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@ -62,7 +65,7 @@ Wright, 1859
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<paragraph id="8B9F36E4FF9AFF972FCAB8B1FEF1FA2E" blockId="1.[151,505,1446,1509]" box="[151,282,1483,1509]" pageId="1" pageNumber="388">
<figureCitation id="131B2A61FF9AFF972FCAB8B1FEF1FA2E" box="[151,282,1483,1509]" captionStart="FIGURE 1" captionStartId="2.[151,250,1874,1898]" captionTargetBox="[286,1301,179,1848]" captionTargetId="figure-23@2.[286,1301,179,1848]" captionTargetPageId="2" captionText="FIGURE 1. Epibiosis of Bimeria vestita Wright, 1859 on Pelogenia fimbriata (Hartman, 1939) (LACM-AHF Poly 1576a). A.Anterior end of P. fimbriata, dorsal view, the asterisk indicates the prostomium, and the arrowhead indicates B. vestita attached to the scale. B. Hydroid colony on the surface of the scale still attached to the dorsum, side view. C. Anterior scale removed. D. Close-up of hydrants. E. Close-up of the colony, arrowhead indicates gonophore. F. Detail of tentacles. G. Nematocyst microbasic eurytele. H. SEM of a hydrant, arrowhead indicates pseudohydrotheca." pageId="1" pageNumber="388">Figs. 1A–H</figureCitation>
<figureCitation id="131B2A61FF9AFF972FCAB8B1FEF1FA2E" box="[151,282,1483,1509]" captionStart="FIGURE 1" captionStartId="2.[151,250,1874,1898]" captionTargetBox="[286,1301,179,1848]" captionTargetId="figure-23@2.[286,1301,179,1848]" captionTargetPageId="2" captionText="FIGURE 1. Epibiosis of Bimeria vestita Wright, 1859 on Pelogenia fimbriata (Hartman, 1939) (LACM-AHF Poly 1576a). A.Anterior end of P. fimbriata, dorsal view, the asterisk indicates the prostomium, and the arrowhead indicates B. vestita attached to the scale. B. Hydroid colony on the surface of the scale still attached to the dorsum, side view. C. Anterior scale removed. D. Close-up of hydrants. E. Close-up of the colony, arrowhead indicates gonophore. F. Detail of tentacles. G. Nematocyst microbasic eurytele. H. SEM of a hydrant, arrowhead indicates pseudohydrotheca." figureDoi="http://doi.org/10.5281/zenodo.14734823" httpUri="https://zenodo.org/record/14734823/files/figure.png" pageId="1" pageNumber="388">Figs. 1A–H</figureCitation>
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@ -115,15 +118,15 @@ above the dorsal line of the worm and extended to the lateral sides of the scale
Mature colony small (
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high), stolonal, monosiphonic, arising from creeping tubular stolons (
<figureCitation id="131B2A61FF9AFF972FC2BBBDFEC9F92A" box="[159,290,1735,1761]" captionStart="FIGURE 1" captionStartId="2.[151,250,1874,1898]" captionTargetBox="[286,1301,179,1848]" captionTargetId="figure-23@2.[286,1301,179,1848]" captionTargetPageId="2" captionText="FIGURE 1. Epibiosis of Bimeria vestita Wright, 1859 on Pelogenia fimbriata (Hartman, 1939) (LACM-AHF Poly 1576a). A.Anterior end of P. fimbriata, dorsal view, the asterisk indicates the prostomium, and the arrowhead indicates B. vestita attached to the scale. B. Hydroid colony on the surface of the scale still attached to the dorsum, side view. C. Anterior scale removed. D. Close-up of hydrants. E. Close-up of the colony, arrowhead indicates gonophore. F. Detail of tentacles. G. Nematocyst microbasic eurytele. H. SEM of a hydrant, arrowhead indicates pseudohydrotheca." pageId="1" pageNumber="388">Figs. 1A–E</figureCitation>
<figureCitation id="131B2A61FF9AFF972FC2BBBDFEC9F92A" box="[159,290,1735,1761]" captionStart="FIGURE 1" captionStartId="2.[151,250,1874,1898]" captionTargetBox="[286,1301,179,1848]" captionTargetId="figure-23@2.[286,1301,179,1848]" captionTargetPageId="2" captionText="FIGURE 1. Epibiosis of Bimeria vestita Wright, 1859 on Pelogenia fimbriata (Hartman, 1939) (LACM-AHF Poly 1576a). A.Anterior end of P. fimbriata, dorsal view, the asterisk indicates the prostomium, and the arrowhead indicates B. vestita attached to the scale. B. Hydroid colony on the surface of the scale still attached to the dorsum, side view. C. Anterior scale removed. D. Close-up of hydrants. E. Close-up of the colony, arrowhead indicates gonophore. F. Detail of tentacles. G. Nematocyst microbasic eurytele. H. SEM of a hydrant, arrowhead indicates pseudohydrotheca." figureDoi="http://doi.org/10.5281/zenodo.14734823" httpUri="https://zenodo.org/record/14734823/files/figure.png" pageId="1" pageNumber="388">Figs. 1A–E</figureCitation>
). Bilayered exoskeleton formed by perisarc and exosarc. Perisarc continuous from hydrorhiza to tentacle base unwrinkled at the base of hydrocaulus. Exosarc thin, thinner over hydranth, tentacle base, and hypostome, forming a pseudohydrotheca (
<figureCitation id="131B2A61FF9AFF972EBABA75FD60F8E2" box="[487,651,1807,1833]" captionStart="FIGURE 1" captionStartId="2.[151,250,1874,1898]" captionTargetBox="[286,1301,179,1848]" captionTargetId="figure-23@2.[286,1301,179,1848]" captionTargetPageId="2" captionText="FIGURE 1. Epibiosis of Bimeria vestita Wright, 1859 on Pelogenia fimbriata (Hartman, 1939) (LACM-AHF Poly 1576a). A.Anterior end of P. fimbriata, dorsal view, the asterisk indicates the prostomium, and the arrowhead indicates B. vestita attached to the scale. B. Hydroid colony on the surface of the scale still attached to the dorsum, side view. C. Anterior scale removed. D. Close-up of hydrants. E. Close-up of the colony, arrowhead indicates gonophore. F. Detail of tentacles. G. Nematocyst microbasic eurytele. H. SEM of a hydrant, arrowhead indicates pseudohydrotheca." pageId="1" pageNumber="388">Figs. 1D, F, H</figureCitation>
<figureCitation id="131B2A61FF9AFF972EBABA75FD60F8E2" box="[487,651,1807,1833]" captionStart="FIGURE 1" captionStartId="2.[151,250,1874,1898]" captionTargetBox="[286,1301,179,1848]" captionTargetId="figure-23@2.[286,1301,179,1848]" captionTargetPageId="2" captionText="FIGURE 1. Epibiosis of Bimeria vestita Wright, 1859 on Pelogenia fimbriata (Hartman, 1939) (LACM-AHF Poly 1576a). A.Anterior end of P. fimbriata, dorsal view, the asterisk indicates the prostomium, and the arrowhead indicates B. vestita attached to the scale. B. Hydroid colony on the surface of the scale still attached to the dorsum, side view. C. Anterior scale removed. D. Close-up of hydrants. E. Close-up of the colony, arrowhead indicates gonophore. F. Detail of tentacles. G. Nematocyst microbasic eurytele. H. SEM of a hydrant, arrowhead indicates pseudohydrotheca." figureDoi="http://doi.org/10.5281/zenodo.14734823" httpUri="https://zenodo.org/record/14734823/files/figure.png" pageId="1" pageNumber="388">Figs. 1D, F, H</figureCitation>
). Hydranth vasiform, merging with short pedicel. Hypostome conical, 10–12 tentacles in two very close whorls (
<figureCitation id="131B2A61FF9AFF972D20BA49FD38F886" box="[637,723,1843,1869]" captionStart="FIGURE 1" captionStartId="2.[151,250,1874,1898]" captionTargetBox="[286,1301,179,1848]" captionTargetId="figure-23@2.[286,1301,179,1848]" captionTargetPageId="2" captionText="FIGURE 1. Epibiosis of Bimeria vestita Wright, 1859 on Pelogenia fimbriata (Hartman, 1939) (LACM-AHF Poly 1576a). A.Anterior end of P. fimbriata, dorsal view, the asterisk indicates the prostomium, and the arrowhead indicates B. vestita attached to the scale. B. Hydroid colony on the surface of the scale still attached to the dorsum, side view. C. Anterior scale removed. D. Close-up of hydrants. E. Close-up of the colony, arrowhead indicates gonophore. F. Detail of tentacles. G. Nematocyst microbasic eurytele. H. SEM of a hydrant, arrowhead indicates pseudohydrotheca." pageId="1" pageNumber="388">Fig. 1F</figureCitation>
<figureCitation id="131B2A61FF9AFF972D20BA49FD38F886" box="[637,723,1843,1869]" captionStart="FIGURE 1" captionStartId="2.[151,250,1874,1898]" captionTargetBox="[286,1301,179,1848]" captionTargetId="figure-23@2.[286,1301,179,1848]" captionTargetPageId="2" captionText="FIGURE 1. Epibiosis of Bimeria vestita Wright, 1859 on Pelogenia fimbriata (Hartman, 1939) (LACM-AHF Poly 1576a). A.Anterior end of P. fimbriata, dorsal view, the asterisk indicates the prostomium, and the arrowhead indicates B. vestita attached to the scale. B. Hydroid colony on the surface of the scale still attached to the dorsum, side view. C. Anterior scale removed. D. Close-up of hydrants. E. Close-up of the colony, arrowhead indicates gonophore. F. Detail of tentacles. G. Nematocyst microbasic eurytele. H. SEM of a hydrant, arrowhead indicates pseudohydrotheca." figureDoi="http://doi.org/10.5281/zenodo.14734823" httpUri="https://zenodo.org/record/14734823/files/figure.png" pageId="1" pageNumber="388">Fig. 1F</figureCitation>
). Gonophore fixed sporosac on hydrorhiza, pedicellate, ovoid, without radial canals, enveloped in the thin exoskeleton (
<figureCitation id="131B2A61FF9AFF972C79BA2DFC90F8BA" box="[804,891,1879,1905]" captionStart="FIGURE 1" captionStartId="2.[151,250,1874,1898]" captionTargetBox="[286,1301,179,1848]" captionTargetId="figure-23@2.[286,1301,179,1848]" captionTargetPageId="2" captionText="FIGURE 1. Epibiosis of Bimeria vestita Wright, 1859 on Pelogenia fimbriata (Hartman, 1939) (LACM-AHF Poly 1576a). A.Anterior end of P. fimbriata, dorsal view, the asterisk indicates the prostomium, and the arrowhead indicates B. vestita attached to the scale. B. Hydroid colony on the surface of the scale still attached to the dorsum, side view. C. Anterior scale removed. D. Close-up of hydrants. E. Close-up of the colony, arrowhead indicates gonophore. F. Detail of tentacles. G. Nematocyst microbasic eurytele. H. SEM of a hydrant, arrowhead indicates pseudohydrotheca." pageId="1" pageNumber="388">Fig. 1E</figureCitation>
<figureCitation id="131B2A61FF9AFF972C79BA2DFC90F8BA" box="[804,891,1879,1905]" captionStart="FIGURE 1" captionStartId="2.[151,250,1874,1898]" captionTargetBox="[286,1301,179,1848]" captionTargetId="figure-23@2.[286,1301,179,1848]" captionTargetPageId="2" captionText="FIGURE 1. Epibiosis of Bimeria vestita Wright, 1859 on Pelogenia fimbriata (Hartman, 1939) (LACM-AHF Poly 1576a). A.Anterior end of P. fimbriata, dorsal view, the asterisk indicates the prostomium, and the arrowhead indicates B. vestita attached to the scale. B. Hydroid colony on the surface of the scale still attached to the dorsum, side view. C. Anterior scale removed. D. Close-up of hydrants. E. Close-up of the colony, arrowhead indicates gonophore. F. Detail of tentacles. G. Nematocyst microbasic eurytele. H. SEM of a hydrant, arrowhead indicates pseudohydrotheca." figureDoi="http://doi.org/10.5281/zenodo.14734823" httpUri="https://zenodo.org/record/14734823/files/figure.png" pageId="1" pageNumber="388">Fig. 1E</figureCitation>
). Nematocysts of tentacles: microbasic eurytele and desmoneme (
<figureCitation id="131B2A61FF9AFF972E01BA01FE5EF85E" box="[348,437,1915,1941]" captionStart="FIGURE 1" captionStartId="2.[151,250,1874,1898]" captionTargetBox="[286,1301,179,1848]" captionTargetId="figure-23@2.[286,1301,179,1848]" captionTargetPageId="2" captionText="FIGURE 1. Epibiosis of Bimeria vestita Wright, 1859 on Pelogenia fimbriata (Hartman, 1939) (LACM-AHF Poly 1576a). A.Anterior end of P. fimbriata, dorsal view, the asterisk indicates the prostomium, and the arrowhead indicates B. vestita attached to the scale. B. Hydroid colony on the surface of the scale still attached to the dorsum, side view. C. Anterior scale removed. D. Close-up of hydrants. E. Close-up of the colony, arrowhead indicates gonophore. F. Detail of tentacles. G. Nematocyst microbasic eurytele. H. SEM of a hydrant, arrowhead indicates pseudohydrotheca." pageId="1" pageNumber="388">Fig. 1G</figureCitation>
<figureCitation id="131B2A61FF9AFF972E01BA01FE5EF85E" box="[348,437,1915,1941]" captionStart="FIGURE 1" captionStartId="2.[151,250,1874,1898]" captionTargetBox="[286,1301,179,1848]" captionTargetId="figure-23@2.[286,1301,179,1848]" captionTargetPageId="2" captionText="FIGURE 1. Epibiosis of Bimeria vestita Wright, 1859 on Pelogenia fimbriata (Hartman, 1939) (LACM-AHF Poly 1576a). A.Anterior end of P. fimbriata, dorsal view, the asterisk indicates the prostomium, and the arrowhead indicates B. vestita attached to the scale. B. Hydroid colony on the surface of the scale still attached to the dorsum, side view. C. Anterior scale removed. D. Close-up of hydrants. E. Close-up of the colony, arrowhead indicates gonophore. F. Detail of tentacles. G. Nematocyst microbasic eurytele. H. SEM of a hydrant, arrowhead indicates pseudohydrotheca." figureDoi="http://doi.org/10.5281/zenodo.14734823" httpUri="https://zenodo.org/record/14734823/files/figure.png" pageId="1" pageNumber="388">Fig. 1G</figureCitation>
).
</paragraph>
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@ -149,7 +152,7 @@ Mendoza-Becerril
).
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FIGURE 1. Epibiosis of

91
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@ -75,7 +78,7 @@ Chinese common name:
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(
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<figureCitation id="13042A3CFFE8857AFF2EF57DFF16F8FA" box="[159,227,1831,1858]" captionStart="FIGURE 4" captionStartId="7.[151,250,1651,1675]" captionTargetBox="[156,1436,230,1625]" captionTargetId="figure-45@7.[151,1436,230,1628]" captionTargetPageId="7" captionText="FIGURE 4. Scydmaenus (s. str.) vestitus (A–E. Male. F. Female). A. Dorsal habitus. B. Meso- and metaventrite. C. Mesotibia. D, E. Aedeagus, dorsal (D) and lateral (E). F. Spermatheca, lateral. Scale bars: 0.5 mm in A, B; 0.2 mm in C–E; 0.05 mm in F." figureDoi="http://doi.org/10.5281/zenodo.14734869" httpUri="https://zenodo.org/record/14734869/files/figure.png" pageId="6" pageNumber="537">Fig. 4</figureCitation>
)
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@ -171,7 +174,7 @@ locality:
; Li, L.-Z.
<emphasis id="B94BEAABFFE9857BFB83F2CDFBABFF17" box="[1074,1118,151,175]" italics="true" pageId="7" pageNumber="538">et al</emphasis>
. 2019: 342;
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<emphasis id="B94BEAABFFE9857BFF26F429FEE6F933" bold="true" box="[151,275,1651,1675]" pageId="7" pageNumber="538">FIGURE 4.</emphasis>
<taxonomicName id="4C3F4D3AFFE9857BFEA8F429FDC2F933" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[281,567,1651,1675]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="538" phylum="Arthropoda" rank="species" species="vestitus">
@ -250,21 +253,21 @@ leg.,
<emphasis id="B94BEAABFFE9857BFF76F537FEB7F83F" bold="true" box="[199,322,1901,1927]" pageId="7" pageNumber="538">Diagnosis.</emphasis>
<emphasis id="B94BEAABFFE9857BFEF9F537FE7CF83F" box="[328,393,1901,1927]" italics="true" pageId="7" pageNumber="538">Male.</emphasis>
Body (
<figureCitation id="13042A3CFFE9857BFE6BF537FDC4F83F" box="[474,561,1901,1927]" captionStart="FIGURE 4" captionStartId="7.[151,250,1651,1675]" captionTargetBox="[156,1436,230,1625]" captionTargetId="figure-45@7.[151,1436,230,1628]" captionTargetPageId="7" captionText="FIGURE 4. Scydmaenus (s. str.) vestitus (A–E. Male. F. Female). A. Dorsal habitus. B. Meso- and metaventrite. C. Mesotibia. D, E. Aedeagus, dorsal (D) and lateral (E). F. Spermatheca, lateral. Scale bars: 0.5 mm in A, B; 0.2 mm in C–E; 0.05 mm in F." pageId="7" pageNumber="538">Fig. 4A</figureCitation>
<figureCitation id="13042A3CFFE9857BFE6BF537FDC4F83F" box="[474,561,1901,1927]" captionStart="FIGURE 4" captionStartId="7.[151,250,1651,1675]" captionTargetBox="[156,1436,230,1625]" captionTargetId="figure-45@7.[151,1436,230,1628]" captionTargetPageId="7" captionText="FIGURE 4. Scydmaenus (s. str.) vestitus (A–E. Male. F. Female). A. Dorsal habitus. B. Meso- and metaventrite. C. Mesotibia. D, E. Aedeagus, dorsal (D) and lateral (E). F. Spermatheca, lateral. Scale bars: 0.5 mm in A, B; 0.2 mm in C–E; 0.05 mm in F." figureDoi="http://doi.org/10.5281/zenodo.14734869" httpUri="https://zenodo.org/record/14734869/files/figure.png" pageId="7" pageNumber="538">Fig. 4A</figureCitation>
) length over
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. Head and pronotum finely and rather sparsely punctate. Head slightly transverse, suboval, tempora much longer than eyes; antenna with antennomere 5 much longer than 4 and 6, 8 transverse, 9–11 greatly enlarged and forming distinct club, 11 subconical. Pronotum bell-shaped, with four large antebasal impressions. Each elytron with round apex, well exposing pygidium. Metaventrite (
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<figureCitation id="13042A3CFFE9857BFAB3F583FAAFF84B" box="[1282,1370,2009,2035]" captionStart="FIGURE 4" captionStartId="7.[151,250,1651,1675]" captionTargetBox="[156,1436,230,1625]" captionTargetId="figure-45@7.[151,1436,230,1628]" captionTargetPageId="7" captionText="FIGURE 4. Scydmaenus (s. str.) vestitus (A–E. Male. F. Female). A. Dorsal habitus. B. Meso- and metaventrite. C. Mesotibia. D, E. Aedeagus, dorsal (D) and lateral (E). F. Spermatheca, lateral. Scale bars: 0.5 mm in A, B; 0.2 mm in C–E; 0.05 mm in F." figureDoi="http://doi.org/10.5281/zenodo.14734869" httpUri="https://zenodo.org/record/14734869/files/figure.png" pageId="7" pageNumber="538">Fig. 4B</figureCitation>
) with large impression at middle, sides of impression forming short, setose ridges posteriorly. Protarsomere 1 broadened, tarsomeres successively narrowing from 1 to 5; mesotrochanter angulate on ventral margin, mesotibia (
<figureCitation id="13042A3CFFE68574FA8AF2E1FA61FF6D" box="[1339,1428,187,213]" captionStart="FIGURE 4" captionStartId="7.[151,250,1651,1675]" captionTargetBox="[156,1436,230,1625]" captionTargetId="figure-45@7.[151,1436,230,1628]" captionTargetPageId="7" captionText="FIGURE 4. Scydmaenus (s. str.) vestitus (A–E. Male. F. Female). A. Dorsal habitus. B. Meso- and metaventrite. C. Mesotibia. D, E. Aedeagus, dorsal (D) and lateral (E). F. Spermatheca, lateral. Scale bars: 0.5 mm in A, B; 0.2 mm in C–E; 0.05 mm in F." pageId="8" pageNumber="539">Fig. 4C</figureCitation>
<figureCitation id="13042A3CFFE68574FA8AF2E1FA61FF6D" box="[1339,1428,187,213]" captionStart="FIGURE 4" captionStartId="7.[151,250,1651,1675]" captionTargetBox="[156,1436,230,1625]" captionTargetId="figure-45@7.[151,1436,230,1628]" captionTargetPageId="7" captionText="FIGURE 4. Scydmaenus (s. str.) vestitus (A–E. Male. F. Female). A. Dorsal habitus. B. Meso- and metaventrite. C. Mesotibia. D, E. Aedeagus, dorsal (D) and lateral (E). F. Spermatheca, lateral. Scale bars: 0.5 mm in A, B; 0.2 mm in C–E; 0.05 mm in F." figureDoi="http://doi.org/10.5281/zenodo.14734869" httpUri="https://zenodo.org/record/14734869/files/figure.png" pageId="8" pageNumber="539">Fig. 4C</figureCitation>
) with long apical spine; metatibia angulate at inner-apical margin. Aedeagus (
<figureCitation id="13042A3CFFE68574FC4AF285FB86FF41" box="[1019,1139,223,249]" captionStart="FIGURE 4" captionStartId="7.[151,250,1651,1675]" captionTargetBox="[156,1436,230,1625]" captionTargetId="figure-45@7.[151,1436,230,1628]" captionTargetPageId="7" captionText="FIGURE 4. Scydmaenus (s. str.) vestitus (A–E. Male. F. Female). A. Dorsal habitus. B. Meso- and metaventrite. C. Mesotibia. D, E. Aedeagus, dorsal (D) and lateral (E). F. Spermatheca, lateral. Scale bars: 0.5 mm in A, B; 0.2 mm in C–E; 0.05 mm in F." pageId="8" pageNumber="539">Fig. 4D, E</figureCitation>
<figureCitation id="13042A3CFFE68574FC4AF285FB86FF41" box="[1019,1139,223,249]" captionStart="FIGURE 4" captionStartId="7.[151,250,1651,1675]" captionTargetBox="[156,1436,230,1625]" captionTargetId="figure-45@7.[151,1436,230,1628]" captionTargetPageId="7" captionText="FIGURE 4. Scydmaenus (s. str.) vestitus (A–E. Male. F. Female). A. Dorsal habitus. B. Meso- and metaventrite. C. Mesotibia. D, E. Aedeagus, dorsal (D) and lateral (E). F. Spermatheca, lateral. Scale bars: 0.5 mm in A, B; 0.2 mm in C–E; 0.05 mm in F." figureDoi="http://doi.org/10.5281/zenodo.14734869" httpUri="https://zenodo.org/record/14734869/files/figure.png" pageId="8" pageNumber="539">Fig. 4D, E</figureCitation>
) elongate, dorso-ventrally symmetric, laterally greatly curved at middle; apical portion of endophallus armature composed of two sclerites pointed at apices, one markedly long and the other much shorter; area above foramen and below apex of median with short and dense setae.
</paragraph>
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<emphasis id="B94BEAABFFE68574FF76F335FEEEFE31" box="[199,283,367,393]" italics="true" pageId="8" pageNumber="539">Female</emphasis>
. Body length slightly shorter than male. Antenna shorter. Metaventrite moderately convex at middle, lacking impression or ridges. Protarsus slender; ventral margin of mesotrochanter rounded; mesotibia lacking apical spine; metatibia lacking apical angulation. Spermatheca (
<figureCitation id="13042A3CFFE68574FCBEF3EDFC97FE69" box="[783,866,439,465]" captionStart="FIGURE 4" captionStartId="7.[151,250,1651,1675]" captionTargetBox="[156,1436,230,1625]" captionTargetId="figure-45@7.[151,1436,230,1628]" captionTargetPageId="7" captionText="FIGURE 4. Scydmaenus (s. str.) vestitus (A–E. Male. F. Female). A. Dorsal habitus. B. Meso- and metaventrite. C. Mesotibia. D, E. Aedeagus, dorsal (D) and lateral (E). F. Spermatheca, lateral. Scale bars: 0.5 mm in A, B; 0.2 mm in C–E; 0.05 mm in F." pageId="8" pageNumber="539">Fig. 4F</figureCitation>
<figureCitation id="13042A3CFFE68574FCBEF3EDFC97FE69" box="[783,866,439,465]" captionStart="FIGURE 4" captionStartId="7.[151,250,1651,1675]" captionTargetBox="[156,1436,230,1625]" captionTargetId="figure-45@7.[151,1436,230,1628]" captionTargetPageId="7" captionText="FIGURE 4. Scydmaenus (s. str.) vestitus (A–E. Male. F. Female). A. Dorsal habitus. B. Meso- and metaventrite. C. Mesotibia. D, E. Aedeagus, dorsal (D) and lateral (E). F. Spermatheca, lateral. Scale bars: 0.5 mm in A, B; 0.2 mm in C–E; 0.05 mm in F." figureDoi="http://doi.org/10.5281/zenodo.14734869" httpUri="https://zenodo.org/record/14734869/files/figure.png" pageId="8" pageNumber="539">Fig. 4F</figureCitation>
) spherical, dome-shaped, with funnel-like structure located on same axis as insertion of accessory gland.
</paragraph>
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85
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@ -73,7 +76,7 @@ Chinese common name:
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<paragraph id="8B8036B9FFEB8579FF26F145FF1EFC81" blockId="5.[151,671,726,825]" box="[151,235,799,825]" pageId="5" pageNumber="536">
(
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<figureCitation id="13042A3CFFEB8579FF2EF145FF17FC81" box="[159,226,799,825]" captionStart="FIGURE 3" captionStartId="6.[151,250,1617,1641]" captionTargetBox="[154,1433,185,1587]" captionTargetId="figure-20@6.[151,1436,181,1593]" captionTargetPageId="6" captionText="FIGURE 3. Scydmaenus (s. str.) chinensis (A–C. Male. D, E. Female). A. Dorsal habitus. B, C. Aedeagus, dorsal (B), and lateral (C). D, E. Spermatheca, dorsal (D) and lateral (E). Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.1 mm in D, E." figureDoi="http://doi.org/10.5281/zenodo.14734867" httpUri="https://zenodo.org/record/14734867/files/figure.png" pageId="5" pageNumber="536">Fig. 3</figureCitation>
)
</paragraph>
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@ -123,7 +126,7 @@ locality:
; Li, L.-Z.
<emphasis id="B94BEAABFFEB8579FB97F1D9FBA1FC22" box="[1062,1108,898,922]" italics="true" pageId="5" pageNumber="536">et al</emphasis>
. 2019: 342;
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<bibRefCitation id="EFAE4B48FFEB8579FB68F1D8FF05FC00" author="Jaloszynski, P." pageId="5" pageNumber="536" pagination="354 - 368" refId="ref5186" refString="Jaloszynski, P. (2021 b) Review of Scydmaenus species of continental China described by Herbert Franz (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4952 (2), 354 - 368. https: // doi. org / 10.11646 / zootaxa. 4952.2.8" type="journal article" year="2021">Jałoszyński 2021b: 355</bibRefCitation>
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.
@ -176,17 +179,17 @@ leg.,
<emphasis id="B94BEAABFFEB8579FF76F608FEB7FBD4" bold="true" box="[199,322,1106,1132]" pageId="5" pageNumber="536">Diagnosis.</emphasis>
<emphasis id="B94BEAABFFEB8579FEFDF609FE78FBD5" box="[332,397,1107,1133]" italics="true" pageId="5" pageNumber="536">Male.</emphasis>
Body (
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<figureCitation id="13042A3CFFEB8579FE57F609FDB7FBD5" box="[486,578,1107,1133]" captionStart="FIGURE 3" captionStartId="6.[151,250,1617,1641]" captionTargetBox="[154,1433,185,1587]" captionTargetId="figure-20@6.[151,1436,181,1593]" captionTargetPageId="6" captionText="FIGURE 3. Scydmaenus (s. str.) chinensis (A–C. Male. D, E. Female). A. Dorsal habitus. B, C. Aedeagus, dorsal (B), and lateral (C). D, E. Spermatheca, dorsal (D) and lateral (E). Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.1 mm in D, E." figureDoi="http://doi.org/10.5281/zenodo.14734867" httpUri="https://zenodo.org/record/14734867/files/figure.png" pageId="5" pageNumber="536">Fig. 3A</figureCitation>
) length approximately
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. Head and pronotum finely and rather sparsely punctate.Head slightly longer than wide, suboval, tempora much longer than eyes; antenna with antennomere 5 much longer than 4 and 6, 7 and 8 transversely subconical, 9–11 greatly enlarged and forming distinct club, 11 subconical. Pronotum bell-shaped, with four large antebasal impressions; lateral margins distinctly convergent posteriorly from broadest point to base. Each elytron with round apex, well exposing pygidium. Metaventrite and legs lacking modifications. Protarsomere 1 broadened, tarsomeres successively narrowing from 1 to 5. Aedeagus (
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<figureCitation id="13042A3CFFEB8579FADEF75DFF29FAFD" captionStart="FIGURE 3" captionStartId="6.[151,250,1617,1641]" captionTargetBox="[154,1433,185,1587]" captionTargetId="figure-20@6.[151,1436,181,1593]" captionTargetPageId="6" captionText="FIGURE 3. Scydmaenus (s. str.) chinensis (A–C. Male. D, E. Female). A. Dorsal habitus. B, C. Aedeagus, dorsal (B), and lateral (C). D, E. Spermatheca, dorsal (D) and lateral (E). Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.1 mm in D, E." figureDoi="http://doi.org/10.5281/zenodo.14734867" httpUri="https://zenodo.org/record/14734867/files/figure.png" pageId="5" pageNumber="536">Fig. 3B, C</figureCitation>
) elongate, dorso-ventrally almost symmetric, laterally moderately curved at middle; endophallus armature composed of broad sclerite bifurcate in apical part, forming two lobes, left lobe shorter than right one, curved at apex, right one gradually narrowing toward apex; areas lateral to ostium with rows of long, curved setae.
</paragraph>
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<emphasis id="B94BEAABFFEB8579FF76F7CDFEEEFA09" box="[199,283,1431,1457]" italics="true" pageId="5" pageNumber="536">Female</emphasis>
. Body length slightly shorter than male. Antenna as long as male. Protarsus slender. Spermatheca (
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<figureCitation id="13042A3CFFEB8579FADEF7CDFF2FFA6D" captionStart="FIGURE 3" captionStartId="6.[151,250,1617,1641]" captionTargetBox="[154,1433,185,1587]" captionTargetId="figure-20@6.[151,1436,181,1593]" captionTargetPageId="6" captionText="FIGURE 3. Scydmaenus (s. str.) chinensis (A–C. Male. D, E. Female). A. Dorsal habitus. B, C. Aedeagus, dorsal (B), and lateral (C). D, E. Spermatheca, dorsal (D) and lateral (E). Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.1 mm in D, E." figureDoi="http://doi.org/10.5281/zenodo.14734867" httpUri="https://zenodo.org/record/14734867/files/figure.png" pageId="5" pageNumber="536">Fig. 3D, E</figureCitation>
) spherical, with broad opening and long funnel-like structure located on same axis as insertion of accessory gland.
</paragraph>
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89
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(
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<figureCitation id="13042A3CFFED857FFF2EF285FF16FF41" box="[159,227,223,249]" captionStart="FIGURE 2" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[170,1417,181,1918]" captionTargetId="figure-20@4.[170,1417,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. Himaloconnus obesus sp. nov., male.A. Dorsal habitus. B. Head venter. C. Mesoventrite. D. Metaventral intercoxal process. E. Metatarsus. F–I. Aedeagus, lateral (F, H) and ventral (G, I). Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.1 mm in E–I." figureDoi="http://doi.org/10.5281/zenodo.14734863" httpUri="https://zenodo.org/record/14734863/files/figure.png" pageId="3" pageNumber="534">Fig. 2</figureCitation>
)
</paragraph>
</subSubSection>
@ -141,14 +144,14 @@ long, apex of median lobe with acute projection at middle, endophallus armature
<emphasis id="B94BEAABFFED857FFF76F0E9FEACFD75" bold="true" box="[199,345,691,717]" pageId="3" pageNumber="534">Description.</emphasis>
<emphasis id="B94BEAABFFED857FFED3F0E9FE69FD75" box="[354,412,691,717]" italics="true" pageId="3" pageNumber="534">Male</emphasis>
. Body (
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<figureCitation id="13042A3CFFED857FFE4AF0E9FDA3FD75" box="[507,598,691,717]" captionStart="FIGURE 2" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[170,1417,181,1918]" captionTargetId="figure-20@4.[170,1417,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. Himaloconnus obesus sp. nov., male.A. Dorsal habitus. B. Head venter. C. Mesoventrite. D. Metaventral intercoxal process. E. Metatarsus. F–I. Aedeagus, lateral (F, H) and ventral (G, I). Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.1 mm in E–I." figureDoi="http://doi.org/10.5281/zenodo.14734863" httpUri="https://zenodo.org/record/14734863/files/figure.png" pageId="3" pageNumber="534">Fig. 2A</figureCitation>
) length
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; head dark brown, pronotum, elytra, antennae and legs reddish-brown, mouthparts and tarsi paler in color. Vertex, sides of head and pronotum and humeri of elytra with long bristles; body dorsum almost impunctate, glabrous, with short and sparse setae found only on disc of elytra.
</paragraph>
<paragraph id="8B8036B9FFED857FFF76F145FF10FBC5" blockId="3.[151,1437,295,2014]" pageId="3" pageNumber="534">
Head (
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<figureCitation id="13042A3CFFED857FFEA4F145FE61FC81" box="[277,404,799,825]" captionStart="FIGURE 2" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[170,1417,181,1918]" captionTargetId="figure-20@4.[170,1417,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. Himaloconnus obesus sp. nov., male.A. Dorsal habitus. B. Head venter. C. Mesoventrite. D. Metaventral intercoxal process. E. Metatarsus. F–I. Aedeagus, lateral (F, H) and ventral (G, I). Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.1 mm in E–I." figureDoi="http://doi.org/10.5281/zenodo.14734863" httpUri="https://zenodo.org/record/14734863/files/figure.png" pageId="3" pageNumber="534">Fig. 2A, B</figureCitation>
) broadest at eyes, length
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, width across eyes
@ -175,14 +178,14 @@ Elytra together oval, broadest approximately at middle, length
</paragraph>
<paragraph id="8B8036B9FFED857FFF76F7D9FA7EFA79" blockId="3.[151,1437,295,2014]" pageId="3" pageNumber="534">
Mesoventrite (
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<figureCitation id="13042A3CFFED857FFEDAF7D9FE37FA25" box="[363,450,1411,1437]" captionStart="FIGURE 2" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[170,1417,181,1918]" captionTargetId="figure-20@4.[170,1417,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. Himaloconnus obesus sp. nov., male.A. Dorsal habitus. B. Head venter. C. Mesoventrite. D. Metaventral intercoxal process. E. Metatarsus. F–I. Aedeagus, lateral (F, H) and ventral (G, I). Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.1 mm in E–I." figureDoi="http://doi.org/10.5281/zenodo.14734863" httpUri="https://zenodo.org/record/14734863/files/figure.png" pageId="3" pageNumber="534">Fig. 2C</figureCitation>
) with broad intercoxal process; large, transversely oval area surrounding coxae greatly impressed and with setose fringes. Metaventrite with intercoxal process (
<figureCitation id="13042A3CFFED857FFC79F7FDFBD5FA79" box="[968,1056,1447,1473]" captionStart="FIGURE 2" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[170,1417,181,1918]" captionTargetId="figure-20@4.[170,1417,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. Himaloconnus obesus sp. nov., male.A. Dorsal habitus. B. Head venter. C. Mesoventrite. D. Metaventral intercoxal process. E. Metatarsus. F–I. Aedeagus, lateral (F, H) and ventral (G, I). Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.1 mm in E–I." pageId="3" pageNumber="534">Fig. 2D</figureCitation>
<figureCitation id="13042A3CFFED857FFC79F7FDFBD5FA79" box="[968,1056,1447,1473]" captionStart="FIGURE 2" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[170,1417,181,1918]" captionTargetId="figure-20@4.[170,1417,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. Himaloconnus obesus sp. nov., male.A. Dorsal habitus. B. Head venter. C. Mesoventrite. D. Metaventral intercoxal process. E. Metatarsus. F–I. Aedeagus, lateral (F, H) and ventral (G, I). Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.1 mm in E–I." figureDoi="http://doi.org/10.5281/zenodo.14734863" httpUri="https://zenodo.org/record/14734863/files/figure.png" pageId="3" pageNumber="534">Fig. 2D</figureCitation>
) formed by pair of acute spines.
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Legs moderately long and slender, unmodified; with small plantulae (
<figureCitation id="13042A3CFFED857FFC63F791FBD3FA5D" box="[978,1062,1483,1509]" captionStart="FIGURE 2" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[170,1417,181,1918]" captionTargetId="figure-20@4.[170,1417,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. Himaloconnus obesus sp. nov., male.A. Dorsal habitus. B. Head venter. C. Mesoventrite. D. Metaventral intercoxal process. E. Metatarsus. F–I. Aedeagus, lateral (F, H) and ventral (G, I). Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.1 mm in E–I." pageId="3" pageNumber="534">Fig. 2E</figureCitation>
<figureCitation id="13042A3CFFED857FFC63F791FBD3FA5D" box="[978,1062,1483,1509]" captionStart="FIGURE 2" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[170,1417,181,1918]" captionTargetId="figure-20@4.[170,1417,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. Himaloconnus obesus sp. nov., male.A. Dorsal habitus. B. Head venter. C. Mesoventrite. D. Metaventral intercoxal process. E. Metatarsus. F–I. Aedeagus, lateral (F, H) and ventral (G, I). Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.1 mm in E–I." figureDoi="http://doi.org/10.5281/zenodo.14734863" httpUri="https://zenodo.org/record/14734863/files/figure.png" pageId="3" pageNumber="534">Fig. 2E</figureCitation>
) between tarsal claws.
</paragraph>
<paragraph id="8B8036B9FFED857FFF76F7B5FECEF906" blockId="3.[151,1437,295,2014]" pageId="3" pageNumber="534">
@ -395,7 +398,7 @@ is distributed in
, while the others are found in the Himalayan region.
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<emphasis id="B94BEAABFFEA8578FF26F5CDFEE7F817" bold="true" box="[151,274,1943,1967]" pageId="4" pageNumber="535">FIGURE 2.</emphasis>
<taxonomicName id="4C3F4D3AFFEA8578FEA6F5CDFE00F817" authorityName="Yin &amp; Zhou" authorityYear="2025" box="[279,501,1943,1967]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="535" phylum="Arthropoda" rank="species" species="obesus" status="sp. nov.">

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@ -75,7 +78,7 @@ Chinese common name:
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<paragraph id="8B8036B9FFEF857DFF26F38AFF1EFE52" blockId="1.[151,829,391,490]" box="[151,235,464,490]" pageId="1" pageNumber="532">
(
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<figureCitation id="13042A3CFFEF857DFF2EF38AFF15FE52" box="[159,224,464,490]" captionStart="FIGURE 1" captionStartId="2.[151,250,1316,1340]" captionTargetBox="[180,1409,189,1287]" captionTargetId="figure-20@2.[177,1411,179,1289]" captionTargetPageId="2" captionText="FIGURE 1. Euconnus (s. str.) wumengshanus sp. nov. (A–E. Male. F. Female). A. Dorsal habitus. B–E. Aedeagus, lateral (B, D) and ventral (C, E). F. Spermatheca, lateral. Scale bars: 0.5 mm in A; 0.1 mm in B–E; 0.05 mm in F." figureDoi="http://doi.org/10.5281/zenodo.14734861" httpUri="https://zenodo.org/record/14734861/files/figure.png" pageId="1" pageNumber="532">Fig. 1</figureCitation>
)
</paragraph>
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@ -184,7 +187,7 @@ Body length approximately
<emphasis id="B94BEAABFFEF857DFF76F64AFEACFB92" bold="true" box="[199,345,1040,1066]" pageId="1" pageNumber="532">Description.</emphasis>
<emphasis id="B94BEAABFFEF857DFEEEF64AFE6CFB92" box="[351,409,1040,1066]" italics="true" pageId="1" pageNumber="532">Male</emphasis>
. Body (
<figureCitation id="13042A3CFFEF857DFE40F64AFDB2FB92" box="[497,583,1040,1066]" captionStart="FIGURE 1" captionStartId="2.[151,250,1316,1340]" captionTargetBox="[180,1409,189,1287]" captionTargetId="figure-20@2.[177,1411,179,1289]" captionTargetPageId="2" captionText="FIGURE 1. Euconnus (s. str.) wumengshanus sp. nov. (A–E. Male. F. Female). A. Dorsal habitus. B–E. Aedeagus, lateral (B, D) and ventral (C, E). F. Spermatheca, lateral. Scale bars: 0.5 mm in A; 0.1 mm in B–E; 0.05 mm in F." pageId="1" pageNumber="532">Fig. 1A</figureCitation>
<figureCitation id="13042A3CFFEF857DFE40F64AFDB2FB92" box="[497,583,1040,1066]" captionStart="FIGURE 1" captionStartId="2.[151,250,1316,1340]" captionTargetBox="[180,1409,189,1287]" captionTargetId="figure-20@2.[177,1411,179,1289]" captionTargetPageId="2" captionText="FIGURE 1. Euconnus (s. str.) wumengshanus sp. nov. (A–E. Male. F. Female). A. Dorsal habitus. B–E. Aedeagus, lateral (B, D) and ventral (C, E). F. Spermatheca, lateral. Scale bars: 0.5 mm in A; 0.1 mm in B–E; 0.05 mm in F." figureDoi="http://doi.org/10.5281/zenodo.14734861" httpUri="https://zenodo.org/record/14734861/files/figure.png" pageId="1" pageNumber="532">Fig. 1A</figureCitation>
) length
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; body uniformly reddish-brown, mouthparts and tarsi paler in color. Posterior margin of vertex, sides of head, and sides and disc of pronotum with scattered thick bristles, with long setae on body surface. Dorsum of body finely punctate.
@ -217,7 +220,7 @@ Elytra suboval and slightly flattened, broadest approximately at middle, length
<paragraph id="8B8036B9FFEF857DFF76F40BFDA0F9D2" blockId="1.[151,1437,536,2039]" box="[199,597,1616,1643]" pageId="1" pageNumber="532">Legs long and slender; unmodified.</paragraph>
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Aedeagus (
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<figureCitation id="13042A3CFFEF857DFEF6F42EFE4AF936" box="[327,447,1652,1678]" captionStart="FIGURE 1" captionStartId="2.[151,250,1316,1340]" captionTargetBox="[180,1409,189,1287]" captionTargetId="figure-20@2.[177,1411,179,1289]" captionTargetPageId="2" captionText="FIGURE 1. Euconnus (s. str.) wumengshanus sp. nov. (A–E. Male. F. Female). A. Dorsal habitus. B–E. Aedeagus, lateral (B, D) and ventral (C, E). F. Spermatheca, lateral. Scale bars: 0.5 mm in A; 0.1 mm in B–E; 0.05 mm in F." figureDoi="http://doi.org/10.5281/zenodo.14734861" httpUri="https://zenodo.org/record/14734861/files/figure.png" pageId="1" pageNumber="532">Fig. 1D–E</figureCitation>
) moderately stout, dorso-ventrally almost symmetric, length
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, in ventral view median lobe with abruptly delimited and long, broad apical projection greatly curved dorsally; compressor plate broad and plate-like in dorso-ventral view, subquadrate with narrowed apical margin, obliquely connected to median lobe in lateral view; lateral projections broad and membranous, curved dorsally, with round apices; ventral wall of median lobe with large, transversely rhomboidal plate, its apical margin with two admesal acute projection; parameres broad and elongate, narrowing from bases toward apices, each with two long setae at apex, two similar long setae at apical 2/5 and several short and thin setae at middle.
@ -227,7 +230,7 @@ Aedeagus (
. Similar to male in habitus and external morphological features. Antennal clubs (
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) slightly shorter than male. Spermatheca (
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) in lateral view transversely oval. Measurements (as for male): body length
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, length/width of head 0.34/
@ -242,7 +245,7 @@ Aedeagus (
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.
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<emphasis id="B94BEAABFFEC857EFF26F77EFEE6FA84" bold="true" box="[151,275,1316,1340]" pageId="2" pageNumber="533">FIGURE 1.</emphasis>
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<subSubSection id="C32B2CCCFFA8FF9BFF6BFD34FEFD608B" box="[151,373,672,699]" pageId="5" pageNumber="376" type="description">
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<figureCitation id="130A63C2FFA8FF9BFF6BFD34FE9F608A" box="[151,279,672,699]" captionStart-0="FIGURES 15–16" captionStart-1="FIGURES 17–22" captionStart-2="FIGURES 23–28" captionStartId-0="5.[151,264,1593,1617]" captionStartId-1="6.[151,264,1649,1673]" captionStartId-2="7.[151,264,1133,1157]" captionTargetBox-0="[151,1436,1154,1569]" captionTargetBox-1="[159,1427,245,1618]" captionTargetBox-2="[159,1433,389,1102]" captionTargetId-0="figure-355@5.[151,1436,1154,1569]" captionTargetId-1="figure-56@6.[151,1435,237,1626]" captionTargetId-2="figure-114@7.[151,1435,381,1110]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURES 15–16. Photographs of Tomocerus (Ocreatomurus) oppositus sp. nov. in ethanol (lateral view): 15, subadult; 16, fully grown adult." captionText-1="FIGURES 17–22. Tomocerus (Ocreatomurus) oppositus sp. nov.: 17, cephalic dorsal macrochaetotaxy; 18, dorsal macrochaetotaxy of Th. II—Abd. V (*macrochaeta present only occasionally); 19, tibiotarsi I, II and III (lateral view, arrows pointing to blunt chaetae); 20, distal end of hind tibiotarsus (outer view, showing size difference between accessory chaetae and pretarsal chaetae); 21, hind claw complex (lateral view, legends to chaetal types as in Fig. 8); 22, tenaculum (anterior view)." captionText-2="FIGURES 23–28. Tomocerus (Ocreatomurus) oppositus sp. nov.: 23, left side of manubrium (dorsal view); 24, distal area of manubrium (dorsal view, arrow pointing to distal corner chaeta); 25, proximal part of dens with basal spines arranged in single row (dorsal view); 26, basal dental spines arranged in double rows (dorsal view); 27, examples of dental spines; 28, left mucro (inner view)." figureDoi-0="http://doi.org/10.5281/zenodo.14735186" figureDoi-1="http://doi.org/10.5281/zenodo.14735188" figureDoi-2="http://doi.org/10.5281/zenodo.14735192" httpUri-0="https://zenodo.org/record/14735186/files/figure.png" httpUri-1="https://zenodo.org/record/14735188/files/figure.png" httpUri-2="https://zenodo.org/record/14735192/files/figure.png" pageId="5" pageNumber="376">Figs 15–28</figureCitation>
,
<tableCitation id="C6B34AFCFFA8FF9BFEDEFD34FEFD608B" box="[290,373,672,698]" captionStart="TABLE 1" captionStartId="5.[152,244,151,175]" captionText="TABLE 1. Main characters distinguishing Tomocerus (Ocreatomurus) spinulus Chen &amp; Christiansen, 1998 (sensu Yu et al. 2018), T. (O.) gyirongensis sp. nov. and T. (O.) oppositus sp. nov." pageId="5" pageNumber="376">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32B2CCCFFA8FF9BFF6BFD7CFA9F6187" pageId="5" pageNumber="376" type="materials_examined">
<paragraph id="8B8E7F47FFA8FF9BFF6BFD7CFA9F6187" blockId="5.[151,1437,744,1130]" pageId="5" pageNumber="376">
<emphasis id="B945A355FFA8FF9BFF6BFD7CFECB6133" bold="true" box="[151,323,744,770]" pageId="5" pageNumber="376">Type material.</emphasis>
<materialsCitation id="3B59751AFFA8FF9BFEB4FD7CFC10617B" collectingDate="2018-08-07" collectionCode="WGS" collectorName="Daoyuan Yu &amp; Yiming Wei &amp; Chunyan Qin" country="China" county="Gyirong County" elevation="2500" latitude="28.368" location="Gyirong Valley" longLatPrecision="1" longitude="85.363014" municipality="Autonomous Region" pageId="5" pageNumber="376" specimenCode="WGS84" specimenCount="1" stateProvince="Xizang" typeStatus="holotype">
<typeStatus id="548AC1E5FFA8FF9BFEB4FD7CFE3D6133" box="[328,437,744,770]" pageId="5" pageNumber="376" type="holotype">Holotype</typeStatus>
: male on slide, coniferous and broad-leaved mixed forest,
<location id="8EEE299CFFA8FF9BFBD3FD7CFB526133" LSID="urn:lsid:plazi:treatment:0398CE51FFA8FF96FF6BFDE8FD0862E4:8EEE299CFFA8FF9BFBD3FD7CFB526133" box="[1071,1242,744,770]" country="China" county="Gyirong County" latitude="28.368" longLatPrecision="1" longitude="85.363014" municipality="Autonomous Region" name="Gyirong Valley" pageId="5" pageNumber="376" stateProvince="Xizang">Gyirong Valley</location>
,
<collectingCounty id="62EF07CBFFA8FF9BFB1EFD7CFA116133" box="[1250,1433,744,770]" pageId="5" pageNumber="376">Gyirong County</collectingCounty>
,
<location id="8EEE299CFFA8FF9BFF6BFC98FF616117" LSID="urn:lsid:plazi:treatment:0398CE51FFA8FF96FF6BFDE8FD0862E4:8EEE299CFFA8FF9BFF6BFC98FF616117" box="[151,233,780,806]" country="China" county="Gyirong County" latitude="28.368" longLatPrecision="1" longitude="85.363014" municipality="Autonomous Region" name="Xigaze" pageId="5" pageNumber="376" stateProvince="Xizang">Xigaze</location>
,
<collectingRegion id="49F5B1A5FFA8FF9BFF04FC98FEC16117" box="[248,329,780,806]" country="China" name="Xizang" pageId="5" pageNumber="376">Xizang</collectingRegion>
<collectingMunicipality id="6BEAE53DFFA8FF9BFEADFC98FDC86117" box="[337,576,780,806]" pageId="5" pageNumber="376">Autonomous Region</collectingMunicipality>
,
<collectingCountry id="F3263FD7FFA8FF9BFDB2FC98FD1C6117" box="[590,660,780,806]" name="China" pageId="5" pageNumber="376">China</collectingCountry>
,
<geoCoordinate id="EE051980FFA8FF9BFD5EFC9AFCB86118" box="[674,816,782,809]" degrees="85" direction="east" minutes="21.781" orientation="longitude" pageId="5" pageNumber="376" precision="1" value="85.363014">85°21.781′E</geoCoordinate>
,
<geoCoordinate id="EE051980FFA8FF9BFCC3FC9AFC466118" box="[831,974,782,809]" degrees="28" direction="north" minutes="22.080" orientation="latitude" pageId="5" pageNumber="376" precision="1" value="28.368">28°22.080′N</geoCoordinate>
(
<specimenCode id="DB97D73CFFA8FF9BFC23FC98FBB76117" box="[991,1087,780,806]" collectionCode="WGS" pageId="5" pageNumber="376">WGS84</specimenCode>
), alt.
<quantity id="4CC9D2A2FFA8FF9BFB7FFC98FB526116" box="[1155,1242,780,807]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="376" unit="m" value="2500.0">
<elevation id="001C9874FFA8FF9BFB7FFC98FB526116" box="[1155,1242,780,807]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="376" unit="m" value="2500.0">2500 m</elevation>
</quantity>
,
<date id="FF8F5987FFA8FF9BFB15FC98FAED6117" box="[1257,1381,780,806]" pageId="5" pageNumber="376" value="2018-08-07">
<collectingDate id="EFCBA06FFFA8FF9BFB15FC98FAED6117" box="[1257,1381,780,806]" pageId="5" pageNumber="376" value="2018-08-07">7.viii.2018</collectingDate>
</date>
, leg.
<collectorName id="26C41A91FFA8FF9BFF6BFCA5FEAD617A" box="[151,293,817,843]" pageId="5" pageNumber="376">Daoyuan Yu</collectorName>
,
<collectorName id="26C41A91FFA8FF9BFECCFCA4FE3F617B" box="[304,439,816,842]" pageId="5" pageNumber="376">Yiming Wei</collectorName>
and
<collectorName id="26C41A91FFA8FF9BFE13FCA4FD0D617B" box="[495,645,816,842]" pageId="5" pageNumber="376">Chunyan Qin</collectorName>
(Sample code 18XZ16)
</materialsCitation>
.
<materialsCitation id="3B59751AFFA8FF9BFC58FCA5FDF5615F" collectingDate="2018-08-07" collectionCode="WGS" collectorName="Daoyuan Yu &amp; Yiming Wei &amp; Chunyan Qin" country="China" county="Gyirong County" elevation="2500" latitude="28.368" location="Gyirong Valley" longLatPrecision="1" longitude="85.363014" municipality="Autonomous Region" pageId="5" pageNumber="376" specimenCount="4" specimenCount-female="3" specimenCount-juvenile="1" stateProvince="Xizang" typeStatus="paratype">
<typeStatus id="548AC1E5FFA8FF9BFC58FCA5FB9D617A" box="[932,1045,817,843]" pageId="5" pageNumber="376" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D37B4CEFFA8FF9BFBDCFCA4FB3F617B" box="[1056,1207,816,842]" count="3" pageId="5" pageNumber="376" type="female">three females</specimenCount>
and
<specimenCount id="9D37B4CEFFA8FF9BFB12FCA5FAF0617B" box="[1262,1400,816,843]" count="1" pageId="5" pageNumber="376" type="juvenile">one juvenile</specimenCount>
on slides,
<quantity id="4CC9D2A2FFA8FF9BFF19FCC0FE95615F" box="[229,285,852,878]" metricMagnitude="-1" metricUnit="m" metricValue="7.62" pageId="5" pageNumber="376" unit="in" value="30.0">30 in</quantity>
ethanol, same data as holotype
</materialsCitation>
;
<materialsCitation id="3B59751AFFA8FF9BFD74FCC1FA9B6187" collectingDate="2018-08-07" collectionCode="NJAU" collectorName="Daoyuan Yu &amp; Yiming Wei &amp; Chunyan Qin &amp; All" country="China" county="Gyirong County" elevation="2765" latitude="28.375383" location="Jipu Village" longLatPrecision="1" longitude="85.32822" municipality="Autonomous Region" pageId="5" pageNumber="376" specimenCount="1" specimenCount-female="1" stateProvince="Xizang" typeStatus="paratype">
<specimenCount id="9D37B4CEFFA8FF9BFD74FCC1FC8B615F" box="[648,771,852,879]" count="1" pageId="5" pageNumber="376" type="female">one female</specimenCount>
on slide,
<quantity id="4CC9D2A2FFA8FF9BFC93FCC0FC20615F" box="[879,936,852,878]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="5" pageNumber="376" unit="in" value="18.0">18 in</quantity>
ethanol, under a wood board in
<location id="8EEE299CFFA8FF9BFAECFCC0FA10615F" LSID="urn:lsid:plazi:treatment:0398CE51FFA8FF96FF6BFDE8FD0862E4:8EEE299CFFA8FF9BFAECFCC0FA10615F" box="[1296,1432,852,878]" country="China" county="Gyirong County" latitude="28.375383" longLatPrecision="1" longitude="85.32822" municipality="Autonomous Region" name="Jipu Village" pageId="5" pageNumber="376" stateProvince="Xizang">Jipu Village</location>
,
<collectingCounty id="62EF07CBFFA8FF9BFF6BFCECFED861A3" box="[151,336,888,914]" pageId="5" pageNumber="376">Gyirong County</collectingCounty>
,
<location id="8EEE299CFFA8FF9BFEA7FCECFE2561A3" LSID="urn:lsid:plazi:treatment:0398CE51FFA8FF96FF6BFDE8FD0862E4:8EEE299CFFA8FF9BFEA7FCECFE2561A3" box="[347,429,888,914]" country="China" county="Gyirong County" latitude="28.375383" longLatPrecision="1" longitude="85.32822" municipality="Autonomous Region" name="Xigaze" pageId="5" pageNumber="376" stateProvince="Xizang">Xigaze</location>
,
<collectingRegion id="49F5B1A5FFA8FF9BFE45FCECFD8261A3" box="[441,522,888,914]" country="China" name="Xizang" pageId="5" pageNumber="376">Xizang</collectingRegion>
<collectingMunicipality id="6BEAE53DFFA8FF9BFDF3FCECFD7361A3" box="[527,763,888,914]" pageId="5" pageNumber="376">Autonomous Region</collectingMunicipality>
,
<collectingCountry id="F3263FD7FFA8FF9BFCFAFCECFCC461A3" box="[774,844,888,914]" name="China" pageId="5" pageNumber="376">China</collectingCountry>
,
<geoCoordinate id="EE051980FFA8FF9BFCA4FCEEFC6D61A4" box="[856,997,890,917]" degrees="85" direction="east" minutes="19.693" orientation="longitude" pageId="5" pageNumber="376" precision="1" value="85.32822">85°19.693′E</geoCoordinate>
,
<geoCoordinate id="EE051980FFA8FF9BFC0DFCEEFB0B61A4" box="[1009,1155,890,917]" degrees="28" direction="north" minutes="22.523" orientation="latitude" pageId="5" pageNumber="376" precision="1" value="28.375383">28°22.523′N</geoCoordinate>
, alt.
<quantity id="4CC9D2A2FFA8FF9BFB45FCECFA9861A2" box="[1209,1296,888,915]" metricMagnitude="3" metricUnit="m" metricValue="2.7649999999999997" pageId="5" pageNumber="376" unit="m" value="2765.0">
<elevation id="001C9874FFA8FF9BFB45FCECFA9861A2" box="[1209,1296,888,915]" metricMagnitude="3" metricUnit="m" metricValue="2.7649999999999997" pageId="5" pageNumber="376" unit="m" value="2765.0">2765 m</elevation>
</quantity>
,
<date id="FF8F5987FFA8FF9BFAE0FCECFA1061A3" box="[1308,1432,888,914]" pageId="5" pageNumber="376" value="2018-08-07">
<collectingDate id="EFCBA06FFFA8FF9BFAE0FCECFA1061A3" box="[1308,1432,888,914]" pageId="5" pageNumber="376" value="2018-08-07">7.viii.2018</collectingDate>
</date>
, leg.
<collectorName id="26C41A91FFA8FF9BFF3BFC09FEDC6186" box="[199,340,925,951]" pageId="5" pageNumber="376">Daoyuan Yu</collectorName>
,
<collectorName id="26C41A91FFA8FF9BFEA2FC08FE6D6187" box="[350,485,924,950]" pageId="5" pageNumber="376">Yiming Wei</collectorName>
and
<collectorName id="26C41A91FFA8FF9BFDE0FC08FD3A6187" box="[540,690,924,950]" pageId="5" pageNumber="376">Chunyan Qin</collectorName>
(Sample code 18XZ15).
<collectorName id="26C41A91FFA8FF9BFC30FC08FC786187" box="[972,1008,924,950]" pageId="5" pageNumber="376">All</collectorName>
types deposited in
<collectionCode id="ED20E782FFA8FF9BFB34FC08FA9B6187" box="[1224,1299,924,950]" pageId="5" pageNumber="376">NJAU</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32B2CCCFFA8FF99FF3BFC54FE886354" lastPageId="7" lastPageNumber="378" pageId="5" pageNumber="376" type="description">
<paragraph id="8B8E7F47FFA8FF9BFF3BFC54FEE2665B" blockId="5.[151,1437,744,1130]" pageId="5" pageNumber="376">
<emphasis id="B945A355FFA8FF9BFF3BFC54FED161EB" bold="true" box="[199,345,960,986]" pageId="5" pageNumber="376">Description.</emphasis>
Body length
<quantity id="4CC9D2A2FFA8FF9BFE12FC54FDFD61EA" box="[494,629,960,987]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="5.1" metricValueMin="3.4" pageId="5" pageNumber="376" unit="mm" value="4.25" valueMax="5.1" valueMin="3.4">3.4–5.1 mm</quantity>
(average 4.3,
<specimenCount id="9D37B4CEFFA8FF9BFCEDFC54FC3561EB" box="[785,957,960,986]" count="5" pageId="5" pageNumber="376" type="generic">five specimens</specimenCount>
). In subadults and small adults, head, body, and coxae deep purple; trochanteres, ventral tube, and manubrium with diffuse purple pigment; Ant. I–II, femora, tibiotarsi, and dens pale or bright yellow; Ant. III basally pale and gradually darker towards apex; Ant. IV purple (
<figureCitation id="130A63C2FFA8FF9BFF5CFBB8FF7B6676" box="[160,243,1068,1095]" captionStart="FIGURES 15–16" captionStartId="5.[151,264,1593,1617]" captionTargetBox="[151,1436,1154,1569]" captionTargetId="figure-355@5.[151,1436,1154,1569]" captionTargetPageId="5" captionText="FIGURES 15–16. Photographs of Tomocerus (Ocreatomurus) oppositus sp. nov. in ethanol (lateral view): 15, subadult; 16, fully grown adult." figureDoi="http://doi.org/10.5281/zenodo.14735186" httpUri="https://zenodo.org/record/14735186/files/figure.png" pageId="5" pageNumber="376">Fig. 15</figureCitation>
). In large adults, purple pigment more intense, becoming dark brown, and extending to Ant. I–II, femora, and dens (
<figureCitation id="130A63C2FFA8FF9BFEF5FBC4FED5665B" box="[265,349,1104,1130]" captionStart="FIGURES 15–16" captionStartId="5.[151,264,1593,1617]" captionTargetBox="[151,1436,1154,1569]" captionTargetId="figure-355@5.[151,1436,1154,1569]" captionTargetPageId="5" captionText="FIGURES 15–16. Photographs of Tomocerus (Ocreatomurus) oppositus sp. nov. in ethanol (lateral view): 15, subadult; 16, fully grown adult." figureDoi="http://doi.org/10.5281/zenodo.14735186" httpUri="https://zenodo.org/record/14735186/files/figure.png" pageId="5" pageNumber="376">Fig. 16</figureCitation>
).
</paragraph>
<caption id="DF4E2FCFFFA8FF9BFF6BF9ADFEC66444" ID-DOI="http://doi.org/10.5281/zenodo.14735186" ID-Zenodo-Dep="14735186" httpUri="https://zenodo.org/record/14735186/files/figure.png" pageId="5" pageNumber="376" startId="5.[151,264,1593,1617]" targetBox="[151,1436,1154,1569]" targetPageId="5" targetType="figure">
<paragraph id="8B8E7F47FFA8FF9BFF6BF9ADFEC66444" blockId="5.[151,1436,1593,1654]" pageId="5" pageNumber="376">
<emphasis id="B945A355FFA8FF9BFF6BF9ADFEDE6460" bold="true" box="[151,342,1593,1617]" pageId="5" pageNumber="376">FIGURES 15–16.</emphasis>
Photographs of
<taxonomicName id="4C3104C4FFA8FF9BFDFEF9AEFC0C6460" authorityName="Yu &amp; Xue &amp; Zhang" authorityYear="2025" box="[514,900,1593,1617]" class="Collembola" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Entomobryomorpha" pageId="5" pageNumber="376" phylum="Arthropoda" rank="species" species="oppositus" status="sp. nov." subGenus="Ocreatomurus">
<emphasis id="B945A355FFA8FF9BFDFEF9AEFDF86460" box="[514,624,1594,1617]" italics="true" pageId="5" pageNumber="376">Tomocerus</emphasis>
(
<emphasis id="B945A355FFA8FF9BFD7DF9AEFC9A6460" box="[641,786,1594,1617]" italics="true" pageId="5" pageNumber="376">Ocreatomurus</emphasis>
)
<emphasis id="B945A355FFA8FF9BFCDFF9AEFC0C6460" box="[803,900,1594,1617]" italics="true" pageId="5" pageNumber="376">oppositus</emphasis>
</taxonomicName>
<emphasis id="B945A355FFA8FF9BFC71F9ADFC576460" bold="true" box="[909,991,1593,1617]" pageId="5" pageNumber="376">
<taxonomicNameLabel id="A2761E2EFFA8FF9BFC71F9ADFC576460" box="[909,991,1593,1617]" pageId="5" pageNumber="376" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in ethanol (lateral view):
<emphasis id="B945A355FFA8FF9BFB0CF9ADFA816460" bold="true" box="[1264,1289,1593,1617]" pageId="5" pageNumber="376">15</emphasis>
, subadult;
<emphasis id="B945A355FFA8FF9BFA81F9ADFA1E6460" bold="true" box="[1405,1430,1593,1617]" pageId="5" pageNumber="376">16</emphasis>
, fully grown adult.
</paragraph>
</caption>
<paragraph id="8B8E7F47FFA8FF9BFF3BF95CFB0D6518" blockId="5.[151,1436,1735,2014]" pageId="5" pageNumber="376">
Intact four-segmented antennae 0.7–0.8 × length of body (average 0.75,
<specimenCount id="9D37B4CEFFA8FF9BFBF9F953FB4864D0" box="[1029,1216,1735,1761]" count="3" pageId="5" pageNumber="376" type="generic">three specimens</specimenCount>
). Length ratio of A nt I: II: III: IV = 1.0: 1.4–1.6: 7.3–8.5: 1.0–1.6. Ant. III occasionally with 1–2 scales at base. Cephalic dorsal macrochaetotaxy as in
<figureCitation id="130A63C2FFA8FF9BFE65F89BFE63651B" box="[409,491,1807,1834]" captionStart="FIGURES 17–22" captionStartId="6.[151,264,1649,1673]" captionTargetBox="[159,1427,245,1618]" captionTargetId="figure-56@6.[151,1435,237,1626]" captionTargetPageId="6" captionText="FIGURES 17–22. Tomocerus (Ocreatomurus) oppositus sp. nov.: 17, cephalic dorsal macrochaetotaxy; 18, dorsal macrochaetotaxy of Th. II—Abd. V (*macrochaeta present only occasionally); 19, tibiotarsi I, II and III (lateral view, arrows pointing to blunt chaetae); 20, distal end of hind tibiotarsus (outer view, showing size difference between accessory chaetae and pretarsal chaetae); 21, hind claw complex (lateral view, legends to chaetal types as in Fig. 8); 22, tenaculum (anterior view)." figureDoi="http://doi.org/10.5281/zenodo.14735188" httpUri="https://zenodo.org/record/14735188/files/figure.png" pageId="5" pageNumber="376">Fig. 17</figureCitation>
. Posterior margin of head with 35–45 chaetae on each side.
</paragraph>
<paragraph id="8B8E7F47FFA8FF9BFF3BF8A0FD5A657F" blockId="5.[151,1436,1735,2014]" box="[199,722,1843,1870]" pageId="5" pageNumber="376">
Pattern of body macrochaetotaxy as in
<figureCitation id="130A63C2FFA8FF9BFD80F8A7FD47657F" box="[636,719,1843,1870]" captionStart="FIGURES 17–22" captionStartId="6.[151,264,1649,1673]" captionTargetBox="[159,1427,245,1618]" captionTargetId="figure-56@6.[151,1435,237,1626]" captionTargetPageId="6" captionText="FIGURES 17–22. Tomocerus (Ocreatomurus) oppositus sp. nov.: 17, cephalic dorsal macrochaetotaxy; 18, dorsal macrochaetotaxy of Th. II—Abd. V (*macrochaeta present only occasionally); 19, tibiotarsi I, II and III (lateral view, arrows pointing to blunt chaetae); 20, distal end of hind tibiotarsus (outer view, showing size difference between accessory chaetae and pretarsal chaetae); 21, hind claw complex (lateral view, legends to chaetal types as in Fig. 8); 22, tenaculum (anterior view)." figureDoi="http://doi.org/10.5281/zenodo.14735188" httpUri="https://zenodo.org/record/14735188/files/figure.png" pageId="5" pageNumber="376">Fig. 18</figureCitation>
.
</paragraph>
<paragraph id="8B8E7F47FFA8FF98FF3BF8C3FDAD62E4" blockId="5.[151,1436,1735,2014]" lastBlockId="6.[151,1436,150,213]" lastPageId="6" lastPageNumber="377" pageId="5" pageNumber="376">
Tibiotarsi I, II, III ventrally with 8–9, 8–10, 6–10 strong chaetae, 5–6, 6–8, 6–9 of them blunt and spine-like, respectively (
<figureCitation id="130A63C2FFA8FF9BFED1F8EFFE0865A7" box="[301,384,1915,1942]" captionStart="FIGURES 17–22" captionStartId="6.[151,264,1649,1673]" captionTargetBox="[159,1427,245,1618]" captionTargetId="figure-56@6.[151,1435,237,1626]" captionTargetPageId="6" captionText="FIGURES 17–22. Tomocerus (Ocreatomurus) oppositus sp. nov.: 17, cephalic dorsal macrochaetotaxy; 18, dorsal macrochaetotaxy of Th. II—Abd. V (*macrochaeta present only occasionally); 19, tibiotarsi I, II and III (lateral view, arrows pointing to blunt chaetae); 20, distal end of hind tibiotarsus (outer view, showing size difference between accessory chaetae and pretarsal chaetae); 21, hind claw complex (lateral view, legends to chaetal types as in Fig. 8); 22, tenaculum (anterior view)." figureDoi="http://doi.org/10.5281/zenodo.14735188" httpUri="https://zenodo.org/record/14735188/files/figure.png" pageId="5" pageNumber="376">Fig. 19</figureCitation>
). Tenent hair 0.79–0.86 × the length of inner edge of unguis; accessory chaetae on tibiotarsus I equal to each other and shorter than pretarsal chaetae; on tibiotarsi II and III, anterior accessory chaeta subequal to or slightly longer than pretarsal chaetae, while posterior accessory chaeta shorter than pretarsal chaetae (
<figureCitation id="130A63C2FFA8FF9BFB0AF857FAC265EF" box="[1270,1354,1987,2014]" captionStart="FIGURES 17–22" captionStartId="6.[151,264,1649,1673]" captionTargetBox="[159,1427,245,1618]" captionTargetId="figure-56@6.[151,1435,237,1626]" captionTargetPageId="6" captionText="FIGURES 17–22. Tomocerus (Ocreatomurus) oppositus sp. nov.: 17, cephalic dorsal macrochaetotaxy; 18, dorsal macrochaetotaxy of Th. II—Abd. V (*macrochaeta present only occasionally); 19, tibiotarsi I, II and III (lateral view, arrows pointing to blunt chaetae); 20, distal end of hind tibiotarsus (outer view, showing size difference between accessory chaetae and pretarsal chaetae); 21, hind claw complex (lateral view, legends to chaetal types as in Fig. 8); 22, tenaculum (anterior view)." figureDoi="http://doi.org/10.5281/zenodo.14735188" httpUri="https://zenodo.org/record/14735188/files/figure.png" pageId="5" pageNumber="376">Fig. 20</figureCitation>
); guard chaetae as long as tenent hair. Unguiculus about 0.57–0.65 × the length of unguis (average 0.61,
<specimenCount id="9D37B4CEFFABFF98FB3EFF03FAF86280" box="[1218,1392,151,177]" count="4" pageId="6" pageNumber="377" type="generic">four specimens</specimenCount>
), its inner edge with one tooth (
<figureCitation id="130A63C2FFABFF98FE39FF2FFD9062E4" box="[453,536,187,213]" captionStart="FIGURES 17–22" captionStartId="6.[151,264,1649,1673]" captionTargetBox="[159,1427,245,1618]" captionTargetId="figure-56@6.[151,1435,237,1626]" captionTargetPageId="6" captionText="FIGURES 17–22. Tomocerus (Ocreatomurus) oppositus sp. nov.: 17, cephalic dorsal macrochaetotaxy; 18, dorsal macrochaetotaxy of Th. II—Abd. V (*macrochaeta present only occasionally); 19, tibiotarsi I, II and III (lateral view, arrows pointing to blunt chaetae); 20, distal end of hind tibiotarsus (outer view, showing size difference between accessory chaetae and pretarsal chaetae); 21, hind claw complex (lateral view, legends to chaetal types as in Fig. 8); 22, tenaculum (anterior view)." figureDoi="http://doi.org/10.5281/zenodo.14735188" httpUri="https://zenodo.org/record/14735188/files/figure.png" pageId="6" pageNumber="377">Fig. 21</figureCitation>
).
</paragraph>
<caption id="DF4E2FCFFFABFF98FF6BF9E5FA0864C7" ID-DOI="http://doi.org/10.5281/zenodo.14735188" ID-Zenodo-Dep="14735188" httpUri="https://zenodo.org/record/14735188/files/figure.png" pageId="6" pageNumber="377" startId="6.[151,264,1649,1673]" targetBox="[159,1427,245,1618]" targetPageId="6" targetType="figure">
<paragraph id="8B8E7F47FFABFF98FF6BF9E5FA0864C7" blockId="6.[151,1437,1649,1782]" pageId="6" pageNumber="377">
<emphasis id="B945A355FFABFF98FF6BF9E5FED664B8" bold="true" box="[151,350,1649,1673]" pageId="6" pageNumber="377">FIGURES 17–22.</emphasis>
<taxonomicName id="4C3104C4FFABFF98FE93F9E6FC8A64B8" authorityName="Yu &amp; Xue &amp; Zhang" authorityYear="2025" box="[367,770,1650,1674]" class="Collembola" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Entomobryomorpha" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="oppositus" status="sp. nov." subGenus="Ocreatomurus">
<emphasis id="B945A355FFABFF98FE93F9E6FE5564B8" box="[367,477,1650,1673]" italics="true" pageId="6" pageNumber="377">Tomocerus</emphasis>
(
<emphasis id="B945A355FFABFF98FE0AF9E6FD0F64B8" box="[502,647,1650,1673]" italics="true" pageId="6" pageNumber="377">Ocreatomurus</emphasis>
)
<emphasis id="B945A355FFABFF98FD5DF9E6FC8A64B8" box="[673,770,1650,1673]" italics="true" pageId="6" pageNumber="377">oppositus</emphasis>
</taxonomicName>
<emphasis id="B945A355FFABFF98FCEEF9E6FCE564BB" bold="true" box="[786,877,1650,1674]" pageId="6" pageNumber="377">
<taxonomicNameLabel id="A2761E2EFFABFF98FCEEF9E6FCE564BB" box="[786,877,1650,1674]" pageId="6" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
:
<emphasis id="B945A355FFABFF98FC79F9E5FC1664B8" bold="true" box="[901,926,1649,1673]" pageId="6" pageNumber="377">17</emphasis>
, cephalic dorsal macrochaetotaxy;
<emphasis id="B945A355FFABFF98FAD2F9E5FACF64B8" bold="true" box="[1326,1351,1649,1673]" pageId="6" pageNumber="377">18</emphasis>
, dorsal macrochaetotaxy of Th. II—Abd. V (*macrochaeta present only occasionally);
<emphasis id="B945A355FFABFF98FC3AF901FC57649C" bold="true" box="[966,991,1685,1709]" pageId="6" pageNumber="377">19</emphasis>
, tibiotarsi I, II and III (lateral view, arrows pointing to blunt chaetae);
<emphasis id="B945A355FFABFF98FE5AF92DFE3764E0" bold="true" box="[422,447,1721,1745]" pageId="6" pageNumber="377">20</emphasis>
, distal end of hind tibiotarsus (outer view, showing size difference between accessory chaetae and pretarsal chaetae);
<emphasis id="B945A355FFABFF98FEAAF949FEE764C4" bold="true" box="[342,367,1757,1781]" pageId="6" pageNumber="377">21</emphasis>
, hind claw complex (lateral view, legends to chaetal types as in Fig. 8);
<emphasis id="B945A355FFABFF98FBB0F949FBED64C4" bold="true" box="[1100,1125,1757,1781]" pageId="6" pageNumber="377">22</emphasis>
, tenaculum (anterior view).
</paragraph>
</caption>
<paragraph id="8B8E7F47FFABFF99FF3BF8DCFE886354" blockId="6.[151,1437,1864,2034]" lastBlockId="7.[151,1436,151,357]" lastPageId="7" lastPageNumber="378" pageId="6" pageNumber="377">
Ventral tube scaled on both faces; anterior face with 35–55 (average 46,
<specimenCount id="9D37B4CEFFABFF98FBDFF8DCFB586553" box="[1059,1232,1864,1890]" count="5" pageId="6" pageNumber="377" type="generic">five specimens</specimenCount>
) chaetae on each side; posterior face with 75–100 (average 89,
<specimenCount id="9D37B4CEFFABFF98FD66F8F8FCCB65B7" box="[666,835,1900,1926]" count="5" pageId="6" pageNumber="377" type="generic">five specimens</specimenCount>
) chaetae; each lateral flap with 90–180 (average 144,
<specimenCount id="9D37B4CEFFABFF98FF6BF804FECB659B" box="[151,323,1936,1962]" count="5" pageId="6" pageNumber="377" type="generic">five specimens</specimenCount>
) chaetae and occasionally 1–3 scales. Anterior face of tenaculum with 10–17 (average 13,
<specimenCount id="9D37B4CEFFABFF98FA8DF804FE8565FF" count="5" pageId="6" pageNumber="377" type="generic">five specimens</specimenCount>
) chaetae and 0–3 scales (
<figureCitation id="130A63C2FFABFF98FDD9F820FDF265FF" box="[549,634,1972,1998]" captionStart="FIGURES 17–22" captionStartId="6.[151,264,1649,1673]" captionTargetBox="[159,1427,245,1618]" captionTargetId="figure-56@6.[151,1435,237,1626]" captionTargetPageId="6" captionText="FIGURES 17–22. Tomocerus (Ocreatomurus) oppositus sp. nov.: 17, cephalic dorsal macrochaetotaxy; 18, dorsal macrochaetotaxy of Th. II—Abd. V (*macrochaeta present only occasionally); 19, tibiotarsi I, II and III (lateral view, arrows pointing to blunt chaetae); 20, distal end of hind tibiotarsus (outer view, showing size difference between accessory chaetae and pretarsal chaetae); 21, hind claw complex (lateral view, legends to chaetal types as in Fig. 8); 22, tenaculum (anterior view)." figureDoi="http://doi.org/10.5281/zenodo.14735188" httpUri="https://zenodo.org/record/14735188/files/figure.png" pageId="6" pageNumber="377">Fig. 22</figureCitation>
). Ratio manubrium: dens: mucro = 3.4–4.1: 4.2–5.2: 1.0. Manubrium laterally with 7–11 strong chaetae; dorsally with 170–280 (average 237,
<specimenCount id="9D37B4CEFFABFF98FC36F84CFBFC65C3" box="[970,1140,2008,2034]" count="5" pageId="6" pageNumber="377" type="generic">five specimens</specimenCount>
) chaetae of different sizes on each chaetal strip, 12–21 (average 16,
<specimenCount id="9D37B4CEFFAAFF99FD85FF03FCAC6280" box="[633,804,151,177]" count="5" pageId="7" pageNumber="378" type="generic">five specimens</specimenCount>
) pseudopores almost arranging in a file and scale strip running from base to about 2/3 length of manubrium (
<figureCitation id="130A63C2FFAAFF99FCFEFF2FFCDE62E4" box="[770,854,187,213]" captionStart="FIGURES 23–28" captionStartId="7.[151,264,1133,1157]" captionTargetBox="[159,1433,389,1102]" captionTargetId="figure-114@7.[151,1435,381,1110]" captionTargetPageId="7" captionText="FIGURES 23–28. Tomocerus (Ocreatomurus) oppositus sp. nov.: 23, left side of manubrium (dorsal view); 24, distal area of manubrium (dorsal view, arrow pointing to distal corner chaeta); 25, proximal part of dens with basal spines arranged in single row (dorsal view); 26, basal dental spines arranged in double rows (dorsal view); 27, examples of dental spines; 28, left mucro (inner view)." figureDoi="http://doi.org/10.5281/zenodo.14735192" httpUri="https://zenodo.org/record/14735192/files/figure.png" pageId="7" pageNumber="378">Fig. 23</figureCitation>
). External distal corner chaeta as large as medium-sized mesochaetae in chaetal strip (
<figureCitation id="130A63C2FFAAFF99FDDFFF4BFDFF62C8" box="[547,631,223,249]" captionStart="FIGURES 23–28" captionStartId="7.[151,264,1133,1157]" captionTargetBox="[159,1433,389,1102]" captionTargetId="figure-114@7.[151,1435,381,1110]" captionTargetPageId="7" captionText="FIGURES 23–28. Tomocerus (Ocreatomurus) oppositus sp. nov.: 23, left side of manubrium (dorsal view); 24, distal area of manubrium (dorsal view, arrow pointing to distal corner chaeta); 25, proximal part of dens with basal spines arranged in single row (dorsal view); 26, basal dental spines arranged in double rows (dorsal view); 27, examples of dental spines; 28, left mucro (inner view)." figureDoi="http://doi.org/10.5281/zenodo.14735192" httpUri="https://zenodo.org/record/14735192/files/figure.png" pageId="7" pageNumber="378">Fig. 24</figureCitation>
). Dental spine formula
<emphasis id="B945A355FFAAFF99FC82FF4BFC3462C9" box="[894,956,223,248]" italics="true" pageId="7" pageNumber="378">sensu</emphasis>
<bibRefCitation id="EFA002B6FFAAFF99FC38FF4BFBE262C8" author="Folsom, J. W." box="[964,1130,223,249]" pageId="7" pageNumber="378" pagination="451 - 472" refId="ref5155" refString="Folsom, J. W. (1913) North American spring-tails of the sub-family Tomocerinae. Proceedings of the United States National Museum, 46, 451 - 472. https: // doi. org / 10.5479 / si. 00963801.46 - 2037.451" type="journal article" year="1913">Folsom (1913)</bibRefCitation>
as 4–6/4–8,
<emphasis id="B945A355FFAAFF99FB05FF4AFA8F62C9" bold="true" box="[1273,1287,222,248]" pageId="7" pageNumber="378">1</emphasis>
, basal spines arranged either in one row (
<figureCitation id="130A63C2FFAAFF99FE2EFE97FDAB632C" box="[466,547,258,285]" captionStart="FIGURES 23–28" captionStartId="7.[151,264,1133,1157]" captionTargetBox="[159,1433,389,1102]" captionTargetId="figure-114@7.[151,1435,381,1110]" captionTargetPageId="7" captionText="FIGURES 23–28. Tomocerus (Ocreatomurus) oppositus sp. nov.: 23, left side of manubrium (dorsal view); 24, distal area of manubrium (dorsal view, arrow pointing to distal corner chaeta); 25, proximal part of dens with basal spines arranged in single row (dorsal view); 26, basal dental spines arranged in double rows (dorsal view); 27, examples of dental spines; 28, left mucro (inner view)." figureDoi="http://doi.org/10.5281/zenodo.14735192" httpUri="https://zenodo.org/record/14735192/files/figure.png" pageId="7" pageNumber="378">Fig. 25</figureCitation>
) or two rows (
<figureCitation id="130A63C2FFAAFF99FD36FE97FC97632C" box="[714,799,259,285]" captionStart="FIGURES 23–28" captionStartId="7.[151,264,1133,1157]" captionTargetBox="[159,1433,389,1102]" captionTargetId="figure-114@7.[151,1435,381,1110]" captionTargetPageId="7" captionText="FIGURES 23–28. Tomocerus (Ocreatomurus) oppositus sp. nov.: 23, left side of manubrium (dorsal view); 24, distal area of manubrium (dorsal view, arrow pointing to distal corner chaeta); 25, proximal part of dens with basal spines arranged in single row (dorsal view); 26, basal dental spines arranged in double rows (dorsal view); 27, examples of dental spines; 28, left mucro (inner view)." figureDoi="http://doi.org/10.5281/zenodo.14735192" httpUri="https://zenodo.org/record/14735192/files/figure.png" pageId="7" pageNumber="378">Fig. 26</figureCitation>
). Denticles on dental spines distributed more densely on ventral side than on dorsal side of spine (
<figureCitation id="130A63C2FFAAFF99FD9EFEB3FD3F6370" box="[610,695,295,321]" captionStart="FIGURES 23–28" captionStartId="7.[151,264,1133,1157]" captionTargetBox="[159,1433,389,1102]" captionTargetId="figure-114@7.[151,1435,381,1110]" captionTargetPageId="7" captionText="FIGURES 23–28. Tomocerus (Ocreatomurus) oppositus sp. nov.: 23, left side of manubrium (dorsal view); 24, distal area of manubrium (dorsal view, arrow pointing to distal corner chaeta); 25, proximal part of dens with basal spines arranged in single row (dorsal view); 26, basal dental spines arranged in double rows (dorsal view); 27, examples of dental spines; 28, left mucro (inner view)." figureDoi="http://doi.org/10.5281/zenodo.14735192" httpUri="https://zenodo.org/record/14735192/files/figure.png" pageId="7" pageNumber="378">Fig. 27</figureCitation>
). Mucro with 4–7 (average six,
<specimenCount id="9D37B4CEFFAAFF99FBE7FEB3FB4C6370" box="[1051,1220,295,321]" count="5" pageId="7" pageNumber="378" type="generic">five specimens</specimenCount>
) intermediate teeth (
<figureCitation id="130A63C2FFAAFF99FF5CFEDFFF7C6354" box="[160,244,331,357]" captionStart="FIGURES 23–28" captionStartId="7.[151,264,1133,1157]" captionTargetBox="[159,1433,389,1102]" captionTargetId="figure-114@7.[151,1435,381,1110]" captionTargetPageId="7" captionText="FIGURES 23–28. Tomocerus (Ocreatomurus) oppositus sp. nov.: 23, left side of manubrium (dorsal view); 24, distal area of manubrium (dorsal view, arrow pointing to distal corner chaeta); 25, proximal part of dens with basal spines arranged in single row (dorsal view); 26, basal dental spines arranged in double rows (dorsal view); 27, examples of dental spines; 28, left mucro (inner view)." figureDoi="http://doi.org/10.5281/zenodo.14735192" httpUri="https://zenodo.org/record/14735192/files/figure.png" pageId="7" pageNumber="378">Fig. 28</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF4E2FCFFFAAFF99FF6BFBF9FE9366C3" ID-DOI="http://doi.org/10.5281/zenodo.14735192" ID-Zenodo-Dep="14735192" httpUri="https://zenodo.org/record/14735192/files/figure.png" pageId="7" pageNumber="378" startId="7.[151,264,1133,1157]" targetBox="[159,1433,389,1102]" targetPageId="7" targetType="figure">
<paragraph id="8B8E7F47FFAAFF99FF6BFBF9FE9366C3" blockId="7.[151,1437,1133,1266]" pageId="7" pageNumber="378">
<emphasis id="B945A355FFAAFF99FF6BFBF9FEDD66B7" bold="true" box="[151,341,1133,1158]" pageId="7" pageNumber="378">FIGURES 23–28.</emphasis>
<taxonomicName id="4C3104C4FFAAFF99FEA0FBFAFD5466B4" authorityName="Yu &amp; Xue &amp; Zhang" authorityYear="2025" box="[348,732,1134,1158]" class="Collembola" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Entomobryomorpha" pageId="7" pageNumber="378" phylum="Arthropoda" rank="species" species="oppositus" status="sp. nov." subGenus="Ocreatomurus">
<emphasis id="B945A355FFAAFF99FEA0FBFAFE4266B4" box="[348,458,1134,1157]" italics="true" pageId="7" pageNumber="378">Tomocerus</emphasis>
(
<emphasis id="B945A355FFAAFF99FE26FBFAFDE366B4" box="[474,619,1134,1157]" italics="true" pageId="7" pageNumber="378">Ocreatomurus</emphasis>
)
<emphasis id="B945A355FFAAFF99FD87FBFAFD5466B4" box="[635,732,1134,1157]" italics="true" pageId="7" pageNumber="378">oppositus</emphasis>
</taxonomicName>
<emphasis id="B945A355FFAAFF99FD1FFBFAFCBD66B7" bold="true" box="[739,821,1134,1158]" pageId="7" pageNumber="378">
<taxonomicNameLabel id="A2761E2EFFAAFF99FD1FFBFAFCBD66B7" box="[739,821,1134,1158]" pageId="7" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
:
<emphasis id="B945A355FFAAFF99FCBFFBFAFCD466B7" bold="true" box="[835,860,1134,1158]" pageId="7" pageNumber="378">23</emphasis>
, left side of manubrium (dorsal view);
<emphasis id="B945A355FFAAFF99FB0DFBFAFA8266B7" bold="true" box="[1265,1290,1134,1158]" pageId="7" pageNumber="378">24</emphasis>
, distal area of manubrium (dorsal view, arrow pointing to distal corner chaeta);
<emphasis id="B945A355FFAAFF99FCD1FB06FCCE669B" bold="true" box="[813,838,1170,1194]" pageId="7" pageNumber="378">25</emphasis>
, proximal part of dens with basal spines arranged in single row (dorsal view);
<emphasis id="B945A355FFAAFF99FEA5FB22FEFA66FF" bold="true" box="[345,370,1206,1230]" pageId="7" pageNumber="378">26</emphasis>
, basal dental spines arranged in double rows (dorsal view);
<emphasis id="B945A355FFAAFF99FC28FB22FC6566FF" bold="true" box="[980,1005,1206,1230]" pageId="7" pageNumber="378">27</emphasis>
, examples of dental spines;
<emphasis id="B945A355FFAAFF99FAF1FB22FAAE66FF" bold="true" box="[1293,1318,1206,1230]" pageId="7" pageNumber="378">28</emphasis>
, left mucro (inner view).
</paragraph>
</caption>
<subSubSection id="C32B2CCCFFAAFF99FF3BFAD0FE5D67DF" pageId="7" pageNumber="378" type="etymology">
<paragraph id="8B8E7F47FFAAFF99FF3BFAD0FDEC6797" blockId="7.[151,1437,1348,2023]" pageId="7" pageNumber="378">
<emphasis id="B945A355FFAAFF99FF3BFAD0FEC6676F" bold="true" box="[199,334,1348,1374]" pageId="7" pageNumber="378">Etymology.</emphasis>
The Latin word
<taxonomicName id="4C3104C4FFAAFF99FDEEFAD1FDF5676F" authorityName="Yu &amp; Xue &amp; Zhang" authorityYear="2025" box="[530,637,1349,1374]" class="Collembola" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Entomobryomorpha" pageId="7" pageNumber="378" phylum="Arthropoda" rank="species" species="oppositus">
<emphasis id="B945A355FFAAFF99FDEEFAD1FDF5676F" box="[530,637,1349,1374]" italics="true" pageId="7" pageNumber="378">oppositus</emphasis>
</taxonomicName>
(masculine adjective) means “opposite”, referencing the unique color pattern of the new species with dark ground and pale appendages, opposite to most other species of the subgenus with pale ground and darker appendages.
</paragraph>
<paragraph id="8B8E7F47FFAAFF99FF3BFA24FE5D67DF" blockId="7.[151,1437,1348,2023]" pageId="7" pageNumber="378">
<emphasis id="B945A355FFAAFF99FF3BFA24FEA267FB" bold="true" box="[199,298,1456,1482]" pageId="7" pageNumber="378">Habitat.</emphasis>
In ground litter, dead logs and live mosses on steep slopes in a closed forest; also under a wood board beside a cottage in a village.
</paragraph>
</subSubSection>
<subSubSection id="C32B2CCCFFAAFF96FF3BFA6CFD0862E4" lastPageId="8" lastPageNumber="379" pageId="7" pageNumber="378" type="discussion">
<paragraph id="8B8E7F47FFAAFF99FF3BFA6CFD5965AF" blockId="7.[151,1437,1348,2023]" pageId="7" pageNumber="378">
<emphasis id="B945A355FFAAFF99FF3BFA6CFEB36423" bold="true" box="[199,315,1528,1554]" pageId="7" pageNumber="378">Remarks.</emphasis>
At first glance, this new species resembles many others of the subgenus
<taxonomicName id="4C3104C4FFAAFF99FB8AFA6DFA146423" authorityName="Yu" authorityYear="2023" box="[1142,1436,1528,1554]" class="Collembola" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Entomobryomorpha" pageId="7" pageNumber="378" phylum="Arthropoda" rank="subGenus" subGenus="Striatomurus">
<emphasis id="B945A355FFAAFF99FB8AFA6DFB786423" box="[1142,1264,1529,1554]" italics="true" pageId="7" pageNumber="378">Tomocerus</emphasis>
(
<emphasis id="B945A355FFAAFF99FAFDFA6CFA1B6423" box="[1281,1427,1528,1554]" italics="true" pageId="7" pageNumber="378">Striatomurus</emphasis>
)
</taxonomicName>
for its dark body color. However, other characters, particularly the compound dental spines, put it in
<taxonomicName id="4C3104C4FFAAFF99FB08F988FA1D6407" authorityName="Yu" authorityYear="2023" box="[1268,1429,1564,1590]" pageId="7" pageNumber="372" rank="subGenus" subGenus="Ocreatomurus">
<emphasis id="B945A355FFAAFF99FB08F988FA1D6407" box="[1268,1429,1564,1590]" italics="true" pageId="7" pageNumber="378">Ocreatomurus</emphasis>
</taxonomicName>
. Within the subgenus,
<taxonomicName id="4C3104C4FFAAFF99FE7BF9D5FDC9646B" authorityName="Yu &amp; Xue &amp; Zhang" authorityYear="2025" box="[391,577,1600,1626]" class="Collembola" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Entomobryomorpha" pageId="7" pageNumber="378" phylum="Arthropoda" rank="species" species="oppositus" status="sp. nov." subGenus="Ocreatomurus">
<emphasis id="B945A355FFAAFF99FE7BF9D5FE15646B" box="[391,413,1601,1626]" italics="true" pageId="7" pageNumber="378">T.</emphasis>
(
<emphasis id="B945A355FFAAFF99FE57F9D4FE4E646B" box="[427,454,1600,1626]" italics="true" pageId="7" pageNumber="378">O.</emphasis>
)
<emphasis id="B945A355FFAAFF99FE2AF9D5FDC9646B" box="[470,577,1601,1626]" italics="true" pageId="7" pageNumber="378">oppositus</emphasis>
</taxonomicName>
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<taxonomicNameLabel id="A2761E2EFFAAFF99FDBBF9D4FD28646B" box="[583,672,1600,1626]" pageId="7" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
possesses a unique color pattern with intensely dark ground and pale appendages, which opposes to most other species with pale ground and darker appendages. A few other species of
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<emphasis id="B945A355FFAAFF99FF6BF91CFEB06493" box="[151,312,1672,1698]" italics="true" pageId="7" pageNumber="378">Ocreatomurus</emphasis>
</taxonomicName>
, such as
<taxonomicName id="4C3104C4FFAAFF99FE5EF91DFB886493" authority="Yu, Zhang &amp; Deharveng, 2014" authorityName="Yu, Zhang &amp; Deharveng" authorityYear="2014" box="[418,1024,1672,1698]" class="Collembola" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Entomobryomorpha" pageId="7" pageNumber="378" phylum="Arthropoda" rank="species" species="postantennalis" subGenus="Ocreatomurus">
<emphasis id="B945A355FFAAFF99FE5EF91DFE3F6493" box="[418,439,1673,1698]" italics="true" pageId="7" pageNumber="378">T.</emphasis>
(
<emphasis id="B945A355FFAAFF99FE34F91CFE6B6493" box="[456,483,1672,1698]" italics="true" pageId="7" pageNumber="378">O.</emphasis>
)
<emphasis id="B945A355FFAAFF99FE09F91CFD106493" box="[501,664,1672,1698]" italics="true" pageId="7" pageNumber="378">postantennalis</emphasis>
<bibRefCitation id="EFA002B6FFAAFF99FD63F91CFB886493" author="Yu, D &amp; Zhang, F. &amp; Deharveng, L." box="[671,1024,1672,1698]" pageId="7" pageNumber="378" pagination="61 - 70" refId="ref5651" refString="Yu, D, Zhang, F. &amp; Deharveng, L. (2014) A peculiar cave species of Tomocerus (Collembola, Tomoceridae, Tomocerinae) from Vietnam, with a discussion of the postantennal organ and prelabral chaetae in Tomocerinae. Zookeys, 408, 61 - 70. https: // doi. org / 10.3897 / zookeys. 408.7030" type="journal article" year="2014">Yu, Zhang &amp; Deharveng, 2014</bibRefCitation>
</taxonomicName>
,
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<emphasis id="B945A355FFAAFF99FBF0F91DFBA96493" box="[1036,1057,1673,1698]" italics="true" pageId="7" pageNumber="378">T.</emphasis>
(
<emphasis id="B945A355FFAAFF99FBCEF91CFBC56493" box="[1074,1101,1672,1698]" italics="true" pageId="7" pageNumber="378">O.</emphasis>
)
<emphasis id="B945A355FFAAFF99FBA2F91CFB1E6493" box="[1118,1174,1672,1698]" italics="true" pageId="7" pageNumber="378">dong</emphasis>
<bibRefCitation id="EFA002B6FFAAFF99FB61F91CFACE6493" author="Yu, D. &amp; Li, Y." box="[1181,1350,1672,1698]" pageId="7" pageNumber="378" pagination="361 - 372" refId="ref5521" refString="Yu, D. &amp; Li, Y. (2016) New troglomorphic species of Tomocerus with well-developed postantennal organs (Collembola: Tomoceridae). Zootaxa, 4162 (2), 361 - 372. http: // doi. org / 10.11646 / zootaxa. 4162.2.10" type="journal article" year="2016">Yu &amp; Li, 2016</bibRefCitation>
</taxonomicName>
,
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<emphasis id="B945A355FFAAFF99FAAEF91DFAEF6493" box="[1362,1383,1673,1698]" italics="true" pageId="7" pageNumber="378">T.</emphasis>
(
<emphasis id="B945A355FFAAFF99FA84F91CFA1B6493" box="[1400,1427,1672,1698]" italics="true" pageId="7" pageNumber="378">O.</emphasis>
)
<emphasis id="B945A355FFAAFF99FF6BF938FE9D64F7" box="[151,277,1708,1734]" italics="true" pageId="7" pageNumber="378">deharvengi</emphasis>
<bibRefCitation id="EFA002B6FFAAFF99FEE0F938FE4A64F7" author="Yu, D. &amp; Li, Y." box="[284,450,1708,1734]" pageId="7" pageNumber="378" pagination="361 - 372" refId="ref5521" refString="Yu, D. &amp; Li, Y. (2016) New troglomorphic species of Tomocerus with well-developed postantennal organs (Collembola: Tomoceridae). Zootaxa, 4162 (2), 361 - 372. http: // doi. org / 10.11646 / zootaxa. 4162.2.10" type="journal article" year="2016">Yu &amp; Li, 2016</bibRefCitation>
</taxonomicName>
and
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<emphasis id="B945A355FFAAFF99FE07F939FD9964F7" box="[507,529,1709,1734]" italics="true" pageId="7" pageNumber="378">T.</emphasis>
(
<emphasis id="B945A355FFAAFF99FDDEF938FDB564F7" box="[546,573,1708,1734]" italics="true" pageId="7" pageNumber="378">O.</emphasis>
)
<emphasis id="B945A355FFAAFF99FDB2F938FD2A64F7" box="[590,674,1708,1734]" italics="true" pageId="7" pageNumber="378">cthulhu</emphasis>
<bibRefCitation id="EFA002B6FFAAFF99FD56F938FCDB64F7" author="Yu, D. &amp; Li, Y." box="[682,851,1708,1734]" pageId="7" pageNumber="378" pagination="361 - 372" refId="ref5521" refString="Yu, D. &amp; Li, Y. (2016) New troglomorphic species of Tomocerus with well-developed postantennal organs (Collembola: Tomoceridae). Zootaxa, 4162 (2), 361 - 372. http: // doi. org / 10.11646 / zootaxa. 4162.2.10" type="journal article" year="2016">Yu &amp; Li, 2016</bibRefCitation>
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, also have dark pigment covering almost the whole body, however, pigment in these species is diffusely distributed, and the whole animals are from light to dark grey, not as intensely dark as
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<emphasis id="B945A355FFAAFF99FE5FF961FE30653F" box="[419,440,1781,1806]" italics="true" pageId="7" pageNumber="378">T.</emphasis>
(
<emphasis id="B945A355FFAAFF99FE34F960FE6B653F" box="[456,483,1780,1806]" italics="true" pageId="7" pageNumber="378">O.</emphasis>
)
<emphasis id="B945A355FFAAFF99FE0FF961FDD6653F" box="[499,606,1781,1806]" italics="true" pageId="7" pageNumber="378">oppositus</emphasis>
</taxonomicName>
<emphasis id="B945A355FFAAFF99FD99F960FD37653F" bold="true" box="[613,703,1780,1806]" pageId="7" pageNumber="378">
<taxonomicNameLabel id="A2761E2EFFAAFF99FD99F960FD37653F" box="[613,703,1780,1806]" pageId="7" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
In spite of the color pattern, this new species is most similar to
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<emphasis id="B945A355FFAAFF99FA74F961FA15653F" box="[1416,1437,1781,1806]" italics="true" pageId="7" pageNumber="378">T.</emphasis>
(
<emphasis id="B945A355FFAAFF99FF5DF88CFF346503" box="[161,188,1816,1842]" italics="true" pageId="7" pageNumber="378">O.</emphasis>
)
<emphasis id="B945A355FFAAFF99FF30F88CFEA16503" box="[204,297,1816,1842]" italics="true" pageId="7" pageNumber="378">spinulus</emphasis>
</taxonomicName>
in the absence of blunt chaetae on manubrium, presence of single large distal dental spine, and small size of denticles on dental spines, but is different from the later mainly in the larger body size, relatively longer antennae (10 %–38 % longer, average 20 %), larger number of blunt inner chaetae on tibiotarsi, and number and position of manubrial prominent chaetae (
<tableCitation id="C6B34AFCFFAAFF99FD90F810FD3765AF" box="[620,703,1924,1950]" captionStart="TABLE 1" captionStartId="5.[152,244,151,175]" captionText="TABLE 1. Main characters distinguishing Tomocerus (Ocreatomurus) spinulus Chen &amp; Christiansen, 1998 (sensu Yu et al. 2018), T. (O.) gyirongensis sp. nov. and T. (O.) oppositus sp. nov." pageId="7" pageNumber="378">Table 1</tableCitation>
).
</paragraph>
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A few variations in macrochaetotaxy exist in
<taxonomicName id="4C3104C4FFAAFF99FD38F83DFC0865F3" authorityName="Yu &amp; Xue &amp; Zhang" authorityYear="2025" box="[708,896,1960,1986]" class="Collembola" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Entomobryomorpha" pageId="7" pageNumber="378" phylum="Arthropoda" rank="species" species="oppositus" status="sp. nov." subGenus="Ocreatomurus">
<emphasis id="B945A355FFAAFF99FD38F83DFD5165F3" box="[708,729,1961,1986]" italics="true" pageId="7" pageNumber="378">T.</emphasis>
(
<emphasis id="B945A355FFAAFF99FD15F83CFC8C65F3" box="[745,772,1960,1986]" italics="true" pageId="7" pageNumber="378">O.</emphasis>
)
<emphasis id="B945A355FFAAFF99FCE9F83DFC0865F3" box="[789,896,1961,1986]" italics="true" pageId="7" pageNumber="378">oppositus</emphasis>
</taxonomicName>
<emphasis id="B945A355FFAAFF99FC74F83CFC6965F3" bold="true" box="[904,993,1960,1986]" pageId="7" pageNumber="378">
<taxonomicNameLabel id="A2761E2EFFAAFF99FC74F83CFC6965F3" box="[904,993,1960,1986]" pageId="7" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
On the head, macrochaeta A3 is absent from the right side in
<typeStatus id="548AC1E5FFAAFF99FE79F858FE6D65D7" box="[389,485,1996,2022]" pageId="7" pageNumber="378" type="paratype">paratype</typeStatus>
#5, and macrochaeta S0 is absent in
<typeStatus id="548AC1E5FFAAFF99FC80F858FC5465D7" box="[892,988,1996,2022]" pageId="7" pageNumber="378" type="paratype">paratype</typeStatus>
#4. On Th. II, macrochaeta m2 is absent from one side in
<typeStatus id="548AC1E5FFA5FF96FEABFF03FE4A6280" box="[343,450,151,177]" pageId="8" pageNumber="379" type="paratype">paratypes</typeStatus>
#2 and #4, whereas chaeta ap5, which is a microchaeta in most types, becomes a small macrochaeta on the left side in
<typeStatus id="548AC1E5FFA5FF96FE09FF2FFDDD62E4" box="[501,597,187,213]" pageId="8" pageNumber="379" type="paratype">paratype</typeStatus>
#6.
</paragraph>
</subSubSection>
</treatment>
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514
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<mods:title id="9E383922A679AF99E1D0BB814C0F0A8F">Two new species of Tomocerus (Ocreatomurus) Yu from Xizang, China (Collembola, Tomoceridae)</mods:title>
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<emphasis id="B945A355FFAFFF9CFF6BFF03FE946280" bold="true" box="[151,284,151,177]" italics="true" pageId="2" pageNumber="373">Tomocerus</emphasis>
(
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)
<emphasis id="B945A355FFAFFF9CFE10FF03FD0D6280" bold="true" box="[492,645,151,177]" italics="true" pageId="2" pageNumber="373">gyirongensis</emphasis>
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<subSubSection id="C32B2CCCFFAFFF9CFF6BFF2FFEEF62E4" box="[151,359,187,213]" pageId="2" pageNumber="373" type="description">
<paragraph id="8B8E7F47FFAFFF9CFF6BFF2FFEEF62E4" blockId="2.[151,746,151,213]" box="[151,359,187,213]" pageId="2" pageNumber="373">
<figureCitation id="130A63C2FFAFFF9CFF6BFF2FFE8062E4" box="[151,264,187,213]" captionStart-0="FIGURES 1–3" captionStart-1="FIGURES 4–9" captionStart-2="FIGURES 10–14" captionStartId-0="2.[151,264,1145,1169]" captionStartId-1="3.[151,264,1573,1597]" captionStartId-2="4.[151,264,1148,1172]" captionTargetBox-0="[151,1436,777,1121]" captionTargetBox-1="[159,1434,186,1545]" captionTargetBox-2="[159,1427,190,1124]" captionTargetId-0="figure-242@2.[151,1436,777,1121]" captionTargetId-1="figure-17@3.[151,1435,181,1549]" captionTargetId-2="figure-30@4.[151,1435,181,1125]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURES 1–3. Photographs of Tomocerus (Ocreatomurus) gyirongensis sp. nov. in ethanol (lateral view): 1, juvenile (arrows indicating positions of diffuse pigment around bothriotricha); 2, subadult; 3, fully grown adult." captionText-1="FIGURES 4–9. Tomocerus (Ocreatomurus) gyirongensis sp. nov.: 4, cephalic dorsal macrochaetotaxy (open circle—chaetal socket); 5, dorsal macrochaetotaxy of Th. II—Abd. V (wavy line—bothriotricha, open circle with a slash—pseudopore); 6, trochantero-femoral organ (inner view); 7, tibiotarsi I, II and III (lateral view, showing tenent hairs and stronger inner chaetae only, other chaetae not shown, arrows pointing to blunt chaetae); 8, hind claw complex (lateral view, showing chaetae on only one side except for accessory chaetae, t—tenent hair, aa—anterior accessory chaeta, ap—posterior accessory chaeta, g—guard chaeta, p—pretarsal chaeta); 9, tenaculum (anterior view)." captionText-2="FIGURES 10–14. Tomocerus (Ocreatomurus) gyirongensis sp. nov.: 10, left side of manubrium (dorsal view, showing lateral chaetae and prominent chaetae, Λ—scale socket); 11, distal area of manubrium (dorsal view, arrow pointing to distal corner chaeta, dashed arrow showing optional smaller chaeta); 12, proximal part of dens (dorsal view, showing dental spines and prominent chaeta); 13, examples of dental spines; 14, right mucro (outer view)." figureDoi-0="http://doi.org/10.5281/zenodo.14735180" figureDoi-1="http://doi.org/10.5281/zenodo.14735182" figureDoi-2="http://doi.org/10.5281/zenodo.14735184" httpUri-0="https://zenodo.org/record/14735180/files/figure.png" httpUri-1="https://zenodo.org/record/14735182/files/figure.png" httpUri-2="https://zenodo.org/record/14735184/files/figure.png" pageId="2" pageNumber="373">Figs 1–14</figureCitation>
,
<tableCitation id="C6B34AFCFFAFFF9CFEE8FF2FFEEF62E4" box="[276,359,187,213]" captionStart="TABLE 1" captionStartId="5.[152,244,151,175]" captionText="TABLE 1. Main characters distinguishing Tomocerus (Ocreatomurus) spinulus Chen &amp; Christiansen, 1998 (sensu Yu et al. 2018), T. (O.) gyirongensis sp. nov. and T. (O.) oppositus sp. nov." pageId="2" pageNumber="373">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32B2CCCFFAFFF9CFF6BFE97FE3063C4" pageId="2" pageNumber="373" type="materials_examined">
<paragraph id="8B8E7F47FFAFFF9CFF6BFE97FE3063C4" blockId="2.[151,1436,259,753]" pageId="2" pageNumber="373">
<emphasis id="B945A355FFAFFF9CFF6BFE97FEC2632C" bold="true" box="[151,330,259,285]" pageId="2" pageNumber="373">Type material.</emphasis>
<materialsCitation id="3B59751AFFAFFF9CFEAAFE97FC536354" collectingDate="2018-08-06" collectionCode="WGS" collectorName="Daoyuan Yu &amp; Yiming Wei &amp; Chunyan Qin" country="China" county="Gyirong County" elevation="2965" latitude="28.4392" location="Gyirong Valley" longLatPrecision="1" longitude="85.25235" municipality="Autonomous Region" pageId="2" pageNumber="373" specimenCode="WGS84" specimenCount="1" stateProvince="Xizang" typeStatus="holotype">
<typeStatus id="548AC1E5FFAFFF9CFEAAFE97FE4B632C" box="[342,451,259,285]" pageId="2" pageNumber="373" type="holotype">Holotype</typeStatus>
: male on slide, coniferous and broad-leaved mixed forest,
<location id="8EEE299CFFAFFF9CFB81FE97FAA7632C" LSID="urn:lsid:plazi:treatment:0398CE51FFAFFF9AFF6BFF03FCB5652C:8EEE299CFFAFFF9CFB81FE97FAA7632C" box="[1149,1327,259,285]" country="China" county="Gyirong County" latitude="28.4392" longLatPrecision="1" longitude="85.25235" municipality="Autonomous Region" name="Gyirong Valley" pageId="2" pageNumber="373" stateProvince="Xizang">Gyirong Valley</location>
,
<collectingCounty id="62EF07CBFFAFFF9CFAC3FE97FF636370" pageId="2" pageNumber="373">Gyirong County</collectingCounty>
,
<location id="8EEE299CFFAFFF9CFF09FEB3FECF6370" LSID="urn:lsid:plazi:treatment:0398CE51FFAFFF9AFF6BFF03FCB5652C:8EEE299CFFAFFF9CFF09FEB3FECF6370" box="[245,327,295,321]" country="China" county="Gyirong County" latitude="28.4392" longLatPrecision="1" longitude="85.25235" municipality="Autonomous Region" name="Xigaze" pageId="2" pageNumber="373" stateProvince="Xizang">Xigaze</location>
,
<collectingRegion id="49F5B1A5FFAFFF9CFEADFEB3FE2A6370" box="[337,418,295,321]" country="China" name="Xizang" pageId="2" pageNumber="373">Xizang</collectingRegion>
<collectingMunicipality id="6BEAE53DFFAFFF9CFE5AFEB3FD196370" box="[422,657,295,321]" pageId="2" pageNumber="373">Autonomous Region</collectingMunicipality>
,
<collectingCountry id="F3263FD7FFAFFF9CFD67FEB3FD696370" box="[667,737,295,321]" name="China" pageId="2" pageNumber="373">China</collectingCountry>
,
<geoCoordinate id="EE051980FFAFFF9CFD17FEBDFCF06375" box="[747,888,297,324]" degrees="85" direction="east" minutes="15.141" orientation="longitude" pageId="2" pageNumber="373" precision="1" value="85.25235">85°15.141′E</geoCoordinate>
,
<geoCoordinate id="EE051980FFAFFF9CFC7FFEBDFB9A6375" box="[899,1042,297,324]" degrees="28" direction="north" minutes="26.352" orientation="latitude" pageId="2" pageNumber="373" precision="1" value="28.4392">28°26.352′N</geoCoordinate>
(
<specimenCode id="DB97D73CFFAFFF9CFBE3FEB3FBF76370" box="[1055,1151,295,321]" collectionCode="WGS" pageId="2" pageNumber="373">WGS84</specimenCode>
), alt.
<quantity id="4CC9D2A2FFAFFF9CFB46FEB3FA996373" box="[1210,1297,295,322]" metricMagnitude="3" metricUnit="m" metricValue="2.965" pageId="2" pageNumber="373" unit="m" value="2965.0">
<elevation id="001C9874FFAFFF9CFB46FEB3FA996373" box="[1210,1297,295,322]" metricMagnitude="3" metricUnit="m" metricValue="2.965" pageId="2" pageNumber="373" unit="m" value="2965.0">2965 m</elevation>
</quantity>
,
<date id="FF8F5987FFAFFF9CFAE0FEB3FA106370" box="[1308,1432,295,321]" pageId="2" pageNumber="373" value="2018-08-06">
<collectingDate id="EFCBA06FFFAFFF9CFAE0FEB3FA106370" box="[1308,1432,295,321]" pageId="2" pageNumber="373" value="2018-08-06">6.viii.2018</collectingDate>
</date>
, leg.
<collectorName id="26C41A91FFAFFF9CFF36FED8FED16357" box="[202,345,332,358]" pageId="2" pageNumber="373">Daoyuan Yu</collectorName>
,
<collectorName id="26C41A91FFAFFF9CFE9AFEDFFE676354" box="[358,495,331,357]" pageId="2" pageNumber="373">Yiming Wei</collectorName>
and
<collectorName id="26C41A91FFAFFF9CFDD7FEDFFD4B6354" box="[555,707,331,357]" pageId="2" pageNumber="373">Chunyan Qin</collectorName>
(Sample code 18XZ13)
</materialsCitation>
.
<typeStatus id="548AC1E5FFAFFF9CFC15FED8FBD26357" box="[1001,1114,332,358]" pageId="2" pageNumber="373" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D37B4CEFFAFFF9CFB9BFED8FB606354" box="[1127,1256,331,358]" count="1" pageId="2" pageNumber="373" type="female">one female</specimenCount>
,
<specimenCount id="9D37B4CEFFAFFF9CFB0AFED8FAE26354" box="[1270,1386,331,358]" count="2" pageId="2" pageNumber="373" type="male">two males</specimenCount>
and
<specimenCount id="9D37B4CEFFAFFF9CFF6BFEE4FEAC63B8" box="[151,292,367,394]" count="1" pageId="2" pageNumber="373" type="juvenile">one juvenile</specimenCount>
on slides,
<materialsCitation id="3B59751AFFAFFF9CFE59FEFBFBE063B8" box="[421,1128,367,394]" collectingDate="2018-08-06" collectionCode="WGS" collectorName="Daoyuan Yu &amp; Yiming Wei &amp; Chunyan Qin" country="China" county="Gyirong County" elevation="2965" latitude="28.4392" location="Gyirong Valley" longLatPrecision="1" longitude="85.25235" municipality="Autonomous Region" pageId="2" pageNumber="373" specimenCount="19" specimenCount-subadult="19" stateProvince="Xizang" typeStatus="paratype">
<specimenCount id="9D37B4CEFFAFFF9CFE59FEFBFDBD63B8" box="[421,565,367,393]" count="19" pageId="2" pageNumber="373" type="subadult">19 subadults</specimenCount>
and juveniles in ethanol, same data as holotype
</materialsCitation>
;
<materialsCitation id="3B59751AFFAFFF9CFB8AFEFBFE3B63C4" collectingDate="2018-08-07" collectionCode="NJAU" collectorName="Daoyuan Yu &amp; Yiming Wei &amp; Chunyan Qin &amp; All" country="China" county="Gyirong County" elevation="2765" latitude="28.375383" location="Jipu Village" longLatPrecision="1" longitude="85.32822" municipality="Autonomous Region" pageId="2" pageNumber="373" specimenCount="4" specimenCount-subadult="4" stateProvince="Xizang" typeStatus="paratype">
<specimenCount id="9D37B4CEFFAFFF9CFB8AFEFBFA9163B8" box="[1142,1305,367,393]" count="4" pageId="2" pageNumber="373" type="subadult">four subadults</specimenCount>
in ethanol, under a wood board in
<location id="8EEE299CFFAFFF9CFE59FE07FDB8639C" LSID="urn:lsid:plazi:treatment:0398CE51FFAFFF9AFF6BFF03FCB5652C:8EEE299CFFAFFF9CFE59FE07FDB8639C" box="[421,560,403,429]" country="China" county="Gyirong County" latitude="28.375383" longLatPrecision="1" longitude="85.32822" municipality="Autonomous Region" name="Jipu Village" pageId="2" pageNumber="373" stateProvince="Xizang">Jipu Village</location>
,
<collectingCounty id="62EF07CBFFAFFF9CFDC1FE07FD7F639C" box="[573,759,403,429]" pageId="2" pageNumber="373">Gyirong County</collectingCounty>
,
<location id="8EEE299CFFAFFF9CFCF8FE07FCDE639C" LSID="urn:lsid:plazi:treatment:0398CE51FFAFFF9AFF6BFF03FCB5652C:8EEE299CFFAFFF9CFCF8FE07FCDE639C" box="[772,854,403,429]" country="China" county="Gyirong County" latitude="28.375383" longLatPrecision="1" longitude="85.32822" municipality="Autonomous Region" name="Xigaze" pageId="2" pageNumber="373" stateProvince="Xizang">Xigaze</location>
,
<collectingRegion id="49F5B1A5FFAFFF9CFC98FE07FC3D639C" box="[868,949,403,429]" country="China" name="Xizang" pageId="2" pageNumber="373">Xizang</collectingRegion>
<collectingMunicipality id="6BEAE53DFFAFFF9CFC40FE07FB22639C" box="[956,1194,403,429]" pageId="2" pageNumber="373">Autonomous Region</collectingMunicipality>
,
<collectingCountry id="F3263FD7FFAFFF9CFB4BFE07FB75639C" box="[1207,1277,403,429]" name="China" pageId="2" pageNumber="373">China</collectingCountry>
,
<geoCoordinate id="EE051980FFAFFF9CFAF6FE01FA106381" box="[1290,1432,405,432]" degrees="85" direction="east" minutes="19.693" orientation="longitude" pageId="2" pageNumber="373" precision="1" value="85.32822">85°19.693′E</geoCoordinate>
,
<geoCoordinate id="EE051980FFAFFF9CFF6BFE2DFEA063E5" box="[151,296,441,468]" degrees="28" direction="north" minutes="22.523" orientation="latitude" pageId="2" pageNumber="373" precision="1" value="28.375383">28°22.523′N</geoCoordinate>
, alt.
