Add updates up until 2024-11-04 10:07:55

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ggserver 2024-11-04 10:12:01 +00:00
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<mods:title id="B26364BD6C8A28023F7C64362489ADA5">Five new species and numerous locality records of Spirobolida millipedes from Madagascar (Diplopoda, Spirobolida, Pachybolidae)</mods:title>
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(deSaussure &amp; Zehntner, 1897)
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. 1 M,
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; 1 M
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, general collecting, coll.
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<emphasis id="B949D804FFC6FFAFFCF62F63D2D1FC18" box="[851,905,871,897]" italics="true" pageId="18" pageNumber="479">et al.</emphasis>
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; 1 F
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,
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,
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,
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near
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,
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, coll.
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,
<collectorName id="26C861C0FFC6FFAFFF452FBBD011FC50" box="[224,329,943,970]" pageId="18" pageNumber="479">J. Köhler</collectorName>
,
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&amp; F.
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; 1 M
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,
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, pet trade,
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2012
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.
</paragraph>
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</treatment>
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View file

@ -0,0 +1,412 @@
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<mods:date id="2C0A7112C5679729E855BF8B002CE4CD">2024</mods:date>
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<heading id="D0CAB37AFFD7FFBEFF322B6DD23CF818" bold="true" box="[151,868,1894,1921]" fontSize="11" level="1" pageId="3" pageNumber="464" reason="1">
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<emphasis id="B949D804FFD7FFBEFF322B6DD09FF818" bold="true" box="[151,455,1894,1921]" italics="true" pageId="3" pageNumber="464">Colossobolus olympiacus</emphasis>
(
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)
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<emphasis id="B949D804FFD7FFBEFF322BC4D025F87E" box="[151,381,1999,2023]" italics="true" pageId="3" pageNumber="464">Spirobolus olympiacus</emphasis>
<bibRefCitation id="EFAC79E7FFD7FFBEFE212BC4D36AF87E" author="Karsch, F." box="[388,562,1999,2023]" pageId="3" pageNumber="464" pagination="1 - 79" refId="ref13686" refString="Karsch, F. (1881) Neue Juliden des Berliner Museums, als Prodromus einer Juliden-Monographie. Zeitschrift fur die gesammten Naturwissenschaftem, 54, 1 - 79." type="journal article" year="1881">Karsch, 1881: 64</bibRefCitation>
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(first description, female)
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<emphasis id="B949D804FFD0FFB9FF322C9CD351FF36" box="[151,521,151,175]" italics="true" pageId="4" pageNumber="465">Spirobolus (Trigoniulus) olympiacus</emphasis>
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deSaussure &amp; Zehntner 1902: 129 (list)
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<emphasis id="B949D804FFD0FFB9FF322CBED0DDFF54" box="[151,389,181,205]" italics="true" pageId="4" pageNumber="465">Trigoniulus olympiacus</emphasis>
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<emphasis id="B949D804FFD0FFB9FF322CDFD0D1FF75" box="[151,393,212,236]" italics="true" pageId="4" pageNumber="465">Spirobolus olympiaceus</emphasis>
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(sic!)
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Wesener
<emphasis id="B949D804FFD0FFB9FD8D2CDED303FF75" box="[552,603,212,236]" italics="true" pageId="4" pageNumber="465">et al.</emphasis>
2009a: 120
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<emphasis id="B949D804FFD0FFB9FF322CF8D0DDFE92" box="[151,389,243,267]" italics="true" pageId="4" pageNumber="465">Trigoniulus olympiacus</emphasis>
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<emphasis id="B949D804FFD0FFB9FF322D32D025FECA" bold="true" box="[151,381,313,339]" pageId="4" pageNumber="465">Material examined</emphasis>
.
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1 M,
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125° ESE Hellville
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, C.
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1 M, 2 F,
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(
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,
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, coll.
<collectorName id="26C861C0FFD0FFB9FF6A2DC2D016FE7A" box="[207,334,457,483]" pageId="4" pageNumber="465">B.L. Fisher</collectorName>
<emphasis id="B949D804FFD0FFB9FEF02DC1D0DEFE7A" box="[341,390,457,483]" italics="true" pageId="4" pageNumber="465">et al</emphasis>
.,
<date id="FF8322D6FFD0FFB9FE3E2DC2D357FE7A" box="[411,527,457,483]" pageId="4" pageNumber="465" value="2013-02-21">
<collectingDate id="EFC7DB3EFFD0FFB9FE3E2DC2D357FE7A" box="[411,527,457,483]" pageId="4" pageNumber="465" value="2013-02-21">21.ii.2013</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<caption id="DF42549EFFD0FFB9FF3228EBD3E4F981" ID-DOI="http://doi.org/10.5281/zenodo.14022750" ID-Zenodo-Dep="14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="4" pageNumber="465" startId="4.[151,250,1248,1272]" targetBox="[417,1171,507,1224]" targetPageId="4" targetType="figure">
<paragraph id="8B820416FFD0FFB9FF3228EBD3E4F981" blockId="4.[151,1437,1248,1560]" pageId="4" pageNumber="465">
<emphasis id="B949D804FFD0FFB9FF3228EBD04DFB61" bold="true" box="[151,277,1248,1272]" pageId="4" pageNumber="465">FIGURE 1.</emphasis>
<taxonomicName id="4C3D7F95FFD0FFB9FEB828EBD39EFB61" authority="(Karsch, 1881)" authorityName="Wesener" authorityYear="2024" baseAuthorityName="Karsch" baseAuthorityYear="1881" box="[285,710,1248,1272]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="4" pageNumber="465" phylum="Arthropoda" rank="species" species="olympiacus" status="comb. nov.">
<emphasis id="B949D804FFD0FFB9FEB828EBD379FB61" box="[285,545,1248,1272]" italics="true" pageId="4" pageNumber="465">Colossobolus olympiacus</emphasis>
(
<bibRefCitation id="EFAC79E7FFD0FFB9FD9428EBD3E6FB61" author="Karsch, F." box="[561,702,1248,1272]" pageId="4" pageNumber="465" pagination="1 - 79" refId="ref13686" refString="Karsch, F. (1881) Neue Juliden des Berliner Museums, als Prodromus einer Juliden-Monographie. Zeitschrift fur die gesammten Naturwissenschaftem, 54, 1 - 79." type="journal article" year="1881">Karsch, 1881</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="B949D804FFD0FFB9FD7728EBD268FB61" bold="true" box="[722,816,1248,1272]" pageId="4" pageNumber="465">
<taxonomicNameLabel id="A27A657FFFD0FFB9FD7728EBD268FB61" box="[722,816,1248,1272]" pageId="4" pageNumber="465" rank="species">n. comb.</taxonomicNameLabel>
</emphasis>
, topotype (
<emphasis id="B949D804FFD0FFB9FC0D28EBD537FB61" bold="true" box="[936,1135,1248,1272]" pageId="4" pageNumber="465">CASENT 9005459</emphasis>
).
<emphasis id="B949D804FFD0FFB9FB2328EBD5C0FB61" bold="true" box="[1158,1176,1248,1272]" pageId="4" pageNumber="465">A</emphasis>
. Multi-layer photograph, male anterior body, lateral view;
<emphasis id="B949D804FFD0FFB9FE5D290FD351FA85" bold="true" box="[504,521,1284,1308]" pageId="4" pageNumber="465">B</emphasis>
. Multi-layer photograph, male leg pairs 17, ventral view;
<emphasis id="B949D804FFD0FFB9FB26290FD5CDFA85" bold="true" box="[1155,1173,1284,1308]" pageId="4" pageNumber="465">C</emphasis>
. Multi-layer photograph, male posterior body end with telson;
<emphasis id="B949D804FFD0FFB9FDBD2923D369FAD9" bold="true" box="[536,561,1320,1344]" pageId="4" pageNumber="465">D.</emphasis>
Right anterior gonopod, anterior view;
<emphasis id="B949D804FFD0FFB9FC6E2923D284FAD9" bold="true" box="[971,988,1320,1344]" pageId="4" pageNumber="465">E</emphasis>
. Right anterior gonopod, posterior view;
<emphasis id="B949D804FFD0FFB9FA2C2923D4C4FAD9" bold="true" box="[1417,1436,1320,1344]" pageId="4" pageNumber="465">F.</emphasis>
Left posterior gonopod, anterior view;
<emphasis id="B949D804FFD0FFB9FD832947D319FAFD" bold="true" box="[550,577,1356,1380]" pageId="4" pageNumber="465">G.</emphasis>
Left posterior gonopod, posterior view;
<emphasis id="B949D804FFD0FFB9FC462947D2A5FAFD" bold="true" box="[995,1021,1356,1380]" pageId="4" pageNumber="465">H.</emphasis>
Male right midbody leg, posterior view.
<emphasis id="B949D804FFD0FFB9FF32297BD068FA11" bold="true" box="[151,304,1392,1416]" pageId="4" pageNumber="465">Abbreviations</emphasis>
: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number.
<emphasis id="B949D804FFD0FFB9FE542A0BD307F981" bold="true" box="[497,607,1536,1560]" pageId="4" pageNumber="465">Scale bars</emphasis>
= 1 mm.
</paragraph>
</caption>
<paragraph id="8B820416FFD0FFB9FF622A61D410F88D" blockId="4.[151,1437,1642,2029]" pageId="4" pageNumber="465">
<emphasis id="B949D804FFD0FFB9FF622A61D00EF91D" bold="true" box="[199,342,1642,1668]" pageId="4" pageNumber="465">Rediagnosis</emphasis>
.
<taxonomicName id="4C3D7F95FFD0FFB9FEC02A61D30DF91D" box="[357,597,1642,1668]" pageId="4" pageNumber="465">
<emphasis id="B949D804FFD0FFB9FEC02A61D0A4F91D" box="[357,508,1642,1668]" italics="true" pageId="4" pageNumber="465">Colossobolus</emphasis>
species
</taxonomicName>
with a finger-shaped coxite branch on the posterior gonopod (
<figureCitation id="13061893FFD0FFB9FABD2A61D4D7F91D" box="[1304,1423,1642,1668]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="4" pageNumber="465">Fig. 1F, G</figureCitation>
), like in
<taxonomicName id="4C3D7F95FFD0FFB9FF4C2A84D360F931" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[233,568,1678,1705]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="4" pageNumber="465" phylum="Arthropoda" rank="species" species="semicyclus">
<emphasis id="B949D804FFD0FFB9FF4C2A84D0DDF931" box="[233,389,1678,1705]" italics="true" pageId="4" pageNumber="465">C. semicyclus</emphasis>
Wesener, 2009
</taxonomicName>
,
<taxonomicName id="4C3D7F95FFD0FFB9FDE02A84D2EDF931" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[581,949,1678,1705]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="4" pageNumber="465" phylum="Arthropoda" rank="species" species="oblongopedus">
<emphasis id="B949D804FFD0FFB9FDE02A84D25DF931" box="[581,773,1678,1705]" italics="true" pageId="4" pageNumber="465">C. oblongopedus</emphasis>
Wesener, 2009
</taxonomicName>
and
<taxonomicName id="4C3D7F95FFD0FFB9FC4A2A84D46BF931" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1007,1331,1678,1705]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="4" pageNumber="465" phylum="Arthropoda" rank="species" species="giganteus">
<emphasis id="B949D804FFD0FFB9FC4A2A84D527F931" box="[1007,1151,1679,1705]" italics="true" pageId="4" pageNumber="465">C. giganteus</emphasis>
Wesener, 2009
</taxonomicName>
. Anterior gonopods lacking a retrorse process on the mesal margin of the telopodite (
<figureCitation id="13061893FFD0FFB9FC562AB9D537F955" box="[1011,1135,1714,1740]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="4" pageNumber="465">Fig. 1D, E</figureCitation>
), like in
<taxonomicName id="4C3D7F95FFD0FFB9FB732AB8D4CDF955" authorityName="Wesener" authorityYear="2009" box="[1238,1429,1714,1741]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="4" pageNumber="465" phylum="Arthropoda" rank="species" species="oblongopedus">
<emphasis id="B949D804FFD0FFB9FB732AB8D4CDF955" box="[1238,1429,1714,1741]" italics="true" pageId="4" pageNumber="465">C. oblongopedus</emphasis>
</taxonomicName>
. Posterior gonopods most similar to those of
<taxonomicName id="4C3D7F95FFD0FFB9FDD12ADCD277F969" authorityName="Wesener" authorityYear="2009" box="[628,815,1750,1777]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="4" pageNumber="465" phylum="Arthropoda" rank="species" species="oblongopedus">
<emphasis id="B949D804FFD0FFB9FDD12ADCD277F969" box="[628,815,1750,1777]" italics="true" pageId="4" pageNumber="465">C. oblongopedus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3D7F95FFD0FFB9FCC42ADCD2B4F969" authorityName="Wesener" authorityYear="2009" box="[865,1004,1751,1777]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="4" pageNumber="465" phylum="Arthropoda" rank="species" species="giganteus">
<emphasis id="B949D804FFD0FFB9FCC42ADCD2B4F969" box="[865,1004,1751,1777]" italics="true" pageId="4" pageNumber="465">C. giganteus</emphasis>
</taxonomicName>
, but differing in the narrowly elongated tip of the lateral branch, as well as the apical part of the mesal branch. Male legs with tarsal pads (
<figureCitation id="13061893FFD0FFB9FB472AF1D463F88D" box="[1250,1339,1786,1812]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="4" pageNumber="465">Fig. 1H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C327579DFFD0FFB8FF622B15D306FC3C" lastPageId="5" lastPageNumber="466" pageId="4" pageNumber="465" type="description">
<paragraph id="8B820416FFD0FFB9FF622B15D23BF8A1" blockId="4.[151,1437,1642,2029]" box="[199,867,1822,1849]" pageId="4" pageNumber="465">
<emphasis id="B949D804FFD0FFB9FF622B15D035F8A1" bold="true" box="[199,365,1822,1848]" pageId="4" pageNumber="465">Redescription</emphasis>
(based on topotypic material from Nosy Be)
</paragraph>
<paragraph id="8B820416FFD0FFB9FF622B48D5B3F8C5" blockId="4.[151,1437,1642,2029]" box="[199,1259,1858,1885]" pageId="4" pageNumber="465">
Measurements: 56+0 body rings plus telson.
<quantity id="4CC5A9F3FFD0FFB9FD1A2B49D244F8C4" box="[703,796,1858,1885]" metricMagnitude="-1" metricUnit="m" metricValue="1.78" pageId="4" pageNumber="465" unit="mm" value="178.0">178 mm</quantity>
long,
<quantity id="4CC5A9F3FFD0FFB9FCC72B49D28CF8C4" box="[866,980,1858,1885]" metricMagnitude="-2" metricUnit="m" metricValue="1.215" pageId="4" pageNumber="465" unit="mm" value="12.15">12.15 mm</quantity>
wide (midbody, widest).
</paragraph>
<paragraph id="8B820416FFD0FFB9FF622B6DD29AF83D" blockId="4.[151,1437,1642,2029]" pageId="4" pageNumber="465">
Colour: mesozonite orange-grey, metazonite black, telson blood red. Ozopores black, head, antenna, mouthparts red. Collum at margins black, surface red (
<figureCitation id="13061893FFD0FFB9FDD02B81D396F83D" box="[629,718,1930,1956]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="4" pageNumber="465">Fig. 1A</figureCitation>
). Legs red to greyish.
</paragraph>
<paragraph id="8B820416FFD0FFB8FF622BA5D3DCFF4C" blockId="4.[151,1437,1642,2029]" lastBlockId="5.[151,1437,151,1257]" lastPageId="5" lastPageNumber="466" pageId="4" pageNumber="465">
Head with 35/36 black ommatidia (
<figureCitation id="13061893FFD0FFB9FDF72BA5D3F3F851" box="[594,683,1966,1992]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="4" pageNumber="465">Fig. 1A</figureCitation>
). Vertical suture not going through whole head, head with the usual 2+2 setiferous foveolae (
<figureCitation id="13061893FFD0FFB9FE122BD9D349F875" box="[439,529,2002,2028]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="4" pageNumber="465">Fig. 1B</figureCitation>
). Gnathochilarium typical for the order, lamellae linguales each with 1+1 seta behind one another, stipes each with three setae (
<figureCitation id="13061893FFD1FFB8FD762C9CD275FF28" box="[723,813,151,177]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="5" pageNumber="466">Fig. 1B</figureCitation>
). Mandible basal lobe not projecting. Antennae short, protruding back to third body ring (
<figureCitation id="13061893FFD1FFB8FD812CB0D323FF4C" box="[548,635,187,213]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="5" pageNumber="466">Fig. 1A</figureCitation>
)
</paragraph>
<paragraph id="8B820416FFD1FFB8FF622CD4D545FF60" blockId="5.[151,1437,151,1257]" box="[199,1053,223,249]" pageId="5" pageNumber="466">Ozopore starting at ring 6, located on suture between meso- and metazonite.</paragraph>
<paragraph id="8B820416FFD1FFB8FF622D08D478FEFC" blockId="5.[151,1437,151,1257]" pageId="5" pageNumber="466">
Legs in male on coxae 3 and 4 with short conical process (
<figureCitation id="13061893FFD1FFB8FCD62D08D294FE84" box="[883,972,259,285]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="5" pageNumber="466">Fig. 1B</figureCitation>
), coxae 57 normal, tarsal pads present, starting at leg 3 (
<figureCitation id="13061893FFD1FFB8FEF62D2CD0F5FED8" box="[339,429,295,321]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="5" pageNumber="466">Fig. 1G</figureCitation>
). Legs 8+ with femur cylindrical, as long as tarsus, podomere length 1&lt;2&lt;3&gt;4=5&lt;6. Each podomere apically with one or two spines, tarsus with wide claw and single short apical spine (
<figureCitation id="13061893FFD1FFB8FB652D40D44FFEFC" box="[1216,1303,331,357]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="5" pageNumber="466">Fig. 1H</figureCitation>
)
</paragraph>
<paragraph id="8B820416FFD1FFB8FF622D64D5DFFE34" blockId="5.[151,1437,151,1257]" pageId="5" pageNumber="466">
Telson: preanal scale very slightly projecting in well-rounded process, not projecting beyond anal valves (
<figureCitation id="13061893FFD1FFB8FACA2D64D1E2FE34" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="5" pageNumber="466">Fig. 1C</figureCitation>
). Anal valves with well-developed lips and large groove. Subanal scale inconspicuous.
</paragraph>
<paragraph id="8B820416FFD1FFB8FF622DBCD0F1FDA4" blockId="5.[151,1437,151,1257]" pageId="5" pageNumber="466">
Anterior gonopod sternite elongated into a wide, broadly rounded-triangular lobe (
<figureCitation id="13061893FFD1FFB8FBC82DBCD59FFE48" box="[1133,1223,439,465]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="5" pageNumber="466">Fig. 1D</figureCitation>
). Mesal process of coxite weakly developed, protruding into a short, wide lobe with a rounded tip (
<figureCitation id="13061893FFD1FFB8FB802DD0D527FE6C" box="[1061,1151,475,501]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="5" pageNumber="466">Fig. 1D</figureCitation>
). Telopodite on posterior side basally with a circular groove (
<figureCitation id="13061893FFD1FFB8FD912DF4D3D4FD80" box="[564,652,511,537]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="5" pageNumber="466">Fig. 1E</figureCitation>
). Telopodite process long and well-rounded (
<figureCitation id="13061893FFD1FFB8FB312DF4D5B3FD80" box="[1172,1259,511,537]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="5" pageNumber="466">Fig. 1E</figureCitation>
), mesal margin laterally sharp (
<figureCitation id="13061893FFD1FFB8FEE32E28D0C3FDA4" box="[326,411,547,573]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="5" pageNumber="466">Fig. 1E</figureCitation>
).
</paragraph>
<paragraph id="8B820416FFD1FFB8FF622E4CD224FCA0" blockId="5.[151,1437,151,1257]" pageId="5" pageNumber="466">
Posterior gonopods telopodites positioned parallel to one another. Coxite relatively short, wide, stout, less than half as long than lateral branch of telopodite (
<figureCitation id="13061893FFD1FFB8FD462E60D205FD1C" box="[739,861,619,645]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="5" pageNumber="466">Fig. 1F, G</figureCitation>
). Latter slightly curved, basally wide, at midpoint tapering, tip again wide and swollen (
<figureCitation id="13061893FFD1FFB8FD9B2E84D3CBFD30" box="[574,659,655,681]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="5" pageNumber="466">Fig. 1F</figureCitation>
). Central membranous area developed as a short, wide, well-rounded lobe (
<figureCitation id="13061893FFD1FFB8FF7D2EB8D076FD54" box="[216,302,691,717]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="5" pageNumber="466">Fig. 1F</figureCitation>
). Mesal main branch short, basal part running parallel to lateral branch. Apical part of main branch slender, extending towards and touching lateral branch (
<figureCitation id="13061893FFD1FFB8FCA42EDCD201FD68" box="[769,857,727,753]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="5" pageNumber="466">Fig. 1G</figureCitation>
). Mesal margin of main branch with a well-rounded, overlapping membranous fold (
<figureCitation id="13061893FFD1FFB8FE592EF0D309FC8C" box="[508,593,763,789]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 17, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="5" pageNumber="466">Fig. 1F</figureCitation>
). Main branch laterally with a small, irregular-shaped membrane (
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). Lateral branch apically strongly tapered, slender (
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).
