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data/91/1B/E6/911BE60E5543FFEC8C03FD49FF61817A.xml
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<mods:titleInfo>
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<mods:title>Incrementing and clarifying the diversity and early evolution of termites (Blattodea: Isoptera)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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||||
</mods:role>
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||||
<mods:namePart>Jouault, Corentin</mods:namePart>
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||||
</mods:name>
|
||||
<mods:name type="personal">
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||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Engel, Michael S.</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Fls</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Legendre, Frédéric</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Huang, Diying</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Grandcolas, Philippe</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Nel, André</mods:namePart>
|
||||
</mods:name>
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||||
<mods:typeOfResource>text</mods:typeOfResource>
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||||
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<mods:part>
|
||||
<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-02-07</mods:number>
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<mods:detail type="volume">
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<mods:number>196</mods:number>
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<mods:classification>journal article</mods:classification>
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||||
<mods:identifier type="DOI">10.1093/zoolinnean/zlac064</mods:identifier>
|
||||
<mods:identifier type="ISSN">0024-4082</mods:identifier>
|
||||
<mods:identifier type="Zenodo-Dep">7195105</mods:identifier>
|
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<mods:identifier type="ZooBank">01556715-E3DE-45E0-B620-ABD11BDB0748</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:911BE60E5543FFEC8C03FD49FF61817A" httpUri="http://treatment.plazi.org/id/911BE60E5543FFEC8C03FD49FF61817A" lastPageNumber="617" pageId="9" pageNumber="617">
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<subSubSection pageId="9" pageNumber="617" type="nomenclature">
|
||||
<paragraph blockId="9.[832,1402,661,715]" pageId="9" pageNumber="617">
|
||||
<collectionCode box="[832,911,661,686]" pageId="9" pageNumber="617">GENUS</collectionCode>
|
||||
<taxonomicName authority="JOUAULT, ENGEL & NEL" authorityName="KRISHNAI JOUAULT, ENGEL & NEL" authorityYear="2022" box="[919,1402,661,686]" class="Insecta" family="Mastotermitidae" genus="Magnifitermes" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="617" phylum="Arthropoda" rank="genus" status="GEN. NOV.">
|
||||
<emphasis bold="true" box="[919,1118,661,684]" italics="true" pageId="9" pageNumber="617">MAGNIFITERMES</emphasis>
|
||||
JOUAULT, ENGEL & NEL
|
||||
</taxonomicName>
|
||||
<emphasis bold="true" box="[1057,1176,691,715]" pageId="9" pageNumber="617">
|
||||
<taxonomicNameLabel box="[1057,1176,691,715]" pageId="9" pageNumber="617">GEN. NOV.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="9" pageNumber="617" type="description">
|
||||
<paragraph blockId="9.[809,1426,732,815]" pageId="9" pageNumber="617">
|
||||
<emphasis box="[809,1076,732,753]" italics="true" pageId="9" pageNumber="617">Zoobank registration:</emphasis>
|
||||
http://zoobank.org/urn:lsid: z o o b a n k. o r g:a c t:
|
||||
<uuid pageId="9" pageNumber="617">B D E A7C 1 1- 0 2 9 6- 4 7F 8- A7 3 5 - 817A4D58F844</uuid>
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||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="9" pageNumber="617" type="etymology">
|
||||
<paragraph blockId="9.[809,1426,840,954]" pageId="9" pageNumber="617">
|
||||
<emphasis box="[809,930,840,861]" italics="true" pageId="9" pageNumber="617">Etymology:</emphasis>
|
||||
The generic name is a combination of the Latin adjective
|
||||
<emphasis box="[914,1036,871,892]" italics="true" pageId="9" pageNumber="617">magnificus</emphasis>
|
||||
, magnificent, and the noun
|
||||
<emphasis box="[1347,1419,871,892]" italics="true" pageId="9" pageNumber="617">termes</emphasis>
|
||||
, meaning ‘maggot’ or ‘termite’, and commonly used as the root for many genera of
|
||||
<taxonomicName authorityName="BRULLE" authorityYear="1832" box="[1067,1159,932,953]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Isoptera" pageId="9" pageNumber="617" phylum="Tracheophyta" rank="order">Isoptera</taxonomicName>
|
||||
. Gender masculine.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="9" pageNumber="617" type="type_taxon">
|
||||
<paragraph blockId="9.[808,1425,978,1031]" pageId="9" pageNumber="617">
|
||||
<emphasis box="[809,952,979,1000]" italics="true" pageId="9" pageNumber="617">
|
||||
<typeStatus box="[809,862,979,1000]" pageId="9" pageNumber="617">Type</typeStatus>
|
||||
species:
|
||||
</emphasis>
|
||||
<taxonomicName authority="Jouault, Engel & Nel" authorityName="Jouault, Engel & Nel" authorityYear="2022" class="Insecta" family="Mastotermitidae" genus="Magnifitermes" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="617" phylum="Arthropoda" rank="species" species="krishnai" status="sp. nov.">
|
||||
<emphasis box="[958,1226,978,1000]" italics="true" pageId="9" pageNumber="617">Magnifitermes krishnai</emphasis>
|
||||
Jouault, Engel & Nel
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel box="[856,939,1010,1031]" pageId="9" pageNumber="617" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="9" pageNumber="617" type="diagnosis">
|
||||
<paragraph blockId="9.[809,1425,1056,1906]" pageId="9" pageNumber="617">
|
||||
<emphasis box="[809,939,1056,1077]" italics="true" pageId="9" pageNumber="617">Diagnosis:</emphasis>
|
||||
Imago. Head massive, square-shaped, dorsally flattened, with posterior corners broadly rounded; epicranial scar present; ocelli present, located just above compound eyes (not separated from eyes by a space greater than their length); postclypeus trapezoidal and flat; anteclypeus anteriorly convex; left mandible with an apical and two distinct marginal teeth, LM1-LM2 interdental space broad, v-shaped, posterior margin of LM1 slightly longer than anterior margin LM2; pronotum massive, trapezoidal, wider than head, anterior margin concave surrounding head, posterior margin broadly convex, with dorsal surface flatly arched in profile, sides convex; tarsi pentamerous, with fifth tarsomere extremely elongate; tibial spur formula 3-4-4; protibia with two additional spines (fa, fb); mesotibia with four(?) additional spines (ma, mb, mc, md); metatibia with at least three additional spines (ha, hb, hc); pretarsal arolium present; forewing scale large, overlapping hindwing scale, claval suture (PCu) broadly arched, basal suture broadly arched; Sc, RA, RP, M heavily sclerotized; Sc with two main branches; R branched within scale; RP running parallel to costal margin, with at least five branches joining costal margin; M slightly closer to RP than to CuA, with at least four secondary branches radiating to apical margin; CuA less sclerotized, with several branches joining lower margin. Hindwing: scale (basal) suture absent; anal lobe (ano-jugal) developed (folded)
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
238
data/91/1B/E6/911BE60E5546FFEB8FF0F907FF1B8168.xml
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238
data/91/1B/E6/911BE60E5546FFEB8FF0F907FF1B8168.xml
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|
|
@ -0,0 +1,238 @@
|
|||
<document ID-DOI="10.1093/zoolinnean/zlac064" ID-ISSN="0024-4082" ID-Zenodo-Dep="7195105" ID-ZooBank="01556715-E3DE-45E0-B620-ABD11BDB0748" approvalRequired="2" approvalRequired_for_treatments="2" checkinTime="1665471643038" checkinUser="plazi" docAuthor="Jouault, Corentin, Engel, Michael S., Fls, Legendre, Frédéric, Huang, Diying, Grandcolas, Philippe & Nel, André" docDate="2022" docId="911BE60E5546FFEB8FF0F907FF1B8168" docLanguage="en" docName="zlac064.pdf" docOrigin="Zoological Journal of the Linnean Society 196" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-.journal_article" docStyleVersion="7" docTitle="Mastotermes myanmarensis Jouault, Engel, Fls, Legendre, Huang, Grandcolas & Nel, 2022, SP. NOV." docType="treatment" docVersion="3" lastPageNumber="621" masterDocId="6D229E76554AFFE58F43FFDCFA5D8608" masterDocTitle="Incrementing and clarifying the diversity and early evolution of termites (Blattodea: Isoptera)" masterLastPageNumber="629" masterPageNumber="608" pageNumber="620" updateTime="1665701992877" updateUser="carolina">
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<mods:title>Incrementing and clarifying the diversity and early evolution of termites (Blattodea: Isoptera)</mods:title>
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<mods:namePart>Jouault, Corentin</mods:namePart>
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<mods:name type="personal">
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|
||||
<mods:roleTerm>Author</mods:roleTerm>
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||||
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|
||||
<mods:namePart>Engel, Michael S.</mods:namePart>
|
||||
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|
||||
<mods:name type="personal">
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||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
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||||
</mods:role>
|
||||
<mods:namePart>Fls</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Legendre, Frédéric</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
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||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
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||||
</mods:role>
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||||
<mods:namePart>Huang, Diying</mods:namePart>
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||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
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||||
<mods:namePart>Grandcolas, Philippe</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
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||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
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||||
</mods:role>
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||||
<mods:namePart>Nel, André</mods:namePart>
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||||
</mods:name>
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||||
<mods:typeOfResource>text</mods:typeOfResource>
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||||
<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zoological Journal of the Linnean Society</mods:title>
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</mods:titleInfo>
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||||
<mods:part>
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||||
<mods:date>2022</mods:date>
|
||||
<mods:detail type="pubDate">
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<mods:number>2022-02-07</mods:number>
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||||
</mods:detail>
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<mods:detail type="volume">
|
||||
<mods:number>196</mods:number>
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<mods:extent unit="page">
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<mods:start>608</mods:start>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1093/zoolinnean/zlac064</mods:identifier>
|
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<mods:identifier type="ISSN">0024-4082</mods:identifier>
|
||||
<mods:identifier type="Zenodo-Dep">7195105</mods:identifier>
|
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<mods:identifier type="ZooBank">01556715-E3DE-45E0-B620-ABD11BDB0748</mods:identifier>
|
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</mods:mods>
|
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7184602" ID-Zenodo-Dep="7184602" LSID="urn:lsid:plazi:treatment:911BE60E5546FFEB8FF0F907FF1B8168" httpUri="http://treatment.plazi.org/id/911BE60E5546FFEB8FF0F907FF1B8168" lastPageId="14" lastPageNumber="621" pageId="12" pageNumber="620">
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<paragraph blockId="12.[179,763,1755,1780]" box="[179,763,1755,1780]" pageId="12" pageNumber="620">
|
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<heading box="[179,763,1755,1780]" centered="true" fontSize="9" level="2" pageId="12" pageNumber="620" reason="2">
|
||||
<taxonomicName authorityName="Jouault & Engel & Fls & Legendre & Huang & Grandcolas & Nel" authorityYear="2022" box="[179,547,1755,1778]" class="Insecta" family="Mastotermitidae" genus="Mastotermes" kingdom="Animalia" order="Blattodea" pageId="12" pageNumber="622" phylum="Arthropoda" rank="species" species="myanmarensis">
|
||||
<emphasis bold="true" box="[179,547,1755,1778]" italics="true" pageId="12" pageNumber="620">MASTOTERMES MYANMARENSIS</emphasis>
|
||||
</taxonomicName>
|
||||
JOUAULT
|
||||
<emphasis bold="true" box="[664,763,1755,1779]" pageId="12" pageNumber="620">
|
||||
<taxonomicNameLabel box="[664,763,1755,1779]" pageId="12" pageNumber="620" rank="species">SP. NOV.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="12" pageNumber="620" type="description">
|
||||
<paragraph blockId="12.[404,537,1794,1819]" box="[404,537,1794,1819]" pageId="12" pageNumber="620">
|
||||
(
|
||||
<figureCitation box="[413,487,1794,1819]" captionStart="Figure 9" captionStartId="13.[145,223,1368,1390]" captionTargetBox="[149,1421,198,1325]" captionTargetId="figure-258@13.[145,1425,195,1329]" captionTargetPageId="13" captionText="Figure 9. Mastotermes myanmarensis sp. nov., holotype IGR.BU-054. A, forewing; B, hindwing. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7195139" httpUri="https://zenodo.org/record/7195139/files/figure.png" pageId="12" pageNumber="620">FIGS 9</figureCitation>
|
||||
,
|
||||
<figureCitation box="[501,529,1794,1819]" captionStart="Figure 10" captionStartId="14.[163,243,1469,1491]" captionTargetBox="[166,1440,199,1425]" captionTargetId="figure-192@14.[163,1443,195,1429]" captionTargetPageId="14" captionText="Figure 10. Mastotermes myanmarensis sp. nov., holotype IGR.BU-054. Line drawings with veins labelled (cross-veins omitted): A, forewing; B, hindwing. A1, first anal vein; CuA, anterior cubitus; M, media; PCu, postcubitus; RA, anterior radius; RP, posterior radius; Sc, subcosta. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7195141" httpUri="https://zenodo.org/record/7195141/files/figure.png" pageId="12" pageNumber="620">10</figureCitation>
|
||||
)
|
||||
</paragraph>
|
||||
<paragraph blockId="12.[163,779,1834,1917]" pageId="12" pageNumber="620">
|
||||
<emphasis box="[163,483,1834,1856]" italics="true" pageId="12" pageNumber="620">Z o o b a n k r e g i s t r a t i o n:</emphasis>
|
||||
h t t p: / / z o o b a n k. o r g /
|
||||
<uri pageId="12" pageNumber="620">
|
||||
urn:lsid:zoobank.org:act:
|
||||
<uuid pageId="12" pageNumber="620">F5CF600D-2068-424E-B255- 9F92051F952B</uuid>
|
||||
</uri>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="12" pageNumber="620" type="materials_examined">
|
||||
<paragraph blockId="12.[827,1442,789,933]" pageId="12" pageNumber="620">
|
||||
<emphasis box="[827,941,789,810]" italics="true" pageId="12" pageNumber="620">Material:</emphasis>
|
||||
<materialsCitation country="France" county="Rennes" location="Geological Department" municipality="Museum" pageId="12" pageNumber="620" specimenCount="1" typeStatus="holotype">
|
||||
<typeStatus box="[951,1063,789,811]" pageId="12" pageNumber="620">Holotype</typeStatus>
|
||||
, IGR.BU-054, preserved in an elongate, pale and ovoid piece of amber measuring 50 × 20 ×
|
||||
<quantity box="[945,1016,850,872]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="620" unit="mm" value="9.0">9 mm</quantity>
|
||||
, housed in the amber collection of the
|
||||
<location LSID="urn:lsid:plazi:treatment:911BE60E5546FFEB8FF0F907FF1B8168:1C6D01C35546FFE98C2CFCADFEC5858E" box="[879,1176,881,903]" country="France" county="Rennes" municipality="Museum" name="Geological Department" pageId="12" pageNumber="620">Geological Department</location>
|
||||
and
|
||||
<collectingMunicipality box="[1246,1353,881,902]" pageId="12" pageNumber="620">Museum</collectingMunicipality>
|
||||
(IGR),
|
||||
<collectingCounty box="[827,913,912,933]" pageId="12" pageNumber="620">Rennes</collectingCounty>
|
||||
,
|
||||
<collectingCountry box="[923,1005,912,933]" name="France" pageId="12" pageNumber="620">France</collectingCountry>
|
||||
.