<quantity id="4CC9D2A2FFAFFF9CFEA1FE23FE3C63E3" box="[349,436,439,466]" metricMagnitude="3" metricUnit="m" metricValue="2.7649999999999997" pageId="2" pageNumber="373" unit="m" value="2765.0">
<elevation id="001C9874FFAFFF9CFEA1FE23FE3C63E3" box="[349,436,439,466]" metricMagnitude="3" metricUnit="m" metricValue="2.7649999999999997" pageId="2" pageNumber="373" unit="m" value="2765.0">2765 m</elevation>
</quantity>
,
<date id="FF8F5987FFAFFF9CFE3CFE23FDB463E0" box="[448,572,439,465]" pageId="2" pageNumber="373" value="2018-08-07">
<collectingDate id="EFCBA06FFFAFFF9CFE3CFE23FDB463E0" box="[448,572,439,465]" pageId="2" pageNumber="373" value="2018-08-07">7.viii.2018</collectingDate>
</date>
, leg.
<collectorName id="26C41A91FFAFFF9CFD8BFE2CFC8B63E3" box="[631,771,440,466]" pageId="2" pageNumber="373">Daoyuan Yu</collectorName>
,
<collectorName id="26C41A91FFAFFF9CFCF1FE23FC1B63E0" box="[781,915,439,465]" pageId="2" pageNumber="373">Yiming Wei</collectorName>
and
<collectorName id="26C41A91FFAFFF9CFC35FE23FBD763E0" box="[969,1119,439,465]" pageId="2" pageNumber="373">Chunyan Qin</collectorName>
(Sample code 18XZ15).
<collectorName id="26C41A91FFAFFF9CFA84FE23FA1463E0" box="[1400,1436,439,465]" pageId="2" pageNumber="373">All</collectorName>
types deposited in
<collectionCode id="ED20E782FFAFFF9CFE94FE4FFE3B63C4" box="[360,435,475,501]" pageId="2" pageNumber="373">NJAU</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32B2CCCFFAFFF9CFF3BFE6BFC6065FF" pageId="2" pageNumber="373" type="description">
<paragraph id="8B8E7F47FFAFFF9CFF3BFE6BFEF560C0" blockId="2.[151,1436,259,753]" pageId="2" pageNumber="373">
<emphasis id="B945A355FFAFFF9CFF3BFE6BFED16028" bold="true" box="[199,345,511,537]" pageId="2" pageNumber="373">Description.</emphasis>
Body length 3.0–
<quantity id="4CC9D2A2FFAFFF9CFDD8FE6BFDF2602B" box="[548,634,511,538]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="2" pageNumber="373" unit="mm" value="4.3">4.3 mm</quantity>
(average 3.8,
<specimenCount id="9D37B4CEFFAFFF9CFCE0FE6BFC416028" box="[796,969,511,537]" count="5" pageId="2" pageNumber="373" type="generic">five specimens</specimenCount>
). Body ground color pale yellow. Ant. III and IV purple. Diffuse reddish-purple pigment present mainly on Ant. I and II, head, coxae, tibiotarsi, ventral tube, and manubrium. In juveniles (
<figureCitation id="130A63C2FFAFFF9CFE12FDD3FDA76050" box="[494,559,583,609]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1145,1169]" captionTargetBox="[151,1436,777,1121]" captionTargetId="figure-242@2.[151,1436,777,1121]" captionTargetPageId="2" captionText="FIGURES 1–3. Photographs of Tomocerus (Ocreatomurus) gyirongensis sp. nov. in ethanol (lateral view): 1, juvenile (arrows indicating positions of diffuse pigment around bothriotricha); 2, subadult; 3, fully grown adult." figureDoi="http://doi.org/10.5281/zenodo.14735180" httpUri="https://zenodo.org/record/14735180/files/figure.png" pageId="2" pageNumber="373">Fig. 1</figureCitation>
), pigment on head distributed more evenly, and light reddish-purple pigment also around bothriotricha on terga, forming diffuse patches on Th. II and spots on other segments. In subadults (
<figureCitation id="130A63C2FFAFFF9CFF5CFD1BFF6B6098" box="[160,227,655,681]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1145,1169]" captionTargetBox="[151,1436,777,1121]" captionTargetId="figure-242@2.[151,1436,777,1121]" captionTargetPageId="2" captionText="FIGURES 1–3. Photographs of Tomocerus (Ocreatomurus) gyirongensis sp. nov. in ethanol (lateral view): 1, juvenile (arrows indicating positions of diffuse pigment around bothriotricha); 2, subadult; 3, fully grown adult." figureDoi="http://doi.org/10.5281/zenodo.14735180" httpUri="https://zenodo.org/record/14735180/files/figure.png" pageId="2" pageNumber="373">Fig. 2</figureCitation>
) and adults (
<figureCitation id="130A63C2FFAFFF9CFE87FD1BFE376098" box="[379,447,655,681]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1145,1169]" captionTargetBox="[151,1436,777,1121]" captionTargetId="figure-242@2.[151,1436,777,1121]" captionTargetPageId="2" captionText="FIGURES 1–3. Photographs of Tomocerus (Ocreatomurus) gyirongensis sp. nov. in ethanol (lateral view): 1, juvenile (arrows indicating positions of diffuse pigment around bothriotricha); 2, subadult; 3, fully grown adult." figureDoi="http://doi.org/10.5281/zenodo.14735180" httpUri="https://zenodo.org/record/14735180/files/figure.png" pageId="2" pageNumber="373">Fig. 3</figureCitation>
), pigment on head concentrating anteriorly around mouth and antennal bases, and the patterns around bothriotricha becoming indistinct. In fully grown adults (
<figureCitation id="130A63C2FFAFFF9CFC17FD27FBA760FC" box="[1003,1071,691,717]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1145,1169]" captionTargetBox="[151,1436,777,1121]" captionTargetId="figure-242@2.[151,1436,777,1121]" captionTargetPageId="2" captionText="FIGURES 1–3. Photographs of Tomocerus (Ocreatomurus) gyirongensis sp. nov. in ethanol (lateral view): 1, juvenile (arrows indicating positions of diffuse pigment around bothriotricha); 2, subadult; 3, fully grown adult." figureDoi="http://doi.org/10.5281/zenodo.14735180" httpUri="https://zenodo.org/record/14735180/files/figure.png" pageId="2" pageNumber="373">Fig. 3</figureCitation>
), purple pigment more intense, becoming brownish.
</paragraph>
<caption id="DF4E2FCFFFAFFF9CFF6BFBEDFBD06684" ID-DOI="http://doi.org/10.5281/zenodo.14735180" ID-Zenodo-Dep="14735180" httpUri="https://zenodo.org/record/14735180/files/figure.png" pageId="2" pageNumber="373" startId="2.[151,264,1145,1169]" targetBox="[151,1436,777,1121]" targetPageId="2" targetType="figure">
<paragraph id="8B8E7F47FFAFFF9CFF6BFBEDFBD06684" blockId="2.[151,1436,1145,1206]" pageId="2" pageNumber="373">
<emphasis id="B945A355FFAFFF9CFF6BFBEDFEB366A0" bold="true" box="[151,315,1145,1169]" pageId="2" pageNumber="373">FIGURES 1–3.</emphasis>
Photographs of
<taxonomicName id="4C3104C4FFAFFF9CFE1DFBEEFC0866A0" authorityName="Yu &amp; Xue &amp; Zhang" authorityYear="2025" box="[481,896,1145,1169]" class="Collembola" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Entomobryomorpha" pageId="2" pageNumber="373" phylum="Arthropoda" rank="species" species="gyirongensis" status="sp. nov." subGenus="Ocreatomurus">
<emphasis id="B945A355FFAFFF9CFE1DFBEEFDC766A0" box="[481,591,1146,1169]" italics="true" pageId="2" pageNumber="373">Tomocerus</emphasis>
(
<emphasis id="B945A355FFAFFF9CFDA3FBEEFD7866A0" box="[607,752,1146,1169]" italics="true" pageId="2" pageNumber="373">Ocreatomurus</emphasis>
)
<emphasis id="B945A355FFAFFF9CFD03FBEEFC0866A0" box="[767,896,1146,1169]" italics="true" pageId="2" pageNumber="373">gyirongensis</emphasis>
</taxonomicName>
<emphasis id="B945A355FFAFFF9CFC7AFBEDFC5E66A0" bold="true" box="[902,982,1145,1169]" pageId="2" pageNumber="373">
<taxonomicNameLabel id="A2761E2EFFAFFF9CFC7AFBEDFC5E66A0" box="[902,982,1145,1169]" pageId="2" pageNumber="373" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in ethanol (lateral view):
<emphasis id="B945A355FFAFFF9CFB21FBEDFB6266A0" bold="true" box="[1245,1258,1145,1169]" pageId="2" pageNumber="373">1</emphasis>
, juvenile (arrows indicating positions of diffuse pigment around bothriotricha);
<emphasis id="B945A355FFAFFF9CFCF7FB09FC906684" bold="true" box="[779,792,1181,1205]" pageId="2" pageNumber="373">2</emphasis>
, subadult;
<emphasis id="B945A355FFAFFF9CFC74FB09FC1D6684" bold="true" box="[904,917,1181,1205]" pageId="2" pageNumber="373">3</emphasis>
, fully grown adult.
</paragraph>
</caption>
<paragraph id="8B8E7F47FFAFFF9CFF3BFA9CFC1A6758" blockId="2.[151,1437,1287,1998]" pageId="2" pageNumber="373">
Intact four-segmented antennae 0.75–0.81 × length of body (average 0.78,
<specimenCount id="9D37B4CEFFAFFF9CFBF6FA9CFB3C6710" box="[1034,1204,1287,1314]" count="2" pageId="2" pageNumber="373" type="generic">two specimens</specimenCount>
). Length ratio of Ant I: II: III: IV = 1.0: 1.6: 8.5–10.1: 0.9–1.2. Ant. III basally with 3–8 scales. Cephalic dorsal macrochaetotaxy as in
<figureCitation id="130A63C2FFAFFF9CFF48FADBFF70675B" box="[180,248,1359,1386]" captionStart="FIGURES 4–9" captionStartId="3.[151,264,1573,1597]" captionTargetBox="[159,1434,186,1545]" captionTargetId="figure-17@3.[151,1435,181,1549]" captionTargetPageId="3" captionText="FIGURES 4–9. Tomocerus (Ocreatomurus) gyirongensis sp. nov.: 4, cephalic dorsal macrochaetotaxy (open circle—chaetal socket); 5, dorsal macrochaetotaxy of Th. II—Abd. V (wavy line—bothriotricha, open circle with a slash—pseudopore); 6, trochantero-femoral organ (inner view); 7, tibiotarsi I, II and III (lateral view, showing tenent hairs and stronger inner chaetae only, other chaetae not shown, arrows pointing to blunt chaetae); 8, hind claw complex (lateral view, showing chaetae on only one side except for accessory chaetae, t—tenent hair, aa—anterior accessory chaeta, ap—posterior accessory chaeta, g—guard chaeta, p—pretarsal chaeta); 9, tenaculum (anterior view)." figureDoi="http://doi.org/10.5281/zenodo.14735182" httpUri="https://zenodo.org/record/14735182/files/figure.png" pageId="2" pageNumber="373">Fig. 4</figureCitation>
. Posterior margin of head with 35–40 chaetae on each side.
</paragraph>
<paragraph id="8B8E7F47FFAFFF9CFF3BFAE0FD4D67BF" blockId="2.[151,1437,1287,1998]" box="[199,709,1395,1422]" pageId="2" pageNumber="373">
Pattern of body macrochaetotaxy as in
<figureCitation id="130A63C2FFAFFF9CFD80FAE7FD4867BF" box="[636,704,1395,1422]" captionStart="FIGURES 4–9" captionStartId="3.[151,264,1573,1597]" captionTargetBox="[159,1434,186,1545]" captionTargetId="figure-17@3.[151,1435,181,1549]" captionTargetPageId="3" captionText="FIGURES 4–9. Tomocerus (Ocreatomurus) gyirongensis sp. nov.: 4, cephalic dorsal macrochaetotaxy (open circle—chaetal socket); 5, dorsal macrochaetotaxy of Th. II—Abd. V (wavy line—bothriotricha, open circle with a slash—pseudopore); 6, trochantero-femoral organ (inner view); 7, tibiotarsi I, II and III (lateral view, showing tenent hairs and stronger inner chaetae only, other chaetae not shown, arrows pointing to blunt chaetae); 8, hind claw complex (lateral view, showing chaetae on only one side except for accessory chaetae, t—tenent hair, aa—anterior accessory chaeta, ap—posterior accessory chaeta, g—guard chaeta, p—pretarsal chaeta); 9, tenaculum (anterior view)." figureDoi="http://doi.org/10.5281/zenodo.14735182" httpUri="https://zenodo.org/record/14735182/files/figure.png" pageId="2" pageNumber="373">Fig. 5</figureCitation>
.
</paragraph>
<paragraph id="8B8E7F47FFAFFF9CFF3BFA03FAC76457" blockId="2.[151,1437,1287,1998]" pageId="2" pageNumber="373">
Single chaeta of femoral organ longer than that of trochanteral organ (
<figureCitation id="130A63C2FFAFFF9CFC24FA03FB946780" box="[984,1052,1431,1457]" captionStart="FIGURES 4–9" captionStartId="3.[151,264,1573,1597]" captionTargetBox="[159,1434,186,1545]" captionTargetId="figure-17@3.[151,1435,181,1549]" captionTargetPageId="3" captionText="FIGURES 4–9. Tomocerus (Ocreatomurus) gyirongensis sp. nov.: 4, cephalic dorsal macrochaetotaxy (open circle—chaetal socket); 5, dorsal macrochaetotaxy of Th. II—Abd. V (wavy line—bothriotricha, open circle with a slash—pseudopore); 6, trochantero-femoral organ (inner view); 7, tibiotarsi I, II and III (lateral view, showing tenent hairs and stronger inner chaetae only, other chaetae not shown, arrows pointing to blunt chaetae); 8, hind claw complex (lateral view, showing chaetae on only one side except for accessory chaetae, t—tenent hair, aa—anterior accessory chaeta, ap—posterior accessory chaeta, g—guard chaeta, p—pretarsal chaeta); 9, tenaculum (anterior view)." figureDoi="http://doi.org/10.5281/zenodo.14735182" httpUri="https://zenodo.org/record/14735182/files/figure.png" pageId="2" pageNumber="373">Fig. 6</figureCitation>
). Tibiotarsi I, II, III ventrally with 8–10, 8–9, 7–9 strong chaetae, 5–8, 7–8, 7–9 of them blunt and spine-like, respectively (
<figureCitation id="130A63C2FFAFFF9CFBA6FA2FFB1667E7" box="[1114,1182,1467,1494]" captionStart="FIGURES 4–9" captionStartId="3.[151,264,1573,1597]" captionTargetBox="[159,1434,186,1545]" captionTargetId="figure-17@3.[151,1435,181,1549]" captionTargetPageId="3" captionText="FIGURES 4–9. Tomocerus (Ocreatomurus) gyirongensis sp. nov.: 4, cephalic dorsal macrochaetotaxy (open circle—chaetal socket); 5, dorsal macrochaetotaxy of Th. II—Abd. V (wavy line—bothriotricha, open circle with a slash—pseudopore); 6, trochantero-femoral organ (inner view); 7, tibiotarsi I, II and III (lateral view, showing tenent hairs and stronger inner chaetae only, other chaetae not shown, arrows pointing to blunt chaetae); 8, hind claw complex (lateral view, showing chaetae on only one side except for accessory chaetae, t—tenent hair, aa—anterior accessory chaeta, ap—posterior accessory chaeta, g—guard chaeta, p—pretarsal chaeta); 9, tenaculum (anterior view)." figureDoi="http://doi.org/10.5281/zenodo.14735182" httpUri="https://zenodo.org/record/14735182/files/figure.png" pageId="2" pageNumber="373">Fig. 7</figureCitation>
). Tenent hair 0.8–1.0 × the length of inner edge of unguis; accessory chaetae on tibiotarsi I and II almost equal to each other, and subequal to or shorter than pretarsal chaetae; on tibiotarsus III, anterior accessory chaeta longer than pretarsal chaetae, while posterior accessory chaeta subequal to pretarsal chaetae; guard chaetae 0.8–0.9 × the length of tenent hair. Unguiculus about 0.56–0.64 × the length of unguis (average 0.58,
<specimenCount id="9D37B4CEFFAFFF9CFD0DF9DFFC166454" box="[753,926,1611,1637]" count="4" pageId="2" pageNumber="373" type="generic">four specimens</specimenCount>
), its inner edge with 0–1 tooth (
<figureCitation id="130A63C2FFAFFF9CFB02F9DFFAC86457" box="[1278,1344,1611,1638]" captionStart="FIGURES 4–9" captionStartId="3.[151,264,1573,1597]" captionTargetBox="[159,1434,186,1545]" captionTargetId="figure-17@3.[151,1435,181,1549]" captionTargetPageId="3" captionText="FIGURES 4–9. Tomocerus (Ocreatomurus) gyirongensis sp. nov.: 4, cephalic dorsal macrochaetotaxy (open circle—chaetal socket); 5, dorsal macrochaetotaxy of Th. II—Abd. V (wavy line—bothriotricha, open circle with a slash—pseudopore); 6, trochantero-femoral organ (inner view); 7, tibiotarsi I, II and III (lateral view, showing tenent hairs and stronger inner chaetae only, other chaetae not shown, arrows pointing to blunt chaetae); 8, hind claw complex (lateral view, showing chaetae on only one side except for accessory chaetae, t—tenent hair, aa—anterior accessory chaeta, ap—posterior accessory chaeta, g—guard chaeta, p—pretarsal chaeta); 9, tenaculum (anterior view)." figureDoi="http://doi.org/10.5281/zenodo.14735182" httpUri="https://zenodo.org/record/14735182/files/figure.png" pageId="2" pageNumber="373">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="8B8E7F47FFAFFF9CFF3BF9FBFC6065FF" blockId="2.[151,1437,1287,1998]" pageId="2" pageNumber="373">
Ventral tube scaled on both faces; anterior face with 28–50 (average 38,
<specimenCount id="9D37B4CEFFAFFF9CFBE6F9FBFB5A64B8" box="[1050,1234,1647,1673]" count="3" pageId="2" pageNumber="373" type="generic">three specimens</specimenCount>
) chaetae on each side; posterior face with 82–122 (average 103,
<specimenCount id="9D37B4CEFFAFFF9CFD53F907FCEE649C" box="[687,870,1683,1709]" count="3" pageId="2" pageNumber="373" type="generic">three specimens</specimenCount>
) chaetae; each lateral flap with 137–170 (average 149,
<specimenCount id="9D37B4CEFFAFFF9CFF2CF923FE0D64E0" box="[208,389,1719,1745]" count="3" pageId="2" pageNumber="373" type="generic">three specimens</specimenCount>
) chaetae and 2–8 scales. Anterior face of tenaculum unscaled, with 14–16 (average 15,
<specimenCount id="9D37B4CEFFAFFF9CFA98F923FE8564C4" count="3" pageId="2" pageNumber="373" type="generic">three specimens</specimenCount>
) chaetae (
<figureCitation id="130A63C2FFAFFF9CFE79F94FFE4164C7" box="[389,457,1755,1782]" captionStart="FIGURES 4–9" captionStartId="3.[151,264,1573,1597]" captionTargetBox="[159,1434,186,1545]" captionTargetId="figure-17@3.[151,1435,181,1549]" captionTargetPageId="3" captionText="FIGURES 4–9. Tomocerus (Ocreatomurus) gyirongensis sp. nov.: 4, cephalic dorsal macrochaetotaxy (open circle—chaetal socket); 5, dorsal macrochaetotaxy of Th. II—Abd. V (wavy line—bothriotricha, open circle with a slash—pseudopore); 6, trochantero-femoral organ (inner view); 7, tibiotarsi I, II and III (lateral view, showing tenent hairs and stronger inner chaetae only, other chaetae not shown, arrows pointing to blunt chaetae); 8, hind claw complex (lateral view, showing chaetae on only one side except for accessory chaetae, t—tenent hair, aa—anterior accessory chaeta, ap—posterior accessory chaeta, g—guard chaeta, p—pretarsal chaeta); 9, tenaculum (anterior view)." figureDoi="http://doi.org/10.5281/zenodo.14735182" httpUri="https://zenodo.org/record/14735182/files/figure.png" pageId="2" pageNumber="373">Fig. 9</figureCitation>
). Ratio manubrium: dens: mucro = 3.3–3.9: 4.4–5.0: 1.0. Manubrium laterally with 9–10 strong chaetae; dorsally with 211–301 (average 252,
<specimenCount id="9D37B4CEFFAFFF9CFCDCF96BFC5C6528" box="[800,980,1791,1817]" count="3" pageId="2" pageNumber="373" type="generic">three specimens</specimenCount>
) chaetae of different sizes on each dorsal chaetal strip, 6–12 (average eight,
<specimenCount id="9D37B4CEFFAFFF9CFDDDF8B7FD46650C" box="[545,718,1827,1853]" count="4" pageId="2" pageNumber="373" type="generic">four specimens</specimenCount>
) pseudopores almost arranging in a file and scale strip running from base to about 3/4 the length of manubrium (
<figureCitation id="130A63C2FFAFFF9CFD3BF8D3FC926553" box="[711,794,1863,1890]" captionStart="FIGURES 10–14" captionStartId="4.[151,264,1148,1172]" captionTargetBox="[159,1427,190,1124]" captionTargetId="figure-30@4.[151,1435,181,1125]" captionTargetPageId="4" captionText="FIGURES 10–14. Tomocerus (Ocreatomurus) gyirongensis sp. nov.: 10, left side of manubrium (dorsal view, showing lateral chaetae and prominent chaetae, Λ—scale socket); 11, distal area of manubrium (dorsal view, arrow pointing to distal corner chaeta, dashed arrow showing optional smaller chaeta); 12, proximal part of dens (dorsal view, showing dental spines and prominent chaeta); 13, examples of dental spines; 14, right mucro (outer view)." figureDoi="http://doi.org/10.5281/zenodo.14735184" httpUri="https://zenodo.org/record/14735184/files/figure.png" pageId="2" pageNumber="373">Fig. 10</figureCitation>
). External distal corner chaeta as large as or shorter than medium-sized mesochaetae in chaetal strip (
<figureCitation id="130A63C2FFAFFF9CFD7CF8FFFD5A65B7" box="[640,722,1899,1926]" captionStart="FIGURES 10–14" captionStartId="4.[151,264,1148,1172]" captionTargetBox="[159,1427,190,1124]" captionTargetId="figure-30@4.[151,1435,181,1125]" captionTargetPageId="4" captionText="FIGURES 10–14. Tomocerus (Ocreatomurus) gyirongensis sp. nov.: 10, left side of manubrium (dorsal view, showing lateral chaetae and prominent chaetae, Λ—scale socket); 11, distal area of manubrium (dorsal view, arrow pointing to distal corner chaeta, dashed arrow showing optional smaller chaeta); 12, proximal part of dens (dorsal view, showing dental spines and prominent chaeta); 13, examples of dental spines; 14, right mucro (outer view)." figureDoi="http://doi.org/10.5281/zenodo.14735184" httpUri="https://zenodo.org/record/14735184/files/figure.png" pageId="2" pageNumber="373">Fig. 11</figureCitation>
). Dental spine formula
<emphasis id="B945A355FFAFFF9CFC29F8F8FB9B65B4" box="[981,1043,1900,1925]" italics="true" pageId="2" pageNumber="373">sensu</emphasis>
<bibRefCitation id="EFA002B6FFAFFF9CFBE5F8FFFB3565B7" author="Folsom, J. W." box="[1049,1213,1899,1926]" pageId="2" pageNumber="373" pagination="451 - 472" refId="ref5155" refString="Folsom, J. W. (1913) North American spring-tails of the sub-family Tomocerinae. Proceedings of the United States National Museum, 46, 451 - 472. https: // doi. org / 10.5479 / si. 00963801.46 - 2037.451" type="journal article" year="1913">Folsom (1913)</bibRefCitation>
as 4–8/3–5,
<emphasis id="B945A355FFAFFF9CFAB4F8FFFADE65B4" bold="true" box="[1352,1366,1899,1925]" pageId="2" pageNumber="373">1</emphasis>
, basal spines arranged in two rows (
<figureCitation id="130A63C2FFAFFF9CFE21F81BFDB9659B" box="[477,561,1935,1962]" captionStart="FIGURES 10–14" captionStartId="4.[151,264,1148,1172]" captionTargetBox="[159,1427,190,1124]" captionTargetId="figure-30@4.[151,1435,181,1125]" captionTargetPageId="4" captionText="FIGURES 10–14. Tomocerus (Ocreatomurus) gyirongensis sp. nov.: 10, left side of manubrium (dorsal view, showing lateral chaetae and prominent chaetae, Λ—scale socket); 11, distal area of manubrium (dorsal view, arrow pointing to distal corner chaeta, dashed arrow showing optional smaller chaeta); 12, proximal part of dens (dorsal view, showing dental spines and prominent chaeta); 13, examples of dental spines; 14, right mucro (outer view)." figureDoi="http://doi.org/10.5281/zenodo.14735184" httpUri="https://zenodo.org/record/14735184/files/figure.png" pageId="2" pageNumber="373">Fig. 12</figureCitation>
). Denticles almost evenly distributed on dental spines except for tips (
<figureCitation id="130A63C2FFAFFF9CFAC0F81BFA07659B" box="[1340,1423,1935,1962]" captionStart="FIGURES 10–14" captionStartId="4.[151,264,1148,1172]" captionTargetBox="[159,1427,190,1124]" captionTargetId="figure-30@4.[151,1435,181,1125]" captionTargetPageId="4" captionText="FIGURES 10–14. Tomocerus (Ocreatomurus) gyirongensis sp. nov.: 10, left side of manubrium (dorsal view, showing lateral chaetae and prominent chaetae, Λ—scale socket); 11, distal area of manubrium (dorsal view, arrow pointing to distal corner chaeta, dashed arrow showing optional smaller chaeta); 12, proximal part of dens (dorsal view, showing dental spines and prominent chaeta); 13, examples of dental spines; 14, right mucro (outer view)." figureDoi="http://doi.org/10.5281/zenodo.14735184" httpUri="https://zenodo.org/record/14735184/files/figure.png" pageId="2" pageNumber="373">Fig. 13</figureCitation>
). Mucro with 4–6 (average five,
<specimenCount id="9D37B4CEFFAFFF9CFE08F827FD1765FC" box="[500,671,1971,1997]" count="4" pageId="2" pageNumber="373" type="generic">four specimens</specimenCount>
) intermediate teeth (
<figureCitation id="130A63C2FFAFFF9CFC7BF827FC5265FF" box="[903,986,1971,1998]" captionStart="FIGURES 10–14" captionStartId="4.[151,264,1148,1172]" captionTargetBox="[159,1427,190,1124]" captionTargetId="figure-30@4.[151,1435,181,1125]" captionTargetPageId="4" captionText="FIGURES 10–14. Tomocerus (Ocreatomurus) gyirongensis sp. nov.: 10, left side of manubrium (dorsal view, showing lateral chaetae and prominent chaetae, Λ—scale socket); 11, distal area of manubrium (dorsal view, arrow pointing to distal corner chaeta, dashed arrow showing optional smaller chaeta); 12, proximal part of dens (dorsal view, showing dental spines and prominent chaeta); 13, examples of dental spines; 14, right mucro (outer view)." figureDoi="http://doi.org/10.5281/zenodo.14735184" httpUri="https://zenodo.org/record/14735184/files/figure.png" pageId="2" pageNumber="373">Fig. 14</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF4E2FCFFFAEFF9DFF6BF9B1FD6E64C0" ID-DOI="http://doi.org/10.5281/zenodo.14735182" ID-Zenodo-Dep="14735182" httpUri="https://zenodo.org/record/14735182/files/figure.png" pageId="3" pageNumber="374" startId="3.[151,264,1573,1597]" targetBox="[159,1434,186,1545]" targetPageId="3" targetType="figure">
<paragraph id="8B8E7F47FFAEFF9DFF6BF9B1FD6E64C0" blockId="3.[151,1436,1573,1777]" pageId="3" pageNumber="374">
<emphasis id="B945A355FFAEFF9DFF6BF9B1FEB4640C" bold="true" box="[151,316,1573,1597]" pageId="3" pageNumber="374">FIGURES 4–9.</emphasis>
<taxonomicName id="4C3104C4FFAEFF9DFEB8F9B2FD6E640C" authorityName="Yu &amp; Xue &amp; Zhang" authorityYear="2025" box="[324,742,1573,1597]" class="Collembola" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Entomobryomorpha" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="gyirongensis" status="sp. nov." subGenus="Ocreatomurus">
<emphasis id="B945A355FFAEFF9DFEB8F9B2FE3A640C" box="[324,434,1574,1597]" italics="true" pageId="3" pageNumber="374">Tomocerus</emphasis>
(
<emphasis id="B945A355FFAEFF9DFE3FF9B1FDDC640D" box="[451,596,1573,1596]" italics="true" pageId="3" pageNumber="374">Ocreatomurus</emphasis>
)
<emphasis id="B945A355FFAEFF9DFD99F9B2FD6E640C" box="[613,742,1574,1597]" italics="true" pageId="3" pageNumber="374">gyirongensis</emphasis>
</taxonomicName>
<emphasis id="B945A355FFAEFF9DFD11F9B1FCB7640C" bold="true" box="[749,831,1573,1597]" pageId="3" pageNumber="374">
<taxonomicNameLabel id="A2761E2EFFAEFF9DFD11F9B1FCB7640C" box="[749,831,1573,1597]" pageId="3" pageNumber="374" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
:
<emphasis id="B945A355FFAEFF9DFCB3F9B1FCD4640C" bold="true" box="[847,860,1573,1597]" pageId="3" pageNumber="374">4</emphasis>
, cephalic dorsal macrochaetotaxy (open circle—chaetal socket);
<emphasis id="B945A355FFAEFF9DFF0CF9DDFF756450" bold="true" box="[240,253,1609,1633]" pageId="3" pageNumber="374">5</emphasis>
, dorsal macrochaetotaxy of Th. II—Abd. V (wavy line—bothriotricha, open circle with a slash—pseudopore);
<emphasis id="B945A355FFAEFF9DFA75F9DDFA1E6450" bold="true" box="[1417,1430,1609,1633]" pageId="3" pageNumber="374">6</emphasis>
, trochantero-femoral organ (inner view);
<emphasis id="B945A355FFAEFF9DFDC8F9F9FDC964B4" bold="true" box="[564,577,1645,1669]" pageId="3" pageNumber="374">7</emphasis>
, tibiotarsi I, II and III (lateral view, showing tenent hairs and stronger inner chaetae only, other chaetae not shown, arrows pointing to blunt chaetae);
<emphasis id="B945A355FFAEFF9DFCCEF905FCB76498" bold="true" box="[818,831,1681,1705]" pageId="3" pageNumber="374">8</emphasis>
, hind claw complex (lateral view, showing chaetae on only one side except for accessory chaetae, t—tenent hair, aa—anterior accessory chaeta, ap—posterior accessory chaeta, g—guard chaeta, p—pretarsal chaeta);
<emphasis id="B945A355FFAEFF9DFE43F94DFE4464C0" bold="true" box="[447,460,1753,1777]" pageId="3" pageNumber="374">9</emphasis>
, tenaculum (anterior view).
</paragraph>
</caption>
<subSubSection id="C32B2CCCFFAEFF9DFF3BF8D7FDFC6594" pageId="3" pageNumber="374" type="etymology">
<paragraph id="8B8E7F47FFAEFF9DFF3BF8D7FCD3656C" blockId="3.[151,1436,1859,1958]" box="[199,859,1859,1886]" pageId="3" pageNumber="374">
<emphasis id="B945A355FFAEFF9DFF3BF8D7FEC6656C" bold="true" box="[199,334,1859,1885]" pageId="3" pageNumber="374">Etymology.</emphasis>
Named after the
<typeStatus id="548AC1E5FFAEFF9DFDECF8D0FDC8656F" box="[528,576,1860,1886]" pageId="3" pageNumber="374">type</typeStatus>
locality: Gyirong Valley.
</paragraph>
<paragraph id="8B8E7F47FFAEFF9DFF3BF8F3FDFC6594" blockId="3.[151,1436,1859,1958]" pageId="3" pageNumber="374">
<emphasis id="B945A355FFAEFF9DFF3BF8F3FEA265B0" bold="true" box="[199,298,1895,1921]" pageId="3" pageNumber="374">Habitat.</emphasis>
In leaf litter, rotten woods and live mosses in a coniferous and broad-leaved mixed forest; also under a wood board beside a cottage in a village.
</paragraph>
</subSubSection>
<caption id="DF4E2FCFFFA9FF9AFF6BFBE8FC336730" ID-DOI="http://doi.org/10.5281/zenodo.14735184" ID-Zenodo-Dep="14735184" httpUri="https://zenodo.org/record/14735184/files/figure.png" pageId="4" pageNumber="375" startId="4.[151,264,1148,1172]" targetBox="[159,1427,190,1124]" targetPageId="4" targetType="figure">
<paragraph id="8B8E7F47FFA9FF9AFF6BFBE8FC336730" blockId="4.[151,1437,1148,1281]" pageId="4" pageNumber="375">
<emphasis id="B945A355FFA9FF9AFF6BFBE8FEDD66A4" bold="true" box="[151,341,1148,1173]" pageId="4" pageNumber="375">FIGURES 10–14.</emphasis>
<taxonomicName id="4C3104C4FFA9FF9AFEA7FBE9FD7266A5" authorityName="Yu &amp; Xue &amp; Zhang" authorityYear="2025" box="[347,762,1149,1173]" class="Collembola" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Entomobryomorpha" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="gyirongensis" status="sp. nov." subGenus="Ocreatomurus">
<emphasis id="B945A355FFA9FF9AFEA7FBE9FE4166A5" box="[347,457,1149,1172]" italics="true" pageId="4" pageNumber="375">Tomocerus</emphasis>
(
<emphasis id="B945A355FFA9FF9AFE25FBE9FDE266A5" box="[473,618,1149,1172]" italics="true" pageId="4" pageNumber="375">Ocreatomurus</emphasis>
)
<emphasis id="B945A355FFA9FF9AFD85FBE9FD7266A5" box="[633,762,1149,1172]" italics="true" pageId="4" pageNumber="375">gyirongensis</emphasis>
</taxonomicName>
<emphasis id="B945A355FFA9FF9AFCFDFBE9FCDA66A4" bold="true" box="[769,850,1149,1173]" pageId="4" pageNumber="375">
<taxonomicNameLabel id="A2761E2EFFA9FF9AFCFDFBE9FCDA66A4" box="[769,850,1149,1173]" pageId="4" pageNumber="375" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
:
<emphasis id="B945A355FFA9FF9AFC9CFBE9FCF166A4" bold="true" box="[864,889,1149,1173]" pageId="4" pageNumber="375">10</emphasis>
, left side of manubrium (dorsal view, showing lateral chaetae and prominent chaetae, Λ—scale socket);
<emphasis id="B945A355FFA9FF9AFD5DFB35FD316688" bold="true" box="[673,697,1185,1209]" pageId="4" pageNumber="375">11</emphasis>
, distal area of manubrium (dorsal view, arrow pointing to distal corner chaeta, dashed arrow showing optional smaller chaeta);
<emphasis id="B945A355FFA9FF9AFD19FB51FD7666EC" bold="true" box="[741,766,1221,1245]" pageId="4" pageNumber="375">12</emphasis>
, proximal part of dens (dorsal view, showing dental spines and prominent chaeta);
<emphasis id="B945A355FFA9FF9AFEA0FB7DFEFD6730" bold="true" box="[348,373,1257,1281]" pageId="4" pageNumber="375">13</emphasis>
, examples of dental spines;
<emphasis id="B945A355FFA9FF9AFD69FB7DFD266730" bold="true" box="[661,686,1257,1281]" pageId="4" pageNumber="375">14</emphasis>
, right mucro (outer view).
</paragraph>
</caption>
<subSubSection id="C32B2CCCFFA9FF9AFF3BFAC7FCB5652C" pageId="4" pageNumber="375" type="discussion">
<paragraph id="8B8E7F47FFA9FF9AFF3BFAC7FCB5652C" blockId="4.[151,1436,1363,1821]" pageId="4" pageNumber="375">
<emphasis id="B945A355FFA9FF9AFF3BFAC7FEB3675C" bold="true" box="[199,315,1363,1389]" pageId="4" pageNumber="375">Remarks.</emphasis>
Within
<taxonomicName id="4C3104C4FFA9FF9AFE66FAC0FD5A675C" box="[410,722,1363,1389]" class="Collembola" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Entomobryomorpha" pageId="4" pageNumber="375" phylum="Arthropoda" rank="subGenus" subGenus="Ocreatomurus">
<emphasis id="B945A355FFA9FF9AFE66FAC0FD9C675C" box="[410,532,1364,1389]" italics="true" pageId="4" pageNumber="375">Tomocerus</emphasis>
(
<emphasis id="B945A355FFA9FF9AFDDAFAC7FD4F675C" box="[550,711,1363,1389]" italics="true" pageId="4" pageNumber="375">Ocreatomurus</emphasis>
)
</taxonomicName>
,
<taxonomicName id="4C3104C4FFA9FF9AFD1DFAC0FC4B675C" authorityName="Yu &amp; Xue &amp; Zhang" authorityYear="2025" box="[737,963,1363,1389]" class="Collembola" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Entomobryomorpha" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="gyirongensis" status="sp. nov." subGenus="Ocreatomurus">
<emphasis id="B945A355FFA9FF9AFD1DFAC0FD7E675C" box="[737,758,1364,1389]" italics="true" pageId="4" pageNumber="375">T.</emphasis>
(
<emphasis id="B945A355FFA9FF9AFCF4FAC7FCAB675C" box="[776,803,1363,1389]" italics="true" pageId="4" pageNumber="375">O.</emphasis>
)
<emphasis id="B945A355FFA9FF9AFCC9FAC0FC4B675C" box="[821,963,1364,1389]" italics="true" pageId="4" pageNumber="375">gyirongensis</emphasis>
</taxonomicName>
<emphasis id="B945A355FFA9FF9AFC30FAC7FBA0675C" bold="true" box="[972,1064,1363,1389]" pageId="4" pageNumber="375">
<taxonomicNameLabel id="A2761E2EFFA9FF9AFC30FAC7FBA0675C" box="[972,1064,1363,1389]" pageId="4" pageNumber="375" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
belongs to a species group with only one large distal dental spine. This group currently contains
<taxonomicName id="4C3104C4FFA9FF9AFC9FFAECFAC167A0" authority="Chen &amp; Christiansen, 1998" authorityName="Chen &amp; Christiansen" authorityYear="1998" box="[867,1353,1399,1425]" class="Collembola" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Entomobryomorpha" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="spinulus" subGenus="Ocreatomurus">
<emphasis id="B945A355FFA9FF9AFC9FFAECFCF067A0" box="[867,888,1400,1425]" italics="true" pageId="4" pageNumber="375">T.</emphasis>
(
<emphasis id="B945A355FFA9FF9AFC7BFAE3FC2A67A0" box="[903,930,1399,1425]" italics="true" pageId="4" pageNumber="375">O.</emphasis>
)
<emphasis id="B945A355FFA9FF9AFC4EFAE3FB8767A0" box="[946,1039,1399,1425]" italics="true" pageId="4" pageNumber="375">spinulus</emphasis>
<bibRefCitation id="EFA002B6FFA9FF9AFBEAFAE3FAC167A0" author="Chen, J. X. &amp; Christiansen, K. A." box="[1046,1353,1399,1425]" pageId="4" pageNumber="375" pagination="51 - 55" refId="ref5063" refString="Chen, J. X. &amp; Christiansen, K. A. (1998) Tomocerus (s. s.) spinulus (Collembola: Entomobryidae), a new species of Chinese springtail. Entomological News, 109, 51 - 55." type="journal article" year="1998">Chen &amp; Christiansen, 1998</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C3104C4FFA9FF9AFAAFFAECFE746784" authority="Yu &amp; Deharveng, 2018" authorityName="Yu &amp; Deharveng" authorityYear="2018" class="Collembola" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Entomobryomorpha" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="leyensis" subGenus="Ocreatomurus">
<emphasis id="B945A355FFA9FF9AFAAFFAECFAE067A0" box="[1363,1384,1400,1425]" italics="true" pageId="4" pageNumber="375">T.</emphasis>
(
<emphasis id="B945A355FFA9FF9AFA84FAE3FA1B67A0" box="[1400,1427,1399,1425]" italics="true" pageId="4" pageNumber="375">O.</emphasis>
)
<emphasis id="B945A355FFA9FF9AFF6BFA0FFF786784" box="[151,240,1435,1461]" italics="true" pageId="4" pageNumber="375">leyensis</emphasis>
<bibRefCitation id="EFA002B6FFA9FF9AFF0BFA0FFE746784" author="Yu, D. &amp; Deharveng, L." box="[247,508,1435,1461]" pageId="4" pageNumber="375" pagination="50 - 60" refId="ref5291" refString="Yu, D. &amp; Deharveng, L. (2018) Two new Chinese species of Tomocerus at different stages of troglomorphic adaptation (Collembola, Tomoceridae). Zootaxa, 4403 (1), 50 - 60. https: // doi. org / 10.11646 / zootaxa. 4403.1.2" type="journal article" year="2018">Yu &amp; Deharveng, 2018</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C3104C4FFA9FF9AFDFBFA08FB806784" authority="Gong, Qin &amp; Yu, 2018" authorityName="Gong, Qin &amp; Yu" authorityYear="2018" box="[519,1032,1435,1461]" class="Collembola" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Entomobryomorpha" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="pseudospinulus" subGenus="Ocreatomurus">
<emphasis id="B945A355FFA9FF9AFDFBFA08FD946784" box="[519,540,1436,1461]" italics="true" pageId="4" pageNumber="375">T.</emphasis>
(
<emphasis id="B945A355FFA9FF9AFDD0FA0FFDCF6784" box="[556,583,1435,1461]" italics="true" pageId="4" pageNumber="375">O.</emphasis>
)
<emphasis id="B945A355FFA9FF9AFDABFA0FFC8C6784" box="[599,772,1435,1461]" italics="true" pageId="4" pageNumber="375">pseudospinulus</emphasis>
<bibRefCitation id="EFA002B6FFA9FF9AFCF7FA0FFB806784" author="Gong, X. &amp; Qin, C. &amp; Yu, D." box="[779,1032,1435,1461]" pageId="4" pageNumber="375" pagination="273 - 282" refId="ref5200" refString="Gong, X., Qin, C. &amp; Yu, D. (2018) Two new species of Tomocerus ocreatus group with a single large distal dental spine (Collembola, Tomoceridae). Zootaxa, 4514 (2), 273 - 282. https: // doi. org / 10.11646 / zootaxa. 4514.2.10" type="journal article" year="2018">Gong, Qin &amp; Yu, 2018</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C3104C4FFA9FF9AFBC1FA08FE9367E8" authority="Gong, Qin &amp; Yu, 2018" authorityName="Gong, Qin &amp; Yu" authorityYear="2018" class="Collembola" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Entomobryomorpha" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="paraspinulus" subGenus="Ocreatomurus">
<emphasis id="B945A355FFA9FF9AFBC1FA08FBDB6784" box="[1085,1107,1436,1461]" italics="true" pageId="4" pageNumber="375">T.</emphasis>
(
<emphasis id="B945A355FFA9FF9AFB9EFA0FFBF56784" box="[1122,1149,1435,1461]" italics="true" pageId="4" pageNumber="375">O.</emphasis>
)
<emphasis id="B945A355FFA9FF9AFB71FA0FFA976784" box="[1165,1311,1435,1461]" italics="true" pageId="4" pageNumber="375">paraspinulus</emphasis>
<bibRefCitation id="EFA002B6FFA9FF9AFADAFA0FFE9367E8" author="Gong, X. &amp; Qin, C. &amp; Yu, D." pageId="4" pageNumber="375" pagination="273 - 282" refId="ref5200" refString="Gong, X., Qin, C. &amp; Yu, D. (2018) Two new species of Tomocerus ocreatus group with a single large distal dental spine (Collembola, Tomoceridae). Zootaxa, 4514 (2), 273 - 282. https: // doi. org / 10.11646 / zootaxa. 4514.2.10" type="journal article" year="2018">Gong, Qin &amp; Yu, 2018</bibRefCitation>
</taxonomicName>
. Among them,
<taxonomicName id="4C3104C4FFA9FF9AFE3EFA54FD1767E8" authorityName="Yu &amp; Xue &amp; Zhang" authorityYear="2025" box="[450,671,1471,1497]" class="Collembola" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Entomobryomorpha" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="gyirongensis" status="sp. nov." subGenus="Ocreatomurus">
<emphasis id="B945A355FFA9FF9AFE3EFA54FE5F67E8" box="[450,471,1472,1497]" italics="true" pageId="4" pageNumber="375">T.</emphasis>
(
<emphasis id="B945A355FFA9FF9AFE1AFA2BFD8A67E8" box="[486,514,1471,1497]" italics="true" pageId="4" pageNumber="375">O.</emphasis>
)
<emphasis id="B945A355FFA9FF9AFDEDFA54FD1767E8" box="[529,671,1472,1497]" italics="true" pageId="4" pageNumber="375">gyirongensis</emphasis>
</taxonomicName>
<emphasis id="B945A355FFA9FF9AFD5AFA2BFD7767E8" bold="true" box="[678,767,1471,1497]" pageId="4" pageNumber="375">
<taxonomicNameLabel id="A2761E2EFFA9FF9AFD5AFA2BFD7767E8" box="[678,767,1471,1497]" pageId="4" pageNumber="375" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is most similar to
<taxonomicName id="4C3104C4FFA9FF9AFC37FA54FBFF67E8" authorityName="Chen &amp; Christiansen" authorityYear="1998" box="[971,1143,1471,1497]" class="Collembola" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Entomobryomorpha" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="spinulus" subGenus="Ocreatomurus">
<emphasis id="B945A355FFA9FF9AFC37FA54FC6867E8" box="[971,992,1472,1497]" italics="true" pageId="4" pageNumber="375">T.</emphasis>
(
<emphasis id="B945A355FFA9FF9AFC13FA2BFB8267E8" box="[1007,1034,1471,1497]" italics="true" pageId="4" pageNumber="375">O.</emphasis>
)
<emphasis id="B945A355FFA9FF9AFBE6FA2BFBFF67E8" box="[1050,1143,1471,1497]" italics="true" pageId="4" pageNumber="375">spinulus</emphasis>
</taxonomicName>
in the presence of pointed prominent chaetae on manubrium and dens, and the small size of denticles on dental spines, but the new species can be distinguished from the later by the following characters (
<tableCitation id="C6B34AFCFFA9FF9AFC84F993FC476410" box="[888,975,1543,1569]" captionStart="TABLE 1" captionStartId="5.[152,244,151,175]" captionText="TABLE 1. Main characters distinguishing Tomocerus (Ocreatomurus) spinulus Chen &amp; Christiansen, 1998 (sensu Yu et al. 2018), T. (O.) gyirongensis sp. nov. and T. (O.) oppositus sp. nov." pageId="4" pageNumber="375">Table 1</tableCitation>
): first,
<taxonomicName id="4C3104C4FFA9FF9AFBDAF99CFA816410" authorityName="Yu &amp; Xue &amp; Zhang" authorityYear="2025" box="[1062,1289,1543,1569]" class="Collembola" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Entomobryomorpha" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="gyirongensis" status="sp. nov." subGenus="Ocreatomurus">
<emphasis id="B945A355FFA9FF9AFBDAF99CFBB36410" box="[1062,1083,1544,1569]" italics="true" pageId="4" pageNumber="375">T.</emphasis>
(
<emphasis id="B945A355FFA9FF9AFBB1F993FBE06410" box="[1101,1128,1543,1569]" italics="true" pageId="4" pageNumber="375">O.</emphasis>
)
<emphasis id="B945A355FFA9FF9AFB87F99CFA816410" box="[1147,1289,1544,1569]" italics="true" pageId="4" pageNumber="375">gyirongensis</emphasis>
</taxonomicName>
<emphasis id="B945A355FFA9FF9AFAEEF993FAE66410" bold="true" box="[1298,1390,1543,1569]" pageId="4" pageNumber="375">
<taxonomicNameLabel id="A2761E2EFFA9FF9AFAEEF993FAE66410" box="[1298,1390,1543,1569]" pageId="4" pageNumber="375" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a larger number of tibiotarsal strong inner chaetae than
<taxonomicName id="4C3104C4FFA9FF9AFCF2F9B8FC366474" authorityName="Chen &amp; Christiansen" authorityYear="1998" box="[782,958,1579,1605]" class="Collembola" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Entomobryomorpha" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="spinulus" subGenus="Ocreatomurus">
<emphasis id="B945A355FFA9FF9AFCF2F9B8FCAB6474" box="[782,803,1580,1605]" italics="true" pageId="4" pageNumber="375">T.</emphasis>
(
<emphasis id="B945A355FFA9FF9AFCC8F9BFFCC76474" box="[820,847,1579,1605]" italics="true" pageId="4" pageNumber="375">O.</emphasis>
)
<emphasis id="B945A355FFA9FF9AFC9DF9BFFC366474" box="[865,958,1579,1605]" italics="true" pageId="4" pageNumber="375">spinulus</emphasis>
</taxonomicName>
; second,
<taxonomicName id="4C3104C4FFA9FF9AFBD7F9B8FA846474" authorityName="Yu &amp; Xue &amp; Zhang" authorityYear="2025" box="[1067,1292,1579,1605]" class="Collembola" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Entomobryomorpha" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="gyirongensis" status="sp. nov." subGenus="Ocreatomurus">
<emphasis id="B945A355FFA9FF9AFBD7F9B8FBC86474" box="[1067,1088,1580,1605]" italics="true" pageId="4" pageNumber="375">T.</emphasis>
(
<emphasis id="B945A355FFA9FF9AFBADF9BFFBE46474" box="[1105,1132,1579,1605]" italics="true" pageId="4" pageNumber="375">O.</emphasis>
)
<emphasis id="B945A355FFA9FF9AFB82F9B8FA846474" box="[1150,1292,1580,1605]" italics="true" pageId="4" pageNumber="375">gyirongensis</emphasis>
</taxonomicName>
<emphasis id="B945A355FFA9FF9AFAE8F9BFFAE76474" bold="true" box="[1300,1391,1579,1605]" pageId="4" pageNumber="375">
<taxonomicNameLabel id="A2761E2EFFA9FF9AFAE8F9BFFAE76474" box="[1300,1391,1579,1605]" pageId="4" pageNumber="375" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has 2 + 2 prominent chaetae on manubrium, while
<taxonomicName id="4C3104C4FFA9FF9AFD59F9C4FCDA6458" authorityName="Chen &amp; Christiansen" authorityYear="1998" box="[677,850,1615,1641]" class="Collembola" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Entomobryomorpha" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="spinulus" subGenus="Ocreatomurus">
<emphasis id="B945A355FFA9FF9AFD59F9C4FD326458" box="[677,698,1616,1641]" italics="true" pageId="4" pageNumber="375">T.</emphasis>
(
<emphasis id="B945A355FFA9FF9AFD36F9DBFD6D6458" box="[714,741,1615,1641]" italics="true" pageId="4" pageNumber="375">O.</emphasis>
)
<emphasis id="B945A355FFA9FF9AFD09F9DBFCDA6458" box="[757,850,1615,1641]" italics="true" pageId="4" pageNumber="375">spinulus</emphasis>
</taxonomicName>
has only 1 + 1 of them (
<emphasis id="B945A355FFA9FF9AFB9BF9C4FB2D6458" box="[1127,1189,1616,1641]" italics="true" pageId="4" pageNumber="375">sensu</emphasis>
Yu
<emphasis id="B945A355FFA9FF9AFB2DF9C4FA816458" box="[1233,1289,1615,1641]" italics="true" pageId="4" pageNumber="375">et al.</emphasis>
2018); third, in
<taxonomicName id="4C3104C4FFA9FF9AFF4FF9E0FE0764BC" authorityName="Yu &amp; Xue &amp; Zhang" authorityYear="2025" box="[179,399,1651,1677]" class="Collembola" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Entomobryomorpha" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="gyirongensis" status="sp. nov." subGenus="Ocreatomurus">
<emphasis id="B945A355FFA9FF9AFF4FF9E0FF4064BC" box="[179,200,1652,1677]" italics="true" pageId="4" pageNumber="375">T.</emphasis>
(
<emphasis id="B945A355FFA9FF9AFF2BF9E7FF7A64BC" box="[215,242,1651,1677]" italics="true" pageId="4" pageNumber="375">O.</emphasis>
)
<emphasis id="B945A355FFA9FF9AFEFDF9E0FE0764BC" box="[257,399,1652,1677]" italics="true" pageId="4" pageNumber="375">gyirongensis</emphasis>
</taxonomicName>
<emphasis id="B945A355FFA9FF9AFE69F9E7FE6664BC" bold="true" box="[405,494,1651,1677]" pageId="4" pageNumber="375">
<taxonomicNameLabel id="A2761E2EFFA9FF9AFE69F9E7FE6664BC" box="[405,494,1651,1677]" pageId="4" pageNumber="375" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the proximal dental spines are arranged strictly in two rows, while in
<taxonomicName id="4C3104C4FFA9FF9AFB0DF9E0FA1464BC" authorityName="Chen &amp; Christiansen" authorityYear="1998" box="[1265,1436,1651,1677]" class="Collembola" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Entomobryomorpha" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="spinulus" subGenus="Ocreatomurus">
<emphasis id="B945A355FFA9FF9AFB0DF9E0FA8E64BC" box="[1265,1286,1652,1677]" italics="true" pageId="4" pageNumber="375">T.</emphasis>
(
<emphasis id="B945A355FFA9FF9AFAE8F9E7FAB864BC" box="[1300,1328,1651,1677]" italics="true" pageId="4" pageNumber="375">O.</emphasis>
)
<emphasis id="B945A355FFA9FF9AFAC3F9E7FA1464BC" box="[1343,1436,1651,1677]" italics="true" pageId="4" pageNumber="375">spinulus</emphasis>
</taxonomicName>
these spines are arranged in a single row. In addition to these major differences, the relative length of antennae in
<taxonomicName id="4C3104C4FFA9FF9AFF6BF928FEFF64E4" authorityName="Yu &amp; Xue &amp; Zhang" authorityYear="2025" box="[151,375,1723,1749]" class="Collembola" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Entomobryomorpha" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="gyirongensis" status="sp. nov." subGenus="Ocreatomurus">
<emphasis id="B945A355FFA9FF9AFF6BF928FF2464E4" box="[151,172,1724,1749]" italics="true" pageId="4" pageNumber="375">T.</emphasis>
(
<emphasis id="B945A355FFA9FF9AFF41F92FFF5064E4" box="[189,216,1723,1749]" italics="true" pageId="4" pageNumber="375">O.</emphasis>
)
<emphasis id="B945A355FFA9FF9AFF15F928FEFF64E4" box="[233,375,1724,1749]" italics="true" pageId="4" pageNumber="375">gyirongensis</emphasis>
</taxonomicName>
<emphasis id="B945A355FFA9FF9AFE83F92FFE5264E4" bold="true" box="[383,474,1723,1749]" pageId="4" pageNumber="375">
<taxonomicNameLabel id="A2761E2EFFA9FF9AFE83F92FFE5264E4" box="[383,474,1723,1749]" pageId="4" pageNumber="375" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is approximately 20–40 % (average 30 %) longer than that in
<taxonomicName id="4C3104C4FFA9FF9AFB61F928FAC464E4" authorityName="Chen &amp; Christiansen" authorityYear="1998" box="[1181,1356,1723,1749]" class="Collembola" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Entomobryomorpha" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="spinulus" subGenus="Ocreatomurus">
<emphasis id="B945A355FFA9FF9AFB61F928FB3A64E4" box="[1181,1202,1724,1749]" italics="true" pageId="4" pageNumber="375">T.</emphasis>
(
<emphasis id="B945A355FFA9FF9AFB3FF92FFB5664E4" box="[1219,1246,1723,1749]" italics="true" pageId="4" pageNumber="375">O.</emphasis>
)
<emphasis id="B945A355FFA9FF9AFB13F92FFAC464E4" box="[1263,1356,1723,1749]" italics="true" pageId="4" pageNumber="375">spinulus</emphasis>
</taxonomicName>
(
<emphasis id="B945A355FFA9FF9AFAA2F928FA1464E4" box="[1374,1436,1724,1749]" italics="true" pageId="4" pageNumber="375">sensu</emphasis>
Yu
<emphasis id="B945A355FFA9FF9AFF41F974FF7D64C8" box="[189,245,1759,1785]" italics="true" pageId="4" pageNumber="375">et al.</emphasis>
2018). Likely due to its larger body size,
<taxonomicName id="4C3104C4FFA9FF9AFD39F974FC2B64C8" authorityName="Yu &amp; Xue &amp; Zhang" authorityYear="2025" box="[709,931,1759,1785]" class="Collembola" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Entomobryomorpha" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="gyirongensis" status="sp. nov." subGenus="Ocreatomurus">
<emphasis id="B945A355FFA9FF9AFD39F974FD5264C8" box="[709,730,1760,1785]" italics="true" pageId="4" pageNumber="375">T.</emphasis>
(
<emphasis id="B945A355FFA9FF9AFD15F94BFC8D64C8" box="[745,773,1759,1785]" italics="true" pageId="4" pageNumber="375">O.</emphasis>
)
<emphasis id="B945A355FFA9FF9AFCE9F974FC2B64C8" box="[789,931,1760,1785]" italics="true" pageId="4" pageNumber="375">gyirongensis</emphasis>
</taxonomicName>
<emphasis id="B945A355FFA9FF9AFC55F94BFB8A64C8" bold="true" box="[937,1026,1759,1785]" pageId="4" pageNumber="375">
<taxonomicNameLabel id="A2761E2EFFA9FF9AFC55F94BFB8A64C8" box="[937,1026,1759,1785]" pageId="4" pageNumber="375" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
also has more chaetae on the ventral tube, tenaculum, and dorsal chaetal strips of the manubrium.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="A815CB242628D1B320C55EAF45F34AB8" ID-DOI="10.11646/zootaxa.5570.2.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="14735032" ID-ZooBank="2769486A-D505-4EAA-89DD-5E0C5894795D" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737737409727" checkinUser="plazi" docAuthor="Raphel, Sheeba &amp; Jose, Joyce" docDate="2025" docId="039E5268FFC0180AFF6F88EAAE13FBDE" docLanguage="en" docName="zootaxa.5570.2.4.pdf" docOrigin="Zootaxa 5570 (2)" docSource="https://doi.org/10.11646/zootaxa.5570.2.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ctenolepisma rothschildi , Silvestri 1907" docType="treatment" docVersion="2" lastPageNumber="321" masterDocId="FFA72A10FFC71806FFF8887DAD31FF98" masterDocTitle="Description of two synanthropic silverfish (Insecta: Zygentoma), Ctenolepisma longicaudatum (Escherich, 1905) and Ctenolepisma rothschildi (Silvestri, 1907), from Kerala, India" masterLastPageNumber="324" masterPageNumber="309" pageNumber="316" updateTime="1737737830484" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D114DA077FCE10CE749AC80D3B0EAF4E">Description of two synanthropic silverfish (Insecta: Zygentoma), Ctenolepisma longicaudatum (Escherich, 1905) and Ctenolepisma rothschildi (Silvestri, 1907), from Kerala, India</mods:title>
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<mods:namePart id="E6F5E462D6AF3EA9FD949DE3A25C83CC">Raphel, Sheeba</mods:namePart>
<mods:affiliation id="2577AB85A8CC9A836340BE3130A81F12">Department of Zoology, St. Thomas College (Autonomous), Thrissur affiliated to the University of Calicut, Kerala, India</mods:affiliation>
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<mods:namePart id="D1D85B46C11F502B53AC294974D415EC">Jose, Joyce</mods:namePart>
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<mods:title id="74FCE566428E7C1DAF39B1EAB3112D17">Zootaxa</mods:title>
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<mods:date id="AAA23EDF61B85CEBE2BE242CEAA3C6E7">2025</mods:date>
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<subSubSection id="C32DB0F5FFC01801FF6F88EAAF4CFF29" box="[151,637,151,178]" pageId="7" pageNumber="316" type="nomenclature">
<paragraph id="8B88E37EFFC01801FF6F88EAAF4CFF29" blockId="7.[151,637,151,214]" box="[151,637,151,178]" pageId="7" pageNumber="316">
<heading id="D0C05412FFC01801FF6F88EAAF4CFF29" bold="true" box="[151,637,151,178]" fontSize="11" level="1" pageId="7" pageNumber="316" reason="1">
<taxonomicName id="4C3798FDFFC01801FF6F88EAAF4CFF29" authority="Silvestri, 1907" authorityName=", Silvestri" authorityYear="1907" box="[151,637,151,178]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="7" pageNumber="316" phylum="Arthropoda" rank="species" species="rothschildi">
<emphasis id="B9433F6CFFC01801FF6F88EAAF4CFF29" bold="true" box="[151,637,151,178]" pageId="7" pageNumber="316">
<emphasis id="B9433F6CFFC01801FF6F88EAACF4FF2A" bold="true" box="[151,453,151,178]" italics="true" pageId="7" pageNumber="316">Ctenolepisma rothschildi</emphasis>
Silvestri, 1907
</emphasis>
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<paragraph id="8B88E37EFFC01801FF6F88C6AC1FFF4E" blockId="7.[151,637,151,214]" box="[151,302,187,214]" pageId="7" pageNumber="316">
(
<figureCitation id="130CFFFBFFC01801FF6788C6ADC8FF4E" box="[159,249,187,214]" captionStart="FIGURES 15‒17" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[210,1376,182,1910]" captionTargetId="figure-17@9.[203,1384,181,1919]" captionTargetPageId="9" captionText="FIGURES 15‒17. Ctenolepisma rothschildi. 15) Dorsal view. 16) Body scales. 17) SEM of labial sensory papillae." figureDoi="http://doi.org/10.5281/zenodo.14735044" httpUri="https://zenodo.org/record/14735044/files/figure.png" pageId="7" pageNumber="316">Figs. 15</figureCitation>
‒27)
</paragraph>
</subSubSection>
<subSubSection id="C32DB0F5FFC0180AFF6F897EAE13FBDE" lastPageId="12" lastPageNumber="321" pageId="7" pageNumber="316" type="materials_examined">
<paragraph id="8B88E37EFFC01801FF6F897EA9BFFEFD" blockId="7.[151,1436,259,358]" pageId="7" pageNumber="316">
<emphasis id="B9433F6CFFC01801FF6F897EACA3FE85" bold="true" box="[151,402,259,285]" pageId="7" pageNumber="316">Materials Examined.</emphasis>
<collectingCountry id="F320A3EEFFC01801FE61897EACE4FE85" box="[409,469,259,285]" name="India" pageId="7" pageNumber="316">India</collectingCountry>
,
<collectingRegion id="49F32D9CFFC01801FE19897EAF1FFE85" box="[481,558,259,285]" country="India" name="Kerala" pageId="7" pageNumber="316">Kerala</collectingRegion>
, specimens from 44 sites (
<tableCitation id="C6B5D6C5FFC01801FCA0897EAE9FFE85" box="[856,942,259,285]" captionStart="TABLE 3" captionStartId="7.[151,239,405,429]" captionText="TABLE 3. List of collection sites of Ctenolepisma rothschildi with geocoordinates." pageId="7" pageNumber="316">Table 3</tableCitation>
), mostly from books and papers, S. Raphel, coll. Specimens in alcohol deposited at Zoological Survey of
<collectingCountry id="F320A3EEFFC01801FCBF895AAEB2FED9" box="[839,899,295,321]" name="India" pageId="7" pageNumber="316">India</collectingCountry>
, Western Ghat Regional Centre,
<collectingCountry id="F320A3EEFFC01801FB0A895AA85CFED9" box="[1266,1389,295,321]" name="India" pageId="7" pageNumber="316">Kozhikode</collectingCountry>
and the registration numbers are ZSI/
<collectionCode id="ED267BBBFFC01801FDF38936AF52FEFD" box="[523,611,331,357]" pageId="7" pageNumber="316">WGRC</collectionCode>
/I.
<collectionCode id="ED267BBBFFC01801FD808936AFBAFEFD" box="[632,651,331,357]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="316">R</collectionCode>
.-INV.26926 and ZSI/
<collectionCode id="ED267BBBFFC01801FC868936AEE6FEFD" box="[894,983,331,357]" pageId="7" pageNumber="316">WGRC</collectionCode>
/I.
<collectionCode id="ED267BBBFFC01801FC148936AECEFEFD" box="[1004,1023,331,357]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="316">R</collectionCode>
.-INV.26927.
</paragraph>
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<paragraph id="8B88E37EFFC01801FF6F89E8A923FE35" blockId="7.[151,1042,405,429]" box="[151,1042,405,429]" pageId="7" pageNumber="316">
<emphasis id="B9433F6CFFC01801FF6F89E8AC3BFE35" bold="true" box="[151,266,405,429]" pageId="7" pageNumber="316">TABLE 3.</emphasis>
List of collection sites of
<taxonomicName id="4C3798FDFFC01801FDD989E8AE1CFE35" authorityName=", Silvestri" authorityYear="1907" box="[545,813,405,429]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="7" pageNumber="316" phylum="Arthropoda" rank="species" species="rothschildi">
<emphasis id="B9433F6CFFC01801FDD989E8AE1CFE35" box="[545,813,405,429]" italics="true" pageId="7" pageNumber="316">Ctenolepisma rothschildi</emphasis>
</taxonomicName>
with geocoordinates.
</paragraph>
</caption>
<paragraph id="8B88E37EFFC01801FF5B89C3A9C8F83E" pageId="7" pageNumber="316">
<table id="F93711DEFFC0E7F9FF5B89C0A807F83F" box="[163,1334,445,1959]" colsContinueFrom="3.[163,1357,193,740]" colsContinueIn="8.[163,1334,202,509]" gridcols="5" gridrows="38" pageId="7" pageNumber="316">
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<th id="76D68840FFC0E7F9FF5B89C0ADF8FE4D" box="[163,201,445,469]" gridcol="0" gridrow="0" pageId="7" pageNumber="316">No.</th>
<th id="76D68840FFC0E7F9FE5C89C0AF39FE4D" box="[420,520,445,469]" gridcol="1" gridrow="0" pageId="7" pageNumber="316">District</th>
<th id="76D68840FFC0E7F9FD5D89C0AE64FE4D" box="[677,853,445,469]" gridcol="2" gridrow="0" pageId="7" pageNumber="316">Location</th>
<th id="76D68840FFC0E7F9FC5E89C0A912FE4D" box="[934,1059,445,469]" gridcol="3" gridrow="0" pageId="7" pageNumber="316">Latitude (N)</th>
<th id="76D68840FFC0E7F9FB5F89C0A807FE4D" box="[1191,1334,445,469]" gridcol="4" gridrow="0" pageId="7" pageNumber="316">Longitude (E)</th>
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<th id="76D68840FFC0E7F9FF5B8998ADF8FE65" box="[163,201,485,509]" gridcol="0" gridrow="1" pageId="7" pageNumber="316">1</th>
<td id="76D68840FFC0E7F9FE5C8998AF39FE65" box="[420,520,485,509]" gridcol="1" gridrow="1" pageId="7" pageNumber="316">Thrissur</td>
<td id="76D68840FFC0E7F9FD5D8998AE64FE65" box="[677,853,485,509]" gridcol="2" gridrow="1" pageId="7" pageNumber="316">Puthur</td>
<td id="76D68840FFC0E7F9FC5E8998A912FE65" box="[934,1059,485,509]" gridcol="3" gridrow="1" pageId="7" pageNumber="316">10.4904</td>
<td id="76D68840FFC0E7F9FB5F8998A807FE65" box="[1191,1334,485,509]" gridcol="4" gridrow="1" pageId="7" pageNumber="316">76.2819</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8A70A807FDBE" box="[163,1334,525,550]" gridrow="2" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8A70ADF8FDBE" box="[163,201,525,550]" gridcol="0" gridrow="2" pageId="7" pageNumber="316">2</th>
<td id="76D68840FFC0E7F9FE5C8A70AF39FDBE" box="[420,520,525,550]" gridcol="1" gridrow="2" pageId="7" pageNumber="316">Thrissur</td>
<td id="76D68840FFC0E7F9FD5D8A70AE64FDBE" box="[677,853,525,550]" gridcol="2" gridrow="2" pageId="7" pageNumber="316">Thozhiyoor</td>
<td id="76D68840FFC0E7F9FC5E8A70A912FDBE" box="[934,1059,525,550]" gridcol="3" gridrow="2" pageId="7" pageNumber="316">10.6317</td>
<td id="76D68840FFC0E7F9FB5F8A70A807FDBE" box="[1191,1334,525,550]" gridcol="4" gridrow="2" pageId="7" pageNumber="316">76.0258</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8A4BA807FDD6" box="[163,1334,566,590]" gridrow="3" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8A4BADF8FDD6" box="[163,201,566,590]" gridcol="0" gridrow="3" pageId="7" pageNumber="316">3</th>
<td id="76D68840FFC0E7F9FE5C8A4BAF39FDD6" box="[420,520,566,590]" gridcol="1" gridrow="3" pageId="7" pageNumber="316">Thrissur</td>
<td id="76D68840FFC0E7F9FD5D8A4BAE64FDD6" box="[677,853,566,590]" gridcol="2" gridrow="3" pageId="7" pageNumber="316">Peringottukara</td>
<td id="76D68840FFC0E7F9FC5E8A4BA912FDD6" box="[934,1059,566,590]" gridcol="3" gridrow="3" pageId="7" pageNumber="316">10.4448</td>
<td id="76D68840FFC0E7F9FB5F8A4BA807FDD6" box="[1191,1334,566,590]" gridcol="4" gridrow="3" pageId="7" pageNumber="316">76.1607</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8A23A807FDEE" box="[163,1334,606,630]" gridrow="4" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8A23ADF8FDEE" box="[163,201,606,630]" gridcol="0" gridrow="4" pageId="7" pageNumber="316">4</th>
<td id="76D68840FFC0E7F9FE5C8A23AF39FDEE" box="[420,520,606,630]" gridcol="1" gridrow="4" pageId="7" pageNumber="316">Thrissur</td>
<td id="76D68840FFC0E7F9FD5D8A23AE64FDEE" box="[677,853,606,630]" gridcol="2" gridrow="4" pageId="7" pageNumber="316">Kattilapoovam</td>
<td id="76D68840FFC0E7F9FC5E8A23A912FDEE" box="[934,1059,606,630]" gridcol="3" gridrow="4" pageId="7" pageNumber="316">10.573</td>
<td id="76D68840FFC0E7F9FB5F8A23A807FDEE" box="[1191,1334,606,630]" gridcol="4" gridrow="4" pageId="7" pageNumber="316">76.2851</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8AFBA807FD06" box="[163,1334,646,670]" gridrow="5" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8AFBADF8FD06" box="[163,201,646,670]" gridcol="0" gridrow="5" pageId="7" pageNumber="316">5</th>
<td id="76D68840FFC0E7F9FE5C8AFBAF39FD06" box="[420,520,646,670]" gridcol="1" gridrow="5" pageId="7" pageNumber="316">Thrissur</td>
<td id="76D68840FFC0E7F9FD5D8AFBAE64FD06" box="[677,853,646,670]" gridcol="2" gridrow="5" pageId="7" pageNumber="316">Kannara</td>
<td id="76D68840FFC0E7F9FC5E8AFBA912FD06" box="[934,1059,646,670]" gridcol="3" gridrow="5" pageId="7" pageNumber="316">10.5356</td>
<td id="76D68840FFC0E7F9FB5F8AFBA807FD06" box="[1191,1334,646,670]" gridcol="4" gridrow="5" pageId="7" pageNumber="316">76.3362</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8AD3A807FD5F" box="[163,1334,686,711]" gridrow="6" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8AD3ADF8FD5F" box="[163,201,686,711]" gridcol="0" gridrow="6" pageId="7" pageNumber="316">6</th>
<td id="76D68840FFC0E7F9FE5C8AD3AF39FD5F" box="[420,520,686,711]" gridcol="1" gridrow="6" pageId="7" pageNumber="316">Thrissur</td>
<td id="76D68840FFC0E7F9FD5D8AD3AE64FD5F" box="[677,853,686,711]" gridcol="2" gridrow="6" pageId="7" pageNumber="316">Kombazha</td>
<td id="76D68840FFC0E7F9FC5E8AD3A912FD5F" box="[934,1059,686,711]" gridcol="3" gridrow="6" pageId="7" pageNumber="316">10.5769</td>
<td id="76D68840FFC0E7F9FB5F8AD3A807FD5F" box="[1191,1334,686,711]" gridcol="4" gridrow="6" pageId="7" pageNumber="316">76.3974</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8AAAA807FD77" box="[163,1334,727,751]" gridrow="7" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8AAAADF8FD77" box="[163,201,727,751]" gridcol="0" gridrow="7" pageId="7" pageNumber="316">7</th>
<td id="76D68840FFC0E7F9FE5C8AAAAF39FD77" box="[420,520,727,751]" gridcol="1" gridrow="7" pageId="7" pageNumber="316">Thrissur</td>
<td id="76D68840FFC0E7F9FD5D8AAAAE64FD77" box="[677,853,727,751]" gridcol="2" gridrow="7" pageId="7" pageNumber="316">Eastfort, Thrissur</td>
<td id="76D68840FFC0E7F9FC5E8AAAA912FD77" box="[934,1059,727,751]" gridcol="3" gridrow="7" pageId="7" pageNumber="316">10.5223</td>
<td id="76D68840FFC0E7F9FB5F8AAAA807FD77" box="[1191,1334,727,751]" gridcol="4" gridrow="7" pageId="7" pageNumber="316">76.2238</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8A82A807FC8F" box="[163,1334,767,791]" gridrow="8" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8A82ADF8FC8F" box="[163,201,767,791]" gridcol="0" gridrow="8" pageId="7" pageNumber="316">8</th>
<td id="76D68840FFC0E7F9FE5C8A82AF39FC8F" box="[420,520,767,791]" gridcol="1" gridrow="8" pageId="7" pageNumber="316">Thrissur</td>
<td id="76D68840FFC0E7F9FD5D8A82AE64FC8F" box="[677,853,767,791]" gridcol="2" gridrow="8" pageId="7" pageNumber="316">Chembukkavu</td>
<td id="76D68840FFC0E7F9FC5E8A82A912FC8F" box="[934,1059,767,791]" gridcol="3" gridrow="8" pageId="7" pageNumber="316">10.5305</td>
<td id="76D68840FFC0E7F9FB5F8A82A807FC8F" box="[1191,1334,767,791]" gridcol="4" gridrow="8" pageId="7" pageNumber="316">76.2204</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8B5AA807FCA7" box="[163,1334,807,831]" gridrow="9" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8B5AADF8FCA7" box="[163,201,807,831]" gridcol="0" gridrow="9" pageId="7" pageNumber="316">9</th>
<td id="76D68840FFC0E7F9FE5C8B5AAF39FCA7" box="[420,520,807,831]" gridcol="1" gridrow="9" pageId="7" pageNumber="316">Thrissur</td>
<td id="76D68840FFC0E7F9FD5D8B5AAE64FCA7" box="[677,853,807,831]" gridcol="2" gridrow="9" pageId="7" pageNumber="316">Echippara</td>
<td id="76D68840FFC0E7F9FC5E8B5AA912FCA7" box="[934,1059,807,831]" gridcol="3" gridrow="9" pageId="7" pageNumber="316">10.4381</td>
<td id="76D68840FFC0E7F9FB5F8B5AA807FCA7" box="[1191,1334,807,831]" gridcol="4" gridrow="9" pageId="7" pageNumber="316">76.4504</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8B32A807FCF0" box="[163,1334,847,872]" gridrow="10" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8B32ADF8FCF0" box="[163,201,847,872]" gridcol="0" gridrow="10" pageId="7" pageNumber="316">10</th>
<td id="76D68840FFC0E7F9FE5C8B32AF39FCF0" box="[420,520,847,872]" gridcol="1" gridrow="10" pageId="7" pageNumber="316">Thrissur</td>
<td id="76D68840FFC0E7F9FD5D8B32AE64FCF0" box="[677,853,847,872]" gridcol="2" gridrow="10" pageId="7" pageNumber="316">Velupadam</td>
<td id="76D68840FFC0E7F9FC5E8B32A912FCF0" box="[934,1059,847,872]" gridcol="3" gridrow="10" pageId="7" pageNumber="316">10.4328</td>
<td id="76D68840FFC0E7F9FB5F8B32A807FCF0" box="[1191,1334,847,872]" gridcol="4" gridrow="10" pageId="7" pageNumber="316">76.3474</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8B05A807FC08" box="[163,1334,888,912]" gridrow="11" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8B05ADF8FC08" box="[163,201,888,912]" gridcol="0" gridrow="11" pageId="7" pageNumber="316">11</th>
<td id="76D68840FFC0E7F9FE5C8B05AF39FC08" box="[420,520,888,912]" gridcol="1" gridrow="11" pageId="7" pageNumber="316">Thrissur</td>
<td id="76D68840FFC0E7F9FD5D8B05AE64FC08" box="[677,853,888,912]" gridcol="2" gridrow="11" pageId="7" pageNumber="316">Madayikonam</td>
<td id="76D68840FFC0E7F9FC5E8B05A912FC08" box="[934,1059,888,912]" gridcol="3" gridrow="11" pageId="7" pageNumber="316">10.3681</td>
<td id="76D68840FFC0E7F9FB5F8B05A807FC08" box="[1191,1334,888,912]" gridcol="4" gridrow="11" pageId="7" pageNumber="316">76.2382</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8BDDA807FC20" box="[163,1334,928,952]" gridrow="12" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8BDDADF8FC20" box="[163,201,928,952]" gridcol="0" gridrow="12" pageId="7" pageNumber="316">12</th>
<td id="76D68840FFC0E7F9FE5C8BDDAF39FC20" box="[420,520,928,952]" gridcol="1" gridrow="12" pageId="7" pageNumber="316">Thrissur</td>
<td id="76D68840FFC0E7F9FD5D8BDDAE64FC20" box="[677,853,928,952]" gridcol="2" gridrow="12" pageId="7" pageNumber="316">Kallettumkara</td>
<td id="76D68840FFC0E7F9FC5E8BDDA912FC20" box="[934,1059,928,952]" gridcol="3" gridrow="12" pageId="7" pageNumber="316">10.3462</td>
<td id="76D68840FFC0E7F9FB5F8BDDA807FC20" box="[1191,1334,928,952]" gridcol="4" gridrow="12" pageId="7" pageNumber="316">76.2786</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8BB5A807FC78" box="[163,1334,968,992]" gridrow="13" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8BB5ADF8FC78" box="[163,201,968,992]" gridcol="0" gridrow="13" pageId="7" pageNumber="316">13</th>
<td id="76D68840FFC0E7F9FE5C8BB5AF39FC78" box="[420,520,968,992]" gridcol="1" gridrow="13" pageId="7" pageNumber="316">Thrissur</td>
<td id="76D68840FFC0E7F9FD5D8BB5AE64FC78" box="[677,853,968,992]" gridcol="2" gridrow="13" pageId="7" pageNumber="316">West Mangad</td>
<td id="76D68840FFC0E7F9FC5E8BB5A912FC78" box="[934,1059,968,992]" gridcol="3" gridrow="13" pageId="7" pageNumber="316">10.6766</td>
<td id="76D68840FFC0E7F9FB5F8BB5A807FC78" box="[1191,1334,968,992]" gridcol="4" gridrow="13" pageId="7" pageNumber="316">76.0513</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8B8DA807FB91" box="[163,1334,1008,1033]" gridrow="14" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8B8DADF8FB91" box="[163,201,1008,1033]" gridcol="0" gridrow="14" pageId="7" pageNumber="316">14</th>
<td id="76D68840FFC0E7F9FE5C8B8DAF39FB91" box="[420,520,1008,1033]" gridcol="1" gridrow="14" pageId="7" pageNumber="316">Thrissur</td>
<td id="76D68840FFC0E7F9FD5D8B8DAE64FB91" box="[677,853,1008,1033]" gridcol="2" gridrow="14" pageId="7" pageNumber="316">Puranattukara</td>
<td id="76D68840FFC0E7F9FC5E8B8DA912FB91" box="[934,1059,1008,1033]" gridcol="3" gridrow="14" pageId="7" pageNumber="316">10.5499</td>
<td id="76D68840FFC0E7F9FB5F8B8DA807FB91" box="[1191,1334,1008,1033]" gridcol="4" gridrow="14" pageId="7" pageNumber="316">76.1564</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8C64A807FBA9" box="[163,1334,1049,1073]" gridrow="15" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8C64ADF8FBA9" box="[163,201,1049,1073]" gridcol="0" gridrow="15" pageId="7" pageNumber="316">15</th>
<td id="76D68840FFC0E7F9FE5C8C64AF39FBA9" box="[420,520,1049,1073]" gridcol="1" gridrow="15" pageId="7" pageNumber="316">Thrissur</td>
<td id="76D68840FFC0E7F9FD5D8C64AE64FBA9" box="[677,853,1049,1073]" gridcol="2" gridrow="15" pageId="7" pageNumber="316">Valarkkavu</td>
<td id="76D68840FFC0E7F9FC5E8C64A912FBA9" box="[934,1059,1049,1073]" gridcol="3" gridrow="15" pageId="7" pageNumber="316">10.506</td>
<td id="76D68840FFC0E7F9FB5F8C64A807FBA9" box="[1191,1334,1049,1073]" gridcol="4" gridrow="15" pageId="7" pageNumber="316">76.2365</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8C3CA807FBC1" box="[163,1334,1089,1113]" gridrow="16" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8C3CADF8FBC1" box="[163,201,1089,1113]" gridcol="0" gridrow="16" pageId="7" pageNumber="316">16</th>
<td id="76D68840FFC0E7F9FE5C8C3CAF39FBC1" box="[420,520,1089,1113]" gridcol="1" gridrow="16" pageId="7" pageNumber="316">Thrissur</td>
<td id="76D68840FFC0E7F9FD5D8C3CAE64FBC1" box="[677,853,1089,1113]" gridcol="2" gridrow="16" pageId="7" pageNumber="316">Padukad</td>
<td id="76D68840FFC0E7F9FC5E8C3CA912FBC1" box="[934,1059,1089,1113]" gridcol="3" gridrow="16" pageId="7" pageNumber="316">10.5621</td>
<td id="76D68840FFC0E7F9FB5F8C3CA807FBC1" box="[1191,1334,1089,1113]" gridcol="4" gridrow="16" pageId="7" pageNumber="316">76.2152</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8C14A807FB19" box="[163,1334,1129,1153]" gridrow="17" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8C14ADF8FB19" box="[163,201,1129,1153]" gridcol="0" gridrow="17" pageId="7" pageNumber="316">17</th>
<td id="76D68840FFC0E7F9FE5C8C14AF39FB19" box="[420,520,1129,1153]" gridcol="1" gridrow="17" pageId="7" pageNumber="316">Palakkad</td>
<td id="76D68840FFC0E7F9FD5D8C14AE64FB19" box="[677,853,1129,1153]" gridcol="2" gridrow="17" pageId="7" pageNumber="316">Alathur</td>
<td id="76D68840FFC0E7F9FC5E8C14A912FB19" box="[934,1059,1129,1153]" gridcol="3" gridrow="17" pageId="7" pageNumber="316">10.6359</td>
<td id="76D68840FFC0E7F9FB5F8C14A807FB19" box="[1191,1334,1129,1153]" gridcol="4" gridrow="17" pageId="7" pageNumber="316">76.5358</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8CECA807FB32" box="[163,1334,1169,1194]" gridrow="18" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8CECADF8FB32" box="[163,201,1169,1194]" gridcol="0" gridrow="18" pageId="7" pageNumber="316">18</th>
<td id="76D68840FFC0E7F9FE5C8CECAF39FB32" box="[420,520,1169,1194]" gridcol="1" gridrow="18" pageId="7" pageNumber="316">Palakkad</td>
<td id="76D68840FFC0E7F9FD5D8CECAE64FB32" box="[677,853,1169,1194]" gridcol="2" gridrow="18" pageId="7" pageNumber="316">Alathur</td>
<td id="76D68840FFC0E7F9FC5E8CECA912FB32" box="[934,1059,1169,1194]" gridcol="3" gridrow="18" pageId="7" pageNumber="316">10.6371</td>
<td id="76D68840FFC0E7F9FB5F8CECA807FB32" box="[1191,1334,1169,1194]" gridcol="4" gridrow="18" pageId="7" pageNumber="316">76.5319</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8CC7A807FB4A" box="[163,1334,1210,1234]" gridrow="19" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8CC7ADF8FB4A" box="[163,201,1210,1234]" gridcol="0" gridrow="19" pageId="7" pageNumber="316">19</th>
<td id="76D68840FFC0E7F9FE5C8CC7AF39FB4A" box="[420,520,1210,1234]" gridcol="1" gridrow="19" pageId="7" pageNumber="316">Palakkad</td>
<td id="76D68840FFC0E7F9FD5D8CC7AE64FB4A" box="[677,853,1210,1234]" gridcol="2" gridrow="19" pageId="7" pageNumber="316">Alathur</td>
<td id="76D68840FFC0E7F9FC5E8CC7A912FB4A" box="[934,1059,1210,1234]" gridcol="3" gridrow="19" pageId="7" pageNumber="316">10.634</td>
<td id="76D68840FFC0E7F9FB5F8CC7A807FB4A" box="[1191,1334,1210,1234]" gridcol="4" gridrow="19" pageId="7" pageNumber="316">76.5311</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8C9FA807FB62" box="[163,1334,1250,1274]" gridrow="20" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8C9FADF8FB62" box="[163,201,1250,1274]" gridcol="0" gridrow="20" pageId="7" pageNumber="316">20</th>
<td id="76D68840FFC0E7F9FE5C8C9FAF39FB62" box="[420,520,1250,1274]" gridcol="1" gridrow="20" pageId="7" pageNumber="316">Palakkad</td>
<td id="76D68840FFC0E7F9FD5D8C9FAE64FB62" box="[677,853,1250,1274]" gridcol="2" gridrow="20" pageId="7" pageNumber="316">Nooradi</td>
<td id="76D68840FFC0E7F9FC5E8C9FA912FB62" box="[934,1059,1250,1274]" gridcol="3" gridrow="20" pageId="7" pageNumber="316">10.505</td>
<td id="76D68840FFC0E7F9FB5F8C9FA807FB62" box="[1191,1334,1250,1274]" gridcol="4" gridrow="20" pageId="7" pageNumber="316">76.6744</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8D77A807FABA" box="[163,1334,1290,1314]" gridrow="21" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8D77ADF8FABA" box="[163,201,1290,1314]" gridcol="0" gridrow="21" pageId="7" pageNumber="316">21</th>
<td id="76D68840FFC0E7F9FE5C8D77AF39FABA" box="[420,520,1290,1314]" gridcol="1" gridrow="21" pageId="7" pageNumber="316">Palakkad</td>
<td id="76D68840FFC0E7F9FD5D8D77AE64FABA" box="[677,853,1290,1314]" gridcol="2" gridrow="21" pageId="7" pageNumber="316">Nenmara</td>
<td id="76D68840FFC0E7F9FC5E8D77A912FABA" box="[934,1059,1290,1314]" gridcol="3" gridrow="21" pageId="7" pageNumber="316">10.595</td>
<td id="76D68840FFC0E7F9FB5F8D77A807FABA" box="[1191,1334,1290,1314]" gridcol="4" gridrow="21" pageId="7" pageNumber="316">76.5969</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8D4FA807FAD3" box="[163,1334,1330,1355]" gridrow="22" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8D4FADF8FAD3" box="[163,201,1330,1355]" gridcol="0" gridrow="22" pageId="7" pageNumber="316">22</th>
<td id="76D68840FFC0E7F9FE5C8D4FAF39FAD3" box="[420,520,1330,1355]" gridcol="1" gridrow="22" pageId="7" pageNumber="316">Palakkad</td>
<td id="76D68840FFC0E7F9FD5D8D4FAE64FAD3" box="[677,853,1330,1355]" gridcol="2" gridrow="22" pageId="7" pageNumber="316">Nenmara</td>
<td id="76D68840FFC0E7F9FC5E8D4FA912FAD3" box="[934,1059,1330,1355]" gridcol="3" gridrow="22" pageId="7" pageNumber="316">10.5954</td>
<td id="76D68840FFC0E7F9FB5F8D4FA807FAD3" box="[1191,1334,1330,1355]" gridcol="4" gridrow="22" pageId="7" pageNumber="316">76.5962</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8D26A807FAEB" box="[163,1334,1371,1395]" gridrow="23" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8D26ADF8FAEB" box="[163,201,1371,1395]" gridcol="0" gridrow="23" pageId="7" pageNumber="316">23</th>
<td id="76D68840FFC0E7F9FE5C8D26AF39FAEB" box="[420,520,1371,1395]" gridcol="1" gridrow="23" pageId="7" pageNumber="316">Palakkad</td>
<td id="76D68840FFC0E7F9FD5D8D26AE64FAEB" box="[677,853,1371,1395]" gridcol="2" gridrow="23" pageId="7" pageNumber="316">Kizhakkenchery</td>
<td id="76D68840FFC0E7F9FC5E8D26A912FAEB" box="[934,1059,1371,1395]" gridcol="3" gridrow="23" pageId="7" pageNumber="316">10.5399</td>
<td id="76D68840FFC0E7F9FB5F8D26A807FAEB" box="[1191,1334,1371,1395]" gridcol="4" gridrow="23" pageId="7" pageNumber="316">76.4679</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8DFEA807FA03" box="[163,1334,1411,1435]" gridrow="24" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8DFEADF8FA03" box="[163,201,1411,1435]" gridcol="0" gridrow="24" pageId="7" pageNumber="316">24</th>
<td id="76D68840FFC0E7F9FE5C8DFEAF39FA03" box="[420,520,1411,1435]" gridcol="1" gridrow="24" pageId="7" pageNumber="316">Kannur</td>
<td id="76D68840FFC0E7F9FD5D8DFEAE64FA03" box="[677,853,1411,1435]" gridcol="2" gridrow="24" pageId="7" pageNumber="316">Padiyur</td>
<td id="76D68840FFC0E7F9FC5E8DFEA912FA03" box="[934,1059,1411,1435]" gridcol="3" gridrow="24" pageId="7" pageNumber="316">11.9944</td>
<td id="76D68840FFC0E7F9FB5F8DFEA807FA03" box="[1191,1334,1411,1435]" gridcol="4" gridrow="24" pageId="7" pageNumber="316">75.619</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8DD6A807FA5B" box="[163,1334,1451,1475]" gridrow="25" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8DD6ADF8FA5B" box="[163,201,1451,1475]" gridcol="0" gridrow="25" pageId="7" pageNumber="316">25</th>
<td id="76D68840FFC0E7F9FE5C8DD6AF39FA5B" box="[420,520,1451,1475]" gridcol="1" gridrow="25" pageId="7" pageNumber="316">Kannur</td>
<td id="76D68840FFC0E7F9FD5D8DD6AE64FA5B" box="[677,853,1451,1475]" gridcol="2" gridrow="25" pageId="7" pageNumber="316">Ullikkal</td>
<td id="76D68840FFC0E7F9FC5E8DD6A912FA5B" box="[934,1059,1451,1475]" gridcol="3" gridrow="25" pageId="7" pageNumber="316">12.0375</td>
<td id="76D68840FFC0E7F9FB5F8DD6A807FA5B" box="[1191,1334,1451,1475]" gridcol="4" gridrow="25" pageId="7" pageNumber="316">75.6703</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8DAEA807FA74" box="[163,1334,1491,1516]" gridrow="26" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8DAEADF8FA74" box="[163,201,1491,1516]" gridcol="0" gridrow="26" pageId="7" pageNumber="316">26</th>
<td id="76D68840FFC0E7F9FE5C8DAEAF39FA74" box="[420,520,1491,1516]" gridcol="1" gridrow="26" pageId="7" pageNumber="316">Kannur</td>
<td id="76D68840FFC0E7F9FD5D8DAEAE64FA74" box="[677,853,1491,1516]" gridcol="2" gridrow="26" pageId="7" pageNumber="316">Kakkad</td>
<td id="76D68840FFC0E7F9FC5E8DAEA912FA74" box="[934,1059,1491,1516]" gridcol="3" gridrow="26" pageId="7" pageNumber="316">11.8894</td>
<td id="76D68840FFC0E7F9FB5F8DAEA807FA74" box="[1191,1334,1491,1516]" gridcol="4" gridrow="26" pageId="7" pageNumber="316">75.3879</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8D86A807F98C" box="[163,1334,1531,1556]" gridrow="27" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8D86ADF8F98C" box="[163,201,1531,1556]" gridcol="0" gridrow="27" pageId="7" pageNumber="316">27</th>
<td id="76D68840FFC0E7F9FE5C8D86AF39F98C" box="[420,520,1531,1556]" gridcol="1" gridrow="27" pageId="7" pageNumber="316">Kannur</td>
<td id="76D68840FFC0E7F9FD5D8D86AE64F98C" box="[677,853,1531,1556]" gridcol="2" gridrow="27" pageId="7" pageNumber="316">Iritty</td>
<td id="76D68840FFC0E7F9FC5E8D86A912F98C" box="[934,1059,1531,1556]" gridcol="3" gridrow="27" pageId="7" pageNumber="316">11.9819</td>
<td id="76D68840FFC0E7F9FB5F8D86A807F98C" box="[1191,1334,1531,1556]" gridcol="4" gridrow="27" pageId="7" pageNumber="316">75.6703</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8E59A807F9A4" box="[163,1334,1572,1596]" gridrow="28" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8E59ADF8F9A4" box="[163,201,1572,1596]" gridcol="0" gridrow="28" pageId="7" pageNumber="316">28</th>
<td id="76D68840FFC0E7F9FE5C8E59AF39F9A4" box="[420,520,1572,1596]" gridcol="1" gridrow="28" pageId="7" pageNumber="316">Idukki</td>
<td id="76D68840FFC0E7F9FD5D8E59AE64F9A4" box="[677,853,1572,1596]" gridcol="2" gridrow="28" pageId="7" pageNumber="316">Painavu</td>
<td id="76D68840FFC0E7F9FC5E8E59A912F9A4" box="[934,1059,1572,1596]" gridcol="3" gridrow="28" pageId="7" pageNumber="316">9.85392</td>
<td id="76D68840FFC0E7F9FB5F8E59A807F9A4" box="[1191,1334,1572,1596]" gridcol="4" gridrow="28" pageId="7" pageNumber="316">76.9481</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8E31A807F9FC" box="[163,1334,1612,1636]" gridrow="29" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8E31ADF8F9FC" box="[163,201,1612,1636]" gridcol="0" gridrow="29" pageId="7" pageNumber="316">29</th>
<td id="76D68840FFC0E7F9FE5C8E31AF39F9FC" box="[420,520,1612,1636]" gridcol="1" gridrow="29" pageId="7" pageNumber="316">Idukki</td>
<td id="76D68840FFC0E7F9FD5D8E31AE64F9FC" box="[677,853,1612,1636]" gridcol="2" gridrow="29" pageId="7" pageNumber="316">Painavu</td>
<td id="76D68840FFC0E7F9FC5E8E31A912F9FC" box="[934,1059,1612,1636]" gridcol="3" gridrow="29" pageId="7" pageNumber="316">9.86493</td>
<td id="76D68840FFC0E7F9FB5F8E31A807F9FC" box="[1191,1334,1612,1636]" gridcol="4" gridrow="29" pageId="7" pageNumber="316">76.9577</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8E09A807F915" box="[163,1334,1652,1677]" gridrow="30" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8E09ADF8F915" box="[163,201,1652,1677]" gridcol="0" gridrow="30" pageId="7" pageNumber="316">30</th>
<td id="76D68840FFC0E7F9FE5C8E09AF39F915" box="[420,520,1652,1677]" gridcol="1" gridrow="30" pageId="7" pageNumber="316">Idukki</td>
<td id="76D68840FFC0E7F9FD5D8E09AE64F915" box="[677,853,1652,1677]" gridcol="2" gridrow="30" pageId="7" pageNumber="316">Cheruthoni</td>
<td id="76D68840FFC0E7F9FC5E8E09A912F915" box="[934,1059,1652,1677]" gridcol="3" gridrow="30" pageId="7" pageNumber="316">9.85885</td>
<td id="76D68840FFC0E7F9FB5F8E09A807F915" box="[1191,1334,1652,1677]" gridcol="4" gridrow="30" pageId="7" pageNumber="316">76.9617</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8EE0A807F92D" box="[163,1334,1693,1717]" gridrow="31" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8EE0ADF8F92D" box="[163,201,1693,1717]" gridcol="0" gridrow="31" pageId="7" pageNumber="316">31</th>
<td id="76D68840FFC0E7F9FE5C8EE0AF39F92D" box="[420,520,1693,1717]" gridcol="1" gridrow="31" pageId="7" pageNumber="316">Idukki</td>
<td id="76D68840FFC0E7F9FD5D8EE0AE64F92D" box="[677,853,1693,1717]" gridcol="2" gridrow="31" pageId="7" pageNumber="316">Upputhode</td>
<td id="76D68840FFC0E7F9FC5E8EE0A912F92D" box="[934,1059,1693,1717]" gridcol="3" gridrow="31" pageId="7" pageNumber="316">9.88916</td>
<td id="76D68840FFC0E7F9FB5F8EE0A807F92D" box="[1191,1334,1693,1717]" gridcol="4" gridrow="31" pageId="7" pageNumber="316">77.0001</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8EB8A807F945" box="[163,1334,1733,1757]" gridrow="32" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8EB8ADF8F945" box="[163,201,1733,1757]" gridcol="0" gridrow="32" pageId="7" pageNumber="316">32</th>
<td id="76D68840FFC0E7F9FE5C8EB8AF39F945" box="[420,520,1733,1757]" gridcol="1" gridrow="32" pageId="7" pageNumber="316">Idukki</td>
<td id="76D68840FFC0E7F9FD5D8EB8AE64F945" box="[677,853,1733,1757]" gridcol="2" gridrow="32" pageId="7" pageNumber="316">Upputhode</td>
<td id="76D68840FFC0E7F9FC5E8EB8A912F945" box="[934,1059,1733,1757]" gridcol="3" gridrow="32" pageId="7" pageNumber="316">9.88379</td>
<td id="76D68840FFC0E7F9FB5F8EB8A807F945" box="[1191,1334,1733,1757]" gridcol="4" gridrow="32" pageId="7" pageNumber="316">77.0067</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8E90A807F89D" box="[163,1334,1773,1797]" gridrow="33" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8E90ADF8F89D" box="[163,201,1773,1797]" gridcol="0" gridrow="33" pageId="7" pageNumber="316">33</th>
<td id="76D68840FFC0E7F9FE5C8E90AF39F89D" box="[420,520,1773,1797]" gridcol="1" gridrow="33" pageId="7" pageNumber="316">Idukki</td>
<td id="76D68840FFC0E7F9FD5D8E90AE64F89D" box="[677,853,1773,1797]" gridcol="2" gridrow="33" pageId="7" pageNumber="316">Upputhode</td>
<td id="76D68840FFC0E7F9FC5E8E90A912F89D" box="[934,1059,1773,1797]" gridcol="3" gridrow="33" pageId="7" pageNumber="316">9.88332</td>
<td id="76D68840FFC0E7F9FB5F8E90A807F89D" box="[1191,1334,1773,1797]" gridcol="4" gridrow="33" pageId="7" pageNumber="316">77.0048</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8F68A807F8B5" box="[163,1334,1813,1837]" gridrow="34" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8F68ADF8F8B5" box="[163,201,1813,1837]" gridcol="0" gridrow="34" pageId="7" pageNumber="316">34</th>
<td id="76D68840FFC0E7F9FE5C8F68AF39F8B5" box="[420,520,1813,1837]" gridcol="1" gridrow="34" pageId="7" pageNumber="316">Idukki</td>
<td id="76D68840FFC0E7F9FD5D8F68AE64F8B5" box="[677,853,1813,1837]" gridcol="2" gridrow="34" pageId="7" pageNumber="316">Thadiyampad</td>
<td id="76D68840FFC0E7F9FC5E8F68A912F8B5" box="[934,1059,1813,1837]" gridcol="3" gridrow="34" pageId="7" pageNumber="316">9.87592</td>
<td id="76D68840FFC0E7F9FB5F8F68A807F8B5" box="[1191,1334,1813,1837]" gridcol="4" gridrow="34" pageId="7" pageNumber="316">76.9684</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8F40A807F8CE" box="[163,1334,1853,1878]" gridrow="35" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8F40ADF8F8CE" box="[163,201,1853,1878]" gridcol="0" gridrow="35" pageId="7" pageNumber="316">35</th>
<td id="76D68840FFC0E7F9FE5C8F40AF39F8CE" box="[420,520,1853,1878]" gridcol="1" gridrow="35" pageId="7" pageNumber="316">Kottayam</td>
<td id="76D68840FFC0E7F9FD5D8F40AE64F8CE" box="[677,853,1853,1878]" gridcol="2" gridrow="35" pageId="7" pageNumber="316">Velloor</td>
<td id="76D68840FFC0E7F9FC5E8F40A912F8CE" box="[934,1059,1853,1878]" gridcol="3" gridrow="35" pageId="7" pageNumber="316">9.82199</td>
<td id="76D68840FFC0E7F9FB5F8F40A807F8CE" box="[1191,1334,1853,1878]" gridcol="4" gridrow="35" pageId="7" pageNumber="316">76.4672</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8F1BA807F8E6" box="[163,1334,1894,1918]" gridrow="36" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8F1BADF8F8E6" box="[163,201,1894,1918]" gridcol="0" gridrow="36" pageId="7" pageNumber="316">36</th>
<td id="76D68840FFC0E7F9FE5C8F1BAF39F8E6" box="[420,520,1894,1918]" gridcol="1" gridrow="36" pageId="7" pageNumber="316">Kottayam</td>
<td id="76D68840FFC0E7F9FD5D8F1BAE64F8E6" box="[677,853,1894,1918]" gridcol="2" gridrow="36" pageId="7" pageNumber="316">Velloor</td>
<td id="76D68840FFC0E7F9FC5E8F1BA912F8E6" box="[934,1059,1894,1918]" gridcol="3" gridrow="36" pageId="7" pageNumber="316">9.8212</td>
<td id="76D68840FFC0E7F9FB5F8F1BA807F8E6" box="[1191,1334,1894,1918]" gridcol="4" gridrow="36" pageId="7" pageNumber="316">76.4672</td>
</tr>
<tr id="3507E13CFFC0E7F9FF5B8FF3A807F83F" box="[163,1334,1934,1959]" gridrow="37" pageId="7" pageNumber="316">
<th id="76D68840FFC0E7F9FF5B8FF3ADF8F83F" box="[163,201,1934,1959]" gridcol="0" gridrow="37" pageId="7" pageNumber="316">37</th>
<td id="76D68840FFC0E7F9FE5C8FF3AF39F83F" box="[420,520,1934,1959]" gridcol="1" gridrow="37" pageId="7" pageNumber="316">Kottayam</td>
<td id="76D68840FFC0E7F9FD5D8FF3AE64F83F" box="[677,853,1934,1959]" gridcol="2" gridrow="37" pageId="7" pageNumber="316">Velloor</td>
<td id="76D68840FFC0E7F9FC5E8FF3A912F83F" box="[934,1059,1934,1959]" gridcol="3" gridrow="37" pageId="7" pageNumber="316">9.82198</td>
<td id="76D68840FFC0E7F9FB5F8FF3A807F83F" box="[1191,1334,1934,1959]" gridcol="4" gridrow="37" pageId="7" pageNumber="316">76.4672</td>
</tr>
</table>
</paragraph>
<paragraph id="8B88E37EFFC01801FBA28FCAA8A0F857" blockId="7.[1114,1425,1975,1999]" box="[1114,1425,1975,1999]" pageId="7" pageNumber="316">
<tableNote id="76D1E2F0FFC01801FBA28FCAA8A0F857" box="[1114,1425,1975,1999]" pageId="7" pageNumber="316" targetBox="[163,1334,445,1959]" targetPageId="7">......continued on the next page</tableNote>
</paragraph>
<caption id="DF48B3F6FFCF180EFF5B88DFACA0FF22" box="[163,401,162,186]" isContinuationCaption="true" pageId="8" pageNumber="317" startId="8.[163,247,162,186]" targetType="table">
<paragraph id="8B88E37EFFCF180EFF5B88DFACA0FF22" blockId="8.[163,401,162,186]" box="[163,401,162,186]" pageId="8" pageNumber="317">
<emphasis id="B9433F6CFFCF180EFF5B88DFAC21FF22" bold="true" box="[163,272,162,186]" pageId="8" pageNumber="317">TABLE 3.</emphasis>
(Continued)
</paragraph>
</caption>
<paragraph id="8B88E37EFFCF180EFF5B88B6A9C8FE64" pageId="8" pageNumber="317">
<table id="F93711DEFFCFE7F9FF5B88B7A807FE65" box="[163,1334,202,509]" colsContinueFrom="7.[163,1334,445,1959]" gridcols="5" gridrows="8" pageId="8" pageNumber="317">
<tr id="3507E13CFFCFE7F9FF5B88B7A807FF7A" box="[163,1334,202,226]" gridrow="0" pageId="8" pageNumber="317">
<th id="76D68840FFCFE7F9FF5B88B7ADF8FF7A" box="[163,201,202,226]" gridcol="0" gridrow="0" pageId="8" pageNumber="317">No.</th>
<th id="76D68840FFCFE7F9FE5C88B7AF16FF7A" box="[420,551,202,226]" gridcol="1" gridrow="0" pageId="8" pageNumber="317">District</th>
<th id="76D68840FFCFE7F9FD5D88B7AE26FF7A" box="[677,791,202,226]" gridcol="2" gridrow="0" pageId="8" pageNumber="317">Location</th>
<th id="76D68840FFCFE7F9FC5E88B7A912FF7A" box="[934,1059,202,226]" gridcol="3" gridrow="0" pageId="8" pageNumber="317">Latitude (N)</th>
<th id="76D68840FFCFE7F9FB5F88B7A807FF7A" box="[1191,1334,202,226]" gridcol="4" gridrow="0" pageId="8" pageNumber="317">Longitude (E)</th>
</tr>
<tr id="3507E13CFFCFE7F9FF5B888EA807FE94" box="[163,1334,243,268]" gridrow="1" pageId="8" pageNumber="317">
<th id="76D68840FFCFE7F9FF5B888EADF8FE94" box="[163,201,243,268]" gridcol="0" gridrow="1" pageId="8" pageNumber="317">38</th>
<td id="76D68840FFCFE7F9FE5C888EAF16FE94" box="[420,551,243,268]" gridcol="1" gridrow="1" pageId="8" pageNumber="317">Kottayam</td>
<td id="76D68840FFCFE7F9FD5D888EAE26FE94" box="[677,791,243,268]" gridcol="2" gridrow="1" pageId="8" pageNumber="317">Kottayam</td>
<td id="76D68840FFCFE7F9FC5E888EA912FE94" box="[934,1059,243,268]" gridcol="3" gridrow="1" pageId="8" pageNumber="317">9.62248</td>
<td id="76D68840FFCFE7F9FB5F888EA807FE94" box="[1191,1334,243,268]" gridcol="4" gridrow="1" pageId="8" pageNumber="317">76.4762</td>
</tr>
<tr id="3507E13CFFCFE7F9FF5B8966A807FEAC" box="[163,1334,283,308]" gridrow="2" pageId="8" pageNumber="317">
<th id="76D68840FFCFE7F9FF5B8966ADF8FEAC" box="[163,201,283,308]" gridcol="0" gridrow="2" pageId="8" pageNumber="317">39</th>
<td id="76D68840FFCFE7F9FE5C8966AF16FEAC" box="[420,551,283,308]" gridcol="1" gridrow="2" pageId="8" pageNumber="317">Kottayam</td>
<td id="76D68840FFCFE7F9FD5D8966AE26FEAC" box="[677,791,283,308]" gridcol="2" gridrow="2" pageId="8" pageNumber="317">Kottayam</td>
<td id="76D68840FFCFE7F9FC5E8966A912FEAC" box="[934,1059,283,308]" gridcol="3" gridrow="2" pageId="8" pageNumber="317">9.59014</td>
<td id="76D68840FFCFE7F9FB5F8966A807FEAC" box="[1191,1334,283,308]" gridcol="4" gridrow="2" pageId="8" pageNumber="317">76.5218</td>
</tr>
<tr id="3507E13CFFCFE7F9FF5B893EA807FEC4" box="[163,1334,323,348]" gridrow="3" pageId="8" pageNumber="317">
<th id="76D68840FFCFE7F9FF5B893EADF8FEC4" box="[163,201,323,348]" gridcol="0" gridrow="3" pageId="8" pageNumber="317">40</th>
<td id="76D68840FFCFE7F9FE5C893EAF16FEC4" box="[420,551,323,348]" gridcol="1" gridrow="3" pageId="8" pageNumber="317">Kottayam</td>
<td id="76D68840FFCFE7F9FD5D893EAE26FEC4" box="[677,791,323,348]" gridcol="2" gridrow="3" pageId="8" pageNumber="317">Kottayam</td>
<td id="76D68840FFCFE7F9FC5E893EA912FEC4" box="[934,1059,323,348]" gridcol="3" gridrow="3" pageId="8" pageNumber="317">9.59016</td>
<td id="76D68840FFCFE7F9FB5F893EA807FEC4" box="[1191,1334,323,348]" gridcol="4" gridrow="3" pageId="8" pageNumber="317">76.5218</td>
</tr>
<tr id="3507E13CFFCFE7F9FF5B8911A807FE1C" box="[163,1334,364,388]" gridrow="4" pageId="8" pageNumber="317">
<th id="76D68840FFCFE7F9FF5B8911ADF8FE1C" box="[163,201,364,388]" gridcol="0" gridrow="4" pageId="8" pageNumber="317">41</th>
<td id="76D68840FFCFE7F9FE5C8911AF16FE1C" box="[420,551,364,388]" gridcol="1" gridrow="4" pageId="8" pageNumber="317">Kottayam</td>
<td id="76D68840FFCFE7F9FD5D8911AE26FE1C" box="[677,791,364,388]" gridcol="2" gridrow="4" pageId="8" pageNumber="317">Thiruvarpu</td>
<td id="76D68840FFCFE7F9FC5E8911A912FE1C" box="[934,1059,364,388]" gridcol="3" gridrow="4" pageId="8" pageNumber="317">9.5821</td>
<td id="76D68840FFCFE7F9FB5F8911A807FE1C" box="[1191,1334,364,388]" gridcol="4" gridrow="4" pageId="8" pageNumber="317">76.4873</td>
</tr>
<tr id="3507E13CFFCFE7F9FF5B89E9A807FE35" box="[163,1334,404,429]" gridrow="5" pageId="8" pageNumber="317">
<th id="76D68840FFCFE7F9FF5B89E9ADF8FE35" box="[163,201,404,429]" gridcol="0" gridrow="5" pageId="8" pageNumber="317">42</th>
<td id="76D68840FFCFE7F9FE5C89E9AF16FE35" box="[420,551,404,429]" gridcol="1" gridrow="5" pageId="8" pageNumber="317">Kottayam</td>
<td id="76D68840FFCFE7F9FD5D89E9AE26FE35" box="[677,791,404,429]" gridcol="2" gridrow="5" pageId="8" pageNumber="317">Kottayam</td>
<td id="76D68840FFCFE7F9FC5E89E9A912FE35" box="[934,1059,404,429]" gridcol="3" gridrow="5" pageId="8" pageNumber="317">9.58211</td>
<td id="76D68840FFCFE7F9FB5F89E9A807FE35" box="[1191,1334,404,429]" gridcol="4" gridrow="5" pageId="8" pageNumber="317">76.4873</td>
</tr>
<tr id="3507E13CFFCFE7F9FF5B89C1A807FE4D" box="[163,1334,444,469]" gridrow="6" pageId="8" pageNumber="317">
<th id="76D68840FFCFE7F9FF5B89C1ADF8FE4D" box="[163,201,444,469]" gridcol="0" gridrow="6" pageId="8" pageNumber="317">43</th>
<td id="76D68840FFCFE7F9FE5C89C1AF16FE4D" box="[420,551,444,469]" gridcol="1" gridrow="6" pageId="8" pageNumber="317">Kottayam</td>
<td id="76D68840FFCFE7F9FD5D89C1AE26FE4D" box="[677,791,444,469]" gridcol="2" gridrow="6" pageId="8" pageNumber="317">Kottayam</td>
<td id="76D68840FFCFE7F9FC5E89C1A912FE4D" box="[934,1059,444,469]" gridcol="3" gridrow="6" pageId="8" pageNumber="317">9.58816</td>
<td id="76D68840FFCFE7F9FB5F89C1A807FE4D" box="[1191,1334,444,469]" gridcol="4" gridrow="6" pageId="8" pageNumber="317">76.5423</td>
</tr>
<tr id="3507E13CFFCFE7F9FF5B8999A807FE65" box="[163,1334,484,509]" gridrow="7" pageId="8" pageNumber="317">
<th id="76D68840FFCFE7F9FF5B8999ADF8FE65" box="[163,201,484,509]" gridcol="0" gridrow="7" pageId="8" pageNumber="317">44</th>
<td id="76D68840FFCFE7F9FE5C8999AF16FE65" box="[420,551,484,509]" gridcol="1" gridrow="7" pageId="8" pageNumber="317">Ernamkulam</td>
<td id="76D68840FFCFE7F9FD5D8999AE26FE65" box="[677,791,484,509]" gridcol="2" gridrow="7" pageId="8" pageNumber="317">Kunnukara</td>
<td id="76D68840FFCFE7F9FC5E8999A912FE65" box="[934,1059,484,509]" gridcol="3" gridrow="7" pageId="8" pageNumber="317">10.1634</td>
<td id="76D68840FFCFE7F9FB5F8999A807FE65" box="[1191,1334,484,509]" gridcol="4" gridrow="7" pageId="8" pageNumber="317">76.3131</td>
</tr>
</table>
</paragraph>
<paragraph id="8B88E37EFFCF180EFF3F8A2FAE15FD65" blockId="8.[151,1437,594,1737]" pageId="8" pageNumber="317">
Elongated body, generally parallel-sided, gradually tapering towards the posterior end (
<figureCitation id="130CFFFBFFCF180EFB748A2FA9EEFDF5" box="[1164,1247,594,621]" captionStart="FIGURES 15‒17" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[210,1376,182,1910]" captionTargetId="figure-17@9.[203,1384,181,1919]" captionTargetPageId="9" captionText="FIGURES 15‒17. Ctenolepisma rothschildi. 15) Dorsal view. 16) Body scales. 17) SEM of labial sensory papillae." figureDoi="http://doi.org/10.5281/zenodo.14735044" httpUri="https://zenodo.org/record/14735044/files/figure.png" pageId="8" pageNumber="317">Fig. 15</figureCitation>
). Body length up to
<quantity id="4CCF4E9BFFCF180EFF4B8A0AAC36FD09" box="[179,263,631,657]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="8" pageNumber="317" unit="mm" value="9.8">9.8 mm</quantity>
(
<specimenCount id="9D3128F7FFCF180EFEEF8A0AAC18FD09" box="[279,297,631,657]" count="1" pageId="8" pageNumber="317" type="female">♀</specimenCount>
) and
<quantity id="4CCF4E9BFFCF180EFE908A0AAC8DFD09" box="[360,444,631,657]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="8" pageNumber="317" unit="mm" value="7.3">7.3 mm</quantity>
(
<specimenCount id="9D3128F7FFCF180EFE338A05ACD3FD0B" box="[459,482,632,659]" count="1" pageId="8" pageNumber="317" type="male">♂</specimenCount>
). Epidermal colour light straw; covered with dark brownish to blackish scales dorsally and whitish yellow scales ventrally. Length of antenna up to
<quantity id="4CCF4E9BFFCF180EFCC28AE7AEACFD2D" box="[826,925,666,693]" metricMagnitude="-3" metricUnit="m" metricValue="7.05" pageId="8" pageNumber="317" unit="mm" value="7.05">7.05 mm</quantity>
and that of cerci and paracercus
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and
<quantity id="4CCF4E9BFFCF180EFF6F8AC3ADDFFD41" box="[151,238,702,729]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="8" pageNumber="317" unit="mm" value="4.5">4.5 mm</quantity>
, respectively. Macrosetae plumose. Scales oval, globular or subtriangular, of varying sizes (
<figureCitation id="130CFFFBFFCF180EFB1E8AC3A806FD40" box="[1254,1335,702,728]" captionStart="FIGURES 15‒17" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[210,1376,182,1910]" captionTargetId="figure-17@9.[203,1384,181,1919]" captionTargetPageId="9" captionText="FIGURES 15‒17. Ctenolepisma rothschildi. 15) Dorsal view. 16) Body scales. 17) SEM of labial sensory papillae." figureDoi="http://doi.org/10.5281/zenodo.14735044" httpUri="https://zenodo.org/record/14735044/files/figure.png" pageId="8" pageNumber="317">Fig. 16</figureCitation>
), usually with 20–30 fine rays or 10-16 widely spaced, thicker rays.
</paragraph>
<paragraph id="8B88E37EFFCF180EFF3F8B7BAE89FC29" blockId="8.[151,1437,594,1737]" pageId="8" pageNumber="317">
Head wider (
<quantity id="4CCF4E9BFFCF180EFEA48B7AACD9FCB9" box="[348,488,775,801]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="8" pageNumber="317" unit="mm" value="1.0" valueMax="1.1" valueMin="0.9">0.9–1.1 mm</quantity>
) than long (
<quantity id="4CCF4E9BFFCF180EFD8D8B7BAE35FCB9" box="[629,772,774,801]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="8" pageNumber="317" unit="mm" value="0.6" valueMax="0.7" valueMin="0.5">0.5–0.7 mm</quantity>
); anterior margin of head with tufts of bifid and pectinate macrochaetae laterally extending to the eye and 1+1 frontal tufts of macrosetae on the frontal margin; clypeus and labrum also with 1+1 tufts of setae. Five-segmented maxillary palps; apical article 5–8.7 times longer than wide and about the same length as the penultimate article (0.9–1.2). Four-segmented labial palps with apical article wider than long (L/
<geoCoordinate id="EE0385B9FFCF180EFF098BEAAC72FC29" box="[241,323,918,945]" degrees="0.75" direction="west" orientation="longitude" pageId="8" pageNumber="317" precision="555" value="-0.75">
W
<quantity id="4CCF4E9BFFCF180EFEEB8BEBAC72FC29" box="[275,323,918,945]" metricMagnitude="-4" metricUnit="m" metricValue="8.049999999999999" metricValueMax="8.6" metricValueMin="7.5" pageId="8" pageNumber="317" unit="mm" value="0.8049999999999999" valueMax="0.86" valueMin="0.75">0.75</quantity>
</geoCoordinate>
–0.86 mm) and with 5 papillae in a single row (
<figureCitation id="130CFFFBFFCF180EFCA08BEBAE9AFC29" box="[856,939,918,945]" captionStart="FIGURES 15‒17" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[210,1376,182,1910]" captionTargetId="figure-17@9.[203,1384,181,1919]" captionTargetPageId="9" captionText="FIGURES 15‒17. Ctenolepisma rothschildi. 15) Dorsal view. 16) Body scales. 17) SEM of labial sensory papillae." figureDoi="http://doi.org/10.5281/zenodo.14735044" httpUri="https://zenodo.org/record/14735044/files/figure.png" pageId="8" pageNumber="317">Fig. 17</figureCitation>
).
