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<document id="13128BE21FE9A3BB58DBACE109FAE0F7" ID-DOI="10.5852/cr-palevol2020v19a5" ID-ISSN="1777-571X" ID-ZooBank="urn:lsid:zoobank.org:pub:DB4BC4A0-E559-49AD-9452-55F1688D99DA" checkinTime="1732276064017" checkinUser="felipe" docAuthor="Collareta, Alberto, Lambert, Olivier, Palomino, Aldo Marcelo Benites, Urbina, Mario & Bianucci, Giovanni" docDate="2020" docId="390387B20B77FFA0C0FDFBA7FBC428AF" docLanguage="en" docName="CRPalevol.19.5.79-100.pdf" docOrigin="Comptes Rendus Palevol 19 (5)" docSource="http://dx.doi.org/10.5852/cr-palevol2020v19a5" docStyle="DocumentStyle:DACF86F01658A8850E13A63D98C287FD.1:CRPalevol.2020-.journal_article" docStyleId="DACF86F01658A8850E13A63D98C287FD" docStyleName="CRPalevol.2020-.journal_article" docStyleVersion="1" docTitle="Platyscaphokogia landinii Collareta & Lambert & Palomino & Urbina & Bianucci 2020, n. sp." docType="treatment" docUuid="FC3BE988-0F09-4262-911C-5B50070D1B97" docUuidSource="ZooBank" docVersion="1" lastPageNumber="84" masterDocId="C53AFFCA0B71FFA7C33CFE35FFC52B41" masterDocTitle="A new physeteroid from the late Miocene of Peru expands the diversity of extinct dwarf and pygmy sperm whales (Cetacea: Odontoceti: Kogiidae)" masterLastPageNumber="100" masterPageNumber="79" pageNumber="83" updateTime="1732276069354" updateUser="felipe" zenodo-license-document="CC-BY-4.0">
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<mods:title id="A537F0F57BE18B4FE27A289DD2960128">A new physeteroid from the late Miocene of Peru expands the diversity of extinct dwarf and pygmy sperm whales (Cetacea: Odontoceti: Kogiidae)</mods:title>
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<mods:namePart id="049172E957A9E5A664CC9B1D95CFB011">Collareta, Alberto</mods:namePart>
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<mods:affiliation id="B1206198DC3CE1F97FFA8CBE3B3F741D">Dipartimento di Scienze della Terra, Università di Pisa, via Santa Maria 53, 56126 Pisa (Italy) alberto. collareta @ unipi. it (corresponding author)</mods:affiliation>
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<mods:namePart id="75F8AD809B6D5AD5E2A7C827DE0FBC63">Lambert, Olivier</mods:namePart>
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<mods:affiliation id="72B9C0CF71C7431B64B6B9E0BBA3537A">Institut Royal des Sciences Naturelles de Belgique, D. O. Terre et Histoire de la Vie, rue Vautier 29, 1000 Brussels (Belgium) Centre de Recherches sur la Paléobiodiversité et les Paléoenvironnements, CR 2 P (CNRS, MNHN, UPMC, Sorbonne Université), Département Origines et Évolution, Muséum national d’Histoire naturelle, case postale 38, 57 rue Cuvier, F- 75231 Paris cedex 05 (France)</mods:affiliation>
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<mods:namePart id="26673ADD99EF3F6282477002EDA40024">Palomino, Aldo Marcelo Benites</mods:namePart>
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<mods:affiliation id="EE40D2D0A2E0AA5E3C7BB62166F19A4B">Departamento de Paleontologia de Vertebrados, Museo de Historia Natural de la Universidad Nacional Mayor de San Marcos, avenida Arenales 1256, Lima 14 (Peru) and Paläontologisches Institut und Museum, Karl-Schmid-Strasse 4, CH- 8006 Zürich (Switzerland) and Center for Tropical Paleoecology and Archeology, Smithsonian Tropical Research Institute, Balboa, Ancón, 0843 - 03092, Panama (Panama)</mods:affiliation>
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<mods:namePart id="835F01148D86B2DB93AF3215C03F7D12">Urbina, Mario</mods:namePart>
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<mods:affiliation id="E260C6EEDA331B8C0EC92128CDD92613">Departamento de Paleontologia de Vertebrados, Museo de Historia Natural de la Universidad Nacional Mayor de San Marcos, avenida Arenales 1256, Lima 14 (Peru)</mods:affiliation>
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<mods:namePart id="49BC03EA590EB4A15EE2BF6EDB9872D9">Bianucci, Giovanni</mods:namePart>
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<mods:affiliation id="745C125542D288268A21B71279020AC5">Dipartimento di Scienze della Terra, Università di Pisa, via Santa Maria 53, 56126 Pisa (Italy)</mods:affiliation>
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<subSubSection id="F9B0652F0B77FFA1C0FDFBA7FADE2EEF" box="[961,1307,1426,1454]" pageId="6" pageNumber="83" type="nomenclature">
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<paragraph id="B11536A40B77FFA1C0FDFBA7FADE2EEF" blockId="6.[961,1307,1426,1454]" box="[961,1307,1426,1454]" pageId="6" pageNumber="83">
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<heading id="EA5D81C80B77FFA1C0FDFBA7FADE2EEF" box="[961,1307,1426,1454]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="83" reason="2">
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<taxonomicName id="76AA4D270B77FFA1C0FDFBA7FB1C2EEC" authority="Collareta & Lambert & Palomino & Urbina & Bianucci, 2020" authorityName="Collareta & Lambert & Palomino & Urbina & Bianucci" authorityYear="2020" box="[961,1241,1426,1453]" class="Mammalia" family="Kogiidae" genus="Platyscaphokogia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="83" phylum="Chordata" rank="species" species="landinii" status="sp. nov.">
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<emphasis id="83DEEAB60B77FFA1C0FDFBA7FB1C2EEC" bold="true" box="[961,1241,1426,1453]" italics="true" pageId="6" pageNumber="83">Platyscaphokogia landinii</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="98ED57CD0B77FFA1C7DCFBA1FADE2EEF" box="[1248,1307,1428,1454]" pageId="6" pageNumber="83" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="B11536A40B77FFA1C71FFB86FB7C2E8C" blockId="6.[1059,1209,1459,1485]" box="[1059,1209,1459,1485]" pageId="6" pageNumber="83">
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(
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<figureCitation id="29912A210B77FFA1C710FB86FB462E8C" box="[1068,1155,1459,1485]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStart-3="FIG" captionStart-4="FIG" captionStartId-0="8.[132,143,1455,1472]" captionStartId-1="9.[134,145,1568,1585]" captionStartId-2="10.[134,145,992,1009]" captionStartId-3="11.[132,143,1544,1561]" captionStartId-4="12.[132,142,1856,1873]" captionTargetBox-0="[143,1444,215,1409]" captionTargetBox-1="[132,1455,214,1526]" captionTargetBox-2="[132,1455,215,953]" captionTargetBox-4="[132,1450,215,1816]" captionTargetId-0="figure-81@8.[170,1408,214,1413]" captionTargetId-1="figure-9@9.[132,1455,214,1512]" captionTargetId-2="figure-14@10.[132,1455,216,876]" captionTargetId-4="figure-88@12.[132,1455,215,1817]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-4="12" captionText-0="FIG. 2. — Cranium, in dorsal view, of the kogiid sperm whale Platyscaphokogia landinii n. gen., n. sp. (MUSM 3405, holotype), from the upper Miocene of the Pisco Formation (East Pisco Basin, southern Peru). A, photograph; B, corresponding explanatory line drawing. The grey-shaded areas correspond to hardened sediment. The dashed line approximates the position of the maxilla-frontal suture. Scale bar: 5 cm." captionText-1="FIG. 3. — Cranium, in ventral view, of the kogiid sperm whale Platyscaphokogia landinii n. gen., n. sp. (MUSM 3405, holotype), from the upper Miocene of the Pisco Formation (East Pisco Basin, southern Peru). A, photograph; B, corresponding explanatory line drawing. The grey-shaded areas correspond to hardened sediment. The dashed line approximates the position of several sutures and borders. Scale bar: 5 cm." captionText-2="FIG. 4. — Cranium, in right lateral view, of the kogiid sperm whale Platyscaphokogia landinii n. gen., n. sp. (MUSM 3405, holotype), from the upper Miocene of the Pisco Formation (East Pisco Basin, southern Peru). A, photograph; B, corresponding explanatory line drawing. Scale bar: 5 cm." captionText-3="FIG. 5. — Cranium, in anterodorsal view, of the kogiid sperm whale Platyscaphokogia landinii n. gen., n. sp. (MUSM 3405, holotype), from the upper Miocene of the Pisco Formation (East Pisco Basin, southern Peru); close-up of the dorsal surface of the neurocranium. Scale bar: 5 cm." captionText-4="FIG. 6. — Cranium of the kogiid sperm whalePlatyscaphokogialandinii n.gen.,n.sp.(MUSM 3291,paratype),from the upper Miocene of the Pisco Formation (East Pisco Basin, southern Peru):A, dorsal view;B, ventral view;C, right lateral view (close-up of the supraorbital region,with indication of bones and sutures). Scale bars:A,B,5 cm;C,2 cm." pageId="6" pageNumber="83">Figs 2-6</figureCitation>
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;
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<figureCitation id="29912A210B77FFA1C7ADFB86FB772E8C" box="[1169,1202,1459,1485]" captionStart="FIG" captionStartId="16.[132,143,1106,1123]" captionTargetBox="[132,1455,215,1063]" captionTargetId="figure-474@16.[132,1455,225,1064]" captionTargetPageId="16" captionText="FIG. 8. — Schematic comparison between the cranial morphologies of the two scaphokogiine genera known to date. A, Right lateral view (photograph and superimposed line drawing) of MUSM 3405, holotype cranium of Platyscaphokogia landinii n. gen., n. sp., from the upper Miocene of the Pisco Formation (East Pisco Basin, southern Peru). B, Right lateral view (photograph and superimposed line drawing) of MUSM 3244, unpublished cranium of Scaphokogia cochlearis, from the upper Miocene of the Pisco Formation (East Pisco Basin, southern Peru). Note the strong differences existing between the two taxa in terms of height and slope of the rostrum. Scale bar: 5 cm." pageId="6" pageNumber="83">8A</figureCitation>
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)
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</paragraph>
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. — The specific name honours Walter Landini, Italian vertebrate palaeontologist, for his substantial contribution to the knowledge and understanding of both the Miocene vertebrate assemblages of the East Pisco Basin (e.g.,
|
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<bibRefCitation id="D53B4B550B77FFA1C7F4F8B2FA4A2DDE" author="LANDINI W. & ALTAMIRANO-SIERRA A. & COLLARETA A. & DI CELMA C. & URBINA M. & BIANUCCI G." box="[1224,1423,1671,1695]" pageId="6" pageNumber="83" pagination="168 - 190" refId="ref19240" refString="LANDINI W., ALTAMIRANO-SIERRA A., COLLARETA A., DI CELMA C., URBINA M. & BIANUCCI G. 2017 a. - The late Miocene elasmobranch assemblage from Cerro Colorado (Pisco Formation, Peru). Journal of South American Earth Sciences 73: 168 - 190. https: // doi. org / 10.1016 / j. jsames. 2016.12.010" type="book chapter" year="2017">
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Landini
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<emphasis id="83DEEAB60B77FFA1C627F8BDFA8E2DDE" box="[1307,1355,1671,1695]" italics="true" pageId="6" pageNumber="83">et al.</emphasis>
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2017a
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</bibRefCitation>
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, b, 2019) and the Italian fossil record of physeteroids (e.g.,
|
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<bibRefCitation id="D53B4B550B77FFA1C603F897FC762D95" author="BIANUCCI G. & LANDINI W." pageId="6" pageNumber="83" pagination="445 - 453" refId="ref15170" refString="BIANUCCI G. & LANDINI W. 1999. - Kogia pusilla from the Middle Pliocene of Tuscany (Italy) and a phylogenetic analysis of the family Kogiidae (Odontoceti, Cetacea). Rivista Italiana di Paleontologia e Stratigrafia 105: 445 - 453. https: // doi. org / 10.13130 / 2039 - 4942 / 5385" type="journal article" year="1999">Bianucci & Landini 1999</bibRefCitation>
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,
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<bibRefCitation id="D53B4B550B77FFA1C083F889FC372D95" author="BIANUCCI G. & LANDINI W." box="[959,1010,1724,1748]" pageId="6" pageNumber="83" pagination="103 - 131" refId="ref15231" refString="BIANUCCI G. & LANDINI W. 2006. - Killer sperm whale: a new basal physeteroid (Mammalia, Cetacea) from the late Miocene of Italy. Zoological Journal of the Linnean Society 148: 103 - 131. https: // 10.1111 / j. 10963642.2006.00228. x" type="journal article" year="2006">2006</bibRefCitation>
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).