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
<paragraph blockId="12.[827,1443,958,1041]" pageId="12" pageNumber="620">
|
||||
<emphasis box="[827,1080,958,979]" italics="true" pageId="12" pageNumber="620">Locality and horizon:</emphasis>
|
||||
Hkamti site, Hkamti District,
|
||||
<collectingRegion box="[827,1008,989,1011]" country="Myanmar" name="Sagaing" pageId="12" pageNumber="620">Sagaing Region</collectingRegion>
|
||||
,
|
||||
<collectingCountry box="[1019,1133,989,1010]" name="Myanmar" pageId="12" pageNumber="620">Myanmar</collectingCountry>
|
||||
; Early Albian (
|
||||
<emphasis box="[1306,1323,989,1010]" italics="true" pageId="12" pageNumber="620">c.</emphasis>
|
||||
110 Mya), Early Cretaceous.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="12" pageNumber="620" type="etymology">
|
||||
<paragraph blockId="12.[827,1444,1066,1180]" pageId="12" pageNumber="620">
|
||||
<emphasis box="[827,956,1066,1087]" italics="true" pageId="12" pageNumber="620">Etymology:</emphasis>
|
||||
The specific epithet refers to the country of origin of the amber piece,
|
||||
<collectingCountry box="[1156,1268,1097,1118]" name="Myanmar" pageId="12" pageNumber="620">Myanmar</collectingCountry>
|
||||
, combining the country name with the Latin toponymic adjectival suffix, –
|
||||
<emphasis box="[917,973,1159,1180]" italics="true" pageId="12" pageNumber="620">ēnsis</emphasis>
|
||||
, meaning ‘from’.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="12" pageNumber="620" type="diagnosis">
|
||||
<paragraph blockId="12.[827,1444,1205,1840]" pageId="12" pageNumber="620">
|
||||
<emphasis box="[827,944,1205,1226]" italics="true" pageId="12" pageNumber="620">Diagnosis:</emphasis>
|
||||
Imago. Wings membranous, long and broad, densely reticulate, with ‘cross-veins’ present; veins Sc, RA, RP and M more heavily sclerotized than CuA, reticulate veins obviously pigmented among radial and medial fields; forewing Sc with two main branches; R with two branches; width of radial field moderate, RA simple, RP with five main branches; medial field encompassing wing apex, M main vein closely parallel to RP main vein, with three main branches, first branch of M arising near apical-third of wing; CuA with five main branches, apical-most branch of CuA terminating into posterior margin and in apical-third of forewing length. Hindwing with large anal lobe; costal space wide; Sc simple, terminating on costal margin at basal-third of hindwing length; RA long, simple and terminating on costal margin before apical-third of hindwing length; RP with two main branches; medial field encompassing wing apex; M separating from RP near wing base, relatively closely parallel to RP, with three main branches, secondary branches present; CuA uniformly branching, terminating just posterior to wing apex.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection lastPageId="13" lastPageNumber="621" pageId="12" pageNumber="620" type="description">
|
||||
<paragraph blockId="12.[827,1442,1865,1917]" lastBlockId="13.[145,763,1450,1901]" lastPageId="13" lastPageNumber="621" pageId="12" pageNumber="620">
|
||||
<emphasis box="[827,971,1865,1886]" italics="true" pageId="12" pageNumber="620">Description:</emphasis>
|
||||
Imago. Wings membranous, reticulate veins present; forewing scale part missing, forewing preserved part at least
|
||||
<quantity box="[424,541,1450,1472]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="13" pageNumber="621" unit="mm" value="10.7">10.70 mm</quantity>
|
||||
long and
|
||||
<quantity box="[660,762,1450,1472]" metricMagnitude="-3" metricUnit="m" metricValue="4.62" pageId="13" pageNumber="621" unit="mm" value="4.62">4.62 mm</quantity>
|
||||
wide, apex rounded, middle section of posterior slightly convex basally then convex; veins Sc, RA, RP and M more heavily sclerotized than CuA and A; forewing Sc short, bifurcate at suture, posterior branch with a faint subbranch at margin; RA with one branch; radial field width moderate, occupying about one-sixth area on average; RP with five branches, proximal branch fading apically before anterior wing margin; M closely parallel to RP, with three main branches; CuA with numerous posterior branches, apical-most branch of CuA terminating on posterior margin well proximal to apex, more proximal branches loosely pectinate, apicalmost of these branches terminating on margin near wing midlength.
|
||||
</paragraph>
|
||||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7195139" ID-Zenodo-Dep="7195139" box="[145,1335,1368,1390]" httpUri="https://zenodo.org/record/7195139/files/figure.png" pageId="13" pageNumber="621" startId="13.[145,223,1368,1390]" targetBox="[149,1421,198,1325]" targetPageId="13">
|
||||
<paragraph blockId="13.[145,1335,1368,1390]" box="[145,1335,1368,1390]" pageId="13" pageNumber="621">
|
||||
<emphasis bold="true" box="[145,249,1368,1390]" pageId="13" pageNumber="621">Figure 9.</emphasis>
|
||||
<taxonomicName authority="Jouault & Engel & Fls & Legendre & Huang & Grandcolas & Nel, 2022" authorityName="Jouault & Engel & Fls & Legendre & Huang & Grandcolas & Nel" authorityYear="2022" box="[260,551,1368,1390]" class="Insecta" family="Mastotermitidae" genus="Mastotermes" kingdom="Animalia" order="Blattodea" pageId="13" pageNumber="621" phylum="Arthropoda" rank="species" species="myanmarensis" status="sp. nov.">
|
||||
<emphasis box="[260,551,1368,1390]" italics="true" pageId="13" pageNumber="621">Mastotermes myanmarensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel box="[558,636,1368,1390]" pageId="13" pageNumber="621" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
, holotype IGR.BU-054. A, forewing; B, hindwing. Scale bars: 2 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph blockId="13.[809,1426,1450,1779]" pageId="13" pageNumber="621">
|
||||
Hindwing with large anal lobe, scale not preserved,
|
||||
<quantity box="[810,925,1481,1502]" metricMagnitude="-2" metricUnit="m" metricValue="1.41" pageId="13" pageNumber="621" unit="mm" value="14.1">14.10 mm</quantity>
|
||||
long and 6.0 mm wide with anal lobe and
|
||||
<quantity box="[810,911,1511,1533]" metricMagnitude="-3" metricUnit="m" metricValue="5.16" pageId="13" pageNumber="621" unit="mm" value="5.16">5.16 mm</quantity>
|
||||
wide without anal lobe; Sc longer than that of forewing; RA long, simple and terminating on costal margin before apical-third of hindwing length; RP with three branches, first branch originating just before point of contact between Sc and wing margin; M with four branches, separating from RP away from wing base, then closely parallel to RP, apical branch of M with a secondary branch; CuA with numerous posterior branches, some dichotomous, terminating posterior to wing apex.
|
||||
</paragraph>
|
||||
<paragraph blockId="13.[809,1067,1803,1825]" box="[809,1067,1803,1825]" pageId="13" pageNumber="621">
|
||||
<heading box="[809,1067,1803,1825]" centered="true" fontSize="9" level="2" pageId="13" pageNumber="621" reason="2">
|
||||
<emphasis box="[809,892,1803,1824]" italics="true" pageId="13" pageNumber="621">Colour:</emphasis>
|
||||
Not preserved.
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection lastPageId="14" lastPageNumber="622" pageId="13" pageNumber="621" type="discussion">
|
||||
<paragraph blockId="13.[809,1424,1850,1903]" lastBlockId="14.[827,1443,1591,1889]" lastPageId="14" lastPageNumber="622" pageId="13" pageNumber="621">
|
||||
<emphasis box="[809,918,1850,1871]" italics="true" pageId="13" pageNumber="621">Remarks:</emphasis>
|
||||
Compared with the known ‘Mid’-Cretaceous fossil and the extant mastotermitids,
|
||||
<taxonomicName authorityName="Jouault & Engel & Fls & Legendre & Huang & Grandcolas & Nel" authorityYear="2022" box="[1219,1424,1881,1902]" class="Insecta" family="Mastotermitidae" genus="Mastotermes" kingdom="Animalia" order="Blattodea" pageId="13" pageNumber="621" phylum="Arthropoda" rank="species" species="myanmarensis">
|
||||
<emphasis box="[1219,1424,1881,1902]" italics="true" pageId="13" pageNumber="621">M. myanmarensis</emphasis>
|
||||
</taxonomicName>
|
||||
stands out in having broadly rounded wings, wider intervals between the longitudinal veins (shared with
|
||||
<taxonomicName authority="Zhao et al., 2019" authorityName="Zhao" authorityYear="2019" box="[162,548,1652,1674]" class="Insecta" family="Mastotermitidae" genus="Mastotermes" kingdom="Animalia" order="Blattodea" pageId="14" pageNumber="622" phylum="Arthropoda" rank="species" species="monostichus">
|
||||
<emphasis box="[162,344,1652,1674]" italics="true" pageId="14" pageNumber="622">M. monostichus</emphasis>
|
||||
<bibRefCitation author="Zhao Z & Eggleton P & Yin X & Gao T & Shih C & Ren D" box="[352,548,1652,1674]" pageId="14" pageNumber="622" pagination="612 - 623" refId="ref14180" refString="Zhao Z, Eggleton P, Yin X, Gao T, Shih C, Ren D. 2019. The oldest known mastotermitids (Blattodea: Termitoidae) and phylogeny of basal termites. Systematic Entomology 44: 612 - 623." type="journal article" year="2019">
|
||||
Zhao
|
||||
<emphasis box="[418,468,1652,1674]" italics="true" pageId="14" pageNumber="622">et al</emphasis>
|
||||
., 2019
|
||||
</bibRefCitation>
|
||||
</taxonomicName>
|
||||
), and a broad space above Sc on the hindwing. Additionally, the space occupied by Sc + RA + RP is extremely wide near the basal suture.
|
||||
<taxonomicName authorityName="Jouault & Engel & Fls & Legendre & Huang & Grandcolas & Nel" authorityYear="2022" box="[374,705,1745,1766]" class="Insecta" family="Mastotermitidae" genus="Mastotermes" kingdom="Animalia" order="Blattodea" pageId="14" pageNumber="622" phylum="Arthropoda" rank="species" species="myanmarensis">
|
||||
<emphasis box="[374,705,1745,1766]" italics="true" pageId="14" pageNumber="622">Mastotermes myanmarensis</emphasis>
|
||||
</taxonomicName>
|
||||
has a hindwing with vein A1 without branches, while the extant
|
||||
<taxonomicName baseAuthorityName="Tillyard" baseAuthorityYear="1931" box="[245,432,1805,1827]" class="Insecta" family="Mastotermitidae" genus="Mastotermes" kingdom="Animalia" order="Blattodea" pageId="14" pageNumber="622" phylum="Arthropoda" rank="species" species="darwiniensis">
|
||||
<emphasis box="[245,432,1805,1827]" italics="true" pageId="14" pageNumber="622">M. darwiniensis</emphasis>
|
||||
</taxonomicName>
|
||||
has a vein A1 with numerous posterior branches (similar to
|
||||
<taxonomicName box="[562,761,1836,1858]" class="Insecta" family="Mastotermitidae" genus="Mastotermes" kingdom="Animalia" order="Blattodea" pageId="14" pageNumber="622" phylum="Arthropoda" rank="species" species="monostichus">
|
||||
<emphasis box="[562,761,1836,1858]" italics="true" pageId="14" pageNumber="622">M. monostichus</emphasis>
|
||||
</taxonomicName>
|
||||
), and the anal veins in the anal lobe are all simple (vs. with some dichotomies in
|
||||
<taxonomicName baseAuthorityName="Tillyard" baseAuthorityYear="1931" box="[1196,1389,1591,1612]" class="Insecta" family="Mastotermitidae" genus="Mastotermes" kingdom="Animalia" order="Blattodea" pageId="14" pageNumber="622" phylum="Arthropoda" rank="species" species="darwiniensis">
|
||||
<emphasis box="[1196,1389,1591,1612]" italics="true" pageId="14" pageNumber="622">M. darwiniensis</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName box="[828,1010,1621,1643]" class="Insecta" family="Mastotermitidae" genus="Mastotermes" kingdom="Animalia" order="Blattodea" pageId="14" pageNumber="622" phylum="Arthropoda" rank="species" species="monostichus">
|
||||
<emphasis box="[828,1010,1621,1643]" italics="true" pageId="14" pageNumber="622">M. monostichus</emphasis>
|
||||
</taxonomicName>
|
||||
).
|
||||
<taxonomicName authorityName="Jouault & Engel & Fls & Legendre & Huang & Grandcolas & Nel" authorityYear="2022" box="[1033,1360,1622,1643]" class="Insecta" family="Mastotermitidae" genus="Mastotermes" kingdom="Animalia" order="Blattodea" pageId="14" pageNumber="622" phylum="Arthropoda" rank="species" species="myanmarensis">
|
||||
<emphasis box="[1033,1360,1622,1643]" italics="true" pageId="14" pageNumber="622">Mastotermes myanmarensis</emphasis>
|
||||
</taxonomicName>
|
||||
can be differentiated from
|
||||
<taxonomicName box="[1064,1248,1652,1674]" class="Insecta" family="Mastotermitidae" genus="Mastotermes" kingdom="Animalia" order="Blattodea" pageId="14" pageNumber="622" phylum="Arthropoda" rank="species" species="monostichus">
|
||||
<emphasis box="[1064,1248,1652,1674]" italics="true" pageId="14" pageNumber="622">M. monostichus</emphasis>
|
||||
</taxonomicName>
|
||||
by the forewing with RA simple, M with only a reduced number of branches and the first dichotomy far from the wing base, CuA with more dichotomous branches; hindwing with a broad space above Sc, and the anal field with more branches.