</paragraph>
<paragraph id="8B88E37EFFCF180EFF3F8BC7AFE1FB49" blockId="8.[151,1437,594,1737]" pageId="8" pageNumber="317">
Thorax wider than abdomen. Maximum thorax width
<quantity id="4CCF4E9BFFCF180EFCE08BC6AE4FFC4D" box="[792,894,955,981]" metricMagnitude="-3" metricUnit="m" metricValue="2.18" pageId="8" pageNumber="317" unit="mm" value="2.18">2.18 mm</quantity>
; maximum thorax length
<quantity id="4CCF4E9BFFCF180EFB6F8BC6A9DEFC4D" box="[1175,1263,955,981]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="8" pageNumber="317" unit="mm" value="3.3">3.3 mm</quantity>
. Pronotal collar with 2–4 rows of bristle combs in the centre and single towards anterolateral margins. Pronotum (
<figureCitation id="130CFFFBFFCF180EFB328BA3A82BFC61" box="[1226,1306,990,1017]" captionStart="FIGURES 18‒20" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[333,1255,187,1916]" captionTargetId="figure-21@10.[327,1261,181,1920]" captionTargetPageId="10" captionText="FIGURES 18‒20. Ctenolepisma rothschildi, bristle combs and trichobothrial areas. 18) Pronotum. 19) Mesonotum. 20) Metanotum." figureDoi="http://doi.org/10.5281/zenodo.14735046" httpUri="https://zenodo.org/record/14735046/files/figure.png" pageId="8" pageNumber="317">Fig. 18</figureCitation>
) with (8–9) + (8–9) bristle combs on the lateral margins composed of 2–4 macrosetae. Mesonotum (
<figureCitation id="130CFFFBFFCF180EFB998C7FA980FB85" box="[1121,1201,1026,1053]" captionStart="FIGURES 18‒20" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[333,1255,187,1916]" captionTargetId="figure-21@10.[327,1261,181,1920]" captionTargetPageId="10" captionText="FIGURES 18‒20. Ctenolepisma rothschildi, bristle combs and trichobothrial areas. 18) Pronotum. 19) Mesonotum. 20) Metanotum." figureDoi="http://doi.org/10.5281/zenodo.14735046" httpUri="https://zenodo.org/record/14735046/files/figure.png" pageId="8" pageNumber="317">Fig. 19</figureCitation>
) with (10–11) + (11– 12) bristle combs composed of 2 to 4 macrosetae; metanotum (
<figureCitation id="130CFFFBFFCF180EFCA78C5BAE80FBD9" box="[863,945,1062,1089]" captionStart="FIGURES 18‒20" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[333,1255,187,1916]" captionTargetId="figure-21@10.[327,1261,181,1920]" captionTargetPageId="10" captionText="FIGURES 18‒20. Ctenolepisma rothschildi, bristle combs and trichobothrial areas. 18) Pronotum. 19) Mesonotum. 20) Metanotum." figureDoi="http://doi.org/10.5281/zenodo.14735046" httpUri="https://zenodo.org/record/14735046/files/figure.png" pageId="8" pageNumber="317">Fig. 20</figureCitation>
) with (8–10) + (9–11) lateral combs of 1–3 macrosetae. All nota with 1+1 posterior combs of 2–3 macrosetae. Trichobothrial areas in pronotum, posterior on inner side of the last (N) comb and the anterior one, on the inner side of N–3 bristle comb. In mesonotum posterior associated to the innerside of last (N) combs and posterior on the outer side of antepenultimate (N–2) comb and to the two last combs (N and N–1) in the metanotum.
</paragraph>
<paragraph id="8B88E37EFFCF180EFF3F8CA6A8A1FAF8" blockId="8.[151,1437,594,1737]" pageId="8" pageNumber="317">
Prosternum (L/
<geoCoordinate id="EE0385B9FFCF180EFE968CA6ACF1FB6D" box="[366,448,1243,1269]" degrees="1.24" direction="west" orientation="longitude" pageId="8" pageNumber="317" precision="555" value="-1.24">W 1.24</geoCoordinate>
) (
<figureCitation id="130CFFFBFFCF180EFE2E8CA7AF14FB6D" box="[470,549,1242,1269]" captionStart="FIGURES 21‒26" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[201,1382,184,1915]" captionTargetId="figure-17@11.[201,1387,181,1920]" captionTargetPageId="11" captionText="FIGURES 21‒26. Ctenolepisma rothschildi. 21) Prosternum. 22) Mesosternum. 23) Metasternum. 24) Urotergite X. 25) Male genital region, coxite IX, styli VIII and IX. 26). Female posterior region and ovipositor." figureDoi="http://doi.org/10.5281/zenodo.14735048" httpUri="https://zenodo.org/record/14735048/files/figure.png" pageId="8" pageNumber="317">Fig. 21</figureCitation>
) and mesosternum (L/
<geoCoordinate id="EE0385B9FFCF180EFCE38CA6AE5CFB6C" box="[795,877,1242,1269]" degrees="1.26" direction="west" orientation="longitude" pageId="8" pageNumber="317" precision="555" value="-1.26">W 1.26</geoCoordinate>
) (
<figureCitation id="130CFFFBFFCF180EFC7A8CA7AEE0FB6D" box="[898,977,1242,1269]" captionStart="FIGURES 21‒26" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[201,1382,184,1915]" captionTargetId="figure-17@11.[201,1387,181,1920]" captionTargetPageId="11" captionText="FIGURES 21‒26. Ctenolepisma rothschildi. 21) Prosternum. 22) Mesosternum. 23) Metasternum. 24) Urotergite X. 25) Male genital region, coxite IX, styli VIII and IX. 26). Female posterior region and ovipositor." figureDoi="http://doi.org/10.5281/zenodo.14735048" httpUri="https://zenodo.org/record/14735048/files/figure.png" pageId="8" pageNumber="317">Fig. 22</figureCitation>
) of approximately equal size and rounded subtriangular shape. Both with 1+1 posterior submarginal combs of 3–4 and 5–6 macrosetae, respectively, and with some marginal setae between comb and margin. Metasternum (
<figureCitation id="130CFFFBFFCF180EFC898D5FAEF4FAA5" box="[881,965,1314,1341]" captionStart="FIGURES 21‒26" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[201,1382,184,1915]" captionTargetId="figure-17@11.[201,1387,181,1920]" captionTargetPageId="11" captionText="FIGURES 21‒26. Ctenolepisma rothschildi. 21) Prosternum. 22) Mesosternum. 23) Metasternum. 24) Urotergite X. 25) Male genital region, coxite IX, styli VIII and IX. 26). Female posterior region and ovipositor." figureDoi="http://doi.org/10.5281/zenodo.14735048" httpUri="https://zenodo.org/record/14735048/files/figure.png" pageId="8" pageNumber="317">Fig. 23</figureCitation>
) broader (width
<quantity id="4CCF4E9BFFCF180EFB718D5EA9C1FAA5" box="[1161,1264,1315,1341]" metricMagnitude="-3" metricUnit="m" metricValue="1.07" pageId="8" pageNumber="317" unit="mm" value="1.07">1.07 mm</quantity>
) than previous segments, with 1+1 posterior submarginal combs of 6 macrosetae and numerous setae along the posterior margin.
</paragraph>
<paragraph id="8B88E37EFFCF180EFF3F8D17AC9AF9A0" blockId="8.[151,1437,594,1737]" pageId="8" pageNumber="317">Protibiae 2.65–2.74 times longer than wide; mesotibiae 3–3.6 times longer than wide and about 1–1.2 times longer than the protibiae; metatibiae 3.25–4.3 times longer than wide and about 1.3–1.6 times longer than the protibiae. Tarsal L/W: foretarsi 7.5, middle tarsi 7.5, hind tarsi 9.3–11. Basal article of tarsi longer than the remaining three together. Pretarsi with two strong, curved lateral claws and a smaller medial claw; coxae with broad, rounded scales along outer margin. Femur with small, narrow scales along proximal two-thirds of outer margin; remaining articles devoid of scales.</paragraph>
<paragraph id="8B88E37EFFCF180EFF3F8E3FA84AF951" blockId="8.[151,1437,594,1737]" pageId="8" pageNumber="317">
Urotergite I with 1+1 infralateral combs of 3–4 macrosetae, urotergites II–
<collectionCode id="ED267BBBFFCF180EFC078E3EA922F9C5" box="[1023,1043,1603,1629]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="317" type="Museum">V</collectionCode>
with 3+3 bristle combs, urotergites
<collectionCode id="ED267BBBFFCF180EFF6F8E1AAD85F919" box="[151,180,1639,1665]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="8" pageNumber="317">VI</collectionCode>
–VIII with 2+2 combs, urotergite IX without combs; urotergite X trapezoidal in shape (
<figureCitation id="130CFFFBFFCF180EFB7F8E1BA9E9F919" box="[1159,1240,1638,1665]" captionStart="FIGURES 21‒26" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[201,1382,184,1915]" captionTargetId="figure-17@11.[201,1387,181,1920]" captionTargetPageId="11" captionText="FIGURES 21‒26. Ctenolepisma rothschildi. 21) Prosternum. 22) Mesosternum. 23) Metasternum. 24) Urotergite X. 25) Male genital region, coxite IX, styli VIII and IX. 26). Female posterior region and ovipositor." figureDoi="http://doi.org/10.5281/zenodo.14735048" httpUri="https://zenodo.org/record/14735048/files/figure.png" pageId="8" pageNumber="317">Fig. 24</figureCitation>
) with 1+1 combs of 3 macrosetae and several marginal setae. Urosternites I and II glabrous, urosternites III–VIII with 1+1 lateral combs of 4–10 macrosetae each. Macrosetal numbers per comb on urotergites and urosternites given in
<tableCitation id="C6B5D6C5FFCF180EFAD98ED3A846F951" box="[1313,1399,1710,1737]" captionStart="TABLE 4" captionStartId="12.[151,239,151,175]" captionText="TABLE 4. Number of macrosetae per bristle comb on urotergites and urosternites in C. longicaudatum and C." pageId="8" pageNumber="317">Table 4</tableCitation>
.
</paragraph>
<caption id="DF48B3F6FFCE180FFF6F8FEAA81BF837" ID-DOI="http://doi.org/10.5281/zenodo.14735044" ID-Zenodo-Dep="14735044" box="[151,1322,1943,1968]" httpUri="https://zenodo.org/record/14735044/files/figure.png" pageId="9" pageNumber="318" startId="9.[151,264,1943,1967]" targetBox="[210,1376,182,1910]" targetPageId="9" targetType="figure">
<paragraph id="8B88E37EFFCE180FFF6F8FEAA81BF837" blockId="9.[151,1322,1943,1968]" box="[151,1322,1943,1968]" pageId="9" pageNumber="318">
<emphasis id="B9433F6CFFCE180FFF6F8FEAAC65F837" bold="true" box="[151,340,1943,1967]" pageId="9" pageNumber="318">FIGURES 15‒17.</emphasis>
<taxonomicName id="4C3798FDFFCE180FFEA28FEAAF68F837" authorityName=", Silvestri" authorityYear="1907" box="[346,601,1943,1967]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="9" pageNumber="318" phylum="Arthropoda" rank="species" species="rothschildi">
<emphasis id="B9433F6CFFCE180FFEA28FEAAF68F837" box="[346,601,1943,1967]" italics="true" pageId="9" pageNumber="318">Ctenolepisma rothschildi</emphasis>
</taxonomicName>
. 15) Dorsal view. 16) Body scales. 17) SEM of labial sensory papillae.
</paragraph>
</caption>
<caption id="DF48B3F6FFCD180CFF6F8FEAAC22F84B" ID-DOI="http://doi.org/10.5281/zenodo.14735046" ID-Zenodo-Dep="14735046" httpUri="https://zenodo.org/record/14735046/files/figure.png" pageId="10" pageNumber="319" startId="10.[151,264,1943,1967]" targetBox="[333,1255,187,1916]" targetPageId="10" targetType="figure">
<paragraph id="8B88E37EFFCD180CFF6F8FEAAC22F84B" blockId="10.[151,1436,1943,2003]" pageId="10" pageNumber="319">
<emphasis id="B9433F6CFFCD180CFF6F8FEAAC68F837" bold="true" box="[151,345,1943,1967]" pageId="10" pageNumber="319">FIGURES 18‒20.</emphasis>
<taxonomicName id="4C3798FDFFCD180CFE9D8FE5AF58F828" authorityName=", Silvestri" authorityYear="1907" box="[357,617,1944,1968]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="10" pageNumber="319" phylum="Arthropoda" rank="species" species="rothschildi">
<emphasis id="B9433F6CFFCD180CFE9D8FE5AF58F828" box="[357,617,1944,1968]" italics="true" pageId="10" pageNumber="319">Ctenolepisma rothschildi</emphasis>
</taxonomicName>
, bristle combs and trichobothrial areas. 18) Pronotum. 19) Mesonotum. 20) Metanotum.
</paragraph>
</caption>
<caption id="DF48B3F6FFCC180DFF6F8FEAA921F84B" ID-DOI="http://doi.org/10.5281/zenodo.14735048" ID-Zenodo-Dep="14735048" httpUri="https://zenodo.org/record/14735048/files/figure.png" pageId="11" pageNumber="320" startId="11.[151,264,1943,1967]" targetBox="[201,1382,184,1915]" targetPageId="11" targetType="figure">
<paragraph id="8B88E37EFFCC180DFF6F8FEAA921F84B" blockId="11.[151,1436,1943,2003]" pageId="11" pageNumber="320">
<emphasis id="B9433F6CFFCC180DFF6F8FEAAC65F837" bold="true" box="[151,340,1943,1967]" pageId="11" pageNumber="320">FIGURES 21‒26.</emphasis>
<emphasis id="B9433F6CFFCC180DFEA38FEAAF50F837" box="[347,609,1943,1967]" italics="true" pageId="11" pageNumber="320">
<taxonomicName id="4C3798FDFFCC180DFEA38FEAAF6DF837" authorityName=", Silvestri" authorityYear="1907" box="[347,604,1943,1967]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="11" pageNumber="320" phylum="Arthropoda" rank="species" species="rothschildi">Ctenolepisma rothschildi</taxonomicName>
.
</emphasis>
21) Prosternum. 22) Mesosternum. 23) Metasternum. 24) Urotergite X. 25) Male genital region, coxite IX, styli VIII and IX. 26). Female posterior region and ovipositor.
</paragraph>
</caption>
<caption id="DF48B3F6FFCB180AFF6F88EAA8ACFF37" box="[151,1437,151,175]" pageId="12" pageNumber="321" startId="12.[151,239,151,175]" targetType="table">
<paragraph id="8B88E37EFFCB180AFF6F88EAA8ACFF37" blockId="12.[151,1437,151,211]" box="[151,1437,151,175]" pageId="12" pageNumber="321">
<emphasis id="B9433F6CFFCB180AFF6F88EAAC20FF37" bold="true" box="[151,273,151,175]" pageId="12" pageNumber="321">TABLE 4.</emphasis>
Number of macrosetae per bristle comb on urotergites and urosternites in
<taxonomicName id="4C3798FDFFCB180AFB8488E5A873FF37" baseAuthorityName="Escherich" baseAuthorityYear="1905" box="[1148,1346,151,175]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="12" pageNumber="321" phylum="Arthropoda" rank="species" species="longicaudatum">
<emphasis id="B9433F6CFFCB180AFB8488E5A873FF37" box="[1148,1346,151,175]" italics="true" pageId="12" pageNumber="321">C. longicaudatum</emphasis>
</taxonomicName>
and C.
</paragraph>
</caption>
<paragraph id="8B88E37EFFCB180AFF6F88C6AC20FF4B" blockId="12.[151,1437,151,211]" box="[151,273,187,211]" pageId="12" pageNumber="321">
<emphasis id="B9433F6CFFCB180AFF6F88C6AC20FF4B" box="[151,273,187,211]" italics="true" pageId="12" pageNumber="321">
<taxonomicName id="4C3798FDFFCB180AFF6F88C6AC3CFF4B" authorityName=", Silvestri" authorityYear="1907" box="[151,269,187,211]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="12" pageNumber="321" phylum="Arthropoda" rank="species" species="rothschildi">rothschildi</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8B88E37EFFCB180AFF5B889EA86BFD2E" pageId="12" pageNumber="321">
<table id="F93711DEFFCBE7F9FF5B889EA8BEFD2E" box="[163,1423,227,694]" gridcols="9" gridrows="12" pageId="12" pageNumber="321">
<tr id="3507E13CFFCBE7F9FF5B889EA8BEFF63" box="[163,1423,227,251]" gridrow="0" pageId="12" pageNumber="321" rowspan-1="1" rowspan-4="1" rowspan-5="1" rowspan-8="1">
<th id="76D68840FFCBE7F9FF5B889EAC31FF63" box="[163,256,227,251]" gridcol="0" gridrow="0" pageId="12" pageNumber="321">
<emphasis id="B9433F6CFFCB180AFF5B889EAC31FF63" bold="true" box="[163,256,227,251]" pageId="12" pageNumber="321">Segment</emphasis>
</th>
<th id="76D68840FFCBE7F9FE38889EAFF4FF63" box="[448,709,227,251]" colspan="2" colspanRight="1" gridcol="2" gridrow="0" pageId="12" pageNumber="321">
<taxonomicName id="4C3798FDFFCB180AFE26889EAFA8FF63" baseAuthorityName="Escherich" baseAuthorityYear="1905" box="[478,665,227,251]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="12" pageNumber="321" phylum="Arthropoda" rank="species" species="longicaudatum">
<emphasis id="B9433F6CFFCB180AFE26889EAFA8FF63" bold="true" box="[478,665,227,251]" italics="true" pageId="12" pageNumber="321">C. longicaudatum</emphasis>
</taxonomicName>
</th>
<th id="76D68840FFCBE7F9FBFD889EA830FF63" box="[1029,1281,227,251]" colspan="2" colspanRight="1" gridcol="6" gridrow="0" pageId="12" pageNumber="321">
<taxonomicName id="4C3798FDFFCB180AFBC1889EA9F9FF63" authority="Urotergite" box="[1081,1224,227,251]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="12" pageNumber="321" phylum="Arthropoda" rank="species" species="rothschildi">
<emphasis id="B9433F6CFFCB180AFBC1889EA9F9FF63" bold="true" box="[1081,1224,227,251]" italics="true" pageId="12" pageNumber="321">C. rothschildi</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="3507E13CFFCBE7F9FF5B8971A8BEFEBC" box="[163,1423,268,292]" gridrow="1" pageId="12" pageNumber="321" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-5="1" rowspan-7="1">
<td id="76D68840FFCBE7F9FE388971AF19FEBC" box="[448,552,268,292]" gridcol="2" gridrow="1" pageId="12" pageNumber="321">Urotergite</td>
<td id="76D68840FFCBE7F9FD1A8971AE65FEBC" box="[738,852,268,292]" gridcol="4" gridrow="1" pageId="12" pageNumber="321">Urosternite</td>
<td id="76D68840FFCBE7F9FBFD8971A95CFEBC" box="[1029,1133,268,292]" gridcol="6" gridrow="1" pageId="12" pageNumber="321">Urotergite</td>
<td id="76D68840FFCBE7F9FAE58971A8BEFEBC" box="[1309,1423,268,292]" gridcol="8" gridrow="1" pageId="12" pageNumber="321">Urosternite</td>
</tr>
<tr id="3507E13CFFCBE7F9FF5B8949A8BEFED4" box="[163,1423,308,332]" gridrow="2" pageId="12" pageNumber="321" rowspan-0="1">
<td id="76D68840FFCBE7F9FED18949AC93FED4" box="[297,418,308,332]" gridcol="1" gridrow="2" pageId="12" pageNumber="321">Infra-lateral</td>
<td id="76D68840FFCBE7F9FE388949AF19FED4" box="[448,552,308,332]" gridcol="2" gridrow="2" pageId="12" pageNumber="321">Lateral</td>
<td id="76D68840FFCBE7F9FDAB8949AFF4FED4" box="[595,709,308,332]" gridcol="3" gridrow="2" pageId="12" pageNumber="321">Submedian</td>
<td id="76D68840FFCBE7F9FD1A8949AE65FED4" box="[738,852,308,332]" gridcol="4" gridrow="2" pageId="12" pageNumber="321">Lateral</td>
<td id="76D68840FFCBE7F9FC7E8949AEFFFED4" box="[902,974,308,332]" gridcol="5" gridrow="2" pageId="12" pageNumber="321">Lateral</td>
<td id="76D68840FFCBE7F9FBFD8949A95CFED4" box="[1029,1133,308,332]" gridcol="6" gridrow="2" pageId="12" pageNumber="321">Sublateral</td>
<td id="76D68840FFCBE7F9FB778949A830FED4" box="[1167,1281,308,332]" gridcol="7" gridrow="2" pageId="12" pageNumber="321">Submedian</td>
<td id="76D68840FFCBE7F9FAE58949A8BEFED4" box="[1309,1423,308,332]" gridcol="8" gridrow="2" pageId="12" pageNumber="321">Lateral</td>
</tr>
<tr id="3507E13CFFCBE7F9FF5B8921A8BEFEEC" box="[163,1423,348,372]" gridrow="3" pageId="12" pageNumber="321">
<th id="76D68840FFCBE7F9FF5B8921AC31FEEC" box="[163,256,348,372]" gridcol="0" gridrow="3" pageId="12" pageNumber="321">
<emphasis id="B9433F6CFFCB180AFF5B8921AD9CFEEC" bold="true" box="[163,173,348,372]" pageId="12" pageNumber="321">I</emphasis>
</th>
<td id="76D68840FFCBE7F9FED18921AC93FEEC" box="[297,418,348,372]" gridcol="1" gridrow="3" pageId="12" pageNumber="321">6–8</td>
<td id="76D68840FFCBE7F9FE388921AF19FEEC" box="[448,552,348,372]" gridcol="2" gridrow="3" pageId="12" pageNumber="321">-</td>
<td id="76D68840FFCBE7F9FDAB8921AFF4FEEC" box="[595,709,348,372]" gridcol="3" gridrow="3" pageId="12" pageNumber="321">-</td>
<td id="76D68840FFCBE7F9FD1A8921AE65FEEC" box="[738,852,348,372]" gridcol="4" gridrow="3" pageId="12" pageNumber="321">-</td>
<td id="76D68840FFCBE7F9FC7E8921AEFFFEEC" box="[902,974,348,372]" gridcol="5" gridrow="3" pageId="12" pageNumber="321">3–4</td>
<td id="76D68840FFCBE7F9FBFD8921A95CFEEC" box="[1029,1133,348,372]" gridcol="6" gridrow="3" pageId="12" pageNumber="321">-</td>
<td id="76D68840FFCBE7F9FB778921A830FEEC" box="[1167,1281,348,372]" gridcol="7" gridrow="3" pageId="12" pageNumber="321">-</td>
<td id="76D68840FFCBE7F9FAE58921A8BEFEEC" box="[1309,1423,348,372]" gridcol="8" gridrow="3" pageId="12" pageNumber="321">-</td>
</tr>
<tr id="3507E13CFFCBE7F9FF5B89F9A8BEFE04" box="[163,1423,388,412]" gridrow="4" pageId="12" pageNumber="321">
<th id="76D68840FFCBE7F9FF5B89F9AC31FE04" box="[163,256,388,412]" gridcol="0" gridrow="4" pageId="12" pageNumber="321">
<emphasis id="B9433F6CFFCB180AFF5B89F9AD86FE04" bold="true" box="[163,183,388,412]" pageId="12" pageNumber="321">II</emphasis>
</th>
<td id="76D68840FFCBE7F9FED189F9AC93FE04" box="[297,418,388,412]" gridcol="1" gridrow="4" pageId="12" pageNumber="321">6–8</td>
<td id="76D68840FFCBE7F9FE3889F9AF19FE04" box="[448,552,388,412]" gridcol="2" gridrow="4" pageId="12" pageNumber="321">5</td>
<td id="76D68840FFCBE7F9FDAB89F9AFF4FE04" box="[595,709,388,412]" gridcol="3" gridrow="4" pageId="12" pageNumber="321">6–7</td>
<td id="76D68840FFCBE7F9FD1A89F9AE65FE04" box="[738,852,388,412]" gridcol="4" gridrow="4" pageId="12" pageNumber="321">-</td>
<td id="76D68840FFCBE7F9FC7E89F9AEFFFE04" box="[902,974,388,412]" gridcol="5" gridrow="4" pageId="12" pageNumber="321">3–4</td>
<td id="76D68840FFCBE7F9FBFD89F9A95CFE04" box="[1029,1133,388,412]" gridcol="6" gridrow="4" pageId="12" pageNumber="321">2</td>
<td id="76D68840FFCBE7F9FB7789F9A830FE04" box="[1167,1281,388,412]" gridcol="7" gridrow="4" pageId="12" pageNumber="321">2</td>
<td id="76D68840FFCBE7F9FAE589F9A8BEFE04" box="[1309,1423,388,412]" gridcol="8" gridrow="4" pageId="12" pageNumber="321">-</td>
</tr>
<tr id="3507E13CFFCBE7F9FF5B89D1A8BEFE5D" box="[163,1423,428,453]" gridrow="5" pageId="12" pageNumber="321">
<th id="76D68840FFCBE7F9FF5B89D1AC31FE5D" box="[163,256,428,453]" gridcol="0" gridrow="5" pageId="12" pageNumber="321">
<emphasis id="B9433F6CFFCB180AFF5B89D1ADF0FE5C" bold="true" box="[163,193,428,452]" pageId="12" pageNumber="321">III</emphasis>
</th>
<td id="76D68840FFCBE7F9FED189D1AC93FE5D" box="[297,418,428,453]" gridcol="1" gridrow="5" pageId="12" pageNumber="321">8</td>
<td id="76D68840FFCBE7F9FE3889D1AF19FE5D" box="[448,552,428,453]" gridcol="2" gridrow="5" pageId="12" pageNumber="321">5</td>
<td id="76D68840FFCBE7F9FDAB89D1AFF4FE5D" box="[595,709,428,453]" gridcol="3" gridrow="5" pageId="12" pageNumber="321">6</td>
<td id="76D68840FFCBE7F9FD1A89D1AE65FE5D" box="[738,852,428,453]" gridcol="4" gridrow="5" pageId="12" pageNumber="321">17</td>
<td id="76D68840FFCBE7F9FC7E89D1AEFFFE5D" box="[902,974,428,453]" gridcol="5" gridrow="5" pageId="12" pageNumber="321">3–4</td>
<td id="76D68840FFCBE7F9FBFD89D1A95CFE5D" box="[1029,1133,428,453]" gridcol="6" gridrow="5" pageId="12" pageNumber="321">3</td>
<td id="76D68840FFCBE7F9FB7789D1A830FE5D" box="[1167,1281,428,453]" gridcol="7" gridrow="5" pageId="12" pageNumber="321">2</td>
<td id="76D68840FFCBE7F9FAE589D1A8BEFE5D" box="[1309,1423,428,453]" gridcol="8" gridrow="5" pageId="12" pageNumber="321">6</td>
</tr>
<tr id="3507E13CFFCBE7F9FF5B89A8A8BEFE75" box="[163,1423,469,493]" gridrow="6" pageId="12" pageNumber="321">
<th id="76D68840FFCBE7F9FF5B89A8AC31FE75" box="[163,256,469,493]" gridcol="0" gridrow="6" pageId="12" pageNumber="321">
<emphasis id="B9433F6CFFCB180AFF5B89A8AD8EFE75" bold="true" box="[163,191,469,493]" pageId="12" pageNumber="321">IV</emphasis>
</th>
<td id="76D68840FFCBE7F9FED189A8AC93FE75" box="[297,418,469,493]" gridcol="1" gridrow="6" pageId="12" pageNumber="321">8</td>
<td id="76D68840FFCBE7F9FE3889A8AF19FE75" box="[448,552,469,493]" gridcol="2" gridrow="6" pageId="12" pageNumber="321">5</td>
<td id="76D68840FFCBE7F9FDAB89A8AFF4FE75" box="[595,709,469,493]" gridcol="3" gridrow="6" pageId="12" pageNumber="321">7</td>
<td id="76D68840FFCBE7F9FD1A89A8AE65FE75" box="[738,852,469,493]" gridcol="4" gridrow="6" pageId="12" pageNumber="321">20</td>
<td id="76D68840FFCBE7F9FC7E89A8AEFFFE75" box="[902,974,469,493]" gridcol="5" gridrow="6" pageId="12" pageNumber="321">4–6</td>
<td id="76D68840FFCBE7F9FBFD89A8A95CFE75" box="[1029,1133,469,493]" gridcol="6" gridrow="6" pageId="12" pageNumber="321">3</td>
<td id="76D68840FFCBE7F9FB7789A8A830FE75" box="[1167,1281,469,493]" gridcol="7" gridrow="6" pageId="12" pageNumber="321">2</td>
<td id="76D68840FFCBE7F9FAE589A8A8BEFE75" box="[1309,1423,469,493]" gridcol="8" gridrow="6" pageId="12" pageNumber="321">8</td>
</tr>
<tr id="3507E13CFFCBE7F9FF5B8980A8BEFD8D" box="[163,1423,509,533]" gridrow="7" pageId="12" pageNumber="321">
<th id="76D68840FFCBE7F9FF5B8980AC31FD8D" box="[163,256,509,533]" gridcol="0" gridrow="7" pageId="12" pageNumber="321">
<emphasis id="B9433F6CFFCB180AFF5B8980AD84FD8D" bold="true" box="[163,181,509,533]" pageId="12" pageNumber="321">V</emphasis>
</th>
<td id="76D68840FFCBE7F9FED18980AC93FD8D" box="[297,418,509,533]" gridcol="1" gridrow="7" pageId="12" pageNumber="321">9–10</td>
<td id="76D68840FFCBE7F9FE388980AF19FD8D" box="[448,552,509,533]" gridcol="2" gridrow="7" pageId="12" pageNumber="321">5–7</td>
<td id="76D68840FFCBE7F9FDAB8980AFF4FD8D" box="[595,709,509,533]" gridcol="3" gridrow="7" pageId="12" pageNumber="321">6</td>
<td id="76D68840FFCBE7F9FD1A8980AE65FD8D" box="[738,852,509,533]" gridcol="4" gridrow="7" pageId="12" pageNumber="321">19-20</td>
<td id="76D68840FFCBE7F9FC7E8980AEFFFD8D" box="[902,974,509,533]" gridcol="5" gridrow="7" pageId="12" pageNumber="321">6</td>
<td id="76D68840FFCBE7F9FBFD8980A95CFD8D" box="[1029,1133,509,533]" gridcol="6" gridrow="7" pageId="12" pageNumber="321">3</td>
<td id="76D68840FFCBE7F9FB778980A830FD8D" box="[1167,1281,509,533]" gridcol="7" gridrow="7" pageId="12" pageNumber="321">2–3</td>
<td id="76D68840FFCBE7F9FAE58980A8BEFD8D" box="[1309,1423,509,533]" gridcol="8" gridrow="7" pageId="12" pageNumber="321">9–10</td>
</tr>
<tr id="3507E13CFFCBE7F9FF5B8A58A8BEFDA5" box="[163,1423,549,573]" gridrow="8" pageId="12" pageNumber="321">
<th id="76D68840FFCBE7F9FF5B8A58AC31FDA5" box="[163,256,549,573]" gridcol="0" gridrow="8" pageId="12" pageNumber="321">
<emphasis id="B9433F6CFFCB180AFF5B8A58AD8EFDA5" bold="true" box="[163,191,549,573]" pageId="12" pageNumber="321">VI</emphasis>
</th>
<td id="76D68840FFCBE7F9FED18A58AC93FDA5" box="[297,418,549,573]" gridcol="1" gridrow="8" pageId="12" pageNumber="321">9–10</td>
<td id="76D68840FFCBE7F9FE388A58AF19FDA5" box="[448,552,549,573]" gridcol="2" gridrow="8" pageId="12" pageNumber="321">5–6</td>
<td id="76D68840FFCBE7F9FDAB8A58AFF4FDA5" box="[595,709,549,573]" gridcol="3" gridrow="8" pageId="12" pageNumber="321">6</td>
<td id="76D68840FFCBE7F9FD1A8A58AE65FDA5" box="[738,852,549,573]" gridcol="4" gridrow="8" pageId="12" pageNumber="321">18-20</td>
<td id="76D68840FFCBE7F9FC7E8A58AEFFFDA5" box="[902,974,549,573]" gridcol="5" gridrow="8" pageId="12" pageNumber="321">5–6</td>
<td id="76D68840FFCBE7F9FBFD8A58A95CFDA5" box="[1029,1133,549,573]" gridcol="6" gridrow="8" pageId="12" pageNumber="321">-</td>
<td id="76D68840FFCBE7F9FB778A58A830FDA5" box="[1167,1281,549,573]" gridcol="7" gridrow="8" pageId="12" pageNumber="321">2–3</td>
<td id="76D68840FFCBE7F9FAE58A58A8BEFDA5" box="[1309,1423,549,573]" gridcol="8" gridrow="8" pageId="12" pageNumber="321">9</td>
</tr>
<tr id="3507E13CFFCBE7F9FF5B8A30A8BEFDFE" box="[163,1423,589,614]" gridrow="9" pageId="12" pageNumber="321">
<th id="76D68840FFCBE7F9FF5B8A30AC31FDFE" box="[163,256,589,614]" gridcol="0" gridrow="9" pageId="12" pageNumber="321">
<emphasis id="B9433F6CFFCB180AFF5B8A30ADF8FDFD" bold="true" box="[163,201,589,613]" pageId="12" pageNumber="321">VII</emphasis>
</th>
<td id="76D68840FFCBE7F9FED18A30AC93FDFE" box="[297,418,589,614]" gridcol="1" gridrow="9" pageId="12" pageNumber="321">-</td>
<td id="76D68840FFCBE7F9FE388A30AF19FDFE" box="[448,552,589,614]" gridcol="2" gridrow="9" pageId="12" pageNumber="321">11</td>
<td id="76D68840FFCBE7F9FDAB8A30AFF4FDFE" box="[595,709,589,614]" gridcol="3" gridrow="9" pageId="12" pageNumber="321">7-8</td>
<td id="76D68840FFCBE7F9FD1A8A30AE65FDFE" box="[738,852,589,614]" gridcol="4" gridrow="9" pageId="12" pageNumber="321">20-21</td>
<td id="76D68840FFCBE7F9FC7E8A30AEFFFDFE" box="[902,974,589,614]" gridcol="5" gridrow="9" pageId="12" pageNumber="321">4–5</td>
<td id="76D68840FFCBE7F9FBFD8A30A95CFDFE" box="[1029,1133,589,614]" gridcol="6" gridrow="9" pageId="12" pageNumber="321">-</td>
<td id="76D68840FFCBE7F9FB778A30A830FDFE" box="[1167,1281,589,614]" gridcol="7" gridrow="9" pageId="12" pageNumber="321">3</td>
<td id="76D68840FFCBE7F9FAE58A30A8BEFDFE" box="[1309,1423,589,614]" gridcol="8" gridrow="9" pageId="12" pageNumber="321">9</td>
</tr>
<tr id="3507E13CFFCBE7F9FF5B8A08A8BEFD16" box="[163,1423,629,654]" gridrow="10" pageId="12" pageNumber="321">
<th id="76D68840FFCBE7F9FF5B8A08AC31FD16" box="[163,256,629,654]" gridcol="0" gridrow="10" pageId="12" pageNumber="321">
<emphasis id="B9433F6CFFCB180AFF5B8A08ADE2FD15" bold="true" box="[163,211,629,653]" pageId="12" pageNumber="321">VIII</emphasis>
</th>
<td id="76D68840FFCBE7F9FED18A08AC93FD16" box="[297,418,629,654]" gridcol="1" gridrow="10" pageId="12" pageNumber="321">-</td>
<td id="76D68840FFCBE7F9FE388A08AF19FD16" box="[448,552,629,654]" gridcol="2" gridrow="10" pageId="12" pageNumber="321">11</td>
<td id="76D68840FFCBE7F9FDAB8A08AFF4FD16" box="[595,709,629,654]" gridcol="3" gridrow="10" pageId="12" pageNumber="321">7</td>
<td id="76D68840FFCBE7F9FD1A8A08AE65FD16" box="[738,852,629,654]" gridcol="4" gridrow="10" pageId="12" pageNumber="321">12-13</td>
<td id="76D68840FFCBE7F9FC7E8A08AEFFFD16" box="[902,974,629,654]" gridcol="5" gridrow="10" pageId="12" pageNumber="321">4–5</td>
<td id="76D68840FFCBE7F9FBFD8A08A95CFD16" box="[1029,1133,629,654]" gridcol="6" gridrow="10" pageId="12" pageNumber="321">-</td>
<td id="76D68840FFCBE7F9FB778A08A830FD16" box="[1167,1281,629,654]" gridcol="7" gridrow="10" pageId="12" pageNumber="321">2</td>
<td id="76D68840FFCBE7F9FAE58A08A8BEFD16" box="[1309,1423,629,654]" gridcol="8" gridrow="10" pageId="12" pageNumber="321">4–6</td>
</tr>
<tr id="3507E13CFFCBE7F9FF5B8AE3A8BEFD2E" box="[163,1423,670,694]" gridrow="11" pageId="12" pageNumber="321">
<th id="76D68840FFCBE7F9FF5B8AE3AC31FD2E" box="[163,256,670,694]" gridcol="0" gridrow="11" pageId="12" pageNumber="321">
<emphasis id="B9433F6CFFCB180AFF5B8AE3AD8EFD2E" bold="true" box="[163,191,670,694]" pageId="12" pageNumber="321">IX</emphasis>
</th>
<td id="76D68840FFCBE7F9FED18AE3AC93FD2E" box="[297,418,670,694]" gridcol="1" gridrow="11" pageId="12" pageNumber="321">-</td>
<td id="76D68840FFCBE7F9FE388AE3AF19FD2E" box="[448,552,670,694]" gridcol="2" gridrow="11" pageId="12" pageNumber="321">-</td>
<td id="76D68840FFCBE7F9FDAB8AE3AFF4FD2E" box="[595,709,670,694]" gridcol="3" gridrow="11" pageId="12" pageNumber="321">-</td>
<td id="76D68840FFCBE7F9FD1A8AE3AE65FD2E" box="[738,852,670,694]" gridcol="4" gridrow="11" pageId="12" pageNumber="321">-</td>
<td id="76D68840FFCBE7F9FC7E8AE3AEFFFD2E" box="[902,974,670,694]" gridcol="5" gridrow="11" pageId="12" pageNumber="321">-</td>
<td id="76D68840FFCBE7F9FBFD8AE3A95CFD2E" box="[1029,1133,670,694]" gridcol="6" gridrow="11" pageId="12" pageNumber="321">-</td>
<td id="76D68840FFCBE7F9FB778AE3A830FD2E" box="[1167,1281,670,694]" gridcol="7" gridrow="11" pageId="12" pageNumber="321">-</td>
<td id="76D68840FFCBE7F9FAE58AE3A8BEFD2E" box="[1309,1423,670,694]" gridcol="8" gridrow="11" pageId="12" pageNumber="321">-</td>
</tr>
</table>
</paragraph>
<paragraph id="8B88E37EFFCB180AFF348B70AE13FBDE" blockId="12.[151,1437,780,1095]" pageId="12" pageNumber="321">
Two pairs of styli in both sexes inserted on abdominal segments VIII and IX, armed apically with several strong spines. Stylets of IX more developed than that of VIII. The ratio length of styli IX/ length of styli VIII in female is about 1–1.67. The ratio length of styli IX/ length of styli VIII in male is about 1.3–1.6. Coxite IX in male with fringe of setae along internal margin; internal process about 2.6–4 times longer than the external process and 1–1.3 (L/W) times as long as broad at its base. Penis with numerous apical setae. Parameres absent (
<figureCitation id="130CFFFBFFCB180AFB748BE1A9D2FC2F" box="[1164,1251,924,951]" captionStart="FIGURES 21‒26" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[201,1382,184,1915]" captionTargetId="figure-17@11.[201,1387,181,1920]" captionTargetPageId="11" captionText="FIGURES 21‒26. Ctenolepisma rothschildi. 21) Prosternum. 22) Mesosternum. 23) Metasternum. 24) Urotergite X. 25) Male genital region, coxite IX, styli VIII and IX. 26). Female posterior region and ovipositor." figureDoi="http://doi.org/10.5281/zenodo.14735048" httpUri="https://zenodo.org/record/14735048/files/figure.png" pageId="12" pageNumber="321">Fig. 25</figureCitation>
). Genital region of female as in
<figureCitation id="130CFFFBFFCB180AFEB48BBDAC91FC42" box="[332,416,960,986]" captionStart="FIGURES 21‒26" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[201,1382,184,1915]" captionTargetId="figure-17@11.[201,1387,181,1920]" captionTargetPageId="11" captionText="FIGURES 21‒26. Ctenolepisma rothschildi. 21) Prosternum. 22) Mesosternum. 23) Metasternum. 24) Urotergite X. 25) Male genital region, coxite IX, styli VIII and IX. 26). Female posterior region and ovipositor." figureDoi="http://doi.org/10.5281/zenodo.14735048" httpUri="https://zenodo.org/record/14735048/files/figure.png" pageId="12" pageNumber="321">Fig. 26</figureCitation>
, coxite IX also with fringe of setae along internal margin, internal process about 0.5–1.5 times longer than wide at its base and 2.2–2.8 times longer than the external process. Ovipositor long with 34–41 divisions and its length measures
<quantity id="4CCF4E9BFFCB180AFDF28C75AF9AFBBB" box="[522,683,1032,1059]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.66" metricValueMin="2.24" pageId="12" pageNumber="321" unit="mm" value="2.45" valueMax="2.66" valueMin="2.24">2.24–2.66 mm</quantity>
from the base of urosternite VIII; surpassing the apex of the internal processes of coxites IX by 4.4–4.8 times coxite IX length.
</paragraph>
</subSubSection>
</treatment>
</document>

306
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<mods:title id="DEB68176852ABC6FAFF8479CBA8F92A0">Description of two synanthropic silverfish (Insecta: Zygentoma), Ctenolepisma longicaudatum (Escherich, 1905) and Ctenolepisma rothschildi (Silvestri, 1907), from Kerala, India</mods:title>
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<mods:namePart id="A8D9476A800472CD669B1F117C77AD95">Raphel, Sheeba</mods:namePart>
<mods:affiliation id="7A987C32E946C624E9499F01B5072E9E">Department of Zoology, St. Thomas College (Autonomous), Thrissur affiliated to the University of Calicut, Kerala, India</mods:affiliation>
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<mods:namePart id="1A2884953CBF1778EE924E97B715D7BC">Jose, Joyce</mods:namePart>
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<treatment id="039E5268FFC51803FF6F8ECDAE5CF8DF" ID-DOI="http://doi.org/10.5281/zenodo.14735030" ID-Zenodo-Dep="14735030" LSID="urn:lsid:plazi:treatment:039E5268FFC51803FF6F8ECDAE5CF8DF" httpUri="http://treatment.plazi.org/id/039E5268FFC51803FF6F8ECDAE5CF8DF" lastPageId="5" lastPageNumber="314" pageId="2" pageNumber="311">
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<paragraph id="8B88E37EFFC51804FF6F8ECDAFE3F952" blockId="2.[151,722,1712,1775]" box="[151,722,1712,1739]" pageId="2" pageNumber="311">
<heading id="D0C05412FFC51804FF6F8ECDAFE3F952" bold="true" box="[151,722,1712,1739]" fontSize="11" level="1" pageId="2" pageNumber="311" reason="1">
<taxonomicName id="4C3798FDFFC51804FF6F8ECDAFE3F952" authority="(Escherich, 1905)" baseAuthorityName="Escherich" baseAuthorityYear="1905" box="[151,722,1712,1739]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="2" pageNumber="311" phylum="Arthropoda" rank="species" species="longicaudatum">
<emphasis id="B9433F6CFFC51804FF6F8ECDAFE3F952" bold="true" box="[151,722,1712,1739]" pageId="2" pageNumber="311">
<emphasis id="B9433F6CFFC51804FF6F8ECDACCBF953" bold="true" box="[151,506,1712,1739]" italics="true" pageId="2" pageNumber="311">Ctenolepisma longicaudatum</emphasis>
(
<bibRefCitation id="EFA69E8FFFC51804FDF18ECDAFFAF952" author="Escherich, K." box="[521,715,1712,1738]" pageId="2" pageNumber="311" refId="ref5515" refString="Escherich, K. (1905) Das System der Lepismatiden. Zoologica. No. 43. Universitat Strassburg, Stuttgart, 164 pp." type="book" year="1905">Escherich,1905</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C32DB0F5FFC51804FF6F8EA8AC11F977" box="[151,288,1749,1775]" pageId="2" pageNumber="311" type="description">
<paragraph id="8B88E37EFFC51804FF6F8EA8AC11F977" blockId="2.[151,722,1712,1775]" box="[151,288,1749,1775]" pageId="2" pageNumber="311">
(
<figureCitation id="130CFFFBFFC51804FF678EA8AC29F977" box="[159,280,1749,1775]" captionStart-0="FIGURES 2‒5" captionStart-1="FIGURES 6‒8" captionStart-2="FIGURES 9‒14" captionStartId-0="4.[151,264,1943,1967]" captionStartId-1="5.[151,264,1334,1358]" captionStartId-2="6.[151,264,1943,1967]" captionTargetBox-0="[210,1378,188,1916]" captionTargetBox-1="[157,1429,186,1304]" captionTargetBox-2="[222,1365,182,1919]" captionTargetId-0="figure-21@4.[203,1384,181,1919]" captionTargetId-1="figure-17@5.[151,1436,181,1310]" captionTargetId-2="figure-21@6.[217,1371,181,1919]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURES 2‒5. Ctenolepisma longicaudatum. 2) Dorsal view. 3) Body scales. 4) Labial palps with five sensory papillae on apical article. 5) SEM of labial palp with more than 10 sensory papillae.Arrow: labial palp." captionText-1="FIGURES 6‒8. Ctenolepisma longicaudatum, bristle combs and trichobothrial areas. 6) Pronotum. 7) Mesonotum. 8) Metanotum." captionText-2="FIGURES 9‒14. Ctenolepisma longicaudatum. 9) Prosternum. 10) Mesosternum. 11) Metasternum. 12) Urotergite X. 13) Male genital area with coxite IX. 14) Female genital area with ovipositor and coxite IX." figureDoi-0="http://doi.org/10.5281/zenodo.14735036" figureDoi-1="http://doi.org/10.5281/zenodo.14735038" figureDoi-2="http://doi.org/10.5281/zenodo.14735040" httpUri-0="https://zenodo.org/record/14735036/files/figure.png" httpUri-1="https://zenodo.org/record/14735038/files/figure.png" httpUri-2="https://zenodo.org/record/14735040/files/figure.png" pageId="2" pageNumber="311">Figs. 2‒14</figureCitation>
)
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<subSubSection id="C32DB0F5FFC51803FF6F8F61AE5CF8DF" lastPageId="5" lastPageNumber="314" pageId="2" pageNumber="311" type="materials_examined">
<paragraph id="8B88E37EFFC51804FF6F8F61A90AF818" blockId="2.[151,1437,1820,1920]" pageId="2" pageNumber="311">
<emphasis id="B9433F6CFFC51804FF6F8F61ACA0F8AE" bold="true" box="[151,401,1820,1846]" pageId="2" pageNumber="311">Materials Examined.</emphasis>
<collectingCountry id="F320A3EEFFC51804FE608F60ACE5F8AF" box="[408,468,1821,1847]" name="India" pageId="2" pageNumber="311">India</collectingCountry>
,
<collectingRegion id="49F32D9CFFC51804FE278F60AF1DF8AF" box="[479,556,1821,1847]" country="India" name="Kerala" pageId="2" pageNumber="311">Kerala</collectingRegion>
, specimens from 13 sites (
<tableCitation id="C6B5D6C5FFC51804FCAA8F60AE96F8AF" box="[850,935,1821,1847]" captionStart="TABLE 2" captionStartId="3.[151,239,152,176]" captionText="TABLE 2. List of collection sites of Ctenolepisma longicaudatum with geocoordinates." pageId="2" pageNumber="311">Table 2</tableCitation>
) among old wooden articles, S. Raphel, coll. Specimens in alcohol deposited at Zoological Survey of
<collectingCountry id="F320A3EEFFC51804FCDE8F3CAE53F8C3" box="[806,866,1857,1883]" name="India" pageId="2" pageNumber="311">India</collectingCountry>
, Western Ghat Regional Centre,
<collectingCountry id="F320A3EEFFC51804FB278F3CA86CF8C3" box="[1247,1373,1857,1883]" name="India" pageId="2" pageNumber="311">Kozhikode</collectingCountry>
, with registration numbers ZSI/
<collectionCode id="ED267BBBFFC51804FE418F18AF20F8E7" box="[441,529,1893,1919]" pageId="2" pageNumber="311">WGRC</collectionCode>
/I.
<collectionCode id="ED267BBBFFC51804FDDD8F18AF09F8E7" box="[549,568,1893,1919]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="311">R</collectionCode>
.-INV.26924 and ZSI/
<collectionCode id="ED267BBBFFC51804FCD48F18AEB4F818" box="[812,901,1893,1920]" pageId="2" pageNumber="311">WGRC</collectionCode>
/I.
<collectionCode id="ED267BBBFFC51804FC628F18AE9CF818" box="[922,941,1893,1920]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="311">R</collectionCode>
.-INV.26925.
</paragraph>
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<paragraph id="8B88E37EFFC41805FF6F88E5A971FF28" blockId="3.[151,1088,152,176]" box="[151,1088,152,176]" pageId="3" pageNumber="312">
<emphasis id="B9433F6CFFC41805FF6F88E5AC3BFF28" bold="true" box="[151,266,152,176]" pageId="3" pageNumber="312">TABLE 2.</emphasis>
List of collection sites of
<taxonomicName id="4C3798FDFFC41805FDD988E5AE6BFF28" baseAuthorityName="Escherich" baseAuthorityYear="1905" box="[545,858,152,176]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="3" pageNumber="312" phylum="Arthropoda" rank="species" species="longicaudatum">
<emphasis id="B9433F6CFFC41805FDD988E5AE6BFF28" box="[545,858,152,176]" italics="true" pageId="3" pageNumber="312">Ctenolepisma longicaudatum</emphasis>
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with geocoordinates.
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<th id="76D68840FFC4E7F9FF5B88BCADF8FF41" box="[163,201,193,217]" gridcol="0" gridrow="0" pageId="3" pageNumber="312">No.</th>
<th id="76D68840FFC4E7F9FEA188BCACC0FF41" box="[345,497,193,217]" gridcol="1" gridrow="0" pageId="3" pageNumber="312">District</th>
<th id="76D68840FFC4E7F9FD9188BCAE3DFF41" box="[617,780,193,217]" gridcol="2" gridrow="0" pageId="3" pageNumber="312">Location</th>
<th id="76D68840FFC4E7F9FC6288BCA926FF41" box="[922,1047,193,217]" gridcol="3" gridrow="0" pageId="3" pageNumber="312">Latitude (N)</th>
<th id="76D68840FFC4E7F9FB4688BCA87CFF41" box="[1214,1357,193,217]" gridcol="4" gridrow="0" pageId="3" pageNumber="312">Longitude (E)</th>
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<tr id="3507E13CFFC4E7F9FF5B8894A87CFE99" box="[163,1357,233,257]" gridrow="1" pageId="3" pageNumber="312">
<th id="76D68840FFC4E7F9FF5B8894ADF8FE99" box="[163,201,233,257]" gridcol="0" gridrow="1" pageId="3" pageNumber="312">1</th>
<td id="76D68840FFC4E7F9FEA18894ACC0FE99" box="[345,497,233,257]" gridcol="1" gridrow="1" pageId="3" pageNumber="312">Thrissur</td>
<td id="76D68840FFC4E7F9FD918894AE3DFE99" box="[617,780,233,257]" gridcol="2" gridrow="1" pageId="3" pageNumber="312">Peechi</td>
<td id="76D68840FFC4E7F9FC628894A926FE99" box="[922,1047,233,257]" gridcol="3" gridrow="1" pageId="3" pageNumber="312">10.527</td>
<td id="76D68840FFC4E7F9FB468894A87CFE99" box="[1214,1357,233,257]" gridcol="4" gridrow="1" pageId="3" pageNumber="312">76.3608</td>
</tr>
<tr id="3507E13CFFC4E7F9FF5B896CA87CFEB1" box="[163,1357,273,297]" gridrow="2" pageId="3" pageNumber="312">
<th id="76D68840FFC4E7F9FF5B896CADF8FEB1" box="[163,201,273,297]" gridcol="0" gridrow="2" pageId="3" pageNumber="312">2</th>
<td id="76D68840FFC4E7F9FEA1896CACC0FEB1" box="[345,497,273,297]" gridcol="1" gridrow="2" pageId="3" pageNumber="312">Thrissur</td>
<td id="76D68840FFC4E7F9FD91896CAE3DFEB1" box="[617,780,273,297]" gridcol="2" gridrow="2" pageId="3" pageNumber="312">Valarkkavu</td>
<td id="76D68840FFC4E7F9FC62896CA926FEB1" box="[922,1047,273,297]" gridcol="3" gridrow="2" pageId="3" pageNumber="312">10.506</td>
<td id="76D68840FFC4E7F9FB46896CA87CFEB1" box="[1214,1357,273,297]" gridcol="4" gridrow="2" pageId="3" pageNumber="312">76.2365</td>
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<tr id="3507E13CFFC4E7F9FF5B8944A87CFECA" box="[163,1357,313,338]" gridrow="3" pageId="3" pageNumber="312">
<th id="76D68840FFC4E7F9FF5B8944ADF8FECA" box="[163,201,313,338]" gridcol="0" gridrow="3" pageId="3" pageNumber="312">3</th>
<td id="76D68840FFC4E7F9FEA18944ACC0FECA" box="[345,497,313,338]" gridcol="1" gridrow="3" pageId="3" pageNumber="312">Thrissur</td>
<td id="76D68840FFC4E7F9FD918944AE3DFECA" box="[617,780,313,338]" gridcol="2" gridrow="3" pageId="3" pageNumber="312">Vaniyampara</td>
<td id="76D68840FFC4E7F9FC628944A926FECA" box="[922,1047,313,338]" gridcol="3" gridrow="3" pageId="3" pageNumber="312">10.5757</td>
<td id="76D68840FFC4E7F9FB468944A87CFECA" box="[1214,1357,313,338]" gridcol="4" gridrow="3" pageId="3" pageNumber="312">76.4006</td>
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<tr id="3507E13CFFC4E7F9FF5B891FA87CFEE2" box="[163,1357,354,378]" gridrow="4" pageId="3" pageNumber="312">
<th id="76D68840FFC4E7F9FF5B891FADF8FEE2" box="[163,201,354,378]" gridcol="0" gridrow="4" pageId="3" pageNumber="312">4</th>
<td id="76D68840FFC4E7F9FEA1891FACC0FEE2" box="[345,497,354,378]" gridcol="1" gridrow="4" pageId="3" pageNumber="312">Thrissur</td>
<td id="76D68840FFC4E7F9FD91891FAE3DFEE2" box="[617,780,354,378]" gridcol="2" gridrow="4" pageId="3" pageNumber="312">Chazhikad</td>
<td id="76D68840FFC4E7F9FC62891FA926FEE2" box="[922,1047,354,378]" gridcol="3" gridrow="4" pageId="3" pageNumber="312">10.2301</td>
<td id="76D68840FFC4E7F9FB46891FA87CFEE2" box="[1214,1357,354,378]" gridcol="4" gridrow="4" pageId="3" pageNumber="312">76.206</td>
</tr>
<tr id="3507E13CFFC4E7F9FF5B89F7A87CFE3A" box="[163,1357,394,418]" gridrow="5" pageId="3" pageNumber="312">
<th id="76D68840FFC4E7F9FF5B89F7ADF8FE3A" box="[163,201,394,418]" gridcol="0" gridrow="5" pageId="3" pageNumber="312">5</th>
<td id="76D68840FFC4E7F9FEA189F7ACC0FE3A" box="[345,497,394,418]" gridcol="1" gridrow="5" pageId="3" pageNumber="312">Thrissur</td>
<td id="76D68840FFC4E7F9FD9189F7AE3DFE3A" box="[617,780,394,418]" gridcol="2" gridrow="5" pageId="3" pageNumber="312">Velupadam</td>
<td id="76D68840FFC4E7F9FC6289F7A926FE3A" box="[922,1047,394,418]" gridcol="3" gridrow="5" pageId="3" pageNumber="312">10.4328</td>
<td id="76D68840FFC4E7F9FB4689F7A87CFE3A" box="[1214,1357,394,418]" gridcol="4" gridrow="5" pageId="3" pageNumber="312">76.3474</td>
</tr>
<tr id="3507E13CFFC4E7F9FF5B89CFA87CFE52" box="[163,1357,434,458]" gridrow="6" pageId="3" pageNumber="312">
<th id="76D68840FFC4E7F9FF5B89CFADF8FE52" box="[163,201,434,458]" gridcol="0" gridrow="6" pageId="3" pageNumber="312">6</th>
<td id="76D68840FFC4E7F9FEA189CFACC0FE52" box="[345,497,434,458]" gridcol="1" gridrow="6" pageId="3" pageNumber="312">Thrissur</td>
<td id="76D68840FFC4E7F9FD9189CFAE3DFE52" box="[617,780,434,458]" gridcol="2" gridrow="6" pageId="3" pageNumber="312">Padukad</td>
<td id="76D68840FFC4E7F9FC6289CFA926FE52" box="[922,1047,434,458]" gridcol="3" gridrow="6" pageId="3" pageNumber="312">10.5621</td>
<td id="76D68840FFC4E7F9FB4689CFA87CFE52" box="[1214,1357,434,458]" gridcol="4" gridrow="6" pageId="3" pageNumber="312">76.2152</td>
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<td id="76D68840FFC4E7F9FEA189A7ACC0FE6A" box="[345,497,474,498]" gridcol="1" gridrow="7" pageId="3" pageNumber="312">Thrissur</td>
<td id="76D68840FFC4E7F9FD9189A7AE3DFE6A" box="[617,780,474,498]" gridcol="2" gridrow="7" pageId="3" pageNumber="312">Wadakkanchery</td>
<td id="76D68840FFC4E7F9FC6289A7A926FE6A" box="[922,1047,474,498]" gridcol="3" gridrow="7" pageId="3" pageNumber="312">10.5236</td>
<td id="76D68840FFC4E7F9FB4689A7A87CFE6A" box="[1214,1357,474,498]" gridcol="4" gridrow="7" pageId="3" pageNumber="312">76.2166</td>
</tr>
<tr id="3507E13CFFC4E7F9FF5B8A7FA87CFD83" box="[163,1357,514,539]" gridrow="8" pageId="3" pageNumber="312">
<th id="76D68840FFC4E7F9FF5B8A7FADF8FD83" box="[163,201,514,539]" gridcol="0" gridrow="8" pageId="3" pageNumber="312">8</th>
<td id="76D68840FFC4E7F9FEA18A7FACC0FD83" box="[345,497,514,539]" gridcol="1" gridrow="8" pageId="3" pageNumber="312">Palakkad</td>
<td id="76D68840FFC4E7F9FD918A7FAE3DFD83" box="[617,780,514,539]" gridcol="2" gridrow="8" pageId="3" pageNumber="312">Alathur</td>
<td id="76D68840FFC4E7F9FC628A7FA926FD83" box="[922,1047,514,539]" gridcol="3" gridrow="8" pageId="3" pageNumber="312">10.6404</td>
<td id="76D68840FFC4E7F9FB468A7FA87CFD83" box="[1214,1357,514,539]" gridcol="4" gridrow="8" pageId="3" pageNumber="312">76.528</td>
</tr>
<tr id="3507E13CFFC4E7F9FF5B8A56A87CFDDB" box="[163,1357,555,579]" gridrow="9" pageId="3" pageNumber="312">
<th id="76D68840FFC4E7F9FF5B8A56ADF8FDDB" box="[163,201,555,579]" gridcol="0" gridrow="9" pageId="3" pageNumber="312">9</th>
<td id="76D68840FFC4E7F9FEA18A56ACC0FDDB" box="[345,497,555,579]" gridcol="1" gridrow="9" pageId="3" pageNumber="312">Palakkad</td>
<td id="76D68840FFC4E7F9FD918A56AE3DFDDB" box="[617,780,555,579]" gridcol="2" gridrow="9" pageId="3" pageNumber="312">Alathur</td>
<td id="76D68840FFC4E7F9FC628A56A926FDDB" box="[922,1047,555,579]" gridcol="3" gridrow="9" pageId="3" pageNumber="312">10.6351</td>
<td id="76D68840FFC4E7F9FB468A56A87CFDDB" box="[1214,1357,555,579]" gridcol="4" gridrow="9" pageId="3" pageNumber="312">76.5331</td>
</tr>
<tr id="3507E13CFFC4E7F9FF5B8A2EA87CFDF3" box="[163,1357,595,619]" gridrow="10" pageId="3" pageNumber="312">
<th id="76D68840FFC4E7F9FF5B8A2EADF8FDF3" box="[163,201,595,619]" gridcol="0" gridrow="10" pageId="3" pageNumber="312">10</th>
<td id="76D68840FFC4E7F9FEA18A2EACC0FDF3" box="[345,497,595,619]" gridcol="1" gridrow="10" pageId="3" pageNumber="312">Ernamkulam</td>
<td id="76D68840FFC4E7F9FD918A2EAE3DFDF3" box="[617,780,595,619]" gridcol="2" gridrow="10" pageId="3" pageNumber="312">Koonammavu</td>
<td id="76D68840FFC4E7F9FC628A2EA926FDF3" box="[922,1047,595,619]" gridcol="3" gridrow="10" pageId="3" pageNumber="312">10.0963</td>
<td id="76D68840FFC4E7F9FB468A2EA87CFDF3" box="[1214,1357,595,619]" gridcol="4" gridrow="10" pageId="3" pageNumber="312">76.2648</td>
</tr>
<tr id="3507E13CFFC4E7F9FF5B8A06A87CFD0B" box="[163,1357,635,659]" gridrow="11" pageId="3" pageNumber="312">
<th id="76D68840FFC4E7F9FF5B8A06ADF8FD0B" box="[163,201,635,659]" gridcol="0" gridrow="11" pageId="3" pageNumber="312">11</th>
<td id="76D68840FFC4E7F9FEA18A06ACC0FD0B" box="[345,497,635,659]" gridcol="1" gridrow="11" pageId="3" pageNumber="312">Pathanamthitta</td>
<td id="76D68840FFC4E7F9FD918A06AE3DFD0B" box="[617,780,635,659]" gridcol="2" gridrow="11" pageId="3" pageNumber="312">Konni</td>
<td id="76D68840FFC4E7F9FC628A06A926FD0B" box="[922,1047,635,659]" gridcol="3" gridrow="11" pageId="3" pageNumber="312">9.23041</td>
<td id="76D68840FFC4E7F9FB468A06A87CFD0B" box="[1214,1357,635,659]" gridcol="4" gridrow="11" pageId="3" pageNumber="312">76.8078</td>
</tr>
<tr id="3507E13CFFC4E7F9FF5B8ADEA87CFD24" box="[163,1357,675,700]" gridrow="12" pageId="3" pageNumber="312">
<th id="76D68840FFC4E7F9FF5B8ADEADF8FD24" box="[163,201,675,700]" gridcol="0" gridrow="12" pageId="3" pageNumber="312">12</th>
<td id="76D68840FFC4E7F9FEA18ADEACC0FD24" box="[345,497,675,700]" gridcol="1" gridrow="12" pageId="3" pageNumber="312">Kannur</td>
<td id="76D68840FFC4E7F9FD918ADEAE3DFD24" box="[617,780,675,700]" gridcol="2" gridrow="12" pageId="3" pageNumber="312">Padiyur</td>
<td id="76D68840FFC4E7F9FC628ADEA926FD24" box="[922,1047,675,700]" gridcol="3" gridrow="12" pageId="3" pageNumber="312">11.9944</td>
<td id="76D68840FFC4E7F9FB468ADEA87CFD24" box="[1214,1357,675,700]" gridcol="4" gridrow="12" pageId="3" pageNumber="312">75.619</td>
</tr>
<tr id="3507E13CFFC4E7F9FF5B8AB1A87CFD7C" box="[163,1357,716,740]" gridrow="13" pageId="3" pageNumber="312">
<th id="76D68840FFC4E7F9FF5B8AB1ADF8FD7C" box="[163,201,716,740]" gridcol="0" gridrow="13" pageId="3" pageNumber="312">13</th>
<td id="76D68840FFC4E7F9FEA18AB1ACC0FD7C" box="[345,497,716,740]" gridcol="1" gridrow="13" pageId="3" pageNumber="312">Idukki</td>
<td id="76D68840FFC4E7F9FD918AB1AE3DFD7C" box="[617,780,716,740]" gridcol="2" gridrow="13" pageId="3" pageNumber="312">Painavu</td>
<td id="76D68840FFC4E7F9FC628AB1A926FD7C" box="[922,1047,716,740]" gridcol="3" gridrow="13" pageId="3" pageNumber="312">9.86493</td>
<td id="76D68840FFC4E7F9FB468AB1A87CFD7C" box="[1214,1357,716,740]" gridcol="4" gridrow="13" pageId="3" pageNumber="312">76.9577</td>
</tr>
</table>
</paragraph>
<paragraph id="8B88E37EFFC41805FF3F8B57AC9BFC28" blockId="3.[151,1437,810,2025]" pageId="3" pageNumber="312">
Body sub-cylindrical, elongated and tapering posteriorly; length up to
<quantity id="4CCF4E9BFFC41805FC208B57A917FCDD" box="[984,1062,810,837]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="312" unit="mm" value="13.0">13 mm</quantity>
(
<figureCitation id="130CFFFBFFC41805FBCD8B57A948FCDC" box="[1077,1145,810,836]" captionStart="FIGURES 2‒5" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[210,1378,188,1916]" captionTargetId="figure-21@4.[203,1384,181,1919]" captionTargetPageId="4" captionText="FIGURES 2‒5. Ctenolepisma longicaudatum. 2) Dorsal view. 3) Body scales. 4) Labial palps with five sensory papillae on apical article. 5) SEM of labial palp with more than 10 sensory papillae.Arrow: labial palp." figureDoi="http://doi.org/10.5281/zenodo.14735036" httpUri="https://zenodo.org/record/14735036/files/figure.png" pageId="3" pageNumber="312">Fig. 2</figureCitation>
). Epidermic pigmentation light golden yellow dorsally, lighter ventrally. Dorsally covered with dark grayish or brownish scales and ventrally with light yellowish scales. Scales rounded with dense parallel ribs; relatively homogeneous structures, only differing in size (
<figureCitation id="130CFFFBFFC41805FEAF8BEBACADFC28" box="[343,412,918,944]" captionStart="FIGURES 2‒5" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[210,1378,188,1916]" captionTargetId="figure-21@4.[203,1384,181,1919]" captionTargetPageId="4" captionText="FIGURES 2‒5. Ctenolepisma longicaudatum. 2) Dorsal view. 3) Body scales. 4) Labial palps with five sensory papillae on apical article. 5) SEM of labial palp with more than 10 sensory papillae.Arrow: labial palp." figureDoi="http://doi.org/10.5281/zenodo.14735036" httpUri="https://zenodo.org/record/14735036/files/figure.png" pageId="3" pageNumber="312">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8B88E37EFFC41805FF3F8BC7A9CAFB11" blockId="3.[151,1437,810,2025]" pageId="3" pageNumber="312">
Head wider (
<quantity id="4CCF4E9BFFC41805FEA48BC7AF32FC4C" box="[348,515,954,981]" metricMagnitude="-3" metricUnit="m" metricValue="1.525" metricValueMax="1.7" metricValueMin="1.35" pageId="3" pageNumber="312" unit="mm" value="1.525" valueMax="1.7" valueMin="1.35">1.35–1.70 mm</quantity>
) than long (
<quantity id="4CCF4E9BFFC41805FD688BC7AE2FFC4C" box="[656,798,954,980]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="3" pageNumber="312" unit="mm" value="1.0" valueMax="1.1" valueMin="0.9">0.9–1.1 mm</quantity>
), with tufts of bifid and pectinate macrosetae on frontal region, 1+1 tufts of macrosetae on clypeus and 1+1 small tufts of macrosetae on labrum. Eyes black with 12 ommatidia, located well behind the antennae. Antennae length
<quantity id="4CCF4E9BFFC41805FC948C7FAEDAFB85" box="[876,1003,1026,1053]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="3" pageNumber="312" unit="mm" value="11.5" valueMax="13.0" valueMin="10.0">10‒13 mm</quantity>
. Five-segmented maxillary palp, with the apical article about 5.9– 6.5 times longer than wide and almost same length as that of the penultimate article. Four-segmented labial palps with distal article wider at the apex than at the base (L/W 1–1.3) and usually with five sensory papillae (
<figureCitation id="130CFFFBFFC41805FEA48C13ACAEFB10" box="[348,415,1134,1160]" captionStart="FIGURES 2‒5" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[210,1378,188,1916]" captionTargetId="figure-21@4.[203,1384,181,1919]" captionTargetPageId="4" captionText="FIGURES 2‒5. Ctenolepisma longicaudatum. 2) Dorsal view. 3) Body scales. 4) Labial palps with five sensory papillae on apical article. 5) SEM of labial palp with more than 10 sensory papillae.Arrow: labial palp." figureDoi="http://doi.org/10.5281/zenodo.14735036" httpUri="https://zenodo.org/record/14735036/files/figure.png" pageId="3" pageNumber="312">Fig. 4</figureCitation>
) but sometimes with more than 10 papillae arranged in a single row (
<figureCitation id="130CFFFBFFC41805FB518C13A9DCFB11" box="[1193,1261,1134,1161]" captionStart="FIGURES 2‒5" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[210,1378,188,1916]" captionTargetId="figure-21@4.[203,1384,181,1919]" captionTargetPageId="4" captionText="FIGURES 2‒5. Ctenolepisma longicaudatum. 2) Dorsal view. 3) Body scales. 4) Labial palps with five sensory papillae on apical article. 5) SEM of labial palp with more than 10 sensory papillae.Arrow: labial palp." figureDoi="http://doi.org/10.5281/zenodo.14735036" httpUri="https://zenodo.org/record/14735036/files/figure.png" pageId="3" pageNumber="312">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="8B88E37EFFC41805FF3F8CEFACB7FA54" blockId="3.[151,1437,810,2025]" pageId="3" pageNumber="312">
Thorax wider than abdomen; thorax width up to
<quantity id="4CCF4E9BFFC41805FD1B8CEFAE79FB35" box="[739,840,1170,1197]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" pageId="3" pageNumber="312" unit="mm" value="3.25">3.25 mm</quantity>
, thorax length up to
<quantity id="4CCF4E9BFFC41805FBD48CEFA9B4FB35" box="[1068,1157,1170,1197]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="3" pageNumber="312" unit="mm" value="4.1">4.1 mm</quantity>
. Pronotal collar with 4–5 rows of bristles in the centre and single row towards the anterolateral side; lateral margin of pronotum with (9–10) + (10–11) bristle combs of 3–6 macrosetae each. Trichobothrial areas of the pronotum situated on inner side of last (N) and the anterior on the inner side of antepenultimate (N-4) bristle-combs (
<figureCitation id="130CFFFBFFC41805FBF48C83A960FA80" box="[1036,1105,1278,1304]" captionStart="FIGURES 6‒8" captionStartId="5.[151,264,1334,1358]" captionTargetBox="[157,1429,186,1304]" captionTargetId="figure-17@5.[151,1436,181,1310]" captionTargetPageId="5" captionText="FIGURES 6‒8. Ctenolepisma longicaudatum, bristle combs and trichobothrial areas. 6) Pronotum. 7) Mesonotum. 8) Metanotum." figureDoi="http://doi.org/10.5281/zenodo.14735038" httpUri="https://zenodo.org/record/14735038/files/figure.png" pageId="3" pageNumber="312">Fig. 6</figureCitation>
). Mesonotum lateral margins with (11–12) + (12–15) bristle combs of 3–6 macrosetae (
<figureCitation id="130CFFFBFFC41805FCEF8D5FAE6BFAA4" box="[791,858,1314,1340]" captionStart="FIGURES 6‒8" captionStartId="5.[151,264,1334,1358]" captionTargetBox="[157,1429,186,1304]" captionTargetId="figure-17@5.[151,1436,181,1310]" captionTargetPageId="5" captionText="FIGURES 6‒8. Ctenolepisma longicaudatum, bristle combs and trichobothrial areas. 6) Pronotum. 7) Mesonotum. 8) Metanotum." figureDoi="http://doi.org/10.5281/zenodo.14735038" httpUri="https://zenodo.org/record/14735038/files/figure.png" pageId="3" pageNumber="312">Fig. 7</figureCitation>
). Trichobothrial areas of the mesonotum situated on last (N) and the anterior on outer side of antepenultimate (N-2) bristle-combs. Metanotum (
<figureCitation id="130CFFFBFFC41805FB588D3BA9D5FAF8" box="[1184,1252,1350,1376]" captionStart="FIGURES 6‒8" captionStartId="5.[151,264,1334,1358]" captionTargetBox="[157,1429,186,1304]" captionTargetId="figure-17@5.[151,1436,181,1310]" captionTargetPageId="5" captionText="FIGURES 6‒8. Ctenolepisma longicaudatum, bristle combs and trichobothrial areas. 6) Pronotum. 7) Mesonotum. 8) Metanotum." figureDoi="http://doi.org/10.5281/zenodo.14735038" httpUri="https://zenodo.org/record/14735038/files/figure.png" pageId="3" pageNumber="312">Fig. 8</figureCitation>
) lateral margins with (10–11) + (10–11) bristle combs of 2–6 macrosetae. Trichobothrial areas of the metanotum located on last (N) and the anterior on outer side of (N-1) bristle-combs. All nota provided with 1+1 bristle combs of 6–7 setae on the posterolateral region.
</paragraph>
<paragraph id="8B88E37EFFC41805FF3F8DAAAF5AF918" blockId="3.[151,1437,810,2025]" pageId="3" pageNumber="312">
Prosternum (L/W: 1.3–1.4) and mesosternum (L/W: 1.1–1.3), slightly longer than wide; metanotum wider than long (L/W: 0.85–0.1) (
<figureCitation id="130CFFFBFFC41805FE608D87ACECF98C" box="[408,477,1530,1556]" captionStart="FIGURES 9‒14" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[222,1365,182,1919]" captionTargetId="figure-21@6.[217,1371,181,1919]" captionTargetPageId="6" captionText="FIGURES 9‒14. Ctenolepisma longicaudatum. 9) Prosternum. 10) Mesosternum. 11) Metasternum. 12) Urotergite X. 13) Male genital area with coxite IX. 14) Female genital area with ovipositor and coxite IX." figureDoi="http://doi.org/10.5281/zenodo.14735040" httpUri="https://zenodo.org/record/14735040/files/figure.png" pageId="3" pageNumber="312">Fig. 9</figureCitation>
). Prosternum with 3–5+ 3–5 bristle combs of 3–10 macrosetae on its subapical area. Mesosternum (
<figureCitation id="130CFFFBFFC41805FEC78E63ACA0F9A0" box="[319,401,1566,1592]" captionStart="FIGURES 9‒14" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[222,1365,182,1919]" captionTargetId="figure-21@6.[217,1371,181,1919]" captionTargetPageId="6" captionText="FIGURES 9‒14. Ctenolepisma longicaudatum. 9) Prosternum. 10) Mesosternum. 11) Metasternum. 12) Urotergite X. 13) Male genital area with coxite IX. 14) Female genital area with ovipositor and coxite IX." figureDoi="http://doi.org/10.5281/zenodo.14735040" httpUri="https://zenodo.org/record/14735040/files/figure.png" pageId="3" pageNumber="312">Fig. 10</figureCitation>
) with 2–5 + 2–5 bristle combs of 4–12 macrosetae on its subapical area. Metasternum (
<figureCitation id="130CFFFBFFC41805FA978E63AD83F9C4" captionStart="FIGURES 9‒14" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[222,1365,182,1919]" captionTargetId="figure-21@6.[217,1371,181,1919]" captionTargetPageId="6" captionText="FIGURES 9‒14. Ctenolepisma longicaudatum. 9) Prosternum. 10) Mesosternum. 11) Metasternum. 12) Urotergite X. 13) Male genital area with coxite IX. 14) Female genital area with ovipositor and coxite IX." figureDoi="http://doi.org/10.5281/zenodo.14735040" httpUri="https://zenodo.org/record/14735040/files/figure.png" pageId="3" pageNumber="312">Fig. 11</figureCitation>
) with 1 + 1 subapical combs of more than 20 macrochaetae, arranged in a single row; few scattered setae also visible between the comb and the margin.
</paragraph>
<paragraph id="8B88E37EFFC41805FF3F8EF6ACAAF8AC" blockId="3.[151,1437,810,2025]" pageId="3" pageNumber="312">
Legs stout; protibiae 2.4–2.8 times longer than wide; mesotibiae 3.1–3.82 times longer than wide and about 1.2–1.3 times longer than the protibiae; metatibiae 4–5 times longer than wide and about 1.87–1.9 times longer than the protibiae. Tarsi L/W
<collectionCode id="ED267BBBFFC41805FE278EAEACC9F975" box="[479,504,1747,1773]" country="Russia" name="Paleontological Institute" pageId="3" pageNumber="312">PI</collectionCode>
6.7–7.5, PII 7.7–12.6,
<collectionCode id="ED267BBBFFC41805FD008EAEAE15F975" box="[760,804,1747,1773]" pageId="3" pageNumber="312">PIII</collectionCode>
10.5–13.66. Pretarsi with two curved lateral claws and a medial claw. Coxae and femora covered ventrally by rounded scales similar to those of the body; remaining articles of leg devoid of scales.
</paragraph>
<paragraph id="8B88E37EFFC41805FF3F8F43A8B1F870" blockId="3.[151,1437,810,2025]" pageId="3" pageNumber="312">
Urotergite I with 1+1 infra-lateral bristle-combs composed of 6–8 macrosetae. Urotergites II–
<collectionCode id="ED267BBBFFC41805FB3D8F42A9D3F8C1" box="[1221,1250,1855,1881]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="3" pageNumber="312">VI</collectionCode>
with 3+3 bristle-combs; infra lateral bristle combs with 5–10 macrosetae, lateral combs with 5–7 macrosetae and sub-median bristle combs with 6–7 macrosetae. Urotergite VII and VIII with 2+2 bristle-combs, each lateral bristle comb composed of 11 macrosetae and sub-median bristle combs with 7–8 macrosetae. Urotergite IX without bristle combs. Urotergite X (
<figureCitation id="130CFFFBFFC41805FF438FB3AC3DF870" box="[187,268,1998,2024]" captionStart="FIGURES 9‒14" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[222,1365,182,1919]" captionTargetId="figure-21@6.[217,1371,181,1919]" captionTargetPageId="6" captionText="FIGURES 9‒14. Ctenolepisma longicaudatum. 9) Prosternum. 10) Mesosternum. 11) Metasternum. 12) Urotergite X. 13) Male genital area with coxite IX. 14) Female genital area with ovipositor and coxite IX." figureDoi="http://doi.org/10.5281/zenodo.14735040" httpUri="https://zenodo.org/record/14735040/files/figure.png" pageId="3" pageNumber="312">Fig. 12</figureCitation>
) trapezoidal in shape with 1+1 bristle-combs of 8–10 macrosetae; few scattered setae towards the tip.
</paragraph>
<caption id="DF48B3F6FFC31802FF6F8FEAA906F84B" ID-DOI="http://doi.org/10.5281/zenodo.14735036" ID-Zenodo-Dep="14735036" httpUri="https://zenodo.org/record/14735036/files/figure.png" pageId="4" pageNumber="313" startId="4.[151,264,1943,1967]" targetBox="[210,1378,188,1916]" targetPageId="4" targetType="figure">
<paragraph id="8B88E37EFFC31802FF6F8FEAA906F84B" blockId="4.[151,1436,1943,2004]" pageId="4" pageNumber="313">
<emphasis id="B9433F6CFFC31802FF6F8FEAAC0CF837" bold="true" box="[151,317,1943,1967]" pageId="4" pageNumber="313">FIGURES 2‒5.</emphasis>
<emphasis id="B9433F6CFFC31802FEBC8FEAAF46F837" box="[324,631,1943,1967]" italics="true" pageId="4" pageNumber="313">
<taxonomicName id="4C3798FDFFC31802FEBC8FEAAF42F837" baseAuthorityName="Escherich" baseAuthorityYear="1905" box="[324,627,1943,1967]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="4" pageNumber="313" phylum="Arthropoda" rank="species" species="longicaudatum">Ctenolepisma longicaudatum</taxonomicName>
.
</emphasis>
2) Dorsal view. 3) Body scales. 4) Labial palps with five sensory papillae on apical article. 5) SEM of labial palp with more than 10 sensory papillae. Arrow: labial palp.
</paragraph>
</caption>
<caption id="DF48B3F6FFC21803FF6F8D4BAC23FAEA" ID-DOI="http://doi.org/10.5281/zenodo.14735038" ID-Zenodo-Dep="14735038" httpUri="https://zenodo.org/record/14735038/files/figure.png" pageId="5" pageNumber="314" startId="5.[151,264,1334,1358]" targetBox="[157,1429,186,1304]" targetPageId="5" targetType="figure">
<paragraph id="8B88E37EFFC21803FF6F8D4BAC23FAEA" blockId="5.[151,1436,1334,1394]" pageId="5" pageNumber="314">
<emphasis id="B9433F6CFFC21803FF6F8D4BAC73FAD6" bold="true" box="[151,322,1334,1358]" pageId="5" pageNumber="314">FIGURES 6‒8.</emphasis>
<taxonomicName id="4C3798FDFFC21803FEB68D4BAF4EFAD6" baseAuthorityName="Escherich" baseAuthorityYear="1905" box="[334,639,1334,1358]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="5" pageNumber="314" phylum="Arthropoda" rank="species" species="longicaudatum">
<emphasis id="B9433F6CFFC21803FEB68D4BAF4EFAD6" box="[334,639,1334,1358]" italics="true" pageId="5" pageNumber="314">Ctenolepisma longicaudatum</emphasis>
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, bristle combs and trichobothrial areas. 6) Pronotum. 7) Mesonotum. 8) Metanotum.
</paragraph>
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Urosternites I and II without setae. Urosternites III–VII with 1+1 lateral combs of 17–20 macrosetae. The lateral combs of urosternite VIII with 12–13 macrosetae. The number of macrosetae per bristle comb on urotergites and urosternites as in
<tableCitation id="C6B5D6C5FFC21803FEA08E70AC9DF9BF" box="[344,428,1549,1575]" captionStart="TABLE 4" captionStartId="12.[151,239,151,175]" captionText="TABLE 4. Number of macrosetae per bristle comb on urotergites and urosternites in C. longicaudatum and C." pageId="5" pageNumber="314">Table 4</tableCitation>
. Both sexes bear two pairs of stylets inserted on abdominal segments VIII and IX (
<figureCitation id="130CFFFBFFC21803FAC38E70AD84F9D3" captionStart="FIGURES 9‒14" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[222,1365,182,1919]" captionTargetId="figure-21@6.[217,1371,181,1919]" captionTargetPageId="6" captionText="FIGURES 9‒14. Ctenolepisma longicaudatum. 9) Prosternum. 10) Mesosternum. 11) Metasternum. 12) Urotergite X. 13) Male genital area with coxite IX. 14) Female genital area with ovipositor and coxite IX." figureDoi="http://doi.org/10.5281/zenodo.14735040" httpUri="https://zenodo.org/record/14735040/files/figure.png" pageId="5" pageNumber="314">Figs. 13, 14</figureCitation>
), armed apically with several strong spines. Ratio of length of styli IX/ length of styli VIII is 1.25–2.
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Coxite IX in male with fringe of setae along internal margin; internal process about 1.8–4.5 times longer than the external process and 1.08–1.18 (L/W) times as long as broad at its base. Penis with numerous apical setae. Parameres absent (
<figureCitation id="130CFFFBFFC21803FE928EE0AC8CF92F" box="[362,445,1693,1719]" captionStart="FIGURES 9‒14" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[222,1365,182,1919]" captionTargetId="figure-21@6.[217,1371,181,1919]" captionTargetPageId="6" captionText="FIGURES 9‒14. Ctenolepisma longicaudatum. 9) Prosternum. 10) Mesosternum. 11) Metasternum. 12) Urotergite X. 13) Male genital area with coxite IX. 14) Female genital area with ovipositor and coxite IX." figureDoi="http://doi.org/10.5281/zenodo.14735040" httpUri="https://zenodo.org/record/14735040/files/figure.png" pageId="5" pageNumber="314">Fig. 13</figureCitation>
).
</paragraph>
<paragraph id="8B88E37EFFC21803FF3F8EBCAE5CF8DF" blockId="5.[151,1437,1477,1863]" pageId="5" pageNumber="314">
Genital region of female shown in
<figureCitation id="130CFFFBFFC21803FDA98EBCAF95F943" box="[593,676,1729,1755]" captionStart="FIGURES 9‒14" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[222,1365,182,1919]" captionTargetId="figure-21@6.[217,1371,181,1919]" captionTargetPageId="6" captionText="FIGURES 9‒14. Ctenolepisma longicaudatum. 9) Prosternum. 10) Mesosternum. 11) Metasternum. 12) Urotergite X. 13) Male genital area with coxite IX. 14) Female genital area with ovipositor and coxite IX." figureDoi="http://doi.org/10.5281/zenodo.14735040" httpUri="https://zenodo.org/record/14735040/files/figure.png" pageId="5" pageNumber="314">Fig. 14</figureCitation>
. Inner process of coxite IX long, triangular and pointed at tip about 2–2.7 times longer than wide at its base and 2.7–5 times longer than the external process. Very long ovipositor with 40–48 divisions and its length measures
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from the base of urosternite VIII; surpassing the apex of the internal processes of coxite IX by 2–3.75 times coxite IX length.
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<taxonomicName id="4C754D01FFA2D679FF3CFAA0C1CC2A63" ID-CoL="46LQW" authority="Bomans &amp; Miyashita, 1997" authorityName="Bomans &amp; Miyashita" authorityYear="1997" box="[151,761,1345,1371]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="356" phylum="Arthropoda" rank="species" species="brochieri">
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<emphasis id="B901EA90FFA2D679FF3CFAA0C2AA2A63" bold="true" box="[151,415,1345,1371]" italics="true" pageId="12" pageNumber="356">Neolucanus brochieri</emphasis>
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Suggested Chinese common name:
<emphasis id="B901EA90FFA2D679FD8CFA87C1FA2A45" bold="true" box="[551,719,1382,1405]" pageId="12" pageNumber="356">DZae圆OiNJƤ</emphasis>
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<figureCitation id="134E2A07FFA2D679FE8FFA68C2502A9B" box="[292,357,1417,1443]" captionStart="FIGURE 2" captionStartId="4.[151,250,1771,1795]" captionTargetBox="[158,1429,186,1740]" captionTargetId="figure-18@4.[151,1436,181,1747]" captionTargetPageId="4" captionText="FIGURE 2. Male habitus of Neolucanus species. A–B, N. horizontalis Xin, Zhong &amp; Qi, new species, paratype, 33.7 mm, mesodont (Jinping, Yunnan); C–D, N. fuscus Didier, 1926, 34.7mm, priodont (Wuming, Guangxi); E–F, N. brochieri Bomans &amp; Miyashita, 1997, 31.4 mm, priodont (Gonshan, Yunnan). A, C, E, dorsal view; B, D, F, ventral view. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14735139" httpUri="https://zenodo.org/record/14735139/files/figure.png" pageId="12" pageNumber="356">2E–F</figureCitation>
; 3M–P; 4I–L; 5M–P; 6D, H, L; 7D
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<emphasis id="B901EA90FFA2D679FF3CFA2FC2472ADE" box="[151,370,1486,1510]" italics="true" pageId="12" pageNumber="356">Neolucanus brochieri</emphasis>
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<materialsCitation id="3B1D3CDFFFA2D679FF3CF9F5C17F2916" box="[151,586,1556,1582]" country="Myanmar" location="Kachin State" pageId="12" pageNumber="356" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="B901EA90FFA2D679FF3CF9F5C2032916" bold="true" box="[151,310,1556,1582]" pageId="12" pageNumber="356">
<typeStatus id="54CE8820FFA2D679FF3CF9F5C3E72916" box="[151,210,1556,1582]" pageId="12" pageNumber="356">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3627612FFA2D679FE97F9F5C29E2916" box="[316,427,1556,1582]" name="Myanmar" pageId="12" pageNumber="356">Myanmar</collectingCountry>
,
<collectingRegion id="49B1F860FFA2D679FE1DF9F5C17F2916" box="[438,586,1556,1582]" country="Myanmar" name="Kachin" pageId="12" pageNumber="356">Kachin State</collectingRegion>
</materialsCitation>
.
</paragraph>
<paragraph id="8BCA3682FFA2D679FF6CF9D9C10F296A" blockId="12.[151,1437,1556,2014]" box="[199,570,1592,1618]" pageId="12" pageNumber="356">
<emphasis id="B901EA90FFA2D679FF6CF9D9C2DA296A" bold="true" box="[199,495,1592,1618]" pageId="12" pageNumber="356">
<typeStatus id="54CE8820FFA2D679FF6CF9D9C237296A" box="[199,258,1592,1618]" pageId="12" pageNumber="356">Type</typeStatus>
material examined.
</emphasis>
None.
</paragraph>
<paragraph id="8BCA3682FFA2D679FF6CF9BDC7BD29DA" blockId="12.[151,1437,1556,2014]" pageId="12" pageNumber="356">
<emphasis id="B901EA90FFA2D679FF6CF9BDC10F294E" bold="true" box="[199,570,1628,1654]" pageId="12" pageNumber="356">Additional material examined.</emphasis>
<materialsCitation id="3B1D3CDFFFA2D679FDEFF9BDC03629A2" collectionCode="CXTZ" country="China" county="Gongshan County" location="Dulongjiang valley" municipality="Nujiang Prefecture" pageId="12" pageNumber="356" specimenCount="2" specimenCount-male="2" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B901EA90FFA2D679FDEFF9BDC1A3294E" bold="true" box="[580,662,1628,1654]" pageId="12" pageNumber="356">
<collectingCountry id="F3627612FFA2D679FDEFF9BDC1A4294E" box="[580,657,1628,1654]" name="China" pageId="12" pageNumber="356">China</collectingCountry>
:
</emphasis>
<specimenCount id="9D73FD0BFFA2D679FD0BF9BDC1CC294E" box="[672,761,1628,1654]" count="2" pageId="12" pageNumber="356" type="male">2 males</specimenCount>
(
<collectionCode id="ED64AE47FFA2D679FCA0F9BDC06F294E" box="[779,858,1628,1654]" pageId="12" pageNumber="356">CXTZ</collectionCode>
),
<collectingRegion id="49B1F860FFA2D679FCC4F9BDC701294E" box="[879,1076,1628,1654]" country="China" name="Yunnan" pageId="12" pageNumber="356">Yunnan Province</collectingRegion>
,
<collectingMunicipality id="6BAEACF8FFA2D679FBE8F9BDC628294E" box="[1091,1309,1628,1654]" pageId="12" pageNumber="356">Nujiang Prefecture</collectingMunicipality>
,
<collectingCounty id="62AB4E0EFFA2D679FA80F9BDC3DE29A2" pageId="12" pageNumber="356">Gongshan County</collectingCounty>
,
<location id="8EAA6059FFA2D679FF51F961C2E029A2" LSID="urn:lsid:plazi:treatment:03DC8794FFA2D678FF3CFAA0C25B2B0D:8EAA6059FFA2D679FF51F961C2E029A2" box="[250,469,1664,1690]" country="China" county="Gongshan County" municipality="Nujiang Prefecture" name="Dulongjiang valley" pageId="12" pageNumber="356" stateProvince="Yunnan">Dulongjiang valley</location>
, 2011, local collector leg
</materialsCitation>
.;
<materialsCitation id="3B1D3CDFFFA2D679FCBCF961C0D92986" collectingDate="2023-07-19" collectionCode="CZHQ" collectorName="Feiyi Xin" country="China" county="Gongshan County" elevation="1800" location="Dulongjiang valley" municipality="Nujiang Prefecture" pageId="12" pageNumber="356" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<specimenCount id="9D73FD0BFFA2D679FCBCF961C05329A2" box="[791,870,1664,1690]" count="1" pageId="12" pageNumber="356" type="male">1 male</specimenCount>
(
<collectionCode id="ED64AE47FFA2D679FCD2F961C0FF29A2" box="[889,970,1664,1690]" pageId="12" pageNumber="356">CZHQ</collectionCode>
),,
<collectingRegion id="49B1F860FFA2D679FC43F961C79B29A2" box="[1000,1198,1664,1690]" country="China" name="Yunnan" pageId="12" pageNumber="356">Yunnan Province</collectingRegion>
,
<collectingMunicipality id="6BAEACF8FFA2D679FB16F961C6AD29A2" box="[1213,1432,1664,1690]" pageId="12" pageNumber="356">Nujiang Prefecture</collectingMunicipality>
,
<collectingCounty id="62AB4E0EFFA2D679FF3CF945C2502986" box="[151,357,1700,1726]" pageId="12" pageNumber="356">Gongshan County</collectingCounty>
,
<location id="8EAA6059FFA2D679FEDAF945C17F2986" LSID="urn:lsid:plazi:treatment:03DC8794FFA2D678FF3CFAA0C25B2B0D:8EAA6059FFA2D679FEDAF945C17F2986" box="[369,586,1700,1726]" country="China" county="Gongshan County" municipality="Nujiang Prefecture" name="Dulongjiang valley" pageId="12" pageNumber="356" stateProvince="Yunnan">Dulongjiang valley</location>
,
<date id="FFCB1042FFA2D679FDFCF945C1D62986" box="[599,739,1700,1726]" pageId="12" pageNumber="356" value="2023-07-19">
<collectingDate id="EF8FE9AAFFA2D679FDFCF945C1D62986" box="[599,739,1700,1726]" pageId="12" pageNumber="356" value="2023-07-19">19.VII.2023</collectingDate>
</date>
,
<quantity id="4C8D9B67FFA2D679FD44F945C00A2986" box="[751,831,1700,1726]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="356" unit="m" value="1800.0">
<elevation id="0058D1B1FFA2D679FD44F945C00A2986" box="[751,831,1700,1726]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="356" unit="m" value="1800.0">1800m</elevation>
</quantity>
,
<collectorName id="26805354FFA2D679FCE7F945C08D2986" box="[844,952,1700,1726]" pageId="12" pageNumber="356">Feiyi Xin</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B1D3CDFFFA2D679FC52F945C7BD29DA" collectingDate="2015-07" collectionCode="CZHQ" collectorName="Xinxiong Deng" country="China" county="Nujiang Prefecture" elevation="2500" location="Pianma Township" municipality="Lushui" pageId="12" pageNumber="356" specimenCount="15" specimenCount-male="15" stateProvince="Yunnan" typeStatus="holotype">
<specimenCount id="9D73FD0BFFA2D679FC52F945C76B2986" box="[1017,1118,1700,1727]" count="15" pageId="12" pageNumber="356" type="male">15 males</specimenCount>
(
<collectionCode id="ED64AE47FFA2D679FBC4F945C7F42986" box="[1135,1217,1700,1726]" pageId="12" pageNumber="356">CZHQ</collectionCode>
),
<collectingRegion id="49B1F860FFA2D679FB7FF945C6AD2986" box="[1236,1432,1700,1726]" country="China" name="Yunnan" pageId="12" pageNumber="356">Yunnan Province</collectingRegion>
,
<collectingCounty id="62AB4E0EFFA2D679FF3CF929C25B29DA" box="[151,366,1736,1762]" pageId="12" pageNumber="356">Nujiang Prefecture</collectingCounty>
,
<collectingMunicipality id="6BAEACF8FFA2D679FED2F929C2F229DA" box="[377,455,1736,1762]" pageId="12" pageNumber="356">Lushui</collectingMunicipality>
city,
<location id="8EAA6059FFA2D679FDAFF929C1E529DA" LSID="urn:lsid:plazi:treatment:03DC8794FFA2D678FF3CFAA0C25B2B0D:8EAA6059FFA2D679FDAFF929C1E529DA" box="[516,720,1736,1762]" country="China" county="Nujiang Prefecture" municipality="Lushui" name="Pianma Township" pageId="12" pageNumber="356" stateProvince="Yunnan">Pianma Township</location>
,
<date id="FFCB1042FFA2D679FD71F929C07629DB" box="[730,835,1736,1763]" pageId="12" pageNumber="356" value="2015-07">
<collectingDate id="EF8FE9AAFFA2D679FD71F929C07629DB" box="[730,835,1736,1763]" pageId="12" pageNumber="356" value="2015-07">VII.2015</collectingDate>
</date>
,
<quantity id="4C8D9B67FFA2D679FCE5F929C0AA29DB" box="[846,927,1736,1763]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="12" pageNumber="356" unit="m" value="2500.0">
<elevation id="0058D1B1FFA2D679FCE5F929C0AA29DB" box="[846,927,1736,1763]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="12" pageNumber="356" unit="m" value="2500.0">2500m</elevation>
</quantity>
,
<collectorName id="26805354FFA2D679FC01F929C76D29DA" box="[938,1112,1736,1762]" pageId="12" pageNumber="356">Xinxiong Deng</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36F6509FFA2D678FF6CF90DC2B82CF1" lastPageId="13" lastPageNumber="357" pageId="12" pageNumber="356" type="description">
<paragraph id="8BCA3682FFA2D679FF6CF90DC160283D" blockId="12.[151,1437,1556,2014]" box="[199,597,1771,1798]" pageId="12" pageNumber="356">
<emphasis id="B901EA90FFA2D679FF6CF90DC160283D" bold="true" box="[199,597,1771,1798]" pageId="12" pageNumber="356">Redescription of the male (n=18).</emphasis>
</paragraph>
<paragraph id="8BCA3682FFA2D679FF6CF8F1C7B92812" blockId="12.[151,1437,1556,2014]" box="[199,1164,1808,1834]" pageId="12" pageNumber="356">
Body length:
<quantity id="4C8D9B67FFA2D679FECBF8F1C1362812" box="[352,515,1808,1834]" metricMagnitude="-2" metricUnit="m" metricValue="3.875" metricValueMax="4.81" metricValueMin="2.94" pageId="12" pageNumber="356" unit="mm" value="38.75" valueMax="48.1" valueMin="29.4">29.4–48.1 mm</quantity>
(
<bibRefCitation id="EFE44B73FFA2D679FDBBF8F1C1A22812" author="Fujita, H." box="[528,663,1808,1834]" pageId="12" pageNumber="356" refId="ref7425" refString="Fujita, H. (2010) The Lucanid Beetles of the World. Mushi-Sha, Tokyo, 472 pp., 248 pls." type="book" year="2010">Fujita 2010</bibRefCitation>
;
<bibRefCitation id="EFE44B73FFA2D679FD0AF8F1C0662812" author="Toshiyuki, T." box="[673,851,1808,1834]" pageId="12" pageNumber="356" pagination="1 - 87" refId="ref7955" refString="Toshiyuki, T. (2021) Illustrated guide to the genus Neolucanus of the world. Be-kuwa, 78, 1 - 87." type="journal article" year="2021">Toshiyuki 2021</bibRefCitation>
;
<bibRefCitation id="EFE44B73FFA2D679FCF6F8F1C0822812" author="Yi, D." box="[861,951,1808,1834]" pageId="12" pageNumber="356" refId="ref7980" refString="Yi, D. (2023) Lucanidae of the World. Design Spacetime, Seoul, 190 pp., 160 pls." type="book" year="2023">Yi 2023</bibRefCitation>
and present paper)
</paragraph>
<paragraph id="8BCA3682FFA2D679FF6CF8D5C26728AE" blockId="12.[151,1437,1556,2014]" pageId="12" pageNumber="356">
<emphasis id="B901EA90FFA2D679FF6CF8D5C2022876" bold="true" box="[199,311,1844,1870]" pageId="12" pageNumber="356">Telodont.</emphasis>
<emphasis id="B901EA90FFA2D679FEEBF8D5C2AC2876" box="[320,409,1844,1870]" italics="true" pageId="12" pageNumber="356">Habitus</emphasis>
(
<figureCitation id="134E2A07FFA2D679FE01F8D5C1162876" box="[426,547,1844,1870]" captionStart="FIGURE 1" captionStartId="3.[151,250,1908,1932]" captionTargetBox="[312,1275,185,1877]" captionTargetId="figure-17@3.[306,1281,179,1883]" captionTargetPageId="3" captionText="FIGURE 1. Male habitus of Neolucanus species. A–B, N. horizontalis Xin, Zhong &amp; Qi, new species, holotype, 36.0 mm, telodont (Lvchun, Yunnan); C–E, N. fuscus Didier, 1926; C–D, 50.9 mm, telodont (Yên Bái, Yên Bái); E, 44.8 mm, telodont (Ziyuan, Guangxi); F–H, N. brochieri Bomans &amp; Miyashita, 1997; F–G, 41.1mm, telodont (Gonshan, Yunnan); H, 41.0 mm, telodont (Gonshan, Yunnan). A, C, E, F, H, dorsal view; B, D, G, ventral view. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14735137" httpUri="https://zenodo.org/record/14735137/files/figure.png" pageId="12" pageNumber="356">Fig. 1F–H</figureCitation>
). Head and pronotum black, elytra black or aurantiaceous with black borders; obviously glossy; body with invisible pubescence in dorsal view, labium with dense setae all legs with sparse yellow setae on surface.
</paragraph>
<paragraph id="8BCA3682FFA2D678FF6CF841C21E2E79" blockId="12.[151,1437,1556,2014]" lastBlockId="13.[151,1437,151,1077]" lastPageId="13" lastPageNumber="357" pageId="12" pageNumber="356">
<emphasis id="B901EA90FFA2D679FF6CF841C2312882" box="[199,260,1952,1978]" italics="true" pageId="12" pageNumber="356">Head</emphasis>
transverse. Anterior ridge raised behind mandible and widely concave on the middle. Canthus long and wide, completely surround the eye. Clypeolabrum (Fig. 3M) tiny and transverse, slightly visible in dorsal view, fused with frons. Mandible (Fig. 3M–N) smooth from base to teeth on inner margin; in lateral view, clearly up-curved and thinnest at basal 1/3, a dorsal tooth behind apex forming an approximately arc-shaped and weakly protrudes beyond upper margin; in dorsal view, slightly curved, thickness almost regular from base to inner teeth, with 2–3 continuous small teeth. Antennal club with 3 antennomeres; antennomeres 8–9 subquadrilateral; antennomere 10 semi-elliptic.
</paragraph>
<paragraph id="8BCA3682FFA3D678FF6CFEAAC2032E95" blockId="13.[151,1437,151,1077]" pageId="13" pageNumber="357">
<emphasis id="B901EA90FFA3D678FF6CFEAAC2032E5C" box="[199,310,331,356]" italics="true" pageId="13" pageNumber="357">Pronotum</emphasis>
(Fig. 3O) transverse, markedly longer and wider than head with slightly raised edges.Anterior margin bisinuate, posterior margin dimly bisinuate. Anterior angles blunt and directed forwards, lateral angles and posterior angles obtuse.
</paragraph>
<paragraph id="8BCA3682FFA3D678FF6CFE56C7302EE9" blockId="13.[151,1437,151,1077]" box="[199,1029,439,465]" pageId="13" pageNumber="357">
<emphasis id="B901EA90FFA3D678FF6CFE56C2382EE9" box="[199,269,439,465]" italics="true" pageId="13" pageNumber="357">Elytra</emphasis>
elongate, anterior margin of elytra clearly narrower than pronotum.
</paragraph>
<paragraph id="8BCA3682FFA3D678FF6CFE3AC0072D05" blockId="13.[151,1437,151,1077]" pageId="13" pageNumber="357">
<emphasis id="B901EA90FFA3D678FF6CFE3AC3C92ECC" box="[199,252,475,500]" italics="true" pageId="13" pageNumber="357">Legs</emphasis>
. Protibia (Fig. 3P) with 4–6 conspicuous sharp and short protuberances along outer margin, apex bifurcate with branches moderately sharp at tip; inner margin with a unclear gap at base. Mesotibia and metatibia straight and outer margin with a terminal spine, without external spines.
</paragraph>
<paragraph id="8BCA3682FFA3D678FF6CFDA6C15B2C01" blockId="13.[151,1437,151,1077]" pageId="13" pageNumber="357">
<emphasis id="B901EA90FFA3D678FF6CFDA6C2D62D59" box="[199,483,583,609]" italics="true" pageId="13" pageNumber="357">Abdomen and aedeagus.</emphasis>
Abdominal tergite VIII (Fig. 5M) subsemi-circular, lateral angles poorly defined. Abdominal ventrite VIII (Fig. 5N) with smooth basal margin. Ventral plate of abdominal segment IX (Fig. 5P) slender without membranous stripe, obviously constricted at base, then widens sharply and significantly, widens gradually to about apical 1/5. Aedeagus (
<figureCitation id="134E2A07FFA3D678FDCFFD52C1CB2DF5" box="[612,766,691,717]" captionStart="FIGURE 6" captionStartId="8.[151,250,1909,1933]" captionTargetBox="[180,1408,181,1879]" captionTargetId="figure-18@8.[174,1414,181,1885]" captionTargetPageId="8" captionText="FIGURE 6. Aedeagus (not including flagellum) of Neolucanus species. A–B, E–F, I–J, N. horizontalis Xin, Zhong &amp; Qi, new species A, E, I, holotype (Lvchun, Yunnan); B, F, J, paratype (Jinping, Yunnan). C, G, K, N. fuscus Didier, 1926 (Ziyuan, Guangxi). D, H, L, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A–D, dorsal view; E–H, ventral view; I–L, lateral view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14735143" httpUri="https://zenodo.org/record/14735143/files/figure.png" pageId="13" pageNumber="357">Fig. 6D, H, L</figureCitation>
) with a significant and relatively slender pair of sclerotized dorsal plates; ventral plate at apical end of basal piece membranous. Paramere relatively narrow, with a obvious curvature at base, apex of paramere curved outward. Penis large, almost as long as paramere in ventral view. Flagellum long, apex of flagellum simple.
</paragraph>
<paragraph id="8BCA3682FFA3D678FF6CFCA3C2B82CF1" blockId="13.[151,1437,151,1077]" pageId="13" pageNumber="357">
<emphasis id="B901EA90FFA3D678FF6CFCA3C2F42C65" bold="true" box="[199,449,834,861]" pageId="13" pageNumber="357">Variation. Mesodont.</emphasis>
Mandible shorter and straighter in dorsal view, with a rounded tooth at base.
<emphasis id="B901EA90FFA3D678FAAEFCA3C6A92C64" bold="true" box="[1285,1436,834,860]" pageId="13" pageNumber="357">Amphiodont</emphasis>
and
<emphasis id="B901EA90FFA3D678FF63FC87C2052CB8" bold="true" box="[200,304,870,896]" pageId="13" pageNumber="357">priodont</emphasis>
(
<figureCitation id="134E2A07FFA3D678FEE9FC86C2832CB9" box="[322,438,871,897]" captionStart="FIGURE 2" captionStartId="4.[151,250,1771,1795]" captionTargetBox="[158,1429,186,1740]" captionTargetId="figure-18@4.[151,1436,181,1747]" captionTargetPageId="4" captionText="FIGURE 2. Male habitus of Neolucanus species. A–B, N. horizontalis Xin, Zhong &amp; Qi, new species, paratype, 33.7 mm, mesodont (Jinping, Yunnan); C–D, N. fuscus Didier, 1926, 34.7mm, priodont (Wuming, Guangxi); E–F, N. brochieri Bomans &amp; Miyashita, 1997, 31.4 mm, priodont (Gonshan, Yunnan). A, C, E, dorsal view; B, D, F, ventral view. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14735139" httpUri="https://zenodo.org/record/14735139/files/figure.png" pageId="13" pageNumber="357">Fig. 2E–F</figureCitation>
). Mandible (
<figureCitation id="134E2A07FFA3D678FDE1FC86C1872CB9" box="[586,690,871,897]" captionStart="FIGURE 4" captionStartId="6.[151,250,1582,1606]" captionTargetBox="[158,1429,188,1551]" captionTargetId="figure-18@6.[151,1436,181,1558]" captionTargetPageId="6" captionText="FIGURE 4. Male fore-head (A–B, E–F, I–J), pronotum (C, G, K), protibia and apical part of profemur (D, H, L) of Neolucanus species.A–D, N. horizontalis Xin, Zhong &amp; Qi, new species, paratype (Jinping, Yunnan); E–H, N. fuscus Didier, 1926 (Wuming, Guangxi); I–L, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, C–D, E, G–H, I, K–L, dorsal view; B, F, J, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14735141" httpUri="https://zenodo.org/record/14735141/files/figure.png" pageId="13" pageNumber="357">Fig. 4I–J</figureCitation>
) slender, straight and only slightly longer than head with unclear consecutive small teeth behind apex, with almost no branch at apex in lateral view; clypeolabrum (
<figureCitation id="134E2A07FFA3D678FB58FC6AC6742C9D" box="[1267,1345,907,933]" captionStart="FIGURE 4" captionStartId="6.[151,250,1582,1606]" captionTargetBox="[158,1429,188,1551]" captionTargetId="figure-18@6.[151,1436,181,1558]" captionTargetPageId="6" captionText="FIGURE 4. Male fore-head (A–B, E–F, I–J), pronotum (C, G, K), protibia and apical part of profemur (D, H, L) of Neolucanus species.A–D, N. horizontalis Xin, Zhong &amp; Qi, new species, paratype (Jinping, Yunnan); E–H, N. fuscus Didier, 1926 (Wuming, Guangxi); I–L, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, C–D, E, G–H, I, K–L, dorsal view; B, F, J, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14735141" httpUri="https://zenodo.org/record/14735141/files/figure.png" pageId="13" pageNumber="357">Fig. 4I</figureCitation>
) clearly visible in dorsal view.
</paragraph>
</subSubSection>
<subSubSection id="C36F6509FFA3D678FF6CFC33C1972CD5" box="[199,674,978,1005]" pageId="13" pageNumber="357" type="diagnosis">
<paragraph id="8BCA3682FFA3D678FF6CFC33C1972CD5" blockId="13.[151,1437,151,1077]" box="[199,674,978,1005]" pageId="13" pageNumber="357">
<emphasis id="B901EA90FFA3D678FF6CFC33C2FB2CD4" bold="true" box="[199,462,978,1004]" pageId="13" pageNumber="357">Differential diagnosis.</emphasis>
See
<tableCitation id="C6F70339FFA3D678FDA8FC32C1632CD5" box="[515,598,979,1005]" captionStart="TABLE 1" captionStartId="14.[151,239,728,752]" captionText="TABLE 1. Male main differences of external characters of Neolucanus horizontalis Xin, Zhong &amp; Qi, new species and its related species." pageId="13" pageNumber="357">Table 1</tableCitation>
and
<tableCitation id="C6F70339FFA3D678FD26FC32C1AB2CD5" box="[653,670,979,1005]" captionStart="TABLE 2" captionStartId="15.[151,239,151,175]" captionText="TABLE 2. Main differences of male abdomen and aedeagus of Neolucanus horizontalis Xin, Zhong &amp; Qi, new species and its related species." pageId="13" pageNumber="357">2</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F6509FFA3D678FF6CFC17C25B2B0D" pageId="13" pageNumber="357" type="distribution">
<paragraph id="8BCA3682FFA3D678FF6CFC17C25B2B0D" blockId="13.[151,1437,151,1077]" pageId="13" pageNumber="357">
<emphasis id="B901EA90FFA3D678FF6CFC17C2542B28" bold="true" box="[199,353,1014,1040]" pageId="13" pageNumber="357">Distribution.</emphasis>
<collectingCountry id="F3627612FFA3D678FEC3FC16C29E2B29" box="[360,427,1015,1041]" name="China" pageId="13" pageNumber="357">China</collectingCountry>
(
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<bibRefCitation id="EFE44B73FFA3D678FCA5FC16C0A02B29" author="Fujita, H." box="[782,917,1015,1041]" pageId="13" pageNumber="357" refId="ref7425" refString="Fujita, H. (2010) The Lucanid Beetles of the World. Mushi-Sha, Tokyo, 472 pp., 248 pls." type="book" year="2010">Fujita 2010</bibRefCitation>
;
<bibRefCitation id="EFE44B73FFA3D678FC0BFC16C7012B29" author="Schenk, K. D." box="[928,1076,1015,1041]" pageId="13" pageNumber="357" pagination="4 - 38" refId="ref7844" refString="Schenk, K. D. (2012) Notes to the Lucanidae of Asia and description of new taxa of the genus Neolucanus. Beetles World, 6, 4 - 38." type="journal article" year="2012">Schenk 2012</bibRefCitation>
,
<bibRefCitation id="EFE44B73FFA3D678FBEBFC16C7492B29" author="Schenk, K. D." box="[1088,1148,1015,1041]" pageId="13" pageNumber="357" pagination="2 - 8" refId="ref7878" refString="Schenk, K. D. (2014) Catalogue of Lucanidae: Genus Neolucanus Thomson, 1862. Beetles World, 10, 2 - 8." type="journal article" year="2014">2014</bibRefCitation>
;
<bibRefCitation id="EFE44B73FFA3D678FB2DFC16C60D2B29" author="Toshiyuki, T." box="[1158,1336,1015,1041]" pageId="13" pageNumber="357" pagination="1 - 87" refId="ref7955" refString="Toshiyuki, T. (2021) Illustrated guide to the genus Neolucanus of the world. Be-kuwa, 78, 1 - 87." type="journal article" year="2021">Toshiyuki 2021</bibRefCitation>
;
<bibRefCitation id="EFE44B73FFA3D678FAE8FC16C6A92B29" author="Yi, D." box="[1347,1436,1015,1041]" pageId="13" pageNumber="357" refId="ref7980" refString="Yi, D. (2023) Lucanidae of the World. Design Spacetime, Seoul, 190 pp., 160 pls." type="book" year="2023">Yi 2023</bibRefCitation>
and present paper).
</paragraph>
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627
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Chinese common name:
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<collectingDate id="EF8FE9AAFFACD677FCECFE8EC08E2EB1" box="[839,955,367,393]" pageId="2" pageNumber="346" value="2024-06-08">8.VI.2024</collectingDate>
</date>
,
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<elevation id="0058D1B1FFACD677FC60FE8EC72E2EB1" box="[971,1051,367,393]" metricMagnitude="3" metricUnit="m" metricValue="1.316" pageId="2" pageNumber="346" unit="m" value="1316.0">1316m</elevation>
</quantity>
, local collector leg
</materialsCitation>
.
<materialsCitation id="3B1D3CDFFFACD677FAA4FE8EC04D2EE9" collectingDate="2023-06-02" collectionCode="CXTZ" country="China" county="Honghe Prefecture" elevation="2400" latitude="22.756666" location="Ma'andi Township" longLatPrecision="21" longitude="103.43917" municipality="Jinping county" pageId="2" pageNumber="346" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<emphasis id="B901EA90FFACD677FAA4FE8EC6B62EB1" bold="true" box="[1295,1411,367,393]" pageId="2" pageNumber="346">
<typeStatus id="54CE8820FFACD677FAA4FE8EC64B2EB1" box="[1295,1406,367,393]" pageId="2" pageNumber="346" type="paratype">Paratype</typeStatus>
,
</emphasis>
<specimenCount id="9D73FD0BFFACD677FA25FE8EC3F82E95" count="1" pageId="2" pageNumber="346" type="male">1 male</specimenCount>
(
<collectionCode id="ED64AE47FFACD677FF74FE72C21B2E95" box="[223,302,403,429]" pageId="2" pageNumber="346">CXTZ</collectionCode>
),
<collectingCountry id="F3627612FFACD677FEE8FE72C2BC2E95" box="[323,393,403,429]" name="China" pageId="2" pageNumber="346">China</collectingCountry>
,
<collectingRegion id="49B1F860FFACD677FE3DFE75C16E2E95" box="[406,603,403,430]" country="China" name="Yunnan" pageId="2" pageNumber="346">Yunnan Province</collectingRegion>
,
<collectingCounty id="62AB4E0EFFACD677FDC2FE72C0742E95" box="[617,833,403,429]" pageId="2" pageNumber="346">Honghe Prefecture</collectingCounty>
,
<collectingMunicipality id="6BAEACF8FFACD677FCE5FE72C0C22E96" box="[846,1015,403,430]" pageId="2" pageNumber="346">Jinping county</collectingMunicipality>
,
<location id="8EAA6059FFACD677FBAEFE72C7EB2E95" LSID="urn:lsid:plazi:treatment:03DC8794FFACD672FF3CFF76C62A28B3:8EAA6059FFACD677FBAEFE72C7EB2E95" box="[1029,1246,403,429]" country="China" county="Honghe Prefecture" latitude="22.756666" longLatPrecision="21" longitude="103.43917" municipality="Jinping county" name="Ma'andi Township" pageId="2" pageNumber="346" stateProvince="Yunnan">Ma’andi Township</location>
,
<location id="8EAA6059FFACD677FB47FE72C6AD2E95" LSID="urn:lsid:plazi:treatment:03DC8794FFACD672FF3CFF76C62A28B3:8EAA6059FFACD677FB47FE72C6AD2E95" box="[1260,1432,403,429]" country="China" county="Honghe Prefecture" latitude="22.756666" longLatPrecision="21" longitude="103.43917" municipality="Jinping county" name="Sitai Mountain" pageId="2" pageNumber="346" stateProvince="Yunnan">Sitai Mountain</location>
,
<geoCoordinate id="EE415045FFACD677FF3CFE56C2162EEA" box="[151,291,439,466]" degrees="22" direction="north" minutes="45" orientation="latitude" pageId="2" pageNumber="346" precision="15" seconds="24" value="22.756666">22°45’24”N</geoCoordinate>
,
<geoCoordinate id="EE415045FFACD677FE85FE56C2F02EE9" box="[302,453,439,465]" degrees="103" direction="east" minutes="26" orientation="longitude" pageId="2" pageNumber="346" precision="15" seconds="21" value="103.43917">103°26’21”E</geoCoordinate>
,
<quantity id="4C8D9B67FFACD677FE7BFE56C1142EE9" box="[464,545,439,465]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="346" unit="m" value="2400.0">
<elevation id="0058D1B1FFACD677FE7BFE56C1142EE9" box="[464,545,439,465]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="346" unit="m" value="2400.0">2400m</elevation>
</quantity>
,
<date id="FFCB1042FFACD677FD87FE56C1952EE9" box="[556,672,439,465]" pageId="2" pageNumber="346" value="2023-06-02">
<collectingDate id="EF8FE9AAFFACD677FD87FE56C1952EE9" box="[556,672,439,465]" pageId="2" pageNumber="346" value="2023-06-02">2.VI.2023</collectingDate>
</date>
, local collector leg
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F6509FFACD677FF6CFE3AC29729A1" pageId="2" pageNumber="346" type="description">
<paragraph id="8BCA3682FFACD677FF6CFE3AC16D2ECD" blockId="2.[151,1437,295,1905]" box="[199,600,475,501]" pageId="2" pageNumber="346">
<emphasis id="B901EA90FFACD677FF6CFE3AC16D2ECD" bold="true" box="[199,600,475,501]" pageId="2" pageNumber="346">
Description of the
<typeStatus id="54CE8820FFACD677FE08FE3AC13E2ECD" box="[419,523,475,501]" pageId="2" pageNumber="346" type="holotype">holotype</typeStatus>
, male.
