Add updates up until 2025-01-28 17:25:08

This commit is contained in:
ggserver 2025-01-28 17:31:15 +00:00
parent a59c1084b7
commit 0cd06fffdf
6 changed files with 1192 additions and 211 deletions

View file

@ -0,0 +1,108 @@
<document id="755E4B3E1E48C8EF9954CE3332806A74" ID-DOI="10.11646/zootaxa.5570.2.8" ID-ISSN="1175-5326" ID-Zenodo-Dep="14735174" ID-ZooBank="8030475D-8EA2-4C77-AD9E-72E3F8888D08" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1737737826885" checkinUser="plazi" docAuthor="Yu, Daoyuan, Xue, Huiying &amp; Zhang, Feng" docDate="2025" docId="0398CE51FFACFF9FFF6BFCD6FD5365CB" docLanguage="en" docName="zootaxa.5570.2.8.pdf" docOrigin="Zootaxa 5570 (2)" docSource="https://doi.org/10.11646/zootaxa.5570.2.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="(Ocreatomurus) Yu 2023" docType="treatment" docVersion="4" lastPageNumber="372" masterDocId="FFA1B629FFADFF9EFFFCFF94FF886231" masterDocTitle="Two new species of Tomocerus (Ocreatomurus) Yu from Xizang, China (Collembola, Tomoceridae)" masterLastPageNumber="379" masterPageNumber="371" pageNumber="372" updateTime="1738085313822" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="2A67147BB71F5075C8269C7E0084E9C9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3B3BD661617B9E11FC76C77A5DBA7E94">
<mods:title id="519963795FCA783B36F326F51969A33A">Two new species of Tomocerus (Ocreatomurus) Yu from Xizang, China (Collembola, Tomoceridae)</mods:title>
</mods:titleInfo>
<mods:name id="1DCC64BD05A39D3B1E42F0F9B46031F0" type="personal">
<mods:role id="EB6A32290D20DCB198A90218904E84D0">
<mods:roleTerm id="50300AE5A698AA7D7FFABE1181EA0CD9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C562A79BD6B678994B20ADB48C00D238">Yu, Daoyuan</mods:namePart>
<mods:affiliation id="3303F0E55791CCD197C118856030E863">Soil Ecology Lab, Department of Ecology, College of Resources and Environmental Sciences, Nanjing Agricultural University, Nanjing, Jiangsu 210095, China</mods:affiliation>
</mods:name>
<mods:name id="4C7136FE63AAEC999AAA5FF25859D9E9" type="personal">
<mods:role id="3E37031F0BFA467EE1F2112339F4F113">
<mods:roleTerm id="95337E31FC5C75EC800F79AA3AE30BA9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="775273BCDC0DDB37AB3F0A9C6D745975">Xue, Huiying</mods:namePart>
<mods:affiliation id="3E7C2B48CA154734190FFCEEEF99FE69">College of Resources and Environment, Xizang Agriculture and Animal Husbandry University, Linzhi, Xizang 860000, China</mods:affiliation>
</mods:name>
<mods:name id="4681AD87CFC026D1E1D67902F79E1278" type="personal">
<mods:role id="47208E07858CF05E846BFACC1ED0E277">
<mods:roleTerm id="E363FD4391235197B684CB0418EDA6B2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3007BA4D1553C2930A682D3B5D441CFB">Zhang, Feng</mods:namePart>
<mods:affiliation id="D29A7670A718AF2116E851D924E69560">Department of Entomology, College of Plant Protection, Nanjing Agricultural University, Nanjing, Jiangsu 210095, China</mods:affiliation>
</mods:name>
<mods:typeOfResource id="03816078DD245D44BAE13247874709D0">text</mods:typeOfResource>
<mods:relatedItem id="2D24D88A197A14CD499C3CE16D39CA54" type="host">
<mods:titleInfo id="1ADCC0B8BF15D149E9DA1D65881620B2">
<mods:title id="AA7479B59F9EF4C3356CFDF879890F9E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="BB455E590FB1071033915406DE4FFAB9">
<mods:date id="A6527BDFBA3C8A981C8BAB82FCD4961D">2025</mods:date>
<mods:detail id="E2E357FFFDEE54E7F119EA596E91F52D" type="pubDate">
<mods:number id="9B7DD15C59BFA673D4D135203984F1DA">2025-01-17</mods:number>
</mods:detail>
<mods:detail id="DF4189DC30127251467CF53E75376EAD" type="volume">
<mods:number id="CC11D5EFF1F4E3DBC5FA8028C51C2734">5570</mods:number>
</mods:detail>
<mods:detail id="85F4DED09672357733558673A840DBA9" type="issue">
<mods:number id="10B181481C1D590836EBC0C6DC94A5EC">2</mods:number>
</mods:detail>
<mods:extent id="28887583B51BC7AA4EFC47F4DA562F32" unit="page">
<mods:start id="077A77F8286F0BD3B2F3A4D3BA4F4EF9">371</mods:start>
<mods:end id="9CAD7E77E55B3DF6192CEB88AFF9551E">379</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="1BAE7FDC3FEDB933902025649EBBE236">
<mods:url id="F1B4F8045033C2CD0832E38D37D1BA7B">https://doi.org/10.11646/zootaxa.5570.2.8</mods:url>
</mods:location>
<mods:classification id="1F301CF2F7A5BC53DF87140A7C9270E5">journal article</mods:classification>
<mods:identifier id="AC3EAD4FA2A8BDC9BD9E19EF771F215B" type="DOI">10.11646/zootaxa.5570.2.8</mods:identifier>
<mods:identifier id="6EBF5BAC765B2238509CB8C279BFB1FE" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A1401D34BCFBA6760A7A9A553B5A3CF8" type="Zenodo-Dep">14735174</mods:identifier>
<mods:identifier id="D9F00DB48D9136FED08915A4B5ED5560" type="ZooBank">8030475D-8EA2-4C77-AD9E-72E3F8888D08</mods:identifier>
</mods:mods>
<treatment id="0398CE51FFACFF9FFF6BFCD6FD5365CB" ID-DOI="http://doi.org/10.5281/zenodo.14756621" ID-Zenodo-Dep="14756621" LSID="urn:lsid:plazi:treatment:0398CE51FFACFF9FFF6BFCD6FD5365CB" httpUri="http://treatment.plazi.org/id/0398CE51FFACFF9FFF6BFCD6FD5365CB" lastPageNumber="372" pageId="1" pageNumber="372">
<subSubSection id="C32B2CCCFFACFF9FFF6BFCD6FC8D616D" box="[151,773,834,861]" pageId="1" pageNumber="372" type="nomenclature">
<paragraph id="8B8E7F47FFACFF9FFF6BFCD6FC8D616D" blockId="1.[151,773,834,861]" box="[151,773,834,861]" pageId="1" pageNumber="372">
<heading id="D0C6C82BFFACFF9FFF6BFCD6FC8D616D" bold="true" box="[151,773,834,861]" fontSize="11" level="1" pageId="1" pageNumber="372" reason="1">
<emphasis id="B945A355FFACFF9FFF6BFCD6FC8D616D" bold="true" box="[151,773,834,861]" pageId="1" pageNumber="372">
Subgenus
<taxonomicName id="4C3104C4FFACFF9FFEEBFCD7FDB0616D" ID-CoL="8ZNDV" authority="Yu, 2023" authorityName="Yu" authorityYear="2023" box="[279,568,834,861]" kingdom="Animalia" pageId="1" pageNumber="372" rank="subGenus" subGenus="Ocreatomurus">
<emphasis id="B945A355FFACFF9FFEEBFCD7FE4D616C" bold="true" box="[279,453,835,861]" italics="true" pageId="1" pageNumber="372">Ocreatomurus</emphasis>
<bibRefCitation id="EFA002B6FFACFF9FFE30FCD7FDB0616D" author="Yu, D. &amp; Deharveng, L. &amp; Lukic, M. &amp; Wei, Y. &amp; Hu, F. &amp; Liu, M." box="[460,568,834,861]" pageId="1" pageNumber="372" refId="ref5424" refString="Yu, D., Deharveng, L., Lukic, M., Wei, Y., Hu, F. &amp; Liu, M. (2023) Corrigendum to &quot; Molecular phylogeny and trait evolution in an ancient terrestrial arthropod lineage: Systematic revision and implications for ecological divergence (Collembola, Tomocerinae) &quot; [Mol. Phylogenet. Evol. 154 (2021) 106995]. Molecular Phylogenetics &amp; Evolution, 189, 107926. https: // doi. org / 10.1016 / j. ympev. 2023.107926" type="journal volume" year="2023">
Yu,
<quantity id="4CC9D2A2FFACFF9FFE01FCD6FDB0616D" box="[509,568,834,860]" metricMagnitude="1" metricUnit="m" metricValue="5.13842" pageId="1" pageNumber="372" unit="in" value="2023.0">2023</quantity>
</bibRefCitation>
</taxonomicName>
in
<bibRefCitation id="EFA002B6FFACFF9FFDA2FCD7FC8D616D" author="Yu, D. &amp; Deharveng, L. &amp; Lukic, M. &amp; Wei, Y. &amp; Hu, F. &amp; Liu, M." box="[606,773,834,861]" pageId="1" pageNumber="372" refId="ref5424" refString="Yu, D., Deharveng, L., Lukic, M., Wei, Y., Hu, F. &amp; Liu, M. (2023) Corrigendum to &quot; Molecular phylogeny and trait evolution in an ancient terrestrial arthropod lineage: Systematic revision and implications for ecological divergence (Collembola, Tomocerinae) &quot; [Mol. Phylogenet. Evol. 154 (2021) 106995]. Molecular Phylogenetics &amp; Evolution, 189, 107926. https: // doi. org / 10.1016 / j. ympev. 2023.107926" type="journal volume" year="2023">
Yu
<emphasis id="B945A355FFACFF9FFD74FCD7FD4B616C" bold="true" box="[648,707,834,861]" italics="true" pageId="1" pageNumber="372">et al.</emphasis>
2023
</bibRefCitation>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32B2CCCFFACFF9FFF6BFC10FD5365CB" pageId="1" pageNumber="372" type="description">
<paragraph id="8B8E7F47FFACFF9FFF6BFC10FE3566AA" blockId="1.[151,1437,900,2043]" pageId="1" pageNumber="372">
<emphasis id="B945A355FFACFF9FFF6BFC10FD3661AF" bold="true" box="[151,702,900,926]" pageId="1" pageNumber="372">Common characters shared by the new species:</emphasis>
Eyes 6+6. Both dorsal and ventral sides of head scaled. Mouthparts normal for
<taxonomicName id="4C3104C4FFACFF9FFEEBFC3DFE1961F3" authorityName="Nicolet" authorityYear="1842" box="[279,401,937,962]" class="Collembola" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Entomobryomorpha" pageId="1" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B945A355FFACFF9FFEEBFC3DFE1961F3" box="[279,401,937,962]" italics="true" pageId="1" pageNumber="372">Tomocerus</emphasis>
</taxonomicName>
. Labral formula 4/5, 5, 4, distal edge of labrum with four papillae ending in curved spines. Mentum with five chaetae, submentum with numerous chaetae. Mandibular head asymmetrical, the left side with four teeth and the right side with five teeth, left molar plate distally with a tapered tooth. Maxillary lamella 5 without beard-like appendage. Maxillary outer lobe with trifurcate palp, one basal chaeta and four sublobal hairs. Both dorsal and ventral sides of Ant. I and Ant. II scaled, Ant. IV unscaled. Cephalic dorsal macrochaetotaxy: anterior area: 2 (A2), 4 (A3, A5); interocular area: 2 (M2), 7 (S0, S2, S5, S6); postocular area: 2+2 (Pa5, Pa6); posterior area: 3+3 (Pa2, Pp3, Pe5).
</paragraph>
<paragraph id="8B8E7F47FFACFF9FFF3BFB30FB16678B" blockId="1.[151,1437,900,2043]" pageId="1" pageNumber="372">Macrochaetae densely arranged along anterior margin of Th. II and sparsely on each tergum, most mesochaetae present laterally and posteriorly on terga. Number of macrochaetae or large mesochaetae in posterior row as 3 (p2, p3, p4), 3 (p1, p3, p5)/3 (m2, m3, m4), 3 (m2, m3, m4), 4 (p1, p3, p6, p7), 2 (p6, p7), 4 (m2, m3, m5, m6) on Th. IIIII/Abd. IV respectively. Th. II behind anterior marginal macrochaetae cluster with a file of macrochaetae, including a3, a4, and a4ae; central macrochaetae a2, a5, m1, m2 and m3 arranged approximately in triangle, m4 lateral to m2. Th. III with anterior macrochaeta a4; Abd. III with two anterior macrochaetae m3 and m6; Abd. IV with antero-lateral macrochaeta m6. Bothriotricha 2 (a6, m6), 1 (m6)/ 0, 0, 1 (a5), 2 (a2, a5), 0, 0 on Th. IIIII/ Abd. IVI. Tergal pseudopores 1, 1/1, 1, 1, 1, 0, 0 from Th. IIIII/Abd. IVI near body axis.</paragraph>
<paragraph id="8B8E7F47FFACFF9FFF3BFA50FD5365CB" blockId="1.[151,1437,900,2043]" pageId="1" pageNumber="372">
Trochantero-femoral organ with 1, 1 slender chaetae. Each tibiotarsus with distal whorl of 11 chaetae, among which the six ventral (inner) chaetae as ordinary chaetae, the five dorsal (outer) chaetae modified into a central clavate tenent hair, a pair of short accessory chaetae and a pair of slender guard chaetae. Unguis slender, with basointernal ridges about 1/41/3 distant from base; lateral teeth pointed, of moderate size. Inner edge of unguis with basal tooth and 46 more distal teeth, sub-basal tooth subequal to others or slightly larger. Unguiculus lanceolate. Rami of tenaculum with 4 + 4 teeth. Manubrium ventrally scaled without chaetae; laterally with large round scales and a row of strong chaetae; dorsally with two longitudinal strips of chaetae, including 2 + 2 pointed prominent chaetae at 1/32/5 and 3/44/5 the length of manubrium from base, respectively. Inner edge of each dorsal chaetal strip with a narrow strip of scales. Dens basally with a pointed prominent dorsal chaeta, without modified outer strong chaetae of
<taxonomicName id="4C3104C4FFACFF9FFE95F89CFE646513" authorityName="Yosii, R" authorityYear="1956" box="[361,492,1800,1826]" class="Collembola" family="Tomoceridae" genus="Plutomurus" kingdom="Animalia" order="Entomobryomorpha" pageId="1" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B945A355FFACFF9FFE95F89CFE646513" box="[361,492,1800,1826]" italics="true" pageId="1" pageNumber="372">Plutomurus</emphasis>
</taxonomicName>
-
<typeStatus id="548AC1E5FFACFF9FFE0FF89DFDAD6512" box="[499,549,1801,1827]" pageId="1" pageNumber="372">type</typeStatus>
or differentiated inner large scales of
<taxonomicName id="4C3104C4FFACFF9FFC16F89CFB3E6513" authorityName="Paclt" authorityYear="1944" box="[1002,1206,1800,1826]" class="Collembola" family="Tomoceridae" genus="Pogonognathellus" kingdom="Animalia" order="Entomobryomorpha" pageId="1" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B945A355FFACFF9FFC16F89CFB3E6513" box="[1002,1206,1800,1826]" italics="true" pageId="1" pageNumber="372">Pogonognathellus</emphasis>
</taxonomicName>
-
<typeStatus id="548AC1E5FFACFF9FFB41F89DFB7A6512" box="[1213,1266,1801,1827]" pageId="1" pageNumber="372">type</typeStatus>
. Dental spines compound, with numerous small denticles on surface. Distal end of dental spine array with a single large spine. Dens dorsally with ordinary chaetae and plumose chaetae, ventrally with dense scales and several apical chaetae. Mucro elongated, with numerous chaetae; apical tooth subequal to or slightly larger than subapical tooth; two dorsal lamellae running from proximal edge of subapical tooth, outer lamella ending in distal edge of inner basal tooth, inner lamella ending at the base of mucro, intermediate teeth located on outer lamella; both basal teeth with proximal lamellae, outer basal tooth with a toothlet.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,90 +1,92 @@
<document id="9642A59CAF72C0D2D14F4B543A06C024" ID-DOI="10.11646/zootaxa.5570.2.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="14735135" ID-ZooBank="8B1BAE9E-FBD3-4AC3-9FA3-9CCC2AA81392" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737737634945" checkinUser="plazi" docAuthor="Xin, Fei-Yi, Zhong, Xiao-Tian, Guo, Wen-Chi, Zhang, Yi-Feng, Qi, Zhi-Hao &amp; Wu, Song-Qing" docDate="2025" docId="03DC8794FFA2D678FF3CFAA0C25B2B0D" docLanguage="en" docName="zootaxa.5570.2.6.pdf" docOrigin="Zootaxa 5570 (2)" docSource="https://doi.org/10.11646/zootaxa.5570.2.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Neolucanus brochieri Bomans &amp; Miyashita 1997" docType="treatment" docVersion="2" lastPageNumber="357" masterDocId="FFE5FFECFFAED675FFABFFE1C3352F38" masterDocTitle="A taxonomic note on Chinese species of the Neolucanus fuscus group with the description of a new species (Coleoptera: Lucanidae: Lucaninae)" masterLastPageNumber="360" masterPageNumber="344" pageNumber="356" updateTime="1737738157217" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="94351F6C78DB6DAB515D5A6200D64A21" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7C8F66972E39D8A8E491092C9DD21502">
<mods:title id="E0726F887DDC5AB544052C049D004923">A taxonomic note on Chinese species of the Neolucanus fuscus group with the description of a new species (Coleoptera: Lucanidae: Lucaninae)</mods:title>
<document id="9401BDA6E18357DC9CA41E5D6ACBFBC1" ID-CLB-Dataset="307889" ID-DOI="10.11646/zootaxa.5570.2.6" ID-GBIF-Dataset="2396024e-a9ed-4584-afe7-9fed779d32e1" ID-ISSN="1175-5326" ID-Zenodo-Dep="14735135" ID-ZooBank="8B1BAE9E-FBD3-4AC3-9FA3-9CCC2AA81392" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1737737634945" checkinUser="plazi" docAuthor="Xin, Fei-Yi, Zhong, Xiao-Tian, Guo, Wen-Chi, Zhang, Yi-Feng, Qi, Zhi-Hao &amp; Wu, Song-Qing" docDate="2025" docId="03DC8794FFA2D678FF3CFAA0C25B2B0D" docLanguage="en" docName="zootaxa.5570.2.6.pdf" docOrigin="Zootaxa 5570 (2)" docSource="https://doi.org/10.11646/zootaxa.5570.2.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Neolucanus brochieri Bomans &amp; Miyashita 1997" docType="treatment" docVersion="5" lastPageNumber="357" masterDocId="FFE5FFECFFAED675FFABFFE1C3352F38" masterDocTitle="A taxonomic note on Chinese species of the Neolucanus fuscus group with the description of a new species (Coleoptera: Lucanidae: Lucaninae)" masterLastPageNumber="360" masterPageNumber="344" pageNumber="356" updateTime="1738085397281" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="EB939C451954975BF556D58D2AFB99B7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8392CD54A0CCC455C1F5D83535272B16">
<mods:title id="ACF37DD3F16D960BF6E0F70B4E729FD0">A taxonomic note on Chinese species of the Neolucanus fuscus group with the description of a new species (Coleoptera: Lucanidae: Lucaninae)</mods:title>
</mods:titleInfo>
<mods:name id="C5953952837E59071E0D0EAA4345AC4C" type="personal">
<mods:role id="0401A36C9E20CC690C690416C9EE2C0F">
<mods:roleTerm id="89DDD9A4592E8FF01F043DF3FF21E64E">Author</mods:roleTerm>
<mods:name id="091A8266E0D94EC14417D4AB03B60C13" type="personal">
<mods:role id="8216256892A8C7F3E19B665D4788E586">
<mods:roleTerm id="3E8FF3BE9080A2FA878FC9FEF0DF19A7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E68195A7F4A24FEC313549BF862D08D1">Xin, Fei-Yi</mods:namePart>
<mods:nameIdentifier id="5240730ACF0427E5FA25D4895247F4DD" type="ORCID">0009-0002-4407-694X</mods:nameIdentifier>
<mods:affiliation id="A8F11076E3C5FE3DE98D934BBCB0AA3E">Fujian Colleges and Universities Engineering Research Institute of Conservation and Utilization of Natural Bioresources, Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Key Laboratory of Integrated Pest Management in Ecological Forests, Fujian Province University, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China.</mods:affiliation>
<mods:namePart id="507B0F141ACB7DDAF0357872B0E2DEAA">Xin, Fei-Yi</mods:namePart>
<mods:nameIdentifier id="EDE6F77BEEB889381641BB0EA4AD316E" type="ORCID">0009-0002-4407-694X</mods:nameIdentifier>
<mods:affiliation id="DB5D5180F059C1AA7E38D1DEBDFAEC6C">Fujian Colleges and Universities Engineering Research Institute of Conservation and Utilization of Natural Bioresources, Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Key Laboratory of Integrated Pest Management in Ecological Forests, Fujian Province University, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China.</mods:affiliation>
</mods:name>
<mods:name id="37997731A3B3AEB200878232EEDC28EB" type="personal">
<mods:role id="B870DDC48D5F249DC2AA354ADD1306F8">
<mods:roleTerm id="293B5824CEEAB41F7524C34379FDE2DD">Author</mods:roleTerm>
<mods:name id="4FD9E934620E5BE23DC667B7DBFF44AB" type="personal">
<mods:role id="023115C69495A86A2043C87594014321">
<mods:roleTerm id="406AACA1456DD0EF3BD2CCE257588217">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C6CD99E572109DB4F597560B08978A50">Zhong, Xiao-Tian</mods:namePart>
<mods:nameIdentifier id="5395EAE891E3BAD82D4816304FB90950" type="ORCID">0009-0003-1423-9274</mods:nameIdentifier>
<mods:affiliation id="9C08E77FAF83D197601AC148787B853F">School of Geoscience and Technology, Southwest Petroleum University, Chengdu 610500, China.</mods:affiliation>
<mods:namePart id="E976E630335F564D86D29712DAE60F5F">Zhong, Xiao-Tian</mods:namePart>
<mods:nameIdentifier id="E232F1BACB5B23E2819309476254D58C" type="ORCID">0009-0003-1423-9274</mods:nameIdentifier>
<mods:affiliation id="26BD78F6320361D5D47D5896D779576B">School of Geoscience and Technology, Southwest Petroleum University, Chengdu 610500, China.</mods:affiliation>
</mods:name>
<mods:name id="AF5F20F44DFF6CBE1CC0A37358731921" type="personal">
<mods:role id="E21DBA809833D49FCDEBEDC7C8DC8B67">
<mods:roleTerm id="B717E631014B159FC166675C7D067963">Author</mods:roleTerm>
<mods:name id="E96CFF4C588EED54D71C3BE964C8DA76" type="personal">
<mods:role id="48FBDBE73DDE830450C7FF1A9FB955FB">
<mods:roleTerm id="07C788FFF6EB351367CE313E2E779E28">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FAFA67AF38E36865E46FC346220FCBD7">Guo, Wen-Chi</mods:namePart>
<mods:nameIdentifier id="752599B0BA978AAFFFADC823D065E52A" type="ORCID">0009-0008-4116-4016</mods:nameIdentifier>
<mods:affiliation id="8E08519DF871BC607D5F642740A972FC">Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China.</mods:affiliation>
<mods:nameIdentifier id="C5504F5B3506E10F8D64E41A40FFA10D" type="email">353600225@qq.com</mods:nameIdentifier>
<mods:namePart id="561E260E9BEFBBADA067BC583A464B4E">Guo, Wen-Chi</mods:namePart>
<mods:nameIdentifier id="320055D1C1E58BD500C8F414EE7BCEF4" type="ORCID">0009-0008-4116-4016</mods:nameIdentifier>
