Add updates up until 2024-08-30 14:26:50

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</subSubSection>
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<paragraph id="8BBE452BF569FFCDC8F63B9EFF5EFD77" blockId="4.[151,658,618,681]" box="[151,237,655,681]" pageId="4" pageNumber="271">
<figureCitation id="133A59AEF569FFCDC8F63B9EFF5EFD77" box="[151,237,655,681]" captionStart="FIGURE 4" captionStartId="7.[152,255,666,690]" captionTargetBox="[151,1436,181,643]" captionTargetId="figure-26@7.[151,1436,181,643]" captionTargetPageId="7" captionText="FIGURE 4. Geographic records of Polyphrix. A. Polyphrix stellata; B. Polyphrix varians. C. Polyphrix misa sp. nov. and Polyphrix tullu sp. nov. Yellow circles represent new records and red represent previous records." figureDoi="http://doi.org/10.5281/zenodo.13618454" httpUri="https://zenodo.org/record/13618454/files/figure.png" pageId="4" pageNumber="271">Fig. 4A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31B16A0F569FFCDC8F63BC7FCF5FCE7" pageId="4" pageNumber="271" type="materials_examined">
<paragraph id="8BBE452BF569FFCDC8F63BC7FCF5FCE7" blockId="4.[151,1437,726,1041]" pageId="4" pageNumber="271">
<emphasis id="B9759939F569FFCDC8F63BC7FECFFD2E" bold="true" box="[151,380,726,752]" pageId="4" pageNumber="271">Material examined</emphasis>
.
<materialsCitation id="3B694F76F569FFCDC9EC3BC6FCA5FCCB" collectingDate="2017-07-01" collectionCode="MD, WIRED, AMAZON, PROYECT, TRAP" country="Peru" elevation="231" location="Albergue Refugio" pageId="4" pageNumber="271" specimenCount="1" specimenCount-female="1" stateProvince="Amazonas">
<specimenCount id="9D078EA2F569FFCDC9EC3BC6FE03FD2F" box="[397,432,727,753]" count="1" pageId="4" pageNumber="271" type="female">1♀</specimenCount>
<collectingCountry id="F31605BBF569FFCDC9A53BC6FDA2FD2F" box="[452,529,727,753]" name="Peru" pageId="4" pageNumber="271">PERU</collectingCountry>
,
<collectionCode id="ED10DDEEF569FFCDCA7F3BC6FDFDFD2F" box="[542,590,727,753]" country="Germany" name="Museum Donaueschingen" pageId="4" pageNumber="271">MD</collectionCode>
,
<location id="8EDE13F0F569FFCDCA3A3BC6FC95FD2F" LSID="urn:lsid:plazi:treatment:03A8F43DF569FFCDC8F63B7BFEB3FBCF:8EDE13F0F569FFCDCA3A3BC6FC95FD2F" box="[603,806,727,753]" country="Peru" name="Albergue Refugio" pageId="4" pageNumber="271" stateProvince="Amazonas">Albergue Refugio</location>
<collectingRegion id="49C58BC9F569FFCDCB4F3BC6FC17FD2F" box="[814,932,727,753]" country="Peru" name="Amazonas" pageId="4" pageNumber="271">Amazonas</collectingRegion>
12º5230”/69º2435”
<quantity id="4CF9E8CEF569FFCDCCC43BC6FB5DFD2F" box="[1189,1262,727,753]" metricMagnitude="2" metricUnit="m" metricValue="2.31" pageId="4" pageNumber="271" unit="m" value="231.0">
<elevation id="002CA218F569FFCDCCC43BC6FB5DFD2F" box="[1189,1262,727,753]" metricMagnitude="2" metricUnit="m" metricValue="2.31" pageId="4" pageNumber="271" unit="m" value="231.0">231 m</elevation>
</quantity>
<date id="FFBF63EBF569FFCDCC993BC6FAC4FD2F" box="[1272,1399,727,753]" pageId="4" pageNumber="271" value="2017-07-01">
<collectingDate id="EFFB9A03F569FFCDCC993BC6FAC4FD2F" box="[1272,1399,727,753]" pageId="4" pageNumber="271" value="2017-07-01">01.vii.2017</collectingDate>
</date>
D. Couceiro //
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PAN
<collectionCode id="ED10DDEEF569FFCDCAA13BEAFCBEFCCB" box="[704,781,763,789]" pageId="4" pageNumber="271">TRAP</collectionCode>
</materialsCitation>
,
<materialsCitation id="3B694F76F569FFCDCB433BEAFB8BFCCB" box="[802,1080,763,789]" collectingDate="2017-07-17" collectionCode="MD, WIRED, AMAZON, PROYECT, TRAP" country="Peru" elevation="231" location="Albergue Refugio" pageId="4" pageNumber="271" specimenCount="1" specimenCount-female="1" stateProvince="Amazonas">
<specimenCount id="9D078EA2F569FFCDCB433BEAFCF6FCCB" box="[802,837,763,789]" count="1" pageId="4" pageNumber="271" type="female">1♀</specimenCount>
idem but
<date id="FFBF63EBF569FFCDCBD43BEAFB8BFCCB" box="[949,1080,763,789]" pageId="4" pageNumber="271" value="2017-07-17">
<collectingDate id="EFFB9A03F569FFCDCBD43BEAFB8BFCCB" box="[949,1080,763,789]" pageId="4" pageNumber="271" value="2017-07-17">17.vii.2017</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="3B694F76F569FFCDCC223BEAFAEBFCCB" box="[1091,1368,763,789]" collectingDate="2017-07-21" collectionCode="MD, WIRED, AMAZON, PROYECT, TRAP" country="Peru" elevation="231" location="Albergue Refugio" pageId="4" pageNumber="271" specimenCount="1" specimenCount-female="1" stateProvince="Amazonas">
<specimenCount id="9D078EA2F569FFCDCC223BEAFBD5FCCB" box="[1091,1126,763,789]" count="1" pageId="4" pageNumber="271" type="female">1♀</specimenCount>
idem but
<date id="FFBF63EBF569FFCDCCB63BEAFAEBFCCB" box="[1239,1368,763,789]" pageId="4" pageNumber="271" value="2017-07-21">
<collectingDate id="EFFB9A03F569FFCDCCB63BEAFAEBFCCB" box="[1239,1368,763,789]" pageId="4" pageNumber="271" value="2017-07-21">21.vii.2017</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="3B694F76F569FFCDCD053BEAFDAAFCE7" collectingDate="2016-11-21" collectionCode="MD, WIRED, AMAZON, PROYECT, TRAP" country="Peru" elevation="231" location="Albergue Refugio" pageId="4" pageNumber="271" specimenCount="2" specimenCount-female="2" stateProvince="Amazonas">
<specimenCount id="9D078EA2F569FFCDCD053BEAFA2FFCCB" box="[1380,1436,763,789]" count="2" pageId="4" pageNumber="271" type="female">2♀♀</specimenCount>
idem but
<date id="FFBF63EBF569FFCDC9603A0EFEC8FCE7" box="[257,379,799,825]" pageId="4" pageNumber="271" value="2016-11-21">
<collectingDate id="EFFB9A03F569FFCDC9603A0EFEC8FCE7" box="[257,379,799,825]" pageId="4" pageNumber="271" value="2016-11-21">21.xi.2016</collectingDate>
</date>
, and idem but
</materialsCitation>
<materialsCitation id="3B694F76F569FFCDCA413A0EFCF1FCE7" box="[544,834,798,825]" collectingDate="2016-10-05" collectionCode="MUSM" country="Peru" pageId="4" pageNumber="271" specimenCount="2" specimenCount-female="2">
<specimenCount id="9D078EA2F569FFCDCA413A0EFDEBFCE7" box="[544,600,799,825]" count="2" pageId="4" pageNumber="271" type="female">2♀♀</specimenCount>
<date id="FFBF63EBF569FFCDCA3E3A0FFD7CFCE7" box="[607,719,798,825]" pageId="4" pageNumber="271" value="2016-10-05">
<collectingDate id="EFFB9A03F569FFCDCA3E3A0FFD7CFCE7" box="[607,719,798,825]" pageId="4" pageNumber="271" value="2016-10-05">05.x.2016</collectingDate>
</date>
(
<collectionCode id="ED10DDEEF569FFCDCABF3A0EFC89FCE7" box="[734,826,799,825]" pageId="4" pageNumber="271">MUSM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31B16A0F569FFCDC8A63A53FCA8FC17" pageId="4" pageNumber="271" type="discussion">
<paragraph id="8BBE452BF569FFCDC8A63A53FCA8FC17" blockId="4.[151,1437,726,1041]" pageId="4" pageNumber="271">
<emphasis id="B9759939F569FFCDC8A63A53FEE1FC82" bold="true" box="[199,338,834,860]" pageId="4" pageNumber="271">Comments:</emphasis>
This species was collected in the same locality as one of the
<typeStatus id="54BAFB89F569FFCDCC6A3A52FBC5FC83" box="[1035,1142,835,861]" pageId="4" pageNumber="271" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="4C013EA8F569FFCDCCFC3A52FAB2FC83" authorityName="Inga &amp; Alvarado" authorityYear="2024" box="[1181,1281,835,861]" class="Insecta" family="Ichneumonidae" genus="Polyphrix" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="271" phylum="Arthropoda" rank="species" species="sefirot" status="sp. nov.">
<emphasis id="B9759939F569FFCDCCFC3A52FAB2FC83" box="[1181,1281,835,861]" italics="true" pageId="4" pageNumber="271">P. sefirot</emphasis>
</taxonomicName>
<emphasis id="B9759939F569FFCDCD693A52FAD0FC83" bold="true" box="[1288,1379,835,861]" pageId="4" pageNumber="271">
<taxonomicNameLabel id="A2462442F569FFCDCD693A52FAD0FC83" box="[1288,1379,835,861]" pageId="4" pageNumber="271" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
This species was collected with yellow pan traps as were the type specimen of the species (
<bibRefCitation id="EF9038DAF569FFCDCCEB3A76FAD0FC5F" author="Santos, B. F. &amp; Aguiar, A. P. &amp; Tedesco, A. M." box="[1162,1379,871,897]" pageId="4" pageNumber="271" pagination="29 - 44" refId="ref5455" refString="Santos, B. F., Aguiar, A. P. &amp; Tedesco, A. M. (2009) Phylogenetic revision and the origin of Polyphrix Townes (Hymenoptera, Ichneumonidae, Cryptinae), with description of a new species. Zootaxa, 2214 (1), 29 - 44. https: // doi. org / 10.11646 / zootaxa. 2214.1.2" type="journal article" year="2009">
Santos
<emphasis id="B9759939F569FFCDCC813A76FAAFFC5F" box="[1248,1308,871,897]" italics="true" pageId="4" pageNumber="271">et al.</emphasis>
2009
</bibRefCitation>
). We installed pan traps from
<date id="FFBF63EBF569FFCDC9C63A9AFDF8FC7B" box="[423,587,907,933]" pageId="4" pageNumber="271" value="2016-02">February 2016</date>
to
<date id="FFBF63EBF569FFCDCA0F3A9AFC90FC7B" box="[622,803,907,933]" pageId="4" pageNumber="271" value="2017-12">December 2017</date>
, but this species was only collected during July, October and November and was absent during the rest of the year.
</paragraph>
</subSubSection>
<subSubSection id="C31B16A0F569FFCDC8A63AC3FEB3FBCF" pageId="4" pageNumber="271" type="distribution">
<paragraph id="8BBE452BF569FFCDC8A63AC3FEB3FBCF" blockId="4.[151,1437,726,1041]" pageId="4" pageNumber="271">
<emphasis id="B9759939F569FFCDC8A63AC3FE72FC33" bold="true" box="[199,449,978,1005]" pageId="4" pageNumber="271">Distribution records.</emphasis>
<collectingCountry id="F31605BBF569FFCDC9A93AC2FDBFFC33" box="[456,524,979,1005]" name="Brazil" pageId="4" pageNumber="271">Brazil</collectingCountry>
(
<bibRefCitation id="EF9038DAF569FFCDCA7A3AC2FD58FC33" author="Santos, B. F. &amp; Aguiar, A. P. &amp; Tedesco, A. M." box="[539,747,979,1005]" pageId="4" pageNumber="271" pagination="29 - 44" refId="ref5455" refString="Santos, B. F., Aguiar, A. P. &amp; Tedesco, A. M. (2009) Phylogenetic revision and the origin of Polyphrix Townes (Hymenoptera, Ichneumonidae, Cryptinae), with description of a new species. Zootaxa, 2214 (1), 29 - 44. https: // doi. org / 10.11646 / zootaxa. 2214.1.2" type="journal article" year="2009">
Santos
<emphasis id="B9759939F569FFCDCA0E3AC2FD14FC33" box="[623,679,979,1005]" italics="true" pageId="4" pageNumber="271">et al.</emphasis>
2009
</bibRefCitation>
). In
<collectingCountry id="F31605BBF569FFCDCB7A3AC2FCFDFC33" box="[795,846,979,1005]" name="Peru" pageId="4" pageNumber="271">Peru</collectingCountry>
is known to occur in the
<collectingRegion id="49C58BC9F569FFCDCC0A3AC2FAA2FC33" box="[1131,1297,979,1005]" country="Peru" name="Madre de Dios" pageId="4" pageNumber="271">Madre de Dios</collectingRegion>
Department (
<figureCitation id="133A59AEF569FFCDC8FE3AE6FF40FBCF" box="[159,243,1015,1041]" captionStart="FIGURE 4" captionStartId="7.[152,255,666,690]" captionTargetBox="[151,1436,181,643]" captionTargetId="figure-26@7.[151,1436,181,643]" captionTargetPageId="7" captionText="FIGURE 4. Geographic records of Polyphrix. A. Polyphrix stellata; B. Polyphrix varians. C. Polyphrix misa sp. nov. and Polyphrix tullu sp. nov. Yellow circles represent new records and red represent previous records." figureDoi="http://doi.org/10.5281/zenodo.13618454" httpUri="https://zenodo.org/record/13618454/files/figure.png" pageId="4" pageNumber="271">Fig 4A</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,278 @@
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<mods:title id="49ABDC49BD8B15F373A6DE681B9379C6">Two new species of the genus Polyphrix Townes 1972 (Hymenoptera, Ichneumonidae, Cryptinae) from Peru</mods:title>
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<mods:namePart id="5EB98B990D455B6F206CEA345EDE73E1">Inga, Lita</mods:namePart>
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<mods:namePart id="95BF6C22A563958BA979359B1A86C375">Alvarado, Mabel</mods:namePart>
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<mods:nameIdentifier id="9AAEC3525BD38C47F81D5E87A987121C" type="email">malvaradog@unmsm.edu.pe</mods:nameIdentifier>
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<paragraph id="8BBE452BF569FFCDC8F63D73FE00FBA3" blockId="4.[151,435,1122,1185]" box="[151,435,1122,1149]" pageId="4" pageNumber="271">
<heading id="D0F6F247F569FFCDC8F63D73FE00FBA3" bold="true" box="[151,435,1122,1149]" fontSize="11" level="1" pageId="4" pageNumber="271" reason="1">
<emphasis id="B9759939F569FFCDC8F63D73FE00FBA3" bold="true" box="[151,435,1122,1149]" pageId="4" pageNumber="271">
<taxonomicName id="4C013EA8F569FFCDC8F63D73FEFDFBA3" authorityName="Inga &amp; Alvarado" authorityYear="2024" box="[151,334,1122,1149]" class="Insecta" family="Ichneumonidae" genus="Polyphrix" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="271" phylum="Arthropoda" rank="species" species="tullu" status="sp. nov.">
<emphasis id="B9759939F569FFCDC8F63D73FEFDFBA3" bold="true" box="[151,334,1122,1149]" italics="true" pageId="4" pageNumber="271">Polyphrix tullu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2462442F569FFCDC9373D72FE00FBA3" box="[342,435,1123,1149]" pageId="4" pageNumber="271" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31B16A0F569FFCDC8F63D96FEBFFB7F" box="[151,268,1159,1185]" pageId="4" pageNumber="271" type="description">
<paragraph id="8BBE452BF569FFCDC8F63D96FEBFFB7F" blockId="4.[151,435,1122,1185]" box="[151,268,1159,1185]" pageId="4" pageNumber="271">
<figureCitation id="133A59AEF569FFCDC8F63D96FF6EFB7F" box="[151,221,1159,1185]" captionStart="FIGURE 3" captionStartId="5.[152,255,1792,1816]" captionTargetBox="[151,1435,181,1768]" captionTargetId="figure-26@5.[151,1435,181,1768]" captionTargetPageId="5" captionText="FIGURE 3. Polyphrix tullu sp. nov., female, holotype. A. habitus, lateral view; B. head, front view; C. pronotum, lateral view; D. mesosoma and metasoma, dorsal view; E. wings; F. ovipositor tip, lateral view. Scale bar = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.13618452" httpUri="https://zenodo.org/record/13618452/files/figure.png" pageId="4" pageNumber="271">Figs 3</figureCitation>
,
<figureCitation id="133A59AEF569FFCDC88B3D96FEBFFB7F" box="[234,268,1159,1185]" captionStart="FIGURE 4" captionStartId="7.[152,255,666,690]" captionTargetBox="[151,1436,181,643]" captionTargetId="figure-26@7.[151,1436,181,643]" captionTargetPageId="7" captionText="FIGURE 4. Geographic records of Polyphrix. A. Polyphrix stellata; B. Polyphrix varians. C. Polyphrix misa sp. nov. and Polyphrix tullu sp. nov. Yellow circles represent new records and red represent previous records." figureDoi="http://doi.org/10.5281/zenodo.13618454" httpUri="https://zenodo.org/record/13618454/files/figure.png" pageId="4" pageNumber="271">4C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31B16A0F569FFCDC8F63DDFFCB4FA8B" pageId="4" pageNumber="271" type="materials_examined">
<paragraph id="8BBE452BF569FFCDC8F63DDFFCB4FA8B" blockId="4.[151,1437,1230,2013]" pageId="4" pageNumber="271">
<emphasis id="B9759939F569FFCDC8F63DDFFE36FB36" bold="true" box="[151,389,1230,1256]" pageId="4" pageNumber="271">Material examined.</emphasis>
<materialsCitation id="3B694F76F569FFCDC9F03DDEFC6BFAD3" collectingDate="2016-06-11" collectionCode="MD, MUSM" collectorName="N. Zenteno" country="Peru" elevation="228" latitude="-12.557139" location="Manu" longLatPrecision="1" longitude="-70.07961" municipality="Los Amigos Biological Station" pageId="4" pageNumber="271" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="B9759939F569FFCDC9F03DDEFDB4FB37" bold="true" box="[401,519,1231,1257]" pageId="4" pageNumber="271">
<typeStatus id="54BAFB89F569FFCDC9F03DDEFDB1FB37" box="[401,514,1231,1257]" pageId="4" pageNumber="271" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="9D078EA2F569FFCDCA723DDEFD85FB37" box="[531,566,1231,1257]" count="1" pageId="4" pageNumber="271" type="female">1♀</specimenCount>
<collectingCountry id="F31605BBF569FFCDCA2D3DDEFD29FB37" box="[588,666,1231,1257]" name="Peru" pageId="4" pageNumber="271">PERU</collectingCountry>
:
<collectionCode id="ED10DDEEF569FFCDCAC83DDEFD69FB37" box="[681,730,1231,1257]" country="Germany" name="Museum Donaueschingen" pageId="4" pageNumber="271">MD</collectionCode>
.
<location id="8EDE13F0F569FFCDCA883DDEFC9EFB37" LSID="urn:lsid:plazi:treatment:03A8F43DF569FFCFC8F63D73FB33FAD3:8EDE13F0F569FFCDCA883DDEFC9EFB37" box="[745,813,1231,1257]" country="Peru" latitude="-12.557139" longLatPrecision="1" longitude="-70.07961" municipality="Los Amigos Biological Station" name="Manu" pageId="4" pageNumber="271">Manu</location>
,
<collectingMunicipality id="6BDADF51F569FFCDCB5C3DDEFB1BFB37" box="[829,1192,1231,1257]" pageId="4" pageNumber="271">Los Amigos Biological Station</collectingMunicipality>
,
<quantity id="4CF9E8CEF569FFCDCCD93DDFFB4CFB37" box="[1208,1279,1230,1257]" metricMagnitude="2" metricUnit="m" metricValue="2.28" pageId="4" pageNumber="271" unit="m" value="228.0">
<elevation id="002CA218F569FFCDCCD93DDFFB4CFB37" box="[1208,1279,1230,1257]" metricMagnitude="2" metricUnit="m" metricValue="2.28" pageId="4" pageNumber="271" unit="m" value="228.0">228m</elevation>
</quantity>
,
<geoCoordinate id="EE3523ECF569FFCDCD653DDFFA2FFB37" box="[1284,1436,1230,1257]" degrees="12" direction="south" minutes="33" orientation="latitude" pageId="4" pageNumber="271" precision="1" seconds="25.7" value="-12.557139">12°3325.7S</geoCoordinate>
<geoCoordinate id="EE3523ECF569FFCDC8F63DE2FE8CFAD3" box="[151,319,1267,1293]" degrees="70" direction="west" minutes="04" orientation="longitude" pageId="4" pageNumber="271" precision="1" seconds="46.6" value="-70.07961">70°0446.6W</geoCoordinate>
,
<date id="FFBF63EBF569FFCDC92E3DE2FE7BFAD3" box="[335,456,1267,1293]" pageId="4" pageNumber="271" value="2016-06-11">
<collectingDate id="EFFB9A03F569FFCDC92E3DE2FE7BFAD3" box="[335,456,1267,1293]" pageId="4" pageNumber="271" value="2016-06-11">11.vi.2016</collectingDate>
</date>
,
<collectingMethod id="52403D3CF569FFCDC9B73DE2FD22FAD3" box="[470,657,1267,1293]" pageId="4" pageNumber="271">Yellow pan trap</collectingMethod>
,
<collectorName id="26F420FDF569FFCDCAC13DE2FC92FAD3" box="[672,801,1267,1293]" pageId="4" pageNumber="271">N. Zenteno</collectorName>
leg.” (
<collectionCode id="ED10DDEEF569FFCDCB153DE2FC63FAD3" box="[884,976,1267,1293]" pageId="4" pageNumber="271">MUSM</collectionCode>
)
</materialsCitation>
.
<emphasis id="B9759939F569FFCDCB843DE2FBEFFAD3" bold="true" box="[997,1116,1267,1293]" pageId="4" pageNumber="271">
<typeStatus id="54BAFB89F569FFCDCB843DE2FBE5FAD3" box="[997,1110,1267,1293]" pageId="4" pageNumber="271" type="paratype">Paratype</typeStatus>
:
</emphasis>
<specimenCount id="9D078EA2F569FFCDCC073DE2FB3AFAD3" box="[1126,1161,1267,1293]" count="1" pageId="4" pageNumber="271" type="female">1♀</specimenCount>
similar to
<typeStatus id="54BAFB89F569FFCDCD6D3DE2FADDFAD3" box="[1292,1390,1267,1293]" pageId="4" pageNumber="271" type="holotype">holotype</typeStatus>
but
<date id="FFBF63EBF569FFCDC8F63C06FEBCFAEF" box="[151,271,1303,1329]" pageId="4" pageNumber="271" value="2016-06-21">21.vi.2016</date>
and
<materialsCitation id="3B694F76F569FFCDC9263C06FE49FA8B" collectingDate="2016-02-24" collectionCode="MD, WIRED, AMAZON, PROYECT" collectorName="Grados" country="Peru" elevation="231" location="Albergue Refugio" pageId="4" pageNumber="271" specimenCount="1" specimenCount-female="1" stateProvince="Amazonas" typeStatus="paratype">
<specimenCount id="9D078EA2F569FFCDC9263C06FED9FAEF" box="[327,362,1303,1329]" count="1" pageId="4" pageNumber="271" type="female">1♀</specimenCount>
<collectingCountry id="F31605BBF569FFCDC91D3C06FE7AFAEF" box="[380,457,1303,1329]" name="Peru" pageId="4" pageNumber="271">PERU</collectingCountry>
,
<collectionCode id="ED10DDEEF569FFCDC9B43C06FDB6FAEF" box="[469,517,1303,1329]" country="Germany" name="Museum Donaueschingen" pageId="4" pageNumber="271">MD</collectionCode>
,
<location id="8EDE13F0F569FFCDCA6E3C06FD6AFAEF" LSID="urn:lsid:plazi:treatment:03A8F43DF569FFCFC8F63D73FB33FAD3:8EDE13F0F569FFCDCA6E3C06FD6AFAEF" box="[527,729,1303,1329]" country="Peru" name="Albergue Refugio" pageId="4" pageNumber="271" stateProvince="Amazonas">Albergue Refugio</location>
<collectingRegion id="49C58BC9F569FFCDCABE3C06FCE6FAEF" box="[735,853,1303,1329]" country="Peru" name="Amazonas" pageId="4" pageNumber="271">Amazonas</collectingRegion>
12º5230”/69º2435”
<quantity id="4CF9E8CEF569FFCDCC303C06FB2AFAEF" box="[1105,1177,1303,1329]" metricMagnitude="2" metricUnit="m" metricValue="2.31" pageId="4" pageNumber="271" unit="m" value="231.0">
<elevation id="002CA218F569FFCDCC303C06FB2AFAEF" box="[1105,1177,1303,1329]" metricMagnitude="2" metricUnit="m" metricValue="2.31" pageId="4" pageNumber="271" unit="m" value="231.0">231 m</elevation>
</quantity>
<date id="FFBF63EBF569FFCDCCC03C06FAA0FAEF" box="[1185,1299,1303,1329]" pageId="4" pageNumber="271" value="2016-02-24">
<collectingDate id="EFFB9A03F569FFCDCCC03C06FAA0FAEF" box="[1185,1299,1303,1329]" pageId="4" pageNumber="271" value="2016-02-24">24.ii.2016</collectingDate>
</date>
J.
<collectorName id="26F420FDF569FFCDCD553C06FA36FAEF" box="[1332,1413,1303,1329]" pageId="4" pageNumber="271">Grados</collectorName>
//
<collectionCode id="ED10DDEEF569FFCDC8F63C2AFE49FA8B" box="[151,506,1339,1365]" pageId="4" pageNumber="271">WIRED AMAZON PROYECT</collectionCode>
</materialsCitation>
PAN
<collectionCode id="ED10DDEEF569FFCDCA5D3C2AFD35FA8B" box="[572,646,1339,1365]" pageId="4" pageNumber="271">TRAP</collectionCode>
<materialsCitation id="3B694F76F569FFCDCAF63C2AFCB0FA8B" box="[663,771,1339,1365]" collectingDate="2016-02-24" collectionCode="MUSM" collectorName="Grados" country="Peru" elevation="231" isTailEnumeration="true" location="Albergue Refugio" pageId="4" pageNumber="271" specimenCount="1" stateProvince="Amazonas" typeStatus="paratype">
(
<collectionCode id="ED10DDEEF569FFCDCAFE3C2AFD48FA8B" box="[671,763,1339,1365]" pageId="4" pageNumber="271">MUSM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31B16A0F569FFCDC8A63C4FFB30FA3B" pageId="4" pageNumber="271" type="diagnosis">
<paragraph id="8BBE452BF569FFCDC8A63C4FFB30FA3B" blockId="4.[151,1437,1230,2013]" pageId="4" pageNumber="271">
<emphasis id="B9759939F569FFCDC8A63C4FFEF1FAA6" bold="true" box="[199,322,1374,1400]" pageId="4" pageNumber="271">Diagnosis.</emphasis>
<taxonomicName id="4C013EA8F569FFCDC92D3C4EFE4BFAA7" authorityName="Inga &amp; Alvarado" authorityYear="2024" box="[332,504,1375,1401]" class="Insecta" family="Ichneumonidae" genus="Polyphrix" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="271" phylum="Arthropoda" rank="species" species="tullu" status="sp. nov.">
<emphasis id="B9759939F569FFCDC92D3C4EFE4BFAA7" box="[332,504,1375,1401]" italics="true" pageId="4" pageNumber="271">Polyphrix tullu</emphasis>
</taxonomicName>
<emphasis id="B9759939F569FFCDCA633C4EFDECFAA7" bold="true" box="[514,607,1375,1401]" pageId="4" pageNumber="271">
<taxonomicNameLabel id="A2462442F569FFCDCA633C4EFDECFAA7" box="[514,607,1375,1401]" pageId="4" pageNumber="271" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
may be distinguished from all other species by the combination of the following characters: pronotum and mesoscutum mostly black, metasomal tergites 28 black with apical white stripe and hind femur orange. It is quite similar to
<taxonomicName id="4C013EA8F569FFCDCAB43CB6FCF7FA1F" authorityName="Tedesco &amp; Santos" authorityYear="2009" box="[725,836,1447,1473]" class="Insecta" family="Ichneumonidae" genus="Polyphrix" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="271" phylum="Arthropoda" rank="species" species="stellata">
<emphasis id="B9759939F569FFCDCAB43CB6FCF7FA1F" box="[725,836,1447,1473]" italics="true" pageId="4" pageNumber="271">P. stellata</emphasis>
</taxonomicName>
but differs in having the metasoma with tergites 28 black with apices white (vs. orange), and pronotum with upper margin black (vs. yellow).
</paragraph>
</subSubSection>
<subSubSection id="C31B16A0F569FFCFC8A63CFFFE36FB1B" lastPageId="6" lastPageNumber="273" pageId="4" pageNumber="271" type="description">
<paragraph id="8BBE452BF569FFCDC8A63CFFFD98F9D7" blockId="4.[151,1437,1230,2013]" box="[199,555,1518,1545]" pageId="4" pageNumber="271">
<emphasis id="B9759939F569FFCDC8A63CFFFD98F9D7" bold="true" box="[199,555,1518,1545]" pageId="4" pageNumber="271">
Description. Female
<typeStatus id="54BAFB89F569FFCDC9DF3CFEFD94F9D7" box="[446,551,1519,1545]" pageId="4" pageNumber="271" type="holotype">holotype</typeStatus>
.
</emphasis>
</paragraph>
<paragraph id="8BBE452BF569FFCDC8A63F02FCA0F963" blockId="4.[151,1437,1230,2013]" pageId="4" pageNumber="271">
Fore wing
<quantity id="4CF9E8CEF569FFCDC9213F03FE2AF9F3" box="[320,409,1554,1581]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="4" pageNumber="271" unit="mm" value="9.5">9.5 mm</quantity>
. Body entirely shiny, scarcely punctate and very sparsely pilose.
<emphasis id="B9759939F569FFCDCC103F02FB1DF9F3" box="[1137,1198,1555,1581]" italics="true" pageId="4" pageNumber="271">Head</emphasis>
(
<figureCitation id="133A59AEF569FFCDCCDC3F02FA82F9F3" box="[1213,1329,1555,1581]" captionStart="FIGURE 3" captionStartId="5.[152,255,1792,1816]" captionTargetBox="[151,1435,181,1768]" captionTargetId="figure-26@5.[151,1435,181,1768]" captionTargetPageId="5" captionText="FIGURE 3. Polyphrix tullu sp. nov., female, holotype. A. habitus, lateral view; B. head, front view; C. pronotum, lateral view; D. mesosoma and metasoma, dorsal view; E. wings; F. ovipositor tip, lateral view. Scale bar = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.13618452" httpUri="https://zenodo.org/record/13618452/files/figure.png" pageId="4" pageNumber="271">Fig. 3B,D</figureCitation>
): antenna with 32 flagellomeres; clypeus scarcely punctate, CWH 1.56, CWW 1,22 apex slightly convex; mandible and palpi sparsely pilose; mandible moderately long, MLW 2.0, MWW 0.58, ventral tooth slightly longer and more robust than dorsal tooth; occipital carina moderately high, sharp, uniformly curved, ending far from hypostomal carina, at distance as large as basal width of mandible; MSM 0.67.
</paragraph>
<paragraph id="8BBE452BF569FFCFC8A63FD6FC96FE57" blockId="4.[151,1437,1230,2013]" lastBlockId="6.[151,1437,151,1293]" lastPageId="6" lastPageNumber="273" pageId="4" pageNumber="271">
<taxonomicName id="4C013EA8F569FFCDC8A63FD6FE8CF93E" box="[199,319,1735,1760]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="271" phylum="Arthropoda" rank="genus">
<emphasis id="B9759939F569FFCDC8A63FD6FE8CF93E" box="[199,319,1735,1760]" italics="true" pageId="4" pageNumber="271">Mesosoma</emphasis>
</taxonomicName>
(
<figureCitation id="133A59AEF569FFCDC9353FD6FDBAF93F" box="[340,521,1735,1761]" captionStart="FIGURE 3" captionStartId="5.[152,255,1792,1816]" captionTargetBox="[151,1435,181,1768]" captionTargetId="figure-26@5.[151,1435,181,1768]" captionTargetPageId="5" captionText="FIGURE 3. Polyphrix tullu sp. nov., female, holotype. A. habitus, lateral view; B. head, front view; C. pronotum, lateral view; D. mesosoma and metasoma, dorsal view; E. wings; F. ovipositor tip, lateral view. Scale bar = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.13618452" httpUri="https://zenodo.org/record/13618452/files/figure.png" pageId="4" pageNumber="271">Figs 3A, 3C,D</figureCitation>
): pronotum with very weak and short wrinkles at posterior margin of collar ventrally; dorsal margin regular, neither swollen nor prominent; mesoscutum subcircular; notaulus almost absent, traceable only in tangent anterior view; lateral carina of scutellum distinct only on anterior 0.2; epicnemial carina restricted to ventral 0.70 of mesopleuron, almost straight, mesopleuron dorsad of carina giving rise to short and very weak transverse strigulation; mesopleural fovea shallow; mesopleural suture straight, narrow and crenulate; median portion of postpectal carina very weak, short and slightly arched forward; metapleuron smooth, shiny, with sparse hairs; justacoxal carina present only as very short subvertical ridge; transverse furrow at base of propodeum centrally 0.7 as long as distance anterior transverse carina to anterior margin of propodeum, measured centrally.
<emphasis id="B9759939F568FFCCC8F63E9EFF7FF876" box="[151,204,1935,1960]" italics="true" pageId="5" pageNumber="272">Legs</emphasis>
: hind pre-apical tarsomere slightly bilobed; FLW 7.50. Propodeum anterior margin centrally concave; SWL 2.33; anterior transverse carina straight, area behind it with 12 or 13 distinct straight transverse wrinkles, either complete or incomplete, posterior ones stronger and sharper.
<emphasis id="B9759939F568FFCCCB243EC6FC3AF82E" box="[837,905,2007,2032]" italics="true" pageId="5" pageNumber="272">Wings</emphasis>
(
<figureCitation id="133A59AEF568FFCCCBF83EC7FBA3F82E" box="[921,1040,2006,2032]" captionStart="FIGURE 3" captionStartId="5.[152,255,1792,1816]" captionTargetBox="[151,1435,181,1768]" captionTargetId="figure-26@5.[151,1435,181,1768]" captionTargetPageId="5" captionText="FIGURE 3. Polyphrix tullu sp. nov., female, holotype. A. habitus, lateral view; B. head, front view; C. pronotum, lateral view; D. mesosoma and metasoma, dorsal view; E. wings; F. ovipositor tip, lateral view. Scale bar = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.13618452" httpUri="https://zenodo.org/record/13618452/files/figure.png" pageId="5" pageNumber="272">Figs 3A,E</figureCitation>
): fore wing vein 1-Rs+M distinctly sinuous, without bulla; crossvein 1m-cu uniformly arched, slightly shorter than 1-Rs+M; vein 1M+Rs anteriorly straight, posteriorly arched; crossvein 1cu-a straight, forming straight angle with M+Cu, ending basad of vein 1M+Rs by about 0.2 its own length; crossvein 2cu-a 0.24 as long as vein 2-Cu; cell 1+2Rs (areolet) of moderate size, APH 0.20, transversely rectangular, AWH 1.50; crossvein 3r-m spectral, distinctly longer than 2r-m; hind wing vein M+Cu forming straight angle with vein M; HW1C 1.14; vein 2-Rs entirely tubular; crossvein 1r-m with bulla at ventral 0.3; veins 1-Rs and 2-Rs distinctly angled; vein Cub straight, reaching about 0.8 of distance to wing margin; vein 2-1A reaching about 0.85 of distance to wing margin.
</paragraph>
<caption id="DF7E15A3F568FFCCC8F93E11FB20F8E2" ID-DOI="http://doi.org/10.5281/zenodo.13618452" ID-Zenodo-Dep="13618452" httpUri="https://zenodo.org/record/13618452/files/figure.png" pageId="5" pageNumber="272" startId="5.[152,255,1792,1816]" targetBox="[151,1435,181,1768]" targetPageId="5" targetType="figure">
<paragraph id="8BBE452BF568FFCCC8F93E11FB20F8E2" blockId="5.[151,1436,1792,1852]" pageId="5" pageNumber="272">
<emphasis id="B9759939F568FFCCC8F93E11FDBCF8C6" bold="true" box="[152,527,1792,1816]" pageId="5" pageNumber="272">
FIGURE 3.
<taxonomicName id="4C013EA8F568FFCCC97C3E11FE09F8C6" authorityName="Inga &amp; Alvarado" authorityYear="2024" box="[285,442,1792,1816]" class="Insecta" family="Ichneumonidae" genus="Polyphrix" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="272" phylum="Arthropoda" rank="species" species="tullu" status="sp. nov.">
<emphasis id="B9759939F568FFCCC97C3E11FE09F8C6" bold="true" box="[285,442,1792,1816]" italics="true" pageId="5" pageNumber="272">Polyphrix tullu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2462442F568FFCCC9A13E11FDBCF8C6" box="[448,527,1792,1816]" pageId="5" pageNumber="272" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female, holotype.
<emphasis id="B9759939F568FFCCCAB03E11FD59F8C6" bold="true" box="[721,746,1792,1816]" pageId="5" pageNumber="272">A.</emphasis>
habitus, lateral view;
<emphasis id="B9759939F568FFCCCBA73E11FC6EF8C6" bold="true" box="[966,989,1792,1816]" pageId="5" pageNumber="272">B.</emphasis>
head, front view;
<emphasis id="B9759939F568FFCCCCF23E11FB18F8C6" bold="true" box="[1171,1195,1792,1816]" pageId="5" pageNumber="272">C.</emphasis>
pronotum, lateral view;
<emphasis id="B9759939F568FFCCC8F63E35FF03F8E2" bold="true" box="[151,176,1828,1852]" pageId="5" pageNumber="272">D.</emphasis>
mesosoma and metasoma, dorsal view;
<emphasis id="B9759939F568FFCCCA283E35FDECF8E2" bold="true" box="[585,607,1828,1852]" pageId="5" pageNumber="272">E.</emphasis>
wings;
<emphasis id="B9759939F568FFCCCAD13E35FD70F8E2" bold="true" box="[688,707,1828,1852]" pageId="5" pageNumber="272">F.</emphasis>
ovipositor tip, lateral view. Scale bar = 1 mm
</paragraph>
</caption>
<paragraph id="8BBE452BF56BFFCFC8A63882FD71FDC7" blockId="6.[151,1437,151,1293]" pageId="6" pageNumber="273">
<emphasis id="B9759939F56BFFCFC8A63882FE8FFE72" box="[199,316,403,428]" italics="true" pageId="6" pageNumber="273">Metasoma</emphasis>
(
<figureCitation id="133A59AEF56BFFCFC92C3882FE53FE73" box="[333,480,403,429]" captionStart="FIGURE 3" captionStartId="5.[152,255,1792,1816]" captionTargetBox="[151,1435,181,1768]" captionTargetId="figure-26@5.[151,1435,181,1768]" captionTargetPageId="5" captionText="FIGURE 3. Polyphrix tullu sp. nov., female, holotype. A. habitus, lateral view; B. head, front view; C. pronotum, lateral view; D. mesosoma and metasoma, dorsal view; E. wings; F. ovipositor tip, lateral view. Scale bar = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.13618452" httpUri="https://zenodo.org/record/13618452/files/figure.png" pageId="6" pageNumber="273">Figs 3A,D,F</figureCitation>
): first tergite elongate, slender, T1LW 3.26, T1WW 2.09; spiracle at center, weakly protuberant; T2LW 1.10, T2WW 1.46; tergites 38 smooth, shiny, pilose; OST 1.14; ovipositor moderately slender, straight, scarcely punctate, laterally strongly compressed; dorsal valve with nodus and notch, ventral valve with eight apical teeth; apex moderately long, pointed.
</paragraph>
<paragraph id="8BBE452BF56BFFCFC8A63B32FE22FC7B" blockId="6.[151,1437,151,1293]" pageId="6" pageNumber="273">
<emphasis id="B9759939F56BFFCFC8A63B32FEBBFDE3" box="[199,264,547,573]" italics="true" pageId="6" pageNumber="273">Color</emphasis>
(
<figureCitation id="133A59AEF56BFFCFC9783B32FEECFDE3" box="[281,351,547,573]" captionStart="FIGURE 3" captionStartId="5.[152,255,1792,1816]" captionTargetBox="[151,1435,181,1768]" captionTargetId="figure-26@5.[151,1435,181,1768]" captionTargetPageId="5" captionText="FIGURE 3. Polyphrix tullu sp. nov., female, holotype. A. habitus, lateral view; B. head, front view; C. pronotum, lateral view; D. mesosoma and metasoma, dorsal view; E. wings; F. ovipositor tip, lateral view. Scale bar = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.13618452" httpUri="https://zenodo.org/record/13618452/files/figure.png" pageId="6" pageNumber="273">Fig. 3</figureCitation>
). Head mostly black, except for flagellomeres T719 white, apically flagellomeres dark brown; supra-clypeal area, clypeus, mouth parts, base of mandible, malar space and para-ocular stripe pale yellow.
<taxonomicName id="4C013EA8F56BFFCFCD423B56FA2FFDBF" box="[1315,1436,583,609]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="271" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
predominantly orange; pronotum black, except for anterior margin, pale yellow; mesoscutum black, centrally with large W-shaped pale-yellow mark; scutellum predominantly pale yellow, ventrally brown; tegula anteriorly pale yellow, posteriorly brown; subalar prominence pale yellow; fore leg predominantly orange, tibia and tarsomere 14 pale yellow, tarsomere 5 brown; mid leg predominantly orange, tarsomere 45 brown; hind leg with coxa and tibia orange, trochanter, trochantellus and femur dark orange, basitarsus pale yellow, tarsomeres 24 off-white tarsomere 5 brown. Metasoma: Tergites usually black; T1 basally orange to apically black, apical stripe pale yellow; T2 basally (at the center) dark orange; T27 basally black with apical and lateral stripes pale yellow; T8 basally black with lateral stripes pale yellow; S13 pale yellow with small black stripes on the sides; ovipositor dark red, sheaths dark brown; wings hyaline.
</paragraph>
<paragraph id="8BBE452BF56BFFCFC8A63ABEFD1AFB7F" blockId="6.[151,1437,151,1293]" pageId="6" pageNumber="273">
<emphasis id="B9759939F56BFFCFC8A63ABEFDAEFC16" box="[199,541,943,969]" italics="true" pageId="6" pageNumber="273">
Variation of female
<typeStatus id="54BAFB89F56BFFCFC9C63ABEFDAAFC16" box="[423,537,943,968]" pageId="6" pageNumber="273" type="paratype">paratypes</typeStatus>
.
</emphasis>
Fore wing length
<quantity id="4CF9E8CEF56BFFCFCA8D3ABEFC36FC17" box="[748,901,943,969]" metricMagnitude="-3" metricUnit="m" metricValue="9.55" metricValueMax="10.2" metricValueMin="8.9" pageId="6" pageNumber="273" unit="mm" value="9.55" valueMax="10.2" valueMin="8.9">8.910.2 mm</quantity>
. They differ from the
<typeStatus id="54BAFB89F56BFFCFCC1F3ABEFB53FC17" box="[1150,1248,943,969]" pageId="6" pageNumber="273" type="holotype">holotype</typeStatus>
in the following features: head with antenna with 33 flagellomeres, CWH 2.002.36, CWW 1.181.26, MLW 1.922.10, MWW 0.540.60, MSM 0.50;
<taxonomicName id="4C013EA8F56BFFCFC9C23AE6FDAFFBCF" box="[419,540,1015,1041]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="271" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
with legs FLW 7.407.44, SWL 1.402.00, area behind it with 8 or 9 distinct straight transverse wrinkles; fore wing APH 0.150.23, AWH 1.502.13; hind wing HW1C 0.891.15; metasoma with T1LW 3.644.33, T1WW 1.501.57, T2LW 1.652.60, T2WW 1.561.85, OST 1.59. Within the coloration varies in having the pronotum with anterior margin pale yellow, solely ventrally (not along the entire margin); one individual with trochantellus and femur brown.
</paragraph>
<paragraph id="8BBE452BF56BFFCFC8A63DBAFE36FB1B" blockId="6.[151,1437,151,1293]" box="[199,389,1195,1221]" pageId="6" pageNumber="273">
<emphasis id="B9759939F56BFFCFC8A63DBAFEBBFB1B" box="[199,264,1195,1221]" italics="true" pageId="6" pageNumber="273">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C31B16A0F56BFFCFC8A63DDFFCFCFB37" box="[199,847,1230,1257]" pageId="6" pageNumber="273" type="distribution">
<paragraph id="8BBE452BF56BFFCFC8A63DDFFCFCFB37" blockId="6.[151,1437,151,1293]" box="[199,847,1230,1257]" pageId="6" pageNumber="273">
<emphasis id="B9759939F56BFFCFC8A63DDFFED2FB36" bold="true" box="[199,353,1230,1256]" pageId="6" pageNumber="273">Distribution.</emphasis>
<collectingCountry id="F31605BBF56BFFCFC9063DDEFE2EFB37" box="[359,413,1231,1257]" name="Peru" pageId="6" pageNumber="273">Peru</collectingCountry>
,
<collectingRegion id="49C58BC9F56BFFCFC9C83DDEFDFCFB37" box="[425,591,1231,1257]" country="Peru" name="Madre de Dios" pageId="6" pageNumber="273">Madre de Dios</collectingRegion>
Department (
<figureCitation id="133A59AEF56BFFCFCA8B3DDEFCF0FB37" box="[746,835,1231,1257]" captionStart="FIGURE 4" captionStartId="7.[152,255,666,690]" captionTargetBox="[151,1436,181,643]" captionTargetId="figure-26@7.[151,1436,181,643]" captionTargetPageId="7" captionText="FIGURE 4. Geographic records of Polyphrix. A. Polyphrix stellata; B. Polyphrix varians. C. Polyphrix misa sp. nov. and Polyphrix tullu sp. nov. Yellow circles represent new records and red represent previous records." figureDoi="http://doi.org/10.5281/zenodo.13618454" httpUri="https://zenodo.org/record/13618454/files/figure.png" pageId="6" pageNumber="273">Fig. 4C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31B16A0F56BFFCFC8A63DE2FB33FAD3" box="[199,1152,1267,1293]" pageId="6" pageNumber="273" type="etymology">
<paragraph id="8BBE452BF56BFFCFC8A63DE2FB33FAD3" blockId="6.[151,1437,151,1293]" box="[199,1152,1267,1293]" pageId="6" pageNumber="273">
<emphasis id="B9759939F56BFFCFC8A63DE2FEFDFAD3" bold="true" box="[199,334,1267,1293]" pageId="6" pageNumber="273">Etymology.</emphasis>
The specific epithet
<emphasis id="B9759939F56BFFCFCA233DE2FDCCFAD3" box="[578,639,1267,1293]" italics="true" pageId="6" pageNumber="273">tullu</emphasis>
means ʻthinʼ in Quechua, noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,185 @@
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<paragraph id="8BBE452BF56AFFCEC8F63A34FF43FCBC" blockId="7.[151,1437,804,866]" pageId="7" pageNumber="274">
<heading id="D0F6F247F56AFFCEC8F63A34FF43FCBC" bold="true" centered="true" fontSize="11" level="1" pageId="7" pageNumber="274" reason="1">
<emphasis id="B9759939F56AFFCEC8F63A34FF43FCBC" bold="true" pageId="7" pageNumber="274">
Key to the species of
<taxonomicName id="4C013EA8F56AFFCEC9C83A35FD34FCE1" ID-CoL="63MYZ" ID-ENA="540460" authority="Townes" authorityName="Townes" authorityYear="1970" box="[425,647,804,831]" class="Insecta" family="Ichneumonidae" genus="Polyphrix" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="274" phylum="Arthropoda" rank="genus">
<emphasis id="B9759939F56AFFCEC9C83A35FD93FCE1" bold="true" box="[425,544,804,831]" italics="true" pageId="7" pageNumber="274">Polyphrix</emphasis>
Townes
</taxonomicName>
(modified from
<bibRefCitation id="EF9038DAF56AFFCECB3A3A34FB0CFCE0" author="Nogueira, L. K. &amp; Aguiar, A. P." box="[859,1215,804,831]" pageId="7" pageNumber="274" pagination="25 - 35" refId="ref5287" refString="Nogueira, L. K. &amp; Aguiar, A. P. (2005) Generic definition, key, and two new species of Polyphrix Townes (Hymenoptera: Ichneumonidae: Cryptinae) from the Atlantic Forest. Zootaxa, 1010 (1), 25 - 35. https: // doi. org / 10.11646 / zootaxa. 1010.1.3" type="journal article" year="2005">Nogueira and Aguiar (2005)</bibRefCitation>
and
<bibRefCitation id="EF9038DAF56AFFCECD633A34FF43FCBC" author="Santos, B. F. &amp; Aguiar, A. P. &amp; Tedesco, A. M." pageId="7" pageNumber="274" pagination="29 - 44" refId="ref5455" refString="Santos, B. F., Aguiar, A. P. &amp; Tedesco, A. M. (2009) Phylogenetic revision and the origin of Polyphrix Townes (Hymenoptera, Ichneumonidae, Cryptinae), with description of a new species. Zootaxa, 2214 (1), 29 - 44. https: // doi. org / 10.11646 / zootaxa. 2214.1.2" type="journal article" year="2009">
Santos
<emphasis id="B9759939F56AFFCECD3E3A34FA2EFCE1" bold="true" box="[1375,1437,804,831]" italics="true" pageId="7" pageNumber="274">et al.</emphasis>
(2009))
</bibRefCitation>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31B16A0F56AFFCEC8F63A9FFA2FFA0F" pageId="7" pageNumber="274" type="key">
<keyStep id="30F55D8EF56AFFCEC8F63A9FFA2FFC25" pageId="7" pageNumber="274">
<paragraph id="8BBE452BF56AFFCEC8F63A9FFA2FFC1E" blockId="7.[151,1437,910,1489]" pageId="7" pageNumber="274">
<keyLead id="30F0E61EF56AFFCEC8F63A9FFA2FFC1E" pageId="7" pageNumber="274">1. Propodeum with one or two pairs of somewhat oval dark spots, a pair behind anterior transverse carina and with or without a pair in front of anterior transverse carina; hind trochanter, and trochantellus dark brown; mesoscutum completely black.... 2</keyLead>
</paragraph>
<paragraph id="8BBE452BF56AFFCEC8F63AD8FA2FFC25" blockId="7.[151,1437,910,1489]" pageId="7" pageNumber="274">
<keyLead id="30F0E61EF56AFFCEC8F63AD8FA2FFC25" pageId="7" pageNumber="274">- Propodeum uniformly orange, without dark spots; hind trochanter and trochantellus entirely yellowish; mesoscutum not completely black..................................................................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F55D8EF56AFFCEC8F63D12FA2FFB72" pageId="7" pageNumber="274">
<paragraph id="8BBE452BF56AFFCEC8F63D12FA2FFB8A" blockId="7.[151,1437,910,1489]" pageId="7" pageNumber="274">
<keyLead id="30F0E61EF56AFFCEC8F63D12FA2FFB8A" pageId="7" pageNumber="274">
2. Paraocular stripe complete; pronotum with strong and distinct submarginal lamellar protuberance; anterior margin of propodeum centrally distinctly concave; ovipositor slender, dorsal valve with nodus, ventral valve with nine apical teeth, apical 34 teeth weak................................................................
<taxonomicName id="4C013EA8F56AFFCECC413D2CFA2FFB8A" authority="Nogueira &amp; Aguiar" authorityName="Nogueira &amp; Aguiar" authorityYear="2005" box="[1056,1436,1085,1108]" class="Insecta" family="Ichneumonidae" genus="Polyphrix" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="274" phylum="Arthropoda" rank="species" species="cristata">
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<emphasis id="B9759939F56AFFCECC413D2CFB62FB8D" bold="true" box="[1056,1233,1085,1107]" italics="true" pageId="7" pageNumber="274">Polyphrix cristata</emphasis>
Nogueira &amp; Aguiar
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBE452BF56AFFCEC8F63D4DFA2FFB72" blockId="7.[151,1437,910,1489]" pageId="7" pageNumber="274">
<keyLead id="30F0E61EF56AFFCEC8F63D4DFA2FFB72" pageId="7" pageNumber="274">
- Paraocular stripe interrupted from supraclypeal area to posterior orbit; pronotum with very weak submarginal protuberance; anterior margin of propodeum weakly sinuous; ovipositor thick, dorsal valve without nodus, ventral valve with seven apical teeth...............................................................
<taxonomicName id="4C013EA8F56AFFCECC723D84FA2FFB72" authority="Nogueira &amp; Aguiar" authorityName="Nogueira &amp; Aguiar" authorityYear="2005" box="[1043,1436,1173,1196]" class="Insecta" family="Ichneumonidae" genus="Polyphrix" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="274" phylum="Arthropoda" rank="species" species="atlantica">
<emphasis id="B9759939F56AFFCECC723D84FA2FFB72" bold="true" box="[1043,1436,1173,1196]" pageId="7" pageNumber="274">
<emphasis id="B9759939F56AFFCECC723D84FB62FB75" bold="true" box="[1043,1233,1173,1195]" italics="true" pageId="7" pageNumber="274">Polyphrix atlantica</emphasis>
Nogueira &amp; Aguiar
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
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<paragraph id="8BBE452BF56AFFCEC8F63DA2FA2FFB16" blockId="7.[151,1437,910,1489]" box="[151,1436,1203,1225]" pageId="7" pageNumber="274">
<keyLead id="30F0E61EF56AFFCEC8F63DA2FA2FFB16" box="[151,1436,1203,1225]" pageId="7" pageNumber="274">
3. Pronotum and mesoscutum completely orange (
<figureCitation id="133A59AEF56AFFCECACD3DA2FCA1FB16" box="[684,786,1203,1225]" captionStart="FIGURE 1" captionStartId="2.[152,255,1919,1943]" captionTargetBox="[151,1435,309,1896]" captionTargetId="figure-93@2.[151,1435,309,1896]" captionTargetPageId="2" captionText="FIGURE 1. Polyphrix misa sp. nov., female, holotype. A. habitus, lateral view; B. head, front view; C. pronotum, lateral view; D. mesosoma and metasoma, dorsal view; E. wings; F. ovipositor tip, lateral view. Scale bar = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.13618448" httpUri="https://zenodo.org/record/13618448/files/figure.png" pageId="7" pageNumber="274">Figs 1A,D</figureCitation>
, &amp;
<figureCitation id="133A59AEF56AFFCECB573DA2FCD8FB16" box="[822,875,1203,1224]" captionStart="FIGURE 2" captionStartId="3.[152,255,1439,1463]" captionTargetBox="[151,1436,181,1415]" captionTargetId="figure-26@3.[151,1436,181,1415]" captionTargetPageId="3" captionText="FIGURE 2. Polyphrix misa sp. nov., male, paratype. A. habitus, lateral view; B. head, front view; C. wings; D. mesosoma and metasoma, dorsal view. Scale bar = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.13618450" httpUri="https://zenodo.org/record/13618450/files/figure.png" pageId="7" pageNumber="274">2A,D</figureCitation>
)............................................ 4
</keyLead>
</paragraph>
<paragraph id="8BBE452BF56AFFCEC8F63DC0FA2FFB38" blockId="7.[151,1437,910,1489]" box="[151,1436,1232,1254]" pageId="7" pageNumber="274">
<keyLead id="30F0E61EF56AFFCEC8F63DC0FA2FFB38" box="[151,1436,1232,1254]" pageId="7" pageNumber="274">
- Pronotum and mesoscutum mostly black, with small yellow marks (
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)................................... 5
</keyLead>
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4. Transverse furrow at base of propodeum, measured centrally, about 1.2 as wide as basal area of propodeum; metafemur entirely yellowish.......................................................................
<taxonomicName id="4C013EA8F56AFFCECCF83C1AFA2FFAFE" ID-CoL="4LDXD" authority="Townes" authorityName="Townes" authorityYear="1970" box="[1177,1436,1291,1313]" class="Insecta" family="Ichneumonidae" genus="Polyphrix" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="274" phylum="Arthropoda" rank="species" species="varians">
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<emphasis id="B9759939F56AFFCECCF83C1AFAFAFAFE" bold="true" box="[1177,1353,1291,1313]" italics="true" pageId="7" pageNumber="274">Polyphrix varians</emphasis>
Townes
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- Transverse furrow at base of propodeum, measured centrally, about 0.7 as wide as basal area of propodeum; metafemur entirely blackish (
<figureCitation id="133A59AEF56AFFCEC95E3C57FE3DFA85" box="[319,398,1350,1372]" captionStart="FIGURE 1" captionStartId="2.[152,255,1919,1943]" captionTargetBox="[151,1435,309,1896]" captionTargetId="figure-93@2.[151,1435,309,1896]" captionTargetPageId="2" captionText="FIGURE 1. Polyphrix misa sp. nov., female, holotype. A. habitus, lateral view; B. head, front view; C. pronotum, lateral view; D. mesosoma and metasoma, dorsal view; E. wings; F. ovipositor tip, lateral view. Scale bar = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.13618448" httpUri="https://zenodo.org/record/13618448/files/figure.png" pageId="7" pageNumber="274">Figs 1A</figureCitation>
,
<figureCitation id="133A59AEF56AFFCEC9F83C57FE04FA85" box="[409,439,1350,1371]" captionStart="FIGURE 2" captionStartId="3.[152,255,1439,1463]" captionTargetBox="[151,1436,181,1415]" captionTargetId="figure-26@3.[151,1436,181,1415]" captionTargetPageId="3" captionText="FIGURE 2. Polyphrix misa sp. nov., male, paratype. A. habitus, lateral view; B. head, front view; C. wings; D. mesosoma and metasoma, dorsal view. Scale bar = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.13618450" httpUri="https://zenodo.org/record/13618450/files/figure.png" pageId="7" pageNumber="274">2A</figureCitation>
)...............................................................
<emphasis id="B9759939F56AFFCECCD93C54FA2FFA85" bold="true" box="[1208,1436,1349,1371]" pageId="7" pageNumber="274">
<taxonomicName id="4C013EA8F56AFFCECCD93C54FAFCFA85" authorityName="Inga &amp; Alvarado" authorityYear="2024" box="[1208,1359,1349,1371]" class="Insecta" family="Ichneumonidae" genus="Polyphrix" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="274" phylum="Arthropoda" rank="species" species="misa" status="sp. nov.">
<emphasis id="B9759939F56AFFCECCD93C54FAFCFA85" bold="true" box="[1208,1359,1349,1371]" italics="true" pageId="7" pageNumber="274">Polyphrix misa</emphasis>
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<taxonomicNameLabel id="A2462442F56AFFCECD353C57FA2FFA85" box="[1364,1436,1350,1371]" pageId="7" pageNumber="274" rank="species">sp. nov</taxonomicNameLabel>
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<keyLead id="30F0E61EF56AFFCEC8F63C72FA2FFA48" pageId="7" pageNumber="274">
5. Metasomal tergites 28 black with apical white stripe (
<figureCitation id="133A59AEF56AFFCECA893C72FCFCFAA6" box="[744,847,1379,1401]" captionStart="FIGURE 3" captionStartId="5.[152,255,1792,1816]" captionTargetBox="[151,1435,181,1768]" captionTargetId="figure-26@5.[151,1435,181,1768]" captionTargetPageId="5" captionText="FIGURE 3. Polyphrix tullu sp. nov., female, holotype. A. habitus, lateral view; B. head, front view; C. pronotum, lateral view; D. mesosoma and metasoma, dorsal view; E. wings; F. ovipositor tip, lateral view. Scale bar = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.13618452" httpUri="https://zenodo.org/record/13618452/files/figure.png" pageId="7" pageNumber="274">Figs 3A,D</figureCitation>
); transverse furrow at base of propodeum, measured centrally, as wide as basal area of propodeum (
<figureCitation id="133A59AEF56AFFCECA253C91FD3CFA48" box="[580,655,1408,1430]" captionStart="FIGURE 3" captionStartId="5.[152,255,1792,1816]" captionTargetBox="[151,1435,181,1768]" captionTargetId="figure-26@5.[151,1435,181,1768]" captionTargetPageId="5" captionText="FIGURE 3. Polyphrix tullu sp. nov., female, holotype. A. habitus, lateral view; B. head, front view; C. pronotum, lateral view; D. mesosoma and metasoma, dorsal view; E. wings; F. ovipositor tip, lateral view. Scale bar = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.13618452" httpUri="https://zenodo.org/record/13618452/files/figure.png" pageId="7" pageNumber="274">Fig. 3D</figureCitation>
)...........................................
<taxonomicName id="4C013EA8F56AFFCECCD33C91FAFBFA48" authorityName="Inga &amp; Alvarado" authorityYear="2024" box="[1202,1352,1408,1430]" class="Insecta" family="Ichneumonidae" genus="Polyphrix" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="274" phylum="Arthropoda" rank="species" species="tullu" status="sp. nov.">Polyphrix tullu</taxonomicName>
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<emphasis id="B9759939F56AFFCECD2F3C90FA25FA48" bold="true" box="[1358,1430,1409,1430]" pageId="7" pageNumber="274">sp. nov</emphasis>
.
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<paragraph id="8BBE452BF56AFFCEC8F63C8FFA2FFA0F" blockId="7.[151,1437,910,1489]" pageId="7" pageNumber="274">
<keyLead id="30F0E61EF56AFFCEC8F63C8FFA2FFA0F" pageId="7" pageNumber="274">
- Metasomal tergites 28 entirely orange; transverse furrow at base of propodeum, measured centrally, about 0.7 as wide as basal area of propodeum.......................................................
<taxonomicName id="4C013EA8F56AFFCECC553CAAFA2FFA0F" authority="Tedesco &amp; Santos" authorityName="Tedesco &amp; Santos" authorityYear="2009" box="[1076,1436,1467,1489]" class="Insecta" family="Ichneumonidae" genus="Polyphrix" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="274" phylum="Arthropoda" rank="species" species="stellata">
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<emphasis id="B9759939F56AFFCECC553CAAFB51FA0F" bold="true" box="[1076,1250,1467,1489]" italics="true" pageId="7" pageNumber="274">Polyphrix stellata</emphasis>
Tedesco &amp; Santos
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View file

@ -0,0 +1,271 @@
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//
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:
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,
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los
<collectorName id="26F420FDF56BFFCFCC023CFEFB0FF9D7" box="[1123,1212,1519,1545]" pageId="6" pageNumber="273">Amigos</collectorName>
<collectionCode id="ED10DDEEF56BFFCFCCA33CFEFAA5F9D7" box="[1218,1302,1519,1545]" pageId="6" pageNumber="273">CICRA</collectionCode>
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14.ix.
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, 208m 12º343”/ 70°546.7” Malaise L. Huerto”
</materialsCitation>
,
<materialsCitation id="3B694F76F56BFFCFCAA13F02FDDFF98F" collectingDate="2012-10-01" collectionCode="LO" collectorName="C. Espinoza" country="Peru" elevation="190" latitude="-5.4521666" location="Alto" longLatPrecision="15" longitude="-76.30945" pageId="6" pageNumber="273" specimenCount="1" specimenCount-female="1" stateProvince="Amazonas">
<specimenCount id="9D078EA2F56BFFCFCAA13F02FD51F9F3" box="[704,738,1555,1581]" count="1" pageId="6" pageNumber="273" type="female">1♀</specimenCount>
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:
<collectionCode id="ED10DDEEF56BFFCFCB2F3F02FCC4F9F3" box="[846,887,1555,1581]" country="Sweden" name="Type Collection" pageId="6" pageNumber="273">LO</collectionCode>
.
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<collectingRegion id="49C58BC9F56BFFCFCBD93F02FB82F9F3" box="[952,1073,1555,1581]" country="Peru" name="Amazonas" pageId="6" pageNumber="273">Amazonas</collectingRegion>
,
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/
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<date id="FFBF63EBF56BFFCFC8F63F26FEB9F98F" box="[151,266,1591,1617]" pageId="6" pageNumber="273" value="2012-10-01">
<collectingDate id="EFFB9A03F56BFFCFC8F63F26FEB9F98F" box="[151,266,1591,1617]" pageId="6" pageNumber="273" value="2012-10-01">01.x.2012</collectingDate>
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.
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,
<collectorName id="26F420FDF56BFFCFC9B43F26FDD1F98F" box="[469,610,1591,1617]" pageId="6" pageNumber="273">C. Espinoza</collectorName>
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,
<materialsCitation id="3B694F76F56BFFCFCA193F26FEE2F9AB" collectingDate="2008-07-18" collectorName="C. Castillo" country="Peru" elevation="118" location="Maynas" pageId="6" pageNumber="273" specimenCount="1" specimenCount-male="1" stateProvince="Loreto">
<specimenCount id="9D078EA2F56BFFCFCA193F26FD28F98F" box="[632,667,1591,1617]" count="1" pageId="6" pageNumber="273" type="male">1♂</specimenCount>
“PE:
<collectingRegion id="49C58BC9F56BFFCFCABE3F26FC9EF98F" box="[735,813,1591,1617]" country="Peru" name="Loreto" pageId="6" pageNumber="273">Loreto</collectingRegion>
,
<location id="8EDE13F0F56BFFCFCB583F26FC21F98F" LSID="urn:lsid:plazi:treatment:03A8F43DF56BFFCFC8F63C4FFDEBF867:8EDE13F0F56BFFCFCB583F26FC21F98F" box="[825,914,1591,1617]" country="Peru" name="Maynas" pageId="6" pageNumber="273" stateProvince="Loreto">Maynas</location>
18M 532239E, 9583762N
<quantity id="4CF9E8CEF56BFFCFCCA93F26FAB9F98F" box="[1224,1290,1591,1617]" metricMagnitude="2" metricUnit="m" metricValue="1.18" pageId="6" pageNumber="273" unit="m" value="118.0">
<elevation id="002CA218F56BFFCFCCA93F26FAB9F98F" box="[1224,1290,1591,1617]" metricMagnitude="2" metricUnit="m" metricValue="1.18" pageId="6" pageNumber="273" unit="m" value="118.0">118m</elevation>
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,
<date id="FFBF63EBF56BFFCFCD773F26FA2BF98F" box="[1302,1432,1591,1617]" pageId="6" pageNumber="273" value="2008-07-18">
<collectingDate id="EFFB9A03F56BFFCFCD773F26FA2BF98F" box="[1302,1432,1591,1617]" pageId="6" pageNumber="273" value="2008-07-18">18.vii.2008</collectingDate>
</date>
,
<collectorName id="26F420FDF56BFFCFC8F63F4AFEA7F9AB" box="[151,276,1627,1653]" pageId="6" pageNumber="273">C. Castillo</collectorName>
”, and
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<materialsCitation id="3B694F76F56BFFCFC9383F4AFAC6F9AB" box="[345,1397,1626,1653]" collectingDate="2008-07-18" collectionCode="MUSM" collectorName="C. Castillo." country="Peru" elevation="118" location="Maynas" pageId="6" pageNumber="273" specimenCount="1" specimenCount-male="1" stateProvince="Loreto">
<specimenCount id="9D078EA2F56BFFCFC9383F4AFECFF9AB" box="[345,380,1627,1653]" count="1" pageId="6" pageNumber="273" type="male">1♂</specimenCount>
“PE:
<collectingRegion id="49C58BC9F56BFFCFC9DE3F4AFDBEF9AB" box="[447,525,1627,1653]" country="Peru" name="Loreto" pageId="6" pageNumber="273">Loreto</collectingRegion>
,
<location id="8EDE13F0F56BFFCFCA763F4AFDC3F9AB" LSID="urn:lsid:plazi:treatment:03A8F43DF56BFFCFC8F63C4FFDEBF867:8EDE13F0F56BFFCFCA763F4AFDC3F9AB" box="[535,624,1627,1653]" country="Peru" name="Maynas" pageId="6" pageNumber="273" stateProvince="Loreto">Maynas</location>
18M 532239E,9583762N
<quantity id="4CF9E8CEF56BFFCFCBFD3F4AFC6DF9AB" box="[924,990,1627,1653]" metricMagnitude="2" metricUnit="m" metricValue="1.18" pageId="6" pageNumber="273" unit="m" value="118.0">
<elevation id="002CA218F56BFFCFCBFD3F4AFC6DF9AB" box="[924,990,1627,1653]" metricMagnitude="2" metricUnit="m" metricValue="1.18" pageId="6" pageNumber="273" unit="m" value="118.0">118m</elevation>
</quantity>
,
<date id="FFBF63EBF56BFFCFCB883F4AFBD8F9AB" box="[1001,1131,1627,1653]" pageId="6" pageNumber="273" value="2008-07-18">
<collectingDate id="EFFB9A03F56BFFCFCB883F4AFBD8F9AB" box="[1001,1131,1627,1653]" pageId="6" pageNumber="273" value="2008-07-18">18.vii.2008</collectingDate>
</date>
,
<collectorName id="26F420FDF56BFFCFCC163F4AFB4BF9AB" box="[1143,1272,1627,1653]" pageId="6" pageNumber="273">C. Castillo.</collectorName>
” (
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)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31B16A0F56BFFCFC8A63F6FFC55F8AF" pageId="6" pageNumber="273" type="discussion">
<paragraph id="8BBE452BF56BFFCFC8A63F6FFC55F8AF" blockId="6.[151,1437,1482,1977]" pageId="6" pageNumber="273">
<emphasis id="B9759939F56BFFCFC8A63F6FFEE1F946" bold="true" box="[199,338,1662,1688]" pageId="6" pageNumber="273">Comments:</emphasis>
<bibRefCitation id="EF9038DAF56BFFCFC9383F6EFDB2F947" author="Townes, H. K." box="[345,513,1663,1689]" pageId="6" pageNumber="273" pagination="1 - 537" refId="ref5561" refString="Townes, H. K. (1970) The genera of Ichneumonidae, Part 2. Memoirs of the American Entomological Institute, 12, 1 - 537." type="journal article" year="1970">Townes (1970)</bibRefCitation>
mentioned that
<taxonomicName id="4C013EA8F56BFFCFCAD73F6EFC95F946" authorityName="Townes" authorityYear="1970" box="[694,806,1663,1688]" class="Insecta" family="Ichneumonidae" genus="Polyphrix" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="273" phylum="Arthropoda" rank="species" species="varians">
<emphasis id="B9759939F56BFFCFCAD73F6EFC95F946" box="[694,806,1663,1688]" italics="true" pageId="6" pageNumber="273">P. varians</emphasis>
</taxonomicName>
, the only known species at that time and
<typeStatus id="54BAFB89F56BFFCFCC973F6EFA95F947" box="[1270,1318,1663,1689]" pageId="6" pageNumber="273">type</typeStatus>
species of the genus, occurred in
<collectingCountry id="F31605BBF56BFFCFC9F03FB2FE77F963" box="[401,452,1699,1725]" name="Peru" pageId="6" pageNumber="273">Peru</collectingCountry>
in the genus description but the
<typeStatus id="54BAFB89F56BFFCFCB483FB2FCEAF963" box="[809,857,1699,1725]" pageId="6" pageNumber="273">type</typeStatus>
material used to described the species was collected in
<collectingCountry id="F31605BBF56BFFCFC8D43FD6FF4AF93F" box="[181,249,1735,1761]" name="Brazil" pageId="6" pageNumber="273">Brazil</collectingCountry>
and no specimen collected from
<collectingCountry id="F31605BBF56BFFCFCA113FD6FD10F93F" box="[624,675,1735,1761]" name="Peru" pageId="6" pageNumber="273">Peru</collectingCountry>
was included. Later, in the list provided by
<bibRefCitation id="EF9038DAF56BFFCFCCF43FD6FAFEF93F" author="Carrasco, Z. F." box="[1173,1357,1735,1761]" pageId="6" pageNumber="273" pagination="324 - 332" refId="ref5260" refString="Carrasco, Z. F. (1972) Catalogo de la familia Ichneumonidae peruanos. Revista Peruana de Entomologia, 15, 324 - 332." type="journal article" year="1972">Carrasco (1972)</bibRefCitation>
, it was mentioned as occurring in
<collectingCountry id="F31605BBF56BFFCFC9A63FFAFE49F8DB" box="[455,506,1771,1797]" name="Peru" pageId="6" pageNumber="273">Peru</collectingCountry>
but acknowledging the lack of data.
<bibRefCitation id="EF9038DAF56BFFCFCBC33FFAFB4EF8DB" author="Rodriguez-Berrio, A. &amp; Bordera, S. &amp; Saaksjarvi, I. E." box="[930,1277,1771,1797]" pageId="6" pageNumber="273" pagination="1 - 44" refId="ref5350" refString="Rodriguez-Berrio, A., Bordera, S. &amp; Saaksjarvi, I. E. (2009) Checklist of Peruvian Ichneumonidae (Insecta, Hymenoptera). Zootaxa, 2303 (1), 1 - 44. https: // doi. org / 10.11646 / zootaxa. 2303.1.1" type="journal article" year="2009">
Rodriguez-Berrio
<emphasis id="B9759939F56BFFCFCC113FFAFB19F8DB" box="[1136,1194,1771,1797]" italics="true" pageId="6" pageNumber="273">et al.</emphasis>
(2009)
</bibRefCitation>
considered in their catalogue of Peruvian
<taxonomicName id="4C013EA8F56BFFCFC9B23E1EFD33F8F7" authorityName="Latreille" authorityYear="1802" box="[467,640,1807,1833]" class="Insecta" family="Ichneumonidae" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="273" phylum="Arthropoda" rank="family">Ichneumonidae</taxonomicName>
as present in
<collectingCountry id="F31605BBF56BFFCFCB413E1EFCE5F8F7" box="[800,854,1807,1833]" name="Peru" pageId="6" pageNumber="273">Peru</collectingCountry>
, ignoring the fact that
<bibRefCitation id="EF9038DAF56BFFCFCC3A3E1EFA2FF8F7" author="Nogueira, L. K. &amp; Aguiar, A. P." box="[1115,1436,1806,1833]" pageId="6" pageNumber="273" pagination="25 - 35" refId="ref5287" refString="Nogueira, L. K. &amp; Aguiar, A. P. (2005) Generic definition, key, and two new species of Polyphrix Townes (Hymenoptera: Ichneumonidae: Cryptinae) from the Atlantic Forest. Zootaxa, 1010 (1), 25 - 35. https: // doi. org / 10.11646 / zootaxa. 1010.1.3" type="journal article" year="2005">Nogueira and Aguiar (2005)</bibRefCitation>
refuted the presence of this species in
<collectingCountry id="F31605BBF56BFFCFCA2C3E22FD37F893" box="[589,644,1843,1869]" name="Peru" pageId="6" pageNumber="273">Peru</collectingCountry>
. Here we found several specimens of
<taxonomicName id="4C013EA8F56BFFCFCC593E22FB1AF892" authorityName="Townes" authorityYear="1970" box="[1080,1193,1843,1868]" class="Insecta" family="Ichneumonidae" genus="Polyphrix" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="273" phylum="Arthropoda" rank="species" species="varians">
<emphasis id="B9759939F56BFFCFCC593E22FB1AF892" box="[1080,1193,1843,1868]" italics="true" pageId="6" pageNumber="273">P. varians</emphasis>
</taxonomicName>
widely distributed in Peruvian Amazon Basin; all specimens were collected between
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112
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</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C31B16A0F56BFFCFC8A63E6BFDEBF867" pageId="6" pageNumber="273" type="distribution">
<paragraph id="8BBE452BF56BFFCFC8A63E6BFDEBF867" blockId="6.[151,1437,1482,1977]" pageId="6" pageNumber="273">
<emphasis id="B9759939F56BFFCFC8A63E6BFE72F84B" bold="true" box="[199,449,1914,1941]" pageId="6" pageNumber="273">Distribution records.</emphasis>
<collectingCountry id="F31605BBF56BFFCFC9A63E6AFDBDF84B" box="[455,526,1915,1941]" name="Brazil" pageId="6" pageNumber="273">Brazil</collectingCountry>
,
<collectingCountry id="F31605BBF56BFFCFCA783E6AFDC3F84B" box="[537,624,1915,1941]" name="Guyana" pageId="6" pageNumber="273">Guyana</collectingCountry>
(
<bibRefCitation id="EF9038DAF56BFFCFCA1E3E6AFCFEF84B" author="Santos, B. F. &amp; Aguiar, A. P. &amp; Tedesco, A. M." box="[639,845,1915,1941]" pageId="6" pageNumber="273" pagination="29 - 44" refId="ref5455" refString="Santos, B. F., Aguiar, A. P. &amp; Tedesco, A. M. (2009) Phylogenetic revision and the origin of Polyphrix Townes (Hymenoptera, Ichneumonidae, Cryptinae), with description of a new species. Zootaxa, 2214 (1), 29 - 44. https: // doi. org / 10.11646 / zootaxa. 2214.1.2" type="journal article" year="2009">
Santos
<emphasis id="B9759939F56BFFCFCAB03E6AFCBAF84B" box="[721,777,1915,1941]" italics="true" pageId="6" pageNumber="273">et al.</emphasis>
2009
</bibRefCitation>
). In
<collectingCountry id="F31605BBF56BFFCFCB1D3E6AFC1CF84B" box="[892,943,1915,1941]" name="Peru" pageId="6" pageNumber="273">Peru</collectingCountry>
it is known to occur in the
<collectingRegion id="49C58BC9F56BFFCFCCBE3E6AFA9BF84B" box="[1247,1320,1915,1941]" country="Peru" name="Cusco" pageId="6" pageNumber="273">Cusco</collectingRegion>
,
<collectingRegion id="49C58BC9F56BFFCFCD533E6AFF7FF867" country="Peru" name="Madre de Dios" pageId="6" pageNumber="273">Madre de Dios</collectingRegion>
and
<collectingRegion id="49C58BC9F56BFFCFC9623E8EFEFDF867" box="[259,334,1951,1977]" country="Peru" name="Loreto" pageId="6" pageNumber="273">Loreto</collectingRegion>
Departments (
<figureCitation id="133A59AEF56BFFCFC9923E8EFDF8F867" box="[499,587,1951,1977]" captionStart="FIGURE 4" captionStartId="7.[152,255,666,690]" captionTargetBox="[151,1436,181,643]" captionTargetId="figure-26@7.[151,1436,181,643]" captionTargetPageId="7" captionText="FIGURE 4. Geographic records of Polyphrix. A. Polyphrix stellata; B. Polyphrix varians. C. Polyphrix misa sp. nov. and Polyphrix tullu sp. nov. Yellow circles represent new records and red represent previous records." figureDoi="http://doi.org/10.5281/zenodo.13618454" httpUri="https://zenodo.org/record/13618454/files/figure.png" pageId="6" pageNumber="273">Fig. 4B</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,327 @@
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<mods:title id="EF0FBFCBDEF0A9C86676DECC5D3B5BB4">Two new species of the genus Polyphrix Townes 1972 (Hymenoptera, Ichneumonidae, Cryptinae) from Peru</mods:title>
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<mods:name id="7F83C3E99FE3E51F75081BDF59FE4FAE" type="personal">
<mods:role id="00FE41F6FF619E3A1ED38A89F1BC785B">
<mods:roleTerm id="DB53D8C28143B7145EB0133859E67F75">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F38DD15B7E2ACFDEF24B5E5A4637DE95">Alvarado, Mabel</mods:namePart>
<mods:nameIdentifier id="D62770AA7B20A61F3A32E2E11BED6E93" type="ORCID">0000-0001-8135-9223</mods:nameIdentifier>
<mods:affiliation id="2AD0DBC9B8D603DDBDA4E249D490E512">Departamento de Entomología, Museo de Historia Natural, Universidad Nacional Mayor de San Marcos, Lima, Perú</mods:affiliation>
<mods:nameIdentifier id="A1AE8683012FF198BC7D7685B870A370" type="email">malvaradog@unmsm.edu.pe</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="9B64DA42647C8B8DB190BAB5F87FA30F">text</mods:typeOfResource>
<mods:relatedItem id="014283018649CBB8DFFD8ADCC16C103B" type="host">
<mods:titleInfo id="3F5836FD8FB173920657F1FCFC195216">
<mods:title id="1659DD3DDB6D018A1AB6213491D5D35B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="0785AE33F643479F82F0B5E4B4FCDCD9">
<mods:date id="710248FE9C6BC9B894A727EA60D537F6">2024</mods:date>
<mods:detail id="0D67BF24F313838848E6746FFDDE1CA4" type="pubDate">
<mods:number id="6D5A47B75B301726F76F7CE3D9E4238B">2024-08-23</mods:number>
</mods:detail>
<mods:detail id="2E7BA24CE118AEA99E001D26ED7C9260" type="volume">
<mods:number id="271131347ABD0C63B031AEEA3AD35D9E">5497</mods:number>
</mods:detail>
<mods:detail id="7A1F79CEFC7C5E9FACA88BD9EBEDA829" type="issue">
<mods:number id="8BE90900EABA6CEBBCC7230D38AE5E7C">2</mods:number>
</mods:detail>
<mods:extent id="1AF9B11114EC9E5A741D61D50761C87C" unit="page">
<mods:start id="5641B58034AF29465D2D82EB2C89920D">267</mods:start>
<mods:end id="A4E0C094A201767F6163D2872937D7F7">275</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="F8A06DBA8BE362EF423A98F3492D3BC5">
<mods:url id="F048CAF20ACF941D49FB28980C84DEAB">http://dx.doi.org/10.11646/zootaxa.5497.2.6</mods:url>
</mods:location>
<mods:classification id="C34272DAD0E085181294FF99DAAD6899">journal article</mods:classification>
<mods:identifier id="EC39C82031994AB8C15E087F16F4EE05" type="DOI">10.11646/zootaxa.5497.2.6</mods:identifier>
<mods:identifier id="CCB16A64B9964F3D5EF04883D0DE9C19" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="0FBD33D794AB6EAC023BB5837516904A" type="Zenodo-Dep">13618446</mods:identifier>
<mods:identifier id="6654EE1B8F28EE8E5E9380A315B0258F" type="ZooBank">77458D3A-B967-44B9-9DD1-A7718ED7953D</mods:identifier>
</mods:mods>
<treatment id="03A8F43DF56CFFCDC8F63BC7FB16FDC7" ID-DOI="http://doi.org/10.5281/zenodo.13618500" ID-Zenodo-Dep="13618500" LSID="urn:lsid:plazi:treatment:03A8F43DF56CFFCDC8F63BC7FB16FDC7" httpUri="http://treatment.plazi.org/id/03A8F43DF56CFFCDC8F63BC7FB16FDC7" lastPageId="4" lastPageNumber="271" pageId="1" pageNumber="268">
<subSubSection id="C31B16A0F56CFFC8C8F63BC7FE00FD2F" box="[151,435,726,753]" pageId="1" pageNumber="268" type="nomenclature">
<paragraph id="8BBE452BF56CFFC8C8F63BC7FE00FD2F" blockId="1.[151,435,726,789]" box="[151,435,726,753]" pageId="1" pageNumber="268">
<heading id="D0F6F247F56CFFC8C8F63BC7FE00FD2F" bold="true" box="[151,435,726,753]" fontSize="11" level="1" pageId="1" pageNumber="268" reason="1">
<emphasis id="B9759939F56CFFC8C8F63BC7FE00FD2F" bold="true" box="[151,435,726,753]" pageId="1" pageNumber="268">
<taxonomicName id="4C013EA8F56CFFC8C8F63BC7FEFDFD2F" authorityName="Inga &amp; Alvarado" authorityYear="2024" box="[151,334,726,753]" class="Insecta" family="Ichneumonidae" genus="Polyphrix" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="268" phylum="Arthropoda" rank="species" species="misa" status="sp. nov.">
<emphasis id="B9759939F56CFFC8C8F63BC7FEFDFD2F" bold="true" box="[151,334,726,753]" italics="true" pageId="1" pageNumber="268">Polyphrix misa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2462442F56CFFC8C9373BC6FE00FD2F" box="[342,435,727,753]" pageId="1" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31B16A0F56CFFC8C8F63BEAFE9BFCCB" box="[151,296,763,789]" pageId="1" pageNumber="268" type="description">
<paragraph id="8BBE452BF56CFFC8C8F63BEAFE9BFCCB" blockId="1.[151,435,726,789]" box="[151,296,763,789]" pageId="1" pageNumber="268">
<figureCitation id="133A59AEF56CFFC8C8F63BEAFF49FCCB" box="[151,250,763,789]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[152,255,1919,1943]" captionStartId-1="3.[152,255,1439,1463]" captionTargetBox-0="[151,1435,309,1896]" captionTargetBox-1="[151,1436,181,1415]" captionTargetId-0="figure-93@2.[151,1435,309,1896]" captionTargetId-1="figure-26@3.[151,1436,181,1415]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Polyphrix misa sp. nov., female, holotype. A. habitus, lateral view; B. head, front view; C. pronotum, lateral view; D. mesosoma and metasoma, dorsal view; E. wings; F. ovipositor tip, lateral view. Scale bar = 1 mm" captionText-1="FIGURE 2. Polyphrix misa sp. nov., male, paratype. A. habitus, lateral view; B. head, front view; C. wings; D. mesosoma and metasoma, dorsal view. Scale bar = 1 mm" figureDoi-0="http://doi.org/10.5281/zenodo.13618448" figureDoi-1="http://doi.org/10.5281/zenodo.13618450" httpUri-0="https://zenodo.org/record/13618448/files/figure.png" httpUri-1="https://zenodo.org/record/13618450/files/figure.png" pageId="1" pageNumber="268">Figs 12</figureCitation>
,
<figureCitation id="133A59AEF56CFFC8C9673BEAFE9BFCCB" box="[262,296,763,789]" captionStart="FIGURE 4" captionStartId="7.[152,255,666,690]" captionTargetBox="[151,1436,181,643]" captionTargetId="figure-26@7.[151,1436,181,643]" captionTargetPageId="7" captionText="FIGURE 4. Geographic records of Polyphrix. A. Polyphrix stellata; B. Polyphrix varians. C. Polyphrix misa sp. nov. and Polyphrix tullu sp. nov. Yellow circles represent new records and red represent previous records." figureDoi="http://doi.org/10.5281/zenodo.13618454" httpUri="https://zenodo.org/record/13618454/files/figure.png" pageId="1" pageNumber="268">4C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31B16A0F56CFFC8C8F63A53FEC0FC33" pageId="1" pageNumber="268" type="materials_examined">
<paragraph id="8BBE452BF56CFFC8C8F63A53FEC0FC33" blockId="1.[151,1437,834,2013]" pageId="1" pageNumber="268">
<emphasis id="B9759939F56CFFC8C8F63A53FE37FC82" bold="true" box="[151,388,834,860]" pageId="1" pageNumber="268">Material examined.</emphasis>
<materialsCitation id="3B694F76F56CFFC8C9EE3A52FC75FC5F" collectingDate="2008-12-12" collectionCode="LO, MUSM" collectorName="C. Castillo" country="Peru" elevation="120" location="Alto Nanay" municipality="Qda Lobillos" pageId="1" pageNumber="268" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="B9759939F56CFFC8C9EE3A52FDB6FC83" bold="true" box="[399,517,835,861]" pageId="1" pageNumber="268">
<typeStatus id="54BAFB89F56CFFC8C9EE3A52FDB3FC83" box="[399,512,835,861]" pageId="1" pageNumber="268" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="9D078EA2F56CFFC8CA713A52FD80FC83" box="[528,563,835,861]" count="1" pageId="1" pageNumber="268" type="female">1♀</specimenCount>
<collectingCountry id="F31605BBF56CFFC8CA293A52FD25FC83" box="[584,662,835,861]" name="Peru" pageId="1" pageNumber="268">PERU</collectingCountry>
:
<collectionCode id="ED10DDEEF56CFFC8CAC53A52FD7EFC83" box="[676,717,835,861]" country="Sweden" name="Type Collection" pageId="1" pageNumber="268">LO</collectionCode>
.
<location id="8EDE13F0F56CFFC8CAB83A52FCD3FC83" LSID="urn:lsid:plazi:treatment:03A8F43DF56CFFCDC8F63BC7FB16FDC7:8EDE13F0F56CFFC8CAB83A52FCD3FC83" box="[729,864,835,861]" country="Peru" municipality="Qda Lobillos" name="Alto Nanay" pageId="1" pageNumber="268">Alto Nanay</location>
,
<collectingMunicipality id="6BDADF51F56CFFC8CB0E3A52FBBBFC83" box="[879,1032,835,861]" pageId="1" pageNumber="268">Qda Lobillos</collectingMunicipality>
,
<quantity id="4CF9E8CEF56CFFC8CC763A52FBE4FC83" box="[1047,1111,835,861]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="268" unit="m" value="120.0">
<elevation id="002CA218F56CFFC8CC763A52FBE4FC83" box="[1047,1111,835,861]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="268" unit="m" value="120.0">120m</elevation>
</quantity>
18M 0563775E, 9610397N
<date id="FFBF63EBF56CFFC8C8F63A76FEAAFC5F" box="[151,281,871,897]" pageId="1" pageNumber="268" value="2008-12-12">
<collectingDate id="EFFB9A03F56CFFC8C8F63A76FEAAFC5F" box="[151,281,871,897]" pageId="1" pageNumber="268" value="2008-12-12">12.xii.2008</collectingDate>
</date>
,
<collectorName id="26F420FDF56CFFC8C9433A76FE2BFC5F" box="[290,408,871,897]" pageId="1" pageNumber="268">C. Castillo</collectorName>
//Aguajal Mixto Colecta manual 11AB” (
<collectionCode id="ED10DDEEF56CFFC8CB033A76FC0DFC5F" box="[866,958,871,897]" pageId="1" pageNumber="268">MUSM</collectionCode>
)
</materialsCitation>
.
<emphasis id="B9759939F56CFFC8CBAE3A76FBF6FC5F" bold="true" box="[975,1093,871,897]" pageId="1" pageNumber="268">
<typeStatus id="54BAFB89F56CFFC8CBAE3A76FBF3FC5F" box="[975,1088,871,897]" pageId="1" pageNumber="268" type="paratype">Paratype</typeStatus>
:
</emphasis>
<specimenCount id="9D078EA2F56CFFC8CC2B3A76FBDFFC5F" box="[1098,1132,871,897]" count="1" pageId="1" pageNumber="268" type="female">1♀</specimenCount>
and
<specimenCount id="9D078EA2F56CFFC8CCFF3A76FB72FC5F" box="[1182,1217,871,897]" count="1" pageId="1" pageNumber="268" type="male">1♂</specimenCount>
similar to
<typeStatus id="54BAFB89F56CFFC8CD523A76FA2BFC5F" box="[1331,1432,871,897]" pageId="1" pageNumber="268" type="holotype">holotype</typeStatus>
,
<materialsCitation id="3B694F76F56CFFC8C8F63A9AFE7CFC17" collectingDate="2008-10-09" collectionCode="LO" collectorName="F. Meza" country="Peru" elevation="174" location="Alto Rio Chambira" municipality="Rio Chambira" pageId="1" pageNumber="268" specimenCode="T32" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D078EA2F56CFFC8C8F63A9AFF09FC7B" box="[151,186,907,933]" count="1" pageId="1" pageNumber="268" type="male">1♂</specimenCount>
<collectingCountry id="F31605BBF56CFFC8C8AA3A9AFEAAFC7B" box="[203,281,907,933]" name="Peru" pageId="1" pageNumber="268">PERU</collectingCountry>
:
<collectionCode id="ED10DDEEF56CFFC8C9423A9AFEFFFC7B" box="[291,332,907,933]" country="Sweden" name="Type Collection" pageId="1" pageNumber="268">LO</collectionCode>
.
<location id="8EDE13F0F56CFFC8C9353A9AFD9AFC7B" LSID="urn:lsid:plazi:treatment:03A8F43DF56CFFCDC8F63BC7FB16FDC7:8EDE13F0F56CFFC8C9353A9AFD9AFC7B" box="[340,553,907,933]" country="Peru" municipality="Rio Chambira" name="Alto Rio Chambira" pageId="1" pageNumber="268">
Alto
<collectingMunicipality id="6BDADF51F56CFFC8C9ED3A9AFD9AFC7B" box="[396,553,907,933]" pageId="1" pageNumber="268">Rio Chambira</collectingMunicipality>
</location>
18M 569924E 9674657N
<quantity id="4CF9E8CEF56CFFC8CB323A9AFC25FC7B" box="[851,918,907,933]" metricMagnitude="2" metricUnit="m" metricValue="1.7399999999999998" pageId="1" pageNumber="268" unit="m" value="174.0">
<elevation id="002CA218F56CFFC8CB323A9AFC25FC7B" box="[851,918,907,933]" metricMagnitude="2" metricUnit="m" metricValue="1.7399999999999998" pageId="1" pageNumber="268" unit="m" value="174.0">174m</elevation>
</quantity>
,
<date id="FFBF63EBF56CFFC8CBC13A9AFBB7FC7B" box="[928,1028,907,933]" pageId="1" pageNumber="268" value="2008-10-09">
<collectingDate id="EFFB9A03F56CFFC8CBC13A9AFBB7FC7B" box="[928,1028,907,933]" pageId="1" pageNumber="268" value="2008-10-09">9.x.2008</collectingDate>
</date>
,
<collectorName id="26F420FDF56CFFC8CC713A9AFBDBFC7B" box="[1040,1128,907,933]" pageId="1" pageNumber="268">F. Meza</collectorName>
// Colinas fuertemnt disect,
<collectingMethod id="52403D3CF56CFFC8C8F63ABEFEA9FC17" box="[151,282,943,969]" pageId="1" pageNumber="268">Yellow trap</collectingMethod>
11
<specimenCode id="DBA7ED50F56CFFC8C95A3ABEFEDFFC17" box="[315,364,943,969]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="268">T32</specimenCode>
Y6”, and
</materialsCitation>
<materialsCitation id="3B694F76F56CFFC8C9B63ABEFEDCFC33" collectingDate="2011-01-19" collectingDateMax="2011-01-21" collectingDateMin="2011-01-19" collectionCode="MD, MUSM" collectorName="J. Costa" country="Peru" elevation="428" latitude="-12.715417" location="Manu" longLatPrecision="1" longitude="-71.24211" municipality="Rio Serjali" pageId="1" pageNumber="268" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D078EA2F56CFFC8C9B63ABEFE49FC17" box="[471,506,943,969]" count="1" pageId="1" pageNumber="268" type="female">1♀</specimenCount>
<collectingCountry id="F31605BBF56CFFC8CA6D3ABEFDE9FC17" box="[524,602,943,969]" name="Peru" pageId="1" pageNumber="268">PERU</collectingCountry>
:
<collectionCode id="ED10DDEEF56CFFC8CA043ABEFD26FC17" box="[613,661,943,969]" country="Germany" name="Museum Donaueschingen" pageId="1" pageNumber="268">MD</collectionCode>
,
<location id="8EDE13F0F56CFFC8CAC03ABEFD56FC17" LSID="urn:lsid:plazi:treatment:03A8F43DF56CFFCDC8F63BC7FB16FDC7:8EDE13F0F56CFFC8CAC03ABEFD56FC17" box="[673,741,943,969]" country="Peru" latitude="-12.715417" longLatPrecision="1" longitude="-71.24211" municipality="Rio Serjali" name="Manu" pageId="1" pageNumber="268">Manu</location>
,
<collectingMunicipality id="6BDADF51F56CFFC8CA913ABEFCD9FC17" box="[752,874,943,969]" pageId="1" pageNumber="268">Rio Serjali</collectingMunicipality>
<geoCoordinate id="EE3523ECF56CFFC8CB103ABFFBBBFC17" box="[881,1032,942,969]" degrees="12" direction="south" minutes="42" orientation="latitude" pageId="1" pageNumber="268" precision="1" seconds="55.5" value="-12.715417">12º4255.5”S</geoCoordinate>
<geoCoordinate id="EE3523ECF56CFFC8CC713ABEFB02FC17" box="[1040,1201,943,969]" degrees="71" direction="west" minutes="14" orientation="longitude" pageId="1" pageNumber="268" precision="1" seconds="31.6" value="-71.24211">71º1431.6”W</geoCoordinate>
<quantity id="4CF9E8CEF56CFFC8CCD83ABEFB48FC17" box="[1209,1275,943,969]" metricMagnitude="2" metricUnit="m" metricValue="4.279999999999999" pageId="1" pageNumber="268" unit="m" value="428.0">
<elevation id="002CA218F56CFFC8CCD83ABEFB48FC17" box="[1209,1275,943,969]" metricMagnitude="2" metricUnit="m" metricValue="4.279999999999999" pageId="1" pageNumber="268" unit="m" value="428.0">428m</elevation>
</quantity>
,
<date id="FFBF63EBF56CFFC8CD663ABEFA2BFC17" box="[1287,1432,943,969]" pageId="1" pageNumber="268" value="2011-01-19" valueMax="2011-01-21" valueMin="2011-01-19">
<collectingDate id="EFFB9A03F56CFFC8CD663ABEFA2BFC17" box="[1287,1432,943,969]" pageId="1" pageNumber="268" value="2011-01-19" valueMax="2011-01-21" valueMin="2011-01-19">19-21.i.2011</collectingDate>
</date>
,
<collectorName id="26F420FDF56CFFC8C8F63AC2FF41FC33" box="[151,242,979,1005]" pageId="1" pageNumber="268">J. Costa</collectorName>
” (
<collectionCode id="ED10DDEEF56CFFC8C96A3AC2FED4FC33" box="[267,359,979,1005]" pageId="1" pageNumber="268">MUSM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31B16A0F56CFFC8C8A63AE7FC13FBA3" pageId="1" pageNumber="268" type="diagnosis">
<paragraph id="8BBE452BF56CFFC8C8A63AE7FC13FBA3" blockId="1.[151,1437,834,2013]" pageId="1" pageNumber="268">
<emphasis id="B9759939F56CFFC8C8A63AE7FEF1FBCE" bold="true" box="[199,322,1014,1040]" pageId="1" pageNumber="268">Diagnosis.</emphasis>
<taxonomicName id="4C013EA8F56CFFC8C92D3AE6FE4AFBCE" authorityName="Inga &amp; Alvarado" authorityYear="2024" box="[332,505,1015,1041]" class="Insecta" family="Ichneumonidae" genus="Polyphrix" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="268" phylum="Arthropoda" rank="species" species="misa" status="sp. nov.">
<emphasis id="B9759939F56CFFC8C92D3AE6FE4AFBCE" box="[332,505,1015,1041]" italics="true" pageId="1" pageNumber="268">Polyphrix misa</emphasis>
</taxonomicName>
<emphasis id="B9759939F56CFFC8CA623AE6FDD3FBCF" bold="true" box="[515,608,1015,1041]" pageId="1" pageNumber="268">
<taxonomicNameLabel id="A2462442F56CFFC8CA623AE6FDD3FBCF" box="[515,608,1015,1041]" pageId="1" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
may be distinguished from all other species by the combination of the following characters: mesosoma completely orange, metasomal tergites 28 black with apical white stripe and hind femur black. It is quite similar to
<taxonomicName id="4C013EA8F56CFFC8CA763D2EFD3BFB86" authorityName="Townes" authorityYear="1970" box="[535,648,1087,1112]" class="Insecta" family="Ichneumonidae" genus="Polyphrix" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="268" phylum="Arthropoda" rank="species" species="varians">
<emphasis id="B9759939F56CFFC8CA763D2EFD3BFB86" box="[535,648,1087,1112]" italics="true" pageId="1" pageNumber="268">P. varians</emphasis>
</taxonomicName>
but differs in having the hind femur black (vs. orange) and the genal orbits pale yellow but interrupted at mid gena (vs. completely white).
</paragraph>
</subSubSection>
<subSubSection id="C31B16A0F56CFFCDC8A63D97FADFFE0F" lastPageId="4" lastPageNumber="271" pageId="1" pageNumber="268" type="description">
<paragraph id="8BBE452BF56CFFC8C8A63D97FD98FB7F" blockId="1.[151,1437,834,2013]" box="[199,555,1158,1185]" pageId="1" pageNumber="268">
<emphasis id="B9759939F56CFFC8C8A63D97FD98FB7F" bold="true" box="[199,555,1158,1185]" pageId="1" pageNumber="268">
Description. Female
<typeStatus id="54BAFB89F56CFFC8C9DF3D96FD94FB7F" box="[446,551,1159,1185]" pageId="1" pageNumber="268" type="holotype">holotype</typeStatus>
.
</emphasis>
</paragraph>
<paragraph id="8BBE452BF56CFFC8C8A63DBAFC33FA8B" blockId="1.[151,1437,834,2013]" pageId="1" pageNumber="268">
Fore wing
<quantity id="4CF9E8CEF56CFFC8C9283DBAFE71FB1B" box="[329,450,1195,1221]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" pageId="1" pageNumber="268" unit="mm" value="10.2">10.20 mm</quantity>
. Body entirely shiny, scarcely punctate and very sparsely pilose.
<emphasis id="B9759939F56CFFC8CCA23DBAFAB3FB1B" box="[1219,1280,1195,1221]" italics="true" pageId="1" pageNumber="268">Head</emphasis>
(
<figureCitation id="133A59AEF56CFFC8CD733DBAFA3CFB1B" box="[1298,1423,1195,1221]" captionStart="FIGURE 1" captionStartId="2.[152,255,1919,1943]" captionTargetBox="[151,1435,309,1896]" captionTargetId="figure-93@2.[151,1435,309,1896]" captionTargetPageId="2" captionText="FIGURE 1. Polyphrix misa sp. nov., female, holotype. A. habitus, lateral view; B. head, front view; C. pronotum, lateral view; D. mesosoma and metasoma, dorsal view; E. wings; F. ovipositor tip, lateral view. Scale bar = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.13618448" httpUri="https://zenodo.org/record/13618448/files/figure.png" pageId="1" pageNumber="268">Figs 1A,B</figureCitation>
): antenna with 33 flagellomeres; clypeus scarcely punctate, CWH 2.00, CWW 1,26 apex slightly convex; mandible and palpi sparsely pilose; mandible moderately long, MLW 2.10, MWW 0.60, ventral tooth slightly longer and more robust than dorsal tooth; occipital carina moderately high, sharp, uniformly curved, ending far from hypostomal carina, at distance as large as basal width of mandible; MSM 0.50.
</paragraph>
<paragraph id="8BBE452BF56CFFC8C8A63C4EFC96F803" blockId="1.[151,1437,834,2013]" pageId="1" pageNumber="268">
<taxonomicName id="4C013EA8F56CFFC8C8A63C4EFE8CFAA6" box="[199,319,1375,1400]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="268" phylum="Arthropoda" rank="genus">
<emphasis id="B9759939F56CFFC8C8A63C4EFE8CFAA6" box="[199,319,1375,1400]" italics="true" pageId="1" pageNumber="268">Mesosoma</emphasis>
</taxonomicName>
(
<figureCitation id="133A59AEF56CFFC8C9313C4EFE48FAA7" box="[336,507,1375,1401]" captionStart="FIGURE 1" captionStartId="2.[152,255,1919,1943]" captionTargetBox="[151,1435,309,1896]" captionTargetId="figure-93@2.[151,1435,309,1896]" captionTargetPageId="2" captionText="FIGURE 1. Polyphrix misa sp. nov., female, holotype. A. habitus, lateral view; B. head, front view; C. pronotum, lateral view; D. mesosoma and metasoma, dorsal view; E. wings; F. ovipositor tip, lateral view. Scale bar = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.13618448" httpUri="https://zenodo.org/record/13618448/files/figure.png" pageId="1" pageNumber="268">Figs 1A, 1C,D</figureCitation>
): pronotum with weak and short wrinkles at posterior margin of collar inferiorly; dorsal margin regular, neither swollen nor prominent; mesoscutum subcircular; notaulus almost absent, traceable only in tangent anterior view; lateral carina of scutellum weak, distinct only on anterior 0.2; epicnemial carina restricted to ventral 0.70 of mesopleuron, almost straight, mesopleuron dorsad of carina giving rise to short and very weak transverse strigulation; mesopleural fovea shallow; mesopleural suture straight, narrow and crenulate; median portion of postpectal carina very weak, short and slightly arched forward; metapleuron smooth, shiny, with sparse hairs; justacoxal carina present only as very short subvertical ridge; transverse furrow at base of propodeum centrally 0.70 as long as distance anterior transverse carina to anterior margin of propodeum, measured centrally.
<emphasis id="B9759939F56CFFC8C8F63F6EFF7FF946" box="[151,204,1663,1688]" italics="true" pageId="1" pageNumber="268">Legs</emphasis>
: hind pre-apical tarsomere not distinctly bilobed; FLW 7.40. Propodeum anterior margin centrally concave; SWL 2.00; anterior transverse carina straight, area behind it with 8 or 9 distinct straight transverse wrinkles, either complete or incomplete, posterior ones stronger and sharper.
<emphasis id="B9759939F56CFFC8CB5A3FD6FCCCF93E" box="[827,895,1735,1760]" italics="true" pageId="1" pageNumber="268">Wings</emphasis>
(
<figureCitation id="133A59AEF56CFFC8CBEC3FD6FBA5F93F" box="[909,1046,1735,1761]" captionStart="FIGURE 1" captionStartId="2.[152,255,1919,1943]" captionTargetBox="[151,1435,309,1896]" captionTargetId="figure-93@2.[151,1435,309,1896]" captionTargetPageId="2" captionText="FIGURE 1. Polyphrix misa sp. nov., female, holotype. A. habitus, lateral view; B. head, front view; C. pronotum, lateral view; D. mesosoma and metasoma, dorsal view; E. wings; F. ovipositor tip, lateral view. Scale bar = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.13618448" httpUri="https://zenodo.org/record/13618448/files/figure.png" pageId="1" pageNumber="268">Figs 1A, 1E</figureCitation>
): fore wing vein 1-Rs+M distinctly sinuous, without bulla; crossvein 1m-cu uniformly arched, slightly shorter than 1-Rs+M; vein 1M+Rs anteriorly straight, posteriorly arched; crossvein 1cu-a straight, forming straight angle with M+Cu, ending basad of vein 1M+Rs by about 0.2 its own length; crossvein 2cu-a 0.25 as long as vein 2-Cu; cell 1+2Rs (areolet) of moderate size, APH 0.15, transversely rectangular, AWH 2.13; crossvein 3r-m spectral, distinctly longer than 2r-m; hind wing vein M+Cu forming straight angle with vein M; HW1C 0.89; vein 2-Rs entirely tubular; crossvein 1r-m with bulla at ventral 0.3; veins 1-Rs and 2-Rs distinctly angled; vein Cub straight, reaching about 0.8 of distance to wing margin; vein 2-1A reaching about 0.85 of distance to wing margin.
</paragraph>
<paragraph id="8BBE452BF56FFFCBC8A63986FD71FEC3" blockId="2.[151,1436,150,285]" pageId="2" pageNumber="269">
<emphasis id="B9759939F56FFFCBC8A63986FE8FFF6E" box="[199,316,151,176]" italics="true" pageId="2" pageNumber="269">Metasoma</emphasis>
(
<figureCitation id="133A59AEF56FFFCBC92B3986FDB3FF6F" box="[330,512,151,177]" captionStart="FIGURE 1" captionStartId="2.[152,255,1919,1943]" captionTargetBox="[151,1435,309,1896]" captionTargetId="figure-93@2.[151,1435,309,1896]" captionTargetPageId="2" captionText="FIGURE 1. Polyphrix misa sp. nov., female, holotype. A. habitus, lateral view; B. head, front view; C. pronotum, lateral view; D. mesosoma and metasoma, dorsal view; E. wings; F. ovipositor tip, lateral view. Scale bar = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.13618448" httpUri="https://zenodo.org/record/13618448/files/figure.png" pageId="2" pageNumber="269">Figs 1A, 1D, 1F</figureCitation>
): first tergite elongate, slender, T1LW 3.64, T1WW 1.57; spiracle at center, slightly protuberant; T2LW 1.65, T2WW 1.85; tergites 38 smooth, shiny, pilose; OST 1.59; ovipositor moderately slender, straight, scarcely punctate, laterally strongly compressed; dorsal valve with nodus and notch, ventral valve with eight apical teeth; apex moderately long, pointed.
</paragraph>
<caption id="DF7E15A3F56FFFCBC8F93E6EFB20F865" ID-DOI="http://doi.org/10.5281/zenodo.13618448" ID-Zenodo-Dep="13618448" httpUri="https://zenodo.org/record/13618448/files/figure.png" pageId="2" pageNumber="269" startId="2.[152,255,1919,1943]" targetBox="[151,1435,309,1896]" targetPageId="2" targetType="figure">
<paragraph id="8BBE452BF56FFFCBC8F93E6EFB20F865" blockId="2.[151,1436,1919,1980]" pageId="2" pageNumber="269">
<emphasis id="B9759939F56FFFCBC8F93E6EFDBCF846" bold="true" box="[152,527,1919,1944]" pageId="2" pageNumber="269">
FIGURE 1.
<taxonomicName id="4C013EA8F56FFFCBC97C3E6EFE09F849" authorityName="Inga &amp; Alvarado" authorityYear="2024" box="[285,442,1919,1943]" class="Insecta" family="Ichneumonidae" genus="Polyphrix" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="269" phylum="Arthropoda" rank="species" species="misa" status="sp. nov.">
<emphasis id="B9759939F56FFFCBC97C3E6EFE09F849" bold="true" box="[285,442,1919,1943]" italics="true" pageId="2" pageNumber="269">Polyphrix misa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2462442F56FFFCBC9A13E91FDBCF846" box="[448,527,1920,1944]" pageId="2" pageNumber="269" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female, holotype.
<emphasis id="B9759939F56FFFCBCAB03E6EFD5AF849" bold="true" box="[721,745,1919,1943]" pageId="2" pageNumber="269">A.</emphasis>
habitus, lateral view;
<emphasis id="B9759939F56FFFCBCBA73E91FC6EF846" bold="true" box="[966,989,1920,1944]" pageId="2" pageNumber="269">B.</emphasis>
head, front view;
<emphasis id="B9759939F56FFFCBCCF23E6EFB18F849" bold="true" box="[1171,1195,1919,1943]" pageId="2" pageNumber="269">C.</emphasis>
pronotum, lateral view;
<emphasis id="B9759939F56FFFCBC8F63EB5FF03F862" bold="true" box="[151,176,1956,1980]" pageId="2" pageNumber="269">D.</emphasis>
mesosoma and metasoma, dorsal view;
<emphasis id="B9759939F56FFFCBCA283EB5FDECF862" bold="true" box="[585,607,1956,1980]" pageId="2" pageNumber="269">E.</emphasis>
wings;
<emphasis id="B9759939F56FFFCBCAD13EB5FD70F862" bold="true" box="[688,707,1956,1980]" pageId="2" pageNumber="269">F.</emphasis>
ovipositor tip, lateral view. Scale bar = 1 mm
</paragraph>
</caption>
<caption id="DF7E15A3F56EFFCAC8F93C8EFD8DFA05" ID-DOI="http://doi.org/10.5281/zenodo.13618450" ID-Zenodo-Dep="13618450" httpUri="https://zenodo.org/record/13618450/files/figure.png" pageId="3" pageNumber="270" startId="3.[152,255,1439,1463]" targetBox="[151,1436,181,1415]" targetPageId="3" targetType="figure">
<paragraph id="8BBE452BF56EFFCAC8F93C8EFD8DFA05" blockId="3.[151,1437,1439,1500]" pageId="3" pageNumber="270">
<emphasis id="B9759939F56EFFCAC8F93C8EFDA2FA66" bold="true" box="[152,529,1439,1464]" pageId="3" pageNumber="270">
FIGURE 2.
<taxonomicName id="4C013EA8F56EFFCAC97C3C8EFE08FA69" authorityName="Inga &amp; Alvarado" authorityYear="2024" box="[285,443,1439,1463]" class="Insecta" family="Ichneumonidae" genus="Polyphrix" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="270" phylum="Arthropoda" rank="species" species="misa" status="sp. nov.">
<emphasis id="B9759939F56EFFCAC97C3C8EFE08FA69" bold="true" box="[285,443,1439,1463]" italics="true" pageId="3" pageNumber="270">Polyphrix misa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2462442F56EFFCAC9A03CB1FDA2FA66" box="[449,529,1440,1464]" pageId="3" pageNumber="270" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male, paratype.
<emphasis id="B9759939F56EFFCACADC3C8EFD65FA69" bold="true" box="[701,726,1439,1463]" pageId="3" pageNumber="270">A.</emphasis>
habitus, lateral view;
<emphasis id="B9759939F56EFFCACBD53CB1FC78FA66" bold="true" box="[948,971,1440,1464]" pageId="3" pageNumber="270">B.</emphasis>
head, front view;
<emphasis id="B9759939F56EFFCACCE03C8EFB29FA69" bold="true" box="[1153,1178,1439,1463]" pageId="3" pageNumber="270">C.</emphasis>
wings;
<emphasis id="B9759939F56EFFCACC883CB1FAB2FA66" bold="true" box="[1257,1281,1440,1464]" pageId="3" pageNumber="270">D.</emphasis>
mesosoma and metasoma, dorsal view. Scale bar = 1 mm
</paragraph>
</caption>
<paragraph id="8BBE452BF56EFFCAC8A63F3FFF42F852" blockId="3.[151,1437,1582,2040]" pageId="3" pageNumber="270">
<emphasis id="B9759939F56EFFCAC8A63F3FFEBBF996" box="[199,264,1582,1608]" italics="true" pageId="3" pageNumber="270">Color</emphasis>
(
<figureCitation id="133A59AEF56EFFCAC9793F3FFEEAF996" box="[280,345,1582,1608]" captionStart="FIGURE 1" captionStartId="2.[152,255,1919,1943]" captionTargetBox="[151,1435,309,1896]" captionTargetId="figure-93@2.[151,1435,309,1896]" captionTargetPageId="2" captionText="FIGURE 1. Polyphrix misa sp. nov., female, holotype. A. habitus, lateral view; B. head, front view; C. pronotum, lateral view; D. mesosoma and metasoma, dorsal view; E. wings; F. ovipositor tip, lateral view. Scale bar = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.13618448" httpUri="https://zenodo.org/record/13618448/files/figure.png" pageId="3" pageNumber="270">Fig. 1</figureCitation>
). Head: scape, pedicel and flagellomeres black except for flagellomeres T717, T6 distal end and T1823 dorsally white, apically flagellomeres dark brown; supra-clypeal area, clypeus, mouth parts, malar space and orbital and gena band (interrupted at mid gena) pale yellow; apex of mandible dark brown; supra-antennal area, occiput, temple and except orbital band, black.
<taxonomicName id="4C013EA8F56EFFCACA873F8BFCD7F96A" box="[742,868,1690,1716]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="271" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
: orange except for fore and mid legs with tibia and tarsomeres 14 pale yellow, tarsomere 5 brown; hind leg with trochantellus, femur and apical third of tarsomere 5 brown, tibia and basitarsus pale yellow, and tarsomeres 24 and tarsomere 5 (except distal third) white. Metasoma: Tergites usually black; T1 basally orange to apically black, apical stripe pale yellow; T2 basally (at the center) dark orange; T27 basally black with apical and lateral stripes pale yellow; T8 basally black with lateral stripes pale yellow; S13 pale yellow with small black stripes on the sides; ovipositor dark red, sheaths dark brown; wings hyaline.
</paragraph>
<paragraph id="8BBE452BF56EFFCDC8A63E87FE2FFF27" blockId="3.[151,1437,1582,2040]" lastBlockId="4.[151,1437,150,537]" lastPageId="4" lastPageNumber="271" pageId="3" pageNumber="270">
<emphasis id="B9759939F56EFFCAC8A63E87FDAEF871" box="[199,541,1942,1968]" italics="true" pageId="3" pageNumber="270">
Variation of female
<typeStatus id="54BAFB89F56EFFCAC9C63E87FDAAF871" box="[423,537,1942,1967]" pageId="3" pageNumber="270" type="paratype">paratypes</typeStatus>
.
</emphasis>
Fore wing length
<quantity id="4CF9E8CEF56EFFCACA8D3E87FC36F86E" box="[748,901,1942,1968]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.08" metricValueMin="0.9199999999999999" pageId="3" pageNumber="270" unit="mm" value="10.0" valueMax="10.8" valueMin="9.2">9.210.8 mm</quantity>
. They differ from the
<typeStatus id="54BAFB89F56EFFCACC1F3E87FB53F86E" box="[1150,1248,1942,1968]" pageId="3" pageNumber="270" type="holotype">holotype</typeStatus>
in the following features: CWH 1.581.80, CWW 1.231.25, MLW 1.922.20, MWW 0.690.70, MSM 0.540.70;
<taxonomicName id="4C013EA8F56EFFCACC8A3EABFAD7F80A" box="[1259,1380,1978,2004]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="271" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
with legs FLW 6.807.64, SWL 1.802.00; fore wing APH 0.160.26, AWH 1.782.00; hind wing HW1C 1.001.15; metasoma with T1LW 3.583.89, T1WW 1.802.00, T2LW 1.601.71, T2WW 1.561.81, OST 1.291.48. Within the coloration orbital and gena band complete or interrupted at middle, antenna with 3334 flagellomeres with T1821 dorsally white.
</paragraph>
<paragraph id="8BBE452BF569FFCDC8A63812FADFFE0F" blockId="4.[151,1437,150,537]" pageId="4" pageNumber="271">
<emphasis id="B9759939F569FFCDC8A63812FEC5FEC2" box="[199,374,259,285]" italics="true" pageId="4" pageNumber="271">
Male
<typeStatus id="54BAFB89F569FFCDC9693812FEC5FEC2" box="[264,374,259,284]" pageId="4" pageNumber="271" type="paratype">paratypes</typeStatus>
</emphasis>
(
<figureCitation id="133A59AEF569FFCDC9E73812FE70FEC3" box="[390,451,259,285]" captionStart="FIGURE 2" captionStartId="3.[152,255,1439,1463]" captionTargetBox="[151,1436,181,1415]" captionTargetId="figure-26@3.[151,1436,181,1415]" captionTargetPageId="3" captionText="FIGURE 2. Polyphrix misa sp. nov., male, paratype. A. habitus, lateral view; B. head, front view; C. wings; D. mesosoma and metasoma, dorsal view. Scale bar = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.13618450" httpUri="https://zenodo.org/record/13618450/files/figure.png" pageId="4" pageNumber="271">Fig 2</figureCitation>
). Fore wing length
<quantity id="4CF9E8CEF569FFCDCAC03813FC98FEC3" box="[673,811,258,285]" metricMagnitude="-3" metricUnit="m" metricValue="8.65" metricValueMax="8.8" metricValueMin="8.5" pageId="4" pageNumber="271" unit="mm" value="8.65" valueMax="8.8" valueMin="8.5">8.58.8 mm</quantity>
. They differ from the females in the following features: CWH 1.772.36, CWW 1.181.28, MLW 1.831.92, MWW 0.540.58, MSM 0.420.50;
<taxonomicName id="4C013EA8F569FFCDCC1A3836FB47FE9F" box="[1147,1268,295,321]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="271" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
with legs FLW 7.448.00, SWL 1.401.50; fore wing APH 0.160.23, AWH 1.502.14; hind wing HW1C 1.151.60; metasoma with T1LW 4.334.41, T1WW 1.311.50, T2LW 2.603.08, T2WW 1.331.56. Antennal coloration differs as follows: pedicel and flagellomeres black except for flagellomeres T522, T6 distal half and T9, 2327 dorsally white or flagellomeres T1021, T6 distal half and T9, 2223 dorsally whit, apically flagellomeres dark brown.
</paragraph>
</subSubSection>
<subSubSection id="C31B16A0F569FFCDC8A638CBFC68FE2B" box="[199,987,474,501]" pageId="4" pageNumber="271" type="distribution">
<paragraph id="8BBE452BF569FFCDC8A638CBFC68FE2B" blockId="4.[151,1437,150,537]" box="[199,987,474,501]" pageId="4" pageNumber="271">
<emphasis id="B9759939F569FFCDC8A638CBFED2FE2A" bold="true" box="[199,353,474,500]" pageId="4" pageNumber="271">Distribution.</emphasis>
<collectingCountry id="F31605BBF569FFCDC90638CAFE2EFE2B" box="[359,413,475,501]" name="Peru" pageId="4" pageNumber="271">Peru</collectingCountry>
,
<collectingRegion id="49C58BC9F569FFCDC9C838CAFE47FE2B" box="[425,500,475,501]" country="Peru" name="Loreto" pageId="4" pageNumber="271">Loreto</collectingRegion>
and
<collectingRegion id="49C58BC9F569FFCDCA4B38CAFD63FE2B" box="[554,720,475,501]" country="Peru" name="Madre de Dios" pageId="4" pageNumber="271">Madre de Dios</collectingRegion>
Departments (
<figureCitation id="133A59AEF569FFCDCB1738CAFC7CFE2B" box="[886,975,475,501]" captionStart="FIGURE 4" captionStartId="7.[152,255,666,690]" captionTargetBox="[151,1436,181,643]" captionTargetId="figure-26@7.[151,1436,181,643]" captionTargetPageId="7" captionText="FIGURE 4. Geographic records of Polyphrix. A. Polyphrix stellata; B. Polyphrix varians. C. Polyphrix misa sp. nov. and Polyphrix tullu sp. nov. Yellow circles represent new records and red represent previous records." figureDoi="http://doi.org/10.5281/zenodo.13618454" httpUri="https://zenodo.org/record/13618454/files/figure.png" pageId="4" pageNumber="271">Fig. 4C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31B16A0F569FFCDC8A638EEFB16FDC7" box="[199,1189,511,537]" pageId="4" pageNumber="271" type="etymology">
<paragraph id="8BBE452BF569FFCDC8A638EEFB16FDC7" blockId="4.[151,1437,150,537]" box="[199,1189,511,537]" pageId="4" pageNumber="271">
<emphasis id="B9759939F569FFCDC8A638EEFEFDFDC7" bold="true" box="[199,334,511,537]" pageId="4" pageNumber="271">Etymology.</emphasis>
The specific epithet
<emphasis id="B9759939F569FFCDCA2338EEFD33FDC7" box="[578,640,511,537]" italics="true" pageId="4" pageNumber="271">misa</emphasis>
means ʻbicolorʼ in Quechua, noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,61 +1,69 @@
<document id="2A50EC204C84E72D9B0B3A57CB65531B" ID-ISSN="2101-0315" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1723047137059" checkinUser="felipe" docAuthor="Keith, Philippe, Hadiaty, Renny, Hubert, Nicolas, Busson, Frédéric &amp; Lord, Clara" docDate="2014" docId="03D25F043100FFACFC8EF90AFE153921" docLanguage="en" docName="Cybium.38.2.133-146.pdf" docOrigin="Cybium 38 (2)" docStyle="DocumentStyle:73DAD84611431DC6606F8B63B2C6039C.1:Cybium.2013-2018.journal_article.type1" docStyleId="73DAD84611431DC6606F8B63B2C6039C" docStyleName="Cybium.2013-2018.journal_article.type1" docStyleVersion="1" docTitle="Lentipes ikeae Keith, Hadiaty, Hubert, Busson &amp; Lord, 2014, n. sp." docType="treatment" docVersion="1" lastPageNumber="145" masterDocId="FFEB277C310AFFA1FFB4FFB0FFA0385E" masterDocTitle="Three new species of Lentipes from Indonesia (Gobiidae)" masterLastPageNumber="146" masterPageNumber="133" pageNumber="143" updateTime="1723047857054" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-4.0">
<mods:mods id="4E35D6490CA15164D954DE48374D6B5C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="EFCCEFDE25EC379643251FB139B09788">Three new species of Lentipes from Indonesia (Gobiidae)</mods:title>
<document id="606A2C6AA558103BA56B36A496166088" ID-DOI="10.26028/cybium/2014-382-004" ID-ISSN="2101-0315" ID-Zenodo-Dep="13618360" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1723047137059" checkinUser="felipe" docAuthor="Keith, Philippe, Hadiaty, Renny K., Hubert, Nicolas, Busson, Frédéric &amp; Lord, Clara" docDate="2014" docId="03D25F043100FFACFC8EF90AFE153921" docLanguage="en" docName="Cybium.38.2.133-146.pdf" docOrigin="Cybium 38 (2)" docSource="http://dx.doi.org/10.26028/cybium/2014-382-004" docStyle="DocumentStyle:73DAD84611431DC6606F8B63B2C6039C.1:Cybium.2013-2018.journal_article.type1" docStyleId="73DAD84611431DC6606F8B63B2C6039C" docStyleName="Cybium.2013-2018.journal_article.type1" docStyleVersion="1" docTitle="Lentipes ikeae Keith, Hadiaty, Hubert, Busson &amp; Lord, 2014, n. sp." docType="treatment" docVersion="3" lastPageNumber="145" masterDocId="FFEB277C310AFFA1FFB4FFB0FFA0385E" masterDocTitle="Three new species of Lentipes from Indonesia (Gobiidae)" masterLastPageNumber="146" masterPageNumber="133" pageNumber="143" updateTime="1725027590525" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="D17AF1BFD46C500F7B2A8F879F5F58E4">Three new species of Lentipes from Indonesia (Gobiidae)</mods:title>
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@ -73,7 +81,7 @@ Keith, Hubert, Busson &amp; Hadiaty
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(
<figureCitation id="1340F2973100FFABFC4BF96AFBFB3EAC" box="[1023,1115,1754,1778]" captionStart-0="Figure 7" captionStart-1="Figure 8" captionStart-2="Figure 9" captionStartId-0="10.[132,195,608,630]" captionStartId-1="11.[132,198,1042,1064]" captionStartId-2="11.[813,877,767,789]" captionTargetBox-0="[132,1455,215,591]" captionTargetBox-1="[151,775,226,1017]" captionTargetBox-2="[827,1453,253,741]" captionTargetId-0="figure-645@10.[132,1455,215,591]" captionTargetId-1="figure-583@11.[132,775,215,1025]" captionTargetId-2="figure-584@11.[813,1456,215,750]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="11" captionText-0="Figure 7. - Lentipes ikeae. A: Male (picture by F. Busson); B:Above, male, Paratype MZB 21387 (picture by N. Hubert; BIF 1730); Below, female (picture by N. Hubert; BIF 1747)." captionText-1="Figure 8. - Diagrammatic illustration of head in Lentipes ikeae (male) showing head pores and sensory papillae. A: Dorsal view; B: Lateral view." captionText-2="Figure 9. - Diagrammatic illustration of urogenital papilla in Lentipes ikeae. A: Male; B: Female. 1: anus; 2: urogenital papilla; 3: anal fin." pageId="10" pageNumber="143">Figs 7-9</figureCitation>
<figureCitation id="1340F2973100FFABFC4BF96AFBFB3EAC" box="[1023,1115,1754,1778]" captionStart-0="Figure 7" captionStart-1="Figure 8" captionStart-2="Figure 9" captionStartId-0="10.[132,195,608,630]" captionStartId-1="11.[132,198,1042,1064]" captionStartId-2="11.[813,877,767,789]" captionTargetBox-0="[132,1455,215,591]" captionTargetBox-1="[151,775,226,1017]" captionTargetBox-2="[827,1453,253,741]" captionTargetId-0="figure-645@10.[132,1455,215,591]" captionTargetId-1="figure-583@11.[132,775,215,1025]" captionTargetId-2="figure-584@11.[813,1456,215,750]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="11" captionText-0="Figure 7. - Lentipes ikeae. A: Male (picture by F. Busson); B:Above, male, Paratype MZB 21387 (picture by N. Hubert; BIF 1730); Below, female (picture by N. Hubert; BIF 1747)." captionText-1="Figure 8. - Diagrammatic illustration of head in Lentipes ikeae (male) showing head pores and sensory papillae. A: Dorsal view; B: Lateral view." captionText-2="Figure 9. - Diagrammatic illustration of urogenital papilla in Lentipes ikeae. A: Male; B: Female. 1: anus; 2: urogenital papilla; 3: anal fin." figureDoi-0="http://doi.org/10.5281/zenodo.13618380" figureDoi-1="http://doi.org/10.5281/zenodo.13618386" figureDoi-2="http://doi.org/10.5281/zenodo.13618382" httpUri-0="https://zenodo.org/record/13618380/files/figure.png" httpUri-1="https://zenodo.org/record/13618386/files/figure.png" httpUri-2="https://zenodo.org/record/13618382/files/figure.png" pageId="10" pageNumber="143">Figs 7-9</figureCitation>
, Tabs V-VII)
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@ -102,7 +110,7 @@ having no enlarged lobes associated with the urogenital papillae or elongate fin
<taxonomicNameLabel id="A23C8F7B3101FFAAFF30FA8BFF1D3D0C" box="[132,189,1339,1362]" pageId="11" pageNumber="144" rank="species">n. sp.</taxonomicNameLabel>
in this paper.
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Figure 9. - Diagrammatic illustration of urogenital papilla in
<taxonomicName id="4C7B95913101FFAAFA36FCB0FC363B72" authorityName="Keith &amp; Hadiaty &amp; Hubert &amp; Busson &amp; Lord" authorityYear="2014" family="Gobiidae" genus="Lentipes" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="ikeae">
@ -115,7 +123,7 @@ Figure 9. - Diagrammatic illustration of urogenital papilla in
: Female. 1: anus; 2: urogenital papilla; 3: anal fin.
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Figure 8. - Diagrammatic illustration of head in
<taxonomicName id="4C7B95913101FFAAFDC0FBA3FCA73C76" authorityName="Keith &amp; Hadiaty &amp; Hubert &amp; Busson &amp; Lord" authorityYear="2014" box="[628,775,1042,1064]" family="Gobiidae" genus="Lentipes" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="145" phylum="Chordata" rank="species" species="ikeae">
@ -259,14 +267,14 @@ rays simple; posterior margin slightly straight. Caudal fin (C) with 13 branched
</paragraph>
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Lateral scales (LS) 30(23-35). They are lightly embedded and mainly cycloid in female. Generally not limited to caudal peduncle, many extend anteriorly along midline between second dorsal and anal fins; some ctenoid scales are on the anteriormost part of the flanks. Males have some ctenoid scales, not strongly developed, along midline and beyond the base of pectoral fins. Cycloid scales on caudal peduncle. Scales in transverse backwards (TRB) series 4 (4-8) and in transverse forward series (TRF) 2 (0-7). Zigzag scales (ZZ) 9 (8-10). Head, breast, nape and belly without scales. Upper jaw teeth distinctly tricuspid anteriorly, males 13 (11-22), females (27-35). Premaxilla in males with 3 (3-6) recurved canines posterior to tricuspid teeth; females without teeth posterior to tricuspid teeth. Teeth in lower jaw recurved and canine in males 2 (2-5), no teeth in females. Cephalic sensory pore system A, B, C, D, F, H, K, L, N and O; pore D singular with all others paired (
<figureCitation id="1340F2973106FFADFE6CFDCAFDB93ACC" box="[472,537,634,658]" captionStart="Figure 8" captionStartId="11.[132,198,1042,1064]" captionTargetBox="[151,775,226,1017]" captionTargetId="figure-583@11.[132,775,215,1025]" captionTargetPageId="11" captionText="Figure 8. - Diagrammatic illustration of head in Lentipes ikeae (male) showing head pores and sensory papillae. A: Dorsal view; B: Lateral view." pageId="12" pageNumber="145">Fig. 8</figureCitation>
<figureCitation id="1340F2973106FFADFE6CFDCAFDB93ACC" box="[472,537,634,658]" captionStart="Figure 8" captionStartId="11.[132,198,1042,1064]" captionTargetBox="[151,775,226,1017]" captionTargetId="figure-583@11.[132,775,215,1025]" captionTargetPageId="11" captionText="Figure 8. - Diagrammatic illustration of head in Lentipes ikeae (male) showing head pores and sensory papillae. A: Dorsal view; B: Lateral view." figureDoi="http://doi.org/10.5281/zenodo.13618386" httpUri="https://zenodo.org/record/13618386/files/figure.png" pageId="12" pageNumber="145">Fig. 8</figureCitation>
); oculoscapular canal divided into anterior and posterior canal between pores H and K. Sensory papillae well developed on head and body.
</paragraph>
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Sexual dimorphism well developed. Second dorsal and anal fin lengths, jaw length, predorsal length, head length and caudal peduncle depth greater in males. Urogenital papilla in males slender and pointed distally without associated lobes or expanded tissue (
<figureCitation id="1340F2973106FFADFEEBFCEAFE133B2C" box="[351,435,858,882]" captionStart="Figure 9" captionStartId="11.[813,877,767,789]" captionTargetBox="[827,1453,253,741]" captionTargetId="figure-584@11.[813,1456,215,750]" captionTargetPageId="11" captionText="Figure 9. - Diagrammatic illustration of urogenital papilla in Lentipes ikeae. A: Male; B: Female. 1: anus; 2: urogenital papilla; 3: anal fin." pageId="12" pageNumber="145">Fig. 9A</figureCitation>
<figureCitation id="1340F2973106FFADFEEBFCEAFE133B2C" box="[351,435,858,882]" captionStart="Figure 9" captionStartId="11.[813,877,767,789]" captionTargetBox="[827,1453,253,741]" captionTargetId="figure-584@11.[813,1456,215,750]" captionTargetPageId="11" captionText="Figure 9. - Diagrammatic illustration of urogenital papilla in Lentipes ikeae. A: Male; B: Female. 1: anus; 2: urogenital papilla; 3: anal fin." figureDoi="http://doi.org/10.5281/zenodo.13618382" httpUri="https://zenodo.org/record/13618382/files/figure.png" pageId="12" pageNumber="145">Fig. 9A</figureCitation>
), urogenital papilla retractable into a sheath-like groove; female urogenital papilla rectangular in appearance (
<figureCitation id="1340F2973106FFADFE3FFC2AFE7E3BEF" box="[395,478,922,946]" captionStart="Figure 9" captionStartId="11.[813,877,767,789]" captionTargetBox="[827,1453,253,741]" captionTargetId="figure-584@11.[813,1456,215,750]" captionTargetPageId="11" captionText="Figure 9. - Diagrammatic illustration of urogenital papilla in Lentipes ikeae. A: Male; B: Female. 1: anus; 2: urogenital papilla; 3: anal fin." pageId="12" pageNumber="145">Fig. 9B</figureCitation>
<figureCitation id="1340F2973106FFADFE3FFC2AFE7E3BEF" box="[395,478,922,946]" captionStart="Figure 9" captionStartId="11.[813,877,767,789]" captionTargetBox="[827,1453,253,741]" captionTargetId="figure-584@11.[813,1456,215,750]" captionTargetPageId="11" captionText="Figure 9. - Diagrammatic illustration of urogenital papilla in Lentipes ikeae. A: Male; B: Female. 1: anus; 2: urogenital papilla; 3: anal fin." figureDoi="http://doi.org/10.5281/zenodo.13618382" httpUri="https://zenodo.org/record/13618382/files/figure.png" pageId="12" pageNumber="145">Fig. 9B</figureCitation>
) and also retractable into a sheath-like groove.
</paragraph>
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@ -283,13 +291,13 @@ Sexual dimorphism well developed. Second dorsal and anal fin lengths, jaw length
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<emphasis id="B90F32003106FFADFF30FA4AFED03E4C" bold="true" box="[132,368,1530,1554]" pageId="12" pageNumber="145">
Colour in life (
<figureCitation id="1340F2973106FFADFE93FA4AFEC83E4C" box="[295,360,1530,1554]" captionStart="Figure 7" captionStartId="10.[132,195,608,630]" captionTargetBox="[132,1455,215,591]" captionTargetId="figure-645@10.[132,1455,215,591]" captionTargetPageId="10" captionText="Figure 7. - Lentipes ikeae. A: Male (picture by F. Busson); B:Above, male, Paratype MZB 21387 (picture by N. Hubert; BIF 1730); Below, female (picture by N. Hubert; BIF 1747)." pageId="12" pageNumber="145">Fig. 7</figureCitation>
<figureCitation id="1340F2973106FFADFE93FA4AFEC83E4C" box="[295,360,1530,1554]" captionStart="Figure 7" captionStartId="10.[132,195,608,630]" captionTargetBox="[132,1455,215,591]" captionTargetId="figure-645@10.[132,1455,215,591]" captionTargetPageId="10" captionText="Figure 7. - Lentipes ikeae. A: Male (picture by F. Busson); B:Above, male, Paratype MZB 21387 (picture by N. Hubert; BIF 1730); Below, female (picture by N. Hubert; BIF 1747)." figureDoi="http://doi.org/10.5281/zenodo.13618380" httpUri="https://zenodo.org/record/13618380/files/figure.png" pageId="12" pageNumber="145">Fig. 7</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8BC4EE123106FFADFF1EF9AAFF7A3F6D" blockId="12.[132,775,1530,1907]" pageId="12" pageNumber="145">
<emphasis id="B90F32003106FFADFF1EF9AAFF4C3E6C" box="[170,236,1562,1586]" italics="true" pageId="12" pageNumber="145">Male.</emphasis>
- Background of body greyish. Background of head and snout greyish. Dorsal and ventral margin of head greyish; opercula dark greyish. Lateral midline greyish. Dorsal fins black with a white margin. Belly bright blue. A yel- low to green patch at pectoral base. The second black dorsal fin with one or two black spots rounded with a white margin. A more or less visible vertical bright red band on caudal peduncle. Caudal and pectoral fins translucent. Anal fin greyish.
- Background of body greyish. Background of head and snout greyish. Dorsal and ventral margin of head greyish; opercula dark greyish. Lateral midline greyish. Dorsal fins black with a white margin. Belly bright blue. A yellow to green patch at pectoral base. The second black dorsal fin with one or two black spots rounded with a white margin. A more or less visible vertical bright red band on caudal peduncle. Caudal and pectoral fins translucent. Anal fin greyish.
</paragraph>
<paragraph id="8BC4EE123106FFADFF1EF88BFEF83F2D" blockId="12.[132,775,1530,1907]" pageId="12" pageNumber="145">
<emphasis id="B90F32003106FFADFF1EF88BFF5D3F0D" box="[170,253,1851,1875]" italics="true" pageId="12" pageNumber="145">Female</emphasis>
@ -379,7 +387,7 @@ the 5
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and 6
<superScript id="7C0E435A3106FFADFB66FCC9FB413BD4" attach="left" box="[1234,1249,889,906]" fontSize="7" pageId="12" pageNumber="145">th</superScript>
spines of D2 long- er, and the coloration in males. Furthermore, it differs from
spines of D2 longer, and the coloration in males. Furthermore, it differs from
<taxonomicName id="4C7B95913106FFADFC99FC0EFC003B8B" authorityName="Allen" authorityYear="1997" box="[813,928,958,981]" family="Gobiidae" genus="Lentipes" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="145" phylum="Chordata" rank="species" species="watsoni">
<emphasis id="B90F32003106FFADFC99FC0EFC003B8B" box="[813,928,958,981]" italics="true" pageId="12" pageNumber="145">L. watsoni</emphasis>
</taxonomicName>
@ -429,8 +437,8 @@ The new species is named for Miss Ike Rachmatika, the late staff of Ichthyology
<collectingCounty id="62A5969E3106FFADFBAEF9BBFB9A3E7E" box="[1050,1082,1547,1568]" pageId="12" pageNumber="145">We</collectingCounty>
wish to thank
<collectingMunicipality id="6BA074683106FFADFB7DF9BBFAD23E7E" box="[1225,1394,1546,1568]" pageId="12" pageNumber="145">Laurent Pouyaud</collectingMunicipality>
, Jean-
<location id="8EA4B8C93106FFADFC99F992FC053E66" LSID="urn:lsid:plazi:treatment:03D25F043100FFACFC8EF90AFE153921:8EA4B8C93106FFADFC99F992FC053E66" box="[813,933,1570,1592]" county="We" municipality="Laurent Pouyaud" name="Paul Toutain" pageId="12" pageNumber="145">Paul Toutain</location>
,
<location id="8EA4B8C93106FFADFC99F992FC053E66" LSID="urn:lsid:plazi:treatment:03D25F043100FFACFC8EF90AFE153921:8EA4B8C93106FFADFC99F992FC053E66" box="[813,933,1570,1592]" county="We" municipality="Laurent Pouyaud" name="Paul Toutain" pageId="12" pageNumber="145">Jean-Paul Toutain</location>
and
<location id="8EA4B8C93106FFADFC66F992FBDF3E69" LSID="urn:lsid:plazi:treatment:03D25F043100FFACFC8EF90AFE153921:8EA4B8C93106FFADFC66F992FBDF3E69" box="[978,1151,1570,1592]" county="We" municipality="Laurent Pouyaud" name="Domenico Caruso" pageId="12" pageNumber="145">Domenico Caruso</location>
for their support.
@ -509,7 +517,7 @@ from
<location id="8EA4B8C93107FFACFDDFFF6AFF723959" LSID="urn:lsid:plazi:treatment:03D25F043100FFACFC8EF90AFE153921:8EA4B8C93107FFACFDDFFF6AFF723959" country="France" county="Fondation" municipality="This" name="Forestry Dept in Kendari" pageId="13" pageNumber="146" stateProvince="Sulawesi Utara">Forestry Dept in Kendari</location>
for the nice field trip in
<location id="8EA4B8C93107FFACFE0FFF41FDEC3959" LSID="urn:lsid:plazi:treatment:03D25F043100FFACFC8EF90AFE153921:8EA4B8C93107FFACFE0FFF41FDEC3959" box="[443,588,241,263]" country="France" county="Fondation" municipality="This" name="Mekongga. We" pageId="13" pageNumber="146" stateProvince="Sulawesi Utara">Mekongga. We</location>
wish to thank RIS- TEK and LIPI for the research permits and supporting letter, and
wish to thank RISTEK and LIPI for the research permits and supporting letter, and
<location id="8EA4B8C93107FFACFF30FE91FF1D3969" LSID="urn:lsid:plazi:treatment:03D25F043100FFACFC8EF90AFE153921:8EA4B8C93107FFACFF30FE91FF1D3969" box="[132,189,289,311]" country="France" county="Fondation" municipality="This" name="Daisy" pageId="13" pageNumber="146" stateProvince="Sulawesi Utara">Daisy</location>
and
<location id="8EA4B8C93107FFACFF59FE91FE253969" LSID="urn:lsid:plazi:treatment:03D25F043100FFACFC8EF90AFE153921:8EA4B8C93107FFACFF59FE91FE253969" box="[237,389,289,311]" country="France" county="Fondation" municipality="This" name="Sopian. Finally" pageId="13" pageNumber="146" stateProvince="Sulawesi Utara">Sopian. Finally</location>

View file

@ -1,61 +1,69 @@
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@ -72,7 +80,7 @@ Keith &amp; Hadiaty
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(
<figureCitation id="1340F297310BFFA0FEEBFC58FE1A3C5F" box="[351,442,1000,1025]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="5.[132,195,484,506]" captionStartId-1="5.[132,196,1370,1392]" captionStartId-2="5.[813,877,1089,1111]" captionTargetBox-0="[132,1455,215,467]" captionTargetBox-1="[144,767,542,1336]" captionTargetBox-2="[825,1453,554,1063]" captionTargetId-0="figure-286@5.[132,1455,215,467]" captionTargetId-1="figure-305@5.[132,775,534,1353]" captionTargetId-2="figure-336@5.[813,1456,534,1072]" captionTargetPageId-0="5" captionTargetPageId-1="5" captionTargetPageId-2="5" captionText-0="Figure 1. - Lentipes mekonggaensis. A: Male; B: Female. (Pictures by R. Hadiaty)." captionText-1="Figure 2. - Diagrammatic illustration of head in Lentipes mekonggaensis (male) showing head pores and sensory papillae. A: Dorsal view; B: Lateral view." captionText-2="Figure 3.- Diagrammatic illustration of urogenital papilla in Lentipes mekonggaensis. A: Male; B: Female. 1: anus; 2: urogenital papilla; 3: anal fin." pageId="1" pageNumber="134">Figs 1-3</figureCitation>
<figureCitation id="1340F297310BFFA0FEEBFC58FE1A3C5F" box="[351,442,1000,1025]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="5.[132,195,484,506]" captionStartId-1="5.[132,196,1370,1392]" captionStartId-2="5.[813,877,1089,1111]" captionTargetBox-0="[132,1455,215,467]" captionTargetBox-1="[144,767,542,1336]" captionTargetBox-2="[825,1453,554,1063]" captionTargetId-0="figure-286@5.[132,1455,215,467]" captionTargetId-1="figure-305@5.[132,775,534,1353]" captionTargetId-2="figure-336@5.[813,1456,534,1072]" captionTargetPageId-0="5" captionTargetPageId-1="5" captionTargetPageId-2="5" captionText-0="Figure 1. - Lentipes mekonggaensis. A: Male; B: Female. (Pictures by R. Hadiaty)." captionText-1="Figure 2. - Diagrammatic illustration of head in Lentipes mekonggaensis (male) showing head pores and sensory papillae. A: Dorsal view; B: Lateral view." captionText-2="Figure 3.- Diagrammatic illustration of urogenital papilla in Lentipes mekonggaensis. A: Male; B: Female. 1: anus; 2: urogenital papilla; 3: anal fin." figureDoi-0="http://doi.org/10.5281/zenodo.13618364" figureDoi-1="http://doi.org/10.5281/zenodo.13618368" figureDoi-2="http://doi.org/10.5281/zenodo.13618366" httpUri-0="https://zenodo.org/record/13618364/files/figure.png" httpUri-1="https://zenodo.org/record/13618368/files/figure.png" httpUri-2="https://zenodo.org/record/13618366/files/figure.png" pageId="1" pageNumber="134">Figs 1-3</figureCitation>
, Tabs I-IV)
</paragraph>
</subSubSection>
@ -402,7 +410,7 @@ SL);
</paragraph>
</subSubSection>
<subSubSection id="C361BD99310BFFA0FF1EF949FC3C3A0A" pageId="1" pageNumber="134" type="description">
<paragraph id="8BC4EE12310BFFA0FF1EF949FCA73F4E" blockId="1.[132,776,1065,1907]" box="[170,775,1784,1809]" pageId="1" pageNumber="134">
<paragraph id="8BC4EE12310BFFA0FF1EF949FAFE394F" blockId="1.[132,776,1065,1907]" lastBlockId="1.[813,1456,217,596]" pageId="1" pageNumber="134">
<taxonomicName id="4C7B9591310BFFA0FF1EF949FEC63F4E" authorityName="Allen" authorityYear="2004" box="[170,358,1785,1808]" family="Gobiidae" genus="Lentipes" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="134" phylum="Chordata" rank="species" species="venustus">
<emphasis id="B90F3200310BFFA0FF1EF949FEC63F4E" box="[170,358,1785,1808]" italics="true" pageId="1" pageNumber="134">Lentipes venustus</emphasis>
</taxonomicName>
@ -410,9 +418,7 @@ SL);
<typeStatus id="54C050B0310BFFA0FDF0F949FD0D3F4F" box="[580,685,1785,1809]" pageId="1" pageNumber="134" type="paratype">paratypes</typeStatus>
)
<specimenCount id="9D7D259B310BFFA0FD0EF949FCA23F4E" box="[698,770,1784,1809]" count="1" pageId="1" pageNumber="134" type="male">1 male</specimenCount>
,
</paragraph>
<paragraph id="8BC4EE12310BFFA0FF30F8A9FAFE394F" blockId="1.[132,776,1065,1907]" lastBlockId="1.[813,1456,217,596]" pageId="1" pageNumber="134">
,
<materialsCitation id="3B13E44F310BFFA0FF30F8A9FB6A38AF" collectingDate="2003-01-30" collectorName="Allen &amp; Stevenson." country="Papua New Guinea" county="Normanby Island Bunama Village" latitude="-10.117784" location="Apatabuia River" longLatPrecision="9" longitude="150.152" municipality="D'Entrecasteaux Islands" pageId="1" pageNumber="134" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D7D259B310BFFA0FF30F8A9FF413F6F" box="[132,225,1817,1841]" count="1" pageId="1" pageNumber="134" type="female">1 female</specimenCount>
(24.0-
@ -457,15 +463,13 @@ MNHN, uncatalogued
</materialsCitation>
.
</paragraph>
<paragraph id="8BC4EE12310BFFA0FCE7FEAAFA0F396F" blockId="1.[813,1456,217,596]" box="[851,1455,281,306]" pageId="1" pageNumber="134">
<paragraph id="8BC4EE12310BFFA0FCE7FEAAFAFE3A4D" blockId="1.[813,1456,217,596]" pageId="1" pageNumber="134">
<taxonomicName id="4C7B9591310BFFA0FCE7FEAAFBFC396F" authorityName="Allen" authorityYear="2001" box="[851,1116,281,305]" family="Gobiidae" genus="Lentipes" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="134" phylum="Chordata" rank="species" species="multiradiatus">
<emphasis id="B90F3200310BFFA0FCE7FEAAFBFC396F" box="[851,1116,281,305]" italics="true" pageId="1" pageNumber="134">Lentipes multiradiatus</emphasis>
</taxonomicName>
. - WAM 32370.003,
<specimenCount id="9D7D259B310BFFA0FAEDFEAAFA0B396F" box="[1369,1451,281,306]" count="1" pageId="1" pageNumber="134" type="male">1 male</specimenCount>
,
</paragraph>
<paragraph id="8BC4EE12310BFFA0FC99FE8AFAFE3A4D" blockId="1.[813,1456,217,596]" pageId="1" pageNumber="134">
,
<materialsCitation id="3B13E44F310BFFA0FC99FE8AFB7A39ED" collectingDate="2003-01-27" collectorName="Allen &amp; Stevenson." country="Papua New Guinea" county="Fergusson Island" latitude="-9.515117" location="Awaetowa River" longLatPrecision="9" longitude="150.86734" municipality="D'Entrecasteaux Islands" pageId="1" pageNumber="134" specimenCount="3" specimenCount-female="3">
<specimenCount id="9D7D259B310BFFA0FC99FE8AFC07390C" box="[813,935,314,338]" count="3" pageId="1" pageNumber="134" type="female">3 females</specimenCount>
(30.0-37.0 mm SL);
@ -624,8 +628,7 @@ MNHN 2013-0652,
<paragraph id="8BC4EE12310BFFA0FC99FB31FC3B3CC7" blockId="1.[813,1455,1153,1241]" box="[813,923,1153,1177]" pageId="1" pageNumber="134">
<emphasis id="B90F3200310BFFA0FC99FB31FC3B3CC7" bold="true" box="[813,923,1153,1177]" pageId="1" pageNumber="134">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BC4EE12310BFFA3FCE6FB11FD6D396F" blockId="1.[813,1455,1153,1241]" lastBlockId="2.[132,717,217,466]" lastPageId="2" lastPageNumber="135" pageId="1" pageNumber="134">The new species has 19-20 pectoral rays, a second dorsal and anal fins I10, and 28-33 lateral scales. The urogenital papilla is retractable into a sheath-like groove and is without lobes or other expanded tissue. The male is characterised by few tricuspid teeth in the upper jaw</paragraph>
<paragraph id="8BC4EE123108FFA3FF30FE89FDA1398F" blockId="2.[132,717,217,466]" pageId="2" pageNumber="135">(10-16), ctenoid scales on anterior body region strongly ossified, the base of the first dorsal fin not reaching the base of the second dorsal fin origin, and a specific body colour, with a bright red head, a red band on caudal peduncle and orange dorsal fins.</paragraph>
<paragraph id="8BC4EE12310BFFA3FCE6FB11FDA1398F" blockId="1.[813,1455,1153,1241]" lastBlockId="2.[132,717,217,466]" lastPageId="2" lastPageNumber="135" pageId="1" pageNumber="134">The new species has 19-20 pectoral rays, a second dorsal and anal fins I10, and 28-33 lateral scales. The urogenital papilla is retractable into a sheath-like groove and is without lobes or other expanded tissue. The male is characterised by few tricuspid teeth in the upper jaw (10-16), ctenoid scales on anterior body region strongly ossified, the base of the first dorsal fin not reaching the base of the second dorsal fin origin, and a specific body colour, with a bright red head, a red band on caudal peduncle and orange dorsal fins.</paragraph>
<tableNote id="769DEF9C3108FFA3FF30FE49FF713AAF" pageId="2" pageNumber="135" targetBox="[765,2434,750,1413]" targetPageId="2">
<paragraph id="8BC4EE123108FFA3FF30FE49FEA83A4F" blockId="2.[132,1405,505,753]" box="[132,264,505,529]" pageId="2" pageNumber="135">
<emphasis id="B90F32003108FFA3FF30FE49FEA83A4F" bold="true" box="[132,264,505,529]" pageId="2" pageNumber="135">Description</emphasis>
@ -909,13 +912,9 @@ females
</tr>
</table>
</paragraph>
<paragraph id="8BC4EE123108FFA3FF1EFB2AFD6D3D2C" blockId="2.[132,718,762,1394]" pageId="2" pageNumber="135">Lateral scales (LS) 32(28-33). No difference in scale number and arrangement between sexes. They are lightly embedded and mainly cycloid in females. Generally limited to caudal peduncle, few may extend anteriorly along midline between second dorsal and anal fins; some ctenoid scales are on the anteriormost part of the flanks. Males have mainly ctenoid scales,</paragraph>
<paragraph id="8BC4EE123108FFA3FF30FACAFD6D3D8D" blockId="2.[132,718,1402,1907]" pageId="2" pageNumber="135">strongly developed with prominent spines on anterior body region, and few cycloid scales on caudal peduncle. Scales in transverse backwards (TRB) series 11(11-</paragraph>
<paragraph id="8BC4EE123108FFA3FF30FA6BFD6D3DAD" blockId="2.[132,718,1402,1907]" box="[132,717,1498,1523]" pageId="2" pageNumber="135">13) and in transverse forward series (TRF) 10(7-12).</paragraph>
<paragraph id="8BC4EE123108FFA3FF30FA4AFD6D3EAC" blockId="2.[132,718,1402,1907]" pageId="2" pageNumber="135">Zigzag scales (ZZ) 12(9-12). Head, breast, nape and belly without scales. Upper jaw teeth distinctly tricuspid anteriorly, males 10(10-16), females (24-32). Premaxilla in males with 3(2-6) recurved canines posterior to tricuspid teeth; females without teeth posterior to tricuspid teeth. Teeth in lower jaw recurved and canine in males 3(3-5); no teeth in females. Cephalic sensory pore system A, B, C, D, F, H, K, L, N and O; pore H</paragraph>
<paragraph id="8BC4EE123108FFA3FF30F94BFDF43F2D" blockId="2.[132,718,1402,1907]" pageId="2" pageNumber="135">
&amp; K sometimes fused. Pore D singular with all others paired (
<figureCitation id="1340F2973108FFA3FF63F8ABFEBA3F6D" box="[215,282,1819,1843]" captionStart="Figure 2" captionStartId="5.[132,196,1370,1392]" captionTargetBox="[144,767,542,1336]" captionTargetId="figure-305@5.[132,775,534,1353]" captionTargetPageId="5" captionText="Figure 2. - Diagrammatic illustration of head in Lentipes mekonggaensis (male) showing head pores and sensory papillae. A: Dorsal view; B: Lateral view." pageId="2" pageNumber="135">Fig. 2</figureCitation>
<paragraph id="8BC4EE123108FFA3FF1EFB2AFDF43F2D" blockId="2.[132,718,762,1394]" lastBlockId="2.[132,718,1402,1907]" pageId="2" pageNumber="135">
Lateral scales (LS) 32(28-33). No difference in scale number and arrangement between sexes. They are lightly embedded and mainly cycloid in females. Generally limited to caudal peduncle, few may extend anteriorly along midline between second dorsal and anal fins; some ctenoid scales are on the anteriormost part of the flanks. Males have mainly ctenoid scales, strongly developed with prominent spines on anterior body region, and few cycloid scales on caudal peduncle. Scales in transverse backwards (TRB) series 11(11- 13) and in transverse forward series (TRF) 10(7-12). Zigzag scales (ZZ) 12(9-12). Head, breast, nape and belly without scales. Upper jaw teeth distinctly tricuspid anteriorly, males 10(10-16), females (24-32). Premaxilla in males with 3(2-6) recurved canines posterior to tricuspid teeth; females without teeth posterior to tricuspid teeth. Teeth in lower jaw recurved and canine in males 3(3-5); no teeth in females. Cephalic sensory pore system A, B, C, D, F, H, K, L, N and O; pore H &amp; K sometimes fused. Pore D singular with all others paired (
<figureCitation id="1340F2973108FFA3FF63F8ABFEBA3F6D" box="[215,282,1819,1843]" captionStart="Figure 2" captionStartId="5.[132,196,1370,1392]" captionTargetBox="[144,767,542,1336]" captionTargetId="figure-305@5.[132,775,534,1353]" captionTargetPageId="5" captionText="Figure 2. - Diagrammatic illustration of head in Lentipes mekonggaensis (male) showing head pores and sensory papillae. A: Dorsal view; B: Lateral view." figureDoi="http://doi.org/10.5281/zenodo.13618368" httpUri="https://zenodo.org/record/13618368/files/figure.png" pageId="2" pageNumber="135">Fig. 2</figureCitation>
); oculoscapular canal divided into anterior and posterior canal between pores H and K. Some cutaneous sensory papillae present on head.
</paragraph>
<caption id="DF04BE9A3109FFA2FF30FF69FD0838B1" box="[132,680,217,239]" pageId="3" pageNumber="136" startId="3.[132,185,217,239]" targetType="table">
@ -1872,9 +1871,9 @@ females
</paragraph>
<paragraph id="8BC4EE12310EFFA5FF1EFB22FE2E3DB0" blockId="4.[132,775,1170,1518]" pageId="4" pageNumber="137">
Sexual dimorphism well developed. Ctenoid scales on anterior body region strongly ossified in males, each with 3-5 prominent spines; scales on posterior part of body with fewer, but larger ctenii than those of females. Urogenital papilla in males slender and pointed distally without associated lobes or expanded tissue (
<figureCitation id="1340F297310EFFA5FE6CFA84FD8C3D12" box="[472,556,1332,1356]" captionStart="Figure 3" captionStartId="5.[813,877,1089,1111]" captionTargetBox="[825,1453,554,1063]" captionTargetId="figure-336@5.[813,1456,534,1072]" captionTargetPageId="5" captionText="Figure 3.- Diagrammatic illustration of urogenital papilla in Lentipes mekonggaensis. A: Male; B: Female. 1: anus; 2: urogenital papilla; 3: anal fin." pageId="4" pageNumber="137">Fig. 3A</figureCitation>
<figureCitation id="1340F297310EFFA5FE6CFA84FD8C3D12" box="[472,556,1332,1356]" captionStart="Figure 3" captionStartId="5.[813,877,1089,1111]" captionTargetBox="[825,1453,554,1063]" captionTargetId="figure-336@5.[813,1456,534,1072]" captionTargetPageId="5" captionText="Figure 3.- Diagrammatic illustration of urogenital papilla in Lentipes mekonggaensis. A: Male; B: Female. 1: anus; 2: urogenital papilla; 3: anal fin." figureDoi="http://doi.org/10.5281/zenodo.13618366" httpUri="https://zenodo.org/record/13618366/files/figure.png" pageId="4" pageNumber="137">Fig. 3A</figureCitation>
), urogenital papilla retractable into a sheath-like groove; female urogenital papilla rectangular in appearance (
<figureCitation id="1340F297310EFFA5FE4CFAC5FDEA3DD2" box="[504,586,1397,1421]" captionStart="Figure 3" captionStartId="5.[813,877,1089,1111]" captionTargetBox="[825,1453,554,1063]" captionTargetId="figure-336@5.[813,1456,534,1072]" captionTargetPageId="5" captionText="Figure 3.- Diagrammatic illustration of urogenital papilla in Lentipes mekonggaensis. A: Male; B: Female. 1: anus; 2: urogenital papilla; 3: anal fin." pageId="4" pageNumber="137">Fig. 3B</figureCitation>
<figureCitation id="1340F297310EFFA5FE4CFAC5FDEA3DD2" box="[504,586,1397,1421]" captionStart="Figure 3" captionStartId="5.[813,877,1089,1111]" captionTargetBox="[825,1453,554,1063]" captionTargetId="figure-336@5.[813,1456,534,1072]" captionTargetPageId="5" captionText="Figure 3.- Diagrammatic illustration of urogenital papilla in Lentipes mekonggaensis. A: Male; B: Female. 1: anus; 2: urogenital papilla; 3: anal fin." figureDoi="http://doi.org/10.5281/zenodo.13618366" httpUri="https://zenodo.org/record/13618366/files/figure.png" pageId="4" pageNumber="137">Fig. 3B</figureCitation>
) and also retractable into a sheath-like groove. Jaw length, head length and caudal peduncle length greater in males. Dorsal, caudal and anal fins longer in males.
</paragraph>
<paragraph id="8BC4EE12310EFFA5FF30F9A6FE253E70" blockId="4.[132,776,1558,1907]" box="[132,389,1558,1582]" pageId="4" pageNumber="137">
@ -1891,7 +1890,7 @@ Sexual dimorphism well developed. Ctenoid scales on anterior body region strongl
<paragraph id="8BC4EE12310EFFA5FC99FA44FBB93E52" blockId="4.[813,1455,1524,1907]" box="[813,1049,1524,1548]" pageId="4" pageNumber="137">
<emphasis id="B90F3200310EFFA5FC99FA44FBB93E52" bold="true" box="[813,1049,1524,1548]" pageId="4" pageNumber="137">
Colour in life (
<figureCitation id="1340F297310EFFA5FC64FA44FBAF3E52" box="[976,1039,1524,1548]" captionStart="Figure 1" captionStartId="5.[132,195,484,506]" captionTargetBox="[132,1455,215,467]" captionTargetId="figure-286@5.[132,1455,215,467]" captionTargetPageId="5" captionText="Figure 1. - Lentipes mekonggaensis. A: Male; B: Female. (Pictures by R. Hadiaty)." pageId="4" pageNumber="137">Fig. 1</figureCitation>
<figureCitation id="1340F297310EFFA5FC64FA44FBAF3E52" box="[976,1039,1524,1548]" captionStart="Figure 1" captionStartId="5.[132,195,484,506]" captionTargetBox="[132,1455,215,467]" captionTargetId="figure-286@5.[132,1455,215,467]" captionTargetPageId="5" captionText="Figure 1. - Lentipes mekonggaensis. A: Male; B: Female. (Pictures by R. Hadiaty)." figureDoi="http://doi.org/10.5281/zenodo.13618364" httpUri="https://zenodo.org/record/13618364/files/figure.png" pageId="4" pageNumber="137">Fig. 1</figureCitation>
)
</emphasis>
</paragraph>
@ -1900,7 +1899,7 @@ Colour in life (
- Background of body greyish. Background of head and snout reddish. Dorsal margin of head greyish, upper lip red. Head ventrally whitish. Lateral midline without a well-marked black subcutaneous band. All ctenoid scales on flanks and caudal peduncle with black margins.
</paragraph>
<paragraph id="8BC4EE12310EFFA5FCE6F97CFCFE3F2D" blockId="4.[813,1455,1524,1907]" pageId="4" pageNumber="137">Bright orange dorsal fins, with a blue-edged black spot in middle of membrane between first two dorsal rays of the second dorsal fin. Caudal and pectoral fins translucent. Anal fin bluish to green in the anterior part and red in the posterior part.</paragraph>
<caption id="DF04BE9A310FFFA4FF30FE54FC3939A4" box="[132,921,484,506]" pageId="5" pageNumber="138" startId="5.[132,195,484,506]" targetBox="[132,1455,215,467]" targetPageId="5" targetType="figure">
<caption id="DF04BE9A310FFFA4FF30FE54FC3939A4" ID-DOI="http://doi.org/10.5281/zenodo.13618364" ID-Zenodo-Dep="13618364" box="[132,921,484,506]" httpUri="https://zenodo.org/record/13618364/files/figure.png" pageId="5" pageNumber="138" startId="5.[132,195,484,506]" targetBox="[132,1455,215,467]" targetPageId="5" targetType="figure">
<paragraph id="8BC4EE12310FFFA4FF30FE54FC3939A4" blockId="5.[132,921,484,506]" box="[132,921,484,506]" pageId="5" pageNumber="138">
Figure 1. -
<taxonomicName id="4C7B9591310FFFA4FF59FE55FE7739A7" authorityName="Keith &amp; Hadiaty &amp; Hubert &amp; Busson &amp; Lord" authorityYear="2014" box="[237,471,484,506]" family="Gobiidae" genus="Lentipes" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="138" phylum="Chordata" rank="species" species="mekonggaensis">
@ -1913,7 +1912,7 @@ Figure 1. -
: Female. (Pictures by R. Hadiaty).
</paragraph>
</caption>
<caption id="DF04BE9A310FFFA4FC99FBF1FC403CD9" pageId="5" pageNumber="138" startId="5.[813,877,1089,1111]" targetBox="[825,1453,554,1063]" targetPageId="5" targetType="figure">
<caption id="DF04BE9A310FFFA4FC99FBF1FC403CD9" ID-DOI="http://doi.org/10.5281/zenodo.13618366" ID-Zenodo-Dep="13618366" httpUri="https://zenodo.org/record/13618366/files/figure.png" pageId="5" pageNumber="138" startId="5.[813,877,1089,1111]" targetBox="[825,1453,554,1063]" targetPageId="5" targetType="figure">
<paragraph id="8BC4EE12310FFFA4FC99FBF1FC403CD9" blockId="5.[813,1455,1089,1159]" pageId="5" pageNumber="138">
Figure 3.- Diagrammatic illustration of urogenital papilla in
<taxonomicName id="4C7B9591310FFFA4FA36FBF2FC5A3C30" authorityName="Keith &amp; Hadiaty &amp; Hubert &amp; Busson &amp; Lord" authorityYear="2014" family="Gobiidae" genus="Lentipes" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="138" phylum="Chordata" rank="species" species="mekonggaensis">
@ -1926,7 +1925,7 @@ Figure 3.- Diagrammatic illustration of urogenital papilla in
: Female. 1: anus; 2: urogenital papilla; 3: anal fin.
</paragraph>
</caption>
<caption id="DF04BE9A310FFFA4FF30FAEAFEF93DFE" pageId="5" pageNumber="138" startId="5.[132,196,1370,1392]" targetBox="[144,767,542,1336]" targetPageId="5" targetType="figure">
<caption id="DF04BE9A310FFFA4FF30FAEAFEF93DFE" ID-DOI="http://doi.org/10.5281/zenodo.13618368" ID-Zenodo-Dep="13618368" httpUri="https://zenodo.org/record/13618368/files/figure.png" pageId="5" pageNumber="138" startId="5.[132,196,1370,1392]" targetBox="[144,767,542,1336]" targetPageId="5" targetType="figure">
<paragraph id="8BC4EE12310FFFA4FF30FAEAFEF93DFE" blockId="5.[132,775,1370,1440]" pageId="5" pageNumber="138">
Figure 2. - Diagrammatic illustration of head in
<taxonomicName id="4C7B9591310FFFA4FDEEFAEBFF6C3DD6" authorityName="Keith &amp; Hadiaty &amp; Hubert &amp; Busson &amp; Lord" authorityYear="2014" family="Gobiidae" genus="Lentipes" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="138" phylum="Chordata" rank="species" species="mekonggaensis">
@ -2062,7 +2061,7 @@ in having fewer scales in lateral series (28-33
30-34).
</paragraph>
</subSubSection>
<caption id="DF04BE9A310CFFA7FF30FDF0FCE13A08" box="[132,833,576,598]" pageId="6" pageNumber="139" startId="6.[132,195,576,598]" targetBox="[132,1455,215,559]" targetPageId="6" targetType="figure">
<caption id="DF04BE9A310CFFA7FF30FDF0FCE13A08" ID-DOI="http://doi.org/10.5281/zenodo.13618372" ID-Zenodo-Dep="13618372" box="[132,833,576,598]" httpUri="https://zenodo.org/record/13618372/files/figure.png" pageId="6" pageNumber="139" startId="6.[132,195,576,598]" targetBox="[132,1455,215,559]" targetPageId="6" targetType="figure">
<paragraph id="8BC4EE12310CFFA7FF30FDF0FCE13A08" blockId="6.[132,833,576,598]" box="[132,833,576,598]" pageId="6" pageNumber="139">
Figure 4. -
<emphasis id="B90F3200310CFFA7FF59FDF1FE093A08" box="[237,425,577,598]" italics="true" pageId="6" pageNumber="139">

View file

@ -1,61 +1,69 @@
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@ -73,7 +81,7 @@ Keith, Hadiaty &amp; Lord
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(
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<figureCitation id="1340F297310CFFA7FBB4FD28FBFA3AEE" box="[1024,1114,664,688]" captionStart-0="Figure 4" captionStart-1="Figure 5" captionStart-2="Figure 6" captionStartId-0="6.[132,195,576,598]" captionStartId-1="6.[132,195,1837,1859]" captionStartId-2="6.[813,877,1837,1859]" captionTargetBox-0="[132,1455,215,559]" captionTargetBox-1="[143,769,1009,1807]" captionTargetBox-2="[825,1453,1303,1813]" captionTargetId-0="figure-253@6.[132,1455,215,559]" captionTargetId-1="figure-270@6.[132,775,998,1820]" captionTargetId-2="figure-299@6.[813,1456,1280,1820]" captionTargetPageId-0="6" captionTargetPageId-1="6" captionTargetPageId-2="6" captionText-0="Figure 4. - Lentipes argenteus. A: Male; B: Female. (Pictures by P. Keith)" captionText-1="Figure 5. - Diagrammatic illustration of head in Lentipes argenteus (male) showing head pores and sensory papillae. A: Dorsal view; B: Lateral view." captionText-2="Figure 6. - Diagrammatic illustration of urogenital papilla in Lentipes argenteus. A: Male; B: Female. 1: anus; 2: urogenital papilla; 3: anal fin." figureDoi-0="http://doi.org/10.5281/zenodo.13618372" figureDoi-1="http://doi.org/10.5281/zenodo.13618374" figureDoi-2="http://doi.org/10.5281/zenodo.13618376" httpUri-0="https://zenodo.org/record/13618372/files/figure.png" httpUri-1="https://zenodo.org/record/13618374/files/figure.png" httpUri-2="https://zenodo.org/record/13618376/files/figure.png" pageId="6" pageNumber="139">Figs 4-6</figureCitation>
, Tabs V-VII)
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@ -104,16 +112,13 @@ and
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, this paper.
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<emphasis id="B90F3200310CFFA7FCE6FC69FBE23BAE" box="[850,1090,984,1008]" italics="true" pageId="6" pageNumber="139">Lentipes dimetrodon</emphasis>
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. - MZB 8001 (
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(
), male (
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SL);
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@ -170,7 +175,7 @@ and not WAM P.31059- 002 as cited by mistake in the original description [
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Figure 5. - Diagrammatic illustration of head in
<taxonomicName id="4C7B9591310CFFA7FDE5F89DFCA73F1C" authorityName="Keith &amp; Hadiaty &amp; Hubert &amp; Busson &amp; Lord" authorityYear="2014" box="[593,775,1837,1858]" family="Gobiidae" genus="Lentipes" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="139" phylum="Chordata" rank="species" species="argenteus">
@ -183,7 +188,7 @@ Figure 5. - Diagrammatic illustration of head in
: Lateral view.
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Figure 6. - Diagrammatic illustration of urogenital papilla in
<taxonomicName id="4C7B9591310CFFA7FA36F89DFC1E3F04" authorityName="Keith &amp; Hadiaty &amp; Hubert &amp; Busson &amp; Lord" authorityYear="2014" family="Gobiidae" genus="Lentipes" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="139" phylum="Chordata" rank="species" species="argenteus">
@ -510,15 +515,13 @@ Table VI. - Meristic counts in studied species of
</tr>
</table>
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<emphasis id="B90F32003102FFA9FF1EFEAAFEFE396F" box="[170,350,282,305]" italics="true" pageId="8" pageNumber="141">Lentipes watsoni</emphasis>
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. - WAM P.31221-001 (
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), male,
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SL,
@ -1255,7 +1258,7 @@ First dorsal fin (D1) with 5-6 flexible spines, second dorsal fin (D2) with one
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and 5
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long- er. Pelvic fins constitute a strong adhesive disc adherent to abdomen between all five rays. Pectoral fin with 16-17 rays, ventralmost 1
longer. Pelvic fins constitute a strong adhesive disc adherent to abdomen between all five rays. Pectoral fin with 16-17 rays, ventralmost 1
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or 2
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@ -1267,14 +1270,14 @@ Lateral scales (LS) 36(35-49). They are lightly embedded and mainly cycloid in f
with M, N and O; pore D missing in
<specimenCount id="9D7D259B3103FFA8FC16FB4BFBA63D4C" box="[930,1030,1274,1298]" count="1" pageId="9" pageNumber="142" type="male">one male</specimenCount>
, but when present singular with all others paired (
<figureCitation id="1340F2973103FFA8FC10FAAAFC453D6C" box="[932,997,1306,1330]" captionStart="Figure 5" captionStartId="6.[132,195,1837,1859]" captionTargetBox="[143,769,1009,1807]" captionTargetId="figure-270@6.[132,775,998,1820]" captionTargetPageId="6" captionText="Figure 5. - Diagrammatic illustration of head in Lentipes argenteus (male) showing head pores and sensory papillae. A: Dorsal view; B: Lateral view." pageId="9" pageNumber="142">Fig. 5</figureCitation>
<figureCitation id="1340F2973103FFA8FC10FAAAFC453D6C" box="[932,997,1306,1330]" captionStart="Figure 5" captionStartId="6.[132,195,1837,1859]" captionTargetBox="[143,769,1009,1807]" captionTargetId="figure-270@6.[132,775,998,1820]" captionTargetPageId="6" captionText="Figure 5. - Diagrammatic illustration of head in Lentipes argenteus (male) showing head pores and sensory papillae. A: Dorsal view; B: Lateral view." figureDoi="http://doi.org/10.5281/zenodo.13618374" httpUri="https://zenodo.org/record/13618374/files/figure.png" pageId="9" pageNumber="142">Fig. 5</figureCitation>
); oculoscapular canal divided into anterior and posterior canal between pores H and K. Some cutaneous sensory papillae present on head and body.
</paragraph>
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Sexual dimorphism well developed. Ctenoid scales on anterior body region ossified in males, each with generally three prominent spines; scales on posterior part of body with fewer, but larger ctenii than those of females. Urogenital papilla in males slender and pointed distally without associated lobes or expanded tissue (
<figureCitation id="1340F2973103FFA8FB34F9AAFB753E6D" box="[1152,1237,1562,1587]" captionStart="Figure 6" captionStartId="6.[813,877,1837,1859]" captionTargetBox="[825,1453,1303,1813]" captionTargetId="figure-299@6.[813,1456,1280,1820]" captionTargetPageId="6" captionText="Figure 6. - Diagrammatic illustration of urogenital papilla in Lentipes argenteus. A: Male; B: Female. 1: anus; 2: urogenital papilla; 3: anal fin." pageId="9" pageNumber="142">Fig. 6A</figureCitation>
<figureCitation id="1340F2973103FFA8FB34F9AAFB753E6D" box="[1152,1237,1562,1587]" captionStart="Figure 6" captionStartId="6.[813,877,1837,1859]" captionTargetBox="[825,1453,1303,1813]" captionTargetId="figure-299@6.[813,1456,1280,1820]" captionTargetPageId="6" captionText="Figure 6. - Diagrammatic illustration of urogenital papilla in Lentipes argenteus. A: Male; B: Female. 1: anus; 2: urogenital papilla; 3: anal fin." figureDoi="http://doi.org/10.5281/zenodo.13618376" httpUri="https://zenodo.org/record/13618376/files/figure.png" pageId="9" pageNumber="142">Fig. 6A</figureCitation>
), urogenital papilla retractable into a sheath-like groove; female urogenital papilla rectangular in appearance (
<figureCitation id="1340F2973103FFA8FB15F9EAFB523E2C" box="[1185,1266,1626,1650]" captionStart="Figure 6" captionStartId="6.[813,877,1837,1859]" captionTargetBox="[825,1453,1303,1813]" captionTargetId="figure-299@6.[813,1456,1280,1820]" captionTargetPageId="6" captionText="Figure 6. - Diagrammatic illustration of urogenital papilla in Lentipes argenteus. A: Male; B: Female. 1: anus; 2: urogenital papilla; 3: anal fin." pageId="9" pageNumber="142">Fig. 6B</figureCitation>
<figureCitation id="1340F2973103FFA8FB15F9EAFB523E2C" box="[1185,1266,1626,1650]" captionStart="Figure 6" captionStartId="6.[813,877,1837,1859]" captionTargetBox="[825,1453,1303,1813]" captionTargetId="figure-299@6.[813,1456,1280,1820]" captionTargetPageId="6" captionText="Figure 6. - Diagrammatic illustration of urogenital papilla in Lentipes argenteus. A: Male; B: Female. 1: anus; 2: urogenital papilla; 3: anal fin." figureDoi="http://doi.org/10.5281/zenodo.13618376" httpUri="https://zenodo.org/record/13618376/files/figure.png" pageId="9" pageNumber="142">Fig. 6B</figureCitation>
) and also retractable into a sheath-like groove. Second dorsal, anal and caudal fin lengths, jaw length and caudal peduncle depth greater in males.
</paragraph>
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@ -1284,7 +1287,7 @@ Sexual dimorphism well developed. Ctenoid scales on anterior body region ossifie
<emphasis id="B90F32003103FFA8FCE6F8AAFC343F6C" box="[850,916,1818,1842]" italics="true" pageId="9" pageNumber="142">Male.</emphasis>
- Background of body greyish. Background of head greyish. Head ventrally greyish. Lateral midline with a marked black subcutaneous band. All scales on flanks and caudal peduncle with black margins. Top of head blackish. Nape greyish. Caudal peduncle pinkish to reddish. Caudal fin rays greyish. Dorsal and anal fins blackish. Pelvic disk without pigment. Pectoral rays greyish. Pectoral fin base greyish.
</paragraph>
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<caption id="DF04BE9A3100FFABFF30FDD0FDAD3AD3" ID-DOI="http://doi.org/10.5281/zenodo.13618380" ID-Zenodo-Dep="13618380" httpUri="https://zenodo.org/record/13618380/files/figure.png" pageId="10" pageNumber="143" startId="10.[132,195,608,630]" targetBox="[132,1455,215,591]" targetPageId="10" targetType="figure">
<paragraph id="8BC4EE123100FFABFF30FDD0FDAD3AD3" blockId="10.[132,1456,608,654]" pageId="10" pageNumber="143">
Figure 7. -
<emphasis id="B90F32003100FFABFF58FDD1FEDD3A2B" box="[236,381,608,630]" italics="true" pageId="10" pageNumber="143">
@ -1304,7 +1307,7 @@ Figure 7. -
<paragraph id="8BC4EE123100FFABFF30FBC9FED03CCF" blockId="10.[132,775,1145,1586]" box="[132,368,1145,1169]" pageId="10" pageNumber="143">
<emphasis id="B90F32003100FFABFF30FBC9FED03CCF" bold="true" box="[132,368,1145,1169]" pageId="10" pageNumber="143">
Colour in life (
<figureCitation id="1340F2973100FFABFE93FBC9FEC83CCF" box="[295,360,1145,1169]" captionStart="Figure 4" captionStartId="6.[132,195,576,598]" captionTargetBox="[132,1455,215,559]" captionTargetId="figure-253@6.[132,1455,215,559]" captionTargetPageId="6" captionText="Figure 4. - Lentipes argenteus. A: Male; B: Female. (Pictures by P. Keith)" pageId="10" pageNumber="143">Fig. 4</figureCitation>
<figureCitation id="1340F2973100FFABFE93FBC9FEC83CCF" box="[295,360,1145,1169]" captionStart="Figure 4" captionStartId="6.[132,195,576,598]" captionTargetBox="[132,1455,215,559]" captionTargetId="figure-253@6.[132,1455,215,559]" captionTargetPageId="6" captionText="Figure 4. - Lentipes argenteus. A: Male; B: Female. (Pictures by P. Keith)" figureDoi="http://doi.org/10.5281/zenodo.13618372" httpUri="https://zenodo.org/record/13618372/files/figure.png" pageId="10" pageNumber="143">Fig. 4</figureCitation>
)
</emphasis>
</paragraph>
@ -1402,7 +1405,7 @@ the 5
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and 6
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spines of D2 long- er, and the coloration in males. Furthermore, it differs from
spines of D2 longer, and the coloration in males. Furthermore, it differs from
<taxonomicName id="4C7B95913100FFABFC99FBCAFC023CCF" authorityName="Allen" authorityYear="1997" box="[813,930,1146,1169]" family="Gobiidae" genus="Lentipes" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="143" phylum="Chordata" rank="species" species="watsoni">
<emphasis id="B90F32003100FFABFC99FBCAFC023CCF" box="[813,930,1146,1169]" italics="true" pageId="10" pageNumber="143">L. watsoni</emphasis>
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View file

@ -1,66 +1,67 @@
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<mods:identifier id="BD48928FBDCC136036FE9162ADA6FA19" type="DOI">10.11646/zootaxa.5497.2.4</mods:identifier>
<mods:identifier id="92225144E0799F366F14423F007CF96D" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03D98784FFA2FFA0FF0F13F02FE98E22" ID-DOI="http://doi.org/10.5281/zenodo.13618352" ID-Zenodo-Dep="13618352" LSID="urn:lsid:plazi:treatment:03D98784FFA2FFA0FF0F13F02FE98E22" httpUri="http://treatment.plazi.org/id/03D98784FFA2FFA0FF0F13F02FE98E22" lastPageNumber="247" pageId="3" pageNumber="247">
<subSubSection id="C36A6519FFA2FFA0FF0F13F02F9589F2" box="[151,746,223,250]" pageId="3" pageNumber="247" type="nomenclature">
<paragraph id="8BCF3692FFA2FFA0FF0F13F02F9589F2" blockId="3.[151,746,223,285]" box="[151,746,223,250]" pageId="3" pageNumber="247">
<heading id="D08781FEFFA2FFA0FF0F13F02F9589F2" bold="true" box="[151,746,223,250]" fontSize="11" level="1" pageId="3" pageNumber="247" reason="1">
@ -80,9 +81,9 @@ Liu
<subSubSection id="C36A6519FFA2FFA0FF0F122C2C3F8816" box="[151,320,259,285]" pageId="3" pageNumber="247" type="description">
<paragraph id="8BCF3692FFA2FFA0FF0F122C2C3F8816" blockId="3.[151,746,223,285]" box="[151,320,259,285]" pageId="3" pageNumber="247">
(
<figureCitation id="134B2A17FFA2FFA0FF07122C2C628816" box="[159,285,259,285]" captionStart="FIGURE 1" captionStartId="2.[152,255,1874,1898]" captionTargetBox="[226,1378,228,1811]" captionTargetId="figure-17@2.[151,1436,179,1848]" captionTargetPageId="2" captionText="FIGURE 1. Habitus views of S. qinghaiensis sp. nov. (A) female; (B) male and S. shanxiensis sp. nov. (C) female; (D) male." pageId="3" pageNumber="247">Figs 1AB</figureCitation>
<figureCitation id="134B2A17FFA2FFA0FF07122C2C628816" box="[159,285,259,285]" captionStart="FIGURE 1" captionStartId="2.[152,255,1874,1898]" captionTargetBox="[226,1378,228,1811]" captionTargetId="figure-17@2.[151,1436,179,1848]" captionTargetPageId="2" captionText="FIGURE 1. Habitus views of S. qinghaiensis sp. nov. (A) female; (B) male and S. shanxiensis sp. nov. (C) female; (D) male." figureDoi="http://doi.org/10.5281/zenodo.13618198" httpUri="https://zenodo.org/record/13618198/files/figure.png" pageId="3" pageNumber="247">Figs 1AB</figureCitation>
,
<figureCitation id="134B2A17FFA2FFA0FEB1122C2C478816" box="[297,312,259,285]" captionStart="FIGURE 2" captionStartId="4.[152,255,1893,1917]" captionTargetBox="[203,1394,219,1821]" captionTargetId="figure-17@4.[151,1436,179,1868]" captionTargetPageId="4" captionText="FIGURE 2. Genitalia of S. qinghaiensis sp. nov. (A) left paramere; (A') top view of left paramere; (B) right paramere; (B) top view of right paramere; (C) aedeagus in ventral views; (DE) endosoma with detached phallotheca in two different views." pageId="3" pageNumber="247">2</figureCitation>
<figureCitation id="134B2A17FFA2FFA0FEB1122C2C478816" box="[297,312,259,285]" captionStart="FIGURE 2" captionStartId="4.[152,255,1893,1917]" captionTargetBox="[203,1394,219,1821]" captionTargetId="figure-17@4.[151,1436,179,1868]" captionTargetPageId="4" captionText="FIGURE 2. Genitalia of S. qinghaiensis sp. nov. (A) left paramere; (A') top view of left paramere; (B) right paramere; (B) top view of right paramere; (C) aedeagus in ventral views; (DE) endosoma with detached phallotheca in two different views." figureDoi="http://doi.org/10.5281/zenodo.13618200" httpUri="https://zenodo.org/record/13618200/files/figure.png" pageId="3" pageNumber="247">2</figureCitation>
)
</paragraph>
</subSubSection>
@ -90,9 +91,9 @@ Liu
<paragraph id="8BCF3692FFA2FFA0FF0F12642F2488FE" blockId="3.[151,1437,331,1833]" pageId="3" pageNumber="247">
<emphasis id="B904EA80FFA2FFA0FF0F12642C6D886E" bold="true" box="[151,274,331,357]" pageId="3" pageNumber="247">Diagnosis.</emphasis>
Recognized by the following combination of characters: relatively short (male 2.34 2.47, female 2.67); dorsum with long, erect, dark, bristle-like setae; head black, with two smooth dark black spots on the inner margins of the eyes; pronotum and hemelytron uniformly dark; second antenna segment cylindrical, not dilated apically in both sexes (
<figureCitation id="134B2A17FFA2FFA0FEDB12982CBD88DA" box="[323,450,439,465]" captionStart="FIGURE 1" captionStartId="2.[152,255,1874,1898]" captionTargetBox="[226,1378,228,1811]" captionTargetId="figure-17@2.[151,1436,179,1848]" captionTargetPageId="2" captionText="FIGURE 1. Habitus views of S. qinghaiensis sp. nov. (A) female; (B) male and S. shanxiensis sp. nov. (C) female; (D) male." pageId="3" pageNumber="247">Fig. 1AB</figureCitation>
<figureCitation id="134B2A17FFA2FFA0FEDB12982CBD88DA" box="[323,450,439,465]" captionStart="FIGURE 1" captionStartId="2.[152,255,1874,1898]" captionTargetBox="[226,1378,228,1811]" captionTargetId="figure-17@2.[151,1436,179,1848]" captionTargetPageId="2" captionText="FIGURE 1. Habitus views of S. qinghaiensis sp. nov. (A) female; (B) male and S. shanxiensis sp. nov. (C) female; (D) male." figureDoi="http://doi.org/10.5281/zenodo.13618198" httpUri="https://zenodo.org/record/13618198/files/figure.png" pageId="3" pageNumber="247">Fig. 1AB</figureCitation>
); aedeagus with two spicules, right sclerite short Y-shaped, left sclerite strap-shaped, flattened, apically dentated (
<figureCitation id="134B2A17FFA2FFA0FE4D12F42F2F88FE" box="[469,592,475,501]" captionStart="FIGURE 2" captionStartId="4.[152,255,1893,1917]" captionTargetBox="[203,1394,219,1821]" captionTargetId="figure-17@4.[151,1436,179,1868]" captionTargetPageId="4" captionText="FIGURE 2. Genitalia of S. qinghaiensis sp. nov. (A) left paramere; (A') top view of left paramere; (B) right paramere; (B) top view of right paramere; (C) aedeagus in ventral views; (DE) endosoma with detached phallotheca in two different views." pageId="3" pageNumber="247">Fig. 2DE</figureCitation>
<figureCitation id="134B2A17FFA2FFA0FE4D12F42F2F88FE" box="[469,592,475,501]" captionStart="FIGURE 2" captionStartId="4.[152,255,1893,1917]" captionTargetBox="[203,1394,219,1821]" captionTargetId="figure-17@4.[151,1436,179,1868]" captionTargetPageId="4" captionText="FIGURE 2. Genitalia of S. qinghaiensis sp. nov. (A) left paramere; (A') top view of left paramere; (B) right paramere; (B) top view of right paramere; (C) aedeagus in ventral views; (DE) endosoma with detached phallotheca in two different views." figureDoi="http://doi.org/10.5281/zenodo.13618200" httpUri="https://zenodo.org/record/13618200/files/figure.png" pageId="3" pageNumber="247">Fig. 2DE</figureCitation>
).
</paragraph>
<paragraph id="8BCF3692FFA2FFA0FF5F12D02EE28B6A" blockId="3.[151,1437,331,1833]" pageId="3" pageNumber="247">
@ -123,25 +124,25 @@ is divided into multiple forks.
<paragraph id="8BCF3692FFA2FFA0FF5F11442F5C8C06" blockId="3.[151,1437,331,1833]" pageId="3" pageNumber="247">
<emphasis id="B904EA80FFA2FFA0FF5F11442CE48B8E" bold="true" box="[199,411,619,645]" pageId="3" pageNumber="247">Description. Male</emphasis>
: Brachypterous, relatively small, total length 2.342.47. COLORATION (
<figureCitation id="134B2A17FFA2FFA0FB57114428588B8E" box="[1231,1319,619,645]" captionStart="FIGURE 1" captionStartId="2.[152,255,1874,1898]" captionTargetBox="[226,1378,228,1811]" captionTargetId="figure-17@2.[151,1436,179,1848]" captionTargetPageId="2" captionText="FIGURE 1. Habitus views of S. qinghaiensis sp. nov. (A) female; (B) male and S. shanxiensis sp. nov. (C) female; (D) male." pageId="3" pageNumber="247">Fig. 1A</figureCitation>
<figureCitation id="134B2A17FFA2FFA0FB57114428588B8E" box="[1231,1319,619,645]" captionStart="FIGURE 1" captionStartId="2.[152,255,1874,1898]" captionTargetBox="[226,1378,228,1811]" captionTargetId="figure-17@2.[151,1436,179,1848]" captionTargetPageId="2" captionText="FIGURE 1. Habitus views of S. qinghaiensis sp. nov. (A) female; (B) male and S. shanxiensis sp. nov. (C) female; (D) male." figureDoi="http://doi.org/10.5281/zenodo.13618198" httpUri="https://zenodo.org/record/13618198/files/figure.png" pageId="3" pageNumber="247">Fig. 1A</figureCitation>
): Dorsum, thoracic pleura, venter, and appendages black; head black, with two smooth dark black spots on the inner margins of the eyes, and sometimes with yellow at the edge of connexivum. SURFACE AND VESTITURE: Dorsum, venter and head except vertex all smooth, inner side of eyes and clypeus highly polished. Head, thorax, hemelytra, genital capsule and abdomen with black long setae. STRUCTURE: Body 2.813.12 × as long as basal width of pronotum. Head: vertical, of nearly equal height and width; vertex flat; eyes relatively small, vertex between inner margins of eyes 1.571.62 × as wide as eye; almost round in lateral; antennal fossa located at distance from ventral margin of eye; antennal segments cylindrical, first segment 1.021.38 × as long as pronotum; second segment 1.241.38 × as long as basal width pronotum, 1.091.30 × as long as width head; third and fourth segments filiform; labium stout, always reaching and slightly surpassing hind coxa. Thorax: Pronotum trapeziform, 1.922.05 × as wide as length, 0.880.96 × as wide as head, with rounded anterior angles, anterior margin slightly convex while posterior margin concave; calli large, raised; mesonotum invisible; scutellum all black, width longer than length; claval commissural margin shorter than length of scutellum; hind femur enlarged. Abdomen: oval, gradually expanding at middle; connexivum upwarped. GENITALIA: Parameres of typical
<taxonomicName id="4C704D11FFA2FFA0FC281710296D8D52" authorityName="A.Costa" authorityYear="1853" box="[944,1042,1087,1113]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="247" phylum="Arthropoda" rank="tribe" tribe="Halticini">Halticini</taxonomicName>
shape (
<figureCitation id="134B2A17FFA2FFA0FBF11710299B8D52" box="[1129,1252,1087,1113]" captionStart="FIGURE 2" captionStartId="4.[152,255,1893,1917]" captionTargetBox="[203,1394,219,1821]" captionTargetId="figure-17@4.[151,1436,179,1868]" captionTargetPageId="4" captionText="FIGURE 2. Genitalia of S. qinghaiensis sp. nov. (A) left paramere; (A') top view of left paramere; (B) right paramere; (B) top view of right paramere; (C) aedeagus in ventral views; (DE) endosoma with detached phallotheca in two different views." pageId="3" pageNumber="247">Fig. 2AB</figureCitation>
<figureCitation id="134B2A17FFA2FFA0FBF11710299B8D52" box="[1129,1252,1087,1113]" captionStart="FIGURE 2" captionStartId="4.[152,255,1893,1917]" captionTargetBox="[203,1394,219,1821]" captionTargetId="figure-17@4.[151,1436,179,1868]" captionTargetPageId="4" captionText="FIGURE 2. Genitalia of S. qinghaiensis sp. nov. (A) left paramere; (A') top view of left paramere; (B) right paramere; (B) top view of right paramere; (C) aedeagus in ventral views; (DE) endosoma with detached phallotheca in two different views." figureDoi="http://doi.org/10.5281/zenodo.13618200" httpUri="https://zenodo.org/record/13618200/files/figure.png" pageId="3" pageNumber="247">Fig. 2AB</figureCitation>
), right paramere larger than left one, with apex flat and concave, basal elongated and narrow, bearing blunt tubercle(
<figureCitation id="134B2A17FFA2FFA0FA9E174C28388D76" box="[1286,1351,1123,1149]" captionStart="FIGURE 2" captionStartId="4.[152,255,1893,1917]" captionTargetBox="[203,1394,219,1821]" captionTargetId="figure-17@4.[151,1436,179,1868]" captionTargetPageId="4" captionText="FIGURE 2. Genitalia of S. qinghaiensis sp. nov. (A) left paramere; (A') top view of left paramere; (B) right paramere; (B) top view of right paramere; (C) aedeagus in ventral views; (DE) endosoma with detached phallotheca in two different views." pageId="3" pageNumber="247">Fig. 2</figureCitation>
<figureCitation id="134B2A17FFA2FFA0FA9E174C28388D76" box="[1286,1351,1123,1149]" captionStart="FIGURE 2" captionStartId="4.[152,255,1893,1917]" captionTargetBox="[203,1394,219,1821]" captionTargetId="figure-17@4.[151,1436,179,1868]" captionTargetPageId="4" captionText="FIGURE 2. Genitalia of S. qinghaiensis sp. nov. (A) left paramere; (A') top view of left paramere; (B) right paramere; (B) top view of right paramere; (C) aedeagus in ventral views; (DE) endosoma with detached phallotheca in two different views." figureDoi="http://doi.org/10.5281/zenodo.13618200" httpUri="https://zenodo.org/record/13618200/files/figure.png" pageId="3" pageNumber="247">Fig. 2</figureCitation>
B'); left paramere Γ- shaped, strong curving, with two small cones(
<figureCitation id="134B2A17FFA2FFA0FCB917A82E1F8DAA" box="[801,864,1159,1185]" captionStart="FIGURE 2" captionStartId="4.[152,255,1893,1917]" captionTargetBox="[203,1394,219,1821]" captionTargetId="figure-17@4.[151,1436,179,1868]" captionTargetPageId="4" captionText="FIGURE 2. Genitalia of S. qinghaiensis sp. nov. (A) left paramere; (A') top view of left paramere; (B) right paramere; (B) top view of right paramere; (C) aedeagus in ventral views; (DE) endosoma with detached phallotheca in two different views." pageId="3" pageNumber="247">Fig. 2</figureCitation>
<figureCitation id="134B2A17FFA2FFA0FCB917A82E1F8DAA" box="[801,864,1159,1185]" captionStart="FIGURE 2" captionStartId="4.[152,255,1893,1917]" captionTargetBox="[203,1394,219,1821]" captionTargetId="figure-17@4.[151,1436,179,1868]" captionTargetPageId="4" captionText="FIGURE 2. Genitalia of S. qinghaiensis sp. nov. (A) left paramere; (A') top view of left paramere; (B) right paramere; (B) top view of right paramere; (C) aedeagus in ventral views; (DE) endosoma with detached phallotheca in two different views." figureDoi="http://doi.org/10.5281/zenodo.13618200" httpUri="https://zenodo.org/record/13618200/files/figure.png" pageId="3" pageNumber="247">Fig. 2</figureCitation>
A'); phallotheca (
<figureCitation id="134B2A17FFA2FFA0FB8617A8290C8DAA" box="[1054,1139,1159,1185]" captionStart="FIGURE 2" captionStartId="4.[152,255,1893,1917]" captionTargetBox="[203,1394,219,1821]" captionTargetId="figure-17@4.[151,1436,179,1868]" captionTargetPageId="4" captionText="FIGURE 2. Genitalia of S. qinghaiensis sp. nov. (A) left paramere; (A') top view of left paramere; (B) right paramere; (B) top view of right paramere; (C) aedeagus in ventral views; (DE) endosoma with detached phallotheca in two different views." pageId="3" pageNumber="247">Fig. 2C</figureCitation>
<figureCitation id="134B2A17FFA2FFA0FB8617A8290C8DAA" box="[1054,1139,1159,1185]" captionStart="FIGURE 2" captionStartId="4.[152,255,1893,1917]" captionTargetBox="[203,1394,219,1821]" captionTargetId="figure-17@4.[151,1436,179,1868]" captionTargetPageId="4" captionText="FIGURE 2. Genitalia of S. qinghaiensis sp. nov. (A) left paramere; (A') top view of left paramere; (B) right paramere; (B) top view of right paramere; (C) aedeagus in ventral views; (DE) endosoma with detached phallotheca in two different views." figureDoi="http://doi.org/10.5281/zenodo.13618200" httpUri="https://zenodo.org/record/13618200/files/figure.png" pageId="3" pageNumber="247">Fig. 2C</figureCitation>
) with membranous ventral wall, apically dentated and equipped with pair of subapical, rounded, sclerotized outgrowths at sides; aedeagus (
<figureCitation id="134B2A17FFA2FFA0FAF717842DA58DE2" captionStart="FIGURE 2" captionStartId="4.[152,255,1893,1917]" captionTargetBox="[203,1394,219,1821]" captionTargetId="figure-17@4.[151,1436,179,1868]" captionTargetPageId="4" captionText="FIGURE 2. Genitalia of S. qinghaiensis sp. nov. (A) left paramere; (A') top view of left paramere; (B) right paramere; (B) top view of right paramere; (C) aedeagus in ventral views; (DE) endosoma with detached phallotheca in two different views." pageId="3" pageNumber="247">Fig. 2DE</figureCitation>
<figureCitation id="134B2A17FFA2FFA0FAF717842DA58DE2" captionStart="FIGURE 2" captionStartId="4.[152,255,1893,1917]" captionTargetBox="[203,1394,219,1821]" captionTargetId="figure-17@4.[151,1436,179,1868]" captionTargetPageId="4" captionText="FIGURE 2. Genitalia of S. qinghaiensis sp. nov. (A) left paramere; (A') top view of left paramere; (B) right paramere; (B) top view of right paramere; (C) aedeagus in ventral views; (DE) endosoma with detached phallotheca in two different views." figureDoi="http://doi.org/10.5281/zenodo.13618200" httpUri="https://zenodo.org/record/13618200/files/figure.png" pageId="3" pageNumber="247">Fig. 2DE</figureCitation>
) with two spicules, right sclerite short Y-shaped, split in apical one-third; left sclerite strap-shaped, flattened, twisted medially, apically dentated.
</paragraph>
<paragraph id="8BCF3692FFA2FFA0FF5F16382ECE8F26" blockId="3.[151,1437,331,1833]" pageId="3" pageNumber="247">
<emphasis id="B904EA80FFA2FFA0FF5F16382C578C3A" bold="true" box="[199,296,1303,1329]" pageId="3" pageNumber="247">Female:</emphasis>
COLORATION (
<figureCitation id="134B2A17FFA2FFA0FE7516382F3E8C3A" box="[493,577,1303,1329]" captionStart="FIGURE 1" captionStartId="2.[152,255,1874,1898]" captionTargetBox="[226,1378,228,1811]" captionTargetId="figure-17@2.[151,1436,179,1848]" captionTargetPageId="2" captionText="FIGURE 1. Habitus views of S. qinghaiensis sp. nov. (A) female; (B) male and S. shanxiensis sp. nov. (C) female; (D) male." pageId="3" pageNumber="247">Fig. 1B</figureCitation>
<figureCitation id="134B2A17FFA2FFA0FE7516382F3E8C3A" box="[493,577,1303,1329]" captionStart="FIGURE 1" captionStartId="2.[152,255,1874,1898]" captionTargetBox="[226,1378,228,1811]" captionTargetId="figure-17@2.[151,1436,179,1848]" captionTargetPageId="2" captionText="FIGURE 1. Habitus views of S. qinghaiensis sp. nov. (A) female; (B) male and S. shanxiensis sp. nov. (C) female; (D) male." figureDoi="http://doi.org/10.5281/zenodo.13618198" httpUri="https://zenodo.org/record/13618198/files/figure.png" pageId="3" pageNumber="247">Fig. 1B</figureCitation>
): Similar to male, but with yellow at the edge of hemelytron. Dorsum, thoracic pleura, venter, and appendages black; head black, with two smooth dark black spots on the inner margins of the eyes. SURFACE AND VESTITURE: Like male. STRUCTURE: Body larger than male, total length 2.67, 3.10 × as long as basal width of pronotum. Head: Vertical, vertex between inner margins of eyes 1.62 × as wide as eye; antennal segments cylindrical, first segment 1.07 × as long as pronotum; second segment 1.05 × as long as basal width of pronotum, 0.98 × as long as width of head; third and fourth segments filiform; labium stout, always reaching and slightly surpassing hind coxa. Thorax: Pronotum 2.09 × as wide as length, 0.93 × as wide as head. Abdomen: much wider width medially than in male, almost equal in length and width.
</paragraph>
</subSubSection>