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</paragraph>
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<materialsCitation id="01C23CF90B77FFA1C011F8D8FB4A2C5E" collectionCode="MUSM" pageId="6" pageNumber="83" specimenCode="MUSM 3405" specimenCount="1" typeStatus="holotype">
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<typeStatus id="6E1188060B77FFA1C011F8D8FC522C42" box="[813,919,1773,1796]" pageId="6" pageNumber="83" type="holotype">HOLOTYPE</typeStatus>
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</smallCapsWord>
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. —
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<specimenCode id="E10C9EDF0B77FFA1C0F8F8D8FB8B2C44" box="[964,1102,1773,1797]" collectionCode="MUSM" pageId="6" pageNumber="83">MUSM 3405</specimenCode>
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, an incomplete skull lacking the tip of the rostrum and the basicranium.
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</paragraph>
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. —
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<collectionCode id="D7BBAE610B77FFA1C084F90DFBC02C11" box="[952,1029,1848,1872]" pageId="6" pageNumber="83">MUSM</collectionCode>
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3291, an incomplete skull, preserving only the rostrum and the right portion of the neurocranium.
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</paragraph>
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<smallCapsWord id="B7F3A0780B77FFA1C011F9B1FC982CD8" baselines="1941,1941" box="[813,861,1924,1947]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="6" pageNumber="83">
|
||||
<collectionCode id="D7BBAE610B77FFA1C011F9B1FC982CD8" box="[813,861,1924,1947]" pageId="6" pageNumber="83">TYPE</collectionCode>
|
||||
</smallCapsWord>
|
||||
<smallCapsWord id="B7F3A0780B77FFA1C05DF9BCFC7F2CDB" baselines="1942" box="[865,954,1929,1946]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="locality" pageId="6" pageNumber="83">
|
||||
<collectionCode id="D7BBAE610B77FFA1C05DF9BCFC7F2CDB" box="[865,954,1929,1946]" pageId="6" pageNumber="83">LOCALITY</collectionCode>
|
||||
</smallCapsWord>
|
||||
. — The
|
||||
<typeStatus id="6E1188060B77FFA1C737F9B6FB9A2CDA" box="[1035,1119,1923,1947]" pageId="6" pageNumber="83" type="holotype">holotype</typeStatus>
|
||||
was found at Cerro Hueco la Zorra,
|
||||
<collectingRegion id="736EF8460B77FFA1C017F9ABFC822CF4" box="[811,839,1950,1973]" country="Peru" name="Ica" pageId="6" pageNumber="83">Ica</collectingRegion>
|
||||
desert,
|
||||
<emphasis id="83DEEAB60B77FFA1C0AEF9AAFC642CF7" box="[914,929,1951,1974]" italics="true" pageId="6" pageNumber="83">c.</emphasis>
|
||||
<quantity id="76529B410B77FFA1C09BF9ABFC202CF7" box="[935,997,1950,1974]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="83" unit="km" value="30.0">30 km</quantity>
|
||||
south of the town of
|
||||
<collectingRegion id="736EF8460B77FFA1C78EF9ABFB142CF4" box="[1202,1233,1950,1973]" country="Peru" name="Ica" pageId="6" pageNumber="83">Ica</collectingRegion>
|
||||
, southern coast of
|
||||
<collectingCountry id="C9BD76340B77FFA1C6BFF9ABFA6A2CF4" box="[1411,1455,1950,1973]" name="Peru" pageId="6" pageNumber="83">Peru</collectingCountry>
|
||||
(
|
||||
<figureCitation id="29912A210B77FFA1C009F98DFCBE2C91" box="[821,891,1976,2000]" captionStart="FIG" captionStartId="5.[132,143,1804,1821]" captionTargetBox="[137,1451,219,1742]" captionTargetId="figure-17@5.[698,1267,784,1355]" captionTargetPageId="5" captionText="FIG. 1. — A, Map of the major structural trends and basins of coastal Peru. The red dashed rectangle outlines the location of the area shown in detail in panel B. Redrawn and modified from Travis et al. (1976) and Thornburg & Kulm (1981); B, Schematic geological map of the East Pisco Basin, showing the areas of Cenozoic outcrop and the location of the sites where the holotype and paratype of the new physeteroid taxon described herein were found (Cerro Hueco la Zorra and Cerro los Quesos; black stars). Redrawn and modified from DeVries & Schrader (1997); C, Schematic, dip-oriented allostratigraphic diagram for the Pisco Formation and the lower Chilcatay and Otuma formations, showing the stratigraphic position of the two kogiid fossils (MUSM 3405 and MUSM 3291) studied in the present work along with 40Ar/39Ar-dated volcanic ash layers (labelled with their field names on the left side of the diagram) and measured stratigraphic sections published in previous works (Bosio et al. 2020b, and references therein). The Pisco Formation exhibits pronounced thinning to the northeast, with the allomembers onlapping onto the basal composite surface (PE0.0) and arranged in a retrogradational pattern, progressively offset to northeast. Modified from Bosio et al. (2020b).Abbreviations: RN, Roca Negra; CTP, Cerro las Tres Piramides; ULL, Ullujaya; CMH, Cerros la Mama y la Hija; CLZ, Cerros Cadena de los Zanjones; ANF, Anfiteatro; CLQ, Cerro los Quesos; CLQw, Cerro los Quesos west; CLB, Cerro la Bruja; HLZ, Cerro Hueco la Zorra; CBC, Cerro Blanco." pageId="6" pageNumber="83">Fig. 1B</figureCitation>
|
||||
); geographic coordinates of the finding site:
|
||||
<geoCoordinate id="D49E50630B77FFA1C625F98DFA6F2C91" box="[1305,1450,1976,2000]" degrees="14" direction="south" minutes="26" orientation="latitude" pageId="6" pageNumber="83" precision="1" seconds="44.35" value="-14.445653">14°26’44.35”S</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="D49E50630B77FFA1C011F9E6FC032CAA" box="[813,966,2003,2027]" degrees="75" direction="west" minutes="42" orientation="longitude" pageId="6" pageNumber="83" precision="1" seconds="24.05" value="-75.70668">75°42’24.05”W</geoCoordinate>
|
||||
.
