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<taxonomicName LSID="http://zoobank.org/C72889DC-5B7A-4366-B703-964353942786" authority="Czaja & Gladstone" authorityName="Czaja & Gladstone" authorityYear="2021" class="Gastropoda" family="Cochliopidae" genus="Phreatoviesca" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phreatoviesca spinosa" order="Littorinimorpha" pageId="0" pageNumber="129" phylum="Mollusca" rank="species" species="spinosa" status="sp. nov.">Phreatoviesca spinosa Czaja & Gladstone</taxonomicName>
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<taxonomicName id="E9A76AEFA39A4DF8C27026E949A8AF83" ID-CoL="8NF5H" LSID="http://zoobank.org/C72889DC-5B7A-4366-B703-964353942786" authority="Czaja & Gladstone" authorityName="Czaja & Gladstone" authorityYear="2021" class="Gastropoda" family="Cochliopidae" genus="Phreatoviesca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phreatoviesca spinosa" order="Littorinimorpha" pageId="0" pageNumber="129" phylum="Mollusca" rank="species" species="spinosa" status="sp. nov.">Phreatoviesca spinosa Czaja & Gladstone</taxonomicName>
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<figureCitation captionStart="Figures 2–13" captionStartId="F2" captionText="Figures 2 - 13. Shells of Phreatoviesca spinosa gen. nov. et sp. nov. 2, 3 holotype, specimen from both sides, UJMC 500 4, 5 paratype 1, specimen from both sides, UJMC 501 6 paratype 2, specimen with a ' corkscrew' - like morphology, UJMC 502 7, 8 paratype 3, specimen from both sides, UJMC 503 9 paratype 4, specimen with smooth whorls and a trumpet-like aperture, UJMC 504 10, 11 paratype 5, specimen with smooth whorls, UJMC 505. Opercula 12, 13 paratype 5, specimen with smooth whorls, UJMC 505. Scale bar: 1 mm." figureDoi="10.3897/subtbiol.39.67799.figures2-13" httpUri="https://binary.pensoft.net/fig/574504" pageId="0" pageNumber="129">Figs 2-13</figureCitation>
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<figureCitation captionStart="Figures 14–17" captionStartId="F3" captionText="Figures 14 - 17. SEM images of Phreatoviesca spinosa gen. nov. et sp. nov. 14 specimen with strong spines, UJMC 506 15 specimen with ribs, UJMC 507 16 specimen with smooth whorls, UJMC 508 17 specimen with smooth whorls, UJMC 509. Scale bar: 1 mm." figureDoi="10.3897/subtbiol.39.67799.figures14-17" httpUri="https://binary.pensoft.net/fig/574505" pageId="0" pageNumber="129">, 14-17</figureCitation>
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<figureCitation captionStart="Figures 18–24" captionStartId="F4" captionText="Figures 18 - 24. SEM images of Phreatoviesca spinosa gen. nov. et sp. nov. 18 shell apex with protoconch, UJMC 508 19 paratype 3, shell apex with protoconch, UJMC 503 20 shell apex with protoconch, UJMC 510 21 paratype 1, apex with protoconch, UJMC 501 22 smooth specimen with body whorl shows irregular, strong marked growth lines, UJMC 508 23 paratype 1, body whorl shows irregular, strong marked growth lines, UJMC 501 24 paratype 2, body whorl shows regularly spaced shovel-shaped spines, UJMC 502." figureDoi="10.3897/subtbiol.39.67799.figures18-24" httpUri="https://binary.pensoft.net/fig/574506" pageId="0" pageNumber="129">, 18-24</figureCitation>
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<subSubSection id="B67B295A1F7E980860D1124CD25BD404" pageId="0" pageNumber="129" type="description">
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<figureCitation id="8752CB78342C865C31E25E617D33336C" captionStart="Figures 2–13" captionStartId="F2" captionText="Figures 2 - 13. Shells of Phreatoviesca spinosa gen. nov. et sp. nov. 2, 3 holotype, specimen from both sides, UJMC 500 4, 5 paratype 1, specimen from both sides, UJMC 501 6 paratype 2, specimen with a ' corkscrew' - like morphology, UJMC 502 7, 8 paratype 3, specimen from both sides, UJMC 503 9 paratype 4, specimen with smooth whorls and a trumpet-like aperture, UJMC 504 10, 11 paratype 5, specimen with smooth whorls, UJMC 505. Opercula 12, 13 paratype 5, specimen with smooth whorls, UJMC 505. Scale bar: 1 mm." figureDoi="10.3897/subtbiol.39.67799.figures2-13" httpUri="https://binary.pensoft.net/fig/574504" pageId="0" pageNumber="129">Figs 2-13</figureCitation>
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<figureCitation id="42A92557C633906E7B01F8F91B12919A" captionStart="Figures 14–17" captionStartId="F3" captionText="Figures 14 - 17. SEM images of Phreatoviesca spinosa gen. nov. et sp. nov. 14 specimen with strong spines, UJMC 506 15 specimen with ribs, UJMC 507 16 specimen with smooth whorls, UJMC 508 17 specimen with smooth whorls, UJMC 509. Scale bar: 1 mm." figureDoi="10.3897/subtbiol.39.67799.figures14-17" httpUri="https://binary.pensoft.net/fig/574505" pageId="0" pageNumber="129">, 14-17</figureCitation>
|
||||
<figureCitation id="7AF97132972CB29463E879AA586E14E0" captionStart="Figures 18–24" captionStartId="F4" captionText="Figures 18 - 24. SEM images of Phreatoviesca spinosa gen. nov. et sp. nov. 18 shell apex with protoconch, UJMC 508 19 paratype 3, shell apex with protoconch, UJMC 503 20 shell apex with protoconch, UJMC 510 21 paratype 1, apex with protoconch, UJMC 501 22 smooth specimen with body whorl shows irregular, strong marked growth lines, UJMC 508 23 paratype 1, body whorl shows irregular, strong marked growth lines, UJMC 501 24 paratype 2, body whorl shows regularly spaced shovel-shaped spines, UJMC 502." figureDoi="10.3897/subtbiol.39.67799.figures18-24" httpUri="https://binary.pensoft.net/fig/574506" pageId="0" pageNumber="129">, 18-24</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="129" type="type locality">
|
||||
<paragraph pageId="0" pageNumber="129">Type locality.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="129">
|
||||
<subSubSection id="D8B0DE684F790F1615EEFA32659C1037" pageId="0" pageNumber="129" type="type locality">
|
||||
<paragraph id="8F2FD1458566C2F9BC9CA07E6259BF9D" pageId="0" pageNumber="129">Type locality.</paragraph>
|
||||
<paragraph id="BCD8077677A1D5E7E1153B695FF0AE82" pageId="0" pageNumber="129">
|
||||
Mexico, Coahuila state,
|
||||
<taxonomicName class="Gastropoda" genus="Viesca" kingdom="Animalia" lsidName="Viesca" pageId="0" pageNumber="129" phylum="Mollusca" rank="genus">Viesca</taxonomicName>
|
||||
<taxonomicName id="BC072D451866B9B287AFCF82E1ECC2F0" class="Gastropoda" genus="Viesca" kingdom="Animalia" lsidName="Viesca" pageId="0" pageNumber="129" phylum="Mollusca" rank="genus">Viesca</taxonomicName>
|
||||
, spring
|
||||
<normalizedToken originalValue="“Túnel">"Tunel</normalizedToken>
|
||||
<normalizedToken id="48C30B5CDB91B0A505FF587CDDFA12D3" originalValue="“Túnel">"Tunel</normalizedToken>
|
||||
7" (
|
||||
<geoCoordinate degrees="25" direction="north" minutes="20" orientation="latitude" precision="15" seconds="38" value="25.34389">25°20'38"N</geoCoordinate>
|
||||
<geoCoordinate id="9DED511ED5823B27F0557B80C5AB6C24" degrees="25" direction="north" minutes="20" orientation="latitude" precision="15" seconds="38" value="25.34389">25°20'38"N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="102" direction="west" minutes="54" orientation="longitude" precision="15" seconds="19" value="-102.90528">102°54'19"W</geoCoordinate>
|
||||
<geoCoordinate id="C0033B4F413FDF04653C92AB94012702" degrees="102" direction="west" minutes="54" orientation="longitude" precision="15" seconds="19" value="-102.90528">102°54'19"W</geoCoordinate>
|
||||
, 1102 m a.s.l.) (Fig.
|
||||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map of the study area with localization of the sampling site near the town of Viesca, Coahuila, Mexico." figureDoi="10.3897/subtbiol.39.67799.figure1" httpUri="https://binary.pensoft.net/fig/574503" pageId="0" pageNumber="129">1</figureCitation>
|
||||
<figureCitation id="4BAE1EA7775592C9F708352E8B743E4B" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map of the study area with localization of the sampling site near the town of Viesca, Coahuila, Mexico." figureDoi="10.3897/subtbiol.39.67799.figure1" httpUri="https://binary.pensoft.net/fig/574503" pageId="0" pageNumber="129">1</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="129" type="types">
|
||||
<paragraph pageId="0" pageNumber="129">Types.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="129">
|
||||
<subSubSection id="67669F4145F62073BEFFF21F9B6DB967" pageId="0" pageNumber="129" type="types">
|
||||
<paragraph id="587C077D65DBEBD3C029575DD3AA8A57" pageId="0" pageNumber="129">Types.</paragraph>
|
||||
<paragraph id="A11C87D88BBAB215F31E9618894C0AA6" pageId="0" pageNumber="129">
|
||||
Holotype (Figs
|
||||
<figureCitation captionStart="Figures 2–13" captionStartId="F2" captionText="Figures 2 - 13. Shells of Phreatoviesca spinosa gen. nov. et sp. nov. 2, 3 holotype, specimen from both sides, UJMC 500 4, 5 paratype 1, specimen from both sides, UJMC 501 6 paratype 2, specimen with a ' corkscrew' - like morphology, UJMC 502 7, 8 paratype 3, specimen from both sides, UJMC 503 9 paratype 4, specimen with smooth whorls and a trumpet-like aperture, UJMC 504 10, 11 paratype 5, specimen with smooth whorls, UJMC 505. Opercula 12, 13 paratype 5, specimen with smooth whorls, UJMC 505. Scale bar: 1 mm." figureDoi="10.3897/subtbiol.39.67799.figures2-13" httpUri="https://binary.pensoft.net/fig/574504" pageId="0" pageNumber="129">2</figureCitation>
|
||||
<figureCitation id="0D68285ADF6C1723C8049281A29815B9" captionStart="Figures 2–13" captionStartId="F2" captionText="Figures 2 - 13. Shells of Phreatoviesca spinosa gen. nov. et sp. nov. 2, 3 holotype, specimen from both sides, UJMC 500 4, 5 paratype 1, specimen from both sides, UJMC 501 6 paratype 2, specimen with a ' corkscrew' - like morphology, UJMC 502 7, 8 paratype 3, specimen from both sides, UJMC 503 9 paratype 4, specimen with smooth whorls and a trumpet-like aperture, UJMC 504 10, 11 paratype 5, specimen with smooth whorls, UJMC 505. Opercula 12, 13 paratype 5, specimen with smooth whorls, UJMC 505. Scale bar: 1 mm." figureDoi="10.3897/subtbiol.39.67799.figures2-13" httpUri="https://binary.pensoft.net/fig/574504" pageId="0" pageNumber="129">2</figureCitation>
|
||||
,
|
||||
<figureCitation captionStart="Figures 2–13" captionStartId="F2" captionText="Figures 2 - 13. Shells of Phreatoviesca spinosa gen. nov. et sp. nov. 2, 3 holotype, specimen from both sides, UJMC 500 4, 5 paratype 1, specimen from both sides, UJMC 501 6 paratype 2, specimen with a ' corkscrew' - like morphology, UJMC 502 7, 8 paratype 3, specimen from both sides, UJMC 503 9 paratype 4, specimen with smooth whorls and a trumpet-like aperture, UJMC 504 10, 11 paratype 5, specimen with smooth whorls, UJMC 505. Opercula 12, 13 paratype 5, specimen with smooth whorls, UJMC 505. Scale bar: 1 mm." figureDoi="10.3897/subtbiol.39.67799.figures2-13" httpUri="https://binary.pensoft.net/fig/574504" pageId="0" pageNumber="129">3</figureCitation>
|
||||
<figureCitation id="5F62297C20D5424A44497E387EC8965F" captionStart="Figures 2–13" captionStartId="F2" captionText="Figures 2 - 13. Shells of Phreatoviesca spinosa gen. nov. et sp. nov. 2, 3 holotype, specimen from both sides, UJMC 500 4, 5 paratype 1, specimen from both sides, UJMC 501 6 paratype 2, specimen with a ' corkscrew' - like morphology, UJMC 502 7, 8 paratype 3, specimen from both sides, UJMC 503 9 paratype 4, specimen with smooth whorls and a trumpet-like aperture, UJMC 504 10, 11 paratype 5, specimen with smooth whorls, UJMC 505. Opercula 12, 13 paratype 5, specimen with smooth whorls, UJMC 505. Scale bar: 1 mm." figureDoi="10.3897/subtbiol.39.67799.figures2-13" httpUri="https://binary.pensoft.net/fig/574504" pageId="0" pageNumber="129">3</figureCitation>
|
||||
), UJMC 500, from type locality, leg. A. Czaja, 15/v/2019. Paratypes, UJMC 501-511, from same lot,>100 dry shells.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="129" type="etymology">
|
||||
<paragraph pageId="0" pageNumber="129">Etymology.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="129">
|
||||
<subSubSection id="A37D92F16351A4440CF4ADEAE5E7DDF8" pageId="0" pageNumber="129" type="etymology">
|
||||
<paragraph id="E006EEE9BD6D51ABB96EA1E3310E9E6C" pageId="0" pageNumber="129">Etymology.</paragraph>
|
||||
<paragraph id="1EF094D679FDFB2CABD5A1ADBEC0A24D" pageId="0" pageNumber="129">
|
||||
Name is derived from the Latin word
|
||||
<taxonomicName authority="Czaja & Gladstone, 2021" authorityName="Czaja & Gladstone" authorityYear="2021" class="Gastropoda" family="Cochliopidae" genus="Phreatoviesca" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phreatoviesca spinosa" order="Littorinimorpha" pageId="0" pageNumber="129" phylum="Mollusca" rank="species" species="spinosa">
|
||||
<emphasis italics="true" pageId="0" pageNumber="129">Phreatoviesca spinosa</emphasis>
|
||||
<taxonomicName id="540A39599C9C5C8D7D5DECEA30091D0B" authorityName="Czaja & Gladstone" authorityYear="2021" class="Gastropoda" family="Cochliopidae" genus="Phreatoviesca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phreatoviesca spinosa" order="Littorinimorpha" pageId="0" pageNumber="129" phylum="Mollusca" rank="species" species="spinosa">
|
||||
<emphasis id="86AC92DB2ED1B2DC753B7B0CD23DA8BE" italics="true" pageId="0" pageNumber="129">Phreatoviesca spinosa</emphasis>
|
||||
</taxonomicName>
|
||||
= having spines.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="129" type="materials_examined">
|
||||
<paragraph pageId="0" pageNumber="129">Referred material.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="129">
|
||||
<subSubSection id="EE5236401A1227BEA658CEFBF6A37414" pageId="0" pageNumber="129" type="materials_examined">
|
||||
<paragraph id="7AFE618F815FBC860C133F2ACE9A7808" pageId="0" pageNumber="129">Referred material.</paragraph>
|
||||
<paragraph id="0849F001770A805383ED716B84A08DCA" pageId="0" pageNumber="129">
|
||||
Coahuila.