View file

@ -1,66 +1,67 @@
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<mods:title id="0C2F6B4E5055CA0ADD9BC9A870D8EC53">Two new species of the brachypterous Scirtetellus (Hemiptera: Heteroptera: Miridae) from China</mods:title>
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@ -80,9 +81,9 @@ Liu
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<paragraph id="8BCF3692FFA4FFA6FF0F13942C3F89DE" blockId="5.[151,734,151,213]" box="[151,320,187,213]" pageId="5" pageNumber="249">
(
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<figureCitation id="134B2A17FFA4FFA6FF0713942C6289DE" box="[159,285,187,213]" captionStart="FIGURE 1" captionStartId="2.[152,255,1874,1898]" captionTargetBox="[226,1378,228,1811]" captionTargetId="figure-17@2.[151,1436,179,1848]" captionTargetPageId="2" captionText="FIGURE 1. Habitus views of S. qinghaiensis sp. nov. (A) female; (B) male and S. shanxiensis sp. nov. (C) female; (D) male." figureDoi="http://doi.org/10.5281/zenodo.13618198" httpUri="https://zenodo.org/record/13618198/files/figure.png" pageId="5" pageNumber="249">Figs 1CD</figureCitation>
,
<figureCitation id="134B2A17FFA4FFA6FEB113942C4889DE" box="[297,311,187,213]" captionStart="FIGURE 3" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[243,1350,275,1889]" captionTargetId="figure-17@6.[214,1373,179,1917]" captionTargetPageId="6" captionText="FIGURE 3. Genitalia of S. shanxiensis sp. nov. (A) left paramere; (A) top view of left paramere; (B) right paramere; (B) top view of right paramere; (C) aedeagus in ventral views; (DE) endosoma with detached phallotheca in two different views; (ad) parts of left sclerite." pageId="5" pageNumber="249">3</figureCitation>
<figureCitation id="134B2A17FFA4FFA6FEB113942C4889DE" box="[297,311,187,213]" captionStart="FIGURE 3" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[243,1350,275,1889]" captionTargetId="figure-17@6.[214,1373,179,1917]" captionTargetPageId="6" captionText="FIGURE 3. Genitalia of S. shanxiensis sp. nov. (A) left paramere; (A) top view of left paramere; (B) right paramere; (B) top view of right paramere; (C) aedeagus in ventral views; (DE) endosoma with detached phallotheca in two different views; (ad) parts of left sclerite." figureDoi="http://doi.org/10.5281/zenodo.13618204" httpUri="https://zenodo.org/record/13618204/files/figure.png" pageId="5" pageNumber="249">3</figureCitation>
)
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@ -90,7 +91,7 @@ Liu
<paragraph id="8BCF3692FFA4FFA6FF0F122C299D886D" blockId="5.[151,1437,259,1725]" pageId="5" pageNumber="249">
<emphasis id="B904EA80FFA4FFA6FF0F122C2C6D8816" bold="true" box="[151,274,259,285]" pageId="5" pageNumber="249">Diagnosis.</emphasis>
Recognized by the following combination of characters: relatively medium size (male 2.752.91, female 3.423.57); head black, with district yellow pattern; pronotum and hemelytron uniformly dark (
<figureCitation id="134B2A17FFA4FFA6FB281208285C884A" box="[1200,1315,295,321]" captionStart="FIGURE 1" captionStartId="2.[152,255,1874,1898]" captionTargetBox="[226,1378,228,1811]" captionTargetId="figure-17@2.[151,1436,179,1848]" captionTargetPageId="2" captionText="FIGURE 1. Habitus views of S. qinghaiensis sp. nov. (A) female; (B) male and S. shanxiensis sp. nov. (C) female; (D) male." pageId="5" pageNumber="249">Fig.1CD</figureCitation>
<figureCitation id="134B2A17FFA4FFA6FB281208285C884A" box="[1200,1315,295,321]" captionStart="FIGURE 1" captionStartId="2.[152,255,1874,1898]" captionTargetBox="[226,1378,228,1811]" captionTargetId="figure-17@2.[151,1436,179,1848]" captionTargetPageId="2" captionText="FIGURE 1. Habitus views of S. qinghaiensis sp. nov. (A) female; (B) male and S. shanxiensis sp. nov. (C) female; (D) male." figureDoi="http://doi.org/10.5281/zenodo.13618198" httpUri="https://zenodo.org/record/13618198/files/figure.png" pageId="5" pageNumber="249">Fig.1CD</figureCitation>
); aedeagus with two spicules, right sclerite long and curved, left sclerite complex, shown in the figure 3 DE.
</paragraph>
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@ -121,23 +122,23 @@ are unforked.
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<emphasis id="B904EA80FFA4FFA6FF5F12F42CE288FE" bold="true" box="[199,413,475,501]" pageId="5" pageNumber="249">Description. Male</emphasis>
: Brachypterous, relatively medium size, total length 2.752.91. COLORATION (
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<figureCitation id="134B2A17FFA4FFA6FAAD12F428F188FE" box="[1333,1422,475,501]" captionStart="FIGURE 1" captionStartId="2.[152,255,1874,1898]" captionTargetBox="[226,1378,228,1811]" captionTargetId="figure-17@2.[151,1436,179,1848]" captionTargetPageId="2" captionText="FIGURE 1. Habitus views of S. qinghaiensis sp. nov. (A) female; (B) male and S. shanxiensis sp. nov. (C) female; (D) male." figureDoi="http://doi.org/10.5281/zenodo.13618198" httpUri="https://zenodo.org/record/13618198/files/figure.png" pageId="5" pageNumber="249">Fig. 1C</figureCitation>
): Dorsum, thoracic pleura, venter, and appendages black; head black with yellow patten, basal two-thirds of mandibular and maxillary plates, and uneven stripes along eyes on vertex, frons and genae, sometimes with yellow at the edge of connexivum. SURFACE AND VESTITURE: Dorsum and venter smooth, head except vertex distinctly polished, shining, abdomen weakly shining. Entire dorsum with dense, adpressed, silvery, moderately flattened scalelike setae; genital capsule and all appendages with dense, erect, distinctly long, dark simple setae. STRUCTURE: Body 3.063.46 × as long as basal width of pronotum. Head: vertical, of nearly equal height and width; vertex flat; eyes relatively small, vertex between inner margins of eyes 1.571.62 × as wide as eye; almost round in lateral; antennal fossa located at a distance from ventral margin of eye; antennal segments cylindrical, first segment 1.211.32 × as long as pronotum; labium stout, always reaching and slightly surpassing hind coxa. Thorax: pronotum trapeziform, 1.792.09 × as wide as length, 0.881.00 × as wide as head, with rounded anterior angles, anterior margin slightly convex while posterior margin concave; calli large, raised; mesonotum invisible; scutellum all black, width is longer than length; claval commissural margin is shorter than length of scutellum. Abdomen: Oval, gradually expanding at middle; connexivum upturned. GENITALIA: parameres of typical
<taxonomicName id="4C704D11FFA4FFA6FC0410802E818AC2" authorityName="A.Costa" authorityYear="1853" box="[924,1022,943,969]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="249" phylum="Arthropoda" rank="tribe" tribe="Halticini">Halticini</taxonomicName>
shape (
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<figureCitation id="134B2A17FFA4FFA6FBCB108029B28AC2" box="[1107,1229,943,969]" captionStart="FIGURE 3" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[243,1350,275,1889]" captionTargetId="figure-17@6.[214,1373,179,1917]" captionTargetPageId="6" captionText="FIGURE 3. Genitalia of S. shanxiensis sp. nov. (A) left paramere; (A) top view of left paramere; (B) right paramere; (B) top view of right paramere; (C) aedeagus in ventral views; (DE) endosoma with detached phallotheca in two different views; (ad) parts of left sclerite." figureDoi="http://doi.org/10.5281/zenodo.13618204" httpUri="https://zenodo.org/record/13618204/files/figure.png" pageId="5" pageNumber="249">Fig. 3AB</figureCitation>
), right paramere is larger than left one, with apex flat and concave, basal elongated and narrow, bearing blunt tubercle (
<figureCitation id="134B2A17FFA4FFA6FA9C10FC28398AE6" box="[1284,1350,979,1005]" captionStart="FIGURE 3" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[243,1350,275,1889]" captionTargetId="figure-17@6.[214,1373,179,1917]" captionTargetPageId="6" captionText="FIGURE 3. Genitalia of S. shanxiensis sp. nov. (A) left paramere; (A) top view of left paramere; (B) right paramere; (B) top view of right paramere; (C) aedeagus in ventral views; (DE) endosoma with detached phallotheca in two different views; (ad) parts of left sclerite." pageId="5" pageNumber="249">Fig. 3</figureCitation>
<figureCitation id="134B2A17FFA4FFA6FA9C10FC28398AE6" box="[1284,1350,979,1005]" captionStart="FIGURE 3" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[243,1350,275,1889]" captionTargetId="figure-17@6.[214,1373,179,1917]" captionTargetPageId="6" captionText="FIGURE 3. Genitalia of S. shanxiensis sp. nov. (A) left paramere; (A) top view of left paramere; (B) right paramere; (B) top view of right paramere; (C) aedeagus in ventral views; (DE) endosoma with detached phallotheca in two different views; (ad) parts of left sclerite." figureDoi="http://doi.org/10.5281/zenodo.13618204" httpUri="https://zenodo.org/record/13618204/files/figure.png" pageId="5" pageNumber="249">Fig. 3</figureCitation>
B'); left paramere sickle-shaped, curving, with two small cones (
<figureCitation id="134B2A17FFA4FFA6FC8910D82E2D8D1A" box="[785,850,1015,1041]" captionStart="FIGURE 3" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[243,1350,275,1889]" captionTargetId="figure-17@6.[214,1373,179,1917]" captionTargetPageId="6" captionText="FIGURE 3. Genitalia of S. shanxiensis sp. nov. (A) left paramere; (A) top view of left paramere; (B) right paramere; (B) top view of right paramere; (C) aedeagus in ventral views; (DE) endosoma with detached phallotheca in two different views; (ad) parts of left sclerite." pageId="5" pageNumber="249">Fig. 3</figureCitation>
<figureCitation id="134B2A17FFA4FFA6FC8910D82E2D8D1A" box="[785,850,1015,1041]" captionStart="FIGURE 3" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[243,1350,275,1889]" captionTargetId="figure-17@6.[214,1373,179,1917]" captionTargetPageId="6" captionText="FIGURE 3. Genitalia of S. shanxiensis sp. nov. (A) left paramere; (A) top view of left paramere; (B) right paramere; (B) top view of right paramere; (C) aedeagus in ventral views; (DE) endosoma with detached phallotheca in two different views; (ad) parts of left sclerite." figureDoi="http://doi.org/10.5281/zenodo.13618204" httpUri="https://zenodo.org/record/13618204/files/figure.png" pageId="5" pageNumber="249">Fig. 3</figureCitation>
A'); phallotheca (
<figureCitation id="134B2A17FFA4FFA6FB8E10D829138D1A" box="[1046,1132,1015,1041]" captionStart="FIGURE 3" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[243,1350,275,1889]" captionTargetId="figure-17@6.[214,1373,179,1917]" captionTargetPageId="6" captionText="FIGURE 3. Genitalia of S. shanxiensis sp. nov. (A) left paramere; (A) top view of left paramere; (B) right paramere; (B) top view of right paramere; (C) aedeagus in ventral views; (DE) endosoma with detached phallotheca in two different views; (ad) parts of left sclerite." pageId="5" pageNumber="249">Fig. 3C</figureCitation>
<figureCitation id="134B2A17FFA4FFA6FB8E10D829138D1A" box="[1046,1132,1015,1041]" captionStart="FIGURE 3" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[243,1350,275,1889]" captionTargetId="figure-17@6.[214,1373,179,1917]" captionTargetPageId="6" captionText="FIGURE 3. Genitalia of S. shanxiensis sp. nov. (A) left paramere; (A) top view of left paramere; (B) right paramere; (B) top view of right paramere; (C) aedeagus in ventral views; (DE) endosoma with detached phallotheca in two different views; (ad) parts of left sclerite." figureDoi="http://doi.org/10.5281/zenodo.13618204" httpUri="https://zenodo.org/record/13618204/files/figure.png" pageId="5" pageNumber="249">Fig. 3C</figureCitation>
) with membranous ventral wall, apically dentated and equipped with pair of subapical, rounded, sclerotized outgrowths at sides; aedeagus with two spicules, right sclerite long and curved, left sclerite complex, divided into multiple forks, we will name them Parts ad; Part a elongated, wider at the apex than at the base, Part b curved in a C-shape with teeth on the inner side, Part c and Part d extended at right angles, shown in the figure 3DE.
</paragraph>
<paragraph id="8BCF3692FFA4FFA6FF5F17842C718C96" blockId="5.[151,1437,259,1725]" pageId="5" pageNumber="249">
<emphasis id="B904EA80FFA4FFA6FF5F17842C578DCE" bold="true" box="[199,296,1195,1221]" pageId="5" pageNumber="249">Female:</emphasis>
COLORATION (
<figureCitation id="134B2A17FFA4FFA6FE6D17842F2F8DCE" box="[501,592,1195,1221]" captionStart="FIGURE 1" captionStartId="2.[152,255,1874,1898]" captionTargetBox="[226,1378,228,1811]" captionTargetId="figure-17@2.[151,1436,179,1848]" captionTargetPageId="2" captionText="FIGURE 1. Habitus views of S. qinghaiensis sp. nov. (A) female; (B) male and S. shanxiensis sp. nov. (C) female; (D) male." pageId="5" pageNumber="249">Fig. 1D</figureCitation>
<figureCitation id="134B2A17FFA4FFA6FE6D17842F2F8DCE" box="[501,592,1195,1221]" captionStart="FIGURE 1" captionStartId="2.[152,255,1874,1898]" captionTargetBox="[226,1378,228,1811]" captionTargetId="figure-17@2.[151,1436,179,1848]" captionTargetPageId="2" captionText="FIGURE 1. Habitus views of S. qinghaiensis sp. nov. (A) female; (B) male and S. shanxiensis sp. nov. (C) female; (D) male." figureDoi="http://doi.org/10.5281/zenodo.13618198" httpUri="https://zenodo.org/record/13618198/files/figure.png" pageId="5" pageNumber="249">Fig. 1D</figureCitation>
): Like male, dorsum, thoracic pleura, venter, and appendages black; head black, with district yellow pattern; SURFACE AND VESTITURE: Similar to male. STRUCTURE: Body larger than male, total length 3.423.57, 3.563.94 ×as long as basal width of pronotum. Head: vertical, vertex between inner margins of eyes 1.721.89 × as wide as eye; antennal segments cylindrical, first segment 1.181.27 × as long as pronotum; labium stout, always reaching and slightly surpassing hind coxa. Thorax: Pronotum 1.982.13 × as wide as length, 0.860.91 × as wide as head. Abdomen: much wider medially than male, almost equal in length and width.
</paragraph>
</subSubSection>

View file

@ -0,0 +1,616 @@
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<mods:title id="80CDBBC845F9278293A293BC44CFC4ED">A second species in the polychaete-associated crab genus Gustavus Ahyong &amp; Ng, 2009 (Decapoda: Brachyura: Aphanodactylidae)</mods:title>
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<mods:namePart id="E9D16243F4570822D84C058847577CBD">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="75829C92F9EC3DC92245A938AAB47864">Anker, Arthur</mods:namePart>
<mods:affiliation id="1B8780ACAF902317626EABCEF9D47DCD">Universidade Federal de Pelotas (UFPEL), Departamento de Ecologia, Zoologia e Genética, Instituto de Biologia, Campus Universitário Capão do Leão, RS, 96010 - 610, Brazil. E-mail &amp; Biological and Environmental Science &amp; Engineering Division, King Abdullah University of Science and Technology (KAUST), Thuwal, Saudi Arabia.</mods:affiliation>
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<paragraph id="8BCD36927A02FFCA43A0FA1840DEF9B5" blockId="1.[151,499,1534,1597]" box="[151,499,1534,1561]" pageId="1" pageNumber="370">
<heading id="D08581FE7A02FFCA43A0FA1840DEF9B5" bold="true" box="[151,499,1534,1561]" fontSize="11" level="1" pageId="1" pageNumber="370" reason="1">
<emphasis id="B906EA807A02FFCA43A0FA1840DEF9B5" bold="true" box="[151,499,1534,1561]" pageId="1" pageNumber="370">
<taxonomicName id="4C724D117A02FFCA43A0FA1840A3F9B5" authorityName="Ng &amp; Anker" authorityYear="2024" box="[151,398,1534,1561]" class="Malacostraca" family="Aphanodactylidae" genus="Gustavus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="species" species="holthuisae" status="sp. nov.">
<emphasis id="B906EA807A02FFCA43A0FA1840A3F9B5" bold="true" box="[151,398,1534,1561]" italics="true" pageId="1" pageNumber="370">Gustavus holthuisae</emphasis>
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<taxonomicNameLabel id="A23557FB7A02FFCA42A2FA1840DEF9B5" box="[405,499,1535,1561]" pageId="1" pageNumber="370" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C36865197A02FFCA43A0F9C4403FF991" box="[151,274,1571,1597]" pageId="1" pageNumber="370" type="description">
<paragraph id="8BCD36927A02FFCA43A0F9C4403FF991" blockId="1.[151,499,1534,1597]" box="[151,274,1571,1597]" pageId="1" pageNumber="370">
(
<figureCitation id="13492A177A02FFCA43A8F9C44024F991" box="[159,265,1571,1597]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="2.[151,250,1918,1942]" captionStartId-1="4.[151,250,1943,1967]" captionStartId-2="5.[151,250,1882,1906]" captionStartId-3="6.[151,250,1851,1875]" captionStartId-4="7.[151,250,1552,1576]" captionStartId-5="8.[151,250,1821,1845]" captionTargetBox-0="[151,1435,181,1894]" captionTargetBox-1="[193,1394,181,1919]" captionTargetBox-2="[151,1436,181,1858]" captionTargetBox-3="[151,1435,181,1828]" captionTargetBox-4="[151,1435,181,1528]" captionTargetBox-5="[158,1429,460,1791]" captionTargetId-0="figure-17@2.[151,1435,181,1894]" captionTargetId-1="figure-17@4.[193,1394,181,1919]" captionTargetId-2="figure-17@5.[151,1436,181,1858]" captionTargetId-3="figure-17@6.[151,1435,181,1828]" captionTargetId-4="figure-17@7.[151,1435,181,1528]" captionTargetId-5="figure-130@8.[151,1435,453,1798]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionText-0="FIGURE 1. Gustavus holthuisae sp. nov., ovigerous female paratype (6.3 × 3.7 mm) from Thuwal, Saudi Arabia, FLMNH UF 72877, crab with terebellid polychaete host shortly after collection; A, general view of worm with crab; B, close-up of crab clinging to worm host. Photographs by A. Anker." captionText-1="FIGURE 2. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876, habitus and colour in life; A, dorsal view; B, ventral view; C, frontal view. Photographs by A. Anker." captionText-2="FIGURE 3. Gustavus holthuisaesp. nov., ovigerous female paratype (7.3 × 4.0 mm) from Thuwal SaudiArabia, ZRC 2023.0580, habitus and colour in life; A, dorsal view; B, ventral view; C, frontal view. Photographs by A. Anker." captionText-3="FIGURE 4. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, C, EG]; ovigerous female paratype (6.3 × 3.7 mm), FLMNH UF 72877, from type locality [B, D]; A, B, habitus in dorsal view; C, D, frontal margin and orbits, dorsal view; E, left G1, ventral view; F, left G1, dorsal view; G, left G2, ventral view." captionText-4="FIGURE 5. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [AD]; ovigerous female paratype (6.3 × 3.7 mm) from type locality, FLMNH UF 72877 [E, F, H]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [G]; A, E, cephalothorax showing orbits, epistome, antennules and antennae, frontal view; B, right chela (partially denuded), outer view; C, male buccal cavity and anterior thoracic sternum, ventral view; D, male anterior thoracic sternum and sternopleonal cavity, showing intact right G1, ventral view; F, right chela (denuded), outer view; G, female pleon; H, female sternopleonal cavity and vulvae, ventral view." captionText-5="FIGURE 6. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, CJ]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [B, KN]; A, B, right Mxp3 (setae denuded), outer view; CF, male P2P5, respectively, dorsal view; G, male pleon; H, left G1, ventral view (setae omitted); I, left G1, dorsal view (setae omitted); J, left G2, ventral view (setae omitted); KN, female P2P5, respectively, dorsal view. Scales: A, B, HJ = 0.2 mm; CG, KN = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13618521" figureDoi-1="http://doi.org/10.5281/zenodo.13618523" figureDoi-2="http://doi.org/10.5281/zenodo.13618525" figureDoi-3="http://doi.org/10.5281/zenodo.13618527" figureDoi-4="http://doi.org/10.5281/zenodo.13618529" figureDoi-5="http://doi.org/10.5281/zenodo.13618533" httpUri-0="https://zenodo.org/record/13618521/files/figure.png" httpUri-1="https://zenodo.org/record/13618523/files/figure.png" httpUri-2="https://zenodo.org/record/13618525/files/figure.png" httpUri-3="https://zenodo.org/record/13618527/files/figure.png" httpUri-4="https://zenodo.org/record/13618529/files/figure.png" httpUri-5="https://zenodo.org/record/13618533/files/figure.png" pageId="1" pageNumber="370">Figs. 16</figureCitation>
)
</paragraph>
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<paragraph id="8BCD36927A02FFCA43A0F98C40E8F82D" blockId="1.[151,1437,1642,1921]" pageId="1" pageNumber="370">
<emphasis id="B906EA807A02FFCA43A0F98C406FF928" bold="true" box="[151,322,1642,1669]" pageId="1" pageNumber="370">Type material</emphasis>
.
<materialsCitation id="3B1A3CCF7A02FFCA4262F98C4511F95D" collectingDate="2023-01-21" collectionCode="FLMNH, KAUST" collectorName="Technology &amp; A. Anker" country="Saudi Arabia" county="King Abdullah Monument" elevation="5" latitude="22.340609" location="Red Sea" longLatPrecision="1" longitude="39.08752" municipality="King Abdullah University of Science" pageId="1" pageNumber="370" specimenCode="UF 72876" specimenCount="1" typeStatus="holotype">
<typeStatus id="54C988307A02FFCA4262F98C40EFF929" box="[341,450,1643,1669]" pageId="1" pageNumber="370" type="holotype">Holotype</typeStatus>
: male (5.3 ×
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) (
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<collectingCountry id="F36576027A02FFCA40EDF98C455EF929" box="[986,1139,1643,1669]" name="Saudi Arabia" pageId="1" pageNumber="370">Saudi Arabia</collectingCountry>
,
<location id="8EAD60497A02FFCA47B5F98C45CEF929" LSID="urn:lsid:plazi:treatment:03DB87847A02FFC143A0FA184013FEED:8EAD60497A02FFCA47B5F98C45CEF929" box="[1154,1251,1643,1669]" country="Saudi Arabia" county="King Abdullah Monument" latitude="22.340609" longLatPrecision="1" longitude="39.08752" municipality="King Abdullah University of Science" name="Red Sea" pageId="1" pageNumber="370">Red Sea</location>
coast, Sta. RS-41,
<location id="8EAD60497A02FFCA43F1F9684033F905" LSID="urn:lsid:plazi:treatment:03DB87847A02FFC143A0FA184013FEED:8EAD60497A02FFCA43F1F9684033F905" box="[198,286,1679,1705]" country="Saudi Arabia" county="King Abdullah Monument" latitude="22.340609" longLatPrecision="1" longitude="39.08752" municipality="King Abdullah University of Science" name="Thuwal" pageId="1" pageNumber="370">Thuwal</location>
,
<collectingMunicipality id="6BA9ACE87A02FFCA4218F96843C9F905" box="[303,740,1679,1705]" pageId="1" pageNumber="370">King Abdullah University of Science</collectingMunicipality>
&amp;
<collectorName id="268753447A02FFCA4025F96842BBF905" box="[786,918,1679,1705]" pageId="1" pageNumber="370">Technology</collectorName>
(
<collectionCode id="ED63AE577A02FFCA409DF9684520F905" box="[938,1037,1679,1705]" pageId="1" pageNumber="370">KAUST</collectionCode>
), near
<collectingCounty id="62AC4E1E7A02FFCA4754F96844B5F905" box="[1123,1432,1679,1705]" pageId="1" pageNumber="370">King Abdullah Monument</collectingCounty>
,
<geoCoordinate id="EE4650557A02FFCA43A0F954401CF961" box="[151,305,1715,1741]" degrees="22.340608" direction="north" orientation="latitude" pageId="1" pageNumber="370" precision="1" value="22.340609">22.340608°N</geoCoordinate>
,
<geoCoordinate id="EE4650557A02FFCA4208F95540F8F961" box="[319,469,1714,1741]" degrees="39.087519" direction="east" orientation="longitude" pageId="1" pageNumber="370" precision="1" value="39.08752">39.087519°E</geoCoordinate>
, sand flat with fine sand near mangrove roots, less than
<quantity id="4C8A9B777A02FFCA4758F955459EF961" box="[1135,1203,1714,1741]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="370" unit="m" value="0.5">
<elevation id="005FD1A17A02FFCA4758F955459EF961" box="[1135,1203,1714,1741]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="370" unit="m" value="0.5">0.5 m</elevation>
</quantity>
at low tide, suction (yabby) pump, in terebellid tube, coll.
<collectorName id="268753447A02FFCA417CF9304398F95D" box="[587,693,1751,1777]" pageId="1" pageNumber="370">A. Anker</collectorName>
,
<date id="FFCC10527A02FFCA41F6F9304251F95D" box="[705,892,1751,1777]" pageId="1" pageNumber="370" value="2023-01-21">
<collectingDate id="EF88E9BA7A02FFCA41F6F9304251F95D" box="[705,892,1751,1777]" pageId="1" pageNumber="370" value="2023-01-21">21 January 2023</collectingDate>
</date>
[AA-22-520/M]
</materialsCitation>
.
<materialsCitation id="3B1A3CCF7A02FFCA4770F93040ECF82D" collectingDate="2023-01-21" collectionCode="FLMNH, RSRC, KAUST" collectorName="Technology &amp; A. Anker" country="Saudi Arabia" county="King Abdullah Monument" elevation="5" latitude="22.340609" location="Red Sea" longLatPrecision="1" longitude="39.08752" municipality="King Abdullah University of Science" pageId="1" pageNumber="370" specimenCode="ZRC 2023.0580, UF 72877, UF 72878" specimenCount="3" typeStatus="paratype">
<typeStatus id="54C988307A02FFCA4770F9304595F95D" box="[1095,1208,1751,1777]" pageId="1" pageNumber="370" type="paratype">Paratypes</typeStatus>
: 1 ovigerous female (7.3 × 4.0 mm) (
<specimenCode id="DBD49EE97A02FFCA427AF91C4325F8B9" box="[333,520,1786,1813]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="370">ZRC 2023.0580</specimenCode>
), same collection data as for the holotype [AA-22-520/F]; 1 ovigerous female (6.3 ×
<quantity id="4C8A9B777A02FFCA4399F8F84028F895" box="[174,261,1823,1849]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="1" pageNumber="370" unit="mm" value="3.6">3.6 mm</quantity>
) (
<collectionCode id="ED63AE577A02FFCA422CF8F84052F895" box="[283,383,1823,1849]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="1" pageNumber="370" type="Museum">FLMNH</collectionCode>
<specimenCode id="DBD49EE97A02FFCA42B2F8F840D7F895" box="[389,506,1823,1849]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="1" pageNumber="370" type="Museum">UF 72877</specimenCode>
), same collection data as for the
<typeStatus id="54C988307A02FFCA4050F8F842E4F895" box="[871,969,1823,1849]" pageId="1" pageNumber="370" type="holotype">holotype</typeStatus>
[AA-22-509; terebellid host, AA-22-508, deposited in
<collectionCode id="ED63AE577A02FFCA4211F8A4405DF8F1" box="[294,368,1859,1885]" pageId="1" pageNumber="370">RSRC</collectionCode>
/
<collectionCode id="ED63AE577A02FFCA424EF8A440F7F8F1" box="[377,474,1859,1885]" pageId="1" pageNumber="370">KAUST</collectionCode>
]; 1 ovigerous female (5.9 ×
<quantity id="4C8A9B777A02FFCA4025F8A44244F8F1" box="[786,873,1859,1885]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="1" pageNumber="370" unit="mm" value="3.4">3.4 mm</quantity>
) (
<collectionCode id="ED63AE577A02FFCA4048F8A442CEF8F1" box="[895,995,1859,1885]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="1" pageNumber="370" type="Museum">FLMNH</collectionCode>
<specimenCode id="DBD49EE97A02FFCA40DDF8A44572F8F1" box="[1002,1119,1859,1885]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="1" pageNumber="370" type="Museum">UF 72878</specimenCode>
), same collection data as for the
<typeStatus id="54C988307A02FFCA43F7F880400FF82D" box="[192,290,1895,1921]" pageId="1" pageNumber="370" type="holotype">holotype</typeStatus>
[AA-22-510]
</materialsCitation>
.
</paragraph>
<caption id="DF0D661A7A01FFC943A0F89943A4F872" ID-DOI="http://doi.org/10.5281/zenodo.13618521" ID-Zenodo-Dep="13618521" httpUri="https://zenodo.org/record/13618521/files/figure.png" pageId="2" pageNumber="371" startId="2.[151,250,1918,1942]" targetBox="[151,1435,181,1894]" targetPageId="2" targetType="figure">
<paragraph id="8BCD36927A01FFC943A0F89943A4F872" blockId="2.[151,1437,1918,2015]" pageId="2" pageNumber="371">
<emphasis id="B906EA807A01FFC943A0F8994022F83A" bold="true" box="[151,271,1918,1942]" pageId="2" pageNumber="371">FIGURE 1</emphasis>
.
<taxonomicName id="4C724D117A01FFC94229F89840C3F83A" authorityName="Ng &amp; Anker" authorityYear="2024" box="[286,494,1918,1942]" class="Malacostraca" family="Aphanodactylidae" genus="Gustavus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="holthuisae" status="sp. nov.">
<emphasis id="B906EA807A01FFC94229F89840C3F83A" box="[286,494,1918,1942]" italics="true" pageId="2" pageNumber="371">Gustavus holthuisae</emphasis>
</taxonomicName>
<emphasis id="B906EA807A01FFC942C1F8994364F83A" bold="true" box="[502,585,1918,1942]" pageId="2" pageNumber="371">
<taxonomicNameLabel id="A23557FB7A01FFC942C1F8994364F83A" box="[502,585,1918,1942]" pageId="2" pageNumber="371" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ovigerous female paratype (6.3 × 3.7 mm) from Thuwal, Saudi Arabia, FLMNH UF 72877, crab with terebellid polychaete host shortly after collection; A, general view of worm with crab; B, close-up of crab clinging to worm host. Photographs by A. Anker.
</paragraph>
</caption>
<paragraph id="8BCD36927A00FFC843F0FF71443BFF54" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="372">
<emphasis id="B906EA807A00FFC843F0FF7140FFFF1C" bold="true" box="[199,466,150,176]" pageId="3" pageNumber="372">Comparative material</emphasis>
.
<taxonomicName id="4C724D117A00FFC842D5FF7042F2FF1C" authority="Ahyong &amp; Ng, 2009" authorityName="Ahyong &amp; Ng" authorityYear="2009" box="[482,991,150,177]" class="Malacostraca" family="Aphanodactylidae" genus="Gustavus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="372" phylum="Arthropoda" rank="species" species="mecognathus">
<emphasis id="B906EA807A00FFC842D5FF7043C5FF1C" box="[482,744,150,177]" italics="true" pageId="3" pageNumber="372">Gustavus mecognathus</emphasis>
<materialsCitation id="3B1A3CCF7A00FFC841D8FF7142F2FF1C" box="[751,991,150,177]" collectingDate="2000-03-20" collectionCode="ZRC" collectorName="G. Paulay" country="Northern Mariana Islands" location="Mariana Islands" pageId="3" pageNumber="372" specimenCode="ZRC 2010.0251, ZRC 2010.0252" specimenCount="2" typeStatus="holotype">
<bibRefCitation id="EFE34B637A00FFC841D8FF7142F2FF1C" author="Ahyong, S. T. &amp; Ng, P. K. L." box="[751,991,150,177]" pageId="3" pageNumber="372" pagination="33 - 47" refId="ref5317" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2009) Aphanodactylidae, a new family of thoracotreme crabs (Crustacea: Brachyura) symbiotic with polychaete worms. Zootaxa, 2289 (1), 33 - 47. https: // doi. org / 10.11646 / zootaxa. 2289.1.3" type="journal article" year="2009">Ahyong &amp; Ng, 2009</bibRefCitation>
</materialsCitation>
</taxonomicName>
.
<typeStatus id="54C988307A00FFC840DBFF714574FF1C" box="[1004,1113,150,176]" pageId="3" pageNumber="372" type="holotype">Holotype</typeStatus>
: male (8.8 ×
<quantity id="4C8A9B777A00FFC847C5FF714466FF1C" box="[1266,1355,150,176]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="3" pageNumber="372" unit="mm" value="6.1">6.1 mm</quantity>
) (
<specimenCode id="DBD49EE97A00FFC84653FF71403EFF79" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="372">ZRC 2010.0251</specimenCode>
);
<typeStatus id="54C988307A00FFC8421EFF5C40A3FF79" box="[297,398,187,213]" pageId="3" pageNumber="372" type="paratype">paratype</typeStatus>
: ovigerous female (13.9 × 8.0 mm) (
<specimenCode id="DBD49EE97A00FFC8400EFF5D42DAFF79" box="[825,1015,186,213]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="372">ZRC 2010.0252</specimenCode>
),
<collectingCountry id="F36576027A00FFC8473BFF5D45E9FF78" box="[1036,1220,186,212]" name="Northern Mariana Islands" pageId="3" pageNumber="372">Mariana Islands</collectingCountry>
,
<collectingCountry id="F36576027A00FFC847E5FF5D4434FF78" box="[1234,1305,186,212]" name="Guam" pageId="3" pageNumber="372">Guam</collectingCountry>
, Southwest Cocos Barrier, near small pass, with large terebellid polychaete worm, coll.
<collectorName id="268753447A00FFC840DEFF394575FF54" box="[1001,1112,222,248]" pageId="3" pageNumber="372">G. Paulay</collectorName>
,
<date id="FFCC10527A00FFC84753FF39443EFF54" box="[1124,1299,222,248]" pageId="3" pageNumber="372" value="2000-03-20">
<collectingDate id="EF88E9BA7A00FFC84753FF39443EFF54" box="[1124,1299,222,248]" pageId="3" pageNumber="372" value="2000-03-20">20 March 2000</collectingDate>
</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C36865197A00FFC843F0FEE54474FE58" pageId="3" pageNumber="372" type="diagnosis">
<paragraph id="8BCD36927A00FFC843F0FEE54474FE58" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="372">
<emphasis id="B906EA807A00FFC843F0FEE54017FEB0" bold="true" box="[199,314,258,284]" pageId="3" pageNumber="372">Diagnosis</emphasis>
. Carapace with metagastric region distinctly transversely depressed in both sexes (
<figureCitation id="13492A177A00FFC847D6FEE5444AFEB0" box="[1249,1383,258,284]" captionStart="FIGURE 2" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[193,1394,181,1919]" captionTargetId="figure-17@4.[193,1394,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876, habitus and colour in life; A, dorsal view; B, ventral view; C, frontal view. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.13618523" httpUri="https://zenodo.org/record/13618523/files/figure.png" pageId="3" pageNumber="372">Figs. 2A, C</figureCitation>
,
<figureCitation id="13492A177A00FFC84644FEE54181FEEC" captionStart="FIGURE 3" captionStartId="5.[151,250,1882,1906]" captionTargetBox="[151,1436,181,1858]" captionTargetId="figure-17@5.[151,1436,181,1858]" captionTargetPageId="5" captionText="FIGURE 3. Gustavus holthuisaesp. nov., ovigerous female paratype (7.3 × 4.0 mm) from Thuwal SaudiArabia, ZRC 2023.0580, habitus and colour in life; A, dorsal view; B, ventral view; C, frontal view. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.13618525" httpUri="https://zenodo.org/record/13618525/files/figure.png" pageId="3" pageNumber="372">3A, C</figureCitation>
,
<figureCitation id="13492A177A00FFC8438EFEC141D2FEEC" box="[185,255,294,320]" captionStart="FIGURE 4" captionStartId="6.[151,250,1851,1875]" captionTargetBox="[151,1435,181,1828]" captionTargetId="figure-17@6.[151,1435,181,1828]" captionTargetPageId="6" captionText="FIGURE 4. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, C, EG]; ovigerous female paratype (6.3 × 3.7 mm), FLMNH UF 72877, from type locality [B, D]; A, B, habitus in dorsal view; C, D, frontal margin and orbits, dorsal view; E, left G1, ventral view; F, left G1, dorsal view; G, left G2, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618527" httpUri="https://zenodo.org/record/13618527/files/figure.png" pageId="3" pageNumber="372">4A, B</figureCitation>
); male frontal width proportionately narrow, median lobes acutely triangular (
<figureCitation id="13492A177A00FFC84745FEC145E7FEEC" box="[1138,1226,294,320]" captionStart="FIGURE 4" captionStartId="6.[151,250,1851,1875]" captionTargetBox="[151,1435,181,1828]" captionTargetId="figure-17@6.[151,1435,181,1828]" captionTargetPageId="6" captionText="FIGURE 4. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, C, EG]; ovigerous female paratype (6.3 × 3.7 mm), FLMNH UF 72877, from type locality [B, D]; A, B, habitus in dorsal view; C, D, frontal margin and orbits, dorsal view; E, left G1, ventral view; F, left G1, dorsal view; G, left G2, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618527" httpUri="https://zenodo.org/record/13618527/files/figure.png" pageId="3" pageNumber="372">Fig. 4C</figureCitation>
,
<figureCitation id="13492A177A00FFC847E0FEC145D1FEED" box="[1239,1276,294,321]" captionStart="FIGURE 5" captionStartId="7.[151,250,1552,1576]" captionTargetBox="[151,1435,181,1528]" captionTargetId="figure-17@7.[151,1435,181,1528]" captionTargetPageId="7" captionText="FIGURE 5. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [AD]; ovigerous female paratype (6.3 × 3.7 mm) from type locality, FLMNH UF 72877 [E, F, H]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [G]; A, E, cephalothorax showing orbits, epistome, antennules and antennae, frontal view; B, right chela (partially denuded), outer view; C, male buccal cavity and anterior thoracic sternum, ventral view; D, male anterior thoracic sternum and sternopleonal cavity, showing intact right G1, ventral view; F, right chela (denuded), outer view; G, female pleon; H, female sternopleonal cavity and vulvae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618529" httpUri="https://zenodo.org/record/13618529/files/figure.png" pageId="3" pageNumber="372">5A</figureCitation>
); female front wider, median lobes low, separated by shallow concavity (
<figureCitation id="13492A177A00FFC84004FEAD42A3FEC8" box="[819,910,330,356]" captionStart="FIGURE 4" captionStartId="6.[151,250,1851,1875]" captionTargetBox="[151,1435,181,1828]" captionTargetId="figure-17@6.[151,1435,181,1828]" captionTargetPageId="6" captionText="FIGURE 4. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, C, EG]; ovigerous female paratype (6.3 × 3.7 mm), FLMNH UF 72877, from type locality [B, D]; A, B, habitus in dorsal view; C, D, frontal margin and orbits, dorsal view; E, left G1, ventral view; F, left G1, dorsal view; G, left G2, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618527" httpUri="https://zenodo.org/record/13618527/files/figure.png" pageId="3" pageNumber="372">Fig. 4D</figureCitation>
,
<figureCitation id="13492A177A00FFC840ACFEAD4293FEC9" box="[923,958,330,357]" captionStart="FIGURE 5" captionStartId="7.[151,250,1552,1576]" captionTargetBox="[151,1435,181,1528]" captionTargetId="figure-17@7.[151,1435,181,1528]" captionTargetPageId="7" captionText="FIGURE 5. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [AD]; ovigerous female paratype (6.3 × 3.7 mm) from type locality, FLMNH UF 72877 [E, F, H]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [G]; A, E, cephalothorax showing orbits, epistome, antennules and antennae, frontal view; B, right chela (partially denuded), outer view; C, male buccal cavity and anterior thoracic sternum, ventral view; D, male anterior thoracic sternum and sternopleonal cavity, showing intact right G1, ventral view; F, right chela (denuded), outer view; G, female pleon; H, female sternopleonal cavity and vulvae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618529" httpUri="https://zenodo.org/record/13618529/files/figure.png" pageId="3" pageNumber="372">5E</figureCitation>
). Mxp3 ischium with inner margin gently concave (
<figureCitation id="13492A177A00FFC84235FE894056FE24" box="[258,379,366,392]" captionStart="FIGURE 6" captionStartId="8.[151,250,1821,1845]" captionTargetBox="[158,1429,460,1791]" captionTargetId="figure-130@8.[151,1435,453,1798]" captionTargetPageId="8" captionText="FIGURE 6. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, CJ]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [B, KN]; A, B, right Mxp3 (setae denuded), outer view; CF, male P2P5, respectively, dorsal view; G, male pleon; H, left G1, ventral view (setae omitted); I, left G1, dorsal view (setae omitted); J, left G2, ventral view (setae omitted); KN, female P2P5, respectively, dorsal view. Scales: A, B, HJ = 0.2 mm; CG, KN = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618533" httpUri="https://zenodo.org/record/13618533/files/figure.png" pageId="3" pageNumber="372">Fig. 6A, B</figureCitation>
).
<collectingRegion id="49B6F8707A00FFC842A7FE8940E7FE24" box="[400,458,366,392]" country="Maldives" name="Maale" pageId="3" pageNumber="372">Male</collectingRegion>
and female P4 and P5 basis-ischium with 2 or 3 spines on flexor margin (
<figureCitation id="13492A177A00FFC84636FE894182FE00" captionStart="FIGURE 6" captionStartId="8.[151,250,1821,1845]" captionTargetBox="[158,1429,460,1791]" captionTargetId="figure-130@8.[151,1435,453,1798]" captionTargetPageId="8" captionText="FIGURE 6. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, CJ]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [B, KN]; A, B, right Mxp3 (setae denuded), outer view; CF, male P2P5, respectively, dorsal view; G, male pleon; H, left G1, ventral view (setae omitted); I, left G1, dorsal view (setae omitted); J, left G2, ventral view (setae omitted); KN, female P2P5, respectively, dorsal view. Scales: A, B, HJ = 0.2 mm; CG, KN = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618533" httpUri="https://zenodo.org/record/13618533/files/figure.png" pageId="3" pageNumber="372">Fig. 6E, F, M, N</figureCitation>
); male and female P4 merus with 1 spine on flexor margin (
<figureCitation id="13492A177A00FFC8407DFE7542EAFE00" box="[842,967,402,428]" captionStart="FIGURE 6" captionStartId="8.[151,250,1821,1845]" captionTargetBox="[158,1429,460,1791]" captionTargetId="figure-130@8.[151,1435,453,1798]" captionTargetPageId="8" captionText="FIGURE 6. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, CJ]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [B, KN]; A, B, right Mxp3 (setae denuded), outer view; CF, male P2P5, respectively, dorsal view; G, male pleon; H, left G1, ventral view (setae omitted); I, left G1, dorsal view (setae omitted); J, left G2, ventral view (setae omitted); KN, female P2P5, respectively, dorsal view. Scales: A, B, HJ = 0.2 mm; CG, KN = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618533" httpUri="https://zenodo.org/record/13618533/files/figure.png" pageId="3" pageNumber="372">Fig. 6E, M</figureCitation>
); male and female P5 merus with 2 spines on flexor margin (
<figureCitation id="13492A177A00FFC84253FE5140C8FE7C" box="[356,485,438,464]" captionStart="FIGURE 6" captionStartId="8.[151,250,1821,1845]" captionTargetBox="[158,1429,460,1791]" captionTargetId="figure-130@8.[151,1435,453,1798]" captionTargetPageId="8" captionText="FIGURE 6. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, CJ]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [B, KN]; A, B, right Mxp3 (setae denuded), outer view; CF, male P2P5, respectively, dorsal view; G, male pleon; H, left G1, ventral view (setae omitted); I, left G1, dorsal view (setae omitted); J, left G2, ventral view (setae omitted); KN, female P2P5, respectively, dorsal view. Scales: A, B, HJ = 0.2 mm; CG, KN = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618533" httpUri="https://zenodo.org/record/13618533/files/figure.png" pageId="3" pageNumber="372">Figs. 6F, N</figureCitation>
).
<collectingRegion id="49B6F8707A00FFC842CDFE514319FE7C" box="[506,564,438,464]" country="Maldives" name="Maale" pageId="3" pageNumber="372">Male</collectingRegion>
pleon with somites 46 immobile, with sutures visible between somites (
<figureCitation id="13492A177A00FFC84658FE514190FE58" captionStart="FIGURE 6" captionStartId="8.[151,250,1821,1845]" captionTargetBox="[158,1429,460,1791]" captionTargetId="figure-130@8.[151,1435,453,1798]" captionTargetPageId="8" captionText="FIGURE 6. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, CJ]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [B, KN]; A, B, right Mxp3 (setae denuded), outer view; CF, male P2P5, respectively, dorsal view; G, male pleon; H, left G1, ventral view (setae omitted); I, left G1, dorsal view (setae omitted); J, left G2, ventral view (setae omitted); KN, female P2P5, respectively, dorsal view. Scales: A, B, HJ = 0.2 mm; CG, KN = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618533" httpUri="https://zenodo.org/record/13618533/files/figure.png" pageId="3" pageNumber="372">Fig. 6G</figureCitation>
). G1 overall relatively stout, apex short, rounded, with sharp subdistal projection (
<figureCitation id="13492A177A00FFC84760FE3D45F9FE59" box="[1111,1236,474,501]" captionStart="FIGURE 4" captionStartId="6.[151,250,1851,1875]" captionTargetBox="[151,1435,181,1828]" captionTargetId="figure-17@6.[151,1435,181,1828]" captionTargetPageId="6" captionText="FIGURE 4. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, C, EG]; ovigerous female paratype (6.3 × 3.7 mm), FLMNH UF 72877, from type locality [B, D]; A, B, habitus in dorsal view; C, D, frontal margin and orbits, dorsal view; E, left G1, ventral view; F, left G1, dorsal view; G, left G2, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618527" httpUri="https://zenodo.org/record/13618527/files/figure.png" pageId="3" pageNumber="372">Figs. 4E, F</figureCitation>
,
<figureCitation id="13492A177A00FFC847E8FE3D4429FE59" box="[1247,1284,474,501]" captionStart="FIGURE 5" captionStartId="7.[151,250,1552,1576]" captionTargetBox="[151,1435,181,1528]" captionTargetId="figure-17@7.[151,1435,181,1528]" captionTargetPageId="7" captionText="FIGURE 5. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [AD]; ovigerous female paratype (6.3 × 3.7 mm) from type locality, FLMNH UF 72877 [E, F, H]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [G]; A, E, cephalothorax showing orbits, epistome, antennules and antennae, frontal view; B, right chela (partially denuded), outer view; C, male buccal cavity and anterior thoracic sternum, ventral view; D, male anterior thoracic sternum and sternopleonal cavity, showing intact right G1, ventral view; F, right chela (denuded), outer view; G, female pleon; H, female sternopleonal cavity and vulvae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618529" httpUri="https://zenodo.org/record/13618529/files/figure.png" pageId="3" pageNumber="372">5D</figureCitation>
,
<figureCitation id="13492A177A00FFC84638FE3D4467FE58" box="[1295,1354,474,500]" captionStart="FIGURE 6" captionStartId="8.[151,250,1821,1845]" captionTargetBox="[158,1429,460,1791]" captionTargetId="figure-130@8.[151,1435,453,1798]" captionTargetPageId="8" captionText="FIGURE 6. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, CJ]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [B, KN]; A, B, right Mxp3 (setae denuded), outer view; CF, male P2P5, respectively, dorsal view; G, male pleon; H, left G1, ventral view (setae omitted); I, left G1, dorsal view (setae omitted); J, left G2, ventral view (setae omitted); KN, female P2P5, respectively, dorsal view. Scales: A, B, HJ = 0.2 mm; CG, KN = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618533" httpUri="https://zenodo.org/record/13618533/files/figure.png" pageId="3" pageNumber="372">6H, I</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36865197A00FFC243F0FE19438DFB69" lastPageId="9" lastPageNumber="378" pageId="3" pageNumber="372" type="description">
<paragraph id="8BCD36927A00FFC843F0FE194465FC09" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="372">
<emphasis id="B906EA807A00FFC843F0FE19407CFDB4" bold="true" box="[199,337,510,536]" pageId="3" pageNumber="372">Description</emphasis>
.
<emphasis id="B906EA807A00FFC84253FE194088FDB4" box="[356,421,510,536]" italics="true" pageId="3" pageNumber="372">
<collectingRegion id="49B6F8707A00FFC84253FE19408CFDB4" box="[356,417,510,536]" country="Maldives" name="Maale" pageId="3" pageNumber="372">Male</collectingRegion>
.
</emphasis>
Carapace subpentagonal, broader than long, width to length ratio 1.56; dorsal surface almost smooth, regions poorly demarcated, with numerous, small, shallow pits (
<figureCitation id="13492A177A00FFC84724FDC5455AFD90" box="[1043,1143,546,572]" captionStart="FIGURE 2" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[193,1394,181,1919]" captionTargetId="figure-17@4.[193,1394,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876, habitus and colour in life; A, dorsal view; B, ventral view; C, frontal view. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.13618523" httpUri="https://zenodo.org/record/13618523/files/figure.png" pageId="3" pageNumber="372">Figs. 2A</figureCitation>
,
<figureCitation id="13492A177A00FFC847B5FDC54585FD90" box="[1154,1192,546,572]" captionStart="FIGURE 4" captionStartId="6.[151,250,1851,1875]" captionTargetBox="[151,1435,181,1828]" captionTargetId="figure-17@6.[151,1435,181,1828]" captionTargetPageId="6" captionText="FIGURE 4. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, C, EG]; ovigerous female paratype (6.3 × 3.7 mm), FLMNH UF 72877, from type locality [B, D]; A, B, habitus in dorsal view; C, D, frontal margin and orbits, dorsal view; E, left G1, ventral view; F, left G1, dorsal view; G, left G2, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618527" httpUri="https://zenodo.org/record/13618527/files/figure.png" pageId="3" pageNumber="372">4A</figureCitation>
); surface and margins finely pubescent; short, dense setae present on front, partially obscuring margin; metagastric region transversely depressed, covered with low, soft, easily detachable setae (
<figureCitation id="13492A177A00FFC84077FD8D42E6FD28" box="[832,971,618,644]" captionStart="FIGURE 2" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[193,1394,181,1919]" captionTargetId="figure-17@4.[193,1394,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876, habitus and colour in life; A, dorsal view; B, ventral view; C, frontal view. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.13618523" httpUri="https://zenodo.org/record/13618523/files/figure.png" pageId="3" pageNumber="372">Figs. 2A, C</figureCitation>
,
<figureCitation id="13492A177A00FFC840EEFD8D452DFD28" box="[985,1024,618,644]" captionStart="FIGURE 4" captionStartId="6.[151,250,1851,1875]" captionTargetBox="[151,1435,181,1828]" captionTargetId="figure-17@6.[151,1435,181,1828]" captionTargetPageId="6" captionText="FIGURE 4. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, C, EG]; ovigerous female paratype (6.3 × 3.7 mm), FLMNH UF 72877, from type locality [B, D]; A, B, habitus in dorsal view; C, D, frontal margin and orbits, dorsal view; E, left G1, ventral view; F, left G1, dorsal view; G, left G2, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618527" httpUri="https://zenodo.org/record/13618527/files/figure.png" pageId="3" pageNumber="372">4A</figureCitation>
); anterolateral margin entire, gently convex laterally, subcristate, continuing onto posterolateral margin but not supraorbital margin (
<figureCitation id="13492A177A00FFC847C4FD694477FD04" box="[1267,1370,654,680]" captionStart="FIGURE 2" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[193,1394,181,1919]" captionTargetId="figure-17@4.[193,1394,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876, habitus and colour in life; A, dorsal view; B, ventral view; C, frontal view. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.13618523" httpUri="https://zenodo.org/record/13618523/files/figure.png" pageId="3" pageNumber="372">Figs. 2A</figureCitation>
,
<figureCitation id="13492A177A00FFC8465DFD6944BDFD04" box="[1386,1424,654,680]" captionStart="FIGURE 4" captionStartId="6.[151,250,1851,1875]" captionTargetBox="[151,1435,181,1828]" captionTargetId="figure-17@6.[151,1435,181,1828]" captionTargetPageId="6" captionText="FIGURE 4. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, C, EG]; ovigerous female paratype (6.3 × 3.7 mm), FLMNH UF 72877, from type locality [B, D]; A, B, habitus in dorsal view; C, D, frontal margin and orbits, dorsal view; E, left G1, ventral view; F, left G1, dorsal view; G, left G2, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618527" httpUri="https://zenodo.org/record/13618527/files/figure.png" pageId="3" pageNumber="372">4A</figureCitation>
). Front deflexed, medially emarginate in dorsal view; median lobes triangular, separated by distinct cleft, triangular in anterior view; lateral tooth distinct in dorsal view (
<figureCitation id="13492A177A00FFC841FAFD31421DFD5C" box="[717,816,726,752]" captionStart="FIGURE 4" captionStartId="6.[151,250,1851,1875]" captionTargetBox="[151,1435,181,1828]" captionTargetId="figure-17@6.[151,1435,181,1828]" captionTargetPageId="6" captionText="FIGURE 4. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, C, EG]; ovigerous female paratype (6.3 × 3.7 mm), FLMNH UF 72877, from type locality [B, D]; A, B, habitus in dorsal view; C, D, frontal margin and orbits, dorsal view; E, left G1, ventral view; F, left G1, dorsal view; G, left G2, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618527" httpUri="https://zenodo.org/record/13618527/files/figure.png" pageId="3" pageNumber="372">Figs. 4C</figureCitation>
,
<figureCitation id="13492A177A00FFC8400CFD31424CFD5D" box="[827,865,726,753]" captionStart="FIGURE 5" captionStartId="7.[151,250,1552,1576]" captionTargetBox="[151,1435,181,1528]" captionTargetId="figure-17@7.[151,1435,181,1528]" captionTargetPageId="7" captionText="FIGURE 5. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [AD]; ovigerous female paratype (6.3 × 3.7 mm) from type locality, FLMNH UF 72877 [E, F, H]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [G]; A, E, cephalothorax showing orbits, epistome, antennules and antennae, frontal view; B, right chela (partially denuded), outer view; C, male buccal cavity and anterior thoracic sternum, ventral view; D, male anterior thoracic sternum and sternopleonal cavity, showing intact right G1, ventral view; F, right chela (denuded), outer view; G, female pleon; H, female sternopleonal cavity and vulvae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618529" httpUri="https://zenodo.org/record/13618529/files/figure.png" pageId="3" pageNumber="372">5A</figureCitation>
). Orbital margin entire, slightly narrowed laterally; infraorbital margin terminating mesially as rounded angle; supraorbital margin terminating mesially as small triangular, ventrally directed tooth (
<figureCitation id="13492A177A00FFC84113FCF943ABFC94" box="[548,646,798,824]" captionStart="FIGURE 4" captionStartId="6.[151,250,1851,1875]" captionTargetBox="[151,1435,181,1828]" captionTargetId="figure-17@6.[151,1435,181,1828]" captionTargetPageId="6" captionText="FIGURE 4. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, C, EG]; ovigerous female paratype (6.3 × 3.7 mm), FLMNH UF 72877, from type locality [B, D]; A, B, habitus in dorsal view; C, D, frontal margin and orbits, dorsal view; E, left G1, ventral view; F, left G1, dorsal view; G, left G2, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618527" httpUri="https://zenodo.org/record/13618527/files/figure.png" pageId="3" pageNumber="372">Figs. 4C</figureCitation>
,
<figureCitation id="13492A177A00FFC841A6FCF9439AFC95" box="[657,695,798,825]" captionStart="FIGURE 5" captionStartId="7.[151,250,1552,1576]" captionTargetBox="[151,1435,181,1528]" captionTargetId="figure-17@7.[151,1435,181,1528]" captionTargetPageId="7" captionText="FIGURE 5. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [AD]; ovigerous female paratype (6.3 × 3.7 mm) from type locality, FLMNH UF 72877 [E, F, H]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [G]; A, E, cephalothorax showing orbits, epistome, antennules and antennae, frontal view; B, right chela (partially denuded), outer view; C, male buccal cavity and anterior thoracic sternum, ventral view; D, male anterior thoracic sternum and sternopleonal cavity, showing intact right G1, ventral view; F, right chela (denuded), outer view; G, female pleon; H, female sternopleonal cavity and vulvae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618529" httpUri="https://zenodo.org/record/13618529/files/figure.png" pageId="3" pageNumber="372">5A</figureCitation>
). Epistome very short, partially sunken medially; posterior margin gently concave, entire (
<figureCitation id="13492A177A00FFC84294FCA5432DFCF1" box="[419,512,834,861]" captionStart="FIGURE 5" captionStartId="7.[151,250,1552,1576]" captionTargetBox="[151,1435,181,1528]" captionTargetId="figure-17@7.[151,1435,181,1528]" captionTargetPageId="7" captionText="FIGURE 5. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [AD]; ovigerous female paratype (6.3 × 3.7 mm) from type locality, FLMNH UF 72877 [E, F, H]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [G]; A, E, cephalothorax showing orbits, epistome, antennules and antennae, frontal view; B, right chela (partially denuded), outer view; C, male buccal cavity and anterior thoracic sternum, ventral view; D, male anterior thoracic sternum and sternopleonal cavity, showing intact right G1, ventral view; F, right chela (denuded), outer view; G, female pleon; H, female sternopleonal cavity and vulvae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618529" httpUri="https://zenodo.org/record/13618529/files/figure.png" pageId="3" pageNumber="372">Fig. 5A</figureCitation>
). Eyes mobile, filling orbit; cornea simple, pigmented (
<figureCitation id="13492A177A00FFC8474AFCA545FBFCF0" box="[1149,1238,834,860]" captionStart="FIGURE 4" captionStartId="6.[151,250,1851,1875]" captionTargetBox="[151,1435,181,1828]" captionTargetId="figure-17@6.[151,1435,181,1828]" captionTargetPageId="6" captionText="FIGURE 4. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, C, EG]; ovigerous female paratype (6.3 × 3.7 mm), FLMNH UF 72877, from type locality [B, D]; A, B, habitus in dorsal view; C, D, frontal margin and orbits, dorsal view; E, left G1, ventral view; F, left G1, dorsal view; G, left G2, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618527" httpUri="https://zenodo.org/record/13618527/files/figure.png" pageId="3" pageNumber="372">Fig. 4C</figureCitation>
,
<figureCitation id="13492A177A00FFC847D3FCA54427FCF1" box="[1252,1290,834,861]" captionStart="FIGURE 5" captionStartId="7.[151,250,1552,1576]" captionTargetBox="[151,1435,181,1528]" captionTargetId="figure-17@7.[151,1435,181,1528]" captionTargetPageId="7" captionText="FIGURE 5. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [AD]; ovigerous female paratype (6.3 × 3.7 mm) from type locality, FLMNH UF 72877 [E, F, H]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [G]; A, E, cephalothorax showing orbits, epistome, antennules and antennae, frontal view; B, right chela (partially denuded), outer view; C, male buccal cavity and anterior thoracic sternum, ventral view; D, male anterior thoracic sternum and sternopleonal cavity, showing intact right G1, ventral view; F, right chela (denuded), outer view; G, female pleon; H, female sternopleonal cavity and vulvae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618529" httpUri="https://zenodo.org/record/13618529/files/figure.png" pageId="3" pageNumber="372">5A</figureCitation>
). Antennular fossae large; antennule folding obliquely, filling fossa (
<figureCitation id="13492A177A00FFC84020FC814259FC2D" box="[791,884,870,897]" captionStart="FIGURE 5" captionStartId="7.[151,250,1552,1576]" captionTargetBox="[151,1435,181,1528]" captionTargetId="figure-17@7.[151,1435,181,1528]" captionTargetPageId="7" captionText="FIGURE 5. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [AD]; ovigerous female paratype (6.3 × 3.7 mm) from type locality, FLMNH UF 72877 [E, F, H]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [G]; A, E, cephalothorax showing orbits, epistome, antennules and antennae, frontal view; B, right chela (partially denuded), outer view; C, male buccal cavity and anterior thoracic sternum, ventral view; D, male anterior thoracic sternum and sternopleonal cavity, showing intact right G1, ventral view; F, right chela (denuded), outer view; G, female pleon; H, female sternopleonal cavity and vulvae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618529" httpUri="https://zenodo.org/record/13618529/files/figure.png" pageId="3" pageNumber="372">Fig. 5A</figureCitation>
). Antenna short, fully entering orbit, with basal antennal article stout, quadrate, not fused to carapace, not reaching distolateral angle of carapace (
<figureCitation id="13492A177A00FFC847D6FC6D4416FC09" box="[1249,1339,906,933]" captionStart="FIGURE 5" captionStartId="7.[151,250,1552,1576]" captionTargetBox="[151,1435,181,1528]" captionTargetId="figure-17@7.[151,1435,181,1528]" captionTargetPageId="7" captionText="FIGURE 5. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [AD]; ovigerous female paratype (6.3 × 3.7 mm) from type locality, FLMNH UF 72877 [E, F, H]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [G]; A, E, cephalothorax showing orbits, epistome, antennules and antennae, frontal view; B, right chela (partially denuded), outer view; C, male buccal cavity and anterior thoracic sternum, ventral view; D, male anterior thoracic sternum and sternopleonal cavity, showing intact right G1, ventral view; F, right chela (denuded), outer view; G, female pleon; H, female sternopleonal cavity and vulvae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618529" httpUri="https://zenodo.org/record/13618529/files/figure.png" pageId="3" pageNumber="372">Fig. 5A</figureCitation>
).
</paragraph>
<paragraph id="8BCD36927A00FFC843F0FC494327FBF4" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="372">
Mxp3 covering most of buccal cavern when closed; ischium distinctly longer than merus, slender, subtriangular, distomesial angle produced into rounded lobe extending mesially beyond level of mesial meral margin, inner margin gently concave; merus smaller than ischium, slightly wider than long; palp articulating on distal margin of merus; basal part of exopod relatively slender, reaching almost to just above merus; exopod flagellum short, not reaching distal margin of merus (
<figureCitation id="13492A177A00FFC84294FBD940D0FBF4" box="[419,509,1086,1112]" captionStart="FIGURE 6" captionStartId="8.[151,250,1821,1845]" captionTargetBox="[158,1429,460,1791]" captionTargetId="figure-130@8.[151,1435,453,1798]" captionTargetPageId="8" captionText="FIGURE 6. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, CJ]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [B, KN]; A, B, right Mxp3 (setae denuded), outer view; CF, male P2P5, respectively, dorsal view; G, male pleon; H, left G1, ventral view (setae omitted); I, left G1, dorsal view (setae omitted); J, left G2, ventral view (setae omitted); KN, female P2P5, respectively, dorsal view. Scales: A, B, HJ = 0.2 mm; CG, KN = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618533" httpUri="https://zenodo.org/record/13618533/files/figure.png" pageId="3" pageNumber="372">Fig. 6A</figureCitation>
).
</paragraph>
<paragraph id="8BCD36927A00FFC843F0FB854075FAF9" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="372">
Chelipeds equal, covered with short, fine setae partially obscuring margins and surface (
<figureCitation id="13492A177A00FFC847E6FB854477FBD0" box="[1233,1370,1122,1148]" captionStart="FIGURE 2" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[193,1394,181,1919]" captionTargetId="figure-17@4.[193,1394,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876, habitus and colour in life; A, dorsal view; B, ventral view; C, frontal view. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.13618523" httpUri="https://zenodo.org/record/13618523/files/figure.png" pageId="3" pageNumber="372">Figs. 2AC</figureCitation>
,
<figureCitation id="13492A177A00FFC8465EFB8544A2FBD0" box="[1385,1423,1122,1148]" captionStart="FIGURE 4" captionStartId="6.[151,250,1851,1875]" captionTargetBox="[151,1435,181,1828]" captionTargetId="figure-17@6.[151,1435,181,1828]" captionTargetPageId="6" captionText="FIGURE 4. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, C, EG]; ovigerous female paratype (6.3 × 3.7 mm), FLMNH UF 72877, from type locality [B, D]; A, B, habitus in dorsal view; C, D, frontal margin and orbits, dorsal view; E, left G1, ventral view; F, left G1, dorsal view; G, left G2, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618527" httpUri="https://zenodo.org/record/13618527/files/figure.png" pageId="3" pageNumber="372">4A</figureCitation>
); merus triangular in cross-section, with plumose setae proximally; carpus smooth, inner angle rounded, with sparser plumose setae; chela surfaces relatively short, surface smooth, except for scattered, low, rounded granules and punctae on lower portion of palm; palm longer than dactylus, ventral margin near pollex concave; pollex with convex occlusal margin, with blunt teeth proximally, distal half with small teeth fused basally, forming blade-like structure; dactylus with occlusal margin armed with 3 low, blunt teeth proximally, distal part unarmed, tip strongly curved (
<figureCitation id="13492A177A00FFC843C4FADD4066FAF9" box="[243,331,1338,1365]" captionStart="FIGURE 5" captionStartId="7.[151,250,1552,1576]" captionTargetBox="[151,1435,181,1528]" captionTargetId="figure-17@7.[151,1435,181,1528]" captionTargetPageId="7" captionText="FIGURE 5. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [AD]; ovigerous female paratype (6.3 × 3.7 mm) from type locality, FLMNH UF 72877 [E, F, H]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [G]; A, E, cephalothorax showing orbits, epistome, antennules and antennae, frontal view; B, right chela (partially denuded), outer view; C, male buccal cavity and anterior thoracic sternum, ventral view; D, male anterior thoracic sternum and sternopleonal cavity, showing intact right G1, ventral view; F, right chela (denuded), outer view; G, female pleon; H, female sternopleonal cavity and vulvae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618529" httpUri="https://zenodo.org/record/13618529/files/figure.png" pageId="3" pageNumber="372">Fig. 5B</figureCitation>
).
</paragraph>
<paragraph id="8BCD36927A00FFC843F0FAB945E9F9FC" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="372">
P2P5 relatively short, P3 longest, P5 shortest; P5 dorsal to other pereopods (
<figureCitation id="13492A177A00FFC84723FAB9455BFAD4" box="[1044,1142,1374,1400]" captionStart="FIGURE 2" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[193,1394,181,1919]" captionTargetId="figure-17@4.[193,1394,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876, habitus and colour in life; A, dorsal view; B, ventral view; C, frontal view. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.13618523" httpUri="https://zenodo.org/record/13618523/files/figure.png" pageId="3" pageNumber="372">Figs. 2A</figureCitation>
,
<figureCitation id="13492A177A00FFC847B7FAB9458BFAD4" box="[1152,1190,1374,1400]" captionStart="FIGURE 4" captionStartId="6.[151,250,1851,1875]" captionTargetBox="[151,1435,181,1828]" captionTargetId="figure-17@6.[151,1435,181,1828]" captionTargetPageId="6" captionText="FIGURE 4. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, C, EG]; ovigerous female paratype (6.3 × 3.7 mm), FLMNH UF 72877, from type locality [B, D]; A, B, habitus in dorsal view; C, D, frontal margin and orbits, dorsal view; E, left G1, ventral view; F, left G1, dorsal view; G, left G2, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618527" httpUri="https://zenodo.org/record/13618527/files/figure.png" pageId="3" pageNumber="372">4A</figureCitation>
); propodus and carpus covered with short, fine tomentum partially obscuring surface and margins; merus with plumose setae along flexor and extensor margins; P2 and P3 merus longer than carpus and propodus combined; P4 and P5 merus subequal to carpus and propodus combined; dactylus short, stout, unarmed laterally, with corneous apex (
<figureCitation id="13492A177A00FFC847F7FA2D4414FA48" box="[1216,1337,1482,1508]" captionStart="FIGURE 6" captionStartId="8.[151,250,1821,1845]" captionTargetBox="[158,1429,460,1791]" captionTargetId="figure-130@8.[151,1435,453,1798]" captionTargetPageId="8" captionText="FIGURE 6. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, CJ]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [B, KN]; A, B, right Mxp3 (setae denuded), outer view; CF, male P2P5, respectively, dorsal view; G, male pleon; H, left G1, ventral view (setae omitted); I, left G1, dorsal view (setae omitted); J, left G2, ventral view (setae omitted); KN, female P2P5, respectively, dorsal view. Scales: A, B, HJ = 0.2 mm; CG, KN = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618533" httpUri="https://zenodo.org/record/13618533/files/figure.png" pageId="3" pageNumber="372">Fig. 6CF</figureCitation>
). P2 and P3 merus with margins almost entire (
<figureCitation id="13492A177A00FFC84178FA0943E2F9A4" box="[591,719,1518,1544]" captionStart="FIGURE 6" captionStartId="8.[151,250,1821,1845]" captionTargetBox="[158,1429,460,1791]" captionTargetId="figure-130@8.[151,1435,453,1798]" captionTargetPageId="8" captionText="FIGURE 6. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, CJ]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [B, KN]; A, B, right Mxp3 (setae denuded), outer view; CF, male P2P5, respectively, dorsal view; G, male pleon; H, left G1, ventral view (setae omitted); I, left G1, dorsal view (setae omitted); J, left G2, ventral view (setae omitted); KN, female P2P5, respectively, dorsal view. Scales: A, B, HJ = 0.2 mm; CG, KN = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618533" httpUri="https://zenodo.org/record/13618533/files/figure.png" pageId="3" pageNumber="372">Fig. 6C, D</figureCitation>
). P4 and P5 ischium each armed with 2 stout spines on flexor margin; P4 merus with 1 larger spine on flexor margin; P5 merus with 2 spines on flexor margin (
<figureCitation id="13492A177A00FFC847FAF9F5446DF980" box="[1229,1344,1554,1580]" captionStart="FIGURE 6" captionStartId="8.[151,250,1821,1845]" captionTargetBox="[158,1429,460,1791]" captionTargetId="figure-130@8.[151,1435,453,1798]" captionTargetPageId="8" captionText="FIGURE 6. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, CJ]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [B, KN]; A, B, right Mxp3 (setae denuded), outer view; CF, male P2P5, respectively, dorsal view; G, male pleon; H, left G1, ventral view (setae omitted); I, left G1, dorsal view (setae omitted); J, left G2, ventral view (setae omitted); KN, female P2P5, respectively, dorsal view. Scales: A, B, HJ = 0.2 mm; CG, KN = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618533" httpUri="https://zenodo.org/record/13618533/files/figure.png" pageId="3" pageNumber="372">Fig. 6E, F</figureCitation>
). P2P5 propodus with 2 or 3 small, fixed spines on distal flexor margin opposing dactylus (
<figureCitation id="13492A177A00FFC84777F9D1459AF9FC" box="[1088,1207,1590,1616]" captionStart="FIGURE 6" captionStartId="8.[151,250,1821,1845]" captionTargetBox="[158,1429,460,1791]" captionTargetId="figure-130@8.[151,1435,453,1798]" captionTargetPageId="8" captionText="FIGURE 6. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, CJ]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [B, KN]; A, B, right Mxp3 (setae denuded), outer view; CF, male P2P5, respectively, dorsal view; G, male pleon; H, left G1, ventral view (setae omitted); I, left G1, dorsal view (setae omitted); J, left G2, ventral view (setae omitted); KN, female P2P5, respectively, dorsal view. Scales: A, B, HJ = 0.2 mm; CG, KN = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618533" httpUri="https://zenodo.org/record/13618533/files/figure.png" pageId="3" pageNumber="372">Fig. 6CF</figureCitation>
).
</paragraph>
<paragraph id="8BCD36927A00FFC843F0F9BD4302F8A8" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="372">
Thoracic sternum relatively wide; sternites 13 completely fused, sternites 1 and 2 demarcated only by row of setae; sternites 3 and 4 partially fused, separated by shallow median groove lined with setae; sternopleonal cavity wide, deep, its distal part subcristate, tip reaching to imaginary line joining subproximal edge of cheliped coxae; proximal third of sternite 5 with rounded tubercle representing pleonal locking mechanism (
<figureCitation id="13492A177A00FFC8479FF9214408F94C" box="[1192,1317,1734,1761]" captionStart="FIGURE 5" captionStartId="7.[151,250,1552,1576]" captionTargetBox="[151,1435,181,1528]" captionTargetId="figure-17@7.[151,1435,181,1528]" captionTargetPageId="7" captionText="FIGURE 5. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [AD]; ovigerous female paratype (6.3 × 3.7 mm) from type locality, FLMNH UF 72877 [E, F, H]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [G]; A, E, cephalothorax showing orbits, epistome, antennules and antennae, frontal view; B, right chela (partially denuded), outer view; C, male buccal cavity and anterior thoracic sternum, ventral view; D, male anterior thoracic sternum and sternopleonal cavity, showing intact right G1, ventral view; F, right chela (denuded), outer view; G, female pleon; H, female sternopleonal cavity and vulvae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618529" httpUri="https://zenodo.org/record/13618529/files/figure.png" pageId="3" pageNumber="372">Fig. 5C, D</figureCitation>
). Thoracic sternite 8 visible when pleon closed.
</paragraph>
<paragraph id="8BCD36927A00FFC843F0F8E94475F8E0" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="372">
Pleon broadly triangular; somites 46 with sutures visible but functionally fused, immobile; somites 13 subequal in width; somites 4 and 5 trapezoidal; somite 6 subrectangular; telson wide, semicircular (
<figureCitation id="13492A177A00FFC847C6F8D54466F8E0" box="[1265,1355,1842,1868]" captionStart="FIGURE 6" captionStartId="8.[151,250,1821,1845]" captionTargetBox="[158,1429,460,1791]" captionTargetId="figure-130@8.[151,1435,453,1798]" captionTargetPageId="8" captionText="FIGURE 6. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, CJ]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [B, KN]; A, B, right Mxp3 (setae denuded), outer view; CF, male P2P5, respectively, dorsal view; G, male pleon; H, left G1, ventral view (setae omitted); I, left G1, dorsal view (setae omitted); J, left G2, ventral view (setae omitted); KN, female P2P5, respectively, dorsal view. Scales: A, B, HJ = 0.2 mm; CG, KN = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618533" httpUri="https://zenodo.org/record/13618533/files/figure.png" pageId="3" pageNumber="372">Fig. 6G</figureCitation>
).
</paragraph>
<paragraph id="8BCD36927A00FFC843F0F8B142E4F870" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="372">
G1 relatively stout, with tip directed mesially (
<figureCitation id="13492A177A00FFC841FFF8B1420CF8DD" box="[712,801,1878,1905]" captionStart="FIGURE 5" captionStartId="7.[151,250,1552,1576]" captionTargetBox="[151,1435,181,1528]" captionTargetId="figure-17@7.[151,1435,181,1528]" captionTargetPageId="7" captionText="FIGURE 5. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [AD]; ovigerous female paratype (6.3 × 3.7 mm) from type locality, FLMNH UF 72877 [E, F, H]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [G]; A, E, cephalothorax showing orbits, epistome, antennules and antennae, frontal view; B, right chela (partially denuded), outer view; C, male buccal cavity and anterior thoracic sternum, ventral view; D, male anterior thoracic sternum and sternopleonal cavity, showing intact right G1, ventral view; F, right chela (denuded), outer view; G, female pleon; H, female sternopleonal cavity and vulvae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618529" httpUri="https://zenodo.org/record/13618529/files/figure.png" pageId="3" pageNumber="372">Fig. 5D</figureCitation>
); distal quarter distinctly bent at about 45° to longitudinal axis; distally tapering gradually to short, rounded apex, with numerous stiff setae; with short, sharp subdistal projection on lower margin; groove for G2 present on mesioventral surface (
<figureCitation id="13492A177A00FFC840D2F8794572F815" box="[997,1119,1950,1977]" captionStart="FIGURE 4" captionStartId="6.[151,250,1851,1875]" captionTargetBox="[151,1435,181,1828]" captionTargetId="figure-17@6.[151,1435,181,1828]" captionTargetPageId="6" captionText="FIGURE 4. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, C, EG]; ovigerous female paratype (6.3 × 3.7 mm), FLMNH UF 72877, from type locality [B, D]; A, B, habitus in dorsal view; C, D, frontal margin and orbits, dorsal view; E, left G1, ventral view; F, left G1, dorsal view; G, left G2, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618527" httpUri="https://zenodo.org/record/13618527/files/figure.png" pageId="3" pageNumber="372">Figs. 4E, F</figureCitation>
,
<figureCitation id="13492A177A00FFC8475DF8794589F814" box="[1130,1188,1950,1976]" captionStart="FIGURE 6" captionStartId="8.[151,250,1821,1845]" captionTargetBox="[158,1429,460,1791]" captionTargetId="figure-130@8.[151,1435,453,1798]" captionTargetPageId="8" captionText="FIGURE 6. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, CJ]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [B, KN]; A, B, right Mxp3 (setae denuded), outer view; CF, male P2P5, respectively, dorsal view; G, male pleon; H, left G1, ventral view (setae omitted); I, left G1, dorsal view (setae omitted); J, left G2, ventral view (setae omitted); KN, female P2P5, respectively, dorsal view. Scales: A, B, HJ = 0.2 mm; CG, KN = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618533" httpUri="https://zenodo.org/record/13618533/files/figure.png" pageId="3" pageNumber="372">6H, I</figureCitation>
). G2 short, about onethird length of G1; distally spatulate, apex bluntly angular (
<figureCitation id="13492A177A00FFC84007F82542B9F870" box="[816,916,1986,2012]" captionStart="FIGURE 4" captionStartId="6.[151,250,1851,1875]" captionTargetBox="[151,1435,181,1828]" captionTargetId="figure-17@6.[151,1435,181,1828]" captionTargetPageId="6" captionText="FIGURE 4. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, C, EG]; ovigerous female paratype (6.3 × 3.7 mm), FLMNH UF 72877, from type locality [B, D]; A, B, habitus in dorsal view; C, D, frontal margin and orbits, dorsal view; E, left G1, ventral view; F, left G1, dorsal view; G, left G2, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618527" httpUri="https://zenodo.org/record/13618527/files/figure.png" pageId="3" pageNumber="372">Figs. 4G</figureCitation>
,
<figureCitation id="13492A177A00FFC840A8F8254291F870" box="[927,956,1986,2012]" captionStart="FIGURE 6" captionStartId="8.[151,250,1821,1845]" captionTargetBox="[158,1429,460,1791]" captionTargetId="figure-130@8.[151,1435,453,1798]" captionTargetPageId="8" captionText="FIGURE 6. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, CJ]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [B, KN]; A, B, right Mxp3 (setae denuded), outer view; CF, male P2P5, respectively, dorsal view; G, male pleon; H, left G1, ventral view (setae omitted); I, left G1, dorsal view (setae omitted); J, left G2, ventral view (setae omitted); KN, female P2P5, respectively, dorsal view. Scales: A, B, HJ = 0.2 mm; CG, KN = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618533" httpUri="https://zenodo.org/record/13618533/files/figure.png" pageId="3" pageNumber="372">6J</figureCitation>
).
</paragraph>
<caption id="DF0D661A7A07FFCF43A0F87045B8F878" ID-DOI="http://doi.org/10.5281/zenodo.13618523" ID-Zenodo-Dep="13618523" httpUri="https://zenodo.org/record/13618523/files/figure.png" pageId="4" pageNumber="373" startId="4.[151,250,1943,1967]" targetBox="[193,1394,181,1919]" targetPageId="4" targetType="figure">
<paragraph id="8BCD36927A07FFCF43A0F87045B8F878" blockId="4.[151,1436,1943,2004]" pageId="4" pageNumber="373">
<emphasis id="B906EA807A07FFCF43A0F870403DF803" bold="true" box="[151,272,1943,1967]" pageId="4" pageNumber="373">FIGURE 2</emphasis>
.
<taxonomicName id="4C724D117A07FFCF4229F87F40DDF81C" authorityName="Ng &amp; Anker" authorityYear="2024" box="[286,496,1944,1968]" class="Malacostraca" family="Aphanodactylidae" genus="Gustavus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="373" phylum="Arthropoda" rank="species" species="holthuisae" status="sp. nov.">
<emphasis id="B906EA807A07FFCF4229F87F40DDF81C" box="[286,496,1944,1968]" italics="true" pageId="4" pageNumber="373">Gustavus holthuisae</emphasis>
</taxonomicName>
<emphasis id="B906EA807A07FFCF42CFF87F437FF81C" bold="true" box="[504,594,1944,1968]" pageId="4" pageNumber="373">
<taxonomicNameLabel id="A23557FB7A07FFCF42CFF87F4360F81C" box="[504,589,1944,1968]" pageId="4" pageNumber="373" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876, habitus and colour in life; A, dorsal view; B, ventral view; C, frontal view. Photographs by A. Anker.
</paragraph>
</caption>
<caption id="DF0D661A7A06FFCE43A0F8BD45B8F83A" ID-DOI="http://doi.org/10.5281/zenodo.13618525" ID-Zenodo-Dep="13618525" httpUri="https://zenodo.org/record/13618525/files/figure.png" pageId="5" pageNumber="374" startId="5.[151,250,1882,1906]" targetBox="[151,1436,181,1858]" targetPageId="5" targetType="figure">
<paragraph id="8BCD36927A06FFCE43A0F8BD45B8F83A" blockId="5.[151,1437,1882,1943]" pageId="5" pageNumber="374">
<emphasis id="B906EA807A06FFCE43A0F8BD4026F8DE" bold="true" box="[151,267,1882,1906]" pageId="5" pageNumber="374">FIGURE 3</emphasis>
.
<taxonomicName id="4C724D117A06FFCE4222F8BC40CFF8DE" authorityName="Ng &amp; Anker" authorityYear="2024" box="[277,482,1882,1906]" class="Malacostraca" family="Aphanodactylidae" genus="Gustavus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="holthuisae" status="sp. nov.">
<emphasis id="B906EA807A06FFCE4222F8BC40CFF8DE" box="[277,482,1882,1906]" italics="true" pageId="5" pageNumber="374">Gustavus holthuisae</emphasis>
</taxonomicName>
<emphasis id="B906EA807A06FFCE42D2F8BC4319F8DF" bold="true" box="[485,564,1883,1907]" pageId="5" pageNumber="374">
<taxonomicNameLabel id="A23557FB7A06FFCE42D2F8BC4319F8DF" box="[485,564,1883,1907]" pageId="5" pageNumber="374" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ovigerous female paratype (7.3 × 4.0 mm) from Thuwal Saudi Arabia, ZRC 2023.0580, habitus and colour in life; A, dorsal view; B, ventral view; C, frontal view. Photographs by A. Anker.
</paragraph>
</caption>
<caption id="DF0D661A7A05FFCD43A0F8DC440FF830" ID-DOI="http://doi.org/10.5281/zenodo.13618527" ID-Zenodo-Dep="13618527" httpUri="https://zenodo.org/record/13618527/files/figure.png" pageId="6" pageNumber="375" startId="6.[151,250,1851,1875]" targetBox="[151,1435,181,1828]" targetPageId="6" targetType="figure">
<paragraph id="8BCD36927A05FFCD43A0F8DC440FF830" blockId="6.[151,1436,1851,1948]" pageId="6" pageNumber="375">
<emphasis id="B906EA807A05FFCD43A0F8DC4021F8F8" bold="true" box="[151,268,1851,1876]" pageId="6" pageNumber="375">FIGURE 4</emphasis>
.
<taxonomicName id="4C724D117A05FFCD422FF8DB40CBF8F8" authorityName="Ng &amp; Anker" authorityYear="2024" box="[280,486,1852,1876]" class="Malacostraca" family="Aphanodactylidae" genus="Gustavus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="375" phylum="Arthropoda" rank="species" species="holthuisae" status="sp. nov.">
<emphasis id="B906EA807A05FFCD422FF8DB40CBF8F8" box="[280,486,1852,1876]" italics="true" pageId="6" pageNumber="375">Gustavus holthuisae</emphasis>
</taxonomicName>
<emphasis id="B906EA807A05FFCD42DCF8DB4317F8F8" bold="true" box="[491,570,1852,1876]" pageId="6" pageNumber="375">
<taxonomicNameLabel id="A23557FB7A05FFCD42DCF8DB4317F8F8" box="[491,570,1852,1876]" pageId="6" pageNumber="375" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, C, EG]; ovigerous female paratype (6.3 × 3.7 mm), FLMNH UF 72877, from type locality [B, D]; A, B, habitus in dorsal view; C, D, frontal margin and orbits, dorsal view; E, left G1, ventral view; F, left G1, dorsal view; G, left G2, ventral view.
</paragraph>
</caption>
<caption id="DF0D661A7A04FFCC43A0F9F742EFF970" ID-DOI="http://doi.org/10.5281/zenodo.13618529" ID-Zenodo-Dep="13618529" httpUri="https://zenodo.org/record/13618529/files/figure.png" pageId="7" pageNumber="376" startId="7.[151,250,1552,1576]" targetBox="[151,1435,181,1528]" targetPageId="7" targetType="figure">
<paragraph id="8BCD36927A04FFCC43A0F9F742EFF970" blockId="7.[151,1436,1552,1756]" pageId="7" pageNumber="376">
<emphasis id="B906EA807A04FFCC43A0F9F7403DF984" bold="true" box="[151,272,1552,1576]" pageId="7" pageNumber="376">FIGURE 5</emphasis>
.
<taxonomicName id="4C724D117A04FFCC4228F9F640DCF984" authorityName="Ng &amp; Anker" authorityYear="2024" box="[287,497,1552,1576]" class="Malacostraca" family="Aphanodactylidae" genus="Gustavus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="376" phylum="Arthropoda" rank="species" species="holthuisae" status="sp. nov.">
<emphasis id="B906EA807A04FFCC4228F9F640DCF984" box="[287,497,1552,1576]" italics="true" pageId="7" pageNumber="376">Gustavus holthuisae</emphasis>
</taxonomicName>
<emphasis id="B906EA807A04FFCC42CDF9F74360F984" bold="true" box="[506,589,1552,1576]" pageId="7" pageNumber="376">
<taxonomicNameLabel id="A23557FB7A04FFCC42CDF9F74360F984" box="[506,589,1552,1576]" pageId="7" pageNumber="376" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [AD]; ovigerous female paratype (6.3 × 3.7 mm) from type locality, FLMNH UF 72877 [E, F, H]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [G]; A, E, cephalothorax showing orbits, epistome, antennules and antennae, frontal view; B, right chela (partially denuded), outer view; C, male buccal cavity and anterior thoracic sternum, ventral view; D, male anterior thoracic sternum and sternopleonal cavity, showing intact right G1, ventral view; F, right chela (denuded), outer view; G, female pleon; H, female sternopleonal cavity and vulvae, ventral view.
</paragraph>
</caption>
<paragraph id="8BCD36927A04FFC343F0F8ED4394FE01" blockId="7.[151,1436,1802,2008]" lastBlockId="8.[151,1437,150,429]" lastPageId="8" lastPageNumber="377" pageId="7" pageNumber="376">
<emphasis id="B906EA807A04FFCC43F0F8ED4036F888" box="[199,283,1802,1828]" italics="true" pageId="7" pageNumber="376">Female</emphasis>
. Carapace transversely ovate, much broader than long, width to length ratio 1.741.83; dorsal surface almost smooth, with numerous small shallow pits, regions poorly demarcated (
<figureCitation id="13492A177A04FFCC471BF8C94599F8E5" box="[1068,1204,1838,1865]" captionStart="FIGURE 3" captionStartId="5.[151,250,1882,1906]" captionTargetBox="[151,1436,181,1858]" captionTargetId="figure-17@5.[151,1436,181,1858]" captionTargetPageId="5" captionText="FIGURE 3. Gustavus holthuisaesp. nov., ovigerous female paratype (7.3 × 4.0 mm) from Thuwal SaudiArabia, ZRC 2023.0580, habitus and colour in life; A, dorsal view; B, ventral view; C, frontal view. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.13618525" httpUri="https://zenodo.org/record/13618525/files/figure.png" pageId="7" pageNumber="376">Figs. 3AC</figureCitation>
,
<figureCitation id="13492A177A04FFCC47F4F8C845CAF8E5" box="[1219,1255,1839,1865]" captionStart="FIGURE 4" captionStartId="6.[151,250,1851,1875]" captionTargetBox="[151,1435,181,1828]" captionTargetId="figure-17@6.[151,1435,181,1828]" captionTargetPageId="6" captionText="FIGURE 4. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, C, EG]; ovigerous female paratype (6.3 × 3.7 mm), FLMNH UF 72877, from type locality [B, D]; A, B, habitus in dorsal view; C, D, frontal margin and orbits, dorsal view; E, left G1, ventral view; F, left G1, dorsal view; G, left G2, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618527" httpUri="https://zenodo.org/record/13618527/files/figure.png" pageId="7" pageNumber="376">4B</figureCitation>
); surface finely pubescent on surface and margins; front with short, dense setae partially obscuring margin; lower lateral margins with longer, dense, plumose setae obscuring surface; metagastric region transversely depressed, covered with low, soft setae, latter easily brushed off (
<figureCitation id="13492A177A04FFCC4106F87D4394F819" box="[561,697,1946,1973]" captionStart="FIGURE 3" captionStartId="5.[151,250,1882,1906]" captionTargetBox="[151,1436,181,1858]" captionTargetId="figure-17@5.[151,1436,181,1858]" captionTargetPageId="5" captionText="FIGURE 3. Gustavus holthuisaesp. nov., ovigerous female paratype (7.3 × 4.0 mm) from Thuwal SaudiArabia, ZRC 2023.0580, habitus and colour in life; A, dorsal view; B, ventral view; C, frontal view. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.13618525" httpUri="https://zenodo.org/record/13618525/files/figure.png" pageId="7" pageNumber="376">Figs. 3A, C</figureCitation>
,
<figureCitation id="13492A177A04FFCC41F0F87C43C6F819" box="[711,747,1947,1973]" captionStart="FIGURE 4" captionStartId="6.[151,250,1851,1875]" captionTargetBox="[151,1435,181,1828]" captionTargetId="figure-17@6.[151,1435,181,1828]" captionTargetPageId="6" captionText="FIGURE 4. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, C, EG]; ovigerous female paratype (6.3 × 3.7 mm), FLMNH UF 72877, from type locality [B, D]; A, B, habitus in dorsal view; C, D, frontal margin and orbits, dorsal view; E, left G1, ventral view; F, left G1, dorsal view; G, left G2, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618527" httpUri="https://zenodo.org/record/13618527/files/figure.png" pageId="7" pageNumber="376">4B</figureCitation>
). Front deflexed, medially gently emarginate in dorsal view, separated by shallow concavity, triangular in anterior view; lateral tooth low but distinct, rounded in dorsal view (
<figureCitation id="13492A177A0BFFC343A8FF704028FF1D" box="[159,261,151,177]" captionStart="FIGURE 4" captionStartId="6.[151,250,1851,1875]" captionTargetBox="[151,1435,181,1828]" captionTargetId="figure-17@6.[151,1435,181,1828]" captionTargetPageId="6" captionText="FIGURE 4. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, C, EG]; ovigerous female paratype (6.3 × 3.7 mm), FLMNH UF 72877, from type locality [B, D]; A, B, habitus in dorsal view; C, D, frontal margin and orbits, dorsal view; E, left G1, ventral view; F, left G1, dorsal view; G, left G2, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618527" httpUri="https://zenodo.org/record/13618527/files/figure.png" pageId="8" pageNumber="377">Figs. 4D</figureCitation>
,
<figureCitation id="13492A177A0BFFC34225FF714019FF1D" box="[274,308,150,177]" captionStart="FIGURE 5" captionStartId="7.[151,250,1552,1576]" captionTargetBox="[151,1435,181,1528]" captionTargetId="figure-17@7.[151,1435,181,1528]" captionTargetPageId="7" captionText="FIGURE 5. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [AD]; ovigerous female paratype (6.3 × 3.7 mm) from type locality, FLMNH UF 72877 [E, F, H]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [G]; A, E, cephalothorax showing orbits, epistome, antennules and antennae, frontal view; B, right chela (partially denuded), outer view; C, male buccal cavity and anterior thoracic sternum, ventral view; D, male anterior thoracic sternum and sternopleonal cavity, showing intact right G1, ventral view; F, right chela (denuded), outer view; G, female pleon; H, female sternopleonal cavity and vulvae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618529" httpUri="https://zenodo.org/record/13618529/files/figure.png" pageId="8" pageNumber="377">5E</figureCitation>
). Orbital margin entire, gently concave, slightly narrowed laterally; infraorbital margin terminating mesially as rounded angle; supraorbital margin terminating mesially as low, slender, ventrally directed tooth (
<figureCitation id="13492A177A0BFFC34653FF5C4191FF55" captionStart="FIGURE 4" captionStartId="6.[151,250,1851,1875]" captionTargetBox="[151,1435,181,1828]" captionTargetId="figure-17@6.[151,1435,181,1828]" captionTargetPageId="6" captionText="FIGURE 4. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, C, EG]; ovigerous female paratype (6.3 × 3.7 mm), FLMNH UF 72877, from type locality [B, D]; A, B, habitus in dorsal view; C, D, frontal margin and orbits, dorsal view; E, left G1, ventral view; F, left G1, dorsal view; G, left G2, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618527" httpUri="https://zenodo.org/record/13618527/files/figure.png" pageId="8" pageNumber="377">Figs. 4D</figureCitation>
,
<figureCitation id="13492A177A0BFFC343FDFF3941C1FF55" box="[202,236,222,249]" captionStart="FIGURE 5" captionStartId="7.[151,250,1552,1576]" captionTargetBox="[151,1435,181,1528]" captionTargetId="figure-17@7.[151,1435,181,1528]" captionTargetPageId="7" captionText="FIGURE 5. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [AD]; ovigerous female paratype (6.3 × 3.7 mm) from type locality, FLMNH UF 72877 [E, F, H]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [G]; A, E, cephalothorax showing orbits, epistome, antennules and antennae, frontal view; B, right chela (partially denuded), outer view; C, male buccal cavity and anterior thoracic sternum, ventral view; D, male anterior thoracic sternum and sternopleonal cavity, showing intact right G1, ventral view; F, right chela (denuded), outer view; G, female pleon; H, female sternopleonal cavity and vulvae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618529" httpUri="https://zenodo.org/record/13618529/files/figure.png" pageId="8" pageNumber="377">5E</figureCitation>
). Anterolateral margin entire, convex laterally, subcristate, continuing onto posterolateral margin but not supraorbital margin, with very shallow, short, transverse groove along proximal third (
<figureCitation id="13492A177A0BFFC347B6FEE445C5FEB1" box="[1153,1256,259,285]" captionStart="FIGURE 3" captionStartId="5.[151,250,1882,1906]" captionTargetBox="[151,1436,181,1858]" captionTargetId="figure-17@5.[151,1436,181,1858]" captionTargetPageId="5" captionText="FIGURE 3. Gustavus holthuisaesp. nov., ovigerous female paratype (7.3 × 4.0 mm) from Thuwal SaudiArabia, ZRC 2023.0580, habitus and colour in life; A, dorsal view; B, ventral view; C, frontal view. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.13618525" httpUri="https://zenodo.org/record/13618525/files/figure.png" pageId="8" pageNumber="377">Figs. 3A</figureCitation>
,
<figureCitation id="13492A177A0BFFC347C0FEE44436FEB1" box="[1271,1307,259,285]" captionStart="FIGURE 4" captionStartId="6.[151,250,1851,1875]" captionTargetBox="[151,1435,181,1828]" captionTargetId="figure-17@6.[151,1435,181,1828]" captionTargetPageId="6" captionText="FIGURE 4. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, C, EG]; ovigerous female paratype (6.3 × 3.7 mm), FLMNH UF 72877, from type locality [B, D]; A, B, habitus in dorsal view; C, D, frontal margin and orbits, dorsal view; E, left G1, ventral view; F, left G1, dorsal view; G, left G2, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618527" httpUri="https://zenodo.org/record/13618527/files/figure.png" pageId="8" pageNumber="377">4B</figureCitation>
). Epistome very short, partly sunken medially; posterior margin gently concave, entire (
<figureCitation id="13492A177A0BFFC34733FEC04570FEED" box="[1028,1117,294,321]" captionStart="FIGURE 5" captionStartId="7.[151,250,1552,1576]" captionTargetBox="[151,1435,181,1528]" captionTargetId="figure-17@7.[151,1435,181,1528]" captionTargetPageId="7" captionText="FIGURE 5. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [AD]; ovigerous female paratype (6.3 × 3.7 mm) from type locality, FLMNH UF 72877 [E, F, H]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [G]; A, E, cephalothorax showing orbits, epistome, antennules and antennae, frontal view; B, right chela (partially denuded), outer view; C, male buccal cavity and anterior thoracic sternum, ventral view; D, male anterior thoracic sternum and sternopleonal cavity, showing intact right G1, ventral view; F, right chela (denuded), outer view; G, female pleon; H, female sternopleonal cavity and vulvae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618529" httpUri="https://zenodo.org/record/13618529/files/figure.png" pageId="8" pageNumber="377">Fig. 5E</figureCitation>
). Eyes mobile, filling orbit; cornea simple, pigmented (
<figureCitation id="13492A177A0BFFC342FBFEAC431CFEC9" box="[460,561,331,357]" captionStart="FIGURE 4" captionStartId="6.[151,250,1851,1875]" captionTargetBox="[151,1435,181,1828]" captionTargetId="figure-17@6.[151,1435,181,1828]" captionTargetPageId="6" captionText="FIGURE 4. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, C, EG]; ovigerous female paratype (6.3 × 3.7 mm), FLMNH UF 72877, from type locality [B, D]; A, B, habitus in dorsal view; C, D, frontal margin and orbits, dorsal view; E, left G1, ventral view; F, left G1, dorsal view; G, left G2, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618527" httpUri="https://zenodo.org/record/13618527/files/figure.png" pageId="8" pageNumber="377">Figs. 4D</figureCitation>
,
<figureCitation id="13492A177A0BFFC34109FEAD434DFEC9" box="[574,608,330,357]" captionStart="FIGURE 5" captionStartId="7.[151,250,1552,1576]" captionTargetBox="[151,1435,181,1528]" captionTargetId="figure-17@7.[151,1435,181,1528]" captionTargetPageId="7" captionText="FIGURE 5. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [AD]; ovigerous female paratype (6.3 × 3.7 mm) from type locality, FLMNH UF 72877 [E, F, H]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [G]; A, E, cephalothorax showing orbits, epistome, antennules and antennae, frontal view; B, right chela (partially denuded), outer view; C, male buccal cavity and anterior thoracic sternum, ventral view; D, male anterior thoracic sternum and sternopleonal cavity, showing intact right G1, ventral view; F, right chela (denuded), outer view; G, female pleon; H, female sternopleonal cavity and vulvae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618529" httpUri="https://zenodo.org/record/13618529/files/figure.png" pageId="8" pageNumber="377">5E</figureCitation>
). Antennular fossae large; antennule folding obliquely, filling fossa (
<figureCitation id="13492A177A0BFFC34658FEAC4194FE25" captionStart="FIGURE 5" captionStartId="7.[151,250,1552,1576]" captionTargetBox="[151,1435,181,1528]" captionTargetId="figure-17@7.[151,1435,181,1528]" captionTargetPageId="7" captionText="FIGURE 5. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [AD]; ovigerous female paratype (6.3 × 3.7 mm) from type locality, FLMNH UF 72877 [E, F, H]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [G]; A, E, cephalothorax showing orbits, epistome, antennules and antennae, frontal view; B, right chela (partially denuded), outer view; C, male buccal cavity and anterior thoracic sternum, ventral view; D, male anterior thoracic sternum and sternopleonal cavity, showing intact right G1, ventral view; F, right chela (denuded), outer view; G, female pleon; H, female sternopleonal cavity and vulvae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618529" httpUri="https://zenodo.org/record/13618529/files/figure.png" pageId="8" pageNumber="377">Fig. 5E</figureCitation>
). Antenna short, completely entering orbit, with basal antennal article stout, quadrate, not fused to carapace, not reaching distolateral angle of carapace (
<figureCitation id="13492A177A0BFFC34162FE744381FE01" box="[597,684,402,429]" captionStart="FIGURE 5" captionStartId="7.[151,250,1552,1576]" captionTargetBox="[151,1435,181,1528]" captionTargetId="figure-17@7.[151,1435,181,1528]" captionTargetPageId="7" captionText="FIGURE 5. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [AD]; ovigerous female paratype (6.3 × 3.7 mm) from type locality, FLMNH UF 72877 [E, F, H]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [G]; A, E, cephalothorax showing orbits, epistome, antennules and antennae, frontal view; B, right chela (partially denuded), outer view; C, male buccal cavity and anterior thoracic sternum, ventral view; D, male anterior thoracic sternum and sternopleonal cavity, showing intact right G1, ventral view; F, right chela (denuded), outer view; G, female pleon; H, female sternopleonal cavity and vulvae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618529" httpUri="https://zenodo.org/record/13618529/files/figure.png" pageId="8" pageNumber="377">Fig. 5E</figureCitation>
).
</paragraph>
<caption id="DF0D661A7A0BFFC343A0F8FA437DF869" ID-DOI="http://doi.org/10.5281/zenodo.13618533" ID-Zenodo-Dep="13618533" httpUri="https://zenodo.org/record/13618533/files/figure.png" pageId="8" pageNumber="377" startId="8.[151,250,1821,1845]" targetBox="[158,1429,460,1791]" targetPageId="8" targetType="figure">
<paragraph id="8BCD36927A0BFFC343A0F8FA437DF869" blockId="8.[151,1437,1821,1989]" pageId="8" pageNumber="377">
<emphasis id="B906EA807A0BFFC343A0F8FA403DF899" bold="true" box="[151,272,1821,1845]" pageId="8" pageNumber="377">FIGURE 6</emphasis>
.
<taxonomicName id="4C724D117A0BFFC34228F8FA40DCF899" authorityName="Ng &amp; Anker" authorityYear="2024" box="[287,497,1821,1845]" class="Malacostraca" family="Aphanodactylidae" genus="Gustavus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="377" phylum="Arthropoda" rank="species" species="holthuisae" status="sp. nov.">
<emphasis id="B906EA807A0BFFC34228F8FA40DCF899" box="[287,497,1821,1845]" italics="true" pageId="8" pageNumber="377">Gustavus holthuisae</emphasis>
</taxonomicName>
<emphasis id="B906EA807A0BFFC342CDF8FA4360F899" bold="true" box="[506,589,1821,1845]" pageId="8" pageNumber="377">
<taxonomicNameLabel id="A23557FB7A0BFFC342CDF8FA4360F899" box="[506,589,1821,1845]" pageId="8" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, CJ]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [B, KN]; A, B, right Mxp3 (setae denuded), outer view; CF, male P2P5, respectively, dorsal view; G, male pleon; H, left G1, ventral view (setae omitted); I, left G1, dorsal view (setae omitted); J, left G2, ventral view (setae omitted); KN, female P2P5, respectively, dorsal view. Scales: A, B, HJ = 0.2 mm; CG, KN = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BCD36927A0AFFC243F0FF70435DFEED" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="378">
Mxp3 covering most of buccal cavern when closed; ischium distinctly longer than merus, slender, subtriangular, distomesial angle produced into rounded lobe extending mesially beyond level of mesial meral margin, mesial margin gently concave; merus smaller than ischium, slightly wider than long, squircular; palp articulating on distal margin of merus; basal part of exopod slender, tip reaching almost to just above merus; exopod flagellum short, not reaching distal margin of merus (
<figureCitation id="13492A177A0AFFC2413DFEC0434EFEED" box="[522,611,295,321]" captionStart="FIGURE 6" captionStartId="8.[151,250,1821,1845]" captionTargetBox="[158,1429,460,1791]" captionTargetId="figure-130@8.[151,1435,453,1798]" captionTargetPageId="8" captionText="FIGURE 6. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, CJ]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [B, KN]; A, B, right Mxp3 (setae denuded), outer view; CF, male P2P5, respectively, dorsal view; G, male pleon; H, left G1, ventral view (setae omitted); I, left G1, dorsal view (setae omitted); J, left G2, ventral view (setae omitted); KN, female P2P5, respectively, dorsal view. Scales: A, B, HJ = 0.2 mm; CG, KN = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618533" httpUri="https://zenodo.org/record/13618533/files/figure.png" pageId="9" pageNumber="378">Fig. 6B</figureCitation>
).
</paragraph>
<paragraph id="8BCD36927A0AFFC243F0FEAC4078FD91" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="378">
Chelipeds equal, densely covered with fine, short, setae partially obscuring margins and surface (
<figureCitation id="13492A177A0AFFC24605FEAC44BAFEC9" box="[1330,1431,331,357]" captionStart="FIGURE 4" captionStartId="6.[151,250,1851,1875]" captionTargetBox="[151,1435,181,1828]" captionTargetId="figure-17@6.[151,1435,181,1828]" captionTargetPageId="6" captionText="FIGURE 4. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, C, EG]; ovigerous female paratype (6.3 × 3.7 mm), FLMNH UF 72877, from type locality [B, D]; A, B, habitus in dorsal view; C, D, frontal margin and orbits, dorsal view; E, left G1, ventral view; F, left G1, dorsal view; G, left G2, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618527" httpUri="https://zenodo.org/record/13618527/files/figure.png" pageId="9" pageNumber="378">Figs. 4B</figureCitation>
,
<figureCitation id="13492A177A0AFFC243A0FE894195FE25" box="[151,184,366,393]" captionStart="FIGURE 5" captionStartId="7.[151,250,1552,1576]" captionTargetBox="[151,1435,181,1528]" captionTargetId="figure-17@7.[151,1435,181,1528]" captionTargetPageId="7" captionText="FIGURE 5. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [AD]; ovigerous female paratype (6.3 × 3.7 mm) from type locality, FLMNH UF 72877 [E, F, H]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [G]; A, E, cephalothorax showing orbits, epistome, antennules and antennae, frontal view; B, right chela (partially denuded), outer view; C, male buccal cavity and anterior thoracic sternum, ventral view; D, male anterior thoracic sternum and sternopleonal cavity, showing intact right G1, ventral view; F, right chela (denuded), outer view; G, female pleon; H, female sternopleonal cavity and vulvae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618529" httpUri="https://zenodo.org/record/13618529/files/figure.png" pageId="9" pageNumber="378">5F</figureCitation>
); merus triangular in cross-section, with plumose setae proximally; carpus smooth, inner angle rounded, with sparse plumose setae; chela somewhat elongate, surfaces smooth except for scattered, low, rounded granules on lower portion of palm; palm distinctly longer than dactylus, with ventral margin near pollex concave; pollex with occlusal margin convex, with blunt teeth proximally, distal two-thirds with small teeth fused basally, forming blade-like structure; dactylus with occlusal margin armed with 2 or 3 low, blunt teeth proximally, distal part unarmed, tip curved (
<figureCitation id="13492A177A0AFFC243C4FDC44065FD91" box="[243,328,546,573]" captionStart="FIGURE 5" captionStartId="7.[151,250,1552,1576]" captionTargetBox="[151,1435,181,1528]" captionTargetId="figure-17@7.[151,1435,181,1528]" captionTargetPageId="7" captionText="FIGURE 5. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [AD]; ovigerous female paratype (6.3 × 3.7 mm) from type locality, FLMNH UF 72877 [E, F, H]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [G]; A, E, cephalothorax showing orbits, epistome, antennules and antennae, frontal view; B, right chela (partially denuded), outer view; C, male buccal cavity and anterior thoracic sternum, ventral view; D, male anterior thoracic sternum and sternopleonal cavity, showing intact right G1, ventral view; F, right chela (denuded), outer view; G, female pleon; H, female sternopleonal cavity and vulvae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618529" httpUri="https://zenodo.org/record/13618529/files/figure.png" pageId="9" pageNumber="378">Fig. 5F</figureCitation>
).
</paragraph>
<paragraph id="8BCD36927A0AFFC243F0FDA141C1FC2D" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="378">
P2P5 relatively short, P3 longest, P5 shortest, dorsal to other pereopods (
<figureCitation id="13492A177A0AFFC240C3FDA0455AFDCD" box="[1012,1143,583,609]" captionStart="FIGURE 3" captionStartId="5.[151,250,1882,1906]" captionTargetBox="[151,1436,181,1858]" captionTargetId="figure-17@5.[151,1436,181,1858]" captionTargetPageId="5" captionText="FIGURE 3. Gustavus holthuisaesp. nov., ovigerous female paratype (7.3 × 4.0 mm) from Thuwal SaudiArabia, ZRC 2023.0580, habitus and colour in life; A, dorsal view; B, ventral view; C, frontal view. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.13618525" httpUri="https://zenodo.org/record/13618525/files/figure.png" pageId="9" pageNumber="378">Figs. 3AC</figureCitation>
,
<figureCitation id="13492A177A0AFFC247B6FDA04588FDCD" box="[1153,1189,583,609]" captionStart="FIGURE 4" captionStartId="6.[151,250,1851,1875]" captionTargetBox="[151,1435,181,1828]" captionTargetId="figure-17@6.[151,1435,181,1828]" captionTargetPageId="6" captionText="FIGURE 4. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, C, EG]; ovigerous female paratype (6.3 × 3.7 mm), FLMNH UF 72877, from type locality [B, D]; A, B, habitus in dorsal view; C, D, frontal margin and orbits, dorsal view; E, left G1, ventral view; F, left G1, dorsal view; G, left G2, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618527" httpUri="https://zenodo.org/record/13618527/files/figure.png" pageId="9" pageNumber="378">4B</figureCitation>
); propodus and carpus covered with short, fine tomentum partially obscuring surface and margins; merus with plumose setae along flexor and extensor margins; P2 and P3 merus longer than carpus and propodus combined; P4 and P5 merus subequal to carpus and propodus combined; dactylus short, stout, unarmed laterally, with corneous apex (
<figureCitation id="13492A177A0AFFC2478CFD544417FD61" box="[1211,1338,691,717]" captionStart="FIGURE 6" captionStartId="8.[151,250,1821,1845]" captionTargetBox="[158,1429,460,1791]" captionTargetId="figure-130@8.[151,1435,453,1798]" captionTargetPageId="8" captionText="FIGURE 6. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, CJ]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [B, KN]; A, B, right Mxp3 (setae denuded), outer view; CF, male P2P5, respectively, dorsal view; G, male pleon; H, left G1, ventral view (setae omitted); I, left G1, dorsal view (setae omitted); J, left G2, ventral view (setae omitted); KN, female P2P5, respectively, dorsal view. Scales: A, B, HJ = 0.2 mm; CG, KN = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618533" httpUri="https://zenodo.org/record/13618533/files/figure.png" pageId="9" pageNumber="378">Fig. 6KN</figureCitation>
). P2 and P3 merus with flexor margin almost smooth except for scattered, very low, sharp granules (
<figureCitation id="13492A177A0AFFC247ACFD30443BFD5D" box="[1179,1302,727,753]" captionStart="FIGURE 6" captionStartId="8.[151,250,1821,1845]" captionTargetBox="[158,1429,460,1791]" captionTargetId="figure-130@8.[151,1435,453,1798]" captionTargetPageId="8" captionText="FIGURE 6. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, CJ]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [B, KN]; A, B, right Mxp3 (setae denuded), outer view; CF, male P2P5, respectively, dorsal view; G, male pleon; H, left G1, ventral view (setae omitted); I, left G1, dorsal view (setae omitted); J, left G2, ventral view (setae omitted); KN, female P2P5, respectively, dorsal view. Scales: A, B, HJ = 0.2 mm; CG, KN = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618533" httpUri="https://zenodo.org/record/13618533/files/figure.png" pageId="9" pageNumber="378">Fig. 6K, L</figureCitation>
). P4 and P5 basis-ischium with 3 sharp, stout, fixed spines on flexor margin (
<figureCitation id="13492A177A0AFFC2404CFD1C42D2FCB9" box="[891,1023,763,789]" captionStart="FIGURE 6" captionStartId="8.[151,250,1821,1845]" captionTargetBox="[158,1429,460,1791]" captionTargetId="figure-130@8.[151,1435,453,1798]" captionTargetPageId="8" captionText="FIGURE 6. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, CJ]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [B, KN]; A, B, right Mxp3 (setae denuded), outer view; CF, male P2P5, respectively, dorsal view; G, male pleon; H, left G1, ventral view (setae omitted); I, left G1, dorsal view (setae omitted); J, left G2, ventral view (setae omitted); KN, female P2P5, respectively, dorsal view. Scales: A, B, HJ = 0.2 mm; CG, KN = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618533" httpUri="https://zenodo.org/record/13618533/files/figure.png" pageId="9" pageNumber="378">Fig. 6M, N</figureCitation>
); P4 merus with 1 large, fixed spine on proximal third of flexor margin (
<figureCitation id="13492A177A0AFFC2411AFCF843A1FC95" box="[557,652,799,825]" captionStart="FIGURE 6" captionStartId="8.[151,250,1821,1845]" captionTargetBox="[158,1429,460,1791]" captionTargetId="figure-130@8.[151,1435,453,1798]" captionTargetPageId="8" captionText="FIGURE 6. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, CJ]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [B, KN]; A, B, right Mxp3 (setae denuded), outer view; CF, male P2P5, respectively, dorsal view; G, male pleon; H, left G1, ventral view (setae omitted); I, left G1, dorsal view (setae omitted); J, left G2, ventral view (setae omitted); KN, female P2P5, respectively, dorsal view. Scales: A, B, HJ = 0.2 mm; CG, KN = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618533" httpUri="https://zenodo.org/record/13618533/files/figure.png" pageId="9" pageNumber="378">Fig. 6M</figureCitation>
); P5 with 2 large spines and 1 sharp, basal tubercle on proximal third of flexor margin (
<figureCitation id="13492A177A0AFFC24253FCA440EDFCF1" box="[356,448,835,861]" captionStart="FIGURE 6" captionStartId="8.[151,250,1821,1845]" captionTargetBox="[158,1429,460,1791]" captionTargetId="figure-130@8.[151,1435,453,1798]" captionTargetPageId="8" captionText="FIGURE 6. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, CJ]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [B, KN]; A, B, right Mxp3 (setae denuded), outer view; CF, male P2P5, respectively, dorsal view; G, male pleon; H, left G1, ventral view (setae omitted); I, left G1, dorsal view (setae omitted); J, left G2, ventral view (setae omitted); KN, female P2P5, respectively, dorsal view. Scales: A, B, HJ = 0.2 mm; CG, KN = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618533" httpUri="https://zenodo.org/record/13618533/files/figure.png" pageId="9" pageNumber="378">Fig. 6N</figureCitation>
). P2P5 propodus with 2 or 3 small, sharp, distoflexor spines opposing dactylus (
<figureCitation id="13492A177A0AFFC24658FCA441CDFC2D" captionStart="FIGURE 6" captionStartId="8.[151,250,1821,1845]" captionTargetBox="[158,1429,460,1791]" captionTargetId="figure-130@8.[151,1435,453,1798]" captionTargetPageId="8" captionText="FIGURE 6. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, CJ]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [B, KN]; A, B, right Mxp3 (setae denuded), outer view; CF, male P2P5, respectively, dorsal view; G, male pleon; H, left G1, ventral view (setae omitted); I, left G1, dorsal view (setae omitted); J, left G2, ventral view (setae omitted); KN, female P2P5, respectively, dorsal view. Scales: A, B, HJ = 0.2 mm; CG, KN = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618533" httpUri="https://zenodo.org/record/13618533/files/figure.png" pageId="9" pageNumber="378">Fig. 6KN</figureCitation>
).
</paragraph>
<paragraph id="8BCD36927A0AFFC243F0FC6C4442FC09" blockId="9.[151,1437,151,2013]" box="[199,1391,906,933]" pageId="9" pageNumber="378">
Thoracic sternum similar in form to that of male in non-sexual characters, but distinctly broader (
<figureCitation id="13492A177A0AFFC2463FFC6C444EFC09" box="[1288,1379,906,933]" captionStart="FIGURE 5" captionStartId="7.[151,250,1552,1576]" captionTargetBox="[151,1435,181,1528]" captionTargetId="figure-17@7.[151,1435,181,1528]" captionTargetPageId="7" captionText="FIGURE 5. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [AD]; ovigerous female paratype (6.3 × 3.7 mm) from type locality, FLMNH UF 72877 [E, F, H]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [G]; A, E, cephalothorax showing orbits, epistome, antennules and antennae, frontal view; B, right chela (partially denuded), outer view; C, male buccal cavity and anterior thoracic sternum, ventral view; D, male anterior thoracic sternum and sternopleonal cavity, showing intact right G1, ventral view; F, right chela (denuded), outer view; G, female pleon; H, female sternopleonal cavity and vulvae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618529" httpUri="https://zenodo.org/record/13618529/files/figure.png" pageId="9" pageNumber="378">Fig. 5H</figureCitation>
).
</paragraph>
<paragraph id="8BCD36927A0AFFC243F0FC4840ABFBBD" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="378">
Pleon wide, ovate, all somites and telson free; somites 3 and 4 widest; telson wide, with strongly convex margins (
<figureCitation id="13492A177A0AFFC243C8FC34404CFC41" box="[255,353,979,1005]" captionStart="FIGURE 3" captionStartId="5.[151,250,1882,1906]" captionTargetBox="[151,1436,181,1858]" captionTargetId="figure-17@5.[151,1436,181,1858]" captionTargetPageId="5" captionText="FIGURE 3. Gustavus holthuisaesp. nov., ovigerous female paratype (7.3 × 4.0 mm) from Thuwal SaudiArabia, ZRC 2023.0580, habitus and colour in life; A, dorsal view; B, ventral view; C, frontal view. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.13618525" httpUri="https://zenodo.org/record/13618525/files/figure.png" pageId="9" pageNumber="378">Figs. 3B</figureCitation>
,
<figureCitation id="13492A177A0AFFC2425CFC3540BFFC41" box="[363,402,978,1005]" captionStart="FIGURE 5" captionStartId="7.[151,250,1552,1576]" captionTargetBox="[151,1435,181,1528]" captionTargetId="figure-17@7.[151,1435,181,1528]" captionTargetPageId="7" captionText="FIGURE 5. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [AD]; ovigerous female paratype (6.3 × 3.7 mm) from type locality, FLMNH UF 72877 [E, F, H]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [G]; A, E, cephalothorax showing orbits, epistome, antennules and antennae, frontal view; B, right chela (partially denuded), outer view; C, male buccal cavity and anterior thoracic sternum, ventral view; D, male anterior thoracic sternum and sternopleonal cavity, showing intact right G1, ventral view; F, right chela (denuded), outer view; G, female pleon; H, female sternopleonal cavity and vulvae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618529" httpUri="https://zenodo.org/record/13618529/files/figure.png" pageId="9" pageNumber="378">5G</figureCitation>
). Vulvae positioned wide on sternite 6; vulvar cover tube-like, operculum soft, covering most of opening (
<figureCitation id="13492A177A0AFFC24228FC104054FBBD" box="[287,377,1014,1041]" captionStart="FIGURE 5" captionStartId="7.[151,250,1552,1576]" captionTargetBox="[151,1435,181,1528]" captionTargetId="figure-17@7.[151,1435,181,1528]" captionTargetPageId="7" captionText="FIGURE 5. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [AD]; ovigerous female paratype (6.3 × 3.7 mm) from type locality, FLMNH UF 72877 [E, F, H]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [G]; A, E, cephalothorax showing orbits, epistome, antennules and antennae, frontal view; B, right chela (partially denuded), outer view; C, male buccal cavity and anterior thoracic sternum, ventral view; D, male anterior thoracic sternum and sternopleonal cavity, showing intact right G1, ventral view; F, right chela (denuded), outer view; G, female pleon; H, female sternopleonal cavity and vulvae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618529" httpUri="https://zenodo.org/record/13618529/files/figure.png" pageId="9" pageNumber="378">Fig. 5H</figureCitation>
).
</paragraph>
<paragraph id="8BCD36927A0AFFC243F0FBFD438DFB69" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="378">
<emphasis id="B906EA807A0AFFC243F0FBFD4015FB98" bold="true" box="[199,312,1050,1076]" pageId="9" pageNumber="378">Variation</emphasis>
. There is a pronounced sexual dimorphism in
<taxonomicName id="4C724D117A0AFFC2405FFBFC457CFB99" authorityName="Ng &amp; Anker" authorityYear="2024" box="[872,1105,1051,1077]" class="Malacostraca" family="Aphanodactylidae" genus="Gustavus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="378" phylum="Arthropoda" rank="species" species="holthuisae" status="sp. nov.">
<emphasis id="B906EA807A0AFFC2405FFBFC457CFB99" box="[872,1105,1051,1077]" italics="true" pageId="9" pageNumber="378">Gustavus holthuisae</emphasis>
</taxonomicName>
<emphasis id="B906EA807A0AFFC2476AFBFC4596FB99" bold="true" box="[1117,1211,1051,1077]" pageId="9" pageNumber="378">
<taxonomicNameLabel id="A23557FB7A0AFFC2476AFBFC4596FB99" box="[1117,1211,1051,1077]" pageId="9" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, with the carapace distinctly different in shape and proportions (cf.
<figureCitation id="13492A177A0AFFC2418EFBD84231FBF5" box="[697,796,1087,1113]" captionStart="FIGURE 2" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[193,1394,181,1919]" captionTargetId="figure-17@4.[193,1394,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876, habitus and colour in life; A, dorsal view; B, ventral view; C, frontal view. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.13618523" httpUri="https://zenodo.org/record/13618523/files/figure.png" pageId="9" pageNumber="378">Figs. 2A</figureCitation>
,
<figureCitation id="13492A177A0AFFC2401FFBD84260FBF5" box="[808,845,1087,1113]" captionStart="FIGURE 3" captionStartId="5.[151,250,1882,1906]" captionTargetBox="[151,1436,181,1858]" captionTargetId="figure-17@5.[151,1436,181,1858]" captionTargetPageId="5" captionText="FIGURE 3. Gustavus holthuisaesp. nov., ovigerous female paratype (7.3 × 4.0 mm) from Thuwal SaudiArabia, ZRC 2023.0580, habitus and colour in life; A, dorsal view; B, ventral view; C, frontal view. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.13618525" httpUri="https://zenodo.org/record/13618525/files/figure.png" pageId="9" pageNumber="378">3A</figureCitation>
,
<figureCitation id="13492A177A0AFFC2406EFBD842B2FBF5" box="[857,927,1087,1113]" captionStart="FIGURE 4" captionStartId="6.[151,250,1851,1875]" captionTargetBox="[151,1435,181,1828]" captionTargetId="figure-17@6.[151,1435,181,1828]" captionTargetPageId="6" captionText="FIGURE 4. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, C, EG]; ovigerous female paratype (6.3 × 3.7 mm), FLMNH UF 72877, from type locality [B, D]; A, B, habitus in dorsal view; C, D, frontal margin and orbits, dorsal view; E, left G1, ventral view; F, left G1, dorsal view; G, left G2, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618527" httpUri="https://zenodo.org/record/13618527/files/figure.png" pageId="9" pageNumber="378">4A, B</figureCitation>
). The largest
<typeStatus id="54C988307A0AFFC2470EFBD845B4FBF5" box="[1081,1177,1087,1113]" pageId="9" pageNumber="378" type="paratype">paratype</typeStatus>
female (7.3 × 4.0 mm, ZRC 2023.0580) is noticeably proportionally wider than the other
<specimenCount id="9D74FD1B7A0AFFC240B9FB844537FBD1" box="[910,1050,1123,1149]" count="2" pageId="9" pageNumber="378" type="female">two females</specimenCount>
, its carapace width to length ratio being 1.83 versus 1.74 and 1.75. The female chelae are proportionally longer than those of the male, with the fingers more elongate and less curved (cf.
<figureCitation id="13492A177A0AFFC24129FB4C43BEFB69" box="[542,659,1194,1221]" captionStart="FIGURE 5" captionStartId="7.[151,250,1552,1576]" captionTargetBox="[151,1435,181,1528]" captionTargetId="figure-17@7.[151,1435,181,1528]" captionTargetPageId="7" captionText="FIGURE 5. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [AD]; ovigerous female paratype (6.3 × 3.7 mm) from type locality, FLMNH UF 72877 [E, F, H]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [G]; A, E, cephalothorax showing orbits, epistome, antennules and antennae, frontal view; B, right chela (partially denuded), outer view; C, male buccal cavity and anterior thoracic sternum, ventral view; D, male anterior thoracic sternum and sternopleonal cavity, showing intact right G1, ventral view; F, right chela (denuded), outer view; G, female pleon; H, female sternopleonal cavity and vulvae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618529" httpUri="https://zenodo.org/record/13618529/files/figure.png" pageId="9" pageNumber="378">Fig. 5B, F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36865197A0AFFC243F0FB2842F5FA9D" pageId="9" pageNumber="378" type="etymology">
<paragraph id="8BCD36927A0AFFC243F0FB2842F5FA9D" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="378">
<emphasis id="B906EA807A0AFFC243F0FB284065FB45" bold="true" box="[199,328,1231,1257]" pageId="9" pageNumber="378">Etymology</emphasis>
. The authors take a great pleasure in naming the species after our friend, the marine biologist Bernadette Holthuis (University of
<collectingRegion id="49B6F8707A0AFFC2411CFB14439DFAA1" box="[555,688,1267,1293]" country="United States of America" name="Washington" pageId="9" pageNumber="378">Washington</collectingRegion>
Friday Harbor Laboratories), who is also a relative of one of the greatest carcinologists of the 20
<superScript id="7C079BDA7A0AFFC242CCFAF14325FA88" attach="left" box="[507,520,1302,1316]" fontSize="6" pageId="9" pageNumber="378">th</superScript>
century, Lipke B. Holthuis (19212008).
</paragraph>
</subSubSection>
<subSubSection id="C36865197A0AFFC243F0FADD4555FA49" pageId="9" pageNumber="378" type="biology_ecology">
<paragraph id="8BCD36927A0AFFC243F0FADD4555FA49" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="378">
<emphasis id="B906EA807A0AFFC243F0FADD400CFAF8" bold="true" box="[199,289,1338,1364]" pageId="9" pageNumber="378">Biology</emphasis>
. All specimens of
<taxonomicName id="4C724D117A0AFFC242D8FADC43FEFAF9" authorityName="Ng &amp; Anker" authorityYear="2024" box="[495,723,1339,1365]" class="Malacostraca" family="Aphanodactylidae" genus="Gustavus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="378" phylum="Arthropoda" rank="species" species="holthuisae" status="sp. nov.">
<emphasis id="B906EA807A0AFFC242D8FADC43FEFAF9" box="[495,723,1339,1365]" italics="true" pageId="9" pageNumber="378">Gustavus holthuisae</emphasis>
</taxonomicName>
<emphasis id="B906EA807A0AFFC241EEFADC421FFAF9" bold="true" box="[729,818,1339,1365]" pageId="9" pageNumber="378">
<taxonomicNameLabel id="A23557FB7A0AFFC241EEFADC421FFAF9" box="[729,818,1339,1365]" pageId="9" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
were collected near mangrove roots (dominant species:
<taxonomicName id="4C724D117A0AFFC243A0FAB840F0FAD5" authority="(Forskal))" baseAuthorityName="Forskal" box="[151,477,1374,1401]" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" kingdom="Plantae" order="Lamiales" pageId="9" pageNumber="378" phylum="Tracheophyta" rank="species" species="marina">
<emphasis id="B906EA807A0AFFC243A0FAB8404CFAD4" box="[151,353,1375,1401]" italics="true" pageId="9" pageNumber="378">Avicennia marina</emphasis>
(Forskål))
</taxonomicName>
, in water less than
<quantity id="4C8A9B777A0AFFC241F5FAB94228FAD5" box="[706,773,1374,1401]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="378" unit="m" value="0.5">0.5 m</quantity>
deep; they were found in tubes of unidentified terebellid worms (
<figureCitation id="13492A177A0AFFC243C1FA644017FA31" box="[246,314,1411,1437]" captionStart="FIGURE 1" captionStartId="2.[151,250,1918,1942]" captionTargetBox="[151,1435,181,1894]" captionTargetId="figure-17@2.[151,1435,181,1894]" captionTargetPageId="2" captionText="FIGURE 1. Gustavus holthuisae sp. nov., ovigerous female paratype (6.3 × 3.7 mm) from Thuwal, Saudi Arabia, FLMNH UF 72877, crab with terebellid polychaete host shortly after collection; A, general view of worm with crab; B, close-up of crab clinging to worm host. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.13618521" httpUri="https://zenodo.org/record/13618521/files/figure.png" pageId="9" pageNumber="378">Fig. 1</figureCitation>
), which were extracted from fine sand/mud with the aid of a metallic suction pump. Although a complete specimen of the terebellid host was collected and deposited in KAUST, its identification to genus level was impossible based on examination of photographs alone (P. Hutchings, pers. comm.).
</paragraph>
</subSubSection>
<subSubSection id="C36865197A0AFFC143F0FA084013FEED" lastPageId="10" lastPageNumber="379" pageId="9" pageNumber="378" type="discussion">
<paragraph id="8BCD36927A0AFFC143F0FA084013FEED" blockId="9.[151,1437,151,2013]" lastBlockId="10.[151,1437,151,321]" lastPageId="10" lastPageNumber="379" pageId="9" pageNumber="378">
<emphasis id="B906EA807A0AFFC243F0FA084019F9A5" bold="true" box="[199,308,1519,1545]" pageId="9" pageNumber="378">Remarks</emphasis>
.
<taxonomicName id="4C724D117A0AFFC24276FA084309F9A5" authorityName="Ng &amp; Anker" authorityYear="2024" box="[321,548,1519,1545]" class="Malacostraca" family="Aphanodactylidae" genus="Gustavus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="378" phylum="Arthropoda" rank="species" species="holthuisae" status="sp. nov.">
<emphasis id="B906EA807A0AFFC24276FA084309F9A5" box="[321,548,1519,1545]" italics="true" pageId="9" pageNumber="378">Gustavus holthuisae</emphasis>
</taxonomicName>
<emphasis id="B906EA807A0AFFC2411EFA0843ACF9A5" bold="true" box="[553,641,1519,1545]" pageId="9" pageNumber="378">
<taxonomicNameLabel id="A23557FB7A0AFFC2411EFA0843ACF9A5" box="[553,641,1519,1545]" pageId="9" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can easily be separated from
<taxonomicName id="4C724D117A0AFFC240F1FA084557F9A5" authorityName="Ahyong &amp; Ng" authorityYear="2009" box="[966,1146,1519,1545]" class="Malacostraca" family="Aphanodactylidae" genus="Gustavus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="378" phylum="Arthropoda" rank="species" species="mecognathus">
<emphasis id="B906EA807A0AFFC240F1FA084557F9A5" box="[966,1146,1519,1545]" italics="true" pageId="9" pageNumber="378">G. mecognathus</emphasis>
</taxonomicName>
by the following features: the metagastric region of the carapace is transversely distinctly depressed in both sexes (
<figureCitation id="13492A177A0AFFC24746F9F445D8F981" box="[1137,1269,1555,1581]" captionStart="FIGURE 2" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[193,1394,181,1919]" captionTargetId="figure-17@4.[193,1394,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876, habitus and colour in life; A, dorsal view; B, ventral view; C, frontal view. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.13618523" httpUri="https://zenodo.org/record/13618523/files/figure.png" pageId="9" pageNumber="378">Figs. 2A, C</figureCitation>
,
<figureCitation id="13492A177A0AFFC24637F9F44468F981" box="[1280,1349,1555,1581]" captionStart="FIGURE 3" captionStartId="5.[151,250,1882,1906]" captionTargetBox="[151,1436,181,1858]" captionTargetId="figure-17@5.[151,1436,181,1858]" captionTargetPageId="5" captionText="FIGURE 3. Gustavus holthuisaesp. nov., ovigerous female paratype (7.3 × 4.0 mm) from Thuwal SaudiArabia, ZRC 2023.0580, habitus and colour in life; A, dorsal view; B, ventral view; C, frontal view. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.13618525" httpUri="https://zenodo.org/record/13618525/files/figure.png" pageId="9" pageNumber="378">3A, C</figureCitation>
,
<figureCitation id="13492A177A0AFFC24667F9F444BEF981" box="[1360,1427,1555,1581]" captionStart="FIGURE 4" captionStartId="6.[151,250,1851,1875]" captionTargetBox="[151,1435,181,1828]" captionTargetId="figure-17@6.[151,1435,181,1828]" captionTargetPageId="6" captionText="FIGURE 4. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, C, EG]; ovigerous female paratype (6.3 × 3.7 mm), FLMNH UF 72877, from type locality [B, D]; A, B, habitus in dorsal view; C, D, frontal margin and orbits, dorsal view; E, left G1, ventral view; F, left G1, dorsal view; G, left G2, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618527" httpUri="https://zenodo.org/record/13618527/files/figure.png" pageId="9" pageNumber="378">4A, B</figureCitation>
) (versus the entire dorsal carapace surface being gently convex, including the metagastric region, in
<taxonomicName id="4C724D117A0AFFC247D7F9D044B8F9FD" authorityName="Ahyong &amp; Ng" authorityYear="2009" box="[1248,1429,1591,1617]" class="Malacostraca" family="Aphanodactylidae" genus="Gustavus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="378" phylum="Arthropoda" rank="species" species="mecognathus">
<emphasis id="B906EA807A0AFFC247D7F9D044B8F9FD" box="[1248,1429,1591,1617]" italics="true" pageId="9" pageNumber="378">G. mecognathus</emphasis>
</taxonomicName>
; cf. Ng &amp;
<bibRefCitation id="EFE34B637A0AFFC243CBF9BC40B5F9D9" author="Ahyong, S. T. &amp; Ng, P. K. L." box="[252,408,1627,1653]" pageId="9" pageNumber="378" pagination="33 - 47" refId="ref5317" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2009) Aphanodactylidae, a new family of thoracotreme crabs (Crustacea: Brachyura) symbiotic with polychaete worms. Zootaxa, 2289 (1), 33 - 47. https: // doi. org / 10.11646 / zootaxa. 2289.1.3" type="journal article" year="2009">Ahyong 2009</bibRefCitation>
: figs. 4A, 5A, 7A); the male frontal margin is proportionally narrower, with the median lobes acutely triangular (
<figureCitation id="13492A177A0AFFC2425BF99840EEF935" box="[364,451,1662,1689]" captionStart="FIGURE 5" captionStartId="7.[151,250,1552,1576]" captionTargetBox="[151,1435,181,1528]" captionTargetId="figure-17@7.[151,1435,181,1528]" captionTargetPageId="7" captionText="FIGURE 5. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [AD]; ovigerous female paratype (6.3 × 3.7 mm) from type locality, FLMNH UF 72877 [E, F, H]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [G]; A, E, cephalothorax showing orbits, epistome, antennules and antennae, frontal view; B, right chela (partially denuded), outer view; C, male buccal cavity and anterior thoracic sternum, ventral view; D, male anterior thoracic sternum and sternopleonal cavity, showing intact right G1, ventral view; F, right chela (denuded), outer view; G, female pleon; H, female sternopleonal cavity and vulvae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618529" httpUri="https://zenodo.org/record/13618529/files/figure.png" pageId="9" pageNumber="378">Fig. 5C</figureCitation>
) (versus broader and with the median lobes lower and more obtuse in
<taxonomicName id="4C724D117A0AFFC247E9F99844B8F935" authorityName="Ahyong &amp; Ng" authorityYear="2009" box="[1246,1429,1663,1689]" class="Malacostraca" family="Aphanodactylidae" genus="Gustavus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="378" phylum="Arthropoda" rank="species" species="mecognathus">
<emphasis id="B906EA807A0AFFC247E9F99844B8F935" box="[1246,1429,1663,1689]" italics="true" pageId="9" pageNumber="378">G. mecognathus</emphasis>
</taxonomicName>
; Ng &amp;
<bibRefCitation id="EFE34B637A0AFFC243EDF944405BF911" author="Ahyong, S. T. &amp; Ng, P. K. L." box="[218,374,1699,1725]" pageId="9" pageNumber="378" pagination="33 - 47" refId="ref5317" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2009) Aphanodactylidae, a new family of thoracotreme crabs (Crustacea: Brachyura) symbiotic with polychaete worms. Zootaxa, 2289 (1), 33 - 47. https: // doi. org / 10.11646 / zootaxa. 2289.1.3" type="journal article" year="2009">Ahyong 2009</bibRefCitation>
: fig. 4A); the female frontal margin has the median lobes separated by a shallow concavity (
<figureCitation id="13492A177A0AFFC24658F9444197F94D" captionStart="FIGURE 5" captionStartId="7.[151,250,1552,1576]" captionTargetBox="[151,1435,181,1528]" captionTargetId="figure-17@7.[151,1435,181,1528]" captionTargetPageId="7" captionText="FIGURE 5. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [AD]; ovigerous female paratype (6.3 × 3.7 mm) from type locality, FLMNH UF 72877 [E, F, H]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [G]; A, E, cephalothorax showing orbits, epistome, antennules and antennae, frontal view; B, right chela (partially denuded), outer view; C, male buccal cavity and anterior thoracic sternum, ventral view; D, male anterior thoracic sternum and sternopleonal cavity, showing intact right G1, ventral view; F, right chela (denuded), outer view; G, female pleon; H, female sternopleonal cavity and vulvae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618529" httpUri="https://zenodo.org/record/13618529/files/figure.png" pageId="9" pageNumber="378">Fig. 5D</figureCitation>
) (versus separated by a distinct V-shaped notch in
<taxonomicName id="4C724D117A0AFFC241C3F9204287F94D" authorityName="Ahyong &amp; Ng" authorityYear="2009" box="[756,938,1735,1761]" class="Malacostraca" family="Aphanodactylidae" genus="Gustavus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="378" phylum="Arthropoda" rank="species" species="mecognathus">
<emphasis id="B906EA807A0AFFC241C3F9204287F94D" box="[756,938,1735,1761]" italics="true" pageId="9" pageNumber="378">G. mecognathus</emphasis>
</taxonomicName>
; cf. Ng &amp;
<bibRefCitation id="EFE34B637A0AFFC24716F9204593F94D" author="Ahyong, S. T. &amp; Ng, P. K. L." box="[1057,1214,1735,1761]" pageId="9" pageNumber="378" pagination="33 - 47" refId="ref5317" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2009) Aphanodactylidae, a new family of thoracotreme crabs (Crustacea: Brachyura) symbiotic with polychaete worms. Zootaxa, 2289 (1), 33 - 47. https: // doi. org / 10.11646 / zootaxa. 2289.1.3" type="journal article" year="2009">Ahyong 2009</bibRefCitation>
: fig. 5A); the Mxp3 ischium has a gently concave inner margin (
<figureCitation id="13492A177A0AFFC2414BF90C43DCF8A9" box="[636,753,1771,1797]" captionStart="FIGURE 6" captionStartId="8.[151,250,1821,1845]" captionTargetBox="[158,1429,460,1791]" captionTargetId="figure-130@8.[151,1435,453,1798]" captionTargetPageId="8" captionText="FIGURE 6. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, CJ]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [B, KN]; A, B, right Mxp3 (setae denuded), outer view; CF, male P2P5, respectively, dorsal view; G, male pleon; H, left G1, ventral view (setae omitted); I, left G1, dorsal view (setae omitted); J, left G2, ventral view (setae omitted); KN, female P2P5, respectively, dorsal view. Scales: A, B, HJ = 0.2 mm; CG, KN = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618533" httpUri="https://zenodo.org/record/13618533/files/figure.png" pageId="9" pageNumber="378">Fig. 6A, B</figureCitation>
) (versus with a distinctly concave margin in
<taxonomicName id="4C724D117A0AFFC247D7F90C44B8F8A9" authorityName="Ahyong &amp; Ng" authorityYear="2009" box="[1248,1429,1771,1797]" class="Malacostraca" family="Aphanodactylidae" genus="Gustavus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="378" phylum="Arthropoda" rank="species" species="mecognathus">
<emphasis id="B906EA807A0AFFC247D7F90C44B8F8A9" box="[1248,1429,1771,1797]" italics="true" pageId="9" pageNumber="378">G. mecognathus</emphasis>
</taxonomicName>
; cf. Ng &amp;
<bibRefCitation id="EFE34B637A0AFFC24236F8E840B2F885" author="Ahyong, S. T. &amp; Ng, P. K. L." box="[257,415,1807,1833]" pageId="9" pageNumber="378" pagination="33 - 47" refId="ref5317" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2009) Aphanodactylidae, a new family of thoracotreme crabs (Crustacea: Brachyura) symbiotic with polychaete worms. Zootaxa, 2289 (1), 33 - 47. https: // doi. org / 10.11646 / zootaxa. 2289.1.3" type="journal article" year="2009">Ahyong 2009</bibRefCitation>
: figs. 4C, 5C); the basis-ischium of the male and female P4 and P5 has two or three spines on the flexor margin (
<figureCitation id="13492A177A0AFFC242BBF8D4436BF8E1" box="[396,582,1843,1869]" captionStart="FIGURE 6" captionStartId="8.[151,250,1821,1845]" captionTargetBox="[158,1429,460,1791]" captionTargetId="figure-130@8.[151,1435,453,1798]" captionTargetPageId="8" captionText="FIGURE 6. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, CJ]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [B, KN]; A, B, right Mxp3 (setae denuded), outer view; CF, male P2P5, respectively, dorsal view; G, male pleon; H, left G1, ventral view (setae omitted); I, left G1, dorsal view (setae omitted); J, left G2, ventral view (setae omitted); KN, female P2P5, respectively, dorsal view. Scales: A, B, HJ = 0.2 mm; CG, KN = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618533" httpUri="https://zenodo.org/record/13618533/files/figure.png" pageId="9" pageNumber="378">Fig. 6E, F, M, N</figureCitation>
) (versus a cluster of three or four small and large spines in
<taxonomicName id="4C724D117A0AFFC247E9F8D444B8F8E1" authorityName="Ahyong &amp; Ng" authorityYear="2009" box="[1246,1429,1843,1869]" class="Malacostraca" family="Aphanodactylidae" genus="Gustavus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="378" phylum="Arthropoda" rank="species" species="mecognathus">
<emphasis id="B906EA807A0AFFC247E9F8D444B8F8E1" box="[1246,1429,1843,1869]" italics="true" pageId="9" pageNumber="378">G. mecognathus</emphasis>
</taxonomicName>
; cf. Ng &amp;
<bibRefCitation id="EFE34B637A0AFFC24231F8B0408BF8DD" author="Ahyong, S. T. &amp; Ng, P. K. L." box="[262,422,1879,1905]" pageId="9" pageNumber="378" pagination="33 - 47" refId="ref5317" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2009) Aphanodactylidae, a new family of thoracotreme crabs (Crustacea: Brachyura) symbiotic with polychaete worms. Zootaxa, 2289 (1), 33 - 47. https: // doi. org / 10.11646 / zootaxa. 2289.1.3" type="journal article" year="2009">Ahyong 2009</bibRefCitation>
: figs. 4G, H, 5G, H); the merus of the male and female P5 has two spines on the flexor margin (
<figureCitation id="13492A177A0AFFC243C2F89C4044F839" box="[245,361,1915,1941]" captionStart="FIGURE 6" captionStartId="8.[151,250,1821,1845]" captionTargetBox="[158,1429,460,1791]" captionTargetId="figure-130@8.[151,1435,453,1798]" captionTargetPageId="8" captionText="FIGURE 6. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, CJ]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [B, KN]; A, B, right Mxp3 (setae denuded), outer view; CF, male P2P5, respectively, dorsal view; G, male pleon; H, left G1, ventral view (setae omitted); I, left G1, dorsal view (setae omitted); J, left G2, ventral view (setae omitted); KN, female P2P5, respectively, dorsal view. Scales: A, B, HJ = 0.2 mm; CG, KN = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618533" httpUri="https://zenodo.org/record/13618533/files/figure.png" pageId="9" pageNumber="378">Fig. 6F, N</figureCitation>
) (versus only one spine in
<taxonomicName id="4C724D117A0AFFC241A0F89C4263F839" authorityName="Ahyong &amp; Ng" authorityYear="2009" box="[663,846,1915,1941]" class="Malacostraca" family="Aphanodactylidae" genus="Gustavus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="378" phylum="Arthropoda" rank="species" species="mecognathus">
<emphasis id="B906EA807A0AFFC241A0F89C4263F839" box="[663,846,1915,1941]" italics="true" pageId="9" pageNumber="378">G. mecognathus</emphasis>
</taxonomicName>
; cf. Ng &amp;
<bibRefCitation id="EFE34B637A0AFFC240F2F89C454FF839" author="Ahyong, S. T. &amp; Ng, P. K. L." box="[965,1122,1915,1941]" pageId="9" pageNumber="378" pagination="33 - 47" refId="ref5317" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2009) Aphanodactylidae, a new family of thoracotreme crabs (Crustacea: Brachyura) symbiotic with polychaete worms. Zootaxa, 2289 (1), 33 - 47. https: // doi. org / 10.11646 / zootaxa. 2289.1.3" type="journal article" year="2009">Ahyong 2009</bibRefCitation>
: figs. 4H, 5H); the merus of the female P4 has only one spine on the flexor margin (
<figureCitation id="13492A177A0AFFC2403BF8784247F815" box="[780,874,1951,1977]" captionStart="FIGURE 6" captionStartId="8.[151,250,1821,1845]" captionTargetBox="[158,1429,460,1791]" captionTargetId="figure-130@8.[151,1435,453,1798]" captionTargetPageId="8" captionText="FIGURE 6. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, CJ]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [B, KN]; A, B, right Mxp3 (setae denuded), outer view; CF, male P2P5, respectively, dorsal view; G, male pleon; H, left G1, ventral view (setae omitted); I, left G1, dorsal view (setae omitted); J, left G2, ventral view (setae omitted); KN, female P2P5, respectively, dorsal view. Scales: A, B, HJ = 0.2 mm; CG, KN = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618533" httpUri="https://zenodo.org/record/13618533/files/figure.png" pageId="9" pageNumber="378">Fig. 6M</figureCitation>
) (versus five spines in
<taxonomicName id="4C724D117A0AFFC24746F8784405F815" authorityName="Ahyong &amp; Ng" authorityYear="2009" box="[1137,1320,1951,1977]" class="Malacostraca" family="Aphanodactylidae" genus="Gustavus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="378" phylum="Arthropoda" rank="species" species="mecognathus">
<emphasis id="B906EA807A0AFFC24746F8784405F815" box="[1137,1320,1951,1977]" italics="true" pageId="9" pageNumber="378">G. mecognathus</emphasis>
</taxonomicName>
; cf. Ng &amp;
<bibRefCitation id="EFE34B637A0AFFC243A0F8244019F871" author="Ahyong, S. T. &amp; Ng, P. K. L." box="[151,308,1987,2013]" pageId="9" pageNumber="378" pagination="33 - 47" refId="ref5317" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2009) Aphanodactylidae, a new family of thoracotreme crabs (Crustacea: Brachyura) symbiotic with polychaete worms. Zootaxa, 2289 (1), 33 - 47. https: // doi. org / 10.11646 / zootaxa. 2289.1.3" type="journal article" year="2009">Ahyong 2009</bibRefCitation>
: fig. 5G); the male pleon is proportionally more elongate, with all sutures distinct and clearly visible, despite the immobility of the somites 46 (
<figureCitation id="13492A177A09FFC141B1FF7043CCFF1D" box="[646,737,151,177]" captionStart="FIGURE 6" captionStartId="8.[151,250,1821,1845]" captionTargetBox="[158,1429,460,1791]" captionTargetId="figure-130@8.[151,1435,453,1798]" captionTargetPageId="8" captionText="FIGURE 6. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, CJ]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [B, KN]; A, B, right Mxp3 (setae denuded), outer view; CF, male P2P5, respectively, dorsal view; G, male pleon; H, left G1, ventral view (setae omitted); I, left G1, dorsal view (setae omitted); J, left G2, ventral view (setae omitted); KN, female P2P5, respectively, dorsal view. Scales: A, B, HJ = 0.2 mm; CG, KN = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618533" httpUri="https://zenodo.org/record/13618533/files/figure.png" pageId="10" pageNumber="379">Fig. 6G</figureCitation>
) (versus with lateral sutures between somites 46 shallow to undiscernible in
<taxonomicName id="4C724D117A09FFC14263FF5C4321FF79" authorityName="Ahyong &amp; Ng" authorityYear="2009" box="[340,524,187,213]" class="Malacostraca" family="Aphanodactylidae" genus="Gustavus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="379" phylum="Arthropoda" rank="species" species="mecognathus">
<emphasis id="B906EA807A09FFC14263FF5C4321FF79" box="[340,524,187,213]" italics="true" pageId="10" pageNumber="379">G. mecognathus</emphasis>
</taxonomicName>
; cf. Ng &amp;
<bibRefCitation id="EFE34B637A09FFC141B0FF5C420BFF79" author="Ahyong, S. T. &amp; Ng, P. K. L." box="[647,806,187,213]" pageId="10" pageNumber="379" pagination="33 - 47" refId="ref5317" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2009) Aphanodactylidae, a new family of thoracotreme crabs (Crustacea: Brachyura) symbiotic with polychaete worms. Zootaxa, 2289 (1), 33 - 47. https: // doi. org / 10.11646 / zootaxa. 2289.1.3" type="journal article" year="2009">Ahyong 2009</bibRefCitation>
: fig. 4J); and, finally, the G1 is overall stouter, with the distal part short and rounded, and with a subdistal projection (
<figureCitation id="13492A177A09FFC14008FF384295FF55" box="[831,952,222,249]" captionStart="FIGURE 5" captionStartId="7.[151,250,1552,1576]" captionTargetBox="[151,1435,181,1528]" captionTargetId="figure-17@7.[151,1435,181,1528]" captionTargetPageId="7" captionText="FIGURE 5. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [AD]; ovigerous female paratype (6.3 × 3.7 mm) from type locality, FLMNH UF 72877 [E, F, H]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [G]; A, E, cephalothorax showing orbits, epistome, antennules and antennae, frontal view; B, right chela (partially denuded), outer view; C, male buccal cavity and anterior thoracic sternum, ventral view; D, male anterior thoracic sternum and sternopleonal cavity, showing intact right G1, ventral view; F, right chela (denuded), outer view; G, female pleon; H, female sternopleonal cavity and vulvae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13618529" httpUri="https://zenodo.org/record/13618529/files/figure.png" pageId="10" pageNumber="379">Figs. 5E, F</figureCitation>
,
<figureCitation id="13492A177A09FFC140F5FF3842D6FF55" box="[962,1019,223,249]" captionStart="FIGURE 6" captionStartId="8.[151,250,1821,1845]" captionTargetBox="[158,1429,460,1791]" captionTargetId="figure-130@8.[151,1435,453,1798]" captionTargetPageId="8" captionText="FIGURE 6. Gustavus holthuisae sp. nov., male holotype (5.3 × 3.4 mm) from Thuwal, Saudi Arabia, FLMNH UF 72876 [A, CJ]; ovigerous female paratype (7.3 × 4.0 mm) from type locality, ZRC 2023.0580 [B, KN]; A, B, right Mxp3 (setae denuded), outer view; CF, male P2P5, respectively, dorsal view; G, male pleon; H, left G1, ventral view (setae omitted); I, left G1, dorsal view (setae omitted); J, left G2, ventral view (setae omitted); KN, female P2P5, respectively, dorsal view. Scales: A, B, HJ = 0.2 mm; CG, KN = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618533" httpUri="https://zenodo.org/record/13618533/files/figure.png" pageId="10" pageNumber="379">6H, I</figureCitation>
) (versus the G1 being relatively more slender, with the distal part more elongate and without a subdistal projection in
<taxonomicName id="4C724D117A09FFC14723FEE445E6FEB1" authorityName="Ahyong &amp; Ng" authorityYear="2009" box="[1044,1227,259,285]" class="Malacostraca" family="Aphanodactylidae" genus="Gustavus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="379" phylum="Arthropoda" rank="species" species="mecognathus">
<emphasis id="B906EA807A09FFC14723FEE445E6FEB1" box="[1044,1227,259,285]" italics="true" pageId="10" pageNumber="379">G. mecognathus</emphasis>
</taxonomicName>
; cf. Ng &amp;
<bibRefCitation id="EFE34B637A09FFC14675FEE441FEFEED" author="Ahyong, S. T. &amp; Ng, P. K. L." pageId="10" pageNumber="379" pagination="33 - 47" refId="ref5317" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2009) Aphanodactylidae, a new family of thoracotreme crabs (Crustacea: Brachyura) symbiotic with polychaete worms. Zootaxa, 2289 (1), 33 - 47. https: // doi. org / 10.11646 / zootaxa. 2289.1.3" type="journal article" year="2009">Ahyong 2009</bibRefCitation>
: fig. 4K).
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<taxonomicName id="4C724D117A02FFCA43A0FE3C4329FE58" ID-CoL="7PHR8" authority="Ahyong &amp; Ng, 2009" authorityName="Ahyong &amp; Ng" authorityYear="2009" box="[151,516,474,501]" class="Malacostraca" family="Aphanodactylidae" genus="Gustavus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B906EA807A02FFCA43A0FE3C4329FE58" bold="true" box="[151,516,474,501]" pageId="1" pageNumber="370">
<emphasis id="B906EA807A02FFCA43A0FE3C402AFE59" bold="true" box="[151,263,475,501]" italics="true" pageId="1" pageNumber="370">Gustavus</emphasis>
<bibRefCitation id="EFE34B637A02FFCA423AFE3D4329FE58" author="Ahyong, S. T. &amp; Ng, P. K. L." box="[269,516,474,501]" pageId="1" pageNumber="370" pagination="33 - 47" refId="ref5317" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2009) Aphanodactylidae, a new family of thoracotreme crabs (Crustacea: Brachyura) symbiotic with polychaete worms. Zootaxa, 2289 (1), 33 - 47. https: // doi. org / 10.11646 / zootaxa. 2289.1.3" type="journal article" year="2009">Ahyong &amp; Ng, 2009</bibRefCitation>
</emphasis>
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<emphasis id="B906EA807A02FFCA43A0FDF84033FD9B" bold="true" box="[151,286,543,567]" pageId="1" pageNumber="370">
<typeStatus id="54C988307A02FFCA43A0FDF841E1FD9B" box="[151,204,543,567]" pageId="1" pageNumber="370">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="4C724D117A02FFCA421DFDC743C0FD9B" ID-CoL="3HMWX" authority="Ahyong &amp; Ng, 2009" authorityName="Ahyong &amp; Ng" authorityYear="2009" box="[298,749,543,568]" class="Malacostraca" family="Aphanodactylidae" genus="Gustavus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="species" species="mecognathus">
<emphasis id="B906EA807A02FFCA421DFDC74339FD9B" box="[298,532,543,567]" italics="true" pageId="1" pageNumber="370">Gustavus mecognathus</emphasis>
<bibRefCitation id="EFE34B637A02FFCA412EFDF843C0FD9B" author="Ahyong, S. T. &amp; Ng, P. K. L." box="[537,749,543,568]" pageId="1" pageNumber="370" pagination="33 - 47" refId="ref5317" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2009) Aphanodactylidae, a new family of thoracotreme crabs (Crustacea: Brachyura) symbiotic with polychaete worms. Zootaxa, 2289 (1), 33 - 47. https: // doi. org / 10.11646 / zootaxa. 2289.1.3" type="journal article" year="2009">Ahyong &amp; Ng, 2009</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BCD36927A02FFCA43A0FD8243D5FDD3" blockId="1.[151,1437,613,928]" box="[151,760,613,640]" pageId="1" pageNumber="370">
<emphasis id="B906EA807A02FFCA43A0FD82408BFDD3" bold="true" box="[151,422,613,639]" pageId="1" pageNumber="370">Other species included</emphasis>
.
<taxonomicName id="4C724D117A02FFCA4284FD8143B5FDD3" authorityName="Ng &amp; Anker" authorityYear="2024" box="[435,664,613,640]" class="Malacostraca" family="Aphanodactylidae" genus="Gustavus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="species" species="holthuisae" status="sp. nov.">
<emphasis id="B906EA807A02FFCA4284FD8143B5FDD3" box="[435,664,613,640]" italics="true" pageId="1" pageNumber="370">Gustavus holthuisae</emphasis>
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<taxonomicNameLabel id="A23557FB7A02FFCA41A9FD8243D5FDD3" box="[670,760,613,639]" pageId="1" pageNumber="370" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C36865197A02FFCA43F0FD6E45CBFC33" pageId="1" pageNumber="370" type="discussion">
<paragraph id="8BCD36927A02FFCA43F0FD6E45CBFC33" blockId="1.[151,1437,613,928]" pageId="1" pageNumber="370">
<emphasis id="B906EA807A02FFCA43F0FD6E4016FD0F" bold="true" box="[199,315,649,675]" pageId="1" pageNumber="370">Remarks.</emphasis>
<bibRefCitation id="EFE34B637A02FFCA4271FD6C40C3FD0F" author="Stevcic, Z." box="[326,494,649,678]" pageId="1" pageNumber="370" pagination="125 - 139" refId="ref6619" refString="Stevcic, Z. (2011) Addition to the reclassification of brachyuran crabs (Crustacea: Decapoda: Brachyura). Part I. New taxa. Natura Croatica, 20, 125 - 139." type="journal article" year="2011">Števčić (2011)</bibRefCitation>
recognised a new tribe,
<taxonomicName id="4C724D117A02FFCA4022FD6E42A8FD0F" authorityName="Števčić" authorityYear="2011" box="[789,901,649,675]" class="Malacostraca" family="Aphanodactylidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="tribe" tribe="Gustavini">Gustavini</taxonomicName>
, placing it under the Aphanodactylinae in his concept of Asthenognathidae Stimpson, 1858. Whereas the morphological features of
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<emphasis id="B906EA807A02FFCA4758FD4945FAFD64" box="[1135,1239,686,712]" italics="true" pageId="1" pageNumber="370">Gustavus</emphasis>
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, in particular the somewhat peculiar Mxp3, are unusual within
<taxonomicName id="4C724D117A02FFCA41A3FD36424EFD47" authorityName="Ahyong &amp; Ng" authorityYear="2009" box="[660,867,721,747]" class="Malacostraca" family="Aphanodactylidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="family">Aphanodactylidae</taxonomicName>
, members of this family are so diverse in form that a suprageneric grouping just for
<taxonomicName id="4C724D117A02FFCA4137FD114345FCBC" authorityName="Ahyong &amp; Ng" authorityYear="2009" box="[512,616,758,784]" class="Malacostraca" family="Aphanodactylidae" genus="Gustavus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B906EA807A02FFCA4137FD114345FCBC" box="[512,616,758,784]" italics="true" pageId="1" pageNumber="370">Gustavus</emphasis>
</taxonomicName>
would not be taxonomically meaningful, even with discovery of a second species of the genus (described below). In addition, the subtriangular Mxp3 ischium of
<taxonomicName id="4C724D117A02FFCA4755FCFD45E7FC98" authorityName="Ahyong &amp; Ng" authorityYear="2009" box="[1122,1226,794,820]" class="Malacostraca" family="Aphanodactylidae" genus="Gustavus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B906EA807A02FFCA4755FCFD45E7FC98" box="[1122,1226,794,820]" italics="true" pageId="1" pageNumber="370">Gustavus</emphasis>
</taxonomicName>
, being slender and distinctly tapering proximally, is a feature also present in
<taxonomicName id="4C724D117A02FFCA4017FCDA45D7FCFB" authority="Naruse &amp; Maenosono, 2012" authorityName="Naruse &amp; Maenosono" authorityYear="2012" box="[800,1274,829,856]" class="Malacostraca" family="Aphanodactylidae" genus="Takedactylus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B906EA807A02FFCA4017FCDA429CFCFB" box="[800,945,829,855]" italics="true" pageId="1" pageNumber="370">Takedactylus</emphasis>
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</taxonomicName>
.
<taxonomicName id="4C724D117A02FFCA4633FCDA44B8FCFB" authorityName="Naruse &amp; Maenosono" authorityYear="2012" box="[1284,1429,829,855]" class="Malacostraca" family="Aphanodactylidae" genus="Takedactylus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B906EA807A02FFCA4633FCDA44B8FCFB" box="[1284,1429,829,855]" italics="true" pageId="1" pageNumber="370">Takedactylus</emphasis>
</taxonomicName>
, however, has a completely different carapace form and much longer walking legs, the latter also with a distinctive dactylus armed with an accessory spine (cf.
<bibRefCitation id="EFE34B637A02FFCA41B1FC614293FC33" author="Naruse, T. &amp; Maenosono, T." box="[646,958,901,928]" pageId="1" pageNumber="370" pagination="225 - 231" refId="ref6099" refString="Naruse, T. &amp; Maenosono, T. (2012) A new genus and species of Aphanodactylidae Ahyong &amp; Ng, 2009 (Decapoda, Brachyura, Pinnotheroidea) from the Ryukyu Islands, Japan. In: Komatsu, H., Okuno, J. &amp; Fukuoka, K. (Eds.), Studies on Eumalacostraca: a homage to Takeda Masatsune. Crustaceana Monographs, 17, 225 - 231. https: // doi. org / 10.1163 / 9789004202894 _ 019" type="journal article" year="2012">Naruse &amp; Maenosono 2012</bibRefCitation>
;
<bibRefCitation id="EFE34B637A02FFCA40FEFC6145F7FC33" author="Naruse, T. &amp; Yoshida, R." box="[969,1242,901,928]" pageId="1" pageNumber="370" pagination="21 - 33" refId="ref6189" refString="Naruse, T. &amp; Yoshida, R. (2018) Two new species of Aphanodactylidae (Crustacea: Decapoda: Brachyura) from the Ryukyu and Ogasawara Islands, Japan, with the establishment of a new genus. Tropical Zoology, 31 (1), 21 - 33. https: // doi. org / 10.1080 / 03946975.2017.1395163" type="journal article" year="2018">Naruse &amp; Yoshida 2018</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,532 @@
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<mods:title id="B9AC13D5E66F55F3F865AB2B1905B32C">Seven new species of Spatulaphorus Rack (Acari: Pygmephoridae) phoretic on scarab beetles (Coleoptera: Scarabaeidae)</mods:title>
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<mods:namePart id="55DAC2719F96E1B7A69CC01EA0598C51">Khaustov, Alexander A.</mods:namePart>
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<mods:affiliation id="C6EBB76D9899190109DB8024FF301D6F">Zoological Institute of Russian Academy of Sciences, Universitetskaya Embankment 1, 199034, Saint Petersburg, Russia</mods:affiliation>
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<mods:namePart id="B0ECC595D33503096A485AB232840B1C">Akhmetova, Lilia A.</mods:namePart>
<mods:affiliation id="2CA803CE202E0E58D6D71BD3F65F4813">Zoological Institute of Russian Academy of Sciences, Universitetskaya Embankment 1, 199034, Saint Petersburg, Russia</mods:affiliation>
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<treatment id="03EB878FFFA3745A81B43F1EFC4FFE8A" LSID="urn:lsid:plazi:treatment:03EB878FFFA3745A81B43F1EFC4FFE8A" httpUri="http://treatment.plazi.org/id/03EB878FFFA3745A81B43F1EFC4FFE8A" lastPageId="9" lastPageNumber="346" pageId="5" pageNumber="342">
<subSubSection id="C3586512FFA3745681B43F1EFD97FD8F" box="[151,585,582,609]" pageId="5" pageNumber="342" type="nomenclature">
<paragraph id="8BFD3699FFA3745681B43F1EFD97FD8F" blockId="5.[151,585,582,645]" box="[151,585,582,609]" pageId="5" pageNumber="342">
<heading id="D0B581F5FFA3745681B43F1EFD97FD8F" bold="true" box="[151,585,582,609]" fontSize="11" level="1" pageId="5" pageNumber="342" reason="1">
<emphasis id="B936EA8BFFA3745681B43F1EFD97FD8F" bold="true" box="[151,585,582,609]" pageId="5" pageNumber="342">
<taxonomicName id="4C424D1AFFA3745681B43F1EFE3AFD8E" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[151,484,582,609]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="342" phylum="Arthropoda" rank="species" species="longisetosus" status="sp. nov.">
<emphasis id="B936EA8BFFA3745681B43F1EFE3AFD8E" bold="true" box="[151,484,582,609]" italics="true" pageId="5" pageNumber="342">Spatulaphorus longisetosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20557F0FFA3745680CF3F1EFD97FD8F" box="[492,585,582,608]" pageId="5" pageNumber="342" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
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<subSubSection id="C3586512FFA3745B81B43F32FE4FF911" lastPageId="8" lastPageNumber="345" pageId="5" pageNumber="342" type="description">
<paragraph id="8BFD3699FFA3745681B43F32FED5FD6A" blockId="5.[151,585,582,645]" box="[151,267,618,645]" pageId="5" pageNumber="342">
(
<figureCitation id="13792A1CFFA3745681BC3F32FEDDFD6A" box="[159,259,618,645]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStart-3="FIGURE 8" captionStartId-0="5.[151,250,1971,1995]" captionStartId-1="6.[151,250,1541,1565]" captionStartId-2="7.[151,250,1947,1971]" captionStartId-3="8.[151,250,1154,1178]" captionTargetBox-0="[151,1435,753,1947]" captionTargetBox-1="[151,1435,421,1518]" captionTargetBox-2="[151,1435,352,1924]" captionTargetBox-3="[151,1436,181,1130]" captionTargetId-0="figure-198@5.[151,1435,740,1947]" captionTargetId-1="figure-137@6.[151,1435,417,1518]" captionTargetId-2="figure-97@7.[151,1435,345,1924]" captionTargetId-3="figure-24@8.[151,1436,181,1130]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionText-0="FIGURE 5. Spatulaphorus longisetosus sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." captionText-1="FIGURE 6. Spatulaphorus longisetosus sp. nov., female: A—left leg I, dorsal aspect; B—left leg II, dorsal aspect." captionText-2="FIGURE 7. Spatulaphorus longisetosus sp. nov., female: A—left leg III, dorsal aspect; B—left leg IV, dorsal aspect." captionText-3="FIGURE 8. DIC micrographs of Spatulaphorus longisetosus sp. nov., female: A—general view dorsally; B—general view ventrally." pageId="5" pageNumber="342">Figs 58</figureCitation>
)
</paragraph>
<paragraph id="8BFD3699FFA3745681B43FEAFAE0FD22" blockId="5.[151,1342,690,717]" box="[151,1342,690,717]" pageId="5" pageNumber="342">
<emphasis id="B936EA8BFFA3745681B43FEAFEFFFD23" bold="true" box="[151,289,690,716]" pageId="5" pageNumber="342">Description</emphasis>
. FEMALE. Body well sclerotized. Length of idiosoma 220 (195225), width 125 (110125).
</paragraph>
<caption id="DF3D6611FFA3745681B43AEBFACAF824" box="[151,1300,1971,1996]" pageId="5" pageNumber="342" startId="5.[151,250,1971,1995]" targetBox="[151,1435,753,1947]" targetPageId="5" targetType="figure">
<paragraph id="8BFD3699FFA3745681B43AEBFACAF824" blockId="5.[151,1300,1971,1996]" box="[151,1300,1971,1996]" pageId="5" pageNumber="342">
<emphasis id="B936EA8BFFA3745681B43AEBFED3F824" bold="true" box="[151,269,1971,1995]" pageId="5" pageNumber="342">FIGURE 5</emphasis>
.
<taxonomicName id="4C424D1AFFA37456803A3AEBFDEFF824" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[281,561,1971,1995]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="342" phylum="Arthropoda" rank="species" species="longisetosus" status="sp. nov.">
<emphasis id="B936EA8BFFA37456803A3AEBFDEFF824" box="[281,561,1971,1995]" italics="true" pageId="5" pageNumber="342">Spatulaphorus longisetosus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFA3745683143AEBFD56F824" bold="true" box="[567,648,1971,1995]" pageId="5" pageNumber="342">
<taxonomicNameLabel id="A20557F0FFA3745683143AEBFD56F824" box="[567,648,1971,1995]" pageId="5" pageNumber="342" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—dorsum of body; B—venter of body. Legs omitted.
</paragraph>
</caption>
<paragraph id="8BFD3699FFA0745581E43DCFFCC9FE66" blockId="6.[151,1437,151,393]" pageId="6" pageNumber="343">
<emphasis id="B936EA8BFFA0745581E43DCFFE88FF5E" box="[199,342,151,177]" italics="true" pageId="6" pageNumber="343">Gnathosoma</emphasis>
. Gnathosomal capsule, excluding palps, subrectangular, length 23 (2123), width 23 (2123). Dorsum with one pair of needle-like cheliceral setae (
<emphasis id="B936EA8BFFA07455823D3DE3FC98FF3A" box="[798,838,187,213]" italics="true" pageId="6" pageNumber="343">cha</emphasis>
) 4 (45). Dorsal median apodeme present, poorly developed. Postpalpal setae (
<emphasis id="B936EA8BFFA0745580FC3D87FE25FF17" box="[479,507,223,248]" italics="true" pageId="6" pageNumber="343">pp</emphasis>
) 3 (3) needle-like, situated laterad setae
<emphasis id="B936EA8BFFA0745582EA3D87FC2FFF16" box="[969,1009,223,249]" italics="true" pageId="6" pageNumber="343">cha</emphasis>
. Ventral gnathosoma with one pair of smooth, weakly blunt-tipped subcapitular setae
<emphasis id="B936EA8BFFA07455838D3C5BFD1CFEF3" box="[686,706,259,284]" italics="true" pageId="6" pageNumber="343">m</emphasis>
7 (67) and round pits
<emphasis id="B936EA8BFFA0745582E93C5BFC06FEF3" box="[970,984,259,284]" italics="true" pageId="6" pageNumber="343">n</emphasis>
situated posteriad
<emphasis id="B936EA8BFFA0745585883C5BFB61FEF3" box="[1195,1215,259,284]" italics="true" pageId="6" pageNumber="343">m</emphasis>
. Palps with smooth and weakly blunt-tipped setae
<emphasis id="B936EA8BFFA0745580C93C7FFDC8FEAE" box="[490,534,295,321]" italics="true" pageId="6" pageNumber="343">dFe</emphasis>
3 (23) and
<emphasis id="B936EA8BFFA0745583813C7FFD0FFEAE" box="[674,721,295,321]" italics="true" pageId="6" pageNumber="343">dGe</emphasis>
7 (67) dorsolaterally. Palps ventrally each with mushroom-like accessory setigenous structure (
<emphasis id="B936EA8BFFA0745580DA3C13FDC3FE8B" box="[505,541,331,356]" italics="true" pageId="6" pageNumber="343">ass</emphasis>
) and tiny solenidion
<emphasis id="B936EA8BFFA0745582293C13FCC0FE8B" box="[778,798,331,356]" italics="true" pageId="6" pageNumber="343">ω</emphasis>
. Palps terminated with small blunt-tipped tibial claw and tiny eupathid-like seta. Pharyngeal pumps poorly visible.
</paragraph>
<caption id="DF3D6611FFA0745581B43B5DFAFBF9F2" box="[151,1317,1541,1566]" pageId="6" pageNumber="343" startId="6.[151,250,1541,1565]" targetBox="[151,1435,421,1518]" targetPageId="6" targetType="figure">
<paragraph id="8BFD3699FFA0745581B43B5DFAFBF9F2" blockId="6.[151,1317,1541,1566]" box="[151,1317,1541,1566]" pageId="6" pageNumber="343">
<emphasis id="B936EA8BFFA0745581B43B5DFED3F9F2" bold="true" box="[151,269,1541,1565]" pageId="6" pageNumber="343">FIGURE 6</emphasis>
.
<taxonomicName id="4C424D1AFFA07455803A3B5EFDEFF9F1" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[281,561,1542,1566]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="343" phylum="Arthropoda" rank="species" species="longisetosus" status="sp. nov.">
<emphasis id="B936EA8BFFA07455803A3B5EFDEFF9F1" box="[281,561,1542,1566]" italics="true" pageId="6" pageNumber="343">Spatulaphorus longisetosus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFA0745583143B5EFD56F9F1" bold="true" box="[567,648,1542,1566]" pageId="6" pageNumber="343">
<taxonomicNameLabel id="A20557F0FFA0745583143B5EFD56F9F1" box="[567,648,1542,1566]" pageId="6" pageNumber="343" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—left leg I, dorsal aspect; B—left leg II, dorsal aspect.
</paragraph>
</caption>
<paragraph id="8BFD3699FFA0745581E43B14FC2BF88D" blockId="6.[151,1437,1612,2034]" pageId="6" pageNumber="343">
<emphasis id="B936EA8BFFA0745581E43B14FE4DF989" box="[199,403,1612,1638]" italics="true" pageId="6" pageNumber="343">Idiosomal dorsum</emphasis>
(
<figureCitation id="13792A1CFFA0745580813B14FDDEF988" box="[418,512,1612,1639]" captionStart="FIGURE 5" captionStartId="5.[151,250,1971,1995]" captionTargetBox="[151,1435,753,1947]" captionTargetId="figure-198@5.[151,1435,740,1947]" captionTargetPageId="5" captionText="FIGURE 5. Spatulaphorus longisetosus sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." pageId="6" pageNumber="343">Figs 5A</figureCitation>
,
<figureCitation id="13792A1CFFA07455832E3B14FDECF989" box="[525,562,1612,1638]" captionStart="FIGURE 8" captionStartId="8.[151,250,1154,1178]" captionTargetBox="[151,1436,181,1130]" captionTargetId="figure-24@8.[151,1436,181,1130]" captionTargetPageId="8" captionText="FIGURE 8. DIC micrographs of Spatulaphorus longisetosus sp. nov., female: A—general view dorsally; B—general view ventrally." pageId="6" pageNumber="343">8A</figureCitation>
). All dorsal shields with big puncta. Prodorsal shield with three pairs of setae (
<emphasis id="B936EA8BFFA0745581823B28FF64F965" box="[161,186,1648,1674]" italics="true" pageId="6" pageNumber="343">v1</emphasis>
,
<emphasis id="B936EA8BFFA0745581EE3B28FF36F965" box="[205,232,1648,1674]" italics="true" pageId="6" pageNumber="343">v2</emphasis>
,
<emphasis id="B936EA8BFFA0745581D93B28FEC1F965" box="[250,287,1648,1674]" italics="true" pageId="6" pageNumber="343">sc2</emphasis>
), one pair of weakly barbed capitate trichobothria (
<emphasis id="B936EA8BFFA0745582A23B28FC79F965" box="[897,935,1648,1674]" italics="true" pageId="6" pageNumber="343">sc1</emphasis>
) and one pair of round stigmata. Stigmata associated with well-developed tracheal trunks.All dorsal idiosomal setae blunt-tipped; seta
<emphasis id="B936EA8BFFA0745585BA3BCCFB6DF941" box="[1177,1203,1684,1710]" italics="true" pageId="6" pageNumber="343">v2</emphasis>
smooth, other dorsal setae distinctly barbed. Alveoli of setae
<emphasis id="B936EA8BFFA07455837F3BE1FDB6F93D" box="[604,616,1721,1746]" italics="true" pageId="6" pageNumber="343">e</emphasis>
and
<emphasis id="B936EA8BFFA0745583823BE0FD77F93D" box="[673,681,1720,1746]" italics="true" pageId="6" pageNumber="343">f</emphasis>
clearly separated. Cupules
<emphasis id="B936EA8BFFA0745582C13BE1FC26F93D" box="[994,1016,1721,1746]" italics="true" pageId="6" pageNumber="343">ia</emphasis>
on tergite D,
<emphasis id="B936EA8BFFA0745585B53BE1FB6CF93D" box="[1174,1202,1721,1746]" italics="true" pageId="6" pageNumber="343">im</emphasis>
on tergite EF and
<emphasis id="B936EA8BFFA0745584A53BE0FA42F93D" box="[1414,1436,1720,1746]" italics="true" pageId="6" pageNumber="343">ih</emphasis>
on tergite H large, round, subequal. Lengths of dorsal setae:
<emphasis id="B936EA8BFFA07455826A3B84FCBDF919" box="[841,867,1756,1782]" italics="true" pageId="6" pageNumber="343">v1</emphasis>
32 (2932),
<emphasis id="B936EA8BFFA0745582DB3B84FBCDF919" box="[1016,1043,1756,1782]" italics="true" pageId="6" pageNumber="343">v2</emphasis>
13 (1314),
<emphasis id="B936EA8BFFA0745585843B84FB13F919" box="[1191,1229,1756,1782]" italics="true" pageId="6" pageNumber="343">sc2</emphasis>
53 (4553),
<emphasis id="B936EA8BFFA0745584423B84FAA2F919" box="[1377,1404,1756,1782]" italics="true" pageId="6" pageNumber="343">c1</emphasis>
52 (4352),
<emphasis id="B936EA8BFFA0745581DE3A58FEC6F8F5" box="[253,280,1792,1818]" italics="true" pageId="6" pageNumber="343">c2</emphasis>
49 (4149),
<emphasis id="B936EA8BFFA0745580843A58FE6BF8F5" box="[423,437,1792,1818]" italics="true" pageId="6" pageNumber="343">d</emphasis>
52 (4552),
<emphasis id="B936EA8BFFA0745583673A59FD8EF8F5" box="[580,592,1793,1818]" italics="true" pageId="6" pageNumber="343">e</emphasis>
26 (2426),
<emphasis id="B936EA8BFFA0745583FC3A58FD39F8F5" box="[735,743,1792,1818]" italics="true" pageId="6" pageNumber="343">f</emphasis>
61 (4861),
<emphasis id="B936EA8BFFA0745582553A58FC4FF8F5" box="[886,913,1792,1818]" italics="true" pageId="6" pageNumber="343">h1</emphasis>
55 (4655),
<emphasis id="B936EA8BFFA0745585023A58FBE3F8F5" box="[1057,1085,1792,1818]" italics="true" pageId="6" pageNumber="343">h2</emphasis>
15 (1516). Distances between setae:
<emphasis id="B936EA8BFFA0745581C13A7CFEFAF8D1" box="[226,292,1828,1854]" italics="true" pageId="6" pageNumber="343">v1v1</emphasis>
26 (2427),
<emphasis id="B936EA8BFFA0745580E23A7CFDDDF8D1" box="[449,515,1828,1854]" italics="true" pageId="6" pageNumber="343">v2v2</emphasis>
41 (3741),
<emphasis id="B936EA8BFFA0745583833A7CFD26F8D1" box="[672,760,1828,1854]" italics="true" pageId="6" pageNumber="343">sc2sc2</emphasis>
41 (3643),
<emphasis id="B936EA8BFFA0745582B63A7CFC06F8D1" box="[917,984,1828,1854]" italics="true" pageId="6" pageNumber="343">c1c1</emphasis>
46 (4048),
<emphasis id="B936EA8BFFA0745585533A7CFB6DF8D1" box="[1136,1203,1828,1854]" italics="true" pageId="6" pageNumber="343">c1c2</emphasis>
22 (2022),
<emphasis id="B936EA8BFFA07455846C3A7CFAA7F8D1" box="[1359,1401,1828,1854]" italics="true" pageId="6" pageNumber="343">dd</emphasis>
61 (5661),
<emphasis id="B936EA8BFFA0745581DD3A11FEFEF88D" box="[254,288,1864,1890]" italics="true" pageId="6" pageNumber="343">ef</emphasis>
10 (810),
<emphasis id="B936EA8BFFA0745580803A10FE1EF88D" box="[419,448,1864,1890]" italics="true" pageId="6" pageNumber="343">ff</emphasis>
52 (4753),
<emphasis id="B936EA8BFFA0745583723A10FD49F88D" box="[593,663,1864,1890]" italics="true" pageId="6" pageNumber="343">h1h1</emphasis>
45 (3845),
<emphasis id="B936EA8BFFA07455820B3A10FCB0F88D" box="[808,878,1864,1890]" italics="true" pageId="6" pageNumber="343">h1h2</emphasis>
11 (1112).
</paragraph>
<paragraph id="8BFD3699FFA0745481E43A34FAFCFEF2" blockId="6.[151,1437,1612,2034]" lastBlockId="7.[151,1437,150,285]" lastPageId="7" lastPageNumber="344" pageId="6" pageNumber="343">
<emphasis id="B936EA8BFFA0745581E43A34FE5DF869" box="[199,387,1900,1926]" italics="true" pageId="6" pageNumber="343">Idiosomal venter</emphasis>
(
<figureCitation id="13792A1CFFA0745580B23A34FE35F868" box="[401,491,1900,1927]" captionStart="FIGURE 5" captionStartId="5.[151,250,1971,1995]" captionTargetBox="[151,1435,753,1947]" captionTargetId="figure-198@5.[151,1435,740,1947]" captionTargetPageId="5" captionText="FIGURE 5. Spatulaphorus longisetosus sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." pageId="6" pageNumber="343">Figs 5B</figureCitation>
,
<figureCitation id="13792A1CFFA0745580D53A34FDC4F869" box="[502,538,1900,1926]" captionStart="FIGURE 8" captionStartId="8.[151,250,1154,1178]" captionTargetBox="[151,1436,181,1130]" captionTargetId="figure-24@8.[151,1436,181,1130]" captionTargetPageId="8" captionText="FIGURE 8. DIC micrographs of Spatulaphorus longisetosus sp. nov., female: A—general view dorsally; B—general view ventrally." pageId="6" pageNumber="343">8B</figureCitation>
). Coxisternal fields III, III and aggenital plate laterally with sparsely distributed big puncta. Setae
<emphasis id="B936EA8BFFA0745580403AC8FE5EF845" box="[355,384,1936,1962]" italics="true" pageId="6" pageNumber="343">1b</emphasis>
pointed and weakly barbed; other ventral setae smooth and blunt-tipped. Bases of setae
<emphasis id="B936EA8BFFA0745584563AC8FA42F845" box="[1397,1436,1936,1962]" italics="true" pageId="6" pageNumber="343">ps3</emphasis>
situated distinctly anteriad bases of setae
<emphasis id="B936EA8BFFA0745583593AECFD61F821" box="[634,703,1972,1998]" italics="true" pageId="6" pageNumber="343">ps1-2.</emphasis>
Ap1 and ap2 well-developed and joined with thick appr; apsej developed only laterally; ap3 weak, not reaching appo; ap4 well developed, exceeding beyond bases of setae
<emphasis id="B936EA8BFFA07455845A3A80FA4BF81D" box="[1401,1429,2008,2034]" italics="true" pageId="6" pageNumber="343">3b</emphasis>
. Ap5 short, joined with appo and reaching bases of setae
<emphasis id="B936EA8BFFA1745482273DCFFCFFFF5E" box="[772,801,151,177]" italics="true" pageId="7" pageNumber="344">4a</emphasis>
. Ags small, cup-like; pgs small, oval. Posterior margin of posterior sternal plate straight in middle part. Posterior margin of aggenital plate rounded. Lengths of ventral setae:
<emphasis id="B936EA8BFFA1745481B43D87FF6AFF16" box="[151,180,223,249]" italics="true" pageId="7" pageNumber="344">1a</emphasis>
11 (1011),
<emphasis id="B936EA8BFFA1745480663D87FEBFFF16" box="[325,353,223,249]" italics="true" pageId="7" pageNumber="344">1b</emphasis>
23 (2225),
<emphasis id="B936EA8BFFA1745480D63D87FDD1FF16" box="[501,527,223,249]" italics="true" pageId="7" pageNumber="344">1с</emphasis>
14 (1214),
<emphasis id="B936EA8BFFA1745483803D87FD60FF16" box="[675,702,223,249]" italics="true" pageId="7" pageNumber="344">2a</emphasis>
16 (1517),
<emphasis id="B936EA8BFFA1745482713D87FCB2FF16" box="[850,876,223,249]" italics="true" pageId="7" pageNumber="344">2с</emphasis>
11 (1112),
<emphasis id="B936EA8BFFA1745482DD3D87FBC4FF16" box="[1022,1050,223,249]" italics="true" pageId="7" pageNumber="344">3a</emphasis>
15 (1415),
<emphasis id="B936EA8BFFA17454858D3D87FB14FF16" box="[1198,1226,223,249]" italics="true" pageId="7" pageNumber="344">3b</emphasis>
14 (1214),
<emphasis id="B936EA8BFFA17454847D3D87FAA6FF16" box="[1374,1400,223,249]" italics="true" pageId="7" pageNumber="344">3c</emphasis>
17 (1517),
<emphasis id="B936EA8BFFA1745481DD3C5BFEC4FEF2" box="[254,282,259,285]" italics="true" pageId="7" pageNumber="344">4a</emphasis>
13 (1213),
<emphasis id="B936EA8BFFA1745480893C5BFE19FEF2" box="[426,455,259,285]" italics="true" pageId="7" pageNumber="344">4b</emphasis>
23 (1923),
<emphasis id="B936EA8BFFA1745483743C5BFDACFEF2" box="[599,626,259,285]" italics="true" pageId="7" pageNumber="344">4c</emphasis>
16 (1516),
<emphasis id="B936EA8BFFA1745482213C5BFCF7FEF2" box="[770,809,259,285]" italics="true" pageId="7" pageNumber="344">ps1</emphasis>
18 (1618),
<emphasis id="B936EA8BFFA1745482993C5BFC3FFEF2" box="[954,993,259,285]" italics="true" pageId="7" pageNumber="344">ps2</emphasis>
18 (1618),
<emphasis id="B936EA8BFFA1745485523C5BFB46FEF2" box="[1137,1176,259,285]" italics="true" pageId="7" pageNumber="344">ps3</emphasis>
15 (1415).
</paragraph>
<caption id="DF3D6611FFA1745481B43AC3FAE2F85C" box="[151,1340,1947,1971]" pageId="7" pageNumber="344" startId="7.[151,250,1947,1971]" targetBox="[151,1435,352,1924]" targetPageId="7" targetType="figure">
<paragraph id="8BFD3699FFA1745481B43AC3FAE2F85C" blockId="7.[151,1340,1947,1971]" box="[151,1340,1947,1971]" pageId="7" pageNumber="344">
<emphasis id="B936EA8BFFA1745481B43AC3FED3F85C" bold="true" box="[151,269,1947,1971]" pageId="7" pageNumber="344">FIGURE 7</emphasis>
.
<taxonomicName id="4C424D1AFFA17454803A3AC3FDEFF85C" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[281,561,1947,1971]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="344" phylum="Arthropoda" rank="species" species="longisetosus" status="sp. nov.">
<emphasis id="B936EA8BFFA17454803A3AC3FDEFF85C" box="[281,561,1947,1971]" italics="true" pageId="7" pageNumber="344">Spatulaphorus longisetosus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFA1745483143AC3FD56F85C" bold="true" box="[567,648,1947,1971]" pageId="7" pageNumber="344">
<taxonomicNameLabel id="A20557F0FFA1745483143AC3FD56F85C" box="[567,648,1947,1971]" pageId="7" pageNumber="344" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—left leg III, dorsal aspect; B—left leg IV, dorsal aspect.
</paragraph>
</caption>
<caption id="DF3D6611FFAE745B81B439DAFF29FB51" pageId="8" pageNumber="345" startId="8.[151,250,1154,1178]" targetBox="[151,1436,181,1130]" targetPageId="8" targetType="figure">
<paragraph id="8BFD3699FFAE745B81B439DAFF29FB51" blockId="8.[151,1436,1154,1214]" pageId="8" pageNumber="345">
<emphasis id="B936EA8BFFAE745B81B439DAFECEFB75" bold="true" box="[151,272,1154,1178]" pageId="8" pageNumber="345">FIGURE 8</emphasis>
. DIC micrographs of
<taxonomicName id="4C424D1AFFAE745B80DA39DAFCCDFB75" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[505,787,1154,1178]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="345" phylum="Arthropoda" rank="species" species="longisetosus" status="sp. nov.">
<emphasis id="B936EA8BFFAE745B80DA39DAFCCDFB75" box="[505,787,1154,1178]" italics="true" pageId="8" pageNumber="345">Spatulaphorus longisetosus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFAE745B823F39DAFCB1FB75" bold="true" box="[796,879,1154,1178]" pageId="8" pageNumber="345">
<taxonomicNameLabel id="A20557F0FFAE745B823F39DAFCB1FB75" box="[796,879,1154,1178]" pageId="8" pageNumber="345" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—general view dorsally; B—general view ventrally.
</paragraph>
</caption>
<paragraph id="8BFD3699FFAE745B81E439B5FAE1F935" blockId="8.[151,1437,1260,2043]" pageId="8" pageNumber="345">
<emphasis id="B936EA8BFFAE745B81E439B5FF22FAE9" box="[199,252,1261,1286]" italics="true" pageId="8" pageNumber="345">Legs</emphasis>
(
<figureCitation id="13792A1CFFAE745B802F39B4FE88FAE9" box="[268,342,1260,1286]" captionStart="FIGURE 6" captionStartId="6.[151,250,1541,1565]" captionTargetBox="[151,1435,421,1518]" captionTargetId="figure-137@6.[151,1435,417,1518]" captionTargetPageId="6" captionText="FIGURE 6. Spatulaphorus longisetosus sp. nov., female: A—left leg I, dorsal aspect; B—left leg II, dorsal aspect." pageId="8" pageNumber="345">Figs 6</figureCitation>
,
<figureCitation id="13792A1CFFAE745B804039B4FEADFAE9" box="[355,371,1260,1286]" captionStart="FIGURE 7" captionStartId="7.[151,250,1947,1971]" captionTargetBox="[151,1435,352,1924]" captionTargetId="figure-97@7.[151,1435,345,1924]" captionTargetPageId="7" captionText="FIGURE 7. Spatulaphorus longisetosus sp. nov., female: A—left leg III, dorsal aspect; B—left leg IV, dorsal aspect." pageId="8" pageNumber="345">7</figureCitation>
). Setation of legs as in
<taxonomicName id="4C424D1AFFAE745B835C39B4FCFBFAE9" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[639,805,1260,1286]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="345" phylum="Arthropoda" rank="species" species="brevisetosus">
<emphasis id="B936EA8BFFAE745B835C39B4FCFBFAE9" box="[639,805,1260,1286]" italics="true" pageId="8" pageNumber="345">S. brevisetosus</emphasis>
</taxonomicName>
. Leg I (
<figureCitation id="13792A1CFFAE745B82A139B4FC00FAE9" box="[898,990,1260,1286]" captionStart="FIGURE 6" captionStartId="6.[151,250,1541,1565]" captionTargetBox="[151,1435,421,1518]" captionTargetId="figure-137@6.[151,1435,417,1518]" captionTargetPageId="6" captionText="FIGURE 6. Spatulaphorus longisetosus sp. nov., female: A—left leg I, dorsal aspect; B—left leg II, dorsal aspect." pageId="8" pageNumber="345">Fig. 6A</figureCitation>
). Tibiotarsus enlarged, with large claw; tibiotarsus with internal oval strongly sclerotized structure near solenidion
<emphasis id="B936EA8BFFAE745B82CB3848FBDBFAC5" box="[1000,1029,1296,1322]" italics="true" pageId="8" pageNumber="345">φ2</emphasis>
. Setae (
<emphasis id="B936EA8BFFAE745B85413849FBAEFAC5" box="[1122,1136,1297,1322]" italics="true" pageId="8" pageNumber="345">u</emphasis>
) consolidated into a horn-like structure opposing claw. Tarsal claw blunt-tipped. Seta
<emphasis id="B936EA8BFFAE745B8210386CFC9FFAA1" box="[819,833,1332,1358]" italics="true" pageId="8" pageNumber="345">d</emphasis>
of femur spatulate, with short subterminal projection. Setae
<emphasis id="B936EA8BFFAE745B81F93800FF35FA9D" box="[218,235,1368,1394]" italics="true" pageId="8" pageNumber="345">l</emphasis>
of femur,
<emphasis id="B936EA8BFFAE745B807D3800FEB4FA9D" box="[350,362,1368,1394]" italics="true" pageId="8" pageNumber="345">k</emphasis>
and
<emphasis id="B936EA8BFFAE745B80833800FE18FA9D" box="[416,454,1368,1394]" italics="true" pageId="8" pageNumber="345">pl”</emphasis>
of tibiotarsus smooth; other leg setae (except eupathidia
<emphasis id="B936EA8BFFAE745B85633800FB89FA9D" box="[1088,1111,1368,1394]" italics="true" pageId="8" pageNumber="345">p</emphasis>
,
<emphasis id="B936EA8BFFAE745B85473800FB5CFA9D" box="[1124,1154,1368,1394]" italics="true" pageId="8" pageNumber="345">p”</emphasis>
,
<emphasis id="B936EA8BFFAE745B85AC3800FB73FA9D" box="[1167,1197,1368,1394]" italics="true" pageId="8" pageNumber="345">tc</emphasis>
,
<emphasis id="B936EA8BFFAE745B85993800FB00FA9D" box="[1210,1246,1368,1394]" italics="true" pageId="8" pageNumber="345">tc”</emphasis>
,
<emphasis id="B936EA8BFFAE745B85C83800FADAFA9D" box="[1259,1284,1368,1394]" italics="true" pageId="8" pageNumber="345">ft</emphasis>
,
<emphasis id="B936EA8BFFAE745B84313800FAEFFA9D" box="[1298,1329,1368,1394]" italics="true" pageId="8" pageNumber="345">ft”</emphasis>
) sparsely barbed. Setae
<emphasis id="B936EA8BFFAE745B80113824FE9DFA79" box="[306,323,1404,1430]" italics="true" pageId="8" pageNumber="345">l</emphasis>
,
<emphasis id="B936EA8BFFAE745B80733824FEB9FA79" box="[336,359,1404,1430]" italics="true" pageId="8" pageNumber="345">l”</emphasis>
of femur, (
<emphasis id="B936EA8BFFAE745B80C13824FE34FA79" box="[482,490,1404,1430]" italics="true" pageId="8" pageNumber="345">l</emphasis>
) of genu and
<emphasis id="B936EA8BFFAE745B83A33824FD52FA79" box="[640,652,1404,1430]" italics="true" pageId="8" pageNumber="345">k</emphasis>
of tibiotarsus blunt-tipped; other leg setae (except eupathidia) pointed. Lengths of solenidia
<emphasis id="B936EA8BFFAE745B80A238F8FE7DFA55" box="[385,419,1440,1466]" italics="true" pageId="8" pageNumber="345">ω1</emphasis>
8 (89),
<emphasis id="B936EA8BFFAE745B832B38F8FDF4FA55" box="[520,554,1440,1466]" italics="true" pageId="8" pageNumber="345">ω2</emphasis>
5 (45),
<emphasis id="B936EA8BFFAE745B83AC38F8FD72FA55" box="[655,684,1440,1466]" italics="true" pageId="8" pageNumber="345">φ1</emphasis>
8 (8),
<emphasis id="B936EA8BFFAE745B83D538F8FCCDFA55" box="[758,787,1440,1466]" italics="true" pageId="8" pageNumber="345">φ2</emphasis>
5 (45); solenidion
<emphasis id="B936EA8BFFAE745B82D738F8FBCFFA55" box="[1012,1041,1440,1466]" italics="true" pageId="8" pageNumber="345">φ1</emphasis>
thick, clavate,
<emphasis id="B936EA8BFFAE745B859838F8FB02FA55" box="[1211,1244,1440,1466]" italics="true" pageId="8" pageNumber="345">ω1</emphasis>
digitiform; other solenidia weakly clavate. Leg II (
<figureCitation id="13792A1CFFAE745B8328389CFDBDFA31" box="[523,611,1476,1502]" captionStart="FIGURE 6" captionStartId="6.[151,250,1541,1565]" captionTargetBox="[151,1435,421,1518]" captionTargetId="figure-137@6.[151,1435,417,1518]" captionTargetPageId="6" captionText="FIGURE 6. Spatulaphorus longisetosus sp. nov., female: A—left leg I, dorsal aspect; B—left leg II, dorsal aspect." pageId="8" pageNumber="345">Fig. 6B</figureCitation>
). Tarsus with thickened basally claws and pad-like empodium. Solenidion
<emphasis id="B936EA8BFFAE745B81B438B1FF75F9ED" box="[151,171,1513,1538]" italics="true" pageId="8" pageNumber="345">ω</emphasis>
5 (56) weakly clavate, solenidion
<emphasis id="B936EA8BFFAE745B831D38B1FD93F9ED" box="[574,589,1513,1538]" italics="true" pageId="8" pageNumber="345">φ</emphasis>
absent. Setae
<emphasis id="B936EA8BFFAE745B83CD38B0FCCAF9ED" box="[750,788,1512,1538]" italics="true" pageId="8" pageNumber="345">pl”</emphasis>
,
<emphasis id="B936EA8BFFAE745B820138B0FCE7F9ED" box="[802,825,1512,1538]" italics="true" pageId="8" pageNumber="345">u</emphasis>
and
<emphasis id="B936EA8BFFAE745B825338B0FC4AF9ED" box="[880,916,1512,1538]" italics="true" pageId="8" pageNumber="345">tc”</emphasis>
of tarsus smooth, other setae barbed; setae
<emphasis id="B936EA8BFFAE745B845C38B0FA43F9ED" box="[1407,1437,1512,1538]" italics="true" pageId="8" pageNumber="345">tc</emphasis>
and
<emphasis id="B936EA8BFFAE745B81E43B54FF33F9C9" box="[199,237,1548,1574]" italics="true" pageId="8" pageNumber="345">pl”</emphasis>
of tarsus spiniform, blunt-tipped; setae
<emphasis id="B936EA8BFFAE745B838C3B54FD63F9C9" box="[687,701,1548,1574]" italics="true" pageId="8" pageNumber="345">d</emphasis>
,
<emphasis id="B936EA8BFFAE745B83EF3B54FD03F9C9" box="[716,733,1548,1574]" italics="true" pageId="8" pageNumber="345">l</emphasis>
, and
<emphasis id="B936EA8BFFAE745B82383B54FCE9F9C9" box="[795,823,1548,1574]" italics="true" pageId="8" pageNumber="345">v”</emphasis>
of femur blunt-tipped; other leg setae pointed. Leg III (
<figureCitation id="13792A1CFFAE745B81BC3B68FF24F9A5" box="[159,250,1584,1610]" captionStart="FIGURE 7" captionStartId="7.[151,250,1947,1971]" captionTargetBox="[151,1435,352,1924]" captionTargetId="figure-97@7.[151,1435,345,1924]" captionTargetPageId="7" captionText="FIGURE 7. Spatulaphorus longisetosus sp. nov., female: A—left leg III, dorsal aspect; B—left leg IV, dorsal aspect." pageId="8" pageNumber="345">Fig. 7A</figureCitation>
). Femur divided into basi- and telofemur. Claws and empodium of same shape as on tarsus II. Seta
<emphasis id="B936EA8BFFAE745B84453B68FAA3F9A5" box="[1382,1405,1584,1610]" italics="true" pageId="8" pageNumber="345">u</emphasis>
of tarsus smooth, other setae barbed; seta
<emphasis id="B936EA8BFFAE745B836D3B0CFDAAF981" box="[590,628,1620,1646]" italics="true" pageId="8" pageNumber="345">pl”</emphasis>
of tarsus spiniform; setae
<emphasis id="B936EA8BFFAE745B82BE3B0CFC6DF981" box="[925,947,1620,1646]" italics="true" pageId="8" pageNumber="345">v</emphasis>
of trochanter,
<emphasis id="B936EA8BFFAE745B85743B0CFBBBF981" box="[1111,1125,1620,1646]" italics="true" pageId="8" pageNumber="345">d</emphasis>
,
<emphasis id="B936EA8BFFAE745B85573B0CFB54F981" box="[1140,1162,1620,1646]" italics="true" pageId="8" pageNumber="345">v</emphasis>
of femur, and
<emphasis id="B936EA8BFFAE745B840C3B0CFA9EF981" box="[1327,1344,1620,1646]" italics="true" pageId="8" pageNumber="345">l</emphasis>
of genu blunt-tipped, other leg setae pointed; solenidion
<emphasis id="B936EA8BFFAE745B83903B21FD1CF97D" box="[691,706,1657,1682]" italics="true" pageId="8" pageNumber="345">φ</emphasis>
absent. Leg IV (
<figureCitation id="13792A1CFFAE745B825F3B20FC10F97D" box="[892,974,1656,1682]" captionStart="FIGURE 7" captionStartId="7.[151,250,1947,1971]" captionTargetBox="[151,1435,352,1924]" captionTargetId="figure-97@7.[151,1435,345,1924]" captionTargetPageId="7" captionText="FIGURE 7. Spatulaphorus longisetosus sp. nov., female: A—left leg III, dorsal aspect; B—left leg IV, dorsal aspect." pageId="8" pageNumber="345">Fig 7B</figureCitation>
). Femur divided into basi- and telofemur. Claws simple, empodium as on tarsi II and III. All setae barbed; setae
<emphasis id="B936EA8BFFAE745B82833BC4FC68F959" box="[928,950,1692,1718]" italics="true" pageId="8" pageNumber="345">v</emphasis>
of trochanter,
<emphasis id="B936EA8BFFAE745B857B3BC4FBB8F959" box="[1112,1126,1692,1718]" italics="true" pageId="8" pageNumber="345">d</emphasis>
,
<emphasis id="B936EA8BFFAE745B85503BC4FB57F959" box="[1139,1161,1692,1718]" italics="true" pageId="8" pageNumber="345">v</emphasis>
of femur,
<emphasis id="B936EA8BFFAE745B85DF3BC4FAC9F959" box="[1276,1303,1692,1718]" italics="true" pageId="8" pageNumber="345">v”</emphasis>
of tibia and
<emphasis id="B936EA8BFFAE745B81B43B98FF63F935" box="[151,189,1728,1754]" italics="true" pageId="8" pageNumber="345">pl”</emphasis>
of tarsus blunt-tipped; seta
<emphasis id="B936EA8BFFAE745B80D73B98FDDBF935" box="[500,517,1728,1754]" italics="true" pageId="8" pageNumber="345">l</emphasis>
of tibia weakly blunt-tipped, other leg setae pointed; solenidion
<emphasis id="B936EA8BFFAE745B85FA3B99FB36F935" box="[1241,1256,1729,1754]" italics="true" pageId="8" pageNumber="345">φ</emphasis>
absent.
</paragraph>
<paragraph id="8BFD3699FFAE745B81EB3BBCFE4FF911" blockId="8.[151,1437,1260,2043]" box="[200,401,1764,1790]" pageId="8" pageNumber="345">MALE unknown.</paragraph>
</subSubSection>
<subSubSection id="C3586512FFAE745B81E43A50FD9BF839" pageId="8" pageNumber="345" type="materials_examined">
<paragraph id="8BFD3699FFAE745B81E43A50FE12F884" blockId="8.[151,1437,1260,2043]" pageId="8" pageNumber="345">
<emphasis id="B936EA8BFFAE745B81E43A50FEAEF8CD" bold="true" box="[199,368,1800,1826]" pageId="8" pageNumber="345">Type material</emphasis>
.
<materialsCitation id="3B2A3CC4FFAE745B80A23A50FE17F884" collectingDate="1984-04" collectingDateMax="1986-09-09" collectingDateMin="1984-04" collectionCode="ZISP, CEMT" country="United States of America" location="Tomball" pageId="8" pageNumber="345" specimenCount="7" specimenCount-female="7" stateProvince="Louisiana" typeStatus="holotype">
Female
<typeStatus id="54F9883BFFAE745B80FE3A50FD9CF8CD" box="[477,578,1800,1826]" pageId="8" pageNumber="345" type="holotype">holotype</typeStatus>
, slide
<collectionCode id="ED53AE5CFFAE745B83AC3A50FD16F8CD" box="[655,712,1800,1826]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="8" pageNumber="345">ZISP</collectionCode>
T-Pygm-011,
<collectingCountry id="F3557609FFAE745B824E3A50FC76F8CD" box="[877,936,1800,1826]" name="United States of America" pageId="8" pageNumber="345">USA</collectingCountry>
,
<collectingRegion id="4986F87BFFAE745B82963A50FBF8F8CD" box="[949,1062,1800,1826]" country="United States of America" name="Louisiana" pageId="8" pageNumber="345">Louisiana</collectingRegion>
,
<date id="FFFC1059FFAE745B85173A50FB50F8CD" box="[1076,1166,1800,1826]" pageId="8" pageNumber="345" value="1984-04">
<collectingDate id="EFB8E9B1FFAE745B85173A50FB50F8CD" box="[1076,1166,1800,1826]" pageId="8" pageNumber="345" value="1984-04">IV.1984</collectingDate>
</date>
(
<collectionCode id="ED53AE5CFFAE745B85BC3A50FB2CF8CD" box="[1183,1266,1800,1826]" pageId="8" pageNumber="345">CEMT</collectionCode>
),
<collectedFrom id="385E7564FFAE745B842B3A51FEB2F8A9" pageId="8" pageNumber="345">
on
<taxonomicName id="4C424D1AFFAE745B840D3A50FEB2F8A9" authority="LeConte" authorityName="LeConte" authorityYear="1847" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="345" phylum="Arthropoda" rank="species" species="difformis">
<emphasis id="B936EA8BFFAE745B840D3A50FF22F8A9" italics="true" pageId="8" pageNumber="345">Phanaeus difformis</emphasis>
LeConte
</taxonomicName>
</collectedFrom>
;
<typeStatus id="54F9883BFFAE745B805F3A75FE32F8A8" box="[380,492,1837,1863]" pageId="8" pageNumber="345" type="paratype">paratypes</typeStatus>
:
<specimenCount id="9D44FD10FFAE745B80D83A74FDB1F8A9" box="[507,623,1836,1863]" count="5" pageId="8" pageNumber="345" type="female">5 females</specimenCount>
, same data;
<specimenCount id="9D44FD10FFAE745B82253A74FCA4F8A9" box="[774,890,1836,1862]" count="2" pageId="8" pageNumber="345" type="female">2 females</specimenCount>
,
<collectingCountry id="F3557609FFAE745B82A93A74FC1BF8A9" box="[906,965,1836,1862]" name="United States of America" pageId="8" pageNumber="345">USA</collectingCountry>
,
<collectingRegion id="4986F87BFFAE745B82F73A75FBC6F8A8" box="[980,1048,1837,1863]" country="United States of America" name="Texas" pageId="8" pageNumber="345">Texas</collectingRegion>
,
<location id="8E9D6042FFAE745B85043A74FB5AF8A9" LSID="urn:lsid:plazi:treatment:03EB878FFFA3745A81B43F1EFC4FFE8A:8E9D6042FFAE745B85043A74FB5AF8A9" box="[1063,1156,1836,1862]" country="United States of America" name="Tomball" pageId="8" pageNumber="345" stateProvince="Louisiana">Tomball</location>
<date id="FFFC1059FFAE745B85B33A74FADFF8A9" box="[1168,1281,1836,1862]" pageId="8" pageNumber="345" value="1986-09-09">
<collectingDate id="EFB8E9B1FFAE745B85B33A74FADFF8A9" box="[1168,1281,1836,1862]" pageId="8" pageNumber="345" value="1986-09-09">9.IX.1986</collectingDate>
</date>
(
<collectionCode id="ED53AE5CFFAE745B84363A74FAB6F8A9" box="[1301,1384,1836,1862]" pageId="8" pageNumber="345">CEMT</collectionCode>
), on
<taxonomicName id="4C424D1AFFAE745B81B43A08FE17F884" authority="MacLeay." authorityName="MacLeay." box="[151,457,1872,1899]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="345" phylum="Arthropoda" rank="species" species="vindex">
<emphasis id="B936EA8BFFAE745B81B43A08FE88F885" box="[151,342,1872,1898]" italics="true" pageId="8" pageNumber="345">Phanaeus vindex</emphasis>
MacLeay
</taxonomicName>
</materialsCitation>
.
</paragraph>
<paragraph id="8BFD3699FFAE745B81E43A2CFD9BF839" blockId="8.[151,1437,1260,2043]" pageId="8" pageNumber="345">
<emphasis id="B936EA8BFFAE745B81E43A2CFE59F861" bold="true" box="[199,391,1908,1934]" pageId="8" pageNumber="345">Type deposition</emphasis>
.
<materialsCitation id="3B2A3CC4FFAE745B80B43A2CFE25F85D" country="Russia" location="Zoological Institute" pageId="8" pageNumber="345" specimenCount="2" stateProvince="Saint Petersburg" typeStatus="holotype">
The
<typeStatus id="54F9883BFFAE745B80EE3A2CFDF1F861" box="[461,559,1908,1934]" pageId="8" pageNumber="345" type="holotype">holotype</typeStatus>
and
<specimenCount id="9D44FD10FFAE745B83483A2DFCD4F860" box="[619,778,1909,1935]" count="2" pageId="8" pageNumber="345" type="generic" typeStatus="paratype">two paratypes</specimenCount>
are deposited in the collection of
<location id="8E9D6042FFAE745B85BF3A2CFAA5F861" LSID="urn:lsid:plazi:treatment:03EB878FFFA3745A81B43F1EFC4FFE8A:8E9D6042FFAE745B85BF3A2CFAA5F861" box="[1180,1403,1908,1934]" country="Russia" name="Zoological Institute" pageId="8" pageNumber="345" stateProvince="Saint Petersburg">Zoological Institute</location>
of RAS,
<collectingRegion id="4986F87BFFAE745B81FC3AC0FE40F85D" box="[223,414,1944,1970]" country="Russia" name="Saint Petersburg" pageId="8" pageNumber="345">Saint Petersburg</collectingRegion>
,
<collectingCountry id="F3557609FFAE745B808E3AC0FE25F85D" box="[429,507,1944,1970]" name="Russia" pageId="8" pageNumber="345">Russia</collectingCountry>
</materialsCitation>
;
<materialsCitation id="3B2A3CC4FFAE745B83293AC0FAC5F85D" box="[522,1307,1944,1971]" country="Russia" location="Tyumen State" pageId="8" pageNumber="345" specimenCount="1" stateProvince="Tyumen'" typeStatus="paratype">
other
<typeStatus id="54F9883BFFAE745B836D3AC1FD67F85C" box="[590,697,1945,1971]" pageId="8" pageNumber="345" type="paratype">paratypes</typeStatus>
are deposited in the collection of the
<collectingRegion id="4986F87BFFAE745B855E3AC1FAC5F85D" box="[1149,1307,1944,1971]" country="Russia" name="Tyumen'" pageId="8" pageNumber="345">Tyumen State</collectingRegion>
</materialsCitation>
<materialsCitation id="3B2A3CC4FFAE745B84053AC0FD9EF839" country="Russia" location="University Museum of Zoology" pageId="8" pageNumber="345" specimenCount="1" stateProvince="Tyumen'" typeStatus="paratype">
<location id="8E9D6042FFAE745B84053AC0FEA1F839" LSID="urn:lsid:plazi:treatment:03EB878FFFA3745A81B43F1EFC4FFE8A:8E9D6042FFAE745B84053AC0FEA1F839" country="Russia" name="University Museum of Zoology" pageId="8" pageNumber="345" stateProvince="Tyumen'">University Museum of Zoology</location>
,
<collectingRegion id="4986F87BFFAE745B80AA3AE5FE39F838" box="[393,487,1981,2007]" country="Russia" name="Tyumen'" pageId="8" pageNumber="345">Tyumen</collectingRegion>
,
<collectingCountry id="F3557609FFAE745B80D13AE4FD9EF839" box="[498,576,1980,2006]" name="Russia" pageId="8" pageNumber="345">Russia</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3586512FFAE745A81E43AB8FEB4FEF2" lastPageId="9" lastPageNumber="346" pageId="8" pageNumber="345" type="diagnosis">
<paragraph id="8BFD3699FFAE745A81E43AB8FEB4FEF2" blockId="8.[151,1437,1260,2043]" lastBlockId="9.[151,1437,151,357]" lastPageId="9" lastPageNumber="346" pageId="8" pageNumber="345">
<emphasis id="B936EA8BFFAE745B81E43AB8FE16F815" bold="true" box="[199,456,2016,2042]" pageId="8" pageNumber="345">Differential diagnosis</emphasis>
. The new species is very similar to
<taxonomicName id="4C424D1AFFAE745B824B3AB8FBD0F815" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[872,1038,2016,2043]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="345" phylum="Arthropoda" rank="species" species="brevisetosus" status="sp. nov.">
<emphasis id="B936EA8BFFAE745B824B3AB8FCA8F815" box="[872,886,2016,2042]" italics="true" pageId="8" pageNumber="345">S</emphasis>
.
<emphasis id="B936EA8BFFAE745B82A63AB8FBD0F815" box="[901,1038,2016,2042]" italics="true" pageId="8" pageNumber="345">brevisetosus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFAE745B85343AB8FBADF815" bold="true" box="[1047,1139,2016,2042]" pageId="8" pageNumber="345">
<taxonomicNameLabel id="A20557F0FFAE745B85343AB8FBADF815" box="[1047,1139,2016,2042]" pageId="8" pageNumber="345" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in having spiniform setae
<emphasis id="B936EA8BFFAF745A81B43DCFFF34FF5E" box="[151,234,151,177]" italics="true" pageId="9" pageNumber="346">pl”, tc</emphasis>
on tarsus II and
<emphasis id="B936EA8BFFAF745A808F3DCFFE0FFF5E" box="[428,465,151,177]" italics="true" pageId="9" pageNumber="346">pl”</emphasis>
on tarsus III and well separated bases of setae
<emphasis id="B936EA8BFFAF745A82CF3DCFFC26FF5F" box="[1004,1016,151,176]" italics="true" pageId="9" pageNumber="346">e</emphasis>
and
<emphasis id="B936EA8BFFAF745A85113DCFFBE4FF5E" box="[1074,1082,151,177]" italics="true" pageId="9" pageNumber="346">f</emphasis>
as well as
<emphasis id="B936EA8BFFAF745A85993DCFFB0BFF5E" box="[1210,1237,151,177]" italics="true" pageId="9" pageNumber="346">h1</emphasis>
and
<emphasis id="B936EA8BFFAF745A842C3DCFFAF5FF5E" box="[1295,1323,151,177]" italics="true" pageId="9" pageNumber="346">h2</emphasis>
. The new species differs from
<taxonomicName id="4C424D1AFFAF745A80593DE3FDC0FF3A" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[378,542,187,213]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="346" phylum="Arthropoda" rank="species" species="brevisetosus">
<emphasis id="B936EA8BFFAF745A80593DE3FE56FF3A" box="[378,392,187,213]" italics="true" pageId="9" pageNumber="346">S</emphasis>
.
<emphasis id="B936EA8BFFAF745A80B63DE3FDC0FF3A" box="[405,542,187,213]" italics="true" pageId="9" pageNumber="346">brevisetosus</emphasis>
</taxonomicName>
in having distinctly longer dorsal idiosomal setae, especially
<emphasis id="B936EA8BFFAF745A85E93DE3FB3BFF3A" box="[1226,1253,187,213]" italics="true" pageId="9" pageNumber="346">c1</emphasis>
,
<emphasis id="B936EA8BFFAF745A85D13DE3FB24FF3A" box="[1266,1274,187,213]" italics="true" pageId="9" pageNumber="346">f</emphasis>
, and
<emphasis id="B936EA8BFFAF745A84153DE3FA8FFF3A" box="[1334,1361,187,213]" italics="true" pageId="9" pageNumber="346">h1</emphasis>
which are longer than the distances between their bases (vs. setae
<emphasis id="B936EA8BFFAF745A82103D87FC90FF16" box="[819,846,223,249]" italics="true" pageId="9" pageNumber="346">c1</emphasis>
,
<emphasis id="B936EA8BFFAF745A827F3D87FCBAFF16" box="[860,868,223,249]" italics="true" pageId="9" pageNumber="346">f</emphasis>
, and
<emphasis id="B936EA8BFFAF745A82813D87FC63FF16" box="[930,957,223,249]" italics="true" pageId="9" pageNumber="346">h1</emphasis>
shorter than distances between their bases in
<taxonomicName id="4C424D1AFFAF745A81973C5BFE87FEF2" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[180,345,259,285]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="346" phylum="Arthropoda" rank="species" species="brevisetosus">
<emphasis id="B936EA8BFFAF745A81973C5BFE87FEF2" box="[180,345,259,285]" italics="true" pageId="9" pageNumber="346">S. brevisetosus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3586512FFAF745A81E43C7FFC4FFE8A" pageId="9" pageNumber="346" type="etymology">
<paragraph id="8BFD3699FFAF745A81E43C7FFC4FFE8A" blockId="9.[151,1437,151,357]" pageId="9" pageNumber="346">
<emphasis id="B936EA8BFFAF745A81E43C7FFE96FEAE" bold="true" box="[199,328,295,321]" pageId="9" pageNumber="346">Etymology</emphasis>
. The name of the new species is a combination of the Latin words:
<emphasis id="B936EA8BFFAF745A85663C7FFB4EFEAE" box="[1093,1168,295,321]" italics="true" pageId="9" pageNumber="346">longus</emphasis>
meaning
<emphasis id="B936EA8BFFAF745A85DC3C7FFAEFFEAE" box="[1279,1329,295,321]" italics="true" pageId="9" pageNumber="346">long</emphasis>
, and
<emphasis id="B936EA8BFFAF745A844C3C7FFA42FEAF" box="[1391,1436,295,320]" italics="true" pageId="9" pageNumber="346">seta</emphasis>
meaning
<emphasis id="B936EA8BFFAF745A81DC3C13FE99FE8A" box="[255,327,331,357]" italics="true" pageId="9" pageNumber="346">bristle</emphasis>
, and refers to relatively long dorsal idiosomal setae.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,84 @@
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<mods:title id="D579454BEFBCD9E4AA5CAD289C5A041D">Seven new species of Spatulaphorus Rack (Acari: Pygmephoridae) phoretic on scarab beetles (Coleoptera: Scarabaeidae)</mods:title>
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<mods:affiliation id="80AF0C940FADE6AE86F2FC0F4C652949">Zoological Institute of Russian Academy of Sciences, Universitetskaya Embankment 1, 199034, Saint Petersburg, Russia</mods:affiliation>
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<heading id="D0B581F5FFA4745181B43C7FFDECFEAE" bold="true" box="[151,562,295,322]" fontSize="11" level="1" pageId="2" pageNumber="339" reason="1">
<emphasis id="B936EA8BFFA4745181B43C7FFDECFEAE" bold="true" box="[151,562,295,322]" pageId="2" pageNumber="339">
Genus
<taxonomicName id="4C424D1AFFA4745181CD3C7FFDECFEAE" authority="Rack, 1993" authorityName="Rack" authorityYear="1993" box="[238,562,295,322]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="339" phylum="Arthropoda" rank="genus">
<emphasis id="B936EA8BFFA4745181CD3C7FFE7EFEAD" bold="true" box="[238,416,295,322]" italics="true" pageId="2" pageNumber="339">Spatulaphorus</emphasis>
Rack, 1993
</taxonomicName>
</emphasis>
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</paragraph>
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<typeStatus id="54F9883BFFA4745181B43C34FF17FE6B" box="[151,201,364,388]" pageId="2" pageNumber="339">Type</typeStatus>
species:
<taxonomicName id="4C424D1AFFA4745180053C34FCF4FE6B" authority="Dastych and Rack, 1993" authorityName="Dastych and Rack" authorityYear="1993" box="[294,810,364,388]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="339" phylum="Arthropoda" rank="species" species="camerikae">
<emphasis id="B936EA8BFFA4745180053C34FDF4FE6B" box="[294,554,364,388]" italics="true" pageId="2" pageNumber="339">Spatulaphorus camerikae</emphasis>
<bibRefCitation id="EFD34B68FFA4745183133C34FCF4FE6B" author="Dastych, H. &amp; Rack, G." box="[560,810,364,388]" pageId="2" pageNumber="339" pagination="265 - 84" refId="ref15517" refString="Dastych, H. &amp; Rack, G. (1993) Spatulaphorus gen. n. and three new species of pygmephorid mites associated with scarab beetles in Botswana and Vietnam (Acari: Heterostigmata). Mitteilungen Hamburgisches Zoologisches Museum und Institut, 90, 265 - 84." type="journal article" year="1993">Dastych and Rack, 1993</bibRefCitation>
</taxonomicName>
, by original designation.
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View file