|
||||
<taxonomicName authorityName="Jouault & Engel & Fls & Legendre & Huang & Grandcolas & Nel" authorityYear="2022" box="[1121,1442,1806,1827]" class="Insecta" family="Mastotermitidae" genus="Mastotermes" kingdom="Animalia" order="Blattodea" pageId="14" pageNumber="622" phylum="Arthropoda" rank="species" species="myanmarensis">
|
||||
<emphasis box="[1121,1442,1806,1827]" italics="true" pageId="14" pageNumber="622">Mastotermes myanmarensis</emphasis>
|
||||
</taxonomicName>
|
||||
also differs from the extant
|
||||
<taxonomicName baseAuthorityName="Tillyard" baseAuthorityYear="1931" box="[1172,1366,1836,1858]" class="Insecta" family="Mastotermitidae" genus="Mastotermes" kingdom="Animalia" order="Blattodea" pageId="14" pageNumber="622" phylum="Arthropoda" rank="species" species="darwiniensis">
|
||||
<emphasis box="[1172,1366,1836,1858]" italics="true" pageId="14" pageNumber="622">M. darwiniensis</emphasis>
|
||||
</taxonomicName>
|
||||
by its temporal range (‘Mid’-Cretaceous vs. extant).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
408
data/91/1B/E6/911BE60E5548FFE08C34F9C4FB7683EB.xml
Normal file
408
data/91/1B/E6/911BE60E5548FFE08C34F9C4FB7683EB.xml
Normal file
|
|
@ -0,0 +1,408 @@
|
|||
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|
||||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||||
<mods:titleInfo>
|
||||
<mods:title>Incrementing and clarifying the diversity and early evolution of termites (Blattodea: Isoptera)</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Jouault, Corentin</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Engel, Michael S.</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Fls</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Legendre, Frédéric</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Huang, Diying</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Grandcolas, Philippe</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Nel, André</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:typeOfResource>text</mods:typeOfResource>
|
||||
<mods:relatedItem type="host">
|
||||
<mods:titleInfo>
|
||||
<mods:title>Zoological Journal of the Linnean Society</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:part>
|
||||
<mods:date>2022</mods:date>
|
||||
<mods:detail type="pubDate">
|
||||
<mods:number>2022-02-07</mods:number>
|
||||
</mods:detail>
|
||||
<mods:detail type="volume">
|
||||
<mods:number>196</mods:number>
|
||||
</mods:detail>
|
||||
<mods:extent unit="page">
|
||||
<mods:start>608</mods:start>
|
||||
<mods:end>629</mods:end>
|
||||
</mods:extent>
|
||||
</mods:part>
|
||||
</mods:relatedItem>
|
||||
<mods:classification>journal article</mods:classification>
|
||||
<mods:identifier type="DOI">10.1093/zoolinnean/zlac064</mods:identifier>
|
||||
<mods:identifier type="ISSN">0024-4082</mods:identifier>
|
||||
<mods:identifier type="Zenodo-Dep">7195105</mods:identifier>
|
||||
<mods:identifier type="ZooBank">01556715-E3DE-45E0-B620-ABD11BDB0748</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment LSID="urn:lsid:plazi:treatment:911BE60E5548FFE08C34F9C4FB7683EB" httpUri="http://treatment.plazi.org/id/911BE60E5548FFE08C34F9C4FB7683EB" lastPageId="5" lastPageNumber="612" pageId="2" pageNumber="610">
|
||||
<subSubSection pageId="2" pageNumber="610" type="nomenclature">
|
||||
<paragraph blockId="2.[887,1382,1560,1615]" pageId="2" pageNumber="610">
|
||||
<taxonomicName authority="JOUAULT, ENGEL," authorityName="JOUAULT, ENGEL, LEGENDRE & NEL" authorityYear="2022" class="Insecta" family="Mastotermitidae" genus="Longitermes" kingdom="Animalia" order="Blattodea" pageId="2" pageNumber="610" phylum="Arthropoda" rank="genus" status="GEN. NOV.">
|
||||
<emphasis bold="true" box="[887,1175,1560,1583]" italics="true" pageId="2" pageNumber="610">LONGITERMES PULCHER</emphasis>
|
||||
JOUAULT, ENGEL, LEGENDRE, & NEL
|
||||
</taxonomicName>
|
||||
<emphasis bold="true" box="[1195,1294,1591,1615]" pageId="2" pageNumber="610">SP. NOV.</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="2" pageNumber="610" type="description">
|
||||
<paragraph blockId="2.[1075,1194,1629,1654]" box="[1075,1194,1629,1654]" pageId="2" pageNumber="610">
|
||||
(
|
||||
<figureCitation box="[1084,1158,1629,1654]" captionStart="Figure 1" captionStartId="3.[145,223,891,913]" captionTargetBox="[304,1264,198,849]" captionTargetId="figure-507@3.[301,1268,195,852]" captionTargetPageId="3" captionText="Figure 1. Longitermes pulcher gen. et sp. nov., holotype IGR.BU-052. Photograph of habitus. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7195107" httpUri="https://zenodo.org/record/7195107/files/figure.png" pageId="2" pageNumber="610">FIGS 1</figureCitation>
|
||||
,
|
||||
<figureCitation box="[1172,1186,1629,1654]" captionStart="Figure 2" captionStartId="4.[164,242,1270,1292]" captionTargetBox="[166,1440,198,1226]" captionTargetId="figure-261@4.[163,1443,195,1230]" captionTargetPageId="4" captionText="Figure 2. Longitermes pulcher gen. et sp. nov., holotype IGR.BU-052. Detailed photographs.A, head; B, mandibles; C, apical part of left foreleg; D, mesotibia; E, metatibia; F, metatibia; G, forewing. la, left apical tooth; lc, lacinia; lci, lacinial incisor; LM1-LM3, left marginal teeth (number corresponding); Lp, labial palp; Mp, maxillary palp; pgl, paraglossa; CuA, anterior cubitus; M, media; PCu, postcubitus; RA, anterior radius; RP, posterior radius; Sc, subcosta. Scale bars: 0.5 mm (A, B, C, D, G); 0.25 mm (E, F)." figureDoi="http://doi.org/10.5281/zenodo.7195111" httpUri="https://zenodo.org/record/7195111/files/figure.png" pageId="2" pageNumber="610">2</figureCitation>
|
||||
)
|
||||
</paragraph>
|
||||
<paragraph blockId="2.[827,1443,1670,1753]" pageId="2" pageNumber="610">
|
||||
<emphasis box="[827,1179,1670,1691]" italics="true" pageId="2" pageNumber="610">
|
||||
<collectionCode box="[827,843,1670,1691]" pageId="2" pageNumber="610">Z</collectionCode>
|
||||
o o b a n k r e g i s t r a t i o n:
|
||||
</emphasis>
|
||||
h t t p: / / z o o b a n k. org/
|
||||
<uri pageId="2" pageNumber="610">
|
||||
urn:lsid:zoobank.org:act:
|
||||
<uuid pageId="2" pageNumber="610">6327861A-3757-4810- BEED-9800D20146C2</uuid>
|
||||
</uri>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection lastPageId="3" lastPageNumber="611" pageId="2" pageNumber="610" type="materials_examined">
|
||||
<paragraph blockId="2.[825,1444,1777,1891]" pageId="2" pageNumber="610">
|
||||
<emphasis box="[827,932,1778,1799]" italics="true" pageId="2" pageNumber="610">Material:</emphasis>
|
||||
<typeStatus box="[938,1042,1778,1800]" pageId="2" pageNumber="610">Holotype</typeStatus>
|
||||
, IGR.BU-052, preserved in an ovoid piece of amber measuring 18 × 12 ×
|
||||
<quantity box="[1250,1317,1808,1830]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="610" unit="mm" value="2.0">2 mm</quantity>
|
||||
, housed in the amber collection of the Geological Department and Museum (IGR), Rennes,
|
||||
<collectingCountry box="[1105,1187,1870,1891]" name="France" pageId="2" pageNumber="610">France</collectingCountry>
|
||||
.
|
||||
</paragraph>
|
||||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7195107" ID-Zenodo-Dep="7195107" box="[145,1298,891,914]" httpUri="https://zenodo.org/record/7195107/files/figure.png" pageId="3" pageNumber="611" startId="3.[145,223,891,913]" targetBox="[304,1264,198,849]" targetPageId="3">
|
||||
<paragraph blockId="3.[145,1298,891,914]" box="[145,1298,891,914]" pageId="3" pageNumber="611">
|
||||
<emphasis bold="true" box="[145,250,891,913]" pageId="3" pageNumber="611">Figure 1.</emphasis>
|
||||
<taxonomicName authority="Jouault & Engel & Fls & Legendre & Huang & Grandcolas & Nel, 2022" authorityName="Jouault & Engel & Fls & Legendre & Huang & Grandcolas & Nel" authorityYear="2022" box="[260,473,892,914]" class="Insecta" family="Mastotermitidae" genus="Longitermes" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="611" phylum="Arthropoda" rank="species" species="pulcher" status="gen. et sp. nov.">
|
||||
<emphasis box="[260,473,892,914]" italics="true" pageId="3" pageNumber="611">Longitermes pulcher</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel box="[479,633,892,914]" pageId="3" pageNumber="611" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
, holotype IGR.BU-052. Photograph of habitus. Scale bar: 1 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph blockId="3.[145,761,952,1035]" pageId="3" pageNumber="611">
|
||||
<emphasis box="[145,384,952,973]" italics="true" pageId="3" pageNumber="611">Locality and horizon:</emphasis>
|
||||
Noije Bum Hill, Hukawng Valley,
|
||||
<collectingRegion box="[145,313,982,1004]" country="Myanmar" name="Kachin" pageId="3" pageNumber="611">Kachin State</collectingRegion>
|
||||
,
|
||||
<collectingCountry box="[329,452,983,1004]" name="Myanmar" pageId="3" pageNumber="611">Myanmar</collectingCountry>
|
||||
; Upper Albian to Lower Cenomanian, ‘Mid’-Cretaceous.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="3" pageNumber="611" type="etymology">
|
||||
<paragraph blockId="3.[145,760,1073,1126]" pageId="3" pageNumber="611">
|
||||
<emphasis box="[145,280,1073,1094]" italics="true" pageId="3" pageNumber="611">Etymology:</emphasis>
|
||||
The specific epithet is from the Latin adjective
|
||||
<taxonomicName authorityName="Jouault & Engel & Fls & Legendre & Huang & Grandcolas & Nel" authorityYear="2022" box="[253,338,1104,1125]" class="Insecta" family="Mastotermitidae" genus="Longitermes" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="611" phylum="Arthropoda" rank="species" species="pulcher">
|
||||
<emphasis box="[253,338,1104,1125]" italics="true" pageId="3" pageNumber="611">pulcher</emphasis>
|
||||
</taxonomicName>
|
||||
, beautiful.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection box="[145,605,1150,1172]" pageId="3" pageNumber="611" type="diagnosis">
|
||||
<paragraph blockId="3.[145,605,1150,1172]" box="[145,605,1150,1172]" pageId="3" pageNumber="611">
|
||||
<emphasis box="[145,264,1151,1172]" italics="true" pageId="3" pageNumber="611">Diagnosis:</emphasis>
|
||||
As for the genus (
|
||||
<emphasis box="[470,589,1150,1172]" italics="true" pageId="3" pageNumber="611">vide supra</emphasis>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection lastPageId="4" lastPageNumber="612" pageId="3" pageNumber="611" type="description">
|
||||
<paragraph blockId="3.[143,762,1197,1894]" lastBlockId="3.[807,1426,952,1894]" pageId="3" pageNumber="611">
|
||||
<emphasis box="[145,287,1198,1219]" italics="true" pageId="3" pageNumber="611">Description:</emphasis>
|
||||
Imago specimen. Body
|
||||
<emphasis box="[573,590,1198,1219]" italics="true" pageId="3" pageNumber="611">c.</emphasis>
|
||||
<quantity box="[599,703,1197,1219]" metricMagnitude="-3" metricUnit="m" metricValue="7.34" pageId="3" pageNumber="611" unit="mm" value="7.34">7.34 mm</quantity>
|
||||
long (as preserved and measured from tip of labrum to abdominal apex). Head elongate,
|
||||
<emphasis box="[563,581,1259,1280]" italics="true" pageId="3" pageNumber="611">c.</emphasis>
|
||||
<quantity box="[592,699,1259,1280]" metricMagnitude="-3" metricUnit="m" metricValue="1.44" pageId="3" pageNumber="611" unit="mm" value="1.44">1.44 mm</quantity>
|
||||
long (measured from posterior margin to tip of labrum) and
|
||||
<emphasis box="[145,162,1320,1341]" italics="true" pageId="3" pageNumber="611">c.</emphasis>
|
||||
<quantity box="[169,270,1320,1341]" metricMagnitude="-4" metricUnit="m" metricValue="8.2" pageId="3" pageNumber="611" unit="mm" value="0.82">0.82 mm</quantity>
|
||||
wide excluding compound eyes (measured beyond eyes), trapezoidal in dorsal view, prognathous, sides slightly convex, postclypeus
|
||||
<emphasis box="[563,580,1382,1403]" italics="true" pageId="3" pageNumber="611">c.</emphasis>
|
||||
<quantity box="[590,697,1381,1403]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="611" unit="mm" value="0.11">0.11 mm</quantity>
|
||||
long, anteclypeus
|
||||
<emphasis box="[304,321,1412,1433]" italics="true" pageId="3" pageNumber="611">c.</emphasis>
|
||||
<quantity box="[333,440,1412,1433]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="611" unit="mm" value="0.11">0.11 mm</quantity>
|
||||
long, labrum lobe-shape, sides nearly parallel, slightly narrower basally, apical margin nearly straight, with few setae; mandibles triangular shaped,
|
||||
<emphasis box="[360,377,1504,1525]" italics="true" pageId="3" pageNumber="611">c.</emphasis>
|
||||
<quantity box="[382,478,1504,1525]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="611" unit="mm" value="0.2">0.20 mm</quantity>
|
||||
high, covered by labrum; left mandible apical tooth elongate, sharp; three marginal teeth, LM1 small, LM1-LM2 interdental space small, v-shaped, LM
|
||||
<subScript attach="left" box="[380,388,1611,1623]" fontSize="5" pageId="3" pageNumber="611">2</subScript>
|
||||
elongate, forming broad cutting edge, LM2-LM3 separated by a notch, LM3 bend toward base of mandible (general aspect similar to left imago mandible of
|
||||
<taxonomicName authorityName="Holmgren" authorityYear="1911" box="[284,453,1688,1709]" class="Insecta" family="Archotermopsidae" genus="Hodotermopsis" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="611" phylum="Arthropoda" rank="genus">
|
||||
<emphasis box="[284,453,1688,1709]" italics="true" pageId="3" pageNumber="611">Hodotermopsis</emphasis>
|
||||
</taxonomicName>
|
||||
); right mandible with long, sharp apical tooth, and at least one marginal tooth of similar shape and length; buccal pieces: glossa with two visible segments, paraglossa with three segments, length from base to apex (in mm) 0.05, 0.06, 0.1; lacinial incisor with two teeth; five maxillary palpomeres visible, length from base to apex (in mm): 0.06, 0.05, 0.08, 0.09, 0.12; three labial palpomeres present, total length as preserved
|
||||
<emphasis box="[1106,1123,983,1004]" italics="true" pageId="3" pageNumber="611">c.</emphasis>
|
||||
<quantity box="[1130,1233,983,1004]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="3" pageNumber="611" unit="mm" value="0.21">0.21 mm</quantity>
|
||||
; compound eyes,
|
||||
<emphasis box="[809,827,1014,1035]" italics="true" pageId="3" pageNumber="611">c.</emphasis>
|
||||
<quantity box="[835,939,1013,1035]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="3" pageNumber="611" unit="mm" value="0.37">0.37 mm</quantity>
|
||||
long, ovoid, situated laterally near head midlength, and separated from posterior head margin by more than their length; ocelli absent; fontanelle absent; antennae moniliform with 14 antennomeres. Pronotum
|
||||
<emphasis box="[930,947,1136,1157]" italics="true" pageId="3" pageNumber="611">c.</emphasis>
|
||||
<quantity box="[954,1054,1136,1158]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" pageId="3" pageNumber="611" unit="mm" value="0.54">0.54 mm</quantity>
|
||||
long and
|
||||
<emphasis box="[1168,1185,1136,1157]" italics="true" pageId="3" pageNumber="611">c.</emphasis>
|
||||
1.0 mm wide, nearly flat with anterior margin concave, anterior corners arched, sides broadly convex and posterior margin convex with a medial indentation, posterior corners broadly arched.
|
||||
</paragraph>
|
||||
<paragraph blockId="3.[807,1426,952,1894]" pageId="3" pageNumber="611">
|
||||
Legs robust; profemur medially swollen,
|
||||
<emphasis box="[1301,1318,1290,1311]" italics="true" pageId="3" pageNumber="611">c.</emphasis>
|
||||
<quantity box="[1325,1425,1289,1311]" metricMagnitude="-4" metricUnit="m" metricValue="6.3" pageId="3" pageNumber="611" unit="mm" value="0.63">0.63 mm</quantity>
|
||||
long, protibia
|
||||
<emphasis box="[975,992,1320,1341]" italics="true" pageId="3" pageNumber="611">c.</emphasis>
|
||||
<quantity box="[1000,1103,1320,1341]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="3" pageNumber="611" unit="mm" value="0.37">0.37 mm</quantity>
|
||||
long, protarsus
|
||||
<emphasis box="[1298,1316,1320,1341]" italics="true" pageId="3" pageNumber="611">c.</emphasis>
|
||||
<quantity box="[1323,1426,1320,1341]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="611" unit="mm" value="0.22">0.22 mm</quantity>
|
||||
long; protibia with three spurs f1, f2, f3; mesofemur medially swollen
|
||||
<emphasis box="[1004,1021,1382,1403]" italics="true" pageId="3" pageNumber="611">c.</emphasis>
|
||||
<quantity box="[1027,1123,1381,1403]" metricMagnitude="-4" metricUnit="m" metricValue="6.3" pageId="3" pageNumber="611" unit="mm" value="0.63">0.63 mm</quantity>
|
||||
long, mesotibia
|
||||
<emphasis box="[1304,1321,1382,1403]" italics="true" pageId="3" pageNumber="611">c.</emphasis>
|
||||
<quantity box="[1327,1424,1381,1403]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="3" pageNumber="611" unit="mm" value="0.42">0.42 mm</quantity>
|
||||
long, mesotarsus
|
||||
<emphasis box="[1027,1045,1412,1433]" italics="true" pageId="3" pageNumber="611">c.</emphasis>
|
||||
<quantity box="[1055,1162,1412,1433]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="611" unit="mm" value="0.22">0.22 mm</quantity>
|
||||
long; mesotibia with four observable spurs m1, m2, m3, m4; metafemur
|
||||
<emphasis box="[1408,1425,1443,1464]" italics="true" pageId="3" pageNumber="611">c.</emphasis>
|
||||
<quantity box="[809,915,1473,1495]" metricMagnitude="-4" metricUnit="m" metricValue="5.1" pageId="3" pageNumber="611" unit="mm" value="0.51">0.51 mm</quantity>
|
||||
long, metatibia
|
||||
<emphasis box="[1115,1133,1474,1495]" italics="true" pageId="3" pageNumber="611">c.</emphasis>
|
||||
<quantity box="[1140,1246,1473,1495]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="3" pageNumber="611" unit="mm" value="0.52">0.52 mm</quantity>
|
||||
long, with two observable spurs h1, h2, mesotarsus
|
||||
<emphasis box="[1237,1254,1504,1525]" italics="true" pageId="3" pageNumber="611">c.</emphasis>
|
||||
<quantity box="[1261,1362,1504,1526]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="611" unit="mm" value="0.25">0.25 mm</quantity>
|
||||
long; all tarsi pentamerous, with arolium. Forewing scale
|
||||
<emphasis box="[1408,1425,1535,1556]" italics="true" pageId="3" pageNumber="611">c.</emphasis>
|
||||
<quantity box="[809,909,1565,1587]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="611" unit="mm" value="0.8">0.80 mm</quantity>
|
||||
long and
|
||||
<quantity box="[1025,1125,1565,1587]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" pageId="3" pageNumber="611" unit="mm" value="0.54">0.54 mm</quantity>
|
||||
wide (measured distally), RP with additional tertiary branches joining costal margin; median (M) weak, closer to cubitus (Cu) than to radial sector (RP), with five branches terminating at lower margin; CuA with six primary and secondary branches extending to lower margin; anal vein absent. Hindwing (folded or damage): humeral suture weak, barely visible; wing venation apparently similar to forewing except anal vein present.