</paragraph>
<paragraph id="8B820416FFD1FFB8FF622F49D306FC3C" blockId="5.[151,1437,151,1257]" pageId="5" pageNumber="466">
<emphasis id="B949D804FFD1FFB8FF622F49D080FCC5" bold="true" box="[199,472,834,860]" pageId="5" pageNumber="466">Intraspecific variation.</emphasis>
Specimens from the Galoko Mountain Chain have only 5254 body rings and are with a length of
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, a width of
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in the male and a length of
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length with a width of
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in the female shorter than those from Nosy Be.
</paragraph>
</subSubSection>
<subSubSection id="C327579DFFD1FFB8FF622FA4D0CDFB70" pageId="5" pageNumber="466" type="discussion">
<paragraph id="8B820416FFD1FFB8FF622FA4D419FC50" blockId="5.[151,1437,151,1257]" box="[199,1345,943,969]" pageId="5" pageNumber="466">
<emphasis id="B949D804FFD1FFB8FF622FA4D063FC50" bold="true" box="[199,315,943,969]" pageId="5" pageNumber="466">Remarks.</emphasis>
At both localities found in sympatry with
<taxonomicName id="4C3D7F95FFD1FFB8FCB02FA4D464FC50" authority="(Pocock, 1893)" baseAuthorityName="Pocock" baseAuthorityYear="1893" box="[789,1340,943,969]" class="Diplopoda" family="Pachybolidae" genus="Aphistogoniulus" kingdom="Animalia" order="Spirobolida" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="erythrocephalus">
<emphasis id="B949D804FFD1FFB8FCB02FA4D5DFFC50" box="[789,1159,943,969]" italics="true" pageId="5" pageNumber="466">Aphistogoniulus erythrocephalus</emphasis>
(
<bibRefCitation id="EFAC79E7FFD1FFB8FB332FA4D46CFC50" author="Pocock, R. I." box="[1174,1332,943,969]" pageId="5" pageNumber="466" pagination="248 - 254" refId="ref14311" refString="Pocock, R. I. (1893) Upon the identity of some of the types of Diplopoda contained in the collection of the British Museum, together with description of some new species of exotic Julidae. Annals and Magazine of Natural History, Series 6, 11, 248 - 254. https: // doi. org / 10.1080 / 00222939308677511" type="journal article" year="1893">Pocock, 1893</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8B820416FFD1FFB8FF622FD8D0CDFB70" blockId="5.[151,1437,151,1257]" pageId="5" pageNumber="466">
<taxonomicName id="4C3D7F95FFD1FFB8FF622FD8D335FC74" authority="Karsch, 1881" authorityName="Karsch" authorityYear="1881" box="[199,621,979,1005]" class="Diplopoda" family="Spirobolidae" genus="Spirobolus" kingdom="Animalia" order="Spirobolida" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="olympiacus">
<emphasis id="B949D804FFD1FFB8FF622FD8D092FC74" box="[199,458,979,1005]" italics="true" pageId="5" pageNumber="466">Spirobolus olympiacus</emphasis>
<bibRefCitation id="EFAC79E7FFD1FFB8FE712FD8D335FC74" author="Karsch, F." box="[468,621,979,1005]" pageId="5" pageNumber="466" pagination="1 - 79" refId="ref13686" refString="Karsch, F. (1881) Neue Juliden des Berliner Museums, als Prodromus einer Juliden-Monographie. Zeitschrift fur die gesammten Naturwissenschaftem, 54, 1 - 79." type="journal article" year="1881">Karsch, 1881</bibRefCitation>
</taxonomicName>
was one of the few remaining
<emphasis id="B949D804FFD1FFB8FC442FD8D527FC74" box="[993,1151,979,1005]" italics="true" pageId="5" pageNumber="466">nomina dubia</emphasis>
of the
<taxonomicName id="4C3D7F95FFD1FFB8FB7D2FD8D403FC74" authorityName="Bollman" authorityYear="1893" box="[1240,1371,979,1005]" class="Diplopoda" kingdom="Animalia" order="Spirobolida" pageId="5" pageNumber="466" phylum="Arthropoda" rank="order">Spirobolida</taxonomicName>
from
<collectingCountry id="F32A4486FFD1FFB8FF322FFCD046FB88" box="[151,286,1015,1041]" name="Madagascar" pageId="5" pageNumber="466">Madagascar</collectingCountry>
(
<bibRefCitation id="EFAC79E7FFD1FFB8FE8B2FFCD37BFB88" author="Wesener T. &amp; Enghoff H. &amp; Sierwald P." box="[302,547,1015,1041]" pageId="5" pageNumber="466" pagination="1 - 128" refId="ref15399" refString="Wesener T., Enghoff H., Sierwald P. (2009 a) Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda). ZooKeys, 19, 1 - 128. https: // doi. org / 10.3897 / zookeys. 19.221" type="journal article" year="2009">
Wesener
<emphasis id="B949D804FFD1FFB8FE3D2FFCD08AFB88" box="[408,466,1015,1041]" italics="true" pageId="5" pageNumber="466">et al.</emphasis>
2009a
</bibRefCitation>
). The short first description is the only known record, based on a female from Nosy Be, and mentions few characters aside from the large size (
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) and the presence of 57 rings. The
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(and only known specimen) seems unfortunately lost. It is no longer in the collections of the museum in Berlin (Mark Judson, pers. comm.), and it is also one of the few not listed in the first Berlin Myriapoda type catalogues, neither under
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nor
<taxonomicName id="4C3D7F95FFD1FFB8FDCD288CD25DFB38" authorityName="Brandt" authorityYear="1833" box="[616,773,1159,1185]" class="Diplopoda" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="466" phylum="Arthropoda" rank="order">Spirostreptida</taxonomicName>
(
<bibRefCitation id="EFAC79E7FFD1FFB8FCB6288CD54EFB38" author="Moritz, M. &amp; Fischer, S. C." box="[787,1046,1159,1185]" pageId="5" pageNumber="466" pagination="323 - 375" refId="ref13815" refString="Moritz, M. &amp; Fischer, S. C. (1974) Typen der Myriapoden-Sammlung des Zoologischen Museums Berlin. I. Diplopoda. Teil 2: Craspedosomatida, Stemmiulida, Spirostreptida. Mitteilungen aus dem Museum fur Naturkunde in Berlin. Zoologisches Museum und Institut fur Spezielle Zoologie, Berlin, 50 (1), 323 - 375. https: // doi. org / 10.1002 / mmnz. 19750510206" type="journal article" year="1974">Moritz &amp; Fischer 1974</bibRefCitation>
,
<bibRefCitation id="EFAC79E7FFD1FFB8FB85288DD504FB38" author="Moritz, M. &amp; Fischer, S. C." box="[1056,1116,1158,1185]" pageId="5" pageNumber="466" pagination="199 - 255" refId="ref13890" refString="Moritz, M. &amp; Fischer, S. C. (1975) Die Typen der Myriapoden-Sammlung des Zoologischen Museums Berlin. I. Diplopoda. Teil 3: Julida, Spirobolida. Mitteilungen aus dem Museum fur Naturkunde in Berlin. Zoologisches Museum und Institut fur Spezielle Zoologie, Berlin, 51 (2), 199 - 255. https: // doi. org / 10.1002 / mmnz. 19750510206" type="journal article" year="1975">1975</bibRefCitation>
). The discovery of a massive
<taxonomicName id="4C3D7F95FFD1FFB8FF3228A0D042FB5C" authorityName="Bollman" authorityYear="1893" box="[151,282,1195,1221]" class="Diplopoda" kingdom="Animalia" order="Spirobolida" pageId="5" pageNumber="466" phylum="Arthropoda" rank="order">Spirobolida</taxonomicName>
from Nosy Be (albeit “only”
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in length) with 56 rings plus telson allows here a redescription of this enigmatic species.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,766 @@
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<mods:identifier id="E60908368738E326951B761DB44327DD" type="DOI">10.11646/zootaxa.5529.3.4</mods:identifier>
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<subSubSection id="C3134895165DFFF542FBFEB96C88FE65" pageId="4" pageNumber="491" type="nomenclature">
<paragraph id="8BB61B1E165DFFF542FBFEB96D95FE40" blockId="4.[151,961,366,429]" box="[151,732,366,393]" pageId="4" pageNumber="491">
<heading id="D0FEAC72165DFFF542FBFEB96D95FE40" bold="true" box="[151,732,366,393]" fontSize="11" level="1" pageId="4" pageNumber="491" reason="1">
<emphasis id="B97DC70C165DFFF542FBFEB96D95FE40" bold="true" box="[151,732,366,393]" pageId="4" pageNumber="491">
<taxonomicName id="4C09609D165DFFF542FBFEB96ED2FE41" authorityName="Bañón &amp; Barros-García &amp; Arronte &amp; Rábade &amp; Rio &amp; Baldó &amp; Carlos" authorityYear="2024" box="[151,411,366,393]" family="Neoscopelidae" genus="Neoscopelus" kingdom="Animalia" order="Myctophiformes" pageId="4" pageNumber="491" phylum="Chordata" rank="species" species="serranoi" status="sp. nov.">
<emphasis id="B97DC70C165DFFF542FBFEB96ED2FE41" bold="true" box="[151,411,366,393]" italics="true" pageId="4" pageNumber="491">Neoscopelus serranoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E7A77165DFFF543CEFEB86D49FE41" box="[418,512,367,393]" pageId="4" pageNumber="491" rank="species">sp. nov.</taxonomicNameLabel>
<figureCitation id="1332079B165DFFF54064FEB96D38FE40" box="[520,625,366,392]" captionStart="FIGURE 1" captionStartId="6.[151,250,1001,1025]" captionTargetBox="[303,1284,250,965]" captionTargetId="figure-48@6.[290,1297,237,978]" captionTargetPageId="6" captionText="FIGURE 1. Neoscopelus serranoi sp. nov. a holotype MHN USC 25200-2, 236 mm TL; b paratype MHN USC 25200-3, 187 mm TL." figureDoi="http://doi.org/10.5281/zenodo.14022809" httpUri="https://zenodo.org/record/14022809/files/figure.png" pageId="4" pageNumber="491">Figure 1</figureCitation>
;
<tableCitation id="C68B2EA5165DFFF540EDFEB86D95FE40" box="[641,732,366,393]" captionStart="TABLE 1" captionStartId="5.[151,239,151,175]" captionTargetPageId="5" captionText="TABLE 1. Morphometric and meristic data of Neoscopelus serranoi sp. nov. based on the examination of 35 specimens from the NE Atlantic (holotype and 34 paratypes) compared with specimens of Neoscopelus microchir from the Western Pacific based on Matsubara (1943) (holotype and 37 paratypes)." pageId="4" pageNumber="491">Table 1</tableCitation>
</emphasis>
</heading>
</paragraph>
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<uri id="FF98171C165DFFF542FBFE456C88FE65" box="[151,961,402,429]" pageId="4" pageNumber="491">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="C3134895165DFFF542FBFE0D6A7AFE3D" box="[151,1331,474,501]" pageId="4" pageNumber="491" type="reference_group">
<paragraph id="8BB61B1E165DFFF542FBFE0D6A7AFE3D" blockId="4.[151,1437,474,2013]" box="[151,1331,474,501]" pageId="4" pageNumber="491">
<emphasis id="B97DC70C165DFFF542FBFE0D6E50FE3C" bold="true" box="[151,281,474,500]" pageId="4" pageNumber="491">References</emphasis>
.
<taxonomicName id="4C09609D165DFFF5434BFE0C6C5DFE3D" ID-CoL="46SS4" authority="(Banon et al., 2002)" baseAuthorityName="Banon" baseAuthorityYear="2002" box="[295,788,475,501]" family="Neoscopelidae" genus="Neoscopelus" kingdom="Animalia" order="Myctophiformes" pageId="4" pageNumber="491" phylum="Chordata" rank="species" species="microchir">
<emphasis id="B97DC70C165DFFF5434BFE0C6D60FE3D" box="[295,553,475,501]" italics="true" pageId="4" pageNumber="491">Neoscopelus microchir</emphasis>
(
<bibRefCitation id="EF9866EF165DFFF54054FE0C6C44FE3D" author="Banon, R. &amp; Rio, J. L. &amp; Pineiro, C. &amp; Casas, M." box="[568,781,475,501]" pageId="4" pageNumber="491" pagination="877 - 880" refId="ref8039" refString="Banon, R., del Rio, J. L., Pineiro, C. &amp; Casas, M. (2002) Occurrence of tropical affinity fishes in Galician waters NW Spain. Journal of the Marine Biological Association of the United Kingdom, 82 (5), 877 - 880. https: // doi. org / 10.1017 / S 0025315402006288" type="journal article" year="2002">
Bañón
<emphasis id="B97DC70C165DFFF540E6FE0C6D8BFE3D" box="[650,706,475,501]" italics="true" pageId="4" pageNumber="491">et al.</emphasis>
, 2002
</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="4C09609D165DFFF5414CFE0C6A66FE3D" ID-CoL="46SS4" authority="(Banon et al., 2022)" baseAuthorityName="Banon" baseAuthorityYear="2022" box="[800,1327,475,501]" family="Neoscopelidae" genus="Neoscopelus" isUncertain="true" kingdom="Animalia" order="Myctophiformes" pageId="4" pageNumber="491" phylum="Chordata" rank="species" species="microchir">
<emphasis id="B97DC70C165DFFF5414CFE0C6B0AFE3D" box="[800,1091,475,501]" italics="true" pageId="4" pageNumber="491">Neoscopelus cf. microchir</emphasis>
(
<bibRefCitation id="EF9866EF165DFFF5463EFE0C6A6EFE3D" author="Banon, R. &amp; de Carlos, A. &amp; Acosta-Morillas, V. &amp; Baldo, F." box="[1106,1319,475,501]" pageId="4" pageNumber="491" refId="ref8262" refString="Banon, R., de Carlos, A., Acosta-Morillas, V. &amp; Baldo, F. (2022) Geographic range expansion and taxonomic notes of the shortfin neoscopelid Neoscopelus cf. microchir (Myctophiformes: Neoscopelidae) in the North-Eastern Atlantic. Journal of Marine Science and Engineering, 10, 954. https: // doi. org / 10.3390 / jmse 10070954" type="journal volume" year="2022">
Bañón
<emphasis id="B97DC70C165DFFF546C8FE0C6B94FE3D" box="[1188,1245,475,501]" italics="true" pageId="4" pageNumber="491">et al.</emphasis>
, 2022
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3134895165DFFF542ABFE286C4EFDA9" pageId="4" pageNumber="491" type="etymology">
<paragraph id="8BB61B1E165DFFF542ABFE286C4EFDA9" blockId="4.[151,1437,474,2013]" pageId="4" pageNumber="491">
<emphasis id="B97DC70C165DFFF542ABFE286E01FDD1" bold="true" box="[199,328,511,537]" pageId="4" pageNumber="491">Etymology</emphasis>
. The name of the new species is dedicated to Dr. Alberto Serrano, from the Instituto Español de Oceanografía (IEO-CSIC), leader of the expedition that carried out the scientific sampling at the
<collectingRegion id="49CDD5FC165DFFF54688FDF46A7CFDF5" box="[1252,1333,547,573]" country="Spain" name="Galicia" pageId="4" pageNumber="491">Galicia</collectingRegion>
bank, an area where this species has been found most frequently.
</paragraph>
</subSubSection>
<subSubSection id="C3134895165DFFF542ABFDBC6E30FBFD" pageId="4" pageNumber="491" type="materials_examined">
<paragraph id="8BB61B1E165DFFF542ABFDBC6AD5FD61" blockId="4.[151,1437,474,2013]" pageId="4" pageNumber="491">
<materialsCitation id="3B611143165DFFF542ABFDBC6AD5FD61" collectingDate="2010-08-17" collectionCode="MHN" elevation="766" latitude="42.666" location="Bank" longLatPrecision="68" longitude="-11.722" pageId="4" pageNumber="491" specimenCode="NSM002" specimenCount="1" stateProvince="Galicia" typeStatus="holotype">
<emphasis id="B97DC70C165DFFF542ABFDBC6E7DFD4D" bold="true" box="[199,308,619,645]" pageId="4" pageNumber="491">
<typeStatus id="54B2A5BC165DFFF542ABFDBC6E7DFD4D" box="[199,308,619,645]" pageId="4" pageNumber="491" type="holotype">Holotype</typeStatus>
</emphasis>
.
<taxonomicName id="4C09609D165DFFF54328FDBC6D73FD4C" authorityName="Bañón &amp; Barros-García &amp; Arronte &amp; Rábade &amp; Rio &amp; Baldó &amp; Carlos" authorityYear="2024" box="[324,570,619,645]" family="Neoscopelidae" genus="Neoscopelus" kingdom="Animalia" order="Myctophiformes" pageId="4" pageNumber="491" phylum="Chordata" rank="species" species="serranoi" status="sp. nov.">
<emphasis id="B97DC70C165DFFF54328FDBC6D73FD4C" box="[324,570,619,645]" italics="true" pageId="4" pageNumber="491">Neoscopelus serranoi</emphasis>
</taxonomicName>
<emphasis id="B97DC70C165DFFF5402FFDBC6DD6FD4D" bold="true" box="[579,671,619,645]" pageId="4" pageNumber="491">
<taxonomicNameLabel id="A24E7A77165DFFF5402FFDBC6DD6FD4D" box="[579,671,619,645]" pageId="4" pageNumber="491" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<collectionCode id="ED1883DB165DFFF540C4FDBC6DA0FD4D" box="[680,745,619,645]" pageId="4" pageNumber="491">MHN</collectionCode>
USC 25200-
<quantity id="4CF1B6FB165DFFF541E9FDBD6B49FD4D" box="[901,1024,618,645]" metricMagnitude="0" metricUnit="m" metricValue="2.2359999999999998" pageId="4" pageNumber="491" unit="mm" value="2236.0">2, 236 mm</quantity>
TL (
<tableCitation id="C68B2EA5165DFFF54656FDBC6BD8FD4D" box="[1082,1169,619,645]" captionStart="TABLE 1" captionStartId="5.[151,239,151,175]" captionTargetPageId="5" captionText="TABLE 1. Morphometric and meristic data of Neoscopelus serranoi sp. nov. based on the examination of 35 specimens from the NE Atlantic (holotype and 34 paratypes) compared with specimens of Neoscopelus microchir from the Western Pacific based on Matsubara (1943) (holotype and 37 paratypes)." pageId="4" pageNumber="491">Table 1</tableCitation>
,
<figureCitation id="1332079B165DFFF546CDFDBC6A46FD4D" box="[1185,1295,619,645]" captionStart="FIGURE 1" captionStartId="6.[151,250,1001,1025]" captionTargetBox="[303,1284,250,965]" captionTargetId="figure-48@6.[290,1297,237,978]" captionTargetPageId="6" captionText="FIGURE 1. Neoscopelus serranoi sp. nov. a holotype MHN USC 25200-2, 236 mm TL; b paratype MHN USC 25200-3, 187 mm TL." figureDoi="http://doi.org/10.5281/zenodo.14022809" httpUri="https://zenodo.org/record/14022809/files/figure.png" pageId="4" pageNumber="491">Figure 1a</figureCitation>
),
<date id="FFB73DDE165DFFF54744FDBC6F9AFD61" pageId="4" pageNumber="491" value="2010-08-17">
<collectingDate id="EFF3C436165DFFF54744FDBC6F9AFD61" pageId="4" pageNumber="491" value="2010-08-17">17 August 2010</collectingDate>
</date>
;
<collectingRegion id="49CDD5FC165DFFF542B2FD586E66FD61" box="[222,303,655,681]" country="Spain" name="Galicia" pageId="4" pageNumber="491">Galicia</collectingRegion>
<location id="8ED64DC5165DFFF5435AFD586E3DFD61" LSID="urn:lsid:plazi:treatment:03A0AA08165DFFF742FBFEB96B23FF1D:8ED64DC5165DFFF5435AFD586E3DFD61" box="[310,372,655,681]" latitude="42.666" longLatPrecision="68" longitude="-11.722" name="Bank" pageId="4" pageNumber="491" stateProvince="Galicia">Bank</location>
,
<geoCoordinate id="EE3D7DD9165DFFF54313FD586EBAFD61" box="[383,499,655,681]" degrees="42.666" direction="north" orientation="latitude" pageId="4" pageNumber="491" precision="55" value="42.666">42.666 ºN</geoCoordinate>
, -
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,
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<quantity id="4CF1B6FB165DFFF540E5FD586D99FD61" box="[649,720,655,681]" metricMagnitude="2" metricUnit="m" metricValue="7.659999999999999" pageId="4" pageNumber="491" unit="m" value="766.0">766 m</quantity>
depth
</elevation>
, sample ID:
<specimenCode id="DBAFB365165DFFF541C9FD586B46FD61" box="[933,1039,655,681]" collectionCode="NSM" country="Canada" httpUri="http://grbio.org/cool/zeys-ppzt" name="Nova Scotia Museum of Natural History" pageId="4" pageNumber="491">NSM002</specimenCode>
; GenBank registration: OR076048.