@ -0,0 +1,654 @@
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<mods:title id="F5996DDF52B7598191B6A2768301566A">Seven new species of Spatulaphorus Rack (Acari: Pygmephoridae) phoretic on scarab beetles (Coleoptera: Scarabaeidae)</mods:title>
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<paragraph id="8BFD3699FFA4745181B43C8BFD96FE02" blockId="2.[151,584,467,529]" box="[151,584,467,494]" pageId="2" pageNumber="339">
<heading id="D0B581F5FFA4745181B43C8BFD96FE02" bold="true" box="[151,584,467,494]" fontSize="11" level="1" pageId="2" pageNumber="339" reason="1">
<emphasis id="B936EA8BFFA4745181B43C8BFD96FE02" bold="true" box="[151,584,467,494]" pageId="2" pageNumber="339">
<taxonomicName id="4C424D1AFFA4745181B43C8BFE3DFE01" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[151,483,467,494]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="339" phylum="Arthropoda" rank="species" species="brevisetosus" status="sp. nov.">
<emphasis id="B936EA8BFFA4745181B43C8BFE3DFE01" bold="true" box="[151,483,467,494]" italics="true" pageId="2" pageNumber="339">Spatulaphorus brevisetosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20557F0FFA4745180C93C8BFD96FE02" box="[490,584,467,493]" pageId="2" pageNumber="339" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C3586512FFA4745781B43CAFFE4FF89E" lastPageId="4" lastPageNumber="341" pageId="2" pageNumber="339" type="description">
<paragraph id="8BFD3699FFA4745181B43CAFFED5FDFE" blockId="2.[151,584,467,529]" box="[151,267,503,529]" pageId="2" pageNumber="339">
(
<figureCitation id="13792A1CFFA4745181BC3CAFFEDDFDFE" box="[159,259,503,529]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="1.[151,250,1956,1980]" captionStartId-1="2.[151,250,1889,1913]" captionStartId-2="3.[151,250,1616,1640]" captionStartId-3="4.[151,250,1773,1797]" captionTargetBox-0="[151,1435,639,1933]" captionTargetBox-1="[151,1430,883,1865]" captionTargetBox-2="[151,1435,388,1593]" captionTargetBox-3="[151,1436,813,1749]" captionTargetId-0="figure-207@1.[151,1435,633,1933]" captionTargetId-1="figure-179@2.[151,1435,877,1865]" captionTargetId-2="figure-150@3.[151,1435,381,1593]" captionTargetId-3="figure-511@4.[151,1436,813,1749]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionText-0="FIGURE 1. Spatulaphorus brevisetosus sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." captionText-1="FIGURE 2. Spatulaphorus brevisetosus sp. nov., female: A—left leg I, dorsal aspect; B—left leg II, dorsal aspect." captionText-2="FIGURE 3. Spatulaphorus brevisetosus sp. nov., female: A—left leg III, dorsal aspect; B—left leg IV, dorsal aspect." captionText-3="FIGURE 4. DIC micrographs of Spatulaphorus brevisetosus sp. nov., female: A—general view dorsally; B—general view ventrally." pageId="2" pageNumber="339">Figs 14</figureCitation>
)
</paragraph>
<paragraph id="8BFD3699FFA4745181B43F67FA9EFDB5" blockId="2.[151,1437,575,854]" box="[151,1344,575,602]" pageId="2" pageNumber="339">
<emphasis id="B936EA8BFFA4745181B43F67FEFFFDB6" bold="true" box="[151,289,575,601]" pageId="2" pageNumber="339">Description</emphasis>
. FEMALE. Body well sclerotized. Length of idiosoma 195 (175255), width 105 (105150).
</paragraph>
<paragraph id="8BFD3699FFA4745181E43F3BFCC9FCBA" blockId="2.[151,1437,575,854]" pageId="2" pageNumber="339">
<emphasis id="B936EA8BFFA4745181E43F3BFE88FD92" box="[199,342,611,637]" italics="true" pageId="2" pageNumber="339">Gnathosoma</emphasis>
. Gnathosomal capsule, excluding palps, subrectangular, length 21 (2123), width 21 (2124). Dorsum with one pair of needle-like cheliceral setae (
<emphasis id="B936EA8BFFA47451823D3FDFFC98FD4E" box="[798,838,647,673]" italics="true" pageId="2" pageNumber="339">cha</emphasis>
) 5 (45). Dorsal median apodeme present, poorly developed. Postpalpal setae (
<emphasis id="B936EA8BFFA4745180FC3FF4FE25FD2A" box="[479,507,684,709]" italics="true" pageId="2" pageNumber="339">pp</emphasis>
) 2 (2) needle-like, situated laterad setae
<emphasis id="B936EA8BFFA4745182EA3FF3FC2FFD2A" box="[969,1009,683,709]" italics="true" pageId="2" pageNumber="339">cha</emphasis>
. Ventral gnathosoma with one pair of smooth, weakly blunt-tipped subcapitular setae
<emphasis id="B936EA8BFFA47451838D3F88FD1CFD06" box="[686,706,720,745]" italics="true" pageId="2" pageNumber="339">m</emphasis>
5 (56) and round pits
<emphasis id="B936EA8BFFA4745182E93F88FC06FD06" box="[970,984,720,745]" italics="true" pageId="2" pageNumber="339">n</emphasis>
situated posteriad
<emphasis id="B936EA8BFFA4745185883F88FB61FD06" box="[1195,1215,720,745]" italics="true" pageId="2" pageNumber="339">m</emphasis>
. Palps with smooth and weakly blunt-tipped setae
<emphasis id="B936EA8BFFA4745180D23FABFDC3FCE2" box="[497,541,755,781]" italics="true" pageId="2" pageNumber="339">dFe</emphasis>
3 (23) and
<emphasis id="B936EA8BFFA4745183933FABFD01FCE2" box="[688,735,755,781]" italics="true" pageId="2" pageNumber="339">dGe</emphasis>
6 (6) dorsolaterally. Palps ventrally each with mushroom-like accessory setigenous structure (
<emphasis id="B936EA8BFFA4745180DA3E40FDC3FCDE" box="[505,541,792,817]" italics="true" pageId="2" pageNumber="339">ass</emphasis>
) and tiny solenidion
<emphasis id="B936EA8BFFA4745182293E40FCC0FCDE" box="[778,798,792,817]" italics="true" pageId="2" pageNumber="339">ω</emphasis>
. Palps terminated with small blunt-tipped tibial claw and tiny eupathid-like seta. Pharyngeal pumps poorly visible.
</paragraph>
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<paragraph id="8BFD3699FFA4745181B43A39FAFAF896" blockId="2.[151,1316,1889,1913]" box="[151,1316,1889,1913]" pageId="2" pageNumber="339">
<emphasis id="B936EA8BFFA4745181B43A39FED3F896" bold="true" box="[151,269,1889,1913]" pageId="2" pageNumber="339">FIGURE 2</emphasis>
.
<taxonomicName id="4C424D1AFFA47451803A3A39FDEEF896" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[281,560,1889,1913]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="339" phylum="Arthropoda" rank="species" species="brevisetosus" status="sp. nov.">
<emphasis id="B936EA8BFFA47451803A3A39FDEEF896" box="[281,560,1889,1913]" italics="true" pageId="2" pageNumber="339">Spatulaphorus brevisetosus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFA4745183153A39FD59F896" bold="true" box="[566,647,1889,1913]" pageId="2" pageNumber="339">
<taxonomicNameLabel id="A20557F0FFA4745183153A39FD59F896" box="[566,647,1889,1913]" pageId="2" pageNumber="339" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—left leg I, dorsal aspect; B—left leg II, dorsal aspect.
</paragraph>
</caption>
<paragraph id="8BFD3699FFA4745181E43AFFFA42F82D" blockId="2.[151,1437,1959,2022]" box="[199,1436,1959,1986]" pageId="2" pageNumber="339">
<emphasis id="B936EA8BFFA4745181E43AFFFE4DF82E" box="[199,403,1959,1985]" italics="true" pageId="2" pageNumber="339">Idiosomal dorsum</emphasis>
(
<figureCitation id="13792A1CFFA4745180813AFFFDDEF82E" box="[418,512,1959,1985]" captionStart="FIGURE 1" captionStartId="1.[151,250,1956,1980]" captionTargetBox="[151,1435,639,1933]" captionTargetId="figure-207@1.[151,1435,633,1933]" captionTargetPageId="1" captionText="FIGURE 1. Spatulaphorus brevisetosus sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." pageId="2" pageNumber="339">Figs 1A</figureCitation>
,
<figureCitation id="13792A1CFFA47451832E3AFFFDEDF82E" box="[525,563,1959,1985]" captionStart="FIGURE 4" captionStartId="4.[151,250,1773,1797]" captionTargetBox="[151,1436,813,1749]" captionTargetId="figure-511@4.[151,1436,813,1749]" captionTargetPageId="4" captionText="FIGURE 4. DIC micrographs of Spatulaphorus brevisetosus sp. nov., female: A—general view dorsally; B—general view ventrally." pageId="2" pageNumber="339">4A</figureCitation>
). All dorsal shields with big puncta. Prodorsal shield with three pairs of setae
</paragraph>
<paragraph id="8BFD3699FFA4745081B43A93FADCFE8A" blockId="2.[151,1437,1959,2022]" lastBlockId="3.[151,1437,151,357]" lastPageId="3" lastPageNumber="340" pageId="2" pageNumber="339">
(
<emphasis id="B936EA8BFFA4745181823A93FF64F80A" box="[161,186,1995,2021]" italics="true" pageId="2" pageNumber="339">v1</emphasis>
,
<emphasis id="B936EA8BFFA4745181EE3A93FF36F80A" box="[205,232,1995,2021]" italics="true" pageId="2" pageNumber="339">v2</emphasis>
,
<emphasis id="B936EA8BFFA4745181D93A93FEC1F80A" box="[250,287,1995,2021]" italics="true" pageId="2" pageNumber="339">sc2</emphasis>
), one pair of weakly barbed capitate trichobothria (
<emphasis id="B936EA8BFFA4745182A23A93FC79F80A" box="[897,935,1995,2021]" italics="true" pageId="2" pageNumber="339">sc1</emphasis>
) and one pair of round stigmata. Stigmata associated with well-developed tracheal trunks. All dorsal idiosomal setae blunt-tipped; setae
<emphasis id="B936EA8BFFA5745085F23DCFFB35FF5E" box="[1233,1259,151,177]" italics="true" pageId="3" pageNumber="340">v2</emphasis>
,
<emphasis id="B936EA8BFFA5745085DF3DCFFAD6FF5F" box="[1276,1288,151,176]" italics="true" pageId="3" pageNumber="340">e</emphasis>
and
<emphasis id="B936EA8BFFA5745084663DCFFABFFF5E" box="[1349,1377,151,177]" italics="true" pageId="3" pageNumber="340">h2</emphasis>
with few minute barbs, other dorsal setae distinctly barbed. Alveoli of setae
<emphasis id="B936EA8BFFA5745082F23DE3FC03FF3B" box="[977,989,187,212]" italics="true" pageId="3" pageNumber="340">e</emphasis>
and
<emphasis id="B936EA8BFFA57450853A3DE3FBFFFF3A" box="[1049,1057,187,213]" italics="true" pageId="3" pageNumber="340">f</emphasis>
clearly separated. Cupules
<emphasis id="B936EA8BFFA5745084423DE3FAA9FF3B" box="[1377,1399,187,212]" italics="true" pageId="3" pageNumber="340">ia</emphasis>
on tergite D,
<emphasis id="B936EA8BFFA57450802B3D87FEFAFF17" box="[264,292,223,248]" italics="true" pageId="3" pageNumber="340">im</emphasis>
on tergite EF and
<emphasis id="B936EA8BFFA5745080D03D87FDD7FF16" box="[499,521,223,249]" italics="true" pageId="3" pageNumber="340">ih</emphasis>
on tergite H large, round, subequal. Lengths of dorsal setae:
<emphasis id="B936EA8BFFA5745085963D87FB11FF16" box="[1205,1231,223,249]" italics="true" pageId="3" pageNumber="340">v1</emphasis>
20 (2025),
<emphasis id="B936EA8BFFA5745084433D87FAA4FF16" box="[1376,1402,223,249]" italics="true" pageId="3" pageNumber="340">v2</emphasis>
11 (1113),
<emphasis id="B936EA8BFFA5745081DF3C5BFEFCFEF2" box="[252,290,259,285]" italics="true" pageId="3" pageNumber="340">sc2</emphasis>
29 (2938),
<emphasis id="B936EA8BFFA5745080913C5BFE12FEF2" box="[434,460,259,285]" italics="true" pageId="3" pageNumber="340">c1</emphasis>
26 (2633),
<emphasis id="B936EA8BFFA57450837A3C5BFDADFEF2" box="[601,627,259,285]" italics="true" pageId="3" pageNumber="340">c2</emphasis>
29 (2932),
<emphasis id="B936EA8BFFA5745082203C5BFCCFFEF2" box="[771,785,259,285]" italics="true" pageId="3" pageNumber="340">d</emphasis>
28 (2833),
<emphasis id="B936EA8BFFA5745082823C5BFC73FEF3" box="[929,941,259,284]" italics="true" pageId="3" pageNumber="340">e</emphasis>
15 (1519),
<emphasis id="B936EA8BFFA57450851E3C5BFB9BFEF2" box="[1085,1093,259,285]" italics="true" pageId="3" pageNumber="340">f</emphasis>
28 (2835),
<emphasis id="B936EA8BFFA5745085F73C5BFB2EFEF2" box="[1236,1264,259,285]" italics="true" pageId="3" pageNumber="340">h1</emphasis>
31 (3140),
<emphasis id="B936EA8BFFA5745084A33C5BFA43FEF2" box="[1408,1437,259,285]" italics="true" pageId="3" pageNumber="340">h2</emphasis>
11 (1115). Distances between setae:
<emphasis id="B936EA8BFFA57450831D3C7FFD5FFEAE" box="[574,641,295,321]" italics="true" pageId="3" pageNumber="340">v1v1</emphasis>
20 (2024),
<emphasis id="B936EA8BFFA5745082313C7FFC8BFEAE" box="[786,853,295,321]" italics="true" pageId="3" pageNumber="340">v2v2</emphasis>
30 (3035),
<emphasis id="B936EA8BFFA5745082C53C7FFBE0FEAE" box="[998,1086,295,321]" italics="true" pageId="3" pageNumber="340">sc2sc2</emphasis>
32 (3238),
<emphasis id="B936EA8BFFA5745085F33C7FFACDFEAE" box="[1232,1299,295,321]" italics="true" pageId="3" pageNumber="340">c1c1</emphasis>
34 (3442),
<emphasis id="B936EA8BFFA5745081B43C13FF04FE8A" box="[151,218,331,357]" italics="true" pageId="3" pageNumber="340">c1c2</emphasis>
19 (1922),
<emphasis id="B936EA8BFFA5745080483C13FE4BFE8A" box="[363,405,331,357]" italics="true" pageId="3" pageNumber="340">dd</emphasis>
44 (4454),
<emphasis id="B936EA8BFFA5745083063C13FD96FE8A" box="[549,584,331,357]" italics="true" pageId="3" pageNumber="340">ef</emphasis>
7 (78),
<emphasis id="B936EA8BFFA57450838D3C13FD12FE8A" box="[686,716,331,357]" italics="true" pageId="3" pageNumber="340">ff</emphasis>
39 (3947),
<emphasis id="B936EA8BFFA57450827F3C13FC7CFE8A" box="[860,930,331,357]" italics="true" pageId="3" pageNumber="340">h1h1</emphasis>
32 (3240),
<emphasis id="B936EA8BFFA5745085103C13FBA7FE8A" box="[1075,1145,331,357]" italics="true" pageId="3" pageNumber="340">h1h2</emphasis>
10 (1011).
</paragraph>
<caption id="DF3D6611FFA5745081B43B08FAE5F988" box="[151,1339,1616,1640]" pageId="3" pageNumber="340" startId="3.[151,250,1616,1640]" targetBox="[151,1435,388,1593]" targetPageId="3" targetType="figure">
<paragraph id="8BFD3699FFA5745081B43B08FAE5F988" blockId="3.[151,1339,1616,1640]" box="[151,1339,1616,1640]" pageId="3" pageNumber="340">
<emphasis id="B936EA8BFFA5745081B43B08FED3F987" bold="true" box="[151,269,1616,1640]" pageId="3" pageNumber="340">FIGURE 3</emphasis>
.
<taxonomicName id="4C424D1AFFA57450803A3B08FDEEF987" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[281,560,1616,1640]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="340" phylum="Arthropoda" rank="species" species="brevisetosus" status="sp. nov.">
<emphasis id="B936EA8BFFA57450803A3B08FDEEF987" box="[281,560,1616,1640]" italics="true" pageId="3" pageNumber="340">Spatulaphorus brevisetosus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFA5745083153B08FD59F987" bold="true" box="[566,647,1616,1640]" pageId="3" pageNumber="340">
<taxonomicNameLabel id="A20557F0FFA5745083153B08FD59F987" box="[566,647,1616,1640]" pageId="3" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—left leg III, dorsal aspect; B—left leg IV, dorsal aspect.
</paragraph>
</caption>
<paragraph id="8BFD3699FFA5745081E43BCEFD6DF81B" blockId="3.[151,1437,1686,2036]" pageId="3" pageNumber="340">
<emphasis id="B936EA8BFFA5745081E43BCEFE58F95F" box="[199,390,1686,1712]" italics="true" pageId="3" pageNumber="340">Idiosomal venter</emphasis>
(
<figureCitation id="13792A1CFFA5745080B43BCEFE2AF95F" box="[407,500,1686,1712]" captionStart="FIGURE 1" captionStartId="1.[151,250,1956,1980]" captionTargetBox="[151,1435,639,1933]" captionTargetId="figure-207@1.[151,1435,633,1933]" captionTargetPageId="1" captionText="FIGURE 1. Spatulaphorus brevisetosus sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." pageId="3" pageNumber="340">Figs 1B</figureCitation>
,
<figureCitation id="13792A1CFFA5745083213BCEFDF8F95F" box="[514,550,1686,1712]" captionStart="FIGURE 4" captionStartId="4.[151,250,1773,1797]" captionTargetBox="[151,1436,813,1749]" captionTargetId="figure-511@4.[151,1436,813,1749]" captionTargetPageId="4" captionText="FIGURE 4. DIC micrographs of Spatulaphorus brevisetosus sp. nov., female: A—general view dorsally; B—general view ventrally." pageId="3" pageNumber="340">4B</figureCitation>
). Coxisternal fields III and aggenital plate laterally with sparsely distributed big puncta. Setae
<emphasis id="B936EA8BFFA5745080403BE2FE5EF93B" box="[355,384,1722,1748]" italics="true" pageId="3" pageNumber="340">1b</emphasis>
pointed and weakly barbed; other ventral setae smooth and blunt-tipped. Bases of setae
<emphasis id="B936EA8BFFA5745084563BE2FA42F93B" box="[1397,1436,1722,1748]" italics="true" pageId="3" pageNumber="340">ps3</emphasis>
situated distinctly anteriad bases of setae
<emphasis id="B936EA8BFFA5745083433B86FD78F917" box="[608,678,1758,1784]" italics="true" pageId="3" pageNumber="340">ps1-2.</emphasis>
Apodemes 1 (ap1) and apodemes 2 (ap2) well-developed and joined with thick prosternal apodeme (appr); sejugal apodeme (apsej) developed only laterally; apodemes 3 (ap3) weak, not reaching poststernal apodeme (appo); apodemes 4 (ap4) well developed, exceeding beyond bases of setae
<emphasis id="B936EA8BFFA57450845A3A7EFA4BF8AF" box="[1401,1429,1830,1856]" italics="true" pageId="3" pageNumber="340">3b</emphasis>
. Apodemes 5 (ap5) short, joined with appo and reaching bases of setae
<emphasis id="B936EA8BFFA5745082893A12FC18F88B" box="[938,966,1866,1892]" italics="true" pageId="3" pageNumber="340">4a</emphasis>
. Anterior genital sclerite (ags) small, cup-like; posterior genital sclerite (pgs) small, oval. Posterior margin of posterior sternal plate weakly concave in middle part. Posterior margin of aggenital plate rounded. Lengths of ventral setae:
<emphasis id="B936EA8BFFA5745082F83ACAFC29F843" box="[987,1015,1938,1964]" italics="true" pageId="3" pageNumber="340">1a</emphasis>
9 (910),
<emphasis id="B936EA8BFFA5745085493ACAFB59F843" box="[1130,1159,1938,1964]" italics="true" pageId="3" pageNumber="340">1b</emphasis>
16 (1618),
<emphasis id="B936EA8BFFA5745084363ACAFAEEF843" box="[1301,1328,1938,1964]" italics="true" pageId="3" pageNumber="340">1с</emphasis>
9 (912),
<emphasis id="B936EA8BFFA5745081B43AEEFF6DF83F" box="[151,179,1974,2000]" italics="true" pageId="3" pageNumber="340">2a</emphasis>
12 (1213),
<emphasis id="B936EA8BFFA5745080673AEEFE80F83F" box="[324,350,1974,2000]" italics="true" pageId="3" pageNumber="340">2с</emphasis>
9 (911),
<emphasis id="B936EA8BFFA5745080F03AEEFE30F83F" box="[467,494,1974,2000]" italics="true" pageId="3" pageNumber="340">3a</emphasis>
10 (1013),
<emphasis id="B936EA8BFFA5745083A33AEEFD42F83F" box="[640,668,1974,2000]" italics="true" pageId="3" pageNumber="340">3b</emphasis>
9 (912),
<emphasis id="B936EA8BFFA5745082323AEEFCF5F83F" box="[785,811,1974,2000]" italics="true" pageId="3" pageNumber="340">3c</emphasis>
11 (1114),
<emphasis id="B936EA8BFFA5745082993AEEFC08F83F" box="[954,982,1974,2000]" italics="true" pageId="3" pageNumber="340">4a</emphasis>
9 (912),
<emphasis id="B936EA8BFFA5745085683AEEFBB9F83F" box="[1099,1127,1974,2000]" italics="true" pageId="3" pageNumber="340">4b</emphasis>
14 (1419),
<emphasis id="B936EA8BFFA5745085DB3AEEFACCF83F" box="[1272,1298,1974,2000]" italics="true" pageId="3" pageNumber="340">4c</emphasis>
10 (1013),
<emphasis id="B936EA8BFFA5745081B43A82FF60F81B" box="[151,190,2010,2036]" italics="true" pageId="3" pageNumber="340">ps1</emphasis>
11 (1113),
<emphasis id="B936EA8BFFA57450806E3A82FEAAF81B" box="[333,372,2010,2036]" italics="true" pageId="3" pageNumber="340">ps2</emphasis>
12 (1214),
<emphasis id="B936EA8BFFA5745083273A82FDF5F81B" box="[516,555,2010,2036]" italics="true" pageId="3" pageNumber="340">ps3</emphasis>
11 (1112).
</paragraph>
<paragraph id="8BFD3699FFA2745781E43DCFFCC5FCFA" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="341">
<emphasis id="B936EA8BFFA2745781E43DCFFF22FF5F" box="[199,252,151,176]" italics="true" pageId="4" pageNumber="341">Legs</emphasis>
(
<figureCitation id="13792A1CFFA2745780283DCFFE8AFF5E" box="[267,340,151,177]" captionStart="FIGURE 2" captionStartId="2.[151,250,1889,1913]" captionTargetBox="[151,1430,883,1865]" captionTargetId="figure-179@2.[151,1435,877,1865]" captionTargetPageId="2" captionText="FIGURE 2. Spatulaphorus brevisetosus sp. nov., female: A—left leg I, dorsal aspect; B—left leg II, dorsal aspect." pageId="4" pageNumber="341">Figs 2</figureCitation>
,
<figureCitation id="13792A1CFFA2745780433DCFFEAEFF5E" box="[352,368,151,177]" captionStart="FIGURE 3" captionStartId="3.[151,250,1616,1640]" captionTargetBox="[151,1435,388,1593]" captionTargetId="figure-150@3.[151,1435,381,1593]" captionTargetPageId="3" captionText="FIGURE 3. Spatulaphorus brevisetosus sp. nov., female: A—left leg III, dorsal aspect; B—left leg IV, dorsal aspect." pageId="4" pageNumber="341">3</figureCitation>
). Leg I (
<figureCitation id="13792A1CFFA2745780F13DCFFDF2FF5E" box="[466,556,151,177]" captionStart="FIGURE 2" captionStartId="2.[151,250,1889,1913]" captionTargetBox="[151,1430,883,1865]" captionTargetId="figure-179@2.[151,1435,877,1865]" captionTargetPageId="2" captionText="FIGURE 2. Spatulaphorus brevisetosus sp. nov., female: A—left leg I, dorsal aspect; B—left leg II, dorsal aspect." pageId="4" pageNumber="341">Fig. 2A</figureCitation>
). Leg setation: Tr 1 (v), Fe 4 (
<emphasis id="B936EA8BFFA2745782AB3DCFFC48FF5E" box="[904,918,151,177]" italics="true" pageId="4" pageNumber="341">d</emphasis>
,
<emphasis id="B936EA8BFFA2745782863DCFFC68FF5E" box="[933,950,151,177]" italics="true" pageId="4" pageNumber="341">l</emphasis>
,
<emphasis id="B936EA8BFFA2745782E73DCFFC05FF5E" box="[964,987,151,177]" italics="true" pageId="4" pageNumber="341">l”</emphasis>
,
<emphasis id="B936EA8BFFA2745782CA3DCFFBDBFF5E" box="[1001,1029,151,177]" italics="true" pageId="4" pageNumber="341">v”</emphasis>
), Ge 2 (
<emphasis id="B936EA8BFFA2745785473DCFFBABFF5E" box="[1124,1141,151,177]" italics="true" pageId="4" pageNumber="341">l</emphasis>
,
<emphasis id="B936EA8BFFA2745785A03DCFFB44FF5E" box="[1155,1178,151,177]" italics="true" pageId="4" pageNumber="341">l”</emphasis>
), TiTa 17(4) (
<emphasis id="B936EA8BFFA27457841A3DCFFA83FF5E" box="[1337,1373,151,177]" italics="true" pageId="4" pageNumber="341">pϛ</emphasis>
,
<emphasis id="B936EA8BFFA2745784483DCFFA4BFF5E" box="[1387,1429,151,177]" italics="true" pageId="4" pageNumber="341">p”ϛ</emphasis>
,
<emphasis id="B936EA8BFFA2745781B43DE3FF1FFF3A" box="[151,193,187,213]" italics="true" pageId="4" pageNumber="341">tcϛ</emphasis>
,
<emphasis id="B936EA8BFFA2745781F33DE3FEDFFF3A" box="[208,257,187,213]" italics="true" pageId="4" pageNumber="341">tc”ϛ</emphasis>
,
<emphasis id="B936EA8BFFA27457802C3DE3FEEAFF3A" box="[271,308,187,213]" italics="true" pageId="4" pageNumber="341">ftϛ</emphasis>
,
<emphasis id="B936EA8BFFA2745780603DE3FEB1FF3A" box="[323,367,187,213]" italics="true" pageId="4" pageNumber="341">ft”ϛ</emphasis>
,
<emphasis id="B936EA8BFFA27457805E3DE3FE57FF3A" box="[381,393,187,213]" italics="true" pageId="4" pageNumber="341">k</emphasis>
,
<emphasis id="B936EA8BFFA2745780BB3DE3FE69FF3A" box="[408,439,187,213]" italics="true" pageId="4" pageNumber="341">pl</emphasis>
,
<emphasis id="B936EA8BFFA2745780E53DE3FE32FF3A" box="[454,492,187,213]" italics="true" pageId="4" pageNumber="341">pl”</emphasis>
,
<emphasis id="B936EA8BFFA2745780D93DE3FDC0FF3A" box="[506,542,187,213]" italics="true" pageId="4" pageNumber="341">pv</emphasis>
,
<emphasis id="B936EA8BFFA27457830E3DE3FD89FF3A" box="[557,599,187,213]" italics="true" pageId="4" pageNumber="341">pv”</emphasis>
,
<emphasis id="B936EA8BFFA2745783453DE3FDAFFF3B" box="[614,625,187,212]" italics="true" pageId="4" pageNumber="341">s</emphasis>
,
<emphasis id="B936EA8BFFA27457835C3DE3FD53FF3A" box="[639,653,187,213]" italics="true" pageId="4" pageNumber="341">d</emphasis>
,
<emphasis id="B936EA8BFFA2745783BF3DE3FD73FF3A" box="[668,685,187,213]" italics="true" pageId="4" pageNumber="341">l</emphasis>
,
<emphasis id="B936EA8BFFA27457839F3DE3FD0DFF3A" box="[700,723,187,213]" italics="true" pageId="4" pageNumber="341">l”</emphasis>
,
<emphasis id="B936EA8BFFA2745783C13DE3FD26FF3A" box="[738,760,187,213]" italics="true" pageId="4" pageNumber="341">v</emphasis>
,
<emphasis id="B936EA8BFFA2745782253DE3FCFCFF3A" box="[774,802,187,213]" italics="true" pageId="4" pageNumber="341">v”</emphasis>
,
<emphasis id="B936EA8BFFA2745782123DE3FC8DFF3A" box="[817,851,187,213]" italics="true" pageId="4" pageNumber="341">ω1</emphasis>
,
<emphasis id="B936EA8BFFA2745782413DE3FC5AFF3A" box="[866,900,187,213]" italics="true" pageId="4" pageNumber="341">ω2</emphasis>
,
<emphasis id="B936EA8BFFA2745782B13DE3FC71FF3A" box="[914,943,187,213]" italics="true" pageId="4" pageNumber="341">φ1</emphasis>
,
<emphasis id="B936EA8BFFA27457829D3DE3FC02FF3A" box="[958,988,187,213]" italics="true" pageId="4" pageNumber="341">φ2</emphasis>
). Tibiotarsus enlarged, with large claw; tibiotarsus internally with oval strongly sclerotized structure near solenidion
<emphasis id="B936EA8BFFA2745782D23D87FBD0FF16" box="[1009,1038,223,249]" italics="true" pageId="4" pageNumber="341">φ2</emphasis>
. Setae (
<emphasis id="B936EA8BFFA27457854B3D87FBA8FF17" box="[1128,1142,223,248]" italics="true" pageId="4" pageNumber="341">u</emphasis>
) consolidated into a horn-like structure opposing claw. Tarsal claw blunt-tipped. Seta
<emphasis id="B936EA8BFFA2745782103C5BFC9FFEF2" box="[819,833,259,285]" italics="true" pageId="4" pageNumber="341">d</emphasis>
of femur spatulate, with short subterminal projection. Setae
<emphasis id="B936EA8BFFA2745781F93C7FFF35FEAE" box="[218,235,295,321]" italics="true" pageId="4" pageNumber="341">l</emphasis>
of femur,
<emphasis id="B936EA8BFFA2745780433C7FFEB2FEAE" box="[352,364,295,321]" italics="true" pageId="4" pageNumber="341">k</emphasis>
and
<emphasis id="B936EA8BFFA2745780813C7FFE1FFEAE" box="[418,449,295,321]" italics="true" pageId="4" pageNumber="341">pl</emphasis>
of tibiotarsus smooth; other leg setae (except eupathidia
<emphasis id="B936EA8BFFA27457851D3C7FFB8BFEAE" box="[1086,1109,295,321]" italics="true" pageId="4" pageNumber="341">p</emphasis>
,
<emphasis id="B936EA8BFFA2745785403C7FFB5EFEAE" box="[1123,1152,295,321]" italics="true" pageId="4" pageNumber="341">p”</emphasis>
,
<emphasis id="B936EA8BFFA2745785AD3C7FFB72FEAE" box="[1166,1196,295,321]" italics="true" pageId="4" pageNumber="341">tc</emphasis>
,
<emphasis id="B936EA8BFFA27457859A3C7FFB03FEAE" box="[1209,1245,295,321]" italics="true" pageId="4" pageNumber="341">tc”</emphasis>
,
<emphasis id="B936EA8BFFA2745785C83C7FFADAFEAE" box="[1259,1284,295,321]" italics="true" pageId="4" pageNumber="341">ft</emphasis>
,
<emphasis id="B936EA8BFFA2745784323C7FFAEEFEAE" box="[1297,1328,295,321]" italics="true" pageId="4" pageNumber="341">ft”</emphasis>
) sparsely barbed. Setae
<emphasis id="B936EA8BFFA2745780103C13FE9AFE8A" box="[307,324,331,357]" italics="true" pageId="4" pageNumber="341">l</emphasis>
,
<emphasis id="B936EA8BFFA2745780723C13FEB6FE8A" box="[337,360,331,357]" italics="true" pageId="4" pageNumber="341">l”</emphasis>
of femur,
<emphasis id="B936EA8BFFA2745780F93C13FE2FFE8A" box="[474,497,331,357]" italics="true" pageId="4" pageNumber="341">l”</emphasis>
of genu and
<emphasis id="B936EA8BFFA27457835C3C13FD55FE8A" box="[639,651,331,357]" italics="true" pageId="4" pageNumber="341">k</emphasis>
of tibiotarsus blunt-tipped; other leg setae (except eupathidia) pointed. Lengths of solenidia
<emphasis id="B936EA8BFFA2745780A53C37FE79FE66" box="[390,423,367,393]" italics="true" pageId="4" pageNumber="341">ω1</emphasis>
8 (89),
<emphasis id="B936EA8BFFA2745783313C37FDEDFE66" box="[530,563,367,393]" italics="true" pageId="4" pageNumber="341">ω2</emphasis>
4 (45),
<emphasis id="B936EA8BFFA2745783BE3C37FD64FE66" box="[669,698,367,393]" italics="true" pageId="4" pageNumber="341">φ1</emphasis>
7 (78),
<emphasis id="B936EA8BFFA2745782063C37FC9CFE66" box="[805,834,367,393]" italics="true" pageId="4" pageNumber="341">φ2</emphasis>
4 (45); Solenidion
<emphasis id="B936EA8BFFA27457850D3C37FB95FE66" box="[1070,1099,367,393]" italics="true" pageId="4" pageNumber="341">φ1</emphasis>
thick, clavate,
<emphasis id="B936EA8BFFA2745785DA3C37FAC5FE66" box="[1273,1307,367,393]" italics="true" pageId="4" pageNumber="341">ω1</emphasis>
digitiform; other solenidia weakly clavate. Leg II (
<figureCitation id="13792A1CFFA27457836F3CCBFD7AFE42" box="[588,676,403,429]" captionStart="FIGURE 2" captionStartId="2.[151,250,1889,1913]" captionTargetBox="[151,1430,883,1865]" captionTargetId="figure-179@2.[151,1435,877,1865]" captionTargetPageId="2" captionText="FIGURE 2. Spatulaphorus brevisetosus sp. nov., female: A—left leg I, dorsal aspect; B—left leg II, dorsal aspect." pageId="4" pageNumber="341">Fig. 2B</figureCitation>
). Leg setation: Tr 1 (
<emphasis id="B936EA8BFFA2745782AE3CCBFC7DFE42" box="[909,931,403,429]" italics="true" pageId="4" pageNumber="341">v</emphasis>
), Fe 3 (
<emphasis id="B936EA8BFFA2745782D83CCBFBD7FE42" box="[1019,1033,403,429]" italics="true" pageId="4" pageNumber="341">d</emphasis>
,
<emphasis id="B936EA8BFFA2745785353CCBFBF9FE42" box="[1046,1063,403,429]" italics="true" pageId="4" pageNumber="341">l</emphasis>
,
<emphasis id="B936EA8BFFA2745785163CCBFB8FFE42" box="[1077,1105,403,429]" italics="true" pageId="4" pageNumber="341">v”</emphasis>
), Ge 1 (
<emphasis id="B936EA8BFFA27457858D3CCBFB61FE42" box="[1198,1215,403,429]" italics="true" pageId="4" pageNumber="341">l</emphasis>
), Ti 4 (
<emphasis id="B936EA8BFFA2745784313CCBFAFEFE42" box="[1298,1312,403,429]" italics="true" pageId="4" pageNumber="341">d</emphasis>
,
<emphasis id="B936EA8BFFA27457840D3CCBFAE1FE42" box="[1326,1343,403,429]" italics="true" pageId="4" pageNumber="341">l</emphasis>
,
<emphasis id="B936EA8BFFA27457846F3CCBFABCFE42" box="[1356,1378,403,429]" italics="true" pageId="4" pageNumber="341">v</emphasis>
,
<emphasis id="B936EA8BFFA2745784533CCBFA55FE42" box="[1392,1419,403,429]" italics="true" pageId="4" pageNumber="341">v”</emphasis>
), Ta 6(1) (
<emphasis id="B936EA8BFFA2745781D83CEFFEFFFE3E" box="[251,289,439,465]" italics="true" pageId="4" pageNumber="341">pl”</emphasis>
,
<emphasis id="B936EA8BFFA27457800C3CEFFE93FE3E" box="[303,333,439,465]" italics="true" pageId="4" pageNumber="341">tc</emphasis>
,
<emphasis id="B936EA8BFFA27457807F3CEFFE5EFE3E" box="[348,384,439,465]" italics="true" pageId="4" pageNumber="341">tc”</emphasis>
,
<emphasis id="B936EA8BFFA2745780AC3CEFFE78FE3E" box="[399,422,439,465]" italics="true" pageId="4" pageNumber="341">u</emphasis>
,
<emphasis id="B936EA8BFFA2745780963CEFFE07FE3E" box="[437,473,439,465]" italics="true" pageId="4" pageNumber="341">pv</emphasis>
,
<emphasis id="B936EA8BFFA2745780CB3CEFFDCCFE3E" box="[488,530,439,465]" italics="true" pageId="4" pageNumber="341">pv”</emphasis>
,
<emphasis id="B936EA8BFFA2745783023CEFFDEBFE3F" box="[545,565,439,464]" italics="true" pageId="4" pageNumber="341">ω</emphasis>
). Tarsus with thickened basally claws and pad-like empodium. Solenidion
<emphasis id="B936EA8BFFA2745784AB3CEFFA42FE3F" box="[1416,1436,439,464]" italics="true" pageId="4" pageNumber="341">ω</emphasis>
5 (5) weakly clavate, solenidion
<emphasis id="B936EA8BFFA2745783263C83FDCAFE1B" box="[517,532,475,500]" italics="true" pageId="4" pageNumber="341">φ</emphasis>
absent. Seta u
<emphasis id="B936EA8BFFA2745783953C83FD61FE1A" box="[694,703,475,501]" italics="true" pageId="4" pageNumber="341"></emphasis>
smooth, other setae barbed; setae
<emphasis id="B936EA8BFFA2745785623C83FB81FE1A" box="[1089,1119,475,501]" italics="true" pageId="4" pageNumber="341">tc</emphasis>
and
<emphasis id="B936EA8BFFA2745785B63C83FB65FE1A" box="[1173,1211,475,501]" italics="true" pageId="4" pageNumber="341">pl”</emphasis>
of tarsus spiniform, blunt-tipped; setae
<emphasis id="B936EA8BFFA2745780513CA7FE5EFDF6" box="[370,384,511,537]" italics="true" pageId="4" pageNumber="341">d</emphasis>
,
<emphasis id="B936EA8BFFA2745780B33CA7FE7FFDF6" box="[400,417,511,537]" italics="true" pageId="4" pageNumber="341">l</emphasis>
, and
<emphasis id="B936EA8BFFA2745780C03CA7FE21FDF6" box="[483,511,511,537]" italics="true" pageId="4" pageNumber="341">v”</emphasis>
of femur blunt-tipped; other leg setae pointed. Leg III (
<figureCitation id="13792A1CFFA2745785A73CA7FB3EFDF6" box="[1156,1248,511,537]" captionStart="FIGURE 3" captionStartId="3.[151,250,1616,1640]" captionTargetBox="[151,1435,388,1593]" captionTargetId="figure-150@3.[151,1435,381,1593]" captionTargetPageId="3" captionText="FIGURE 3. Spatulaphorus brevisetosus sp. nov., female: A—left leg III, dorsal aspect; B—left leg IV, dorsal aspect." pageId="4" pageNumber="341">Fig. 3A</figureCitation>
). Femur divided into basi- and telofemur. Leg setation: Tr 1 (
<emphasis id="B936EA8BFFA2745783A53F7BFD42FDD2" box="[646,668,547,573]" italics="true" pageId="4" pageNumber="341">v</emphasis>
), Fe 2 (
<emphasis id="B936EA8BFFA2745783D63F7BFCDDFDD2" box="[757,771,547,573]" italics="true" pageId="4" pageNumber="341">d</emphasis>
,
<emphasis id="B936EA8BFFA2745782323F7BFCF9FDD2" box="[785,807,547,573]" italics="true" pageId="4" pageNumber="341">v</emphasis>
), Ge 1 (
<emphasis id="B936EA8BFFA2745782A73F7BFC4BFDD2" box="[900,917,547,573]" italics="true" pageId="4" pageNumber="341">l</emphasis>
), Ti 4 (
<emphasis id="B936EA8BFFA2745782CA3F7BFC29FDD2" box="[1001,1015,547,573]" italics="true" pageId="4" pageNumber="341">d</emphasis>
,
<emphasis id="B936EA8BFFA2745785263F7BFBC8FDD2" box="[1029,1046,547,573]" italics="true" pageId="4" pageNumber="341">l</emphasis>
,
<emphasis id="B936EA8BFFA2745785073F7BFBE4FDD2" box="[1060,1082,547,573]" italics="true" pageId="4" pageNumber="341">v</emphasis>
,
<emphasis id="B936EA8BFFA27457856B3F7BFBBDFDD2" box="[1096,1123,547,573]" italics="true" pageId="4" pageNumber="341">v”</emphasis>
), Ta 6 (
<emphasis id="B936EA8BFFA27457859F3F7BFB3FFDD2" box="[1212,1249,547,573]" italics="true" pageId="4" pageNumber="341">pl”</emphasis>
,
<emphasis id="B936EA8BFFA2745785CC3F7BFAD3FDD2" box="[1263,1293,547,573]" italics="true" pageId="4" pageNumber="341">tc</emphasis>
,
<emphasis id="B936EA8BFFA2745784393F7BFAE0FDD2" box="[1306,1342,547,573]" italics="true" pageId="4" pageNumber="341">tc”</emphasis>
,
<emphasis id="B936EA8BFFA27457846F3F7BFABDFDD2" box="[1356,1379,547,573]" italics="true" pageId="4" pageNumber="341">u</emphasis>
,
<emphasis id="B936EA8BFFA2745784523F7BFA4BFDD2" box="[1393,1429,547,573]" italics="true" pageId="4" pageNumber="341">pv</emphasis>
,
<emphasis id="B936EA8BFFA2745781B43F1FFF1FFD8E" box="[151,193,583,609]" italics="true" pageId="4" pageNumber="341">pv”</emphasis>
). Claws and empodium of same shape as on tarsus II. Setae
<emphasis id="B936EA8BFFA2745782433F1FFCA8FD8E" box="[864,886,583,609]" italics="true" pageId="4" pageNumber="341">v</emphasis>
of trochanter and
<emphasis id="B936EA8BFFA2745785623F1FFB86FD8E" box="[1089,1112,583,609]" italics="true" pageId="4" pageNumber="341">u</emphasis>
of tarsus smooth, other setae barbed; seta
<emphasis id="B936EA8BFFA2745780003F33FE97FD6A" box="[291,329,619,645]" italics="true" pageId="4" pageNumber="341">pl”</emphasis>
of tarsus spiniform; setae
<emphasis id="B936EA8BFFA27457834F3F33FD5CFD6A" box="[620,642,619,645]" italics="true" pageId="4" pageNumber="341">v</emphasis>
of trochanter,
<emphasis id="B936EA8BFFA2745782003F33FCEFFD6A" box="[803,817,619,645]" italics="true" pageId="4" pageNumber="341">d</emphasis>
,
<emphasis id="B936EA8BFFA27457821D3F33FC8AFD6A" box="[830,852,619,645]" italics="true" pageId="4" pageNumber="341">v</emphasis>
of femur, and
<emphasis id="B936EA8BFFA2745782D63F33FBD8FD6A" box="[1013,1030,619,645]" italics="true" pageId="4" pageNumber="341">l</emphasis>
of genu blunt-tipped, other leg setae pointed; solenidion
<emphasis id="B936EA8BFFA27457805B3FD7FE59FD47" box="[376,391,655,680]" italics="true" pageId="4" pageNumber="341">φ</emphasis>
absent. Leg IV (
<figureCitation id="13792A1CFFA2745783653FD7FD46FD46" box="[582,664,655,681]" captionStart="FIGURE 3" captionStartId="3.[151,250,1616,1640]" captionTargetBox="[151,1435,388,1593]" captionTargetId="figure-150@3.[151,1435,381,1593]" captionTargetPageId="3" captionText="FIGURE 3. Spatulaphorus brevisetosus sp. nov., female: A—left leg III, dorsal aspect; B—left leg IV, dorsal aspect." pageId="4" pageNumber="341">Fig 3B</figureCitation>
). Femur divided into basi- and telofemur. Leg setation: Tr 1 (
<emphasis id="B936EA8BFFA2745784713FD7FAB6FD46" box="[1362,1384,655,681]" italics="true" pageId="4" pageNumber="341">v</emphasis>
), Fe 2 (
<emphasis id="B936EA8BFFA2745781953FEBFF1AFD22" box="[182,196,691,717]" italics="true" pageId="4" pageNumber="341">d</emphasis>
,
<emphasis id="B936EA8BFFA2745781F03FEBFF37FD22" box="[211,233,691,717]" italics="true" pageId="4" pageNumber="341">v</emphasis>
), Ge 0, Ti 4 (
<emphasis id="B936EA8BFFA2745780A63FEBFE4DFD22" box="[389,403,691,717]" italics="true" pageId="4" pageNumber="341">d</emphasis>
,
<emphasis id="B936EA8BFFA2745780813FEBFE6DFD22" box="[418,435,691,717]" italics="true" pageId="4" pageNumber="341">l</emphasis>
,
<emphasis id="B936EA8BFFA2745780E13FEBFE06FD22" box="[450,472,691,717]" italics="true" pageId="4" pageNumber="341">v</emphasis>
,
<emphasis id="B936EA8BFFA2745780C43FEBFDDDFD22" box="[487,515,691,717]" italics="true" pageId="4" pageNumber="341">v”</emphasis>
), Ta 6 (
<emphasis id="B936EA8BFFA27457837E3FEBFD5DFD22" box="[605,643,691,717]" italics="true" pageId="4" pageNumber="341">pl”</emphasis>
,
<emphasis id="B936EA8BFFA2745783B13FEBFD6EFD22" box="[658,688,691,717]" italics="true" pageId="4" pageNumber="341">tc</emphasis>
,
<emphasis id="B936EA8BFFA27457839D3FEBFD3CFD22" box="[702,738,691,717]" italics="true" pageId="4" pageNumber="341">tc”</emphasis>
,
<emphasis id="B936EA8BFFA2745783D23FEBFCD6FD22" box="[753,776,691,717]" italics="true" pageId="4" pageNumber="341">u</emphasis>
,
<emphasis id="B936EA8BFFA2745782343FEBFCE5FD22" box="[791,827,691,717]" italics="true" pageId="4" pageNumber="341">pv</emphasis>
,
<emphasis id="B936EA8BFFA2745782693FEBFCADFD22" box="[842,883,691,717]" italics="true" pageId="4" pageNumber="341">pv”</emphasis>
). Claws simple, empodium as on tarsi II and III. Seta
<emphasis id="B936EA8BFFA2745781EC3F8FFF3BFD1E" box="[207,229,727,753]" italics="true" pageId="4" pageNumber="341">v</emphasis>
of trochanter smooth, other setae barbed; setae
<emphasis id="B936EA8BFFA2745782233F8FFCC8FD1E" box="[768,790,727,753]" italics="true" pageId="4" pageNumber="341">v</emphasis>
of trochanter,
<emphasis id="B936EA8BFFA2745782983F8FFC17FD1E" box="[955,969,727,753]" italics="true" pageId="4" pageNumber="341">d</emphasis>
,
<emphasis id="B936EA8BFFA2745782FB3F8FFC30FD1E" box="[984,1006,727,753]" italics="true" pageId="4" pageNumber="341">v</emphasis>
of femur,
<emphasis id="B936EA8BFFA2745785473F8FFBA1FD1E" box="[1124,1151,727,753]" italics="true" pageId="4" pageNumber="341">v”</emphasis>
of tibia and
<emphasis id="B936EA8BFFA27457842C3F8FFAEBFD1E" box="[1295,1333,727,753]" italics="true" pageId="4" pageNumber="341">pl”</emphasis>
of tarsus blunt-tipped, other leg setae pointed; solenidion
<emphasis id="B936EA8BFFA2745783963FA3FD1AFCFB" box="[693,708,763,788]" italics="true" pageId="4" pageNumber="341">φ</emphasis>
absent.
</paragraph>
<caption id="DF3D6611FFA2745781B43BB5FF29F8C6" pageId="4" pageNumber="341" startId="4.[151,250,1773,1797]" targetBox="[151,1436,813,1749]" targetPageId="4" targetType="figure">
<paragraph id="8BFD3699FFA2745781B43BB5FF29F8C6" blockId="4.[151,1436,1773,1833]" pageId="4" pageNumber="341">
<emphasis id="B936EA8BFFA2745781B43BB5FECEF8EA" bold="true" box="[151,272,1773,1797]" pageId="4" pageNumber="341">FIGURE 4</emphasis>
. DIC micrographs of
<taxonomicName id="4C424D1AFFA2745780DA3BB5FCCCF8EA" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[505,786,1773,1797]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="341" phylum="Arthropoda" rank="species" species="brevisetosus" status="sp. nov.">
<emphasis id="B936EA8BFFA2745780DA3BB5FCCCF8EA" box="[505,786,1773,1797]" italics="true" pageId="4" pageNumber="341">Spatulaphorus brevisetosus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFA2745782383BB5FCB1F8EA" bold="true" box="[795,879,1773,1797]" pageId="4" pageNumber="341">
<taxonomicNameLabel id="A20557F0FFA2745782383BB5FCB1F8EA" box="[795,879,1773,1797]" pageId="4" pageNumber="341" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—general view dorsally; B—general view ventrally.
</paragraph>
</caption>
<paragraph id="8BFD3699FFA2745781EB3A0FFE4FF89E" blockId="4.[151,1436,1879,1978]" box="[200,401,1879,1905]" pageId="4" pageNumber="341">MALE unknown.</paragraph>
</subSubSection>
<subSubSection id="C3586512FFA2745681E43A23FD9BFF17" lastPageId="5" lastPageNumber="342" pageId="4" pageNumber="341" type="materials_examined">
<paragraph id="8BFD3699FFA2745781E43A23FC3EF856" blockId="4.[151,1436,1879,1978]" pageId="4" pageNumber="341">
<emphasis id="B936EA8BFFA2745781E43A23FEAFF87A" bold="true" box="[199,369,1915,1941]" pageId="4" pageNumber="341">Type material</emphasis>
. Female
<materialsCitation id="3B2A3CC4FFA2745780C33A23FD85F855" collectingDate="1970-07-08" collectingDateMax="1970-07-09" collectingDateMin="1970-07-08" collectionCode="ZISP, CEMT" country="United States of America" location="Blackbeard Island" pageId="4" pageNumber="341" specimenCount="1" typeStatus="holotype">
<typeStatus id="54F9883BFFA2745780C33A23FD9BF87A" box="[480,581,1915,1941]" pageId="4" pageNumber="341" type="holotype">holotype</typeStatus>
, slide
<collectionCode id="ED53AE5CFFA2745783B73A23FD13F87A" box="[660,717,1915,1941]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="4" pageNumber="341">ZISP</collectionCode>
T-Pygm-010,
<collectingCountry id="F3557609FFA2745782563A23FC6EF87A" box="[885,944,1915,1941]" name="United States of America" pageId="4" pageNumber="341">USA</collectingCountry>
, Georgia,
<location id="8E9D6042FFA2745785093A23FB22F87A" LSID="urn:lsid:plazi:treatment:03EB878FFFA4745681B43C8BFC49FE1A:8E9D6042FFA2745785093A23FB22F87A" box="[1066,1276,1915,1941]" country="United States of America" name="Blackbeard Island" pageId="4" pageNumber="341">Blackbeard Island</location>
.
<date id="FFFC1059FFA2745784293A23FA42F87A" box="[1290,1436,1915,1941]" pageId="4" pageNumber="341" value="1970-07-08" valueMax="1970-07-09" valueMin="1970-07-08">
<collectingDate id="EFB8E9B1FFA2745784293A23FA42F87A" box="[1290,1436,1915,1941]" pageId="4" pageNumber="341" value="1970-07-08" valueMax="1970-07-09" valueMin="1970-07-08">8-9.VII.1970</collectingDate>
</date>
(
<collectionCode id="ED53AE5CFFA2745781BC3AC7FF2CF856" box="[159,242,1951,1977]" pageId="4" pageNumber="341">CEMT</collectionCode>
),
<collectedFrom id="385E7564FFA2745780253AF8FD85F855" box="[262,603,1951,1978]" pageId="4" pageNumber="341">
on
<taxonomicName id="4C424D1AFFA27457800A3AC7FD85F855" authority="MacLeay" authorityName="MacLeay" authorityYear="1819" box="[297,603,1951,1978]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="341" phylum="Arthropoda" rank="species" species="igneus">
<emphasis id="B936EA8BFFA27457800A3AC7FE39F856" box="[297,487,1951,1977]" italics="true" pageId="4" pageNumber="341">Phanaeus igneus</emphasis>
MacLeay
</taxonomicName>
</collectedFrom>
</materialsCitation>
;
<materialsCitation id="3B2A3CC4FFA2745783443AF8FC05F856" box="[615,987,1951,1978]" collectingDate="1970-07-08" collectingDateMax="1970-07-09" collectingDateMin="1970-07-08" collectionCode="Male" country="United States of America" location="Blackbeard Island" pageId="4" pageNumber="341" specimenCount="17" specimenCount-female="17" typeStatus="paratype">
<typeStatus id="54F9883BFFA2745783443AF8FD09F855" box="[615,727,1952,1978]" pageId="4" pageNumber="341" type="paratype">paratypes</typeStatus>
:
<specimenCount id="9D44FD10FFA2745783C23AC7FC80F856" box="[737,862,1951,1977]" count="17" pageId="4" pageNumber="341" type="female">17 females</specimenCount>
, same data
</materialsCitation>
.
</paragraph>
<paragraph id="8BFD3699FFA3745681E43DCEFD9BFF17" blockId="5.[151,1437,150,501]" pageId="5" pageNumber="342">
<emphasis id="B936EA8BFFA3745681E43DCEFE58FF5F" bold="true" box="[199,390,150,176]" pageId="5" pageNumber="342">Type deposition</emphasis>
.
<materialsCitation id="3B2A3CC4FFA3745680B43DCEFE25FF3B" country="Russia" location="Zoological Institute" pageId="5" pageNumber="342" specimenCount="4" stateProvince="Saint Petersburg" typeStatus="holotype">
The
<typeStatus id="54F9883BFFA3745680EF3DCEFDF0FF5F" box="[460,558,150,176]" pageId="5" pageNumber="342" type="holotype">holotype</typeStatus>
and
<specimenCount id="9D44FD10FFA3745683493DCEFCD3FF5E" box="[618,781,150,177]" count="4" pageId="5" pageNumber="342" type="generic" typeStatus="paratype">four paratypes</specimenCount>
are deposited in the collection of
<location id="8E9D6042FFA3745685BF3DCEFAA5FF5F" LSID="urn:lsid:plazi:treatment:03EB878FFFA4745681B43C8BFC49FE1A:8E9D6042FFA3745685BF3DCEFAA5FF5F" box="[1180,1403,150,176]" country="Russia" name="Zoological Institute" pageId="5" pageNumber="342" stateProvince="Saint Petersburg">Zoological Institute</location>
of RAS,
<collectingRegion id="4986F87BFFA3745681FC3DE2FE40FF3B" box="[223,414,186,212]" country="Russia" name="Saint Petersburg" pageId="5" pageNumber="342">Saint Petersburg</collectingRegion>
,
<collectingCountry id="F3557609FFA37456808E3DE2FE25FF3B" box="[429,507,186,212]" name="Russia" pageId="5" pageNumber="342">Russia</collectingCountry>
</materialsCitation>
;
<materialsCitation id="3B2A3CC4FFA3745683293DE2FAC5FF3B" box="[522,1307,186,213]" country="Russia" location="Tyumen State" pageId="5" pageNumber="342" specimenCount="1" stateProvince="Tyumen'" typeStatus="paratype">
other
<typeStatus id="54F9883BFFA37456836D3DE3FD67FF3A" box="[590,697,187,213]" pageId="5" pageNumber="342" type="paratype">paratypes</typeStatus>
are deposited in the collection of the
<collectingRegion id="4986F87BFFA37456855E3DE3FAC5FF3B" box="[1149,1307,186,213]" country="Russia" name="Tyumen'" pageId="5" pageNumber="342">Tyumen State</collectingRegion>
</materialsCitation>
<materialsCitation id="3B2A3CC4FFA3745684053DE2FD9EFF17" country="Russia" location="University Museum of Zoology" pageId="5" pageNumber="342" specimenCount="1" stateProvince="Tyumen'" typeStatus="paratype">
<location id="8E9D6042FFA3745684053DE2FEA1FF17" LSID="urn:lsid:plazi:treatment:03EB878FFFA4745681B43C8BFC49FE1A:8E9D6042FFA3745684053DE2FEA1FF17" country="Russia" name="University Museum of Zoology" pageId="5" pageNumber="342" stateProvince="Tyumen'">University Museum of Zoology</location>
,
<collectingRegion id="4986F87BFFA3745680AA3D87FE39FF16" box="[393,487,223,249]" country="Russia" name="Tyumen'" pageId="5" pageNumber="342">Tyumen</collectingRegion>
,
<collectingCountry id="F3557609FFA3745680D13D86FD9EFF17" box="[498,576,222,248]" name="Russia" pageId="5" pageNumber="342">Russia</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3586512FFA3745681E43C5AFEC0FE43" pageId="5" pageNumber="342" type="diagnosis">
<paragraph id="8BFD3699FFA3745681E43C5AFEC0FE43" blockId="5.[151,1437,150,501]" pageId="5" pageNumber="342">
<emphasis id="B936EA8BFFA3745681E43C5AFE19FEF3" bold="true" box="[199,455,258,284]" pageId="5" pageNumber="342">Differential diagnosis</emphasis>
. The new species is most similar to
<taxonomicName id="4C424D1AFFA3745682413C5BFB36FEF3" authority="Trejo-Palacios, 2023" authorityName="Trejo-Palacios" authorityYear="2023" box="[866,1256,258,285]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="342" phylum="Arthropoda" rank="species" species="mexicanus">
<emphasis id="B936EA8BFFA3745682413C5BFCAEFEF2" box="[866,880,259,285]" italics="true" pageId="5" pageNumber="342">S</emphasis>
.
<emphasis id="B936EA8BFFA37456825D3C5BFC2AFEF3" box="[894,1012,259,284]" italics="true" pageId="5" pageNumber="342">mexicanus</emphasis>
<bibRefCitation id="EFD34B68FFA3745682DF3C5AFB36FEF3" author="Trejo-Palacios, S. J. &amp; Toledo-Hernandez, V. H. &amp; Paredes-Leon, R. &amp; Corona-Lopez, A. M." box="[1020,1256,258,284]" pageId="5" pageNumber="342" pagination="77 - 87" refId="ref16938" refString="Trejo-Palacios, S. J., Toledo-Hernandez, V. H., Paredes-Leon, R. &amp; Corona-Lopez, A. M. (2023) A new species and new records of Pygmephoridae (Acari: Heterostigmata) phoretic on Coleoptera from Mexico. Systematic &amp; Applied Acarology, 28 (1), 77 - 87. https: // doi. org / 10.11158 / saa. 28.1.8" type="journal article" year="2023">Trejo-Palacios, 2023</bibRefCitation>
</taxonomicName>
, described from
<collectingCountry id="F3557609FFA3745681B43C7EFF33FEAF" box="[151,237,294,320]" name="Mexico" pageId="5" pageNumber="342">Mexico</collectingCountry>
(
<bibRefCitation id="EFD34B68FFA3745681DF3C7EFDF8FEAF" author="Trejo-Palacios, S. J. &amp; Toledo-Hernandez, V. H. &amp; Paredes-Leon, R. &amp; Corona-Lopez, A. M." box="[252,550,294,320]" pageId="5" pageNumber="342" pagination="77 - 87" refId="ref16938" refString="Trejo-Palacios, S. J., Toledo-Hernandez, V. H., Paredes-Leon, R. &amp; Corona-Lopez, A. M. (2023) A new species and new records of Pygmephoridae (Acari: Heterostigmata) phoretic on Coleoptera from Mexico. Systematic &amp; Applied Acarology, 28 (1), 77 - 87. https: // doi. org / 10.11158 / saa. 28.1.8" type="journal article" year="2023">
Trejo-Palacios
<emphasis id="B936EA8BFFA37456808A3C7FFE3CFEAF" box="[425,482,294,320]" italics="true" pageId="5" pageNumber="342">et al.</emphasis>
2023
</bibRefCitation>
), in having spiniform setae
<emphasis id="B936EA8BFFA3745682423C7EFC6AFEAE" box="[865,948,294,321]" italics="true" pageId="5" pageNumber="342">pl”, tc</emphasis>
on tarsus II and
<emphasis id="B936EA8BFFA3745685573C7EFB44FEAF" box="[1140,1178,294,320]" italics="true" pageId="5" pageNumber="342">pl”</emphasis>
on tarsus III. The new species differs from
<taxonomicName id="4C424D1AFFA3745680A23C13FDCBFE8B" authorityName="Trejo-Palacios" authorityYear="2023" box="[385,533,331,357]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="342" phylum="Arthropoda" rank="species" species="mexicanus">
<emphasis id="B936EA8BFFA3745680A23C13FE51FE8A" box="[385,399,331,357]" italics="true" pageId="5" pageNumber="342">S</emphasis>
.
<emphasis id="B936EA8BFFA3745680BC3C13FDCBFE8B" box="[415,533,331,356]" italics="true" pageId="5" pageNumber="342">mexicanus</emphasis>
</taxonomicName>
in having setae
<emphasis id="B936EA8BFFA3745683F03C13FD01FE8B" box="[723,735,331,356]" italics="true" pageId="5" pageNumber="342">e</emphasis>
and
<emphasis id="B936EA8BFFA37456823A3C12FCFFFE8B" box="[793,801,330,356]" italics="true" pageId="5" pageNumber="342">f</emphasis>
as well as
<emphasis id="B936EA8BFFA3745682803C12FC61FE8B" box="[931,959,330,356]" italics="true" pageId="5" pageNumber="342">h1</emphasis>
and
<emphasis id="B936EA8BFFA3745682DA3C12FBCBFE8B" box="[1017,1045,330,356]" italics="true" pageId="5" pageNumber="342">h2</emphasis>
distinctly separated (vs. alveoli of setae
<emphasis id="B936EA8BFFA3745681F43C37FF3DFE67" box="[215,227,367,392]" italics="true" pageId="5" pageNumber="342">e</emphasis>
and
<emphasis id="B936EA8BFFA37456803F3C36FEFAFE67" box="[284,292,366,392]" italics="true" pageId="5" pageNumber="342">f</emphasis>
as well as
<emphasis id="B936EA8BFFA3745680803C36FE61FE67" box="[419,447,366,392]" italics="true" pageId="5" pageNumber="342">h1</emphasis>
and
<emphasis id="B936EA8BFFA3745680D43C36FDCAFE67" box="[503,532,366,392]" italics="true" pageId="5" pageNumber="342">h2</emphasis>
contiguous in
<taxonomicName id="4C424D1AFFA37456839F3C37FC91FE67" authorityName="Trejo-Palacios" authorityYear="2023" box="[700,847,367,393]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="342" phylum="Arthropoda" rank="species" species="mexicanus">
<emphasis id="B936EA8BFFA37456839F3C37FD14FE66" box="[700,714,367,393]" italics="true" pageId="5" pageNumber="342">S</emphasis>
.
<emphasis id="B936EA8BFFA3745683FA3C37FC91FE67" box="[729,847,367,392]" italics="true" pageId="5" pageNumber="342">mexicanus</emphasis>
</taxonomicName>
) and in having smooth setae
<emphasis id="B936EA8BFFA3745685BD3C36FB05FE67" box="[1182,1243,366,392]" italics="true" pageId="5" pageNumber="342">ps1-3</emphasis>
(vs. barbed in
<taxonomicName id="4C424D1AFFA3745684A43C37FED3FE43" authorityName="Trejo-Palacios" authorityYear="2023" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="342" phylum="Arthropoda" rank="species" species="mexicanus">
<emphasis id="B936EA8BFFA3745684A43C37FED3FE43" italics="true" pageId="5" pageNumber="342">S. mexicanus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3586512FFA3745681E43CEEFC49FE1A" pageId="5" pageNumber="342" type="etymology">
<paragraph id="8BFD3699FFA3745681E43CEEFC49FE1A" blockId="5.[151,1437,150,501]" pageId="5" pageNumber="342">
<emphasis id="B936EA8BFFA3745681E43CEEFE96FE3F" bold="true" box="[199,328,438,464]" pageId="5" pageNumber="342">Etymology</emphasis>
. The name of the new species is a combination of to Latin words:
<emphasis id="B936EA8BFFA3745685613CEEFB5BFE3F" box="[1090,1157,438,464]" italics="true" pageId="5" pageNumber="342">brevis</emphasis>
meaning
<emphasis id="B936EA8BFFA3745685D53CEEFAEEFE3F" box="[1270,1328,438,464]" italics="true" pageId="5" pageNumber="342">short</emphasis>
, and
<emphasis id="B936EA8BFFA37456844C3CEFFA42FE3F" box="[1391,1436,439,464]" italics="true" pageId="5" pageNumber="342">seta</emphasis>
meaning
<emphasis id="B936EA8BFFA3745681DC3C82FE99FE1B" box="[255,327,474,500]" italics="true" pageId="5" pageNumber="342">bristle</emphasis>
, and refers to relatively short dorsal idiosomal setae.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,514 @@
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<mods:title id="E4D474276FE5E2991144850A11598F1D">Seven new species of Spatulaphorus Rack (Acari: Pygmephoridae) phoretic on scarab beetles (Coleoptera: Scarabaeidae)</mods:title>
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<mods:affiliation id="4531BE9AF4FCAD7E2E94E3A43D0880A4">Zoological Institute of Russian Academy of Sciences, Universitetskaya Embankment 1, 199034, Saint Petersburg, Russia</mods:affiliation>
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<paragraph id="8BFD3699FFAB745E81B43F1EFDB6FD8E" blockId="13.[151,616,582,645]" box="[151,616,582,609]" pageId="13" pageNumber="350">
<heading id="D0B581F5FFAB745E81B43F1EFDB6FD8E" bold="true" box="[151,616,582,609]" fontSize="11" level="1" pageId="13" pageNumber="350" reason="1">
<emphasis id="B936EA8BFFAB745E81B43F1EFDB6FD8E" bold="true" box="[151,616,582,609]" pageId="13" pageNumber="350">
<taxonomicName id="4C424D1AFFAB745E81B43F1EFDDDFD8E" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[151,515,582,609]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="13" pageNumber="350" phylum="Arthropoda" rank="species" species="venezuelaensis" status="sp. nov.">
<emphasis id="B936EA8BFFAB745E81B43F1EFDDDFD8E" bold="true" box="[151,515,582,609]" italics="true" pageId="13" pageNumber="350">Spatulaphorus venezuelaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20557F0FFAB745E83283F1FFDB6FD8E" box="[523,616,583,609]" pageId="13" pageNumber="350" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C3586512FFAB744381B43F33FE4FFBBF" lastPageId="16" lastPageNumber="353" pageId="13" pageNumber="350" type="description">
<paragraph id="8BFD3699FFAB745E81B43F33FEF9FD6A" blockId="13.[151,616,582,645]" box="[151,295,619,645]" pageId="13" pageNumber="350">
(
<figureCitation id="13792A1CFFAB745E81BC3F33FEC1FD6A" box="[159,287,619,645]" captionStart-0="FIGURE 13" captionStart-1="FIGURE 14" captionStart-2="FIGURE 15" captionStart-3="FIGURE 16" captionStartId-0="13.[151,250,1900,1924]" captionStartId-1="14.[151,250,1368,1392]" captionStartId-2="15.[151,250,1443,1467]" captionStartId-3="16.[151,250,972,996]" captionTargetBox-0="[151,1435,746,1877]" captionTargetBox-1="[151,1435,348,1344]" captionTargetBox-2="[151,1435,189,1420]" captionTargetBox-3="[151,1436,181,948]" captionTargetId-0="figure-203@13.[151,1435,741,1877]" captionTargetId-1="figure-105@14.[151,1435,345,1344]" captionTargetId-2="figure-17@15.[151,1435,181,1420]" captionTargetId-3="figure-24@16.[151,1436,181,948]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="15" captionTargetPageId-3="16" captionText-0="FIGURE 13. Spatulaphorus venezuelaensis sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." captionText-1="FIGURE 14. Spatulaphorus venezuelaensis sp. nov., female: A—right leg I, dorsal aspect; B—right leg II, dorsal aspect." captionText-2="FIGURE 15. Spatulaphorus venezuelaensis sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." captionText-3="FIGURE 16. DIC micrographs of Spatulaphorus venezuelaensis sp. nov., female: A—general view dorsally; B—general view ventrally." pageId="13" pageNumber="350">Figs 1316</figureCitation>
)
</paragraph>
<paragraph id="8BFD3699FFAB745E81B43FEAFA9EFD22" blockId="13.[151,1344,690,717]" box="[151,1344,690,717]" pageId="13" pageNumber="350">
<emphasis id="B936EA8BFFAB745E81B43FEAFEFFFD23" bold="true" box="[151,289,690,716]" pageId="13" pageNumber="350">Description</emphasis>
. FEMALE. Body well sclerotized. Length of idiosoma 205 (175255), width 130 (105150).
</paragraph>
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<paragraph id="8BFD3699FFAB745E81B43A34FAE4F86A" blockId="13.[151,1338,1900,1925]" box="[151,1338,1900,1925]" pageId="13" pageNumber="350">
<emphasis id="B936EA8BFFAB745E81B43A34FEC7F86A" bold="true" box="[151,281,1900,1925]" pageId="13" pageNumber="350">FIGURE 13</emphasis>
.
<taxonomicName id="4C424D1AFFAB745E80053A35FD89F86A" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[294,599,1901,1925]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="13" pageNumber="350" phylum="Arthropoda" rank="species" species="venezuelaensis" status="sp. nov.">
<emphasis id="B936EA8BFFAB745E80053A35FD89F86A" box="[294,599,1901,1925]" italics="true" pageId="13" pageNumber="350">Spatulaphorus venezuelaensis</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFAB745E837E3A35FD70F86A" bold="true" box="[605,686,1901,1925]" pageId="13" pageNumber="350">
<taxonomicNameLabel id="A20557F0FFAB745E837E3A35FD70F86A" box="[605,686,1901,1925]" pageId="13" pageNumber="350" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—dorsum of body; B—venter of body. Legs omitted.
</paragraph>
</caption>
<paragraph id="8BFD3699FFAB745E81E43AEBFA43F822" blockId="13.[151,1437,1971,2034]" box="[199,1437,1971,1998]" pageId="13" pageNumber="350">
<emphasis id="B936EA8BFFAB745E81E43AEBFE88F822" box="[199,342,1971,1997]" italics="true" pageId="13" pageNumber="350">Gnathosoma</emphasis>
. Gnathosomal capsule, excluding palps, subrectangular, length 24 (2227), width 25 (2327).
</paragraph>
<paragraph id="8BFD3699FFAB745D81B43A8FFD3BFEAE" blockId="13.[151,1437,1971,2034]" lastBlockId="14.[151,1437,151,321]" lastPageId="14" lastPageNumber="351" pageId="13" pageNumber="350">
Dorsum with one pair of needle-like cheliceral setae (
<emphasis id="B936EA8BFFAB745E823D3A8FFC98F81E" box="[798,838,2007,2033]" italics="true" pageId="13" pageNumber="350">cha</emphasis>
) 5 (45). Dorsal median apodeme present, poorly developed. Postpalpal setae (
<emphasis id="B936EA8BFFA8745D80F93DCFFE28FF5F" box="[474,502,151,176]" italics="true" pageId="14" pageNumber="351">pp</emphasis>
) 4 (34) needle-like, situated laterad setae
<emphasis id="B936EA8BFFA8745D82F03DCFFC25FF5E" box="[979,1019,151,177]" italics="true" pageId="14" pageNumber="351">cha</emphasis>
. Ventral gnathosoma with one pair of smooth, pointed subcapitular setae
<emphasis id="B936EA8BFFA8745D83003DE3FDE9FF3B" box="[547,567,187,212]" italics="true" pageId="14" pageNumber="351">m</emphasis>
7 (67) and round pits
<emphasis id="B936EA8BFFA8745D82633DE3FC90FF3B" box="[832,846,187,212]" italics="true" pageId="14" pageNumber="351">n</emphasis>
situated posteriad
<emphasis id="B936EA8BFFA8745D85013DE3FBE8FF3B" box="[1058,1078,187,212]" italics="true" pageId="14" pageNumber="351">m</emphasis>
. Palps with smooth and weakly blunt-tipped setae
<emphasis id="B936EA8BFFA8745D80443D87FE4DFF16" box="[359,403,223,249]" italics="true" pageId="14" pageNumber="351">dFe</emphasis>
4 (34) and
<emphasis id="B936EA8BFFA8745D83003D87FD8CFF16" box="[547,594,223,249]" italics="true" pageId="14" pageNumber="351">dGe</emphasis>
7 (78) dorsolaterally. Palps ventrally each with mushroom-like accessory setigenous structure (
<emphasis id="B936EA8BFFA8745D80AC3C5BFE6DFEF3" box="[399,435,259,284]" italics="true" pageId="14" pageNumber="351">ass</emphasis>
) and tiny solenidion
<emphasis id="B936EA8BFFA8745D83933C5BFD1AFEF3" box="[688,708,259,284]" italics="true" pageId="14" pageNumber="351">ω</emphasis>
. Palps terminated with small blunt-tipped tibial claw and tiny eupathid-like seta. Pharyngeal pumps poorly visible.
</paragraph>
<caption id="DF3D6611FFA8745D81B43800FAB9FA80" box="[151,1383,1368,1392]" pageId="14" pageNumber="351" startId="14.[151,250,1368,1392]" targetBox="[151,1435,348,1344]" targetPageId="14" targetType="figure">
<paragraph id="8BFD3699FFA8745D81B43800FAB9FA80" blockId="14.[151,1383,1368,1392]" box="[151,1383,1368,1392]" pageId="14" pageNumber="351">
<emphasis id="B936EA8BFFA8745D81B43800FEC7FA9F" bold="true" box="[151,281,1368,1392]" pageId="14" pageNumber="351">FIGURE 14</emphasis>
.
<taxonomicName id="4C424D1AFFA8745D80053800FD89FA9F" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[294,599,1368,1392]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="14" pageNumber="351" phylum="Arthropoda" rank="species" species="venezuelaensis" status="sp. nov.">
<emphasis id="B936EA8BFFA8745D80053800FD89FA9F" box="[294,599,1368,1392]" italics="true" pageId="14" pageNumber="351">Spatulaphorus venezuelaensis</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFA8745D837E3800FD70FA9F" bold="true" box="[605,686,1368,1392]" pageId="14" pageNumber="351">
<taxonomicNameLabel id="A20557F0FFA8745D837E3800FD70FA9F" box="[605,686,1368,1392]" pageId="14" pageNumber="351" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—right leg I, dorsal aspect; B—right leg II, dorsal aspect.
</paragraph>
</caption>
<paragraph id="8BFD3699FFA8745D81E438C6FC04F95B" blockId="14.[151,1437,1438,2040]" pageId="14" pageNumber="351">
<emphasis id="B936EA8BFFA8745D81E438C6FE4BFA57" box="[199,405,1438,1464]" italics="true" pageId="14" pageNumber="351">Idiosomal dorsum</emphasis>
(
<figureCitation id="13792A1CFFA8745D808438C6FDCBFA57" box="[423,533,1438,1464]" captionStart="FIGURE 13" captionStartId="13.[151,250,1900,1924]" captionTargetBox="[151,1435,746,1877]" captionTargetId="figure-203@13.[151,1435,741,1877]" captionTargetPageId="13" captionText="FIGURE 13. Spatulaphorus venezuelaensis sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." pageId="14" pageNumber="351">Figs 13A</figureCitation>
,
<figureCitation id="13792A1CFFA8745D830738C6FD89FA57" box="[548,599,1438,1464]" captionStart="FIGURE 16" captionStartId="16.[151,250,972,996]" captionTargetBox="[151,1436,181,948]" captionTargetId="figure-24@16.[151,1436,181,948]" captionTargetPageId="16" captionText="FIGURE 16. DIC micrographs of Spatulaphorus venezuelaensis sp. nov., female: A—general view dorsally; B—general view ventrally." pageId="14" pageNumber="351">16A</figureCitation>
). All dorsal shields with big puncta. Prodorsal shield with three pairs of setae (
<emphasis id="B936EA8BFFA8745D81C3389AFF24FA33" box="[224,250,1474,1500]" italics="true" pageId="14" pageNumber="351">v1</emphasis>
,
<emphasis id="B936EA8BFFA8745D802A389AFEFDFA33" box="[265,291,1474,1500]" italics="true" pageId="14" pageNumber="351">v2</emphasis>
,
<emphasis id="B936EA8BFFA8745D8012389AFE89FA33" box="[305,343,1474,1500]" italics="true" pageId="14" pageNumber="351">sc2</emphasis>
), one pair of weakly barbed capitate trichobothria (
<emphasis id="B936EA8BFFA8745D82B8389AFC1FFA33" box="[923,961,1474,1500]" italics="true" pageId="14" pageNumber="351">sc1</emphasis>
) and one pair of round stigmata. Stigmata associated with well-developed tracheal trunks. All dorsal idiosomal setae blunt-tipped and barbed. Alveoli of setae
<emphasis id="B936EA8BFFA8745D81B43B53FF7DF9CB" box="[151,163,1547,1572]" italics="true" pageId="14" pageNumber="351">e</emphasis>
and
<emphasis id="B936EA8BFFA8745D81FF3B52FF3AF9CB" box="[220,228,1546,1572]" italics="true" pageId="14" pageNumber="351">f</emphasis>
distinctly separated; setae
<emphasis id="B936EA8BFFA8745D83353B53FDFCF9CB" box="[534,546,1547,1572]" italics="true" pageId="14" pageNumber="351">e</emphasis>
located slightly anteriad
<emphasis id="B936EA8BFFA8745D82613B52FC94F9CB" box="[834,842,1546,1572]" italics="true" pageId="14" pageNumber="351">f</emphasis>
. Cupules
<emphasis id="B936EA8BFFA8745D829E3B53FC0DF9CB" box="[957,979,1547,1572]" italics="true" pageId="14" pageNumber="351">ia</emphasis>
on tergite D,
<emphasis id="B936EA8BFFA8745D85513B53FB50F9CB" box="[1138,1166,1547,1572]" italics="true" pageId="14" pageNumber="351">im</emphasis>
on tergite EF and
<emphasis id="B936EA8BFFA8745D84413B52FAA6F9CB" box="[1378,1400,1546,1572]" italics="true" pageId="14" pageNumber="351">ih</emphasis>
on tergite H large, round, subequal. Lengths of dorsal setae:
<emphasis id="B936EA8BFFA8745D82153B76FC8EF9A7" box="[822,848,1582,1608]" italics="true" pageId="14" pageNumber="351">v1</emphasis>
34 (2840),
<emphasis id="B936EA8BFFA8745D82C83B76FBD8F9A7" box="[1003,1030,1582,1608]" italics="true" pageId="14" pageNumber="351">v2</emphasis>
17 (1619),
<emphasis id="B936EA8BFFA8745D85833B76FB18F9A7" box="[1184,1222,1582,1608]" italics="true" pageId="14" pageNumber="351">sc2</emphasis>
61 (4476),
<emphasis id="B936EA8BFFA8745D84433B76FAA5F9A7" box="[1376,1403,1582,1608]" italics="true" pageId="14" pageNumber="351">c1</emphasis>
53 (4059),
<emphasis id="B936EA8BFFA8745D81DE3B0AFEC6F983" box="[253,280,1618,1644]" italics="true" pageId="14" pageNumber="351">c2</emphasis>
50 (3962),
<emphasis id="B936EA8BFFA8745D80843B0AFE6BF983" box="[423,437,1618,1644]" italics="true" pageId="14" pageNumber="351">d</emphasis>
57 (4170),
<emphasis id="B936EA8BFFA8745D83673B0BFD8EF983" box="[580,592,1619,1644]" italics="true" pageId="14" pageNumber="351">e</emphasis>
31 (2233),
<emphasis id="B936EA8BFFA8745D83FC3B0AFD39F983" box="[735,743,1618,1644]" italics="true" pageId="14" pageNumber="351">f</emphasis>
70 (5187),
<emphasis id="B936EA8BFFA8745D82553B0AFC4FF983" box="[886,913,1618,1644]" italics="true" pageId="14" pageNumber="351">h1</emphasis>
60 (4675),
<emphasis id="B936EA8BFFA8745D85023B0AFBE3F983" box="[1057,1085,1618,1644]" italics="true" pageId="14" pageNumber="351">h2</emphasis>
21 (1221). Distances between setae:
<emphasis id="B936EA8BFFA8745D81C13B2EFEFAF97F" box="[226,292,1654,1681]" italics="true" pageId="14" pageNumber="351">v1v1</emphasis>
25 (2229),
<emphasis id="B936EA8BFFA8745D80E23B2EFDDDF97F" box="[449,515,1654,1680]" italics="true" pageId="14" pageNumber="351">v2v2</emphasis>
44 (3649),
<emphasis id="B936EA8BFFA8745D83833B2EFD26F97F" box="[672,760,1654,1680]" italics="true" pageId="14" pageNumber="351">sc2sc2</emphasis>
44 (3649),
<emphasis id="B936EA8BFFA8745D82B63B2EFC06F97F" box="[917,984,1654,1681]" italics="true" pageId="14" pageNumber="351">c1c1</emphasis>
44 (3649),
<emphasis id="B936EA8BFFA8745D85533B2EFB6DF97F" box="[1136,1203,1654,1681]" italics="true" pageId="14" pageNumber="351">c1c2</emphasis>
24 (1926),
<emphasis id="B936EA8BFFA8745D846C3B2EFAA7F97F" box="[1359,1401,1654,1681]" italics="true" pageId="14" pageNumber="351">dd</emphasis>
70 (5476),
<emphasis id="B936EA8BFFA8745D81DD3BC3FEFEF95B" box="[254,288,1690,1717]" italics="true" pageId="14" pageNumber="351">ef</emphasis>
8 (611),
<emphasis id="B936EA8BFFA8745D80B73BC2FE6FF95B" box="[404,433,1690,1717]" italics="true" pageId="14" pageNumber="351">ff</emphasis>
56 (4160),
<emphasis id="B936EA8BFFA8745D83613BC2FD56F95B" box="[578,648,1690,1717]" italics="true" pageId="14" pageNumber="351">h1h1</emphasis>
51 (3753),
<emphasis id="B936EA8BFFA8745D823A3BC2FC81F95B" box="[793,863,1690,1717]" italics="true" pageId="14" pageNumber="351">h1h2</emphasis>
10 (910).
</paragraph>
<paragraph id="8BFD3699FFA8745D81E43BE6FF29F817" blockId="14.[151,1437,1438,2040]" pageId="14" pageNumber="351">
<emphasis id="B936EA8BFFA8745D81E43BE6FE5AF937" box="[199,388,1726,1752]" italics="true" pageId="14" pageNumber="351">Idiosomal venter</emphasis>
(
<figureCitation id="13792A1CFFA8745D80B03BE6FE22F937" box="[403,508,1726,1752]" captionStart="FIGURE 13" captionStartId="13.[151,250,1900,1924]" captionTargetBox="[151,1435,746,1877]" captionTargetId="figure-203@13.[151,1435,741,1877]" captionTargetPageId="13" captionText="FIGURE 13. Spatulaphorus venezuelaensis sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." pageId="14" pageNumber="351">Figs 13B</figureCitation>
,
<figureCitation id="13792A1CFFA8745D832B3BE6FDE4F937" box="[520,570,1726,1752]" captionStart="FIGURE 16" captionStartId="16.[151,250,972,996]" captionTargetBox="[151,1436,181,948]" captionTargetId="figure-24@16.[151,1436,181,948]" captionTargetPageId="16" captionText="FIGURE 16. DIC micrographs of Spatulaphorus venezuelaensis sp. nov., female: A—general view dorsally; B—general view ventrally." pageId="14" pageNumber="351">16B</figureCitation>
). Coxisternal fields III and aggenital plate laterally with sparsely distributed big puncta. Setae
<emphasis id="B936EA8BFFA8745D807C3BBAFEA2F913" box="[351,380,1762,1788]" italics="true" pageId="14" pageNumber="351">1b</emphasis>
pointed; other ventral setae blunt-tipped; setae
<emphasis id="B936EA8BFFA8745D82B13BBAFC70F913" box="[914,942,1762,1788]" italics="true" pageId="14" pageNumber="351">1b</emphasis>
and
<emphasis id="B936EA8BFFA8745D82C63BBAFBD2F913" box="[997,1036,1762,1788]" italics="true" pageId="14" pageNumber="351">ps3</emphasis>
barbed, other ventral setae smooth. Bases of setae
<emphasis id="B936EA8BFFA8745D80613A5EFEB7F8CF" box="[322,361,1798,1824]" italics="true" pageId="14" pageNumber="351">ps3</emphasis>
situated distinctly anteriad bases of setae
<emphasis id="B936EA8BFFA8745D826E3A5EFC4CF8CF" box="[845,914,1798,1824]" italics="true" pageId="14" pageNumber="351">ps1-2.</emphasis>
Ap1 and ap2 well-developed and joined with thick appr; apsej developed only laterally; ap3 weak, not reaching appo; ap4 well developed, exceeding beyond bases of setae
<emphasis id="B936EA8BFFA8745D801B3A16FE8AF887" box="[312,340,1870,1896]" italics="true" pageId="14" pageNumber="351">3b</emphasis>
. Ap5 short, joined with appo and reaching bases of setae
<emphasis id="B936EA8BFFA8745D82F93A16FC28F887" box="[986,1014,1870,1896]" italics="true" pageId="14" pageNumber="351">4a</emphasis>
. Ags small, cup-like; pgs small, oval. Posterior margin of posterior sternal plate straight in middle part. Posterior margin of aggenital plate rounded. Lengths of ventral setae:
<emphasis id="B936EA8BFFA8745D80903ACEFE0EF85F" box="[435,464,1942,1968]" italics="true" pageId="14" pageNumber="351">1a</emphasis>
10 (912),
<emphasis id="B936EA8BFFA8745D83703ACEFDB1F85F" box="[595,623,1942,1968]" italics="true" pageId="14" pageNumber="351">1b</emphasis>
20 (1723),
<emphasis id="B936EA8BFFA8745D82223ACEFCC5F85F" box="[769,795,1942,1968]" italics="true" pageId="14" pageNumber="351">1с</emphasis>
14 (1015),
<emphasis id="B936EA8BFFA8745D828E3ACEFC16F85F" box="[941,968,1942,1968]" italics="true" pageId="14" pageNumber="351">2a</emphasis>
13 (1214),
<emphasis id="B936EA8BFFA8745D85793ACEFBAAF85F" box="[1114,1140,1942,1968]" italics="true" pageId="14" pageNumber="351">2с</emphasis>
11 (911),
<emphasis id="B936EA8BFFA8745D85D53ACEFACCF85F" box="[1270,1298,1942,1968]" italics="true" pageId="14" pageNumber="351">3a</emphasis>
15 (1015),
<emphasis id="B936EA8BFFA8745D81B43AE2FF6DF83B" box="[151,179,1978,2004]" italics="true" pageId="14" pageNumber="351">3b</emphasis>
13 (1113),
<emphasis id="B936EA8BFFA8745D80663AE2FE81F83B" box="[325,351,1978,2004]" italics="true" pageId="14" pageNumber="351">3c</emphasis>
16 (1217),
<emphasis id="B936EA8BFFA8745D80D13AE2FDD0F83B" box="[498,526,1978,2004]" italics="true" pageId="14" pageNumber="351">4a</emphasis>
13 (1014),
<emphasis id="B936EA8BFFA8745D83823AE2FD63F83B" box="[673,701,1978,2004]" italics="true" pageId="14" pageNumber="351">4b</emphasis>
19 (1524),
<emphasis id="B936EA8BFFA8745D826C3AE2FCB4F83B" box="[847,874,1978,2004]" italics="true" pageId="14" pageNumber="351">4c</emphasis>
14 (1116),
<emphasis id="B936EA8BFFA8745D82D83AE2FBFCF83B" box="[1019,1058,1978,2004]" italics="true" pageId="14" pageNumber="351">ps1</emphasis>
11 (712),
<emphasis id="B936EA8BFFA8745D85853AE2FB13F83B" box="[1190,1229,1978,2004]" italics="true" pageId="14" pageNumber="351">ps2</emphasis>
10 (813),
<emphasis id="B936EA8BFFA8745D84713AE2FAA7F83B" box="[1362,1401,1978,2004]" italics="true" pageId="14" pageNumber="351">ps3</emphasis>
21 (1623).
</paragraph>
<caption id="DF3D6611FFA9745C81B438FBFAA0FA54" box="[151,1406,1443,1468]" pageId="15" pageNumber="352" startId="15.[151,250,1443,1467]" targetBox="[151,1435,189,1420]" targetPageId="15" targetType="figure">
<paragraph id="8BFD3699FFA9745C81B438FBFAA0FA54" blockId="15.[151,1406,1443,1468]" box="[151,1406,1443,1468]" pageId="15" pageNumber="352">
<emphasis id="B936EA8BFFA9745C81B438FBFEC7FA53" bold="true" box="[151,281,1443,1468]" pageId="15" pageNumber="352">FIGURE 15</emphasis>
.
<taxonomicName id="4C424D1AFFA9745C800538FCFD89FA53" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[294,599,1444,1468]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="15" pageNumber="352" phylum="Arthropoda" rank="species" species="venezuelaensis" status="sp. nov.">
<emphasis id="B936EA8BFFA9745C800538FCFD89FA53" box="[294,599,1444,1468]" italics="true" pageId="15" pageNumber="352">Spatulaphorus venezuelaensis</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFA9745C837E38FCFD70FA53" bold="true" box="[605,686,1444,1468]" pageId="15" pageNumber="352">
<taxonomicNameLabel id="A20557F0FFA9745C837E38FCFD70FA53" box="[605,686,1444,1468]" pageId="15" pageNumber="352" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect.
</paragraph>
</caption>
<paragraph id="8BFD3699FFA9745C81E438B3FDC4F837" blockId="15.[151,1437,1514,2008]" pageId="15" pageNumber="352">
<emphasis id="B936EA8BFFA9745C81E438B3FF22F9EB" box="[199,252,1515,1540]" italics="true" pageId="15" pageNumber="352">Legs</emphasis>
(
<figureCitation id="13792A1CFFA9745C802938B2FE81F9EB" box="[266,351,1514,1540]" captionStart="FIGURE 14" captionStartId="14.[151,250,1368,1392]" captionTargetBox="[151,1435,348,1344]" captionTargetId="figure-105@14.[151,1435,345,1344]" captionTargetPageId="14" captionText="FIGURE 14. Spatulaphorus venezuelaensis sp. nov., female: A—right leg I, dorsal aspect; B—right leg II, dorsal aspect." pageId="15" pageNumber="352">Figs 14</figureCitation>
,
<figureCitation id="13792A1CFFA9745C804938B2FE57F9EA" box="[362,393,1514,1541]" captionStart="FIGURE 15" captionStartId="15.[151,250,1443,1467]" captionTargetBox="[151,1435,189,1420]" captionTargetId="figure-17@15.[151,1435,181,1420]" captionTargetPageId="15" captionText="FIGURE 15. Spatulaphorus venezuelaensis sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." pageId="15" pageNumber="352">15</figureCitation>
). Setation of legs as in
<taxonomicName id="4C424D1AFFA9745C83A438B2FCF5F9EB" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[647,811,1514,1540]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="15" pageNumber="352" phylum="Arthropoda" rank="species" species="brevisetosus">
<emphasis id="B936EA8BFFA9745C83A438B2FCF5F9EB" box="[647,811,1514,1540]" italics="true" pageId="15" pageNumber="352">S. brevisetosus</emphasis>
</taxonomicName>
. Leg I (
<figureCitation id="13792A1CFFA9745C82A138B2FC37F9EB" box="[898,1001,1514,1540]" captionStart="FIGURE 14" captionStartId="14.[151,250,1368,1392]" captionTargetBox="[151,1435,348,1344]" captionTargetId="figure-105@14.[151,1435,345,1344]" captionTargetPageId="14" captionText="FIGURE 14. Spatulaphorus venezuelaensis sp. nov., female: A—right leg I, dorsal aspect; B—right leg II, dorsal aspect." pageId="15" pageNumber="352">Fig. 14A</figureCitation>
). Tibiotarsus enlarged, with large claw; tibiotarsus internally with oval strongly sclerotized structure near solenidion
<emphasis id="B936EA8BFFA9745C82D23B56FBD0F9C7" box="[1009,1038,1550,1576]" italics="true" pageId="15" pageNumber="352">φ2</emphasis>
. Setae (
<emphasis id="B936EA8BFFA9745C854B3B57FBA8F9C7" box="[1128,1142,1551,1576]" italics="true" pageId="15" pageNumber="352">u</emphasis>
) consolidated into a horn-like structure opposing claw. Tarsal claw thick and blunt-tipped. Seta
<emphasis id="B936EA8BFFA9745C828F3B6AFC64F9A3" box="[940,954,1586,1612]" italics="true" pageId="15" pageNumber="352">d</emphasis>
of femur spatulate, with short subterminal projection. Setae
<emphasis id="B936EA8BFFA9745C80783B0EFEB2F99F" box="[347,364,1622,1648]" italics="true" pageId="15" pageNumber="352">l</emphasis>
of femur and
<emphasis id="B936EA8BFFA9745C832E3B0EFDC7F99F" box="[525,537,1622,1648]" italics="true" pageId="15" pageNumber="352">k</emphasis>
of tibiotarsus smooth, other leg setae (except eupathidia
<emphasis id="B936EA8BFFA9745C85BF3B0EFB6DF99F" box="[1180,1203,1622,1648]" italics="true" pageId="15" pageNumber="352">p</emphasis>
,
<emphasis id="B936EA8BFFA9745C85E13B0EFB3EF99F" box="[1218,1248,1622,1648]" italics="true" pageId="15" pageNumber="352">p”</emphasis>
,
<emphasis id="B936EA8BFFA9745C85CD3B0EFAD2F99F" box="[1262,1292,1622,1648]" italics="true" pageId="15" pageNumber="352">tc</emphasis>
,
<emphasis id="B936EA8BFFA9745C84393B0EFAE0F99F" box="[1306,1342,1622,1648]" italics="true" pageId="15" pageNumber="352">tc”</emphasis>
,
<emphasis id="B936EA8BFFA9745C846F3B0EFABBF99F" box="[1356,1381,1622,1648]" italics="true" pageId="15" pageNumber="352">ft</emphasis>
,
<emphasis id="B936EA8BFFA9745C84573B0EFA4DF99F" box="[1396,1427,1622,1648]" italics="true" pageId="15" pageNumber="352">ft”</emphasis>
) sparsely barbed. Setae
<emphasis id="B936EA8BFFA9745C80B73B22FE7BF97B" box="[404,421,1658,1684]" italics="true" pageId="15" pageNumber="352">l</emphasis>
,
<emphasis id="B936EA8BFFA9745C80913B22FE17F97B" box="[434,457,1658,1684]" italics="true" pageId="15" pageNumber="352">l”</emphasis>
of femur, (
<emphasis id="B936EA8BFFA9745C83603B22FD95F97B" box="[579,587,1658,1684]" italics="true" pageId="15" pageNumber="352">l</emphasis>
) of genu and
<emphasis id="B936EA8BFFA9745C83C13B22FD30F97B" box="[738,750,1658,1684]" italics="true" pageId="15" pageNumber="352">k</emphasis>
of tibiotarsus blunt-tipped; other leg setae (except eupathidia) pointed. Lengths of solenidia
<emphasis id="B936EA8BFFA9745C80C03BC6FDDAF957" box="[483,516,1694,1720]" italics="true" pageId="15" pageNumber="352">ω1</emphasis>
9 (710),
<emphasis id="B936EA8BFFA9745C835B3BC6FD47F957" box="[632,665,1694,1720]" italics="true" pageId="15" pageNumber="352">ω2</emphasis>
5 (35),
<emphasis id="B936EA8BFFA9745C83DD3BC6FCC5F957" box="[766,795,1694,1720]" italics="true" pageId="15" pageNumber="352">φ1</emphasis>
7 (78),
<emphasis id="B936EA8BFFA9745C82A23BC6FC41F957" box="[897,927,1694,1720]" italics="true" pageId="15" pageNumber="352">φ2</emphasis>
4 (45); solenidion
<emphasis id="B936EA8BFFA9745C85A33BC6FB7FF957" box="[1152,1185,1694,1720]" italics="true" pageId="15" pageNumber="352">ω1</emphasis>
digitiform; solenidion
<emphasis id="B936EA8BFFA9745C81B43B9AFF6AF933" box="[151,180,1730,1756]" italics="true" pageId="15" pageNumber="352">φ1</emphasis>
thick, clavate; other solenidia weakly clavate. Leg II (
<figureCitation id="13792A1CFFA9745C822A3B9AFCB0F933" box="[777,878,1730,1756]" captionStart="FIGURE 14" captionStartId="14.[151,250,1368,1392]" captionTargetBox="[151,1435,348,1344]" captionTargetId="figure-105@14.[151,1435,345,1344]" captionTargetPageId="14" captionText="FIGURE 14. Spatulaphorus venezuelaensis sp. nov., female: A—right leg I, dorsal aspect; B—right leg II, dorsal aspect." pageId="15" pageNumber="352">Fig. 14B</figureCitation>
). Tarsus with thickened basally claws and pad-like empodium. Solenidion
<emphasis id="B936EA8BFFA9745C80BE3BBFFE6FF8EF" box="[413,433,1767,1792]" italics="true" pageId="15" pageNumber="352">ω</emphasis>
5 (46) weakly clavate, solenidion
<emphasis id="B936EA8BFFA9745C821C3BBFFC90F8EF" box="[831,846,1767,1792]" italics="true" pageId="15" pageNumber="352">φ</emphasis>
absent. All setae barbed; seta
<emphasis id="B936EA8BFFA9745C85BD3BBEFB62F8EF" box="[1182,1212,1766,1792]" italics="true" pageId="15" pageNumber="352">tc</emphasis>
of tarsus spiniform, blunt-tipped and curved; setae
<emphasis id="B936EA8BFFA9745C80C83A52FE27F8CB" box="[491,505,1802,1828]" italics="true" pageId="15" pageNumber="352">d</emphasis>
,
<emphasis id="B936EA8BFFA9745C83253A52FDC9F8CB" box="[518,535,1802,1828]" italics="true" pageId="15" pageNumber="352">l</emphasis>
, and
<emphasis id="B936EA8BFFA9745C83713A52FDB0F8CB" box="[594,622,1802,1828]" italics="true" pageId="15" pageNumber="352">v”</emphasis>
of femur and
<emphasis id="B936EA8BFFA9745C822B3A52FCC7F8CB" box="[776,793,1802,1828]" italics="true" pageId="15" pageNumber="352">l</emphasis>
of genu blunt-tipped; other leg setae pointed. Leg III (
<figureCitation id="13792A1CFFA9745C844C3A52FF12F8A6" captionStart="FIGURE 15" captionStartId="15.[151,250,1443,1467]" captionTargetBox="[151,1435,189,1420]" captionTargetId="figure-17@15.[151,1435,181,1420]" captionTargetPageId="15" captionText="FIGURE 15. Spatulaphorus venezuelaensis sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." pageId="15" pageNumber="352">Fig. 15A</figureCitation>
). Femur divided into basi- and telofemur. Claws and empodium of same shape as on tarsus II. Seta
<emphasis id="B936EA8BFFA9745C843C3A76FAE8F8A7" box="[1311,1334,1838,1864]" italics="true" pageId="15" pageNumber="352">u</emphasis>
of tarsus smooth, other setae barbed; setae
<emphasis id="B936EA8BFFA9745C832F3A0AFDFCF883" box="[524,546,1874,1900]" italics="true" pageId="15" pageNumber="352">v</emphasis>
of trochanter,
<emphasis id="B936EA8BFFA9745C83E03A0AFD0FF883" box="[707,721,1874,1900]" italics="true" pageId="15" pageNumber="352">d</emphasis>
,
<emphasis id="B936EA8BFFA9745C83FD3A0AFD2AF883" box="[734,756,1874,1900]" italics="true" pageId="15" pageNumber="352">v</emphasis>
of femur, and
<emphasis id="B936EA8BFFA9745C82B73A0AFC7BF883" box="[916,933,1874,1900]" italics="true" pageId="15" pageNumber="352">l</emphasis>
of genu blunt-tipped, other leg setae pointed; solenidion
<emphasis id="B936EA8BFFA9745C80313A2FFEFFF87F" box="[274,289,1911,1936]" italics="true" pageId="15" pageNumber="352">φ</emphasis>
absent. Leg IV (
<figureCitation id="13792A1CFFA9745C80FA3A2EFDE6F87E" box="[473,568,1910,1937]" captionStart="FIGURE 15" captionStartId="15.[151,250,1443,1467]" captionTargetBox="[151,1435,189,1420]" captionTargetId="figure-17@15.[151,1435,181,1420]" captionTargetPageId="15" captionText="FIGURE 15. Spatulaphorus venezuelaensis sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." pageId="15" pageNumber="352">Fig 15B</figureCitation>
). Femur divided into basi- and telofemur. Claws simple, empodium as on tarsi II and III. All setae barbed; setae
<emphasis id="B936EA8BFFA9745C83293AC2FDFEF85B" box="[522,544,1946,1972]" italics="true" pageId="15" pageNumber="352">v</emphasis>
of trochanter,
<emphasis id="B936EA8BFFA9745C83E03AC2FD0FF85B" box="[707,721,1946,1972]" italics="true" pageId="15" pageNumber="352">d</emphasis>
,
<emphasis id="B936EA8BFFA9745C83FC3AC2FD2BF85B" box="[735,757,1946,1972]" italics="true" pageId="15" pageNumber="352">v</emphasis>
of femur,
<emphasis id="B936EA8BFFA9745C824A3AC2FC5AF85B" box="[873,900,1946,1972]" italics="true" pageId="15" pageNumber="352">v”</emphasis>
of tibia and
<emphasis id="B936EA8BFFA9745C85323AC2FBE9F85B" box="[1041,1079,1946,1972]" italics="true" pageId="15" pageNumber="352">pl”</emphasis>
of tarsus blunt-tipped, other leg setae pointed; solenidion
<emphasis id="B936EA8BFFA9745C80963AE7FE1AF837" box="[437,452,1983,2008]" italics="true" pageId="15" pageNumber="352">φ</emphasis>
absent.
</paragraph>
<caption id="DF3D6611FFB6744381B43E94FF29FBE7" pageId="16" pageNumber="353" startId="16.[151,250,972,996]" targetBox="[151,1436,181,948]" targetPageId="16" targetType="figure">
<paragraph id="8BFD3699FFB6744381B43E94FF29FBE7" blockId="16.[151,1436,972,1032]" pageId="16" pageNumber="353">
<emphasis id="B936EA8BFFB6744381B43E94FEC7FC0B" bold="true" box="[151,281,972,996]" pageId="16" pageNumber="353">FIGURE 16</emphasis>
. DIC micrographs of
<taxonomicName id="4C424D1AFFB6744380DB3E94FCF7FC0B" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[504,809,972,996]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="16" pageNumber="353" phylum="Arthropoda" rank="species" species="venezuelaensis" status="sp. nov.">
<emphasis id="B936EA8BFFB6744380DB3E94FCF7FC0B" box="[504,809,972,996]" italics="true" pageId="16" pageNumber="353">Spatulaphorus venezuelaensis</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFB67443820C3E94FCA1FC0B" bold="true" box="[815,895,972,996]" pageId="16" pageNumber="353">
<taxonomicNameLabel id="A20557F0FFB67443820C3E94FCA1FC0B" box="[815,895,972,996]" pageId="16" pageNumber="353" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—general view dorsally; B—general view ventrally.
</paragraph>
</caption>
<paragraph id="8BFD3699FFB6744381EB396FFE4FFBBF" blockId="16.[151,1437,1078,1537]" box="[200,401,1078,1105]" pageId="16" pageNumber="353">MALE unknown.</paragraph>
</subSubSection>
<subSubSection id="C3586512FFB6744381E43902FD9BFAC7" pageId="16" pageNumber="353" type="materials_examined">
<paragraph id="8BFD3699FFB6744381E43902FD04FB52" blockId="16.[151,1437,1078,1537]" pageId="16" pageNumber="353">
<emphasis id="B936EA8BFFB6744381E43902FEB0FB9B" bold="true" box="[199,366,1114,1140]" pageId="16" pageNumber="353">Type material</emphasis>
.
<materialsCitation id="3B2A3CC4FFB67443805E3902FD0CFB53" collectingDate="2001-04" collectingDateMax="2006-08-06" collectingDateMin="2001-04" collectionCode="ZISP, CEMT" country="Venezuela" location="Santa Barbara" municipality="Female" pageId="16" pageNumber="353" specimenCount="10" specimenCount-female="10" stateProvince="Zulia" typeStatus="holotype">
<collectingMunicipality id="6B99ACE3FFB67443805E3902FE11FB9B" box="[381,463,1114,1140]" pageId="16" pageNumber="353">Female</collectingMunicipality>
<typeStatus id="54F9883BFFB6744380F43902FDE2FB9B" box="[471,572,1114,1140]" pageId="16" pageNumber="353" type="holotype">holotype</typeStatus>
, slide
<collectionCode id="ED53AE5CFFB6744383A73902FD63FB9B" box="[644,701,1114,1140]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="16" pageNumber="353">ZISP</collectionCode>
T-Pygm-013,
<collectingCountry id="F3557609FFB67443827D3902FC0AFB9B" box="[862,980,1114,1140]" name="Venezuela" pageId="16" pageNumber="353">Venezuela</collectingCountry>
,
<collectingRegion id="4986F87BFFB6744382C33902FBC0FB9B" box="[992,1054,1114,1140]" country="Venezuela" name="Zulia" pageId="16" pageNumber="353">Zulia</collectingRegion>
,
<location id="8E9D6042FFB6744385093902FB17FB9B" LSID="urn:lsid:plazi:treatment:03EB878FFFAB744381B43F1EFB03F9EF:8E9D6042FFB6744385093902FB17FB9B" box="[1066,1225,1114,1140]" country="Venezuela" municipality="Female" name="Santa Barbara" pageId="16" pageNumber="353" stateProvince="Zulia">Santa Bárbara</location>
del
<collectingRegion id="4986F87BFFB6744385D93902FAE6FB9B" box="[1274,1336,1114,1140]" country="Venezuela" name="Zulia" pageId="16" pageNumber="353">Zulia</collectingRegion>
,
<date id="FFFC1059FFB6744384603902FA42FB9B" box="[1347,1436,1114,1140]" pageId="16" pageNumber="353" value="2001-04">
<collectingDate id="EFB8E9B1FFB6744384603902FA42FB9B" box="[1347,1436,1114,1140]" pageId="16" pageNumber="353" value="2001-04">IV.2001</collectingDate>
</date>
(
<collectionCode id="ED53AE5CFFB6744381BC3926FF2CFB77" box="[159,242,1150,1176]" pageId="16" pageNumber="353">CEMT</collectionCode>
),
<collectedFrom id="385E7564FFB6744380243927FD80FB77" box="[263,606,1150,1177]" pageId="16" pageNumber="353">
on
<taxonomicName id="4C424D1AFFB67443800F3926FD80FB77" authority="Harold" authorityName="Harold" authorityYear="1868" box="[300,606,1150,1176]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="353" phylum="Arthropoda" rank="species" species="prasinus">
<emphasis id="B936EA8BFFB67443800F3926FDDDFB77" box="[300,515,1150,1176]" italics="true" pageId="16" pageNumber="353">Phanaeus prasinus</emphasis>
Harold
</taxonomicName>
</collectedFrom>
;
<typeStatus id="54F9883BFFB6744383483927FD05FB76" box="[619,731,1151,1177]" pageId="16" pageNumber="353" type="paratype">paratypes</typeStatus>
:
<specimenCount id="9D44FD10FFB6744383C43927FC89FB77" box="[743,855,1150,1177]" count="7" pageId="16" pageNumber="353" type="female">7 females</specimenCount>
, same data;
<specimenCount id="9D44FD10FFB6744382C53926FB88FB77" box="[998,1110,1150,1176]" count="3" pageId="16" pageNumber="353" type="female">3 females</specimenCount>
,
<collectingCountry id="F3557609FFB6744385403926FB07FB77" box="[1123,1241,1150,1176]" name="Venezuela" pageId="16" pageNumber="353">Venezuela</collectingCountry>
, Boliar, Anacoco Edo,
<date id="FFFC1059FFB6744381F139FAFEBBFB53" box="[210,357,1186,1212]" pageId="16" pageNumber="353" value="2006-08-06">
<collectingDate id="EFB8E9B1FFB6744381F139FAFEBBFB53" box="[210,357,1186,1212]" pageId="16" pageNumber="353" value="2006-08-06">06.VIII.2006</collectingDate>
</date>
(
<collectionCode id="ED53AE5CFFB67443805039FAFE18FB53" box="[371,454,1186,1212]" pageId="16" pageNumber="353">CEMT</collectionCode>
),
<collectedFrom id="385E7564FFB6744380F939FBFD0CFB53" box="[474,722,1186,1213]" pageId="16" pageNumber="353">
on
<taxonomicName id="4C424D1AFFB6744380DE39FAFD0CFB53" authorityName="Harold" authorityYear="1868" box="[509,722,1186,1212]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="353" phylum="Arthropoda" rank="species" species="prasinus">
<emphasis id="B936EA8BFFB6744380DE39FAFD0CFB53" box="[509,722,1186,1212]" italics="true" pageId="16" pageNumber="353">Phanaeus prasinus</emphasis>
</taxonomicName>
</collectedFrom>
</materialsCitation>
.
</paragraph>
<paragraph id="8BFD3699FFB6744381E4399EFD9BFAC7" blockId="16.[151,1437,1078,1537]" pageId="16" pageNumber="353">
<emphasis id="B936EA8BFFB6744381E4399EFE58FB0F" bold="true" box="[199,390,1222,1248]" pageId="16" pageNumber="353">Type deposition</emphasis>
.
<materialsCitation id="3B2A3CC4FFB6744380B6399EFE25FAEB" country="Russia" location="Zoological Institute" pageId="16" pageNumber="353" specimenCount="3" stateProvince="Saint Petersburg" typeStatus="holotype">
The
<typeStatus id="54F9883BFFB6744380E9399EFDF2FB0F" box="[458,556,1222,1248]" pageId="16" pageNumber="353" type="holotype">holotype</typeStatus>
and
<specimenCount id="9D44FD10FFB674438344399EFCCDFB0E" box="[615,787,1222,1249]" count="3" pageId="16" pageNumber="353" type="generic" typeStatus="paratype">three paratypes</specimenCount>
are deposited in the collection of
<location id="8E9D6042FFB6744385BD399EFAA2FB0F" LSID="urn:lsid:plazi:treatment:03EB878FFFAB744381B43F1EFB03F9EF:8E9D6042FFB6744385BD399EFAA2FB0F" box="[1182,1404,1222,1248]" country="Russia" name="Zoological Institute" pageId="16" pageNumber="353" stateProvince="Saint Petersburg">Zoological Institute</location>
of RAS,
<collectingRegion id="4986F87BFFB6744381FC39B2FE40FAEB" box="[223,414,1258,1284]" country="Russia" name="Saint Petersburg" pageId="16" pageNumber="353">Saint Petersburg</collectingRegion>
,
<collectingCountry id="F3557609FFB67443808E39B2FE25FAEB" box="[429,507,1258,1284]" name="Russia" pageId="16" pageNumber="353">Russia</collectingCountry>
</materialsCitation>
;
<materialsCitation id="3B2A3CC4FFB67443832939B2FAC5FAEB" box="[522,1307,1258,1285]" country="Russia" location="Tyumen State" pageId="16" pageNumber="353" specimenCount="1" stateProvince="Tyumen'" typeStatus="paratype">
other
<typeStatus id="54F9883BFFB67443836D39B3FD67FAEA" box="[590,697,1259,1285]" pageId="16" pageNumber="353" type="paratype">paratypes</typeStatus>
are deposited in the collection of the
<collectingRegion id="4986F87BFFB67443855E39B3FAC5FAEB" box="[1149,1307,1258,1285]" country="Russia" name="Tyumen'" pageId="16" pageNumber="353">Tyumen State</collectingRegion>
</materialsCitation>
<materialsCitation id="3B2A3CC4FFB67443840539B2FD9EFAC7" country="Russia" location="University Museum of Zoology" pageId="16" pageNumber="353" specimenCount="1" stateProvince="Tyumen'" typeStatus="paratype">
<location id="8E9D6042FFB67443840539B2FEA1FAC7" LSID="urn:lsid:plazi:treatment:03EB878FFFAB744381B43F1EFB03F9EF:8E9D6042FFB67443840539B2FEA1FAC7" country="Russia" name="University Museum of Zoology" pageId="16" pageNumber="353" stateProvince="Tyumen'">University Museum of Zoology</location>
,
<collectingRegion id="4986F87BFFB6744380AA3857FE39FAC6" box="[393,487,1295,1321]" country="Russia" name="Tyumen'" pageId="16" pageNumber="353">Tyumen</collectingRegion>
,
<collectingCountry id="F3557609FFB6744380D13856FD9EFAC7" box="[498,576,1294,1320]" name="Russia" pageId="16" pageNumber="353">Russia</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3586512FFB6744381E4386AFD1BFA33" pageId="16" pageNumber="353" type="diagnosis">
<paragraph id="8BFD3699FFB6744381E4386AFD1BFA33" blockId="16.[151,1437,1078,1537]" pageId="16" pageNumber="353">
<emphasis id="B936EA8BFFB6744381E4386AFE16FAA3" bold="true" box="[199,456,1330,1356]" pageId="16" pageNumber="353">Differential diagnosis</emphasis>
. The new species is very similar to
<taxonomicName id="4C424D1AFFB674438248386BFAFAFAA3" authority="Khaustov and Frolov, 2021" authorityName="Khaustov and Frolov" authorityYear="2021" box="[875,1316,1330,1357]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="16" pageNumber="353" phylum="Arthropoda" rank="species" species="porosus">
<emphasis id="B936EA8BFFB674438248386BFCA7FAA2" box="[875,889,1331,1357]" italics="true" pageId="16" pageNumber="353">S</emphasis>
.
<emphasis id="B936EA8BFFB6744382AA386BFC3FFAA3" box="[905,993,1331,1356]" italics="true" pageId="16" pageNumber="353">porosus</emphasis>
<bibRefCitation id="EFD34B68FFB6744382C9386AFAFAFAA3" author="Khaustov, A. A. &amp; Frolov, A. V." box="[1002,1316,1330,1356]" pageId="16" pageNumber="353" pagination="333 - 345" refId="ref16337" refString="Khaustov, A. A. &amp; Frolov, A. V. (2021) Two new species and a new record of Spatulaphorus (Acari &quot; Pygmephoridae) phoretic on scarab beetles (Coleoptera: Scarabaeidae) from French Guiana. Acarina, 29 (2), 333 - 345. https: // doi. org / 10.21684 / 0132 - 8077 - 2021 - 29 - 2 - 233 - 245" type="journal article" year="2021">Khaustov and Frolov, 2021</bibRefCitation>
</taxonomicName>
, described from
<collectingCountry id="F3557609FFB6744381F4380EFEA0FA9F" box="[215,382,1366,1392]" name="French Guiana" pageId="16" pageNumber="353">French Guiana</collectingCountry>
(
<bibRefCitation id="EFD34B68FFB6744380AC380EFD6FFA9F" author="Khaustov, A. A. &amp; Frolov, A. V." box="[399,689,1366,1392]" pageId="16" pageNumber="353" pagination="333 - 345" refId="ref16337" refString="Khaustov, A. A. &amp; Frolov, A. V. (2021) Two new species and a new record of Spatulaphorus (Acari &quot; Pygmephoridae) phoretic on scarab beetles (Coleoptera: Scarabaeidae) from French Guiana. Acarina, 29 (2), 333 - 345. https: // doi. org / 10.21684 / 0132 - 8077 - 2021 - 29 - 2 - 233 - 245" type="journal article" year="2021">Khaustov &amp; Frolov 2021</bibRefCitation>
) in having spiniform seta
<emphasis id="B936EA8BFFB6744382C2380FFC21FA9E" box="[993,1023,1367,1393]" italics="true" pageId="16" pageNumber="353">tc</emphasis>
on tarsus II, presence of big puncta on dorsal and ventral idiosomal sclerites, enlarged tibiotarsus I and thickened basally claws on tarsi II and III. The new species differs from
<taxonomicName id="4C424D1AFFB67443809738C7FDF6FA57" authorityName="Khaustov and Frolov" authorityYear="2021" box="[436,552,1439,1465]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="16" pageNumber="353" phylum="Arthropoda" rank="species" species="porosus">
<emphasis id="B936EA8BFFB67443809738C7FE1CFA56" box="[436,450,1439,1465]" italics="true" pageId="16" pageNumber="353">S</emphasis>
.
<emphasis id="B936EA8BFFB6744380F338C7FDF6FA57" box="[464,552,1439,1464]" italics="true" pageId="16" pageNumber="353">porosus</emphasis>
</taxonomicName>
in having setae
<emphasis id="B936EA8BFFB6744383C338C7FD32FA57" box="[736,748,1439,1464]" italics="true" pageId="16" pageNumber="353">e</emphasis>
situated slightly anteriad bases of setae
<emphasis id="B936EA8BFFB67443859138C6FB64FA57" box="[1202,1210,1438,1464]" italics="true" pageId="16" pageNumber="353">f</emphasis>
(vs. setae
<emphasis id="B936EA8BFFB67443841238C7FAE3FA57" box="[1329,1341,1439,1464]" italics="true" pageId="16" pageNumber="353">e</emphasis>
situated distinctly posteriad bases of setae
<emphasis id="B936EA8BFFB674438336389AFDC3FA33" box="[533,541,1474,1500]" italics="true" pageId="16" pageNumber="353">f</emphasis>
in
<taxonomicName id="4C424D1AFFB674438362389BFD6BFA33" authorityName="Khaustov and Frolov" authorityYear="2021" box="[577,693,1475,1501]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="16" pageNumber="353" phylum="Arthropoda" rank="species" species="porosus">
<emphasis id="B936EA8BFFB674438362389BFD6BFA33" box="[577,693,1475,1501]" italics="true" pageId="16" pageNumber="353">S. porosus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3586512FFB6744381E438BEFB03F9EF" box="[199,1245,1510,1537]" pageId="16" pageNumber="353" type="etymology">
<paragraph id="8BFD3699FFB6744381E438BEFB03F9EF" blockId="16.[151,1437,1078,1537]" box="[199,1245,1510,1537]" pageId="16" pageNumber="353">
<emphasis id="B936EA8BFFB6744381E438BEFE96F9EF" bold="true" box="[199,328,1510,1536]" pageId="16" pageNumber="353">Etymology</emphasis>
. The name of the new species refers to its geographical distribution in
<collectingCountry id="F3557609FFB67443854138BEFB07F9EF" box="[1122,1241,1510,1536]" name="Venezuela" pageId="16" pageNumber="353">Venezuela</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="1227CC38B685C2552C2422E437FE9BB3" ID-DOI="10.11646/zootaxa.5497.3.2" ID-ISSN="1175-5326" ID-ZooBank="C107D617-5F70-42C5-8FD4-E5C2DE0E48B1" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725026357643" checkinUser="plazi" docAuthor="Khaustov, Alexander A., Frolov, Andrey V. &amp; Akhmetova, Lilia A." docDate="2024" docId="03EB878FFFAF745E81B43CEEFB9DFE1A" docLanguage="en" docName="zootaxa.5497.3.2.pdf" docOrigin="Zootaxa 5497 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5497.3.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Spatulaphorus spinisetus Khaustov, Frolov &amp; Akhmetova, 2024, sp. nov." docType="treatment" docVersion="1" lastPageNumber="350" masterDocId="FFD2FFF7FFA6745381233D58FFDEFFEF" masterDocTitle="Seven new species of Spatulaphorus Rack (Acari: Pygmephoridae) phoretic on scarab beetles (Coleoptera: Scarabaeidae)" masterLastPageNumber="368" masterPageNumber="337" pageNumber="346" updateTime="1725027619610" updateUser="GgImagineBatch" zenodo-license-document="CLOSED">
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<mods:title id="0655BCF18706DF77F740B829F2221C3F">Seven new species of Spatulaphorus Rack (Acari: Pygmephoridae) phoretic on scarab beetles (Coleoptera: Scarabaeidae)</mods:title>
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<mods:namePart id="F43D872B8309E8D6374447713D84A5B4">Khaustov, Alexander A.</mods:namePart>
<mods:affiliation id="0092C83B43B7A8CA3ABAF258ABC07732">Tyumen State University, Tyumen, 6 Volodarskogo Str., 625003 Russia.</mods:affiliation>
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<mods:namePart id="4DEDA5BA7CDCE880F315775DB277CE6C">Frolov, Andrey V.</mods:namePart>
<mods:affiliation id="CA1C24B8B579DD814E126E491C02859C">Zoological Institute of Russian Academy of Sciences, Universitetskaya Embankment 1, 199034, Saint Petersburg, Russia</mods:affiliation>
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<mods:namePart id="198A3B7A65F012DFEF4F89BD9CD646CB">Akhmetova, Lilia A.</mods:namePart>
<mods:affiliation id="F8811FD8FA6ABA0EE438037A9C215000">Zoological Institute of Russian Academy of Sciences, Universitetskaya Embankment 1, 199034, Saint Petersburg, Russia</mods:affiliation>
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<mods:title id="8E209AB492FB452183DC1DDD12BCC892">Zootaxa</mods:title>
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<mods:date id="BF9717A5D50812A4F10449B516D78E61">2024</mods:date>
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<mods:number id="44371874A0191B1E1DD0AD0413EBD0A9">2024-08-26</mods:number>
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<mods:number id="6F7CC63DD4AFADC0B73FB9828766FCA3">5497</mods:number>
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<subSubSection id="C3586512FFAF745A81B43CEEFDF2FE3E" box="[151,556,438,465]" pageId="9" pageNumber="346" type="nomenclature">
<paragraph id="8BFD3699FFAF745A81B43CEEFDF2FE3E" blockId="9.[151,556,438,501]" box="[151,556,438,465]" pageId="9" pageNumber="346">
<heading id="D0B581F5FFAF745A81B43CEEFDF2FE3E" bold="true" box="[151,556,438,465]" fontSize="11" level="1" pageId="9" pageNumber="346" reason="1">
<emphasis id="B936EA8BFFAF745A81B43CEEFDF2FE3E" bold="true" box="[151,556,438,465]" pageId="9" pageNumber="346">
<taxonomicName id="4C424D1AFFAF745A81B43CEEFE19FE3E" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[151,455,438,465]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="346" phylum="Arthropoda" rank="species" species="spinisetus" status="sp. nov.">
<emphasis id="B936EA8BFFAF745A81B43CEEFE19FE3E" bold="true" box="[151,455,438,465]" italics="true" pageId="9" pageNumber="346">Spatulaphorus spinisetus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20557F0FFAF745A80EC3CEFFDF2FE3E" box="[463,556,439,465]" pageId="9" pageNumber="346" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3586512FFAF745F81B43C83FE4FF9FC" lastPageId="12" lastPageNumber="349" pageId="9" pageNumber="346" type="description">
<paragraph id="8BFD3699FFAF745A81B43C83FEC7FE1A" blockId="9.[151,556,438,501]" box="[151,281,475,501]" pageId="9" pageNumber="346">
(
<figureCitation id="13792A1CFFAF745A81BC3C83FECFFE1A" box="[159,273,475,501]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStart-2="FIGURE 11" captionStart-3="FIGURE 12" captionStartId-0="9.[151,250,1795,1819]" captionStartId-1="10.[151,250,1311,1335]" captionStartId-2="11.[151,250,1860,1884]" captionStartId-3="12.[151,250,1422,1446]" captionTargetBox-0="[151,1435,610,1772]" captionTargetBox-1="[151,1435,241,1288]" captionTargetBox-2="[151,1418,190,1836]" captionTargetBox-3="[151,1436,633,1399]" captionTargetId-0="figure-146@9.[151,1435,597,1772]" captionTargetId-1="figure-50@10.[151,1435,237,1288]" captionTargetId-2="figure-17@11.[151,1436,181,1836]" captionTargetId-3="figure-287@12.[151,1436,633,1399]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="12" captionText-0="FIGURE 9. Spatulaphorus spinisetus sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." captionText-1="FIGURE 10. Spatulaphorus spinisetus sp. nov., female: A—right leg I, dorsal aspect; B—right leg II, dorsal aspect." captionText-2="FIGURE 11. Spatulaphorus spinisetus sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." captionText-3="FIGURE 12. DIC micrographs of Spatulaphorus spinisetus sp. nov., female: A—general view dorsally; B—general view ventrally." pageId="9" pageNumber="346">Figs 912</figureCitation>
)
</paragraph>
<paragraph id="8BFD3699FFAF745A81B43F7AFAE1FDD2" blockId="9.[151,1343,546,573]" box="[151,1343,546,573]" pageId="9" pageNumber="346">
<emphasis id="B936EA8BFFAF745A81B43F7AFEFFFDD3" bold="true" box="[151,289,546,572]" pageId="9" pageNumber="346">Description</emphasis>
. FEMALE. Body well sclerotized. Length of idiosoma 245 (190265), width 155 (105155).
</paragraph>
<caption id="DF3D6611FFAF745A81B43A5BFB25F8F4" box="[151,1275,1795,1820]" pageId="9" pageNumber="346" startId="9.[151,250,1795,1819]" targetBox="[151,1435,610,1772]" targetPageId="9" targetType="figure">
<paragraph id="8BFD3699FFAF745A81B43A5BFB25F8F4" blockId="9.[151,1275,1795,1820]" box="[151,1275,1795,1820]" pageId="9" pageNumber="346">
<emphasis id="B936EA8BFFAF745A81B43A5BFED3F8F4" bold="true" box="[151,269,1795,1819]" pageId="9" pageNumber="346">FIGURE 9</emphasis>
.
<taxonomicName id="4C424D1AFFAF745A803A3A5BFDC6F8F4" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[281,536,1795,1819]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="346" phylum="Arthropoda" rank="species" species="spinisetus" status="sp. nov.">
<emphasis id="B936EA8BFFAF745A803A3A5BFDC6F8F4" box="[281,536,1795,1819]" italics="true" pageId="9" pageNumber="346">Spatulaphorus spinisetus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFAF745A833D3A5BFDB0F8F4" bold="true" box="[542,622,1795,1819]" pageId="9" pageNumber="346">
<taxonomicNameLabel id="A20557F0FFAF745A833D3A5BFDB0F8F4" box="[542,622,1795,1819]" pageId="9" pageNumber="346" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—dorsum of body; B—venter of body. Legs omitted.
</paragraph>
</caption>
<paragraph id="8BFD3699FFAF745981E43A11FCC9FF3A" blockId="9.[151,1437,1865,2036]" lastBlockId="10.[151,1436,151,213]" lastPageId="10" lastPageNumber="347" pageId="9" pageNumber="346">
<emphasis id="B936EA8BFFAF745A81E43A11FE88F88C" box="[199,342,1865,1891]" italics="true" pageId="9" pageNumber="346">Gnathosoma</emphasis>
. Gnathosomal capsule, excluding palps, subrectangular, length 21 (1922), width 22 (1922). Dorsum with one pair of needle-like cheliceral setae (
<emphasis id="B936EA8BFFAF745A823D3A35FC98F868" box="[798,838,1901,1927]" italics="true" pageId="9" pageNumber="346">cha</emphasis>
) 6 (56). Dorsal median apodeme present, poorly developed. Postpalpal setae (
<emphasis id="B936EA8BFFAF745A80FC3ACAFE25F844" box="[479,507,1938,1963]" italics="true" pageId="9" pageNumber="346">pp</emphasis>
) 2 (2) needle-like, situated laterad setae
<emphasis id="B936EA8BFFAF745A82EA3AC9FC2FF844" box="[969,1009,1937,1963]" italics="true" pageId="9" pageNumber="346">cha</emphasis>
. Ventral gnathosoma with one pair of smooth, weakly blunt-tipped subcapitular setae
<emphasis id="B936EA8BFFAF745A838D3AEEFD1CF820" box="[686,706,1974,1999]" italics="true" pageId="9" pageNumber="346">m</emphasis>
6 (67) and round pits
<emphasis id="B936EA8BFFAF745A82E93AEEFC06F820" box="[970,984,1974,1999]" italics="true" pageId="9" pageNumber="346">n</emphasis>
situated posteriad
<emphasis id="B936EA8BFFAF745A85883AEEFB61F820" box="[1195,1215,1974,1999]" italics="true" pageId="9" pageNumber="346">m</emphasis>
. Palps with smooth and weakly blunt-tipped setae
<emphasis id="B936EA8BFFAF745A80C93A81FDC8F81C" box="[490,534,2009,2035]" italics="true" pageId="9" pageNumber="346">dFe</emphasis>
4 (34) and
<emphasis id="B936EA8BFFAF745A83813A81FD0FF81C" box="[674,721,2009,2035]" italics="true" pageId="9" pageNumber="346">dGe</emphasis>
8 (78) dorsolaterally. Palps ventrally each with mushroom-like accessory setigenous structure (
<emphasis id="B936EA8BFFAC745980DA3DCFFDC3FF5F" box="[505,541,151,176]" italics="true" pageId="10" pageNumber="347">ass</emphasis>
) and tiny solenidion
<emphasis id="B936EA8BFFAC745982293DCFFCC0FF5F" box="[778,798,151,176]" italics="true" pageId="10" pageNumber="347">ω</emphasis>
. Palps terminated with small blunt-tipped tibial claw and tiny eupathid-like seta. Pharyngeal pumps poorly visible.
</paragraph>
<caption id="DF3D6611FFAC745981B43847FAEAFAD8" box="[151,1332,1311,1335]" pageId="10" pageNumber="347" startId="10.[151,250,1311,1335]" targetBox="[151,1435,241,1288]" targetPageId="10" targetType="figure">
<paragraph id="8BFD3699FFAC745981B43847FAEAFAD8" blockId="10.[151,1332,1311,1335]" box="[151,1332,1311,1335]" pageId="10" pageNumber="347">
<emphasis id="B936EA8BFFAC745981B43847FEC7FAD8" bold="true" box="[151,281,1311,1335]" pageId="10" pageNumber="347">FIGURE 10</emphasis>
.
<taxonomicName id="4C424D1AFFAC745980053847FDFAFAD8" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[294,548,1311,1335]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="347" phylum="Arthropoda" rank="species" species="spinisetus" status="sp. nov.">
<emphasis id="B936EA8BFFAC745980053847FDFAFAD8" box="[294,548,1311,1335]" italics="true" pageId="10" pageNumber="347">Spatulaphorus spinisetus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFAC745983093847FDA5FAD8" bold="true" box="[554,635,1311,1335]" pageId="10" pageNumber="347">
<taxonomicNameLabel id="A20557F0FFAC745983093847FDA5FAD8" box="[554,635,1311,1335]" pageId="10" pageNumber="347" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—right leg I, dorsal aspect; B—right leg II, dorsal aspect.
</paragraph>
</caption>
<paragraph id="8BFD3699FFAC745981E4383DFC95F994" blockId="10.[151,1437,1381,1984]" pageId="10" pageNumber="347">
<emphasis id="B936EA8BFFAC745981E4383DFE48FA90" box="[199,406,1381,1407]" italics="true" pageId="10" pageNumber="347">Idiosomal dorsum</emphasis>
(
<figureCitation id="13792A1CFFAC7459808A383DFDD4FA90" box="[425,522,1381,1407]" captionStart="FIGURE 9" captionStartId="9.[151,250,1795,1819]" captionTargetBox="[151,1435,610,1772]" captionTargetId="figure-146@9.[151,1435,597,1772]" captionTargetPageId="9" captionText="FIGURE 9. Spatulaphorus spinisetus sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." pageId="10" pageNumber="347">Figs 9A</figureCitation>
,
<figureCitation id="13792A1CFFAC7459833A383DFD93FA90" box="[537,589,1381,1407]" captionStart="FIGURE 12" captionStartId="12.[151,250,1422,1446]" captionTargetBox="[151,1436,633,1399]" captionTargetId="figure-287@12.[151,1436,633,1399]" captionTargetPageId="12" captionText="FIGURE 12. DIC micrographs of Spatulaphorus spinisetus sp. nov., female: A—general view dorsally; B—general view ventrally." pageId="10" pageNumber="347">12A</figureCitation>
). All dorsal shields with big puncta. Prodorsal shield with three pairs of setae (
<emphasis id="B936EA8BFFAC745981C338D1FF24FA4C" box="[224,250,1417,1443]" italics="true" pageId="10" pageNumber="347">v1</emphasis>
,
<emphasis id="B936EA8BFFAC7459802A38D1FEFDFA4C" box="[265,291,1417,1443]" italics="true" pageId="10" pageNumber="347">v2</emphasis>
,
<emphasis id="B936EA8BFFAC7459801238D1FE89FA4C" box="[305,343,1417,1443]" italics="true" pageId="10" pageNumber="347">sc2</emphasis>
), one pair of weakly barbed capitate trichobothria (
<emphasis id="B936EA8BFFAC745982B838D1FC1FFA4C" box="[923,961,1417,1443]" italics="true" pageId="10" pageNumber="347">sc1</emphasis>
) and one pair of round stigmata. Stigmata associated with well-developed tracheal trunks.All dorsal idiosomal setae blunt-tipped; seta
<emphasis id="B936EA8BFFAC745985BA38F5FB6DFA28" box="[1177,1203,1453,1479]" italics="true" pageId="10" pageNumber="347">v2</emphasis>
smooth, other dorsal setae distinctly barbed.Alveoli of setae
<emphasis id="B936EA8BFFAC7459836D388AFD84FA04" box="[590,602,1490,1515]" italics="true" pageId="10" pageNumber="347">e</emphasis>
and
<emphasis id="B936EA8BFFAC745983AC3889FD49FA04" box="[655,663,1489,1515]" italics="true" pageId="10" pageNumber="347">f</emphasis>
almost contiguous. Cupules
<emphasis id="B936EA8BFFAC745982F6388AFC35FA04" box="[981,1003,1490,1515]" italics="true" pageId="10" pageNumber="347">ia</emphasis>
on tergite D,
<emphasis id="B936EA8BFFAC745985A3388AFB42FA04" box="[1152,1180,1490,1515]" italics="true" pageId="10" pageNumber="347">im</emphasis>
on tergite EF and
<emphasis id="B936EA8BFFAC745984463889FAA5FA04" box="[1381,1403,1489,1515]" italics="true" pageId="10" pageNumber="347">ih</emphasis>
on tergite H large, round, subequal. Lengths of dorsal setae:
<emphasis id="B936EA8BFFAC7459823138ADFCF2F9E0" box="[786,812,1525,1551]" italics="true" pageId="10" pageNumber="347">v1</emphasis>
26 (2127),
<emphasis id="B936EA8BFFAC7459829838ADFC08F9E0" box="[955,982,1525,1551]" italics="true" pageId="10" pageNumber="347">v2</emphasis>
9 (810),
<emphasis id="B936EA8BFFAC7459856B38ADFBB0F9E0" box="[1096,1134,1525,1551]" italics="true" pageId="10" pageNumber="347">sc2</emphasis>
47 (3849),
<emphasis id="B936EA8BFFAC745985DF38ADFAC9F9E0" box="[1276,1303,1525,1551]" italics="true" pageId="10" pageNumber="347">c1</emphasis>
32 (2734),
<emphasis id="B936EA8BFFAC745981B43B41FF6CF9DC" box="[151,178,1561,1587]" italics="true" pageId="10" pageNumber="347">c2</emphasis>
45 (3648),
<emphasis id="B936EA8BFFAC745980663B41FE8DF9DC" box="[325,339,1561,1587]" italics="true" pageId="10" pageNumber="347">d</emphasis>
39 (3039),
<emphasis id="B936EA8BFFAC745980C43B42FE2DF9DC" box="[487,499,1562,1587]" italics="true" pageId="10" pageNumber="347">e</emphasis>
24 (2027),
<emphasis id="B936EA8BFFAC745983A43B41FD51F9DC" box="[647,655,1561,1587]" italics="true" pageId="10" pageNumber="347">f</emphasis>
41 (3242),
<emphasis id="B936EA8BFFAC745982003B41FCE0F9DC" box="[803,830,1561,1587]" italics="true" pageId="10" pageNumber="347">h1</emphasis>
43 (3444),
<emphasis id="B936EA8BFFAC745982F03B41FC31F9DC" box="[979,1007,1561,1587]" italics="true" pageId="10" pageNumber="347">h2</emphasis>
24 (1924). Distances between setae:
<emphasis id="B936EA8BFFAC745981B43B65FF04F9B8" box="[151,218,1597,1624]" italics="true" pageId="10" pageNumber="347">v1v1</emphasis>
27 (2429),
<emphasis id="B936EA8BFFAC745980533B65FE6CF9B8" box="[368,434,1597,1623]" italics="true" pageId="10" pageNumber="347">v2v2</emphasis>
45 (4049),
<emphasis id="B936EA8BFFAC7459836A3B65FD7FF9B8" box="[585,673,1597,1623]" italics="true" pageId="10" pageNumber="347">sc2sc2</emphasis>
56 (4661),
<emphasis id="B936EA8BFFAC745982143B65FCA4F9B8" box="[823,890,1597,1624]" italics="true" pageId="10" pageNumber="347">c1c1</emphasis>
44 (3547),
<emphasis id="B936EA8BFFAC7459852D3B65FB8FF9B8" box="[1038,1105,1597,1624]" italics="true" pageId="10" pageNumber="347">c1c2</emphasis>
32 (2432),
<emphasis id="B936EA8BFFAC745985C43B65FACFF9B8" box="[1255,1297,1597,1624]" italics="true" pageId="10" pageNumber="347">dd</emphasis>
76 (6580),
<emphasis id="B936EA8BFFAC745981B43B3AFF64F994" box="[151,186,1633,1660]" italics="true" pageId="10" pageNumber="347">ef</emphasis>
5 (45),
<emphasis id="B936EA8BFFAC745980033B39FEE0F994" box="[288,318,1633,1660]" italics="true" pageId="10" pageNumber="347">ff</emphasis>
73 (5875),
<emphasis id="B936EA8BFFAC745980ED3B39FDCAF994" box="[462,532,1633,1660]" italics="true" pageId="10" pageNumber="347">h1h1</emphasis>
62 (4762),
<emphasis id="B936EA8BFFAC745983863B39FD35F994" box="[677,747,1633,1660]" italics="true" pageId="10" pageNumber="347">h1h2</emphasis>
7 (78).
</paragraph>
<paragraph id="8BFD3699FFAC745981E43BDDFE3DF82F" blockId="10.[151,1437,1381,1984]" pageId="10" pageNumber="347">
<emphasis id="B936EA8BFFAC745981E43BDDFE57F970" box="[199,393,1669,1695]" italics="true" pageId="10" pageNumber="347">Idiosomal venter</emphasis>
(
<figureCitation id="13792A1CFFAC745980BF3BDDFE22F970" box="[412,508,1669,1695]" captionStart="FIGURE 9" captionStartId="9.[151,250,1795,1819]" captionTargetBox="[151,1435,610,1772]" captionTargetId="figure-146@9.[151,1435,597,1772]" captionTargetPageId="9" captionText="FIGURE 9. Spatulaphorus spinisetus sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." pageId="10" pageNumber="347">Figs 9B</figureCitation>
,
<figureCitation id="13792A1CFFAC7459832E3BDDFDE1F970" box="[525,575,1669,1695]" captionStart="FIGURE 12" captionStartId="12.[151,250,1422,1446]" captionTargetBox="[151,1436,633,1399]" captionTargetId="figure-287@12.[151,1436,633,1399]" captionTargetPageId="12" captionText="FIGURE 12. DIC micrographs of Spatulaphorus spinisetus sp. nov., female: A—general view dorsally; B—general view ventrally." pageId="10" pageNumber="347">12B</figureCitation>
). Coxisternal fields I-II, III-IV and aggenital plate laterally with sparsely distributed big puncta. Setae
<emphasis id="B936EA8BFFAC745980C53BF1FDDCF92C" box="[486,514,1705,1731]" italics="true" pageId="10" pageNumber="347">1b</emphasis>
pointed; other ventral setae blunt-tipped; setae
<emphasis id="B936EA8BFFAC745985063BF1FB9CF92C" box="[1061,1090,1705,1731]" italics="true" pageId="10" pageNumber="347">1b</emphasis>
and
<emphasis id="B936EA8BFFAC7459855F3BF1FB7DF92C" box="[1148,1187,1705,1731]" italics="true" pageId="10" pageNumber="347">ps3</emphasis>
weakly barbed, other ventral setae smooth. Bases of setae
<emphasis id="B936EA8BFFAC745983053B95FD93F908" box="[550,589,1741,1767]" italics="true" pageId="10" pageNumber="347">ps3</emphasis>
situated distinctly anteriad bases of setae
<emphasis id="B936EA8BFFAC745985353B95FB82F908" box="[1046,1116,1741,1767]" italics="true" pageId="10" pageNumber="347">ps1-2.</emphasis>
Ap1 and ap2 well-developed and joined with thick appr; apsej developed only laterally; ap3 weak, not reaching appo; ap4 well developed, exceeding beyond bases of setae
<emphasis id="B936EA8BFFAC745983323A4DFDF3F8C0" box="[529,557,1813,1839]" italics="true" pageId="10" pageNumber="347">3b</emphasis>
. Ap5 short, joined with appo and reaching bases of setae
<emphasis id="B936EA8BFFAC7459859C3A4DFB02F8C0" box="[1215,1244,1813,1839]" italics="true" pageId="10" pageNumber="347">4a</emphasis>
. Ags small, cup-like; pgs small, oval. Posterior margin of posterior sternal plate straight in middle part. Posterior margin of aggenital plate weakly concave. Lengths of ventral setae:
<emphasis id="B936EA8BFFAC745983983A05FD09F898" box="[699,727,1885,1911]" italics="true" pageId="10" pageNumber="347">1a</emphasis>
9 (810),
<emphasis id="B936EA8BFFAC7459826C3A05FCB2F898" box="[847,876,1885,1911]" italics="true" pageId="10" pageNumber="347">1b</emphasis>
20 (1722),
<emphasis id="B936EA8BFFAC745985233A05FBC4F898" box="[1024,1050,1885,1911]" italics="true" pageId="10" pageNumber="347">1с</emphasis>
11 (1012),
<emphasis id="B936EA8BFFAC7459858D3A05FB14F898" box="[1198,1226,1885,1911]" italics="true" pageId="10" pageNumber="347">2a</emphasis>
12 (1114),
<emphasis id="B936EA8BFFAC7459847E3A05FAA9F898" box="[1373,1399,1885,1911]" italics="true" pageId="10" pageNumber="347">2с</emphasis>
13 (1113),
<emphasis id="B936EA8BFFAC745981DE3AD9FEC7F874" box="[253,281,1921,1947]" italics="true" pageId="10" pageNumber="347">3a</emphasis>
14 (1114),
<emphasis id="B936EA8BFFAC7459808A3AD9FE1BF874" box="[425,453,1921,1947]" italics="true" pageId="10" pageNumber="347">3b</emphasis>
12 (1013),
<emphasis id="B936EA8BFFAC745983753AD9FDAEF874" box="[598,624,1921,1947]" italics="true" pageId="10" pageNumber="347">3c</emphasis>
16 (1216),
<emphasis id="B936EA8BFFAC745982213AD9FCC0F874" box="[770,798,1921,1947]" italics="true" pageId="10" pageNumber="347">4a</emphasis>
12 (1012),
<emphasis id="B936EA8BFFAC7459828C3AD9FC15F874" box="[943,971,1921,1947]" italics="true" pageId="10" pageNumber="347">4b</emphasis>
19 (1520),
<emphasis id="B936EA8BFFAC7459857F3AD9FBA8F874" box="[1116,1142,1921,1947]" italics="true" pageId="10" pageNumber="347">4c</emphasis>
14 (1216),
<emphasis id="B936EA8BFFAC745984243AD9FAF0F874" box="[1287,1326,1921,1947]" italics="true" pageId="10" pageNumber="347">ps1</emphasis>
9 (710),
<emphasis id="B936EA8BFFAC745981B43AFDFF60F850" box="[151,190,1957,1983]" italics="true" pageId="10" pageNumber="347">ps2</emphasis>
9 (810),
<emphasis id="B936EA8BFFAC745980103AFDFE84F850" box="[307,346,1957,1983]" italics="true" pageId="10" pageNumber="347">ps3</emphasis>
19 (1519).
</paragraph>
<caption id="DF3D6611FFAD745881B43A1CFA94F8B3" box="[151,1354,1860,1884]" pageId="11" pageNumber="348" startId="11.[151,250,1860,1884]" targetBox="[151,1418,190,1836]" targetPageId="11" targetType="figure">
<paragraph id="8BFD3699FFAD745881B43A1CFA94F8B3" blockId="11.[151,1354,1860,1884]" box="[151,1354,1860,1884]" pageId="11" pageNumber="348">
<emphasis id="B936EA8BFFAD745881B43A1CFEC6F8B3" bold="true" box="[151,280,1860,1884]" pageId="11" pageNumber="348">FIGURE 11</emphasis>
.
<taxonomicName id="4C424D1AFFAD745880063A1CFDFDF8B3" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[293,547,1860,1884]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="348" phylum="Arthropoda" rank="species" species="spinisetus" status="sp. nov.">
<emphasis id="B936EA8BFFAD745880063A1CFDFDF8B3" box="[293,547,1860,1884]" italics="true" pageId="11" pageNumber="348">Spatulaphorus spinisetus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFAD7458830A3A1CFDA4F8B3" bold="true" box="[553,634,1860,1884]" pageId="11" pageNumber="348">
<taxonomicNameLabel id="A20557F0FFAD7458830A3A1CFDA4F8B3" box="[553,634,1860,1884]" pageId="11" pageNumber="348" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect.
</paragraph>
</caption>
<paragraph id="8BFD3699FFAD745F81E43AD3FF23FD8E" blockId="11.[151,1437,1930,2029]" lastBlockId="12.[151,1437,151,609]" lastPageId="12" lastPageNumber="349" pageId="11" pageNumber="348">
<emphasis id="B936EA8BFFAD745881E43AD3FF22F84B" box="[199,252,1931,1956]" italics="true" pageId="11" pageNumber="348">Legs</emphasis>
(
<figureCitation id="13792A1CFFAD7458802D3AD2FEB9F84B" box="[270,359,1930,1956]" captionStart="FIGURE 10" captionStartId="10.[151,250,1311,1335]" captionTargetBox="[151,1435,241,1288]" captionTargetId="figure-50@10.[151,1435,237,1288]" captionTargetPageId="10" captionText="FIGURE 10. Spatulaphorus spinisetus sp. nov., female: A—right leg I, dorsal aspect; B—right leg II, dorsal aspect." pageId="11" pageNumber="348">Figs 10</figureCitation>
,
<figureCitation id="13792A1CFFAD745880553AD2FE4DF84B" box="[374,403,1930,1956]" captionStart="FIGURE 11" captionStartId="11.[151,250,1860,1884]" captionTargetBox="[151,1418,190,1836]" captionTargetId="figure-17@11.[151,1436,181,1836]" captionTargetPageId="11" captionText="FIGURE 11. Spatulaphorus spinisetus sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." pageId="11" pageNumber="348">11</figureCitation>
). Setation of legs as in
<taxonomicName id="4C424D1AFFAD7458838B3AD3FC8EF84B" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[680,848,1930,1957]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="348" phylum="Arthropoda" rank="species" species="brevisetosus">
<emphasis id="B936EA8BFFAD7458838B3AD3FC8EF84B" box="[680,848,1930,1957]" italics="true" pageId="11" pageNumber="348">S. brevisetosus</emphasis>
</taxonomicName>
. Leg I (
<figureCitation id="13792A1CFFAD745882923AD2FBC2F84B" box="[945,1052,1930,1956]" captionStart="FIGURE 10" captionStartId="10.[151,250,1311,1335]" captionTargetBox="[151,1435,241,1288]" captionTargetId="figure-50@10.[151,1435,237,1288]" captionTargetPageId="10" captionText="FIGURE 10. Spatulaphorus spinisetus sp. nov., female: A—right leg I, dorsal aspect; B—right leg II, dorsal aspect." pageId="11" pageNumber="348">Fig. 10A</figureCitation>
). Tibiotarsus enlarged, with large claw; tibiotarsus internally with oval strongly sclerotized structure near solenidion
<emphasis id="B936EA8BFFAD745885623AF6FB81F827" box="[1089,1119,1966,1992]" italics="true" pageId="11" pageNumber="348">φ2</emphasis>
. Setae (
<emphasis id="B936EA8BFFAD7458859F3AF7FB14F827" box="[1212,1226,1967,1992]" italics="true" pageId="11" pageNumber="348">u</emphasis>
) consolidated into a horn-like structure opposing claw. Tarsal claw very thick and blunt-tipped. Seta
<emphasis id="B936EA8BFFAD7458851B3A8AFB98F803" box="[1080,1094,2002,2028]" italics="true" pageId="11" pageNumber="348">d</emphasis>
of femur spatulate, with short subterminal projection. Setae
<emphasis id="B936EA8BFFAA745F80C43DCFFE26FF5E" box="[487,504,151,177]" italics="true" pageId="12" pageNumber="349">l</emphasis>
of femur and
<emphasis id="B936EA8BFFAA745F83BB3DCFFD7AFF5E" box="[664,676,151,177]" italics="true" pageId="12" pageNumber="349">k</emphasis>
of tibiotarsus smooth, other leg setae (except eupathidia
<emphasis id="B936EA8BFFAA745F84053DCFFAE3FF5E" box="[1318,1341,151,177]" italics="true" pageId="12" pageNumber="349">p</emphasis>
,
<emphasis id="B936EA8BFFAA745F846F3DCFFAB7FF5E" box="[1356,1385,151,177]" italics="true" pageId="12" pageNumber="349">p”</emphasis>
,
<emphasis id="B936EA8BFFAA745F845B3DCFFA48FF5E" box="[1400,1430,151,177]" italics="true" pageId="12" pageNumber="349">tc</emphasis>
,
<emphasis id="B936EA8BFFAA745F81B43DE3FF65FF3A" box="[151,187,187,213]" italics="true" pageId="12" pageNumber="349">tc”</emphasis>
,
<emphasis id="B936EA8BFFAA745F81EB3DE3FF3FFF3A" box="[200,225,187,213]" italics="true" pageId="12" pageNumber="349">ft</emphasis>
,
<emphasis id="B936EA8BFFAA745F81CE3DE3FED3FF3A" box="[237,269,187,213]" italics="true" pageId="12" pageNumber="349">ft”</emphasis>
) sparsely barbed. Setae
<emphasis id="B936EA8BFFAA745F833A3DE3FDF4FF3A" box="[537,554,187,213]" italics="true" pageId="12" pageNumber="349">l</emphasis>
,
<emphasis id="B936EA8BFFAA745F83143DE3FD90FF3A" box="[567,590,187,213]" italics="true" pageId="12" pageNumber="349">l”</emphasis>
of femur,
<emphasis id="B936EA8BFFAA745F83E33DE3FD08FF3A" box="[704,726,187,213]" italics="true" pageId="12" pageNumber="349">l”</emphasis>
of genu and
<emphasis id="B936EA8BFFAA745F82473DE3FCAEFF3A" box="[868,880,187,213]" italics="true" pageId="12" pageNumber="349">k</emphasis>
of tibiotarsus blunt-tipped; other leg setae (except eupathidia) pointed. Lengths of solenidia
<emphasis id="B936EA8BFFAA745F834E3D87FD50FF16" box="[621,654,223,249]" italics="true" pageId="12" pageNumber="349">ω1</emphasis>
6 (67),
<emphasis id="B936EA8BFFAA745F83D43D87FCC7FF16" box="[759,793,223,249]" italics="true" pageId="12" pageNumber="349">ω2</emphasis>
5 (5),
<emphasis id="B936EA8BFFAA745F82463D87FC5CFF16" box="[869,898,223,249]" italics="true" pageId="12" pageNumber="349">φ1</emphasis>
7 (67),
<emphasis id="B936EA8BFFAA745F82C83D87FBD7FF16" box="[1003,1033,223,249]" italics="true" pageId="12" pageNumber="349">φ2</emphasis>
5 (45); solenidia
<emphasis id="B936EA8BFFAA745F85FC3D87FADEFF16" box="[1247,1280,223,249]" italics="true" pageId="12" pageNumber="349">ω1</emphasis>
and
<emphasis id="B936EA8BFFAA745F841B3D87FA8BFF16" box="[1336,1365,223,249]" italics="true" pageId="12" pageNumber="349">φ1</emphasis>
thick, clavate; other solenidia weakly clavate. Leg II (
<figureCitation id="13792A1CFFAA745F83873C5BFCD7FEF2" box="[676,777,259,285]" captionStart="FIGURE 10" captionStartId="10.[151,250,1311,1335]" captionTargetBox="[151,1435,241,1288]" captionTargetId="figure-50@10.[151,1435,237,1288]" captionTargetPageId="10" captionText="FIGURE 10. Spatulaphorus spinisetus sp. nov., female: A—right leg I, dorsal aspect; B—right leg II, dorsal aspect." pageId="12" pageNumber="349">Fig. 10B</figureCitation>
). Tarsus with thickened basally claws and pad-like empodium. Solenidion
<emphasis id="B936EA8BFFAA745F806E3C7FFEBFFEAF" box="[333,353,295,320]" italics="true" pageId="12" pageNumber="349">ω</emphasis>
6 (56) weakly clavate, solenidion
<emphasis id="B936EA8BFFAA745F83DD3C7FFCD3FEAF" box="[766,781,295,320]" italics="true" pageId="12" pageNumber="349">φ</emphasis>
absent. Setae
<emphasis id="B936EA8BFFAA745F82973C7FFC04FEAE" box="[948,986,295,321]" italics="true" pageId="12" pageNumber="349">pl”</emphasis>
,
<emphasis id="B936EA8BFFAA745F82CA3C7FFBD9FEAE" box="[1001,1031,295,321]" italics="true" pageId="12" pageNumber="349">tc</emphasis>
,
<emphasis id="B936EA8BFFAA745F85343C7FFBE4FEAE" box="[1047,1082,295,321]" italics="true" pageId="12" pageNumber="349">tc”</emphasis>
and
<emphasis id="B936EA8BFFAA745F85563C7FFB52FEAE" box="[1141,1164,295,321]" italics="true" pageId="12" pageNumber="349">u</emphasis>
of tarsus smooth, other setae barbed; setae
<emphasis id="B936EA8BFFAA745F804C3C13FE53FE8A" box="[367,397,331,357]" italics="true" pageId="12" pageNumber="349">tc</emphasis>
and
<emphasis id="B936EA8BFFAA745F80E03C13FE36FE8A" box="[451,488,331,357]" italics="true" pageId="12" pageNumber="349">pl”</emphasis>
of tarsus spiniform, blunt-tipped; setae
<emphasis id="B936EA8BFFAA745F82863C13FC6DFE8A" box="[933,947,331,357]" italics="true" pageId="12" pageNumber="349">d</emphasis>
,
<emphasis id="B936EA8BFFAA745F82E23C13FC0CFE8A" box="[961,978,331,357]" italics="true" pageId="12" pageNumber="349">l</emphasis>
, and
<emphasis id="B936EA8BFFAA745F852C3C13FBF4FE8A" box="[1039,1066,331,357]" italics="true" pageId="12" pageNumber="349">v”</emphasis>
of femur blunt-tipped; setae (
<emphasis id="B936EA8BFFAA745F845B3C13FA4CFE8B" box="[1400,1426,331,356]" italics="true" pageId="12" pageNumber="349">pv</emphasis>
) of tarsus weakly foliate distally; other leg setae pointed. Leg III (
<figureCitation id="13792A1CFFAA745F824B3C37FC11FE66" box="[872,975,367,393]" captionStart="FIGURE 11" captionStartId="11.[151,250,1860,1884]" captionTargetBox="[151,1418,190,1836]" captionTargetId="figure-17@11.[151,1436,181,1836]" captionTargetPageId="11" captionText="FIGURE 11. Spatulaphorus spinisetus sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." pageId="12" pageNumber="349">Fig. 11A</figureCitation>
). Femur divided into basi- and telofemur. Claws and empodium of same shape as on tarsus II. Setae
<emphasis id="B936EA8BFFAA745F82133CCBFC9DFE42" box="[816,835,403,429]" italics="true" pageId="12" pageNumber="349">v</emphasis>
of trochanter,
<emphasis id="B936EA8BFFAA745F82CA3CCBFBD9FE42" box="[1001,1031,403,429]" italics="true" pageId="12" pageNumber="349">tc</emphasis>
,
<emphasis id="B936EA8BFFAA745F85363CCBFBE5FE42" box="[1045,1083,403,429]" italics="true" pageId="12" pageNumber="349">pl”</emphasis>
, and
<emphasis id="B936EA8BFFAA745F85593CCBFB4FFE42" box="[1146,1169,403,429]" italics="true" pageId="12" pageNumber="349">u</emphasis>
of tarsus smooth, other setae barbed; setae
<emphasis id="B936EA8BFFAA745F804C3CEFFE53FE3E" box="[367,397,439,465]" italics="true" pageId="12" pageNumber="349">tc</emphasis>
and
<emphasis id="B936EA8BFFAA745F80E13CEFFE36FE3E" box="[450,488,439,465]" italics="true" pageId="12" pageNumber="349">pl”</emphasis>
of tarsus spiniform; setae
<emphasis id="B936EA8BFFAA745F822E3CEFFCFDFE3E" box="[781,803,439,465]" italics="true" pageId="12" pageNumber="349">v</emphasis>
of trochanter,
<emphasis id="B936EA8BFFAA745F82E53CEFFC0AFE3E" box="[966,980,439,465]" italics="true" pageId="12" pageNumber="349">d</emphasis>
,
<emphasis id="B936EA8BFFAA745F82C23CEFFC29FE3E" box="[993,1015,439,465]" italics="true" pageId="12" pageNumber="349">v</emphasis>
of femur, and
<emphasis id="B936EA8BFFAA745F85B93CEFFB75FE3E" box="[1178,1195,439,465]" italics="true" pageId="12" pageNumber="349">l</emphasis>
of genu blunt-tipped, other leg setae pointed; solenidion
<emphasis id="B936EA8BFFAA745F830D3C83FDE3FE1B" box="[558,573,475,500]" italics="true" pageId="12" pageNumber="349">φ</emphasis>
absent. Leg IV (
<figureCitation id="13792A1CFFAA745F82253C83FCB6FE1A" box="[774,872,475,501]" captionStart="FIGURE 11" captionStartId="11.[151,250,1860,1884]" captionTargetBox="[151,1418,190,1836]" captionTargetId="figure-17@11.[151,1436,181,1836]" captionTargetPageId="11" captionText="FIGURE 11. Spatulaphorus spinisetus sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." pageId="12" pageNumber="349">Fig 11B</figureCitation>
). Femur divided into basi- and telofemur. Claws simple, empodium as on tarsi II and III. Setae
<emphasis id="B936EA8BFFAA745F83B83CA7FD6FFDF6" box="[667,689,511,537]" italics="true" pageId="12" pageNumber="349">v</emphasis>
of trochanter and
<emphasis id="B936EA8BFFAA745F825F3CA7FC7CFDF6" box="[892,930,511,537]" italics="true" pageId="12" pageNumber="349">pl”</emphasis>
of tarsus smooth, other setae barbed; setae
<emphasis id="B936EA8BFFAA745F84A53CA7FA42FDF6" box="[1414,1436,511,537]" italics="true" pageId="12" pageNumber="349">v</emphasis>
of trochanter,
<emphasis id="B936EA8BFFAA745F80113F7BFE9EFDD2" box="[306,320,547,573]" italics="true" pageId="12" pageNumber="349">d</emphasis>
,
<emphasis id="B936EA8BFFAA745F806E3F7BFEBDFDD2" box="[333,355,547,573]" italics="true" pageId="12" pageNumber="349">v</emphasis>
of femur,
<emphasis id="B936EA8BFFAA745F80F63F7BFE2FFDD2" box="[469,497,547,573]" italics="true" pageId="12" pageNumber="349">v”</emphasis>
of tibia blunt-tipped; seta
<emphasis id="B936EA8BFFAA745F82363F7BFCE5FDD2" box="[789,827,547,573]" italics="true" pageId="12" pageNumber="349">pl”</emphasis>
of tarsus spiniform, other leg setae pointed; solenidion
<emphasis id="B936EA8BFFAA745F81B43F1FFF78FD8F" box="[151,166,583,608]" italics="true" pageId="12" pageNumber="349">φ</emphasis>
absent.
</paragraph>
<caption id="DF3D6611FFAA745F81B438D6FF29FA25" pageId="12" pageNumber="349" startId="12.[151,250,1422,1446]" targetBox="[151,1436,633,1399]" targetPageId="12" targetType="figure">
<paragraph id="8BFD3699FFAA745F81B438D6FF29FA25" blockId="12.[151,1436,1422,1482]" pageId="12" pageNumber="349">
<emphasis id="B936EA8BFFAA745F81B438D6FEC3FA49" bold="true" box="[151,285,1422,1446]" pageId="12" pageNumber="349">FIGURE 12</emphasis>
. DIC micrographs of
<taxonomicName id="4C424D1AFFAA745F832938D6FCD2FA49" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[522,780,1422,1446]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="12" pageNumber="349" phylum="Arthropoda" rank="species" species="spinisetus" status="sp. nov.">
<emphasis id="B936EA8BFFAA745F832938D6FCD2FA49" box="[522,780,1422,1446]" italics="true" pageId="12" pageNumber="349">Spatulaphorus spinisetus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFAA745F823538D7FCB4FA48" bold="true" box="[790,874,1423,1447]" pageId="12" pageNumber="349">
<taxonomicNameLabel id="A20557F0FFAA745F823538D7FCB4FA48" box="[790,874,1423,1447]" pageId="12" pageNumber="349" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—general view dorsally; B—general view ventrally.
</paragraph>
</caption>
<paragraph id="8BFD3699FFAA745F81EB38A1FE4FF9FC" blockId="12.[151,1436,1529,1987]" box="[200,401,1529,1555]" pageId="12" pageNumber="349">MALE unknown.</paragraph>
</subSubSection>
<subSubSection id="C3586512FFAA745F81E43B45FD9BF904" pageId="12" pageNumber="349" type="materials_examined">
<paragraph id="8BFD3699FFAA745F81E43B45FE56F990" blockId="12.[151,1436,1529,1987]" pageId="12" pageNumber="349">
<emphasis id="B936EA8BFFAA745F81E43B45FEB5F9D9" bold="true" box="[199,363,1564,1591]" pageId="12" pageNumber="349">Type material</emphasis>
.
<materialsCitation id="3B2A3CC4FFAA745F80543B45FE5FF990" collectingDate="2004-01-19" collectingDateMax="2005-01" collectingDateMin="2004-01-19" collectionCode="ZISP, CEMT" country="Brazil" location="Campos do Jordao" municipality="Female" pageId="12" pageNumber="349" specimenCount="36" specimenCount-female="36" stateProvince="Sao Paulo" typeStatus="holotype">
<collectingMunicipality id="6B99ACE3FFAA745F80543B45FE17F9D8" box="[375,457,1565,1591]" pageId="12" pageNumber="349">Female</collectingMunicipality>
<typeStatus id="54F9883BFFAA745F80EE3B45FDECF9D8" box="[461,562,1565,1591]" pageId="12" pageNumber="349" type="holotype">holotype</typeStatus>
, slide
<collectionCode id="ED53AE5CFFAA745F83573B45FD73F9D8" box="[628,685,1565,1591]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="12" pageNumber="349">ZISP</collectionCode>
T-Pygm-012,
<collectingCountry id="F3557609FFAA745F826B3B45FC51F9D8" box="[840,911,1565,1591]" name="Brazil" pageId="12" pageNumber="349">Brazil</collectingCountry>
,
<collectingRegion id="4986F87BFFAA745F82BB3B45FBD7F9D8" box="[920,1033,1565,1591]" country="Brazil" name="Sao Paulo" pageId="12" pageNumber="349">São Paulo</collectingRegion>
,
<location id="8E9D6042FFAA745F85313B45FB3EF9D8" LSID="urn:lsid:plazi:treatment:03EB878FFFAF745E81B43CEEFB9DFE1A:8E9D6042FFAA745F85313B45FB3EF9D8" box="[1042,1248,1565,1591]" country="Brazil" municipality="Female" name="Campos do Jordao" pageId="12" pageNumber="349" stateProvince="Sao Paulo">Campos do Jordão</location>
,
<date id="FFFC1059FFAA745F85CB3B44FAEFF9D8" box="[1256,1329,1564,1591]" pageId="12" pageNumber="349" value="2005-01">
<collectingDate id="EFB8E9B1FFAA745F85CB3B44FAEFF9D8" box="[1256,1329,1564,1591]" pageId="12" pageNumber="349" value="2005-01">I.2005</collectingDate>
</date>
(
<collectionCode id="ED53AE5CFFAA745F841E3B45FA4EF9D8" box="[1341,1424,1565,1591]" pageId="12" pageNumber="349">CEMT</collectionCode>
),
<collectedFrom id="385E7564FFAA745F81B43B19FDD2F9B4" box="[151,524,1601,1627]" pageId="12" pageNumber="349">
on
<taxonomicName id="4C424D1AFFAA745F819E3B19FDD2F9B4" authority="(Lucas)" baseAuthorityName="Lucas" baseAuthorityYear="1859" box="[189,524,1601,1627]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="349" phylum="Arthropoda" rank="species" species="buqueti">
<emphasis id="B936EA8BFFAA745F819E3B19FE74F9B4" box="[189,426,1601,1627]" italics="true" pageId="12" pageNumber="349">Dichotomius buqueti</emphasis>
(Lucas)
</taxonomicName>
</collectedFrom>
;
<typeStatus id="54F9883BFFAA745F83393B19FD54F9B4" box="[538,650,1601,1627]" pageId="12" pageNumber="349" type="paratype">paratypes</typeStatus>
:
<specimenCount id="9D44FD10FFAA745F83B43B19FCC9F9B4" box="[663,791,1601,1627]" count="10" pageId="12" pageNumber="349" type="female">10 females</specimenCount>
, same data;
<specimenCount id="9D44FD10FFAA745F828B3B19FBF6F9B4" box="[936,1064,1601,1627]" count="26" pageId="12" pageNumber="349" type="female">26 females</specimenCount>
, same locality but
<date id="FFFC1059FFAA745F84273B19FAAFF9B4" box="[1284,1393,1601,1627]" pageId="12" pageNumber="349" value="2004-01-19">
<collectingDate id="EFB8E9B1FFAA745F84273B19FAAFF9B4" box="[1284,1393,1601,1627]" pageId="12" pageNumber="349" value="2004-01-19">19.I.2004</collectingDate>
</date>
,
<collectedFrom id="385E7564FFAA745F84A33B19FE5FF990" pageId="12" pageNumber="349">
on
<taxonomicName id="4C424D1AFFAA745F81B43B3DFE5FF990" baseAuthorityName="Lucas" baseAuthorityYear="1859" box="[151,385,1637,1663]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="349" phylum="Arthropoda" rank="species" species="buqueti">
<emphasis id="B936EA8BFFAA745F81B43B3DFE5FF990" box="[151,385,1637,1663]" italics="true" pageId="12" pageNumber="349">Dichotomius buqueti</emphasis>
</taxonomicName>
</collectedFrom>
</materialsCitation>
.
</paragraph>
<paragraph id="8BFD3699FFAA745F81E43BD1FD9BF904" blockId="12.[151,1436,1529,1987]" pageId="12" pageNumber="349">
<emphasis id="B936EA8BFFAA745F81E43BD1FE58F94D" bold="true" box="[199,390,1672,1699]" pageId="12" pageNumber="349">Type deposition</emphasis>
.
<materialsCitation id="3B2A3CC4FFAA745F80B43BD1FE25F928" country="Russia" location="Zoological Institute" pageId="12" pageNumber="349" specimenCount="4" stateProvince="Saint Petersburg" typeStatus="holotype">
The
<typeStatus id="54F9883BFFAA745F80EF3BD1FDF0F94C" box="[460,558,1673,1699]" pageId="12" pageNumber="349" type="holotype">holotype</typeStatus>
and
<specimenCount id="9D44FD10FFAA745F83493BD1FCD3F94C" box="[618,781,1673,1699]" count="4" pageId="12" pageNumber="349" type="generic" typeStatus="paratype">four paratypes</specimenCount>
are deposited in the collection of
<location id="8E9D6042FFAA745F85BF3BD1FAA5F94C" LSID="urn:lsid:plazi:treatment:03EB878FFFAF745E81B43CEEFB9DFE1A:8E9D6042FFAA745F85BF3BD1FAA5F94C" box="[1180,1403,1673,1699]" country="Russia" name="Zoological Institute" pageId="12" pageNumber="349" stateProvince="Saint Petersburg">Zoological Institute</location>
of RAS,
<collectingRegion id="4986F87BFFAA745F81FC3BF5FE40F928" box="[223,414,1709,1735]" country="Russia" name="Saint Petersburg" pageId="12" pageNumber="349">Saint Petersburg</collectingRegion>
,
<collectingCountry id="F3557609FFAA745F808E3BF5FE25F928" box="[429,507,1709,1735]" name="Russia" pageId="12" pageNumber="349">Russia</collectingCountry>
</materialsCitation>
;
<materialsCitation id="3B2A3CC4FFAA745F83293BF5FAC5F928" box="[522,1307,1709,1735]" country="Russia" location="Tyumen State" pageId="12" pageNumber="349" specimenCount="1" stateProvince="Tyumen'" typeStatus="paratype">
other
<typeStatus id="54F9883BFFAA745F836D3BF5FD67F928" box="[590,697,1709,1735]" pageId="12" pageNumber="349" type="paratype">paratypes</typeStatus>
are deposited in the collection of the
<collectingRegion id="4986F87BFFAA745F855E3BF5FAC5F928" box="[1149,1307,1709,1735]" country="Russia" name="Tyumen'" pageId="12" pageNumber="349">Tyumen State</collectingRegion>
</materialsCitation>
<materialsCitation id="3B2A3CC4FFAA745F84053BF5FD9EF904" country="Russia" location="University Museum of Zoology" pageId="12" pageNumber="349" specimenCount="1" stateProvince="Tyumen'" typeStatus="paratype">
<location id="8E9D6042FFAA745F84053BF5FEA1F904" LSID="urn:lsid:plazi:treatment:03EB878FFFAF745E81B43CEEFB9DFE1A:8E9D6042FFAA745F84053BF5FEA1F904" country="Russia" name="University Museum of Zoology" pageId="12" pageNumber="349" stateProvince="Tyumen'">University Museum of Zoology</location>
,
<collectingRegion id="4986F87BFFAA745F80AA3B89FE39F904" box="[393,487,1745,1771]" country="Russia" name="Tyumen'" pageId="12" pageNumber="349">Tyumen</collectingRegion>
,
<collectingCountry id="F3557609FFAA745F80D13B89FD9EF904" box="[498,576,1745,1771]" name="Russia" pageId="12" pageNumber="349">Russia</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3586512FFAA745F81E43BACFE26F82C" pageId="12" pageNumber="349" type="diagnosis">
<paragraph id="8BFD3699FFAA745F81E43BACFE26F82C" blockId="12.[151,1436,1529,1987]" pageId="12" pageNumber="349">
<emphasis id="B936EA8BFFAA745F81E43BACFE15F8E1" bold="true" box="[199,459,1780,1806]" pageId="12" pageNumber="349">Differential diagnosis</emphasis>
. The new species is very similar to
<taxonomicName id="4C424D1AFFAA745F82A73BADFADAF8E0" authority="(Mahunka, 1980)" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" baseAuthorityName="Mahunka" baseAuthorityYear="1980" box="[900,1284,1781,1807]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="12" pageNumber="349" phylum="Arthropoda" rank="species" species="imbricatipes" status="comb. nov.">
<emphasis id="B936EA8BFFAA745F82A73BADFC4CF8E0" box="[900,914,1781,1807]" italics="true" pageId="12" pageNumber="349">S</emphasis>
.
<emphasis id="B936EA8BFFAA745F82863BADFBEFF8E0" box="[933,1073,1781,1807]" italics="true" pageId="12" pageNumber="349">imbricatipes</emphasis>
(
<bibRefCitation id="EFD34B68FFAA745F85673BADFB22F8E0" author="Mahunka, S." box="[1092,1276,1781,1807]" pageId="12" pageNumber="349" pagination="377 - 399" refId="ref16707" refString="Mahunka, S. (1980) Data to the knowledge of mites preserved in the &quot; Berlese collection &quot; (Acari: Tarsonemida, Oribatida). I. Acta Zoologica Academiae Scientiarum Hungaricae, 26, 377 - 399." type="journal article" year="1980">Mahunka, 1980</bibRefCitation>
)
</taxonomicName>
<emphasis id="B936EA8BFFAA745F84333BADFA4BF8E0" bold="true" box="[1296,1429,1781,1807]" pageId="12" pageNumber="349">
<taxonomicNameLabel id="A20557F0FFAA745F84333BADFA4BF8E0" box="[1296,1429,1781,1807]" pageId="12" pageNumber="349" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, described from
<collectingCountry id="F3557609FFAA745F80663A41FE6AF8DC" box="[325,436,1817,1843]" name="Argentina" pageId="12" pageNumber="349">Argentina</collectingCountry>
(
<bibRefCitation id="EFD34B68FFAA745F80E13A41FDB0F8DC" author="Mahunka, S." box="[450,622,1817,1843]" pageId="12" pageNumber="349" pagination="377 - 399" refId="ref16707" refString="Mahunka, S. (1980) Data to the knowledge of mites preserved in the &quot; Berlese collection &quot; (Acari: Tarsonemida, Oribatida). I. Acta Zoologica Academiae Scientiarum Hungaricae, 26, 377 - 399." type="journal article" year="1980">Mahunka 1980</bibRefCitation>
) in having spiniform setae
<emphasis id="B936EA8BFFAA745F82BD3A41FC31F8DC" box="[926,1007,1817,1843]" italics="true" pageId="12" pageNumber="349">pl”, tc</emphasis>
on tarsi II and III, enlarged tibiotarsus I and basally thickened claws on tarsi II and III. The new species differs from
<taxonomicName id="4C424D1AFFAA745F85323A65FB64F8B8" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" baseAuthorityName="Mahunka" baseAuthorityYear="1980" box="[1041,1210,1853,1879]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="12" pageNumber="349" phylum="Arthropoda" rank="species" species="imbricatipes">
<emphasis id="B936EA8BFFAA745F85323A65FBC1F8B8" box="[1041,1055,1853,1879]" italics="true" pageId="12" pageNumber="349">S</emphasis>
.
<emphasis id="B936EA8BFFAA745F850D3A65FB64F8B8" box="[1070,1210,1853,1879]" italics="true" pageId="12" pageNumber="349">imbricatipes</emphasis>
</taxonomicName>
in having distinctly longer dorsal idiosomal setae, especially
<emphasis id="B936EA8BFFAA745F834E3A39FD57F894" box="[621,649,1889,1915]" italics="true" pageId="12" pageNumber="349">h1</emphasis>
which are longer than the distance between their bases (vs. setae
<emphasis id="B936EA8BFFAA745F84A33A39FA42F894" box="[1408,1436,1889,1915]" italics="true" pageId="12" pageNumber="349">h1</emphasis>
shorter than distance between their bases in
<taxonomicName id="4C424D1AFFAA745F83A03ADDFCF4F870" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" baseAuthorityName="Mahunka" baseAuthorityYear="1980" box="[643,810,1925,1951]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="12" pageNumber="349" phylum="Arthropoda" rank="species" species="imbricatipes">
<emphasis id="B936EA8BFFAA745F83A03ADDFCF4F870" box="[643,810,1925,1951]" italics="true" pageId="12" pageNumber="349">S. imbricatipes</emphasis>
</taxonomicName>
) and setae
<emphasis id="B936EA8BFFAA745F828B3ADDFC1DF870" box="[936,963,1925,1951]" italics="true" pageId="12" pageNumber="349">c1</emphasis>
distinctly shorter than
<emphasis id="B936EA8BFFAA745F85E03ADDFB00F870" box="[1219,1246,1925,1951]" italics="true" pageId="12" pageNumber="349">c2</emphasis>
(vs. setae
<emphasis id="B936EA8BFFAA745F84703ADDFAB0F870" box="[1363,1390,1925,1951]" italics="true" pageId="12" pageNumber="349">c1</emphasis>
and
<emphasis id="B936EA8BFFAA745F81B43AF1FF6CF82C" box="[151,178,1961,1987]" italics="true" pageId="12" pageNumber="349">c2</emphasis>
subequal in
<taxonomicName id="4C424D1AFFAA745F80633AF1FE36F82C" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" baseAuthorityName="Mahunka" baseAuthorityYear="1980" box="[320,488,1961,1987]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="12" pageNumber="349" phylum="Arthropoda" rank="species" species="imbricatipes">
<emphasis id="B936EA8BFFAA745F80633AF1FE36F82C" box="[320,488,1961,1987]" italics="true" pageId="12" pageNumber="349">S. imbricatipes</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3586512FFAB745E81E43DCFFC60FF3A" pageId="13" pageNumber="350" type="etymology">
<paragraph id="8BFD3699FFAB745E81E43DCFFC60FF3A" blockId="13.[151,1437,151,501]" pageId="13" pageNumber="350">
<emphasis id="B936EA8BFFAB745E81E43DCFFE96FF5E" bold="true" box="[199,328,151,177]" pageId="13" pageNumber="350">Etymology</emphasis>
. The name of the new species is a combination of two Latin words:
<emphasis id="B936EA8BFFAB745E85613DCFFB7CFF5F" box="[1090,1186,151,176]" italics="true" pageId="13" pageNumber="350">spinosus</emphasis>
meaning
<emphasis id="B936EA8BFFAB745E84333DCFFAB8FF5F" box="[1296,1382,151,176]" italics="true" pageId="13" pageNumber="350">spinous</emphasis>
, and
<emphasis id="B936EA8BFFAB745E81B43DE3FF1AFF3B" box="[151,196,187,212]" italics="true" pageId="13" pageNumber="350">seta</emphasis>
meaning
<emphasis id="B936EA8BFFAB745E80103DE3FEA5FF3A" box="[307,379,187,213]" italics="true" pageId="13" pageNumber="350">bristle</emphasis>
, and refers to the spiniform setae on tarsi II and III.
</paragraph>
</subSubSection>
<subSubSection id="C3586512FFAB745E81E43D87FB9DFE1A" pageId="13" pageNumber="350" type="discussion">
<paragraph id="8BFD3699FFAB745E81E43D87FB9DFE1A" blockId="13.[151,1437,151,501]" pageId="13" pageNumber="350">
<emphasis id="B936EA8BFFAB745E81E43D87FEEEFF16" bold="true" box="[199,304,223,249]" pageId="13" pageNumber="350">Remark.</emphasis>
<bibRefCitation id="EFD34B68FFAB745E80153D87FE2EFF16" author="Mahunka, S." box="[310,496,223,249]" pageId="13" pageNumber="350" pagination="377 - 399" refId="ref16707" refString="Mahunka, S. (1980) Data to the knowledge of mites preserved in the &quot; Berlese collection &quot; (Acari: Tarsonemida, Oribatida). I. Acta Zoologica Academiae Scientiarum Hungaricae, 26, 377 - 399." type="journal article" year="1980">Mahunka (1980)</bibRefCitation>
described
<emphasis id="B936EA8BFFAB745E83453D87FC73FF16" box="[614,941,223,249]" italics="true" pageId="13" pageNumber="350">
<taxonomicName id="4C424D1AFFAB745E83453D87FCC5FF16" box="[614,795,223,249]" genus="Pygmephorellus" pageId="13" pageNumber="367" rank="genus">Pygmephorellus</taxonomicName>
<taxonomicName id="4C424D1AFFAB745E82023D87FC73FF16" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" baseAuthorityName="Mahunka" baseAuthorityYear="1980" box="[801,941,223,249]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="13" pageNumber="350" phylum="Arthropoda" rank="species" species="imbricatipes">imbricatipes</taxonomicName>
</emphasis>
based on specimens deposited in the Berlese collection and collected on
<taxonomicName id="4C424D1AFFAB745E80FB3C5BFDAEFEF2" box="[472,624,259,285]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="350" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B936EA8BFFAB745E80FB3C5BFD98FEF2" box="[472,582,259,285]" italics="true" pageId="13" pageNumber="350">Phanaeus</emphasis>
sp.
</taxonomicName>
in
<collectingCountry id="F3557609FFAB745E83BA3C5BFCD2FEF2" box="[665,780,259,285]" name="Argentina" pageId="13" pageNumber="350">Argentina</collectingCountry>
. In the description he did not describe the chaetotaxy of legs. However, illustrations of legs IIV show character states typical for
<taxonomicName id="4C424D1AFFAB745E82C23C7FFB5AFEAE" authorityName="Rack" authorityYear="1993" box="[993,1156,295,321]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="13" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="B936EA8BFFAB745E82C23C7FFB5AFEAE" box="[993,1156,295,321]" italics="true" pageId="13" pageNumber="350">Spatulaphorus</emphasis>
</taxonomicName>
, namely the presence of only two setae on genu I, only one seta on genu II and genu IV without setae. Also, setae
<emphasis id="B936EA8BFFAB745E85A73C13FB75FE8A" box="[1156,1195,331,357]" italics="true" pageId="13" pageNumber="350">ps3</emphasis>
are located distinctly anteriad setae
<emphasis id="B936EA8BFFAB745E80143C37FEA4FE66" box="[311,378,367,393]" italics="true" pageId="13" pageNumber="350">ps12</emphasis>
which is characteristic for most Neotropical
<taxonomicName id="4C424D1AFFAB745E82503C37FBABFE66" box="[883,1141,367,393]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" kingdom="Animalia" order="Trombidiformes" pageId="13" pageNumber="350" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B936EA8BFFAB745E82503C37FBC8FE66" box="[883,1046,367,393]" italics="true" pageId="13" pageNumber="350">Spatulaphorus</emphasis>
species.
</taxonomicName>
<bibRefCitation id="EFD34B68FFAB745E855F3C37FA42FE66" author="Khaustov, A. A. &amp; Trach, V. A." box="[1148,1436,367,393]" pageId="13" pageNumber="350" pagination="480 - 485" refId="ref16479" refString="Khaustov, A. A. &amp; Trach, V. A. (2012) A new species and new records of mites of the genus Spatulaphorus Rack (Acari: Heterostigmata: Pygmephoridae) from Ukraine. International Journal of Acarology, 38, 480 - 485. https: // doi. org / 10.1080 / 01647954.2012.677475" type="journal article" year="2012">Khaustov &amp; Trach (2012)</bibRefCitation>
and
<bibRefCitation id="EFD34B68FFAB745E81E53CCBFE35FE42" author="Khaustov, A. A. &amp; Frolov, A. V." box="[198,491,403,429]" pageId="13" pageNumber="350" pagination="501 - 521" refId="ref16128" refString="Khaustov, A. A. &amp; Frolov, A. V. (2017) New species of heterostigmatic mites (Acari: Heterostigmata: Athyreacaridae, Dolichocybidae, Pygmephoridae) associated with scarab beetles (Coleoptera: Geotrupidae, Scarabaeidae) from Brazil. Zootaxa, 4294 (5), 501 - 521. https: // doi. org / 10.11646 / zootaxa. 4294.5.1" type="journal article" year="2017">Khaustov &amp; Frolov (2017</bibRefCitation>
;
<bibRefCitation id="EFD34B68FFAB745E80D53CCBFDF1FE42" author="Khaustov, A. A. &amp; Frolov, A. V." box="[502,559,403,429]" pageId="13" pageNumber="350" pagination="333 - 345" refId="ref16337" refString="Khaustov, A. A. &amp; Frolov, A. V. (2021) Two new species and a new record of Spatulaphorus (Acari &quot; Pygmephoridae) phoretic on scarab beetles (Coleoptera: Scarabaeidae) from French Guiana. Acarina, 29 (2), 333 - 345. https: // doi. org / 10.21684 / 0132 - 8077 - 2021 - 29 - 2 - 233 - 245" type="journal article" year="2021">2021</bibRefCitation>
) suggested that
<taxonomicName id="4C424D1AFFAB745E83C73CCBFC55FE42" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" baseAuthorityName="Mahunka" baseAuthorityYear="1980" box="[740,907,403,429]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="13" pageNumber="350" phylum="Arthropoda" rank="species" species="imbricatipes">
<emphasis id="B936EA8BFFAB745E83C73CCBFC55FE42" box="[740,907,403,429]" italics="true" pageId="13" pageNumber="350">P. imbricatipes</emphasis>
</taxonomicName>
could be a species of the genus
<taxonomicName id="4C424D1AFFAB745E85D13CCBFA4BFE42" authorityName="Rack" authorityYear="1993" box="[1266,1429,403,429]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="13" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="B936EA8BFFAB745E85D13CCBFA4BFE42" box="[1266,1429,403,429]" italics="true" pageId="13" pageNumber="350">Spatulaphorus</emphasis>
</taxonomicName>
. Discovery of the closely related species
<taxonomicName id="4C424D1AFFAB745E83483CEFFC58FE3F" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[619,902,439,465]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="13" pageNumber="350" phylum="Arthropoda" rank="species" species="spinisetus" status="sp. nov.">
<emphasis id="B936EA8BFFAB745E83483CEFFC58FE3F" box="[619,902,439,465]" italics="true" pageId="13" pageNumber="350">Spatulaphorus spinisetus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFAB745E82B23CEFFC33FE3E" bold="true" box="[913,1005,439,465]" pageId="13" pageNumber="350">
<taxonomicNameLabel id="A20557F0FFAB745E82B23CEFFC33FE3E" box="[913,1005,439,465]" pageId="13" pageNumber="350" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
confirms this opinion. Therefore, we move
<emphasis id="B936EA8BFFAB745E81FF3C83FDF9FE1A" box="[220,551,475,501]" italics="true" pageId="13" pageNumber="350">
<taxonomicName id="4C424D1AFFAB745E81FF3C83FE4FFE1A" box="[220,401,475,501]" genus="Pygmephorellus" pageId="13" pageNumber="367" rank="genus">Pygmephorellus</taxonomicName>
imbricaripes
</emphasis>
to the genus
<taxonomicName id="4C424D1AFFAB745E839E3C83FCBEFE1A" authorityName="Rack" authorityYear="1993" box="[701,864,475,501]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="13" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="B936EA8BFFAB745E839E3C83FCBEFE1A" box="[701,864,475,501]" italics="true" pageId="13" pageNumber="350">Spatulaphorus</emphasis>
</taxonomicName>
(new combination).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,472 @@
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<mods:title id="DAEAFCC591D228A74CD45029D0B9446B">Seven new species of Spatulaphorus Rack (Acari: Pygmephoridae) phoretic on scarab beetles (Coleoptera: Scarabaeidae)</mods:title>
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<mods:namePart id="672EDFCA9DA04CF14C8D1ED3C62FC615">Khaustov, Alexander A.</mods:namePart>
<mods:affiliation id="91EC6E52768EA5E99B86AA18112AA7EB">Tyumen State University, Tyumen, 6 Volodarskogo Str., 625003 Russia.</mods:affiliation>
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<mods:affiliation id="72DA4B9A1BD6FD8376907C7EE802DA07">Zoological Institute of Russian Academy of Sciences, Universitetskaya Embankment 1, 199034, Saint Petersburg, Russia</mods:affiliation>
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<mods:affiliation id="BCCD752CA7E2ED90EBE54D366D8A6955">Zoological Institute of Russian Academy of Sciences, Universitetskaya Embankment 1, 199034, Saint Petersburg, Russia</mods:affiliation>
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<subSubSection id="C3586512FFB2744781B43BF9FDC3F953" box="[151,541,1697,1724]" pageId="20" pageNumber="357" type="nomenclature">
<paragraph id="8BFD3699FFB2744781B43BF9FDC3F953" blockId="20.[151,541,1697,1760]" box="[151,541,1697,1724]" pageId="20" pageNumber="357">
<heading id="D0B581F5FFB2744781B43BF9FDC3F953" bold="true" box="[151,541,1697,1724]" fontSize="11" level="1" pageId="20" pageNumber="357" reason="1">
<emphasis id="B936EA8BFFB2744781B43BF9FDC3F953" bold="true" box="[151,541,1697,1724]" pageId="20" pageNumber="357">
<taxonomicName id="4C424D1AFFB2744781B43BF9FE66F953" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[151,440,1697,1724]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="20" pageNumber="357" phylum="Arthropoda" rank="species" species="reductus" status="sp. nov.">
<emphasis id="B936EA8BFFB2744781B43BF9FE66F953" bold="true" box="[151,440,1697,1724]" italics="true" pageId="20" pageNumber="357">Spatulaphorus reductus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20557F0FFB2744780E33BFAFDC3F953" box="[448,541,1698,1724]" pageId="20" pageNumber="357" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3586512FFB2744481B43B9EFE4FF8C3" lastPageId="23" lastPageNumber="360" pageId="20" pageNumber="357" type="description">
<paragraph id="8BFD3699FFB2744781B43B9EFEF9F90F" blockId="20.[151,541,1697,1760]" box="[151,295,1734,1760]" pageId="20" pageNumber="357">
(
<figureCitation id="13792A1CFFB2744781BC3B9EFEC1F90F" box="[159,287,1734,1760]" captionStart-0="FIGURE 21" captionStart-1="FIGURE 22" captionStart-2="FIGURE 23" captionStart-3="FIGURE 24" captionStartId-0="21.[151,250,1357,1381]" captionStartId-1="22.[151,250,1442,1466]" captionStartId-2="23.[151,250,1740,1764]" captionStartId-3="24.[151,247,1089,1113]" captionTargetBox-0="[151,1435,245,1333]" captionTargetBox-1="[151,1417,185,1419]" captionTargetBox-2="[160,1427,190,1716]" captionTargetBox-3="[151,1436,181,1066]" captionTargetId-0="figure-41@21.[151,1435,237,1333]" captionTargetId-1="figure-24@22.[151,1436,181,1419]" captionTargetId-2="figure-17@23.[151,1436,181,1716]" captionTargetId-3="figure-24@24.[151,1436,181,1066]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionTargetPageId-2="23" captionTargetPageId-3="24" captionText-0="FIGURE 21. Spatulaphorus reductus sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." captionText-1="FIGURE 22. Spatulaphorus reductus sp. nov., female: A—right leg I, dorsal aspect; B—left leg II, dorsal aspect." captionText-2="FIGURE 23. Spatulaphorus reductus sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." captionText-3="FIGURE 24. DIC micrographs of Spatulaphorus reductus sp. nov., female:A—general view dorsally; B—general view ventrally." pageId="20" pageNumber="357">Figs 2124</figureCitation>
)
</paragraph>
<paragraph id="8BFD3699FFB2744781B43A55FA81F8C7" blockId="20.[151,1437,1805,2012]" box="[151,1375,1805,1832]" pageId="20" pageNumber="357">
<emphasis id="B936EA8BFFB2744781B43A55FEFFF8C8" bold="true" box="[151,289,1805,1831]" pageId="20" pageNumber="357">Description</emphasis>
. FEMALE. Body weakly sclerotized. Length of idiosoma 155 (150175), width 105 (100110).
</paragraph>
<paragraph id="8BFD3699FFB2744681E43A6AFD94FF3A" blockId="20.[151,1437,1805,2012]" lastBlockId="21.[151,1436,151,213]" lastPageId="21" lastPageNumber="358" pageId="20" pageNumber="357">
<emphasis id="B936EA8BFFB2744781E43A6AFE88F8A3" box="[199,342,1842,1868]" italics="true" pageId="20" pageNumber="357">Gnathosoma</emphasis>
. Gnathosomal capsule, excluding palps, subrectangular, length 19 (1921), width 23 (2023). Dorsum with one pair of pointed cheliceral setae (
<emphasis id="B936EA8BFFB2744783DF3A0EFCFAF89F" box="[764,804,1878,1904]" italics="true" pageId="20" pageNumber="357">cha</emphasis>
) 10 (610). Dorsal median apodeme present, poorly developed. Postpalpal setae (
<emphasis id="B936EA8BFFB2744780FC3A22FE25F87C" box="[479,507,1914,1939]" italics="true" pageId="20" pageNumber="357">pp</emphasis>
) 4 (5) baculiform, situated laterad setae
<emphasis id="B936EA8BFFB2744782EB3A22FC2EF87B" box="[968,1008,1914,1940]" italics="true" pageId="20" pageNumber="357">cha</emphasis>
. Ventral gnathosoma with one pair of smooth, pointed subcapitular setae
<emphasis id="B936EA8BFFB2744783013AC6FDE8F858" box="[546,566,1950,1975]" italics="true" pageId="20" pageNumber="357">m</emphasis>
8 (89) and round pits
<emphasis id="B936EA8BFFB27447821D3AC6FC92F858" box="[830,844,1950,1975]" italics="true" pageId="20" pageNumber="357">n</emphasis>
situated posteriad
<emphasis id="B936EA8BFFB2744785033AC6FBEAF858" box="[1056,1076,1950,1975]" italics="true" pageId="20" pageNumber="357">m</emphasis>
. Palps with smooth and pointed setae
<emphasis id="B936EA8BFFB2744781F43A9AFEDDF833" box="[215,259,1986,2012]" italics="true" pageId="20" pageNumber="357">dFe</emphasis>
7 (67) and
<emphasis id="B936EA8BFFB2744780B63A9AFE1AF833" box="[405,452,1986,2012]" italics="true" pageId="20" pageNumber="357">dGe</emphasis>
8 (813) dorsolaterally. Palps ventrally each with mushroom-like accessory setigenous structure (
<emphasis id="B936EA8BFFB3744680283DCFFEF1FF5F" box="[267,303,151,176]" italics="true" pageId="21" pageNumber="358">ass</emphasis>
) and tiny solenidion
<emphasis id="B936EA8BFFB3744683063DCFFDE7FF5F" box="[549,569,151,176]" italics="true" pageId="21" pageNumber="358">ω</emphasis>
. Palps terminated with small blunt-tipped tibial claw and tiny eupathid-like seta. Pharyngeal pumps poorly visible.
</paragraph>
<caption id="DF3D6611FFB3744681B43815FB24FA8A" box="[151,1274,1357,1381]" pageId="21" pageNumber="358" startId="21.[151,250,1357,1381]" targetBox="[151,1435,245,1333]" targetPageId="21" targetType="figure">
<paragraph id="8BFD3699FFB3744681B43815FB24FA8A" blockId="21.[151,1274,1357,1381]" box="[151,1274,1357,1381]" pageId="21" pageNumber="358">
<emphasis id="B936EA8BFFB3744681B43815FEC7FA8A" bold="true" box="[151,281,1357,1381]" pageId="21" pageNumber="358">FIGURE 21</emphasis>
.
<taxonomicName id="4C424D1AFFB3744680053815FDC8FA8A" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[294,534,1357,1381]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="21" pageNumber="358" phylum="Arthropoda" rank="species" species="reductus" status="sp. nov.">
<emphasis id="B936EA8BFFB3744680053815FDC8FA8A" box="[294,534,1357,1381]" italics="true" pageId="21" pageNumber="358">Spatulaphorus reductus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFB37446833E3815FDB3FA8B" bold="true" box="[541,621,1357,1380]" pageId="21" pageNumber="358">
<taxonomicNameLabel id="A20557F0FFB37446833E3815FDB3FA8B" box="[541,621,1357,1380]" pageId="21" pageNumber="358" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—dorsum of body; B—venter of body. Legs omitted.
</paragraph>
</caption>
<paragraph id="8BFD3699FFB3744681E438CBFBB2F945" blockId="21.[151,1437,1427,1958]" pageId="21" pageNumber="358">
<emphasis id="B936EA8BFFB3744681E438CBFE4AFA42" box="[199,404,1427,1453]" italics="true" pageId="21" pageNumber="358">Idiosomal dorsum</emphasis>
(
<figureCitation id="13792A1CFFB37446808738CBFDCCFA42" box="[420,530,1427,1453]" captionStart="FIGURE 21" captionStartId="21.[151,250,1357,1381]" captionTargetBox="[151,1435,245,1333]" captionTargetId="figure-41@21.[151,1435,237,1333]" captionTargetPageId="21" captionText="FIGURE 21. Spatulaphorus reductus sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." pageId="21" pageNumber="358">Figs 21A</figureCitation>
,
<figureCitation id="13792A1CFFB37446833D38CBFD8DFA42" box="[542,595,1427,1453]" captionStart="FIGURE 24" captionStartId="24.[151,247,1089,1113]" captionTargetBox="[151,1436,181,1066]" captionTargetId="figure-24@24.[151,1436,181,1066]" captionTargetPageId="24" captionText="FIGURE 24. DIC micrographs of Spatulaphorus reductus sp. nov., female:A—general view dorsally; B—general view ventrally." pageId="21" pageNumber="358">24A</figureCitation>
). All dorsal shields smooth. Prodorsal shield with three pairs of setae (
<emphasis id="B936EA8BFFB37446845838CBFA4BFA42" box="[1403,1429,1427,1453]" italics="true" pageId="21" pageNumber="358">v1</emphasis>
,
<emphasis id="B936EA8BFFB3744681B438EFFF6CFA3E" box="[151,178,1463,1489]" italics="true" pageId="21" pageNumber="358">v2</emphasis>
,
<emphasis id="B936EA8BFFB3744681E338EFFF38FA3E" box="[192,230,1463,1489]" italics="true" pageId="21" pageNumber="358">sc2</emphasis>
), one pair of weakly barbed capitate trichobothria (
<emphasis id="B936EA8BFFB37446820D38EFFC8AFA3E" box="[814,852,1463,1489]" italics="true" pageId="21" pageNumber="358">sc1</emphasis>
) and one pair of oval stigmata. Stigmata associated with well-developed tracheal trunks. Setae
<emphasis id="B936EA8BFFB3744683A43884FD4DFA1A" box="[647,659,1500,1525]" italics="true" pageId="21" pageNumber="358">e</emphasis>
,
<emphasis id="B936EA8BFFB3744683803883FD61FA1A" box="[675,703,1499,1525]" italics="true" pageId="21" pageNumber="358">h1</emphasis>
and
<emphasis id="B936EA8BFFB3744683D83883FCC9FA1A" box="[763,791,1499,1525]" italics="true" pageId="21" pageNumber="358">h2</emphasis>
pointed; other dorsal setae blunt-tipped; all dorsal setae barbed. Alveoli of setae
<emphasis id="B936EA8BFFB37446808E3B58FE67F9F6" box="[429,441,1536,1561]" italics="true" pageId="21" pageNumber="358">e</emphasis>
and
<emphasis id="B936EA8BFFB3744680D138A7FE24F9F6" box="[498,506,1535,1561]" italics="true" pageId="21" pageNumber="358">f</emphasis>
distinctly separated. Cupules
<emphasis id="B936EA8BFFB3744682733B58FCB8F9F6" box="[848,870,1536,1561]" italics="true" pageId="21" pageNumber="358">ia</emphasis>
on tergite D,
<emphasis id="B936EA8BFFB3744685263B58FBFFF9F6" box="[1029,1057,1536,1561]" italics="true" pageId="21" pageNumber="358">im</emphasis>
on tergite EF and
<emphasis id="B936EA8BFFB3744685D538A7FAD2F9F6" box="[1270,1292,1535,1561]" italics="true" pageId="21" pageNumber="358">ih</emphasis>
on tergite H large, round;
<emphasis id="B936EA8BFFB3744680133B7CFE92F9D2" box="[304,332,1572,1597]" italics="true" pageId="21" pageNumber="358">im</emphasis>
much larger than subequal
<emphasis id="B936EA8BFFB3744683A93B7CFD7EF9D2" box="[650,672,1572,1597]" italics="true" pageId="21" pageNumber="358">ia</emphasis>
and
<emphasis id="B936EA8BFFB3744683FA3B7BFD31F9D2" box="[729,751,1571,1597]" italics="true" pageId="21" pageNumber="358">ih</emphasis>
. Lengths of dorsal setae:
<emphasis id="B936EA8BFFB3744685343B7BFBEFF9D2" box="[1047,1073,1571,1597]" italics="true" pageId="21" pageNumber="358">v1</emphasis>
21 (2022),
<emphasis id="B936EA8BFFB3744685E43B7BFB3CF9D2" box="[1223,1250,1571,1597]" italics="true" pageId="21" pageNumber="358">v2</emphasis>
16 (1517),
<emphasis id="B936EA8BFFB3744684543B7BFA43F9D2" box="[1399,1437,1571,1597]" italics="true" pageId="21" pageNumber="358">sc2</emphasis>
30 (3033),
<emphasis id="B936EA8BFFB3744680023B1FFEE2F98E" box="[289,316,1607,1633]" italics="true" pageId="21" pageNumber="358">c1</emphasis>
21 (2123),
<emphasis id="B936EA8BFFB3744680EA3B1FFE3AF98E" box="[457,484,1607,1633]" italics="true" pageId="21" pageNumber="358">c2</emphasis>
33 (3033),
<emphasis id="B936EA8BFFB3744683563B1FFD5DF98E" box="[629,643,1607,1633]" italics="true" pageId="21" pageNumber="358">d</emphasis>
24 (2425),
<emphasis id="B936EA8BFFB3744682373B10FCFEF98E" box="[788,800,1608,1633]" italics="true" pageId="21" pageNumber="358">e</emphasis>
20 (1920),
<emphasis id="B936EA8BFFB3744682923B1FFC67F98E" box="[945,953,1607,1633]" italics="true" pageId="21" pageNumber="358">f</emphasis>
25 (2527),
<emphasis id="B936EA8BFFB3744685693B1FFBBBF98E" box="[1098,1125,1607,1633]" italics="true" pageId="21" pageNumber="358">h1</emphasis>
36 (3336),
<emphasis id="B936EA8BFFB3744685D53B1FFACDF98E" box="[1270,1299,1607,1633]" italics="true" pageId="21" pageNumber="358">h2</emphasis>
20 (1320). Distances between setae:
<emphasis id="B936EA8BFFB37446809F3B33FE20F96A" box="[444,510,1643,1670]" italics="true" pageId="21" pageNumber="358">v1v1</emphasis>
19 (1820),
<emphasis id="B936EA8BFFB3744683B63B33FD09F96A" box="[661,727,1643,1669]" italics="true" pageId="21" pageNumber="358">v2v2</emphasis>
34 (3235),
<emphasis id="B936EA8BFFB37446824D3B33FC18F96A" box="[878,966,1643,1669]" italics="true" pageId="21" pageNumber="358">sc2sc2</emphasis>
32 (3236),
<emphasis id="B936EA8BFFB37446857E3B33FB7EF96A" box="[1117,1184,1643,1670]" italics="true" pageId="21" pageNumber="358">c1c1</emphasis>
32 (3235),
<emphasis id="B936EA8BFFB3744684173B33FAA9F96A" box="[1332,1399,1643,1670]" italics="true" pageId="21" pageNumber="358">c1c2</emphasis>
20 (1921),
<emphasis id="B936EA8BFFB3744681DD3BD7FEF6F946" box="[254,296,1679,1706]" italics="true" pageId="21" pageNumber="358">dd</emphasis>