|
||||
<taxonomicName class="Gastropoda" genus="Viesca" kingdom="Animalia" lsidName="Viesca" pageId="0" pageNumber="129" phylum="Mollusca" rank="genus">Viesca</taxonomicName>
|
||||
<taxonomicName id="E7D1B9AF8D6CF9FF8D49B3F3303A2F4F" class="Gastropoda" genus="Viesca" kingdom="Animalia" lsidName="Viesca" pageId="0" pageNumber="129" phylum="Mollusca" rank="genus">Viesca</taxonomicName>
|
||||
, Spring
|
||||
<normalizedToken originalValue="“Túnel">"Tunel</normalizedToken>
|
||||
<normalizedToken id="9CA1DF5EFEEA4562899AABFEB3E1E216" originalValue="“Túnel">"Tunel</normalizedToken>
|
||||
7", UJMC 500-511, A. Czaja, J. L.
|
||||
<normalizedToken originalValue="Estrada-Rodríguez">Estrada-Rodriguez</normalizedToken>
|
||||
<date value="2015-06-10">10/vi/2015</date>
|
||||
<normalizedToken id="119CCBB8C504D29A6D7D60EBCE86FBFB" originalValue="Estrada-Rodríguez">Estrada-Rodriguez</normalizedToken>
|
||||
<date id="83AEC21260D8EC55ACB60C99CDD51230" value="2015-06-10">10/vi/2015</date>
|
||||
and
|
||||
<date value="2019-05-15">15/v/2019</date>
|
||||
<date id="FB442F53CA02594CF569C82D11B55891" value="2019-05-15">15/v/2019</date>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="129" type="diagnosis">
|
||||
<paragraph pageId="0" pageNumber="129">Diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="129">Like for the genus.</paragraph>
|
||||
<subSubSection id="C70EF321664BC5D1C1A42033C0351F12" pageId="0" pageNumber="129" type="diagnosis">
|
||||
<paragraph id="AA69F30AD97160E8F56AEEEE35AA5F18" pageId="0" pageNumber="129">Diagnosis.</paragraph>
|
||||
<paragraph id="E1E46A94CDDF528EF7BEB1D541A0220E" pageId="0" pageNumber="129">Like for the genus.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="129" type="description">
|
||||
<paragraph pageId="0" pageNumber="129">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="129">
|
||||
Shell small, conical, white or colorless, sometime with rests of light brown periostracum, yielding diversity in shell form, with 4-
|
||||
<normalizedToken originalValue="5½">51/2</normalizedToken>
|
||||
<subSubSection id="E18E7E46FF00D81075E883AF9A8D1197" pageId="0" pageNumber="129" type="description">
|
||||
<paragraph id="820A4404A5EA45E00B54331B08F4921E" pageId="0" pageNumber="129">Description.</paragraph>
|
||||
<paragraph id="68B2C168FD548E0503620E444AB20F4C" pageId="0" pageNumber="129">
|
||||
Shell small, conical, white or colorless, sometime with rests of light brown periostracum, yielding diversity in shell form, with 4-
|
||||
<normalizedToken id="286E05DEF6654C8A0DA8ABAE2F80A9E5" originalValue="5½">51/2</normalizedToken>
|
||||
rounded whorls (usually 5), whorls increasing in radius, the first three whorls never uncoiled, subsequent whorls open coiled, body whorl always uncoiled, some specimens show a
|
||||
<normalizedToken originalValue="‘corkscrew’">'corkscrew'</normalizedToken>
|
||||
<normalizedToken id="EC5478380BD1D83D2265996F023ACC7A" originalValue="‘corkscrew’">'corkscrew'</normalizedToken>
|
||||
-like morphology (Figs
|
||||
<figureCitation captionStart="Figures 2–13" captionStartId="F2" captionText="Figures 2 - 13. Shells of Phreatoviesca spinosa gen. nov. et sp. nov. 2, 3 holotype, specimen from both sides, UJMC 500 4, 5 paratype 1, specimen from both sides, UJMC 501 6 paratype 2, specimen with a ' corkscrew' - like morphology, UJMC 502 7, 8 paratype 3, specimen from both sides, UJMC 503 9 paratype 4, specimen with smooth whorls and a trumpet-like aperture, UJMC 504 10, 11 paratype 5, specimen with smooth whorls, UJMC 505. Opercula 12, 13 paratype 5, specimen with smooth whorls, UJMC 505. Scale bar: 1 mm." figureDoi="10.3897/subtbiol.39.67799.figures2-13" httpUri="https://binary.pensoft.net/fig/574504" pageId="0" pageNumber="129">3</figureCitation>
|
||||
<figureCitation id="4BB36AB8B18A8A5D9C5D24C0B7036ED8" captionStart="Figures 2–13" captionStartId="F2" captionText="Figures 2 - 13. Shells of Phreatoviesca spinosa gen. nov. et sp. nov. 2, 3 holotype, specimen from both sides, UJMC 500 4, 5 paratype 1, specimen from both sides, UJMC 501 6 paratype 2, specimen with a ' corkscrew' - like morphology, UJMC 502 7, 8 paratype 3, specimen from both sides, UJMC 503 9 paratype 4, specimen with smooth whorls and a trumpet-like aperture, UJMC 504 10, 11 paratype 5, specimen with smooth whorls, UJMC 505. Opercula 12, 13 paratype 5, specimen with smooth whorls, UJMC 505. Scale bar: 1 mm." figureDoi="10.3897/subtbiol.39.67799.figures2-13" httpUri="https://binary.pensoft.net/fig/574504" pageId="0" pageNumber="129">3</figureCitation>
|
||||
,
|
||||
<figureCitation captionStart="Figures 2–13" captionStartId="F2" captionText="Figures 2 - 13. Shells of Phreatoviesca spinosa gen. nov. et sp. nov. 2, 3 holotype, specimen from both sides, UJMC 500 4, 5 paratype 1, specimen from both sides, UJMC 501 6 paratype 2, specimen with a ' corkscrew' - like morphology, UJMC 502 7, 8 paratype 3, specimen from both sides, UJMC 503 9 paratype 4, specimen with smooth whorls and a trumpet-like aperture, UJMC 504 10, 11 paratype 5, specimen with smooth whorls, UJMC 505. Opercula 12, 13 paratype 5, specimen with smooth whorls, UJMC 505. Scale bar: 1 mm." figureDoi="10.3897/subtbiol.39.67799.figures2-13" httpUri="https://binary.pensoft.net/fig/574504" pageId="0" pageNumber="129">6</figureCitation>
|
||||
<figureCitation id="5ADB35A4549EDE36A5FAD3EDB628F335" captionStart="Figures 2–13" captionStartId="F2" captionText="Figures 2 - 13. Shells of Phreatoviesca spinosa gen. nov. et sp. nov. 2, 3 holotype, specimen from both sides, UJMC 500 4, 5 paratype 1, specimen from both sides, UJMC 501 6 paratype 2, specimen with a ' corkscrew' - like morphology, UJMC 502 7, 8 paratype 3, specimen from both sides, UJMC 503 9 paratype 4, specimen with smooth whorls and a trumpet-like aperture, UJMC 504 10, 11 paratype 5, specimen with smooth whorls, UJMC 505. Opercula 12, 13 paratype 5, specimen with smooth whorls, UJMC 505. Scale bar: 1 mm." figureDoi="10.3897/subtbiol.39.67799.figures2-13" httpUri="https://binary.pensoft.net/fig/574504" pageId="0" pageNumber="129">6</figureCitation>
|
||||
), suture deep; teleoconch sculptured with irregular, strong marked growth lines and with ribs (Figs
|
||||
<figureCitation captionStart="Figures 2–13" captionStartId="F2" captionText="Figures 2 - 13. Shells of Phreatoviesca spinosa gen. nov. et sp. nov. 2, 3 holotype, specimen from both sides, UJMC 500 4, 5 paratype 1, specimen from both sides, UJMC 501 6 paratype 2, specimen with a ' corkscrew' - like morphology, UJMC 502 7, 8 paratype 3, specimen from both sides, UJMC 503 9 paratype 4, specimen with smooth whorls and a trumpet-like aperture, UJMC 504 10, 11 paratype 5, specimen with smooth whorls, UJMC 505. Opercula 12, 13 paratype 5, specimen with smooth whorls, UJMC 505. Scale bar: 1 mm." figureDoi="10.3897/subtbiol.39.67799.figures2-13" httpUri="https://binary.pensoft.net/fig/574504" pageId="0" pageNumber="129">4</figureCitation>
|
||||
<figureCitation id="B311C744EE6A70F486FDEA5ED2D7AA14" captionStart="Figures 2–13" captionStartId="F2" captionText="Figures 2 - 13. Shells of Phreatoviesca spinosa gen. nov. et sp. nov. 2, 3 holotype, specimen from both sides, UJMC 500 4, 5 paratype 1, specimen from both sides, UJMC 501 6 paratype 2, specimen with a ' corkscrew' - like morphology, UJMC 502 7, 8 paratype 3, specimen from both sides, UJMC 503 9 paratype 4, specimen with smooth whorls and a trumpet-like aperture, UJMC 504 10, 11 paratype 5, specimen with smooth whorls, UJMC 505. Opercula 12, 13 paratype 5, specimen with smooth whorls, UJMC 505. Scale bar: 1 mm." figureDoi="10.3897/subtbiol.39.67799.figures2-13" httpUri="https://binary.pensoft.net/fig/574504" pageId="0" pageNumber="129">4</figureCitation>
|
||||
,
|
||||
<figureCitation captionStart="Figures 2–13" captionStartId="F2" captionText="Figures 2 - 13. Shells of Phreatoviesca spinosa gen. nov. et sp. nov. 2, 3 holotype, specimen from both sides, UJMC 500 4, 5 paratype 1, specimen from both sides, UJMC 501 6 paratype 2, specimen with a ' corkscrew' - like morphology, UJMC 502 7, 8 paratype 3, specimen from both sides, UJMC 503 9 paratype 4, specimen with smooth whorls and a trumpet-like aperture, UJMC 504 10, 11 paratype 5, specimen with smooth whorls, UJMC 505. Opercula 12, 13 paratype 5, specimen with smooth whorls, UJMC 505. Scale bar: 1 mm." figureDoi="10.3897/subtbiol.39.67799.figures2-13" httpUri="https://binary.pensoft.net/fig/574504" pageId="0" pageNumber="129">5</figureCitation>
|
||||
<figureCitation id="C7B70109672839D5CED4EF2706D95CD6" captionStart="Figures 2–13" captionStartId="F2" captionText="Figures 2 - 13. Shells of Phreatoviesca spinosa gen. nov. et sp. nov. 2, 3 holotype, specimen from both sides, UJMC 500 4, 5 paratype 1, specimen from both sides, UJMC 501 6 paratype 2, specimen with a ' corkscrew' - like morphology, UJMC 502 7, 8 paratype 3, specimen from both sides, UJMC 503 9 paratype 4, specimen with smooth whorls and a trumpet-like aperture, UJMC 504 10, 11 paratype 5, specimen with smooth whorls, UJMC 505. Opercula 12, 13 paratype 5, specimen with smooth whorls, UJMC 505. Scale bar: 1 mm." figureDoi="10.3897/subtbiol.39.67799.figures2-13" httpUri="https://binary.pensoft.net/fig/574504" pageId="0" pageNumber="129">5</figureCitation>