<mods:affiliation id="16E21445BA4A65CBEC1E9041B9F20196">Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China.</mods:affiliation>
<mods:nameIdentifier id="B83BBC066D4139840A5B9721074B3527" type="email">353600225@qq.com</mods:nameIdentifier>
</mods:name>
<mods:name id="97B9DC3A17950BE65F6032E4552B19F7" type="personal">
<mods:role id="A9FCE0D6D471D568C07F9B2FE779604E">
<mods:roleTerm id="895236B5465B3FA2006963E6D9B91E2F">Author</mods:roleTerm>
<mods:name id="49F635BF4470D4F69F07F2CFA67A81B8" type="personal">
<mods:role id="6A1F378D4746CD160583CC4E305F92CF">
<mods:roleTerm id="17518D863B8777EAA577CE054366B55E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3E5850FC07D82BAA23083FCA45D65FC0">Zhang, Yi-Feng</mods:namePart>
<mods:nameIdentifier id="5973E2D7719E33C62D5815EBC361E0B7" type="ORCID">0000-0003-3264-9472</mods:nameIdentifier>
<mods:affiliation id="CA6F9EFF50B95B5205EF6D6B1EC6D37D">Fujian Academy of Forestry, Fuzhou 350012, China.</mods:affiliation>
<mods:nameIdentifier id="8017826A756DC5AE693BA8D53B1258E1" type="email">zhang_yifeng@foxmail.com</mods:nameIdentifier>
<mods:namePart id="3FBD9FD257409AFE965F1EB396854758">Zhang, Yi-Feng</mods:namePart>
<mods:nameIdentifier id="5CE1CDEDD37FA71D55563D288602F35E" type="ORCID">0000-0003-3264-9472</mods:nameIdentifier>
<mods:affiliation id="4035C212797BF72E64B3E447EBD053FC">Fujian Academy of Forestry, Fuzhou 350012, China.</mods:affiliation>
<mods:nameIdentifier id="AD6D92F0160D8D736903C624CB86B373" type="email">zhang_yifeng@foxmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="7631DEFD3E700A157B7D8B9DB39A595C" type="personal">
<mods:role id="47A08134307583A66099DE00384C7A87">
<mods:roleTerm id="EB42754B02F5C2C338CBA2CAA3132EA4">Author</mods:roleTerm>
<mods:name id="9F1C6510999037B35B6E3C42982C9E09" type="personal">
<mods:role id="1AC540968C99D7F6B5B60232A060E651">
<mods:roleTerm id="49B4FB506C22CCDC102CAF397B9C8BF9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="26DD9577C2EEBF1ED6DA9C0186069FE5">Qi, Zhi-Hao</mods:namePart>
<mods:nameIdentifier id="EFD654C18153CC0119DDD56A5B7FA369" type="ORCID">0000-0001-8596-0913</mods:nameIdentifier>
<mods:affiliation id="677EBFB7833F4F0A0DFA26516CB46C19">Key Laboratory of Integrated Pest Management in Ecological Forests, Fujian Province University, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Fujian Academy of Forestry, Fuzhou 350012, China.</mods:affiliation>
<mods:namePart id="24306A36720619B8866EEAA3299BAF00">Qi, Zhi-Hao</mods:namePart>
<mods:nameIdentifier id="5CD488BE48FBFBFF5BC445EAF3185A61" type="ORCID">0000-0001-8596-0913</mods:nameIdentifier>
<mods:affiliation id="6223E7BF2D6110B3C45D3EEA01757A9B">Key Laboratory of Integrated Pest Management in Ecological Forests, Fujian Province University, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Fujian Academy of Forestry, Fuzhou 350012, China.</mods:affiliation>
</mods:name>
<mods:name id="A41D8B3DF85498FDCC29595AD589E264" type="personal">
<mods:role id="E8AD644AFBA0906FBC8213DB901DD820">
<mods:roleTerm id="40B15AD6BF479527C74B4A8A9E0D088C">Author</mods:roleTerm>
<mods:name id="3CA880C2068D09C86CDA2BB81109C5AD" type="personal">
<mods:role id="C838FDE8167438CA55D9B8D7980DB354">
<mods:roleTerm id="4724A1D8F7D5E123DD7031E63A78E5BA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="44DEF4C04BD27C345F74593A7E2290B2">Wu, Song-Qing</mods:namePart>
<mods:nameIdentifier id="6312805A084E58AEA80B5ABD42C566B3" type="ORCID">0000-0002-2763-5378</mods:nameIdentifier>
<mods:affiliation id="2C1833A3183803A739C97B0E1DB726A5">Fujian Colleges and Universities Engineering Research Institute of Conservation and Utilization of Natural Bioresources, Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Key Laboratory of Integrated Pest Management in Ecological Forests, Fujian Province University, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China.</mods:affiliation>
<mods:namePart id="966B3E5CC54059DD5CD0040469408BD8">Wu, Song-Qing</mods:namePart>
<mods:nameIdentifier id="5A4713906DA883B8947CC87A99D0583D" type="ORCID">0000-0002-2763-5378</mods:nameIdentifier>
<mods:affiliation id="529B04D96D54FDEC3DC6AD7598A29FFF">Fujian Colleges and Universities Engineering Research Institute of Conservation and Utilization of Natural Bioresources, Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Key Laboratory of Integrated Pest Management in Ecological Forests, Fujian Province University, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="EE06CDD6937427874C8AABD823CA7179">text</mods:typeOfResource>
<mods:relatedItem id="EA0061C5BC81432DB69D92A0530F39D7" type="host">
<mods:titleInfo id="8A4FFE949F6153144D61BA485238FFCB">
<mods:title id="9D017F92B15841A44292CF8F4929979C">Zootaxa</mods:title>
<mods:typeOfResource id="A71D4C045C5BD0F97C8941D9185216FE">text</mods:typeOfResource>
<mods:relatedItem id="634DB63C060CFC52984D8073C49F67D1" type="host">
<mods:titleInfo id="8A8381C306D970CA584F761DCF2396EB">
<mods:title id="24B6268C40B987129A0D059421C0B9E7">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="69B115142FC862918528E5D06CAD0A54">
<mods:date id="6D594EA60B58EB22724302C7FBB249DA">2025</mods:date>
<mods:detail id="49919AE7BED4B06E87A19A59088CEE5A" type="pubDate">
<mods:number id="C580A0B7EF7975BEE26E19AD3CED5D67">2025-01-17</mods:number>
<mods:part id="64B7AF34D014EEB33E601B1421050CFD">
<mods:date id="9109CAF08DA2673C554318F82F12DA13">2025</mods:date>
<mods:detail id="A89E6C6793E152B184C565F8EDB0F5ED" type="pubDate">
<mods:number id="D5049C9DB0F5555C03CECCBD94B1D167">2025-01-17</mods:number>
</mods:detail>
<mods:detail id="86E416765932C0BD5E2983439171825C" type="volume">
<mods:number id="7DE55A75B9C26798D7F78B894D9A6569">5570</mods:number>
<mods:detail id="9BE0245B2833C49225B2CBE28FCD3DC7" type="volume">
<mods:number id="69C6D1B8A8919B0C5FBD0CA6FD7C0C8C">5570</mods:number>
</mods:detail>
<mods:detail id="41ECB08B6BF32745774E5E70FB657BFB" type="issue">
<mods:number id="E50ACF51CFC8B5682E3BE81AC442B06A">2</mods:number>
<mods:detail id="80B86ABAE7BF06ECE631F5E1969FA447" type="issue">
<mods:number id="8CEAD12AE3BB6A8B7A60B7F4FCE57D68">2</mods:number>
</mods:detail>
<mods:extent id="7715CF71CFC6C4D22A01DCCE7B5D4550" unit="page">
<mods:start id="8D405915AD66A4256E8B228B30F08932">344</mods:start>
<mods:end id="A03B400840D6F900ADEC5EB4BD2CA116">360</mods:end>
<mods:extent id="2292770C66FED6A2C7D93DD56C413B0C" unit="page">
<mods:start id="2FDF066A0DB59974BADD267251E9BD24">344</mods:start>
<mods:end id="FE660A54B4957C038E8248F559981AA5">360</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="B76249B88D86DA2AF06DAA8453E77BEA">
<mods:url id="BB16CF5F0E0628A44CD976FBDCE0CB5A">https://doi.org/10.11646/zootaxa.5570.2.6</mods:url>
<mods:location id="B78E7A3CEC3AAB80ED494DBD182FEB79">
<mods:url id="7FDF442CC6281A756302950B98DAAB6C">https://doi.org/10.11646/zootaxa.5570.2.6</mods:url>
</mods:location>
<mods:classification id="2FA50256718C840C0A818F54478BEF15">journal article</mods:classification>
<mods:identifier id="B61A0B56C99B0B0B955B0454BE594873" type="DOI">10.11646/zootaxa.5570.2.6</mods:identifier>
<mods:identifier id="0ABAFA40F6526BE43C13A6F4FEEFA319" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="77FF4AFB02FF5B7A03E330B219577F4B" type="Zenodo-Dep">14735135</mods:identifier>
<mods:identifier id="ED519009E5CFC4908D5824BC154C3C34" type="ZooBank">8B1BAE9E-FBD3-4AC3-9FA3-9CCC2AA81392</mods:identifier>
<mods:classification id="2539556012DC5A618B46FBB6B8484859">journal article</mods:classification>
<mods:identifier id="0C402C50F2858BBEDBEC0D6767C41F37" type="CLB-Dataset">307889</mods:identifier>
<mods:identifier id="81A6B2376C4807B1A728DB2EA810C4EA" type="DOI">10.11646/zootaxa.5570.2.6</mods:identifier>
<mods:identifier id="D40BF5999A3AD3D9F556FC7D4A6FF8DC" type="GBIF-Dataset">2396024e-a9ed-4584-afe7-9fed779d32e1</mods:identifier>
<mods:identifier id="A38F625EDA69C5F4229337C34C3708D4" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A4753F785F937A61C09EEF46EFCCBE8A" type="Zenodo-Dep">14735135</mods:identifier>
<mods:identifier id="AA4D8D3B6E7BE5A919AEB5F48BF0A8E5" type="ZooBank">8B1BAE9E-FBD3-4AC3-9FA3-9CCC2AA81392</mods:identifier>
</mods:mods>
<treatment id="03DC8794FFA2D678FF3CFAA0C25B2B0D" LSID="urn:lsid:plazi:treatment:03DC8794FFA2D678FF3CFAA0C25B2B0D" httpUri="http://treatment.plazi.org/id/03DC8794FFA2D678FF3CFAA0C25B2B0D" lastPageId="13" lastPageNumber="357" pageId="12" pageNumber="356">
<treatment id="03DC8794FFA2D678FF3CFAA0C25B2B0D" ID-DOI="http://doi.org/10.5281/zenodo.14756557" ID-Zenodo-Dep="14756557" LSID="urn:lsid:plazi:treatment:03DC8794FFA2D678FF3CFAA0C25B2B0D" httpUri="http://treatment.plazi.org/id/03DC8794FFA2D678FF3CFAA0C25B2B0D" lastPageId="13" lastPageNumber="357" pageId="12" pageNumber="356" scope_family="Lucanidae" scope_order="Coleoptera" scope_subFamily="Lucaninae">
<subSubSection id="C36F6509FFA2D679FF3CFAA0C1FA2A45" pageId="12" pageNumber="356" type="nomenclature">
<paragraph id="8BCA3682FFA2D679FF3CFAA0C1CC2A63" blockId="12.[151,761,1345,1444]" box="[151,761,1345,1371]" pageId="12" pageNumber="356">
<heading id="D08281EEFFA2D679FF3CFAA0C1CC2A63" bold="true" box="[151,761,1345,1371]" fontSize="11" level="1" pageId="12" pageNumber="356" reason="1">
@ -106,7 +108,11 @@ Suggested Chinese common name:
<figureCitation id="134E2A07FFA2D679FF3CFA68C22D2A9B" box="[151,280,1417,1443]" captionStart="FIGURE 1" captionStartId="3.[151,250,1908,1932]" captionTargetBox="[312,1275,185,1877]" captionTargetId="figure-17@3.[306,1281,179,1883]" captionTargetPageId="3" captionText="FIGURE 1. Male habitus of Neolucanus species. AB, N. horizontalis Xin, Zhong &amp; Qi, new species, holotype, 36.0 mm, telodont (Lvchun, Yunnan); CE, N. fuscus Didier, 1926; CD, 50.9 mm, telodont (Yên Bái, Yên Bái); E, 44.8 mm, telodont (Ziyuan, Guangxi); FH, N. brochieri Bomans &amp; Miyashita, 1997; FG, 41.1mm, telodont (Gonshan, Yunnan); H, 41.0 mm, telodont (Gonshan, Yunnan). A, C, E, F, H, dorsal view; B, D, G, ventral view. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14735137" httpUri="https://zenodo.org/record/14735137/files/figure.png" pageId="12" pageNumber="356">Figs. 1FH</figureCitation>
;
<figureCitation id="134E2A07FFA2D679FE8FFA68C2502A9B" box="[292,357,1417,1443]" captionStart="FIGURE 2" captionStartId="4.[151,250,1771,1795]" captionTargetBox="[158,1429,186,1740]" captionTargetId="figure-18@4.[151,1436,181,1747]" captionTargetPageId="4" captionText="FIGURE 2. Male habitus of Neolucanus species. AB, N. horizontalis Xin, Zhong &amp; Qi, new species, paratype, 33.7 mm, mesodont (Jinping, Yunnan); CD, N. fuscus Didier, 1926, 34.7mm, priodont (Wuming, Guangxi); EF, N. brochieri Bomans &amp; Miyashita, 1997, 31.4 mm, priodont (Gonshan, Yunnan). A, C, E, dorsal view; B, D, F, ventral view. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14735139" httpUri="https://zenodo.org/record/14735139/files/figure.png" pageId="12" pageNumber="356">2EF</figureCitation>
; 3MP; 4IL; 5MP; 6D, H, L; 7D
;
<figureCitation id="134E2A07FFA2D679FEC4FA68C28D2A9B" box="[367,440,1417,1443]" captionStart="FIGURE 3" captionStartId="5.[151,250,1908,1932]" captionTargetBox="[225,1363,187,1879]" captionTargetId="figure-17@5.[219,1368,181,1885]" captionTargetPageId="5" captionText="FIGURE 3. Male fore-head (AB, EF, IJ, MN), pronotum (C, G, K, O), protibia and apical part of profemur (D, H, L, P) of Neolucanus species. AD, N. lehmanni Baba, 1995 (Imitation of Baba, 1995); EH, N. horizontalis Xin, Zhong &amp; Qi, new species, holotype (Lvchun, Yunnan); IL, N. fuscus Didier, 1926 (Ziyuan, Guangxi); MP, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, CD, E, GH, I, KL, M, OP, dorsal view; B, F, J, N lateral view." figureDoi="http://doi.org/10.5281/zenodo.14756551" httpUri="https://zenodo.org/record/14756551/files/figure.png" pageId="12" pageNumber="356">3MP</figureCitation>
; 4IL;
<figureCitation id="134E2A07FFA2D679FDA3FA68C1652A9C" box="[520,592,1417,1444]" captionStart="FIGURE 5" captionStartId="7.[151,250,1799,1823]" captionTargetBox="[157,1434,185,1769]" captionTargetId="figure-17@7.[151,1436,181,1775]" captionTargetPageId="7" captionText="FIGURE 5. Male abdominal tergite VIII (A, E, I, M), abdominal ventrite VIII (B, F, J, N) and abdominal segment IX (CD, G H, KL, OP) of Neolucanus species. AH, N. horizontalis Xin, Zhong &amp; Qi, new species AD, holotype (Lvchun, Yunnan); EH, paratype (Jinping, Yunnan). IL, N. fuscus Didier, 1926 (Ziyuan, Guangxi). MP, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, C, E, G, I, K, M, O, dorsal view; B, D, F, H, J, L, N, P, ventral view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14756558" httpUri="https://zenodo.org/record/14756558/files/figure.png" pageId="12" pageNumber="356">5MP</figureCitation>
; 6D, H, L; 7D
</paragraph>
</subSubSection>
<subSubSection id="C36F6509FFA2D679FF3CFA2FC1842ADE" box="[151,689,1486,1510]" pageId="12" pageNumber="356" type="reference_group">
@ -238,11 +244,17 @@ and present paper)
</paragraph>
<paragraph id="8BCA3682FFA2D678FF6CF841C21E2E79" blockId="12.[151,1437,1556,2014]" lastBlockId="13.[151,1437,151,1077]" lastPageId="13" lastPageNumber="357" pageId="12" pageNumber="356">
<emphasis id="B901EA90FFA2D679FF6CF841C2312882" box="[199,260,1952,1978]" italics="true" pageId="12" pageNumber="356">Head</emphasis>
transverse. Anterior ridge raised behind mandible and widely concave on the middle. Canthus long and wide, completely surround the eye. Clypeolabrum (Fig. 3M) tiny and transverse, slightly visible in dorsal view, fused with frons. Mandible (Fig. 3MN) smooth from base to teeth on inner margin; in lateral view, clearly up-curved and thinnest at basal 1/3, a dorsal tooth behind apex forming an approximately arc-shaped and weakly protrudes beyond upper margin; in dorsal view, slightly curved, thickness almost regular from base to inner teeth, with 23 continuous small teeth. Antennal club with 3 antennomeres; antennomeres 89 subquadrilateral; antennomere 10 semi-elliptic.
transverse. Anterior ridge raised behind mandible and widely concave on the middle. Canthus long and wide, completely surround the eye. Clypeolabrum (
<figureCitation id="134E2A07FFA2D679FD66F825C01228E6" box="[717,807,1988,2014]" captionStart="FIGURE 3" captionStartId="5.[151,250,1908,1932]" captionTargetBox="[225,1363,187,1879]" captionTargetId="figure-17@5.[219,1368,181,1885]" captionTargetPageId="5" captionText="FIGURE 3. Male fore-head (AB, EF, IJ, MN), pronotum (C, G, K, O), protibia and apical part of profemur (D, H, L, P) of Neolucanus species. AD, N. lehmanni Baba, 1995 (Imitation of Baba, 1995); EH, N. horizontalis Xin, Zhong &amp; Qi, new species, holotype (Lvchun, Yunnan); IL, N. fuscus Didier, 1926 (Ziyuan, Guangxi); MP, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, CD, E, GH, I, KL, M, OP, dorsal view; B, F, J, N lateral view." figureDoi="http://doi.org/10.5281/zenodo.14756551" httpUri="https://zenodo.org/record/14756551/files/figure.png" pageId="12" pageNumber="356">Fig. 3M</figureCitation>
) tiny and transverse, slightly visible in dorsal view, fused with frons. Mandible (
<figureCitation id="134E2A07FFA3D678FE33FF76C12D2F89" box="[408,536,151,177]" captionStart="FIGURE 3" captionStartId="5.[151,250,1908,1932]" captionTargetBox="[225,1363,187,1879]" captionTargetId="figure-17@5.[219,1368,181,1885]" captionTargetPageId="5" captionText="FIGURE 3. Male fore-head (AB, EF, IJ, MN), pronotum (C, G, K, O), protibia and apical part of profemur (D, H, L, P) of Neolucanus species. AD, N. lehmanni Baba, 1995 (Imitation of Baba, 1995); EH, N. horizontalis Xin, Zhong &amp; Qi, new species, holotype (Lvchun, Yunnan); IL, N. fuscus Didier, 1926 (Ziyuan, Guangxi); MP, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, CD, E, GH, I, KL, M, OP, dorsal view; B, F, J, N lateral view." figureDoi="http://doi.org/10.5281/zenodo.14756551" httpUri="https://zenodo.org/record/14756551/files/figure.png" pageId="13" pageNumber="357">Fig. 3MN</figureCitation>
) smooth from base to teeth on inner margin; in lateral view, clearly up-curved and thinnest at basal 1/3, a dorsal tooth behind apex forming an approximately arc-shaped and weakly protrudes beyond upper margin; in dorsal view, slightly curved, thickness almost regular from base to inner teeth, with 23 continuous small teeth. Antennal club with 3 antennomeres; antennomeres 89 subquadrilateral; antennomere 10 semi-elliptic.
</paragraph>
<paragraph id="8BCA3682FFA3D678FF6CFEAAC2032E95" blockId="13.[151,1437,151,1077]" pageId="13" pageNumber="357">
<emphasis id="B901EA90FFA3D678FF6CFEAAC2032E5C" box="[199,310,331,356]" italics="true" pageId="13" pageNumber="357">Pronotum</emphasis>
(Fig. 3O) transverse, markedly longer and wider than head with slightly raised edges.Anterior margin bisinuate, posterior margin dimly bisinuate. Anterior angles blunt and directed forwards, lateral angles and posterior angles obtuse.
(
<figureCitation id="134E2A07FFA3D678FEEFFEAAC2AF2E5D" box="[324,410,331,357]" captionStart="FIGURE 3" captionStartId="5.[151,250,1908,1932]" captionTargetBox="[225,1363,187,1879]" captionTargetId="figure-17@5.[219,1368,181,1885]" captionTargetPageId="5" captionText="FIGURE 3. Male fore-head (AB, EF, IJ, MN), pronotum (C, G, K, O), protibia and apical part of profemur (D, H, L, P) of Neolucanus species. AD, N. lehmanni Baba, 1995 (Imitation of Baba, 1995); EH, N. horizontalis Xin, Zhong &amp; Qi, new species, holotype (Lvchun, Yunnan); IL, N. fuscus Didier, 1926 (Ziyuan, Guangxi); MP, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, CD, E, GH, I, KL, M, OP, dorsal view; B, F, J, N lateral view." figureDoi="http://doi.org/10.5281/zenodo.14756551" httpUri="https://zenodo.org/record/14756551/files/figure.png" pageId="13" pageNumber="357">Fig. 3O</figureCitation>
) transverse, markedly longer and wider than head with slightly raised edges.Anterior margin bisinuate, posterior margin dimly bisinuate. Anterior angles blunt and directed forwards, lateral angles and posterior angles obtuse.
</paragraph>
<paragraph id="8BCA3682FFA3D678FF6CFE56C7302EE9" blockId="13.[151,1437,151,1077]" box="[199,1029,439,465]" pageId="13" pageNumber="357">
<emphasis id="B901EA90FFA3D678FF6CFE56C2382EE9" box="[199,269,439,465]" italics="true" pageId="13" pageNumber="357">Elytra</emphasis>
@ -250,11 +262,19 @@ elongate, anterior margin of elytra clearly narrower than pronotum.
</paragraph>
<paragraph id="8BCA3682FFA3D678FF6CFE3AC0072D05" blockId="13.[151,1437,151,1077]" pageId="13" pageNumber="357">
<emphasis id="B901EA90FFA3D678FF6CFE3AC3C92ECC" box="[199,252,475,500]" italics="true" pageId="13" pageNumber="357">Legs</emphasis>
. Protibia (Fig. 3P) with 46 conspicuous sharp and short protuberances along outer margin, apex bifurcate with branches moderately sharp at tip; inner margin with a unclear gap at base. Mesotibia and metatibia straight and outer margin with a terminal spine, without external spines.
. Protibia (
<figureCitation id="134E2A07FFA3D678FEDBFE3AC2F42ECD" box="[368,449,475,501]" captionStart="FIGURE 3" captionStartId="5.[151,250,1908,1932]" captionTargetBox="[225,1363,187,1879]" captionTargetId="figure-17@5.[219,1368,181,1885]" captionTargetPageId="5" captionText="FIGURE 3. Male fore-head (AB, EF, IJ, MN), pronotum (C, G, K, O), protibia and apical part of profemur (D, H, L, P) of Neolucanus species. AD, N. lehmanni Baba, 1995 (Imitation of Baba, 1995); EH, N. horizontalis Xin, Zhong &amp; Qi, new species, holotype (Lvchun, Yunnan); IL, N. fuscus Didier, 1926 (Ziyuan, Guangxi); MP, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, CD, E, GH, I, KL, M, OP, dorsal view; B, F, J, N lateral view." figureDoi="http://doi.org/10.5281/zenodo.14756551" httpUri="https://zenodo.org/record/14756551/files/figure.png" pageId="13" pageNumber="357">Fig. 3P</figureCitation>
) with 46 conspicuous sharp and short protuberances along outer margin, apex bifurcate with branches moderately sharp at tip; inner margin with a unclear gap at base. Mesotibia and metatibia straight and outer margin with a terminal spine, without external spines.