43 (4352),
<emphasis id="B936EA8BFFB37446809B3BC8FE05F946" box="[440,475,1679,1706]" italics="true" pageId="21" pageNumber="358">ef</emphasis>
6 (69),
<emphasis id="B936EA8BFFB3744683623BD7FD81F946" box="[577,607,1679,1706]" italics="true" pageId="21" pageNumber="358">ff</emphasis>
36 (3639),
<emphasis id="B936EA8BFFB3744683D33BD7FCEBF946" box="[752,821,1679,1706]" italics="true" pageId="21" pageNumber="358">h1h1</emphasis>
37 (3643),
<emphasis id="B936EA8BFFB3744682E53BD7FBD2F946" box="[966,1036,1679,1706]" italics="true" pageId="21" pageNumber="358">h1h2</emphasis>
5 (56).
</paragraph>
<paragraph id="8BFD3699FFB3744681E43BEBFAFCF849" blockId="21.[151,1437,1427,1958]" pageId="21" pageNumber="358">
<emphasis id="B936EA8BFFB3744681E43BEBFE5DF922" box="[199,387,1715,1741]" italics="true" pageId="21" pageNumber="358">Idiosomal venter</emphasis>
(
<figureCitation id="13792A1CFFB3744680B23BEBFE24F922" box="[401,506,1715,1741]" captionStart="FIGURE 21" captionStartId="21.[151,250,1357,1381]" captionTargetBox="[151,1435,245,1333]" captionTargetId="figure-41@21.[151,1435,237,1333]" captionTargetPageId="21" captionText="FIGURE 21. Spatulaphorus reductus sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." pageId="21" pageNumber="358">Figs 21B</figureCitation>
,
<figureCitation id="13792A1CFFB3744683273BEBFDE9F922" box="[516,567,1715,1741]" captionStart="FIGURE 24" captionStartId="24.[151,247,1089,1113]" captionTargetBox="[151,1436,181,1066]" captionTargetId="figure-24@24.[151,1436,181,1066]" captionTargetPageId="24" captionText="FIGURE 24. DIC micrographs of Spatulaphorus reductus sp. nov., female:A—general view dorsally; B—general view ventrally." pageId="21" pageNumber="358">24B</figureCitation>
). All ventral plates smooth.All ventral setae pointed; setae
<emphasis id="B936EA8BFFB3744685E63BEBFADDF922" box="[1221,1283,1715,1741]" italics="true" pageId="21" pageNumber="358">ps1-3</emphasis>
barbed, other ventral setae smooth. Bases of setae
<emphasis id="B936EA8BFFB3744683123B8FFD86F91E" box="[561,600,1751,1777]" italics="true" pageId="21" pageNumber="358">ps3</emphasis>
situated slightly anteriad bases of setae
<emphasis id="B936EA8BFFB3744685393B8FFB86F91E" box="[1050,1112,1751,1777]" italics="true" pageId="21" pageNumber="358">ps1-2</emphasis>
; alveoli of setae
<emphasis id="B936EA8BFFB37446843B3B8FFAE1F91E" box="[1304,1343,1751,1777]" italics="true" pageId="21" pageNumber="358">ps1</emphasis>
and
<emphasis id="B936EA8BFFB3744684563B8FFA42F91E" box="[1397,1436,1751,1777]" italics="true" pageId="21" pageNumber="358">ps2</emphasis>
clearly separated. Ap1 and ap2 well-developed and joined with thick appr; apsej absent; ap3 not evident; ap4 well developed, exceeding beyond bases of setae
<emphasis id="B936EA8BFFB3744683A03A47FD41F8D6" box="[643,671,1823,1849]" italics="true" pageId="21" pageNumber="358">3b</emphasis>
; ap5 absent.Ags small, cup-like; pgs small, oval. Posterior margin of posterior sternal plate straight in middle part. Posterior margin of aggenital plate rounded. Lengths of ventral setae:
<emphasis id="B936EA8BFFB3744681B43A3FFF6AF86E" box="[151,180,1895,1921]" italics="true" pageId="21" pageNumber="358">1a</emphasis>
12 (1112),
<emphasis id="B936EA8BFFB3744680653A3FFEBCF86E" box="[326,354,1895,1921]" italics="true" pageId="21" pageNumber="358">1b</emphasis>
14 (1416),
<emphasis id="B936EA8BFFB3744680D53A3FFDCEF86E" box="[502,528,1895,1921]" italics="true" pageId="21" pageNumber="358">1с</emphasis>
13 (1315),
<emphasis id="B936EA8BFFB3744683873A3FFD61F86E" box="[676,703,1895,1921]" italics="true" pageId="21" pageNumber="358">2a</emphasis>
11 (1113),
<emphasis id="B936EA8BFFB3744682723A3FFCB5F86E" box="[849,875,1895,1921]" italics="true" pageId="21" pageNumber="358">2с</emphasis>
12 (1112),
<emphasis id="B936EA8BFFB3744682DD3A3FFBC4F86E" box="[1022,1050,1895,1921]" italics="true" pageId="21" pageNumber="358">3a</emphasis>
14 (1314),
<emphasis id="B936EA8BFFB37446858D3A3FFB14F86E" box="[1198,1226,1895,1921]" italics="true" pageId="21" pageNumber="358">3b</emphasis>
12 (1213),
<emphasis id="B936EA8BFFB37446847D3A3FFAA6F86E" box="[1374,1400,1895,1921]" italics="true" pageId="21" pageNumber="358">3c</emphasis>
14 (1314),
<emphasis id="B936EA8BFFB3744681DD3AD3FEC4F84A" box="[254,282,1931,1957]" italics="true" pageId="21" pageNumber="358">4a</emphasis>
13 (1013),
<emphasis id="B936EA8BFFB3744680893AD3FE19F84A" box="[426,455,1931,1957]" italics="true" pageId="21" pageNumber="358">4b</emphasis>
15 (1516),
<emphasis id="B936EA8BFFB3744683743AD3FDACF84A" box="[599,626,1931,1957]" italics="true" pageId="21" pageNumber="358">4c</emphasis>
16 (1216),
<emphasis id="B936EA8BFFB3744682213AD3FCF7F84A" box="[770,809,1931,1957]" italics="true" pageId="21" pageNumber="358">ps1</emphasis>
13 (1213),
<emphasis id="B936EA8BFFB3744682993AD3FC3FF84A" box="[954,993,1931,1957]" italics="true" pageId="21" pageNumber="358">ps2</emphasis>
13 (1213),
<emphasis id="B936EA8BFFB3744685523AD3FB46F84A" box="[1137,1176,1931,1957]" italics="true" pageId="21" pageNumber="358">ps3</emphasis>
16 (1516).
</paragraph>
<caption id="DF3D6611FFB0744581B438FAFAC7FA55" box="[151,1305,1442,1467]" pageId="22" pageNumber="359" startId="22.[151,250,1442,1466]" targetBox="[151,1417,185,1419]" targetPageId="22" targetType="figure">
<paragraph id="8BFD3699FFB0744581B438FAFAC7FA55" blockId="22.[151,1305,1442,1467]" box="[151,1305,1442,1467]" pageId="22" pageNumber="359">
<emphasis id="B936EA8BFFB0744581B438FAFEC7FA55" bold="true" box="[151,281,1442,1466]" pageId="22" pageNumber="359">FIGURE 22</emphasis>
.
<taxonomicName id="4C424D1AFFB07445800538FBFDC8FA54" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[294,534,1443,1467]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="22" pageNumber="359" phylum="Arthropoda" rank="species" species="reductus" status="sp. nov.">
<emphasis id="B936EA8BFFB07445800538FBFDC8FA54" box="[294,534,1443,1467]" italics="true" pageId="22" pageNumber="359">Spatulaphorus reductus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFB07445833E38FBFDB3FA54" bold="true" box="[541,621,1443,1467]" pageId="22" pageNumber="359">
<taxonomicNameLabel id="A20557F0FFB07445833E38FBFDB3FA54" box="[541,621,1443,1467]" pageId="22" pageNumber="359" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—right leg I, dorsal aspect; B—left leg II, dorsal aspect.
</paragraph>
</caption>
<paragraph id="8BFD3699FFB0744581E438B2FE05F85C" blockId="22.[151,1437,1513,1971]" pageId="22" pageNumber="359">
<emphasis id="B936EA8BFFB0744581E438B2FF22F9EC" box="[199,252,1514,1539]" italics="true" pageId="22" pageNumber="359">Legs</emphasis>
(
<figureCitation id="13792A1CFFB07445802A38B1FE80F9EC" box="[265,350,1513,1539]" captionStart="FIGURE 22" captionStartId="22.[151,250,1442,1466]" captionTargetBox="[151,1417,185,1419]" captionTargetId="figure-24@22.[151,1436,181,1419]" captionTargetPageId="22" captionText="FIGURE 22. Spatulaphorus reductus sp. nov., female: A—right leg I, dorsal aspect; B—left leg II, dorsal aspect." pageId="22" pageNumber="359">Figs 22</figureCitation>
,
<figureCitation id="13792A1CFFB07445804438B1FE59F9EC" box="[359,391,1513,1539]" captionStart="FIGURE 23" captionStartId="23.[151,250,1740,1764]" captionTargetBox="[160,1427,190,1716]" captionTargetId="figure-17@23.[151,1436,181,1716]" captionTargetPageId="23" captionText="FIGURE 23. Spatulaphorus reductus sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." pageId="22" pageNumber="359">23</figureCitation>
). Setation of legs as in
<taxonomicName id="4C424D1AFFB07445835D38B1FCFFF9EC" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[638,801,1513,1539]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="22" pageNumber="359" phylum="Arthropoda" rank="species" species="brevisetosus">
<emphasis id="B936EA8BFFB07445835D38B1FCFFF9EC" box="[638,801,1513,1539]" italics="true" pageId="22" pageNumber="359">S. brevisetosus</emphasis>
</taxonomicName>
except presence of solenidia
<emphasis id="B936EA8BFFB07445854238B2FBAEF9EC" box="[1121,1136,1514,1539]" italics="true" pageId="22" pageNumber="359">φ</emphasis>
on tibiae II-III and absence of seta
<emphasis id="B936EA8BFFB0744581C43B55FED2F9C8" box="[231,268,1549,1575]" italics="true" pageId="22" pageNumber="359">pl”</emphasis>
on tarsus IV. Leg I (
<figureCitation id="13792A1CFFB0744580CE3B55FD88F9C8" box="[493,598,1549,1575]" captionStart="FIGURE 22" captionStartId="22.[151,250,1442,1466]" captionTargetBox="[151,1417,185,1419]" captionTargetId="figure-24@22.[151,1436,181,1419]" captionTargetPageId="22" captionText="FIGURE 22. Spatulaphorus reductus sp. nov., female: A—right leg I, dorsal aspect; B—left leg II, dorsal aspect." pageId="22" pageNumber="359">Fig. 22A</figureCitation>
). Tibiotarsus enlarged, with large and thick claw; tibiotarsus internally with oval strongly sclerotized structure near solenidion
<emphasis id="B936EA8BFFB0744583F83B69FD27F9A4" box="[731,761,1585,1611]" italics="true" pageId="22" pageNumber="359">φ2</emphasis>
. Setae (
<emphasis id="B936EA8BFFB07445827B3B6AFCB8F9A4" box="[856,870,1586,1611]" italics="true" pageId="22" pageNumber="359">u</emphasis>
) consolidated into a horn-like structure opposing claw. Tarsal claw blunt-tipped. Seta
<emphasis id="B936EA8BFFB0744583113B0DFD9EF980" box="[562,576,1621,1647]" italics="true" pageId="22" pageNumber="359">d</emphasis>
of femur spatulate, with short subterminal projection. Setae
<emphasis id="B936EA8BFFB0744585D33B0DFADFF980" box="[1264,1281,1621,1647]" italics="true" pageId="22" pageNumber="359">l</emphasis>
of femur and
<emphasis id="B936EA8BFFB0744581B43B21FF7DF97C" box="[151,163,1657,1683]" italics="true" pageId="22" pageNumber="359">k</emphasis>
of tibiotarsus smooth, other leg setae (except eupathidia
<emphasis id="B936EA8BFFB0744582133B21FC99F97C" box="[816,839,1657,1683]" italics="true" pageId="22" pageNumber="359">p</emphasis>
,
<emphasis id="B936EA8BFFB0744582753B21FCAAF97C" box="[854,884,1657,1683]" italics="true" pageId="22" pageNumber="359">p”</emphasis>
,
<emphasis id="B936EA8BFFB0744582A03B21FC7FF97C" box="[899,929,1657,1683]" italics="true" pageId="22" pageNumber="359">tc</emphasis>
,
<emphasis id="B936EA8BFFB0744582933B21FC0AF97C" box="[944,980,1657,1683]" italics="true" pageId="22" pageNumber="359">tc”</emphasis>
,
<emphasis id="B936EA8BFFB0744582C73B21FC23F97C" box="[996,1021,1657,1683]" italics="true" pageId="22" pageNumber="359">ft</emphasis>
,
<emphasis id="B936EA8BFFB07445852F3B21FBF5F97C" box="[1036,1067,1657,1683]" italics="true" pageId="22" pageNumber="359">ft”</emphasis>
) barbed. Setae
<emphasis id="B936EA8BFFB0744585FE3B21FB35F97C" box="[1245,1259,1657,1683]" italics="true" pageId="22" pageNumber="359">l</emphasis>
of femur and
<emphasis id="B936EA8BFFB0744584B33B21FA42F97C" box="[1424,1436,1657,1683]" italics="true" pageId="22" pageNumber="359">k</emphasis>
of tibiotarsus weakly blunt-tipped; other leg setae (except eupathidia) pointed. Lengths of solenidia
<emphasis id="B936EA8BFFB0744585DB3BC5FAC4F958" box="[1272,1306,1693,1719]" italics="true" pageId="22" pageNumber="359">ω1</emphasis>
6 (6),
<emphasis id="B936EA8BFFB0744584463BC5FA59F958" box="[1381,1415,1693,1719]" italics="true" pageId="22" pageNumber="359">ω2</emphasis>
3 (3),
<emphasis id="B936EA8BFFB0744581E53B99FF3DF934" box="[198,227,1729,1755]" italics="true" pageId="22" pageNumber="359">φ1</emphasis>
5 (5),
<emphasis id="B936EA8BFFB07445800D3B99FE92F934" box="[302,332,1729,1755]" italics="true" pageId="22" pageNumber="359">φ2</emphasis>
4 (4); solenidia
<emphasis id="B936EA8BFFB0744583203B99FDFAF934" box="[515,548,1729,1755]" italics="true" pageId="22" pageNumber="359">ω1</emphasis>
and
<emphasis id="B936EA8BFFB0744583783B99FDA6F934" box="[603,632,1729,1755]" italics="true" pageId="22" pageNumber="359">φ1</emphasis>
thick, clavate; other solenidia weakly clavate. Leg II (
<figureCitation id="13792A1CFFB0744585F93B99FA9FF934" box="[1242,1345,1729,1755]" captionStart="FIGURE 22" captionStartId="22.[151,250,1442,1466]" captionTargetBox="[151,1417,185,1419]" captionTargetId="figure-24@22.[151,1436,181,1419]" captionTargetPageId="22" captionText="FIGURE 22. Spatulaphorus reductus sp. nov., female: A—right leg I, dorsal aspect; B—left leg II, dorsal aspect." pageId="22" pageNumber="359">Fig. 22B</figureCitation>
). Tarsus with simple claws and pad-like empodium. Solenidion
<emphasis id="B936EA8BFFB0744582213BBEFCC8F910" box="[770,790,1766,1791]" italics="true" pageId="22" pageNumber="359">ω</emphasis>
6 (56) weakly clavate, solenidion
<emphasis id="B936EA8BFFB0744585843BBEFB68F910" box="[1191,1206,1766,1791]" italics="true" pageId="22" pageNumber="359">φ</emphasis>
2 (2) very short. All leg setae barbed and pointed. Leg III (
<figureCitation id="13792A1CFFB0744583663A51FD70F8CC" box="[581,686,1801,1827]" captionStart="FIGURE 23" captionStartId="23.[151,250,1740,1764]" captionTargetBox="[160,1427,190,1716]" captionTargetId="figure-17@23.[151,1436,181,1716]" captionTargetPageId="23" captionText="FIGURE 23. Spatulaphorus reductus sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." pageId="22" pageNumber="359">Fig. 23A</figureCitation>
). Femur divided into basi- and telofemur. Claws and empodium of same shape as on tarsus II. All leg setae barbed and pointed; solenidion
<emphasis id="B936EA8BFFB0744582EE3A76FC02F8A8" box="[973,988,1838,1863]" italics="true" pageId="22" pageNumber="359">φ</emphasis>
very small (less than 0.5). Leg IV (
<figureCitation id="13792A1CFFB0744584563A75FF14F884" captionStart="FIGURE 23" captionStartId="23.[151,250,1740,1764]" captionTargetBox="[160,1427,190,1716]" captionTargetId="figure-17@23.[151,1436,181,1716]" captionTargetPageId="23" captionText="FIGURE 23. Spatulaphorus reductus sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." pageId="22" pageNumber="359">Fig 23B</figureCitation>
). Femur divided into basi- and telofemur. Claws and empodium as on tarsi II and III. All leg setae barbed and pointed; external part of solenidion
<emphasis id="B936EA8BFFB0744583123A2EFD9EF860" box="[561,576,1910,1935]" italics="true" pageId="22" pageNumber="359">φ</emphasis>
absent, but subcuticular very small cup-shaped structure situated at typical insertion point of solenidion.
</paragraph>
<caption id="DF3D6611FFB1744481B43B94FAE0F90C" box="[151,1342,1740,1764]" pageId="23" pageNumber="360" startId="23.[151,250,1740,1764]" targetBox="[160,1427,190,1716]" targetPageId="23" targetType="figure">
<paragraph id="8BFD3699FFB1744481B43B94FAE0F90C" blockId="23.[151,1342,1740,1764]" box="[151,1342,1740,1764]" pageId="23" pageNumber="360">
<emphasis id="B936EA8BFFB1744481B43B94FEC7F90B" bold="true" box="[151,281,1740,1764]" pageId="23" pageNumber="360">FIGURE 23</emphasis>
.
<taxonomicName id="4C424D1AFFB1744480053B94FDC8F90B" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[294,534,1740,1764]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="23" pageNumber="360" phylum="Arthropoda" rank="species" species="reductus" status="sp. nov.">
<emphasis id="B936EA8BFFB1744480053B94FDC8F90B" box="[294,534,1740,1764]" italics="true" pageId="23" pageNumber="360">Spatulaphorus reductus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFB17444833E3B94FDB3F90B" bold="true" box="[541,621,1740,1764]" pageId="23" pageNumber="360">
<taxonomicNameLabel id="A20557F0FFB17444833E3B94FDB3F90B" box="[541,621,1740,1764]" pageId="23" pageNumber="360" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect.
</paragraph>
</caption>
<paragraph id="8BFD3699FFB1744481EB3A4AFE4FF8C3" blockId="23.[151,1436,1810,1981]" box="[200,401,1810,1836]" pageId="23" pageNumber="360">MALE unknown.</paragraph>
</subSubSection>
<subSubSection id="C3586512FFB1744B81E43A6EFD9BFB05" lastPageId="24" lastPageNumber="361" pageId="23" pageNumber="360" type="materials_examined">
<paragraph id="8BFD3699FFB1744481E43A6EFC10F852" blockId="23.[151,1436,1810,1981]" pageId="23" pageNumber="360">
<emphasis id="B936EA8BFFB1744481E43A6EFEAFF8BF" bold="true" box="[199,369,1846,1872]" pageId="23" pageNumber="360">Type material</emphasis>
. Female
<typeStatus id="54F9883BFFB1744480FD3A6EFD9DF8BF" box="[478,579,1846,1872]" pageId="23" pageNumber="360" type="holotype">holotype</typeStatus>
, slide
<collectionCode id="ED53AE5CFFB1744483AC3A6EFD16F8BF" box="[655,712,1846,1872]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="23" pageNumber="360">ZISP</collectionCode>
T-Pygm-015
<collectingCountry id="F3557609FFB1744482453A6EFC06F8BF" box="[870,984,1846,1872]" name="Argentina" pageId="23" pageNumber="360">Argentina</collectingCountry>
,
<collectingRegion id="4986F87BFFB1744482C53A6EFB10F8BE" box="[998,1230,1846,1873]" country="Argentina" name="Santiago del Estero" pageId="23" pageNumber="360">Santiago Del Estero</collectingRegion>
, Salado River, on
<taxonomicName id="4C424D1AFFB1744481B43A03FE21F89B" authority="(Harold)" baseAuthorityName="Harold" baseAuthorityYear="1868" box="[151,511,1882,1908]" class="Insecta" family="Scarabaeidae" genus="Parataenius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="360" phylum="Arthropoda" rank="species" species="simulator">
<emphasis id="B936EA8BFFB1744481B43A03FE4DF89B" box="[151,403,1882,1908]" italics="true" pageId="23" pageNumber="360">Parataenius simulator</emphasis>
(Harold)
</taxonomicName>
;
<typeStatus id="54F9883BFFB1744483283A03FDA5F89A" box="[523,635,1883,1909]" pageId="23" pageNumber="360" type="paratype">paratypes</typeStatus>
: same locality data,
<specimenCount id="9D44FD10FFB1744482413A02FC03F89B" box="[866,989,1882,1908]" count="47" pageId="23" pageNumber="360" type="female">47 females</specimenCount>
on 9
<taxonomicName id="4C424D1AFFB17444853C3A03FAC4F89B" baseAuthorityName="Harold" baseAuthorityYear="1868" box="[1055,1306,1882,1908]" class="Insecta" family="Scarabaeidae" genus="Parataenius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="360" phylum="Arthropoda" rank="species" species="simulator">
<emphasis id="B936EA8BFFB17444853C3A03FAC4F89B" box="[1055,1306,1882,1908]" italics="true" pageId="23" pageNumber="360">Parataenius simulator</emphasis>
</taxonomicName>
beetles;
<specimenCount id="9D44FD10FFB1744484A33A02FF30F877" count="38" pageId="23" pageNumber="360" type="female">38 females</specimenCount>
on 7
<taxonomicName id="4C424D1AFFB17444800D3A26FDE0F877" authority="Harold" authorityName="Harold" authorityYear="1867" box="[302,574,1918,1944]" class="Insecta" family="Scarabaeidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="360" phylum="Arthropoda" rank="species" species="picinus">
<emphasis id="B936EA8BFFB17444800D3A26FE36F877" box="[302,488,1918,1944]" italics="true" pageId="23" pageNumber="360">Ataenius picinus</emphasis>
Harold
</taxonomicName>
beetles;
<specimenCount id="9D44FD10FFB1744483823A26FCD0F877" box="[673,782,1918,1944]" count="8" pageId="23" pageNumber="360" type="female">8 females</specimenCount>
on 2
<taxonomicName id="4C424D1AFFB17444826D3A26FBA6F877" authority="Schmidt" authorityName="Schmidt" authorityYear="1916" box="[846,1144,1918,1944]" class="Insecta" family="Scarabaeidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="360" phylum="Arthropoda" rank="species" species="clavatus">
<emphasis id="B936EA8BFFB17444826D3A26FBCAF877" box="[846,1044,1918,1944]" italics="true" pageId="23" pageNumber="360">Ataenius clavatus</emphasis>
Schmidt
</taxonomicName>
beetles;
<specimenCount id="9D44FD10FFB1744485FE3A26FA97F877" box="[1245,1353,1918,1945]" count="5" pageId="23" pageNumber="360" type="female">5 females</specimenCount>
on one
<taxonomicName id="4C424D1AFFB1744481B43AFAFE2FF853" authority="(Blanchard)" baseAuthorityName="Blanchard" baseAuthorityYear="1846" box="[151,497,1954,1980]" class="Insecta" family="Scarabaeidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="360" phylum="Arthropoda" rank="species" species="platensis">
<emphasis id="B936EA8BFFB1744481B43AFAFEBAF853" box="[151,356,1954,1980]" italics="true" pageId="23" pageNumber="360">Ataenius platensis</emphasis>
(Blanchard)
</taxonomicName>
beetle. All host beetles are housed in
<collectionCode id="ED53AE5CFFB1744482BB3AFBFC14F852" box="[920,970,1955,1981]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="23" pageNumber="360" type="Museum">ZIN</collectionCode>
.
</paragraph>
<caption id="DF3D6611FFBE744B81B43919FA42FBB5" box="[151,1436,1089,1114]" pageId="24" pageNumber="361" startId="24.[151,247,1089,1113]" targetBox="[151,1436,181,1066]" targetPageId="24" targetType="figure">
<paragraph id="8BFD3699FFBE744B81B43919FA42FBB5" blockId="24.[151,1436,1089,1114]" box="[151,1436,1089,1114]" pageId="24" pageNumber="361">
<emphasis id="B936EA8BFFBE744B81B43919FEC8FBB6" bold="true" box="[151,278,1089,1113]" pageId="24" pageNumber="361">FIGURE 24</emphasis>
. DIC micrographs of
<taxonomicName id="4C424D1AFFBE744B80CD391AFD09FBB5" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[494,727,1090,1114]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="24" pageNumber="361" phylum="Arthropoda" rank="species" species="reductus" status="sp. nov.">
<emphasis id="B936EA8BFFBE744B80CD391AFD09FBB5" box="[494,727,1090,1114]" italics="true" pageId="24" pageNumber="361">Spatulaphorus reductus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFBE744B83FE391AFCF5FBB5" bold="true" box="[733,811,1090,1114]" pageId="24" pageNumber="361">
<taxonomicNameLabel id="A20557F0FFBE744B83FE391AFCF5FBB5" box="[733,811,1090,1114]" pageId="24" pageNumber="361" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—general view dorsally; B—general view ventrally.
</paragraph>
</caption>
<paragraph id="8BFD3699FFBE744B81E439D0FD9BFB05" blockId="24.[151,1437,1160,1438]" pageId="24" pageNumber="361">
<emphasis id="B936EA8BFFBE744B81E439D0FE58FB4D" bold="true" box="[199,390,1160,1186]" pageId="24" pageNumber="361">Type deposition</emphasis>
.
<materialsCitation id="3B2A3CC4FFBE744B80B439D0FE25FB29" country="Russia" location="Zoological Institute" pageId="24" pageNumber="361" specimenCount="4" stateProvince="Saint Petersburg" typeStatus="holotype">
The
<typeStatus id="54F9883BFFBE744B80EF39D0FDF0FB4D" box="[460,558,1160,1186]" pageId="24" pageNumber="361" type="holotype">holotype</typeStatus>
and
<specimenCount id="9D44FD10FFBE744B834939D0FCD3FB4D" box="[618,781,1160,1186]" count="4" pageId="24" pageNumber="361" type="generic" typeStatus="paratype">four paratypes</specimenCount>
are deposited in the collection of
<location id="8E9D6042FFBE744B85BF39D0FAA5FB4D" LSID="urn:lsid:plazi:treatment:03EB878FFFB2744B81B43BF9FE00FA71:8E9D6042FFBE744B85BF39D0FAA5FB4D" box="[1180,1403,1160,1186]" country="Russia" name="Zoological Institute" pageId="24" pageNumber="361" stateProvince="Saint Petersburg">Zoological Institute</location>
of RAS,
<collectingRegion id="4986F87BFFBE744B81FC39F4FE40FB29" box="[223,414,1196,1222]" country="Russia" name="Saint Petersburg" pageId="24" pageNumber="361">Saint Petersburg</collectingRegion>
,
<collectingCountry id="F3557609FFBE744B808E39F4FE25FB29" box="[429,507,1196,1222]" name="Russia" pageId="24" pageNumber="361">Russia</collectingCountry>
</materialsCitation>
;
<materialsCitation id="3B2A3CC4FFBE744B832939F4FAC5FB29" box="[522,1307,1196,1222]" country="Russia" location="Tyumen State" pageId="24" pageNumber="361" specimenCount="1" stateProvince="Tyumen'" typeStatus="paratype">
other
<typeStatus id="54F9883BFFBE744B836D39F4FD67FB29" box="[590,697,1196,1222]" pageId="24" pageNumber="361" type="paratype">paratypes</typeStatus>
are deposited in the collection of the
<collectingRegion id="4986F87BFFBE744B855E39F4FAC5FB29" box="[1149,1307,1196,1222]" country="Russia" name="Tyumen'" pageId="24" pageNumber="361">Tyumen State</collectingRegion>
</materialsCitation>
<materialsCitation id="3B2A3CC4FFBE744B840539F4FD9EFB05" country="Russia" location="University Museum of Zoology" pageId="24" pageNumber="361" specimenCount="1" stateProvince="Tyumen'" typeStatus="paratype">
<location id="8E9D6042FFBE744B840539F4FEA1FB05" LSID="urn:lsid:plazi:treatment:03EB878FFFB2744B81B43BF9FE00FA71:8E9D6042FFBE744B840539F4FEA1FB05" country="Russia" name="University Museum of Zoology" pageId="24" pageNumber="361" stateProvince="Tyumen'">University Museum of Zoology</location>
,
<collectingRegion id="4986F87BFFBE744B80AA3988FE39FB05" box="[393,487,1232,1258]" country="Russia" name="Tyumen'" pageId="24" pageNumber="361">Tyumen</collectingRegion>
,
<collectingCountry id="F3557609FFBE744B80D13988FD9EFB05" box="[498,576,1232,1258]" name="Russia" pageId="24" pageNumber="361">Russia</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3586512FFBE744B81E439ACFB2EFAB9" pageId="24" pageNumber="361" type="diagnosis">
<paragraph id="8BFD3699FFBE744B81E439ACFB2EFAB9" blockId="24.[151,1437,1160,1438]" pageId="24" pageNumber="361">
<emphasis id="B936EA8BFFBE744B81E439ACFE18FAE1" bold="true" box="[199,454,1268,1294]" pageId="24" pageNumber="361">Differential diagnosis</emphasis>
. The new species significantly differs from other species of the genus
<taxonomicName id="4C424D1AFFBE744B85FF39ACFAA1FAE1" authorityName="Rack" authorityYear="1993" box="[1244,1407,1268,1294]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="24" pageNumber="361" phylum="Arthropoda" rank="genus">
<emphasis id="B936EA8BFFBE744B85FF39ACFAA1FAE1" box="[1244,1407,1268,1294]" italics="true" pageId="24" pageNumber="361">Spatulaphorus</emphasis>
</taxonomicName>
in having tarsus IV with five setae (
<emphasis id="B936EA8BFFBE744B83293840FDEEFADD" box="[522,560,1304,1330]" italics="true" pageId="24" pageNumber="361">pl”</emphasis>
absent) and the alveoli of setae
<emphasis id="B936EA8BFFBE744B82B43840FC60FADD" box="[919,958,1304,1330]" italics="true" pageId="24" pageNumber="361">ps1</emphasis>
and
<emphasis id="B936EA8BFFBE744B82D73840FBC5FADD" box="[1012,1051,1304,1330]" italics="true" pageId="24" pageNumber="361">ps2</emphasis>
clearly separated. In other species of
<taxonomicName id="4C424D1AFFBE744B81953864FE87FAB9" authorityName="Rack" authorityYear="1993" box="[182,345,1340,1366]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="24" pageNumber="361" phylum="Arthropoda" rank="genus">
<emphasis id="B936EA8BFFBE744B81953864FE87FAB9" box="[182,345,1340,1366]" italics="true" pageId="24" pageNumber="361">Spatulaphorus</emphasis>
</taxonomicName>
tarsus IV has six setae (
<emphasis id="B936EA8BFFBE744B834A3864FD50FAB9" box="[617,654,1340,1366]" italics="true" pageId="24" pageNumber="361">pl”</emphasis>
present) and the alveoli of setae
<emphasis id="B936EA8BFFBE744B82DC3864FBE3FAB9" box="[1023,1085,1340,1366]" italics="true" pageId="24" pageNumber="361">ps1-2</emphasis>
are contiguous.
</paragraph>
</subSubSection>
<subSubSection id="C3586512FFBE744B81E43838FE00FA71" pageId="24" pageNumber="361" type="etymology">
<paragraph id="8BFD3699FFBE744B81E43838FE00FA71" blockId="24.[151,1437,1160,1438]" pageId="24" pageNumber="361">
<emphasis id="B936EA8BFFBE744B81E43838FE96FA95" bold="true" box="[199,328,1376,1402]" pageId="24" pageNumber="361">Etymology</emphasis>
. The name of the new species is derived from Latin
<taxonomicName id="4C424D1AFFBE744B82A63838FC3AFA95" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[901,996,1376,1402]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="24" pageNumber="361" phylum="Arthropoda" rank="species" species="reductus">
<emphasis id="B936EA8BFFBE744B82A63838FC3AFA95" box="[901,996,1376,1402]" italics="true" pageId="24" pageNumber="361">reductus</emphasis>
</taxonomicName>
meaning
<emphasis id="B936EA8BFFBE744B85733838FB77FA95" box="[1104,1193,1376,1402]" italics="true" pageId="24" pageNumber="361">reduced</emphasis>
, and refers to reduced number of setae on tarsus IV.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,500 @@
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<mods:title id="8FCCB450EEA7F20D6CB541CCB0E4E802">Seven new species of Spatulaphorus Rack (Acari: Pygmephoridae) phoretic on scarab beetles (Coleoptera: Scarabaeidae)</mods:title>
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<mods:namePart id="C0F0191AB2CDC53EC8BD9BAAFAD1918A">Khaustov, Alexander A.</mods:namePart>
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<mods:affiliation id="90F6AEFE8013B3474A518FCECA163494">Zoological Institute of Russian Academy of Sciences, Universitetskaya Embankment 1, 199034, Saint Petersburg, Russia</mods:affiliation>
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<mods:namePart id="502300D2AA467B6214FF1EF82E5C5029">Akhmetova, Lilia A.</mods:namePart>
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<treatment id="03EB878FFFB6744781B43B0AFC07FE1A" LSID="urn:lsid:plazi:treatment:03EB878FFFB6744781B43B0AFC07FE1A" httpUri="http://treatment.plazi.org/id/03EB878FFFB6744781B43B0AFC07FE1A" lastPageId="20" lastPageNumber="357" pageId="16" pageNumber="353">
<subSubSection id="C3586512FFB6744381B43B0AFDD4F983" box="[151,522,1618,1645]" pageId="16" pageNumber="353" type="nomenclature">
<paragraph id="8BFD3699FFB6744381B43B0AFDD4F983" blockId="16.[151,522,1618,1680]" box="[151,522,1618,1645]" pageId="16" pageNumber="353">
<heading id="D0B581F5FFB6744381B43B0AFDD4F983" bold="true" box="[151,522,1618,1645]" fontSize="11" level="1" pageId="16" pageNumber="353" reason="1">
<emphasis id="B936EA8BFFB6744381B43B0AFDD4F983" bold="true" box="[151,522,1618,1645]" pageId="16" pageNumber="353">
<taxonomicName id="4C424D1AFFB6744381B43B0AFE7BF983" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[151,421,1618,1645]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="16" pageNumber="353" phylum="Arthropoda" rank="species" species="incisus" status="sp. nov.">
<emphasis id="B936EA8BFFB6744381B43B0AFE7BF983" bold="true" box="[151,421,1618,1645]" italics="true" pageId="16" pageNumber="353">Spatulaphorus incisus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20557F0FFB67443808F3B0AFDD4F983" box="[428,522,1618,1644]" pageId="16" pageNumber="353" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3586512FFB6744081B43B2EFE4FF86B" lastPageId="19" lastPageNumber="356" pageId="16" pageNumber="353" type="description">
<paragraph id="8BFD3699FFB6744381B43B2EFEF9F97F" blockId="16.[151,522,1618,1680]" box="[151,295,1654,1680]" pageId="16" pageNumber="353">
(
<figureCitation id="13792A1CFFB6744381BC3B2EFEFEF97F" box="[159,288,1654,1680]" captionStart-0="FIGURE 17" captionStart-1="FIGURE 18" captionStart-2="FIGURE 19" captionStart-3="FIGURE 20" captionStartId-0="17.[151,250,1496,1520]" captionStartId-1="18.[151,250,1430,1454]" captionStartId-2="19.[151,250,1827,1851]" captionStartId-3="20.[151,250,1555,1579]" captionTargetBox-0="[151,1435,187,1472]" captionTargetBox-1="[151,1436,276,1407]" captionTargetBox-2="[174,1413,189,1804]" captionTargetBox-3="[151,1436,525,1532]" captionTargetId-0="figure-17@17.[151,1435,181,1472]" captionTargetId-1="figure-87@18.[151,1436,273,1407]" captionTargetId-2="figure-17@19.[151,1436,181,1804]" captionTargetId-3="figure-181@20.[151,1436,525,1532]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionTargetPageId-2="19" captionTargetPageId-3="20" captionText-0="FIGURE 17. Spatulaphorus incisus sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." captionText-1="FIGURE 18. Spatulaphorus incisus sp. nov., female: A—right leg I, dorsal aspect; B—right leg II, dorsal aspect." captionText-2="FIGURE 19. Spatulaphorus incisus sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." captionText-3="FIGURE 20. DIC micrographs of Spatulaphorus incisus sp. nov., female: A—general view dorsally; B—general view ventrally." pageId="16" pageNumber="353">Figs 1720</figureCitation>
)
</paragraph>
<paragraph id="8BFD3699FFB6744381B43BE6FAE1F936" blockId="16.[151,1437,1726,2005]" box="[151,1343,1726,1753]" pageId="16" pageNumber="353">
<emphasis id="B936EA8BFFB6744381B43BE6FEFFF937" bold="true" box="[151,289,1726,1752]" pageId="16" pageNumber="353">Description</emphasis>
. FEMALE. Body well sclerotized. Length of idiosoma 180 (180205), width 105 (105125).
</paragraph>
<paragraph id="8BFD3699FFB6744381E43BBAFCC9F83B" blockId="16.[151,1437,1726,2005]" pageId="16" pageNumber="353">
<emphasis id="B936EA8BFFB6744381E43BBAFE88F913" box="[199,342,1762,1788]" italics="true" pageId="16" pageNumber="353">Gnathosoma</emphasis>
. Gnathosomal capsule, excluding palps, subrectangular, length 17 (1720), width 18 (1820). Dorsum with one pair of needle-like cheliceral setae (
<emphasis id="B936EA8BFFB67443823D3A5EFC98F8CF" box="[798,838,1798,1824]" italics="true" pageId="16" pageNumber="353">cha</emphasis>
) 3 (34). Dorsal median apodeme present, poorly developed. Postpalpal setae (
<emphasis id="B936EA8BFFB6744380C13A73FE20F8AB" box="[482,510,1835,1860]" italics="true" pageId="16" pageNumber="353">pp</emphasis>
) 2 (2) spiniform, situated laterad setae
<emphasis id="B936EA8BFFB6744382E03A72FC35F8AB" box="[963,1003,1834,1860]" italics="true" pageId="16" pageNumber="353">cha</emphasis>
. Ventral gnathosoma with one pair of smooth, pointed subcapitular setae
<emphasis id="B936EA8BFFB6744383003A17FDE9F887" box="[547,567,1871,1896]" italics="true" pageId="16" pageNumber="353">m</emphasis>
7 (67) and round pits
<emphasis id="B936EA8BFFB6744382633A17FC90F887" box="[832,846,1871,1896]" italics="true" pageId="16" pageNumber="353">n</emphasis>
situated posteriad
<emphasis id="B936EA8BFFB6744385013A17FBE8F887" box="[1058,1078,1871,1896]" italics="true" pageId="16" pageNumber="353">m</emphasis>
. Palps with smooth and weakly blunt-tipped setae
<emphasis id="B936EA8BFFB67443804D3A2AFE44F863" box="[366,410,1906,1932]" italics="true" pageId="16" pageNumber="353">dFe</emphasis>
3 (3) and pointed
<emphasis id="B936EA8BFFB67443835F3A2AFD75F863" box="[636,683,1906,1932]" italics="true" pageId="16" pageNumber="353">dGe</emphasis>
8 (89) dorsolaterally. Palps ventrally each with mushroom-like accessory setigenous structure (
<emphasis id="B936EA8BFFB6744380DA3ACFFDC3F85F" box="[505,541,1943,1968]" italics="true" pageId="16" pageNumber="353">ass</emphasis>
) and tiny solenidion
<emphasis id="B936EA8BFFB6744382293ACFFCC0F85F" box="[778,798,1943,1968]" italics="true" pageId="16" pageNumber="353">ω</emphasis>
. Palps terminated with small blunt-tipped tibial claw and tiny eupathid-like seta. Pharyngeal pumps poorly visible.
</paragraph>
<caption id="DF3D6611FFB7744281B43880FB34FA1F" box="[151,1258,1496,1521]" pageId="17" pageNumber="354" startId="17.[151,250,1496,1520]" targetBox="[151,1435,187,1472]" targetPageId="17" targetType="figure">
<paragraph id="8BFD3699FFB7744281B43880FB34FA1F" blockId="17.[151,1258,1496,1521]" box="[151,1258,1496,1521]" pageId="17" pageNumber="354">
<emphasis id="B936EA8BFFB7744281B43880FEC7FA1F" bold="true" box="[151,281,1496,1520]" pageId="17" pageNumber="354">FIGURE 17</emphasis>
.
<taxonomicName id="4C424D1AFFB7744280053880FDD8FA1F" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[294,518,1496,1520]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="17" pageNumber="354" phylum="Arthropoda" rank="species" species="incisus" status="sp. nov.">
<emphasis id="B936EA8BFFB7744280053880FDD8FA1F" box="[294,518,1496,1520]" italics="true" pageId="17" pageNumber="354">Spatulaphorus incisus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFB77442832E3880FD83FA1F" bold="true" box="[525,605,1496,1520]" pageId="17" pageNumber="354">
<taxonomicNameLabel id="A20557F0FFB77442832E3880FD83FA1F" box="[525,605,1496,1520]" pageId="17" pageNumber="354" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—dorsum of body; B—venter of body. Legs omitted.
</paragraph>
</caption>
<paragraph id="8BFD3699FFB7744281E43B46FC86F8DA" blockId="17.[151,1437,1566,2025]" pageId="17" pageNumber="354">
<emphasis id="B936EA8BFFB7744281E43B46FE4BF9D7" box="[199,405,1566,1592]" italics="true" pageId="17" pageNumber="354">Idiosomal dorsum</emphasis>
(
<figureCitation id="13792A1CFFB7744280843B46FDCBF9D6" box="[423,533,1566,1593]" captionStart="FIGURE 17" captionStartId="17.[151,250,1496,1520]" captionTargetBox="[151,1435,187,1472]" captionTargetId="figure-17@17.[151,1435,181,1472]" captionTargetPageId="17" captionText="FIGURE 17. Spatulaphorus incisus sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." pageId="17" pageNumber="354">Figs 17A</figureCitation>
,
<figureCitation id="13792A1CFFB7744283073B47FD86F9D6" box="[548,600,1567,1593]" captionStart="FIGURE 20" captionStartId="20.[151,250,1555,1579]" captionTargetBox="[151,1436,525,1532]" captionTargetId="figure-181@20.[151,1436,525,1532]" captionTargetPageId="20" captionText="FIGURE 20. DIC micrographs of Spatulaphorus incisus sp. nov., female: A—general view dorsally; B—general view ventrally." pageId="17" pageNumber="354">20A</figureCitation>
). All dorsal shields with big puncta. Prodorsal shield with three pairs of setae (
<emphasis id="B936EA8BFFB7744281C33B1BFF24F9B2" box="[224,250,1603,1629]" italics="true" pageId="17" pageNumber="354">v1</emphasis>
,
<emphasis id="B936EA8BFFB77442802A3B1BFEFDF9B2" box="[265,291,1603,1629]" italics="true" pageId="17" pageNumber="354">v2</emphasis>
,
<emphasis id="B936EA8BFFB7744280123B1BFE89F9B2" box="[305,343,1603,1629]" italics="true" pageId="17" pageNumber="354">sc2</emphasis>
), one pair of weakly barbed capitate trichobothria (
<emphasis id="B936EA8BFFB7744282B83B1BFC1FF9B2" box="[923,961,1603,1629]" italics="true" pageId="17" pageNumber="354">sc1</emphasis>
) and one pair of round stigmata. Stigmata associated with well-developed tracheal trunks. All dorsal idiosomal setae blunt-tipped; setae
<emphasis id="B936EA8BFFB7744285FA3B3FFB2AF96E" box="[1241,1268,1639,1665]" italics="true" pageId="17" pageNumber="354">v2</emphasis>
smooth, other dorsal setae barbed. Alveoli of setae
<emphasis id="B936EA8BFFB7744283173BD3FD9EF94B" box="[564,576,1675,1700]" italics="true" pageId="17" pageNumber="354">e</emphasis>
and
<emphasis id="B936EA8BFFB77442835B3BD3FD5EF94A" box="[632,640,1675,1701]" italics="true" pageId="17" pageNumber="354">f</emphasis>
almost contiguous. Cupules
<emphasis id="B936EA8BFFB7744282E63BD3FC05F94B" box="[965,987,1675,1700]" italics="true" pageId="17" pageNumber="354">ia</emphasis>
on tergite D,
<emphasis id="B936EA8BFFB7744285553BD3FB4CF94B" box="[1142,1170,1675,1700]" italics="true" pageId="17" pageNumber="354">im</emphasis>
on tergite EF and
<emphasis id="B936EA8BFFB7744284403BD2FAA7F94B" box="[1379,1401,1674,1700]" italics="true" pageId="17" pageNumber="354">ih</emphasis>
on tergite H large, round, subequal. Lengths of dorsal setae:
<emphasis id="B936EA8BFFB7744282313BF7FCF2F926" box="[786,812,1711,1737]" italics="true" pageId="17" pageNumber="354">v1</emphasis>
15 (1520),
<emphasis id="B936EA8BFFB7744282983BF7FC08F926" box="[955,982,1711,1737]" italics="true" pageId="17" pageNumber="354">v2</emphasis>
9 (910),
<emphasis id="B936EA8BFFB77442856B3BF7FBB0F926" box="[1096,1134,1711,1737]" italics="true" pageId="17" pageNumber="354">sc2</emphasis>
28 (2632),
<emphasis id="B936EA8BFFB7744285DF3BF7FAC9F926" box="[1276,1303,1711,1737]" italics="true" pageId="17" pageNumber="354">c1</emphasis>
23 (2326),
<emphasis id="B936EA8BFFB7744281B43B8BFF6CF902" box="[151,178,1747,1773]" italics="true" pageId="17" pageNumber="354">c2</emphasis>
28 (2630),
<emphasis id="B936EA8BFFB7744280663B8AFE8DF903" box="[325,339,1746,1772]" italics="true" pageId="17" pageNumber="354">d</emphasis>
29 (2832),
<emphasis id="B936EA8BFFB7744280C43B8BFE2DF903" box="[487,499,1747,1772]" italics="true" pageId="17" pageNumber="354">e</emphasis>
19 (1924),
<emphasis id="B936EA8BFFB7744283A43B8BFD51F902" box="[647,655,1747,1773]" italics="true" pageId="17" pageNumber="354">f</emphasis>
33 (3139),
<emphasis id="B936EA8BFFB7744282003B8AFCE0F903" box="[803,830,1746,1772]" italics="true" pageId="17" pageNumber="354">h1</emphasis>
37 (3743),
<emphasis id="B936EA8BFFB7744282F03B8AFC31F903" box="[979,1007,1746,1772]" italics="true" pageId="17" pageNumber="354">h2</emphasis>
18 (1824). Distances between setae:
<emphasis id="B936EA8BFFB7744281B43BAFFF04F8FE" box="[151,218,1783,1809]" italics="true" pageId="17" pageNumber="354">v1v1</emphasis>
18 (1820),
<emphasis id="B936EA8BFFB7744280533BAFFE6CF8FE" box="[368,434,1783,1809]" italics="true" pageId="17" pageNumber="354">v2v2</emphasis>
33 (3339),
<emphasis id="B936EA8BFFB77442836A3BAFFD7FF8FE" box="[585,673,1783,1809]" italics="true" pageId="17" pageNumber="354">sc2sc2</emphasis>
38 (3845),
<emphasis id="B936EA8BFFB7744282143BAFFCA4F8FE" box="[823,890,1783,1809]" italics="true" pageId="17" pageNumber="354">c1c1</emphasis>
31 (3137),
<emphasis id="B936EA8BFFB77442852D3BAFFB8FF8FE" box="[1038,1105,1783,1809]" italics="true" pageId="17" pageNumber="354">c1c2</emphasis>
22 (2225),
<emphasis id="B936EA8BFFB7744285C43BAEFACFF8FF" box="[1255,1297,1782,1809]" italics="true" pageId="17" pageNumber="354">dd</emphasis>
51 (5162),
<emphasis id="B936EA8BFFB7744281B43A43FF64F8DA" box="[151,186,1819,1845]" italics="true" pageId="17" pageNumber="354">ef</emphasis>
5 (56),
<emphasis id="B936EA8BFFB7744280033A43FEE0F8DA" box="[288,318,1819,1845]" italics="true" pageId="17" pageNumber="354">ff</emphasis>
43 (4357),
<emphasis id="B936EA8BFFB7744280ED3A42FDCAF8DB" box="[462,532,1818,1845]" italics="true" pageId="17" pageNumber="354">h1h1</emphasis>
30 (3041),
<emphasis id="B936EA8BFFB7744283863A42FD35F8DB" box="[677,747,1818,1845]" italics="true" pageId="17" pageNumber="354">h1h2</emphasis>
8 (811).
</paragraph>
<paragraph id="8BFD3699FFB7744181E43A66FA46FF16" blockId="17.[151,1437,1566,2025]" lastBlockId="18.[151,1437,151,249]" lastPageId="18" lastPageNumber="355" pageId="17" pageNumber="354">
<emphasis id="B936EA8BFFB7744281E43A66FE5DF8B7" box="[199,387,1854,1880]" italics="true" pageId="17" pageNumber="354">Idiosomal venter</emphasis>
(
<figureCitation id="13792A1CFFB7744280B33A66FE27F8B6" box="[400,505,1854,1881]" captionStart="FIGURE 17" captionStartId="17.[151,250,1496,1520]" captionTargetBox="[151,1435,187,1472]" captionTargetId="figure-17@17.[151,1435,181,1472]" captionTargetPageId="17" captionText="FIGURE 17. Spatulaphorus incisus sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." pageId="17" pageNumber="354">Figs 17B</figureCitation>
,
<figureCitation id="13792A1CFFB7744283203A67FDE8F8B6" box="[515,566,1855,1881]" captionStart="FIGURE 20" captionStartId="20.[151,250,1555,1579]" captionTargetBox="[151,1436,525,1532]" captionTargetId="figure-181@20.[151,1436,525,1532]" captionTargetPageId="20" captionText="FIGURE 20. DIC micrographs of Spatulaphorus incisus sp. nov., female: A—general view dorsally; B—general view ventrally." pageId="17" pageNumber="354">20B</figureCitation>
). Coxisternal fields I-II laterally with sparsely distributed big puncta. Setae
<emphasis id="B936EA8BFFB77442845A3A66FA48F8B7" box="[1401,1430,1854,1880]" italics="true" pageId="17" pageNumber="354">1b</emphasis>
,
<emphasis id="B936EA8BFFB7744281B43A3BFF6FF892" box="[151,177,1891,1917]" italics="true" pageId="17" pageNumber="354">2c</emphasis>
, and
<emphasis id="B936EA8BFFB7744281CD3A3AFED4F893" box="[238,266,1890,1916]" italics="true" pageId="17" pageNumber="354">4b</emphasis>
pointed; other ventral setae blunt-tipped; setae
<emphasis id="B936EA8BFFB7744282393A3AFCE8F893" box="[794,822,1890,1916]" italics="true" pageId="17" pageNumber="354">1b</emphasis>
,
<emphasis id="B936EA8BFFB7744282603A3BFC83F892" box="[835,861,1891,1917]" italics="true" pageId="17" pageNumber="354">2c</emphasis>
, and
<emphasis id="B936EA8BFFB7744282B93A3AFC68F893" box="[922,950,1890,1916]" italics="true" pageId="17" pageNumber="354">4b</emphasis>
with few weak barbs; setae
<emphasis id="B936EA8BFFB7744285CC3A3BFAF3F892" box="[1263,1325,1891,1917]" italics="true" pageId="17" pageNumber="354">ps1-3</emphasis>
distinctly barbed, other ventral setae smooth. Bases of setae
<emphasis id="B936EA8BFFB7744283FC3ADFFCD8F84E" box="[735,774,1927,1953]" italics="true" pageId="17" pageNumber="354">ps3</emphasis>
situated distinctly anteriad bases of setae
<emphasis id="B936EA8BFFB7744285CE3ADFFAEDF84E" box="[1261,1331,1927,1953]" italics="true" pageId="17" pageNumber="354">ps1-2.</emphasis>
Ap1 and ap2 well-developed and joined with thick appr; apsej developed only laterally; ap3 weak, not reaching appo; ap4 well developed, exceeding beyond bases of setae
<emphasis id="B936EA8BFFB7744283E03A96FD01F807" box="[707,735,1998,2024]" italics="true" pageId="17" pageNumber="354">3b</emphasis>
. Ap5 very short and thin. Ags small, cup-like; pgs small, oval. Posterior margin of posterior sternal plate deeply incised in middle part. Posterior margin of aggenital plate rounded. Lengths of ventral setae:
<emphasis id="B936EA8BFFB4744180863DE3FE1EFF3A" box="[421,448,187,213]" italics="true" pageId="18" pageNumber="355">1a</emphasis>
8 (811),
<emphasis id="B936EA8BFFB47441830D3DE3FD97FF3A" box="[558,585,187,213]" italics="true" pageId="18" pageNumber="355">1b</emphasis>
17 (1719),
<emphasis id="B936EA8BFFB4744183F03DE3FD32FF3A" box="[723,748,187,213]" italics="true" pageId="18" pageNumber="355">1с</emphasis>
10 (1012),
<emphasis id="B936EA8BFFB4744182553DE3FC4FFF3A" box="[886,913,187,213]" italics="true" pageId="18" pageNumber="355">2a</emphasis>
11 (1113),
<emphasis id="B936EA8BFFB47441853B3DE3FBECFF3A" box="[1048,1074,187,213]" italics="true" pageId="18" pageNumber="355">2с</emphasis>
12 (1217),
<emphasis id="B936EA8BFFB47441859F3DE3FB09FF3A" box="[1212,1239,187,213]" italics="true" pageId="18" pageNumber="355">3a</emphasis>
13 (1315),
<emphasis id="B936EA8BFFB4744184433DE3FAA5FF3A" box="[1376,1403,187,213]" italics="true" pageId="18" pageNumber="355">3b</emphasis>
10 (1014),
<emphasis id="B936EA8BFFB4744181D93D87FECDFF16" box="[250,275,223,249]" italics="true" pageId="18" pageNumber="355">3c</emphasis>
12 (1214),
<emphasis id="B936EA8BFFB4744180BD3D87FE67FF16" box="[414,441,223,249]" italics="true" pageId="18" pageNumber="355">4a</emphasis>
12 (1214),
<emphasis id="B936EA8BFFB4744183673D87FD81FF16" box="[580,607,223,249]" italics="true" pageId="18" pageNumber="355">4b</emphasis>
21 (2129),
<emphasis id="B936EA8BFFB4744183CA3D87FCDCFF16" box="[745,770,223,249]" italics="true" pageId="18" pageNumber="355">4c</emphasis>
13 (1315),
<emphasis id="B936EA8BFFB4744182AD3D87FC6DFF16" box="[910,947,223,249]" italics="true" pageId="18" pageNumber="355">ps1</emphasis>
18 (1820),
<emphasis id="B936EA8BFFB47441851D3D87FBBDFF16" box="[1086,1123,223,249]" italics="true" pageId="18" pageNumber="355">ps2</emphasis>
18 (1820),
<emphasis id="B936EA8BFFB4744185CD3D87FACDFF16" box="[1262,1299,223,249]" italics="true" pageId="18" pageNumber="355">ps3</emphasis>
19 (1922).
</paragraph>
<caption id="DF3D6611FFB4744181B438CEFAC9FA41" box="[151,1303,1430,1455]" pageId="18" pageNumber="355" startId="18.[151,250,1430,1454]" targetBox="[151,1436,276,1407]" targetPageId="18" targetType="figure">
<paragraph id="8BFD3699FFB4744181B438CEFAC9FA41" blockId="18.[151,1303,1430,1455]" box="[151,1303,1430,1455]" pageId="18" pageNumber="355">
<emphasis id="B936EA8BFFB4744181B438CEFEC7FA41" bold="true" box="[151,281,1430,1454]" pageId="18" pageNumber="355">FIGURE 18</emphasis>
.
<taxonomicName id="4C424D1AFFB47441800538CFFDD8FA41" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[294,518,1431,1455]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="18" pageNumber="355" phylum="Arthropoda" rank="species" species="incisus" status="sp. nov.">
<emphasis id="B936EA8BFFB47441800538CFFDD8FA41" box="[294,518,1431,1455]" italics="true" pageId="18" pageNumber="355">Spatulaphorus incisus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFB47441832E38CFFD83FA40" bold="true" box="[525,605,1431,1455]" pageId="18" pageNumber="355">
<taxonomicNameLabel id="A20557F0FFB47441832E38CFFD83FA40" box="[525,605,1431,1455]" pageId="18" pageNumber="355" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—right leg I, dorsal aspect; B—right leg II, dorsal aspect.
</paragraph>
</caption>
<paragraph id="8BFD3699FFB4744181E43886FB8DF800" blockId="18.[151,1437,1501,2031]" pageId="18" pageNumber="355">
<emphasis id="B936EA8BFFB4744181E43886FF22FA18" box="[199,252,1502,1527]" italics="true" pageId="18" pageNumber="355">Legs</emphasis>
(
<figureCitation id="13792A1CFFB47441802F3885FEBAFA18" box="[268,356,1501,1527]" captionStart="FIGURE 18" captionStartId="18.[151,250,1430,1454]" captionTargetBox="[151,1436,276,1407]" captionTargetId="figure-87@18.[151,1436,273,1407]" captionTargetPageId="18" captionText="FIGURE 18. Spatulaphorus incisus sp. nov., female: A—right leg I, dorsal aspect; B—right leg II, dorsal aspect." pageId="18" pageNumber="355">Figs 18</figureCitation>
,
<figureCitation id="13792A1CFFB4744180523885FE4EFA18" box="[369,400,1501,1527]" captionStart="FIGURE 19" captionStartId="19.[151,250,1827,1851]" captionTargetBox="[174,1413,189,1804]" captionTargetId="figure-17@19.[151,1436,181,1804]" captionTargetPageId="19" captionText="FIGURE 19. Spatulaphorus incisus sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." pageId="18" pageNumber="355">19</figureCitation>
). Setation of legs as in
<taxonomicName id="4C424D1AFFB4744183B93885FC9EFA18" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[666,832,1501,1527]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="18" pageNumber="355" phylum="Arthropoda" rank="species" species="brevisetosus">
<emphasis id="B936EA8BFFB4744183B93885FC9EFA18" box="[666,832,1501,1527]" italics="true" pageId="18" pageNumber="355">S. brevisetosus</emphasis>
</taxonomicName>
. Leg I (
<figureCitation id="13792A1CFFB4744182BE3885FBD9FA18" box="[925,1031,1501,1527]" captionStart="FIGURE 18" captionStartId="18.[151,250,1430,1454]" captionTargetBox="[151,1436,276,1407]" captionTargetId="figure-87@18.[151,1436,273,1407]" captionTargetPageId="18" captionText="FIGURE 18. Spatulaphorus incisus sp. nov., female: A—right leg I, dorsal aspect; B—right leg II, dorsal aspect." pageId="18" pageNumber="355">Fig. 18A</figureCitation>
). Tibiotarsus slightly enlarged, with large and long claw; tibiotarsus internally with oval strongly sclerotized structure near solenidion
<emphasis id="B936EA8BFFB4744184253B59FAFAF9F4" box="[1286,1316,1537,1563]" italics="true" pageId="18" pageNumber="355">φ2</emphasis>
. Setae (
<emphasis id="B936EA8BFFB4744184A63B5AFA4DF9F4" box="[1413,1427,1538,1563]" italics="true" pageId="18" pageNumber="355">u</emphasis>
) consolidated into a bifid horn-like structure opposing claw. Tarsal claw weakly blunt-tipped. Seta
<emphasis id="B936EA8BFFB47441843E3B7DFAF5F9D0" box="[1309,1323,1573,1599]" italics="true" pageId="18" pageNumber="355">d</emphasis>
of femur spatulate, with short subterminal projection. Setae (
<emphasis id="B936EA8BFFB4744183FD3B11FD38F98C" box="[734,742,1609,1635]" italics="true" pageId="18" pageNumber="355">l</emphasis>
) of genu and
<emphasis id="B936EA8BFFB4744182A63B11FC7EF98C" box="[901,928,1609,1635]" italics="true" pageId="18" pageNumber="355">v”</emphasis>
of tibiotarsus weakly barbed, other leg setae (except eupathidia
<emphasis id="B936EA8BFFB47441804D3B35FE5BF968" box="[366,389,1645,1671]" italics="true" pageId="18" pageNumber="355">p</emphasis>
,
<emphasis id="B936EA8BFFB4744180B73B35FE6FF968" box="[404,433,1645,1671]" italics="true" pageId="18" pageNumber="355">p”</emphasis>
,
<emphasis id="B936EA8BFFB4744180E33B35FE00F968" box="[448,478,1645,1671]" italics="true" pageId="18" pageNumber="355">tc</emphasis>
,
<emphasis id="B936EA8BFFB4744180CF3B35FDCEF968" box="[492,528,1645,1671]" italics="true" pageId="18" pageNumber="355">tc”</emphasis>
,
<emphasis id="B936EA8BFFB47441833C3B35FDE6F968" box="[543,568,1645,1671]" italics="true" pageId="18" pageNumber="355">ft</emphasis>
,
<emphasis id="B936EA8BFFB4744183653B35FDB8F968" box="[582,614,1645,1671]" italics="true" pageId="18" pageNumber="355">ft”</emphasis>
) smooth. Setae
<emphasis id="B936EA8BFFB47441823F3B35FCF4F968" box="[796,810,1645,1671]" italics="true" pageId="18" pageNumber="355">l</emphasis>
of femur, (
<emphasis id="B936EA8BFFB4744182883B35FC6DF968" box="[939,947,1645,1671]" italics="true" pageId="18" pageNumber="355">l</emphasis>
) of genu and
<emphasis id="B936EA8BFFB4744185723B35FB83F968" box="[1105,1117,1645,1671]" italics="true" pageId="18" pageNumber="355">k</emphasis>
of tibiotarsus weakly blunt-tipped; other leg setae (except eupathidia) pointed. Lengths of solenidia
<emphasis id="B936EA8BFFB4744182E23BC9FC3CF944" box="[961,994,1681,1707]" italics="true" pageId="18" pageNumber="355">ω1</emphasis>
6 (67),
<emphasis id="B936EA8BFFB47441856A3BC9FBB5F944" box="[1097,1131,1681,1707]" italics="true" pageId="18" pageNumber="355">ω2</emphasis>
4 (4),
<emphasis id="B936EA8BFFB4744185953BC9FB0DF944" box="[1206,1235,1681,1707]" italics="true" pageId="18" pageNumber="355">φ1</emphasis>
6 (67),
<emphasis id="B936EA8BFFB4744184193BC9FA86F944" box="[1338,1368,1681,1707]" italics="true" pageId="18" pageNumber="355">φ2</emphasis>
4 (4); solenidion
<emphasis id="B936EA8BFFB4744180313BEDFEEDF920" box="[274,307,1717,1743]" italics="true" pageId="18" pageNumber="355">ω1</emphasis>
digitiform; solenidion
<emphasis id="B936EA8BFFB4744183113BEDFD91F920" box="[562,591,1717,1743]" italics="true" pageId="18" pageNumber="355">φ1</emphasis>
thick, clavate; other solenidia weakly clavate. Leg II (
<figureCitation id="13792A1CFFB4744185863BEDFAD4F920" box="[1189,1290,1717,1743]" captionStart="FIGURE 18" captionStartId="18.[151,250,1430,1454]" captionTargetBox="[151,1436,276,1407]" captionTargetId="figure-87@18.[151,1436,273,1407]" captionTargetPageId="18" captionText="FIGURE 18. Spatulaphorus incisus sp. nov., female: A—right leg I, dorsal aspect; B—right leg II, dorsal aspect." pageId="18" pageNumber="355">Fig. 18B</figureCitation>
). Tarsus with thickened basally claws and pad-like empodium. Solenidion
<emphasis id="B936EA8BFFB4744182643B82FC85F91C" box="[839,859,1754,1779]" italics="true" pageId="18" pageNumber="355">ω</emphasis>
5 (56) weakly clavate, solenidion
<emphasis id="B936EA8BFFB4744185D13B82FADFF91C" box="[1266,1281,1754,1779]" italics="true" pageId="18" pageNumber="355">φ</emphasis>
absent. Setae
<emphasis id="B936EA8BFFB4744181B43BA5FF76F8F8" box="[151,168,1789,1815]" italics="true" pageId="18" pageNumber="355">l</emphasis>
of femur,
<emphasis id="B936EA8BFFB47441803D3BA5FEEBF8F8" box="[286,309,1789,1815]" italics="true" pageId="18" pageNumber="355">u</emphasis>
, and
<emphasis id="B936EA8BFFB4744180573BA5FE46F8F8" box="[372,408,1789,1815]" italics="true" pageId="18" pageNumber="355">tc”</emphasis>
of tarsus smooth, other setae barbed; setae
<emphasis id="B936EA8BFFB4744182A13BA5FC4EF8F8" box="[898,912,1789,1815]" italics="true" pageId="18" pageNumber="355">d</emphasis>
and
<emphasis id="B936EA8BFFB4744182E43BA5FC06F8F8" box="[967,984,1789,1815]" italics="true" pageId="18" pageNumber="355">l</emphasis>
of femur weakly blunt-tipped; other leg setae pointed. Leg III (
<figureCitation id="13792A1CFFB4744180BA3A79FDDCF8D4" box="[409,514,1825,1851]" captionStart="FIGURE 19" captionStartId="19.[151,250,1827,1851]" captionTargetBox="[174,1413,189,1804]" captionTargetId="figure-17@19.[151,1436,181,1804]" captionTargetPageId="19" captionText="FIGURE 19. Spatulaphorus incisus sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." pageId="18" pageNumber="355">Fig. 19A</figureCitation>
). Femur divided into basi- and telofemur. Claws and empodium of same shape as on tarsus II. Setae
<emphasis id="B936EA8BFFB4744180453A1DFEA2F8B0" box="[358,380,1861,1887]" italics="true" pageId="18" pageNumber="355">v</emphasis>
of trochanter and
<emphasis id="B936EA8BFFB47441836B3A1DFD81F8B0" box="[584,607,1861,1887]" italics="true" pageId="18" pageNumber="355">u</emphasis>
of tarsus smooth, other setae barbed; setae
<emphasis id="B936EA8BFFB4744185663A1DFB85F8B0" box="[1093,1115,1861,1887]" italics="true" pageId="18" pageNumber="355">v</emphasis>
of trochanter,
<emphasis id="B936EA8BFFB4744185DE3A1DFAD5F8B0" box="[1277,1291,1861,1887]" italics="true" pageId="18" pageNumber="355">d</emphasis>
,
<emphasis id="B936EA8BFFB47441843A3A1DFAF1F8B0" box="[1305,1327,1861,1887]" italics="true" pageId="18" pageNumber="355">v</emphasis>
of femur, and
<emphasis id="B936EA8BFFB4744181E53A31FF09F86C" box="[198,215,1897,1923]" italics="true" pageId="18" pageNumber="355">l</emphasis>
of genu blunt-tipped, other leg setae pointed; solenidion
<emphasis id="B936EA8BFFB4744182723A32FCBEF86C" box="[849,864,1898,1923]" italics="true" pageId="18" pageNumber="355">φ</emphasis>
absent. Leg IV (
<figureCitation id="13792A1CFFB47441853B3A31FBA9F86C" box="[1048,1143,1897,1923]" captionStart="FIGURE 19" captionStartId="19.[151,250,1827,1851]" captionTargetBox="[174,1413,189,1804]" captionTargetId="figure-17@19.[151,1436,181,1804]" captionTargetPageId="19" captionText="FIGURE 19. Spatulaphorus incisus sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." pageId="18" pageNumber="355">Fig 19B</figureCitation>
). Femur divided into basi- and telofemur. Claws simple, empodium as on tarsi II and III. Setae
<emphasis id="B936EA8BFFB4744182AC3AD5FC7BF848" box="[911,933,1933,1959]" italics="true" pageId="18" pageNumber="355">v</emphasis>
of trochanter and
<emphasis id="B936EA8BFFB4744185533AD5FB48F848" box="[1136,1174,1933,1959]" italics="true" pageId="18" pageNumber="355">pl”</emphasis>
of tarsus smooth; other setae barbed; setae
<emphasis id="B936EA8BFFB4744180533AE9FE58F824" box="[368,390,1969,1995]" italics="true" pageId="18" pageNumber="355">v</emphasis>
of trochanter,
<emphasis id="B936EA8BFFB4744183093AE9FDE6F824" box="[554,568,1969,1995]" italics="true" pageId="18" pageNumber="355">d</emphasis>
,
<emphasis id="B936EA8BFFB4744183653AE9FD82F824" box="[582,604,1969,1995]" italics="true" pageId="18" pageNumber="355">v</emphasis>
of femur,
<emphasis id="B936EA8BFFB4744183F13AE9FD3DF824" box="[722,739,1969,1995]" italics="true" pageId="18" pageNumber="355">l</emphasis>
,
<emphasis id="B936EA8BFFB4744183D23AE9FCD2F824" box="[753,780,1969,1995]" italics="true" pageId="18" pageNumber="355">v”</emphasis>
of tibia and
<emphasis id="B936EA8BFFB4744182B83AE9FC1FF824" box="[923,961,1969,1995]" italics="true" pageId="18" pageNumber="355">pl”</emphasis>
of tarsus blunt-tipped; tips of setae
<emphasis id="B936EA8BFFB4744184703AE9FAB4F824" box="[1363,1386,1969,1995]" italics="true" pageId="18" pageNumber="355">u</emphasis>
,
<emphasis id="B936EA8BFFB47441845B3AE9FA42F824" box="[1400,1436,1969,1995]" italics="true" pageId="18" pageNumber="355">tc”</emphasis>
and
<emphasis id="B936EA8BFFB4744181E43A8DFF2EF800" box="[199,240,2005,2031]" italics="true" pageId="18" pageNumber="355">pv”</emphasis>
slightly thickened and flattened; other leg setae pointed; solenidion
<emphasis id="B936EA8BFFB4744182CD3A8EFC23F800" box="[1006,1021,2006,2031]" italics="true" pageId="18" pageNumber="355">φ</emphasis>
absent.
</paragraph>
<caption id="DF3D6611FFB5744081B43A7BFAF0F8D4" box="[151,1326,1827,1852]" pageId="19" pageNumber="356" startId="19.[151,250,1827,1851]" targetBox="[174,1413,189,1804]" targetPageId="19" targetType="figure">
<paragraph id="8BFD3699FFB5744081B43A7BFAF0F8D4" blockId="19.[151,1326,1827,1852]" box="[151,1326,1827,1852]" pageId="19" pageNumber="356">
<emphasis id="B936EA8BFFB5744081B43A7BFEC7F8D4" bold="true" box="[151,281,1827,1851]" pageId="19" pageNumber="356">FIGURE 19</emphasis>
.
<taxonomicName id="4C424D1AFFB5744080053A7BFDD8F8D4" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[294,518,1827,1851]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="19" pageNumber="356" phylum="Arthropoda" rank="species" species="incisus" status="sp. nov.">
<emphasis id="B936EA8BFFB5744080053A7BFDD8F8D4" box="[294,518,1827,1851]" italics="true" pageId="19" pageNumber="356">Spatulaphorus incisus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFB57440832E3A7CFD83F8D3" bold="true" box="[525,605,1828,1852]" pageId="19" pageNumber="356">
<taxonomicNameLabel id="A20557F0FFB57440832E3A7CFD83F8D3" box="[525,605,1828,1852]" pageId="19" pageNumber="356" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect.
</paragraph>
</caption>
<paragraph id="8BFD3699FFB5744081EB3A32FE4FF86B" blockId="19.[151,1436,1898,2032]" box="[200,401,1898,1924]" pageId="19" pageNumber="356">MALE unknown.</paragraph>
</subSubSection>
<subSubSection id="C3586512FFB5744781E43AD6FD9BFF3A" lastPageId="20" lastPageNumber="357" pageId="19" pageNumber="356" type="materials_examined">
<paragraph id="8BFD3699FFB5744081E43AD6FBF1F823" blockId="19.[151,1436,1898,2032]" pageId="19" pageNumber="356">
<emphasis id="B936EA8BFFB5744081E43AD6FEB0F848" bold="true" box="[199,366,1933,1960]" pageId="19" pageNumber="356">Type material</emphasis>
. Female
<materialsCitation id="3B2A3CC4FFB5744080F63AD6FD75F823" collectingDate="2017-02-26" collectionCode="ZISP, ESEC, CEMT" country="Brazil" location="Rio Ronuro" pageId="19" pageNumber="356" specimenCount="1" stateProvince="Mato Grosso" typeStatus="holotype">
<typeStatus id="54F9883BFFB5744080F63AD6FDE4F847" box="[469,570,1934,1960]" pageId="19" pageNumber="356" type="holotype">holotype</typeStatus>
, slide
<collectionCode id="ED53AE5CFFB5744083A33AD6FD67F847" box="[640,697,1934,1960]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="19" pageNumber="356">ZISP</collectionCode>
T-Pygm-014,
<collectingCountry id="F3557609FFB5744082793AD6FC7FF847" box="[858,929,1934,1960]" name="Brazil" pageId="19" pageNumber="356">Brazil</collectingCountry>
,
<collectingRegion id="4986F87BFFB57440828F3AD6FB9EF847" box="[940,1088,1934,1960]" country="Brazil" name="Mato Grosso" pageId="19" pageNumber="356">Mato Grosso</collectingRegion>
,
<collectionCode id="ED53AE5CFFB5744085683AD6FB51F847" box="[1099,1167,1934,1960]" country="Egypt" httpUri="http://grbio.org/cool/4cts-0w01" name="Entomological Society of Egypt" pageId="19" pageNumber="356">ESEC</collectionCode>
<location id="8E9D6042FFB5744085B53AD6FAC3F847" LSID="urn:lsid:plazi:treatment:03EB878FFFB6744781B43B0AFC07FE1A:8E9D6042FFB5744085B53AD6FAC3F847" box="[1174,1309,1934,1960]" country="Brazil" name="Rio Ronuro" pageId="19" pageNumber="356" stateProvince="Mato Grosso">Rio Ronuro</location>
,
<date id="FFFC1059FFB57440840B3AD6FA42F847" box="[1320,1436,1934,1960]" pageId="19" pageNumber="356" value="2017-02-26">
<collectingDate id="EFB8E9B1FFB57440840B3AD6FA42F847" box="[1320,1436,1934,1960]" pageId="19" pageNumber="356" value="2017-02-26">26.II.2017</collectingDate>
</date>
(
<collectionCode id="ED53AE5CFFB5744081BC3AEAFF2CF823" box="[159,242,1970,1996]" pageId="19" pageNumber="356">CEMT</collectionCode>
),
<collectedFrom id="385E7564FFB5744080253AEAFD75F823" box="[262,683,1970,1996]" pageId="19" pageNumber="356">
on
<taxonomicName id="4C424D1AFFB57440800A3AEAFD75F823" authority="(MacLeay)" baseAuthorityName="MacLeay" box="[297,683,1970,1996]" class="Insecta" family="Scarabaeidae" genus="Sulcophanaeus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="356" phylum="Arthropoda" rank="species" species="faunus">
<emphasis id="B936EA8BFFB57440800A3AEAFDFAF823" box="[297,548,1970,1996]" italics="true" pageId="19" pageNumber="356">Sulcophanaeus faunus</emphasis>
(MacLeay)
</taxonomicName>
</collectedFrom>
</materialsCitation>
;
<materialsCitation id="3B2A3CC4FFB5744083953AEAFBF5F823" box="[694,1067,1970,1996]" collectingDate="2017-02-26" collectionCode="Male" country="Brazil" location="Rio Ronuro" pageId="19" pageNumber="356" specimenCount="42" specimenCount-female="42" stateProvince="Mato Grosso" typeStatus="paratype">
<typeStatus id="54F9883BFFB5744083953AEAFCF8F823" box="[694,806,1970,1996]" pageId="19" pageNumber="356" type="paratype">paratypes</typeStatus>
:
<specimenCount id="9D44FD10FFB5744082133AEAFC73F823" box="[816,941,1970,1996]" count="42" pageId="19" pageNumber="356" type="female">42 females</specimenCount>
, same data
</materialsCitation>
.
</paragraph>
<paragraph id="8BFD3699FFB5744781E43A8EFD9BFF3A" blockId="19.[151,1436,1898,2032]" lastBlockId="20.[151,1437,151,501]" lastPageId="20" lastPageNumber="357" pageId="19" pageNumber="356">
<emphasis id="B936EA8BFFB5744081E43A8EFE58F800" bold="true" box="[199,390,2005,2032]" pageId="19" pageNumber="356">Type deposition</emphasis>
.
<materialsCitation id="3B2A3CC4FFB5744780B43A8EFE25FF5E" country="Russia" lastPageId="20" lastPageNumber="357" location="Zoological Institute" pageId="19" pageNumber="356" specimenCount="4" stateProvince="Saint Petersburg" typeStatus="holotype">
The
<typeStatus id="54F9883BFFB5744080EF3A8EFDF0F81F" box="[460,558,2006,2032]" pageId="19" pageNumber="356" type="holotype">holotype</typeStatus>
and
<specimenCount id="9D44FD10FFB5744083493A8EFCD3F81F" box="[618,781,2006,2032]" count="4" pageId="19" pageNumber="356" type="generic" typeStatus="paratype">four paratypes</specimenCount>
are deposited in the collection of
<location id="8E9D6042FFB5744085BF3A8EFAA5F81F" LSID="urn:lsid:plazi:treatment:03EB878FFFB6744781B43B0AFC07FE1A:8E9D6042FFB5744085BF3A8EFAA5F81F" box="[1180,1403,2006,2032]" country="Russia" name="Zoological Institute" pageId="19" pageNumber="356" stateProvince="Saint Petersburg">Zoological Institute</location>
of RAS,
<collectingRegion id="4986F87BFFB2744781FC3DCFFE40FF5E" box="[223,414,151,177]" country="Russia" name="Saint Petersburg" pageId="20" pageNumber="357">Saint Petersburg</collectingRegion>
,
<collectingCountry id="F3557609FFB27447808E3DCFFE25FF5E" box="[429,507,151,177]" name="Russia" pageId="20" pageNumber="357">Russia</collectingCountry>
</materialsCitation>
;
<materialsCitation id="3B2A3CC4FFB2744783293DCFFAC5FF5E" box="[522,1307,151,177]" country="Russia" location="Tyumen State" pageId="20" pageNumber="357" specimenCount="1" stateProvince="Tyumen'" typeStatus="paratype">
other
<typeStatus id="54F9883BFFB27447836D3DCFFD67FF5E" box="[590,697,151,177]" pageId="20" pageNumber="357" type="paratype">paratypes</typeStatus>
are deposited in the collection of the
<collectingRegion id="4986F87BFFB27447855E3DCFFAC5FF5E" box="[1149,1307,151,177]" country="Russia" name="Tyumen'" pageId="20" pageNumber="357">Tyumen State</collectingRegion>
</materialsCitation>
<materialsCitation id="3B2A3CC4FFB2744784053DCFFD9EFF3A" country="Russia" location="University Museum of Zoology" pageId="20" pageNumber="357" specimenCount="1" stateProvince="Tyumen'" typeStatus="paratype">
<location id="8E9D6042FFB2744784053DCFFEA1FF3A" LSID="urn:lsid:plazi:treatment:03EB878FFFB6744781B43B0AFC07FE1A:8E9D6042FFB2744784053DCFFEA1FF3A" country="Russia" name="University Museum of Zoology" pageId="20" pageNumber="357" stateProvince="Tyumen'">University Museum of Zoology</location>
,
<collectingRegion id="4986F87BFFB2744780AA3DE3FE39FF3A" box="[393,487,187,213]" country="Russia" name="Tyumen'" pageId="20" pageNumber="357">Tyumen</collectingRegion>
,
<collectingCountry id="F3557609FFB2744780D13DE3FD9EFF3A" box="[498,576,187,213]" name="Russia" pageId="20" pageNumber="357">Russia</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3586512FFB2744781E43D86FD70FE42" pageId="20" pageNumber="357" type="diagnosis">
<paragraph id="8BFD3699FFB2744781E43D86FD70FE42" blockId="20.[151,1437,151,501]" pageId="20" pageNumber="357">
<emphasis id="B936EA8BFFB2744781E43D86FE1BFF17" bold="true" box="[199,453,222,248]" pageId="20" pageNumber="357">Differential diagnosis</emphasis>
. The new species is very similar to
<taxonomicName id="4C424D1AFFB2744782713D87FAF9FF16" authority="Khaustov and Frolov, 2017" authorityName="Khaustov and Frolov" authorityYear="2017" box="[850,1319,223,249]" class="Insecta" family="Scarabaeidae" genus="Sulcophanaeus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="357" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="B936EA8BFFB2744782713D87FCBEFF16" box="[850,864,223,249]" italics="true" pageId="20" pageNumber="357">S</emphasis>
.
<emphasis id="B936EA8BFFB27447824D3D87FC31FF16" box="[878,1007,223,249]" italics="true" pageId="20" pageNumber="357">brasiliensis</emphasis>
<bibRefCitation id="EFD34B68FFB2744782D63D87FAF9FF16" author="Khaustov, A. A. &amp; Frolov, A. V." box="[1013,1319,223,249]" pageId="20" pageNumber="357" pagination="501 - 521" refId="ref16128" refString="Khaustov, A. A. &amp; Frolov, A. V. (2017) New species of heterostigmatic mites (Acari: Heterostigmata: Athyreacaridae, Dolichocybidae, Pygmephoridae) associated with scarab beetles (Coleoptera: Geotrupidae, Scarabaeidae) from Brazil. Zootaxa, 4294 (5), 501 - 521. https: // doi. org / 10.11646 / zootaxa. 4294.5.1" type="journal article" year="2017">Khaustov and Frolov, 2017</bibRefCitation>
</taxonomicName>
, described from
<collectingCountry id="F3557609FFB2744781F73C5BFEC6FEF2" box="[212,280,259,285]" name="Brazil" pageId="20" pageNumber="357">Brazil</collectingCountry>
(
<bibRefCitation id="EFD34B68FFB2744780053C5BFD9DFEF2" author="Khaustov, A. A. &amp; Frolov, A. V." box="[294,579,259,285]" pageId="20" pageNumber="357" pagination="501 - 521" refId="ref16128" refString="Khaustov, A. A. &amp; Frolov, A. V. (2017) New species of heterostigmatic mites (Acari: Heterostigmata: Athyreacaridae, Dolichocybidae, Pygmephoridae) associated with scarab beetles (Coleoptera: Geotrupidae, Scarabaeidae) from Brazil. Zootaxa, 4294 (5), 501 - 521. https: // doi. org / 10.11646 / zootaxa. 4294.5.1" type="journal article" year="2017">Khaustov &amp; Frolov 2017</bibRefCitation>
), in having unmodified setae on tarsi II and III, presence of big puncta on the dorsal idiosomal sclerites, and basally thickened claws on tarsi II and III. The new species differs from
<taxonomicName id="4C424D1AFFB2744784223C7FFA42FEAE" authorityName="Khaustov and Frolov" authorityYear="2017" box="[1281,1436,295,321]" class="Insecta" family="Scarabaeidae" genus="Sulcophanaeus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="357" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="B936EA8BFFB2744784223C7FFAD1FEAE" box="[1281,1295,295,321]" italics="true" pageId="20" pageNumber="357">S</emphasis>
.
<emphasis id="B936EA8BFFB2744784383C7FFA42FEAE" box="[1307,1436,295,321]" italics="true" pageId="20" pageNumber="357">brasiliensis</emphasis>
</taxonomicName>
in having setae
<emphasis id="B936EA8BFFB2744780653C13FE5AFE8A" box="[326,388,331,357]" italics="true" pageId="20" pageNumber="357">ps1-2</emphasis>
barbed and weakly blunt-tipped (vs. setae
<emphasis id="B936EA8BFFB2744782473C13FC7CFE8A" box="[868,930,331,357]" italics="true" pageId="20" pageNumber="357">ps1-2</emphasis>
smooth and pointed in
<taxonomicName id="4C424D1AFFB2744785893C13FA99FE8A" authorityName="Khaustov and Frolov" authorityYear="2017" box="[1194,1351,331,357]" class="Insecta" family="Scarabaeidae" genus="Sulcophanaeus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="357" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="B936EA8BFFB2744785893C13FA99FE8A" box="[1194,1351,331,357]" italics="true" pageId="20" pageNumber="357">S. brasiliensis</emphasis>
</taxonomicName>
) and in having an unusually deeply incised posterior margin of the posterior sternal plate (vs. posterior margin of posterior sternal plate slightly concave in
<taxonomicName id="4C424D1AFFB2744783223CCBFD40FE42" authorityName="Khaustov and Frolov" authorityYear="2017" box="[513,670,403,429]" class="Insecta" family="Scarabaeidae" genus="Sulcophanaeus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="357" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="B936EA8BFFB2744783223CCBFD40FE42" box="[513,670,403,429]" italics="true" pageId="20" pageNumber="357">S. brasiliensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3586512FFB2744781E43CEFFC07FE1A" pageId="20" pageNumber="357" type="etymology">
<paragraph id="8BFD3699FFB2744781E43CEFFC07FE1A" blockId="20.[151,1437,151,501]" pageId="20" pageNumber="357">
<emphasis id="B936EA8BFFB2744781E43CEFFE96FE3E" bold="true" box="[199,328,439,465]" pageId="20" pageNumber="357">Etymology</emphasis>
. The name of the new species is derived from Latin
<taxonomicName id="4C424D1AFFB2744782993CEFFBD6FE3F" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[954,1032,439,464]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="20" pageNumber="357" phylum="Arthropoda" rank="species" species="incisus">
<emphasis id="B936EA8BFFB2744782993CEFFBD6FE3F" box="[954,1032,439,464]" italics="true" pageId="20" pageNumber="357">incisus</emphasis>
</taxonomicName>
meaning
<emphasis id="B936EA8BFFB27447855F3CEFFB15FE3E" box="[1148,1227,439,465]" italics="true" pageId="20" pageNumber="357">incised</emphasis>
, and refers to the unusual deep incision in the posterior margin of the posterior sternal plate.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,485 @@
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<mods:title id="D875C878093D04D61BC29678274550ED">Seven new species of Spatulaphorus Rack (Acari: Pygmephoridae) phoretic on scarab beetles (Coleoptera: Scarabaeidae)</mods:title>
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<subSubSection id="C3586512FFBE744B81B438B7FD52F9E5" box="[151,652,1519,1546]" pageId="24" pageNumber="361" type="nomenclature">
<paragraph id="8BFD3699FFBE744B81B438B7FD52F9E5" blockId="24.[151,652,1519,1583]" box="[151,652,1519,1546]" pageId="24" pageNumber="361">
<heading id="D0B581F5FFBE744B81B438B7FD52F9E5" bold="true" box="[151,652,1519,1546]" fontSize="11" level="1" pageId="24" pageNumber="361" reason="1">
<emphasis id="B936EA8BFFBE744B81B438B7FD52F9E5" bold="true" box="[151,652,1519,1546]" pageId="24" pageNumber="361">
<taxonomicName id="4C424D1AFFBE744B81B438B7FDF9F9E5" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[151,551,1519,1546]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="24" pageNumber="361" phylum="Arthropoda" rank="species" species="madagascariensis" status="sp. nov.">
<emphasis id="B936EA8BFFBE744B81B438B7FDF9F9E5" bold="true" box="[151,551,1519,1546]" italics="true" pageId="24" pageNumber="361">Spatulaphorus madagascariensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20557F0FFBE744B830C38A8FD52F9E5" box="[559,652,1520,1546]" pageId="24" pageNumber="361" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C3586512FFBE744881B43B4CFE4FF92E" lastPageId="27" lastPageNumber="364" pageId="24" pageNumber="361" type="description">
<paragraph id="8BFD3699FFBE744B81B43B4CFEF9F9C0" blockId="24.[151,652,1519,1583]" box="[151,295,1556,1583]" pageId="24" pageNumber="361">
(
<figureCitation id="13792A1CFFBE744B81BC3B4CFEC1F9C0" box="[159,287,1556,1583]" captionStart-0="FIGURE 25" captionStart-1="FIGURE 26" captionStart-2="FIGURE 27" captionStart-3="FIGURE 28" captionStartId-0="25.[151,250,1592,1616]" captionStartId-1="26.[151,250,1281,1305]" captionStartId-2="27.[151,250,1597,1621]" captionStartId-3="28.[151,250,1196,1220]" captionTargetBox-0="[151,1435,394,1569]" captionTargetBox-1="[151,1435,187,1252]" captionTargetBox-2="[166,1419,221,1542]" captionTargetBox-3="[151,1436,273,1172]" captionTargetId-0="figure-137@25.[151,1435,381,1569]" captionTargetId-1="figure-24@26.[151,1435,181,1257]" captionTargetId-2="figure-17@27.[151,1436,204,1559]" captionTargetId-3="figure-69@28.[151,1436,273,1172]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionTargetPageId-2="27" captionTargetPageId-3="28" captionText-0="FIGURE 25. Spatulaphorus madagascariensis sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." captionText-1="FIGURE 26. Spatulaphorus madagascariensis sp. nov., female: A—right leg I, dorsal aspect; B—right leg II, dorsal aspect." captionText-2="FIGURE 27. Spatulaphorus madagascariensis sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." captionText-3="FIGURE 28. DIC micrographs of Spatulaphorus madagascariensis sp. nov., female: A—general view dorsally; B—general view ventrally." pageId="24" pageNumber="361">Figs 2528</figureCitation>
)
</paragraph>
<paragraph id="8BFD3699FFBE744B81B43B04FA80F998" blockId="24.[151,1437,1628,2014]" box="[151,1374,1628,1655]" pageId="24" pageNumber="361">
<emphasis id="B936EA8BFFBE744B81B43B04FEFFF999" bold="true" box="[151,289,1628,1654]" pageId="24" pageNumber="361">Description</emphasis>
. FEMALE. Body weakly sclerotized. Length of idiosoma 185 (170190), width 110 (105115).
</paragraph>
<paragraph id="8BFD3699FFBE744B81E43BD8FEE4F89D" blockId="24.[151,1437,1628,2014]" pageId="24" pageNumber="361">
<emphasis id="B936EA8BFFBE744B81E43BD8FE88F975" box="[199,342,1664,1690]" italics="true" pageId="24" pageNumber="361">Gnathosoma</emphasis>
. Gnathosomal capsule, excluding palps, subrectangular, length 21 (2021), width 21 (2021). Dorsum with one pair of needle-like cheliceral setae (
<emphasis id="B936EA8BFFBE744B83DB3BFCFCFEF951" box="[760,800,1700,1726]" italics="true" pageId="24" pageNumber="361">cha</emphasis>
) 6 (6). Dorsal median apodeme absent. Postpalpal setae (
<emphasis id="B936EA8BFFBE744B81823B91FF63F90D" box="[161,189,1737,1762]" italics="true" pageId="24" pageNumber="361">pp</emphasis>
) 2 (2) spiniform, situated laterad setae
<emphasis id="B936EA8BFFBE744B83583B90FD7DF90D" box="[635,675,1736,1762]" italics="true" pageId="24" pageNumber="361">cha</emphasis>
. Ventral gnathosoma with one pair of smooth, pointed subcapitular setae
<emphasis id="B936EA8BFFBE744B81FB3BB5FF32F8E9" box="[216,236,1773,1798]" italics="true" pageId="24" pageNumber="361">m</emphasis>
8 (89) and round pits
<emphasis id="B936EA8BFFBE744B80DF3BB5FDD4F8E9" box="[508,522,1773,1798]" italics="true" pageId="24" pageNumber="361">n</emphasis>
situated posteriad
<emphasis id="B936EA8BFFBE744B83C23BB5FD2BF8E9" box="[737,757,1773,1798]" italics="true" pageId="24" pageNumber="361">m</emphasis>
. Palps with smooth and pointed setae
<emphasis id="B936EA8BFFBE744B858E3BB4FB07F8E9" box="[1197,1241,1772,1798]" italics="true" pageId="24" pageNumber="361">dFe</emphasis>
6 (67) and
<emphasis id="B936EA8BFFBE744B844D3BB4FA43F8E9" box="[1390,1437,1772,1798]" italics="true" pageId="24" pageNumber="361">dGe</emphasis>
12 (1113) dorsolaterally. Palps ventrally each with mushroom-like accessory setigenous structure (
<emphasis id="B936EA8BFFBE744B842A3A49FAF3F8C5" box="[1289,1325,1809,1834]" italics="true" pageId="24" pageNumber="361">ass</emphasis>
) and tiny solenidion
<emphasis id="B936EA8BFFBE744B80373A6DFEF6F8A1" box="[276,296,1845,1870]" italics="true" pageId="24" pageNumber="361">ω</emphasis>
. Palps terminated with small blunt-tipped tibial claw and tiny eupathid-like seta. Pharyngeal pumps poorly visible.
</paragraph>
<paragraph id="8BFD3699FFBE744A81E43A24FEE3FE8A" blockId="24.[151,1437,1628,2014]" lastBlockId="25.[151,1436,151,357]" lastPageId="25" lastPageNumber="362" pageId="24" pageNumber="361">
<emphasis id="B936EA8BFFBE744B81E43A24FE4AF879" box="[199,404,1916,1942]" italics="true" pageId="24" pageNumber="361">Idiosomal dorsum</emphasis>
(
<figureCitation id="13792A1CFFBE744B80873A24FDCCF878" box="[420,530,1916,1943]" captionStart="FIGURE 25" captionStartId="25.[151,250,1592,1616]" captionTargetBox="[151,1435,394,1569]" captionTargetId="figure-137@25.[151,1435,381,1569]" captionTargetPageId="25" captionText="FIGURE 25. Spatulaphorus madagascariensis sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." pageId="24" pageNumber="361">Figs 25A</figureCitation>
,
<figureCitation id="13792A1CFFBE744B833C3A24FD8AF879" box="[543,596,1916,1942]" captionStart="FIGURE 28" captionStartId="28.[151,250,1196,1220]" captionTargetBox="[151,1436,273,1172]" captionTargetId="figure-69@28.[151,1436,273,1172]" captionTargetPageId="28" captionText="FIGURE 28. DIC micrographs of Spatulaphorus madagascariensis sp. nov., female: A—general view dorsally; B—general view ventrally." pageId="24" pageNumber="361">28A</figureCitation>
). All dorsal shields with large puncta as illustrated in
<figureCitation id="13792A1CFFBE744B859F3A24FAFAF878" box="[1212,1316,1916,1943]" captionStart="FIGURE 25" captionStartId="25.[151,250,1592,1616]" captionTargetBox="[151,1435,394,1569]" captionTargetId="figure-137@25.[151,1435,381,1569]" captionTargetPageId="25" captionText="FIGURE 25. Spatulaphorus madagascariensis sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." pageId="24" pageNumber="361">Fig. 25A</figureCitation>
. Prodorsal shield with three pairs of setae (
<emphasis id="B936EA8BFFBE744B83233AF8FDC4F855" box="[512,538,1952,1978]" italics="true" pageId="24" pageNumber="361">v1</emphasis>
,
<emphasis id="B936EA8BFFBE744B830B3AF8FD9DF855" box="[552,579,1952,1978]" italics="true" pageId="24" pageNumber="361">v2</emphasis>
,
<emphasis id="B936EA8BFFBE744B83723AF8FDA9F855" box="[593,631,1952,1978]" italics="true" pageId="24" pageNumber="361">sc2</emphasis>
), one pair of weakly barbed capitate trichobothria (
<emphasis id="B936EA8BFFBE744B859F3AF8FB3FF855" box="[1212,1249,1952,1978]" italics="true" pageId="24" pageNumber="361">sc1</emphasis>
) and one pair of round stigmata. Stigmata associated with well-developed tracheal trunks. Setae
<emphasis id="B936EA8BFFBE744B85053A9CFB9EF831" box="[1062,1088,1988,2014]" italics="true" pageId="24" pageNumber="361">v1</emphasis>
,
<emphasis id="B936EA8BFFBE744B85723A9CFBB5F831" box="[1105,1131,1988,2014]" italics="true" pageId="24" pageNumber="361">v2</emphasis>
,
<emphasis id="B936EA8BFFBE744B85583A9CFB7FF831" box="[1147,1185,1988,2014]" italics="true" pageId="24" pageNumber="361">sc2</emphasis>
and
<emphasis id="B936EA8BFFBE744B85F83A9CFB2BF831" box="[1243,1269,1988,2014]" italics="true" pageId="24" pageNumber="361">c2</emphasis>
pointed; other dorsal setae blunt-tipped; setae
<emphasis id="B936EA8BFFBF744A832B3DCFFDCAFF5F" box="[520,532,151,176]" italics="true" pageId="25" pageNumber="362">e</emphasis>
and
<emphasis id="B936EA8BFFBF744A83703DCFFDB1FF5E" box="[595,623,151,177]" italics="true" pageId="25" pageNumber="362">h2</emphasis>
smooth; other dorsal setae barbed. Alveoli of setae
<emphasis id="B936EA8BFFBF744A85F53DCFFB3CFF5F" box="[1238,1250,151,176]" italics="true" pageId="25" pageNumber="362">e</emphasis>
and
<emphasis id="B936EA8BFFBF744A84023DCFFAF7FF5E" box="[1313,1321,151,177]" italics="true" pageId="25" pageNumber="362">f</emphasis>
distinctly separated. Cupules
<emphasis id="B936EA8BFFBF744A80573DE3FE54FF3B" box="[372,394,187,212]" italics="true" pageId="25" pageNumber="362">ia</emphasis>
on tergite D,
<emphasis id="B936EA8BFFBF744A83093DE3FD98FF3B" box="[554,582,187,212]" italics="true" pageId="25" pageNumber="362">im</emphasis>
on tergite EF and
<emphasis id="B936EA8BFFBF744A823F3DE3FCECFF3A" box="[796,818,187,213]" italics="true" pageId="25" pageNumber="362">ih</emphasis>
on tergite H large, round, subequal. Lengths of dorsal setae:
<emphasis id="B936EA8BFFBF744A81FC3D87FF27FF16" box="[223,249,223,249]" italics="true" pageId="25" pageNumber="362">v1</emphasis>
29 (2629),
<emphasis id="B936EA8BFFBF744A80AD3D87FE77FF16" box="[398,425,223,249]" italics="true" pageId="25" pageNumber="362">v2</emphasis>
21 (1822),
<emphasis id="B936EA8BFFBF744A831E3D87FDBDFF16" box="[573,611,223,249]" italics="true" pageId="25" pageNumber="362">sc2</emphasis>
43 (4357),
<emphasis id="B936EA8BFFBF744A83D43D87FCCCFF16" box="[759,786,223,249]" italics="true" pageId="25" pageNumber="362">c1</emphasis>
30 (2734),
<emphasis id="B936EA8BFFBF744A82813D87FC63FF16" box="[930,957,223,249]" italics="true" pageId="25" pageNumber="362">c2</emphasis>
43 (4049),
<emphasis id="B936EA8BFFBF744A85723D87FB81FF16" box="[1105,1119,223,249]" italics="true" pageId="25" pageNumber="362">d</emphasis>
32 (3240),
<emphasis id="B936EA8BFFBF744A85D73D87FADEFF17" box="[1268,1280,223,248]" italics="true" pageId="25" pageNumber="362">e</emphasis>
16 (1516),
<emphasis id="B936EA8BFFBF744A84B73D87FA42FF16" box="[1428,1436,223,249]" italics="true" pageId="25" pageNumber="362">f</emphasis>
36 (3444),
<emphasis id="B936EA8BFFBF744A80023C5BFEE3FEF2" box="[289,317,259,285]" italics="true" pageId="25" pageNumber="362">h1</emphasis>
43 (3944),
<emphasis id="B936EA8BFFBF744A80EC3C5BFE35FEF2" box="[463,491,259,285]" italics="true" pageId="25" pageNumber="362">h2</emphasis>
12 (1214). Distances between setae:
<emphasis id="B936EA8BFFBF744A82B83C5BFC00FEF2" box="[923,990,259,285]" italics="true" pageId="25" pageNumber="362">v1v1</emphasis>
20 (2021),
<emphasis id="B936EA8BFFBF744A854C3C5BFB6CFEF2" box="[1135,1202,259,285]" italics="true" pageId="25" pageNumber="362">v2v2</emphasis>
33 (3336),
<emphasis id="B936EA8BFFBF744A84673C5BFA42FEF2" box="[1348,1436,259,285]" italics="true" pageId="25" pageNumber="362">sc2sc2</emphasis>
38 (3738),
<emphasis id="B936EA8BFFBF744A80073C7FFEB9FEAE" box="[292,359,295,321]" italics="true" pageId="25" pageNumber="362">c1c1</emphasis>
31 (3137),
<emphasis id="B936EA8BFFBF744A80D93C7FFDE3FEAE" box="[506,573,295,321]" italics="true" pageId="25" pageNumber="362">c1c2</emphasis>
24 (2224),
<emphasis id="B936EA8BFFBF744A83F23C7FFD25FEAE" box="[721,763,295,321]" italics="true" pageId="25" pageNumber="362">dd</emphasis>
56 (5357),
<emphasis id="B936EA8BFFBF744A82AC3C7FFC6FFEAE" box="[911,945,295,321]" italics="true" pageId="25" pageNumber="362">ef</emphasis>
5 (56),
<emphasis id="B936EA8BFFBF744A85393C7FFBE6FEAE" box="[1050,1080,295,321]" italics="true" pageId="25" pageNumber="362">ff</emphasis>
51 (4752),
<emphasis id="B936EA8BFFBF744A85E83C7FFACFFEAE" box="[1227,1297,295,321]" italics="true" pageId="25" pageNumber="362">h1h1</emphasis>
41 (3742),
<emphasis id="B936EA8BFFBF744A81B43C13FF03FE8A" box="[151,221,331,357]" italics="true" pageId="25" pageNumber="362">h1h2</emphasis>
6 (68).
</paragraph>
<caption id="DF3D6611FFBF744A81B43B60FA85F9BF" box="[151,1371,1592,1617]" pageId="25" pageNumber="362" startId="25.[151,250,1592,1616]" targetBox="[151,1435,394,1569]" targetPageId="25" targetType="figure">
<paragraph id="8BFD3699FFBF744A81B43B60FA85F9BF" blockId="25.[151,1371,1592,1617]" box="[151,1371,1592,1617]" pageId="25" pageNumber="362">
<emphasis id="B936EA8BFFBF744A81B43B60FEC7F9BF" bold="true" box="[151,281,1592,1616]" pageId="25" pageNumber="362">FIGURE 25</emphasis>
.
<taxonomicName id="4C424D1AFFBF744A80053B60FDA9F9BF" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[294,631,1592,1616]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="25" pageNumber="362" phylum="Arthropoda" rank="species" species="madagascariensis" status="sp. nov.">
<emphasis id="B936EA8BFFBF744A80053B60FDA9F9BF" box="[294,631,1592,1616]" italics="true" pageId="25" pageNumber="362">Spatulaphorus madagascariensis</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFBF744A835E3B60FD10F9BF" bold="true" box="[637,718,1592,1616]" pageId="25" pageNumber="362">
<taxonomicNameLabel id="A20557F0FFBF744A835E3B60FD10F9BF" box="[637,718,1592,1616]" pageId="25" pageNumber="362" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—dorsum of body; B—venter of body. Legs omitted.
</paragraph>
</caption>
<paragraph id="8BFD3699FFBF744A81E43B26FDBBF833" blockId="25.[151,1437,1662,2013]" pageId="25" pageNumber="362">
<emphasis id="B936EA8BFFBF744A81E43B26FE56F977" box="[199,392,1662,1688]" italics="true" pageId="25" pageNumber="362">Idiosomal venter</emphasis>
(
<figureCitation id="13792A1CFFBF744A80B93B26FDD9F976" box="[410,519,1662,1689]" captionStart="FIGURE 25" captionStartId="25.[151,250,1592,1616]" captionTargetBox="[151,1435,394,1569]" captionTargetId="figure-137@25.[151,1435,381,1569]" captionTargetPageId="25" captionText="FIGURE 25. Spatulaphorus madagascariensis sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." pageId="25" pageNumber="362">Figs 25B</figureCitation>
,
<figureCitation id="13792A1CFFBF744A83353B27FD97F976" box="[534,585,1663,1689]" captionStart="FIGURE 28" captionStartId="28.[151,250,1196,1220]" captionTargetBox="[151,1436,273,1172]" captionTargetId="figure-69@28.[151,1436,273,1172]" captionTargetPageId="28" captionText="FIGURE 28. DIC micrographs of Spatulaphorus madagascariensis sp. nov., female: A—general view dorsally; B—general view ventrally." pageId="25" pageNumber="362">28B</figureCitation>
). Anterior and posterior sternal plates with very small, hardly discernible puncta; aggenital and pseudanal plates with slightly larger puncta. Setae
<emphasis id="B936EA8BFFBF744A82E83BFAFC36F953" box="[971,1000,1698,1724]" italics="true" pageId="25" pageNumber="362">1b</emphasis>
,
<emphasis id="B936EA8BFFBF744A82D53BFBFBCEF952" box="[1014,1040,1699,1725]" italics="true" pageId="25" pageNumber="362">2c</emphasis>
,
<emphasis id="B936EA8BFFBF744A853C3BFAFBE5F953" box="[1055,1083,1698,1724]" italics="true" pageId="25" pageNumber="362">4b</emphasis>
, and
<emphasis id="B936EA8BFFBF744A85593BFBFB66F952" box="[1146,1208,1699,1725]" italics="true" pageId="25" pageNumber="362">ps1-2</emphasis>
pointed; other setae weakly blunt-tipped; setae
<emphasis id="B936EA8BFFBF744A80E23B9EFE03F90F" box="[449,477,1734,1760]" italics="true" pageId="25" pageNumber="362">1b</emphasis>
weakly barbed, other ventral setae smooth. Bases of setae
<emphasis id="B936EA8BFFBF744A85433B9FFB59F90E" box="[1120,1159,1735,1761]" italics="true" pageId="25" pageNumber="362">ps3</emphasis>
situated slightly anteriad bases of setae
<emphasis id="B936EA8BFFBF744A80153BB3FEAAF8EA" box="[310,372,1771,1797]" italics="true" pageId="25" pageNumber="362">ps1-2</emphasis>
; setae
<emphasis id="B936EA8BFFBF744A80E23BB3FE39F8EA" box="[449,487,1771,1797]" italics="true" pageId="25" pageNumber="362">ps1</emphasis>
usually shorter than
<emphasis id="B936EA8BFFBF744A83EC3BB3FD28F8EA" box="[719,758,1771,1797]" italics="true" pageId="25" pageNumber="362">ps2</emphasis>
; setae
<emphasis id="B936EA8BFFBF744A82603BB3FC5FF8EA" box="[835,897,1771,1797]" italics="true" pageId="25" pageNumber="362">ps1-2</emphasis>
subequal in some specimens. Ap1 and ap2 well-developed and joined with thick appr; apsej absent; ap3 very thin, hardly discernible; ap4 well developed, exceeding beyond bases of setae
<emphasis id="B936EA8BFFBF744A80B23A6AFE73F8A3" box="[401,429,1842,1868]" italics="true" pageId="25" pageNumber="362">3b</emphasis>
; ap5 well developed and exceeding bases of trochanter IV. Ags small, cup-like; pgs small, oval. Posterior margin of posterior sternal plate weakly concave in middle part. Posterior margin of aggenital plate with short tongue-like elongation. Lengths of ventral setae:
<emphasis id="B936EA8BFFBF744A82143A23FC8AF87A" box="[823,852,1915,1941]" italics="true" pageId="25" pageNumber="362">1a</emphasis>
10 (911),
<emphasis id="B936EA8BFFBF744A82F53A22FC2DF87B" box="[982,1011,1914,1940]" italics="true" pageId="25" pageNumber="362">1b</emphasis>
19 (1719),
<emphasis id="B936EA8BFFBF744A85A73A23FB40F87A" box="[1156,1182,1915,1941]" italics="true" pageId="25" pageNumber="362">1с</emphasis>
12 (912),
<emphasis id="B936EA8BFFBF744A84013A23FAE0F87A" box="[1314,1342,1915,1941]" italics="true" pageId="25" pageNumber="362">2a</emphasis>
11 (11),
<emphasis id="B936EA8BFFBF744A81B43AC7FF6FF856" box="[151,177,1951,1977]" italics="true" pageId="25" pageNumber="362">2с</emphasis>
13 (1315),
<emphasis id="B936EA8BFFBF744A80673AC7FEBEF856" box="[324,352,1951,1977]" italics="true" pageId="25" pageNumber="362">3a</emphasis>
13 (1213),
<emphasis id="B936EA8BFFBF744A80D13AC6FDD0F857" box="[498,526,1950,1976]" italics="true" pageId="25" pageNumber="362">3b</emphasis>
11 (1011),
<emphasis id="B936EA8BFFBF744A83BD3AC7FD66F856" box="[670,696,1951,1977]" italics="true" pageId="25" pageNumber="362">3c</emphasis>
12 (1213),
<emphasis id="B936EA8BFFBF744A82693AC7FCB9F856" box="[842,871,1951,1977]" italics="true" pageId="25" pageNumber="362">4a</emphasis>
10 (1011),
<emphasis id="B936EA8BFFBF744A82D43AC6FBCAF857" box="[1015,1044,1950,1976]" italics="true" pageId="25" pageNumber="362">4b</emphasis>
16 (1520),
<emphasis id="B936EA8BFFBF744A85863AC7FB1EF856" box="[1189,1216,1951,1977]" italics="true" pageId="25" pageNumber="362">4c</emphasis>
14 (1215),
<emphasis id="B936EA8BFFBF744A84713AC7FAA7F856" box="[1362,1401,1951,1977]" italics="true" pageId="25" pageNumber="362">ps1</emphasis>
12 (1112),
<emphasis id="B936EA8BFFBF744A81DE3A9BFEFAF832" box="[253,292,1987,2013]" italics="true" pageId="25" pageNumber="362">ps2</emphasis>
15 (1315),
<emphasis id="B936EA8BFFBF744A80973A9BFE05F832" box="[436,475,1987,2013]" italics="true" pageId="25" pageNumber="362">ps3</emphasis>
16 (1617).
</paragraph>
<caption id="DF3D6611FFBC744981B43859FA59FAF6" box="[151,1415,1281,1305]" pageId="26" pageNumber="363" startId="26.[151,250,1281,1305]" targetBox="[151,1435,187,1252]" targetPageId="26" targetType="figure">
<paragraph id="8BFD3699FFBC744981B43859FA59FAF6" blockId="26.[151,1415,1281,1305]" box="[151,1415,1281,1305]" pageId="26" pageNumber="363">
<emphasis id="B936EA8BFFBC744981B43859FEC7FAF6" bold="true" box="[151,281,1281,1305]" pageId="26" pageNumber="363">FIGURE 26</emphasis>
.
<taxonomicName id="4C424D1AFFBC744980053859FDA9FAF6" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[294,631,1281,1305]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="26" pageNumber="363" phylum="Arthropoda" rank="species" species="madagascariensis" status="sp. nov.">
<emphasis id="B936EA8BFFBC744980053859FDA9FAF6" box="[294,631,1281,1305]" italics="true" pageId="26" pageNumber="363">Spatulaphorus madagascariensis</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFBC7449835E3859FD10FAF6" bold="true" box="[637,718,1281,1305]" pageId="26" pageNumber="363">
<taxonomicNameLabel id="A20557F0FFBC7449835E3859FD10FAF6" box="[637,718,1281,1305]" pageId="26" pageNumber="363" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—right leg I, dorsal aspect; B—right leg II, dorsal aspect.
</paragraph>
</caption>
<paragraph id="8BFD3699FFBC744981E43810FE05F84E" blockId="26.[151,1437,1351,1953]" pageId="26" pageNumber="363">
<emphasis id="B936EA8BFFBC744981E43810FF22FA8E" box="[199,252,1352,1377]" italics="true" pageId="26" pageNumber="363">Legs</emphasis>
(
<figureCitation id="13792A1CFFBC74498028381FFEBCFA8E" box="[267,354,1351,1377]" captionStart="FIGURE 26" captionStartId="26.[151,250,1281,1305]" captionTargetBox="[151,1435,187,1252]" captionTargetId="figure-24@26.[151,1435,181,1257]" captionTargetPageId="26" captionText="FIGURE 26. Spatulaphorus madagascariensis sp. nov., female: A—right leg I, dorsal aspect; B—right leg II, dorsal aspect." pageId="26" pageNumber="363">Figs 26</figureCitation>
,
<figureCitation id="13792A1CFFBC7449804D3810FE50FA8D" box="[366,398,1352,1378]" captionStart="FIGURE 27" captionStartId="27.[151,250,1597,1621]" captionTargetBox="[166,1419,221,1542]" captionTargetId="figure-17@27.[151,1436,204,1559]" captionTargetPageId="27" captionText="FIGURE 27. Spatulaphorus madagascariensis sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." pageId="26" pageNumber="363">27</figureCitation>
). Setation of legs as in
<taxonomicName id="4C424D1AFFBC744983B23810FCE8FA8E" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[657,822,1351,1378]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="26" pageNumber="363" phylum="Arthropoda" rank="species" species="brevisetosus">
<emphasis id="B936EA8BFFBC744983B23810FCE8FA8E" box="[657,822,1351,1378]" italics="true" pageId="26" pageNumber="363">S. brevisetosus</emphasis>
</taxonomicName>
except presence of solenidion
<emphasis id="B936EA8BFFBC744985B23810FB7EFA8E" box="[1169,1184,1352,1377]" italics="true" pageId="26" pageNumber="363">φ</emphasis>
on tibia II. Leg I (
<figureCitation id="13792A1CFFBC7449844C381FFF12FA6A" captionStart="FIGURE 26" captionStartId="26.[151,250,1281,1305]" captionTargetBox="[151,1435,187,1252]" captionTargetId="figure-24@26.[151,1435,181,1257]" captionTargetPageId="26" captionText="FIGURE 26. Spatulaphorus madagascariensis sp. nov., female: A—right leg I, dorsal aspect; B—right leg II, dorsal aspect." pageId="26" pageNumber="363">Fig. 26A</figureCitation>
). Tibiotarsus enlarged, with large, thick and strongly curved claw; tibiotarsus internally without oval strongly sclerotized structure. Setae (
<emphasis id="B936EA8BFFBC744980F038C8FE3FFA46" box="[467,481,1424,1449]" italics="true" pageId="26" pageNumber="363">u</emphasis>
) consolidated into a bifid horn-like structure opposing claw. Tarsal claw blunt-tipped. Seta
<emphasis id="B936EA8BFFBC744981ED38EBFF02FA22" box="[206,220,1459,1485]" italics="true" pageId="26" pageNumber="363">d</emphasis>
of femur spatulate, with short subterminal projection. Setae
<emphasis id="B936EA8BFFBC7449825C38EBFC4EFA22" box="[895,912,1459,1485]" italics="true" pageId="26" pageNumber="363">l</emphasis>
of femur and
<emphasis id="B936EA8BFFBC7449850D38EBFBE4FA22" box="[1070,1082,1459,1485]" italics="true" pageId="26" pageNumber="363">k</emphasis>
of tibiotarsus smooth, other leg setae (except eupathidia
<emphasis id="B936EA8BFFBC7449808E3880FE1AFA1D" box="[429,452,1496,1522]" italics="true" pageId="26" pageNumber="363">p</emphasis>
,
<emphasis id="B936EA8BFFBC744980F03880FE2FFA1D" box="[467,497,1496,1522]" italics="true" pageId="26" pageNumber="363">p”</emphasis>
,
<emphasis id="B936EA8BFFBC744980DC3880FDC3FA1D" box="[511,541,1496,1522]" italics="true" pageId="26" pageNumber="363">tc</emphasis>
,
<emphasis id="B936EA8BFFBC744983083880FD91FA1D" box="[555,591,1496,1522]" italics="true" pageId="26" pageNumber="363">tc”</emphasis>
,
<emphasis id="B936EA8BFFBC7449837D388FFDA9FA1E" box="[606,631,1495,1521]" italics="true" pageId="26" pageNumber="363">ft</emphasis>
,
<emphasis id="B936EA8BFFBC744983A6388FFD7AFA1E" box="[645,676,1495,1521]" italics="true" pageId="26" pageNumber="363">ft”</emphasis>
) barbed. Setae
<emphasis id="B936EA8BFFBC74498276388FFCBDFA1E" box="[853,867,1495,1521]" italics="true" pageId="26" pageNumber="363">l</emphasis>
of femur and
<emphasis id="B936EA8BFFBC74498527388FFBCEFA1E" box="[1028,1040,1495,1521]" italics="true" pageId="26" pageNumber="363">k</emphasis>
of tibiotarsus weakly blunt-tipped; other leg setae (except eupathidia) pointed. Lengths of solenidia
<emphasis id="B936EA8BFFBC7449824D38A4FC4EF9F9" box="[878,912,1532,1558]" italics="true" pageId="26" pageNumber="363">ω1</emphasis>
8 (8),
<emphasis id="B936EA8BFFBC744982F838A4FC23F9F9" box="[987,1021,1532,1558]" italics="true" pageId="26" pageNumber="363">ω2</emphasis>
5 (5),
<emphasis id="B936EA8BFFBC7449856A38A4FBB8F9F9" box="[1097,1126,1532,1558]" italics="true" pageId="26" pageNumber="363">φ1</emphasis>
8 (8),
<emphasis id="B936EA8BFFBC7449859238A4FB11F9F9" box="[1201,1231,1532,1558]" italics="true" pageId="26" pageNumber="363">φ2</emphasis>
6 (56); solenidia
<emphasis id="B936EA8BFFBC744981B43B78FF67F9D5" box="[151,185,1568,1594]" italics="true" pageId="26" pageNumber="363">ω1</emphasis>
and
<emphasis id="B936EA8BFFBC744981D33B78FED3F9D5" box="[240,269,1568,1594]" italics="true" pageId="26" pageNumber="363">φ1</emphasis>
thick, clavate; other solenidia weakly clavate. Leg II (
<figureCitation id="13792A1CFFBC744982503B47FC04F9D6" box="[883,986,1567,1593]" captionStart="FIGURE 26" captionStartId="26.[151,250,1281,1305]" captionTargetBox="[151,1435,187,1252]" captionTargetId="figure-24@26.[151,1435,181,1257]" captionTargetPageId="26" captionText="FIGURE 26. Spatulaphorus madagascariensis sp. nov., female: A—right leg I, dorsal aspect; B—right leg II, dorsal aspect." pageId="26" pageNumber="363">Fig. 26B</figureCitation>
). Tarsus with simple claws and pad-like empodium. Solenidion
<emphasis id="B936EA8BFFBC744980BC3B1CFE6DF9B2" box="[415,435,1604,1629]" italics="true" pageId="26" pageNumber="363">ω</emphasis>
7 (7) weakly clavate, solenidion
<emphasis id="B936EA8BFFBC7449820F3B1CFCE5F9B2" box="[812,827,1604,1629]" italics="true" pageId="26" pageNumber="363">φ</emphasis>
2 (2) very short. Seta
<emphasis id="B936EA8BFFBC744985193B1CFB8FF9B1" box="[1082,1105,1604,1630]" italics="true" pageId="26" pageNumber="363">u</emphasis>
of tarsus smooth; other setae leg setae barbed; seta
<emphasis id="B936EA8BFFBC744980B03B3FFE7FF96E" box="[403,417,1639,1665]" italics="true" pageId="26" pageNumber="363">d</emphasis>
of femur blunt-tipped; tips of setae (
<emphasis id="B936EA8BFFBC7449826A3B30FCBDF96E" box="[841,867,1640,1665]" italics="true" pageId="26" pageNumber="363">pv</emphasis>
) of tarsus slightly foliate; other leg setae pointed. Leg III (
<figureCitation id="13792A1CFFBC744981DA3BD3FEBAF949" box="[249,356,1675,1702]" captionStart="FIGURE 27" captionStartId="27.[151,250,1597,1621]" captionTargetBox="[166,1419,221,1542]" captionTargetId="figure-17@27.[151,1436,204,1559]" captionTargetPageId="27" captionText="FIGURE 27. Spatulaphorus madagascariensis sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." pageId="26" pageNumber="363">Fig. 27A</figureCitation>
). Femur divided into basi- and telofemur. Claws and empodium of same shape as on tarsus II. Seta
<emphasis id="B936EA8BFFBC744981EC3BE8FF38F925" box="[207,230,1712,1738]" italics="true" pageId="26" pageNumber="363">u</emphasis>
of tarsus smooth; other leg setae barbed; setae
<emphasis id="B936EA8BFFBC744982223BF7FCD1F926" box="[769,783,1711,1737]" italics="true" pageId="26" pageNumber="363">d</emphasis>
of femur and
<emphasis id="B936EA8BFFBC744982923BF7FC1CF926" box="[945,962,1711,1737]" italics="true" pageId="26" pageNumber="363">l</emphasis>
of genu blunt-tipped; tips of setae (
<emphasis id="B936EA8BFFBC7449847A3BE8FAADF926" box="[1369,1395,1712,1737]" italics="true" pageId="26" pageNumber="363">pv</emphasis>
) of tarsus slightly flattened; other leg setae pointed; external part of solenidion
<emphasis id="B936EA8BFFBC744982D43B8CFBD8F902" box="[1015,1030,1748,1773]" italics="true" pageId="26" pageNumber="363">φ</emphasis>
absent, but subcuticular very small cup-shaped structure situated at typical insertion point of solenidion. Leg IV (
<figureCitation id="13792A1CFFBC7449852E3BAFFBB0F8FD" box="[1037,1134,1783,1810]" captionStart="FIGURE 27" captionStartId="27.[151,250,1597,1621]" captionTargetBox="[166,1419,221,1542]" captionTargetId="figure-17@27.[151,1436,204,1559]" captionTargetPageId="27" captionText="FIGURE 27. Spatulaphorus madagascariensis sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." pageId="26" pageNumber="363">Fig 27B</figureCitation>
). Femur divided into basi- and telofemur. Claws and empodium as on tarsi II and III. Seta
<emphasis id="B936EA8BFFBC744982793A43FC5EF8DA" box="[858,896,1819,1845]" italics="true" pageId="26" pageNumber="363">pl”</emphasis>
of tarsus smooth; other leg setae barbed; setae
<emphasis id="B936EA8BFFBC744984AD3A43FA42F8DA" box="[1422,1436,1819,1845]" italics="true" pageId="26" pageNumber="363">d</emphasis>
of femur,
<emphasis id="B936EA8BFFBC744980253A18FEFCF8B5" box="[262,290,1856,1882]" italics="true" pageId="26" pageNumber="363">v”</emphasis>
of tibia,
<emphasis id="B936EA8BFFBC744980AA3A67FE71F8B6" box="[393,431,1855,1881]" italics="true" pageId="26" pageNumber="363">pl”</emphasis>
and
<emphasis id="B936EA8BFFBC744980C53A18FDD4F8B5" box="[486,522,1856,1882]" italics="true" pageId="26" pageNumber="363">tc”</emphasis>
of tarsus blunt-tipped; setae (
<emphasis id="B936EA8BFFBC744982793A18FCAAF8B6" box="[858,884,1856,1881]" italics="true" pageId="26" pageNumber="363">pv</emphasis>
) of tarsus slightly foliate distally; other leg setae pointed; external part of solenidion
<emphasis id="B936EA8BFFBC744983123A3CFD9EF892" box="[561,576,1892,1917]" italics="true" pageId="26" pageNumber="363">φ</emphasis>
absent, but subcuticular very small cup-shaped structure situated at typical insertion point of solenidion.
</paragraph>
<caption id="DF3D6611FFBD744881B43B65FF00F996" pageId="27" pageNumber="364" startId="27.[151,250,1597,1621]" targetBox="[166,1419,221,1542]" targetPageId="27" targetType="figure">
<paragraph id="8BFD3699FFBD744881B43B65FF00F996" blockId="27.[151,1436,1597,1657]" pageId="27" pageNumber="364">
<emphasis id="B936EA8BFFBD744881B43B65FEC0F9BA" bold="true" box="[151,286,1597,1621]" pageId="27" pageNumber="364">FIGURE 27</emphasis>
.
<taxonomicName id="4C424D1AFFBD744880133B65FD58F9BA" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[304,646,1597,1621]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="27" pageNumber="364" phylum="Arthropoda" rank="species" species="madagascariensis" status="sp. nov.">
<emphasis id="B936EA8BFFBD744880133B65FD58F9BA" box="[304,646,1597,1621]" italics="true" pageId="27" pageNumber="364">Spatulaphorus madagascariensis</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFBD744883B23B65FD39F9BA" bold="true" box="[657,743,1597,1621]" pageId="27" pageNumber="364">
<taxonomicNameLabel id="A20557F0FFBD744883B23B65FD39F9BA" box="[657,743,1597,1621]" pageId="27" pageNumber="364" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect.
</paragraph>
</caption>
<paragraph id="8BFD3699FFBD744881EB3BF0FE4FF92E" blockId="27.[151,1437,1703,2018]" box="[200,401,1703,1730]" pageId="27" pageNumber="364">MALE unknown.</paragraph>
</subSubSection>
<subSubSection id="C3586512FFBD744881E43B94FD9BF89A" pageId="27" pageNumber="364" type="materials_examined">
<paragraph id="8BFD3699FFBD744881E43B94FB17F8E6" blockId="27.[151,1437,1703,2018]" pageId="27" pageNumber="364">
<emphasis id="B936EA8BFFBD744881E43B94FEACF90A" bold="true" box="[199,370,1739,1766]" pageId="27" pageNumber="364">Type material</emphasis>
. Female
<materialsCitation id="3B2A3CC4FFBD744880C23B93FC9BF8E6" collectionCode="ZISP, ZIN" country="Madagascar" location="Adringitra Range" municipality="Andohariana" pageId="27" pageNumber="364" specimenCount="1" typeStatus="holotype">
<typeStatus id="54F9883BFFBD744880C23B93FD98F90A" box="[481,582,1739,1765]" pageId="27" pageNumber="364" type="holotype">holotype</typeStatus>
, slide
<collectionCode id="ED53AE5CFFBD744883B73B94FD13F909" box="[660,717,1740,1766]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="27" pageNumber="364">ZISP</collectionCode>
T-Pygm-016,
<collectingCountry id="F3557609FFBD744882563B93FC20F90A" box="[885,1022,1739,1765]" name="Madagascar" pageId="27" pageNumber="364">Madagascar</collectingCountry>
,
<location id="8E9D6042FFBD744885283B93FB0BF909" LSID="urn:lsid:plazi:treatment:03EB878FFFBE744F81B438B7FB10FF16:8E9D6042FFBD744885283B93FB0BF909" box="[1035,1237,1739,1766]" country="Madagascar" municipality="Andohariana" name="Adringitra Range" pageId="27" pageNumber="364">Adringitra Range</location>
,
<collectingMunicipality id="6B99ACE3FFBD744885C03B93FAA9F90A" box="[1251,1399,1739,1765]" pageId="27" pageNumber="364">Andohariana</collectingMunicipality>
, 9- XI/
<date id="FFFC1059FFBD744881E33BA8FE96F8E5" box="[192,328,1776,1802]" pageId="27" pageNumber="364" value="1979-12-10">10-XII.1979</date>
(
<collectionCode id="ED53AE5CFFBD744880743BA8FE55F8E5" box="[343,395,1776,1802]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="27" pageNumber="364" type="Museum">ZIN</collectionCode>
),
<collectedFrom id="385E7564FFBD744880BD3BA8FC9BF8E6" box="[414,837,1775,1802]" pageId="27" pageNumber="364">
on
<taxonomicName id="4C424D1AFFBD744880E23BB7FC9BF8E6" authority="Orbigny" authorityName="Orbigny" authorityYear="1915" box="[449,837,1775,1801]" class="Insecta" family="Scarabaeidae" genus="Helictopleurus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="364" phylum="Arthropoda" rank="species" species="corruscus">
<emphasis id="B936EA8BFFBD744880E23BB7FD02F8E6" box="[449,732,1775,1801]" italics="true" pageId="27" pageNumber="364">Helictopleurus corruscus</emphasis>
Orbigny
</taxonomicName>
</collectedFrom>
</materialsCitation>
;
<materialsCitation id="3B2A3CC4FFBD744882733BA8FB1BF8E6" box="[848,1221,1775,1802]" collectionCode="Male" country="Madagascar" location="Adringitra Range" municipality="Andohariana" pageId="27" pageNumber="364" specimenCount="75" specimenCount-female="75" typeStatus="paratype">
<typeStatus id="54F9883BFFBD744882733BA8FC1EF8E5" box="[848,960,1776,1802]" pageId="27" pageNumber="364" type="paratype">paratypes</typeStatus>
:
<specimenCount id="9D44FD10FFBD744882E93BB7FB99F8E6" box="[970,1095,1775,1802]" count="75" pageId="27" pageNumber="364" type="female">75 females</specimenCount>
, same data
</materialsCitation>
.
</paragraph>
<paragraph id="8BFD3699FFBD744881E43A4CFD9BF89A" blockId="27.[151,1437,1703,2018]" pageId="27" pageNumber="364">
<emphasis id="B936EA8BFFBD744881E43A4CFE58F8C2" bold="true" box="[199,390,1811,1838]" pageId="27" pageNumber="364">Type deposition</emphasis>
.
<materialsCitation id="3B2A3CC4FFBD744880B53A4BFE25F8BE" country="Russia" location="Zoological Institute" pageId="27" pageNumber="364" specimenCount="9" stateProvince="Saint Petersburg" typeStatus="holotype">
The
<typeStatus id="54F9883BFFBD744880EF3A4BFDF0F8C2" box="[460,558,1811,1837]" pageId="27" pageNumber="364" type="holotype">holotype</typeStatus>
and
<specimenCount id="9D44FD10FFBD7448834A3A4BFCD0F8C1" box="[617,782,1811,1838]" count="9" pageId="27" pageNumber="364" type="generic" typeStatus="paratype">nine paratypes</specimenCount>
are deposited in the collection of
<location id="8E9D6042FFBD744885BE3A4BFAA5F8C2" LSID="urn:lsid:plazi:treatment:03EB878FFFBE744F81B438B7FB10FF16:8E9D6042FFBD744885BE3A4BFAA5F8C2" box="[1181,1403,1811,1837]" country="Russia" name="Zoological Institute" pageId="27" pageNumber="364" stateProvince="Saint Petersburg">Zoological Institute</location>
of RAS,
<collectingRegion id="4986F87BFFBD744881FC3A6FFE40F8BE" box="[223,414,1847,1873]" country="Russia" name="Saint Petersburg" pageId="27" pageNumber="364">Saint Petersburg</collectingRegion>
,
<collectingCountry id="F3557609FFBD7448808E3A6FFE25F8BE" box="[429,507,1847,1873]" name="Russia" pageId="27" pageNumber="364">Russia</collectingCountry>
</materialsCitation>
;
<materialsCitation id="3B2A3CC4FFBD744883293A6FFAC5F8BD" box="[522,1307,1847,1874]" country="Russia" location="Tyumen State" pageId="27" pageNumber="364" specimenCount="1" stateProvince="Tyumen'" typeStatus="paratype">
other
<typeStatus id="54F9883BFFBD7448836D3A60FD67F8BD" box="[590,697,1848,1874]" pageId="27" pageNumber="364" type="paratype">paratypes</typeStatus>
are deposited in the collection of the
<collectingRegion id="4986F87BFFBD7448855E3A60FAC5F8BD" box="[1149,1307,1848,1874]" country="Russia" name="Tyumen'" pageId="27" pageNumber="364">Tyumen State</collectingRegion>
</materialsCitation>
<materialsCitation id="3B2A3CC4FFBD744884053A6FFD9EF89A" country="Russia" location="University Museum of Zoology" pageId="27" pageNumber="364" specimenCount="1" stateProvince="Tyumen'" typeStatus="paratype">
<location id="8E9D6042FFBD744884053A6FFEA1F89A" LSID="urn:lsid:plazi:treatment:03EB878FFFBE744F81B438B7FB10FF16:8E9D6042FFBD744884053A6FFEA1F89A" country="Russia" name="University Museum of Zoology" pageId="27" pageNumber="364" stateProvince="Tyumen'">University Museum of Zoology</location>
,
<collectingRegion id="4986F87BFFBD744880AA3A04FE39F899" box="[393,487,1884,1910]" country="Russia" name="Tyumen'" pageId="27" pageNumber="364">Tyumen</collectingRegion>
,
<collectingCountry id="F3557609FFBD744880D13A03FD9EF89A" box="[498,576,1883,1909]" name="Russia" pageId="27" pageNumber="364">Russia</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3586512FFBD744F81E43A27FC77FF3A" lastPageId="28" lastPageNumber="365" pageId="27" pageNumber="364" type="diagnosis">
<paragraph id="8BFD3699FFBD744F81E43A27FC77FF3A" blockId="27.[151,1437,1703,2018]" lastBlockId="28.[151,1436,151,249]" lastPageId="28" lastPageNumber="365" pageId="27" pageNumber="364">
<emphasis id="B936EA8BFFBD744881E43A27FE18F876" bold="true" box="[199,454,1919,1945]" pageId="27" pageNumber="364">Differential diagnosis</emphasis>
. The new species is most similar to
<taxonomicName id="4C424D1AFFBD7448827D3AD8FC16F876" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[862,968,1920,1946]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="27" pageNumber="364" phylum="Arthropoda" rank="species" species="incisus">
<emphasis id="B936EA8BFFBD7448827D3AD8FC16F876" box="[862,968,1920,1946]" italics="true" pageId="27" pageNumber="364">S. incisus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFBD744882EC3AD8FBF6F875" bold="true" box="[975,1064,1920,1946]" pageId="27" pageNumber="364">sp, nov.</emphasis>
in having an enlarged tibiotarsus I, simple claws on tarsi II and III and presence of solenidion
<emphasis id="B936EA8BFFBD744882183AFCFC94F852" box="[827,842,1956,1981]" italics="true" pageId="27" pageNumber="364">φ</emphasis>
on tibia II. The new species differs from
<taxonomicName id="4C424D1AFFBD744884353AFCFA5EF852" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[1302,1408,1956,1982]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="27" pageNumber="364" phylum="Arthropoda" rank="species" species="incisus">
<emphasis id="B936EA8BFFBD744884353AFCFA5EF852" box="[1302,1408,1956,1982]" italics="true" pageId="27" pageNumber="364">S. incisus</emphasis>
</taxonomicName>
in having well developed ap5 exceeding trochanters IV (vs. ap5 very short and thin in
<taxonomicName id="4C424D1AFFBD744885613A90FB72F80E" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[1090,1196,1992,2018]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="27" pageNumber="364" phylum="Arthropoda" rank="species" species="incisus">
<emphasis id="B936EA8BFFBD744885613A90FB72F80E" box="[1090,1196,1992,2018]" italics="true" pageId="27" pageNumber="364">S. incisus</emphasis>
</taxonomicName>
); posterior margin of posterior sternal plate almost straight medially (vs. posterior margin of posterior sternal plate deeply incised in
<taxonomicName id="4C424D1AFFBA744F84A43DCFFF3BFF3B" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="28" pageNumber="365" phylum="Arthropoda" rank="species" species="incisus">
<emphasis id="B936EA8BFFBA744F84A43DCFFF3BFF3B" italics="true" pageId="28" pageNumber="365">S. incisus</emphasis>
</taxonomicName>
); and setae
<emphasis id="B936EA8BFFBA744F80483DE3FE77FF3A" box="[363,425,187,213]" italics="true" pageId="28" pageNumber="365">ps1-3</emphasis>
smooth (vs. setae
<emphasis id="B936EA8BFFBA744F835A3DE3FD69FF3A" box="[633,695,187,213]" italics="true" pageId="28" pageNumber="365">ps1-3</emphasis>
barbed in
<taxonomicName id="4C424D1AFFBA744F820D3DE3FC46FF3B" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[814,920,187,213]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="28" pageNumber="365" phylum="Arthropoda" rank="species" species="incisus">
<emphasis id="B936EA8BFFBA744F820D3DE3FC46FF3B" box="[814,920,187,213]" italics="true" pageId="28" pageNumber="365">S. incisus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3586512FFBA744F81E43D87FB10FF16" box="[199,1230,223,249]" pageId="28" pageNumber="365" type="etymology">
<paragraph id="8BFD3699FFBA744F81E43D87FB10FF16" blockId="28.[151,1436,151,249]" box="[199,1230,223,249]" pageId="28" pageNumber="365">
<emphasis id="B936EA8BFFBA744F81E43D87FE96FF16" bold="true" box="[199,328,223,249]" pageId="28" pageNumber="365">Etymology</emphasis>
. The name of the new species refers to geographical distribution in
<collectingCountry id="F3557609FFBA744F85623D87FB14FF16" box="[1089,1226,223,249]" name="Madagascar" pageId="28" pageNumber="365">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,295 @@
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<mods:title id="DE7B474BA02A995ECD301199F928E800">Vermiophis cangshanensis sp. nov., a new wormlion fly from Yunnan, China, with notes on its immature stages and biology (Diptera: Vermileonidae)</mods:title>
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<mods:namePart id="57954D1793054A41DAA9ECE6A1FBE2AC">Li, Jia-Ling</mods:namePart>
<mods:affiliation id="8AA5C9B4878B2C45A6143A31E0749628">College of Agriculture and Biological Science, Dali University, Dali, Yunnan 671003, P. R. China &amp; Cangshan Forest Ecosystem Observation and Research Station of Yunnan Province, Dali University, Dali, Yunnan 671003, P. R. China</mods:affiliation>
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<mods:namePart id="DF4616D02134DF6F5863DACB7BA9842B">Zhao, Can-Jun</mods:namePart>
<mods:affiliation id="9D9872651E3D3D277BA8134AF8D61AA8">Biodiversity Conservation and Breeding Center (Dali) of Northwestern Yunnan, Bureau of the Cangshan-Erhai National Nature Reserve, Dali, Yunnan 671000, P. R. China</mods:affiliation>
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<mods:location id="A37AF952E28EFD97CB3CE0629CE8ACA2">
<mods:url id="9CF8ACBA972605B7856C75D315DE5A6E">http://dx.doi.org/10.11646/zootaxa.5497.3.5</mods:url>
</mods:location>
<mods:classification id="4F8C67836D3FEB456C33D4F92F80BEFC">journal article</mods:classification>
<mods:identifier id="E8134D5ACA91C7EBC169464693A911F5" type="DOI">10.11646/zootaxa.5497.3.5</mods:identifier>
<mods:identifier id="E8AE52DCB45E24D7754DE434841B9910" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="BE7FCA27105052B8601C5BCE0FF3026F" type="Zenodo-Dep">13618558</mods:identifier>
<mods:identifier id="484E8DB742039A9BCDCFB3666CE5D372" type="ZooBank">833269F0-7A0A-48DE-B76C-60A87FA8F529</mods:identifier>
</mods:mods>
<treatment id="03FABF64FFBE2270FF60EEFCFF21FF68" LSID="urn:lsid:plazi:treatment:03FABF64FFBE2270FF60EEFCFF21FF68" httpUri="http://treatment.plazi.org/id/03FABF64FFBE2270FF60EEFCFF21FF68" lastPageId="4" lastPageNumber="404" pageId="1" pageNumber="401">
<subSubSection id="C3495DF9FFBE2275FF60EEFCFCE9FC50" box="[151,808,978,1005]" pageId="1" pageNumber="401" type="nomenclature">
<paragraph id="8BEC0E72FFBE2275FF60EEFCFCE9FC50" blockId="1.[151,808,978,1041]" box="[151,808,978,1005]" pageId="1" pageNumber="401">
<heading id="D0A4B91EFFBE2275FF60EEFCFCE9FC50" bold="true" box="[151,808,978,1005]" fontSize="11" level="1" pageId="1" pageNumber="401" reason="1">
<emphasis id="B927D260FFBE2275FF60EEFCFCE9FC50" bold="true" box="[151,808,978,1005]" pageId="1" pageNumber="401">
<taxonomicName id="4C5375F1FFBE2275FF60EEFCFD7FFC50" authority="Li, Zhao &amp; Wang" authorityName="Li, Zhao &amp; Wang" authorityYear="2024" box="[151,702,978,1005]" class="Insecta" family="Vermileonidae" genus="Vermiophis" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="401" phylum="Arthropoda" rank="species" species="cangshanensis" status="sp. nov.">
<emphasis id="B927D260FFBE2275FF60EEFCFE18FC50" bold="true" box="[151,473,978,1005]" italics="true" pageId="1" pageNumber="401">Vermiophis cangshanensis</emphasis>
Li, Zhao &amp; Wang
</taxonomicName>
,
<taxonomicNameLabel id="A2146F1BFFBE2275FD3CEEFDFCE9FC50" box="[715,808,979,1005]" pageId="1" pageNumber="401" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3495DF9FFBE2275FF60EED9FECAFBAC" box="[151,267,1014,1041]" pageId="1" pageNumber="401" type="description">
<paragraph id="8BEC0E72FFBE2275FF60EED9FECAFBAC" blockId="1.[151,808,978,1041]" box="[151,267,1014,1041]" pageId="1" pageNumber="401">
(
<figureCitation id="136812F7FFBE2275FF68EED9FEC3FBAC" box="[159,258,1014,1041]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="1.[151,250,1845,1869]" captionStartId-1="2.[151,250,1798,1822]" captionStartId-2="3.[151,250,1920,1944]" captionStartId-3="4.[151,250,1755,1779]" captionStartId-4="5.[151,250,1342,1366]" captionTargetBox-0="[226,1362,1075,1815]" captionTargetBox-1="[194,1436,668,1758]" captionTargetBox-2="[209,1379,682,1884]" captionTargetBox-3="[186,1401,686,1719]" captionTargetBox-4="[151,1436,345,1319]" captionTargetId-0="figure-270@1.[213,1375,1062,1818]" captionTargetId-1="figure-213@2.[151,1436,633,1775]" captionTargetId-2="figure-253@3.[196,1392,669,1897]" captionTargetId-3="figure-181@4.[173,1413,669,1731]" captionTargetId-4="figure-106@5.[151,1436,345,1319]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionTargetPageId-4="5" captionText-0="FIGURE 1. Habitus of Vermiophis cangshanensis sp. nov. Paratypes.A. Male, dorsal view; B. Female, dorsal view." captionText-1="FIGURE 2. Morphological characters of Vermiophis cangshanensis sp. nov., male. Paratypes. A. Habitus, dorsal view; B. Right antenna, dorsal view; CE. Head, left-lateral, dorsal, and ventral views, respectively; F. Right wing, dorsal view." captionText-2="FIGURE 3. Male genitalia of Vermiophis cangshanensis sp. nov. Paratype.A. T9 (epandrium),dorsal view; BE.Hypopygium, ventral, caudal, dorsal, and left-lateral views, respectively.Abbreviations: ae, aedeagus; aap, aedeagal apodeme; ce, cercus; dbr, dorsal bridge; gap, gonocoxital apodeme; gcx, gonocoxite; gs, gonostylus; vp, ventral plane." captionText-3="FIGURE 4. Immature stages of Vermiophis cangshanensis sp. nov. A. Egg; BD. Last instar larva, ventral, left-lateral, and dorsal views, respectively; E.Terminal abdomen of last instar larva showing the posterior spiracles, dorso-posterior view; F.Pupa, right-lateral view. Abbreviations: com, comb-shaped spines; lcp, locomotory papillae; prl, proleg; psp, posterior spiracles." captionText-4="FIGURE 5. Habitat of Vermiophis cangshanensis sp. nov. A. Cangshan Mountain; B, C. Larval colony; D. A larva showing its typical Ω-shaped defensive posture." figureDoi-0="http://doi.org/10.5281/zenodo.13618560" figureDoi-1="http://doi.org/10.5281/zenodo.13618562" figureDoi-2="http://doi.org/10.5281/zenodo.13618564" figureDoi-3="http://doi.org/10.5281/zenodo.13618566" figureDoi-4="http://doi.org/10.5281/zenodo.13618568" httpUri-0="https://zenodo.org/record/13618560/files/figure.png" httpUri-1="https://zenodo.org/record/13618562/files/figure.png" httpUri-2="https://zenodo.org/record/13618564/files/figure.png" httpUri-3="https://zenodo.org/record/13618566/files/figure.png" httpUri-4="https://zenodo.org/record/13618568/files/figure.png" pageId="1" pageNumber="401">Figs 15</figureCitation>
)
</paragraph>
</subSubSection>
<caption id="DF2C5EFAFFBE2275FF60EA1BFA84F8F0" ID-DOI="http://doi.org/10.5281/zenodo.13618560" ID-Zenodo-Dep="13618560" box="[151,1349,1844,1869]" httpUri="https://zenodo.org/record/13618560/files/figure.png" pageId="1" pageNumber="401" startId="1.[151,250,1845,1869]" targetBox="[226,1362,1075,1815]" targetPageId="1" targetType="figure">
<paragraph id="8BEC0E72FFBE2275FF60EA1BFA84F8F0" blockId="1.[151,1349,1844,1869]" box="[151,1349,1844,1869]" pageId="1" pageNumber="401">
<emphasis id="B927D260FFBE2275FF60EA1BFD3FF8F0" bold="true" box="[151,766,1844,1869]" pageId="1" pageNumber="401">
FIGURE 1. Habitus of
<taxonomicName id="4C5375F1FFBE2275FE66EA1AFD66F8F1" authorityName="Li &amp; Zhao &amp; Wang" authorityYear="2024" box="[401,679,1844,1868]" class="Insecta" family="Vermileonidae" genus="Vermiophis" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="401" phylum="Arthropoda" rank="species" species="cangshanensis" status="sp. nov.">
<emphasis id="B927D260FFBE2275FE66EA1AFD66F8F1" bold="true" box="[401,679,1844,1868]" italics="true" pageId="1" pageNumber="401">Vermiophis cangshanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2146F1BFFBE2275FD5AEA1BFD3FF8F0" box="[685,766,1845,1869]" pageId="1" pageNumber="401" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Paratypes. A. Male, dorsal view; B. Female, dorsal view.
</paragraph>
</caption>
<subSubSection id="C3495DF9FFBE2276FF60EAB9FEFEFED8" lastPageId="2" lastPageNumber="402" pageId="1" pageNumber="401" type="diagnosis">
<paragraph id="8BEC0E72FFBE2276FF60EAB9FEFEFED8" blockId="1.[151,1436,1943,2042]" lastBlockId="2.[151,1437,151,609]" lastPageId="2" lastPageNumber="402" pageId="1" pageNumber="401">
<emphasis id="B927D260FFBE2275FF60EAB9FED3F80C" bold="true" box="[151,274,1943,1969]" pageId="1" pageNumber="401">Diagnosis.</emphasis>
By characters such as the short proboscis, the eight-segmented antennae without arista, the gradually tapering flagellomeres (
<bibRefCitation id="EFC27383FFBE2275FE5CEA95FD6FF868" author="Nagatomi, A. &amp; Yang, C. - K. &amp; Yang, D." box="[427,686,1979,2005]" pageId="1" pageNumber="401" pagination="1 - 154" refId="ref4234" refString="Nagatomi, A., Yang, C. - K. &amp; Yang, D. (1999) The Chinese species and the world genera of Vermileonidae (Diptera). Tropics, Monograph Series, 1, 1 - 154. https: // doi. org / 10.3759 / tropics. MonographNo. 1 _ 1" type="journal article" year="1999">
Nagatomi
<emphasis id="B927D260FFBE2275FDDFEA92FDA7F868" box="[552,614,1979,2005]" italics="true" pageId="1" pageNumber="401">et al.</emphasis>
1999
</bibRefCitation>
), and the general morphology of genitalia, we are confident in assigning this species into the genus
<emphasis id="B927D260FFBE2275FDC5EAF1FD79F844" box="[562,696,2015,2041]" italics="true" pageId="1" pageNumber="401">
<taxonomicName id="4C5375F1FFBE2275FDC5EAF1FD75F844" authorityName="Yang" authorityYear="1979" box="[562,692,2015,2041]" class="Insecta" family="Vermileonidae" genus="Vermiophis" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="401" phylum="Arthropoda" rank="genus">Vermiophis</taxonomicName>
.
</emphasis>
From its general appearance,
<taxonomicName id="4C5375F1FFBE2275FBF0EACEFB04F844" authorityName="Li &amp; Zhao &amp; Wang" authorityYear="2024" box="[1031,1221,2015,2041]" class="Insecta" family="Vermileonidae" genus="Vermiophis" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="401" phylum="Arthropoda" rank="species" species="cangshanensis" status="sp. nov.">
<emphasis id="B927D260FFBE2275FBF0EACEFB04F844" box="[1031,1221,2015,2041]" italics="true" pageId="1" pageNumber="401">V. cangshanensis</emphasis>
</taxonomicName>
<emphasis id="B927D260FFBE2275FB3BEAF1FAE7F844" bold="true" box="[1228,1318,2015,2041]" pageId="1" pageNumber="401">
<taxonomicNameLabel id="A2146F1BFFBE2275FB3BEAF1FAE7F844" box="[1228,1318,2015,2041]" pageId="1" pageNumber="401" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resembles
<taxonomicName id="4C5375F1FFBD2276FF60EDB9FEDEFF0C" authorityName="Yang &amp; Chen" authorityYear="1987" box="[151,287,151,177]" class="Insecta" family="Vermileonidae" genus="Vermiophis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="402" phylum="Arthropoda" rank="species" species="tibetensis">
<emphasis id="B927D260FFBD2276FF60EDB9FEDEFF0C" box="[151,287,151,177]" italics="true" pageId="2" pageNumber="402">V. tibetensis</emphasis>
</taxonomicName>
mostly among its congeners. However,
<taxonomicName id="4C5375F1FFBD2276FD1BEDB9FC6DFF0C" authorityName="Li &amp; Zhao &amp; Wang" authorityYear="2024" box="[748,940,151,177]" class="Insecta" family="Vermileonidae" genus="Vermiophis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="402" phylum="Arthropoda" rank="species" species="cangshanensis">
<emphasis id="B927D260FFBD2276FD1BEDB9FC6DFF0C" box="[748,940,151,177]" italics="true" pageId="2" pageNumber="402">V. cangshanensis</emphasis>
</taxonomicName>
can be differentiated from the latter by the following characters: 1) antennae with last flagellomere longer than the preceding two combined (
<emphasis id="B927D260FFBD2276FB00ED95FACFFF69" box="[1271,1294,187,212]" italics="true" pageId="2" pageNumber="402">vs</emphasis>
. shorter); 2) wings with R
<subScript id="17D70C37FFBD2276FED8EDDFFEF6FF42" attach="left" box="[303,311,241,255]" fontSize="6" pageId="2" pageNumber="402">5</subScript>
nearly straight (
<emphasis id="B927D260FFBD2276FE04EDF1FDCBFF45" box="[499,522,223,248]" italics="true" pageId="2" pageNumber="402">vs</emphasis>
. greatly curved); cell m
<subScript id="17D70C37FFBD2276FCEBEDDFFCE5FF42" attach="left" box="[796,804,241,255]" fontSize="6" pageId="2" pageNumber="402">3</subScript>
open (
<emphasis id="B927D260FFBD2276FC83EDF1FC4AFF45" box="[884,907,223,248]" italics="true" pageId="2" pageNumber="402">vs</emphasis>
. closed); M
<subScript id="17D70C37FFBD2276FBE2EDDFFBEFFF42" attach="left" box="[1045,1070,241,255]" fontSize="6" pageId="2" pageNumber="402">1+2</subScript>
furcating at m-m (
<emphasis id="B927D260FFBD2276FAF0EDF1FADFFF45" box="[1287,1310,223,248]" italics="true" pageId="2" pageNumber="402">vs</emphasis>
. before mm); M
<subScript id="17D70C37FFBD2276FF28EC3BFF38FE9E" attach="left" box="[223,249,277,291]" fontSize="6" pageId="2" pageNumber="402">3+4</subScript>
furcating before m-cu (
<emphasis id="B927D260FFBD2276FDF0EC2DFDDFFEA1" box="[519,542,259,284]" italics="true" pageId="2" pageNumber="402">vs</emphasis>
. at m-cu); 3) dorsomedial-posterior projection of fused gonocoxites absent (
<emphasis id="B927D260FFBD2276FA89EC2DFA54FEA1" box="[1406,1429,259,284]" italics="true" pageId="2" pageNumber="402">vs</emphasis>
. present); and 4) stout gonostylus approximately 1.4 times as long as its width (
<emphasis id="B927D260FFBD2276FBEBEC09FBF2FEFD" box="[1052,1075,295,320]" italics="true" pageId="2" pageNumber="402">vs</emphasis>
. slender and about 1.8 times as long as width).
</paragraph>
</subSubSection>
<subSubSection id="C3495DF9FFBD2276FF30EC41FE5CFE48" pageId="2" pageNumber="402" type="materials_examined">
<paragraph id="8BEC0E72FFBD2276FF30EC41FE5CFE48" blockId="2.[151,1437,151,609]" pageId="2" pageNumber="402">
<emphasis id="B927D260FFBD2276FF30EC41FEB8FE35" bold="true" box="[199,377,366,393]" pageId="2" pageNumber="402">Type material.</emphasis>
<materialsCitation id="3B3B042FFFBD2276FE73EC41FB30FE10" collectingDate="2022-11-01" collectorName="Dali Bai Autonomous Prefecture &amp; Dali City" country="China" latitude="100.13941" location="Cangshan Mountain" longLatPrecision="1" longitude="25.661617" pageId="2" pageNumber="402" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<collectingCountry id="F3444EE2FFBD2276FE73EC41FE0AFE34" box="[388,459,367,393]" name="China" pageId="2" pageNumber="402">China</collectingCountry>
:
<collectingRegion id="4997C090FFBD2276FE2EEC41FDF2FE34" box="[473,563,367,393]" country="China" name="Yunnan" pageId="2" pageNumber="402">Yunnan</collectingRegion>
:
<typeStatus id="54E8B0D0FFBD2276FDB6EC41FD6FFE34" box="[577,686,367,393]" pageId="2" pageNumber="402" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D55C5FBFFBD2276FD4BEC41FD10FE34" box="[700,721,367,393]" count="1" pageId="2" pageNumber="402" type="male"></specimenCount>
(DALUVer001),
<collectorName id="26A66BA4FFBD2276FC68EC41FADDFE34" box="[927,1308,367,393]" pageId="2" pageNumber="402">Dali Bai Autonomous Prefecture</collectorName>
,
<collectorName id="26A66BA4FFBD2276FADBEC41FA59FE34" box="[1324,1432,367,393]" pageId="2" pageNumber="402">Dali City</collectorName>
, eastern slope of the
<location id="8E8C58A9FFBD2276FE80ECBDFD9BFE10" LSID="urn:lsid:plazi:treatment:03FABF64FFBE2270FF60EEFCFF21FF68:8E8C58A9FFBD2276FE80ECBDFD9BFE10" box="[375,602,403,429]" country="China" latitude="100.13941" longLatPrecision="1" longitude="25.661617" name="Cangshan Mountain" pageId="2" pageNumber="402" stateProvince="Yunnan">Cangshan Mountain</location>
<geoCoordinate id="EE6768B5FFBD2276FD96ECBDFCC9FE10" box="[609,776,403,429]" degrees="100.139413" direction="north" orientation="latitude" pageId="2" pageNumber="402" precision="1" value="100.13941">100.139413°N</geoCoordinate>
,
<geoCoordinate id="EE6768B5FFBD2276FCE4ECBCFC68FE10" box="[787,937,402,429]" degrees="25.661618" direction="east" orientation="longitude" pageId="2" pageNumber="402" precision="1" value="25.661617">25.661618°E</geoCoordinate>
, collected as larva,
<date id="FFED28B2FFBD2276FB72ECBDFB30FE10" box="[1157,1265,403,429]" pageId="2" pageNumber="402" value="2022-11-01">
<collectingDate id="EFA9D15AFFBD2276FB72ECBDFB30FE10" box="[1157,1265,403,429]" pageId="2" pageNumber="402" value="2022-11-01">1.xi.2022</collectingDate>
</date>
</materialsCitation>
, adult emerged
<date id="FFED28B2FFBD2276FF60EC99FEC3FE6C" box="[151,258,439,465]" pageId="2" pageNumber="402" value="2023-04-07">7.iv.2023</date>
, collected and reared by Jia-Ling Li.
<materialsCitation id="3B3B042FFFBD2276FD6BEC99FEFAFE48" collectingDate="2021-06" collectorName="Dali Bai Autonomous Prefecture &amp; Dali City" country="China" latitude="100.13941" location="Cangshan Mountain" longLatPrecision="1" longitude="25.661617" pageId="2" pageNumber="402" specimenCount="23" specimenCount-female="12" specimenCount-male="11" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="54E8B0D0FFBD2276FD6BEC99FCCCFE6C" box="[668,781,439,465]" pageId="2" pageNumber="402" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D55C5FBFFBD2276FCE0EC99FC88FE6C" box="[791,841,439,465]" count="11" pageId="2" pageNumber="402" type="male">11♂</specimenCount>
<specimenCount id="9D55C5FBFFBD2276FCBEEC99FCB6FE6C" box="[841,887,439,465]" count="12" pageId="2" pageNumber="402" type="female">12♀</specimenCount>
(DALUVer002024), same data except collected during
<date id="FFED28B2FFBD2276FF10ECF5FEFAFE48" box="[231,315,475,501]" pageId="2" pageNumber="402" value="2021-06">
<collectingDate id="EFA9D15AFFBD2276FF10ECF5FEFAFE48" box="[231,315,475,501]" pageId="2" pageNumber="402" value="2021-06">vi.2021</collectingDate>
</date>
</materialsCitation>
<materialsCitation id="3B3B042FFFBD2276FEBCECF5FE58FE48" box="[331,409,475,501]" collectingDate="2023-05" collectorName="Dali Bai Autonomous Prefecture &amp; Dali City" country="China" isTailEnumeration="true" latitude="100.13941" location="Cangshan Mountain" longLatPrecision="1" longitude="25.661617" pageId="2" pageNumber="402" specimenCount="1" stateProvince="Yunnan" typeStatus="paratype">
<date id="FFED28B2FFBD2276FEBCECF5FE58FE48" box="[331,409,475,501]" pageId="2" pageNumber="402" value="2023-05">
<collectingDate id="EFA9D15AFFBD2276FEBCECF5FE58FE48" box="[331,409,475,501]" pageId="2" pageNumber="402" value="2023-05">v.2023</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3495DF9FFBD2276FF30ECD1FBA5FDA4" box="[199,1124,511,537]" pageId="2" pageNumber="402" type="etymology">
<paragraph id="8BEC0E72FFBD2276FF30ECD1FBA5FDA4" blockId="2.[151,1437,151,609]" box="[199,1124,511,537]" pageId="2" pageNumber="402">
<emphasis id="B927D260FFBD2276FF30ECD1FE8FFDA4" bold="true" box="[199,334,511,537]" pageId="2" pageNumber="402">Etymology.</emphasis>
The new species is named after the
<typeStatus id="54E8B0D0FFBD2276FD14ECD1FCD2FDA4" box="[739,787,511,537]" pageId="2" pageNumber="402">type</typeStatus>
locality, Cangshan Mountain.
</paragraph>
</subSubSection>
<subSubSection id="C3495DF9FFBD2270FF30EF0DFF21FF68" lastPageId="4" lastPageNumber="404" pageId="2" pageNumber="402" type="description">
<paragraph id="8BEC0E72FFBD2276FF30EF0DFADAFDDC" blockId="2.[151,1437,151,609]" pageId="2" pageNumber="402">
<emphasis id="B927D260FFBD2276FF30EF0DFEBFFD80" bold="true" box="[199,382,547,573]" pageId="2" pageNumber="402">Measurements.</emphasis>
Male. Body length: 12.0
<quantity id="4CABA397FFBD2276FD5AEF0CFCD5FD80" box="[685,788,546,573]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="2" pageNumber="402" unit="mm" value="14.5">14.5 mm</quantity>
, wing length: 8.0
<quantity id="4CABA397FFBD2276FC1BEF0DFB92FD80" box="[1004,1107,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="1.3599999999999999" pageId="2" pageNumber="402" unit="mm" value="13.6">13.6 mm</quantity>
, wing width:
<quantity id="4CABA397FFBD2276FB04EF0CFABDFD80" box="[1267,1404,546,573]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" metricValueMax="3.3" metricValueMin="2.5" pageId="2" pageNumber="402" unit="mm" value="2.9" valueMax="3.3" valueMin="2.5">2.53.3 mm</quantity>
(n =12). Female. Body length:
<quantity id="4CABA397FFBD2276FE24EF69FDB9FDDC" box="[467,632,583,609]" metricMagnitude="-2" metricUnit="m" metricValue="1.335" metricValueMax="1.46" metricValueMin="1.21" pageId="2" pageNumber="402" unit="mm" value="13.35" valueMax="14.6" valueMin="12.1">12.114.6 mm</quantity>
, wing length:
<quantity id="4CABA397FFBD2276FCEFEF69FC6EFDDC" box="[792,943,583,609]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.46" metricValueMin="0.94" pageId="2" pageNumber="402" unit="mm" value="12.0" valueMax="14.6" valueMin="9.4">9.414.6 mm</quantity>
, wing width:
<quantity id="4CABA397FFBD2276FBBFEF68FB7BFDDC" box="[1096,1210,582,609]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" metricValueMax="5.0" metricValueMin="2.8" pageId="2" pageNumber="402" unit="mm" value="3.9" valueMax="5.0" valueMin="2.8">2.85 mm</quantity>
(n =12).
</paragraph>
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<paragraph id="8BEC0E72FFBD2276FF60EA28FA8BF8FF" blockId="2.[151,1436,1798,1858]" pageId="2" pageNumber="402">
<emphasis id="B927D260FFBD2276FF60EA28FBCEF8A3" bold="true" box="[151,1039,1798,1822]" pageId="2" pageNumber="402">
FIGURE 2. Morphological characters of
<taxonomicName id="4C5375F1FFBD2276FDA0EA28FCACF8A3" authorityName="Li &amp; Zhao &amp; Wang" authorityYear="2024" box="[599,877,1798,1822]" class="Insecta" family="Vermileonidae" genus="Vermiophis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="402" phylum="Arthropoda" rank="species" species="cangshanensis" status="sp. nov.">
<emphasis id="B927D260FFBD2276FDA0EA28FCACF8A3" bold="true" box="[599,877,1798,1822]" italics="true" pageId="2" pageNumber="402">Vermiophis cangshanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2146F1BFFBD2276FC82EA28FC09F8A3" box="[885,968,1798,1822]" pageId="2" pageNumber="402" rank="species">sp. nov.</taxonomicNameLabel>
, male.
</emphasis>
Paratypes. A. Habitus, dorsal view; B. Right antenna, dorsal view; CE. Head, left-lateral, dorsal, and ventral views, respectively; F. Right wing, dorsal view.
</paragraph>
</caption>
<paragraph id="8BEC0E72FFBD2277FF30EABAFD4BFD38" blockId="2.[151,1436,1940,2039]" lastBlockId="3.[151,1437,151,645]" lastPageId="3" lastPageNumber="403" pageId="2" pageNumber="402">
<emphasis id="B927D260FFBD2276FF30EABAFE20F812" bold="true" box="[199,481,1940,1967]" pageId="2" pageNumber="402">Description-male. Head</emphasis>
(
<figureCitation id="136812F7FFBD2276FE18EABAFDF5F812" box="[495,564,1940,1967]" captionStart="FIGURE 1" captionStartId="1.[151,250,1845,1869]" captionTargetBox="[226,1362,1075,1815]" captionTargetId="figure-270@1.[213,1375,1062,1818]" captionTargetPageId="1" captionText="FIGURE 1. Habitus of Vermiophis cangshanensis sp. nov. Paratypes.A. Male, dorsal view; B. Female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13618560" httpUri="https://zenodo.org/record/13618560/files/figure.png" pageId="2" pageNumber="402">Figs 1</figureCitation>
,
<figureCitation id="136812F7FFBD2276FDB6EABBFD45F812" box="[577,644,1941,1967]" captionStart="FIGURE 2" captionStartId="2.[151,250,1798,1822]" captionTargetBox="[194,1436,668,1758]" captionTargetId="figure-213@2.[151,1436,633,1775]" captionTargetPageId="2" captionText="FIGURE 2. Morphological characters of Vermiophis cangshanensis sp. nov., male. Paratypes. A. Habitus, dorsal view; B. Right antenna, dorsal view; CE. Head, left-lateral, dorsal, and ventral views, respectively; F. Right wing, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13618562" httpUri="https://zenodo.org/record/13618562/files/figure.png" pageId="2" pageNumber="402">2AE</figureCitation>
). Antennae with scape and pedicel yellowish brown. Scape nearly three times as long as pedicel. Flagellum eight-segmented, gradually darkening from second flagellomere towards last two. Proboscis and maxillary palps yellow. Vertex and frons dark brown and glabrous. Ocellar tubercle black with dense and short setae. Occiput and genae black and covered with yellowish long setae.
<emphasis id="B927D260FFBC2277FB96EDB9FB7BFF0C" bold="true" box="[1121,1210,151,177]" pageId="3" pageNumber="403">Thorax</emphasis>
(
<figureCitation id="136812F7FFBC2277FB3FEDB9FACFFF0C" box="[1224,1294,151,177]" captionStart="FIGURE 1" captionStartId="1.[151,250,1845,1869]" captionTargetBox="[226,1362,1075,1815]" captionTargetId="figure-270@1.[213,1375,1062,1818]" captionTargetPageId="1" captionText="FIGURE 1. Habitus of Vermiophis cangshanensis sp. nov. Paratypes.A. Male, dorsal view; B. Female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13618560" httpUri="https://zenodo.org/record/13618560/files/figure.png" pageId="3" pageNumber="403">Figs 1</figureCitation>
,
<figureCitation id="136812F7FFBC2277FAEBEDB9FA81FF0C" box="[1308,1344,151,177]" captionStart="FIGURE 2" captionStartId="2.[151,250,1798,1822]" captionTargetBox="[194,1436,668,1758]" captionTargetId="figure-213@2.[151,1436,633,1775]" captionTargetPageId="2" captionText="FIGURE 2. Morphological characters of Vermiophis cangshanensis sp. nov., male. Paratypes. A. Habitus, dorsal view; B. Right antenna, dorsal view; CE. Head, left-lateral, dorsal, and ventral views, respectively; F. Right wing, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13618562" httpUri="https://zenodo.org/record/13618562/files/figure.png" pageId="3" pageNumber="403">2A</figureCitation>
). Meso- and metanotum mostly shining black. Mesonotum with two yellowish brown stripes ending before scutellum. Legs: Fore- and midlegs mostly yellowish brown. Hindleg with femur dark brown except yellowish brown middle and apical portions; tibia dark brown except yellowish brown base and apex; tarsus light brown.
<emphasis id="B927D260FFBC2277FB51EC2CFB26FEA1" bold="true" box="[1190,1255,258,284]" pageId="3" pageNumber="403">Wing</emphasis>
(
<figureCitation id="136812F7FFBC2277FB01EC2DFAFCFEA0" box="[1270,1341,259,285]" captionStart="FIGURE 1" captionStartId="1.[151,250,1845,1869]" captionTargetBox="[226,1362,1075,1815]" captionTargetId="figure-270@1.[213,1375,1062,1818]" captionTargetPageId="1" captionText="FIGURE 1. Habitus of Vermiophis cangshanensis sp. nov. Paratypes.A. Male, dorsal view; B. Female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13618560" httpUri="https://zenodo.org/record/13618560/files/figure.png" pageId="3" pageNumber="403">Figs 1</figureCitation>
,
<figureCitation id="136812F7FFBC2277FABBEC2DFA4CFEA0" box="[1356,1421,259,285]" captionStart="FIGURE 2" captionStartId="2.[151,250,1798,1822]" captionTargetBox="[194,1436,668,1758]" captionTargetId="figure-213@2.[151,1436,633,1775]" captionTargetPageId="2" captionText="FIGURE 2. Morphological characters of Vermiophis cangshanensis sp. nov., male. Paratypes. A. Habitus, dorsal view; B. Right antenna, dorsal view; CE. Head, left-lateral, dorsal, and ventral views, respectively; F. Right wing, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13618562" httpUri="https://zenodo.org/record/13618562/files/figure.png" pageId="3" pageNumber="403">2A, F</figureCitation>
). Wing membrane hyaline with pale yellowish tinge in basal portion and grayish in distal portion; veins dark brown. Dark brown patterns in distal portion of cell br, base of cell d, and along CuA. R
<subScript id="17D70C37FFBC2277FBCAEC73FB97FED6" attach="left" box="[1085,1110,349,363]" fontSize="6" pageId="3" pageNumber="403">2+3</subScript>
curved anteriorly at end; R
<subScript id="17D70C37FFBC2277FA63EC73FA5DFED6" attach="left" box="[1428,1436,349,363]" fontSize="6" pageId="3" pageNumber="403">5</subScript>
nearly straight; M
<subScript id="17D70C37FFBC2277FEA8ECAFFEB9FE32" attach="left" box="[351,376,385,399]" fontSize="6" pageId="3" pageNumber="403">1+2</subScript>
furcated at m-m; M
<subScript id="17D70C37FFBC2277FDA0ECAFFDB0FE32" attach="left" box="[599,625,385,399]" fontSize="6" pageId="3" pageNumber="403">3+4</subScript>
furcated before m-cu; cell m
<subScript id="17D70C37FFBC2277FC44ECAFFC7AFE32" attach="left" box="[947,955,385,399]" fontSize="6" pageId="3" pageNumber="403">3</subScript>
slightly open (closed in a few individuals), cell cua open. Halteres club-shaped.
<emphasis id="B927D260FFBC2277FDC5ECBCFD66FE11" bold="true" box="[562,679,402,428]" pageId="3" pageNumber="403">Abdomen</emphasis>
(
<figureCitation id="136812F7FFBC2277FD41ECBDFD3DFE10" box="[694,764,403,429]" captionStart="FIGURE 1" captionStartId="1.[151,250,1845,1869]" captionTargetBox="[226,1362,1075,1815]" captionTargetId="figure-270@1.[213,1375,1062,1818]" captionTargetPageId="1" captionText="FIGURE 1. Habitus of Vermiophis cangshanensis sp. nov. Paratypes.A. Male, dorsal view; B. Female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13618560" httpUri="https://zenodo.org/record/13618560/files/figure.png" pageId="3" pageNumber="403">Figs 1</figureCitation>
,
<figureCitation id="136812F7FFBC2277FCFDECBDFCEFFE10" box="[778,814,403,429]" captionStart="FIGURE 2" captionStartId="2.[151,250,1798,1822]" captionTargetBox="[194,1436,668,1758]" captionTargetId="figure-213@2.[151,1436,633,1775]" captionTargetPageId="2" captionText="FIGURE 2. Morphological characters of Vermiophis cangshanensis sp. nov., male. Paratypes. A. Habitus, dorsal view; B. Right antenna, dorsal view; CE. Head, left-lateral, dorsal, and ventral views, respectively; F. Right wing, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13618562" httpUri="https://zenodo.org/record/13618562/files/figure.png" pageId="3" pageNumber="403">2A</figureCitation>
). Yellowish to dark brown, usually lighter in proximate segments. Segment III narrowest.
<emphasis id="B927D260FFBC2277FDEEEC98FD5DFE6D" bold="true" box="[537,668,438,464]" pageId="3" pageNumber="403">Terminalia</emphasis>
(
<figureCitation id="136812F7FFBC2277FD5BEC99FD30FE6C" box="[684,753,439,465]" captionStart="FIGURE 3" captionStartId="3.[151,250,1920,1944]" captionTargetBox="[209,1379,682,1884]" captionTargetId="figure-253@3.[196,1392,669,1897]" captionTargetPageId="3" captionText="FIGURE 3. Male genitalia of Vermiophis cangshanensis sp. nov. Paratype.A. T9 (epandrium),dorsal view; BE.Hypopygium, ventral, caudal, dorsal, and left-lateral views, respectively.Abbreviations: ae, aedeagus; aap, aedeagal apodeme; ce, cercus; dbr, dorsal bridge; gap, gonocoxital apodeme; gcx, gonocoxite; gs, gonostylus; vp, ventral plane." figureDoi="http://doi.org/10.5281/zenodo.13618564" httpUri="https://zenodo.org/record/13618564/files/figure.png" pageId="3" pageNumber="403">Fig. 3</figureCitation>
). T9 (epandrium) subtrapezoidal with terminal emargination and a pair of postro-lateral processes. Ventral plane subtrapezoidal, and wider than long. Cercus short and broad, expanded basally. S10 (sternum X) subsemicircular and concealed by T
<quantity id="4CABA397FFBC2277FC42ECD1FC1EFDA4" box="[949,991,511,537]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="3" pageNumber="403" unit="in" value="9.0">9 in</quantity>
dorsal view. Fused gonocoxites lacking dorsomedial-posterior projection; posterolateral ventral part long and tapering towards apex. Gonostylus broad, approximately 1.4 times as long as its width, with distal-inner potion slightly elongated and pointed. Aedeagus with apex greatly bending ventrad in lateral view.
</paragraph>
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<paragraph id="8BEC0E72FFBC2277FF60EAAEFB81F85C" blockId="3.[151,1437,1920,2017]" pageId="3" pageNumber="403">
<emphasis id="B927D260FFBC2277FF60EAAEFECAF825" bold="true" box="[151,267,1920,1944]" pageId="3" pageNumber="403">FIGURE 3</emphasis>
.
<emphasis id="B927D260FFBC2277FEE2EAAFFCF1F824" bold="true" box="[277,816,1920,1945]" pageId="3" pageNumber="403">
Male genitalia of
<taxonomicName id="4C5375F1FFBC2277FE3DEAAEFD1CF825" authorityName="Li &amp; Zhao &amp; Wang" authorityYear="2024" box="[458,733,1920,1944]" class="Insecta" family="Vermileonidae" genus="Vermiophis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="403" phylum="Arthropoda" rank="species" species="cangshanensis" status="sp. nov.">
<emphasis id="B927D260FFBC2277FE3DEAAEFD1CF825" bold="true" box="[458,733,1920,1944]" italics="true" pageId="3" pageNumber="403">Vermiophis cangshanensis</emphasis>
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<taxonomicNameLabel id="A2146F1BFFBC2277FD15EAAFFCF1F824" box="[738,816,1921,1945]" pageId="3" pageNumber="403" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Paratype.A. T9 (epandrium), dorsal view; BE. Hypopygium, ventral, caudal, dorsal, and left-lateral views, respectively. Abbreviations: ae, aedeagus; aap, aedeagal apodeme; ce, cercus; dbr, dorsal bridge; gap, gonocoxital apodeme; gcx, gonocoxite; gs, gonostylus; vp, ventral plane.
</paragraph>
</caption>
<paragraph id="8BEC0E72FFBB2270FF30EDB8FF21FF68" blockId="4.[151,1436,150,213]" pageId="4" pageNumber="404">
<emphasis id="B927D260FFBB2270FF30EDB8FE70FF0D" bold="true" box="[199,433,150,176]" pageId="4" pageNumber="404">Description-female.</emphasis>
Habitus (
<figureCitation id="136812F7FFBB2270FDD0EDB9FD41FF0C" box="[551,640,151,177]" captionStart="FIGURE 1" captionStartId="1.[151,250,1845,1869]" captionTargetBox="[226,1362,1075,1815]" captionTargetId="figure-270@1.[213,1375,1062,1818]" captionTargetPageId="1" captionText="FIGURE 1. Habitus of Vermiophis cangshanensis sp. nov. Paratypes.A. Male, dorsal view; B. Female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13618560" httpUri="https://zenodo.org/record/13618560/files/figure.png" pageId="4" pageNumber="404">Fig. 1B</figureCitation>
) similar to males, but yellow parts in hindlegs longer than those in males.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,126 @@
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<emphasis id="93FEEA97FFE25D4299A6FEDC1B92A5FC" bold="true" box="[151,468,1415,1442]" pageId="3" pageNumber="304">
<emphasis id="93FEEA97FFE25D4299A6FEDC1B75A5FF" bold="true" box="[151,307,1415,1442]" italics="true" pageId="3" pageNumber="304">Lathys arabs</emphasis>
Simon, 1910
</emphasis>
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<emphasis id="93FEEA97FFE25D4299A6FE971B5DA5B9" box="[151,283,1484,1508]" italics="true" pageId="3" pageNumber="304">Lathys arabs</emphasis>
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;
<treatmentCitation id="202B1094FFE25D429819FE971B9FA5B9" author="Simon, E." box="[296,473,1484,1508]" page="278" pageId="3" pageNumber="304" year="1911">
<bibRefCitation id="C51B4B74FFE25D429819FE971B9FA5B9" author="Simon, E." box="[296,473,1484,1508]" pageId="3" pageNumber="304" pagination="265 - 332" refId="ref23418" refString="Simon, E. (1911) Catalogue raisonne des arachnides du nord de l'Afrique (1 re partie). Annales de la Societe Entomologique de France, 79 (3), 265 - 332." type="journal article" year="1911">Simon 1911: 278</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFE25D4299A6FD4A1E30A60D" pageId="3" pageNumber="304" type="materials_examined">
<paragraph id="A1353685FFE25D4299A6FD4A1E6AA671" blockId="3.[151,1225,1553,1652]" box="[151,1068,1553,1580]" pageId="3" pageNumber="304">
<emphasis id="93FEEA97FFE25D4299A6FD4A1BC5A676" bold="true" box="[151,387,1553,1579]" pageId="3" pageNumber="304">Material examined:</emphasis>
BERRIANE: Sidi Mbarek, hand collecting,
<specimenCount id="B78CFD0CFFE25D429A49FD4919E4A671" box="[888,930,1554,1580]" count="1" pageId="3" pageNumber="304" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFE25D429A81FD491E61A671" box="[944,1063,1554,1580]" pageId="3" pageNumber="304" value="2021-02-18">18.II.2021</date>
.
</paragraph>
<paragraph id="A1353685FFE25D4299F6FD6E1E30A60D" blockId="3.[151,1225,1553,1652]" box="[199,1142,1589,1616]" pageId="3" pageNumber="304">
<emphasis id="93FEEA97FFE25D4299F6FD6E1BE8A612" bold="true" box="[199,430,1589,1616]" pageId="3" pageNumber="304">Previous records in</emphasis>
<materialsCitation id="11E23CD8FFE25D429882FD6E1E34A60D" box="[435,1138,1589,1616]" country="Algeria" location="Biskra" pageId="3" pageNumber="304" specimenCount="1" stateProvince="Biskra">
<emphasis id="93FEEA97FFE25D429882FD6E1853A612" bold="true" box="[435,533,1589,1615]" pageId="3" pageNumber="304">
<collectingCountry id="D99D7615FFE25D429882FD6E1856A612" box="[435,528,1589,1615]" name="Algeria" pageId="3" pageNumber="304">Algeria</collectingCountry>
:
</emphasis>
Known from
<collectingRegion id="634EF867FFE25D429B82FD6D18B9A60D" box="[691,767,1590,1616]" country="Algeria" name="Biskra" pageId="3" pageNumber="304">Biskra</collectingRegion>
,
<collectingRegion id="634EF867FFE25D429A3BFD6D1916A60D" box="[778,848,1590,1616]" country="Algeria" name="Djelfa" pageId="3" pageNumber="304">Djelfa</collectingRegion>
and
<collectingRegion id="634EF867FFE25D429AB6FD6D1989A60D" box="[903,975,1590,1616]" country="Algeria" name="M'Sila" pageId="3" pageNumber="304">Msila</collectingRegion>
(
<bibRefCitation id="C51B4B74FFE25D429AECFD6D1E2CA60D" author="Simon, E." box="[989,1130,1590,1616]" pageId="3" pageNumber="304" pagination="265 - 332" refId="ref23418" refString="Simon, E. (1911) Catalogue raisonne des arachnides du nord de l'Afrique (1 re partie). Annales de la Societe Entomologique de France, 79 (3), 265 - 332." type="journal article" year="1911">Simon 1911</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFE25D4299F6FD021E8FA629" box="[199,1225,1625,1652]" pageId="3" pageNumber="304" type="distribution">
<paragraph id="A1353685FFE25D4299F6FD021E8FA629" blockId="3.[151,1225,1553,1652]" box="[199,1225,1625,1652]" pageId="3" pageNumber="304">
<emphasis id="93FEEA97FFE25D4299F6FD021B25A62E" bold="true" box="[199,355,1625,1651]" pageId="3" pageNumber="304">Distribution:</emphasis>
<collectingCountry id="D99D7615FFE25D429859FD011BF9A629" box="[360,447,1626,1652]" name="Algeria" pageId="3" pageNumber="304">Algeria</collectingCountry>
,
<collectingCountry id="D99D7615FFE25D4298FBFD011866A629" box="[458,544,1626,1652]" name="Tunisia" pageId="3" pageNumber="304">Tunisia</collectingCountry>
,
<collectingCountry id="D99D7615FFE25D429B1AFD011818A629" box="[555,606,1626,1652]" name="Italy" pageId="3" pageNumber="304">Italy</collectingCountry>
(
<collectingRegion id="634EF867FFE25D429B5CFD0118F5A629" box="[621,691,1626,1652]" country="Italy" name="Sicilia" pageId="3" pageNumber="304">Sicily</collectingRegion>
),
<collectingCountry id="D99D7615FFE25D429BF6FD01195FA629" box="[711,793,1626,1652]" name="Greece" pageId="3" pageNumber="304">Greece</collectingCountry>
,
<collectingCountry id="D99D7615FFE25D429A15FD011933A629" box="[804,885,1626,1652]" name="Cyprus" pageId="3" pageNumber="304">Cyprus</collectingCountry>
(
<bibRefCitation id="C51B4B74FFE25D429AB2FD011EFBA629" author="World Spider Catalog" box="[899,1213,1626,1652]" pageId="3" pageNumber="304" refId="ref24111" refString="World Spider Catalog (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern, Bern. Available from: https: // wsc. nmbe. ch / (accessed 1 June 2024) https: // doi. org / 10.24436 / 2" type="book" year="2024">World Spider Catalog 2024</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,149 @@
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<paragraph id="A1353685FFE45D4499A6F9AE1909A352" blockId="5.[151,847,757,784]" box="[151,847,757,784]" pageId="5" pageNumber="306">
<heading id="FA7D81E9FFE45D4499A6F9AE1909A352" bold="true" box="[151,847,757,784]" fontSize="11" level="1" pageId="5" pageNumber="306" reason="1">
<emphasis id="93FEEA97FFE45D4499A6F9AE1909A352" bold="true" box="[151,847,757,784]" pageId="5" pageNumber="306">
<taxonomicName id="668A4D06FFE45D4499A6F9AE185DA352" ID-CoL="3TT97" authority="(Dalmas, 1919)" baseAuthorityName="Dalmas" baseAuthorityYear="1919" box="[151,539,757,784]" class="Arachnida" family="Gnaphosidae" genus="Leptopilos" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="pupa">
<emphasis id="93FEEA97FFE45D4499A6F9AE1B1EA34D" bold="true" box="[151,344,757,784]" italics="true" pageId="5" pageNumber="306">Leptopilos pupa</emphasis>
(
<bibRefCitation id="C51B4B74FFE45D449859F9AE1855A352" author="Dalmas, R. de" box="[360,531,757,783]" pageId="5" pageNumber="306" pagination="243 - 250" refId="ref20441" refString="Dalmas, R. de (1919) Catalogue des araignees du genre Leptodrassus (Gnaphosidae) d'apres les materiaux de la collection E. Simon au Museum d'Histoire naturelle. Bulletin du Museum National d'Histoire Naturelle de Paris, 25, 243 - 250. https: // doi. org / 10.5962 / bhl. part. 7927" type="journal article" year="1919">Dalmas, 1919</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="39B12A00FFE45D449B1AF9AE18EEA352" box="[555,680,757,783]" captionStart="FIGURE 3" captionStartId="5.[152,255,1599,1623]" captionTargetBox="[151,1436,1085,1575]" captionTargetId="figure-197@5.[151,1436,1085,1575]" captionTargetPageId="5" captionText="FIGURE 3ac. Leptopilos pupa, (a) Dorsal view of the male, (b) Dorsal view of the female, (c) Ventral view of the female, (c)." pageId="5" pageNumber="306">Figs. 3ac</figureCitation>
&amp; figs. 4ah)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFE45D4499A6F8611B90A32C" pageId="5" pageNumber="306" type="reference_group">
<paragraph id="A1353685FFE45D4499A6F8611895A30F" blockId="5.[151,723,826,881]" box="[151,723,826,850]" pageId="5" pageNumber="306">
<treatmentCitationGroup id="819A11ABFFE45D4499A6F8611895A30F" box="[151,723,826,850]" pageId="5" pageNumber="306">
<taxonomicName id="668A4D06FFE45D4499A6F8611B1EA30F" authorityName="De Dalmas" authorityYear="1919" box="[151,344,826,850]" class="Arachnida" family="Gnaphosidae" genus="Leptodrassus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="pupa">
<emphasis id="93FEEA97FFE45D4499A6F8611B1EA30F" box="[151,344,826,850]" italics="true" pageId="5" pageNumber="306">Leptodrassus pupa</emphasis>
</taxonomicName>
;
<treatmentCitation id="202B1094FFE45D449854F8611867A30F" author="Dalmas, R. de" box="[357,545,826,850]" page="248" pageId="5" pageNumber="306" year="1919">
<bibRefCitation id="C51B4B74FFE45D449854F8611867A30F" author="Dalmas, R. de" box="[357,545,826,850]" pageId="5" pageNumber="306" pagination="243 - 250" refId="ref20441" refString="Dalmas, R. de (1919) Catalogue des araignees du genre Leptodrassus (Gnaphosidae) d'apres les materiaux de la collection E. Simon au Museum d'Histoire naturelle. Bulletin du Museum National d'Histoire Naturelle de Paris, 25, 243 - 250. https: // doi. org / 10.5962 / bhl. part. 7927" type="journal article" year="1919">Dalmas 1919: 248</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="202B1094FFE45D449B1DF8611889A30F" author="Levy, G." box="[556,719,826,850]" page="446" pageId="5" pageNumber="306" year="1999">
<bibRefCitation id="C51B4B74FFE45D449B1DF8611889A30F" author="Levy, G." box="[556,719,826,850]" pageId="5" pageNumber="306" pagination="427 - 452" refId="ref21470" refString="Levy, G. (1999) Spiders of six uncommon drassodine genera (Araneae: Gnaphosidae) from Israel. Israel Journal of Zoology, 45, 427 - 452." type="journal article" year="1999">Levy 1999: 446</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A1353685FFE45D4499A6F8031B90A32C" blockId="5.[151,723,826,881]" box="[151,470,856,881]" pageId="5" pageNumber="306">
<treatmentCitationGroup id="819A11ABFFE45D4499A6F8031B90A32C" box="[151,470,856,881]" pageId="5" pageNumber="306">
<taxonomicName id="668A4D06FFE45D4499A6F8031B7DA32D" ID-CoL="3TT97" baseAuthorityName="Dalmas" baseAuthorityYear="1919" box="[151,315,856,880]" class="Arachnida" family="Gnaphosidae" genus="Leptopilos" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="pupa">
<emphasis id="93FEEA97FFE45D4499A6F8031B7DA32D" box="[151,315,856,880]" italics="true" pageId="5" pageNumber="306">Leptopilos pupa</emphasis>
</taxonomicName>
;
<treatmentCitation id="202B1094FFE45D449878F8021B94A32C" author="Levy, G." box="[329,466,857,881]" page="9" pageId="5" pageNumber="306" year="2009">
<bibRefCitation id="C51B4B74FFE45D449878F8021B94A32C" author="Levy, G." box="[329,466,857,881]" pageId="5" pageNumber="306" pagination="1 - 49" refId="ref21502" refString="Levy, G. (2009) New ground-spider genera and species with annexed checklist of the Gnaphosidae (Araneae) of Israel. Zootaxa, 2066 (1), 1 - 49. https: // doi. org / 10.11646 / zootaxa. 2066.1.1" type="journal article" year="2009">Levy 2009: 9</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFE45D4499A6F8C51E3FA380" pageId="5" pageNumber="306" type="materials_examined">
<paragraph id="A1353685FFE45D4499A6F8C51EB0A3E4" blockId="5.[151,1270,926,1061]" box="[151,1270,926,953]" pageId="5" pageNumber="306">
<emphasis id="93FEEA97FFE45D4499A6F8C51BC5A3E5" bold="true" box="[151,387,926,952]" pageId="5" pageNumber="306">Material examined:</emphasis>
BERRIANE: Sidi Mbarek, hand collecting,
<specimenCount id="B78CFD0CFFE45D449A49F8C419E4A3E4" box="[888,930,927,953]" count="1" pageId="5" pageNumber="306" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFE45D449A81F8C41E68A3E4" box="[944,1070,927,953]" pageId="5" pageNumber="306" value="2021-04-23">23.IV.2021</date>
;
<materialsCitation id="11E23CD8FFE45D449D0BF8C41EB4A3E4" box="[1082,1266,927,953]" collectingDate="2021-04-29" isTailEnumeration="true" pageId="5" pageNumber="306" specimenCount="1" specimenCount-male="1">
<specimenCount id="B78CFD0CFFE45D449D0BF8C41E20A3E4" box="[1082,1126,927,953]" count="1" pageId="5" pageNumber="306" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFE45D449D43F8C41EB4A3E4" box="[1138,1266,927,953]" pageId="5" pageNumber="306" value="2021-04-29">
<collectingDate id="C570E9ADFFE45D449D43F8C41EB4A3E4" box="[1138,1266,927,953]" pageId="5" pageNumber="306" value="2021-04-29">29.IV.2021</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="A1353685FFE45D4499F6F8991E3FA380" blockId="5.[151,1270,926,1061]" box="[199,1145,962,989]" pageId="5" pageNumber="306">
<emphasis id="93FEEA97FFE45D4499F6F8991853A381" bold="true" box="[199,533,962,989]" pageId="5" pageNumber="306">
Previous records in
<collectingCountry id="D99D7615FFE45D449882F8991856A381" box="[435,528,962,988]" name="Algeria" pageId="5" pageNumber="306">Algeria</collectingCountry>
:
</emphasis>
this is a new genus and species record for the country.
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFE45D4499F6F8BD1967A45C" box="[199,801,998,1025]" pageId="5" pageNumber="306" type="distribution">
<paragraph id="A1353685FFE45D4499F6F8BD1967A45C" blockId="5.[151,1270,926,1061]" box="[199,801,998,1025]" pageId="5" pageNumber="306">
<emphasis id="93FEEA97FFE45D4499F6F8BD1B25A45D" bold="true" box="[199,355,998,1024]" pageId="5" pageNumber="306">Distribution:</emphasis>
The species is known only from
<collectingCountry id="D99D7615FFE45D449BE6F8BC1958A45C" box="[727,798,999,1025]" name="Egypt" pageId="5" pageNumber="306">Egypt</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFE45D4499F6FF511E1CA478" box="[199,1114,1034,1061]" pageId="5" pageNumber="306" type="discussion">
<paragraph id="A1353685FFE45D4499F6FF511E1CA478" blockId="5.[151,1270,926,1061]" box="[199,1114,1034,1061]" pageId="5" pageNumber="306">
<emphasis id="93FEEA97FFE45D4499F6FF511B01A479" bold="true" box="[199,327,1034,1060]" pageId="5" pageNumber="306">Comment:</emphasis>
This is the first record of the Genus
<taxonomicName id="668A4D06FFE45D449BD3FF50199AA478" authority="Levy, 2009" authorityName="Levy" authorityYear="2009" box="[738,988,1035,1061]" class="Arachnida" family="Gnaphosidae" genus="Leptopilos" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis id="93FEEA97FFE45D449BD3FF50191EA478" box="[738,856,1035,1061]" italics="true" pageId="5" pageNumber="306">Leptopilos</emphasis>
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Levy,
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<emphasis id="93FEEA97FFE45D4499A6FA351BE2A1D4" bold="true" box="[151,420,366,393]" italics="true" pageId="5" pageNumber="306">Haplodrassus dentifer</emphasis>
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Bosmans
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2018: 15
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BERRIANE: Balouh, hand collecting,
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. EL ATTEUF: Palm grove of El Atteuf, hand collecting,
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<emphasis id="93FEEA97FFE45D4499F6F93E1BE8A222" bold="true" box="[199,430,613,639]" pageId="5" pageNumber="306">Previous records in</emphasis>
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<collectingCountry id="D99D7615FFE45D449882F93E1856A222" box="[435,528,613,639]" name="Algeria" pageId="5" pageNumber="306">Algeria</collectingCountry>
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(
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Bosmans
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<emphasis id="93FEEA97FFE45D4499F6F9D21B25A2FE" bold="true" box="[199,355,649,675]" pageId="5" pageNumber="306">Distribution:</emphasis>
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,
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,
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<emphasis id="93FEEA97FFE45D4499A6FD961BBAA6B5" bold="true" box="[151,508,1741,1768]" italics="true" pageId="5" pageNumber="306">Marinarozelotes holosericeus</emphasis>
(
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)
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<paragraph id="A1353685FFE45D4499A6FC4918A4A777" blockId="5.[151,738,1810,1834]" box="[151,738,1810,1834]" pageId="5" pageNumber="306">
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<taxonomicName id="668A4D06FFE45D4499A6FC491BEFA777" ID-CoL="7CMH5" baseAuthorityName="Simon" baseAuthorityYear="1878" box="[151,425,1810,1834]" class="Arachnida" family="Gnaphosidae" genus="Trachyzelotes" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="holosericeus">
<emphasis id="93FEEA97FFE45D4499A6FC491BEFA777" box="[151,425,1810,1834]" italics="true" pageId="5" pageNumber="306">Trachyzelotes holosericeus</emphasis>
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<treatmentCitation id="202B1094FFE45D449887FC491898A777" author="Platnick, N. I. &amp; Murphy, J. A." box="[438,734,1810,1834]" page="17" pageId="5" pageNumber="306" year="1984">
<bibRefCitation id="C51B4B74FFE45D449887FC491898A777" author="Platnick, N. I. &amp; Murphy, J. A." box="[438,734,1810,1834]" pageId="5" pageNumber="306" pagination="1 - 30" refId="ref22383" refString="Platnick, N. I. &amp; Murphy, J. A. (1984) A revision of the spider genera Trachyzelotes and Urozelotes (Araneae, Gnaphosidae). American Museum Novitates, 2792, 1 - 30." type="journal article" year="1984">Platnick &amp; Murphy 1984: 17</bibRefCitation>
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<emphasis id="93FEEA97FFE45D4499A6FC031BC5A72F" bold="true" box="[151,387,1880,1906]" pageId="5" pageNumber="306">Material examined:</emphasis>
EL GUERRARA: El Amied, pitfall traps,
<specimenCount id="B78CFD0CFFE45D449A53FC0319C8A72F" box="[866,910,1880,1906]" count="2" pageId="5" pageNumber="306" type="male">2 ♂</specimenCount>
,
<date id="D5341045FFE45D449AABFC031E4DA72F" box="[922,1035,1880,1906]" pageId="5" pageNumber="306" value="2020-04-01">1.IV.2020</date>
.
</paragraph>
<paragraph id="A1353685FFE45D4499F6FC271927A7E7" blockId="5.[151,1273,1880,1978]" pageId="5" pageNumber="306">
<emphasis id="93FEEA97FFE45D4499F6FC271853A7CB" bold="true" box="[199,533,1916,1942]" pageId="5" pageNumber="306">
Previous records in
<collectingCountry id="D99D7615FFE45D449882FC271856A7CB" box="[435,528,1916,1942]" name="Algeria" pageId="5" pageNumber="306">Algeria</collectingCountry>
:
</emphasis>
Known from
<collectingRegion id="634EF867FFE45D449B82FC2718BBA7CB" box="[691,765,1916,1942]" country="Algeria" name="M'Sila" pageId="5" pageNumber="306">Msila</collectingRegion>
,
<collectingRegion id="634EF867FFE45D449A38FC27191EA7CB" box="[777,856,1916,1942]" country="Algeria" name="Naama" pageId="5" pageNumber="306">Naama</collectingRegion>
and
<collectingRegion id="634EF867FFE45D449ABFFC271980A7CB" box="[910,966,1916,1942]" country="Algeria" name="Oran" pageId="5" pageNumber="306">Oran</collectingRegion>
(
<bibRefCitation id="C51B4B74FFE45D449AE4FC271EB7A7CB" author="Platnick, N. I. &amp; Murphy, J. A." box="[981,1265,1916,1942]" pageId="5" pageNumber="306" pagination="1 - 30" refId="ref22383" refString="Platnick, N. I. &amp; Murphy, J. A. (1984) A revision of the spider genera Trachyzelotes and Urozelotes (Araneae, Gnaphosidae). American Museum Novitates, 2792, 1 - 30." type="journal article" year="1984">Platnick &amp; Murphy 1984</bibRefCitation>
)
<emphasis id="93FEEA97FFE45D4499F6FCFB1B25A7E7" bold="true" box="[199,355,1952,1978]" pageId="5" pageNumber="306">Distribution:</emphasis>
Mediterranean (
<bibRefCitation id="C51B4B74FFE45D449B2AFCFB1913A7E7" author="World Spider Catalog" box="[539,853,1952,1978]" pageId="5" pageNumber="306" refId="ref24111" refString="World Spider Catalog (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern, Bern. Available from: https: // wsc. nmbe. ch / (accessed 1 June 2024) https: // doi. org / 10.24436 / 2" type="book" year="2024">World Spider Catalog 2024</bibRefCitation>
).
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View file