|
||||
), spiny shells with whorls with a peripheral slightly pronounced carinae, ribs at the carina are modified into regularly spaced shovel-shaped spines (Figs
|
||||
<figureCitation captionStart="Figures 14–17" captionStartId="F3" captionText="Figures 14 - 17. SEM images of Phreatoviesca spinosa gen. nov. et sp. nov. 14 specimen with strong spines, UJMC 506 15 specimen with ribs, UJMC 507 16 specimen with smooth whorls, UJMC 508 17 specimen with smooth whorls, UJMC 509. Scale bar: 1 mm." figureDoi="10.3897/subtbiol.39.67799.figures14-17" httpUri="https://binary.pensoft.net/fig/574505" pageId="0" pageNumber="129">14</figureCitation>
|
||||
<figureCitation id="1A6B3489DAA85307794A221DE78FDB4C" captionStart="Figures 14–17" captionStartId="F3" captionText="Figures 14 - 17. SEM images of Phreatoviesca spinosa gen. nov. et sp. nov. 14 specimen with strong spines, UJMC 506 15 specimen with ribs, UJMC 507 16 specimen with smooth whorls, UJMC 508 17 specimen with smooth whorls, UJMC 509. Scale bar: 1 mm." figureDoi="10.3897/subtbiol.39.67799.figures14-17" httpUri="https://binary.pensoft.net/fig/574505" pageId="0" pageNumber="129">14</figureCitation>
|
||||
,
|
||||
<figureCitation captionStart="Figures 18–24" captionStartId="F4" captionText="Figures 18 - 24. SEM images of Phreatoviesca spinosa gen. nov. et sp. nov. 18 shell apex with protoconch, UJMC 508 19 paratype 3, shell apex with protoconch, UJMC 503 20 shell apex with protoconch, UJMC 510 21 paratype 1, apex with protoconch, UJMC 501 22 smooth specimen with body whorl shows irregular, strong marked growth lines, UJMC 508 23 paratype 1, body whorl shows irregular, strong marked growth lines, UJMC 501 24 paratype 2, body whorl shows regularly spaced shovel-shaped spines, UJMC 502." figureDoi="10.3897/subtbiol.39.67799.figures18-24" httpUri="https://binary.pensoft.net/fig/574506" pageId="0" pageNumber="129">24</figureCitation>
|
||||
<figureCitation id="6EE09267D974109EF3A99A5D82386371" captionStart="Figures 18–24" captionStartId="F4" captionText="Figures 18 - 24. SEM images of Phreatoviesca spinosa gen. nov. et sp. nov. 18 shell apex with protoconch, UJMC 508 19 paratype 3, shell apex with protoconch, UJMC 503 20 shell apex with protoconch, UJMC 510 21 paratype 1, apex with protoconch, UJMC 501 22 smooth specimen with body whorl shows irregular, strong marked growth lines, UJMC 508 23 paratype 1, body whorl shows irregular, strong marked growth lines, UJMC 501 24 paratype 2, body whorl shows regularly spaced shovel-shaped spines, UJMC 502." figureDoi="10.3897/subtbiol.39.67799.figures18-24" httpUri="https://binary.pensoft.net/fig/574506" pageId="0" pageNumber="129">24</figureCitation>
|
||||
), transition protoconch/teleoconch distinct, marked by a change in surface texture from pitted to axial growth lines, whorls rapidly increasing in diameter, first two whorls smooth, without carina or spines, the last three whorls with increasing number spines (up to 40 on the body whorl, but usually less than 30), spiral lines beginning at the end of protoconch, a few specimens with smooth whorls without any sculpture but with thickened axial growth lines, some (smooth) specimens with a varix just behind the aperture (Fig.
|
||||
<figureCitation captionStart="Figures 2–13" captionStartId="F2" captionText="Figures 2 - 13. Shells of Phreatoviesca spinosa gen. nov. et sp. nov. 2, 3 holotype, specimen from both sides, UJMC 500 4, 5 paratype 1, specimen from both sides, UJMC 501 6 paratype 2, specimen with a ' corkscrew' - like morphology, UJMC 502 7, 8 paratype 3, specimen from both sides, UJMC 503 9 paratype 4, specimen with smooth whorls and a trumpet-like aperture, UJMC 504 10, 11 paratype 5, specimen with smooth whorls, UJMC 505. Opercula 12, 13 paratype 5, specimen with smooth whorls, UJMC 505. Scale bar: 1 mm." figureDoi="10.3897/subtbiol.39.67799.figures2-13" httpUri="https://binary.pensoft.net/fig/574504" pageId="0" pageNumber="129">13</figureCitation>
|
||||
<figureCitation id="E5CDFCBC3C682622C3E480A60B2DDAF0" captionStart="Figures 2–13" captionStartId="F2" captionText="Figures 2 - 13. Shells of Phreatoviesca spinosa gen. nov. et sp. nov. 2, 3 holotype, specimen from both sides, UJMC 500 4, 5 paratype 1, specimen from both sides, UJMC 501 6 paratype 2, specimen with a ' corkscrew' - like morphology, UJMC 502 7, 8 paratype 3, specimen from both sides, UJMC 503 9 paratype 4, specimen with smooth whorls and a trumpet-like aperture, UJMC 504 10, 11 paratype 5, specimen with smooth whorls, UJMC 505. Opercula 12, 13 paratype 5, specimen with smooth whorls, UJMC 505. Scale bar: 1 mm." figureDoi="10.3897/subtbiol.39.67799.figures2-13" httpUri="https://binary.pensoft.net/fig/574504" pageId="0" pageNumber="129">13</figureCitation>
|
||||
), body whorl large, apertures large, ovate to subrounded, often trumpet-like (Fig.
|
||||
<figureCitation captionStart="Figures 2–13" captionStartId="F2" captionText="Figures 2 - 13. Shells of Phreatoviesca spinosa gen. nov. et sp. nov. 2, 3 holotype, specimen from both sides, UJMC 500 4, 5 paratype 1, specimen from both sides, UJMC 501 6 paratype 2, specimen with a ' corkscrew' - like morphology, UJMC 502 7, 8 paratype 3, specimen from both sides, UJMC 503 9 paratype 4, specimen with smooth whorls and a trumpet-like aperture, UJMC 504 10, 11 paratype 5, specimen with smooth whorls, UJMC 505. Opercula 12, 13 paratype 5, specimen with smooth whorls, UJMC 505. Scale bar: 1 mm." figureDoi="10.3897/subtbiol.39.67799.figures2-13" httpUri="https://binary.pensoft.net/fig/574504" pageId="0" pageNumber="129">9</figureCitation>
|
||||
<figureCitation id="0F2D41B9768ABBC16BA8C35DFE7189A8" captionStart="Figures 2–13" captionStartId="F2" captionText="Figures 2 - 13. Shells of Phreatoviesca spinosa gen. nov. et sp. nov. 2, 3 holotype, specimen from both sides, UJMC 500 4, 5 paratype 1, specimen from both sides, UJMC 501 6 paratype 2, specimen with a ' corkscrew' - like morphology, UJMC 502 7, 8 paratype 3, specimen from both sides, UJMC 503 9 paratype 4, specimen with smooth whorls and a trumpet-like aperture, UJMC 504 10, 11 paratype 5, specimen with smooth whorls, UJMC 505. Opercula 12, 13 paratype 5, specimen with smooth whorls, UJMC 505. Scale bar: 1 mm." figureDoi="10.3897/subtbiol.39.67799.figures2-13" httpUri="https://binary.pensoft.net/fig/574504" pageId="0" pageNumber="129">9</figureCitation>
|
||||
). Protoconch with coarsely honeycomb-like pits, the basal and outer lip rounded and thin, some smooth specimens with trumpet-like peristome, umbiculus deep or, in corkscrew-like specimens, almost without umbiculus; Opercula not preserved.
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="129">Shell measurements</emphasis>
|
||||
<emphasis id="1143E7764027B25BC263CA51F354068D" bold="true" italics="true" pageId="0" pageNumber="129">Shell measurements</emphasis>
|
||||
(mean
|
||||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||||
<normalizedToken id="CF164C9E21F249D5FA88DAACC7B8FB66" originalValue="±">+/-</normalizedToken>
|
||||
standard deviation in parentheses;
|
||||
<emphasis italics="true" pageId="0" pageNumber="129">n</emphasis>
|
||||
<emphasis id="16AF21BF02B65C527895C2A4FD012EE5" italics="true" pageId="0" pageNumber="129">n</emphasis>
|
||||
= 17): SH 2.08 (0.31) mm, SW 1.24 (0.17) mm, AH 0.79 (0.09) mm, AW 0.61 (0.08) mm, WN 4.93 (0.44) whorls; HBW 1.23 (0.21) mm. Paratypes from the type locality.
|
||||
</paragraph>
|
||||
<caption doi="10.3897/subtbiol.39.67799.figures2-13" httpUri="https://binary.pensoft.net/fig/574504" pageId="0" pageNumber="129" start="Figures 2–13" startId="F2">
|
||||
<paragraph pageId="0" pageNumber="129">
|
||||
<emphasis bold="true" pageId="0" pageNumber="129">Figures 2-13.</emphasis>
|
||||
<caption id="30328926DBF2B3DA41972B0DCC3270E5" doi="10.3897/subtbiol.39.67799.figures2-13" httpUri="https://binary.pensoft.net/fig/574504" pageId="0" pageNumber="129" start="Figures 2–13" startId="F2">
|
||||
<paragraph id="C3645218FD49DBAB37CA40CFD4ABD8A1" pageId="0" pageNumber="129">
|
||||
<emphasis id="785AC7E1636F06667C2C4F79013C096E" bold="true" pageId="0" pageNumber="129">Figures 2-13.</emphasis>
|
||||
Shells of
|
||||
<taxonomicName authority="Czaja & Gladstone, 2021" authorityName="Czaja & Gladstone" authorityYear="2021" class="Gastropoda" family="Cochliopidae" genus="Phreatoviesca" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phreatoviesca spinosa" order="Littorinimorpha" pageId="0" pageNumber="129" phylum="Mollusca" rank="species" species="spinosa">
|
||||
<emphasis italics="true" pageId="0" pageNumber="129">Phreatoviesca spinosa</emphasis>
|
||||
<taxonomicName id="63C3F586236EE637C747849B163E63E4" authorityName="Czaja & Gladstone" authorityYear="2021" class="Gastropoda" family="Cochliopidae" genus="Phreatoviesca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phreatoviesca spinosa" order="Littorinimorpha" pageId="0" pageNumber="129" phylum="Mollusca" rank="species" species="spinosa">
|
||||
<emphasis id="442622D3FF293842ADBB29E168847A7E" italics="true" pageId="0" pageNumber="129">Phreatoviesca spinosa</emphasis>
|
||||
</taxonomicName>
|
||||
gen. nov. et sp. nov.