</paragraph>
<paragraph id="8BCA3682FFA3D678FF6CFDA6C15B2C01" blockId="13.[151,1437,151,1077]" pageId="13" pageNumber="357">
<emphasis id="B901EA90FFA3D678FF6CFDA6C2D62D59" box="[199,483,583,609]" italics="true" pageId="13" pageNumber="357">Abdomen and aedeagus.</emphasis>
Abdominal tergite VIII (Fig. 5M) subsemi-circular, lateral angles poorly defined. Abdominal ventrite VIII (Fig. 5N) with smooth basal margin. Ventral plate of abdominal segment IX (Fig. 5P) slender without membranous stripe, obviously constricted at base, then widens sharply and significantly, widens gradually to about apical 1/5. Aedeagus (
Abdominal tergite VIII (
<figureCitation id="134E2A07FFA3D678FCA1FDA6C05F2D59" box="[778,874,582,609]" captionStart="FIGURE 5" captionStartId="7.[151,250,1799,1823]" captionTargetBox="[157,1434,185,1769]" captionTargetId="figure-17@7.[151,1436,181,1775]" captionTargetPageId="7" captionText="FIGURE 5. Male abdominal tergite VIII (A, E, I, M), abdominal ventrite VIII (B, F, J, N) and abdominal segment IX (CD, G H, KL, OP) of Neolucanus species. AH, N. horizontalis Xin, Zhong &amp; Qi, new species AD, holotype (Lvchun, Yunnan); EH, paratype (Jinping, Yunnan). IL, N. fuscus Didier, 1926 (Ziyuan, Guangxi). MP, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, C, E, G, I, K, M, O, dorsal view; B, D, F, H, J, L, N, P, ventral view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14756558" httpUri="https://zenodo.org/record/14756558/files/figure.png" pageId="13" pageNumber="357">Fig. 5M</figureCitation>
) subsemi-circular, lateral angles poorly defined. Abdominal ventrite VIII (
<figureCitation id="134E2A07FFA3D678FE6AFD8AC12E2DBD" box="[449,539,618,645]" captionStart="FIGURE 5" captionStartId="7.[151,250,1799,1823]" captionTargetBox="[157,1434,185,1769]" captionTargetId="figure-17@7.[151,1436,181,1775]" captionTargetPageId="7" captionText="FIGURE 5. Male abdominal tergite VIII (A, E, I, M), abdominal ventrite VIII (B, F, J, N) and abdominal segment IX (CD, G H, KL, OP) of Neolucanus species. AH, N. horizontalis Xin, Zhong &amp; Qi, new species AD, holotype (Lvchun, Yunnan); EH, paratype (Jinping, Yunnan). IL, N. fuscus Didier, 1926 (Ziyuan, Guangxi). MP, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, C, E, G, I, K, M, O, dorsal view; B, D, F, H, J, L, N, P, ventral view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14756558" httpUri="https://zenodo.org/record/14756558/files/figure.png" pageId="13" pageNumber="357">Fig. 5N</figureCitation>
) with smooth basal margin. Ventral plate of abdominal segment IX (
<figureCitation id="134E2A07FFA3D678FA95FD8AC6A12DBD" box="[1342,1428,618,645]" captionStart="FIGURE 5" captionStartId="7.[151,250,1799,1823]" captionTargetBox="[157,1434,185,1769]" captionTargetId="figure-17@7.[151,1436,181,1775]" captionTargetPageId="7" captionText="FIGURE 5. Male abdominal tergite VIII (A, E, I, M), abdominal ventrite VIII (B, F, J, N) and abdominal segment IX (CD, G H, KL, OP) of Neolucanus species. AH, N. horizontalis Xin, Zhong &amp; Qi, new species AD, holotype (Lvchun, Yunnan); EH, paratype (Jinping, Yunnan). IL, N. fuscus Didier, 1926 (Ziyuan, Guangxi). MP, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, C, E, G, I, K, M, O, dorsal view; B, D, F, H, J, L, N, P, ventral view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14756558" httpUri="https://zenodo.org/record/14756558/files/figure.png" pageId="13" pageNumber="357">Fig. 5P</figureCitation>
) slender without membranous stripe, obviously constricted at base, then widens sharply and significantly, widens gradually to about apical 1/5. Aedeagus (
<figureCitation id="134E2A07FFA3D678FDCFFD52C1CB2DF5" box="[612,766,691,717]" captionStart="FIGURE 6" captionStartId="8.[151,250,1909,1933]" captionTargetBox="[180,1408,181,1879]" captionTargetId="figure-18@8.[174,1414,181,1885]" captionTargetPageId="8" captionText="FIGURE 6. Aedeagus (not including flagellum) of Neolucanus species. AB, EF, IJ, N. horizontalis Xin, Zhong &amp; Qi, new species A, E, I, holotype (Lvchun, Yunnan); B, F, J, paratype (Jinping, Yunnan). C, G, K, N. fuscus Didier, 1926 (Ziyuan, Guangxi). D, H, L, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). AD, dorsal view; EH, ventral view; IL, lateral view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14735143" httpUri="https://zenodo.org/record/14735143/files/figure.png" pageId="13" pageNumber="357">Fig. 6D, H, L</figureCitation>
) with a significant and relatively slender pair of sclerotized dorsal plates; ventral plate at apical end of basal piece membranous. Paramere relatively narrow, with a obvious curvature at base, apex of paramere curved outward. Penis large, almost as long as paramere in ventral view. Flagellum long, apex of flagellum simple.
</paragraph>
@ -277,9 +297,9 @@ and
<paragraph id="8BCA3682FFA3D678FF6CFC33C1972CD5" blockId="13.[151,1437,151,1077]" box="[199,674,978,1005]" pageId="13" pageNumber="357">
<emphasis id="B901EA90FFA3D678FF6CFC33C2FB2CD4" bold="true" box="[199,462,978,1004]" pageId="13" pageNumber="357">Differential diagnosis.</emphasis>
See
<tableCitation id="C6F70339FFA3D678FDA8FC32C1632CD5" box="[515,598,979,1005]" captionStart="TABLE 1" captionStartId="14.[151,239,728,752]" captionText="TABLE 1. Male main differences of external characters of Neolucanus horizontalis Xin, Zhong &amp; Qi, new species and its related species." pageId="13" pageNumber="357">Table 1</tableCitation>
<tableCitation id="C6F70339FFA3D678FDA8FC32C1632CD5" box="[515,598,979,1005]" captionStart="TABLE 1" captionStartId="14.[151,239,728,752]" captionTargetPageId="14" captionText="TABLE 1. Male main differences of external characters of Neolucanus horizontalis Xin, Zhong &amp; Qi, new species and its related species." httpUri="http://table.plazi.org/id/DF0A660AFFA0D67BFF3CFD39C26E2C2C" pageId="13" pageNumber="357" tableUuid="DF0A660AFFA0D67BFF3CFD39C26E2C2C">Table 1</tableCitation>
and
<tableCitation id="C6F70339FFA3D678FD26FC32C1AB2CD5" box="[653,670,979,1005]" captionStart="TABLE 2" captionStartId="15.[151,239,151,175]" captionText="TABLE 2. Main differences of male abdomen and aedeagus of Neolucanus horizontalis Xin, Zhong &amp; Qi, new species and its related species." pageId="13" pageNumber="357">2</tableCitation>
<tableCitation id="C6F70339FFA3D678FD26FC32C1AB2CD5" box="[653,670,979,1005]" captionStart="TABLE 2" captionStartId="15.[151,239,151,175]" captionTargetPageId="15" captionText="TABLE 2. Main differences of male abdomen and aedeagus of Neolucanus horizontalis Xin, Zhong &amp; Qi, new species and its related species." httpUri="http://table.plazi.org/id/DF0A660AFFA1D67AFF3CFF76C2BD2FEB" pageId="13" pageNumber="357" tableUuid="DF0A660AFFA1D67AFF3CFF76C2BD2FEB">2</tableCitation>
.
</paragraph>
</subSubSection>

View file

@ -0,0 +1,214 @@
<document id="988325A8F6A7A6DF01465A9E12409228" ID-CLB-Dataset="307889" ID-DOI="10.11646/zootaxa.5570.2.6" ID-GBIF-Dataset="2396024e-a9ed-4584-afe7-9fed779d32e1" ID-ISSN="1175-5326" ID-Zenodo-Dep="14735135" ID-ZooBank="8B1BAE9E-FBD3-4AC3-9FA3-9CCC2AA81392" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1737737634945" checkinUser="plazi" docAuthor="Xin, Fei-Yi, Zhong, Xiao-Tian, Guo, Wen-Chi, Zhang, Yi-Feng, Qi, Zhi-Hao &amp; Wu, Song-Qing" docDate="2025" docId="03DC8794FFA3D678FF3CFB66C6A9297F" docLanguage="en" docName="zootaxa.5570.2.6.pdf" docOrigin="Zootaxa 5570 (2)" docSource="https://doi.org/10.11646/zootaxa.5570.2.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Neolucanus fuscus Didier 1926" docType="key" docVersion="5" lastPageNumber="357" masterDocId="FFE5FFECFFAED675FFABFFE1C3352F38" masterDocTitle="A taxonomic note on Chinese species of the Neolucanus fuscus group with the description of a new species (Coleoptera: Lucanidae: Lucaninae)" masterLastPageNumber="360" masterPageNumber="344" pageNumber="357" updateTime="1738085397281" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="EE0A4F2C8595FDAEF397825C175B1796" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2CF76FD6DC1126F54346F320475AB6FB">
<mods:title id="74CBB3BBD1CC390168A685A62AF1B17E">A taxonomic note on Chinese species of the Neolucanus fuscus group with the description of a new species (Coleoptera: Lucanidae: Lucaninae)</mods:title>
</mods:titleInfo>
<mods:name id="07E5422662D76B165EE5A65A707D231E" type="personal">
<mods:role id="5B4363B87CECA074FEF3D8D8CC5A32AD">
<mods:roleTerm id="C9432A99892A9084D6226DC306D3E363">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C3F6B2FEDE15476A4C3A79B3FFA87CEE">Xin, Fei-Yi</mods:namePart>
<mods:nameIdentifier id="A3CAED6120102780F0EC6B3CAD688019" type="ORCID">0009-0002-4407-694X</mods:nameIdentifier>
<mods:affiliation id="37B7E712789F695C14DB9569E52C0766">Fujian Colleges and Universities Engineering Research Institute of Conservation and Utilization of Natural Bioresources, Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Key Laboratory of Integrated Pest Management in Ecological Forests, Fujian Province University, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China.</mods:affiliation>
</mods:name>
<mods:name id="0C20B1177722557A66AAA32805674424" type="personal">
<mods:role id="F61A58765FE517AB28C32F7274DE81FB">
<mods:roleTerm id="D0D0DE63A8A619CDB8C6AD3D44F5BBE7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8329C1EA63F6488C8DD9618A2CEC570D">Zhong, Xiao-Tian</mods:namePart>
<mods:nameIdentifier id="53FA781567148214873D70FF81B40909" type="ORCID">0009-0003-1423-9274</mods:nameIdentifier>
<mods:affiliation id="4B58229F0827B40660F0E9F6C473487B">School of Geoscience and Technology, Southwest Petroleum University, Chengdu 610500, China.</mods:affiliation>
</mods:name>
<mods:name id="DFD6F0B66CF7DB12683342F7DFEBE807" type="personal">
<mods:role id="76C9D86384872C03163FFCEB3351A506">
<mods:roleTerm id="4EE8C28BF0362B2D0B8AF79A174D849E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="44C6D6BA533F5C7602568E31A026D301">Guo, Wen-Chi</mods:namePart>
<mods:nameIdentifier id="810E4F0CB9C6837EB37CC0E6FE697A06" type="ORCID">0009-0008-4116-4016</mods:nameIdentifier>
<mods:affiliation id="88891FF60ACD2094549E87FD05722D5C">Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China.</mods:affiliation>
<mods:nameIdentifier id="563DCACEC2B6F4784B3EEBB50DBC9C08" type="email">353600225@qq.com</mods:nameIdentifier>
</mods:name>
<mods:name id="2043E537862BDA24EC5DA3F79ED86FFA" type="personal">
<mods:role id="28123AE677A53319A4A2D1BB0B6765DB">
<mods:roleTerm id="7F6AF2649BD277DEB136DB7C6F883063">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FB1CC8B7039E3BBDA5BD954450882BE2">Zhang, Yi-Feng</mods:namePart>
<mods:nameIdentifier id="A5494B1DE4A717E45836A40E72CA1FC5" type="ORCID">0000-0003-3264-9472</mods:nameIdentifier>
<mods:affiliation id="E0F1F14043517F786F957B9D59396D3A">Fujian Academy of Forestry, Fuzhou 350012, China.</mods:affiliation>
<mods:nameIdentifier id="18D41D9853CB7347BE636CFB47B0134E" type="email">zhang_yifeng@foxmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="440554D141B76910C6CDFB389033E217" type="personal">
<mods:role id="C2C07078A04C7FAC41CA133668E50FD0">
<mods:roleTerm id="16442C3E5F16467E475ED27C871123F8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BEE23096C1BEF3F6C218406941ADD5C9">Qi, Zhi-Hao</mods:namePart>
<mods:nameIdentifier id="1B70A881836BE40F2C4ABC4E0C3D0CB6" type="ORCID">0000-0001-8596-0913</mods:nameIdentifier>
<mods:affiliation id="7DCE83C000B49D1231AFEA32B0F8AAAF">Key Laboratory of Integrated Pest Management in Ecological Forests, Fujian Province University, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Fujian Academy of Forestry, Fuzhou 350012, China.</mods:affiliation>
</mods:name>
<mods:name id="C841C1D1FD336610B45A9752CF7E295B" type="personal">
<mods:role id="CFBA85655C22AFCCE04866A0CBA95C98">
<mods:roleTerm id="6F543B2FEDA42E156CDC14D48628DDD5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B1AAD243449D84C2E815CAD4EEF8BD36">Wu, Song-Qing</mods:namePart>
<mods:nameIdentifier id="B4CBDF646339D8D1EB664474BFF3167B" type="ORCID">0000-0002-2763-5378</mods:nameIdentifier>
<mods:affiliation id="F5DA87F6EE836B61666188EB73FA1127">Fujian Colleges and Universities Engineering Research Institute of Conservation and Utilization of Natural Bioresources, Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Key Laboratory of Integrated Pest Management in Ecological Forests, Fujian Province University, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="81C23C7383B2A17DBFA29178DD77A465">text</mods:typeOfResource>
<mods:relatedItem id="20709D90BC0780FC68BF8205954A3096" type="host">
<mods:titleInfo id="F0826284666DB7DEA61306DAD946A19A">
<mods:title id="116EBA108021B8F8B7B5019457DF5630">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="EA736B3B802955D444EBEF0CD4B27A0E">
<mods:date id="54EEEDE19B5C8308E7EE56D187FBD974">2025</mods:date>
<mods:detail id="70DBCE8CC12CA184FBD824669931767A" type="pubDate">
<mods:number id="329DF43A3394BC805AF2016C1B47514C">2025-01-17</mods:number>
</mods:detail>
<mods:detail id="C959C9F9C6C9A9168FE2A7567F67E606" type="volume">
<mods:number id="D57AE184078DD9B9185AA66F890BDC05">5570</mods:number>
</mods:detail>
<mods:detail id="072C5F4FC14E0500BC424F346CD643C0" type="issue">
<mods:number id="24E063A59F70748BA8934253E9918991">2</mods:number>
</mods:detail>
<mods:extent id="AA34AFCBEF58EDECF145AAD7D92DF4F2" unit="page">
<mods:start id="27C5F948DB33ACDD5A1C4D4CEDA82525">344</mods:start>
<mods:end id="4CC2B8C0CF83D332CA2B65DC0C407D44">360</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="0B52C5151FE008469A50C6C5FF4DDB29">
<mods:url id="9922B6A5698ABEDF0314C79605A445B8">https://doi.org/10.11646/zootaxa.5570.2.6</mods:url>
</mods:location>
<mods:classification id="25A9E1E1A4B9BDC04AA85D151C757876">journal article</mods:classification>
<mods:identifier id="FFF1B39F4FBB145BF7E620C4BEE75373" type="CLB-Dataset">307889</mods:identifier>
<mods:identifier id="8E025CCDA1509A753B9126633CFE3FF1" type="DOI">10.11646/zootaxa.5570.2.6</mods:identifier>
<mods:identifier id="E832067B39E17697F5433CE9B97A8B5F" type="GBIF-Dataset">2396024e-a9ed-4584-afe7-9fed779d32e1</mods:identifier>
<mods:identifier id="22FD3A595D74EC9D35B99FC254C79375" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="5227C71E5A83E2D8E4752C6509305FA2" type="Zenodo-Dep">14735135</mods:identifier>
<mods:identifier id="18442B75640C216E3AE3E77E42DFED0F" type="ZooBank">8B1BAE9E-FBD3-4AC3-9FA3-9CCC2AA81392</mods:identifier>
</mods:mods>
<treatment id="03DC8794FFA3D678FF3CFB66C6A9297F" ID-DOI="http://doi.org/10.5281/zenodo.14756560" ID-Zenodo-Dep="14756560" LSID="urn:lsid:plazi:treatment:03DC8794FFA3D678FF3CFB66C6A9297F" httpUri="http://treatment.plazi.org/id/03DC8794FFA3D678FF3CFB66C6A9297F" lastPageNumber="357" pageId="13" pageNumber="357" scope_family="Lucanidae" scope_order="Coleoptera" scope_subFamily="Lucaninae">
<subSubSection id="C36F6509FFA3D678FF3CFB66C2F22BFC" pageId="13" pageNumber="357" type="nomenclature">
<paragraph id="8BCA3682FFA3D678FF3CFB66C2F22BFC" blockId="13.[151,1436,1158,1221]" pageId="13" pageNumber="357">
<heading id="D08281EEFFA3D678FF3CFB66C2F22BFC" bold="true" centered="true" fontSize="11" level="1" pageId="13" pageNumber="357" reason="1">
<emphasis id="B901EA90FFA3D678FF3CFB66C2F22BFC" bold="true" pageId="13" pageNumber="357">
Key to identification of telodont males of the
<taxonomicName id="4C754D01FFA3D678FD61FB67C0852B98" ID-CoL="46LRJ" authorityName="Didier" authorityYear="1926" box="[714,944,1158,1185]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="357" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="B901EA90FFA3D678FD61FB67C0852B98" bold="true" box="[714,944,1158,1185]" italics="true" pageId="13" pageNumber="357">Neolucanus fuscus</emphasis>
</taxonomicName>
group (
<bibRefCitation id="EFE44B73FFA3D678FBB9FB67C7922B98" author="Fujita, H." box="[1042,1191,1158,1184]" pageId="13" pageNumber="357" refId="ref7425" refString="Fujita, H. (2010) The Lucanid Beetles of the World. Mushi-Sha, Tokyo, 472 pp., 248 pls." type="book" year="2010">Fujita 2010</bibRefCitation>
;
<bibRefCitation id="EFE44B73FFA3D678FB18FB67C6462B98" author="Toshiyuki, T." box="[1203,1395,1158,1184]" pageId="13" pageNumber="357" pagination="1 - 87" refId="ref7955" refString="Toshiyuki, T. (2021) Illustrated guide to the genus Neolucanus of the world. Be-kuwa, 78, 1 - 87." type="journal article" year="2021">Toshiyuki 2021</bibRefCitation>
;
<bibRefCitation id="EFE44B73FFA3D678FA2BFB67C3E72BFC" author="Yi, D." pageId="13" pageNumber="357" refId="ref7980" refString="Yi, D. (2023) Lucanidae of the World. Design Spacetime, Seoul, 190 pp., 160 pls." type="book" year="2023">Yi 2023</bibRefCitation>
and present paper)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36F6509FFA3D678FF3CFB11C6A9297F" pageId="13" pageNumber="357" type="key">
<key id="FFE461B3FFA3D678FF3CFB11C6A9297F" pageId="13" pageNumber="357">
<keyStep id="30812E27FFA3D678FF3CFB11C6A92A07" pageId="13" pageNumber="357">
<paragraph id="8BCA3682FFA3D678FF3CFB11C6A82A1B" blockId="13.[151,1437,1264,1608]" pageId="13" pageNumber="357">
<keyLead id="308495B7FFA3D678FF3CFB11C6A82A1B" pageId="13" pageNumber="357">
1 Mandible almost horizontal with approximate right-angle fork behind the apex in lateral view (
<figureCitation id="134E2A07FFA3D678FBD9FB11C78D2A3D" box="[1138,1208,1264,1286]" captionStart="FIGURE 3" captionStartId="5.[151,250,1908,1932]" captionTargetBox="[225,1363,187,1879]" captionTargetId="figure-17@5.[219,1368,181,1885]" captionTargetPageId="5" captionText="FIGURE 3. Male fore-head (AB, EF, IJ, MN), pronotum (C, G, K, O), protibia and apical part of profemur (D, H, L, P) of Neolucanus species. AD, N. lehmanni Baba, 1995 (Imitation of Baba, 1995); EH, N. horizontalis Xin, Zhong &amp; Qi, new species, holotype (Lvchun, Yunnan); IL, N. fuscus Didier, 1926 (Ziyuan, Guangxi); MP, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, CD, E, GH, I, KL, M, OP, dorsal view; B, F, J, N lateral view." figureDoi="http://doi.org/10.5281/zenodo.14756551" httpUri="https://zenodo.org/record/14756551/files/figure.png" pageId="13" pageNumber="357">Fig. 3F</figureCitation>
).........................................................................
<taxonomicName id="4C754D01FFA3D678FC2EFAECC6292A1B" authority="Xin, Zhong &amp; Qi" authorityName="Xin, Zhong &amp; Qi" authorityYear="2025" box="[901,1308,1293,1315]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="357" phylum="Arthropoda" rank="species" species="horizontalis" status="sp. nov.">
<emphasis id="B901EA90FFA3D678FC2EFAECC75A2A1A" box="[901,1135,1293,1314]" italics="true" pageId="13" pageNumber="357">Neolucanus horizontalis</emphasis>
Xin, Zhong &amp; Qi
</taxonomicName>
,
<emphasis id="B901EA90FFA3D678FA8DFAECC6A82A1B" bold="true" box="[1318,1437,1293,1315]" pageId="13" pageNumber="357">
<taxonomicNameLabel id="A23257EBFFA3D678FA8DFAECC6A82A1B" box="[1318,1437,1293,1315]" pageId="13" pageNumber="357" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BCA3682FFA3D678FF3CFACAC6A92A07" blockId="13.[151,1437,1264,1608]" box="[151,1436,1322,1344]" pageId="13" pageNumber="357">
<keyLead id="308495B7FFA3D678FF3CFACAC6A92A07" box="[151,1436,1322,1344]" pageId="13" pageNumber="357">
Mandible up-curved with acute-angle fork behind the apex in lateral view (
<figureCitation id="134E2A07FFA3D678FC18FACBC70E2A07" box="[947,1083,1322,1344]" captionStart="FIGURE 3" captionStartId="5.[151,250,1908,1932]" captionTargetBox="[225,1363,187,1879]" captionTargetId="figure-17@5.[219,1368,181,1885]" captionTargetPageId="5" captionText="FIGURE 3. Male fore-head (AB, EF, IJ, MN), pronotum (C, G, K, O), protibia and apical part of profemur (D, H, L, P) of Neolucanus species. AD, N. lehmanni Baba, 1995 (Imitation of Baba, 1995); EH, N. horizontalis Xin, Zhong &amp; Qi, new species, holotype (Lvchun, Yunnan); IL, N. fuscus Didier, 1926 (Ziyuan, Guangxi); MP, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, CD, E, GH, I, KL, M, OP, dorsal view; B, F, J, N lateral view." figureDoi="http://doi.org/10.5281/zenodo.14756551" httpUri="https://zenodo.org/record/14756551/files/figure.png" pageId="13" pageNumber="357">Figs. 3B, J, N</figureCitation>
)............................ 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30812E27FFA3D678FF3CFAA9C6A92A42" pageId="13" pageNumber="357">
<paragraph id="8BCA3682FFA3D678FF3CFAA9C6A92A65" blockId="13.[151,1437,1264,1608]" box="[151,1436,1352,1374]" pageId="13" pageNumber="357">
<keyLead id="308495B7FFA3D678FF3CFAA9C6A92A65" box="[151,1436,1352,1374]" pageId="13" pageNumber="357">
2 Base of inner margin of protibia with a noticeable gap (
<figureCitation id="134E2A07FFA3D678FD5DFAA9C0752A65" box="[758,832,1352,1374]" captionStart="FIGURE 3" captionStartId="5.[151,250,1908,1932]" captionTargetBox="[225,1363,187,1879]" captionTargetId="figure-17@5.[219,1368,181,1885]" captionTargetPageId="5" captionText="FIGURE 3. Male fore-head (AB, EF, IJ, MN), pronotum (C, G, K, O), protibia and apical part of profemur (D, H, L, P) of Neolucanus species. AD, N. lehmanni Baba, 1995 (Imitation of Baba, 1995); EH, N. horizontalis Xin, Zhong &amp; Qi, new species, holotype (Lvchun, Yunnan); IL, N. fuscus Didier, 1926 (Ziyuan, Guangxi); MP, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, CD, E, GH, I, KL, M, OP, dorsal view; B, F, J, N lateral view." figureDoi="http://doi.org/10.5281/zenodo.14756551" httpUri="https://zenodo.org/record/14756551/files/figure.png" pageId="13" pageNumber="357">Fig. 3L</figureCitation>
)................................................. 3
</keyLead>
</paragraph>
<paragraph id="8BCA3682FFA3D678FF3CFA84C6A92A42" blockId="13.[151,1437,1264,1608]" box="[151,1436,1381,1403]" pageId="13" pageNumber="357">
<keyLead id="308495B7FFA3D678FF3CFA84C6A92A42" box="[151,1436,1381,1403]" pageId="13" pageNumber="357">
Base of inner margin of protibia direct or with a unclear gap (
<figureCitation id="134E2A07FFA3D678FC9BFA84C08E2A42" box="[816,955,1381,1403]" captionStart="FIGURE 3" captionStartId="5.[151,250,1908,1932]" captionTargetBox="[225,1363,187,1879]" captionTargetId="figure-17@5.[219,1368,181,1885]" captionTargetPageId="5" captionText="FIGURE 3. Male fore-head (AB, EF, IJ, MN), pronotum (C, G, K, O), protibia and apical part of profemur (D, H, L, P) of Neolucanus species. AD, N. lehmanni Baba, 1995 (Imitation of Baba, 1995); EH, N. horizontalis Xin, Zhong &amp; Qi, new species, holotype (Lvchun, Yunnan); IL, N. fuscus Didier, 1926 (Ziyuan, Guangxi); MP, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, CD, E, GH, I, KL, M, OP, dorsal view; B, F, J, N lateral view." figureDoi="http://doi.org/10.5281/zenodo.14756551" httpUri="https://zenodo.org/record/14756551/files/figure.png" pageId="13" pageNumber="357">Figs. 3D, H, P</figureCitation>
)...................................... 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30812E27FFA3D678FF3CFA63C6A92A8D" pageId="13" pageNumber="357">
<paragraph id="8BCA3682FFA3D678FF3CFA63C6A92AA0" blockId="13.[151,1437,1264,1608]" box="[151,1436,1410,1432]" pageId="13" pageNumber="357">
<keyLead id="308495B7FFA3D678FF3CFA63C6A92AA0" box="[151,1436,1410,1432]" pageId="13" pageNumber="357">
3 Mandible slightly curved inward and generally straight in dorsal view (
<figureCitation id="134E2A07FFA3D678FC26FA63C0E52AAF" box="[909,976,1410,1432]" captionStart="FIGURE 3" captionStartId="5.[151,250,1908,1932]" captionTargetBox="[225,1363,187,1879]" captionTargetId="figure-17@5.[219,1368,181,1885]" captionTargetPageId="5" captionText="FIGURE 3. Male fore-head (AB, EF, IJ, MN), pronotum (C, G, K, O), protibia and apical part of profemur (D, H, L, P) of Neolucanus species. AD, N. lehmanni Baba, 1995 (Imitation of Baba, 1995); EH, N. horizontalis Xin, Zhong &amp; Qi, new species, holotype (Lvchun, Yunnan); IL, N. fuscus Didier, 1926 (Ziyuan, Guangxi); MP, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, CD, E, GH, I, KL, M, OP, dorsal view; B, F, J, N lateral view." figureDoi="http://doi.org/10.5281/zenodo.14756551" httpUri="https://zenodo.org/record/14756551/files/figure.png" pageId="13" pageNumber="357">Fig. 3I</figureCitation>
).............