</emphasis>
</paragraph>
<paragraph id="8BCA3682FFACD677FF6CFDE1C63B2D05" blockId="2.[151,1437,295,1905]" pageId="2" pageNumber="346">
<emphasis id="B901EA90FFACD677FF6CFDE1C2442D21" box="[199,369,512,537]" italics="true" pageId="2" pageNumber="346">Measurements.</emphasis>
Body length (36.0 mm). Length of particular body parts: head (
<quantity id="4C8D9B67FFACD677FBE1FE1EC7922D21" box="[1098,1191,511,537]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="2" pageNumber="346" unit="mm" value="3.8">3.8 mm</quantity>
), mandible (
<quantity id="4C8D9B67FFACD677FA9FFE1EC6A52D21" box="[1332,1424,511,537]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="2" pageNumber="346" unit="mm" value="8.2">8.2 mm</quantity>
), pronotum (7.0 mm), elytra (
<quantity id="4C8D9B67FFACD677FE7BFDC2C10C2D05" box="[464,569,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="1.63" pageId="2" pageNumber="346" unit="mm" value="16.3">16.3 mm</quantity>
). Width: head (
<quantity id="4C8D9B67FFACD677FD4FFDC2C00A2D05" box="[740,831,547,573]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="2" pageNumber="346" unit="mm" value="9.1">9.1 mm</quantity>
), pronotum (
<quantity id="4C8D9B67FFACD677FC65FDC2C7022D05" box="[974,1079,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="2" pageNumber="346" unit="mm" value="12.3">12.3 mm</quantity>
), elytra (
<quantity id="4C8D9B67FFACD677FB33FDC2C6372D05" box="[1176,1282,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="2" pageNumber="346" unit="mm" value="12.3">12.3 mm</quantity>
).
</paragraph>
<paragraph id="8BCA3682FFACD677FF6CFDA6C0992DBD" blockId="2.[151,1437,295,1905]" pageId="2" pageNumber="346">
<emphasis id="B901EA90FFACD677FF6CFDA6C2152D59" box="[199,288,583,609]" italics="true" pageId="2" pageNumber="346">Habitus</emphasis>
(
<figureCitation id="134E2A07FFACD677FE84FDA6C29F2D59" box="[303,426,583,609]" captionStart="FIGURE 1" captionStartId="3.[151,250,1908,1932]" captionTargetBox="[312,1275,185,1877]" captionTargetId="figure-17@3.[306,1281,179,1883]" captionTargetPageId="3" captionText="FIGURE 1. Male habitus of Neolucanus species. A–B, N. horizontalis Xin, Zhong &amp; Qi, new species, holotype, 36.0 mm, telodont (Lvchun, Yunnan); C–E, N. fuscus Didier, 1926; C–D, 50.9 mm, telodont (Yên Bái, Yên Bái); E, 44.8 mm, telodont (Ziyuan, Guangxi); F–H, N. brochieri Bomans &amp; Miyashita, 1997; F–G, 41.1mm, telodont (Gonshan, Yunnan); H, 41.0 mm, telodont (Gonshan, Yunnan). A, C, E, F, H, dorsal view; B, D, G, ventral view. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14735137" httpUri="https://zenodo.org/record/14735137/files/figure.png" pageId="2" pageNumber="346">Fig. 1A–B</figureCitation>
). Head and pronotum black, elytra kermesinus, with slightly glossy; whole body with tiny and almost invisible pubescence, all legs with yellow setae on surface.
</paragraph>
<paragraph id="8BCA3682FFACD677FF6CFD6EC6572CB9" blockId="2.[151,1437,295,1905]" pageId="2" pageNumber="346">
<emphasis id="B901EA90FFACD677FF6CFD6EC2312D91" box="[199,260,655,681]" italics="true" pageId="2" pageNumber="346">Head</emphasis>
transverse. Anterior ridge raised behind mandible and widely concave on the middle. Canthus long and wide, completely surround the eye. Clypeolabrum (Fig. 3E) tiny and subtriangle, visible in dorsal view, fused with frons, apex sharp. Mandible (Fig. 3E–F) about 2.2 times longer than head, inner smooth from base to teeth; in lateral view, almost horizontal, a dorsal tooth behind apex forming an approximately vertical right-angle and slightly protrudes beyond upper margin; in dorsal view, slightly curved, base thick, becomes thicker firstly and then thinner from base to teeth, with 2 continuous subtriangle inner teeth behind apex, proximal one smaller, apex of mandible sharp. Antennal club with 3 antennomeres; antennomeres 8–9 subquadrilateral; antennomere 10 semi-elliptic.
</paragraph>
<paragraph id="8BCA3682FFACD677FF6CFC6DC02B2CD5" blockId="2.[151,1437,295,1905]" pageId="2" pageNumber="346">
<emphasis id="B901EA90FFACD677FF6CFC6DC2032C9D" box="[199,310,908,933]" italics="true" pageId="2" pageNumber="346">Pronotum</emphasis>
(Fig. 3G) transverse, 1.8 times wider than long, markedly longer and wider than head with slightly raised edges. Anterior margin bisinuate, posterior margin slightly bisinuate. Anterior angles rounded and directed forwards, lateral angles obtuse and posterior angles blunt.
</paragraph>
<paragraph id="8BCA3682FFACD677FF6CFC16C0932B29" blockId="2.[151,1437,295,1905]" box="[199,934,1015,1042]" pageId="2" pageNumber="346">
<taxonomicName id="4C754D01FFACD677FF6CFC16C2002B29" box="[199,309,1015,1041]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="2" pageNumber="346" phylum="Arthropoda" rank="genus">
<emphasis id="B901EA90FFACD677FF6CFC16C2002B29" box="[199,309,1015,1041]" italics="true" pageId="2" pageNumber="346">Scutellum</emphasis>
</taxonomicName>
subtriangular, punctate, width almost as long as length.
</paragraph>
<paragraph id="8BCA3682FFACD677FF6CFBFDC3EF2B45" blockId="2.[151,1437,295,1905]" pageId="2" pageNumber="346">
<emphasis id="B901EA90FFACD677FF6CFBFDC3C92B0D" box="[199,252,1052,1077]" italics="true" pageId="2" pageNumber="346">Legs</emphasis>
. Protibia (Fig. 3H) with 2 conspicuous sharp protuberances along outer margin, apex bifurcate with branches moderately sharp at tip. Mesotibia and metatibia straight and smooth at outer margin with a terminal spine.
</paragraph>
<paragraph id="8BCA3682FFACD677FF6CFB66C62D2B9A" blockId="2.[151,1437,295,1905]" box="[199,1304,1159,1186]" pageId="2" pageNumber="346">
<emphasis id="B901EA90FFACD677FF6CFB66C2382B99" box="[199,269,1159,1185]" italics="true" pageId="2" pageNumber="346">Elytra</emphasis>
elongate, 1.3 times longer than wide, anterior margin significantly narrower than pronotum.
</paragraph>
<paragraph id="8BCA3682FFACD677FF6CFB4AC11E2ADD" blockId="2.[151,1437,295,1905]" pageId="2" pageNumber="346">
<emphasis id="B901EA90FFACD677FF6CFB4AC2D12BFD" box="[199,484,1195,1221]" italics="true" pageId="2" pageNumber="346">Abdomen and aedeagus.</emphasis>
Abdominal tergite VIII (Fig. 5A) nearly semilune, lateral angles poorly defined. Abdominal ventrite VIII (Fig. 5B) with 2 slightly protruding tips on basal margin near middle. Ventral plate of abdominal segment IX (Fig. 5D) elongate without membranous stripe, basal part extremely narrow, then suddenly became wider, widens gradually to apex and constricted near apex. Aedeagus (
<figureCitation id="134E2A07FFACD677FBB8FAF6C7912A09" box="[1043,1188,1303,1330]" captionStart="FIGURE 6" captionStartId="8.[151,250,1909,1933]" captionTargetBox="[180,1408,181,1879]" captionTargetId="figure-18@8.[174,1414,181,1885]" captionTargetPageId="8" captionText="FIGURE 6. Aedeagus (not including flagellum) of Neolucanus species. A–B, E–F, I–J, N. horizontalis Xin, Zhong &amp; Qi, new species A, E, I, holotype (Lvchun, Yunnan); B, F, J, paratype (Jinping, Yunnan). C, G, K, N. fuscus Didier, 1926 (Ziyuan, Guangxi). D, H, L, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A–D, dorsal view; E–H, ventral view; I–L, lateral view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14735143" httpUri="https://zenodo.org/record/14735143/files/figure.png" pageId="2" pageNumber="346">Fig. 6A, E, I</figureCitation>
) in dorsal view about 3.2 times longer than wide. Basal piece elongated, weakly longer than parameres in dorsal view, with a significant and relatively slender pair of sclerotized dorsal plates, and almost as wide as paramere; ventral plate at apical end of basal piece membranous. Paramere relatively narrow, with a small inner curvature at base, apex of paramere curved outward. Penis about 0.9 times length of paramere in ventral view. Flagellum long, about 1.6 times longer than aedeagus, apex of flagellum simple.
</paragraph>
<paragraph id="8BCA3682FFACD677FF6CFA0EC21B294D" blockId="2.[151,1437,295,1905]" pageId="2" pageNumber="346">
<emphasis id="B901EA90FFACD677FF6CFA0EC1912931" bold="true" box="[199,676,1519,1545]" pageId="2" pageNumber="346">
Variation of
<typeStatus id="54CE8820FFACD677FECEFA0EC2E52931" box="[357,464,1519,1545]" pageId="2" pageNumber="346" type="paratype">paratype</typeStatus>
(n=1). Mesodont
</emphasis>
(
<figureCitation id="134E2A07FFACD677FD1CFA0EC0012931" box="[695,820,1519,1545]" captionStart="FIGURE 2" captionStartId="4.[151,250,1771,1795]" captionTargetBox="[158,1429,186,1740]" captionTargetId="figure-18@4.[151,1436,181,1747]" captionTargetPageId="4" captionText="FIGURE 2. Male habitus of Neolucanus species. A–B, N. horizontalis Xin, Zhong &amp; Qi, new species, paratype, 33.7 mm, mesodont (Jinping, Yunnan); C–D, N. fuscus Didier, 1926, 34.7mm, priodont (Wuming, Guangxi); E–F, N. brochieri Bomans &amp; Miyashita, 1997, 31.4 mm, priodont (Gonshan, Yunnan). A, C, E, dorsal view; B, D, F, ventral view. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14735139" httpUri="https://zenodo.org/record/14735139/files/figure.png" pageId="2" pageNumber="346">Fig. 2A–B</figureCitation>
). Mandible 1.8 times longer than head (
<figureCitation id="134E2A07FFACD677FABAFA0EC6BA2931" box="[1297,1423,1519,1545]" captionStart="FIGURE 4" captionStartId="6.[151,250,1582,1606]" captionTargetBox="[158,1429,188,1551]" captionTargetId="figure-18@6.[151,1436,181,1558]" captionTargetPageId="6" captionText="FIGURE 4. Male fore-head (A–B, E–F, I–J), pronotum (C, G, K), protibia and apical part of profemur (D, H, L) of Neolucanus species.A–D, N. horizontalis Xin, Zhong &amp; Qi, new species, paratype (Jinping, Yunnan); E–H, N. fuscus Didier, 1926 (Wuming, Guangxi); I–L, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, C–D, E, G–H, I, K–L, dorsal view; B, F, J, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14735141" httpUri="https://zenodo.org/record/14735141/files/figure.png" pageId="2" pageNumber="346">Fig. 4A–B</figureCitation>
), shorter and straighter in dorsal view, dorsal tooth almost not protrudes beyond upper margin in lateral view, with 3 indistinct smaller inner teeth and a large blunt tooth at base. Protibia (
<figureCitation id="134E2A07FFACD677FC14F9D6C7222969" box="[959,1047,1591,1617]" captionStart="FIGURE 4" captionStartId="6.[151,250,1582,1606]" captionTargetBox="[158,1429,188,1551]" captionTargetId="figure-18@6.[151,1436,181,1558]" captionTargetPageId="6" captionText="FIGURE 4. Male fore-head (A–B, E–F, I–J), pronotum (C, G, K), protibia and apical part of profemur (D, H, L) of Neolucanus species.A–D, N. horizontalis Xin, Zhong &amp; Qi, new species, paratype (Jinping, Yunnan); E–H, N. fuscus Didier, 1926 (Wuming, Guangxi); I–L, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, C–D, E, G–H, I, K–L, dorsal view; B, F, J, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14735141" httpUri="https://zenodo.org/record/14735141/files/figure.png" pageId="2" pageNumber="346">Fig. 4D</figureCitation>
) with 3 sharp protuberances along outer margin.
</paragraph>
<paragraph id="8BCA3682FFACD677FF6CF99EC29729A1" blockId="2.[151,1437,295,1905]" box="[199,418,1663,1689]" pageId="2" pageNumber="346">
<emphasis id="B901EA90FFACD677FF6CF99EC21029A1" bold="true" box="[199,293,1663,1689]" pageId="2" pageNumber="346">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C36F6509FFACD677FF6CF942C2E5283D" pageId="2" pageNumber="346" type="diagnosis">
<paragraph id="8BCA3682FFACD677FF6CF942C2E5283D" blockId="2.[151,1437,295,1905]" pageId="2" pageNumber="346">
<emphasis id="B901EA90FFACD677FF6CF942C2F82985" bold="true" box="[199,461,1699,1725]" pageId="2" pageNumber="346">Differential diagnosis.</emphasis>
The new species is similar to
<taxonomicName id="4C754D01FFACD677FCBCF945C7242985" authority="Didier, 1926" authorityName="Didier" authorityYear="1926" box="[791,1041,1699,1726]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="346" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="B901EA90FFACD677FCBCF945C01F2985" box="[791,810,1700,1725]" italics="true" pageId="2" pageNumber="346">N</emphasis>
.
<emphasis id="B901EA90FFACD677FC9CF942C0482985" box="[823,893,1699,1725]" italics="true" pageId="2" pageNumber="346">fuscus</emphasis>
<bibRefCitation id="EFE44B73FFACD677FC28F942C7242985" author="Didier, R." box="[899,1041,1699,1725]" pageId="2" pageNumber="346" pagination="210 - 214" refId="ref7377" refString="Didier, R. (1926) Descriptions sommaires de Lucanides nouveaux, du genre Neolucanus, de la faune indo-chinoise. Bulletin de la Societe Entomologique de France, 1926, 210 - 214. https: // doi. org / 10.3406 / bsef. 1926.27736" type="journal article" year="1926">Didier, 1926</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C754D01FFACD677FBB0F945C3E729D9" authority="Bomans &amp; Miyashita, 1997" authorityName="Bomans &amp; Miyashita" authorityYear="1997" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="346" phylum="Arthropoda" rank="species" species="brochieri">
<emphasis id="B901EA90FFACD677FBB0F945C71B2985" box="[1051,1070,1700,1725]" italics="true" pageId="2" pageNumber="346">N</emphasis>
.
<emphasis id="B901EA90FFACD677FB90F942C7972985" box="[1083,1186,1699,1725]" italics="true" pageId="2" pageNumber="346">brochieri</emphasis>
<bibRefCitation id="EFE44B73FFACD677FB03F945C3E729D9" author="Bomans, H. E. &amp; Miyashita, T." pageId="2" pageNumber="346" pagination="2 - 3" refId="ref7344" refString="Bomans, H. E. &amp; Miyashita, T. (1997) Description de trois nouvelles especes de Lucanidae du nord Birmanie. Besoiro, 4, 2 - 3." type="journal article" year="1997">Bomans &amp; Miyashita, 1997</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C754D01FFACD677FF4BF929C2CD29DA" authority="Baba, 1995" authorityName="Baba" authorityYear="1995" box="[224,504,1735,1762]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="346" phylum="Arthropoda" rank="species" species="lehmanni">
<emphasis id="B901EA90FFACD677FF4BF929C3C629D9" box="[224,243,1736,1761]" italics="true" pageId="2" pageNumber="346">N</emphasis>
.
<emphasis id="B901EA90FFACD677FEAFF926C25929D9" box="[260,364,1735,1761]" italics="true" pageId="2" pageNumber="346">lehmanni</emphasis>
<bibRefCitation id="EFE44B73FFACD677FEDDF926C2CD29DA" author="Baba, M." box="[374,504,1735,1762]" pageId="2" pageNumber="346" pagination="8 - 11" refId="ref7312" refString="Baba, M. (1995) A new species of the genus Neolucanus Tomson, 1862 from south Burma (Myanmar). Gekkan-Mushi, 292, 8 - 11." type="journal article" year="1995">
Baba,
<quantity id="4C8D9B67FFACD677FE6BF926C2CD29DA" box="[448,504,1735,1762]" metricMagnitude="1" metricUnit="m" metricValue="5.0673" pageId="2" pageNumber="346" unit="in" value="1995.0">1995</quantity>
</bibRefCitation>
</taxonomicName>
in general appearance, but the following set of characters could help to identify precisely (
<tableCitation id="C6F70339FFACD677FEA0F90AC26A283D" box="[267,351,1771,1797]" captionStart="TABLE 1" captionStartId="14.[151,239,728,752]" captionText="TABLE 1. Male main differences of external characters of Neolucanus horizontalis Xin, Zhong &amp; Qi, new species and its related species." pageId="2" pageNumber="346">Table 1</tableCitation>
,
<tableCitation id="C6F70339FFACD677FEC6F90AC2F6283D" box="[365,451,1771,1797]" captionStart="TABLE 2" captionStartId="15.[151,239,151,175]" captionText="TABLE 2. Main differences of male abdomen and aedeagus of Neolucanus horizontalis Xin, Zhong &amp; Qi, new species and its related species." pageId="2" pageNumber="346">Table 2</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36F6509FFACD677FF6CF8EEC2082875" pageId="2" pageNumber="346" type="etymology">
<paragraph id="8BCA3682FFACD677FF6CF8EEC2082875" blockId="2.[151,1437,295,1905]" pageId="2" pageNumber="346">
<emphasis id="B901EA90FFACD677FF6CF8EEC27B2811" bold="true" box="[199,334,1807,1833]" pageId="2" pageNumber="346">Etymology.</emphasis>
This new species is named after the morphological features of its horizontal mandible when viewed in lateral view.
</paragraph>
</subSubSection>
<subSubSection id="C36F6509FFACD670FF6CF8B6C74D28C1" lastPageId="5" lastPageNumber="349" pageId="2" pageNumber="346" type="distribution">
<paragraph id="8BCA3682FFACD677FF6CF8B6C1142849" blockId="2.[151,1437,295,1905]" box="[199,545,1879,1905]" pageId="2" pageNumber="346">
<emphasis id="B901EA90FFACD677FF6CF8B6C2542849" bold="true" box="[199,353,1879,1905]" pageId="2" pageNumber="346">Distribution.</emphasis>
<collectingCountry id="F3627612FFACD677FECCF8B6C29F2849" box="[359,426,1879,1905]" name="China" pageId="2" pageNumber="346">China</collectingCountry>
(
<collectingRegion id="49B1F860FFACD677FE12F8B6C1202849" box="[441,533,1879,1905]" country="China" name="Yunnan" pageId="2" pageNumber="346">Yunnan</collectingRegion>
).
</paragraph>
<caption id="DF0A660AFFADD676FF3CF895C7AC28C0" ID-DOI="http://doi.org/10.5281/zenodo.14735137" ID-Zenodo-Dep="14735137" httpUri="https://zenodo.org/record/14735137/files/figure.png" pageId="3" pageNumber="347" startId="3.[151,250,1908,1932]" targetBox="[312,1275,185,1877]" targetPageId="3" targetType="figure">
<paragraph id="8BCA3682FFADD676FF3CF895C7AC28C0" blockId="3.[151,1436,1908,2041]" pageId="3" pageNumber="347">
<emphasis id="B901EA90FFADD676FF3CF895C22028B4" bold="true" box="[151,277,1908,1932]" pageId="3" pageNumber="347">FIGURE 1.</emphasis>
Male habitus of
<taxonomicName id="4C754D01FFADD676FE62F894C18128B4" box="[457,692,1908,1933]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="347" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="B901EA90FFADD676FE62F894C17428B5" box="[457,577,1909,1933]" italics="true" pageId="3" pageNumber="347">Neolucanus</emphasis>
species.
<emphasis id="B901EA90FFADD676FD0AF895C18128B4" bold="true" box="[673,692,1908,1932]" pageId="3" pageNumber="347">A</emphasis>
</taxonomicName>
<emphasis id="B901EA90FFADD676FD1FF895C1E228B4" bold="true" box="[692,727,1908,1932]" pageId="3" pageNumber="347">–B,</emphasis>
<taxonomicName id="4C754D01FFADD676FD4BF894C70C28B5" authority="Xin, Zhong &amp; Qi" authorityName="Xin, Zhong &amp; Qi" authorityYear="2025" box="[736,1081,1909,1933]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="347" phylum="Arthropoda" rank="species" species="horizontalis" status="new species">
<emphasis id="B901EA90FFADD676FD4BF894C1C428B4" box="[736,753,1909,1932]" italics="true" pageId="3" pageNumber="347">N</emphasis>
.
<emphasis id="B901EA90FFADD676FCABF894C04C28B5" box="[768,889,1909,1933]" italics="true" pageId="3" pageNumber="347">horizontalis</emphasis>
Xin, Zhong &amp; Qi
</taxonomicName>
,
<emphasis id="B901EA90FFADD676FBEEF894C7F128B4" bold="true" box="[1093,1220,1908,1933]" pageId="3" pageNumber="347">
<taxonomicNameLabel id="A23257EBFFADD676FBEEF894C7F128B4" box="[1093,1220,1908,1933]" pageId="3" pageNumber="347" rank="species">new species</taxonomicNameLabel>
</emphasis>
, holotype, 36.0 mm, telodont (Lvchun, Yunnan);
<emphasis id="B901EA90FFADD676FE13F879C2DA2888" bold="true" box="[440,495,1944,1968]" pageId="3" pageNumber="347">C–E,</emphasis>
<taxonomicName id="4C754D01FFADD676FE5DF878C1EA2889" authority="Didier, 1926" authorityName="Didier" authorityYear="1926" box="[502,735,1945,1969]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="347" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="B901EA90FFADD676FE5DF878C1322888" box="[502,519,1945,1968]" italics="true" pageId="3" pageNumber="347">N</emphasis>
.
<emphasis id="B901EA90FFADD676FDBEF878C1612889" box="[533,596,1945,1969]" italics="true" pageId="3" pageNumber="347">fuscus</emphasis>
<bibRefCitation id="EFE44B73FFADD676FDF0F878C1EA2889" author="Didier, R." box="[603,735,1945,1969]" pageId="3" pageNumber="347" pagination="210 - 214" refId="ref7377" refString="Didier, R. (1926) Descriptions sommaires de Lucanides nouveaux, du genre Neolucanus, de la faune indo-chinoise. Bulletin de la Societe Entomologique de France, 1926, 210 - 214. https: // doi. org / 10.3406 / bsef. 1926.27736" type="journal article" year="1926">Didier, 1926</bibRefCitation>
</taxonomicName>
;
<emphasis id="B901EA90FFADD676FD41F879C0142888" bold="true" box="[746,801,1944,1968]" pageId="3" pageNumber="347">C–D,</emphasis>
50.9 mm, telodont (Yên Bái, Yên Bái);
<emphasis id="B901EA90FFADD676FB69F878C7EC2889" bold="true" box="[1218,1241,1945,1969]" pageId="3" pageNumber="347">E,</emphasis>
44.8 mm, telodont (Ziyuan, Guangxi);
<emphasis id="B901EA90FFADD676FEC8F85CC2AC28ED" bold="true" box="[355,409,1981,2005]" pageId="3" pageNumber="347">F–H,</emphasis>
<taxonomicName id="4C754D01FFADD676FE0AF85CC07728ED" authority="Bomans &amp; Miyashita, 1997" authorityName="Bomans &amp; Miyashita" authorityYear="1997" box="[417,834,1981,2005]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="347" phylum="Arthropoda" rank="species" species="brochieri">
<emphasis id="B901EA90FFADD676FE0AF85CC28728EC" box="[417,434,1981,2004]" italics="true" pageId="3" pageNumber="347">N</emphasis>
.
<emphasis id="B901EA90FFADD676FE14F85CC12928ED" box="[447,540,1981,2005]" italics="true" pageId="3" pageNumber="347">brochieri</emphasis>
<bibRefCitation id="EFE44B73FFADD676FD8FF85CC07728ED" author="Bomans, H. E. &amp; Miyashita, T." box="[548,834,1981,2005]" pageId="3" pageNumber="347" pagination="2 - 3" refId="ref7344" refString="Bomans, H. E. &amp; Miyashita, T. (1997) Description de trois nouvelles especes de Lucanidae du nord Birmanie. Besoiro, 4, 2 - 3." type="journal article" year="1997">Bomans &amp; Miyashita, 1997</bibRefCitation>
</taxonomicName>
;
<emphasis id="B901EA90FFADD676FCE6F85DC0B628EC" bold="true" box="[845,899,1980,2004]" pageId="3" pageNumber="347">F–G,</emphasis>
41.1mm, telodont (Gonshan, Yunnan);
<emphasis id="B901EA90FFADD676FAB1F85CC60628ED" bold="true" box="[1306,1331,1981,2005]" pageId="3" pageNumber="347">H,</emphasis>
41.0 mm, telodont (Gonshan, Yunnan).
<emphasis id="B901EA90FFADD676FE6AF801C16528C1" bold="true" box="[449,592,2016,2041]" pageId="3" pageNumber="347">A, C, E, F, H,</emphasis>
dorsal view;
<emphasis id="B901EA90FFADD676FD71F800C00528C0" bold="true" box="[730,816,2016,2041]" pageId="3" pageNumber="347">B, D, G,</emphasis>
ventral view. Scale bar = 10.0 mm.
</paragraph>
</caption>
<caption id="DF0A660AFFAAD671FF3CF90AC6412872" ID-DOI="http://doi.org/10.5281/zenodo.14735139" ID-Zenodo-Dep="14735139" httpUri="https://zenodo.org/record/14735139/files/figure.png" pageId="4" pageNumber="348" startId="4.[151,250,1771,1795]" targetBox="[158,1429,186,1740]" targetPageId="4" targetType="figure">
<paragraph id="8BCA3682FFAAD671FF3CF90AC6412872" blockId="4.[151,1437,1771,1867]" pageId="4" pageNumber="348">
<emphasis id="B901EA90FFAAD671FF3CF90AC220283B" bold="true" box="[151,277,1771,1795]" pageId="4" pageNumber="348">FIGURE 2.</emphasis>
Male habitus of
<taxonomicName id="4C754D01FFAAD671FE61F90AC180283B" box="[458,693,1771,1795]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="348" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="B901EA90FFAAD671FE61F90AC177283B" box="[458,578,1771,1795]" italics="true" pageId="4" pageNumber="348">Neolucanus</emphasis>
species.
<emphasis id="B901EA90FFAAD671FD09F90AC180283B" bold="true" box="[674,693,1771,1795]" pageId="4" pageNumber="348">A</emphasis>
</taxonomicName>
<emphasis id="B901EA90FFAAD671FD1EF90AC1ED283B" bold="true" box="[693,728,1771,1795]" pageId="4" pageNumber="348">–B,</emphasis>
<taxonomicName id="4C754D01FFAAD671FD4AF90DC70F283B" authority="Xin, Zhong &amp; Qi" authorityName="Xin, Zhong &amp; Qi" authorityYear="2025" box="[737,1082,1771,1795]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="348" phylum="Arthropoda" rank="species" species="horizontalis" status="new species">
<emphasis id="B901EA90FFAAD671FD4AF90DC1C7283B" box="[737,754,1772,1795]" italics="true" pageId="4" pageNumber="348">N</emphasis>
.
<emphasis id="B901EA90FFAAD671FCABF90AC04C283B" box="[768,889,1771,1795]" italics="true" pageId="4" pageNumber="348">horizontalis</emphasis>
Xin, Zhong &amp; Qi
</taxonomicName>
,
<emphasis id="B901EA90FFAAD671FBEDF90AC7F0283B" bold="true" box="[1094,1221,1771,1795]" pageId="4" pageNumber="348">
<taxonomicNameLabel id="A23257EBFFAAD671FBEDF90AC7F0283B" box="[1094,1221,1771,1795]" pageId="4" pageNumber="348" rank="species">new species</taxonomicNameLabel>
</emphasis>
, paratype, 33.7 mm, mesodont (Jinping, Yunnan);
<emphasis id="B901EA90FFAAD671FE14F8EEC2C2281F" bold="true" box="[447,503,1807,1831]" pageId="4" pageNumber="348">C–D,</emphasis>
<taxonomicName id="4C754D01FFAAD671FE57F8F1C1EB281F" authority="Didier, 1926" authorityName="Didier" authorityYear="1926" box="[508,734,1807,1831]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="348" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="B901EA90FFAAD671FE57F8F1C138281F" box="[508,525,1808,1831]" italics="true" pageId="4" pageNumber="348">N</emphasis>
.
<emphasis id="B901EA90FFAAD671FDB2F8EEC16D281F" box="[537,600,1807,1831]" italics="true" pageId="4" pageNumber="348">fuscus</emphasis>
<bibRefCitation id="EFE44B73FFAAD671FDF5F8EEC1EB281F" author="Didier, R." box="[606,734,1807,1831]" pageId="4" pageNumber="348" pagination="210 - 214" refId="ref7377" refString="Didier, R. (1926) Descriptions sommaires de Lucanides nouveaux, du genre Neolucanus, de la faune indo-chinoise. Bulletin de la Societe Entomologique de France, 1926, 210 - 214. https: // doi. org / 10.3406 / bsef. 1926.27736" type="journal article" year="1926">Didier, 1926</bibRefCitation>
</taxonomicName>
, 34.7mm, priodont (Wuming, Guangxi);
<emphasis id="B901EA90FFAAD671FBD1F8EEC79E281F" bold="true" box="[1146,1195,1807,1831]" pageId="4" pageNumber="348">E–F,</emphasis>
<taxonomicName id="4C754D01FFAAD671FB1AF8F1C20B2873" authority="Bomans &amp; Miyashita, 1997" authorityName="Bomans &amp; Miyashita" authorityYear="1997" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="348" phylum="Arthropoda" rank="species" species="brochieri">
<emphasis id="B901EA90FFAAD671FB1AF8F1C7F7281F" box="[1201,1218,1808,1831]" italics="true" pageId="4" pageNumber="348">N</emphasis>
.
<emphasis id="B901EA90FFAAD671FB66F8EEC61F281F" box="[1229,1322,1807,1831]" italics="true" pageId="4" pageNumber="348">brochieri</emphasis>
<bibRefCitation id="EFE44B73FFAAD671FA9BF8EEC20B2873" author="Bomans, H. E. &amp; Miyashita, T." pageId="4" pageNumber="348" pagination="2 - 3" refId="ref7344" refString="Bomans, H. E. &amp; Miyashita, T. (1997) Description de trois nouvelles especes de Lucanidae du nord Birmanie. Besoiro, 4, 2 - 3." type="journal article" year="1997">Bomans &amp; Miyashita, 1997</bibRefCitation>
</taxonomicName>
, 31.4 mm, priodont (Gonshan, Yunnan).
<emphasis id="B901EA90FFAAD671FD77F8D2C0042873" bold="true" box="[732,817,1843,1867]" pageId="4" pageNumber="348">A, C, E,</emphasis>
dorsal view;
<emphasis id="B901EA90FFAAD671FC10F8D2C73E2873" bold="true" box="[955,1035,1843,1867]" pageId="4" pageNumber="348">B, D, F,</emphasis>
ventral view. Scale bar = 10.0 mm.
</paragraph>
</caption>
<paragraph id="8BCA3682FFABD670FF3CF895C74D28C1" blockId="5.[151,1437,1908,2041]" pageId="5" pageNumber="349">
<emphasis id="B901EA90FFABD670FF3CF895C22628B4" bold="true" box="[151,275,1908,1932]" pageId="5" pageNumber="349">FIGURE 3.</emphasis>
Male fore-head
<emphasis id="B901EA90FFABD670FE10F895C19228B4" bold="true" box="[443,679,1908,1933]" pageId="5" pageNumber="349">(A–B, E–F, I–J, M–N)</emphasis>
, pronotum
<emphasis id="B901EA90FFABD670FCB6F895C09728B4" bold="true" box="[797,930,1908,1933]" pageId="5" pageNumber="349">(C, G, K, O)</emphasis>
, protibia and apical part of profemur
<emphasis id="B901EA90FFABD670FAB4F895C6A928B4" bold="true" box="[1311,1436,1908,1933]" pageId="5" pageNumber="349">(D, H, L, P)</emphasis>
of
<taxonomicName id="4C754D01FFABD670FF18F878C2AF2888" box="[179,410,1944,1969]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="349" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="B901EA90FFABD670FF18F878C21E2889" box="[179,299,1945,1969]" italics="true" pageId="5" pageNumber="349">Neolucanus</emphasis>
species.
<emphasis id="B901EA90FFABD670FE23F879C2AF2888" bold="true" box="[392,410,1944,1968]" pageId="5" pageNumber="349">A</emphasis>
</taxonomicName>
<emphasis id="B901EA90FFABD670FE31F879C28C2888" bold="true" box="[410,441,1944,1968]" pageId="5" pageNumber="349">–D</emphasis>
,
<taxonomicName id="4C754D01FFABD670FE6DF878C18B2888" authority="Baba, 1995" authorityName="Baba" authorityYear="1995" box="[454,702,1944,1969]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="349" phylum="Arthropoda" rank="species" species="lehmanni">
<emphasis id="B901EA90FFABD670FE6DF878C2E22888" box="[454,471,1945,1968]" italics="true" pageId="5" pageNumber="349">N</emphasis>
.
<emphasis id="B901EA90FFABD670FE4EF878C1762889" box="[485,579,1945,1969]" italics="true" pageId="5" pageNumber="349">lehmanni</emphasis>
<bibRefCitation id="EFE44B73FFABD670FDE1F878C18B2888" author="Baba, M." box="[586,702,1944,1969]" pageId="5" pageNumber="349" pagination="8 - 11" refId="ref7312" refString="Baba, M. (1995) A new species of the genus Neolucanus Tomson, 1862 from south Burma (Myanmar). Gekkan-Mushi, 292, 8 - 11." type="journal article" year="1995">Baba, 1995</bibRefCitation>
</taxonomicName>
(Imitation of
<bibRefCitation id="EFE44B73FFABD670FCE6F878C0F12888" author="Baba, M." box="[845,964,1944,1969]" pageId="5" pageNumber="349" pagination="8 - 11" refId="ref7312" refString="Baba, M. (1995) A new species of the genus Neolucanus Tomson, 1862 from south Burma (Myanmar). Gekkan-Mushi, 292, 8 - 11." type="journal article" year="1995">
<collectingRegion id="49B1F860FFABD670FCE6F878C0B12889" box="[845,900,1945,1969]" country="Myanmar" name="Chin" pageId="5" pageNumber="349">Baba</collectingRegion>
, 1995
</bibRefCitation>
);
<emphasis id="B901EA90FFABD670FC7CF878C73C2889" bold="true" box="[983,1033,1945,1969]" pageId="5" pageNumber="349">E–H</emphasis>
,
<taxonomicName id="4C754D01FFABD670FBBEF878C6532889" authority="Xin, Zhong &amp; Qi" authorityName="Xin, Zhong &amp; Qi" authorityYear="2025" box="[1045,1382,1945,1969]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="349" phylum="Arthropoda" rank="species" species="horizontalis" status="new species">
<emphasis id="B901EA90FFABD670FBBEF878C7132888" box="[1045,1062,1945,1968]" italics="true" pageId="5" pageNumber="349">N</emphasis>
.
<emphasis id="B901EA90FFABD670FB98F878C7992889" box="[1075,1196,1945,1969]" italics="true" pageId="5" pageNumber="349">horizontalis</emphasis>
Xin, Zhong &amp; Qi
</taxonomicName>
,
<emphasis id="B901EA90FFABD670FADAF878C3D728EC" bold="true" pageId="5" pageNumber="349">
<taxonomicNameLabel id="A23257EBFFABD670FADAF878C3D728EC" pageId="5" pageNumber="349" rank="species">new species</taxonomicNameLabel>
</emphasis>
,
<typeStatus id="54CE8820FFABD670FF44F85CC27D28ED" box="[239,328,1981,2005]" pageId="5" pageNumber="349" type="holotype">holotype</typeStatus>
(Lvchun,
<collectingRegion id="49B1F860FFABD670FE04F85CC13428ED" box="[431,513,1981,2005]" country="China" name="Yunnan" pageId="5" pageNumber="349">Yunnan</collectingRegion>
);
<emphasis id="B901EA90FFABD670FDB8F85CC10E28ED" bold="true" box="[531,571,1981,2005]" pageId="5" pageNumber="349">I–L</emphasis>
,
<taxonomicName id="4C754D01FFABD670FDECF85CC01F28EC" authority="Didier, 1926" authorityName="Didier" authorityYear="1926" box="[583,810,1980,2005]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="349" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="B901EA90FFABD670FDECF85CC16D28EC" box="[583,600,1981,2004]" italics="true" pageId="5" pageNumber="349">N</emphasis>
.
<emphasis id="B901EA90FFABD670FDCEF85CC19128ED" box="[613,676,1981,2005]" italics="true" pageId="5" pageNumber="349">fuscus</emphasis>
<bibRefCitation id="EFE44B73FFABD670FD00F85CC01F28EC" author="Didier, R." box="[683,810,1980,2005]" pageId="5" pageNumber="349" pagination="210 - 214" refId="ref7377" refString="Didier, R. (1926) Descriptions sommaires de Lucanides nouveaux, du genre Neolucanus, de la faune indo-chinoise. Bulletin de la Societe Entomologique de France, 1926, 210 - 214. https: // doi. org / 10.3406 / bsef. 1926.27736" type="journal article" year="1926">Didier, 1926</bibRefCitation>
</taxonomicName>
(Ziyuan,
<collectingRegion id="49B1F860FFABD670FC26F85CC0DC28ED" box="[909,1001,1981,2005]" country="China" name="Guangxi" pageId="5" pageNumber="349">Guangxi</collectingRegion>
);
<emphasis id="B901EA90FFABD670FC51F85CC71B28ED" bold="true" box="[1018,1070,1981,2005]" pageId="5" pageNumber="349">M–P</emphasis>
,
<taxonomicName id="4C754D01FFABD670FB90F85CC3FF28C1" authority="Bomans &amp; Miyashita, 1997" authorityName="Bomans &amp; Miyashita" authorityYear="1997" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="349" phylum="Arthropoda" rank="species" species="brochieri">
<emphasis id="B901EA90FFABD670FB90F85CC77928EC" box="[1083,1100,1981,2004]" italics="true" pageId="5" pageNumber="349">N</emphasis>
.
<emphasis id="B901EA90FFABD670FBF2F85CC78328ED" box="[1113,1206,1981,2005]" italics="true" pageId="5" pageNumber="349">brochieri</emphasis>
<bibRefCitation id="EFE44B73FFABD670FB16F85CC3FF28C1" author="Bomans, H. E. &amp; Miyashita, T." pageId="5" pageNumber="349" pagination="2 - 3" refId="ref7344" refString="Bomans, H. E. &amp; Miyashita, T. (1997) Description de trois nouvelles especes de Lucanidae du nord Birmanie. Besoiro, 4, 2 - 3." type="journal article" year="1997">Bomans &amp; Miyashita, 1997</bibRefCitation>
</taxonomicName>
(Gonshan,
<collectingRegion id="49B1F860FFABD670FE95F800C2A528C1" box="[318,400,2017,2041]" country="China" name="Yunnan" pageId="5" pageNumber="349">Yunnan</collectingRegion>
).
<emphasis id="B901EA90FFABD670FE0AF801C03C28C0" bold="true" box="[417,777,2016,2041]" pageId="5" pageNumber="349">A, C–D, E, G–H, I, K–L, M, O–P,</emphasis>
dorsal view;
<emphasis id="B901EA90FFABD670FC39F800C0C028C1" bold="true" box="[914,1013,2017,2041]" pageId="5" pageNumber="349">B, F, J, N</emphasis>
lateral view.
</paragraph>
</subSubSection>
<caption id="DF0A660AFFA8D673FF3CF9CFC221298A" ID-DOI="http://doi.org/10.5281/zenodo.14735141" ID-Zenodo-Dep="14735141" httpUri="https://zenodo.org/record/14735141/files/figure.png" pageId="6" pageNumber="350" startId="6.[151,250,1582,1606]" targetBox="[158,1429,188,1551]" targetPageId="6" targetType="figure">
<paragraph id="8BCA3682FFA8D673FF3CF9CFC221298A" blockId="6.[151,1437,1582,1714]" pageId="6" pageNumber="350">
<emphasis id="B901EA90FFA8D673FF3CF9CFC227297E" bold="true" box="[151,274,1582,1606]" pageId="6" pageNumber="350">FIGURE 4.</emphasis>
Male fore-head
<emphasis id="B901EA90FFA8D673FE1CF9CFC169297E" bold="true" box="[439,604,1582,1607]" pageId="6" pageNumber="350">(A–B, E–F, I–J)</emphasis>
, pronotum
<emphasis id="B901EA90FFA8D673FD64F9CFC004297E" bold="true" box="[719,817,1582,1606]" pageId="6" pageNumber="350">(C, G, K)</emphasis>
, protibia and apical part of profemur
<emphasis id="B901EA90FFA8D673FB0DF9CFC633297E" bold="true" box="[1190,1286,1582,1607]" pageId="6" pageNumber="350">(D, H, L)</emphasis>
of
<taxonomicName id="4C754D01FFA8D673FA8EF9CFC3C82952" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="350" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="B901EA90FFA8D673FA8EF9CFC6A8297E" box="[1317,1437,1582,1606]" italics="true" pageId="6" pageNumber="350">Neolucanus</emphasis>
species.
<emphasis id="B901EA90FFA8D673FF40F9B3C3C82952" bold="true" box="[235,253,1618,1642]" pageId="6" pageNumber="350">A</emphasis>
</taxonomicName>
<emphasis id="B901EA90FFA8D673FF56F9B3C2282952" bold="true" box="[253,285,1618,1642]" pageId="6" pageNumber="350">–D</emphasis>
,
<taxonomicName id="4C754D01FFA8D673FE83F9B2C15B2952" authority="Xin, Zhong &amp; Qi" authorityName="Xin, Zhong &amp; Qi" authorityYear="2025" box="[296,622,1618,1643]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="350" phylum="Arthropoda" rank="species" species="horizontalis" status="new species">
<emphasis id="B901EA90FFA8D673FE83F9B2C20C2952" box="[296,313,1619,1642]" italics="true" pageId="6" pageNumber="350">N</emphasis>
.
<emphasis id="B901EA90FFA8D673FEEFF9B3C2882952" box="[324,445,1618,1642]" italics="true" pageId="6" pageNumber="350">horizontalis</emphasis>
Xin, Zhong &amp; Qi
</taxonomicName>
,
<emphasis id="B901EA90FFA8D673FDDDF9B2C1C72952" bold="true" box="[630,754,1618,1643]" pageId="6" pageNumber="350">
<taxonomicNameLabel id="A23257EBFFA8D673FDDDF9B2C1C72952" box="[630,754,1618,1643]" pageId="6" pageNumber="350" rank="species">new species</taxonomicNameLabel>
</emphasis>
, paratype (Jinping, Yunnan);
<emphasis id="B901EA90FFA8D673FBBCF9B2C77C2953" bold="true" box="[1047,1097,1619,1643]" pageId="6" pageNumber="350">E–H</emphasis>
,
<taxonomicName id="4C754D01FFA8D673FBFFF9B2C6042952" authority="Didier, 1926" authorityName="Didier" authorityYear="1926" box="[1108,1329,1618,1643]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="350" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="B901EA90FFA8D673FBFFF9B2C7502952" box="[1108,1125,1619,1642]" italics="true" pageId="6" pageNumber="350">N</emphasis>
.
<emphasis id="B901EA90FFA8D673FBDBF9B2C79A2953" box="[1136,1199,1619,1643]" italics="true" pageId="6" pageNumber="350">fuscus</emphasis>
<bibRefCitation id="EFE44B73FFA8D673FB1FF9B3C6042952" author="Didier, R." box="[1204,1329,1618,1642]" pageId="6" pageNumber="350" pagination="210 - 214" refId="ref7377" refString="Didier, R. (1926) Descriptions sommaires de Lucanides nouveaux, du genre Neolucanus, de la faune indo-chinoise. Bulletin de la Societe Entomologique de France, 1926, 210 - 214. https: // doi. org / 10.3406 / bsef. 1926.27736" type="journal article" year="1926">Didier, 1926</bibRefCitation>
</taxonomicName>
(Wuming, Guangxi);
<emphasis id="B901EA90FFA8D673FEAEF996C21829B7" bold="true" box="[261,301,1655,1679]" pageId="6" pageNumber="350">I–L</emphasis>
,
<taxonomicName id="4C754D01FFA8D673FE90F996C1EF29B7" authority="Bomans &amp; Miyashita, 1997" authorityName="Bomans &amp; Miyashita" authorityYear="1997" box="[315,730,1654,1679]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="350" phylum="Arthropoda" rank="species" species="brochieri">
<emphasis id="B901EA90FFA8D673FE90F996C27929B6" box="[315,332,1655,1678]" italics="true" pageId="6" pageNumber="350">N</emphasis>
.
<emphasis id="B901EA90FFA8D673FEF2F997C28329B6" box="[345,438,1654,1678]" italics="true" pageId="6" pageNumber="350">brochieri</emphasis>
<bibRefCitation id="EFE44B73FFA8D673FE15F996C1EF29B7" author="Bomans, H. E. &amp; Miyashita, T." box="[446,730,1654,1679]" pageId="6" pageNumber="350" pagination="2 - 3" refId="ref7344" refString="Bomans, H. E. &amp; Miyashita, T. (1997) Description de trois nouvelles especes de Lucanidae du nord Birmanie. Besoiro, 4, 2 - 3." type="journal article" year="1997">Bomans &amp; Miyashita, 1997</bibRefCitation>
</taxonomicName>
(Gonshan, Yunnan).
<emphasis id="B901EA90FFA8D673FC1FF997C7F629B7" bold="true" box="[948,1219,1654,1679]" pageId="6" pageNumber="350">A, C–D, E, G–H, I, K–L,</emphasis>
dorsal view;
<emphasis id="B901EA90FFA8D673FAFBF996C6A929B7" bold="true" box="[1360,1436,1655,1679]" pageId="6" pageNumber="350">B, F, J,</emphasis>
lateral view.
</paragraph>
</caption>
<subSubSection id="C36F6509FFA8D673FF6CF8E5C0072827" box="[199,818,1796,1823]" pageId="6" pageNumber="350" type="biology_ecology">
<paragraph id="8BCA3682FFA8D673FF6CF8E5C0072827" blockId="6.[151,1436,1796,1895]" box="[199,818,1796,1823]" pageId="6" pageNumber="350">
<emphasis id="B901EA90FFA8D673FF6CF8E5C2122826" bold="true" box="[199,295,1796,1822]" pageId="6" pageNumber="350">Biology.</emphasis>
This species can be attracted to artificial light.
</paragraph>
</subSubSection>
<subSubSection id="C36F6509FFA8D672FF6CF8C8C62A28B3" lastPageId="7" lastPageNumber="351" pageId="6" pageNumber="350" type="discussion">
<paragraph id="8BCA3682FFA8D673FF6CF8C8C6B0285F" blockId="6.[151,1436,1796,1895]" pageId="6" pageNumber="350">
<emphasis id="B901EA90FFA8D673FF6CF8C8C20E287B" bold="true" box="[199,315,1833,1859]" pageId="6" pageNumber="350">Remarks.</emphasis>
Many species within the
<taxonomicName id="4C754D01FFA8D673FDE4F8C8C181287B" authorityName="Didier" authorityYear="1926" box="[591,692,1833,1859]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="350" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="B901EA90FFA8D673FDE4F8C8C157287A" box="[591,610,1833,1858]" italics="true" pageId="6" pageNumber="350">N</emphasis>
.
<emphasis id="B901EA90FFA8D673FDC5F8C8C181287B" box="[622,692,1833,1859]" italics="true" pageId="6" pageNumber="350">fuscus</emphasis>
</taxonomicName>
group exhibit individuals with elytra featuring prominent yellowish areas. Therefore, we hypothesize that similar individuals will also be found within the newly discovered species.
</paragraph>
<paragraph id="8BCA3682FFA9D672FF3CF8E6C62A28B3" blockId="7.[151,1437,1799,1931]" pageId="7" pageNumber="351">
<emphasis id="B901EA90FFA9D672FF3CF8E6C2272827" bold="true" box="[151,274,1799,1823]" pageId="7" pageNumber="351">FIGURE 5.</emphasis>
Male abdominal tergite VIII (
<emphasis id="B901EA90FFA9D672FD94F8E6C1932827" bold="true" box="[575,678,1799,1823]" pageId="7" pageNumber="351">A, E, I, M</emphasis>
), abdominal ventrite VIII (
<emphasis id="B901EA90FFA9D672FC1DF8E6C7202827" bold="true" box="[950,1045,1799,1823]" pageId="7" pageNumber="351">B, F, J, N</emphasis>
) and abdominal segment IX (
<emphasis id="B901EA90FFA9D672FA94F8E6C213287B" bold="true" pageId="7" pageNumber="351">C–D, G– H, K–L, O–P</emphasis>
) of
<taxonomicName id="4C754D01FFA9D672FEFBF8CAC103287B" box="[336,566,1835,1859]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="351" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="B901EA90FFA9D672FEFBF8CAC2FD287B" box="[336,456,1835,1859]" italics="true" pageId="7" pageNumber="351">Neolucanus</emphasis>
species.
<emphasis id="B901EA90FFA9D672FD8FF8CAC103287B" bold="true" box="[548,566,1835,1859]" pageId="7" pageNumber="351">A</emphasis>
</taxonomicName>
<emphasis id="B901EA90FFA9D672FD9CF8CAC168287B" bold="true" box="[567,605,1835,1859]" pageId="7" pageNumber="351">–H,</emphasis>
<taxonomicName id="4C754D01FFA9D672FDCFF8CDC086287B" authority="Xin, Zhong &amp; Qi" authorityName="Xin, Zhong &amp; Qi" authorityYear="2025" box="[612,947,1835,1859]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="351" phylum="Arthropoda" rank="species" species="horizontalis" status="new species">
<emphasis id="B901EA90FFA9D672FDCFF8CDC140287B" box="[612,629,1836,1859]" italics="true" pageId="7" pageNumber="351">N</emphasis>
.
<emphasis id="B901EA90FFA9D672FD29F8CAC1CE287B" box="[642,763,1835,1859]" italics="true" pageId="7" pageNumber="351">horizontalis</emphasis>
Xin, Zhong &amp; Qi
</taxonomicName>
,
<emphasis id="B901EA90FFA9D672FC16F8CAC74D287B" bold="true" box="[957,1144,1835,1859]" pageId="7" pageNumber="351">
<taxonomicNameLabel id="A23257EBFFA9D672FC16F8CAC70F287B" box="[957,1082,1835,1859]" pageId="7" pageNumber="351" rank="species">new species</taxonomicNameLabel>
A–D,
</emphasis>
<typeStatus id="54CE8820FFA9D672FBD5F8CAC7E2287B" box="[1150,1239,1835,1859]" pageId="7" pageNumber="351" type="holotype">holotype</typeStatus>
(Lvchun,
<collectingRegion id="49B1F860FFA9D672FA94F8CAC6A4287B" box="[1343,1425,1835,1859]" country="China" name="Yunnan" pageId="7" pageNumber="351">Yunnan</collectingRegion>
);
<emphasis id="B901EA90FFA9D672FF3CF8AEC3FA285F" bold="true" box="[151,207,1871,1895]" pageId="7" pageNumber="351">E–H,</emphasis>
<typeStatus id="54CE8820FFA9D672FF7FF8B1C21E2850" box="[212,299,1872,1896]" pageId="7" pageNumber="351" type="paratype">paratype</typeStatus>
(Jinping,
<collectingRegion id="49B1F860FFA9D672FE25F8AEC2D5285F" box="[398,480,1871,1895]" country="China" name="Yunnan" pageId="7" pageNumber="351">Yunnan</collectingRegion>
).
<emphasis id="B901EA90FFA9D672FE44F8AEC128285F" bold="true" box="[495,541,1871,1895]" pageId="7" pageNumber="351">I–L,</emphasis>
<taxonomicName id="4C754D01FFA9D672FD89F8B1C034285F" authority="Didier, 1926" authorityName="Didier" authorityYear="1926" box="[546,769,1871,1895]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="351" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="B901EA90FFA9D672FD89F8B1C106285F" box="[546,563,1872,1895]" italics="true" pageId="7" pageNumber="351">N</emphasis>
.
<emphasis id="B901EA90FFA9D672FD95F8AEC148285F" box="[574,637,1871,1895]" italics="true" pageId="7" pageNumber="351">fuscus</emphasis>
<bibRefCitation id="EFE44B73FFA9D672FD29F8AEC034285F" author="Didier, R." box="[642,769,1871,1895]" pageId="7" pageNumber="351" pagination="210 - 214" refId="ref7377" refString="Didier, R. (1926) Descriptions sommaires de Lucanides nouveaux, du genre Neolucanus, de la faune indo-chinoise. Bulletin de la Societe Entomologique de France, 1926, 210 - 214. https: // doi. org / 10.3406 / bsef. 1926.27736" type="journal article" year="1926">Didier, 1926</bibRefCitation>
</taxonomicName>
(Ziyuan,
<collectingRegion id="49B1F860FFA9D672FCCAF8AEC088285F" box="[865,957,1871,1895]" country="China" name="Guangxi" pageId="7" pageNumber="351">Guangxi</collectingRegion>
).
<emphasis id="B901EA90FFA9D672FC67F8AEC731285F" bold="true" box="[972,1028,1871,1895]" pageId="7" pageNumber="351">M–P,</emphasis>
<taxonomicName id="4C754D01FFA9D672FBA2F8B1C6A8285F" authority="Bomans &amp; Miyashita, 1997" authorityName="Bomans &amp; Miyashita" authorityYear="1997" box="[1033,1437,1871,1895]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="351" phylum="Arthropoda" rank="species" species="brochieri">
<emphasis id="B901EA90FFA9D672FBA2F8B1C72F285F" box="[1033,1050,1872,1895]" italics="true" pageId="7" pageNumber="351">N</emphasis>
.
<emphasis id="B901EA90FFA9D672FB8EF8AEC7B7285F" box="[1061,1154,1871,1895]" italics="true" pageId="7" pageNumber="351">brochieri</emphasis>
<bibRefCitation id="EFE44B73FFA9D672FB23F8B1C6A8285F" author="Bomans, H. E. &amp; Miyashita, T." box="[1160,1437,1871,1895]" pageId="7" pageNumber="351" pagination="2 - 3" refId="ref7344" refString="Bomans, H. E. &amp; Miyashita, T. (1997) Description de trois nouvelles especes de Lucanidae du nord Birmanie. Besoiro, 4, 2 - 3." type="journal article" year="1997">Bomans &amp; Miyashita, 1997</bibRefCitation>
</taxonomicName>
(Gonshan,
<collectingRegion id="49B1F860FFA9D672FEAEF892C26228B3" box="[261,343,1907,1931]" country="China" name="Yunnan" pageId="7" pageNumber="351">Yunnan</collectingRegion>
).
<emphasis id="B901EA90FFA9D672FEC3F892C16C28B3" bold="true" box="[360,601,1907,1931]" pageId="7" pageNumber="351">A, C, E, G, I, K, M, O,</emphasis>
dorsal view;
<emphasis id="B901EA90FFA9D672FD49F892C0F628B3" bold="true" box="[738,963,1907,1931]" pageId="7" pageNumber="351">B, D, F, H, J, L, N, P,</emphasis>
ventral view. Scale bar = 1.0 mm.
</paragraph>
</subSubSection>
</treatment>
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688
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<paragraph id="8BF736DEFF82DD5BFF1D828435A63C6A" blockId="14.[151,975,510,573]" box="[151,975,510,537]" pageId="14" pageNumber="415">
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<taxonomicName id="4C484D5DFF82DD5BFF1D828435A63C6A" ID-CoL="47SLT" authority="Wildekamp, Watters &amp; Sainthouse, 2002" authorityName="Wildekamp, Watters &amp; Sainthouse" authorityYear="2002" box="[151,975,510,537]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="14" pageNumber="415" phylum="Chordata" rank="species" species="geminus">
<emphasis id="B93CEACCFF82DD5BFF1D828435A63C6A" bold="true" box="[151,975,510,537]" pageId="14" pageNumber="415">
<emphasis id="B93CEACCFF82DD5BFF1D828437A23C6B" bold="true" box="[151,459,510,537]" italics="true" pageId="14" pageNumber="415">Nothobranchius geminus</emphasis>
<bibRefCitation id="EFD94B2FFF82DD5BFE5B828435A63C6A" author="Wildekamp, R. H. &amp; Watters, B. R. &amp; Sainthouse, I. F. N." box="[465,975,510,537]" pageId="14" pageNumber="415" pagination="1 - 10" refId="ref35525" refString="Wildekamp, R. H., Watters, B. R. &amp; Sainthouse, I. F. N. (2002) Two new species of annual fishes of the genus Nothobranchius (Cyprinodontiformes: Aplocheilidae) from the Kilombero River basin, Tanzania. Ichthyological Exploration of Freshwaters, 13 (1), 1 - 10." type="journal article" year="2002">Wildekamp, Watters &amp; Sainthouse, 2002</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3526555FF82DD5BFF1D815937423C4F" box="[151,299,546,573]" pageId="14" pageNumber="415" type="description">
<paragraph id="8BF736DEFF82DD5BFF1D815937423C4F" blockId="14.[151,975,510,573]" box="[151,299,546,573]" pageId="14" pageNumber="415">
(
<figureCitation id="13732A5BFF82DD5BFF158159374A3C4F" box="[159,291,546,573]" captionStart="FIGURE 5" captionStartId="16.[151,250,1708,1732]" captionTargetBox="[151,1435,882,1682]" captionTargetId="figure-326@16.[151,1435,882,1682]" captionTargetPageId="16" captionText="FIGURE 5. Wild-caught specimens of Nothobranchius geminus and examples of typical habitat.A. N. geminus Ifakara TAN 95- 4, male, Kilombero River drainage, type locality; B. N. geminus TSTS 2010-11, female, Kilombero River drainage (photograph by B. Nagy); C. Location Ifakara TAN 95-4, type locality, heavily vegetated shallow habitat at margin of the Kibasira Swamp (photograph June 7, 1995), hosting N. geminus, N. lucius, N. kilomberoensis and N. lourensi. D. Location Lupiro TAN 00-15, a typical heavily vegetated shallow habitat, part of a stream system flowing into the Kibasira Swamp (photograph June 11, 2000), hosting N. geminus, N. kilomberoensis, N. lucius and N. lourensi. Photographs by B.R. Watters except where otherwise noted." pageId="14" pageNumber="415">Figs. 5A–B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3526555FF82DD5BFF1D8110378B3D28" pageId="14" pageNumber="415" type="materials_examined">
<paragraph id="8BF736DEFF82DD5BFF1D811032773CBE" blockId="14.[151,1437,618,2038]" pageId="14" pageNumber="415">
<emphasis id="B93CEACCFF82DD5BFF1D811037EB3CF6" bold="true" box="[151,386,618,644]" pageId="14" pageNumber="415">Material examined.</emphasis>
<materialsCitation id="3B203C83FF82DD5BFE01811035C73CF6" box="[395,942,618,645]" collectionCode="RMCA" pageId="14" pageNumber="415" specimenCode="RMCA 97-83" specimenCount="1" typeStatus="holotype">
<specimenCode id="DBEE9EA5FF82DD5BFE01811034433CF6" box="[395,554,618,644]" collectionCode="RMCA" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="14" pageNumber="415" type="Museum">RMCA 97-83</specimenCode>
-P-1,
<typeStatus id="54F3887CFF82DD5BFDEF811034A33CF6" box="[613,714,618,644]" pageId="14" pageNumber="415" type="holotype">holotype</typeStatus>
, male,
<quantity id="4CB09B3BFF82DD5BFC97811035EB3CF7" box="[797,898,618,645]" metricMagnitude="-2" metricUnit="m" metricValue="3.2299999999999995" pageId="14" pageNumber="415" unit="mm" value="32.3">32.3 mm</quantity>
SL
</materialsCitation>
; –
<materialsCitation id="3B203C83FF82DD5BFC43811037463CBE" collectionCode="RMCA" collectorName="B. Watters" country="Tanzania" latitude="-8.167367" location="Kilombero" longLatPrecision="1" longitude="36.693516" municipality="Ifakara" pageId="14" pageNumber="415" specimenCode="RMCA 97-83" specimenCount="16" specimenCount-female="7" specimenCount-male="9" typeStatus="paratype">
<specimenCode id="DBEE9EA5FF82DD5BFC438110320F3CF6" box="[969,1126,618,644]" collectionCode="RMCA" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="14" pageNumber="415" type="Museum">RMCA 97-83</specimenCode>
-P-2-17,
<typeStatus id="54F3887CFF82DD5BFB438111335E3CF7" box="[1225,1335,619,645]" pageId="14" pageNumber="415" type="paratype">paratypes</typeStatus>
,
<specimenCount id="9D4EFD57FF82DD5BFACE811033F53CF6" box="[1348,1436,618,644]" count="9" pageId="14" pageNumber="415" type="male">9 males</specimenCount>
<quantity id="4CB09B3BFF82DD5BFF1D81F437533CDA" box="[151,314,654,680]" metricMagnitude="-2" metricUnit="m" metricValue="3.03" metricValueMax="3.2299999999999995" metricValueMin="2.83" pageId="14" pageNumber="415" unit="mm" value="30.299999999999997" valueMax="32.3" valueMin="28.3">28.3–32.3 mm</quantity>
SL,
<specimenCount id="9D4EFD57FF82DD5BFEE581F437B73CDA" box="[367,478,654,680]" count="7" pageId="14" pageNumber="415" type="female">7 females</specimenCount>
,
<quantity id="4CB09B3BFF82DD5BFE6381F434E53CDA" box="[489,652,654,681]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" metricValueMax="3.02" metricValueMin="2.58" pageId="14" pageNumber="415" unit="mm" value="28.0" valueMax="30.2" valueMin="25.8">25.8–30.2 mm</quantity>
SL;
<collectingCountry id="F35F764EFF82DD5BFD4881F435423CDA" box="[706,811,654,680]" name="Tanzania" pageId="14" pageNumber="415">Tanzania</collectingCountry>
:
<location id="8E976005FF82DD5BFCBF81F435C73CDA" LSID="urn:lsid:plazi:treatment:03E187C8FF82DD44FF1D828433FD385F:8E976005FF82DD5BFCBF81F435C73CDA" box="[821,942,654,680]" country="Tanzania" latitude="-8.167367" longLatPrecision="1" longitude="36.693516" municipality="Ifakara" name="Kilombero" pageId="14" pageNumber="415">Kilombero</location>
drainage,
<collectingMunicipality id="6B93ACA4FF82DD5BFBAE81F4321E3CDA" box="[1060,1143,654,680]" pageId="14" pageNumber="415">Ifakara</collectingMunicipality>
;
<geoCoordinate id="EE7C5019FF82DD5BFB0881F4337C3CDA" box="[1154,1301,654,680]" degrees="08" direction="south" minutes="10.042" orientation="latitude" pageId="14" pageNumber="415" precision="1" value="-8.167367">08°10.042’ S</geoCoordinate>
,
<geoCoordinate id="EE7C5019FF82DD5BFAA881F436C23CBE" degrees="36" direction="east" minutes="41.611" orientation="longitude" pageId="14" pageNumber="415" precision="1" value="36.693516">36°41.611’ E</geoCoordinate>
;
<collectorName id="26BD5308FF82DD5BFF3D81C837463CBE" box="[183,303,690,716]" pageId="14" pageNumber="415">B. Watters</collectorName>
</materialsCitation>
,
<materialsCitation id="3B203C83FF82DD5BFEB081C832723CBE" box="[314,1051,689,716]" collectionCode="R" collectorName="Wildekamp, I &amp; June" country="Tanzania" pageId="14" pageNumber="415" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED59AE1BFF82DD5BFEB081C837393CBE" box="[314,336,690,716]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="14" pageNumber="415">R</collectionCode>
.
<collectorName id="26BD5308FF82DD5BFED081CB379F3CBE" box="[346,502,689,716]" pageId="14" pageNumber="415">Wildekamp, I</collectorName>
. Sainthouse; 7
<collectorName id="26BD5308FF82DD5BFD2A81C834BF3CBE" box="[672,726,690,716]" pageId="14" pageNumber="415">June</collectorName>
, 1995; field code: TAN 95-4
</materialsCitation>
.
</paragraph>
<paragraph id="8BF736DEFF82DD5BFF4D81AF378B3D28" blockId="14.[151,1437,618,2038]" pageId="14" pageNumber="415">
<emphasis id="B93CEACCFF82DD5BFF4D81AF345C3C9D" bold="true" box="[199,565,725,751]" pageId="14" pageNumber="415">Additional material examined.</emphasis>
<materialsCitation id="3B203C83FF82DD5BFDB681AF32F83D61" collectingDate="2002-06-09" collectorName="B. Watters &amp; B. Cooper &amp; O. Schmidt" country="Tanzania" latitude="-8.148033" location="Kilombero" longLatPrecision="1" longitude="36.415184" municipality="Narubungo" pageId="14" pageNumber="415" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D4EFD57FF82DD5BFDB681AF34E33C9D" box="[572,650,725,751]" count="1" pageId="14" pageNumber="415" type="male">1 male</specimenCount>
,
<quantity id="4CB09B3BFF82DD5BFD1C81AF34933C82" box="[662,762,725,752]" metricMagnitude="-2" metricUnit="m" metricValue="2.81" pageId="14" pageNumber="415" unit="mm" value="28.1">28.1 mm</quantity>
SL;
<collectingCountry id="F35F764EFF82DD5BFCBB81AF35F33C9D" box="[817,922,725,751]" name="Tanzania" pageId="14" pageNumber="415">Tanzania</collectingCountry>
:
<location id="8E976005FF82DD5BFC2E81AF32743C9D" LSID="urn:lsid:plazi:treatment:03E187C8FF82DD44FF1D828433FD385F:8E976005FF82DD5BFC2E81AF32743C9D" box="[932,1053,725,751]" country="Tanzania" latitude="-8.148033" longLatPrecision="1" longitude="36.415184" municipality="Narubungo" name="Kilombero" pageId="14" pageNumber="415">Kilombero</location>
drainage,
<collectingMunicipality id="6B93ACA4FF82DD5BFB1E81AF337F3C9D" box="[1172,1302,725,751]" pageId="14" pageNumber="415">Narubungo</collectingMunicipality>
;
<geoCoordinate id="EE7C5019FF82DD5BFAAB81AF36C13D61" degrees="08" direction="south" minutes="08.882" orientation="latitude" pageId="14" pageNumber="415" precision="1" value="-8.148033">08°08.882’ S</geoCoordinate>
,
<geoCoordinate id="EE7C5019FF82DD5BFF3C818337243D66" box="[182,333,761,788]" degrees="36" direction="east" minutes="24.911" orientation="longitude" pageId="14" pageNumber="415" precision="1" value="36.415184">36°24.911’ E</geoCoordinate>
;
<collectorName id="26BD5308FF82DD5BFED0818037BA3D66" box="[346,467,762,788]" pageId="14" pageNumber="415">B. Watters</collectorName>
,
<collectorName id="26BD5308FF82DD5BFE558180343C3D61" box="[479,597,761,788]" pageId="14" pageNumber="415">B. Cooper</collectorName>
,
<collectorName id="26BD5308FF82DD5BFDE88183348F3D61" box="[610,742,761,787]" pageId="14" pageNumber="415">O. Schmidt</collectorName>
;
<date id="FFF6101EFF82DD5BFD79818335E83D61" box="[755,897,761,788]" pageId="14" pageNumber="415" value="2002-06-09">
<collectingDate id="EFB2E9F6FF82DD5BFD79818335E83D61" box="[755,897,761,788]" pageId="14" pageNumber="415" value="2002-06-09">9 June 2002</collectingDate>
</date>
; field code: TAN 02-24
</materialsCitation>
. –
<materialsCitation id="3B203C83FF82DD5BFB25818337B63D28" collectingDate="2000-06-12" collectorName="B. Watters &amp; B. Cooper &amp; O. Schmidt" country="Tanzania" latitude="-8.552917" location="Kilombero" longLatPrecision="1" longitude="36.50408" municipality="Iragua" pageId="14" pageNumber="415" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D4EFD57FF82DD5BFB25818332933D61" box="[1199,1274,761,787]" count="1" pageId="14" pageNumber="415" type="male">1 male</specimenCount>
,
<quantity id="4CB09B3BFF82DD5BFA8D818333053D66" box="[1287,1388,761,788]" metricMagnitude="-2" metricUnit="m" metricValue="3.01" pageId="14" pageNumber="415" unit="mm" value="30.1">30.1 mm</quantity>
SL;
<collectingCountry id="F35F764EFF82DD5BFF1D806737693D45" box="[151,256,797,823]" name="Tanzania" pageId="14" pageNumber="415">Tanzania</collectingCountry>
:
<location id="8E976005FF82DD5BFE87806737EF3D45" LSID="urn:lsid:plazi:treatment:03E187C8FF82DD44FF1D828433FD385F:8E976005FF82DD5BFE87806737EF3D45" box="[269,390,797,823]" country="Tanzania" latitude="-8.552917" longLatPrecision="1" longitude="36.50408" municipality="Iragua" name="Kilombero" pageId="14" pageNumber="415">Kilombero</location>
drainage,
<collectingMunicipality id="6B93ACA4FF82DD5BFD8A806734223D45" box="[512,587,797,823]" pageId="14" pageNumber="415">Iragua</collectingMunicipality>
;
<geoCoordinate id="EE7C5019FF82DD5BFDD3806734863D45" box="[601,751,797,824]" degrees="08" direction="south" minutes="33.175" orientation="latitude" pageId="14" pageNumber="415" precision="1" value="-8.552917">08°33.175’ S</geoCoordinate>
,
<geoCoordinate id="EE7C5019FF82DD5BFD74806735FF3D45" box="[766,918,797,824]" degrees="36" direction="east" minutes="30.245" orientation="longitude" pageId="14" pageNumber="415" precision="1" value="36.50408">36°30.245’ E</geoCoordinate>
;
<collectorName id="26BD5308FF82DD5BFC2E806732763D45" box="[932,1055,797,823]" pageId="14" pageNumber="415">B. Watters</collectorName>
,
<collectorName id="26BD5308FF82DD5BFBA6806732CA3D45" box="[1068,1187,797,823]" pageId="14" pageNumber="415">B. Cooper</collectorName>
,
<collectorName id="26BD5308FF82DD5BFB3B8067335F3D45" box="[1201,1334,797,823]" pageId="14" pageNumber="415">O. Schmidt</collectorName>
;
<date id="FFF6101EFF82DD5BFACE806736BA3D29" pageId="14" pageNumber="415" value="2000-06-12">
<collectingDate id="EFB2E9F6FF82DD5BFACE806736BA3D29" pageId="14" pageNumber="415" value="2000-06-12">12 June 2000</collectingDate>
</date>
; field code: TAN 00-16
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3526555FF82DD5BFF4D801E34DE3BAC" pageId="14" pageNumber="415" type="diagnosis">
<paragraph id="8BF736DEFF82DD5BFF4D801E34DE3BAC" blockId="14.[151,1437,618,2038]" pageId="14" pageNumber="415">
<emphasis id="B93CEACCFF82DD5BFF4D801E372B3D0C" bold="true" box="[199,322,868,894]" pageId="14" pageNumber="415">Diagnosis.</emphasis>
<taxonomicName id="4C484D5DFF82DD5BFECF801E34333D0C" authorityName="Wildekamp, Watters &amp; Sainthouse" authorityYear="2002" box="[325,602,868,894]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="14" pageNumber="415" phylum="Chordata" rank="species" species="geminus">
<emphasis id="B93CEACCFF82DD5BFECF801E34333D0C" box="[325,602,868,894]" italics="true" pageId="14" pageNumber="415">Nothobranchius geminus</emphasis>
</taxonomicName>
belongs to the
<taxonomicName id="4C484D5DFF82DD5BFD72801F35023D0C" box="[760,875,869,894]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="14" pageNumber="415" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF82DD5BFD72801F35023D0C" box="[760,875,869,894]" italics="true" pageId="14" pageNumber="415">N.janpapi</emphasis>
</taxonomicName>
species group within the subgenus
<taxonomicName id="4C484D5DFF82DD5BFB6F801E33F43D0C" authorityName="Wildekamp" authorityYear="1977" box="[1253,1437,868,894]" class="Actinopterygii" family="Nothobranchiidae" genus="Aphyobranchius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyprinodontiformes" pageId="14" pageNumber="415" phylum="Chordata" rank="genus">
<emphasis id="B93CEACCFF82DD5BFB6F801E33F43D0C" box="[1253,1437,868,894]" italics="true" pageId="14" pageNumber="415">Aphyobranchius</emphasis>
</taxonomicName>
by presenting the following combination of characters in the male: light blue body colouration with narrow red scale margins and yellow-brown to yellow-green dorsum stripe; rearward positioned dorsal fin with predorsal length of 61.2–70.8 % SL and predorsal length
<quantity id="4CB09B3BFF82DD5BFDC880B534A93D9B" box="[578,704,975,1002]" metricMagnitude="0" metricUnit="m" metricValue="3.0099" metricValueMax="3.175" metricValueMin="2.8448" pageId="14" pageNumber="415" unit="in" value="118.5" valueMax="125.0" valueMin="112.0">112–125 in</quantity>
% of preanal length; dorsal fin relatively small with low number of rays, 11–13; cephalic sensory system at supraorbital level in an undivided, slightly curved, almost flat, shallow groove with three exposed neuromasts and pronounced lobes (
<figureCitation id="13732A5BFF82DD5BFCCC876D35E63A43" box="[838,911,1047,1073]" captionStart="FIGURE 3" captionStartId="10.[151,250,1307,1331]" captionTargetBox="[156,1428,706,1276]" captionTargetId="figure-280@10.[151,1436,702,1281]" captionTargetPageId="10" captionText="FIGURE 3. Cephalic structure: a, Nothobranchius janpapi; b, N. luekei; c, N. geminus; d, N. lourensi and N. fuscotaeniatus. Key to label abbreviations: an, anterior nostril; pn, posterior nostril; f, frontal neuromast; so, supraorbital canal; st, supratemporal canal." pageId="14" pageNumber="415">Fig 3c</figureCitation>
). Both sexes with central 2–3 rays of the caudal fin elongated forming a posterior lobe (vs. central lobe absent).
<taxonomicName id="4C484D5DFF82DD5BFCEF874132173A26" authorityName="Wildekamp, Watters &amp; Sainthouse" authorityYear="2002" box="[869,1150,1083,1109]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="14" pageNumber="415" phylum="Chordata" rank="species" species="geminus">
<emphasis id="B93CEACCFF82DD5BFCEF874132173A26" box="[869,1150,1083,1109]" italics="true" pageId="14" pageNumber="415">Nothobranchius geminus</emphasis>
</taxonomicName>
is distinguished from the other members of the
<taxonomicName id="4C484D5DFF82DD5BFE1B872534603A0A" box="[401,521,1119,1144]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="14" pageNumber="415" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF82DD5BFE1B872534603A0A" box="[401,521,1119,1144]" italics="true" pageId="14" pageNumber="415">N. janpapi</emphasis>
</taxonomicName>
species group by male colouration of anal fin with slender pale yellow subdistal band and narrow red margin (vs. slender orange-red distal band with narrow light blue margin in
<taxonomicName id="4C484D5DFF82DD5BFB4D87F933543AEE" box="[1223,1341,1155,1180]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="14" pageNumber="415" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF82DD5BFB4D87F933543AEE" box="[1223,1341,1155,1180]" italics="true" pageId="14" pageNumber="415">N. janpapi</emphasis>
</taxonomicName>
); caudal fin plain red to orange-red with narrow black distal margin (vs. base and proximal zone olive, and with light grey to translucent subdistal band and narrow dark grey distal band in
<taxonomicName id="4C484D5DFF82DD5BFCD987B035A03A91" box="[851,969,1226,1251]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="14" pageNumber="415" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF82DD5BFCD987B035A03A91" box="[851,969,1226,1251]" italics="true" pageId="14" pageNumber="415">N. janpapi</emphasis>
</taxonomicName>
, and base and proximal zone grey to light red with red subdistal band and narrow black distal band in
<taxonomicName id="4C484D5DFF82DD5BFCCA879435CD3B75" box="[832,932,1261,1287]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="14" pageNumber="415" phylum="Chordata" rank="species" species="luekei">
<emphasis id="B93CEACCFF82DD5BFCCA879435CD3B75" box="[832,932,1261,1287]" italics="true" pageId="14" pageNumber="415">N. luekei</emphasis>
</taxonomicName>
); pelvic fin yellow to red with narrow black margin (vs. plain orange-red with narrow light blue distal band in
<taxonomicName id="4C484D5DFF82DD5BFCF48668359C3B59" box="[894,1013,1298,1323]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="14" pageNumber="415" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF82DD5BFCF48668359C3B59" box="[894,1013,1298,1323]" italics="true" pageId="14" pageNumber="415">N. janpapi</emphasis>
</taxonomicName>
, and light blue with narrow red distal margin in
<taxonomicName id="4C484D5DFF82DD5BFE81864C37063B3D" box="[267,367,1333,1359]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="14" pageNumber="415" phylum="Chordata" rank="species" species="luekei">
<emphasis id="B93CEACCFF82DD5BFE81864C37063B3D" box="[267,367,1333,1359]" italics="true" pageId="14" pageNumber="415">N. luekei</emphasis>
</taxonomicName>
); by morphometric characters of head depth 66–72 % HL (vs.
<quantity id="4CB09B3BFF82DD5BFBBC864F32F03B3D" box="[1078,1177,1333,1359]" metricMagnitude="0" metricUnit="m" metricValue="1.9685" metricValueMax="2.0574" metricValueMin="1.8796" pageId="14" pageNumber="415" unit="in" value="77.5" valueMax="81.0" valueMin="74.0">74–81 in</quantity>
<taxonomicName id="4C484D5DFF82DD5BFB2B864C33713B3D" box="[1185,1304,1334,1359]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="14" pageNumber="415" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF82DD5BFB2B864C33713B3D" box="[1185,1304,1334,1359]" italics="true" pageId="14" pageNumber="415">N. janpapi</emphasis>
</taxonomicName>
, and
<quantity id="4CB09B3BFF82DD5BFADC864F36C43B00" metricMagnitude="0" metricUnit="m" metricValue="1.4986000000000002" metricValueMax="1.6002000000000003" metricValueMin="1.3969999999999998" pageId="14" pageNumber="415" unit="in" value="59.0" valueMax="63.0" valueMin="55.0">55–63 in</quantity>
<taxonomicName id="4C484D5DFF82DD5BFF3E862337713B00" box="[180,280,1368,1394]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="14" pageNumber="415" phylum="Chordata" rank="species" species="luekei">
<emphasis id="B93CEACCFF82DD5BFF3E862337713B00" box="[180,280,1368,1394]" italics="true" pageId="14" pageNumber="415">N. luekei</emphasis>
</taxonomicName>
) and caudal peduncle length 121–130 as % of its depth (vs.
<quantity id="4CB09B3BFF82DD5BFC4B862332293B00" box="[961,1088,1368,1395]" metricMagnitude="0" metricUnit="m" metricValue="3.4671" metricValueMax="3.556" metricValueMin="3.3781999999999996" pageId="14" pageNumber="415" unit="in" value="136.5" valueMax="140.0" valueMin="133.0">133–140 in</quantity>
<taxonomicName id="4C484D5DFF82DD5BFBCD862332D73B00" box="[1095,1214,1369,1394]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="14" pageNumber="415" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF82DD5BFBCD862332D73B00" box="[1095,1214,1369,1394]" italics="true" pageId="14" pageNumber="415">N. janpapi</emphasis>
</taxonomicName>
, and
<quantity id="4CB09B3BFF82DD5BFB76862233123B00" box="[1276,1403,1368,1395]" metricMagnitude="0" metricUnit="m" metricValue="3.8608" metricValueMax="3.9624" metricValueMin="3.7592" pageId="14" pageNumber="415" unit="in" value="152.0" valueMax="156.0" valueMin="148.0">148–156 in</quantity>
<taxonomicName id="4C484D5DFF82DD5BFA09862336B33BE4" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="14" pageNumber="415" phylum="Chordata" rank="species" species="luekei">
<emphasis id="B93CEACCFF82DD5BFA09862336B33BE4" italics="true" pageId="14" pageNumber="415">N. luekei</emphasis>
</taxonomicName>
); and, it differs from
<taxonomicName id="4C484D5DFF82DD5BFE4D860734543BE4" box="[455,573,1405,1430]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="14" pageNumber="415" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF82DD5BFE4D860734543BE4" box="[455,573,1405,1430]" italics="true" pageId="14" pageNumber="415">N. janpapi</emphasis>
</taxonomicName>
by dorsal-fin base length 15.6–18.6 % SL (vs. 18.8–25.1). Further, the female differs in the colouration element of small, sparsely distributed red-brown spots across mid- to upper parts of flanks and head (vs. absent in
<taxonomicName id="4C484D5DFF82DD5BFE1486BE347C3BAF" box="[414,533,1476,1501]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="14" pageNumber="415" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF82DD5BFE1486BE347C3BAF" box="[414,533,1476,1501]" italics="true" pageId="14" pageNumber="415">N. janpapi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C484D5DFF82DD5BFDC186BE34C73BAC" box="[587,686,1476,1502]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="14" pageNumber="415" phylum="Chordata" rank="species" species="luekei">
<emphasis id="B93CEACCFF82DD5BFDC186BE34C73BAC" box="[587,686,1476,1502]" italics="true" pageId="14" pageNumber="415">N. luekei</emphasis>
</taxonomicName>
)
</paragraph>
</subSubSection>
<subSubSection id="C3526555FF82DD44FF4D869D342E3DD7" lastPageId="17" lastPageNumber="418" pageId="14" pageNumber="415" type="description">
<paragraph id="8BF736DEFF82DD5BFF4D869D34F43854" blockId="14.[151,1437,618,2038]" pageId="14" pageNumber="415">
<emphasis id="B93CEACCFF82DD5BFF4D869D37303873" bold="true" box="[199,345,1511,1537]" pageId="14" pageNumber="415">Description.</emphasis>
General body features illustrated by
<figureCitation id="13732A5BFF82DD5BFC9A869D35DC3870" box="[784,949,1511,1538]" captionStart="FIGURE 5" captionStartId="16.[151,250,1708,1732]" captionTargetBox="[151,1435,882,1682]" captionTargetId="figure-326@16.[151,1435,882,1682]" captionTargetPageId="16" captionText="FIGURE 5. Wild-caught specimens of Nothobranchius geminus and examples of typical habitat.A. N. geminus Ifakara TAN 95- 4, male, Kilombero River drainage, type locality; B. N. geminus TSTS 2010-11, female, Kilombero River drainage (photograph by B. Nagy); C. Location Ifakara TAN 95-4, type locality, heavily vegetated shallow habitat at margin of the Kibasira Swamp (photograph June 7, 1995), hosting N. geminus, N. lucius, N. kilomberoensis and N. lourensi. D. Location Lupiro TAN 00-15, a typical heavily vegetated shallow habitat, part of a stream system flowing into the Kibasira Swamp (photograph June 11, 2000), hosting N. geminus, N. kilomberoensis, N. lucius and N. lourensi. Photographs by B.R. Watters except where otherwise noted." pageId="14" pageNumber="415">Figures 5A–B</figureCitation>
. Morphometric and meristic characters of examined material are summarized in
<tableCitation id="C6CA0365FF82DD5BFDC9857134F13854" box="[579,664,1547,1574]" captionStart="TABLE 5" captionStartId="15.[151,239,151,175]" captionText="TABLE 5. Morphometric and meristic data of Nothobranchius geminus. SD, standard deviation." pageId="14" pageNumber="415">Table 5</tableCitation>
.
</paragraph>
<paragraph id="8BF736DEFF82DD5BFF4D85553737396D" blockId="14.[151,1437,618,2038]" pageId="14" pageNumber="415">
Male: Small
<taxonomicName id="4C484D5DFF82DD5BFEEB8555341C383B" box="[353,629,1583,1609]" pageId="14" pageNumber="415">
<emphasis id="B93CEACCFF82DD5BFEEB8555347C383B" box="[353,533,1583,1609]" italics="true" pageId="14" pageNumber="415">Nothobranchius</emphasis>
species
</taxonomicName>
, maximum recorded size
<quantity id="4CB09B3BFF82DD5BFC2C85553266383B" box="[934,1039,1583,1609]" metricMagnitude="-2" metricUnit="m" metricValue="3.2299999999999995" pageId="14" pageNumber="415" unit="mm" value="32.3">32.3 mm</quantity>
SL. General body shape laterally compressed. Greatest body depth at vertical in front of pelvic-fin origin (25.3–30.2 % SL). Greatest body width at pectoral-fin base with body progressively narrowing towards caudal-fin base. Dorsal profile slightly concave from tip of snout to nape and straight to slightly convex from nape to dorsal-fin origin, straight to slightly concave on caudal peduncle. Ventral profile convex from lower jaw to base of last anal-fin ray, straight to slightly concave on caudal peduncle. Caudal peduncle moderately deep, length 121–130 % of its depth. Anus situated directly in front of anal-fin origin.
</paragraph>
<paragraph id="8BF736DEFF82DD5BFF4D845335F93984" blockId="14.[151,1437,618,2038]" pageId="14" pageNumber="415">Head moderately long (27.9–32.4 % SL), laterally compressed, deeper than wide (head width 73–83 % of its depth). Snout distinctly pointed, smaller than eye diameter. Mouth supraterminal, slightly oblique in profile. Jaws subequal, lower jaw longer than upper, posterior end of rictus at same level or slightly ventral to centre of eye. Premaxilla and dentary with many irregularly distributed conical teeth on inner rows and slightly curved teeth on outer row. Orbit large, eye diameter 28–34 % HL, mainly in anterior half of head (snout to eye end length 49–58 % HL). Branchiostegal membrane projecting posteriorly from opercle.</paragraph>
<caption id="DF376656FF83DD5AFF1D83ED32F63EDD" box="[151,1183,151,176]" pageId="15" pageNumber="416" startId="15.[151,239,151,175]" targetType="table">
<paragraph id="8BF736DEFF83DD5AFF1D83ED32F63EDD" blockId="15.[151,1183,151,176]" box="[151,1183,151,176]" pageId="15" pageNumber="416">
<emphasis id="B93CEACCFF83DD5AFF1D83ED37633EC2" bold="true" box="[151,266,151,176]" pageId="15" pageNumber="416">TABLE 5.</emphasis>
Morphometric and meristic data of
<taxonomicName id="4C484D5DFF83DD5AFD0183ED35FE3EDD" authorityName="Wildekamp, Watters &amp; Sainthouse" authorityYear="2002" box="[651,919,151,175]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="15" pageNumber="416" phylum="Chordata" rank="species" species="geminus">
<emphasis id="B93CEACCFF83DD5AFD0183ED35FE3EDD" box="[651,919,151,175]" italics="true" pageId="15" pageNumber="416">Nothobranchius geminus</emphasis>
</taxonomicName>
. SD, standard deviation.
</paragraph>
</caption>
<paragraph id="8BF736DEFF83DD5AFD7E83BA32B738BB" pageId="15" pageNumber="416">
<table id="F948C47EFF8322AAFF2983BA333638BB" box="[163,1375,192,1737]" colsContinueIn="22.[153,1394,192,1665]" gridcols="7" gridrows="36" pageId="15" pageNumber="416">
<tr id="3578349CFF8322AAFF2983BA33363EAA" box="[163,1375,192,216]" gridrow="0" pageId="15" pageNumber="416" rowspan-0="1" rowspan-3="1" rowspan-4="1" rowspan-6="1">
<th id="76A95DE0FF8322AAFDF983BA353B3EAA" box="[627,850,192,216]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="15" pageNumber="416">males</th>
<th id="76A95DE0FF8322AAFB2F83BA32973EAA" box="[1189,1278,192,216]" gridcol="5" gridrow="0" pageId="15" pageNumber="416">females</th>
</tr>
<tr id="3578349CFF8322AAFF29839233363F72" box="[163,1375,232,256]" gridrow="1" pageId="15" pageNumber="416" rowspan-0="1" rowspan-3="1" rowspan-4="1" rowspan-6="1">
<td id="76A95DE0FF8322AAFDF98392353B3F72" box="[627,850,232,256]" colspan="2" colspanRight="1" gridcol="1" gridrow="1" pageId="15" pageNumber="416">(n = 12)</td>
<td id="76A95DE0FF8322AAFB2F839232973F72" box="[1189,1278,232,256]" gridcol="5" gridrow="1" pageId="15" pageNumber="416">(n = 7)</td>
</tr>
<tr id="3578349CFF8322AAFF29824333363F23" box="[163,1375,313,337]" gridrow="2" pageId="15" pageNumber="416" rowspan-0="1">
<td id="76A95DE0FF8322AAFDF98243349B3F23" box="[627,754,313,337]" gridcol="1" gridrow="2" pageId="15" pageNumber="416">range</td>
<td id="76A95DE0FF8322AAFC938243353B3F23" box="[793,850,313,337]" gridcol="2" gridrow="2" pageId="15" pageNumber="416">mean</td>
<td id="76A95DE0FF8322AAFCF1824335F23F23" box="[891,923,313,337]" gridcol="3" gridrow="2" pageId="15" pageNumber="416">SD</td>
<td id="76A95DE0FF8322AAFB8D824332EC3F23" box="[1031,1157,313,337]" gridcol="4" gridrow="2" pageId="15" pageNumber="416">range</td>
<td id="76A95DE0FF8322AAFB2F824332973F23" box="[1189,1278,313,337]" gridcol="5" gridrow="2" pageId="15" pageNumber="416">mean</td>
<td id="76A95DE0FF8322AAFAB5824333363F23" box="[1343,1375,313,337]" gridcol="6" gridrow="2" pageId="15" pageNumber="416">SD</td>
</tr>
<tr id="3578349CFF8322AAFF29821B33363F0B" box="[163,1375,353,377]" gridrow="3" pageId="15" pageNumber="416" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1">
<th id="76A95DE0FF8322AAFF29821B342A3F0B" box="[163,579,353,377]" gridcol="0" gridrow="3" pageId="15" pageNumber="416">Standard length (mm)</th>
<td id="76A95DE0FF8322AAFDF9821B349B3F0B" box="[627,754,353,377]" gridcol="1" gridrow="3" pageId="15" pageNumber="416">28.1–32.3</td>
<td id="76A95DE0FF8322AAFB8D821B32EC3F0B" box="[1031,1157,353,377]" gridcol="4" gridrow="3" pageId="15" pageNumber="416">25.8–30.2</td>
</tr>
<tr id="3578349CFF8322AAFF2982F333363FD3" box="[163,1375,393,417]" gridrow="4" pageId="15" pageNumber="416">
<th id="76A95DE0FF8322AAFF2982F333363FD3" box="[163,1375,393,417]" colspan="7" colspanRight="6" gridcol="0" gridrow="4" pageId="15" pageNumber="416">
<emphasis id="B93CEACCFF83DD5AFF2982F337AB3FD3" bold="true" box="[163,450,393,417]" pageId="15" pageNumber="416">Percent of standard length</emphasis>
</th>
</tr>
<tr id="3578349CFF8322AAFF2982CB33363FBB" box="[163,1375,433,457]" gridrow="5" pageId="15" pageNumber="416">
<th id="76A95DE0FF8322AAFF2982CB342A3FBB" box="[163,579,433,457]" gridcol="0" gridrow="5" pageId="15" pageNumber="416">Total length</th>
<td id="76A95DE0FF8322AAFDF982CB349B3FBB" box="[627,754,433,457]" gridcol="1" gridrow="5" pageId="15" pageNumber="416">125.5–132.5</td>
<td id="76A95DE0FF8322AAFC9382CB353B3FBB" box="[793,850,433,457]" gridcol="2" gridrow="5" pageId="15" pageNumber="416">128</td>
<td id="76A95DE0FF8322AAFCF182CB35F23FBB" box="[891,923,433,457]" gridcol="3" gridrow="5" pageId="15" pageNumber="416">1.8</td>
<td id="76A95DE0FF8322AAFB8D82CB32EC3FBB" box="[1031,1157,433,457]" gridcol="4" gridrow="5" pageId="15" pageNumber="416">123.4–131.1</td>
<td id="76A95DE0FF8322AAFB2F82CB32973FBB" box="[1189,1278,433,457]" gridcol="5" gridrow="5" pageId="15" pageNumber="416">128</td>
<td id="76A95DE0FF8322AAFAB582CB33363FBB" box="[1343,1375,433,457]" gridcol="6" gridrow="5" pageId="15" pageNumber="416">3.0</td>
</tr>
<tr id="3578349CFF8322AAFF2982A333363F80" box="[163,1375,473,498]" gridrow="6" pageId="15" pageNumber="416">
<th id="76A95DE0FF8322AAFF2982A3342A3F80" box="[163,579,473,498]" gridcol="0" gridrow="6" pageId="15" pageNumber="416">Body depth at pelvic-fin origin</th>
<td id="76A95DE0FF8322AAFDF982A3349B3F80" box="[627,754,473,498]" gridcol="1" gridrow="6" pageId="15" pageNumber="416">25.3–30.2</td>
<td id="76A95DE0FF8322AAFC9382A3353B3F80" box="[793,850,473,498]" gridcol="2" gridrow="6" pageId="15" pageNumber="416">28.6</td>
<td id="76A95DE0FF8322AAFCF182A335F23F80" box="[891,923,473,498]" gridcol="3" gridrow="6" pageId="15" pageNumber="416">1.5</td>
<td id="76A95DE0FF8322AAFB8D82A332EC3F80" box="[1031,1157,473,498]" gridcol="4" gridrow="6" pageId="15" pageNumber="416">22.0–26.7</td>
<td id="76A95DE0FF8322AAFB2F82A332973F80" box="[1189,1278,473,498]" gridcol="5" gridrow="6" pageId="15" pageNumber="416">24.7</td>
<td id="76A95DE0FF8322AAFAB582A333363F80" box="[1343,1375,473,498]" gridcol="6" gridrow="6" pageId="15" pageNumber="416">1.8</td>
</tr>
<tr id="3578349CFF8322AAFF29817833363C68" box="[163,1375,514,538]" gridrow="7" pageId="15" pageNumber="416">
<th id="76A95DE0FF8322AAFF298178342A3C68" box="[163,579,514,538]" gridcol="0" gridrow="7" pageId="15" pageNumber="416">Head length</th>
<td id="76A95DE0FF8322AAFDF98178349B3C68" box="[627,754,514,538]" gridcol="1" gridrow="7" pageId="15" pageNumber="416">27.9–32.4</td>
<td id="76A95DE0FF8322AAFC938178353B3C68" box="[793,850,514,538]" gridcol="2" gridrow="7" pageId="15" pageNumber="416">29.1</td>
<td id="76A95DE0FF8322AAFCF1817835F23C68" box="[891,923,514,538]" gridcol="3" gridrow="7" pageId="15" pageNumber="416">1.3</td>
<td id="76A95DE0FF8322AAFB8D817832EC3C68" box="[1031,1157,514,538]" gridcol="4" gridrow="7" pageId="15" pageNumber="416">23.2–30.2</td>
<td id="76A95DE0FF8322AAFB2F817832973C68" box="[1189,1278,514,538]" gridcol="5" gridrow="7" pageId="15" pageNumber="416">26.5</td>
<td id="76A95DE0FF8322AAFAB5817833363C68" box="[1343,1375,514,538]" gridcol="6" gridrow="7" pageId="15" pageNumber="416">2.2</td>
</tr>
<tr id="3578349CFF8322AAFF29815033363C30" box="[163,1375,554,578]" gridrow="8" pageId="15" pageNumber="416">
<th id="76A95DE0FF8322AAFF298150342A3C30" box="[163,579,554,578]" gridcol="0" gridrow="8" pageId="15" pageNumber="416">Preanal length</th>
<td id="76A95DE0FF8322AAFDF98150349B3C30" box="[627,754,554,578]" gridcol="1" gridrow="8" pageId="15" pageNumber="416">52.5–59.8</td>
<td id="76A95DE0FF8322AAFC938150353B3C30" box="[793,850,554,578]" gridcol="2" gridrow="8" pageId="15" pageNumber="416">56.1</td>
<td id="76A95DE0FF8322AAFCF1815035F23C30" box="[891,923,554,578]" gridcol="3" gridrow="8" pageId="15" pageNumber="416">1.8</td>
<td id="76A95DE0FF8322AAFB8D815032EC3C30" box="[1031,1157,554,578]" gridcol="4" gridrow="8" pageId="15" pageNumber="416">58.2–63.1</td>
<td id="76A95DE0FF8322AAFB2F815032973C30" box="[1189,1278,554,578]" gridcol="5" gridrow="8" pageId="15" pageNumber="416">60.9</td>
<td id="76A95DE0FF8322AAFAB5815033363C30" box="[1343,1375,554,578]" gridcol="6" gridrow="8" pageId="15" pageNumber="416">2.2</td>
</tr>
<tr id="3578349CFF8322AAFF29812833363C18" box="[163,1375,594,618]" gridrow="9" pageId="15" pageNumber="416">
<th id="76A95DE0FF8322AAFF298128342A3C18" box="[163,579,594,618]" gridcol="0" gridrow="9" pageId="15" pageNumber="416">Predorsal length</th>
<td id="76A95DE0FF8322AAFDF98128349B3C18" box="[627,754,594,618]" gridcol="1" gridrow="9" pageId="15" pageNumber="416">61.2–70.8</td>
<td id="76A95DE0FF8322AAFC938128353B3C18" box="[793,850,594,618]" gridcol="2" gridrow="9" pageId="15" pageNumber="416">66.3</td>
<td id="76A95DE0FF8322AAFCF1812835F23C18" box="[891,923,594,618]" gridcol="3" gridrow="9" pageId="15" pageNumber="416">2.4</td>
<td id="76A95DE0FF8322AAFB8D812832EC3C18" box="[1031,1157,594,618]" gridcol="4" gridrow="9" pageId="15" pageNumber="416">66.1–70.0</td>
<td id="76A95DE0FF8322AAFB2F812832973C18" box="[1189,1278,594,618]" gridcol="5" gridrow="9" pageId="15" pageNumber="416">67.4</td>
<td id="76A95DE0FF8322AAFAB5812833363C18" box="[1343,1375,594,618]" gridcol="6" gridrow="9" pageId="15" pageNumber="416">1.3</td>
</tr>
<tr id="3578349CFF8322AAFF29810033363CE1" box="[163,1375,634,659]" gridrow="10" pageId="15" pageNumber="416">
<th id="76A95DE0FF8322AAFF298100342A3CE1" box="[163,579,634,659]" gridcol="0" gridrow="10" pageId="15" pageNumber="416">Prepelvic length</th>
<td id="76A95DE0FF8322AAFDF98100349B3CE1" box="[627,754,634,659]" gridcol="1" gridrow="10" pageId="15" pageNumber="416">40.5–45.6</td>
<td id="76A95DE0FF8322AAFC938100353B3CE1" box="[793,850,634,659]" gridcol="2" gridrow="10" pageId="15" pageNumber="416">43.5</td>
<td id="76A95DE0FF8322AAFCF1810035F23CE1" box="[891,923,634,659]" gridcol="3" gridrow="10" pageId="15" pageNumber="416">1.4</td>
<td id="76A95DE0FF8322AAFB8D810032EC3CE1" box="[1031,1157,634,659]" gridcol="4" gridrow="10" pageId="15" pageNumber="416">43.4–49.3</td>
<td id="76A95DE0FF8322AAFB2F810032973CE1" box="[1189,1278,634,659]" gridcol="5" gridrow="10" pageId="15" pageNumber="416">46.2</td>
<td id="76A95DE0FF8322AAFAB5810033363CE1" box="[1343,1375,634,659]" gridcol="6" gridrow="10" pageId="15" pageNumber="416">2.2</td>
</tr>
<tr id="3578349CFF8322AAFF2981D933363CC9" box="[163,1375,675,699]" gridrow="11" pageId="15" pageNumber="416">
<th id="76A95DE0FF8322AAFF2981D9342A3CC9" box="[163,579,675,699]" gridcol="0" gridrow="11" pageId="15" pageNumber="416">Prepectoral length</th>
<td id="76A95DE0FF8322AAFDF981D9349B3CC9" box="[627,754,675,699]" gridcol="1" gridrow="11" pageId="15" pageNumber="416">26.0–32.7</td>
<td id="76A95DE0FF8322AAFC9381D9353B3CC9" box="[793,850,675,699]" gridcol="2" gridrow="11" pageId="15" pageNumber="416">29.3</td>
<td id="76A95DE0FF8322AAFCF181D935F23CC9" box="[891,923,675,699]" gridcol="3" gridrow="11" pageId="15" pageNumber="416">1.7</td>
<td id="76A95DE0FF8322AAFB8D81D932EC3CC9" box="[1031,1157,675,699]" gridcol="4" gridrow="11" pageId="15" pageNumber="416">22.3–27.4</td>
<td id="76A95DE0FF8322AAFB2F81D932973CC9" box="[1189,1278,675,699]" gridcol="5" gridrow="11" pageId="15" pageNumber="416">25.1</td>
<td id="76A95DE0FF8322AAFAB581D933363CC9" box="[1343,1375,675,699]" gridcol="6" gridrow="11" pageId="15" pageNumber="416">1.9</td>
</tr>
<tr id="3578349CFF8322AAFF2981B133363C91" box="[163,1375,715,739]" gridrow="12" pageId="15" pageNumber="416">
<th id="76A95DE0FF8322AAFF2981B1342A3C91" box="[163,579,715,739]" gridcol="0" gridrow="12" pageId="15" pageNumber="416">Caudal peduncle length</th>
<td id="76A95DE0FF8322AAFDF981B1349B3C91" box="[627,754,715,739]" gridcol="1" gridrow="12" pageId="15" pageNumber="416">15.9–21.1</td>
<td id="76A95DE0FF8322AAFC9381B1353B3C91" box="[793,850,715,739]" gridcol="2" gridrow="12" pageId="15" pageNumber="416">18.6</td>
<td id="76A95DE0FF8322AAFCF181B135F23C91" box="[891,923,715,739]" gridcol="3" gridrow="12" pageId="15" pageNumber="416">1.3</td>
<td id="76A95DE0FF8322AAFB8D81B132EC3C91" box="[1031,1157,715,739]" gridcol="4" gridrow="12" pageId="15" pageNumber="416">18.2–21.4</td>
<td id="76A95DE0FF8322AAFB2F81B132973C91" box="[1189,1278,715,739]" gridcol="5" gridrow="12" pageId="15" pageNumber="416">20.2</td>
<td id="76A95DE0FF8322AAFAB581B133363C91" box="[1343,1375,715,739]" gridcol="6" gridrow="12" pageId="15" pageNumber="416">1.0</td>
</tr>
<tr id="3578349CFF8322AAFF29818933363D79" box="[163,1375,755,779]" gridrow="13" pageId="15" pageNumber="416">
<th id="76A95DE0FF8322AAFF298189342A3D79" box="[163,579,755,779]" gridcol="0" gridrow="13" pageId="15" pageNumber="416">Caudal peduncle depth</th>
<td id="76A95DE0FF8322AAFDF98189349B3D79" box="[627,754,755,779]" gridcol="1" gridrow="13" pageId="15" pageNumber="416">13.0–16.2</td>
<td id="76A95DE0FF8322AAFC938189353B3D79" box="[793,850,755,779]" gridcol="2" gridrow="13" pageId="15" pageNumber="416">14.7</td>
<td id="76A95DE0FF8322AAFCF1818935F23D79" box="[891,923,755,779]" gridcol="3" gridrow="13" pageId="15" pageNumber="416">1.0</td>
<td id="76A95DE0FF8322AAFB8D818932EC3D79" box="[1031,1157,755,779]" gridcol="4" gridrow="13" pageId="15" pageNumber="416">11.6–13.6</td>
<td id="76A95DE0FF8322AAFB2F818932973D79" box="[1189,1278,755,779]" gridcol="5" gridrow="13" pageId="15" pageNumber="416">12.7</td>
<td id="76A95DE0FF8322AAFAB5818933363D79" box="[1343,1375,755,779]" gridcol="6" gridrow="13" pageId="15" pageNumber="416">0.6</td>
</tr>
<tr id="3578349CFF8322AAFF29806133363D46" box="[163,1375,795,820]" gridrow="14" pageId="15" pageNumber="416">
<th id="76A95DE0FF8322AAFF298061342A3D46" box="[163,579,795,820]" gridcol="0" gridrow="14" pageId="15" pageNumber="416">Dorsal-fin base length</th>
<td id="76A95DE0FF8322AAFDF98061349B3D46" box="[627,754,795,820]" gridcol="1" gridrow="14" pageId="15" pageNumber="416">15.6–18.6</td>
<td id="76A95DE0FF8322AAFC938061353B3D46" box="[793,850,795,820]" gridcol="2" gridrow="14" pageId="15" pageNumber="416">17.5</td>
<td id="76A95DE0FF8322AAFCF1806135F23D46" box="[891,923,795,820]" gridcol="3" gridrow="14" pageId="15" pageNumber="416">0.8</td>
<td id="76A95DE0FF8322AAFB8D806132EC3D46" box="[1031,1157,795,820]" gridcol="4" gridrow="14" pageId="15" pageNumber="416">14.6–16.1</td>
<td id="76A95DE0FF8322AAFB2F806132973D46" box="[1189,1278,795,820]" gridcol="5" gridrow="14" pageId="15" pageNumber="416">15.4</td>
<td id="76A95DE0FF8322AAFAB5806133363D46" box="[1343,1375,795,820]" gridcol="6" gridrow="14" pageId="15" pageNumber="416">0.6</td>
</tr>
<tr id="3578349CFF8322AAFF29803933363D2E" box="[163,1375,835,860]" gridrow="15" pageId="15" pageNumber="416">
<th id="76A95DE0FF8322AAFF298039342A3D2E" box="[163,579,835,860]" gridcol="0" gridrow="15" pageId="15" pageNumber="416">Anal-fin base length</th>
<td id="76A95DE0FF8322AAFDF98039349B3D2E" box="[627,754,835,860]" gridcol="1" gridrow="15" pageId="15" pageNumber="416">25.6–28.3</td>
<td id="76A95DE0FF8322AAFC938039353B3D2E" box="[793,850,835,860]" gridcol="2" gridrow="15" pageId="15" pageNumber="416">26.9</td>
<td id="76A95DE0FF8322AAFCF1803935F23D2E" box="[891,923,835,860]" gridcol="3" gridrow="15" pageId="15" pageNumber="416">0.9</td>
<td id="76A95DE0FF8322AAFB8D803932EC3D2E" box="[1031,1157,835,860]" gridcol="4" gridrow="15" pageId="15" pageNumber="416">17.9–21.3</td>
<td id="76A95DE0FF8322AAFB2F803932973D2E" box="[1189,1278,835,860]" gridcol="5" gridrow="15" pageId="15" pageNumber="416">20.0</td>
<td id="76A95DE0FF8322AAFAB5803933363D2E" box="[1343,1375,835,860]" gridcol="6" gridrow="15" pageId="15" pageNumber="416">1.2</td>
</tr>
<tr id="3578349CFF8322AAFF29801633363DF6" box="[163,1375,876,900]" gridrow="16" pageId="15" pageNumber="416">
<th id="76A95DE0FF8322AAFF298016342A3DF6" box="[163,579,876,900]" gridcol="0" gridrow="16" pageId="15" pageNumber="416">Caudal-fin length</th>
<td id="76A95DE0FF8322AAFDF98016349B3DF6" box="[627,754,876,900]" gridcol="1" gridrow="16" pageId="15" pageNumber="416">25.5–32.5</td>
<td id="76A95DE0FF8322AAFC938016353B3DF6" box="[793,850,876,900]" gridcol="2" gridrow="16" pageId="15" pageNumber="416">28.0</td>
<td id="76A95DE0FF8322AAFCF1801635F23DF6" box="[891,923,876,900]" gridcol="3" gridrow="16" pageId="15" pageNumber="416">1.8</td>
<td id="76A95DE0FF8322AAFB8D801632EC3DF6" box="[1031,1157,876,900]" gridcol="4" gridrow="16" pageId="15" pageNumber="416">23.4–31.1</td>
<td id="76A95DE0FF8322AAFB2F801632973DF6" box="[1189,1278,876,900]" gridcol="5" gridrow="16" pageId="15" pageNumber="416">28.0</td>
<td id="76A95DE0FF8322AAFAB5801633363DF6" box="[1343,1375,876,900]" gridcol="6" gridrow="16" pageId="15" pageNumber="416">3.0</td>
</tr>
<tr id="3578349CFF8322AAFF2980EE33363DDE" box="[163,1375,916,940]" gridrow="17" pageId="15" pageNumber="416">
<th id="76A95DE0FF8322AAFF2980EE33363DDE" box="[163,1375,916,940]" colspan="7" colspanRight="6" gridcol="0" gridrow="17" pageId="15" pageNumber="416">
<emphasis id="B93CEACCFF83DD5AFF2980EE37FC3DDE" bold="true" box="[163,405,916,940]" pageId="15" pageNumber="416">Percent of head length</emphasis>
</th>
</tr>
<tr id="3578349CFF8322AAFF2980C633363DA6" box="[163,1375,956,980]" gridrow="18" pageId="15" pageNumber="416">
<th id="76A95DE0FF8322AAFF2980C6342A3DA6" box="[163,579,956,980]" gridcol="0" gridrow="18" pageId="15" pageNumber="416">Head width</th>
<td id="76A95DE0FF8322AAFDF980C6349B3DA6" box="[627,754,956,980]" gridcol="1" gridrow="18" pageId="15" pageNumber="416">51–60</td>
<td id="76A95DE0FF8322AAFC9380C6353B3DA6" box="[793,850,956,980]" gridcol="2" gridrow="18" pageId="15" pageNumber="416">53.8</td>
<td id="76A95DE0FF8322AAFCF180C635F23DA6" box="[891,923,956,980]" gridcol="3" gridrow="18" pageId="15" pageNumber="416">2.4</td>
<td id="76A95DE0FF8322AAFB8D80C632EC3DA6" box="[1031,1157,956,980]" gridcol="4" gridrow="18" pageId="15" pageNumber="416">45–52</td>
<td id="76A95DE0FF8322AAFB2F80C632973DA6" box="[1189,1278,956,980]" gridcol="5" gridrow="18" pageId="15" pageNumber="416">48.9</td>
<td id="76A95DE0FF8322AAFAB580C633363DA6" box="[1343,1375,956,980]" gridcol="6" gridrow="18" pageId="15" pageNumber="416">2.7</td>
</tr>
<tr id="3578349CFF8322AAFF29809E33363D8F" box="[163,1375,996,1021]" gridrow="19" pageId="15" pageNumber="416">
<th id="76A95DE0FF8322AAFF29809E342A3D8F" box="[163,579,996,1021]" gridcol="0" gridrow="19" pageId="15" pageNumber="416">Head depth</th>
<td id="76A95DE0FF8322AAFDF9809E349B3D8F" box="[627,754,996,1021]" gridcol="1" gridrow="19" pageId="15" pageNumber="416">66–72</td>
<td id="76A95DE0FF8322AAFC93809E353B3D8F" box="[793,850,996,1021]" gridcol="2" gridrow="19" pageId="15" pageNumber="416">69.4</td>
<td id="76A95DE0FF8322AAFCF1809E35F23D8F" box="[891,923,996,1021]" gridcol="3" gridrow="19" pageId="15" pageNumber="416">1.8</td>
<td id="76A95DE0FF8322AAFB8D809E32EC3D8F" box="[1031,1157,996,1021]" gridcol="4" gridrow="19" pageId="15" pageNumber="416">58–65</td>
<td id="76A95DE0FF8322AAFB2F809E32973D8F" box="[1189,1278,996,1021]" gridcol="5" gridrow="19" pageId="15" pageNumber="416">60.8</td>
<td id="76A95DE0FF8322AAFAB5809E33363D8F" box="[1343,1375,996,1021]" gridcol="6" gridrow="19" pageId="15" pageNumber="416">2.8</td>
</tr>
<tr id="3578349CFF8322AAFF29877733363A57" box="[163,1375,1037,1061]" gridrow="20" pageId="15" pageNumber="416">
<th id="76A95DE0FF8322AAFF298777342A3A57" box="[163,579,1037,1061]" gridcol="0" gridrow="20" pageId="15" pageNumber="416">Interorbital width</th>
<td id="76A95DE0FF8322AAFDF98777349B3A57" box="[627,754,1037,1061]" gridcol="1" gridrow="20" pageId="15" pageNumber="416">41–50</td>
<td id="76A95DE0FF8322AAFC938777353B3A57" box="[793,850,1037,1061]" gridcol="2" gridrow="20" pageId="15" pageNumber="416">44.3</td>
<td id="76A95DE0FF8322AAFCF1877735F23A57" box="[891,923,1037,1061]" gridcol="3" gridrow="20" pageId="15" pageNumber="416">2.7</td>
<td id="76A95DE0FF8322AAFB8D877732EC3A57" box="[1031,1157,1037,1061]" gridcol="4" gridrow="20" pageId="15" pageNumber="416">38–45</td>
<td id="76A95DE0FF8322AAFB2F877732973A57" box="[1189,1278,1037,1061]" gridcol="5" gridrow="20" pageId="15" pageNumber="416">41.2</td>
<td id="76A95DE0FF8322AAFAB5877733363A57" box="[1343,1375,1037,1061]" gridcol="6" gridrow="20" pageId="15" pageNumber="416">3.0</td>
</tr>
<tr id="3578349CFF8322AAFF29874F33363A3F" box="[163,1375,1077,1101]" gridrow="21" pageId="15" pageNumber="416">
<th id="76A95DE0FF8322AAFF29874F342A3A3F" box="[163,579,1077,1101]" gridcol="0" gridrow="21" pageId="15" pageNumber="416">Postorbital length</th>
<td id="76A95DE0FF8322AAFDF9874F349B3A3F" box="[627,754,1077,1101]" gridcol="1" gridrow="21" pageId="15" pageNumber="416">42–51</td>
<td id="76A95DE0FF8322AAFC93874F353B3A3F" box="[793,850,1077,1101]" gridcol="2" gridrow="21" pageId="15" pageNumber="416">45.3</td>
<td id="76A95DE0FF8322AAFCF1874F35F23A3F" box="[891,923,1077,1101]" gridcol="3" gridrow="21" pageId="15" pageNumber="416">2.8</td>
<td id="76A95DE0FF8322AAFB8D874F32EC3A3F" box="[1031,1157,1077,1101]" gridcol="4" gridrow="21" pageId="15" pageNumber="416">40–49</td>
<td id="76A95DE0FF8322AAFB2F874F32973A3F" box="[1189,1278,1077,1101]" gridcol="5" gridrow="21" pageId="15" pageNumber="416">43.2</td>
<td id="76A95DE0FF8322AAFAB5874F33363A3F" box="[1343,1375,1077,1101]" gridcol="6" gridrow="21" pageId="15" pageNumber="416">2.8</td>
</tr>
<tr id="3578349CFF8322AAFF29872733363A07" box="[163,1375,1117,1141]" gridrow="22" pageId="15" pageNumber="416">
<th id="76A95DE0FF8322AAFF298727342A3A07" box="[163,579,1117,1141]" gridcol="0" gridrow="22" pageId="15" pageNumber="416">Suborbital depth</th>
<td id="76A95DE0FF8322AAFDF98727349B3A07" box="[627,754,1117,1141]" gridcol="1" gridrow="22" pageId="15" pageNumber="416">10–14</td>
<td id="76A95DE0FF8322AAFC938727353B3A07" box="[793,850,1117,1141]" gridcol="2" gridrow="22" pageId="15" pageNumber="416">11.8</td>
<td id="76A95DE0FF8322AAFCF1872735F23A07" box="[891,923,1117,1141]" gridcol="3" gridrow="22" pageId="15" pageNumber="416">1.0</td>
<td id="76A95DE0FF8322AAFB8D872732EC3A07" box="[1031,1157,1117,1141]" gridcol="4" gridrow="22" pageId="15" pageNumber="416">12–15</td>
<td id="76A95DE0FF8322AAFB2F872732973A07" box="[1189,1278,1117,1141]" gridcol="5" gridrow="22" pageId="15" pageNumber="416">13.3</td>
<td id="76A95DE0FF8322AAFAB5872733363A07" box="[1343,1375,1117,1141]" gridcol="6" gridrow="22" pageId="15" pageNumber="416">0.9</td>
</tr>
<tr id="3578349CFF8322AAFF2987FF33363AEC" box="[163,1375,1157,1182]" gridrow="23" pageId="15" pageNumber="416">
<th id="76A95DE0FF8322AAFF2987FF342A3AEC" box="[163,579,1157,1182]" gridcol="0" gridrow="23" pageId="15" pageNumber="416">Eye diameter</th>
<td id="76A95DE0FF8322AAFDF987FF349B3AEC" box="[627,754,1157,1182]" gridcol="1" gridrow="23" pageId="15" pageNumber="416">28–34</td>
<td id="76A95DE0FF8322AAFC9387FF353B3AEC" box="[793,850,1157,1182]" gridcol="2" gridrow="23" pageId="15" pageNumber="416">30.4</td>
<td id="76A95DE0FF8322AAFCF187FF35F23AEC" box="[891,923,1157,1182]" gridcol="3" gridrow="23" pageId="15" pageNumber="416">2.2</td>
<td id="76A95DE0FF8322AAFB8D87FF32EC3AEC" box="[1031,1157,1157,1182]" gridcol="4" gridrow="23" pageId="15" pageNumber="416">28–35</td>
<td id="76A95DE0FF8322AAFB2F87FF32973AEC" box="[1189,1278,1157,1182]" gridcol="5" gridrow="23" pageId="15" pageNumber="416">32.4</td>
<td id="76A95DE0FF8322AAFAB587FF33363AEC" box="[1343,1375,1157,1182]" gridcol="6" gridrow="23" pageId="15" pageNumber="416">2.3</td>
</tr>
<tr id="3578349CFF8322AAFF2987D433363AB4" box="[163,1375,1198,1222]" gridrow="24" pageId="15" pageNumber="416">
<th id="76A95DE0FF8322AAFF2987D4342A3AB4" box="[163,579,1198,1222]" gridcol="0" gridrow="24" pageId="15" pageNumber="416">Snout to eye end length</th>
<td id="76A95DE0FF8322AAFDF987D4349B3AB4" box="[627,754,1198,1222]" gridcol="1" gridrow="24" pageId="15" pageNumber="416">49–58</td>
<td id="76A95DE0FF8322AAFC9387D4353B3AB4" box="[793,850,1198,1222]" gridcol="2" gridrow="24" pageId="15" pageNumber="416">54.7</td>
<td id="76A95DE0FF8322AAFCF187D435F23AB4" box="[891,923,1198,1222]" gridcol="3" gridrow="24" pageId="15" pageNumber="416">2.8</td>
<td id="76A95DE0FF8322AAFB8D87D432EC3AB4" box="[1031,1157,1198,1222]" gridcol="4" gridrow="24" pageId="15" pageNumber="416">51–60</td>
<td id="76A95DE0FF8322AAFB2F87D432973AB4" box="[1189,1278,1198,1222]" gridcol="5" gridrow="24" pageId="15" pageNumber="416">56.8</td>
<td id="76A95DE0FF8322AAFAB587D433363AB4" box="[1343,1375,1198,1222]" gridcol="6" gridrow="24" pageId="15" pageNumber="416">2.8</td>
</tr>
<tr id="3578349CFF8322AAFF2987AC33363A9C" box="[163,1375,1238,1262]" gridrow="25" pageId="15" pageNumber="416">
<th id="76A95DE0FF8322AAFF2987AC342A3A9C" box="[163,579,1238,1262]" gridcol="0" gridrow="25" pageId="15" pageNumber="416">Snout length</th>
<td id="76A95DE0FF8322AAFDF987AC349B3A9C" box="[627,754,1238,1262]" gridcol="1" gridrow="25" pageId="15" pageNumber="416">21–28</td>
<td id="76A95DE0FF8322AAFC9387AC353B3A9C" box="[793,850,1238,1262]" gridcol="2" gridrow="25" pageId="15" pageNumber="416">25.3</td>
<td id="76A95DE0FF8322AAFCF187AC35F23A9C" box="[891,923,1238,1262]" gridcol="3" gridrow="25" pageId="15" pageNumber="416">2.1</td>
<td id="76A95DE0FF8322AAFB8D87AC32EC3A9C" box="[1031,1157,1238,1262]" gridcol="4" gridrow="25" pageId="15" pageNumber="416">22–29</td>
<td id="76A95DE0FF8322AAFB2F87AC32973A9C" box="[1189,1278,1238,1262]" gridcol="5" gridrow="25" pageId="15" pageNumber="416">25.9</td>
<td id="76A95DE0FF8322AAFAB587AC33363A9C" box="[1343,1375,1238,1262]" gridcol="6" gridrow="25" pageId="15" pageNumber="416">2.8</td>
</tr>
<tr id="3578349CFF8322AAFF29878433363B64" box="[163,1375,1278,1302]" gridrow="26" pageId="15" pageNumber="416">
<th id="76A95DE0FF8322AAFF29878433363B64" box="[163,1375,1278,1302]" colspan="7" colspanRight="6" gridcol="0" gridrow="26" pageId="15" pageNumber="416">
<emphasis id="B93CEACCFF83DD5AFF29878437A33B64" bold="true" box="[163,458,1278,1302]" pageId="15" pageNumber="416">Other morphometric ratios</emphasis>
</th>
</tr>
<tr id="3578349CFF8322AAFF29865D33363B11" box="[163,1375,1319,1379]" gridrow="27" pageId="15" pageNumber="416">
<th id="76A95DE0FF8322AAFF29865D342A3B11" box="[163,579,1319,1379]" gridcol="0" gridrow="27" pageId="15" pageNumber="416">Predorsal length in % of preanal length</th>
<td id="76A95DE0FF8322AAFDF9865D349B3B11" box="[627,754,1319,1379]" gridcol="1" gridrow="27" pageId="15" pageNumber="416">112–125</td>
<td id="76A95DE0FF8322AAFC93865D353B3B11" box="[793,850,1319,1379]" gridcol="2" gridrow="27" pageId="15" pageNumber="416">118.3</td>
<td id="76A95DE0FF8322AAFCF1865D35F23B11" box="[891,923,1319,1379]" gridcol="3" gridrow="27" pageId="15" pageNumber="416">3.6</td>
<td id="76A95DE0FF8322AAFB8D865D32EC3B11" box="[1031,1157,1319,1379]" gridcol="4" gridrow="27" pageId="15" pageNumber="416">106–114</td>
<td id="76A95DE0FF8322AAFB2F865D32973B11" box="[1189,1278,1319,1379]" gridcol="5" gridrow="27" pageId="15" pageNumber="416">110.7</td>
<td id="76A95DE0FF8322AAFAB5865D33363B11" box="[1343,1375,1319,1379]" gridcol="6" gridrow="27" pageId="15" pageNumber="416">3.8</td>
</tr>
<tr id="3578349CFF8322AAFF29860933363BDD" box="[163,1375,1395,1455]" gridrow="28" pageId="15" pageNumber="416">
<th id="76A95DE0FF8322AAFF298609342A3BDD" box="[163,579,1395,1455]" gridcol="0" gridrow="28" pageId="15" pageNumber="416">Caudal peduncle length in % of its depth</th>
<td id="76A95DE0FF8322AAFDF98609349B3BDD" box="[627,754,1395,1455]" gridcol="1" gridrow="28" pageId="15" pageNumber="416">121–130</td>
<td id="76A95DE0FF8322AAFC938609353B3BDD" box="[793,850,1395,1455]" gridcol="2" gridrow="28" pageId="15" pageNumber="416">127.2</td>
<td id="76A95DE0FF8322AAFCF1860935F23BDD" box="[891,923,1395,1455]" gridcol="3" gridrow="28" pageId="15" pageNumber="416">3.3</td>
<td id="76A95DE0FF8322AAFB8D860932EC3BDD" box="[1031,1157,1395,1455]" gridcol="4" gridrow="28" pageId="15" pageNumber="416">157–164</td>
<td id="76A95DE0FF8322AAFB2F860932973BDD" box="[1189,1278,1395,1455]" gridcol="5" gridrow="28" pageId="15" pageNumber="416">158.6</td>
<td id="76A95DE0FF8322AAFAB5860933363BDD" box="[1343,1375,1395,1455]" gridcol="6" gridrow="28" pageId="15" pageNumber="416">2.5</td>
</tr>
<tr id="3578349CFF8322AAFF2986C533363BA5" box="[163,1375,1471,1495]" gridrow="29" pageId="15" pageNumber="416">
<th id="76A95DE0FF8322AAFF2986C5342A3BA5" box="[163,579,1471,1495]" gridcol="0" gridrow="29" pageId="15" pageNumber="416">Head width in % of its depth</th>
<td id="76A95DE0FF8322AAFDF986C5349B3BA5" box="[627,754,1471,1495]" gridcol="1" gridrow="29" pageId="15" pageNumber="416">73–83</td>
<td id="76A95DE0FF8322AAFC9386C5353B3BA5" box="[793,850,1471,1495]" gridcol="2" gridrow="29" pageId="15" pageNumber="416">77.6</td>
<td id="76A95DE0FF8322AAFCF186C535F23BA5" box="[891,923,1471,1495]" gridcol="3" gridrow="29" pageId="15" pageNumber="416">3.3</td>
<td id="76A95DE0FF8322AAFB8D86C532EC3BA5" box="[1031,1157,1471,1495]" gridcol="4" gridrow="29" pageId="15" pageNumber="416">78–85</td>
<td id="76A95DE0FF8322AAFB2F86C532973BA5" box="[1189,1278,1471,1495]" gridcol="5" gridrow="29" pageId="15" pageNumber="416">80.4</td>
<td id="76A95DE0FF8322AAFAB586C533363BA5" box="[1343,1375,1471,1495]" gridcol="6" gridrow="29" pageId="15" pageNumber="416">2.5</td>
</tr>
<tr id="3578349CFF8322AAFF29869D33363872" box="[163,1375,1511,1536]" gridrow="30" pageId="15" pageNumber="416" rowspan-3="1" rowspan-6="1">
<th id="76A95DE0FF8322AAFF29869D342A3872" box="[163,579,1511,1536]" gridcol="0" gridrow="30" pageId="15" pageNumber="416">
<emphasis id="B93CEACCFF83DD5AFF29869D376E3B8D" bold="true" box="[163,263,1511,1535]" pageId="15" pageNumber="416">Meristics</emphasis>
</th>
<td id="76A95DE0FF8322AAFDF9869D349B3872" box="[627,754,1511,1536]" gridcol="1" gridrow="30" pageId="15" pageNumber="416">range</td>
<td id="76A95DE0FF8322AAFC93869D353B3872" box="[793,850,1511,1536]" gridcol="2" gridrow="30" pageId="15" pageNumber="416">mode</td>
<td id="76A95DE0FF8322AAFB8D869D32EC3872" box="[1031,1157,1511,1536]" gridcol="4" gridrow="30" pageId="15" pageNumber="416">range</td>
<td id="76A95DE0FF8322AAFB2F869D32973872" box="[1189,1278,1511,1536]" gridcol="5" gridrow="30" pageId="15" pageNumber="416">mode</td>
</tr>
<tr id="3578349CFF8322AAFF2985753336385A" box="[163,1375,1551,1576]" gridrow="31" pageId="15" pageNumber="416" rowspan-3="1" rowspan-6="1">
<th id="76A95DE0FF8322AAFF298575342A385A" box="[163,579,1551,1576]" gridcol="0" gridrow="31" pageId="15" pageNumber="416">Dorsal-fin rays</th>
<td id="76A95DE0FF8322AAFDF98575349B385A" box="[627,754,1551,1576]" gridcol="1" gridrow="31" pageId="15" pageNumber="416">11–13</td>
<td id="76A95DE0FF8322AAFC938575353B385A" box="[793,850,1551,1576]" gridcol="2" gridrow="31" pageId="15" pageNumber="416">11</td>
<td id="76A95DE0FF8322AAFB8D857532EC385A" box="[1031,1157,1551,1576]" gridcol="4" gridrow="31" pageId="15" pageNumber="416">11–13</td>
<td id="76A95DE0FF8322AAFB2F85753297385A" box="[1189,1278,1551,1576]" gridcol="5" gridrow="31" pageId="15" pageNumber="416">12</td>
</tr>
<tr id="3578349CFF8322AAFF29854233363822" box="[163,1375,1592,1616]" gridrow="32" pageId="15" pageNumber="416" rowspan-3="1" rowspan-6="1">
<th id="76A95DE0FF8322AAFF298542342A3822" box="[163,579,1592,1616]" gridcol="0" gridrow="32" pageId="15" pageNumber="416">Anal-fin rays</th>
<td id="76A95DE0FF8322AAFDF98542349B3822" box="[627,754,1592,1616]" gridcol="1" gridrow="32" pageId="15" pageNumber="416">17–21</td>
<td id="76A95DE0FF8322AAFC938542353B3822" box="[793,850,1592,1616]" gridcol="2" gridrow="32" pageId="15" pageNumber="416">18</td>
<td id="76A95DE0FF8322AAFB8D854232EC3822" box="[1031,1157,1592,1616]" gridcol="4" gridrow="32" pageId="15" pageNumber="416">17–20</td>
<td id="76A95DE0FF8322AAFB2F854232973822" box="[1189,1278,1592,1616]" gridcol="5" gridrow="32" pageId="15" pageNumber="416">18</td>
</tr>
<tr id="3578349CFF8322AAFF29851A3336380A" box="[163,1375,1632,1656]" gridrow="33" pageId="15" pageNumber="416" rowspan-3="1" rowspan-6="1">
<th id="76A95DE0FF8322AAFF29851A342A380A" box="[163,579,1632,1656]" gridcol="0" gridrow="33" pageId="15" pageNumber="416">Scales mid-longitudinal series</th>
<td id="76A95DE0FF8322AAFDF9851A349B380A" box="[627,754,1632,1656]" gridcol="1" gridrow="33" pageId="15" pageNumber="416">26–28</td>
<td id="76A95DE0FF8322AAFC93851A353B380A" box="[793,850,1632,1656]" gridcol="2" gridrow="33" pageId="15" pageNumber="416">27</td>
<td id="76A95DE0FF8322AAFB8D851A32EC380A" box="[1031,1157,1632,1656]" gridcol="4" gridrow="33" pageId="15" pageNumber="416">26–28</td>
<td id="76A95DE0FF8322AAFB2F851A3297380A" box="[1189,1278,1632,1656]" gridcol="5" gridrow="33" pageId="15" pageNumber="416">27</td>
</tr>
<tr id="3578349CFF8322AAFF2985F2333638D2" box="[163,1375,1672,1696]" gridrow="34" pageId="15" pageNumber="416" rowspan-3="1" rowspan-6="1">
<th id="76A95DE0FF8322AAFF2985F2342A38D2" box="[163,579,1672,1696]" gridcol="0" gridrow="34" pageId="15" pageNumber="416">Scales transverse</th>
<td id="76A95DE0FF8322AAFDF985F2349B38D2" box="[627,754,1672,1696]" gridcol="1" gridrow="34" pageId="15" pageNumber="416">11–11</td>
<td id="76A95DE0FF8322AAFC9385F2353B38D2" box="[793,850,1672,1696]" gridcol="2" gridrow="34" pageId="15" pageNumber="416">11</td>
<td id="76A95DE0FF8322AAFB8D85F232EC38D2" box="[1031,1157,1672,1696]" gridcol="4" gridrow="34" pageId="15" pageNumber="416">11–11</td>
<td id="76A95DE0FF8322AAFB2F85F2329738D2" box="[1189,1278,1672,1696]" gridcol="5" gridrow="34" pageId="15" pageNumber="416">11</td>
</tr>
<tr id="3578349CFF8322AAFF2985CA333638BB" box="[163,1375,1712,1737]" gridrow="35" pageId="15" pageNumber="416" rowspan-3="1" rowspan-6="1">
<th id="76A95DE0FF8322AAFF2985CA342A38BB" box="[163,579,1712,1737]" gridcol="0" gridrow="35" pageId="15" pageNumber="416">Scales circumpeduncular</th>
<td id="76A95DE0FF8322AAFDF985CA349B38BB" box="[627,754,1712,1737]" gridcol="1" gridrow="35" pageId="15" pageNumber="416">12–12</td>
<td id="76A95DE0FF8322AAFC9385CA353B38BB" box="[793,850,1712,1737]" gridcol="2" gridrow="35" pageId="15" pageNumber="416">12</td>
<td id="76A95DE0FF8322AAFB8D85CA32EC38BB" box="[1031,1157,1712,1737]" gridcol="4" gridrow="35" pageId="15" pageNumber="416">12–12</td>
<td id="76A95DE0FF8322AAFB2F85CA329738BB" box="[1189,1278,1712,1737]" gridcol="5" gridrow="35" pageId="15" pageNumber="416">12</td>
</tr>
</table>
</paragraph>
<paragraph id="8BF736DEFF83DD45FF4D8582350F3EA7" blockId="15.[151,1437,1784,2027]" lastBlockId="16.[151,1437,151,861]" lastPageId="16" lastPageNumber="417" pageId="15" pageNumber="416">Dorsal-fin origin markedly posterior to anal-fin origin (predorsal length 112–125 as % of preanal length), both fins originating posterior to mid-length of body (predorsal length 61.2–70.8 % SL; preanal length 52.5–59.8 % SL). Dorsal fin relatively small with low number of rays, 11–13; extremity rounded. Anal fin trapezoidal in shape, with 17–21 rays; lower margin moderately concave in profile. Dorsal and anal fins with short fin-ray extensions. Pectoral fin subtriangular, insertion slightly posterior to margin of opercular opening, base slightly oblique, upper fin rays positioned slightly anterior to lower fin rays, with tip reaching or slightly overlapping base of pelvic fin. Pelvic fin subabdominal, origin at about mid-length of body, short, base medially separated, with tip reaching urogenital papilla. Caudal fin rounded, with 13–16 branched rays, plus 3–4 dorsal and ventral procurrent rays; central 2–3 rays of fin relatively slightly elongated resulting in mild central lobe.</paragraph>
<paragraph id="8BF736DEFF9CDD45FF4D83A535983F33" blockId="16.[151,1437,151,861]" pageId="16" pageNumber="417">Scales cycloid, body and head entirely scaled, except for ventral surface of head. No scales on base of dorsal and anal fins. Scales in mid-longitudinal series 26–28 plus 2–3 small scales on caudal-fin base. Transverse rows of scales in front of dorsal-fin origin 11; scale rows around caudal peduncle 12.</paragraph>
<paragraph id="8BF736DEFF9CDD45FF4D823137CB3CF7" blockId="16.[151,1437,151,861]" pageId="16" pageNumber="417">
Cephalic squamation pattern G-type. Frontal neuromast in single shallow groove behind upper lip. Cephalic sensory system at supraorbital level in an undivided, slightly curved zigzag pattern, almost flat shallow groove with three exposed neuromasts and pronounced lobes (
<figureCitation id="13732A5BFF9CDD45FD7582E935263FDF" box="[767,847,403,429]" captionStart="FIGURE 3" captionStartId="10.[151,250,1307,1331]" captionTargetBox="[156,1428,706,1276]" captionTargetId="figure-280@10.[151,1436,702,1281]" captionTargetPageId="10" captionText="FIGURE 3. Cephalic structure: a, Nothobranchius janpapi; b, N. luekei; c, N. geminus; d, N. lourensi and N. fuscotaeniatus. Key to label abbreviations: an, anterior nostril; pn, posterior nostril; f, frontal neuromast; so, supraorbital canal; st, supratemporal canal." pageId="16" pageNumber="417">Fig. 3c</figureCitation>
), whereas at supratemporal level with three exposed neuromasts in undivided, slightly curved shallow groove. Preorbital canal in shallow groove with two exposed neuromasts; infraorbital level with series of about 20 small buttons at ventral and posterior margin of orbit; postorbital canal in deep groove with one exposed neuromast. Preopercular system with six exposed neuromasts in deep groove on preopercle portion, whereas in shallow groove with about ten small buttons on ventral portion. Mandibular canal in shallow groove with about six small neuromasts. One neuromast on each scale along trunk mid-longitudinal series.