|
||||
</paragraph>
|
||||
<caption id="E5D5662C0B76FFA0C3BAFEEEFD632A5B" pageId="7" pageNumber="84" startId="7.[134,143,219,236]" targetType="table">
|
||||
<paragraph id="B11536A40B76FFA0C3BAFEEEFD632A5B" blockId="7.[132,775,219,282]" pageId="7" pageNumber="84">
|
||||
<smallCapsWord id="B7F3A0780B76FFA0C3BAFEEEFF772BAD" baselines="232,233" box="[134,178,219,236]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Table" pageId="7" pageNumber="84">TABLE</smallCapsWord>
|
||||
1. — Measurements (in mm) of the skulls of
|
||||
<taxonomicName id="76AA4D270B76FFA0C10FFEEEFCC22BAD" authority="Collareta & Lambert & Palomino & Urbina & Bianucci, 2020" authorityName="Collareta & Lambert & Palomino & Urbina & Bianucci" authorityYear="2020" box="[563,775,219,236]" class="Mammalia" family="Kogiidae" genus="Platyscaphokogia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="84" phylum="Chordata" rank="species" species="landinii" status="n. gen., n. sp.">
|
||||
<emphasis id="83DEEAB60B76FFA0C10FFEEEFCC22BAD" box="[563,775,219,236]" italics="true" pageId="7" pageNumber="84">Platyscaphokogia landinii</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="98ED57CD0B76FFA0C3BAFEC7FF3A2A42" box="[134,255,242,259]" pageId="7" pageNumber="84" rank="species">n. gen., n. sp.</taxonomicNameLabel>
|
||||
, from the upper Miocene of the Pisco Formation (East Pisco Basin, southern Peru). Estimated dimensions are preceded by ‘
|
||||
<emphasis id="83DEEAB60B76FFA0C1AEFF3CFD592A5B" box="[658,668,265,282]" italics="true" pageId="7" pageNumber="84">e</emphasis>
|
||||
’.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="B11536A40B76FFA0C2C0FF61FE852F7C" pageId="7" pageNumber="84">
|
||||
<table id="C3AAC4040B760058C3BAFF61FD3A2F7C" box="[134,767,340,1085]" gridcols="3" gridrows="30" pageId="7" pageNumber="84">
|
||||
<tr id="0F9A34E60B760058C3BAFF61FD3A2A29" box="[134,767,340,360]" gridrow="0" pageId="7" pageNumber="84" rowspan-0="1">
|
||||
<th id="4C4B5D9A0B760058C2C0FF61FDB02A29" box="[508,629,340,360]" gridcol="1" gridrow="0" pageId="7" pageNumber="84">
|
||||
<emphasis id="83DEEAB60B76FFA0C2C0FF61FDB02A29" bold="true" box="[508,629,340,360]" pageId="7" pageNumber="84">MUSM 3405</emphasis>
|
||||
</th>
|
||||
<th id="4C4B5D9A0B760058C1BAFF61FD3A2A29" box="[646,767,340,360]" gridcol="2" gridrow="0" pageId="7" pageNumber="84">
|
||||
<emphasis id="83DEEAB60B76FFA0C1BAFF61FD3A2A29" bold="true" box="[646,767,340,360]" pageId="7" pageNumber="84">MUSM 3291</emphasis>
|
||||
</th>
|
||||
</tr>
|
||||
<tr id="0F9A34E60B760058C3BAFF5BFD3A2AC3" box="[134,767,366,386]" gridrow="1" pageId="7" pageNumber="84" rowspan-0="1">
|
||||
<td id="4C4B5D9A0B760058C2C0FF5BFDB02AC3" box="[508,629,366,386]" gridcol="1" gridrow="1" pageId="7" pageNumber="84">
|
||||
<emphasis id="83DEEAB60B76FFA0C13AFF5BFDAE2AC3" bold="true" box="[518,619,366,386]" pageId="7" pageNumber="84">(holotype)</emphasis>
|
||||
</td>
|
||||
<td id="4C4B5D9A0B760058C1BAFF5BFD3A2AC3" box="[646,767,366,386]" gridcol="2" gridrow="1" pageId="7" pageNumber="84">
|
||||
<emphasis id="83DEEAB60B76FFA0C1B3FF5BFD332AC3" bold="true" box="[655,758,366,386]" pageId="7" pageNumber="84">(paratype)</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="0F9A34E60B760058C3BAFFA4FD3A2AE4" box="[134,767,401,421]" gridrow="2" pageId="7" pageNumber="84">
|
||||
<th id="4C4B5D9A0B760058C3BAFFA4FE242AE4" box="[134,481,401,421]" gridcol="0" gridrow="2" pageId="7" pageNumber="84">Maximum preserved length of skull</th>
|
||||
<td id="4C4B5D9A0B760058C2C0FFA4FDB02AE4" box="[508,629,401,421]" gridcol="1" gridrow="2" pageId="7" pageNumber="84">550</td>
|
||||
<td id="4C4B5D9A0B760058C1BAFFA4FD3A2AE4" box="[646,767,401,421]" gridcol="2" gridrow="2" pageId="7" pageNumber="84">395</td>
|
||||
</tr>
|
||||
<tr id="0F9A34E60B760058C3BAFF9CFD3A2AFC" box="[134,767,425,445]" gridrow="3" pageId="7" pageNumber="84">
|
||||
<th id="4C4B5D9A0B760058C3BAFF9CFE242AFC" box="[134,481,425,445]" gridcol="0" gridrow="3" pageId="7" pageNumber="84">Length of rostrum</th>
|
||||
<td id="4C4B5D9A0B760058C2C0FF9CFDB02AFC" box="[508,629,425,445]" gridcol="1" gridrow="3" pageId="7" pageNumber="84">300</td>
|
||||
<td id="4C4B5D9A0B760058C1BAFF9CFD3A2AFC" box="[646,767,425,445]" gridcol="2" gridrow="3" pageId="7" pageNumber="84">270</td>
|
||||
</tr>
|
||||
<tr id="0F9A34E60B760058C3BAFFF4FD3A2A94" box="[134,767,449,469]" gridrow="4" pageId="7" pageNumber="84">
|
||||
<th id="4C4B5D9A0B760058C3BAFFF4FE242A94" box="[134,481,449,469]" gridcol="0" gridrow="4" pageId="7" pageNumber="84">Width of rostrum at base</th>
|
||||
<td id="4C4B5D9A0B760058C2C0FFF4FDB02A94" box="[508,629,449,469]" gridcol="1" gridrow="4" pageId="7" pageNumber="84">158</td>
|
||||
<td id="4C4B5D9A0B760058C1BAFFF4FD3A2A94" box="[646,767,449,469]" gridcol="2" gridrow="4" pageId="7" pageNumber="84">167</td>
|
||||
</tr>
|
||||
<tr id="0F9A34E60B760058C3BAFFECFD3A2AAC" box="[134,767,473,493]" gridrow="5" pageId="7" pageNumber="84">
|
||||
<th id="4C4B5D9A0B760058C3BAFFECFE242AAC" box="[134,481,473,493]" gridcol="0" gridrow="5" pageId="7" pageNumber="84">Height of rostrum at base</th>
|
||||
<td id="4C4B5D9A0B760058C2C0FFECFDB02AAC" box="[508,629,473,493]" gridcol="1" gridrow="5" pageId="7" pageNumber="84">70</td>
|
||||
<td id="4C4B5D9A0B760058C1BAFFECFD3A2AAC" box="[646,767,473,493]" gridcol="2" gridrow="5" pageId="7" pageNumber="84">–</td>
|
||||
</tr>
|
||||
<tr id="0F9A34E60B760058C3BAFFC7FD3A2947" box="[134,767,498,518]" gridrow="6" pageId="7" pageNumber="84">
|
||||
<th id="4C4B5D9A0B760058C3BAFFC7FE242947" box="[134,481,498,518]" gridcol="0" gridrow="6" pageId="7" pageNumber="84">Width of rostrum at mid length</th>
|
||||
<td id="4C4B5D9A0B760058C2C0FFC7FDB02947" box="[508,629,498,518]" gridcol="1" gridrow="6" pageId="7" pageNumber="84">106</td>
|
||||
<td id="4C4B5D9A0B760058C1BAFFC7FD3A2947" box="[646,767,498,518]" gridcol="2" gridrow="6" pageId="7" pageNumber="84">120</td>
|
||||
</tr>
|
||||
<tr id="0F9A34E60B760058C3BAFC3FFD3A295F" box="[134,767,522,542]" gridrow="7" pageId="7" pageNumber="84">
|
||||
<th id="4C4B5D9A0B760058C3BAFC3FFE24295F" box="[134,481,522,542]" gridcol="0" gridrow="7" pageId="7" pageNumber="84">Height of rostrum at mid length</th>
|
||||
<td id="4C4B5D9A0B760058C2C0FC3FFDB0295F" box="[508,629,522,542]" gridcol="1" gridrow="7" pageId="7" pageNumber="84">45</td>
|
||||
<td id="4C4B5D9A0B760058C1BAFC3FFD3A295F" box="[646,767,522,542]" gridcol="2" gridrow="7" pageId="7" pageNumber="84">53</td>
|
||||
</tr>
|
||||
<tr id="0F9A34E60B760058C3BAFC14FD3A2977" box="[134,767,545,566]" gridrow="8" pageId="7" pageNumber="84">
|
||||
<th id="4C4B5D9A0B760058C3BAFC14FE242977" box="[134,481,545,566]" gridcol="0" gridrow="8" pageId="7" pageNumber="84">Width of rostrum at ¾ length,</th>
|
||||
<td id="4C4B5D9A0B760058C2C0FC14FDB02977" box="[508,629,545,566]" gridcol="1" gridrow="8" pageId="7" pageNumber="84">70</td>
|
||||
<td id="4C4B5D9A0B760058C1BAFC14FD3A2977" box="[646,767,545,566]" gridcol="2" gridrow="8" pageId="7" pageNumber="84">79</td>
|
||||
</tr>
|
||||
<tr id="0F9A34E60B760058C3BAFC0EFD3A290E" box="[134,767,571,591]" gridrow="9" pageId="7" pageNumber="84" rowspan-1="1" rowspan-2="1">
|
||||
<th id="4C4B5D9A0B760058C3BAFC0EFE24290E" box="[134,481,571,591]" gridcol="0" gridrow="9" pageId="7" pageNumber="84">measured from posterior end</th>
|
||||
</tr>
|
||||
<tr id="0F9A34E60B760058C3BAFC66FD3A2926" box="[134,767,595,615]" gridrow="10" pageId="7" pageNumber="84">
|
||||
<th id="4C4B5D9A0B760058C3BAFC66FE242926" box="[134,481,595,615]" gridcol="0" gridrow="10" pageId="7" pageNumber="84">Height of rostrum at ¾ length,</th>
|
||||
<td id="4C4B5D9A0B760058C2C0FC66FDB02926" box="[508,629,595,615]" gridcol="1" gridrow="10" pageId="7" pageNumber="84">39</td>
|
||||
<td id="4C4B5D9A0B760058C1BAFC66FD3A2926" box="[646,767,595,615]" gridcol="2" gridrow="10" pageId="7" pageNumber="84">42</td>
|
||||
</tr>
|
||||
<tr id="0F9A34E60B760058C3BAFC58FD3A29C0" box="[134,767,621,641]" gridrow="11" pageId="7" pageNumber="84" rowspan-1="1" rowspan-2="1">
|
||||
<th id="4C4B5D9A0B760058C3BAFC58FE2429C0" box="[134,481,621,641]" gridcol="0" gridrow="11" pageId="7" pageNumber="84">measured from posterior end</th>
|
||||
</tr>
|
||||
<tr id="0F9A34E60B760058C3BAFCB0FD3A29D8" box="[134,767,645,665]" gridrow="12" pageId="7" pageNumber="84">
|
||||
<th id="4C4B5D9A0B760058C3BAFCB0FE2429D8" box="[134,481,645,665]" gridcol="0" gridrow="12" pageId="7" pageNumber="84">Width of mesorostral groove at apex</th>
|
||||
<td id="4C4B5D9A0B760058C2C0FCB0FDB029D8" box="[508,629,645,665]" gridcol="1" gridrow="12" pageId="7" pageNumber="84">38</td>
|
||||
<td id="4C4B5D9A0B760058C1BAFCB0FD3A29D8" box="[646,767,645,665]" gridcol="2" gridrow="12" pageId="7" pageNumber="84">37</td>
|
||||
</tr>
|
||||
<tr id="0F9A34E60B760058C3BAFCAAFD3A29F2" box="[134,767,671,691]" gridrow="13" pageId="7" pageNumber="84" rowspan-1="1" rowspan-2="1">
|
||||
<th id="4C4B5D9A0B760058C3BAFCAAFE2429F2" box="[134,481,671,691]" gridcol="0" gridrow="13" pageId="7" pageNumber="84">of rostrum</th>
|
||||
</tr>
|
||||
<tr id="0F9A34E60B760058C3BAFC82FD3A298A" box="[134,767,695,715]" gridrow="14" pageId="7" pageNumber="84">
|
||||
<th id="4C4B5D9A0B760058C3BAFC82FE24298A" box="[134,481,695,715]" gridcol="0" gridrow="14" pageId="7" pageNumber="84">Maximum width of dorsal opening</th>
|
||||
<td id="4C4B5D9A0B760058C2C0FC82FDB0298A" box="[508,629,695,715]" gridcol="1" gridrow="14" pageId="7" pageNumber="84">23</td>
|
||||
<td id="4C4B5D9A0B760058C1BAFC82FD3A298A" box="[646,767,695,715]" gridcol="2" gridrow="14" pageId="7" pageNumber="84">28</td>
|
||||
</tr>
|
||||
<tr id="0F9A34E60B760058C3BAFCE5FD3A29A5" box="[134,767,720,740]" gridrow="15" pageId="7" pageNumber="84" rowspan-1="1" rowspan-2="1">
|
||||