<taxonomicName id="4C754D01FFA3D678FBC0FA63C6A92AA0" ID-CoL="46LRJ" authority="Didier, 1926" authorityName="Didier" authorityYear="1926" box="[1131,1436,1410,1432]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="357" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="B901EA90FFA3D678FBC0FA63C62B2AAF" box="[1131,1310,1410,1431]" italics="true" pageId="13" pageNumber="357">Neolucanus fuscus</emphasis>
<bibRefCitation id="EFE44B73FFA3D678FA8FFA63C6A92AA0" author="Didier, R." box="[1316,1436,1410,1432]" pageId="13" pageNumber="357" pagination="210 - 214" refId="ref7377" refString="Didier, R. (1926) Descriptions sommaires de Lucanides nouveaux, du genre Neolucanus, de la faune indo-chinoise. Bulletin de la Societe Entomologique de France, 1926, 210 - 214. https: // doi. org / 10.3406 / bsef. 1926.27736" type="journal article" year="1926">Didier, 1926</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCA3682FFA3D678FF3CFA41C6A92A8D" blockId="13.[151,1437,1264,1608]" box="[151,1436,1439,1462]" pageId="13" pageNumber="357">
<keyLead id="308495B7FFA3D678FF3CFA41C6A92A8D" box="[151,1436,1439,1462]" pageId="13" pageNumber="357"> Mandible violently curved inward, forming a circular arc in dorsal view.......................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30812E27FFA3D678FF3CFA5CC6A92935" pageId="13" pageNumber="357">
<paragraph id="8BCA3682FFA3D678FF3CFA5CC6A92AD7" blockId="13.[151,1437,1264,1608]" pageId="13" pageNumber="357">
<keyLead id="308495B7FFA3D678FF3CFA5CC6A92AD7" pageId="13" pageNumber="357">
4 Base of inner margin of protibia with a unclear gap; dorsal tooth tiny (
<figureCitation id="134E2A07FFA3D678FC2FFA5CC0DF2AEA" box="[900,1002,1469,1491]" captionStart="FIGURE 3" captionStartId="5.[151,250,1908,1932]" captionTargetBox="[225,1363,187,1879]" captionTargetId="figure-17@5.[219,1368,181,1885]" captionTargetPageId="5" captionText="FIGURE 3. Male fore-head (AB, EF, IJ, MN), pronotum (C, G, K, O), protibia and apical part of profemur (D, H, L, P) of Neolucanus species. AD, N. lehmanni Baba, 1995 (Imitation of Baba, 1995); EH, N. horizontalis Xin, Zhong &amp; Qi, new species, holotype (Lvchun, Yunnan); IL, N. fuscus Didier, 1926 (Ziyuan, Guangxi); MP, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, CD, E, GH, I, KL, M, OP, dorsal view; B, F, J, N lateral view." figureDoi="http://doi.org/10.5281/zenodo.14756551" httpUri="https://zenodo.org/record/14756551/files/figure.png" pageId="13" pageNumber="357">Fig. 3N, P</figureCitation>
)............................................................................................... ..
<taxonomicName id="4C754D01FFA3D678FC16FA3BC6A92AD7" ID-CoL="46LQW" authority="Bomans &amp; Miyashita, 1997" authorityName="Bomans &amp; Miyashita" authorityYear="1997" box="[957,1436,1498,1520]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="357" phylum="Arthropoda" rank="species" species="brochieri">
<emphasis id="B901EA90FFA3D678FC16FA3BC7B92AD7" box="[957,1164,1498,1519]" italics="true" pageId="13" pageNumber="357">Neolucanus brochieri</emphasis>
<bibRefCitation id="EFE44B73FFA3D678FB38FA3AC6A92AD7" author="Bomans, H. E. &amp; Miyashita, T." box="[1171,1436,1498,1520]" pageId="13" pageNumber="357" pagination="2 - 3" refId="ref7344" refString="Bomans, H. E. &amp; Miyashita, T. (1997) Description de trois nouvelles especes de Lucanidae du nord Birmanie. Besoiro, 4, 2 - 3." type="journal article" year="1997">Bomans &amp; Miyashita, 1997</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCA3682FFA3D678FF3CFA19C6A92935" blockId="13.[151,1437,1264,1608]" box="[151,1436,1527,1550]" pageId="13" pageNumber="357">
<keyLead id="308495B7FFA3D678FF3CFA19C6A92935" box="[151,1436,1527,1550]" pageId="13" pageNumber="357">
Base of inner margin of protibia direct; dorsal tooth relatively long (
<figureCitation id="134E2A07FFA3D678FCDBFA19C0E22935" box="[880,983,1528,1550]" captionStart="FIGURE 3" captionStartId="5.[151,250,1908,1932]" captionTargetBox="[225,1363,187,1879]" captionTargetId="figure-17@5.[219,1368,181,1885]" captionTargetPageId="5" captionText="FIGURE 3. Male fore-head (AB, EF, IJ, MN), pronotum (C, G, K, O), protibia and apical part of profemur (D, H, L, P) of Neolucanus species. AD, N. lehmanni Baba, 1995 (Imitation of Baba, 1995); EH, N. horizontalis Xin, Zhong &amp; Qi, new species, holotype (Lvchun, Yunnan); IL, N. fuscus Didier, 1926 (Ziyuan, Guangxi); MP, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, CD, E, GH, I, KL, M, OP, dorsal view; B, F, J, N lateral view." figureDoi="http://doi.org/10.5281/zenodo.14756551" httpUri="https://zenodo.org/record/14756551/files/figure.png" pageId="13" pageNumber="357">Fig. 3B, D</figureCitation>
)..........
<taxonomicName id="4C754D01FFA3D678FBF3FA19C6A92935" ID-CoL="46LS8" authority="Baba, 1995" authorityName="Baba" authorityYear="1995" box="[1112,1436,1527,1550]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="357" phylum="Arthropoda" rank="species" species="lehmanni">
<emphasis id="B901EA90FFA3D678FBF3FA19C61C2935" box="[1112,1321,1528,1549]" italics="true" pageId="13" pageNumber="357">Neolucanus lehmanni</emphasis>
<bibRefCitation id="EFE44B73FFA3D678FA84FA19C6A92935" author="Baba, M." box="[1327,1436,1527,1550]" pageId="13" pageNumber="357" pagination="8 - 11" refId="ref7312" refString="Baba, M. (1995) A new species of the genus Neolucanus Tomson, 1862 from south Burma (Myanmar). Gekkan-Mushi, 292, 8 - 11." type="journal article" year="1995">Baba, 1995</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30812E27FFA3D678FF3CF9F4C6A9297F" pageId="13" pageNumber="357">
<paragraph id="8BCA3682FFA3D678FF3CF9F4C6A92912" blockId="13.[151,1437,1264,1608]" box="[151,1436,1557,1579]" pageId="13" pageNumber="357">
<keyLead id="308495B7FFA3D678FF3CF9F4C6A92912" box="[151,1436,1557,1579]" pageId="13" pageNumber="357">
5 Mandible violently up-curved.................................................
<taxonomicName id="4C754D01FFA3D678FBFBF9F4C6A92912" ID-CoL="46LR7" authority="Didier, 1927" authorityName="Didier" authorityYear="1927" box="[1104,1436,1557,1579]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="357" phylum="Arthropoda" rank="species" species="delicatus">
<emphasis id="B901EA90FFA3D678FBFBF9F4C62B2912" box="[1104,1310,1557,1578]" italics="true" pageId="13" pageNumber="357">Neolucanus delicatus</emphasis>
Didier, 1927
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCA3682FFA3D678FF3CF9D2C6A9297F" blockId="13.[151,1437,1264,1608]" box="[151,1436,1586,1608]" pageId="13" pageNumber="357">
<keyLead id="308495B7FFA3D678FF3CF9D2C6A9297F" box="[151,1436,1586,1608]" pageId="13" pageNumber="357">
Mandible not violently up-curved.............................................
<taxonomicName id="4C754D01FFA3D678FBFBF9D3C6A9297F" ID-CoL="46LRT" authority="Okuda, 2009" authorityName="Okuda" authorityYear="2009" box="[1104,1436,1586,1608]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="357" phylum="Arthropoda" rank="species" species="inaharai">
<emphasis id="B901EA90FFA3D678FBFBF9D3C62E297F" box="[1104,1307,1586,1607]" italics="true" pageId="13" pageNumber="357">Neolucanus inaharai</emphasis>
Okuda, 2009
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,514 @@
<document id="547E8460BF306B0F418EE0A7681459D5" ID-CLB-Dataset="307889" ID-DOI="10.11646/zootaxa.5570.2.6" ID-GBIF-Dataset="2396024e-a9ed-4584-afe7-9fed779d32e1" ID-ISSN="1175-5326" ID-Zenodo-Dep="14735135" ID-ZooBank="8B1BAE9E-FBD3-4AC3-9FA3-9CCC2AA81392" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1737737634945" checkinUser="plazi" docAuthor="Xin, Fei-Yi, Zhong, Xiao-Tian, Guo, Wen-Chi, Zhang, Yi-Feng, Qi, Zhi-Hao &amp; Wu, Song-Qing" docDate="2025" docId="03DC8794FFA7D67FFF3CF977C07B2881" docLanguage="en" docName="zootaxa.5570.2.6.pdf" docOrigin="Zootaxa 5570 (2)" docSource="https://doi.org/10.11646/zootaxa.5570.2.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Neolucanus fuscus Didier 1926" docType="treatment" docVersion="5" lastPageNumber="354" masterDocId="FFE5FFECFFAED675FFABFFE1C3352F38" masterDocTitle="A taxonomic note on Chinese species of the Neolucanus fuscus group with the description of a new species (Coleoptera: Lucanidae: Lucaninae)" masterLastPageNumber="360" masterPageNumber="344" pageNumber="353" updateTime="1738085397281" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="D5CAC153F36A155CBA54F2B9BFD01071" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="673351FFD62C9D8BB59EAA3E43214649">
<mods:title id="CEA7327E0E499630790212530793AFF5">A taxonomic note on Chinese species of the Neolucanus fuscus group with the description of a new species (Coleoptera: Lucanidae: Lucaninae)</mods:title>
</mods:titleInfo>
<mods:name id="BC156C6B380448E1AF85DC3785A31553" type="personal">
<mods:role id="B5C769A4FC25F1FF297EFA450406DFFB">
<mods:roleTerm id="1176B63099A24B8BE1BF9F15AC63B013">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D2235CFB3529F3DC4F1B9CF530FA55B7">Xin, Fei-Yi</mods:namePart>
<mods:nameIdentifier id="731116568C395A15FDD2A15299063EB2" type="ORCID">0009-0002-4407-694X</mods:nameIdentifier>
<mods:affiliation id="A074B261FBA85FA8A890EEDBFFCF014B">Fujian Colleges and Universities Engineering Research Institute of Conservation and Utilization of Natural Bioresources, Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Key Laboratory of Integrated Pest Management in Ecological Forests, Fujian Province University, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China.</mods:affiliation>
</mods:name>
<mods:name id="9E9284C023B8CB578CE80860D0BD2E48" type="personal">
<mods:role id="A8871A5BD620FF71838335BF50DCFD7E">
<mods:roleTerm id="DFE42CFB64A4223F787DEB6414A5B992">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7EC89C00D9270CE83C8FE42956FF08B5">Zhong, Xiao-Tian</mods:namePart>
<mods:nameIdentifier id="DE452C8C1A54E2324C0A2CA004C35182" type="ORCID">0009-0003-1423-9274</mods:nameIdentifier>
<mods:affiliation id="8B19F6DB823C23EE14D0760BB6478452">School of Geoscience and Technology, Southwest Petroleum University, Chengdu 610500, China.</mods:affiliation>
</mods:name>
<mods:name id="131CB1A5F6967AFE78B2C696BEA5829F" type="personal">
<mods:role id="6DE97FBD24772B2FC5F90396EECE3759">
<mods:roleTerm id="E3957F561BAD596623F8E1DAA8A83438">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1C921D8E281FDFD67DE77F5064AF5173">Guo, Wen-Chi</mods:namePart>
<mods:nameIdentifier id="95FD1C3671C2693A4A74489BDAE65E6F" type="ORCID">0009-0008-4116-4016</mods:nameIdentifier>
<mods:affiliation id="0DFA4B009E2AF1502ABAD44D961E6B3E">Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China.</mods:affiliation>
<mods:nameIdentifier id="9BE5307068AB96073F920CE1D6BF9414" type="email">353600225@qq.com</mods:nameIdentifier>
</mods:name>
<mods:name id="10744249E897DA27407359AF50B080F7" type="personal">
<mods:role id="EA0DD857071D175CD34E767A7E0DFD59">
<mods:roleTerm id="A1EFB7C478B7E2F161185047F6B7246E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5706F062684795A2A6FB8DA17AFAA052">Zhang, Yi-Feng</mods:namePart>
<mods:nameIdentifier id="E35CC88DE44BFC5190565F7C25D0635F" type="ORCID">0000-0003-3264-9472</mods:nameIdentifier>
<mods:affiliation id="39A99F9F08C074887A07D756897D1452">Fujian Academy of Forestry, Fuzhou 350012, China.</mods:affiliation>
<mods:nameIdentifier id="8062020BD2995D0DDBAB46554F8C1F0C" type="email">zhang_yifeng@foxmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="C045A59B83F0D00177CA18695371FD16" type="personal">
<mods:role id="2452A77282541137A01835C71542D238">
<mods:roleTerm id="B99730BDA2C9F8D8CF52343CD05FEAC5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="555F9A3CE972E76BD64B892A77FF0270">Qi, Zhi-Hao</mods:namePart>
<mods:nameIdentifier id="C8854215DCEF7DDB1E3A7C3559E07BC7" type="ORCID">0000-0001-8596-0913</mods:nameIdentifier>
<mods:affiliation id="CF1B8A8EAFE469220E5DD1E92DA5D9AD">Key Laboratory of Integrated Pest Management in Ecological Forests, Fujian Province University, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Fujian Academy of Forestry, Fuzhou 350012, China.</mods:affiliation>
</mods:name>
<mods:name id="3B99E3A1AD400E94778004E8E5A5DD6D" type="personal">
<mods:role id="B5395496862DCB1604036B64AC1FA543">
<mods:roleTerm id="014AB6025AC3C4990F5FD5F4162A086D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="378FAFE098E602BDF4540C09BBC93729">Wu, Song-Qing</mods:namePart>
<mods:nameIdentifier id="71FB9AA916F5A070D534099213055632" type="ORCID">0000-0002-2763-5378</mods:nameIdentifier>
<mods:affiliation id="805E9596221E6DD81E030899A1ED6695">Fujian Colleges and Universities Engineering Research Institute of Conservation and Utilization of Natural Bioresources, Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Key Laboratory of Integrated Pest Management in Ecological Forests, Fujian Province University, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="E8CE7F7592864706845830DA59DC4C03">text</mods:typeOfResource>
<mods:relatedItem id="05E402BD59A2B913152C3EAB34702CEA" type="host">
<mods:titleInfo id="642990C869674CBFF51391B9F902B8FD">
<mods:title id="9827FB614527F2B9D394DB5D679C1BFB">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A712570D07519B10B67E13A2E4B083DA">
<mods:date id="C409F3B7295CF683AE82733827102AA0">2025</mods:date>
<mods:detail id="943E84B2E5A0BDB63A78EFE957EBC598" type="pubDate">
<mods:number id="64A604309CD99F719B07204F8AD414A5">2025-01-17</mods:number>
</mods:detail>
<mods:detail id="BBF2799B439B247D1C987ADD720A0DEA" type="volume">
<mods:number id="7BEE8D867DF96CBF69D89B87B993CB04">5570</mods:number>
</mods:detail>
<mods:detail id="559AC104BA1CA69E4D249A8A81DEEE85" type="issue">
<mods:number id="AB8231B64C9486E8E3A7862108F5B17C">2</mods:number>
</mods:detail>
<mods:extent id="1E340ED44B9CD4AA290DC03BAF802A35" unit="page">
<mods:start id="E6DE46F5A89E51C2EF0258692D1EA674">344</mods:start>
<mods:end id="D46F8170D1A8E613C7F079A5F5C724D8">360</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="37BF2990F6ACF8AB28B0AAA588561239">
<mods:url id="4B9C2CF3E8B14F400916EED29C59711A">https://doi.org/10.11646/zootaxa.5570.2.6</mods:url>
</mods:location>
<mods:classification id="ADCE11CB6B56A1CC979BCD31EBCD7A8B">journal article</mods:classification>
<mods:identifier id="4E5734E7BB484D3C2880E1B3A3689913" type="CLB-Dataset">307889</mods:identifier>
<mods:identifier id="D29452ABFE89356D6538E4D25A8A5780" type="DOI">10.11646/zootaxa.5570.2.6</mods:identifier>
<mods:identifier id="C9FEF9674DBAF3F2E4F0046ED20E7EB1" type="GBIF-Dataset">2396024e-a9ed-4584-afe7-9fed779d32e1</mods:identifier>
<mods:identifier id="3AAD26B607B0E65EC376003E9F7A26BC" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="FBE41CC12B128C057FF7BB625854F0FD" type="Zenodo-Dep">14735135</mods:identifier>
<mods:identifier id="0D3F6FAC83CAE879692CDBD4C7A2F9FA" type="ZooBank">8B1BAE9E-FBD3-4AC3-9FA3-9CCC2AA81392</mods:identifier>
</mods:mods>
<treatment id="03DC8794FFA7D67FFF3CF977C07B2881" ID-DOI="http://doi.org/10.5281/zenodo.14756553" ID-Zenodo-Dep="14756553" LSID="urn:lsid:plazi:treatment:03DC8794FFA7D67FFF3CF977C07B2881" httpUri="http://treatment.plazi.org/id/03DC8794FFA7D67FFF3CF977C07B2881" lastPageId="10" lastPageNumber="354" pageId="9" pageNumber="353" scope_family="Lucanidae" scope_order="Coleoptera" scope_subFamily="Lucaninae">
<subSubSection id="C36F6509FFA7D67CFF3CF977C18629EB" pageId="9" pageNumber="353" type="nomenclature">
<paragraph id="8BCA3682FFA7D67CFF3CF977C1292989" blockId="9.[151,815,1686,1786]" box="[151,540,1686,1713]" pageId="9" pageNumber="353">
<heading id="D08281EEFFA7D67CFF3CF977C1292989" bold="true" box="[151,540,1686,1713]" fontSize="11" level="1" pageId="9" pageNumber="353" reason="1">
<taxonomicName id="4C754D01FFA7D67CFF3CF977C1292989" ID-CoL="46LRJ" authority="Didier, 1926" authorityName="Didier" authorityYear="1926" box="[151,540,1686,1713]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="353" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="B901EA90FFA7D67CFF3CF977C1292989" bold="true" box="[151,540,1686,1713]" pageId="9" pageNumber="353">
<emphasis id="B901EA90FFA7D67CFF3CF977C24B2989" bold="true" box="[151,382,1686,1713]" italics="true" pageId="9" pageNumber="353">Neolucanus fuscus</emphasis>
<bibRefCitation id="EFE44B73FFA7D67CFE2EF976C1292989" author="Didier, R." box="[389,540,1687,1713]" pageId="9" pageNumber="353" pagination="210 - 214" refId="ref7377" refString="Didier, R. (1926) Descriptions sommaires de Lucanides nouveaux, du genre Neolucanus, de la faune indo-chinoise. Bulletin de la Societe Entomologique de France, 1926, 210 - 214. https: // doi. org / 10.3406 / bsef. 1926.27736" type="journal article" year="1926">Didier, 1926</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BCA3682FFA7D67CFF3CF95AC18629EB" blockId="9.[151,815,1686,1786]" box="[151,691,1723,1750]" pageId="9" pageNumber="353">
Suggested Chinese common name:
<emphasis id="B901EA90FFA7D67CFD8CF95DC18629EB" bold="true" box="[551,691,1724,1747]" pageId="9" pageNumber="353">Ḃ圆OiNJƤ</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C36F6509FFA7D67CFF3CF93EC01A29C1" box="[151,815,1759,1786]" pageId="9" pageNumber="353" type="description">
<paragraph id="8BCA3682FFA7D67CFF3CF93EC01A29C1" blockId="9.[151,815,1686,1786]" box="[151,815,1759,1786]" pageId="9" pageNumber="353">
<figureCitation id="134E2A07FFA7D67CFF3CF93EC22D29C1" box="[151,280,1759,1785]" captionStart="FIGURE 1" captionStartId="3.[151,250,1908,1932]" captionTargetBox="[312,1275,185,1877]" captionTargetId="figure-17@3.[306,1281,179,1883]" captionTargetPageId="3" captionText="FIGURE 1. Male habitus of Neolucanus species. AB, N. horizontalis Xin, Zhong &amp; Qi, new species, holotype, 36.0 mm, telodont (Lvchun, Yunnan); CE, N. fuscus Didier, 1926; CD, 50.9 mm, telodont (Yên Bái, Yên Bái); E, 44.8 mm, telodont (Ziyuan, Guangxi); FH, N. brochieri Bomans &amp; Miyashita, 1997; FG, 41.1mm, telodont (Gonshan, Yunnan); H, 41.0 mm, telodont (Gonshan, Yunnan). A, C, E, F, H, dorsal view; B, D, G, ventral view. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14735137" httpUri="https://zenodo.org/record/14735137/files/figure.png" pageId="9" pageNumber="353">Figs. 1CE</figureCitation>
;
<figureCitation id="134E2A07FFA7D67CFE8FF93EC25F29C1" box="[292,362,1759,1785]" captionStart="FIGURE 2" captionStartId="4.[151,250,1771,1795]" captionTargetBox="[158,1429,186,1740]" captionTargetId="figure-18@4.[151,1436,181,1747]" captionTargetPageId="4" captionText="FIGURE 2. Male habitus of Neolucanus species. AB, N. horizontalis Xin, Zhong &amp; Qi, new species, paratype, 33.7 mm, mesodont (Jinping, Yunnan); CD, N. fuscus Didier, 1926, 34.7mm, priodont (Wuming, Guangxi); EF, N. brochieri Bomans &amp; Miyashita, 1997, 31.4 mm, priodont (Gonshan, Yunnan). A, C, E, dorsal view; B, D, F, ventral view. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14735139" httpUri="https://zenodo.org/record/14735139/files/figure.png" pageId="9" pageNumber="353">2CD</figureCitation>
;
<figureCitation id="134E2A07FFA7D67CFEDDF93EC29A29C1" box="[374,431,1759,1785]" captionStart="FIGURE 3" captionStartId="5.[151,250,1908,1932]" captionTargetBox="[225,1363,187,1879]" captionTargetId="figure-17@5.[219,1368,181,1885]" captionTargetPageId="5" captionText="FIGURE 3. Male fore-head (AB, EF, IJ, MN), pronotum (C, G, K, O), protibia and apical part of profemur (D, H, L, P) of Neolucanus species. AD, N. lehmanni Baba, 1995 (Imitation of Baba, 1995); EH, N. horizontalis Xin, Zhong &amp; Qi, new species, holotype (Lvchun, Yunnan); IL, N. fuscus Didier, 1926 (Ziyuan, Guangxi); MP, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, CD, E, GH, I, KL, M, OP, dorsal view; B, F, J, N lateral view." figureDoi="http://doi.org/10.5281/zenodo.14756551" httpUri="https://zenodo.org/record/14756551/files/figure.png" pageId="9" pageNumber="353">3IL</figureCitation>
; 4EH;
<figureCitation id="134E2A07FFA7D67CFDA0F93EC17029C2" box="[523,581,1759,1786]" captionStart="FIGURE 5" captionStartId="7.[151,250,1799,1823]" captionTargetBox="[157,1434,185,1769]" captionTargetId="figure-17@7.[151,1436,181,1775]" captionTargetPageId="7" captionText="FIGURE 5. Male abdominal tergite VIII (A, E, I, M), abdominal ventrite VIII (B, F, J, N) and abdominal segment IX (CD, G H, KL, OP) of Neolucanus species. AH, N. horizontalis Xin, Zhong &amp; Qi, new species AD, holotype (Lvchun, Yunnan); EH, paratype (Jinping, Yunnan). IL, N. fuscus Didier, 1926 (Ziyuan, Guangxi). MP, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, C, E, G, I, K, M, O, dorsal view; B, D, F, H, J, L, N, P, ventral view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14756558" httpUri="https://zenodo.org/record/14756558/files/figure.png" pageId="9" pageNumber="353">5IL</figureCitation>
; 6C, G, K; 7C; 8AJ
</paragraph>
</subSubSection>
<subSubSection id="C36F6509FFA7D67CFF3CF8C2C04328CB" pageId="9" pageNumber="353" type="reference_group">
<paragraph id="8BCA3682FFA7D67CFF3CF8C2C1212804" blockId="9.[151,1436,1827,2036]" box="[151,532,1827,1852]" pageId="9" pageNumber="353">
<treatmentCitationGroup id="AB6511ACFFA7D67CFF3CF8C2C1212804" box="[151,532,1827,1852]" pageId="9" pageNumber="353">
<treatmentCitation id="0AD41093FFA7D67CFF3CF8C2C1252804" author="Didier, R." box="[151,528,1827,1852]" page="210" pageId="9" pageNumber="353" year="1926">
<taxonomicName id="4C754D01FFA7D67CFF3CF8C2C1252804" ID-CoL="46LRJ" authority="Didier, 1926: 210" authorityName="Didier" authorityPageNumber="210" authorityYear="1926" box="[151,528,1827,1852]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="353" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="B901EA90FFA7D67CFF3CF8C2C2612804" box="[151,340,1827,1852]" italics="true" pageId="9" pageNumber="353">Neolucanus fuscus</emphasis>
<bibRefCitation id="EFE44B73FFA7D67CFEF0F8C2C1252804" author="Didier, R." box="[347,528,1827,1852]" pageId="9" pageNumber="353" pagination="210 - 214" refId="ref7377" refString="Didier, R. (1926) Descriptions sommaires de Lucanides nouveaux, du genre Neolucanus, de la faune indo-chinoise. Bulletin de la Societe Entomologique de France, 1926, 210 - 214. https: // doi. org / 10.3406 / bsef. 1926.27736" type="journal article" year="1926">Didier, 1926: 210</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCA3682FFA7D67CFF3CF8A3C2C92841" blockId="9.[151,1436,1827,2036]" pageId="9" pageNumber="353">
<treatmentCitationGroup id="AB6511ACFFA7D67CFF3CF8A3C2C92841" pageId="9" pageNumber="353">
<taxonomicName id="4C754D01FFA7D67CFF3CF8A3C25C2862" ID-CoL="7XGC4" authorityName="Lacroix" authorityYear="1972" box="[151,361,1858,1882]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="353" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="B901EA90FFA7D67CFF3CF8A3C25C2862" box="[151,361,1858,1882]" italics="true" pageId="9" pageNumber="353">Neolucanus armatus</emphasis>
</taxonomicName>
:
<treatmentCitation id="0AD41093FFA7D67CFEDCF8A3C1012862" author="Lacroix, J. P." box="[375,564,1858,1882]" page="111" pageId="9" pageNumber="353" year="1972">
<bibRefCitation id="EFE44B73FFA7D67CFEDCF8A3C1012862" author="Lacroix, J. P." box="[375,564,1858,1882]" pageId="9" pageNumber="353" pagination="111 - 126" refId="ref7540" refString="Lacroix, J. P. (1972) Neolucanus nouveaux ou peu connus. Annales de la Societe Entomologique de France, New Series, 8 (1), 111 - 126. https: // doi. org / 10.1080 / 21686351.1972.12278925" type="journal article" year="1972">Lacroix 1972: 111</bibRefCitation>
</treatmentCitation>
, fig. 13,
<typeStatus id="54CE8820FFA7D67CFD37F8A2C1FD2863" box="[668,712,1859,1883]" pageId="9" pageNumber="353">type</typeStatus>
locality:
<collectingCountry id="F3627612FFA7D67CFC83F8A3C0B52862" box="[808,896,1858,1882]" name="Vietnam" pageId="9" pageNumber="353">Vietnam</collectingCountry>
, Phố Hà Nội Province, Tam Dao;
<treatmentCitation id="0AD41093FFA7D67CFB75F8A2C6A92862" author="Nguyen, T. Q." box="[1246,1436,1858,1883]" page="382" pageId="9" pageNumber="353" year="2013">
<bibRefCitation id="EFE44B73FFA7D67CFB75F8A2C6A92862" author="Nguyen, T. Q." box="[1246,1436,1858,1883]" pageId="9" pageNumber="353" pagination="377 - 384" refId="ref7754" refString="Nguyen, T. Q. (2013) Description of a new species of the genus Neolucanus Thomson, 1862 (Coleoptera: Lucanidae) from central Vietnam. Zootaxa, 3741 (3), 377 - 384. https: // doi. org / 10.11646 / zootaxa. 3741.3.6" type="journal article" year="2013">Nguyen 2013: 382</bibRefCitation>
</treatmentCitation>
(synonymized with
<taxonomicName id="4C754D01FFA7D67CFE3BF883C2D82841" ID-CoL="46LRJ" authorityName="Didier" authorityYear="1926" box="[400,493,1889,1913]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="353" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="B901EA90FFA7D67CFE3BF883C2942841" box="[400,417,1890,1913]" italics="true" pageId="9" pageNumber="353">N</emphasis>
.