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<emphasis id="93FEEA97FFE55D4599A6FDD21BCFA6F9" bold="true" box="[151,393,1673,1700]" italics="true" pageId="4" pageNumber="305">Drassodes lutescens</emphasis>
(C. L. Koch, 1839)
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<emphasis id="93FEEA97FFE55D4599A6FD951B20A6BB" box="[151,358,1742,1766]" italics="true" pageId="4" pageNumber="305">Drassodes lutescens</emphasis>
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Abrous-Kherbouche
<emphasis id="93FEEA97FFE55D459C58FD951FDBA6BB" box="[1385,1437,1742,1766]" italics="true" pageId="4" pageNumber="305">et al.</emphasis>
1997: 79
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<bibRefCitation id="C51B4B74FFE55D459802FDB718C9A759" author="Brague-Bouragba, N. &amp; Brague, A. &amp; Dellouli, S. &amp; Lieutier, F." box="[307,655,1772,1796]" pageId="4" pageNumber="305" pagination="923 - 939" refId="ref20102" refString="Brague-Bouragba, N., Brague, A., Dellouli, S. &amp; Lieutier, F. (2007) Comparaison des peuplements de Coleopteres et d'Araignees en zone reboisee et en zone steppique dans une region presaharienne d'Algerie. Comptes Rendus Biologies, 330 (2007), 923 - 939. https: // doi. org / 10.1016 / j. crvi. 2007.09.004" type="journal article" year="2007">
Brague-Bouragba
<emphasis id="93FEEA97FFE55D4598DEFDB61863A759" box="[495,549,1772,1796]" italics="true" pageId="4" pageNumber="305">et al.</emphasis>
2007:925
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Mansouri
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2019: 4
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Bouragba
<emphasis id="93FEEA97FFE55D459D3CFDB61E05A759" box="[1037,1091,1772,1796]" italics="true" pageId="4" pageNumber="305">et al.</emphasis>
2020: 29
</bibRefCitation>
</treatmentCitation>
;
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<bibRefCitation id="C51B4B74FFE55D459D87FDB71BC7A77E" author="Chaib, S. &amp; Kherbouche-Abrous, O." pageId="4" pageNumber="305" pagination="129 - 138" refId="ref20171" refString="Chaib, S. &amp; Kherbouche-Abrous, O. (2021) Ecologie et phenologie des Gnaphosidae (Araneae, Arachnida) dans la region de Tikjda (Parc National du Djurdjura, Algerie). Bulletin de la Societe Zoologique de France, 146 (4), 129 - 138." type="journal article" year="2021">Chaib &amp; Kherbouche-Abrous 2021: 132</bibRefCitation>
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.
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</paragraph>
<paragraph id="A1353685FFE55D4599A6FC711967A71F" blockId="4.[151,1437,1741,1858]" box="[151,801,1833,1858]" pageId="4" pageNumber="305">
<treatmentCitationGroup id="819A11ABFFE55D4599A6FC711967A71F" box="[151,801,1833,1858]" pageId="4" pageNumber="305">
<taxonomicName id="668A4D06FFE55D4599A6FC711B2AA71F" ID-CoL="37LQ5" authorityName="Denis" authorityYear="1937" box="[151,364,1834,1858]" class="Arachnida" family="Gnaphosidae" genus="Drassodes" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="persimilis">
<emphasis id="93FEEA97FFE55D4599A6FC711B2AA71F" box="[151,364,1834,1858]" italics="true" pageId="4" pageNumber="305">Drassodes persimilis</emphasis>
</taxonomicName>
;
<treatmentCitation id="202B1094FFE55D45984BFC711878A71C" author="Denis, J." box="[378,574,1833,1858]" page="1035" pageId="4" pageNumber="305" year="1937">
<bibRefCitation id="C51B4B74FFE55D45984BFC711878A71C" author="Denis, J." box="[378,574,1833,1858]" pageId="4" pageNumber="305" pagination="1027 - 1060" refId="ref20639" refString="Denis, J. (1937 b) On a collection of spiders from Algeria. Proceedings of the Zoological Society of London, 106 (4), 1027 - 1060. https: // doi. org / 10.1111 / j. 1469 - 7998.1936. tb 06301. x" type="journal article" year="1937">Denis 1937b: 1035</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="202B1094FFE55D459B79FC71195BA71F" author="Di Franco, F." box="[584,797,1834,1858]" page="201" pageId="4" pageNumber="305" year="1994">
<bibRefCitation id="C51B4B74FFE55D459B79FC71195BA71F" author="Di Franco, F." box="[584,797,1834,1858]" pageId="4" pageNumber="305" pagination="193 - 211" refId="ref20785" refString="Di Franco, F. (1994) Contributo alla conoscenza degli Gnaphosidae (Arachnida, Araneae) del Maghreb. Animalia, 19, 193 - 211." type="journal article" year="1994">Di Franco 1994: 201</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="E990650EFFE55D4499A6FC341B8AA088" lastPageId="5" lastPageNumber="306" pageId="4" pageNumber="305" type="materials_examined">
<paragraph id="A1353685FFE55D4599A6FC341B96A7F2" blockId="4.[151,1436,1903,2038]" pageId="4" pageNumber="305">
<emphasis id="93FEEA97FFE55D4599A6FC341BC2A7D4" bold="true" box="[151,388,1903,1929]" pageId="4" pageNumber="305">Material examined:</emphasis>
EL GUERRARA: Aghzou, Pitfall traps,
<specimenCount id="B78CFD0CFFE55D459A66FC2B19C2A7D7" box="[855,900,1904,1930]" count="1" pageId="4" pageNumber="305" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFE55D459AA0FC2B1E4FA7D7" box="[913,1033,1904,1930]" pageId="4" pageNumber="305" value="2020-02-22">22.II.2020</date>
. MANSOURA: Mansoura Kedima, pitfall traps,
<specimenCount id="B78CFD0CFFE55D459817FCCF1B14A7F3" box="[294,338,1940,1966]" count="2" pageId="4" pageNumber="305" type="male">2 ♂</specimenCount>
,
<date id="D5341045FFE55D45986FFCCF1B8AA7F2" box="[350,460,1940,1967]" pageId="4" pageNumber="305" value="2015-01-10">10.I.2015</date>
.
</paragraph>
<paragraph id="A1353685FFE55D4499F6FCEC1B8AA088" blockId="4.[151,1436,1903,2038]" lastBlockId="5.[151,1436,151,285]" lastPageId="5" lastPageNumber="306" pageId="4" pageNumber="305">
<emphasis id="93FEEA97FFE55D4599F6FCEC185AA78C" bold="true" box="[199,540,1975,2002]" pageId="4" pageNumber="305">
Previous records in
<collectingCountry id="D99D7615FFE55D45988BFCEC1851A78C" box="[442,535,1975,2001]" name="Algeria" pageId="4" pageNumber="305">Algeria</collectingCountry>
:
</emphasis>
<collectingRegion id="634EF867FFE55D459B15FCE31838A78F" box="[548,638,1976,2002]" country="Algeria" name="Annaba" pageId="4" pageNumber="305">Annaba</collectingRegion>
,
<collectingRegion id="634EF867FFE55D459BBDFCE31896A78F" box="[652,720,1976,2002]" country="Algeria" name="Batna" pageId="4" pageNumber="305">Batna</collectingRegion>
,
<collectingRegion id="634EF867FFE55D459BECFCE31961A78F" box="[733,807,1976,2002]" country="Algeria" name="Bejaia" pageId="4" pageNumber="305">Béjaïa</collectingRegion>
,
<collectingRegion id="634EF867FFE55D459A04FCE31932A78F" box="[821,884,1976,2002]" country="Algeria" name="Blida" pageId="4" pageNumber="305">Blida</collectingRegion>
,
<collectingRegion id="634EF867FFE55D459AB3FCE31997A78F" box="[898,977,1976,2002]" country="Algeria" name="Bouira" pageId="4" pageNumber="305">Bouira</collectingRegion>
,
<collectingRegion id="634EF867FFE55D459AEEFCE31E2EA78F" box="[991,1128,1976,2002]" country="Algeria" name="Constantine" pageId="4" pageNumber="305">Constantine</collectingRegion>
,
<collectingRegion id="634EF867FFE55D459D44FCE31EF8A78F" box="[1141,1214,1976,2002]" country="Algeria" name="Djelfa" pageId="4" pageNumber="305">Djelfa</collectingRegion>
,
<collectingRegion id="634EF867FFE55D459DFAFCE31F58A78F" box="[1227,1310,1976,2002]" country="Algeria" name="El Tarf" pageId="4" pageNumber="305">El Tarf</collectingRegion>
,
<collectingRegion id="634EF867FFE55D459C1AFCE31FDEA78F" box="[1323,1432,1976,2002]" country="Algeria" name="Laghouat" pageId="4" pageNumber="305">Laghouat</collectingRegion>
,
<collectingRegion id="634EF867FFE55D4599A6FC871AA0A7AB" box="[151,230,2012,2038]" country="Algeria" name="Medea" pageId="4" pageNumber="305">Médéa</collectingRegion>
,
<collectingRegion id="634EF867FFE55D4599C7FC871B68A7AB" box="[246,302,2012,2038]" country="Algeria" name="Mila" pageId="4" pageNumber="305">Mila</collectingRegion>
,
<collectingRegion id="634EF867FFE55D45980CFC871B33A7AB" box="[317,373,2012,2038]" country="Algeria" name="Setif" pageId="4" pageNumber="305">Sétif</collectingRegion>
,
<collectingRegion id="634EF867FFE55D4598B5FC87184FA7AB" box="[388,521,2012,2038]" country="Algeria" name="Tizi Ouzou" pageId="4" pageNumber="305">Tizi Ouzou</collectingRegion>
,
<collectingRegion id="634EF867FFE55D459B28FC87183DA7AB" box="[537,635,2012,2038]" country="Algeria" name="Tlemcen" pageId="4" pageNumber="305">Tlemcen</collectingRegion>
(
<bibRefCitation id="C51B4B74FFE55D459BBFFC871959A7AB" author="Simon, E." box="[654,799,2012,2038]" pageId="4" pageNumber="305" pagination="82 - 87" refId="ref23337" refString="Simon, E. (1899) Liste des arachnides recueillis en Algerie par M. P. Lesne et descrition d'une espece nouvelle. Bulletin du Museum d'Histoire Naturelle Paris, 5, 82 - 87." type="journal article" year="1899">Simon 1899</bibRefCitation>
;
<bibRefCitation id="C51B4B74FFE55D459A01FC8719F9A7AB" author="Simon, E." box="[816,959,2012,2038]" pageId="4" pageNumber="305" pagination="414 - 419" refId="ref23456" refString="Simon, E. (1912) Araneae. In: Jeannel, R. &amp; Peyerimhof, P. de (eds.). Recoltes entomologiques dans les Beni-Snassen (Maroc orientale). Annales de la Societe Entomologique de France, 80 (3), 414 - 419." type="journal article" year="1912">Simon 1912</bibRefCitation>
;
<bibRefCitation id="C51B4B74FFE55D459AE1FC871E20A7AB" author="Denis, J." box="[976,1126,2012,2038]" pageId="4" pageNumber="305" pagination="1027 - 1060" refId="ref20639" refString="Denis, J. (1937 b) On a collection of spiders from Algeria. Proceedings of the Zoological Society of London, 106 (4), 1027 - 1060. https: // doi. org / 10.1111 / j. 1469 - 7998.1936. tb 06301. x" type="journal article" year="1937">Denis 1937b</bibRefCitation>
;
<bibRefCitation id="C51B4B74FFE55D459D47FC871F74A7AB" author="Di Franco, F." box="[1142,1330,2012,2038]" pageId="4" pageNumber="305" pagination="193 - 211" refId="ref20785" refString="Di Franco, F. (1994) Contributo alla conoscenza degli Gnaphosidae (Arachnida, Araneae) del Maghreb. Animalia, 19, 193 - 211." type="journal article" year="1994">Di Franco 1994</bibRefCitation>
;
<bibRefCitation id="C51B4B74FFE55D449C71FC871BE8A0EC" author="Abrous-Kherbouche, O. &amp; Jocque, R. &amp; Maelfait, J. - P." lastPageId="5" lastPageNumber="306" pageId="4" pageNumber="305" pagination="71 - 90" refId="ref18469" refString="Abrous-Kherbouche, O., Jocque, R. &amp; Maelfait, J. - P. (1997) Les effets du paturage intensif sur l'araneofaune dans la region de Tala-Guilef (Parc National du Djurdjura, Algerie). Annales de la Societe Entomologique de Belgique, 133, 71 - 90." type="journal article" year="1997">
Abrous-Kherbouche
<emphasis id="93FEEA97FFE45D44981AFBCC1B2EA0EC" box="[299,360,151,177]" italics="true" pageId="5" pageNumber="306">et al.</emphasis>
1997
</bibRefCitation>
;
<bibRefCitation id="C51B4B74FFE45D44988FFBCC1955A0EC" author="Brague-Bouragba, N. &amp; Brague, A. &amp; Dellouli, S. &amp; Lieutier, F." box="[446,787,151,177]" pageId="5" pageNumber="306" pagination="923 - 939" refId="ref20102" refString="Brague-Bouragba, N., Brague, A., Dellouli, S. &amp; Lieutier, F. (2007) Comparaison des peuplements de Coleopteres et d'Araignees en zone reboisee et en zone steppique dans une region presaharienne d'Algerie. Comptes Rendus Biologies, 330 (2007), 923 - 939. https: // doi. org / 10.1016 / j. crvi. 2007.09.004" type="journal article" year="2007">
Brague-Bouragba
<emphasis id="93FEEA97FFE45D449BA0FBCC188BA0EC" box="[657,717,151,177]" italics="true" pageId="5" pageNumber="306">et al.</emphasis>
2007
</bibRefCitation>
;
<bibRefCitation id="C51B4B74FFE45D449A15FBCC1E5BA0EC" author="Mansouri, H. &amp; Ould Rouis, S. &amp; Kherbouche-Abrous, O. &amp; Ould Rouis, A. &amp; Beladjal, L." box="[804,1053,151,177]" pageId="5" pageNumber="306" pagination="1 - 13" refId="ref21811" refString="Mansouri, H., Ould Rouis, S., Kherbouche-Abrous, O., Ould Rouis, A. &amp; Beladjal, L. (2019) Effects of anthropogenic factors on spider communities (Arthropoda: Araneae) in Chrea National park (Blida, Algeria). African Journal of Ecology, 2019 (00), 1 - 13. https: // doi. org / 10.1111 / aje. 12701" type="journal article" year="2019">
Mansouri
<emphasis id="93FEEA97FFE45D449AABFBCC1991A0EC" box="[922,983,151,177]" italics="true" pageId="5" pageNumber="306">et al.</emphasis>
2019
</bibRefCitation>
;
<bibRefCitation id="C51B4B74FFE45D449D1CFBCC1F6FA0EC" author="Bouragba, N. &amp; Bouragba, M. &amp; Djouklafi, A. &amp; Amraoui, S. &amp; Brague, A. &amp; Beladjal, L." box="[1069,1321,151,177]" pageId="5" pageNumber="306" pagination="25 - 34" refId="ref19910" refString="Bouragba, N., Bouragba, M., Djouklafi, A., Amraoui, S., Brague, A. &amp; Beladjal, L. (2020) Spider (Araneae) and beetle (Coleoptera) communities structure in the North African streamside zone (Zahrez Gharbi, Djelfa, Algeria) after reforestation. African Entomology, 28 (1), 25 - 34." type="journal article" year="2020">
Bouragba
<emphasis id="93FEEA97FFE45D449D94FBCC1EA4A0EC" box="[1189,1250,151,177]" italics="true" pageId="5" pageNumber="306">et al.</emphasis>
2020
</bibRefCitation>
;
<bibRefCitation id="C51B4B74FFE45D449C09FBCC1BF9A088" author="Chaib, S. &amp; Kherbouche-Abrous, O." pageId="5" pageNumber="306" pagination="129 - 138" refId="ref20171" refString="Chaib, S. &amp; Kherbouche-Abrous, O. (2021) Ecologie et phenologie des Gnaphosidae (Araneae, Arachnida) dans la region de Tikjda (Parc National du Djurdjura, Algerie). Bulletin de la Societe Zoologique de France, 146 (4), 129 - 138." type="journal article" year="2021">Chaib &amp; Kherbouche-Abrous 2021</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFE45D4499F6FB851BC8A140" pageId="5" pageNumber="306" type="distribution">
<paragraph id="A1353685FFE45D4499F6FB851BC8A140" blockId="5.[151,1436,151,285]" pageId="5" pageNumber="306">
<emphasis id="93FEEA97FFE45D4499F6FB851B25A0A5" bold="true" box="[199,355,222,248]" pageId="5" pageNumber="306">Distribution:</emphasis>
Mediterranean,
<collectingCountry id="D99D7615FFE45D449B29FB841833A0A4" box="[536,629,223,249]" name="Ukraine" pageId="5" pageNumber="306">Ukraine</collectingCountry>
, Caucasus,
<collectingCountry id="D99D7615FFE45D449BC4FB841906A0A4" box="[757,832,223,249]" name="Russia" pageId="5" pageNumber="306">Russia</collectingCountry>
(Europe) to Central Asia,
<collectingCountry id="D99D7615FFE45D449D51FB841ED6A0A4" box="[1120,1168,223,249]" name="Iran" pageId="5" pageNumber="306">Iran</collectingCountry>
,
<collectingCountry id="D99D7615FFE45D449DABFB841EBAA0A4" box="[1178,1276,223,249]" name="Pakistan" pageId="5" pageNumber="306">Pakistan</collectingCountry>
,
<collectingCountry id="D99D7615FFE45D449C34FB841F0EA0A4" box="[1285,1352,223,249]" name="Nepal" pageId="5" pageNumber="306">Nepal</collectingCountry>
(
<bibRefCitation id="C51B4B74FFE45D449C64FB841BC4A140" author="World Spider Catalog" pageId="5" pageNumber="306" refId="ref24111" refString="World Spider Catalog (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern, Bern. Available from: https: // wsc. nmbe. ch / (accessed 1 June 2024) https: // doi. org / 10.24436 / 2" type="book" year="2024">World Spider Catalog 2024</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<paragraph id="A1353685FFE65D4699A6F903185BA22F" blockId="7.[151,541,600,627]" box="[151,541,600,627]" pageId="7" pageNumber="308">
<heading id="FA7D81E9FFE65D4699A6F903185BA22F" bold="true" box="[151,541,600,627]" fontSize="11" level="1" pageId="7" pageNumber="308" reason="1">
<taxonomicName id="668A4D06FFE65D4699A6F903185BA22F" ID-CoL="47P2Y" authority="Dalmas, 1921" authorityName="Dalmas" authorityYear="1921" box="[151,541,600,627]" class="Arachnida" family="Gnaphosidae" genus="Nomisia" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="castanea">
<emphasis id="93FEEA97FFE65D4699A6F903185BA22F" bold="true" box="[151,541,600,627]" pageId="7" pageNumber="308">
<emphasis id="93FEEA97FFE65D4699A6F9031B28A22E" bold="true" box="[151,366,600,627]" italics="true" pageId="7" pageNumber="308">Nomisia castanea</emphasis>
<bibRefCitation id="C51B4B74FFE65D469844F902185BA22F" author="Dalmas, R. de" box="[373,541,600,627]" pageId="7" pageNumber="308" pagination="233 - 328" refId="ref20501" refString="Dalmas, R. de (1921) Monographie des araignees de la section des Pterotricha (Aran. Gnaphosidae). Annales de la Societe Entomologique de France, 89, 233 - 328." type="journal article" year="1921">Dalmas, 1921</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFE65D4699A6F9C51940A289" pageId="7" pageNumber="308" type="reference_group">
<paragraph id="A1353685FFE65D4699A6F9C51940A289" blockId="7.[151,1436,669,724]" pageId="7" pageNumber="308">
<treatmentCitationGroup id="819A11ABFFE65D4699A6F9C51940A289" pageId="7" pageNumber="308">
<taxonomicName id="668A4D06FFE65D4699A6F9C51B0CA2E8" ID-CoL="47P2Y" authorityName="Dalmas" authorityYear="1921" box="[151,330,670,693]" class="Arachnida" family="Gnaphosidae" genus="Nomisia" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="castanea">
<emphasis id="93FEEA97FFE65D4699A6F9C51B0CA2E8" box="[151,330,670,693]" italics="true" pageId="7" pageNumber="308">Nomisia castanea</emphasis>
</taxonomicName>
;
<treatmentCitation id="202B1094FFE65D469867F9C61856A2E8" author="Dalmas, R. de" box="[342,528,669,693]" page="284" pageId="7" pageNumber="308" year="1921">
<bibRefCitation id="C51B4B74FFE65D469867F9C61856A2E8" author="Dalmas, R. de" box="[342,528,669,693]" pageId="7" pageNumber="308" pagination="233 - 328" refId="ref20501" refString="Dalmas, R. de (1921) Monographie des araignees de la section des Pterotricha (Aran. Gnaphosidae). Annales de la Societe Entomologique de France, 89, 233 - 328." type="journal article" year="1921">Dalmas 1921: 284</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="202B1094FFE65D469B28F9C618AFA2E8" author="Di Franco, F." box="[537,745,669,693]" page="196" pageId="7" pageNumber="308" year="1994">
<bibRefCitation id="C51B4B74FFE65D469B28F9C618AFA2E8" author="Di Franco, F." box="[537,745,669,693]" pageId="7" pageNumber="308" pagination="193 - 211" refId="ref20785" refString="Di Franco, F. (1994) Contributo alla conoscenza degli Gnaphosidae (Arachnida, Araneae) del Maghreb. Animalia, 19, 193 - 211." type="journal article" year="1994">Di Franco 1994: 196</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="202B1094FFE65D469BC3F9C61E02A2E8" author="Brague-Bouragba, N. &amp; Brague, A. &amp; Dellouli, S. &amp; Lieutier, F." box="[754,1092,669,693]" page="937" pageId="7" pageNumber="308" year="2007">
<bibRefCitation id="C51B4B74FFE65D469BC3F9C61E02A2E8" author="Brague-Bouragba, N. &amp; Brague, A. &amp; Dellouli, S. &amp; Lieutier, F." box="[754,1092,669,693]" pageId="7" pageNumber="308" pagination="923 - 939" refId="ref20102" refString="Brague-Bouragba, N., Brague, A., Dellouli, S. &amp; Lieutier, F. (2007) Comparaison des peuplements de Coleopteres et d'Araignees en zone reboisee et en zone steppique dans une region presaharienne d'Algerie. Comptes Rendus Biologies, 330 (2007), 923 - 939. https: // doi. org / 10.1016 / j. crvi. 2007.09.004" type="journal article" year="2007">
Brague-Bouragba
<emphasis id="93FEEA97FFE65D469A9AF9C5199AA2E8" box="[939,988,669,693]" italics="true" pageId="7" pageNumber="308">et al.</emphasis>
2007:937
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="202B1094FFE65D469D7DF9C61F77A2E8" author="Mansouri, H. &amp; Ould Rouis, S. &amp; Kherbouche-Abrous, O. &amp; Ould Rouis, A. &amp; Beladjal, L." box="[1100,1329,669,693]" page="5" pageId="7" pageNumber="308" year="2019">
<bibRefCitation id="C51B4B74FFE65D469D7DF9C61F77A2E8" author="Mansouri, H. &amp; Ould Rouis, S. &amp; Kherbouche-Abrous, O. &amp; Ould Rouis, A. &amp; Beladjal, L." box="[1100,1329,669,693]" pageId="7" pageNumber="308" pagination="1 - 13" refId="ref21811" refString="Mansouri, H., Ould Rouis, S., Kherbouche-Abrous, O., Ould Rouis, A. &amp; Beladjal, L. (2019) Effects of anthropogenic factors on spider communities (Arthropoda: Araneae) in Chrea National park (Blida, Algeria). African Journal of Ecology, 2019 (00), 1 - 13. https: // doi. org / 10.1111 / aje. 12701" type="journal article" year="2019">
Mansouri
<emphasis id="93FEEA97FFE65D469D83F9C51EA5A2E8" box="[1202,1251,669,693]" italics="true" pageId="7" pageNumber="308">et al.</emphasis>
2019:5
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="202B1094FFE65D469C0BF9C61B1AA289" author="Bouragba, N. &amp; Bouragba, M. &amp; Djouklafi, A. &amp; Amraoui, S. &amp; Brague, A. &amp; Beladjal, L." page="29" pageId="7" pageNumber="308" year="2020">
<bibRefCitation id="C51B4B74FFE65D469C0BF9C61B1AA289" author="Bouragba, N. &amp; Bouragba, M. &amp; Djouklafi, A. &amp; Amraoui, S. &amp; Brague, A. &amp; Beladjal, L." pageId="7" pageNumber="308" pagination="25 - 34" refId="ref19910" refString="Bouragba, N., Bouragba, M., Djouklafi, A., Amraoui, S., Brague, A. &amp; Beladjal, L. (2020) Spider (Araneae) and beetle (Coleoptera) communities structure in the North African streamside zone (Zahrez Gharbi, Djelfa, Algeria) after reforestation. African Entomology, 28 (1), 25 - 34." type="journal article" year="2020">
Bouragba
<emphasis id="93FEEA97FFE65D4699F6F9E71ABCA289" box="[199,250,700,724]" italics="true" pageId="7" pageNumber="308">et al.</emphasis>
2020: 29
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="202B1094FFE65D469856F9E71944A289" author="Chaib, S. &amp; Kherbouche-Abrous, O." box="[359,770,700,724]" page="132" pageId="7" pageNumber="308" year="2021">
<bibRefCitation id="C51B4B74FFE65D469856F9E71944A289" author="Chaib, S. &amp; Kherbouche-Abrous, O." box="[359,770,700,724]" pageId="7" pageNumber="308" pagination="129 - 138" refId="ref20171" refString="Chaib, S. &amp; Kherbouche-Abrous, O. (2021) Ecologie et phenologie des Gnaphosidae (Araneae, Arachnida) dans la region de Tikjda (Parc National du Djurdjura, Algerie). Bulletin de la Societe Zoologique de France, 146 (4), 129 - 138." type="journal article" year="2021">Chaib &amp; Kherbouche-Abrous 2021: 132</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFE65D4699A6F85A183FA565" pageId="7" pageNumber="308" type="materials_examined">
<paragraph id="A1353685FFE65D4699A6F85A1F46A490" blockId="7.[151,1437,769,1372]" pageId="7" pageNumber="308">
<emphasis id="93FEEA97FFE65D4699A6F85A1BC5A346" bold="true" box="[151,387,769,795]" pageId="7" pageNumber="308">Material examined:</emphasis>
BERRIANE: Balouh, hand collecting,
<specimenCount id="B78CFD0CFFE65D469A70F859192DA341" box="[833,875,770,796]" count="3" pageId="7" pageNumber="308" type="female">3 ♀</specimenCount>
,
<date id="D5341045FFE65D469A4BF85919A5A341" box="[890,995,770,796]" pageId="7" pageNumber="308" value="2021-02-02">2.II.2021</date>
;
<specimenCount id="B78CFD0CFFE65D469AC1F8591E5DA341" box="[1008,1051,770,796]" count="2" pageId="7" pageNumber="308" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFE65D469D18F8591EE7A341" box="[1065,1185,770,796]" pageId="7" pageNumber="308" value="2021-02-12">12.II.2021</date>
;
<specimenCount id="B78CFD0CFFE65D469D9CF8591E9EA341" box="[1197,1240,770,796]" count="4" pageId="7" pageNumber="308" type="female">4 ♀</specimenCount>
,
<date id="D5341045FFE65D469DD6F8591F18A341" box="[1255,1374,770,796]" pageId="7" pageNumber="308" value="2021-02-18">18.II.2021</date>
;
<specimenCount id="B78CFD0CFFE65D469C5AF8591FD3A341" box="[1387,1429,770,796]" count="2" pageId="7" pageNumber="308" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFE65D4699A6F87D1B4CA31C" box="[151,266,806,833]" pageId="7" pageNumber="308" value="2021-03-05">5.III.2021</date>
;
<specimenCount id="B78CFD0CFFE65D469827F87D1B06A31D" box="[278,320,806,832]" count="1" pageId="7" pageNumber="308" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFE65D46987EF87D1B88A31D" box="[335,462,806,832]" pageId="7" pageNumber="308" value="2021-04-10">10.IV.2021</date>
;
<specimenCount id="B78CFD0CFFE65D4698EBF87D1842A31D" box="[474,516,806,832]" count="2" pageId="7" pageNumber="308" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFE65D469B23F87D18D7A31D" box="[530,657,806,832]" pageId="7" pageNumber="308" value="2021-04-17">17.IV.2021</date>
;
<specimenCount id="B78CFD0CFFE65D469BACF87D1881A31D" box="[669,711,806,832]" count="3" pageId="7" pageNumber="308" type="female">3 ♀</specimenCount>
,
<date id="D5341045FFE65D469BE4F87D1912A31D" box="[725,852,806,832]" pageId="7" pageNumber="308" value="2021-04-30">30.IV.2021</date>
, pitfall traps,
<specimenCount id="B78CFD0CFFE65D469ADFF87D1E5EA31D" box="[1006,1048,806,832]" count="1" pageId="7" pageNumber="308" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFE65D469D16F87D1EDFA31C" box="[1063,1177,806,833]" pageId="7" pageNumber="308" value="2021-03-05">5.III.2021</date>
;
<specimenCount id="B78CFD0CFFE65D469D97F87D1E96A31D" box="[1190,1232,806,832]" count="1" pageId="7" pageNumber="308" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFE65D469DEFF87D1F26A31D" box="[1246,1376,806,832]" pageId="7" pageNumber="308" value="2021-03-12">12.III.2021</date>
;
<specimenCount id="B78CFD0CFFE65D469C5AF87D1FD3A31D" box="[1387,1429,806,832]" count="1" pageId="7" pageNumber="308" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFE65D4699A6F8111B5EA339" box="[151,280,842,868]" pageId="7" pageNumber="308" value="2021-03-19">19.III.2021</date>
;
<specimenCount id="B78CFD0CFFE65D469814F8111B16A339" box="[293,336,842,868]" count="2" pageId="7" pageNumber="308" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFE65D46986EF8111BA6A338" box="[351,480,842,869]" pageId="7" pageNumber="308" value="2021-03-25">25.III.2021</date>
;
<specimenCount id="B78CFD0CFFE65D4698DCF811185EA339" box="[493,536,842,868]" count="1" pageId="7" pageNumber="308" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFE65D469B16F81118E0A339" box="[551,678,842,868]" pageId="7" pageNumber="308" value="2021-04-17">17.IV.2021</date>
.
<materialsCitation id="11E23CD8FFE65D469B83F81119BBA339" box="[690,1021,842,868]" location="Sidi Mbarek" pageId="7" pageNumber="308" specimenCount="1">
<location id="A455605EFFE65D469B83F8111907A339" LSID="urn:lsid:plazi:treatment:29238793FFE65D4699A6F9031986A501:A455605EFFE65D469B83F8111907A339" box="[690,833,842,868]" name="Sidi Mbarek" pageId="7" pageNumber="308">Sidi Mbarek</location>
, hand collecting
</materialsCitation>
,
<specimenCount id="B78CFD0CFFE65D469D38F8111E72A339" box="[1033,1076,842,868]" count="1" pageId="7" pageNumber="308" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFE65D469D72F8111EEBA338" box="[1091,1197,842,869]" pageId="7" pageNumber="308" value="2021-02-05">5.II.2021</date>
;
<specimenCount id="B78CFD0CFFE65D469D8BF8111EA3A339" box="[1210,1253,842,868]" count="1" pageId="7" pageNumber="308" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFE65D469DC5F8111F18A339" box="[1268,1374,842,868]" pageId="7" pageNumber="308" value="2021-02-06">6.II.2021</date>
;
<specimenCount id="B78CFD0CFFE65D469C5BF8111FD3A339" box="[1386,1429,842,868]" count="1" pageId="7" pageNumber="308" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFE65D4699A6F8351B47A3D5" box="[151,257,878,904]" pageId="7" pageNumber="308" value="2021-02-08">8.II.2021</date>
;
<specimenCount id="B78CFD0CFFE65D46983DF8351B73A3D5" box="[268,309,878,904]" count="2" pageId="7" pageNumber="308" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFE65D469873F8351BFEA3D5" box="[322,440,878,904]" pageId="7" pageNumber="308" value="2021-02-11">11.II.2021</date>
;
<specimenCount id="B78CFD0CFFE65D4698F2F8351BABA3D5" box="[451,493,878,904]" count="2" pageId="7" pageNumber="308" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFE65D4698CBF8351837A3D5" box="[506,625,878,904]" pageId="7" pageNumber="308" value="2021-02-18">18.II.2021</date>
;
<specimenCount id="B78CFD0CFFE65D469B4DF83518E3A3D5" box="[636,677,878,904]" count="1" pageId="7" pageNumber="308" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFE65D469B83F835196CA3D4" box="[690,810,878,905]" pageId="7" pageNumber="308" value="2021-02-25">25.II.2021</date>
;
<specimenCount id="B78CFD0CFFE65D469A04F8351918A3D5" box="[821,862,878,904]" count="1" pageId="7" pageNumber="308" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFE65D469A5AF8351998A3D5" box="[875,990,878,904]" pageId="7" pageNumber="308" value="2021-03-04">4.III.2021</date>
;
<specimenCount id="B78CFD0CFFE65D469AD8F8351E54A3D5" box="[1001,1042,878,904]" count="1" pageId="7" pageNumber="308" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFE65D469D2EF8351ED9A3D5" box="[1055,1183,878,904]" pageId="7" pageNumber="308" value="2021-03-11">11.III.2021</date>
;
<specimenCount id="B78CFD0CFFE65D469D9BF8351E95A3D5" box="[1194,1235,878,904]" count="1" pageId="7" pageNumber="308" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFE65D469DD1F8351F27A3D4" box="[1248,1377,878,905]" pageId="7" pageNumber="308" value="2021-03-25">25.III.2021</date>
;
<specimenCount id="B78CFD0CFFE65D469C5DF8351FD3A3D5" box="[1388,1429,878,904]" count="3" pageId="7" pageNumber="308" type="female">3 ♀</specimenCount>
,
<date id="D5341045FFE65D4699A6F8C91B50A3F1" box="[151,278,914,940]" pageId="7" pageNumber="308" value="2021-04-23">23.IV.2021</date>
;
<specimenCount id="B78CFD0CFFE65D46982EF8C91B00A3F1" box="[287,326,914,940]" count="1" pageId="7" pageNumber="308" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFE65D469860F8C91B96A3F1" box="[337,464,914,940]" pageId="7" pageNumber="308" value="2021-04-29">29.IV.2021</date>
.
<materialsCitation id="11E23CD8FFE65D4698E9F8C91E5FA3F1" box="[472,1049,914,940]" location="Grazil" pageId="7" pageNumber="308" specimenCount="1">
DHAYET BENDHAHOUA:
<location id="A455605EFFE65D469A2CF8C91920A3F1" LSID="urn:lsid:plazi:treatment:29238793FFE65D4699A6F9031986A501:A455605EFFE65D469A2CF8C91920A3F1" box="[797,870,914,940]" name="Grazil" pageId="7" pageNumber="308">Grazil</location>
, hand collecting
</materialsCitation>
,
<specimenCount id="B78CFD0CFFE65D469D10F8C91E0EA3F1" box="[1057,1096,914,940]" count="7" pageId="7" pageNumber="308" type="female">7 ♀</specimenCount>
,
<date id="D5341045FFE65D469D62F8C91E97A3F0" box="[1107,1233,914,941]" pageId="7" pageNumber="308" value="2021-12-05">5.XII.2021</date>
;
<specimenCount id="B78CFD0CFFE65D469DEBF8C91F47A3F1" box="[1242,1281,914,940]" count="1" pageId="7" pageNumber="308" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFE65D469C3DF8C91FD1A3F0" box="[1292,1431,914,941]" pageId="7" pageNumber="308" value="2021-12-15">15.XII.2021</date>
;
<specimenCount id="B78CFD0CFFE65D4699A6F8ED1A84A38D" box="[151,194,950,976]" count="2" pageId="7" pageNumber="308" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFE65D4699E0F8ED1B1BA38D" box="[209,349,950,976]" pageId="7" pageNumber="308" value="2021-12-26">26.XII.2021</date>
;
<specimenCount id="B78CFD0CFFE65D46985BF8ED1BD3A38D" box="[362,405,950,976]" count="8" pageId="7" pageNumber="308" type="female">8 ♀</specimenCount>
,
<date id="D5341045FFE65D469895F8ED1842A38C" box="[420,516,950,977]" pageId="7" pageNumber="308" value="2022-01-05">5.I.2022</date>
;
<specimenCount id="B78CFD0CFFE65D469B20F8ED187AA38D" box="[529,572,950,976]" count="3" pageId="7" pageNumber="308" type="female">3 ♀</specimenCount>
,
<date id="D5341045FFE65D469B7AF8ED18FFA38C" box="[587,697,950,977]" pageId="7" pageNumber="308" value="2022-01-15">15.I.2022</date>
;
<specimenCount id="B78CFD0CFFE65D469BF7F8ED18B5A38D" box="[710,755,950,976]" count="1" pageId="7" pageNumber="308" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFE65D469A31F8ED1931A38C" box="[768,887,950,977]" pageId="7" pageNumber="308" value="2022-02-15">15.II.2022</date>
;
<specimenCount id="B78CFD0CFFE65D469AB5F8ED19F7A38D" box="[900,945,950,976]" count="1" pageId="7" pageNumber="308" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFE65D469A8FF8ED1E73A38C" box="[958,1077,950,977]" pageId="7" pageNumber="308" value="2022-02-25">25.II.2022</date>
, pitfall traps,
<specimenCount id="B78CFD0CFFE65D469DE3F8ED1EBBA38D" box="[1234,1277,950,976]" count="1" pageId="7" pageNumber="308" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFE65D469C3DF8ED1FDEA38D" box="[1292,1432,950,976]" pageId="7" pageNumber="308" value="2021-12-26">26.XII.2021</date>
;
<materialsCitation id="11E23CD8FFE65D4699A6F8811BF5A3A9" box="[151,435,986,1012]" location="Oumdjer" pageId="7" pageNumber="308" specimenCount="1">
<location id="A455605EFFE65D4699A6F8811ABAA3A9" LSID="urn:lsid:plazi:treatment:29238793FFE65D4699A6F9031986A501:A455605EFFE65D4699A6F8811ABAA3A9" box="[151,252,986,1012]" name="Oumdjer" pageId="7" pageNumber="308">Oumdjer</location>
, hand collecting
</materialsCitation>
,
<specimenCount id="B78CFD0CFFE65D46988CF8811BA0A3A9" box="[445,486,986,1012]" count="1" pageId="7" pageNumber="308" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFE65D4698C2F8811837A3A8" box="[499,625,986,1013]" pageId="7" pageNumber="308" value="2021-12-05">5.XII.2021</date>
.
<materialsCitation id="11E23CD8FFE65D469B4BF8811E93A3A9" box="[634,1237,986,1012]" collectorName="El Atteuf" location="Palm" pageId="7" pageNumber="308" specimenCount="1">
EL ATTEUF:
<location id="A455605EFFE65D469A24F8811909A3A9" LSID="urn:lsid:plazi:treatment:29238793FFE65D4699A6F9031986A501:A455605EFFE65D469A24F8811909A3A9" box="[789,847,986,1012]" name="Palm" pageId="7" pageNumber="308">Palm</location>
grove of
<collectorName id="0C7F5353FFE65D469A86F8811E59A3A9" box="[951,1055,986,1012]" pageId="7" pageNumber="308">El Atteuf</collectorName>
, hand collecting
</materialsCitation>
,
<specimenCount id="B78CFD0CFFE65D469DD1F8811F4EA3A9" box="[1248,1288,986,1012]" count="1" pageId="7" pageNumber="308" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFE65D469C24F8811FDEA3A9" box="[1301,1432,986,1012]" pageId="7" pageNumber="308" value="2014-11-20">20.XI.2014</date>
;
<specimenCount id="B78CFD0CFFE65D4699A6F8A51A86A445" box="[151,192,1022,1048]" count="2" pageId="7" pageNumber="308" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFE65D4699FFF8A51B7AA444" box="[206,316,1022,1049]" pageId="7" pageNumber="308" value="2015-01-25">25.I.2015</date>
;
<specimenCount id="B78CFD0CFFE65D469876F8A51B36A445" box="[327,368,1022,1049]" count="5" pageId="7" pageNumber="308" type="female">5 ♀</specimenCount>
,
<date id="D5341045FFE65D46984CF8A51BA1A444" box="[381,487,1022,1049]" pageId="7" pageNumber="308" value="2015-02-05">5.II.2015</date>
;
<specimenCount id="B78CFD0CFFE65D4698C3F8A5185DA445" box="[498,539,1022,1048]" count="6" pageId="7" pageNumber="308" type="female">6 ♀</specimenCount>
,
<date id="D5341045FFE65D469B19F8A518E6A444" box="[552,672,1022,1049]" pageId="7" pageNumber="308" value="2015-02-16">16.II.2015</date>
;
<specimenCount id="B78CFD0CFFE65D469B9AF8A51892A445" box="[683,724,1022,1048]" count="1" pageId="7" pageNumber="308" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFE65D469BD3F8A5191FA444" box="[738,857,1022,1049]" pageId="7" pageNumber="308" value="2015-02-25">25.II.2015</date>
;
<specimenCount id="B78CFD0CFFE65D469A55F8A519C1A444" box="[868,903,1022,1049]" count="4" pageId="7" pageNumber="308" type="female">4 ♀</specimenCount>
,
<date id="D5341045FFE65D469ABDF8A51E40A444" box="[908,1030,1022,1049]" pageId="7" pageNumber="308" value="2015-03-06">6.III.2015</date>
,
<specimenCount id="B78CFD0CFFE65D469D23F8A51E7DA445" box="[1042,1083,1022,1048]" count="3" pageId="7" pageNumber="308" type="female">3 ♀</specimenCount>
,
<date id="D5341045FFE65D469D79F8A51E8FA444" box="[1096,1225,1022,1049]" pageId="7" pageNumber="308" value="2015-03-25">25.III.2015</date>
;
<specimenCount id="B78CFD0CFFE65D469DE5F8A51F4DA445" box="[1236,1291,1022,1048]" count="13" pageId="7" pageNumber="308" type="female">13 ♀</specimenCount>
,
<date id="D5341045FFE65D469C29F8A51FDEA444" box="[1304,1432,1022,1049]" pageId="7" pageNumber="308" value="2015-04-05">05.IV.2015</date>
;
<specimenCount id="B78CFD0CFFE65D4699A6FF791A87A461" box="[151,193,1058,1084]" count="3" pageId="7" pageNumber="308" type="female">3 ♀</specimenCount>
,
<date id="D5341045FFE65D4699E1FF791B7CA460" box="[208,314,1058,1085]" pageId="7" pageNumber="308" value="2023-02-05">5.II.2023</date>
;
<specimenCount id="B78CFD0CFFE65D469874FF791B29A461" box="[325,367,1058,1084]" count="2" pageId="7" pageNumber="308" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFE65D46984FFF791BB0A460" box="[382,502,1058,1085]" pageId="7" pageNumber="308" value="2023-02-15">15.II.2023</date>
;
<specimenCount id="B78CFD0CFFE65D469B30FF79186DA461" box="[513,555,1058,1084]" count="8" pageId="7" pageNumber="308" type="female">8 ♀</specimenCount>
,
<date id="D5341045FFE65D469B0BFF7918F7A460" box="[570,689,1058,1085]" pageId="7" pageNumber="308" value="2023-02-25">25.II.2023</date>
;
<specimenCount id="B78CFD0CFFE65D469B8CFF7918A1A461" box="[701,743,1058,1084]" count="1" pageId="7" pageNumber="308" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFE65D469BC7FF79192FA460" box="[758,873,1058,1085]" pageId="7" pageNumber="308" value="2023-03-05">5.III.2023</date>
;
<specimenCount id="B78CFD0CFFE65D469A44FF7919E6A461" box="[885,928,1058,1084]" count="4" pageId="7" pageNumber="308" type="female">4 ♀</specimenCount>
;
<date id="D5341045FFE65D469A9FFF791E69A460" box="[942,1071,1058,1085]" pageId="7" pageNumber="308" value="2023-03-15">15.III.2023</date>
;
<specimenCount id="B78CFD0CFFE65D469D0AFF791E23A461" box="[1083,1125,1058,1084]" count="3" pageId="7" pageNumber="308" type="female">3 ♀</specimenCount>
,
<date id="D5341045FFE65D469D42FF791EB5A460" box="[1139,1267,1058,1085]" pageId="7" pageNumber="308" value="2023-03-25">25.III.2023</date>
, pitfall traps,
<specimenCount id="B78CFD0CFFE65D469CBFFF791AEAA43D" count="1" pageId="7" pageNumber="308" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFE65D46998BFF1D1B77A43C" box="[186,305,1094,1121]" pageId="7" pageNumber="308" value="2015-02-16">16.II.2015</date>
;
<specimenCount id="B78CFD0CFFE65D46980CFF1D1B20A43D" box="[317,358,1094,1120]" count="2" pageId="7" pageNumber="308" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFE65D469842FF1D1BB2A43C" box="[371,500,1094,1121]" pageId="7" pageNumber="308" value="2015-03-15">15.III.2015</date>
,
<specimenCount id="B78CFD0CFFE65D4698CEFF1D186EA43D" box="[511,552,1094,1120]" count="4" pageId="7" pageNumber="308" type="female">4 ♀</specimenCount>
,
<date id="D5341045FFE65D469B07FF1D18F1A43C" box="[566,695,1094,1121]" pageId="7" pageNumber="308" value="2015-03-25">25.III.2015</date>
;
<specimenCount id="B78CFD0CFFE65D469BF3FF1D18AAA43D" box="[706,748,1094,1120]" count="3" pageId="7" pageNumber="308" type="female">3 ♀</specimenCount>
,
<date id="D5341045FFE65D469BC8FF1D193EA43C" box="[761,888,1094,1121]" pageId="7" pageNumber="308" value="2015-04-05">05.IV.2015</date>
;
<specimenCount id="B78CFD0CFFE65D469AB2FF1D19EBA43D" box="[899,941,1094,1120]" count="1" pageId="7" pageNumber="308" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFE65D469A8BFF1D1E62A43C" box="[954,1060,1094,1121]" pageId="7" pageNumber="308" value="2023-02-05">5.II.2023</date>
;
<specimenCount id="B78CFD0CFFE65D469D1EFF1D1E1FA43D" box="[1071,1113,1094,1120]" count="2" pageId="7" pageNumber="308" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFE65D469D57FF1D1E98A43C" box="[1126,1246,1094,1121]" pageId="7" pageNumber="308" value="2023-02-15">15.II.2023</date>
;
<specimenCount id="B78CFD0CFFE65D469DD8FF1D1F54A43D" box="[1257,1298,1094,1120]" count="2" pageId="7" pageNumber="308" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFE65D469C11FF1D1FD1A43C" box="[1312,1431,1094,1121]" pageId="7" pageNumber="308" value="2023-02-25">25.II.2023</date>
;
<specimenCount id="B78CFD0CFFE65D4699A6FF311A84A4D9" box="[151,194,1130,1156]" count="1" pageId="7" pageNumber="308" type="female">1 ♀</specimenCount>
;
<date id="D5341045FFE65D4699FEFF311B04A4D8" box="[207,322,1130,1157]" pageId="7" pageNumber="308" value="2023-03-05">5.III.2023</date>
;
<specimenCount id="B78CFD0CFFE65D46987CFF311B31A4D9" box="[333,375,1130,1156]" count="1" pageId="7" pageNumber="308" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFE65D4698B5FF311840A4D8" box="[388,518,1130,1157]" pageId="7" pageNumber="308" value="2023-03-25">25.III.2023</date>
. EL GUERRARA: El Amied, hand collecting,
<specimenCount id="B78CFD0CFFE65D469D21FF311E7CA4D9" box="[1040,1082,1130,1156]" count="2" pageId="7" pageNumber="308" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFE65D469D76FF311EF7A4D9" box="[1095,1201,1130,1156]" pageId="7" pageNumber="308" value="2020-02-07">7.II.2020</date>
.
<materialsCitation id="11E23CD8FFE65D469D8AFF311B03A4F5" location="Souareg" pageId="7" pageNumber="308" specimenCount="1">
METLILI:
<location id="A455605EFFE65D469C08FF311FDEA4D9" LSID="urn:lsid:plazi:treatment:29238793FFE65D4699A6F9031986A501:A455605EFFE65D469C08FF311FDEA4D9" box="[1337,1432,1130,1156]" name="Souareg" pageId="7" pageNumber="308">Souareg</location>
, hand collecting
</materialsCitation>
,
<specimenCount id="B78CFD0CFFE65D469861FFD51B3DA4F5" box="[336,379,1166,1192]" count="1" pageId="7" pageNumber="308" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFE65D4698B6FFD5184CA4F4" box="[391,522,1166,1193]" pageId="7" pageNumber="308" value="2021-11-25">25.XI.2021</date>
;
<specimenCount id="B78CFD0CFFE65D469B25FFD51806A4F5" box="[532,576,1166,1192]" count="1" pageId="7" pageNumber="308" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFE65D469B7AFFD5188FA4F4" box="[587,713,1166,1193]" pageId="7" pageNumber="308" value="2021-12-05">5.XII.2021</date>
;
<specimenCount id="B78CFD0CFFE65D469BE5FFD518BBA4F5" box="[724,765,1166,1192]" count="4" pageId="7" pageNumber="308" type="female">4 ♀</specimenCount>
,
<date id="D5341045FFE65D469A3AFFD51938A4F4" box="[779,894,1166,1193]" pageId="7" pageNumber="308" value="2022-03-05">5.III.2022</date>
;
<specimenCount id="B78CFD0CFFE65D469AB8FFD519F4A4F5" box="[905,946,1166,1192]" count="4" pageId="7" pageNumber="308" type="female">4 ♀</specimenCount>
,
<date id="D5341045FFE65D469AF1FFD51E07A4F4" box="[960,1089,1166,1193]" pageId="7" pageNumber="308" value="2022-03-25">25.III.2022</date>
;
<materialsCitation id="11E23CD8FFE65D469D7DFFD51FDEA4F5" box="[1100,1432,1166,1192]" location="Guemgouma" pageId="7" pageNumber="308" specimenCount="1">
<location id="A455605EFFE65D469D7DFFD51EA6A4F5" LSID="urn:lsid:plazi:treatment:29238793FFE65D4699A6F9031986A501:A455605EFFE65D469D7DFFD51EA6A4F5" box="[1100,1248,1166,1192]" name="Guemgouma" pageId="7" pageNumber="308">Guemgouma</location>
, hand collecting
</materialsCitation>
,
<specimenCount id="B78CFD0CFFE65D4699A6FFE91A87A491" box="[151,193,1202,1228]" count="2" pageId="7" pageNumber="308" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFE65D4699FEFFE91B17A490" box="[207,337,1202,1229]" pageId="7" pageNumber="308" value="2022-03-15">15.III.2022</date>
;
<specimenCount id="B78CFD0CFFE65D46986DFFE91BC0A491" box="[348,390,1202,1228]" count="1" pageId="7" pageNumber="308" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFE65D4698A5FFE91852A490" box="[404,532,1202,1229]" pageId="7" pageNumber="308" value="2022-03-25">25.III.2022</date>
, pitfall traps,
<specimenCount id="B78CFD0CFFE65D469B9FFFE9189EA491" box="[686,728,1202,1228]" count="1" pageId="7" pageNumber="308" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFE65D469BD7FFE91934A490" box="[742,882,1202,1229]" pageId="7" pageNumber="308" value="2021-12-25">25.XII.2021</date>
;
<specimenCount id="B78CFD0CFFE65D469A4FFFE919EEA491" box="[894,936,1202,1228]" count="1" pageId="7" pageNumber="308" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFE65D469A87FFE91E71A490" box="[950,1079,1202,1229]" pageId="7" pageNumber="308" value="2022-03-15">15.III.2022</date>
;
<specimenCount id="B78CFD0CFFE65D469D72FFE91E2BA491" box="[1091,1133,1202,1228]" count="1" pageId="7" pageNumber="308" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFE65D469D4AFFE91EBBA490" box="[1147,1277,1202,1229]" pageId="7" pageNumber="308" value="2022-03-25">25.III.2022</date>
.
</paragraph>
<paragraph id="A1353685FFE65D4699F6FF8E183FA565" blockId="7.[151,1437,769,1372]" pageId="7" pageNumber="308">
<emphasis id="93FEEA97FFE65D4699F6FF8E184BA4B2" bold="true" box="[199,525,1237,1264]" pageId="7" pageNumber="308">
Previous records in
<collectingCountry id="D99D7615FFE65D46989AFF8E184EA4B2" box="[427,520,1237,1263]" name="Algeria" pageId="7" pageNumber="308">Algeria</collectingCountry>
:
</emphasis>
Sahari,
<collectingRegion id="634EF867FFE65D469B55FF8D18EEA4AD" box="[612,680,1238,1264]" country="Algeria" name="Batna" pageId="7" pageNumber="308">Batna</collectingRegion>
,
<collectingRegion id="634EF867FFE65D469B80FF8D18BBA4AD" box="[689,765,1238,1264]" country="Algeria" name="Biskra" pageId="7" pageNumber="308">Biskra</collectingRegion>
,
<collectingRegion id="634EF867FFE65D469A34FF8D1902A4AD" box="[773,836,1238,1264]" country="Algeria" name="Blida" pageId="7" pageNumber="308">Blida</collectingRegion>
,
<collectingRegion id="634EF867FFE65D469A7CFF8D19DAA4AD" box="[845,924,1238,1264]" country="Algeria" name="Bouira" pageId="7" pageNumber="308">Bouira</collectingRegion>
,
<collectingRegion id="634EF867FFE65D469A95FF8D1E6BA4AD" box="[932,1069,1238,1264]" country="Algeria" name="Constantine" pageId="7" pageNumber="308">Constantine</collectingRegion>
,
<collectingRegion id="634EF867FFE65D469D04FF8D1E38A4AD" box="[1077,1150,1238,1264]" country="Algeria" name="Djelfa" pageId="7" pageNumber="308">Djelfa</collectingRegion>
,
<collectingRegion id="634EF867FFE65D469DB6FF8D1E9FA4AD" box="[1159,1241,1238,1264]" country="Algeria" name="Naama" pageId="7" pageNumber="308">Naama</collectingRegion>
,
<collectingRegion id="634EF867FFE65D469DD0FF8D1F64A4AD" box="[1249,1314,1238,1264]" country="Algeria" name="Saida" pageId="7" pageNumber="308">Saïda</collectingRegion>
,
<collectingRegion id="634EF867FFE65D469C1BFF8D1FDAA4AD" box="[1322,1436,1238,1264]" country="Algeria" name="Tissemsilt" pageId="7" pageNumber="308">Tissemsilt</collectingRegion>
and
<collectingRegion id="634EF867FFE65D4699F9FFA11B6CA549" box="[200,298,1274,1300]" country="Algeria" name="Tlemcen" pageId="7" pageNumber="308">Tlemcen</collectingRegion>
(
<bibRefCitation id="C51B4B74FFE65D46980AFFA11B91A549" author="Dalmas, R. de" box="[315,471,1274,1300]" pageId="7" pageNumber="308" pagination="233 - 328" refId="ref20501" refString="Dalmas, R. de (1921) Monographie des araignees de la section des Pterotricha (Aran. Gnaphosidae). Annales de la Societe Entomologique de France, 89, 233 - 328." type="journal article" year="1921">Dalmas 1921</bibRefCitation>
;
<bibRefCitation id="C51B4B74FFE65D4698D4FFA118DAA549" author="Di Franco, F." box="[485,668,1274,1300]" pageId="7" pageNumber="308" pagination="193 - 211" refId="ref20785" refString="Di Franco, F. (1994) Contributo alla conoscenza degli Gnaphosidae (Arachnida, Araneae) del Maghreb. Animalia, 19, 193 - 211." type="journal article" year="1994">Di Franco 1994</bibRefCitation>
; Brague-Bragba
<emphasis id="93FEEA97FFE65D469A6FFFA019DFA549" box="[862,921,1274,1300]" italics="true" pageId="7" pageNumber="308">et al.</emphasis>
2007;
<bibRefCitation id="C51B4B74FFE65D469ADAFFA11E98A549" author="Mansouri, H. &amp; Ould Rouis, S. &amp; Kherbouche-Abrous, O. &amp; Ould Rouis, A. &amp; Beladjal, L." box="[1003,1246,1274,1300]" pageId="7" pageNumber="308" pagination="1 - 13" refId="ref21811" refString="Mansouri, H., Ould Rouis, S., Kherbouche-Abrous, O., Ould Rouis, A. &amp; Beladjal, L. (2019) Effects of anthropogenic factors on spider communities (Arthropoda: Araneae) in Chrea National park (Blida, Algeria). African Journal of Ecology, 2019 (00), 1 - 13. https: // doi. org / 10.1111 / aje. 12701" type="journal article" year="2019">
Mansouri
<emphasis id="93FEEA97FFE65D469D6EFFA01EDCA549" box="[1119,1178,1274,1300]" italics="true" pageId="7" pageNumber="308">et al.</emphasis>
2019
</bibRefCitation>
;
<bibRefCitation id="C51B4B74FFE65D469DDDFFA11A95A565" author="Bouragba, N. &amp; Bouragba, M. &amp; Djouklafi, A. &amp; Amraoui, S. &amp; Brague, A. &amp; Beladjal, L." pageId="7" pageNumber="308" pagination="25 - 34" refId="ref19910" refString="Bouragba, N., Bouragba, M., Djouklafi, A., Amraoui, S., Brague, A. &amp; Beladjal, L. (2020) Spider (Araneae) and beetle (Coleoptera) communities structure in the North African streamside zone (Zahrez Gharbi, Djelfa, Algeria) after reforestation. African Entomology, 28 (1), 25 - 34." type="journal article" year="2020">
Bouragba
<emphasis id="93FEEA97FFE65D469C53FFA01FDBA549" box="[1378,1437,1274,1300]" italics="true" pageId="7" pageNumber="308">et al.</emphasis>
2020
</bibRefCitation>
;
<bibRefCitation id="C51B4B74FFE65D4699EFFE45182AA565" author="Chaib, S. &amp; Kherbouche-Abrous, O." box="[222,620,1310,1336]" pageId="7" pageNumber="308" pagination="129 - 138" refId="ref20171" refString="Chaib, S. &amp; Kherbouche-Abrous, O. (2021) Ecologie et phenologie des Gnaphosidae (Araneae, Arachnida) dans la region de Tikjda (Parc National du Djurdjura, Algerie). Bulletin de la Societe Zoologique de France, 146 (4), 129 - 138." type="journal article" year="2021">Chaib &amp; Kherbouche-Abrous 2021</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFE65D4699F6FE1A1986A501" box="[199,960,1345,1372]" pageId="7" pageNumber="308" type="distribution">
<paragraph id="A1353685FFE65D4699F6FE1A1986A501" blockId="7.[151,1437,769,1372]" box="[199,960,1345,1372]" pageId="7" pageNumber="308">
<emphasis id="93FEEA97FFE65D4699F6FE1A1B25A506" bold="true" box="[199,355,1345,1371]" pageId="7" pageNumber="308">Distribution:</emphasis>
<collectingCountry id="D99D7615FFE65D469859FE191BF9A501" box="[360,447,1346,1372]" name="Algeria" pageId="7" pageNumber="308">Algeria</collectingCountry>
,
<collectingCountry id="D99D7615FFE65D4698FBFE191866A501" box="[458,544,1346,1372]" name="Tunisia" pageId="7" pageNumber="308">Tunisia</collectingCountry>
,
<collectingCountry id="D99D7615FFE65D469B1AFE19182AA501" box="[555,620,1346,1372]" name="Libya" pageId="7" pageNumber="308">Libya</collectingCountry>
(
<bibRefCitation id="C51B4B74FFE65D469B4BFE1919F2A501" author="World Spider Catalog" box="[634,948,1346,1372]" pageId="7" pageNumber="308" refId="ref24111" refString="World Spider Catalog (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern, Bern. Available from: https: // wsc. nmbe. ch / (accessed 1 June 2024) https: // doi. org / 10.24436 / 2" type="book" year="2024">World Spider Catalog 2024</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,217 @@
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<mods:title id="A5E236B7319FF5A248A0DA6D28ECD8C9">Spiders of arid lands: The Ghardaïa region (Northern Sahara) with seven new records for Algeria</mods:title>
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<mods:affiliation id="6031D33DEC39B4286BC14E73AE9C43E5">Laboratoire de Valorisation et Conservation des Ecosystèmes Arides (LVCEA). Faculty of Natural, Life and Earth Sciences, University of Ghardaïa, BP 455, 47000 Ghardaïa, Algeria</mods:affiliation>
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<mods:affiliation id="4D44540C4576DF316B3FC3FEFAAB9D5E">Terrestrial Ecology Unit, Ledeganckstraat 35, B- 9000 Gent, Belgium</mods:affiliation>
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<subSubSection id="E990650EFFE65D4699A6FBCC187DA0EC" box="[151,571,151,177]" pageId="7" pageNumber="308" type="nomenclature">
<paragraph id="A1353685FFE65D4699A6FBCC187DA0EC" blockId="7.[151,571,151,177]" box="[151,571,151,177]" pageId="7" pageNumber="308">
<heading id="FA7D81E9FFE65D4699A6FBCC187DA0EC" bold="true" box="[151,571,151,177]" fontSize="11" level="1" pageId="7" pageNumber="308" reason="1">
<taxonomicName id="668A4D06FFE65D4699A6FBCC187DA0EC" ID-CoL="47P2W" authority="(L. Koch, 1872)" baseAuthorityName="L. Koch" baseAuthorityYear="1872" box="[151,571,151,177]" class="Arachnida" family="Gnaphosidae" genus="Nomisia" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="aussereri">
<emphasis id="93FEEA97FFE65D4699A6FBCC187DA0EC" bold="true" box="[151,571,151,177]" pageId="7" pageNumber="308">
<emphasis id="93FEEA97FFE65D4699A6FBCC1B35A0EC" bold="true" box="[151,371,151,177]" italics="true" pageId="7" pageNumber="308">Nomisia aussereri</emphasis>
(L. Koch, 1872)
</emphasis>
</taxonomicName>
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</paragraph>
</subSubSection>
<subSubSection id="E990650EFFE65D4699A6FB871F2DA14F" pageId="7" pageNumber="308" type="reference_group">
<paragraph id="A1353685FFE65D4699A6FB871897A0A9" blockId="7.[151,1387,219,274]" box="[151,721,219,244]" pageId="7" pageNumber="308">
<treatmentCitationGroup id="819A11ABFFE65D4699A6FB871897A0A9" box="[151,721,219,244]" pageId="7" pageNumber="308">
<taxonomicName id="668A4D06FFE65D4699A6FB871B1DA0AE" ID-CoL="47P3H" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1874" box="[151,347,220,243]" class="Arachnida" family="Gnaphosidae" genus="Nomisia" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="marginata">
<emphasis id="93FEEA97FFE65D4699A6FB871B1DA0AE" box="[151,347,220,243]" italics="true" pageId="7" pageNumber="308">Nomisia marginata</emphasis>
</taxonomicName>
;
<treatmentCitation id="202B1094FFE65D469859FB871863A0AE" author="Dalmas, R. de" box="[360,549,219,244]" page="296" pageId="7" pageNumber="308" year="1921">
<bibRefCitation id="C51B4B74FFE65D469859FB871863A0AE" author="Dalmas, R. de" box="[360,549,219,244]" pageId="7" pageNumber="308" pagination="233 - 328" refId="ref20501" refString="Dalmas, R. de (1921) Monographie des araignees de la section des Pterotricha (Aran. Gnaphosidae). Annales de la Societe Entomologique de France, 89, 233 - 328." type="journal article" year="1921">Dalmas 1921: 296</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="202B1094FFE65D469B1EFB87188BA0A9" author="Denis, J." box="[559,717,219,244]" page="48" pageId="7" pageNumber="308" year="1945">
<bibRefCitation id="C51B4B74FFE65D469B1EFB87188BA0A9" author="Denis, J." box="[559,717,219,244]" pageId="7" pageNumber="308" pagination="41 - 57" refId="ref20691" refString="Denis, J. (1945) Descriptions d'araignees nord-africaines. Bulletin de la Societe d'Histoire Naturelle de Toulouse, 79 (1), 41 - 57." type="journal article" year="1945">Denis 1945: 48</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A1353685FFE65D4699A6FBA01F2DA14F" blockId="7.[151,1387,219,274]" box="[151,1387,250,274]" pageId="7" pageNumber="308">
<treatmentCitationGroup id="819A11ABFFE65D4699A6FBA01F2DA14F" box="[151,1387,250,274]" pageId="7" pageNumber="308">
<taxonomicName id="668A4D06FFE65D4699A6FBA01B09A14F" ID-CoL="47P2W" baseAuthorityName="L. Koch" baseAuthorityYear="1872" box="[151,335,251,274]" class="Arachnida" family="Gnaphosidae" genus="Nomisia" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="aussereri">
<emphasis id="93FEEA97FFE65D4699A6FBA01B09A14F" box="[151,335,251,274]" italics="true" pageId="7" pageNumber="308">Nomisia aussereri</emphasis>
</taxonomicName>
;
<treatmentCitation id="202B1094FFE65D46986CFBA1185FA14F" author="Dalmas, R. de" box="[349,537,250,274]" page="297" pageId="7" pageNumber="308" year="1921">
<bibRefCitation id="C51B4B74FFE65D46986CFBA1185FA14F" author="Dalmas, R. de" box="[349,537,250,274]" pageId="7" pageNumber="308" pagination="233 - 328" refId="ref20501" refString="Dalmas, R. de (1921) Monographie des araignees de la section des Pterotricha (Aran. Gnaphosidae). Annales de la Societe Entomologique de France, 89, 233 - 328." type="journal article" year="1921">Dalmas 1921: 297</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="202B1094FFE65D469B15FBA118BFA14F" author="Di Franco, F." box="[548,761,250,274]" page="196" pageId="7" pageNumber="308" year="1994">
<bibRefCitation id="C51B4B74FFE65D469B15FBA118BFA14F" author="Di Franco, F." box="[548,761,250,274]" pageId="7" pageNumber="308" pagination="193 - 211" refId="ref20785" refString="Di Franco, F. (1994) Contributo alla conoscenza degli Gnaphosidae (Arachnida, Araneae) del Maghreb. Animalia, 19, 193 - 211." type="journal article" year="1994">Di Franco 1994: 196</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="202B1094FFE65D469A33FBA11E2AA14F" author="Abrous-Kherbouche, O. &amp; Jocque, R. &amp; Maelfait, J. - P." box="[770,1132,250,274]" page="80" pageId="7" pageNumber="308" year="1997">
<bibRefCitation id="C51B4B74FFE65D469A33FBA11E2AA14F" author="Abrous-Kherbouche, O. &amp; Jocque, R. &amp; Maelfait, J. - P." box="[770,1132,250,274]" pageId="7" pageNumber="308" pagination="71 - 90" refId="ref18469" refString="Abrous-Kherbouche, O., Jocque, R. &amp; Maelfait, J. - P. (1997) Les effets du paturage intensif sur l'araneofaune dans la region de Tala-Guilef (Parc National du Djurdjura, Algerie). Annales de la Societe Entomologique de Belgique, 133, 71 - 90." type="journal article" year="1997">
Abrous-Kherbouche
<emphasis id="93FEEA97FFE65D469AE6FBA01E4DA14F" box="[983,1035,250,274]" italics="true" pageId="7" pageNumber="308">et al.</emphasis>
1997: 80
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="202B1094FFE65D469D46FBA11F21A14F" author="Mansouri, H. &amp; Ould Rouis, S. &amp; Kherbouche-Abrous, O. &amp; Ould Rouis, A. &amp; Beladjal, L." box="[1143,1383,250,274]" page="4" pageId="7" pageNumber="308" year="2019">
<bibRefCitation id="C51B4B74FFE65D469D46FBA11F21A14F" author="Mansouri, H. &amp; Ould Rouis, S. &amp; Kherbouche-Abrous, O. &amp; Ould Rouis, A. &amp; Beladjal, L." box="[1143,1383,250,274]" pageId="7" pageNumber="308" pagination="1 - 13" refId="ref21811" refString="Mansouri, H., Ould Rouis, S., Kherbouche-Abrous, O., Ould Rouis, A. &amp; Beladjal, L. (2019) Effects of anthropogenic factors on spider communities (Arthropoda: Araneae) in Chrea National park (Blida, Algeria). African Journal of Ecology, 2019 (00), 1 - 13. https: // doi. org / 10.1111 / aje. 12701" type="journal article" year="2019">
Mansouri
<emphasis id="93FEEA97FFE65D469DEEFBA01F54A14F" box="[1247,1298,250,274]" italics="true" pageId="7" pageNumber="308">et al.</emphasis>
2019: 4
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFE65D4699A6FA1B1F03A19A" pageId="7" pageNumber="308" type="materials_examined">
<paragraph id="A1353685FFE65D4699A6FA1B1F5BA122" blockId="7.[151,1437,320,527]" pageId="7" pageNumber="308">
<emphasis id="93FEEA97FFE65D4699A6FA1B1BC4A107" bold="true" box="[151,386,320,346]" pageId="7" pageNumber="308">Material examined:</emphasis>
<materialsCitation id="11E23CD8FFE65D4698B6FA1A1E75A106" box="[391,1075,320,347]" collectingDate="2014-01-07" location="Mansoura Kedima" pageId="7" pageNumber="308" specimenCount="1" specimenCount-female="1">
MANSOURA:
<location id="A455605EFFE65D469B04FA1A1941A107" LSID="urn:lsid:plazi:treatment:29238793FFE65D4699A6FBCC191EA252:A455605EFFE65D469B04FA1A1941A107" box="[565,775,320,347]" name="Mansoura Kedima" pageId="7" pageNumber="308">Mansoura Kedima</location>
,
<collectingMethod id="78CB4E92FFE65D469A20FA1B19D4A106" box="[785,914,320,347]" pageId="7" pageNumber="308">pitfall traps</collectingMethod>
,
<specimenCount id="B78CFD0CFFE65D469AACFA1A1980A106" box="[925,966,321,347]" count="1" pageId="7" pageNumber="308" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFE65D469AE3FA1A1E75A106" box="[978,1075,321,347]" pageId="7" pageNumber="308" value="2014-01-07">
<collectingDate id="C570E9ADFFE65D469AE3FA1A1E75A106" box="[978,1075,321,347]" pageId="7" pageNumber="308" value="2014-01-07">7.I.2014</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="11E23CD8FFE65D469D0CFA1A1EA4A106" box="[1085,1250,320,347]" collectingDate="2015-01-12" isTailEnumeration="true" location="Mansoura Kedima" pageId="7" pageNumber="308" specimenCount="1" specimenCount-female="1">
<specimenCount id="B78CFD0CFFE65D469D0CFA1A1E20A106" box="[1085,1126,321,347]" count="1" pageId="7" pageNumber="308" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFE65D469D42FA1B1EA4A106" box="[1139,1250,320,347]" pageId="7" pageNumber="308" value="2015-01-12">
<collectingDate id="C570E9ADFFE65D469D42FA1B1EA4A106" box="[1139,1250,320,347]" pageId="7" pageNumber="308" value="2015-01-12">12.I.2015</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="11E23CD8FFE65D469DDAFA1A1815A122" collectingDate="2015-03-07" location="Oued Sebseb" pageId="7" pageNumber="308" specimenCount="1" specimenCount-female="1">
SEBSEB:
<location id="A455605EFFE65D469C51FA1B1AAFA123" LSID="urn:lsid:plazi:treatment:29238793FFE65D4699A6FBCC191EA252:A455605EFFE65D469C51FA1B1AAFA123" name="Oued Sebseb" pageId="7" pageNumber="308">Oued Sebseb</location>
, hand collecting,
<specimenCount id="B78CFD0CFFE65D46989CFA3F1B97A122" box="[429,465,356,383]" count="1" pageId="7" pageNumber="308" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFE65D4698EEFA3F1815A122" box="[479,595,356,383]" pageId="7" pageNumber="308" value="2015-03-07">
<collectingDate id="C570E9ADFFE65D4698EEFA3F1815A122" box="[479,595,356,383]" pageId="7" pageNumber="308" value="2015-03-07">7.III.2015</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="11E23CD8FFE65D469B6CFA3E1F5FA122" box="[605,1305,356,383]" collectingDate="2015-02-14" location="Dune of Zelfana" pageId="7" pageNumber="308" specimenCount="2" specimenCount-female="2">
ZELFANA:
<location id="A455605EFFE65D469BDBFA3E19E2A123" LSID="urn:lsid:plazi:treatment:29238793FFE65D4699A6FBCC191EA252:A455605EFFE65D469BDBFA3E19E2A123" box="[746,932,356,383]" name="Dune of Zelfana" pageId="7" pageNumber="308">Dune of Zelfana</location>
, hand collecting,
<specimenCount id="B78CFD0CFFE65D469D58FA3E1ED5A122" box="[1129,1171,357,383]" count="2" pageId="7" pageNumber="308" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFE65D469D90FA3F1F5FA122" box="[1185,1305,356,383]" pageId="7" pageNumber="308" value="2015-02-14">
<collectingDate id="C570E9ADFFE65D469D90FA3F1F5FA122" box="[1185,1305,356,383]" pageId="7" pageNumber="308" value="2015-02-14">14.II.2015</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="A1353685FFE65D4699F6FAD31F03A19A" blockId="7.[151,1437,320,527]" pageId="7" pageNumber="308">
<emphasis id="93FEEA97FFE65D4699F6FAD31858A1FF" bold="true" box="[199,542,392,419]" pageId="7" pageNumber="308">
Previous records in
<collectingCountry id="D99D7615FFE65D46988DFAD3185FA1FF" box="[444,537,392,418]" name="Algeria" pageId="7" pageNumber="308">Algeria</collectingCountry>
:
</emphasis>
Without precise locality,
<collectingRegion id="634EF867FFE65D469A77FAD319E6A1FF" box="[838,928,392,418]" country="Algeria" name="Annaba" pageId="7" pageNumber="308">Annaba</collectingRegion>
,
<collectingRegion id="634EF867FFE65D469A9FFAD219B4A1FE" box="[942,1010,393,419]" country="Algeria" name="Batna" pageId="7" pageNumber="308">Batna</collectingRegion>
,
<collectingRegion id="634EF867FFE65D469D31FAD31E0AA1FF" box="[1024,1100,392,418]" country="Algeria" name="Biskra" pageId="7" pageNumber="308">Biskra</collectingRegion>
,
<collectingRegion id="634EF867FFE65D469D6BFAD31EDFA1FF" box="[1114,1177,392,418]" country="Algeria" name="Blida" pageId="7" pageNumber="308">Blida</collectingRegion>
,
<collectingRegion id="634EF867FFE65D469D96FAD31E9FA1FF" box="[1191,1241,392,418]" country="Algeria" name="Jijel" pageId="7" pageNumber="308">Jijel</collectingRegion>
,
<collectingRegion id="634EF867FFE65D469DD6FAD31F73A1FF" box="[1255,1333,392,418]" country="Algeria" name="Skikda" pageId="7" pageNumber="308">Skikda</collectingRegion>
and
<collectingRegion id="634EF867FFE65D469C41FAD31AA4A19A" country="Algeria" name="Tizi Ouzou" pageId="7" pageNumber="308">Tizi Ouzou</collectingRegion>
(
<bibRefCitation id="C51B4B74FFE65D4699C1FAF71BCCA19A" author="Dalmas, R. de" box="[240,394,428,455]" pageId="7" pageNumber="308" pagination="233 - 328" refId="ref20501" refString="Dalmas, R. de (1921) Monographie des araignees de la section des Pterotricha (Aran. Gnaphosidae). Annales de la Societe Entomologique de France, 89, 233 - 328." type="journal article" year="1921">Dalmas 1921</bibRefCitation>
;
<bibRefCitation id="C51B4B74FFE65D4698A4FAF7185FA19A" author="Denis, J." box="[405,537,428,455]" pageId="7" pageNumber="308" pagination="41 - 57" refId="ref20691" refString="Denis, J. (1945) Descriptions d'araignees nord-africaines. Bulletin de la Societe d'Histoire Naturelle de Toulouse, 79 (1), 41 - 57." type="journal article" year="1945">Denis 1945</bibRefCitation>
;
<bibRefCitation id="C51B4B74FFE65D469B14FAF71892A19A" author="Di Franco, F." box="[549,724,428,455]" pageId="7" pageNumber="308" pagination="193 - 211" refId="ref20785" refString="Di Franco, F. (1994) Contributo alla conoscenza degli Gnaphosidae (Arachnida, Araneae) del Maghreb. Animalia, 19, 193 - 211." type="journal article" year="1994">Di Franco 1994</bibRefCitation>
<bibRefCitation id="C51B4B74FFE65D469BEBFAF71E79A19A" author="Abrous-Kherbouche, O. &amp; Jocque, R. &amp; Maelfait, J. - P." box="[730,1087,428,455]" pageId="7" pageNumber="308" pagination="71 - 90" refId="ref18469" refString="Abrous-Kherbouche, O., Jocque, R. &amp; Maelfait, J. - P. (1997) Les effets du paturage intensif sur l'araneofaune dans la region de Tala-Guilef (Parc National du Djurdjura, Algerie). Annales de la Societe Entomologique de Belgique, 133, 71 - 90." type="journal article" year="1997">
Abrous-Kherbouche
<emphasis id="93FEEA97FFE65D469AF4FAF619B8A19B" box="[965,1022,428,454]" italics="true" pageId="7" pageNumber="308">et al.</emphasis>
1997
</bibRefCitation>
;
<bibRefCitation id="C51B4B74FFE65D469D7AFAF71F7FA19A" author="Mansouri, H. &amp; Ould Rouis, S. &amp; Kherbouche-Abrous, O. &amp; Ould Rouis, A. &amp; Beladjal, L." box="[1099,1337,428,455]" pageId="7" pageNumber="308" pagination="1 - 13" refId="ref21811" refString="Mansouri, H., Ould Rouis, S., Kherbouche-Abrous, O., Ould Rouis, A. &amp; Beladjal, L. (2019) Effects of anthropogenic factors on spider communities (Arthropoda: Araneae) in Chrea National park (Blida, Algeria). African Journal of Ecology, 2019 (00), 1 - 13. https: // doi. org / 10.1111 / aje. 12701" type="journal article" year="2019">
Mansouri
<emphasis id="93FEEA97FFE65D469D8CFAF61EB0A19B" box="[1213,1270,428,454]" italics="true" pageId="7" pageNumber="308">et al.</emphasis>
2019
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFE65D4699F6FA8B191EA252" pageId="7" pageNumber="308" type="distribution">
<paragraph id="A1353685FFE65D4699F6FA8B191EA252" blockId="7.[151,1437,320,527]" pageId="7" pageNumber="308">
<emphasis id="93FEEA97FFE65D4699F6FA8B1B25A1B7" bold="true" box="[199,355,464,490]" pageId="7" pageNumber="308">Distribution:</emphasis>
Mediterranean, Eastern Europe,
<collectingCountry id="D99D7615FFE65D469BFDFA8B195BA1B7" box="[716,797,464,490]" name="Turkey" pageId="7" pageNumber="308">Turkey</collectingCountry>
, Middle East, Caucasus,
<collectingCountry id="D99D7615FFE65D469D1CFA8B1E3EA1B7" box="[1069,1144,464,490]" name="Russia" pageId="7" pageNumber="308">Russia</collectingCountry>
(Europe to South Siberia),
<collectingCountry id="D99D7615FFE65D4699A6FAAF1B5BA253" box="[151,285,500,526]" name="Kazakhstan" pageId="7" pageNumber="308">Kazakhstan</collectingCountry>
, Central Asia,
<collectingCountry id="D99D7615FFE65D4698F0FAAF1842A253" box="[449,516,500,526]" name="China" pageId="7" pageNumber="308">China</collectingCountry>
(
<bibRefCitation id="C51B4B74FFE65D469B23FAAF190AA252" author="World Spider Catalog" box="[530,844,500,527]" pageId="7" pageNumber="308" refId="ref24111" refString="World Spider Catalog (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern, Bern. Available from: https: // wsc. nmbe. ch / (accessed 1 June 2024) https: // doi. org / 10.24436 / 2" type="book" year="2024">World Spider Catalog 2024</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,145 @@
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<mods:title id="050D08CBB20689BB8830A6746BBA13A0">Spiders of arid lands: The Ghardaïa region (Northern Sahara) with seven new records for Algeria</mods:title>
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<mods:namePart id="0C95821F9924E11F505E7399F691248E">Alioua, Youcef</mods:namePart>
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<mods:namePart id="C09879177B6BCA2B8593E8FFDFCBC6D3">Bosmans, Robert</mods:namePart>
<mods:affiliation id="59AEB7E92A0B0EB73169E6AE4D1FC873">Terrestrial Ecology Unit, Ledeganckstraat 35, B- 9000 Gent, Belgium</mods:affiliation>
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<mods:date id="6ED1FB81C5074E21AAC569595AB1B801">2024</mods:date>
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<subSubSection id="E990650EFFE65D4699A6FEF51849A59A" box="[151,527,1453,1480]" pageId="7" pageNumber="308" type="nomenclature">
<paragraph id="A1353685FFE65D4699A6FEF51849A59A" blockId="7.[151,527,1453,1480]" box="[151,527,1453,1480]" pageId="7" pageNumber="308">
<heading id="FA7D81E9FFE65D4699A6FEF51849A59A" bold="true" box="[151,527,1453,1480]" fontSize="11" level="1" pageId="7" pageNumber="308" reason="1">
<taxonomicName id="668A4D06FFE65D4699A6FEF51849A59A" ID-CoL="47P3X" authority="(Pavesi, 1880)" baseAuthorityName="Pavesi" baseAuthorityYear="1880" box="[151,527,1453,1480]" class="Arachnida" family="Gnaphosidae" genus="Nomisia" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="recepta">
<emphasis id="93FEEA97FFE65D4699A6FEF51849A59A" bold="true" box="[151,527,1453,1480]" pageId="7" pageNumber="308">
<emphasis id="93FEEA97FFE65D4699A6FEF51B1AA595" bold="true" box="[151,348,1454,1480]" italics="true" pageId="7" pageNumber="308">Nomisia recepta</emphasis>
(
<bibRefCitation id="C51B4B74FFE65D46985AFEF61841A59A" author="Pavesi, P." box="[363,519,1453,1479]" pageId="7" pageNumber="308" pagination="283 - 388" refId="ref22217" refString="Pavesi, P. (1880) Studi sugli Aracnidi africani. I. Aracnidi di Tunisia. Annali del Museo Civico di Storia Naturale di Genova, 15, 283 - 388." type="journal article" year="1880">Pavesi, 1880</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFE65D4699A6FEA81894A657" box="[151,722,1522,1546]" pageId="7" pageNumber="308" type="reference_group">
<paragraph id="A1353685FFE65D4699A6FEA81894A657" blockId="7.[151,722,1522,1546]" box="[151,722,1522,1546]" pageId="7" pageNumber="308">
<treatmentCitationGroup id="819A11ABFFE65D4699A6FEA81894A657" box="[151,722,1522,1546]" pageId="7" pageNumber="308">
<taxonomicName id="668A4D06FFE65D4699A6FEA81B7BA657" ID-CoL="47P3X" baseAuthorityName="Pavesi" baseAuthorityYear="1880" box="[151,317,1523,1546]" class="Arachnida" family="Gnaphosidae" genus="Nomisia" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="recepta">
<emphasis id="93FEEA97FFE65D4699A6FEA81B7BA657" box="[151,317,1523,1546]" italics="true" pageId="7" pageNumber="308">Nomisia recepta</emphasis>
</taxonomicName>
;
<treatmentCitation id="202B1094FFE65D46987BFEA91841A657" author="Dalmas, R. de" box="[330,519,1522,1546]" page="283" pageId="7" pageNumber="308" year="1921">
<bibRefCitation id="C51B4B74FFE65D46987BFEA91841A657" author="Dalmas, R. de" box="[330,519,1522,1546]" pageId="7" pageNumber="308" pagination="233 - 328" refId="ref20501" refString="Dalmas, R. de (1921) Monographie des araignees de la section des Pterotricha (Aran. Gnaphosidae). Annales de la Societe Entomologique de France, 89, 233 - 328." type="journal article" year="1921">Dalmas 1921: 283</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="202B1094FFE65D469B23FEA91888A657" author="Chatzaki, M." box="[530,718,1522,1546]" page="15" pageId="7" pageNumber="308" year="2010">
<bibRefCitation id="C51B4B74FFE65D469B23FEA91888A657" author="Chatzaki, M." box="[530,718,1522,1546]" pageId="7" pageNumber="308" pagination="1 - 22" refId="ref20290" refString="Chatzaki, M. (2010) A revision of the genus Nomisia in Greece and neighboring regions with the description of two new species. Zootaxa, 2501 (1), 1 - 22. https: // doi. org / 10.11646 / zootaxa. 2501.1.1" type="journal article" year="2010">Chatzaki 2010: 15</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFE65D4699A6FD631E9CA6C6" pageId="7" pageNumber="308" type="materials_examined">
<paragraph id="A1353685FFE65D4699A6FD631942A62A" blockId="7.[151,1436,1592,1727]" pageId="7" pageNumber="308">
<emphasis id="93FEEA97FFE65D4699A6FD631BC3A60F" bold="true" box="[151,389,1592,1618]" pageId="7" pageNumber="308">Material examined:</emphasis>
BERRIANE: Balouh, hand collecting,
<specimenCount id="B78CFD0CFFE65D469A79FD621932A60E" box="[840,884,1593,1619]" count="2" pageId="7" pageNumber="308" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFE65D469AB5FD621E45A60E" box="[900,1027,1593,1619]" pageId="7" pageNumber="308" value="2021-04-24">24.IV.2021</date>
;
<materialsCitation id="11E23CD8FFE65D469D21FD621E8DA60E" box="[1040,1227,1593,1619]" collectingDate="2021-04-30" isTailEnumeration="true" pageId="7" pageNumber="308" specimenCount="1" specimenCount-male="1">
<specimenCount id="B78CFD0CFFE65D469D21FD621E78A60E" box="[1040,1086,1593,1619]" count="1" pageId="7" pageNumber="308" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFE65D469D7DFD621E8DA60E" box="[1100,1227,1593,1619]" pageId="7" pageNumber="308" value="2021-04-30">
<collectingDate id="C570E9ADFFE65D469D7DFD621E8DA60E" box="[1100,1227,1593,1619]" pageId="7" pageNumber="308" value="2021-04-30">30.IV.2021</collectingDate>
</date>
</materialsCitation>
, pitfall traps,
<specimenCount id="B78CFD0CFFE65D469C58FD621FD3A60E" box="[1385,1429,1593,1619]" count="1" pageId="7" pageNumber="308" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFE65D4699A6FD061B50A62A" box="[151,278,1629,1655]" pageId="7" pageNumber="308" value="2021-04-17">17.IV.2021</date>
. Sidi Mbarek, pitfall traps,
<specimenCount id="B78CFD0CFFE65D469B78FD061833A62A" box="[585,629,1629,1655]" count="1" pageId="7" pageNumber="308" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFE65D469BB0FD061947A62A" box="[641,769,1629,1655]" pageId="7" pageNumber="308" value="2021-04-23">23.IV.2021</date>
.
</paragraph>
<paragraph id="A1353685FFE65D4699F6FDDB1E9CA6C6" blockId="7.[151,1436,1592,1727]" box="[199,1242,1664,1691]" pageId="7" pageNumber="308">
<emphasis id="93FEEA97FFE65D4699F6FDDB1853A6C7" bold="true" box="[199,533,1664,1691]" pageId="7" pageNumber="308">
Previous records in
<collectingCountry id="D99D7615FFE65D469882FDDB1856A6C7" box="[435,528,1664,1690]" name="Algeria" pageId="7" pageNumber="308">Algeria</collectingCountry>
:
</emphasis>
Known from
<collectingRegion id="634EF867FFE65D469B82FDDA18B2A6C6" box="[691,756,1665,1691]" country="Algeria" name="Batna" pageId="7" pageNumber="308">Batna</collectingRegion>
and
<collectingRegion id="634EF867FFE65D469A1AFDDB1935A6C7" box="[811,883,1664,1690]" country="Algeria" name="M'Sila" pageId="7" pageNumber="308">Msila</collectingRegion>
(
<bibRefCitation id="C51B4B74FFE65D469AB0FDDB1E5CA6C6" author="Dalmas, R. de" box="[897,1050,1664,1691]" pageId="7" pageNumber="308" pagination="233 - 328" refId="ref20501" refString="Dalmas, R. de (1921) Monographie des araignees de la section des Pterotricha (Aran. Gnaphosidae). Annales de la Societe Entomologique de France, 89, 233 - 328." type="journal article" year="1921">Dalmas 1921</bibRefCitation>
;
<bibRefCitation id="C51B4B74FFE65D469D17FDDB1E88A6C6" author="Chatzaki, M." box="[1062,1230,1664,1691]" pageId="7" pageNumber="308" pagination="1 - 22" refId="ref20290" refString="Chatzaki, M. (2010) A revision of the genus Nomisia in Greece and neighboring regions with the description of two new species. Zootaxa, 2501 (1), 1 - 22. https: // doi. org / 10.11646 / zootaxa. 2501.1.1" type="journal article" year="2010">Chatzaki 2010</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFE65D4699F6FDFF1F77A6E2" box="[199,1329,1700,1727]" pageId="7" pageNumber="308" type="distribution">
<paragraph id="A1353685FFE65D4699F6FDFF1F77A6E2" blockId="7.[151,1436,1592,1727]" box="[199,1329,1700,1727]" pageId="7" pageNumber="308">
<emphasis id="93FEEA97FFE65D4699F6FDFF1B25A6E3" bold="true" box="[199,355,1700,1726]" pageId="7" pageNumber="308">Distribution:</emphasis>
<collectingCountry id="D99D7615FFE65D469858FDFF1BF9A6E3" box="[361,447,1700,1726]" name="Tunisia" pageId="7" pageNumber="308">Tunisia</collectingCountry>
,
<collectingCountry id="D99D7615FFE65D4698F8FDFF1866A6E3" box="[457,544,1700,1726]" name="Algeria" pageId="7" pageNumber="308">Algeria</collectingCountry>
,
<collectingCountry id="D99D7615FFE65D469B1AFDFF1818A6E3" box="[555,606,1700,1726]" name="Italy" pageId="7" pageNumber="308">Italy</collectingCountry>
(mainland,
<collectingRegion id="634EF867FFE65D469BD4FDFF196CA6E3" box="[741,810,1700,1726]" country="Italy" name="Sicilia" pageId="7" pageNumber="308">Sicily</collectingRegion>
),
<collectingCountry id="D99D7615FFE65D469A0CFDFF19C7A6E3" box="[829,897,1700,1726]" name="Malta" pageId="7" pageNumber="308">Malta</collectingCountry>
,
<collectingCountry id="D99D7615FFE65D469ABCFDFE1998A6E2" box="[909,990,1701,1727]" name="Cyprus" pageId="7" pageNumber="308">Cyprus</collectingCountry>
(
<bibRefCitation id="C51B4B74FFE65D469ADDFDFF1F63A6E2" author="World Spider Catalog" box="[1004,1317,1700,1727]" pageId="7" pageNumber="308" refId="ref24111" refString="World Spider Catalog (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern, Bern. Available from: https: // wsc. nmbe. ch / (accessed 1 June 2024) https: // doi. org / 10.24436 / 2" type="book" year="2024">World Spider Catalog 2024</bibRefCitation>
).
</paragraph>
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EL ATTEUF: Palm grove of El Atteuf, pitfall traps,
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Previous records in
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Known only from
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<emphasis id="93FEEA97FFEB5D4B99A6FB871B0EA0AE" box="[151,328,220,244]" italics="true" pageId="10" pageNumber="311">Zelotes inauratus</emphasis>
</taxonomicName>
;
<treatmentCitation id="202B1094FFEB5D4B9864FB871865A0AE" author="Marinaro, J. - Y." box="[341,547,219,244]" page="695" pageId="10" pageNumber="311" year="1968">
<bibRefCitation id="C51B4B74FFEB5D4B9864FB871865A0AE" author="Marinaro, J. - Y." box="[341,547,219,244]" pageId="10" pageNumber="311" pagination="687 - 704" refId="ref21888" refString="Marinaro, J. - Y. (1968) Les araignees d'Afrique du Nord. I. Sur une collection de Drassidae a peigne metatarsal d'Algerie. Bulletin de la Societe Zoologique de France, 92, 687 - 704." type="journal article" year="1968">Marinaro 1968: 695</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFEB5D4B99A6FA7A1F6DA191" pageId="10" pageNumber="311" type="materials_examined">
<paragraph id="A1353685FFEB5D4B99A6FA7A1F28A1F5" blockId="10.[151,1437,289,532]" pageId="10" pageNumber="311">
<emphasis id="93FEEA97FFEB5D4B99A6FA7A1BC6A166" bold="true" box="[151,384,289,315]" pageId="10" pageNumber="311">Material examined:</emphasis>
BERRIANE: Balouh,
<specimenCount id="B78CFD0CFFEB5D4B9B4BFA7918E4A161" box="[634,674,290,316]" count="1" pageId="10" pageNumber="311" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFEB5D4B9B9CFA79196BA161" box="[685,813,290,316]" pageId="10" pageNumber="311" value="2021-04-10">10.IV.2021</date>
;
<materialsCitation id="11E23CD8FFEB5D4B9A07FA791E5AA161" box="[822,1052,290,316]" collectingDate="2021-04-24" isTailEnumeration="true" pageId="10" pageNumber="311" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="B78CFD0CFFEB5D4B9A07FA791919A161" box="[822,863,290,316]" count="1" pageId="10" pageNumber="311" type="male">1 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFEB5D4B9A58FA7919D7A161" box="[873,913,290,316]" count="1" pageId="10" pageNumber="311" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFEB5D4B9AADFA791E5AA161" box="[924,1052,290,316]" pageId="10" pageNumber="311" value="2021-04-24">
<collectingDate id="C570E9ADFFEB5D4B9AADFA791E5AA161" box="[924,1052,290,316]" pageId="10" pageNumber="311" value="2021-04-24">24.IV.2021</collectingDate>
</date>
</materialsCitation>
. Sidi Mbarek, hand collecting,
<specimenCount id="B78CFD0CFFEB5D4B9C5FFA791FD1A161" box="[1390,1431,290,316]" count="1" pageId="10" pageNumber="311" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFEB5D4B99A6FA1D1B48A13D" box="[151,270,326,352]" pageId="10" pageNumber="311" value="2021-02-11">11.II.2021</date>
;
<materialsCitation id="11E23CD8FFEB5D4B9828FA1D1B84A13D" box="[281,450,326,352]" collectingDate="2021-04-09" isTailEnumeration="true" pageId="10" pageNumber="311" specimenCount="1" specimenCount-male="1">
<specimenCount id="B78CFD0CFFEB5D4B9828FA1D1B03A13D" box="[281,325,326,352]" count="1" pageId="10" pageNumber="311" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFEB5D4B9860FA1D1B84A13D" box="[337,450,326,352]" pageId="10" pageNumber="311" value="2021-04-09">
<collectingDate id="C570E9ADFFEB5D4B9860FA1D1B84A13D" box="[337,450,326,352]" pageId="10" pageNumber="311" value="2021-04-09">9.IV.2021</collectingDate>
</date>
</materialsCitation>
. DHAYET BENDHAHOUA: Grazil, hand collecting,
<specimenCount id="B78CFD0CFFEB5D4B9D15FA1D1E16A13D" box="[1060,1104,326,352]" count="2" pageId="10" pageNumber="311" type="male">2 ♂</specimenCount>
,
<date id="D5341045FFEB5D4B9D6DFA1D1E95A13C" box="[1116,1235,326,353]" pageId="10" pageNumber="311" value="2022-02-15">15.II.2022</date>
;
<materialsCitation id="11E23CD8FFEB5D4B9DEEFA1D1FDEA13C" box="[1247,1432,326,353]" collectingDate="2022-03-15" isTailEnumeration="true" pageId="10" pageNumber="311" specimenCount="1" specimenCount-female="1">
<specimenCount id="B78CFD0CFFEB5D4B9DEEFA1D1F4FA13D" box="[1247,1289,326,352]" count="1" pageId="10" pageNumber="311" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFEB5D4B9C26FA1D1FDEA13C" box="[1303,1432,326,353]" pageId="10" pageNumber="311" value="2022-03-15">
<collectingDate id="C570E9ADFFEB5D4B9C26FA1D1FDEA13C" box="[1303,1432,326,353]" pageId="10" pageNumber="311" value="2022-03-15">15.III.2022</collectingDate>
</date>
</materialsCitation>
;
<specimenCount id="B78CFD0CFFEB5D4B99A6FA311A83A1D9" box="[151,197,362,388]" count="1" pageId="10" pageNumber="311" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFEB5D4B99E3FA311B15A1D8" box="[210,339,362,389]" pageId="10" pageNumber="311" value="2022-03-25">25.III.2022</date>
; Oumdjer, hand collecting,
<specimenCount id="B78CFD0CFFEB5D4B9BBEFA3118FDA1D9" box="[655,699,362,388]" count="1" pageId="10" pageNumber="311" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFEB5D4B9BFBFA311975A1D8" box="[714,819,362,389]" pageId="10" pageNumber="311" value="2022-02-05">5.II.2022</date>
;
<materialsCitation id="11E23CD8FFEB5D4B9A70FA3119B2A1D8" box="[833,1012,362,389]" collectingDate="2022-02-25" isTailEnumeration="true" pageId="10" pageNumber="311" specimenCount="1" specimenCount-female="1">
<specimenCount id="B78CFD0CFFEB5D4B9A70FA31192AA1D9" box="[833,876,362,388]" count="1" pageId="10" pageNumber="311" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFEB5D4B9A4DFA3119B2A1D8" box="[892,1012,362,389]" pageId="10" pageNumber="311" value="2022-02-25">
<collectingDate id="C570E9ADFFEB5D4B9A4DFA3119B2A1D8" box="[892,1012,362,389]" pageId="10" pageNumber="311" value="2022-02-25">25.II.2022</collectingDate>
</date>
</materialsCitation>
;
<specimenCount id="B78CFD0CFFEB5D4B9D30FA311E68A1D9" box="[1025,1070,362,388]" count="1" pageId="10" pageNumber="311" type="male">1 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFEB5D4B9D0DFA311E21A1D9" box="[1084,1127,362,388]" count="1" pageId="10" pageNumber="311" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFEB5D4B9D46FA311EBEA1D8" box="[1143,1272,362,389]" pageId="10" pageNumber="311" value="2022-03-15">15.III.2022</date>
. EL ATTEUF: Palm grove of El Atteuf, pitfall traps,
<specimenCount id="B78CFD0CFFEB5D4B9B0EFAD5182DA1F5" box="[575,619,398,424]" count="2" pageId="10" pageNumber="311" type="male">2 ♂</specimenCount>
,
<date id="D5341045FFEB5D4B9B46FAD518B0A1F4" box="[631,758,398,425]" pageId="10" pageNumber="311" value="2015-04-18">18.IV.2015</date>
. EL GUERRARA: Aghzou, pitfall traps
<specimenCount id="B78CFD0CFFEB5D4B9D8FFAD51EACA1F5" box="[1214,1258,398,424]" count="1" pageId="10" pageNumber="311" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFEB5D4B9DC7FAD51F2CA1F5" box="[1270,1386,398,424]" pageId="10" pageNumber="311" value="2020-03-03">3.III.2020</date>
.
</paragraph>
<paragraph id="A1353685FFEB5D4B99F6FAEA1F6DA191" blockId="10.[151,1437,289,532]" box="[199,1323,433,460]" pageId="10" pageNumber="311">
<emphasis id="93FEEA97FFEB5D4B99F6FAEA1853A196" bold="true" box="[199,533,433,460]" pageId="10" pageNumber="311">
Previous records in
<collectingCountry id="D99D7615FFEB5D4B9882FAEA1856A196" box="[435,528,433,459]" name="Algeria" pageId="10" pageNumber="311">Algeria</collectingCountry>
:
</emphasis>
Until now, it is known only from Still in El Megaier (
<bibRefCitation id="C51B4B74FFEB5D4B9D40FAE91F58A191" author="Marinaro, J. - Y." box="[1137,1310,434,460]" pageId="10" pageNumber="311" pagination="687 - 704" refId="ref21888" refString="Marinaro, J. - Y. (1968) Les araignees d'Afrique du Nord. I. Sur une collection de Drassidae a peigne metatarsal d'Algerie. Bulletin de la Societe Zoologique de France, 92, 687 - 704." type="journal article" year="1968">Marinaro 1968</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFEB5D4B99F6FA8E1F76A249" pageId="10" pageNumber="311" type="distribution">
<paragraph id="A1353685FFEB5D4B99F6FA8E1F76A249" blockId="10.[151,1437,289,532]" pageId="10" pageNumber="311">
<emphasis id="93FEEA97FFEB5D4B99F6FA8E1B25A1B2" bold="true" box="[199,355,469,495]" pageId="10" pageNumber="311">Distribution:</emphasis>
North Africa to
<collectingCountry id="D99D7615FFEB5D4B9B1CFA8D18C0A1AD" box="[557,646,470,496]" name="Senegal" pageId="10" pageNumber="311">Senegal</collectingCountry>
and
<collectingCountry id="D99D7615FFEB5D4B9BF4FA8D1957A1AD" box="[709,785,470,496]" name="Kenya" pageId="10" pageNumber="311">Kenya</collectingCountry>
,
<collectingCountry id="D99D7615FFEB5D4B9A11FA8D19C4A1AD" box="[800,898,470,496]" name="Portugal" pageId="10" pageNumber="311">Portugal</collectingCountry>
,
<collectingCountry id="D99D7615FFEB5D4B9AA0FA8D19A6A1AD" box="[913,992,470,496]" name="France" pageId="10" pageNumber="311">France</collectingCountry>
,
<collectingCountry id="D99D7615FFEB5D4B9AC1FA8D1E79A1AD" box="[1008,1087,470,496]" name="Greece" pageId="10" pageNumber="311">Greece</collectingCountry>
(
<collectingRegion id="634EF867FFEB5D4B9D63FA8D1ED2A1AD" box="[1106,1172,470,496]" country="Greece" name="Kriti" pageId="10" pageNumber="311">Crete</collectingRegion>
),
<collectingCountry id="D99D7615FFEB5D4B9D9DFA8D1EBBA1AD" box="[1196,1277,470,496]" name="Turkey" pageId="10" pageNumber="311">Turkey</collectingCountry>
,
<collectingCountry id="D99D7615FFEB5D4B9C3DFA8D1F0BA1AD" box="[1292,1357,470,496]" name="Israel" pageId="10" pageNumber="311">Israel</collectingCountry>
,
<collectingCountry id="D99D7615FFEB5D4B9C6DFA8D1AA0A249" name="Saudi Arabia" pageId="10" pageNumber="311">Saudi Arabia</collectingCountry>
,
<collectingCountry id="D99D7615FFEB5D4B99C0FAA11B64A249" box="[241,290,506,532]" name="Iran" pageId="10" pageNumber="311">Iran</collectingCountry>
. Introduced to
<collectingRegion id="634EF867FFEB5D4B98FBFAA11858A249" box="[458,542,506,532]" country="United States of America" name="Hawaii" pageId="10" pageNumber="311">Hawaii</collectingRegion>
,
<collectingCountry id="D99D7615FFEB5D4B9B19FAA11825A249" box="[552,611,506,532]" name="United States of America" pageId="10" pageNumber="311">USA</collectingCountry>
,
<collectingCountry id="D99D7615FFEB5D4B9B5FFAA11880A249" box="[622,710,506,532]" name="Mexico" pageId="10" pageNumber="311">Mexico</collectingCountry>
,
<collectingCountry id="D99D7615FFEB5D4B9BE3FAA1194EA249" box="[722,776,506,532]" name="Peru" pageId="10" pageNumber="311">Peru</collectingCountry>
,
<collectingCountry id="D99D7615FFEB5D4B9A23FAA119C1A249" box="[786,903,506,532]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="10" pageNumber="311">Ascension</collectingCountry>
Islands (
<bibRefCitation id="C51B4B74FFEB5D4B9ADAFAA11F63A249" author="World Spider Catalog" box="[1003,1317,506,532]" pageId="10" pageNumber="311" refId="ref24111" refString="World Spider Catalog (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern, Bern. Available from: https: // wsc. nmbe. ch / (accessed 1 June 2024) https: // doi. org / 10.24436 / 2" type="book" year="2024">World Spider Catalog 2024</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,159 @@
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<heading id="FA7D81E9FFEC5D4C99A6F9B0181AA358" bold="true" box="[151,604,747,774]" fontSize="11" level="1" pageId="13" pageNumber="314" reason="1">
<taxonomicName id="668A4D06FFEC5D4C99A6F9B0181AA358" ID-CoL="G4DH" authority="(Blackwall, 1841)" baseAuthorityName="Blackwall" baseAuthorityYear="1841" box="[151,604,747,774]" class="Arachnida" family="Linyphiidae" genus="Araeoncus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="humilis">
<emphasis id="93FEEA97FFEC5D4C99A6F9B0181AA358" bold="true" box="[151,604,747,774]" pageId="13" pageNumber="314">
<emphasis id="93FEEA97FFEC5D4C99A6F9B01B3AA35B" bold="true" box="[151,380,747,774]" italics="true" pageId="13" pageNumber="314">Araeoncus humilis</emphasis>
(Blackwall, 1841)
</emphasis>
</taxonomicName>
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<emphasis id="93FEEA97FFEC5D4C99A6F86B1B13A315" box="[151,341,816,840]" italics="true" pageId="13" pageNumber="314">Araeoncus humilis</emphasis>
</taxonomicName>
;
<treatmentCitation id="202B1094FFEC5D4C9853F86B1855A31A" author="Simon, E." box="[354,531,815,840]" page="636" pageId="13" pageNumber="314" year="1884">
<bibRefCitation id="C51B4B74FFEC5D4C9853F86B1855A31A" author="Simon, E." box="[354,531,815,840]" pageId="13" pageNumber="314" pagination="180 - 885" refId="ref23149" refString="Simon, E. (1884) n. k. In: Les arachnides de France. Tome Cinquieme, Deuxieme et Troisieme Partie. Roret, Paris, pp. 180 - 885." type="book chapter" year="1884">Simon 1884: 636</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="202B1094FFEC5D4C9B2FF86B18A4A315" author="Denis, J." box="[542,738,816,840]" page="1042" pageId="13" pageNumber="314" year="1937">
<bibRefCitation id="C51B4B74FFEC5D4C9B2FF86B18A4A315" author="Denis, J." box="[542,738,816,840]" pageId="13" pageNumber="314" pagination="1027 - 1060" refId="ref20639" refString="Denis, J. (1937 b) On a collection of spiders from Algeria. Proceedings of the Zoological Society of London, 106 (4), 1027 - 1060. https: // doi. org / 10.1111 / j. 1469 - 7998.1936. tb 06301. x" type="journal article" year="1937">Denis 1937b: 1042</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="202B1094FFEC5D4C9BDDF86B19FEA31A" author="Bosmans, R." box="[748,952,815,839]" page="125" pageId="13" pageNumber="314" year="1996">
<bibRefCitation id="C51B4B74FFEC5D4C9BDDF86B19FEA31A" author="Bosmans, R." box="[748,952,815,839]" pageId="13" pageNumber="314" pagination="123 - 151" refId="ref19317" refString="Bosmans, R. (1996) The genera Araeoncus Simon, Delorripis Simon and Diplocephalus Bertkau in Northern Africa (Araneae: Linyphiidae: Erigoninae). Studies on North African Linyphiidae VII. Belgian Journal of Zoology, 126 (2), 123 - 151." type="journal article" year="1996">Bosmans 1996: 125</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="202B1094FFEC5D4C9AF2F86B1F46A31A" author="Bosmans, R. &amp; Chergui, F." box="[963,1280,815,840]" page="356" pageId="13" pageNumber="314" year="1994">
<bibRefCitation id="C51B4B74FFEC5D4C9AF2F86B1F46A31A" author="Bosmans, R. &amp; Chergui, F." box="[963,1280,815,840]" pageId="13" pageNumber="314" pagination="341 - 358" refId="ref19264" refString="Bosmans, R. &amp; Chergui, F. (1994) The genus Mecopisthes Simon in North Africa (Araneae: Linyphiidae: Erigoninae). Studies on North African Linyphiidae VII. Bulletin &amp; Annales de la Societe Entomologique de Belgique, 129 (4), 341 - 358." type="journal article" year="1994">Bosmans &amp; Chergui 1994: 356</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A1353685FFEC5D4C99A6F8141800A33B" blockId="13.[151,1284,815,870]" box="[151,582,846,870]" pageId="13" pageNumber="314">
<treatmentCitationGroup id="819A11ABFFEC5D4C99A6F8141800A33B" box="[151,582,846,870]" pageId="13" pageNumber="314">
<taxonomicName id="668A4D06FFEC5D4C99A6F8141BCCA33B" ID-CoL="G4DR" authorityName="Denis" authorityYear="1954" box="[151,394,846,870]" class="Arachnida" family="Linyphiidae" genus="Araeoncus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="meridionalis">
<emphasis id="93FEEA97FFEC5D4C99A6F8141BCCA33B" box="[151,394,846,870]" italics="true" pageId="13" pageNumber="314">Araeoncus meridionalis</emphasis>
</taxonomicName>
;
<treatmentCitation id="202B1094FFEC5D4C98A6F8151804A33B" author="Denis, J." box="[407,578,846,870]" page="313" pageId="13" pageNumber="314" year="1954">
<bibRefCitation id="C51B4B74FFEC5D4C98A6F8151804A33B" author="Denis, J." box="[407,578,846,870]" pageId="13" pageNumber="314" pagination="311 - 324" refId="ref20753" refString="Denis, J. (1954) Araignees recueillies par P. Remy du Sud-Algerien au Hoggar. Bulletin de la Societe Zoologique de France, 78, 311 - 324." type="journal article" year="1954">Denis 1954: 313</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFEC5D4C99A6F8CF19ADA3AB" pageId="13" pageNumber="314" type="materials_examined">
<paragraph id="A1353685FFEC5D4C99A6F8CF1EE9A3F2" blockId="13.[151,1436,916,1087]" box="[151,1199,916,943]" pageId="13" pageNumber="314">
<emphasis id="93FEEA97FFEC5D4C99A6F8CF1BC5A3F3" bold="true" box="[151,387,916,942]" pageId="13" pageNumber="314">Material examined:</emphasis>
EL ATTEUF: Palm grove of El Atteuf, hand collecting,
<specimenCount id="B78CFD0CFFEC5D4C9ACAF8CE1E63A3F2" box="[1019,1061,917,943]" count="1" pageId="13" pageNumber="314" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFEC5D4C9D02F8CF1EEDA3F2" box="[1075,1195,916,943]" pageId="13" pageNumber="314" value="2015-02-25">25.II.2015</date>
.
</paragraph>
<paragraph id="A1353685FFEC5D4C99F6F8E319ADA3AB" blockId="13.[151,1436,916,1087]" pageId="13" pageNumber="314">
<emphasis id="93FEEA97FFEC5D4C99F6F8E31858A38F" bold="true" box="[199,542,952,979]" pageId="13" pageNumber="314">
Previous records in
<collectingCountry id="D99D7615FFEC5D4C988DF8E3185FA38F" box="[444,537,952,978]" name="Algeria" pageId="13" pageNumber="314">Algeria</collectingCountry>
:
</emphasis>
<collectingRegion id="634EF867FFEC5D4C9B16F8E3182FA38F" box="[551,617,952,978]" country="Algeria" name="Alger" pageId="13" pageNumber="314">Alger</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C9B47F8E218FCA38E" box="[630,698,953,979]" country="Algeria" name="Batna" pageId="13" pageNumber="314">Batna</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C9BF9F8E31905A38F" box="[712,835,952,978]" country="Algeria" name="El Bayadh" pageId="13" pageNumber="314">El Bayadh</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C9A63F8E319E3A38F" box="[850,933,952,978]" country="Algeria" name="El Tarf" pageId="13" pageNumber="314">El Tarf</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C9A82F8E31E66A38F" box="[947,1056,952,978]" country="Algeria" name="Laghouat" pageId="13" pageNumber="314">Laghouat</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C9D1FF8E31E20A38F" box="[1070,1126,952,978]" country="Algeria" name="Mila" pageId="13" pageNumber="314">Mila</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C9D45F8E31EF8A38F" box="[1140,1214,952,978]" country="Algeria" name="M'Sila" pageId="13" pageNumber="314">Msila</collectingRegion>
, Touggourt (
<bibRefCitation id="C51B4B74FFEC5D4C9C60F8E31A94A3AA" author="Simon, E." pageId="13" pageNumber="314" pagination="180 - 885" refId="ref23149" refString="Simon, E. (1884) n. k. In: Les arachnides de France. Tome Cinquieme, Deuxieme et Troisieme Partie. Roret, Paris, pp. 180 - 885." type="book chapter" year="1884">Simon 1884</bibRefCitation>
;
<bibRefCitation id="C51B4B74FFEC5D4C99EFF8871B36A3AB" author="Denis, J." box="[222,368,988,1014]" pageId="13" pageNumber="314" pagination="1027 - 1060" refId="ref20639" refString="Denis, J. (1937 b) On a collection of spiders from Algeria. Proceedings of the Zoological Society of London, 106 (4), 1027 - 1060. https: // doi. org / 10.1111 / j. 1469 - 7998.1936. tb 06301. x" type="journal article" year="1937">Denis 1937b</bibRefCitation>
;
<bibRefCitation id="C51B4B74FFEC5D4C984AF8871BB8A3AA" author="Denis, J." box="[379,510,988,1015]" pageId="13" pageNumber="314" pagination="311 - 324" refId="ref20753" refString="Denis, J. (1954) Araignees recueillies par P. Remy du Sud-Algerien au Hoggar. Bulletin de la Societe Zoologique de France, 78, 311 - 324." type="journal article" year="1954">Denis 1954</bibRefCitation>
;
<bibRefCitation id="C51B4B74FFEC5D4C9B3BF8861976A3AA" author="Bosmans, R. &amp; Chergui, F." box="[522,816,988,1015]" pageId="13" pageNumber="314" pagination="341 - 358" refId="ref19264" refString="Bosmans, R. &amp; Chergui, F. (1994) The genus Mecopisthes Simon in North Africa (Araneae: Linyphiidae: Erigoninae). Studies on North African Linyphiidae VII. Bulletin &amp; Annales de la Societe Entomologique de Belgique, 129 (4), 341 - 358." type="journal article" year="1994">Bosmans &amp; Chergui 1994</bibRefCitation>
;
<bibRefCitation id="C51B4B74FFEC5D4C9A0DF88619A5A3AB" author="Bosmans, R." box="[828,995,988,1015]" pageId="13" pageNumber="314" pagination="123 - 151" refId="ref19317" refString="Bosmans, R. (1996) The genera Araeoncus Simon, Delorripis Simon and Diplocephalus Bertkau in Northern Africa (Araneae: Linyphiidae: Erigoninae). Studies on North African Linyphiidae VII. Belgian Journal of Zoology, 126 (2), 123 - 151." type="journal article" year="1996">Bosmans 1996</bibRefCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFEC5D4C99F6FF5B1BA2A462" pageId="13" pageNumber="314" type="distribution">
<paragraph id="A1353685FFEC5D4C99F6FF5B1BA2A462" blockId="13.[151,1436,916,1087]" pageId="13" pageNumber="314">
<emphasis id="93FEEA97FFEC5D4C99F6FF5B1B25A447" bold="true" box="[199,355,1024,1050]" pageId="13" pageNumber="314">Distribution:</emphasis>
Europe, North Africa,
<collectingCountry id="D99D7615FFEC5D4C9B55FF5B18E9A447" box="[612,687,1024,1050]" name="Russia" pageId="13" pageNumber="314">Russia</collectingCountry>
(Europe to South Siberia),
<collectingCountry id="D99D7615FFEC5D4C9AD0FF5A1E57A446" box="[993,1041,1025,1051]" name="Iran" pageId="13" pageNumber="314">Iran</collectingCountry>
,
<collectingCountry id="D99D7615FFEC5D4C9D2DFF5A1E19A446" box="[1052,1119,1025,1051]" name="Japan" pageId="13" pageNumber="314">Japan</collectingCountry>
. Introduced to
<collectingCountry id="D99D7615FFEC5D4C9C37FF5A1FDAA447" box="[1286,1436,1024,1051]" name="New Zealand" pageId="13" pageNumber="314">New Zealand</collectingCountry>
(
<bibRefCitation id="C51B4B74FFEC5D4C99AFFF7F1B9EA462" author="World Spider Catalog" box="[158,472,1060,1087]" pageId="13" pageNumber="314" refId="ref24111" refString="World Spider Catalog (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern, Bern. Available from: https: // wsc. nmbe. ch / (accessed 1 June 2024) https: // doi. org / 10.24436 / 2" type="book" year="2024">World Spider Catalog 2024</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,211 @@
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<paragraph id="A1353685FFEC5D4C99A6FB8418E5A0A4" blockId="13.[151,675,223,250]" box="[151,675,223,250]" pageId="13" pageNumber="314">
<heading id="FA7D81E9FFEC5D4C99A6FB8418E5A0A4" bold="true" box="[151,675,223,250]" fontSize="11" level="1" pageId="13" pageNumber="314" reason="1">
<taxonomicName id="668A4D06FFEC5D4C99A6FB8418E5A0A4" ID-CoL="BBRJ" authority="Wunderlich, 1980" authorityName="Wunderlich" authorityYear="1980" box="[151,675,223,250]" class="Arachnida" family="Linyphiidae" genus="Agyneta" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="pseudorurestris">
<emphasis id="93FEEA97FFEC5D4C99A6FB8418E5A0A4" bold="true" box="[151,675,223,250]" pageId="13" pageNumber="314">
<emphasis id="93FEEA97FFEC5D4C99A6FB841BFBA0A7" bold="true" box="[151,445,223,250]" italics="true" pageId="13" pageNumber="314">Agyneta pseudorurestris</emphasis>
Wunderlich, 1980
</emphasis>
</taxonomicName>
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</paragraph>
</subSubSection>
<subSubSection id="E990650EFFEC5D4C99A6FA7F18D9A124" pageId="13" pageNumber="314" type="reference_group">
<paragraph id="A1353685FFEC5D4C99A6FA7F1E3FA161" blockId="13.[151,1145,291,377]" box="[151,1145,291,316]" pageId="13" pageNumber="314">
<treatmentCitationGroup id="819A11ABFFEC5D4C99A6FA7F1E3FA161" box="[151,1145,291,316]" pageId="13" pageNumber="314">
<taxonomicName id="668A4D06FFEC5D4C99A6FA7F1B1AA166" baseAuthorityName="C.L.Koch" baseAuthorityYear="1836" box="[151,348,292,315]" class="Arachnida" family="Linyphiidae" genus="Microneta" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="rurestris">
<emphasis id="93FEEA97FFEC5D4C99A6FA7F1B1AA166" box="[151,348,292,315]" italics="true" pageId="13" pageNumber="314">Microneta rurestris</emphasis>
</taxonomicName>
;
<treatmentCitation id="202B1094FFEC5D4C985BFA7F185EA166" author="Simon, E." box="[362,536,291,316]" page="436" pageId="13" pageNumber="314" year="1884">
<bibRefCitation id="C51B4B74FFEC5D4C985BFA7F185EA166" author="Simon, E." box="[362,536,291,316]" pageId="13" pageNumber="314" pagination="180 - 885" refId="ref23149" refString="Simon, E. (1884) n. k. In: Les arachnides de France. Tome Cinquieme, Deuxieme et Troisieme Partie. Roret, Paris, pp. 180 - 885." type="book chapter" year="1884">Simon 1884: 436</bibRefCitation>
</treatmentCitation>
(Misidentification);
<treatmentCitation id="202B1094FFEC5D4C9BDAFA7F19EBA161" author="Denis, J." box="[747,941,292,316]" page="1044" pageId="13" pageNumber="314" year="1937">
<bibRefCitation id="C51B4B74FFEC5D4C9BDAFA7F19EBA161" author="Denis, J." box="[747,941,292,316]" pageId="13" pageNumber="314" pagination="1027 - 1060" refId="ref20639" refString="Denis, J. (1937 b) On a collection of spiders from Algeria. Proceedings of the Zoological Society of London, 106 (4), 1027 - 1060. https: // doi. org / 10.1111 / j. 1469 - 7998.1936. tb 06301. x" type="journal article" year="1937">Denis 1937b: 1044</bibRefCitation>
</treatmentCitation>
(Misidentification).
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</paragraph>
<paragraph id="A1353685FFEC5D4C99A6FA18183FA107" blockId="13.[151,1145,291,377]" box="[151,633,322,346]" pageId="13" pageNumber="314">
<treatmentCitationGroup id="819A11ABFFEC5D4C99A6FA18183FA107" box="[151,633,322,346]" pageId="13" pageNumber="314">
<taxonomicName id="668A4D06FFEC5D4C99A6FA181BDDA107" baseAuthorityName="Wunderlich" baseAuthorityYear="1980" box="[151,411,322,346]" class="Arachnida" family="Linyphiidae" genus="Meioneta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="pseudorurestris">
<emphasis id="93FEEA97FFEC5D4C99A6FA181BDDA107" box="[151,411,322,346]" italics="true" pageId="13" pageNumber="314">Meioneta pseudorurestris</emphasis>
</taxonomicName>
;
<treatmentCitation id="202B1094FFEC5D4C9898FA181833A107" author="Bosmans, R." box="[425,629,322,346]" page="140" pageId="13" pageNumber="314" year="2006">
<bibRefCitation id="C51B4B74FFEC5D4C9898FA181833A107" author="Bosmans, R." box="[425,629,322,346]" pageId="13" pageNumber="314" pagination="125 - 161" refId="ref19501" refString="Bosmans, R. (2006) Contribution to the knowledge of the Linyphiidae of the Maghreb. Part XI. Miscellaneous linyphiid genera and additions (Araneae: Linyphiidae: Linyphiinae). Bulletin &amp; Annales de la Societe Entomologique de Belgique, 141, 125 - 161." type="journal article" year="2006">Bosmans 2006: 140</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A1353685FFEC5D4C99A6FA3A18D9A124" blockId="13.[151,1145,291,377]" box="[151,671,353,377]" pageId="13" pageNumber="314">
<treatmentCitationGroup id="819A11ABFFEC5D4C99A6FA3A18D9A124" box="[151,671,353,377]" pageId="13" pageNumber="314">
<taxonomicName id="668A4D06FFEC5D4C99A6FA3A1BC9A124" ID-CoL="BBRJ" authorityName="Wunderlich" authorityYear="1980" box="[151,399,353,377]" class="Arachnida" family="Linyphiidae" genus="Agyneta" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="pseudorurestris">
<emphasis id="93FEEA97FFEC5D4C99A6FA3A1BC9A124" box="[151,399,353,377]" italics="true" pageId="13" pageNumber="314">Agyneta pseudorurestris</emphasis>
</taxonomicName>
;
<treatmentCitation id="202B1094FFEC5D4C98ADFA3A18DDA124" author="Boucherit, S. &amp; Bouragba, N. &amp; Douaoui, A. &amp; Bosmans, R." box="[412,667,353,377]" page="63" pageId="13" pageNumber="314" year="2020">
<bibRefCitation id="C51B4B74FFEC5D4C98ADFA3A18DDA124" author="Boucherit, S. &amp; Bouragba, N. &amp; Douaoui, A. &amp; Bosmans, R." box="[412,667,353,377]" pageId="13" pageNumber="314" pagination="60 - 70" refId="ref19840" refString="Boucherit, S., Bouragba, N., Douaoui, A. &amp; Bosmans, R. (2020) Araignees de la plaine alluviale du Haut-Cheliff (Algerie), avec la citation de quelques especes nouvelles ou rares pour l'Algerie (Araneae: Gnaphosidae, Miturgidae). Nieuwsbrief van de Belgische Arachnologische Vereniging, 35 (1 - 2), 60 - 70." type="journal article" year="2020">
Boucherit
<emphasis id="93FEEA97FFEC5D4C9B37FA39187CA124" box="[518,570,353,377]" italics="true" pageId="13" pageNumber="314">et al.</emphasis>
2020: 63
</bibRefCitation>
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.
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</paragraph>
</subSubSection>
<subSubSection id="E990650EFFEC5D4C99A6FAFC1B59A228" pageId="13" pageNumber="314" type="materials_examined">
<paragraph id="A1353685FFEC5D4C99A6FAFC1E6AA19C" blockId="13.[151,1437,423,665]" box="[151,1068,423,450]" pageId="13" pageNumber="314">
<emphasis id="93FEEA97FFEC5D4C99A6FAFC1BC5A19C" bold="true" box="[151,387,423,449]" pageId="13" pageNumber="314">Material examined:</emphasis>
BERRIANE: Sidi Mbarek, hand collecting,
<specimenCount id="B78CFD0CFFEC5D4C9A49FAFC19E4A19F" box="[888,930,423,450]" count="1" pageId="13" pageNumber="314" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFEC5D4C9A81FAFC1E61A19C" box="[944,1063,423,449]" pageId="13" pageNumber="314" value="2021-02-18">18.II.2021</date>
.
</paragraph>
<paragraph id="A1353685FFEC5D4C99F6FA901B59A228" blockId="13.[151,1437,423,665]" pageId="13" pageNumber="314">
<emphasis id="93FEEA97FFEC5D4C99F6FA90185AA1B8" bold="true" box="[199,540,459,485]" pageId="13" pageNumber="314">
Previous records in
<collectingCountry id="D99D7615FFEC5D4C988BFA901851A1B8" box="[442,535,459,485]" name="Algeria" pageId="13" pageNumber="314">Algeria</collectingCountry>
:
</emphasis>
<collectingRegion id="634EF867FFEC5D4C9B15FA9018DEA1B8" box="[548,664,459,485]" country="Algeria" name="Ain Defla" pageId="13" pageNumber="314">Aïn Defla</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C9B95FA901920A1B8" box="[676,870,459,485]" country="Algeria" name="Ain Temouchent" pageId="13" pageNumber="314">Aïn Témouchent</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C9A43FA9019F2A1B8" box="[882,948,459,485]" country="Algeria" name="Alger" pageId="13" pageNumber="314">Alger</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C9A8EFA901E5FA1B8" box="[959,1049,459,485]" country="Algeria" name="Annaba" pageId="13" pageNumber="314">Annaba</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C9D17FA901E36A1B8" box="[1062,1136,459,485]" country="Algeria" name="Bejaia" pageId="13" pageNumber="314">Béjaïa</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C9D4FFA901E8CA1B8" box="[1150,1226,459,485]" country="Algeria" name="Biskra" pageId="13" pageNumber="314">Biskra</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C9DE6FA901F50A1B8" box="[1239,1302,459,485]" country="Algeria" name="Blida" pageId="13" pageNumber="314">Blida</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C9C15FA901AB2A254" country="Algeria" name="Bordj Bou Arreridj" pageId="13" pageNumber="314">Bordj Bou Arréridj</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C9833FAB41B17A254" box="[258,337,495,521]" country="Algeria" name="Bouira" pageId="13" pageNumber="314">Bouira</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C986EFAB41BA7A254" box="[351,481,495,521]" country="Algeria" name="Boumerdes" pageId="13" pageNumber="314">Boumerdès</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C98DEFAB41876A254" box="[495,560,495,521]" country="Algeria" name="Chlef" pageId="13" pageNumber="314">Chlef</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C9B0FFAB41881A254" box="[574,711,495,521]" country="Algeria" name="Constantine" pageId="13" pageNumber="314">Constantine</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C9BE4FAB41958A254" box="[725,798,495,521]" country="Algeria" name="Djelfa" pageId="13" pageNumber="314">Djelfa</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C9A1DFAB419E0A254" box="[812,934,495,521]" country="Algeria" name="El Bayadh" pageId="13" pageNumber="314">El Bayadh</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C9A84FAB41E50A254" box="[949,1046,495,521]" country="Algeria" name="El Oued" pageId="13" pageNumber="314">El Oued</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C9D14FAB41E3EA254" box="[1061,1144,495,521]" country="Algeria" name="El Tarf" pageId="13" pageNumber="314">El Tarf</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C9DB7FAB41EB6A254" box="[1158,1264,495,521]" country="Algeria" name="Ghardaia" pageId="13" pageNumber="314">Ghardaïa</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C9DCCFAB41F1FA254" box="[1277,1369,495,521]" country="Algeria" name="Guelma" pageId="13" pageNumber="314">Guelma</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C9C57FAB41FDEA254" box="[1382,1432,495,521]" country="Algeria" name="Jijel" pageId="13" pageNumber="314">Jijel</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C99A6F9481B42A270" box="[151,260,531,557]" country="Algeria" name="Laghouat" pageId="13" pageNumber="314">Laghouat</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C9821F9481B0EA270" box="[272,328,531,557]" country="Algeria" name="Mila" pageId="13" pageNumber="314">Mila</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C9864F9481BD9A270" box="[341,415,531,557]" country="Algeria" name="M'Sila" pageId="13" pageNumber="314">Msila</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C989DF9481BA1A270" box="[428,487,531,557]" country="Algeria" name="Oran" pageId="13" pageNumber="314">Oran</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C98C5F948181FA270" box="[500,601,531,557]" country="Algeria" name="Relizane" pageId="13" pageNumber="314">Relizane</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C9B54F94818E0A270" box="[613,678,531,557]" country="Algeria" name="Saida" pageId="13" pageNumber="314">Saïda</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C9B82F94818ADA270" box="[691,747,531,557]" country="Algeria" name="Setif" pageId="13" pageNumber="314">Sétif</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C9BC6F94819E3A270" box="[759,933,531,557]" country="Algeria" name="Sidi Bel Abbes" pageId="13" pageNumber="314">Sidi Bel Abbès</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C9A83F9481E45A270" box="[946,1027,531,557]" country="Algeria" name="Skikda" pageId="13" pageNumber="314">Skikda</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C9D3EF9481E2AA270" box="[1039,1132,531,557]" country="Algeria" name="Tebessa" pageId="13" pageNumber="314">Tébessa</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C9D49F9481EFBA270" box="[1144,1213,531,557]" country="Algeria" name="Tiaret" pageId="13" pageNumber="314">Tiaret</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C9DF8F9481F51A270" box="[1225,1303,531,557]" country="Algeria" name="Tipaza" pageId="13" pageNumber="314">Tipaza</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C9C12F9481FDEA270" box="[1315,1432,531,557]" country="Algeria" name="Tissemsilt" pageId="13" pageNumber="314">Tissemsilt</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C99A6F96C1B51A20C" box="[151,279,567,593]" country="Algeria" name="Tizi Ouzou" pageId="13" pageNumber="314">Tizi Ouzou</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C9813F96C1BC2A20C" box="[290,388,567,593]" country="Algeria" name="Tlemcen" pageId="13" pageNumber="314">Tlemcen</collectingRegion>
(
<bibRefCitation id="C51B4B74FFEC5D4C98A3F96C185DA20C" author="Simon, E." box="[402,539,567,593]" pageId="13" pageNumber="314" pagination="180 - 885" refId="ref23149" refString="Simon, E. (1884) n. k. In: Les arachnides de France. Tome Cinquieme, Deuxieme et Troisieme Partie. Roret, Paris, pp. 180 - 885." type="book chapter" year="1884">Simon 1884</bibRefCitation>
(misidentification),
<bibRefCitation id="C51B4B74FFEC5D4C9A31F96C19C8A20C" author="Denis, J." box="[768,910,567,593]" pageId="13" pageNumber="314" pagination="1027 - 1060" refId="ref20639" refString="Denis, J. (1937 b) On a collection of spiders from Algeria. Proceedings of the Zoological Society of London, 106 (4), 1027 - 1060. https: // doi. org / 10.1111 / j. 1469 - 7998.1936. tb 06301. x" type="journal article" year="1937">Denis 1937b</bibRefCitation>
(misidentification);
<bibRefCitation id="C51B4B74FFEC5D4C9D42F9631F5AA20C" author="Bosmans, R." box="[1139,1308,567,594]" pageId="13" pageNumber="314" pagination="125 - 161" refId="ref19501" refString="Bosmans, R. (2006) Contribution to the knowledge of the Linyphiidae of the Maghreb. Part XI. Miscellaneous linyphiid genera and additions (Araneae: Linyphiidae: Linyphiinae). Bulletin &amp; Annales de la Societe Entomologique de Belgique, 141, 125 - 161." type="journal article" year="2006">Bosmans 2006</bibRefCitation>
; Boucherit.
<emphasis id="93FEEA97FFEC5D4C99A6F9071A89A228" box="[151,207,603,629]" italics="true" pageId="13" pageNumber="314">et al.</emphasis>
2020).
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFEC5D4C99F6F9241927A2C4" box="[199,865,639,665]" pageId="13" pageNumber="314" type="distribution">
<paragraph id="A1353685FFEC5D4C99F6F9241927A2C4" blockId="13.[151,1437,423,665]" box="[199,865,639,665]" pageId="13" pageNumber="314">
<emphasis id="93FEEA97FFEC5D4C99F6F9241B25A2C4" bold="true" box="[199,355,639,665]" pageId="13" pageNumber="314">Distribution:</emphasis>
Mediterranean (
<bibRefCitation id="C51B4B74FFEC5D4C9B2AF9241913A2C4" author="World Spider Catalog" box="[539,853,639,665]" pageId="13" pageNumber="314" refId="ref24111" refString="World Spider Catalog (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern, Bern. Available from: https: // wsc. nmbe. ch / (accessed 1 June 2024) https: // doi. org / 10.24436 / 2" type="book" year="2024">World Spider Catalog 2024</bibRefCitation>
).
</paragraph>
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</treatment>
</document>