|
||||
<emphasis bold="true" pageId="0" pageNumber="129">2, 3</emphasis>
|
||||
<emphasis id="24DB4E5B3C614B8A5DFEC88E977349C2" bold="true" pageId="0" pageNumber="129">2, 3</emphasis>
|
||||
holotype, specimen from both sides, UJMC 500
|
||||
<emphasis bold="true" pageId="0" pageNumber="129">4, 5</emphasis>
|
||||
<emphasis id="EE407B1A184A0A08E565E722B113CC42" bold="true" pageId="0" pageNumber="129">4, 5</emphasis>
|
||||
paratype 1, specimen from both sides, UJMC 501
|
||||
<emphasis bold="true" pageId="0" pageNumber="129">6</emphasis>
|
||||
<emphasis id="7162BCA1BDC0F0032D94B45AA6456699" bold="true" pageId="0" pageNumber="129">6</emphasis>
|
||||
paratype 2, specimen with a
|
||||
<normalizedToken originalValue="‘corkscrew’">'corkscrew'</normalizedToken>
|
||||
<normalizedToken id="4FB7DFB0FBE170F275FF6A10238E9B1A" originalValue="‘corkscrew’">'corkscrew'</normalizedToken>
|
||||
-like morphology, UJMC 502
|
||||
<emphasis bold="true" pageId="0" pageNumber="129">7, 8</emphasis>
|
||||
<emphasis id="BB9DD695E864CE456B22601D5C50FADB" bold="true" pageId="0" pageNumber="129">7, 8</emphasis>
|
||||
paratype 3, specimen from both sides, UJMC 503
|
||||
<emphasis bold="true" pageId="0" pageNumber="129">9</emphasis>
|
||||
<emphasis id="82EB484FC0937C9C0AD88657EE860654" bold="true" pageId="0" pageNumber="129">9</emphasis>
|
||||
paratype 4, specimen with smooth whorls and a trumpet-like aperture, UJMC 504
|
||||
<emphasis bold="true" pageId="0" pageNumber="129">10, 11</emphasis>
|
||||
<emphasis id="A7DAB593107E7E052ABD411E88ED4197" bold="true" pageId="0" pageNumber="129">10, 11</emphasis>
|
||||
paratype 5, specimen with smooth whorls, UJMC 505. Opercula
|
||||
<emphasis bold="true" pageId="0" pageNumber="129">12, 13</emphasis>
|
||||
<emphasis id="2DEB31881D2059D065CA7DB84A823B04" bold="true" pageId="0" pageNumber="129">12, 13</emphasis>
|
||||
paratype 5, specimen with smooth whorls, UJMC 505. Scale bar: 1 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="129" type="description">
|
||||
<paragraph pageId="0" pageNumber="129">Measurements of holotype.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="129">
|
||||
<subSubSection id="D1060536497273508A904EC730158B66" pageId="0" pageNumber="129" type="description">
|
||||
<paragraph id="BC1EBEEC695517C5C6752E347BD7D694" pageId="0" pageNumber="129">Measurements of holotype.</paragraph>
|
||||
<paragraph id="AA17D4471104F098A85421B2A9176431" pageId="0" pageNumber="129">
|
||||
WN
|
||||
<normalizedToken originalValue="5¼">51/4</normalizedToken>
|
||||
<normalizedToken id="149757D4F1DFD47F1637E0920B70152C" originalValue="5¼">51/4</normalizedToken>
|
||||
whorls; SH 2.26 mm; SW 1.41 mm; AH 0.86 mm; AW 0.67 mm, HBW 1.46 mm.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="129" type="habitat">
|
||||
<paragraph pageId="0" pageNumber="129">Habitat.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="129">
|
||||
<subSubSection id="E78CE4CC3D1B06141EAA7CDCBEBEBEE6" pageId="0" pageNumber="129" type="habitat">
|
||||
<paragraph id="4119B749667305343E9F8C4EBE9272C9" pageId="0" pageNumber="129">Habitat.</paragraph>
|
||||
<paragraph id="E7266064746489323286CD3F024CB5C6" pageId="0" pageNumber="129">
|
||||
The new species was found exclusively in one spring near
|
||||
<taxonomicName class="Gastropoda" genus="Viesca" kingdom="Animalia" lsidName="Viesca" pageId="0" pageNumber="129" phylum="Mollusca" rank="genus">Viesca</taxonomicName>
|
||||
<taxonomicName id="B5779D18073ABFF527B7757E93077F2D" class="Gastropoda" genus="Viesca" kingdom="Animalia" lsidName="Viesca" pageId="0" pageNumber="129" phylum="Mollusca" rank="genus">Viesca</taxonomicName>
|
||||
, Coahuila. The original habitat was probably the outlet of a cave, were the species likely inhabited interstitial waters.
|
||||
</paragraph>
|
||||
<caption doi="10.3897/subtbiol.39.67799.figures14-17" httpUri="https://binary.pensoft.net/fig/574505" pageId="0" pageNumber="129" start="Figures 14–17" startId="F3">
|
||||
<paragraph pageId="0" pageNumber="129">
|
||||
<emphasis bold="true" pageId="0" pageNumber="129">Figures 14-17.</emphasis>
|
||||
<caption id="1DD4919335723FF0F421E2800201452D" doi="10.3897/subtbiol.39.67799.figures14-17" httpUri="https://binary.pensoft.net/fig/574505" pageId="0" pageNumber="129" start="Figures 14–17" startId="F3">
|
||||
<paragraph id="0B533D02A10A80EF7AB9E10519E6E640" pageId="0" pageNumber="129">
|
||||
<emphasis id="C8F48768ADED7B5780131EAC2A0F9BDC" bold="true" pageId="0" pageNumber="129">Figures 14-17.</emphasis>
|
||||
SEM images of
|
||||
<taxonomicName authority="Czaja & Gladstone, 2021" authorityName="Czaja & Gladstone" authorityYear="2021" class="Gastropoda" family="Cochliopidae" genus="Phreatoviesca" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phreatoviesca spinosa" order="Littorinimorpha" pageId="0" pageNumber="129" phylum="Mollusca" rank="species" species="spinosa">
|
||||
<emphasis italics="true" pageId="0" pageNumber="129">Phreatoviesca spinosa</emphasis>
|
||||
<taxonomicName id="0B48C1E8EFADB88BFA5659356EC11C4A" authorityName="Czaja & Gladstone" authorityYear="2021" class="Gastropoda" family="Cochliopidae" genus="Phreatoviesca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phreatoviesca spinosa" order="Littorinimorpha" pageId="0" pageNumber="129" phylum="Mollusca" rank="species" species="spinosa">
|
||||
<emphasis id="5EE46F566155987835FDD4CABE10DF42" italics="true" pageId="0" pageNumber="129">Phreatoviesca spinosa</emphasis>
|
||||
</taxonomicName>
|
||||
gen. nov. et sp. nov.
|
||||
<emphasis bold="true" pageId="0" pageNumber="129">14</emphasis>
|
||||
<emphasis id="A0030934786545B53489F3A1E53B8D57" bold="true" pageId="0" pageNumber="129">14</emphasis>
|
||||
specimen with strong spines, UJMC 506
|
||||
<emphasis bold="true" pageId="0" pageNumber="129">15</emphasis>
|
||||
<emphasis id="E79BA94FA631B1DBE596B6B38B87958C" bold="true" pageId="0" pageNumber="129">15</emphasis>
|
||||
specimen with ribs, UJMC 507
|
||||
<emphasis bold="true" pageId="0" pageNumber="129">16</emphasis>
|
||||
<emphasis id="E57156FE13D2B8FF07517B6B3D332D88" bold="true" pageId="0" pageNumber="129">16</emphasis>
|
||||
specimen with smooth whorls, UJMC 508
|
||||
<emphasis bold="true" pageId="0" pageNumber="129">17</emphasis>
|
||||
<emphasis id="2C60DE1EF45AEECCEE33ED13A6E919DF" bold="true" pageId="0" pageNumber="129">17</emphasis>
|
||||
specimen with smooth whorls, UJMC 509. Scale bar: 1 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="129" type="distribution">
|
||||
<paragraph pageId="0" pageNumber="129">Distribution.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="129">
|
||||
<subSubSection id="AD83F80835454DAC4FA797FBB966B45E" pageId="0" pageNumber="129" type="distribution">
|
||||
<paragraph id="05F50BC14B528E42CADD136944817EB3" pageId="0" pageNumber="129">Distribution.</paragraph>
|
||||
<paragraph id="5BB3407F2062A0DD2CA7533D4CB47F45" pageId="0" pageNumber="129">
|
||||
A microendemic species, only in spring
|
||||
<normalizedToken originalValue="“Túnel">"Tunel</normalizedToken>
|
||||
<normalizedToken id="B2D5BB228D8A62AF020898A4CF12D2FC" originalValue="“Túnel">"Tunel</normalizedToken>
|
||||
7", near the town of
|
||||
<taxonomicName class="Gastropoda" genus="Viesca" kingdom="Animalia" lsidName="Viesca" pageId="0" pageNumber="129" phylum="Mollusca" rank="genus">Viesca</taxonomicName>
|
||||
<taxonomicName id="CBE2F138502D43D61A60290A1B95D147" class="Gastropoda" genus="Viesca" kingdom="Animalia" lsidName="Viesca" pageId="0" pageNumber="129" phylum="Mollusca" rank="genus">Viesca</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="129" type="remarks">
|
||||
<paragraph pageId="0" pageNumber="129">Remarks.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="129">
|
||||
<subSubSection id="0404D576FB397B716D6734F13503AB57" pageId="0" pageNumber="129" type="remarks">
|
||||
<paragraph id="3FBB18A76804FE2B69BE603D7957240C" pageId="0" pageNumber="129">Remarks.</paragraph>
|
||||
<paragraph id="A49ED01C39C1C0518992AF047D634C4C" pageId="0" pageNumber="129">
|
||||
The open coiled last whorl, shovel-shaped spines and a protoconch with coarsely honeycomb-like pits are the most evident characteristics which differentiated the shells of
|
||||
<taxonomicName authority="Czaja & Gladstone, 2021" authorityName="Czaja & Gladstone" authorityYear="2021" class="Gastropoda" family="Cochliopidae" genus="Phreatoviesca" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phreatoviesca" order="Littorinimorpha" pageId="0" pageNumber="129" phylum="Mollusca" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="129">Phreatoviesca</emphasis>
|
||||
<taxonomicName id="357405C5D70F2D2DE8EA11245D98A9DB" authorityName="Czaja & Gladstone" authorityYear="2021" class="Gastropoda" family="Cochliopidae" genus="Phreatoviesca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phreatoviesca" order="Littorinimorpha" pageId="0" pageNumber="129" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="3AA95F09C08B7087BDFF68267FEB1664" italics="true" pageId="0" pageNumber="129">Phreatoviesca</emphasis>
|
||||
</taxonomicName>
|
||||
gen. nov. et. sp. nov. from shells of all other described stygobiotic gastropods in North America. We considered these shell features as derived characters (apomorphy) of a new clade, most likely within the family
|
||||
<taxonomicName authorityName="Tryon" authorityYear="1866" class="Gastropoda" family="Cochliopidae" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="0" pageNumber="129" phylum="Mollusca" rank="family">Cochliopidae</taxonomicName>
|
||||
<taxonomicName id="BD0AC0CAE19E768432A108FFDED918FC" authorityName="Tryon" authorityYear="1866" class="Gastropoda" family="Cochliopidae" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="0" pageNumber="129" phylum="Mollusca" rank="family">Cochliopidae</taxonomicName>
|
||||
. The SEM imagines of the two different morphotypes (smooth and spinous) from
|
||||
<taxonomicName class="Gastropoda" genus="Viesca" kingdom="Animalia" lsidName="Viesca" pageId="0" pageNumber="129" phylum="Mollusca" rank="genus">Viesca</taxonomicName>
|
||||
<taxonomicName id="A6E68F94017F600DAA2A03A81F2B7050" class="Gastropoda" genus="Viesca" kingdom="Animalia" lsidName="Viesca" pageId="0" pageNumber="129" phylum="Mollusca" rank="genus">Viesca</taxonomicName>
|
||||
show that both have identical coarsely honeycomb-like pitted protoconchs (Figs
|
||||
<figureCitation captionStart="Figures 18–24" captionStartId="F4" captionText="Figures 18 - 24. SEM images of Phreatoviesca spinosa gen. nov. et sp. nov. 18 shell apex with protoconch, UJMC 508 19 paratype 3, shell apex with protoconch, UJMC 503 20 shell apex with protoconch, UJMC 510 21 paratype 1, apex with protoconch, UJMC 501 22 smooth specimen with body whorl shows irregular, strong marked growth lines, UJMC 508 23 paratype 1, body whorl shows irregular, strong marked growth lines, UJMC 501 24 paratype 2, body whorl shows regularly spaced shovel-shaped spines, UJMC 502." figureDoi="10.3897/subtbiol.39.67799.figures18-24" httpUri="https://binary.pensoft.net/fig/574506" pageId="0" pageNumber="129">18</figureCitation>
|
||||
<figureCitation id="0FF1D16A74F6AE4AD1A30BF21B51C4E5" captionStart="Figures 18–24" captionStartId="F4" captionText="Figures 18 - 24. SEM images of Phreatoviesca spinosa gen. nov. et sp. nov. 18 shell apex with protoconch, UJMC 508 19 paratype 3, shell apex with protoconch, UJMC 503 20 shell apex with protoconch, UJMC 510 21 paratype 1, apex with protoconch, UJMC 501 22 smooth specimen with body whorl shows irregular, strong marked growth lines, UJMC 508 23 paratype 1, body whorl shows irregular, strong marked growth lines, UJMC 501 24 paratype 2, body whorl shows regularly spaced shovel-shaped spines, UJMC 502." figureDoi="10.3897/subtbiol.39.67799.figures18-24" httpUri="https://binary.pensoft.net/fig/574506" pageId="0" pageNumber="129">18</figureCitation>
|
||||
,
|
||||
<figureCitation captionStart="Figures 18–24" captionStartId="F4" captionText="Figures 18 - 24. SEM images of Phreatoviesca spinosa gen. nov. et sp. nov. 18 shell apex with protoconch, UJMC 508 19 paratype 3, shell apex with protoconch, UJMC 503 20 shell apex with protoconch, UJMC 510 21 paratype 1, apex with protoconch, UJMC 501 22 smooth specimen with body whorl shows irregular, strong marked growth lines, UJMC 508 23 paratype 1, body whorl shows irregular, strong marked growth lines, UJMC 501 24 paratype 2, body whorl shows regularly spaced shovel-shaped spines, UJMC 502." figureDoi="10.3897/subtbiol.39.67799.figures18-24" httpUri="https://binary.pensoft.net/fig/574506" pageId="0" pageNumber="129">19</figureCitation>