<emphasis id="B901EA90FFA7D67CFE05F880C2D82841" box="[430,493,1889,1913]" italics="true" pageId="9" pageNumber="353">fuscus</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCA3682FFA7D67CFF3CF89EC2C9288E" blockId="9.[151,1436,1827,2036]" pageId="9" pageNumber="353">
<treatmentCitationGroup id="AB6511ACFFA7D67CFF3CF89EC2C9288E" pageId="9" pageNumber="353">
<taxonomicName id="4C754D01FFA7D67CFF3CF89EC24F28AF" ID-CoL="7XGCH" authorityName="Lacroix" authorityYear="1978" box="[151,378,1919,1943]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="353" phylum="Arthropoda" rank="species" species="sianoukei">
<emphasis id="B901EA90FFA7D67CFF3CF89EC24F28AF" box="[151,378,1919,1943]" italics="true" pageId="9" pageNumber="353">Neolucanus sianoukei</emphasis>
</taxonomicName>
:
<treatmentCitation id="0AD41093FFA7D67CFE2CF89EC17E28A0" author="Lacroix, J. P." box="[391,587,1919,1944]" page="270" pageId="9" pageNumber="353" year="1978">
<bibRefCitation id="EFE44B73FFA7D67CFE2CF89EC17E28A0" author="Lacroix, J. P." box="[391,587,1919,1944]" pageId="9" pageNumber="353" pagination="249 - 294" refId="ref7585" refString="Lacroix, J. P. (1978) Contribution a l'etudes des Coleopteres Lucanides du globe. Deux genres nouveaux et onze especes inedites (Chiasognathinae, Lucaninae, Chalcodinae, Cladognathinae, Dorcinae). Bulletin et Annales de la Societe Royale d'Entomologie de Belgique, 114, 249 - 294." type="journal article" year="1978">Lacroix 1978: 270</bibRefCitation>
</treatmentCitation>
, fig. 1,
<typeStatus id="54CE8820FFA7D67CFD09F861C1FB28A0" box="[674,718,1920,1944]" pageId="9" pageNumber="353">type</typeStatus>
locality:
<collectingCountry id="F3627612FFA7D67CFC9FF89EC0B928AF" box="[820,908,1919,1943]" name="Vietnam" pageId="9" pageNumber="353">Vietnam</collectingCountry>
, Lâm Đồng Province, Ðà Lạt;
<treatmentCitation id="0AD41093FFA7D67CFB72F861C6A928A0" author="Nguyen, T. Q." box="[1241,1436,1920,1944]" page="382" pageId="9" pageNumber="353" year="2013">
<bibRefCitation id="EFE44B73FFA7D67CFB72F861C6A928A0" author="Nguyen, T. Q." box="[1241,1436,1920,1944]" pageId="9" pageNumber="353" pagination="377 - 384" refId="ref7754" refString="Nguyen, T. Q. (2013) Description of a new species of the genus Neolucanus Thomson, 1862 (Coleoptera: Lucanidae) from central Vietnam. Zootaxa, 3741 (3), 377 - 384. https: // doi. org / 10.11646 / zootaxa. 3741.3.6" type="journal article" year="2013">Nguyen 2013: 382</bibRefCitation>
</treatmentCitation>
(synonymized with
<taxonomicName id="4C754D01FFA7D67CFE3BF87EC2D8288E" ID-CoL="46LRJ" authorityName="Didier" authorityYear="1926" box="[400,493,1950,1974]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="353" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="B901EA90FFA7D67CFE3BF87EC294288E" box="[400,417,1951,1974]" italics="true" pageId="9" pageNumber="353">N</emphasis>
.
<emphasis id="B901EA90FFA7D67CFE05F87FC2D8288E" box="[430,493,1950,1974]" italics="true" pageId="9" pageNumber="353">fuscus</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCA3682FFA7D67CFF3CF85CC67B28ED" blockId="9.[151,1436,1827,2036]" box="[151,1358,1981,2005]" pageId="9" pageNumber="353">
<treatmentCitationGroup id="AB6511ACFFA7D67CFF3CF85CC67B28ED" box="[151,1358,1981,2005]" pageId="9" pageNumber="353">
<taxonomicName id="4C754D01FFA7D67CFF3CF85CC24D28ED" ID-CoL="46LTS" authorityName="Schenk" authorityYear="2012" box="[151,376,1981,2005]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="353" phylum="Arthropoda" rank="species" species="zhongguo">
<emphasis id="B901EA90FFA7D67CFF3CF85CC24D28ED" box="[151,376,1981,2005]" italics="true" pageId="9" pageNumber="353">Neolucanus zhongguo</emphasis>
</taxonomicName>
:
<materialsCitation id="3B1D3CDFFFA7D67CFE2EF85CC67F28ED" box="[389,1354,1981,2005]" country="China" location="Dayao Mountain" municipality="Liuzhou City" pageId="9" pageNumber="353" specimenCount="1" stateProvince="Guangxi">
<treatmentCitation id="0AD41093FFA7D67CFE2EF85CC11128ED" author="Schenk, K. D." box="[389,548,1981,2005]" page="3" pageId="9" pageNumber="353" year="2012">
<bibRefCitation id="EFE44B73FFA7D67CFE2EF85CC11128ED" author="Schenk, K. D." box="[389,548,1981,2005]" pageId="9" pageNumber="353" pagination="4 - 38" refId="ref7844" refString="Schenk, K. D. (2012) Notes to the Lucanidae of Asia and description of new taxa of the genus Neolucanus. Beetles World, 6, 4 - 38." type="journal article" year="2012">Schenk 2012: 3</bibRefCitation>
</treatmentCitation>
, fig. 1,
<typeStatus id="54CE8820FFA7D67CFDDAF85CC1A828ED" box="[625,669,1981,2005]" pageId="9" pageNumber="353">type</typeStatus>
locality:
<collectingCountry id="F3627612FFA7D67CFD56F85CC00928ED" box="[765,828,1981,2005]" name="China" pageId="9" pageNumber="353">China</collectingCountry>
,
<collectingRegion id="49B1F860FFA7D67CFCEDF85CC73428ED" box="[838,1025,1981,2005]" country="China" name="Guangxi" pageId="9" pageNumber="353">Guangxi Province</collectingRegion>
,
<collectingMunicipality id="6BAEACF8FFA7D67CFBA1F85CC7A728ED" box="[1034,1170,1981,2005]" pageId="9" pageNumber="353">Liuzhou City</collectingMunicipality>
,
<location id="8EAA6059FFA7D67CFB37F85CC67F28ED" LSID="urn:lsid:plazi:treatment:03DC8794FFA7D67FFF3CF977C07B2881:8EAA6059FFA7D67CFB37F85CC67F28ED" box="[1180,1354,1981,2005]" country="China" municipality="Liuzhou City" name="Dayao Mountain" pageId="9" pageNumber="353" stateProvince="Guangxi">Dayao Mountain</location>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCA3682FFA7D67CFF3CF83AC04328CB" blockId="9.[151,1436,1827,2036]" box="[151,886,2011,2036]" pageId="9" pageNumber="353">
<treatmentCitationGroup id="AB6511ACFFA7D67CFF3CF83AC04328CB" box="[151,886,2011,2036]" pageId="9" pageNumber="353">
<taxonomicName id="4C754D01FFA7D67CFF3CF83AC28B28CB" ID-CoL="5JV8L" authorityName="Schenk" authorityYear="2012" box="[151,446,2011,2036]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="353" phylum="Arthropoda" rank="subSpecies" species="fuscus" subSpecies="zhongguo">
<emphasis id="B901EA90FFA7D67CFF3CF83AC28B28CB" box="[151,446,2011,2036]" italics="true" pageId="9" pageNumber="353">Neolucanus fuscus zhongguo</emphasis>
</taxonomicName>
:
<treatmentCitation id="0AD41093FFA7D67CFE60F83AC14228CB" author="Schenk, K. D." box="[459,631,2011,2036]" page="35" pageId="9" pageNumber="353" year="2014">
<bibRefCitation id="EFE44B73FFA7D67CFE60F83AC14228CB" author="Schenk, K. D." box="[459,631,2011,2036]" pageId="9" pageNumber="353" pagination="2 - 8" refId="ref7878" refString="Schenk, K. D. (2014) Catalogue of Lucanidae: Genus Neolucanus Thomson, 1862. Beetles World, 10, 2 - 8." type="journal article" year="2014">Schenk 2014: 35</bibRefCitation>
</treatmentCitation>
, fig. 42,
<emphasis id="B901EA90FFA7D67CFD7BF83DC04528CC" bold="true" box="[720,880,2012,2036]" pageId="9" pageNumber="353">new synonymy</emphasis>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36F6509FFA4D67FFF3CFF76C21F2E95" pageId="10" pageNumber="354" type="materials_examined">
<paragraph id="8BCA3682FFA4D67FFF3CFF76C1862F89" blockId="10.[151,1437,150,1977]" box="[151,691,150,177]" pageId="10" pageNumber="354">
<materialsCitation id="3B1D3CDFFFA4D67FFF3CFF76C19B2F89" box="[151,686,150,177]" country="Vietnam" location="Tuyen Quang Province" pageId="10" pageNumber="354" specimenCount="1" stateProvince="Tuyen Quang" typeStatus="holotype">
<emphasis id="B901EA90FFA4D67FFF3CFF76C2032F88" bold="true" box="[151,310,150,177]" pageId="10" pageNumber="354">
<typeStatus id="54CE8820FFA4D67FFF3CFF76C3E72F89" box="[151,210,151,177]" pageId="10" pageNumber="354">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3627612FFA4D67FFE97FF76C2AB2F89" box="[316,414,151,177]" name="Vietnam" pageId="10" pageNumber="354">Vietnam</collectingCountry>
,
<collectingRegion id="49B1F860FFA4D67FFE03FF76C19B2F89" box="[424,686,151,177]" country="Vietnam" name="Tuyen Quang" pageId="10" pageNumber="354">Tuyên Quang Province</collectingRegion>
</materialsCitation>
.
</paragraph>
<paragraph id="8BCA3682FFA4D67FFF6CFF5AC10F2FED" blockId="10.[151,1437,150,1977]" box="[199,570,186,213]" pageId="10" pageNumber="354">
<emphasis id="B901EA90FFA4D67FFF6CFF5AC2DA2FEC" bold="true" box="[199,495,186,213]" pageId="10" pageNumber="354">
<typeStatus id="54CE8820FFA4D67FFF6CFF5AC2372FED" box="[199,258,187,213]" pageId="10" pageNumber="354">Type</typeStatus>
material examined.
</emphasis>
None.
</paragraph>
<paragraph id="8BCA3682FFA4D67FFF6CFF3FC21F2E95" blockId="10.[151,1437,150,1977]" pageId="10" pageNumber="354">
<emphasis id="B901EA90FFA4D67FFF6CFF3FC1092FC0" bold="true" box="[199,572,222,248]" pageId="10" pageNumber="354">Additional material examined.</emphasis>
<materialsCitation id="3B1D3CDFFFA4D67FFDEDFF3FC2AB2E25" collectingDate="2021-06" collectionCode="CFYX" collectorName="You Xiao" country="China" county="Ziyuan County" location="Guilin City" pageId="10" pageNumber="354" specimenCount="2" specimenCount-male="2" stateProvince="Guangxi" typeStatus="holotype">
<emphasis id="B901EA90FFA4D67FFDEDFF3FC1AC2FC0" bold="true" box="[582,665,222,248]" pageId="10" pageNumber="354">
<collectingCountry id="F3627612FFA4D67FFDEDFF3FC1A62FC0" box="[582,659,222,248]" name="China" pageId="10" pageNumber="354">China</collectingCountry>
:
</emphasis>
<specimenCount id="9D73FD0BFFA4D67FFD0FFF3EC1C82FC1" box="[676,765,223,249]" count="2" pageId="10" pageNumber="354" type="male">2 males</specimenCount>
(
<collectionCode id="ED64AE47FFA4D67FFCBBFF3EC0552FC1" box="[784,864,223,249]" pageId="10" pageNumber="354">CFYX</collectionCode>
),
<collectingRegion id="49B1F860FFA4D67FFCDCFF3EC77C2FC1" box="[887,1097,223,249]" country="China" name="Guangxi" pageId="10" pageNumber="354">Guangxi Province</collectingRegion>
,
<location id="8EAA6059FFA4D67FFBF3FF3EC7EE2FC1" LSID="urn:lsid:plazi:treatment:03DC8794FFA7D67FFF3CF977C07B2881:8EAA6059FFA4D67FFBF3FF3EC7EE2FC1" box="[1112,1243,223,249]" country="China" county="Ziyuan County" name="Guilin City" pageId="10" pageNumber="354" stateProvince="Guangxi">Guilin City</location>
,
<collectingCounty id="62AB4E0EFFA4D67FFB41FF3EC6AC2FC1" box="[1258,1433,223,249]" pageId="10" pageNumber="354">Ziyuan County</collectingCounty>
,
<date id="FFCB1042FFA4D67FFF3CFEE2C3C22E25" box="[151,247,259,285]" pageId="10" pageNumber="354" value="2021-06">
<collectingDate id="EF8FE9AAFFA4D67FFF3CFEE2C3C22E25" box="[151,247,259,285]" pageId="10" pageNumber="354" value="2021-06">VI.2021</collectingDate>
</date>
,
<collectorName id="26805354FFA4D67FFEABFEE2C25E2E25" box="[256,363,259,285]" pageId="10" pageNumber="354">You Xiao</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B1D3CDFFFA4D67FFE02FEE2C2442E79" collectingDate="2023-06-03" collectionCode="CFYX" country="China" county="Wuming County" location="Tianping" municipality="Nanning City" pageId="10" pageNumber="354" specimenCount="2" specimenCount-male="2" stateProvince="Guangxi" typeStatus="holotype">
<specimenCount id="9D73FD0BFFA4D67FFE02FEE2C2CA2E25" box="[425,511,259,285]" count="2" pageId="10" pageNumber="354" type="male">2 males</specimenCount>
(
<collectionCode id="ED64AE47FFA4D67FFDA5FEE2C16B2E25" box="[526,606,259,285]" pageId="10" pageNumber="354">CFYX</collectionCode>
),
<collectingRegion id="49B1F860FFA4D67FFDDAFEE2C00B2E25" box="[625,830,259,285]" country="China" name="Guangxi" pageId="10" pageNumber="354">Guangxi Province</collectingRegion>
,
<collectingMunicipality id="6BAEACF8FFA4D67FFCE1FEE2C0D72E25" box="[842,994,259,285]" pageId="10" pageNumber="354">Nanning City</collectingMunicipality>
,
<collectingCounty id="62AB4E0EFFA4D67FFC47FEE2C79D2E25" box="[1004,1192,259,285]" pageId="10" pageNumber="354">Wuming County</collectingCounty>
,
<location id="8EAA6059FFA4D67FFB19FEE2C62C2E25" LSID="urn:lsid:plazi:treatment:03DC8794FFA7D67FFF3CF977C07B2881:8EAA6059FFA4D67FFB19FEE2C62C2E25" box="[1202,1305,259,285]" country="China" county="Wuming County" municipality="Nanning City" name="Tianping" pageId="10" pageNumber="354" stateProvince="Guangxi">Tianping</location>
,
<date id="FFCB1042FFA4D67FFA8FFEE2C6AD2E25" box="[1316,1432,259,285]" pageId="10" pageNumber="354" value="2023-06-03">
<collectingDate id="EF8FE9AAFFA4D67FFA8FFEE2C6AD2E25" box="[1316,1432,259,285]" pageId="10" pageNumber="354" value="2023-06-03">3.VI.2023</collectingDate>
</date>
, local collector leg.;
</materialsCitation>
<materialsCitation id="3B1D3CDFFFA4D67FFED2FEC6C6092E79" box="[377,1340,295,321]" collectingDate="2010-06" collectionCode="CZHQ" country="China" location="Daming Mountain" pageId="10" pageNumber="354" specimenCount="1" specimenCount-male="1" stateProvince="Guangxi" typeStatus="holotype">
<specimenCount id="9D73FD0BFFA4D67FFED2FEC6C2F32E79" box="[377,454,295,321]" count="1" pageId="10" pageNumber="354" type="male">1 male</specimenCount>
(
<collectionCode id="ED64AE47FFA4D67FFE7DFEC6C11D2E79" box="[470,552,295,321]" pageId="10" pageNumber="354">CZHQ</collectionCode>
),
<collectingRegion id="49B1F860FFA4D67FFD96FEC6C03E2E79" box="[573,779,295,321]" country="China" name="Guangxi" pageId="10" pageNumber="354">Guangxi Province</collectingRegion>
,
<location id="8EAA6059FFA4D67FFCB3FEC6C0DE2E79" LSID="urn:lsid:plazi:treatment:03DC8794FFA7D67FFF3CF977C07B2881:8EAA6059FFA4D67FFCB3FEC6C0DE2E79" box="[792,1003,295,321]" country="China" name="Daming Mountain" pageId="10" pageNumber="354" stateProvince="Guangxi">Daming Mountain</location>
,
<date id="FFCB1042FFA4D67FFC5DFEC6C7632E79" box="[1014,1110,295,321]" pageId="10" pageNumber="354" value="2010-06">
<collectingDate id="EF8FE9AAFFA4D67FFC5DFEC6C7632E79" box="[1014,1110,295,321]" pageId="10" pageNumber="354" value="2010-06">VI.2010</collectingDate>
</date>
, local collector leg.;
</materialsCitation>
<materialsCitation id="3B1D3CDFFFA4D67FFAEEFEC6C7502E5D" collectingDate="2013-07-02" collectingDateMax="2013-07-08" collectingDateMin="2013-07-02" collectionCode="CZHQ" collectorName="Yanquan Lu" country="China" elevation="1300" location="Daming Mountain" pageId="10" pageNumber="354" specimenCount="4" specimenCount-male="4" stateProvince="Guangxi" typeStatus="holotype">
<specimenCount id="9D73FD0BFFA4D67FFAEEFEC6C6A92E79" box="[1349,1436,295,321]" count="4" pageId="10" pageNumber="354" type="male">4 males</specimenCount>
(
<collectionCode id="ED64AE47FFA4D67FFF34FEAAC3C42E5D" box="[159,241,331,357]" pageId="10" pageNumber="354">CZHQ</collectionCode>
),
<collectingRegion id="49B1F860FFA4D67FFEA8FEAAC2FA2E5D" box="[259,463,331,357]" country="China" name="Guangxi" pageId="10" pageNumber="354">Guangxi Province</collectingRegion>
,
<location id="8EAA6059FFA4D67FFE72FEAAC19C2E5D" LSID="urn:lsid:plazi:treatment:03DC8794FFA7D67FFF3CF977C07B2881:8EAA6059FFA4D67FFE72FEAAC19C2E5D" box="[473,681,331,357]" country="China" name="Daming Mountain" pageId="10" pageNumber="354" stateProvince="Guangxi">Daming Mountain</location>
,
<date id="FFCB1042FFA4D67FFD19FEAAC0792E5D" box="[690,844,331,357]" pageId="10" pageNumber="354" value="2013-07-02" valueMax="2013-07-08" valueMin="2013-07-02">
<collectingDate id="EF8FE9AAFFA4D67FFD19FEAAC0792E5D" box="[690,844,331,357]" pageId="10" pageNumber="354" value="2013-07-02" valueMax="2013-07-08" valueMin="2013-07-02">28.VII.2013</collectingDate>
</date>
,
<quantity id="4C8D9B67FFA4D67FFCFDFEAAC0932E5D" box="[854,934,331,357]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="10" pageNumber="354" unit="m" value="1300.0">
<elevation id="0058D1B1FFA4D67FFCFDFEAAC0932E5D" box="[854,934,331,357]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="10" pageNumber="354" unit="m" value="1300.0">1300m</elevation>
</quantity>
,
<collectorName id="26805354FFA4D67FFC04FEAAC7032E5D" box="[943,1078,331,357]" pageId="10" pageNumber="354">Yanquan Lu</collectorName>
leg.
