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="340" phylum="Arthropoda" rank="species" species="longiclava">
<emphasis id="B927EAB61B0EDE37436D5B64D0F4FA15" bold="true" box="[1124,1255,1393,1415]" italics="true" pageId="15" pageNumber="340">A. longiclava</emphasis>
Wolfrum
</taxonomicName>
(
<figureCitation id="13682A211B0EDE3742465B64D186FA15" box="[1359,1429,1393,1415]" captionStart="FIGURES 27–31" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[209,1378,184,1919]" captionTargetId="figure-17@11.[197,1390,181,1919]" captionTargetPageId="11" captionText="FIGURES 27–31. Malagasy Apatenia species, habitus in dorsal view. 27, A. longiclava Wolfrum, 1955 (male); 28, A. fallax Frieser, 2010 (female); 29–31, A. mesostigma Wolfrum, 1961 (female): 29, habitus in dorsal view; 30 and 31, colour of elytral spot in different viewing angles." figureDoi="http://doi.org/10.5281/zenodo.14239883" httpUri="https://zenodo.org/record/14239883/files/figure.png" pageId="15" pageNumber="340">Fig. 27</figureCitation>
)
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