|
||||
<figureCitation id="C5F91F1C40CDD07544D1272428016D3C" captionStart="Figures 18–24" captionStartId="F4" captionText="Figures 18 - 24. SEM images of Phreatoviesca spinosa gen. nov. et sp. nov. 18 shell apex with protoconch, UJMC 508 19 paratype 3, shell apex with protoconch, UJMC 503 20 shell apex with protoconch, UJMC 510 21 paratype 1, apex with protoconch, UJMC 501 22 smooth specimen with body whorl shows irregular, strong marked growth lines, UJMC 508 23 paratype 1, body whorl shows irregular, strong marked growth lines, UJMC 501 24 paratype 2, body whorl shows regularly spaced shovel-shaped spines, UJMC 502." figureDoi="10.3897/subtbiol.39.67799.figures18-24" httpUri="https://binary.pensoft.net/fig/574506" pageId="0" pageNumber="129">19</figureCitation>
|
||||
) and also the details of the shell wall microstructure with fine growth lines are similar (Figs
|
||||
<figureCitation captionStart="Figures 18–24" captionStartId="F4" captionText="Figures 18 - 24. SEM images of Phreatoviesca spinosa gen. nov. et sp. nov. 18 shell apex with protoconch, UJMC 508 19 paratype 3, shell apex with protoconch, UJMC 503 20 shell apex with protoconch, UJMC 510 21 paratype 1, apex with protoconch, UJMC 501 22 smooth specimen with body whorl shows irregular, strong marked growth lines, UJMC 508 23 paratype 1, body whorl shows irregular, strong marked growth lines, UJMC 501 24 paratype 2, body whorl shows regularly spaced shovel-shaped spines, UJMC 502." figureDoi="10.3897/subtbiol.39.67799.figures18-24" httpUri="https://binary.pensoft.net/fig/574506" pageId="0" pageNumber="129">22</figureCitation>
|
||||
<figureCitation id="25EA5ACBA844E53FBC95348A1330225A" captionStart="Figures 18–24" captionStartId="F4" captionText="Figures 18 - 24. SEM images of Phreatoviesca spinosa gen. nov. et sp. nov. 18 shell apex with protoconch, UJMC 508 19 paratype 3, shell apex with protoconch, UJMC 503 20 shell apex with protoconch, UJMC 510 21 paratype 1, apex with protoconch, UJMC 501 22 smooth specimen with body whorl shows irregular, strong marked growth lines, UJMC 508 23 paratype 1, body whorl shows irregular, strong marked growth lines, UJMC 501 24 paratype 2, body whorl shows regularly spaced shovel-shaped spines, UJMC 502." figureDoi="10.3897/subtbiol.39.67799.figures18-24" httpUri="https://binary.pensoft.net/fig/574506" pageId="0" pageNumber="129">22</figureCitation>
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||||
,
|
||||
<figureCitation captionStart="Figures 18–24" captionStartId="F4" captionText="Figures 18 - 24. SEM images of Phreatoviesca spinosa gen. nov. et sp. nov. 18 shell apex with protoconch, UJMC 508 19 paratype 3, shell apex with protoconch, UJMC 503 20 shell apex with protoconch, UJMC 510 21 paratype 1, apex with protoconch, UJMC 501 22 smooth specimen with body whorl shows irregular, strong marked growth lines, UJMC 508 23 paratype 1, body whorl shows irregular, strong marked growth lines, UJMC 501 24 paratype 2, body whorl shows regularly spaced shovel-shaped spines, UJMC 502." figureDoi="10.3897/subtbiol.39.67799.figures18-24" httpUri="https://binary.pensoft.net/fig/574506" pageId="0" pageNumber="129">23</figureCitation>
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<figureCitation id="6DCEE1B35B6149D3EBA434E8DEDCB6F0" captionStart="Figures 18–24" captionStartId="F4" captionText="Figures 18 - 24. SEM images of Phreatoviesca spinosa gen. nov. et sp. nov. 18 shell apex with protoconch, UJMC 508 19 paratype 3, shell apex with protoconch, UJMC 503 20 shell apex with protoconch, UJMC 510 21 paratype 1, apex with protoconch, UJMC 501 22 smooth specimen with body whorl shows irregular, strong marked growth lines, UJMC 508 23 paratype 1, body whorl shows irregular, strong marked growth lines, UJMC 501 24 paratype 2, body whorl shows regularly spaced shovel-shaped spines, UJMC 502." figureDoi="10.3897/subtbiol.39.67799.figures18-24" httpUri="https://binary.pensoft.net/fig/574506" pageId="0" pageNumber="129">23</figureCitation>
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). Therefore, we consider these two morphotypes as belonging to the same species. There is no significant difference in shells measurements between smooth and spiny morphotypes and therefore sexual dimorphism is unlikely. Moreover, most of the shells have strong spines and only less than 5% of the morphotypes collected are smooth. Two morphotypes (one smooth and other with lamelliform costae) not associated with sexual dimorphism, were reported also from shells of the subterranean genus
|
||||
<taxonomicName authorityName="D. W. Taylor" authorityYear="1966" class="Gastropoda" family="Cochliopidae" genus="Paludiscala" kingdom="Animalia" lsidName="Paludiscala" order="Littorinimorpha" pageId="0" pageNumber="129" phylum="Mollusca" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="129">Paludiscala</emphasis>
|
||||
<taxonomicName id="773AB7DA959E4F8624A575D4298FE396" authorityName="D. W. Taylor" authorityYear="1966" class="Gastropoda" family="Cochliopidae" genus="Paludiscala" kingdom="Animalia" lsidName="Paludiscala" order="Littorinimorpha" pageId="0" pageNumber="129" phylum="Mollusca" rank="genus">
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<emphasis id="86EB1A439DAF6553D00606E00DA6DF75" italics="true" pageId="0" pageNumber="129">Paludiscala</emphasis>
|
||||
</taxonomicName>
|
||||
Taylor, 1966, described from the neighboring Cuatro
|
||||
<normalizedToken originalValue="Ciénegas">Cienegas</normalizedToken>
|
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<normalizedToken id="AEC96582F650C575E0C2581EE8969F46" originalValue="Ciénegas">Cienegas</normalizedToken>
|
||||
Basin (Hershler, 1985). Interestingly, our material is conchologically similar to members of the stygobiotic and stygophilic genus
|
||||
<taxonomicName authorityName="Ancey" authorityYear="1888" class="Gastropoda" family="Cochliopidae" genus="Pyrgophorus" kingdom="Animalia" lsidName="Pyrgophorus" order="Littorinimorpha" pageId="0" pageNumber="129" phylum="Mollusca" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="129">Pyrgophorus</emphasis>
|
||||
<taxonomicName id="B9D8E0A5CF6C186A41A9E2EA63D9D55E" authorityName="Ancey" authorityYear="1888" class="Gastropoda" family="Cochliopidae" genus="Pyrgophorus" kingdom="Animalia" lsidName="Pyrgophorus" order="Littorinimorpha" pageId="0" pageNumber="129" phylum="Mollusca" rank="genus">
|
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<emphasis id="41A3F81469EF9AA575BF7ED3EEFE34BF" italics="true" pageId="0" pageNumber="129">Pyrgophorus</emphasis>
|
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</taxonomicName>
|
||||
Ancey, 1888 in Mexico, which show similar shovel-shaped spines (
|
||||
<bibRefCitation DOI="https://doi.org/10.3897/subtbiol.29.32779" author="Grego, J" journalOrPublisher="Subterranean Biology" pageId="0" pageNumber="129" pagination="79 - 88" refId="B17" refString="Grego, J, Angyal, D, Beltran, LAL, 2019. First record of subterranean freshwater gastropods (Mollusca, Gastropoda, Cochliopidae) from the cenotes of Yucatan state. Subterranean Biology 29: 79 - 88, DOI: https://doi.org/10.3897/subtbiol.29.32779" title="First record of subterranean freshwater gastropods (Mollusca, Gastropoda, Cochliopidae) from the cenotes of Yucatan state." url="https://doi.org/10.3897/subtbiol.29.32779" volume="29" year="2019">Grego et al. 2019</bibRefCitation>
|
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<bibRefCitation id="9A880DD0ED3E97CB4099ADEA2F160DCD" DOI="https://doi.org/10.3897/subtbiol.29.32779" author="Grego, J" journalOrPublisher="Subterranean Biology" pageId="0" pageNumber="129" pagination="79 - 88" refId="B17" refString="Grego, J, Angyal, D, Beltran, LAL, 2019. First record of subterranean freshwater gastropods (Mollusca, Gastropoda, Cochliopidae) from the cenotes of Yucatan state. Subterranean Biology 29: 79 - 88, DOI: https://doi.org/10.3897/subtbiol.29.32779" title="First record of subterranean freshwater gastropods (Mollusca, Gastropoda, Cochliopidae) from the cenotes of Yucatan state." url="https://doi.org/10.3897/subtbiol.29.32779" volume="29" year="2019">Grego et al. 2019</bibRefCitation>
|
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). This resemblance is surely an evolutionary convergence and result from living in subterranean habitats.
|
||||
</paragraph>
|
||||
<caption doi="10.3897/subtbiol.39.67799.figures18-24" httpUri="https://binary.pensoft.net/fig/574506" pageId="0" pageNumber="129" start="Figures 18–24" startId="F4">
|
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<paragraph pageId="0" pageNumber="129">
|
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<emphasis bold="true" pageId="0" pageNumber="129">Figures 18-24.</emphasis>
|
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<caption id="F35B1AE2E19A55B576F5B45F8FC224F7" doi="10.3897/subtbiol.39.67799.figures18-24" httpUri="https://binary.pensoft.net/fig/574506" pageId="0" pageNumber="129" start="Figures 18–24" startId="F4">
|
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<paragraph id="78C0B54BA9BAB0F621F72EFD5B593E8F" pageId="0" pageNumber="129">
|
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<emphasis id="6B145D6ECA4705205C1C0CE3B3A82E51" bold="true" pageId="0" pageNumber="129">Figures 18-24.</emphasis>
|
||||
SEM images of
|
||||
<taxonomicName authority="Czaja & Gladstone, 2021" authorityName="Czaja & Gladstone" authorityYear="2021" class="Gastropoda" family="Cochliopidae" genus="Phreatoviesca" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phreatoviesca spinosa" order="Littorinimorpha" pageId="0" pageNumber="129" phylum="Mollusca" rank="species" species="spinosa">
|
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<emphasis italics="true" pageId="0" pageNumber="129">Phreatoviesca spinosa</emphasis>
|
||||
<taxonomicName id="8ABA3D09845C9A05A71399FAC2F15C0D" authorityName="Czaja & Gladstone" authorityYear="2021" class="Gastropoda" family="Cochliopidae" genus="Phreatoviesca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phreatoviesca spinosa" order="Littorinimorpha" pageId="0" pageNumber="129" phylum="Mollusca" rank="species" species="spinosa">
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<emphasis id="7DC3697C2A0DB63E42CA8D8B540DC53C" italics="true" pageId="0" pageNumber="129">Phreatoviesca spinosa</emphasis>
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</taxonomicName>
|
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gen. nov. et sp. nov.
|
||||
<emphasis bold="true" pageId="0" pageNumber="129">18</emphasis>
|
||||
<emphasis id="7CCBD8338E78DFBEA58EFDF660F99F11" bold="true" pageId="0" pageNumber="129">18</emphasis>
|
||||
shell apex with protoconch, UJMC 508
|
||||
<emphasis bold="true" pageId="0" pageNumber="129">19</emphasis>
|
||||
<emphasis id="ACC6054FE311B907E39776501893695F" bold="true" pageId="0" pageNumber="129">19</emphasis>
|
||||
paratype 3, shell apex with protoconch, UJMC 503
|
||||
<emphasis bold="true" pageId="0" pageNumber="129">20</emphasis>
|
||||
<emphasis id="204EF7DAAF1AEACC1C80DE247228CC1C" bold="true" pageId="0" pageNumber="129">20</emphasis>
|
||||
shell apex with protoconch, UJMC 510
|
||||
<emphasis bold="true" pageId="0" pageNumber="129">21</emphasis>
|
||||
<emphasis id="22037AEB65A19F90C659FBB0196411CC" bold="true" pageId="0" pageNumber="129">21</emphasis>
|
||||
paratype 1, apex with protoconch, UJMC 501
|
||||
<emphasis bold="true" pageId="0" pageNumber="129">22</emphasis>
|
||||
<emphasis id="508BA396D59DB7C3B8ACDAB0EB78A5FA" bold="true" pageId="0" pageNumber="129">22</emphasis>
|
||||
smooth specimen with body whorl shows irregular, strong marked growth lines, UJMC 508
|
||||
<emphasis bold="true" pageId="0" pageNumber="129">23</emphasis>
|
||||
<emphasis id="D7F2991D0081962ACDA3D85287361445" bold="true" pageId="0" pageNumber="129">23</emphasis>
|
||||
paratype 1, body whorl shows irregular, strong marked growth lines, UJMC 501
|
||||
<emphasis bold="true" pageId="0" pageNumber="129">24</emphasis>
|
||||
<emphasis id="1515EDB5AC86D7FBA5357EA4EB6D2E01" bold="true" pageId="0" pageNumber="129">24</emphasis>
|
||||
paratype 2, body whorl shows regularly spaced shovel-shaped spines, UJMC 502.