</materialsCitation>
<materialsCitation id="3B1D3CDFFFA4D67FFBC1FEABC1B92EB1" collectionCode="CXTZ" country="Vietnam" location="Yen Bai Province" pageId="10" pageNumber="354" specimenCount="8" specimenCount-male="8" stateProvince="Yen Bai" typeStatus="holotype">
<emphasis id="B901EA90FFA4D67FFBC1FEABC7EC2E5C" bold="true" box="[1130,1241,330,356]" pageId="10" pageNumber="354">
<collectingCountry id="F3627612FFA4D67FFBC1FEABC7E12E5C" box="[1130,1236,330,356]" name="Vietnam" pageId="10" pageNumber="354">Vietnam</collectingCountry>
:
</emphasis>
<specimenCount id="9D73FD0BFFA4D67FFB74FEAAC6012E5D" box="[1247,1332,331,357]" count="8" pageId="10" pageNumber="354" type="male">8 males</specimenCount>
(
<collectionCode id="ED64AE47FFA4D67FFAEAFEAAC6A52E5D" box="[1345,1424,331,357]" pageId="10" pageNumber="354">CXTZ</collectionCode>
),
<collectingRegion id="49B1F860FFA4D67FFF3CFE8EC2512EB1" box="[151,356,367,393]" country="Vietnam" name="Yen Bai" pageId="10" pageNumber="354">Yên Bái Province</collectingRegion>
, 2023, local collector leg
</materialsCitation>
.;
<materialsCitation id="3B1D3CDFFFA4D67FFD35FE8EC66D2EB1" box="[670,1368,367,393]" collectingDate="2024-06" collectionCode="CZHQ" country="Vietnam" location="City" pageId="10" pageNumber="354" specimenCount="18" specimenCount-male="18" stateProvince="Yen Bai" typeStatus="holotype">
<specimenCount id="9D73FD0BFFA4D67FFD35FE8EC0312EB1" box="[670,772,367,393]" count="18" pageId="10" pageNumber="354" type="male">18 males</specimenCount>
(
<collectionCode id="ED64AE47FFA4D67FFCBEFE8EC0522EB1" box="[789,871,367,393]" pageId="10" pageNumber="354">CZHQ</collectionCode>
),
<collectingRegion id="49B1F860FFA4D67FFCD0FE8EC77D2EB1" box="[891,1096,367,393]" country="Vietnam" name="Yen Bai" pageId="10" pageNumber="354">Yên Bái Province</collectingRegion>
, Yên Bái
<location id="8EAA6059FFA4D67FFB17FE8EC7D82EB1" LSID="urn:lsid:plazi:treatment:03DC8794FFA7D67FFF3CF977C07B2881:8EAA6059FFA4D67FFB17FE8EC7D82EB1" box="[1212,1261,367,393]" country="Vietnam" name="City" pageId="10" pageNumber="354" stateProvince="Yen Bai">City</location>
,
<date id="FFCB1042FFA4D67FFB52FE8EC66D2EB1" box="[1273,1368,367,393]" pageId="10" pageNumber="354" value="2024-06">
<collectingDate id="EF8FE9AAFFA4D67FFB52FE8EC66D2EB1" box="[1273,1368,367,393]" pageId="10" pageNumber="354" value="2024-06">VI.2024</collectingDate>
</date>
</materialsCitation>
, local collector leg.
</paragraph>
</subSubSection>
<subSubSection id="C36F6509FFA4D67FFF6CFE57C0892931" pageId="10" pageNumber="354" type="description">
<paragraph id="8BCA3682FFA4D67FFF6CFE57C1602EE8" blockId="10.[151,1437,150,1977]" box="[199,597,438,465]" pageId="10" pageNumber="354">
<emphasis id="B901EA90FFA4D67FFF6CFE57C1602EE8" bold="true" box="[199,597,438,465]" pageId="10" pageNumber="354">Redescription of the male (n=35).</emphasis>
</paragraph>
<paragraph id="8BCA3682FFA4D67FFF6CFE3AC7B92ECD" blockId="10.[151,1437,150,1977]" box="[199,1164,474,501]" pageId="10" pageNumber="354">
Body length:
<quantity id="4C8D9B67FFA4D67FFECBFE3BC1362ECD" box="[352,515,474,501]" metricMagnitude="-2" metricUnit="m" metricValue="4.3149999999999995" metricValueMax="5.71" metricValueMin="2.92" pageId="10" pageNumber="354" unit="mm" value="43.15" valueMax="57.1" valueMin="29.2">29.257.1 mm</quantity>
(
<bibRefCitation id="EFE44B73FFA4D67FFDBBFE3AC1A22ECD" author="Fujita, H." box="[528,663,475,501]" pageId="10" pageNumber="354" refId="ref7425" refString="Fujita, H. (2010) The Lucanid Beetles of the World. Mushi-Sha, Tokyo, 472 pp., 248 pls." type="book" year="2010">Fujita 2010</bibRefCitation>
;
<bibRefCitation id="EFE44B73FFA4D67FFD0AFE3AC0662ECD" author="Toshiyuki, T." box="[673,851,475,501]" pageId="10" pageNumber="354" pagination="1 - 87" refId="ref7955" refString="Toshiyuki, T. (2021) Illustrated guide to the genus Neolucanus of the world. Be-kuwa, 78, 1 - 87." type="journal article" year="2021">Toshiyuki 2021</bibRefCitation>
;
<bibRefCitation id="EFE44B73FFA4D67FFCF6FE3AC0822ECD" author="Yi, D." box="[861,951,475,501]" pageId="10" pageNumber="354" refId="ref7980" refString="Yi, D. (2023) Lucanidae of the World. Design Spacetime, Seoul, 190 pp., 160 pls." type="book" year="2023">Yi 2023</bibRefCitation>
and present paper)
</paragraph>
<paragraph id="8BCA3682FFA4D67FFF6CFE1EC3C52D59" blockId="10.[151,1437,150,1977]" pageId="10" pageNumber="354">
<emphasis id="B901EA90FFA4D67FFF6CFE1EC2022D21" bold="true" box="[199,311,511,537]" pageId="10" pageNumber="354">Telodont.</emphasis>
<emphasis id="B901EA90FFA4D67FFE97FE1EC2A02D21" box="[316,405,511,537]" italics="true" pageId="10" pageNumber="354">Habitus</emphasis>
(
<figureCitation id="134E2A07FFA4D67FFE08FE1EC12C2D21" box="[419,537,511,537]" captionStart="FIGURE 1" captionStartId="3.[151,250,1908,1932]" captionTargetBox="[312,1275,185,1877]" captionTargetId="figure-17@3.[306,1281,179,1883]" captionTargetPageId="3" captionText="FIGURE 1. Male habitus of Neolucanus species. AB, N. horizontalis Xin, Zhong &amp; Qi, new species, holotype, 36.0 mm, telodont (Lvchun, Yunnan); CE, N. fuscus Didier, 1926; CD, 50.9 mm, telodont (Yên Bái, Yên Bái); E, 44.8 mm, telodont (Ziyuan, Guangxi); FH, N. brochieri Bomans &amp; Miyashita, 1997; FG, 41.1mm, telodont (Gonshan, Yunnan); H, 41.0 mm, telodont (Gonshan, Yunnan). A, C, E, F, H, dorsal view; B, D, G, ventral view. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14735137" httpUri="https://zenodo.org/record/14735137/files/figure.png" pageId="10" pageNumber="354">Fig. 1CE</figureCitation>
). Head and pronotum black, elytra black or large areas yellowish; luster relatively matte; body with invisible pubescence in dorsal view, labium with dense setae, all legs with sparse yellow setae on surface.
</paragraph>
<paragraph id="8BCA3682FFA4D67FFF6CFD8AC0492C65" blockId="10.[151,1437,150,1977]" pageId="10" pageNumber="354">
<emphasis id="B901EA90FFA4D67FFF6CFD8AC2312DBD" box="[199,260,619,645]" italics="true" pageId="10" pageNumber="354">Head</emphasis>
transverse. Anterior ridge raised behind mandible and widely concave on the middle. Canthus long and wide, completely surround the eye. Clypeolabrum (
<figureCitation id="134E2A07FFA4D67FFCBAFD6EC06A2D91" box="[785,863,655,681]" captionStart="FIGURE 3" captionStartId="5.[151,250,1908,1932]" captionTargetBox="[225,1363,187,1879]" captionTargetId="figure-17@5.[219,1368,181,1885]" captionTargetPageId="5" captionText="FIGURE 3. Male fore-head (AB, EF, IJ, MN), pronotum (C, G, K, O), protibia and apical part of profemur (D, H, L, P) of Neolucanus species. AD, N. lehmanni Baba, 1995 (Imitation of Baba, 1995); EH, N. horizontalis Xin, Zhong &amp; Qi, new species, holotype (Lvchun, Yunnan); IL, N. fuscus Didier, 1926 (Ziyuan, Guangxi); MP, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, CD, E, GH, I, KL, M, OP, dorsal view; B, F, J, N lateral view." figureDoi="http://doi.org/10.5281/zenodo.14756551" httpUri="https://zenodo.org/record/14756551/files/figure.png" pageId="10" pageNumber="354">Fig. 3I</figureCitation>
) almost invisible in dorsal view, fused with frons. Mandible (
<figureCitation id="134E2A07FFA4D67FFEB8FD52C24C2DF5" box="[275,377,691,717]" captionStart="FIGURE 3" captionStartId="5.[151,250,1908,1932]" captionTargetBox="[225,1363,187,1879]" captionTargetId="figure-17@5.[219,1368,181,1885]" captionTargetPageId="5" captionText="FIGURE 3. Male fore-head (AB, EF, IJ, MN), pronotum (C, G, K, O), protibia and apical part of profemur (D, H, L, P) of Neolucanus species. AD, N. lehmanni Baba, 1995 (Imitation of Baba, 1995); EH, N. horizontalis Xin, Zhong &amp; Qi, new species, holotype (Lvchun, Yunnan); IL, N. fuscus Didier, 1926 (Ziyuan, Guangxi); MP, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, CD, E, GH, I, KL, M, OP, dorsal view; B, F, J, N lateral view." figureDoi="http://doi.org/10.5281/zenodo.14756551" httpUri="https://zenodo.org/record/14756551/files/figure.png" pageId="10" pageNumber="354">Fig. 3IJ</figureCitation>
) smooth from base to teeth on inner margin; in lateral view, clearly up-curved, a dorsal tooth behind apex forming an approximately acute-angle and significantly protrudes beyond upper margin, and protrudes to almost same length as lower tooth; in dorsal view, slightly curved, thickness almost regular from base to inner teeth, with 47 continuous small teeth behind apex and getting smaller apically. Antennal club with 3 antennomeres; antennomeres 89 subquadrilateral; antennomere 10 semi-elliptic.
</paragraph>
<paragraph id="8BCA3682FFA4D67FFF6CFC86C2E52CF1" blockId="10.[151,1437,150,1977]" pageId="10" pageNumber="354">
<emphasis id="B901EA90FFA4D67FFF6CFC86C2032CB8" box="[199,310,871,896]" italics="true" pageId="10" pageNumber="354">Pronotum</emphasis>
(
<figureCitation id="134E2A07FFA4D67FFEEFFC86C2AC2CB9" box="[324,409,871,897]" captionStart="FIGURE 3" captionStartId="5.[151,250,1908,1932]" captionTargetBox="[225,1363,187,1879]" captionTargetId="figure-17@5.[219,1368,181,1885]" captionTargetPageId="5" captionText="FIGURE 3. Male fore-head (AB, EF, IJ, MN), pronotum (C, G, K, O), protibia and apical part of profemur (D, H, L, P) of Neolucanus species. AD, N. lehmanni Baba, 1995 (Imitation of Baba, 1995); EH, N. horizontalis Xin, Zhong &amp; Qi, new species, holotype (Lvchun, Yunnan); IL, N. fuscus Didier, 1926 (Ziyuan, Guangxi); MP, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, CD, E, GH, I, KL, M, OP, dorsal view; B, F, J, N lateral view." figureDoi="http://doi.org/10.5281/zenodo.14756551" httpUri="https://zenodo.org/record/14756551/files/figure.png" pageId="10" pageNumber="354">Fig. 3K</figureCitation>
) transverse, markedly longer and wider than head with slightly raised edges.Anterior margin bisinuate, posterior margin slightly bisinuate. Anterior angles relatively sharp and directed forwards, lateral angles and posterior angles obtuse.
</paragraph>
<paragraph id="8BCA3682FFA4D67FFF6CFC32C0F72CD5" blockId="10.[151,1437,150,1977]" box="[199,962,979,1005]" pageId="10" pageNumber="354">
<emphasis id="B901EA90FFA4D67FFF6CFC32C2382CD5" box="[199,269,979,1005]" italics="true" pageId="10" pageNumber="354">Elytra</emphasis>
elongate, anterior margin obviously narrower than pronotum.
</paragraph>
<paragraph id="8BCA3682FFA4D67FFF6CFC16C7E62B61" blockId="10.[151,1437,150,1977]" pageId="10" pageNumber="354">
<emphasis id="B901EA90FFA4D67FFF6CFC16C3C92B28" box="[199,252,1015,1040]" italics="true" pageId="10" pageNumber="354">Legs</emphasis>
. Protibia (
<figureCitation id="134E2A07FFA4D67FFED8FC16C2FD2B29" box="[371,456,1015,1041]" captionStart="FIGURE 3" captionStartId="5.[151,250,1908,1932]" captionTargetBox="[225,1363,187,1879]" captionTargetId="figure-17@5.[219,1368,181,1885]" captionTargetPageId="5" captionText="FIGURE 3. Male fore-head (AB, EF, IJ, MN), pronotum (C, G, K, O), protibia and apical part of profemur (D, H, L, P) of Neolucanus species. AD, N. lehmanni Baba, 1995 (Imitation of Baba, 1995); EH, N. horizontalis Xin, Zhong &amp; Qi, new species, holotype (Lvchun, Yunnan); IL, N. fuscus Didier, 1926 (Ziyuan, Guangxi); MP, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, CD, E, GH, I, KL, M, OP, dorsal view; B, F, J, N lateral view." figureDoi="http://doi.org/10.5281/zenodo.14756551" httpUri="https://zenodo.org/record/14756551/files/figure.png" pageId="10" pageNumber="354">Fig. 3L</figureCitation>
) with 23 conspicuous weakly sharp protuberances along outer margin, apex bifurcate with branches weakly sharp at tip, and base with a gap. Profemur (
<figureCitation id="134E2A07FFA4D67FFC2BFBFAC0E12B0D" box="[896,980,1051,1077]" captionStart="FIGURE 3" captionStartId="5.[151,250,1908,1932]" captionTargetBox="[225,1363,187,1879]" captionTargetId="figure-17@5.[219,1368,181,1885]" captionTargetPageId="5" captionText="FIGURE 3. Male fore-head (AB, EF, IJ, MN), pronotum (C, G, K, O), protibia and apical part of profemur (D, H, L, P) of Neolucanus species. AD, N. lehmanni Baba, 1995 (Imitation of Baba, 1995); EH, N. horizontalis Xin, Zhong &amp; Qi, new species, holotype (Lvchun, Yunnan); IL, N. fuscus Didier, 1926 (Ziyuan, Guangxi); MP, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, CD, E, GH, I, KL, M, OP, dorsal view; B, F, J, N lateral view." figureDoi="http://doi.org/10.5281/zenodo.14756551" httpUri="https://zenodo.org/record/14756551/files/figure.png" pageId="10" pageNumber="354">Fig. 3L</figureCitation>
) with a semi-circular protrusion at apex. Mesotibia and metatibia straight and outer margin with a terminal spine, without external spines.
</paragraph>
<paragraph id="8BCA3682FFA4D67FFF6CFB82C2692A41" blockId="10.[151,1437,150,1977]" pageId="10" pageNumber="354">
<emphasis id="B901EA90FFA4D67FFF6CFB82C2D02B45" box="[199,485,1123,1149]" italics="true" pageId="10" pageNumber="354">Abdomen and aedeagus.</emphasis>
Abdominal tergite VIII (
<figureCitation id="134E2A07FFA4D67FFCB8FB82C0512B45" box="[787,868,1122,1149]" captionStart="FIGURE 5" captionStartId="7.[151,250,1799,1823]" captionTargetBox="[157,1434,185,1769]" captionTargetId="figure-17@7.[151,1436,181,1775]" captionTargetPageId="7" captionText="FIGURE 5. Male abdominal tergite VIII (A, E, I, M), abdominal ventrite VIII (B, F, J, N) and abdominal segment IX (CD, G H, KL, OP) of Neolucanus species. AH, N. horizontalis Xin, Zhong &amp; Qi, new species AD, holotype (Lvchun, Yunnan); EH, paratype (Jinping, Yunnan). IL, N. fuscus Didier, 1926 (Ziyuan, Guangxi). MP, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, C, E, G, I, K, M, O, dorsal view; B, D, F, H, J, L, N, P, ventral view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14756558" httpUri="https://zenodo.org/record/14756558/files/figure.png" pageId="10" pageNumber="354">Fig. 5I</figureCitation>
) nearly semilune, lateral angles poorly defined. Abdominal ventrite VIII (
<figureCitation id="134E2A07FFA4D67FFE6AFB66C1272B99" box="[449,530,1158,1185]" captionStart="FIGURE 5" captionStartId="7.[151,250,1799,1823]" captionTargetBox="[157,1434,185,1769]" captionTargetId="figure-17@7.[151,1436,181,1775]" captionTargetPageId="7" captionText="FIGURE 5. Male abdominal tergite VIII (A, E, I, M), abdominal ventrite VIII (B, F, J, N) and abdominal segment IX (CD, G H, KL, OP) of Neolucanus species. AH, N. horizontalis Xin, Zhong &amp; Qi, new species AD, holotype (Lvchun, Yunnan); EH, paratype (Jinping, Yunnan). IL, N. fuscus Didier, 1926 (Ziyuan, Guangxi). MP, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, C, E, G, I, K, M, O, dorsal view; B, D, F, H, J, L, N, P, ventral view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14756558" httpUri="https://zenodo.org/record/14756558/files/figure.png" pageId="10" pageNumber="354">Fig. 5J</figureCitation>
) with 2 obvious protruding tips on basal margin near middle. Ventral plate of abdominal segment IX (
<figureCitation id="134E2A07FFA4D67FFE0FFB4AC2CD2BFD" box="[420,504,1194,1221]" captionStart="FIGURE 5" captionStartId="7.[151,250,1799,1823]" captionTargetBox="[157,1434,185,1769]" captionTargetId="figure-17@7.[151,1436,181,1775]" captionTargetPageId="7" captionText="FIGURE 5. Male abdominal tergite VIII (A, E, I, M), abdominal ventrite VIII (B, F, J, N) and abdominal segment IX (CD, G H, KL, OP) of Neolucanus species. AH, N. horizontalis Xin, Zhong &amp; Qi, new species AD, holotype (Lvchun, Yunnan); EH, paratype (Jinping, Yunnan). IL, N. fuscus Didier, 1926 (Ziyuan, Guangxi). MP, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, C, E, G, I, K, M, O, dorsal view; B, D, F, H, J, L, N, P, ventral view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14756558" httpUri="https://zenodo.org/record/14756558/files/figure.png" pageId="10" pageNumber="354">Fig. 5L</figureCitation>
) wide without membranous stripe, an ambiguous protrusion on both side near base, basal 2/3 almost equally wide, then widens towards apex, and gradually constricted near apex. Aedeagus (
<figureCitation id="134E2A07FFA4D67FFAEBFB2EC3E72A35" captionStart="FIGURE 6" captionStartId="8.[151,250,1909,1933]" captionTargetBox="[180,1408,181,1879]" captionTargetId="figure-18@8.[174,1414,181,1885]" captionTargetPageId="8" captionText="FIGURE 6. Aedeagus (not including flagellum) of Neolucanus species. AB, EF, IJ, N. horizontalis Xin, Zhong &amp; Qi, new species A, E, I, holotype (Lvchun, Yunnan); B, F, J, paratype (Jinping, Yunnan). C, G, K, N. fuscus Didier, 1926 (Ziyuan, Guangxi). D, H, L, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). AD, dorsal view; EH, ventral view; IL, lateral view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14735143" httpUri="https://zenodo.org/record/14735143/files/figure.png" pageId="10" pageNumber="354">Fig. 6C, G, K</figureCitation>
) with a large triangular pair of sclerotized dorsal plates and almost intersecting; ventral plate at apical end of basal piece with a sclerotized protruded part. Paramere wide, with relatively large inner curvature at base, apex of paramere curved outward. Penis about 0.65 times length of paramere in ventral view. Flagellum long, apex of flagellum simple.
</paragraph>
<paragraph id="8BCA3682FFA4D67FFF6CFA63C0892931" blockId="10.[151,1437,150,1977]" pageId="10" pageNumber="354">
<emphasis id="B901EA90FFA4D67FFF6CFA63C2F62AA5" bold="true" box="[199,451,1410,1437]" pageId="10" pageNumber="354">Variation. Mesodont.</emphasis>
Mandible only weakly longer than head, with a bifurcated sharp large tooth at base and continuous small teeth, the largest one at about proximal 1/3.
<emphasis id="B901EA90FFA4D67FFCCEFA47C0C92AF8" bold="true" box="[869,1020,1446,1472]" pageId="10" pageNumber="354">Amphiodont</emphasis>
and
<emphasis id="B901EA90FFA4D67FFB93FA47C7952AF8" bold="true" box="[1080,1184,1446,1472]" pageId="10" pageNumber="354">priodont</emphasis>
(
<figureCitation id="134E2A07FFA4D67FFB19FA46C60E2AF9" box="[1202,1339,1447,1473]" captionStart="FIGURE 2" captionStartId="4.[151,250,1771,1795]" captionTargetBox="[158,1429,186,1740]" captionTargetId="figure-18@4.[151,1436,181,1747]" captionTargetPageId="4" captionText="FIGURE 2. Male habitus of Neolucanus species. AB, N. horizontalis Xin, Zhong &amp; Qi, new species, paratype, 33.7 mm, mesodont (Jinping, Yunnan); CD, N. fuscus Didier, 1926, 34.7mm, priodont (Wuming, Guangxi); EF, N. brochieri Bomans &amp; Miyashita, 1997, 31.4 mm, priodont (Gonshan, Yunnan). A, C, E, dorsal view; B, D, F, ventral view. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14735139" httpUri="https://zenodo.org/record/14735139/files/figure.png" pageId="10" pageNumber="354">Figs. 2CD</figureCitation>
;
<figureCitation id="134E2A07FFA4D67FFAE2FA46C6B82AF9" box="[1353,1421,1447,1473]" captionStart="FIGURE 8" captionStartId="11.[151,250,1874,1898]" captionTargetBox="[238,1361,182,1847]" captionTargetId="figure-17@11.[226,1361,181,1850]" captionTargetPageId="11" captionText="FIGURE 8. An amphiodont male of Neolucanus fuscus Didier, 1926 (36.5 mm, Wuming, Guangxi). AB, habitus; C, abdominal tergite VIII; D, abdominal ventrite VIII; FG, abdominal segment IX; E, HJ, aedeagus. A, C, F, H, dorsal view; B, DE, G, J, ventral view; I, lateral view. Scale bar for AB = 10.0 mm, for CJ = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14735148" httpUri="https://zenodo.org/record/14735148/files/figure.png" pageId="10" pageNumber="354">8AB</figureCitation>
). Mandible weakly shorter than head with unclear consecutive small teeth on inner side, without branch at apex in lateral view; clypeolabrum (
<figureCitation id="134E2A07FFA4D67FFE78FA0EC1012931" box="[467,564,1519,1545]" captionStart="FIGURE 4" captionStartId="6.[151,250,1582,1606]" captionTargetBox="[158,1429,188,1551]" captionTargetId="figure-18@6.[151,1436,181,1558]" captionTargetPageId="6" captionText="FIGURE 4. Male fore-head (AB, EF, IJ), pronotum (C, G, K), protibia and apical part of profemur (D, H, L) of Neolucanus species.AD, N. horizontalis Xin, Zhong &amp; Qi, new species, paratype (Jinping, Yunnan); EH, N. fuscus Didier, 1926 (Wuming, Guangxi); IL, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, CD, E, GH, I, KL, dorsal view; B, F, J, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14735141" httpUri="https://zenodo.org/record/14735141/files/figure.png" pageId="10" pageNumber="354">Figs. 4E</figureCitation>
;
<figureCitation id="134E2A07FFA4D67FFDEBFA0EC1562931" box="[576,611,1519,1545]" captionStart="FIGURE 8" captionStartId="11.[151,250,1874,1898]" captionTargetBox="[238,1361,182,1847]" captionTargetId="figure-17@11.[226,1361,181,1850]" captionTargetPageId="11" captionText="FIGURE 8. An amphiodont male of Neolucanus fuscus Didier, 1926 (36.5 mm, Wuming, Guangxi). AB, habitus; C, abdominal tergite VIII; D, abdominal ventrite VIII; FG, abdominal segment IX; E, HJ, aedeagus. A, C, F, H, dorsal view; B, DE, G, J, ventral view; I, lateral view. Scale bar for AB = 10.0 mm, for CJ = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14735148" httpUri="https://zenodo.org/record/14735148/files/figure.png" pageId="10" pageNumber="354">8A</figureCitation>
) clearly visible in dorsal view.