</materialsCitation>
</paragraph>
<paragraph id="8BB61B1E165DFFF542ABFD646BD2FC01" blockId="4.[151,1437,474,2013]" pageId="4" pageNumber="491">
<emphasis id="B97DC70C165DFFF542ABFD646E76FD05" bold="true" box="[199,319,691,717]" pageId="4" pageNumber="491">
<typeStatus id="54B2A5BC165DFFF542ABFD646E76FD05" box="[199,319,691,717]" pageId="4" pageNumber="491" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<taxonomicName id="4C09609D165DFFF54323FD646D0FFD04" authorityName="Bañón &amp; Barros-García &amp; Arronte &amp; Rábade &amp; Rio &amp; Baldó &amp; Carlos" authorityYear="2024" box="[335,582,691,717]" family="Neoscopelidae" genus="Neoscopelus" kingdom="Animalia" order="Myctophiformes" pageId="4" pageNumber="491" phylum="Chordata" rank="species" species="serranoi" status="sp. nov.">
<emphasis id="B97DC70C165DFFF54323FD646D0FFD04" box="[335,582,691,717]" italics="true" pageId="4" pageNumber="491">Neoscopelus serranoi</emphasis>
</taxonomicName>
<emphasis id="B97DC70C165DFFF54023FD646DE2FD05" bold="true" box="[591,683,691,717]" pageId="4" pageNumber="491">
<taxonomicNameLabel id="A24E7A77165DFFF54023FD646DE2FD05" box="[591,683,691,717]" pageId="4" pageNumber="491" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<collectionCode id="ED1883DB165DFFF540D9FD646DBFFD05" box="[693,758,691,717]" pageId="4" pageNumber="491">MHN</collectionCode>
USC 25200-3 (
<figureCitation id="1332079B165DFFF541C2FD646B56FD05" box="[942,1055,691,717]" captionStart="FIGURE 1" captionStartId="6.[151,250,1001,1025]" captionTargetBox="[303,1284,250,965]" captionTargetId="figure-48@6.[290,1297,237,978]" captionTargetPageId="6" captionText="FIGURE 1. Neoscopelus serranoi sp. nov. a holotype MHN USC 25200-2, 236 mm TL; b paratype MHN USC 25200-3, 187 mm TL." figureDoi="http://doi.org/10.5281/zenodo.14022809" httpUri="https://zenodo.org/record/14022809/files/figure.png" pageId="4" pageNumber="491">Figure 1b</figureCitation>
),
<quantity id="4CF1B6FB165DFFF54654FD646BD1FD05" box="[1080,1176,691,717]" metricMagnitude="-1" metricUnit="m" metricValue="1.8699999999999999" pageId="4" pageNumber="491" unit="mm" value="187.0">187 mm</quantity>
TL,
<date id="FFB73DDE165DFFF546BFFD656AD1FD05" box="[1235,1432,690,717]" pageId="4" pageNumber="491" value="2022-10-15">15 October 2022</date>
; Cantabrian Sea,
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, -
<geoCoordinate id="EE3D7DD9165DFFF5438BFD006D1EFD39" box="[487,599,727,753]" degrees="3.808" direction="west" orientation="longitude" pageId="4" pageNumber="491" precision="55" value="-3.808">3.808 ºW</geoCoordinate>
,
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depth, sample ID:
<collectionCode id="ED1883DB165DFFF541F4FD006C9EFD39" box="[920,983,727,753]" country="Canada" httpUri="http://grbio.org/cool/zeys-ppzt" name="Nova Scotia Museum of Natural History" pageId="4" pageNumber="491">NSM</collectionCode>
004; GenBank registration: OR076047;
<taxonomicName id="4C09609D165DFFF542FBFD2C6E91FCDD" authority="IIPB" authorityName="IIPB" box="[151,472,763,789]" family="Neoscopelidae" genus="Neoscopelus" kingdom="Animalia" order="Myctophiformes" pageId="4" pageNumber="491" phylum="Chordata" rank="species" species="microchir">
<emphasis id="B97DC70C165DFFF542FBFD2C6ED3FCDD" box="[151,410,763,789]" italics="true" pageId="4" pageNumber="491">Neoscopelus microchir</emphasis>
IIPB
</taxonomicName>
7/2001 to
<collectionCode id="ED1883DB165DFFF54039FD2C6DC3FCDD" box="[597,650,763,789]" country="Spain" httpUri="http://grbio.org/cool/55k9-48j7" name="Instituto de Ciencias del Mar" pageId="4" pageNumber="491">IIPB</collectionCode>
38/2001,
<date id="FFB73DDE165DFFF54091FD2C6CD6FCDD" box="[765,927,763,789]" pageId="4" pageNumber="491" value="1999-04-10">10 April 1999</date>
,
<collectingRegion id="49CDD5FC165DFFF541C7FD2C6CB5FCDD" box="[939,1020,763,789]" country="Spain" name="Galicia" pageId="4" pageNumber="491">Galicia</collectingRegion>
Bank,
<geoCoordinate id="EE3D7DD9165DFFF54623FD2C6BF2FCDD" box="[1103,1211,763,789]" degrees="42.666" direction="north" orientation="latitude" pageId="4" pageNumber="491" precision="55" value="42.666">42.666 N</geoCoordinate>
, -
<geoCoordinate id="EE3D7DD9165DFFF546A3FD2C6A09FCDD" box="[1231,1344,763,789]" degrees="11.700" direction="west" orientation="longitude" pageId="4" pageNumber="491" precision="55" value="-11.7">11.700 W</geoCoordinate>
,
<quantity id="4CF1B6FB165DFFF54720FD2C6AD1FCDD" box="[1356,1432,763,789]" metricMagnitude="2" metricUnit="m" metricValue="7.64" pageId="4" pageNumber="491" unit="m" value="764.0">764 m</quantity>
;
<taxonomicName id="4C09609D165DFFF542FBFCC86EF1FCF1" box="[151,440,799,825]" family="Neoscopelidae" genus="Neoscopelus" isUncertain="true" kingdom="Animalia" order="Myctophiformes" pageId="4" pageNumber="491" phylum="Chordata" rank="species" species="microchir">
<emphasis id="B97DC70C165DFFF542FBFCC86EF1FCF1" box="[151,440,799,825]" italics="true" pageId="4" pageNumber="491">Neoscopelus cf microchir</emphasis>
</taxonomicName>
,
<date id="FFB73DDE165DFFF543A5FCC86D36FCF1" box="[457,639,799,825]" pageId="4" pageNumber="491" value="2010-08-17">17 August 2010</date>
<collectionCode id="ED1883DB165DFFF540E4FCC86D80FCF1" box="[648,713,799,825]" pageId="4" pageNumber="491">MHN</collectionCode>
USC 25200, Porcupine Bank,
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,
<geoCoordinate id="EE3D7DD9165DFFF546AAFCC86A0FFCF1" box="[1222,1350,799,825]" degrees="13.2322" direction="west" orientation="longitude" pageId="4" pageNumber="491" precision="5" value="-13.2322">13.2322 W</geoCoordinate>
,
<quantity id="4CF1B6FB165DFFF5473FFCC96AD5FCF1" box="[1363,1436,798,825]" metricMagnitude="2" metricUnit="m" metricValue="7.05" pageId="4" pageNumber="491" unit="m" value="705.0">705 m</quantity>
depth, sample ID: PCB026; GenBank registration: ON774728;
<taxonomicName id="4C09609D165DFFF5410AFC946A79FC95" authority="MHN USC" authorityName="MHN USC" box="[870,1328,835,861]" family="Neoscopelidae" genus="Neoscopelus" kingdom="Animalia" order="Myctophiformes" pageId="4" pageNumber="491" phylum="Chordata" rank="species" species="macrolepidotus">
<emphasis id="B97DC70C165DFFF5410AFC946BE0FC95" box="[870,1193,835,861]" italics="true" pageId="4" pageNumber="491">Neoscopelus macrolepidotus</emphasis>
MHN USC
</taxonomicName>
25211-
<quantity id="4CF1B6FB165DFFF547E2FC956FBFFC49" metricMagnitude="0" metricUnit="m" metricValue="1.209" pageId="4" pageNumber="491" unit="mm" value="1209.0">1, 209 mm</quantity>
TL,
<date id="FFB73DDE165DFFF54343FCB06EAEFC49" box="[303,487,871,897]" pageId="4" pageNumber="491" value="2010-08-18">18 August 2010</date>
,
<collectingRegion id="49CDD5FC165DFFF54398FCB06D0CFC49" box="[500,581,871,897]" country="Spain" name="Galicia" pageId="4" pageNumber="491">Galicia</collectingRegion>
Bank,
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, -
<geoCoordinate id="EE3D7DD9165DFFF54175FCB06CC3FC49" box="[793,906,871,897]" degrees="11.689" direction="west" orientation="longitude" pageId="4" pageNumber="491" precision="55" value="-11.689">11.689ºW</geoCoordinate>
,
<quantity id="4CF1B6FB165DFFF541FBFCB06C96FC49" box="[919,991,871,897]" metricMagnitude="2" metricUnit="m" metricValue="7.659999999999999" pageId="4" pageNumber="491" unit="m" value="766.0">766 m</quantity>
depth, sample ID:
<collectionCode id="ED1883DB165DFFF546D7FCB06BB2FC49" box="[1211,1275,871,897]" country="Canada" httpUri="http://grbio.org/cool/zeys-ppzt" name="Nova Scotia Museum of Natural History" pageId="4" pageNumber="491">NSM</collectionCode>
001; GenBank registration: OR076049;
<collectionCode id="ED1883DB165DFFF543DDFC5C6EBBFC6D" box="[433,498,907,933]" pageId="4" pageNumber="491">MHN</collectionCode>
USC 25211-
<quantity id="4CF1B6FB165DFFF540E5FC5D6C48FC6D" box="[649,769,906,933]" metricMagnitude="0" metricUnit="m" metricValue="2.167" pageId="4" pageNumber="491" unit="mm" value="2167.0">2, 167 mm</quantity>
TL,
<date id="FFB73DDE165DFFF54155FC5C6C9DFC6D" box="[825,980,907,933]" pageId="4" pageNumber="491" value="2022-06-16">16 June 2022</date>
; Grand Banks,
<geoCoordinate id="EE3D7DD9165DFFF546EEFC5C6A4CFC6D" box="[1154,1285,907,933]" degrees="43.1067" direction="north" orientation="latitude" pageId="4" pageNumber="491" precision="5" value="43.1067">43.1067 ºN</geoCoordinate>
, -
<geoCoordinate id="EE3D7DD9165DFFF5477BFC5C6AD1FC6D" box="[1303,1432,907,933]" degrees="49.4962" direction="west" orientation="longitude" pageId="4" pageNumber="491" precision="5" value="-49.4962">49.4962ºW</geoCoordinate>
,
<quantity id="4CF1B6FB165DFFF542FBFC786F97FC01" box="[151,222,943,969]" metricMagnitude="2" metricUnit="m" metricValue="7.029999999999999" pageId="4" pageNumber="491" unit="m" value="703.0">703 m</quantity>
depth, ample ID:
<collectionCode id="ED1883DB165DFFF543C5FC786EA0FC01" box="[425,489,943,969]" country="Canada" httpUri="http://grbio.org/cool/zeys-ppzt" name="Nova Scotia Museum of Natural History" pageId="4" pageNumber="491">NSM</collectionCode>
003; GenBank registration: OR076050 (
<tableCitation id="C68B2EA5165DFFF541CBFC786CB2FC01" box="[935,1019,943,969]" captionStart="TABLE 1" captionStartId="5.[151,239,151,175]" captionTargetPageId="5" captionText="TABLE 1. Morphometric and meristic data of Neoscopelus serranoi sp. nov. based on the examination of 35 specimens from the NE Atlantic (holotype and 34 paratypes) compared with specimens of Neoscopelus microchir from the Western Pacific based on Matsubara (1943) (holotype and 37 paratypes)." pageId="4" pageNumber="491">Table 1</tableCitation>
, Appendix 2).
</paragraph>
<paragraph id="8BB61B1E165DFFF542ABFC056E30FBFD" blockId="4.[151,1437,474,2013]" pageId="4" pageNumber="491">
<emphasis id="B97DC70C165DFFF542ABFC056E9FFC24" bold="true" box="[199,470,978,1004]" pageId="4" pageNumber="491">Material comparative.</emphasis>
<taxonomicName id="4C09609D165DFFF5438DFC046C76FC25" authority="MHN USC" authorityName="MHN USC" box="[481,831,979,1005]" family="Neoscopelidae" genus="Neoscopelus" kingdom="Animalia" order="Myctophiformes" pageId="4" pageNumber="491" phylum="Chordata" rank="species" species="macrolepidotus">
<emphasis id="B97DC70C165DFFF5438DFC046DFBFC25" box="[481,690,979,1005]" italics="true" pageId="4" pageNumber="491">N. macrolepidotus</emphasis>
MHN USC
</taxonomicName>
25211-
<quantity id="4CF1B6FB165DFFF541CCFC056B50FC25" box="[928,1049,978,1005]" metricMagnitude="0" metricUnit="m" metricValue="1.209" pageId="4" pageNumber="491" unit="mm" value="1209.0">1, 209 mm</quantity>
TL,
<date id="FFB73DDE165DFFF5463AFC046A5BFC25" box="[1110,1298,979,1005]" pageId="4" pageNumber="491" value="2010-08-18">18 August 2010</date>
,
<geoCoordinate id="EE3D7DD9165DFFF5474DFC046AD1FC25" box="[1313,1432,979,1005]" degrees="42.687" direction="north" orientation="latitude" pageId="4" pageNumber="491" precision="55" value="42.687">42.687 ºN</geoCoordinate>
, -
<geoCoordinate id="EE3D7DD9165DFFF542F3FC206E50FBD9" box="[159,281,1015,1041]" degrees="11.689" direction="west" orientation="longitude" pageId="4" pageNumber="491" precision="55" value="-11.689">11.689 ºW</geoCoordinate>
,
<quantity id="4CF1B6FB165DFFF54348FC206E24FBD9" box="[292,365,1015,1041]" metricMagnitude="2" metricUnit="m" metricValue="7.659999999999999" pageId="4" pageNumber="491" unit="m" value="766.0">766 m</quantity>
depth;
<taxonomicName id="4C09609D165DFFF543AFFC206C53FBD9" authority="MHN USC" authorityName="MHN USC" box="[451,794,1015,1041]" family="Neoscopelidae" genus="Neoscopelus" kingdom="Animalia" order="Myctophiformes" pageId="4" pageNumber="491" phylum="Chordata" rank="species" species="macrolepidotus">
<emphasis id="B97DC70C165DFFF543AFFC206DDBFBD9" box="[451,658,1015,1041]" italics="true" pageId="4" pageNumber="491">N. macrolepidotus</emphasis>
MHN USC
</taxonomicName>
25211-
<quantity id="4CF1B6FB165DFFF5411FFC216CA5FBD9" box="[883,1004,1014,1041]" metricMagnitude="0" metricUnit="m" metricValue="2.167" pageId="4" pageNumber="491" unit="mm" value="2167.0">2, 167 mm</quantity>
TL,
<date id="FFB73DDE165DFFF54649FC206BF6FBD9" box="[1061,1215,1015,1041]" pageId="4" pageNumber="491" value="2022-06-16">16 June 2022</date>
,
<geoCoordinate id="EE3D7DD9165DFFF546A0FC206A55FBD9" box="[1228,1308,1015,1041]" degrees="43.106" direction="north" orientation="latitude" pageId="4" pageNumber="491" precision="55" value="43.106">43.106</geoCoordinate>
,
<geoCoordinate id="EE3D7DD9165DFFF54744FC206A37FBD9" box="[1320,1406,1015,1041]" degrees="49.496" direction="west" orientation="longitude" pageId="4" pageNumber="491" precision="55" value="-49.496">
-
<quantity id="4CF1B6FB165DFFF54743FC206A37FBD9" box="[1327,1406,1015,1041]" metricMagnitude="1" metricUnit="m" metricValue="4.9496" pageId="4" pageNumber="491" unit="m" value="49.496">49.496</quantity>
</geoCoordinate>
m depth (Appendix 3).
</paragraph>
</subSubSection>
<subSubSection id="C3134895165DFFF742ABFBE96B23FF1D" lastPageId="6" lastPageNumber="493" pageId="4" pageNumber="491" type="diagnosis">
<paragraph id="8BB61B1E165DFFF542ABFBE96CFFFB21" blockId="4.[151,1437,474,2013]" pageId="4" pageNumber="491">
<emphasis id="B97DC70C165DFFF542ABFBE96E0BFB90" bold="true" box="[199,322,1086,1112]" pageId="4" pageNumber="491">Diagnosis.</emphasis>
The new species belongs to the genus
<taxonomicName id="4C09609D165DFFF5417CFBE86CD7FB91" box="[784,926,1087,1113]" family="Neoscopelidae" genus="Neoscopelus" kingdom="Animalia" order="Myctophiformes" pageId="4" pageNumber="491" phylum="Chordata" rank="genus">
<emphasis id="B97DC70C165DFFF5417CFBE86CD7FB91" box="[784,926,1087,1113]" italics="true" pageId="4" pageNumber="491">Neoscopelus</emphasis>
</taxonomicName>
as defined by
<bibRefCitation id="EF9866EF165DFFF54622FBE86A56FB91" author="Nafpaktitis, B. G." box="[1102,1311,1087,1113]" pageId="4" pageNumber="491" refId="ref11273" refString="Nafpaktitis, B. G. (1977) Family Neoscopelidae. In: Gibbs, R. H., Berry, F. H., Bohlke, J. E., Cohen, D. M., Collette, B. B., Eschmeyer, W. N., Mead, G. W., Merriman, D., Pietsch, T. W. &amp; Parr, A. E. (Eds.), Fishes of the western North Atlantic. Vol. 7. Yale University, New Haven, Connecticut, pp. 1 - 12." type="book" year="1977">Nafpaktitis (1977)</bibRefCitation>
by having a body fusiform and compressed; head conical and large; large jaws, extended to the back of the orbit; dentary, premaxilla, palatine, and mesopterygoid with villiform teeth; large eyes, its diameter contained 5 times or less in the head length; elongate pectorals, extending posteriorly beyond the anus; adipose fin present; ventrolateral rows of photophores and a series of light organs on the periphery of the tongue.
</paragraph>
<paragraph id="8BB61B1E165DFFF542ABFB256C50FAB1" blockId="4.[151,1437,474,2013]" pageId="4" pageNumber="491">
<emphasis id="B97DC70C165DFFF542ABFB256E99FAC4" bold="true" box="[199,464,1266,1292]" pageId="4" pageNumber="491">Differential diagnosis.</emphasis>
<taxonomicName id="4C09609D165DFFF543B5FB246D86FAC4" authorityName="Bañón &amp; Barros-García &amp; Arronte &amp; Rábade &amp; Rio &amp; Baldó &amp; Carlos" authorityYear="2024" box="[473,719,1267,1293]" family="Neoscopelidae" genus="Neoscopelus" kingdom="Animalia" order="Myctophiformes" pageId="4" pageNumber="491" phylum="Chordata" rank="species" species="serranoi" status="sp. nov.">
<emphasis id="B97DC70C165DFFF543B5FB246D86FAC4" box="[473,719,1267,1293]" italics="true" pageId="4" pageNumber="491">Neoscopelus serranoi</emphasis>
</taxonomicName>
<emphasis id="B97DC70C165DFFF540B4FB246C7DFAC5" bold="true" box="[728,820,1267,1293]" pageId="4" pageNumber="491">
<taxonomicNameLabel id="A24E7A77165DFFF540B4FB246C7DFAC5" box="[728,820,1267,1293]" pageId="4" pageNumber="491" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is morphologically very similar to
<taxonomicName id="4C09609D165DFFF546A0FB246A15FAC5" box="[1228,1372,1267,1293]" family="Neoscopelidae" genus="Neoscopelus" kingdom="Animalia" order="Myctophiformes" pageId="4" pageNumber="491" phylum="Chordata" rank="species" species="microchir">
<emphasis id="B97DC70C165DFFF546A0FB246A15FAC5" box="[1228,1372,1267,1293]" italics="true" pageId="4" pageNumber="491">N. microchir</emphasis>
</taxonomicName>
from which it differs subtly.