|
||||
</paragraph>
|
||||
</caption>
|
||||
|
|
|
|||
|
|
@ -1,210 +1,210 @@
|
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|
||||
<subSubSection id="861B1EB040BBD1D29BB399C9A489EBC5" pageId="0" pageNumber="61" type="nomenclature">
|
||||
<paragraph id="4EA693A9BE074BB2F38AFEBD1B337C56" pageId="0" pageNumber="61">
|
||||
<taxonomicName id="AD0AF95B5EC689D2C380903A6D6662D1" ID-CoL="8MHQC" LSID="8CD5F85F-6B59-5FA0-8552-C1864653A65E" authority="Habu & Baba, 1960" authorityName="Habu & Baba" authorityYear="1960" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Nialoe) asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus" subGenus="Nialoe">Pterostichus (Nialoe) asahinus Habu & Baba, 1960</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="61" type="description">
|
||||
<paragraph pageId="0" pageNumber="61">
|
||||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Pterostichus asahinus male from Miomote, on the right bank of the Miomotegawa River A habitus dorsal view B-E Endophallus left lateral (B) ventral (C) right lateral (D) and dorsal (E) views F-H right paramere left lateral (F) apical (G) and dorsal (H) views. go, gonopore; lal, left apical lobe; lpl, left preapical lobe; rpl, right preapical lobe." figureDoi="10.3897/subtbiol.43.80969.figure2" httpUri="https://binary.pensoft.net/fig/690341" pageId="0" pageNumber="61">Fig. 2</figureCitation>
|
||||
<subSubSection id="8957434F722AAEB2C6A61C264493DE76" pageId="0" pageNumber="61" type="description">
|
||||
<paragraph id="C30A92733759E17FAB5BA1B928C747A7" pageId="0" pageNumber="61">
|
||||
<figureCitation id="393B0757F055E7F6957DD689FE23CBA1" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Pterostichus asahinus male from Miomote, on the right bank of the Miomotegawa River A habitus dorsal view B-E Endophallus left lateral (B) ventral (C) right lateral (D) and dorsal (E) views F-H right paramere left lateral (F) apical (G) and dorsal (H) views. go, gonopore; lal, left apical lobe; lpl, left preapical lobe; rpl, right preapical lobe." figureDoi="10.3897/subtbiol.43.80969.figure2" httpUri="https://binary.pensoft.net/fig/690341" pageId="0" pageNumber="61">Fig. 2</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="61" type="reference_group">
|
||||
<paragraph pageId="0" pageNumber="61">
|
||||
<taxonomicName class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Paralianoe) macrogenys subsp. asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="subSpecies" species="macrogenys" subGenus="Paralianoe" subSpecies="asahinus">Pterostichus (Paralianoe) macrogenys asahinus</taxonomicName>
|
||||
<subSubSection id="47ABBA715AF85FDA7FA0FDE1DE373F85" pageId="0" pageNumber="61" type="reference_group">
|
||||
<paragraph id="0FEF55F9230AE96CC971BED422D1EB63" pageId="0" pageNumber="61">
|
||||
<taxonomicName id="95AAF06B8A7248265C60325104AB4362" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Paralianoe) macrogenys subsp. asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="subSpecies" species="macrogenys" subGenus="Paralianoe" subSpecies="asahinus">Pterostichus (Paralianoe) macrogenys asahinus</taxonomicName>
|
||||
:
|
||||
<bibRefCitation author="Habu, A" journalOrPublisher="Entomological Review Japan" pageId="0" pageNumber="61" refId="B5" refString="Habu, A, Baba, K, 1960. A new subspecies of Pterostichus macrogenys Bates from Niigata Prefecture, Japan. Akitu 9: 62." title="A new subspecies of Pterostichus macrogenys Bates from Niigata Prefecture, Japan. Akitu 9: 62." year="1960">Habu and Baba (1960)</bibRefCitation>
|
||||
<bibRefCitation id="7EC78C595846E55EDEA53305F77E44FC" author="Habu, A" journalOrPublisher="Entomological Review Japan" pageId="0" pageNumber="61" refId="B5" refString="Habu, A, Baba, K, 1960. A new subspecies of Pterostichus macrogenys Bates from Niigata Prefecture, Japan. Akitu 9: 62." title="A new subspecies of Pterostichus macrogenys Bates from Niigata Prefecture, Japan. Akitu 9: 62." year="1960">Habu and Baba (1960)</bibRefCitation>
|
||||
: 62 (original description), holotype ♀: "Mt. Dorokujin, Mts. Asahi, Niigata Pref." [Miomote, Mt.
|
||||
<normalizedToken originalValue="Dôrokujinpô">Dorokujinpo</normalizedToken>
|
||||
<normalizedToken id="C2A7EEC58DD77660FF701C54444D5D91" originalValue="Dôrokujinpô">Dorokujinpo</normalizedToken>
|
||||
, Murakami-shi, Niigata Prefecture, Japan];
|
||||
<bibRefCitation author="Habu, A" journalOrPublisher="Transactions of Essa Entomological Society" pageId="0" pageNumber="61" pagination="1 - 58" refId="B6" refString="Habu, A, Baba, K, 1972. List of Carabidae (excluding Carabini) from Niigata Prefecture, Japan. Transactions of Essa Entomological Society 41: 1 - 58" title="List of Carabidae (excluding Carabini) from Niigata Prefecture, Japan." volume="41" year="1972">Habu and Baba (1972)</bibRefCitation>
|
||||
<bibRefCitation id="C705C02C2E02BF7D3472F26AD822877E" author="Habu, A" journalOrPublisher="Transactions of Essa Entomological Society" pageId="0" pageNumber="61" pagination="1 - 58" refId="B6" refString="Habu, A, Baba, K, 1972. List of Carabidae (excluding Carabini) from Niigata Prefecture, Japan. Transactions of Essa Entomological Society 41: 1 - 58" title="List of Carabidae (excluding Carabini) from Niigata Prefecture, Japan." volume="41" year="1972">Habu and Baba (1972)</bibRefCitation>
|
||||
: 19.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="61">
|
||||
<taxonomicName authorityName="Habu & Baba" authorityYear="1960" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Paralianoe) asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus" subGenus="Paralianoe">Pterostichus (Paralianoe) asahinus</taxonomicName>
|
||||
<paragraph id="1F059D92ED1D788F750A8D21F62F6123" pageId="0" pageNumber="61">
|
||||
<taxonomicName id="2D8F9DCC603CABF1B8F01EA5950E3094" authorityName="Habu & Baba" authorityYear="1960" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Paralianoe) asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus" subGenus="Paralianoe">Pterostichus (Paralianoe) asahinus</taxonomicName>
|
||||
:
|
||||
<bibRefCitation author="Habu, A" journalOrPublisher="Entomological Review Japan" pageId="0" pageNumber="61" pagination="13 - 17" refId="B4" refString="Habu, A, 1977. On two Pterostichus species found in caves in Niigata Prefecture, Japan (Coleoptera, Carabidae). Entomological Review Japan 30: 13 - 17" title="On two Pterostichus species found in caves in Niigata Prefecture, Japan (Coleoptera, Carabidae)." volume="30" year="1977">Habu (1977)</bibRefCitation>
|
||||
<bibRefCitation id="9F8F5EFBF96CAF2535C3C8E8D8900D84" author="Habu, A" journalOrPublisher="Entomological Review Japan" pageId="0" pageNumber="61" pagination="13 - 17" refId="B4" refString="Habu, A, 1977. On two Pterostichus species found in caves in Niigata Prefecture, Japan (Coleoptera, Carabidae). Entomological Review Japan 30: 13 - 17" title="On two Pterostichus species found in caves in Niigata Prefecture, Japan (Coleoptera, Carabidae)." volume="30" year="1977">Habu (1977)</bibRefCitation>
|
||||
: 14 (part).
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="61">
|
||||
<taxonomicName authorityName="Sasakawa & Itô" authorityYear="2022" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus macrogenys" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="macrogenys">Pterostichus macrogenys</taxonomicName>
|
||||
<paragraph id="BB4EF9BE8F6D30B281F8033CB2CB180D" pageId="0" pageNumber="61">
|
||||
<taxonomicName id="408FC0438DD280BF43B3DB7C2252BE2C" authorityName="Sasakawa & Itô" authorityYear="2022" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus macrogenys" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="macrogenys">Pterostichus macrogenys</taxonomicName>
|
||||
:
|
||||
<bibRefCitation author="Tanaka, K" editor="Ueno, S" journalOrPublisher="Vol. II. Hoikusha, Osaka" pageId="0" pageNumber="61" pagination="105 - 138" refId="B21" refString="Tanaka, K, 1985. Carabidae (Pterostichinae, Zabrinae). In: Ueno, S, Kurosawa, Y, Sato, M, Eds., The Coleoptera of Japan in Color. Vol. II. Hoikusha, Osaka: 105 - 138" title="Carabidae (Pterostichinae, Zabrinae)." volumeTitle="The Coleoptera of Japan in Color." year="1985">Tanaka (1985)</bibRefCitation>
|
||||
<bibRefCitation id="56F391B676CEEF807776ED94C0AE077B" author="Tanaka, K" editor="Ueno, S" journalOrPublisher="Vol. II. Hoikusha, Osaka" pageId="0" pageNumber="61" pagination="105 - 138" refId="B21" refString="Tanaka, K, 1985. Carabidae (Pterostichinae, Zabrinae). In: Ueno, S, Kurosawa, Y, Sato, M, Eds., The Coleoptera of Japan in Color. Vol. II. Hoikusha, Osaka: 105 - 138" title="Carabidae (Pterostichinae, Zabrinae)." volumeTitle="The Coleoptera of Japan in Color." year="1985">Tanaka (1985)</bibRefCitation>
|
||||
: 114 (part?).