</paragraph>
</subSubSection>
<subSubSection id="C36F6509FFA4D67FFF6CF9F3C1972915" box="[199,674,1554,1581]" pageId="10" pageNumber="354" type="diagnosis">
<paragraph id="8BCA3682FFA4D67FFF6CF9F3C1972915" blockId="10.[151,1437,150,1977]" box="[199,674,1554,1581]" pageId="10" pageNumber="354">
<emphasis id="B901EA90FFA4D67FFF6CF9F3C2FB2914" bold="true" box="[199,462,1554,1580]" pageId="10" pageNumber="354">Differential diagnosis.</emphasis>
See
<tableCitation id="C6F70339FFA4D67FFDA8F9F2C1632915" box="[515,598,1555,1581]" captionStart="TABLE 1" captionStartId="14.[151,239,728,752]" captionTargetPageId="14" captionText="TABLE 1. Male main differences of external characters of Neolucanus horizontalis Xin, Zhong &amp; Qi, new species and its related species." httpUri="http://table.plazi.org/id/DF0A660AFFA0D67BFF3CFD39C26E2C2C" pageId="10" pageNumber="354" tableUuid="DF0A660AFFA0D67BFF3CFD39C26E2C2C">Table 1</tableCitation>
and
<tableCitation id="C6F70339FFA4D67FFD26F9F2C1AB2915" box="[653,670,1555,1581]" captionStart="TABLE 2" captionStartId="15.[151,239,151,175]" captionTargetPageId="15" captionText="TABLE 2. Main differences of male abdomen and aedeagus of Neolucanus horizontalis Xin, Zhong &amp; Qi, new species and its related species." httpUri="http://table.plazi.org/id/DF0A660AFFA1D67AFF3CFF76C2BD2FEB" pageId="10" pageNumber="354" tableUuid="DF0A660AFFA1D67AFF3CFF76C2BD2FEB">2</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F6509FFA4D67FFF6CF9D7C73C294D" pageId="10" pageNumber="354" type="distribution">
<paragraph id="8BCA3682FFA4D67FFF6CF9D7C73C294D" blockId="10.[151,1437,150,1977]" pageId="10" pageNumber="354">
<emphasis id="B901EA90FFA4D67FFF6CF9D7C2542968" bold="true" box="[199,353,1590,1616]" pageId="10" pageNumber="354">Distribution.</emphasis>
<collectingCountry id="F3627612FFA4D67FFEC2F9D6C2992969" box="[361,428,1591,1617]" name="China" pageId="10" pageNumber="354">China</collectingCountry>
(
<collectingRegion id="49B1F860FFA4D67FFE16F9D6C1172969" box="[445,546,1591,1617]" country="China" name="Guizhou" pageId="10" pageNumber="354">Guizhou</collectingRegion>
,
<collectingRegion id="49B1F860FFA4D67FFD85F9D6C1A42969" box="[558,657,1591,1617]" country="China" name="Guangxi" pageId="10" pageNumber="354">Guangxi</collectingRegion>
,
<collectingRegion id="49B1F860FFA4D67FFD35F9D6C1CC2969" box="[670,761,1591,1617]" country="China" name="Yunnan" pageId="10" pageNumber="354">Yunnan</collectingRegion>
);
<collectingCountry id="F3627612FFA4D67FFCA7F9D6C05E2969" box="[780,875,1591,1617]" name="Vietnam" pageId="10" pageNumber="354">Vietnam</collectingCountry>
(
<collectingRegion id="49B1F860FFA4D67FFCD7F9D6C0E92969" box="[892,988,1591,1617]" country="Vietnam" name="Lao Cai" pageId="10" pageNumber="354">Lào Cai</collectingRegion>
,
<collectingRegion id="49B1F860FFA4D67FFC43F9D6C74A2969" box="[1000,1151,1591,1617]" country="Vietnam" name="Thai Nguyen" pageId="10" pageNumber="354">Thái Nguyên</collectingRegion>
,
<collectingRegion id="49B1F860FFA4D67FFB27F9D6C61D2969" box="[1164,1320,1591,1617]" country="Vietnam" name="Tuyen Quang" pageId="10" pageNumber="354">Tuyên Quang</collectingRegion>
,
<collectingRegion id="49B1F860FFA4D67FFA9EF9D6C6A12969" box="[1333,1428,1591,1617]" country="Vietnam" name="Yen Bai" pageId="10" pageNumber="354">Yên Bái</collectingRegion>
) (
<bibRefCitation id="EFE44B73FFA4D67FFF35F9BAC210294D" author="Fujita, H." box="[158,293,1627,1653]" pageId="10" pageNumber="354" refId="ref7425" refString="Fujita, H. (2010) The Lucanid Beetles of the World. Mushi-Sha, Tokyo, 472 pp., 248 pls." type="book" year="2010">Fujita 2010</bibRefCitation>
;
<bibRefCitation id="EFE44B73FFA4D67FFE9BF9BAC2F1294D" author="Schenk, K. D." box="[304,452,1627,1653]" pageId="10" pageNumber="354" pagination="4 - 38" refId="ref7844" refString="Schenk, K. D. (2012) Notes to the Lucanidae of Asia and description of new taxa of the genus Neolucanus. Beetles World, 6, 4 - 38." type="journal article" year="2012">Schenk 2012</bibRefCitation>
,
<bibRefCitation id="EFE44B73FFA4D67FFE64F9BAC13E294D" author="Schenk, K. D." box="[463,523,1627,1653]" pageId="10" pageNumber="354" pagination="2 - 8" refId="ref7878" refString="Schenk, K. D. (2014) Catalogue of Lucanidae: Genus Neolucanus Thomson, 1862. Beetles World, 10, 2 - 8." type="journal article" year="2014">2014</bibRefCitation>
;
<bibRefCitation id="EFE44B73FFA4D67FFDBEF9BAC1F2294D" author="Toshiyuki, T." box="[533,711,1627,1653]" pageId="10" pageNumber="354" pagination="1 - 87" refId="ref7955" refString="Toshiyuki, T. (2021) Illustrated guide to the genus Neolucanus of the world. Be-kuwa, 78, 1 - 87." type="journal article" year="2021">Toshiyuki 2021</bibRefCitation>
;
<bibRefCitation id="EFE44B73FFA4D67FFD79F9BAC01E294D" author="Yi, D." box="[722,811,1627,1653]" pageId="10" pageNumber="354" refId="ref7980" refString="Yi, D. (2023) Lucanidae of the World. Design Spacetime, Seoul, 190 pp., 160 pls." type="book" year="2023">Yi 2023</bibRefCitation>
and present paper).
</paragraph>
</subSubSection>
<subSubSection id="C36F6509FFA4D67FFF6CF99EC07B2881" pageId="10" pageNumber="354" type="discussion">
<paragraph id="8BCA3682FFA4D67FFF6CF99EC07B2985" blockId="10.[151,1437,150,1977]" pageId="10" pageNumber="354">
<emphasis id="B901EA90FFA4D67FFF6CF99EC20E29A1" bold="true" box="[199,315,1663,1689]" pageId="10" pageNumber="354">Remarks.</emphasis>
A notable feature observed is a gap at the base of the protibia, which can be folded to fit snugly into a semi-circular protrusion located at the apex of the profemur.
</paragraph>
<paragraph id="8BCA3682FFA4D67FFF6CF926C07B2881" blockId="10.[151,1437,150,1977]" pageId="10" pageNumber="354">
In our examination of telodont males of
<taxonomicName id="4C754D01FFA4D67FFD30F926C03129D9" authorityName="Didier" authorityYear="1926" box="[667,772,1735,1761]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="354" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="B901EA90FFA4D67FFD30F926C03129D9" box="[667,772,1735,1761]" italics="true" pageId="10" pageNumber="354">N. fuscus</emphasis>
</taxonomicName>
from northern
<collectingCountry id="F3627612FFA4D67FFC1EF926C72129D9" box="[949,1044,1735,1761]" name="Vietnam" pageId="10" pageNumber="354">Vietnam</collectingCountry>
and southern
<collectingCountry id="F3627612FFA4D67FFB13F926C7CB29D9" box="[1208,1278,1735,1761]" name="China" pageId="10" pageNumber="354">China</collectingCountry>
, we found no discernible differences between the two populations in terms of external features or male genitalia. Furthermore, we inspected an ampriodont male (
<figureCitation id="134E2A07FFA4D67FFD97F8EEC1872811" box="[572,690,1807,1833]" captionStart="FIGURE 8" captionStartId="11.[151,250,1874,1898]" captionTargetBox="[238,1361,182,1847]" captionTargetId="figure-17@11.[226,1361,181,1850]" captionTargetPageId="11" captionText="FIGURE 8. An amphiodont male of Neolucanus fuscus Didier, 1926 (36.5 mm, Wuming, Guangxi). AB, habitus; C, abdominal tergite VIII; D, abdominal ventrite VIII; FG, abdominal segment IX; E, HJ, aedeagus. A, C, F, H, dorsal view; B, DE, G, J, ventral view; I, lateral view. Scale bar for AB = 10.0 mm, for CJ = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14735148" httpUri="https://zenodo.org/record/14735148/files/figure.png" pageId="10" pageNumber="354">Fig. 8AJ</figureCitation>
) from Nanning,
<collectingRegion id="49B1F860FFA4D67FFCD4F8EEC7672811" box="[895,1106,1807,1833]" country="China" name="Guangxi" pageId="10" pageNumber="354">Guangxi Province</collectingRegion>
,
<collectingCountry id="F3627612FFA4D67FFBC8F8EEC79C2811" box="[1123,1193,1807,1833]" name="China" pageId="10" pageNumber="354">China</collectingCountry>
, which exhibited no external distinctions from the
<typeStatus id="54CE8820FFA4D67FFE4EF8D2C1722875" box="[485,583,1843,1869]" pageId="10" pageNumber="354" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C754D01FFA4D67FFDC0F8D2C0702875" authorityName="Schenk" authorityYear="2012" box="[619,837,1843,1869]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="354" phylum="Arthropoda" rank="subSpecies" species="fuscus" subSpecies="zhongguo">
<emphasis id="B901EA90FFA4D67FFDC0F8D2C14B2874" box="[619,638,1843,1868]" italics="true" pageId="10" pageNumber="354">N</emphasis>
.
<emphasis id="B901EA90FFA4D67FFD27F8D2C0702875" box="[652,837,1843,1869]" italics="true" pageId="10" pageNumber="354">fuscus zhongguo</emphasis>
</taxonomicName>
(see fig.
<quantity id="4C8D9B67FFA4D67FFC07F8D2C0E32875" box="[940,982,1843,1869]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="10" pageNumber="354" unit="in" value="1.0">1 in</quantity>
<bibRefCitation id="EFE44B73FFA4D67FFC77F8D2C7472875" author="Schenk, K. D." box="[988,1138,1843,1869]" pageId="10" pageNumber="354" pagination="4 - 38" refId="ref7844" refString="Schenk, K. D. (2012) Notes to the Lucanidae of Asia and description of new taxa of the genus Neolucanus. Beetles World, 6, 4 - 38." type="journal article" year="2012">Schenk 2012</bibRefCitation>
). Besides, the distance and geographical conditions of the interconnected distribution areas do not present any significant isolation that would justify the differentiation of
<taxonomicName id="4C754D01FFA4D67FFE77F89AC17028AD" authorityName="Didier" authorityYear="1926" box="[476,581,1915,1941]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="354" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="B901EA90FFA4D67FFE77F89AC2DA28AC" box="[476,495,1915,1940]" italics="true" pageId="10" pageNumber="354">N</emphasis>
.
<emphasis id="B901EA90FFA4D67FFE54F89AC17028AD" box="[511,581,1915,1941]" italics="true" pageId="10" pageNumber="354">fuscus</emphasis>
</taxonomicName>
into subspecies. Therefore, we propose that
<taxonomicName id="4C754D01FFA4D67FFBEDF89AC61028AD" authorityName="Schenk" authorityYear="2012" box="[1094,1317,1915,1941]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="354" phylum="Arthropoda" rank="subSpecies" species="fuscus" subSpecies="zhongguo">
<emphasis id="B901EA90FFA4D67FFBEDF89AC76C28AC" box="[1094,1113,1915,1940]" italics="true" pageId="10" pageNumber="354">N</emphasis>
.
<emphasis id="B901EA90FFA4D67FFBC2F89AC61028AD" box="[1129,1317,1915,1941]" italics="true" pageId="10" pageNumber="354">fuscus zhongguo</emphasis>
</taxonomicName>
should be considered a new junior synonym of the nominate subspecies.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,90 +1,92 @@
<document id="A0B3E9E496D31C768DF757B200F285D1" ID-DOI="10.11646/zootaxa.5570.2.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="14735135" ID-ZooBank="8B1BAE9E-FBD3-4AC3-9FA3-9CCC2AA81392" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737737634945" checkinUser="plazi" docAuthor="Xin, Fei-Yi, Zhong, Xiao-Tian, Guo, Wen-Chi, Zhang, Yi-Feng, Qi, Zhi-Hao &amp; Wu, Song-Qing" docDate="2025" docId="03DC8794FFACD672FF3CFF76C62A28B3" docLanguage="en" docName="zootaxa.5570.2.6.pdf" docOrigin="Zootaxa 5570 (2)" docSource="https://doi.org/10.11646/zootaxa.5570.2.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Neolucanus horizontalis Xin, Zhong &amp; Qi 2025, new species" docType="treatment" docVersion="2" lastPageNumber="351" masterDocId="FFE5FFECFFAED675FFABFFE1C3352F38" masterDocTitle="A taxonomic note on Chinese species of the Neolucanus fuscus group with the description of a new species (Coleoptera: Lucanidae: Lucaninae)" masterLastPageNumber="360" masterPageNumber="344" pageNumber="346" updateTime="1737738157217" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="D3F2919A7A25002A7110161EB4F83341" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CF0C7CC3B4DEB912CF4135A224949435">
<mods:title id="1D043243A19DC89F55980BD126046443">A taxonomic note on Chinese species of the Neolucanus fuscus group with the description of a new species (Coleoptera: Lucanidae: Lucaninae)</mods:title>
<document id="A613BBE026EED9438881813DAB0C5287" ID-CLB-Dataset="307889" ID-DOI="10.11646/zootaxa.5570.2.6" ID-GBIF-Dataset="2396024e-a9ed-4584-afe7-9fed779d32e1" ID-ISSN="1175-5326" ID-Zenodo-Dep="14735135" ID-ZooBank="8B1BAE9E-FBD3-4AC3-9FA3-9CCC2AA81392" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1737737634945" checkinUser="plazi" docAuthor="Xin, Fei-Yi, Zhong, Xiao-Tian, Guo, Wen-Chi, Zhang, Yi-Feng, Qi, Zhi-Hao &amp; Wu, Song-Qing" docDate="2025" docId="03DC8794FFACD673FF3CFF76C6B0285F" docLanguage="en" docName="zootaxa.5570.2.6.pdf" docOrigin="Zootaxa 5570 (2)" docSource="https://doi.org/10.11646/zootaxa.5570.2.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Neolucanus horizontalis Xin, Zhong &amp; Qi 2025, new species" docType="treatment" docVersion="3" lastPageNumber="350" masterDocId="FFE5FFECFFAED675FFABFFE1C3352F38" masterDocTitle="A taxonomic note on Chinese species of the Neolucanus fuscus group with the description of a new species (Coleoptera: Lucanidae: Lucaninae)" masterLastPageNumber="360" masterPageNumber="344" pageNumber="346" updateTime="1738085397281" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="1C6A4101CFC20180DC8E317354761BDD" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="641164A8C057173337611177124B8D6A">
<mods:title id="8585E02DA17D36F03968A91ACCB1C500">A taxonomic note on Chinese species of the Neolucanus fuscus group with the description of a new species (Coleoptera: Lucanidae: Lucaninae)</mods:title>
</mods:titleInfo>
<mods:name id="144153DA80089BA17F466DF27877CFED" type="personal">
<mods:role id="C064A4F92EE27E6AD4C7A585FC8BEC00">
<mods:roleTerm id="4E1353862E58628933EC3A9151529FA3">Author</mods:roleTerm>
<mods:name id="14D61A93FAC1DD7897B25073AE2A9BBE" type="personal">
<mods:role id="132F49B86339BBA8653B1C508C25053F">
<mods:roleTerm id="1B5CEFDFC329A80C12D43C709DFAFB5A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F39DFA16FEB19493B6B8D7C0CA2A4D2D">Xin, Fei-Yi</mods:namePart>
<mods:nameIdentifier id="197B962391C615B7DB3E439A37CE2F17" type="ORCID">0009-0002-4407-694X</mods:nameIdentifier>
<mods:affiliation id="160898452FCC4A17E2F9C9620D89EFA9">Fujian Colleges and Universities Engineering Research Institute of Conservation and Utilization of Natural Bioresources, Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Key Laboratory of Integrated Pest Management in Ecological Forests, Fujian Province University, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China.</mods:affiliation>
<mods:namePart id="85D3D5DDFAA9EC23074CAE8C12023211">Xin, Fei-Yi</mods:namePart>
<mods:nameIdentifier id="70C46814A906F888B38224F6D8B9D875" type="ORCID">0009-0002-4407-694X</mods:nameIdentifier>
<mods:affiliation id="D9103795AB1DF8770A624EDA3C42ACA2">Fujian Colleges and Universities Engineering Research Institute of Conservation and Utilization of Natural Bioresources, Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Key Laboratory of Integrated Pest Management in Ecological Forests, Fujian Province University, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China.</mods:affiliation>
</mods:name>
<mods:name id="2805478CE27B121FFC47735E01637B38" type="personal">
<mods:role id="7BADEC5C33863255AD0427A364AE6E17">
<mods:roleTerm id="EE7C9A8CCDEE225B216C0F94185AC887">Author</mods:roleTerm>
<mods:name id="F9326432C907927590EDA768A0C39223" type="personal">
<mods:role id="F369DF47F5B56101CECE9075185BAB6C">
<mods:roleTerm id="64BDCA807563539EC424C4574BBDB02D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F5ECFDA05BB5F91A76BFB1C02EE9C452">Zhong, Xiao-Tian</mods:namePart>
<mods:nameIdentifier id="85C93D45CBE73158D551CCC503EFC0D7" type="ORCID">0009-0003-1423-9274</mods:nameIdentifier>
<mods:affiliation id="12E3EB8E07096F2E32025163CF703BB2">School of Geoscience and Technology, Southwest Petroleum University, Chengdu 610500, China.</mods:affiliation>
<mods:namePart id="DEB1B0F43A0335A12D761A0F96A2DBA6">Zhong, Xiao-Tian</mods:namePart>
<mods:nameIdentifier id="6DA3FC4BC7030D7ECB042E051DA75752" type="ORCID">0009-0003-1423-9274</mods:nameIdentifier>
<mods:affiliation id="4248D2CFF80AE49594E6C0D8114B7782">School of Geoscience and Technology, Southwest Petroleum University, Chengdu 610500, China.</mods:affiliation>
</mods:name>
<mods:name id="D72AF184FA664415DB3FD0EC8199B555" type="personal">
<mods:role id="4CA29A06CCC32C898A1B54B801933222">
<mods:roleTerm id="6C9A2E140170A77C8C0BDA47C078C1F0">Author</mods:roleTerm>
<mods:name id="4CBF7E9B1899CCB8C1A98523481078C8" type="personal">
<mods:role id="1813994DD4A4A5DAC961A42AD80EAE42">
<mods:roleTerm id="7D4A7AE1AC953BAF1B6078C15E6A3A25">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A11B10F848D1550FBE5D9F3BEFA1190D">Guo, Wen-Chi</mods:namePart>
<mods:nameIdentifier id="7E8AEDFBBFBB7F4B8083FF5C8FE3CF0F" type="ORCID">0009-0008-4116-4016</mods:nameIdentifier>
<mods:affiliation id="4BD8547D69524CE176ADE63AFD644922">Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China.</mods:affiliation>
<mods:nameIdentifier id="17D712B3C1432311F263BD37921FAE98" type="email">353600225@qq.com</mods:nameIdentifier>
<mods:namePart id="D53CD5ABC14BD02C10FBDEFA31767B23">Guo, Wen-Chi</mods:namePart>
<mods:nameIdentifier id="28DAFC8EFF020D28C425FE68D08B5288" type="ORCID">0009-0008-4116-4016</mods:nameIdentifier>
<mods:affiliation id="7FB00BE1E844A494D4B3A3ED82C94818">Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China.</mods:affiliation>
<mods:nameIdentifier id="D4A510A884429C05D39F17793C4C5A26" type="email">353600225@qq.com</mods:nameIdentifier>
</mods:name>
<mods:name id="9A76BA655E7A694C1B164E04774A87E6" type="personal">
<mods:role id="690AFE52C0C0F63E7CCA48D5C431D156">
<mods:roleTerm id="14E4023D8DD7EB3C7762FCD8972F5C1E">Author</mods:roleTerm>
<mods:name id="696B55103E355D8BFF94D92C8F9CF95B" type="personal">
<mods:role id="290B2C4FD5B32D113AA1D8B4A74A4925">
<mods:roleTerm id="8A82F360DA128EB6A4D788E9020C3ED3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A3D64AA6B502A84E252F58C6EBE251A3">Zhang, Yi-Feng</mods:namePart>
<mods:nameIdentifier id="7266B4C42A07D055B3B5787C8FCA8E0B" type="ORCID">0000-0003-3264-9472</mods:nameIdentifier>
<mods:affiliation id="F7DCC2462B3009FFD5E5DC07BCDA5077">Fujian Academy of Forestry, Fuzhou 350012, China.</mods:affiliation>
<mods:nameIdentifier id="145A8FD94FC1478EEB2BE3908FC0707C" type="email">zhang_yifeng@foxmail.com</mods:nameIdentifier>
<mods:namePart id="EC637F358D377FE71430A8F6F1D1A2C6">Zhang, Yi-Feng</mods:namePart>
<mods:nameIdentifier id="B2EDDC567480222BB60BBB94AC94484A" type="ORCID">0000-0003-3264-9472</mods:nameIdentifier>
<mods:affiliation id="8A09170E509E9BE0830DEC2A13E809ED">Fujian Academy of Forestry, Fuzhou 350012, China.</mods:affiliation>
<mods:nameIdentifier id="0D1EBE97AF8A8058D24290CB368D397E" type="email">zhang_yifeng@foxmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="89B6009AEEEBE49EB3175308C5E3E1BD" type="personal">
<mods:role id="FFC403D8C94BD670E97D6F91ED58CFB1">
<mods:roleTerm id="7D425DD34155806DF352BFEF77550505">Author</mods:roleTerm>
<mods:name id="BEB6F8ECCDF2062544C28F73F5803BF5" type="personal">
<mods:role id="AA57482707C4AF72773FABF72452FAFD">
<mods:roleTerm id="4317959CA9B1CDC563342B84F2C8AEDB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8E9C076CD9597BDED8A89893EBDBA317">Qi, Zhi-Hao</mods:namePart>
<mods:nameIdentifier id="6C320BD91FFBC715248B6304D3A26786" type="ORCID">0000-0001-8596-0913</mods:nameIdentifier>
<mods:affiliation id="51E0C6DDAE435D76E79BFFADCB89DED0">Key Laboratory of Integrated Pest Management in Ecological Forests, Fujian Province University, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Fujian Academy of Forestry, Fuzhou 350012, China.</mods:affiliation>
<mods:namePart id="11C2B2178FD5777E1DA949A46C0144E0">Qi, Zhi-Hao</mods:namePart>
<mods:nameIdentifier id="8DA83DE99FDF1AEEADB7B22D5DA64FBE" type="ORCID">0000-0001-8596-0913</mods:nameIdentifier>
<mods:affiliation id="0B53D6A8D759D7ED0944928507EA004D">Key Laboratory of Integrated Pest Management in Ecological Forests, Fujian Province University, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Fujian Academy of Forestry, Fuzhou 350012, China.</mods:affiliation>
</mods:name>
<mods:name id="E1E389285CA52F2F06AFB85F9B02EC73" type="personal">
<mods:role id="DD6CF3C15EA9CC8232E1F91515440218">
<mods:roleTerm id="B572CEAD870ADDE864B46BBA034938AB">Author</mods:roleTerm>
<mods:name id="04D77BDA58FF2E2EE2AADC208D521EB0" type="personal">
<mods:role id="1F6856E29E315BCDD62E013337995CD8">
<mods:roleTerm id="3A6975E7B3FAFCCFB293CBE74D18A4DE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7693BA36907AAF86785A22FC52CE3F79">Wu, Song-Qing</mods:namePart>
<mods:nameIdentifier id="5B1BDD88E966ABAB952E37864CE1BEAF" type="ORCID">0000-0002-2763-5378</mods:nameIdentifier>
<mods:affiliation id="DE7FA07C2B5A9D555EF19A11A4BB5BD1">Fujian Colleges and Universities Engineering Research Institute of Conservation and Utilization of Natural Bioresources, Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Key Laboratory of Integrated Pest Management in Ecological Forests, Fujian Province University, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China.</mods:affiliation>