|
||||
</paragraph>
|
||||
<paragraph blockId="3.[807,1426,952,1894]" lastBlockId="4.[163,780,1447,1561]" lastPageId="4" lastPageNumber="612" pageId="3" pageNumber="611">
|
||||
Abdomen at least 4.0 mm long (damaged during the fossilization process) with at least ten observable segments; abdominal segments apparently slightly wider near midlength. Cerci multi-merous
|
||||
<emphasis box="[655,672,1479,1500]" italics="true" pageId="4" pageNumber="612">c.</emphasis>
|
||||
<quantity box="[679,779,1478,1500]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="4" pageNumber="612" unit="mm" value="0.16">0.16 mm</quantity>
|
||||
long, five(?) cercomeres (sometimes fused so not clearly countable).
|
||||
</paragraph>
|
||||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7195111" ID-Zenodo-Dep="7195111" httpUri="https://zenodo.org/record/7195111/files/figure.png" pageId="4" pageNumber="612" startId="4.[164,242,1270,1292]" targetBox="[166,1440,198,1226]" targetPageId="4">
|
||||
<paragraph blockId="4.[162,1443,1270,1409]" pageId="4" pageNumber="612">
|
||||
<emphasis bold="true" box="[164,267,1270,1292]" pageId="4" pageNumber="612">Figure 2.</emphasis>
|
||||
<taxonomicName authority="Jouault & Engel & Fls & Legendre & Huang & Grandcolas & Nel, 2022" authorityName="Jouault & Engel & Fls & Legendre & Huang & Grandcolas & Nel" authorityYear="2022" box="[276,487,1270,1292]" class="Insecta" family="Mastotermitidae" genus="Longitermes" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="612" phylum="Arthropoda" rank="species" species="pulcher" status="gen. et sp. nov.">
|
||||
<emphasis box="[276,487,1270,1292]" italics="true" pageId="4" pageNumber="612">Longitermes pulcher</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel box="[492,643,1270,1292]" pageId="4" pageNumber="612" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
, holotype IGR.BU-052. Detailed photographs. A, head; B, mandibles; C, apical part of left foreleg; D, mesotibia; E, metatibia; F, metatibia; G, forewing. la, left apical tooth; lc, lacinia; lci, lacinial incisor; LM1-LM3, left marginal teeth (number corresponding); Lp, labial palp; Mp, maxillary palp; pgl, paraglossa; CuA, anterior cubitus; M, media; PCu, postcubitus; RA, anterior radius; RP, posterior radius; Sc, subcosta. Scale bars: 0.5 mm (A, B, C, D, G); 0.25 mm (E, F).
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph blockId="4.[163,421,1602,1624]" box="[163,421,1602,1624]" pageId="4" pageNumber="612">
|
||||
<heading box="[163,421,1602,1624]" centered="true" fontSize="9" level="2" pageId="4" pageNumber="612" reason="2">
|
||||
<emphasis box="[163,246,1602,1623]" italics="true" pageId="4" pageNumber="612">Colour:</emphasis>
|
||||
Not preserved.
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection lastPageId="5" lastPageNumber="613" pageId="4" pageNumber="612" type="discussion">
|
||||
<paragraph blockId="4.[161,780,1662,1899]" lastBlockId="5.[145,762,197,1507]" lastPageId="5" lastPageNumber="613" pageId="4" pageNumber="612">
|
||||
<emphasis box="[163,552,1662,1684]" italics="true" pageId="4" pageNumber="612">
|
||||
Remarks:
|
||||
<taxonomicName authorityName="Jouault & Engel & Fls & Legendre & Huang & Grandcolas & Nel" authorityYear="2022" box="[292,552,1662,1684]" class="Insecta" family="Mastotermitidae" genus="Longitermes" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="612" phylum="Arthropoda" rank="species" species="pulcher">Longitermes pulcher</taxonomicName>
|
||||
</emphasis>
|
||||
is unique among Cretaceous termites, with an unprecedented combination of diagnostic characters making it close to the family
|
||||
<taxonomicName box="[324,540,1754,1776]" class="Insecta" family="Archotermopsidae" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="612" phylum="Arthropoda" rank="family">Archotermopsidae</taxonomicName>
|
||||
, as treated in
|
||||
<bibRefCitation author="Jiang RX & Zhang HR & Eldredge KT & Song XB & Li YY & Tihelka E & Huang DY & Wang S & Engel MS & Cai C" pageId="4" pageNumber="612" pagination="374 - 389" refId="ref13160" refString="Jiang RX, Zhang HR, Eldredge KT, Song XB, Li YY, Tihelka E, Huang DY, Wang S, Engel MS, Cai C. 2021. Further evidence of Cretaceous termitophily: description of new termite hosts of the trichopseniine Cretotrichopsenius (Coleoptera: Staphylinidae), with emendations to the classification of lower termites (Isoptera). Palaeoentomology 4: 374 - 389." type="journal article" year="2021">
|
||||
Jiang
|
||||
<emphasis box="[163,228,1785,1806]" italics="true" pageId="4" pageNumber="612">et al.</emphasis>
|
||||
(2021)
|
||||
</bibRefCitation>
|
||||
, i.e. encompassing
|
||||
<taxonomicName authorityName="Desneux" authorityYear="1904" box="[577,771,1785,1806]" class="Insecta" family="Archotermopsidae" genus="Archotermopsis" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="612" phylum="Arthropoda" rank="genus">
|
||||
<emphasis box="[577,771,1785,1806]" italics="true" pageId="4" pageNumber="612">Archotermopsis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName authorityName="Holmgren" authorityYear="1911" box="[163,335,1815,1836]" class="Insecta" family="Archotermopsidae" genus="Hodotermopsis" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="612" phylum="Arthropoda" rank="genus">
|
||||
<emphasis box="[163,335,1815,1836]" italics="true" pageId="4" pageNumber="612">Hodotermopsis</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName authorityName="Emerson" authorityYear="1933" box="[394,546,1816,1837]" class="Insecta" family="Archotermopsidae" genus="Zootermopsis" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="612" phylum="Arthropoda" rank="genus">
|
||||
<emphasis box="[394,546,1816,1837]" italics="true" pageId="4" pageNumber="612">Zootermopsis</emphasis>
|
||||
</taxonomicName>
|
||||
. Here we follow the new system of
|
||||
<bibRefCitation author="Wang M & Hellemans S & Sobotnik J & Arora J & Bucek A & Sillam-Dusses D & Clitheroe C & Lu T & Lo N & Engel MS & Roisin Y & Evans TA & Bourguignon T" box="[322,512,1846,1868]" pageId="4" pageNumber="612" pagination="12548" refId="ref14024" refString="Wang M, Hellemans S, Sobotnik J, Arora J, Bucek A, Sillam-Dusses D, Clitheroe C, Lu T, Lo N, Engel MS, Roisin Y, Evans TA, Bourguignon T. 2022. Phylogeny, biogeography, and classification of Teletisoptera (Blattaria: Isoptera). Systematic Entomology. In Press. Doi: 10.1111 / syen. 12548" type="book chapter" year="2022">
|
||||
Wang
|
||||
<emphasis box="[391,443,1846,1868]" italics="true" pageId="4" pageNumber="612">et al.</emphasis>
|
||||
(2022
|
||||
</bibRefCitation>
|
||||
: table 1), which recently resolved paraphyly of
|
||||
<taxonomicName box="[418,628,1877,1899]" class="Insecta" family="Archotermopsidae" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="612" phylum="Arthropoda" rank="family">Archotermopsidae</taxonomicName>
|
||||
by elevating Hodotermopsinae to family rank. In fact,
|
||||
<taxonomicName authorityName="Jouault & Engel & Fls & Legendre & Huang & Grandcolas & Nel" authorityYear="2022" box="[1325,1443,1447,1469]" class="Insecta" family="Mastotermitidae" genus="Longitermes" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="612" phylum="Arthropoda" rank="species" species="pulcher">
|
||||
<emphasis box="[1325,1443,1447,1469]" italics="true" pageId="4" pageNumber="612">L. pulcher</emphasis>
|
||||
</taxonomicName>
|
||||
shares with
|
||||
<taxonomicName box="[970,1181,1478,1500]" class="Insecta" family="Archotermopsidae" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="612" phylum="Arthropoda" rank="family">Archotermopsidae</taxonomicName>
|
||||
and
|
||||
<taxonomicName box="[1240,1443,1478,1500]" family="Hodotermopsidae" pageId="4" pageNumber="627" rank="family">Hodotermopsidae</taxonomicName>
|
||||
the lack of a fontanelle; a Y-shaped ecdysial line present; large compound eyes; ocelli absent; pronotum faintly arched, slightly wider than head; left mandible with an apical tooth and three marginal teeth. Legs: five tarsomeres (some cryptic); tibial spur formula 3-4-2; arolium present. Cerci with five(?) cercomeres. Forewing: costal margin flat to faintly arched; humeral suture well defined, almost straight; Sc, RA, RP sclerotized, RP with the main branch bending to apex of wing, five to six secondary branches and additional tertiary branches joining costal margin; M weak, with secondary branches terminating at lower margin; Cu with primary and secondary branches extending to lower margin; anal vein absent. The wing venation with the radial field encompassing most of the wing apex and the shape of the left mandibular teeth, nearly similar to that of
|
||||
<taxonomicName authorityName="Holmgren" authorityYear="1911" box="[442,617,289,310]" class="Insecta" family="Archotermopsidae" genus="Hodotermopsis" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="613" phylum="Arthropoda" rank="genus">
|
||||
<emphasis box="[442,617,289,310]" italics="true" pageId="5" pageNumber="613">Hodotermopsis</emphasis>
|
||||
</taxonomicName>
|
||||
, strengthen the taxonomic placement. However, the phylogenetic position of the new genus is hard to ascertain and it may be a stem representative of
|
||||
<taxonomicName box="[508,705,381,403]" family="Hodotermopsidae" pageId="5" pageNumber="627" rank="family">Hodotermopsidae</taxonomicName>
|
||||
or of
|
||||
<taxonomicName authority="(sensu Wang et al., 2022)" baseAuthorityName="Wang" baseAuthorityYear="2022" box="[145,643,412,434]" class="Insecta" family="Archotermopsidae" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="613" phylum="Arthropoda" rank="family">
|
||||
Archotermopsidae (
|
||||
<emphasis box="[368,431,412,433]" italics="true" pageId="5" pageNumber="613">sensu</emphasis>
|
||||
<bibRefCitation author="Wang M & Hellemans S & Sobotnik J & Arora J & Bucek A & Sillam-Dusses D & Clitheroe C & Lu T & Lo N & Engel MS & Roisin Y & Evans TA & Bourguignon T" box="[437,633,412,434]" pageId="5" pageNumber="613" pagination="12548" refId="ref14024" refString="Wang M, Hellemans S, Sobotnik J, Arora J, Bucek A, Sillam-Dusses D, Clitheroe C, Lu T, Lo N, Engel MS, Roisin Y, Evans TA, Bourguignon T. 2022. Phylogeny, biogeography, and classification of Teletisoptera (Blattaria: Isoptera). Systematic Entomology. In Press. Doi: 10.1111 / syen. 12548" type="book chapter" year="2022">
|
||||
Wang
|
||||
<emphasis box="[509,566,412,434]" italics="true" pageId="5" pageNumber="613">et al.</emphasis>
|
||||
, 2022
|
||||
</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
. Since the new specimen differs from the genera
|
||||
<taxonomicName authorityName="Desneux" authorityYear="1904" box="[585,761,442,463]" class="Insecta" family="Archotermopsidae" genus="Archotermopsis" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="613" phylum="Arthropoda" rank="genus">
|
||||
<emphasis box="[585,761,442,463]" italics="true" pageId="5" pageNumber="613">Archotermopsis</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName authorityName="Emerson" authorityYear="1933" box="[201,362,474,495]" class="Insecta" family="Archotermopsidae" genus="Zootermopsis" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="613" phylum="Arthropoda" rank="genus">
|
||||
<emphasis box="[201,362,474,495]" italics="true" pageId="5" pageNumber="613">Zootermopsis</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<taxonomicName authority="sensu Wang et al., 2022" authorityName="Wang" authorityYear="2022" class="Insecta" family="Archotermopsidae" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="613" phylum="Arthropoda" rank="family">
|
||||
Archotermopsidae
|
||||
<emphasis box="[616,683,474,495]" italics="true" pageId="5" pageNumber="613">sensu</emphasis>
|
||||
<bibRefCitation author="Wang M & Hellemans S & Sobotnik J & Arora J & Bucek A & Sillam-Dusses D & Clitheroe C & Lu T & Lo N & Engel MS & Roisin Y & Evans TA & Bourguignon T" pageId="5" pageNumber="613" pagination="12548" refId="ref14024" refString="Wang M, Hellemans S, Sobotnik J, Arora J, Bucek A, Sillam-Dusses D, Clitheroe C, Lu T, Lo N, Engel MS, Roisin Y, Evans TA, Bourguignon T. 2022. Phylogeny, biogeography, and classification of Teletisoptera (Blattaria: Isoptera). Systematic Entomology. In Press. Doi: 10.1111 / syen. 12548" type="book chapter" year="2022">
|
||||
Wang
|
||||
<emphasis box="[145,204,504,526]" italics="true" pageId="5" pageNumber="613">et al.</emphasis>
|
||||
, 2022
|
||||
</bibRefCitation>
|
||||
</taxonomicName>
|
||||
), at least based on its mandibular shape with LM2 elongate and forming a broad cutting edge (vs. sharp and not forming a cutting edge in the aforementioned genera) (
|
||||
<bibRefCitation author="Krishna K & Grimaldi DA & Krishna V & Engel MS" box="[450,691,596,618]" pageId="5" pageNumber="613" refId="ref13371" refString="Krishna K, Grimaldi DA, Krishna V, Engel MS. 2013. Treatise on the Isoptera of the world. Bulletin of the American Museum of Natural History 377: 19891 - 12433." type="journal volume" year="2013">
|
||||
Krishna
|
||||
<emphasis box="[557,618,596,618]" italics="true" pageId="5" pageNumber="613">et al.</emphasis>
|
||||
, 2013
|
||||
</bibRefCitation>
|
||||
), and resembles more the genus
|
||||
<taxonomicName authorityName="Holmgren" authorityYear="1911" box="[456,628,626,647]" class="Insecta" family="Archotermopsidae" genus="Hodotermopsis" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="613" phylum="Arthropoda" rank="genus">
|
||||
<emphasis box="[456,628,626,647]" italics="true" pageId="5" pageNumber="613">Hodotermopsis</emphasis>
|
||||
</taxonomicName>
|
||||
in its wing venation and mandibular configuration, we presume that
|
||||
<taxonomicName authorityName="JOUAULT, ENGEL" authorityYear="2022" box="[200,339,688,709]" class="Insecta" family="Mastotermitidae" genus="Longitermes" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="613" phylum="Arthropoda" rank="genus">
|
||||
<emphasis box="[200,339,688,709]" italics="true" pageId="5" pageNumber="613">Longitermes</emphasis>
|
||||
</taxonomicName>
|
||||
is probably a stem-Hodotermopsidae (
|
||||
<emphasis box="[153,217,719,740]" italics="true" pageId="5" pageNumber="613">sensu</emphasis>
|
||||
<bibRefCitation author="Wang M & Hellemans S & Sobotnik J & Arora J & Bucek A & Sillam-Dusses D & Clitheroe C & Lu T & Lo N & Engel MS & Roisin Y & Evans TA & Bourguignon T" box="[224,423,718,740]" pageId="5" pageNumber="613" pagination="12548" refId="ref14024" refString="Wang M, Hellemans S, Sobotnik J, Arora J, Bucek A, Sillam-Dusses D, Clitheroe C, Lu T, Lo N, Engel MS, Roisin Y, Evans TA, Bourguignon T. 2022. Phylogeny, biogeography, and classification of Teletisoptera (Blattaria: Isoptera). Systematic Entomology. In Press. Doi: 10.1111 / syen. 12548" type="book chapter" year="2022">
|
||||
Wang
|
||||
<emphasis box="[297,354,718,740]" italics="true" pageId="5" pageNumber="613">et al.</emphasis>
|
||||
, 2022
|
||||
</bibRefCitation>
|
||||
).