<th id="4C4B5D9A0B760058C3BAFCE5FE2429A5" box="[134,481,720,740]" gridcol="0" gridrow="15" pageId="7" pageNumber="84">of mesorostral groove posterior</th>
|
||||
</tr>
|
||||
<tr id="0F9A34E60B760058C3BAFCDFFD3A29BF" box="[134,767,746,766]" gridrow="16" pageId="7" pageNumber="84" rowspan-1="1" rowspan-2="1">
|
||||
<th id="4C4B5D9A0B760058C3BAFCDFFE2429BF" box="[134,481,746,766]" gridcol="0" gridrow="16" pageId="7" pageNumber="84">to dorsomedial contact between</th>
|
||||
</tr>
|
||||
<tr id="0F9A34E60B760058C3BAFD31FD3A2859" box="[134,767,772,792]" gridrow="17" pageId="7" pageNumber="84" rowspan-1="1" rowspan-2="1">
|
||||
<th id="4C4B5D9A0B760058C3BAFD31FE242859" box="[134,481,772,792]" gridcol="0" gridrow="17" pageId="7" pageNumber="84">premaxillae along rostrum</th>
|
||||
</tr>
|
||||
<tr id="0F9A34E60B760058C3BAFD29FD3A2871" box="[134,767,796,816]" gridrow="18" pageId="7" pageNumber="84">
|
||||
<th id="4C4B5D9A0B760058C3BAFD29FE242871" box="[134,481,796,816]" gridcol="0" gridrow="18" pageId="7" pageNumber="84">Distance from tip of rostrum to right</th>
|
||||
<td id="4C4B5D9A0B760058C2C0FD29FDB02871" box="[508,629,796,816]" gridcol="1" gridrow="18" pageId="7" pageNumber="84">320</td>
|
||||
<td id="4C4B5D9A0B760058C1BAFD29FD3A2871" box="[646,767,796,816]" gridcol="2" gridrow="18" pageId="7" pageNumber="84">
|
||||
<emphasis id="83DEEAB60B76FFA0C199FD29FD752871" box="[677,688,796,816]" italics="true" pageId="7" pageNumber="84">e</emphasis>
|
||||
290
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="0F9A34E60B760058C3BAFD00FD3A2808" box="[134,767,821,841]" gridrow="19" pageId="7" pageNumber="84" rowspan-1="1" rowspan-2="1">
|
||||
<th id="4C4B5D9A0B760058C3BAFD00FE242808" box="[134,481,821,841]" gridcol="0" gridrow="19" pageId="7" pageNumber="84">external naris</th>
|
||||
</tr>
|
||||
<tr id="0F9A34E60B760058C3BAFD78FD3A2820" box="[134,767,845,865]" gridrow="20" pageId="7" pageNumber="84">
|
||||
<th id="4C4B5D9A0B760058C3BAFD78FE242820" box="[134,481,845,865]" gridcol="0" gridrow="20" pageId="7" pageNumber="84">Greatest preorbital width</th>
|
||||
<td id="4C4B5D9A0B760058C2C0FD78FDB02820" box="[508,629,845,865]" gridcol="1" gridrow="20" pageId="7" pageNumber="84">
|
||||
<emphasis id="83DEEAB60B76FFA0C119FD78FDF52820" box="[549,560,845,865]" italics="true" pageId="7" pageNumber="84">e</emphasis>
|
||||
250
|
||||
</td>
|
||||
<td id="4C4B5D9A0B760058C1BAFD78FD3A2820" box="[646,767,845,865]" gridcol="2" gridrow="20" pageId="7" pageNumber="84">
|
||||
<emphasis id="83DEEAB60B76FFA0C199FD78FD752820" box="[677,688,845,865]" italics="true" pageId="7" pageNumber="84">e</emphasis>
|
||||
245
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="0F9A34E60B760058C3BAFD53FD3A283B" box="[134,767,870,890]" gridrow="21" pageId="7" pageNumber="84">
|
||||
<th id="4C4B5D9A0B760058C3BAFD53FE24283B" box="[134,481,870,890]" gridcol="0" gridrow="21" pageId="7" pageNumber="84">Least supraorbital width</th>
|
||||
<td id="4C4B5D9A0B760058C2C0FD53FDB0283B" box="[508,629,870,890]" gridcol="1" gridrow="21" pageId="7" pageNumber="84">235</td>
|
||||
<td id="4C4B5D9A0B760058C1BAFD53FD3A283B" box="[646,767,870,890]" gridcol="2" gridrow="21" pageId="7" pageNumber="84">e235</td>
|
||||
</tr>
|
||||
<tr id="0F9A34E60B760058C3BAFD4BFD3A28D3" box="[134,767,894,914]" gridrow="22" pageId="7" pageNumber="84">
|
||||
<th id="4C4B5D9A0B760058C3BAFD4BFE2428D3" box="[134,481,894,914]" gridcol="0" gridrow="22" pageId="7" pageNumber="84">Greatest postorbital width</th>
|
||||
<td id="4C4B5D9A0B760058C2C0FD4BFDB028D3" box="[508,629,894,914]" gridcol="1" gridrow="22" pageId="7" pageNumber="84">
|
||||
<emphasis id="83DEEAB60B76FFA0C119FD4BFDF528D3" box="[549,560,894,914]" italics="true" pageId="7" pageNumber="84">e</emphasis>
|
||||
255
|
||||
</td>
|
||||
<td id="4C4B5D9A0B760058C1BAFD4BFD3A28D3" box="[646,767,894,914]" gridcol="2" gridrow="22" pageId="7" pageNumber="84">
|
||||
<emphasis id="83DEEAB60B76FFA0C199FD4BFD7528D3" box="[677,688,894,914]" italics="true" pageId="7" pageNumber="84">e</emphasis>
|
||||
250
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="0F9A34E60B760058C3BAFDA3FD3A28EB" box="[134,767,918,938]" gridrow="23" pageId="7" pageNumber="84">
|
||||
<th id="4C4B5D9A0B760058C3BAFDA3FE2428EB" box="[134,481,918,938]" gridcol="0" gridrow="23" pageId="7" pageNumber="84">Width of left external bony naris</th>
|
||||
<td id="4C4B5D9A0B760058C2C0FDA3FDB028EB" box="[508,629,918,938]" gridcol="1" gridrow="23" pageId="7" pageNumber="84">16</td>
|
||||
<td id="4C4B5D9A0B760058C1BAFDA3FD3A28EB" box="[646,767,918,938]" gridcol="2" gridrow="23" pageId="7" pageNumber="84">–</td>
|
||||
</tr>
|
||||
<tr id="0F9A34E60B760058C3BAFD9BFD3A2883" box="[134,767,942,962]" gridrow="24" pageId="7" pageNumber="84">
|
||||
<th id="4C4B5D9A0B760058C3BAFD9BFE242883" box="[134,481,942,962]" gridcol="0" gridrow="24" pageId="7" pageNumber="84">Width of right external bony naris</th>
|
||||
<td id="4C4B5D9A0B760058C2C0FD9BFDB02883" box="[508,629,942,962]" gridcol="1" gridrow="24" pageId="7" pageNumber="84">11</td>
|
||||
<td id="4C4B5D9A0B760058C1BAFD9BFD3A2883" box="[646,767,942,962]" gridcol="2" gridrow="24" pageId="7" pageNumber="84">–</td>
|
||||
</tr>
|
||||
<tr id="0F9A34E60B760058C3BAFDF3FD3A289B" box="[134,767,966,986]" gridrow="25" pageId="7" pageNumber="84">
|
||||
<th id="4C4B5D9A0B760058C3BAFDF3FE24289B" box="[134,481,966,986]" gridcol="0" gridrow="25" pageId="7" pageNumber="84">Greatest width across external bony</th>
|
||||
<td id="4C4B5D9A0B760058C2C0FDF3FDB0289B" box="[508,629,966,986]" gridcol="1" gridrow="25" pageId="7" pageNumber="84">38</td>
|
||||
<td id="4C4B5D9A0B760058C1BAFDF3FD3A289B" box="[646,767,966,986]" gridcol="2" gridrow="25" pageId="7" pageNumber="84">–</td>
|
||||
</tr>
|
||||
<tr id="0F9A34E60B760058C3BAFDD5FD3A28B5" box="[134,767,992,1012]" gridrow="26" pageId="7" pageNumber="84" rowspan-1="1" rowspan-2="1">
|
||||
<th id="4C4B5D9A0B760058C3BAFDD5FE2428B5" box="[134,481,992,1012]" gridcol="0" gridrow="26" pageId="7" pageNumber="84">nares</th>
|
||||
</tr>
|
||||
<tr id="0F9A34E60B760058C3BAFDCDFD3A2F4D" box="[134,767,1016,1036]" gridrow="27" pageId="7" pageNumber="84">
|
||||
<th id="4C4B5D9A0B760058C3BAFDCDFE242F4D" box="[134,481,1016,1036]" gridcol="0" gridrow="27" pageId="7" pageNumber="84">Greatest width across premaxillae</th>
|
||||
<td id="4C4B5D9A0B760058C2C0FDCDFDB02F4D" box="[508,629,1016,1036]" gridcol="1" gridrow="27" pageId="7" pageNumber="84">89</td>
|
||||
<td id="4C4B5D9A0B760058C1BAFDCDFD3A2F4D" box="[646,767,1016,1036]" gridcol="2" gridrow="27" pageId="7" pageNumber="84">–</td>
|
||||
</tr>
|
||||
<tr id="0F9A34E60B760058C3BAFA25FD3A2F65" box="[134,767,1040,1060]" gridrow="28" pageId="7" pageNumber="84">
|
||||
<th id="4C4B5D9A0B760058C3BAFA25FE242F65" box="[134,481,1040,1060]" gridcol="0" gridrow="28" pageId="7" pageNumber="84">Width across premaxillae at mid</th>
|
||||
<td id="4C4B5D9A0B760058C2C0FA25FDB02F65" box="[508,629,1040,1060]" gridcol="1" gridrow="28" pageId="7" pageNumber="84">63</td>
|
||||
<td id="4C4B5D9A0B760058C1BAFA25FD3A2F65" box="[646,767,1040,1060]" gridcol="2" gridrow="28" pageId="7" pageNumber="84">62</td>
|
||||
</tr>
|
||||
<tr id="0F9A34E60B760058C3BAFA1CFD3A2F7C" box="[134,767,1065,1085]" gridrow="29" pageId="7" pageNumber="84" rowspan-1="1" rowspan-2="1">
|
||||
<th id="4C4B5D9A0B760058C3BAFA1CFE242F7C" box="[134,481,1065,1085]" gridcol="0" gridrow="29" pageId="7" pageNumber="84">length of rostrum</th>
|
||||
</tr>
|
||||
</table>
|
||||
</paragraph>
|
||||
<paragraph id="B11536A40B76FFA0C3B8FA42FEAC2EA6" blockId="7.[131,777,1142,1511]" pageId="7" pageNumber="84">
|
||||
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|
||||
<collectionCode id="D7BBAE610B76FFA0C3B8FA42FF702FCD" box="[132,181,1143,1166]" pageId="7" pageNumber="84">TYPE</collectionCode>
|
||||
</smallCapsWord>
|
||||
<smallCapsWord id="B7F3A0780B76FFA0C380FA49FED22FCC" baselines="1161" box="[188,279,1148,1165]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="horizon" pageId="7" pageNumber="84">
|
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<collectionCode id="D7BBAE610B76FFA0C380FA49FED22FCC" box="[188,279,1148,1165]" pageId="7" pageNumber="84">HORIZON</collectionCode>
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|
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. — The
|
||||
<typeStatus id="6E1188060B76FFA0C253FA43FE002FCF" box="[367,453,1142,1166]" pageId="7" pageNumber="84" type="holotype">holotype</typeStatus>
|
||||
was found in deposits belonging to the silty upper portion of the P2 allomember of the Pisco Formation (as defined by
|
||||
<bibRefCitation id="D53B4B550B76FFA0C25DFA9EFDF42F82" author="DI CELMA C. & MALINVERNO E. & BOSIO G. & COLLARETA A. & GARIBOLDI K. & GIONCADA A. & MOLLI G. & BASSO D. & VARAS-MALCA R. M. & PIERANTONI P. P. & VILLA I. M. & LAMBERT O. & LANDINI W. & SARTI G. & CANTALAMESSA G. & URBINA M. & BIANUCCI G." box="[353,561,1195,1219]" pageId="7" pageNumber="84" pagination="255 - 274" refId="ref17197" refString="DI CELMA C., MALINVERNO E., BOSIO G., COLLARETA A., GARIBOLDI K., GIONCADA A., MOLLI G., BASSO D., VARAS-MALCA R. M., PIERANTONI P. P., VILLA I. M., LAMBERT O., LANDINI W., SARTI G., CANTALAMESSA G., URBINA M. & BIANUCCI G. 2017. - Sequence stratigraphy and paleontology of the upper Miocene Pisco Formation along the western side of the lower Ica valley (Ica Desert, Peru). Rivista Italiana di Paleontologia e Stratigrafia 123: 255 - 274. https: // doi. org / 10.13130 / 2039 - 4942 / 8373" type="journal article" year="2017">
|
||||
Di Celma
|
||||
<emphasis id="83DEEAB60B76FFA0C2F5FA99FE3D2F82" box="[457,504,1195,1219]" italics="true" pageId="7" pageNumber="84">et al.</emphasis>
|
||||
2017
|
||||
</bibRefCitation>
|
||||
, 2018a) (
|
||||