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="61">
|
||||
<taxonomicName authorityName="Habu & Baba" authorityYear="1960" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Nialoe) asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus" subGenus="Nialoe">Pterostichus (Nialoe) asahinus</taxonomicName>
|
||||
<paragraph id="2EBEA10A4406AD46F7CC8500716DA05F" pageId="0" pageNumber="61">
|
||||
<taxonomicName id="680A06BFA7C707493255C9A76BF73096" authorityName="Habu & Baba" authorityYear="1960" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Nialoe) asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus" subGenus="Nialoe">Pterostichus (Nialoe) asahinus</taxonomicName>
|
||||
:
|
||||
<bibRefCitation author="Bousquet, Y" editor="Loebl, I" journalOrPublisher="1, Archostemata-Myxophaga-Adephaga, Revised and Updated Edition. Brill, Leiden" pageId="0" pageNumber="61" pagination="675 - 755" refId="B2" refString="Bousquet, Y, 2017. Tribe Pterostichini Bonelli, 1810. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera, Vol. 1, Archostemata-Myxophaga-Adephaga, Revised and Updated Edition. Brill, Leiden: 675 - 755" title="Tribe Pterostichini Bonelli, 1810." volumeTitle="Catalogue of Palaearctic Coleoptera, Vol." year="2017">Bousquet (2017)</bibRefCitation>
|
||||
<bibRefCitation id="0FDC3CA1797737440E0AD8080A6973F0" author="Bousquet, Y" editor="Loebl, I" journalOrPublisher="1, Archostemata-Myxophaga-Adephaga, Revised and Updated Edition. Brill, Leiden" pageId="0" pageNumber="61" pagination="675 - 755" refId="B2" refString="Bousquet, Y, 2017. Tribe Pterostichini Bonelli, 1810. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera, Vol. 1, Archostemata-Myxophaga-Adephaga, Revised and Updated Edition. Brill, Leiden: 675 - 755" title="Tribe Pterostichini Bonelli, 1810." volumeTitle="Catalogue of Palaearctic Coleoptera, Vol." year="2017">Bousquet (2017)</bibRefCitation>
|
||||
: 724.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="61">
|
||||
<taxonomicName authorityName="Sasakawa" authorityYear="2005" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Nialoe) falcispinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="falcispinus" subGenus="Nialoe">Pterostichus (Nialoe) falcispinus</taxonomicName>
|
||||
<paragraph id="FA4CC34011D4666FE20EEA8E83A259F1" pageId="0" pageNumber="61">
|
||||
<taxonomicName id="3DB29D4FBB63206EF2841C6F0E10960C" authorityName="Sasakawa" authorityYear="2005" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Nialoe) falcispinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="falcispinus" subGenus="Nialoe">Pterostichus (Nialoe) falcispinus</taxonomicName>
|
||||
:
|
||||
<bibRefCitation author="Sasakawa, K" journalOrPublisher="Biogeography" pageId="0" pageNumber="61" pagination="69 - 78" refId="B12" refString="Sasakawa, K, 2005. Pterostichus macrogenys Bates, 1883 (Coleoptera, Carabidae) and its allied species of northern Japan. Biogeography 7: 69 - 78" title="Pterostichus macrogenys Bates, 1883 (Coleoptera, Carabidae) and its allied species of northern Japan." volume="7" year="2005">Sasakawa (2005)</bibRefCitation>
|
||||
<bibRefCitation id="B092E54EEB450F612D3570608EF2E9FC" author="Sasakawa, K" journalOrPublisher="Biogeography" pageId="0" pageNumber="61" pagination="69 - 78" refId="B12" refString="Sasakawa, K, 2005. Pterostichus macrogenys Bates, 1883 (Coleoptera, Carabidae) and its allied species of northern Japan. Biogeography 7: 69 - 78" title="Pterostichus macrogenys Bates, 1883 (Coleoptera, Carabidae) and its allied species of northern Japan." volume="7" year="2005">Sasakawa (2005)</bibRefCitation>
|
||||
: 75 (original description), holotype ♂: "Cave Ishikiri, Nakajo-Machi, N-Echigo" [Ishikiri Cave, Mt. Ishikiriyama, Haguro, Tainai-shi, Niigata Prefecture, Japan];
|
||||
<bibRefCitation author="Bousquet, Y" editor="Loebl, I" journalOrPublisher="1, Archostemata-Myxophaga-Adephaga, Revised and Updated Edition. Brill, Leiden" pageId="0" pageNumber="61" pagination="675 - 755" refId="B2" refString="Bousquet, Y, 2017. Tribe Pterostichini Bonelli, 1810. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera, Vol. 1, Archostemata-Myxophaga-Adephaga, Revised and Updated Edition. Brill, Leiden: 675 - 755" title="Tribe Pterostichini Bonelli, 1810." volumeTitle="Catalogue of Palaearctic Coleoptera, Vol." year="2017">Bousquet (2017)</bibRefCitation>
|
||||
<bibRefCitation id="3CB245FA7C1FE7E401BB593BF9ADF34D" author="Bousquet, Y" editor="Loebl, I" journalOrPublisher="1, Archostemata-Myxophaga-Adephaga, Revised and Updated Edition. Brill, Leiden" pageId="0" pageNumber="61" pagination="675 - 755" refId="B2" refString="Bousquet, Y, 2017. Tribe Pterostichini Bonelli, 1810. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera, Vol. 1, Archostemata-Myxophaga-Adephaga, Revised and Updated Edition. Brill, Leiden: 675 - 755" title="Tribe Pterostichini Bonelli, 1810." volumeTitle="Catalogue of Palaearctic Coleoptera, Vol." year="2017">Bousquet (2017)</bibRefCitation>
|
||||
: 724;
|
||||
<bibRefCitation DOI="https://doi.org/10.3390/insects11110803" author="Sasakawa, K" journalOrPublisher="Applied Entomology and Zoology" pageId="0" pageNumber="61" refId="B17" refString="Sasakawa, K, Mitsuduka, Y, Ito, H, 2020. Unexpected species identities and interspecific relationships in a subterranean beetle lineage, the Pterostichus macrogenys species group (Coleoptera, Carabidae), revealed by fine-scale field sampling and detailed morphological comparisons. Insects 11: 803. https://doi.org/10.3390/insects11110803" title="Unexpected species identities and interspecific relationships in a subterranean beetle lineage, the Pterostichus macrogenys species group (Coleoptera, Carabidae), revealed by fine-scale field sampling and detailed morphological comparisons. Insects 11: 803." url="https://doi.org/10.3390/insects11110803" year="2020">Sasakawa et al. (2020)</bibRefCitation>
|
||||
<bibRefCitation id="D3FE0FAC5D74A198E0132972CDDA1BDA" DOI="https://doi.org/10.3390/insects11110803" author="Sasakawa, K" journalOrPublisher="Applied Entomology and Zoology" pageId="0" pageNumber="61" refId="B17" refString="Sasakawa, K, Mitsuduka, Y, Ito, H, 2020. Unexpected species identities and interspecific relationships in a subterranean beetle lineage, the Pterostichus macrogenys species group (Coleoptera, Carabidae), revealed by fine-scale field sampling and detailed morphological comparisons. Insects 11: 803. https://doi.org/10.3390/insects11110803" title="Unexpected species identities and interspecific relationships in a subterranean beetle lineage, the Pterostichus macrogenys species group (Coleoptera, Carabidae), revealed by fine-scale field sampling and detailed morphological comparisons. Insects 11: 803." url="https://doi.org/10.3390/insects11110803" year="2020">Sasakawa et al. (2020)</bibRefCitation>
|
||||
: 7. Syn. nov.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="61" type="specimen examined">
|
||||
<paragraph pageId="0" pageNumber="61">Specimen examined.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="61">
|
||||
<subSubSection id="72096F1ED0BFA2B0D5B7AA786549A1FB" pageId="0" pageNumber="61" type="specimen examined">
|
||||
<paragraph id="C4BB171E575DD5696609B2CFD4583485" pageId="0" pageNumber="61">Specimen examined.</paragraph>
|
||||
<paragraph id="4AEBD80FC77B29C48B3D90869D728EEF" pageId="0" pageNumber="61">
|
||||
1♂, Miomote, alt. 276 m, on the right bank of Miomotegawa River, Murakami-shi, Niigata Prefecture, Japan (
|
||||
<geoCoordinate degrees="38.273211" direction="north" orientation="latitude" precision="1" value="38.273212">38.273211°N</geoCoordinate>
|
||||
<geoCoordinate id="BCE1BEB97536BA98466CF9B540BE310C" degrees="38.273211" direction="north" orientation="latitude" precision="1" value="38.273212">38.273211°N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="139.779922" direction="east" orientation="longitude" precision="1" value="139.77992">139.779922°E</geoCoordinate>
|
||||
<geoCoordinate id="CAEF9F21A139E7E54FE23D1AFFE1E075" degrees="139.779922" direction="east" orientation="longitude" precision="1" value="139.77992">139.779922°E</geoCoordinate>
|
||||
), 12.vi.-17.vii.2021,
|
||||
<normalizedToken originalValue="Hirotarô">Hirotaro</normalizedToken>
|
||||
<normalizedToken originalValue="Itô">Ito</normalizedToken>
|
||||
<normalizedToken id="9C56742F357865B79F9E586B499B7B66" originalValue="Hirotarô">Hirotaro</normalizedToken>
|
||||
<normalizedToken id="788573952A565B45C8A30F5FFB20CF21" originalValue="Itô">Ito</normalizedToken>
|
||||
leg., in the collection of HI.
|
||||
</paragraph>
|
||||
<caption doi="10.3897/subtbiol.43.80969.figure2" httpUri="https://binary.pensoft.net/fig/690341" pageId="0" pageNumber="61" start="Figure 2" startId="F2">
|
||||
<paragraph pageId="0" pageNumber="61">
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">Figure 2.</emphasis>
|
||||
<taxonomicName authorityName="Habu & Baba" authorityYear="1960" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="61">Pterostichus asahinus</emphasis>
|
||||
<caption id="C530FDC105DF6C651F5F9D95AAE28BD4" doi="10.3897/subtbiol.43.80969.figure2" httpUri="https://binary.pensoft.net/fig/690341" pageId="0" pageNumber="61" start="Figure 2" startId="F2">
|
||||
<paragraph id="63180431A8A196F322249253BF9C0E2F" pageId="0" pageNumber="61">
|
||||
<emphasis id="77CC9E034167B20AFF189C63FE09F64D" bold="true" pageId="0" pageNumber="61">Figure 2.</emphasis>
|
||||
<taxonomicName id="5E3F006BFC7C346B7841E94012D485D9" authorityName="Habu & Baba" authorityYear="1960" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus">
|
||||
<emphasis id="CDCDF320E6B4F04B80CA42BDD6A4F8F5" italics="true" pageId="0" pageNumber="61">Pterostichus asahinus</emphasis>
|
||||
</taxonomicName>
|
||||
male from Miomote, on the right bank of the Miomotegawa River
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">A</emphasis>
|
||||
<emphasis id="5A52D1363C0A9DE67567F41DF343BD4D" bold="true" pageId="0" pageNumber="61">A</emphasis>
|
||||
habitus dorsal view
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">B-E</emphasis>
|
||||
<emphasis id="5E31E372E1D28CB3DA3B334CBABA9446" bold="true" pageId="0" pageNumber="61">B-E</emphasis>
|
||||
Endophallus left lateral (
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">B</emphasis>
|
||||
<emphasis id="270EAFA9DE01BB2C46B12392296C65D7" bold="true" pageId="0" pageNumber="61">B</emphasis>
|
||||
) ventral (
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">C</emphasis>
|
||||
<emphasis id="0C9D21BCD388C6D44A8B0588BB8F199A" bold="true" pageId="0" pageNumber="61">C</emphasis>
|
||||
) right lateral (
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">D</emphasis>
|
||||
<emphasis id="CFC343AA6851C5F1FB3B34BCDA11917E" bold="true" pageId="0" pageNumber="61">D</emphasis>
|
||||
) and dorsal (
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">E</emphasis>
|
||||
<emphasis id="75ADEDA4467C6A6EE63BA27175E2FC5E" bold="true" pageId="0" pageNumber="61">E</emphasis>
|
||||
) views
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">F-H</emphasis>
|
||||
<emphasis id="8F6FE4635CC00D7544DDBB327FB09FEB" bold="true" pageId="0" pageNumber="61">F-H</emphasis>
|
||||
right paramere left lateral (
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">F</emphasis>
|
||||
<emphasis id="F039D909F18071805910AFDB3E5EC1C1" bold="true" pageId="0" pageNumber="61">F</emphasis>
|
||||
) apical (
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">G</emphasis>
|
||||
<emphasis id="594EFA5D434E3ED57403DAE2E17EB1C6" bold="true" pageId="0" pageNumber="61">G</emphasis>
|
||||
) and dorsal (
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">H</emphasis>
|
||||
<emphasis id="90384B4940F0FCCAA8C382EF30915A42" bold="true" pageId="0" pageNumber="61">H</emphasis>
|
||||
) views.
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">go</emphasis>
|
||||
<emphasis id="A8FFBEC35A379035961E8C21E2F42449" bold="true" pageId="0" pageNumber="61">go</emphasis>
|
||||
, gonopore;
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">lal</emphasis>
|
||||
<emphasis id="0B4A2DA98A2215E40478C17FFF937155" bold="true" pageId="0" pageNumber="61">lal</emphasis>
|
||||
, left apical lobe;
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">lpl</emphasis>
|
||||
<emphasis id="DEB7972B8224460D4D62407D9EE06541" bold="true" pageId="0" pageNumber="61">lpl</emphasis>
|
||||
, left preapical lobe;
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">rpl</emphasis>
|
||||
<emphasis id="17D8C0C24D946BFEA40838978CAD3DF5" bold="true" pageId="0" pageNumber="61">rpl</emphasis>
|
||||
, right preapical lobe.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="61" type="notes">
|
||||
<paragraph pageId="0" pageNumber="61">Notes.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="61">
|
||||
<bibRefCitation DOI="https://doi.org/10.3390/insects11110803" author="Sasakawa, K" journalOrPublisher="Applied Entomology and Zoology" pageId="0" pageNumber="61" refId="B17" refString="Sasakawa, K, Mitsuduka, Y, Ito, H, 2020. Unexpected species identities and interspecific relationships in a subterranean beetle lineage, the Pterostichus macrogenys species group (Coleoptera, Carabidae), revealed by fine-scale field sampling and detailed morphological comparisons. Insects 11: 803. https://doi.org/10.3390/insects11110803" title="Unexpected species identities and interspecific relationships in a subterranean beetle lineage, the Pterostichus macrogenys species group (Coleoptera, Carabidae), revealed by fine-scale field sampling and detailed morphological comparisons. Insects 11: 803." url="https://doi.org/10.3390/insects11110803" year="2020">Sasakawa et al. (2020)</bibRefCitation>
|
||||
<subSubSection id="548F36E7DB8E63985EE1A15792D6006F" pageId="0" pageNumber="61" type="notes">
|
||||
<paragraph id="13CBE3F09CD0FC405424C15CA8E9D513" pageId="0" pageNumber="61">Notes.</paragraph>
|
||||
<paragraph id="99DAEED5CA60976833F46137E027074B" pageId="0" pageNumber="61">
|
||||
<bibRefCitation id="FFCE57C2F8FED6D18C494D21A6A13EF5" DOI="https://doi.org/10.3390/insects11110803" author="Sasakawa, K" journalOrPublisher="Applied Entomology and Zoology" pageId="0" pageNumber="61" refId="B17" refString="Sasakawa, K, Mitsuduka, Y, Ito, H, 2020. Unexpected species identities and interspecific relationships in a subterranean beetle lineage, the Pterostichus macrogenys species group (Coleoptera, Carabidae), revealed by fine-scale field sampling and detailed morphological comparisons. Insects 11: 803. https://doi.org/10.3390/insects11110803" title="Unexpected species identities and interspecific relationships in a subterranean beetle lineage, the Pterostichus macrogenys species group (Coleoptera, Carabidae), revealed by fine-scale field sampling and detailed morphological comparisons. Insects 11: 803." url="https://doi.org/10.3390/insects11110803" year="2020">Sasakawa et al. (2020)</bibRefCitation>
|
||||
suggested that
|
||||
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="61">P. asahinus</emphasis>
|
||||
<taxonomicName id="1C2FC4793F2A71812B8971CE9FF7D068" class="Insecta" kingdom="Animalia" lsidName="P. asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus">
|
||||
<emphasis id="81A1D4B254BBB521CBFB58F1E73F342E" italics="true" pageId="0" pageNumber="61">P. asahinus</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. falcispinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="falcispinus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="61">P. falcispinus</emphasis>
|
||||
<taxonomicName id="E59ABAD79ACF52444610E86ED69B7692" class="Insecta" kingdom="Animalia" lsidName="P. falcispinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="falcispinus">
|
||||
<emphasis id="9B61B23B1F0D346B1546DBC0658B7086" italics="true" pageId="0" pageNumber="61">P. falcispinus</emphasis>
|
||||
</taxonomicName>
|
||||
might be conspecific. However, this hypothesis remained untested because males from the type locality of
|
||||
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="61">P. asahinus</emphasis>
|
||||
<taxonomicName id="D71D3548982F3ED827F1F6EB18653CC6" class="Insecta" kingdom="Animalia" lsidName="P. asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus">
|
||||
<emphasis id="FAA19F3FFCB610ECD53CC5DBB2B6EF8E" italics="true" pageId="0" pageNumber="61">P. asahinus</emphasis>
|
||||
</taxonomicName>
|
||||
have not been examined. The male specimen examined here was obtained from a valley southwest of Mt.