|
||||
<taxonomicName authorityName="Holmgren" authorityYear="1911" box="[446,617,718,739]" class="Insecta" family="Archotermopsidae" genus="Hodotermopsis" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="613" phylum="Arthropoda" rank="genus">
|
||||
<emphasis box="[446,617,718,739]" italics="true" pageId="5" pageNumber="613">Hodotermopsis</emphasis>
|
||||
</taxonomicName>
|
||||
differs from the new specimen in possessing small eyes, while they are large in
|
||||
<taxonomicName authorityName="JOUAULT, ENGEL" authorityYear="2022" box="[366,515,780,801]" class="Insecta" family="Mastotermitidae" genus="Longitermes" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="613" phylum="Arthropoda" rank="genus">
|
||||
<emphasis box="[366,515,780,801]" italics="true" pageId="5" pageNumber="613">Longitermes</emphasis>
|
||||
</taxonomicName>
|
||||
, but also in having numerous antennomeres (at least 23), while the new specimen only possesses 14 (the antenna might not be complete but the number of antennomeres seems to be lower than in
|
||||
<taxonomicName authorityName="Holmgren" authorityYear="1911" box="[385,562,902,923]" class="Insecta" family="Archotermopsidae" genus="Hodotermopsis" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="613" phylum="Arthropoda" rank="genus">
|
||||
<emphasis box="[385,562,902,923]" italics="true" pageId="5" pageNumber="613">Hodotermopsis</emphasis>
|
||||
</taxonomicName>
|
||||
) (
|
||||
<bibRefCitation author="Krishna K & Grimaldi DA & Krishna V & Engel MS" pageId="5" pageNumber="613" refId="ref13371" refString="Krishna K, Grimaldi DA, Krishna V, Engel MS. 2013. Treatise on the Isoptera of the world. Bulletin of the American Museum of Natural History 377: 19891 - 12433." type="journal volume" year="2013">
|
||||
Krishna
|
||||
<emphasis box="[694,755,902,924]" italics="true" pageId="5" pageNumber="613">et al.</emphasis>
|
||||
, 2013
|
||||
</bibRefCitation>
|
||||
). Recently, several families were erected based on Cretaceous specimens from
|
||||
<collectingRegion box="[529,615,964,986]" country="Myanmar" name="Kachin" pageId="5" pageNumber="613">Kachin</collectingRegion>
|
||||
amber (e.g. Melqartitermitidae and Mylacrotermitidae) but our specimen cannot be placed in any of them (
|
||||
<bibRefCitation author="Jiang RX & Zhang HR & Eldredge KT & Song XB & Li YY & Tihelka E & Huang DY & Wang S & Engel MS & Cai C" pageId="5" pageNumber="613" pagination="374 - 389" refId="ref13160" refString="Jiang RX, Zhang HR, Eldredge KT, Song XB, Li YY, Tihelka E, Huang DY, Wang S, Engel MS, Cai C. 2021. Further evidence of Cretaceous termitophily: description of new termite hosts of the trichopseniine Cretotrichopsenius (Coleoptera: Staphylinidae), with emendations to the classification of lower termites (Isoptera). Palaeoentomology 4: 374 - 389." type="journal article" year="2021">
|
||||
Jiang
|
||||
<emphasis box="[700,755,1025,1047]" italics="true" pageId="5" pageNumber="613">et al.</emphasis>
|
||||
, 2021
|
||||
</bibRefCitation>
|
||||
). Its broad radial field precludes affinities with the Melqartitermitidae and the presence of a welldefined forewing basal suture demarcating the scale precludes affinities with Mylacrotermitidae (
|
||||
<bibRefCitation author="Jiang RX & Zhang HR & Eldredge KT & Song XB & Li YY & Tihelka E & Huang DY & Wang S & Engel MS & Cai C" pageId="5" pageNumber="613" pagination="374 - 389" refId="ref13160" refString="Jiang RX, Zhang HR, Eldredge KT, Song XB, Li YY, Tihelka E, Huang DY, Wang S, Engel MS, Cai C. 2021. Further evidence of Cretaceous termitophily: description of new termite hosts of the trichopseniine Cretotrichopsenius (Coleoptera: Staphylinidae), with emendations to the classification of lower termites (Isoptera). Palaeoentomology 4: 374 - 389." type="journal article" year="2021">
|
||||
Jiang
|
||||
<emphasis box="[145,202,1178,1200]" italics="true" pageId="5" pageNumber="613">et al.</emphasis>
|
||||
, 2021
|
||||
</bibRefCitation>
|
||||
). Similarly, the new specimen differs from representatives of the family
|
||||
<taxonomicName box="[500,723,1209,1231]" family="Krishnatermitidae" pageId="5" pageNumber="613" rank="family">Krishnatermitidae</taxonomicName>
|
||||
, at least, because of its forewing with PCu straight (vs. arched in
|
||||
<taxonomicName box="[260,428,1270,1291]" class="Insecta" genus="Krishnatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="613" phylum="Arthropoda" rank="genus">
|
||||
<emphasis box="[260,428,1270,1291]" italics="true" pageId="5" pageNumber="613">Krishnatermes</emphasis>
|
||||
</taxonomicName>
|
||||
), RP encompassing the wing apex (variable in
|
||||
<taxonomicName box="[337,501,1301,1322]" class="Insecta" genus="Krishnatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="613" phylum="Arthropoda" rank="genus">
|
||||
<emphasis box="[337,501,1301,1322]" italics="true" pageId="5" pageNumber="613">Krishnatermes</emphasis>
|
||||
</taxonomicName>
|
||||
) and its hindwing with a defined scale (vs. absent in
|
||||
<taxonomicName box="[472,636,1332,1353]" class="Insecta" genus="Krishnatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="613" phylum="Arthropoda" rank="genus">
|
||||
<emphasis box="[472,636,1332,1353]" italics="true" pageId="5" pageNumber="613">Krishnatermes</emphasis>
|
||||
</taxonomicName>
|
||||
). However, the habitus of
|
||||
<taxonomicName authorityName="JOUAULT, ENGEL" authorityYear="2022" box="[314,455,1363,1384]" class="Insecta" family="Mastotermitidae" genus="Longitermes" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="613" phylum="Arthropoda" rank="genus">
|
||||
<emphasis box="[314,455,1363,1384]" italics="true" pageId="5" pageNumber="613">Longitermes</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName box="[515,684,1362,1383]" class="Insecta" genus="Krishnatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="613" phylum="Arthropoda" rank="genus">
|
||||
<emphasis box="[515,684,1362,1383]" italics="true" pageId="5" pageNumber="613">Krishnatermes</emphasis>
|
||||
</taxonomicName>
|
||||
are at first sight similar, which, if not symplesiomorphic, may indicate a proximity between the two genera and a hypothetical proximity of
|
||||
<taxonomicName box="[478,651,1454,1475]" class="Insecta" genus="Krishnatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="613" phylum="Arthropoda" rank="genus">
|
||||
<emphasis box="[478,651,1454,1475]" italics="true" pageId="5" pageNumber="613">Krishnatermes</emphasis>
|
||||
</taxonomicName>
|
||||
with the Teletisoptera.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
137
data/91/1B/E6/911BE60E5548FFE78CC5FEA1FEBB83C9.xml
Normal file
137
data/91/1B/E6/911BE60E5548FFE78CC5FEA1FEBB83C9.xml
Normal file
|
|
@ -0,0 +1,137 @@
|
|||
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
|
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<mods:title>Incrementing and clarifying the diversity and early evolution of termites (Blattodea: Isoptera)</mods:title>
|
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</mods:titleInfo>
|
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<mods:name type="personal">
|
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<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
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</mods:role>
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<mods:namePart>Jouault, Corentin</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
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||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Engel, Michael S.</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Fls</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Legendre, Frédéric</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Huang, Diying</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Grandcolas, Philippe</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Nel, André</mods:namePart>
|
||||
</mods:name>
|
||||
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<mods:title>Zoological Journal of the Linnean Society</mods:title>
|
||||
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|
||||
<mods:part>
|
||||
<mods:date>2022</mods:date>
|
||||
<mods:detail type="pubDate">
|
||||
<mods:number>2022-02-07</mods:number>
|
||||
</mods:detail>
|
||||
<mods:detail type="volume">
|
||||
<mods:number>196</mods:number>
|
||||
</mods:detail>
|
||||
<mods:extent unit="page">
|
||||
<mods:start>608</mods:start>
|
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|
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|
||||
</mods:relatedItem>
|
||||
<mods:classification>journal article</mods:classification>
|
||||
<mods:identifier type="DOI">10.1093/zoolinnean/zlac064</mods:identifier>
|
||||
<mods:identifier type="ISSN">0024-4082</mods:identifier>
|
||||
<mods:identifier type="Zenodo-Dep">7195105</mods:identifier>
|
||||
<mods:identifier type="ZooBank">01556715-E3DE-45E0-B620-ABD11BDB0748</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment LSID="urn:lsid:plazi:treatment:911BE60E5548FFE78CC5FEA1FEBB83C9" httpUri="http://treatment.plazi.org/id/911BE60E5548FFE78CC5FEA1FEBB83C9" lastPageNumber="610" pageId="2" pageNumber="610">
|
||||
<subSubSection pageId="2" pageNumber="610" type="nomenclature">
|
||||
<paragraph blockId="2.[902,1367,381,437]" pageId="2" pageNumber="610">
|
||||
<collectionCode box="[902,982,381,406]" pageId="2" pageNumber="610">GENUS</collectionCode>
|
||||
<taxonomicName authorityName="JOUAULT, ENGEL, LEGENDRE, NEL" authorityYear="2022" class="Insecta" family="Mastotermitidae" genus="Longitermes" kingdom="Animalia" order="Blattodea" pageId="2" pageNumber="610" phylum="Arthropoda" rank="genus" status="GEN. NOV.">
|
||||
<emphasis bold="true" box="[989,1160,382,405]" italics="true" pageId="2" pageNumber="610">LONGITERMES</emphasis>
|
||||
JOUAULT, ENGEL,
|
||||
<collectionCode box="[968,1092,412,437]" pageId="2" pageNumber="610">LEGENDRE</collectionCode>
|
||||
& NEL
|
||||
</taxonomicName>
|
||||
<emphasis bold="true" box="[1182,1301,412,436]" pageId="2" pageNumber="610">
|
||||
<taxonomicNameLabel box="[1182,1301,412,436]" pageId="2" pageNumber="610">GEN. NOV.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="2" pageNumber="610" type="description">
|
||||
<paragraph blockId="2.[827,1443,452,536]" pageId="2" pageNumber="610">
|
||||
<emphasis box="[827,1179,452,474]" italics="true" pageId="2" pageNumber="610">
|
||||
<collectionCode box="[827,843,453,474]" pageId="2" pageNumber="610">Z</collectionCode>
|
||||
o o b a n k r e g i s t r a t i o n:
|
||||
</emphasis>
|
||||
h t t p: / / z o o b a n k. org/
|
||||
<uri pageId="2" pageNumber="610">
|
||||
urn:lsid:zoobank.org:act:
|
||||
<uuid pageId="2" pageNumber="610">AB77601A-A6D5-4CB9- 9236-DEC7FF2144B7</uuid>
|
||||
</uri>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="2" pageNumber="610" type="type_taxon">
|
||||
<paragraph blockId="2.[827,1441,560,613]" pageId="2" pageNumber="610">
|
||||
<emphasis box="[827,984,561,582]" italics="true" pageId="2" pageNumber="610">
|
||||
<typeStatus box="[827,883,561,582]" pageId="2" pageNumber="610">Type</typeStatus>
|
||||
species:
|
||||
</emphasis>
|
||||
<taxonomicName authority="Jouault, Engel, Legendre & Nel" authorityName="Jouault, Engel, Legendre & Nel" authorityYear="2022" class="Insecta" family="Mastotermitidae" genus="Longitermes" kingdom="Animalia" order="Blattodea" pageId="2" pageNumber="610" phylum="Arthropoda" rank="species" species="pulcher" status="sp. nov.">
|
||||
<emphasis box="[994,1245,560,582]" italics="true" pageId="2" pageNumber="610">Longitermes pulcher</emphasis>
|
||||
Jouault, Engel, Legendre & Nel
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel box="[1014,1098,591,612]" pageId="2" pageNumber="610" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="2" pageNumber="610" type="etymology">
|
||||
<paragraph blockId="2.[827,1443,638,752]" pageId="2" pageNumber="610">
|
||||
<emphasis box="[827,955,638,659]" italics="true" pageId="2" pageNumber="610">Etymology:</emphasis>
|
||||
The generic name is a combination of the Latin
|
||||
<emphasis box="[900,977,668,689]" italics="true" pageId="2" pageNumber="610">longus</emphasis>
|
||||
, long, referring to the elongate head of the new species, and
|
||||
<emphasis box="[1074,1150,700,721]" italics="true" pageId="2" pageNumber="610">termes</emphasis>
|
||||
, termite, commonly used to refer to
|
||||
<taxonomicName authorityName="BRULLE" authorityYear="1832" box="[946,1043,730,751]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Isoptera" pageId="2" pageNumber="610" phylum="Tracheophyta" rank="order">Isoptera</taxonomicName>
|
||||
. Gender masculine.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="2" pageNumber="610" type="diagnosis">
|
||||
<paragraph blockId="2.[827,1444,776,1473]" pageId="2" pageNumber="610">
|
||||
<emphasis box="[827,943,777,798]" italics="true" pageId="2" pageNumber="610">Diagnosis:</emphasis>
|
||||
Imago. Head elongate, with anterior corners rounded orthogonal and posterior corners broadly rounded; occiput and frons convex; epicranial ecdysial line visible; ocelli absent; postclypeus trapezoidal; anteclypeus small; left mandible apical tooth elongate, sharp, three marginal teeth, LM
|
||||
<subScript attach="right" box="[1069,1077,945,957]" fontSize="5" pageId="2" pageNumber="610">1</subScript>
|
||||
-LM
|
||||
<subScript attach="left" box="[1127,1135,945,957]" fontSize="5" pageId="2" pageNumber="610">2</subScript>
|
||||
interdental space small, v-shaped, LM1 small, LM2 elongate, forming broad cutting edge, LM2-LM3 separated by a notch, LM3 bent toward base of mandible; right mandible with long, sharp apical tooth, and at least one marginal tooth of similar shape and length; pronotum slightly wider than head, anterior margin concave, posterior margin broadly convex with a median concavity, sides broadly convex; tarsi pentamerous (fourth tarsomere cryptic); tibial spur formula 3-4-2; all tibiae with strong setae but no additional spines; arolium present; forewing with scale large, with humeral margin nearly straight, claval suture straight (PCu), basal suture faintly arched; Sc, RA, RP heavily sclerotized; R branched within scale; RP nearly encompassing wing apex, with main branch bending to wing apex, with five secondary branches; M and CuA weaker than other surrounding veins; CuA with three branches in scales; anal veins absent; hindwing with delineated scale.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
118
data/91/1B/E6/911BE60E554FFFE08F98F957F9BF85F0.xml
Normal file
118
data/91/1B/E6/911BE60E554FFFE08F98F957F9BF85F0.xml
Normal file
|
|
@ -0,0 +1,118 @@
|
|||
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Incrementing and clarifying the diversity and early evolution of termites (Blattodea: Isoptera)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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||||
<mods:namePart>Jouault, Corentin</mods:namePart>
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</mods:name>
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||||
<mods:name type="personal">
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<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Engel, Michael S.</mods:namePart>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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||||