<taxonomicName id="4C09609D165DFFF543CEFAC06DD1FAF8" authorityName="Bañón &amp; Barros-García &amp; Arronte &amp; Rábade &amp; Rio &amp; Baldó &amp; Carlos" authorityYear="2024" box="[418,664,1303,1329]" family="Neoscopelidae" genus="Neoscopelus" kingdom="Animalia" order="Myctophiformes" pageId="4" pageNumber="491" phylum="Chordata" rank="species" species="serranoi" status="sp. nov.">
<emphasis id="B97DC70C165DFFF543CEFAC06DD1FAF8" box="[418,664,1303,1329]" italics="true" pageId="4" pageNumber="491">Neoscopelus serranoi</emphasis>
</taxonomicName>
<emphasis id="B97DC70C165DFFF540CDFAC06DB5FAF9" bold="true" box="[673,764,1303,1329]" pageId="4" pageNumber="491">
<taxonomicNameLabel id="A24E7A77165DFFF540CDFAC06DB5FAF9" box="[673,764,1303,1329]" pageId="4" pageNumber="491" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<typeStatus id="54B2A5BC165DFFF541FBFAC06C8EFAF9" box="[919,967,1303,1329]" pageId="4" pageNumber="491">type</typeStatus>
specimens of
<taxonomicName id="4C09609D165DFFF54603FAC06BB7FAF9" box="[1135,1278,1303,1329]" family="Neoscopelidae" genus="Neoscopelus" kingdom="Animalia" order="Myctophiformes" pageId="4" pageNumber="491" phylum="Chordata" rank="species" species="microchir">
<emphasis id="B97DC70C165DFFF54603FAC06BB7FAF9" box="[1135,1278,1303,1329]" italics="true" pageId="4" pageNumber="491">N. microchir</emphasis>
</taxonomicName>
, according to
<bibRefCitation id="EF9866EF165DFFF542FBFAEC6E29FA9D" author="Matsubara, K." box="[151,352,1339,1365]" pageId="4" pageNumber="491" pagination="37 - 82" refId="ref10823" refString="Matsubara, K. (1943) Ichthyological annotations from the depth of the Sea of Japan, I-VII. Journal of the Sigenkagaku Kenkyusyo, 1, 37 - 82." type="journal article" year="1943">Matsubara (1943)</bibRefCitation>
, in having a shorter anal-fin base (25.734 vs 33.743.7% HL) and a shorter pelvic fin (40.459.6, mean±95% IC=50.1±1.78 vs 61.365.8% HL). (
<tableCitation id="C68B2EA5165DFFF540D8FA886C41FAB1" box="[692,776,1375,1401]" captionStart="TABLE 1" captionStartId="5.[151,239,151,175]" captionTargetPageId="5" captionText="TABLE 1. Morphometric and meristic data of Neoscopelus serranoi sp. nov. based on the examination of 35 specimens from the NE Atlantic (holotype and 34 paratypes) compared with specimens of Neoscopelus microchir from the Western Pacific based on Matsubara (1943) (holotype and 37 paratypes)." pageId="4" pageNumber="491">Table 1</tableCitation>
).
</paragraph>
<paragraph id="8BB61B1E165DFFF542ABFA546EBCFA09" blockId="4.[151,1437,474,2013]" pageId="4" pageNumber="491">
It differs from
<taxonomicName id="4C09609D165DFFF54307FA546D70FA55" box="[363,569,1411,1437]" family="Neoscopelidae" genus="Neoscopelus" kingdom="Animalia" order="Myctophiformes" pageId="4" pageNumber="491" phylum="Chordata" rank="species" species="macrolepidotus">
<emphasis id="B97DC70C165DFFF54307FA546D70FA55" box="[363,569,1411,1437]" italics="true" pageId="4" pageNumber="491">N. macrolepidotus</emphasis>
</taxonomicName>
in having a shorter anal-fin base (25.734 vs 37.944.4 % HL), and more LO photophores (2024 vs 1215).
</paragraph>
<paragraph id="8BB61B1E165DFFF542ABFA1C6B66F9E5" blockId="4.[151,1437,474,2013]" pageId="4" pageNumber="491">
It differs from
<taxonomicName id="4C09609D165DFFF5431AFA1C6EBBFA2C" authorityName="Arai" authorityYear="1969" box="[374,498,1483,1508]" family="Neoscopelidae" genus="Neoscopelus" kingdom="Animalia" order="Myctophiformes" pageId="4" pageNumber="491" phylum="Chordata" rank="species" species="porosus">
<emphasis id="B97DC70C165DFFF5431AFA1C6EBBFA2C" box="[374,498,1483,1508]" italics="true" pageId="4" pageNumber="491">N. porosus</emphasis>
</taxonomicName>
in having a greater eye (20.328.8 vs 17.421.6% HL), longer pre-anal distance (72.279 vs 64.571.2% SL), shorter anal-fin base (25.734 vs 44.851.8 % HL), fewer gillrakers (1415 vs 17 22), and fewer LO photophores (2024 vs 3540) (
<bibRefCitation id="EF9866EF165DFFF540BFF9C46C06F9E5" author="Arai, R." box="[723,847,1555,1581]" pageId="4" pageNumber="491" pagination="465 - 471" refId="ref7997" refString="Arai, R. (1969) A new iniomous fish of the genus Neoscopelus from Suruga Bay, Japan. Bulletin of the National Museum of Nature and Science, Tokyo, 12 (3), 465 - 471." type="journal article" year="1969">Arai, 1969</bibRefCitation>
;
<bibRefCitation id="EF9866EF165DFFF54136F9C46B6BF9E5" author="Wada, H. &amp; Ohtomi, J. &amp; Motomura, H." box="[858,1058,1555,1581]" pageId="4" pageNumber="491" pagination="24 - 30" refId="ref12630" refString="Wada, H., Ohtomi, J. &amp; Motomura, H. (2021) The northernmost and second Japanese records of Diretmoides veriginae (Beryciformes: Diretmidae) from Suruga Bay and off the Satsuma Peninsula, and the first records of Neoscopelus porosus (Myctophiformes: Neoscopelidae) from Kagoshima Prefecture. Ichthy, Natural History of Fishes of Japan, 8, 24 - 30. https: // doi. org / 10.34583 / ichthy. 8.0 _ 24" type="journal article" year="2021">
Wada
<emphasis id="B97DC70C165DFFF541CCF9C46C9BF9E5" box="[928,978,1555,1581]" italics="true" pageId="4" pageNumber="491">et al</emphasis>
., 2021
</bibRefCitation>
).
</paragraph>
<paragraph id="8BB61B1E165DFFF542ABF9E16A11F9BD" blockId="4.[151,1437,474,2013]" pageId="4" pageNumber="491">
<emphasis id="B97DC70C165DFFF542ABF9E16EABF998" bold="true" box="[199,482,1590,1616]" pageId="4" pageNumber="491">Morphological revision.</emphasis>
A revision of the morphological characters reported in the ichthyological literature for the two Atlantic species,
<taxonomicName id="4C09609D165DFFF543DCF98C6D37F9BD" box="[432,638,1627,1653]" family="Neoscopelidae" genus="Neoscopelus" kingdom="Animalia" order="Myctophiformes" pageId="4" pageNumber="491" phylum="Chordata" rank="species" species="macrolepidotus">
<emphasis id="B97DC70C165DFFF543DCF98C6D37F9BD" box="[432,638,1627,1653]" italics="true" pageId="4" pageNumber="491">N. macrolepidotus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C09609D165DFFF540D8F98C6C0BF9BD" box="[692,834,1627,1653]" family="Neoscopelidae" genus="Neoscopelus" kingdom="Animalia" order="Myctophiformes" pageId="4" pageNumber="491" phylum="Chordata" rank="species" species="microchir">
<emphasis id="B97DC70C165DFFF540D8F98C6C0BF9BD" box="[692,834,1627,1653]" italics="true" pageId="4" pageNumber="491">N. microchir</emphasis>
</taxonomicName>
is presented in the
<tableCitation id="C68B2EA5165DFFF54677F98C6B31F9BD" box="[1051,1144,1627,1653]" captionStart="TABLE 2" captionStartId="6.[151,239,1145,1169]" captionTargetPageId="6" captionText="TABLE 2. Differences in meristic and morphometric characters in Neoscopelus macrolepidotus specimens between the Western North Pacific (Jordan &amp; Starks, 1904; Matsubara, 1943; Wang &amp; Shao, 2006; Tatsuta et al., 2014), Western Atlantic (Nafpaktitis, 1977; Uyeno et al., 1983; Rodríguez &amp; Sang, 1986; Roa-Varón et al., 2007) and Eastern Atlantic (Johnson, 1863; Maul, 1951; Lloris, 1986; Bañón-Díaz et al., 1997; Ordines et al., 2017) in percentage of standard length (%SL) and/or head length (%HL)." pageId="4" pageNumber="491">Tables 2</tableCitation>
and
<tableCitation id="C68B2EA5165DFFF546C3F98C6BF7F9BD" box="[1199,1214,1627,1653]" captionStart="TABLE 3" captionStartId="7.[151,239,715,739]" captionTargetPageId="7" captionText="TABLE 3. Differences in meristic and morphometric characters in Neoscopelus microchir specimens between the Western North Pacific (Matsubara 1943), Western Atlantic (Nafpaktitis, 1977; Roa-Varón et al., 2007) and Eastern Atlantic (Maul, 1951, 1976; Furnestin, 1958; Bañón et al., 2002, 2022) in percentage of standard length (%SL) and/or head length (%HL)." pageId="4" pageNumber="491">3</tableCitation>
, respectively.
</paragraph>
<paragraph id="8BB61B1E165DFFF542ABF9A96C31F929" blockId="4.[151,1437,474,2013]" pageId="4" pageNumber="491">
<emphasis id="B97DC70C165DFFF542ABF9A96E8BF950" bold="true" box="[199,450,1662,1688]" pageId="4" pageNumber="491">Species Delimitation.</emphasis>
The results of the DNA-based species delimitation analyses are summarised in
<figureCitation id="1332079B165DFFF54724F9A86FEEF975" captionStart="FIGURE 2" captionStartId="8.[151,250,1679,1703]" captionTargetBox="[252,1333,190,1647]" captionTargetId="figure-19@8.[248,1340,181,1655]" captionTargetPageId="8" captionText="FIGURE 2. Consensus phylogeny (Maximum Likelihood and Bayesian Inference) of Neoscopelus based on COI gene sequences with the results of six different molecular approaches: BIN, Barcode Index Number; ASAP, Assemble Species by Automatic Partitioning; bPTP, Bayesian Poisson Tree Process; mPTP, multiple Poisson Tree Process; GMYC, General Mixed Yule-Coalescent model and mGMYC, multiple General Mixed Yule-Coalescent model. The proposed Molecular Operational Taxonomic Units (MOTUs) for each analysis are represented as boxes. The colours in the branches indicate the proposed MOTUs in this study. In the first node of each MOTU, both boostrap values (&gt;75) and posterior probabilities (&gt;0.9) are shown. Each individual contains the putative morphospecies, repository and geographic codes: NEA—Northeast Atlantic; JAM— Jamaica; ESA—Eastern South Africa; TWN—Taiwan; WAU—Western Australia; SCS—South China Sea; IO—Indian Ocean; PNG—Papua New Guinea; ESA—Eastern South Africa; TAS—Tasmania; GME—Gulf of Mexico; NZL—New Zealand; NWA—Northwest Atlantic." figureDoi="http://doi.org/10.5281/zenodo.14022811" httpUri="https://zenodo.org/record/14022811/files/figure.png" pageId="4" pageNumber="491">Figures 2</figureCitation>
,
<figureCitation id="1332079B165DFFF542D8F9746F8AF975" box="[180,195,1699,1725]" captionStart="FIGURE 3" captionStartId="9.[151,250,843,867]" captionTargetBox="[286,1300,184,814]" captionTargetId="figure-17@9.[282,1305,181,819]" captionTargetPageId="9" captionText="FIGURE 3. Map of sampling locations of the Neoscopelus microchir species complex and the proposed Neoscopelus serranoi sp. nov. in red. The diameter of the circle is indicative of the sample size, and the colours correspond to the MOTUs detected in the phylogenetic analysis." figureDoi="http://doi.org/10.5281/zenodo.14022813" httpUri="https://zenodo.org/record/14022813/files/figure.png" pageId="4" pageNumber="491">3</figureCitation>
,
<figureCitation id="1332079B165DFFF542BDF9746F96F975" box="[209,223,1699,1725]" captionStart="FIGURE 4" captionStartId="9.[151,250,1661,1685]" captionTargetBox="[286,1300,1002,1632]" captionTargetId="figure-64@9.[282,1305,999,1637]" captionTargetPageId="9" captionText="FIGURE 4. Map of sampling locations of the Neoscopelus macrolepidotus species complex and the single specimen of Neoscopelus porosus (in purple). The diameter of the circle is indicative of the sample size, and the colours correspond to the MOTUs detected in the phylogenetic analysis." figureDoi="http://doi.org/10.5281/zenodo.14022815" httpUri="https://zenodo.org/record/14022815/files/figure.png" pageId="4" pageNumber="491">4</figureCitation>
and Appendix 4. All analyses have confirmed hidden diversity within the
<taxonomicName id="4C09609D165DFFF54644F9746A46F975" box="[1064,1295,1699,1725]" pageId="4" pageNumber="491">
<emphasis id="B97DC70C165DFFF54644F9746BFFF975" box="[1064,1206,1699,1725]" italics="true" pageId="4" pageNumber="491">Neoscopelus</emphasis>
species
</taxonomicName>
except for a single individual of
<taxonomicName id="4C09609D165DFFF54314F9106EB8F928" authorityName="Arai" authorityYear="1969" box="[376,497,1735,1760]" family="Neoscopelidae" genus="Neoscopelus" kingdom="Animalia" order="Myctophiformes" pageId="4" pageNumber="491" phylum="Chordata" rank="species" species="porosus">
<emphasis id="B97DC70C165DFFF54314F9106EB8F928" box="[376,497,1735,1760]" italics="true" pageId="4" pageNumber="491">N. porosus</emphasis>
</taxonomicName>
(GBGCA 11195-15) from
<collectingCountry id="F31E5B8E165DFFF54173F9106C3CF929" box="[799,885,1735,1761]" name="Taiwan" pageId="4" pageNumber="491">Taiwan</collectingCountry>
.
</paragraph>
<paragraph id="8BB61B1E165DFFF542ABF93C6D4BF814" blockId="4.[151,1437,474,2013]" pageId="4" pageNumber="491">
The new
<emphasis id="B97DC70C165DFFF54346F93C6E13F8CD" box="[298,346,1771,1797]" italics="true" pageId="4" pageNumber="491">COI</emphasis>
sequences labelled as
<taxonomicName id="4C09609D165DFFF54020F93C6C75F8CC" authorityName="Bañón &amp; Barros-García &amp; Arronte &amp; Rábade &amp; Rio &amp; Baldó &amp; Carlos" authorityYear="2024" box="[588,828,1771,1797]" family="Neoscopelidae" genus="Neoscopelus" kingdom="Animalia" order="Myctophiformes" pageId="4" pageNumber="491" phylum="Chordata" rank="species" species="serranoi" status="sp. nov.">
<emphasis id="B97DC70C165DFFF54020F93C6C75F8CC" box="[588,828,1771,1797]" italics="true" pageId="4" pageNumber="491">Neoscopelus serranoi</emphasis>
</taxonomicName>
<emphasis id="B97DC70C165DFFF5412CF93C6CDEF8CD" bold="true" box="[832,919,1771,1797]" pageId="4" pageNumber="491">
<taxonomicNameLabel id="A24E7A77165DFFF5412CF93C6CDEF8CD" box="[832,919,1771,1797]" pageId="4" pageNumber="491" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
were all assembled into an independent MOTU in four of the six analyses (BIN, ASAP, bPTP and mPTP) including four individuals mined from the repositories and assigned to
<taxonomicName id="4C09609D165DFFF5433BF8E46EA1F885" box="[343,488,1843,1869]" family="Neoscopelidae" genus="Neoscopelus" kingdom="Animalia" order="Myctophiformes" pageId="4" pageNumber="491" phylum="Chordata" rank="species" species="microchir">
<emphasis id="B97DC70C165DFFF5433BF8E46EA1F885" box="[343,488,1843,1869]" italics="true" pageId="4" pageNumber="491">N. microchir</emphasis>
</taxonomicName>
(MFLE001-12 from
<collectingCountry id="F31E5B8E165DFFF54088F8E46C08F885" box="[740,833,1843,1869]" name="Jamaica" pageId="4" pageNumber="491">Jamaica</collectingCountry>
, KF489668KF489669 from
<collectingCountry id="F31E5B8E165DFFF546F6F8E46A64F885" box="[1178,1325,1843,1869]" name="South Africa" pageId="4" pageNumber="491">South Africa</collectingCountry>
Southern Indian Ocean and PORCU026-22 from Northeast Atlantic) (
<figureCitation id="1332079B165DFFF54152F8806CE2F8B9" box="[830,939,1879,1905]" captionStart="FIGURE 2" captionStartId="8.[151,250,1679,1703]" captionTargetBox="[252,1333,190,1647]" captionTargetId="figure-19@8.[248,1340,181,1655]" captionTargetPageId="8" captionText="FIGURE 2. Consensus phylogeny (Maximum Likelihood and Bayesian Inference) of Neoscopelus based on COI gene sequences with the results of six different molecular approaches: BIN, Barcode Index Number; ASAP, Assemble Species by Automatic Partitioning; bPTP, Bayesian Poisson Tree Process; mPTP, multiple Poisson Tree Process; GMYC, General Mixed Yule-Coalescent model and mGMYC, multiple General Mixed Yule-Coalescent model. The proposed Molecular Operational Taxonomic Units (MOTUs) for each analysis are represented as boxes. The colours in the branches indicate the proposed MOTUs in this study. In the first node of each MOTU, both boostrap values (&gt;75) and posterior probabilities (&gt;0.9) are shown. Each individual contains the putative morphospecies, repository and geographic codes: NEA—Northeast Atlantic; JAM— Jamaica; ESA—Eastern South Africa; TWN—Taiwan; WAU—Western Australia; SCS—South China Sea; IO—Indian Ocean; PNG—Papua New Guinea; ESA—Eastern South Africa; TAS—Tasmania; GME—Gulf of Mexico; NZL—New Zealand; NWA—Northwest Atlantic." figureDoi="http://doi.org/10.5281/zenodo.14022811" httpUri="https://zenodo.org/record/14022811/files/figure.png" pageId="4" pageNumber="491">Figures 2</figureCitation>
,
<figureCitation id="1332079B165DFFF541D4F8806C81F8B9" box="[952,968,1879,1905]" captionStart="FIGURE 3" captionStartId="9.[151,250,843,867]" captionTargetBox="[286,1300,184,814]" captionTargetId="figure-17@9.[282,1305,181,819]" captionTargetPageId="9" captionText="FIGURE 3. Map of sampling locations of the Neoscopelus microchir species complex and the proposed Neoscopelus serranoi sp. nov. in red. The diameter of the circle is indicative of the sample size, and the colours correspond to the MOTUs detected in the phylogenetic analysis." figureDoi="http://doi.org/10.5281/zenodo.14022813" httpUri="https://zenodo.org/record/14022813/files/figure.png" pageId="4" pageNumber="491">3</figureCitation>
). GMYC and mGMYC further subdivide this cluster in two groups, one with eighteen
<taxonomicName id="4C09609D165DFFF540F4F8AC6CC5F85C" authorityName="Bañón &amp; Barros-García &amp; Arronte &amp; Rábade &amp; Rio &amp; Baldó &amp; Carlos" authorityYear="2024" box="[664,908,1915,1941]" family="Neoscopelidae" genus="Neoscopelus" kingdom="Animalia" order="Myctophiformes" pageId="4" pageNumber="491" phylum="Chordata" rank="species" species="serranoi" status="sp. nov.">
<emphasis id="B97DC70C165DFFF540F4F8AC6CC5F85C" box="[664,908,1915,1941]" italics="true" pageId="4" pageNumber="491">Neoscopelus serranoi</emphasis>
</taxonomicName>
<emphasis id="B97DC70C165DFFF541F8F8AC6CA6F85D" bold="true" box="[916,1007,1915,1941]" pageId="4" pageNumber="491">
<taxonomicNameLabel id="A24E7A77165DFFF541F8F8AC6CA6F85D" box="[916,1007,1915,1941]" pageId="4" pageNumber="491" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<specimenCount id="9D0FD097165DFFF54644F8AC6B3AF85C" box="[1064,1139,1915,1941]" count="1" pageId="4" pageNumber="491" type="generic">
one
<taxonomicName id="4C09609D165DFFF54634F8AC6B3AF85C" box="[1112,1139,1915,1940]" family="Neoscopelidae" genus="Neoscopelus" kingdom="Animalia" order="Myctophiformes" pageId="4" pageNumber="491" phylum="Chordata" rank="species" species="microchir">
<emphasis id="B97DC70C165DFFF54634F8AC6B3AF85C" box="[1112,1139,1915,1940]" italics="true" pageId="4" pageNumber="491">N.</emphasis>
</taxonomicName>
</specimenCount>
microchir (MFLE001-12) sequences and the second with eight
<taxonomicName id="4C09609D165DFFF54059F8486C60F870" authorityName="Bañón &amp; Barros-García &amp; Arronte &amp; Rábade &amp; Rio &amp; Baldó &amp; Carlos" authorityYear="2024" box="[565,809,1951,1977]" family="Neoscopelidae" genus="Neoscopelus" kingdom="Animalia" order="Myctophiformes" pageId="4" pageNumber="491" phylum="Chordata" rank="species" species="serranoi" status="sp. nov.">
<emphasis id="B97DC70C165DFFF54059F8486C60F870" box="[565,809,1951,1977]" italics="true" pageId="4" pageNumber="491">Neoscopelus serranoi</emphasis>
</taxonomicName>
<emphasis id="B97DC70C165DFFF5415CF8486CC3F871" bold="true" box="[816,906,1951,1977]" pageId="4" pageNumber="491">
<taxonomicNameLabel id="A24E7A77165DFFF5415CF8486CC3F871" box="[816,906,1951,1977]" pageId="4" pageNumber="491" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<specimenCount id="9D0FD097165DFFF541ACF8486B53F870" box="[960,1050,1951,1977]" count="3" pageId="4" pageNumber="491" type="generic">
three
<taxonomicName id="4C09609D165DFFF54193F8486B53F870" box="[1023,1050,1951,1976]" family="Neoscopelidae" genus="Neoscopelus" kingdom="Animalia" order="Myctophiformes" pageId="4" pageNumber="491" phylum="Chordata" rank="species" species="microchir">
<emphasis id="B97DC70C165DFFF54193F8486B53F870" box="[1023,1050,1951,1976]" italics="true" pageId="4" pageNumber="491">N.</emphasis>
</taxonomicName>
</specimenCount>
microchir sequences (KF489668 KF489669 and PORCU026-22).