</paragraph>
<paragraph id="8BF736DEFF9CDD45FF4D81F534E33D2F" blockId="16.[151,1437,151,861]" pageId="16" pageNumber="417">
Female: Smaller than male, maximum recorded size
<quantity id="4CB09B3BFF9CDD45FCA081F535F83CDB" box="[810,913,655,681]" metricMagnitude="-2" metricUnit="m" metricValue="3.02" pageId="16" pageNumber="417" unit="mm" value="30.2">30.2 mm</quantity>
SL. Branchiostegal membrane not projecting posteriorly from opercle. Dorsal fin rounded with low number of fin rays, 11–13. Caudal fin basically rounded but with central 2–3 rays slightly elongated resulting in pointed posterior margin. Anal fin subtriangular, tip rounded, central rays longer and more rigid.Anal-fin base length smaller than in males (17.9–21.3 % SL vs. 25.6–28.3). Head depth smaller than in males (58–65 % HL vs. 66–72). Caudal peduncle shallower than in males (caudal peduncle length 157–164 % of its depth vs. 121–130).
</paragraph>
<caption id="DF376656FF9CDD45FF1D85D633FB390B" pageId="16" pageNumber="417" startId="16.[151,250,1708,1732]" targetBox="[151,1435,882,1682]" targetPageId="16" targetType="figure">
<paragraph id="8BF736DEFF9CDD45FF1D85D633FB390B" blockId="16.[151,1436,1708,1913]" pageId="16" pageNumber="417">
<emphasis id="B93CEACCFF9CDD45FF1D85D6377838B7" bold="true" box="[151,273,1708,1733]" pageId="16" pageNumber="417">FIGURE 5.</emphasis>
Wild-caught specimens of
<taxonomicName id="4C484D5DFF9CDD45FDAB85D7357438B6" authorityName="Wildekamp, Watters &amp; Sainthouse" authorityYear="2002" box="[545,797,1709,1733]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="geminus">
<emphasis id="B93CEACCFF9CDD45FDAB85D7357438B6" box="[545,797,1709,1733]" italics="true" pageId="16" pageNumber="417">Nothobranchius geminus</emphasis>
</taxonomicName>
and examples of typical habitat.
<emphasis id="B93CEACCFF9CDD45FBEE85D6321438B6" bold="true" box="[1124,1149,1708,1732]" pageId="16" pageNumber="417">A.</emphasis>
<taxonomicName id="4C484D5DFF9CDD45FB0885D7331F38B7" authority="Ifakara TAN" authorityName="Ifakara TAN" box="[1154,1398,1709,1733]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="geminus">
<emphasis id="B93CEACCFF9CDD45FB0885D7329B38B6" box="[1154,1266,1709,1732]" italics="true" pageId="16" pageNumber="417">N. geminus</emphasis>
Ifakara TAN
</taxonomicName>
95- 4, male, Kilombero River drainage, type locality;
<emphasis id="B93CEACCFF9CDD45FD0685AB34CD389B" bold="true" box="[652,676,1745,1769]" pageId="16" pageNumber="417">B.</emphasis>
<taxonomicName id="4C484D5DFF9CDD45FD2385AB35F0389B" authority="TSTS 2010" authorityName="TSTS" authorityYear="2010" box="[681,921,1745,1769]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="geminus">
<emphasis id="B93CEACCFF9CDD45FD2385AB3573389A" box="[681,794,1745,1768]" italics="true" pageId="16" pageNumber="417">N. geminus</emphasis>
TSTS 2010
</taxonomicName>
-11, female, Kilombero River drainage (photograph by B. Nagy);
<emphasis id="B93CEACCFF9CDD45FEA9858E3752397E" bold="true" box="[291,315,1780,1804]" pageId="16" pageNumber="417">C.</emphasis>
Location Ifakara TAN 95-4, type locality, heavily vegetated shallow habitat at margin of the Kibasira Swamp (photograph June 7, 1995), hosting
<taxonomicName id="4C484D5DFF9CDD45FE75846334183942" authorityName="TSTS" authorityYear="2010" box="[511,625,1817,1840]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="geminus">
<emphasis id="B93CEACCFF9CDD45FE75846334183942" box="[511,625,1817,1840]" italics="true" pageId="16" pageNumber="417">N. geminus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C484D5DFF9CDD45FDF4846334B13943" authorityName="Wildekamp, Shidlovskiy &amp; Watters" authorityYear="2009" box="[638,728,1817,1841]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="lucius">
<emphasis id="B93CEACCFF9CDD45FDF4846334B13943" box="[638,728,1817,1841]" italics="true" pageId="16" pageNumber="417">N. lucius</emphasis>
</taxonomicName>
,
<taxonomicName id="4C484D5DFF9CDD45FD6E846335F33943" authorityName="Wildekamp, Watters &amp; Sainthouse" authorityYear="2002" box="[740,922,1817,1841]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="kilomberoensis">
<emphasis id="B93CEACCFF9CDD45FD6E846335F33943" box="[740,922,1817,1841]" italics="true" pageId="16" pageNumber="417">N. kilomberoensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C484D5DFF9CDD45FC41846332523943" authorityName=", Wildekamp" authorityYear="1977" box="[971,1083,1817,1841]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="lourensi">
<emphasis id="B93CEACCFF9CDD45FC41846332523943" box="[971,1083,1817,1841]" italics="true" pageId="16" pageNumber="417">N. lourensi</emphasis>
</taxonomicName>
.
<emphasis id="B93CEACCFF9CDD45FBCD846332093943" bold="true" box="[1095,1120,1817,1841]" pageId="16" pageNumber="417">D.</emphasis>
Location Lupiro TAN 00-15, a typical heavily vegetated shallow habitat, part of a stream system flowing into the Kibasira Swamp (photograph June 11, 2000), hosting
<taxonomicName id="4C484D5DFF9CDD45FF62841B3733390A" authorityName="TSTS" authorityYear="2010" box="[232,346,1889,1912]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="geminus">
<emphasis id="B93CEACCFF9CDD45FF62841B3733390A" box="[232,346,1889,1912]" italics="true" pageId="16" pageNumber="417">N. geminus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C484D5DFF9CDD45FEED841B3475390B" authorityName="Wildekamp, Watters &amp; Sainthouse" authorityYear="2002" box="[359,540,1889,1913]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="kilomberoensis">
<emphasis id="B93CEACCFF9CDD45FEED841B3475390B" box="[359,540,1889,1913]" italics="true" pageId="16" pageNumber="417">N. kilomberoensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C484D5DFF9CDD45FDA3841B34ED390B" authorityName="Wildekamp, Shidlovskiy &amp; Watters" authorityYear="2009" box="[553,644,1889,1913]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="lucius">
<emphasis id="B93CEACCFF9CDD45FDA3841B34ED390B" box="[553,644,1889,1913]" italics="true" pageId="16" pageNumber="417">N. lucius</emphasis>
</taxonomicName>
and
<taxonomicName id="4C484D5DFF9CDD45FD3F841B354D390B" authorityName=", Wildekamp" authorityYear="1977" box="[693,804,1889,1913]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="lourensi">
<emphasis id="B93CEACCFF9CDD45FD3F841B354D390B" box="[693,804,1889,1913]" italics="true" pageId="16" pageNumber="417">N. lourensi</emphasis>
</taxonomicName>
. Photographs by B.R. Watters except where otherwise noted.
</paragraph>
</caption>
<paragraph id="8BF736DEFF9CDD44FF4D84DD343F3D67" blockId="16.[151,1436,1959,2021]" lastBlockId="17.[151,1437,151,1581]" lastPageId="17" lastPageNumber="418" pageId="16" pageNumber="417">
<emphasis id="B93CEACCFF9CDD45FF4D84DD373639B3" bold="true" box="[199,351,1959,1985]" pageId="16" pageNumber="417">Colouration.</emphasis>
Live male (
<figureCitation id="13732A5BFF9CDD45FE7E84DD343D39B0" box="[500,596,1959,1986]" captionStart="FIGURE 5" captionStartId="16.[151,250,1708,1732]" captionTargetBox="[151,1435,882,1682]" captionTargetId="figure-326@16.[151,1435,882,1682]" captionTargetPageId="16" captionText="FIGURE 5. Wild-caught specimens of Nothobranchius geminus and examples of typical habitat.A. N. geminus Ifakara TAN 95- 4, male, Kilombero River drainage, type locality; B. N. geminus TSTS 2010-11, female, Kilombero River drainage (photograph by B. Nagy); C. Location Ifakara TAN 95-4, type locality, heavily vegetated shallow habitat at margin of the Kibasira Swamp (photograph June 7, 1995), hosting N. geminus, N. lucius, N. kilomberoensis and N. lourensi. D. Location Lupiro TAN 00-15, a typical heavily vegetated shallow habitat, part of a stream system flowing into the Kibasira Swamp (photograph June 11, 2000), hosting N. geminus, N. kilomberoensis, N. lucius and N. lourensi. Photographs by B.R. Watters except where otherwise noted." pageId="16" pageNumber="417">Fig. 5A</figureCitation>
): Scales on the trunk and head light iridescent blue to blue-green with prominent red margins, forming a regular reticulation pattern. Scales on the abdomen are faint blue to cream with less pronounced margins. Scales on the dorsum anterior to the dorsal fin have yellow-brown centres forming a narrow zone. Snout, frontal and dorsal portions of the head and throat are yellow. Operculum yellow with sparse blue to blue-green and red markings. Exposed part of the branchiostegal membrane is yellow with white edging. Iris golden yellow to golden blue-green, reflective pale blue when viewed from above. Dorsal fin light blue-green in the basal and proximal zones grading to yellow-green through the medial zone and into the distal zone, overlaid by irregular red spots and streaks in crude parallelism with fin rays in the proximal and submedial zones but arranged somewhat concentrically conforming to the fin margin in the medial and posterior zones; the distal zone comprises a solid red band that may be followed by a very narrow black margin. Short extensions to the dorsal and anal fins are black where the similarly coloured margin is present but are otherwise red. Anal fin background colour is light blue in the basal zone and light blue or yellow in the proximal zone grading to yellow through the medial and subdistal zones; the proximal and medial zones are overlaid by red spots that commonly merge to form stripes aligned parallel to the fin rays. The density of these red markings is variable at both the inter-and intrapopulation levels to the extent that they may be present in the proximal zone but grade outwards to absent in the medial zone. There is always a narrow red margin to the anal fin and short fin-ray extensions may be red or black as discussed above. The caudal fin is red to orange-red with a narrow dark grey to black margin most prominent anteriorly; fin rays may be a darker red than the intervening membrane. Pelvic fins are yellow to light blue with a variable density of red spots, and a red marginal zone that may also show incomplete black edging. Pectoral fins are dominantly hyaline or pale yellow with narrow and irregular blue margins.
</paragraph>
<paragraph id="8BF736DEFF9DDD44FF4D8065342E3DD7" blockId="17.[151,1437,151,1581]" pageId="17" pageNumber="418">
Live female (
<figureCitation id="13732A5BFF9DDD44FED1806537DA3D4B" box="[347,435,798,825]" captionStart="FIGURE 5" captionStartId="16.[151,250,1708,1732]" captionTargetBox="[151,1435,882,1682]" captionTargetId="figure-326@16.[151,1435,882,1682]" captionTargetPageId="16" captionText="FIGURE 5. Wild-caught specimens of Nothobranchius geminus and examples of typical habitat.A. N. geminus Ifakara TAN 95- 4, male, Kilombero River drainage, type locality; B. N. geminus TSTS 2010-11, female, Kilombero River drainage (photograph by B. Nagy); C. Location Ifakara TAN 95-4, type locality, heavily vegetated shallow habitat at margin of the Kibasira Swamp (photograph June 7, 1995), hosting N. geminus, N. lucius, N. kilomberoensis and N. lourensi. D. Location Lupiro TAN 00-15, a typical heavily vegetated shallow habitat, part of a stream system flowing into the Kibasira Swamp (photograph June 11, 2000), hosting N. geminus, N. kilomberoensis, N. lucius and N. lourensi. Photographs by B.R. Watters except where otherwise noted." pageId="17" pageNumber="418">Fig. 5B</figureCitation>
): The trunk and head are overall light grey with light blue to blue-green iridescence on the scales which have pale grey edgings; small red-brown spots are sparsely distributed across the mid- to upper parts of the flanks and head, being more common over the upper body. The operculum is iridescent very pale blue-green. All fins are hyaline. The iris is golden.
</paragraph>
</subSubSection>
<subSubSection id="C3526555FF9DDD44FF4D80D433FD385F" pageId="17" pageNumber="418" type="distribution">
<paragraph id="8BF736DEFF9DDD44FF4D80D434A03AB7" blockId="17.[151,1437,151,1581]" pageId="17" pageNumber="418">
<emphasis id="B93CEACCFF9DDD44FF4D80D437083DBA" bold="true" box="[199,353,942,968]" pageId="17" pageNumber="418">Distribution.</emphasis>
<taxonomicName id="4C484D5DFF9DDD44FEE280D534EB3DBA" authorityName="TSTS" authorityYear="2010" box="[360,642,943,969]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="17" pageNumber="418" phylum="Chordata" rank="species" species="geminus">
<emphasis id="B93CEACCFF9DDD44FEE280D534EB3DBA" box="[360,642,943,969]" italics="true" pageId="17" pageNumber="418">Nothobranchius geminus</emphasis>
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is endemic to seasonal freshwater habitats in central-eastern
<collectingCountry id="F35F764EFF9DDD44FAB280D533F43DBB" box="[1336,1437,943,969]" name="Tanzania" pageId="17" pageNumber="418">Tanzania</collectingCountry>
and has been found in the seasonal marginal and peripheral zones of the Kibasira Swamp in the Kilombero River valley. The latter system represents a fairly remote component of the Rufiji River system, connected to it through the Ulanga River, the downstream continuation of the Kilombero River. Known sites are associated with the northeastern part of the Swamp, south-southwest and west-southwest of the town of Ifakara (
<bibRefCitation id="EFD94B2FFF9DDD44FBCB874532853A2B" author="Watters, B. R." box="[1089,1260,1087,1113]" pageId="17" pageNumber="418" pagination="131 - 153" refId="ref35043" refString="Watters, B. R. (2016 b) Nothobranchius fishes of the Kibasira Swamp. Journal of the American Killifish Association, 49 (5 - 6), 131 - 153." type="journal article" year="2016">Watters 2016b</bibRefCitation>
). However, the Kibasira Swamp is a vast entity and undoubtedly the distribution of this species is significantly greater than has presently been reported and is expected to also extend throughout the less accessible southwestern part of the Swamp that is yet to be fully investigated (
<figureCitation id="13732A5BFF9DDD44FDFD87D134D13AB7" box="[631,696,1195,1221]" captionStart="FIGURE 1" captionStartId="2.[151,250,1762,1786]" captionTargetBox="[151,1436,374,1736]" captionTargetId="figure-118@2.[151,1436,374,1736]" captionTargetPageId="2" captionText="FIGURE 1. Distribution of species belonging to the subgenus Aphyobranchius, all of which occur in the east-central and southern coastal lowlands of Tanzania: N. janpapi (yellow-filled circle), N. luekei (red-filled triangle), N. geminus (orange-filled diamond), N. lourensi (green-filled square), N. fuscotaeniatus (blue-filled inverted triangle), N. cf. janpapi (dark magenta-filled circle). The inset map shows the area of distribution in a somewhat broader geographic context, to include the distribution of N. willerti (red-filled star) in the lower Tana River area of northeastern coastal Kenya, a species that herein is considered to belong to the subgenus Adiniops, rather than Aphyobranchius to which it was ascribed in the original description by Wildekamp (1992). Note that individual symbols on the map may represent more than one locality or population where the sites are close to one another and could not reasonably be resolved at the scale of the map." pageId="17" pageNumber="418">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8BF736DEFF9DDD44FF4D87B433FD385F" blockId="17.[151,1437,151,1581]" pageId="17" pageNumber="418">
<emphasis id="B93CEACCFF9DDD44FF4D87B437D53A9B" bold="true" box="[199,444,1230,1257]" pageId="17" pageNumber="418">Conservation status.</emphasis>
<taxonomicName id="4C484D5DFF9DDD44FE4987B534B53A9A" authorityName="TSTS" authorityYear="2010" box="[451,732,1231,1257]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="17" pageNumber="418" phylum="Chordata" rank="species" species="geminus">
<emphasis id="B93CEACCFF9DDD44FE4987B534B53A9A" box="[451,732,1231,1257]" italics="true" pageId="17" pageNumber="418">Nothobranchius geminus</emphasis>
</taxonomicName>
is listed as Vulnerable B1ab(iii) (
<bibRefCitation id="EFD94B2FFF9DDD44FBDC87B533373A9B" author="Nagy, B. &amp; Watters, B." box="[1110,1374,1231,1257]" pageId="17" pageNumber="418" refId="ref33700" refString="Nagy, B. &amp; Watters, B. (2019 c) Nothobranchius geminus. The IUCN Red List of Threatened Species, 2019, e. T 61296 A 47244197. https: // doi. org / 10.2305 / IUCN. UK. 2019 - 3. RLTS. T 61296 A 47244197. en" type="journal volume" year="2019">Nagy &amp; Watters 2019c</bibRefCitation>
). The known extent of occurrence (EOO) is
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², with a maximum estimate of around
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². Within its restricted area of distribution, the occurrence of the species is fragmented and currently it is known from only five threat-based locations. However, it is expected to occur at up to 10 threat-based locations within the drainage of the Kilombero River. This species is expected to also inhabit the southwestern parts of the Kibasira Swamp that are yet to be fully investigated; all presently known sites are in the northeastern parts of the swamp. In combination with its annual life cycle in seasonal habitats, its dependence on the clay-rich vertisol-type substratum for embryonic development makes it more subject to threats, as is the case with all other members of the subgenus. This category is assigned based on the upper estimates of the EOO and number of threat-based locations as best estimates. Notably, drainage modifications associated with road construction has resulted in the destruction of the
<typeStatus id="54F3887CFF9DDD44FBD8856932EB385F" box="[1106,1154,1555,1581]" pageId="17" pageNumber="418">type</typeStatus>
locality for this species.
</paragraph>
</subSubSection>
</treatment>
</document>

665
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<emphasis id="B93CEACCFF86DD5FFF1D86B737C43B9A" bold="true" box="[151,429,1485,1512]" italics="true" pageId="10" pageNumber="411">Nothobranchius luekei</emphasis>
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(
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)
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<emphasis id="B93CEACCFF86DD5FFF1D854337EA3821" bold="true" box="[151,387,1593,1619]" pageId="10" pageNumber="411">Material examined.</emphasis>
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,
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SL,
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SL;
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;
<date id="FFF6101EFF86DD5FFCFD8524327A380A" box="[887,1043,1630,1656]" pageId="10" pageNumber="411" value="2000-06-16">
<collectingDate id="EFB2E9F6FF86DD5FFCFD8524327A380A" box="[887,1043,1630,1656]" pageId="10" pageNumber="411" value="2000-06-16">16 June 2000</collectingDate>
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; field code: TAN 00-23
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; –
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SL,
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SL;
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<geoCoordinate id="EE7C5019FF86DD5FFB5785F8331C38EE" box="[1245,1397,1666,1692]" degrees="39" direction="east" minutes="10.299" orientation="longitude" pageId="10" pageNumber="411" precision="1" value="39.17165">39°10.299’ E</geoCoordinate>
;
<collectorName id="26BD5308FF86DD5FFA0985F836BE38B2" pageId="10" pageNumber="411">B. Nagy</collectorName>
;
<date id="FFF6101EFF86DD5FFF6985DC37F638B2" box="[227,415,1702,1728]" pageId="10" pageNumber="411" value="2009-01-13">
<collectingDate id="EFB2E9F6FF86DD5FFF6985DC37F638B2" box="[227,415,1702,1728]" pageId="10" pageNumber="411" value="2009-01-13">13 January 2009</collectingDate>
</date>
; field code: TZN 09-9
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3526555FF86DD5EFF4D85B3333E3C6B" lastPageId="11" lastPageNumber="412" pageId="10" pageNumber="411" type="diagnosis">
<paragraph id="8BF736DEFF86DD5EFF4D85B3333E3C6B" blockId="10.[151,1437,1593,2016]" lastBlockId="11.[151,1437,151,537]" lastPageId="11" lastPageNumber="412" pageId="10" pageNumber="411">
<emphasis id="B93CEACCFF86DD5FFF4D85B3372B3891" bold="true" box="[199,322,1737,1763]" pageId="10" pageNumber="411">Diagnosis.</emphasis>
<taxonomicName id="4C484D5DFF86DD5FFECD85B0342D3896" authorityName="Seegers" authorityYear="1984" box="[327,580,1738,1764]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="411" phylum="Chordata" rank="species" species="luekei">
<emphasis id="B93CEACCFF86DD5FFECD85B0342D3896" box="[327,580,1738,1764]" italics="true" pageId="10" pageNumber="411">Nothobranchius luekei</emphasis>
</taxonomicName>
belongs to the
<taxonomicName id="4C484D5DFF86DD5FFD6085B035363891" box="[746,863,1738,1763]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="411" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF86DD5FFD6085B035363891" box="[746,863,1738,1763]" italics="true" pageId="10" pageNumber="411">N. janpapi</emphasis>
</taxonomicName>
species group within the subgenus
<taxonomicName id="4C484D5DFF86DD5FFB6F85B033F43896" authorityName="Wildekamp" authorityYear="1977" box="[1253,1437,1738,1764]" class="Actinopterygii" family="Nothobranchiidae" genus="Aphyobranchius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="411" phylum="Chordata" rank="genus">
<emphasis id="B93CEACCFF86DD5FFB6F85B033F43896" box="[1253,1437,1738,1764]" italics="true" pageId="10" pageNumber="411">Aphyobranchius</emphasis>
</taxonomicName>
by presenting the following combination of characters in the male: light blue body colouration with narrow red scale margins and yellow-brown to yellow-green dorsum stripe; rearward positioned dorsal fin with predorsal length of 68–73.0 % SL and predorsal length 117–122 as % of preanal length; dorsal fin relatively small with low number of rays, 11–13; cephalic sensory system at supraorbital level in an undivided, slightly curved, almost flat, shallow groove with three exposed neuromasts and pronounced lobes (
<figureCitation id="13732A5BFF86DD5FFC378404327B39EA" box="[957,1042,1918,1944]" captionStart="FIGURE 3" captionStartId="10.[151,250,1307,1331]" captionTargetBox="[156,1428,706,1276]" captionTargetId="figure-280@10.[151,1436,702,1281]" captionTargetPageId="10" captionText="FIGURE 3. Cephalic structure: a, Nothobranchius janpapi; b, N. luekei; c, N. geminus; d, N. lourensi and N. fuscotaeniatus. Key to label abbreviations: an, anterior nostril; pn, posterior nostril; f, frontal neuromast; so, supraorbital canal; st, supratemporal canal." pageId="10" pageNumber="411">Fig. 3b</figureCitation>
). Both sexes with central 2–3 rays of the caudal fin slightly elongated forming a posterior lobe (vs. central lobe absent).
<taxonomicName id="4C484D5DFF86DD5FFBF484D833E939CE" authorityName="Seegers" authorityYear="1984" box="[1150,1408,1954,1980]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="411" phylum="Chordata" rank="species" species="luekei">
<emphasis id="B93CEACCFF86DD5FFBF484D833E939CE" box="[1150,1408,1954,1980]" italics="true" pageId="10" pageNumber="411">Nothobranchius luekei</emphasis>
</taxonomicName>
is distinguished from the other members of the
<taxonomicName id="4C484D5DFF86DD5FFD1884BC356139AD" box="[658,776,1990,2015]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="411" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF86DD5FFD1884BC356139AD" box="[658,776,1990,2015]" italics="true" pageId="10" pageNumber="411">N. janpapi</emphasis>
</taxonomicName>
species group by male colouration of anal fin with slender yellow subdistal band and narrow red margin (vs. slender orange-red distal band with narrow light blue margin in
<taxonomicName id="4C484D5DFF87DD5EFF1D83C137663EA6" box="[151,271,187,212]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF87DD5EFF1D83C137663EA6" box="[151,271,187,212]" italics="true" pageId="11" pageNumber="412">N. janpapi</emphasis>
</taxonomicName>
); caudal fin base and proximal zone grey to light red with slender red subdistal band and narrow black distal band (vs. base and proximal zone olive, and with light grey to translucent subdistal band and narrow dark grey distal band in
<taxonomicName id="4C484D5DFF87DD5EFEB8827937CE3F6E" box="[306,423,259,284]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF87DD5EFEB8827937CE3F6E" box="[306,423,259,284]" italics="true" pageId="11" pageNumber="412">N. janpapi</emphasis>
</taxonomicName>
, and plain red to orange-red with narrow black distal margin in
<taxonomicName id="4C484D5DFF87DD5EFBEC8279328B3F6E" authorityName="Wildekamp, Watters and Sainthouse" authorityYear="2002" box="[1126,1250,259,284]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="geminus">
<emphasis id="B93CEACCFF87DD5EFBEC8279328B3F6E" box="[1126,1250,259,284]" italics="true" pageId="11" pageNumber="412">N. geminus</emphasis>
</taxonomicName>
); pelvic fin light blue with yellow subdistal band and narrow red distal band (vs. plain orange-red with narrow light blue distal band in
<taxonomicName id="4C484D5DFF87DD5EFF39823137403F16" box="[179,297,331,356]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF87DD5EFF39823137403F16" box="[179,297,331,356]" italics="true" pageId="11" pageNumber="412">N. janpapi</emphasis>
</taxonomicName>
, and yellow to red with narrow black distal band in
<taxonomicName id="4C484D5DFF87DD5EFCEC8231358B3F16" authorityName="Wildekamp, Watters and Sainthouse" authorityYear="2002" box="[870,994,331,356]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="geminus">
<emphasis id="B93CEACCFF87DD5EFCEC8231358B3F16" box="[870,994,331,356]" italics="true" pageId="11" pageNumber="412">N. geminus</emphasis>
</taxonomicName>
); as well as morphometric characters in males of body depth at pelvic-fin origin 21.5–23.6 % SL (vs.
<quantity id="4CB09B3BFF87DD5EFCB5821435A23FFB" box="[831,971,366,393]" metricMagnitude="-1" metricUnit="m" metricValue="7.0485" metricValueMax="7.6708" metricValueMin="6.4262" pageId="11" pageNumber="412" unit="in" value="27.75" valueMax="30.2" valueMin="25.3">25.3–30.2 in</quantity>
<taxonomicName id="4C484D5DFF87DD5EFC5B821532273FFA" authorityName="Wildekamp, Watters and Sainthouse" authorityYear="2002" box="[977,1102,367,392]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="geminus">
<emphasis id="B93CEACCFF87DD5EFC5B821532273FFA" box="[977,1102,367,392]" italics="true" pageId="11" pageNumber="412">N. geminus</emphasis>
</taxonomicName>
and 27.4–31.0 in
<taxonomicName id="4C484D5DFF87DD5EFA9C821533E53FFA" box="[1302,1420,367,392]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF87DD5EFA9C821533E53FFA" box="[1302,1420,367,392]" italics="true" pageId="11" pageNumber="412">N. janpapi</emphasis>
</taxonomicName>
); head depth 55–63% HL (vs.
<quantity id="4CB09B3BFF87DD5EFE5F82E934513FDF" box="[469,568,403,429]" metricMagnitude="0" metricUnit="m" metricValue="1.7526" metricValueMax="1.8288" metricValueMin="1.6764" pageId="11" pageNumber="412" unit="in" value="69.0" valueMax="72.0" valueMin="66.0">66–72 in</quantity>
<taxonomicName id="4C484D5DFF87DD5EFDB482E934D23FDE" authorityName="Wildekamp, Watters and Sainthouse" authorityYear="2002" box="[574,699,403,428]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="geminus">
<emphasis id="B93CEACCFF87DD5EFDB482E934D23FDE" box="[574,699,403,428]" italics="true" pageId="11" pageNumber="412">N. geminus</emphasis>
</taxonomicName>
and
<quantity id="4CB09B3BFF87DD5EFD7B82E9353A3FDF" box="[753,851,403,429]" metricMagnitude="0" metricUnit="m" metricValue="1.9685" metricValueMax="2.0574" metricValueMin="1.8796" pageId="11" pageNumber="412" unit="in" value="77.5" valueMax="81.0" valueMin="74.0">74–81 in</quantity>
<taxonomicName id="4C484D5DFF87DD5EFCD382E935A63FDE" box="[857,975,403,428]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF87DD5EFCD382E935A63FDE" box="[857,975,403,428]" italics="true" pageId="11" pageNumber="412">N. janpapi</emphasis>
</taxonomicName>
); head width 87–97 as % of its depth (vs.
<quantity id="4CB09B3BFF87DD5EFF1D82CD36933FA3" box="[151,250,439,465]" metricMagnitude="0" metricUnit="m" metricValue="1.9812" metricValueMax="2.1082" metricValueMin="1.8542" pageId="11" pageNumber="412" unit="in" value="78.0" valueMax="83.0" valueMin="73.0">73–83 in</quantity>
<taxonomicName id="4C484D5DFF87DD5EFE8A82CD37173FA2" authorityName="Wildekamp, Watters and Sainthouse" authorityYear="2002" box="[256,382,439,464]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="geminus">
<emphasis id="B93CEACCFF87DD5EFE8A82CD37173FA2" box="[256,382,439,464]" italics="true" pageId="11" pageNumber="412">N. geminus</emphasis>
</taxonomicName>
and
<quantity id="4CB09B3BFF87DD5EFE3E82CD347F3FA3" box="[436,534,439,465]" metricMagnitude="0" metricUnit="m" metricValue="1.9685" metricValueMax="2.0574" metricValueMin="1.8796" pageId="11" pageNumber="412" unit="in" value="77.5" valueMax="81.0" valueMin="74.0">74–81 in</quantity>
<taxonomicName id="4C484D5DFF87DD5EFD9782CD34FA3FA2" box="[541,659,439,464]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF87DD5EFD9782CD34FA3FA2" box="[541,659,439,464]" italics="true" pageId="11" pageNumber="412">N. janpapi</emphasis>
</taxonomicName>
); and caudal peduncle length 148–156 as % of its depth (vs.
<quantity id="4CB09B3BFF87DD5EFAB082CD36C43F87" metricMagnitude="0" metricUnit="m" metricValue="3.1877" metricValueMax="3.3019999999999996" metricValueMin="3.0734" pageId="11" pageNumber="412" unit="in" value="125.5" valueMax="130.0" valueMin="121.0">121–130 in</quantity>
<taxonomicName id="4C484D5DFF87DD5EFF3F82A1375A3F86" authorityName="Wildekamp, Watters and Sainthouse" authorityYear="2002" box="[181,307,475,500]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="geminus">
<emphasis id="B93CEACCFF87DD5EFF3F82A1375A3F86" box="[181,307,475,500]" italics="true" pageId="11" pageNumber="412">N. geminus</emphasis>
</taxonomicName>
and
<quantity id="4CB09B3BFF87DD5EFEE182A137823F87" box="[363,491,475,501]" metricMagnitude="0" metricUnit="m" metricValue="3.4671" metricValueMax="3.556" metricValueMin="3.3781999999999996" pageId="11" pageNumber="412" unit="in" value="136.5" valueMax="140.0" valueMin="133.0">133–140 in</quantity>
<taxonomicName id="4C484D5DFF87DD5EFE7882A134003F86" box="[498,617,475,500]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF87DD5EFE7882A134003F86" box="[498,617,475,500]" italics="true" pageId="11" pageNumber="412">N. janpapi</emphasis>
</taxonomicName>
). Further, it differs from
<taxonomicName id="4C484D5DFF87DD5EFC0C82A135973F86" box="[902,1022,475,500]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF87DD5EFC0C82A135973F86" box="[902,1022,475,500]" italics="true" pageId="11" pageNumber="412">N. janpapi</emphasis>
</taxonomicName>
by predorsal length 68.0–73.0 % SL (vs. 63.3–67.9 % SL); head width 52–56 % HL (vs. 58–65 %); and suborbital depth 8–12 % HL (vs. 16–20).
</paragraph>
</subSubSection>
<caption id="DF376656FF87DD5EFF1D861232B83863" pageId="11" pageNumber="412" startId="11.[151,250,1384,1408]" targetBox="[151,1435,558,1358]" targetPageId="11" targetType="figure">
<paragraph id="8BF736DEFF87DD5EFF1D861232B83863" blockId="11.[151,1437,1384,1553]" pageId="11" pageNumber="412">
<emphasis id="B93CEACCFF87DD5EFF1D8612377D3BF3" bold="true" box="[151,276,1384,1409]" pageId="11" pageNumber="412">FIGURE 4.</emphasis>
Wild-caught specimens of
<taxonomicName id="4C484D5DFF87DD5EFDA78613357D3BF3" authorityName="Seegers" authorityYear="1984" box="[557,788,1385,1409]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="luekei">
<emphasis id="B93CEACCFF87DD5EFDA78613357D3BF3" box="[557,788,1385,1409]" italics="true" pageId="11" pageNumber="412">Nothobranchius luekei</emphasis>
</taxonomicName>
and example of typical habitat.
<emphasis id="B93CEACCFF87DD5EFBEA861232103BF2" bold="true" box="[1120,1145,1384,1408]" pageId="11" pageNumber="412">A.</emphasis>
<taxonomicName id="4C484D5DFF87DD5EFB0A861333F53BF3" authority="Mbezi River TAN" authorityName="Mbezi River TAN" box="[1152,1436,1385,1409]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="luekei">
<emphasis id="B93CEACCFF87DD5EFB0A861332B23BF3" box="[1152,1243,1385,1409]" italics="true" pageId="11" pageNumber="412">N. luekei</emphasis>
Mbezi River TAN
</taxonomicName>
00-23, male, Mbezi River drainage, in immediate vicinity of the type locality;
<emphasis id="B93CEACCFF87DD5EFC5986F735833BD7" bold="true" box="[979,1002,1421,1445]" pageId="11" pageNumber="412">B.</emphasis>
<taxonomicName id="4C484D5DFF87DD5EFC7E86F733773BD7" authority="Luhule River TAN" authorityName="Luhule River TAN" box="[1012,1310,1421,1445]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="luekei">
<emphasis id="B93CEACCFF87DD5EFC7E86F732393BD7" box="[1012,1104,1421,1445]" italics="true" pageId="11" pageNumber="412">N. luekei</emphasis>
Luhule River TAN
</taxonomicName>
02-8, male, Luhule/Luhute River drainage;
<emphasis id="B93CEACCFF87DD5EFE5F86CA37873BBA" bold="true" box="[469,494,1456,1480]" pageId="11" pageNumber="412">C.</emphasis>
<taxonomicName id="4C484D5DFF87DD5EFE7E86CB35613BBA" authority="Mbezi River TZN" authorityName="Mbezi River TZN" box="[500,776,1457,1481]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="luekei">
<emphasis id="B93CEACCFF87DD5EFE7E86CB34243BBB" box="[500,589,1457,1481]" italics="true" pageId="11" pageNumber="412">N. luekei</emphasis>
Mbezi River TZN
</taxonomicName>
09-9, female, Mbezi River drainage (photograph by B. Nagy);
<emphasis id="B93CEACCFF87DD5EFA0E86CB33F53BBB" bold="true" box="[1412,1436,1457,1481]" pageId="11" pageNumber="412">D.</emphasis>
Location Luhule River TAN 02-8, a typical heavily vegetated shallow habitat (photograph June 2, 2002), hosting
<taxonomicName id="4C484D5DFF87DD5EFA9786AF331E3B9F" box="[1309,1399,1493,1517]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="luekei">
<emphasis id="B93CEACCFF87DD5EFA9786AF331E3B9F" box="[1309,1399,1493,1517]" italics="true" pageId="11" pageNumber="412">N. luekei</emphasis>
</taxonomicName>
,
<taxonomicName id="4C484D5DFF87DD5EFA0F86AF36BD3863" authorityName="Wildekamp, Shidlovskiy &amp; Watters" authorityYear="2009" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="lucius">
<emphasis id="B93CEACCFF87DD5EFA0F86AF36BD3863" italics="true" pageId="11" pageNumber="412">N. lucius</emphasis>
</taxonomicName>
,
<taxonomicName id="4C484D5DFF87DD5EFF6A868337193863" authorityName="Seegers" authorityYear="1986" box="[224,368,1529,1553]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="rubripinnis">
<emphasis id="B93CEACCFF87DD5EFF6A868337193863" box="[224,368,1529,1553]" italics="true" pageId="11" pageNumber="412">N. rubripinnis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C484D5DFF87DD5EFE2B8683340D3863" authorityName="Costa" authorityYear="2009" box="[417,612,1529,1553]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="ruudwildekampi">
<emphasis id="B93CEACCFF87DD5EFE2B8683340D3863" box="[417,612,1529,1553]" italics="true" pageId="11" pageNumber="412">N. ruudwildekampi</emphasis>
</taxonomicName>
. Photographs by B.R. Watters except where otherwise noted.
</paragraph>
</caption>
<subSubSection id="C3526555FF87DD58FF4D854533E938C3" lastPageId="13" lastPageNumber="414" pageId="11" pageNumber="412" type="description">
<paragraph id="8BF736DEFF87DD5EFF4D8545341D380F" blockId="11.[151,1437,1599,2021]" pageId="11" pageNumber="412">
<emphasis id="B93CEACCFF87DD5EFF4D85453730382B" bold="true" box="[199,345,1599,1625]" pageId="11" pageNumber="412">Description.</emphasis>
General body features illustrated by
<figureCitation id="13732A5BFF87DD5EFC9A854535DC382B" box="[784,949,1599,1625]" captionStart="FIGURE 4" captionStartId="11.[151,250,1384,1408]" captionTargetBox="[151,1435,558,1358]" captionTargetId="figure-251@11.[151,1435,558,1358]" captionTargetPageId="11" captionText="FIGURE 4. Wild-caught specimens of Nothobranchius luekei and example of typical habitat. A. N. luekei Mbezi River TAN 00-23, male, Mbezi River drainage, in immediate vicinity of the type locality; B. N. luekei Luhule River TAN 02-8, male, Luhule/Luhute River drainage; C. N. luekei Mbezi River TZN 09-9, female, Mbezi River drainage (photograph by B. Nagy); D. Location Luhule River TAN 02-8, a typical heavily vegetated shallow habitat (photograph June 2, 2002), hosting N. luekei, N. lucius, N. rubripinnis and N. ruudwildekampi. Photographs by B.R. Watters except where otherwise noted." pageId="11" pageNumber="412">Figures 4A–C</figureCitation>
. Morphometric and meristic characters of examined material summarized in
<tableCitation id="C6CA0365FF87DD5EFD9085193419380F" box="[538,624,1635,1661]" captionStart="TABLE 4" captionStartId="12.[151,239,260,284]" captionText="TABLE 4. Morphometric and meristic data of Nothobranchius luekei. SD, standard deviation." pageId="11" pageNumber="412">Table 4</tableCitation>
.
</paragraph>
<paragraph id="8BF736DEFF87DD5EFF4D85FD35693927" blockId="11.[151,1437,1599,2021]" pageId="11" pageNumber="412">
Male: Small
<taxonomicName id="4C484D5DFF87DD5EFEDD85FD340F38D3" box="[343,614,1671,1697]" pageId="11" pageNumber="412">
<emphasis id="B93CEACCFF87DD5EFEDD85FD346238D3" box="[343,523,1671,1697]" italics="true" pageId="11" pageNumber="412">Nothobranchius</emphasis>
species
</taxonomicName>
, maximum size of examined specimens
<quantity id="4CB09B3BFF87DD5EFBAD85FD32E338D3" box="[1063,1162,1671,1697]" metricMagnitude="-2" metricUnit="m" metricValue="2.62" pageId="11" pageNumber="412" unit="mm" value="26.2">26.2 mm</quantity>
SL. General body shape laterally compressed. Greatest body depth at vertical in front of pelvic-fin origin (21.5–23.6 % SL). Greatest body width at pectoral-fin base with body progressively narrowing towards caudal-fin base. Dorsal profile straight from tip of snout to dorsal-fin origin, slightly concave on caudal peduncle. Ventral profile convex from lower jaw to base of last anal-fin ray, straight to slightly concave on caudal peduncle. Caudal peduncle deep, length 148–156 % of its depth. Anus situated directly in front of anal-fin origin.
</paragraph>
<paragraph id="8BF736DEFF87DD59FF4D842535F93EA7" blockId="11.[151,1437,1599,2021]" lastBlockId="12.[151,1436,150,213]" lastPageId="12" lastPageNumber="413" pageId="11" pageNumber="412">Head moderately long (27.1–31.1 % SL), laterally compressed, slightly deeper than wide (head width 87–97 % of its depth). Snout distinctly pointed, smaller than eye diameter. Mouth supraterminal, slightly oblique in profile. Jaws subequal, lower jaw longer than upper, posterior end of rictus at same level or slightly ventral to centre of eye. Premaxilla and dentary with many irregularly distributed conical teeth on inner rows and slightly curved teeth on outer row. Orbit large, eye diameter 32–37 % HL, mainly in anterior half of head (snout to eye end length 52–58 % HL). Branchiostegal membrane projecting posteriorly from opercle.</paragraph>
<caption id="DF376656FF80DD59FF1D827E32EF3F6E" box="[151,1158,260,285]" pageId="12" pageNumber="413" startId="12.[151,239,260,284]" targetType="table">
<paragraph id="8BF736DEFF80DD59FF1D827E32EF3F6E" blockId="12.[151,1158,260,285]" box="[151,1158,260,285]" pageId="12" pageNumber="413">
<emphasis id="B93CEACCFF80DD59FF1D827E37633F6E" bold="true" box="[151,266,260,284]" pageId="12" pageNumber="413">TABLE 4.</emphasis>
Morphometric and meristic data of
<taxonomicName id="4C484D5DFF80DD59FD01827E35173F6E" box="[651,894,260,284]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="luekei">
<emphasis id="B93CEACCFF80DD59FD01827E35173F6E" box="[651,894,260,284]" italics="true" pageId="12" pageNumber="413">Nothobranchius luekei</emphasis>
</taxonomicName>
. SD, standard deviation.
</paragraph>
</caption>
<paragraph id="8BF736DEFF80DD59FC948257328F389C" pageId="12" pageNumber="413">
<table id="F948C47EFF8022AAFF298257330D389C" box="[163,1380,301,1774]" colsContinueIn="22.[153,1394,192,1665]" gridcols="7" gridrows="36" pageId="12" pageNumber="413">
<tr id="3578349CFF8022AAFF298257330D3F37" box="[163,1380,301,325]" gridrow="0" pageId="12" pageNumber="413" rowspan-0="1" rowspan-3="1" rowspan-4="1" rowspan-6="1">
<th id="76A95DE0FF8022AAFD3B825735EC3F37" box="[689,901,301,325]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="12" pageNumber="413">males</th>
<th id="76A95DE0FF8022AAFB288257336F3F37" box="[1186,1286,301,325]" gridcol="5" gridrow="0" pageId="12" pageNumber="413">females</th>
</tr>
<tr id="3578349CFF8022AAFF29822F330D3F1F" box="[163,1380,341,365]" gridrow="1" pageId="12" pageNumber="413" rowspan-0="1" rowspan-3="1" rowspan-4="1" rowspan-6="1">
<td id="76A95DE0FF8022AAFD3B822F35EC3F1F" box="[689,901,341,365]" colspan="2" colspanRight="1" gridcol="1" gridrow="1" pageId="12" pageNumber="413">(n = 7)</td>
<td id="76A95DE0FF8022AAFB28822F336F3F1F" box="[1186,1286,341,365]" gridcol="5" gridrow="1" pageId="12" pageNumber="413">(n = 2)</td>
</tr>
<tr id="3578349CFF8022AAFF2982DC330D3FCC" box="[163,1380,422,446]" gridrow="2" pageId="12" pageNumber="413" rowspan-0="1">
<td id="76A95DE0FF8022AAFD3B82DC35593FCC" box="[689,816,422,446]" gridcol="1" gridrow="2" pageId="12" pageNumber="413">range</td>
<td id="76A95DE0FF8022AAFCC682DC35EC3FCC" box="[844,901,422,446]" gridcol="2" gridrow="2" pageId="12" pageNumber="413">mean</td>
<td id="76A95DE0FF8022AAFC2882DC35AB3FCC" box="[930,962,422,446]" gridcol="3" gridrow="2" pageId="12" pageNumber="413">SD</td>
<td id="76A95DE0FF8022AAFB8B82DC32E93FCC" box="[1025,1152,422,446]" gridcol="4" gridrow="2" pageId="12" pageNumber="413">range</td>
<td id="76A95DE0FF8022AAFB2882DC336F3FCC" box="[1186,1286,422,446]" gridcol="5" gridrow="2" pageId="12" pageNumber="413">mean</td>
<td id="76A95DE0FF8022AAFACE82DC330D3FCC" box="[1348,1380,422,446]" gridcol="6" gridrow="2" pageId="12" pageNumber="413">SD</td>
</tr>
<tr id="3578349CFF8022AAFF2982B4330D3F94" box="[163,1380,462,486]" gridrow="3" pageId="12" pageNumber="413" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1">
<th id="76A95DE0FF8022AAFF2982B434FF3F94" box="[163,662,462,486]" gridcol="0" gridrow="3" pageId="12" pageNumber="413">Standard length (mm)</th>
<td id="76A95DE0FF8022AAFD3B82B435593F94" box="[689,816,462,486]" gridcol="1" gridrow="3" pageId="12" pageNumber="413">21.2–26.2</td>
<td id="76A95DE0FF8022AAFB8B82B432E93F94" box="[1025,1152,462,486]" gridcol="4" gridrow="3" pageId="12" pageNumber="413">21.3–24.6</td>
</tr>
<tr id="3578349CFF8022AAFF29828F330D3C7C" box="[163,1380,501,526]" gridrow="4" pageId="12" pageNumber="413">
<th id="76A95DE0FF8022AAFF29828F330D3C7C" box="[163,1380,501,526]" colspan="7" colspanRight="6" gridcol="0" gridrow="4" pageId="12" pageNumber="413">
<emphasis id="B93CEACCFF80DD59FF29828C37AB3C7C" bold="true" box="[163,450,501,526]" pageId="12" pageNumber="413">Percent of standard length</emphasis>
</th>
</tr>
<tr id="3578349CFF8022AAFF298164330D3C44" box="[163,1380,542,566]" gridrow="5" pageId="12" pageNumber="413">
<th id="76A95DE0FF8022AAFF29816434FF3C44" box="[163,662,542,566]" gridcol="0" gridrow="5" pageId="12" pageNumber="413">Total length</th>
<td id="76A95DE0FF8022AAFD3B816435593C44" box="[689,816,542,566]" gridcol="1" gridrow="5" pageId="12" pageNumber="413">121.5–128.8</td>
<td id="76A95DE0FF8022AAFCC6816435EC3C44" box="[844,901,542,566]" gridcol="2" gridrow="5" pageId="12" pageNumber="413">125.6</td>
<td id="76A95DE0FF8022AAFC28816435AB3C44" box="[930,962,542,566]" gridcol="3" gridrow="5" pageId="12" pageNumber="413">2.4</td>
<td id="76A95DE0FF8022AAFB8B816432E93C44" box="[1025,1152,542,566]" gridcol="4" gridrow="5" pageId="12" pageNumber="413">124.8–128.6</td>
<td id="76A95DE0FF8022AAFB288164336F3C44" box="[1186,1286,542,566]" gridcol="5" gridrow="5" pageId="12" pageNumber="413">126.7</td>
<td id="76A95DE0FF8022AAFACE8164330D3C44" box="[1348,1380,542,566]" gridcol="6" gridrow="5" pageId="12" pageNumber="413">2.7</td>
</tr>
<tr id="3578349CFF8022AAFF29813C330D3C2C" box="[163,1380,582,606]" gridrow="6" pageId="12" pageNumber="413">
<th id="76A95DE0FF8022AAFF29813C34FF3C2C" box="[163,662,582,606]" gridcol="0" gridrow="6" pageId="12" pageNumber="413">Body depth at pelvic-fin origin</th>
<td id="76A95DE0FF8022AAFD3B813C35593C2C" box="[689,816,582,606]" gridcol="1" gridrow="6" pageId="12" pageNumber="413">21.5–23.6</td>
<td id="76A95DE0FF8022AAFCC6813C35EC3C2C" box="[844,901,582,606]" gridcol="2" gridrow="6" pageId="12" pageNumber="413">23.0</td>
<td id="76A95DE0FF8022AAFC28813C35AB3C2C" box="[930,962,582,606]" gridcol="3" gridrow="6" pageId="12" pageNumber="413">0.8</td>
<td id="76A95DE0FF8022AAFB8B813C32E93C2C" box="[1025,1152,582,606]" gridcol="4" gridrow="6" pageId="12" pageNumber="413">19.5–23.0</td>
<td id="76A95DE0FF8022AAFB28813C336F3C2C" box="[1186,1286,582,606]" gridcol="5" gridrow="6" pageId="12" pageNumber="413">21.3</td>
<td id="76A95DE0FF8022AAFACE813C330D3C2C" box="[1348,1380,582,606]" gridcol="6" gridrow="6" pageId="12" pageNumber="413">2.5</td>
</tr>
<tr id="3578349CFF8022AAFF298114330D3CF5" box="[163,1380,622,647]" gridrow="7" pageId="12" pageNumber="413">
<th id="76A95DE0FF8022AAFF29811434FF3CF5" box="[163,662,622,647]" gridcol="0" gridrow="7" pageId="12" pageNumber="413">Head length</th>
<td id="76A95DE0FF8022AAFD3B811435593CF5" box="[689,816,622,647]" gridcol="1" gridrow="7" pageId="12" pageNumber="413">27.1–31.1</td>
<td id="76A95DE0FF8022AAFCC6811435EC3CF5" box="[844,901,622,647]" gridcol="2" gridrow="7" pageId="12" pageNumber="413">29.3</td>
<td id="76A95DE0FF8022AAFC28811435AB3CF5" box="[930,962,622,647]" gridcol="3" gridrow="7" pageId="12" pageNumber="413">1.4</td>
<td id="76A95DE0FF8022AAFB8B811432E93CF5" box="[1025,1152,622,647]" gridcol="4" gridrow="7" pageId="12" pageNumber="413">25.6–29.6</td>
<td id="76A95DE0FF8022AAFB288114336F3CF5" box="[1186,1286,622,647]" gridcol="5" gridrow="7" pageId="12" pageNumber="413">27.6</td>
<td id="76A95DE0FF8022AAFACE8114330D3CF5" box="[1348,1380,622,647]" gridcol="6" gridrow="7" pageId="12" pageNumber="413">2.8</td>
</tr>
<tr id="3578349CFF8022AAFF2981ED330D3CDD" box="[163,1380,663,687]" gridrow="8" pageId="12" pageNumber="413">
<th id="76A95DE0FF8022AAFF2981ED34FF3CDD" box="[163,662,663,687]" gridcol="0" gridrow="8" pageId="12" pageNumber="413">Preanal length</th>
<td id="76A95DE0FF8022AAFD3B81ED35593CDD" box="[689,816,663,687]" gridcol="1" gridrow="8" pageId="12" pageNumber="413">55.7–60.5</td>
<td id="76A95DE0FF8022AAFCC681ED35EC3CDD" box="[844,901,663,687]" gridcol="2" gridrow="8" pageId="12" pageNumber="413">58.1</td>
<td id="76A95DE0FF8022AAFC2881ED35AB3CDD" box="[930,962,663,687]" gridcol="3" gridrow="8" pageId="12" pageNumber="413">1.4</td>
<td id="76A95DE0FF8022AAFB8B81ED32E93CDD" box="[1025,1152,663,687]" gridcol="4" gridrow="8" pageId="12" pageNumber="413">59.6–61.4</td>
<td id="76A95DE0FF8022AAFB2881ED336F3CDD" box="[1186,1286,663,687]" gridcol="5" gridrow="8" pageId="12" pageNumber="413">60.5</td>
<td id="76A95DE0FF8022AAFACE81ED330D3CDD" box="[1348,1380,663,687]" gridcol="6" gridrow="8" pageId="12" pageNumber="413">1.2</td>
</tr>
<tr id="3578349CFF8022AAFF2981C5330D3CA5" box="[163,1380,703,727]" gridrow="9" pageId="12" pageNumber="413">
<th id="76A95DE0FF8022AAFF2981C534FF3CA5" box="[163,662,703,727]" gridcol="0" gridrow="9" pageId="12" pageNumber="413">Predorsal length</th>
<td id="76A95DE0FF8022AAFD3B81C535593CA5" box="[689,816,703,727]" gridcol="1" gridrow="9" pageId="12" pageNumber="413">68.0–73.0</td>
<td id="76A95DE0FF8022AAFCC681C535EC3CA5" box="[844,901,703,727]" gridcol="2" gridrow="9" pageId="12" pageNumber="413">69.6</td>
<td id="76A95DE0FF8022AAFC2881C535AB3CA5" box="[930,962,703,727]" gridcol="3" gridrow="9" pageId="12" pageNumber="413">1.7</td>
<td id="76A95DE0FF8022AAFB8B81C532E93CA5" box="[1025,1152,703,727]" gridcol="4" gridrow="9" pageId="12" pageNumber="413">69.5–69.9</td>
<td id="76A95DE0FF8022AAFB2881C5336F3CA5" box="[1186,1286,703,727]" gridcol="5" gridrow="9" pageId="12" pageNumber="413">69.7</td>
<td id="76A95DE0FF8022AAFACE81C5330D3CA5" box="[1348,1380,703,727]" gridcol="6" gridrow="9" pageId="12" pageNumber="413">0.3</td>
</tr>
<tr id="3578349CFF8022AAFF29819D330D3C8D" box="[163,1380,743,767]" gridrow="10" pageId="12" pageNumber="413">
<th id="76A95DE0FF8022AAFF29819D34FF3C8D" box="[163,662,743,767]" gridcol="0" gridrow="10" pageId="12" pageNumber="413">Prepelvic length</th>
<td id="76A95DE0FF8022AAFD3B819D35593C8D" box="[689,816,743,767]" gridcol="1" gridrow="10" pageId="12" pageNumber="413">42.5–48.8</td>
<td id="76A95DE0FF8022AAFCC6819D35EC3C8D" box="[844,901,743,767]" gridcol="2" gridrow="10" pageId="12" pageNumber="413">45.8</td>
<td id="76A95DE0FF8022AAFC28819D35AB3C8D" box="[930,962,743,767]" gridcol="3" gridrow="10" pageId="12" pageNumber="413">2.0</td>
<td id="76A95DE0FF8022AAFB8B819D32E93C8D" box="[1025,1152,743,767]" gridcol="4" gridrow="10" pageId="12" pageNumber="413">46.9–48.0</td>
<td id="76A95DE0FF8022AAFB28819D336F3C8D" box="[1186,1286,743,767]" gridcol="5" gridrow="10" pageId="12" pageNumber="413">47.5</td>
<td id="76A95DE0FF8022AAFACE819D330D3C8D" box="[1348,1380,743,767]" gridcol="6" gridrow="10" pageId="12" pageNumber="413">0.7</td>
</tr>
<tr id="3578349CFF8022AAFF298075330D3D5A" box="[163,1380,783,808]" gridrow="11" pageId="12" pageNumber="413">
<th id="76A95DE0FF8022AAFF29807534FF3D5A" box="[163,662,783,808]" gridcol="0" gridrow="11" pageId="12" pageNumber="413">Prepectoral length</th>
<td id="76A95DE0FF8022AAFD3B807535593D5A" box="[689,816,783,808]" gridcol="1" gridrow="11" pageId="12" pageNumber="413">28.2–33.0</td>
<td id="76A95DE0FF8022AAFCC6807535EC3D5A" box="[844,901,783,808]" gridcol="2" gridrow="11" pageId="12" pageNumber="413">30.9</td>
<td id="76A95DE0FF8022AAFC28807535AB3D5A" box="[930,962,783,808]" gridcol="3" gridrow="11" pageId="12" pageNumber="413">1.7</td>
<td id="76A95DE0FF8022AAFB8B807532E93D5A" box="[1025,1152,783,808]" gridcol="4" gridrow="11" pageId="12" pageNumber="413">28.5–31.5</td>
<td id="76A95DE0FF8022AAFB288075336F3D5A" box="[1186,1286,783,808]" gridcol="5" gridrow="11" pageId="12" pageNumber="413">30.0</td>
<td id="76A95DE0FF8022AAFACE8075330D3D5A" box="[1348,1380,783,808]" gridcol="6" gridrow="11" pageId="12" pageNumber="413">2.1</td>
</tr>
<tr id="3578349CFF8022AAFF298042330D3D22" box="[163,1380,824,848]" gridrow="12" pageId="12" pageNumber="413">
<th id="76A95DE0FF8022AAFF29804234FF3D22" box="[163,662,824,848]" gridcol="0" gridrow="12" pageId="12" pageNumber="413">Caudal peduncle length</th>
<td id="76A95DE0FF8022AAFD3B804235593D22" box="[689,816,824,848]" gridcol="1" gridrow="12" pageId="12" pageNumber="413">17.7–20.1</td>
<td id="76A95DE0FF8022AAFCC6804235EC3D22" box="[844,901,824,848]" gridcol="2" gridrow="12" pageId="12" pageNumber="413">19.0</td>
<td id="76A95DE0FF8022AAFC28804235AB3D22" box="[930,962,824,848]" gridcol="3" gridrow="12" pageId="12" pageNumber="413">0.9</td>
<td id="76A95DE0FF8022AAFB8B804232E93D22" box="[1025,1152,824,848]" gridcol="4" gridrow="12" pageId="12" pageNumber="413">18.8–20.3</td>
<td id="76A95DE0FF8022AAFB288042336F3D22" box="[1186,1286,824,848]" gridcol="5" gridrow="12" pageId="12" pageNumber="413">19.6</td>
<td id="76A95DE0FF8022AAFACE8042330D3D22" box="[1348,1380,824,848]" gridcol="6" gridrow="12" pageId="12" pageNumber="413">1.1</td>
</tr>
<tr id="3578349CFF8022AAFF29801A330D3D0A" box="[163,1380,864,888]" gridrow="13" pageId="12" pageNumber="413">
<th id="76A95DE0FF8022AAFF29801A34FF3D0A" box="[163,662,864,888]" gridcol="0" gridrow="13" pageId="12" pageNumber="413">Caudal peduncle depth</th>
<td id="76A95DE0FF8022AAFD3B801A35593D0A" box="[689,816,864,888]" gridcol="1" gridrow="13" pageId="12" pageNumber="413">11.6–13.1</td>
<td id="76A95DE0FF8022AAFCC6801A35EC3D0A" box="[844,901,864,888]" gridcol="2" gridrow="13" pageId="12" pageNumber="413">12.5</td>
<td id="76A95DE0FF8022AAFC28801A35AB3D0A" box="[930,962,864,888]" gridcol="3" gridrow="13" pageId="12" pageNumber="413">0.6</td>
<td id="76A95DE0FF8022AAFB8B801A32E93D0A" box="[1025,1152,864,888]" gridcol="4" gridrow="13" pageId="12" pageNumber="413">11.3–12.6</td>
<td id="76A95DE0FF8022AAFB28801A336F3D0A" box="[1186,1286,864,888]" gridcol="5" gridrow="13" pageId="12" pageNumber="413">11.9</td>
<td id="76A95DE0FF8022AAFACE801A330D3D0A" box="[1348,1380,864,888]" gridcol="6" gridrow="13" pageId="12" pageNumber="413">0.9</td>
</tr>
<tr id="3578349CFF8022AAFF2980F2330D3DD2" box="[163,1380,904,928]" gridrow="14" pageId="12" pageNumber="413">
<th id="76A95DE0FF8022AAFF2980F234FF3DD2" box="[163,662,904,928]" gridcol="0" gridrow="14" pageId="12" pageNumber="413">Dorsal-fin base length</th>
<td id="76A95DE0FF8022AAFD3B80F235593DD2" box="[689,816,904,928]" gridcol="1" gridrow="14" pageId="12" pageNumber="413">16.3–19.8</td>
<td id="76A95DE0FF8022AAFCC680F235EC3DD2" box="[844,901,904,928]" gridcol="2" gridrow="14" pageId="12" pageNumber="413">18.0</td>
<td id="76A95DE0FF8022AAFC2880F235AB3DD2" box="[930,962,904,928]" gridcol="3" gridrow="14" pageId="12" pageNumber="413">1.1</td>
<td id="76A95DE0FF8022AAFB8B80F232E93DD2" box="[1025,1152,904,928]" gridcol="4" gridrow="14" pageId="12" pageNumber="413">15.0–16.9</td>
<td id="76A95DE0FF8022AAFB2880F2336F3DD2" box="[1186,1286,904,928]" gridcol="5" gridrow="14" pageId="12" pageNumber="413">16.0</td>
<td id="76A95DE0FF8022AAFACE80F2330D3DD2" box="[1348,1380,904,928]" gridcol="6" gridrow="14" pageId="12" pageNumber="413">1.3</td>
</tr>
<tr id="3578349CFF8022AAFF2980CA330D3DBB" box="[163,1380,944,969]" gridrow="15" pageId="12" pageNumber="413">
<th id="76A95DE0FF8022AAFF2980CA34FF3DBB" box="[163,662,944,969]" gridcol="0" gridrow="15" pageId="12" pageNumber="413">Anal-fin base length</th>
<td id="76A95DE0FF8022AAFD3B80CA35593DBB" box="[689,816,944,969]" gridcol="1" gridrow="15" pageId="12" pageNumber="413">21.9–26.3</td>
<td id="76A95DE0FF8022AAFCC680CA35EC3DBB" box="[844,901,944,969]" gridcol="2" gridrow="15" pageId="12" pageNumber="413">24.4</td>
<td id="76A95DE0FF8022AAFC2880CA35AB3DBB" box="[930,962,944,969]" gridcol="3" gridrow="15" pageId="12" pageNumber="413">1.6</td>
<td id="76A95DE0FF8022AAFB8B80CA32E93DBB" box="[1025,1152,944,969]" gridcol="4" gridrow="15" pageId="12" pageNumber="413">19.7–20.3</td>
<td id="76A95DE0FF8022AAFB2880CA336F3DBB" box="[1186,1286,944,969]" gridcol="5" gridrow="15" pageId="12" pageNumber="413">20.0</td>
<td id="76A95DE0FF8022AAFACE80CA330D3DBB" box="[1348,1380,944,969]" gridcol="6" gridrow="15" pageId="12" pageNumber="413">0.4</td>
</tr>
<tr id="3578349CFF8022AAFF2980A3330D3D83" box="[163,1380,985,1009]" gridrow="16" pageId="12" pageNumber="413">
<th id="76A95DE0FF8022AAFF2980A334FF3D83" box="[163,662,985,1009]" gridcol="0" gridrow="16" pageId="12" pageNumber="413">Caudal-fin length</th>
<td id="76A95DE0FF8022AAFD3B80A335593D83" box="[689,816,985,1009]" gridcol="1" gridrow="16" pageId="12" pageNumber="413">21.5–28.8</td>
<td id="76A95DE0FF8022AAFCC680A335EC3D83" box="[844,901,985,1009]" gridcol="2" gridrow="16" pageId="12" pageNumber="413">25.6</td>
<td id="76A95DE0FF8022AAFC2880A335AB3D83" box="[930,962,985,1009]" gridcol="3" gridrow="16" pageId="12" pageNumber="413">2.4</td>
<td id="76A95DE0FF8022AAFB8B80A332E93D83" box="[1025,1152,985,1009]" gridcol="4" gridrow="16" pageId="12" pageNumber="413">24.4–29.1</td>
<td id="76A95DE0FF8022AAFB2880A3336F3D83" box="[1186,1286,985,1009]" gridcol="5" gridrow="16" pageId="12" pageNumber="413">26.7</td>
<td id="76A95DE0FF8022AAFACE80A3330D3D83" box="[1348,1380,985,1009]" gridcol="6" gridrow="16" pageId="12" pageNumber="413">3.3</td>
</tr>
<tr id="3578349CFF8022AAFF29877B330D3A6B" box="[163,1380,1025,1049]" gridrow="17" pageId="12" pageNumber="413">
<th id="76A95DE0FF8022AAFF29877B330D3A6B" box="[163,1380,1025,1049]" colspan="7" colspanRight="6" gridcol="0" gridrow="17" pageId="12" pageNumber="413">
<emphasis id="B93CEACCFF80DD59FF29877B37FC3A6B" bold="true" box="[163,405,1025,1049]" pageId="12" pageNumber="413">Percent of head length</emphasis>
</th>
</tr>
<tr id="3578349CFF8022AAFF298753330D3A33" box="[163,1380,1065,1089]" gridrow="18" pageId="12" pageNumber="413">
<th id="76A95DE0FF8022AAFF29875334FF3A33" box="[163,662,1065,1089]" gridcol="0" gridrow="18" pageId="12" pageNumber="413">Head width</th>
<td id="76A95DE0FF8022AAFD3B875335593A33" box="[689,816,1065,1089]" gridcol="1" gridrow="18" pageId="12" pageNumber="413">52–56</td>
<td id="76A95DE0FF8022AAFCC6875335EC3A33" box="[844,901,1065,1089]" gridcol="2" gridrow="18" pageId="12" pageNumber="413">54.0</td>
<td id="76A95DE0FF8022AAFC28875335AB3A33" box="[930,962,1065,1089]" gridcol="3" gridrow="18" pageId="12" pageNumber="413">1.4</td>
<td id="76A95DE0FF8022AAFB8B875332E93A33" box="[1025,1152,1065,1089]" gridcol="4" gridrow="18" pageId="12" pageNumber="413">51–54</td>
<td id="76A95DE0FF8022AAFB288753336F3A33" box="[1186,1286,1065,1089]" gridcol="5" gridrow="18" pageId="12" pageNumber="413">52.4</td>
<td id="76A95DE0FF8022AAFACE8753330D3A33" box="[1348,1380,1065,1089]" gridcol="6" gridrow="18" pageId="12" pageNumber="413">2.2</td>
</tr>
<tr id="3578349CFF8022AAFF29872B330D3A18" box="[163,1380,1105,1130]" gridrow="19" pageId="12" pageNumber="413">
<th id="76A95DE0FF8022AAFF29872B34FF3A18" box="[163,662,1105,1130]" gridcol="0" gridrow="19" pageId="12" pageNumber="413">Head depth</th>
<td id="76A95DE0FF8022AAFD3B872B35593A18" box="[689,816,1105,1130]" gridcol="1" gridrow="19" pageId="12" pageNumber="413">55–63</td>
<td id="76A95DE0FF8022AAFCC6872B35EC3A18" box="[844,901,1105,1130]" gridcol="2" gridrow="19" pageId="12" pageNumber="413">59.2</td>
<td id="76A95DE0FF8022AAFC28872B35AB3A18" box="[930,962,1105,1130]" gridcol="3" gridrow="19" pageId="12" pageNumber="413">2.9</td>
<td id="76A95DE0FF8022AAFB8B872B32E93A18" box="[1025,1152,1105,1130]" gridcol="4" gridrow="19" pageId="12" pageNumber="413">57–60</td>
<td id="76A95DE0FF8022AAFB28872B336F3A18" box="[1186,1286,1105,1130]" gridcol="5" gridrow="19" pageId="12" pageNumber="413">58.7</td>
<td id="76A95DE0FF8022AAFACE872B330D3A18" box="[1348,1380,1105,1130]" gridcol="6" gridrow="19" pageId="12" pageNumber="413">2.2</td>
</tr>
<tr id="3578349CFF8022AAFF298700330D3AE0" box="[163,1380,1146,1170]" gridrow="20" pageId="12" pageNumber="413">
<th id="76A95DE0FF8022AAFF29870034FF3AE0" box="[163,662,1146,1170]" gridcol="0" gridrow="20" pageId="12" pageNumber="413">Interorbital width</th>
<td id="76A95DE0FF8022AAFD3B870035593AE0" box="[689,816,1146,1170]" gridcol="1" gridrow="20" pageId="12" pageNumber="413">44–49</td>
<td id="76A95DE0FF8022AAFCC6870035EC3AE0" box="[844,901,1146,1170]" gridcol="2" gridrow="20" pageId="12" pageNumber="413">46.4</td>
<td id="76A95DE0FF8022AAFC28870035AB3AE0" box="[930,962,1146,1170]" gridcol="3" gridrow="20" pageId="12" pageNumber="413">1.9</td>
<td id="76A95DE0FF8022AAFB8B870032E93AE0" box="[1025,1152,1146,1170]" gridcol="4" gridrow="20" pageId="12" pageNumber="413">41–44</td>
<td id="76A95DE0FF8022AAFB288700336F3AE0" box="[1186,1286,1146,1170]" gridcol="5" gridrow="20" pageId="12" pageNumber="413">42.9</td>
<td id="76A95DE0FF8022AAFACE8700330D3AE0" box="[1348,1380,1146,1170]" gridcol="6" gridrow="20" pageId="12" pageNumber="413">2.2</td>
</tr>
<tr id="3578349CFF8022AAFF2987D8330D3AC8" box="[163,1380,1186,1210]" gridrow="21" pageId="12" pageNumber="413">
<th id="76A95DE0FF8022AAFF2987D834FF3AC8" box="[163,662,1186,1210]" gridcol="0" gridrow="21" pageId="12" pageNumber="413">Postorbital length</th>
<td id="76A95DE0FF8022AAFD3B87D835593AC8" box="[689,816,1186,1210]" gridcol="1" gridrow="21" pageId="12" pageNumber="413">42–48</td>
<td id="76A95DE0FF8022AAFCC687D835EC3AC8" box="[844,901,1186,1210]" gridcol="2" gridrow="21" pageId="12" pageNumber="413">45.2</td>
<td id="76A95DE0FF8022AAFC2887D835AB3AC8" box="[930,962,1186,1210]" gridcol="3" gridrow="21" pageId="12" pageNumber="413">2.3</td>
<td id="76A95DE0FF8022AAFB8B87D832E93AC8" box="[1025,1152,1186,1210]" gridcol="4" gridrow="21" pageId="12" pageNumber="413">46–48</td>
<td id="76A95DE0FF8022AAFB2887D8336F3AC8" box="[1186,1286,1186,1210]" gridcol="5" gridrow="21" pageId="12" pageNumber="413">46.8</td>
<td id="76A95DE0FF8022AAFACE87D8330D3AC8" box="[1348,1380,1186,1210]" gridcol="6" gridrow="21" pageId="12" pageNumber="413">1.1</td>
</tr>
<tr id="3578349CFF8022AAFF2987B0330D3A90" box="[163,1380,1226,1250]" gridrow="22" pageId="12" pageNumber="413">
<th id="76A95DE0FF8022AAFF2987B034FF3A90" box="[163,662,1226,1250]" gridcol="0" gridrow="22" pageId="12" pageNumber="413">Suborbital depth</th>
<td id="76A95DE0FF8022AAFD3B87B035593A90" box="[689,816,1226,1250]" gridcol="1" gridrow="22" pageId="12" pageNumber="413">8–12</td>
<td id="76A95DE0FF8022AAFCC687B035EC3A90" box="[844,901,1226,1250]" gridcol="2" gridrow="22" pageId="12" pageNumber="413">9.9</td>
<td id="76A95DE0FF8022AAFC2887B035AB3A90" box="[930,962,1226,1250]" gridcol="3" gridrow="22" pageId="12" pageNumber="413">1.5</td>
<td id="76A95DE0FF8022AAFB8B87B032E93A90" box="[1025,1152,1226,1250]" gridcol="4" gridrow="22" pageId="12" pageNumber="413">6–8</td>
<td id="76A95DE0FF8022AAFB2887B0336F3A90" box="[1186,1286,1226,1250]" gridcol="5" gridrow="22" pageId="12" pageNumber="413">7.1</td>
<td id="76A95DE0FF8022AAFACE87B0330D3A90" box="[1348,1380,1226,1250]" gridcol="6" gridrow="22" pageId="12" pageNumber="413">1.1</td>
</tr>
<tr id="3578349CFF8022AAFF298788330D3B79" box="[163,1380,1266,1291]" gridrow="23" pageId="12" pageNumber="413">
<th id="76A95DE0FF8022AAFF29878834FF3B79" box="[163,662,1266,1291]" gridcol="0" gridrow="23" pageId="12" pageNumber="413">Eye diameter</th>
<td id="76A95DE0FF8022AAFD3B878835593B79" box="[689,816,1266,1291]" gridcol="1" gridrow="23" pageId="12" pageNumber="413">32–37</td>
<td id="76A95DE0FF8022AAFCC6878835EC3B79" box="[844,901,1266,1291]" gridcol="2" gridrow="23" pageId="12" pageNumber="413">34.2</td>
<td id="76A95DE0FF8022AAFC28878835AB3B79" box="[930,962,1266,1291]" gridcol="3" gridrow="23" pageId="12" pageNumber="413">1.5</td>
<td id="76A95DE0FF8022AAFB8B878832E93B79" box="[1025,1152,1266,1291]" gridcol="4" gridrow="23" pageId="12" pageNumber="413">33–35</td>
<td id="76A95DE0FF8022AAFB288788336F3B79" box="[1186,1286,1266,1291]" gridcol="5" gridrow="23" pageId="12" pageNumber="413">34.1</td>
<td id="76A95DE0FF8022AAFACE8788330D3B79" box="[1348,1380,1266,1291]" gridcol="6" gridrow="23" pageId="12" pageNumber="413">1.1</td>
</tr>
<tr id="3578349CFF8022AAFF298661330D3B41" box="[163,1380,1307,1331]" gridrow="24" pageId="12" pageNumber="413">
<th id="76A95DE0FF8022AAFF29866134FF3B41" box="[163,662,1307,1331]" gridcol="0" gridrow="24" pageId="12" pageNumber="413">Snout to eye end length</th>
<td id="76A95DE0FF8022AAFD3B866135593B41" box="[689,816,1307,1331]" gridcol="1" gridrow="24" pageId="12" pageNumber="413">52–58</td>
<td id="76A95DE0FF8022AAFCC6866135EC3B41" box="[844,901,1307,1331]" gridcol="2" gridrow="24" pageId="12" pageNumber="413">54.8</td>
<td id="76A95DE0FF8022AAFC28866135AB3B41" box="[930,962,1307,1331]" gridcol="3" gridrow="24" pageId="12" pageNumber="413">2.3</td>
<td id="76A95DE0FF8022AAFB8B866132E93B41" box="[1025,1152,1307,1331]" gridcol="4" gridrow="24" pageId="12" pageNumber="413">51–54</td>
<td id="76A95DE0FF8022AAFB288661336F3B41" box="[1186,1286,1307,1331]" gridcol="5" gridrow="24" pageId="12" pageNumber="413">52.4</td>
<td id="76A95DE0FF8022AAFACE8661330D3B41" box="[1348,1380,1307,1331]" gridcol="6" gridrow="24" pageId="12" pageNumber="413">2.2</td>
</tr>
<tr id="3578349CFF8022AAFF298639330D3B29" box="[163,1380,1347,1371]" gridrow="25" pageId="12" pageNumber="413">
<th id="76A95DE0FF8022AAFF29863934FF3B29" box="[163,662,1347,1371]" gridcol="0" gridrow="25" pageId="12" pageNumber="413">Snout length</th>
<td id="76A95DE0FF8022AAFD3B863935593B29" box="[689,816,1347,1371]" gridcol="1" gridrow="25" pageId="12" pageNumber="413">20–25</td>
<td id="76A95DE0FF8022AAFCC6863935EC3B29" box="[844,901,1347,1371]" gridcol="2" gridrow="25" pageId="12" pageNumber="413">21.7</td>
<td id="76A95DE0FF8022AAFC28863935AB3B29" box="[930,962,1347,1371]" gridcol="3" gridrow="25" pageId="12" pageNumber="413">2.1</td>
<td id="76A95DE0FF8022AAFB8B863932E93B29" box="[1025,1152,1347,1371]" gridcol="4" gridrow="25" pageId="12" pageNumber="413">19–21</td>
<td id="76A95DE0FF8022AAFB288639336F3B29" box="[1186,1286,1347,1371]" gridcol="5" gridrow="25" pageId="12" pageNumber="413">19.8</td>
<td id="76A95DE0FF8022AAFACE8639330D3B29" box="[1348,1380,1347,1371]" gridcol="6" gridrow="25" pageId="12" pageNumber="413">1.1</td>
</tr>
<tr id="3578349CFF8022AAFF298611330D3BF1" box="[163,1380,1387,1411]" gridrow="26" pageId="12" pageNumber="413">
<th id="76A95DE0FF8022AAFF298611330D3BF1" box="[163,1380,1387,1411]" colspan="7" colspanRight="6" gridcol="0" gridrow="26" pageId="12" pageNumber="413">
<emphasis id="B93CEACCFF80DD59FF29861137A33BF1" bold="true" box="[163,458,1387,1411]" pageId="12" pageNumber="413">Other morphometric ratios</emphasis>
</th>
</tr>
<tr id="3578349CFF8022AAFF2986E9330D3BDE" box="[163,1380,1427,1452]" gridrow="27" pageId="12" pageNumber="413">
<th id="76A95DE0FF8022AAFF2986E934FF3BDE" box="[163,662,1427,1452]" gridcol="0" gridrow="27" pageId="12" pageNumber="413">Predorsal length in % of preanal length</th>
<td id="76A95DE0FF8022AAFD3B86E935593BDE" box="[689,816,1427,1452]" gridcol="1" gridrow="27" pageId="12" pageNumber="413">117–122</td>
<td id="76A95DE0FF8022AAFCC686E935EC3BDE" box="[844,901,1427,1452]" gridcol="2" gridrow="27" pageId="12" pageNumber="413">119.8</td>
<td id="76A95DE0FF8022AAFC2886E935AB3BDE" box="[930,962,1427,1452]" gridcol="3" gridrow="27" pageId="12" pageNumber="413">1.7</td>
<td id="76A95DE0FF8022AAFB8B86E932E93BDE" box="[1025,1152,1427,1452]" gridcol="4" gridrow="27" pageId="12" pageNumber="413">114–117</td>
<td id="76A95DE0FF8022AAFB2886E9336F3BDE" box="[1186,1286,1427,1452]" gridcol="5" gridrow="27" pageId="12" pageNumber="413">115.2</td>
<td id="76A95DE0FF8022AAFACE86E9330D3BDE" box="[1348,1380,1427,1452]" gridcol="6" gridrow="27" pageId="12" pageNumber="413">1.9</td>
</tr>
<tr id="3578349CFF8022AAFF2986C6330D3BA6" box="[163,1380,1468,1492]" gridrow="28" pageId="12" pageNumber="413">
<th id="76A95DE0FF8022AAFF2986C634FF3BA6" box="[163,662,1468,1492]" gridcol="0" gridrow="28" pageId="12" pageNumber="413">Caudal peduncle length in % of its depth</th>
<td id="76A95DE0FF8022AAFD3B86C635593BA6" box="[689,816,1468,1492]" gridcol="1" gridrow="28" pageId="12" pageNumber="413">148–156</td>
<td id="76A95DE0FF8022AAFCC686C635EC3BA6" box="[844,901,1468,1492]" gridcol="2" gridrow="28" pageId="12" pageNumber="413">152.1</td>
<td id="76A95DE0FF8022AAFC2886C635AB3BA6" box="[930,962,1468,1492]" gridcol="3" gridrow="28" pageId="12" pageNumber="413">2.4</td>
<td id="76A95DE0FF8022AAFB8B86C632E93BA6" box="[1025,1152,1468,1492]" gridcol="4" gridrow="28" pageId="12" pageNumber="413">161–167</td>
<td id="76A95DE0FF8022AAFB2886C6336F3BA6" box="[1186,1286,1468,1492]" gridcol="5" gridrow="28" pageId="12" pageNumber="413">164</td>
<td id="76A95DE0FF8022AAFACE86C6330D3BA6" box="[1348,1380,1468,1492]" gridcol="6" gridrow="28" pageId="12" pageNumber="413">3.8</td>
</tr>
<tr id="3578349CFF8022AAFF29869E330D3B8E" box="[163,1380,1508,1532]" gridrow="29" pageId="12" pageNumber="413">
<th id="76A95DE0FF8022AAFF29869E34FF3B8E" box="[163,662,1508,1532]" gridcol="0" gridrow="29" pageId="12" pageNumber="413">Head width in % of its depth</th>
<td id="76A95DE0FF8022AAFD3B869E35593B8E" box="[689,816,1508,1532]" gridcol="1" gridrow="29" pageId="12" pageNumber="413">87–97</td>
<td id="76A95DE0FF8022AAFCC6869E35EC3B8E" box="[844,901,1508,1532]" gridcol="2" gridrow="29" pageId="12" pageNumber="413">91.3</td>
<td id="76A95DE0FF8022AAFC28869E35AB3B8E" box="[930,962,1508,1532]" gridcol="3" gridrow="29" pageId="12" pageNumber="413">3.2</td>
<td id="76A95DE0FF8022AAFB8B869E32E93B8E" box="[1025,1152,1508,1532]" gridcol="4" gridrow="29" pageId="12" pageNumber="413">89–90</td>
<td id="76A95DE0FF8022AAFB28869E336F3B8E" box="[1186,1286,1508,1532]" gridcol="5" gridrow="29" pageId="12" pageNumber="413">89.2</td>
<td id="76A95DE0FF8022AAFACE869E330D3B8E" box="[1348,1380,1508,1532]" gridcol="6" gridrow="29" pageId="12" pageNumber="413">0.4</td>
</tr>
<tr id="3578349CFF8022AAFF298576330D3857" box="[163,1380,1548,1573]" gridrow="30" pageId="12" pageNumber="413" rowspan-3="1" rowspan-6="1">
<th id="76A95DE0FF8022AAFF29857634FF3857" box="[163,662,1548,1573]" gridcol="0" gridrow="30" pageId="12" pageNumber="413">
<emphasis id="B93CEACCFF80DD59FF298576376E3856" bold="true" box="[163,263,1548,1572]" pageId="12" pageNumber="413">Meristics</emphasis>
</th>
<td id="76A95DE0FF8022AAFD3B857635593857" box="[689,816,1548,1573]" gridcol="1" gridrow="30" pageId="12" pageNumber="413">range</td>
<td id="76A95DE0FF8022AAFCC6857635EC3857" box="[844,901,1548,1573]" gridcol="2" gridrow="30" pageId="12" pageNumber="413">mode</td>
<td id="76A95DE0FF8022AAFB8B857632E93857" box="[1025,1152,1548,1573]" gridcol="4" gridrow="30" pageId="12" pageNumber="413">range</td>
<td id="76A95DE0FF8022AAFB288576336F3857" box="[1186,1286,1548,1573]" gridcol="5" gridrow="30" pageId="12" pageNumber="413">mode</td>
</tr>
<tr id="3578349CFF8022AAFF29854E330D383F" box="[163,1380,1588,1613]" gridrow="31" pageId="12" pageNumber="413" rowspan-3="1" rowspan-6="1">
<th id="76A95DE0FF8022AAFF29854E34FF383F" box="[163,662,1588,1613]" gridcol="0" gridrow="31" pageId="12" pageNumber="413">Dorsal-fin rays</th>
<td id="76A95DE0FF8022AAFD3B854E3559383F" box="[689,816,1588,1613]" gridcol="1" gridrow="31" pageId="12" pageNumber="413">11–13</td>
<td id="76A95DE0FF8022AAFCC6854E35EC383F" box="[844,901,1588,1613]" gridcol="2" gridrow="31" pageId="12" pageNumber="413">11</td>
<td id="76A95DE0FF8022AAFB8B854E32E9383F" box="[1025,1152,1588,1613]" gridcol="4" gridrow="31" pageId="12" pageNumber="413">11–12</td>
<td id="76A95DE0FF8022AAFB28854E336F383F" box="[1186,1286,1588,1613]" gridcol="5" gridrow="31" pageId="12" pageNumber="413">-</td>
</tr>
<tr id="3578349CFF8022AAFF298527330D3807" box="[163,1380,1629,1653]" gridrow="32" pageId="12" pageNumber="413" rowspan-3="1" rowspan-6="1">
<th id="76A95DE0FF8022AAFF29852734FF3807" box="[163,662,1629,1653]" gridcol="0" gridrow="32" pageId="12" pageNumber="413">Anal-fin rays</th>
<td id="76A95DE0FF8022AAFD3B852735593807" box="[689,816,1629,1653]" gridcol="1" gridrow="32" pageId="12" pageNumber="413">17–19</td>
<td id="76A95DE0FF8022AAFCC6852735EC3807" box="[844,901,1629,1653]" gridcol="2" gridrow="32" pageId="12" pageNumber="413">18</td>
<td id="76A95DE0FF8022AAFB8B852732E93807" box="[1025,1152,1629,1653]" gridcol="4" gridrow="32" pageId="12" pageNumber="413">17–18</td>
<td id="76A95DE0FF8022AAFB288527336F3807" box="[1186,1286,1629,1653]" gridcol="5" gridrow="32" pageId="12" pageNumber="413">-</td>
</tr>
<tr id="3578349CFF8022AAFF2985FF330D38EF" box="[163,1380,1669,1693]" gridrow="33" pageId="12" pageNumber="413" rowspan-3="1" rowspan-6="1">
<th id="76A95DE0FF8022AAFF2985FF34FF38EF" box="[163,662,1669,1693]" gridcol="0" gridrow="33" pageId="12" pageNumber="413">Scales mid-longitudinal series</th>
<td id="76A95DE0FF8022AAFD3B85FF355938EF" box="[689,816,1669,1693]" gridcol="1" gridrow="33" pageId="12" pageNumber="413">26–28</td>
<td id="76A95DE0FF8022AAFCC685FF35EC38EF" box="[844,901,1669,1693]" gridcol="2" gridrow="33" pageId="12" pageNumber="413">27</td>
<td id="76A95DE0FF8022AAFB8B85FF32E938EF" box="[1025,1152,1669,1693]" gridcol="4" gridrow="33" pageId="12" pageNumber="413">27–28</td>
<td id="76A95DE0FF8022AAFB2885FF336F38EF" box="[1186,1286,1669,1693]" gridcol="5" gridrow="33" pageId="12" pageNumber="413">-</td>
</tr>
<tr id="3578349CFF8022AAFF2985D7330D38B7" box="[163,1380,1709,1733]" gridrow="34" pageId="12" pageNumber="413" rowspan-3="1" rowspan-6="1">
<th id="76A95DE0FF8022AAFF2985D734FF38B7" box="[163,662,1709,1733]" gridcol="0" gridrow="34" pageId="12" pageNumber="413">Scales transverse</th>
<td id="76A95DE0FF8022AAFD3B85D7355938B7" box="[689,816,1709,1733]" gridcol="1" gridrow="34" pageId="12" pageNumber="413">10–10</td>
<td id="76A95DE0FF8022AAFCC685D735EC38B7" box="[844,901,1709,1733]" gridcol="2" gridrow="34" pageId="12" pageNumber="413">10</td>
<td id="76A95DE0FF8022AAFB8B85D732E938B7" box="[1025,1152,1709,1733]" gridcol="4" gridrow="34" pageId="12" pageNumber="413">10–10</td>
<td id="76A95DE0FF8022AAFB2885D7336F38B7" box="[1186,1286,1709,1733]" gridcol="5" gridrow="34" pageId="12" pageNumber="413">10</td>
</tr>
<tr id="3578349CFF8022AAFF2985AF330D389C" box="[163,1380,1749,1774]" gridrow="35" pageId="12" pageNumber="413" rowspan-3="1" rowspan-6="1">
<th id="76A95DE0FF8022AAFF2985AF34FF389C" box="[163,662,1749,1774]" gridcol="0" gridrow="35" pageId="12" pageNumber="413">Scales circumpeduncular</th>
<td id="76A95DE0FF8022AAFD3B85AF3559389C" box="[689,816,1749,1774]" gridcol="1" gridrow="35" pageId="12" pageNumber="413">12–12</td>
<td id="76A95DE0FF8022AAFCC685AF35EC389C" box="[844,901,1749,1774]" gridcol="2" gridrow="35" pageId="12" pageNumber="413">12</td>
<td id="76A95DE0FF8022AAFB8B85AF32E9389C" box="[1025,1152,1749,1774]" gridcol="4" gridrow="35" pageId="12" pageNumber="413">12–12</td>
<td id="76A95DE0FF8022AAFB2885AF336F389C" box="[1186,1286,1749,1774]" gridcol="5" gridrow="35" pageId="12" pageNumber="413">12</td>
</tr>
</table>
</paragraph>
<paragraph id="8BF736DEFF80DD58FF4D846733F53E8A" blockId="12.[151,1437,1821,2027]" lastBlockId="13.[151,1437,151,2036]" lastPageId="13" lastPageNumber="414" pageId="12" pageNumber="413">Dorsal-fin origin markedly posterior to anal-fin origin (predorsal length 117–122 as % of preanal length), both fins originating posterior to mid-length of body (predorsal length 68.0–73.0 % SL; preanal length 55.7–60.5 % SL). Dorsal fin relatively small with low number of rays, 11–13; extremity rounded. Anal fin trapezoidal in shape, with 17–19 rays; lower margin quite strongly concave in profile, resulting in mildly lobate rear part of fin. Dorsal and anal fins with short fin-ray extensions. Pectoral fin subtriangular, insertion slightly posterior to margin of opercular opening, base slightly oblique, upper fin rays positioned slightly anterior to lower fin rays, with tip reaching or slightly overlapping base of pelvic fin. Pelvic fin subabdominal, origin at about mid-length of body, short, base medially separated, with tip reaching urogenital papilla. Caudal fin rounded, with 13–16 branched rays, plus 3–4 dorsal and ventral procurrent rays; central 2–3 rays of fin relatively slightly elongated resulting in mild central lobe.</paragraph>
<paragraph id="8BF736DEFF81DD58FF4D8278359B3F11" blockId="13.[151,1437,151,2036]" pageId="13" pageNumber="414">Scales cycloid, body and head entirely scaled, except for ventral surface of head. No scales on base of dorsal and anal fins. Scales in mid-longitudinal series 26–28 plus 2–3 small scales on caudal-fin base. Transverse rows of scales in front of dorsal-fin origin 10; scale rows around caudal peduncle 12.</paragraph>
<paragraph id="8BF736DEFF81DD58FF4D821737CB3CD7" blockId="13.[151,1437,151,2036]" pageId="13" pageNumber="414">
Cephalic squamation pattern G-type. Frontal neuromast in single shallow groove behind upper lip. Cephalic sensory system at supraorbital level in an undivided, slightly curved zigzag pattern, almost flat, shallow groove with three exposed neuromasts and pronounced lobes (
<figureCitation id="13732A5BFF81DD58FD7482CE35393FBC" box="[766,848,436,462]" captionStart="FIGURE 3" captionStartId="10.[151,250,1307,1331]" captionTargetBox="[156,1428,706,1276]" captionTargetId="figure-280@10.[151,1436,702,1281]" captionTargetPageId="10" captionText="FIGURE 3. Cephalic structure: a, Nothobranchius janpapi; b, N. luekei; c, N. geminus; d, N. lourensi and N. fuscotaeniatus. Key to label abbreviations: an, anterior nostril; pn, posterior nostril; f, frontal neuromast; so, supraorbital canal; st, supratemporal canal." pageId="13" pageNumber="414">Fig. 3b</figureCitation>
), whereas at supratemporal level with three exposed neuromasts in slightly curved arrangement on flat surface, with pronounced lobes. Preorbital canal in deep groove with two exposed neuromasts; infraorbital level with series of about 20 small buttons at ventral and posterior margin of orbit; postorbital canal in deep groove with one exposed neuromast. Preopercular system with six exposed neuromasts in deep groove on preopercle portion, whereas in shallow groove with six small neuromasts on ventral portion. Mandibular canal in shallow groove with about 20 small buttons. One neuromast on each scale along trunk mid-longitudinal series.
</paragraph>
<paragraph id="8BF736DEFF81DD58FF4D81D535DC3D0E" blockId="13.[151,1437,151,2036]" pageId="13" pageNumber="414">
Female: Smaller than male, maximum recorded size
<quantity id="4CB09B3BFF81DD58FCA081D535F83CBB" box="[810,913,687,713]" metricMagnitude="-2" metricUnit="m" metricValue="2.46" pageId="13" pageNumber="414" unit="mm" value="24.6">24.6 mm</quantity>
SL. Branchiostegal membrane not projecting posteriorly from opercle. Dorsal fin rounded with low number of fin rays, 11–12. Caudal fin basically rounded but with central 2–3 rays elongated resulting in pointed posterior margin. Anal fin subtriangular, tip rounded, central rays longer and more rigid; posterior lower margin may be very slightly concave, mimicking in very mild way, that of male. Anal-fin base length smaller than in male (19.7–20.3 % SL vs. 21.9–26.3). Caudal peduncle shallower than in males (caudal peduncle length 161–167 % of its depth vs. 148–156).
</paragraph>
<paragraph id="8BF736DEFF81DD58FF4D80FF32E83834" blockId="13.[151,1437,151,2036]" pageId="13" pageNumber="414">
<emphasis id="B93CEACCFF81DD58FF4D80FF37363DED" bold="true" box="[199,351,901,927]" pageId="13" pageNumber="414">Colouration.</emphasis>
Live male (
<figureCitation id="13732A5BFF81DD58FE6580FF34133DED" box="[495,634,901,927]" captionStart="FIGURE 4" captionStartId="11.[151,250,1384,1408]" captionTargetBox="[151,1435,558,1358]" captionTargetId="figure-251@11.[151,1435,558,1358]" captionTargetPageId="11" captionText="FIGURE 4. Wild-caught specimens of Nothobranchius luekei and example of typical habitat. A. N. luekei Mbezi River TAN 00-23, male, Mbezi River drainage, in immediate vicinity of the type locality; B. N. luekei Luhule River TAN 02-8, male, Luhule/Luhute River drainage; C. N. luekei Mbezi River TZN 09-9, female, Mbezi River drainage (photograph by B. Nagy); D. Location Luhule River TAN 02-8, a typical heavily vegetated shallow habitat (photograph June 2, 2002), hosting N. luekei, N. lucius, N. rubripinnis and N. ruudwildekampi. Photographs by B.R. Watters except where otherwise noted." pageId="13" pageNumber="414">Figs. 4A–B</figureCitation>
): Scales on the trunk and head are light iridescent blue to blue-green with red margins, forming a regular reticulation pattern. Scales on the abdomen are faint blue to cream with less pronounced margins. Scales on the dorsum anterior to the dorsal fin have yellow-green centres forming a narrow stripe. Snout, and frontal and dorsal portions of head, are yellow-green (Mbezi population) or grey (Luhule/Luhute population); the throat is yellow-green. Operculum has blue to blue-green, and lesser red, markings with intervening yellow areas. Exposed part of the branchiostegal membrane is yellow with white edging. Iris is blue to blue-green. Dorsal fin background light blue-green in the basal and proximal zones grading to yellow through the medial zone and into the distal zone; background overlaid by irregular red spots and patches that become coarser in the medial and distal zones (Luhule/Luhute) that may also take the form of streaks parallel to fin rays (Mbezi), merging in the distal zone as an irregular marginal band. Anal fin background comprises light blue basal and proximal zones grading to blue-green through the medial zone to yellow in the subdistal zone. In the Luhule/Luhute population the anal fin background is overlaid by narrow irregular red bands over the proximal, sub-medial, medial and distal zones, the latter forming a distinct marginal band; in the Mbezi population the background is overlaid by prominent red streaks arranged parallel to fin rays that end at the yellow subdistal zone and there is a red marginal band in the distal zone. Basal, proximal and medial zones of the caudal fin are grey (Luhule/Luhute) or light red (Mbezi), with red spots and streaks, the latter aligned parallel to the fin rays; in the Luhule/Luhute population the distal zone comprises a broad red band followed by a black marginal band; in the Mbezi population the distal zone grades from a relatively dark red outwards to a lighter red, with no black margin. Pelvic fins mimic, in part, the pattern of anal fin with light blue overlaid by red markings, followed by an irregular red band or red spots, a yellow band and red edging. Pectoral fins are dominantly hyaline or pale yellow with prominent blue margins.
</paragraph>
<paragraph id="8BF736DEFF81DD58FF4D852A33E938C3" blockId="13.[151,1437,151,2036]" pageId="13" pageNumber="414">
Live female (
<figureCitation id="13732A5BFF81DD58FED5852A37D33818" box="[351,442,1616,1642]" captionStart="FIGURE 4" captionStartId="11.[151,250,1384,1408]" captionTargetBox="[151,1435,558,1358]" captionTargetId="figure-251@11.[151,1435,558,1358]" captionTargetPageId="11" captionText="FIGURE 4. Wild-caught specimens of Nothobranchius luekei and example of typical habitat. A. N. luekei Mbezi River TAN 00-23, male, Mbezi River drainage, in immediate vicinity of the type locality; B. N. luekei Luhule River TAN 02-8, male, Luhule/Luhute River drainage; C. N. luekei Mbezi River TZN 09-9, female, Mbezi River drainage (photograph by B. Nagy); D. Location Luhule River TAN 02-8, a typical heavily vegetated shallow habitat (photograph June 2, 2002), hosting N. luekei, N. lucius, N. rubripinnis and N. ruudwildekampi. Photographs by B.R. Watters except where otherwise noted." pageId="13" pageNumber="414">Fig. 4C</figureCitation>
): Trunk and head are light grey, darker on the dorsum and lighter ventrally. Scales have narrow darker grey margins on the flanks, becoming broader on the back; scale centres on the trunk above the abdomen are iridescent pale blue; the operculum is iridescent blue-green. All fins are hyaline. The iris is golden.