</mods:role>
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<mods:namePart>Fls</mods:namePart>
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||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Legendre, Frédéric</mods:namePart>
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||||
</mods:name>
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||||
<mods:name type="personal">
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||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
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||||
</mods:role>
|
||||
<mods:namePart>Huang, Diying</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Grandcolas, Philippe</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Nel, André</mods:namePart>
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||||
</mods:name>
|
||||
<mods:typeOfResource>text</mods:typeOfResource>
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||||
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|
||||
<mods:part>
|
||||
<mods:date>2022</mods:date>
|
||||
<mods:detail type="pubDate">
|
||||
<mods:number>2022-02-07</mods:number>
|
||||
</mods:detail>
|
||||
<mods:detail type="volume">
|
||||
<mods:number>196</mods:number>
|
||||
</mods:detail>
|
||||
<mods:extent unit="page">
|
||||
<mods:start>608</mods:start>
|
||||
<mods:end>629</mods:end>
|
||||
</mods:extent>
|
||||
</mods:part>
|
||||
</mods:relatedItem>
|
||||
<mods:classification>journal article</mods:classification>
|
||||
<mods:identifier type="DOI">10.1093/zoolinnean/zlac064</mods:identifier>
|
||||
<mods:identifier type="ISSN">0024-4082</mods:identifier>
|
||||
<mods:identifier type="ZooBank">01556715-E3DE-45E0-B620-ABD11BDB0748</mods:identifier>
|
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</mods:mods>
|
||||
<treatment LSID="urn:lsid:plazi:treatment:911BE60E554FFFE08F98F957F9BF85F0" httpUri="http://treatment.plazi.org/id/911BE60E554FFFE08F98F957F9BF85F0" lastPageNumber="613" pageId="5" pageNumber="613">
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<subSubSection box="[219,686,1675,1700]" pageId="5" pageNumber="613" type="nomenclature">
|
||||
<paragraph blockId="5.[219,686,1675,1700]" box="[219,686,1675,1700]" pageId="5" pageNumber="613">
|
||||
<heading box="[219,686,1675,1700]" centered="true" fontSize="9" level="2" pageId="5" pageNumber="613" reason="2">
|
||||
GENUS
|
||||
<taxonomicName authority="ZHAO" authorityName="ZHAO" box="[306,539,1675,1700]" class="Insecta" family="Mastotermitidae" genus="Anisotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="613" phylum="Arthropoda" rank="genus">
|
||||
<emphasis box="[306,469,1676,1699]" italics="true" pageId="5" pageNumber="613">ANISOTERMES</emphasis>
|
||||
ZHAO
|
||||
</taxonomicName>
|
||||
<emphasis box="[546,610,1679,1699]" italics="true" pageId="5" pageNumber="613">ET AL</emphasis>
|
||||
., 2019
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="5" pageNumber="613" type="discussion">
|
||||
<paragraph blockId="5.[145,761,1716,1799]" pageId="5" pageNumber="613">
|
||||
<emphasis box="[145,343,1716,1737]" italics="true" pageId="5" pageNumber="613">Included species:</emphasis>
|
||||
<taxonomicName authority="Zhao, Eggleton & Ren 2019" authorityName="Zhao, Eggleton & Ren" authorityYear="2019" class="Insecta" family="Mastotermitidae" genus="Anisotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="613" phylum="Arthropoda" rank="species" species="xiai">
|
||||
<emphasis box="[351,547,1716,1737]" italics="true" pageId="5" pageNumber="613">Anisotermes xiai</emphasis>
|
||||
<bibRefCitation author="Zhao Z & Eggleton P & Yin X & Gao T & Shih C & Ren D" pageId="5" pageNumber="613" pagination="612 - 623" refId="ref14180" refString="Zhao Z, Eggleton P, Yin X, Gao T, Shih C, Ren D. 2019. The oldest known mastotermitids (Blattodea: Termitoidae) and phylogeny of basal termites. Systematic Entomology 44: 612 - 623." type="journal article" year="2019">Zhao, Eggleton & Ren 2019</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(
|
||||
<typeStatus box="[268,318,1747,1768]" pageId="5" pageNumber="613">type</typeStatus>
|
||||
species) and
|
||||
<taxonomicName authority="Jouault, Engel, Huang & Nel" authorityName="Jouault, Engel, Huang & Nel" authorityYear="2022" class="Insecta" family="Mastotermitidae" genus="Anisotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="613" phylum="Arthropoda" rank="species" species="bourguignoni" status="sp. nov.">
|
||||
<emphasis box="[475,659,1746,1768]" italics="true" pageId="5" pageNumber="613">A. bourguignoni</emphasis>
|
||||
Jouault, Engel, Huang & Nel
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel box="[383,465,1778,1799]" pageId="5" pageNumber="613" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="5" pageNumber="613" type="diagnosis">
|
||||
<paragraph blockId="5.[145,760,1824,1907]" lastBlockId="5.[809,1425,197,1016]" pageId="5" pageNumber="613">
|
||||
<emphasis box="[145,389,1824,1845]" italics="true" pageId="5" pageNumber="613">Emended diagnosis:</emphasis>
|
||||
Head approximately rounded; ocelli absent; fontanelle absent; Y-shaped ecdysial line absent; mandible not exceeding labrum, with la and ra conspicuously longer than LM1 and RM1; antenna moniliform, with about 25 antennomeres, terminal eight antennomeres tapered; compound eyes hemispheric, lying at head midlength; pronotum definitely saddle-shaped, slightly wider than head in dorsal view; lateral margin weakly upcurved; procoxal carina present; additional tibial spines present on all legs; tibial spur formula 3-4-4; tarsi wholly pentamerous; arolium variable (absent in
|
||||
<taxonomicName authorityName="Zhao, Eggleton & Ren" authorityYear="2019" box="[1303,1378,443,464]" class="Insecta" family="Mastotermitidae" genus="Anisotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="613" phylum="Arthropoda" rank="species" species="xiai">
|
||||
<emphasis box="[1303,1378,443,464]" italics="true" pageId="5" pageNumber="613">A. xiai</emphasis>
|
||||
</taxonomicName>
|
||||
but present in
|
||||
<taxonomicName authorityName="Jouault, Engel, Huang & Nel" authorityYear="2022" box="[947,1149,473,494]" class="Insecta" family="Mastotermitidae" genus="Anisotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="613" phylum="Arthropoda" rank="species" species="bourguignoni">
|
||||
<emphasis box="[947,1149,473,494]" italics="true" pageId="5" pageNumber="613">A. bourguignoni</emphasis>
|
||||
</taxonomicName>
|
||||
); wings membranous, densely reticulate; forewing scale large, overlapping hindwing base, basal suture convex, veins Sc, RA, RP and M more heavily pigmented than CuA. Forewing: all major veins originate within scale; Sc simple or double; RP with five to six main branches, terminating on costal margin anterior to wing apex; radial field of moderate width, parallel to costal margin; M running about midway between RP and CuA, terminating on posterior margin; medial field relatively narrow; CuA branched, terminating about same distance with Rs to wing apex; PCu (claval suture) arched, meeting basal suture on posterior margin. Hindwing without basal suture, large anal lobe present; M first inferior branch fading away, terminating in center of hindwing; cerci short, trimerous or tetramerous (with additional indented rings); abdominal styli absent in female but present in male.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
360
data/91/1B/E6/911BE60E554FFFEC8C02FB96FEC68424.xml
Normal file
360
data/91/1B/E6/911BE60E554FFFEC8C02FB96FEC68424.xml
Normal file
|
|
@ -0,0 +1,360 @@
|
|||
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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|
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|
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|
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|
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|
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|
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|
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|
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</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
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|
||||
</mods:role>
|
||||
<mods:namePart>Fls</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Legendre, Frédéric</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
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|
||||
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|
||||
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|
||||
<mods:namePart>Huang, Diying</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Grandcolas, Philippe</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
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||||
<mods:namePart>Nel, André</mods:namePart>
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|
||||
<mods:part>
|
||||
<mods:date>2022</mods:date>
|
||||
<mods:detail type="pubDate">
|
||||
<mods:number>2022-02-07</mods:number>
|
||||
</mods:detail>
|
||||
<mods:detail type="volume">
|
||||
<mods:number>196</mods:number>
|
||||
</mods:detail>
|
||||
<mods:extent unit="page">
|
||||
<mods:start>608</mods:start>
|
||||
<mods:end>629</mods:end>
|
||||
</mods:extent>
|
||||
</mods:part>
|
||||
</mods:relatedItem>
|
||||
<mods:classification>journal article</mods:classification>
|
||||
<mods:identifier type="DOI">10.1093/zoolinnean/zlac064</mods:identifier>
|
||||
<mods:identifier type="ISSN">0024-4082</mods:identifier>
|
||||
<mods:identifier type="Zenodo-Dep">7195105</mods:identifier>
|
||||
<mods:identifier type="ZooBank">01556715-E3DE-45E0-B620-ABD11BDB0748</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment LSID="urn:lsid:plazi:treatment:911BE60E554FFFEC8C02FB96FEC68424" httpUri="http://treatment.plazi.org/id/911BE60E554FFFEC8C02FB96FEC68424" lastPageId="9" lastPageNumber="617" pageId="5" pageNumber="613">
|
||||
<subSubSection pageId="5" pageNumber="613" type="nomenclature">
|
||||
<paragraph blockId="5.[833,1401,1098,1154]" pageId="5" pageNumber="613">
|
||||
<taxonomicName authorityName="Jouault, Engel, Huang & Nel" authorityYear="2022" box="[833,1194,1098,1122]" class="Insecta" family="Mastotermitidae" genus="Anisotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="617" phylum="Arthropoda" rank="species" species="bourguignoni">
|
||||
<emphasis bold="true" box="[833,1194,1098,1122]" italics="true" pageId="5" pageNumber="613">ANISOTERMES BOURGUIGNONI</emphasis>
|
||||
</taxonomicName>
|
||||
JOUAULT, ENGEL,
|
||||
<heading box="[980,1253,1129,1154]" centered="true" fontSize="9" level="2" pageId="5" pageNumber="613" reason="3">
|
||||
HUANG & NEL
|
||||
<emphasis bold="true" box="[1155,1253,1129,1153]" pageId="5" pageNumber="613">SP. NOV.</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="5" pageNumber="613" type="description">
|
||||
<paragraph blockId="5.[1057,1176,1167,1192]" box="[1057,1176,1167,1192]" pageId="5" pageNumber="613">
|
||||
(
|
||||
<figureCitation box="[1065,1168,1167,1192]" captionStart-0="Figure 3" captionStart-1="Figure 4" captionStart-2="Figure 5" captionStartId-0="6.[163,243,1214,1236]" captionStartId-1="7.[145,221,1760,1782]" captionStartId-2="8.[163,243,1711,1733]" captionTargetBox-0="[322,1282,198,1172]" captionTargetBox-1="[199,1370,198,1744]" captionTargetBox-2="[166,1440,198,1668]" captionTargetId-0="figure-306@6.[319,1285,195,1175]" captionTargetId-1="figure-27@7.[196,1373,195,1747]" captionTargetId-2="figure-27@8.[163,1443,195,1671]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="Figure 3. Anisotermes bourguignoni sp. nov., holotype NIGP177751. Photographs of habitus. A, dorsal view; B, ventral view. Scale bars: 1 mm." captionText-1="Figure 4. Anisotermes bourguignoni sp. nov., holotype NIGP177751. Detailed photographs. A, head and pronoun in dorsal view; B, head in ventral view; C, high-resolution X-ray microtomography reconstruction of head in ventral view; D, high-resolution X-ray microtomographyreconstruction ofhead inanterolateral view;E, line drawing ofmandibles;F,line drawingof Mastotermesdarwinensis mandibles.gl,glossa;la,leftapical tooth;lb,labrum;lc,lacinia;lci,lacinialincisor;LM1-LM3,leftmarginalteeth(numbercorresponding); Lp, labial palp; mb, mandibles; Mp, maxillary palp; mt, mentum; pc, postclypeus;pgl, paraglossa; ra, right apical tooth; RM1-RM2, right" captionText-2="Figure 5. Anisotermes bourguignoni sp. nov., holotype NIGP177751. Detailed photographs. A, protarus; B, mesotarsus; C, apex of metatibia; D, line drawing of scale venation with names of veins labelled; E, scale; F, cerci in ventral view with arrows pointing to cercomeres; G, cerci in dorsal view; H, interpretative drawing of protibia; I, interpretative drawing of mesotibiae; J, interpretative drawing of metatibiae. CuA, anterior cubitus; MA, media; PCu, postcubitus; RA, anterior radius; RP, posterior radius; Sc, subcosta; for the nomenclature of tibial spines and spurs refers to Krishna et al., 2013. Scale bars: 0.2 mm (A, F, G); 0.5 mm (B, C, D, E, G, H, I, J)." figureDoi-0="http://doi.org/10.5281/zenodo.7195119" figureDoi-1="http://doi.org/10.5281/zenodo.7195127" figureDoi-2="http://doi.org/10.5281/zenodo.7195129" httpUri-0="https://zenodo.org/record/7195119/files/figure.png" httpUri-1="https://zenodo.org/record/7195127/files/figure.png" httpUri-2="https://zenodo.org/record/7195129/files/figure.png" pageId="5" pageNumber="613">FIGS 3–5</figureCitation>
|
||||
)
|
||||
</paragraph>
|
||||
<paragraph blockId="5.[809,1425,1208,1291]" pageId="5" pageNumber="613">
|
||||
<emphasis box="[809,1161,1208,1230]" italics="true" pageId="5" pageNumber="613">
|
||||
<collectionCode box="[809,825,1209,1230]" pageId="5" pageNumber="613">Z</collectionCode>
|
||||
o o b a n k r e g i s t r a t i o n:
|
||||
</emphasis>
|
||||
h t t p: / / z o o b a n k. org/
|
||||
<uri pageId="5" pageNumber="613">
|
||||
urn:lsid:zoobank.org:act:
|
||||
<uuid pageId="5" pageNumber="613">A406BDD6-D799-4DB8- 93D0-4C2E61C71F1A</uuid>
|
||||
</uri>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="5" pageNumber="613" type="materials_examined">
|
||||
<paragraph blockId="5.[809,1425,1316,1460]" pageId="5" pageNumber="613">
|
||||
<emphasis box="[809,915,1316,1337]" italics="true" pageId="5" pageNumber="613">
|
||||
<typeStatus box="[809,909,1316,1337]" pageId="5" pageNumber="613">Holotype</typeStatus>
|
||||
:
|
||||
</emphasis>
|
||||
Specimen identifier NIGP177751 (a complete dealate specimen preserved in a rounded amber piece), housed in the collection of the Nanjing Institute of Geology and Palaeontology Chinese Academy of Sciences (NIGPAS), Nanjing,
|
||||
<collectingCountry box="[1143,1216,1438,1460]" name="China" pageId="5" pageNumber="613">China</collectingCountry>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="5" pageNumber="613" type="etymology">
|
||||
<paragraph blockId="5.[809,1425,1485,1568]" pageId="5" pageNumber="613">
|
||||
<emphasis box="[809,939,1485,1506]" italics="true" pageId="5" pageNumber="613">Etymology:</emphasis>
|
||||
The specific epithet honours Dr Thomas Bourguignon for his work on termite evolution. It is to be treated as a noun in the genitive case.