<figureCitation id="29912A210B76FFA0C1AEFA9EFD3E2F82" box="[658,763,1195,1219]" captionStart="FIG" captionStartId="5.[132,143,1804,1821]" captionTargetBox="[137,1451,219,1742]" captionTargetId="figure-17@5.[698,1267,784,1355]" captionTargetPageId="5" captionText="FIG. 1. — A, Map of the major structural trends and basins of coastal Peru. The red dashed rectangle outlines the location of the area shown in detail in panel B. Redrawn and modified from Travis et al. (1976) and Thornburg & Kulm (1981); B, Schematic geological map of the East Pisco Basin, showing the areas of Cenozoic outcrop and the location of the sites where the holotype and paratype of the new physeteroid taxon described herein were found (Cerro Hueco la Zorra and Cerro los Quesos; black stars). Redrawn and modified from DeVries & Schrader (1997); C, Schematic, dip-oriented allostratigraphic diagram for the Pisco Formation and the lower Chilcatay and Otuma formations, showing the stratigraphic position of the two kogiid fossils (MUSM 3405 and MUSM 3291) studied in the present work along with 40Ar/39Ar-dated volcanic ash layers (labelled with their field names on the left side of the diagram) and measured stratigraphic sections published in previous works (Bosio et al. 2020b, and references therein). The Pisco Formation exhibits pronounced thinning to the northeast, with the allomembers onlapping onto the basal composite surface (PE0.0) and arranged in a retrogradational pattern, progressively offset to northeast. Modified from Bosio et al. (2020b).Abbreviations: RN, Roca Negra; CTP, Cerro las Tres Piramides; ULL, Ullujaya; CMH, Cerros la Mama y la Hija; CLZ, Cerros Cadena de los Zanjones; ANF, Anfiteatro; CLQ, Cerro los Quesos; CLQw, Cerro los Quesos west; CLB, Cerro la Bruja; HLZ, Cerro Hueco la Zorra; CBC, Cerro Blanco." pageId="7" pageNumber="84">Fig. 1B, C</figureCitation>
|
||||
). Stratigraphically,
|
||||
<collectionCode id="D7BBAE610B76FFA0C210FAF3FEBC2F9F" box="[300,377,1222,1246]" pageId="7" pageNumber="84">MUSM</collectionCode>
|
||||
3405 is located between the P2-8 and the P2-9 horizons of
|
||||
<bibRefCitation id="D53B4B550B76FFA0C217FAD5FDC72FB9" author="DI CELMA C. & MALINVERNO E. & BOSIO G. & COLLARETA A. & GARIBOLDI K. & GIONCADA A. & MOLLI G. & BASSO D. & VARAS-MALCA R. M. & PIERANTONI P. P. & VILLA I. M. & LAMBERT O. & LANDINI W. & SARTI G. & CANTALAMESSA G. & URBINA M. & BIANUCCI G." box="[299,514,1248,1272]" pageId="7" pageNumber="84" pagination="255 - 274" refId="ref17197" refString="DI CELMA C., MALINVERNO E., BOSIO G., COLLARETA A., GARIBOLDI K., GIONCADA A., MOLLI G., BASSO D., VARAS-MALCA R. M., PIERANTONI P. P., VILLA I. M., LAMBERT O., LANDINI W., SARTI G., CANTALAMESSA G., URBINA M. & BIANUCCI G. 2017. - Sequence stratigraphy and paleontology of the upper Miocene Pisco Formation along the western side of the lower Ica valley (Ica Desert, Peru). Rivista Italiana di Paleontologia e Stratigrafia 123: 255 - 274. https: // doi. org / 10.13130 / 2039 - 4942 / 8373" type="journal article" year="2017">
|
||||
Di Celma
|
||||
<emphasis id="83DEEAB60B76FFA0C2AEFAD4FE042FB9" box="[402,449,1248,1272]" italics="true" pageId="7" pageNumber="84">et al.</emphasis>
|
||||
(2017
|
||||
</bibRefCitation>
|
||||
, 2018a), which are prominent and laterally continuous ledges at the top of sheer cliff faces. At the nearby locality of Cerro Blanco, a tephra layer placed just below the P2-8 horizon has been recently dated to 7.084 ± 0.044 Ma by means of Ar-Ar radiometric analyses (
|
||||
<bibRefCitation id="D53B4B550B76FFA0C121FB7FFD162E23" author="BOSIO G. & MALINVERNO E. & VILLA I. M. & DI CELMA C. & GARIBOLDI K. & GIONCADA A. & BARBERINI B. & URBINA M. & BIANUCCI G." box="[541,723,1354,1378]" pageId="7" pageNumber="84" pagination="213 - 247" refId="ref16115" refString="BOSIO G., MALINVERNO E., VILLA I. M., DI CELMA C., GARIBOLDI K., GIONCADA A., BARBERINI B., URBINA M. & BIANUCCI G. 2020 b. - Tephrochronology and chronostratigraphy of the Miocene Chilcatay and Pisco formations (East Pisco Basin, Peru). Newsletters on Stratigraphy 53 (2): 213 - 247. https: // doi. org / 10.1127 / nos / 2019 / 0525" type="journal article" year="2020">
|
||||
Bosio
|
||||
<emphasis id="83DEEAB60B76FFA0C160FB7EFD482E23" box="[604,653,1354,1378]" italics="true" pageId="7" pageNumber="84">et al.</emphasis>
|
||||
2020b
|
||||
</bibRefCitation>
|
||||
). An age younger than 7.084 Ma is thus proposed for the horizon from which the
|
||||
<typeStatus id="6E1188060B76FFA0C3D5FB4AFEFA2ED6" box="[233,319,1407,1431]" pageId="7" pageNumber="84" type="holotype">holotype</typeStatus>
|
||||
was collected. Considering also that no marine deposit younger than the Miocene is known from the western side of the lower
|
||||
<collectingRegion id="736EF8460B76FFA0C3C2FB81FEDF2E8A" box="[254,282,1460,1483]" country="Peru" name="Ica" pageId="7" pageNumber="84">Ica</collectingRegion>
|
||||
valley,
|
||||
<collectionCode id="D7BBAE610B76FFA0C25DFB81FE6B2E8D" box="[353,430,1460,1484]" pageId="7" pageNumber="84">MUSM</collectionCode>
|
||||
3405 is here regarded as Messinian (latest Miocene) in age.
|
||||
</paragraph>
|
||||
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<collectionCode id="D7BBAE610B76FFA0C3B8F827FF262D69" box="[132,227,1554,1577]" pageId="7" pageNumber="84">LOCALITY</collectionCode>
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<collectionCode id="D7BBAE610B76FFA0C3D7F822FED02D69" box="[235,277,1559,1576]" pageId="7" pageNumber="84">AND</collectionCode>
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<collectionCode id="D7BBAE610B76FFA0C220F822FEB22D69" box="[284,375,1559,1576]" pageId="7" pageNumber="84">HORIZON</collectionCode>
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<collectionCode id="D7BBAE610B76FFA0C2ECF822FDEE2D69" box="[464,555,1559,1576]" pageId="7" pageNumber="84">PARATYPE</collectionCode>
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. — The
|
||||
<typeStatus id="6E1188060B76FFA0C1B4F827FD192D6B" box="[648,732,1554,1578]" pageId="7" pageNumber="84" type="paratype">paratype</typeStatus>
|
||||
was found at Cerro los Quesos (
|
||||
<collectingRegion id="736EF8460B76FFA0C2A8F819FE772D05" box="[404,434,1580,1604]" country="Peru" name="Ica" pageId="7" pageNumber="84">Ica</collectingRegion>
|
||||
desert,
|
||||
<emphasis id="83DEEAB60B76FFA0C2C1F819FDC82D02" box="[509,525,1580,1603]" italics="true" pageId="7" pageNumber="84">c.</emphasis>
|
||||
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|
||||
south of the town of
|
||||
<collectingRegion id="736EF8460B76FFA0C3A2F873FF782D1C" box="[158,189,1606,1629]" country="Peru" name="Ica" pageId="7" pageNumber="84">Ica</collectingRegion>
|
||||
, southern coast of
|
||||
<collectingCountry id="C9BD76340B76FFA0C24FF873FE5A2D1C" box="[371,415,1606,1629]" name="Peru" pageId="7" pageNumber="84">Peru</collectingCountry>
|
||||
(
|
||||
<figureCitation id="29912A210B76FFA0C292F873FDD12D1F" box="[430,532,1606,1630]" captionStart="FIG" captionStartId="5.[132,143,1804,1821]" captionTargetBox="[137,1451,219,1742]" captionTargetId="figure-17@5.[698,1267,784,1355]" captionTargetPageId="5" captionText="FIG. 1. — A, Map of the major structural trends and basins of coastal Peru. The red dashed rectangle outlines the location of the area shown in detail in panel B. Redrawn and modified from Travis et al. (1976) and Thornburg & Kulm (1981); B, Schematic geological map of the East Pisco Basin, showing the areas of Cenozoic outcrop and the location of the sites where the holotype and paratype of the new physeteroid taxon described herein were found (Cerro Hueco la Zorra and Cerro los Quesos; black stars). Redrawn and modified from DeVries & Schrader (1997); C, Schematic, dip-oriented allostratigraphic diagram for the Pisco Formation and the lower Chilcatay and Otuma formations, showing the stratigraphic position of the two kogiid fossils (MUSM 3405 and MUSM 3291) studied in the present work along with 40Ar/39Ar-dated volcanic ash layers (labelled with their field names on the left side of the diagram) and measured stratigraphic sections published in previous works (Bosio et al. 2020b, and references therein). The Pisco Formation exhibits pronounced thinning to the northeast, with the allomembers onlapping onto the basal composite surface (PE0.0) and arranged in a retrogradational pattern, progressively offset to northeast. Modified from Bosio et al. (2020b).Abbreviations: RN, Roca Negra; CTP, Cerro las Tres Piramides; ULL, Ullujaya; CMH, Cerros la Mama y la Hija; CLZ, Cerros Cadena de los Zanjones; ANF, Anfiteatro; CLQ, Cerro los Quesos; CLQw, Cerro los Quesos west; CLB, Cerro la Bruja; HLZ, Cerro Hueco la Zorra; CBC, Cerro Blanco." pageId="7" pageNumber="84">Fig. 1B, C</figureCitation>
|
||||
); geographic coordinates of the finding site:
|
||||
<geoCoordinate id="D49E50630B76FFA0C274F855FE192D39" box="[328,476,1632,1656]" degrees="14" direction="south" minutes="30" orientation="latitude" pageId="7" pageNumber="84" precision="1" seconds="54.00" value="-14.515">14°30’54.00”S</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="D49E50630B76FFA0C2D4F855FD412D39" box="[488,644,1632,1656]" degrees="75" direction="west" minutes="43" orientation="longitude" pageId="7" pageNumber="84" precision="1" seconds="08.90" value="-75.71914">75°43’08.90”W</geoCoordinate>
|
||||
), in deposits belonging to the P2 allomember of the Pisco Formation.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="F9B0652F0B76FFA0C3B8F8A0FBC428AF" pageId="7" pageNumber="84" type="description">
|
||||
<paragraph id="B11536A40B76FFA0C3B8F8A0FBCA2A7F" blockId="7.[131,776,1553,2027]" lastBlockId="7.[811,1457,215,504]" pageId="7" pageNumber="84">
|
||||
MUSM 3291 was indicated with the field number O14 and identified as belonging to “aff.