</paragraph>
</subSubSection>
<subSubSection id="C3526555FF81DD5BFF4D85C1356E3FDF" lastPageId="14" lastPageNumber="415" pageId="13" pageNumber="414" type="distribution">
<paragraph id="8BF736DEFF81DD58FF4D85C1326D3916" blockId="13.[151,1437,151,2036]" pageId="13" pageNumber="414">
<emphasis id="B93CEACCFF81DD58FF4D85C1370838A7" bold="true" box="[199,353,1723,1749]" pageId="13" pageNumber="414">Distribution.</emphasis>
<taxonomicName id="4C484D5DFF81DD58FEE185C1340738A7" box="[363,622,1723,1749]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="13" pageNumber="414" phylum="Chordata" rank="species" species="luekei">
<emphasis id="B93CEACCFF81DD58FEE185C1340738A7" box="[363,622,1723,1749]" italics="true" pageId="13" pageNumber="414">Nothobranchius luekei</emphasis>
</taxonomicName>
is endemic to seasonal freshwater habitats in coastal
<collectingCountry id="F35F764EFF81DD58FB6D85C1332538A7" box="[1255,1356,1723,1749]" name="Tanzania" pageId="13" pageNumber="414">Tanzania</collectingCountry>
and is relatively sparsely distributed over a restricted range. Confirmed sites occur in association with the Mbezi and Luhule/Luhute river systems in the near coastal region south of
<collectingRegion id="498CF83CFF81DD58FCEF84793262396E" box="[869,1035,1794,1821]" country="Tanzania" name="Dar es Salaam" pageId="13" pageNumber="414">Dar es Salaam</collectingRegion>
. It appears to be confined to the so-called Mbezi Triangle (
<emphasis id="B93CEACCFF81DD58FE14845D37B53932" box="[414,476,1831,1856]" italics="true" pageId="13" pageNumber="414">sensu</emphasis>
<bibRefCitation id="EFD94B2FFF81DD58FE69845D34D03932" author="Watters, B. R. &amp; Cooper, B. J. &amp; Wildekamp, R. H." box="[483,697,1830,1857]" pageId="13" pageNumber="414" pagination="5 - 6" refId="ref35078" refString="Watters, B. R., Cooper, B. J. &amp; Wildekamp, R. H. (2007) Description of Nothobranchius cardinalis spec. nov. (Cyprinodontiformes: Aplocheilidae), an annual fish from the Mbwemkuru River basin, Tanzania. Journal of the American Killifish Association, 40 (5 - 6), 129 - 145. [issue printed in 2008 = year of species description]" type="journal article" year="2007">
Watters
<emphasis id="B93CEACCFF81DD58FDB5845D34193932" box="[575,624,1830,1856]" italics="true" pageId="13" pageNumber="414">et al</emphasis>
. 2007
</bibRefCitation>
;
<bibRefCitation id="EFD94B2FFF81DD58FD4F845D35343932" author="Watters, B. R." box="[709,861,1830,1857]" pageId="13" pageNumber="414" pagination="37 - 76" refId="ref34941" refString="Watters, B. R. (2009) The ecology and distribution of Nothobranchius fishes. Journal of the American Killifish Association, 42 (2), 37 - 76." type="journal article" year="2009">Watters 2009</bibRefCitation>
), a region that is separated to varying degrees from the Ruvu and Rufiji systems by the Pugu and Mtoti hills, respectively (
<figureCitation id="13732A5BFF81DD58FC388430359A3916" box="[946,1011,1866,1892]" captionStart="FIGURE 1" captionStartId="2.[151,250,1762,1786]" captionTargetBox="[151,1436,374,1736]" captionTargetId="figure-118@2.[151,1436,374,1736]" captionTargetPageId="2" captionText="FIGURE 1. Distribution of species belonging to the subgenus Aphyobranchius, all of which occur in the east-central and southern coastal lowlands of Tanzania: N. janpapi (yellow-filled circle), N. luekei (red-filled triangle), N. geminus (orange-filled diamond), N. lourensi (green-filled square), N. fuscotaeniatus (blue-filled inverted triangle), N. cf. janpapi (dark magenta-filled circle). The inset map shows the area of distribution in a somewhat broader geographic context, to include the distribution of N. willerti (red-filled star) in the lower Tana River area of northeastern coastal Kenya, a species that herein is considered to belong to the subgenus Adiniops, rather than Aphyobranchius to which it was ascribed in the original description by Wildekamp (1992). Note that individual symbols on the map may represent more than one locality or population where the sites are close to one another and could not reasonably be resolved at the scale of the map." pageId="13" pageNumber="414">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8BF736DEFF81DD5BFF4D8417356E3FDF" blockId="13.[151,1437,151,2036]" lastBlockId="14.[151,1436,151,429]" lastPageId="14" lastPageNumber="415" pageId="13" pageNumber="414">
<emphasis id="B93CEACCFF81DD58FF4D841737D639FA" bold="true" box="[199,447,1901,1928]" pageId="13" pageNumber="414">Conservation status.</emphasis>
<taxonomicName id="4C484D5DFF81DD58FE40841734A539F5" box="[458,716,1901,1927]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="13" pageNumber="414" phylum="Chordata" rank="species" species="luekei">
<emphasis id="B93CEACCFF81DD58FE40841734A539F5" box="[458,716,1901,1927]" italics="true" pageId="13" pageNumber="414">Nothobranchius luekei</emphasis>
</taxonomicName>
has been assessed as Endangered B1ab(iii)+2ab(iii) (
<bibRefCitation id="EFD94B2FFF81DD58FAB78414375439D9" author="Nagy, B. &amp; Watters, B." pageId="13" pageNumber="414" refId="ref33642" refString="Nagy, B. &amp; Watters, B. (2019 b) Nothobranchius luekei. The IUCN Red List of Threatened Species, 2019, e. T 60336 A 47183695. https: // doi. org / 10.2305 / IUCN. UK. 2019 - 3. RLTS. T 60336 A 47183695. en" type="journal volume" year="2019">Nagy &amp; Watters 2019b</bibRefCitation>
). The known extent of occurrence (EOO) is
<quantity id="4CB09B3BFF81DD58FCAF84EB351239DC" box="[805,891,1937,1966]" metricMagnitude="5" metricUnit="m" metricValue="2.38" pageId="13" pageNumber="414" unit="km" value="238.0">238 km</quantity>
², with a maximum estimate of around
<quantity id="4CB09B3BFF81DD58FAAE84EB33E639DC" box="[1316,1423,1937,1966]" metricMagnitude="6" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="414" unit="km" value="1000.0">1,000 km</quantity>
², whereas the area of occupancy (AOO) is
<quantity id="4CB09B3BFF81DD58FDE884CF34C339A0" box="[610,682,1973,2002]" metricMagnitude="4" metricUnit="m" metricValue="2.4" pageId="13" pageNumber="414" unit="km" value="24.0">24 km</quantity>
², with a maximum estimate of around
<quantity id="4CB09B3BFF81DD58FBDD84CF32C739A0" box="[1111,1198,1973,2002]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="414" unit="km" value="100.0">100 km</quantity>
². Within its restricted area of distribution, the occurrence of the species is fragmented and currently it is known from only five threat-based locations. As observed directly by the authors during the course of repeated visits to localities over a number of years, there is increasing pressure on land resources due to the expansion of agriculture in both principal areas where this species occurs. Such expansion of agricultural activity is resulting in habitat degradation and modification, rendering them unsuitable to support the semi-annual life cycle of the species and by altering conditions of the habitat niche preferred by this species. This category is assigned based on using the upper estimates of the EOO, AOO, and number of locations as best estimates. As presently known, this species is restricted to the Mbezi Triangle (
<emphasis id="B93CEACCFF82DD5BFF2B821536B63FFA" box="[161,223,367,392]" italics="true" pageId="14" pageNumber="415">sensu</emphasis>
<bibRefCitation id="EFD94B2FFF82DD5BFF69821537DA3FFB" author="Watters, B. R. &amp; Cooper, B. J. &amp; Wildekamp, R. H." box="[227,435,367,393]" pageId="14" pageNumber="415" pagination="5 - 6" refId="ref35078" refString="Watters, B. R., Cooper, B. J. &amp; Wildekamp, R. H. (2007) Description of Nothobranchius cardinalis spec. nov. (Cyprinodontiformes: Aplocheilidae), an annual fish from the Mbwemkuru River basin, Tanzania. Journal of the American Killifish Association, 40 (5 - 6), 129 - 145. [issue printed in 2008 = year of species description]" type="journal article" year="2007">
Watters
<emphasis id="B93CEACCFF82DD5BFEB7821537053FFB" box="[317,364,367,393]" italics="true" pageId="14" pageNumber="415">et al</emphasis>
. 2007
</bibRefCitation>
;
<bibRefCitation id="EFD94B2FFF82DD5BFE37821534383FFB" author="Watters, B. R." box="[445,593,367,393]" pageId="14" pageNumber="415" pagination="37 - 76" refId="ref34941" refString="Watters, B. R. (2009) The ecology and distribution of Nothobranchius fishes. Journal of the American Killifish Association, 42 (2), 37 - 76." type="journal article" year="2009">Watters 2009</bibRefCitation>
) in coastal
<collectingCountry id="F35F764EFF82DD5BFD408215355B3FFB" box="[714,818,367,393]" name="Tanzania" pageId="14" pageNumber="415">Tanzania</collectingCountry>
, and its distribution will remain restricted by geographic confinement and configuration of the drainage systems.
</paragraph>
</subSubSection>
</treatment>
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<mods:number id="5E2A3D1CC870C97800B439EE82D63C68">5570</mods:number>
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<mods:number id="9CE0D910AD9E9016BA08FEBE5C5769E8">3</mods:number>
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<mods:extent id="9D0CFE60EDA59FF1560968926797B751" unit="page">
<mods:start id="C06810BADD6750C8B96E70B2185039A4">401</mods:start>
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<mods:location id="1A1AA84B77CF5CAE61A31977A9454757">
<mods:url id="4D94AA0BA00D7EBFBE55B0432642272E">https://doi.org/10.11646/zootaxa.5570.3.1</mods:url>
</mods:location>
<mods:classification id="9677AA73F73B15A5CECFBE09B78257EA">journal article</mods:classification>
<mods:identifier id="5DAA1FDF97F66DC1D86970040CA6EF4E" type="DOI">10.11646/zootaxa.5570.3.1</mods:identifier>
<mods:identifier id="B7671D8930E7D958ADF57B2297F151D2" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="6B8476E24E5EAB23EC468550033A8C4B" type="ZooBank">7C1CE642-C779-40B8-A689-1C364888F500</mods:identifier>
</mods:mods>
<treatment id="03E187C8FF8ADD5FFF1D818034D33CDB" LSID="urn:lsid:plazi:treatment:03E187C8FF8ADD5FFF1D818034D33CDB" httpUri="http://treatment.plazi.org/id/03E187C8FF8ADD5FFF1D818034D33CDB" lastPageId="10" lastPageNumber="411" pageId="6" pageNumber="407">
<subSubSection id="C3526555FF8ADD53FF1D818034CB3D66" box="[151,674,762,789]" pageId="6" pageNumber="407" type="nomenclature">
<paragraph id="8BF736DEFF8ADD53FF1D818034CB3D66" blockId="6.[151,674,762,825]" box="[151,674,762,789]" pageId="6" pageNumber="407">
<heading id="D0BF81B2FF8ADD53FF1D818034CB3D66" bold="true" box="[151,674,762,789]" fontSize="11" level="1" pageId="6" pageNumber="407" reason="1">
<taxonomicName id="4C484D5DFF8ADD53FF1D818034CB3D66" ID-CoL="47SM5" authority="Wildekamp, 1977" authorityName="Wildekamp" authorityYear="1977" box="[151,674,762,789]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF8ADD53FF1D818034CB3D66" bold="true" box="[151,674,762,789]" pageId="6" pageNumber="407">
<emphasis id="B93CEACCFF8ADD53FF1D818037D63D67" bold="true" box="[151,447,762,789]" italics="true" pageId="6" pageNumber="407">Nothobranchius janpapi</emphasis>
<bibRefCitation id="EFD94B2FFF8ADD53FE4C818034CB3D66" author="Wildekamp, R. H." box="[454,674,762,788]" pageId="6" pageNumber="407" pagination="326 - 331" refId="ref35401" refString="Wildekamp, R. H. (1977) Nothobranchius lourensi sp. nov. und Nothobranchius janpapi spec. nov., zwei neue Rivulinen aus Ost Afrika. Das Aquarium, 11 (98), 326 - 331." type="journal article" year="1977">Wildekamp, 1977</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3526555FF8ADD53FF1D806537443D4B" box="[151,301,799,825]" pageId="6" pageNumber="407" type="description">
<paragraph id="8BF736DEFF8ADD53FF1D806537443D4B" blockId="6.[151,674,762,825]" box="[151,301,799,825]" pageId="6" pageNumber="407">
(
<figureCitation id="13732A5BFF8ADD53FF158065374D3D4B" box="[159,292,799,825]" captionStart="FIGURE 2" captionStartId="9.[151,250,1410,1434]" captionTargetBox="[151,1435,179,1384]" captionTargetId="figure-19@9.[151,1435,179,1384]" captionTargetPageId="9" captionText="FIGURE 2. Wild-caught specimens of Nothobranchius janpapi and examples of typical habitat. A. N. janpapi Ndundu TAN 95- 10, male, lower Rufiji River drainage; B. N. janpapi Kwaraza TAN 95-14, male, lower Ruvu River drainage, in vicinity of the type locality; C. N. janpapi Soga TAN 97-45, male, Ruvu River drainage; D. N. janpapi Ruvu River TZN 09-4, female, lower Ruvu River drainage (photograph by B. Nagy); E. Location Ruvu River TAN 00-20, a typical thickly vegetated shallow habitat on the floodplain of the lower Ruvu River (photograph June 14, 2000), hosting N. janpapi, N. melanospilus, N. annectens, N. ocellatus and N. eggersi; F. Location Kikale TAN 02-10, thickly vegetated residual pool at a culvert in a seasonal stream course, peripheral to the northern margin of the Rufiji River delta (photograph June 3, 2002), hosting N. janpapi, N. annectens and N. melanospilus. Photographs by B.R. Watters except where otherwise noted." pageId="6" pageNumber="407">Figs. 2A–D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3526555FF8ADD53FF1D801C35763A9B" pageId="6" pageNumber="407" type="materials_examined">
<paragraph id="8BF736DEFF8ADD53FF1D801C32933DBB" blockId="6.[151,1437,870,2013]" pageId="6" pageNumber="407">
<emphasis id="B93CEACCFF8ADD53FF1D801C37E83DF2" bold="true" box="[151,385,870,896]" pageId="6" pageNumber="407">Material examined.</emphasis>
<materialsCitation id="3B203C83FF8ADD53FE0D801D35D93DF3" box="[391,944,871,897]" collectionCode="RMCA" pageId="6" pageNumber="407" specimenCode="RMCA 77-16" specimenCount="1" typeStatus="holotype">
<specimenCode id="DBEE9EA5FF8ADD53FE0D801D344B3DF3" box="[391,546,871,897]" collectionCode="RMCA" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="6" pageNumber="407" type="Museum">RMCA 77-16</specimenCode>
-P-19,
<typeStatus id="54F3887CFF8ADD53FDE7801D34BB3DF3" box="[621,722,871,897]" pageId="6" pageNumber="407" type="holotype">holotype</typeStatus>
, male,
<quantity id="4CB09B3BFF8ADD53FCAB801D35EC3DF3" box="[801,901,871,897]" metricMagnitude="-2" metricUnit="m" metricValue="3.37" pageId="6" pageNumber="407" unit="mm" value="33.7">33.7 mm</quantity>
SL
</materialsCitation>
; –
<materialsCitation id="3B203C83FF8ADD53FC43801D33F13DF3" box="[969,1432,871,897]" collectionCode="RMCA" pageId="6" pageNumber="407" specimenCode="RMCA 77-16" specimenCount="1" typeStatus="holotype">
<specimenCode id="DBEE9EA5FF8ADD53FC43801D320B3DF3" box="[969,1122,871,897]" collectionCode="RMCA" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="6" pageNumber="407" type="Museum">RMCA 77-16</specimenCode>
-P-20, female,
<quantity id="4CB09B3BFF8ADD53FA83801D33043DF3" box="[1289,1389,871,897]" metricMagnitude="-2" metricUnit="m" metricValue="2.8899999999999997" pageId="6" pageNumber="407" unit="mm" value="28.9">28.9 mm</quantity>
SL
</materialsCitation>
; –
<materialsCitation id="3B203C83FF8ADD53FF2F80F136943DBB" collectionCode="RMCA" collectorName="Kwaraza" country="Tanzania" latitude="-6.6966667" location="Ruvu" longLatPrecision="924" longitude="38.7" pageId="6" pageNumber="407" specimenCode="RMCA 77-16" specimenCount="2" specimenCount-female="2" typeStatus="holotype">
<specimenCode id="DBEE9EA5FF8ADD53FF2F80F1372A3DD7" box="[165,323,907,933]" collectionCode="RMCA" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="6" pageNumber="407" type="Museum">RMCA 77-16</specimenCode>
-P-21-23,
<specimenCount id="9D4EFD57FF8ADD53FE3C80F134403DD7" box="[438,553,907,933]" count="2" pageId="6" pageNumber="407" type="female">2 females</specimenCount>
,
<quantity id="4CB09B3BFF8ADD53FDBD80F134B43DD7" box="[567,733,907,933]" metricMagnitude="-2" metricUnit="m" metricValue="2.84" metricValueMax="2.86" metricValueMin="2.82" pageId="6" pageNumber="407" unit="mm" value="28.4" valueMax="28.6" valueMin="28.2">28.2–28.6 mm</quantity>
SL;
<collectingCountry id="F35F764EFF8ADD53FC9380F135EB3DD7" box="[793,898,907,933]" name="Tanzania" pageId="6" pageNumber="407">Tanzania</collectingCountry>
:
<location id="8E976005FF8ADD53FC1A80F135A43DD7" LSID="urn:lsid:plazi:treatment:03E187C8FF8ADD5FFF1D818034D33CDB:8E976005FF8ADD53FC1A80F135A43DD7" box="[912,973,907,933]" country="Tanzania" latitude="-6.6966667" longLatPrecision="924" longitude="38.7" name="Ruvu" pageId="6" pageNumber="407">Ruvu</location>
drainage,
<collectorName id="26BD5308FF8ADD53FBC280F132C73DD7" box="[1096,1198,907,933]" pageId="6" pageNumber="407">Kwaraza</collectorName>
, approx.
<geoCoordinate id="EE7C5019FF8ADD53FA9680F133FE3DD7" box="[1308,1431,907,933]" degrees="06" direction="south" minutes="41.8" orientation="latitude" pageId="6" pageNumber="407" precision="92" value="-6.6966667">06°41.8’ S</geoCoordinate>
,
<geoCoordinate id="EE7C5019FF8ADD53FF1D80D536943DBB" box="[151,253,943,969]" degrees="38" direction="east" minutes="42" orientation="longitude" pageId="6" pageNumber="407" precision="925" value="38.7">38°42’ E</geoCoordinate>
</materialsCitation>
;
<materialsCitation id="3B203C83FF8ADD53FE8380D537C73DBB" box="[265,430,943,969]" collectionCode="R" collectorName="Wildekamp" country="Tanzania" pageId="6" pageNumber="407" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED59AE1BFF8ADD53FE8380D537763DBB" box="[265,287,943,969]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="407">R</collectionCode>
.
<collectorName id="26BD5308FF8ADD53FEA380D537C73DBB" box="[297,430,943,969]" pageId="6" pageNumber="407">Wildekamp</collectorName>
</materialsCitation>
,
<materialsCitation id="3B203C83FF8ADD53FE3280D534D43DBB" box="[440,701,943,969]" collectionCode="T" collectorName="Steinfort, J &amp; Lourens" country="Tanzania" pageId="6" pageNumber="407" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED59AE1BFF8ADD53FE3280D537A33DBB" box="[440,458,943,969]" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="407">T</collectionCode>
.
<collectorName id="26BD5308FF8ADD53FE5F80D5343A3DBB" box="[469,595,943,969]" pageId="6" pageNumber="407">Steinfort, J</collectorName>
.
<collectorName id="26BD5308FF8ADD53FDD480D534D43DBB" box="[606,701,943,969]" pageId="6" pageNumber="407">Lourens</collectorName>
</materialsCitation>
,
<materialsCitation id="3B203C83FF8ADD53FD4D80D532F73DBB" box="[711,1182,943,969]" collectingDate="1976-06-12" collectionCode="R" country="Tanzania" location="Haarlem" pageId="6" pageNumber="407" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED59AE1BFF8ADD53FD4D80D534B43DBB" box="[711,733,943,969]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="407">R</collectionCode>
. van
<location id="8E976005FF8ADD53FC9D80D535123DBB" LSID="urn:lsid:plazi:treatment:03E187C8FF8ADD5FFF1D818034D33CDB:8E976005FF8ADD53FC9D80D535123DBB" box="[791,891,943,969]" country="Tanzania" name="Haarlem" pageId="6" pageNumber="407">Haarlem</location>
;
<date id="FFF6101EFF8ADD53FC0D80D532763DBB" box="[903,1055,943,969]" pageId="6" pageNumber="407" value="1976-06-12">
<collectingDate id="EFB2E9F6FF8ADD53FC0D80D532763DBB" box="[903,1055,943,969]" pageId="6" pageNumber="407" value="1976-06-12">12 June 1976</collectingDate>
</date>
; field code
</materialsCitation>
:
<materialsCitation id="3B203C83FF8ADD53FB2280D5329F3DBB" box="[1192,1270,943,969]" collectionCode="T" country="Tanzania" pageId="6" pageNumber="407" specimenCode="T 76-8" specimenCount="1" typeStatus="holotype">
<specimenCode id="DBEE9EA5FF8ADD53FB2280D5329F3DBB" box="[1192,1270,943,969]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="407">T 76-8</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BF736DEFF8ADD53FF4D80A835763A9B" blockId="6.[151,1437,870,2013]" pageId="6" pageNumber="407">
<emphasis id="B93CEACCFF8ADD53FF4D80A834573D9E" bold="true" box="[199,574,978,1004]" pageId="6" pageNumber="407">Additional material examined.</emphasis>
<materialsCitation id="3B203C83FF8ADD53FDC380A935003A63" collectingDate="2009-01-12" collectorName="B. Nagy" country="Tanzania" latitude="-6.6802998" location="Ruvu" longLatPrecision="1" longitude="38.66382" municipality="Kwaraza" pageId="6" pageNumber="407" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D4EFD57FF8ADD53FDC380A934F53D9F" box="[585,668,979,1005]" count="1" pageId="6" pageNumber="407" type="male">1 male</specimenCount>
,
<quantity id="4CB09B3BFF8ADD53FD2680A9357C3D9F" box="[684,789,979,1005]" metricMagnitude="-2" metricUnit="m" metricValue="2.77" pageId="6" pageNumber="407" unit="mm" value="27.7">27.7 mm</quantity>
SL;
<collectingCountry id="F35F764EFF8ADD53FCDE80A935D43D9F" box="[852,957,979,1005]" name="Tanzania" pageId="6" pageNumber="407">Tanzania</collectingCountry>
:
<location id="8E976005FF8ADD53FC4680A932603D9F" LSID="urn:lsid:plazi:treatment:03E187C8FF8ADD5FFF1D818034D33CDB:8E976005FF8ADD53FC4680A932603D9F" box="[972,1033,979,1005]" country="Tanzania" latitude="-6.6802998" longLatPrecision="1" longitude="38.66382" municipality="Kwaraza" name="Ruvu" pageId="6" pageNumber="407">Ruvu</location>
drainage,
<collectingMunicipality id="6B93ACA4FF8ADD53FB0D80A932873D9F" box="[1159,1262,979,1005]" pageId="6" pageNumber="407">Kwaraza</collectingMunicipality>
;
<geoCoordinate id="EE7C5019FF8ADD53FB7480A933FE3D9F" box="[1278,1431,979,1005]" degrees="06" direction="south" minutes="40.818" orientation="latitude" pageId="6" pageNumber="407" precision="1" value="-6.6802998">06°40.818’ S</geoCoordinate>
,
<geoCoordinate id="EE7C5019FF8ADD53FF1D808D37463A63" box="[151,303,1015,1041]" degrees="38" direction="east" minutes="39.829" orientation="longitude" pageId="6" pageNumber="407" precision="1" value="38.66382">38°39.829’ E</geoCoordinate>
;
<collectorName id="26BD5308FF8ADD53FEB6808D37F43A63" box="[316,413,1015,1041]" pageId="6" pageNumber="407">B. Nagy</collectorName>
;
<date id="FFF6101EFF8ADD53FE20808D34013A63" box="[426,616,1015,1041]" pageId="6" pageNumber="407" value="2009-01-12">
<collectingDate id="EFB2E9F6FF8ADD53FE20808D34013A63" box="[426,616,1015,1041]" pageId="6" pageNumber="407" value="2009-01-12">12 January 2009</collectingDate>
</date>
; field code: TZN 09-4
</materialsCitation>
. –
<materialsCitation id="3B203C83FF8ADD53FC0D808D36DF3A2B" collectingDate="1998-05-29" collectorName="B. Watters &amp; J. Rosenstock &amp; M. Agnew &amp; Larsen" country="Tanzania" latitude="-6.8356667" location="Ruvu" longLatPrecision="13" longitude="38.84667" municipality="Soga" pageId="6" pageNumber="407" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D4EFD57FF8ADD53FC0D808D35BB3A63" box="[903,978,1015,1041]" count="1" pageId="6" pageNumber="407" type="male">1 male</specimenCount>
,
<quantity id="4CB09B3BFF8ADD53FC55808D322D3A63" box="[991,1092,1015,1041]" metricMagnitude="-2" metricUnit="m" metricValue="2.84" pageId="6" pageNumber="407" unit="mm" value="28.4">28.4 mm</quantity>
SL;
<collectingCountry id="F35F764EFF8ADD53FBF6808D328C3A63" box="[1148,1253,1015,1041]" name="Tanzania" pageId="6" pageNumber="407">Tanzania</collectingCountry>
:
<location id="8E976005FF8ADD53FB7A808D33443A63" LSID="urn:lsid:plazi:treatment:03E187C8FF8ADD5FFF1D818034D33CDB:8E976005FF8ADD53FB7A808D33443A63" box="[1264,1325,1015,1041]" country="Tanzania" latitude="-6.8356667" longLatPrecision="13" longitude="38.84667" municipality="Soga" name="Ruvu" pageId="6" pageNumber="407">Ruvu</location>
drainage,
<collectingMunicipality id="6B93ACA4FF8ADD53FF1D876136BB3A47" box="[151,210,1051,1077]" pageId="6" pageNumber="407">Soga</collectingMunicipality>
;
<geoCoordinate id="EE7C5019FF8ADD53FF578760370B3A47" box="[221,354,1050,1077]" degrees="06" direction="south" minutes="50.14" orientation="latitude" pageId="6" pageNumber="407" precision="9" value="-6.8356667">06°50.14’ S</geoCoordinate>
,
<geoCoordinate id="EE7C5019FF8ADD53FEE78760379C3A47" box="[365,501,1050,1077]" degrees="38" direction="east" minutes="50.80" orientation="longitude" pageId="6" pageNumber="407" precision="9" value="38.84667">38°50.80’ E</geoCoordinate>
;
<collectorName id="26BD5308FF8ADD53FD8A8761341E3A47" box="[512,631,1051,1077]" pageId="6" pageNumber="407">B. Watters</collectorName>
,
<collectorName id="26BD5308FF8ADD53FD0B876135743A47" box="[641,797,1051,1077]" pageId="6" pageNumber="407">J. Rosenstock</collectorName>
,
<collectorName id="26BD5308FF8ADD53FCAD876135F73A47" box="[807,926,1051,1077]" pageId="6" pageNumber="407">M. Agnew</collectorName>
, F.
<collectorName id="26BD5308FF8ADD53FC488761327B3A47" box="[962,1042,1051,1077]" pageId="6" pageNumber="407">Larsen</collectorName>
;
<date id="FFF6101EFF8ADD53FB96876132DA3A47" box="[1052,1203,1051,1077]" pageId="6" pageNumber="407" value="1998-05-29">
<collectingDate id="EFB2E9F6FF8ADD53FB96876132DA3A47" box="[1052,1203,1051,1077]" pageId="6" pageNumber="407" value="1998-05-29">29 May 1998</collectingDate>
</date>
; field code: TAN 98- 14
</materialsCitation>
. –
<materialsCitation id="3B203C83FF8ADD53FF59874534FB3A0F" collectingDate="1997-06-12" collectionCode="R" collectorName="B. Watters &amp; Wildekamp, B &amp; Cooper" country="Tanzania" latitude="-6.8365" location="Ruvu" longLatPrecision="13" longitude="38.846333" municipality="Soga" pageId="6" pageNumber="407" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D4EFD57FF8ADD53FF59874537743A2B" box="[211,285,1087,1113]" count="1" pageId="6" pageNumber="407" type="male">1 male</specimenCount>
,
<quantity id="4CB09B3BFF8ADD53FEA2874537E23A2B" box="[296,395,1087,1113]" metricMagnitude="-2" metricUnit="m" metricValue="3.26" pageId="6" pageNumber="407" unit="mm" value="32.6">32.6 mm</quantity>
SL;
<collectingCountry id="F35F764EFF8ADD53FE4A874534403A2B" box="[448,553,1087,1113]" name="Tanzania" pageId="6" pageNumber="407">Tanzania</collectingCountry>
:
<location id="8E976005FF8ADD53FDBE874534183A2B" LSID="urn:lsid:plazi:treatment:03E187C8FF8ADD5FFF1D818034D33CDB:8E976005FF8ADD53FDBE874534183A2B" box="[564,625,1087,1113]" country="Tanzania" latitude="-6.8365" longLatPrecision="13" longitude="38.846333" municipality="Soga" name="Ruvu" pageId="6" pageNumber="407">Ruvu</location>
drainage,
<collectingMunicipality id="6B93ACA4FF8ADD53FD6F874535493A2B" box="[741,800,1087,1113]" pageId="6" pageNumber="407">Soga</collectingMunicipality>
;
<geoCoordinate id="EE7C5019FF8ADD53FCA6874435D83A2B" box="[812,945,1086,1113]" degrees="06" direction="south" minutes="50.19" orientation="latitude" pageId="6" pageNumber="407" precision="9" value="-6.8365">06°50.19’ S</geoCoordinate>
,
<geoCoordinate id="EE7C5019FF8ADD53FC378744322C3A2B" box="[957,1093,1086,1113]" degrees="38" direction="east" minutes="50.78" orientation="longitude" pageId="6" pageNumber="407" precision="9" value="38.846333">38°50.78’ E</geoCoordinate>
;
<collectorName id="26BD5308FF8ADD53FBDB874532A03A2B" box="[1105,1225,1087,1113]" pageId="6" pageNumber="407">B. Watters</collectorName>
,
<collectionCode id="ED59AE1BFF8ADD53FB59874532803A2B" box="[1235,1257,1087,1113]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="407">R</collectionCode>
.
<collectorName id="26BD5308FF8ADD53FB79874533F13A2B" box="[1267,1432,1087,1113]" pageId="6" pageNumber="407">Wildekamp, B</collectorName>
.
<collectorName id="26BD5308FF8ADD53FF1D871936843A0F" box="[151,237,1123,1149]" pageId="6" pageNumber="407">Cooper</collectorName>
;
<date id="FFF6101EFF8ADD53FF7D871937E53A0F" box="[247,396,1123,1149]" pageId="6" pageNumber="407" value="1997-06-12">
<collectingDate id="EFB2E9F6FF8ADD53FF7D871937E53A0F" box="[247,396,1123,1149]" pageId="6" pageNumber="407" value="1997-06-12">12 June 1997</collectingDate>
</date>
; field code: TAN 97-45
</materialsCitation>
. –
<materialsCitation id="3B203C83FF8ADD53FD27871932E83AD3" collectingDate="2003-05-29" collectorName="B. Watters &amp; B. Cooper &amp; W. Bishopp" country="Tanzania" latitude="-8.051833" location="Rufiji" longLatPrecision="1" longitude="39.0085" municipality="Kitonga" pageId="6" pageNumber="407" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D4EFD57FF8ADD53FD278719349F3A0F" box="[685,758,1123,1149]" count="1" pageId="6" pageNumber="407" type="male">1 male</specimenCount>
, 32.0 mm SL;
<collectingCountry id="F35F764EFF8ADD53FC1F871935973A0F" box="[917,1022,1123,1149]" name="Tanzania" pageId="6" pageNumber="407">Tanzania</collectingCountry>
:
<location id="8E976005FF8ADD53FB8D871932213A0F" LSID="urn:lsid:plazi:treatment:03E187C8FF8ADD5FFF1D818034D33CDB:8E976005FF8ADD53FB8D871932213A0F" box="[1031,1096,1123,1149]" country="Tanzania" latitude="-8.051833" longLatPrecision="1" longitude="39.0085" municipality="Kitonga" name="Rufiji" pageId="6" pageNumber="407">Rufiji</location>
drainage,
<collectingMunicipality id="6B93ACA4FF8ADD53FB30871933713A0F" box="[1210,1304,1123,1149]" pageId="6" pageNumber="407">Kitonga</collectingMunicipality>
;
<geoCoordinate id="EE7C5019FF8ADD53FAA8871936C13AD3" degrees="08" direction="south" minutes="03.110" orientation="latitude" pageId="6" pageNumber="407" precision="1" value="-8.051833">08°03.110’ S</geoCoordinate>
,
<geoCoordinate id="EE7C5019FF8ADD53FF3E87FC37203AD3" box="[180,329,1158,1185]" degrees="39" direction="east" minutes="00.510" orientation="longitude" pageId="6" pageNumber="407" precision="1" value="39.0085">39°00.510’ E</geoCoordinate>
;
<collectorName id="26BD5308FF8ADD53FEDE87FD37A23AD3" box="[340,459,1159,1185]" pageId="6" pageNumber="407">B. Watters</collectorName>
,
<collectorName id="26BD5308FF8ADD53FE5F87FD34203AD3" box="[469,585,1159,1185]" pageId="6" pageNumber="407">B. Cooper</collectorName>
,
<collectorName id="26BD5308FF8ADD53FDD887FD34B13AD3" box="[594,728,1159,1185]" pageId="6" pageNumber="407">W. Bishopp</collectorName>
;
<date id="FFF6101EFF8ADD53FD6887FD35103AD3" box="[738,889,1159,1185]" pageId="6" pageNumber="407" value="2003-05-29">
<collectingDate id="EFB2E9F6FF8ADD53FD6887FD35103AD3" box="[738,889,1159,1185]" pageId="6" pageNumber="407" value="2003-05-29">29 May 2003</collectingDate>
</date>
; field code: TAN 03-22
</materialsCitation>
. –
<specimenCount id="9D4EFD57FF8ADD53FB1087FC32993AD3" box="[1178,1264,1158,1185]" count="5" pageId="6" pageNumber="407" type="male">5 males</specimenCount>
,
<quantity id="4CB09B3BFF8ADD53FB7187FC33F43AD3" box="[1275,1437,1158,1185]" metricMagnitude="-2" metricUnit="m" metricValue="2.3899999999999997" metricValueMax="2.56" metricValueMin="2.2199999999999998" pageId="6" pageNumber="407" unit="mm" value="23.9" valueMax="25.6" valueMin="22.2">22.2–25.6 mm</quantity>
SL,
<materialsCitation id="3B203C83FF8ADD53FF4F87D135723A9B" collectingDate="2002-06-04" collectorName="B. Watters &amp; B. Cooper &amp; O. Schmidt" country="Tanzania" latitude="-7.8012166" location="Ruhoi" longLatPrecision="1" longitude="38.812317" municipality="Ngulakula" pageId="6" pageNumber="407" specimenCount="2" specimenCount-female="2">
<specimenCount id="9D4EFD57FF8ADD53FF4F87D1375A3AB7" box="[197,307,1195,1221]" count="2" pageId="6" pageNumber="407" type="female">2 females</specimenCount>
,
<quantity id="4CB09B3BFF8ADD53FEB487D037893AB7" box="[318,480,1194,1221]" metricMagnitude="-2" metricUnit="m" metricValue="2.23" metricValueMax="2.35" metricValueMin="2.11" pageId="6" pageNumber="407" unit="mm" value="22.3" valueMax="23.5" valueMin="21.1">21.1–23.5 mm</quantity>
SL;
<collectingCountry id="F35F764EFF8ADD53FD9E87D134143AB7" box="[532,637,1195,1221]" name="Tanzania" pageId="6" pageNumber="407">Tanzania</collectingCountry>
:
<location id="8E976005FF8ADD53FD0D87D134A23AB7" LSID="urn:lsid:plazi:treatment:03E187C8FF8ADD5FFF1D818034D33CDB:8E976005FF8ADD53FD0D87D134A23AB7" box="[647,715,1195,1221]" country="Tanzania" latitude="-7.8012166" longLatPrecision="1" longitude="38.812317" municipality="Ngulakula" name="Ruhoi" pageId="6" pageNumber="407">Ruhoi</location>
drainage,
<collectingMunicipality id="6B93ACA4FF8ADD53FCB587D135DE3AB7" box="[831,951,1195,1221]" pageId="6" pageNumber="407">Ngulakula</collectingMunicipality>
;
<geoCoordinate id="EE7C5019FF8ADD53FC4887D1323D3AB7" box="[962,1108,1195,1221]" degrees="07" direction="south" minutes="48.073" orientation="latitude" pageId="6" pageNumber="407" precision="1" value="-7.8012166">07°48.073’ S</geoCoordinate>
,
<geoCoordinate id="EE7C5019FF8ADD53FBEA87D1329F3AB7" box="[1120,1270,1195,1221]" degrees="38" direction="east" minutes="48.739" orientation="longitude" pageId="6" pageNumber="407" precision="1" value="38.812317">38°48.739’ E</geoCoordinate>
;
<collectorName id="26BD5308FF8ADD53FA8B87D133113AB7" box="[1281,1400,1195,1221]" pageId="6" pageNumber="407">B. Watters</collectorName>
,
<collectorName id="26BD5308FF8ADD53FA0887D136823A9B" pageId="6" pageNumber="407">B. Cooper</collectorName>
,
<collectorName id="26BD5308FF8ADD53FF7C87B537133A9B" box="[246,378,1231,1257]" pageId="6" pageNumber="407">O. Schmidt</collectorName>
;
<date id="FFF6101EFF8ADD53FE0F87B534793A9B" box="[389,528,1231,1257]" pageId="6" pageNumber="407" value="2002-06-04">
<collectingDate id="EFB2E9F6FF8ADD53FE0F87B534793A9B" box="[389,528,1231,1257]" pageId="6" pageNumber="407" value="2002-06-04">4 June 2002</collectingDate>
</date>
; field code: TAN 02-13
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3526555FF8ADD53FF4D8788321839E7" pageId="6" pageNumber="407" type="diagnosis">
<paragraph id="8BF736DEFF8ADD53FF4D8788321839E7" blockId="6.[151,1437,870,2013]" pageId="6" pageNumber="407">
<emphasis id="B93CEACCFF8ADD53FF4D8788372B3B7E" bold="true" box="[199,322,1266,1292]" pageId="6" pageNumber="407">Diagnosis.</emphasis>
<taxonomicName id="4C484D5DFF8ADD53FECC8789343D3B7E" authorityName="Wildekamp" authorityYear="1977" box="[326,596,1267,1293]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF8ADD53FECC8789343D3B7E" box="[326,596,1267,1293]" italics="true" pageId="6" pageNumber="407">Nothobranchius janpapi</emphasis>
</taxonomicName>
belongs to the
<taxonomicName id="4C484D5DFF8ADD53FD7E878935013B7E" authorityName="Wildekamp" authorityYear="1977" box="[756,872,1267,1292]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF8ADD53FD7E878935013B7E" box="[756,872,1267,1292]" italics="true" pageId="6" pageNumber="407">N. janpapi</emphasis>
</taxonomicName>
species group within the subgenus
<taxonomicName id="4C484D5DFF8ADD53FB6F878933F43B7F" authorityName="Wildekamp" authorityYear="1977" box="[1253,1437,1267,1293]" class="Actinopterygii" family="Nothobranchiidae" genus="Aphyobranchius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyprinodontiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="genus">
<emphasis id="B93CEACCFF8ADD53FB6F878933F43B7F" box="[1253,1437,1267,1293]" italics="true" pageId="6" pageNumber="407">Aphyobranchius</emphasis>
</taxonomicName>
by presenting the following combination of characters in the male: light blue body colouration with narrow red scale margins and yellow-brown to yellow-green dorsum stripe; rearward positioned dorsal fin with predorsal length of 63.3–67.9 % SL and predorsal length 109–115 as percentage of preanal length; dorsal fin relatively small with low number of rays, 12–14; cephalic sensory system at supraorbital level in an undivided, slightly curved, almost flat, shallow groove with three exposed neuromasts and pronounced lobes (
<figureCitation id="13732A5BFF8ADD53FC2786DD35963BB3" box="[941,1023,1447,1473]" captionStart="FIGURE 3" captionStartId="10.[151,250,1307,1331]" captionTargetBox="[156,1428,706,1276]" captionTargetId="figure-280@10.[151,1436,702,1281]" captionTargetPageId="10" captionText="FIGURE 3. Cephalic structure: a, Nothobranchius janpapi; b, N. luekei; c, N. geminus; d, N. lourensi and N. fuscotaeniatus. Key to label abbreviations: an, anterior nostril; pn, posterior nostril; f, frontal neuromast; so, supraorbital canal; st, supratemporal canal." pageId="6" pageNumber="407">Fig. 3a</figureCitation>
). Both sexes with central 2–3 rays of the caudal fin slightly elongated forming a mild posterior lobe (vs. central lobe absent).
<taxonomicName id="4C484D5DFF8ADD53FBFB86B133EA3B96" authorityName="Wildekamp" authorityYear="1977" box="[1137,1411,1483,1509]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF8ADD53FBFB86B133EA3B96" box="[1137,1411,1483,1509]" italics="true" pageId="6" pageNumber="407">Nothobranchius janpapi</emphasis>
</taxonomicName>
is distinguished from the other members of the
<taxonomicName id="4C484D5DFF8ADD53FD1886953561387A" authorityName="Wildekamp" authorityYear="1977" box="[658,776,1519,1544]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF8ADD53FD1886953561387A" box="[658,776,1519,1544]" italics="true" pageId="6" pageNumber="407">N. janpapi</emphasis>
</taxonomicName>
species group by male colouration of anal fin with slender orange-red distal band and narrow light blue margin (vs. slender yellow subdistal band with narrow red distal band); caudal fin base and proximal zone olive, with light grey to translucent subdistal band and narrow dark grey distal band (vs. base and proximal zone grey to light red with slender red subdistal band and narrow black distal band in
<taxonomicName id="4C484D5DFF8ADD53FF1D8505369438EB" authorityName="Seegers" authorityYear="1984" box="[151,253,1663,1689]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="luekei">
<emphasis id="B93CEACCFF8ADD53FF1D8505369438EB" box="[151,253,1663,1689]" italics="true" pageId="6" pageNumber="407">N. luekei</emphasis>
</taxonomicName>
, and caudal fin plain red to orange-red with narrow black distal margin in
<taxonomicName id="4C484D5DFF8ADD53FBD1850532B238EA" authorityName="Wildekamp, Watters and Sainthouse" authorityYear="2002" box="[1115,1243,1663,1688]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="geminus">
<emphasis id="B93CEACCFF8ADD53FBD1850532B238EA" box="[1115,1243,1663,1688]" italics="true" pageId="6" pageNumber="407">N. geminus</emphasis>
</taxonomicName>
); dorsal fin with narrow light-blue distal band (vs. slender red in
<taxonomicName id="4C484D5DFF8ADD53FD3785D9354B38CF" authorityName="Seegers" authorityYear="1984" box="[701,802,1699,1725]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="luekei">
<emphasis id="B93CEACCFF8ADD53FD3785D9354B38CF" box="[701,802,1699,1725]" italics="true" pageId="6" pageNumber="407">N. luekei</emphasis>
</taxonomicName>
and narrow black in
<taxonomicName id="4C484D5DFF8ADD53FB9C85D932FC38CE" authorityName="Wildekamp, Watters and Sainthouse" authorityYear="2002" box="[1046,1173,1699,1724]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="geminus">
<emphasis id="B93CEACCFF8ADD53FB9C85D932FC38CE" box="[1046,1173,1699,1724]" italics="true" pageId="6" pageNumber="407">N. geminus</emphasis>
</taxonomicName>
); pelvic fin orange-red with narrow light blue distal band (vs. light blue with narrow red distal margin in
<taxonomicName id="4C484D5DFF8ADD53FBA485BD32F83893" authorityName="Seegers" authorityYear="1984" box="[1070,1169,1735,1761]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="luekei">
<emphasis id="B93CEACCFF8ADD53FBA485BD32F83893" box="[1070,1169,1735,1761]" italics="true" pageId="6" pageNumber="407">N. luekei</emphasis>
</taxonomicName>
, and yellow to red with narrow black margin in
<taxonomicName id="4C484D5DFF8ADD53FE2C8591344C3976" authorityName="Wildekamp, Watters and Sainthouse" authorityYear="2002" box="[422,549,1771,1796]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="geminus">
<emphasis id="B93CEACCFF8ADD53FE2C8591344C3976" box="[422,549,1771,1796]" italics="true" pageId="6" pageNumber="407">N. geminus</emphasis>
</taxonomicName>
); as well as morphometric characters in males of head depth 74–81 % HL (vs.
<quantity id="4CB09B3BFF8ADD53FF1D84753690395B" box="[151,249,1807,1833]" metricMagnitude="0" metricUnit="m" metricValue="1.7526" metricValueMax="1.8288" metricValueMin="1.6764" pageId="6" pageNumber="407" unit="in" value="69.0" valueMax="72.0" valueMin="66.0">66–72 in</quantity>
<taxonomicName id="4C484D5DFF8ADD53FF7584753715395A" authorityName="Wildekamp, Watters and Sainthouse" authorityYear="2002" box="[255,380,1807,1832]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="geminus">
<emphasis id="B93CEACCFF8ADD53FF7584753715395A" box="[255,380,1807,1832]" italics="true" pageId="6" pageNumber="407">N. geminus</emphasis>
</taxonomicName>
and
<quantity id="4CB09B3BFF8ADD53FE3B8474347A395B" box="[433,531,1806,1833]" metricMagnitude="0" metricUnit="m" metricValue="1.4986000000000002" metricValueMax="1.6002000000000003" metricValueMin="1.3969999999999998" pageId="6" pageNumber="407" unit="in" value="59.0" valueMax="63.0" valueMin="55.0">55–63 in</quantity>
<taxonomicName id="4C484D5DFF8ADD53FD9284753412395B" authorityName="Seegers" authorityYear="1984" box="[536,635,1807,1833]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="luekei">
<emphasis id="B93CEACCFF8ADD53FD9284753412395B" box="[536,635,1807,1833]" italics="true" pageId="6" pageNumber="407">N. luekei</emphasis>
</taxonomicName>
); suborbital depth 16–20 % HL (vs.
<quantity id="4CB09B3BFF8ADD53FB8584753218395B" box="[1039,1137,1807,1833]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" metricValueMax="3.556" metricValueMin="2.54" pageId="6" pageNumber="407" unit="in" value="12.0" valueMax="14.0" valueMin="10.0">10–14 in</quantity>
<taxonomicName id="4C484D5DFF8ADD53FBFD8475329D395A" authorityName="Wildekamp, Watters and Sainthouse" authorityYear="2002" box="[1143,1268,1807,1832]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="geminus">
<emphasis id="B93CEACCFF8ADD53FBFD8475329D395A" box="[1143,1268,1807,1832]" italics="true" pageId="6" pageNumber="407">N. geminus</emphasis>
</taxonomicName>
and
<quantity id="4CB09B3BFF8ADD53FAA384753314395B" box="[1321,1405,1807,1833]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" metricValueMax="3.048" metricValueMin="2.032" pageId="6" pageNumber="407" unit="in" value="10.0" valueMax="12.0" valueMin="8.0">8–12 in</quantity>
<taxonomicName id="4C484D5DFF8ADD53FA09847536B3393F" authorityName="Seegers" authorityYear="1984" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="luekei">
<emphasis id="B93CEACCFF8ADD53FA09847536B3393F" italics="true" pageId="6" pageNumber="407">N. luekei</emphasis>
</taxonomicName>
); and caudal peduncle length
<quantity id="4CB09B3BFF8ADD53FDAE844934C8393F" box="[548,673,1843,1869]" metricMagnitude="0" metricUnit="m" metricValue="3.4671" metricValueMax="3.556" metricValueMin="3.3781999999999996" pageId="6" pageNumber="407" unit="in" value="136.5" valueMax="140.0" valueMin="133.0">133–140 in</quantity>
% of its depth (vs.
<quantity id="4CB09B3BFF8ADD53FCF984493598393F" box="[883,1009,1843,1869]" metricMagnitude="0" metricUnit="m" metricValue="3.1877" metricValueMax="3.3019999999999996" metricValueMin="3.0734" pageId="6" pageNumber="407" unit="in" value="125.5" valueMax="130.0" valueMin="121.0">121–130 in</quantity>
<taxonomicName id="4C484D5DFF8ADD53FC7C8449321A393E" authorityName="Wildekamp, Watters and Sainthouse" authorityYear="2002" box="[1014,1139,1843,1868]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="geminus">
<emphasis id="B93CEACCFF8ADD53FC7C8449321A393E" box="[1014,1139,1843,1868]" italics="true" pageId="6" pageNumber="407">N. geminus</emphasis>
</taxonomicName>
and
<quantity id="4CB09B3BFF8ADD53FB2D8448334D393F" box="[1191,1316,1842,1869]" metricMagnitude="0" metricUnit="m" metricValue="3.8608" metricValueMax="3.9624" metricValueMin="3.7592" pageId="6" pageNumber="407" unit="in" value="152.0" valueMax="156.0" valueMin="148.0">148–156 in</quantity>
<taxonomicName id="4C484D5DFF8ADD53FAA0844933E5393F" authorityName="Seegers" authorityYear="1984" box="[1322,1420,1843,1869]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="luekei">
<emphasis id="B93CEACCFF8ADD53FAA0844933E5393F" box="[1322,1420,1843,1869]" italics="true" pageId="6" pageNumber="407">N. luekei</emphasis>
</taxonomicName>
). Further, it differs from
<taxonomicName id="4C484D5DFF8ADD53FE17842D34723902" authorityName="Wildekamp, Watters and Sainthouse" authorityYear="2002" box="[413,539,1879,1904]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="geminus">
<emphasis id="B93CEACCFF8ADD53FE17842D34723902" box="[413,539,1879,1904]" italics="true" pageId="6" pageNumber="407">N. geminus</emphasis>
</taxonomicName>
by dorsal-fin base length 18.8–25.1 % SL vs. 15.6–18.6, and from
<taxonomicName id="4C484D5DFF8ADD53FA9E842D33103903" authorityName="Seegers" authorityYear="1984" box="[1300,1401,1879,1905]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="luekei">
<emphasis id="B93CEACCFF8ADD53FA9E842D33103903" box="[1300,1401,1879,1905]" italics="true" pageId="6" pageNumber="407">N. luekei</emphasis>
</taxonomicName>
by predorsal length 63.3–67.9 % SL vs. 68.0–73.0, and head width 58–65 % HL vs. 52–56.
</paragraph>
</subSubSection>
<subSubSection id="C3526555FF8ADD5CFF4D84E4322D3954" lastPageId="9" lastPageNumber="410" pageId="6" pageNumber="407" type="description">
<paragraph id="8BF736DEFF8ADD53FF4D84E434F439AF" blockId="6.[151,1437,870,2013]" pageId="6" pageNumber="407">
<emphasis id="B93CEACCFF8ADD53FF4D84E4373039CA" bold="true" box="[199,345,1950,1976]" pageId="6" pageNumber="407">Description.</emphasis>
General body features illustrated by
<figureCitation id="13732A5BFF8ADD53FC8584E535DF39CB" box="[783,950,1951,1977]" captionStart="FIGURE 2" captionStartId="9.[151,250,1410,1434]" captionTargetBox="[151,1435,179,1384]" captionTargetId="figure-19@9.[151,1435,179,1384]" captionTargetPageId="9" captionText="FIGURE 2. Wild-caught specimens of Nothobranchius janpapi and examples of typical habitat. A. N. janpapi Ndundu TAN 95- 10, male, lower Rufiji River drainage; B. N. janpapi Kwaraza TAN 95-14, male, lower Ruvu River drainage, in vicinity of the type locality; C. N. janpapi Soga TAN 97-45, male, Ruvu River drainage; D. N. janpapi Ruvu River TZN 09-4, female, lower Ruvu River drainage (photograph by B. Nagy); E. Location Ruvu River TAN 00-20, a typical thickly vegetated shallow habitat on the floodplain of the lower Ruvu River (photograph June 14, 2000), hosting N. janpapi, N. melanospilus, N. annectens, N. ocellatus and N. eggersi; F. Location Kikale TAN 02-10, thickly vegetated residual pool at a culvert in a seasonal stream course, peripheral to the northern margin of the Rufiji River delta (photograph June 3, 2002), hosting N. janpapi, N. annectens and N. melanospilus. Photographs by B.R. Watters except where otherwise noted." pageId="6" pageNumber="407">Figures 2A–D</figureCitation>
. Morphometric and meristic characters of examined material are summarized in
<tableCitation id="C6CA0365FF8ADD53FDC984B934F139AF" box="[579,664,1987,2013]" captionStart="TABLE 3" captionStartId="7.[151,239,151,175]" captionText="TABLE 3. Morphometric and meristic data of Nothobranchius janpapi. SD, standard deviation." pageId="6" pageNumber="407">Table 3</tableCitation>
.
</paragraph>
<caption id="DF376656FF8BDD52FF1D83ED32F13EDD" box="[151,1176,151,175]" pageId="7" pageNumber="408" startId="7.[151,239,151,175]" targetType="table">
<paragraph id="8BF736DEFF8BDD52FF1D83ED32F13EDD" blockId="7.[151,1176,151,175]" box="[151,1176,151,175]" pageId="7" pageNumber="408">
<emphasis id="B93CEACCFF8BDD52FF1D83ED37633EDD" bold="true" box="[151,266,151,175]" pageId="7" pageNumber="408">TABLE 3.</emphasis>
Morphometric and meristic data of
<taxonomicName id="4C484D5DFF8BDD52FD0183ED35E63EDD" authorityName="Wildekamp" authorityYear="1977" box="[651,911,151,175]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="408" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF8BDD52FD0183ED35E63EDD" box="[651,911,151,175]" italics="true" pageId="7" pageNumber="408">Nothobranchius janpapi</emphasis>
</taxonomicName>
. SD, standard deviation.
</paragraph>
</caption>
<paragraph id="8BF736DEFF8BDD52FCE283C5337B38F2" pageId="7" pageNumber="408">
<table id="F948C47EFF8B22AAFF1383C5331938F2" box="[153,1392,191,1664]" gridcols="7" gridrows="36" pageId="7" pageNumber="408">
<tr id="3578349CFF8B22AAFF1383C533193EA5" box="[153,1392,191,215]" gridrow="0" pageId="7" pageNumber="408" rowspan-0="1" rowspan-3="1" rowspan-6="1">
<th id="76A95DE0FF8B22AAFD6C83C535A93EA5" box="[742,960,191,215]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="7" pageNumber="408">males</th>
<th id="76A95DE0FF8B22AAFBEE83C5335B3EA5" box="[1124,1330,191,215]" colspan="2" colspanRight="1" gridcol="4" gridrow="0" pageId="7" pageNumber="408">females</th>
</tr>
<tr id="3578349CFF8B22AAFF13839D33193F72" box="[153,1392,231,256]" gridrow="1" pageId="7" pageNumber="408" rowspan-0="1" rowspan-3="1" rowspan-6="1">
<td id="76A95DE0FF8B22AAFD6C839D35A93F72" box="[742,960,231,256]" colspan="2" colspanRight="1" gridcol="1" gridrow="1" pageId="7" pageNumber="408">(n = 10)</td>
<td id="76A95DE0FF8B22AAFBEE839D335B3F72" box="[1124,1330,231,256]" colspan="2" colspanRight="1" gridcol="4" gridrow="1" pageId="7" pageNumber="408">(n = 5)</td>
</tr>
<tr id="3578349CFF8B22AAFF13824233193F23" box="[153,1392,312,337]" gridrow="2" pageId="7" pageNumber="408" rowspan-0="1">
<td id="76A95DE0FF8B22AAFD6C8242350C3F23" box="[742,869,312,337]" gridcol="1" gridrow="2" pageId="7" pageNumber="408">range</td>
<td id="76A95DE0FF8B22AAFC0D824235A93F23" box="[903,960,312,337]" gridcol="2" gridrow="2" pageId="7" pageNumber="408">mean</td>
<td id="76A95DE0FF8B22AAFC6A824232693F23" box="[992,1024,312,337]" gridcol="3" gridrow="2" pageId="7" pageNumber="408">SD</td>
<td id="76A95DE0FF8B22AAFBEE8242328A3F23" box="[1124,1251,312,337]" gridcol="4" gridrow="2" pageId="7" pageNumber="408">range</td>
<td id="76A95DE0FF8B22AAFB738242335B3F23" box="[1273,1330,312,337]" gridcol="5" gridrow="2" pageId="7" pageNumber="408">mean</td>
<td id="76A95DE0FF8B22AAFADA824233193F23" box="[1360,1392,312,337]" gridcol="6" gridrow="2" pageId="7" pageNumber="408">SD</td>
</tr>
<tr id="3578349CFF8B22AAFF13821A33193F0A" box="[153,1392,352,376]" gridrow="3" pageId="7" pageNumber="408" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1">
<th id="76A95DE0FF8B22AAFF13821A34BF3F0A" box="[153,726,352,376]" gridcol="0" gridrow="3" pageId="7" pageNumber="408">Standard length (mm)</th>
<td id="76A95DE0FF8B22AAFD6C821A350C3F0A" box="[742,869,352,376]" gridcol="1" gridrow="3" pageId="7" pageNumber="408">22.2–33.7</td>
<td id="76A95DE0FF8B22AAFBEE821A328A3F0A" box="[1124,1251,352,376]" gridcol="4" gridrow="3" pageId="7" pageNumber="408">21.1–28.9</td>
</tr>
<tr id="3578349CFF8B22AAFF1382F233193FD2" box="[153,1392,392,416]" gridrow="4" pageId="7" pageNumber="408">
<th id="76A95DE0FF8B22AAFF1382F233193FD2" box="[153,1392,392,416]" colspan="7" colspanRight="6" gridcol="0" gridrow="4" pageId="7" pageNumber="408">
<emphasis id="B93CEACCFF8BDD52FF1382F237D13FD2" bold="true" box="[153,440,392,416]" pageId="7" pageNumber="408">Percent of standard length</emphasis>
</th>
</tr>
<tr id="3578349CFF8B22AAFF1382CB33193FBB" box="[153,1392,433,457]" gridrow="5" pageId="7" pageNumber="408">
<th id="76A95DE0FF8B22AAFF1382CB34BF3FBB" box="[153,726,433,457]" gridcol="0" gridrow="5" pageId="7" pageNumber="408">Total length</th>
<td id="76A95DE0FF8B22AAFD6C82CB350C3FBB" box="[742,869,433,457]" gridcol="1" gridrow="5" pageId="7" pageNumber="408">124.9–130.6</td>
<td id="76A95DE0FF8B22AAFC0D82CB35A93FBB" box="[903,960,433,457]" gridcol="2" gridrow="5" pageId="7" pageNumber="408">127.3</td>
<td id="76A95DE0FF8B22AAFC6A82CB32693FBB" box="[992,1024,433,457]" gridcol="3" gridrow="5" pageId="7" pageNumber="408">1.9</td>
<td id="76A95DE0FF8B22AAFBEE82CB328A3FBB" box="[1124,1251,433,457]" gridcol="4" gridrow="5" pageId="7" pageNumber="408">122.8–128.4</td>
<td id="76A95DE0FF8B22AAFB7382CB335B3FBB" box="[1273,1330,433,457]" gridcol="5" gridrow="5" pageId="7" pageNumber="408">124.8</td>
<td id="76A95DE0FF8B22AAFADA82CB33193FBB" box="[1360,1392,433,457]" gridcol="6" gridrow="5" pageId="7" pageNumber="408">2.2</td>
</tr>
<tr id="3578349CFF8B22AAFF1382A333193F83" box="[153,1392,473,497]" gridrow="6" pageId="7" pageNumber="408">
<th id="76A95DE0FF8B22AAFF1382A334BF3F83" box="[153,726,473,497]" gridcol="0" gridrow="6" pageId="7" pageNumber="408">Body depth at pelvic-fin origin</th>
<td id="76A95DE0FF8B22AAFD6C82A3350C3F83" box="[742,869,473,497]" gridcol="1" gridrow="6" pageId="7" pageNumber="408">27.4–31.0</td>
<td id="76A95DE0FF8B22AAFC0D82A335A93F83" box="[903,960,473,497]" gridcol="2" gridrow="6" pageId="7" pageNumber="408">29.0</td>
<td id="76A95DE0FF8B22AAFC6A82A332693F83" box="[992,1024,473,497]" gridcol="3" gridrow="6" pageId="7" pageNumber="408">1.1</td>
<td id="76A95DE0FF8B22AAFBEE82A3328A3F83" box="[1124,1251,473,497]" gridcol="4" gridrow="6" pageId="7" pageNumber="408">25.2–30.1</td>
<td id="76A95DE0FF8B22AAFB7382A3335B3F83" box="[1273,1330,473,497]" gridcol="5" gridrow="6" pageId="7" pageNumber="408">26.3</td>
<td id="76A95DE0FF8B22AAFADA82A333193F83" box="[1360,1392,473,497]" gridcol="6" gridrow="6" pageId="7" pageNumber="408">2.1</td>
</tr>
<tr id="3578349CFF8B22AAFF13817B33193C6B" box="[153,1392,513,537]" gridrow="7" pageId="7" pageNumber="408">
<th id="76A95DE0FF8B22AAFF13817B34BF3C6B" box="[153,726,513,537]" gridcol="0" gridrow="7" pageId="7" pageNumber="408">Head length</th>
<td id="76A95DE0FF8B22AAFD6C817B350C3C6B" box="[742,869,513,537]" gridcol="1" gridrow="7" pageId="7" pageNumber="408">25.6–30.6</td>
<td id="76A95DE0FF8B22AAFC0D817B35A93C6B" box="[903,960,513,537]" gridcol="2" gridrow="7" pageId="7" pageNumber="408">27.9</td>
<td id="76A95DE0FF8B22AAFC6A817B32693C6B" box="[992,1024,513,537]" gridcol="3" gridrow="7" pageId="7" pageNumber="408">1.7</td>
<td id="76A95DE0FF8B22AAFBEE817B328A3C6B" box="[1124,1251,513,537]" gridcol="4" gridrow="7" pageId="7" pageNumber="408">25.5–28.0</td>
<td id="76A95DE0FF8B22AAFB73817B335B3C6B" box="[1273,1330,513,537]" gridcol="5" gridrow="7" pageId="7" pageNumber="408">26.7</td>
<td id="76A95DE0FF8B22AAFADA817B33193C6B" box="[1360,1392,513,537]" gridcol="6" gridrow="7" pageId="7" pageNumber="408">1.0</td>
</tr>
<tr id="3578349CFF8B22AAFF13815333193C30" box="[153,1392,553,578]" gridrow="8" pageId="7" pageNumber="408">
<th id="76A95DE0FF8B22AAFF13815334BF3C30" box="[153,726,553,578]" gridcol="0" gridrow="8" pageId="7" pageNumber="408">Preanal length</th>
<td id="76A95DE0FF8B22AAFD6C8153350C3C30" box="[742,869,553,578]" gridcol="1" gridrow="8" pageId="7" pageNumber="408">55.5–59.9</td>
<td id="76A95DE0FF8B22AAFC0D815335A93C30" box="[903,960,553,578]" gridcol="2" gridrow="8" pageId="7" pageNumber="408">57.5</td>
<td id="76A95DE0FF8B22AAFC6A815332693C30" box="[992,1024,553,578]" gridcol="3" gridrow="8" pageId="7" pageNumber="408">1.4</td>
<td id="76A95DE0FF8B22AAFBEE8153328A3C30" box="[1124,1251,553,578]" gridcol="4" gridrow="8" pageId="7" pageNumber="408">60.5–65.2</td>
<td id="76A95DE0FF8B22AAFB738153335B3C30" box="[1273,1330,553,578]" gridcol="5" gridrow="8" pageId="7" pageNumber="408">62.7</td>
<td id="76A95DE0FF8B22AAFADA815333193C30" box="[1360,1392,553,578]" gridcol="6" gridrow="8" pageId="7" pageNumber="408">2.3</td>
</tr>
<tr id="3578349CFF8B22AAFF13812833193C18" box="[153,1392,594,618]" gridrow="9" pageId="7" pageNumber="408">
<th id="76A95DE0FF8B22AAFF13812834BF3C18" box="[153,726,594,618]" gridcol="0" gridrow="9" pageId="7" pageNumber="408">Predorsal length</th>
<td id="76A95DE0FF8B22AAFD6C8128350C3C18" box="[742,869,594,618]" gridcol="1" gridrow="9" pageId="7" pageNumber="408">63.3–67.9</td>
<td id="76A95DE0FF8B22AAFC0D812835A93C18" box="[903,960,594,618]" gridcol="2" gridrow="9" pageId="7" pageNumber="408">65.1</td>
<td id="76A95DE0FF8B22AAFC6A812832693C18" box="[992,1024,594,618]" gridcol="3" gridrow="9" pageId="7" pageNumber="408">1.5</td>
<td id="76A95DE0FF8B22AAFBEE8128328A3C18" box="[1124,1251,594,618]" gridcol="4" gridrow="9" pageId="7" pageNumber="408">63.6–69.2</td>
<td id="76A95DE0FF8B22AAFB738128335B3C18" box="[1273,1330,594,618]" gridcol="5" gridrow="9" pageId="7" pageNumber="408">67.1</td>
<td id="76A95DE0FF8B22AAFADA812833193C18" box="[1360,1392,594,618]" gridcol="6" gridrow="9" pageId="7" pageNumber="408">2.3</td>
</tr>
<tr id="3578349CFF8B22AAFF13810033193CE0" box="[153,1392,634,658]" gridrow="10" pageId="7" pageNumber="408">
<th id="76A95DE0FF8B22AAFF13810034BF3CE0" box="[153,726,634,658]" gridcol="0" gridrow="10" pageId="7" pageNumber="408">Prepelvic length</th>
<td id="76A95DE0FF8B22AAFD6C8100350C3CE0" box="[742,869,634,658]" gridcol="1" gridrow="10" pageId="7" pageNumber="408">41.1–45.5</td>
<td id="76A95DE0FF8B22AAFC0D810035A93CE0" box="[903,960,634,658]" gridcol="2" gridrow="10" pageId="7" pageNumber="408">43.2</td>
<td id="76A95DE0FF8B22AAFC6A810032693CE0" box="[992,1024,634,658]" gridcol="3" gridrow="10" pageId="7" pageNumber="408">1.2</td>
<td id="76A95DE0FF8B22AAFBEE8100328A3CE0" box="[1124,1251,634,658]" gridcol="4" gridrow="10" pageId="7" pageNumber="408">44.4–48.4</td>
<td id="76A95DE0FF8B22AAFB738100335B3CE0" box="[1273,1330,634,658]" gridcol="5" gridrow="10" pageId="7" pageNumber="408">46.5</td>
<td id="76A95DE0FF8B22AAFADA810033193CE0" box="[1360,1392,634,658]" gridcol="6" gridrow="10" pageId="7" pageNumber="408">1.5</td>
</tr>
<tr id="3578349CFF8B22AAFF1381D833193CC8" box="[153,1392,674,698]" gridrow="11" pageId="7" pageNumber="408">
<th id="76A95DE0FF8B22AAFF1381D834BF3CC8" box="[153,726,674,698]" gridcol="0" gridrow="11" pageId="7" pageNumber="408">Prepectoral length</th>
<td id="76A95DE0FF8B22AAFD6C81D8350C3CC8" box="[742,869,674,698]" gridcol="1" gridrow="11" pageId="7" pageNumber="408">26.7–32.4</td>
<td id="76A95DE0FF8B22AAFC0D81D835A93CC8" box="[903,960,674,698]" gridcol="2" gridrow="11" pageId="7" pageNumber="408">29.1</td>
<td id="76A95DE0FF8B22AAFC6A81D832693CC8" box="[992,1024,674,698]" gridcol="3" gridrow="11" pageId="7" pageNumber="408">1.9</td>
<td id="76A95DE0FF8B22AAFBEE81D8328A3CC8" box="[1124,1251,674,698]" gridcol="4" gridrow="11" pageId="7" pageNumber="408">26.2–28.4</td>
<td id="76A95DE0FF8B22AAFB7381D8335B3CC8" box="[1273,1330,674,698]" gridcol="5" gridrow="11" pageId="7" pageNumber="408">27.2</td>
<td id="76A95DE0FF8B22AAFADA81D833193CC8" box="[1360,1392,674,698]" gridcol="6" gridrow="11" pageId="7" pageNumber="408">1.0</td>
</tr>
<tr id="3578349CFF8B22AAFF1381B033193C91" box="[153,1392,714,739]" gridrow="12" pageId="7" pageNumber="408">
<th id="76A95DE0FF8B22AAFF1381B034BF3C91" box="[153,726,714,739]" gridcol="0" gridrow="12" pageId="7" pageNumber="408">Caudal peduncle length</th>
<td id="76A95DE0FF8B22AAFD6C81B0350C3C91" box="[742,869,714,739]" gridcol="1" gridrow="12" pageId="7" pageNumber="408">17.5–20.8</td>
<td id="76A95DE0FF8B22AAFC0D81B035A93C91" box="[903,960,714,739]" gridcol="2" gridrow="12" pageId="7" pageNumber="408">18.9</td>
<td id="76A95DE0FF8B22AAFC6A81B032693C91" box="[992,1024,714,739]" gridcol="3" gridrow="12" pageId="7" pageNumber="408">1.0</td>
<td id="76A95DE0FF8B22AAFBEE81B0328A3C91" box="[1124,1251,714,739]" gridcol="4" gridrow="12" pageId="7" pageNumber="408">20.1–22.3</td>
<td id="76A95DE0FF8B22AAFB7381B0335B3C91" box="[1273,1330,714,739]" gridcol="5" gridrow="12" pageId="7" pageNumber="408">21.1</td>
<td id="76A95DE0FF8B22AAFADA81B033193C91" box="[1360,1392,714,739]" gridcol="6" gridrow="12" pageId="7" pageNumber="408">0.9</td>
</tr>
<tr id="3578349CFF8B22AAFF13818933193D79" box="[153,1392,755,779]" gridrow="13" pageId="7" pageNumber="408">
<th id="76A95DE0FF8B22AAFF13818934BF3D79" box="[153,726,755,779]" gridcol="0" gridrow="13" pageId="7" pageNumber="408">Caudal peduncle depth</th>
<td id="76A95DE0FF8B22AAFD6C8189350C3D79" box="[742,869,755,779]" gridcol="1" gridrow="13" pageId="7" pageNumber="408">12.7–15.0</td>
<td id="76A95DE0FF8B22AAFC0D818935A93D79" box="[903,960,755,779]" gridcol="2" gridrow="13" pageId="7" pageNumber="408">13.8</td>
<td id="76A95DE0FF8B22AAFC6A818932693D79" box="[992,1024,755,779]" gridcol="3" gridrow="13" pageId="7" pageNumber="408">0.7</td>
<td id="76A95DE0FF8B22AAFBEE8189328A3D79" box="[1124,1251,755,779]" gridcol="4" gridrow="13" pageId="7" pageNumber="408">10.7–11.8</td>
<td id="76A95DE0FF8B22AAFB738189335B3D79" box="[1273,1330,755,779]" gridcol="5" gridrow="13" pageId="7" pageNumber="408">11.2</td>
<td id="76A95DE0FF8B22AAFADA818933193D79" box="[1360,1392,755,779]" gridcol="6" gridrow="13" pageId="7" pageNumber="408">0.4</td>
</tr>
<tr id="3578349CFF8B22AAFF13806133193D41" box="[153,1392,795,819]" gridrow="14" pageId="7" pageNumber="408">
<th id="76A95DE0FF8B22AAFF13806134BF3D41" box="[153,726,795,819]" gridcol="0" gridrow="14" pageId="7" pageNumber="408">Dorsal-fin base length</th>
<td id="76A95DE0FF8B22AAFD6C8061350C3D41" box="[742,869,795,819]" gridcol="1" gridrow="14" pageId="7" pageNumber="408">18.8–25.1</td>
<td id="76A95DE0FF8B22AAFC0D806135A93D41" box="[903,960,795,819]" gridcol="2" gridrow="14" pageId="7" pageNumber="408">21.8</td>
<td id="76A95DE0FF8B22AAFC6A806132693D41" box="[992,1024,795,819]" gridcol="3" gridrow="14" pageId="7" pageNumber="408">2.2</td>
<td id="76A95DE0FF8B22AAFBEE8061328A3D41" box="[1124,1251,795,819]" gridcol="4" gridrow="14" pageId="7" pageNumber="408">17.0–19.0</td>
<td id="76A95DE0FF8B22AAFB738061335B3D41" box="[1273,1330,795,819]" gridcol="5" gridrow="14" pageId="7" pageNumber="408">17.7</td>
<td id="76A95DE0FF8B22AAFADA806133193D41" box="[1360,1392,795,819]" gridcol="6" gridrow="14" pageId="7" pageNumber="408">0.7</td>
</tr>
<tr id="3578349CFF8B22AAFF13803933193D29" box="[153,1392,835,859]" gridrow="15" pageId="7" pageNumber="408">
<th id="76A95DE0FF8B22AAFF13803934BF3D29" box="[153,726,835,859]" gridcol="0" gridrow="15" pageId="7" pageNumber="408">Anal-fin base length</th>
<td id="76A95DE0FF8B22AAFD6C8039350C3D29" box="[742,869,835,859]" gridcol="1" gridrow="15" pageId="7" pageNumber="408">22.8–27.1</td>
<td id="76A95DE0FF8B22AAFC0D803935A93D29" box="[903,960,835,859]" gridcol="2" gridrow="15" pageId="7" pageNumber="408">25.0</td>
<td id="76A95DE0FF8B22AAFC6A803932693D29" box="[992,1024,835,859]" gridcol="3" gridrow="15" pageId="7" pageNumber="408">1.4</td>
<td id="76A95DE0FF8B22AAFBEE8039328A3D29" box="[1124,1251,835,859]" gridcol="4" gridrow="15" pageId="7" pageNumber="408">18.7–20.4</td>
<td id="76A95DE0FF8B22AAFB738039335B3D29" box="[1273,1330,835,859]" gridcol="5" gridrow="15" pageId="7" pageNumber="408">19.4</td>
<td id="76A95DE0FF8B22AAFADA803933193D29" box="[1360,1392,835,859]" gridcol="6" gridrow="15" pageId="7" pageNumber="408">0.7</td>
</tr>
<tr id="3578349CFF8B22AAFF13801133193DF6" box="[153,1392,875,900]" gridrow="16" pageId="7" pageNumber="408">
<th id="76A95DE0FF8B22AAFF13801134BF3DF6" box="[153,726,875,900]" gridcol="0" gridrow="16" pageId="7" pageNumber="408">Caudal-fin length</th>
<td id="76A95DE0FF8B22AAFD6C8011350C3DF6" box="[742,869,875,900]" gridcol="1" gridrow="16" pageId="7" pageNumber="408">23.9–30.6</td>
<td id="76A95DE0FF8B22AAFC0D801135A93DF6" box="[903,960,875,900]" gridcol="2" gridrow="16" pageId="7" pageNumber="408">27.2</td>
<td id="76A95DE0FF8B22AAFC6A801132693DF6" box="[992,1024,875,900]" gridcol="3" gridrow="16" pageId="7" pageNumber="408">2.1</td>
<td id="76A95DE0FF8B22AAFBEE8011328A3DF6" box="[1124,1251,875,900]" gridcol="4" gridrow="16" pageId="7" pageNumber="408">22.8–28.4</td>
<td id="76A95DE0FF8B22AAFB738011335B3DF6" box="[1273,1330,875,900]" gridcol="5" gridrow="16" pageId="7" pageNumber="408">24.8</td>
<td id="76A95DE0FF8B22AAFADA801133193DF6" box="[1360,1392,875,900]" gridcol="6" gridrow="16" pageId="7" pageNumber="408">2.2</td>
</tr>
<tr id="3578349CFF8B22AAFF1380E933193DDE" box="[153,1392,915,940]" gridrow="17" pageId="7" pageNumber="408">
<th id="76A95DE0FF8B22AAFF1380E933193DDE" box="[153,1392,915,940]" colspan="7" colspanRight="6" gridcol="0" gridrow="17" pageId="7" pageNumber="408">
<emphasis id="B93CEACCFF8BDD52FF1380EE37E23DDE" bold="true" box="[153,395,915,940]" pageId="7" pageNumber="408">Percent of head length</emphasis>
</th>
</tr>
<tr id="3578349CFF8B22AAFF1380C633193DA6" box="[153,1392,956,980]" gridrow="18" pageId="7" pageNumber="408">
<th id="76A95DE0FF8B22AAFF1380C634BF3DA6" box="[153,726,956,980]" gridcol="0" gridrow="18" pageId="7" pageNumber="408">Head width</th>
<td id="76A95DE0FF8B22AAFD6C80C6350C3DA6" box="[742,869,956,980]" gridcol="1" gridrow="18" pageId="7" pageNumber="408">58–65</td>
<td id="76A95DE0FF8B22AAFC0D80C635A93DA6" box="[903,960,956,980]" gridcol="2" gridrow="18" pageId="7" pageNumber="408">60.1</td>
<td id="76A95DE0FF8B22AAFC6A80C632693DA6" box="[992,1024,956,980]" gridcol="3" gridrow="18" pageId="7" pageNumber="408">2.2</td>
<td id="76A95DE0FF8B22AAFBEE80C6328A3DA6" box="[1124,1251,956,980]" gridcol="4" gridrow="18" pageId="7" pageNumber="408">57–63</td>
<td id="76A95DE0FF8B22AAFB7380C6335B3DA6" box="[1273,1330,956,980]" gridcol="5" gridrow="18" pageId="7" pageNumber="408">59.7</td>
<td id="76A95DE0FF8B22AAFADA80C633193DA6" box="[1360,1392,956,980]" gridcol="6" gridrow="18" pageId="7" pageNumber="408">2.0</td>
</tr>
<tr id="3578349CFF8B22AAFF13809E33193D8E" box="[153,1392,996,1020]" gridrow="19" pageId="7" pageNumber="408">
<th id="76A95DE0FF8B22AAFF13809E34BF3D8E" box="[153,726,996,1020]" gridcol="0" gridrow="19" pageId="7" pageNumber="408">Head depth</th>
<td id="76A95DE0FF8B22AAFD6C809E350C3D8E" box="[742,869,996,1020]" gridcol="1" gridrow="19" pageId="7" pageNumber="408">74–81</td>
<td id="76A95DE0FF8B22AAFC0D809E35A93D8E" box="[903,960,996,1020]" gridcol="2" gridrow="19" pageId="7" pageNumber="408">77.4</td>
<td id="76A95DE0FF8B22AAFC6A809E32693D8E" box="[992,1024,996,1020]" gridcol="3" gridrow="19" pageId="7" pageNumber="408">2.4</td>
<td id="76A95DE0FF8B22AAFBEE809E328A3D8E" box="[1124,1251,996,1020]" gridcol="4" gridrow="19" pageId="7" pageNumber="408">67–69</td>
<td id="76A95DE0FF8B22AAFB73809E335B3D8E" box="[1273,1330,996,1020]" gridcol="5" gridrow="19" pageId="7" pageNumber="408">68.1</td>
<td id="76A95DE0FF8B22AAFADA809E33193D8E" box="[1360,1392,996,1020]" gridcol="6" gridrow="19" pageId="7" pageNumber="408">1.3</td>
</tr>
<tr id="3578349CFF8B22AAFF13877633193A57" box="[153,1392,1036,1061]" gridrow="20" pageId="7" pageNumber="408">
<th id="76A95DE0FF8B22AAFF13877634BF3A57" box="[153,726,1036,1061]" gridcol="0" gridrow="20" pageId="7" pageNumber="408">Interorbital width</th>
<td id="76A95DE0FF8B22AAFD6C8776350C3A57" box="[742,869,1036,1061]" gridcol="1" gridrow="20" pageId="7" pageNumber="408">46–55</td>
<td id="76A95DE0FF8B22AAFC0D877635A93A57" box="[903,960,1036,1061]" gridcol="2" gridrow="20" pageId="7" pageNumber="408">49.9</td>
<td id="76A95DE0FF8B22AAFC6A877632693A57" box="[992,1024,1036,1061]" gridcol="3" gridrow="20" pageId="7" pageNumber="408">2.7</td>
<td id="76A95DE0FF8B22AAFBEE8776328A3A57" box="[1124,1251,1036,1061]" gridcol="4" gridrow="20" pageId="7" pageNumber="408">38–42</td>
<td id="76A95DE0FF8B22AAFB738776335B3A57" box="[1273,1330,1036,1061]" gridcol="5" gridrow="20" pageId="7" pageNumber="408">40.7</td>
<td id="76A95DE0FF8B22AAFADA877633193A57" box="[1360,1392,1036,1061]" gridcol="6" gridrow="20" pageId="7" pageNumber="408">1.7</td>
</tr>
<tr id="3578349CFF8B22AAFF13874E33193A3F" box="[153,1392,1076,1101]" gridrow="21" pageId="7" pageNumber="408">
<th id="76A95DE0FF8B22AAFF13874E34BF3A3F" box="[153,726,1076,1101]" gridcol="0" gridrow="21" pageId="7" pageNumber="408">Postorbital length</th>
<td id="76A95DE0FF8B22AAFD6C874E350C3A3F" box="[742,869,1076,1101]" gridcol="1" gridrow="21" pageId="7" pageNumber="408">41–49</td>
<td id="76A95DE0FF8B22AAFC0D874E35A93A3F" box="[903,960,1076,1101]" gridcol="2" gridrow="21" pageId="7" pageNumber="408">45.9</td>
<td id="76A95DE0FF8B22AAFC6A874E32693A3F" box="[992,1024,1076,1101]" gridcol="3" gridrow="21" pageId="7" pageNumber="408">2.8</td>
<td id="76A95DE0FF8B22AAFBEE874E328A3A3F" box="[1124,1251,1076,1101]" gridcol="4" gridrow="21" pageId="7" pageNumber="408">47–53</td>
<td id="76A95DE0FF8B22AAFB73874E335B3A3F" box="[1273,1330,1076,1101]" gridcol="5" gridrow="21" pageId="7" pageNumber="408">49.4</td>
<td id="76A95DE0FF8B22AAFADA874E33193A3F" box="[1360,1392,1076,1101]" gridcol="6" gridrow="21" pageId="7" pageNumber="408">2.3</td>
</tr>
<tr id="3578349CFF8B22AAFF13872733193A07" box="[153,1392,1117,1141]" gridrow="22" pageId="7" pageNumber="408">
<th id="76A95DE0FF8B22AAFF13872734BF3A07" box="[153,726,1117,1141]" gridcol="0" gridrow="22" pageId="7" pageNumber="408">Suborbital depth</th>
<td id="76A95DE0FF8B22AAFD6C8727350C3A07" box="[742,869,1117,1141]" gridcol="1" gridrow="22" pageId="7" pageNumber="408">16–20</td>
<td id="76A95DE0FF8B22AAFC0D872735A93A07" box="[903,960,1117,1141]" gridcol="2" gridrow="22" pageId="7" pageNumber="408">17.4</td>
<td id="76A95DE0FF8B22AAFC6A872732693A07" box="[992,1024,1117,1141]" gridcol="3" gridrow="22" pageId="7" pageNumber="408">1.8</td>
<td id="76A95DE0FF8B22AAFBEE8727328A3A07" box="[1124,1251,1117,1141]" gridcol="4" gridrow="22" pageId="7" pageNumber="408">11–14</td>
<td id="76A95DE0FF8B22AAFB738727335B3A07" box="[1273,1330,1117,1141]" gridcol="5" gridrow="22" pageId="7" pageNumber="408">12.6</td>
<td id="76A95DE0FF8B22AAFADA872733193A07" box="[1360,1392,1117,1141]" gridcol="6" gridrow="22" pageId="7" pageNumber="408">1.1</td>
</tr>
<tr id="3578349CFF8B22AAFF1387FF33193AEC" box="[153,1392,1157,1182]" gridrow="23" pageId="7" pageNumber="408">
<th id="76A95DE0FF8B22AAFF1387FF34BF3AEC" box="[153,726,1157,1182]" gridcol="0" gridrow="23" pageId="7" pageNumber="408">Eye diameter</th>
<td id="76A95DE0FF8B22AAFD6C87FF350C3AEC" box="[742,869,1157,1182]" gridcol="1" gridrow="23" pageId="7" pageNumber="408">34–38</td>
<td id="76A95DE0FF8B22AAFC0D87FF35A93AEC" box="[903,960,1157,1182]" gridcol="2" gridrow="23" pageId="7" pageNumber="408">35.6</td>
<td id="76A95DE0FF8B22AAFC6A87FF32693AEC" box="[992,1024,1157,1182]" gridcol="3" gridrow="23" pageId="7" pageNumber="408">1.5</td>
<td id="76A95DE0FF8B22AAFBEE87FF328A3AEC" box="[1124,1251,1157,1182]" gridcol="4" gridrow="23" pageId="7" pageNumber="408">33–38</td>
<td id="76A95DE0FF8B22AAFB7387FF335B3AEC" box="[1273,1330,1157,1182]" gridcol="5" gridrow="23" pageId="7" pageNumber="408">35.8</td>
<td id="76A95DE0FF8B22AAFADA87FF33193AEC" box="[1360,1392,1157,1182]" gridcol="6" gridrow="23" pageId="7" pageNumber="408">1.8</td>
</tr>
<tr id="3578349CFF8B22AAFF1387D733193AB4" box="[153,1392,1197,1222]" gridrow="24" pageId="7" pageNumber="408">
<th id="76A95DE0FF8B22AAFF1387D734BF3AB4" box="[153,726,1197,1222]" gridcol="0" gridrow="24" pageId="7" pageNumber="408">Snout to eye end length</th>
<td id="76A95DE0FF8B22AAFD6C87D7350C3AB4" box="[742,869,1197,1222]" gridcol="1" gridrow="24" pageId="7" pageNumber="408">51–59</td>
<td id="76A95DE0FF8B22AAFC0D87D735A93AB4" box="[903,960,1197,1222]" gridcol="2" gridrow="24" pageId="7" pageNumber="408">54.1</td>
<td id="76A95DE0FF8B22AAFC6A87D732693AB4" box="[992,1024,1197,1222]" gridcol="3" gridrow="24" pageId="7" pageNumber="408">2.8</td>
<td id="76A95DE0FF8B22AAFBEE87D7328A3AB4" box="[1124,1251,1197,1222]" gridcol="4" gridrow="24" pageId="7" pageNumber="408">48–53</td>
<td id="76A95DE0FF8B22AAFB7387D7335B3AB4" box="[1273,1330,1197,1222]" gridcol="5" gridrow="24" pageId="7" pageNumber="408">50.6</td>
<td id="76A95DE0FF8B22AAFADA87D733193AB4" box="[1360,1392,1197,1222]" gridcol="6" gridrow="24" pageId="7" pageNumber="408">2.3</td>
</tr>
<tr id="3578349CFF8B22AAFF1387AF33193A9C" box="[153,1392,1237,1262]" gridrow="25" pageId="7" pageNumber="408">
<th id="76A95DE0FF8B22AAFF1387AF34BF3A9C" box="[153,726,1237,1262]" gridcol="0" gridrow="25" pageId="7" pageNumber="408">Snout length</th>
<td id="76A95DE0FF8B22AAFD6C87AF350C3A9C" box="[742,869,1237,1262]" gridcol="1" gridrow="25" pageId="7" pageNumber="408">21–30</td>
<td id="76A95DE0FF8B22AAFC0D87AF35A93A9C" box="[903,960,1237,1262]" gridcol="2" gridrow="25" pageId="7" pageNumber="408">23.1</td>
<td id="76A95DE0FF8B22AAFC6A87AF32693A9C" box="[992,1024,1237,1262]" gridcol="3" gridrow="25" pageId="7" pageNumber="408">2.8</td>
<td id="76A95DE0FF8B22AAFBEE87AF328A3A9C" box="[1124,1251,1237,1262]" gridcol="4" gridrow="25" pageId="7" pageNumber="408">24–28</td>
<td id="76A95DE0FF8B22AAFB7387AF335B3A9C" box="[1273,1330,1237,1262]" gridcol="5" gridrow="25" pageId="7" pageNumber="408">25.9</td>
<td id="76A95DE0FF8B22AAFADA87AF33193A9C" box="[1360,1392,1237,1262]" gridcol="6" gridrow="25" pageId="7" pageNumber="408">1.4</td>
</tr>
<tr id="3578349CFF8B22AAFF13878733193B67" box="[153,1392,1277,1301]" gridrow="26" pageId="7" pageNumber="408">
<th id="76A95DE0FF8B22AAFF13878733193B67" box="[153,1392,1277,1301]" colspan="7" colspanRight="6" gridcol="0" gridrow="26" pageId="7" pageNumber="408">
<emphasis id="B93CEACCFF8BDD52FF13878737A93B67" bold="true" box="[153,448,1277,1301]" pageId="7" pageNumber="408">Other morphometric ratios</emphasis>
</th>
</tr>
<tr id="3578349CFF8B22AAFF13865C33193B4C" box="[153,1392,1318,1342]" gridrow="27" pageId="7" pageNumber="408">
<th id="76A95DE0FF8B22AAFF13865C34BF3B4C" box="[153,726,1318,1342]" gridcol="0" gridrow="27" pageId="7" pageNumber="408">Predorsal length in % of preanal length</th>
<td id="76A95DE0FF8B22AAFD6C865C350C3B4C" box="[742,869,1318,1342]" gridcol="1" gridrow="27" pageId="7" pageNumber="408">109–115</td>
<td id="76A95DE0FF8B22AAFC0D865C35A93B4C" box="[903,960,1318,1342]" gridcol="2" gridrow="27" pageId="7" pageNumber="408">113.2</td>
<td id="76A95DE0FF8B22AAFC6A865C32693B4C" box="[992,1024,1318,1342]" gridcol="3" gridrow="27" pageId="7" pageNumber="408">1.9</td>
<td id="76A95DE0FF8B22AAFBEE865C328A3B4C" box="[1124,1251,1318,1342]" gridcol="4" gridrow="27" pageId="7" pageNumber="408">105–109</td>
<td id="76A95DE0FF8B22AAFB73865C335B3B4C" box="[1273,1330,1318,1342]" gridcol="5" gridrow="27" pageId="7" pageNumber="408">107.2</td>
<td id="76A95DE0FF8B22AAFADA865C33193B4C" box="[1360,1392,1318,1342]" gridcol="6" gridrow="27" pageId="7" pageNumber="408">1.9</td>
</tr>
<tr id="3578349CFF8B22AAFF13863433193B14" box="[153,1392,1358,1382]" gridrow="28" pageId="7" pageNumber="408">
<th id="76A95DE0FF8B22AAFF13863434BF3B14" box="[153,726,1358,1382]" gridcol="0" gridrow="28" pageId="7" pageNumber="408">Caudal peduncle length in % of its depth</th>
<td id="76A95DE0FF8B22AAFD6C8634350C3B14" box="[742,869,1358,1382]" gridcol="1" gridrow="28" pageId="7" pageNumber="408">133–140</td>
<td id="76A95DE0FF8B22AAFC0D863435A93B14" box="[903,960,1358,1382]" gridcol="2" gridrow="28" pageId="7" pageNumber="408">137.4</td>
<td id="76A95DE0FF8B22AAFC6A863432693B14" box="[992,1024,1358,1382]" gridcol="3" gridrow="28" pageId="7" pageNumber="408">2.7</td>
<td id="76A95DE0FF8B22AAFBEE8634328A3B14" box="[1124,1251,1358,1382]" gridcol="4" gridrow="28" pageId="7" pageNumber="408">184–190</td>
<td id="76A95DE0FF8B22AAFB738634335B3B14" box="[1273,1330,1358,1382]" gridcol="5" gridrow="28" pageId="7" pageNumber="408">187.7</td>
<td id="76A95DE0FF8B22AAFADA863433193B14" box="[1360,1392,1358,1382]" gridcol="6" gridrow="28" pageId="7" pageNumber="408">2.2</td>
</tr>
<tr id="3578349CFF8B22AAFF13860C33193BFD" box="[153,1392,1398,1423]" gridrow="29" pageId="7" pageNumber="408">
<th id="76A95DE0FF8B22AAFF13860C34BF3BFD" box="[153,726,1398,1423]" gridcol="0" gridrow="29" pageId="7" pageNumber="408">Head width in % of its depth</th>
<td id="76A95DE0FF8B22AAFD6C860C350C3BFD" box="[742,869,1398,1423]" gridcol="1" gridrow="29" pageId="7" pageNumber="408">74–81</td>
<td id="76A95DE0FF8B22AAFC0D860C35A93BFD" box="[903,960,1398,1423]" gridcol="2" gridrow="29" pageId="7" pageNumber="408">77.6</td>
<td id="76A95DE0FF8B22AAFC6A860C32693BFD" box="[992,1024,1398,1423]" gridcol="3" gridrow="29" pageId="7" pageNumber="408">2.3</td>
<td id="76A95DE0FF8B22AAFBEE860C328A3BFD" box="[1124,1251,1398,1423]" gridcol="4" gridrow="29" pageId="7" pageNumber="408">85–90</td>
<td id="76A95DE0FF8B22AAFB73860C335B3BFD" box="[1273,1330,1398,1423]" gridcol="5" gridrow="29" pageId="7" pageNumber="408">87.7</td>
<td id="76A95DE0FF8B22AAFADA860C33193BFD" box="[1360,1392,1398,1423]" gridcol="6" gridrow="29" pageId="7" pageNumber="408">2.3</td>
</tr>
<tr id="3578349CFF8B22AAFF1386E433193BC5" box="[153,1392,1438,1463]" gridrow="30" pageId="7" pageNumber="408" rowspan-3="1" rowspan-6="1">
<th id="76A95DE0FF8B22AAFF1386E434BF3BC5" box="[153,726,1438,1463]" gridcol="0" gridrow="30" pageId="7" pageNumber="408">
<emphasis id="B93CEACCFF8BDD52FF1386E436943BC4" bold="true" box="[153,253,1438,1462]" pageId="7" pageNumber="408">Meristics</emphasis>
</th>
<td id="76A95DE0FF8B22AAFD6C86E4350C3BC5" box="[742,869,1438,1463]" gridcol="1" gridrow="30" pageId="7" pageNumber="408">range</td>
<td id="76A95DE0FF8B22AAFC0D86E435A93BC5" box="[903,960,1438,1463]" gridcol="2" gridrow="30" pageId="7" pageNumber="408">mode</td>
<td id="76A95DE0FF8B22AAFBEE86E4328A3BC5" box="[1124,1251,1438,1463]" gridcol="4" gridrow="30" pageId="7" pageNumber="408">range</td>
<td id="76A95DE0FF8B22AAFB7386E4335B3BC5" box="[1273,1330,1438,1463]" gridcol="5" gridrow="30" pageId="7" pageNumber="408">mode</td>
</tr>
<tr id="3578349CFF8B22AAFF1386BD33193B92" box="[153,1392,1479,1504]" gridrow="31" pageId="7" pageNumber="408" rowspan-3="1" rowspan-6="1">
<th id="76A95DE0FF8B22AAFF1386BD34BF3B92" box="[153,726,1479,1504]" gridcol="0" gridrow="31" pageId="7" pageNumber="408">Dorsal-fin rays</th>
<td id="76A95DE0FF8B22AAFD6C86BD350C3B92" box="[742,869,1479,1504]" gridcol="1" gridrow="31" pageId="7" pageNumber="408">12–14</td>
<td id="76A95DE0FF8B22AAFC0D86BD35A93B92" box="[903,960,1479,1504]" gridcol="2" gridrow="31" pageId="7" pageNumber="408">13</td>
<td id="76A95DE0FF8B22AAFBEE86BD328A3B92" box="[1124,1251,1479,1504]" gridcol="4" gridrow="31" pageId="7" pageNumber="408">12–14</td>
<td id="76A95DE0FF8B22AAFB7386BD335B3B92" box="[1273,1330,1479,1504]" gridcol="5" gridrow="31" pageId="7" pageNumber="408">12</td>
</tr>
<tr id="3578349CFF8B22AAFF1386953319387A" box="[153,1392,1519,1544]" gridrow="32" pageId="7" pageNumber="408" rowspan-3="1" rowspan-6="1">
<th id="76A95DE0FF8B22AAFF13869534BF387A" box="[153,726,1519,1544]" gridcol="0" gridrow="32" pageId="7" pageNumber="408">Anal-fin rays</th>
<td id="76A95DE0FF8B22AAFD6C8695350C387A" box="[742,869,1519,1544]" gridcol="1" gridrow="32" pageId="7" pageNumber="408">16–19</td>
<td id="76A95DE0FF8B22AAFC0D869535A9387A" box="[903,960,1519,1544]" gridcol="2" gridrow="32" pageId="7" pageNumber="408">18</td>
<td id="76A95DE0FF8B22AAFBEE8695328A387A" box="[1124,1251,1519,1544]" gridcol="4" gridrow="32" pageId="7" pageNumber="408">17–18</td>
<td id="76A95DE0FF8B22AAFB738695335B387A" box="[1273,1330,1519,1544]" gridcol="5" gridrow="32" pageId="7" pageNumber="408">17</td>
</tr>
<tr id="3578349CFF8B22AAFF13856D33193842" box="[153,1392,1559,1584]" gridrow="33" pageId="7" pageNumber="408" rowspan-3="1" rowspan-6="1">
<th id="76A95DE0FF8B22AAFF13856D34BF3842" box="[153,726,1559,1584]" gridcol="0" gridrow="33" pageId="7" pageNumber="408">Scales mid-longitudinal series</th>
<td id="76A95DE0FF8B22AAFD6C856D350C3842" box="[742,869,1559,1584]" gridcol="1" gridrow="33" pageId="7" pageNumber="408">26–28</td>
<td id="76A95DE0FF8B22AAFC0D856D35A93842" box="[903,960,1559,1584]" gridcol="2" gridrow="33" pageId="7" pageNumber="408">28</td>
<td id="76A95DE0FF8B22AAFBEE856D328A3842" box="[1124,1251,1559,1584]" gridcol="4" gridrow="33" pageId="7" pageNumber="408">26–28</td>
<td id="76A95DE0FF8B22AAFB73856D335B3842" box="[1273,1330,1559,1584]" gridcol="5" gridrow="33" pageId="7" pageNumber="408">27</td>
</tr>
<tr id="3578349CFF8B22AAFF13853A3319382A" box="[153,1392,1600,1624]" gridrow="34" pageId="7" pageNumber="408" rowspan-3="1" rowspan-6="1">
<th id="76A95DE0FF8B22AAFF13853A34BF382A" box="[153,726,1600,1624]" gridcol="0" gridrow="34" pageId="7" pageNumber="408">Scales transverse</th>
<td id="76A95DE0FF8B22AAFD6C853A350C382A" box="[742,869,1600,1624]" gridcol="1" gridrow="34" pageId="7" pageNumber="408">11</td>
<td id="76A95DE0FF8B22AAFC0D853A35A9382A" box="[903,960,1600,1624]" gridcol="2" gridrow="34" pageId="7" pageNumber="408">11</td>
<td id="76A95DE0FF8B22AAFBEE853A328A382A" box="[1124,1251,1600,1624]" gridcol="4" gridrow="34" pageId="7" pageNumber="408">11</td>
<td id="76A95DE0FF8B22AAFB73853A335B382A" box="[1273,1330,1600,1624]" gridcol="5" gridrow="34" pageId="7" pageNumber="408">11</td>
</tr>
<tr id="3578349CFF8B22AAFF138512331938F2" box="[153,1392,1640,1664]" gridrow="35" pageId="7" pageNumber="408" rowspan-3="1" rowspan-6="1">
<th id="76A95DE0FF8B22AAFF13851234BF38F2" box="[153,726,1640,1664]" gridcol="0" gridrow="35" pageId="7" pageNumber="408">Scales circumpeduncular</th>
<td id="76A95DE0FF8B22AAFD6C8512350C38F2" box="[742,869,1640,1664]" gridcol="1" gridrow="35" pageId="7" pageNumber="408">12</td>
<td id="76A95DE0FF8B22AAFC0D851235A938F2" box="[903,960,1640,1664]" gridcol="2" gridrow="35" pageId="7" pageNumber="408">12</td>
<td id="76A95DE0FF8B22AAFBEE8512328A38F2" box="[1124,1251,1640,1664]" gridcol="4" gridrow="35" pageId="7" pageNumber="408">12</td>
<td id="76A95DE0FF8B22AAFB738512335B38F2" box="[1273,1330,1640,1664]" gridcol="5" gridrow="35" pageId="7" pageNumber="408">12</td>
</tr>
</table>
</paragraph>
<paragraph id="8BF736DEFF8BDD52FF4D85D532B5390F" blockId="7.[151,1437,1711,2026]" pageId="7" pageNumber="408">
Male: Small
<taxonomicName id="4C484D5DFF8BDD52FED585D5341A38BB" box="[351,627,1711,1737]" pageId="7" pageNumber="408">
<emphasis id="B93CEACCFF8BDD52FED585D5347A38BB" box="[351,531,1711,1737]" italics="true" pageId="7" pageNumber="408">Nothobranchius</emphasis>
species
</taxonomicName>
, maximum examined size
<quantity id="4CB09B3BFF8BDD52FC2185CA327A38B8" box="[939,1043,1712,1738]" metricMagnitude="-2" metricUnit="m" metricValue="3.37" pageId="7" pageNumber="408" unit="mm" value="33.7">33.7 mm</quantity>
SL. General body shape laterally compressed. Greatest body depth at vertical in front of pelvic-fin origin (27.4–31.0 % SL). Greatest body width at pectoral-fin base with body progressively narrowing towards caudal-fin base. Dorsal profile slightly convex to straight from tip of snout to dorsal-fin origin, straight to slightly concave on caudal peduncle. Ventral profile convex from lower jaw to base of last anal-fin ray, straight to slightly concave on caudal peduncle. Caudal peduncle moderately deep, length 133–140 % of its depth. Anus situated directly in front of anal-fin origin.
</paragraph>
<paragraph id="8BF736DEFF8BDD5DFF4D84FD35F93E8A" blockId="7.[151,1437,1711,2026]" lastBlockId="8.[151,1437,151,2035]" lastPageId="8" lastPageNumber="409" pageId="7" pageNumber="408">Head moderately long (25.6–30.6 % SL), laterally compressed, deeper than wide (head width 74–81 % of its depth). Snout distinctly pointed, smaller than eye diameter. Mouth supraterminal, slightly oblique in profile. Jaws subequal, lower jaw longer than upper, posterior end of rictus at same level or slightly ventral to centre of eye. Premaxilla and dentary with many irregularly distributed conical teeth on inner rows and slightly curved teeth on outer row. Orbit large, eye diameter 34–38 % HL, mainly in anterior half of head (snout to eye end length 51–59 % HL). Branchiostegal membrane projecting posteriorly from opercle.</paragraph>
<paragraph id="8BF736DEFF84DD5DFF4D827832F63C48" blockId="8.[151,1437,151,2035]" pageId="8" pageNumber="409">Dorsal-fin origin markedly posterior to anal-fin origin (predorsal length 109–115 as % of preanal length), both fins originating posterior to mid-length of body (predorsal length 63.3–67.9 % SL; preanal length 55.5–59.9 % SL). Dorsal fin relatively small with low number of rays,12–14; extremity rounded. Anal fin trapezoidal in shape, with 16–19 rays; lower margin mildly concave in profile. Dorsal and anal fins usually with short fin-ray extensions. Pectoral fin subtriangular, insertion slightly posterior to margin of opercular opening, base slightly oblique, upper fin rays positioned slightly anterior to lower fin rays, with tip reaching or slightly overlapping base of pelvic fin. Pelvic fin subabdominal, origin at about mid-length of body, short, base medially separated, with tip reaching urogenital papilla. Caudal fin rounded, with 13–18 branched rays, plus 2–3 dorsal and ventral procurrent rays; central 2–3 rays of fin commonly relatively slightly elongated resulting in mild central lobe.</paragraph>
<paragraph id="8BF736DEFF84DD5DFF4D813932243CD7" blockId="8.[151,1437,151,2035]" pageId="8" pageNumber="409">Scales cycloid, body and head entirely scaled, except for ventral surface of head. No scales on base of dorsal and anal fins. Scales in mid-longitudinal series 26–28 plus two or three small scales on caudal-fin base. Transverse rows of scales in front of dorsal-fin origin 11; scale rows around caudal peduncle 12.</paragraph>
<paragraph id="8BF736DEFF84DD5DFF4D81D534403D94" blockId="8.[151,1437,151,2035]" pageId="8" pageNumber="409">
Cephalic squamation pattern G-type. Frontal neuromast in single shallow groove behind upper lip. Cephalic sensory system at supraorbital level in an undivided, slightly curved, almost flat, shallow groove with three exposed neuromasts and pronounced lobes (
<figureCitation id="13732A5BFF84DD5DFDB9818C34E03D62" box="[563,649,758,784]" captionStart="FIGURE 3" captionStartId="10.[151,250,1307,1331]" captionTargetBox="[156,1428,706,1276]" captionTargetId="figure-280@10.[151,1436,702,1281]" captionTargetPageId="10" captionText="FIGURE 3. Cephalic structure: a, Nothobranchius janpapi; b, N. luekei; c, N. geminus; d, N. lourensi and N. fuscotaeniatus. Key to label abbreviations: an, anterior nostril; pn, posterior nostril; f, frontal neuromast; so, supraorbital canal; st, supratemporal canal." pageId="8" pageNumber="409">Fig. 3a</figureCitation>
), whereas at supratemporal level with three exposed neuromasts in slightly curved arrangement on flat surface, with pronounced lobes. Preorbital canal in shallow groove with 2–3 exposed neuromasts; infraorbital level with series of about 15 small buttons at ventral and posterior margin of orbit; postorbital canal in shallow groove with one exposed long neuromast. Preopercular system with six exposed neuromasts in deep groove on preopercle portion, whereas in shallow groove with four exposed neuromasts on ventral portion. Mandibular canal in shallow groove with about ten small buttons. One neuromast on each scale along trunk mid-longitudinal series.
</paragraph>
<paragraph id="8BF736DEFF84DD5DFF4D808A352F3B76" blockId="8.[151,1437,151,2035]" pageId="8" pageNumber="409">
Female: Smaller than male, maximum recorded size
<quantity id="4CB09B3BFF84DD5DFCA0808A35F83A79" box="[810,913,1008,1035]" metricMagnitude="-2" metricUnit="m" metricValue="2.8899999999999997" pageId="8" pageNumber="409" unit="mm" value="28.9">28.9 mm</quantity>
SL. Branchiostegal membrane not projecting posteriorly from opercle. Dorsal fin rounded with low number of rays, 12–14. Caudal fin basically rounded but with central 2–3 rays slightly elongated resulting in mildly pointed posterior margin. Anal fin subtriangular, tip rounded, central rays longer and more rigid. Anal fin positioned more posteriorly than in males (preanal length 60.5–65.2 % SL vs. 55.5–59.9). Anal-fin base length smaller than in males (18.7–20.4 % SL vs. 22.8–27.1). Head depth smaller than in males (67–69 % HL vs. 74–81; suborbital depth 11–14 % HL vs. 16–20), whereas head width
<quantity id="4CB09B3BFF84DD5DFB7587D933083ACF" box="[1279,1377,1187,1214]" metricMagnitude="0" metricUnit="m" metricValue="2.2225" metricValueMax="2.286" metricValueMin="2.159" pageId="8" pageNumber="409" unit="in" value="87.5" valueMax="90.0" valueMin="85.0">85–90 in</quantity>
% of its depth vs. 74–81. Caudal peduncle shallower than in male (caudal peduncle depth 10.7–11.8 % SL vs. 12.7–15.0; caudal peduncle length 184–190 % of its depth vs. 133–140).
</paragraph>
<paragraph id="8BF736DEFF84DD5DFF4D867435263981" blockId="8.[151,1437,151,2035]" pageId="8" pageNumber="409">
<emphasis id="B93CEACCFF84DD5DFF4D867437363B5A" bold="true" box="[199,351,1294,1320]" pageId="8" pageNumber="409">Colouration.</emphasis>
Live male (
<figureCitation id="13732A5BFF84DD5DFE608674341A3B5A" box="[490,627,1294,1320]" captionStart="FIGURE 2" captionStartId="9.[151,250,1410,1434]" captionTargetBox="[151,1435,179,1384]" captionTargetId="figure-19@9.[151,1435,179,1384]" captionTargetPageId="9" captionText="FIGURE 2. Wild-caught specimens of Nothobranchius janpapi and examples of typical habitat. A. N. janpapi Ndundu TAN 95- 10, male, lower Rufiji River drainage; B. N. janpapi Kwaraza TAN 95-14, male, lower Ruvu River drainage, in vicinity of the type locality; C. N. janpapi Soga TAN 97-45, male, Ruvu River drainage; D. N. janpapi Ruvu River TZN 09-4, female, lower Ruvu River drainage (photograph by B. Nagy); E. Location Ruvu River TAN 00-20, a typical thickly vegetated shallow habitat on the floodplain of the lower Ruvu River (photograph June 14, 2000), hosting N. janpapi, N. melanospilus, N. annectens, N. ocellatus and N. eggersi; F. Location Kikale TAN 02-10, thickly vegetated residual pool at a culvert in a seasonal stream course, peripheral to the northern margin of the Rufiji River delta (photograph June 3, 2002), hosting N. janpapi, N. annectens and N. melanospilus. Photographs by B.R. Watters except where otherwise noted." pageId="8" pageNumber="409">Figs. 2A–C</figureCitation>
): Scales on trunk and head light iridescent blue with narrow red to less commonly grey-brown margins, forming a regular reticulation pattern. Scales on abdomen faint blue to white with less pronounced margins. Scales on the dorsum anterior to dorsal fin may have yellow-brown centres. Snout, frontal and dorsal portions of the head are yellow-brown; the throat is yellow. Operculum has light blue markings. Exposed part of branchiostegal membrane is yellow with white edging. Iris is blue to blue-green. Dorsal fin background is green to yellow-green grading to grey in the distal zone with a light blue margin that may be white in the very short fin-ray extensions; the background is overlaid by small red spots that may be denser and larger in the basal and proximal zones, becoming smaller and, in some populations, dark grey in the medial and distal zones. Anal fin background comprises a light blue basal zone grading to yellow-brown or yellow-green in the proximal and medial zones which are overlaid by prominent red stripes over fin rays; in some populations these red stripes may be partially broken into irregular spots. The development of red stripes is variable at both the inter- and intrapopulation level; in some they are narrow, allowing the yellow-brown or yellow-green background to dominate, while in others they are broad and dense resulting in an almost solid red fin and merging with the commonly present orange-red distal zone. A narrow, often only partially developed, light blue margin is usually present on the anal fin. Basal, proximal and medial zones of the caudal fin are olive grading into a light grey to translucent subdistal zone that, in turn, grades into a darker grey outer component. Faint red spots may be present on the basal, proximal and submedial zones and may be arranged concentrically conforming to the general outer shape of the fin; less commonly, there may be radiating red stripes parallel to fin rays. Many populations also show orange-red markings along the lower, and more rarely upper, marginal parts of the distal zone of the caudal fin, extending from the fin base and decreasing anteriorly. Pelvic fins are orange-red with poorly developed blue-white edges. Pectoral fins are dominantly hyaline with poorly developed, and usually only partial, blue margins.
</paragraph>
<caption id="DF376656FF85DD5CFF1D86F835E238E4" pageId="9" pageNumber="410" startId="9.[151,250,1410,1434]" targetBox="[151,1435,179,1384]" targetPageId="9" targetType="figure">
<paragraph id="8BF736DEFF85DD5CFF1D86F835E238E4" blockId="9.[151,1437,1410,1686]" pageId="9" pageNumber="410">
<emphasis id="B93CEACCFF85DD5CFF1D86F8377B3BE8" bold="true" box="[151,274,1410,1434]" pageId="9" pageNumber="410">FIGURE 2.</emphasis>
Wild-caught specimens of
<taxonomicName id="4C484D5DFF85DD5CFDA886F835713BE8" authorityName="Wildekamp" authorityYear="1977" box="[546,792,1410,1434]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="410" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF85DD5CFDA886F835713BE8" box="[546,792,1410,1434]" italics="true" pageId="9" pageNumber="410">Nothobranchius janpapi</emphasis>
</taxonomicName>
and examples of typical habitat.
<emphasis id="B93CEACCFF85DD5CFBEA86F832103BE8" bold="true" box="[1120,1145,1410,1434]" pageId="9" pageNumber="410">A.</emphasis>
<taxonomicName id="4C484D5DFF85DD5CFBF486F9331C3BE8" authority="Ndundu TAN" authorityName="Ndundu TAN" box="[1150,1397,1410,1434]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="410" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF85DD5CFBF486F9328E3BE8" box="[1150,1255,1411,1434]" italics="true" pageId="9" pageNumber="410">N. janpapi</emphasis>
Ndundu TAN
</taxonomicName>
95- 10, male, lower Rufiji River drainage;
<emphasis id="B93CEACCFF85DD5CFDA886DC34503BCC" bold="true" box="[546,569,1446,1470]" pageId="9" pageNumber="410">B.</emphasis>
<taxonomicName id="4C484D5DFF85DD5CFDCA86DD352D3BCC" authority="Kwaraza TAN" authorityName="Kwaraza TAN" box="[576,836,1446,1470]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="410" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF85DD5CFDCA86DD34C23BCC" box="[576,683,1447,1470]" italics="true" pageId="9" pageNumber="410">N. janpapi</emphasis>
Kwaraza TAN
</taxonomicName>
95-14, male, lower Ruvu River drainage, in vicinity of the type locality;
<emphasis id="B93CEACCFF85DD5CFEA986B037553B90" bold="true" box="[291,316,1482,1506]" pageId="9" pageNumber="410">C.</emphasis>
<taxonomicName id="4C484D5DFF85DD5CFEC986B134493B90" authority="Soga TAN" authorityName="Soga TAN" box="[323,544,1482,1506]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="410" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF85DD5CFEC986B137C73B90" box="[323,430,1483,1506]" italics="true" pageId="9" pageNumber="410">N. janpapi</emphasis>
Soga TAN
</taxonomicName>
97-45, male, Ruvu River drainage;
<emphasis id="B93CEACCFF85DD5CFC0586B035C13B90" bold="true" box="[911,936,1482,1506]" pageId="9" pageNumber="410">D.</emphasis>
<taxonomicName id="4C484D5DFF85DD5CFC2486B132A73B90" authority="Ruvu River TZN" authorityName="Ruvu River TZN" box="[942,1230,1482,1506]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="410" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF85DD5CFC2486B132703B90" box="[942,1049,1483,1506]" italics="true" pageId="9" pageNumber="410">N. janpapi</emphasis>
Ruvu River TZN
</taxonomicName>
09-4, female, lower Ruvu River drainage (photograph by B. Nagy);
<emphasis id="B93CEACCFF85DD5CFDF1869434FB3874" bold="true" box="[635,658,1518,1542]" pageId="9" pageNumber="410">E.</emphasis>
Location Ruvu River TAN 00-20, a typical thickly vegetated shallow habitat on the floodplain of the lower Ruvu River (photograph June 14, 2000), hosting
<taxonomicName id="4C484D5DFF85DD5CFC428569325D3858" box="[968,1076,1555,1578]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="410" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF85DD5CFC428569325D3858" box="[968,1076,1555,1578]" italics="true" pageId="9" pageNumber="410">N. janpapi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C484D5DFF85DD5CFBC88569328F3858" authorityName="TZH" authorityYear="2018" box="[1090,1254,1554,1578]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="410" phylum="Chordata" rank="species" species="melanospilus">
<emphasis id="B93CEACCFF85DD5CFBC88569328F3858" box="[1090,1254,1554,1578]" italics="true" pageId="9" pageNumber="410">N. melanospilus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C484D5DFF85DD5CFB7E8569331E3858" authorityName="Watters, Wildekamp &amp; Cooper" authorityYear="1998" box="[1268,1399,1555,1578]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="410" phylum="Chordata" rank="species" species="annectens">
<emphasis id="B93CEACCFF85DD5CFB7E8569331E3858" box="[1268,1399,1555,1578]" italics="true" pageId="9" pageNumber="410">N. annectens</emphasis>
</taxonomicName>
,
<taxonomicName id="4C484D5DFF85DD5CFA0F8569369B383C" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="410" phylum="Chordata" rank="species" species="ocellatus">
<emphasis id="B93CEACCFF85DD5CFA0F8569369B383C" italics="true" pageId="9" pageNumber="410">N. ocellatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C484D5DFF85DD5CFEA8854D37E3383C" authorityName="Seegers" authorityYear="1982" box="[290,394,1591,1614]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="410" phylum="Chordata" rank="species" species="eggersi">
<emphasis id="B93CEACCFF85DD5CFEA8854D37E3383C" box="[290,394,1591,1614]" italics="true" pageId="9" pageNumber="410">N. eggersi</emphasis>
</taxonomicName>
;
<emphasis id="B93CEACCFF85DD5CFE1C854C37C0383C" bold="true" box="[406,425,1590,1614]" pageId="9" pageNumber="410">F.</emphasis>
Location Kikale TAN 02-10, thickly vegetated residual pool at a culvert in a seasonal stream course, peripheral to the northern margin of the Rufiji River delta (photograph June 3, 2002), hosting
<taxonomicName id="4C484D5DFF85DD5CFBDD852132AB3800" box="[1111,1218,1627,1650]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="410" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF85DD5CFBDD852132AB3800" box="[1111,1218,1627,1650]" italics="true" pageId="9" pageNumber="410">N. janpapi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C484D5DFF85DD5CFB458521333B3800" authorityName="Watters, Wildekamp &amp; Cooper" authorityYear="1998" box="[1231,1362,1627,1650]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="410" phylum="Chordata" rank="species" species="annectens">
<emphasis id="B93CEACCFF85DD5CFB458521333B3800" box="[1231,1362,1627,1650]" italics="true" pageId="9" pageNumber="410">N. annectens</emphasis>
</taxonomicName>
and
<taxonomicName id="4C484D5DFF85DD5CFA0F8521377438E4" authorityName="TZH" authorityYear="2018" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="410" phylum="Chordata" rank="species" species="melanospilus">
<emphasis id="B93CEACCFF85DD5CFA0F8521377438E4" italics="true" pageId="9" pageNumber="410">N. melanospilus</emphasis>
</taxonomicName>
. Photographs by B.R. Watters except where otherwise noted.
</paragraph>
</caption>
<paragraph id="8BF736DEFF85DD5CFF4D85BE322D3954" blockId="9.[151,1437,1732,2011]" pageId="9" pageNumber="410">
Live female (
<figureCitation id="13732A5BFF85DD5CFEE885BE37D638AC" box="[354,447,1732,1758]" captionStart="FIGURE 2" captionStartId="9.[151,250,1410,1434]" captionTargetBox="[151,1435,179,1384]" captionTargetId="figure-19@9.[151,1435,179,1384]" captionTargetPageId="9" captionText="FIGURE 2. Wild-caught specimens of Nothobranchius janpapi and examples of typical habitat. A. N. janpapi Ndundu TAN 95- 10, male, lower Rufiji River drainage; B. N. janpapi Kwaraza TAN 95-14, male, lower Ruvu River drainage, in vicinity of the type locality; C. N. janpapi Soga TAN 97-45, male, Ruvu River drainage; D. N. janpapi Ruvu River TZN 09-4, female, lower Ruvu River drainage (photograph by B. Nagy); E. Location Ruvu River TAN 00-20, a typical thickly vegetated shallow habitat on the floodplain of the lower Ruvu River (photograph June 14, 2000), hosting N. janpapi, N. melanospilus, N. annectens, N. ocellatus and N. eggersi; F. Location Kikale TAN 02-10, thickly vegetated residual pool at a culvert in a seasonal stream course, peripheral to the northern margin of the Rufiji River delta (photograph June 3, 2002), hosting N. janpapi, N. annectens and N. melanospilus. Photographs by B.R. Watters except where otherwise noted." pageId="9" pageNumber="410">Fig. 2D</figureCitation>
): Trunk and head light grey, darker on the dorsum and lighter ventrally. Scales on the flanks have narrow darker grey-brown margins; scale centres on the trunk above the abdomen are iridescent pale blue; the operculum is iridescent light blue. All fins hyaline. The iris is light golden.