|
||||
</paragraph>
|
||||
<paragraph blockId="5.[809,1424,1593,1676]" pageId="5" pageNumber="613">
|
||||
<emphasis box="[809,1016,1593,1615]" italics="true" pageId="5" pageNumber="613">Locality and age:</emphasis>
|
||||
Noije Bum Hill, Hukawng Valley,
|
||||
<collectingRegion box="[809,983,1624,1646]" country="Myanmar" name="Kachin" pageId="5" pageNumber="613">Kachin State</collectingRegion>
|
||||
,
|
||||
<collectingCountry box="[1002,1128,1624,1645]" name="Myanmar" pageId="5" pageNumber="613">Myanmar</collectingCountry>
|
||||
; Late Albian to Early Cenomanian, ‘Mid’-Cretaceous.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection lastPageId="6" lastPageNumber="614" pageId="5" pageNumber="613" type="diagnosis">
|
||||
<paragraph blockId="5.[809,1425,1701,1907]" lastBlockId="6.[163,779,1317,1769]" lastPageId="6" lastPageNumber="614" pageId="5" pageNumber="613">
|
||||
<emphasis box="[809,942,1702,1723]" italics="true" pageId="5" pageNumber="613">Diagnosis:</emphasis>
|
||||
Imago. Head rounded, with anterior corners slightly orthogonal and posterior corners rounded; left mandible with la longer than LM1, both sharp; right mandible with an apical and two marginal teeth, ra shorter than RM
|
||||
<subScript attach="left" box="[1100,1108,1839,1851]" fontSize="5" pageId="5" pageNumber="613">1</subScript>
|
||||
, RM
|
||||
<subScript attach="left" box="[1161,1169,1839,1851]" fontSize="5" pageId="5" pageNumber="613">1</subScript>
|
||||
and RM
|
||||
<subScript attach="left" box="[1265,1273,1839,1851]" fontSize="5" pageId="5" pageNumber="613">2</subScript>
|
||||
separated by a sharp angle; occiput and frons convex; postclypeus trapezoidal; anteclypeus small; ocelli absent; pronotum massive, trapezoidal, wider than head, anterior margin concave surrounding head, posterior margin broadly convex, with dorsal surface flatly arched in profile, sides convex; tarsi pentamerous; tibial spur formula 3-4-4; protibia with two additional spine (fa, fb); mesotibia with six additional spines (ma-mf); metatibia with at least three additional spines (ha-hc); arolium present; forewing with scale large, with humeral margin welldefined and slightly arched, claval suture strongly curved (PCu), basal suture convex; Sc, RA, RP and CuA heavily sclerotized; Sc branched within scale and going outside of scale; R branched within scale; M weaker than other surrounding veins; CuA with three branches in scales; PCu strongly curved; anal veins not discernible within scale.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7195119" ID-Zenodo-Dep="7195119" httpUri="https://zenodo.org/record/7195119/files/figure.png" pageId="6" pageNumber="614" startId="6.[163,243,1214,1236]" targetBox="[322,1282,198,1172]" targetPageId="6">
|
||||
<paragraph blockId="6.[163,1442,1214,1266]" pageId="6" pageNumber="614">
|
||||
<emphasis bold="true" box="[163,270,1214,1236]" pageId="6" pageNumber="614">Figure 3.</emphasis>
|
||||
<taxonomicName authority="Jouault & Engel & Fls & Legendre & Huang & Grandcolas & Nel, 2022" authorityName="Jouault & Engel & Fls & Legendre & Huang & Grandcolas & Nel" authorityYear="2022" box="[282,562,1214,1236]" class="Insecta" family="Mastotermitidae" genus="Anisotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="614" phylum="Arthropoda" rank="species" species="bourguignoni" status="sp. nov.">
|
||||
<emphasis box="[282,562,1214,1236]" italics="true" pageId="6" pageNumber="614">Anisotermes bourguignoni</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel box="[570,651,1214,1236]" pageId="6" pageNumber="614" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
, holotype NIGP177751. Photographs of habitus. A, dorsal view; B, ventral view. Scale bars: 1 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<subSubSection lastPageId="9" lastPageNumber="617" pageId="6" pageNumber="614" type="description">
|
||||
<paragraph blockId="6.[163,781,1808,1891]" lastBlockId="9.[141,763,197,863]" lastPageId="9" lastPageNumber="617" pageId="6" pageNumber="614">
|
||||
<emphasis box="[163,305,1809,1830]" italics="true" pageId="6" pageNumber="614">Description:</emphasis>
|
||||
Imago. Body
|
||||
<emphasis box="[470,487,1809,1830]" italics="true" pageId="6" pageNumber="614">c.</emphasis>
|
||||
<quantity box="[495,577,1808,1829]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="614" unit="mm" value="10.0">10 mm</quantity>
|
||||
long (from tip of labrum to abdomen apex). Head robust,
|
||||
<emphasis box="[649,666,1839,1860]" italics="true" pageId="6" pageNumber="614">c.</emphasis>
|
||||
<quantity box="[675,780,1839,1860]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="6" pageNumber="614" unit="mm" value="2.2">2.20 mm</quantity>
|
||||
long (measured from under the pronotum to the tip of the labrum) and
|
||||
<emphasis box="[1019,1036,1318,1339]" italics="true" pageId="6" pageNumber="614">c.</emphasis>
|
||||
<quantity box="[1043,1143,1317,1339]" metricMagnitude="-3" metricUnit="m" metricValue="1.93" pageId="6" pageNumber="614" unit="mm" value="1.93">1.93 mm</quantity>
|
||||
wide excluding compound eyes, square-shaped in dorsal view, hypognathous, sides slightly convex, anterolateral corner slightly orthogonal, posterolateral corners rounded, Y-shaped ecdysial scar absent; tentorial pit visible (ttp); postclypeus (pc)
|
||||
<quantity box="[1019,1120,1471,1492]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="6" pageNumber="614" unit="mm" value="0.32">0.32 mm</quantity>
|
||||
long, trapezoidal; labrum
|
||||
<emphasis box="[1426,1444,1471,1492]" italics="true" pageId="6" pageNumber="614">c.</emphasis>
|
||||
<quantity box="[827,923,1501,1523]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="614" unit="mm" value="0.5">0.50 mm</quantity>
|
||||
long, lobe-shape, narrower basally; mandibles massive, triangular shaped, at least
|
||||
<quantity box="[1269,1375,1532,1554]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="6" pageNumber="614" unit="mm" value="0.53">0.53 mm</quantity>
|
||||
high; buccal pieces with glossa (gl), paraglossa (gl), stipe (st), mentum (mt), submentum (sm) visible but not clearly describable; five maxillary palpomeres, combined length as preserved
|
||||
<emphasis box="[1219,1237,1655,1676]" italics="true" pageId="6" pageNumber="614">c.</emphasis>
|
||||
<quantity box="[1248,1345,1655,1676]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="614" unit="mm" value="1.2">1.2 mm</quantity>
|
||||
; four(?) labial palpomeres present, with three discernible palpomeres, combined length
|
||||
<emphasis box="[1180,1198,1717,1738]" italics="true" pageId="6" pageNumber="614">c.</emphasis>
|
||||
<quantity box="[1206,1312,1716,1738]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="6" pageNumber="614" unit="mm" value="0.75">0.75 mm</quantity>
|
||||
; compound eyes,
|
||||
<emphasis box="[889,906,1747,1768]" italics="true" pageId="6" pageNumber="614">c.</emphasis>
|
||||
<quantity box="[913,1014,1747,1769]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="6" pageNumber="614" unit="mm" value="0.45">0.45 mm</quantity>
|
||||
long, circular, situated laterally near head midlength, and separated from posterior head margin by more than their length; fontanelle not visible and probably absent; antenna moniliform with at least 22 antennomeres. Pronotum
|
||||
<emphasis box="[1260,1277,1870,1891]" italics="true" pageId="6" pageNumber="614">c.</emphasis>
|
||||
<quantity box="[1285,1386,1869,1891]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="614" unit="mm" value="1.6">1.60 mm</quantity>
|
||||
long marginal teeth (number corresponding); sm, submentum; st, stipe; ttp, tentorial pit. Scale bars
|
||||
<quantity box="[1039,1112,1907,1929]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="615" unit="mm" value="0.5">0.5 mm</quantity>
|
||||
(A, C, D);
|
||||
<quantity box="[1205,1261,1907,1929]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="615" unit="mm" value="1.0">1 mm</quantity>
|
||||
(B). and
|
||||
<emphasis box="[195,212,198,219]" italics="true" pageId="9" pageNumber="617">c.</emphasis>
|
||||
<quantity box="[220,320,197,219]" metricMagnitude="-3" metricUnit="m" metricValue="2.73" pageId="9" pageNumber="617" unit="mm" value="2.73">2.73 mm</quantity>
|
||||
wide, overlapping head dorsally, sides slightly upturned.
|
||||
</paragraph>
|
||||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7195127" ID-Zenodo-Dep="7195127" httpUri="https://zenodo.org/record/7195127/files/figure.png" pageId="7" pageNumber="615" startId="7.[145,221,1760,1782]" targetBox="[199,1370,198,1744]" targetPageId="7">
|
||||
<paragraph blockId="7.[145,1426,1760,1899]" pageId="7" pageNumber="615">
|
||||
<emphasis bold="true" box="[145,245,1760,1782]" pageId="7" pageNumber="615">Figure 4.</emphasis>
|
||||
<taxonomicName authority="Jouault & Engel & Fls & Legendre & Huang & Grandcolas & Nel, 2022" authorityName="Jouault & Engel & Fls & Legendre & Huang & Grandcolas & Nel" authorityYear="2022" box="[255,517,1760,1782]" class="Insecta" family="Mastotermitidae" genus="Anisotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="615" phylum="Arthropoda" rank="species" species="bourguignoni" status="sp. nov.">
|
||||
<emphasis box="[255,517,1760,1782]" italics="true" pageId="7" pageNumber="615">Anisotermes bourguignoni</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel box="[522,597,1760,1782]" pageId="7" pageNumber="615" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
, holotype NIGP177751. Detailed photographs. A, head and pronoun in dorsal view; B, head in ventral view; C, high-resolution X-ray microtomography reconstruction of head in ventral view; D, high-resolution X-ray microtomography reconstruction of head in anterolateral view; E, line drawing of mandibles; F, line drawing of
|
||||
<taxonomicName authorityName="Froggatt" authorityYear="1897" box="[1182,1424,1819,1841]" class="Insecta" family="Mastotermitidae" genus="Mastotermes" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="615" phylum="Arthropoda" rank="species" species="darwinensis">
|
||||
<emphasis box="[1182,1424,1819,1841]" italics="true" pageId="7" pageNumber="615">Mastotermes darwinensis</emphasis>
|
||||
</taxonomicName>
|
||||
mandibles.gl, glossa; la, left apical tooth; lb, labrum; lc, lacinia; lci,lacinial incisor; LM1-LM3, left marginal teeth (number corresponding); Lp, labial palp; mb, mandibles; Mp, maxillary palp; mt, mentum; pc, postclypeus; pgl, paraglossa; ra, right apical tooth; RM1-RM2, right
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7195129" ID-Zenodo-Dep="7195129" httpUri="https://zenodo.org/record/7195129/files/figure.png" pageId="8" pageNumber="616" startId="8.[163,243,1711,1733]" targetBox="[166,1440,198,1668]" targetPageId="8">
|
||||
<paragraph blockId="8.[162,1444,1711,1880]" pageId="8" pageNumber="616">
|
||||
<emphasis bold="true" box="[163,269,1711,1733]" pageId="8" pageNumber="616">Figure 5.</emphasis>
|
||||
<taxonomicName authority="Jouault & Engel & Fls & Legendre & Huang & Grandcolas & Nel, 2022" authorityName="Jouault & Engel & Fls & Legendre & Huang & Grandcolas & Nel" authorityYear="2022" box="[282,562,1711,1733]" class="Insecta" family="Mastotermitidae" genus="Anisotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="616" phylum="Arthropoda" rank="species" species="bourguignoni" status="sp. nov.">
|
||||
<emphasis box="[282,562,1711,1733]" italics="true" pageId="8" pageNumber="616">Anisotermes bourguignoni</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel box="[569,650,1711,1733]" pageId="8" pageNumber="616" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
, holotype NIGP177751. Detailed photographs. A, protarus; B, mesotarsus; C, apex of metatibia; D, line drawing of scale venation with names of veins labelled; E, scale; F, cerci in ventral view with arrows pointing to cercomeres; G, cerci in dorsal view; H, interpretative drawing of protibia; I, interpretative drawing of mesotibiae; J, interpretative drawing of metatibiae. CuA, anterior cubitus; MA, media; PCu, postcubitus; RA, anterior radius; RP, posterior radius; Sc, subcosta; for the nomenclature of tibial spines and spurs refers to
|
||||
<bibRefCitation author="Krishna K & Grimaldi DA & Krishna V & Engel MS" box="[1175,1376,1829,1851]" pageId="8" pageNumber="616" refId="ref13371" refString="Krishna K, Grimaldi DA, Krishna V, Engel MS. 2013. Treatise on the Isoptera of the world. Bulletin of the American Museum of Natural History 377: 19891 - 12433." type="journal volume" year="2013">
|
||||
Krishna
|
||||
<emphasis box="[1266,1315,1829,1851]" italics="true" pageId="8" pageNumber="616">et al.</emphasis>
|
||||
, 2013
|
||||
</bibRefCitation>
|
||||
. Scale bars: 0.2 mm (A, F, G); 0.5 mm (B, C, D, E, G, H, I, J).