|
||||
<taxonomicName id="76AA4D270B76FFA0C256F885FE392D89" box="[362,508,1712,1736]" class="Mammalia" family="Kogiidae" genus="Scaphokogia" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="84" phylum="Chordata" rank="species" species="undetermined">
|
||||
<emphasis id="83DEEAB60B76FFA0C256F885FE192D89" box="[362,476,1712,1736]" italics="true" pageId="7" pageNumber="84">Scaphokogia</emphasis>
|
||||
sp.
|
||||
</taxonomicName>
|
||||
” in the fossil map published by
|
||||
<bibRefCitation id="D53B4B550B76FFA0C398F8FFFE432DA3" author="BIANUCCI G. & DI CELMA C. & COLLARETA A. & LANDINI W. & POST K. & TINELLI C. & MUIZON C. DE & BOSIO G. & GARIBOLDI K. & GIONCADA A. & MALINVERNO E. & CANTALAMESSA G. & ALTAMIRANO-SIERRA A. & SALAS-GISMONDI R. & URBINA M. & LAMBERT O." box="[164,390,1738,1762]" pageId="7" pageNumber="84" pagination="1037 - 1046" refId="ref15405" refString="BIANUCCI G., DI CELMA C., COLLARETA A., LANDINI W., POST K., TINELLI C., MUIZON C. DE, BOSIO G., GARIBOLDI K., GIONCADA A., MALINVERNO E., CANTALAMESSA G., ALTAMIRANO-SIERRA A., SALAS-GISMONDI R., URBINA M. & LAMBERT O. 2016 a. - Fossil marine vertebrates of Cerro Los Quesos: Distribution of cetaceans, seals, crocodiles, seabirds, sharks, and bony fish in a late Miocene locality of the Pisco Basin, Peru. Journal of Maps 12: 1037 - 1046. https: // doi. org / 10.1080 / 17445647.2015.1115785" type="journal article" year="2016">
|
||||
Bianucci
|
||||
<emphasis id="83DEEAB60B76FFA0C23CF8FEFEF52DA3" box="[256,304,1738,1762]" italics="true" pageId="7" pageNumber="84">et al.</emphasis>
|
||||
(2016a)
|
||||
</bibRefCitation>
|
||||
. The horizon from which the
|
||||
<typeStatus id="6E1188060B76FFA0C18FF8FEFCC32DA2" box="[691,774,1739,1763]" pageId="7" pageNumber="84" type="paratype">paratype</typeStatus>
|
||||
was collected belongs to the stratal package known as “Member F” (
|
||||
<bibRefCitation id="D53B4B550B76FFA0C3B0F8CAFEAC2C56" author="DI CELMA C. & MALINVERNO E. & CANTALAMESSA G. & GIONCADA A. & BOSIO G. & VILLA I. M. & GARIBOLDI K. & RUSTICHELLI A. & PIERANTONI P. P. & LANDINI W. & TINELLI C. & COLLARETA A. & BIANUCCI G." box="[140,361,1791,1815]" pageId="7" pageNumber="84" pagination="1020 - 1028" refId="ref17012" refString="DI CELMA C., MALINVERNO E., CANTALAMESSA G., GIONCADA A., BOSIO G., VILLA I. M., GARIBOLDI K., RUSTICHELLI A., PIERANTONI P. P., LANDINI W., TINELLI C., COLLARETA A. & BIANUCCI G. 2016 a. - Stratigraphic framework of the late Miocene Pisco Formation at Cerro Los Quesos (Ica Desert, Peru). Journal of Maps 12: 1020 - 1028. https: // doi. org / 10.108 0 / 17445647.2015.1115783." type="journal article" year="2016">
|
||||
Di Celma
|
||||
<emphasis id="83DEEAB60B76FFA0C3CAF935FEE32C56" box="[246,294,1791,1815]" italics="true" pageId="7" pageNumber="84">et al.</emphasis>
|
||||
2016a
|
||||
</bibRefCitation>
|
||||
), within the silty upper portion of the P2 allomember; it is further located
|
||||
<emphasis id="83DEEAB60B76FFA0C2F5F92FFE1D2C70" box="[457,472,1818,1841]" italics="true" pageId="7" pageNumber="84">c.</emphasis>
|
||||
<quantity id="76529B410B76FFA0C2E2F92CFDD72C70" box="[478,530,1817,1841]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="7" pageNumber="84" unit="m" value="35.0">35 m</quantity>
|
||||
above a tephra layer, the Mono key bed of
|
||||
<bibRefCitation id="D53B4B550B76FFA0C20AF901FDE52C0D" author="DI CELMA C. & MALINVERNO E. & CANTALAMESSA G. & GIONCADA A. & BOSIO G. & VILLA I. M. & GARIBOLDI K. & RUSTICHELLI A. & PIERANTONI P. P. & LANDINI W. & TINELLI C. & COLLARETA A. & BIANUCCI G." box="[310,544,1844,1868]" pageId="7" pageNumber="84" pagination="1020 - 1028" refId="ref17012" refString="DI CELMA C., MALINVERNO E., CANTALAMESSA G., GIONCADA A., BOSIO G., VILLA I. M., GARIBOLDI K., RUSTICHELLI A., PIERANTONI P. P., LANDINI W., TINELLI C., COLLARETA A. & BIANUCCI G. 2016 a. - Stratigraphic framework of the late Miocene Pisco Formation at Cerro Los Quesos (Ica Desert, Peru). Journal of Maps 12: 1020 - 1028. https: // doi. org / 10.108 0 / 17445647.2015.1115783." type="journal article" year="2016">
|
||||
Di Celma
|
||||
<emphasis id="83DEEAB60B76FFA0C2A1F900FE082C0D" box="[413,461,1844,1868]" italics="true" pageId="7" pageNumber="84">et al.</emphasis>
|
||||
(2016a)
|
||||
</bibRefCitation>
|
||||
, that has been dated to 6.93 ± 0.09 Ma by means of Ar-Ar radiometric analyses. A stratigraphically higher volcanic ash bed, cropping out in the uppermost portion of the Cerro Los Quesos succession (i.e.,
|
||||
<quantity id="76529B410B76FFA0C150F9B1FD652CDA" box="[620,672,1924,1948]" metricMagnitude="1" metricUnit="m" metricValue="1.7" pageId="7" pageNumber="84" unit="m" value="17.0">17 m</quantity>
|
||||
above the
|
||||
<typeStatus id="6E1188060B76FFA0C3B9F9ABFF1A2CF7" box="[133,223,1950,1974]" pageId="7" pageNumber="84" type="paratype">paratype</typeStatus>
|
||||
), has been dated to an age equal or older than 6.71 ± 0.02 Ma (
|
||||
<bibRefCitation id="D53B4B550B76FFA0C3D8F98DFE7C2C91" author="DI CELMA C. & MALINVERNO E. & CANTALAMESSA G. & GIONCADA A. & BOSIO G. & VILLA I. M. & GARIBOLDI K. & RUSTICHELLI A. & PIERANTONI P. P. & LANDINI W. & TINELLI C. & COLLARETA A. & BIANUCCI G." box="[228,441,1976,2000]" pageId="7" pageNumber="84" pagination="1020 - 1028" refId="ref17012" refString="DI CELMA C., MALINVERNO E., CANTALAMESSA G., GIONCADA A., BOSIO G., VILLA I. M., GARIBOLDI K., RUSTICHELLI A., PIERANTONI P. P., LANDINI W., TINELLI C., COLLARETA A. & BIANUCCI G. 2016 a. - Stratigraphic framework of the late Miocene Pisco Formation at Cerro Los Quesos (Ica Desert, Peru). Journal of Maps 12: 1020 - 1028. https: // doi. org / 10.108 0 / 17445647.2015.1115783." type="journal article" year="2016">
|
||||
Di Celma
|
||||
<emphasis id="83DEEAB60B76FFA0C274F98CFEB32C91" box="[328,374,1976,2000]" italics="true" pageId="7" pageNumber="84">et al.</emphasis>
|
||||
2016a
|
||||
</bibRefCitation>
|
||||
). As such, the horizon from which the
|
||||
<typeStatus id="6E1188060B76FFA0C395F9E6FF392CAA" box="[169,252,2003,2027]" pageId="7" pageNumber="84" type="paratype">paratype</typeStatus>
|
||||
originates can be constrained to a short time interval within the Messinian, i.e., between 6.93 Ma and 6.71 Ma. Such an age estimate finds support in the biostratigraphic data, which indicate an age younger than 7.35 Ma for this portion of Member F (
|
||||
<bibRefCitation id="D53B4B550B76FFA0C009FF13FBC42A7F" author="GARIBOLDI K. & BOSIO G. & MALINVERNO E. & GIONCADA A. & DI CELMA C. & VILLA I. M. & URBINA M. & BIANUCCI G." box="[821,1025,294,318]" pageId="7" pageNumber="84" pagination="417 - 444" refId="ref18233" refString="GARIBOLDI K., BOSIO G., MALINVERNO E., GIONCADA A., DI CELMA C. & VILLA I. M., URBINA M. & BIANUCCI G. 2017. - Biostratigraphy, geochronology and sedimentation rates of the upper Miocene Pisco Formation at two important marine vertebrate fossil-bearing sites of southern Peru. Newsletters on Stratigraphy 50: 417 - 444. https: // doi. org / 10.1127 / nos / 2017 / 0345" type="journal article" year="2017">
|
||||
Gariboldi
|
||||
<emphasis id="83DEEAB60B76FFA0C0A6FF12FC0C2A7F" box="[922,969,294,318]" italics="true" pageId="7" pageNumber="84">et al.</emphasis>
|
||||
2017
|
||||
</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="B11536A40B76FFA0C011FF74FBC32AB9" blockId="7.[811,1457,215,504]" pageId="7" pageNumber="84">
|
||||
Several fossil vertebrates from Cerro Los Quesos were found in strata belonging to Member F, including mysticetes (balaenopterids and cetotheriids), odontocetes (e.g., the
|
||||
<typeStatus id="6E1188060B76FFA0C7B2FF43FB212ACF" box="[1166,1252,374,398]" pageId="7" pageNumber="84" type="holotype">holotype</typeStatus>
|
||||
of the beaked whale
|
||||
<taxonomicName id="76AA4D270B76FFA0C011FFA5FB8F2AE9" authorityName="Bianucci, Di Celma, Urbina & Lambert" authorityYear="2016" box="[813,1098,400,424]" class="Mammalia" family="Hyperoodontidae" genus="Chavinziphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="84" phylum="Chordata" rank="species" species="maxillocristatus">
|
||||
<emphasis id="83DEEAB60B76FFA0C011FFA5FB8F2AE9" box="[813,1098,400,424]" italics="true" pageId="7" pageNumber="84">Chavinziphius maxillocristatus</emphasis>
|
||||
</taxonomicName>
|
||||
, the macroraptorial stem physeteroid
|
||||
<taxonomicName id="76AA4D270B76FFA0C017FF9FFC012A83" box="[811,964,426,450]" class="Mammalia" genus="Acrophyseter" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="84" phylum="Chordata" rank="species" species="undetermined">
|
||||
<emphasis id="83DEEAB60B76FFA0C017FF9FFC5B2A83" box="[811,926,426,450]" italics="true" pageId="7" pageNumber="84">Acrophyseter</emphasis>
|
||||
sp.