</paragraph>
</subSubSection>
<subSubSection id="C3526555FF85DD5FFF4D844A34D33CDB" lastPageId="10" lastPageNumber="411" pageId="9" pageNumber="410" type="distribution">
<paragraph id="8BF736DEFF85DD5FFF4D844A32153FFB" blockId="9.[151,1437,1732,2011]" lastBlockId="10.[151,1437,151,681]" lastPageId="10" lastPageNumber="411" pageId="9" pageNumber="410">
<emphasis id="B93CEACCFF85DD5CFF4D844A37083938" bold="true" box="[199,353,1840,1866]" pageId="9" pageNumber="410">Distribution.</emphasis>
<taxonomicName id="4C484D5DFF85DD5CFEE0844A34173938" box="[362,638,1840,1866]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="410" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF85DD5CFEE0844A34173938" box="[362,638,1840,1866]" italics="true" pageId="9" pageNumber="410">Nothobranchius janpapi</emphasis>
</taxonomicName>
is endemic to seasonal freshwater habitats in coastal
<collectingCountry id="F35F764EFF85DD5CFB63844A33273938" box="[1257,1358,1840,1866]" name="Tanzania" pageId="9" pageNumber="410">Tanzania</collectingCountry>
and is the most common, but second most widespread, member of the
<taxonomicName id="4C484D5DFF85DD5CFCE8842E3273391C" authorityName="Wildekamp" authorityYear="1977" box="[866,1050,1876,1902]" class="Actinopterygii" family="Nothobranchiidae" genus="Aphyobranchius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
<emphasis id="B93CEACCFF85DD5CFCE8842E3273391C" box="[866,1050,1876,1902]" italics="true" pageId="9" pageNumber="410">Aphyobranchius</emphasis>
</taxonomicName>
subgenus, occurring mainly in the Ruvu and Rufiji river systems (
<figureCitation id="13732A5BFF85DD5CFE738402345339E0" box="[505,570,1912,1938]" captionStart="FIGURE 1" captionStartId="2.[151,250,1762,1786]" captionTargetBox="[151,1436,374,1736]" captionTargetId="figure-118@2.[151,1436,374,1736]" captionTargetPageId="2" captionText="FIGURE 1. Distribution of species belonging to the subgenus Aphyobranchius, all of which occur in the east-central and southern coastal lowlands of Tanzania: N. janpapi (yellow-filled circle), N. luekei (red-filled triangle), N. geminus (orange-filled diamond), N. lourensi (green-filled square), N. fuscotaeniatus (blue-filled inverted triangle), N. cf. janpapi (dark magenta-filled circle). The inset map shows the area of distribution in a somewhat broader geographic context, to include the distribution of N. willerti (red-filled star) in the lower Tana River area of northeastern coastal Kenya, a species that herein is considered to belong to the subgenus Adiniops, rather than Aphyobranchius to which it was ascribed in the original description by Wildekamp (1992). Note that individual symbols on the map may represent more than one locality or population where the sites are close to one another and could not reasonably be resolved at the scale of the map." pageId="9" pageNumber="410">Fig. 1</figureCitation>
). It occurs in the middle and lower reaches of the latter and throughout most of the former but dominantly in the lower reaches. The westernmost locality known to date is in the Kisaki area on the floodplain of the Mgeta River, a tributary of the upper Ruvu River system. There is currently a lack of range data available for the middle reaches of the Ruvu River, but this can probably be attributed to a lack of investigation in that area. The northern extent of distribution is in the southern part of the Wami River delta where it merges with the coastal plains associated with the Ruvu River. The southernmost occurrences are in the coastal floodplains immediately south of the Rufiji delta, probably extending to the lower Matandu River system where a population of what has provisionally been referred to as “
<taxonomicName id="4C484D5DFF86DD5FFD15825D35533F32" box="[671,826,295,321]" family="Nothobranchiidae" genus="Nothobranchius" isUncertain="true" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="411" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF86DD5FFD15825D34D33F32" box="[671,698,295,320]" italics="true" pageId="10" pageNumber="411">N.</emphasis>
cf.
<emphasis id="B93CEACCFF86DD5FFD6E825D35533F32" box="[740,826,295,320]" italics="true" pageId="10" pageNumber="411">janpapi</emphasis>
</taxonomicName>
” is present. The affiliation of this southern population is presently being studied. A single, relatively isolated, occurrence is also known from the Luhule/Luhute River floodplain within the so-called Mbezi Triangle (
<emphasis id="B93CEACCFF86DD5FFD3B821534863FFA" box="[689,751,367,392]" italics="true" pageId="10" pageNumber="411">sensu</emphasis>
<bibRefCitation id="EFD94B2FFF86DD5FFD7C821535A53FFB" author="Watters, B. R. &amp; Cooper, B. J. &amp; Wildekamp, R. H." box="[758,972,367,393]" pageId="10" pageNumber="411" pagination="5 - 6" refId="ref35078" refString="Watters, B. R., Cooper, B. J. &amp; Wildekamp, R. H. (2007) Description of Nothobranchius cardinalis spec. nov. (Cyprinodontiformes: Aplocheilidae), an annual fish from the Mbwemkuru River basin, Tanzania. Journal of the American Killifish Association, 40 (5 - 6), 129 - 145. [issue printed in 2008 = year of species description]" type="journal article" year="2007">
Watters
<emphasis id="B93CEACCFF86DD5FFCD8821535EA3FFB" box="[850,899,367,393]" italics="true" pageId="10" pageNumber="411">et al</emphasis>
. 2007
</bibRefCitation>
;
<bibRefCitation id="EFD94B2FFF86DD5FFC5D821532193FFB" author="Watters, B. R." box="[983,1136,367,393]" pageId="10" pageNumber="411" pagination="37 - 76" refId="ref34941" refString="Watters, B. R. (2009) The ecology and distribution of Nothobranchius fishes. Journal of the American Killifish Association, 42 (2), 37 - 76." type="journal article" year="2009">Watters 2009</bibRefCitation>
).
</paragraph>
<paragraph id="8BF736DEFF86DD5FFF4D82E834D33CDB" blockId="10.[151,1437,151,681]" pageId="10" pageNumber="411">
<emphasis id="B93CEACCFF86DD5FFF4D82E837D53FDF" bold="true" box="[199,444,402,429]" pageId="10" pageNumber="411">Conservation status.</emphasis>
<taxonomicName id="4C484D5DFF86DD5FFE4982E934BC3FDE" box="[451,725,403,429]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="411" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF86DD5FFE4982E934BC3FDE" box="[451,725,403,429]" italics="true" pageId="10" pageNumber="411">Nothobranchius janpapi</emphasis>
</taxonomicName>
has been evaluated as Least Concern (
<bibRefCitation id="EFD94B2FFF86DD5FFB0382E933F83FDF" author="Nagy, B. &amp; Watters, B." box="[1161,1425,403,429]" pageId="10" pageNumber="411" refId="ref33584" refString="Nagy, B. &amp; Watters, B. (2019 a) Nothobranchius janpapi. The IUCN Red List of Threatened Species, 2019, e. T 60368 A 47185381. https: // doi. org / 10.2305 / IUCN. UK. 2019 - 3. RLTS. T 60368 A 47185381. en" type="journal volume" year="2019">Nagy &amp; Watters 2019a</bibRefCitation>
). The known extent of occurrence (EOO) is
<quantity id="4CB09B3BFF86DD5FFDFE82CD34863FA1" box="[628,751,439,467]" metricMagnitude="7" metricUnit="m" metricValue="2.279" pageId="10" pageNumber="411" unit="km" value="22790.0">22,790 km</quantity>
². Within its area of distribution, the occurrence of the species is fragmented but it seems to be relatively widespread, and it is known from more than 30 threat-based locations in several river drainages. These factors make it the least threatened member of the
<taxonomicName id="4C484D5DFF86DD5FFBE18285334A3C6B" authorityName="Wildekamp" authorityYear="1977" box="[1131,1315,511,537]" class="Actinopterygii" family="Nothobranchiidae" genus="Aphyobranchius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="411" phylum="Chordata" rank="genus">
<emphasis id="B93CEACCFF86DD5FFBE18285334A3C6B" box="[1131,1315,511,537]" italics="true" pageId="10" pageNumber="411">Aphyobranchius</emphasis>
</taxonomicName>
subgenus. However, there is increasing habitat degradation due to expansion of agriculture and urbanisation across its range, including the conversion of some seasonal habitats into permanent ponds (e.g. in the Kwaraza area of the lower Ruvu River system, where the
<typeStatus id="54F3887CFF86DD5FFE64811134773CF7" box="[494,542,619,645]" pageId="10" pageNumber="411">type</typeStatus>
locality occurs), thereby modifying them in ways that render them unsuitable to support the semi-annual life cycle of the species.
</paragraph>
</subSubSection>
</treatment>
</document>

604
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<heading id="D0BF81B2FF99DD40FF1D876034AA3A46" bold="true" box="[151,707,1050,1077]" fontSize="11" level="1" pageId="21" pageNumber="422" reason="1">
<taxonomicName id="4C484D5DFF99DD40FF1D876034AA3A46" ID-CoL="47SLQ" authority="Seegers, 1997" authorityName="Seegers" authorityYear="1997" box="[151,707,1050,1077]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="21" pageNumber="422" phylum="Chordata" rank="species" species="fuscotaeniatus">
<emphasis id="B93CEACCFF99DD40FF1D876034AA3A46" bold="true" box="[151,707,1050,1077]" pageId="21" pageNumber="422">
<emphasis id="B93CEACCFF99DD40FF1D8760347D3A46" bold="true" box="[151,532,1050,1077]" italics="true" pageId="21" pageNumber="422">Nothobranchius fuscotaeniatus</emphasis>
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<paragraph id="8BF736DEFF99DD40FF1D874537423A2B" blockId="21.[151,707,1050,1113]" box="[151,299,1087,1113]" pageId="21" pageNumber="422">
(
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)
</paragraph>
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<subSubSection id="C3526555FF99DD40FF1D87FC331C3B43" pageId="21" pageNumber="422" type="materials_examined">
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<emphasis id="B93CEACCFF99DD40FF1D87FC37E83AD2" bold="true" box="[151,385,1158,1184]" pageId="21" pageNumber="422">Material examined.</emphasis>
<materialsCitation id="3B203C83FF99DD40FE0287FD35623AB7" collectionCode="RMCA" collectorName="B. Watters" country="Tanzania" latitude="-8.052167" location="Rufiji" longLatPrecision="13" longitude="39.008835" municipality="Kitonga" pageId="21" pageNumber="422" specimenCode="RMCA 1997.082" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCode id="DBEE9EA5FF99DD40FE0287FD34243AD3" box="[392,589,1159,1185]" collectionCode="RMCA" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="21" pageNumber="422" type="Museum">RMCA 1997.082</specimenCode>
.P.0032-0033,
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SL,
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SL;
<collectingCountry id="F35F764EFF99DD40FB6D87FD33393AD3" box="[1255,1360,1159,1185]" name="Tanzania" pageId="21" pageNumber="422">Tanzania</collectingCountry>
:
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drainage,
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;
<collectorName id="26BD5308FF99DD40FD1987D135623AB7" box="[659,779,1195,1221]" pageId="21" pageNumber="422">B. Watters</collectorName>
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,
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<collectionCode id="ED59AE1BFF99DD40FC9F87D135423AB7" box="[789,811,1195,1221]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="21" pageNumber="422">R</collectionCode>
.
<collectorName id="26BD5308FF99DD40FCBF87D135B33AB7" box="[821,986,1195,1221]" pageId="21" pageNumber="422">Wildekamp, B</collectorName>
. Cooper;
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<collectingDate id="EFB2E9F6FF99DD40FBCF87D132B43AB7" box="[1093,1245,1195,1221]" pageId="21" pageNumber="422" value="1997-05-31">31 May 1997</collectingDate>
</date>
; field code: TAN 97-9
</materialsCitation>
.
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<emphasis id="B93CEACCFF99DD40FF4D8788345D3B7E" bold="true" box="[199,564,1266,1292]" pageId="21" pageNumber="422">Additional material examined.</emphasis>
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,
<quantity id="4CB09B3BFF99DD40FD158788352B3B7F" box="[671,834,1266,1293]" metricMagnitude="-2" metricUnit="m" metricValue="3.925" metricValueMax="4.34" metricValueMin="3.51" pageId="21" pageNumber="422" unit="mm" value="39.25" valueMax="43.4" valueMin="35.1">35.1–43.4 mm</quantity>
SL,
<materialsCitation id="3B203C83FF99DD40FCFD8789331B3B43" collectingDate="1997-05-31" collectionCode="R" collectorName="B. Watters &amp; Wildekamp, B" country="Tanzania" latitude="-8.052167" location="Rufiji" longLatPrecision="13" longitude="39.008835" municipality="Kitonga" pageId="21" pageNumber="422" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D4EFD57FF99DD40FCFD878935B13B7F" box="[887,984,1267,1293]" count="1" pageId="21" pageNumber="422" type="female">1 female</specimenCount>
<quantity id="4CB09B3BFF99DD40FC6A8788322D3B7F" box="[992,1092,1266,1293]" metricMagnitude="-2" metricUnit="m" metricValue="2.59" pageId="21" pageNumber="422" unit="mm" value="25.9">25.9 mm</quantity>
SL;
<collectingCountry id="F35F764EFF99DD40FBF38789328B3B7F" box="[1145,1250,1267,1293]" name="Tanzania" pageId="21" pageNumber="422">Tanzania</collectingCountry>
:
<location id="8E976005FF99DD40FB66878933443B7F" LSID="urn:lsid:plazi:treatment:03E187C8FF99DD4DFF1D876034A6393F:8E976005FF99DD40FB66878933443B7F" box="[1260,1325,1267,1293]" country="Tanzania" latitude="-8.052167" longLatPrecision="13" longitude="39.008835" municipality="Kitonga" name="Rufiji" pageId="21" pageNumber="422">Rufiji</location>
drainage,
<collectingMunicipality id="6B93ACA4FF99DD40FF1D866D369C3B43" box="[151,245,1303,1329]" pageId="21" pageNumber="422">Kitonga</collectingMunicipality>
;
<geoCoordinate id="EE7C5019FF99DD40FE8A866D37EF3B43" box="[256,390,1303,1329]" degrees="08" direction="south" minutes="03.13" orientation="latitude" pageId="21" pageNumber="422" precision="9" value="-8.052167">08°03.13’ S</geoCoordinate>
,
<geoCoordinate id="EE7C5019FF99DD40FE18866C34723B43" box="[402,539,1302,1329]" degrees="39" direction="east" minutes="00.53" orientation="longitude" pageId="21" pageNumber="422" precision="9" value="39.008835">39°00.53’ E</geoCoordinate>
;
<collectorName id="26BD5308FF99DD40FDAD866D34F63B43" box="[551,671,1303,1329]" pageId="21" pageNumber="422">B. Watters</collectorName>
,
<collectionCode id="ED59AE1BFF99DD40FD20866D34A93B43" box="[682,704,1303,1329]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="21" pageNumber="422">R</collectionCode>
.
<collectorName id="26BD5308FF99DD40FD40866D35063B43" box="[714,879,1303,1329]" pageId="21" pageNumber="422">Wildekamp, B</collectorName>
. Cooper;
<date id="FFF6101EFF99DD40FC51866D321A3B43" box="[987,1139,1303,1329]" pageId="21" pageNumber="422" value="1997-05-31">
<collectingDate id="EFB2E9F6FF99DD40FC51866D321A3B43" box="[987,1139,1303,1329]" pageId="21" pageNumber="422" value="1997-05-31">31 May 1997</collectingDate>
</date>
; field code: TAN 97-9
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3526555FF99DD40FF4D8640335D38CF" pageId="21" pageNumber="422" type="diagnosis">
<paragraph id="8BF736DEFF99DD40FF4D8640335D38CF" blockId="21.[151,1437,1158,2013]" pageId="21" pageNumber="422">
<emphasis id="B93CEACCFF99DD40FF4D8640372B3B26" bold="true" box="[199,322,1338,1364]" pageId="21" pageNumber="422">Diagnosis.</emphasis>
<taxonomicName id="4C484D5DFF99DD40FEC1864134C33B27" authorityName="Seegers" authorityYear="1997" box="[331,682,1339,1365]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="21" pageNumber="422" phylum="Chordata" rank="species" species="fuscotaeniatus">
<emphasis id="B93CEACCFF99DD40FEC1864134C33B27" box="[331,682,1339,1365]" italics="true" pageId="21" pageNumber="422">Nothobranchius fuscotaeniatus</emphasis>
</taxonomicName>
is here assigned to the
<taxonomicName id="4C484D5DFF99DD40FC37864132533B27" box="[957,1082,1339,1365]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="21" pageNumber="422" phylum="Chordata" rank="species" species="lourensi">
<emphasis id="B93CEACCFF99DD40FC37864132533B27" box="[957,1082,1339,1365]" italics="true" pageId="21" pageNumber="422">N. lourensi</emphasis>
</taxonomicName>
species group of the subgenus
<taxonomicName id="4C484D5DFF99DD40FF1D862537263B0B" authorityName="Wildekamp" authorityYear="1977" box="[151,335,1375,1401]" class="Actinopterygii" family="Nothobranchiidae" genus="Aphyobranchius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyprinodontiformes" pageId="21" pageNumber="422" phylum="Chordata" rank="genus">
<emphasis id="B93CEACCFF99DD40FF1D862537263B0B" box="[151,335,1375,1401]" italics="true" pageId="21" pageNumber="422">Aphyobranchius</emphasis>
</taxonomicName>
by presenting the following combination of characters in males, compared to those of other subgenera: anal fin trapezoidal in shape with a concave lower margin (vs. rounded with convex lower margin); caudal fin rounded with 2–3 central rays slightly elongated forming a mild central lobe (vs. central lobe absent); spawning behaviour in mid-water levels (vs. spawning directly into the substrate), and molecular phylogenetic relationships.
<taxonomicName id="4C484D5DFF99DD40FF1D8695379E387B" authorityName="Seegers" authorityYear="1997" box="[151,503,1519,1545]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="21" pageNumber="422" phylum="Chordata" rank="species" species="fuscotaeniatus">
<emphasis id="B93CEACCFF99DD40FF1D8695379E387B" box="[151,503,1519,1545]" italics="true" pageId="21" pageNumber="422">Nothobranchius fuscotaeniatus</emphasis>
</taxonomicName>
is further distinguished from all the other members of the genus
<taxonomicName id="4C484D5DFF99DD40FB62869533F5387B" authorityName="Peters" authorityYear="1868" box="[1256,1436,1519,1545]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="21" pageNumber="422" phylum="Chordata" rank="genus">
<emphasis id="B93CEACCFF99DD40FB62869533F5387B" box="[1256,1436,1519,1545]" italics="true" pageId="21" pageNumber="422">Nothobranchius</emphasis>
</taxonomicName>
by the unique diagnostic characters of male colouration comprising a green-blue body with 10–12 distinct red-brown crossbars on entire trunk that extend into dorsal and anal fins; anal and dorsal fins green-blue with crossbars extending from trunk into the proximal and medial parts and with broad black distal band; caudal fin with slender yellow to yellow-green medial band and broad black distal band; as well as female colouration with a unique pattern of light blue and red-brown crossbars on the body and a light blue iridescence on the dorsal and anal fins.
</paragraph>
</subSubSection>
<subSubSection id="C3526555FF99DD4DFF4D85BC34383B43" lastPageId="24" lastPageNumber="425" pageId="21" pageNumber="422" type="description">
<paragraph id="8BF736DEFF99DD40FF4D85BC34F43977" blockId="21.[151,1437,1158,2013]" pageId="21" pageNumber="422">
<emphasis id="B93CEACCFF99DD40FF4D85BC37303892" bold="true" box="[199,345,1734,1760]" pageId="21" pageNumber="422">Description.</emphasis>
General body features illustrated by
<figureCitation id="13732A5BFF99DD40FC9A85BD35DD3893" box="[784,948,1735,1761]" captionStart="FIGURE 7" captionStartId="23.[151,250,1818,1842]" captionTargetBox="[151,1436,974,1796]" captionTargetId="figure-379@23.[151,1436,974,1796]" captionTargetPageId="23" captionText="FIGURE 7. Wild-caught specimens of Nothobranchius fuscotaeniatus and only known locality. A. N. fuscotaeniatus Kitonga north TAN 97-9, male, Rufiji River drainage; B. N. fuscotaeniatus Kitonga north TAN 97-9, female, Rufiji River drainage; C. N. fuscotaeniatus Kitonga north TAN 97-9, Rufiji River floodplain, type locality, shallow with dense vegetation (photograph May 31, 1997), hosting N. fuscotaeniatus, N. janpapi and N. lourensi; D. Same site as in “C”, photograph by B. Nagy in 2017. Multiple visits to the site subsequent to the original discovery in 1997 showed an absence of the species, due to substantial changes in the habitat, primarily the loss of dense vegetation and contamination of the habitat by soil washed off the sinceabandoned adjacent road. Photographs by B.R. Watters except where otherwise noted." pageId="21" pageNumber="422">Figures 7A–B</figureCitation>
. Morphometric and meristic characters of examined material are summarized in
<tableCitation id="C6CA0365FF99DD40FDC9859134F03977" box="[579,665,1771,1797]" captionStart="TABLE 7" captionStartId="22.[151,239,151,175]" captionText="TABLE 7. Morphometric and meristic data of Nothobranchius fuscotaeniatus. SD, standard deviation." pageId="21" pageNumber="422">Table 7</tableCitation>
.
</paragraph>
<paragraph id="8BF736DEFF99DD40FF4D8475327D39AF" blockId="21.[151,1437,1158,2013]" pageId="21" pageNumber="422">
Male: Medium-sized
<taxonomicName id="4C484D5DFF99DD40FE3F847534AA395B" box="[437,707,1807,1833]" pageId="21" pageNumber="422">
<emphasis id="B93CEACCFF99DD40FE3F84753400395B" box="[437,617,1807,1833]" italics="true" pageId="21" pageNumber="422">Nothobranchius</emphasis>
species
</taxonomicName>
, maximum recorded size
<quantity id="4CB09B3BFF99DD40FC5184753254395B" box="[987,1085,1807,1833]" metricMagnitude="-2" metricUnit="m" metricValue="4.34" pageId="21" pageNumber="422" unit="mm" value="43.4">43.4 mm</quantity>
SL. General body shape robust, laterally compressed and deep. Greatest body depth at vertical passing through pelvic-fin origin, 32.5–35.3 % SL. Greatest body width at pectoral-fin base with body progressively narrowing towards caudal-fin base. Dorsal profile convex from tip of snout to dorsal-fin origin, straight to slightly concave on caudal peduncle. Ventral profile convex from lower jaw to base of last anal-fin ray, straight to slightly concave on caudal peduncle. Caudal peduncle deep, length 146–153 % of its depth. Anus situated directly in front of anal-fin origin.
</paragraph>
<caption id="DF376656FF9ADD43FF1D83ED32893EDD" box="[151,1248,151,176]" pageId="22" pageNumber="423" startId="22.[151,239,151,175]" targetType="table">
<paragraph id="8BF736DEFF9ADD43FF1D83ED32893EDD" blockId="22.[151,1248,151,176]" box="[151,1248,151,176]" pageId="22" pageNumber="423">
<emphasis id="B93CEACCFF9ADD43FF1D83ED37633EC2" bold="true" box="[151,266,151,176]" pageId="22" pageNumber="423">TABLE 7.</emphasis>
Morphometric and meristic data of
<taxonomicName id="4C484D5DFF9ADD43FD0183ED35B13EC2" authorityName="Seegers" authorityYear="1997" box="[651,984,151,176]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="22" pageNumber="423" phylum="Chordata" rank="species" species="fuscotaeniatus">
<emphasis id="B93CEACCFF9ADD43FD0183ED35B13EC2" box="[651,984,151,176]" italics="true" pageId="22" pageNumber="423">Nothobranchius fuscotaeniatus</emphasis>
</taxonomicName>
. SD, standard deviation.
</paragraph>
</caption>
<paragraph id="8BF736DEFF9ADD43FCD883BA336A38F3" pageId="22" pageNumber="423">
<table id="F948C47EFF9A22AAFF1383BA331B38F3" box="[153,1394,192,1665]" colsContinueFrom="18.[153,1385,553,2026]" gridcols="7" gridrows="36" pageId="22" pageNumber="423">
<tr id="3578349CFF9A22AAFF1383BA331B3EAA" box="[153,1394,192,216]" gridrow="0" pageId="22" pageNumber="423" rowspan-0="1" rowspan-3="1" rowspan-4="1" rowspan-6="1">
<th id="76A95DE0FF9A22AAFD5383BA35C73EAA" box="[729,942,192,216]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="22" pageNumber="423">males</th>
<th id="76A95DE0FF9A22AAFB4783BA334A3EAA" box="[1229,1315,192,216]" gridcol="5" gridrow="0" pageId="22" pageNumber="423">females</th>
</tr>
<tr id="3578349CFF9A22AAFF138392331B3F72" box="[153,1394,232,256]" gridrow="1" pageId="22" pageNumber="423" rowspan-0="1" rowspan-3="1" rowspan-4="1" rowspan-6="1">
<td id="76A95DE0FF9A22AAFD53839235C73F72" box="[729,942,232,256]" colspan="2" colspanRight="1" gridcol="1" gridrow="1" pageId="22" pageNumber="423">(n = 3)</td>
<td id="76A95DE0FF9A22AAFB478392334A3F72" box="[1229,1315,232,256]" gridcol="5" gridrow="1" pageId="22" pageNumber="423">(n = 2)</td>
</tr>
<tr id="3578349CFF9A22AAFF138243331B3F23" box="[153,1394,313,337]" gridrow="2" pageId="22" pageNumber="423" rowspan-0="1">
<td id="76A95DE0FF9A22AAFD538243353E3F23" box="[729,855,313,337]" gridcol="1" gridrow="2" pageId="22" pageNumber="423">range</td>
<td id="76A95DE0FF9A22AAFCFF824335C73F23" box="[885,942,313,337]" gridcol="2" gridrow="2" pageId="22" pageNumber="423">mean</td>
<td id="76A95DE0FF9A22AAFC4F8243358C3F23" box="[965,997,313,337]" gridcol="3" gridrow="2" pageId="22" pageNumber="423">SD</td>
<td id="76A95DE0FF9A22AAFBC4824332DA3F23" box="[1102,1203,313,337]" gridcol="4" gridrow="2" pageId="22" pageNumber="423">range</td>
<td id="76A95DE0FF9A22AAFB478243334A3F23" box="[1229,1315,313,337]" gridcol="5" gridrow="2" pageId="22" pageNumber="423">mean</td>
<td id="76A95DE0FF9A22AAFAD88243331B3F23" box="[1362,1394,313,337]" gridcol="6" gridrow="2" pageId="22" pageNumber="423">SD</td>
</tr>
<tr id="3578349CFF9A22AAFF13821B331B3F0B" box="[153,1394,353,377]" gridrow="3" pageId="22" pageNumber="423" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1">
<th id="76A95DE0FF9A22AAFF13821B34CF3F0B" box="[153,678,353,377]" gridcol="0" gridrow="3" pageId="22" pageNumber="423">Standard length</th>
<td id="76A95DE0FF9A22AAFD53821B353E3F0B" box="[729,855,353,377]" gridcol="1" gridrow="3" pageId="22" pageNumber="423">35.1–43.4</td>
<td id="76A95DE0FF9A22AAFBC4821B32DA3F0B" box="[1102,1203,353,377]" gridcol="4" gridrow="3" pageId="22" pageNumber="423">24.2–25.9</td>
</tr>
<tr id="3578349CFF9A22AAFF1382F3331B3FD3" box="[153,1394,393,417]" gridrow="4" pageId="22" pageNumber="423">
<th id="76A95DE0FF9A22AAFF1382F334CF3FD3" box="[153,678,393,417]" gridcol="0" gridrow="4" pageId="22" pageNumber="423">
<emphasis id="B93CEACCFF9ADD43FF1382F337D13FD3" bold="true" box="[153,440,393,417]" pageId="22" pageNumber="423">Percent of standard length</emphasis>
</th>
<td id="76A95DE0FF9A22AAFD5382F3353E3FD3" box="[729,855,393,417]" gridcol="1" gridrow="4" pageId="22" pageNumber="423">min–max</td>
<td id="76A95DE0FF9A22AAFCFF82F335C73FD3" box="[885,942,393,417]" gridcol="2" gridrow="4" pageId="22" pageNumber="423">mean</td>
<td id="76A95DE0FF9A22AAFC4F82F3358C3FD3" box="[965,997,393,417]" gridcol="3" gridrow="4" pageId="22" pageNumber="423">SD</td>
<td id="76A95DE0FF9A22AAFBC482F332DA3FD3" box="[1102,1203,393,417]" gridcol="4" gridrow="4" pageId="22" pageNumber="423">min–max</td>
<td id="76A95DE0FF9A22AAFB4782F3334A3FD3" box="[1229,1315,393,417]" gridcol="5" gridrow="4" pageId="22" pageNumber="423">mean</td>
<td id="76A95DE0FF9A22AAFAD882F3331B3FD3" box="[1362,1394,393,417]" gridcol="6" gridrow="4" pageId="22" pageNumber="423">SD</td>
</tr>
<tr id="3578349CFF9A22AAFF1382CB331B3FBB" box="[153,1394,433,457]" gridrow="5" pageId="22" pageNumber="423">
<th id="76A95DE0FF9A22AAFF1382CB34CF3FBB" box="[153,678,433,457]" gridcol="0" gridrow="5" pageId="22" pageNumber="423">Total length</th>
<td id="76A95DE0FF9A22AAFD5382CB353E3FBB" box="[729,855,433,457]" gridcol="1" gridrow="5" pageId="22" pageNumber="423">123.3–124.8</td>
<td id="76A95DE0FF9A22AAFCFF82CB35C73FBB" box="[885,942,433,457]" gridcol="2" gridrow="5" pageId="22" pageNumber="423">124.2</td>
<td id="76A95DE0FF9A22AAFC4F82CB358C3FBB" box="[965,997,433,457]" gridcol="3" gridrow="5" pageId="22" pageNumber="423">0.8</td>
<td id="76A95DE0FF9A22AAFBC482CB334A3FBB" box="[1102,1315,433,457]" colspan="2" colspanRight="1" gridcol="4" gridrow="5" pageId="22" pageNumber="423">123.9–124.4 124.2</td>
<td id="76A95DE0FF9A22AAFAD882CB331B3FBB" box="[1362,1394,433,457]" gridcol="6" gridrow="5" pageId="22" pageNumber="423">0.3</td>
</tr>
<tr id="3578349CFF9A22AAFF1382A3331B3F80" box="[153,1394,473,498]" gridrow="6" pageId="22" pageNumber="423">
<th id="76A95DE0FF9A22AAFF1382A334CF3F80" box="[153,678,473,498]" gridcol="0" gridrow="6" pageId="22" pageNumber="423">Body depth at pelvic-fin origin</th>
<td id="76A95DE0FF9A22AAFD5382A3353E3F80" box="[729,855,473,498]" gridcol="1" gridrow="6" pageId="22" pageNumber="423">32.5–35.3</td>
<td id="76A95DE0FF9A22AAFCFF82A335C73F80" box="[885,942,473,498]" gridcol="2" gridrow="6" pageId="22" pageNumber="423">33.6</td>
<td id="76A95DE0FF9A22AAFC4F82A3358C3F80" box="[965,997,473,498]" gridcol="3" gridrow="6" pageId="22" pageNumber="423">1.5</td>
<td id="76A95DE0FF9A22AAFBC482A332DA3F80" box="[1102,1203,473,498]" gridcol="4" gridrow="6" pageId="22" pageNumber="423">31.7–33.1</td>
<td id="76A95DE0FF9A22AAFB4782A3334A3F80" box="[1229,1315,473,498]" gridcol="5" gridrow="6" pageId="22" pageNumber="423">32.4</td>
<td id="76A95DE0FF9A22AAFAD882A3331B3F80" box="[1362,1394,473,498]" gridcol="6" gridrow="6" pageId="22" pageNumber="423">1.0</td>
</tr>
<tr id="3578349CFF9A22AAFF138178331B3C68" box="[153,1394,514,538]" gridrow="7" pageId="22" pageNumber="423">
<th id="76A95DE0FF9A22AAFF13817834CF3C68" box="[153,678,514,538]" gridcol="0" gridrow="7" pageId="22" pageNumber="423">Head length</th>
<td id="76A95DE0FF9A22AAFD538178353E3C68" box="[729,855,514,538]" gridcol="1" gridrow="7" pageId="22" pageNumber="423">30.4–33.6</td>
<td id="76A95DE0FF9A22AAFCFF817835C73C68" box="[885,942,514,538]" gridcol="2" gridrow="7" pageId="22" pageNumber="423">32.5</td>
<td id="76A95DE0FF9A22AAFC4F8178358C3C68" box="[965,997,514,538]" gridcol="3" gridrow="7" pageId="22" pageNumber="423">1.9</td>
<td id="76A95DE0FF9A22AAFBC4817832DA3C68" box="[1102,1203,514,538]" gridcol="4" gridrow="7" pageId="22" pageNumber="423">33.2–37.2</td>
<td id="76A95DE0FF9A22AAFB478178334A3C68" box="[1229,1315,514,538]" gridcol="5" gridrow="7" pageId="22" pageNumber="423">35.2</td>
<td id="76A95DE0FF9A22AAFAD88178331B3C68" box="[1362,1394,514,538]" gridcol="6" gridrow="7" pageId="22" pageNumber="423">2.8</td>
</tr>
<tr id="3578349CFF9A22AAFF138150331B3C30" box="[153,1394,554,578]" gridrow="8" pageId="22" pageNumber="423">
<th id="76A95DE0FF9A22AAFF13815034CF3C30" box="[153,678,554,578]" gridcol="0" gridrow="8" pageId="22" pageNumber="423">Preanal length</th>
<td id="76A95DE0FF9A22AAFD538150353E3C30" box="[729,855,554,578]" gridcol="1" gridrow="8" pageId="22" pageNumber="423">52.3–55.5</td>
<td id="76A95DE0FF9A22AAFCFF815035C73C30" box="[885,942,554,578]" gridcol="2" gridrow="8" pageId="22" pageNumber="423">53.6</td>
<td id="76A95DE0FF9A22AAFC4F8150358C3C30" box="[965,997,554,578]" gridcol="3" gridrow="8" pageId="22" pageNumber="423">1.7</td>
<td id="76A95DE0FF9A22AAFBC4815032DA3C30" box="[1102,1203,554,578]" gridcol="4" gridrow="8" pageId="22" pageNumber="423">62.2–66.9</td>
<td id="76A95DE0FF9A22AAFB478150334A3C30" box="[1229,1315,554,578]" gridcol="5" gridrow="8" pageId="22" pageNumber="423">64.6</td>
<td id="76A95DE0FF9A22AAFAD88150331B3C30" box="[1362,1394,554,578]" gridcol="6" gridrow="8" pageId="22" pageNumber="423">3.4</td>
</tr>
<tr id="3578349CFF9A22AAFF138128331B3C18" box="[153,1394,594,618]" gridrow="9" pageId="22" pageNumber="423">
<th id="76A95DE0FF9A22AAFF13812834CF3C18" box="[153,678,594,618]" gridcol="0" gridrow="9" pageId="22" pageNumber="423">Predorsal length</th>
<td id="76A95DE0FF9A22AAFD538128353E3C18" box="[729,855,594,618]" gridcol="1" gridrow="9" pageId="22" pageNumber="423">53.0–55.5</td>
<td id="76A95DE0FF9A22AAFCFF812835C73C18" box="[885,942,594,618]" gridcol="2" gridrow="9" pageId="22" pageNumber="423">53.8</td>
<td id="76A95DE0FF9A22AAFC4F8128358C3C18" box="[965,997,594,618]" gridcol="3" gridrow="9" pageId="22" pageNumber="423">1.5</td>
<td id="76A95DE0FF9A22AAFBC4812832DA3C18" box="[1102,1203,594,618]" gridcol="4" gridrow="9" pageId="22" pageNumber="423">61.8–66.1</td>
<td id="76A95DE0FF9A22AAFB478128334A3C18" box="[1229,1315,594,618]" gridcol="5" gridrow="9" pageId="22" pageNumber="423">63.9</td>
<td id="76A95DE0FF9A22AAFAD88128331B3C18" box="[1362,1394,594,618]" gridcol="6" gridrow="9" pageId="22" pageNumber="423">3.1</td>
</tr>
<tr id="3578349CFF9A22AAFF138100331B3CE1" box="[153,1394,634,659]" gridrow="10" pageId="22" pageNumber="423">
<th id="76A95DE0FF9A22AAFF13810034CF3CE1" box="[153,678,634,659]" gridcol="0" gridrow="10" pageId="22" pageNumber="423">Prepelvic length</th>
<td id="76A95DE0FF9A22AAFD538100353E3CE1" box="[729,855,634,659]" gridcol="1" gridrow="10" pageId="22" pageNumber="423">42.7–43.8</td>
<td id="76A95DE0FF9A22AAFCFF810035C73CE1" box="[885,942,634,659]" gridcol="2" gridrow="10" pageId="22" pageNumber="423">43.1</td>
<td id="76A95DE0FF9A22AAFC4F8100358C3CE1" box="[965,997,634,659]" gridcol="3" gridrow="10" pageId="22" pageNumber="423">0.6</td>
<td id="76A95DE0FF9A22AAFBC4810032DA3CE1" box="[1102,1203,634,659]" gridcol="4" gridrow="10" pageId="22" pageNumber="423">41.3–43.2</td>
<td id="76A95DE0FF9A22AAFB478100334A3CE1" box="[1229,1315,634,659]" gridcol="5" gridrow="10" pageId="22" pageNumber="423">42.3</td>
<td id="76A95DE0FF9A22AAFAD88100331B3CE1" box="[1362,1394,634,659]" gridcol="6" gridrow="10" pageId="22" pageNumber="423">1.4</td>
</tr>
<tr id="3578349CFF9A22AAFF1381D9331B3CC9" box="[153,1394,675,699]" gridrow="11" pageId="22" pageNumber="423">
<th id="76A95DE0FF9A22AAFF1381D934CF3CC9" box="[153,678,675,699]" gridcol="0" gridrow="11" pageId="22" pageNumber="423">Prepectoral length</th>
<td id="76A95DE0FF9A22AAFD5381D9353E3CC9" box="[729,855,675,699]" gridcol="1" gridrow="11" pageId="22" pageNumber="423">30.4–33.6</td>
<td id="76A95DE0FF9A22AAFCFF81D935C73CC9" box="[885,942,675,699]" gridcol="2" gridrow="11" pageId="22" pageNumber="423">32.5</td>
<td id="76A95DE0FF9A22AAFC4F81D9358C3CC9" box="[965,997,675,699]" gridcol="3" gridrow="11" pageId="22" pageNumber="423">1.9</td>
<td id="76A95DE0FF9A22AAFBC481D932DA3CC9" box="[1102,1203,675,699]" gridcol="4" gridrow="11" pageId="22" pageNumber="423">33.2–37.2</td>
<td id="76A95DE0FF9A22AAFB4781D9334A3CC9" box="[1229,1315,675,699]" gridcol="5" gridrow="11" pageId="22" pageNumber="423">35.2</td>
<td id="76A95DE0FF9A22AAFAD881D9331B3CC9" box="[1362,1394,675,699]" gridcol="6" gridrow="11" pageId="22" pageNumber="423">2.8</td>
</tr>
<tr id="3578349CFF9A22AAFF1381B1331B3C91" box="[153,1394,715,739]" gridrow="12" pageId="22" pageNumber="423">
<th id="76A95DE0FF9A22AAFF1381B134CF3C91" box="[153,678,715,739]" gridcol="0" gridrow="12" pageId="22" pageNumber="423">Caudal peduncle length</th>
<td id="76A95DE0FF9A22AAFD5381B1353E3C91" box="[729,855,715,739]" gridcol="1" gridrow="12" pageId="22" pageNumber="423">20.0–21.7</td>
<td id="76A95DE0FF9A22AAFCFF81B135C73C91" box="[885,942,715,739]" gridcol="2" gridrow="12" pageId="22" pageNumber="423">21.1</td>
<td id="76A95DE0FF9A22AAFC4F81B1358C3C91" box="[965,997,715,739]" gridcol="3" gridrow="12" pageId="22" pageNumber="423">0.9</td>
<td id="76A95DE0FF9A22AAFBC481B132DA3C91" box="[1102,1203,715,739]" gridcol="4" gridrow="12" pageId="22" pageNumber="423">20.5–24.8</td>
<td id="76A95DE0FF9A22AAFB4781B1334A3C91" box="[1229,1315,715,739]" gridcol="5" gridrow="12" pageId="22" pageNumber="423">22.6</td>
<td id="76A95DE0FF9A22AAFAD881B1331B3C91" box="[1362,1394,715,739]" gridcol="6" gridrow="12" pageId="22" pageNumber="423">3.1</td>
</tr>
<tr id="3578349CFF9A22AAFF138189331B3D79" box="[153,1394,755,779]" gridrow="13" pageId="22" pageNumber="423">
<th id="76A95DE0FF9A22AAFF13818934CF3D79" box="[153,678,755,779]" gridcol="0" gridrow="13" pageId="22" pageNumber="423">Caudal peduncle depth</th>
<td id="76A95DE0FF9A22AAFD538189353E3D79" box="[729,855,755,779]" gridcol="1" gridrow="13" pageId="22" pageNumber="423">13.1–14.8</td>
<td id="76A95DE0FF9A22AAFCFF818935C73D79" box="[885,942,755,779]" gridcol="2" gridrow="13" pageId="22" pageNumber="423">14.0</td>
<td id="76A95DE0FF9A22AAFC4F8189358C3D79" box="[965,997,755,779]" gridcol="3" gridrow="13" pageId="22" pageNumber="423">0.8</td>
<td id="76A95DE0FF9A22AAFBC4818932DA3D79" box="[1102,1203,755,779]" gridcol="4" gridrow="13" pageId="22" pageNumber="423">13.1–16.5</td>
<td id="76A95DE0FF9A22AAFB478189334A3D79" box="[1229,1315,755,779]" gridcol="5" gridrow="13" pageId="22" pageNumber="423">14.8</td>
<td id="76A95DE0FF9A22AAFAD88189331B3D79" box="[1362,1394,755,779]" gridcol="6" gridrow="13" pageId="22" pageNumber="423">2.4</td>
</tr>
<tr id="3578349CFF9A22AAFF138061331B3D46" box="[153,1394,795,820]" gridrow="14" pageId="22" pageNumber="423">
<th id="76A95DE0FF9A22AAFF13806134CF3D46" box="[153,678,795,820]" gridcol="0" gridrow="14" pageId="22" pageNumber="423">Dorsal-fin base length</th>
<td id="76A95DE0FF9A22AAFD538061353E3D46" box="[729,855,795,820]" gridcol="1" gridrow="14" pageId="22" pageNumber="423">26.2–29.6</td>
<td id="76A95DE0FF9A22AAFCFF806135C73D46" box="[885,942,795,820]" gridcol="2" gridrow="14" pageId="22" pageNumber="423">27.9</td>
<td id="76A95DE0FF9A22AAFC4F8061358C3D46" box="[965,997,795,820]" gridcol="3" gridrow="14" pageId="22" pageNumber="423">1.7</td>
<td id="76A95DE0FF9A22AAFBC4806132DA3D46" box="[1102,1203,795,820]" gridcol="4" gridrow="14" pageId="22" pageNumber="423">23.9–27.7</td>
<td id="76A95DE0FF9A22AAFB478061334A3D46" box="[1229,1315,795,820]" gridcol="5" gridrow="14" pageId="22" pageNumber="423">25.8</td>
<td id="76A95DE0FF9A22AAFAD88061331B3D46" box="[1362,1394,795,820]" gridcol="6" gridrow="14" pageId="22" pageNumber="423">2.7</td>
</tr>
<tr id="3578349CFF9A22AAFF138039331B3D2E" box="[153,1394,835,860]" gridrow="15" pageId="22" pageNumber="423">
<th id="76A95DE0FF9A22AAFF13803934CF3D2E" box="[153,678,835,860]" gridcol="0" gridrow="15" pageId="22" pageNumber="423">Anal-fin base length</th>
<td id="76A95DE0FF9A22AAFD538039353E3D2E" box="[729,855,835,860]" gridcol="1" gridrow="15" pageId="22" pageNumber="423">24.3–26.7</td>
<td id="76A95DE0FF9A22AAFCFF803935C73D2E" box="[885,942,835,860]" gridcol="2" gridrow="15" pageId="22" pageNumber="423">25.8</td>
<td id="76A95DE0FF9A22AAFC4F8039358C3D2E" box="[965,997,835,860]" gridcol="3" gridrow="15" pageId="22" pageNumber="423">1.3</td>
<td id="76A95DE0FF9A22AAFBC4803932DA3D2E" box="[1102,1203,835,860]" gridcol="4" gridrow="15" pageId="22" pageNumber="423">18.5–21.9</td>
<td id="76A95DE0FF9A22AAFB478039334A3D2E" box="[1229,1315,835,860]" gridcol="5" gridrow="15" pageId="22" pageNumber="423">20.2</td>
<td id="76A95DE0FF9A22AAFAD88039331B3D2E" box="[1362,1394,835,860]" gridcol="6" gridrow="15" pageId="22" pageNumber="423">2.4</td>
</tr>
<tr id="3578349CFF9A22AAFF138016331B3DF6" box="[153,1394,876,900]" gridrow="16" pageId="22" pageNumber="423">
<th id="76A95DE0FF9A22AAFF13801634CF3DF6" box="[153,678,876,900]" gridcol="0" gridrow="16" pageId="22" pageNumber="423">Caudal-fin length</th>
<td id="76A95DE0FF9A22AAFD538016353E3DF6" box="[729,855,876,900]" gridcol="1" gridrow="16" pageId="22" pageNumber="423">23.3–24.8</td>
<td id="76A95DE0FF9A22AAFCFF801635C73DF6" box="[885,942,876,900]" gridcol="2" gridrow="16" pageId="22" pageNumber="423">24.2</td>
<td id="76A95DE0FF9A22AAFC4F8016358C3DF6" box="[965,997,876,900]" gridcol="3" gridrow="16" pageId="22" pageNumber="423">0.8</td>
<td id="76A95DE0FF9A22AAFBC4801632DA3DF6" box="[1102,1203,876,900]" gridcol="4" gridrow="16" pageId="22" pageNumber="423">23.9–24.4</td>
<td id="76A95DE0FF9A22AAFB478016334A3DF6" box="[1229,1315,876,900]" gridcol="5" gridrow="16" pageId="22" pageNumber="423">24.2</td>
<td id="76A95DE0FF9A22AAFAD88016331B3DF6" box="[1362,1394,876,900]" gridcol="6" gridrow="16" pageId="22" pageNumber="423">0.3</td>
</tr>
<tr id="3578349CFF9A22AAFF1380EE331B3DDE" box="[153,1394,916,940]" gridrow="17" pageId="22" pageNumber="423">
<th id="76A95DE0FF9A22AAFF1380EE331B3DDE" box="[153,1394,916,940]" colspan="7" colspanRight="6" gridcol="0" gridrow="17" pageId="22" pageNumber="423">
<emphasis id="B93CEACCFF9ADD43FF1380EE37E23DDE" bold="true" box="[153,395,916,940]" pageId="22" pageNumber="423">Percent of head length</emphasis>
</th>
</tr>
<tr id="3578349CFF9A22AAFF1380C6331B3DA6" box="[153,1394,956,980]" gridrow="18" pageId="22" pageNumber="423">
<th id="76A95DE0FF9A22AAFF1380C634CF3DA6" box="[153,678,956,980]" gridcol="0" gridrow="18" pageId="22" pageNumber="423">Head width</th>
<td id="76A95DE0FF9A22AAFD5380C6353E3DA6" box="[729,855,956,980]" gridcol="1" gridrow="18" pageId="22" pageNumber="423">55–58</td>
<td id="76A95DE0FF9A22AAFCFF80C635C73DA6" box="[885,942,956,980]" gridcol="2" gridrow="18" pageId="22" pageNumber="423">56.9</td>
<td id="76A95DE0FF9A22AAFC4F80C6358C3DA6" box="[965,997,956,980]" gridcol="3" gridrow="18" pageId="22" pageNumber="423">1.8</td>
<td id="76A95DE0FF9A22AAFBC480C632DA3DA6" box="[1102,1203,956,980]" gridcol="4" gridrow="18" pageId="22" pageNumber="423">59–63</td>
<td id="76A95DE0FF9A22AAFB4780C6334A3DA6" box="[1229,1315,956,980]" gridcol="5" gridrow="18" pageId="22" pageNumber="423">60.8</td>
<td id="76A95DE0FF9A22AAFAD880C6331B3DA6" box="[1362,1394,956,980]" gridcol="6" gridrow="18" pageId="22" pageNumber="423">2.8</td>
</tr>
<tr id="3578349CFF9A22AAFF13809E331B3D8F" box="[153,1394,996,1021]" gridrow="19" pageId="22" pageNumber="423">
<th id="76A95DE0FF9A22AAFF13809E34CF3D8F" box="[153,678,996,1021]" gridcol="0" gridrow="19" pageId="22" pageNumber="423">Head depth</th>
<td id="76A95DE0FF9A22AAFD53809E353E3D8F" box="[729,855,996,1021]" gridcol="1" gridrow="19" pageId="22" pageNumber="423">85–86</td>
<td id="76A95DE0FF9A22AAFCFF809E35C73D8F" box="[885,942,996,1021]" gridcol="2" gridrow="19" pageId="22" pageNumber="423">85.8</td>
<td id="76A95DE0FF9A22AAFC4F809E358C3D8F" box="[965,997,996,1021]" gridcol="3" gridrow="19" pageId="22" pageNumber="423">1.1</td>
<td id="76A95DE0FF9A22AAFBC4809E32DA3D8F" box="[1102,1203,996,1021]" gridcol="4" gridrow="19" pageId="22" pageNumber="423">80–81</td>
<td id="76A95DE0FF9A22AAFB47809E334A3D8F" box="[1229,1315,996,1021]" gridcol="5" gridrow="19" pageId="22" pageNumber="423">80.7</td>
<td id="76A95DE0FF9A22AAFAD8809E331B3D8F" box="[1362,1394,996,1021]" gridcol="6" gridrow="19" pageId="22" pageNumber="423">1.0</td>
</tr>
<tr id="3578349CFF9A22AAFF138777331B3A57" box="[153,1394,1037,1061]" gridrow="20" pageId="22" pageNumber="423">
<th id="76A95DE0FF9A22AAFF13877734CF3A57" box="[153,678,1037,1061]" gridcol="0" gridrow="20" pageId="22" pageNumber="423">Interorbital width</th>
<td id="76A95DE0FF9A22AAFD538777353E3A57" box="[729,855,1037,1061]" gridcol="1" gridrow="20" pageId="22" pageNumber="423">45–49</td>
<td id="76A95DE0FF9A22AAFCFF877735C73A57" box="[885,942,1037,1061]" gridcol="2" gridrow="20" pageId="22" pageNumber="423">46.7</td>
<td id="76A95DE0FF9A22AAFC4F8777358C3A57" box="[965,997,1037,1061]" gridcol="3" gridrow="20" pageId="22" pageNumber="423">2.3</td>
<td id="76A95DE0FF9A22AAFBC4877732DA3A57" box="[1102,1203,1037,1061]" gridcol="4" gridrow="20" pageId="22" pageNumber="423">45–48</td>
<td id="76A95DE0FF9A22AAFB478777334A3A57" box="[1229,1315,1037,1061]" gridcol="5" gridrow="20" pageId="22" pageNumber="423">46.6</td>
<td id="76A95DE0FF9A22AAFAD88777331B3A57" box="[1362,1394,1037,1061]" gridcol="6" gridrow="20" pageId="22" pageNumber="423">1.7</td>
</tr>
<tr id="3578349CFF9A22AAFF13874F331B3A3F" box="[153,1394,1077,1101]" gridrow="21" pageId="22" pageNumber="423">
<th id="76A95DE0FF9A22AAFF13874F34CF3A3F" box="[153,678,1077,1101]" gridcol="0" gridrow="21" pageId="22" pageNumber="423">Postorbital length</th>
<td id="76A95DE0FF9A22AAFD53874F353E3A3F" box="[729,855,1077,1101]" gridcol="1" gridrow="21" pageId="22" pageNumber="423">50–53</td>
<td id="76A95DE0FF9A22AAFCFF874F35C73A3F" box="[885,942,1077,1101]" gridcol="2" gridrow="21" pageId="22" pageNumber="423">50.9</td>
<td id="76A95DE0FF9A22AAFC4F874F358C3A3F" box="[965,997,1077,1101]" gridcol="3" gridrow="21" pageId="22" pageNumber="423">1.6</td>
<td id="76A95DE0FF9A22AAFBC4874F32DA3A3F" box="[1102,1203,1077,1101]" gridcol="4" gridrow="21" pageId="22" pageNumber="423">51–52</td>
<td id="76A95DE0FF9A22AAFB47874F334A3A3F" box="[1229,1315,1077,1101]" gridcol="5" gridrow="21" pageId="22" pageNumber="423">51.7</td>
<td id="76A95DE0FF9A22AAFAD8874F331B3A3F" box="[1362,1394,1077,1101]" gridcol="6" gridrow="21" pageId="22" pageNumber="423">0.7</td>
</tr>
<tr id="3578349CFF9A22AAFF138727331B3A07" box="[153,1394,1117,1141]" gridrow="22" pageId="22" pageNumber="423">
<th id="76A95DE0FF9A22AAFF13872734CF3A07" box="[153,678,1117,1141]" gridcol="0" gridrow="22" pageId="22" pageNumber="423">Suborbital depth</th>
<td id="76A95DE0FF9A22AAFD538727353E3A07" box="[729,855,1117,1141]" gridcol="1" gridrow="22" pageId="22" pageNumber="423">12–16</td>
<td id="76A95DE0FF9A22AAFCFF872735C73A07" box="[885,942,1117,1141]" gridcol="2" gridrow="22" pageId="22" pageNumber="423">14.4</td>
<td id="76A95DE0FF9A22AAFC4F8727358C3A07" box="[965,997,1117,1141]" gridcol="3" gridrow="22" pageId="22" pageNumber="423">2.5</td>
<td id="76A95DE0FF9A22AAFBC4872732DA3A07" box="[1102,1203,1117,1141]" gridcol="4" gridrow="22" pageId="22" pageNumber="423">9–12</td>
<td id="76A95DE0FF9A22AAFB478727334A3A07" box="[1229,1315,1117,1141]" gridcol="5" gridrow="22" pageId="22" pageNumber="423">10.8</td>
<td id="76A95DE0FF9A22AAFAD88727331B3A07" box="[1362,1394,1117,1141]" gridcol="6" gridrow="22" pageId="22" pageNumber="423">2.1</td>
</tr>
<tr id="3578349CFF9A22AAFF1387FF331B3AEC" box="[153,1394,1157,1182]" gridrow="23" pageId="22" pageNumber="423">
<th id="76A95DE0FF9A22AAFF1387FF34CF3AEC" box="[153,678,1157,1182]" gridcol="0" gridrow="23" pageId="22" pageNumber="423">Eye diameter</th>
<td id="76A95DE0FF9A22AAFD5387FF353E3AEC" box="[729,855,1157,1182]" gridcol="1" gridrow="23" pageId="22" pageNumber="423">33–36</td>
<td id="76A95DE0FF9A22AAFCFF87FF35C73AEC" box="[885,942,1157,1182]" gridcol="2" gridrow="23" pageId="22" pageNumber="423">34.0</td>
<td id="76A95DE0FF9A22AAFC4F87FF358C3AEC" box="[965,997,1157,1182]" gridcol="3" gridrow="23" pageId="22" pageNumber="423">1.4</td>
<td id="76A95DE0FF9A22AAFBC487FF32DA3AEC" box="[1102,1203,1157,1182]" gridcol="4" gridrow="23" pageId="22" pageNumber="423">32–36</td>
<td id="76A95DE0FF9A22AAFB4787FF334A3AEC" box="[1229,1315,1157,1182]" gridcol="5" gridrow="23" pageId="22" pageNumber="423">34.1</td>
<td id="76A95DE0FF9A22AAFAD887FF331B3AEC" box="[1362,1394,1157,1182]" gridcol="6" gridrow="23" pageId="22" pageNumber="423">2.7</td>
</tr>
<tr id="3578349CFF9A22AAFF1387D4331B3AB4" box="[153,1394,1198,1222]" gridrow="24" pageId="22" pageNumber="423">
<th id="76A95DE0FF9A22AAFF1387D434CF3AB4" box="[153,678,1198,1222]" gridcol="0" gridrow="24" pageId="22" pageNumber="423">Snout to eye end length</th>
<td id="76A95DE0FF9A22AAFD5387D4353E3AB4" box="[729,855,1198,1222]" gridcol="1" gridrow="24" pageId="22" pageNumber="423">47–50</td>
<td id="76A95DE0FF9A22AAFCFF87D435C73AB4" box="[885,942,1198,1222]" gridcol="2" gridrow="24" pageId="22" pageNumber="423">48.8</td>
<td id="76A95DE0FF9A22AAFC4F87D4358C3AB4" box="[965,997,1198,1222]" gridcol="3" gridrow="24" pageId="22" pageNumber="423">1.4</td>
<td id="76A95DE0FF9A22AAFBC487D432DA3AB4" box="[1102,1203,1198,1222]" gridcol="4" gridrow="24" pageId="22" pageNumber="423">48–49</td>
<td id="76A95DE0FF9A22AAFB4787D4334A3AB4" box="[1229,1315,1198,1222]" gridcol="5" gridrow="24" pageId="22" pageNumber="423">48.3</td>
<td id="76A95DE0FF9A22AAFAD887D4331B3AB4" box="[1362,1394,1198,1222]" gridcol="6" gridrow="24" pageId="22" pageNumber="423">0.9</td>
</tr>
<tr id="3578349CFF9A22AAFF1387AC331B3A9C" box="[153,1394,1238,1262]" gridrow="25" pageId="22" pageNumber="423">
<th id="76A95DE0FF9A22AAFF1387AC34CF3A9C" box="[153,678,1238,1262]" gridcol="0" gridrow="25" pageId="22" pageNumber="423">Snout length</th>
<td id="76A95DE0FF9A22AAFD5387AC353E3A9C" box="[729,855,1238,1262]" gridcol="1" gridrow="25" pageId="22" pageNumber="423">14–18</td>
<td id="76A95DE0FF9A22AAFCFF87AC35C73A9C" box="[885,942,1238,1262]" gridcol="2" gridrow="25" pageId="22" pageNumber="423">16.1</td>
<td id="76A95DE0FF9A22AAFC4F87AC358C3A9C" box="[965,997,1238,1262]" gridcol="3" gridrow="25" pageId="22" pageNumber="423">2.2</td>
<td id="76A95DE0FF9A22AAFBC487AC32DA3A9C" box="[1102,1203,1238,1262]" gridcol="4" gridrow="25" pageId="22" pageNumber="423">17</td>
<td id="76A95DE0FF9A22AAFB4787AC334A3A9C" box="[1229,1315,1238,1262]" gridcol="5" gridrow="25" pageId="22" pageNumber="423">17.1</td>
<td id="76A95DE0FF9A22AAFAD887AC331B3A9C" box="[1362,1394,1238,1262]" gridcol="6" gridrow="25" pageId="22" pageNumber="423">0.5</td>
</tr>
<tr id="3578349CFF9A22AAFF138784331B3B64" box="[153,1394,1278,1302]" gridrow="26" pageId="22" pageNumber="423">
<th id="76A95DE0FF9A22AAFF138784331B3B64" box="[153,1394,1278,1302]" colspan="7" colspanRight="6" gridcol="0" gridrow="26" pageId="22" pageNumber="423">
<emphasis id="B93CEACCFF9ADD43FF13878437A93B64" bold="true" box="[153,448,1278,1302]" pageId="22" pageNumber="423">Other morphometric ratios</emphasis>
</th>
</tr>
<tr id="3578349CFF9A22AAFF13865D331B3B4D" box="[153,1394,1319,1343]" gridrow="27" pageId="22" pageNumber="423">
<th id="76A95DE0FF9A22AAFF13865D34CF3B4D" box="[153,678,1319,1343]" gridcol="0" gridrow="27" pageId="22" pageNumber="423">Predorsal length in % of preanal length</th>
<td id="76A95DE0FF9A22AAFD53865D353E3B4D" box="[729,855,1319,1343]" gridcol="1" gridrow="27" pageId="22" pageNumber="423">100–101</td>
<td id="76A95DE0FF9A22AAFCFF865D35C73B4D" box="[885,942,1319,1343]" gridcol="2" gridrow="27" pageId="22" pageNumber="423">100.4</td>
<td id="76A95DE0FF9A22AAFC4F865D358C3B4D" box="[965,997,1319,1343]" gridcol="3" gridrow="27" pageId="22" pageNumber="423">0.8</td>
<td id="76A95DE0FF9A22AAFBC4865D32DA3B4D" box="[1102,1203,1319,1343]" gridcol="4" gridrow="27" pageId="22" pageNumber="423">99</td>
<td id="76A95DE0FF9A22AAFB47865D334A3B4D" box="[1229,1315,1319,1343]" gridcol="5" gridrow="27" pageId="22" pageNumber="423">99.1</td>
<td id="76A95DE0FF9A22AAFAD8865D331B3B4D" box="[1362,1394,1319,1343]" gridcol="6" gridrow="27" pageId="22" pageNumber="423">0.4</td>
</tr>
<tr id="3578349CFF9A22AAFF138635331B3B15" box="[153,1394,1359,1383]" gridrow="28" pageId="22" pageNumber="423">
<th id="76A95DE0FF9A22AAFF13863534CF3B15" box="[153,678,1359,1383]" gridcol="0" gridrow="28" pageId="22" pageNumber="423">Caudal peduncle length in % of its depth</th>
<td id="76A95DE0FF9A22AAFD538635353E3B15" box="[729,855,1359,1383]" gridcol="1" gridrow="28" pageId="22" pageNumber="423">146–153</td>
<td id="76A95DE0FF9A22AAFCFF863535C73B15" box="[885,942,1359,1383]" gridcol="2" gridrow="28" pageId="22" pageNumber="423">150.6</td>
<td id="76A95DE0FF9A22AAFC4F8635358C3B15" box="[965,997,1359,1383]" gridcol="3" gridrow="28" pageId="22" pageNumber="423">3.8</td>
<td id="76A95DE0FF9A22AAFBC4863532DA3B15" box="[1102,1203,1359,1383]" gridcol="4" gridrow="28" pageId="22" pageNumber="423">150–156</td>
<td id="76A95DE0FF9A22AAFB478635334A3B15" box="[1229,1315,1359,1383]" gridcol="5" gridrow="28" pageId="22" pageNumber="423">152.9</td>
<td id="76A95DE0FF9A22AAFAD88635331B3B15" box="[1362,1394,1359,1383]" gridcol="6" gridrow="28" pageId="22" pageNumber="423">4.2</td>
</tr>
<tr id="3578349CFF9A22AAFF13860D331B3BFD" box="[153,1394,1399,1423]" gridrow="29" pageId="22" pageNumber="423">
<th id="76A95DE0FF9A22AAFF13860D34CF3BFD" box="[153,678,1399,1423]" gridcol="0" gridrow="29" pageId="22" pageNumber="423">Head width in % of its depth</th>
<td id="76A95DE0FF9A22AAFD53860D353E3BFD" box="[729,855,1399,1423]" gridcol="1" gridrow="29" pageId="22" pageNumber="423">64–69</td>
<td id="76A95DE0FF9A22AAFCFF860D35C73BFD" box="[885,942,1399,1423]" gridcol="2" gridrow="29" pageId="22" pageNumber="423">66.3</td>
<td id="76A95DE0FF9A22AAFC4F860D358C3BFD" box="[965,997,1399,1423]" gridcol="3" gridrow="29" pageId="22" pageNumber="423">2.7</td>
<td id="76A95DE0FF9A22AAFBC4860D32DA3BFD" box="[1102,1203,1399,1423]" gridcol="4" gridrow="29" pageId="22" pageNumber="423">74–77</td>
<td id="76A95DE0FF9A22AAFB47860D334A3BFD" box="[1229,1315,1399,1423]" gridcol="5" gridrow="29" pageId="22" pageNumber="423">75.4</td>
<td id="76A95DE0FF9A22AAFAD8860D331B3BFD" box="[1362,1394,1399,1423]" gridcol="6" gridrow="29" pageId="22" pageNumber="423">2.5</td>
</tr>
<tr id="3578349CFF9A22AAFF1386E5331B3BCA" box="[153,1394,1439,1464]" gridrow="30" pageId="22" pageNumber="423" rowspan-3="1" rowspan-6="1">
<th id="76A95DE0FF9A22AAFF1386E534CF3BCA" box="[153,678,1439,1464]" gridcol="0" gridrow="30" pageId="22" pageNumber="423">
<emphasis id="B93CEACCFF9ADD43FF1386E536943BC5" bold="true" box="[153,253,1439,1463]" pageId="22" pageNumber="423">Meristics</emphasis>
</th>
<td id="76A95DE0FF9A22AAFD5386E5353E3BCA" box="[729,855,1439,1464]" gridcol="1" gridrow="30" pageId="22" pageNumber="423">range</td>
<td id="76A95DE0FF9A22AAFCFF86E535C73BCA" box="[885,942,1439,1464]" gridcol="2" gridrow="30" pageId="22" pageNumber="423">mode</td>
<td id="76A95DE0FF9A22AAFBC486E532DA3BCA" box="[1102,1203,1439,1464]" gridcol="4" gridrow="30" pageId="22" pageNumber="423">min–max</td>
<td id="76A95DE0FF9A22AAFB4786E5334A3BCA" box="[1229,1315,1439,1464]" gridcol="5" gridrow="30" pageId="22" pageNumber="423">mode</td>
</tr>
<tr id="3578349CFF9A22AAFF1386BD331B3B92" box="[153,1394,1479,1504]" gridrow="31" pageId="22" pageNumber="423" rowspan-3="1" rowspan-6="1">
<th id="76A95DE0FF9A22AAFF1386BD34CF3B92" box="[153,678,1479,1504]" gridcol="0" gridrow="31" pageId="22" pageNumber="423">Dorsal-fin rays</th>
<td id="76A95DE0FF9A22AAFD5386BD353E3B92" box="[729,855,1479,1504]" gridcol="1" gridrow="31" pageId="22" pageNumber="423">15–16</td>
<td id="76A95DE0FF9A22AAFCFF86BD35C73B92" box="[885,942,1479,1504]" gridcol="2" gridrow="31" pageId="22" pageNumber="423">16</td>
<td id="76A95DE0FF9A22AAFBC486BD32DA3B92" box="[1102,1203,1479,1504]" gridcol="4" gridrow="31" pageId="22" pageNumber="423">15</td>
<td id="76A95DE0FF9A22AAFB4786BD334A3B92" box="[1229,1315,1479,1504]" gridcol="5" gridrow="31" pageId="22" pageNumber="423">15</td>
</tr>
<tr id="3578349CFF9A22AAFF13868A331B387A" box="[153,1394,1520,1544]" gridrow="32" pageId="22" pageNumber="423" rowspan-3="1" rowspan-6="1">
<th id="76A95DE0FF9A22AAFF13868A34CF387A" box="[153,678,1520,1544]" gridcol="0" gridrow="32" pageId="22" pageNumber="423">Anal-fin rays</th>
<td id="76A95DE0FF9A22AAFD53868A353E387A" box="[729,855,1520,1544]" gridcol="1" gridrow="32" pageId="22" pageNumber="423">17–18</td>
<td id="76A95DE0FF9A22AAFCFF868A35C7387A" box="[885,942,1520,1544]" gridcol="2" gridrow="32" pageId="22" pageNumber="423">17</td>
<td id="76A95DE0FF9A22AAFBC4868A32DA387A" box="[1102,1203,1520,1544]" gridcol="4" gridrow="32" pageId="22" pageNumber="423">17</td>
<td id="76A95DE0FF9A22AAFB47868A334A387A" box="[1229,1315,1520,1544]" gridcol="5" gridrow="32" pageId="22" pageNumber="423">17</td>
</tr>
<tr id="3578349CFF9A22AAFF138562331B3842" box="[153,1394,1560,1584]" gridrow="33" pageId="22" pageNumber="423" rowspan-3="1" rowspan-6="1">
<th id="76A95DE0FF9A22AAFF13856234CF3842" box="[153,678,1560,1584]" gridcol="0" gridrow="33" pageId="22" pageNumber="423">Scales mid-longitudinal series</th>
<td id="76A95DE0FF9A22AAFD538562353E3842" box="[729,855,1560,1584]" gridcol="1" gridrow="33" pageId="22" pageNumber="423">25–26</td>
<td id="76A95DE0FF9A22AAFCFF856235C73842" box="[885,942,1560,1584]" gridcol="2" gridrow="33" pageId="22" pageNumber="423">26</td>
<td id="76A95DE0FF9A22AAFBC4856232DA3842" box="[1102,1203,1560,1584]" gridcol="4" gridrow="33" pageId="22" pageNumber="423">25</td>
<td id="76A95DE0FF9A22AAFB478562334A3842" box="[1229,1315,1560,1584]" gridcol="5" gridrow="33" pageId="22" pageNumber="423">25</td>
</tr>
<tr id="3578349CFF9A22AAFF13853A331B382A" box="[153,1394,1600,1624]" gridrow="34" pageId="22" pageNumber="423" rowspan-3="1" rowspan-6="1">
<th id="76A95DE0FF9A22AAFF13853A34CF382A" box="[153,678,1600,1624]" gridcol="0" gridrow="34" pageId="22" pageNumber="423">Scales transverse</th>
<td id="76A95DE0FF9A22AAFD53853A353E382A" box="[729,855,1600,1624]" gridcol="1" gridrow="34" pageId="22" pageNumber="423">10</td>
<td id="76A95DE0FF9A22AAFCFF853A35C7382A" box="[885,942,1600,1624]" gridcol="2" gridrow="34" pageId="22" pageNumber="423">10</td>
<td id="76A95DE0FF9A22AAFBC4853A32DA382A" box="[1102,1203,1600,1624]" gridcol="4" gridrow="34" pageId="22" pageNumber="423">10</td>
<td id="76A95DE0FF9A22AAFB47853A334A382A" box="[1229,1315,1600,1624]" gridcol="5" gridrow="34" pageId="22" pageNumber="423">10</td>
</tr>
<tr id="3578349CFF9A22AAFF138512331B38F3" box="[153,1394,1640,1665]" gridrow="35" pageId="22" pageNumber="423" rowspan-3="1" rowspan-6="1">
<th id="76A95DE0FF9A22AAFF13851234CF38F3" box="[153,678,1640,1665]" gridcol="0" gridrow="35" pageId="22" pageNumber="423">Scales circumpeduncular</th>
<td id="76A95DE0FF9A22AAFD538512353E38F3" box="[729,855,1640,1665]" gridcol="1" gridrow="35" pageId="22" pageNumber="423">12</td>
<td id="76A95DE0FF9A22AAFCFF851235C738F3" box="[885,942,1640,1665]" gridcol="2" gridrow="35" pageId="22" pageNumber="423">12</td>
<td id="76A95DE0FF9A22AAFBC4851232DA38F3" box="[1102,1203,1640,1665]" gridcol="4" gridrow="35" pageId="22" pageNumber="423">12</td>
<td id="76A95DE0FF9A22AAFB478512334A38F3" box="[1229,1315,1640,1665]" gridcol="5" gridrow="35" pageId="22" pageNumber="423">12</td>
</tr>
</table>
</paragraph>
<paragraph id="8BF736DEFF9ADD43FF4D85CA3220390C" blockId="22.[151,1437,1712,2027]" pageId="22" pageNumber="423">Head moderately long (30.4–33.6 % SL), laterally compressed, deeper than wide (head width 64–69 % of its depth). Snout distinctly pointed, smaller than eye diameter. Mouth supraterminal, slightly oblique in profile. Jaws subequal, lower jaw longer than upper, posterior end of rictus at same level or slightly ventral to centre of eye. Premaxilla and dentary with many irregularly distributed conical teeth on inner rows and slightly curved teeth on outer row. Orbit large, eye diameter 33–36 % HL, positioned entirely in anterior half of head (snout to eye end length 47–50 % HL). Branchiostegal membrane projecting posteriorly from opercle.</paragraph>
<paragraph id="8BF736DEFF9ADD42FF4D84F2350F3F13" blockId="22.[151,1437,1712,2027]" lastBlockId="23.[151,1437,151,953]" lastPageId="23" lastPageNumber="424" pageId="22" pageNumber="423">
Dorsal-fin origin at same vertical positioning as anal-fin origin (mean predorsal length
<quantity id="4CB09B3BFF9ADD43FB0184F2336039D0" box="[1163,1289,1928,1954]" metricMagnitude="0" metricUnit="m" metricValue="2.5527" metricValueMax="2.5654" metricValueMin="2.54" pageId="22" pageNumber="423" unit="in" value="100.5" valueMax="101.0" valueMin="100.0">100–101 in</quantity>
% of preanal length), both fins originating slightly posterior to mid-length of body (predorsal length 53.0–55.5 % SL; preanal length 52.3–55.5 % SL). Dorsal fin with 15–16 rays, extremity rounded. Anal fin trapezoidal in shape, with 17–18 rays; lower margin moderately concave in profile. Dorsal and anal fins with short fin-ray extensions. Pectoral fin subtriangular, insertion slightly posterior to margin of opercular opening, base slightly oblique, upper fin rays positioned slightly anterior to lower fin rays, with tip reaching or slightly overlapping base of pelvic fin. Pelvic fin subabdominal, origin at about mid-length of body, short, base medially separated, with tip reaching urogenital papilla. Caudal fin rounded, with 16–18 branched rays, plus 3–4 dorsal and ventral procurrent rays; central 2–3 rays of fin relatively slightly elongated resulting in mild central lobe.
</paragraph>
<paragraph id="8BF736DEFF9BDD42FF4D821032273FB8" blockId="23.[151,1437,151,953]" pageId="23" pageNumber="424">Scales cycloid, body and head entirely scaled, except for ventral surface of head. No scales on base of dorsal and anal fins. Scales in mid-longitudinal series 25–26 plus two or three small scales on caudal-fin base. Transverse rows of scales in front of dorsal-fin origin 10; scale rows around caudal peduncle 12.</paragraph>
<paragraph id="8BF736DEFF9BDD42FF4D82A937043C96" blockId="23.[151,1437,151,953]" pageId="23" pageNumber="424">
Cephalic squamation pattern G-type. Two frontal neuromasts behind upper lip, posterior in shallow groove. Cephalic sensory system at supraorbital level in divided shallow groove with one and three exposed neuromasts, in anterior and posterior portions, respectively (
<figureCitation id="13732A5BFF9BDD42FD00816034B53C46" box="[650,732,538,564]" captionStart="FIGURE 3" captionStartId="10.[151,250,1307,1331]" captionTargetBox="[156,1428,706,1276]" captionTargetId="figure-280@10.[151,1436,702,1281]" captionTargetPageId="10" captionText="FIGURE 3. Cephalic structure: a, Nothobranchius janpapi; b, N. luekei; c, N. geminus; d, N. lourensi and N. fuscotaeniatus. Key to label abbreviations: an, anterior nostril; pn, posterior nostril; f, frontal neuromast; so, supraorbital canal; st, supratemporal canal." pageId="23" pageNumber="424">Fig. 3d</figureCitation>
), whereas at supratemporal level with four exposed neuromasts in undivided, slightly curved shallow groove. Preorbital canal in shallow groove with three exposed neuromasts; infraorbital level with series of about dozen small buttons at ventral and posterior margin of orbit; postorbital canal in deep groove with one exposed neuromast. Preopercular system with seven exposed neuromasts in deep groove. Mandibular canal in shallow groove with three small neuromasts. One neuromast on each scale along trunk mid-longitudinal series.
</paragraph>
<paragraph id="8BF736DEFF9BDD42FF4D819735E73DCB" blockId="23.[151,1437,151,953]" pageId="23" pageNumber="424">
Female: Smaller than male, maximum recorded size
<quantity id="4CB09B3BFF9BDD42FCA0819735F83D7A" box="[810,913,749,776]" metricMagnitude="-2" metricUnit="m" metricValue="2.59" pageId="23" pageNumber="424" unit="mm" value="25.9">25.9 mm</quantity>
SL. Branchiostegal membrane not projecting posteriorly from opercle. Dorsal and caudal fins rounded. Anal fin subtriangular, tip rounded, central rays longer and more rigid. Dorsal fin positioned more posteriorly than in males (predorsal length 61.8–66.1 % SL vs 53.0–55.5). Anal fin positioned more posteriorly than in males (preanal length 62.2–66.9 % SL vs. 52.3–55.5), and base length smaller than in males (18.5–21.9 % SL vs. 24.3–26.7). Head width larger than in males (59–63 % HL vs. 55–58) whereas head depth smaller than in males (80–81 % HL vs. 85–86).
</paragraph>
<caption id="DF376656FF9BDD42FF1D846032683980" pageId="23" pageNumber="424" startId="23.[151,250,1818,1842]" targetBox="[151,1436,974,1796]" targetPageId="23" targetType="figure">
<paragraph id="8BF736DEFF9BDD42FF1D846032683980" blockId="23.[151,1437,1818,2034]" pageId="23" pageNumber="424">
<emphasis id="B93CEACCFF9BDD42FF1D8460377D3940" bold="true" box="[151,276,1818,1842]" pageId="23" pageNumber="424">FIGURE 7.</emphasis>
Wild-caught specimens of
<taxonomicName id="4C484D5DFF9BDD42FDA6846035013940" authorityName="Seegers" authorityYear="1997" box="[556,872,1818,1842]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="23" pageNumber="424" phylum="Chordata" rank="species" species="fuscotaeniatus">
<emphasis id="B93CEACCFF9BDD42FDA6846035013940" box="[556,872,1818,1842]" italics="true" pageId="23" pageNumber="424">Nothobranchius fuscotaeniatus</emphasis>
</taxonomicName>
and only known locality.
<emphasis id="B93CEACCFF9BDD42FBF9846032E53940" bold="true" box="[1139,1164,1818,1842]" pageId="23" pageNumber="424">A.</emphasis>
<taxonomicName id="4C484D5DFF9BDD42FB19846133F43940" authority="Kitonga" authorityName="Kitonga" box="[1171,1437,1818,1842]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="23" pageNumber="424" phylum="Chordata" rank="species" species="fuscotaeniatus">
<emphasis id="B93CEACCFF9BDD42FB198461332A3940" box="[1171,1347,1818,1842]" italics="true" pageId="23" pageNumber="424">N. fuscotaeniatus</emphasis>
Kitonga
</taxonomicName>
north TAN 97-9, male, Rufiji River drainage;
<emphasis id="B93CEACCFF9BDD42FDE7844034ED3920" bold="true" box="[621,644,1850,1874]" pageId="23" pageNumber="424">B.</emphasis>
<taxonomicName id="4C484D5DFF9BDD42FD01844135FD3920" authority="Kitonga" authorityName="Kitonga" box="[651,916,1850,1874]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="23" pageNumber="424" phylum="Chordata" rank="species" species="fuscotaeniatus">
<emphasis id="B93CEACCFF9BDD42FD01844135523920" box="[651,827,1850,1874]" italics="true" pageId="23" pageNumber="424">N. fuscotaeniatus</emphasis>
Kitonga
</taxonomicName>
north TAN 97-9, female, Rufiji River drainage;
<emphasis id="B93CEACCFF9BDD42FA0E844033F53920" bold="true" box="[1412,1436,1850,1874]" pageId="23" pageNumber="424">C.</emphasis>
<taxonomicName id="4C484D5DFF9BDD42FF1D842137CB3900" authority="Kitonga" authorityName="Kitonga" box="[151,418,1882,1906]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="23" pageNumber="424" phylum="Chordata" rank="species" species="fuscotaeniatus">
<emphasis id="B93CEACCFF9BDD42FF1D842137213900" box="[151,328,1882,1906]" italics="true" pageId="23" pageNumber="424">N. fuscotaeniatus</emphasis>
Kitonga
</taxonomicName>
north TAN 97-9, Rufiji River floodplain, type locality, shallow with dense vegetation (photograph May 31, 1997), hosting
<taxonomicName id="4C484D5DFF9BDD42FE018401345239E0" box="[395,571,1914,1938]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="23" pageNumber="424" phylum="Chordata" rank="species" species="fuscotaeniatus">
<emphasis id="B93CEACCFF9BDD42FE018401345239E0" box="[395,571,1914,1938]" italics="true" pageId="23" pageNumber="424">N. fuscotaeniatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C484D5DFF9BDD42FDC2840134DA39E0" box="[584,691,1915,1938]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="23" pageNumber="424" phylum="Chordata" rank="species" species="janpapi">
<emphasis id="B93CEACCFF9BDD42FDC2840134DA39E0" box="[584,691,1915,1938]" italics="true" pageId="23" pageNumber="424">N. janpapi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C484D5DFF9BDD42FD6F8401353C39E0" box="[741,853,1914,1938]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="23" pageNumber="424" phylum="Chordata" rank="species" species="lourensi">
<emphasis id="B93CEACCFF9BDD42FD6F8401353C39E0" box="[741,853,1914,1938]" italics="true" pageId="23" pageNumber="424">N. lourensi</emphasis>
</taxonomicName>
;
<emphasis id="B93CEACCFF9BDD42FCE88400351239E0" bold="true" box="[866,891,1914,1938]" pageId="23" pageNumber="424">D.</emphasis>
Same site as in “C”, photograph by B. Nagy in 2017. Multiple visits to the site subsequent to the original discovery in 1997 showed an absence of the species, due to substantial changes in the habitat, primarily the loss of dense vegetation and contamination of the habitat by soil washed off the sinceabandoned adjacent road. Photographs by B.R. Watters except where otherwise noted.
</paragraph>
</caption>
<paragraph id="8BF736DEFF94DD4DFF4D83EC35233DD7" blockId="24.[151,1437,150,1869]" pageId="24" pageNumber="425">
<emphasis id="B93CEACCFF94DD4DFF4D83EC37363EC2" bold="true" box="[199,351,150,176]" pageId="24" pageNumber="425">Colouration.</emphasis>
Live male (
<figureCitation id="13732A5BFF94DD4DFE6E83ED34563EC3" box="[484,575,151,177]" captionStart="FIGURE 7" captionStartId="23.[151,250,1818,1842]" captionTargetBox="[151,1436,974,1796]" captionTargetId="figure-379@23.[151,1436,974,1796]" captionTargetPageId="23" captionText="FIGURE 7. Wild-caught specimens of Nothobranchius fuscotaeniatus and only known locality. A. N. fuscotaeniatus Kitonga north TAN 97-9, male, Rufiji River drainage; B. N. fuscotaeniatus Kitonga north TAN 97-9, female, Rufiji River drainage; C. N. fuscotaeniatus Kitonga north TAN 97-9, Rufiji River floodplain, type locality, shallow with dense vegetation (photograph May 31, 1997), hosting N. fuscotaeniatus, N. janpapi and N. lourensi; D. Same site as in “C”, photograph by B. Nagy in 2017. Multiple visits to the site subsequent to the original discovery in 1997 showed an absence of the species, due to substantial changes in the habitat, primarily the loss of dense vegetation and contamination of the habitat by soil washed off the sinceabandoned adjacent road. Photographs by B.R. Watters except where otherwise noted." pageId="24" pageNumber="425">Fig. 7A</figureCitation>
): The body has a series of as many as 12 prominent dark red-brown crossbars resulting from broad scale margins. Scales in the relatively wider intervening sections are a brilliant iridescent blue to blue-green with very narrow dark red-brown edgings. The snout is blue-green to yellow with irregular dark markings including some on the lips. The throat and operculum are yellow to blue-green. The exposed part of the branchiostegal membrane is yellow with black edging. Scales on the dorsum anterior to the dorsal fin may have medium brown centres forming a vague stripe between the dorsal fin and snout. The barred colour pattern of the trunk extends onto the abdomen with intervening scales grading into light green, with only vague dark edgings, that may extend posteriorly to the caudal fin base. The iris is blue-green. The dorsal fin background colour is blue-green in the basal and proximal zones grading to yellow-green through the medial zone; the background is overlaid by irregular dark red-brown to black spots and short crude bars that appear as extensions of the dark bars on the body, becoming broken up and smaller outwards; the distal zone comprises a black marginal band with very narrow light blue to blue-green edging. The anal fin is somewhat similar in colour pattern to the dorsal fin; the basal zone is blue grading to blue-green in the proximal zone through to yellow-green in the medial zone. The basal, proximal and submedial zones are overlaid by dark red-brown to black irregular markings that represent extensions of the dark bars on the body; these irregular markings usually do not extend into outer part of the medial zone but, when they do, they appear broken and sparse; the distal zone comprises a broad black marginal band with very narrow light blue to blue-green edging that is less well developed than in the dorsal fin and may only be present on the very short fin-ray extensions. The caudal fin displays a prominent concentrically-banded pattern; the basal zone is blue-green followed by a narrow arc-like dark red-brown to black band in the proximal zone, followed by a yellow-green band and another, broader, black band and then a yellow to yellow-green band; the distal zone comprises a broad black band. Pelvic fins are blue-green with a black marginal zone. Pectoral fins are hyaline with, in some specimens, very poorly developed and usually only partial, light blue margins.
</paragraph>
<paragraph id="8BF736DEFF94DD4DFF4D80D534383B43" blockId="24.[151,1437,150,1869]" pageId="24" pageNumber="425">
Live female (
<figureCitation id="13732A5BFF94DD4DFED780D537DF3DBB" box="[349,438,943,969]" captionStart="FIGURE 7" captionStartId="23.[151,250,1818,1842]" captionTargetBox="[151,1436,974,1796]" captionTargetId="figure-379@23.[151,1436,974,1796]" captionTargetPageId="23" captionText="FIGURE 7. Wild-caught specimens of Nothobranchius fuscotaeniatus and only known locality. A. N. fuscotaeniatus Kitonga north TAN 97-9, male, Rufiji River drainage; B. N. fuscotaeniatus Kitonga north TAN 97-9, female, Rufiji River drainage; C. N. fuscotaeniatus Kitonga north TAN 97-9, Rufiji River floodplain, type locality, shallow with dense vegetation (photograph May 31, 1997), hosting N. fuscotaeniatus, N. janpapi and N. lourensi; D. Same site as in “C”, photograph by B. Nagy in 2017. Multiple visits to the site subsequent to the original discovery in 1997 showed an absence of the species, due to substantial changes in the habitat, primarily the loss of dense vegetation and contamination of the habitat by soil washed off the sinceabandoned adjacent road. Photographs by B.R. Watters except where otherwise noted." pageId="24" pageNumber="425">Fig. 7B</figureCitation>
): The body has up to 10 narrow cross-bars formed by very dark red-brown or black scale margins. These are similar to those shown by the male but are less well defined in the female and more distinct anteriorly. Scales in the intervening parts of the body have iridescent light blue to blue-green centres and yellow-brown scale margins. Scales on the dorsum are yellow-brown. The head is yellow-brown with some irregular dark red-brown to black markings; the throat is pale yellow-brown. The lower part of the operculum is yellow and there is a patch immediately posterior to the operculum that has broad bright yellow scale margins. Scales on the abdomen are light yellow-brown, some showing iridescent light blue centres and bright yellow margins. The dorsal and anal fins have yellow-brown fin rays with the intervening membrane being iridescent blue-green; the distal component is grey; some dark spots are sparsely present over most of the fins and those at or near the fin bases show a crude extensional arrangement with the bars on the body. The caudal, pectoral and pelvic fins are hyaline. The iris is golden grading outwards to blue-green.
</paragraph>
</subSubSection>
<subSubSection id="C3526555FF94DD4DFF4D864034A6393F" pageId="24" pageNumber="425" type="distribution">
<paragraph id="8BF736DEFF94DD4DFF4D8640325C3B97" blockId="24.[151,1437,150,1869]" pageId="24" pageNumber="425">
<emphasis id="B93CEACCFF94DD4DFF4D864037083B26" bold="true" box="[199,353,1338,1364]" pageId="24" pageNumber="425">Distribution.</emphasis>
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<emphasis id="B93CEACCFF94DD4DFEED864134AA3B27" box="[359,707,1339,1365]" italics="true" pageId="24" pageNumber="425">Nothobranchius fuscotaeniatus</emphasis>
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is known from a single site only, comprising an originally thickly vegetated ditch on the immediate west side of a now defunct road, about
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southwards from the south bank terminus for the old ferry across the Rufiji River (
<figureCitation id="13732A5BFF94DD4DFD5986F9357F3BEF" box="[723,790,1411,1437]" captionStart="FIGURE 1" captionStartId="2.[151,250,1762,1786]" captionTargetBox="[151,1436,374,1736]" captionTargetId="figure-118@2.[151,1436,374,1736]" captionTargetPageId="2" captionText="FIGURE 1. Distribution of species belonging to the subgenus Aphyobranchius, all of which occur in the east-central and southern coastal lowlands of Tanzania: N. janpapi (yellow-filled circle), N. luekei (red-filled triangle), N. geminus (orange-filled diamond), N. lourensi (green-filled square), N. fuscotaeniatus (blue-filled inverted triangle), N. cf. janpapi (dark magenta-filled circle). The inset map shows the area of distribution in a somewhat broader geographic context, to include the distribution of N. willerti (red-filled star) in the lower Tana River area of northeastern coastal Kenya, a species that herein is considered to belong to the subgenus Adiniops, rather than Aphyobranchius to which it was ascribed in the original description by Wildekamp (1992). Note that individual symbols on the map may represent more than one locality or population where the sites are close to one another and could not reasonably be resolved at the scale of the map." pageId="24" pageNumber="425">Fig. 1</figureCitation>
). It is, therefore, within the lower Rufiji River drainage. Since the original discovery of this species by the first author and colleagues in May, 1997, and also soon after by
<bibRefCitation id="EFD94B2FFF94DD4DFF1D86B1372B3B97" author="Seegers, L." box="[151,322,1483,1509]" pageId="24" pageNumber="425" refId="ref34429" refString="Seegers, L. (1997) Killifishes of the World. Old World, Killis II. (Apolocheilus, Epiplatys, Nothobranchius) Aqualog. Verlag A. C. S. GmbH, Morfelden-Waldorf, 112 pp." type="book" year="1997">Seegers (1997)</bibRefCitation>
, numerous visits to that site have been unable to find it there again.
</paragraph>
<paragraph id="8BF736DEFF94DD4DFF4D869434A6393F" blockId="24.[151,1437,150,1869]" pageId="24" pageNumber="425">
<emphasis id="B93CEACCFF94DD4DFF4D869437A9387B" bold="true" box="[199,448,1518,1545]" pageId="24" pageNumber="425">Conservation status.</emphasis>
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<emphasis id="B93CEACCFF94DD4DFE4186953545387B" box="[459,812,1519,1545]" italics="true" pageId="24" pageNumber="425">Nothobranchius fuscotaeniatus</emphasis>
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is listed as Critically Endangered B2ab(iii) (
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). The species is currently known only from the
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locality with an area of occupancy (AOO) of
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. Several surveys in the years following the original discovery were unable to detect its presence at either the
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locality or elsewhere in the drainage of the Rufiji River, whereas other congeners were found in abundance at the
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locality and in the general area. As shown in
<bibRefCitation id="EFD94B2FFF94DD4DFD6C8505358338EB" author="Nagy, B. &amp; Watters, B. R." box="[742,1002,1663,1689]" pageId="24" pageNumber="425" pagination="199 - 216" refId="ref33874" refString="Nagy, B. &amp; Watters, B. R. (2021) A review of the conservation status of seasonal Nothobranchius fishes (Teleostei: Cyprinodontiformes), a genus with a high level of threat, inhabiting ephemeral wetland habitats in Africa. Aquatic Conservation: Marine and Freshwater Ecosystems, 32 (1), 199 - 216. https: // doi. org / 10.1002 / aqc. 3741" type="journal article" year="2021">Nagy &amp; Watters (2021)</bibRefCitation>
and
<figureCitation id="13732A5BFF94DD4DFB95850532F938EB" box="[1055,1168,1663,1689]" captionStart="FIGURE 7" captionStartId="23.[151,250,1818,1842]" captionTargetBox="[151,1436,974,1796]" captionTargetId="figure-379@23.[151,1436,974,1796]" captionTargetPageId="23" captionText="FIGURE 7. Wild-caught specimens of Nothobranchius fuscotaeniatus and only known locality. A. N. fuscotaeniatus Kitonga north TAN 97-9, male, Rufiji River drainage; B. N. fuscotaeniatus Kitonga north TAN 97-9, female, Rufiji River drainage; C. N. fuscotaeniatus Kitonga north TAN 97-9, Rufiji River floodplain, type locality, shallow with dense vegetation (photograph May 31, 1997), hosting N. fuscotaeniatus, N. janpapi and N. lourensi; D. Same site as in “C”, photograph by B. Nagy in 2017. Multiple visits to the site subsequent to the original discovery in 1997 showed an absence of the species, due to substantial changes in the habitat, primarily the loss of dense vegetation and contamination of the habitat by soil washed off the sinceabandoned adjacent road. Photographs by B.R. Watters except where otherwise noted." pageId="24" pageNumber="425">Figure 7D</figureCitation>
herein, the
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locality has been significantly modified since the species was first discovered in 1997. Extensive local farming around the
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locality represents increasing pressure on land resources that has resulted in habitat degradation in the area. The species is adapted to specific habitat conditions by occupying a niche among the densely vegetated marginal zones of the pool and is likely, therefore, to be very highly susceptible to habitat modifications by agriculture, which invariably results in habitats becoming more open.
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<emphasis id="B93CEACCFF9DDD44FF1D850437AE38EB" bold="true" box="[151,455,1662,1689]" italics="true" pageId="17" pageNumber="418">Nothobranchius lourensi</emphasis>
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<subSubSection id="C3526555FF9DDD44FF1D85D9374438CF" box="[151,301,1699,1725]" pageId="17" pageNumber="418" type="description">
<paragraph id="8BF736DEFF9DDD44FF1D85D9374438CF" blockId="17.[151,682,1662,1725]" box="[151,301,1699,1725]" pageId="17" pageNumber="418">
(
<figureCitation id="13732A5BFF9DDD44FF1585D9374D38CF" box="[159,292,1699,1725]" captionStart="FIGURE 6" captionStartId="19.[151,250,1409,1433]" captionTargetBox="[151,1435,179,1383]" captionTargetId="figure-19@19.[151,1435,179,1383]" captionTargetPageId="19" captionText="FIGURE 6. Wild-caught specimens of Nothobranchius lourensi and examples of typical habitat. A. N. lourensi Ifakara TAN 95-4, male, Kilombero River drainage; B. N. lourensi Kitonga north TAN 97-9, male, lower Rufiji River drainage; C. N. lourensi TSTS 10-12, male, Kilombero River drainage; D. N. lourensi TSTS 10-12, female, Kilombero River drainage; E. Location Ihango TAN 02-25, a typical thickly vegetated shallow habitat on the floodplain of the Kilombero River, Kibasira Swamp (photograph June 9, 2002), hosting N. lourensi and N. lucius; F. Location Narubungo TAN 02-24, very thickly vegetated residual pool alongside a railway embankment, Kilombero River drainage, Kibasira Swamp (photograph June 9, 2002), hosting N. lourensi, N. geminus, N. kilomberoensis and N. lucius. See also FIGUREs 5C and D, and FIGURE 7C for further examples of seasonal pools that host N. lourensi. Photographs by B.R. Watters (A, B, E, F) and B. Nagy (C, D)." pageId="17" pageNumber="418">Figs. 6A–D</figureCitation>
)
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; –
<materialsCitation id="3B203C83FF9DDD44FC43859132AC395B" collectionCode="RMCA" country="Tanzania" latitude="-6.6983333" location="Kwaraza village" longLatPrecision="129" longitude="38.705" pageId="17" pageNumber="418" specimenCode="RMCA 77-16" specimenCount="9" specimenCount-female="4" specimenCount-male="5" typeStatus="holotype">
<specimenCode id="DBEE9EA5FF9DDD44FC438591320A3977" box="[969,1123,1771,1797]" collectionCode="RMCA" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="17" pageNumber="418" type="Museum">RMCA 77-16</specimenCode>
-P-29-33,
<specimenCount id="9D4EFD57FF9DDD44FB5E859033443977" box="[1236,1325,1770,1797]" count="5" pageId="17" pageNumber="418" type="male">5 males</specimenCount>
,
<quantity id="4CB09B3BFF9DDD44FAB3859133F43977" box="[1337,1437,1771,1797]" metricMagnitude="-2" metricUnit="m" metricValue="3.9299999999999997" pageId="17" pageNumber="418" unit="mm" value="39.3">39.3 mm</quantity>
SL,
<specimenCount id="9D4EFD57FF9DDD44FF4284753750395B" box="[200,313,1807,1833]" count="4" pageId="17" pageNumber="418" type="female">4 females</specimenCount>
,
<quantity id="4CB09B3BFF9DDD44FECC84743782395B" box="[326,491,1806,1833]" metricMagnitude="-2" metricUnit="m" metricValue="3.29" metricValueMax="3.54" metricValueMin="3.04" pageId="17" pageNumber="418" unit="mm" value="32.9" valueMax="35.4" valueMin="30.4">30.4–35.4 mm</quantity>
SL;
<collectingCountry id="F35F764EFF9DDD44FDAE847534E4395B" box="[548,653,1807,1833]" name="Tanzania" pageId="17" pageNumber="418">Tanzania</collectingCountry>
:
<location id="8E976005FF9DDD44FD108475353F395B" LSID="urn:lsid:plazi:treatment:03E187C8FF9DDD40FF1D850434623DBB:8E976005FF9DDD44FD108475353F395B" box="[666,854,1807,1833]" country="Tanzania" latitude="-6.6983333" longLatPrecision="129" longitude="38.705" name="Kwaraza village" pageId="17" pageNumber="418">Kwaraza village</location>
; approx.
<geoCoordinate id="EE7C5019FF9DDD44FC4B84753253395B" box="[961,1082,1807,1833]" degrees="06" direction="south" minutes="41.9" orientation="latitude" pageId="17" pageNumber="418" precision="92" value="-6.6983333">06°41.9’ S</geoCoordinate>
,
<geoCoordinate id="EE7C5019FF9DDD44FBC3847532AC395B" box="[1097,1221,1807,1833]" degrees="38" direction="east" minutes="42.3" orientation="longitude" pageId="17" pageNumber="418" precision="92" value="38.705">38°42.3’ E</geoCoordinate>
</materialsCitation>
;
<materialsCitation id="3B203C83FF9DDD44FB5984753313395B" box="[1235,1402,1807,1833]" collectionCode="R" country="Tanzania" location="Wildekamp" pageId="17" pageNumber="418" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED59AE1BFF9DDD44FB5984753280395B" box="[1235,1257,1807,1833]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="418">R</collectionCode>
.
<location id="8E976005FF9DDD44FB7F84753313395B" LSID="urn:lsid:plazi:treatment:03E187C8FF9DDD40FF1D850434623DBB:8E976005FF9DDD44FB7F84753313395B" box="[1269,1402,1807,1833]" country="Tanzania" name="Wildekamp" pageId="17" pageNumber="418">Wildekamp</location>
</materialsCitation>
,
<materialsCitation id="3B203C83FF9DDD44FA0C84753695393F" collectionCode="T" country="Tanzania" location="Steinfort" pageId="17" pageNumber="418" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED59AE1BFF9DDD44FA0C847533F1395B" box="[1414,1432,1807,1833]" name="Tavera, Department of Geology and Geophysics" pageId="17" pageNumber="418">T</collectionCode>
.
<location id="8E976005FF9DDD44FF1D84493695393F" LSID="urn:lsid:plazi:treatment:03E187C8FF9DDD40FF1D850434623DBB:8E976005FF9DDD44FF1D84493695393F" box="[151,252,1843,1869]" country="Tanzania" name="Steinfort" pageId="17" pageNumber="418">Steinfort</location>
</materialsCitation>
,
<materialsCitation id="3B203C83FF9DDD44FE8D844934B7393F" box="[263,734,1843,1869]" collectingDate="1976-06-12" collectionCode="R" collectorName="Haarlem" country="Tanzania" pageId="17" pageNumber="418" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED59AE1BFF9DDD44FE8D84493774393F" box="[263,285,1843,1869]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="418">R</collectionCode>
. van
<collectorName id="26BD5308FF9DDD44FEDD844937D2393F" box="[343,443,1843,1869]" pageId="17" pageNumber="418">Haarlem</collectorName>
;
<date id="FFF6101EFF9DDD44FE4C84493436393F" box="[454,607,1843,1869]" pageId="17" pageNumber="418" value="1976-06-12">
<collectingDate id="EFB2E9F6FF9DDD44FE4C84493436393F" box="[454,607,1843,1869]" pageId="17" pageNumber="418" value="1976-06-12">12 June 1976</collectingDate>
</date>
; field code
</materialsCitation>
:
<materialsCitation id="3B203C83FF9DDD44FD628449355F393F" box="[744,822,1843,1869]" collectionCode="T" country="Tanzania" pageId="17" pageNumber="418" specimenCode="T 76-8" specimenCount="1" typeStatus="holotype">
<specimenCode id="DBEE9EA5FF9DDD44FD628449355F393F" box="[744,822,1843,1869]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="17" pageNumber="418">T 76-8</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BF736DEFF9DDD44FF4D842C32DF39E7" blockId="17.[151,1437,1770,2013]" pageId="17" pageNumber="418">
<emphasis id="B93CEACCFF9DDD44FF4D842C34593902" bold="true" box="[199,560,1878,1904]" pageId="17" pageNumber="418">Additional material examined.</emphasis>
<materialsCitation id="3B203C83FF9DDD44FDBF842D32DA39E7" collectingDate="1995-06-07" collectionCode="R" collectorName="B. Watters &amp; Wildekamp, I" country="Tanzania" latitude="-8.167367" location="Kilombero" longLatPrecision="1" longitude="36.693516" municipality="Ifakara" pageId="17" pageNumber="418" specimenCount="3" specimenCount-male="3">
<specimenCount id="9D4EFD57FF9DDD44FDBF842D34E53903" box="[565,652,1879,1905]" count="3" pageId="17" pageNumber="418" type="male">3 males</specimenCount>
,
<quantity id="4CB09B3BFF9DDD44FD1C842C355E3903" box="[662,823,1878,1905]" metricMagnitude="-2" metricUnit="m" metricValue="3.405" metricValueMax="3.56" metricValueMin="3.25" pageId="17" pageNumber="418" unit="mm" value="34.05" valueMax="35.6" valueMin="32.5">32.5–35.6 mm</quantity>
SL;
<collectingCountry id="F35F764EFF9DDD44FCE2842D35B83903" box="[872,977,1879,1905]" name="Tanzania" pageId="17" pageNumber="418">Tanzania</collectingCountry>
:
<location id="8E976005FF9DDD44FC50842D323A3903" LSID="urn:lsid:plazi:treatment:03E187C8FF9DDD40FF1D850434623DBB:8E976005FF9DDD44FC50842D323A3903" box="[986,1107,1879,1905]" country="Tanzania" latitude="-8.167367" longLatPrecision="1" longitude="36.693516" municipality="Ifakara" name="Kilombero" pageId="17" pageNumber="418">Kilombero</location>
drainage,
<collectingMunicipality id="6B93ACA4FF9DDD44FB4F842D33713903" box="[1221,1304,1879,1905]" pageId="17" pageNumber="418">Ifakara</collectingMunicipality>
;
<geoCoordinate id="EE7C5019FF9DDD44FAAB842D36C139E7" degrees="08" direction="south" minutes="10.042" orientation="latitude" pageId="17" pageNumber="418" precision="1" value="-8.167367">08°10.042’ S</geoCoordinate>
,
<geoCoordinate id="EE7C5019FF9DDD44FF3F8401372339E7" box="[181,330,1915,1941]" degrees="36" direction="east" minutes="41.611" orientation="longitude" pageId="17" pageNumber="418" precision="1" value="36.693516">36°41.611’ E</geoCoordinate>
;
<collectorName id="26BD5308FF9DDD44FEDC840137A739E7" box="[342,462,1915,1941]" pageId="17" pageNumber="418">B. Watters</collectorName>
,
<collectionCode id="ED59AE1BFF9DDD44FE538401378639E7" box="[473,495,1915,1941]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="418">R</collectionCode>
.
<collectorName id="26BD5308FF9DDD44FE73840134FC39E7" box="[505,661,1915,1941]" pageId="17" pageNumber="418">Wildekamp, I</collectorName>
. Sainthouse;
<date id="FFF6101EFF9DDD44FCA0840135DD39E7" box="[810,948,1914,1941]" pageId="17" pageNumber="418" value="1995-06-07">
<collectingDate id="EFB2E9F6FF9DDD44FCA0840135DD39E7" box="[810,948,1914,1941]" pageId="17" pageNumber="418" value="1995-06-07">7 June 1995</collectingDate>
</date>
; field code: TAN 95-4
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3526555FF9DDD47FF4D84E432B63FFB" lastPageId="18" lastPageNumber="419" pageId="17" pageNumber="418" type="diagnosis">
<paragraph id="8BF736DEFF9DDD47FF4D84E432B63FFB" blockId="17.[151,1437,1770,2013]" lastBlockId="18.[151,1436,151,465]" lastPageId="18" lastPageNumber="419" pageId="17" pageNumber="418">
<emphasis id="B93CEACCFF9DDD44FF4D84E4372B39CA" bold="true" box="[199,322,1950,1976]" pageId="17" pageNumber="418">Diagnosis.</emphasis>
<taxonomicName id="4C484D5DFF9DDD44FECC84E5343339CB" authorityName=", Wildekamp" authorityYear="1977" box="[326,602,1951,1977]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="17" pageNumber="418" phylum="Chordata" rank="species" species="lourensi">
<emphasis id="B93CEACCFF9DDD44FECC84E5343339CB" box="[326,602,1951,1977]" italics="true" pageId="17" pageNumber="418">Nothobranchius lourensi</emphasis>
</taxonomicName>
is distinguished from all the other members of the genus
<taxonomicName id="4C484D5DFF9DDD44FB4D84E5331239CB" authorityName="Peters" authorityYear="1868" box="[1223,1403,1951,1977]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="17" pageNumber="418" phylum="Chordata" rank="genus">
<emphasis id="B93CEACCFF9DDD44FB4D84E5331239CB" box="[1223,1403,1951,1977]" italics="true" pageId="17" pageNumber="418">Nothobranchius</emphasis>
</taxonomicName>
by the unique combination of diagnostic characters of male colouration comprising: blue-green scales with dark brown posterior scale margins, creating a reticulation on the body, with a chevron pattern on the posterior part of trunk; light blue throat; anal and caudal fins with 8–12 distinct narrow to slender bands on the entire fin, perpendicular to fin rays, alternating red-brown and light blue to light yellow, with iridescent light blue to yellow subdistal band and dark grey to black distal band; dorsal-fin anterior half with irregular red brown spots and stripes parallel to fin rays, whereas posterior half with alternating red-brown and light blue to light yellow narrow to slender bands, perpendicular to fin rays; as well as having an anal fin with posterior lower margin straight to mildly concave. Also, the female has a grey-brown body, with a dark grey chevron pattern on the posterior half of trunk.
</paragraph>
</subSubSection>
<subSubSection id="C3526555FF9EDD40FF4D82E835693FFB" lastPageId="21" lastPageNumber="422" pageId="18" pageNumber="419" type="description">
<paragraph id="8BF736DEFF9EDD47FF4D82E834F43FA3" blockId="18.[151,1436,151,465]" pageId="18" pageNumber="419">
<emphasis id="B93CEACCFF9EDD47FF4D82E837303FDE" bold="true" box="[199,345,402,428]" pageId="18" pageNumber="419">Description.</emphasis>
General body features illustrated by
<figureCitation id="13732A5BFF9EDD47FC8582E935DC3FDF" box="[783,949,403,429]" captionStart="FIGURE 6" captionStartId="19.[151,250,1409,1433]" captionTargetBox="[151,1435,179,1383]" captionTargetId="figure-19@19.[151,1435,179,1383]" captionTargetPageId="19" captionText="FIGURE 6. Wild-caught specimens of Nothobranchius lourensi and examples of typical habitat. A. N. lourensi Ifakara TAN 95-4, male, Kilombero River drainage; B. N. lourensi Kitonga north TAN 97-9, male, lower Rufiji River drainage; C. N. lourensi TSTS 10-12, male, Kilombero River drainage; D. N. lourensi TSTS 10-12, female, Kilombero River drainage; E. Location Ihango TAN 02-25, a typical thickly vegetated shallow habitat on the floodplain of the Kilombero River, Kibasira Swamp (photograph June 9, 2002), hosting N. lourensi and N. lucius; F. Location Narubungo TAN 02-24, very thickly vegetated residual pool alongside a railway embankment, Kilombero River drainage, Kibasira Swamp (photograph June 9, 2002), hosting N. lourensi, N. geminus, N. kilomberoensis and N. lucius. See also FIGUREs 5C and D, and FIGURE 7C for further examples of seasonal pools that host N. lourensi. Photographs by B.R. Watters (A, B, E, F) and B. Nagy (C, D)." pageId="18" pageNumber="419">Figures 6A–D</figureCitation>
. Morphometric and meristic characters of examined material are summarized in
<tableCitation id="C6CA0365FF9EDD47FDC982CD34F03FA3" box="[579,665,439,465]" captionStart="TABLE 6" captionStartId="18.[151,239,512,536]" captionText="TABLE 6. Morphometric and meristic data of Nothobranchius lourensi. SD, standard deviation." pageId="18" pageNumber="419">Table 6</tableCitation>
.
</paragraph>
<caption id="DF376656FF9EDD47FF1D817A32F43C6A" box="[151,1181,512,537]" pageId="18" pageNumber="419" startId="18.[151,239,512,536]" targetType="table">
<paragraph id="8BF736DEFF9EDD47FF1D817A32F43C6A" blockId="18.[151,1181,512,537]" box="[151,1181,512,537]" pageId="18" pageNumber="419">
<emphasis id="B93CEACCFF9EDD47FF1D817A37633C6A" bold="true" box="[151,266,512,536]" pageId="18" pageNumber="419">TABLE 6.</emphasis>
Morphometric and meristic data of
<taxonomicName id="4C484D5DFF9EDD47FD01817A35FD3C6A" authorityName=", Wildekamp" authorityYear="1977" box="[651,916,512,536]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="18" pageNumber="419" phylum="Chordata" rank="species" species="lourensi">
<emphasis id="B93CEACCFF9EDD47FD01817A35FD3C6A" box="[651,916,512,536]" italics="true" pageId="18" pageNumber="419">Nothobranchius lourensi</emphasis>
</taxonomicName>
. SD, standard deviation.