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@ -0,0 +1,153 @@
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<heading id="FA7D81E9FFEC5D4C99A6FDBF1949A6A3" bold="true" box="[151,783,1764,1791]" fontSize="11" level="1" pageId="13" pageNumber="314" reason="1">
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<emphasis id="93FEEA97FFEC5D4C99A6FDBF1BCDA6A2" bold="true" box="[151,395,1764,1791]" italics="true" pageId="13" pageNumber="314">Styloctetor romanus</emphasis>
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<emphasis id="93FEEA97FFEC5D4C99A6FC721B25A71C" box="[151,355,1833,1857]" italics="true" pageId="13" pageNumber="314">Styloctetor romanus</emphasis>
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;
<treatmentCitation id="202B1094FFEC5D4C9B1DFC7218BFA71C" author="Bosmans, R." box="[556,761,1833,1857]" page="139" pageId="13" pageNumber="314" year="2007">
<bibRefCitation id="C51B4B74FFEC5D4C9B1DFC7218BFA71C" author="Bosmans, R." box="[556,761,1833,1857]" pageId="13" pageNumber="314" pagination="117 - 163" refId="ref19551" refString="Bosmans, R. (2007) Contribution to the knowledge of the Linyphiidae of the Maghreb. Part XII. Miscellaneous erigonine genera and additional records (Araneae: Linyphiidae: Erigoninae). Bulletin &amp; Annales de la Societe Entomologique de Belgique, 143, 117 - 163." type="journal article" year="2007">Bosmans 2007: 139</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="202B1094FFEC5D4C9A32FC721E5CA71C" author="Bouseksou, S. &amp; Kherbouche-Abrous, O. &amp; Beladjal, L." box="[771,1050,1833,1857]" page="259" pageId="13" pageNumber="314" year="2015">
<bibRefCitation id="C51B4B74FFEC5D4C9A32FC721E5CA71C" author="Bouseksou, S. &amp; Kherbouche-Abrous, O. &amp; Beladjal, L." box="[771,1050,1833,1857]" pageId="13" pageNumber="314" pagination="257 - 264" refId="ref19982" refString="Bouseksou, S., Kherbouche-Abrous, O. &amp; Beladjal, L. (2015) Ecology of Araneae (Arthropoda, Arachnida) populations in two agroecosystems: wheat and oilseed rape in the Mitidja plain (Algeria). Vie et milieu - Life and Environment, 65 (4), 257 - 264." type="journal article" year="2015">
Bouseksou
<emphasis id="93FEEA97FFEC5D4C9A49FC7119EAA71C" box="[888,940,1833,1857]" italics="true" pageId="13" pageNumber="314">et al.</emphasis>
2015: 259
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFEC5D4C99A6FC341BDCA78F" pageId="13" pageNumber="314" type="materials_examined">
<paragraph id="A1353685FFEC5D4C99A6FC341EA1A7D7" blockId="13.[151,1436,1903,2002]" box="[151,1255,1903,1930]" pageId="13" pageNumber="314">
<emphasis id="93FEEA97FFEC5D4C99A6FC341BC5A7D4" bold="true" box="[151,387,1903,1929]" pageId="13" pageNumber="314">Material examined:</emphasis>
BERRIANE: Sidi Mbarek, hand collecting,
<specimenCount id="B78CFD0CFFEC5D4C9A49FC3419E4A7D7" box="[888,930,1903,1930]" count="1" pageId="13" pageNumber="314" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFEC5D4C9A81FC341E61A7D4" box="[944,1063,1903,1929]" pageId="13" pageNumber="314" value="2021-02-18">18.II.2021</date>
;
<materialsCitation id="11E23CD8FFEC5D4C9D02FC341EA5A7D7" box="[1075,1251,1903,1930]" collectingDate="2021-02-25" isTailEnumeration="true" pageId="13" pageNumber="314" specimenCount="2" specimenCount-male="2">
<specimenCount id="B78CFD0CFFEC5D4C9D02FC341E19A7D7" box="[1075,1119,1903,1930]" count="2" pageId="13" pageNumber="314" type="male">2 ♂</specimenCount>
,
<date id="D5341045FFEC5D4C9D5AFC341EA5A7D7" box="[1131,1251,1903,1930]" pageId="13" pageNumber="314" value="2021-02-25">
<collectingDate id="C570E9ADFFEC5D4C9D5AFC341EA5A7D7" box="[1131,1251,1903,1930]" pageId="13" pageNumber="314" value="2021-02-25">25.II.2021</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="A1353685FFEC5D4C99F6FCC81BDCA78F" blockId="13.[151,1436,1903,2002]" pageId="13" pageNumber="314">
<emphasis id="93FEEA97FFEC5D4C99F6FCC81864A7F0" bold="true" box="[199,546,1939,1965]" pageId="13" pageNumber="314">
Previous records in
<collectingCountry id="D99D7615FFEC5D4C98F1FCC8185BA7F0" box="[448,541,1939,1965]" name="Algeria" pageId="13" pageNumber="314">Algeria</collectingCountry>
:
</emphasis>
Without precise locality,
<collectingRegion id="634EF867FFEC5D4C9A60FCC819D6A7F0" box="[849,912,1939,1965]" country="Algeria" name="Blida" pageId="13" pageNumber="314">Blida</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C9A91FCC819ACA7F0" box="[928,1002,1939,1965]" country="Algeria" name="M'Sila" pageId="13" pageNumber="314">Msila</collectingRegion>
,
<collectingRegion id="634EF867FFEC5D4C9AC8FCC81E71A7F0" box="[1017,1079,1939,1965]" country="Algeria" name="Saida" pageId="13" pageNumber="314">Saïda</collectingRegion>
(
<bibRefCitation id="C51B4B74FFEC5D4C9D7BFCC81E9DA7F0" author="Simon, E." box="[1098,1243,1939,1965]" pageId="13" pageNumber="314" pagination="309 - 532" refId="ref23547" refString="Simon, E. (1926) n. k. In: Les arachnides de France. Synopsis generale et catalogue des especes francaises de l'ordre des Araneae. Tome VI. 2 e partie. Roret, Paris, pp. 309 - 532." type="book chapter" year="1926">Simon 1926</bibRefCitation>
;
<bibRefCitation id="C51B4B74FFEC5D4C9DDAFCCF1FDEA7F0" author="Bosmans, R." box="[1259,1432,1939,1966]" pageId="13" pageNumber="314" pagination="117 - 163" refId="ref19551" refString="Bosmans, R. (2007) Contribution to the knowledge of the Linyphiidae of the Maghreb. Part XII. Miscellaneous erigonine genera and additional records (Araneae: Linyphiidae: Erigoninae). Bulletin &amp; Annales de la Societe Entomologique de Belgique, 143, 117 - 163." type="journal article" year="2007">Bosmans 2007</bibRefCitation>
,
<bibRefCitation id="C51B4B74FFEC5D4C99A6FCEC1BD4A78F" author="Bouseksou, S. &amp; Kherbouche-Abrous, O. &amp; Beladjal, L." box="[151,402,1975,2002]" pageId="13" pageNumber="314" pagination="257 - 264" refId="ref19982" refString="Bouseksou, S., Kherbouche-Abrous, O. &amp; Beladjal, L. (2015) Ecology of Araneae (Arthropoda, Arachnida) populations in two agroecosystems: wheat and oilseed rape in the Mitidja plain (Algeria). Vie et milieu - Life and Environment, 65 (4), 257 - 264." type="journal article" year="2015">
Bouseksou
<emphasis id="93FEEA97FFEC5D4C9828FCE31B17A78C" box="[281,337,1975,2001]" italics="true" pageId="13" pageNumber="314">et al.</emphasis>
2015
</bibRefCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFEF5D4F99F6FBCD1829A088" pageId="14" pageNumber="315" type="distribution">
<paragraph id="A1353685FFEF5D4F99F6FBCD1FDAA0EC" blockId="14.[151,1436,150,213]" box="[199,1436,150,177]" pageId="14" pageNumber="315">
<emphasis id="93FEEA97FFEF5D4F99F6FBCD1B25A0ED" bold="true" box="[199,355,150,176]" pageId="14" pageNumber="315">Distribution:</emphasis>
Europe, North Africa,
<collectingCountry id="D99D7615FFEF5D4F9B54FBCC18F0A0EC" box="[613,694,151,177]" name="Turkey" pageId="14" pageNumber="315">Turkey</collectingCountry>
, Caucasus,
<collectingCountry id="D99D7615FFEF5D4F9A08FBCC19C2A0EC" box="[825,900,151,177]" name="Russia" pageId="14" pageNumber="315">Russia</collectingCountry>
(Europe to Far East),
<collectingCountry id="D99D7615FFEF5D4F9D4FFBCC1F42A0EC" box="[1150,1284,151,177]" name="Kazakhstan" pageId="14" pageNumber="315">Kazakhstan</collectingCountry>
,
<collectingCountry id="D99D7615FFEF5D4F9C3EFBCC1F79A0EC" box="[1295,1343,151,177]" name="Iran" pageId="14" pageNumber="315">Iran</collectingCountry>
, Central
</paragraph>
<paragraph id="A1353685FFEF5D4F99A6FBE01829A088" blockId="14.[151,1436,150,213]" box="[151,623,187,213]" pageId="14" pageNumber="315">
Asia,
<collectingCountry id="D99D7615FFEF5D4F99E8FBE01B5AA088" box="[217,284,187,213]" name="China" pageId="14" pageNumber="315">China</collectingCountry>
(
<bibRefCitation id="C51B4B74FFEF5D4F981BFBE01825A088" author="World Spider Catalog" box="[298,611,187,213]" pageId="14" pageNumber="315" refId="ref24111" refString="World Spider Catalog (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern, Bern. Available from: https: // wsc. nmbe. ch / (accessed 1 June 2024) https: // doi. org / 10.24436 / 2" type="book" year="2024">World Spider Catalog 2024</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,430 @@
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<emphasis id="93FEEA97FFEE5D4E99A6F9391B03A227" box="[151,325,610,634]" italics="true" pageId="15" pageNumber="316">Pardosa proxima</emphasis>
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<treatmentCitation id="202B1094FFEE5D4E9863F9391BB2A227" author="Simon, E." box="[338,500,610,634]" page="85" pageId="15" pageNumber="316" year="1899">
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</treatmentCitation>
;
<treatmentCitation id="202B1094FFEE5D4E98CFF9391887A227" author="Denis, J." box="[510,705,610,634]" page="1053" pageId="15" pageNumber="316" year="1937">
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;
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<bibRefCitation id="C51B4B74FFEE5D4E9BF9F9391E69A227" author="Abrous-Kherbouche, O. &amp; Jocque, R. &amp; Maelfait, J. - P." box="[712,1071,610,634]" pageId="15" pageNumber="316" pagination="71 - 90" refId="ref18469" refString="Abrous-Kherbouche, O., Jocque, R. &amp; Maelfait, J. - P. (1997) Les effets du paturage intensif sur l'araneofaune dans la region de Tala-Guilef (Parc National du Djurdjura, Algerie). Annales de la Societe Entomologique de Belgique, 133, 71 - 90." type="journal article" year="1997">
Abrous-Kherbouche
<emphasis id="93FEEA97FFEE5D4E9AACF9381989A227" box="[925,975,610,634]" italics="true" pageId="15" pageNumber="316">et al.</emphasis>
1997: 79
</bibRefCitation>
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;
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<bibRefCitation id="C51B4B74FFEE5D4E9D08F9391F0BA227" author="Bouseksou, S. &amp; Kherbouche-Abrous, O. &amp; Beladjal, L." box="[1081,1357,610,634]" pageId="15" pageNumber="316" pagination="257 - 264" refId="ref19982" refString="Bouseksou, S., Kherbouche-Abrous, O. &amp; Beladjal, L. (2015) Ecology of Araneae (Arthropoda, Arachnida) populations in two agroecosystems: wheat and oilseed rape in the Mitidja plain (Algeria). Vie et milieu - Life and Environment, 65 (4), 257 - 264." type="journal article" year="2015">
Bouseksou
<emphasis id="93FEEA97FFEE5D4E9D9FF9381EA6A227" box="[1198,1248,610,634]" italics="true" pageId="15" pageNumber="316">et al.</emphasis>
2015: 260
</bibRefCitation>
</treatmentCitation>
;
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<bibRefCitation id="C51B4B74FFEE5D4E9C67F9391B2FA2C4" author="Touchi, W. &amp; Kherbouche-Abrous, O. &amp; Saadi, A. &amp; Beladjal, L." pageId="15" pageNumber="316" pagination="269 - 282" refId="ref23866" refString="Touchi, W., Kherbouche-Abrous, O., Saadi, A. &amp; Beladjal, L. (2018) Spider communities (Arthropoda, Araneae) in different pine forests of Zeralda game reserve (Algiers, Algeria): Taxonomy and biodiversity. Revue d'ecologie (Terre et vie), 73 (3), 269 - 282. https: // doi. org / 10.3406 / revec. 2018.1934" type="journal article" year="2018">
Touchi
<emphasis id="93FEEA97FFEE5D4E99F6F9D91ABCA2C4" box="[199,250,641,665]" italics="true" pageId="15" pageNumber="316">et al.</emphasis>
2018: 277
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="202B1094FFEE5D4E9845F9DA1835A2C4" author="Boucherit, S. &amp; Bouragba, N. &amp; Douaoui, A. &amp; Bosmans, R." box="[372,627,641,665]" page="63" pageId="15" pageNumber="316" year="2020">
<bibRefCitation id="C51B4B74FFEE5D4E9845F9DA1835A2C4" author="Boucherit, S. &amp; Bouragba, N. &amp; Douaoui, A. &amp; Bosmans, R." box="[372,627,641,665]" pageId="15" pageNumber="316" pagination="60 - 70" refId="ref19840" refString="Boucherit, S., Bouragba, N., Douaoui, A. &amp; Bosmans, R. (2020) Araignees de la plaine alluviale du Haut-Cheliff (Algerie), avec la citation de quelques especes nouvelles ou rares pour l'Algerie (Araneae: Gnaphosidae, Miturgidae). Nieuwsbrief van de Belgische Arachnologische Vereniging, 35 (1 - 2), 60 - 70." type="journal article" year="2020">
Boucherit
<emphasis id="93FEEA97FFEE5D4E98EFF9D91857A2C4" box="[478,529,641,665]" italics="true" pageId="15" pageNumber="316">et al.</emphasis>
2020: 63
</bibRefCitation>
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.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFEE5D4E99A6F99C1ECDA4CC" pageId="15" pageNumber="316" type="materials_examined">
<paragraph id="A1353685FFEE5D4E99A6F99C1B06A417" blockId="15.[151,1436,711,1241]" pageId="15" pageNumber="316">
<emphasis id="93FEEA97FFEE5D4E99A6F99C1BC8A2BC" bold="true" box="[151,398,711,737]" pageId="15" pageNumber="316">Material examined:</emphasis>
BERRIANE: Balouh, hand collecting,
<specimenCount id="B78CFD0CFFEE5D4E9A51F99C19D6A2BF" box="[864,912,711,738]" count="1" pageId="15" pageNumber="316" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFEE5D4E9A90F99C1E4DA2BC" box="[929,1035,711,737]" pageId="15" pageNumber="316" value="2021-02-04">4.II.2021</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9D2AF9931E0AA2BF" box="[1051,1100,712,738]" count="7" pageId="15" pageNumber="316" type="male">7 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFEE5D4E9D6DF99C1ECDA2BF" box="[1116,1163,711,738]" count="8" pageId="15" pageNumber="316" type="female">8 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E9DACF99C1F53A2BC" box="[1181,1301,711,737]" pageId="15" pageNumber="316" value="2021-02-12">12.II.2021</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9C14F99C1F10A2BF" box="[1317,1366,711,738]" count="1" pageId="15" pageNumber="316" type="male">1 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFEE5D4E9C56F99C1FD3A2BF" box="[1383,1429,711,738]" count="1" pageId="15" pageNumber="316" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E99A6F9B01B49A358" box="[151,271,747,773]" pageId="15" pageNumber="316" value="2021-02-16">16.II.2021</date>
;
<specimenCount id="B78CFD0CFFEE5D4E982CF9B01B0AA35B" box="[285,332,747,774]" count="1" pageId="15" pageNumber="316" type="male">1 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFEE5D4E986AF9B01BC1A35B" box="[347,391,747,774]" count="2" pageId="15" pageNumber="316" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E98A9F9B01856A358" box="[408,528,747,773]" pageId="15" pageNumber="316" value="2021-02-19">19.II.2021</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9B2FF9B0180CA35B" box="[542,586,747,774]" count="1" pageId="15" pageNumber="316" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E9B6AF9B01895A358" box="[603,723,747,773]" pageId="15" pageNumber="316" value="2021-02-26">26.II.2021</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9BD0F9B01948A35B" box="[737,782,747,774]" count="1" pageId="15" pageNumber="316" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E9A2FF9B019D7A35B" box="[798,913,747,774]" pageId="15" pageNumber="316" value="2021-03-05">5.III.2021</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9A91F9B0198AA35B" box="[928,972,747,774]" count="1" pageId="15" pageNumber="316" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E9AECF9B01E18A358" box="[989,1118,747,773]" pageId="15" pageNumber="316" value="2021-03-19">19.III.2021</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9D5DF9B01EDDA35B" box="[1132,1179,747,774]" count="2" pageId="15" pageNumber="316" type="male">2 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFEE5D4E9D9BF9B01E90A35B" box="[1194,1238,747,774]" count="4" pageId="15" pageNumber="316" type="female">4 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E9DD6F9B01F1CA35B" box="[1255,1370,747,774]" pageId="15" pageNumber="316" value="2021-03-05">5.III.2021</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9C59F9B01FD1A35B" box="[1384,1431,747,774]" count="1" pageId="15" pageNumber="316" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFEE5D4E99A6F8541B5FA374" box="[151,281,783,809]" pageId="15" pageNumber="316" value="2021-03-12">12.III.2021</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9814F8541B14A377" box="[293,338,783,810]" count="1" pageId="15" pageNumber="316" type="male">1 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFEE5D4E986FF8541BCFA377" box="[350,393,783,810]" count="4" pageId="15" pageNumber="316" type="female">4 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E98A9F854185CA374" box="[408,538,783,809]" pageId="15" pageNumber="316" value="2021-03-26">26.III.2021</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9B17F8541815A377" box="[550,595,783,810]" count="1" pageId="15" pageNumber="316" type="male">1 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFEE5D4E9B6EF84B18CCA377" box="[607,650,784,810]" count="7" pageId="15" pageNumber="316" type="female">7 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E9BA8F854194DA374" box="[665,779,783,809]" pageId="15" pageNumber="316" value="2021-04-03">3.IV.2021</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9A26F8541905A377" box="[791,835,783,810]" count="9" pageId="15" pageNumber="316" type="male">9 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFEE5D4E9A61F854193DA377" box="[848,891,783,810]" count="6" pageId="15" pageNumber="316" type="female">6 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E9ABBF8541E4FA374" box="[906,1033,783,809]" pageId="15" pageNumber="316" value="2021-04-10">10.IV.2021</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9D27F84B1E04A377" box="[1046,1090,784,810]" count="7" pageId="15" pageNumber="316" type="male">7 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFEE5D4E9D7EF8541E3CA377" box="[1103,1146,783,810]" count="9" pageId="15" pageNumber="316" type="female">9 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E9DB8F8541F4EA374" box="[1161,1288,783,809]" pageId="15" pageNumber="316" value="2021-04-17">17.IV.2021</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9C25F8541F09A377" box="[1300,1359,783,810]" count="12" pageId="15" pageNumber="316" type="male">12 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFEE5D4E9C6DF8541FD3A377" box="[1372,1429,783,810]" count="13" pageId="15" pageNumber="316" type="female">13 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E99A6F8681B51A310" box="[151,279,819,845]" pageId="15" pageNumber="316" value="2021-04-24">24.IV.2021</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9815F8681B14A313" box="[292,338,819,846]" count="1" pageId="15" pageNumber="316" type="male">1 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFEE5D4E9851F8681BCBA313" box="[352,397,819,846]" count="8" pageId="15" pageNumber="316" type="female">8 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E98ACF868185AA310" box="[413,540,819,845]" pageId="15" pageNumber="316" value="2021-04-26">26.IV.2021</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9B1BF8681823A313" box="[554,613,819,846]" count="11" pageId="15" pageNumber="316" type="male">11 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFEE5D4E9B42F86F18D9A313" box="[627,671,820,846]" count="7" pageId="15" pageNumber="316" type="female">7 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E9B9EF8681968A310" box="[687,814,819,845]" pageId="15" pageNumber="316" value="2021-04-30">30.IV.2021</date>
, pitfall traps,
<specimenCount id="B78CFD0CFFEE5D4E9AFFF86819BCA313" box="[974,1018,819,846]" count="4" pageId="15" pageNumber="316" type="female">4 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E9D3BF8681EC7A310" box="[1034,1153,819,845]" pageId="15" pageNumber="316" value="2021-02-12">12.II.2021</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9DA1F8681EF8A313" box="[1168,1214,819,846]" count="2" pageId="15" pageNumber="316" type="male">2 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFEE5D4E9DFDF8681EBEA313" box="[1228,1272,819,846]" count="5" pageId="15" pageNumber="316" type="female">5 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E9C39F8681FC6A310" box="[1288,1408,819,845]" pageId="15" pageNumber="316" value="2021-02-19">19.II.2021</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9CBFF8681AE8A32F" count="2" pageId="15" pageNumber="316" type="male">2 ♂</specimenCount>
,
<date id="D5341045FFEE5D4E998DF80C1B72A32C" box="[188,308,855,881]" pageId="15" pageNumber="316" value="2021-02-26">26.II.2021</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9873F80C1B29A32F" box="[322,367,855,882]" count="1" pageId="15" pageNumber="316" type="male">1 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFEE5D4E984CF80C1BEFA32F" box="[381,425,855,882]" count="1" pageId="15" pageNumber="316" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E9888F80C186AA32F" box="[441,556,855,882]" pageId="15" pageNumber="316" value="2021-03-05">5.III.2021</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9B0BF80C182EA32F" box="[570,616,855,882]" count="1" pageId="15" pageNumber="316" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFEE5D4E9B47F80C18B1A32C" box="[630,759,855,881]" pageId="15" pageNumber="316" value="2021-03-12">12.III.2021</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9A34F80C1975A32F" box="[773,819,855,882]" count="1" pageId="15" pageNumber="316" type="male">1 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFEE5D4E9A70F80C192BA32F" box="[833,877,855,882]" count="1" pageId="15" pageNumber="316" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E9A4DF80C19B8A32C" box="[892,1022,855,881]" pageId="15" pageNumber="316" value="2021-03-26">26.III.2021</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9D3AF80C1E71A32F" box="[1035,1079,855,882]" count="3" pageId="15" pageNumber="316" type="female">3 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E9D76F80C1EFFA32C" box="[1095,1209,855,881]" pageId="15" pageNumber="316" value="2021-04-03">3.IV.2021</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9DF7F80C1EB2A32F" box="[1222,1268,855,882]" count="3" pageId="15" pageNumber="316" type="male">3 ♂</specimenCount>
,
<date id="D5341045FFEE5D4E9C33F80C1FC7A32C" box="[1282,1409,855,881]" pageId="15" pageNumber="316" value="2021-04-10">10.IV.2021</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9CBFF80C1AE8A3CB" count="1" pageId="15" pageNumber="316" type="male">1 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFEE5D4E998DF8201AAEA3CB" box="[188,232,891,918]" count="2" pageId="15" pageNumber="316" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E99C6F8201B30A3C8" box="[247,374,891,917]" pageId="15" pageNumber="316" value="2021-04-17">17.IV.2021</date>
;
<specimenCount id="B78CFD0CFFEE5D4E98B5F8201BE9A3CB" box="[388,431,891,918]" count="1" pageId="15" pageNumber="316" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E988EF8201878A3C8" box="[447,574,891,917]" pageId="15" pageNumber="316" value="2021-04-24">24.IV.2021</date>
.
<materialsCitation id="11E23CD8FFEE5D4E9B7AF82019D1A3C8" box="[587,919,891,917]" location="Sidi Mbarek" pageId="15" pageNumber="316" specimenCount="1">
<location id="A455605EFFEE5D4E9B7AF820189CA3C8" LSID="urn:lsid:plazi:treatment:29238793FFEE5D4E99A6F9461881A484:A455605EFFEE5D4E9B7AF820189CA3C8" box="[587,730,891,917]" name="Sidi Mbarek" pageId="15" pageNumber="316">Sidi Mbarek</location>
, hand collecting
</materialsCitation>
,
<specimenCount id="B78CFD0CFFEE5D4E9A95F8201994A3CB" box="[932,978,891,918]" count="4" pageId="15" pageNumber="316" type="male">4 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFEE5D4E9AD1F8271E4DA3CB" box="[992,1035,892,918]" count="7" pageId="15" pageNumber="316" type="female">7 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E9D2AF8201EC3A3C8" box="[1051,1157,891,917]" pageId="15" pageNumber="316" value="2021-02-06">6.II.2021</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9DA3F8201E86A3CB" box="[1170,1216,891,918]" count="1" pageId="15" pageNumber="316" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFEE5D4E9DFFF8201F03A3C8" box="[1230,1349,891,917]" pageId="15" pageNumber="316" value="2021-02-18">18.II.2021</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9C62F8201FC6A3CB" box="[1363,1408,891,918]" count="1" pageId="15" pageNumber="316" type="male">1 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFEE5D4E9CBFF8201AEAA3E7" count="2" pageId="15" pageNumber="316" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E998AF8C41B75A3E7" box="[187,307,927,954]" pageId="15" pageNumber="316" value="2021-02-25">25.II.2021</date>
;
<specimenCount id="B78CFD0CFFEE5D4E980EF8C41B2DA3E7" box="[319,363,927,954]" count="1" pageId="15" pageNumber="316" type="male">1 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFEE5D4E9849F8C41BE4A3E7" box="[376,418,927,954]" count="2" pageId="15" pageNumber="316" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E9880F8C41862A3E4" box="[433,548,927,953]" pageId="15" pageNumber="316" value="2021-03-04">4.III.2021</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9B01F8C4181BA3E7" box="[560,605,927,954]" count="1" pageId="15" pageNumber="316" type="male">1 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFEE5D4E9B58F8C418D2A3E7" box="[617,660,927,954]" count="2" pageId="15" pageNumber="316" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E9B93F8C41965A3E4" box="[674,803,927,953]" pageId="15" pageNumber="316" value="2021-03-18">18.III.2021</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9A01F8C4191AA3E7" box="[816,860,927,954]" count="1" pageId="15" pageNumber="316" type="male">1 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFEE5D4E9A58F8C419D5A3E7" box="[873,915,927,954]" count="3" pageId="15" pageNumber="316" type="female">3 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E9A93F8C41E54A3E4" box="[930,1042,927,953]" pageId="15" pageNumber="316" value="2021-04-02">2.IV.2021</date>
, pitfall traps,
<specimenCount id="B78CFD0CFFEE5D4E9D9CF8C41E9EA3E7" box="[1197,1240,927,954]" count="1" pageId="15" pageNumber="316" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E9DD6F8C41F18A3E4" box="[1255,1374,927,953]" pageId="15" pageNumber="316" value="2021-02-18">18.II.2021</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9C5BF8C41FD3A3E7" box="[1386,1429,927,954]" count="1" pageId="15" pageNumber="316" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E99A6F8981B51A380" box="[151,279,963,989]" pageId="15" pageNumber="316" value="2021-03-11">11.III.2021</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9814F8981B16A383" box="[293,336,963,990]" count="1" pageId="15" pageNumber="316" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E9851F8981BA7A383" box="[352,481,963,990]" pageId="15" pageNumber="316" value="2021-03-25">25.III.2021</date>
;
<specimenCount id="B78CFD0CFFEE5D4E98DFF898185CA383" box="[494,538,963,990]" count="2" pageId="15" pageNumber="316" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E9B1BF89818DDA380" box="[554,667,963,989]" pageId="15" pageNumber="316" value="2021-04-02">2.IV.2021</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9B99F8981890A383" box="[680,726,963,990]" count="1" pageId="15" pageNumber="316" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFEE5D4E9BD2F8981913A380" box="[739,853,963,989]" pageId="15" pageNumber="316" value="2021-04-09">9.IV.2021</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9A53F89819D6A383" box="[866,912,963,990]" count="1" pageId="15" pageNumber="316" type="male">1 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFEE5D4E9AACF898198FA383" box="[925,969,963,990]" count="1" pageId="15" pageNumber="316" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E9AE8F8981E1EA380" box="[985,1112,963,989]" pageId="15" pageNumber="316" value="2021-04-16">16.IV.2021</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9D54F8981ED5A383" box="[1125,1171,963,990]" count="1" pageId="15" pageNumber="316" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFEE5D4E9D91F8981F66A380" box="[1184,1312,963,989]" pageId="15" pageNumber="316" value="2021-04-23">23.IV.2021</date>
.
<materialsCitation id="11E23CD8FFEE5D4E9C1DF8981834A45C" location="Grazil" pageId="15" pageNumber="316" specimenCount="1">
DHAYET BENDHAHOUA:
<location id="A455605EFFEE5D4E985CF8BC1BF0A45C" LSID="urn:lsid:plazi:treatment:29238793FFEE5D4E99A6F9461881A484:A455605EFFEE5D4E985CF8BC1BF0A45C" box="[365,438,999,1025]" name="Grazil" pageId="15" pageNumber="316">Grazil</location>
, hand collecting
</materialsCitation>
,
<specimenCount id="B78CFD0CFFEE5D4E9B4FF8BC18EAA45F" box="[638,684,999,1026]" count="1" pageId="15" pageNumber="316" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFEE5D4E9B88F8BC195FA45F" box="[697,793,999,1026]" pageId="15" pageNumber="316" value="2022-01-05">5.I.2022</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9A17F8BC1917A45F" box="[806,849,999,1026]" count="1" pageId="15" pageNumber="316" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E9A50F8BC199FA45F" box="[865,985,999,1026]" pageId="15" pageNumber="316" value="2022-02-25">25.II.2022</date>
.
<materialsCitation id="11E23CD8FFEE5D4E9AD4F8B31B01A478" collectorName="El Atteuf" location="Palm" pageId="15" pageNumber="316" specimenCount="1">
EL ATTEUF:
<location id="A455605EFFEE5D4E9DB4F8BC1EF9A45C" LSID="urn:lsid:plazi:treatment:29238793FFEE5D4E99A6F9461881A484:A455605EFFEE5D4E9DB4F8BC1EF9A45C" box="[1157,1215,999,1025]" name="Palm" pageId="15" pageNumber="316">Palm</location>
grove of
<collectorName id="0C7F5353FFEE5D4E9C1FF8BC1FDFA45C" box="[1326,1433,999,1025]" pageId="15" pageNumber="316">El Atteuf</collectorName>
, hand collecting
</materialsCitation>
,
<specimenCount id="B78CFD0CFFEE5D4E9862FF501B38A47B" box="[339,382,1035,1062]" count="2" pageId="15" pageNumber="316" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E98BCFF501846A47B" box="[397,512,1035,1062]" pageId="15" pageNumber="316" value="2015-03-06">6.III.2015</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9B3CFF501871A47B" box="[525,567,1035,1062]" count="2" pageId="15" pageNumber="316" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E9B77FF50188EA47B" box="[582,712,1035,1062]" pageId="15" pageNumber="316" value="2015-03-15">15.III.2015</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9BE5FF5018B9A47B" box="[724,767,1035,1062]" count="1" pageId="15" pageNumber="316" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E9A3FFF501939A47B" box="[782,895,1035,1062]" pageId="15" pageNumber="316" value="2015-04-05">5.IV.2015</date>
, pitfall traps,
<specimenCount id="B78CFD0CFFEE5D4E9D2AFF501E00A47B" box="[1051,1094,1035,1062]" count="2" pageId="15" pageNumber="316" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E9D64FF501E8FA47B" box="[1109,1225,1035,1062]" pageId="15" pageNumber="316" value="2015-03-06">6.III.2015</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9DE4FF501F46A47B" box="[1237,1280,1035,1062]" count="1" pageId="15" pageNumber="316" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E9C3EFF501FC4A47B" box="[1295,1410,1035,1062]" pageId="15" pageNumber="316" value="2015-03-06">6.III.2015</date>
;
<specimenCount id="B78CFD0CFFEE5D4E9CBFFF501AEAA417" count="4" pageId="15" pageNumber="316" type="female">4 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E998BFF741B7AA417" box="[186,316,1071,1098]" pageId="15" pageNumber="316" value="2015-03-25">25.III.2015</date>
.
</paragraph>
<paragraph id="A1353685FFEE5D4E99F6FF081ECDA4CC" blockId="15.[151,1436,711,1241]" pageId="15" pageNumber="316">
<emphasis id="93FEEA97FFEE5D4E99F6FF08185FA430" bold="true" box="[199,537,1107,1133]" pageId="15" pageNumber="316">
Previous records in
<collectingCountry id="D99D7615FFEE5D4E9886FF081852A430" box="[439,532,1107,1133]" name="Algeria" pageId="15" pageNumber="316">Algeria</collectingCountry>
:
</emphasis>
<collectingRegion id="634EF867FFEE5D4E9B2EFF0818D4A430" box="[543,658,1107,1133]" country="Algeria" name="Ain Defla" pageId="15" pageNumber="316">Aïn Defla</collectingRegion>
,
<collectingRegion id="634EF867FFEE5D4E9BACFF081899A430" box="[669,735,1107,1133]" country="Algeria" name="Alger" pageId="15" pageNumber="316">Alger</collectingRegion>
,
<collectingRegion id="634EF867FFEE5D4E9BDAFF08196CA430" box="[747,810,1107,1133]" country="Algeria" name="Blida" pageId="15" pageNumber="316">Blida</collectingRegion>
,
<collectingRegion id="634EF867FFEE5D4E9A06FF081929A430" box="[823,879,1107,1133]" country="Algeria" name="Mila" pageId="15" pageNumber="316">Mila</collectingRegion>
,
<collectingRegion id="634EF867FFEE5D4E9A4BFF0819BFA430" box="[890,1017,1107,1133]" country="Algeria" name="Tizi Ouzou" pageId="15" pageNumber="316">Tizi Ouzou</collectingRegion>
(
<bibRefCitation id="C51B4B74FFEE5D4E9D38FF081ED1A430" author="Simon, E." box="[1033,1175,1107,1133]" pageId="15" pageNumber="316" pagination="82 - 87" refId="ref23337" refString="Simon, E. (1899) Liste des arachnides recueillis en Algerie par M. P. Lesne et descrition d'une espece nouvelle. Bulletin du Museum d'Histoire Naturelle Paris, 5, 82 - 87." type="journal article" year="1899">Simon 1899</bibRefCitation>
;
<bibRefCitation id="C51B4B74FFEE5D4E9D92FF081F70A430" author="Denis, J." box="[1187,1334,1107,1133]" pageId="15" pageNumber="316" pagination="1027 - 1060" refId="ref20639" refString="Denis, J. (1937 b) On a collection of spiders from Algeria. Proceedings of the Zoological Society of London, 106 (4), 1027 - 1060. https: // doi. org / 10.1111 / j. 1469 - 7998.1936. tb 06301. x" type="journal article" year="1937">Denis 1937b</bibRefCitation>
,
<bibRefCitation id="C51B4B74FFEE5D4E9C71FF081BE7A4CC" author="Abrous-Kherbouche, O. &amp; Jocque, R. &amp; Maelfait, J. - P." pageId="15" pageNumber="316" pagination="71 - 90" refId="ref18469" refString="Abrous-Kherbouche, O., Jocque, R. &amp; Maelfait, J. - P. (1997) Les effets du paturage intensif sur l'araneofaune dans la region de Tala-Guilef (Parc National du Djurdjura, Algerie). Annales de la Societe Entomologique de Belgique, 133, 71 - 90." type="journal article" year="1997">
Abrous-Kherbouche
<emphasis id="93FEEA97FFEE5D4E9816FF231B19A4CC" box="[295,351,1143,1169]" italics="true" pageId="15" pageNumber="316">et al.</emphasis>
1997
</bibRefCitation>
;
<bibRefCitation id="C51B4B74FFEE5D4E989CFF2C18EFA4CF" author="Bouseksou, S. &amp; Kherbouche-Abrous, O. &amp; Beladjal, L." box="[429,681,1143,1170]" pageId="15" pageNumber="316" pagination="257 - 264" refId="ref19982" refString="Bouseksou, S., Kherbouche-Abrous, O. &amp; Beladjal, L. (2015) Ecology of Araneae (Arthropoda, Arachnida) populations in two agroecosystems: wheat and oilseed rape in the Mitidja plain (Algeria). Vie et milieu - Life and Environment, 65 (4), 257 - 264." type="journal article" year="2015">
Bouseksou
<emphasis id="93FEEA97FFEE5D4E9B1EFF231821A4CC" box="[559,615,1143,1169]" italics="true" pageId="15" pageNumber="316">et al.</emphasis>
2015
</bibRefCitation>
;
<bibRefCitation id="C51B4B74FFEE5D4E9B85FF2C19C4A4CC" author="Touchi, W. &amp; Kherbouche-Abrous, O. &amp; Saadi, A. &amp; Beladjal, L." box="[692,898,1143,1169]" pageId="15" pageNumber="316" pagination="269 - 282" refId="ref23866" refString="Touchi, W., Kherbouche-Abrous, O., Saadi, A. &amp; Beladjal, L. (2018) Spider communities (Arthropoda, Araneae) in different pine forests of Zeralda game reserve (Algiers, Algeria): Taxonomy and biodiversity. Revue d'ecologie (Terre et vie), 73 (3), 269 - 282. https: // doi. org / 10.3406 / revec. 2018.1934" type="journal article" year="2018">
Touchi
<emphasis id="93FEEA97FFEE5D4E9A39FF231907A4CC" box="[776,833,1143,1169]" italics="true" pageId="15" pageNumber="316">et al.</emphasis>
2018
</bibRefCitation>
;
<bibRefCitation id="C51B4B74FFEE5D4E9ABFFF2C1E39A4CC" author="Boucherit, S. &amp; Bouragba, N. &amp; Douaoui, A. &amp; Bosmans, R." box="[910,1151,1143,1169]" pageId="15" pageNumber="316" pagination="60 - 70" refId="ref19840" refString="Boucherit, S., Bouragba, N., Douaoui, A. &amp; Bosmans, R. (2020) Araignees de la plaine alluviale du Haut-Cheliff (Algerie), avec la citation de quelques especes nouvelles ou rares pour l'Algerie (Araneae: Gnaphosidae, Miturgidae). Nieuwsbrief van de Belgische Arachnologische Vereniging, 35 (1 - 2), 60 - 70." type="journal article" year="2020">
Boucherit
<emphasis id="93FEEA97FFEE5D4E9D32FF231E7AA4CC" box="[1027,1084,1143,1169]" italics="true" pageId="15" pageNumber="316">et al.</emphasis>
2020
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFEE5D4E99F6FFC01881A484" pageId="15" pageNumber="316" type="distribution">
<paragraph id="A1353685FFEE5D4E99F6FFC01881A484" blockId="15.[151,1436,711,1241]" pageId="15" pageNumber="316">
<emphasis id="93FEEA97FFEE5D4E99F6FFC01B25A4E8" bold="true" box="[199,355,1179,1205]" pageId="15" pageNumber="316">Distribution:</emphasis>
Macaronesia, northern Africa, Europe, Caucasus,
<collectingCountry id="D99D7615FFEE5D4E9AA7FFC019A7A4E8" box="[918,993,1179,1205]" name="Russia" pageId="15" pageNumber="316">Russia</collectingCountry>
(Europe to Far East),
<collectingCountry id="D99D7615FFEE5D4E9DE9FFC01F18A4E8" box="[1240,1374,1179,1205]" name="Kazakhstan" pageId="15" pageNumber="316">Kazakhstan</collectingCountry>
,
<collectingCountry id="D99D7615FFEE5D4E9C59FFC71FDEA4EB" box="[1384,1432,1180,1206]" name="Iran" pageId="15" pageNumber="316">Iran</collectingCountry>
, Central Asia,
<collectingCountry id="D99D7615FFEE5D4E9801FFE41B35A484" box="[304,371,1215,1241]" name="China" pageId="15" pageNumber="316">China</collectingCountry>
(
<bibRefCitation id="C51B4B74FFEE5D4E98B3FFE418FDA484" author="World Spider Catalog" box="[386,699,1215,1241]" pageId="15" pageNumber="316" refId="ref24111" refString="World Spider Catalog (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern, Bern. Available from: https: // wsc. nmbe. ch / (accessed 1 June 2024) https: // doi. org / 10.24436 / 2" type="book" year="2024">World Spider Catalog 2024</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,146 @@
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<mods:title id="51F75CD8BB682C4FA31301114CB814CA">Spiders of arid lands: The Ghardaïa region (Northern Sahara) with seven new records for Algeria</mods:title>
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<mods:namePart id="A68FC8DD2EFD3F5CC99DF31DF7824569">Alioua, Youcef</mods:namePart>
<mods:affiliation id="244FE09EECA3DB6834EF5B9EFB2380D0">Laboratoire de Valorisation et Conservation des Ecosystèmes Arides (LVCEA). Faculty of Natural, Life and Earth Sciences, University of Ghardaïa, BP 455, 47000 Ghardaïa, Algeria</mods:affiliation>
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<mods:namePart id="6F97B92E2A64BFEDD3A7843EAC66A52F">Bosmans, Robert</mods:namePart>
<mods:affiliation id="5C6BCF7BC37B43F633972419BE8C5264">Terrestrial Ecology Unit, Ledeganckstraat 35, B- 9000 Gent, Belgium</mods:affiliation>
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<treatment id="29238793FFEE5D4E99A6FBCC1EDEA191" LSID="urn:lsid:plazi:treatment:29238793FFEE5D4E99A6FBCC1EDEA191" httpUri="http://treatment.plazi.org/id/29238793FFEE5D4E99A6FBCC1EDEA191" lastPageNumber="316" pageId="15" pageNumber="316">
<subSubSection id="E990650EFFEE5D4E99A6FBCC1849A0EC" box="[151,527,151,178]" pageId="15" pageNumber="316" type="nomenclature">
<paragraph id="A1353685FFEE5D4E99A6FBCC1849A0EC" blockId="15.[151,527,151,178]" box="[151,527,151,178]" pageId="15" pageNumber="316">
<heading id="FA7D81E9FFEE5D4E99A6FBCC1849A0EC" bold="true" box="[151,527,151,178]" fontSize="11" level="1" pageId="15" pageNumber="316" reason="1">
<taxonomicName id="668A4D06FFEE5D4E99A6FBCC1849A0EC" ID-CoL="4DJR7" authority="Roewer, 1959" authorityName="Roewer" authorityYear="1959" box="[151,527,151,178]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="gefsana">
<emphasis id="93FEEA97FFEE5D4E99A6FBCC1849A0EC" bold="true" box="[151,527,151,178]" pageId="15" pageNumber="316">
<emphasis id="93FEEA97FFEE5D4E99A6FBCC1B27A0EC" bold="true" box="[151,353,151,178]" italics="true" pageId="15" pageNumber="316">Pardosa gefsana</emphasis>
<bibRefCitation id="C51B4B74FFEE5D4E9859FBCC1849A0EC" author="Roewer, C. F." box="[360,527,151,177]" pageId="15" pageNumber="316" pagination="1 - 518" refId="ref22907" refString="Roewer, C. F. (1959) Araneae Lycosaeformia II (Lycosidae). Exploration du Parc National de l'Upemba, Mission G. F. de Witte, 55, 1 - 518." type="journal article" year="1959">Roewer, 1959</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFEE5D4E99A6FB8719D8A0AE" box="[151,926,219,244]" pageId="15" pageNumber="316" type="reference_group">
<paragraph id="A1353685FFEE5D4E99A6FB8719D8A0AE" blockId="15.[151,926,219,244]" box="[151,926,219,244]" pageId="15" pageNumber="316">
<treatmentCitationGroup id="819A11ABFFEE5D4E99A6FB8719D8A0AE" box="[151,926,219,244]" pageId="15" pageNumber="316">
<taxonomicName id="668A4D06FFEE5D4E99A6FB871B07A0A9" ID-CoL="4DJR7" authorityName="Roewer" authorityYear="1959" box="[151,321,220,244]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="gefsana">
<emphasis id="93FEEA97FFEE5D4E99A6FB871B07A0A9" box="[151,321,220,244]" italics="true" pageId="15" pageNumber="316">Pardosa gefsana</emphasis>
</taxonomicName>
;
<treatmentCitation id="202B1094FFEE5D4E987CFB8718E5A0A9" author="Alderweireldt, M. &amp; Jocque, R." box="[333,675,220,244]" page="89" pageId="15" pageNumber="316" year="1992">
<bibRefCitation id="C51B4B74FFEE5D4E987CFB8718E5A0A9" author="Alderweireldt, M. &amp; Jocque, R." box="[333,675,220,244]" pageId="15" pageNumber="316" pagination="73 - 113" refId="ref18583" refString="Alderweireldt, M. &amp; Jocque, R. (1992) A review of the nebulosa - group of Pardosa Koch 1847 in Africa, a complex with some highly variable species (Araneae Lycosidae). Tropical Zoology, 5 (1), 73 - 113. https: // doi. org / 10.1080 / 03946975.1992.10539183" type="journal article" year="1992">Alderweireldt &amp; Jocqué 1992: 89</bibRefCitation>
</treatmentCitation>
; Alioua
<emphasis id="93FEEA97FFEE5D4E9BC9FB87196DA0A9" box="[760,811,220,244]" italics="true" pageId="15" pageNumber="316">et al.</emphasis>
2022: 167.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFEE5D4E99A6FA7A1EA7A1F5" pageId="15" pageNumber="316" type="materials_examined">
<paragraph id="A1353685FFEE5D4E99A6FA7A1B13A13D" blockId="15.[151,1436,289,460]" pageId="15" pageNumber="316">
<emphasis id="93FEEA97FFEE5D4E99A6FA7A1BC5A166" bold="true" box="[151,387,289,315]" pageId="15" pageNumber="316">Material examined:</emphasis>
EL GUERRARA: Aghzou, hand collecting,
<specimenCount id="B78CFD0CFFEE5D4E9A49FA7919E4A161" box="[888,930,290,316]" count="1" pageId="15" pageNumber="316" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E9A9EFA791E76A161" box="[943,1072,290,316]" pageId="15" pageNumber="316" value="2020-03-11">11.III.2020</date>
. El Amied, hand collecting,
<specimenCount id="B78CFD0CFFEE5D4E9C5AFA791FD1A161" box="[1387,1431,290,316]" count="1" pageId="15" pageNumber="316" type="male">1 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFEE5D4E99A6FA1D1A87A13D" box="[151,193,326,352]" count="1" pageId="15" pageNumber="316" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFEE5D4E99FEFA1D1B16A13D" box="[207,336,326,352]" pageId="15" pageNumber="316" value="2020-03-12">12.III.2020</date>
.
</paragraph>
<paragraph id="A1353685FFEE5D4E99F6FA321EA7A1F5" blockId="15.[151,1436,289,460]" pageId="15" pageNumber="316">
<emphasis id="93FEEA97FFEE5D4E99F6FA321BF2A1DE" bold="true" box="[199,436,361,388]" pageId="15" pageNumber="316">Previous records in</emphasis>
<materialsCitation id="11E23CD8FFEE5D4E988CFA321AA8A1F5" collectorName="El Meniaa" country="Algeria" location="Ain Temouchent" pageId="15" pageNumber="316" specimenCount="1" stateProvince="Ain Temouchent">
<emphasis id="93FEEA97FFEE5D4E988CFA321859A1DE" bold="true" box="[445,543,361,387]" pageId="15" pageNumber="316">
<collectingCountry id="D99D7615FFEE5D4E988CFA32185CA1DE" box="[445,538,361,387]" name="Algeria" pageId="15" pageNumber="316">Algeria</collectingCountry>
:
</emphasis>
<collectingRegion id="634EF867FFEE5D4E9B19FA3118ADA1D9" box="[552,747,362,388]" country="Algeria" name="Ain Temouchent" pageId="15" pageNumber="316">Aïn Témouchent</collectingRegion>
,
<collectingRegion id="634EF867FFEE5D4E9BC8FA31190CA1D9" box="[761,842,362,388]" country="Algeria" name="Bechar" pageId="15" pageNumber="316">Béchar</collectingRegion>
,
<collectingRegion id="634EF867FFEE5D4E9A69FA3119E4A1D9" box="[856,930,362,388]" country="Algeria" name="Bejaia" pageId="15" pageNumber="316">Béjaïa</collectingRegion>
,
<collectingRegion id="634EF867FFEE5D4E9A80FA3119BBA1D9" box="[945,1021,362,388]" country="Algeria" name="Biskra" pageId="15" pageNumber="316">Biskra</collectingRegion>
,
<collectingRegion id="634EF867FFEE5D4E9D3AFA311E1CA1D9" box="[1035,1114,362,388]" country="Algeria" name="Bouira" pageId="15" pageNumber="316">Bouira</collectingRegion>
,
<collectingRegion id="634EF867FFEE5D4E9D58FA311EADA1D9" box="[1129,1259,362,388]" country="Algeria" name="Boumerdes" pageId="15" pageNumber="316">Boumerdès</collectingRegion>
,
<collectingRegion id="634EF867FFEE5D4E9DC8FA311F33A1D9" box="[1273,1397,362,388]" country="Algeria" name="El Bayadh" pageId="15" pageNumber="316">El Bayadh</collectingRegion>
,
<collectorName id="0C7F5353FFEE5D4E9CB2FA311AA8A1F5" pageId="15" pageNumber="316">El Meniaâ</collectorName>
</materialsCitation>
,
<collectingRegion id="634EF867FFEE5D4E99C8FAD51B0CA1F5" box="[249,330,398,424]" country="Algeria" name="El Tarf" pageId="15" pageNumber="316">El Tarf</collectingRegion>
,
<collectingRegion id="634EF867FFEE5D4E9864FAD51BD9A1F5" box="[341,415,398,424]" country="Algeria" name="M'Sila" pageId="15" pageNumber="316">Msila</collectingRegion>
, Timimoun,
<collectingRegion id="634EF867FFEE5D4E9B05FAD518DCA1F5" box="[564,666,398,424]" country="Algeria" name="Tlemcen" pageId="15" pageNumber="316">Tlemcen</collectingRegion>
. (
<bibRefCitation id="C51B4B74FFEE5D4E9B9AFAD519BAA1F5" author="Alderweireldt, M. &amp; Jocque, R." box="[683,1020,398,424]" pageId="15" pageNumber="316" pagination="73 - 113" refId="ref18583" refString="Alderweireldt, M. &amp; Jocque, R. (1992) A review of the nebulosa - group of Pardosa Koch 1847 in Africa, a complex with some highly variable species (Araneae Lycosidae). Tropical Zoology, 5 (1), 73 - 113. https: // doi. org / 10.1080 / 03946975.1992.10539183" type="journal article" year="1992">Alderweireldt &amp; Jocqué 1992</bibRefCitation>
; Alioua
<emphasis id="93FEEA97FFEE5D4E9D68FAD41ED4A1F5" box="[1113,1170,398,424]" italics="true" pageId="15" pageNumber="316">et al.</emphasis>
2022).
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFEE5D4E99F6FAEA1EDEA191" box="[199,1176,433,460]" pageId="15" pageNumber="316" type="distribution">
<paragraph id="A1353685FFEE5D4E99F6FAEA1EDEA191" blockId="15.[151,1436,289,460]" box="[199,1176,433,460]" pageId="15" pageNumber="316">
<emphasis id="93FEEA97FFEE5D4E99F6FAEA1B25A196" bold="true" box="[199,355,433,459]" pageId="15" pageNumber="316">Distribution:</emphasis>
<collectingCountry id="D99D7615FFEE5D4E985BFAE91BEBA191" box="[362,429,434,460]" name="Spain" pageId="15" pageNumber="316">Spain</collectingCountry>
,
<collectingCountry id="D99D7615FFEE5D4E9889FAE91BADA191" box="[440,491,434,460]" name="Italy" pageId="15" pageNumber="316">Italy</collectingCountry>
(
<collectingRegion id="634EF867FFEE5D4E98CBFAE91878A191" box="[506,574,434,460]" country="Italy" name="Sicilia" pageId="15" pageNumber="316">Sicily</collectingRegion>
,
<collectingRegion id="634EF867FFEE5D4E9B78FAE918EDA191" box="[585,683,434,460]" country="Italy" name="Sardegna" pageId="15" pageNumber="316">Sardinia</collectingRegion>
), North Africa (
<bibRefCitation id="C51B4B74FFEE5D4E9A6BFAE91ED6A191" author="World Spider Catalog" box="[858,1168,434,460]" pageId="15" pageNumber="316" refId="ref24111" refString="World Spider Catalog (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern, Bern. Available from: https: // wsc. nmbe. ch / (accessed 1 June 2024) https: // doi. org / 10.24436 / 2" type="book" year="2024">World Spider Catalog 2024</bibRefCitation>
)
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,158 @@
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<mods:title id="A8585E6A6E73B6E640A5B91BCA2265CB">Spiders of arid lands: The Ghardaïa region (Northern Sahara) with seven new records for Algeria</mods:title>
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<mods:namePart id="39940A3087B005486BFA0D9A529A38FB">Alioua, Youcef</mods:namePart>
<mods:affiliation id="8979261D484AAA2E7DAB57DFB1A60EFC">Laboratoire de Valorisation et Conservation des Ecosystèmes Arides (LVCEA). Faculty of Natural, Life and Earth Sciences, University of Ghardaïa, BP 455, 47000 Ghardaïa, Algeria</mods:affiliation>
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<mods:namePart id="17D0E561C30A7B047FB289FB0EF83AB2">Bosmans, Robert</mods:namePart>
<mods:affiliation id="E2E5C9EFCC966291957704F785D6DF99">Terrestrial Ecology Unit, Ledeganckstraat 35, B- 9000 Gent, Belgium</mods:affiliation>
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<treatment id="29238793FFEF5D4F99A6F8071F5FA430" LSID="urn:lsid:plazi:treatment:29238793FFEF5D4F99A6F8071F5FA430" httpUri="http://treatment.plazi.org/id/29238793FFEF5D4F99A6F8071F5FA430" lastPageNumber="315" pageId="14" pageNumber="315">
<subSubSection id="E990650EFFEF5D4F99A6F807186BA32B" box="[151,557,860,886]" pageId="14" pageNumber="315" type="nomenclature">
<paragraph id="A1353685FFEF5D4F99A6F807186BA32B" blockId="14.[151,557,860,886]" box="[151,557,860,886]" pageId="14" pageNumber="315">
<heading id="FA7D81E9FFEF5D4F99A6F807186BA32B" bold="true" box="[151,557,860,886]" fontSize="11" level="1" pageId="14" pageNumber="315" reason="1">
<taxonomicName id="668A4D06FFEF5D4F99A6F807186BA32B" ID-CoL="3DHPR" authority="(Audouin, 1826)" baseAuthorityName="Audouin" baseAuthorityYear="1826" box="[151,557,860,886]" class="Arachnida" family="Lycosidae" genus="Evippa" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="arenaria">
<emphasis id="93FEEA97FFEF5D4F99A6F807186BA32B" bold="true" box="[151,557,860,886]" pageId="14" pageNumber="315">
<emphasis id="93FEEA97FFEF5D4F99A6F8071B1DA32B" bold="true" box="[151,347,860,886]" italics="true" pageId="14" pageNumber="315">Evippa arenaria</emphasis>
(Audouin, 1826)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFEF5D4F99A6F8FA18BEA3E5" box="[151,760,928,953]" pageId="14" pageNumber="315" type="reference_group">
<paragraph id="A1353685FFEF5D4F99A6F8FA18BEA3E5" blockId="14.[151,760,928,953]" box="[151,760,928,953]" pageId="14" pageNumber="315">
<treatmentCitationGroup id="819A11ABFFEF5D4F99A6F8FA18BEA3E5" box="[151,760,928,953]" pageId="14" pageNumber="315">
<taxonomicName id="668A4D06FFEF5D4F99A6F8FA1B7BA3E5" ID-CoL="3DHPR" baseAuthorityName="Audouin" baseAuthorityYear="1826" box="[151,317,929,952]" class="Arachnida" family="Lycosidae" genus="Evippa" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="arenaria">
<emphasis id="93FEEA97FFEF5D4F99A6F8FA1B7BA3E5" box="[151,317,929,952]" italics="true" pageId="14" pageNumber="315">Evippa arenaria</emphasis>
</taxonomicName>
;
<treatmentCitation id="202B1094FFEF5D4F987BF8FB1BA9A3E5" author="Simon, E." box="[330,495,928,953]" page="85" pageId="14" pageNumber="315" year="1899">
<bibRefCitation id="C51B4B74FFEF5D4F987BF8FB1BA9A3E5" author="Simon, E." box="[330,495,928,953]" pageId="14" pageNumber="315" pagination="82 - 87" refId="ref23337" refString="Simon, E. (1899) Liste des arachnides recueillis en Algerie par M. P. Lesne et descrition d'une espece nouvelle. Bulletin du Museum d'Histoire Naturelle Paris, 5, 82 - 87." type="journal article" year="1899">Simon 1899: 85</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="202B1094FFEF5D4F98C9F8FB18B2A3E5" author="Alderweireldt, M." box="[504,756,928,953]" page="363" pageId="14" pageNumber="315" year="1991">
<bibRefCitation id="C51B4B74FFEF5D4F98C9F8FB18B2A3E5" author="Alderweireldt, M." box="[504,756,928,953]" pageId="14" pageNumber="315" pagination="359 - 381" refId="ref18524" refString="Alderweireldt, M. (1991) A revision of the African representatives of the wolf spider genus Evippa Simon, 1882 (Araneae, Lycosidae) with notes on allied species and genera. Journal of Natural History, 25 (2), 359 - 381. https: // doi. org / 10.1080 / 00222939100770261" type="journal article" year="1991">Alderweireldt 1991: 363</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFEF5D4F99A6F8BD1F27A414" pageId="14" pageNumber="315" type="materials_examined">
<paragraph id="A1353685FFEF5D4F99A6F8BD1F42A478" blockId="14.[151,1436,998,1133]" pageId="14" pageNumber="315">
<emphasis id="93FEEA97FFEF5D4F99A6F8BD1BC5A45D" bold="true" box="[151,387,998,1024]" pageId="14" pageNumber="315">Material examined:</emphasis>
DHAYET BENDHAHOUA: Oumdjer, hand collecting,
<specimenCount id="B78CFD0CFFEF5D4F9D30F8BC1E6AA45C" box="[1025,1068,999,1025]" count="1" pageId="14" pageNumber="315" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFEF5D4F9D0BF8BD1EE5A45C" box="[1082,1187,998,1025]" pageId="14" pageNumber="315" value="2022-02-05">5.II.2022</date>
;
<specimenCount id="B78CFD0CFFEF5D4F9D81F8BC1E9AA45C" box="[1200,1244,999,1025]" count="2" pageId="14" pageNumber="315" type="male">2 ♂</specimenCount>
,
<date id="D5341045FFEF5D4F9DD9F8BD1F27A45C" box="[1256,1377,998,1025]" pageId="14" pageNumber="315" value="2022-02-15">15.II.2022</date>
.
<specimenCount id="B78CFD0CFFEF5D4F9C5AF8BC1FD3A45C" box="[1387,1429,999,1025]" count="1" pageId="14" pageNumber="315" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFEF5D4F99A6FF511B4DA478" box="[151,267,1034,1061]" pageId="14" pageNumber="315" value="2022-03-05">5.III.2022</date>
.
<materialsCitation id="11E23CD8FFEF5D4F9824FF50193DA478" box="[277,891,1035,1061]" collectorName="El Atteuf" location="Palm" pageId="14" pageNumber="315" specimenCount="1">
EL ATTEUF:
<location id="A455605EFFEF5D4F9882FF501BABA478" LSID="urn:lsid:plazi:treatment:29238793FFEF5D4F99A6F8071F5FA430:A455605EFFEF5D4F9882FF501BABA478" box="[435,493,1035,1061]" name="Palm" pageId="14" pageNumber="315">Palm</location>
grove of
<collectorName id="0C7F5353FFEF5D4F9B69FF501884A478" box="[600,706,1035,1061]" pageId="14" pageNumber="315">El Atteuf</collectorName>
, hand collecting
</materialsCitation>
,
<specimenCount id="B78CFD0CFFEF5D4F9AB7FF5019F4A478" box="[902,946,1035,1061]" count="4" pageId="14" pageNumber="315" type="male">4 ♂</specimenCount>
,
<date id="D5341045FFEF5D4F9A8FFF511E7BA478" box="[958,1085,1034,1061]" pageId="14" pageNumber="315" value="2015-04-18">18.IV.2015</date>
;
<specimenCount id="B78CFD0CFFEF5D4F9D78FF501E35A478" box="[1097,1139,1035,1061]" count="1" pageId="14" pageNumber="315" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFEF5D4F9DB0FF511F46A478" box="[1153,1280,1034,1061]" pageId="14" pageNumber="315" value="2023-04-15">15.IV.2023</date>
.
</paragraph>
<paragraph id="A1353685FFEF5D4F99F6FF751F27A414" blockId="14.[151,1436,998,1133]" box="[199,1377,1070,1097]" pageId="14" pageNumber="315">
<emphasis id="93FEEA97FFEF5D4F99F6FF751853A415" bold="true" box="[199,533,1070,1097]" pageId="14" pageNumber="315">
Previous records in
<collectingCountry id="D99D7615FFEF5D4F9882FF751856A415" box="[435,528,1070,1096]" name="Algeria" pageId="14" pageNumber="315">Algeria</collectingCountry>
:
</emphasis>
<collectingRegion id="634EF867FFEF5D4F9B2DFF741823A414" box="[540,613,1071,1097]" country="Algeria" name="Djelfa" pageId="14" pageNumber="315">Djelfa</collectingRegion>
,
<collectingRegion id="634EF867FFEF5D4F9B41FF7418EEA414" box="[624,680,1071,1097]" country="Algeria" name="Illizi" pageId="14" pageNumber="315">Illizi</collectingRegion>
,
<collectingRegion id="634EF867FFEF5D4F9B82FF741966A414" box="[691,800,1071,1097]" country="Algeria" name="Laghouat" pageId="14" pageNumber="315">Laghouat</collectingRegion>
,
<collectingRegion id="634EF867FFEF5D4F9A1AFF741933A414" box="[811,885,1071,1097]" country="Algeria" name="M'Sila" pageId="14" pageNumber="315">Msila</collectingRegion>
,
<collectingRegion id="634EF867FFEF5D4F9AB1FF741989A414" box="[896,975,1071,1097]" country="Algeria" name="Naama" pageId="14" pageNumber="315">Naama</collectingRegion>
(
<bibRefCitation id="C51B4B74FFEF5D4F9AEFFF741E2DA414" author="Simon, E." box="[990,1131,1071,1097]" pageId="14" pageNumber="315" pagination="82 - 87" refId="ref23337" refString="Simon, E. (1899) Liste des arachnides recueillis en Algerie par M. P. Lesne et descrition d'une espece nouvelle. Bulletin du Museum d'Histoire Naturelle Paris, 5, 82 - 87." type="journal article" year="1899">Simon 1899</bibRefCitation>
;
<bibRefCitation id="C51B4B74FFEF5D4F9D44FF741F12A414" author="Alderweireldt, M." box="[1141,1364,1071,1097]" pageId="14" pageNumber="315" pagination="359 - 381" refId="ref18524" refString="Alderweireldt, M. (1991) A revision of the African representatives of the wolf spider genus Evippa Simon, 1882 (Araneae, Lycosidae) with notes on allied species and genera. Journal of Natural History, 25 (2), 359 - 381. https: // doi. org / 10.1080 / 00222939100770261" type="journal article" year="1991">Alderweireldt 1991</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFEF5D4F99F6FF091F5FA430" box="[199,1305,1106,1133]" pageId="14" pageNumber="315" type="distribution">
<paragraph id="A1353685FFEF5D4F99F6FF091F5FA430" blockId="14.[151,1436,998,1133]" box="[199,1305,1106,1133]" pageId="14" pageNumber="315">
<emphasis id="93FEEA97FFEF5D4F99F6FF091B25A431" bold="true" box="[199,355,1106,1132]" pageId="14" pageNumber="315">Distribution:</emphasis>
<collectingCountry id="D99D7615FFEF5D4F9859FF081BF9A430" box="[360,447,1107,1133]" name="Algeria" pageId="14" pageNumber="315">Algeria</collectingCountry>
,
<collectingCountry id="D99D7615FFEF5D4F98FBFF081866A430" box="[458,544,1107,1133]" name="Tunisia" pageId="14" pageNumber="315">Tunisia</collectingCountry>
,
<collectingCountry id="D99D7615FFEF5D4F9B1AFF081829A430" box="[555,623,1107,1133]" name="Libya" pageId="14" pageNumber="315">Libya</collectingCountry>
,
<collectingCountry id="D99D7615FFEF5D4F9B4BFF0818FEA430" box="[634,696,1107,1133]" name="Chad" pageId="14" pageNumber="315">Chad</collectingCountry>
,
<collectingCountry id="D99D7615FFEF5D4F9BF2FF08194FA430" box="[707,777,1107,1133]" name="Egypt" pageId="14" pageNumber="315">Egypt</collectingCountry>
,
<collectingCountry id="D99D7615FFEF5D4F9A25FF081913A430" box="[788,853,1107,1133]" name="Israel" pageId="14" pageNumber="315">Israel</collectingCountry>
,
<collectingCountry id="D99D7615FFEF5D4F9A51FF081983A430" box="[864,965,1107,1133]" name="Palestine" pageId="14" pageNumber="315">Palestine</collectingCountry>
(
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@ -0,0 +1,194 @@
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<emphasis id="93FEEA97FFEF5D4F99A6FA351B84A1D4" bold="true" box="[151,450,366,393]" italics="true" pageId="14" pageNumber="315">Mesiotelus mauritanicus</emphasis>
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<emphasis id="93FEEA97FFEF5D4F99A6FAE81BD5A196" box="[151,403,435,459]" italics="true" pageId="14" pageNumber="315">Mesiotelus mauritanicus</emphasis>
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<bibRefCitation id="C51B4B74FFEF5D4F9B50FAE81961A196" author="Denis, J." box="[609,807,435,459]" pageId="14" pageNumber="315" pagination="1027 - 1060" refId="ref20639" refString="Denis, J. (1937 b) On a collection of spiders from Algeria. Proceedings of the Zoological Society of London, 106 (4), 1027 - 1060. https: // doi. org / 10.1111 / j. 1469 - 7998.1936. tb 06301. x" type="journal article" year="1937">Denis 1937b: 1051</bibRefCitation>
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<emphasis id="93FEEA97FFEF5D4F9D39FAEF1E7AA196" box="[1032,1084,435,459]" italics="true" pageId="14" pageNumber="315">et al.</emphasis>
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Brague-Bouragba
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2007: 938
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Bouseksou
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2015: 260
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Mansouri
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2019: 5
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Bouragba
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2020: 29
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<emphasis id="93FEEA97FFEF5D4F99A6F9431BC5A26F" bold="true" box="[151,387,536,562]" pageId="14" pageNumber="315">Material examined:</emphasis>
BERRIANE: Sidi Mbarek, hand collecting,
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,
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,
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.
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Previous records in
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:
</emphasis>
Without precise locality,
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,
<collectingRegion id="634EF867FFEF5D4F9AA1F9671989A20B" box="[912,975,572,598]" country="Algeria" name="Blida" pageId="14" pageNumber="315">Blida</collectingRegion>
,
<collectingRegion id="634EF867FFEF5D4F9AECF9671E60A20B" box="[989,1062,572,598]" country="Algeria" name="Djelfa" pageId="14" pageNumber="315">Djelfa</collectingRegion>
,
<collectingRegion id="634EF867FFEF5D4F9D02F9671E2DA20B" box="[1075,1131,572,598]" country="Algeria" name="Mila" pageId="14" pageNumber="315">Mila</collectingRegion>
,
<collectingRegion id="634EF867FFEF5D4F9D49F9671EBEA20B" box="[1144,1272,572,598]" country="Algeria" name="Tizi Ouzou" pageId="14" pageNumber="315">Tizi Ouzou</collectingRegion>
(
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Abrous-Kherbouche
<emphasis id="93FEEA97FFEF5D4F9B2DF93A1815A227" box="[540,595,608,634]" italics="true" pageId="14" pageNumber="315">et al.</emphasis>
1997
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Brague-Bouragba
<emphasis id="93FEEA97FFEF5D4F9A5DF93A19E2A227" box="[876,932,608,634]" italics="true" pageId="14" pageNumber="315">et al.</emphasis>
2007
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Bouseksou
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2015
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Mansouri
<emphasis id="93FEEA97FFEF5D4F9C54F93A1FDBA227" box="[1381,1437,608,634]" italics="true" pageId="14" pageNumber="315">et al.</emphasis>
2019
</bibRefCitation>
;
<bibRefCitation id="C51B4B74FFEF5D4F99EFF9DF1B8DA2C3" author="Bouragba, N. &amp; Bouragba, M. &amp; Djouklafi, A. &amp; Amraoui, S. &amp; Brague, A. &amp; Beladjal, L." box="[222,459,644,670]" pageId="14" pageNumber="315" pagination="25 - 34" refId="ref19910" refString="Bouragba, N., Bouragba, M., Djouklafi, A., Amraoui, S., Brague, A. &amp; Beladjal, L. (2020) Spider (Araneae) and beetle (Coleoptera) communities structure in the North African streamside zone (Zahrez Gharbi, Djelfa, Algeria) after reforestation. African Entomology, 28 (1), 25 - 34." type="journal article" year="2020">
Bouragba
<emphasis id="93FEEA97FFEF5D4F9863F9DE1BCCA2C3" box="[338,394,644,670]" italics="true" pageId="14" pageNumber="315">et al.</emphasis>
2020
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<emphasis id="93FEEA97FFEF5D4F99F6F9F31B25A29F" bold="true" box="[199,355,680,706]" pageId="14" pageNumber="315">Distribution:</emphasis>
Mediterranean (
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EL ATTEUF: Palm grove of El Atteuf, pitfall traps,
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Previous records in
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North Africa,
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<taxonomicName id="668A4D06FFF05D5099A6F8D61883A3FA" ID-CoL="4QG7K" authority="(Muster, 2007)" baseAuthorityName="Muster" baseAuthorityYear="2007" box="[151,709,909,936]" class="Arachnida" family="Philodromidae" genus="Pulchellodromus" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="lamellipalpis">
<emphasis id="93FEEA97FFF05D5099A6F8D61883A3FA" bold="true" box="[151,709,909,936]" pageId="17" pageNumber="318">
<emphasis id="93FEEA97FFF05D5099A6F8D6184FA3F5" bold="true" box="[151,521,909,936]" italics="true" pageId="17" pageNumber="318">Pulchellodromus lamellipalpis</emphasis>
(
<bibRefCitation id="C51B4B74FFF05D509B26F8D618FBA3FA" author="Muster, C. &amp; Bosmans, R. &amp; Thaler, K." box="[535,701,909,935]" pageId="17" pageNumber="318" pagination="39 - 72" refId="ref22110" refString="Muster, C., Bosmans, R. &amp; Thaler, K. (2007) The Philodromus pulchellus - group in the Mediterranean: taxonomic revision, phylogenetic analysis and biogeography (Araneae: Philodromidae). Invertebrate Systematics, 21 (1), 39 - 72. https: // doi. org / 10.1071 / IS 06014" type="journal article" year="2007">Muster, 2007</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFF05D5099A6F88918DEA3B7" box="[151,664,978,1002]" pageId="17" pageNumber="318" type="reference_group">
<paragraph id="A1353685FFF05D5099A6F88918DEA3B7" blockId="17.[151,664,978,1002]" box="[151,664,978,1002]" pageId="17" pageNumber="318">
<treatmentCitationGroup id="819A11ABFFF05D5099A6F88918DEA3B7" box="[151,664,978,1002]" pageId="17" pageNumber="318">
<taxonomicName id="668A4D06FFF05D5099A6F8891BE5A3B7" ID-CoL="9396B" authorityName="Muster" authorityYear="2007" box="[151,419,978,1002]" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="lamellipalpis">
<emphasis id="93FEEA97FFF05D5099A6F8891BE5A3B7" box="[151,419,978,1002]" italics="true" pageId="17" pageNumber="318">Philodromus lamellipalpis</emphasis>
</taxonomicName>
;
<treatmentCitation id="202B1094FFF05D509880F88818D2A3B7" author="Muster, C. &amp; Bosmans, R. &amp; Thaler, K." box="[433,660,978,1002]" page="55" pageId="17" pageNumber="318" year="2007">
<bibRefCitation id="C51B4B74FFF05D509880F88818D2A3B7" author="Muster, C. &amp; Bosmans, R. &amp; Thaler, K." box="[433,660,978,1002]" pageId="17" pageNumber="318" pagination="39 - 72" refId="ref22110" refString="Muster, C., Bosmans, R. &amp; Thaler, K. (2007) The Philodromus pulchellus - group in the Mediterranean: taxonomic revision, phylogenetic analysis and biogeography (Araneae: Philodromidae). Invertebrate Systematics, 21 (1), 39 - 72. https: // doi. org / 10.1071 / IS 06014" type="journal article" year="2007">
Muster
<emphasis id="93FEEA97FFF05D5098CEF8881875A3B7" box="[511,563,978,1002]" italics="true" pageId="17" pageNumber="318">et al.</emphasis>
2007: 55
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFF05D5099A6FF4319D1A427" pageId="17" pageNumber="318" type="materials_examined">
<paragraph id="A1353685FFF05D5099A6FF431E31A46E" blockId="17.[151,1347,1048,1147]" box="[151,1143,1048,1075]" pageId="17" pageNumber="318">
<emphasis id="93FEEA97FFF05D5099A6FF431BC5A46F" bold="true" box="[151,387,1048,1074]" pageId="17" pageNumber="318">Material examined:</emphasis>
MANSOURA: Mansoura Kedima: hand collecting,
<specimenCount id="B78CFD0CFFF05D509AE0FF4319BDA46E" box="[977,1019,1048,1075]" count="1" pageId="17" pageNumber="318" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF05D509D38FF431E35A46E" box="[1033,1139,1048,1075]" pageId="17" pageNumber="318" value="2015-02-07">7.II.2015</date>
.
</paragraph>
<paragraph id="A1353685FFF05D5099F6FF6719D1A427" blockId="17.[151,1347,1048,1147]" pageId="17" pageNumber="318">
<emphasis id="93FEEA97FFF05D5099F6FF671853A40B" bold="true" box="[199,533,1084,1110]" pageId="17" pageNumber="318">
Previous records in
<collectingCountry id="D99D7615FFF05D509882FF671856A40B" box="[435,528,1084,1110]" name="Algeria" pageId="17" pageNumber="318">Algeria</collectingCountry>
:
</emphasis>
This species is known previously only from
<collectingRegion id="634EF867FFF05D509D3CFF661E1AA40A" box="[1037,1116,1085,1111]" country="Algeria" name="Naama" pageId="17" pageNumber="318">Naama</collectingRegion>
(
<bibRefCitation id="C51B4B74FFF05D509D5AFF671F7AA40B" author="Muster, C. &amp; Bosmans, R. &amp; Thaler, K." box="[1131,1340,1084,1110]" pageId="17" pageNumber="318" pagination="39 - 72" refId="ref22110" refString="Muster, C., Bosmans, R. &amp; Thaler, K. (2007) The Philodromus pulchellus - group in the Mediterranean: taxonomic revision, phylogenetic analysis and biogeography (Araneae: Philodromidae). Invertebrate Systematics, 21 (1), 39 - 72. https: // doi. org / 10.1071 / IS 06014" type="journal article" year="2007">
Muster
<emphasis id="93FEEA97FFF05D509DF2FF661EBDA40B" box="[1219,1275,1084,1110]" italics="true" pageId="17" pageNumber="318">et al.</emphasis>
2007
</bibRefCitation>
)
<emphasis id="93FEEA97FFF05D5099F6FF3B1B25A427" bold="true" box="[199,355,1120,1146]" pageId="17" pageNumber="318">Distribution:</emphasis>
Endemic to Algeria (
<bibRefCitation id="C51B4B74FFF05D509B63FF3B19CDA427" author="World Spider Catalog" box="[594,907,1120,1146]" pageId="17" pageNumber="318" refId="ref24111" refString="World Spider Catalog (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern, Bern. Available from: https: // wsc. nmbe. ch / (accessed 1 June 2024) https: // doi. org / 10.24436 / 2" type="book" year="2024">World Spider Catalog 2024</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,159 @@
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<subSubSection id="E990650EFFF05D5099A6FF971960A4BB" box="[151,806,1228,1255]" pageId="17" pageNumber="318" type="nomenclature">
<paragraph id="A1353685FFF05D5099A6FF971960A4BB" blockId="17.[151,806,1228,1255]" box="[151,806,1228,1255]" pageId="17" pageNumber="318">
<heading id="FA7D81E9FFF05D5099A6FF971960A4BB" bold="true" box="[151,806,1228,1255]" fontSize="11" level="1" pageId="17" pageNumber="318" reason="1">
<taxonomicName id="668A4D06FFF05D5099A6FF971960A4BB" ID-CoL="4QG7Q" authority="(Muster &amp; Bosmans, 2007)" baseAuthorityName="Muster &amp; Bosmans" baseAuthorityYear="2007" box="[151,806,1228,1255]" class="Arachnida" family="Philodromidae" genus="Pulchellodromus" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="pardalis">
<emphasis id="93FEEA97FFF05D5099A6FF971960A4BB" bold="true" box="[151,806,1228,1255]" pageId="17" pageNumber="318">
<emphasis id="93FEEA97FFF05D5099A6FF971B96A4BA" bold="true" box="[151,464,1228,1255]" italics="true" pageId="17" pageNumber="318">Pulchellodromus pardalis</emphasis>
(
<bibRefCitation id="C51B4B74FFF05D5098EFFF97195BA4BB" author="Muster, C. &amp; Bosmans, R. &amp; Thaler, K." box="[478,797,1228,1254]" pageId="17" pageNumber="318" pagination="39 - 72" refId="ref22110" refString="Muster, C., Bosmans, R. &amp; Thaler, K. (2007) The Philodromus pulchellus - group in the Mediterranean: taxonomic revision, phylogenetic analysis and biogeography (Araneae: Philodromidae). Invertebrate Systematics, 21 (1), 39 - 72. https: // doi. org / 10.1071 / IS 06014" type="journal article" year="2007">Muster &amp; Bosmans, 2007</bibRefCitation>
)
</emphasis>
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<paragraph id="A1353685FFF05D5099A6FE4A18C4A574" blockId="17.[151,642,1297,1321]" box="[151,642,1297,1321]" pageId="17" pageNumber="318">
<treatmentCitationGroup id="819A11ABFFF05D5099A6FE4A18C4A574" box="[151,642,1297,1321]" pageId="17" pageNumber="318">
<taxonomicName id="668A4D06FFF05D5099A6FE4A1B35A574" ID-CoL="9396L" authorityName="Muster &amp; Bosmans" authorityYear="2007" box="[151,371,1297,1321]" class="Arachnida" family="Philodromidae" genus="Philodromus" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="pardalis">
<emphasis id="93FEEA97FFF05D5099A6FE4A1B35A574" box="[151,371,1297,1321]" italics="true" pageId="17" pageNumber="318">Philodromus pardalis</emphasis>
</taxonomicName>
; in
<treatmentCitation id="202B1094FFF05D5098ABFE4A1838A574" author="Muster, C. &amp; Bosmans, R. &amp; Thaler, K." box="[410,638,1297,1321]" page="59" pageId="17" pageNumber="318" year="2007">
<bibRefCitation id="C51B4B74FFF05D5098ABFE4A1838A574" author="Muster, C. &amp; Bosmans, R. &amp; Thaler, K." box="[410,638,1297,1321]" pageId="17" pageNumber="318" pagination="39 - 72" refId="ref22110" refString="Muster, C., Bosmans, R. &amp; Thaler, K. (2007) The Philodromus pulchellus - group in the Mediterranean: taxonomic revision, phylogenetic analysis and biogeography (Araneae: Philodromidae). Invertebrate Systematics, 21 (1), 39 - 72. https: // doi. org / 10.1071 / IS 06014" type="journal article" year="2007">
Muster
<emphasis id="93FEEA97FFF05D5098D8FE49185AA574" box="[489,540,1297,1321]" italics="true" pageId="17" pageNumber="318">et al.</emphasis>
2007: 59
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.
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</paragraph>
</subSubSection>
<subSubSection id="E990650EFFF05D5099A6FE0C1A9FA65C" pageId="17" pageNumber="318" type="materials_examined">
<paragraph id="A1353685FFF05D5099A6FE0C198CA5E7" blockId="17.[151,1437,1367,1574]" pageId="17" pageNumber="318">
<emphasis id="93FEEA97FFF05D5099A6FE0C1BC6A52C" bold="true" box="[151,384,1367,1393]" pageId="17" pageNumber="318">Material examined:</emphasis>
DHAYET BENDHAHOUA: Grazil, hand collecting,
<specimenCount id="B78CFD0CFFF05D509AE0FE0C19BFA52F" box="[977,1017,1367,1394]" count="1" pageId="17" pageNumber="318" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF05D509D35FE0C1E3EA52F" box="[1028,1144,1367,1394]" pageId="17" pageNumber="318" value="2022-03-05">5.III.2022</date>
. MANSOURA: Mansoura Kedima, hand collecting,
<specimenCount id="B78CFD0CFFF05D5098F5FE201BB4A5CB" box="[452,498,1403,1430]" count="3" pageId="17" pageNumber="318" type="female">3 ♀</specimenCount>
,
<date id="D5341045FFF05D509B35FE201828A5C8" box="[516,622,1403,1429]" pageId="17" pageNumber="318" value="2017-02-04">4.II.2017</date>
. METLILI: Guemgouma, hand collecting,
<specimenCount id="B78CFD0CFFF05D509D55FE201ED4A5CB" box="[1124,1170,1403,1430]" count="1" pageId="17" pageNumber="318" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF05D509D95FE201F5EA5CB" box="[1188,1304,1403,1430]" pageId="17" pageNumber="318" value="2022-03-05">5.III.2022</date>
;
<materialsCitation id="11E23CD8FFF05D509C16FE201B5EA5E7" collectingDate="2022-03-25" isTailEnumeration="true" pageId="17" pageNumber="318" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="B78CFD0CFFF05D509C16FE201F11A5CB" box="[1319,1367,1403,1430]" count="1" pageId="17" pageNumber="318" type="male">1 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFF05D509C56FE201FD3A5CB" box="[1383,1429,1403,1430]" count="1" pageId="17" pageNumber="318" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF05D5099A6FEC41B5EA5E7" box="[151,280,1439,1466]" pageId="17" pageNumber="318" value="2022-03-25">
<collectingDate id="C570E9ADFFF05D5099A6FEC41B5EA5E7" box="[151,280,1439,1466]" pageId="17" pageNumber="318" value="2022-03-25">25.III.2022</collectingDate>
</date>
</materialsCitation>
; Souareg, hand collecting,
<specimenCount id="B78CFD0CFFF05D509B79FEC41834A5E7" box="[584,626,1439,1466]" count="2" pageId="17" pageNumber="318" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFF05D509BB1FEC41947A5E7" box="[640,769,1439,1466]" pageId="17" pageNumber="318" value="2022-03-15">15.III.2022</date>
;
<materialsCitation id="11E23CD8FFF05D509A3DFEC41980A5E7" box="[780,966,1439,1466]" collectingDate="2022-03-25" isTailEnumeration="true" pageId="17" pageNumber="318" specimenCount="1" specimenCount-female="1">
<specimenCount id="B78CFD0CFFF05D509A3DFEC41970A5E7" box="[780,822,1439,1466]" count="1" pageId="17" pageNumber="318" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF05D509A75FEC41980A5E7" box="[836,966,1439,1466]" pageId="17" pageNumber="318" value="2022-03-25">
<collectingDate id="C570E9ADFFF05D509A75FEC41980A5E7" box="[836,966,1439,1466]" pageId="17" pageNumber="318" value="2022-03-25">25.III.2022</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="A1353685FFF05D5099F6FE981A9FA65C" blockId="17.[151,1437,1367,1574]" pageId="17" pageNumber="318">
<emphasis id="93FEEA97FFF05D5099F6FE981BE9A580" bold="true" box="[199,431,1475,1501]" pageId="17" pageNumber="318">Previous records in</emphasis>
<materialsCitation id="11E23CD8FFF05D509887FE981A9FA65C" country="Algeria" location="Ghardaia" pageId="17" pageNumber="318" specimenCount="1" stateProvince="Ghardaia">
<emphasis id="93FEEA97FFF05D509887FE98185EA580" bold="true" box="[438,536,1475,1501]" pageId="17" pageNumber="318">
<collectingCountry id="D99D7615FFF05D509887FE981855A580" box="[438,531,1475,1501]" name="Algeria" pageId="17" pageNumber="318">Algeria</collectingCountry>
:
</emphasis>
This species is known previously only from
<collectingRegion id="634EF867FFF05D509D24FE981E3AA580" box="[1045,1148,1475,1501]" country="Algeria" name="Ghardaia" pageId="17" pageNumber="318">Ghardaïa</collectingRegion>
and
<collectingRegion id="634EF867FFF05D509D85FE981EBAA580" box="[1204,1276,1475,1501]" country="Algeria" name="M'Sila" pageId="17" pageNumber="318">Msila</collectingRegion>
(
<bibRefCitation id="C51B4B74FFF05D509C3BFE981A97A65C" author="Muster, C. &amp; Bosmans, R. &amp; Thaler, K." pageId="17" pageNumber="318" pagination="39 - 72" refId="ref22110" refString="Muster, C., Bosmans, R. &amp; Thaler, K. (2007) The Philodromus pulchellus - group in the Mediterranean: taxonomic revision, phylogenetic analysis and biogeography (Araneae: Philodromidae). Invertebrate Systematics, 21 (1), 39 - 72. https: // doi. org / 10.1071 / IS 06014" type="journal article" year="2007">
Muster
<emphasis id="93FEEA97FFF05D509C52FE9F1FDBA580" box="[1379,1437,1475,1501]" italics="true" pageId="17" pageNumber="318">et al.</emphasis>
2007
</bibRefCitation>
)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFF05D5099F6FD501E77A678" box="[199,1073,1547,1574]" pageId="17" pageNumber="318" type="distribution">
<paragraph id="A1353685FFF05D5099F6FD501E77A678" blockId="17.[151,1437,1367,1574]" box="[199,1073,1547,1574]" pageId="17" pageNumber="318">
<emphasis id="93FEEA97FFF05D5099F6FD501B25A678" bold="true" box="[199,355,1547,1573]" pageId="17" pageNumber="318">Distribution:</emphasis>
<collectingCountry id="D99D7615FFF05D50985BFD501B8AA678" box="[362,460,1547,1573]" name="Portugal" pageId="17" pageNumber="318">Portugal</collectingCountry>
,
<collectingCountry id="D99D7615FFF05D5098E6FD50185CA678" box="[471,538,1547,1573]" name="Spain" pageId="17" pageNumber="318">Spain</collectingCountry>
,
<collectingCountry id="D99D7615FFF05D509B12FD501831A678" box="[547,631,1547,1573]" name="Algeria" pageId="17" pageNumber="318">Algeria</collectingCountry>
to
<collectingCountry id="D99D7615FFF05D509BAAFD571898A67B" box="[667,734,1548,1574]" name="Egypt" pageId="17" pageNumber="318">Egypt</collectingCountry>
(
<bibRefCitation id="C51B4B74FFF05D509BDDFD501E63A678" author="World Spider Catalog" box="[748,1061,1547,1573]" pageId="17" pageNumber="318" refId="ref24111" refString="World Spider Catalog (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern, Bern. Available from: https: // wsc. nmbe. ch / (accessed 1 June 2024) https: // doi. org / 10.24436 / 2" type="book" year="2024">World Spider Catalog 2024</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,469 @@
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<subSubSection id="E990650EFFF05D5099A6FD2D1853A6CC" box="[151,533,1654,1681]" pageId="17" pageNumber="318" type="nomenclature">
<paragraph id="A1353685FFF05D5099A6FD2D1853A6CC" blockId="17.[151,533,1654,1681]" box="[151,533,1654,1681]" pageId="17" pageNumber="318">
<heading id="FA7D81E9FFF05D5099A6FD2D1853A6CC" bold="true" box="[151,533,1654,1681]" fontSize="11" level="1" pageId="17" pageNumber="318" reason="1">
<taxonomicName id="668A4D06FFF05D5099A6FD2D1853A6CC" ID-CoL="5644S" authority="Simon, 1870" authorityName="Simon" authorityYear="1870" box="[151,533,1654,1681]" class="Arachnida" family="Philodromidae" genus="Thanatus" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="vulgaris">
<emphasis id="93FEEA97FFF05D5099A6FD2D1853A6CC" bold="true" box="[151,533,1654,1681]" pageId="17" pageNumber="318">
<emphasis id="93FEEA97FFF05D5099A6FD2D1B32A6CC" bold="true" box="[151,372,1654,1681]" italics="true" pageId="17" pageNumber="318">Thanatus vulgaris</emphasis>
Simon, 1870
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<treatmentCitationGroup id="819A11ABFFF05D5099A6FDE01BFEA6AF" pageId="17" pageNumber="318">
<taxonomicName id="668A4D06FFF05D5099A6FDE01B16A68E" ID-CoL="5644S" authorityName="Simon" authorityYear="1870" box="[151,336,1723,1747]" class="Arachnida" family="Philodromidae" genus="Thanatus" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="vulgaris">
<emphasis id="93FEEA97FFF05D5099A6FDE01B16A68E" box="[151,336,1723,1747]" italics="true" pageId="17" pageNumber="318">Thanatus vulgaris</emphasis>
</taxonomicName>
;
<treatmentCitation id="202B1094FFF05D50986FFDE01843A68E" author="Simon, E." box="[350,517,1723,1748]" page="85" pageId="17" pageNumber="318" year="1899">
<bibRefCitation id="C51B4B74FFF05D50986FFDE01843A68E" author="Simon, E." box="[350,517,1723,1748]" pageId="17" pageNumber="318" pagination="82 - 87" refId="ref23337" refString="Simon, E. (1899) Liste des arachnides recueillis en Algerie par M. P. Lesne et descrition d'une espece nouvelle. Bulletin du Museum d'Histoire Naturelle Paris, 5, 82 - 87." type="journal article" year="1899">Simon 1899: 85</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="202B1094FFF05D509B21FDE01882A68E" author="Strand, E." box="[528,708,1723,1748]" page="135" pageId="17" pageNumber="318" year="1907">
<bibRefCitation id="C51B4B74FFF05D509B21FDE01882A68E" author="Strand, E." box="[528,708,1723,1748]" pageId="17" pageNumber="318" pagination="103 - 147" refId="ref23684" refString="Strand, E. (1907) Nordafrikanische, hauptsachlich von Carlo Freiherr von Erlanger gesammelte Thomisiden. Jahrbucher des Nassauischen Vereins fur Naturkunde, 60, 103 - 147. https: // doi. org / 10.5962 / bhl. part. 27907" type="journal article" year="1907">Strand 1907: 135</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="202B1094FFF05D509BFEFDE019D3A689" author="Denis, J." box="[719,917,1723,1748]" page="1048" pageId="17" pageNumber="318" year="1937">
<bibRefCitation id="C51B4B74FFF05D509BFEFDE019D3A689" author="Denis, J." box="[719,917,1723,1748]" pageId="17" pageNumber="318" pagination="1027 - 1060" refId="ref20639" refString="Denis, J. (1937 b) On a collection of spiders from Algeria. Proceedings of the Zoological Society of London, 106 (4), 1027 - 1060. https: // doi. org / 10.1111 / j. 1469 - 7998.1936. tb 06301. x" type="journal article" year="1937">Denis 1937b: 1048</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="202B1094FFF05D509AAEFDE01EC5A689" author="Alioua, Y. &amp; Bissati, S. &amp; Kherbouche, O. &amp; Bosmans, R." box="[927,1155,1723,1748]" page="37" pageId="17" pageNumber="318" year="2016">
<bibRefCitation id="C51B4B74FFF05D509AAEFDE01EC5A689" author="Alioua, Y. &amp; Bissati, S. &amp; Kherbouche, O. &amp; Bosmans, R." box="[927,1155,1723,1748]" pageId="17" pageNumber="318" pagination="33 - 40" refId="ref18687" refString="Alioua, Y., Bissati, S., Kherbouche, O. &amp; Bosmans, R. (2016) Spiders of Sebkhet El Melah (Northern Sahara, Algeria): review and new records. Serket, 15 (1), 33 - 40." type="journal article" year="2016">
Alioua
<emphasis id="93FEEA97FFF05D509ADAFDE71E59A68E" box="[1003,1055,1723,1747]" italics="true" pageId="17" pageNumber="318">et al.</emphasis>
2016: 37
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="202B1094FFF05D509DBFFDE01FC9A68E" author="Boucherit, S. &amp; Bouragba, N. &amp; Douaoui, A. &amp; Bosmans, R." box="[1166,1423,1723,1748]" page="63" pageId="17" pageNumber="318" year="2020">
<bibRefCitation id="C51B4B74FFF05D509DBFFDE01FC9A68E" author="Boucherit, S. &amp; Bouragba, N. &amp; Douaoui, A. &amp; Bosmans, R." box="[1166,1423,1723,1748]" pageId="17" pageNumber="318" pagination="60 - 70" refId="ref19840" refString="Boucherit, S., Bouragba, N., Douaoui, A. &amp; Bosmans, R. (2020) Araignees de la plaine alluviale du Haut-Cheliff (Algerie), avec la citation de quelques especes nouvelles ou rares pour l'Algerie (Araneae: Gnaphosidae, Miturgidae). Nieuwsbrief van de Belgische Arachnologische Vereniging, 35 (1 - 2), 60 - 70." type="journal article" year="2020">
Boucherit
<emphasis id="93FEEA97FFF05D509DCBFDE71F68A68E" box="[1274,1326,1723,1747]" italics="true" pageId="17" pageNumber="318">et al.</emphasis>
2020: 63
</bibRefCitation>
</treatmentCitation>
; Alioua
<emphasis id="93FEEA97FFF05D509823FD801B00A6AF" box="[274,326,1754,1778]" italics="true" pageId="17" pageNumber="318">et al.</emphasis>
2022: 167.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFF05D5399A6FC7B1998A244" lastPageId="18" lastPageNumber="319" pageId="17" pageNumber="318" type="materials_examined">
<paragraph id="A1353685FFF05D5399A6FC7B1910A18C" blockId="17.[151,1436,1824,2031]" lastBlockId="18.[151,1437,150,645]" lastPageId="18" lastPageNumber="319" pageId="17" pageNumber="318">
<emphasis id="93FEEA97FFF05D5099A6FC7B1BC2A767" bold="true" box="[151,388,1824,1850]" pageId="17" pageNumber="318">Material examined:</emphasis>
BERRIANE: Balouh, hand collecting,
<specimenCount id="B78CFD0CFFF05D509A77FC7B1934A766" box="[838,882,1824,1851]" count="1" pageId="17" pageNumber="318" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF05D509AB0FC7B19ADA767" box="[897,1003,1824,1850]" pageId="17" pageNumber="318" value="2021-02-04">4.II.2021</date>
;
<specimenCount id="B78CFD0CFFF05D509AC9FC7B1E62A766" box="[1016,1060,1824,1851]" count="1" pageId="17" pageNumber="318" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF05D509D02FC7B1EDBA767" box="[1075,1181,1824,1850]" pageId="17" pageNumber="318" value="2021-02-06">6.II.2021</date>
;
<specimenCount id="B78CFD0CFFF05D509D9BFC7B1E93A766" box="[1194,1237,1824,1851]" count="1" pageId="17" pageNumber="318" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF05D509DD4FC7B1F1BA767" box="[1253,1373,1824,1850]" pageId="17" pageNumber="318" value="2021-02-19">19.II.2021</date>
;
<specimenCount id="B78CFD0CFFF05D509C5BFC7B1FD3A766" box="[1386,1429,1824,1851]" count="1" pageId="17" pageNumber="318" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF05D5099A6FC1F1B4CA702" box="[151,266,1860,1887]" pageId="17" pageNumber="318" value="2021-03-05">5.III.2021</date>
;
<specimenCount id="B78CFD0CFFF05D509827FC1F1B07A702" box="[278,321,1860,1887]" count="1" pageId="17" pageNumber="318" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFF05D50987CFC1F1BF8A703" box="[333,446,1860,1886]" pageId="17" pageNumber="318" value="2021-04-03">3.IV.2021</date>
;
<specimenCount id="B78CFD0CFFF05D5098F8FC1F1BB3A702" box="[457,501,1860,1887]" count="2" pageId="17" pageNumber="318" type="male">2 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFF05D509B30FC1F186CA702" box="[513,554,1860,1887]" count="1" pageId="17" pageNumber="318" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF05D509B09FC1F18F1A703" box="[568,695,1860,1886]" pageId="17" pageNumber="318" value="2021-04-10">10.IV.2021</date>
;
<specimenCount id="B78CFD0CFFF05D509BF3FC1F18AAA702" box="[706,748,1860,1887]" count="1" pageId="17" pageNumber="318" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF05D509BCBFC1F193FA703" box="[762,889,1860,1886]" pageId="17" pageNumber="318" value="2021-04-24">24.IV.2021</date>
;
<specimenCount id="B78CFD0CFFF05D509AB5FC1F19E8A702" box="[900,942,1860,1887]" count="1" pageId="17" pageNumber="318" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF05D509A8AFC1F1E7DA703" box="[955,1083,1860,1886]" pageId="17" pageNumber="318" value="2021-04-30">30.IV.2021</date>
; Pifall traps,
<specimenCount id="B78CFD0CFFF05D509DFCFC1F1EBFA702" box="[1229,1273,1860,1887]" count="1" pageId="17" pageNumber="318" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFF05D509C34FC1F1FC5A703" box="[1285,1411,1860,1886]" pageId="17" pageNumber="318" value="2021-04-10">10.IV.2021</date>
,
<specimenCount id="B78CFD0CFFF05D509CBFFC1F1AEAA7DE" count="1" pageId="17" pageNumber="318" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF05D509988FC331B7FA7DF" box="[185,313,1896,1922]" pageId="17" pageNumber="318" value="2021-04-24">24.IV.2021</date>
. Sidi Mbarek, hand collecting,;
<specimenCount id="B78CFD0CFFF05D509B90FC33188CA7DE" box="[673,714,1896,1923]" count="1" pageId="17" pageNumber="318" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF05D509BE9FC331904A7DF" box="[728,834,1896,1922]" pageId="17" pageNumber="318" value="2021-02-06">6.II.2021</date>
;
<specimenCount id="B78CFD0CFFF05D509A7DFC33193EA7DE" box="[844,888,1896,1923]" count="2" pageId="17" pageNumber="318" type="male">2 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFF05D509AB2FC3319EAA7DE" box="[899,940,1896,1923]" count="3" pageId="17" pageNumber="318" type="female">3 ♀</specimenCount>
,
<date id="D5341045FFF05D509A8BFC331E76A7DF" box="[954,1072,1896,1922]" pageId="17" pageNumber="318" value="2021-02-11">11.II.2021</date>
;
<specimenCount id="B78CFD0CFFF05D509D0AFC331E21A7DE" box="[1083,1127,1896,1923]" count="1" pageId="17" pageNumber="318" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFF05D509D43FC331EA3A7DF" box="[1138,1253,1896,1922]" pageId="17" pageNumber="318" value="2021-03-04">4.III.2021</date>
;
<specimenCount id="B78CFD0CFFF05D509DC1FC331F5FA7DE" box="[1264,1305,1896,1923]" count="1" pageId="17" pageNumber="318" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF05D509C16FC331FDEA7DF" box="[1319,1432,1896,1922]" pageId="17" pageNumber="318" value="2021-04-09">9.IV.2021</date>
;
<specimenCount id="B78CFD0CFFF05D5099A6FCD71A83A7FA" box="[151,197,1932,1959]" count="1" pageId="17" pageNumber="318" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFF05D5099E3FCD71B17A7FB" box="[210,337,1932,1958]" pageId="17" pageNumber="318" value="2021-04-23">23.IV.2021</date>
;
<specimenCount id="B78CFD0CFFF05D50986FFCD71BCAA7FA" box="[350,396,1932,1959]" count="2" pageId="17" pageNumber="318" type="male">2 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFF05D5098A8FCD71B83A7FA" box="[409,453,1932,1959]" count="1" pageId="17" pageNumber="318" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF05D5098E5FCD71815A7FB" box="[468,595,1932,1958]" pageId="17" pageNumber="318" value="2021-04-29">29.IV.2021</date>
, pitfall traps,
<specimenCount id="B78CFD0CFFF05D509BC0FCD71958A7FA" box="[753,798,1932,1959]" count="1" pageId="17" pageNumber="318" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFF05D509A1DFCD719E4A7FB" box="[812,930,1932,1958]" pageId="17" pageNumber="318" value="2021-02-11">11.II.2021</date>
;
<specimenCount id="B78CFD0CFFF05D509A9EFCD7199BA7FA" box="[943,989,1932,1959]" count="1" pageId="17" pageNumber="318" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFF05D509ADBFCD71E24A7FB" box="[1002,1122,1932,1958]" pageId="17" pageNumber="318" value="2021-02-18">18.II.2021</date>
;
<specimenCount id="B78CFD0CFFF05D509D5EFCD71EDBA7FA" box="[1135,1181,1932,1959]" count="1" pageId="17" pageNumber="318" type="male">1 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFF05D509D9BFCD71E90A7FA" box="[1194,1238,1932,1959]" count="2" pageId="17" pageNumber="318" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFF05D509DD4FCD71F1BA7FA" box="[1253,1373,1932,1959]" pageId="17" pageNumber="318" value="2021-02-25">25.II.2021</date>
;
<specimenCount id="B78CFD0CFFF05D509C5BFCD71FD1A7FA" box="[1386,1431,1932,1959]" count="1" pageId="17" pageNumber="318" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFF05D5099A6FCEB1B4CA797" box="[151,266,1968,1994]" pageId="17" pageNumber="318" value="2021-03-04">4.III.2021</date>
;
<specimenCount id="B78CFD0CFFF05D509829FCEB1B03A796" box="[280,325,1968,1995]" count="1" pageId="17" pageNumber="318" type="male">1 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFF05D509862FCEB1B38A796" box="[339,382,1968,1995]" count="1" pageId="17" pageNumber="318" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF05D5098BFFCEB1849A797" box="[398,527,1968,1994]" pageId="17" pageNumber="318" value="2021-03-18">18.III.2021</date>
;
<specimenCount id="B78CFD0CFFF05D509B2DFCEB180EA796" box="[540,584,1968,1995]" count="2" pageId="17" pageNumber="318" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFF05D509B69FCEB188FA797" box="[600,713,1968,1994]" pageId="17" pageNumber="318" value="2021-04-02">2.IV.2021</date>
;
<specimenCount id="B78CFD0CFFF05D509BE7FCEB1942A796" box="[726,772,1968,1995]" count="1" pageId="17" pageNumber="318" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFF05D509A20FCEB19D6A797" box="[785,912,1968,1994]" pageId="17" pageNumber="318" value="2021-04-16">16.IV.2021</date>
;
<specimenCount id="B78CFD0CFFF05D509AAFFCEB198FA796" box="[926,969,1968,1995]" count="1" pageId="17" pageNumber="318" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF05D509AE8FCEB1E1EA797" box="[985,1112,1968,1994]" pageId="17" pageNumber="318" value="2021-04-23">23.IV.2021</date>
;
<specimenCount id="B78CFD0CFFF05D509D54FCEB1ED5A796" box="[1125,1171,1968,1995]" count="2" pageId="17" pageNumber="318" type="male">2 ♂</specimenCount>
,
<date id="D5341045FFF05D509D91FCEB1F66A797" box="[1184,1312,1968,1994]" pageId="17" pageNumber="318" value="2021-04-29">29.IV.2021</date>
. DHAYET BENDHAHOUA: Grazil, hand collecting,
<specimenCount id="B78CFD0CFFF05D509BB3FC8F18F6A7B2" box="[642,688,2004,2031]" count="2" pageId="17" pageNumber="318" type="male">2 ♂</specimenCount>
,
<date id="D5341045FFF05D509B8FFC8F197AA7B2" box="[702,828,2004,2031]" pageId="17" pageNumber="318" value="2021-12-05">5.XII.2021</date>
;
<specimenCount id="B78CFD0CFFF05D509A7BFC8F193EA7B2" box="[842,888,2004,2031]" count="1" pageId="17" pageNumber="318" type="male">1 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFF05D509AB7FC8F19F5A7B2" box="[902,947,2004,2031]" count="1" pageId="17" pageNumber="318" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF05D509AF2FC8F1E65A7B2" box="[963,1059,2004,2031]" pageId="17" pageNumber="318" value="2022-01-05">5.I.2022</date>
;
<specimenCount id="B78CFD0CFFF05D509D00FC8F1E1BA7B2" box="[1073,1117,2004,2031]" count="1" pageId="17" pageNumber="318" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF05D509D5CFC8F1EABA7B2" box="[1133,1261,2004,2031]" pageId="17" pageNumber="318" value="2022-03-15">15.III.2022</date>
, pitfall traps,
<specimenCount id="B78CFD0CFFF05D539CBFFC8F1AEAA0EC" count="1" lastPageId="18" lastPageNumber="319" pageId="17" pageNumber="318" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF35D53998DFBCD1B7AA0EC" box="[188,316,150,177]" pageId="18" pageNumber="319" value="2022-03-15">15.III.2022</date>
, Oumdjer, hand collecting,
<specimenCount id="B78CFD0CFFF35D539B4BFBCC18EEA0EC" box="[634,680,151,177]" count="1" pageId="18" pageNumber="319" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFF35D539B87FBCD1959A0EC" box="[694,799,150,177]" pageId="18" pageNumber="319" value="2022-02-05">5.II.2022</date>
;
<specimenCount id="B78CFD0CFFF35D539A1CFBCC191FA0EC" box="[813,857,151,177]" count="1" pageId="18" pageNumber="319" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF35D539A58FBCD19ACA0EC" box="[873,1002,150,177]" pageId="18" pageNumber="319" value="2022-03-15">15.III.2022</date>
;
<specimenCount id="B78CFD0CFFF35D539AC9FBCC1E65A0EC" box="[1016,1059,151,177]" count="2" pageId="18" pageNumber="319" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFF35D539D02FBCD1EF3A0EC" box="[1075,1205,150,177]" pageId="18" pageNumber="319" value="2022-03-25">25.III.2022</date>
. EL ATTEUF: Palm grove of El Atteuf, hand collecting,
<specimenCount id="B78CFD0CFFF35D539B1EFBE0181AA088" box="[559,604,187,213]" count="1" pageId="18" pageNumber="319" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFF35D539B58FBE018A6A088" box="[617,736,187,213]" pageId="18" pageNumber="319" value="2014-02-20">20.II.2014</date>
;
<specimenCount id="B78CFD0CFFF35D539BDCFBE0195CA088" box="[749,794,187,213]" count="1" pageId="18" pageNumber="319" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFF35D539A16FBE119F5A088" box="[807,947,186,213]" pageId="18" pageNumber="319" value="2015-12-15">15.XII.2015</date>
;
<specimenCount id="B78CFD0CFFF35D539A8EFBE019AAA088" box="[959,1004,187,213]" count="1" pageId="18" pageNumber="319" type="male">1 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFF35D539AC8FBE01E62A088" box="[1017,1060,187,213]" count="2" pageId="18" pageNumber="319" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFF35D539D02FBE11ED5A088" box="[1075,1171,186,213]" pageId="18" pageNumber="319" value="2015-01-05">5.I.2015</date>
;
<specimenCount id="B78CFD0CFFF35D539D91FBE01E8BA088" box="[1184,1229,187,213]" count="1" pageId="18" pageNumber="319" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFF35D539DEBFBE11F0EA088" box="[1242,1352,186,213]" pageId="18" pageNumber="319" value="2015-01-15">15.I.2015</date>
;
<specimenCount id="B78CFD0CFFF35D539C64FBE01FC7A088" box="[1365,1409,187,213]" count="2" pageId="18" pageNumber="319" type="male">2 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFF35D539CBFFBE01AEAA0A4" count="2" pageId="18" pageNumber="319" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFF35D53998AFB851B74A0A4" box="[187,306,222,249]" pageId="18" pageNumber="319" value="2015-02-16">16.II.2015</date>
;
<specimenCount id="B78CFD0CFFF35D53980FFB841B2FA0A4" box="[318,361,223,249]" count="2" pageId="18" pageNumber="319" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFF35D539846FB851BA8A0A4" box="[375,494,222,249]" pageId="18" pageNumber="319" value="2015-02-25">25.II.2015</date>
,
<specimenCount id="B78CFD0CFFF35D5398CBFB841862A0A4" box="[506,548,223,249]" count="3" pageId="18" pageNumber="319" type="female">3 ♀</specimenCount>
,
<date id="D5341045FFF35D539B02FB8518E0A0A4" box="[563,678,222,249]" pageId="18" pageNumber="319" value="2015-03-06">6.III.2015</date>
;
<specimenCount id="B78CFD0CFFF35D539B83FB841898A0A4" box="[690,734,223,249]" count="2" pageId="18" pageNumber="319" type="male">2 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFF35D539BDBFB841949A0A4" box="[746,783,222,249]" count="7" pageId="18" pageNumber="319" type="female">7 ♀</specimenCount>
,
<date id="D5341045FFF35D539A25FB8519DBA0A4" box="[788,925,222,249]" pageId="18" pageNumber="319" value="2015-03-15">15.III.2015</date>
;
<specimenCount id="B78CFD0CFFF35D539A98FB841993A0A4" box="[937,981,223,249]" count="8" pageId="18" pageNumber="319" type="male">8 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFF35D539AD0FB841E5CA0A4" box="[993,1050,223,249]" count="10" pageId="18" pageNumber="319" type="female">10 ♀</specimenCount>
,
<date id="D5341045FFF35D539D19FB851EECA0A4" box="[1064,1194,222,249]" pageId="18" pageNumber="319" value="2015-03-25">25.III.2015</date>
;
<specimenCount id="B78CFD0CFFF35D539D84FB841EA4A0A4" box="[1205,1250,223,249]" count="1" pageId="18" pageNumber="319" type="male">1 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFF35D539DDFFB841F5EA0A4" box="[1262,1304,223,249]" count="4" pageId="18" pageNumber="319" type="female">4 ♀</specimenCount>
,
<date id="D5341045FFF35D539C16FB851FDEA0A4" box="[1319,1432,222,249]" pageId="18" pageNumber="319" value="2015-04-05">5.IV.2015</date>
;
<specimenCount id="B78CFD0CFFF35D5399A6FA581A83A140" box="[151,197,259,285]" count="1" pageId="18" pageNumber="319" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFF35D5399E3FA591B0FA140" box="[210,329,258,285]" pageId="18" pageNumber="319" value="2023-02-25">25.II.2023</date>
;
<specimenCount id="B78CFD0CFFF35D539866FA581BC2A140" box="[343,388,259,285]" count="1" pageId="18" pageNumber="319" type="male">1 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFF35D5398A0FA581BFBA140" box="[401,445,259,285]" count="1" pageId="18" pageNumber="319" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF35D5398FDFA591808A140" box="[460,590,258,285]" pageId="18" pageNumber="319" value="2023-03-25">25.III.2023</date>
;
<specimenCount id="B78CFD0CFFF35D539B6BFA5818C3A140" box="[602,645,259,285]" count="1" pageId="18" pageNumber="319" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF35D539BA4FA591955A140" box="[661,787,258,285]" pageId="18" pageNumber="319" value="2023-04-15">15.IV.2023</date>
, pitfall traps,
<specimenCount id="B78CFD0CFFF35D539A80FA581998A140" box="[945,990,259,285]" count="1" pageId="18" pageNumber="319" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFF35D539ADDFA591E25A140" box="[1004,1123,258,285]" pageId="18" pageNumber="319" value="2015-02-16">16.II.2015</date>
,
<specimenCount id="B78CFD0CFFF35D539D41FA581EDBA140" box="[1136,1181,259,285]" count="1" pageId="18" pageNumber="319" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFF35D539D9BFA591F64A140" box="[1194,1314,258,285]" pageId="18" pageNumber="319" value="2015-02-25">25.II.2015</date>
;
<specimenCount id="B78CFD0CFFF35D539C1EFA581F1BA140" box="[1327,1373,259,285]" count="4" pageId="18" pageNumber="319" type="male">4 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFF35D539C5BFA581FD3A140" box="[1386,1429,259,285]" count="1" pageId="18" pageNumber="319" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF35D5399A6FA7D1B4DA11C" box="[151,267,294,321]" pageId="18" pageNumber="319" value="2015-03-06">6.III.2015</date>
;
<specimenCount id="B78CFD0CFFF35D539826FA7C1B15A11C" box="[279,339,295,321]" count="10" pageId="18" pageNumber="319" type="male">10 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFF35D539851FA7C1BCDA11C" box="[352,395,295,321]" count="3" pageId="18" pageNumber="319" type="female">3 ♀</specimenCount>
,
<date id="D5341045FFF35D5398AAFA7D185AA11C" box="[411,540,294,321]" pageId="18" pageNumber="319" value="2015-03-15">15.III.2015</date>
;
<specimenCount id="B78CFD0CFFF35D539B18FA7C1810A11C" box="[553,598,295,321]" count="8" pageId="18" pageNumber="319" type="male">8 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFF35D539B52FA7C18C9A11C" box="[611,655,295,321]" count="2" pageId="18" pageNumber="319" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFF35D539BAFFA7D1966A11C" box="[670,800,294,321]" pageId="18" pageNumber="319" value="2015-03-25">25.III.2015</date>
;
<specimenCount id="B78CFD0CFFF35D539A1DFA7C192EA11C" box="[812,872,295,321]" count="19" pageId="18" pageNumber="319" type="male">19 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFF35D539A44FA7C19E6A11C" box="[885,928,295,321]" count="2" pageId="18" pageNumber="319" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFF35D539A81FA7D1E67A11C" box="[944,1057,294,321]" pageId="18" pageNumber="319" value="2015-04-05">5.IV.2015</date>
;
<specimenCount id="B78CFD0CFFF35D539D1FFA7C1E1DA11C" box="[1070,1115,295,321]" count="3" pageId="18" pageNumber="319" type="male">3 ♂</specimenCount>
,
<date id="D5341045FFF35D539D59FA7D1EA1A11C" box="[1128,1255,294,321]" pageId="18" pageNumber="319" value="2015-04-18">18.IV.2015</date>
.
<specimenCount id="B78CFD0CFFF35D539DC5FA7C1F59A11C" box="[1268,1311,295,321]" count="1" pageId="18" pageNumber="319" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF35D539C1FFA7D1FDEA11C" box="[1326,1432,294,321]" pageId="18" pageNumber="319" value="2023-02-05">5.II.2023</date>
. EL GUERRARA: Aghzou, hand collecting,
<specimenCount id="B78CFD0CFFF35D539BA9FA101883A138" box="[664,709,331,357]" count="1" pageId="18" pageNumber="319" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF35D539BE6FA101902A138" box="[727,836,331,357]" pageId="18" pageNumber="319" value="2020-01-31">31.I.2020</date>
,
<specimenCount id="B78CFD0CFFF35D539A62FA1019C7A138" box="[851,897,331,357]" count="1" pageId="18" pageNumber="319" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF35D539AA3FA111E4FA138" box="[914,1033,330,357]" pageId="18" pageNumber="319" value="2020-02-15">15.II.2020</date>
;
<specimenCount id="B78CFD0CFFF35D539D28FA101E0EA138" box="[1049,1096,331,357]" count="1" pageId="18" pageNumber="319" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFF35D539D69FA101E8DA138" box="[1112,1227,331,357]" pageId="18" pageNumber="319" value="2020-03-03">3.III.2020</date>
;
<specimenCount id="B78CFD0CFFF35D539DEBFA101F4FA138" box="[1242,1289,331,357]" count="1" pageId="18" pageNumber="319" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFF35D539C29FA101FDEA138" box="[1304,1432,331,357]" pageId="18" pageNumber="319" value="2020-03-11">11.III.2020</date>
, pitfall traps,
<specimenCount id="B78CFD0CFFF35D53981AFA341B1FA1D4" box="[299,345,367,393]" count="2" pageId="18" pageNumber="319" type="male">2 ♂</specimenCount>
,
<date id="D5341045FFF35D539859FA341B94A1D4" box="[360,466,367,393]" pageId="18" pageNumber="319" value="2020-02-08">8.II.2020</date>
;
<specimenCount id="B78CFD0CFFF35D5398D1FA341848A1D4" box="[480,526,367,393]" count="2" pageId="18" pageNumber="319" type="male">2 ♂</specimenCount>
,
<date id="D5341045FFF35D539B2CFA3418D2A1D4" box="[541,660,367,393]" pageId="18" pageNumber="319" value="2020-02-22">22.II.2020</date>
;
<specimenCount id="B78CFD0CFFF35D539B92FA341897A1D4" box="[675,721,367,393]" count="3" pageId="18" pageNumber="319" type="male">3 ♂</specimenCount>
,
<date id="D5341045FFF35D539BD1FA341915A1D4" box="[736,851,367,393]" pageId="18" pageNumber="319" value="2020-03-03">3.III.2020</date>
;
<specimenCount id="B78CFD0CFFF35D539A50FA3419C9A1D4" box="[865,911,367,393]" count="2" pageId="18" pageNumber="319" type="male">2 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFF35D539AAFFA34198CA1D4" box="[926,970,367,393]" count="2" pageId="18" pageNumber="319" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFF35D539AEAFA341E1BA1D4" box="[987,1117,367,393]" pageId="18" pageNumber="319" value="2020-03-17">17.III.2020</date>
. El Amied, hand collecting,
<specimenCount id="B78CFD0CFFF35D5399A6FAC81A82A1F0" box="[151,196,403,429]" count="3" pageId="18" pageNumber="319" type="female">3 ♀</specimenCount>
,
<date id="D5341045FFF35D5399E4FAC91B0AA1F0" box="[213,332,402,429]" pageId="18" pageNumber="319" value="2020-02-15">15.II.2020</date>
, pitfall traps,
<specimenCount id="B78CFD0CFFF35D5398DEFAC81858A1F0" box="[495,542,403,429]" count="1" pageId="18" pageNumber="319" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFF35D539B1CFAC818E3A1F0" box="[557,677,403,429]" pageId="18" pageNumber="319" value="2020-02-22">22.II.2020</date>
. MANSOURA: Mansoura Kedima, pitfall traps,
<specimenCount id="B78CFD0CFFF35D539DEDFAC81F4DA1F0" box="[1244,1291,403,429]" count="1" pageId="18" pageNumber="319" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFF35D539C2BFAC81FDFA1F0" box="[1306,1433,403,429]" pageId="18" pageNumber="319" value="2014-12-08">8.XII.2014</date>
. METLILI: Guemgouma, hand collecting,
<specimenCount id="B78CFD0CFFF35D539B5FFAEC18DCA18C" box="[622,666,439,465]" count="1" pageId="18" pageNumber="319" type="male">1 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFF35D539B97FAEC1896A18C" box="[678,720,439,465]" count="1" pageId="18" pageNumber="319" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF35D539BEFFAED1914A18C" box="[734,850,438,465]" pageId="18" pageNumber="319" value="2022-03-05">5.III.2022</date>
.
</paragraph>
<paragraph id="A1353685FFF35D5399F6FA811998A244" blockId="18.[151,1437,150,645]" pageId="18" pageNumber="319">
<emphasis id="93FEEA97FFF35D5399F6FA811852A1A9" bold="true" box="[199,532,474,501]" pageId="18" pageNumber="319">
Previous records in
<collectingCountry id="D99D7615FFF35D539883FA811849A1A9" box="[434,527,474,500]" name="Algeria" pageId="18" pageNumber="319">Algeria</collectingCountry>
:
</emphasis>
<collectingRegion id="634EF867FFF35D539B28FA8018CCA1A8" box="[537,650,475,501]" country="Algeria" name="Ain Defla" pageId="18" pageNumber="319">Aïn Defla</collectingRegion>
,
<collectingRegion id="634EF867FFF35D539BA2FA801893A1A8" box="[659,725,475,501]" country="Algeria" name="Alger" pageId="18" pageNumber="319">Alger</collectingRegion>
, El Meniaâ,
<collectingRegion id="634EF867FFF35D539A51FA80198BA1A8" box="[864,973,475,501]" country="Algeria" name="Laghouat" pageId="18" pageNumber="319">Laghouat</collectingRegion>
,
<collectingRegion id="634EF867FFF35D539AE6FA801E49A1A8" box="[983,1039,475,501]" country="Algeria" name="Mila" pageId="18" pageNumber="319">Mila</collectingRegion>
,
<collectingRegion id="634EF867FFF35D539D2BFA801E2FA1A8" box="[1050,1129,475,501]" country="Algeria" name="Naama" pageId="18" pageNumber="319">Naâma</collectingRegion>
(
<bibRefCitation id="C51B4B74FFF35D539D46FA801F45A1A8" author="Simon, E." box="[1143,1283,475,501]" pageId="18" pageNumber="319" pagination="82 - 87" refId="ref23337" refString="Simon, E. (1899) Liste des arachnides recueillis en Algerie par M. P. Lesne et descrition d'une espece nouvelle. Bulletin du Museum d'Histoire Naturelle Paris, 5, 82 - 87." type="journal article" year="1899">Simon 1899</bibRefCitation>
,
<bibRefCitation id="C51B4B74FFF35D539C3FFA801FDEA1A8" author="Strand, E." box="[1294,1432,475,501]" pageId="18" pageNumber="319" pagination="103 - 147" refId="ref23684" refString="Strand, E. (1907) Nordafrikanische, hauptsachlich von Carlo Freiherr von Erlanger gesammelte Thomisiden. Jahrbucher des Nassauischen Vereins fur Naturkunde, 60, 103 - 147. https: // doi. org / 10.5962 / bhl. part. 27907" type="journal article" year="1907">Strand 1907</bibRefCitation>
,
<bibRefCitation id="C51B4B74FFF35D5399A6FAA41B6EA244" author="Denis, J." box="[151,296,511,537]" pageId="18" pageNumber="319" pagination="1027 - 1060" refId="ref20639" refString="Denis, J. (1937 b) On a collection of spiders from Algeria. Proceedings of the Zoological Society of London, 106 (4), 1027 - 1060. https: // doi. org / 10.1111 / j. 1469 - 7998.1936. tb 06301. x" type="journal article" year="1937">Denis 1937b</bibRefCitation>
,
<bibRefCitation id="C51B4B74FFF35D539803FAA41BB9A244" author="Alioua, Y. &amp; Bissati, S. &amp; Kherbouche, O. &amp; Bosmans, R." box="[306,511,511,537]" pageId="18" pageNumber="319" pagination="33 - 40" refId="ref18687" refString="Alioua, Y., Bissati, S., Kherbouche, O. &amp; Bosmans, R. (2016) Spiders of Sebkhet El Melah (Northern Sahara, Algeria): review and new records. Serket, 15 (1), 33 - 40." type="journal article" year="2016">
Alioua
<emphasis id="93FEEA97FFF35D5398B4FAA41BFBA244" box="[389,445,511,537]" italics="true" pageId="18" pageNumber="319">et al.</emphasis>
2016
</bibRefCitation>
,
<bibRefCitation id="C51B4B74FFF35D539B3BFAA418BCA244" author="Boucherit, S. &amp; Bouragba, N. &amp; Douaoui, A. &amp; Bosmans, R." box="[522,762,511,537]" pageId="18" pageNumber="319" pagination="60 - 70" refId="ref19840" refString="Boucherit, S., Bouragba, N., Douaoui, A. &amp; Bosmans, R. (2020) Araignees de la plaine alluviale du Haut-Cheliff (Algerie), avec la citation de quelques especes nouvelles ou rares pour l'Algerie (Araneae: Gnaphosidae, Miturgidae). Nieuwsbrief van de Belgische Arachnologische Vereniging, 35 (1 - 2), 60 - 70." type="journal article" year="2020">
Boucherit
<emphasis id="93FEEA97FFF35D539BB1FAA418FEA244" box="[640,696,511,537]" italics="true" pageId="18" pageNumber="319">et al.</emphasis>
2020
</bibRefCitation>
, Alioua
<emphasis id="93FEEA97FFF35D539A67FAA419C9A244" box="[854,911,511,537]" italics="true" pageId="18" pageNumber="319">et al.</emphasis>
2022).
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFF35D5399F6F9791B78A2D8" pageId="18" pageNumber="319" type="distribution">
<paragraph id="A1353685FFF35D5399F6F9791B78A2D8" blockId="18.[151,1437,150,645]" pageId="18" pageNumber="319">
<emphasis id="93FEEA97FFF35D5399F6F9791B25A261" bold="true" box="[199,355,546,572]" pageId="18" pageNumber="319">Distribution:</emphasis>
Europe, North Africa,
<collectingCountry id="D99D7615FFF35D539B4EF9781896A260" box="[639,720,547,573]" name="Turkey" pageId="18" pageNumber="319">Turkey</collectingCountry>
, Caucasus,
<collectingCountry id="D99D7615FFF35D539A51F97819EDA260" box="[864,939,547,573]" name="Russia" pageId="18" pageNumber="319">Russia</collectingCountry>
(Europe to Far East), Middle East,
<collectingCountry id="D99D7615FFF35D539C59F9781FDEA260" box="[1384,1432,547,573]" name="Iran" pageId="18" pageNumber="319">Iran</collectingCountry>
,
<collectingCountry id="D99D7615FFF35D5399A6F91C1B5BA23C" box="[151,285,583,609]" name="Kazakhstan" pageId="18" pageNumber="319">Kazakhstan</collectingCountry>
, Central Asia,
<collectingCountry id="D99D7615FFF35D53988FF91C1842A23C" box="[446,516,583,609]" name="China" pageId="18" pageNumber="319">China</collectingCountry>
,
<collectingCountry id="D99D7615FFF35D539B3FF91C1813A23C" box="[526,597,583,609]" name="South Korea" pageId="18" pageNumber="319">Korea</collectingCountry>
,
<collectingCountry id="D99D7615FFF35D539B6EF91C18E4A23C" box="[607,674,583,609]" name="Japan" pageId="18" pageNumber="319">Japan</collectingCountry>
. Introduced to North America,
<collectingCountry id="D99D7615FFF35D539AC8F91C1ECFA23C" box="[1017,1161,583,609]" name="South Africa" pageId="18" pageNumber="319">South Africa</collectingCountry>
,
<collectingCountry id="D99D7615FFF35D539DA3F91C1EBFA23C" box="[1170,1273,583,609]" name="Australia" pageId="18" pageNumber="319">Australia</collectingCountry>
(
<bibRefCitation id="C51B4B74FFF35D539C37F91C1B74A2D8" author="World Spider Catalog" pageId="18" pageNumber="319" refId="ref24111" refString="World Spider Catalog (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern, Bern. Available from: https: // wsc. nmbe. ch / (accessed 1 June 2024) https: // doi. org / 10.24436 / 2" type="book" year="2024">World Spider Catalog 2024</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,163 @@
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<paragraph id="A1353685FFF15D5199A6FE0719A3A52B" blockId="16.[151,997,1372,1399]" box="[151,997,1372,1399]" pageId="16" pageNumber="317">
<heading id="FA7D81E9FFF15D5199A6FE0719A3A52B" bold="true" box="[151,997,1372,1399]" fontSize="11" level="1" pageId="16" pageNumber="317" reason="1">
<emphasis id="93FEEA97FFF15D5199A6FE0719A3A52B" bold="true" box="[151,997,1372,1399]" pageId="16" pageNumber="317">
<taxonomicName id="668A4D06FFF15D5199A6FE0718F1A52A" ID-CoL="48V3R" authority="Wunderlich, 1995" authorityName="Wunderlich" authorityYear="1995" box="[151,695,1372,1399]" class="Arachnida" family="Oecobiidae" genus="Oecobius" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="paulomaculatus">
<emphasis id="93FEEA97FFF15D5199A6FE071B97A52A" bold="true" box="[151,465,1372,1399]" italics="true" pageId="16" pageNumber="317">Oecobius paulomaculatus</emphasis>
<bibRefCitation id="C51B4B74FFF15D5198E9FE0618F1A52A" author="Wunderlich, J." box="[472,695,1373,1399]" pageId="16" pageNumber="317" pagination="585 - 608" refId="ref24159" refString="Wunderlich, J. (1995) Zu Taxonomie und Biogeographie der Arten der Gattung Oecobius Lucas 1846, mit Neubeschreibungen aus der Mediterraneis und von der Arabischen Halbinsel (Arachnida: Araneae: Oecobiidae). Beitrage zur Araneologie, 4 (1994), 585 - 608." type="journal article" year="1995">Wunderlich, 1995</bibRefCitation>
</taxonomicName>
(
<figureCitation id="39B12A00FFF15D519BF7FE071902A52A" box="[710,836,1372,1399]" captionStart="FIGURE 8" captionStartId="17.[152,255,695,719]" captionTargetBox="[151,1436,181,671]" captionTargetId="figure-17@17.[151,1436,181,671]" captionTargetPageId="17" captionText="FIGURE 8ac. Oecobius paulomaculatus, (a) Dorsal view of the male, (b) Dorsal view of the female, (c) Ventral view of the female." pageId="16" pageNumber="317">Figs. 8ac</figureCitation>
&amp; figs. 9af)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFF15D5199A6FEFA18D0A5E4" box="[151,662,1441,1465]" pageId="16" pageNumber="317" type="reference_group">
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<treatmentCitationGroup id="819A11ABFFF15D5199A6FEFA18D0A5E4" box="[151,662,1441,1465]" pageId="16" pageNumber="317">
<taxonomicName id="668A4D06FFF15D5199A6FEFA1BD9A5E4" ID-CoL="48V3R" authorityName="Wunderlich" authorityYear="1995" box="[151,415,1441,1465]" class="Arachnida" family="Oecobiidae" genus="Oecobius" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="paulomaculatus">
<emphasis id="93FEEA97FFF15D5199A6FEFA1BD9A5E4" box="[151,415,1441,1465]" italics="true" pageId="16" pageNumber="317">Oecobius paulomaculatus</emphasis>
</taxonomicName>
;
<treatmentCitation id="202B1094FFF15D51989DFEFA18D4A5E4" author="Wunderlich, J." box="[428,658,1441,1465]" page="596" pageId="16" pageNumber="317" year="1995">
<bibRefCitation id="C51B4B74FFF15D51989DFEFA18D4A5E4" author="Wunderlich, J." box="[428,658,1441,1465]" pageId="16" pageNumber="317" pagination="585 - 608" refId="ref24159" refString="Wunderlich, J. (1995) Zu Taxonomie und Biogeographie der Arten der Gattung Oecobius Lucas 1846, mit Neubeschreibungen aus der Mediterraneis und von der Arabischen Halbinsel (Arachnida: Araneae: Oecobiidae). Beitrage zur Araneologie, 4 (1994), 585 - 608." type="journal article" year="1995">Wunderlich 1995: 596</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFF15D5199A6FEBC1833A6CF" pageId="16" pageNumber="317" type="materials_examined">
<paragraph id="A1353685FFF15D5199A6FEBC1E95A617" blockId="16.[151,1436,1511,1790]" pageId="16" pageNumber="317">
<emphasis id="93FEEA97FFF15D5199A6FEBC1BC6A65C" bold="true" box="[151,384,1511,1537]" pageId="16" pageNumber="317">Material examined:</emphasis>
BERRIANE:Balouh, hand collecting,
<specimenCount id="B78CFD0CFFF15D519A1EFEB3191EA65F" box="[815,856,1512,1538]" count="4" pageId="16" pageNumber="317" type="male">4 ♂</specimenCount>
,
<date id="D5341045FFF15D519A53FEB3199FA65F" box="[866,985,1512,1538]" pageId="16" pageNumber="317" value="2021-02-12">12.II.2021</date>
;
<specimenCount id="B78CFD0CFFF15D519AD2FEB31E4DA65F" box="[995,1035,1512,1538]" count="1" pageId="16" pageNumber="317" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF15D519D27FEB31ED3A65F" box="[1046,1173,1512,1538]" pageId="16" pageNumber="317" value="2021-04-10">10.IV.2021</date>
;
<specimenCount id="B78CFD0CFFF15D519DAEFEB31E8EA65F" box="[1183,1224,1512,1538]" count="1" pageId="16" pageNumber="317" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFF15D519DE3FEB31F16A65F" box="[1234,1360,1512,1538]" pageId="16" pageNumber="317" value="2021-04-17">17.IV.2021</date>
, pitfall traps,
<specimenCount id="B78CFD0CFFF15D5199EDFD571B40A67B" box="[220,262,1548,1574]" count="1" pageId="16" pageNumber="317" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF15D519825FD501BCAA678" box="[276,396,1547,1573]" pageId="16" pageNumber="317" value="2021-02-26">26.II.2021</date>
. Sidi Mbarek, hand collecting,
<specimenCount id="B78CFD0CFFF15D519BDBFD571951A67B" box="[746,791,1548,1574]" count="1" pageId="16" pageNumber="317" type="male">1 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFF15D519A12FD57190BA67B" box="[803,845,1548,1574]" count="1" pageId="16" pageNumber="317" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF15D519A6AFD571997A67B" box="[859,977,1548,1574]" pageId="16" pageNumber="317" value="2021-02-11">11.II.2021</date>
;
<specimenCount id="B78CFD0CFFF15D519AEFFD571E4CA67B" box="[990,1034,1548,1574]" count="2" pageId="16" pageNumber="317" type="male">2 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFF15D519D27FD571E06A67B" box="[1046,1088,1548,1574]" count="4" pageId="16" pageNumber="317" type="female">4 ♀</specimenCount>
,
<date id="D5341045FFF15D519D7FFD571EF9A67B" box="[1102,1215,1548,1574]" pageId="16" pageNumber="317" value="2021-04-09">9.IV.2021</date>
;
<specimenCount id="B78CFD0CFFF15D519DFAFD571EB3A67B" box="[1227,1269,1548,1574]" count="1" pageId="16" pageNumber="317" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF15D519C32FD501FC4A678" box="[1283,1410,1547,1573]" pageId="16" pageNumber="317" value="2021-04-16">16.IV.2021</date>
;
<specimenCount id="B78CFD0CFFF15D519CBFFD571AEAA617" count="3" pageId="16" pageNumber="317" type="female">3 ♀</specimenCount>
,
<date id="D5341045FFF15D51998BFD6B1B7FA617" box="[186,313,1584,1610]" pageId="16" pageNumber="317" value="2021-04-23">23.IV.2021</date>
;
<specimenCount id="B78CFD0CFFF15D519874FD6B1B29A617" box="[325,367,1584,1610]" count="2" pageId="16" pageNumber="317" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFF15D51984CFD6B1BBAA617" box="[381,508,1584,1610]" pageId="16" pageNumber="317" value="2021-04-29">29.IV.2021</date>
. DHAYET BENDHAHOUA: Grazil, pitfall traps,
<specimenCount id="B78CFD0CFFF15D519D05FD6B1E26A617" box="[1076,1120,1584,1610]" count="1" pageId="16" pageNumber="317" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFF15D519D5DFD741E95A617" box="[1132,1235,1583,1610]" pageId="16" pageNumber="317" value="2022-02-05">5.II.2022</date>
</paragraph>
<paragraph id="A1353685FFF15D5199F6FD081833A6CF" blockId="16.[151,1436,1511,1790]" pageId="16" pageNumber="317">
<emphasis id="93FEEA97FFF15D5199F6FD08185EA630" bold="true" box="[199,536,1619,1646]" pageId="16" pageNumber="317">
Previous records in
<collectingCountry id="D99D7615FFF15D519887FD081855A630" box="[438,531,1619,1645]" name="Algeria" pageId="16" pageNumber="317">Algeria</collectingCountry>
:
</emphasis>
This species is endemic to
<collectingRegion id="634EF867FFF15D519A63FD0819FFA630" box="[850,953,1619,1645]" country="Algeria" name="Ghardaia" pageId="16" pageNumber="317">Ghardaïa</collectingRegion>
(
<bibRefCitation id="C51B4B74FFF15D519AF9FD081ED7A633" author="Wunderlich, J." box="[968,1169,1619,1646]" pageId="16" pageNumber="317" pagination="585 - 608" refId="ref24159" refString="Wunderlich, J. (1995) Zu Taxonomie und Biogeographie der Arten der Gattung Oecobius Lucas 1846, mit Neubeschreibungen aus der Mediterraneis und von der Arabischen Halbinsel (Arachnida: Araneae: Oecobiidae). Beitrage zur Araneologie, 4 (1994), 585 - 608." type="journal article" year="1995">Wunderlich 1995</bibRefCitation>
). Until now it was only known from the palm yard of Beni Isguen.
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFF15D5199F6FDC01917A684" pageId="16" pageNumber="317" type="distribution">
<paragraph id="A1353685FFF15D5199F6FDC01917A684" blockId="16.[151,1436,1511,1790]" pageId="16" pageNumber="317">
<emphasis id="93FEEA97FFF15D5199F6FDC01B25A6E8" bold="true" box="[199,355,1691,1717]" pageId="16" pageNumber="317">Distribution:</emphasis>
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(
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). Besides Beni Isguen, the species has now also been collected in palm groves of Berriane and Dhayet Bendhahoua.
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<paragraph id="A1353685FFF15D5199F6FDB81E27A6A0" blockId="16.[151,1436,1511,1790]" box="[199,1121,1763,1790]" pageId="16" pageNumber="317">
<emphasis id="93FEEA97FFF15D5199F6FDB81B01A6A0" bold="true" box="[199,327,1763,1789]" pageId="16" pageNumber="317">Comment:</emphasis>
This study presents the photos of
<taxonomicName id="668A4D06FFF15D519BF7FDBF19DDA6A0" authorityName="Wunderlich" authorityYear="1995" box="[710,923,1763,1790]" class="Arachnida" family="Oecobiidae" genus="Oecobius" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="paulomaculatus">
<emphasis id="93FEEA97FFF15D519BF7FDBF189CA6A3" box="[710,730,1764,1790]" italics="true" pageId="16" pageNumber="317">O</emphasis>
.
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View file