</paragraph>
<caption id="DF764B96165CFFF442FBFF406C0DFF30" pageId="5" pageNumber="492" startId="5.[151,239,151,175]" targetBox="[163,1403,263,1575]" targetIsTable="true" targetPageId="5" targetType="table">
<paragraph id="8BB61B1E165CFFF442FBFF406C0DFF30" blockId="5.[151,1437,151,248]" pageId="5" pageNumber="492">
<emphasis id="B97DC70C165CFFF442FBFF406E43FF67" bold="true" box="[151,266,151,175]" pageId="5" pageNumber="492">TABLE 1.</emphasis>
Morphometric and meristic data of
<taxonomicName id="4C09609D165CFFF440E0FF406C3DFF67" authorityName="Bañón &amp; Barros-García &amp; Arronte &amp; Rábade &amp; Rio &amp; Baldó &amp; Carlos" authorityYear="2024" box="[652,884,151,175]" family="Neoscopelidae" genus="Neoscopelus" kingdom="Animalia" order="Myctophiformes" pageId="5" pageNumber="492" phylum="Chordata" rank="species" species="serranoi" status="sp. nov.">
<emphasis id="B97DC70C165CFFF440E0FF406C3DFF67" box="[652,884,151,175]" italics="true" pageId="5" pageNumber="492">Neoscopelus serranoi</emphasis>
</taxonomicName>
<emphasis id="B97DC70C165CFFF44117FF4F6C98FF78" bold="true" box="[891,977,152,176]" pageId="5" pageNumber="492">
<taxonomicNameLabel id="A24E7A77165CFFF44117FF4F6C98FF78" box="[891,977,152,176]" pageId="5" pageNumber="492" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
based on the examination of 35 specimens from the NE Atlantic (holotype and 34 paratypes) compared with specimens of
<taxonomicName id="4C09609D165CFFF44184FF6C6B94FF1B" box="[1000,1245,187,211]" family="Neoscopelidae" genus="Neoscopelus" kingdom="Animalia" order="Myctophiformes" pageId="5" pageNumber="492" phylum="Chordata" rank="species" species="microchir">
<emphasis id="B97DC70C165CFFF44184FF6C6B94FF1B" box="[1000,1245,187,211]" italics="true" pageId="5" pageNumber="492">Neoscopelus microchir</emphasis>
</taxonomicName>
from the Western Pacific based on
<bibRefCitation id="EF9866EF165CFFF44321FF086D42FF30" author="Matsubara, K." box="[333,523,223,248]" pageId="5" pageNumber="492" pagination="37 - 82" refId="ref10823" refString="Matsubara, K. (1943) Ichthyological annotations from the depth of the Sea of Japan, I-VII. Journal of the Sigenkagaku Kenkyusyo, 1, 37 - 82." type="journal article" year="1943">Matsubara (1943)</bibRefCitation>
(holotype and 37 paratypes).
</paragraph>
</caption>
<paragraph id="8BB61B1E165CFFF4401BFED06A2EF9EF" pageId="5" pageNumber="492">
<table id="F909E9BE165C000E42CFFED06A32F9EF" box="[163,1403,263,1575]" gridcols="6" gridrows="33" pageId="5" pageNumber="492">
<tr id="3539195C165C000E42CFFED06A32FED7" box="[163,1403,263,287]" gridrow="0" pageId="5" pageNumber="492" rowspan-0="1">
<th id="76E87020165C000E4062FED06B5BFED7" box="[526,1042,263,287]" colspan="3" colspanRight="2" gridcol="1" gridrow="0" pageId="5" pageNumber="492">
<emphasis id="B97DC70C165CFFF4401BFED06CE7FED7" bold="true" box="[631,942,263,287]" pageId="5" pageNumber="492">
<taxonomicName id="4C09609D165CFFF4401BFED06C1EFED7" authorityName="Bañón &amp; Barros-García &amp; Arronte &amp; Rábade &amp; Rio &amp; Baldó &amp; Carlos" authorityYear="2024" box="[631,855,263,287]" family="Neoscopelidae" genus="Neoscopelus" kingdom="Animalia" order="Myctophiformes" pageId="5" pageNumber="492" phylum="Chordata" rank="species" species="serranoi" status="sp. nov.">
<emphasis id="B97DC70C165CFFF4401BFED06C1EFED7" bold="true" box="[631,855,263,287]" italics="true" pageId="5" pageNumber="492">Neoscopelus serranoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E7A77165CFFF44131FED06CE7FED7" box="[861,942,263,287]" pageId="5" pageNumber="492" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<th id="76E87020165C000E4604FED06A32FED7" box="[1128,1403,263,287]" colspan="2" colspanRight="1" gridcol="4" gridrow="0" pageId="5" pageNumber="492">
<taxonomicName id="4C09609D165CFFF44618FED06A2DFED7" box="[1140,1380,263,287]" family="Neoscopelidae" genus="Neoscopelus" kingdom="Animalia" order="Myctophiformes" pageId="5" pageNumber="492" phylum="Chordata" rank="species" species="microchir">
<emphasis id="B97DC70C165CFFF44618FED06A2DFED7" bold="true" box="[1140,1380,263,287]" italics="true" pageId="5" pageNumber="492">Neoscopelus microchir</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="3539195C165C000E42CFFEE76A32FE80" box="[163,1403,304,328]" gridrow="1" pageId="5" pageNumber="492" rowspan-0="1">
<td id="76E87020165C000E4062FEE76D25FE80" box="[526,620,304,328]" gridcol="1" gridrow="1" pageId="5" pageNumber="492">Holotype</td>
<td id="76E87020165C000E40C8FEE76C3BFE80" box="[676,882,304,328]" gridcol="2" gridrow="1" pageId="5" pageNumber="492">Paratypes Range (n)</td>
<td id="76E87020165C000E41C1FEE76B5BFE80" box="[941,1042,304,328]" gridcol="3" gridrow="1" pageId="5" pageNumber="492">Mean±SE</td>
<td id="76E87020165C000E4604FEE76B8FFE80" box="[1128,1222,304,328]" gridcol="4" gridrow="1" pageId="5" pageNumber="492">Holotype</td>
<td id="76E87020165C000E4779FEE76A32FE80" box="[1301,1403,304,328]" gridcol="5" gridrow="1" pageId="5" pageNumber="492">Paratypes</td>
</tr>
<tr id="3539195C165C000E42CFFE8F6A32FEB8" box="[163,1403,344,368]" gridrow="2" pageId="5" pageNumber="492" rowspan-4="1" rowspan-5="1">
<th id="76E87020165C000E42CFFE8F6EE2FEB8" box="[163,427,344,368]" gridcol="0" gridrow="2" pageId="5" pageNumber="492">Total length (mm)</th>
<td id="76E87020165C000E4062FE8F6D25FEB8" box="[526,620,344,368]" gridcol="1" gridrow="2" pageId="5" pageNumber="492">236</td>
<td id="76E87020165C000E40C8FE8F6C3BFEB8" box="[676,882,344,368]" gridcol="2" gridrow="2" pageId="5" pageNumber="492">187323(34)</td>
<td id="76E87020165C000E41C1FE8F6B5BFEB8" box="[941,1042,344,368]" gridcol="3" gridrow="2" pageId="5" pageNumber="492">247±5.7</td>
</tr>
<tr id="3539195C165C000E42CFFE576A32FE50" box="[163,1403,384,408]" gridrow="3" pageId="5" pageNumber="492" rowspan-5="1">
<th id="76E87020165C000E42CFFE576EE2FE50" box="[163,427,384,408]" gridcol="0" gridrow="3" pageId="5" pageNumber="492">Standard length (mm)</th>
<td id="76E87020165C000E4062FE576D25FE50" box="[526,620,384,408]" gridcol="1" gridrow="3" pageId="5" pageNumber="492">192</td>
<td id="76E87020165C000E40C8FE576C3BFE50" box="[676,882,384,408]" gridcol="2" gridrow="3" pageId="5" pageNumber="492">150259(34)</td>
<td id="76E87020165C000E41C1FE576B5BFE50" box="[941,1042,384,408]" gridcol="3" gridrow="3" pageId="5" pageNumber="492">198±4.9</td>
<td id="76E87020165C000E4604FE576B8FFE50" box="[1128,1222,384,408]" gridcol="4" gridrow="3" pageId="5" pageNumber="492">146.8</td>
</tr>
<tr id="3539195C165C000E42CFFE7F6A32FE08" box="[163,1403,424,448]" gridrow="4" pageId="5" pageNumber="492">
<th id="76E87020165C000E42CFFE7F6A32FE08" box="[163,1403,424,448]" colspan="6" colspanRight="5" gridcol="0" gridrow="4" pageId="5" pageNumber="492">
<emphasis id="B97DC70C165CFFF442CFFE7E6FB0FE08" box="[163,249,424,448]" italics="true" pageId="5" pageNumber="492">As % SL</emphasis>
</th>
</tr>
<tr id="3539195C165C000E42CFFE076A32FE21" box="[163,1403,464,489]" gridrow="5" pageId="5" pageNumber="492">
<th id="76E87020165C000E42CFFE076EE2FE21" box="[163,427,464,489]" gridcol="0" gridrow="5" pageId="5" pageNumber="492">Head length</th>
<td id="76E87020165C000E4062FE076D25FE21" box="[526,620,464,489]" gridcol="1" gridrow="5" pageId="5" pageNumber="492">29.7</td>
<td id="76E87020165C000E40C8FE076C3BFE21" box="[676,882,464,489]" gridcol="2" gridrow="5" pageId="5" pageNumber="492">29.734.5(34)</td>
<td id="76E87020165C000E41C1FE076B5BFE21" box="[941,1042,464,489]" gridcol="3" gridrow="5" pageId="5" pageNumber="492">31.8±0.2</td>
<td id="76E87020165C000E4604FE076B8FFE21" box="[1128,1222,464,489]" gridcol="4" gridrow="5" pageId="5" pageNumber="492">30.3</td>
<td id="76E87020165C000E4779FE076A32FE21" box="[1301,1403,464,489]" gridcol="5" gridrow="5" pageId="5" pageNumber="492">28.233</td>
</tr>
<tr id="3539195C165C000E42CFFE2E6A32FDD9" box="[163,1403,505,529]" gridrow="6" pageId="5" pageNumber="492">
<th id="76E87020165C000E42CFFE2E6EE2FDD9" box="[163,427,505,529]" gridcol="0" gridrow="6" pageId="5" pageNumber="492">Pre-dorsal-fin length</th>
<td id="76E87020165C000E4062FE2E6D25FDD9" box="[526,620,505,529]" gridcol="1" gridrow="6" pageId="5" pageNumber="492">41.1</td>
<td id="76E87020165C000E40C8FE2E6C3BFDD9" box="[676,882,505,529]" gridcol="2" gridrow="6" pageId="5" pageNumber="492">4148(34)</td>
<td id="76E87020165C000E41C1FE2E6B5BFDD9" box="[941,1042,505,529]" gridcol="3" gridrow="6" pageId="5" pageNumber="492">43.2±0.3</td>
<td id="76E87020165C000E4604FE2E6B8FFDD9" box="[1128,1222,505,529]" gridcol="4" gridrow="6" pageId="5" pageNumber="492">41.5</td>
<td id="76E87020165C000E4779FE2E6A32FDD9" box="[1301,1403,505,529]" gridcol="5" gridrow="6" pageId="5" pageNumber="492">39.844.1</td>
</tr>
<tr id="3539195C165C000E42CFFDF66A32FDF1" box="[163,1403,545,569]" gridrow="7" pageId="5" pageNumber="492">
<th id="76E87020165C000E42CFFDF66EE2FDF1" box="[163,427,545,569]" gridcol="0" gridrow="7" pageId="5" pageNumber="492">Pre-anal-fin length</th>
<td id="76E87020165C000E4062FDF66D25FDF1" box="[526,620,545,569]" gridcol="1" gridrow="7" pageId="5" pageNumber="492">76.6</td>
<td id="76E87020165C000E40C8FDF66C3BFDF1" box="[676,882,545,569]" gridcol="2" gridrow="7" pageId="5" pageNumber="492">72.279(34)</td>
<td id="76E87020165C000E41C1FDF66B5BFDF1" box="[941,1042,545,569]" gridcol="3" gridrow="7" pageId="5" pageNumber="492">75.9±0.3</td>
<td id="76E87020165C000E4604FDF66B8FFDF1" box="[1128,1222,545,569]" gridcol="4" gridrow="7" pageId="5" pageNumber="492">73.5</td>
<td id="76E87020165C000E4779FDF66A32FDF1" box="[1301,1403,545,569]" gridcol="5" gridrow="7" pageId="5" pageNumber="492">70.476.9</td>
</tr>
<tr id="3539195C165C000E42CFFD9E6A32FDA9" box="[163,1403,585,609]" gridrow="8" pageId="5" pageNumber="492" rowspan-3="1">
<th id="76E87020165C000E42CFFD9E6EE2FDA9" box="[163,427,585,609]" gridcol="0" gridrow="8" pageId="5" pageNumber="492">Pre-pectoral-fin length</th>
<td id="76E87020165C000E4062FD9E6D25FDA9" box="[526,620,585,609]" gridcol="1" gridrow="8" pageId="5" pageNumber="492">31.8</td>
<td id="76E87020165C000E40C8FD9E6C3BFDA9" box="[676,882,585,609]" gridcol="2" gridrow="8" pageId="5" pageNumber="492">29.931.1(3)</td>
<td id="76E87020165C000E4604FD9E6B8FFDA9" box="[1128,1222,585,609]" gridcol="4" gridrow="8" pageId="5" pageNumber="492">31.6</td>
<td id="76E87020165C000E4779FD9E6A32FDA9" box="[1301,1403,585,609]" gridcol="5" gridrow="8" pageId="5" pageNumber="492">29.835</td>
</tr>
<tr id="3539195C165C000E42CFFDA66A32FD42" box="[163,1403,625,650]" gridrow="9" pageId="5" pageNumber="492">
<th id="76E87020165C000E42CFFDA66EE2FD42" box="[163,427,625,650]" gridcol="0" gridrow="9" pageId="5" pageNumber="492">Pre-pelvic-fin length</th>
<td id="76E87020165C000E4062FDA66D25FD42" box="[526,620,625,650]" gridcol="1" gridrow="9" pageId="5" pageNumber="492">44.3</td>
<td id="76E87020165C000E40C8FDA66C3BFD42" box="[676,882,625,650]" gridcol="2" gridrow="9" pageId="5" pageNumber="492">40.447.5(34)</td>
<td id="76E87020165C000E41C1FDA66B5BFD42" box="[941,1042,625,650]" gridcol="3" gridrow="9" pageId="5" pageNumber="492">45±0.3</td>
<td id="76E87020165C000E4604FDA66B8FFD42" box="[1128,1222,625,650]" gridcol="4" gridrow="9" pageId="5" pageNumber="492">42</td>
<td id="76E87020165C000E4779FDA66A32FD42" box="[1301,1403,625,650]" gridcol="5" gridrow="9" pageId="5" pageNumber="492">41.547.8</td>
</tr>
<tr id="3539195C165C000E42CFFD4D6A32FD7A" box="[163,1403,666,690]" gridrow="10" pageId="5" pageNumber="492">
<th id="76E87020165C000E42CFFD4D6EE2FD7A" box="[163,427,666,690]" gridcol="0" gridrow="10" pageId="5" pageNumber="492">Pre-adipose length</th>
<td id="76E87020165C000E4062FD4D6D25FD7A" box="[526,620,666,690]" gridcol="1" gridrow="10" pageId="5" pageNumber="492">79.7</td>
<td id="76E87020165C000E40C8FD4D6C3BFD7A" box="[676,882,666,690]" gridcol="2" gridrow="10" pageId="5" pageNumber="492">78.384.8(34)</td>
<td id="76E87020165C000E41C1FD4D6B5BFD7A" box="[941,1042,666,690]" gridcol="3" gridrow="10" pageId="5" pageNumber="492">81.1±0.2</td>
<td id="76E87020165C000E4604FD4D6B8FFD7A" box="[1128,1222,666,690]" gridcol="4" gridrow="10" pageId="5" pageNumber="492">80</td>
<td id="76E87020165C000E4779FD4D6A32FD7A" box="[1301,1403,666,690]" gridcol="5" gridrow="10" pageId="5" pageNumber="492">76.982.6</td>
</tr>
<tr id="3539195C165C000E42CFFD156A32FD12" box="[163,1403,706,730]" gridrow="11" pageId="5" pageNumber="492">
<th id="76E87020165C000E42CFFD156EE2FD12" box="[163,427,706,730]" gridcol="0" gridrow="11" pageId="5" pageNumber="492">Body depth (maximum)</th>
<td id="76E87020165C000E4062FD156D25FD12" box="[526,620,706,730]" gridcol="1" gridrow="11" pageId="5" pageNumber="492">22.9</td>
<td id="76E87020165C000E40C8FD156C3BFD12" box="[676,882,706,730]" gridcol="2" gridrow="11" pageId="5" pageNumber="492">18.928.1(34)</td>
<td id="76E87020165C000E41C1FD156B5BFD12" box="[941,1042,706,730]" gridcol="3" gridrow="11" pageId="5" pageNumber="492">24.3±0.4</td>
<td id="76E87020165C000E4604FD156B8FFD12" box="[1128,1222,706,730]" gridcol="4" gridrow="11" pageId="5" pageNumber="492">22.3</td>
<td id="76E87020165C000E4779FD156A32FD12" box="[1301,1403,706,730]" gridcol="5" gridrow="11" pageId="5" pageNumber="492">21.727.9</td>
</tr>
<tr id="3539195C165C000E42CFFD3D6A32FCCA" box="[163,1403,746,770]" gridrow="12" pageId="5" pageNumber="492" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="76E87020165C000E42CFFD3D6EE2FCCA" box="[163,427,746,770]" gridcol="0" gridrow="12" pageId="5" pageNumber="492">Interdorsal-adipose length</th>
<td id="76E87020165C000E4062FD3D6D25FCCA" box="[526,620,746,770]" gridcol="1" gridrow="12" pageId="5" pageNumber="492">20.3</td>
</tr>
<tr id="3539195C165C000E42CFFCC46A32FCE3" box="[163,1403,787,811]" gridrow="13" pageId="5" pageNumber="492">
<th id="76E87020165C000E42CFFCC46A32FCE3" box="[163,1403,787,811]" colspan="6" colspanRight="5" gridcol="0" gridrow="13" pageId="5" pageNumber="492">
<emphasis id="B97DC70C165CFFF442CFFCC46FB7FCE2" box="[163,254,787,811]" italics="true" pageId="5" pageNumber="492">As % HL</emphasis>
</th>
</tr>
<tr id="3539195C165C000E42CFFCEC6A32FC9B" box="[163,1403,827,851]" gridrow="14" pageId="5" pageNumber="492" rowspan-4="1">
<th id="76E87020165C000E42CFFCEC6EE2FC9B" box="[163,427,827,851]" gridcol="0" gridrow="14" pageId="5" pageNumber="492">Pre-orbital length</th>
<td id="76E87020165C000E4062FCEC6D25FC9B" box="[526,620,827,851]" gridcol="1" gridrow="14" pageId="5" pageNumber="492">24.6</td>
<td id="76E87020165C000E40C8FCEC6C3BFC9B" box="[676,882,827,851]" gridcol="2" gridrow="14" pageId="5" pageNumber="492">18.928.2(34)</td>
<td id="76E87020165C000E41C1FCEC6B5BFC9B" box="[941,1042,827,851]" gridcol="3" gridrow="14" pageId="5" pageNumber="492">23.9±0.4</td>
<td id="76E87020165C000E4779FCEC6A32FC9B" box="[1301,1403,827,851]" gridcol="5" gridrow="14" pageId="5" pageNumber="492">2226.3</td>
</tr>
<tr id="3539195C165C000E42CFFCB46A32FCB3" box="[163,1403,867,891]" gridrow="15" pageId="5" pageNumber="492" rowspan-4="1" rowspan-5="1">
<th id="76E87020165C000E42CFFCB46EE2FCB3" box="[163,427,867,891]" gridcol="0" gridrow="15" pageId="5" pageNumber="492">Post-orbital length</th>
<td id="76E87020165C000E4062FCB46D25FCB3" box="[526,620,867,891]" gridcol="1" gridrow="15" pageId="5" pageNumber="492">52.6</td>