</paragraph>
</caption>
<paragraph id="8BF736DEFF9EDD47FCB38153329E3998" pageId="18" pageNumber="419">
<table id="F948C47EFF9E22AAFF13815333003998" box="[153,1385,553,2026]" colsContinueIn="22.[153,1394,192,1665]" gridcols="7" gridrows="36" pageId="18" pageNumber="419">
<tr id="3578349CFF9E22AAFF13815333003C33" box="[153,1385,553,577]" gridrow="0" pageId="18" pageNumber="419" rowspan-0="1" rowspan-3="1" rowspan-4="1" rowspan-6="1">
<th id="76A95DE0FF9E22AAFD4F815335E63C33" box="[709,911,553,577]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="18" pageNumber="419">males</th>
<th id="76A95DE0FF9E22AAFB378153337E3C33" box="[1213,1303,553,577]" gridcol="5" gridrow="0" pageId="18" pageNumber="419">females</th>
</tr>
<tr id="3578349CFF9E22AAFF13812B33003C1B" box="[153,1385,593,617]" gridrow="1" pageId="18" pageNumber="419" rowspan-0="1" rowspan-3="1" rowspan-4="1" rowspan-6="1">
<td id="76A95DE0FF9E22AAFD4F812B35E63C1B" box="[709,911,593,617]" colspan="2" colspanRight="1" gridcol="1" gridrow="1" pageId="18" pageNumber="419">(n = 5)</td>
<td id="76A95DE0FF9E22AAFB37812B337E3C1B" box="[1213,1303,593,617]" gridcol="5" gridrow="1" pageId="18" pageNumber="419">(n = 4)</td>
</tr>
<tr id="3578349CFF9E22AAFF1381D833003CC8" box="[153,1385,674,698]" gridrow="2" pageId="18" pageNumber="419" rowspan-0="1">
<td id="76A95DE0FF9E22AAFD4F81D8352D3CC8" box="[709,836,674,698]" gridcol="1" gridrow="2" pageId="18" pageNumber="419">range</td>
<td id="76A95DE0FF9E22AAFCDC81D835E63CC8" box="[854,911,674,698]" gridcol="2" gridrow="2" pageId="18" pageNumber="419">mean</td>
<td id="76A95DE0FF9E22AAFC3A81D835B93CC8" box="[944,976,674,698]" gridcol="3" gridrow="2" pageId="18" pageNumber="419">SD</td>
<td id="76A95DE0FF9E22AAFBA481D832FD3CC8" box="[1070,1172,674,698]" gridcol="4" gridrow="2" pageId="18" pageNumber="419">range</td>
<td id="76A95DE0FF9E22AAFB3781D8337E3CC8" box="[1213,1303,674,698]" gridcol="5" gridrow="2" pageId="18" pageNumber="419">mean</td>
<td id="76A95DE0FF9E22AAFAC381D833003CC8" box="[1353,1385,674,698]" gridcol="6" gridrow="2" pageId="18" pageNumber="419">SD</td>
</tr>
<tr id="3578349CFF9E22AAFF1381B333003C90" box="[153,1385,713,738]" gridrow="3" pageId="18" pageNumber="419" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1">
<th id="76A95DE0FF9E22AAFF1381B334E33C90" box="[153,650,713,738]" gridcol="0" gridrow="3" pageId="18" pageNumber="419">Standard length (mm)</th>
<td id="76A95DE0FF9E22AAFD4F81B3352D3C90" box="[709,836,713,738]" gridcol="1" gridrow="3" pageId="18" pageNumber="419">32.5–40.6</td>
<td id="76A95DE0FF9E22AAFBA481B332FD3C90" box="[1070,1172,713,738]" gridcol="4" gridrow="3" pageId="18" pageNumber="419">30.4–35.4</td>
</tr>
<tr id="3578349CFF9E22AAFF13818B33003D78" box="[153,1385,753,778]" gridrow="4" pageId="18" pageNumber="419">
<th id="76A95DE0FF9E22AAFF13818B34E33D78" box="[153,650,753,778]" gridcol="0" gridrow="4" pageId="18" pageNumber="419">
<emphasis id="B93CEACCFF9EDD47FF13818837D13D78" bold="true" box="[153,440,753,778]" pageId="18" pageNumber="419">Percent of standard length</emphasis>
</th>
<td id="76A95DE0FF9E22AAFD4F818B352D3D78" box="[709,836,753,778]" gridcol="1" gridrow="4" pageId="18" pageNumber="419">min–max</td>
<td id="76A95DE0FF9E22AAFCDC818B35E63D78" box="[854,911,753,778]" gridcol="2" gridrow="4" pageId="18" pageNumber="419">mean</td>
<td id="76A95DE0FF9E22AAFC3A818B35B93D78" box="[944,976,753,778]" gridcol="3" gridrow="4" pageId="18" pageNumber="419">SD</td>
<td id="76A95DE0FF9E22AAFBA4818B32FD3D78" box="[1070,1172,753,778]" gridcol="4" gridrow="4" pageId="18" pageNumber="419">min–max</td>
<td id="76A95DE0FF9E22AAFB37818B337E3D78" box="[1213,1303,753,778]" gridcol="5" gridrow="4" pageId="18" pageNumber="419">mean</td>
<td id="76A95DE0FF9E22AAFAC3818B33003D78" box="[1353,1385,753,778]" gridcol="6" gridrow="4" pageId="18" pageNumber="419">SD</td>
</tr>
<tr id="3578349CFF9E22AAFF13806033003D40" box="[153,1385,794,818]" gridrow="5" pageId="18" pageNumber="419">
<th id="76A95DE0FF9E22AAFF13806034E33D40" box="[153,650,794,818]" gridcol="0" gridrow="5" pageId="18" pageNumber="419">Total length</th>
<td id="76A95DE0FF9E22AAFD4F8060352D3D40" box="[709,836,794,818]" gridcol="1" gridrow="5" pageId="18" pageNumber="419">120.2–126.5</td>
<td id="76A95DE0FF9E22AAFCDC806035E63D40" box="[854,911,794,818]" gridcol="2" gridrow="5" pageId="18" pageNumber="419">123.1</td>
<td id="76A95DE0FF9E22AAFC3A806035B93D40" box="[944,976,794,818]" gridcol="3" gridrow="5" pageId="18" pageNumber="419">2.4</td>
<td id="76A95DE0FF9E22AAFBA48060337E3D40" box="[1070,1303,794,818]" colspan="2" colspanRight="1" gridcol="4" gridrow="5" pageId="18" pageNumber="419">120.6–125.0 122.6</td>
<td id="76A95DE0FF9E22AAFAC3806033003D40" box="[1353,1385,794,818]" gridcol="6" gridrow="5" pageId="18" pageNumber="419">1.8</td>
</tr>
<tr id="3578349CFF9E22AAFF13803833003D28" box="[153,1385,834,858]" gridrow="6" pageId="18" pageNumber="419">
<th id="76A95DE0FF9E22AAFF13803834E33D28" box="[153,650,834,858]" gridcol="0" gridrow="6" pageId="18" pageNumber="419">Body depth at pelvic-fin origin</th>
<td id="76A95DE0FF9E22AAFD4F8038352D3D28" box="[709,836,834,858]" gridcol="1" gridrow="6" pageId="18" pageNumber="419">28.6–34.0</td>
<td id="76A95DE0FF9E22AAFCDC803835E63D28" box="[854,911,834,858]" gridcol="2" gridrow="6" pageId="18" pageNumber="419">30.7</td>
<td id="76A95DE0FF9E22AAFC3A803835B93D28" box="[944,976,834,858]" gridcol="3" gridrow="6" pageId="18" pageNumber="419">2.2</td>
<td id="76A95DE0FF9E22AAFBA4803832FD3D28" box="[1070,1172,834,858]" gridcol="4" gridrow="6" pageId="18" pageNumber="419">28.2–34.9</td>
<td id="76A95DE0FF9E22AAFB378038337E3D28" box="[1213,1303,834,858]" gridcol="5" gridrow="6" pageId="18" pageNumber="419">31.7</td>
<td id="76A95DE0FF9E22AAFAC3803833003D28" box="[1353,1385,834,858]" gridcol="6" gridrow="6" pageId="18" pageNumber="419">3.0</td>
</tr>
<tr id="3578349CFF9E22AAFF13801033003DF1" box="[153,1385,874,899]" gridrow="7" pageId="18" pageNumber="419">
<th id="76A95DE0FF9E22AAFF13801034E33DF1" box="[153,650,874,899]" gridcol="0" gridrow="7" pageId="18" pageNumber="419">Head length</th>
<td id="76A95DE0FF9E22AAFD4F8010352D3DF1" box="[709,836,874,899]" gridcol="1" gridrow="7" pageId="18" pageNumber="419">29.2–32.8</td>
<td id="76A95DE0FF9E22AAFCDC801035E63DF1" box="[854,911,874,899]" gridcol="2" gridrow="7" pageId="18" pageNumber="419">30.7</td>
<td id="76A95DE0FF9E22AAFC3A801035B93DF1" box="[944,976,874,899]" gridcol="3" gridrow="7" pageId="18" pageNumber="419">1.5</td>
<td id="76A95DE0FF9E22AAFBA4801032FD3DF1" box="[1070,1172,874,899]" gridcol="4" gridrow="7" pageId="18" pageNumber="419">31.7–33.6</td>
<td id="76A95DE0FF9E22AAFB378010337E3DF1" box="[1213,1303,874,899]" gridcol="5" gridrow="7" pageId="18" pageNumber="419">32.6</td>
<td id="76A95DE0FF9E22AAFAC3801033003DF1" box="[1353,1385,874,899]" gridcol="6" gridrow="7" pageId="18" pageNumber="419">0.9</td>
</tr>
<tr id="3578349CFF9E22AAFF1380E933003DD9" box="[153,1385,915,939]" gridrow="8" pageId="18" pageNumber="419">
<th id="76A95DE0FF9E22AAFF1380E934E33DD9" box="[153,650,915,939]" gridcol="0" gridrow="8" pageId="18" pageNumber="419">Preanal length</th>
<td id="76A95DE0FF9E22AAFD4F80E9352D3DD9" box="[709,836,915,939]" gridcol="1" gridrow="8" pageId="18" pageNumber="419">53.7–57.6</td>
<td id="76A95DE0FF9E22AAFCDC80E935E63DD9" box="[854,911,915,939]" gridcol="2" gridrow="8" pageId="18" pageNumber="419">55.5</td>
<td id="76A95DE0FF9E22AAFC3A80E935B93DD9" box="[944,976,915,939]" gridcol="3" gridrow="8" pageId="18" pageNumber="419">1.7</td>
<td id="76A95DE0FF9E22AAFBA480E932FD3DD9" box="[1070,1172,915,939]" gridcol="4" gridrow="8" pageId="18" pageNumber="419">61.7–67.1</td>
<td id="76A95DE0FF9E22AAFB3780E9337E3DD9" box="[1213,1303,915,939]" gridcol="5" gridrow="8" pageId="18" pageNumber="419">65.3</td>
<td id="76A95DE0FF9E22AAFAC380E933003DD9" box="[1353,1385,915,939]" gridcol="6" gridrow="8" pageId="18" pageNumber="419">2.5</td>
</tr>
<tr id="3578349CFF9E22AAFF1380C133003DA1" box="[153,1385,955,979]" gridrow="9" pageId="18" pageNumber="419">
<th id="76A95DE0FF9E22AAFF1380C134E33DA1" box="[153,650,955,979]" gridcol="0" gridrow="9" pageId="18" pageNumber="419">Predorsal length</th>
<td id="76A95DE0FF9E22AAFD4F80C1352D3DA1" box="[709,836,955,979]" gridcol="1" gridrow="9" pageId="18" pageNumber="419">52.3–56.2</td>
<td id="76A95DE0FF9E22AAFCDC80C135E63DA1" box="[854,911,955,979]" gridcol="2" gridrow="9" pageId="18" pageNumber="419">54.8</td>
<td id="76A95DE0FF9E22AAFC3A80C135B93DA1" box="[944,976,955,979]" gridcol="3" gridrow="9" pageId="18" pageNumber="419">1.6</td>
<td id="76A95DE0FF9E22AAFBA480C132FD3DA1" box="[1070,1172,955,979]" gridcol="4" gridrow="9" pageId="18" pageNumber="419">55.4–60.9</td>
<td id="76A95DE0FF9E22AAFB3780C1337E3DA1" box="[1213,1303,955,979]" gridcol="5" gridrow="9" pageId="18" pageNumber="419">58.0</td>
<td id="76A95DE0FF9E22AAFAC380C133003DA1" box="[1353,1385,955,979]" gridcol="6" gridrow="9" pageId="18" pageNumber="419">2.3</td>
</tr>
<tr id="3578349CFF9E22AAFF13809933003D89" box="[153,1385,995,1019]" gridrow="10" pageId="18" pageNumber="419">
<th id="76A95DE0FF9E22AAFF13809934E33D89" box="[153,650,995,1019]" gridcol="0" gridrow="10" pageId="18" pageNumber="419">Prepelvic length</th>
<td id="76A95DE0FF9E22AAFD4F8099352D3D89" box="[709,836,995,1019]" gridcol="1" gridrow="10" pageId="18" pageNumber="419">40.7–44.6</td>
<td id="76A95DE0FF9E22AAFCDC809935E63D89" box="[854,911,995,1019]" gridcol="2" gridrow="10" pageId="18" pageNumber="419">43.0</td>
<td id="76A95DE0FF9E22AAFC3A809935B93D89" box="[944,976,995,1019]" gridcol="3" gridrow="10" pageId="18" pageNumber="419">1.4</td>
<td id="76A95DE0FF9E22AAFBA4809932FD3D89" box="[1070,1172,995,1019]" gridcol="4" gridrow="10" pageId="18" pageNumber="419">44.6–50.0</td>
<td id="76A95DE0FF9E22AAFB378099337E3D89" box="[1213,1303,995,1019]" gridcol="5" gridrow="10" pageId="18" pageNumber="419">48.4</td>
<td id="76A95DE0FF9E22AAFAC3809933003D89" box="[1353,1385,995,1019]" gridcol="6" gridrow="10" pageId="18" pageNumber="419">2.5</td>
</tr>
<tr id="3578349CFF9E22AAFF13877133003A56" box="[153,1385,1035,1060]" gridrow="11" pageId="18" pageNumber="419">
<th id="76A95DE0FF9E22AAFF13877134E33A56" box="[153,650,1035,1060]" gridcol="0" gridrow="11" pageId="18" pageNumber="419">Prepectoral length</th>
<td id="76A95DE0FF9E22AAFD4F8771352D3A56" box="[709,836,1035,1060]" gridcol="1" gridrow="11" pageId="18" pageNumber="419">29.5–33.1</td>
<td id="76A95DE0FF9E22AAFCDC877135E63A56" box="[854,911,1035,1060]" gridcol="2" gridrow="11" pageId="18" pageNumber="419">31.0</td>
<td id="76A95DE0FF9E22AAFC3A877135B93A56" box="[944,976,1035,1060]" gridcol="3" gridrow="11" pageId="18" pageNumber="419">1.5</td>
<td id="76A95DE0FF9E22AAFBA4877132FD3A56" box="[1070,1172,1035,1060]" gridcol="4" gridrow="11" pageId="18" pageNumber="419">31.9–34.2</td>
<td id="76A95DE0FF9E22AAFB378771337E3A56" box="[1213,1303,1035,1060]" gridcol="5" gridrow="11" pageId="18" pageNumber="419">32.9</td>
<td id="76A95DE0FF9E22AAFAC3877133003A56" box="[1353,1385,1035,1060]" gridcol="6" gridrow="11" pageId="18" pageNumber="419">1.1</td>
</tr>
<tr id="3578349CFF9E22AAFF13874E33003A3E" box="[153,1385,1076,1100]" gridrow="12" pageId="18" pageNumber="419">
<th id="76A95DE0FF9E22AAFF13874E34E33A3E" box="[153,650,1076,1100]" gridcol="0" gridrow="12" pageId="18" pageNumber="419">Caudal peduncle length</th>
<td id="76A95DE0FF9E22AAFD4F874E352D3A3E" box="[709,836,1076,1100]" gridcol="1" gridrow="12" pageId="18" pageNumber="419">18.6–21.1</td>
<td id="76A95DE0FF9E22AAFCDC874E35E63A3E" box="[854,911,1076,1100]" gridcol="2" gridrow="12" pageId="18" pageNumber="419">20.2</td>
<td id="76A95DE0FF9E22AAFC3A874E35B93A3E" box="[944,976,1076,1100]" gridcol="3" gridrow="12" pageId="18" pageNumber="419">1.0</td>
<td id="76A95DE0FF9E22AAFBA4874E32FD3A3E" box="[1070,1172,1076,1100]" gridcol="4" gridrow="12" pageId="18" pageNumber="419">19.7–21.7</td>
<td id="76A95DE0FF9E22AAFB37874E337E3A3E" box="[1213,1303,1076,1100]" gridcol="5" gridrow="12" pageId="18" pageNumber="419">20.6</td>
<td id="76A95DE0FF9E22AAFAC3874E33003A3E" box="[1353,1385,1076,1100]" gridcol="6" gridrow="12" pageId="18" pageNumber="419">1.0</td>
</tr>
<tr id="3578349CFF9E22AAFF13872633003A06" box="[153,1385,1116,1140]" gridrow="13" pageId="18" pageNumber="419">
<th id="76A95DE0FF9E22AAFF13872634E33A06" box="[153,650,1116,1140]" gridcol="0" gridrow="13" pageId="18" pageNumber="419">Caudal peduncle depth</th>
<td id="76A95DE0FF9E22AAFD4F8726352D3A06" box="[709,836,1116,1140]" gridcol="1" gridrow="13" pageId="18" pageNumber="419">12.7–14.6</td>
<td id="76A95DE0FF9E22AAFCDC872635E63A06" box="[854,911,1116,1140]" gridcol="2" gridrow="13" pageId="18" pageNumber="419">13.7</td>
<td id="76A95DE0FF9E22AAFC3A872635B93A06" box="[944,976,1116,1140]" gridcol="3" gridrow="13" pageId="18" pageNumber="419">0.8</td>
<td id="76A95DE0FF9E22AAFBA4872632FD3A06" box="[1070,1172,1116,1140]" gridcol="4" gridrow="13" pageId="18" pageNumber="419">11.5–12.3</td>
<td id="76A95DE0FF9E22AAFB378726337E3A06" box="[1213,1303,1116,1140]" gridcol="5" gridrow="13" pageId="18" pageNumber="419">11.8</td>
<td id="76A95DE0FF9E22AAFAC3872633003A06" box="[1353,1385,1116,1140]" gridcol="6" gridrow="13" pageId="18" pageNumber="419">0.4</td>
</tr>
<tr id="3578349CFF9E22AAFF1387FE33003AEE" box="[153,1385,1156,1180]" gridrow="14" pageId="18" pageNumber="419">
<th id="76A95DE0FF9E22AAFF1387FE34E33AEE" box="[153,650,1156,1180]" gridcol="0" gridrow="14" pageId="18" pageNumber="419">Dorsal-fin base length</th>
<td id="76A95DE0FF9E22AAFD4F87FE352D3AEE" box="[709,836,1156,1180]" gridcol="1" gridrow="14" pageId="18" pageNumber="419">24.6–28.0</td>
<td id="76A95DE0FF9E22AAFCDC87FE35E63AEE" box="[854,911,1156,1180]" gridcol="2" gridrow="14" pageId="18" pageNumber="419">26.3</td>
<td id="76A95DE0FF9E22AAFC3A87FE35B93AEE" box="[944,976,1156,1180]" gridcol="3" gridrow="14" pageId="18" pageNumber="419">1.5</td>
<td id="76A95DE0FF9E22AAFBA487FE32FD3AEE" box="[1070,1172,1156,1180]" gridcol="4" gridrow="14" pageId="18" pageNumber="419">22.9–26.3</td>
<td id="76A95DE0FF9E22AAFB3787FE337E3AEE" box="[1213,1303,1156,1180]" gridcol="5" gridrow="14" pageId="18" pageNumber="419">25.1</td>
<td id="76A95DE0FF9E22AAFAC387FE33003AEE" box="[1353,1385,1156,1180]" gridcol="6" gridrow="14" pageId="18" pageNumber="419">1.5</td>
</tr>
<tr id="3578349CFF9E22AAFF1387D633003AB7" box="[153,1385,1196,1221]" gridrow="15" pageId="18" pageNumber="419">
<th id="76A95DE0FF9E22AAFF1387D634E33AB7" box="[153,650,1196,1221]" gridcol="0" gridrow="15" pageId="18" pageNumber="419">Anal-fin base length</th>
<td id="76A95DE0FF9E22AAFD4F87D6352D3AB7" box="[709,836,1196,1221]" gridcol="1" gridrow="15" pageId="18" pageNumber="419">23.4–26.7</td>
<td id="76A95DE0FF9E22AAFCDC87D635E63AB7" box="[854,911,1196,1221]" gridcol="2" gridrow="15" pageId="18" pageNumber="419">24.9</td>
<td id="76A95DE0FF9E22AAFC3A87D635B93AB7" box="[944,976,1196,1221]" gridcol="3" gridrow="15" pageId="18" pageNumber="419">1.4</td>
<td id="76A95DE0FF9E22AAFBA487D632FD3AB7" box="[1070,1172,1196,1221]" gridcol="4" gridrow="15" pageId="18" pageNumber="419">18.6–21.4</td>
<td id="76A95DE0FF9E22AAFB3787D6337E3AB7" box="[1213,1303,1196,1221]" gridcol="5" gridrow="15" pageId="18" pageNumber="419">20.6</td>
<td id="76A95DE0FF9E22AAFAC387D633003AB7" box="[1353,1385,1196,1221]" gridcol="6" gridrow="15" pageId="18" pageNumber="419">1.4</td>
</tr>
<tr id="3578349CFF9E22AAFF1387AF33003A9F" box="[153,1385,1237,1261]" gridrow="16" pageId="18" pageNumber="419">
<th id="76A95DE0FF9E22AAFF1387AF34E33A9F" box="[153,650,1237,1261]" gridcol="0" gridrow="16" pageId="18" pageNumber="419">Caudal-fin length</th>
<td id="76A95DE0FF9E22AAFD4F87AF352D3A9F" box="[709,836,1237,1261]" gridcol="1" gridrow="16" pageId="18" pageNumber="419">20.2–26.5</td>
<td id="76A95DE0FF9E22AAFCDC87AF35E63A9F" box="[854,911,1237,1261]" gridcol="2" gridrow="16" pageId="18" pageNumber="419">23.1</td>
<td id="76A95DE0FF9E22AAFC3A87AF35B93A9F" box="[944,976,1237,1261]" gridcol="3" gridrow="16" pageId="18" pageNumber="419">2.4</td>
<td id="76A95DE0FF9E22AAFBA487AF32FD3A9F" box="[1070,1172,1237,1261]" gridcol="4" gridrow="16" pageId="18" pageNumber="419">20.6–25.0</td>
<td id="76A95DE0FF9E22AAFB3787AF337E3A9F" box="[1213,1303,1237,1261]" gridcol="5" gridrow="16" pageId="18" pageNumber="419">22.6</td>
<td id="76A95DE0FF9E22AAFAC387AF33003A9F" box="[1353,1385,1237,1261]" gridcol="6" gridrow="16" pageId="18" pageNumber="419">1.8</td>
</tr>
<tr id="3578349CFF9E22AAFF13878733003B67" box="[153,1385,1277,1301]" gridrow="17" pageId="18" pageNumber="419">
<th id="76A95DE0FF9E22AAFF13878733003B67" box="[153,1385,1277,1301]" colspan="7" colspanRight="6" gridcol="0" gridrow="17" pageId="18" pageNumber="419">
<emphasis id="B93CEACCFF9EDD47FF13878737E23B67" bold="true" box="[153,395,1277,1301]" pageId="18" pageNumber="419">Percent of head length</emphasis>
</th>
</tr>
<tr id="3578349CFF9E22AAFF13865F33003B4F" box="[153,1385,1317,1341]" gridrow="18" pageId="18" pageNumber="419">
<th id="76A95DE0FF9E22AAFF13865F34E33B4F" box="[153,650,1317,1341]" gridcol="0" gridrow="18" pageId="18" pageNumber="419">Head width</th>
<td id="76A95DE0FF9E22AAFD4F865F352D3B4F" box="[709,836,1317,1341]" gridcol="1" gridrow="18" pageId="18" pageNumber="419">54–59</td>
<td id="76A95DE0FF9E22AAFCDC865F35E63B4F" box="[854,911,1317,1341]" gridcol="2" gridrow="18" pageId="18" pageNumber="419">56.3</td>
<td id="76A95DE0FF9E22AAFC3A865F35B93B4F" box="[944,976,1317,1341]" gridcol="3" gridrow="18" pageId="18" pageNumber="419">1.9</td>
<td id="76A95DE0FF9E22AAFBA4865F32FD3B4F" box="[1070,1172,1317,1341]" gridcol="4" gridrow="18" pageId="18" pageNumber="419">59–63</td>
<td id="76A95DE0FF9E22AAFB37865F337E3B4F" box="[1213,1303,1317,1341]" gridcol="5" gridrow="18" pageId="18" pageNumber="419">60.9</td>
<td id="76A95DE0FF9E22AAFAC3865F33003B4F" box="[1353,1385,1317,1341]" gridcol="6" gridrow="18" pageId="18" pageNumber="419">1.9</td>
</tr>
<tr id="3578349CFF9E22AAFF13863733003B14" box="[153,1385,1357,1382]" gridrow="19" pageId="18" pageNumber="419">
<th id="76A95DE0FF9E22AAFF13863734E33B14" box="[153,650,1357,1382]" gridcol="0" gridrow="19" pageId="18" pageNumber="419">Head depth</th>
<td id="76A95DE0FF9E22AAFD4F8637352D3B14" box="[709,836,1357,1382]" gridcol="1" gridrow="19" pageId="18" pageNumber="419">83–87</td>
<td id="76A95DE0FF9E22AAFCDC863735E63B14" box="[854,911,1357,1382]" gridcol="2" gridrow="19" pageId="18" pageNumber="419">84.8</td>
<td id="76A95DE0FF9E22AAFC3A863735B93B14" box="[944,976,1357,1382]" gridcol="3" gridrow="19" pageId="18" pageNumber="419">1.2</td>
<td id="76A95DE0FF9E22AAFBA4863732FD3B14" box="[1070,1172,1357,1382]" gridcol="4" gridrow="19" pageId="18" pageNumber="419">79–81</td>
<td id="76A95DE0FF9E22AAFB378637337E3B14" box="[1213,1303,1357,1382]" gridcol="5" gridrow="19" pageId="18" pageNumber="419">80.1</td>
<td id="76A95DE0FF9E22AAFAC3863733003B14" box="[1353,1385,1357,1382]" gridcol="6" gridrow="19" pageId="18" pageNumber="419">0.8</td>
</tr>
<tr id="3578349CFF9E22AAFF13860C33003BFC" box="[153,1385,1398,1422]" gridrow="20" pageId="18" pageNumber="419">
<th id="76A95DE0FF9E22AAFF13860C34E33BFC" box="[153,650,1398,1422]" gridcol="0" gridrow="20" pageId="18" pageNumber="419">Interorbital width</th>
<td id="76A95DE0FF9E22AAFD4F860C352D3BFC" box="[709,836,1398,1422]" gridcol="1" gridrow="20" pageId="18" pageNumber="419">33–40</td>
<td id="76A95DE0FF9E22AAFCDC860C35E63BFC" box="[854,911,1398,1422]" gridcol="2" gridrow="20" pageId="18" pageNumber="419">37.2</td>
<td id="76A95DE0FF9E22AAFC3A860C35B93BFC" box="[944,976,1398,1422]" gridcol="3" gridrow="20" pageId="18" pageNumber="419">2.8</td>
<td id="76A95DE0FF9E22AAFBA4860C32FD3BFC" box="[1070,1172,1398,1422]" gridcol="4" gridrow="20" pageId="18" pageNumber="419">31–34</td>
<td id="76A95DE0FF9E22AAFB37860C337E3BFC" box="[1213,1303,1398,1422]" gridcol="5" gridrow="20" pageId="18" pageNumber="419">32.3</td>
<td id="76A95DE0FF9E22AAFAC3860C33003BFC" box="[1353,1385,1398,1422]" gridcol="6" gridrow="20" pageId="18" pageNumber="419">1.0</td>
</tr>
<tr id="3578349CFF9E22AAFF1386E433003BC4" box="[153,1385,1438,1462]" gridrow="21" pageId="18" pageNumber="419">
<th id="76A95DE0FF9E22AAFF1386E434E33BC4" box="[153,650,1438,1462]" gridcol="0" gridrow="21" pageId="18" pageNumber="419">Postorbital length</th>
<td id="76A95DE0FF9E22AAFD4F86E4352D3BC4" box="[709,836,1438,1462]" gridcol="1" gridrow="21" pageId="18" pageNumber="419">50–55</td>
<td id="76A95DE0FF9E22AAFCDC86E435E63BC4" box="[854,911,1438,1462]" gridcol="2" gridrow="21" pageId="18" pageNumber="419">52.7</td>
<td id="76A95DE0FF9E22AAFC3A86E435B93BC4" box="[944,976,1438,1462]" gridcol="3" gridrow="21" pageId="18" pageNumber="419">2.0</td>
<td id="76A95DE0FF9E22AAFBA486E432FD3BC4" box="[1070,1172,1438,1462]" gridcol="4" gridrow="21" pageId="18" pageNumber="419">50–53</td>
<td id="76A95DE0FF9E22AAFB3786E4337E3BC4" box="[1213,1303,1438,1462]" gridcol="5" gridrow="21" pageId="18" pageNumber="419">51.5</td>
<td id="76A95DE0FF9E22AAFAC386E433003BC4" box="[1353,1385,1438,1462]" gridcol="6" gridrow="21" pageId="18" pageNumber="419">1.6</td>
</tr>
<tr id="3578349CFF9E22AAFF1386BC33003BAC" box="[153,1385,1478,1502]" gridrow="22" pageId="18" pageNumber="419">
<th id="76A95DE0FF9E22AAFF1386BC34E33BAC" box="[153,650,1478,1502]" gridcol="0" gridrow="22" pageId="18" pageNumber="419">Suborbital depth</th>
<td id="76A95DE0FF9E22AAFD4F86BC352D3BAC" box="[709,836,1478,1502]" gridcol="1" gridrow="22" pageId="18" pageNumber="419">12–15</td>
<td id="76A95DE0FF9E22AAFCDC86BC35E63BAC" box="[854,911,1478,1502]" gridcol="2" gridrow="22" pageId="18" pageNumber="419">13.5</td>
<td id="76A95DE0FF9E22AAFC3A86BC35B93BAC" box="[944,976,1478,1502]" gridcol="3" gridrow="22" pageId="18" pageNumber="419">1.4</td>
<td id="76A95DE0FF9E22AAFBA486BC32FD3BAC" box="[1070,1172,1478,1502]" gridcol="4" gridrow="22" pageId="18" pageNumber="419">10–12</td>
<td id="76A95DE0FF9E22AAFB3786BC337E3BAC" box="[1213,1303,1478,1502]" gridcol="5" gridrow="22" pageId="18" pageNumber="419">10.8</td>
<td id="76A95DE0FF9E22AAFAC386BC33003BAC" box="[1353,1385,1478,1502]" gridcol="6" gridrow="22" pageId="18" pageNumber="419">1.1</td>
</tr>
<tr id="3578349CFF9E22AAFF13869433003875" box="[153,1385,1518,1543]" gridrow="23" pageId="18" pageNumber="419">
<th id="76A95DE0FF9E22AAFF13869434E33875" box="[153,650,1518,1543]" gridcol="0" gridrow="23" pageId="18" pageNumber="419">Eye diameter</th>
<td id="76A95DE0FF9E22AAFD4F8694352D3875" box="[709,836,1518,1543]" gridcol="1" gridrow="23" pageId="18" pageNumber="419">27–36</td>
<td id="76A95DE0FF9E22AAFCDC869435E63875" box="[854,911,1518,1543]" gridcol="2" gridrow="23" pageId="18" pageNumber="419">32.0</td>
<td id="76A95DE0FF9E22AAFC3A869435B93875" box="[944,976,1518,1543]" gridcol="3" gridrow="23" pageId="18" pageNumber="419">3.2</td>
<td id="76A95DE0FF9E22AAFBA4869432FD3875" box="[1070,1172,1518,1543]" gridcol="4" gridrow="23" pageId="18" pageNumber="419">28–32</td>
<td id="76A95DE0FF9E22AAFB378694337E3875" box="[1213,1303,1518,1543]" gridcol="5" gridrow="23" pageId="18" pageNumber="419">29.8</td>
<td id="76A95DE0FF9E22AAFAC3869433003875" box="[1353,1385,1518,1543]" gridcol="6" gridrow="23" pageId="18" pageNumber="419">1.4</td>
</tr>
<tr id="3578349CFF9E22AAFF13856D3300385D" box="[153,1385,1559,1583]" gridrow="24" pageId="18" pageNumber="419">
<th id="76A95DE0FF9E22AAFF13856D34E3385D" box="[153,650,1559,1583]" gridcol="0" gridrow="24" pageId="18" pageNumber="419">Snout to eye end length</th>
<td id="76A95DE0FF9E22AAFD4F856D352D385D" box="[709,836,1559,1583]" gridcol="1" gridrow="24" pageId="18" pageNumber="419">45–50</td>
<td id="76A95DE0FF9E22AAFCDC856D35E6385D" box="[854,911,1559,1583]" gridcol="2" gridrow="24" pageId="18" pageNumber="419">47.3</td>
<td id="76A95DE0FF9E22AAFC3A856D35B9385D" box="[944,976,1559,1583]" gridcol="3" gridrow="24" pageId="18" pageNumber="419">2.0</td>
<td id="76A95DE0FF9E22AAFBA4856D32FD385D" box="[1070,1172,1559,1583]" gridcol="4" gridrow="24" pageId="18" pageNumber="419">47–51</td>
<td id="76A95DE0FF9E22AAFB37856D337E385D" box="[1213,1303,1559,1583]" gridcol="5" gridrow="24" pageId="18" pageNumber="419">48.5</td>
<td id="76A95DE0FF9E22AAFAC3856D3300385D" box="[1353,1385,1559,1583]" gridcol="6" gridrow="24" pageId="18" pageNumber="419">1.6</td>
</tr>
<tr id="3578349CFF9E22AAFF13854533003825" box="[153,1385,1599,1623]" gridrow="25" pageId="18" pageNumber="419">
<th id="76A95DE0FF9E22AAFF13854534E33825" box="[153,650,1599,1623]" gridcol="0" gridrow="25" pageId="18" pageNumber="419">Snout length</th>
<td id="76A95DE0FF9E22AAFD4F8545352D3825" box="[709,836,1599,1623]" gridcol="1" gridrow="25" pageId="18" pageNumber="419">20–29</td>
<td id="76A95DE0FF9E22AAFCDC854535E63825" box="[854,911,1599,1623]" gridcol="2" gridrow="25" pageId="18" pageNumber="419">24.5</td>
<td id="76A95DE0FF9E22AAFC3A854535B93825" box="[944,976,1599,1623]" gridcol="3" gridrow="25" pageId="18" pageNumber="419">3.1</td>
<td id="76A95DE0FF9E22AAFBA4854532FD3825" box="[1070,1172,1599,1623]" gridcol="4" gridrow="25" pageId="18" pageNumber="419">25–30</td>
<td id="76A95DE0FF9E22AAFB378545337E3825" box="[1213,1303,1599,1623]" gridcol="5" gridrow="25" pageId="18" pageNumber="419">27.7</td>
<td id="76A95DE0FF9E22AAFAC3854533003825" box="[1353,1385,1599,1623]" gridcol="6" gridrow="25" pageId="18" pageNumber="419">2.1</td>
</tr>
<tr id="3578349CFF9E22AAFF13851D3300380D" box="[153,1385,1639,1663]" gridrow="26" pageId="18" pageNumber="419">
<th id="76A95DE0FF9E22AAFF13851D3300380D" box="[153,1385,1639,1663]" colspan="7" colspanRight="6" gridcol="0" gridrow="26" pageId="18" pageNumber="419">
<emphasis id="B93CEACCFF9EDD47FF13851D37A9380D" bold="true" box="[153,448,1639,1663]" pageId="18" pageNumber="419">Other morphometric ratios</emphasis>
</th>
</tr>
<tr id="3578349CFF9E22AAFF1385F5330038DA" box="[153,1385,1679,1704]" gridrow="27" pageId="18" pageNumber="419">
<th id="76A95DE0FF9E22AAFF1385F534E338DA" box="[153,650,1679,1704]" gridcol="0" gridrow="27" pageId="18" pageNumber="419">Predorsal length in % of preanal length</th>
<td id="76A95DE0FF9E22AAFD4F85F5352D38DA" box="[709,836,1679,1704]" gridcol="1" gridrow="27" pageId="18" pageNumber="419">94–105</td>
<td id="76A95DE0FF9E22AAFCDC85F535E638DA" box="[854,911,1679,1704]" gridcol="2" gridrow="27" pageId="18" pageNumber="419">98.8</td>
<td id="76A95DE0FF9E22AAFC3A85F535B938DA" box="[944,976,1679,1704]" gridcol="3" gridrow="27" pageId="18" pageNumber="419">4.4</td>
<td id="76A95DE0FF9E22AAFBA485F532FD38DA" box="[1070,1172,1679,1704]" gridcol="4" gridrow="27" pageId="18" pageNumber="419">86–91</td>
<td id="76A95DE0FF9E22AAFB3785F5337E38DA" box="[1213,1303,1679,1704]" gridcol="5" gridrow="27" pageId="18" pageNumber="419">88.8</td>
<td id="76A95DE0FF9E22AAFAC385F5330038DA" box="[1353,1385,1679,1704]" gridcol="6" gridrow="27" pageId="18" pageNumber="419">2.3</td>
</tr>
<tr id="3578349CFF9E22AAFF1385C2330038A2" box="[153,1385,1720,1744]" gridrow="28" pageId="18" pageNumber="419">
<th id="76A95DE0FF9E22AAFF1385C234E338A2" box="[153,650,1720,1744]" gridcol="0" gridrow="28" pageId="18" pageNumber="419">Caudal peduncle length in % of its depth</th>
<td id="76A95DE0FF9E22AAFD4F85C2352D38A2" box="[709,836,1720,1744]" gridcol="1" gridrow="28" pageId="18" pageNumber="419">144–152</td>
<td id="76A95DE0FF9E22AAFCDC85C235E638A2" box="[854,911,1720,1744]" gridcol="2" gridrow="28" pageId="18" pageNumber="419">147.5</td>
<td id="76A95DE0FF9E22AAFC3A85C235B938A2" box="[944,976,1720,1744]" gridcol="3" gridrow="28" pageId="18" pageNumber="419">3.9</td>
<td id="76A95DE0FF9E22AAFBA485C232FD38A2" box="[1070,1172,1720,1744]" gridcol="4" gridrow="28" pageId="18" pageNumber="419">171–177</td>
<td id="76A95DE0FF9E22AAFB3785C2337E38A2" box="[1213,1303,1720,1744]" gridcol="5" gridrow="28" pageId="18" pageNumber="419">173.8</td>
<td id="76A95DE0FF9E22AAFAC385C2330038A2" box="[1353,1385,1720,1744]" gridcol="6" gridrow="28" pageId="18" pageNumber="419">3.1</td>
</tr>
<tr id="3578349CFF9E22AAFF13859A3300388A" box="[153,1385,1760,1784]" gridrow="29" pageId="18" pageNumber="419">
<th id="76A95DE0FF9E22AAFF13859A34E3388A" box="[153,650,1760,1784]" gridcol="0" gridrow="29" pageId="18" pageNumber="419">Head width in % of its depth</th>
<td id="76A95DE0FF9E22AAFD4F859A352D388A" box="[709,836,1760,1784]" gridcol="1" gridrow="29" pageId="18" pageNumber="419">64–70</td>
<td id="76A95DE0FF9E22AAFCDC859A35E6388A" box="[854,911,1760,1784]" gridcol="2" gridrow="29" pageId="18" pageNumber="419">66.3</td>
<td id="76A95DE0FF9E22AAFC3A859A35B9388A" box="[944,976,1760,1784]" gridcol="3" gridrow="29" pageId="18" pageNumber="419">2.4</td>
<td id="76A95DE0FF9E22AAFBA4859A32FD388A" box="[1070,1172,1760,1784]" gridcol="4" gridrow="29" pageId="18" pageNumber="419">74–78</td>
<td id="76A95DE0FF9E22AAFB37859A337E388A" box="[1213,1303,1760,1784]" gridcol="5" gridrow="29" pageId="18" pageNumber="419">76.1</td>
<td id="76A95DE0FF9E22AAFAC3859A3300388A" box="[1353,1385,1760,1784]" gridcol="6" gridrow="29" pageId="18" pageNumber="419">2.1</td>
</tr>
<tr id="3578349CFF9E22AAFF13847233003953" box="[153,1385,1800,1825]" gridrow="30" pageId="18" pageNumber="419" rowspan-3="1" rowspan-6="1">
<th id="76A95DE0FF9E22AAFF13847234E33953" box="[153,650,1800,1825]" gridcol="0" gridrow="30" pageId="18" pageNumber="419">
<emphasis id="B93CEACCFF9EDD47FF13847236943952" bold="true" box="[153,253,1800,1824]" pageId="18" pageNumber="419">Meristics</emphasis>
</th>
<td id="76A95DE0FF9E22AAFD4F8472352D3953" box="[709,836,1800,1825]" gridcol="1" gridrow="30" pageId="18" pageNumber="419">range</td>
<td id="76A95DE0FF9E22AAFCDC847235E63953" box="[854,911,1800,1825]" gridcol="2" gridrow="30" pageId="18" pageNumber="419">mode</td>
<td id="76A95DE0FF9E22AAFBA4847232FD3953" box="[1070,1172,1800,1825]" gridcol="4" gridrow="30" pageId="18" pageNumber="419">min–max</td>
<td id="76A95DE0FF9E22AAFB378472337E3953" box="[1213,1303,1800,1825]" gridcol="5" gridrow="30" pageId="18" pageNumber="419">mode</td>
</tr>
<tr id="3578349CFF9E22AAFF13844A3300393B" box="[153,1385,1840,1865]" gridrow="31" pageId="18" pageNumber="419" rowspan-3="1" rowspan-6="1">
<th id="76A95DE0FF9E22AAFF13844A34E3393B" box="[153,650,1840,1865]" gridcol="0" gridrow="31" pageId="18" pageNumber="419">Dorsal-fin rays</th>
<td id="76A95DE0FF9E22AAFD4F844A352D393B" box="[709,836,1840,1865]" gridcol="1" gridrow="31" pageId="18" pageNumber="419">14–16</td>
<td id="76A95DE0FF9E22AAFCDC844A35E6393B" box="[854,911,1840,1865]" gridcol="2" gridrow="31" pageId="18" pageNumber="419">16</td>
<td id="76A95DE0FF9E22AAFBA4844A32FD393B" box="[1070,1172,1840,1865]" gridcol="4" gridrow="31" pageId="18" pageNumber="419">15–17</td>
<td id="76A95DE0FF9E22AAFB37844A337E393B" box="[1213,1303,1840,1865]" gridcol="5" gridrow="31" pageId="18" pageNumber="419">15</td>
</tr>
<tr id="3578349CFF9E22AAFF13842333003903" box="[153,1385,1881,1905]" gridrow="32" pageId="18" pageNumber="419" rowspan-3="1" rowspan-6="1">
<th id="76A95DE0FF9E22AAFF13842334E33903" box="[153,650,1881,1905]" gridcol="0" gridrow="32" pageId="18" pageNumber="419">Anal-fin rays</th>
<td id="76A95DE0FF9E22AAFD4F8423352D3903" box="[709,836,1881,1905]" gridcol="1" gridrow="32" pageId="18" pageNumber="419">17–20</td>
<td id="76A95DE0FF9E22AAFCDC842335E63903" box="[854,911,1881,1905]" gridcol="2" gridrow="32" pageId="18" pageNumber="419">17</td>
<td id="76A95DE0FF9E22AAFBA4842332FD3903" box="[1070,1172,1881,1905]" gridcol="4" gridrow="32" pageId="18" pageNumber="419">17–19</td>
<td id="76A95DE0FF9E22AAFB378423337E3903" box="[1213,1303,1881,1905]" gridcol="5" gridrow="32" pageId="18" pageNumber="419">17</td>
</tr>
<tr id="3578349CFF9E22AAFF1384FB330039EB" box="[153,1385,1921,1945]" gridrow="33" pageId="18" pageNumber="419" rowspan-3="1" rowspan-6="1">
<th id="76A95DE0FF9E22AAFF1384FB34E339EB" box="[153,650,1921,1945]" gridcol="0" gridrow="33" pageId="18" pageNumber="419">Scales mid-longitudinal series</th>
<td id="76A95DE0FF9E22AAFD4F84FB352D39EB" box="[709,836,1921,1945]" gridcol="1" gridrow="33" pageId="18" pageNumber="419">28–30</td>
<td id="76A95DE0FF9E22AAFCDC84FB35E639EB" box="[854,911,1921,1945]" gridcol="2" gridrow="33" pageId="18" pageNumber="419">29</td>
<td id="76A95DE0FF9E22AAFBA484FB32FD39EB" box="[1070,1172,1921,1945]" gridcol="4" gridrow="33" pageId="18" pageNumber="419">27–29</td>
<td id="76A95DE0FF9E22AAFB3784FB337E39EB" box="[1213,1303,1921,1945]" gridcol="5" gridrow="33" pageId="18" pageNumber="419">29</td>
</tr>
<tr id="3578349CFF9E22AAFF1384D3330039B3" box="[153,1385,1961,1985]" gridrow="34" pageId="18" pageNumber="419" rowspan-3="1" rowspan-6="1">
<th id="76A95DE0FF9E22AAFF1384D334E339B3" box="[153,650,1961,1985]" gridcol="0" gridrow="34" pageId="18" pageNumber="419">Scales transverse</th>
<td id="76A95DE0FF9E22AAFD4F84D3352D39B3" box="[709,836,1961,1985]" gridcol="1" gridrow="34" pageId="18" pageNumber="419">10</td>
<td id="76A95DE0FF9E22AAFCDC84D335E639B3" box="[854,911,1961,1985]" gridcol="2" gridrow="34" pageId="18" pageNumber="419">10</td>
<td id="76A95DE0FF9E22AAFBA484D332FD39B3" box="[1070,1172,1961,1985]" gridcol="4" gridrow="34" pageId="18" pageNumber="419">10</td>
<td id="76A95DE0FF9E22AAFB3784D3337E39B3" box="[1213,1303,1961,1985]" gridcol="5" gridrow="34" pageId="18" pageNumber="419">10</td>
</tr>
<tr id="3578349CFF9E22AAFF1384AB33003998" box="[153,1385,2001,2026]" gridrow="35" pageId="18" pageNumber="419" rowspan-3="1" rowspan-6="1">
<th id="76A95DE0FF9E22AAFF1384AB34E33998" box="[153,650,2001,2026]" gridcol="0" gridrow="35" pageId="18" pageNumber="419">Scales circumpeduncular</th>
<td id="76A95DE0FF9E22AAFD4F84AB352D3998" box="[709,836,2001,2026]" gridcol="1" gridrow="35" pageId="18" pageNumber="419">12</td>
<td id="76A95DE0FF9E22AAFCDC84AB35E63998" box="[854,911,2001,2026]" gridcol="2" gridrow="35" pageId="18" pageNumber="419">12</td>
<td id="76A95DE0FF9E22AAFBA484AB32FD3998" box="[1070,1172,2001,2026]" gridcol="4" gridrow="35" pageId="18" pageNumber="419">12</td>
<td id="76A95DE0FF9E22AAFB3784AB337E3998" box="[1213,1303,2001,2026]" gridcol="5" gridrow="35" pageId="18" pageNumber="419">12</td>
</tr>
</table>
</paragraph>
<caption id="DF376656FF9FDD46FF1D86FB32E838E7" pageId="19" pageNumber="420" startId="19.[151,250,1409,1433]" targetBox="[151,1435,179,1383]" targetPageId="19" targetType="figure">
<paragraph id="8BF736DEFF9FDD46FF1D86FB32E838E7" blockId="19.[151,1437,1409,1686]" pageId="19" pageNumber="420">
<emphasis id="B93CEACCFF9FDD46FF1D86FB377D3BEB" bold="true" box="[151,276,1409,1433]" pageId="19" pageNumber="420">FIGURE 6.</emphasis>
Wild-caught specimens of
<taxonomicName id="4C484D5DFF9FDD46FDA486FB35423BEB" authorityName=", Wildekamp" authorityYear="1977" box="[558,811,1409,1433]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="19" pageNumber="420" phylum="Chordata" rank="species" species="lourensi">
<emphasis id="B93CEACCFF9FDD46FDA486FB35423BEB" box="[558,811,1409,1433]" italics="true" pageId="19" pageNumber="420">Nothobranchius lourensi</emphasis>
</taxonomicName>
and examples of typical habitat.
<emphasis id="B93CEACCFF9FDD46FB0986FB32F23BEB" bold="true" box="[1155,1179,1409,1433]" pageId="19" pageNumber="420">A.</emphasis>
<taxonomicName id="4C484D5DFF9FDD46FB2986F833F53BEB" authority="Ifakara TAN" authorityName="Ifakara TAN" box="[1187,1436,1409,1433]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="19" pageNumber="420" phylum="Chordata" rank="species" species="lourensi">
<emphasis id="B93CEACCFF9FDD46FB2986F8337D3BEB" box="[1187,1300,1409,1433]" italics="true" pageId="19" pageNumber="420">N. lourensi</emphasis>
Ifakara TAN
</taxonomicName>
95-4, male, Kilombero River drainage;
<emphasis id="B93CEACCFF9FDD46FDAB86DF34503BCF" bold="true" box="[545,569,1445,1469]" pageId="19" pageNumber="420">B.</emphasis>
<taxonomicName id="4C484D5DFF9FDD46FDB486DC356A3BCF" authority="Kitonga" authorityName="Kitonga" box="[574,771,1445,1469]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="19" pageNumber="420" phylum="Chordata" rank="species" species="lourensi">
<emphasis id="B93CEACCFF9FDD46FDB486DC34C53BCF" box="[574,684,1445,1469]" italics="true" pageId="19" pageNumber="420">N. lourensi</emphasis>
Kitonga
</taxonomicName>
north TAN 97-9, male, lower Rufiji River drainage;
<emphasis id="B93CEACCFF9FDD46FA9A86DF33413BCF" bold="true" box="[1296,1320,1445,1469]" pageId="19" pageNumber="420">C.</emphasis>
<taxonomicName id="4C484D5DFF9FDD46FAA486DC36BB3B93" authority="TSTS" authorityName="TSTS" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="19" pageNumber="420" phylum="Chordata" rank="species" species="lourensi">
<emphasis id="B93CEACCFF9FDD46FAA486DC33F53BCF" box="[1326,1436,1445,1469]" italics="true" pageId="19" pageNumber="420">N. lourensi</emphasis>
TSTS
</taxonomicName>
10-12, male, Kilombero River drainage;
<emphasis id="B93CEACCFF9FDD46FD0F86B334F43B93" bold="true" box="[645,669,1481,1505]" pageId="19" pageNumber="420">D.</emphasis>
<taxonomicName id="4C484D5DFF9FDD46FD2C86B035353B93" authority="TSTS" authorityName="TSTS" box="[678,860,1481,1505]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="19" pageNumber="420" phylum="Chordata" rank="species" species="lourensi">
<emphasis id="B93CEACCFF9FDD46FD2C86B035713B93" box="[678,792,1481,1505]" italics="true" pageId="19" pageNumber="420">N. lourensi</emphasis>
TSTS
</taxonomicName>
10-12, female, Kilombero River drainage;
<emphasis id="B93CEACCFF9FDD46FAA886B333503B93" bold="true" box="[1314,1337,1481,1505]" pageId="19" pageNumber="420">E.</emphasis>
Location Ihango TAN 02-25, a typical thickly vegetated shallow habitat on the floodplain of the Kilombero River, Kibasira Swamp (photograph June 9, 2002), hosting
<taxonomicName id="4C484D5DFF9FDD46FE7085683401385B" box="[506,616,1553,1577]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="19" pageNumber="420" phylum="Chordata" rank="species" species="lourensi">
<emphasis id="B93CEACCFF9FDD46FE7085683401385B" box="[506,616,1553,1577]" italics="true" pageId="19" pageNumber="420">N. lourensi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C484D5DFF9FDD46FD1D85683498385B" authorityName="Wildekamp, Shidlovskiy &amp; Watters" authorityYear="2009" box="[663,753,1553,1577]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="19" pageNumber="420" phylum="Chordata" rank="species" species="lucius">
<emphasis id="B93CEACCFF9FDD46FD1D85683498385B" box="[663,753,1553,1577]" italics="true" pageId="19" pageNumber="420">N. lucius</emphasis>
</taxonomicName>
;
<emphasis id="B93CEACCFF9FDD46FD76856B3566385B" bold="true" box="[764,783,1553,1577]" pageId="19" pageNumber="420">F.</emphasis>
Location Narubungo TAN 02-24, very thickly vegetated residual pool alongside a railway embankment, Kilombero River drainage, Kibasira Swamp (photograph June 9, 2002), hosting
<taxonomicName id="4C484D5DFF9FDD46FA0F854C36803803" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="19" pageNumber="420" phylum="Chordata" rank="species" species="lourensi">
<emphasis id="B93CEACCFF9FDD46FA0F854C36803803" italics="true" pageId="19" pageNumber="420">N. lourensi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C484D5DFF9FDD46FF7D852037003803" authorityName="TSTS" authorityYear="2010" box="[247,361,1626,1649]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="19" pageNumber="420" phylum="Chordata" rank="species" species="geminus">
<emphasis id="B93CEACCFF9FDD46FF7D852037003803" box="[247,361,1626,1649]" italics="true" pageId="19" pageNumber="420">N. geminus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C484D5DFF9FDD46FEFC852034453803" authorityName="Wildekamp, Watters &amp; Sainthouse" authorityYear="2002" box="[374,556,1625,1649]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="19" pageNumber="420" phylum="Chordata" rank="species" species="kilomberoensis">
<emphasis id="B93CEACCFF9FDD46FEFC852034453803" box="[374,556,1625,1649]" italics="true" pageId="19" pageNumber="420">N. kilomberoensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C484D5DFF9FDD46FDD5852034D33803" authorityName="Wildekamp, Shidlovskiy &amp; Watters" authorityYear="2009" box="[607,698,1625,1649]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="19" pageNumber="420" phylum="Chordata" rank="species" species="lucius">
<emphasis id="B93CEACCFF9FDD46FDD5852034D33803" box="[607,698,1625,1649]" italics="true" pageId="19" pageNumber="420">N. lucius</emphasis>
</taxonomicName>
. See also FIGUREs 5C and D, and FIGURE 7C for further examples of seasonal pools that host
<taxonomicName id="4C484D5DFF9FDD46FE078504379438E7" box="[397,509,1661,1685]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="19" pageNumber="420" phylum="Chordata" rank="species" species="lourensi">
<emphasis id="B93CEACCFF9FDD46FE078504379438E7" box="[397,509,1661,1685]" italics="true" pageId="19" pageNumber="420">N. lourensi</emphasis>
</taxonomicName>
. Photographs by B.R. Watters (A, B, E, F) and B. Nagy (C, D).
</paragraph>
</caption>
<paragraph id="8BF736DEFF9FDD46FF4D85B9352939E3" blockId="19.[151,1436,1731,2010]" pageId="19" pageNumber="420">
Male: Medium-sized
<taxonomicName id="4C484D5DFF9FDD46FE3F85B934AA38AF" box="[437,707,1731,1757]" pageId="19" pageNumber="420">
<emphasis id="B93CEACCFF9FDD46FE3F85B9340038AF" box="[437,617,1731,1757]" italics="true" pageId="19" pageNumber="420">Nothobranchius</emphasis>
species
</taxonomicName>
, maximum recorded size
<quantity id="4CB09B3BFF9FDD46FC5185B9325438AC" box="[987,1085,1731,1758]" metricMagnitude="-2" metricUnit="m" metricValue="4.06" pageId="19" pageNumber="420" unit="mm" value="40.6">40.6 mm</quantity>
SL. General body shape robust, laterally compressed and deep. Greatest body depth at pelvic-fin origin, 28.6–34.0 % SL. Greatest body width at pectoral-fin base with body progressively narrowing towards caudal-fin base. Dorsal profile convex from tip of snout to dorsal-fin origin, straight to slightly concave on caudal peduncle. Ventral profile convex from lower jaw to base of last anal-fin ray, straight to slightly concave on caudal peduncle. Caudal peduncle deep, length 144–152 % of its depth. Anus situated directly in front of anal-fin origin.
</paragraph>
<paragraph id="8BF736DEFF9FDD41FF4D84E132CF3F6E" blockId="19.[151,1436,1731,2010]" lastBlockId="20.[151,1437,151,2036]" lastPageId="20" lastPageNumber="421" pageId="19" pageNumber="420">Head moderately long (29.2–32.8 % SL), laterally compressed, deeper than wide (head width 64–70 % of its depth). Snout distinctly pointed, smaller than eye diameter. Mouth supraterminal, slightly oblique in profile. Jaws subequal, lower jaw longer than upper, posterior end of rictus at same level or slightly ventral to centre of eye. Premaxilla and dentary with many irregularly distributed conical teeth on inner rows and slightly curved teeth on outer row. Orbit moderately large, eye diameter 27–36 % HL, positioned entirely in anterior half of head (snout to eye end length 45–50 % HL). Branchiostegal membrane projecting posteriorly from opercle.</paragraph>
<paragraph id="8BF736DEFF98DD41FF4D825F351F3C2C" blockId="20.[151,1437,151,2036]" pageId="20" pageNumber="421">Dorsal-fin origin at about same vertical positioning as anal-fin origin (mean predorsal length 94–105 as % of preanal length), both fins originating slightly posterior to mid-length of body (predorsal length 52.3–56.2 % SL; preanal length 53.7–57.6 % SL). Dorsal fin with 14–16 rays, extremity rounded. Anal fin anterior margin and anterior part of lower margin rounded, posterior part of lower margin straight to mildly concave, angular to posterior margin, with 17–20 rays. Dorsal and anal fins with short fin-ray extensions. Pectoral fin subtriangular, insertion slightly posterior to margin of opercular opening, base slightly oblique, upper fin rays positioned slightly anterior to lower fin rays, with tip reaching or slightly overlapping base of pelvic fin. Pelvic fin subabdominal, origin at about mid-length of body, short, base medially separated, with tip reaching urogenital papilla. Caudal fin rounded, with 14–18 branched rays, plus 3–4 dorsal and ventral procurrent rays.</paragraph>
<paragraph id="8BF736DEFF98DD41FF4D811D32273CBB" blockId="20.[151,1437,151,2036]" pageId="20" pageNumber="421">Scales cycloid, body and head entirely scaled, except for ventral surface of head. No scales on base of dorsal and anal fins. Scales in mid-longitudinal series 28–30 plus two or three small scales on caudal-fin base. Transverse rows of scales in front of dorsal-fin origin 10; scale rows around caudal peduncle 12.</paragraph>
<paragraph id="8BF736DEFF98DD41FF4D81A837043D94" blockId="20.[151,1437,151,2036]" pageId="20" pageNumber="421">
Cephalic squamation pattern G-type. Two frontal neuromasts behind upper lip, posterior in shallow groove. Cephalic sensory system at supraorbital level in divided shallow groove with one and three exposed neuromasts, in anterior and posterior portions, respectively (
<figureCitation id="13732A5BFF98DD41FD00806034B53D46" box="[650,732,794,820]" captionStart="FIGURE 3" captionStartId="10.[151,250,1307,1331]" captionTargetBox="[156,1428,706,1276]" captionTargetId="figure-280@10.[151,1436,702,1281]" captionTargetPageId="10" captionText="FIGURE 3. Cephalic structure: a, Nothobranchius janpapi; b, N. luekei; c, N. geminus; d, N. lourensi and N. fuscotaeniatus. Key to label abbreviations: an, anterior nostril; pn, posterior nostril; f, frontal neuromast; so, supraorbital canal; st, supratemporal canal." pageId="20" pageNumber="421">Fig. 3d</figureCitation>
), whereas at supratemporal level with four exposed neuromasts in undivided, slightly curved, shallow groove. Preorbital canal in shallow groove with three exposed neuromasts; infraorbital level with series of about dozen small buttons at ventral and posterior margin of orbit; postorbital canal in deep groove with one exposed neuromast. Preopercular system with seven exposed neuromasts in deep groove. Mandibular canal in shallow groove with three small neuromasts. One neuromast on each scale along trunk mid-longitudinal series.
</paragraph>
<paragraph id="8BF736DEFF98DD41FF4D808A33E63AEB" blockId="20.[151,1437,151,2036]" pageId="20" pageNumber="421">
Female: Slightly smaller than male, maximum recorded size
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SL. Branchiostegal membrane not projecting posteriorly from opercle. Dorsal and caudal fins rounded, former with 15–17 rays. Anal fin subtriangular, tip rounded, central rays longer and more rigid. Anal fin positioned more posteriorly than in males (preanal length 61.7–67.1 % SL vs. 53.7–57.6), and base length smaller than in male (18.6–21.4 % SL vs. 23.4–26.7). Head width larger than in males (59–63 % HL vs. 54–59) whereas head depth smaller than in males (79–81 % HL vs. 83–87).
</paragraph>
<paragraph id="8BF736DEFF98DD40FF4D87D937D73E8B" blockId="20.[151,1437,151,2036]" lastBlockId="21.[151,1437,151,969]" lastPageId="21" lastPageNumber="422" pageId="20" pageNumber="421">
<emphasis id="B93CEACCFF98DD41FF4D87D937363ACF" bold="true" box="[199,351,1187,1213]" pageId="20" pageNumber="421">Colouration.</emphasis>
Live male (
<figureCitation id="13732A5BFF98DD41FE6D87D934063ACF" box="[487,623,1187,1213]" captionStart="FIGURE 6" captionStartId="19.[151,250,1409,1433]" captionTargetBox="[151,1435,179,1383]" captionTargetId="figure-19@19.[151,1435,179,1383]" captionTargetPageId="19" captionText="FIGURE 6. Wild-caught specimens of Nothobranchius lourensi and examples of typical habitat. A. N. lourensi Ifakara TAN 95-4, male, Kilombero River drainage; B. N. lourensi Kitonga north TAN 97-9, male, lower Rufiji River drainage; C. N. lourensi TSTS 10-12, male, Kilombero River drainage; D. N. lourensi TSTS 10-12, female, Kilombero River drainage; E. Location Ihango TAN 02-25, a typical thickly vegetated shallow habitat on the floodplain of the Kilombero River, Kibasira Swamp (photograph June 9, 2002), hosting N. lourensi and N. lucius; F. Location Narubungo TAN 02-24, very thickly vegetated residual pool alongside a railway embankment, Kilombero River drainage, Kibasira Swamp (photograph June 9, 2002), hosting N. lourensi, N. geminus, N. kilomberoensis and N. lucius. See also FIGUREs 5C and D, and FIGURE 7C for further examples of seasonal pools that host N. lourensi. Photographs by B.R. Watters (A, B, E, F) and B. Nagy (C, D)." pageId="20" pageNumber="421">Figs. 6A–C</figureCitation>
): Scales on trunk and head are iridescent blue to blue-green with narrow brown to red-brown margins, forming a regular reticulation pattern with, on the posterior part of the body, a weakly developed diagonally-barred or chevron pattern. Scales on abdomen light blue to white with less pronounced margins that are paler red-brown to yellow-brown in colour. Scales on the dorsum anterior to dorsal fin have broader margins and may have yellow-tinged centres. Snout, frontal and dorsal portions of the head are mainly brown although some populations show red lips; the throat is pale blue to blue-green with a yellow tinge. Operculum has light blue to blue-green scale centres with red-brown margins in the upper part, grading to yellow margins in the lower part. Exposed part of branchiostegal membrane is yellow to white. Iris is blue to blue-green. The dorsal fin background is iridescent yellow-brown to golden colour and, in some populations, may show a red tinge in the basal and proximal zones and some blue green colour in the distal zone. The background is overlaid by dark red-brown to maroon spots that are arranged in such a way as to coalesce in part, forming irregular concentric bands or, in some specimens, lines parallel to the fin rays; the latter linear arrangement is more prominent in the basal and proximal zones but may extend through the medial zone. A narrow dark red to maroon marginal band is invariably present and some specimens also show light blue tips to short fin-ray extensions. The anal fin background comprises a light blue to grey basal zone grading to blue-green, yellow-green or yellow in the remainder of the fin. The background is overlaid by 5–7 dark maroon bands, the most distal of which forms a marginal band. The innermost bands in the proximal and submedial zones may be relatively indistinct and short, where the others are distinct and prominent extending fully across the fin in a pattern ranging from straight to curved, more or less parallel to the distal margin. Background colour of the caudal fin is variable: it can be yellow in the basal, proximal and medial zones grading to a lighter yellow or blue in the subdistal zone; or blue-green or blue in the basal, proximal and medial zones, grading to blue in the subdistal zone; or blue or yellow throughout the basal to subdistal zones. The background is overlaid by 4–5 concentric bars conforming to the general outer shape of the fin comprising, essentially, rows of coalescent dark maroon spots. There is a distinct black marginal band most commonly preceded by a submarginal zone relatively free of maroon markings and which appears as a distinct band showing the background colour. Pelvic fins mimic, in a simpler fashion, the colour pattern of the anal fin with maroon markings, arranged in a crude banding, on a yellow, blue-green or blue background. Pectoral fins are dominantly hyaline with poorly developed, and usually only partial, blue margins.
</paragraph>
<paragraph id="8BF736DEFF99DD40FF4D827935693FFB" blockId="21.[151,1437,151,969]" pageId="21" pageNumber="422">
Live female (
<figureCitation id="13732A5BFF99DD40FEE8827937D63F6F" box="[354,447,259,285]" captionStart="FIGURE 6" captionStartId="19.[151,250,1409,1433]" captionTargetBox="[151,1435,179,1383]" captionTargetId="figure-19@19.[151,1435,179,1383]" captionTargetPageId="19" captionText="FIGURE 6. Wild-caught specimens of Nothobranchius lourensi and examples of typical habitat. A. N. lourensi Ifakara TAN 95-4, male, Kilombero River drainage; B. N. lourensi Kitonga north TAN 97-9, male, lower Rufiji River drainage; C. N. lourensi TSTS 10-12, male, Kilombero River drainage; D. N. lourensi TSTS 10-12, female, Kilombero River drainage; E. Location Ihango TAN 02-25, a typical thickly vegetated shallow habitat on the floodplain of the Kilombero River, Kibasira Swamp (photograph June 9, 2002), hosting N. lourensi and N. lucius; F. Location Narubungo TAN 02-24, very thickly vegetated residual pool alongside a railway embankment, Kilombero River drainage, Kibasira Swamp (photograph June 9, 2002), hosting N. lourensi, N. geminus, N. kilomberoensis and N. lucius. See also FIGUREs 5C and D, and FIGURE 7C for further examples of seasonal pools that host N. lourensi. Photographs by B.R. Watters (A, B, E, F) and B. Nagy (C, D)." pageId="21" pageNumber="422">Fig. 6D</figureCitation>
): Trunk and head light grey, darker on the dorsum and lighter ventrally. Scales on the flanks have narrow darker grey-brown margins which may form a faint reticulated pattern on the posterior part of the body; scale centres on the trunk above the abdomen are iridescent pale blue; the operculum is iridescent light blue or yellow. All fins hyaline. The iris is pale golden.
</paragraph>
</subSubSection>
<subSubSection id="C3526555FF99DD40FF4D82E834623DBB" pageId="21" pageNumber="422" type="distribution">
<paragraph id="8BF736DEFF99DD40FF4D82E8337E3CBF" blockId="21.[151,1437,151,969]" pageId="21" pageNumber="422">
<emphasis id="B93CEACCFF99DD40FF4D82E837083FDE" bold="true" box="[199,353,402,428]" pageId="21" pageNumber="422">Distribution.</emphasis>
<taxonomicName id="4C484D5DFF99DD40FEE082E934EB3FDF" box="[362,642,403,429]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="21" pageNumber="422" phylum="Chordata" rank="species" species="lourensi">
<emphasis id="B93CEACCFF99DD40FEE082E934EB3FDF" box="[362,642,403,429]" italics="true" pageId="21" pageNumber="422">Nothobranchius lourensi</emphasis>
</taxonomicName>
is endemic to seasonal freshwater habitats in coastal
<collectingCountry id="F35F764EFF99DD40FB6082E933263FDF" box="[1258,1359,403,429]" name="Tanzania" pageId="21" pageNumber="422">Tanzania</collectingCountry>
and is the most widespread member of the
<taxonomicName id="4C484D5DFF99DD40FDC882CD34933FA3" authorityName="Wildekamp" authorityYear="1977" box="[578,762,439,465]" class="Actinopterygii" family="Nothobranchiidae" genus="Aphyobranchius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyprinodontiformes" pageId="21" pageNumber="422" phylum="Chordata" rank="genus">
<emphasis id="B93CEACCFF99DD40FDC882CD34933FA3" box="[578,762,439,465]" italics="true" pageId="21" pageNumber="422">Aphyobranchius</emphasis>
</taxonomicName>
subgenus, occurring most abundantly in the marginal and peripheral zones of the Kibasira Swamp in the Kilombero River valley but relatively sparsely elsewhere (
<figureCitation id="13732A5BFF99DD40FAC282A133E23F87" box="[1352,1419,475,501]" captionStart="FIGURE 1" captionStartId="2.[151,250,1762,1786]" captionTargetBox="[151,1436,374,1736]" captionTargetId="figure-118@2.[151,1436,374,1736]" captionTargetPageId="2" captionText="FIGURE 1. Distribution of species belonging to the subgenus Aphyobranchius, all of which occur in the east-central and southern coastal lowlands of Tanzania: N. janpapi (yellow-filled circle), N. luekei (red-filled triangle), N. geminus (orange-filled diamond), N. lourensi (green-filled square), N. fuscotaeniatus (blue-filled inverted triangle), N. cf. janpapi (dark magenta-filled circle). The inset map shows the area of distribution in a somewhat broader geographic context, to include the distribution of N. willerti (red-filled star) in the lower Tana River area of northeastern coastal Kenya, a species that herein is considered to belong to the subgenus Adiniops, rather than Aphyobranchius to which it was ascribed in the original description by Wildekamp (1992). Note that individual symbols on the map may represent more than one locality or population where the sites are close to one another and could not reasonably be resolved at the scale of the map." pageId="21" pageNumber="422">Fig. 1</figureCitation>
). The Kilombero system represents a fairly remote component of the Rufiji River system, connected to it through the Ulanga River, the downstream continuation of the Kilombero River. Relatively rare, isolated, single site occurrences have also been reported from: the lower Ruvu River near Kwaraza, representing the
<typeStatus id="54F3887CFF99DD40FBCD813D321E3C13" box="[1095,1143,583,609]" pageId="21" pageNumber="422">type</typeStatus>
location; a couple of sites in the lower Rufiji River system, one associated with the Ruhoi River and another situated
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south of the main course of the Rufiji River near where it approaches the delta; a single occurrence is also known from the Luhule/ Luhute River floodplain within the so-called Mbezi Triangle (
<emphasis id="B93CEACCFF99DD40FCC781C935E23CBE" box="[845,907,691,716]" italics="true" pageId="21" pageNumber="422">sensu</emphasis>
<bibRefCitation id="EFD94B2FFF99DD40FC1881C9320E3CBF" author="Watters, B. R. &amp; Cooper, B. J. &amp; Wildekamp, R. H." box="[914,1127,691,717]" pageId="21" pageNumber="422" pagination="5 - 6" refId="ref35078" refString="Watters, B. R., Cooper, B. J. &amp; Wildekamp, R. H. (2007) Description of Nothobranchius cardinalis spec. nov. (Cyprinodontiformes: Aplocheilidae), an annual fish from the Mbwemkuru River basin, Tanzania. Journal of the American Killifish Association, 40 (5 - 6), 129 - 145. [issue printed in 2008 = year of species description]" type="journal article" year="2007">
Watters
<emphasis id="B93CEACCFF99DD40FC6781C932763CBF" box="[1005,1055,691,717]" italics="true" pageId="21" pageNumber="422">et al</emphasis>
. 2007
</bibRefCitation>
;
<bibRefCitation id="EFD94B2FFF99DD40FBF981C933623CBF" author="Watters, B. R." box="[1139,1291,691,717]" pageId="21" pageNumber="422" pagination="37 - 76" refId="ref34941" refString="Watters, B. R. (2009) The ecology and distribution of Nothobranchius fishes. Journal of the American Killifish Association, 42 (2), 37 - 76." type="journal article" year="2009">Watters 2009</bibRefCitation>
).
</paragraph>
<paragraph id="8BF736DEFF99DD40FF4D81AC34623DBB" blockId="21.[151,1437,151,969]" pageId="21" pageNumber="422">
<emphasis id="B93CEACCFF99DD40FF4D81AC37A83C83" bold="true" box="[199,449,726,753]" pageId="21" pageNumber="422">Conservation status.</emphasis>
<taxonomicName id="4C484D5DFF99DD40FE4481AD34833C83" box="[462,746,727,753]" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="21" pageNumber="422" phylum="Chordata" rank="species" species="lourensi">
<emphasis id="B93CEACCFF99DD40FE4481AD34833C83" box="[462,746,727,753]" italics="true" pageId="21" pageNumber="422">Nothobranchius lourensi</emphasis>
</taxonomicName>
has been evaluated as Near Threatened (
<bibRefCitation id="EFD94B2FFF99DD40FB5081AD368B3D67" author="Nagy, B. &amp; Watters, B." pageId="21" pageNumber="422" refId="ref33758" refString="Nagy, B. &amp; Watters, B. (2019 d) Nothobranchius lourensi. The IUCN Red List of Threatened Species, 2019, e. T 61288 A 47243786. https: // doi. org / 10.2305 / IUCN. UK. 2019 - 3. RLTS. T 61288 A 47243786. en" type="journal volume" year="2019">Nagy &amp; Watters 2019d</bibRefCitation>
). The known extent of occurrence (EOO) is
<quantity id="4CB09B3BFF99DD40FD68818135363D65" box="[738,863,763,791]" metricMagnitude="7" metricUnit="m" metricValue="2.8006" pageId="21" pageNumber="422" unit="km" value="28006.0">28,006 km</quantity>
². Within its area of distribution, the occurrence of the species is currently known from more than 10 threat-based locations, with disjunct populations in several river drainages. There is increasing habitat degradation due primarily to expansion of agriculture in the area, including the conversion of some seasonal habitats into permanent ponds (e.g. in the Kwaraza area of the lower Ruvu River system, where the
<typeStatus id="54F3887CFF99DD40FEED80F137FE3DD7" box="[359,407,907,933]" pageId="21" pageNumber="422">type</typeStatus>
locality occurs), thereby modifying them in ways that render them unsuitable to support the seasonal life cycle of the species.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="D6C1967CCA3F60E9055BD268DC5293D9">Description of two new species of Rhaphidophora (Orthoptera: Rhaphidophoridae: Rhaphidophorinae) from Yunnan, China</mods:title>
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<mods:namePart id="58BD5E9275FC8C8AA9F8A727E07B1ACD">Yu, Tingting</mods:namePart>
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<mods:affiliation id="73E24075FC43396569D80FF4EC26C960">College of Life Sciences &amp; Technology, Inner Mongolia Normal University, Hohhot 010022, China.</mods:affiliation>
<mods:nameIdentifier id="50CD54CD442E64BF4963C598B8ACEC3E" type="email">yutingtingnmg@outlook.com</mods:nameIdentifier>
<mods:namePart id="9D20D415E773AD9B09217B41E31F3E16">Yu, Tingting</mods:namePart>
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<mods:namePart id="D9C56B80189E5EC756CC63C8CD3068DE">Liu, Mingyu</mods:namePart>
<mods:affiliation id="BC32BE9D1C60553F92B4AD81DB85CF97">College of Agricultural and Life Sciences, Kunming University, Kunming 650214, China.</mods:affiliation>
<mods:namePart id="FE7DD828F3CA53824DE525397AFEEB34">Liu, Mingyu</mods:namePart>
<mods:affiliation id="6D44CDB8B77EEE7D020AB23B0DCD393B">College of Agricultural and Life Sciences, Kunming University, Kunming 650214, China.</mods:affiliation>
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<mods:namePart id="023A814393E1E2A29E58179410AAD5D6">Bian, Xun</mods:namePart>
<mods:affiliation id="7DD6636A6B20568408E9878E8D7DBAE3">Key Laboratory of Ecology of Rare and Endangered Species and Environmental Protection (Guangxi Normal University), Ministry of Education, Guilin 541006, China.</mods:affiliation>
<mods:namePart id="4D477FEC0D9B0F2508F7006811F14DFA">Bian, Xun</mods:namePart>
<mods:affiliation id="673E8FECAD2AA408D0928F710B599D05">Key Laboratory of Ecology of Rare and Endangered Species and Environmental Protection (Guangxi Normal University), Ministry of Education, Guilin 541006, China.</mods:affiliation>
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<mods:namePart id="F924D0ACE209477E39143A899F486101">Zhang, Bin</mods:namePart>
<mods:affiliation id="BFA1331AFE860E30C27AFE75C5BC659C">College of Life Sciences &amp; Technology, Inner Mongolia Normal University, Hohhot 010022, China.</mods:affiliation>
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<heading id="D0A32A35FFD0B102FF6088A8FDF7FA34" bold="true" box="[151,595,1410,1437]" fontSize="11" level="1" pageId="3" pageNumber="397" reason="1">
@ -83,7 +86,7 @@
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<paragraph id="8BEB9D59FFD0B102FF6088E1FEB8FA4C" blockId="3.[151,595,1410,1509]" box="[151,284,1483,1509]" pageId="3" pageNumber="397">
<figureCitation id="136F81DCFFD0B102FF6088E1FEB8FA4C" box="[151,284,1483,1509]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="4.[151,250,1327,1351]" captionStartId-1="5.[151,250,1795,1819]" captionTargetBox-0="[151,1436,179,1300]" captionTargetBox-1="[151,1436,597,1772]" captionTargetId-0="figure-18@4.[151,1436,179,1300]" captionTargetId-1="figure-202@5.[151,1436,597,1772]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 3. Rhaphidophora minutispina sp. nov. Male: A. head in frontal view; B–C. head and thoraces: B. dorsal view, C. lateral view; D–E. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; G–K. apex of abdomen: G. apical view, H. apico-lateral view, I. lateral view, J–K. ventral view." captionText-1="FIGURE 4. Rhaphidophora minutispina sp. nov. Female: A. head in frontal view; B–C. head and thorax: B. dorsal view, C. lateral view; D–E. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; G–H. apex of abdomen: G. dorsal view, H. lateral view; I. subgenital plate in ventral view; J. seventh abdominal sternite and subgenital plate in ventral view." pageId="3" pageNumber="397">Figures 3–4</figureCitation>
<figureCitation id="136F81DCFFD0B102FF6088E1FEB8FA4C" box="[151,284,1483,1509]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="4.[151,250,1327,1351]" captionStartId-1="5.[151,250,1795,1819]" captionTargetBox-0="[151,1436,179,1300]" captionTargetBox-1="[151,1436,597,1772]" captionTargetId-0="figure-18@4.[151,1436,179,1300]" captionTargetId-1="figure-202@5.[151,1436,597,1772]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 3. Rhaphidophora minutispina sp. nov. Male: A. head in frontal view; B–C. head and thoraces: B. dorsal view, C. lateral view; D–E. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; G–K. apex of abdomen: G. apical view, H. apico-lateral view, I. lateral view, J–K. ventral view." captionText-1="FIGURE 4. Rhaphidophora minutispina sp. nov. Female: A. head in frontal view; B–C. head and thorax: B. dorsal view, C. lateral view; D–E. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; G–H. apex of abdomen: G. dorsal view, H. lateral view; I. subgenital plate in ventral view; J. seventh abdominal sternite and subgenital plate in ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.14734845" figureDoi-1="http://doi.org/10.5281/zenodo.14734847" httpUri-0="https://zenodo.org/record/14734845/files/figure.png" httpUri-1="https://zenodo.org/record/14734847/files/figure.png" pageId="3" pageNumber="397">Figures 3–4</figureCitation>
</paragraph>
<paragraph id="8BEB9D59FFD0B102FF608B38FB66F8AC" blockId="3.[151,1437,1554,2013]" pageId="3" pageNumber="397">
<emphasis id="B920414BFFD0B102FF608B38FED7F984" bold="true" box="[151,371,1554,1581]" pageId="3" pageNumber="397">Description. Male.</emphasis>
@ -92,26 +95,26 @@ Body medium.
Head and pronotum as other species of the genus.
<emphasis id="B920414BFFD0B102FB528B39FB45F985" box="[1189,1249,1555,1580]" italics="true" pageId="3" pageNumber="397">Legs.</emphasis>
Fore coxae with 1 spine; femora with 1 short ventral spine on internal margin; tibiae ventrally with 1 internal and 2 external spines, apices with 1 pair of spines on ventral surface. Middle femora with 1 pair of apical spines; tibiae dorsally with 2 pairs of spines, ventral surface with 1–3 pairs of spines, apices with 1 pair of spines on dorsal and ventral surfaces separately. Hind femora with 1 short internal spine (
<figureCitation id="136F81DCFFD0B102FD018B89FCF4F914" box="[758,848,1699,1725]" captionStart="FIGURE 3" captionStartId="4.[151,250,1327,1351]" captionTargetBox="[151,1436,179,1300]" captionTargetId="figure-18@4.[151,1436,179,1300]" captionTargetPageId="4" captionText="FIGURE 3. Rhaphidophora minutispina sp. nov. Male: A. head in frontal view; B–C. head and thoraces: B. dorsal view, C. lateral view; D–E. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; G–K. apex of abdomen: G. apical view, H. apico-lateral view, I. lateral view, J–K. ventral view." pageId="3" pageNumber="397">Fig. 3E</figureCitation>
<figureCitation id="136F81DCFFD0B102FD018B89FCF4F914" box="[758,848,1699,1725]" captionStart="FIGURE 3" captionStartId="4.[151,250,1327,1351]" captionTargetBox="[151,1436,179,1300]" captionTargetId="figure-18@4.[151,1436,179,1300]" captionTargetPageId="4" captionText="FIGURE 3. Rhaphidophora minutispina sp. nov. Male: A. head in frontal view; B–C. head and thoraces: B. dorsal view, C. lateral view; D–E. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; G–K. apex of abdomen: G. apical view, H. apico-lateral view, I. lateral view, J–K. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734845" httpUri="https://zenodo.org/record/14734845/files/figure.png" pageId="3" pageNumber="397">Fig. 3E</figureCitation>
); tibiae dorsally with 19–25 internal spines, 21–23 external spines, and 1 pair of small subapical spines, apices with 1 pair of dorsal spines and 2 pairs of ventral spines; basitarsi with 4–6 small spines and 1 apical spine on dorsal surface along the midline (
<figureCitation id="136F81DCFFD0B102FB978BC1FB11F8AC" box="[1120,1205,1771,1797]" captionStart="FIGURE 3" captionStartId="4.[151,250,1327,1351]" captionTargetBox="[151,1436,179,1300]" captionTargetId="figure-18@4.[151,1436,179,1300]" captionTargetPageId="4" captionText="FIGURE 3. Rhaphidophora minutispina sp. nov. Male: A. head in frontal view; B–C. head and thoraces: B. dorsal view, C. lateral view; D–E. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; G–K. apex of abdomen: G. apical view, H. apico-lateral view, I. lateral view, J–K. ventral view." pageId="3" pageNumber="397">Fig. 3F</figureCitation>
<figureCitation id="136F81DCFFD0B102FB978BC1FB11F8AC" box="[1120,1205,1771,1797]" captionStart="FIGURE 3" captionStartId="4.[151,250,1327,1351]" captionTargetBox="[151,1436,179,1300]" captionTargetId="figure-18@4.[151,1436,179,1300]" captionTargetPageId="4" captionText="FIGURE 3. Rhaphidophora minutispina sp. nov. Male: A. head in frontal view; B–C. head and thoraces: B. dorsal view, C. lateral view; D–E. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; G–K. apex of abdomen: G. apical view, H. apico-lateral view, I. lateral view, J–K. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734845" httpUri="https://zenodo.org/record/14734845/files/figure.png" pageId="3" pageNumber="397">Fig. 3F</figureCitation>
).
</paragraph>
<paragraph id="8BEB9D59FFD0B102FF308A25FAE8F874" blockId="3.[151,1437,1554,2013]" pageId="3" pageNumber="397">
<emphasis id="B920414BFFD0B102FF308A25FE9DF880" box="[199,313,1807,1833]" italics="true" pageId="3" pageNumber="397">Abdomen.</emphasis>
Posterior margin of tenth abdominal tergite faintly concave in middle. Dorsal surface of epiproct with obviously raised lateral margins, between them concave (
<figureCitation id="136F81DCFFD0B102FCD28A19FCD9F8E4" box="[805,893,1843,1869]" captionStart="FIGURE 3" captionStartId="4.[151,250,1327,1351]" captionTargetBox="[151,1436,179,1300]" captionTargetId="figure-18@4.[151,1436,179,1300]" captionTargetPageId="4" captionText="FIGURE 3. Rhaphidophora minutispina sp. nov. Male: A. head in frontal view; B–C. head and thoraces: B. dorsal view, C. lateral view; D–E. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; G–K. apex of abdomen: G. apical view, H. apico-lateral view, I. lateral view, J–K. ventral view." pageId="3" pageNumber="397">Fig. 3H</figureCitation>
<figureCitation id="136F81DCFFD0B102FCD28A19FCD9F8E4" box="[805,893,1843,1869]" captionStart="FIGURE 3" captionStartId="4.[151,250,1327,1351]" captionTargetBox="[151,1436,179,1300]" captionTargetId="figure-18@4.[151,1436,179,1300]" captionTargetPageId="4" captionText="FIGURE 3. Rhaphidophora minutispina sp. nov. Male: A. head in frontal view; B–C. head and thoraces: B. dorsal view, C. lateral view; D–E. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; G–K. apex of abdomen: G. apical view, H. apico-lateral view, I. lateral view, J–K. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734845" httpUri="https://zenodo.org/record/14734845/files/figure.png" pageId="3" pageNumber="397">Fig. 3H</figureCitation>
), and then prolonged into a pair of wide spines (
<figureCitation id="136F81DCFFD0B102FF688A7DFF53F8D8" box="[159,247,1879,1905]" captionStart="FIGURE 3" captionStartId="4.[151,250,1327,1351]" captionTargetBox="[151,1436,179,1300]" captionTargetId="figure-18@4.[151,1436,179,1300]" captionTargetPageId="4" captionText="FIGURE 3. Rhaphidophora minutispina sp. nov. Male: A. head in frontal view; B–C. head and thoraces: B. dorsal view, C. lateral view; D–E. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; G–K. apex of abdomen: G. apical view, H. apico-lateral view, I. lateral view, J–K. ventral view." pageId="3" pageNumber="397">Fig. 3G</figureCitation>
<figureCitation id="136F81DCFFD0B102FF688A7DFF53F8D8" box="[159,247,1879,1905]" captionStart="FIGURE 3" captionStartId="4.[151,250,1327,1351]" captionTargetBox="[151,1436,179,1300]" captionTargetId="figure-18@4.[151,1436,179,1300]" captionTargetPageId="4" captionText="FIGURE 3. Rhaphidophora minutispina sp. nov. Male: A. head in frontal view; B–C. head and thoraces: B. dorsal view, C. lateral view; D–E. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; G–K. apex of abdomen: G. apical view, H. apico-lateral view, I. lateral view, J–K. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734845" httpUri="https://zenodo.org/record/14734845/files/figure.png" pageId="3" pageNumber="397">Fig. 3G</figureCitation>
), the internal margin of the spine with 1 minute spine about the middle area (
<figureCitation id="136F81DCFFD0B102FB918A7DFB19F8D8" box="[1126,1213,1879,1905]" captionStart="FIGURE 3" captionStartId="4.[151,250,1327,1351]" captionTargetBox="[151,1436,179,1300]" captionTargetId="figure-18@4.[151,1436,179,1300]" captionTargetPageId="4" captionText="FIGURE 3. Rhaphidophora minutispina sp. nov. Male: A. head in frontal view; B–C. head and thoraces: B. dorsal view, C. lateral view; D–E. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; G–K. apex of abdomen: G. apical view, H. apico-lateral view, I. lateral view, J–K. ventral view." pageId="3" pageNumber="397">Fig. 3H</figureCitation>
<figureCitation id="136F81DCFFD0B102FB918A7DFB19F8D8" box="[1126,1213,1879,1905]" captionStart="FIGURE 3" captionStartId="4.[151,250,1327,1351]" captionTargetBox="[151,1436,179,1300]" captionTargetId="figure-18@4.[151,1436,179,1300]" captionTargetPageId="4" captionText="FIGURE 3. Rhaphidophora minutispina sp. nov. Male: A. head in frontal view; B–C. head and thoraces: B. dorsal view, C. lateral view; D–E. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; G–K. apex of abdomen: G. apical view, H. apico-lateral view, I. lateral view, J–K. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734845" httpUri="https://zenodo.org/record/14734845/files/figure.png" pageId="3" pageNumber="397">Fig. 3H</figureCitation>
); ventral surface of epiproct transverse, posterolateral area with 1 thin spine on each side, its apices directing forward and acute (
<figureCitation id="136F81DCFFD0B102FA988A51FF15F810" captionStart="FIGURE 3" captionStartId="4.[151,250,1327,1351]" captionTargetBox="[151,1436,179,1300]" captionTargetId="figure-18@4.[151,1436,179,1300]" captionTargetPageId="4" captionText="FIGURE 3. Rhaphidophora minutispina sp. nov. Male: A. head in frontal view; B–C. head and thoraces: B. dorsal view, C. lateral view; D–E. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; G–K. apex of abdomen: G. apical view, H. apico-lateral view, I. lateral view, J–K. ventral view." pageId="3" pageNumber="397">Fig. 3I</figureCitation>
<figureCitation id="136F81DCFFD0B102FA988A51FF15F810" captionStart="FIGURE 3" captionStartId="4.[151,250,1327,1351]" captionTargetBox="[151,1436,179,1300]" captionTargetId="figure-18@4.[151,1436,179,1300]" captionTargetPageId="4" captionText="FIGURE 3. Rhaphidophora minutispina sp. nov. Male: A. head in frontal view; B–C. head and thoraces: B. dorsal view, C. lateral view; D–E. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; G–K. apex of abdomen: G. apical view, H. apico-lateral view, I. lateral view, J–K. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734845" httpUri="https://zenodo.org/record/14734845/files/figure.png" pageId="3" pageNumber="397">Fig. 3I</figureCitation>
). Subgenital plate wider than long, lateral margins convex, posterior margin truncate; styli conical, about two times as long as subgenital plate, inserted on the posterolateral areas of the plate (
<figureCitation id="136F81DCFFD0B102FBDF8AE9FB26F874" box="[1064,1154,1987,2013]" captionStart="FIGURE 3" captionStartId="4.[151,250,1327,1351]" captionTargetBox="[151,1436,179,1300]" captionTargetId="figure-18@4.[151,1436,179,1300]" captionTargetPageId="4" captionText="FIGURE 3. Rhaphidophora minutispina sp. nov. Male: A. head in frontal view; B–C. head and thoraces: B. dorsal view, C. lateral view; D–E. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; G–K. apex of abdomen: G. apical view, H. apico-lateral view, I. lateral view, J–K. ventral view." pageId="3" pageNumber="397">Fig. 3K</figureCitation>
<figureCitation id="136F81DCFFD0B102FBDF8AE9FB26F874" box="[1064,1154,1987,2013]" captionStart="FIGURE 3" captionStartId="4.[151,250,1327,1351]" captionTargetBox="[151,1436,179,1300]" captionTargetId="figure-18@4.[151,1436,179,1300]" captionTargetPageId="4" captionText="FIGURE 3. Rhaphidophora minutispina sp. nov. Male: A. head in frontal view; B–C. head and thoraces: B. dorsal view, C. lateral view; D–E. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; G–K. apex of abdomen: G. apical view, H. apico-lateral view, I. lateral view, J–K. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734845" httpUri="https://zenodo.org/record/14734845/files/figure.png" pageId="3" pageNumber="397">Fig. 3K</figureCitation>
), apices subacute.
</paragraph>
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<paragraph id="8BEB9D59FFD7B105FF608805FC7CFA26" blockId="4.[151,1436,1326,1423]" pageId="4" pageNumber="398">
<emphasis id="B920414BFFD7B105FF608805FCA4FAEE" bold="true" box="[151,768,1326,1351]" pageId="4" pageNumber="398">
FIGURE 3.
@ -151,15 +154,15 @@ ventral view.
<paragraph id="8BEB9D59FFD7B105FF3088CBFD49F922" blockId="4.[151,1436,1505,2036]" pageId="4" pageNumber="398">
<emphasis id="B920414BFFD7B105FF3088CBFE81FA52" bold="true" box="[199,293,1505,1531]" pageId="4" pageNumber="398">Female.</emphasis>
Tenth abdominal tergite concave in middle (
<figureCitation id="136F81DCFFD7B105FCCE88CBFC37FA52" box="[825,915,1505,1531]" captionStart="FIGURE 4" captionStartId="5.[151,250,1795,1819]" captionTargetBox="[151,1436,597,1772]" captionTargetId="figure-202@5.[151,1436,597,1772]" captionTargetPageId="5" captionText="FIGURE 4. Rhaphidophora minutispina sp. nov. Female: A. head in frontal view; B–C. head and thorax: B. dorsal view, C. lateral view; D–E. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; G–H. apex of abdomen: G. dorsal view, H. lateral view; I. subgenital plate in ventral view; J. seventh abdominal sternite and subgenital plate in ventral view." pageId="4" pageNumber="398">Fig. 4G</figureCitation>
<figureCitation id="136F81DCFFD7B105FCCE88CBFC37FA52" box="[825,915,1505,1531]" captionStart="FIGURE 4" captionStartId="5.[151,250,1795,1819]" captionTargetBox="[151,1436,597,1772]" captionTargetId="figure-202@5.[151,1436,597,1772]" captionTargetPageId="5" captionText="FIGURE 4. Rhaphidophora minutispina sp. nov. Female: A. head in frontal view; B–C. head and thorax: B. dorsal view, C. lateral view; D–E. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; G–H. apex of abdomen: G. dorsal view, H. lateral view; I. subgenital plate in ventral view; J. seventh abdominal sternite and subgenital plate in ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734847" httpUri="https://zenodo.org/record/14734847/files/figure.png" pageId="4" pageNumber="398">Fig. 4G</figureCitation>
). Epiproct roughly triangular, dorsal surface concave along middle, posterior margin rounded (
<figureCitation id="136F81DCFFD7B105FD4B8B2FFCB5F9B6" box="[700,785,1541,1567]" captionStart="FIGURE 4" captionStartId="5.[151,250,1795,1819]" captionTargetBox="[151,1436,597,1772]" captionTargetId="figure-202@5.[151,1436,597,1772]" captionTargetPageId="5" captionText="FIGURE 4. Rhaphidophora minutispina sp. nov. Female: A. head in frontal view; B–C. head and thorax: B. dorsal view, C. lateral view; D–E. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; G–H. apex of abdomen: G. dorsal view, H. lateral view; I. subgenital plate in ventral view; J. seventh abdominal sternite and subgenital plate in ventral view." pageId="4" pageNumber="398">Fig. 4G</figureCitation>
<figureCitation id="136F81DCFFD7B105FD4B8B2FFCB5F9B6" box="[700,785,1541,1567]" captionStart="FIGURE 4" captionStartId="5.[151,250,1795,1819]" captionTargetBox="[151,1436,597,1772]" captionTargetId="figure-202@5.[151,1436,597,1772]" captionTargetPageId="5" captionText="FIGURE 4. Rhaphidophora minutispina sp. nov. Female: A. head in frontal view; B–C. head and thorax: B. dorsal view, C. lateral view; D–E. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; G–H. apex of abdomen: G. dorsal view, H. lateral view; I. subgenital plate in ventral view; J. seventh abdominal sternite and subgenital plate in ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734847" httpUri="https://zenodo.org/record/14734847/files/figure.png" pageId="4" pageNumber="398">Fig. 4G</figureCitation>
). Cerci slender. Seventh abdominal sternite longer than the preceding sternites. Subgenital plate semi-circular, separated by a prolonged membranous zone from the preceding sternite (
<figureCitation id="136F81DCFFD7B105FF0B8B67FEE9F9CE" box="[252,333,1613,1639]" captionStart="FIGURE 4" captionStartId="5.[151,250,1795,1819]" captionTargetBox="[151,1436,597,1772]" captionTargetId="figure-202@5.[151,1436,597,1772]" captionTargetPageId="5" captionText="FIGURE 4. Rhaphidophora minutispina sp. nov. Female: A. head in frontal view; B–C. head and thorax: B. dorsal view, C. lateral view; D–E. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; G–H. apex of abdomen: G. dorsal view, H. lateral view; I. subgenital plate in ventral view; J. seventh abdominal sternite and subgenital plate in ventral view." pageId="4" pageNumber="398">Fig. 4I</figureCitation>
<figureCitation id="136F81DCFFD7B105FF0B8B67FEE9F9CE" box="[252,333,1613,1639]" captionStart="FIGURE 4" captionStartId="5.[151,250,1795,1819]" captionTargetBox="[151,1436,597,1772]" captionTargetId="figure-202@5.[151,1436,597,1772]" captionTargetPageId="5" captionText="FIGURE 4. Rhaphidophora minutispina sp. nov. Female: A. head in frontal view; B–C. head and thorax: B. dorsal view, C. lateral view; D–E. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; G–H. apex of abdomen: G. dorsal view, H. lateral view; I. subgenital plate in ventral view; J. seventh abdominal sternite and subgenital plate in ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734847" httpUri="https://zenodo.org/record/14734847/files/figure.png" pageId="4" pageNumber="398">Fig. 4I</figureCitation>
), posterior margin with 1 median concavity (
<figureCitation id="136F81DCFFD7B105FCAE8B67FC08F9CE" box="[857,940,1613,1639]" captionStart="FIGURE 4" captionStartId="5.[151,250,1795,1819]" captionTargetBox="[151,1436,597,1772]" captionTargetId="figure-202@5.[151,1436,597,1772]" captionTargetPageId="5" captionText="FIGURE 4. Rhaphidophora minutispina sp. nov. Female: A. head in frontal view; B–C. head and thorax: B. dorsal view, C. lateral view; D–E. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; G–H. apex of abdomen: G. dorsal view, H. lateral view; I. subgenital plate in ventral view; J. seventh abdominal sternite and subgenital plate in ventral view." pageId="4" pageNumber="398">Fig. 4J</figureCitation>
<figureCitation id="136F81DCFFD7B105FCAE8B67FC08F9CE" box="[857,940,1613,1639]" captionStart="FIGURE 4" captionStartId="5.[151,250,1795,1819]" captionTargetBox="[151,1436,597,1772]" captionTargetId="figure-202@5.[151,1436,597,1772]" captionTargetPageId="5" captionText="FIGURE 4. Rhaphidophora minutispina sp. nov. Female: A. head in frontal view; B–C. head and thorax: B. dorsal view, C. lateral view; D–E. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; G–H. apex of abdomen: G. dorsal view, H. lateral view; I. subgenital plate in ventral view; J. seventh abdominal sternite and subgenital plate in ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734847" httpUri="https://zenodo.org/record/14734847/files/figure.png" pageId="4" pageNumber="398">Fig. 4J</figureCitation>
). Ovipositor about as long as hind femora, gradually upcurved behind the middle area (
<figureCitation id="136F81DCFFD7B105FD718B5BFD79F925" box="[646,733,1649,1676]" captionStart="FIGURE 4" captionStartId="5.[151,250,1795,1819]" captionTargetBox="[151,1436,597,1772]" captionTargetId="figure-202@5.[151,1436,597,1772]" captionTargetPageId="5" captionText="FIGURE 4. Rhaphidophora minutispina sp. nov. Female: A. head in frontal view; B–C. head and thorax: B. dorsal view, C. lateral view; D–E. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; G–H. apex of abdomen: G. dorsal view, H. lateral view; I. subgenital plate in ventral view; J. seventh abdominal sternite and subgenital plate in ventral view." pageId="4" pageNumber="398">Fig. 4H</figureCitation>
<figureCitation id="136F81DCFFD7B105FD718B5BFD79F925" box="[646,733,1649,1676]" captionStart="FIGURE 4" captionStartId="5.[151,250,1795,1819]" captionTargetBox="[151,1436,597,1772]" captionTargetId="figure-202@5.[151,1436,597,1772]" captionTargetPageId="5" captionText="FIGURE 4. Rhaphidophora minutispina sp. nov. Female: A. head in frontal view; B–C. head and thorax: B. dorsal view, C. lateral view; D–E. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; G–H. apex of abdomen: G. dorsal view, H. lateral view; I. subgenital plate in ventral view; J. seventh abdominal sternite and subgenital plate in ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734847" httpUri="https://zenodo.org/record/14734847/files/figure.png" pageId="4" pageNumber="398">Fig. 4H</figureCitation>
).
</paragraph>
<paragraph id="8BEB9D59FFD7B105FF308BBFFA9BF906" blockId="4.[151,1436,1505,2036]" box="[199,1343,1685,1711]" pageId="4" pageNumber="398">
@ -435,9 +438,9 @@ The new species differs from
Shen, Wang &amp; He, 2024
</taxonomicName>
by the internal margin of male epiproct bearing a minuscule spine on the upper plate (
<figureCitation id="136F81DCFFD6B104FC4C8CB9FBB7FE04" box="[955,1043,403,429]" captionStart="FIGURE 3" captionStartId="4.[151,250,1327,1351]" captionTargetBox="[151,1436,179,1300]" captionTargetId="figure-18@4.[151,1436,179,1300]" captionTargetPageId="4" captionText="FIGURE 3. Rhaphidophora minutispina sp. nov. Male: A. head in frontal view; B–C. head and thoraces: B. dorsal view, C. lateral view; D–E. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; G–K. apex of abdomen: G. apical view, H. apico-lateral view, I. lateral view, J–K. ventral view." pageId="5" pageNumber="399">Fig. 3G</figureCitation>
<figureCitation id="136F81DCFFD6B104FC4C8CB9FBB7FE04" box="[955,1043,403,429]" captionStart="FIGURE 3" captionStartId="4.[151,250,1327,1351]" captionTargetBox="[151,1436,179,1300]" captionTargetId="figure-18@4.[151,1436,179,1300]" captionTargetPageId="4" captionText="FIGURE 3. Rhaphidophora minutispina sp. nov. Male: A. head in frontal view; B–C. head and thoraces: B. dorsal view, C. lateral view; D–E. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; G–K. apex of abdomen: G. apical view, H. apico-lateral view, I. lateral view, J–K. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734845" httpUri="https://zenodo.org/record/14734845/files/figure.png" pageId="5" pageNumber="399">Fig. 3G</figureCitation>
), and male subgenital plate with 1 pair of longer and subconical styli (
<figureCitation id="136F81DCFFD6B104FDD38C9DFDDAFE78" box="[548,638,439,465]" captionStart="FIGURE 3" captionStartId="4.[151,250,1327,1351]" captionTargetBox="[151,1436,179,1300]" captionTargetId="figure-18@4.[151,1436,179,1300]" captionTargetPageId="4" captionText="FIGURE 3. Rhaphidophora minutispina sp. nov. Male: A. head in frontal view; B–C. head and thoraces: B. dorsal view, C. lateral view; D–E. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; G–K. apex of abdomen: G. apical view, H. apico-lateral view, I. lateral view, J–K. ventral view." pageId="5" pageNumber="399">Fig. 3K</figureCitation>
<figureCitation id="136F81DCFFD6B104FDD38C9DFDDAFE78" box="[548,638,439,465]" captionStart="FIGURE 3" captionStartId="4.[151,250,1327,1351]" captionTargetBox="[151,1436,179,1300]" captionTargetId="figure-18@4.[151,1436,179,1300]" captionTargetPageId="4" captionText="FIGURE 3. Rhaphidophora minutispina sp. nov. Male: A. head in frontal view; B–C. head and thoraces: B. dorsal view, C. lateral view; D–E. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; G–K. apex of abdomen: G. apical view, H. apico-lateral view, I. lateral view, J–K. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734845" httpUri="https://zenodo.org/record/14734845/files/figure.png" pageId="5" pageNumber="399">Fig. 3K</figureCitation>
).
</paragraph>
</subSubSection>

147
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<paragraph id="8BEB9D59FFD3B101FF608AF4FEB8F851" blockId="0.[151,616,1941,2040]" box="[151,284,2014,2040]" pageId="0" pageNumber="394">
<figureCitation id="136F81DCFFD3B101FF608AF4FEB8F851" box="[151,284,2014,2040]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="1.[151,250,1896,1920]" captionStartId-1="2.[151,250,1516,1540]" captionTargetBox-0="[158,1430,179,1870]" captionTargetBox-1="[151,1436,181,1492]" captionTargetId-0="figure-27@1.[158,1430,179,1870]" captionTargetId-1="figure-18@2.[151,1436,181,1492]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; E–F. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; H–L. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." captionText-1="FIGURE 2. Rhaphidophora condoprocera sp. nov. Female: A. head in frontal view; B–C. head and thorax: B. dorsal view, C. lateral view; D. apices of hind femora in internal view; E. tarsus of hind leg in lateral view; F–G. apex of abdomen: F. dorsal view, G. lateral view; H. seventh abdominal sternite and subgenital plate in ventral view." pageId="0" pageNumber="394">Figures 1–2</figureCitation>
<figureCitation id="136F81DCFFD3B101FF608AF4FEB8F851" box="[151,284,2014,2040]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="1.[151,250,1896,1920]" captionStartId-1="2.[151,250,1516,1540]" captionTargetBox-0="[158,1430,179,1870]" captionTargetBox-1="[151,1436,181,1492]" captionTargetId-0="figure-27@1.[158,1430,179,1870]" captionTargetId-1="figure-18@2.[151,1436,181,1492]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; E–F. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; H–L. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." captionText-1="FIGURE 2. Rhaphidophora condoprocera sp. nov. Female: A. head in frontal view; B–C. head and thorax: B. dorsal view, C. lateral view; D. apices of hind femora in internal view; E. tarsus of hind leg in lateral view; F–G. apex of abdomen: F. dorsal view, G. lateral view; H. seventh abdominal sternite and subgenital plate in ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.14734837" figureDoi-1="http://doi.org/10.5281/zenodo.14734841" httpUri-0="https://zenodo.org/record/14734837/files/figure.png" httpUri-1="https://zenodo.org/record/14734841/files/figure.png" pageId="0" pageNumber="394">Figures 1–2</figureCitation>
</paragraph>
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FIGURE 1.
@ -126,7 +129,7 @@ ventral and slightly lateral view,
ventral view.
</paragraph>
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FIGURE 2.
@ -162,57 +165,57 @@ seventh abdominal sternite and subgenital plate in ventral view.
Body medium.
<emphasis id="B920414BFFD1B103FDDA8BB4FDCEF911" box="[557,618,1694,1720]" italics="true" pageId="2" pageNumber="396">Head</emphasis>
. Face smooth (
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<figureCitation id="136F81DCFFD1B103FCEE8BB4FCD0F911" box="[793,884,1694,1720]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; E–F. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; H–L. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734837" httpUri="https://zenodo.org/record/14734837/files/figure.png" pageId="2" pageNumber="396">Fig. 1A</figureCitation>
). Fastigium verticis projecting forward in lateral view (
<figureCitation id="136F81DCFFD1B103FF178BE8FE9EF975" box="[224,314,1730,1756]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; E–F. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; H–L. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." pageId="2" pageNumber="396">Fig. 1B</figureCitation>
<figureCitation id="136F81DCFFD1B103FF178BE8FE9EF975" box="[224,314,1730,1756]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; E–F. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; H–L. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734837" httpUri="https://zenodo.org/record/14734837/files/figure.png" pageId="2" pageNumber="396">Fig. 1B</figureCitation>
), dorsal surface with a longitudinal median furrow, apices which divide fastigium verticis into 2 distinct rostral tubercles and slightly separated from each other (
<figureCitation id="136F81DCFFD1B103FC9C8BCCFC66F8A9" box="[875,962,1766,1792]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; E–F. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; H–L. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." pageId="2" pageNumber="396">Fig. 1B</figureCitation>
<figureCitation id="136F81DCFFD1B103FC9C8BCCFC66F8A9" box="[875,962,1766,1792]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; E–F. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; H–L. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734837" httpUri="https://zenodo.org/record/14734837/files/figure.png" pageId="2" pageNumber="396">Fig. 1B</figureCitation>
). Eyes ovoid, protruding forward; median ocellus slightly larger than lateral ocelli (
<figureCitation id="136F81DCFFD1B103FD9C8A20FD62F88D" box="[619,710,1802,1828]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; E–F. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; H–L. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." pageId="2" pageNumber="396">Fig. 1A</figureCitation>
<figureCitation id="136F81DCFFD1B103FD9C8A20FD62F88D" box="[619,710,1802,1828]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; E–F. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; H–L. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734837" httpUri="https://zenodo.org/record/14734837/files/figure.png" pageId="2" pageNumber="396">Fig. 1A</figureCitation>
); lateral ocelli large, nearly occupying all the lateral margins of rostral tubercles (
<figureCitation id="136F81DCFFD1B103FE968A04FE1EF8E1" box="[353,442,1838,1864]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; E–F. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; H–L. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." pageId="2" pageNumber="396">Fig. 1B</figureCitation>
<figureCitation id="136F81DCFFD1B103FE968A04FE1EF8E1" box="[353,442,1838,1864]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; E–F. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; H–L. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734837" httpUri="https://zenodo.org/record/14734837/files/figure.png" pageId="2" pageNumber="396">Fig. 1B</figureCitation>
). Apical segments of maxillary palpi about two times as long as subapical ones (
<figureCitation id="136F81DCFFD1B103FA988A04FF1DF8C5" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; E–F. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; H–L. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." pageId="2" pageNumber="396">Fig. 1B</figureCitation>
<figureCitation id="136F81DCFFD1B103FA988A04FF1DF8C5" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; E–F. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; H–L. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734837" httpUri="https://zenodo.org/record/14734837/files/figure.png" pageId="2" pageNumber="396">Fig. 1B</figureCitation>
).
</paragraph>
<paragraph id="8BEB9D59FFD1B102FF308A5CFC80FECC" blockId="2.[151,1437,1694,2009]" lastBlockId="3.[151,1437,151,1329]" lastPageId="3" lastPageNumber="397" pageId="2" pageNumber="396">
<emphasis id="B920414BFFD1B103FF308A5CFE84F839" box="[199,288,1910,1936]" italics="true" pageId="2" pageNumber="396">Thorax.</emphasis>
Anterior margin of pronotum slightly projecting, posterior margin widely rounded, protruding backward (
<figureCitation id="136F81DCFFD1B103FF688AB0FF57F81D" box="[159,243,1946,1972]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; E–F. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; H–L. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." pageId="2" pageNumber="396">Fig. 1C</figureCitation>
<figureCitation id="136F81DCFFD1B103FF688AB0FF57F81D" box="[159,243,1946,1972]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; E–F. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; H–L. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734837" httpUri="https://zenodo.org/record/14734837/files/figure.png" pageId="2" pageNumber="396">Fig. 1C</figureCitation>
); lateral lobes longer than high, ventral margin undulate (
<figureCitation id="136F81DCFFD1B103FCA88AB0FC12F81D" box="[863,950,1946,1972]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; E–F. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; H–L. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." pageId="2" pageNumber="396">Fig. 1D</figureCitation>
<figureCitation id="136F81DCFFD1B103FCA88AB0FC12F81D" box="[863,950,1946,1972]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; E–F. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; H–L. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734837" httpUri="https://zenodo.org/record/14734837/files/figure.png" pageId="2" pageNumber="396">Fig. 1D</figureCitation>
). Posterior margin of mesonotum protruding backward, posterior margin of metanotum faintly protruding.
<emphasis id="B920414BFFD1B103FCB88A95FC2FF871" box="[847,907,1983,2008]" italics="true" pageId="2" pageNumber="396">Legs.</emphasis>
Fore coxae swollen with 1 stout spine; femora with 1 internal spine; tibiae ventrally with 1 internal and 1–2 external spines, apices with 1 pair of spines on ventral surface. Middle femora with 1 pair of apical spines; tibiae dorsally with 2 pairs of spines, ventral surface with 1–3 pairs of spines, apices with 1 pair of spines on dorsal and ventral surfaces separately. Apices of hind femora with 1 short internal spine (
<figureCitation id="136F81DCFFD0B102FE888C29FE70FEB4" box="[383,468,259,285]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; E–F. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; H–L. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." pageId="3" pageNumber="397">Fig. 1E</figureCitation>
<figureCitation id="136F81DCFFD0B102FE888C29FE70FEB4" box="[383,468,259,285]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; E–F. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; H–L. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734837" httpUri="https://zenodo.org/record/14734837/files/figure.png" pageId="3" pageNumber="397">Fig. 1E</figureCitation>
); tibiae dorsally with 21–23 internal spines, 19–25 external spines and 1 pair of small subapical spines, apices with 1 pair of dorsal and 2 pairs of ventral spines; basitarsi with 4–6 small spines and 1 large apical spine on dorsal surface along the midline (
<figureCitation id="136F81DCFFD0B102FD4A8C61FCB7FECC" box="[701,787,331,357]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; E–F. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; H–L. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." pageId="3" pageNumber="397">Fig. 1G</figureCitation>
<figureCitation id="136F81DCFFD0B102FD4A8C61FCB7FECC" box="[701,787,331,357]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; E–F. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; H–L. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734837" httpUri="https://zenodo.org/record/14734837/files/figure.png" pageId="3" pageNumber="397">Fig. 1G</figureCitation>
).
</paragraph>
<paragraph id="8BEB9D59FFD0B102FF308C45FCA9FDC8" blockId="3.[151,1437,151,1329]" pageId="3" pageNumber="397">
<emphasis id="B920414BFFD0B102FF308C45FE9DFE20" box="[199,313,367,393]" italics="true" pageId="3" pageNumber="397">Abdomen.</emphasis>
Apex of abdomen curving downwards. Posterior margin of ninth abdominal tergite concave, tenth abdominal tergite narrow. Epiproct prolonged and gradually curving downward, basal half with lateral margins convex and converging, apical area slightly widened, bilobate in dorsal view (
<figureCitation id="136F81DCFFD0B102FBE68C9DFBCDFE78" box="[1041,1129,439,465]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; E–F. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; H–L. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." pageId="3" pageNumber="397">Fig. 1H</figureCitation>
<figureCitation id="136F81DCFFD0B102FBE68C9DFBCDFE78" box="[1041,1129,439,465]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; E–F. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; H–L. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734837" httpUri="https://zenodo.org/record/14734837/files/figure.png" pageId="3" pageNumber="397">Fig. 1H</figureCitation>
), its ventral surface with 1 depressed process (
<figureCitation id="136F81DCFFD0B102FE838CF1FE6FFE5C" box="[372,459,475,501]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; E–F. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; H–L. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." pageId="3" pageNumber="397">Fig. 1L</figureCitation>
<figureCitation id="136F81DCFFD0B102FE838CF1FE6FFE5C" box="[372,459,475,501]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; E–F. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; H–L. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734837" httpUri="https://zenodo.org/record/14734837/files/figure.png" pageId="3" pageNumber="397">Fig. 1L</figureCitation>
); ventral surface of epiproct with 1 pair of stout spines, its apices directing outwards and faintly curved forward (
<figureCitation id="136F81DCFFD0B102FE228CD5FD8BFDB0" box="[469,559,511,537]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; E–F. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; H–L. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." pageId="3" pageNumber="397">Fig. 1K</figureCitation>
<figureCitation id="136F81DCFFD0B102FE228CD5FD8BFDB0" box="[469,559,511,537]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; E–F. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; H–L. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734837" httpUri="https://zenodo.org/record/14734837/files/figure.png" pageId="3" pageNumber="397">Fig. 1K</figureCitation>
). Paraprocts prolonged into a spine, the apical half incurved. Subgenital plate wider than long, posterior margin almost straight; styli gradually upcurved before the middle (
<figureCitation id="136F81DCFFD0B102FB4B8F09FAACFD94" box="[1212,1288,547,573]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; E–F. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; H–L. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." pageId="3" pageNumber="397">Fig. 1I</figureCitation>
<figureCitation id="136F81DCFFD0B102FB4B8F09FAACFD94" box="[1212,1288,547,573]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; E–F. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; H–L. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734837" httpUri="https://zenodo.org/record/14734837/files/figure.png" pageId="3" pageNumber="397">Fig. 1I</figureCitation>
), inserted on posterolateral areas of the plate (
<figureCitation id="136F81DCFFD0B102FDF38F6DFDFDFDC8" box="[516,601,583,609]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; E–F. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; H–L. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." pageId="3" pageNumber="397">Fig. 1L</figureCitation>
<figureCitation id="136F81DCFFD0B102FDF38F6DFDFDFDC8" box="[516,601,583,609]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; E–F. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; H–L. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734837" httpUri="https://zenodo.org/record/14734837/files/figure.png" pageId="3" pageNumber="397">Fig. 1L</figureCitation>
), apices obtuse.
</paragraph>
<paragraph id="8BEB9D59FFD0B102FF308F41FCB7FD58" blockId="3.[151,1437,151,1329]" pageId="3" pageNumber="397">
<emphasis id="B920414BFFD0B102FF308F41FE81FD2C" bold="true" box="[199,293,619,645]" pageId="3" pageNumber="397">Female.</emphasis>
Tenth abdominal tergite with posterior margin concave in middle (
<figureCitation id="136F81DCFFD0B102FBDF8F41FBDBFD2C" box="[1064,1151,619,645]" captionStart="FIGURE 2" captionStartId="2.[151,250,1516,1540]" captionTargetBox="[151,1436,181,1492]" captionTargetId="figure-18@2.[151,1436,181,1492]" captionTargetPageId="2" captionText="FIGURE 2. Rhaphidophora condoprocera sp. nov. Female: A. head in frontal view; B–C. head and thorax: B. dorsal view, C. lateral view; D. apices of hind femora in internal view; E. tarsus of hind leg in lateral view; F–G. apex of abdomen: F. dorsal view, G. lateral view; H. seventh abdominal sternite and subgenital plate in ventral view." pageId="3" pageNumber="397">Fig. 2F</figureCitation>
<figureCitation id="136F81DCFFD0B102FBDF8F41FBDBFD2C" box="[1064,1151,619,645]" captionStart="FIGURE 2" captionStartId="2.[151,250,1516,1540]" captionTargetBox="[151,1436,181,1492]" captionTargetId="figure-18@2.[151,1436,181,1492]" captionTargetPageId="2" captionText="FIGURE 2. Rhaphidophora condoprocera sp. nov. Female: A. head in frontal view; B–C. head and thorax: B. dorsal view, C. lateral view; D. apices of hind femora in internal view; E. tarsus of hind leg in lateral view; F–G. apex of abdomen: F. dorsal view, G. lateral view; H. seventh abdominal sternite and subgenital plate in ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734841" httpUri="https://zenodo.org/record/14734841/files/figure.png" pageId="3" pageNumber="397">Fig. 2F</figureCitation>
). Epiproct semielliptical, dorsal surface furrowed along the midline, posterior margin obtuse (
<figureCitation id="136F81DCFFD0B102FC358FA5FBB9FD00" box="[962,1053,655,681]" captionStart="FIGURE 2" captionStartId="2.[151,250,1516,1540]" captionTargetBox="[151,1436,181,1492]" captionTargetId="figure-18@2.[151,1436,181,1492]" captionTargetPageId="2" captionText="FIGURE 2. Rhaphidophora condoprocera sp. nov. Female: A. head in frontal view; B–C. head and thorax: B. dorsal view, C. lateral view; D. apices of hind femora in internal view; E. tarsus of hind leg in lateral view; F–G. apex of abdomen: F. dorsal view, G. lateral view; H. seventh abdominal sternite and subgenital plate in ventral view." pageId="3" pageNumber="397">Fig. 2F</figureCitation>
<figureCitation id="136F81DCFFD0B102FC358FA5FBB9FD00" box="[962,1053,655,681]" captionStart="FIGURE 2" captionStartId="2.[151,250,1516,1540]" captionTargetBox="[151,1436,181,1492]" captionTargetId="figure-18@2.[151,1436,181,1492]" captionTargetPageId="2" captionText="FIGURE 2. Rhaphidophora condoprocera sp. nov. Female: A. head in frontal view; B–C. head and thorax: B. dorsal view, C. lateral view; D. apices of hind femora in internal view; E. tarsus of hind leg in lateral view; F–G. apex of abdomen: F. dorsal view, G. lateral view; H. seventh abdominal sternite and subgenital plate in ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734841" httpUri="https://zenodo.org/record/14734841/files/figure.png" pageId="3" pageNumber="397">Fig. 2F</figureCitation>
). Cerci slender. Subgenital plate transverse, posterior margin undulating and faintly projecting in middle (
<figureCitation id="136F81DCFFD0B102FC3E8F99FBBBFD64" box="[969,1055,691,717]" captionStart="FIGURE 2" captionStartId="2.[151,250,1516,1540]" captionTargetBox="[151,1436,181,1492]" captionTargetId="figure-18@2.[151,1436,181,1492]" captionTargetPageId="2" captionText="FIGURE 2. Rhaphidophora condoprocera sp. nov. Female: A. head in frontal view; B–C. head and thorax: B. dorsal view, C. lateral view; D. apices of hind femora in internal view; E. tarsus of hind leg in lateral view; F–G. apex of abdomen: F. dorsal view, G. lateral view; H. seventh abdominal sternite and subgenital plate in ventral view." pageId="3" pageNumber="397">Fig. 2H</figureCitation>
<figureCitation id="136F81DCFFD0B102FC3E8F99FBBBFD64" box="[969,1055,691,717]" captionStart="FIGURE 2" captionStartId="2.[151,250,1516,1540]" captionTargetBox="[151,1436,181,1492]" captionTargetId="figure-18@2.[151,1436,181,1492]" captionTargetPageId="2" captionText="FIGURE 2. Rhaphidophora condoprocera sp. nov. Female: A. head in frontal view; B–C. head and thorax: B. dorsal view, C. lateral view; D. apices of hind femora in internal view; E. tarsus of hind leg in lateral view; F–G. apex of abdomen: F. dorsal view, G. lateral view; H. seventh abdominal sternite and subgenital plate in ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734841" httpUri="https://zenodo.org/record/14734841/files/figure.png" pageId="3" pageNumber="397">Fig. 2H</figureCitation>
). Ovipositor about half as long as hind femora, gradually upcurved, apices acute (
<figureCitation id="136F81DCFFD0B102FD5B8FFDFCA6FD58" box="[684,770,727,753]" captionStart="FIGURE 2" captionStartId="2.[151,250,1516,1540]" captionTargetBox="[151,1436,181,1492]" captionTargetId="figure-18@2.[151,1436,181,1492]" captionTargetPageId="2" captionText="FIGURE 2. Rhaphidophora condoprocera sp. nov. Female: A. head in frontal view; B–C. head and thorax: B. dorsal view, C. lateral view; D. apices of hind femora in internal view; E. tarsus of hind leg in lateral view; F–G. apex of abdomen: F. dorsal view, G. lateral view; H. seventh abdominal sternite and subgenital plate in ventral view." pageId="3" pageNumber="397">Fig. 2G</figureCitation>
<figureCitation id="136F81DCFFD0B102FD5B8FFDFCA6FD58" box="[684,770,727,753]" captionStart="FIGURE 2" captionStartId="2.[151,250,1516,1540]" captionTargetBox="[151,1436,181,1492]" captionTargetId="figure-18@2.[151,1436,181,1492]" captionTargetPageId="2" captionText="FIGURE 2. Rhaphidophora condoprocera sp. nov. Female: A. head in frontal view; B–C. head and thorax: B. dorsal view, C. lateral view; D. apices of hind femora in internal view; E. tarsus of hind leg in lateral view; F–G. apex of abdomen: F. dorsal view, G. lateral view; H. seventh abdominal sternite and subgenital plate in ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734841" httpUri="https://zenodo.org/record/14734841/files/figure.png" pageId="3" pageNumber="397">Fig. 2G</figureCitation>
).
</paragraph>
<paragraph id="8BEB9D59FFD0B102FF308FD0FBB1FC90" blockId="3.[151,1437,151,1329]" pageId="3" pageNumber="397">
@ -341,9 +344,9 @@ The new species differs from other species of
<emphasis id="B920414BFFD0B102FC628949FBE1FBD4" box="[917,1093,1123,1149]" italics="true" pageId="3" pageNumber="397">Rhaphidophora</emphasis>
</taxonomicName>
by male epiproct prolonged downwards and backwards apical area with three swellings: two dorsad and one ventrad, apical margin of each swelling undulant (
<figureCitation id="136F81DCFFD0B102FE868981FE1AFB6C" box="[369,446,1195,1221]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; E–F. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; H–L. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." pageId="3" pageNumber="397">Fig. 1I</figureCitation>
<figureCitation id="136F81DCFFD0B102FE868981FE1AFB6C" box="[369,446,1195,1221]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; E–F. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; H–L. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734837" httpUri="https://zenodo.org/record/14734837/files/figure.png" pageId="3" pageNumber="397">Fig. 1I</figureCitation>
), ventral surface of epiproct with 1 pair of stout and widely separated spines; posterior margin of female subgenital plate faintly projecting in middle (
<figureCitation id="136F81DCFFD0B102FCAE89E5FC0BFB40" box="[857,943,1231,1257]" captionStart="FIGURE 2" captionStartId="2.[151,250,1516,1540]" captionTargetBox="[151,1436,181,1492]" captionTargetId="figure-18@2.[151,1436,181,1492]" captionTargetPageId="2" captionText="FIGURE 2. Rhaphidophora condoprocera sp. nov. Female: A. head in frontal view; B–C. head and thorax: B. dorsal view, C. lateral view; D. apices of hind femora in internal view; E. tarsus of hind leg in lateral view; F–G. apex of abdomen: F. dorsal view, G. lateral view; H. seventh abdominal sternite and subgenital plate in ventral view." pageId="3" pageNumber="397">Fig. 2H</figureCitation>
<figureCitation id="136F81DCFFD0B102FCAE89E5FC0BFB40" box="[857,943,1231,1257]" captionStart="FIGURE 2" captionStartId="2.[151,250,1516,1540]" captionTargetBox="[151,1436,181,1492]" captionTargetId="figure-18@2.[151,1436,181,1492]" captionTargetPageId="2" captionText="FIGURE 2. Rhaphidophora condoprocera sp. nov. Female: A. head in frontal view; B–C. head and thorax: B. dorsal view, C. lateral view; D. apices of hind femora in internal view; E. tarsus of hind leg in lateral view; F–G. apex of abdomen: F. dorsal view, G. lateral view; H. seventh abdominal sternite and subgenital plate in ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734841" httpUri="https://zenodo.org/record/14734841/files/figure.png" pageId="3" pageNumber="397">Fig. 2H</figureCitation>
).
</paragraph>
</subSubSection>