|
||||
<normalizedToken originalValue="Dôrokujinpô">Dorokujinpo</normalizedToken>
|
||||
<normalizedToken id="E451D16D69926DB16ABA81ED288F9A37" originalValue="Dôrokujinpô">Dorokujinpo</normalizedToken>
|
||||
, which can virtually be regarded as the type locality (Fig.
|
||||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of the Pterostichus macrogenys species group in northern Chubu, northern Kanto, and southern Tohoku Districts, Honshu, based on specimens with unambiguous identities, modified from Sasakawa et al. (2020) and Sato (2021) 1 P. chokaisanus Sasakawa, 2009 2 P. kurikomasanus Sasakawa, 2005 3 P. asahinus Habu and Baba, 1960 4 P. takadateyamanus Sasakawa, 2009 5 P. adatarasanus Sasakawa, 2005 6 P. gassanus Sasakawa, 2009 7 P. shinbodakensis Sasakawa & Ito, 2017 8 P. awashimaensis sp. nov. 9 P. iwakiensis Sasakawa, 2009 10 P. eboshiyamanus Sasakawa, 2009 11 P. tateishiyamanus Sasakawa & Ito, 2017 12 P. monolineatus Sasakawa, Mitsuduka, & Ito, 2020 13 P. yamizosanus Sasakawa, 2005 14 P. yahikosanus Sasakawa, 2009 15 P. ohsawacavus Sasakawa, 2005 16 P. sumondakensis Sasakawa, 2005 17 P. isolatus Sasakawa, 2005 18 P. nakamiyorinus Morita, Ohkawa & Kurihara, 2013 19 P. momuranus Morita, Ohkawa & Kurihara, 2013 20 P. macrogenys Bates, 1883 21 P. nagasawai Ito & Ogai, 2015. Red letters denote the type localities of each species. The blue letter indicates where the male specimen of P. asahinus newly examined here was collected." figureDoi="10.3897/subtbiol.43.80969.figure1" httpUri="https://binary.pensoft.net/fig/690340" pageId="0" pageNumber="61">1</figureCitation>
|
||||
<figureCitation id="DCD2E592295600B4358C8033E282E551" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of the Pterostichus macrogenys species group in northern Chubu, northern Kanto, and southern Tohoku Districts, Honshu, based on specimens with unambiguous identities, modified from Sasakawa et al. (2020) and Sato (2021) 1 P. chokaisanus Sasakawa, 2009 2 P. kurikomasanus Sasakawa, 2005 3 P. asahinus Habu and Baba, 1960 4 P. takadateyamanus Sasakawa, 2009 5 P. adatarasanus Sasakawa, 2005 6 P. gassanus Sasakawa, 2009 7 P. shinbodakensis Sasakawa & Ito, 2017 8 P. awashimaensis sp. nov. 9 P. iwakiensis Sasakawa, 2009 10 P. eboshiyamanus Sasakawa, 2009 11 P. tateishiyamanus Sasakawa & Ito, 2017 12 P. monolineatus Sasakawa, Mitsuduka, & Ito, 2020 13 P. yamizosanus Sasakawa, 2005 14 P. yahikosanus Sasakawa, 2009 15 P. ohsawacavus Sasakawa, 2005 16 P. sumondakensis Sasakawa, 2005 17 P. isolatus Sasakawa, 2005 18 P. nakamiyorinus Morita, Ohkawa & Kurihara, 2013 19 P. momuranus Morita, Ohkawa & Kurihara, 2013 20 P. macrogenys Bates, 1883 21 P. nagasawai Ito & Ogai, 2015. Red letters denote the type localities of each species. The blue letter indicates where the male specimen of P. asahinus newly examined here was collected." figureDoi="10.3897/subtbiol.43.80969.figure1" httpUri="https://binary.pensoft.net/fig/690340" pageId="0" pageNumber="61">1</figureCitation>
|
||||
). The structures of the endophallus and right paramere of this specimen are identical to those of the eastern type of
|
||||
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. falcispinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="falcispinus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="61">P. falcispinus</emphasis>
|
||||
<taxonomicName id="B41238298508BE887CDCA0CC00620A31" class="Insecta" kingdom="Animalia" lsidName="P. falcispinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="falcispinus">
|
||||
<emphasis id="19EB4568B64EF843840664D754FEE54E" italics="true" pageId="0" pageNumber="61">P. falcispinus</emphasis>
|
||||
</taxonomicName>
|
||||
in
|
||||
<bibRefCitation DOI="https://doi.org/10.3390/insects11110803" author="Sasakawa, K" journalOrPublisher="Applied Entomology and Zoology" pageId="0" pageNumber="61" refId="B17" refString="Sasakawa, K, Mitsuduka, Y, Ito, H, 2020. Unexpected species identities and interspecific relationships in a subterranean beetle lineage, the Pterostichus macrogenys species group (Coleoptera, Carabidae), revealed by fine-scale field sampling and detailed morphological comparisons. Insects 11: 803. https://doi.org/10.3390/insects11110803" title="Unexpected species identities and interspecific relationships in a subterranean beetle lineage, the Pterostichus macrogenys species group (Coleoptera, Carabidae), revealed by fine-scale field sampling and detailed morphological comparisons. Insects 11: 803." url="https://doi.org/10.3390/insects11110803" year="2020">Sasakawa et al. (2020)</bibRefCitation>
|
||||
<bibRefCitation id="5A9901DF8F95663E96AD7A7D6795A3AE" DOI="https://doi.org/10.3390/insects11110803" author="Sasakawa, K" journalOrPublisher="Applied Entomology and Zoology" pageId="0" pageNumber="61" refId="B17" refString="Sasakawa, K, Mitsuduka, Y, Ito, H, 2020. Unexpected species identities and interspecific relationships in a subterranean beetle lineage, the Pterostichus macrogenys species group (Coleoptera, Carabidae), revealed by fine-scale field sampling and detailed morphological comparisons. Insects 11: 803. https://doi.org/10.3390/insects11110803" title="Unexpected species identities and interspecific relationships in a subterranean beetle lineage, the Pterostichus macrogenys species group (Coleoptera, Carabidae), revealed by fine-scale field sampling and detailed morphological comparisons. Insects 11: 803." url="https://doi.org/10.3390/insects11110803" year="2020">Sasakawa et al. (2020)</bibRefCitation>
|
||||
. Its body length is intermediate between that of the eastern and western types of
|
||||
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. falcispinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="falcispinus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="61">P. falcispinus</emphasis>
|
||||
<taxonomicName id="15D598B8812832DC7A5E727EA11AAAAD" class="Insecta" kingdom="Animalia" lsidName="P. falcispinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="falcispinus">
|
||||
<emphasis id="EDAC239DA8C5136EAE651B4B2CF6D2C6" italics="true" pageId="0" pageNumber="61">P. falcispinus</emphasis>
|
||||
</taxonomicName>
|
||||
as follows: body length from mandible apices to elytral end (BLm) 16.32 mm, that from anterior margin of labrum (BLl) 14.72 mm, and that from clypeal apex (BLc) 14.26 mm. Based on these results, we regard
|
||||
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="61">P. asahinus</emphasis>
|
||||
<taxonomicName id="EC582803158B1A838C851F1A47D732AB" class="Insecta" kingdom="Animalia" lsidName="P. asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus">
|
||||
<emphasis id="1C6CE692801B7ABFA746AD06387132A9" italics="true" pageId="0" pageNumber="61">P. asahinus</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. falcispinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="falcispinus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="61">P. falcispinus</emphasis>
|
||||
<taxonomicName id="2B5A579630DD1D4E3A2EB8B081008196" class="Insecta" kingdom="Animalia" lsidName="P. falcispinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="falcispinus">
|
||||
<emphasis id="B60EA8A39C94A3D547E21BAF4F6E4DA8" italics="true" pageId="0" pageNumber="61">P. falcispinus</emphasis>
|
||||
</taxonomicName>
|
||||
as conspecific. The results of the discriminant analysis support this conclusion. The
|
||||
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="61">P. asahinus</emphasis>
|
||||
<taxonomicName id="EA142E96EBC1ECB51C9DF3B28626E5B2" class="Insecta" kingdom="Animalia" lsidName="P. asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus">
|
||||
<emphasis id="E2346FE6EC75BA6F3C67A344B86B2ADF" italics="true" pageId="0" pageNumber="61">P. asahinus</emphasis>
|
||||
</taxonomicName>
|
||||
holotype was classified in this newly defined
|
||||
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="61">P. asahinus</emphasis>
|
||||
<taxonomicName id="6A87BD3C14AA0F52EF4AA856309EAA78" class="Insecta" kingdom="Animalia" lsidName="P. asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus">
|
||||
<emphasis id="7CDBCADA82C88EF46CEA0D1CD279DF1B" italics="true" pageId="0" pageNumber="61">P. asahinus</emphasis>
|
||||
</taxonomicName>
|
||||
in the discriminant function analysis, and was within the area of
|
||||
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="61">P. asahinus</emphasis>
|
||||
<taxonomicName id="963053FF9C22785DDE3F0AEFA0656630" class="Insecta" kingdom="Animalia" lsidName="P. asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus">
|
||||
<emphasis id="DDDA5B5D2C6207B18282B7FFB0DE903C" italics="true" pageId="0" pageNumber="61">P. asahinus</emphasis>
|
||||
</taxonomicName>
|
||||
on the scatterplot of the first two canonical variates (Fig.
|
||||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Scatterplot of the first two canonical variates obtained from the discriminant analysis of four measurements of the female pronotum." figureDoi="10.3897/subtbiol.43.80969.figure3" httpUri="https://binary.pensoft.net/fig/690342" pageId="0" pageNumber="61">3</figureCitation>
|
||||
<figureCitation id="E55AC29C51F45B2201611131D3C8D199" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Scatterplot of the first two canonical variates obtained from the discriminant analysis of four measurements of the female pronotum." figureDoi="10.3897/subtbiol.43.80969.figure3" httpUri="https://binary.pensoft.net/fig/690342" pageId="0" pageNumber="61">3</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption doi="10.3897/subtbiol.43.80969.figure3" httpUri="https://binary.pensoft.net/fig/690342" pageId="0" pageNumber="61" start="Figure 3" startId="F3">
|
||||
<paragraph pageId="0" pageNumber="61">
|
||||
<emphasis bold="true" pageId="0" pageNumber="61">Figure 3.</emphasis>
|
||||
<caption id="72CF9B3270DD4E0DAE1C6A9E9F0EB1B6" doi="10.3897/subtbiol.43.80969.figure3" httpUri="https://binary.pensoft.net/fig/690342" pageId="0" pageNumber="61" start="Figure 3" startId="F3">
|
||||
<paragraph id="5FC33BCDEE140833096E753CA012C09D" pageId="0" pageNumber="61">
|
||||
<emphasis id="8B6983CF2932C65FECA19B9D7CDAF4C6" bold="true" pageId="0" pageNumber="61">Figure 3.</emphasis>
|
||||
Scatterplot of the first two canonical variates obtained from the discriminant analysis of four measurements of the female pronotum.
|
||||
</paragraph>
|
||||
</caption>
|
||||
|
|
|
|||
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