|
||||
</taxonomicName>
|
||||
, and the phocoenid cf.
|
||||
<taxonomicName id="76AA4D270B76FFA0C78FFF9EFAB52A83" authorityName="Muizon" authorityYear="1986" box="[1203,1392,426,450]" class="Mammalia" family="Phocoenidae" genus="Lomacetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="84" phylum="Chordata" rank="species" species="ginsburgi">
|
||||
<emphasis id="83DEEAB60B76FFA0C78FFF9EFAB52A83" box="[1203,1392,426,450]" italics="true" pageId="7" pageNumber="84">Lomacetus ginsburgi</emphasis>
|
||||
</taxonomicName>
|
||||
), pinnipeds, crocodiles, seabirds, and sharks (
|
||||
<bibRefCitation id="D53B4B550B76FFA0C781FFF0FA4B2A9C" author="BIANUCCI G. & DI CELMA C. & COLLARETA A. & LANDINI W. & POST K. & TINELLI C. & MUIZON C. DE & BOSIO G. & GARIBOLDI K. & GIONCADA A. & MALINVERNO E. & CANTALAMESSA G. & ALTAMIRANO-SIERRA A. & SALAS-GISMONDI R. & URBINA M. & LAMBERT O." box="[1213,1422,453,477]" pageId="7" pageNumber="84" pagination="1037 - 1046" refId="ref15405" refString="BIANUCCI G., DI CELMA C., COLLARETA A., LANDINI W., POST K., TINELLI C., MUIZON C. DE, BOSIO G., GARIBOLDI K., GIONCADA A., MALINVERNO E., CANTALAMESSA G., ALTAMIRANO-SIERRA A., SALAS-GISMONDI R., URBINA M. & LAMBERT O. 2016 a. - Fossil marine vertebrates of Cerro Los Quesos: Distribution of cetaceans, seals, crocodiles, seabirds, sharks, and bony fish in a late Miocene locality of the Pisco Basin, Peru. Journal of Maps 12: 1037 - 1046. https: // doi. org / 10.1080 / 17445647.2015.1115785" type="journal article" year="2016">
|
||||
Bianucci
|
||||
<emphasis id="83DEEAB60B76FFA0C626FFF3FA8F2A9C" box="[1306,1354,453,477]" italics="true" pageId="7" pageNumber="84">et al.</emphasis>
|
||||
2016a
|
||||
</bibRefCitation>
|
||||
, b;
|
||||
<bibRefCitation id="D53B4B550B76FFA0C011FFEAFC3D2AB9" author="LAMBERT O. & BIANUCCI G. & MUIZON C. DE" box="[813,1016,479,504]" pageId="7" pageNumber="84" pagination="404 - 474" refId="ref19034" refString="LAMBERT O., BIANUCCI G., & MUIZON C. DE 2017 a. - Macroraptorial sperm whales (Cetacea, Odontoceti, Physeteroidea) from the Miocene of Peru. Zoological Journal of the Linnean Society 179: 404 - 474. https: // doi. org / 10.1111 / zoj. 12456" type="journal article" year="2017">
|
||||
Lambert
|
||||
<emphasis id="83DEEAB60B76FFA0C0BAFFD5FC702AB6" box="[902,949,479,503]" italics="true" pageId="7" pageNumber="84">et al.</emphasis>
|
||||
2017a
|
||||
</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="B11536A40B76FFA0C011FC25FBC02824" blockId="7.[811,1457,527,869]" pageId="7" pageNumber="84">
|
||||
<smallCapsWord id="B7F3A0780B76FFA0C011FC25FC762967" baselines="545,546" box="[813,947,528,551]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Differential" pageId="7" pageNumber="84">DIFFERENTIAL</smallCapsWord>
|
||||
<smallCapsWord id="B7F3A0780B76FFA0C084FC20FBD92967" baselines="546" box="[952,1052,533,550]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="diagnosis" pageId="7" pageNumber="84">DIAGNOSIS</smallCapsWord>
|
||||
. —
|
||||
<taxonomicName id="76AA4D270B76FFA0C778FC3AFAE82966" authority="Collareta & Lambert & Palomino & Urbina & Bianucci, 2020" authorityName="Collareta & Lambert & Palomino & Urbina & Bianucci" authorityYear="2020" box="[1092,1325,527,551]" class="Mammalia" family="Kogiidae" genus="Platyscaphokogia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="84" phylum="Chordata" rank="species" species="landinii" status="n. gen., n. sp.">
|
||||
<emphasis id="83DEEAB60B76FFA0C778FC3AFAE82966" box="[1092,1325,527,551]" italics="true" pageId="7" pageNumber="84">Platyscaphokogia landinii</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="98ED57CD0B76FFA0C60EFC25FA752966" box="[1330,1456,528,552]" pageId="7" pageNumber="84" rank="species">n. gen., n. sp.</taxonomicNameLabel>
|
||||
differs from
|
||||
<taxonomicName id="76AA4D270B76FFA0C097FC1CFB452900" authorityName="Muizon" authorityYear="1988" box="[939,1152,553,577]" class="Mammalia" family="Kogiidae" genus="Scaphokogia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="7" pageNumber="84" phylum="Chordata" rank="species" species="cochlearis">
|
||||
<emphasis id="83DEEAB60B76FFA0C097FC1CFB452900" box="[939,1152,553,577]" italics="true" pageId="7" pageNumber="84">Scaphokogia cochlearis</emphasis>
|
||||
</taxonomicName>
|
||||
by the following combination of characters: in dorsal view, skull roughly drop-shaped, due to the remarkable tapering of the rostrum towards the anterior tip of the skull; sagittal facial crest longer, less displaced laterally, dividing the supracranial basin in two different sub-basins; supracranial basin moderately shallower and slightly more anterodorsally oriented; in lateral view, rostrum vertically much thinner (i.e., dorsoventrally compressed) and not exceeding in height the lateral maxillary crests at its base; long axis of the rostrum projecting anteriorly rather than anteroventrally; in ventral view, presence of larger, well-individualised, likely functional upper dental alveoli (right maxilla presenting 10 alveoli).