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BERRIANE: Balouh, hand collecting,
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Known only from
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2022).
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Algeria, Tunisia, Egypt (
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<paragraph id="A1353685FFF45D5499A6FF341844A4DB" blockId="21.[151,514,1134,1158]" box="[151,514,1134,1158]" pageId="21" pageNumber="322">
<treatmentCitationGroup id="819A11ABFFF45D5499A6FF341844A4DB" box="[151,514,1134,1158]" pageId="21" pageNumber="322">
<taxonomicName id="668A4D06FFF45D5499A6FF341B0AA4DB" ID-CoL="6RMQM" baseAuthorityName="Simon" baseAuthorityYear="1868" box="[151,332,1134,1158]" class="Arachnida" family="Salticidae" genus="Mogrus" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="neglectus">
<emphasis id="93FEEA97FFF45D5499A6FF341B0AA4DB" box="[151,332,1134,1158]" italics="true" pageId="21" pageNumber="322">Mogrus neglectus</emphasis>
</taxonomicName>
;
<treatmentCitation id="202B1094FFF45D549868FF351BB8A4DB" author="Simon, E." box="[345,510,1134,1158]" page="85" pageId="21" pageNumber="322" year="1899">
<bibRefCitation id="C51B4B74FFF45D549868FF351BB8A4DB" author="Simon, E." box="[345,510,1134,1158]" pageId="21" pageNumber="322" pagination="82 - 87" refId="ref23337" refString="Simon, E. (1899) Liste des arachnides recueillis en Algerie par M. P. Lesne et descrition d'une espece nouvelle. Bulletin du Museum d'Histoire Naturelle Paris, 5, 82 - 87." type="journal article" year="1899">Simon 1899: 85</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFF45D5499A6FFEF198CA54A" pageId="21" pageNumber="322" type="materials_examined">
<paragraph id="A1353685FFF45D5499A6FFEF1B02A4AE" blockId="21.[151,1436,1204,1375]" pageId="21" pageNumber="322">
<emphasis id="93FEEA97FFF45D5499A6FFEF1BC5A493" bold="true" box="[151,387,1204,1230]" pageId="21" pageNumber="322">Material examined:</emphasis>
EL GUERRARA: Aghzou, hand collecting,
<specimenCount id="B78CFD0CFFF45D549A46FFEE19E5A492" box="[887,931,1205,1231]" count="1" pageId="21" pageNumber="322" type="male">1 ♂</specimenCount>
,
<specimenCount id="B78CFD0CFFF45D549A9EFFEE199FA492" box="[943,985,1205,1231]" count="1" pageId="21" pageNumber="322" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF45D549AD7FFEE1E21A492" box="[998,1127,1205,1231]" pageId="21" pageNumber="322" value="2020-03-27">27.III.2020</date>
. El Amied, hand collecting,
<specimenCount id="B78CFD0CFFF45D5499A6FF821A87A4AE" box="[151,193,1241,1267]" count="1" pageId="21" pageNumber="322" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF45D5499FEFF821B07A4AE" box="[207,321,1241,1267]" pageId="21" pageNumber="322" value="2020-04-01">1.IV.2020</date>
.
</paragraph>
<paragraph id="A1353685FFF45D5499F6FFA7198CA54A" blockId="21.[151,1436,1204,1375]" box="[199,970,1276,1303]" pageId="21" pageNumber="322">
<emphasis id="93FEEA97FFF45D5499F6FFA71853A54B" bold="true" box="[199,533,1276,1303]" pageId="21" pageNumber="322">
Previous records in
<collectingCountry id="D99D7615FFF45D549882FFA71856A54B" box="[435,528,1276,1302]" name="Algeria" pageId="21" pageNumber="322">Algeria</collectingCountry>
:
</emphasis>
Known only from
<collectingRegion id="634EF867FFF45D549BDDFFA71967A54B" box="[748,801,1276,1302]" country="Algeria" name="Setif" pageId="21" pageNumber="322">Sétif</collectingRegion>
(
<bibRefCitation id="C51B4B74FFF45D549A1EFFA719FBA54A" author="Simon, E." box="[815,957,1276,1303]" pageId="21" pageNumber="322" pagination="82 - 87" refId="ref23337" refString="Simon, E. (1899) Liste des arachnides recueillis en Algerie par M. P. Lesne et descrition d'une espece nouvelle. Bulletin du Museum d'Histoire Naturelle Paris, 5, 82 - 87." type="journal article" year="1899">Simon 1899</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFF45D5499F6FE7B1B78A502" pageId="21" pageNumber="322" type="distribution">
<paragraph id="A1353685FFF45D5499F6FE7B1B78A502" blockId="21.[151,1436,1204,1375]" pageId="21" pageNumber="322">
<emphasis id="93FEEA97FFF45D5499F6FE7B1B25A567" bold="true" box="[199,355,1312,1338]" pageId="21" pageNumber="322">Distribution:</emphasis>
North
<collectingCountry id="D99D7615FFF45D549889FE7B1871A567" box="[440,567,1312,1338]" name="Macedonia" pageId="21" pageNumber="322">Macedonia</collectingCountry>
,
<collectingCountry id="D99D7615FFF45D549B74FE7A18D1A566" box="[581,663,1313,1339]" name="Greece" pageId="21" pageNumber="322">Greece</collectingCountry>
,
<collectingCountry id="D99D7615FFF45D549B94FE7B18B0A567" box="[677,758,1312,1338]" name="Turkey" pageId="21" pageNumber="322">Turkey</collectingCountry>
,
<collectingCountry id="D99D7615FFF45D549A35FE7A191EA566" box="[772,856,1313,1339]" name="Cyprus" pageId="21" pageNumber="322">Cyprus</collectingCountry>
,
<collectingCountry id="D99D7615FFF45D549A57FE7B19E1A567" box="[870,935,1312,1338]" name="Israel" pageId="21" pageNumber="322">Israel</collectingCountry>
, Caucasus,
<collectingCountry id="D99D7615FFF45D549D01FE7A1E26A566" box="[1072,1120,1313,1339]" name="Iran" pageId="21" pageNumber="322">Iran</collectingCountry>
,
<collectingCountry id="D99D7615FFF45D549D5FFE7B1EB7A567" box="[1134,1265,1312,1338]" name="Kazakhstan" pageId="21" pageNumber="322">Kazakhstan</collectingCountry>
(
<bibRefCitation id="C51B4B74FFF45D549C33FE7B1B74A502" author="World Spider Catalog" pageId="21" pageNumber="322" refId="ref24111" refString="World Spider Catalog (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern, Bern. Available from: https: // wsc. nmbe. ch / (accessed 1 June 2024) https: // doi. org / 10.24436 / 2" type="book" year="2024">World Spider Catalog 2024</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,127 @@
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<mods:title id="0B16C3BD700E69E0E23BDA7B67BA8622">Spiders of arid lands: The Ghardaïa region (Northern Sahara) with seven new records for Algeria</mods:title>
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<mods:namePart id="06FDA1B7F4C6945E3C29EA9C75E94E4D">Bosmans, Robert</mods:namePart>
<mods:affiliation id="220CD7C909D0EA1D12CFFC4ED7837AE7">Terrestrial Ecology Unit, Ledeganckstraat 35, B- 9000 Gent, Belgium</mods:affiliation>
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<subSubSection id="E990650EFFF55D5599A6FD9918C4A686" box="[151,642,1729,1756]" pageId="20" pageNumber="321" type="nomenclature">
<paragraph id="A1353685FFF55D5599A6FD9918C4A686" blockId="20.[151,642,1729,1756]" box="[151,642,1729,1756]" pageId="20" pageNumber="321">
<heading id="FA7D81E9FFF55D5599A6FD9918C4A686" bold="true" box="[151,642,1729,1756]" fontSize="11" level="1" pageId="20" pageNumber="321" reason="1">
<taxonomicName id="668A4D06FFF55D5599A6FD9918C4A686" ID-CoL="3PB48" authority="Alicata &amp; Cantarella, 1994" authorityName="Alicata &amp; Cantarella" authorityYear="1994" box="[151,642,1729,1756]" class="Arachnida" family="Salticidae" genus="Icius" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="simoni">
<emphasis id="93FEEA97FFF55D5599A6FD9918C4A686" bold="true" box="[151,642,1729,1756]" pageId="20" pageNumber="321">
<emphasis id="93FEEA97FFF55D5599A6FD991B6BA681" bold="true" box="[151,301,1730,1756]" italics="true" pageId="20" pageNumber="321">Icius simoni</emphasis>
<bibRefCitation id="C51B4B74FFF55D559803FD9A18C4A686" author="Alicata, P. &amp; Cantarella, T." box="[306,642,1729,1755]" pageId="20" pageNumber="321" pagination="111 - 131" refId="ref18643" refString="Alicata, P. &amp; Cantarella, T. (1994) The Euro-mediterranean species of Icius (Araneae, Salticidae): a critical revision and description of two new species. Animalia, 20 (1993), 111 - 131." type="journal article" year="1994">Alicata &amp; Cantarella, 1994</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFF55D5599A6FC5C196AA743" box="[151,812,1798,1823]" pageId="20" pageNumber="321" type="reference_group">
<paragraph id="A1353685FFF55D5599A6FC5C196AA743" blockId="20.[151,812,1798,1823]" box="[151,812,1798,1823]" pageId="20" pageNumber="321">
<treatmentCitationGroup id="819A11ABFFF55D5599A6FC5C196AA743" box="[151,812,1798,1823]" pageId="20" pageNumber="321">
<taxonomicName id="668A4D06FFF55D5599A6FC5C1B55A743" ID-CoL="3PB48" authorityName="Alicata &amp; Cantarella" authorityYear="1994" box="[151,275,1799,1822]" class="Arachnida" family="Salticidae" genus="Icius" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="simoni">
<emphasis id="93FEEA97FFF55D5599A6FC5C1B55A743" box="[151,275,1799,1822]" italics="true" pageId="20" pageNumber="321">Icius simoni</emphasis>
</taxonomicName>
;
<treatmentCitation id="202B1094FFF55D55982FFC5D1819A743" author="Alicata, P. &amp; Cantarella, T." box="[286,607,1798,1822]" page="127" pageId="20" pageNumber="321" year="1994">
<bibRefCitation id="C51B4B74FFF55D55982FFC5D1819A743" author="Alicata, P. &amp; Cantarella, T." box="[286,607,1798,1822]" pageId="20" pageNumber="321" pagination="111 - 131" refId="ref18643" refString="Alicata, P. &amp; Cantarella, T. (1994) The Euro-mediterranean species of Icius (Araneae, Salticidae): a critical revision and description of two new species. Animalia, 20 (1993), 111 - 131." type="journal article" year="1994">Alicata &amp; Cantarella 1994: 127</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="202B1094FFF55D559B5BFC5C196EA743" author="Logunov, D. V." box="[618,808,1798,1823]" page="68" pageId="20" pageNumber="321" year="2015">
<bibRefCitation id="C51B4B74FFF55D559B5BFC5C196EA743" author="Logunov, D. V." box="[618,808,1798,1823]" pageId="20" pageNumber="321" pagination="33 - 85" refId="ref21696" refString="Logunov, D. V. (2015) Taxonomic-faunistic notes on the jumping spiders of the Mediterranean (Aranei: Salticidae). Arthropoda Selecta, 24 (1), 33 - 85. https: // doi. org / 10.15298 / arthsel. 24.1.03" type="journal article" year="2015">Logunov 2015: 68</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFF55D5599A6FC171956A78F" pageId="20" pageNumber="321" type="materials_examined">
<paragraph id="A1353685FFF55D5599A6FC171FDAA7D7" blockId="20.[151,1436,1868,2002]" pageId="20" pageNumber="321">
<emphasis id="93FEEA97FFF55D5599A6FC171BC5A73B" bold="true" box="[151,387,1868,1894]" pageId="20" pageNumber="321">Material examined:</emphasis>
EL ATTEUF: Palm grove of El Atteuf, pitfall traps;
<specimenCount id="B78CFD0CFFF55D559AE1FC1719BCA73A" box="[976,1018,1868,1895]" count="1" pageId="20" pageNumber="321" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFF55D559D39FC171EC6A73A" box="[1032,1152,1868,1895]" pageId="20" pageNumber="321" value="2015-02-16">16.II.2015</date>
;
<materialsCitation id="11E23CD8FFF55D559DBAFC171FDEA7D7" collectingDate="2015-03-06" country="Algeria" county="Previous" location="Algeria" municipality="The" pageId="20" pageNumber="321" specimenCount="2" specimenCount-female="2" stateProvince="Biskra">
<specimenCount id="B78CFD0CFFF55D559DBAFC171EF3A73A" box="[1163,1205,1868,1895]" count="2" pageId="20" pageNumber="321" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFF55D559DF2FC171F71A73A" box="[1219,1335,1868,1895]" pageId="20" pageNumber="321" value="2015-03-06">
<collectingDate id="C570E9ADFFF55D559DF2FC171F71A73A" box="[1219,1335,1868,1895]" pageId="20" pageNumber="321" value="2015-03-06">6.III.2015</collectingDate>
</date>
.
<emphasis id="93FEEA97FFF55D5599F6FC2B185DA7D7" bold="true" box="[199,539,1904,1930]" pageId="20" pageNumber="321">
<collectingCounty id="48544E09FFF55D5599F6FC2B1B69A7D7" box="[199,303,1904,1930]" pageId="20" pageNumber="321">Previous</collectingCounty>
records in
<collectingCountry id="D99D7615FFF55D559888FC2B1850A7D7" box="[441,534,1904,1930]" name="Algeria" pageId="20" pageNumber="321">Algeria</collectingCountry>
:
</emphasis>
<collectingMunicipality id="4151ACFFFFF55D559B15FC2B1816A7D7" box="[548,592,1904,1930]" pageId="20" pageNumber="321">The</collectingMunicipality>
species is endemic to
<location id="A455605EFFF55D559A62FC2B19E1A7D7" LSID="urn:lsid:plazi:treatment:29238793FFF55D5599A6FD991956A78F:A455605EFFF55D559A62FC2B19E1A7D7" box="[851,935,1904,1930]" country="Algeria" county="Previous" municipality="The" name="Algeria" pageId="20" pageNumber="321" stateProvince="Biskra">Algeria</location>
and known previously from
<collectingRegion id="634EF867FFF55D559DC4FC2B1F07A7D7" box="[1269,1345,1904,1930]" country="Algeria" name="Biskra" pageId="20" pageNumber="321">Biskra</collectingRegion>
,
<collectingRegion id="634EF867FFF55D559C7FFC2B1FDEA7D7" box="[1358,1432,1904,1930]" country="Algeria" name="M'Sila" pageId="20" pageNumber="321">Msila</collectingRegion>
</materialsCitation>
,
</paragraph>
<paragraph id="A1353685FFF55D5599A6FCCF1956A78F" blockId="20.[151,1436,1868,2002]" pageId="20" pageNumber="321">
<collectingRegion id="634EF867FFF55D5599A6FCCF1ABFA7F3" box="[151,249,1940,1966]" country="Algeria" name="Tlemcen" pageId="20" pageNumber="321">Tlemcen</collectingRegion>
(
<bibRefCitation id="C51B4B74FFF55D559839FCCF1875A7F3" author="Alicata, P. &amp; Cantarella, T." box="[264,563,1940,1966]" pageId="20" pageNumber="321" pagination="111 - 131" refId="ref18643" refString="Alicata, P. &amp; Cantarella, T. (1994) The Euro-mediterranean species of Icius (Araneae, Salticidae): a critical revision and description of two new species. Animalia, 20 (1993), 111 - 131." type="journal article" year="1994">Alicata &amp; Cantarella 1994</bibRefCitation>
;
<bibRefCitation id="C51B4B74FFF55D559B0EFCCE18A3A7F2" author="Logunov, D. V." box="[575,741,1940,1967]" pageId="20" pageNumber="321" pagination="33 - 85" refId="ref21696" refString="Logunov, D. V. (2015) Taxonomic-faunistic notes on the jumping spiders of the Mediterranean (Aranei: Salticidae). Arthropoda Selecta, 24 (1), 33 - 85. https: // doi. org / 10.15298 / arthsel. 24.1.03" type="journal article" year="2015">Logunov 2015</bibRefCitation>
)
<emphasis id="93FEEA97FFF55D5599F6FCE31B25A78F" bold="true" box="[199,355,1976,2002]" pageId="20" pageNumber="321">Distribution:</emphasis>
<collectingCountry id="D99D7615FFF55D559859FCE31BFAA78F" box="[360,444,1976,2002]" name="Algeria" pageId="20" pageNumber="321">Algeria</collectingCountry>
(
<bibRefCitation id="C51B4B74FFF55D5598FBFCE31942A78F" author="World Spider Catalog" box="[458,772,1976,2002]" pageId="20" pageNumber="321" refId="ref24111" refString="World Spider Catalog (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern, Bern. Available from: https: // wsc. nmbe. ch / (accessed 1 June 2024) https: // doi. org / 10.24436 / 2" type="book" year="2024">World Spider Catalog 2024</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="506BDC81A3B72DD00E57C034F33CA318">Spiders of arid lands: The Ghardaïa region (Northern Sahara) with seven new records for Algeria</mods:title>
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<taxonomicName id="668A4D06FFF85D5899A6FD401976A66B" ID-CoL="4ZMP9" authority="(O. Pickard-Cambridge, 1872)" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1872" box="[151,816,1563,1590]" class="Arachnida" family="Theridiidae" genus="Steatoda" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="erigoniformis">
<emphasis id="93FEEA97FFF85D5899A6FD401BEAA668" bold="true" box="[151,428,1563,1590]" italics="true" pageId="25" pageNumber="326">Steatoda erigoniformis</emphasis>
(O.
<bibRefCitation id="C51B4B74FFF85D5898D3FD47196EA66B" author="Pickard-Cambridge, O." box="[482,808,1564,1590]" pageId="25" pageNumber="326" pagination="212 - 354" refId="ref22251" refString="Pickard-Cambridge, O. (1872) General list of the spiders of Palestine and Syria, with descriptions of numerous new species, and characters of two new genera. Proceedings of the Zoological Society of London, 40 (1), 212 - 354. https: // doi. org / 10.1111 / j. 1469 - 7998.1872. tb 00489. x" type="journal article" year="1872">Pickard-Cambridge, 1872</bibRefCitation>
)
</taxonomicName>
(
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)
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<taxonomicName id="668A4D06FFF85D5899A6FD3B1BC3A624" authorityName="O.P.-Cambridge" authorityYear="1872" box="[151,389,1632,1657]" class="Arachnida" family="Theridiidae" genus="Theridion" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="erigoniforme">
<emphasis id="93FEEA97FFF85D5899A6FD3B1BC3A624" box="[151,389,1632,1657]" italics="true" pageId="25" pageNumber="326">Theridion erigoniforme</emphasis>
</taxonomicName>
; O.
<treatmentCitation id="202B1094FFF85D589883FD3B18AEA624" author="Pickard-Cambridge, O." box="[434,744,1632,1657]" page="284" pageId="25" pageNumber="326" year="1872">
<bibRefCitation id="C51B4B74FFF85D589883FD3B18AEA624" author="Pickard-Cambridge, O." box="[434,744,1632,1657]" pageId="25" pageNumber="326" pagination="212 - 354" refId="ref22251" refString="Pickard-Cambridge, O. (1872) General list of the spiders of Palestine and Syria, with descriptions of numerous new species, and characters of two new genera. Proceedings of the Zoological Society of London, 40 (1), 212 - 354. https: // doi. org / 10.1111 / j. 1469 - 7998.1872. tb 00489. x" type="journal article" year="1872">Pickard-Cambridge 1872: 284</bibRefCitation>
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.
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</paragraph>
<paragraph id="A1353685FFF85D5899A6FD2418E1A6CA" blockId="25.[151,1051,1632,1841]" box="[151,679,1663,1687]" pageId="25" pageNumber="326">
<treatmentCitationGroup id="819A11ABFFF85D5899A6FD2418E1A6CA" box="[151,679,1663,1687]" pageId="25" pageNumber="326">
<taxonomicName id="668A4D06FFF85D5899A6FD241B07A6CA" ID-CoL="4ZMRT" authorityName="O. Pickard-Cambridge" authorityYear="1876" box="[151,321,1663,1687]" class="Arachnida" family="Theridiidae" genus="Steatoda" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="signata">
<emphasis id="93FEEA97FFF85D5899A6FD241B07A6CA" box="[151,321,1663,1687]" italics="true" pageId="25" pageNumber="326">Steatoda signata</emphasis>
</taxonomicName>
; O.
<treatmentCitation id="202B1094FFF85D58985CFD2418E5A6CA" author="Pickard-Cambridge, O." box="[365,675,1663,1687]" page="568" pageId="25" pageNumber="326" year="1876">
<bibRefCitation id="C51B4B74FFF85D58985CFD2418E5A6CA" author="Pickard-Cambridge, O." box="[365,675,1663,1687]" pageId="25" pageNumber="326" pagination="541 - 630" refId="ref22318" refString="Pickard-Cambridge, O. (1876) Catalogue of a collection of spiders made in Egypt, with descriptions of new species and characters of a new genus. Proceedings of the Zoological Society of London, 44 (3), 541 - 630. https: // doi. org / 10.1111 / j. 1096 - 3642.1876. tb 02595. x" type="journal article" year="1876">Pickard-Cambridge 1876: 568</bibRefCitation>
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.
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</paragraph>
<paragraph id="A1353685FFF85D5899A6FDC51851A6EB" blockId="25.[151,1051,1632,1841]" box="[151,535,1694,1718]" pageId="25" pageNumber="326">
<treatmentCitationGroup id="819A11ABFFF85D5899A6FDC51851A6EB" box="[151,535,1694,1718]" pageId="25" pageNumber="326">
<taxonomicName id="668A4D06FFF85D5899A6FDC51B13A6E8" authorityName="Simon" authorityYear="1881" box="[151,341,1694,1718]" class="Arachnida" family="Theridiidae" genus="Crustulina" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="signata">
<emphasis id="93FEEA97FFF85D5899A6FDC51B13A6E8" box="[151,341,1694,1718]" italics="true" pageId="25" pageNumber="326">Crustulina signata</emphasis>
</taxonomicName>
;
<treatmentCitation id="202B1094FFF85D589853FDC51855A6EB" author="Simon, E." box="[354,531,1694,1718]" page="161" pageId="25" pageNumber="326" year="1881">
<bibRefCitation id="C51B4B74FFF85D589853FDC51855A6EB" author="Simon, E." box="[354,531,1694,1718]" pageId="25" pageNumber="326" refId="ref23124" refString="Simon, E. (1881) Les arachnides de France. Tome Cinquieme, Premiere Partie. Roret, Paris, 180 pp." type="book" year="1881">Simon 1881: 161</bibRefCitation>
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.
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</paragraph>
<paragraph id="A1353685FFF85D5899A6FDE718F9A688" blockId="25.[151,1051,1632,1841]" box="[151,703,1724,1749]" pageId="25" pageNumber="326">
<treatmentCitationGroup id="819A11ABFFF85D5899A6FDE718F9A688" box="[151,703,1724,1749]" pageId="25" pageNumber="326">
<taxonomicName id="668A4D06FFF85D5899A6FDE71B88A689" authorityName="Keyserling" authorityYear="1884" box="[151,462,1724,1748]" class="Arachnida" family="Theridiidae" genus="Lithyphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="septemmaculatus">
<emphasis id="93FEEA97FFF85D5899A6FDE71B88A689" box="[151,462,1724,1748]" italics="true" pageId="25" pageNumber="326">Lithyphantes septemmaculatus</emphasis>
</taxonomicName>
;
<treatmentCitation id="202B1094FFF85D5898EDFDE718FDA688" author="Keyserling, E." box="[476,699,1724,1749]" page="141" pageId="25" pageNumber="326" year="1884">
<bibRefCitation id="C51B4B74FFF85D5898EDFDE718FDA688" author="Keyserling, E." box="[476,699,1724,1749]" pageId="25" pageNumber="326" refId="ref21026" refString="Keyserling, E. (1884) Die Spinnen Amerikas II. Halfte. Theridiidae. Bauer &amp; Raspe, Nurnberg, 222 pp. https: // doi. org / 10.5962 / bhl. title. 64832" type="book" year="1884">Keyserling 1884: 141</bibRefCitation>
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.
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</paragraph>
<paragraph id="A1353685FFF85D5899A6FD8018CFA6AE" blockId="25.[151,1051,1632,1841]" box="[151,649,1755,1779]" pageId="25" pageNumber="326">
<treatmentCitationGroup id="819A11ABFFF85D5899A6FD8018CFA6AE" box="[151,649,1755,1779]" pageId="25" pageNumber="326">
<taxonomicName id="668A4D06FFF85D5899A6FD801BDEA6AE" authorityName="Schenkel" authorityYear="1937" box="[151,408,1755,1779]" class="Arachnida" family="Theridiidae" genus="Asagenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="erigoniformis">
<emphasis id="93FEEA97FFF85D5899A6FD801BDEA6AE" box="[151,408,1755,1779]" italics="true" pageId="25" pageNumber="326">Asagenella erigoniformis</emphasis>
</taxonomicName>
;
<treatmentCitation id="202B1094FFF85D589894FD8018C3A6AE" author="Levi, H. W. &amp; Levi, L. R." box="[421,645,1755,1779]" page="60" pageId="25" pageNumber="326" year="1962">
<bibRefCitation id="C51B4B74FFF85D589894FD8018C3A6AE" author="Levi, H. W. &amp; Levi, L. R." box="[421,645,1755,1779]" pageId="25" pageNumber="326" pagination="1 - 71" refId="ref21288" refString="Levi, H. W. &amp; Levi, L. R. (1962) The genera of the spider family Theridiidae. Bulletin of the Museum of Comparative Zoology, 127, 1 - 71." type="journal article" year="1962">Levi &amp; Levi 1962: 60</bibRefCitation>
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.
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</paragraph>
<paragraph id="A1353685FFF85D5899A6FDA11865A74F" blockId="25.[151,1051,1632,1841]" box="[151,547,1786,1810]" pageId="25" pageNumber="326">
<treatmentCitationGroup id="819A11ABFFF85D5899A6FDA11865A74F" box="[151,547,1786,1810]" pageId="25" pageNumber="326">
<taxonomicName id="668A4D06FFF85D5899A6FDA11BC6A74F" ID-CoL="4ZMP9" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1872" box="[151,384,1786,1810]" class="Arachnida" family="Theridiidae" genus="Steatoda" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="erigoniformis">
<emphasis id="93FEEA97FFF85D5899A6FDA11BC6A74F" box="[151,384,1786,1810]" italics="true" pageId="25" pageNumber="326">Steatoda erigoniformis</emphasis>
</taxonomicName>
;
<treatmentCitation id="202B1094FFF85D5898BCFDA11858A74F" author="Levi, H. W. &amp; Levi, L. R." box="[397,542,1786,1810]" page="25" pageId="25" pageNumber="326" year="1962">
<bibRefCitation id="C51B4B74FFF85D5898BCFDA11858A74F" author="Levi, H. W. &amp; Levi, L. R." box="[397,542,1786,1810]" pageId="25" pageNumber="326" pagination="1 - 71" refId="ref21288" refString="Levi, H. W. &amp; Levi, L. R. (1962) The genera of the spider family Theridiidae. Bulletin of the Museum of Comparative Zoology, 127, 1 - 71." type="journal article" year="1962">Levi 1962: 25</bibRefCitation>
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.
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</paragraph>
<paragraph id="A1353685FFF85D5899A6FC421E5DA76C" blockId="25.[151,1051,1632,1841]" box="[151,1051,1816,1841]" pageId="25" pageNumber="326">
For a complete list of taxonomic references, please see the
<bibRefCitation id="C51B4B74FFF85D589BDCFC431E50A76C" author="World Spider Catalog" box="[749,1046,1816,1841]" pageId="25" pageNumber="326" refId="ref24111" refString="World Spider Catalog (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern, Bern. Available from: https: // wsc. nmbe. ch / (accessed 1 June 2024) https: // doi. org / 10.24436 / 2" type="book" year="2024">World Spider Catalog (2024)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFF85D5B99A6FC041BF9A088" lastPageId="26" lastPageNumber="327" pageId="25" pageNumber="326" type="description">
<paragraph id="A1353685FFF85D5899A6FC041FDAA7B8" blockId="25.[151,1437,1887,2021]" pageId="25" pageNumber="326">
In
<collectingCountry id="D99D7615FFF85D589989FC041B49A724" box="[184,271,1887,1913]" name="Algeria" pageId="25" pageNumber="326">Algeria</collectingCountry>
, the genus
<taxonomicName id="668A4D06FFF85D5898A7FC0418FBA724" authority="Sundevall, 1833" authorityName="Sundevall" authorityYear="1833" box="[406,701,1887,1913]" class="Arachnida" family="Theridiidae" genus="Steatoda" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="326" phylum="Arthropoda" rank="genus">
<emphasis id="93FEEA97FFF85D5898A7FC041BBEA724" box="[406,504,1887,1913]" italics="true" pageId="25" pageNumber="326">Steatoda</emphasis>
Sundevall, 1833
</taxonomicName>
is represented by 8 species:
<taxonomicName id="668A4D06FFF85D589D25FC041FD1A724" authority="(De Geer, 1778)" baseAuthorityName="De Geer" baseAuthorityYear="1778" box="[1044,1431,1887,1913]" class="Arachnida" family="Theridiidae" genus="Steatoda" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="albomaculata">
<emphasis id="93FEEA97FFF85D589D25FC041E64A724" box="[1044,1058,1887,1913]" italics="true" pageId="25" pageNumber="326">S</emphasis>
.
<emphasis id="93FEEA97FFF85D589D02FC041E8BA724" box="[1075,1229,1887,1913]" italics="true" pageId="25" pageNumber="326">albomaculata</emphasis>
(De Geer, 1778)
</taxonomicName>
,
<taxonomicName id="668A4D06FFF85D5899A6FCD81BAEA7C0" authority="(Thorell, 1875)" baseAuthorityName="Thorell" baseAuthorityYear="1875" box="[151,488,1922,1949]" class="Arachnida" family="Theridiidae" genus="Steatoda" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="ephippiata">
<emphasis id="93FEEA97FFF85D5899A6FCD81AE3A7C0" box="[151,165,1923,1949]" italics="true" pageId="25" pageNumber="326">S</emphasis>
.
<emphasis id="93FEEA97FFF85D589987FCD81B68A7C0" box="[182,302,1923,1949]" italics="true" pageId="25" pageNumber="326">ephippiata</emphasis>
(
<bibRefCitation id="C51B4B74FFF85D58980EFCD81B99A7C0" author="Thorell, T." box="[319,479,1922,1949]" pageId="25" pageNumber="326" pagination="1 - 204" refId="ref23836" refString="Thorell, T. (1875) Descriptions of several European and North African spiders. Kongliga Svenska Vetenskaps-Akademiens Handlingar, 13 (5), 1 - 204." type="journal article" year="1875">Thorell, 1875</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="668A4D06FFF85D5898C7FCD8190EA7C0" authority="(C. L. Koch, 1838)" baseAuthorityName="C. L. Koch" baseAuthorityYear="1838" box="[502,840,1923,1949]" class="Arachnida" family="Theridiidae" genus="Steatoda" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="grossa">
<emphasis id="93FEEA97FFF85D5898C7FCD81842A7C0" box="[502,516,1923,1949]" italics="true" pageId="25" pageNumber="326">S</emphasis>
.
<emphasis id="93FEEA97FFF85D589B24FCD81819A7C1" box="[533,607,1923,1948]" italics="true" pageId="25" pageNumber="326">grossa</emphasis>
(C. L. Koch, 1838)
</taxonomicName>
,
<taxonomicName id="668A4D06FFF85D589A66FCD81EE2A7C0" authority="(Simon, 1873)" baseAuthorityName="Simon" baseAuthorityYear="1873" box="[855,1188,1923,1949]" class="Arachnida" family="Theridiidae" genus="Steatoda" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="latifasciata">
<emphasis id="93FEEA97FFF85D589A66FCD81923A7C0" box="[855,869,1923,1949]" italics="true" pageId="25" pageNumber="326">S</emphasis>
.
<emphasis id="93FEEA97FFF85D589A47FCD819B2A7C0" box="[886,1012,1923,1949]" italics="true" pageId="25" pageNumber="326">latifasciata</emphasis>
(Simon, 1873)
</taxonomicName>
,
<taxonomicName id="668A4D06FFF85D589D82FCD81A9DA79C" authority="(Thorell, 1875)" baseAuthorityName="Thorell" baseAuthorityYear="1875" class="Arachnida" family="Theridiidae" genus="Steatoda" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="moerens">
<emphasis id="93FEEA97FFF85D589D82FCD81E87A7C0" box="[1203,1217,1923,1949]" italics="true" pageId="25" pageNumber="326">S</emphasis>
.
<emphasis id="93FEEA97FFF85D589DE3FCD81F76A7C1" box="[1234,1328,1923,1948]" italics="true" pageId="25" pageNumber="326">moerens</emphasis>
(
<bibRefCitation id="C51B4B74FFF85D589C70FCD81A94A79C" author="Thorell, T." pageId="25" pageNumber="326" pagination="1 - 204" refId="ref23836" refString="Thorell, T. (1875) Descriptions of several European and North African spiders. Kongliga Svenska Vetenskaps-Akademiens Handlingar, 13 (5), 1 - 204." type="journal article" year="1875">Thorell, 1875</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="668A4D06FFF85D5899DBFCFC1857A79C" authority="(Thorell, 1875)" baseAuthorityName="Thorell" baseAuthorityYear="1875" box="[234,529,1958,1985]" class="Arachnida" family="Theridiidae" genus="Steatoda" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="nobilis">
<emphasis id="93FEEA97FFF85D5899DBFCFC1ABEA79C" box="[234,248,1959,1985]" italics="true" pageId="25" pageNumber="326">S</emphasis>
.
<emphasis id="93FEEA97FFF85D58983BFCFC1B10A79C" box="[266,342,1959,1985]" italics="true" pageId="25" pageNumber="326">nobilis</emphasis>
(
<bibRefCitation id="C51B4B74FFF85D589859FCFC184FA79C" author="Thorell, T." box="[360,521,1958,1985]" pageId="25" pageNumber="326" pagination="1 - 204" refId="ref23836" refString="Thorell, T. (1875) Descriptions of several European and North African spiders. Kongliga Svenska Vetenskaps-Akademiens Handlingar, 13 (5), 1 - 204." type="journal article" year="1875">Thorell, 1875</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="668A4D06FFF85D589B10FCFC19E8A79C" authority="(Walckenaer, 1805)" baseAuthorityName="Walckenaer" baseAuthorityYear="1805" box="[545,942,1958,1985]" class="Arachnida" family="Theridiidae" genus="Steatoda" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="paykulliana">
<emphasis id="93FEEA97FFF85D589B10FCFC1869A79C" box="[545,559,1959,1985]" italics="true" pageId="25" pageNumber="326">S</emphasis>
.
<emphasis id="93FEEA97FFF85D589B70FCFC1883A79C" box="[577,709,1959,1985]" italics="true" pageId="25" pageNumber="326">paykulliana</emphasis>
(Walckenaer, 1805)
</taxonomicName>
and
<taxonomicName id="668A4D06FFF85D589ADDFCFC1F3FA79C" authority="(Walckenaer, 1802)" baseAuthorityName="Walckenaer" baseAuthorityYear="1802" box="[1004,1401,1959,1985]" class="Arachnida" family="Theridiidae" genus="Steatoda" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="triangulosa">
<emphasis id="93FEEA97FFF85D589ADDFCFC19BCA79C" box="[1004,1018,1959,1985]" italics="true" pageId="25" pageNumber="326">S</emphasis>
.
<emphasis id="93FEEA97FFF85D589D3AFCFC1ECAA79C" box="[1035,1164,1959,1985]" italics="true" pageId="25" pageNumber="326">triangulosa</emphasis>
(Walckenaer, 1802)
</taxonomicName>
.
<taxonomicName id="668A4D06FFF85D589CB6FCFC1B69A7B8" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1872" class="Arachnida" family="Theridiidae" genus="Steatoda" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="erigoniformis">
<emphasis id="93FEEA97FFF85D589CB6FCFC1FD3A79C" box="[1415,1429,1959,1985]" italics="true" pageId="25" pageNumber="326">S</emphasis>
.
<emphasis id="93FEEA97FFF85D5899A6FC901B69A7B8" box="[151,303,1995,2021]" italics="true" pageId="25" pageNumber="326">erigoniformis</emphasis>
</taxonomicName>
occurs in many countries over the world, in the Middle East, East Mediterranean, to reach the far
</paragraph>
<paragraph id="A1353685FFFB5D5B99A6FBCC1BF9A088" blockId="26.[151,1437,151,393]" pageId="26" pageNumber="327">
East in
<collectingCountry id="D99D7615FFFB5D5B99DCFBCC1B76A0EC" box="[237,304,151,177]" name="China" pageId="26" pageNumber="327">China</collectingCountry>
and
<collectingCountry id="D99D7615FFFB5D5B9859FBCC1BEEA0EC" box="[360,424,151,177]" name="Japan" pageId="26" pageNumber="327">Japan</collectingCountry>
(
<bibRefCitation id="C51B4B74FFFB5D5B9886FBCC18B5A0EC" author="World Spider Catalog" box="[439,755,151,177]" pageId="26" pageNumber="327" refId="ref24111" refString="World Spider Catalog (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern, Bern. Available from: https: // wsc. nmbe. ch / (accessed 1 June 2024) https: // doi. org / 10.24436 / 2" type="book" year="2024">World Spider Catalog 2024</bibRefCitation>
), but in North Africa, the species is known only from
<collectingCountry id="D99D7615FFFB5D5B9C68FBCC1FDAA0EC" box="[1369,1436,151,177]" name="Egypt" pageId="26" pageNumber="327">Egypt</collectingCountry>
(
<bibRefCitation id="C51B4B74FFFB5D5B99AEFBE01BF4A088" author="Bosmans, R. &amp; Herve, C." box="[159,434,187,213]" pageId="26" pageNumber="327" pagination="58 - 64" refId="ref19706" refString="Bosmans, R. &amp; Herve, C. (2021) Less is more: eight new synonyms in Mediterranean spiders (Araneae), with a new Pelecopsis species from Tunisia (Linyphiidae). Arachnologische Mitteilungen, 61, 58 - 64. https: // doi. org / 10.30963 / aramit 6109" type="journal article" year="2021">Bosmans &amp; Hervé 2021</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFFB5D5B99F6FB85196EA11C" pageId="26" pageNumber="327" type="materials_examined">
<paragraph id="A1353685FFFB5D5B99F6FB851802A140" blockId="26.[151,1437,151,393]" pageId="26" pageNumber="327">
<emphasis id="93FEEA97FFFB5D5B99F6FB851BF4A0A5" bold="true" box="[199,434,222,248]" pageId="26" pageNumber="327">Material examined:</emphasis>
EL GUERRARA: Aghzou, pitfall traps,
<specimenCount id="B78CFD0CFFFB5D5B9A47FB8419E7A0A4" box="[886,929,223,249]" count="7" pageId="26" pageNumber="327" type="male">7 ♂</specimenCount>
,
<date id="D5341045FFFB5D5B9A9DFB841E65A0A4" box="[940,1059,223,249]" pageId="26" pageNumber="327" value="2020-02-22">22.II.2020</date>
;
<materialsCitation id="11E23CD8FFFB5D5B9D1FFB841E91A0A4" box="[1070,1239,223,249]" collectingDate="2020-03-03" isTailEnumeration="true" pageId="26" pageNumber="327" specimenCount="1" specimenCount-male="1">
<specimenCount id="B78CFD0CFFFB5D5B9D1FFB841E1FA0A4" box="[1070,1113,223,249]" count="1" pageId="26" pageNumber="327" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFFB5D5B9D55FB841E91A0A4" box="[1124,1239,223,249]" pageId="26" pageNumber="327" value="2020-03-03">
<collectingDate id="C570E9ADFFFB5D5B9D55FB841E91A0A4" box="[1124,1239,223,249]" pageId="26" pageNumber="327" value="2020-03-03">3.III.2020</collectingDate>
</date>
</materialsCitation>
;
<specimenCount id="B78CFD0CFFFB5D5B9DD0FB841F4AA0A4" box="[1249,1292,223,249]" count="2" pageId="26" pageNumber="327" type="male">2 ♂</specimenCount>
,
<date id="D5341045FFFB5D5B9C26FB841FDFA0A4" box="[1303,1433,223,249]" pageId="26" pageNumber="327" value="2020-03-17">17.III.2020</date>
. El Amied, pitfall traps,
<specimenCount id="B78CFD0CFFFB5D5B98AFFA581B8CA140" box="[414,458,259,285]" count="1" pageId="26" pageNumber="327" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFFB5D5B98E7FA581806A140" box="[470,576,259,285]" pageId="26" pageNumber="327" value="2020-02-07">7.II.2020</date>
.
</paragraph>
<paragraph id="A1353685FFFB5D5B99F6FA7D196EA11C" blockId="26.[151,1437,151,393]" box="[199,808,294,321]" pageId="26" pageNumber="327">
<emphasis id="93FEEA97FFFB5D5B99F6FA7D1853A11D" bold="true" box="[199,533,294,321]" pageId="26" pageNumber="327">
Previous records in
<collectingCountry id="D99D7615FFFB5D5B9882FA7D1856A11D" box="[435,528,294,320]" name="Algeria" pageId="26" pageNumber="327">Algeria</collectingCountry>
:
</emphasis>
New record for Algeria.
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFFB5D5B99F6FA111E4AA1D4" pageId="26" pageNumber="327" type="distribution">
<paragraph id="A1353685FFFB5D5B99F6FA111E4AA1D4" blockId="26.[151,1437,151,393]" pageId="26" pageNumber="327">
<emphasis id="93FEEA97FFFB5D5B99F6FA111B25A139" bold="true" box="[199,355,330,356]" pageId="26" pageNumber="327">Distribution:</emphasis>
East Mediterranean, Middle East, Caucasus,
<collectingCountry id="D99D7615FFFB5D5B9A45FA1019F6A138" box="[884,944,331,357]" name="India" pageId="26" pageNumber="327">India</collectingCountry>
,
<collectingCountry id="D99D7615FFFB5D5B9A8EFA101E43A138" box="[959,1029,331,357]" name="China" pageId="26" pageNumber="327">China</collectingCountry>
,
<collectingCountry id="D99D7615FFFB5D5B9D22FA101E1CA138" box="[1043,1114,331,357]" name="South Korea" pageId="26" pageNumber="327">Korea</collectingCountry>
,
<collectingCountry id="D99D7615FFFB5D5B9D59FA101EEDA138" box="[1128,1195,331,357]" name="Japan" pageId="26" pageNumber="327">Japan</collectingCountry>
. Introduced to
<collectingCountry id="D99D7615FFFB5D5B9C6CFA101FDEA138" box="[1373,1432,331,357]" name="United States of America" pageId="26" pageNumber="327">USA</collectingCountry>
, Caribbean,
<collectingCountry id="D99D7615FFFB5D5B9829FA341BC8A1D4" box="[280,398,367,393]" name="Venezuela" pageId="26" pageNumber="327">Venezuela</collectingCountry>
,
<collectingCountry id="D99D7615FFFB5D5B98A8FA341858A1D4" box="[409,542,367,393]" name="Cape Verde" pageId="26" pageNumber="327">Cabo Verde</collectingCountry>
,
<collectingCountry id="D99D7615FFFB5D5B9B1BFA3418FEA1D4" box="[554,696,367,393]" name="South Africa" pageId="26" pageNumber="327">South Africa</collectingCountry>
(
<bibRefCitation id="C51B4B74FFFB5D5B9BF7FA341E46A1D4" author="World Spider Catalog" box="[710,1024,367,393]" pageId="26" pageNumber="327" refId="ref24111" refString="World Spider Catalog (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern, Bern. Available from: https: // wsc. nmbe. ch / (accessed 1 June 2024) https: // doi. org / 10.24436 / 2" type="book" year="2024">World Spider Catalog 2024</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,124 @@
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<subSubSection id="E990650EFFFA5D5A99A6FC791B9CA761" box="[151,474,1826,1853]" pageId="27" pageNumber="328" type="nomenclature">
<paragraph id="A1353685FFFA5D5A99A6FC791B9CA761" blockId="27.[151,474,1826,1853]" box="[151,474,1826,1853]" pageId="27" pageNumber="328">
<heading id="FA7D81E9FFFA5D5A99A6FC791B9CA761" bold="true" box="[151,474,1826,1853]" fontSize="11" level="1" pageId="27" pageNumber="328" reason="1">
<taxonomicName id="668A4D06FFFA5D5A99A6FC791B9CA761" ID-CoL="95GCR" authority="(Simon, 1885)" baseAuthorityName="Simon" baseAuthorityYear="1885" box="[151,474,1826,1853]" class="Arachnida" family="Sparassidae" genus="Olios" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="pictus">
<emphasis id="93FEEA97FFFA5D5A99A6FC791B9CA761" bold="true" box="[151,474,1826,1853]" pageId="27" pageNumber="328">
<emphasis id="93FEEA97FFFA5D5A99A6FC791B60A760" bold="true" box="[151,294,1826,1853]" italics="true" pageId="27" pageNumber="328">Olios pictus</emphasis>
(
<bibRefCitation id="C51B4B74FFFA5D5A9804FC791B97A761" author="Simon, E." box="[309,465,1826,1852]" pageId="27" pageNumber="328" pagination="1 - 55" refId="ref23185" refString="Simon, E. (1885) Etudes sur les Arachnides recueillis en Tunisie en 1883 et 1884 par MM. A. Letourneux, M. Sedillot et Valery Mayet, membres de la mission de l'Exploration scientifique de la Tunisie. In: Exploration scientifique de la Tunisie, publiee sous les auspices du Ministere de l'instruction publique. Zoologie - Arachnides. Imprimerie nationale, Paris, pp. 1 - 55 pp." type="book chapter" year="1885">Simon, 1885</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFFA5D5A99A6FC331BB6A722" box="[151,496,1895,1920]" pageId="27" pageNumber="328" type="reference_group">
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<treatmentCitationGroup id="819A11ABFFFA5D5A99A6FC331BB6A722" box="[151,496,1895,1920]" pageId="27" pageNumber="328">
<taxonomicName id="668A4D06FFFA5D5A99A6FC331B7AA722" ID-CoL="47PGR" authorityName="Simon" authorityYear="1885" box="[151,316,1896,1919]" class="Arachnida" family="Sparassidae" genus="Nonianus" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="pictus">
<emphasis id="93FEEA97FFFA5D5A99A6FC331B7AA722" box="[151,316,1896,1919]" italics="true" pageId="27" pageNumber="328">Nonianus pictus</emphasis>
</taxonomicName>
;
<treatmentCitation id="202B1094FFFA5D5A9878FC331BAAA722" author="Levy, G." box="[329,492,1895,1920]" page="138" pageId="27" pageNumber="328" year="1989">
<bibRefCitation id="C51B4B74FFFA5D5A9878FC331BAAA722" author="Levy, G." box="[329,492,1895,1920]" pageId="27" pageNumber="328" pagination="127 - 176" refId="ref21326" refString="Levy, G. (1989) The family of huntsman spiders in Israel with annotations on species of the Middle East (Araneae: Sparassidae). Journal of Zoology, London, 217 (1), 127 - 176. https: // doi. org / 10.1111 / j. 1469 - 7998.1989. tb 02480. x" type="journal article" year="1989">Levy 1989: 138</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFFA5D5A99A6FCF61E9EA7B6" pageId="27" pageNumber="328" type="materials_examined">
<paragraph id="A1353685FFFA5D5A99A6FCF61E9EA7B6" blockId="27.[151,1240,1965,2028]" pageId="27" pageNumber="328">
<emphasis id="93FEEA97FFFA5D5A99A6FCF61BC5A79A" bold="true" box="[151,387,1965,1991]" pageId="27" pageNumber="328">Material examined:</emphasis>
<materialsCitation id="11E23CD8FFFA5D5A98BBFCF61E95A7B6" collectingDate="2015-02-07" country="Algeria" county="Algeria" location="Previous" municipality="Known" pageId="27" pageNumber="328" specimenCount="1" specimenCount-female="1">
MANSOURA: Mansoura Kedima: hand collecting,
<specimenCount id="B78CFD0CFFFA5D5A9AE0FCF619BDA795" box="[977,1019,1965,1992]" count="1" pageId="27" pageNumber="328" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFFA5D5A9D38FCF61E35A795" box="[1033,1139,1965,1992]" pageId="27" pageNumber="328" value="2015-02-07">
<collectingDate id="C570E9ADFFFA5D5A9D38FCF61E35A795" box="[1033,1139,1965,1992]" pageId="27" pageNumber="328" value="2015-02-07">7.II.2015</collectingDate>
</date>
.
<emphasis id="93FEEA97FFFA5D5A99F6FC8A1853A7B6" bold="true" box="[199,533,2001,2027]" pageId="27" pageNumber="328">
<location id="A455605EFFFA5D5A99F6FC8A1B69A7B6" LSID="urn:lsid:plazi:treatment:29238793FFFA5D5D99A6FC791E89A0EC:A455605EFFFA5D5A99F6FC8A1B69A7B6" box="[199,303,2001,2027]" country="Algeria" county="Algeria" municipality="Known" name="Previous" pageId="27" pageNumber="328">Previous</location>
records in
<collectingCountry id="D99D7615FFFA5D5A9882FC8A1856A7B6" box="[435,528,2001,2027]" name="Algeria" pageId="27" pageNumber="328">Algeria</collectingCountry>
:
</emphasis>
<collectingMunicipality id="4151ACFFFFFA5D5A9B2DFC891828A7B1" box="[540,622,2002,2028]" pageId="27" pageNumber="328">Known</collectingMunicipality>
from
<collectingCounty id="48544E09FFFA5D5A9B83FC8A1940A7B6" box="[690,774,2001,2027]" pageId="27" pageNumber="328">Algeria</collectingCounty>
but without precise locality (
<bibRefCitation id="C51B4B74FFFA5D5A9D7CFC8A1E8DA7B6" author="Levy, G." box="[1101,1227,2001,2027]" pageId="27" pageNumber="328" pagination="127 - 176" refId="ref21326" refString="Levy, G. (1989) The family of huntsman spiders in Israel with annotations on species of the Middle East (Araneae: Sparassidae). Journal of Zoology, London, 217 (1), 127 - 176. https: // doi. org / 10.1111 / j. 1469 - 7998.1989. tb 02480. x" type="journal article" year="1989">Levy 1989</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFFD5D5D99F6FBCD1E89A0EC" box="[199,1231,150,177]" pageId="28" pageNumber="329" type="distribution">
<paragraph id="A1353685FFFD5D5D99F6FBCD1E89A0EC" blockId="28.[199,1231,150,177]" box="[199,1231,150,177]" pageId="28" pageNumber="329">
<emphasis id="93FEEA97FFFD5D5D99F6FBCD1B25A0ED" bold="true" box="[199,355,150,176]" pageId="28" pageNumber="329">Distribution:</emphasis>
<collectingCountry id="D99D7615FFFD5D5D985BFBCC1B94A0EC" box="[362,466,151,177]" name="Morocco" pageId="28" pageNumber="329">Morocco</collectingCountry>
,
<collectingCountry id="D99D7615FFFD5D5D98EAFBCC1874A0EC" box="[475,562,151,177]" name="Algeria" pageId="28" pageNumber="329">Algeria</collectingCountry>
,
<collectingCountry id="D99D7615FFFD5D5D9B0CFBCC18D5A0EC" box="[573,659,151,177]" name="Tunisia" pageId="28" pageNumber="329">Tunisia</collectingCountry>
,
<collectingCountry id="D99D7615FFFD5D5D9BAFFBCC1899A0EC" box="[670,735,151,177]" name="Israel" pageId="28" pageNumber="329">Israel</collectingCountry>
,
<collectingCountry id="D99D7615FFFD5D5D9BDBFBCC193DA0EC" box="[746,891,151,177]" name="Saudi Arabia" pageId="28" pageNumber="329">Saudi Arabia</collectingCountry>
(
<bibRefCitation id="C51B4B74FFFD5D5D9ABBFBCC1E85A0EC" author="World Spider Catalog" box="[906,1219,151,177]" pageId="28" pageNumber="329" refId="ref24111" refString="World Spider Catalog (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern, Bern. Available from: https: // wsc. nmbe. ch / (accessed 1 June 2024) https: // doi. org / 10.24436 / 2" type="book" year="2024">World Spider Catalog 2024</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,195 @@
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<mods:title id="C0EDAA7572D0CD187433CC67DF810D67">Spiders of arid lands: The Ghardaïa region (Northern Sahara) with seven new records for Algeria</mods:title>
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<mods:namePart id="4365B8E36430B99AD77511C91031E62F">Alioua, Youcef</mods:namePart>
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<mods:namePart id="676B9D1254D3104BF97A12536244C1A1">Bosmans, Robert</mods:namePart>
<mods:affiliation id="91709E40026BAFD5AFF56C628F6BA1AF">Terrestrial Ecology Unit, Ledeganckstraat 35, B- 9000 Gent, Belgium</mods:affiliation>
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<subSubSection id="E990650EFFFC5D5C99A6FAD11851A1F9" box="[151,535,394,421]" pageId="29" pageNumber="330" type="nomenclature">
<paragraph id="A1353685FFFC5D5C99A6FAD11851A1F9" blockId="29.[151,535,394,421]" box="[151,535,394,421]" pageId="29" pageNumber="330">
<heading id="FA7D81E9FFFC5D5C99A6FAD11851A1F9" bold="true" box="[151,535,394,421]" fontSize="11" level="1" pageId="29" pageNumber="330" reason="1">
<taxonomicName id="668A4D06FFFC5D5C99A6FAD11851A1F9" ID-CoL="5D6CR" authority="Simon, 1914" authorityName="Simon" authorityYear="1914" box="[151,535,394,421]" class="Arachnida" family="Zodariidae" genus="Zodarion" isUncertain="true" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="330" phylum="Arthropoda" rank="species" species="pusio">
<emphasis id="93FEEA97FFFC5D5C99A6FAD11851A1F9" bold="true" box="[151,535,394,421]" pageId="29" pageNumber="330">
<emphasis id="93FEEA97FFFC5D5C99A6FAD11B41A1F8" bold="true" box="[151,263,394,421]" italics="true" pageId="29" pageNumber="330">Zodarion</emphasis>
cf.
<emphasis id="93FEEA97FFFC5D5C9805FAD01B33A1F8" bold="true" box="[308,373,395,421]" italics="true" pageId="29" pageNumber="330">pusio</emphasis>
<bibRefCitation id="C51B4B74FFFC5D5C984CFAD11851A1F9" author="Simon, E." box="[381,535,394,420]" pageId="29" pageNumber="330" refId="ref23509" refString="Simon, E. (1914) Les arachnides de France. Synopsis generale et catalogue des especes francaises de l'ordre des Araneae. Tome VI. 1 re partie. Roret, Paris, 308 pp." type="book" year="1914">Simon, 1914</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFFC5D5C99A6FA941E5DA278" pageId="29" pageNumber="330" type="reference_group">
<paragraph id="A1353685FFFC5D5C99A6FA94184BA1BA" blockId="29.[151,1051,463,549]" box="[151,525,463,487]" pageId="29" pageNumber="330">
<treatmentCitationGroup id="819A11ABFFFC5D5C99A6FA94184BA1BA" box="[151,525,463,487]" pageId="29" pageNumber="330">
<taxonomicName id="668A4D06FFFC5D5C99A6FA941B15A1BA" ID-CoL="5D6CQ" authorityName="Denis" authorityYear="1937" box="[151,339,463,487]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="330" phylum="Arthropoda" rank="species" species="punicum">
<emphasis id="93FEEA97FFFC5D5C99A6FA941B15A1BA" box="[151,339,463,487]" italics="true" pageId="29" pageNumber="330">Zodarion punicum</emphasis>
</taxonomicName>
;
<treatmentCitation id="202B1094FFFC5D5C9851FA94184FA1BA" author="Denis, J." box="[352,521,463,487]" page="28" pageId="29" pageNumber="330" year="1937">
<bibRefCitation id="C51B4B74FFFC5D5C9851FA94184FA1BA" author="Denis, J." box="[352,521,463,487]" pageId="29" pageNumber="330" pagination="1 - 50" refId="ref20611" refString="Denis, J. (1937 a) Contribution a l'etude des araignees du genre Zodarion Walckenaer. Festschrift Embrik Strand, 3, 1 - 50." type="journal article" year="1937">Denis 1937a: 28</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A1353685FFFC5D5C99A6FAB51857A25B" blockId="29.[151,1051,463,549]" box="[151,529,494,518]" pageId="29" pageNumber="330">
<treatmentCitationGroup id="819A11ABFFFC5D5C99A6FAB51857A25B" box="[151,529,494,518]" pageId="29" pageNumber="330">
<taxonomicName id="668A4D06FFFC5D5C99A6FAB51B75A258" ID-CoL="5D6CR" authorityName="Simon" authorityYear="1914" box="[151,307,494,518]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="330" phylum="Arthropoda" rank="species" species="pusio">
<emphasis id="93FEEA97FFFC5D5C99A6FAB51B75A258" box="[151,307,494,518]" italics="true" pageId="29" pageNumber="330">Zodarion pusio</emphasis>
</taxonomicName>
;
<treatmentCitation id="202B1094FFFC5D5C9871FAB5184AA25B" author="Bosmans, R." box="[320,524,494,518]" page="274" pageId="29" pageNumber="330" year="1997">
<bibRefCitation id="C51B4B74FFFC5D5C9871FAB5184AA25B" author="Bosmans, R." box="[320,524,494,518]" pageId="29" pageNumber="330" pagination="265 - 294" refId="ref19366" refString="Bosmans, R. (1997) Revision of the genus Zodarion Walckenaer, 1833, part II. Western and Central Europe, including Italy (Araneae: Zodariidae). Bulletin of the British Arachnological Society, 10 (8), 265 - 294." type="journal article" year="1997">Bosmans 1997: 274</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A1353685FFFC5D5C99A6F9561E5DA278" blockId="29.[151,1051,463,549]" box="[151,1051,524,549]" pageId="29" pageNumber="330">
For a complete list of taxonomic references, please see the
<bibRefCitation id="C51B4B74FFFC5D5C9BDCF9571E50A278" author="World Spider Catalog" box="[749,1046,524,549]" pageId="29" pageNumber="330" refId="ref24111" refString="World Spider Catalog (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern, Bern. Available from: https: // wsc. nmbe. ch / (accessed 1 June 2024) https: // doi. org / 10.24436 / 2" type="book" year="2024">World Spider Catalog (2024)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFFC5D5C99A6F9081BCAA2CC" pageId="29" pageNumber="330" type="description">
<paragraph id="A1353685FFFC5D5C99A6F9081BCAA2CC" blockId="29.[151,1437,595,873]" pageId="29" pageNumber="330">
Although
<taxonomicName id="668A4D06FFFC5D5C9836F9081B26A231" authorityName="Simon" authorityYear="1914" box="[263,352,595,621]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="330" phylum="Arthropoda" rank="species" species="pusio">
<emphasis id="93FEEA97FFFC5D5C9836F9081B51A231" box="[263,279,595,620]" italics="true" pageId="29" pageNumber="330">Z</emphasis>
.
<emphasis id="93FEEA97FFFC5D5C9812F9081B26A231" box="[291,352,595,620]" italics="true" pageId="29" pageNumber="330">pusio</emphasis>
</taxonomicName>
is known to exist in several countries in Europe, the species is reported only from
<collectingCountry id="D99D7615FFFC5D5C9DD6F9081F7CA230" box="[1255,1338,595,621]" name="Tunisia" pageId="29" pageNumber="330">Tunisia</collectingCountry>
in North Africa (
<bibRefCitation id="C51B4B74FFFC5D5C99DCF92C1BC6A2CC" author="Denis, J." box="[237,384,631,657]" pageId="29" pageNumber="330" pagination="1 - 50" refId="ref20611" refString="Denis, J. (1937 a) Contribution a l'etude des araignees du genre Zodarion Walckenaer. Festschrift Embrik Strand, 3, 1 - 50." type="journal article" year="1937">Denis 1937a</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFFC5D5C99F6F9C11FDBA284" pageId="29" pageNumber="330" type="materials_examined">
<paragraph id="A1353685FFFC5D5C99F6F9C11FDBA284" blockId="29.[151,1437,595,873]" pageId="29" pageNumber="330">
<emphasis id="93FEEA97FFFC5D5C99F6F9C11BF4A2E9" bold="true" box="[199,434,666,692]" pageId="29" pageNumber="330">Material examined:</emphasis>
BERRIANE: Balouh, hand collecting;
<specimenCount id="B78CFD0CFFFC5D5C9A57F9C019D7A2E8" box="[870,913,667,693]" count="1" pageId="29" pageNumber="330" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFFC5D5C9AADF9C01E5DA2E8" box="[924,1051,667,693]" pageId="29" pageNumber="330" value="2021-04-10">10.IV.2021</date>
;
<materialsCitation id="11E23CD8FFFC5D5C9D14F9C01E9CA2E8" box="[1061,1242,667,693]" collectingDate="2021-04-24" isTailEnumeration="true" pageId="29" pageNumber="330" specimenCount="1" specimenCount-female="1">
<specimenCount id="B78CFD0CFFFC5D5C9D14F9C01E08A2E8" box="[1061,1102,667,693]" count="1" pageId="29" pageNumber="330" type="female">1 ♀</specimenCount>
,
<date id="D5341045FFFC5D5C9D6BF9C01E9CA2E8" box="[1114,1242,667,693]" pageId="29" pageNumber="330" value="2021-04-24">
<collectingDate id="C570E9ADFFFC5D5C9D6BF9C01E9CA2E8" box="[1114,1242,667,693]" pageId="29" pageNumber="330" value="2021-04-24">24.IV.2021</collectingDate>
</date>
</materialsCitation>
;
<specimenCount id="B78CFD0CFFFC5D5C9DD5F9C01F4BA2E8" box="[1252,1293,667,693]" count="2" pageId="29" pageNumber="330" type="female">2 ♀</specimenCount>
,
<date id="D5341045FFFC5D5C9C28F9C01FDFA2E8" box="[1305,1433,667,693]" pageId="29" pageNumber="330" value="2021-04-30">30.IV.2021</date>
. Sidi Mbarek, hand collecting,
<specimenCount id="B78CFD0CFFFC5D5C98DBF9E41850A284" box="[490,534,703,729]" count="1" pageId="29" pageNumber="330" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFFC5D5C9B13F9E418DEA284" box="[546,664,703,729]" pageId="29" pageNumber="330" value="2021-02-11">11.II.2021</date>
. EL ATTEUF: Palm grove of El Atteuf, pitfall traps;
<specimenCount id="B78CFD0CFFFC5D5C9DD8F9E41F53A284" box="[1257,1301,703,729]" count="1" pageId="29" pageNumber="330" type="male">1 ♂</specimenCount>
,
<date id="D5341045FFFC5D5C9C10F9E51FDEA284" box="[1313,1432,702,729]" pageId="29" pageNumber="330" value="2015-02-16">16.II.2015</date>
.
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFFC5D5C99F6F9B91FDDA2A0" box="[199,1435,738,765]" pageId="29" pageNumber="330" type="distribution">
<paragraph id="A1353685FFFC5D5C99F6F9B91FDDA2A0" blockId="29.[151,1437,595,873]" box="[199,1435,738,765]" pageId="29" pageNumber="330">
<emphasis id="93FEEA97FFFC5D5C99F6F9B91B25A2A1" bold="true" box="[199,355,738,764]" pageId="29" pageNumber="330">Distribution:</emphasis>
<collectingCountry id="D99D7615FFFC5D5C985BF9B81BFFA2A0" box="[362,441,739,765]" name="France" pageId="29" pageNumber="330">France</collectingCountry>
,
<collectingCountry id="D99D7615FFFC5D5C98F5F9B81BBEA2A0" box="[452,504,739,765]" name="Italy" pageId="29" pageNumber="330">Italy</collectingCountry>
,
<collectingCountry id="D99D7615FFFC5D5C9B32F9B8182EA2A0" box="[515,616,739,765]" name="Slovenia" pageId="29" pageNumber="330">Slovenia</collectingCountry>
,
<collectingCountry id="D99D7615FFFC5D5C9B42F9B8188EA2A0" box="[627,712,739,765]" name="Croatia" pageId="29" pageNumber="330">Croatia</collectingCountry>
,
<collectingCountry id="D99D7615FFFC5D5C9BE5F9B819ACA2A0" box="[724,1002,739,765]" name="Bosnia and Herzegovina" pageId="29" pageNumber="330">Bosnia and Herzegovina</collectingCountry>
,
<collectingCountry id="D99D7615FFFC5D5C9AC4F9B81E0EA2A0" box="[1013,1096,739,765]" name="Tunisia" pageId="29" pageNumber="330">Tunisia</collectingCountry>
(
<bibRefCitation id="C51B4B74FFFC5D5C9D67F9B81FC9A2A0" author="World Spider Catalog" box="[1110,1423,739,765]" pageId="29" pageNumber="330" refId="ref24111" refString="World Spider Catalog (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern, Bern. Available from: https: // wsc. nmbe. ch / (accessed 1 June 2024) https: // doi. org / 10.24436 / 2" type="book" year="2024">World Spider Catalog 2024</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E990650EFFFC5D5C99F6F85D1F6DA438" pageId="29" pageNumber="330" type="discussion">
<paragraph id="A1353685FFFC5D5C99F6F85D1BFFA334" blockId="29.[151,1437,595,873]" pageId="29" pageNumber="330">
<emphasis id="93FEEA97FFFC5D5C99F6F85D1B01A37D" bold="true" box="[199,327,774,800]" pageId="29" pageNumber="330">Comment:</emphasis>
We have noticed some differences at the end of the retrolateral apophysis of the male pedipalp in comparison to the description of
<bibRefCitation id="C51B4B74FFFC5D5C9B3DF870188EA318" author="Bosmans, R." box="[524,712,811,837]" pageId="29" pageNumber="330" pagination="265 - 294" refId="ref19366" refString="Bosmans, R. (1997) Revision of the genus Zodarion Walckenaer, 1833, part II. Western and Central Europe, including Italy (Araneae: Zodariidae). Bulletin of the British Arachnological Society, 10 (8), 265 - 294." type="journal article" year="1997">Bosmans (1997)</bibRefCitation>
. This result is the subject of a work on the revision of the genus
<taxonomicName id="668A4D06FFFC5D5C99A6F8141AB9A334" box="[151,255,847,873]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="330" phylum="Arthropoda" rank="genus">
<emphasis id="93FEEA97FFFC5D5C99A6F8141AB9A334" box="[151,255,847,873]" italics="true" pageId="29" pageNumber="330">Zodarion</emphasis>
</taxonomicName>
in the Maghreb.
</paragraph>
<paragraph id="A1353685FFFC5D5C99A6F8CC1927A388" blockId="29.[151,1437,918,1125]" pageId="29" pageNumber="330">
Among this study, a total number of 2037 adult spiders grouped into 1065 males and
<specimenCount id="B78CFD0CFFFC5D5C9D55F8CC1EA8A3EC" box="[1124,1262,919,945]" count="972" pageId="29" pageNumber="330" type="female">972 females</specimenCount>
was identified, they were classified into 21 families, 55 genera, and 70 species.
</paragraph>
<paragraph id="A1353685FFFC5D5C99F6F8841F6DA438" blockId="29.[151,1437,918,1125]" pageId="29" pageNumber="330">
The family of
<taxonomicName id="668A4D06FFFC5D5C9841F8841844A3A4" authorityName="Banks" authorityYear="1892" box="[368,514,991,1017]" class="Arachnida" family="Gnaphosidae" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="330" phylum="Arthropoda" rank="family">Gnaphosidae</taxonomicName>
was the richest in species with 20, followed by
<taxonomicName id="668A4D06FFFC5D5C9D07F8841EE2A3A4" authorityName="Blackwall" authorityYear="1841" box="[1078,1188,991,1017]" class="Arachnida" family="Salticidae" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="330" phylum="Arthropoda" rank="family">Salticidae</taxonomicName>
with 12 species. The families of
<taxonomicName id="668A4D06FFFC5D5C9827FF581BCEA440" authorityName="Sundevall" authorityYear="1833" box="[278,392,1027,1053]" class="Arachnida" family="Lycosidae" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="330" phylum="Arthropoda" rank="family">Lycosidae</taxonomicName>
and
<taxonomicName id="668A4D06FFFC5D5C988CFF581878A440" authorityName="Sundevall" authorityYear="1833" box="[445,574,1027,1053]" class="Arachnida" family="Theridiidae" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="330" phylum="Arthropoda" rank="family">Theridiidae</taxonomicName>
contained seven species for each of them. The Lyniphiidae is the only family that was represented by four species.
<taxonomicName id="668A4D06FFFC5D5C9B7EFF7C18B2A41C" authorityName="Thorell" authorityYear="1869" box="[591,756,1063,1089]" class="Arachnida" family="Philodromidae" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="330" phylum="Arthropoda" rank="family">Philodromidae</taxonomicName>
with three species. There were two species in each of the
<taxonomicName id="668A4D06FFFC5D5C99A6FF101B4CA438" authorityName="Clerck" authorityYear="1757" box="[151,266,1099,1125]" class="Arachnida" family="Araneidae" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="330" phylum="Arthropoda" rank="family">Araneidae</taxonomicName>
and
<taxonomicName id="668A4D06FFFC5D5C9871FF101B82A438" authorityName="Sundevall" authorityYear="1833" box="[320,452,1099,1125]" class="Arachnida" family="Thomisidae" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="330" phylum="Arthropoda" rank="family">Thomisidae</taxonomicName>
families, whereas there was only one species per family in the other families.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,202 @@
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<taxonomicName id="2AA34D28CA55FFA3FF7200D7FDC5FF1B" ID-CoL="6BMMM" authority="Bernhauer, 1915" authorityName="Bernhauer (FMNH" authorityYear="1915" box="[151,595,151,178]" class="Insecta" family="Staphylinidae" genus="Ctenopeuca" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="261" phylum="Arthropoda" rank="species" species="heynei">
<emphasis id="DFD7EAB9CA55FFA3FF7200D7FDC5FF1B" bold="true" box="[151,595,151,178]" pageId="6" pageNumber="261">
<emphasis id="DFD7EAB9CA55FFA3FF7200D7FEE8FF18" bold="true" box="[151,382,151,178]" italics="true" pageId="6" pageNumber="261">Ctenopeuca heynei</emphasis>
<bibRefCitation id="89324B5ACA55FFA3FE6000D7FDC5FF1B" author="Bernhauer, M." box="[389,595,151,177]" pageId="6" pageNumber="261" pagination="291 - 301" refId="ref5053" refString="Bernhauer, M. (1915) Zur Staphyliniden-Fauna von Sudamerika. Stettiner Entomologische Zeitung, 76, 291 - 301. https: // doi. org / 10.1002 / mmnd. 191519150207" type="journal article" year="1915">Bernhauer, 1915</bibRefCitation>
</emphasis>
</taxonomicName>
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<paragraph id="ED1C36ABCA55FFA3FF7200FBFEF0FF7F" blockId="6.[151,595,151,213]" box="[151,358,187,213]" pageId="6" pageNumber="261">
(
<figureCitation id="75982A2ECA55FFA3FF7A00FBFE9DFF7F" box="[159,267,187,213]" captionStart="FIGURES 16" captionStartId="3.[152,270,1658,1682]" captionTargetBox="[151,1436,181,1635]" captionTargetId="figure-22@3.[151,1436,181,1635]" captionTargetPageId="3" captionText="FIGURES 16. Habitus of Ctenopeuca. 13) Lectotype of C. heynei Bernhauer (FMNH): 1) lateral view (ST3 indicates spine of tergum III, SS3 indicates spine of sternum III, ST4 indicates spine of tergum IV, SS4 indicates spine of sternum IV, ST7 indicates spine of tergum VII), 2) dorsal view, 3) ventral view. 13) Lectotype of C. romani Bernhauer (FMNH): 4) lateral view, 5) dorsal view, 6) ventral view. Scale bars 3.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13618396" httpUri="https://zenodo.org/record/13618396/files/figure.png" pageId="6" pageNumber="261">Figs. 13</figureCitation>
,
<figureCitation id="75982A2ECA55FFA3FEF200FBFEC8FF7F" box="[279,350,187,213]" captionStart="FIGURES 2127" captionStartId="7.[152,270,1943,1967]" captionTargetBox="[238,1348,187,1913]" captionTargetId="figure-22@7.[232,1354,181,1919]" captionTargetPageId="7" captionText="FIGURES 2127. Ctenopeuca heynei: 21) Tergum VIII, male, dorsal view; 22) Sternum VIII, male, dorsal view; 23) Median lobe, ventral view; 24) Median lobe, lateral view; 25) Paramere; 26) Tergum VIII, female, dorsal view; 27) Sternum VIII, female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13618408" httpUri="https://zenodo.org/record/13618408/files/figure.png" pageId="6" pageNumber="261">2127</figureCitation>
)
</paragraph>
<paragraph id="ED1C36ABCA55FFA3FF720144FDEBFDB3" blockId="6.[151,1437,259,1222]" pageId="6" pageNumber="261">
Type material.
<materialsCitation id="5DCB3CF6CA55FFA3FEA40144FDEBFDB3" collectionCode="FMNH, T, FIELD, MUSEUM" collectorName="M. Bernhauer &amp; A. Heyne &amp; S. Ware" country="Peru" elevation="1000" location="Chanchamayo" pageId="6" pageNumber="261" specimenCount="2" typeStatus="lectotype">
<typeStatus id="32188809CA55FFA3FEA40144FE27FEB4" box="[321,433,260,286]" pageId="6" pageNumber="261" type="lectotype">Lectotype</typeStatus>
(
<collectionCode id="8BB2AE6ECA55FFA3FE240143FD8EFEB7" box="[449,536,259,285]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="6" pageNumber="261" type="Museum">FMNH</collectionCode>
), male, here designated (
<figureCitation id="75982A2ECA55FFA3FCD40143FC0BFEB7" box="[817,925,259,285]" captionStart="FIGURES 16" captionStartId="3.[152,270,1658,1682]" captionTargetBox="[151,1436,181,1635]" captionTargetId="figure-22@3.[151,1436,181,1635]" captionTargetPageId="3" captionText="FIGURES 16. Habitus of Ctenopeuca. 13) Lectotype of C. heynei Bernhauer (FMNH): 1) lateral view (ST3 indicates spine of tergum III, SS3 indicates spine of sternum III, ST4 indicates spine of tergum IV, SS4 indicates spine of sternum IV, ST7 indicates spine of tergum VII), 2) dorsal view, 3) ventral view. 13) Lectotype of C. romani Bernhauer (FMNH): 4) lateral view, 5) dorsal view, 6) ventral view. Scale bars 3.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13618396" httpUri="https://zenodo.org/record/13618396/files/figure.png" pageId="6" pageNumber="261">Figs. 13</figureCitation>
). Labels: 1) “Chicago NHMus/
<collectorName id="4056537DCA55FFA3FAE40143FA02FEB7" box="[1281,1428,259,285]" pageId="6" pageNumber="261">M.Bernhauer</collectorName>
/ Collection” [white label, printed in black]; 2) “
<quantity id="2A5B9B4ECA55FFA3FD7D0167FD7CFEEB" box="[664,746,295,321]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="261" unit="m" value="1000.0">
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” [white label, manuscript]; 3) “
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/
<collectingCountry id="95B4763BCA55FFA3FB0A0167FAB3FEEB" box="[1263,1317,295,321]" name="Peru" pageId="6" pageNumber="261">Peru</collectingCountry>
.
<collectorName id="4056537DCA55FFA3FACE0167FA04FEEB" box="[1323,1426,295,321]" pageId="6" pageNumber="261">A.Heyne</collectorName>
” [white label, printed in black]; 4) “
<taxonomicName id="2AA34D28CA55FFA3FDD8010CFD55FECC" box="[573,707,332,358]" class="Insecta" family="Staphylinidae" genus="Ctenopeuca" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="DFD7EAB9CA55FFA3FDD8010CFD55FECC" box="[573,707,332,358]" italics="true" pageId="6" pageNumber="261">Ctenopeuca</emphasis>
</taxonomicName>
/
<taxonomicName id="2AA34D28CA55FFA3FD2E010BFCE2FECF" authority="Bernh." authorityName="Bernhauer (FMNH" authorityYear="1915" box="[715,884,331,357]" class="Insecta" family="Staphylinidae" genus="Ctenopeuca" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="261" phylum="Arthropoda" rank="species" species="heynei">Heynei Bernh.</taxonomicName>
/typus” [white old label, manuscript]; 5) [male symbol, white label, manuscript]; 6) “
<collectionCode id="8BB2AE6ECA55FFA3FDA9012FFD34FE23" box="[588,674,367,393]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="6" pageNumber="261" type="Museum">FMNH</collectionCode>
” [white label, printed in black]; 7) “
<typeStatus id="32188809CA55FFA3FBA9012FFB2DFE23" box="[1100,1211,367,393]" pageId="6" pageNumber="261" type="lectotype">Lectotype</typeStatus>
/
<taxonomicName id="2AA34D28CA55FFA3FB210130FADCFE20" box="[1220,1354,368,394]" class="Insecta" family="Staphylinidae" genus="Ctenopeuca" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="DFD7EAB9CA55FFA3FB210130FADCFE20" box="[1220,1354,368,394]" italics="true" pageId="6" pageNumber="261">Ctenopeuca</emphasis>
</taxonomicName>
/heynei [male symbol + Greek letter alpha]/des. K.
<collectionCode id="8BB2AE6ECA55FFA3FD9101D3FD13FE07" box="[628,645,403,429]" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="261">T</collectionCode>
.Eldredge 2012” [red label, the first and last line (except 2012) printed in black, the rest manuscript]; 8) “QR Code/FMNHINS/3982449/
<collectionCode id="8BB2AE6ECA55FFA3FC9801F7FBC6FE7B" box="[893,1104,439,465]" pageId="6" pageNumber="261">FIELD MUSEUM</collectionCode>
/Pinned” [white label, printed in black]; 9) “PHOTOGRAPHED/
<collectorName id="4056537DCA55FFA3FDC3019BFDEFFE5C" box="[550,633,475,502]" pageId="6" pageNumber="261">S. Ware</collectorName>
2021” [white label, prinited in black]. Note: Bernahuer (1915) did not specify how many specimens were studied.
</materialsCitation>
</paragraph>
<paragraph id="ED1C36ABCA55FFA3FF220263FDBAFD97" blockId="6.[151,1437,259,1222]" box="[199,556,547,573]" pageId="6" pageNumber="261">
<materialsCitation id="5DCB3CF6CA55FFA3FF220263FDBEFD97" box="[199,552,547,573]" collectionCode="FMNH" pageId="6" pageNumber="261" specimenCount="1" typeStatus="paralectotype">
<typeStatus id="32188809CA55FFA3FF220263FEC9FD97" box="[199,351,547,573]" pageId="6" pageNumber="261" type="paralectotype">Paralectotype</typeStatus>
(
<collectionCode id="8BB2AE6ECA55FFA3FE8B0263FE53FD97" box="[366,453,547,573]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="6" pageNumber="261" type="Museum">FMNH</collectionCode>
), female
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A5B96520CA55FFA3FF220207FB73FD67" pageId="6" pageNumber="261" type="diagnosis">
<paragraph id="ED1C36ABCA55FFA3FF220207FB73FD67" blockId="6.[151,1437,259,1222]" pageId="6" pageNumber="261">
<emphasis id="DFD7EAB9CA55FFA3FF220207FED4FDCB" bold="true" box="[199,322,583,609]" pageId="6" pageNumber="261">Diagnosis.</emphasis>
<taxonomicName id="2AA34D28CA55FFA3FEAC0208FDB6FDCB" authorityName="Bernhauer (FMNH" authorityYear="1915" box="[329,544,583,610]" class="Insecta" family="Staphylinidae" genus="Ctenopeuca" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="261" phylum="Arthropoda" rank="species" species="heynei">
<emphasis id="DFD7EAB9CA55FFA3FEAC0208FDB6FDCB" box="[329,544,583,610]" italics="true" pageId="6" pageNumber="261">Ctenopeuca heynei</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2AA34D28CA55FFA3FD5D0208FCBFFDCB" authorityName="Bernhauer (FMNH" authorityYear="1928" box="[696,809,584,610]" class="Insecta" family="Staphylinidae" genus="Ctenopeuca" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="261" phylum="Arthropoda" rank="species" species="romani">
<emphasis id="DFD7EAB9CA55FFA3FD5D0208FCBFFDCB" box="[696,809,584,610]" italics="true" pageId="6" pageNumber="261">C. romani</emphasis>
</taxonomicName>
by having head and pronotum with the same color and abdominal segments IIIIV slightly lighter than VVII (
<figureCitation id="75982A2ECA55FFA3FCFF022BFC11FD2F" box="[794,903,619,645]" captionStart="FIGURES 16" captionStartId="3.[152,270,1658,1682]" captionTargetBox="[151,1436,181,1635]" captionTargetId="figure-22@3.[151,1436,181,1635]" captionTargetPageId="3" captionText="FIGURES 16. Habitus of Ctenopeuca. 13) Lectotype of C. heynei Bernhauer (FMNH): 1) lateral view (ST3 indicates spine of tergum III, SS3 indicates spine of sternum III, ST4 indicates spine of tergum IV, SS4 indicates spine of sternum IV, ST7 indicates spine of tergum VII), 2) dorsal view, 3) ventral view. 13) Lectotype of C. romani Bernhauer (FMNH): 4) lateral view, 5) dorsal view, 6) ventral view. Scale bars 3.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13618396" httpUri="https://zenodo.org/record/13618396/files/figure.png" pageId="6" pageNumber="261">Figs. 13</figureCitation>
); antennomere 4 quadrate; posterior margin of abdominal tergum VIII of male with a prominent tooth on each side of serrate region (
<figureCitation id="75982A2ECA55FFA3FBB902CFFB39FD03" box="[1116,1199,655,681]" captionStart="FIGURES 2127" captionStartId="7.[152,270,1943,1967]" captionTargetBox="[238,1348,187,1913]" captionTargetId="figure-22@7.[232,1354,181,1919]" captionTargetPageId="7" captionText="FIGURES 2127. Ctenopeuca heynei: 21) Tergum VIII, male, dorsal view; 22) Sternum VIII, male, dorsal view; 23) Median lobe, ventral view; 24) Median lobe, lateral view; 25) Paramere; 26) Tergum VIII, female, dorsal view; 27) Sternum VIII, female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13618408" httpUri="https://zenodo.org/record/13618408/files/figure.png" pageId="6" pageNumber="261">Fig. 21</figureCitation>
); posterior margin of abdominal tergum VIII of female with a slightly tooth on each side of emarginate region (
<figureCitation id="75982A2ECA55FFA3FB6102F3FB4EFD67" box="[1156,1240,691,717]" captionStart="FIGURES 2127" captionStartId="7.[152,270,1943,1967]" captionTargetBox="[238,1348,187,1913]" captionTargetId="figure-22@7.[232,1354,181,1919]" captionTargetPageId="7" captionText="FIGURES 2127. Ctenopeuca heynei: 21) Tergum VIII, male, dorsal view; 22) Sternum VIII, male, dorsal view; 23) Median lobe, ventral view; 24) Median lobe, lateral view; 25) Paramere; 26) Tergum VIII, female, dorsal view; 27) Sternum VIII, female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13618408" httpUri="https://zenodo.org/record/13618408/files/figure.png" pageId="6" pageNumber="261">Fig. 26</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A5B96520CA55FFA3FF220297FE9DFB9F" pageId="6" pageNumber="261" type="description">
<paragraph id="ED1C36ABCA55FFA3FF220297FE9DFB9F" blockId="6.[151,1437,259,1222]" pageId="6" pageNumber="261">
<emphasis id="DFD7EAB9CA55FFA3FF220297FEE3FD5B" bold="true" box="[199,373,727,753]" pageId="6" pageNumber="261">Redescription.</emphasis>
Male. Body length 6.0 mm, humeral width 1.0 mm. Head, antennomeres 411, pronotum, postero-lateral angle region of elytra and abdominal segments IVVII brown to dark brown, the rest of body yellowish to light brown (
<figureCitation id="75982A2ECA55FFA3FE5B035FFD93FC93" box="[446,517,799,825]" captionStart="FIGURES 16" captionStartId="3.[152,270,1658,1682]" captionTargetBox="[151,1436,181,1635]" captionTargetId="figure-22@3.[151,1436,181,1635]" captionTargetPageId="3" captionText="FIGURES 16. Habitus of Ctenopeuca. 13) Lectotype of C. heynei Bernhauer (FMNH): 1) lateral view (ST3 indicates spine of tergum III, SS3 indicates spine of sternum III, ST4 indicates spine of tergum IV, SS4 indicates spine of sternum IV, ST7 indicates spine of tergum VII), 2) dorsal view, 3) ventral view. 13) Lectotype of C. romani Bernhauer (FMNH): 4) lateral view, 5) dorsal view, 6) ventral view. Scale bars 3.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13618396" httpUri="https://zenodo.org/record/13618396/files/figure.png" pageId="6" pageNumber="261">Fig. 2</figureCitation>
). Antennomere 4 quadrate. Abdominal spine of tergum IV somewhat quadrate in lateral view (
<figureCitation id="75982A2ECA55FFA3FECB0303FEE4FCF7" box="[302,370,835,861]" captionStart="FIGURES 16" captionStartId="3.[152,270,1658,1682]" captionTargetBox="[151,1436,181,1635]" captionTargetId="figure-22@3.[151,1436,181,1635]" captionTargetPageId="3" captionText="FIGURES 16. Habitus of Ctenopeuca. 13) Lectotype of C. heynei Bernhauer (FMNH): 1) lateral view (ST3 indicates spine of tergum III, SS3 indicates spine of sternum III, ST4 indicates spine of tergum IV, SS4 indicates spine of sternum IV, ST7 indicates spine of tergum VII), 2) dorsal view, 3) ventral view. 13) Lectotype of C. romani Bernhauer (FMNH): 4) lateral view, 5) dorsal view, 6) ventral view. Scale bars 3.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13618396" httpUri="https://zenodo.org/record/13618396/files/figure.png" pageId="6" pageNumber="261">Fig. 1</figureCitation>
, ST4); spine of sternum III three times longer than the spine of sternum IV (
<figureCitation id="75982A2ECA55FFA3FB0C0303FAB8FCF7" box="[1257,1326,835,861]" captionStart="FIGURES 16" captionStartId="3.[152,270,1658,1682]" captionTargetBox="[151,1436,181,1635]" captionTargetId="figure-22@3.[151,1436,181,1635]" captionTargetPageId="3" captionText="FIGURES 16. Habitus of Ctenopeuca. 13) Lectotype of C. heynei Bernhauer (FMNH): 1) lateral view (ST3 indicates spine of tergum III, SS3 indicates spine of sternum III, ST4 indicates spine of tergum IV, SS4 indicates spine of sternum IV, ST7 indicates spine of tergum VII), 2) dorsal view, 3) ventral view. 13) Lectotype of C. romani Bernhauer (FMNH): 4) lateral view, 5) dorsal view, 6) ventral view. Scale bars 3.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13618396" httpUri="https://zenodo.org/record/13618396/files/figure.png" pageId="6" pageNumber="261">Fig. 1</figureCitation>
, SS3 and SS4); tergum VIII with posterior margin serrate on mesal three-fifth, a prominent tooth on each side of serrate region (
<figureCitation id="75982A2ECA55FFA3FF0803CBFED6FC0F" box="[237,320,907,933]" captionStart="FIGURES 2127" captionStartId="7.[152,270,1943,1967]" captionTargetBox="[238,1348,187,1913]" captionTargetId="figure-22@7.[232,1354,181,1919]" captionTargetPageId="7" captionText="FIGURES 2127. Ctenopeuca heynei: 21) Tergum VIII, male, dorsal view; 22) Sternum VIII, male, dorsal view; 23) Median lobe, ventral view; 24) Median lobe, lateral view; 25) Paramere; 26) Tergum VIII, female, dorsal view; 27) Sternum VIII, female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13618408" httpUri="https://zenodo.org/record/13618408/files/figure.png" pageId="6" pageNumber="261">Fig. 21</figureCitation>
); sternum VIII with posterior margin strongly projected medially (
<figureCitation id="75982A2ECA55FFA3FBC703CBFBE0FC0F" box="[1058,1142,907,933]" captionStart="FIGURES 2127" captionStartId="7.[152,270,1943,1967]" captionTargetBox="[238,1348,187,1913]" captionTargetId="figure-22@7.[232,1354,181,1919]" captionTargetPageId="7" captionText="FIGURES 2127. Ctenopeuca heynei: 21) Tergum VIII, male, dorsal view; 22) Sternum VIII, male, dorsal view; 23) Median lobe, ventral view; 24) Median lobe, lateral view; 25) Paramere; 26) Tergum VIII, female, dorsal view; 27) Sternum VIII, female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13618408" httpUri="https://zenodo.org/record/13618408/files/figure.png" pageId="6" pageNumber="261">Fig. 22</figureCitation>
); median lobe of aedeagus strongly curved paramerally in lateral view (
<figureCitation id="75982A2ECA55FFA3FD6103EFFD40FC63" box="[644,726,943,969]" captionStart="FIGURES 2127" captionStartId="7.[152,270,1943,1967]" captionTargetBox="[238,1348,187,1913]" captionTargetId="figure-22@7.[232,1354,181,1919]" captionTargetPageId="7" captionText="FIGURES 2127. Ctenopeuca heynei: 21) Tergum VIII, male, dorsal view; 22) Sternum VIII, male, dorsal view; 23) Median lobe, ventral view; 24) Median lobe, lateral view; 25) Paramere; 26) Tergum VIII, female, dorsal view; 27) Sternum VIII, female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13618408" httpUri="https://zenodo.org/record/13618408/files/figure.png" pageId="6" pageNumber="261">Fig. 24</figureCitation>
), apical half with ventral face evenly curved. Female. Similar to male, except tergum VIII with posterior margin emarginate on mesal three-fifth, a prominent tooth on each side of emarginate region (
<figureCitation id="75982A2ECA55FFA3FE9403B7FE50FBBB" box="[369,454,1015,1041]" captionStart="FIGURES 2127" captionStartId="7.[152,270,1943,1967]" captionTargetBox="[238,1348,187,1913]" captionTargetId="figure-22@7.[232,1354,181,1919]" captionTargetPageId="7" captionText="FIGURES 2127. Ctenopeuca heynei: 21) Tergum VIII, male, dorsal view; 22) Sternum VIII, male, dorsal view; 23) Median lobe, ventral view; 24) Median lobe, lateral view; 25) Paramere; 26) Tergum VIII, female, dorsal view; 27) Sternum VIII, female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13618408" httpUri="https://zenodo.org/record/13618408/files/figure.png" pageId="6" pageNumber="261">Fig. 26</figureCitation>
); sternum VIII with posterior margin slightly projected medially (
<figureCitation id="75982A2ECA55FFA3FB4D03B7FB6AFBBB" box="[1192,1276,1015,1041]" captionStart="FIGURES 2127" captionStartId="7.[152,270,1943,1967]" captionTargetBox="[238,1348,187,1913]" captionTargetId="figure-22@7.[232,1354,181,1919]" captionTargetPageId="7" captionText="FIGURES 2127. Ctenopeuca heynei: 21) Tergum VIII, male, dorsal view; 22) Sternum VIII, male, dorsal view; 23) Median lobe, ventral view; 24) Median lobe, lateral view; 25) Paramere; 26) Tergum VIII, female, dorsal view; 27) Sternum VIII, female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13618408" httpUri="https://zenodo.org/record/13618408/files/figure.png" pageId="6" pageNumber="261">Fig. 27</figureCitation>
); spermatheca not found.
</paragraph>
</subSubSection>
<subSubSection id="A5B96520CA55FFA3FF22047FFEBFFB6C" pageId="6" pageNumber="261" type="distribution">
<paragraph id="ED1C36ABCA55FFA3FF22047FFC9FFBF3" blockId="6.[151,1437,259,1222]" box="[199,777,1087,1114]" pageId="6" pageNumber="261">
<emphasis id="DFD7EAB9CA55FFA3FF22047FFEF7FBF3" bold="true" box="[199,353,1087,1113]" pageId="6" pageNumber="261">Distribution.</emphasis>
<collectingCountry id="95B4763BCA55FFA3FE820400FE09FBF0" box="[359,415,1088,1114]" name="Peru" pageId="6" pageNumber="261">Peru</collectingCountry>
:
<collectingRegion id="2F67F849CA55FFA3FE4F047FFE71FBF3" box="[426,487,1087,1113]" country="Peru" name="Junin" pageId="6" pageNumber="261">Junín</collectingRegion>
(Chanchamayo) (
<figureCitation id="75982A2ECA55FFA3FD4A047FFC97FBF3" box="[687,769,1087,1113]" captionStart="FIGURE 34" captionStartId="9.[152,255,1373,1397]" captionTargetBox="[151,1436,181,1350]" captionTargetId="figure-22@9.[151,1436,181,1350]" captionTargetPageId="9" captionText="FIGURE 34. Map of Ctenopeuca distribution. The green spot indicates C. heynei (not exact record), the pink spot indicates C. romani (not exact record), and the red spot indicates C. romani (exact record)." figureDoi="http://doi.org/10.5281/zenodo.13618416" httpUri="https://zenodo.org/record/13618416/files/figure.png" pageId="6" pageNumber="261">Fig. 34</figureCitation>
)
</paragraph>
<paragraph id="ED1C36ABCA55FFA3FF220423FEBFFB6C" blockId="6.[151,1437,259,1222]" pageId="6" pageNumber="261">
<emphasis id="DFD7EAB9CA55FFA3FF220423FE1EFBD7" bold="true" box="[199,392,1123,1149]" pageId="6" pageNumber="261">Biological notes.</emphasis>
<materialsCitation id="5DCB3CF6CA55FFA3FE680423FEB2FB6C" country="Peru" county="The" location="Only" municipality="Chanchamayo" pageId="6" pageNumber="261" specimenCount="1" stateProvince="Junin" typeStatus="lectotype">
<location id="E87C6070CA55FFA3FE680423FE53FBD7" LSID="urn:lsid:plazi:treatment:650A87BDCA55FFA3FF7200D7FEBFFB6C:E87C6070CA55FFA3FE680423FE53FBD7" box="[397,453,1123,1149]" country="Peru" county="The" municipality="Chanchamayo" name="Only" pageId="6" pageNumber="261" stateProvince="Junin">Only</location>
the type material is known and collected from
<collectingMunicipality id="0D78ACD1CA55FFA3FC210423FBF1FBD7" box="[964,1127,1123,1149]" pageId="6" pageNumber="261">Chanchamayo</collectingMunicipality>
, presumably in the province in northern
<collectingRegion id="2F67F849CA55FFA3FEFC04C7FE31FB0B" box="[281,423,1159,1185]" country="Peru" name="Junin" pageId="6" pageNumber="261">Junín region</collectingRegion>
,
<collectingCountry id="95B4763BCA55FFA3FE5704C8FE7FFB08" box="[434,489,1160,1186]" name="Peru" pageId="6" pageNumber="261">Peru</collectingCountry>
.
<collectingCounty id="047D4E27CA55FFA3FE1604C7FD89FB0B" box="[499,543,1159,1185]" pageId="6" pageNumber="261">The</collectingCounty>
<typeStatus id="32188809CA55FFA3FDC004C7FD1AFB0B" box="[549,652,1159,1185]" pageId="6" pageNumber="261" type="lectotype">lectotype</typeStatus>
has a label indicating that the specimen was collected at an altitude of 1000 meters
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,288 @@
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<mods:title id="32975251BB494F249EA0D2A1B6514314">Review of Ctenopeuca Bernhauer, a spiny, pipevine flower-associated rove beetle from South America (Coleoptera, Staphylinidae, Aleocharinae, Oxypodini)</mods:title>
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<subSubSection id="A5B96520CA55FFA3FF720557FDCAFA9B" box="[151,604,1303,1329]" pageId="6" pageNumber="261" type="nomenclature">
<paragraph id="ED1C36ABCA55FFA3FF720557FDCAFA9B" blockId="6.[151,604,1303,1365]" box="[151,604,1303,1329]" pageId="6" pageNumber="261">
<heading id="B65481C7CA55FFA3FF720557FDCAFA9B" bold="true" box="[151,604,1303,1329]" fontSize="11" level="1" pageId="6" pageNumber="261" reason="1">
<taxonomicName id="2AA34D28CA55FFA3FF720557FDCAFA9B" ID-CoL="6BMMN" authority="Bernhauer, 1928" authorityName="Bernhauer (FMNH" authorityYear="1928" box="[151,604,1303,1329]" class="Insecta" family="Staphylinidae" genus="Ctenopeuca" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="261" phylum="Arthropoda" rank="species" species="romani">
<emphasis id="DFD7EAB9CA55FFA3FF720557FDCAFA9B" bold="true" box="[151,604,1303,1329]" pageId="6" pageNumber="261">
<emphasis id="DFD7EAB9CA55FFA3FF720557FE10FA9B" bold="true" box="[151,390,1303,1329]" italics="true" pageId="6" pageNumber="261">Ctenopeuca romani</emphasis>
<bibRefCitation id="89324B5ACA55FFA3FE680557FDCAFA9B" author="Bernhauer, M." box="[397,604,1303,1329]" pageId="6" pageNumber="261" pagination="45 - 47" refId="ref5087" refString="Bernhauer, M. (1928) Neue Staphyliniden aus der Ausbeute der schwedischen Amazonasexpeditionem. Entomologisk Tidsrift, 49, 45 - 47." type="journal article" year="1928">Bernhauer, 1928</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A5B96520CA55FFA3FF72057BFE49F883" pageId="6" pageNumber="261" type="description">
<paragraph id="ED1C36ABCA55FFA3FF72057BFEE2FAFF" blockId="6.[151,604,1303,1365]" box="[151,372,1339,1365]" pageId="6" pageNumber="261">
(
<figureCitation id="75982A2ECA55FFA3FF7A057BFE8FFAFF" box="[159,281,1339,1365]" captionStart-0="FIGURES 16" captionStart-1="FIGURES 716" captionStart-2="FIGURES 1720" captionStartId-0="3.[152,270,1658,1682]" captionStartId-1="4.[152,270,1873,1897]" captionStartId-2="5.[152,270,1490,1514]" captionTargetBox-0="[151,1436,181,1635]" captionTargetBox-1="[219,1366,181,1849]" captionTargetBox-2="[151,1436,181,1466]" captionTargetId-0="figure-22@3.[151,1436,181,1635]" captionTargetId-1="figure-24@4.[219,1369,181,1849]" captionTargetId-2="figure-22@5.[151,1436,181,1466]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURES 16. Habitus of Ctenopeuca. 13) Lectotype of C. heynei Bernhauer (FMNH): 1) lateral view (ST3 indicates spine of tergum III, SS3 indicates spine of sternum III, ST4 indicates spine of tergum IV, SS4 indicates spine of sternum IV, ST7 indicates spine of tergum VII), 2) dorsal view, 3) ventral view. 13) Lectotype of C. romani Bernhauer (FMNH): 4) lateral view, 5) dorsal view, 6) ventral view. Scale bars 3.0 mm." captionText-1="FIGURES 716. Ctenopeuca romani: 7) Antenna; 8) Labrum, dorsal view; 9) Detail of labrum, dorsal view (bold arrow indicates a-sensilla, thin arrow indicates b-sensilla); 10) Right mandible, dorsal view; 11) Left mandible, dorsal view; 12) Maxilla, dorsal view; 13) Labium, ventral view; 14) Head, ventral view (arrow indicates infraocular carina); 15) Meso- and metaventrite, ventral view; 16) Abdomen, dorsal view (ST3 indicates spine of tergum III, SS3 indicates spine of sternum III, ST4 indicates spine of tergum IV, SS4 indicates spine of sternum IV, ST7 indicates spine of tergum VII)." captionText-2="FIGURES 1720. Ctenopeuca romani: 17) Median lobe, lateral view; 18) Median lobe, ventro-lateral view; 19) Apex of median lobe, lateral view; 20) Paramere, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.13618396" figureDoi-1="http://doi.org/10.5281/zenodo.13618400" figureDoi-2="http://doi.org/10.5281/zenodo.13618404" httpUri-0="https://zenodo.org/record/13618396/files/figure.png" httpUri-1="https://zenodo.org/record/13618400/files/figure.png" httpUri-2="https://zenodo.org/record/13618404/files/figure.png" pageId="6" pageNumber="261">Figs. 420</figureCitation>
,
<figureCitation id="75982A2ECA55FFA3FEC0057BFEFBFAFF" box="[293,365,1339,1365]" captionStart="FIGURES 2833" captionStartId="8.[152,270,1865,1889]" captionTargetBox="[158,1429,194,1834]" captionTargetId="figure-24@8.[151,1436,181,1841]" captionTargetPageId="8" captionText="FIGURES 2833. Ctenopeuca romani: 28) Tergum VIII, male, dorsal view; 29) Sternum VIII, male, dorsal view; 30) Median lobe, lateral view; 31) Spermatheca; 32) Tergum VIII, female, dorsal view; 33) Sternum VIII, female, dorsal view. Scale bars 0.6mm." figureDoi="http://doi.org/10.5281/zenodo.13618412" httpUri="https://zenodo.org/record/13618412/files/figure.png" pageId="6" pageNumber="261">2833</figureCitation>
)
</paragraph>
<paragraph id="ED1C36ABCA55FFA3FF7205C4FB22F933" blockId="6.[151,1437,1411,1978]" pageId="6" pageNumber="261">
Type material.
<materialsCitation id="5DCB3CF6CA55FFA3FEA605C4FB22F933" collectionCode="T, FMNH, FIELD, MUSEUM" collectorName="Sv. Amaz. &amp; M. Bernhauer &amp; S. Ware" location="Labels" pageId="6" pageNumber="261" specimenCount="2" stateProvince="Bahia" typeStatus="lectotype">
<typeStatus id="32188809CA55FFA3FEA605C4FE25FA34" box="[323,435,1412,1438]" pageId="6" pageNumber="261" type="lectotype">Lectotype</typeStatus>
here designated (
<figureCitation id="75982A2ECA55FFA3FD9B05C3FD7BFA37" box="[638,749,1411,1437]" captionStart="FIGURES 16" captionStartId="3.[152,270,1658,1682]" captionTargetBox="[151,1436,181,1635]" captionTargetId="figure-22@3.[151,1436,181,1635]" captionTargetPageId="3" captionText="FIGURES 16. Habitus of Ctenopeuca. 13) Lectotype of C. heynei Bernhauer (FMNH): 1) lateral view (ST3 indicates spine of tergum III, SS3 indicates spine of sternum III, ST4 indicates spine of tergum IV, SS4 indicates spine of sternum IV, ST7 indicates spine of tergum VII), 2) dorsal view, 3) ventral view. 13) Lectotype of C. romani Bernhauer (FMNH): 4) lateral view, 5) dorsal view, 6) ventral view. Scale bars 3.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13618396" httpUri="https://zenodo.org/record/13618396/files/figure.png" pageId="6" pageNumber="261">Figs. 46</figureCitation>
).
<location id="E87C6070CA55FFA3FCE605C3FCC4FA37" LSID="urn:lsid:plazi:treatment:650A87BDCA55FFACFF720557FE01F870:E87C6070CA55FFA3FCE605C3FCC4FA37" box="[771,850,1411,1437]" name="Labels" pageId="6" pageNumber="261" stateProvince="Bahia">Labels</location>
: 1) “
<collectingRegion id="2F67F849CA55FFA3FC6D05C3FC5CFA37" box="[904,970,1411,1437]" country="Brazil" name="Bahia" pageId="6" pageNumber="261">Bahia</collectingRegion>
/Iguassú/Roman” [white label, printed in black]; 2) “
<collectorName id="4056537DCA55FFA3FEF305E7FE1CFA6B" box="[278,394,1447,1473]" pageId="6" pageNumber="261">Sv. Amaz.</collectorName>
/Exp. Roman” [white label, printed in black]; 3) “Chicago NHMus/
<collectorName id="4056537DCA55FFA3FB6505E7FA85FA6B" box="[1152,1299,1447,1473]" pageId="6" pageNumber="261">M.Bernhauer</collectorName>
/Collection” [white label, printed in black]; 4) “10 juli” [white label, numbers manuscript and letters printed in black]; 5) “
<taxonomicName id="2AA34D28CA55FFA3FF4105B0FEBCF9A0" box="[164,298,1520,1546]" class="Insecta" family="Staphylinidae" genus="Ctenopeuca" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="DFD7EAB9CA55FFA3FF4105B0FEBCF9A0" box="[164,298,1520,1546]" italics="true" pageId="6" pageNumber="261">Ctenopeuca</emphasis>
</taxonomicName>
/Romani./Bernh. Cotypus” [white old label, manuscript]; 6) [male simbol, white label, manuscript]; 7) “
<typeStatus id="32188809CA55FFA3FF410653FE85F987" box="[164,275,1555,1581]" pageId="6" pageNumber="261" type="lectotype">Lectotype</typeStatus>
/
<taxonomicName id="2AA34D28CA55FFA3FEFE0654FE37F984" box="[283,417,1556,1582]" class="Insecta" family="Staphylinidae" genus="Ctenopeuca" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="DFD7EAB9CA55FFA3FEFE0654FE37F984" box="[283,417,1556,1582]" italics="true" pageId="6" pageNumber="261">Ctenopeuca</emphasis>
</taxonomicName>
/romani [male simbol]/des. K.
<collectionCode id="8BB2AE6ECA55FFA3FD020653FD6EF987" box="[743,760,1555,1581]" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="261">T</collectionCode>
.Eldredge 2013” [red label, the first and last line (except 2013) printed in black, the rest manuscript]; 8) “
<collectionCode id="8BB2AE6ECA55FFA3FDBB0677FD22F9FB" box="[606,692,1591,1617]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="6" pageNumber="261" type="Museum">FMNH</collectionCode>
” [white label, printed in black]; 9) “QR Code/FMNHINS/3982449/
<collectionCode id="8BB2AE6ECA55FFA3FF72061CFEFAF9DF" box="[151,364,1627,1654]" pageId="6" pageNumber="261">FIELD MUSEUM</collectionCode>
/Pinned” [white label, printed in black]; 10) “PHOTOGRAPHED/
<collectorName id="4056537DCA55FFA3FB81061BFB21F9DC" box="[1124,1207,1627,1654]" pageId="6" pageNumber="261">S. Ware</collectorName>
2021” [white label, prinited in black]. Note: Bernahuer (1928) did not specify how many specimens were studied.
</materialsCitation>
</paragraph>
<paragraph id="ED1C36ABCA55FFA3FF2206E3FDBAF917" blockId="6.[151,1437,1411,1978]" box="[199,556,1699,1725]" pageId="6" pageNumber="261">
<materialsCitation id="5DCB3CF6CA55FFA3FF2206E3FDBEF917" box="[199,552,1699,1725]" collectionCode="FMNH" pageId="6" pageNumber="261" specimenCount="1" typeStatus="paralectotype">
<typeStatus id="32188809CA55FFA3FF2206E3FEC9F917" box="[199,351,1699,1725]" pageId="6" pageNumber="261" type="paralectotype">Paralectotype</typeStatus>
(
<collectionCode id="8BB2AE6ECA55FFA3FE8B06E3FE53F917" box="[366,453,1699,1725]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="6" pageNumber="261" type="Museum">FMNH</collectionCode>
), female
</materialsCitation>
.
</paragraph>
<paragraph id="ED1C36ABCA55FFA3FF220687FE49F883" blockId="6.[151,1437,1411,1978]" pageId="6" pageNumber="261">
Additional material.
<materialsCitation id="5DCB3CF6CA55FFA3FE490687FE4DF883" collectingDate="2010-11-21" collectionCode="CESP" collectorName="Rio das Contas" country="Brazil" latitude="-13.527361" location="Bahia" longLatPrecision="1" longitude="-41.73275" pageId="6" pageNumber="261" specimenCount="105" stateProvince="Bahia">
<specimenCount id="FBA5FD22CA55FFA3FE490687FDC6F94B" box="[428,592,1735,1762]" count="105" pageId="6" pageNumber="261" type="generic">105 specimens</specimenCount>
deposited in
<collectionCode id="8BB2AE6ECA55FFA3FD3A0687FCB7F94B" box="[735,801,1735,1761]" pageId="6" pageNumber="261">CESP</collectionCode>
from
<collectingCountry id="95B4763BCA55FFA3FC840687FC3EF94B" box="[865,936,1735,1761]" name="Brazil" pageId="6" pageNumber="261">Brazil</collectingCountry>
,
<collectingRegion id="2F67F849CA55FFA3FC570687FC60F94B" box="[946,1014,1735,1761]" country="Brazil" name="Bahia" pageId="6" pageNumber="261">Bahia</collectingRegion>
,
<collectorName id="4056537DCA55FFA3FC1A0687FB3EF94B" box="[1023,1192,1735,1761]" pageId="6" pageNumber="261">Rio das Contas</collectorName>
, Umbuzeiro, Chapada Diamantina,
<geoCoordinate id="8897506CCA55FFA3FEC206ABFE56F8AC" box="[295,448,1771,1798]" degrees="13" direction="south" minutes="31" orientation="latitude" pageId="6" pageNumber="261" precision="1" seconds="38.5" value="-13.527361">S13°3138.5”</geoCoordinate>
<geoCoordinate id="8897506CCA55FFA3FE2306ACFDFBF8AC" box="[454,621,1771,1798]" degrees="41" direction="west" minutes="43" orientation="longitude" pageId="6" pageNumber="261" precision="1" seconds="57.9" value="-41.73275">W41°4357.9”</geoCoordinate>
,
<date id="991D106BCA55FFA3FD9D06ABFD63F8AF" box="[632,757,1771,1797]" pageId="6" pageNumber="261" value="2010-11-21">
<collectingDate id="8959E983CA55FFA3FD9D06ABFD63F8AF" box="[632,757,1771,1797]" pageId="6" pageNumber="261" value="2010-11-21">21/xi/2010</collectingDate>
</date>
, collected in
<taxonomicName id="2AA34D28CA55FFA3FC6106ABFE3CF883" authority="Mart. &amp; Zucc., J. Hipolito, I. Manimann &amp; I. Abreu" authorityName="Mart. &amp; Zucc., J. Hipolito, I. Manimann &amp; I. Abreu" class="Magnoliopsida" family="Aristolochiaceae" genus="Aristolochia" kingdom="Plantae" order="Piperales" pageId="6" pageNumber="261" phylum="Tracheophyta" rank="species" species="gigantea">Aristolochia gigantea Mart. &amp; Zucc., J.Hipolito, I. Manimann &amp; I. Abreu</taxonomicName>
, col
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A5B96520CA55FFA3FF220773FAEFF813" pageId="6" pageNumber="261" type="diagnosis">
<paragraph id="ED1C36ABCA55FFA3FF220773FAEFF813" blockId="6.[151,1437,1411,1978]" pageId="6" pageNumber="261">
<emphasis id="DFD7EAB9CA55FFA3FF220773FED4F8E7" bold="true" box="[199,322,1843,1869]" pageId="6" pageNumber="261">Diagnosis.</emphasis>
<taxonomicName id="2AA34D28CA55FFA3FEAE0774FDBCF8E7" authorityName="Bernhauer (FMNH" authorityYear="1928" box="[331,554,1844,1870]" class="Insecta" family="Staphylinidae" genus="Ctenopeuca" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="261" phylum="Arthropoda" rank="species" species="romani">
<emphasis id="DFD7EAB9CA55FFA3FEAE0774FDBCF8E7" box="[331,554,1844,1870]" italics="true" pageId="6" pageNumber="261">Ctenopeuca romani</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2AA34D28CA55FFA3FD200774FCA7F8E7" authorityName="Bernhauer (FMNH" authorityYear="1915" box="[709,817,1843,1870]" class="Insecta" family="Staphylinidae" genus="Ctenopeuca" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="261" phylum="Arthropoda" rank="species" species="heynei">
<emphasis id="DFD7EAB9CA55FFA3FD200774FCA7F8E7" box="[709,817,1843,1870]" italics="true" pageId="6" pageNumber="261">C. heynei</emphasis>
</taxonomicName>
by having head darker than pronotum and abdominal segments IIIIV conspicuously lighter than VVII (
<figureCitation id="75982A2ECA55FFA3FD160717FCAAF8D8" box="[755,828,1879,1906]" captionStart="FIGURES 16" captionStartId="3.[152,270,1658,1682]" captionTargetBox="[151,1436,181,1635]" captionTargetId="figure-22@3.[151,1436,181,1635]" captionTargetPageId="3" captionText="FIGURES 16. Habitus of Ctenopeuca. 13) Lectotype of C. heynei Bernhauer (FMNH): 1) lateral view (ST3 indicates spine of tergum III, SS3 indicates spine of sternum III, ST4 indicates spine of tergum IV, SS4 indicates spine of sternum IV, ST7 indicates spine of tergum VII), 2) dorsal view, 3) ventral view. 13) Lectotype of C. romani Bernhauer (FMNH): 4) lateral view, 5) dorsal view, 6) ventral view. Scale bars 3.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13618396" httpUri="https://zenodo.org/record/13618396/files/figure.png" pageId="6" pageNumber="261">Fig. 5</figureCitation>
); antennomere 4 wider than long (
<figureCitation id="75982A2ECA55FFA3FB330717FA89F8DB" box="[1238,1311,1879,1905]" captionStart="FIGURES 716" captionStartId="4.[152,270,1873,1897]" captionTargetBox="[219,1366,181,1849]" captionTargetId="figure-24@4.[219,1369,181,1849]" captionTargetPageId="4" captionText="FIGURES 716. Ctenopeuca romani: 7) Antenna; 8) Labrum, dorsal view; 9) Detail of labrum, dorsal view (bold arrow indicates a-sensilla, thin arrow indicates b-sensilla); 10) Right mandible, dorsal view; 11) Left mandible, dorsal view; 12) Maxilla, dorsal view; 13) Labium, ventral view; 14) Head, ventral view (arrow indicates infraocular carina); 15) Meso- and metaventrite, ventral view; 16) Abdomen, dorsal view (ST3 indicates spine of tergum III, SS3 indicates spine of sternum III, ST4 indicates spine of tergum IV, SS4 indicates spine of sternum IV, ST7 indicates spine of tergum VII)." figureDoi="http://doi.org/10.5281/zenodo.13618400" httpUri="https://zenodo.org/record/13618400/files/figure.png" pageId="6" pageNumber="261">Fig. 7</figureCitation>
); posterior margin of abdominal tergum VIII of male without a prominent tooth on each side of serrate region (
<figureCitation id="75982A2ECA55FFA3FAD2073BFA19F83F" box="[1335,1423,1915,1941]" captionStart="FIGURES 2833" captionStartId="8.[152,270,1865,1889]" captionTargetBox="[158,1429,194,1834]" captionTargetId="figure-24@8.[151,1436,181,1841]" captionTargetPageId="8" captionText="FIGURES 2833. Ctenopeuca romani: 28) Tergum VIII, male, dorsal view; 29) Sternum VIII, male, dorsal view; 30) Median lobe, lateral view; 31) Spermatheca; 32) Tergum VIII, female, dorsal view; 33) Sternum VIII, female, dorsal view. Scale bars 0.6mm." figureDoi="http://doi.org/10.5281/zenodo.13618412" httpUri="https://zenodo.org/record/13618412/files/figure.png" pageId="6" pageNumber="261">Fig. 28</figureCitation>
); posterior margin of abdominal tergum VIII of female without tooth on each side of emarginate region (
<figureCitation id="75982A2ECA55FFA3FAFD07DFFAFAF813" box="[1304,1388,1951,1977]" captionStart="FIGURES 2833" captionStartId="8.[152,270,1865,1889]" captionTargetBox="[158,1429,194,1834]" captionTargetId="figure-24@8.[151,1436,181,1841]" captionTargetPageId="8" captionText="FIGURES 2833. Ctenopeuca romani: 28) Tergum VIII, male, dorsal view; 29) Sternum VIII, male, dorsal view; 30) Median lobe, lateral view; 31) Spermatheca; 32) Tergum VIII, female, dorsal view; 33) Sternum VIII, female, dorsal view. Scale bars 0.6mm." figureDoi="http://doi.org/10.5281/zenodo.13618412" httpUri="https://zenodo.org/record/13618412/files/figure.png" pageId="6" pageNumber="261">Fig. 32</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="B9DC6623CA54FFA2FF7D07D7FEF2F85D" ID-DOI="http://doi.org/10.5281/zenodo.13618408" ID-Zenodo-Dep="13618408" httpUri="https://zenodo.org/record/13618408/files/figure.png" pageId="7" pageNumber="262" startId="7.[152,270,1943,1967]" targetBox="[238,1348,187,1913]" targetPageId="7" targetType="figure">
<paragraph id="ED1C36ABCA54FFA2FF7D07D7FEF2F85D" blockId="7.[151,1436,1943,2039]" pageId="7" pageNumber="262">
<emphasis id="DFD7EAB9CA54FFA2FF7D07D7FECDF805" bold="true" box="[152,347,1943,1967]" pageId="7" pageNumber="262">FIGURES 2127.</emphasis>
<taxonomicName id="2AA34D28CA54FFA2FE8407D8FDB5F805" authorityName="Bernhauer (FMNH" authorityYear="1915" box="[353,547,1943,1967]" class="Insecta" family="Staphylinidae" genus="Ctenopeuca" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="262" phylum="Arthropoda" rank="species" species="heynei">
<emphasis id="DFD7EAB9CA54FFA2FE8407D8FDB5F805" box="[353,547,1943,1967]" italics="true" pageId="7" pageNumber="262">Ctenopeuca heynei</emphasis>
</taxonomicName>
: 21) Tergum VIII, male, dorsal view; 22) Sternum VIII, male, dorsal view; 23) Median lobe, ventral view; 24) Median lobe, lateral view; 25) Paramere; 26) Tergum VIII, female, dorsal view; 27) Sternum VIII, female, dorsal view.
</paragraph>
</caption>
<caption id="B9DC6623CA5BFFADFF7D0709FF70F803" ID-DOI="http://doi.org/10.5281/zenodo.13618412" ID-Zenodo-Dep="13618412" httpUri="https://zenodo.org/record/13618412/files/figure.png" pageId="8" pageNumber="263" startId="8.[152,270,1865,1889]" targetBox="[158,1429,194,1834]" targetPageId="8" targetType="figure">
<paragraph id="ED1C36ABCA5BFFADFF7D0709FF70F803" blockId="8.[151,1436,1865,1961]" pageId="8" pageNumber="263">
<emphasis id="DFD7EAB9CA5BFFADFF7D0709FECCF8CB" bold="true" box="[152,346,1865,1889]" pageId="8" pageNumber="263">FIGURES 2833.</emphasis>
<taxonomicName id="2AA34D28CA5BFFADFE85070AFDBEF8CB" authorityName="Bernhauer (FMNH" authorityYear="1928" box="[352,552,1866,1889]" class="Insecta" family="Staphylinidae" genus="Ctenopeuca" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="263" phylum="Arthropoda" rank="species" species="romani">
<emphasis id="DFD7EAB9CA5BFFADFE85070AFDBEF8CB" box="[352,552,1866,1889]" italics="true" pageId="8" pageNumber="263">Ctenopeuca romani</emphasis>
</taxonomicName>
: 28) Tergum VIII, male, dorsal view; 29) Sternum VIII, male, dorsal view; 30) Median lobe, lateral view; 31) Spermatheca; 32) Tergum VIII, female, dorsal view; 33) Sternum VIII, female, dorsal view. Scale bars 0.6mm.
</paragraph>
</caption>
<caption id="B9DC6623CA5AFFACFF7D051DFC39FA30" ID-DOI="http://doi.org/10.5281/zenodo.13618416" ID-Zenodo-Dep="13618416" httpUri="https://zenodo.org/record/13618416/files/figure.png" pageId="9" pageNumber="264" startId="9.[152,255,1373,1397]" targetBox="[151,1436,181,1350]" targetPageId="9" targetType="figure">
<paragraph id="ED1C36ABCA5AFFACFF7D051DFC39FA30" blockId="9.[151,1437,1373,1434]" pageId="9" pageNumber="264">
<emphasis id="DFD7EAB9CA5AFFACFF7D051DFEB3FADF" bold="true" box="[152,293,1373,1397]" pageId="9" pageNumber="264">FIGURE 34.</emphasis>
Map of
<taxonomicName id="2AA34D28CA5AFFACFE9E051EFE62FADF" box="[379,500,1374,1397]" class="Insecta" family="Staphylinidae" genus="Ctenopeuca" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="DFD7EAB9CA5AFFACFE9E051EFE62FADF" box="[379,500,1374,1397]" italics="true" pageId="9" pageNumber="264">Ctenopeuca</emphasis>
</taxonomicName>
distribution. The green spot indicates
<taxonomicName id="2AA34D28CA5AFFACFC92051EFC40FADC" authorityName="Bernhauer (FMNH" authorityYear="1915" box="[887,982,1374,1398]" class="Insecta" family="Staphylinidae" genus="Ctenopeuca" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="264" phylum="Arthropoda" rank="species" species="heynei">
<emphasis id="DFD7EAB9CA5AFFACFC92051EFC40FADC" box="[887,982,1374,1398]" italics="true" pageId="9" pageNumber="264">C. heynei</emphasis>
</taxonomicName>
(not exact record), the pink spot indicates
<taxonomicName id="2AA34D28CA5AFFACFA60051EFF49FA33" authorityName="Bernhauer (FMNH" authorityYear="1928" class="Insecta" family="Staphylinidae" genus="Ctenopeuca" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="264" phylum="Arthropoda" rank="species" species="romani">
<emphasis id="DFD7EAB9CA5AFFACFA60051EFF49FA33" italics="true" pageId="9" pageNumber="264">C. romani</emphasis>
</taxonomicName>
(not exact record), and the red spot indicates
<taxonomicName id="2AA34D28CA5AFFACFD4B05C2FC82FA33" authorityName="Bernhauer (FMNH" authorityYear="1928" box="[686,788,1410,1433]" class="Insecta" family="Staphylinidae" genus="Ctenopeuca" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="264" phylum="Arthropoda" rank="species" species="romani">
<emphasis id="DFD7EAB9CA5AFFACFD4B05C2FC82FA33" box="[686,788,1410,1433]" italics="true" pageId="9" pageNumber="264">C. romani</emphasis>
</taxonomicName>
(exact record).
</paragraph>
</caption>
<subSubSection id="A5B96520CA5AFFACFF2205ACFC47F88C" pageId="9" pageNumber="264" type="description">
<paragraph id="ED1C36ABCA5AFFACFF2205ACFC47F88C" blockId="9.[151,1437,1516,2010]" pageId="9" pageNumber="264">
<emphasis id="DFD7EAB9CA5AFFACFF2205ACFEE3F9AC" bold="true" box="[199,373,1516,1542]" pageId="9" pageNumber="264">Redescription.</emphasis>
Male. Body length, mean
<quantity id="2A5B9B4ECA5AFFACFD7C05ACFD68F9AC" box="[665,766,1516,1543]" metricMagnitude="-3" metricUnit="m" metricValue="5.57" pageId="9" pageNumber="264" unit="mm" value="5.57">5.57 mm</quantity>
, standard deviation
<quantity id="2A5B9B4ECA5AFFACFC3905ACFBD7F9AC" box="[988,1089,1516,1542]" metricMagnitude="-4" metricUnit="m" metricValue="8.6" pageId="9" pageNumber="264" unit="mm" value="0.86">0.86 mm</quantity>
. Humeral width 1.0 mm. Head, antennomeres 411, posterolateral angle region of elytra and abdominal segments VVII brown to dark brown, the rest of body yellowish to light brown (
<figureCitation id="75982A2ECA5AFFACFD640674FD5FF9E4" box="[641,713,1588,1614]" captionStart="FIGURES 16" captionStartId="3.[152,270,1658,1682]" captionTargetBox="[151,1436,181,1635]" captionTargetId="figure-22@3.[151,1436,181,1635]" captionTargetPageId="3" captionText="FIGURES 16. Habitus of Ctenopeuca. 13) Lectotype of C. heynei Bernhauer (FMNH): 1) lateral view (ST3 indicates spine of tergum III, SS3 indicates spine of sternum III, ST4 indicates spine of tergum IV, SS4 indicates spine of sternum IV, ST7 indicates spine of tergum VII), 2) dorsal view, 3) ventral view. 13) Lectotype of C. romani Bernhauer (FMNH): 4) lateral view, 5) dorsal view, 6) ventral view. Scale bars 3.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13618396" httpUri="https://zenodo.org/record/13618396/files/figure.png" pageId="9" pageNumber="264">Fig. 2</figureCitation>
). Antennomere 4 wider than long (
<figureCitation id="75982A2ECA5AFFACFBB80674FB35F9E4" box="[1117,1187,1588,1614]" captionStart="FIGURES 716" captionStartId="4.[152,270,1873,1897]" captionTargetBox="[219,1366,181,1849]" captionTargetId="figure-24@4.[219,1369,181,1849]" captionTargetPageId="4" captionText="FIGURES 716. Ctenopeuca romani: 7) Antenna; 8) Labrum, dorsal view; 9) Detail of labrum, dorsal view (bold arrow indicates a-sensilla, thin arrow indicates b-sensilla); 10) Right mandible, dorsal view; 11) Left mandible, dorsal view; 12) Maxilla, dorsal view; 13) Labium, ventral view; 14) Head, ventral view (arrow indicates infraocular carina); 15) Meso- and metaventrite, ventral view; 16) Abdomen, dorsal view (ST3 indicates spine of tergum III, SS3 indicates spine of sternum III, ST4 indicates spine of tergum IV, SS4 indicates spine of sternum IV, ST7 indicates spine of tergum VII)." figureDoi="http://doi.org/10.5281/zenodo.13618400" httpUri="https://zenodo.org/record/13618400/files/figure.png" pageId="9" pageNumber="264">Fig. 7</figureCitation>
). Abdominal spine of tergum IV triangular in lateral view (
<figureCitation id="75982A2ECA5AFFACFDD90618FD06F9D8" box="[572,656,1624,1650]" captionStart="FIGURES 716" captionStartId="4.[152,270,1873,1897]" captionTargetBox="[219,1366,181,1849]" captionTargetId="figure-24@4.[219,1369,181,1849]" captionTargetPageId="4" captionText="FIGURES 716. Ctenopeuca romani: 7) Antenna; 8) Labrum, dorsal view; 9) Detail of labrum, dorsal view (bold arrow indicates a-sensilla, thin arrow indicates b-sensilla); 10) Right mandible, dorsal view; 11) Left mandible, dorsal view; 12) Maxilla, dorsal view; 13) Labium, ventral view; 14) Head, ventral view (arrow indicates infraocular carina); 15) Meso- and metaventrite, ventral view; 16) Abdomen, dorsal view (ST3 indicates spine of tergum III, SS3 indicates spine of sternum III, ST4 indicates spine of tergum IV, SS4 indicates spine of sternum IV, ST7 indicates spine of tergum VII)." figureDoi="http://doi.org/10.5281/zenodo.13618400" httpUri="https://zenodo.org/record/13618400/files/figure.png" pageId="9" pageNumber="264">Fig. 16</figureCitation>
, ST4); spine of sternum III slightly longer than the spine of sternum IV (
<figureCitation id="75982A2ECA5AFFACFF21063CFE81F93C" box="[196,279,1660,1686]" captionStart="FIGURES 716" captionStartId="4.[152,270,1873,1897]" captionTargetBox="[219,1366,181,1849]" captionTargetId="figure-24@4.[219,1369,181,1849]" captionTargetPageId="4" captionText="FIGURES 716. Ctenopeuca romani: 7) Antenna; 8) Labrum, dorsal view; 9) Detail of labrum, dorsal view (bold arrow indicates a-sensilla, thin arrow indicates b-sensilla); 10) Right mandible, dorsal view; 11) Left mandible, dorsal view; 12) Maxilla, dorsal view; 13) Labium, ventral view; 14) Head, ventral view (arrow indicates infraocular carina); 15) Meso- and metaventrite, ventral view; 16) Abdomen, dorsal view (ST3 indicates spine of tergum III, SS3 indicates spine of sternum III, ST4 indicates spine of tergum IV, SS4 indicates spine of sternum IV, ST7 indicates spine of tergum VII)." figureDoi="http://doi.org/10.5281/zenodo.13618400" httpUri="https://zenodo.org/record/13618400/files/figure.png" pageId="9" pageNumber="264">Fig. 16</figureCitation>
, SS3 and SS4); tergum VIII with posterior margin emarginate and serrate (
<figureCitation id="75982A2ECA5AFFACFB82063CFB2AF93C" box="[1127,1212,1660,1686]" captionStart="FIGURES 2833" captionStartId="8.[152,270,1865,1889]" captionTargetBox="[158,1429,194,1834]" captionTargetId="figure-24@8.[151,1436,181,1841]" captionTargetPageId="8" captionText="FIGURES 2833. Ctenopeuca romani: 28) Tergum VIII, male, dorsal view; 29) Sternum VIII, male, dorsal view; 30) Median lobe, lateral view; 31) Spermatheca; 32) Tergum VIII, female, dorsal view; 33) Sternum VIII, female, dorsal view. Scale bars 0.6mm." figureDoi="http://doi.org/10.5281/zenodo.13618412" httpUri="https://zenodo.org/record/13618412/files/figure.png" pageId="9" pageNumber="264">Fig. 28</figureCitation>
); sternum VIII with posterior margin projected medially (
<figureCitation id="75982A2ECA5AFFACFDD906E0FD04F910" box="[572,658,1696,1722]" captionStart="FIGURES 2833" captionStartId="8.[152,270,1865,1889]" captionTargetBox="[158,1429,194,1834]" captionTargetId="figure-24@8.[151,1436,181,1841]" captionTargetPageId="8" captionText="FIGURES 2833. Ctenopeuca romani: 28) Tergum VIII, male, dorsal view; 29) Sternum VIII, male, dorsal view; 30) Median lobe, lateral view; 31) Spermatheca; 32) Tergum VIII, female, dorsal view; 33) Sternum VIII, female, dorsal view. Scale bars 0.6mm." figureDoi="http://doi.org/10.5281/zenodo.13618412" httpUri="https://zenodo.org/record/13618412/files/figure.png" pageId="9" pageNumber="264">Fig. 29</figureCitation>
); median lobe of aedeagus curved paramerally in lateral view, apical half with ventral face straight at basal two-third and curved at the apical third (
<figureCitation id="75982A2ECA5AFFACFBFF0684FBE6F974" box="[1050,1136,1732,1758]" captionStart="FIGURES 2833" captionStartId="8.[152,270,1865,1889]" captionTargetBox="[158,1429,194,1834]" captionTargetId="figure-24@8.[151,1436,181,1841]" captionTargetPageId="8" captionText="FIGURES 2833. Ctenopeuca romani: 28) Tergum VIII, male, dorsal view; 29) Sternum VIII, male, dorsal view; 30) Median lobe, lateral view; 31) Spermatheca; 32) Tergum VIII, female, dorsal view; 33) Sternum VIII, female, dorsal view. Scale bars 0.6mm." figureDoi="http://doi.org/10.5281/zenodo.13618412" httpUri="https://zenodo.org/record/13618412/files/figure.png" pageId="9" pageNumber="264">Fig. 30</figureCitation>
). Female. Similar to male, except tergum VIII with posterior margin not serrate (
<figureCitation id="75982A2ECA5AFFACFD0906A8FCD6F8A8" box="[748,832,1768,1794]" captionStart="FIGURES 2833" captionStartId="8.[152,270,1865,1889]" captionTargetBox="[158,1429,194,1834]" captionTargetId="figure-24@8.[151,1436,181,1841]" captionTargetPageId="8" captionText="FIGURES 2833. Ctenopeuca romani: 28) Tergum VIII, male, dorsal view; 29) Sternum VIII, male, dorsal view; 30) Median lobe, lateral view; 31) Spermatheca; 32) Tergum VIII, female, dorsal view; 33) Sternum VIII, female, dorsal view. Scale bars 0.6mm." figureDoi="http://doi.org/10.5281/zenodo.13618412" httpUri="https://zenodo.org/record/13618412/files/figure.png" pageId="9" pageNumber="264">Fig. 32</figureCitation>
); sternum VIII with posterior margin curved (
<figureCitation id="75982A2ECA5AFFACFADE06A8FA18F8A8" box="[1339,1422,1768,1794]" captionStart="FIGURES 2833" captionStartId="8.[152,270,1865,1889]" captionTargetBox="[158,1429,194,1834]" captionTargetId="figure-24@8.[151,1436,181,1841]" captionTargetPageId="8" captionText="FIGURES 2833. Ctenopeuca romani: 28) Tergum VIII, male, dorsal view; 29) Sternum VIII, male, dorsal view; 30) Median lobe, lateral view; 31) Spermatheca; 32) Tergum VIII, female, dorsal view; 33) Sternum VIII, female, dorsal view. Scale bars 0.6mm." figureDoi="http://doi.org/10.5281/zenodo.13618412" httpUri="https://zenodo.org/record/13618412/files/figure.png" pageId="9" pageNumber="264">Fig. 33</figureCitation>
); spermatheca L-shaped, capsule globose and without coiled duct (
<figureCitation id="75982A2ECA5AFFACFC95074CFC52F88C" box="[880,964,1804,1830]" captionStart="FIGURES 2833" captionStartId="8.[152,270,1865,1889]" captionTargetBox="[158,1429,194,1834]" captionTargetId="figure-24@8.[151,1436,181,1841]" captionTargetPageId="8" captionText="FIGURES 2833. Ctenopeuca romani: 28) Tergum VIII, male, dorsal view; 29) Sternum VIII, male, dorsal view; 30) Median lobe, lateral view; 31) Spermatheca; 32) Tergum VIII, female, dorsal view; 33) Sternum VIII, female, dorsal view. Scale bars 0.6mm." figureDoi="http://doi.org/10.5281/zenodo.13618412" httpUri="https://zenodo.org/record/13618412/files/figure.png" pageId="9" pageNumber="264">Fig. 31</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A5B96520CA5AFFACFF220770FE01F870" pageId="9" pageNumber="264" type="distribution">
<paragraph id="ED1C36ABCA5AFFACFF220770FD5DF8E0" blockId="9.[151,1437,1516,2010]" box="[199,715,1840,1866]" pageId="9" pageNumber="264">
<emphasis id="DFD7EAB9CA5AFFACFF220770FEF7F8E0" bold="true" box="[199,353,1840,1866]" pageId="9" pageNumber="264">Distribution.</emphasis>
<collectingCountry id="95B4763BCA5AFFACFE820770FE26F8E0" box="[359,432,1840,1866]" name="Brazil" pageId="9" pageNumber="264">Brazil</collectingCountry>
:
<collectingRegion id="2F67F849CA5AFFACFE5E0770FE6AF8E0" box="[443,508,1840,1866]" country="Brazil" name="Bahia" pageId="9" pageNumber="264">Bahia</collectingRegion>
(Iguaçú) (
<figureCitation id="75982A2ECA5AFFACFD940770FD54F8E0" box="[625,706,1840,1866]" captionStart="FIGURE 34" captionStartId="9.[152,255,1373,1397]" captionTargetBox="[151,1436,181,1350]" captionTargetId="figure-22@9.[151,1436,181,1350]" captionTargetPageId="9" captionText="FIGURE 34. Map of Ctenopeuca distribution. The green spot indicates C. heynei (not exact record), the pink spot indicates C. romani (not exact record), and the red spot indicates C. romani (exact record)." figureDoi="http://doi.org/10.5281/zenodo.13618416" httpUri="https://zenodo.org/record/13618416/files/figure.png" pageId="9" pageNumber="264">Fig. 34</figureCitation>
)
</paragraph>
<paragraph id="ED1C36ABCA5AFFACFF220714FE01F870" blockId="9.[151,1437,1516,2010]" pageId="9" pageNumber="264">
<emphasis id="DFD7EAB9CA5AFFACFF220714FE11F8C4" bold="true" box="[199,391,1876,1902]" pageId="9" pageNumber="264">Biological notes.</emphasis>
<taxonomicName id="2AA34D28CA5AFFACFE6E0714FDF3F8C4" authorityName="Bernhauer (FMNH" authorityYear="1928" box="[395,613,1876,1902]" class="Insecta" family="Staphylinidae" genus="Ctenopeuca" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="264" phylum="Arthropoda" rank="species" species="romani">
<emphasis id="DFD7EAB9CA5AFFACFE6E0714FDF3F8C4" box="[395,613,1876,1902]" italics="true" pageId="9" pageNumber="264">Ctenopeuca romani</emphasis>
</taxonomicName>
is associated to the flower of
<taxonomicName id="2AA34D28CA5AFFACFC450714FBBCF8C4" box="[928,1066,1876,1902]" class="Magnoliopsida" family="Aristolochiaceae" genus="Aristolochia" kingdom="Plantae" order="Piperales" pageId="9" pageNumber="264" phylum="Tracheophyta" rank="genus">
<emphasis id="DFD7EAB9CA5AFFACFC450714FBBCF8C4" box="[928,1066,1876,1902]" italics="true" pageId="9" pageNumber="264">Aristolochia</emphasis>
</taxonomicName>
L. (
<taxonomicName id="2AA34D28CA5AFFACFBB60714FA85F8C4" box="[1107,1299,1876,1902]" class="Magnoliopsida" family="Aristolochiaceae" kingdom="Plantae" order="Piperales" pageId="9" pageNumber="264" phylum="Tracheophyta" rank="family">Aristolochiaceae</taxonomicName>
) (
<bibRefCitation id="89324B5ACA5AFFACFAC30714FF46F838" author="Bernhauer, M." pageId="9" pageNumber="264" pagination="45 - 47" refId="ref5087" refString="Bernhauer, M. (1928) Neue Staphyliniden aus der Ausbeute der schwedischen Amazonasexpeditionem. Entomologisk Tidsrift, 49, 45 - 47." type="journal article" year="1928">Bernhauer 1928</bibRefCitation>
) and in 2015 all specimens were collected in
<taxonomicName id="2AA34D28CA5AFFACFD010738FB80F838" authority="Mart. &amp; Zucc." authorityName="Mart. &amp; Zucc." box="[740,1046,1912,1938]" class="Magnoliopsida" family="Aristolochiaceae" genus="Aristolochia" kingdom="Plantae" order="Piperales" pageId="9" pageNumber="264" phylum="Tracheophyta" rank="species" species="gigantea">
<emphasis id="DFD7EAB9CA5AFFACFD010738FCF1F838" box="[740,871,1912,1938]" italics="true" pageId="9" pageNumber="264">A. gigantea</emphasis>
Mart. &amp; Zucc.
</taxonomicName>
The host is an endemic liana that occurs in the East and South of
<collectingCountry id="95B4763BCA5AFFACFE1807DCFDD2F81C" box="[509,580,1948,1974]" name="Brazil" pageId="9" pageNumber="264">Brazil</collectingCountry>
, such as
<collectingRegion id="2F67F849CA5AFFACFD4D07DCFD7AF81C" box="[680,748,1948,1974]" country="Brazil" name="Bahia" pageId="9" pageNumber="264">Bahia</collectingRegion>
,
<collectingRegion id="2F67F849CA5AFFACFD1D07DCFC07F81C" box="[760,913,1948,1974]" country="Brazil" name="Minas Gerais" pageId="9" pageNumber="264">Minas Gerais</collectingRegion>
,
<collectingRegion id="2F67F849CA5AFFACFC7807DCFBD6F81C" box="[925,1088,1948,1974]" country="Brazil" name="Espirito Santo" pageId="9" pageNumber="264">Espirito Santo</collectingRegion>
,
<collectingRegion id="2F67F849CA5AFFACFBAE07DCFB66F81C" box="[1099,1264,1948,1974]" country="Brazil" name="Rio de Janeiro" pageId="9" pageNumber="264">Rio de Janeiro</collectingRegion>
,
<collectingRegion id="2F67F849CA5AFFACFB1E07DCFAFBF81C" box="[1275,1389,1948,1974]" country="Brazil" name="Sao Paulo" pageId="9" pageNumber="264">São Paulo</collectingRegion>
and
<collectingRegion id="2F67F849CA5AFFACFF720780FF75F870" box="[151,227,1984,2010]" country="Brazil" name="Parana" pageId="9" pageNumber="264">Paraná</collectingRegion>
(
<bibRefCitation id="89324B5ACA5AFFACFF110780FE1AF870" author="Reflora" box="[244,396,1984,2010]" pageId="9" pageNumber="264" refId="ref6114" refString="Reflora (2023) Aristolochiaceae in Flora e Funga do Brasil. Jardim Botanico do Rio de Janeiro. Available from: https: // floradobrasil. jbrj. gov. br / FB 15749 (accessed 18 October 2023)" type="url" year="2023">Reflora 2023</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,227 @@
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<mods:identifier id="A4815066503AD01EB09FC96B7F8127D0" type="ZooBank">59DCFD3A-ED77-48C9-BEE5-412E07CDF522</mods:identifier>
</mods:mods>
<treatment id="8318C873FFC2FFC378CE94AF2401FA10" ID-DOI="http://doi.org/10.5281/zenodo.13618574" ID-Zenodo-Dep="13618574" LSID="urn:lsid:zoobank.org:act:1A12AC30-BFF5-4FAF-8D09-0E92A6CB0275" httpUri="http://treatment.plazi.org/id/8318C873FFC2FFC378CE94AF2401FA10" lastPageId="3" lastPageNumber="288" pageId="2" pageNumber="287">
<subSubSection id="43AB2AEEFFC2FFC278CE94AF2492FCAB" box="[151,767,151,178]" pageId="2" pageNumber="287" type="nomenclature">
<paragraph id="0B0E7965FFC2FFC278CE94AF2492FCAB" blockId="2.[151,767,151,213]" box="[151,767,151,178]" pageId="2" pageNumber="287">
<heading id="5046CE09FFC2FFC278CE94AF2492FCAB" bold="true" box="[151,767,151,178]" fontSize="11" level="1" pageId="2" pageNumber="287" reason="1">
<emphasis id="39C5A577FFC2FFC278CE94AF2492FCAB" bold="true" box="[151,767,151,178]" pageId="2" pageNumber="287">
<taxonomicName id="CCB102E6FFC2FFC278CE94AF24F7FCAB" authority="Santos and Mazariegos" authorityName="Santos and Mazariegos" authorityYear="2024" box="[151,666,151,178]" class="Insecta" family="Aulacidae" genus="Aulacus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="287" phylum="Arthropoda" rank="species" species="gracielae" status="sp. nov.">
<emphasis id="39C5A577FFC2FFC278CE94AF271CFCA8" bold="true" box="[151,369,151,178]" italics="true" pageId="2" pageNumber="287">Aulacus gracielae</emphasis>
Santos and Mazariegos
</taxonomicName>
<taxonomicNameLabel id="22F6180CFFC2FFC27AF894AF2492FCAB" box="[673,767,151,177]" pageId="2" pageNumber="287" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="43AB2AEEFFC2FFC278CE9483276DFCCF" box="[151,256,187,213]" pageId="2" pageNumber="287" type="description">
<paragraph id="0B0E7965FFC2FFC278CE9483276DFCCF" blockId="2.[151,767,151,213]" box="[151,256,187,213]" pageId="2" pageNumber="287">
<figureCitation id="938A65E0FFC2FFC278CE94832696FCCF" box="[151,251,187,213]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="2.[152,255,1929,1953]" captionStartId-1="3.[152,255,982,1006]" captionTargetBox-0="[151,1436,921,1905]" captionTargetBox-1="[151,1436,181,956]" captionTargetId-0="figure-309@2.[151,1436,921,1905]" captionTargetId-1="figure-27@3.[151,1436,181,958]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 2: Aulacus gracielae Santos &amp; Mazariegos sp. n., female; A, lateral habitus; B, head in dorsal view; C, head in frontal view; D, head and mesosoma in lateral view; E, mesosoma in dorsal view." captionText-1="FIGURE 3: Distribution map for the species of Aulacus recorded in Colombia. The red star represents the location of the Mesenia-Paramillo nature reserve, and consequently the locality of the new record of Aulacus costaricensis Smith and the record of Aulacus gracielae Santos &amp; Mazariegos sp. nov." figureDoi-0="http://doi.org/10.5281/zenodo.13618462" figureDoi-1="http://doi.org/10.5281/zenodo.13618464" httpUri-0="https://zenodo.org/record/13618462/files/figure.png" httpUri-1="https://zenodo.org/record/13618464/files/figure.png" pageId="2" pageNumber="287">Figs 23</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="43AB2AEEFFC2FFC278CE953B2295FD5B" pageId="2" pageNumber="287" type="materials_examined">
<paragraph id="0B0E7965FFC2FFC278CE953B2295FD5B" blockId="2.[151,1436,259,897]" pageId="2" pageNumber="287">
<emphasis id="39C5A577FFC2FFC278CE953B27E8FD07" bold="true" box="[151,389,259,285]" pageId="2" pageNumber="287">Examined material.</emphasis>
<materialsCitation id="BBD97338FFC2FFC279D5953B2295FD5B" collectingDate="2020-08" collectionCode="CEUA" collectorName="L. Mazariegos" country="Colombia" county="Jardin" elevation="2280" latitude="5.519166" location="Olinguito" longLatPrecision="1" longitude="-75.86218" municipality="Mesenia-Paramillo nature reserve" pageId="2" pageNumber="287" specimenCount="1" specimenCount-female="1" stateProvince="Antioquia" typeStatus="holotype">
<emphasis id="39C5A577FFC2FFC279D5953B2446FD07" bold="true" box="[396,555,259,285]" pageId="2" pageNumber="287">
<typeStatus id="D40AC7C7FFC2FFC279D5953B2446FD07" box="[396,555,259,285]" pageId="2" pageNumber="287" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
<specimenCount id="1DB7B2ECFFC2FFC27A6A953C2424FD04" box="[563,585,260,286]" count="1" pageId="2" pageNumber="287" type="female"></specimenCount>
.
<collectingCountry id="73A639F5FFC2FFC27A0E953B24A5FD07" box="[599,712,259,285]" name="Colombia" pageId="2" pageNumber="287">Colombia</collectingCountry>
,
<collectingRegion id="C975B787FFC2FFC27A8B953B252AFD07" box="[722,839,259,285]" country="Colombia" name="Antioquia" pageId="2" pageNumber="287">Antioquia</collectingRegion>
:
<collectingCounty id="E26F01E9FFC2FFC27B0B953B25F7FD07" box="[850,922,259,285]" pageId="2" pageNumber="287">Jardín</collectingCounty>
;
<collectingMunicipality id="EB6AE31FFFC2FFC27BFF953B2349FD04" box="[934,1316,259,286]" pageId="2" pageNumber="287">Mesenia-Paramillo nature reserve</collectingMunicipality>
;
<location id="0E6E2FBEFFC2FFC27D68953B23F1FD07" LSID="urn:lsid:plazi:treatment:8318C873FFC2FFC378CE94AF2401FA10:0E6E2FBEFFC2FFC27D68953B23F1FD07" box="[1329,1436,259,285]" country="Colombia" county="Jardin" latitude="5.519166" longLatPrecision="1" longitude="-75.86218" municipality="Mesenia-Paramillo nature reserve" name="Olinguito" pageId="2" pageNumber="287" stateProvince="Antioquia">Olinguito</location>
sector;
<geoCoordinate id="6E851FA2FFC2FFC278B0951F2738FD58" box="[233,341,295,322]" degrees="5.519166" direction="north" orientation="latitude" pageId="2" pageNumber="287" precision="1" value="5.519166">5.519166</geoCoordinate>
,
<geoCoordinate id="6E851FA2FFC2FFC27939951F2789FD58" box="[352,484,295,322]" degrees="75.862185" direction="west" orientation="longitude" pageId="2" pageNumber="287" precision="1" value="-75.86218">-75.862185</geoCoordinate>
;
<quantity id="CC49D480FFC2FFC279B6951F242AFD58" box="[495,583,295,322]" metricMagnitude="3" metricUnit="m" metricValue="2.28" pageId="2" pageNumber="287" unit="m" value="2280.0">
<elevation id="809C9E56FFC2FFC279B6951F242AFD58" box="[495,583,295,322]" metricMagnitude="3" metricUnit="m" metricValue="2.28" pageId="2" pageNumber="287" unit="m" value="2280.0">2280 m</elevation>
</quantity>
;
<collectingMethod id="D2F00172FFC2FFC27A0B951F248CFD58" box="[594,737,295,322]" pageId="2" pageNumber="287">Malaise trap</collectingMethod>
;
<date id="7F0F5FA5FFC2FFC27AB5951F2538FD5B" box="[748,853,295,321]" pageId="2" pageNumber="287" value="2020-08">
<collectingDate id="6F4BA64DFFC2FFC27AB5951F2538FD5B" box="[748,853,295,321]" pageId="2" pageNumber="287" value="2020-08">viii.2020</collectingDate>
</date>
;
<collectorName id="A6441CB3FFC2FFC27B399510226FFD5B" box="[864,1026,295,322]" pageId="2" pageNumber="287">L. Mazariegos</collectorName>
and col. gp. (
<collectionCode id="6DA0E1A0FFC2FFC27CC3951F2280FD5B" box="[1178,1261,295,321]" pageId="2" pageNumber="287">CEUA</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="43AB2AEEFFC2FFC2789E9573255DFDCB" pageId="2" pageNumber="287" type="diagnosis">
<paragraph id="0B0E7965FFC2FFC2789E9573255DFDCB" blockId="2.[151,1436,259,897]" pageId="2" pageNumber="287">
<emphasis id="39C5A577FFC2FFC2789E9573272FFD7F" bold="true" box="[199,322,331,357]" pageId="2" pageNumber="287">Diagnosis.</emphasis>
Female.
<taxonomicName id="CCB102E6FFC2FFC279F295732458FD7F" authorityName="Santos &amp; Mazariegos" authorityYear="2024" box="[427,565,331,357]" class="Insecta" family="Aulacidae" genus="Aulacus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="287" phylum="Arthropoda" rank="species" species="gracielae" status="sp. nov.">
<emphasis id="39C5A577FFC2FFC279F295732458FD7F" box="[427,565,331,357]" italics="true" pageId="2" pageNumber="287">A. gracielae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22F6180CFFC2FFC27A64957424F8FD7C" box="[573,661,332,358]" pageId="2" pageNumber="287" rank="species">sp. nov.</taxonomicNameLabel>
is distinct from all other known species of
<taxonomicName id="CCB102E6FFC2FFC27CDA957322B0FD7F" authorityName="Jurine" authorityYear="1807" box="[1155,1245,331,357]" class="Insecta" family="Aulacidae" genus="Aulacus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="39C5A577FFC2FFC27CDA957322B0FD7F" box="[1155,1245,331,357]" italics="true" pageId="2" pageNumber="287">Aulacus</emphasis>
</taxonomicName>
according to the following combination of morphological characteristics: head black; mesosoma reddish-orange; all femurs and tibias black and tarsi mostly white; fore wings hyaline with a black apical spot and metasoma mostly black, with petiole reddish-orange; metanotum with transverse carinae.
</paragraph>
</subSubSection>
<subSubSection id="43AB2AEEFFC2FFC3789E95E327E5F8F0" lastPageId="3" lastPageNumber="288" pageId="2" pageNumber="287" type="description">
<paragraph id="0B0E7965FFC2FFC2789E95E32322FE9F" blockId="2.[151,1436,259,897]" pageId="2" pageNumber="287">
<emphasis id="39C5A577FFC2FFC2789E95E327D3FDEF" bold="true" box="[199,446,475,501]" pageId="2" pageNumber="287">Description. Female.</emphasis>
Body
<quantity id="CC49D480FFC2FFC27A5095E3241DFDEC" box="[521,624,475,502]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="2" pageNumber="287" unit="mm" value="10.3">10.3 mm</quantity>
; Fore wing
<quantity id="CC49D480FFC2FFC27AAC95E32523FDEC" box="[757,846,475,502]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="2" pageNumber="287" unit="mm" value="9.2">9.2 mm</quantity>
; Ovipositor
<quantity id="CC49D480FFC2FFC27B8195E32253FDEC" box="[984,1086,475,502]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="2" pageNumber="287" unit="mm" value="11.7">11.7 mm</quantity>
.
<emphasis id="39C5A577FFC2FFC27C1095E322E7FDEF" box="[1097,1162,475,501]" italics="true" pageId="2" pageNumber="287">Color</emphasis>
. Head black; mesosoma and pro, mid and hind coxae reddish orange, trochanters reddish, legs from trochantellus black, except mid-basitarsus whiteish, and hind three most proximal tarsomeres white; Fore wing hyaline, slightly amber at the proximal half and with black apex; stigma and veins black, more brownish at the hyaline area; metasoma predominantly black, with the T1 partially reddish orange at its anterior half; ovipositor sheath black with white band at its distal third.
</paragraph>
<paragraph id="0B0E7965FFC2FFC2789E96B724BFFEEB" blockId="2.[151,1436,259,897]" pageId="2" pageNumber="287">
<emphasis id="39C5A577FFC2FFC2789E96B72769FEB3" box="[199,260,655,681]" italics="true" pageId="2" pageNumber="287">Head</emphasis>
. Integument smooth and shining, without punctures on frons, with fine silvery pubescence, densest on clypeus, malar space, mandibular base and gena; lower interocular distance 0.7 × eye height (
<quantity id="CC49D480FFC2FFC27CF3968B2369FED7" box="[1194,1284,691,717]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="287" unit="mm" value="1.7">1.7 mm</quantity>
); malar space 0.3 × eye height; antennal length 3.2 × head width.
</paragraph>
<paragraph id="0B0E7965FFC2FFC2789E96C42252FF9B" blockId="2.[151,1436,259,897]" pageId="2" pageNumber="287">
<emphasis id="39C5A577FFC2FFC2789E96C4272AFF0F" box="[199,327,764,789]" italics="true" pageId="2" pageNumber="287">Mesosoma.</emphasis>
Integument with areas finely punctuate with whitish setae at the pronotum, scutellum, metanotum, propodeum, and mainly on pro-, meso- and metapleuron, and legs; polished areas at the meso- and metapleuron, and posteriorly on the pro-, mid- and hind coxa (
<figureCitation id="938A65E0FFC2FFC27AF7977B256AFF47" box="[686,775,835,861]" captionStart="FIGURE 2" captionStartId="2.[152,255,1929,1953]" captionTargetBox="[151,1436,921,1905]" captionTargetId="figure-309@2.[151,1436,921,1905]" captionTargetPageId="2" captionText="FIGURE 2: Aulacus gracielae Santos &amp; Mazariegos sp. n., female; A, lateral habitus; B, head in dorsal view; C, head in frontal view; D, head and mesosoma in lateral view; E, mesosoma in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13618462" httpUri="https://zenodo.org/record/13618462/files/figure.png" pageId="2" pageNumber="287">Fig. 2A</figureCitation>
); mesoscutum median and lateral lobes, and mesoscutellum transversely carinate; ventral portion of the mesopleuron and propodeum reticulate.
</paragraph>
<caption id="5FCE29EDFFC2FFC278C193B125E5FBDF" ID-DOI="http://doi.org/10.5281/zenodo.13618462" ID-Zenodo-Dep="13618462" httpUri="https://zenodo.org/record/13618462/files/figure.png" pageId="2" pageNumber="287" startId="2.[152,255,1929,1953]" targetBox="[151,1436,921,1905]" targetPageId="2" targetType="figure">
<paragraph id="0B0E7965FFC2FFC278C193B125E5FBDF" blockId="2.[151,1437,1929,1989]" pageId="2" pageNumber="287">
<emphasis id="39C5A577FFC2FFC278C193B12777FBBB" bold="true" box="[152,282,1929,1953]" pageId="2" pageNumber="287">FIGURE 2:</emphasis>
<taxonomicName id="CCB102E6FFC2FFC2794693B124DFFBBB" authority="Santos &amp; Mazariegos" authorityName="Santos &amp; Mazariegos" authorityYear="2024" box="[287,690,1929,1953]" class="Insecta" family="Aulacidae" genus="Aulacus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="287" phylum="Arthropoda" rank="species" species="gracielae" status="sp. nov.">
<emphasis id="39C5A577FFC2FFC2794693B127BBFBBB" box="[287,470,1929,1953]" italics="true" pageId="2" pageNumber="287">Aulacus gracielae</emphasis>
Santos &amp; Mazariegos
</taxonomicName>
<emphasis id="39C5A577FFC2FFC27AE193B1249DFBBB" bold="true" box="[696,752,1929,1953]" pageId="2" pageNumber="287">
<taxonomicNameLabel id="22F6180CFFC2FFC27AE193B1249DFBBB" box="[696,752,1929,1953]" pageId="2" pageNumber="287" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, female; A, lateral habitus; B, head in dorsal view; C, head in frontal view; D, head and mesosoma in lateral view; E, mesosoma in dorsal view.
</paragraph>
</caption>
<caption id="5FCE29EDFFC3FFC378C197EE2484F82C" ID-DOI="http://doi.org/10.5281/zenodo.13618464" ID-Zenodo-Dep="13618464" httpUri="https://zenodo.org/record/13618464/files/figure.png" pageId="3" pageNumber="288" startId="3.[152,255,982,1006]" targetBox="[151,1436,181,956]" targetPageId="3" targetType="figure">
<paragraph id="0B0E7965FFC3FFC378C197EE2484F82C" blockId="3.[151,1436,982,1078]" pageId="3" pageNumber="288">
<emphasis id="39C5A577FFC3FFC378C197EE2770FFF4" bold="true" box="[152,285,982,1006]" pageId="3" pageNumber="288">FIGURE 3:</emphasis>
Distribution map for the species of
<taxonomicName id="CCB102E6FFC3FFC37AC597EE2483FFF4" authorityName="Jurine" authorityYear="1807" box="[668,750,982,1006]" class="Insecta" family="Aulacidae" genus="Aulacus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="39C5A577FFC3FFC37AC597EE2483FFF4" box="[668,750,982,1006]" italics="true" pageId="3" pageNumber="288">Aulacus</emphasis>
</taxonomicName>
recorded in Colombia. The red star represents the location of the Mesenia-Paramillo nature reserve, and consequently the locality of the new record of
<taxonomicName id="CCB102E6FFC3FFC37C7897C2232BF808" authority="Smith" authorityName="Smith" authorityYear="2008" box="[1057,1350,1018,1042]" class="Insecta" family="Aulacidae" genus="Aulacus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="288" phylum="Arthropoda" rank="species" species="costaricensis">
<emphasis id="39C5A577FFC3FFC37C7897C2236DF808" box="[1057,1280,1018,1042]" italics="true" pageId="3" pageNumber="288">Aulacus costaricensis</emphasis>
Smith
</taxonomicName>
and the record of
<taxonomicName id="CCB102E6FFC3FFC378A3902624FFF82C" authority="Santos &amp; Mazariegos" authorityName="Santos &amp; Mazariegos" authorityYear="2024" box="[250,658,1054,1078]" class="Insecta" family="Aulacidae" genus="Aulacus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="288" phylum="Arthropoda" rank="species" species="gracielae" status="sp. nov.">
<emphasis id="39C5A577FFC3FFC378A3902627DFF82C" box="[250,434,1054,1078]" italics="true" pageId="3" pageNumber="288">Aulacus gracielae</emphasis>
Santos &amp; Mazariegos
</taxonomicName>
<taxonomicNameLabel id="22F6180CFFC3FFC37AC190262484F82C" box="[664,745,1054,1078]" pageId="3" pageNumber="288" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="0B0E7965FFC3FFC3789E90B12797F8DD" blockId="3.[151,1437,1160,1547]" pageId="3" pageNumber="288">
<emphasis id="39C5A577FFC3FFC3789E90B12751F8B8" box="[199,316,1161,1186]" italics="true" pageId="3" pageNumber="288">Metasoma</emphasis>
. Integument polished with tiny golden setae on T6, T7, and from S2 to S6; ovipositor sheath with subapical white band (
<figureCitation id="938A65E0FFC3FFC379CA90942780F8DD" box="[403,493,1196,1223]" captionStart="FIGURE 2" captionStartId="2.[152,255,1929,1953]" captionTargetBox="[151,1436,921,1905]" captionTargetId="figure-309@2.[151,1436,921,1905]" captionTargetPageId="2" captionText="FIGURE 2: Aulacus gracielae Santos &amp; Mazariegos sp. n., female; A, lateral habitus; B, head in dorsal view; C, head in frontal view; D, head and mesosoma in lateral view; E, mesosoma in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13618462" httpUri="https://zenodo.org/record/13618462/files/figure.png" pageId="3" pageNumber="288">Fig. 2A</figureCitation>
).
</paragraph>
<paragraph id="0B0E7965FFC3FFC3789E90E927E5F8F0" blockId="3.[151,1437,1160,1547]" box="[199,392,1232,1259]" pageId="3" pageNumber="288">
<emphasis id="39C5A577FFC3FFC3789E90E92766F8F1" bold="true" box="[199,267,1233,1259]" pageId="3" pageNumber="288">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="43AB2AEEFFC3FFC3789E90CD2716F914" box="[199,379,1268,1295]" pageId="3" pageNumber="288" type="biology_ecology">
<paragraph id="0B0E7965FFC3FFC3789E90CD2716F914" blockId="3.[151,1437,1160,1547]" box="[199,379,1268,1295]" pageId="3" pageNumber="288">
<emphasis id="39C5A577FFC3FFC3789E90CD276BF915" bold="true" box="[199,262,1269,1295]" pageId="3" pageNumber="288">Host.</emphasis>
Unknown
</paragraph>
</subSubSection>
<subSubSection id="43AB2AEEFFC3FFC3789E9120240CF928" box="[199,609,1304,1330]" pageId="3" pageNumber="288" type="distribution">
<paragraph id="0B0E7965FFC3FFC3789E9120240CF928" blockId="3.[151,1437,1160,1547]" box="[199,609,1304,1330]" pageId="3" pageNumber="288">
<emphasis id="39C5A577FFC3FFC3789E9120270EF928" bold="true" box="[199,355,1304,1330]" pageId="3" pageNumber="288">Distribution:</emphasis>
<collectingCountry id="73A639F5FFC3FFC37933912027B5F928" box="[362,472,1304,1330]" name="Colombia" pageId="3" pageNumber="288">Colombia</collectingCountry>
(
<collectingRegion id="C975B787FFC3FFC379BE91202436F928" box="[487,603,1304,1330]" country="Colombia" name="Antioquia" pageId="3" pageNumber="288">Antioquia</collectingRegion>
)
</paragraph>
</subSubSection>
<subSubSection id="43AB2AEEFFC3FFC3789E91052401FA10" pageId="3" pageNumber="288" type="etymology">
<paragraph id="0B0E7965FFC3FFC3789E91052401FA10" blockId="3.[151,1437,1160,1547]" pageId="3" pageNumber="288">
<emphasis id="39C5A577FFC3FFC3789E91052723F94D" bold="true" box="[199,334,1341,1367]" pageId="3" pageNumber="288">Etymology.</emphasis>
The epiphet
<taxonomicName id="CCB102E6FFC3FFC379B09105240DF94C" authorityName="Santos &amp; Mazariegos" authorityYear="2024" box="[489,608,1340,1367]" class="Insecta" family="Aulacidae" genus="Aulacus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="288" phylum="Arthropoda" rank="species" species="gracielae">
<emphasis id="39C5A577FFC3FFC379AF9104240DF94C" box="[502,608,1340,1366]" italics="true" pageId="3" pageNumber="288">gracielae</emphasis>
</taxonomicName>
” is in honour of the groundbreaking contributions and resilience of Dr. Graciela Hurtado de Mazariegos (1921-2020), OB/GYN and the second female doctor in Colombias history. She practised her profession for six decades until the age of 92, and by challenging preconceived notions, she paved the way for other women to enter the medical field. In a distinctive and well-deserved homage to her enduring legacy, we name this
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after her, highlighting the interconnectedness of various disciplines and honours the multifaceted impact of women in science.
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(synonymization);
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Head black; mesonotum reddish orange with propleuron and pronotum black; fore wing yellowish with a broad black band at center and black apex; metasoma reddish orange with apical 4 segments and ovipositor sheath black. Vertex with large and widely spaced punctures; mesonotum transversely carinate; notauli meeting on transscutal articulation; hind coxa shining, without carinae or punctures; ovipositor length 2.2 × fore wing length.
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<emphasis id="39C5A577FFC4FFC478CE94AF27ADFCAB" bold="true" box="[151,448,151,178]" pageId="4" pageNumber="289">
<emphasis id="39C5A577FFC4FFC478CE94AF2703FCA8" bold="true" box="[151,366,151,178]" italics="true" pageId="4" pageNumber="289">Aulacus cephalus</emphasis>
Smith
</emphasis>
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</paragraph>
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<taxonomicName id="CCB102E6FFC4FFC478CE94E42459FCEE" ID-CoL="JRHL" authority="Smith, 2005: 222 - 224" authorityName="Smith" authorityPageNumber="222 - 224" authorityYear="2005" box="[151,564,219,244]" class="Insecta" family="Aulacidae" genus="Aulacus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="289" phylum="Arthropoda" rank="species" species="cephalus">
<emphasis id="39C5A577FFC4FFC478CE94E42724FCEE" box="[151,329,220,244]" italics="true" pageId="4" pageNumber="289">Aulacus cephalus</emphasis>
<bibRefCitation id="6F200494FFC4FFC4790994E42459FCEE" author="Smith, D. R." box="[336,564,219,244]" pageId="4" pageNumber="289" pagination="217 - 253" refId="ref3686" refString="Smith, D. R. (2005) Aulacidae (Hymenoptera) of northern South America, emphasizing Colombia. Transactions of the American Entomological Society, 131, 217 - 253. [https: // www. jstor. org / stable / 25078885]" type="journal article" year="2005">Smith, 2005: 222224</bibRefCitation>
</taxonomicName>
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, Figs 712
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="43AB2AEEFFC4FFC478CE95192201FD9E" pageId="4" pageNumber="289" type="diagnosis">
<paragraph id="0B0E7965FFC4FFC478CE95192201FD9E" blockId="4.[151,1436,289,424]" pageId="4" pageNumber="289">
<emphasis id="39C5A577FFC4FFC478CE9519277FFD21" bold="true" box="[151,274,289,315]" pageId="4" pageNumber="289">Diagnosis.</emphasis>
Body black; fore wing hyaline with black apex; ovipositor sheath with a white band. Head elongate behind eyes; vertex with large widely spaced punctures; mesosoma densely rugosely punctate; hind coxa shining, without transverse carinae or punctures; ovipositor length subequal to fore wing length.
</paragraph>
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<subSubSection id="43AB2AEEFFC4FFC4789E95B52429FDB2" box="[199,580,397,424]" pageId="4" pageNumber="289" type="distribution">
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<emphasis id="39C5A577FFC4FFC4789E95B5270CFDBD" bold="true" box="[199,353,397,423]" pageId="4" pageNumber="289">Distribution.</emphasis>
<collectingCountry id="73A639F5FFC4FFC4793E95B627B8FDB2" box="[359,469,398,424]" name="Colombia" pageId="4" pageNumber="289">Colombia</collectingCountry>
(
<collectingRegion id="C975B787FFC4FFC479BC95B62451FDB2" box="[485,572,398,424]" country="Colombia" name="Bolivar" pageId="4" pageNumber="289">Bolívar</collectingRegion>
)
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View file