<td id="76E87020165C000E40C8FCB46C3BFCB3" box="[676,882,867,891]" gridcol="2" gridrow="15" pageId="5" pageNumber="492">46.856.7(34)</td>
<td id="76E87020165C000E41C1FCB46B5BFCB3" box="[941,1042,867,891]" gridcol="3" gridrow="15" pageId="5" pageNumber="492">51.5±0.4</td>
</tr>
<tr id="3539195C165C000E42CFFC5C6A32FC6C" box="[163,1403,907,932]" gridrow="16" pageId="5" pageNumber="492">
<th id="76E87020165C000E42CFFC5C6EE2FC6C" box="[163,427,907,932]" gridcol="0" gridrow="16" pageId="5" pageNumber="492">Eye diameter</th>
<td id="76E87020165C000E4062FC5C6D25FC6C" box="[526,620,907,932]" gridcol="1" gridrow="16" pageId="5" pageNumber="492">21.1</td>
<td id="76E87020165C000E40C8FC5C6C3BFC6C" box="[676,882,907,932]" gridcol="2" gridrow="16" pageId="5" pageNumber="492">20.328.8(34)</td>
<td id="76E87020165C000E41C1FC5C6B5BFC6C" box="[941,1042,907,932]" gridcol="3" gridrow="16" pageId="5" pageNumber="492">24.7±0.4</td>
<td id="76E87020165C000E4604FC5C6B8FFC6C" box="[1128,1222,907,932]" gridcol="4" gridrow="16" pageId="5" pageNumber="492">24</td>
<td id="76E87020165C000E4779FC5C6A32FC6C" box="[1301,1403,907,932]" gridcol="5" gridrow="16" pageId="5" pageNumber="492">21.227.3</td>
</tr>
<tr id="3539195C165C000E42CFFC646A32FC04" box="[163,1403,947,972]" gridrow="17" pageId="5" pageNumber="492">
<th id="76E87020165C000E42CFFC646EE2FC04" box="[163,427,947,972]" gridcol="0" gridrow="17" pageId="5" pageNumber="492">Interorbital width</th>
<td id="76E87020165C000E4062FC646D25FC04" box="[526,620,947,972]" gridcol="1" gridrow="17" pageId="5" pageNumber="492">29.8</td>
<td id="76E87020165C000E40C8FC646C3BFC04" box="[676,882,947,972]" gridcol="2" gridrow="17" pageId="5" pageNumber="492">20.734(34)</td>
<td id="76E87020165C000E41C1FC646B5BFC04" box="[941,1042,947,972]" gridcol="3" gridrow="17" pageId="5" pageNumber="492">24.3±0.4</td>
<td id="76E87020165C000E4604FC646B8FFC04" box="[1128,1222,947,972]" gridcol="4" gridrow="17" pageId="5" pageNumber="492">30.3</td>
<td id="76E87020165C000E4779FC646A32FC04" box="[1301,1403,947,972]" gridcol="5" gridrow="17" pageId="5" pageNumber="492">25.833</td>
</tr>
<tr id="3539195C165C000E42CFFC0B6A32FC3C" box="[163,1403,988,1012]" gridrow="18" pageId="5" pageNumber="492">
<th id="76E87020165C000E42CFFC0B6EE2FC3C" box="[163,427,988,1012]" gridcol="0" gridrow="18" pageId="5" pageNumber="492">Upper jaw length</th>
<td id="76E87020165C000E4062FC0B6D25FC3C" box="[526,620,988,1012]" gridcol="1" gridrow="18" pageId="5" pageNumber="492">54.4</td>
<td id="76E87020165C000E40C8FC0B6C3BFC3C" box="[676,882,988,1012]" gridcol="2" gridrow="18" pageId="5" pageNumber="492">45.760(34)</td>
<td id="76E87020165C000E41C1FC0B6B5BFC3C" box="[941,1042,988,1012]" gridcol="3" gridrow="18" pageId="5" pageNumber="492">52.7±0.5</td>
<td id="76E87020165C000E4604FC0B6B8FFC3C" box="[1128,1222,988,1012]" gridcol="4" gridrow="18" pageId="5" pageNumber="492">51.8</td>
<td id="76E87020165C000E4779FC0B6A32FC3C" box="[1301,1403,988,1012]" gridcol="5" gridrow="18" pageId="5" pageNumber="492">5156.8</td>
</tr>
<tr id="3539195C165C000E42CFFBD36A32FBD4" box="[163,1403,1028,1052]" gridrow="19" pageId="5" pageNumber="492">
<th id="76E87020165C000E42CFFBD36EE2FBD4" box="[163,427,1028,1052]" gridcol="0" gridrow="19" pageId="5" pageNumber="492">Dorsal-fin base</th>
<td id="76E87020165C000E4062FBD36D25FBD4" box="[526,620,1028,1052]" gridcol="1" gridrow="19" pageId="5" pageNumber="492">59.6</td>
<td id="76E87020165C000E40C8FBD36C3BFBD4" box="[676,882,1028,1052]" gridcol="2" gridrow="19" pageId="5" pageNumber="492">44.259.6(34)</td>
<td id="76E87020165C000E41C1FBD36B5BFBD4" box="[941,1042,1028,1052]" gridcol="3" gridrow="19" pageId="5" pageNumber="492">51.6± 0.6</td>
<td id="76E87020165C000E4604FBD36B8FFBD4" box="[1128,1222,1028,1052]" gridcol="4" gridrow="19" pageId="5" pageNumber="492">51.8</td>
<td id="76E87020165C000E4779FBD36A32FBD4" box="[1301,1403,1028,1052]" gridcol="5" gridrow="19" pageId="5" pageNumber="492">45.260.3</td>
</tr>
<tr id="3539195C165C000E42CFFBFB6A32FB8C" box="[163,1403,1068,1092]" gridrow="20" pageId="5" pageNumber="492">
<th id="76E87020165C000E42CFFBFB6EE2FB8C" box="[163,427,1068,1092]" gridcol="0" gridrow="20" pageId="5" pageNumber="492">Anal-fin base</th>
<td id="76E87020165C000E4062FBFB6D25FB8C" box="[526,620,1068,1092]" gridcol="1" gridrow="20" pageId="5" pageNumber="492">31.6</td>
<td id="76E87020165C000E40C8FBFB6C3BFB8C" box="[676,882,1068,1092]" gridcol="2" gridrow="20" pageId="5" pageNumber="492">25.734(34)</td>
<td id="76E87020165C000E41C1FBFB6B5BFB8C" box="[941,1042,1068,1092]" gridcol="3" gridrow="20" pageId="5" pageNumber="492">31±0.4</td>
<td id="76E87020165C000E4604FBFB6B8FFB8C" box="[1128,1222,1068,1092]" gridcol="4" gridrow="20" pageId="5" pageNumber="492">40</td>
<td id="76E87020165C000E4779FBFB6A32FB8C" box="[1301,1403,1068,1092]" gridcol="5" gridrow="20" pageId="5" pageNumber="492">33.743.7</td>
</tr>
<tr id="3539195C165C000E42CFFB836A32FBA5" box="[163,1403,1108,1133]" gridrow="21" pageId="5" pageNumber="492">
<th id="76E87020165C000E42CFFB836EE2FBA5" box="[163,427,1108,1133]" gridcol="0" gridrow="21" pageId="5" pageNumber="492">Pectoral-fin length</th>
<td id="76E87020165C000E4062FB836D25FBA5" box="[526,620,1108,1133]" gridcol="1" gridrow="21" pageId="5" pageNumber="492">107</td>
<td id="76E87020165C000E40C8FB836C3BFBA5" box="[676,882,1108,1133]" gridcol="2" gridrow="21" pageId="5" pageNumber="492">84.4111.7(33)</td>
<td id="76E87020165C000E41C1FB836B5BFBA5" box="[941,1042,1108,1133]" gridcol="3" gridrow="21" pageId="5" pageNumber="492">99.9±1.2</td>
<td id="76E87020165C000E4604FB836B8FFBA5" box="[1128,1222,1108,1133]" gridcol="4" gridrow="21" pageId="5" pageNumber="492">87</td>
<td id="76E87020165C000E4779FB836A32FBA5" box="[1301,1403,1108,1133]" gridcol="5" gridrow="21" pageId="5" pageNumber="492">82100</td>
</tr>
<tr id="3539195C165C000E42CFFBAA6A32FB5D" box="[163,1403,1149,1173]" gridrow="22" pageId="5" pageNumber="492" rowspan-4="1">
<th id="76E87020165C000E42CFFBAA6EE2FB5D" box="[163,427,1149,1173]" gridcol="0" gridrow="22" pageId="5" pageNumber="492">Pelvic-fin length</th>
<td id="76E87020165C000E4062FBAA6D25FB5D" box="[526,620,1149,1173]" gridcol="1" gridrow="22" pageId="5" pageNumber="492">59.6</td>
<td id="76E87020165C000E40C8FBAA6C3BFB5D" box="[676,882,1149,1173]" gridcol="2" gridrow="22" pageId="5" pageNumber="492">40.459.6(34)</td>
<td id="76E87020165C000E41C1FBAA6B5BFB5D" box="[941,1042,1149,1173]" gridcol="3" gridrow="22" pageId="5" pageNumber="492">50.1±0.9</td>
<td id="76E87020165C000E4779FBAA6A32FB5D" box="[1301,1403,1149,1173]" gridcol="5" gridrow="22" pageId="5" pageNumber="492">61.365.8</td>
</tr>
<tr id="3539195C165C000E42CFFB726A32FB75" box="[163,1403,1189,1213]" gridrow="23" pageId="5" pageNumber="492" rowspan-3="1">
<th id="76E87020165C000E42CFFB726EE2FB75" box="[163,427,1189,1213]" gridcol="0" gridrow="23" pageId="5" pageNumber="492">Caudal-peduncle depth</th>
<td id="76E87020165C000E4062FB726D25FB75" box="[526,620,1189,1213]" gridcol="1" gridrow="23" pageId="5" pageNumber="492">33.6</td>
<td id="76E87020165C000E40C8FB726C3BFB75" box="[676,882,1189,1213]" gridcol="2" gridrow="23" pageId="5" pageNumber="492">33.635.1(3)</td>
<td id="76E87020165C000E4604FB726B8FFB75" box="[1128,1222,1189,1213]" gridcol="4" gridrow="23" pageId="5" pageNumber="492">36.4</td>
<td id="76E87020165C000E4779FB726A32FB75" box="[1301,1403,1189,1213]" gridcol="5" gridrow="23" pageId="5" pageNumber="492">31.141.8</td>
</tr>
<tr id="3539195C165C000E42CFFB196A32FB2D" box="[163,1403,1230,1253]" gridrow="24" pageId="5" pageNumber="492">
<th id="76E87020165C000E42CFFB196A32FB2D" box="[163,1403,1230,1253]" colspan="6" colspanRight="5" gridcol="0" gridrow="24" pageId="5" pageNumber="492">
<emphasis id="B97DC70C165CFFF442CFFB196FBEFB2D" box="[163,247,1230,1253]" italics="true" pageId="5" pageNumber="492">Meristic</emphasis>
</th>
</tr>
<tr id="3539195C165C000E42CFFB226A32FAC6" box="[163,1403,1269,1294]" gridrow="25" pageId="5" pageNumber="492" rowspan-3="1">
<th id="76E87020165C000E42CFFB226EE2FAC6" box="[163,427,1269,1294]" gridcol="0" gridrow="25" pageId="5" pageNumber="492">Dorsal fin rays</th>
<td id="76E87020165C000E4062FB226D25FAC6" box="[526,620,1269,1294]" gridcol="1" gridrow="25" pageId="5" pageNumber="492">14</td>
<td id="76E87020165C000E40C8FB226C3BFAC6" box="[676,882,1269,1294]" gridcol="2" gridrow="25" pageId="5" pageNumber="492">1314</td>
<td id="76E87020165C000E4604FB226B8FFAC6" box="[1128,1222,1269,1294]" gridcol="4" gridrow="25" pageId="5" pageNumber="492">13</td>
<td id="76E87020165C000E4779FB226A32FAC6" box="[1301,1403,1269,1294]" gridcol="5" gridrow="25" pageId="5" pageNumber="492">1213</td>
</tr>
<tr id="3539195C165C000E42CFFAC96A32FAFE" box="[163,1403,1310,1334]" gridrow="26" pageId="5" pageNumber="492" rowspan-3="1">
<th id="76E87020165C000E42CFFAC96EE2FAFE" box="[163,427,1310,1334]" gridcol="0" gridrow="26" pageId="5" pageNumber="492">Anal fin rays</th>
<td id="76E87020165C000E4062FAC96D25FAFE" box="[526,620,1310,1334]" gridcol="1" gridrow="26" pageId="5" pageNumber="492">11</td>
<td id="76E87020165C000E40C8FAC96C3BFAFE" box="[676,882,1310,1334]" gridcol="2" gridrow="26" pageId="5" pageNumber="492">1012</td>
<td id="76E87020165C000E4604FAC96B8FFAFE" box="[1128,1222,1310,1334]" gridcol="4" gridrow="26" pageId="5" pageNumber="492">12</td>
<td id="76E87020165C000E4779FAC96A32FAFE" box="[1301,1403,1310,1334]" gridcol="5" gridrow="26" pageId="5" pageNumber="492">1113</td>
</tr>
<tr id="3539195C165C000E42CFFA916A32FA96" box="[163,1403,1350,1374]" gridrow="27" pageId="5" pageNumber="492" rowspan-3="1">
<th id="76E87020165C000E42CFFA916EE2FA96" box="[163,427,1350,1374]" gridcol="0" gridrow="27" pageId="5" pageNumber="492">Pectoral fin rays</th>
<td id="76E87020165C000E4062FA916D25FA96" box="[526,620,1350,1374]" gridcol="1" gridrow="27" pageId="5" pageNumber="492">17</td>
<td id="76E87020165C000E40C8FA916C3BFA96" box="[676,882,1350,1374]" gridcol="2" gridrow="27" pageId="5" pageNumber="492">1618</td>
<td id="76E87020165C000E4604FA916B8FFA96" box="[1128,1222,1350,1374]" gridcol="4" gridrow="27" pageId="5" pageNumber="492">16</td>
<td id="76E87020165C000E4779FA916A32FA96" box="[1301,1403,1350,1374]" gridcol="5" gridrow="27" pageId="5" pageNumber="492">1516</td>
</tr>
<tr id="3539195C165C000E42CFFAB96A32FA4F" box="[163,1403,1390,1415]" gridrow="28" pageId="5" pageNumber="492" rowspan-3="1">
<th id="76E87020165C000E42CFFAB96EE2FA4F" box="[163,427,1390,1415]" gridcol="0" gridrow="28" pageId="5" pageNumber="492">Pelvic fin rays</th>
<td id="76E87020165C000E4062FAB96D25FA4F" box="[526,620,1390,1415]" gridcol="1" gridrow="28" pageId="5" pageNumber="492">9</td>
<td id="76E87020165C000E40C8FAB96C3BFA4F" box="[676,882,1390,1415]" gridcol="2" gridrow="28" pageId="5" pageNumber="492">9</td>
<td id="76E87020165C000E4604FAB96B8FFA4F" box="[1128,1222,1390,1415]" gridcol="4" gridrow="28" pageId="5" pageNumber="492">9</td>
<td id="76E87020165C000E4779FAB96A32FA4F" box="[1301,1403,1390,1415]" gridcol="5" gridrow="28" pageId="5" pageNumber="492">9</td>
</tr>
<tr id="3539195C165C000E42CFFA416A32FA67" box="[163,1403,1430,1455]" gridrow="29" pageId="5" pageNumber="492" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="76E87020165C000E42CFFA416EE2FA67" box="[163,427,1430,1455]" gridcol="0" gridrow="29" pageId="5" pageNumber="492">Branchiostegal rays</th>
<td id="76E87020165C000E4062FA416D25FA67" box="[526,620,1430,1455]" gridcol="1" gridrow="29" pageId="5" pageNumber="492">9</td>
<td id="76E87020165C000E40C8FA416C3BFA67" box="[676,882,1430,1455]" gridcol="2" gridrow="29" pageId="5" pageNumber="492">9</td>
</tr>
<tr id="3539195C165C000E42CFFA686A32FA1F" box="[163,1403,1471,1495]" gridrow="30" pageId="5" pageNumber="492" rowspan-3="1">
<th id="76E87020165C000E42CFFA686EE2FA1F" box="[163,427,1471,1495]" gridcol="0" gridrow="30" pageId="5" pageNumber="492">Gill rakers</th>
<td id="76E87020165C000E4062FA686D25FA1F" box="[526,620,1471,1495]" gridcol="1" gridrow="30" pageId="5" pageNumber="492">14</td>
<td id="76E87020165C000E40C8FA686C3BFA1F" box="[676,882,1471,1495]" gridcol="2" gridrow="30" pageId="5" pageNumber="492">1415</td>
<td id="76E87020165C000E4604FA686B8FFA1F" box="[1128,1222,1471,1495]" gridcol="4" gridrow="30" pageId="5" pageNumber="492">16</td>
<td id="76E87020165C000E4779FA686A32FA1F" box="[1301,1403,1471,1495]" gridcol="5" gridrow="30" pageId="5" pageNumber="492">1418</td>
</tr>
<tr id="3539195C165C000E42CFFA306A32FA37" box="[163,1403,1511,1535]" gridrow="31" pageId="5" pageNumber="492" rowspan-3="1">
<th id="76E87020165C000E42CFFA306EE2FA37" box="[163,427,1511,1535]" gridcol="0" gridrow="31" pageId="5" pageNumber="492">Is phtophores</th>
<td id="76E87020165C000E4062FA306D25FA37" box="[526,620,1511,1535]" gridcol="1" gridrow="31" pageId="5" pageNumber="492">9</td>
<td id="76E87020165C000E40C8FA306C3BFA37" box="[676,882,1511,1535]" gridcol="2" gridrow="31" pageId="5" pageNumber="492">89</td>
<td id="76E87020165C000E4604FA306B8FFA37" box="[1128,1222,1511,1535]" gridcol="4" gridrow="31" pageId="5" pageNumber="492">9</td>
<td id="76E87020165C000E4779FA306A32FA37" box="[1301,1403,1511,1535]" gridcol="5" gridrow="31" pageId="5" pageNumber="492">810</td>
</tr>
<tr id="3539195C165C000E42CFF9D86A32F9EF" box="[163,1403,1551,1575]" gridrow="32" pageId="5" pageNumber="492" rowspan-3="1">
<th id="76E87020165C000E42CFF9D86EE2F9EF" box="[163,427,1551,1575]" gridcol="0" gridrow="32" pageId="5" pageNumber="492">LO photophores</th>
<td id="76E87020165C000E4062F9D86D25F9EF" box="[526,620,1551,1575]" gridcol="1" gridrow="32" pageId="5" pageNumber="492">22</td>
<td id="76E87020165C000E40C8F9D86C3BF9EF" box="[676,882,1551,1575]" gridcol="2" gridrow="32" pageId="5" pageNumber="492">2124</td>
<td id="76E87020165C000E4604F9D86B8FF9EF" box="[1128,1222,1551,1575]" gridcol="4" gridrow="32" pageId="5" pageNumber="492">24</td>
<td id="76E87020165C000E4779F9D86A32F9EF" box="[1301,1403,1551,1575]" gridcol="5" gridrow="32" pageId="5" pageNumber="492">2026</td>
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</paragraph>
<paragraph id="8BB61B1E165CFFF442ABF9AC6A71F915" blockId="5.[151,1437,1659,2010]" pageId="5" pageNumber="492">
For the remaining
<taxonomicName id="4C09609D165CFFF443F4F9AB6D6FF95D" box="[408,550,1659,1685]" family="Neoscopelidae" genus="Neoscopelus" kingdom="Animalia" order="Myctophiformes" pageId="5" pageNumber="492" phylum="Chordata" rank="species" species="microchir">
<emphasis id="B97DC70C165CFFF443F4F9AB6D6FF95D" box="[408,550,1659,1685]" italics="true" pageId="5" pageNumber="492">N. microchir</emphasis>
</taxonomicName>
, the DNA analyses revealed three independent MOTUs consistently supported by all analyses; the first with eleven individuals from
<collectingCountry id="F31E5B8E165CFFF44086F9486C76F971" box="[746,831,1695,1721]" name="Taiwan" pageId="5" pageNumber="492">Taiwan</collectingCountry>
, the second with six sequences from
<collectingCountry id="F31E5B8E165CFFF446B4F9486A0BF971" box="[1240,1346,1695,1721]" name="Australia" pageId="5" pageNumber="492">Australia</collectingCountry>
, and the third with four individuals, three of them from the
<collectingCountry id="F31E5B8E165CFFF440A3F9146C5BF915" box="[719,786,1731,1757]" name="China" pageId="5" pageNumber="492">China</collectingCountry>
Sea and the last (AP002921) of unknown origin.