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph blockId="9.[141,763,197,863]" pageId="9" pageNumber="617">
|
||||
Legs robust; procoxa
|
||||
<emphasis box="[423,441,259,280]" italics="true" pageId="9" pageNumber="617">c.</emphasis>
|
||||
<quantity box="[449,554,259,280]" metricMagnitude="-3" metricUnit="m" metricValue="1.38" pageId="9" pageNumber="617" unit="mm" value="1.38">1.38 mm</quantity>
|
||||
long, profemur
|
||||
<emphasis box="[746,763,259,280]" italics="true" pageId="9" pageNumber="617">c.</emphasis>
|
||||
<quantity box="[145,237,289,311]" metricMagnitude="-3" metricUnit="m" metricValue="1.51" pageId="9" pageNumber="617" unit="mm" value="1.51">1.51mm</quantity>
|
||||
long, protibia
|
||||
<emphasis box="[390,407,290,311]" italics="true" pageId="9" pageNumber="617">c.</emphasis>
|
||||
<quantity box="[413,505,289,311]" metricMagnitude="-3" metricUnit="m" metricValue="1.41" pageId="9" pageNumber="617" unit="mm" value="1.41">1.41mm</quantity>
|
||||
long, protarsus at least
|
||||
<quantity box="[143,246,320,342]" metricMagnitude="-4" metricUnit="m" metricValue="7.6" pageId="9" pageNumber="617" unit="mm" value="0.76">0.76 mm</quantity>
|
||||
long; protibia with three spurs f1, f2, f3 and additional spines; mesocoxa
|
||||
<emphasis box="[456,473,351,372]" italics="true" pageId="9" pageNumber="617">c.</emphasis>
|
||||
<quantity box="[478,572,351,372]" metricMagnitude="-3" metricUnit="m" metricValue="1.13" pageId="9" pageNumber="617" unit="mm" value="1.13">1.13 mm</quantity>
|
||||
long, mesofemur medially swollen
|
||||
<emphasis box="[339,356,382,403]" italics="true" pageId="9" pageNumber="617">c.</emphasis>
|
||||
<quantity box="[362,460,381,403]" metricMagnitude="-3" metricUnit="m" metricValue="1.57" pageId="9" pageNumber="617" unit="mm" value="1.57">1.57 mm</quantity>
|
||||
long, mesotibia
|
||||
<emphasis box="[640,657,382,403]" italics="true" pageId="9" pageNumber="617">c.</emphasis>
|
||||
<quantity box="[663,761,381,403]" metricMagnitude="-3" metricUnit="m" metricValue="1.72" pageId="9" pageNumber="617" unit="mm" value="1.72">1.72 mm</quantity>
|
||||
long, mesotarsus
|
||||
<emphasis box="[344,361,413,434]" italics="true" pageId="9" pageNumber="617">c.</emphasis>
|
||||
<quantity box="[368,468,412,434]" metricMagnitude="-4" metricUnit="m" metricValue="6.4" pageId="9" pageNumber="617" unit="mm" value="0.64">0.64 mm</quantity>
|
||||
long; mesotibia with four observable spurs m1, m2, m3, m4 and additional spines; metacoxa at least
|
||||
<quantity box="[360,464,473,495]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="9" pageNumber="617" unit="mm" value="1.1">1.10 mm</quantity>
|
||||
long, metafemur at least
|
||||
<quantity box="[146,236,504,525]" metricMagnitude="-3" metricUnit="m" metricValue="2.24" pageId="9" pageNumber="617" unit="mm" value="2.24">2.24mm</quantity>
|
||||
long, metatibia at least
|
||||
<quantity box="[491,585,504,526]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="617" unit="mm" value="1.5">1.50 mm</quantity>
|
||||
long, with three observable spurs h1, h2, h3, h4 and additional spines; all spurs flattened with serration; all tarsi pentamerous; pretarsal arolium present. Forewing scale
|
||||
<emphasis box="[636,653,597,618]" italics="true" pageId="9" pageNumber="617">c.</emphasis>
|
||||
<quantity box="[661,762,596,618]" metricMagnitude="-3" metricUnit="m" metricValue="2.36" pageId="9" pageNumber="617" unit="mm" value="2.36">2.36 mm</quantity>
|
||||
long and
|
||||
<quantity box="[248,345,627,648]" metricMagnitude="-3" metricUnit="m" metricValue="1.41" pageId="9" pageNumber="617" unit="mm" value="1.41">1.41 mm</quantity>
|
||||
wide (measured distally), sclerotized, especially in anal area, with sparse setation. Hindwing scale not visible but proximal part of hindwing clearly cut beyond basal suture.
|
||||
</paragraph>
|
||||
<paragraph blockId="9.[141,763,197,863]" pageId="9" pageNumber="617">
|
||||
Abdomen at least.6.0mm long with at least11observable segments; abdominal segments widest at midlength. Cerci multimerous
|
||||
<emphasis box="[297,313,811,832]" italics="true" pageId="9" pageNumber="617">c.</emphasis>
|
||||
<quantity box="[320,420,810,832]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="9" pageNumber="617" unit="mm" value="0.52">0.52 mm</quantity>
|
||||
long, five to seven cercomeres (sometimes fused so not clearly countable).
|
||||
</paragraph>
|
||||
<paragraph blockId="9.[145,403,888,910]" box="[145,403,888,910]" pageId="9" pageNumber="617">
|
||||
<heading box="[145,403,888,910]" centered="true" fontSize="9" level="2" pageId="9" pageNumber="617" reason="2">
|
||||
<emphasis box="[145,228,888,909]" italics="true" pageId="9" pageNumber="617">Colour:</emphasis>
|
||||
Not preserved.
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="9" pageNumber="617" type="discussion">
|
||||
<paragraph blockId="9.[145,762,934,1907]" lastBlockId="9.[809,1428,197,556]" pageId="9" pageNumber="617">
|
||||
<emphasis box="[145,599,934,956]" italics="true" pageId="9" pageNumber="617">
|
||||
Remarks:
|
||||
<taxonomicName authorityName="Jouault & Engel & Fls & Legendre & Huang & Grandcolas & Nel" authorityYear="2022" box="[271,599,934,956]" class="Insecta" family="Mastotermitidae" genus="Anisotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="617" phylum="Arthropoda" rank="species" species="bourguignoni">Anisotermes bourguignoni</taxonomicName>
|
||||
</emphasis>
|
||||
differs from most other Cretaceous ‘lower’
|
||||
<taxonomicName authorityName="BRULLE" authorityYear="1832" box="[513,612,966,987]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Isoptera" pageId="9" pageNumber="617" phylum="Tracheophyta" rank="order">Isoptera</taxonomicName>
|
||||
in having a plesiomorphic character in scale configuration (e.g. fine reticulation between veins in the scale), tarsi pentamerous (plesiomorphic character), a tibial spur formula of 3-4-4 and an overall massive trapezoidal-shaped pronotum. The wholly pentamerous tarsi and enriched tibial spur formula tend to imply an early diverged position within
|
||||
<taxonomicName authorityName="BRULLE" authorityYear="1832" box="[226,322,1180,1201]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Isoptera" pageId="9" pageNumber="617" phylum="Tracheophyta" rank="order">Isoptera</taxonomicName>
|
||||
, among groups like
|
||||
<taxonomicName box="[549,757,1180,1202]" class="Insecta" family="Archotermopsidae" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="617" phylum="Arthropoda" rank="family">Archotermopsidae</taxonomicName>
|
||||
,
|
||||
<taxonomicName box="[145,333,1210,1232]" class="Insecta" family="Hodotermitidae" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="617" phylum="Arthropoda" rank="family">Hodotermitidae</taxonomicName>
|
||||
,
|
||||
<taxonomicName authorityName="DESNEUX" authorityYear="1904" box="[346,543,1210,1232]" class="Insecta" family="Mastotermitidae" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="617" phylum="Arthropoda" rank="family">Mastotermitidae</taxonomicName>
|
||||
and their various extinct relatives. Additionally, its general habitus, and particularly its large pronotum, suggests that
|
||||
<taxonomicName authorityName="Jouault & Engel & Fls & Legendre & Huang & Grandcolas & Nel" authorityYear="2022" box="[145,321,1302,1324]" class="Insecta" family="Mastotermitidae" genus="Anisotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="617" phylum="Arthropoda" rank="species" species="bourguignoni">
|
||||
<emphasis box="[145,321,1302,1324]" italics="true" pageId="9" pageNumber="617">A. bourguignoni</emphasis>
|
||||
</taxonomicName>
|
||||
is related to
|
||||
<taxonomicName authorityName="DESNEUX" authorityYear="1904" box="[465,649,1302,1324]" class="Insecta" family="Mastotermitidae" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="617" phylum="Arthropoda" rank="family">Mastotermitidae</taxonomicName>
|
||||
or to taxa belonging to the ‘
|
||||
<taxonomicName authorityName="Lacasa-Ruiz & Martínez-Delclòs" authorityYear="1986" box="[342,473,1334,1355]" class="Insecta" family="Hodotermitidae" genus="Meiatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="617" phylum="Arthropoda" rank="genus">
|
||||
<emphasis box="[342,473,1334,1355]" italics="true" pageId="9" pageNumber="617">Meiatermes</emphasis>
|
||||
</taxonomicName>
|
||||
-grade’. Interestingly, the hindwings of
|
||||
<taxonomicName authorityName="Jouault & Engel & Fls & Legendre & Huang & Grandcolas & Nel" authorityYear="2022" box="[283,460,1364,1385]" class="Insecta" family="Mastotermitidae" genus="Anisotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="617" phylum="Arthropoda" rank="species" species="bourguignoni">
|
||||
<emphasis box="[283,460,1364,1385]" italics="true" pageId="9" pageNumber="617">A. bourguignoni</emphasis>
|
||||
</taxonomicName>
|
||||
were cut slightly distal from the hindwing basal scale suture, while in the forewing the cut follows the basal suture. The hindwing basal scale suture was probably not fragile enough to break alone after the mating swarm, as it normally happens in a majority of extant termites. This suggests that the suture of the hindwing is rudimentary (symplesiomorphy) as in
|
||||
<taxonomicName authority="(Tillyard, 1931)" baseAuthorityName="Tillyard" baseAuthorityYear="1931" box="[175,651,1578,1600]" class="Insecta" family="Mastotermitidae" genus="Mastotermes" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="617" phylum="Arthropoda" rank="species" species="darwiniensis">
|
||||
<emphasis box="[175,469,1578,1600]" italics="true" pageId="9" pageNumber="617">Mastotermes darwiniensis</emphasis>
|
||||
(
|
||||
<bibRefCitation author="Tillyard RJ" box="[484,642,1578,1600]" pageId="9" pageNumber="617" pagination="371 - 390" refId="ref13990" refString="Tillyard RJ. 1931. The wing venation of the order Isoptera: introduction and the family Mastotermitidae. Proceedings of the Linnean Society of New South Wales 56: 371 - 390." type="journal article" year="1931">Tillyard, 1931</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, and that the new specimen could be related to
|
||||
<taxonomicName authorityName="DESNEUX" authorityYear="1904" box="[568,758,1609,1631]" class="Insecta" family="Mastotermitidae" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="617" phylum="Arthropoda" rank="family">Mastotermitidae</taxonomicName>
|
||||
. Another character suggesting placement of
|
||||
<taxonomicName box="[626,760,1640,1661]" class="Insecta" family="Mastotermitidae" genus="Anisotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="617" phylum="Arthropoda" rank="genus">
|
||||
<emphasis box="[626,760,1640,1661]" italics="true" pageId="9" pageNumber="617">Anisotermes</emphasis>
|
||||
</taxonomicName>
|
||||
within
|
||||
<taxonomicName authorityName="DESNEUX" authorityYear="1904" box="[229,424,1670,1692]" class="Insecta" family="Mastotermitidae" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="617" phylum="Arthropoda" rank="family">Mastotermitidae</taxonomicName>
|
||||
is the mandibular shape. In fact,
|
||||
<taxonomicName box="[204,346,1701,1722]" class="Insecta" family="Mastotermitidae" genus="Anisotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="617" phylum="Arthropoda" rank="genus">
|
||||
<emphasis box="[204,346,1701,1722]" italics="true" pageId="9" pageNumber="617">Anisotermes</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName authorityName="FROGGATT" authorityYear="1897" box="[407,555,1702,1723]" class="Insecta" family="Mastotermitidae" genus="Mastotermes" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="617" phylum="Arthropoda" rank="genus">
|
||||
<emphasis box="[407,555,1702,1723]" italics="true" pageId="9" pageNumber="617">Mastotermes</emphasis>
|
||||
</taxonomicName>
|
||||
exhibit a similar configuration of teeth on both mandibles, and length variations (RM1 conspicuously longer than ra vs. equal in length) are usually indicative of different genera from the same family. Additionally, this placement is strengthened by the diagnostic characters of the new species: ocelli absent, procoxal carina present, arolium present and the aforementioned scale characters. The absence of ocelli indicates affinities with
|
||||
<taxonomicName box="[1088,1276,228,250]" class="Insecta" family="Mastotermitidae" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="624" phylum="Arthropoda" rank="subFamily" subFamily="Idanotermitinae">Idanotermitinae</taxonomicName>
|
||||
, but the new fossil differs from the genus
|
||||
<taxonomicName authority="Engel, 2008" authorityName="Engel" authorityYear="2008" box="[1139,1425,258,280]" class="Insecta" family="Mastotermitidae" genus="Idanotermes" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="617" phylum="Arthropoda" rank="genus">
|
||||
<emphasis box="[1139,1281,258,279]" italics="true" pageId="9" pageNumber="617">Idanotermes</emphasis>
|
||||
<bibRefCitation author="Engel MS" box="[1289,1425,258,280]" pageId="9" pageNumber="617" pagination="194 - 203" refId="ref12740" refString="Engel MS. 2008. Two new termites in Baltic amber (Isoptera). Journal of the Kansas Entomological Society 81 (3): 194 - 203." type="journal article" year="2008">
|
||||
Engel,
|
||||
<quantity box="[1370,1425,259,280]" metricMagnitude="1" metricUnit="m" metricValue="5.10032" pageId="9" pageNumber="617" unit="in" value="2008.0">2008</quantity>
|
||||
</bibRefCitation>
|
||||
</taxonomicName>
|
||||
in at least its different tibial spur formula (i.e. 3-4-4 vs. 3-4-3). The genus
|
||||
<taxonomicName box="[1059,1200,320,341]" class="Insecta" family="Mastotermitidae" genus="Anisotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="617" phylum="Arthropoda" rank="genus">
|
||||
<emphasis box="[1059,1200,320,341]" italics="true" pageId="9" pageNumber="617">Anisotermes</emphasis>
|
||||
</taxonomicName>
|
||||
is here tentatively considered as a member of the
|
||||
<taxonomicName box="[1161,1346,350,372]" class="Insecta" family="Mastotermitidae" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="624" phylum="Arthropoda" rank="subFamily" subFamily="Idanotermitinae">Idanotermitinae</taxonomicName>
|
||||
(
|
||||
<bibRefCitation author="Jiang RX & Zhang HR & Eldredge KT & Song XB & Li YY & Tihelka E & Huang DY & Wang S & Engel MS & Cai C" pageId="9" pageNumber="617" pagination="374 - 389" refId="ref13160" refString="Jiang RX, Zhang HR, Eldredge KT, Song XB, Li YY, Tihelka E, Huang DY, Wang S, Engel MS, Cai C. 2021. Further evidence of Cretaceous termitophily: description of new termite hosts of the trichopseniine Cretotrichopsenius (Coleoptera: Staphylinidae), with emendations to the classification of lower termites (Isoptera). Palaeoentomology 4: 374 - 389." type="journal article" year="2021">
|
||||
Jiang
|
||||
<emphasis box="[809,868,381,403]" italics="true" pageId="9" pageNumber="617">et al.</emphasis>
|
||||
, 2021: 379
|
||||
</bibRefCitation>
|
||||
); we confirm this placement mainly based on the presence of the anal lobe (characteristics of
|
||||
<taxonomicName authorityName="DESNEUX" authorityYear="1904" box="[809,996,442,464]" class="Insecta" family="Mastotermitidae" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="617" phylum="Arthropoda" rank="family">Mastotermitidae</taxonomicName>
|
||||
within the
|
||||
<taxonomicName authorityName="BRULLE" authorityYear="1832" box="[1126,1219,443,464]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Isoptera" pageId="9" pageNumber="617" phylum="Tracheophyta" rank="order">Isoptera</taxonomicName>
|
||||
) and based on the absence of ocelli. The new species differs from
|
||||
<taxonomicName authorityName="Zhao, Eggleton & Ren" authorityYear="2019" box="[1319,1390,473,495]" class="Insecta" family="Mastotermitidae" genus="Anisotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="617" phylum="Arthropoda" rank="species" species="xiai">
|
||||
<emphasis box="[1319,1390,473,495]" italics="true" pageId="9" pageNumber="617">A. xiai</emphasis>
|
||||
</taxonomicName>
|
||||
by,
|
||||
<emphasis box="[809,908,504,525]" italics="true" pageId="9" pageNumber="617">inter alia</emphasis>
|
||||
, the presence of the pretarsal arolium and more additional spines along the tibiae.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
Loading…
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Reference in a new issue