@ -0,0 +1,131 @@
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<taxonomicName id="CCB102E6FFC4FFC478CE95C12782FE09" authority="Smith" authorityName="Smith" authorityYear="2008" box="[151,495,505,532]" class="Insecta" family="Aulacidae" genus="Aulacus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="289" phylum="Arthropoda" rank="species" species="costaricensis">
<emphasis id="39C5A577FFC4FFC478CE95C12782FE09" bold="true" box="[151,495,505,532]" pageId="4" pageNumber="289">
<emphasis id="39C5A577FFC4FFC478CE95C127F0FE0E" bold="true" box="[151,413,505,532]" italics="true" pageId="4" pageNumber="289">Aulacus costaricensis</emphasis>
Smith
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<bibRefCitation id="6F200494FFC4FFC4792396062428FE4C" author="Smith, D. R." box="[378,581,574,598]" pageId="4" pageNumber="289" pagination="267 - 355" refId="ref3734" refString="Smith, D. R. (2008) Aulacidae of the southwestern United States, Mexico, and Central America. Beitrage zur Entomologie, 58, 267 - 355. https: // doi. org / 10.21248 / contrib. entomol. 58.2.267 - 355" type="journal article" year="2008">Smith, 2008: 1618</bibRefCitation>
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, Figs 1821
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<emphasis id="39C5A577FFC4FFC478CE96BC277FFE84" bold="true" box="[151,274,644,670]" pageId="4" pageNumber="289">Diagnosis.</emphasis>
Body black, only tarsi mostly whitish; fore wing hyaline with apex black. Head smooth and shining; vertex with inconspicuous fine punctures; mesoscutum shining, with coarse transverse carinae which are stronger posteriorly, but few punctures; hind coxa smooth and shining.
</paragraph>
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<emphasis id="39C5A577FFC4FFC4789E96C827D8FF10" bold="true" box="[199,437,752,778]" pageId="4" pageNumber="289">Examined material.</emphasis>
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,
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,
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:
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;
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sector;
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;
<collectingMethod id="D2F00172FFC4FFC47A5B972C24FDFF35" box="[514,656,788,815]" pageId="4" pageNumber="289">Malaise trap</collectingMethod>
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and col. gp. (
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.
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<emphasis id="39C5A577FFC4FFC4789E9700270DFF49" bold="true" box="[199,352,824,851]" pageId="4" pageNumber="289">Distribution.</emphasis>
<collectingCountry id="73A639F5FFC4FFC4793E970027B8FF48" box="[359,469,824,850]" name="Colombia" pageId="4" pageNumber="289">Colombia</collectingCountry>
(
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<collectingCountry id="73A639F5FFC4FFC47A3797012485FF48" box="[622,744,824,851]" name="Costa Rica" pageId="4" pageNumber="289">Costa Rica</collectingCountry>
(Puntaneras)
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<taxonomicName id="CCB102E6FFC4FFC478CE979C27D9FFA4" ID-CoL="JRK5" authority="Smith" authorityName="Smith" authorityYear="2005" box="[151,436,932,959]" class="Insecta" family="Aulacidae" genus="Aulacus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="289" phylum="Arthropoda" rank="species" species="ochreus">
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<emphasis id="39C5A577FFC4FFC478CE979C270FFFA5" bold="true" box="[151,354,932,959]" italics="true" pageId="4" pageNumber="289">Aulacus ochreus</emphasis>
Smith
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, Figs 1318;
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, Figs 510.
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<emphasis id="39C5A577FFC4FFC478CE9017277FF853" bold="true" box="[151,274,1071,1097]" pageId="4" pageNumber="289">Diagnosis.</emphasis>
Mostly yellow; fore wing hyaline with apex black. Head in dorsal view narrow behind eyes; eye close to posterior margin of head in lateral view; head and mesosoma with large and widely spaced punctures; notauli V-shaped meeting at transscutal articulation. Hind coxa shining, without carinae or punctures; ovipositor 2.2 × fore wing length.
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<emphasis id="39C5A577FFC4FFC4789E9087270CF8C3" bold="true" box="[199,353,1215,1241]" pageId="4" pageNumber="289">Distribution.</emphasis>
<collectingCountry id="73A639F5FFC4FFC4793E908727B8F8C3" box="[359,469,1215,1241]" name="Colombia" pageId="4" pageNumber="289">Colombia</collectingCountry>
(
<collectingRegion id="C975B787FFC4FFC479BC90872405F8C3" box="[485,616,1215,1241]" country="Colombia" name="Magdalena" pageId="4" pageNumber="289">Magdalena</collectingRegion>
);
<collectingCountry id="73A639F5FFC4FFC47A2290872498F8C3" box="[635,757,1215,1241]" name="Costa Rica" pageId="4" pageNumber="289">Costa Rica</collectingCountry>
(
<collectingRegion id="C975B787FFC4FFC47B5C908725E5F8C3" box="[773,904,1215,1241]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="289">Guanacaste</collectingRegion>
)
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<emphasis id="2DD7EA89FFA10E18FF22FD88CD71FDEF" bold="true" box="[151,276,597,624]" italics="true" pageId="11" pageNumber="392">Ephemera</emphasis>
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<figureCitation id="87982A1EFFA10E18FF22FDA6CD5DFD0A" box="[151,312,635,661]" captionStart-0="FIGURES 3134" captionStart-1="FIGURES 3537" captionStartId-0="12.[151,261,1943,1967]" captionStartId-1="13.[151,264,1943,1967]" captionTargetBox-0="[256,1331,181,1919]" captionTargetBox-1="[286,1301,181,1919]" captionTargetId-0="figure-18@12.[256,1331,181,1919]" captionTargetId-1="figure-16@13.[286,1301,181,1919]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionText-0="FIGURES 3134. Egg morphology of Ephemera (Sinephemera) shengmi: 3132, general shape and arrangement of the micropyles; 33, proximal part of micropylar canal on chorion surface; 34, thickness on the layers forming the eggshell and chorion. AL-Ex—complex adhesive-extrachorion layer; AL—adhesive layer; C—chorion; MO—micropylar opening; mp—micropyle." captionText-1="FIGURES 3537. Egg morphology of Ephemera (Sinephemera) shengmi: 3536, egg surface without complex extrachorionadhesive layer; 37, proximal part of micropylar canal on chorion surface. AL-Ex—complex adhesive-extrachorion layer; CS—chorion surface; MO—micropylar opening; mp—micropyle." pageId="11" pageNumber="392">Figures 3137</figureCitation>
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,
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, Przhevalsky spit,
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</subSubSection>
<subSubSection id="57B96510FFA10E18FF72FCE8CD0DFA8F" pageId="11" pageNumber="392" type="distribution">
<paragraph id="1F1C369BFFA10E18FF72FCE8CF63FCD0" blockId="11.[151,1437,709,1856]" box="[199,774,821,848]" pageId="11" pageNumber="392">
<emphasis id="2DD7EA89FFA10E18FF72FCE8CD3FFCCF" bold="true" box="[199,346,821,848]" italics="true" pageId="11" pageNumber="392">Distribution.</emphasis>
South of the Russian Far East,
<collectingCountry id="67B4760BFFA10E18FD0EFCE8CF67FCD0" box="[699,770,821,847]" name="China" pageId="11" pageNumber="392">China</collectingCountry>
.
</paragraph>
<paragraph id="1F1C369BFFA10E18FF72FC86CE53FC7A" blockId="11.[151,1437,709,1856]" pageId="11" pageNumber="392">
The egg is oval (
<figureCitation id="87982A1EFFA10E18FE36FC86CE60FCEA" box="[387,517,859,885]" captionStart="FIGURES 3134" captionStartId="12.[151,261,1943,1967]" captionTargetBox="[256,1331,181,1919]" captionTargetId="figure-18@12.[256,1331,181,1919]" captionTargetPageId="12" captionText="FIGURES 3134. Egg morphology of Ephemera (Sinephemera) shengmi: 3132, general shape and arrangement of the micropyles; 33, proximal part of micropylar canal on chorion surface; 34, thickness on the layers forming the eggshell and chorion. AL-Ex—complex adhesive-extrachorion layer; AL—adhesive layer; C—chorion; MO—micropylar opening; mp—micropyle." pageId="11" pageNumber="392">Figs 3132</figureCitation>
,
<figureCitation id="87982A1EFFA10E18FDA5FC87CE3EFCEA" box="[528,603,858,885]" captionStart="FIGURES 3537" captionStartId="13.[151,264,1943,1967]" captionTargetBox="[286,1301,181,1919]" captionTargetId="figure-16@13.[286,1301,181,1919]" captionTargetPageId="13" captionText="FIGURES 3537. Egg morphology of Ephemera (Sinephemera) shengmi: 3536, egg surface without complex extrachorionadhesive layer; 37, proximal part of micropylar canal on chorion surface. AL-Ex—complex adhesive-extrachorion layer; CS—chorion surface; MO—micropylar opening; mp—micropyle." pageId="11" pageNumber="392">3536</figureCitation>
). Dimensions: 224.0255.0 µm in length (239.9 µm) and 130.0142.0 µm in width (136.3 µm). The extrachorion-adhesive layer is thin, 0.51.2 μm; therefore, the surface of the adhesive layer partially follows the sculpture of the chorion (
<figureCitation id="87982A1EFFA10E18FD5FFC78CF0BFC20" box="[746,878,933,959]" captionStart="FIGURES 3134" captionStartId="12.[151,261,1943,1967]" captionTargetBox="[256,1331,181,1919]" captionTargetId="figure-18@12.[256,1331,181,1919]" captionTargetPageId="12" captionText="FIGURES 3134. Egg morphology of Ephemera (Sinephemera) shengmi: 3132, general shape and arrangement of the micropyles; 33, proximal part of micropylar canal on chorion surface; 34, thickness on the layers forming the eggshell and chorion. AL-Ex—complex adhesive-extrachorion layer; AL—adhesive layer; C—chorion; MO—micropylar opening; mp—micropyle." pageId="11" pageNumber="392">Figs 3132</figureCitation>
,
<figureCitation id="87982A1EFFA10E18FCCEFC78CFFFFC5F" box="[891,922,933,960]" captionStart="FIGURES 3537" captionStartId="13.[151,264,1943,1967]" captionTargetBox="[286,1301,181,1919]" captionTargetId="figure-16@13.[286,1301,181,1919]" captionTargetPageId="13" captionText="FIGURES 3537. Egg morphology of Ephemera (Sinephemera) shengmi: 3536, egg surface without complex extrachorionadhesive layer; 37, proximal part of micropylar canal on chorion surface. AL-Ex—complex adhesive-extrachorion layer; CS—chorion surface; MO—micropylar opening; mp—micropyle." pageId="11" pageNumber="392">35</figureCitation>
). In the equatorial area, there are one to two micropyles per egg (
<figureCitation id="87982A1EFFA10E18FEC8FC16CD9BFC7A" box="[381,510,971,997]" captionStart="FIGURES 3134" captionStartId="12.[151,261,1943,1967]" captionTargetBox="[256,1331,181,1919]" captionTargetId="figure-18@12.[256,1331,181,1919]" captionTargetPageId="12" captionText="FIGURES 3134. Egg morphology of Ephemera (Sinephemera) shengmi: 3132, general shape and arrangement of the micropyles; 33, proximal part of micropylar canal on chorion surface; 34, thickness on the layers forming the eggshell and chorion. AL-Ex—complex adhesive-extrachorion layer; AL—adhesive layer; C—chorion; MO—micropylar opening; mp—micropyle." pageId="11" pageNumber="392">Figs 3132</figureCitation>
,
<figureCitation id="87982A1EFFA10E18FDBCFC17CE4DFC7A" box="[521,552,970,997]" captionStart="FIGURES 3537" captionStartId="13.[151,264,1943,1967]" captionTargetBox="[286,1301,181,1919]" captionTargetId="figure-16@13.[286,1301,181,1919]" captionTargetPageId="13" captionText="FIGURES 3537. Egg morphology of Ephemera (Sinephemera) shengmi: 3536, egg surface without complex extrachorionadhesive layer; 37, proximal part of micropylar canal on chorion surface. AL-Ex—complex adhesive-extrachorion layer; CS—chorion surface; MO—micropylar opening; mp—micropyle." pageId="11" pageNumber="392">35</figureCitation>
).
</paragraph>
<paragraph id="1F1C369BFFA10E18FF72FC2CC965FB5A" blockId="11.[151,1437,709,1856]" pageId="11" pageNumber="392">
Eggs are characterized by a linear-type micropyle (
<bibRefCitation id="7B324B6AFFA10E18FCBCFC2DC84DFB95" author="Koss, R. W. &amp; Edmunds, G. F. Jr." box="[777,1064,1008,1034]" pageId="11" pageNumber="392" pagination="267 - 349" refId="ref7161" refString="Koss, R. W. &amp; Edmunds, G. F. Jr. (1974) Ephemeroptera eggs and their contribution to phylogenetic studies of the order. Zoological Journal of the Linnean Society, 55, 267 - 349. https: // doi. org / 10.1111 / j. 1096 - 3642.1974. tb 01648. x" type="journal article" year="1974">Koss and Edmunds 1974</bibRefCitation>
). In this
<typeStatus id="C0188839FFA10E18FB39FC2CC8D9FB94" box="[1164,1212,1009,1035]" pageId="11" pageNumber="392">type</typeStatus>
of micropyle, there is no sperm guide, and the micropylar canal is relatively short and protrudes over both the adhesive and chorion surfaces (
<figureCitation id="87982A1EFFA10E18FEB0FBE6CDE9FBCA" box="[261,396,1083,1109]" captionStart="FIGURES 3134" captionStartId="12.[151,261,1943,1967]" captionTargetBox="[256,1331,181,1919]" captionTargetId="figure-18@12.[256,1331,181,1919]" captionTargetPageId="12" captionText="FIGURES 3134. Egg morphology of Ephemera (Sinephemera) shengmi: 3132, general shape and arrangement of the micropyles; 33, proximal part of micropylar canal on chorion surface; 34, thickness on the layers forming the eggshell and chorion. AL-Ex—complex adhesive-extrachorion layer; AL—adhesive layer; C—chorion; MO—micropylar opening; mp—micropyle." pageId="11" pageNumber="392">Figs 3132</figureCitation>
). The micropylar canal, tunnel-shaped with thick walls, is relatively short, 814 μm long and 710 μm wide (
<figureCitation id="87982A1EFFA10E18FECCFBBDCDA8FBE5" box="[377,461,1120,1146]" captionStart="FIGURES 3134" captionStartId="12.[151,261,1943,1967]" captionTargetBox="[256,1331,181,1919]" captionTargetId="figure-18@12.[256,1331,181,1919]" captionTargetPageId="12" captionText="FIGURES 3134. Egg morphology of Ephemera (Sinephemera) shengmi: 3132, general shape and arrangement of the micropyles; 33, proximal part of micropylar canal on chorion surface; 34, thickness on the layers forming the eggshell and chorion. AL-Ex—complex adhesive-extrachorion layer; AL—adhesive layer; C—chorion; MO—micropylar opening; mp—micropyle." pageId="11" pageNumber="392">Fig. 33</figureCitation>
). The micropylar opening is nearly round, 3.5 μm wide and 3.0 μm high (
<figureCitation id="87982A1EFFA10E18FAB9FBBDC906FBE5" box="[1292,1379,1120,1146]" captionStart="FIGURES 3134" captionStartId="12.[151,261,1943,1967]" captionTargetBox="[256,1331,181,1919]" captionTargetId="figure-18@12.[256,1331,181,1919]" captionTargetPageId="12" captionText="FIGURES 3134. Egg morphology of Ephemera (Sinephemera) shengmi: 3132, general shape and arrangement of the micropyles; 33, proximal part of micropylar canal on chorion surface; 34, thickness on the layers forming the eggshell and chorion. AL-Ex—complex adhesive-extrachorion layer; AL—adhesive layer; C—chorion; MO—micropylar opening; mp—micropyle." pageId="11" pageNumber="392">Figs 33</figureCitation>
,
<figureCitation id="87982A1EFFA10E18FAC5FBBDC9F5FBE5" box="[1392,1424,1120,1146]" captionStart="FIGURES 3537" captionStartId="13.[151,264,1943,1967]" captionTargetBox="[286,1301,181,1919]" captionTargetId="figure-16@13.[286,1301,181,1919]" captionTargetPageId="13" captionText="FIGURES 3537. Egg morphology of Ephemera (Sinephemera) shengmi: 3536, egg surface without complex extrachorionadhesive layer; 37, proximal part of micropylar canal on chorion surface. AL-Ex—complex adhesive-extrachorion layer; CS—chorion surface; MO—micropylar opening; mp—micropyle." pageId="11" pageNumber="392">37</figureCitation>
). The sculpture of the chorion consists of discontinuous long and short ridged, curved, and broken filaments, the distribution and arrangement of which regularly cover the entire surface of the chorion (
<figureCitation id="87982A1EFFA10E18FBC5FB76C896FB5A" box="[1136,1267,1194,1221]" captionStart="FIGURES 3537" captionStartId="13.[151,264,1943,1967]" captionTargetBox="[286,1301,181,1919]" captionTargetId="figure-16@13.[286,1301,181,1919]" captionTargetPageId="13" captionText="FIGURES 3537. Egg morphology of Ephemera (Sinephemera) shengmi: 3536, egg surface without complex extrachorionadhesive layer; 37, proximal part of micropylar canal on chorion surface. AL-Ex—complex adhesive-extrachorion layer; CS—chorion surface; MO—micropylar opening; mp—micropyle." pageId="11" pageNumber="392">Figs 3537</figureCitation>
).
</paragraph>
<paragraph id="1F1C369BFFA10E18FF72FB12CEBBFB75" blockId="11.[151,1437,709,1856]" box="[199,734,1231,1259]" pageId="11" pageNumber="392">
<taxonomicName id="D8A34D18FFA10E18FF72FB12CEBBFB75" authority="Eaton, 1892" authorityName="Eaton" authorityYear="1892" box="[199,734,1231,1259]" class="Insecta" family="Ephemeridae" genus="Ephemera" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="392" phylum="Arthropoda" rank="species" species="strigata" subGenus="Sinephemera">
<emphasis id="2DD7EA89FFA10E18FF72FB12CE2AFB75" bold="true" box="[199,591,1231,1258]" pageId="11" pageNumber="392">
<emphasis id="2DD7EA89FFA10E18FF72FB12CD5AFB75" bold="true" box="[199,319,1231,1258]" italics="true" pageId="11" pageNumber="392">Ephemera</emphasis>
(
<emphasis id="2DD7EA89FFA10E18FEFAFB12CD82FB75" bold="true" box="[335,487,1231,1258]" italics="true" pageId="11" pageNumber="392">Sinephemera</emphasis>
)
<emphasis id="2DD7EA89FFA10E18FE4DFB0DCE2AFB75" bold="true" box="[504,591,1232,1258]" italics="true" pageId="11" pageNumber="392">strigata</emphasis>
</emphasis>
<bibRefCitation id="7B324B6AFFA10E18FDE3FB0CCEBBFB75" author="Eaton, A. E." box="[598,734,1232,1259]" pageId="11" pageNumber="392" pagination="302 - 303" refId="ref6967" refString="Eaton, A. E. (1892) On two new and some other Japanese species of Ephemeridae. Entomologist's Monthly Magazine, Series 2, 3 (28), 302 - 303." type="journal article" year="1892">Eaton, 1892</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="1F1C369BFFA10E18FF72FB28CD0DFA8F" blockId="11.[151,1437,709,1856]" box="[199,360,1269,1296]" pageId="11" pageNumber="392">
<figureCitation id="87982A1EFFA10E18FF72FB28CD0DFA8F" box="[199,360,1269,1296]" captionStart-0="FIGURES 3841" captionStart-1="FIGURES 4245" captionStartId-0="15.[151,264,1943,1967]" captionStartId-1="16.[151,264,1943,1967]" captionTargetBox-0="[284,1303,181,1919]" captionTargetBox-1="[289,1299,181,1919]" captionTargetId-0="figure-16@15.[284,1303,181,1919]" captionTargetId-1="figure-18@16.[289,1299,181,1919]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="FIGURES 3841. Egg morphology of Ephemera (Sinephemera) strigata: 3839, general shape and arrangement of the micropyles; 40, micropyle in the complex extrachorion-adhesive layer; 41, chorion thickness. AL-Ex—complex adhesive-extrachorion layer; C—chorion; CM—canal micropylar; MO—micropylar opening; SG—sperm guide." captionText-1="FIGURES 4245. Egg morphology of Ephemera (Sinephemera) strigata: 42, remains of the adhesive layer on the egg surface; 4344, proximal part of micropylar canal on chorion surface; 45, chorionic sculpturing. AL-Ex—complex adhesive-extrachorion layer; CS—chorion surface; CM—canal micropylar; MO—micropylar opening." pageId="11" pageNumber="392">Figures 3845</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="57B96510FFA10E18FF72FAC7CF91FAE0" pageId="11" pageNumber="392" type="materials_examined">
<paragraph id="1F1C369BFFA10E18FF72FAC7CF91FAE0" blockId="11.[151,1437,709,1856]" pageId="11" pageNumber="392">
<emphasis id="2DD7EA89FFA10E18FF72FAC7CDCDFAAB" bold="true" box="[199,424,1306,1332]" pageId="11" pageNumber="392">Material examined</emphasis>
.
<materialsCitation id="AFCB3CC6FFA10E18FE01FAC6CF95FAE0" collectingDate="2003-10-06" collectingDateMax="2021-06-24" collectingDateMin="2003-10-06" collectionCode="T" country="Russia" county="Khasansky district" location="Fish Hatchery" municipality="Barabashevka River" pageId="11" pageNumber="392" specimenCount="8" specimenCount-female="8" stateProvince="Primorskiy">
<collectingCountry id="67B4760BFFA10E18FE01FAC6CE66FAAA" box="[436,515,1307,1333]" name="Russia" pageId="11" pageNumber="392">Russia</collectingCountry>
:
<collectingRegion id="DD67F879FFA10E18FDB9FAC6CEDFFAAA" box="[524,698,1307,1333]" country="Russia" name="Primorskiy" pageId="11" pageNumber="392">Primorsky Krai</collectingRegion>
,
<collectingCounty id="F67D4E17FFA10E18FD76FAC6CFF3FAAA" box="[707,918,1307,1333]" pageId="11" pageNumber="392">Khasansky district</collectingCounty>
:
<collectingMunicipality id="FF78ACE1FFA10E18FC2BFAC6C8E4FAAA" box="[926,1153,1307,1333]" pageId="11" pageNumber="392">Barabashevka River</collectingMunicipality>
, below the
<location id="1A7C6040FFA10E18FB4EFAC6C9FDFAAA" LSID="urn:lsid:plazi:treatment:970A878DFFA10E18FF22FD88CF36F8DF:1A7C6040FFA10E18FB4EFAC6C9FDFAAA" box="[1275,1432,1307,1333]" country="Russia" county="Khasansky district" municipality="Barabashevka River" name="Fish Hatchery" pageId="11" pageNumber="392" stateProvince="Primorskiy">Fish Hatchery</location>
,
<specimenCount id="09A5FD12FFA10E18FF22FA9DCCDFFAC5" box="[151,186,1344,1370]" count="4" pageId="11" pageNumber="392" type="female">4♀</specimenCount>
adults,
<date id="6B1D105BFFA10E18FEA5FA9DCDF5FAC5" box="[272,400,1344,1370]" pageId="11" pageNumber="392" value="2003-10-06">
<collectingDate id="7B59E9B3FFA10E18FEA5FA9DCDF5FAC5" box="[272,400,1344,1370]" pageId="11" pageNumber="392" value="2003-10-06">10.06.2003</collectingDate>
</date>
,
<collectionCode id="79B2AE5EFFA10E18FE2FFA9CCDC9FAC4" box="[410,428,1345,1371]" name="Tavera, Department of Geology and Geophysics" pageId="11" pageNumber="392">T</collectionCode>
. Tiunova;
<location id="1A7C6040FFA10E18FDAAFA9DCF67FAC5" LSID="urn:lsid:plazi:treatment:970A878DFFA10E18FF22FD88CF36F8DF:1A7C6040FFA10E18FDAAFA9DCF67FAC5" box="[543,770,1344,1370]" country="Russia" county="Khasansky district" municipality="Barabashevka River" name="Barabashevka River" pageId="11" pageNumber="392" stateProvince="Primorskiy">Barabashevka River</location>
, above the Fish Hatchery,
<date id="6B1D105BFFA10E18FB90FA9DC8C3FAC5" box="[1061,1190,1344,1370]" pageId="11" pageNumber="392" value="2021-06-24">
<collectingDate id="7B59E9B3FFA10E18FB90FA9DC8C3FAC5" box="[1061,1190,1344,1370]" pageId="11" pageNumber="392" value="2021-06-24">24.06.2021</collectingDate>
</date>
,
<specimenCount id="09A5FD12FFA10E18FB05FA9DC8B7FAC5" box="[1200,1234,1344,1370]" count="1" pageId="11" pageNumber="392" type="female">1♀</specimenCount>
adult,
<collectionCode id="79B2AE5EFFA10E18FAA8FA9CC94AFAC4" box="[1309,1327,1345,1371]" name="Tavera, Department of Geology and Geophysics" pageId="11" pageNumber="392">T</collectionCode>
. Tiunova;
<location id="1A7C6040FFA10E18FF22FAB8CD02FAE0" LSID="urn:lsid:plazi:treatment:970A878DFFA10E18FF22FD88CF36F8DF:1A7C6040FFA10E18FF22FAB8CD02FAE0" box="[151,359,1381,1407]" country="Russia" county="Khasansky district" municipality="Barabashevka River" name="Ryazanovka River" pageId="11" pageNumber="392" stateProvince="Primorskiy">Ryazanovka River</location>
, below the Okhotbaza,
<date id="6B1D105BFFA10E18FDDBFAB8CE8BFAE0" box="[622,750,1381,1407]" pageId="11" pageNumber="392" value="2003-11-06">
<collectingDate id="7B59E9B3FFA10E18FDDBFAB8CE8BFAE0" box="[622,750,1381,1407]" pageId="11" pageNumber="392" value="2003-11-06">11.06.2003</collectingDate>
</date>
,
<specimenCount id="09A5FD12FFA10E18FD4CFAB8CF79FAE0" box="[761,796,1381,1407]" count="3" pageId="11" pageNumber="392" type="female">3♀</specimenCount>
adults,
<collectionCode id="79B2AE5EFFA10E18FCC1FABBCFE3FA1F" box="[884,902,1382,1408]" name="Tavera, Department of Geology and Geophysics" pageId="11" pageNumber="392">T</collectionCode>
. Tiunova
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="57B96510FFA10E18FF72FA57CF36F8DF" pageId="11" pageNumber="392" type="distribution">
<paragraph id="1F1C369BFFA10E18FF72FA57C87DFA3A" blockId="11.[151,1437,709,1856]" box="[199,1048,1418,1445]" pageId="11" pageNumber="392">
<emphasis id="2DD7EA89FFA10E18FF72FA57CD3FFA3A" bold="true" box="[199,346,1418,1445]" italics="true" pageId="11" pageNumber="392">Distribution.</emphasis>
East Siberia, Far East
<collectingCountry id="67B4760BFFA10E18FDECFA56CEC2FA3A" box="[601,679,1419,1445]" name="Russia" pageId="11" pageNumber="392">Russia</collectingCountry>
,
<collectingCountry id="67B4760BFFA10E18FD07FA56CE90FA3A" box="[690,757,1419,1445]" name="Japan" pageId="11" pageNumber="392">Japan</collectingCountry>
,
<collectingCountry id="67B4760BFFA10E18FCB5FA56CF15FA3A" box="[768,880,1419,1445]" name="Mongolia" pageId="11" pageNumber="392">Mongolia</collectingCountry>
,
<collectingCountry id="67B4760BFFA10E18FCCEFA56CFA7FA3A" box="[891,962,1419,1445]" name="South Korea" pageId="11" pageNumber="392">Korea</collectingCountry>
,
<collectingCountry id="67B4760BFFA10E18FC78FA56C871FA3A" box="[973,1044,1419,1445]" name="China" pageId="11" pageNumber="392">China</collectingCountry>
.
</paragraph>
<paragraph id="1F1C369BFFA10E18FF72FA6DC889F98A" blockId="11.[151,1437,709,1856]" pageId="11" pageNumber="392">
The egg has been described by
<bibRefCitation id="7B324B6AFFA10E18FD9DFA6DCEB5FA55" author="Okazaki, H." box="[552,720,1456,1482]" pageId="11" pageNumber="392" pagination="8 - 10" refId="ref7347" refString="Okazaki, H. (1981) On the eggs of Japanese mayflies (1). Biology of Inland Waters, 2, 8 - 10." type="journal article" year="1981">Okazaki (1981</bibRefCitation>
, p. 9: fig. 6; 1984, p. 21: fig. 15) and
<bibRefCitation id="7B324B6AFFA10E18FBC2FA6DC919FA55" author="Tojo, K. &amp; Machida, R." box="[1143,1404,1456,1482]" pageId="11" pageNumber="392" pagination="573 - 579" refId="ref7457" refString="Tojo, K. &amp; Machida, R. (1998) Egg Structures of Japanes Ephemeridae species (Ephemeroptera). Entomological Science, 1 (4), 573 - 579." type="journal article" year="1998">Tojo &amp; Machida (1998</bibRefCitation>
, p. 575: fig. 2). The authors write that the eggs of
<taxonomicName id="D8A34D18FFA10E18FD15FA0BCF73FA70" box="[672,790,1494,1519]" class="Insecta" family="Ephemeridae" genus="Ephemera" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="392" phylum="Arthropoda" rank="species" species="strigata">
<emphasis id="2DD7EA89FFA10E18FD15FA0BCF73FA70" box="[672,790,1494,1519]" italics="true" pageId="11" pageNumber="392">E. strigata</emphasis>
</taxonomicName>
and
<taxonomicName id="D8A34D18FFA10E18FCF8FA0BCFABFA70" box="[845,974,1494,1519]" class="Insecta" family="Ephemeridae" genus="Ephemera" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="392" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="2DD7EA89FFA10E18FCF8FA0BCFABFA70" box="[845,974,1494,1519]" italics="true" pageId="11" pageNumber="392">E. japonica</emphasis>
</taxonomicName>
are very similar in egg size and shape, in the thickness of the adhesive layer, in the structure of the chorion, and in the absence of micropyle.
</paragraph>
<paragraph id="1F1C369BFFA10E18FF72F9FDCF36F8DF" blockId="11.[151,1437,709,1856]" pageId="11" pageNumber="392">
According to our data, the egg is oval (
<figureCitation id="87982A1EFFA10E18FD3FF9FDCF6BF9A5" box="[650,782,1568,1594]" captionStart="FIGURES 3841" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[284,1303,181,1919]" captionTargetId="figure-16@15.[284,1303,181,1919]" captionTargetPageId="15" captionText="FIGURES 3841. Egg morphology of Ephemera (Sinephemera) strigata: 3839, general shape and arrangement of the micropyles; 40, micropyle in the complex extrachorion-adhesive layer; 41, chorion thickness. AL-Ex—complex adhesive-extrachorion layer; C—chorion; CM—canal micropylar; MO—micropylar opening; SG—sperm guide." pageId="11" pageNumber="392">Figs 3839</figureCitation>
). Dimensions: 197.0239.0 µm in length (215.2 µm) and 116.0139.0 µm in width (126.6 µm). The extrachorion-adhesive layer covering the egg is presented as a thin membrane (
<figureCitation id="87982A1EFFA10E18FEA9F9B6CD0AF91A" box="[284,367,1643,1669]" captionStart="FIGURES 4245" captionStartId="16.[151,264,1943,1967]" captionTargetBox="[289,1299,181,1919]" captionTargetId="figure-18@16.[289,1299,181,1919]" captionTargetPageId="16" captionText="FIGURES 4245. Egg morphology of Ephemera (Sinephemera) strigata: 42, remains of the adhesive layer on the egg surface; 4344, proximal part of micropylar canal on chorion surface; 45, chorionic sculpturing. AL-Ex—complex adhesive-extrachorion layer; CS—chorion surface; CM—canal micropylar; MO—micropylar opening." pageId="11" pageNumber="392">Fig. 42</figureCitation>
). The adhesive layer is almost smooth (
<figureCitation id="87982A1EFFA10E18FC97F9B6CFAEF91A" box="[802,971,1643,1669]" captionStart="FIGURES 3841" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[284,1303,181,1919]" captionTargetId="figure-16@15.[284,1303,181,1919]" captionTargetPageId="15" captionText="FIGURES 3841. Egg morphology of Ephemera (Sinephemera) strigata: 3839, general shape and arrangement of the micropyles; 40, micropyle in the complex extrachorion-adhesive layer; 41, chorion thickness. AL-Ex—complex adhesive-extrachorion layer; C—chorion; CM—canal micropylar; MO—micropylar opening; SG—sperm guide." pageId="11" pageNumber="392">Figs 3839, 41</figureCitation>
). There are one to two micropyles per egg in the equatorial area (
<figureCitation id="87982A1EFFA10E18FE27F94DCE70F935" box="[402,533,1680,1706]" captionStart="FIGURES 3841" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[284,1303,181,1919]" captionTargetId="figure-16@15.[284,1303,181,1919]" captionTargetPageId="15" captionText="FIGURES 3841. Egg morphology of Ephemera (Sinephemera) strigata: 3839, general shape and arrangement of the micropyles; 40, micropyle in the complex extrachorion-adhesive layer; 41, chorion thickness. AL-Ex—complex adhesive-extrachorion layer; C—chorion; CM—canal micropylar; MO—micropylar opening; SG—sperm guide." pageId="11" pageNumber="392">Figs 3839</figureCitation>
). The micropyle is of the “tagenoform
<typeStatus id="C0188839FFA10E18FC7CF94DCF9BF935" box="[969,1022,1680,1706]" pageId="11" pageNumber="392">type</typeStatus>
,” with a drop-shaped, poorly defined sperm guide, 1418 μm long and 1618 μm wide (
<figureCitation id="87982A1EFFA10E18FD78F968CF44F950" box="[717,801,1717,1743]" captionStart="FIGURES 3841" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[284,1303,181,1919]" captionTargetId="figure-16@15.[284,1303,181,1919]" captionTargetPageId="15" captionText="FIGURES 3841. Egg morphology of Ephemera (Sinephemera) strigata: 3839, general shape and arrangement of the micropyles; 40, micropyle in the complex extrachorion-adhesive layer; 41, chorion thickness. AL-Ex—complex adhesive-extrachorion layer; C—chorion; CM—canal micropylar; MO—micropylar opening; SG—sperm guide." pageId="11" pageNumber="392">Fig. 40</figureCitation>
). The micropylar canal is 9.815.6. µm long and 5.57.2 µm wide; the entrance to the micropylar canal is tunnel-shaped, with thick walls; it protrudes above the adhesive layer and chorion (
<figureCitation id="87982A1EFFA10E18FEDFF8DDCDA4F885" box="[362,449,1792,1818]" captionStart="FIGURES 3841" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[284,1303,181,1919]" captionTargetId="figure-16@15.[284,1303,181,1919]" captionTargetPageId="15" captionText="FIGURES 3841. Egg morphology of Ephemera (Sinephemera) strigata: 3839, general shape and arrangement of the micropyles; 40, micropyle in the complex extrachorion-adhesive layer; 41, chorion thickness. AL-Ex—complex adhesive-extrachorion layer; C—chorion; CM—canal micropylar; MO—micropylar opening; SG—sperm guide." pageId="11" pageNumber="392">Figs 40</figureCitation>
,
<figureCitation id="87982A1EFFA10E18FE79F8DDCD89F885" box="[460,492,1792,1818]" captionStart="FIGURES 4245" captionStartId="16.[151,264,1943,1967]" captionTargetBox="[289,1299,181,1919]" captionTargetId="figure-18@16.[289,1299,181,1919]" captionTargetPageId="16" captionText="FIGURES 4245. Egg morphology of Ephemera (Sinephemera) strigata: 42, remains of the adhesive layer on the egg surface; 4344, proximal part of micropylar canal on chorion surface; 45, chorionic sculpturing. AL-Ex—complex adhesive-extrachorion layer; CS—chorion surface; CM—canal micropylar; MO—micropylar opening." pageId="11" pageNumber="392">43</figureCitation>
). The micropylar opening is rounded to 2.53.0 µm wide and 2.52.9 µm high (
<figureCitation id="87982A1EFFA10E18FADEF8DDCC87F8A0" captionStart="FIGURES 4245" captionStartId="16.[151,264,1943,1967]" captionTargetBox="[289,1299,181,1919]" captionTargetId="figure-18@16.[289,1299,181,1919]" captionTargetPageId="16" captionText="FIGURES 4245. Egg morphology of Ephemera (Sinephemera) strigata: 42, remains of the adhesive layer on the egg surface; 4344, proximal part of micropylar canal on chorion surface; 45, chorionic sculpturing. AL-Ex—complex adhesive-extrachorion layer; CS—chorion surface; CM—canal micropylar; MO—micropylar opening." pageId="11" pageNumber="392">Figs 4344</figureCitation>
). The chorionic surface is finely wrinkled (
<figureCitation id="87982A1EFFA10E18FD76F8F8CF23F8DF" box="[707,838,1829,1856]" captionStart="FIGURES 4245" captionStartId="16.[151,264,1943,1967]" captionTargetBox="[289,1299,181,1919]" captionTargetId="figure-18@16.[289,1299,181,1919]" captionTargetPageId="16" captionText="FIGURES 4245. Egg morphology of Ephemera (Sinephemera) strigata: 42, remains of the adhesive layer on the egg surface; 4344, proximal part of micropylar canal on chorion surface; 45, chorionic sculpturing. AL-Ex—complex adhesive-extrachorion layer; CS—chorion surface; CM—canal micropylar; MO—micropylar opening." pageId="11" pageNumber="392">Figs 43, 45</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,197 @@
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<taxonomicName id="D8A34D18FFAB0E12FF22FC1BCF62FC7F" ID-CoL="6FP7W" authority="McLachlan, 1875" authorityName="McLachlan" authorityYear="1875" box="[151,775,966,993]" class="Insecta" family="Ephemeridae" genus="Ephemera" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="382" phylum="Arthropoda" rank="species" species="orientalis" subGenus="Ephemera">
<emphasis id="2DD7EA89FFAB0E12FF22FC1BCF62FC7F" bold="true" box="[151,775,966,993]" pageId="1" pageNumber="382">
<emphasis id="2DD7EA89FFAB0E12FF22FC1BCD71FC7E" bold="true" box="[151,276,966,993]" italics="true" pageId="1" pageNumber="382">Ephemera</emphasis>
(
<emphasis id="2DD7EA89FFAB0E12FE93FC1BCDC6FC7E" bold="true" box="[294,419,966,993]" italics="true" pageId="1" pageNumber="382">Ephemera</emphasis>
)
<emphasis id="2DD7EA89FFAB0E12FE01FC1BCE43FC7E" bold="true" box="[436,550,966,993]" italics="true" pageId="1" pageNumber="382">orientalis</emphasis>
<bibRefCitation id="7B324B6AFFAB0E12FD9BFC1ACF62FC7F" author="McLachlan, R." box="[558,775,966,993]" pageId="1" pageNumber="382" pagination="167 - 190" refId="ref7290" refString="McLachlan, R. (1875) A sketch of our present knowledge of the neuropterous fauna of Japan (excluding Odonata and Trichoptera). Transactions of the Entomological Society, London, 167 - 190. https: // doi. org / 10.1111 / j. 1365 - 2311.1875. tb 01906. x" type="book chapter" year="1875">McLachlan, 1875</bibRefCitation>
</emphasis>
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<paragraph id="1F1C369BFFAB0E12FF22FC31CD79FB99" blockId="1.[151,775,966,1030]" box="[151,284,1004,1030]" pageId="1" pageNumber="382">
<figureCitation id="87982A1EFFAB0E12FF22FC31CD79FB99" box="[151,284,1004,1030]" captionStart-0="FIGURES 14" captionStart-1="FIGURES 58" captionStartId-0="2.[151,264,1943,1967]" captionStartId-1="3.[151,264,1943,1967]" captionTargetBox-0="[246,1341,181,1919]" captionTargetBox-1="[232,1355,181,1919]" captionTargetId-0="figure-18@2.[246,1341,181,1919]" captionTargetId-1="figure-16@3.[232,1355,181,1919]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 14. Egg morphology of Ephemera (Ephemera) orientalis: 12, general shape; 34, micropyle of the complex extrachorion-adhesive layer. CM—canal micropylar; MO—micropylar opening; mp—micropyle; SG—sperm guide." captionText-1="FIGURES 58. Egg morphology of Ephemera (Ephemera) orientalis: 56, egg surface without complex extrachorion-adhesive layer; 7, proximal part of micropylar canal on chorion surface; 8, thickness of the layer forming the eggshell. AL-Ex—complex adhesive-extrachorion layer; CS—chorion surface; MO—micropylar opening; mp—micropyle." pageId="1" pageNumber="382">Figures 18</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="57B96510FFAB0E12FF22FBEBC938FB04" pageId="1" pageNumber="382" type="materials_examined">
<paragraph id="1F1C369BFFAB0E12FF22FBEBC938FB04" blockId="1.[151,1437,1078,1889]" pageId="1" pageNumber="382">
<emphasis id="2DD7EA89FFAB0E12FF22FBEBCD12FBCF" bold="true" box="[151,375,1078,1104]" pageId="1" pageNumber="382">Material examined</emphasis>
.
<materialsCitation id="AFCB3CC6FFAB0E12FE31FBEAC9F9FBCE" box="[388,1436,1079,1105]" country="Russia" location="Kabarga River" municipality="Ussuri River" pageId="1" pageNumber="382" specimenCount="1" stateProvince="Primorskiy">
<collectingCountry id="67B4760BFFAB0E12FE31FBEACDB6FBCE" box="[388,467,1079,1105]" name="Russia" pageId="1" pageNumber="382">Russia</collectingCountry>
:
<collectingRegion id="DD67F879FFAB0E12FE6EFBEACEEFFBCE" box="[475,650,1079,1105]" country="Russia" name="Primorskiy" pageId="1" pageNumber="382">Primorsky Krai</collectingRegion>
,
<collectingMunicipality id="FF78ACE1FFAB0E12FD26FBEACF7AFBCE" box="[659,799,1079,1105]" pageId="1" pageNumber="382">Ussuri River</collectingMunicipality>
basin,
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, below the road bridge, Vladivostok
</materialsCitation>
<materialsCitation id="AFCB3CC6FFAB0E12FF19FB81CD01FB04" collectingDate="2000-06-17" collectingDateMax="2005-06-15" collectingDateMin="2000-06-17" collectionCode="T" country="Russia" location="Stepanovka village" municipality="Ussuri River" pageId="1" pageNumber="382" specimenCount="5" specimenCount-female="5" stateProvince="Khabarovsk">
<collectingRegion id="DD67F879FFAB0E12FF19FB81CD56FBE9" box="[172,307,1116,1142]" country="Russia" name="Khabarovsk" pageId="1" pageNumber="382">Khabarovsk</collectingRegion>
highway,
<date id="6B1D105BFFAB0E12FE1DFB81CE4DFBE9" box="[424,552,1116,1142]" pageId="1" pageNumber="382" value="2000-06-17">
<collectingDate id="7B59E9B3FFAB0E12FE1DFB81CE4DFBE9" box="[424,552,1116,1142]" pageId="1" pageNumber="382" value="2000-06-17">17.06.2000</collectingDate>
</date>
,
<specimenCount id="09A5FD12FFAB0E12FD81FB81CE33FBE9" box="[564,598,1116,1142]" count="3" pageId="1" pageNumber="382" type="female">3♀</specimenCount>
adults,
<collectionCode id="79B2AE5EFFAB0E12FD1BFB80CEA5FBE8" box="[686,704,1117,1143]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="382">T</collectionCode>
. Tiunova;
<collectingMunicipality id="FF78ACE1FFAB0E12FC83FB81CFA3FBE9" box="[822,966,1116,1142]" pageId="1" pageNumber="382">Ussuri River</collectingMunicipality>
, near
<location id="1A7C6040FFAB0E12FBBCFB81C887FBE9" LSID="urn:lsid:plazi:treatment:970A878DFFAB0E12FF22FC1BCF63F8FE:1A7C6040FFAB0E12FBBCFB81C887FBE9" box="[1033,1250,1116,1142]" country="Russia" municipality="Ussuri River" name="Stepanovka village" pageId="1" pageNumber="382" stateProvince="Khabarovsk">Stepanovka village</location>
,
<date id="6B1D105BFFAB0E12FB58FB81C90BFBE8" box="[1261,1390,1116,1143]" pageId="1" pageNumber="382" value="2005-06-15">
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,
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adults,
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. Tiunova
</materialsCitation>
;
<materialsCitation id="AFCB3CC6FFAB0E12FEDAFB5CC93CFB04" box="[367,1369,1153,1180]" collectingDate="2005-06-30" collectionCode="T" country="Russia" county="Khabarovsky Krai" location="Bystrinsk village" municipality="River" pageId="1" pageNumber="382" specimenCount="1" specimenCount-female="1" stateProvince="Amur">
<collectingCounty id="F67D4E17FFAB0E12FEDAFB5CCE24FB04" box="[367,577,1153,1179]" pageId="1" pageNumber="382">Khabarovsky Krai</collectingCounty>
,
<collectingRegion id="DD67F879FFAB0E12FDFFFB5FCEEEFB03" box="[586,651,1154,1180]" country="Russia" name="Amur" pageId="1" pageNumber="382">Amur</collectingRegion>
<collectingMunicipality id="FF78ACE1FFAB0E12FD27FB5CCEB7FB04" box="[658,722,1153,1179]" pageId="1" pageNumber="382">River</collectingMunicipality>
, near
<location id="1A7C6040FFAB0E12FCA1FB5CCFB3FB04" LSID="urn:lsid:plazi:treatment:970A878DFFAB0E12FF22FC1BCF63F8FE:1A7C6040FFAB0E12FCA1FB5CCFB3FB04" box="[788,982,1153,1179]" country="Russia" county="Khabarovsky Krai" municipality="River" name="Bystrinsk village" pageId="1" pageNumber="382" stateProvince="Amur">Bystrinsk village</location>
,
<date id="6B1D105BFFAB0E12FC54FB5CC807FB03" box="[993,1122,1153,1180]" pageId="1" pageNumber="382" value="2005-06-30">
<collectingDate id="7B59E9B3FFAB0E12FC54FB5CC807FB03" box="[993,1122,1153,1180]" pageId="1" pageNumber="382" value="2005-06-30">30.06.2005</collectingDate>
</date>
,
<specimenCount id="09A5FD12FFAB0E12FBD8FB5CC8F5FB04" box="[1133,1168,1153,1179]" count="1" pageId="1" pageNumber="382" type="female">1♀</specimenCount>
adult,
<collectionCode id="79B2AE5EFFAB0E12FB69FB5FC88BFB03" box="[1244,1262,1154,1180]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="382">T</collectionCode>
. Tiunova
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="57B96510FFAB0E12FF72FB7BCF63F8FE" pageId="1" pageNumber="382" type="distribution">
<paragraph id="1F1C369BFFAB0E12FF72FB7BC9EAFB5E" blockId="1.[151,1437,1078,1889]" box="[199,1423,1190,1217]" pageId="1" pageNumber="382">
<emphasis id="2DD7EA89FFAB0E12FF72FB7BCD3FFB5F" box="[199,346,1190,1217]" italics="true" pageId="1" pageNumber="382">
<emphasis id="2DD7EA89FFAB0E12FF72FB7BCD36FB5E" bold="true" box="[199,339,1190,1217]" italics="true" pageId="1" pageNumber="382">Distribution</emphasis>
.
</emphasis>
Siberia, Far-East
<collectingCountry id="67B4760BFFAB0E12FD91FB7ACE17FB5E" box="[548,626,1191,1217]" name="Russia" pageId="1" pageNumber="382">Russia</collectingCountry>
,
<collectingCountry id="67B4760BFFAB0E12FDC8FB7ACEA5FB5E" box="[637,704,1191,1217]" name="Japan" pageId="1" pageNumber="382">Japan</collectingCountry>
,
<collectingCountry id="67B4760BFFAB0E12FD7FFB7ACF5FFB5E" box="[714,826,1191,1217]" name="Mongolia" pageId="1" pageNumber="382">Mongolia</collectingCountry>
,
<collectingCountry id="67B4760BFFAB0E12FCF0FB7ACFE9FB5E" box="[837,908,1191,1217]" name="South Korea" pageId="1" pageNumber="382">Korea</collectingCountry>
, N.E.
<collectingCountry id="67B4760BFFAB0E12FC67FB7AC87DFB5E" box="[978,1048,1191,1217]" name="China" pageId="1" pageNumber="382">China</collectingCountry>
,
<collectingCountry id="67B4760BFFAB0E12FB96FB7AC8C3FB5E" box="[1059,1190,1191,1217]" name="Kazakhstan" pageId="1" pageNumber="382">Kazakhstan</collectingCountry>
(Irtysh River basin).
</paragraph>
<paragraph id="1F1C369BFFAB0E12FF72FB11C97FFAE4" blockId="1.[151,1437,1078,1889]" pageId="1" pageNumber="382">
The egg has been described by
<bibRefCitation id="7B324B6AFFAB0E12FD9EFB11CEB1FB79" author="Okazaki, H." box="[555,724,1228,1254]" pageId="1" pageNumber="382" pagination="8 - 10" refId="ref7347" refString="Okazaki, H. (1981) On the eggs of Japanese mayflies (1). Biology of Inland Waters, 2, 8 - 10." type="journal article" year="1981">Okazaki (1981</bibRefCitation>
, p. 9: fig. 5; 1984, p. 21: fig. 16),
<bibRefCitation id="7B324B6AFFAB0E12FBEDFB11C971FB79" author="Su, C. - R. &amp; Zhu, C. - D." box="[1112,1300,1228,1254]" pageId="1" pageNumber="382" pagination="121 - 123" refId="ref7405" refString="Su, C. - R. &amp; Zhu, C. - D. (1997) Observation on the chorion structure of four species of mayfly by scanning electron microscope. Journal of Nanjing Normal University (Nature Science), 20 (4), 121 - 123." type="journal article" year="1997">Su &amp; Zhu (1997</bibRefCitation>
, p. 122: fig. 70), and
<bibRefCitation id="7B324B6AFFAB0E12FF4CFB2CCD98FA94" author="Tojo, K. &amp; Machida, R." box="[249,509,1265,1291]" pageId="1" pageNumber="382" pagination="573 - 579" refId="ref7457" refString="Tojo, K. &amp; Machida, R. (1998) Egg Structures of Japanes Ephemeridae species (Ephemeroptera). Entomological Science, 1 (4), 573 - 579." type="journal article" year="1998">Tojo &amp; Machida (1998</bibRefCitation>
, p. 576: fig. 3). From the above descriptions, the following can be emphasized: the egg is rectangular in shape, with a length of 187.0 µm and a width of 120.8 µm. In the work of Tojo and Machida, the egg is ellipsoidal in shape, about 200x100 µm, the adhesive layer is thick (34 µm), smooth, the chorion has well-defined reticulation, the opening diameter is 11.5 µm, and the sperm guide is hood-like in shape.
</paragraph>
<paragraph id="1F1C369BFFAB0E12FF72FA5ACF63F8FE" blockId="1.[151,1437,1078,1889]" pageId="1" pageNumber="382">
According to our data, the egg has an oval shape close to quadrangular with rounded corners (
<figureCitation id="87982A1EFFAB0E12FA90FA5AC9EAFA3E" box="[1317,1423,1415,1441]" captionStart="FIGURES 14" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[246,1341,181,1919]" captionTargetId="figure-18@2.[246,1341,181,1919]" captionTargetPageId="2" captionText="FIGURES 14. Egg morphology of Ephemera (Ephemera) orientalis: 12, general shape; 34, micropyle of the complex extrachorion-adhesive layer. CM—canal micropylar; MO—micropylar opening; mp—micropyle; SG—sperm guide." pageId="1" pageNumber="382">Figs 12</figureCitation>
). Dimensions: 173.0206.0 µm in length (191.3 µm) and 104.1123.4 µm in width (111.0 µm). The attachment structure is a complex extrachorion-adhesive layer (AL-Ex) that covers the entire surface of the chorion (CS) (
<figureCitation id="87982A1EFFAB0E12FADFFA0CCCA6F98E" captionStart="FIGURES 14" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[246,1341,181,1919]" captionTargetId="figure-18@2.[246,1341,181,1919]" captionTargetPageId="2" captionText="FIGURES 14. Egg morphology of Ephemera (Ephemera) orientalis: 12, general shape; 34, micropyle of the complex extrachorion-adhesive layer. CM—canal micropylar; MO—micropylar opening; mp—micropyle; SG—sperm guide." pageId="1" pageNumber="382">Figs 12</figureCitation>
,
<figureCitation id="87982A1EFFAB0E12FF65FA2BCC9BF98E" box="[208,254,1526,1553]" captionStart="FIGURES 58" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[232,1355,181,1919]" captionTargetId="figure-16@3.[232,1355,181,1919]" captionTargetPageId="3" captionText="FIGURES 58. Egg morphology of Ephemera (Ephemera) orientalis: 56, egg surface without complex extrachorion-adhesive layer; 7, proximal part of micropylar canal on chorion surface; 8, thickness of the layer forming the eggshell. AL-Ex—complex adhesive-extrachorion layer; CS—chorion surface; MO—micropylar opening; mp—micropyle." pageId="1" pageNumber="382">56</figureCitation>
). Its thickness for
<taxonomicName id="D8A34D18FFAB0E12FE7BFA2ACE3CF98E" authorityName="McLachlan" authorityYear="1875" box="[462,601,1527,1553]" class="Insecta" family="Ephemeridae" genus="Ephemera" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="382" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="2DD7EA89FFAB0E12FE7BFA2ACE3CF98E" box="[462,601,1527,1553]" italics="true" pageId="1" pageNumber="382">E. orientalis</emphasis>
</taxonomicName>
is 1.6 1.8 µm (
<figureCitation id="87982A1EFFAB0E12FCA0FA2ACF3FF98E" box="[789,858,1527,1553]" captionStart="FIGURES 58" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[232,1355,181,1919]" captionTargetId="figure-16@3.[232,1355,181,1919]" captionTargetPageId="3" captionText="FIGURES 58. Egg morphology of Ephemera (Ephemera) orientalis: 56, egg surface without complex extrachorion-adhesive layer; 7, proximal part of micropylar canal on chorion surface; 8, thickness of the layer forming the eggshell. AL-Ex—complex adhesive-extrachorion layer; CS—chorion surface; MO—micropylar opening; mp—micropyle." pageId="1" pageNumber="382">Fig. 8</figureCitation>
). The surface of the adhesive layer (AL) is slightly bumpy (
<figureCitation id="87982A1EFFAB0E12FF4DF9C1CD04F9A9" box="[248,353,1564,1590]" captionStart="FIGURES 14" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[246,1341,181,1919]" captionTargetId="figure-18@2.[246,1341,181,1919]" captionTargetPageId="2" captionText="FIGURES 14. Egg morphology of Ephemera (Ephemera) orientalis: 12, general shape; 34, micropyle of the complex extrachorion-adhesive layer. CM—canal micropylar; MO—micropylar opening; mp—micropyle; SG—sperm guide." pageId="1" pageNumber="382">Figs 12</figureCitation>
). There are one or two micropyles (mp) per egg in the equatorial area (
<figureCitation id="87982A1EFFAB0E12FB06F9C1C979F9A9" box="[1203,1308,1564,1590]" captionStart="FIGURES 14" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[246,1341,181,1919]" captionTargetId="figure-18@2.[246,1341,181,1919]" captionTargetPageId="2" captionText="FIGURES 14. Egg morphology of Ephemera (Ephemera) orientalis: 12, general shape; 34, micropyle of the complex extrachorion-adhesive layer. CM—canal micropylar; MO—micropylar opening; mp—micropyle; SG—sperm guide." pageId="1" pageNumber="382">Figs 12</figureCitation>
,
<figureCitation id="87982A1EFFAB0E12FA99F9C1C93FF9A8" box="[1324,1370,1564,1591]" captionStart="FIGURES 58" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[232,1355,181,1919]" captionTargetId="figure-16@3.[232,1355,181,1919]" captionTargetPageId="3" captionText="FIGURES 58. Egg morphology of Ephemera (Ephemera) orientalis: 56, egg surface without complex extrachorion-adhesive layer; 7, proximal part of micropylar canal on chorion surface; 8, thickness of the layer forming the eggshell. AL-Ex—complex adhesive-extrachorion layer; CS—chorion surface; MO—micropylar opening; mp—micropyle." pageId="1" pageNumber="382">56</figureCitation>
). The micropyles are “tagenoform-type,” with a well-expressed sperm guide (SG) oval and relatively deep (
<figureCitation id="87982A1EFFAB0E12FA92F99CC9F5F9C4" box="[1319,1424,1601,1627]" captionStart="FIGURES 14" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[246,1341,181,1919]" captionTargetId="figure-18@2.[246,1341,181,1919]" captionTargetPageId="2" captionText="FIGURES 14. Egg morphology of Ephemera (Ephemera) orientalis: 12, general shape; 34, micropyle of the complex extrachorion-adhesive layer. CM—canal micropylar; MO—micropylar opening; mp—micropyle; SG—sperm guide." pageId="1" pageNumber="382">Figs 34</figureCitation>
). Sperm guide is 5.810.4 µm long and 5.06.1 µm wide. It should be noted that the sperm guide is only clearly visible on the adhesive surface of the egg. Once the adhesive layer has been removed, the sperm guide is not visible on the chorion (
<figureCitation id="87982A1EFFAB0E12FEFCF96CCDCBF954" box="[329,430,1713,1739]" captionStart="FIGURES 14" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[246,1341,181,1919]" captionTargetId="figure-18@2.[246,1341,181,1919]" captionTargetPageId="2" captionText="FIGURES 14. Egg morphology of Ephemera (Ephemera) orientalis: 12, general shape; 34, micropyle of the complex extrachorion-adhesive layer. CM—canal micropylar; MO—micropylar opening; mp—micropyle; SG—sperm guide." pageId="1" pageNumber="382">Figs 3, 4</figureCitation>
,
<figureCitation id="87982A1EFFAB0E12FE0FF96CCDAFF954" box="[442,458,1713,1739]" captionStart="FIGURES 58" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[232,1355,181,1919]" captionTargetId="figure-16@3.[232,1355,181,1919]" captionTargetPageId="3" captionText="FIGURES 58. Egg morphology of Ephemera (Ephemera) orientalis: 56, egg surface without complex extrachorion-adhesive layer; 7, proximal part of micropylar canal on chorion surface; 8, thickness of the layer forming the eggshell. AL-Ex—complex adhesive-extrachorion layer; CS—chorion surface; MO—micropylar opening; mp—micropyle." pageId="1" pageNumber="382">6</figureCitation>
). The micropylar canal, 1218 µm long and 46 µm wide, protrudes slightly above the adhesive layer (
<figureCitation id="87982A1EFFAB0E12FEFFF90ACDD4F96E" box="[330,433,1751,1777]" captionStart="FIGURES 14" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[246,1341,181,1919]" captionTargetId="figure-18@2.[246,1341,181,1919]" captionTargetPageId="2" captionText="FIGURES 14. Egg morphology of Ephemera (Ephemera) orientalis: 12, general shape; 34, micropyle of the complex extrachorion-adhesive layer. CM—canal micropylar; MO—micropylar opening; mp—micropyle; SG—sperm guide." pageId="1" pageNumber="382">Figs 23</figureCitation>
,
<figureCitation id="87982A1EFFAB0E12FE0BF90BCDABF96E" box="[446,462,1750,1777]" captionStart="FIGURES 58" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[232,1355,181,1919]" captionTargetId="figure-16@3.[232,1355,181,1919]" captionTargetPageId="3" captionText="FIGURES 58. Egg morphology of Ephemera (Ephemera) orientalis: 56, egg surface without complex extrachorion-adhesive layer; 7, proximal part of micropylar canal on chorion surface; 8, thickness of the layer forming the eggshell. AL-Ex—complex adhesive-extrachorion layer; CS—chorion surface; MO—micropylar opening; mp—micropyle." pageId="1" pageNumber="382">5</figureCitation>
). As shown previously (
<bibRefCitation id="7B324B6AFFAB0E12FD56F90AC87EF96E" author="Ubero-Pascal, N. A. &amp; Puig, M. A." box="[739,1051,1751,1777]" pageId="1" pageNumber="382" pagination="15 - 29" refId="ref7527" refString="Ubero-Pascal, N. A. &amp; Puig, M. A. (2007) Egg morphology update based on new chorionic data of Potamanthus luteus (Linnaeus), Ephemera danica Muller and Oligoneuriella rhenana (Imhoff) (Insecta, Ephemeroptera) obtained by scanning electron microscopy. Zootaxa, 1465 (1), 15 - 29. https: // doi. org / 10.11646 / zootaxa. 1465.1.2" type="journal article" year="2007">Ubero-Pascal &amp; Puig 2007</bibRefCitation>
), features of the chorion structure are visible on SEM only after removal of the adhesive layer. For
<emphasis id="2DD7EA89FFAB0E12FCCEF920C868F889" box="[891,1037,1788,1814]" italics="true" pageId="1" pageNumber="382">
<taxonomicName id="D8A34D18FFAB0E12FCCEF920C86CF889" authorityName="McLachlan" authorityYear="1875" box="[891,1033,1788,1814]" class="Insecta" family="Ephemeridae" genus="Ephemera" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="382" phylum="Arthropoda" rank="species" species="orientalis">E. orientalis</taxonomicName>
,
</emphasis>
the surface of the chorion is finely wrinkled (
<figureCitation id="87982A1EFFAB0E12FEBFF8FCCD17F8A3" box="[266,370,1825,1852]" captionStart="FIGURES 58" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[232,1355,181,1919]" captionTargetId="figure-16@3.[232,1355,181,1919]" captionTargetPageId="3" captionText="FIGURES 58. Egg morphology of Ephemera (Ephemera) orientalis: 56, egg surface without complex extrachorion-adhesive layer; 7, proximal part of micropylar canal on chorion surface; 8, thickness of the layer forming the eggshell. AL-Ex—complex adhesive-extrachorion layer; CS—chorion surface; MO—micropylar opening; mp—micropyle." pageId="1" pageNumber="382">Figs 56</figureCitation>
). The thickened proximal part of the tunnel-type micropylar canal and micropylar opening are clearly visible on the surface of the chorion. (
<figureCitation id="87982A1EFFAB0E12FD26F89ACE9CF8FE" box="[659,761,1863,1889]" captionStart="FIGURES 58" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[232,1355,181,1919]" captionTargetId="figure-16@3.[232,1355,181,1919]" captionTargetPageId="3" captionText="FIGURES 58. Egg morphology of Ephemera (Ephemera) orientalis: 56, egg surface without complex extrachorion-adhesive layer; 7, proximal part of micropylar canal on chorion surface; 8, thickness of the layer forming the eggshell. AL-Ex—complex adhesive-extrachorion layer; CS—chorion surface; MO—micropylar opening; mp—micropyle." pageId="1" pageNumber="382">Figs 67</figureCitation>
).
</paragraph>
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<paragraph id="1F1C369BFFAC0E15FF22FC86CF50FCEA" blockId="6.[151,821,859,922]" box="[151,821,859,886]" pageId="6" pageNumber="387">
<heading id="445481F7FFAC0E15FF22FC86CF50FCEA" bold="true" box="[151,821,859,886]" fontSize="11" level="1" pageId="6" pageNumber="387" reason="1">
<taxonomicName id="D8A34D18FFAC0E15FF22FC86CF50FCEA" ID-CoL="6FP9P" authority="Tshernova, 1973" authorityName="Tshernova" authorityYear="1973" box="[151,821,859,886]" class="Insecta" family="Ephemeridae" genus="Ephemera" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="387" phylum="Arthropoda" rank="species" species="transbajkalica" subGenus="Ephemera">
<emphasis id="2DD7EA89FFAC0E15FF22FC86CF50FCEA" bold="true" box="[151,821,859,886]" pageId="6" pageNumber="387">
<emphasis id="2DD7EA89FFAC0E15FF22FC86CD71FCE9" bold="true" box="[151,276,859,886]" italics="true" pageId="6" pageNumber="387">Ephemera</emphasis>
(
<emphasis id="2DD7EA89FFAC0E15FE93FC86CDC6FCE9" bold="true" box="[294,419,859,886]" italics="true" pageId="6" pageNumber="387">Ephemera</emphasis>
)
<emphasis id="2DD7EA89FFAC0E15FE01FC86CE04FCE9" bold="true" box="[436,609,859,886]" italics="true" pageId="6" pageNumber="387">transbajkalica</emphasis>
<bibRefCitation id="7B324B6AFFAC0E15FDDDFC86CF50FCEA" author="Tshernova, O. A." box="[616,821,859,885]" pageId="6" pageNumber="387" pagination="324 - 339" refId="ref7491" refString="Tshernova, O. A. (1973) Palearctic mayfly species of the genus Ephemera (Ephemeroptera, Ephemeridae). Revue d'Entomologie de l'URSS, 52 (2), 324 - 339." type="journal article" year="1973">Tshernova, 1973</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="57B96510FFAC0E15FF22FC5DCD5DFC05" box="[151,312,896,922]" pageId="6" pageNumber="387" type="description">
<paragraph id="1F1C369BFFAC0E15FF22FC5DCD5DFC05" blockId="6.[151,821,859,922]" box="[151,312,896,922]" pageId="6" pageNumber="387">
<figureCitation id="87982A1EFFAC0E15FF22FC5DCD5DFC05" box="[151,312,896,922]" captionStart-0="FIGURES 1619" captionStart-1="FIGURES 2023" captionStartId-0="7.[151,264,1943,1967]" captionStartId-1="8.[151,264,1905,1929]" captionTargetBox-0="[219,1369,181,1919]" captionTargetBox-1="[287,1299,151,1889]" captionTargetId-0="figure-16@7.[219,1369,181,1919]" captionTargetId-1="figure-18@8.[287,1299,151,1889]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURES 1619. Egg morphology of Ephemera (Ephemera) transbajkalica: 1617, general shape; 1819, micropyle in the complex extrachorion-adhesive layer. AL-Ex—complex adhesive-extrachorion layer; CM—canal micropylar; mp—micropyle; SG—sperm guide." captionText-1="FIGURES 2023. Egg morphology of Ephemera (Ephemera) transbajkalica: 20, egg surface without complex extrachorionadhesive layer; 21, thickness on the layer forming the eggshell; 22, proximal part of micropylar canal on chorion surface; 23, chorionic sculpturing. AL-Ex—complex adhesive-extrachorion layer; CS—chorion surface; MO—micropylar opening; mp—micropyle." pageId="6" pageNumber="387">Figures 1623</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="57B96510FFAC0E15FF22FC16CF29FBAF" pageId="6" pageNumber="387" type="materials_examined">
<paragraph id="1F1C369BFFAC0E15FF22FC16CF29FBAF" blockId="6.[151,1436,971,1408]" pageId="6" pageNumber="387">
<emphasis id="2DD7EA89FFAC0E15FF22FC16CD1CFC7A" bold="true" box="[151,377,971,997]" pageId="6" pageNumber="387">Material examined</emphasis>
.
<materialsCitation id="AFCB3CC6FFAC0E15FE32FC16CEC4FB95" collectingDate="2004-06-16" collectionCode="T" country="Russia" county="Selemdzha River" location="Nora River" municipality="Burunda River" pageId="6" pageNumber="387" specimenCount="2" specimenCount-female="2" stateProvince="Amur">
<collectingCountry id="67B4760BFFAC0E15FE32FC16CDB3FC7A" box="[391,470,971,997]" name="Russia" pageId="6" pageNumber="387">Russia</collectingCountry>
:
<collectingRegion id="DD67F879FFAC0E15FE6AFC16CE16FC7A" box="[479,627,971,997]" country="Russia" name="Amur" pageId="6" pageNumber="387">Amur Oblast</collectingRegion>
,
<collectingCounty id="F67D4E17FFAC0E15FDCBFC16CF58FC7A" box="[638,829,971,997]" pageId="6" pageNumber="387">Selemdzha River</collectingCounty>
basin,
<collectingMunicipality id="FF78ACE1FFAC0E15FC38FC16C851FC7A" box="[909,1076,971,997]" pageId="6" pageNumber="387">Burunda River</collectingMunicipality>
, mouth, tributary of
<location id="1A7C6040FFAC0E15FAACFC11C9FDFC7A" LSID="urn:lsid:plazi:treatment:970A878DFFAC0E15FF22FC86C92DFA1F:1A7C6040FFAC0E15FAACFC11C9FDFC7A" box="[1305,1432,971,998]" country="Russia" county="Selemdzha River" municipality="Burunda River" name="Nora River" pageId="6" pageNumber="387" stateProvince="Amur">Nora River</location>
,
<date id="6B1D105BFFAC0E15FF22FC2DCD7DFB95" box="[151,280,1008,1034]" pageId="6" pageNumber="387" value="2004-06-16">
<collectingDate id="7B59E9B3FFAC0E15FF22FC2DCD7DFB95" box="[151,280,1008,1034]" pageId="6" pageNumber="387" value="2004-06-16">16.06.2004</collectingDate>
</date>
,
<specimenCount id="09A5FD12FFAC0E15FE97FC2CCD20FB94" box="[290,325,1009,1035]" count="2" pageId="6" pageNumber="387" type="female">2♀</specimenCount>
adults,
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. Tiunova; Republic of
</materialsCitation>
<materialsCitation id="AFCB3CC6FFAC0E15FD12FC2DCF2DFBAF" collectingDate="2006-01-08" collectionCode="T" country="Russia" county="Aldan River" location="Ungrinsky reserve" municipality="Ungra River" pageId="6" pageNumber="387" specimenCount="2" specimenCount-female="2" stateProvince="Sakha">
<collectingRegion id="DD67F879FFAC0E15FD12FC2DCE8EFB95" box="[679,747,1008,1034]" country="Russia" name="Sakha" pageId="6" pageNumber="387">Sakha</collectingRegion>
(Yakutia),
<collectingCounty id="F67D4E17FFAC0E15FCD0FC2DCF8BFB95" box="[869,1006,1008,1034]" pageId="6" pageNumber="387">Aldan River</collectingCounty>
basin,
<collectingMunicipality id="FF78ACE1FFAC0E15FB89FC2CC8A2FB95" box="[1084,1223,1008,1035]" pageId="6" pageNumber="387">Ungra River</collectingMunicipality>
, base of
<location id="1A7C6040FFAC0E15FA93FC2DCC8DFBAF" LSID="urn:lsid:plazi:treatment:970A878DFFAC0E15FF22FC86C92DFA1F:1A7C6040FFAC0E15FA93FC2DCC8DFBAF" country="Russia" county="Aldan River" municipality="Ungra River" name="Ungrinsky reserve" pageId="6" pageNumber="387" stateProvince="Sakha">Ungrinsky reserve</location>
“Yukhta,”
<date id="6B1D105BFFAC0E15FED0FBCBCD83FBAF" box="[357,486,1046,1072]" pageId="6" pageNumber="387" value="2006-01-08">
<collectingDate id="7B59E9B3FFAC0E15FED0FBCBCD83FBAF" box="[357,486,1046,1072]" pageId="6" pageNumber="387" value="2006-01-08">01.08.2006</collectingDate>
</date>
,
<specimenCount id="09A5FD12FFAC0E15FE44FBCBCE71FBAF" box="[497,532,1046,1072]" count="2" pageId="6" pageNumber="387" type="female">2♀</specimenCount>
adults (reared),
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. Tiunova
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="57B96510FFAC0E15FF72FBE6C92DFA1F" pageId="6" pageNumber="387" type="distribution">
<paragraph id="1F1C369BFFAC0E15FF72FBE6CF42FBCA" blockId="6.[151,1436,971,1408]" box="[199,807,1083,1110]" pageId="6" pageNumber="387">
<emphasis id="2DD7EA89FFAC0E15FF72FBE6CD3FFBC9" bold="true" box="[199,346,1083,1110]" italics="true" pageId="6" pageNumber="387">Distribution.</emphasis>
East Siberia, Far East
<collectingCountry id="67B4760BFFAC0E15FDECFBE6CEC2FBCA" box="[601,679,1083,1109]" name="Russia" pageId="6" pageNumber="387">Russia</collectingCountry>
,
<collectingCountry id="67B4760BFFAC0E15FD07FBE6CF46FBCA" box="[690,803,1083,1109]" name="Mongolia" pageId="6" pageNumber="387">Mongolia</collectingCountry>
.
</paragraph>
<paragraph id="1F1C369BFFAC0E15FF72FBBDC92DFA1F" blockId="6.[151,1436,971,1408]" pageId="6" pageNumber="387">
The egg is oval-shaped (
<figureCitation id="87982A1EFFAC0E15FE6CFBBDCE39FBE5" box="[473,604,1120,1146]" captionStart="FIGURES 1619" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[219,1369,181,1919]" captionTargetId="figure-16@7.[219,1369,181,1919]" captionTargetPageId="7" captionText="FIGURES 1619. Egg morphology of Ephemera (Ephemera) transbajkalica: 1617, general shape; 1819, micropyle in the complex extrachorion-adhesive layer. AL-Ex—complex adhesive-extrachorion layer; CM—canal micropylar; mp—micropyle; SG—sperm guide." pageId="6" pageNumber="387">Figs 1617</figureCitation>
). Dimensions: 220.0226.0 µm in length (220.8 µm) and 123.0129.0 µm in width (124.6 µm). The thickness of the extrachorion-adhesive layer is 1.31.7 µm (
<figureCitation id="87982A1EFFAC0E15FBF0FB5BC8A1FB3F" box="[1093,1220,1158,1184]" captionStart="FIGURES 2023" captionStartId="8.[151,264,1905,1929]" captionTargetBox="[287,1299,151,1889]" captionTargetId="figure-18@8.[287,1299,151,1889]" captionTargetPageId="8" captionText="FIGURES 2023. Egg morphology of Ephemera (Ephemera) transbajkalica: 20, egg surface without complex extrachorionadhesive layer; 21, thickness on the layer forming the eggshell; 22, proximal part of micropylar canal on chorion surface; 23, chorionic sculpturing. AL-Ex—complex adhesive-extrachorion layer; CS—chorion surface; MO—micropylar opening; mp—micropyle." pageId="6" pageNumber="387">Figs 21, 23</figureCitation>
). The surface of the adhesive layer is slightly roughened; almost smooth (
<figureCitation id="87982A1EFFAC0E15FD53FB76CF03FB5A" box="[742,870,1195,1221]" captionStart="FIGURES 1619" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[219,1369,181,1919]" captionTargetId="figure-16@7.[219,1369,181,1919]" captionTargetPageId="7" captionText="FIGURES 1619. Egg morphology of Ephemera (Ephemera) transbajkalica: 1617, general shape; 1819, micropyle in the complex extrachorion-adhesive layer. AL-Ex—complex adhesive-extrachorion layer; CM—canal micropylar; mp—micropyle; SG—sperm guide." pageId="6" pageNumber="387">Figs 16, 18</figureCitation>
). There are one or two micropyles in the equatorial area (
<figureCitation id="87982A1EFFAC0E15FF60FB0DCD35FB75" box="[213,336,1232,1258]" captionStart="FIGURES 1619" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[219,1369,181,1919]" captionTargetId="figure-16@7.[219,1369,181,1919]" captionTargetPageId="7" captionText="FIGURES 1619. Egg morphology of Ephemera (Ephemera) transbajkalica: 1617, general shape; 1819, micropyle in the complex extrachorion-adhesive layer. AL-Ex—complex adhesive-extrachorion layer; CM—canal micropylar; mp—micropyle; SG—sperm guide." pageId="6" pageNumber="387">Figs1617</figureCitation>
); micropyles are “tagenoform-type”; sperm guide well expressed (10.019.6 µm long, 4.27.2 µm wide), elongated, oval, with pointed distal margin, not deep (
<figureCitation id="87982A1EFFAC0E15FCFDFB2BCFA9FA8F" box="[840,972,1270,1296]" captionStart="FIGURES 1619" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[219,1369,181,1919]" captionTargetId="figure-16@7.[219,1369,181,1919]" captionTargetPageId="7" captionText="FIGURES 1619. Egg morphology of Ephemera (Ephemera) transbajkalica: 1617, general shape; 1819, micropyle in the complex extrachorion-adhesive layer. AL-Ex—complex adhesive-extrachorion layer; CM—canal micropylar; mp—micropyle; SG—sperm guide." pageId="6" pageNumber="387">Figs 1819</figureCitation>
). Micropylar canal: 4.06.0 µm long and 2.37.1 µm wide; weakly protrudes above the adhesive layer (
<figureCitation id="87982A1EFFAC0E15FCFBFAC6CFB5FAAA" box="[846,976,1307,1333]" captionStart="FIGURES 1619" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[219,1369,181,1919]" captionTargetId="figure-16@7.[219,1369,181,1919]" captionTargetPageId="7" captionText="FIGURES 1619. Egg morphology of Ephemera (Ephemera) transbajkalica: 1617, general shape; 1819, micropyle in the complex extrachorion-adhesive layer. AL-Ex—complex adhesive-extrachorion layer; CM—canal micropylar; mp—micropyle; SG—sperm guide." pageId="6" pageNumber="387">Figs 16, 18</figureCitation>
). The chorionic surface is covered by the regular mesh units penta- or hexagonal cells, each with a flat, almost smooth bottom and a convex protuberance in the middle (
<figureCitation id="87982A1EFFAC0E15FEA8FABBCD17FA1F" box="[285,370,1382,1408]" captionStart="FIGURES 2023" captionStartId="8.[151,264,1905,1929]" captionTargetBox="[287,1299,151,1889]" captionTargetId="figure-18@8.[287,1299,151,1889]" captionTargetPageId="8" captionText="FIGURES 2023. Egg morphology of Ephemera (Ephemera) transbajkalica: 20, egg surface without complex extrachorionadhesive layer; 21, thickness on the layer forming the eggshell; 22, proximal part of micropylar canal on chorion surface; 23, chorionic sculpturing. AL-Ex—complex adhesive-extrachorion layer; CS—chorion surface; MO—micropylar opening; mp—micropyle." pageId="6" pageNumber="387">Fig. 23</figureCitation>
). The proximal part of the micropylar opening elevates above the chorion (
<figureCitation id="87982A1EFFAC0E15FB0CFABBC959FA1F" box="[1209,1340,1382,1408]" captionStart="FIGURES 2023" captionStartId="8.[151,264,1905,1929]" captionTargetBox="[287,1299,151,1889]" captionTargetId="figure-18@8.[287,1299,151,1889]" captionTargetPageId="8" captionText="FIGURES 2023. Egg morphology of Ephemera (Ephemera) transbajkalica: 20, egg surface without complex extrachorionadhesive layer; 21, thickness on the layer forming the eggshell; 22, proximal part of micropylar canal on chorion surface; 23, chorionic sculpturing. AL-Ex—complex adhesive-extrachorion layer; CS—chorion surface; MO—micropylar opening; mp—micropyle." pageId="6" pageNumber="387">Figs 20, 22</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,175 @@
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<treatment id="970A878DFFAC0E15FF22FF4ACDEEFC98" LSID="urn:lsid:plazi:treatment:970A878DFFAC0E15FF22FF4ACDEEFC98" httpUri="http://treatment.plazi.org/id/970A878DFFAC0E15FF22FF4ACDEEFC98" lastPageNumber="387" pageId="6" pageNumber="387">
<subSubSection id="57B96510FFAC0E15FF22FF4ACF5EFF2E" box="[151,827,151,178]" pageId="6" pageNumber="387" type="nomenclature">
<paragraph id="1F1C369BFFAC0E15FF22FF4ACF5EFF2E" blockId="6.[151,827,151,215]" box="[151,827,151,178]" pageId="6" pageNumber="387">
<heading id="445481F7FFAC0E15FF22FF4ACF5EFF2E" bold="true" box="[151,827,151,178]" fontSize="11" level="1" pageId="6" pageNumber="387" reason="1">
<taxonomicName id="D8A34D18FFAC0E15FF22FF4ACF5EFF2E" ID-CoL="6FP9X" authority="Matsumura, 1911" authorityName="Matsumura" authorityYear="1911" box="[151,827,151,178]" class="Insecta" family="Ephemeridae" genus="Ephemera" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="387" phylum="Arthropoda" rank="species" species="sachalinensis" subGenus="Ephemera">
<emphasis id="2DD7EA89FFAC0E15FF22FF4ACF5EFF2E" bold="true" box="[151,827,151,178]" pageId="6" pageNumber="387">
<emphasis id="2DD7EA89FFAC0E15FF22FF4ACD71FF2D" bold="true" box="[151,276,151,178]" italics="true" pageId="6" pageNumber="387">Ephemera</emphasis>
(
<emphasis id="2DD7EA89FFAC0E15FE93FF4ACDC6FF2D" bold="true" box="[294,419,151,178]" italics="true" pageId="6" pageNumber="387">Ephemera</emphasis>
)
<emphasis id="2DD7EA89FFAC0E15FE01FF4ACE32FF2D" bold="true" box="[436,599,151,178]" italics="true" pageId="6" pageNumber="387">sachalinensis</emphasis>
Matsumura, 1911
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="57B96510FFAC0E15FF22FF61CD4FFF48" box="[151,298,188,215]" pageId="6" pageNumber="387" type="description">
<paragraph id="1F1C369BFFAC0E15FF22FF61CD4FFF48" blockId="6.[151,827,151,215]" box="[151,298,188,215]" pageId="6" pageNumber="387">
<figureCitation id="87982A1EFFAC0E15FF22FF61CD4FFF48" box="[151,298,188,215]" captionStart-0="FIGURES 911" captionStart-1="FIGURES 1215" captionStartId-0="4.[151,264,1898,1922]" captionStartId-1="5.[151,264,1943,1967]" captionTargetBox-0="[265,1322,151,1889]" captionTargetBox-1="[204,1384,181,1919]" captionTargetId-0="figure-18@4.[265,1322,151,1889]" captionTargetId-1="figure-16@5.[204,1384,181,1919]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURES 911. Egg morphology of Ephemera (Ephemera) sachalinensis: 9, general shape; 1011, micropyle in the complex extrachorion-adhesive layer. AL-Ex—complex adhesive-extrachorion layer; MO—micropylar opening; mp—micropyle; CM—canal micropylar; SG— sperm guide." captionText-1="FIGURES 1215. Egg morphology of Ephemera (Ephemera) sachalinensis: 1213, egg surface without complex extrachorionadhesive layer; 14, proximal part of micropylar canal on chorion surface; 15, thickness on the layer forming the eggshell. AL-Ex—complex adhesive-extrachorion layer; CS—chorion surface; MO—micropylar opening; mp—micropyle." pageId="6" pageNumber="387">Figures 915</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="57B96510FFAC0E15FF22FEDAC969FE0E" pageId="6" pageNumber="387" type="materials_examined">
<paragraph id="1F1C369BFFAC0E15FF22FEDAC969FE0E" blockId="6.[151,1437,263,775]" pageId="6" pageNumber="387">
<emphasis id="2DD7EA89FFAC0E15FF22FEDACDE3FEBE" bold="true" box="[151,390,263,289]" pageId="6" pageNumber="387">Material examined.</emphasis>
<materialsCitation id="AFCB3CC6FFAC0E15FE26FEDACDF6FEF3" collectingDate="2003-11-06" collectingDateMax="2007-03-08" collectingDateMin="2003-11-06" collectionCode="T" country="Russia" county="Khasansky District" location="Fish Hatchery" municipality="Barabashevka River" pageId="6" pageNumber="387" specimenCount="4" specimenCount-female="4" stateProvince="Primorskiy">
<collectingCountry id="67B4760BFFAC0E15FE26FEDACD87FEBE" box="[403,482,263,289]" name="Russia" pageId="6" pageNumber="387">Russia</collectingCountry>
:
<collectingRegion id="DD67F879FFAC0E15FE47FEDACECDFEBE" box="[498,680,263,289]" country="Russia" name="Primorskiy" pageId="6" pageNumber="387">Primorsky Krai</collectingRegion>
,
<collectingCounty id="F67D4E17FFAC0E15FD0CFEDACFF2FEBE" box="[697,919,263,289]" pageId="6" pageNumber="387">Khasansky District</collectingCounty>
,
<collectingMunicipality id="FF78ACE1FFAC0E15FC1DFEDAC8F7FEBE" box="[936,1170,263,289]" pageId="6" pageNumber="387">Barabashevka River</collectingMunicipality>
, below
<location id="1A7C6040FFAC0E15FB41FEDAC9FCFEBE" LSID="urn:lsid:plazi:treatment:970A878DFFAC0E15FF22FF4ACDEEFC98:1A7C6040FFAC0E15FB41FEDAC9FCFEBE" box="[1268,1433,263,289]" country="Russia" county="Khasansky District" municipality="Barabashevka River" name="Fish Hatchery" pageId="6" pageNumber="387" stateProvince="Primorskiy">Fish Hatchery</location>
,
<date id="6B1D105BFFAC0E15FF22FEF0CD7DFED8" box="[151,280,301,327]" pageId="6" pageNumber="387" value="2007-03-08">
<collectingDate id="7B59E9B3FFAC0E15FF22FEF0CD7DFED8" box="[151,280,301,327]" pageId="6" pageNumber="387" value="2007-03-08">03.08.2007</collectingDate>
</date>
,
<specimenCount id="09A5FD12FFAC0E15FE9DFEF0CD2EFED8" box="[296,331,301,327]" count="2" pageId="6" pageNumber="387" type="female">2♀</specimenCount>
adults,
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. Tiunova;
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, below Okhotbaza,
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<collectingDate id="7B59E9B3FFAC0E15FBB7FEF1C8E4FED9" box="[1026,1153,300,326]" pageId="6" pageNumber="387" value="2003-11-06">11.06.2003</collectingDate>
</date>
,
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adults,
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. Tiunova;
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</materialsCitation>
,
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<collectingRegion id="DD67F879FFAC0E15FE10FE8FCE62FEF3" box="[421,519,338,364]" country="Russia" name="Sakhalin" pageId="6" pageNumber="387">Sakhalin</collectingRegion>
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,
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, eastern part,
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<collectingDate id="7B59E9B3FFAC0E15FC5AFE8FC80AFEF3" box="[1007,1135,338,364]" pageId="6" pageNumber="387" value="2002-07-18">18.07.2002</collectingDate>
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,
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adults,
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. Teslenko;
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</materialsCitation>
,
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<collectingRegion id="DD67F879FFAC0E15FEC0FEAACDD3FE0E" box="[373,438,375,401]" country="Russia" name="Amur" pageId="6" pageNumber="387">Amur</collectingRegion>
<collectingCounty id="F67D4E17FFAC0E15FE08FEAACD9EFE0E" box="[445,507,375,401]" pageId="6" pageNumber="387">River</collectingCounty>
basin,
<collectingMunicipality id="FF78ACE1FFAC0E15FDFEFEA5CE86FE0E" box="[587,739,375,402]" pageId="6" pageNumber="387">Bureya River</collectingMunicipality>
, below
<location id="1A7C6040FFAC0E15FC8FFEAAC863FE0E" LSID="urn:lsid:plazi:treatment:970A878DFFAC0E15FF22FF4ACDEEFC98:1A7C6040FFAC0E15FC8FFEAAC863FE0E" box="[826,1030,375,401]" country="Russia" county="River" municipality="Bureya River" name="Kulikovka village" pageId="6" pageNumber="387" stateProvince="Amur">Kulikovka village</location>
,
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adults,
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. Tiunova
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="57B96510FFAC0E15FF72FE41CDEEFC98" pageId="6" pageNumber="387" type="distribution">
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<emphasis id="2DD7EA89FFAC0E15FF72FE41CD3FFE29" box="[199,346,412,439]" italics="true" pageId="6" pageNumber="387">
<emphasis id="2DD7EA89FFAC0E15FF72FE41CD36FE28" bold="true" box="[199,339,412,439]" italics="true" pageId="6" pageNumber="387">Distribution</emphasis>
.
</emphasis>
East Siberia, Far East
<collectingCountry id="67B4760BFFAC0E15FDECFE41CEC2FE29" box="[601,679,412,438]" name="Russia" pageId="6" pageNumber="387">Russia</collectingCountry>
,
<collectingCountry id="67B4760BFFAC0E15FD07FE41CF47FE29" box="[690,802,412,438]" name="Mongolia" pageId="6" pageNumber="387">Mongolia</collectingCountry>
,
<collectingCountry id="67B4760BFFAC0E15FC98FE40CF11FE28" box="[813,884,413,439]" name="South Korea" pageId="6" pageNumber="387">Korea</collectingCountry>
,
<collectingCountry id="67B4760BFFAC0E15FCCAFE41CFA0FE29" box="[895,965,412,438]" name="China" pageId="6" pageNumber="387">China</collectingCountry>
,
<collectingCountry id="67B4760BFFAC0E15FC65FE41C836FE29" box="[976,1107,412,438]" name="Kazakhstan" pageId="6" pageNumber="387">Kazakhstan</collectingCountry>
(Irtysh River basin).
</paragraph>
<paragraph id="1F1C369BFFAC0E15FF72FE1FCDEEFC98" blockId="6.[151,1437,263,775]" pageId="6" pageNumber="387">
The egg has an oval shape close to quadrangular with rounded corners (
<figureCitation id="87982A1EFFAC0E15FC47FE1FC852FE43" box="[1010,1079,450,476]" captionStart="FIGURES 911" captionStartId="4.[151,264,1898,1922]" captionTargetBox="[265,1322,151,1889]" captionTargetId="figure-18@4.[265,1322,151,1889]" captionTargetPageId="4" captionText="FIGURES 911. Egg morphology of Ephemera (Ephemera) sachalinensis: 9, general shape; 1011, micropyle in the complex extrachorion-adhesive layer. AL-Ex—complex adhesive-extrachorion layer; MO—micropylar opening; mp—micropyle; CM—canal micropylar; SG— sperm guide." pageId="6" pageNumber="387">Fig. 9</figureCitation>
). Dimensions: 204.0230.0 µm in length (213.5 µm) and 118.0132.0 µm in width (125.8 µm). The thickness of the extrachorion-adhesive layer is 1.93.0 µm (
<figureCitation id="87982A1EFFAC0E15FEF6FDD1CDF2FDB8" box="[323,407,524,551]" captionStart="FIGURES 1215" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[204,1384,181,1919]" captionTargetId="figure-16@5.[204,1384,181,1919]" captionTargetPageId="5" captionText="FIGURES 1215. Egg morphology of Ephemera (Ephemera) sachalinensis: 1213, egg surface without complex extrachorionadhesive layer; 14, proximal part of micropylar canal on chorion surface; 15, thickness on the layer forming the eggshell. AL-Ex—complex adhesive-extrachorion layer; CS—chorion surface; MO—micropylar opening; mp—micropyle." pageId="6" pageNumber="387">Fig. 15</figureCitation>
). The surface of the adhesive layer is shagreen or roughened (
<figureCitation id="87982A1EFFAC0E15FBE0FDD1C8FFFDB8" box="[1109,1178,524,551]" captionStart="FIGURES 911" captionStartId="4.[151,264,1898,1922]" captionTargetBox="[265,1322,151,1889]" captionTargetId="figure-18@4.[265,1322,151,1889]" captionTargetPageId="4" captionText="FIGURES 911. Egg morphology of Ephemera (Ephemera) sachalinensis: 9, general shape; 1011, micropyle in the complex extrachorion-adhesive layer. AL-Ex—complex adhesive-extrachorion layer; MO—micropylar opening; mp—micropyle; CM—canal micropylar; SG— sperm guide." pageId="6" pageNumber="387">Fig. 9</figureCitation>
). There are one or two micropyles per egg in the equatorial area (
<figureCitation id="87982A1EFFAC0E15FDC5FDEFCEDCFDD3" box="[624,697,562,588]" captionStart="FIGURES 911" captionStartId="4.[151,264,1898,1922]" captionTargetBox="[265,1322,151,1889]" captionTargetId="figure-18@4.[265,1322,151,1889]" captionTargetPageId="4" captionText="FIGURES 911. Egg morphology of Ephemera (Ephemera) sachalinensis: 9, general shape; 1011, micropyle in the complex extrachorion-adhesive layer. AL-Ex—complex adhesive-extrachorion layer; MO—micropylar opening; mp—micropyle; CM—canal micropylar; SG— sperm guide." pageId="6" pageNumber="387">Figs 9</figureCitation>
,
<figureCitation id="87982A1EFFAC0E15FD70FDEFCE81FDD3" box="[709,740,562,588]" captionStart="FIGURES 1215" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[204,1384,181,1919]" captionTargetId="figure-16@5.[204,1384,181,1919]" captionTargetPageId="5" captionText="FIGURES 1215. Egg morphology of Ephemera (Ephemera) sachalinensis: 1213, egg surface without complex extrachorionadhesive layer; 14, proximal part of micropylar canal on chorion surface; 15, thickness on the layer forming the eggshell. AL-Ex—complex adhesive-extrachorion layer; CS—chorion surface; MO—micropylar opening; mp—micropyle." pageId="6" pageNumber="387">13</figureCitation>
). Micropyles are “tagenoform-type” (
<figureCitation id="87982A1EFFAC0E15FB3FFDEFC89AFDD3" box="[1162,1279,562,588]" captionStart="FIGURES 911" captionStartId="4.[151,264,1898,1922]" captionTargetBox="[265,1322,151,1889]" captionTargetId="figure-18@4.[265,1322,151,1889]" captionTargetPageId="4" captionText="FIGURES 911. Egg morphology of Ephemera (Ephemera) sachalinensis: 9, general shape; 1011, micropyle in the complex extrachorion-adhesive layer. AL-Ex—complex adhesive-extrachorion layer; MO—micropylar opening; mp—micropyle; CM—canal micropylar; SG— sperm guide." pageId="6" pageNumber="387">Figs 910</figureCitation>
), sperm guide weakly expressed (6.49.7 µm long, 3.05.0 µm wide), almost rectangular, elongated, distal margin not closed (
<figureCitation id="87982A1EFFAC0E15FADEFD8ACC85FD08" captionStart="FIGURES 911" captionStartId="4.[151,264,1898,1922]" captionTargetBox="[265,1322,151,1889]" captionTargetId="figure-18@4.[265,1322,151,1889]" captionTargetPageId="4" captionText="FIGURES 911. Egg morphology of Ephemera (Ephemera) sachalinensis: 9, general shape; 1011, micropyle in the complex extrachorion-adhesive layer. AL-Ex—complex adhesive-extrachorion layer; MO—micropylar opening; mp—micropyle; CM—canal micropylar; SG— sperm guide." pageId="6" pageNumber="387">Figs 1011</figureCitation>
). The micropylar canal, about 5 μm long, does not protrude above the adhesive layer (
<figureCitation id="87982A1EFFAC0E15FB11FDA1C97CFD08" box="[1188,1305,636,663]" captionStart="FIGURES 911" captionStartId="4.[151,264,1898,1922]" captionTargetBox="[265,1322,151,1889]" captionTargetId="figure-18@4.[265,1322,151,1889]" captionTargetPageId="4" captionText="FIGURES 911. Egg morphology of Ephemera (Ephemera) sachalinensis: 9, general shape; 1011, micropyle in the complex extrachorion-adhesive layer. AL-Ex—complex adhesive-extrachorion layer; MO—micropylar opening; mp—micropyle; CM—canal micropylar; SG— sperm guide." pageId="6" pageNumber="387">Figs 910</figureCitation>
). Chorionic sculpturing consists of interrupted broken ridges, whose distribution and arrangement extend regularly over the whole chorion surface (
<figureCitation id="87982A1EFFAC0E15FE2BFD1ACE44FD7E" box="[414,545,711,737]" captionStart="FIGURES 1215" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[204,1384,181,1919]" captionTargetId="figure-16@5.[204,1384,181,1919]" captionTargetPageId="5" captionText="FIGURES 1215. Egg morphology of Ephemera (Ephemera) sachalinensis: 1213, egg surface without complex extrachorionadhesive layer; 14, proximal part of micropylar canal on chorion surface; 15, thickness on the layer forming the eggshell. AL-Ex—complex adhesive-extrachorion layer; CS—chorion surface; MO—micropylar opening; mp—micropyle." pageId="6" pageNumber="387">Figs 1213</figureCitation>
). The proximal part of the micropylar opening (MO) does not elevate above the chorion (
<figureCitation id="87982A1EFFAC0E15FF49FD31CD1BFC98" box="[252,382,748,775]" captionStart="FIGURES 1215" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[204,1384,181,1919]" captionTargetId="figure-16@5.[204,1384,181,1919]" captionTargetPageId="5" captionText="FIGURES 1215. Egg morphology of Ephemera (Ephemera) sachalinensis: 1213, egg surface without complex extrachorionadhesive layer; 14, proximal part of micropylar canal on chorion surface; 15, thickness on the layer forming the eggshell. AL-Ex—complex adhesive-extrachorion layer; CS—chorion surface; MO—micropylar opening; mp—micropyle." pageId="6" pageNumber="387">Figs 1314</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,246 @@
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<taxonomicName id="D8A34D18FFAC0E15FF22FA09CF7AFA71" authority="McLachlan, 1875" authorityName="McLachlan" authorityYear="1875" box="[151,799,1492,1519]" class="Insecta" family="Ephemeridae" genus="Ephemera" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="387" phylum="Arthropoda" rank="species" species="japonica" subGenus="Sinephemera">
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<emphasis id="2DD7EA89FFAC0E15FF22FA09CD71FA70" bold="true" box="[151,276,1492,1519]" italics="true" pageId="6" pageNumber="387">Ephemera</emphasis>
(
<emphasis id="2DD7EA89FFAC0E15FE93FA09CDA3FA70" bold="true" box="[294,454,1492,1519]" italics="true" pageId="6" pageNumber="387">Sinephemera</emphasis>
)
<emphasis id="2DD7EA89FFAC0E15FE62FA09CE5AFA71" bold="true" box="[471,575,1492,1518]" italics="true" pageId="6" pageNumber="387">japonica</emphasis>
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<subSubSection id="57B96510FFAC0E15FF22FA27CD5CF98B" box="[151,313,1530,1556]" pageId="6" pageNumber="387" type="description">
<paragraph id="1F1C369BFFAC0E15FF22FA27CD5CF98B" blockId="6.[151,799,1492,1556]" box="[151,313,1530,1556]" pageId="6" pageNumber="387">
<figureCitation id="87982A1EFFAC0E15FF22FA27CD5CF98B" box="[151,313,1530,1556]" captionStart-0="FIGURES 2426" captionStart-1="FIGURES 2730" captionStartId-0="9.[151,264,1943,1967]" captionStartId-1="10.[151,264,1943,1967]" captionTargetBox-0="[282,1306,181,1919]" captionTargetBox-1="[277,1310,181,1919]" captionTargetId-0="figure-16@9.[282,1306,181,1919]" captionTargetId-1="figure-18@10.[277,1310,181,1919]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURES 2426. Egg morphology of Ephemera (Sinephemera) japonica: 2425, remains of the adhesive layer on the egg surface; 26, chorion thickness. AL-Ex—complex adhesive-extrachorion layer; AL—adhesive layer; C—chorion." captionText-1="FIGURES 2730. Egg morphology of Ephemera (Sinephemera) japonica: 2728, egg shapes; 29, micropyle in the complex extrachorion-adhesive layer; 30, proximal part of micropylar canal on chorion surface. AL—adhesive layer; CM—canal micropylar; MO—micropylar opening; CS—chorion surface; mp—micropyle; SG—sperm guide." pageId="6" pageNumber="387">Figures 2430</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="57B96510FFAC0E15FF22F999CE7BF96B" pageId="6" pageNumber="387" type="materials_examined">
<paragraph id="1F1C369BFFAC0E15FF22F999CE7BF96B" blockId="6.[151,1437,1604,2005]" pageId="6" pageNumber="387">
<emphasis id="2DD7EA89FFAC0E15FF22F999CD1BF9C1" bold="true" box="[151,382,1604,1630]" pageId="6" pageNumber="387">Material examined</emphasis>
.
<materialsCitation id="AFCB3CC6FFAC0E15FE25F999CE7FF96B" collectingDate="1994-04-08" collectingDateMax="2001-07-21" collectingDateMin="1994-04-08" collectionCode="V, T" country="Russia" county="Sakhalinskaya Oblast" location="Belaya River" municipality="Island" pageId="6" pageNumber="387" specimenCount="8" specimenCount-female="8" stateProvince="Sakhalin">
<collectingCountry id="67B4760BFFAC0E15FE25F999CDBAF9C1" box="[400,479,1604,1630]" name="Russia" pageId="6" pageNumber="387">Russia</collectingCountry>
:
<collectingCounty id="F67D4E17FFAC0E15FE5BF999CE8DF9C1" box="[494,744,1604,1630]" pageId="6" pageNumber="387">Sakhalinskaya Oblast</collectingCounty>
:
<collectingRegion id="DD67F879FFAC0E15FD42F999CF3CF9C1" box="[759,857,1604,1630]" country="Russia" name="Sakhalin" pageId="6" pageNumber="387">Sakhalin</collectingRegion>
<collectingMunicipality id="FF78ACE1FFAC0E15FCD1F999CFCEF9C1" box="[868,939,1604,1630]" pageId="6" pageNumber="387">Island</collectingMunicipality>
,
<location id="1A7C6040FFAC0E15FC0EF999C831F9C1" LSID="urn:lsid:plazi:treatment:970A878DFFAC0E18FF22FA09CE36FD9E:1A7C6040FFAC0E15FC0EF999C831F9C1" box="[955,1108,1604,1630]" country="Russia" county="Sakhalinskaya Oblast" municipality="Island" name="Belaya River" pageId="6" pageNumber="387" stateProvince="Sakhalin">Belaya River</location>
,
<quantity id="D85B9B7EFFAC0E15FBD6F998C8C4F9C1" box="[1123,1185,1604,1631]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="387" unit="km" value="2.0">2 km</quantity>
southeast of “Sokol” station,
<date id="6B1D105BFFAC0E15FF47F9B7CD16F91B" box="[242,371,1642,1668]" pageId="6" pageNumber="387" value="2001-07-21">
<collectingDate id="7B59E9B3FFAC0E15FF47F9B7CD16F91B" box="[242,371,1642,1668]" pageId="6" pageNumber="387" value="2001-07-21">21.07.2001</collectingDate>
</date>
,
<specimenCount id="09A5FD12FFAC0E15FE35F9B7CDC6F91B" box="[384,419,1642,1668]" count="1" pageId="6" pageNumber="387" type="female">1♀</specimenCount>
adult,
<collectionCode id="79B2AE5EFFAC0E15FE46F9B7CE62F91B" box="[499,519,1642,1668]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="387" type="Museum">V</collectionCode>
. Teslenko;
<collectingRegion id="DD67F879FFAC0E15FD3CF9B7CE8EF91B" box="[649,747,1642,1668]" country="Russia" name="Sakhalin" pageId="6" pageNumber="387">Sakhalin</collectingRegion>
Island, unnamed stream,
<quantity id="D85B9B7EFFAC0E15FBA6F9B7C839F91B" box="[1043,1116,1642,1669]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="6" pageNumber="387" unit="km" value="25.0">25 km</quantity>
south of
<location id="1A7C6040FFAC0E15FB7FF9B7C9FDF91B" LSID="urn:lsid:plazi:treatment:970A878DFFAC0E18FF22FA09CE36FD9E:1A7C6040FFAC0E15FB7FF9B7C9FDF91B" box="[1226,1432,1642,1668]" country="Russia" county="Sakhalinskaya Oblast" municipality="Island" name="Vostochny village" pageId="6" pageNumber="387" stateProvince="Sakhalin">Vostochny village</location>
,
<date id="6B1D105BFFAC0E15FF22F952CD7DF936" box="[151,280,1679,1705]" pageId="6" pageNumber="387" value="2001-01-08">
<collectingDate id="7B59E9B3FFAC0E15FF22F952CD7DF936" box="[151,280,1679,1705]" pageId="6" pageNumber="387" value="2001-01-08">01.08.2001</collectingDate>
</date>
,
<specimenCount id="09A5FD12FFAC0E15FE93F952CD2CF936" box="[294,329,1679,1705]" count="1" pageId="6" pageNumber="387" type="female">1♀</specimenCount>
adult,
<collectionCode id="79B2AE5EFFAC0E15FE29F952CDD5F936" box="[412,432,1679,1705]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="387" type="Museum">V</collectionCode>
.Teslenko;
<location id="1A7C6040FFAC0E15FD9FF94DCEA5F936" LSID="urn:lsid:plazi:treatment:970A878DFFAC0E18FF22FA09CE36FD9E:1A7C6040FFAC0E15FD9FF94DCEA5F936" box="[554,704,1679,1706]" country="Russia" county="Sakhalinskaya Oblast" municipality="Island" name="Iturup Island" pageId="6" pageNumber="387" stateProvince="Sakhalin">Iturup Island</location>
,
<location id="1A7C6040FFAC0E15FD7BF952CF05F935" LSID="urn:lsid:plazi:treatment:970A878DFFAC0E18FF22FA09CE36FD9E:1A7C6040FFAC0E15FD7BF952CF05F935" box="[718,864,1679,1706]" country="Russia" county="Sakhalinskaya Oblast" municipality="Island" name="Ketoviy Bay" pageId="6" pageNumber="387" stateProvince="Sakhalin">Ketoviy Bay</location>
,
<location id="1A7C6040FFAC0E15FCDAF952C825F936" LSID="urn:lsid:plazi:treatment:970A878DFFAC0E18FF22FA09CE36FD9E:1A7C6040FFAC0E15FCDAF952C825F936" box="[879,1088,1679,1705]" country="Russia" county="Sakhalinskaya Oblast" municipality="Island" name="Podoshevka River" pageId="6" pageNumber="387" stateProvince="Sakhalin">Podoshevka River</location>
,
<locationDeviation id="0EAE2FB9FFAC0E15FBFBF952C9FDF936" box="[1102,1432,1679,1706]" location="Fish Hatchery" pageId="6" pageNumber="387">
<quantity id="D85B9B7EFFAC0E15FBFBF952C8FAF936" box="[1102,1183,1679,1706]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="387" unit="km" value="1.5">1.5 km</quantity>
above Fish Hatchery
</locationDeviation>
,
<date id="6B1D105BFFAC0E15FF22F968CD72F950" box="[151,279,1717,1743]" pageId="6" pageNumber="387" value="1997-07-29">
<collectingDate id="7B59E9B3FFAC0E15FF22F968CD72F950" box="[151,279,1717,1743]" pageId="6" pageNumber="387" value="1997-07-29">29.07.1997</collectingDate>
</date>
,
<specimenCount id="09A5FD12FFAC0E15FE97F968CD20F950" box="[290,325,1717,1743]" count="4" pageId="6" pageNumber="387" type="female">4♀</specimenCount>
adults,
<collectionCode id="79B2AE5EFFAC0E15FE2EF968CDCAF950" box="[411,431,1717,1743]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="387" type="Museum">V</collectionCode>
. Teslenko;
<location id="1A7C6040FFAC0E15FD9EF969CEBBF951" LSID="urn:lsid:plazi:treatment:970A878DFFAC0E18FF22FA09CE36FD9E:1A7C6040FFAC0E15FD9EF969CEBBF951" box="[555,734,1716,1742]" country="Russia" county="Sakhalinskaya Oblast" municipality="Island" name="Kunashir Island" pageId="6" pageNumber="387" stateProvince="Sakhalin">Kunashir Island</location>
,
<location id="1A7C6040FFAC0E15FD5CF968CFE9F951" LSID="urn:lsid:plazi:treatment:970A878DFFAC0E18FF22FA09CE36FD9E:1A7C6040FFAC0E15FD5CF968CFE9F951" box="[745,908,1716,1743]" country="Russia" county="Sakhalinskaya Oblast" municipality="Island" name="Lesnaya River" pageId="6" pageNumber="387" stateProvince="Sakhalin">Lesnaya River</location>
, about
<quantity id="D85B9B7EFFAC0E15FC6EF968C876F951" box="[987,1043,1716,1743]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="387" unit="km" value="1.0">1 km</quantity>
above the mouth of Kislyy Stream,
<date id="6B1D105BFFAC0E15FF22F907CD7DF96B" box="[151,280,1754,1780]" pageId="6" pageNumber="387" value="1994-04-08">
<collectingDate id="7B59E9B3FFAC0E15FF22F907CD7DF96B" box="[151,280,1754,1780]" pageId="6" pageNumber="387" value="1994-04-08">04.08.1994</collectingDate>
</date>
,
<specimenCount id="09A5FD12FFAC0E15FE96F907CD23F96B" box="[291,326,1754,1780]" count="2" pageId="6" pageNumber="387" type="female">2♀</specimenCount>
adults,
<collectionCode id="79B2AE5EFFAC0E15FE2BF907CDD5F96B" box="[414,432,1754,1780]" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="387">T</collectionCode>
. Tiunova
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="57B96510FFAC0E18FF72F922CE36FD9E" lastPageId="11" lastPageNumber="392" pageId="6" pageNumber="387" type="distribution">
<paragraph id="1F1C369BFFAC0E15FF72F922CED3F886" blockId="6.[151,1437,1604,2005]" box="[199,694,1791,1818]" pageId="6" pageNumber="387">
<emphasis id="2DD7EA89FFAC0E15FF72F922CD3FF885" bold="true" box="[199,346,1791,1818]" italics="true" pageId="6" pageNumber="387">Distribution.</emphasis>
Far East
<collectingCountry id="67B4760BFFAC0E15FE70F922CE76F886" box="[453,531,1791,1817]" name="Russia" pageId="6" pageNumber="387">Russia</collectingCountry>
,
<collectingCountry id="67B4760BFFAC0E15FDA8F8DDCE05F885" box="[541,608,1792,1818]" name="Japan" pageId="6" pageNumber="387">Japan</collectingCountry>
,
<collectingCountry id="67B4760BFFAC0E15FDDEF922CED7F886" box="[619,690,1791,1817]" name="China" pageId="6" pageNumber="387">China</collectingCountry>
.
</paragraph>
<paragraph id="1F1C369BFFAC0E15FF72F8F9C839F84B" blockId="6.[151,1437,1604,2005]" pageId="6" pageNumber="387">
The egg was described previously by
<bibRefCitation id="7B324B6AFFAC0E15FDDBF8F8CFE0F8A0" author="Koss, R. W. &amp; Edmunds, G. F. Jr." box="[622,901,1828,1855]" pageId="6" pageNumber="387" pagination="267 - 349" refId="ref7161" refString="Koss, R. W. &amp; Edmunds, G. F. Jr. (1974) Ephemeroptera eggs and their contribution to phylogenetic studies of the order. Zoological Journal of the Linnean Society, 55, 267 - 349. https: // doi. org / 10.1111 / j. 1096 - 3642.1974. tb 01648. x" type="journal article" year="1974">Koss &amp; Edmunds (1974)</bibRefCitation>
and
<bibRefCitation id="7B324B6AFFAC0E15FC0EF8F9C8DAF8A0" author="Tojo, K. &amp; Machida, R." box="[955,1215,1828,1855]" pageId="6" pageNumber="387" pagination="573 - 579" refId="ref7457" refString="Tojo, K. &amp; Machida, R. (1998) Egg Structures of Japanes Ephemeridae species (Ephemeroptera). Entomological Science, 1 (4), 573 - 579." type="journal article" year="1998">Tojo &amp; Machida (1998</bibRefCitation>
, p. 574: fig. 1). The description notes: the shape of the egg is ellipsoidal, about 200x100 µm; the adhesive layer is very thin (about 0.1 µm); the surface of the chorion has an ill-developed reticulation (by
<bibRefCitation id="7B324B6AFFAC0E15FC2CF8B2C8F7F816" author="Tojo, K. &amp; Machida, R." box="[921,1170,1903,1929]" pageId="6" pageNumber="387" pagination="573 - 579" refId="ref7457" refString="Tojo, K. &amp; Machida, R. (1998) Egg Structures of Japanes Ephemeridae species (Ephemeroptera). Entomological Science, 1 (4), 573 - 579." type="journal article" year="1998">Tojo &amp; Machida 1998</bibRefCitation>
) or smooth (by
<bibRefCitation id="7B324B6AFFAC0E15FAF2F8ADCD21F830" author="Koss, R. W. &amp; Edmunds, G. F. Jr." pageId="6" pageNumber="387" pagination="267 - 349" refId="ref7161" refString="Koss, R. W. &amp; Edmunds, G. F. Jr. (1974) Ephemeroptera eggs and their contribution to phylogenetic studies of the order. Zoological Journal of the Linnean Society, 55, 267 - 349. https: // doi. org / 10.1111 / j. 1096 - 3642.1974. tb 01648. x" type="journal article" year="1974">Koss &amp; Edmunds 1974</bibRefCitation>
); micropyle, one per egg; the sperm guide is undeveloped; micropylar canal according to Koss and Edmunds (elongated, 4255 µm long); according to Tojo and Machida (about 20 µm).
</paragraph>
<caption id="4BDC6613FFAD0E14FF22F84AC885F867" pageId="7" pageNumber="388" startId="7.[151,264,1943,1967]" targetBox="[219,1369,181,1919]" targetPageId="7" targetType="figure">
<paragraph id="1F1C369BFFAD0E14FF22F84ACE70F84B" blockId="7.[151,1436,1943,2040]" pageId="7" pageNumber="388">
<emphasis id="2DD7EA89FFAD0E14FF22F84ACD30F830" bold="true" box="[151,341,1943,1967]" pageId="7" pageNumber="388">FIGURES 1619.</emphasis>
Egg morphology of
<taxonomicName id="D8A34D18FFAD0E14FD98F845CFD5F82F" authorityName="Tshernova" authorityYear="1973" box="[557,944,1944,1968]" class="Insecta" family="Ephemeridae" genus="Ephemera" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="388" phylum="Arthropoda" rank="species" species="transbajkalica" subGenus="Ephemera">
<emphasis id="2DD7EA89FFAD0E14FD98F845CEF0F82F" box="[557,661,1944,1968]" italics="true" pageId="7" pageNumber="388">Ephemera</emphasis>
(
<emphasis id="2DD7EA89FFAD0E14FD10F845CF68F82F" box="[677,781,1944,1968]" italics="true" pageId="7" pageNumber="388">Ephemera</emphasis>
)
<emphasis id="2DD7EA89FFAD0E14FCABF845CFD5F82F" box="[798,944,1944,1968]" italics="true" pageId="7" pageNumber="388">transbajkalica</emphasis>
</taxonomicName>
: 1617, general shape; 1819, micropyle in the complex extrachorion-adhesive layer.
</paragraph>
<paragraph id="1F1C369BFFAD0E14FF22F83DC885F867" blockId="7.[151,1436,1943,2040]" box="[151,1248,2016,2040]" pageId="7" pageNumber="388">AL-Ex—complex adhesive-extrachorion layer; CM—canal micropylar; mp—micropyle; SG—sperm guide.</paragraph>
</caption>
<caption id="4BDC6613FFA20E1BFF22F8ACC978F869" pageId="8" pageNumber="389" startId="8.[151,264,1905,1929]" targetBox="[287,1299,151,1889]" targetPageId="8" targetType="figure">
<paragraph id="1F1C369BFFA20E1BFF22F8ACCD12F84D" blockId="8.[151,1436,1905,2038]" pageId="8" pageNumber="389">
<emphasis id="2DD7EA89FFA20E1BFF22F8ACCD30F815" bold="true" box="[151,341,1905,1930]" pageId="8" pageNumber="389">FIGURES 2023.</emphasis>
Egg morphology of
<taxonomicName id="D8A34D18FFA20E1BFD99F8AFCFCBF815" authorityName="Tshernova" authorityYear="1973" box="[556,942,1906,1930]" class="Insecta" family="Ephemeridae" genus="Ephemera" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="389" phylum="Arthropoda" rank="species" species="transbajkalica" subGenus="Ephemera">
<emphasis id="2DD7EA89FFA20E1BFD99F8AFCEF1F815" box="[556,660,1906,1930]" italics="true" pageId="8" pageNumber="389">Ephemera</emphasis>
(
<emphasis id="2DD7EA89FFA20E1BFD11F8AFCF69F815" box="[676,780,1906,1930]" italics="true" pageId="8" pageNumber="389">Ephemera</emphasis>
)
<emphasis id="2DD7EA89FFA20E1BFCA9F8AFCFCBF815" box="[796,942,1906,1930]" italics="true" pageId="8" pageNumber="389">transbajkalica</emphasis>
</taxonomicName>
: 20, egg surface without complex extrachorionadhesive layer; 21, thickness on the layer forming the eggshell; 22, proximal part of micropylar canal on chorion surface; 23, chorionic sculpturing.
</paragraph>
<paragraph id="1F1C369BFFA20E1BFF22F803C978F869" blockId="8.[151,1436,1905,2038]" box="[151,1309,2014,2038]" pageId="8" pageNumber="389">AL-Ex—complex adhesive-extrachorion layer; CS—chorion surface; MO—micropylar opening; mp—micropyle.</paragraph>
</caption>
<caption id="4BDC6613FFA30E1AFF22F84ACFB1F867" pageId="9" pageNumber="390" startId="9.[151,264,1943,1967]" targetBox="[282,1306,181,1919]" targetPageId="9" targetType="figure">
<paragraph id="1F1C369BFFA30E1AFF22F84ACDA8F84B" blockId="9.[151,1437,1943,2040]" pageId="9" pageNumber="390">
<emphasis id="2DD7EA89FFA30E1AFF22F84ACD33F830" bold="true" box="[151,342,1943,1967]" pageId="9" pageNumber="390">FIGURES 2426.</emphasis>
Egg morphology of
<taxonomicName id="D8A34D18FFA30E1AFD84F845CFFDF830" box="[561,920,1944,1968]" class="Insecta" family="Ephemeridae" genus="Ephemera" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="390" phylum="Arthropoda" rank="species" species="japonica" subGenus="Sinephemera">
<emphasis id="2DD7EA89FFA30E1AFD84F845CEFCF82F" box="[561,665,1944,1968]" italics="true" pageId="9" pageNumber="390">Ephemera</emphasis>
(
<emphasis id="2DD7EA89FFA30E1AFD1FF845CF4BF82F" box="[682,814,1944,1968]" italics="true" pageId="9" pageNumber="390">Sinephemera</emphasis>
)
<emphasis id="2DD7EA89FFA30E1AFC8AF845CFFDF830" box="[831,920,1944,1967]" italics="true" pageId="9" pageNumber="390">japonica</emphasis>
</taxonomicName>
: 2425, remains of the adhesive layer on the egg surface; 26, chorion thickness.
</paragraph>
<paragraph id="1F1C369BFFA30E1AFF22F83DCFB1F867" blockId="9.[151,1437,1943,2040]" box="[151,980,2016,2040]" pageId="9" pageNumber="390">AL-Ex—complex adhesive-extrachorion layer; AL—adhesive layer; C—chorion.</paragraph>
</caption>
<caption id="4BDC6613FFA00E19FF22F84AC9FCF867" pageId="10" pageNumber="391" startId="10.[151,264,1943,1967]" targetBox="[277,1310,181,1919]" targetPageId="10" targetType="figure">
<paragraph id="1F1C369BFFA00E19FF22F84AC860F84B" blockId="10.[151,1436,1943,2040]" pageId="10" pageNumber="391">
<emphasis id="2DD7EA89FFA00E19FF22F84ACD33F830" bold="true" box="[151,342,1943,1967]" pageId="10" pageNumber="391">FIGURES 2730.</emphasis>
Egg morphology of
<taxonomicName id="D8A34D18FFA00E19FD9BF845CFF6F830" box="[558,915,1944,1968]" class="Insecta" family="Ephemeridae" genus="Ephemera" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="391" phylum="Arthropoda" rank="species" species="japonica" subGenus="Sinephemera">
<emphasis id="2DD7EA89FFA00E19FD9BF845CEF3F82F" box="[558,662,1944,1968]" italics="true" pageId="10" pageNumber="391">Ephemera</emphasis>
(
<emphasis id="2DD7EA89FFA00E19FD13F845CF4FF82F" box="[678,810,1944,1968]" italics="true" pageId="10" pageNumber="391">Sinephemera</emphasis>
)
<emphasis id="2DD7EA89FFA00E19FC8FF845CFF6F830" box="[826,915,1944,1967]" italics="true" pageId="10" pageNumber="391">japonica</emphasis>
</taxonomicName>
: 2728, egg shapes; 29, micropyle in the complex extrachorion-adhesive layer; 30, proximal part of micropylar canal on chorion surface.
</paragraph>
<paragraph id="1F1C369BFFA00E19FF22F83DC9FCF867" blockId="10.[151,1436,1943,2040]" box="[151,1433,2016,2040]" pageId="10" pageNumber="391">AL—adhesive layer; CM—canal micropylar; MO—micropylar opening; CS—chorion surface; mp—micropyle; SG—sperm guide.</paragraph>
</caption>
<paragraph id="1F1C369BFFA10E18FF7BFF4ACE36FD9E" blockId="11.[151,1437,150,513]" pageId="11" pageNumber="392">
According to our data, the egg is oval, more often ovoid in shape (
<figureCitation id="87982A1EFFA10E18FC7FFF4AC828FF2E" box="[970,1101,150,177]" captionStart="FIGURES 2426" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[282,1306,181,1919]" captionTargetId="figure-16@9.[282,1306,181,1919]" captionTargetPageId="9" captionText="FIGURES 2426. Egg morphology of Ephemera (Sinephemera) japonica: 2425, remains of the adhesive layer on the egg surface; 26, chorion thickness. AL-Ex—complex adhesive-extrachorion layer; AL—adhesive layer; C—chorion." pageId="11" pageNumber="392">Figs 2425</figureCitation>
,
<figureCitation id="87982A1EFFA10E18FBEFFF4AC8C1FF2E" box="[1114,1188,151,177]" captionStart="FIGURES 2730" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[277,1310,181,1919]" captionTargetId="figure-18@10.[277,1310,181,1919]" captionTargetPageId="10" captionText="FIGURES 2730. Egg morphology of Ephemera (Sinephemera) japonica: 2728, egg shapes; 29, micropyle in the complex extrachorion-adhesive layer; 30, proximal part of micropylar canal on chorion surface. AL—adhesive layer; CM—canal micropylar; MO—micropylar opening; CS—chorion surface; mp—micropyle; SG—sperm guide." pageId="11" pageNumber="392">2728</figureCitation>
). Dimensions: 293.0 334.0 µm in length (315.4 µm) and 173197 µm in width (180.1 µm). The extrachorion-adhesive layer covering the egg is thin, smoothly amorphous (
<figureCitation id="87982A1EFFA10E18FDF7FF3CCEA7FF63" box="[578,706,225,252]" captionStart="FIGURES 2426" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[282,1306,181,1919]" captionTargetId="figure-16@9.[282,1306,181,1919]" captionTargetPageId="9" captionText="FIGURES 2426. Egg morphology of Ephemera (Sinephemera) japonica: 2425, remains of the adhesive layer on the egg surface; 26, chorion thickness. AL-Ex—complex adhesive-extrachorion layer; AL—adhesive layer; C—chorion." pageId="11" pageNumber="392">Fig. 2425</figureCitation>
). In the equatorial area, there is one micropyle per egg (
<figureCitation id="87982A1EFFA10E18FA8EFF3CC9F5FF64" box="[1339,1424,225,251]" captionStart="FIGURES 2730" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[277,1310,181,1919]" captionTargetId="figure-18@10.[277,1310,181,1919]" captionTargetPageId="10" captionText="FIGURES 2730. Egg morphology of Ephemera (Sinephemera) japonica: 2728, egg shapes; 29, micropyle in the complex extrachorion-adhesive layer; 30, proximal part of micropylar canal on chorion surface. AL—adhesive layer; CM—canal micropylar; MO—micropylar opening; CS—chorion surface; mp—micropyle; SG—sperm guide." pageId="11" pageNumber="392">Fig. 27</figureCitation>
). The micropyle is of the “tagenoform
<typeStatus id="C0188839FFA10E18FD8DFEDACE08FEBE" box="[568,621,263,289]" pageId="11" pageNumber="392">type</typeStatus>
,” with a sharply expanding, rounded, poorly defined sperm guide, 1416 μm long and 2124 μm wide (
<figureCitation id="87982A1EFFA10E18FE42FEF1CE1BFED9" box="[503,638,300,326]" captionStart="FIGURES 2730" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[277,1310,181,1919]" captionTargetId="figure-18@10.[277,1310,181,1919]" captionTargetPageId="10" captionText="FIGURES 2730. Egg morphology of Ephemera (Sinephemera) japonica: 2728, egg shapes; 29, micropyle in the complex extrachorion-adhesive layer; 30, proximal part of micropylar canal on chorion surface. AL—adhesive layer; CM—canal micropylar; MO—micropylar opening; CS—chorion surface; mp—micropyle; SG—sperm guide." pageId="11" pageNumber="392">Figs 27, 29</figureCitation>
). As in other species, the sperm guide is visible only on the adhesive surface of the egg. After the removal of the adhesive layer, only the micropylar canal is visible (
<figureCitation id="87982A1EFFA10E18FB6DFE8CC938FEF4" box="[1240,1373,337,363]" captionStart="FIGURES 2730" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[277,1310,181,1919]" captionTargetId="figure-18@10.[277,1310,181,1919]" captionTargetPageId="10" captionText="FIGURES 2730. Egg morphology of Ephemera (Sinephemera) japonica: 2728, egg shapes; 29, micropyle in the complex extrachorion-adhesive layer; 30, proximal part of micropylar canal on chorion surface. AL—adhesive layer; CM—canal micropylar; MO—micropylar opening; CS—chorion surface; mp—micropyle; SG—sperm guide." pageId="11" pageNumber="392">Figs 28, 30</figureCitation>
). The micropylar canal is 1630 μm long and 38 μm wide, protruding prominently above the adhesive layer and chorion (
<figureCitation id="87982A1EFFA10E18FF2AFE41CD46FE29" box="[159,291,412,438]" captionStart="FIGURES 2730" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[277,1310,181,1919]" captionTargetId="figure-18@10.[277,1310,181,1919]" captionTargetPageId="10" captionText="FIGURES 2730. Egg morphology of Ephemera (Sinephemera) japonica: 2728, egg shapes; 29, micropyle in the complex extrachorion-adhesive layer; 30, proximal part of micropylar canal on chorion surface. AL—adhesive layer; CM—canal micropylar; MO—micropylar opening; CS—chorion surface; mp—micropyle; SG—sperm guide." pageId="11" pageNumber="392">Figs 2930</figureCitation>
). The surface of the chorion is smooth (
<figureCitation id="87982A1EFFA10E18FD68FE41CF55FE29" box="[733,816,412,438]" captionStart="FIGURES 2730" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[277,1310,181,1919]" captionTargetId="figure-18@10.[277,1310,181,1919]" captionTargetPageId="10" captionText="FIGURES 2730. Egg morphology of Ephemera (Sinephemera) japonica: 2728, egg shapes; 29, micropyle in the complex extrachorion-adhesive layer; 30, proximal part of micropylar canal on chorion surface. AL—adhesive layer; CM—canal micropylar; MO—micropylar opening; CS—chorion surface; mp—micropyle; SG—sperm guide." pageId="11" pageNumber="392">Fig. 28</figureCitation>
), or with a weakly expressed very fine reticulation, and the proximal part of the tunnel-type micropylar canal and a micropylar opening 23 μm wide are clearly visible on the surface of the chorion (
<figureCitation id="87982A1EFFA10E18FE71FE3ACE22FD9E" box="[452,583,487,513]" captionStart="FIGURES 2730" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[277,1310,181,1919]" captionTargetId="figure-18@10.[277,1310,181,1919]" captionTargetPageId="10" captionText="FIGURES 2730. Egg morphology of Ephemera (Sinephemera) japonica: 2728, egg shapes; 29, micropyle in the complex extrachorion-adhesive layer; 30, proximal part of micropylar canal on chorion surface. AL—adhesive layer; CM—canal micropylar; MO—micropylar opening; CS—chorion surface; mp—micropyle; SG—sperm guide." pageId="11" pageNumber="392">Figs 28, 30</figureCitation>
).
</paragraph>
</subSubSection>
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Archaeoniscidae
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<emphasis id="06FDBF35FF87FF8CFDFAFBFDADADFBD4" bold="true" box="[598,645,1087,1113]" italics="true" pageId="7" pageNumber="416">non</emphasis>
<bibRefCitation id="50181ED6FF87FF8CFD21FBFDAC00FBD4" author="Haack, W." box="[653,808,1087,1113]" pageId="7" pageNumber="416" pagination="73 - 102" refId="ref10737" refString="Haack, W. (1918) Uber einen Isopoden aus dem Sepulit des weslichen Osnings (Archaeoniscus Brodei Milne-Edw.). Jahrbuch der Koniglichen Preussichen Geologischen Landesanstalt, 39, 73 - 102, pl. 4." type="journal article" year="1918">Haack, 1918</bibRefCitation>
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<emphasis id="06FDBF35FF87FF8CFF3BFB45AE2AFB2C" box="[151,258,1159,1185]" italics="true" pageId="7" pageNumber="416">Remarks:</emphasis>
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is, as far as can be determined, up to now always credited to
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but the name first appears in a table in
<bibRefCitation id="50181ED6FF87FF8CFE1BFB6EADF8FB4B" author="von Ammon, L." box="[439,720,1195,1222]" pageId="7" pageNumber="416" pagination="507 - 551" refId="ref14671" refString="von Ammon, L. (1882) Ein Beitrag zur Kenntniss der vorweltlichen Asseln. Sitzungsberichte der Mathematisch-Physikalischen Classe der Koniglich Bayrischen Akademie der Wissenschaften, 1882, 507 - 551, pls. 1 - 4." type="journal article" year="1882">von Ammon (1882: 546)</bibRefCitation>
as Archaeoniscinae within Cymothoïnae (
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, =
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). Authorship for this family-level taxon must therefore go to von Ammon.
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<mods:identifier id="D7EFDCED30120238803E33E0C9993DB7" type="Zenodo-Dep">13618434</mods:identifier>
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</mods:mods>
<treatment id="BC7A9D777A37FFC50ABBFF0C59ED035C" ID-DOI="http://doi.org/10.5281/zenodo.13618498" ID-Zenodo-Dep="13618498" LSID="urn:lsid:plazi:treatment:BC7A9D777A37FFC50ABBFF0C59ED035C" httpUri="http://treatment.plazi.org/id/BC7A9D777A37FFC50ABBFF0C59ED035C" lastPageId="7" lastPageNumber="283" pageId="2" pageNumber="278">
<subSubSection id="7CC97FEA7A37FFC00ABBFF0C5ADE00F8" box="[151,468,223,250]" pageId="2" pageNumber="278" type="nomenclature">
<paragraph id="346C2C617A37FFC00ABBFF0C5ADE00F8" blockId="2.[151,468,223,286]" box="[151,468,223,250]" pageId="2" pageNumber="278">
<heading id="6F249B0D7A37FFC00ABBFF0C5ADE00F8" bold="true" box="[151,468,223,250]" fontSize="11" level="1" pageId="2" pageNumber="278" reason="1">
<emphasis id="06A7F0737A37FFC00ABBFF0C5ADE00F8" bold="true" box="[151,468,223,250]" pageId="2" pageNumber="278">
<taxonomicName id="F3D357E27A37FFC00ABBFF0C5A6700FB" authorityName="Pinedo-Garcia &amp; Hamada &amp; Nascimento" authorityYear="2024" box="[151,365,223,250]" class="Insecta" family="Elmidae" genus="Hintonelmis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="278" phylum="Arthropoda" rank="species" species="spila" status="sp. nov.">
<emphasis id="06A7F0737A37FFC00ABBFF0C5A6700FB" bold="true" box="[151,365,223,250]" italics="true" pageId="2" pageNumber="278">Hintonelmis spila</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1D944D087A37FFC00B59FF0C5ADE00F8" box="[373,468,223,249]" pageId="2" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7CC97FEA7A37FFC00ABBFED05A01011F" box="[151,267,259,286]" pageId="2" pageNumber="278" type="description">
<paragraph id="346C2C617A37FFC00ABBFED05A01011F" blockId="2.[151,468,223,286]" box="[151,267,259,286]" pageId="2" pageNumber="278">
(
<figureCitation id="ACE830E47A37FFC00AB3FED05A08011F" box="[159,258,259,286]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="1.[151,250,1399,1423]" captionStartId-1="2.[151,250,1920,1944]" captionStartId-2="3.[151,250,1578,1602]" captionStartId-3="5.[151,250,1919,1943]" captionStartId-4="6.[151,250,1937,1961]" captionTargetBox-0="[176,1412,417,1375]" captionTargetBox-1="[204,1383,309,1897]" captionTargetBox-2="[203,1384,181,1554]" captionTargetBox-3="[277,1310,181,1894]" captionTargetBox-4="[269,1317,181,1905]" captionTargetId-0="figure-100@1.[176,1412,417,1375]" captionTargetId-1="figure-32@2.[204,1383,309,1897]" captionTargetId-2="figure-26@3.[203,1384,181,1554]" captionTargetId-3="figure-26@5.[277,1310,181,1894]" captionTargetId-4="figure-22@6.[269,1317,181,1913]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="5" captionTargetPageId-4="6" captionText-0="FIGURE 1. Map of South America with detailed area indicating the distributional records and general view of the sampled streams for Hintonelmis spila sp. nov. (Coleoptera: Elmidae)." captionText-1="FIGURE 2. Hintonelmis spila sp. nov. (Coleoptera: Elmidae). Adult male, habitus. (A) Dorsal view. (B) Ventral view. (C) Lateral view. (D) Front view. Scale bars = 0.5 mm." captionText-2="FIGURE 3. Hintonelmis spila sp. nov. (Coleoptera: Elmidae). Adult, mouthparts. (A) Labrum, dorsal view. (B) Maxilla, dorsal view. (C) Right mandible, dorsal view. (D) Right mandible, ventral view. (E) Labium, ventral view. Scale bars = 0.1 mm." captionText-3="FIGURE 4. Hintonelmis spila sp. nov. (Coleoptera: Elmidae).Adult, male. (A) Genitalia, ventral view. (B) Tergite VIII, ventral view. (C) Sternite VIII, dorsal view. (D). Segment IX, ventral view. (E) Metathoracic wing. Scale bars = 0.1 mm." captionText-4="FIGURE 5. Hintonelmis spila sp. nov. (Coleoptera: Elmidae). Adult, female. (A) Ovipositor, ventral view. (B) Tergite VIII, dorsal view, with the anterior region highlighted. (C) Sternite VIII, dorsal view. Scale bars = 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13618436" figureDoi-1="http://doi.org/10.5281/zenodo.13618438" figureDoi-2="http://doi.org/10.5281/zenodo.13618440" figureDoi-3="http://doi.org/10.5281/zenodo.13618442" figureDoi-4="http://doi.org/10.5281/zenodo.13618444" httpUri-0="https://zenodo.org/record/13618436/files/figure.png" httpUri-1="https://zenodo.org/record/13618438/files/figure.png" httpUri-2="https://zenodo.org/record/13618440/files/figure.png" httpUri-3="https://zenodo.org/record/13618442/files/figure.png" httpUri-4="https://zenodo.org/record/13618444/files/figure.png" pageId="2" pageNumber="278">Figs 15</figureCitation>
)
</paragraph>
</subSubSection>
<caption id="60AC7CE97A37FFC00ABBF853599607BD" ID-DOI="http://doi.org/10.5281/zenodo.13618438" ID-Zenodo-Dep="13618438" httpUri="https://zenodo.org/record/13618438/files/figure.png" pageId="2" pageNumber="278" startId="2.[151,250,1920,1944]" targetBox="[204,1383,309,1897]" targetPageId="2" targetType="figure">
<paragraph id="346C2C617A37FFC00ABBF853599607BD" blockId="2.[151,1436,1920,1981]" pageId="2" pageNumber="278">
<emphasis id="06A7F0737A37FFC00ABBF8535A1C0798" bold="true" box="[151,278,1920,1945]" pageId="2" pageNumber="278">FIGURE 2.</emphasis>
<taxonomicName id="F3D357E27A37FFC00B33F8525ADE0798" authorityName="Pinedo-Garcia &amp; Hamada &amp; Nascimento" authorityYear="2024" box="[287,468,1921,1945]" class="Insecta" family="Elmidae" genus="Hintonelmis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="278" phylum="Arthropoda" rank="species" species="spila" status="sp. nov.">
<emphasis id="06A7F0737A37FFC00B33F8525ADE0798" box="[287,468,1921,1945]" italics="true" pageId="2" pageNumber="278">Hintonelmis spila</emphasis>
</taxonomicName>
<emphasis id="06A7F0737A37FFC00BF2F852593B0798" bold="true" box="[478,561,1921,1945]" pageId="2" pageNumber="278">
<taxonomicNameLabel id="1D944D087A37FFC00BF2F852593B0798" box="[478,561,1921,1945]" pageId="2" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<taxonomicName id="F3D357E27A37FFC0086DF85259BC0798" authorityName="Linnaeus" authorityYear="1758" box="[577,694,1921,1945]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="278" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
:
<taxonomicName id="F3D357E27A37FFC008EEF85258110798" authorityName="Curtis" authorityYear="1830" box="[706,795,1921,1945]" class="Insecta" family="Elmidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="278" phylum="Arthropoda" rank="family">Elmidae</taxonomicName>
). Adult male, habitus. (A) Dorsal view. (B) Ventral view. (C) Lateral view. (D) Front view. Scale bars = 0.5 mm.
</paragraph>
</caption>
<caption id="60AC7CE97A36FFC10ABBF9F95E6E0664" ID-DOI="http://doi.org/10.5281/zenodo.13618440" ID-Zenodo-Dep="13618440" httpUri="https://zenodo.org/record/13618440/files/figure.png" pageId="3" pageNumber="279" startId="3.[151,250,1578,1602]" targetBox="[203,1384,181,1554]" targetPageId="3" targetType="figure">
<paragraph id="346C2C617A36FFC10ABBF9F95E6E0664" blockId="3.[151,1436,1578,1638]" pageId="3" pageNumber="279">
<emphasis id="06A7F0737A36FFC10ABBF9F95A180643" bold="true" box="[151,274,1578,1602]" pageId="3" pageNumber="279">FIGURE 3.</emphasis>
<taxonomicName id="F3D357E27A36FFC10B34F9F95AC10643" authorityName="Pinedo-Garcia &amp; Hamada &amp; Nascimento" authorityYear="2024" box="[280,459,1578,1602]" class="Insecta" family="Elmidae" genus="Hintonelmis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="279" phylum="Arthropoda" rank="species" species="spila" status="sp. nov.">
<emphasis id="06A7F0737A36FFC10B34F9F95AC10643" box="[280,459,1578,1602]" italics="true" pageId="3" pageNumber="279">Hintonelmis spila</emphasis>
</taxonomicName>
<emphasis id="06A7F0737A36FFC10BFDF9F9592B0643" bold="true" box="[465,545,1578,1602]" pageId="3" pageNumber="279">
<taxonomicNameLabel id="1D944D087A36FFC10BFDF9F9592B0643" box="[465,545,1578,1602]" pageId="3" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<taxonomicName id="F3D357E27A36FFC10802F9F959A90643" authorityName="Linnaeus" authorityYear="1758" box="[558,675,1578,1602]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="279" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
:
<taxonomicName id="F3D357E27A36FFC10880F9F958080643" authorityName="Curtis" authorityYear="1830" box="[684,770,1578,1602]" class="Insecta" family="Elmidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="279" phylum="Arthropoda" rank="family">Elmidae</taxonomicName>
). Adult, mouthparts. (A) Labrum, dorsal view. (B) Maxilla, dorsal view. (C) Right mandible, dorsal view. (D) Right mandible, ventral view. (E) Labium, ventral view. Scale bars = 0.1 mm.
</paragraph>
</caption>
<subSubSection id="7CC97FEA7A36FFC10ABBF9475F54073F" pageId="3" pageNumber="279" type="diagnosis">
<paragraph id="346C2C617A36FFC10ABBF9475F54073F" blockId="3.[151,1437,1684,2034]" pageId="3" pageNumber="279">
<emphasis id="06A7F0737A36FFC10ABBF9475A1806AF" bold="true" box="[151,274,1684,1710]" pageId="3" pageNumber="279">Diagnosis.</emphasis>
Body elongated; head with short yellow setae; antennal coloration uniform. Pronotum with a pair of short, strongly raised sublateral carinae along basal third. Elytra with yellowish-brown patches (one pair near the humeral region, the second pair shorter, approximately in apical 3/5, and a patch at the apex along the elytral suture). Phallobase about 3× longer than wide, approximately 0.74× the length of the penis. Parameres 3/5 the length of the penis. Penis 6.2× longer than wide, tapering towards apex on distal 1/3, apex rounded.
</paragraph>
</subSubSection>
<subSubSection id="7CC97FEA7A36FFC60AEBF89B5BDD0608" lastPageId="4" lastPageNumber="280" pageId="3" pageNumber="279" type="description">
<paragraph id="346C2C617A36FFC10AEBF89B5E340787" blockId="3.[151,1437,1684,2034]" pageId="3" pageNumber="279">
<emphasis id="06A7F0737A36FFC10AEBF89B5A530763" bold="true" box="[199,345,1864,1890]" pageId="3" pageNumber="279">Description.</emphasis>
Male (
<figureCitation id="ACE830E47A36FFC10B9CF89B59040763" box="[432,526,1864,1890]" captionStart="FIGURE 2" captionStartId="2.[151,250,1920,1944]" captionTargetBox="[204,1383,309,1897]" captionTargetId="figure-32@2.[204,1383,309,1897]" captionTargetPageId="2" captionText="FIGURE 2. Hintonelmis spila sp. nov. (Coleoptera: Elmidae). Adult male, habitus. (A) Dorsal view. (B) Ventral view. (C) Lateral view. (D) Front view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618438" httpUri="https://zenodo.org/record/13618438/files/figure.png" pageId="3" pageNumber="279">Fig. 2A</figureCitation>
). Body length =
<quantity id="F32B81847A36FFC108F5F89B586F0762" box="[729,869,1864,1891]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="3.2" metricValueMin="2.1" pageId="3" pageNumber="279" unit="mm" value="2.65" valueMax="3.2" valueMin="2.1">2.13.2 mm</quantity>
, body mean width =
<quantity id="F32B81847A36FFC10E48F89B5FE50762" box="[1124,1263,1864,1891]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="3" pageNumber="279" unit="mm" value="1.0" valueMax="1.2" valueMin="0.8">0.81.2 mm</quantity>
(n = 5). Body elongated; coloration brown with yellowish-brown patches on pronotum and elytra, legs yellowish brown.
</paragraph>
<paragraph id="346C2C617A36FFC60AEBF843584D01D0" blockId="3.[151,1437,1684,2034]" lastBlockId="4.[151,1437,150,2013]" lastPageId="4" lastPageNumber="280" pageId="3" pageNumber="279">
<emphasis id="06A7F0737A36FFC10AEBF8435A0D07AB" bold="true" box="[199,263,1936,1962]" pageId="3" pageNumber="279">Head</emphasis>
(
<figureCitation id="ACE830E47A36FFC10B3AF8435A7A07AB" box="[278,368,1936,1962]" captionStart="FIGURE 3" captionStartId="3.[151,250,1578,1602]" captionTargetBox="[203,1384,181,1554]" captionTargetId="figure-26@3.[203,1384,181,1554]" captionTargetPageId="3" captionText="FIGURE 3. Hintonelmis spila sp. nov. (Coleoptera: Elmidae). Adult, mouthparts. (A) Labrum, dorsal view. (B) Maxilla, dorsal view. (C) Right mandible, dorsal view. (D) Right mandible, ventral view. (E) Labium, ventral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13618440" httpUri="https://zenodo.org/record/13618440/files/figure.png" pageId="3" pageNumber="279">Fig. 3D</figureCitation>
). Antennae uniformly yellowish brown. Labrum (
<figureCitation id="ACE830E47A36FFC109B1F84358F207AB" box="[925,1016,1936,1962]" captionStart="FIGURE 3" captionStartId="3.[151,250,1578,1602]" captionTargetBox="[203,1384,181,1554]" captionTargetId="figure-26@3.[203,1384,181,1554]" captionTargetPageId="3" captionText="FIGURE 3. Hintonelmis spila sp. nov. (Coleoptera: Elmidae). Adult, mouthparts. (A) Labrum, dorsal view. (B) Maxilla, dorsal view. (C) Right mandible, dorsal view. (D) Right mandible, ventral view. (E) Labium, ventral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13618440" httpUri="https://zenodo.org/record/13618440/files/figure.png" pageId="3" pageNumber="279">Fig. 3A</figureCitation>
): subrectangular, with lateral margins rounded; anterolateral margin with long setae; surface with long setae on distal half and short setae on basal half. Mandibles (
<figureCitation id="ACE830E47A36FFC10B33F80B5A9507F3" box="[287,415,2008,2034]" captionStart="FIGURE 3" captionStartId="3.[151,250,1578,1602]" captionTargetBox="[203,1384,181,1554]" captionTargetId="figure-26@3.[203,1384,181,1554]" captionTargetPageId="3" captionText="FIGURE 3. Hintonelmis spila sp. nov. (Coleoptera: Elmidae). Adult, mouthparts. (A) Labrum, dorsal view. (B) Maxilla, dorsal view. (C) Right mandible, dorsal view. (D) Right mandible, ventral view. (E) Labium, ventral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13618440" httpUri="https://zenodo.org/record/13618440/files/figure.png" pageId="3" pageNumber="279">Fig. 3C, D</figureCitation>
): lateral lobe subtriangular, slightly projected; posterolateral angle subtriangular; condyle 1.5× wider than long, semicircular. Maxilla (
<figureCitation id="ACE830E47A31FFC608BEFF4459E600B0" box="[658,748,151,177]" captionStart="FIGURE 3" captionStartId="3.[151,250,1578,1602]" captionTargetBox="[203,1384,181,1554]" captionTargetId="figure-26@3.[203,1384,181,1554]" captionTargetPageId="3" captionText="FIGURE 3. Hintonelmis spila sp. nov. (Coleoptera: Elmidae). Adult, mouthparts. (A) Labrum, dorsal view. (B) Maxilla, dorsal view. (C) Right mandible, dorsal view. (D) Right mandible, ventral view. (E) Labium, ventral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13618440" httpUri="https://zenodo.org/record/13618440/files/figure.png" pageId="4" pageNumber="280">Fig. 3B</figureCitation>
): cardo longer than wide; stipes subtriangular; palpomere IV elongated, 1.5× longer than each of palpomeres IIII, truncate at apex; galea with apical segment subcylindrical, almost 2× longer than basal one, lateral margin with apex rounded, with long and thin setae; lacinia subrectangular, 2× longer than wide, lateral margin with long setae. Labium (3E): mentum 4× wider than long, lateral margins straight, anterolateral angles sharp; postmentum subtriangular, wider than long; prementum subrectangular, base and apex convex, 5× wider than long; ligula subrectangular, 2× wider than long, anterolateral region projected laterally and with median spines, ventral surface with two transverse rows of spines on disc; labial palpus with three palpomeres, palpomere III subcylindrical, 2× wider than the sum of the two basal ones combined, with rounded apex, sensory pores present, external margin with short setae.
</paragraph>
<paragraph id="346C2C617A31FFC60AEBFE085BDA03EC" blockId="4.[151,1437,150,2013]" pageId="4" pageNumber="280">
<emphasis id="06A7F0737A31FFC60AEBFE085A2D01F4" bold="true" box="[199,295,475,501]" pageId="4" pageNumber="280">Thorax.</emphasis>
Pronotum (
<figureCitation id="ACE830E47A31FFC60B81FE08590C01F4" box="[429,518,475,501]" captionStart="FIGURE 2" captionStartId="2.[151,250,1920,1944]" captionTargetBox="[204,1383,309,1897]" captionTargetId="figure-32@2.[204,1383,309,1897]" captionTargetPageId="2" captionText="FIGURE 2. Hintonelmis spila sp. nov. (Coleoptera: Elmidae). Adult male, habitus. (A) Dorsal view. (B) Ventral view. (C) Lateral view. (D) Front view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618438" httpUri="https://zenodo.org/record/13618438/files/figure.png" pageId="4" pageNumber="280">Fig. 2A</figureCitation>
) longer than wide (0.6/
<quantity id="F32B81847A31FFC60922FE09586201F4" box="[782,872,474,501]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="280" unit="mm" value="0.5">0.5 mm</quantity>
); with short yellow setae; lateral margins sinuous; anterolateral angles rounded; anterior margin broadly convex; posterolateral angles acute; posterior margin trisinuate; anterior portion narrower than posterior one; pair of short, strongly raised sublateral carinae on basal 1/3; median longitudinal impression on disc shallow, short; with a transverse impression on the anterior 2/5 extending from one lateral margin to the other; broad, yellowish-brown patch on medial region extending from anterior to posterior region. Elytra (
<figureCitation id="ACE830E47A31FFC60B69FD5C5A9502A8" box="[325,415,655,681]" captionStart="FIGURE 2" captionStartId="2.[151,250,1920,1944]" captionTargetBox="[204,1383,309,1897]" captionTargetId="figure-32@2.[204,1383,309,1897]" captionTargetPageId="2" captionText="FIGURE 2. Hintonelmis spila sp. nov. (Coleoptera: Elmidae). Adult male, habitus. (A) Dorsal view. (B) Ventral view. (C) Lateral view. (D) Front view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618438" httpUri="https://zenodo.org/record/13618438/files/figure.png" pageId="4" pageNumber="280">Fig. 2A</figureCitation>
) subparallel, 1.9× longer than wide (length =
<quantity id="F32B81847A31FFC6099DFD5D5F0602A8" box="[945,1036,654,681]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="280" unit="mm" value="1.5">1.5 mm</quantity>
; maximum width =
<quantity id="F32B81847A31FFC60EDAFD5C5E5802A8" box="[1270,1362,655,681]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="280" unit="mm" value="0.8">0.8 mm</quantity>
); apex truncate; with yellowish-brown patches (one pair elongated, near humeral region, second pair shorter, approximately on apical 3/5, and apical patch along elytral suture). Scutellum cordiform, slightly longer than wide. Metathoracic wings present (
<figureCitation id="ACE830E47A31FFC60B6DFD285A920314" box="[321,408,763,789]" captionStart="FIGURE 4" captionStartId="5.[151,250,1919,1943]" captionTargetBox="[277,1310,181,1894]" captionTargetId="figure-26@5.[277,1310,181,1894]" captionTargetPageId="5" captionText="FIGURE 4. Hintonelmis spila sp. nov. (Coleoptera: Elmidae).Adult, male. (A) Genitalia, ventral view. (B) Tergite VIII, ventral view. (C) Sternite VIII, dorsal view. (D). Segment IX, ventral view. (E) Metathoracic wing. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13618442" httpUri="https://zenodo.org/record/13618442/files/figure.png" pageId="4" pageNumber="280">Fig. 4E</figureCitation>
). Prosternum (
<figureCitation id="ACE830E47A31FFC60817FD28599B0314" box="[571,657,763,789]" captionStart="FIGURE 2" captionStartId="2.[151,250,1920,1944]" captionTargetBox="[204,1383,309,1897]" captionTargetId="figure-32@2.[204,1383,309,1897]" captionTargetPageId="2" captionText="FIGURE 2. Hintonelmis spila sp. nov. (Coleoptera: Elmidae). Adult male, habitus. (A) Dorsal view. (B) Ventral view. (C) Lateral view. (D) Front view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618438" httpUri="https://zenodo.org/record/13618438/files/figure.png" pageId="4" pageNumber="280">Fig. 2B</figureCitation>
) 1.36× wider than long, anterior margin concave, anterolateral angles acute. Prosternal process (
<figureCitation id="ACE830E47A31FFC60B91FCCC591E0338" box="[445,532,799,825]" captionStart="FIGURE 2" captionStartId="2.[151,250,1920,1944]" captionTargetBox="[204,1383,309,1897]" captionTargetId="figure-32@2.[204,1383,309,1897]" captionTargetPageId="2" captionText="FIGURE 2. Hintonelmis spila sp. nov. (Coleoptera: Elmidae). Adult male, habitus. (A) Dorsal view. (B) Ventral view. (C) Lateral view. (D) Front view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618438" httpUri="https://zenodo.org/record/13618438/files/figure.png" pageId="4" pageNumber="280">Fig. 2B</figureCitation>
) shorter than wide (0.17/
<quantity id="F32B81847A31FFC6091CFCCC58920338" box="[816,920,799,825]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="4" pageNumber="280" unit="mm" value="0.23">0.23 mm</quantity>
). Mesoventrite (
<figureCitation id="ACE830E47A31FFC60E63FCCC5FAF0338" box="[1103,1189,799,825]" captionStart="FIGURE 2" captionStartId="2.[151,250,1920,1944]" captionTargetBox="[204,1383,309,1897]" captionTargetId="figure-32@2.[204,1383,309,1897]" captionTargetPageId="2" captionText="FIGURE 2. Hintonelmis spila sp. nov. (Coleoptera: Elmidae). Adult male, habitus. (A) Dorsal view. (B) Ventral view. (C) Lateral view. (D) Front view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618438" httpUri="https://zenodo.org/record/13618438/files/figure.png" pageId="4" pageNumber="280">Fig. 2B</figureCitation>
) wider (
<quantity id="F32B81847A31FFC60F28FCCC5E570338" box="[1284,1373,799,825]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="280" unit="mm" value="0.3">0.3 mm</quantity>
) than long (
<quantity id="F32B81847A31FFC60AF7FC905A33035C" box="[219,313,835,861]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="280" unit="mm" value="0.2">0.2 mm</quantity>
); posterior margin concave. Metaventrite (
<figureCitation id="ACE830E47A31FFC60908FC905877035C" box="[804,893,835,861]" captionStart="FIGURE 2" captionStartId="2.[151,250,1920,1944]" captionTargetBox="[204,1383,309,1897]" captionTargetId="figure-32@2.[204,1383,309,1897]" captionTargetPageId="2" captionText="FIGURE 2. Hintonelmis spila sp. nov. (Coleoptera: Elmidae). Adult male, habitus. (A) Dorsal view. (B) Ventral view. (C) Lateral view. (D) Front view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618438" httpUri="https://zenodo.org/record/13618438/files/figure.png" pageId="4" pageNumber="280">Fig. 2B</figureCitation>
) with disc flat; anterior margin convex; lateral margins covered by tomentum, discrimen on posterior 4/5; surface with few, short setae; posterior region in front of coxae with pair of transverse arched sulci. Posterior leg (
<figureCitation id="ACE830E47A31FFC60936FC58587C03A4" box="[794,886,907,933]" captionStart="FIGURE 2" captionStartId="2.[151,250,1920,1944]" captionTargetBox="[204,1383,309,1897]" captionTargetId="figure-32@2.[204,1383,309,1897]" captionTargetPageId="2" captionText="FIGURE 2. Hintonelmis spila sp. nov. (Coleoptera: Elmidae). Adult male, habitus. (A) Dorsal view. (B) Ventral view. (C) Lateral view. (D) Front view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618438" httpUri="https://zenodo.org/record/13618438/files/figure.png" pageId="4" pageNumber="280">Fig. 2D</figureCitation>
). Femora 4× longer than wide. Tibia 9.7× longer than wide. Apical tarsomere 2× longer than the four basal ones combined. Claws large, stout, with feebly dentate base.
</paragraph>
<paragraph id="346C2C617A31FFC60AEBFC255951050C" blockId="4.[151,1437,150,2013]" pageId="4" pageNumber="280">
<emphasis id="06A7F0737A31FFC60AEBFC255A360411" bold="true" box="[199,316,1014,1040]" pageId="4" pageNumber="280">Abdomen</emphasis>
(
<figureCitation id="ACE830E47A31FFC60B66FC245AA80410" box="[330,418,1015,1041]" captionStart="FIGURE 2" captionStartId="2.[151,250,1920,1944]" captionTargetBox="[204,1383,309,1897]" captionTargetId="figure-32@2.[204,1383,309,1897]" captionTargetPageId="2" captionText="FIGURE 2. Hintonelmis spila sp. nov. (Coleoptera: Elmidae). Adult male, habitus. (A) Dorsal view. (B) Ventral view. (C) Lateral view. (D) Front view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13618438" httpUri="https://zenodo.org/record/13618438/files/figure.png" pageId="4" pageNumber="280">Fig. 2B</figureCitation>
). Ventrites IV with tomentum on disc; ventrite V with yellowish setae on posterior margin. Tergite VIII (
<figureCitation id="ACE830E47A31FFC60B07FBC85A8B0434" box="[299,385,1051,1077]" captionStart="FIGURE 4" captionStartId="5.[151,250,1919,1943]" captionTargetBox="[277,1310,181,1894]" captionTargetId="figure-26@5.[277,1310,181,1894]" captionTargetPageId="5" captionText="FIGURE 4. Hintonelmis spila sp. nov. (Coleoptera: Elmidae).Adult, male. (A) Genitalia, ventral view. (B) Tergite VIII, ventral view. (C) Sternite VIII, dorsal view. (D). Segment IX, ventral view. (E) Metathoracic wing. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13618442" httpUri="https://zenodo.org/record/13618442/files/figure.png" pageId="4" pageNumber="280">Fig. 4B</figureCitation>
) as broad as long, posterior margin rounded, with median setae; surface with microspines and median setae. Sternite VIII (
<figureCitation id="ACE830E47A31FFC60BFDFBEC59230458" box="[465,553,1087,1113]" captionStart="FIGURE 4" captionStartId="5.[151,250,1919,1943]" captionTargetBox="[277,1310,181,1894]" captionTargetId="figure-26@5.[277,1310,181,1894]" captionTargetPageId="5" captionText="FIGURE 4. Hintonelmis spila sp. nov. (Coleoptera: Elmidae).Adult, male. (A) Genitalia, ventral view. (B) Tergite VIII, ventral view. (C) Sternite VIII, dorsal view. (D). Segment IX, ventral view. (E) Metathoracic wing. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13618442" httpUri="https://zenodo.org/record/13618442/files/figure.png" pageId="4" pageNumber="280">Fig. 4C</figureCitation>
): 2× wider than long; surface with short setae and microspines; anterior median projection about 0.5× longer than the body length of sternite. Segment IX (
<figureCitation id="ACE830E47A31FFC609F2FBB05F3C047C" box="[990,1078,1123,1149]" captionStart="FIGURE 4" captionStartId="5.[151,250,1919,1943]" captionTargetBox="[277,1310,181,1894]" captionTargetId="figure-26@5.[277,1310,181,1894]" captionTargetPageId="5" captionText="FIGURE 4. Hintonelmis spila sp. nov. (Coleoptera: Elmidae).Adult, male. (A) Genitalia, ventral view. (B) Tergite VIII, ventral view. (C) Sternite VIII, dorsal view. (D). Segment IX, ventral view. (E) Metathoracic wing. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13618442" httpUri="https://zenodo.org/record/13618442/files/figure.png" pageId="4" pageNumber="280">Fig. 4D</figureCitation>
) subtriangular; apex bifurcated, lateral margins with sensorial pores. Genitalia (
<figureCitation id="ACE830E47A31FFC60899FB54581804A0" box="[693,786,1159,1185]" captionStart="FIGURE 4" captionStartId="5.[151,250,1919,1943]" captionTargetBox="[277,1310,181,1894]" captionTargetId="figure-26@5.[277,1310,181,1894]" captionTargetPageId="5" captionText="FIGURE 4. Hintonelmis spila sp. nov. (Coleoptera: Elmidae).Adult, male. (A) Genitalia, ventral view. (B) Tergite VIII, ventral view. (C) Sternite VIII, dorsal view. (D). Segment IX, ventral view. (E) Metathoracic wing. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13618442" httpUri="https://zenodo.org/record/13618442/files/figure.png" pageId="4" pageNumber="280">Fig. 4A</figureCitation>
). Phallobase 3× longer than wide, 0.74× length of penis. Parameres 3/5 length of the penis, wider at base, tapering towards apex; apex acute. Penis 6.2× longer than wide; tapering towards apex on distal 1/3, apex slightly rounded, with sensory micropores; lateral margin slightly sinuous; basolateral process 2× longer than wide.
</paragraph>
<paragraph id="346C2C617A31FFC60AEBFAC45F6B0530" blockId="4.[151,1437,150,2013]" box="[199,1121,1302,1329]" pageId="4" pageNumber="280">
<emphasis id="06A7F0737A31FFC60AEBFAC45A2F0530" bold="true" box="[199,293,1303,1329]" pageId="4" pageNumber="280">Female.</emphasis>
Similar to male. Body length =
<quantity id="F32B81847A31FFC608A2FAC4581D0530" box="[654,791,1303,1329]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.2" metricValueMin="2.3" pageId="4" pageNumber="280" unit="mm" value="2.75" valueMax="3.2" valueMin="2.3">2.33.2 mm</quantity>
, width =
<quantity id="F32B81847A31FFC609ADFAC45F020530" box="[897,1032,1303,1329]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="4" pageNumber="280" unit="mm" value="1.0" valueMax="1.2" valueMin="0.8">0.81.2 mm</quantity>
(n = 5).
</paragraph>
<paragraph id="346C2C617A31FFC60AEBFAE85BDD0608" blockId="4.[151,1437,150,2013]" pageId="4" pageNumber="280">
Tergite VIII (
<figureCitation id="ACE830E47A31FFC60B73FAE85ABD0554" box="[351,439,1338,1365]" captionStart="FIGURE 5" captionStartId="6.[151,250,1937,1961]" captionTargetBox="[269,1317,181,1905]" captionTargetId="figure-22@6.[269,1317,181,1913]" captionTargetPageId="6" captionText="FIGURE 5. Hintonelmis spila sp. nov. (Coleoptera: Elmidae). Adult, female. (A) Ovipositor, ventral view. (B) Tergite VIII, dorsal view, with the anterior region highlighted. (C) Sternite VIII, dorsal view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13618444" httpUri="https://zenodo.org/record/13618444/files/figure.png" pageId="4" pageNumber="280">Fig. 5B</figureCitation>
) wider than long, posterior margin rounded, with short setae; surface with microspines and long setae. Sternite VIII (
<figureCitation id="ACE830E47A31FFC60BCFFA8C59330578" box="[483,569,1374,1401]" captionStart="FIGURE 5" captionStartId="6.[151,250,1937,1961]" captionTargetBox="[269,1317,181,1905]" captionTargetId="figure-22@6.[269,1317,181,1913]" captionTargetPageId="6" captionText="FIGURE 5. Hintonelmis spila sp. nov. (Coleoptera: Elmidae). Adult, female. (A) Ovipositor, ventral view. (B) Tergite VIII, dorsal view, with the anterior region highlighted. (C) Sternite VIII, dorsal view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13618444" httpUri="https://zenodo.org/record/13618444/files/figure.png" pageId="4" pageNumber="280">Fig. 5C</figureCitation>
) 1.7× wider than long; surface with median setae; anterior median projection 4.3× longer than the remaining part of sternite; posterior margin with short setae, surface with short setae and microspines. Ovipositor (
<figureCitation id="ACE830E47A31FFC60B9BFA74591905C0" box="[439,531,1446,1473]" captionStart="FIGURE 5" captionStartId="6.[151,250,1937,1961]" captionTargetBox="[269,1317,181,1905]" captionTargetId="figure-22@6.[269,1317,181,1913]" captionTargetPageId="6" captionText="FIGURE 5. Hintonelmis spila sp. nov. (Coleoptera: Elmidae). Adult, female. (A) Ovipositor, ventral view. (B) Tergite VIII, dorsal view, with the anterior region highlighted. (C) Sternite VIII, dorsal view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13618444" httpUri="https://zenodo.org/record/13618444/files/figure.png" pageId="4" pageNumber="280">Fig. 5A</figureCitation>
): valvifer dilated on posterior 1/3. Apical coxite 4.6× longer than wide, surface with sensory micropores; 1.9× longer than basal coxite, with sensory microspores. Stylus divergent, apex with short setae.
</paragraph>
</subSubSection>
<subSubSection id="7CC97FEA7A31FFC50AEBF9C058BF0188" lastPageId="7" lastPageNumber="283" pageId="4" pageNumber="280" type="etymology">
<paragraph id="346C2C617A31FFC60AEBF9C05F5E062C" blockId="4.[151,1437,150,2013]" box="[199,1108,1555,1581]" pageId="4" pageNumber="280">
<emphasis id="06A7F0737A31FFC60AEBF9C05A44062C" bold="true" box="[199,334,1555,1581]" pageId="4" pageNumber="280">Etymology.</emphasis>
From the Greek word
<emphasis id="06A7F0737A31FFC60862F9C0597C062C" box="[590,630,1555,1581]" italics="true" pageId="4" pageNumber="280">spil</emphasis>
(= stain), a reference to the elytra patches.
</paragraph>
<paragraph id="346C2C617A31FFC50AEBF9E558BF0188" blockId="4.[151,1437,150,2013]" lastBlockId="7.[151,1437,151,861]" lastPageId="7" lastPageNumber="283" pageId="4" pageNumber="280">
<emphasis id="06A7F0737A31FFC60AEBF9E55AA30650" bold="true" box="[199,425,1590,1617]" pageId="4" pageNumber="280">Comparative notes</emphasis>
.
<taxonomicName id="F3D357E27A31FFC60B94F9E459740650" authorityName="Pinedo-Garcia &amp; Hamada &amp; Nascimento" authorityYear="2024" box="[440,638,1591,1617]" class="Insecta" family="Elmidae" genus="Hintonelmis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="280" phylum="Arthropoda" rank="species" species="spila" status="sp. nov.">
<emphasis id="06A7F0737A31FFC60B94F9E459740650" box="[440,638,1591,1617]" italics="true" pageId="4" pageNumber="280">Hintonelmis spila</emphasis>
</taxonomicName>
<emphasis id="06A7F0737A31FFC608AAF9E459D50650" bold="true" box="[646,735,1591,1617]" pageId="4" pageNumber="280">
<taxonomicNameLabel id="1D944D087A31FFC608AAF9E459D50650" box="[646,735,1591,1617]" pageId="4" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resembles
<emphasis id="06A7F0737A31FFC60972F9E458F80651" box="[862,1010,1591,1616]" italics="true" pageId="4" pageNumber="280">
<taxonomicName id="F3D357E27A31FFC60972F9E458E40651" authorityName="Fernandes, Passos &amp; Hamada" authorityYear="2010" box="[862,1006,1591,1616]" class="Insecta" family="Elmidae" genus="Hintonelmis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="280" phylum="Arthropoda" rank="species" species="anamarie">H. anamarie</taxonomicName>
,
</emphasis>
especially in coloration pattern of the elytra with five patches (one pair near the humeral region, one pair approximately on apical 3/5, and a patch at the apex along the elytral suture). However, both species can be easily differentiated as follows: in
<taxonomicName id="F3D357E27A31FFC60EF9F9AC5E960698" authorityName="Pinedo-Garcia &amp; Hamada &amp; Nascimento" authorityYear="2024" box="[1237,1436,1663,1689]" class="Insecta" family="Elmidae" genus="Hintonelmis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="280" phylum="Arthropoda" rank="species" species="spila" status="sp. nov.">
<emphasis id="06A7F0737A31FFC60EF9F9AC5E960698" box="[1237,1436,1663,1689]" italics="true" pageId="4" pageNumber="280">Hintonelmis spila</emphasis>
</taxonomicName>
<emphasis id="06A7F0737A31FFC60ABBF9705BF806BC" bold="true" box="[151,242,1699,1725]" pageId="4" pageNumber="280">
<taxonomicNameLabel id="1D944D087A31FFC60ABBF9705BF806BC" box="[151,242,1699,1725]" pageId="4" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, the antenna has a uniform coloration, whereas in
<taxonomicName id="F3D357E27A31FFC6091DF97058B506BD" authorityName="Fernandes, Passos &amp; Hamada" authorityYear="2010" box="[817,959,1699,1724]" class="Insecta" family="Elmidae" genus="Hintonelmis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="280" phylum="Arthropoda" rank="species" species="anamarie">
<emphasis id="06A7F0737A31FFC6091DF97058B506BD" box="[817,959,1699,1724]" italics="true" pageId="4" pageNumber="280">H. anamarie</emphasis>
</taxonomicName>
the antenna is bicolored; the pronotum of the new species has a broad, yellowish-brown patch in the medial region extending from the anterior to the posterior region, whereas in
<taxonomicName id="F3D357E27A31FFC60B43F9385AF70705" authorityName="Fernandes, Passos &amp; Hamada" authorityYear="2010" box="[367,509,1771,1796]" class="Insecta" family="Elmidae" genus="Hintonelmis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="280" phylum="Arthropoda" rank="species" species="anamarie">
<emphasis id="06A7F0737A31FFC60B43F9385AF70705" box="[367,509,1771,1796]" italics="true" pageId="4" pageNumber="280">H. anamarie</emphasis>
</taxonomicName>
this patch is more rounded and located anteriorly in the medial region. The shape of the patches in the humeral region of the elytra can be helpful to differentiate both species: in the new species, the patches are elongated and do not reach the anterolateral margin of the elytra, whereas in
<taxonomicName id="F3D357E27A31FFC60E8AF8E05E3E074D" authorityName="Fernandes, Passos &amp; Hamada" authorityYear="2010" box="[1190,1332,1843,1868]" class="Insecta" family="Elmidae" genus="Hintonelmis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="280" phylum="Arthropoda" rank="species" species="anamarie">
<emphasis id="06A7F0737A31FFC60E8AF8E05E3E074D" box="[1190,1332,1843,1868]" italics="true" pageId="4" pageNumber="280">H. anamarie</emphasis>
</taxonomicName>
, they are rounded, extending diagonally from the anterolateral margin. Additionally, the male genitalia of both species can be differentiated as follows: in
<taxonomicName id="F3D357E27A31FFC60BD1F8A859CF0794" authorityName="Pinedo-Garcia &amp; Hamada &amp; Nascimento" authorityYear="2024" box="[509,709,1915,1941]" class="Insecta" family="Elmidae" genus="Hintonelmis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="280" phylum="Arthropoda" rank="species" species="spila" status="sp. nov.">
<emphasis id="06A7F0737A31FFC60BD1F8A859CF0794" box="[509,709,1915,1941]" italics="true" pageId="4" pageNumber="280">Hintonelmis spila</emphasis>
</taxonomicName>
<emphasis id="06A7F0737A31FFC608E2F8A858200794" bold="true" box="[718,810,1915,1941]" pageId="4" pageNumber="280">
<taxonomicNameLabel id="1D944D087A31FFC608E2F8A858200794" box="[718,810,1915,1941]" pageId="4" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, the parameres are 3/5 the length of the penis, and the phallobase is shorter than the penis; whereas in
<taxonomicName id="F3D357E27A31FFC6089CF84C583707B9" authorityName="Fernandes, Passos &amp; Hamada" authorityYear="2010" box="[688,829,1951,1976]" class="Insecta" family="Elmidae" genus="Hintonelmis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="280" phylum="Arthropoda" rank="species" species="anamarie">
<emphasis id="06A7F0737A31FFC6089CF84C583707B9" box="[688,829,1951,1976]" italics="true" pageId="4" pageNumber="280">H. anamarie</emphasis>
</taxonomicName>
, the parameres are 4/5 the length of the penis, and the phallobase is nearly the same length as the penis.
<taxonomicName id="F3D357E27A31FFC60895F810587107DC" authorityName="Pinedo-Garcia &amp; Hamada &amp; Nascimento" authorityYear="2024" box="[697,891,1987,2013]" class="Insecta" family="Elmidae" genus="Hintonelmis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="280" phylum="Arthropoda" rank="species" species="spila" status="sp. nov.">
<emphasis id="06A7F0737A31FFC60895F810587107DC" box="[697,891,1987,2013]" italics="true" pageId="4" pageNumber="280">Hintonelmis spila</emphasis>
</taxonomicName>
<emphasis id="06A7F0737A31FFC609AEF81058D007DC" bold="true" box="[898,986,1987,2013]" pageId="4" pageNumber="280">
<taxonomicNameLabel id="1D944D087A31FFC609AEF81058D007DC" box="[898,986,1987,2013]" pageId="4" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
also shares with
<taxonomicName id="F3D357E27A31FFC60EBBF8105E0F07DC" authorityName="Polizei &amp; Barclay" authorityYear="2018" box="[1175,1285,1987,2013]" class="Insecta" family="Elmidae" genus="Hintonelmis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="280" phylum="Arthropoda" rank="species" species="roellae">
<emphasis id="06A7F0737A31FFC60EBBF8105E0F07DC" box="[1175,1285,1987,2013]" italics="true" pageId="4" pageNumber="280">H. roellae</emphasis>
</taxonomicName>
the antennae with uniform coloration, yellowish-brown, and elongated patches on the elytra. However, the anterior patches in
<taxonomicName id="F3D357E27A32FFC50ABBFF685A0200D4" authorityName="Polizei &amp; Barclay" authorityYear="2018" box="[151,264,187,213]" class="Insecta" family="Elmidae" genus="Hintonelmis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="283" phylum="Arthropoda" rank="species" species="roellae">
<emphasis id="06A7F0737A32FFC50ABBFF685A0200D4" box="[151,264,187,213]" italics="true" pageId="7" pageNumber="283">H. roellae</emphasis>
</taxonomicName>
are more elongated, extending beyond the distal half of the elytra (in the new species, these patches do not extend beyond the distal half). In addition,
<taxonomicName id="F3D357E27A32FFC50886FF0C581100F8" authorityName="Polizei &amp; Barclay" authorityYear="2018" box="[682,795,223,249]" class="Insecta" family="Elmidae" genus="Hintonelmis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="283" phylum="Arthropoda" rank="species" species="roellae">
<emphasis id="06A7F0737A32FFC50886FF0C581100F8" box="[682,795,223,249]" italics="true" pageId="7" pageNumber="283">H. roellae</emphasis>
</taxonomicName>
has a long and deep longitudinal impression on the disc, whereas
<taxonomicName id="F3D357E27A32FFC50AD4FED05A45011C" authorityName="Pinedo-Garcia &amp; Hamada &amp; Nascimento" authorityYear="2024" box="[248,335,259,285]" class="Insecta" family="Elmidae" genus="Hintonelmis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="283" phylum="Arthropoda" rank="species" species="spila" status="sp. nov.">
<emphasis id="06A7F0737A32FFC50AD4FED05A45011C" box="[248,335,259,285]" italics="true" pageId="7" pageNumber="283">H. spila</emphasis>
</taxonomicName>
<emphasis id="06A7F0737A32FFC50B79FED05AA7011C" bold="true" box="[341,429,259,285]" pageId="7" pageNumber="283">
<taxonomicNameLabel id="1D944D087A32FFC50B79FED05AA7011C" box="[341,429,259,285]" pageId="7" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a short and shallow one. Finally, the male genitalia of both species can be differentiated as follows: in
<taxonomicName id="F3D357E27A32FFC50B1AFEF45AF60140" authorityName="Pinedo-Garcia &amp; Hamada &amp; Nascimento" authorityYear="2024" box="[310,508,295,321]" class="Insecta" family="Elmidae" genus="Hintonelmis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="283" phylum="Arthropoda" rank="species" species="spila" status="sp. nov.">
<emphasis id="06A7F0737A32FFC50B1AFEF45AF60140" box="[310,508,295,321]" italics="true" pageId="7" pageNumber="283">Hintonelmis spila</emphasis>
</taxonomicName>
<emphasis id="06A7F0737A32FFC50828FEF459540140" bold="true" box="[516,606,295,321]" pageId="7" pageNumber="283">
<taxonomicNameLabel id="1D944D087A32FFC50828FEF459540140" box="[516,606,295,321]" pageId="7" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, the parameres are 3/5 the length of the penis, the phallobase is 3× longer than wide, and the penis is 6.2× longer than wide; in
<taxonomicName id="F3D357E27A32FFC508C9FE98587B0165" authorityName="Fernandes, Passos &amp; Hamada" authorityYear="2010" box="[741,881,331,356]" class="Insecta" family="Elmidae" genus="Hintonelmis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="283" phylum="Arthropoda" rank="species" species="anamarie">
<emphasis id="06A7F0737A32FFC508C9FE98587B0165" box="[741,881,331,356]" italics="true" pageId="7" pageNumber="283">H. anamarie</emphasis>
</taxonomicName>
, the parameres are 4/5 the length of the penis, the phallobase is 2× longer than wide and the penis is 4× longer than wide.
</paragraph>
</subSubSection>
<caption id="60AC7CE97A30FFC70ABBF8AC5E1B07BB" ID-DOI="http://doi.org/10.5281/zenodo.13618442" ID-Zenodo-Dep="13618442" httpUri="https://zenodo.org/record/13618442/files/figure.png" pageId="5" pageNumber="281" startId="5.[151,250,1919,1943]" targetBox="[277,1310,181,1894]" targetPageId="5" targetType="figure">
<paragraph id="346C2C617A30FFC70ABBF8AC5E1B07BB" blockId="5.[151,1436,1919,1979]" pageId="5" pageNumber="281">
<emphasis id="06A7F0737A30FFC70ABBF8AC5A180796" bold="true" box="[151,274,1919,1943]" pageId="5" pageNumber="281">FIGURE 4.</emphasis>
<taxonomicName id="F3D357E27A30FFC70B34F8AC5AC00796" authorityName="Pinedo-Garcia &amp; Hamada &amp; Nascimento" authorityYear="2024" box="[280,458,1919,1943]" class="Insecta" family="Elmidae" genus="Hintonelmis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="spila" status="sp. nov.">
<emphasis id="06A7F0737A30FFC70B34F8AC5AC00796" box="[280,458,1919,1943]" italics="true" pageId="5" pageNumber="281">Hintonelmis spila</emphasis>
</taxonomicName>
<emphasis id="06A7F0737A30FFC70BFCF8AC59150796" bold="true" box="[464,543,1919,1943]" pageId="5" pageNumber="281">
<taxonomicNameLabel id="1D944D087A30FFC70BFCF8AC59150796" box="[464,543,1919,1943]" pageId="5" pageNumber="281" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<taxonomicName id="F3D357E27A30FFC70800F8AC59AB0796" authorityName="Linnaeus" authorityYear="1758" box="[556,673,1919,1943]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
:
<taxonomicName id="F3D357E27A30FFC70885F8AC58080796" authorityName="Curtis" authorityYear="1830" box="[681,770,1919,1943]" class="Insecta" family="Elmidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="family">Elmidae</taxonomicName>
). Adult, male. (A) Genitalia, ventral view. (B) Tergite VIII, ventral view. (C) Sternite VIII, dorsal view. (D). Segment IX, ventral view. (E) Metathoracic wing. Scale bars = 0.1 mm.
</paragraph>
</caption>
<caption id="60AC7CE97A33FFC40ABBF8425F9307CC" ID-DOI="http://doi.org/10.5281/zenodo.13618444" ID-Zenodo-Dep="13618444" httpUri="https://zenodo.org/record/13618444/files/figure.png" pageId="6" pageNumber="282" startId="6.[151,250,1937,1961]" targetBox="[269,1317,181,1905]" targetPageId="6" targetType="figure">
<paragraph id="346C2C617A33FFC40ABBF8425F9307CC" blockId="6.[151,1436,1937,1998]" pageId="6" pageNumber="282">
<emphasis id="06A7F0737A33FFC40ABBF8425A1F07AB" bold="true" box="[151,277,1937,1962]" pageId="6" pageNumber="282">FIGURE 5.</emphasis>
<taxonomicName id="F3D357E27A33FFC40B31F8425AD807A8" authorityName="Pinedo-Garcia &amp; Hamada &amp; Nascimento" authorityYear="2024" box="[285,466,1937,1961]" class="Insecta" family="Elmidae" genus="Hintonelmis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="282" phylum="Arthropoda" rank="species" species="spila" status="sp. nov.">
<emphasis id="06A7F0737A33FFC40B31F8425AD807A8" box="[285,466,1937,1961]" italics="true" pageId="6" pageNumber="282">Hintonelmis spila</emphasis>
</taxonomicName>
<emphasis id="06A7F0737A33FFC40BF6F841592607AB" bold="true" box="[474,556,1938,1962]" pageId="6" pageNumber="282">
<taxonomicNameLabel id="1D944D087A33FFC40BF6F841592607AB" box="[474,556,1938,1962]" pageId="6" pageNumber="282" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<taxonomicName id="F3D357E27A33FFC40810F84259BB07A8" authorityName="Linnaeus" authorityYear="1758" box="[572,689,1937,1961]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="282" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
:
<taxonomicName id="F3D357E27A33FFC40890F842581F07A8" authorityName="Curtis" authorityYear="1830" box="[700,789,1937,1961]" class="Insecta" family="Elmidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="282" phylum="Arthropoda" rank="family">Elmidae</taxonomicName>
). Adult, female. (A) Ovipositor, ventral view. (B) Tergite VIII, dorsal view, with the anterior region highlighted. (C) Sternite VIII, dorsal view. Scale bars = 0.1 mm.
</paragraph>
</caption>
<subSubSection id="7CC97FEA7A32FFC50AEBFE415E1701AC" box="[199,1309,402,429]" pageId="7" pageNumber="283" type="distribution">
<paragraph id="346C2C617A32FFC50AEBFE415E1701AC" blockId="7.[151,1437,151,861]" box="[199,1309,402,429]" pageId="7" pageNumber="283">
<emphasis id="06A7F0737A32FFC50AEBFE415A6B01AD" bold="true" box="[199,353,402,428]" pageId="7" pageNumber="283">Distribution.</emphasis>
<collectingCountry id="4CC46CF17A32FFC50B4BFE405AA101AC" box="[359,427,403,429]" name="Brazil" pageId="7" pageNumber="283">Brazil</collectingCountry>
(
<collectingRegion id="F617E2837A32FFC50B97FE405AFC01AC" box="[443,502,403,429]" country="Brazil" name="Acre" pageId="7" pageNumber="283">Acre</collectingRegion>
, Amazonas,
<collectingRegion id="F617E2837A32FFC508AFFE4059F901AC" box="[643,755,403,429]" country="Brazil" name="Rondonia" pageId="7" pageNumber="283">Rondônia</collectingRegion>
, and Pará states),
<collectingCountry id="4CC46CF17A32FFC50990FE405F1C01AC" box="[956,1046,403,429]" name="Guyana" pageId="7" pageNumber="283">Guyana</collectingCountry>
, and
<collectingCountry id="4CC46CF17A32FFC50E7CFE405FB101AC" box="[1104,1211,403,429]" name="Suriname" pageId="7" pageNumber="283">Suriname</collectingCountry>
(
<figureCitation id="ACE830E47A32FFC50EE7FE405E0601AC" box="[1227,1292,403,429]" captionStart="FIGURE 1" captionStartId="1.[151,250,1399,1423]" captionTargetBox="[176,1412,417,1375]" captionTargetId="figure-100@1.[176,1412,417,1375]" captionTargetPageId="1" captionText="FIGURE 1. Map of South America with detailed area indicating the distributional records and general view of the sampled streams for Hintonelmis spila sp. nov. (Coleoptera: Elmidae)." figureDoi="http://doi.org/10.5281/zenodo.13618436" httpUri="https://zenodo.org/record/13618436/files/figure.png" pageId="7" pageNumber="283">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="7CC97FEA7A32FFC50AEBFE6459ED035C" pageId="7" pageNumber="283" type="materials_examined">
<paragraph id="346C2C617A32FFC50AEBFE6459160218" blockId="7.[151,1437,151,861]" pageId="7" pageNumber="283">
<emphasis id="06A7F0737A32FFC50AEBFE645A7601D1" bold="true" box="[199,380,438,465]" pageId="7" pageNumber="283">Type material.</emphasis>
<materialsCitation id="84BB263C7A32FFC50BA5FE6459160218" collectingDate="2022-06-15" collectionCode="R" collectorName="B. Pinedo-Garcia &amp; J. O. Silva &amp; G. Desiderio &amp; A. Pes &amp; H. L. Ferreira" country="Brazil" latitude="-8.009639" location="Japiim river" longLatPrecision="1" longitude="-69.78758" municipality="Mancio Lima" pageId="7" pageNumber="283" specimenCount="1" stateProvince="Acre" typeStatus="holotype">
<emphasis id="06A7F0737A32FFC50BA5FE645AF701D0" bold="true" box="[393,509,439,465]" pageId="7" pageNumber="283">
<typeStatus id="EB6892C37A32FFC50BA5FE645AF201D0" box="[393,504,439,465]" pageId="7" pageNumber="283" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male.
<collectingCountry id="4CC46CF17A32FFC50875FE6459CB01D0" box="[601,705,439,465]" name="Brazil" pageId="7" pageNumber="283">BRAZIL</collectingCountry>
,
<emphasis id="06A7F0737A32FFC508FFFE64580D01D0" box="[723,775,439,465]" italics="true" pageId="7" pageNumber="283">
<collectingRegion id="F617E2837A32FFC508FFFE64580D01D0" box="[723,775,439,465]" country="Brazil" name="Acre" pageId="7" pageNumber="283">Acre</collectingRegion>
</emphasis>
.
<collectingMunicipality id="D408B61B7A32FFC50937FE6458B001D0" box="[795,954,439,465]" pageId="7" pageNumber="283">Mâncio Lima</collectingMunicipality>
,
<location id="310C7ABA7A32FFC509E2FE645F5401D0" LSID="urn:lsid:plazi:treatment:BC7A9D777A37FFC50ABBFF0C59ED035C:310C7ABA7A32FFC509E2FE645F5401D0" box="[974,1118,439,465]" country="Brazil" latitude="-8.009639" longLatPrecision="1" longitude="-69.78758" municipality="Mancio Lima" name="Japiim river" pageId="7" pageNumber="283" stateProvince="Acre">Japiim river</location>
,
<date id="406D0AA17A32FFC50E43FE655FE301D0" box="[1135,1257,438,465]" pageId="7" pageNumber="283" value="2022-06-15">
<collectingDate id="5029F3497A32FFC50E43FE655FE301D0" box="[1135,1257,438,465]" pageId="7" pageNumber="283" value="2022-06-15">15.vi.2022</collectingDate>
</date>
,
<geoCoordinate id="51E74AA67A32FFC50ED0FE645E9201D0" box="[1276,1432,439,465]" degrees="07" direction="south" minutes="60" orientation="latitude" pageId="7" pageNumber="283" precision="1" seconds="34.7" value="-8.009639">07°6034.7”S</geoCoordinate>
,
<geoCoordinate id="51E74AA67A32FFC50ABBFE095A3001F4" box="[151,314,474,501]" degrees="69" direction="west" minutes="47" orientation="longitude" pageId="7" pageNumber="283" precision="1" seconds="15.3" value="-69.78758">69°4715.3”W</geoCoordinate>
;
<collectionCode id="52C2B4A47A32FFC50B63FE085A6C01F4" box="[335,358,475,501]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="283">R</collectionCode>
.
<collectorName id="992649B77A32FFC50B46FE08593E01F4" box="[362,564,475,501]" pageId="7" pageNumber="283">B. Pinedo-Garcia</collectorName>
,
<collectorName id="992649B77A32FFC50869FE0859B101F4" box="[581,699,475,501]" pageId="7" pageNumber="283">J.O. Silva</collectorName>
,
<collectorName id="992649B77A32FFC508E1FE08586F01F4" box="[717,869,475,501]" pageId="7" pageNumber="283">G. Desiderio</collectorName>
,
<collectorName id="992649B77A32FFC50958FE0858CC01F4" box="[884,966,475,501]" pageId="7" pageNumber="283">A. Pes</collectorName>
,
<collectorName id="992649B77A32FFC509FBFE085F7A01F4" box="[983,1136,475,501]" pageId="7" pageNumber="283">H.L. Ferreira</collectorName>
<emphasis id="06A7F0737A32FFC50E51FE085F9501F4" box="[1149,1183,475,501]" italics="true" pageId="7" pageNumber="283">leg</emphasis>
. (INPA-body pinned, mouthparts and genitalia on slide).
</materialsCitation>
</paragraph>
<paragraph id="346C2C617A32FFC50AEBFDF059ED035C" blockId="7.[151,1437,151,861]" pageId="7" pageNumber="283">
<materialsCitation id="84BB263C7A32FFC50AEBFDF059E9035C" collectingDate="2010-08-27" collectingDateMax="2021-08-06" collectingDateMin="2010-08-27" collectionCode="R" collectorName="L. S. Santos &amp; J. A. Rafael &amp; N. Hamada &amp; A. Short" country="Guyana" county="Same" latitude="2.1693695" location="Liberdade River" longLatPrecision="1" longitude="-56.78986" municipality="Rondonia. Urupa River" pageId="7" pageNumber="283" specimenCount="55" specimenCount-female="15" specimenCount-male="40" stateProvince="Sipaliwini" typeStatus="paratype">
<emphasis id="06A7F0737A32FFC50AEBFDF05A4C023C" bold="true" box="[199,326,547,573]" pageId="7" pageNumber="283">
<typeStatus id="EB6892C37A32FFC50AEBFDF05A4B023C" box="[199,321,547,573]" pageId="7" pageNumber="283" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingCounty id="DD0D54ED7A32FFC50B7CFDF05A84023C" box="[336,398,547,573]" pageId="7" pageNumber="283">Same</collectingCounty>
data as holotype (
<specimenCount id="22D5E7E87A32FFC5084BFDF059A9023C" box="[615,675,547,573]" count="10" pageId="7" pageNumber="283" type="male">10 ♂</specimenCount>
and
<specimenCount id="22D5E7E87A32FFC508CCFDF05811023C" box="[736,795,547,573]" count="10" pageId="7" pageNumber="283" type="female">10 ♀</specimenCount>
in INPA-alcohol), (
<specimenCount id="22D5E7E87A32FFC50E2FFDF05F35023C" box="[1027,1087,547,573]" count="10" pageId="7" pageNumber="283" type="male">10 ♂</specimenCount>
and
<specimenCount id="22D5E7E87A32FFC50E50FDF15FA3023C" box="[1148,1193,546,573]" count="5" pageId="7" pageNumber="283" type="female">5 ♀</specimenCount>
in MZUSP-alcohol).
<collectingMunicipality id="D408B61B7A32FFC50ABBFD945AAF0260" box="[151,421,583,609]" pageId="7" pageNumber="283">
<emphasis id="06A7F0737A32FFC50ABBFD945A0E0260" box="[151,260,583,609]" italics="true" pageId="7" pageNumber="283">Rondônia</emphasis>
. Urupá River
</collectingMunicipality>
,
<date id="406D0AA17A32FFC50B9FFD94593F0260" box="[435,565,583,609]" pageId="7" pageNumber="283" value="2021-07-31">
<collectingDate id="5029F3497A32FFC50B9FFD94593F0260" box="[435,565,583,609]" pageId="7" pageNumber="283" value="2021-07-31">31.vii.2021</collectingDate>
</date>
,
<geoCoordinate id="51E74AA67A32FFC5086FFD9459D70260" box="[579,733,583,609]" degrees="11" direction="south" minutes="02" orientation="latitude" pageId="7" pageNumber="283" precision="1" seconds="11.0" value="-11.036389">11°0211.0”S</geoCoordinate>
,
<geoCoordinate id="51E74AA67A32FFC508C7FD9458870260" box="[747,909,583,609]" degrees="62" direction="west" minutes="08" orientation="longitude" pageId="7" pageNumber="283" precision="1" seconds="42.0" value="-62.145">62°0842.0”W</geoCoordinate>
,
<collectorName id="992649B77A32FFC509B2FD945F2B0260" box="[926,1057,583,609]" pageId="7" pageNumber="283">L.S. Santos</collectorName>
<emphasis id="06A7F0737A32FFC50E07FD945F930260" box="[1067,1177,583,609]" italics="true" pageId="7" pageNumber="283">et al. leg.</emphasis>
(
<quantity id="F32B81847A32FFC50E87FD945FE20260" box="[1195,1256,583,609]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" pageId="7" pageNumber="283" unit="in" value="16.0">16 in</quantity>
INPA-alcohol).
<emphasis id="06A7F0737A32FFC50ABBFDB85A000284" box="[151,266,619,645]" italics="true" pageId="7" pageNumber="283">Amazonas</emphasis>
. Ipixuna,
<location id="310C7ABA7A32FFC50BAEFDB859370284" LSID="urn:lsid:plazi:treatment:BC7A9D777A37FFC50ABBFF0C59ED035C:310C7ABA7A32FFC50BAEFDB859370284" box="[386,573,619,645]" country="Guyana" county="Same" latitude="2.1693695" longLatPrecision="1" longitude="-56.78986" municipality="Rondonia. Urupa River" name="Liberdade River" pageId="7" pageNumber="283" stateProvince="Sipaliwini">Liberdade River</location>
,
<date id="406D0AA17A32FFC50860FDB859EC0284" box="[588,742,619,645]" pageId="7" pageNumber="283" value="2011-05-12" valueMax="2011-05-13" valueMin="2011-05-12">
<collectingDate id="5029F3497A32FFC50860FDB859EC0284" box="[588,742,619,645]" pageId="7" pageNumber="283" value="2011-05-12" valueMax="2011-05-13" valueMin="2011-05-12">1213.v.2011</collectingDate>
</date>
, 07°2146”7S,
<geoCoordinate id="51E74AA67A32FFC509BBFDB95F2E0284" box="[919,1060,618,645]" degrees="71" direction="west" minutes="52" orientation="longitude" pageId="7" pageNumber="283" precision="15" seconds="07" value="-71.868614">71°5207”W</geoCoordinate>
,
<collectorName id="992649B77A32FFC50E19FDB85FBF0284" box="[1077,1205,619,645]" pageId="7" pageNumber="283">J.A. Rafael</collectorName>
<emphasis id="06A7F0737A32FFC50EECFDB85E2D0284" box="[1216,1319,619,645]" italics="true" pageId="7" pageNumber="283">et al. leg</emphasis>
. (
<specimenCount id="22D5E7E87A32FFC50F6CFDB85E760284" box="[1344,1404,619,645]" count="10" pageId="7" pageNumber="283" type="male">10 ♂</specimenCount>
in
<emphasis id="06A7F0737A32FFC50ABBFD5C5BD102A8" box="[151,219,655,681]" italics="true" pageId="7" pageNumber="283">INPA-</emphasis>
pinned). Gregorio River, 2021.
<date id="406D0AA17A32FFC50817FD5C598202A8" box="[571,648,655,681]" pageId="7" pageNumber="283" value="2011-05">
<collectingDate id="5029F3497A32FFC50817FD5C598202A8" box="[571,648,655,681]" pageId="7" pageNumber="283" value="2011-05">v.2011</collectingDate>
</date>
,
<geoCoordinate id="51E74AA67A32FFC508B9FD5C583A02A8" box="[661,816,655,681]" degrees="07" direction="south" minutes="10" orientation="latitude" pageId="7" pageNumber="283" precision="1" seconds="11.7" value="-7.1699166">07°1011.7”S</geoCoordinate>
,
<geoCoordinate id="51E74AA67A32FFC50912FD5C58EA02A8" box="[830,992,655,681]" degrees="70" direction="west" minutes="49" orientation="longitude" pageId="7" pageNumber="283" precision="1" seconds="10.3" value="-70.81953">70°4910.3”W</geoCoordinate>
,
<collectorName id="992649B77A32FFC509C3FD5C5F6502A8" box="[1007,1135,655,681]" pageId="7" pageNumber="283">J.A. Rafael</collectorName>
<emphasis id="06A7F0737A32FFC50E54FD5C5FD702A8" box="[1144,1245,655,681]" italics="true" pageId="7" pageNumber="283">et al. leg</emphasis>
. (
<specimenCount id="22D5E7E87A32FFC50EDAFD5C5E3A02A8" box="[1270,1328,655,681]" count="10" pageId="7" pageNumber="283" type="male">10 ♂</specimenCount>
in
<emphasis id="06A7F0737A32FFC50F74FD5C5BE202CC" italics="true" pageId="7" pageNumber="283">INPApinned</emphasis>
).
<emphasis id="06A7F0737A32FFC50AD2FD605A3C02CC" box="[254,310,691,717]" italics="true" pageId="7" pageNumber="283">Pará</emphasis>
. Goianesia, Capim River,
<date id="406D0AA17A32FFC50847FD61580802CC" box="[619,770,690,717]" pageId="7" pageNumber="283" value="2021-08-05" valueMax="2021-08-06" valueMin="2021-08-05">
<collectingDate id="5029F3497A32FFC50847FD61580802CC" box="[619,770,690,717]" pageId="7" pageNumber="283" value="2021-08-05" valueMax="2021-08-06" valueMin="2021-08-05">56.viii.2021</collectingDate>
</date>
,
<geoCoordinate id="51E74AA67A32FFC5093DFD61589502CC" box="[785,927,690,717]" degrees="3" direction="south" minutes="42" orientation="latitude" pageId="7" pageNumber="283" precision="1" seconds="35.3" value="-3.7098055">3°4235.3”S</geoCoordinate>
,
<geoCoordinate id="51E74AA67A32FFC50981FD605F4502CC" box="[941,1103,691,717]" degrees="48" direction="west" minutes="33" orientation="longitude" pageId="7" pageNumber="283" precision="1" seconds="42.4" value="-48.561775">48°3342.4”W</geoCoordinate>
,
<collectorName id="992649B77A32FFC50E4CFD605FE902CC" box="[1120,1251,691,717]" pageId="7" pageNumber="283">N. Hamada</collectorName>
<emphasis id="06A7F0737A32FFC50EC1FD605E5E02CC" box="[1261,1364,691,717]" italics="true" pageId="7" pageNumber="283">et al. leg</emphasis>
. (
<quantity id="F32B81847A32FFC50F42FD605E9602CC" box="[1390,1436,691,717]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="7" pageNumber="283" unit="in" value="6.0">6 in</quantity>
INPA-alcohol).
<collectingCountry id="4CC46CF17A32FFC50B61FD045ACD02F0" box="[333,455,727,753]" name="Guyana" pageId="7" pageNumber="283">GUYANA</collectingCountry>
.
<emphasis id="06A7F0737A32FFC50BFAFD04593502F1" box="[470,575,726,752]" italics="true" pageId="7" pageNumber="283">Region 6</emphasis>
, Upper Berbice, Base camp
<quantity id="F32B81847A32FFC509BAFD0458CF02F0" box="[918,965,727,753]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="7" pageNumber="283" unit="in" value="1.0">1 in</quantity>
basecamp,
<date id="406D0AA17A32FFC50E7DFD055FFC02F0" box="[1105,1270,726,753]" pageId="7" pageNumber="283" value="2014-09-21" valueMax="2014-09-25" valueMin="2014-09-21">
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pinned)
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(
<figureCitation id="D4BF2A69FFD37248A7D930706F18C9C2" box="[159,267,888,914]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="3.[151,250,1946,1970]" captionStartId-1="4.[151,250,1444,1468]" captionTargetBox-0="[156,1432,777,1922]" captionTargetBox-1="[491,1095,181,1421]" captionTargetId-0="figure-299@3.[156,1432,777,1922]" captionTargetId-1="figure-19@4.[491,1095,181,1421]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 1. Cambeva damnata sp. nov., holotype, UFRJ 7026, 12967, 46.5 mm SL. (a) lateral view; (b) dorsal view; (c) ventral view." captionText-1="FIGURE 2. Ontogenetic variation of colouration in Cambeva damnata sp. nov., paratypes, UFRJ 12983: (a) 20.9 mm SL; (b) 21.4; (c) 28.3; (d) 30.7; (e) 33.9 mm SL; (f) 43.2 mm SL; (g) 44.3 mm SL; (h) 53.5 mm SL." pageId="1" pageNumber="427">Figs. 12</figureCitation>
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<taxonomicName id="8B844D6FFFD37248A7D130B16F3CC980" box="[151,303,952,977]" class="Actinopterygii" family="Trichomycteridae" genus="Cambeva" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="427" phylum="Chordata" rank="species" species="undefined-5">
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sp. 5
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(
<bibRefCitation id="28154B1DFFD37248A67A30B16FFCC981" author="Costa, W. J. E. M. &amp; Mattos, J. L. O. &amp; Azevedo-Santos, V. M. &amp; Feltrin, C. R. M. &amp; Amorim, P. F. &amp; Ottoni, F. P. &amp; Vilardo, P. J. &amp; Katz, A. M." box="[316,495,953,977]" pageId="1" pageNumber="427" refId="ref4915" refString="Costa, W. J. E. M., Mattos, J. L. O., Azevedo-Santos, V. M., Feltrin, C. R. M., Amorim, P. F., Ottoni, F. P., Vilardo, P. J. &amp; Katz, A. M. (2024) A poorly known catfish clade in an endangered Neotropical biodiversity hotspot: relationships and distribution patterns of the Cambeva variegata group (Siluriformes: Trichomycteridae). Fishes, 9, 116. https: // doi. org / 10.3390 / fishes 9040116" type="journal volume" year="2024">
Costa
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2024
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: fig. 1 [relationships], fig. 2 [cephalic latero-sensory system], figs. 3C, F, 4C, 5C, 7E, G, 8C [osteology], fig. 6 [distribution]).
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<location id="495B6037FFD37248A6F637356C6FCE07" LSID="urn:lsid:plazi:treatment:C42D87FAFFD3724FA7D130386FD0C89D:495B6037FFD37248A6F637356C6FCE07" box="[432,636,1085,1111]" country="Brazil" county="Nova Lima Municipality" latitude="-19.974722" longLatPrecision="21" longitude="-43.860832" municipality="Corrego do Jambreiro" name="Rio Sao Francisco" pageId="1" pageNumber="427" stateProvince="Minas Gerais">Rio São Francisco</location>
basin, in the treck to
<location id="495B6037FFD37248A42C37356A60CE07" LSID="urn:lsid:plazi:treatment:C42D87FAFFD3724FA7D130386FD0C89D:495B6037FFD37248A42C37356A60CE07" box="[874,1139,1085,1112]" country="Brazil" county="Nova Lima Municipality" latitude="-19.974722" longLatPrecision="21" longitude="-43.860832" municipality="Corrego do Jambreiro" name="Cachoeira do Boa Vista" pageId="1" pageNumber="427" stateProvince="Minas Gerais">Cachoeira do Boa Vista</location>
,
<geoCoordinate id="29B0502BFFD37248A33B37356B11CE08" box="[1149,1282,1085,1112]" degrees="19" direction="south" minutes="58" orientation="latitude" pageId="1" pageNumber="427" precision="15" seconds="29" value="-19.974722">19°5829”S</geoCoordinate>
<geoCoordinate id="29B0502BFFD37248A24E37356B84CE08" box="[1288,1431,1085,1112]" degrees="43" direction="west" minutes="51" orientation="longitude" pageId="1" pageNumber="427" precision="15" seconds="39" value="-43.860832">43°5139”W</geoCoordinate>
, about
<elevation id="C7A9D1DFFFD37248A79A37696F5FCE2B" box="[220,332,1121,1148]" metricMagnitude="2" metricUnit="m" metricValue="7.65" pageId="1" pageNumber="427" unit="m" value="765.0">
<quantity id="8B7C9B09FFD37248A79A37696F30CE2B" box="[220,291,1121,1148]" metricMagnitude="2" metricUnit="m" metricValue="7.65" pageId="1" pageNumber="427" unit="m" value="765.0">765 m</quantity>
asl
</elevation>
;
<collectorName id="E171533AFFD37248A61137696FC2CE2B" box="[343,465,1121,1147]" pageId="1" pageNumber="427">A.M. Katz</collectorName>
,
<collectorName id="E171533AFFD37248A69D37696C4ACE2B" box="[475,601,1121,1147]" pageId="1" pageNumber="427">F. P. Ottoni</collectorName>
and
<collectorName id="E171533AFFD37248A5C937696D0ACE2B" box="[655,793,1121,1147]" pageId="1" pageNumber="427">P. J. Vilardo</collectorName>
,
<date id="383A102CFFD37248A46237696DAFCE2B" box="[804,956,1121,1147]" pageId="1" pageNumber="427" value="2022-06-11">
<collectingDate id="287EE9C4FFD37248A46237696DAFCE2B" box="[804,956,1121,1147]" pageId="1" pageNumber="427" value="2022-06-11">11 June 2022</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="4C3B36ECFFD37248A781378D6F27CF01" blockId="1.[151,1437,1049,2041]" pageId="1" pageNumber="427">
<materialsCitation id="FCEC3CB1FFD37248A781378D6F22CF01" collectingDate="2022-06-12" collectorName="Rio das Velhas" country="Brazil" elevation="730" latitude="-19.97389" location="Rio Sao Francisco" longLatPrecision="21" longitude="-43.796387" municipality="Ribeirao da Prata" pageId="1" pageNumber="427" specimenCount="76" stateProvince="Minas Gerais" typeStatus="paratype">
<emphasis id="7EF0EAFEFFD37248A781378D6F5BCECF" bold="true" box="[199,328,1157,1183]" pageId="1" pageNumber="427">
<typeStatus id="933F884EFFD37248A781378D6F50CECF" box="[199,323,1157,1183]" pageId="1" pageNumber="427" type="paratype">Paratypes</typeStatus>
:
</emphasis>
UFRJ 12967,
<specimenCount id="5A82FD65FFD37248A6AF378D6C32CECF" box="[489,545,1157,1183]" count="6" pageId="1" pageNumber="427" type="generic">6 ex.</specimenCount>
, 18.0
<quantity id="8B7C9B09FFD37248A52D378C6CDDCECF" box="[619,718,1156,1183]" metricMagnitude="-2" metricUnit="m" metricValue="3.55" pageId="1" pageNumber="427" unit="mm" value="35.5">35.5 mm</quantity>
SL; UFRJ 12968,
<specimenCount id="5A82FD65FFD37248A4DB378D6DF0CECF" box="[925,995,1157,1183]" count="22" pageId="1" pageNumber="427" type="generic">22 ex.</specimenCount>
,
<quantity id="8B7C9B09FFD37248A4A8378C6A83CECF" box="[1006,1168,1156,1183]" metricMagnitude="-2" metricUnit="m" metricValue="3.0799999999999996" metricValueMax="4.54" metricValueMin="1.6199999999999999" pageId="1" pageNumber="427" unit="mm" value="30.799999999999997" valueMax="45.4" valueMin="16.2">16.245.4 mm</quantity>
SL; UFRJ 12983,
<specimenCount id="5A82FD65FFD37248A226378D6B84CECF" box="[1376,1431,1157,1183]" count="9" pageId="1" pageNumber="427" type="generic">9 ex.</specimenCount>
,
<quantity id="8B7C9B09FFD37248A7D137A06F28CE93" box="[151,315,1192,1219]" metricMagnitude="-2" metricUnit="m" metricValue="3.72" metricValueMax="5.35" metricValueMin="2.09" pageId="1" pageNumber="427" unit="mm" value="37.2" valueMax="53.5" valueMin="20.9">20.953.5 mm</quantity>
SL; UFRJ 13654,
<specimenCount id="5A82FD65FFD37248A55537A16C58CE93" box="[531,587,1193,1219]" count="4" pageId="1" pageNumber="427" type="generic">4 ex.</specimenCount>
(C&amp;S),
<quantity id="8B7C9B09FFD37248A5EA37A06D42CE93" box="[684,849,1192,1219]" metricMagnitude="-2" metricUnit="m" metricValue="4.435" metricValueMax="4.65" metricValueMin="4.22" pageId="1" pageNumber="427" unit="mm" value="44.35" valueMax="46.5" valueMin="42.2">42.246.5 mm</quantity>
SL; CICCAA 07950,
<specimenCount id="5A82FD65FFD37248A30937A16A85CE93" box="[1103,1174,1193,1219]" count="10" pageId="1" pageNumber="427" type="generic">10 ex.</specimenCount>
, 26.0
<quantity id="8B7C9B09FFD37248A3A537A06B5BCE93" box="[1251,1352,1192,1219]" metricMagnitude="-2" metricUnit="m" metricValue="4.59" pageId="1" pageNumber="427" unit="mm" value="45.9">45.9 mm</quantity>
SL; all collected with the
<typeStatus id="933F884EFFD37248A62037C46FDFCEB6" box="[358,460,1228,1254]" pageId="1" pageNumber="427" type="holotype">holotype</typeStatus>
. UFRJ 12965,
<specimenCount id="5A82FD65FFD37248A53537C56CBFCEB7" box="[627,684,1229,1255]" count="7" pageId="1" pageNumber="427" type="generic">7 ex.</specimenCount>
,
<quantity id="8B7C9B09FFD37248A5FE37C46D48CEB7" box="[696,859,1228,1255]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" metricValueMax="3.06" metricValueMin="1.54" pageId="1" pageNumber="427" unit="mm" value="23.0" valueMax="30.6" valueMin="15.4">15.430.6 mm</quantity>
SL; UFRJ 12966,
<specimenCount id="5A82FD65FFD37248A36837C46A66CEB7" box="[1070,1141,1228,1255]" count="15" pageId="1" pageNumber="427" type="generic">15 ex.</specimenCount>
, 26.0
<quantity id="8B7C9B09FFD37248A38737C46B37CEB7" box="[1217,1316,1228,1255]" metricMagnitude="-2" metricUnit="m" metricValue="3.96" pageId="1" pageNumber="427" unit="mm" value="39.6">39.6 mm</quantity>
SL; UFRJ 12982,
<specimenCount id="5A82FD65FFD37248A7A837F86F39CF5A" box="[238,298,1264,1290]" count="3" pageId="1" pageNumber="427" type="generic">3 ex.</specimenCount>
,
<quantity id="8B7C9B09FFD37248A67F37F86FCCCF5A" box="[313,479,1264,1290]" metricMagnitude="-2" metricUnit="m" metricValue="3.9299999999999997" metricValueMax="4.64" metricValueMin="3.22" pageId="1" pageNumber="427" unit="mm" value="39.3" valueMax="46.4" valueMin="32.2">32.246.4 mm</quantity>
SL;
<collectingCountry id="3493767CFFD37248A55D37F86C77CF5A" box="[539,612,1264,1290]" name="Brazil" pageId="1" pageNumber="427">Brazil</collectingCountry>
:
<collectingRegion id="8E40F80EFFD37248A53437F86D42CF5A" box="[626,849,1264,1290]" country="Brazil" name="Minas Gerais" pageId="1" pageNumber="427">Minas Gerais State</collectingRegion>
: Raposos Municipality:
<collectingMunicipality id="AC5FAC96FFD37248A32D37F86B25CF5A" box="[1131,1334,1264,1290]" pageId="1" pageNumber="427">Ribeirão da Prata</collectingMunicipality>
,
<collectorName id="E171533AFFD37248A20237F86EF1CF7E" pageId="1" pageNumber="427">Rio das Velhas</collectorName>
drainage,
<location id="495B6037FFD37248A61E361C6C35CF7E" LSID="urn:lsid:plazi:treatment:C42D87FAFFD3724FA7D130386FD0C89D:495B6037FFD37248A61E361C6C35CF7E" box="[344,550,1300,1326]" country="Brazil" latitude="-19.97389" longLatPrecision="21" longitude="-43.796387" municipality="Ribeirao da Prata" name="Rio Sao Francisco" pageId="1" pageNumber="427" stateProvince="Minas Gerais">Rio São Francisco</location>
basin,
<geoCoordinate id="29B0502BFFD37248A531361B6CE8CF7E" box="[631,763,1299,1326]" degrees="19" direction="south" minutes="58" orientation="latitude" pageId="1" pageNumber="427" precision="15" seconds="26" value="-19.97389">19°5826”S</geoCoordinate>
<geoCoordinate id="29B0502BFFD37248A445361C6D81CF7E" box="[771,914,1300,1326]" degrees="43" direction="west" minutes="47" orientation="longitude" pageId="1" pageNumber="427" precision="15" seconds="47" value="-43.796387">43°4747”W</geoCoordinate>
, about
<elevation id="C7A9D1DFFFD37248A4A5361C6A40CF7E" box="[995,1107,1300,1326]" metricMagnitude="2" metricUnit="m" metricValue="7.3" pageId="1" pageNumber="427" unit="m" value="730.0">
<quantity id="8B7C9B09FFD37248A4A5361C6A39CF7E" box="[995,1066,1300,1326]" metricMagnitude="2" metricUnit="m" metricValue="7.3" pageId="1" pageNumber="427" unit="m" value="730.0">730 m</quantity>
asl
</elevation>
; same collectors as
<typeStatus id="933F884EFFD37248A275361C6B8BCF7E" box="[1331,1432,1300,1326]" pageId="1" pageNumber="427" type="holotype">holotype</typeStatus>
,
<date id="383A102CFFD37248A7D1363F6F22CF01" box="[151,305,1335,1362]" pageId="1" pageNumber="427" value="2022-06-12">
<collectingDate id="287EE9C4FFD37248A7D1363F6F22CF01" box="[151,305,1335,1362]" pageId="1" pageNumber="427" value="2022-06-12">12 June 2022</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="049E6567FFD37248A78136536A32CD72" pageId="1" pageNumber="427" type="diagnosis">
<paragraph id="4C3B36ECFFD37248A78136536A32CD72" blockId="1.[151,1437,1049,2041]" pageId="1" pageNumber="427">
<emphasis id="7EF0EAFEFFD37248A78136536F57CF25" bold="true" box="[199,324,1371,1397]" pageId="1" pageNumber="427">Diagnosis:</emphasis>
<taxonomicName id="8B844D6FFFD37248A60C36536C0ECF25" authorityName="Costa &amp; Azevedo-Santos &amp; Ottoni &amp; Vilardo &amp; Katz" authorityYear="2024" box="[330,541,1371,1397]" class="Actinopterygii" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="427" phylum="Chordata" rank="species" species="damnata">
<emphasis id="7EF0EAFEFFD37248A60C36536C0ECF25" box="[330,541,1371,1397]" italics="true" pageId="1" pageNumber="427">Cambeva damnata</emphasis>
</taxonomicName>
is distinguished from all other congeners, except species of the CVG clade (
<emphasis id="7EF0EAFEFFD37248A23536546B85CF25" box="[1395,1430,1372,1397]" italics="true" pageId="1" pageNumber="427">i.e.</emphasis>
,
<taxonomicName id="8B844D6FFFD37248A7D136776F0ECFC9" authorityName="Costa" authorityYear="1992" box="[151,285,1407,1433]" class="Actinopterygii" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="427" phylum="Chordata" rank="species" species="concolor">
<emphasis id="7EF0EAFEFFD37248A7D136776F0ECFC9" box="[151,285,1407,1433]" italics="true" pageId="1" pageNumber="427">C. concolor</emphasis>
</taxonomicName>
and
<taxonomicName id="8B844D6FFFD37248A61336776FF1CFC9" authorityName="Costa" authorityYear="1992" box="[341,482,1407,1433]" class="Actinopterygii" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="427" phylum="Chordata" rank="species" species="variegata">
<emphasis id="7EF0EAFEFFD37248A61336776FF1CFC9" box="[341,482,1407,1433]" italics="true" pageId="1" pageNumber="427">C. variegata</emphasis>
</taxonomicName>
), by the presence of a prominent adipose-like skin crest on the dorsal margin of the caudal peduncle (vs. skin crest absent), an interrupted supraorbital canal (vs. continuous), and a small premaxilla, smaller than the maxilla, with an accentuated constriction on its lateral portion (vs. premaxilla larger than the maxilla or with similar length, without a lateral constriction).
<taxonomicName id="8B844D6FFFD37248A5B036E26DD9CC54" authorityName="Costa &amp; Azevedo-Santos &amp; Ottoni &amp; Vilardo &amp; Katz" authorityYear="2024" box="[758,970,1514,1540]" class="Actinopterygii" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="427" phylum="Chordata" rank="species" species="damnata">
<emphasis id="7EF0EAFEFFD37248A5B036E26DD9CC54" box="[758,970,1514,1540]" italics="true" pageId="1" pageNumber="427">Cambeva damnata</emphasis>
</taxonomicName>
differs from all other species of CVG by having more interopercular odontodes (3744 vs. 2631); from
<taxonomicName id="8B844D6FFFD37248A42A35066DE0CC78" authorityName="Costa" authorityYear="1992" box="[876,1011,1550,1576]" class="Actinopterygii" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="427" phylum="Chordata" rank="species" species="concolor">
<emphasis id="7EF0EAFEFFD37248A42A35066DE0CC78" box="[876,1011,1550,1576]" italics="true" pageId="1" pageNumber="427">C. concolor</emphasis>
</taxonomicName>
and
<taxonomicName id="8B844D6FFFD37248A36D35066AABCC78" authorityName="Costa" authorityYear="1992" box="[1067,1208,1550,1576]" class="Actinopterygii" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="427" phylum="Chordata" rank="species" species="variegata">
<emphasis id="7EF0EAFEFFD37248A36D35066AABCC78" box="[1067,1208,1550,1576]" italics="true" pageId="1" pageNumber="427">C. variegata</emphasis>
</taxonomicName>
by having a minute pectoral-fin filament, about 10 % of the pectoral-fin length or less (vs. about 2030 %) and the dorsal-fin origin at the vertical through the centrum of the 16th vertebra (vs. 17th); from
<taxonomicName id="8B844D6FFFD37248A4D1355E6A08CC3F" authorityName="Costa" authorityYear="1992" box="[919,1051,1621,1648]" class="Actinopterygii" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="427" phylum="Chordata" rank="species" species="concolor">
<emphasis id="7EF0EAFEFFD37248A4D1355E6A08CC3F" box="[919,1051,1621,1648]" italics="true" pageId="1" pageNumber="427">C. concolor</emphasis>
</taxonomicName>
by having a spotted colour pattern (vs. homogeneous pale yellow) and a longer pectoral fin (pectoral-fin length 15.117.7 % SL vs. 12.313.7 % SL); and from
<taxonomicName id="8B844D6FFFD37248A64435956F9FCCE6" authorityName="Costa" authorityYear="1992" box="[258,396,1693,1719]" class="Actinopterygii" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="427" phylum="Chordata" rank="species" species="variegata">
<emphasis id="7EF0EAFEFFD37248A64435956F9FCCE6" box="[258,396,1693,1719]" italics="true" pageId="1" pageNumber="427">C. variegata</emphasis>
</taxonomicName>
by having a more slender body, more conspicuous in specimens between about 45 and
<quantity id="8B7C9B09FFD37248A20935946B8ECCE7" box="[1359,1437,1692,1719]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="427" unit="mm" value="50.0">50 mm</quantity>
SL (body depth 12.814.8 % SL in
<taxonomicName id="8B844D6FFFD37248A56035C96CBACC8A" authorityName="Costa &amp; Azevedo-Santos &amp; Ottoni &amp; Vilardo &amp; Katz" authorityYear="2024" box="[550,681,1728,1755]" class="Actinopterygii" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="427" phylum="Chordata" rank="species" species="damnata">
<emphasis id="7EF0EAFEFFD37248A56035C96CBACC8A" box="[550,681,1728,1755]" italics="true" pageId="1" pageNumber="427">C. damnata</emphasis>
</taxonomicName>
vs. 16.621.2 % SL in
<taxonomicName id="8B844D6FFFD37248A4F435C96A2DCC8A" authorityName="Costa" authorityYear="1992" box="[946,1086,1729,1755]" class="Actinopterygii" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="427" phylum="Chordata" rank="species" species="variegata">
<emphasis id="7EF0EAFEFFD37248A4F435C96A2DCC8A" box="[946,1086,1729,1755]" italics="true" pageId="1" pageNumber="427">C. variegata</emphasis>
</taxonomicName>
), and no sexual dimorphism in colour pattern (vs. sexual dimorphism in colour pattern, with males paler, without large dark brown blotches, and females with large darker spots highly contrasting with light yellow interspaces).
</paragraph>
</subSubSection>
<subSubSection id="049E6567FFD3724AA78134236B75C8F9" lastPageId="3" lastPageNumber="429" pageId="1" pageNumber="427" type="description">
<paragraph id="4C3B36ECFFD37248A78134236B1BCDA9" blockId="1.[151,1437,1049,2041]" pageId="1" pageNumber="427">
<emphasis id="7EF0EAFEFFD37248A78134236F48CD15" bold="true" box="[199,347,1835,1861]" pageId="1" pageNumber="427">Description:</emphasis>
Morphometric data in
<tableCitation id="01060357FFD37248A52434246CA4CD16" box="[610,695,1836,1862]" captionStart="TABLE 1" captionStartId="2.[152,244,151,175]" captionText="TABLE 1. Morphometric data of Cambeva damnata sp. nov." pageId="1" pageNumber="427">Table 1</tableCitation>
. Body relatively slender, anterior portion of trunk subcylindrical, posterior one compressed. Greatest body depth situated at midway of distance between pectoral and the pelvic-fin bases. Dorsal profile of trunk slightly convex between nape and dorsal-fin origin, about straight at dorsal-fin base insertion, weakly convex on caudal peduncle. Ventral profile of trunk approximately straight. Anus and urogenital papilla opening located at vertical through area just anterior to middle of dorsal-fin base. Lateral line canal of trunk short, restricted to anterior-most part of flank, with two pores. Thirty-five or 36 vertebrae and 11 ribs.
</paragraph>
<caption id="18FB6664FFD0724BA7DE339F6D38CAE0" box="[152,811,151,176]" pageId="2" pageNumber="428" startId="2.[152,244,151,175]" targetType="table">
<paragraph id="4C3B36ECFFD0724BA7DE339F6D38CAE0" blockId="2.[152,811,151,176]" box="[152,811,151,176]" pageId="2" pageNumber="428">
<emphasis id="7EF0EAFEFFD0724BA7DE339F6F1CCAFF" bold="true" box="[152,271,151,175]" pageId="2" pageNumber="428">TABLE 1.</emphasis>
Morphometric data of
<taxonomicName id="8B844D6FFFD0724BA540339F6CDCCAFF" authorityName="Costa &amp; Azevedo-Santos &amp; Ottoni &amp; Vilardo &amp; Katz" authorityYear="2024" box="[518,719,151,175]" class="Actinopterygii" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="428" phylum="Chordata" rank="species" species="damnata" status="sp. nov.">
<emphasis id="7EF0EAFEFFD0724BA540339F6CDCCAFF" box="[518,719,151,175]" italics="true" pageId="2" pageNumber="428">Cambeva damnata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65C35785FFD0724BA59033906D38CAE0" box="[726,811,152,176]" pageId="2" pageNumber="428" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="4C3B36ECFFD0724BA53833C86AD9CE17" pageId="2" pageNumber="428">
<table id="3E84C44CFFD08DB6A7E533C86AD9CE17" box="[163,1226,192,1095]" gridcols="4" gridrows="22" pageId="2" pageNumber="428">
<tr id="F2B434AEFFD08DB6A7E533C86AD9CA88" box="[163,1226,192,216]" gridrow="0" pageId="2" pageNumber="428" rowspan-0="1">
<th id="B1655DD2FFD08DB6A53833C86CCFCA88" box="[638,732,192,216]" gridcol="1" gridrow="0" pageId="2" pageNumber="428">Holotype</th>
<th id="B1655DD2FFD08DB6A4D533C86A51CA88" box="[915,1090,192,216]" gridcol="2" gridrow="0" pageId="2" pageNumber="428">Paratypes (n=10)</th>
<th id="B1655DD2FFD08DB6A3EC33C86AD9CA88" box="[1194,1226,192,216]" gridcol="3" gridrow="0" pageId="2" pageNumber="428">SD</th>
</tr>
<tr id="F2B434AEFFD08DB6A7E533E46AD9CB55" box="[163,1226,236,261]" gridrow="1" pageId="2" pageNumber="428" rowspan-3="1">
<th id="B1655DD2FFD08DB6A7E533E46F9FCB55" box="[163,396,236,261]" gridcol="0" gridrow="1" pageId="2" pageNumber="428">Standard length (SL)</th>
<td id="B1655DD2FFD08DB6A53833E46CCFCB55" box="[638,732,236,261]" gridcol="1" gridrow="1" pageId="2" pageNumber="428">46.5</td>
<td id="B1655DD2FFD08DB6A4D533E46A51CB55" box="[915,1090,236,261]" gridcol="2" gridrow="1" pageId="2" pageNumber="428">39.953.5</td>
</tr>
<tr id="F2B434AEFFD08DB6A7E5322D6AD9CB6D" box="[163,1226,293,317]" gridrow="2" pageId="2" pageNumber="428">
<th id="B1655DD2FFD08DB6A7E5322D6AD9CB6D" box="[163,1226,293,317]" colspan="4" colspanRight="3" gridcol="0" gridrow="2" pageId="2" pageNumber="428">
<emphasis id="7EF0EAFEFFD0724BA7E5322D6FF4CB6D" bold="true" box="[163,487,293,317]" pageId="2" pageNumber="428">Percentage of standard length</emphasis>
</th>
</tr>
<tr id="F2B434AEFFD08DB6A7E532456AD9CB35" box="[163,1226,333,357]" gridrow="3" pageId="2" pageNumber="428">
<th id="B1655DD2FFD08DB6A7E532456F9FCB35" box="[163,396,333,357]" gridcol="0" gridrow="3" pageId="2" pageNumber="428">Body depth</th>
<td id="B1655DD2FFD08DB6A53832456CCFCB35" box="[638,732,333,357]" gridcol="1" gridrow="3" pageId="2" pageNumber="428">13.2</td>
<td id="B1655DD2FFD08DB6A4D532456A51CB35" box="[915,1090,333,357]" gridcol="2" gridrow="3" pageId="2" pageNumber="428">12.815.3</td>
<td id="B1655DD2FFD08DB6A3EC32456AD9CB35" box="[1194,1226,333,357]" gridcol="3" gridrow="3" pageId="2" pageNumber="428">0.8</td>
</tr>
<tr id="F2B434AEFFD08DB6A7E5327D6AD9CBDD" box="[163,1226,373,397]" gridrow="4" pageId="2" pageNumber="428">
<th id="B1655DD2FFD08DB6A7E5327D6F9FCBDD" box="[163,396,373,397]" gridcol="0" gridrow="4" pageId="2" pageNumber="428">Caudal peduncle depth</th>
<td id="B1655DD2FFD08DB6A538327D6CCFCBDD" box="[638,732,373,397]" gridcol="1" gridrow="4" pageId="2" pageNumber="428">11.0</td>
<td id="B1655DD2FFD08DB6A4D5327D6A51CBDD" box="[915,1090,373,397]" gridcol="2" gridrow="4" pageId="2" pageNumber="428">11.112.6</td>
<td id="B1655DD2FFD08DB6A3EC327D6AD9CBDD" box="[1194,1226,373,397]" gridcol="3" gridrow="4" pageId="2" pageNumber="428">0.6</td>
</tr>
<tr id="F2B434AEFFD08DB6A7E532956AD9CBE6" box="[163,1226,413,438]" gridrow="5" pageId="2" pageNumber="428">
<th id="B1655DD2FFD08DB6A7E532956F9FCBE6" box="[163,396,413,438]" gridcol="0" gridrow="5" pageId="2" pageNumber="428">Body width</th>
<td id="B1655DD2FFD08DB6A53832956CCFCBE6" box="[638,732,413,438]" gridcol="1" gridrow="5" pageId="2" pageNumber="428">10.2</td>
<td id="B1655DD2FFD08DB6A4D532956A51CBE6" box="[915,1090,413,438]" gridcol="2" gridrow="5" pageId="2" pageNumber="428">10.111.6</td>
<td id="B1655DD2FFD08DB6A3EC32956AD9CBE6" box="[1194,1226,413,438]" gridcol="3" gridrow="5" pageId="2" pageNumber="428">0.5</td>
</tr>
<tr id="F2B434AEFFD08DB6A7E532CE6AD9CB8E" box="[163,1226,454,478]" gridrow="6" pageId="2" pageNumber="428">
<th id="B1655DD2FFD08DB6A7E532CE6F9FCB8E" box="[163,396,454,478]" gridcol="0" gridrow="6" pageId="2" pageNumber="428">Caudal peduncle width</th>
<td id="B1655DD2FFD08DB6A53832CE6CCFCB8E" box="[638,732,454,478]" gridcol="1" gridrow="6" pageId="2" pageNumber="428">3.8</td>
<td id="B1655DD2FFD08DB6A4D532CE6A51CB8E" box="[915,1090,454,478]" gridcol="2" gridrow="6" pageId="2" pageNumber="428">3.15.2</td>
<td id="B1655DD2FFD08DB6A3EC32CE6AD9CB8E" box="[1194,1226,454,478]" gridcol="3" gridrow="6" pageId="2" pageNumber="428">0.6</td>
</tr>
<tr id="F2B434AEFFD08DB6A7E532E66AD9C856" box="[163,1226,494,518]" gridrow="7" pageId="2" pageNumber="428">
<th id="B1655DD2FFD08DB6A7E532E66F9FC856" box="[163,396,494,518]" gridcol="0" gridrow="7" pageId="2" pageNumber="428">Pre-dorsal length</th>
<td id="B1655DD2FFD08DB6A53832E66CCFC856" box="[638,732,494,518]" gridcol="1" gridrow="7" pageId="2" pageNumber="428">59.0</td>
<td id="B1655DD2FFD08DB6A4D532E66A51C856" box="[915,1090,494,518]" gridcol="2" gridrow="7" pageId="2" pageNumber="428">58.263.0</td>
<td id="B1655DD2FFD08DB6A3EC32E66AD9C856" box="[1194,1226,494,518]" gridcol="3" gridrow="7" pageId="2" pageNumber="428">1.4</td>
</tr>
<tr id="F2B434AEFFD08DB6A7E5311E6AD9C87E" box="[163,1226,534,558]" gridrow="8" pageId="2" pageNumber="428">
<th id="B1655DD2FFD08DB6A7E5311E6F9FC87E" box="[163,396,534,558]" gridcol="0" gridrow="8" pageId="2" pageNumber="428">Pre-pelvic length</th>
<td id="B1655DD2FFD08DB6A538311E6CCFC87E" box="[638,732,534,558]" gridcol="1" gridrow="8" pageId="2" pageNumber="428">53.9</td>
<td id="B1655DD2FFD08DB6A4D5311E6A51C87E" box="[915,1090,534,558]" gridcol="2" gridrow="8" pageId="2" pageNumber="428">51.659.6</td>
<td id="B1655DD2FFD08DB6A3EC311E6AD9C87E" box="[1194,1226,534,558]" gridcol="3" gridrow="8" pageId="2" pageNumber="428">2.1</td>
</tr>
<tr id="F2B434AEFFD08DB6A7E531366AD9C807" box="[163,1226,574,599]" gridrow="9" pageId="2" pageNumber="428">
<th id="B1655DD2FFD08DB6A7E531366F9FC807" box="[163,396,574,599]" gridcol="0" gridrow="9" pageId="2" pageNumber="428">Dorsal-fin base length</th>
<td id="B1655DD2FFD08DB6A53831366CCFC807" box="[638,732,574,599]" gridcol="1" gridrow="9" pageId="2" pageNumber="428">14.5</td>
<td id="B1655DD2FFD08DB6A4D531366A51C807" box="[915,1090,574,599]" gridcol="2" gridrow="9" pageId="2" pageNumber="428">13.315.5</td>
<td id="B1655DD2FFD08DB6A3EC31366AD9C807" box="[1194,1226,574,599]" gridcol="3" gridrow="9" pageId="2" pageNumber="428">0.7</td>
</tr>
<tr id="F2B434AEFFD08DB6A7E5316F6AD9C82F" box="[163,1226,615,639]" gridrow="10" pageId="2" pageNumber="428">
<th id="B1655DD2FFD08DB6A7E5316F6F9FC82F" box="[163,396,615,639]" gridcol="0" gridrow="10" pageId="2" pageNumber="428">Anal-fin base length</th>
<td id="B1655DD2FFD08DB6A538316F6CCFC82F" box="[638,732,615,639]" gridcol="1" gridrow="10" pageId="2" pageNumber="428">8.5</td>
<td id="B1655DD2FFD08DB6A4D5316F6A51C82F" box="[915,1090,615,639]" gridcol="2" gridrow="10" pageId="2" pageNumber="428">8.19.7</td>
<td id="B1655DD2FFD08DB6A3EC316F6AD9C82F" box="[1194,1226,615,639]" gridcol="3" gridrow="10" pageId="2" pageNumber="428">0.5</td>
</tr>
<tr id="F2B434AEFFD08DB6A7E531876AD9C8F7" box="[163,1226,655,679]" gridrow="11" pageId="2" pageNumber="428">
<th id="B1655DD2FFD08DB6A7E531876F9FC8F7" box="[163,396,655,679]" gridcol="0" gridrow="11" pageId="2" pageNumber="428">Caudal-fin length</th>
<td id="B1655DD2FFD08DB6A53831876CCFC8F7" box="[638,732,655,679]" gridcol="1" gridrow="11" pageId="2" pageNumber="428">18.3</td>
<td id="B1655DD2FFD08DB6A4D531876A51C8F7" box="[915,1090,655,679]" gridcol="2" gridrow="11" pageId="2" pageNumber="428">18.220.6</td>
<td id="B1655DD2FFD08DB6A3EC31876AD9C8F7" box="[1194,1226,655,679]" gridcol="3" gridrow="11" pageId="2" pageNumber="428">0.8</td>
</tr>
<tr id="F2B434AEFFD08DB6A7E531BF6AD9C89F" box="[163,1226,695,719]" gridrow="12" pageId="2" pageNumber="428">
<th id="B1655DD2FFD08DB6A7E531BF6F9FC89F" box="[163,396,695,719]" gridcol="0" gridrow="12" pageId="2" pageNumber="428">Pectoral-fin length</th>
<td id="B1655DD2FFD08DB6A53831BF6CCFC89F" box="[638,732,695,719]" gridcol="1" gridrow="12" pageId="2" pageNumber="428">15.6</td>
<td id="B1655DD2FFD08DB6A4D531BF6A51C89F" box="[915,1090,695,719]" gridcol="2" gridrow="12" pageId="2" pageNumber="428">15.117.7</td>
<td id="B1655DD2FFD08DB6A3EC31BF6AD9C89F" box="[1194,1226,695,719]" gridcol="3" gridrow="12" pageId="2" pageNumber="428">0.8</td>
</tr>
<tr id="F2B434AEFFD08DB6A7E531D76AD9C8A8" box="[163,1226,735,760]" gridrow="13" pageId="2" pageNumber="428">
<th id="B1655DD2FFD08DB6A7E531D76F9FC8A8" box="[163,396,735,760]" gridcol="0" gridrow="13" pageId="2" pageNumber="428">Pelvic-fin length</th>
<td id="B1655DD2FFD08DB6A53831D76CCFC8A8" box="[638,732,735,760]" gridcol="1" gridrow="13" pageId="2" pageNumber="428">12.2</td>
<td id="B1655DD2FFD08DB6A4D531D76A51C8A8" box="[915,1090,735,760]" gridcol="2" gridrow="13" pageId="2" pageNumber="428">11.113.2</td>
<td id="B1655DD2FFD08DB6A3EC31D76AD9C8A8" box="[1194,1226,735,760]" gridcol="3" gridrow="13" pageId="2" pageNumber="428">0.7</td>
</tr>
<tr id="F2B434AEFFD08DB6A7E530006AD9C970" box="[163,1226,776,800]" gridrow="14" pageId="2" pageNumber="428">
<th id="B1655DD2FFD08DB6A7E530006F9FC970" box="[163,396,776,800]" gridcol="0" gridrow="14" pageId="2" pageNumber="428">Head length</th>
<td id="B1655DD2FFD08DB6A53830006CCFC970" box="[638,732,776,800]" gridcol="1" gridrow="14" pageId="2" pageNumber="428">21.9</td>
<td id="B1655DD2FFD08DB6A4D530006A51C970" box="[915,1090,776,800]" gridcol="2" gridrow="14" pageId="2" pageNumber="428">21.124.5</td>
<td id="B1655DD2FFD08DB6A3EC30006AD9C970" box="[1194,1226,776,800]" gridcol="3" gridrow="14" pageId="2" pageNumber="428">0.9</td>
</tr>
<tr id="F2B434AEFFD08DB6A7E530356AD9C905" box="[163,1226,829,853]" gridrow="15" pageId="2" pageNumber="428">
<th id="B1655DD2FFD08DB6A7E530356AD9C905" box="[163,1226,829,853]" colspan="4" colspanRight="3" gridcol="0" gridrow="15" pageId="2" pageNumber="428">
<emphasis id="7EF0EAFEFFD0724BA7E530356FA9C905" bold="true" box="[163,442,829,853]" pageId="2" pageNumber="428">Percentage of head length</emphasis>
</th>
</tr>
<tr id="F2B434AEFFD08DB6A7E5306D6AD9C92E" box="[163,1226,869,894]" gridrow="16" pageId="2" pageNumber="428">
<th id="B1655DD2FFD08DB6A7E5306D6F9FC92E" box="[163,396,869,894]" gridcol="0" gridrow="16" pageId="2" pageNumber="428">Head depth</th>
<td id="B1655DD2FFD08DB6A538306D6CCFC92E" box="[638,732,869,894]" gridcol="1" gridrow="16" pageId="2" pageNumber="428">44.2</td>
<td id="B1655DD2FFD08DB6A4D5306D6A51C92E" box="[915,1090,869,894]" gridcol="2" gridrow="16" pageId="2" pageNumber="428">41.248.9</td>
<td id="B1655DD2FFD08DB6A3EC306D6AD9C92E" box="[1194,1226,869,894]" gridcol="3" gridrow="16" pageId="2" pageNumber="428">2.4</td>
</tr>
<tr id="F2B434AEFFD08DB6A7E530866AD9C9F6" box="[163,1226,910,934]" gridrow="17" pageId="2" pageNumber="428">
<th id="B1655DD2FFD08DB6A7E530866F9FC9F6" box="[163,396,910,934]" gridcol="0" gridrow="17" pageId="2" pageNumber="428">Head width</th>
<td id="B1655DD2FFD08DB6A53830866CCFC9F6" box="[638,732,910,934]" gridcol="1" gridrow="17" pageId="2" pageNumber="428">84.9</td>
<td id="B1655DD2FFD08DB6A4D530866A51C9F6" box="[915,1090,910,934]" gridcol="2" gridrow="17" pageId="2" pageNumber="428">75.887.2</td>
<td id="B1655DD2FFD08DB6A3EC30866AD9C9F6" box="[1194,1226,910,934]" gridcol="3" gridrow="17" pageId="2" pageNumber="428">3.7</td>
</tr>
<tr id="F2B434AEFFD08DB6A7E530BE6AD9C99E" box="[163,1226,950,974]" gridrow="18" pageId="2" pageNumber="428">
<th id="B1655DD2FFD08DB6A7E530BE6F9FC99E" box="[163,396,950,974]" gridcol="0" gridrow="18" pageId="2" pageNumber="428">Snout length</th>
<td id="B1655DD2FFD08DB6A53830BE6CCFC99E" box="[638,732,950,974]" gridcol="1" gridrow="18" pageId="2" pageNumber="428">47.2</td>
<td id="B1655DD2FFD08DB6A4D530BE6A51C99E" box="[915,1090,950,974]" gridcol="2" gridrow="18" pageId="2" pageNumber="428">42.547.5</td>
<td id="B1655DD2FFD08DB6A3EC30BE6AD9C99E" box="[1194,1226,950,974]" gridcol="3" gridrow="18" pageId="2" pageNumber="428">1.8</td>
</tr>
<tr id="F2B434AEFFD08DB6A7E530D66AD9C9A6" box="[163,1226,990,1014]" gridrow="19" pageId="2" pageNumber="428">
<th id="B1655DD2FFD08DB6A7E530D66F9FC9A6" box="[163,396,990,1014]" gridcol="0" gridrow="19" pageId="2" pageNumber="428">Interorbital width</th>
<td id="B1655DD2FFD08DB6A53830D66CCFC9A6" box="[638,732,990,1014]" gridcol="1" gridrow="19" pageId="2" pageNumber="428">22.6</td>
<td id="B1655DD2FFD08DB6A4D530D66A51C9A6" box="[915,1090,990,1014]" gridcol="2" gridrow="19" pageId="2" pageNumber="428">18.124.7</td>
<td id="B1655DD2FFD08DB6A3EC30D66AD9C9A6" box="[1194,1226,990,1014]" gridcol="3" gridrow="19" pageId="2" pageNumber="428">1.8</td>
</tr>
<tr id="F2B434AEFFD08DB6A7E5370E6AD9CE4F" box="[163,1226,1030,1055]" gridrow="20" pageId="2" pageNumber="428">
<th id="B1655DD2FFD08DB6A7E5370E6F9FCE4F" box="[163,396,1030,1055]" gridcol="0" gridrow="20" pageId="2" pageNumber="428">Preorbital length</th>
<td id="B1655DD2FFD08DB6A538370E6CCFCE4F" box="[638,732,1030,1055]" gridcol="1" gridrow="20" pageId="2" pageNumber="428">14.7</td>
<td id="B1655DD2FFD08DB6A4D5370E6A51CE4F" box="[915,1090,1030,1055]" gridcol="2" gridrow="20" pageId="2" pageNumber="428">12.416.9</td>
<td id="B1655DD2FFD08DB6A3EC370E6AD9CE4F" box="[1194,1226,1030,1055]" gridcol="3" gridrow="20" pageId="2" pageNumber="428">1.4</td>
</tr>
<tr id="F2B434AEFFD08DB6A7E537276AD9CE17" box="[163,1226,1071,1095]" gridrow="21" pageId="2" pageNumber="428">
<th id="B1655DD2FFD08DB6A7E537276F9FCE17" box="[163,396,1071,1095]" gridcol="0" gridrow="21" pageId="2" pageNumber="428">Eye diameter</th>
<td id="B1655DD2FFD08DB6A53837276CCFCE17" box="[638,732,1071,1095]" gridcol="1" gridrow="21" pageId="2" pageNumber="428">11.1</td>
<td id="B1655DD2FFD08DB6A4D537276A51CE17" box="[915,1090,1071,1095]" gridcol="2" gridrow="21" pageId="2" pageNumber="428">10.916.8</td>
<td id="B1655DD2FFD08DB6A3EC37276AD9CE17" box="[1194,1226,1071,1095]" gridcol="3" gridrow="21" pageId="2" pageNumber="428">1.8</td>
</tr>
</table>
</paragraph>
<paragraph id="4C3B36ECFFD0724BA78137956FA5CC13" blockId="2.[151,1437,1181,2035]" pageId="2" pageNumber="428">Head sub-trapezoidal in dorsal view, with anterior profile of snout slightly convex in dorsal view. Eye moderately large, dorsally positioned on head, nearer posterior margin of opercle than snout tip. Distance between anterior and posterior nostrils shorter than distance between posterior nostril and orbital rim. Barbels moderately long; tip of nasal barbel reaching area between orbit and opercular patch of odontodes, tip of maxillary and rictal barbels reaching posterior portion of interopercular patch of odontodes or area just posterior to it. Mouth subterminal. Lateral fleshy lobes of mouth relatively large, its largest length about half length of lower jaw excluding lobes. Jaw teeth incisiform, arranged in two rows. Total of 1115 teeth on outer premaxillary row, 12 or 13 on inner row, nine or ten teeth on outer dentary row, 1720 on inner row. Ventral surface of head covered by minute skin papillae. Branchial membrane thickened, attached to isthmus only at its anterior-most point, in ventral midline. Total of seven branchiostegal rays. Odontodes always conical, straight and arranged in irregular longitudinal in interopercle, straight to slightly curved and arranged in transverse rows in opercle. Total of 1315 opercular odontodes, 3744 interopercular odontodes.</paragraph>
<paragraph id="4C3B36ECFFD0724AA78135456F9BCA85" blockId="2.[151,1437,1181,2035]" lastBlockId="3.[151,1437,150,753]" lastPageId="3" lastPageNumber="429" pageId="2" pageNumber="428">Fins well-developed, with thickened bases. Dorsal fin relatively long and deep, subtriangular, with 12 (ii + II + 8) rays. Dorsal-fin origin at midway between nape and caudal-fin base. Anal fin smaller than the dorsal fin, subtriangular, with nine (ii + II + 5) rays. Anal-fin origin at vertical through posterior portion of dorsal-fin base, at base of sixth branched ray. Base of first dorsal-fin ray at vertical through centrum of 16th vertebra, base of first anal-fin ray at vertical through centrum of 22nd vertebra. Pectoral fin subtriangular in dorsal view. First pectoral-fin slightly extending beyond fin membrane to form short terminal filament, its length about 10 % of pectoral-fin length or less. Eight (I + 7) pectoral-fin rays. Pelvic fin sub-oval, its posterior extremity reaching vertical through middle of dorsal-fin base, surpassing anus position. Pelvic-fin bases medially separated by interspace corresponding to about half pelvic-fin base width. Five (I + 4) pelvic-fin rays. Caudal fin subtruncate with posterior corners rounded. Thirteen (I + 11 + I) principal caudal-fin rays, 1719 (xvi-xviii + I) dorsal procurrent rays, and nine or ten (viiiix + I) ventral procurrent rays; one or two loose minute rays positioned anteriorly to dorsal procurrent rays, within prominent skin crest on dorsal surface of caudal peduncle. Caudal skeleton comprising two triangular hypural plates, dorsal plate, corresponding to the hypural bones 3 + 4 + 5, and ventral plate corresponding to hypural bones 1 + 2 and parhypural.</paragraph>
<paragraph id="4C3B36ECFFD1724AA78133D76F90C8D5" blockId="3.[151,1437,150,753]" pageId="3" pageNumber="429">
<emphasis id="7EF0EAFEFFD1724AA78133D76F89CAA9" box="[199,410,223,249]" italics="true" pageId="3" pageNumber="429">Colouration in life</emphasis>
(
<figureCitation id="D4BF2A69FFD1724AA6EC33D76FE3CAA9" box="[426,496,223,249]" captionStart="FIGURE 2" captionStartId="4.[151,250,1444,1468]" captionTargetBox="[491,1095,181,1421]" captionTargetId="figure-19@4.[491,1095,181,1421]" captionTargetPageId="4" captionText="FIGURE 2. Ontogenetic variation of colouration in Cambeva damnata sp. nov., paratypes, UFRJ 12983: (a) 20.9 mm SL; (b) 21.4; (c) 28.3; (d) 30.7; (e) 33.9 mm SL; (f) 43.2 mm SL; (g) 44.3 mm SL; (h) 53.5 mm SL." pageId="3" pageNumber="429">Fig. 2</figureCitation>
). In specimens above about
<quantity id="8B7C9B09FFD1724AA47533D76D90CAA9" box="[819,899,223,249]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="429" unit="mm" value="40.0">40 mm</quantity>
SL (
<figureCitation id="D4BF2A69FFD1724AA4FC33D76A38CAA9" box="[954,1067,223,249]" captionStart="FIGURE 2" captionStartId="4.[151,250,1444,1468]" captionTargetBox="[491,1095,181,1421]" captionTargetId="figure-19@4.[491,1095,181,1421]" captionTargetPageId="4" captionText="FIGURE 2. Ontogenetic variation of colouration in Cambeva damnata sp. nov., paratypes, UFRJ 12983: (a) 20.9 mm SL; (b) 21.4; (c) 28.3; (d) 30.7; (e) 33.9 mm SL; (f) 43.2 mm SL; (g) 44.3 mm SL; (h) 53.5 mm SL." pageId="3" pageNumber="429">Figs 2fh</figureCitation>
), flank light yellowish grey with large dark brown blotches, irregularly shaped, often coalesced with neighbouring spots. Dorsum predominantly dark brown, contrasting with light yellowish grey spots. Venter and ventral surface of head yellowish white, without vestige of dark pigmentation. Dorsal and lateral regions of head predominantly dark brown, with unpigmented area on infraorbital region, extending to entire interopercular patch of odontodes and posterior portion of opercular patch of odontodes. Nasal barbel dark grey, maxillary and rictal barbels light grey. Iris dark brown, with narrow yellow circle around pupil. Fins hyaline, with concentration of melanophores on pectoral and unpaired fins, and anterobasal portion of adipose-like skin crest of caudal peduncle. Specimens between about 20 and
<quantity id="8B7C9B09FFD1724AA3E232D26AE1CBA5" box="[1188,1266,474,501]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="429" unit="mm" value="35.0">35 mm</quantity>
SL (
<figureCitation id="D4BF2A69FFD1724AA26032D36B87CBA5" box="[1318,1428,475,501]" captionStart="FIGURE 2" captionStartId="4.[151,250,1444,1468]" captionTargetBox="[491,1095,181,1421]" captionTargetId="figure-19@4.[491,1095,181,1421]" captionTargetPageId="4" captionText="FIGURE 2. Ontogenetic variation of colouration in Cambeva damnata sp. nov., paratypes, UFRJ 12983: (a) 20.9 mm SL; (b) 21.4; (c) 28.3; (d) 30.7; (e) 33.9 mm SL; (f) 43.2 mm SL; (g) 44.3 mm SL; (h) 53.5 mm SL." pageId="3" pageNumber="429">Figs 2ae</figureCitation>
) differ from larger ones by flank spots being smaller, sub-rectangular, arranged in longitudinal row on flank midline line of in another one on area between dorsum and flank, often each one alternating longitudinally with small pale golden dots. In specimens about
<quantity id="8B7C9B09FFD1724AA542314E6C41C831" box="[516,594,582,609]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="429" unit="mm" value="15.0">15 mm</quantity>
SL, dark pigmentation of body restricted to narrow black stripe along flank longitudinal midline.
</paragraph>
<paragraph id="4C3B36ECFFD1724AA78131876B75C8F9" blockId="3.[151,1437,150,753]" box="[199,1382,655,681]" pageId="3" pageNumber="429">
<emphasis id="7EF0EAFEFFD1724AA78131876FDBC8F9" box="[199,456,655,681]" italics="true" pageId="3" pageNumber="429">Colouration in alcohol</emphasis>
(
<figureCitation id="D4BF2A69FFD1724AA69131876C0AC8F9" box="[471,537,655,681]" captionStart="FIGURE 1" captionStartId="3.[151,250,1946,1970]" captionTargetBox="[156,1432,777,1922]" captionTargetId="figure-299@3.[156,1432,777,1922]" captionTargetPageId="3" captionText="FIGURE 1. Cambeva damnata sp. nov., holotype, UFRJ 7026, 12967, 46.5 mm SL. (a) lateral view; (b) dorsal view; (c) ventral view." pageId="3" pageNumber="429">Fig. 1</figureCitation>
). Similar to live specimens, but brown marks paler and golden dots absent.
</paragraph>
</subSubSection>
<subSubSection id="049E6567FFD1724AA78131BB6B7DC8A1" pageId="3" pageNumber="429" type="etymology">
<paragraph id="4C3B36ECFFD1724AA78131BB6B7DC8A1" blockId="3.[151,1437,150,753]" pageId="3" pageNumber="429">
<emphasis id="7EF0EAFEFFD1724AA78131BB6F41C89D" bold="true" box="[199,338,691,717]" pageId="3" pageNumber="429">Etymology:</emphasis>
From the Latin, the name
<taxonomicName id="8B844D6FFFD1724AA53D31BB6CCEC89D" authorityName="Costa &amp; Azevedo-Santos &amp; Ottoni &amp; Vilardo &amp; Katz" authorityYear="2024" box="[635,733,691,717]" class="Actinopterygii" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="429" phylum="Chordata" rank="species" species="damnata">
<emphasis id="7EF0EAFEFFD1724AA53D31BB6CCEC89D" box="[635,733,691,717]" italics="true" pageId="3" pageNumber="429">damnata</emphasis>
</taxonomicName>
means condemned and refers to the risks that the species faces with the plans for mining activities at the Serra do Curral and urban expansion (see conservation notes below).
</paragraph>
</subSubSection>
<caption id="18FB6664FFD1724AA7D134926EDDCD86" pageId="3" pageNumber="429" startId="3.[151,250,1946,1970]" targetBox="[156,1432,777,1922]" targetPageId="3" targetType="figure">
<paragraph id="4C3B36ECFFD1724AA7D134926EDDCD86" blockId="3.[151,1436,1946,2006]" pageId="3" pageNumber="429">
<emphasis id="7EF0EAFEFFD1724AA7D134926F01CDE2" bold="true" box="[151,274,1946,1970]" pageId="3" pageNumber="429">FIGURE 1.</emphasis>
<taxonomicName id="8B844D6FFFD1724AA65134926FC5CDE2" authorityName="Costa &amp; Azevedo-Santos &amp; Ottoni &amp; Vilardo &amp; Katz" authorityYear="2024" box="[279,470,1946,1970]" class="Actinopterygii" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="429" phylum="Chordata" rank="species" species="damnata" status="sp. nov.">
<emphasis id="7EF0EAFEFFD1724AA65134926FC5CDE2" box="[279,470,1946,1970]" italics="true" pageId="3" pageNumber="429">Cambeva damnata</emphasis>
</taxonomicName>
<emphasis id="7EF0EAFEFFD1724AA69D34926C39CDE2" bold="true" box="[475,554,1946,1970]" pageId="3" pageNumber="429">
<taxonomicNameLabel id="65C35785FFD1724AA69D34926C39CDE2" box="[475,554,1946,1970]" pageId="3" pageNumber="429" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, UFRJ 7026, 12967, 46.5 mm SL. (a) lateral view; (b) dorsal view; (c) ventral view.
</paragraph>
</caption>
<caption id="18FB6664FFD6724DA7D136AC6A2FCFB1" pageId="4" pageNumber="430" startId="4.[151,250,1444,1468]" targetBox="[491,1095,181,1421]" targetPageId="4" targetType="figure">
<paragraph id="4C3B36ECFFD6724DA7D136AC6A2FCFB1" blockId="4.[151,1436,1444,1505]" pageId="4" pageNumber="430">
<emphasis id="7EF0EAFEFFD6724DA7D136AC6F00CFED" bold="true" box="[151,275,1444,1469]" pageId="4" pageNumber="430">FIGURE 2.</emphasis>
Ontogenetic variation of colouration in
<taxonomicName id="8B844D6FFFD6724DA5E836AD6D7ECFED" authorityName="Costa &amp; Azevedo-Santos &amp; Ottoni &amp; Vilardo &amp; Katz" authorityYear="2024" box="[686,877,1445,1469]" class="Actinopterygii" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="430" phylum="Chordata" rank="species" species="damnata" status="sp. nov.">
<emphasis id="7EF0EAFEFFD6724DA5E836AD6D7ECFED" box="[686,877,1445,1469]" italics="true" pageId="4" pageNumber="430">Cambeva damnata</emphasis>
</taxonomicName>
<emphasis id="7EF0EAFEFFD6724DA43236AD6DD6CFED" bold="true" box="[884,965,1445,1469]" pageId="4" pageNumber="430">
<taxonomicNameLabel id="65C35785FFD6724DA43236AD6DD6CFED" box="[884,965,1445,1469]" pageId="4" pageNumber="430" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratypes, UFRJ 12983: (a) 20.9 mm SL; (b) 21.4; (c) 28.3; (d) 30.7; (e) 33.9 mm SL; (f) 43.2 mm SL; (g) 44.3 mm SL; (h) 53.5 mm SL.
</paragraph>
</caption>
<subSubSection id="049E6567FFD6724FA781353B6FD0C89D" lastPageId="6" lastPageNumber="432" pageId="4" pageNumber="430" type="distribution">
<paragraph id="4C3B36ECFFD6724DA781353B6FDBCC8D" blockId="4.[151,1437,1587,2009]" pageId="4" pageNumber="430">
<emphasis id="7EF0EAFEFFD6724DA781353B6FFDCC1D" bold="true" box="[199,494,1587,1613]" pageId="4" pageNumber="430">Distribution and habitat</emphasis>
:
<taxonomicName id="8B844D6FFFD6724DA6B9353B6CC0CC1D" authorityName="Costa &amp; Azevedo-Santos &amp; Ottoni &amp; Vilardo &amp; Katz" authorityYear="2024" box="[511,723,1587,1613]" class="Actinopterygii" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="430" phylum="Chordata" rank="species" species="damnata">
<emphasis id="7EF0EAFEFFD6724DA6B9353B6CC0CC1D" box="[511,723,1587,1613]" italics="true" pageId="4" pageNumber="430">Cambeva damnata</emphasis>
</taxonomicName>
is known from two small urban streams that are tributaries of the Rio das Velhas drainage, Rio São Francisco basin, in
<collectingRegion id="8E40F80EFFD6724DA45B355F6DE5CC21" box="[797,1014,1623,1649]" country="Brazil" name="Minas Gerais" pageId="4" pageNumber="430">Minas Gerais State</collectingRegion>
, south-eastern
<collectingCountry id="3493767CFFD6724DA3D9355F6AFBCC21" box="[1183,1256,1623,1649]" name="Brazil" pageId="4" pageNumber="430">Brazil</collectingCountry>
: the Córrego do Jambreiro in the Nova Lima Municipality, the
<typeStatus id="933F884EFFD6724DA5D835736CDDCCC5" box="[670,718,1659,1685]" pageId="4" pageNumber="430">type</typeStatus>
locality, and the Ribeirão da Prata, in the Raposos Municipality. Nova Lima and Raposos are neighbouring municipalities, with these two collection sites being at a straight line distance of
<quantity id="8B7C9B09FFD6724DA65E35CB6F75CC8D" box="[280,358,1731,1757]" metricMagnitude="3" metricUnit="m" metricValue="6.7" pageId="4" pageNumber="430" unit="km" value="6.7">6.7 km</quantity>
(
<figureCitation id="D4BF2A69FFD6724DA63335CB6FA9CC8D" box="[373,442,1731,1757]" captionStart="FIGURE 3" captionStartId="5.[151,250,1036,1060]" captionTargetBox="[154,1428,185,1009]" captionTargetId="figure-30@5.[153,1434,181,1012]" captionTargetPageId="5" captionText="FIGURE 3. Geographical distribution of the Cambeva variegata species group: (1) Cambeva concolor (Costa, 1992); (2) Cambeva variegata (Costa, 1992); (3) Cambeva damnata sp. nov. Stars indicate type localities." pageId="4" pageNumber="430">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="4C3B36ECFFD6724DA78135EF6A41CD89" blockId="4.[151,1437,1587,2009]" pageId="4" pageNumber="430">
The Córrego do Jambreiro (
<figureCitation id="D4BF2A69FFD6724DA54535EF6C68CD51" box="[515,635,1767,1793]" captionStart="FIGURE 4" captionStartId="5.[151,250,1978,2002]" captionTargetBox="[257,1331,1150,1955]" captionTargetId="figure-62@5.[257,1331,1150,1955]" captionTargetPageId="5" captionText="FIGURE 4. Biotopes of Cambeva damnata sp. nov. in (a,b) Córrego do Jambreiro, Município de Nova Lima, (c,d) Ribeirão da Prata, Município de Raposos: (a, c) out of water general view; (b,d) underwater view." pageId="4" pageNumber="430">Figs. 4a, b</figureCitation>
) is located at the Serra do Curral, a small mountain area that is part of the Serra do Espinhaço. At the
<typeStatus id="933F884EFFD6724DA6BC34036C39CD75" box="[506,554,1803,1829]" pageId="4" pageNumber="430">type</typeStatus>
locality, about
<quantity id="8B7C9B09FFD6724DA59C34036D31CD75" box="[730,802,1803,1830]" metricMagnitude="2" metricUnit="m" metricValue="7.65" pageId="4" pageNumber="430" unit="m" value="765.0">765 m</quantity>
asl, the water flow is fast to moderate and the substrate is predominantly composed of gravel and rocks of different sizes. Along the stream course, there were some areas where the substrate was composed of sand and leaf litter, especially in areas with slower water flow. Along the entire collecting site, the bank was covered by riparian forest. The specimens were captured using the following collection procedure: one collector lifted and moved the rocks and gravel using his hands and feet, while another collector lifted the hand net. This suggests that the species lives buried under gravel and rocks.
</paragraph>
<caption id="18FB6664FFD7724CA7D137046A73CE18" pageId="5" pageNumber="431" startId="5.[151,250,1036,1060]" targetBox="[154,1428,185,1009]" targetPageId="5" targetType="figure">
<paragraph id="4C3B36ECFFD7724CA7D137046A73CE18" blockId="5.[151,1436,1036,1097]" pageId="5" pageNumber="431">
<emphasis id="7EF0EAFEFFD7724CA7D137046F05CE74" bold="true" box="[151,278,1036,1060]" pageId="5" pageNumber="431">FIGURE 3.</emphasis>
Geographical distribution of the
<taxonomicName id="8B844D6FFFD7724CA53F37046D50CE74" authorityName="Costa" authorityYear="1992" box="[633,835,1036,1060]" class="Actinopterygii" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="431" phylum="Chordata" rank="species" species="variegata">
<emphasis id="7EF0EAFEFFD7724CA53F37046D50CE74" box="[633,835,1036,1060]" italics="true" pageId="5" pageNumber="431">Cambeva variegata</emphasis>
</taxonomicName>
species group: (1)
<taxonomicName id="8B844D6FFFD7724CA35537046B62CE75" authority="(Costa, 1992)" baseAuthorityName="Costa" baseAuthorityYear="1992" box="[1043,1393,1036,1061]" class="Actinopterygii" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="431" phylum="Chordata" rank="species" species="concolor">
<emphasis id="7EF0EAFEFFD7724CA35537046AC4CE74" box="[1043,1239,1036,1060]" italics="true" pageId="5" pageNumber="431">Cambeva concolor</emphasis>
(
<bibRefCitation id="28154B1DFFD7724CA3AE37056B79CE75" author="Costa, W. J. E. M." box="[1256,1386,1037,1061]" pageId="5" pageNumber="431" pagination="101 - 110" refId="ref4561" refString="Costa, W. J. E. M. (1992) Description de huit nouvelles especes du genre Trichomycterus (Siluriformes: Trichomycteridae), du Bresil oriental. Revue francaise d'Aquariologie et. Herpetologie, 18, 101 - 110." type="journal article" year="1992">Costa, 1992</bibRefCitation>
)
</taxonomicName>
; (2)
<taxonomicName id="8B844D6FFFD7724CA7D137386FE1CE19" authority="(Costa, 1992)" baseAuthorityName="Costa" baseAuthorityYear="1992" box="[151,498,1072,1097]" class="Actinopterygii" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="431" phylum="Chordata" rank="species" species="variegata">
<emphasis id="7EF0EAFEFFD7724CA7D137386F4DCE18" box="[151,350,1072,1096]" italics="true" pageId="5" pageNumber="431">Cambeva variegata</emphasis>
(
<bibRefCitation id="28154B1DFFD7724CA62A37396FF9CE19" author="Costa, W. J. E. M." box="[364,490,1073,1097]" pageId="5" pageNumber="431" pagination="101 - 110" refId="ref4561" refString="Costa, W. J. E. M. (1992) Description de huit nouvelles especes du genre Trichomycterus (Siluriformes: Trichomycteridae), du Bresil oriental. Revue francaise d'Aquariologie et. Herpetologie, 18, 101 - 110." type="journal article" year="1992">Costa, 1992</bibRefCitation>
)
</taxonomicName>
; (3)
<taxonomicName id="8B844D6FFFD7724CA56637386CCCCE18" authorityName="Costa &amp; Azevedo-Santos &amp; Ottoni &amp; Vilardo &amp; Katz" authorityYear="2024" box="[544,735,1072,1096]" class="Actinopterygii" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="431" phylum="Chordata" rank="species" species="damnata" status="sp. nov.">
<emphasis id="7EF0EAFEFFD7724CA56637386CCCCE18" box="[544,735,1072,1096]" italics="true" pageId="5" pageNumber="431">Cambeva damnata</emphasis>
</taxonomicName>
<emphasis id="7EF0EAFEFFD7724CA5A337396D26CE19" bold="true" box="[741,821,1073,1097]" pageId="5" pageNumber="431">
<taxonomicNameLabel id="65C35785FFD7724CA5A337396D26CE19" box="[741,821,1073,1097]" pageId="5" pageNumber="431" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Stars indicate type localities.
</paragraph>
</caption>
<caption id="18FB6664FFD7724CA7D134B26DE9CDA6" pageId="5" pageNumber="431" startId="5.[151,250,1978,2002]" targetBox="[257,1331,1150,1955]" targetPageId="5" targetType="figure">
<paragraph id="4C3B36ECFFD7724CA7D134B26DE9CDA6" blockId="5.[151,1436,1978,2039]" pageId="5" pageNumber="431">
<emphasis id="7EF0EAFEFFD7724CA7D134B26F01CD82" bold="true" box="[151,274,1978,2002]" pageId="5" pageNumber="431">FIGURE 4.</emphasis>
Biotopes of
<taxonomicName id="8B844D6FFFD7724CA6D534B26C41CD82" authorityName="Costa &amp; Azevedo-Santos &amp; Ottoni &amp; Vilardo &amp; Katz" authorityYear="2024" box="[403,594,1978,2002]" class="Actinopterygii" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="431" phylum="Chordata" rank="species" species="damnata" status="sp. nov.">
<emphasis id="7EF0EAFEFFD7724CA6D534B26C41CD82" box="[403,594,1978,2002]" italics="true" pageId="5" pageNumber="431">Cambeva damnata</emphasis>
</taxonomicName>
<emphasis id="7EF0EAFEFFD7724CA51E34B36CBBCD83" bold="true" box="[600,680,1979,2003]" pageId="5" pageNumber="431">
<taxonomicNameLabel id="65C35785FFD7724CA51E34B36CBBCD83" box="[600,680,1979,2003]" pageId="5" pageNumber="431" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in (a,b) Córrego do Jambreiro, Município de Nova Lima, (c,d) Ribeirão da Prata, Município de Raposos: (a, c) out of water general view; (b,d) underwater view.
</paragraph>
</caption>
<paragraph id="4C3B36ECFFD4724FA781339F6A98CB11" blockId="6.[151,1436,151,717]" pageId="6" pageNumber="432">
The collecting site at Ribeirão da Prata (
<figureCitation id="D4BF2A69FFD4724FA5D0339F6D3DCAE1" box="[662,814,151,177]" captionStart="FIGURE 4" captionStartId="5.[151,250,1978,2002]" captionTargetBox="[257,1331,1150,1955]" captionTargetId="figure-62@5.[257,1331,1150,1955]" captionTargetPageId="5" captionText="FIGURE 4. Biotopes of Cambeva damnata sp. nov. in (a,b) Córrego do Jambreiro, Município de Nova Lima, (c,d) Ribeirão da Prata, Município de Raposos: (a, c) out of water general view; (b,d) underwater view." pageId="6" pageNumber="432">Figures 4c, d</figureCitation>
) is located at
<quantity id="8B7C9B09FFD4724FA497339F6A09CAE1" box="[977,1050,151,177]" metricMagnitude="2" metricUnit="m" metricValue="7.3" pageId="6" pageNumber="432" unit="m" value="730.0">730 m</quantity>
asl, has moderate water flow and substrate composed of mud, sand, leaf litter, tree trunks, gravel and rocks, depending on the area. This site is situated in an area with more intense urbanization process, with houses occupying banks and surroundings. Consequently, sewage is dumped directly into the stream, and the stream banks are heavily deforested. The specimens were found inhabiting areas of gravel and rocks, and they were collected similarly as described above.
</paragraph>
<paragraph id="4C3B36ECFFD4724FA78132426FD0C89D" blockId="6.[151,1436,151,717]" pageId="6" pageNumber="432">
<emphasis id="7EF0EAFEFFD4724FA78132426FBDCB35" bold="true" box="[199,430,330,357]" pageId="6" pageNumber="432">Conservation notes</emphasis>
:
<taxonomicName id="8B844D6FFFD4724FA6F932436C80CB35" authorityName="Costa &amp; Azevedo-Santos &amp; Ottoni &amp; Vilardo &amp; Katz" authorityYear="2024" box="[447,659,331,357]" class="Actinopterygii" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="432" phylum="Chordata" rank="species" species="damnata">
<emphasis id="7EF0EAFEFFD4724FA6F932436C80CB35" box="[447,659,331,357]" italics="true" pageId="6" pageNumber="432">Cambeva damnata</emphasis>
</taxonomicName>
occurs in a region subject to strong pressure from human activities, such as mining, deforestation, and urban expansion. The Córrego do Jambreiro holds its headwaters in the Serra do Curral (see above), a mountain subjected to mining activities. The impoundment is known as the Serra do Taquaril Mining Complex (Complexo Minerário Serra do Taquaril, in Portuguese), and the headwaters of a tributary of Córrego do Jambreiro is very close to the area that will receive the influence of mining activities. Mining is responsible for siltation, destruction of riparian vegetation, and water contamination downstream (
<bibRefCitation id="28154B1DFFD4724FA3A532F76F0AC86D" author="Azevedo-Santos, V. M. &amp; Arcifa, M. S. &amp; Brito, M. F. &amp; Agostinho, A. A. &amp; Hughes, R. M. &amp; Vitule, J. R. S. &amp; Simberloff, D. &amp; Olden, J. D. &amp; Pelicice, F. M." pageId="6" pageNumber="432" refId="ref4277" refString="Azevedo-Santos, V. M., Arcifa, M. S., Brito, M. F., Agostinho, A. A., Hughes, R. M., Vitule, J. R. S., Simberloff, D., Olden, J. D. &amp; Pelicice, F. M. (2021) Negative impacts of mining on Neotropical freshwater fishes. Neotropical Ichthyology, 19, e 210001. https: // doi. org / 10.1590 / 1982 - 0224 - 2021 - 0001" type="journal volume" year="2021">
Azevedo-Santos
<emphasis id="7EF0EAFEFFD4724FA7D1312B6EC0C86D" box="[151,211,547,573]" italics="true" pageId="6" pageNumber="432">et al.</emphasis>
2021
</bibRefCitation>
). Therefore, plans for mining on the Serra do Curral may put populations of
<taxonomicName id="8B844D6FFFD4724FA3E7312B6B3BC86D" authorityName="Costa &amp; Azevedo-Santos &amp; Ottoni &amp; Vilardo &amp; Katz" authorityYear="2024" box="[1185,1320,547,573]" class="Actinopterygii" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="432" phylum="Chordata" rank="species" species="damnata">
<emphasis id="7EF0EAFEFFD4724FA3E7312B6B3BC86D" box="[1185,1320,547,573]" italics="true" pageId="6" pageNumber="432">C. damnata</emphasis>
</taxonomicName>
at risk of extinction. On the other hand, the Ribeirão da Prata area is under an intense process of urban expansion, which presently is a strong regional threat (
<bibRefCitation id="28154B1DFFD4724FA56931636D78C8D5" author="Azevedo-Santos, V. M. &amp; Daga, V. S. &amp; Tonella, L. H. &amp; Ruaro, R. &amp; Arcifa, M. S. &amp; Fearnside, P. M. &amp; Giarrizzo, T." box="[559,875,619,645]" pageId="6" pageNumber="432" refId="ref4375" refString="Azevedo-Santos, V. M., Daga, V. S., Tonella, L. H., Ruaro, R., Arcifa, M. S., Fearnside, P. M. &amp; Giarrizzo, T. (2023) Brazil's urban ecosystems threatened by law. Land Use Policy, 131, 106721. https: // doi. org / 10.1016 / j. landusepol. 2023.106721" type="journal volume" year="2023">
Azevedo-Santos
<emphasis id="7EF0EAFEFFD4724FA5A931636D34C8D5" box="[751,807,619,645]" italics="true" pageId="6" pageNumber="432">et al.</emphasis>
2023
</bibRefCitation>
). Therefore, since the only two localities where
<taxonomicName id="8B844D6FFFD4724FA2C531636EEAC8F9" authorityName="Costa &amp; Azevedo-Santos &amp; Ottoni &amp; Vilardo &amp; Katz" authorityYear="2024" class="Actinopterygii" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="432" phylum="Chordata" rank="species" species="damnata">
<emphasis id="7EF0EAFEFFD4724FA2C531636EEAC8F9" italics="true" pageId="6" pageNumber="432">C. damnata</emphasis>
</taxonomicName>
is known to occur are being dramatically affected by anthropogenic features, this species may be seriously threatened with extinction.
</paragraph>
</subSubSection>
</treatment>
</document>