</paragraph>
<paragraph id="8BB61B1E165CFFF742ABF9306B23FF1D" blockId="5.[151,1437,1659,2010]" lastBlockId="6.[151,1436,151,213]" lastPageId="6" lastPageNumber="493" pageId="5" pageNumber="492">
Several MOTUs have also been found in
<taxonomicName id="4C09609D165CFFF440DFF93F6CCCF8C9" box="[691,901,1767,1793]" family="Neoscopelidae" genus="Neoscopelus" kingdom="Animalia" order="Myctophiformes" pageId="5" pageNumber="492" phylum="Chordata" rank="species" species="macrolepidotus">
<emphasis id="B97DC70C165CFFF440DFF93F6CCCF8C9" box="[691,901,1767,1793]" italics="true" pageId="5" pageNumber="492">N. macrolepidotus</emphasis>
</taxonomicName>
, ranging from three to five depending on the analysis. The sequences assigned to
<taxonomicName id="4C09609D165CFFF44059F8DB6C4DF8ED" box="[565,772,1803,1829]" family="Neoscopelidae" genus="Neoscopelus" kingdom="Animalia" order="Myctophiformes" pageId="5" pageNumber="492" phylum="Chordata" rank="species" species="macrolepidotus">
<emphasis id="B97DC70C165CFFF44059F8DB6C4DF8ED" box="[565,772,1803,1829]" italics="true" pageId="5" pageNumber="492">N. macrolepidotus</emphasis>
</taxonomicName>
are mostly subdivided into two large groups supported by all the analyses (
<figureCitation id="1332079B165CFFF4433EF8F86EF6F881" box="[338,447,1839,1865]" captionStart="FIGURE 2" captionStartId="8.[151,250,1679,1703]" captionTargetBox="[252,1333,190,1647]" captionTargetId="figure-19@8.[248,1340,181,1655]" captionTargetPageId="8" captionText="FIGURE 2. Consensus phylogeny (Maximum Likelihood and Bayesian Inference) of Neoscopelus based on COI gene sequences with the results of six different molecular approaches: BIN, Barcode Index Number; ASAP, Assemble Species by Automatic Partitioning; bPTP, Bayesian Poisson Tree Process; mPTP, multiple Poisson Tree Process; GMYC, General Mixed Yule-Coalescent model and mGMYC, multiple General Mixed Yule-Coalescent model. The proposed Molecular Operational Taxonomic Units (MOTUs) for each analysis are represented as boxes. The colours in the branches indicate the proposed MOTUs in this study. In the first node of each MOTU, both boostrap values (&gt;75) and posterior probabilities (&gt;0.9) are shown. Each individual contains the putative morphospecies, repository and geographic codes: NEA—Northeast Atlantic; JAM— Jamaica; ESA—Eastern South Africa; TWN—Taiwan; WAU—Western Australia; SCS—South China Sea; IO—Indian Ocean; PNG—Papua New Guinea; ESA—Eastern South Africa; TAS—Tasmania; GME—Gulf of Mexico; NZL—New Zealand; NWA—Northwest Atlantic." figureDoi="http://doi.org/10.5281/zenodo.14022811" httpUri="https://zenodo.org/record/14022811/files/figure.png" pageId="5" pageNumber="492">Figures 2</figureCitation>
,
<figureCitation id="1332079B165CFFF443A7F8F86E95F881" box="[459,476,1839,1865]" captionStart="FIGURE 4" captionStartId="9.[151,250,1661,1685]" captionTargetBox="[286,1300,1002,1632]" captionTargetId="figure-64@9.[282,1305,999,1637]" captionTargetPageId="9" captionText="FIGURE 4. Map of sampling locations of the Neoscopelus macrolepidotus species complex and the single specimen of Neoscopelus porosus (in purple). The diameter of the circle is indicative of the sample size, and the colours correspond to the MOTUs detected in the phylogenetic analysis." figureDoi="http://doi.org/10.5281/zenodo.14022815" httpUri="https://zenodo.org/record/14022815/files/figure.png" pageId="5" pageNumber="492">4</figureCitation>
). The first comprises eight individuals of the Indo-Pacific area and one from the Gulf of Mexico (MG856810). The second group includes between 10 and 12 individuals distributed worldwide. Both groups represent 85% of the
<taxonomicName id="4C09609D165CFFF443B3F8AF6DE7F859" box="[479,686,1911,1937]" family="Neoscopelidae" genus="Neoscopelus" kingdom="Animalia" order="Myctophiformes" pageId="5" pageNumber="492" phylum="Chordata" rank="species" species="macrolepidotus">
<emphasis id="B97DC70C165CFFF443B3F8AF6DE7F859" box="[479,686,1911,1937]" italics="true" pageId="5" pageNumber="492">N. macrolepidotus</emphasis>
</taxonomicName>
individuals used in the present study. Similarly, a single sequence from the Indian Ocean (FOAQ886-22) is always clustered individually. On the other hand, there are two sequences (FMVIC257-08 from
<collectingCountry id="F31E5B8E165CFFF443E3F8686EBFF811" box="[399,502,1983,2009]" name="Australia" pageId="5" pageNumber="492">Australia</collectingCountry>
and AP012238) that are included in one of the majority MOTU in three analyses (ASAP, mPTP and mGMYC) and are considered as two independent MOTUs for bPTP. BIN and GMYC included AP012238 in a shared cluster but considered FMVIC257-08 as an independent species.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,54 +1,56 @@
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<collectorName id="38465302BB21FFB4FE1EE234FDA3F96B" box="[456,597,1886,1913]" pageId="6" pageNumber="442">Wm. Clarke</collectorName> <collectorName id="38465302BB21FFB4FE1EE234FDA3F96B" box="[456,597,1886,1913]" pageId="6" pageNumber="442">Wm. Clarke</collectorName>
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; ;
<materialsCitation id="25DB3C89BB21FFB4FDB7E234FC96F96B" box="[609,864,1886,1913]" collectionCode="NMNH" country="Ecuador" pageId="6" pageNumber="442" specimenCode="NMNH 31472" specimenCount="1" specimenCount-male="1"> <materialsCitation id="25DB3C89BB21FFB4FDB7E234FB27F96B" box="[609,1233,1886,1913]" collectingDate="1939-04-29" collectionCode="NMNH" collectorName="Wm. Clarke" country="Ecuador" location="Banos" pageId="6" pageNumber="442" specimenCode="NMNH 31472" specimenCount="1" specimenCount-male="1">
<specimenCode id="C5159EAFBB21FFB4FDB7E234FCF1F96B" box="[609,775,1886,1913]" collectionCode="NMNH" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="442" type="Museum">NMNH 31472</specimenCode> <specimenCode id="C5159EAFBB21FFB4FDB7E234FCF1F96B" box="[609,775,1886,1913]" collectionCode="NMNH" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="442" type="Museum">NMNH 31472</specimenCode>
, ,
<specimenCount id="83B5FD5DBB21FFB4FCC4E235FC96F96B" box="[786,864,1886,1912]" count="1" pageId="6" pageNumber="442" type="male">1 male</specimenCount> <specimenCount id="83B5FD5DBB21FFB4FCC4E235FC96F96B" box="[786,864,1886,1912]" count="1" pageId="6" pageNumber="442" type="male">1 male</specimenCount>
</materialsCitation>
, ,
<materialsCitation id="25DB3C89BB21FFB4FCBAE234FB27F96B" box="[876,1233,1886,1913]" collectingDate="1939-04-29" collectorName="Wm. Clarke" country="Ecuador" location="Banos" pageId="6" pageNumber="442" specimenCount="1">
<location id="906C600FBB21FFB4FCBAE234FC43F96A" LSID="urn:lsid:plazi:treatment:1D1A87C2BB21FFBAFF41E3A5FD71FC97:906C600FBB21FFB4FCBAE234FC43F96A" box="[876,949,1887,1913]" country="Ecuador" name="Banos" pageId="6" pageNumber="442">Baños</location> <location id="906C600FBB21FFB4FCBAE234FC43F96A" LSID="urn:lsid:plazi:treatment:1D1A87C2BB21FFBAFF41E3A5FD71FC97:906C600FBB21FFB4FCBAE234FC43F96A" box="[876,949,1887,1913]" country="Ecuador" name="Banos" pageId="6" pageNumber="442">Baños</location>
, ,
<date id="E10D1014BB21FFB4FC17E235FBCFF96B" box="[961,1081,1886,1912]" pageId="6" pageNumber="442" value="1939-04-29"> <date id="E10D1014BB21FFB4FC17E235FBCFF96B" box="[961,1081,1886,1912]" pageId="6" pageNumber="442" value="1939-04-29">
@ -130,13 +130,11 @@ and
<collectorName id="38465302BB21FFB4FB92E234FB27F96B" box="[1092,1233,1886,1913]" pageId="6" pageNumber="442">Wm. Clarke</collectorName> <collectorName id="38465302BB21FFB4FB92E234FB27F96B" box="[1092,1233,1886,1913]" pageId="6" pageNumber="442">Wm. Clarke</collectorName>
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; ;
<materialsCitation id="25DB3C89BB21FFB4FB0BE234FEF2F98F" collectionCode="NMNH" country="Ecuador" pageId="6" pageNumber="442" specimenCode="NMNH 31472" specimenCount="1"> <materialsCitation id="25DB3C89BB21FFB4FB0BE234FD82F98F" collectingDate="1938-10-19" collectionCode="NMNH" collectorName="Wm. Clarke" country="Ecuador" location="Banos" pageId="6" pageNumber="442" specimenCode="NMNH 31472" specimenCount="1">
<specimenCode id="C5159EAFBB21FFB4FB0BE234FA75F96B" box="[1245,1411,1886,1913]" collectionCode="NMNH" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="442" type="Museum">NMNH 31472</specimenCode> <specimenCode id="C5159EAFBB21FFB4FB0BE234FA75F96B" box="[1245,1411,1886,1913]" collectionCode="NMNH" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="442" type="Museum">NMNH 31472</specimenCode>
, ,
<specimenCount id="83B5FD5DBB21FFB4FA58E235FEF2F98F" count="1" pageId="6" pageNumber="442" type="generic">1 specimen</specimenCount> <specimenCount id="83B5FD5DBB21FFB4FA58E235FEF2F98F" count="1" pageId="6" pageNumber="442" type="generic">1 specimen</specimenCount>
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<materialsCitation id="25DB3C89BB21FFB4FEC7E2E8FD82F98F" box="[273,628,1922,1949]" collectingDate="1938-10-19" collectorName="Wm. Clarke" country="Ecuador" location="Banos" pageId="6" pageNumber="442" specimenCount="1">
<location id="906C600FBB21FFB4FEC7E2E8FEACF98E" LSID="urn:lsid:plazi:treatment:1D1A87C2BB21FFBAFF41E3A5FD71FC97:906C600FBB21FFB4FEC7E2E8FEACF98E" box="[273,346,1923,1949]" country="Ecuador" name="Banos" pageId="6" pageNumber="442">Baños</location> <location id="906C600FBB21FFB4FEC7E2E8FEACF98E" LSID="urn:lsid:plazi:treatment:1D1A87C2BB21FFBAFF41E3A5FD71FC97:906C600FBB21FFB4FEC7E2E8FEACF98E" box="[273,346,1923,1949]" country="Ecuador" name="Banos" pageId="6" pageNumber="442">Baños</location>
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<date id="E10D1014BB21FFB4FEB1E2E9FE2FF98F" box="[359,473,1922,1948]" pageId="6" pageNumber="442" value="1938-10-19"> <date id="E10D1014BB21FFB4FEB1E2E9FE2FF98F" box="[359,473,1922,1948]" pageId="6" pageNumber="442" value="1938-10-19">
@ -146,13 +144,11 @@ and
<collectorName id="38465302BB21FFB4FE30E2E8FD82F98F" box="[486,628,1922,1949]" pageId="6" pageNumber="442">Wm. Clarke</collectorName> <collectorName id="38465302BB21FFB4FE30E2E8FD82F98F" box="[486,628,1922,1949]" pageId="6" pageNumber="442">Wm. Clarke</collectorName>
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<materialsCitation id="25DB3C89BB21FFB4FD54E2E8FC51F98F" box="[642,935,1922,1949]" collectionCode="NMNH" country="Ecuador" pageId="6" pageNumber="442" specimenCode="NMNH 31472" specimenCount="4" specimenCount-female="4"> <materialsCitation id="25DB3C89BB21FFB4FD54E2E8FED5F9D3" collectingDate="1937-09-15" collectionCode="NMNH" collectorName="Wm. Clarke" country="Ecuador" location="Rio Blanco" municipality="Prov." pageId="6" pageNumber="442" specimenCode="NMNH 31472" specimenCount="4" specimenCount-female="4" stateProvince="Tungurahua">
<specimenCode id="C5159EAFBB21FFB4FD54E2E8FCDFF98F" box="[642,809,1922,1949]" collectionCode="NMNH" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="442" type="Museum">NMNH 31472</specimenCode> <specimenCode id="C5159EAFBB21FFB4FD54E2E8FCDFF98F" box="[642,809,1922,1949]" collectionCode="NMNH" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="442" type="Museum">NMNH 31472</specimenCode>
, ,
<specimenCount id="83B5FD5DBB21FFB4FCE0E2E9FC51F98F" box="[822,935,1922,1948]" count="4" pageId="6" pageNumber="442" type="female">4 females</specimenCount> <specimenCount id="83B5FD5DBB21FFB4FCE0E2E9FC51F98F" box="[822,935,1922,1948]" count="4" pageId="6" pageNumber="442" type="female">4 females</specimenCount>
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<materialsCitation id="25DB3C89BB21FFB4FC62E2E8FED5F9D3" collectingDate="1937-09-15" collectorName="Wm. Clarke" country="Ecuador" location="Rio Blanco" municipality="Prov." pageId="6" pageNumber="442" specimenCount="1" stateProvince="Tungurahua">
<collectingMunicipality id="7568ACAEBB21FFB4FC62E2E8FC19F98E" box="[948,1007,1923,1949]" pageId="6" pageNumber="442">Prov.</collectingMunicipality> <collectingMunicipality id="7568ACAEBB21FFB4FC62E2E8FC19F98E" box="[948,1007,1923,1949]" pageId="6" pageNumber="442">Prov.</collectingMunicipality>
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@ -165,13 +161,11 @@ and
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<specimenCode id="C5159EAFBB21FFB4FEF9E2CCFE22F9D2" box="[303,468,1958,1985]" collectionCode="NMNH" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="442" type="Museum">NMNH 03285</specimenCode> <specimenCode id="C5159EAFBB21FFB4FEF9E2CCFE22F9D2" box="[303,468,1958,1985]" collectionCode="NMNH" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="442" type="Museum">NMNH 03285</specimenCode>
, ,
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<materialsCitation id="25DB3C89BB21FFB4FD99E2CCFB8CF9D3" box="[591,1146,1958,1985]" collectingDate="1976-01-24" collectorName="P. J. Sparrylr" country="Ecuador" location="Banos" municipality="Prov." pageId="6" pageNumber="442" specimenCount="1" stateProvince="Tungurahua">
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View file

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male (here designated) and male (here designated)
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@ -72,17 +74,18 @@
<subSubSection id="DDA9655FBB2FFFBAFF41E628FD20FD92" pageId="8" pageNumber="444" type="materials_examined"> <subSubSection id="DDA9655FBB2FFFBAFF41E628FD20FD92" pageId="8" pageNumber="444" type="materials_examined">
<paragraph id="950C36D4BB2FFFBAFF41E628FD20FD92" blockId="8.[151,1437,834,1005]" pageId="8" pageNumber="444"> <paragraph id="950C36D4BB2FFFBAFF41E628FD20FD92" blockId="8.[151,1437,834,1005]" pageId="8" pageNumber="444">
<emphasis id="A7C7EAC6BB2FFFBAFF41E628FE3DFD4F" bold="true" box="[151,459,834,861]" pageId="8" pageNumber="444">Type material examined:</emphasis> <emphasis id="A7C7EAC6BB2FFFBAFF41E628FE3DFD4F" bold="true" box="[151,459,834,861]" pageId="8" pageNumber="444">Type material examined:</emphasis>
<materialsCitation id="25DB3C89BB2FFFBAFE01E628FD20FD92" collectingDate="1955-02-13" collectionCode="CAS" collectorName="Banos, N. &amp; E. I Schlinger &amp; E. S. Ross." country="Ecuador" location="Tungurahua" pageId="8" pageNumber="444" specimenCode="CAS 15215" specimenCount="1" stateProvince="Tungurahua" typeStatus="holotype"> <materialsCitation id="25DB3C89BB2FFFBAFE01E628FD20FD92" collectingDate="1955-02-13" collectionCode="CAS" collectorName="E. I Schlinger &amp; E. S. Ross." country="Ecuador" location="Tungurahua" pageId="8" pageNumber="444" specimenCode="CAS 15215" specimenCount="1" stateProvince="Tungurahua" typeStatus="holotype">
<typeStatus id="4A088876BB2FFFBAFE01E628FDC9FD4E" box="[471,575,835,861]" pageId="8" pageNumber="444" type="holotype">Holotype</typeStatus> <typeStatus id="4A088876BB2FFFBAFE01E628FDC9FD4E" box="[471,575,835,861]" pageId="8" pageNumber="444" type="holotype">Holotype</typeStatus>
male <specimenCount id="83B5FD5DBB2FFFBAFD9DE628FD77FD4E" box="[587,641,835,861]" pageId="8" pageNumber="444" type="male">male</specimenCount>
<specimenCode id="C5159EAFBB2FFFBAFD5BE628FCE3FD4E" box="[653,789,834,861]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="8" pageNumber="444" type="Museum">CAS 15215</specimenCode> <specimenCode id="C5159EAFBB2FFFBAFD5BE628FCE3FD4E" box="[653,789,834,861]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="8" pageNumber="444" type="Museum">CAS 15215</specimenCode>
(12-169), (12-169),
<collectingCountry id="EDA47644BB2FFFBAFC43E628FC02FD4E" box="[917,1012,835,861]" name="Ecuador" pageId="8" pageNumber="444">Ecuador</collectingCountry> <collectingCountry id="EDA47644BB2FFFBAFC43E628FC02FD4E" box="[917,1012,835,861]" name="Ecuador" pageId="8" pageNumber="444">Ecuador</collectingCountry>
, ,
<locationDeviation id="84BE2FF6BB2FFFBAFBD2E628FA6FFD4E" box="[1028,1433,835,861]" pageId="8" pageNumber="444">
<quantity id="524B9B31BB2FFFBAFBD2E628FB9EFD4E" box="[1028,1128,835,861]" metricMagnitude="4" metricUnit="m" metricValue="1.4484096000000002" metricValueMax="1.9312128000000002" metricValueMin="0.9656064000000001" pageId="8" pageNumber="444" unit="mi" value="9.0" valueMax="12.0" valueMin="6.0">6-12 mi.</quantity> <quantity id="524B9B31BB2FFFBAFBD2E628FB9EFD4E" box="[1028,1128,835,861]" metricMagnitude="4" metricUnit="m" metricValue="1.4484096000000002" metricValueMax="1.9312128000000002" metricValueMin="0.9656064000000001" pageId="8" pageNumber="444" unit="mi" value="9.0" valueMax="12.0" valueMin="6.0">6-12 mi.</quantity>
SW. SW. Baños, N. slope Mt
<collectorName id="38465302BB2FFFBAFB78E628FAD4FD4E" box="[1198,1314,835,861]" pageId="8" pageNumber="444">Baños, N.</collectorName> </locationDeviation>
slope Mt. .
<collectingRegion id="5777F836BB2FFFBAFF41E60CFED6FD92" box="[151,288,871,897]" country="Ecuador" name="Tungurahua" pageId="8" pageNumber="444">Tungurahua</collectingRegion> <collectingRegion id="5777F836BB2FFFBAFF41E60CFED6FD92" box="[151,288,871,897]" country="Ecuador" name="Tungurahua" pageId="8" pageNumber="444">Tungurahua</collectingRegion>
, ,
<date id="E10D1014BB2FFFBAFEFDE60DFE56FD92" box="[299,416,870,897]" pageId="8" pageNumber="444" value="1955-02-13"> <date id="E10D1014BB2FFFBAFEFDE60DFE56FD92" box="[299,416,870,897]" pageId="8" pageNumber="444" value="1955-02-13">
@ -96,7 +99,7 @@ slope Mt.