|
||||
</paragraph>
|
||||
<paragraph id="B11536A40B76FFA0C011FDA0FAC728EE" blockId="7.[813,1455,917,1006]" box="[813,1282,917,943]" pageId="7" pageNumber="84">DESCRIPTION, COMPARISONS, AND REMARKS</paragraph>
|
||||
<paragraph id="B11536A40B76FFA0C011FD80FBC428AF" blockId="7.[813,1455,917,1006]" pageId="7" pageNumber="84">
|
||||
The following description is based on the
|
||||
<typeStatus id="6E1188060B76FFA0C7C2FD80FAA5288E" box="[1278,1376,949,975]" pageId="7" pageNumber="84" type="holotype">holotype</typeStatus>
|
||||
, unless otherwise specified.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
145
data/39/03/87/390387B20B77FFA1C0D2FDA0FA5A2E1F.xml
Normal file
145
data/39/03/87/390387B20B77FFA1C0D2FDA0FA5A2E1F.xml
Normal file
|
|
@ -0,0 +1,145 @@
|
|||
<document id="379DD00B0039B9F311FF3836F7FAC1EC" ID-DOI="10.5852/cr-palevol2020v19a5" ID-ISSN="1777-571X" ID-ZooBank="urn:lsid:zoobank.org:pub:DB4BC4A0-E559-49AD-9452-55F1688D99DA" checkinTime="1732276064017" checkinUser="felipe" docAuthor="Collareta, Alberto, Lambert, Olivier, Palomino, Aldo Marcelo Benites, Urbina, Mario & Bianucci, Giovanni" docDate="2020" docId="390387B20B77FFA1C0D2FDA0FA5A2E1F" docLanguage="en" docName="CRPalevol.19.5.79-100.pdf" docOrigin="Comptes Rendus Palevol 19 (5)" docSource="http://dx.doi.org/10.5852/cr-palevol2020v19a5" docStyle="DocumentStyle:DACF86F01658A8850E13A63D98C287FD.1:CRPalevol.2020-.journal_article" docStyleId="DACF86F01658A8850E13A63D98C287FD" docStyleName="CRPalevol.2020-.journal_article" docStyleVersion="1" docTitle="Platyscaphokogia Collareta & Lambert & Palomino & Urbina & Bianucci 2020, n. gen." docType="treatment" docUuid="F764D6F6-9E50-4CA8-8205-941A2E7CE7E9" docUuidSource="ZooBank" docVersion="1" lastPageNumber="83" masterDocId="C53AFFCA0B71FFA7C33CFE35FFC52B41" masterDocTitle="A new physeteroid from the late Miocene of Peru expands the diversity of extinct dwarf and pygmy sperm whales (Cetacea: Odontoceti: Kogiidae)" masterLastPageNumber="100" masterPageNumber="79" pageNumber="83" updateTime="1732276069354" updateUser="felipe" zenodo-license-document="CC-BY-4.0">
|
||||
<mods:mods id="0661C15627E211B732DC8C47D310C251" xmlns:mods="http://www.loc.gov/mods/v3">
|
||||
<mods:titleInfo id="3EF38AA42ECA8FF57EBDDAF363D58F5B">
|
||||
<mods:title id="1C118B024BF6C1B3B59823D7F3BA3349">A new physeteroid from the late Miocene of Peru expands the diversity of extinct dwarf and pygmy sperm whales (Cetacea: Odontoceti: Kogiidae)</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:name id="DEA213D26761CF9F4EED42183FD13EB5" type="personal">
|
||||
<mods:role id="F3BC714C0FD1C3F20BACEA502CC8DC8C">
|
||||
<mods:roleTerm id="E14679442F26FAC7F51C92DDE7B14269">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="9C066D54C64D39FE68C89A49CEF171FC">Collareta, Alberto</mods:namePart>
|
||||
<mods:affiliation id="9662D49E2075B70B01CE0AB4C8F95D7F">Dipartimento di Scienze della Terra, Università di Pisa, via Santa Maria 53, 56126 Pisa (Italy) alberto. collareta @ unipi. it (corresponding author)</mods:affiliation>
|
||||
<mods:nameIdentifier id="CB3B70C09B27023E28138C6200AD0670" type="email">alberto.collareta@unipi.it</mods:nameIdentifier>
|
||||
</mods:name>
|
||||
<mods:name id="449F98C057646870CE4974E559A11B0E" type="personal">
|
||||
<mods:role id="223113F8E5F2DD1D6FB4637C5079EC50">
|
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<mods:roleTerm id="EF9406484768E852AF2686A26631534B">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="2F2F9E2566B6650210A479E8A703424F">Lambert, Olivier</mods:namePart>
|
||||
<mods:affiliation id="5A0FFC418B2715A1279FA68F583C4B25">Institut Royal des Sciences Naturelles de Belgique, D. O. Terre et Histoire de la Vie, rue Vautier 29, 1000 Brussels (Belgium) Centre de Recherches sur la Paléobiodiversité et les Paléoenvironnements, CR 2 P (CNRS, MNHN, UPMC, Sorbonne Université), Département Origines et Évolution, Muséum national d’Histoire naturelle, case postale 38, 57 rue Cuvier, F- 75231 Paris cedex 05 (France)</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:name id="39D63BDC1D2C361C3BC134865A385933" type="personal">
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</mods:role>
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<mods:namePart id="7230677BBA0B8B2916D228135DFC4CE3">Palomino, Aldo Marcelo Benites</mods:namePart>
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||||
<mods:affiliation id="D425564976749331EEDB0326F1330163">Departamento de Paleontologia de Vertebrados, Museo de Historia Natural de la Universidad Nacional Mayor de San Marcos, avenida Arenales 1256, Lima 14 (Peru) and Paläontologisches Institut und Museum, Karl-Schmid-Strasse 4, CH- 8006 Zürich (Switzerland) and Center for Tropical Paleoecology and Archeology, Smithsonian Tropical Research Institute, Balboa, Ancón, 0843 - 03092, Panama (Panama)</mods:affiliation>
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</mods:name>
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<mods:roleTerm id="AF173EB60FDD760C09027B09739AE28F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="466DB021CAEF071BCAC417AB1A323C69">Urbina, Mario</mods:namePart>
|
||||
<mods:affiliation id="70BF0613C291CE13B36C340DA2C564B2">Departamento de Paleontologia de Vertebrados, Museo de Historia Natural de la Universidad Nacional Mayor de San Marcos, avenida Arenales 1256, Lima 14 (Peru)</mods:affiliation>
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</mods:name>
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<mods:roleTerm id="4950063261F51F76B16A5BE2AF5DF292">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="1D543AA3FEE970AA902880B23B04BC55">Bianucci, Giovanni</mods:namePart>
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<mods:affiliation id="746E424EEF593EAC1B64CF95C41A412F">Dipartimento di Scienze della Terra, Università di Pisa, via Santa Maria 53, 56126 Pisa (Italy)</mods:affiliation>
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<mods:typeOfResource id="89A2A06CAD3138A7B15AD58155E57C97">text</mods:typeOfResource>
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<mods:title id="B1C7E15EBE27E599724D0426188EBF94">Comptes Rendus Palevol</mods:title>
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</mods:titleInfo>
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<mods:part id="D25BBCB429B6DE677C7ED9091EBEF174">
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<mods:date id="D6353E7538021369C061C67782A87439">2020</mods:date>
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<mods:number id="3B140874BDAD1FFD627D4931067CB0D1">2020-10-05</mods:number>
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<mods:number id="C8D6F6668642BFC33E400412C0BFBBF3">19</mods:number>
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</mods:detail>
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<mods:detail id="8BD0E7E3A5ACCF10C4A3E5253AF38CB9" type="issue">
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<mods:number id="EFAE987F8601F9B7896832D4EB365EA6">5</mods:number>
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<mods:start id="22B2799E538111C3E99DE8AAE53C9E40">79</mods:start>
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<mods:end id="367431756B33600866DD6D9E1170504E">100</mods:end>
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<mods:location id="A302EA86069015FC608134A8AC2DF25D">
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<mods:url id="E8575E7BBBFBE3AE5E61068365938226">http://dx.doi.org/10.5852/cr-palevol2020v19a5</mods:url>
|
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</mods:location>
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<mods:classification id="A3F44BCAC1AC6BAB9D067C2B84B823EA">journal article</mods:classification>
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<mods:identifier id="8CA6ABF0C330EB3D9A2E9DBE271B5395" type="DOI">10.5852/cr-palevol2020v19a5</mods:identifier>
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<mods:identifier id="1142EB6FD727F9308B067027D60F4242" type="ISSN">1777-571X</mods:identifier>
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<subSubSection id="F9B0652F0B77FFA1C0D2FDA0FA6A28A9" pageId="6" pageNumber="83" type="nomenclature">
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<paragraph id="B11536A40B77FFA1C0D2FDA0FB2B28EE" blockId="6.[1006,1262,917,943]" box="[1006,1262,917,943]" pageId="6" pageNumber="83">
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<taxonomicName id="76AA4D270B77FFA1C0D2FDA0FB5928EE" authority="Collareta & Lambert & Palomino & Urbina & Bianucci, 2020" authorityName="Collareta & Lambert & Palomino & Urbina & Bianucci" authorityYear="2020" box="[1006,1180,917,943]" class="Mammalia" family="Kogiidae" genus="Platyscaphokogia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="83" phylum="Chordata" rank="genus" status="gen. nov.">
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<emphasis id="83DEEAB60B77FFA1C0D2FDA0FB5928EE" box="[1006,1180,917,943]" italics="true" pageId="6" pageNumber="83">Platyscaphokogia</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="B11536A40B77FFA1C011FDE2FA6A28A9" blockId="6.[813,1455,983,1000]" box="[813,1455,983,1000]" pageId="6" pageNumber="83">
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<uri id="C53B3AA60B77FFA1C011FDE2FA6A28A9" box="[813,1455,983,1000]" pageId="6" pageNumber="83">
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urn:lsid:zoobank.org:act:
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<uuid id="C50C0C710B77FFA1C737FDE2FA6A28A9" box="[1035,1455,983,1000]" pageId="6" pageNumber="83">F764D6F6-9E50-4CA8-8205-941A2E7CE7E9</uuid>
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</uri>
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</subSubSection>
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<subSubSection id="F9B0652F0B77FFA1C011FA2CFBD22FDB" pageId="6" pageNumber="83" type="etymology">
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. — From the combination of the Ancient Greek adjective “platús” (πλατύς), meaning “flat”, with “
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<taxonomicName id="76AA4D270B77FFA1C63CFA06FAB62F0A" authorityName="Muizon" authorityYear="1988" box="[1280,1395,1075,1099]" class="Mammalia" family="Kogiidae" genus="Scaphokogia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="83" phylum="Chordata" rank="genus">
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<emphasis id="83DEEAB60B77FFA1C63CFA06FAB62F0A" box="[1280,1395,1075,1099]" italics="true" pageId="6" pageNumber="83">Scaphokogia</emphasis>
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</taxonomicName>
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”, i.e., the name of the
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genus of
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<taxonomicName id="76AA4D270B77FFA1C75BFA78FAC32F24" authorityName="Muizon" authorityYear="1988" box="[1127,1286,1101,1125]" class="Mammalia" family="Kogiidae" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="96" phylum="Chordata" rank="subFamily" subFamily="Scaphokogiinae">Scaphokogiinae</taxonomicName>
|
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, for highlighting the dorsoventrally flattened morphology of the rostrum of the new form. Gender feminine.
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</paragraph>
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</subSubSection>
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<subSubSection id="F9B0652F0B77FFA1C011FA87FC772FA5" pageId="6" pageNumber="83" type="materials_examined">
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. —
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<taxonomicName id="76AA4D270B77FFA1C78EFA87FA742F8B" authority="Collareta & Lambert & Palomino & Urbina & Bianucci, 2020" authorityName="Collareta & Lambert & Palomino & Urbina & Bianucci" authorityYear="2020" box="[1202,1457,1202,1226]" class="Mammalia" family="Kogiidae" genus="Platyscaphokogia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="83" phylum="Chordata" rank="species" species="landinii" status="n. gen., n. sp.">
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<emphasis id="83DEEAB60B77FFA1C78EFA87FA742F8B" box="[1202,1457,1202,1226]" italics="true" pageId="6" pageNumber="83">Platyscaphokogia landinii</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="98ED57CD0B77FFA1C011FAF8FC772FA5" box="[813,946,1229,1253]" pageId="6" pageNumber="83" rank="species">n. gen., n. sp.</taxonomicNameLabel>
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</paragraph>
|
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</subSubSection>
|
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<subSubSection id="F9B0652F0B77FFA1C011FAC9FA5A2E1F" pageId="6" pageNumber="83" type="diagnosis">
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||||
. — Same as for the
|
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species until other species are described.
|
||||
</paragraph>
|
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<paragraph id="B11536A40B77FFA1C011FB72FA5A2E1F" blockId="6.[813,1439,1350,1374]" box="[813,1439,1350,1374]" pageId="6" pageNumber="83">
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<smallCapsWord id="B7F3A0780B77FFA1C011FB72FCB82E1C" baselines="1368,1369" box="[813,893,1351,1374]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Known" pageId="6" pageNumber="83">KNOWN</smallCapsWord>
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<smallCapsWord id="B7F3A0780B77FFA1C0BFFB79FC062E1C" baselines="1369" box="[899,963,1356,1373]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="range" pageId="6" pageNumber="83">RANGE</smallCapsWord>
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||||
. — Late Miocene (Messinian) of southern
|
||||
<collectingCountry id="C9BD76340B77FFA1C657FB72FA5E2E1F" box="[1387,1435,1351,1374]" name="Peru" pageId="6" pageNumber="83">Peru</collectingCountry>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
Loading…
Reference in a new issue