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection id="DDA9655FBB2FFFB8FF11E6E1FAC6FFE6" lastPageId="10" lastPageNumber="446" pageId="8" pageNumber="444" type="description"> <subSubSection id="DDA9655FBB2FFFB8FF11E6E1FAC6FFE6" lastPageId="10" lastPageNumber="446" pageId="8" pageNumber="444" type="description">
<paragraph id="950C36D4BB2FFFBAFF11E6E1FA6AFDFE" blockId="8.[151,1437,834,1005]" pageId="8" pageNumber="444"> <paragraph id="950C36D4BB2FFFB8FF11E6E1FAC6FFE6" blockId="8.[151,1437,834,1005]" lastBlockId="10.[151,1437,151,681]" lastPageId="10" lastPageNumber="446" pageId="8" pageNumber="444">
<emphasis id="A7C7EAC6BB2FFFBAFF11E6E1FE08FDB6" bold="true" box="[199,510,906,933]" pageId="8" pageNumber="444"> <emphasis id="A7C7EAC6BB2FFFBAFF11E6E1FE08FDB6" bold="true" box="[199,510,906,933]" pageId="8" pageNumber="444">
Redescription of Redescription of
<typeStatus id="4A088876BB2FFFBAFE47E6E0FE0CFDB6" box="[401,506,907,933]" pageId="8" pageNumber="444" type="holotype">holotype</typeStatus> <typeStatus id="4A088876BB2FFFBAFE47E6E0FE0CFDB6" box="[401,506,907,933]" pageId="8" pageNumber="444" type="holotype">holotype</typeStatus>
@ -108,8 +111,17 @@ from anterior margin of cephalic plate to posterior margin of tergite 21 (
<figureCitation id="0D882A51BB2FFFBAFF15E6C4FEBEFDDA" box="[195,328,943,969]" captionStart="FIGURES 2021" captionStartId="8.[151,264,1948,1972]" captionTargetBox="[151,1435,1029,1925]" captionTargetId="figure-358@8.[151,1435,1029,1925]" captionTargetPageId="8" captionText="FIGURES 2021. Otostigmus (P.) lavanus Chamberlin, 1957 syn. nov. Holotype CAS 15215 (12-169), male. 20. General view dorsally. 21. General view ventral." figureDoi="http://doi.org/10.5281/zenodo.14022709" httpUri="https://zenodo.org/record/14022709/files/figure.png" pageId="8" pageNumber="444">Figs 20, 21</figureCitation> <figureCitation id="0D882A51BB2FFFBAFF15E6C4FEBEFDDA" box="[195,328,943,969]" captionStart="FIGURES 2021" captionStartId="8.[151,264,1948,1972]" captionTargetBox="[151,1435,1029,1925]" captionTargetId="figure-358@8.[151,1435,1029,1925]" captionTargetPageId="8" captionText="FIGURES 2021. Otostigmus (P.) lavanus Chamberlin, 1957 syn. nov. Holotype CAS 15215 (12-169), male. 20. General view dorsally. 21. General view ventral." figureDoi="http://doi.org/10.5281/zenodo.14022709" httpUri="https://zenodo.org/record/14022709/files/figure.png" pageId="8" pageNumber="444">Figs 20, 21</figureCitation>
). Antennae damaged; antennae with 11 articles (left) and 12 (right); first two basal and basal part of third glabrous [17 articles]. Cephalic plate smooth, without sutures and depressions [not described] ( ). Antennae damaged; antennae with 11 articles (left) and 12 (right); first two basal and basal part of third glabrous [17 articles]. Cephalic plate smooth, without sutures and depressions [not described] (
<figureCitation id="0D882A51BB2FFFBAFAEFE6B8FA66FDFE" box="[1337,1424,979,1005]" captionStart="FIGURES 2225" captionStartId="9.[151,264,1607,1631]" captionTargetBox="[151,1436,181,1583]" captionTargetId="figure-16@9.[151,1436,181,1583]" captionTargetPageId="9" captionText="FIGURES 2225. Otostigmus (P.) lavanus Chamberlin, 1957 syn. nov. Holotype CAS 15215 (12-169), male. 22. Cephalic plate and tergite 1, dorsally. 23. Head, forcipular segment, and sternites 1 and 2, ventrally. 24. Tergites 20 and 21, dorsally. 25. Sternites 20 and 21, ventrally." figureDoi="http://doi.org/10.5281/zenodo.14022711" httpUri="https://zenodo.org/record/14022711/files/figure.png" pageId="8" pageNumber="444">Fig. 22</figureCitation> <figureCitation id="0D882A51BB2FFFBAFAEFE6B8FA66FDFE" box="[1337,1424,979,1005]" captionStart="FIGURES 2225" captionStartId="9.[151,264,1607,1631]" captionTargetBox="[151,1436,181,1583]" captionTargetId="figure-16@9.[151,1436,181,1583]" captionTargetPageId="9" captionText="FIGURES 2225. Otostigmus (P.) lavanus Chamberlin, 1957 syn. nov. Holotype CAS 15215 (12-169), male. 22. Cephalic plate and tergite 1, dorsally. 23. Head, forcipular segment, and sternites 1 and 2, ventrally. 24. Tergites 20 and 21, dorsally. 25. Sternites 20 and 21, ventrally." figureDoi="http://doi.org/10.5281/zenodo.14022711" httpUri="https://zenodo.org/record/14022711/files/figure.png" pageId="8" pageNumber="444">Fig. 22</figureCitation>
). ). Coxosternal tooth-plates wider than long, with 4+4 teeth and a long bristle (
<figureCitation id="0D882A51BB2EFFBBFC0CE391FBDAF907" box="[986,1068,1786,1812]" captionStart="FIGURES 2225" captionStartId="9.[151,264,1607,1631]" captionTargetBox="[151,1436,181,1583]" captionTargetId="figure-16@9.[151,1436,181,1583]" captionTargetPageId="9" captionText="FIGURES 2225. Otostigmus (P.) lavanus Chamberlin, 1957 syn. nov. Holotype CAS 15215 (12-169), male. 22. Cephalic plate and tergite 1, dorsally. 23. Head, forcipular segment, and sternites 1 and 2, ventrally. 24. Tergites 20 and 21, dorsally. 25. Sternites 20 and 21, ventrally." figureDoi="http://doi.org/10.5281/zenodo.14022711" httpUri="https://zenodo.org/record/14022711/files/figure.png" pageId="9" pageNumber="445">Fig. 23</figureCitation>
); coxosternite with a short paired sutures at base of tooth-plates, this suture is bifurcate in the sides and a short median longitudinal suture [without sutures]. Trochanteroprefemoral process large, with the margin with three short tubercles and with tip pointed [not described]. Tergites 620 with complete paramedian sutures; tergites 921 with a very shallow median longitudinal keel [last tergites]; tergites 14 smooth; tergites 521 rugose on lateral sides, the rugosity increasing from the anterior to posterior tergites; tergites 921 margined [only tergite 21]. Posterior margin of tergite 21 very slightly angulate, with a shallow median longitudinal depression in the posterior border (
<figureCitation id="0D882A51BB2EFFBBFBC9E2B9FB85F9FF" box="[1055,1139,2002,2028]" captionStart="FIGURES 2225" captionStartId="9.[151,264,1607,1631]" captionTargetBox="[151,1436,181,1583]" captionTargetId="figure-16@9.[151,1436,181,1583]" captionTargetPageId="9" captionText="FIGURES 2225. Otostigmus (P.) lavanus Chamberlin, 1957 syn. nov. Holotype CAS 15215 (12-169), male. 22. Cephalic plate and tergite 1, dorsally. 23. Head, forcipular segment, and sternites 1 and 2, ventrally. 24. Tergites 20 and 21, dorsally. 25. Sternites 20 and 21, ventrally." figureDoi="http://doi.org/10.5281/zenodo.14022711" httpUri="https://zenodo.org/record/14022711/files/figure.png" pageId="9" pageNumber="445">Fig. 24</figureCitation>
); tergite 21 with a shallow short median longitudinal keel. Sternites 419 a small rounded depression in the center. Sternite 21 with posterior margin concave [triangular], shorter than precedent, converging posteriorly, with a shallow median longitudinal depression [without median sulcus]. Coxopleuron without spines. Pore-field covers almost entire coxopleuron, only the posterior end poreless. Posterior part of coxopleuron with a very short, angulate protuberance. Leg 1 without femoral and tibial spur; left leg 1 with two tarsal spurs and right leg with one; legs 1(2)19 with one tarsal spur, 20 and 21 without. Coxae 19 with a short protuberance as a very short appendix [coxa with a mesocaudal corner] (
<figureCitation id="0D882A51BB2DFFB8FAB9E420FF40FF9A" captionStart="FIGURES 2225" captionStartId="9.[151,264,1607,1631]" captionTargetBox="[151,1436,181,1583]" captionTargetId="figure-16@9.[151,1436,181,1583]" captionTargetPageId="9" captionText="FIGURES 2225. Otostigmus (P.) lavanus Chamberlin, 1957 syn. nov. Holotype CAS 15215 (12-169), male. 22. Cephalic plate and tergite 1, dorsally. 23. Head, forcipular segment, and sternites 1 and 2, ventrally. 24. Tergites 20 and 21, dorsally. 25. Sternites 20 and 21, ventrally." figureDoi="http://doi.org/10.5281/zenodo.14022711" httpUri="https://zenodo.org/record/14022711/files/figure.png" pageId="10" pageNumber="446">Fig. 25</figureCitation>
); coxae 20 with a larger appendix curved posteriorly [coxae of the twentieth legs with a conspicuous process, curved distally and extending a little beyond the last sternite]. This appendix exceeds sternite 21 (
<figureCitation id="0D882A51BB2DFFB8FB18E4F8FAD4FFBE" box="[1230,1314,402,429]" captionStart="FIGURES 2225" captionStartId="9.[151,264,1607,1631]" captionTargetBox="[151,1436,181,1583]" captionTargetId="figure-16@9.[151,1436,181,1583]" captionTargetPageId="9" captionText="FIGURES 2225. Otostigmus (P.) lavanus Chamberlin, 1957 syn. nov. Holotype CAS 15215 (12-169), male. 22. Cephalic plate and tergite 1, dorsally. 23. Head, forcipular segment, and sternites 1 and 2, ventrally. 24. Tergites 20 and 21, dorsally. 25. Sternites 20 and 21, ventrally." figureDoi="http://doi.org/10.5281/zenodo.14022711" httpUri="https://zenodo.org/record/14022711/files/figure.png" pageId="10" pageNumber="446">Fig. 25</figureCitation>
). Prefemur of ultimate legs without spines, with a digitiform appendix almost the same length as the prefemur; the digitiform appendix is dorsoventrally flattened and with a tuft of hair at its tip. This tuft of hair is localized dorsally.
</paragraph> </paragraph>
</subSubSection>
<caption id="C1CC665CBB2FFFBAFF41E2F7FE00F9CA" ID-DOI="http://doi.org/10.5281/zenodo.14022709" ID-Zenodo-Dep="14022709" httpUri="https://zenodo.org/record/14022709/files/figure.png" pageId="8" pageNumber="444" startId="8.[151,264,1948,1972]" targetBox="[151,1435,1029,1925]" targetPageId="8" targetType="figure"> <caption id="C1CC665CBB2FFFBAFF41E2F7FE00F9CA" ID-DOI="http://doi.org/10.5281/zenodo.14022709" ID-Zenodo-Dep="14022709" httpUri="https://zenodo.org/record/14022709/files/figure.png" pageId="8" pageNumber="444" startId="8.[151,264,1948,1972]" targetBox="[151,1435,1029,1925]" targetPageId="8" targetType="figure">
<paragraph id="950C36D4BB2FFFBAFF41E2F7FE00F9CA" blockId="8.[151,1436,1948,2009]" pageId="8" pageNumber="444"> <paragraph id="950C36D4BB2FFFBAFF41E2F7FE00F9CA" blockId="8.[151,1436,1948,2009]" pageId="8" pageNumber="444">
<emphasis id="A7C7EAC6BB2FFFBAFF41E2F7FEA5F9A6" bold="true" box="[151,339,1948,1973]" pageId="8" pageNumber="444">FIGURES 2021.</emphasis> <emphasis id="A7C7EAC6BB2FFFBAFF41E2F7FEA5F9A6" bold="true" box="[151,339,1948,1973]" pageId="8" pageNumber="444">FIGURES 2021.</emphasis>
@ -160,18 +172,6 @@ Tergites 20 and 21, dorsally.
Sternites 20 and 21, ventrally. Sternites 20 and 21, ventrally.
</paragraph> </paragraph>
</caption> </caption>
<paragraph id="950C36D4BB2EFFB8FF41E391FAC6FFE6" blockId="9.[151,1437,1786,2028]" lastBlockId="10.[151,1437,151,681]" lastPageId="10" lastPageNumber="446" pageId="9" pageNumber="445">
Coxosternal tooth-plates wider than long, with 4+4 teeth and a long bristle (
<figureCitation id="0D882A51BB2EFFBBFC0CE391FBDAF907" box="[986,1068,1786,1812]" captionStart="FIGURES 2225" captionStartId="9.[151,264,1607,1631]" captionTargetBox="[151,1436,181,1583]" captionTargetId="figure-16@9.[151,1436,181,1583]" captionTargetPageId="9" captionText="FIGURES 2225. Otostigmus (P.) lavanus Chamberlin, 1957 syn. nov. Holotype CAS 15215 (12-169), male. 22. Cephalic plate and tergite 1, dorsally. 23. Head, forcipular segment, and sternites 1 and 2, ventrally. 24. Tergites 20 and 21, dorsally. 25. Sternites 20 and 21, ventrally." figureDoi="http://doi.org/10.5281/zenodo.14022711" httpUri="https://zenodo.org/record/14022711/files/figure.png" pageId="9" pageNumber="445">Fig. 23</figureCitation>
); coxosternite with a short paired sutures at base of tooth-plates, this suture is bifurcate in the sides and a short median longitudinal suture [without sutures]. Trochanteroprefemoral process large, with the margin with three short tubercles and with tip pointed [not described]. Tergites 620 with complete paramedian sutures; tergites 921 with a very shallow median longitudinal keel [last tergites]; tergites 14 smooth; tergites 521 rugose on lateral sides, the rugosity increasing from the anterior to posterior tergites; tergites 921 margined [only tergite 21]. Posterior margin of tergite 21 very slightly angulate, with a shallow median longitudinal depression in the posterior border (
<figureCitation id="0D882A51BB2EFFBBFBC9E2B9FB85F9FF" box="[1055,1139,2002,2028]" captionStart="FIGURES 2225" captionStartId="9.[151,264,1607,1631]" captionTargetBox="[151,1436,181,1583]" captionTargetId="figure-16@9.[151,1436,181,1583]" captionTargetPageId="9" captionText="FIGURES 2225. Otostigmus (P.) lavanus Chamberlin, 1957 syn. nov. Holotype CAS 15215 (12-169), male. 22. Cephalic plate and tergite 1, dorsally. 23. Head, forcipular segment, and sternites 1 and 2, ventrally. 24. Tergites 20 and 21, dorsally. 25. Sternites 20 and 21, ventrally." figureDoi="http://doi.org/10.5281/zenodo.14022711" httpUri="https://zenodo.org/record/14022711/files/figure.png" pageId="9" pageNumber="445">Fig. 24</figureCitation>
); tergite 21 with a shallow short median longitudinal keel. Sternites 419 a small rounded depression in the center. Sternite 21 with posterior margin concave [triangular], shorter than precedent, converging posteriorly, with a shallow median longitudinal depression [without median sulcus]. Coxopleuron without spines. Pore-field covers almost entire coxopleuron, only the posterior end poreless. Posterior part of coxopleuron with a very short, angulate protuberance. Leg 1 without femoral and tibial spur; left leg 1 with two tarsal spurs and right leg with one; legs 1(2)19 with one tarsal spur, 20 and 21 without. Coxae 19 with a short protuberance as a very short appendix [coxa with a mesocaudal corner] (
<figureCitation id="0D882A51BB2DFFB8FAB9E420FF40FF9A" captionStart="FIGURES 2225" captionStartId="9.[151,264,1607,1631]" captionTargetBox="[151,1436,181,1583]" captionTargetId="figure-16@9.[151,1436,181,1583]" captionTargetPageId="9" captionText="FIGURES 2225. Otostigmus (P.) lavanus Chamberlin, 1957 syn. nov. Holotype CAS 15215 (12-169), male. 22. Cephalic plate and tergite 1, dorsally. 23. Head, forcipular segment, and sternites 1 and 2, ventrally. 24. Tergites 20 and 21, dorsally. 25. Sternites 20 and 21, ventrally." figureDoi="http://doi.org/10.5281/zenodo.14022711" httpUri="https://zenodo.org/record/14022711/files/figure.png" pageId="10" pageNumber="446">Fig. 25</figureCitation>
); coxae 20 with a larger appendix curved posteriorly [coxae of the twentieth legs with a conspicuous process, curved distally and extending a little beyond the last sternite]. This appendix exceeds sternite 21 (
<figureCitation id="0D882A51BB2DFFB8FB18E4F8FAD4FFBE" box="[1230,1314,402,429]" captionStart="FIGURES 2225" captionStartId="9.[151,264,1607,1631]" captionTargetBox="[151,1436,181,1583]" captionTargetId="figure-16@9.[151,1436,181,1583]" captionTargetPageId="9" captionText="FIGURES 2225. Otostigmus (P.) lavanus Chamberlin, 1957 syn. nov. Holotype CAS 15215 (12-169), male. 22. Cephalic plate and tergite 1, dorsally. 23. Head, forcipular segment, and sternites 1 and 2, ventrally. 24. Tergites 20 and 21, dorsally. 25. Sternites 20 and 21, ventrally." figureDoi="http://doi.org/10.5281/zenodo.14022711" httpUri="https://zenodo.org/record/14022711/files/figure.png" pageId="10" pageNumber="446">Fig. 25</figureCitation>
). Prefemur of ultimate legs without spines, with a digitiform appendix almost the same length as the prefemur; the digitiform appendix is dorsoventrally flattened and with a tuft of hair at its tip. This tuft of hair is localized dorsally.
</paragraph>
</subSubSection>
<subSubSection id="DDA9655FBB2DFFB8FF11E494FE52FCBA" pageId="10" pageNumber="446" type="discussion"> <subSubSection id="DDA9655FBB2DFFB8FF11E494FE52FCBA" pageId="10" pageNumber="446" type="discussion">
<paragraph id="950C36D4BB2DFFB8FF11E494FE52FCBA" blockId="10.[151,1437,151,681]" pageId="10" pageNumber="446"> <paragraph id="950C36D4BB2DFFB8FF11E494FE52FCBA" blockId="10.[151,1437,151,681]" pageId="10" pageNumber="446">
<emphasis id="A7C7EAC6BB2DFFB8FF11E494FECDFC0A" bold="true" box="[199,315,511,537]" pageId="10" pageNumber="446">Remarks.</emphasis> <emphasis id="A7C7EAC6BB2DFFB8FF11E494FECDFC0A" bold="true" box="[199,315,511,537]" pageId="10" pageNumber="446">Remarks.</emphasis>

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@ -1,54 +1,56 @@
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<mods:title id="ED6146E98F6825081109D592FB449D2A">A monograph of the Oligochaete family Alluroididae</mods:title> <mods:title id="DE406E74C0F447E2C0E5A89D9334211F">A monograph of the Oligochaete family Alluroididae</mods:title>
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@ -77,9 +79,9 @@ sp.
<bibRefCitation id="8B1D4B787943FFE6CF0FF883FDCFF8A8" author="Coates, K. A. &amp; Stacey, D. F." box="[321,587,1792,1816]" pageId="21" pageNumber="422" pagination="79 - 84" refId="ref22798" refString="Coates, K. A. &amp; Stacey, D. F. (1994) Oligochaetes (Naididae, Tubificidae, Enchytraeidae and Alluroididae) of Guyana, Peru and Ecuador. Hydrobiologia, 278, 79 - 84. https: // doi. org / 10.1007 / BF 00142313" type="journal article" year="1994">Coates &amp; Stacey 1994: 81</bibRefCitation> <bibRefCitation id="8B1D4B787943FFE6CF0FF883FDCFF8A8" author="Coates, K. A. &amp; Stacey, D. F." box="[321,587,1792,1816]" pageId="21" pageNumber="422" pagination="79 - 84" refId="ref22798" refString="Coates, K. A. &amp; Stacey, D. F. (1994) Oligochaetes (Naididae, Tubificidae, Enchytraeidae and Alluroididae) of Guyana, Peru and Ecuador. Hydrobiologia, 278, 79 - 84. https: // doi. org / 10.1007 / BF 00142313" type="journal article" year="1994">Coates &amp; Stacey 1994: 81</bibRefCitation>
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, ,
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, ,
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@ -91,7 +93,7 @@ sp. 2
. .
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: :
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and and
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. .