<mods:namePart id="842F94A03537198B0C1AEF9E608E36B3">Wu, Song-Qing</mods:namePart>
<mods:nameIdentifier id="091EC11C85D9D74205C19B0CBB39820D" type="ORCID">0000-0002-2763-5378</mods:nameIdentifier>
<mods:affiliation id="71D58610AD4D6B4B4098A130BF8BAAB2">Fujian Colleges and Universities Engineering Research Institute of Conservation and Utilization of Natural Bioresources, Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Key Laboratory of Integrated Pest Management in Ecological Forests, Fujian Province University, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="85DB91CE18F3F8ED8B69E42CC5E50800">text</mods:typeOfResource>
<mods:relatedItem id="B152D614B77036C65B57A0BCE6F8A133" type="host">
<mods:titleInfo id="B250DF38DB4E8003A204DC7F0C5F38BA">
<mods:title id="130EB74649978BF47AC1CE1555018296">Zootaxa</mods:title>
<mods:typeOfResource id="ADA347311075C6A52D72C67AE8F58E64">text</mods:typeOfResource>
<mods:relatedItem id="166FA7A535DCACD6915A5EA45B95F524" type="host">
<mods:titleInfo id="33C1259AB0F98E58E97CAF0CCCB84D91">
<mods:title id="A2B1917F4BAD8C157F807895EB8DF883">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="73E5DCCA61B4911288190C6CA0AF0A9A">
<mods:date id="A33F8A231DF49A1A9CAE1981DEB7CD58">2025</mods:date>
<mods:detail id="9D3F446B1B2A650EF9DCFED51910BC43" type="pubDate">
<mods:number id="F06AB9F641EF2EF9E4AF580F9EDA8599">2025-01-17</mods:number>
<mods:part id="D0CFAA8587577EA16B79A3595768B043">
<mods:date id="2860F547992BB92C74E945BD9F7808EF">2025</mods:date>
<mods:detail id="046ED4D4179C8573C07D27A8BDDA60EE" type="pubDate">
<mods:number id="4392097AA90C031B7C8471E9FE5DD986">2025-01-17</mods:number>
</mods:detail>
<mods:detail id="9CEB4B729E8C74D32DC86FA48747B1E9" type="volume">
<mods:number id="E43D162FEE8EBEDE6D3100822423CA67">5570</mods:number>
<mods:detail id="9881FC0126202A394A467121476A18F2" type="volume">
<mods:number id="C7A9A735CC512DFEF6AC32A249D8A88F">5570</mods:number>
</mods:detail>
<mods:detail id="A753F9286D7901501F4AC712E174C442" type="issue">
<mods:number id="E18AD9BB7438FA32844B4B137E0D5EC0">2</mods:number>
<mods:detail id="D97584FF4FA961BE30FA493032B6BB06" type="issue">
<mods:number id="E9159F62D4BE0FD9527C368A183F0773">2</mods:number>
</mods:detail>
<mods:extent id="86FB603E912539BD1FF7331F7EFF7AD4" unit="page">
<mods:start id="8CFF7FD4D8DA2FA1FB9501E23E35B465">344</mods:start>
<mods:end id="2BF660355214ACF7583C5445FAB9AE90">360</mods:end>
<mods:extent id="9FBEC90C1E666728F662CE1D81823004" unit="page">
<mods:start id="9CDD0C6D84A4C10215386468167DDB8E">344</mods:start>
<mods:end id="03A493729CD2070293AEC5CFBCB64302">360</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="79A3AB46896FC7DB53FF9725DF689584">
<mods:url id="53ADD3BD8BB0F2DF209DB2550B1351CD">https://doi.org/10.11646/zootaxa.5570.2.6</mods:url>
<mods:location id="26E9A55AD515ED11B8A8BEF3228174D7">
<mods:url id="D44DDB43A77E0964CAB877D4018CF9C2">https://doi.org/10.11646/zootaxa.5570.2.6</mods:url>
</mods:location>
<mods:classification id="321CA525BFEE67DECEAA55FB4350CA34">journal article</mods:classification>
<mods:identifier id="E1E051035CCDBAD65C6C6E90217E5A84" type="DOI">10.11646/zootaxa.5570.2.6</mods:identifier>
<mods:identifier id="57B337F67E9B86DCABCAC30637C356C8" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="C3F0E0FC3609AC4A3F13E7F2E1F30BF2" type="Zenodo-Dep">14735135</mods:identifier>
<mods:identifier id="D021EC5441BC84B4208881A834BE2FEF" type="ZooBank">8B1BAE9E-FBD3-4AC3-9FA3-9CCC2AA81392</mods:identifier>
<mods:classification id="100F576173849174298502472F5CD757">journal article</mods:classification>
<mods:identifier id="2B109E087EB6A82293CD438DE8D62672" type="CLB-Dataset">307889</mods:identifier>
<mods:identifier id="FCE7AAEF17BCB687EEE4D00FA3FC9475" type="DOI">10.11646/zootaxa.5570.2.6</mods:identifier>
<mods:identifier id="B409005F59193E1CD0FF679419752FC5" type="GBIF-Dataset">2396024e-a9ed-4584-afe7-9fed779d32e1</mods:identifier>
<mods:identifier id="1E62B9269344F04C43A17F692593D116" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="84EF1992F8029C95AFF1AF5B96E74C5F" type="Zenodo-Dep">14735135</mods:identifier>
<mods:identifier id="3522B6C6BB20B3B6AADAE579FE975597" type="ZooBank">8B1BAE9E-FBD3-4AC3-9FA3-9CCC2AA81392</mods:identifier>
</mods:mods>
<treatment id="03DC8794FFACD672FF3CFF76C62A28B3" LSID="urn:lsid:plazi:treatment:03DC8794FFACD672FF3CFF76C62A28B3" httpUri="http://treatment.plazi.org/id/03DC8794FFACD672FF3CFF76C62A28B3" lastPageId="7" lastPageNumber="351" pageId="2" pageNumber="346">
<treatment id="03DC8794FFACD673FF3CFF76C6B0285F" ID-DOI="http://doi.org/10.5281/zenodo.14756547" ID-Zenodo-Dep="14756547" LSID="urn:lsid:plazi:treatment:03DC8794FFACD673FF3CFF76C6B0285F" httpUri="http://treatment.plazi.org/id/03DC8794FFACD673FF3CFF76C6B0285F" lastPageId="6" lastPageNumber="350" pageId="2" pageNumber="346" scope_family="Lucanidae" scope_order="Coleoptera" scope_subFamily="Lucaninae">
<subSubSection id="C36F6509FFACD677FF3CFF76C1602FEB" pageId="2" pageNumber="346" type="nomenclature">
<paragraph id="8BCA3682FFACD677FF3CFF76C00C2F89" blockId="2.[151,879,151,250]" box="[151,825,151,178]" pageId="2" pageNumber="346">
<heading id="D08281EEFFACD677FF3CFF76C00C2F89" bold="true" box="[151,825,151,178]" fontSize="11" level="1" pageId="2" pageNumber="346" reason="1">
@ -108,7 +110,11 @@ Chinese common name:
<figureCitation id="134E2A07FFACD677FF3CFF3EC2292FC1" box="[151,284,223,249]" captionStart="FIGURE 1" captionStartId="3.[151,250,1908,1932]" captionTargetBox="[312,1275,185,1877]" captionTargetId="figure-17@3.[306,1281,179,1883]" captionTargetPageId="3" captionText="FIGURE 1. Male habitus of Neolucanus species. AB, N. horizontalis Xin, Zhong &amp; Qi, new species, holotype, 36.0 mm, telodont (Lvchun, Yunnan); CE, N. fuscus Didier, 1926; CD, 50.9 mm, telodont (Yên Bái, Yên Bái); E, 44.8 mm, telodont (Ziyuan, Guangxi); FH, N. brochieri Bomans &amp; Miyashita, 1997; FG, 41.1mm, telodont (Gonshan, Yunnan); H, 41.0 mm, telodont (Gonshan, Yunnan). A, C, E, F, H, dorsal view; B, D, G, ventral view. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14735137" httpUri="https://zenodo.org/record/14735137/files/figure.png" pageId="2" pageNumber="346">Figs. 1AB</figureCitation>
;
<figureCitation id="134E2A07FFACD677FE8CFF3EC2582FC1" box="[295,365,223,249]" captionStart="FIGURE 2" captionStartId="4.[151,250,1771,1795]" captionTargetBox="[158,1429,186,1740]" captionTargetId="figure-18@4.[151,1436,181,1747]" captionTargetPageId="4" captionText="FIGURE 2. Male habitus of Neolucanus species. AB, N. horizontalis Xin, Zhong &amp; Qi, new species, paratype, 33.7 mm, mesodont (Jinping, Yunnan); CD, N. fuscus Didier, 1926, 34.7mm, priodont (Wuming, Guangxi); EF, N. brochieri Bomans &amp; Miyashita, 1997, 31.4 mm, priodont (Gonshan, Yunnan). A, C, E, dorsal view; B, D, F, ventral view. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14735139" httpUri="https://zenodo.org/record/14735139/files/figure.png" pageId="2" pageNumber="346">2AB</figureCitation>
; 3EH; 4AD; 5AH; 6AB, EF, IJ; 7AB
;
<figureCitation id="134E2A07FFACD677FED2FF3EC28B2FC1" box="[377,446,223,249]" captionStart="FIGURE 3" captionStartId="5.[151,250,1908,1932]" captionTargetBox="[225,1363,187,1879]" captionTargetId="figure-17@5.[219,1368,181,1885]" captionTargetPageId="5" captionText="FIGURE 3. Male fore-head (AB, EF, IJ, MN), pronotum (C, G, K, O), protibia and apical part of profemur (D, H, L, P) of Neolucanus species. AD, N. lehmanni Baba, 1995 (Imitation of Baba, 1995); EH, N. horizontalis Xin, Zhong &amp; Qi, new species, holotype (Lvchun, Yunnan); IL, N. fuscus Didier, 1926 (Ziyuan, Guangxi); MP, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, CD, E, GH, I, KL, M, OP, dorsal view; B, F, J, N lateral view." figureDoi="http://doi.org/10.5281/zenodo.14756551" httpUri="https://zenodo.org/record/14756551/files/figure.png" pageId="2" pageNumber="346">3EH</figureCitation>
; 4AD;
<figureCitation id="134E2A07FFACD677FDB7FF3EC1512FC2" box="[540,612,223,250]" captionStart="FIGURE 5" captionStartId="7.[151,250,1799,1823]" captionTargetBox="[157,1434,185,1769]" captionTargetId="figure-17@7.[151,1436,181,1775]" captionTargetPageId="7" captionText="FIGURE 5. Male abdominal tergite VIII (A, E, I, M), abdominal ventrite VIII (B, F, J, N) and abdominal segment IX (CD, G H, KL, OP) of Neolucanus species. AH, N. horizontalis Xin, Zhong &amp; Qi, new species AD, holotype (Lvchun, Yunnan); EH, paratype (Jinping, Yunnan). IL, N. fuscus Didier, 1926 (Ziyuan, Guangxi). MP, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, C, E, G, I, K, M, O, dorsal view; B, D, F, H, J, L, N, P, ventral view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14756558" httpUri="https://zenodo.org/record/14756558/files/figure.png" pageId="2" pageNumber="346">5AH</figureCitation>
; 6AB, EF, IJ; 7AB
</paragraph>
</subSubSection>
<subSubSection id="C36F6509FFACD677FF3CFEC6C0492EE9" pageId="2" pageNumber="346" type="materials_examined">
@ -126,7 +132,7 @@ locality.
,
<collectingCounty id="62AB4E0EFFACD677FC90FEC6C0DE2E79" box="[827,1003,295,321]" pageId="2" pageNumber="346">Lvchun County</collectingCounty>
,
<location id="8EAA6059FFACD677FC5DFEC6C7B82E7A" LSID="urn:lsid:plazi:treatment:03DC8794FFACD672FF3CFF76C62A28B3:8EAA6059FFACD677FC5DFEC6C7B82E7A" box="[1014,1165,295,322]" country="China" county="Lvchun County" municipality="Honghe Prefecture" name="Pinghe Town" pageId="2" pageNumber="346" stateProvince="Yunnan">Pinghe Town</location>
<location id="8EAA6059FFACD677FC5DFEC6C7B82E7A" LSID="urn:lsid:plazi:treatment:03DC8794FFACD673FF3CFF76C6B0285F:8EAA6059FFACD677FC5DFEC6C7B82E7A" box="[1014,1165,295,322]" country="China" county="Lvchun County" municipality="Honghe Prefecture" name="Pinghe Town" pageId="2" pageNumber="346" stateProvince="Yunnan">Pinghe Town</location>
</materialsCitation>
.
</paragraph>
@ -148,7 +154,7 @@ male (
,
<collectingCounty id="62AB4E0EFFACD677FF3CFE8EC2792EB1" box="[151,332,367,393]" pageId="2" pageNumber="346">Lvchun County</collectingCounty>
,
<location id="8EAA6059FFACD677FEF0FE8EC2C32EB2" LSID="urn:lsid:plazi:treatment:03DC8794FFACD672FF3CFF76C62A28B3:8EAA6059FFACD677FEF0FE8EC2C32EB2" box="[347,502,367,394]" country="China" county="Lvchun County" latitude="22.819445" longLatPrecision="21" longitude="102.523895" municipality="Honghe Prefecture" name="Pinghe Town" pageId="2" pageNumber="346" stateProvince="Yunnan">Pinghe Town</location>
<location id="8EAA6059FFACD677FEF0FE8EC2C32EB2" LSID="urn:lsid:plazi:treatment:03DC8794FFACD673FF3CFF76C6B0285F:8EAA6059FFACD677FEF0FE8EC2C32EB2" box="[347,502,367,394]" country="China" county="Lvchun County" latitude="22.819445" longLatPrecision="21" longitude="102.523895" municipality="Honghe Prefecture" name="Pinghe Town" pageId="2" pageNumber="346" stateProvince="Yunnan">Pinghe Town</location>
,
<geoCoordinate id="EE415045FFACD677FDAEFE8EC1A42EB1" box="[517,657,367,393]" degrees="22" direction="north" minutes="49" orientation="latitude" pageId="2" pageNumber="346" precision="15" seconds="10" value="22.819445">22°4910”N</geoCoordinate>
,
@ -181,9 +187,9 @@ male (
,
<collectingMunicipality id="6BAEACF8FFACD677FCE5FE72C0C22E96" box="[846,1015,403,430]" pageId="2" pageNumber="346">Jinping county</collectingMunicipality>
,
<location id="8EAA6059FFACD677FBAEFE72C7EB2E95" LSID="urn:lsid:plazi:treatment:03DC8794FFACD672FF3CFF76C62A28B3:8EAA6059FFACD677FBAEFE72C7EB2E95" box="[1029,1246,403,429]" country="China" county="Honghe Prefecture" latitude="22.756666" longLatPrecision="21" longitude="103.43917" municipality="Jinping county" name="Ma'andi Township" pageId="2" pageNumber="346" stateProvince="Yunnan">Maandi Township</location>
<location id="8EAA6059FFACD677FBAEFE72C7EB2E95" LSID="urn:lsid:plazi:treatment:03DC8794FFACD673FF3CFF76C6B0285F:8EAA6059FFACD677FBAEFE72C7EB2E95" box="[1029,1246,403,429]" country="China" county="Honghe Prefecture" latitude="22.756666" longLatPrecision="21" longitude="103.43917" municipality="Jinping county" name="Ma'andi Township" pageId="2" pageNumber="346" stateProvince="Yunnan">Maandi Township</location>
,
<location id="8EAA6059FFACD677FB47FE72C6AD2E95" LSID="urn:lsid:plazi:treatment:03DC8794FFACD672FF3CFF76C62A28B3:8EAA6059FFACD677FB47FE72C6AD2E95" box="[1260,1432,403,429]" country="China" county="Honghe Prefecture" latitude="22.756666" longLatPrecision="21" longitude="103.43917" municipality="Jinping county" name="Sitai Mountain" pageId="2" pageNumber="346" stateProvince="Yunnan">Sitai Mountain</location>
<location id="8EAA6059FFACD677FB47FE72C6AD2E95" LSID="urn:lsid:plazi:treatment:03DC8794FFACD673FF3CFF76C6B0285F:8EAA6059FFACD677FB47FE72C6AD2E95" box="[1260,1432,403,429]" country="China" county="Honghe Prefecture" latitude="22.756666" longLatPrecision="21" longitude="103.43917" municipality="Jinping county" name="Sitai Mountain" pageId="2" pageNumber="346" stateProvince="Yunnan">Sitai Mountain</location>
,
<geoCoordinate id="EE415045FFACD677FF3CFE56C2162EEA" box="[151,291,439,466]" degrees="22" direction="north" minutes="45" orientation="latitude" pageId="2" pageNumber="346" precision="15" seconds="24" value="22.756666">22°4524”N</geoCoordinate>
,
@ -233,11 +239,17 @@ Body length (36.0 mm). Length of particular body parts: head (
</paragraph>
<paragraph id="8BCA3682FFACD677FF6CFD6EC6572CB9" blockId="2.[151,1437,295,1905]" pageId="2" pageNumber="346">
<emphasis id="B901EA90FFACD677FF6CFD6EC2312D91" box="[199,260,655,681]" italics="true" pageId="2" pageNumber="346">Head</emphasis>
transverse. Anterior ridge raised behind mandible and widely concave on the middle. Canthus long and wide, completely surround the eye. Clypeolabrum (Fig. 3E) tiny and subtriangle, visible in dorsal view, fused with frons, apex sharp. Mandible (Fig. 3EF) about 2.2 times longer than head, inner smooth from base to teeth; in lateral view, almost horizontal, a dorsal tooth behind apex forming an approximately vertical right-angle and slightly protrudes beyond upper margin; in dorsal view, slightly curved, base thick, becomes thicker firstly and then thinner from base to teeth, with 2 continuous subtriangle inner teeth behind apex, proximal one smaller, apex of mandible sharp. Antennal club with 3 antennomeres; antennomeres 89 subquadrilateral; antennomere 10 semi-elliptic.
transverse. Anterior ridge raised behind mandible and widely concave on the middle. Canthus long and wide, completely surround the eye. Clypeolabrum (
<figureCitation id="134E2A07FFACD677FD73FD52C0182DF5" box="[728,813,691,717]" captionStart="FIGURE 3" captionStartId="5.[151,250,1908,1932]" captionTargetBox="[225,1363,187,1879]" captionTargetId="figure-17@5.[219,1368,181,1885]" captionTargetPageId="5" captionText="FIGURE 3. Male fore-head (AB, EF, IJ, MN), pronotum (C, G, K, O), protibia and apical part of profemur (D, H, L, P) of Neolucanus species. AD, N. lehmanni Baba, 1995 (Imitation of Baba, 1995); EH, N. horizontalis Xin, Zhong &amp; Qi, new species, holotype (Lvchun, Yunnan); IL, N. fuscus Didier, 1926 (Ziyuan, Guangxi); MP, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, CD, E, GH, I, KL, M, OP, dorsal view; B, F, J, N lateral view." figureDoi="http://doi.org/10.5281/zenodo.14756551" httpUri="https://zenodo.org/record/14756551/files/figure.png" pageId="2" pageNumber="346">Fig. 3E</figureCitation>
) tiny and subtriangle, visible in dorsal view, fused with frons, apex sharp. Mandible (
<figureCitation id="134E2A07FFACD677FE70FD36C1792DC9" box="[475,588,727,753]" captionStart="FIGURE 3" captionStartId="5.[151,250,1908,1932]" captionTargetBox="[225,1363,187,1879]" captionTargetId="figure-17@5.[219,1368,181,1885]" captionTargetPageId="5" captionText="FIGURE 3. Male fore-head (AB, EF, IJ, MN), pronotum (C, G, K, O), protibia and apical part of profemur (D, H, L, P) of Neolucanus species. AD, N. lehmanni Baba, 1995 (Imitation of Baba, 1995); EH, N. horizontalis Xin, Zhong &amp; Qi, new species, holotype (Lvchun, Yunnan); IL, N. fuscus Didier, 1926 (Ziyuan, Guangxi); MP, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, CD, E, GH, I, KL, M, OP, dorsal view; B, F, J, N lateral view." figureDoi="http://doi.org/10.5281/zenodo.14756551" httpUri="https://zenodo.org/record/14756551/files/figure.png" pageId="2" pageNumber="346">Fig. 3EF</figureCitation>
) about 2.2 times longer than head, inner smooth from base to teeth; in lateral view, almost horizontal, a dorsal tooth behind apex forming an approximately vertical right-angle and slightly protrudes beyond upper margin; in dorsal view, slightly curved, base thick, becomes thicker firstly and then thinner from base to teeth, with 2 continuous subtriangle inner teeth behind apex, proximal one smaller, apex of mandible sharp. Antennal club with 3 antennomeres; antennomeres 89 subquadrilateral; antennomere 10 semi-elliptic.
</paragraph>
<paragraph id="8BCA3682FFACD677FF6CFC6DC02B2CD5" blockId="2.[151,1437,295,1905]" pageId="2" pageNumber="346">
<emphasis id="B901EA90FFACD677FF6CFC6DC2032C9D" box="[199,310,908,933]" italics="true" pageId="2" pageNumber="346">Pronotum</emphasis>
(Fig. 3G) transverse, 1.8 times wider than long, markedly longer and wider than head with slightly raised edges. Anterior margin bisinuate, posterior margin slightly bisinuate. Anterior angles rounded and directed forwards, lateral angles obtuse and posterior angles blunt.
(
<figureCitation id="134E2A07FFACD677FEEDFC6AC2AA2C9D" box="[326,415,907,933]" captionStart="FIGURE 3" captionStartId="5.[151,250,1908,1932]" captionTargetBox="[225,1363,187,1879]" captionTargetId="figure-17@5.[219,1368,181,1885]" captionTargetPageId="5" captionText="FIGURE 3. Male fore-head (AB, EF, IJ, MN), pronotum (C, G, K, O), protibia and apical part of profemur (D, H, L, P) of Neolucanus species. AD, N. lehmanni Baba, 1995 (Imitation of Baba, 1995); EH, N. horizontalis Xin, Zhong &amp; Qi, new species, holotype (Lvchun, Yunnan); IL, N. fuscus Didier, 1926 (Ziyuan, Guangxi); MP, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, CD, E, GH, I, KL, M, OP, dorsal view; B, F, J, N lateral view." figureDoi="http://doi.org/10.5281/zenodo.14756551" httpUri="https://zenodo.org/record/14756551/files/figure.png" pageId="2" pageNumber="346">Fig. 3G</figureCitation>
) transverse, 1.8 times wider than long, markedly longer and wider than head with slightly raised edges. Anterior margin bisinuate, posterior margin slightly bisinuate. Anterior angles rounded and directed forwards, lateral angles obtuse and posterior angles blunt.
</paragraph>
<paragraph id="8BCA3682FFACD677FF6CFC16C0932B29" blockId="2.[151,1437,295,1905]" box="[199,934,1015,1042]" pageId="2" pageNumber="346">
<taxonomicName id="4C754D01FFACD677FF6CFC16C2002B29" box="[199,309,1015,1041]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="2" pageNumber="346" phylum="Arthropoda" rank="genus">
@ -247,7 +259,9 @@ subtriangular, punctate, width almost as long as length.
</paragraph>
<paragraph id="8BCA3682FFACD677FF6CFBFDC3EF2B45" blockId="2.[151,1437,295,1905]" pageId="2" pageNumber="346">
<emphasis id="B901EA90FFACD677FF6CFBFDC3C92B0D" box="[199,252,1052,1077]" italics="true" pageId="2" pageNumber="346">Legs</emphasis>
. Protibia (Fig. 3H) with 2 conspicuous sharp protuberances along outer margin, apex bifurcate with branches moderately sharp at tip. Mesotibia and metatibia straight and smooth at outer margin with a terminal spine.
. Protibia (
<figureCitation id="134E2A07FFACD677FED0FBFAC2E22B0D" box="[379,471,1051,1077]" captionStart="FIGURE 3" captionStartId="5.[151,250,1908,1932]" captionTargetBox="[225,1363,187,1879]" captionTargetId="figure-17@5.[219,1368,181,1885]" captionTargetPageId="5" captionText="FIGURE 3. Male fore-head (AB, EF, IJ, MN), pronotum (C, G, K, O), protibia and apical part of profemur (D, H, L, P) of Neolucanus species. AD, N. lehmanni Baba, 1995 (Imitation of Baba, 1995); EH, N. horizontalis Xin, Zhong &amp; Qi, new species, holotype (Lvchun, Yunnan); IL, N. fuscus Didier, 1926 (Ziyuan, Guangxi); MP, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, CD, E, GH, I, KL, M, OP, dorsal view; B, F, J, N lateral view." figureDoi="http://doi.org/10.5281/zenodo.14756551" httpUri="https://zenodo.org/record/14756551/files/figure.png" pageId="2" pageNumber="346">Fig. 3H</figureCitation>
) with 2 conspicuous sharp protuberances along outer margin, apex bifurcate with branches moderately sharp at tip. Mesotibia and metatibia straight and smooth at outer margin with a terminal spine.
</paragraph>
<paragraph id="8BCA3682FFACD677FF6CFB66C62D2B9A" blockId="2.[151,1437,295,1905]" box="[199,1304,1159,1186]" pageId="2" pageNumber="346">
<emphasis id="B901EA90FFACD677FF6CFB66C2382B99" box="[199,269,1159,1185]" italics="true" pageId="2" pageNumber="346">Elytra</emphasis>
@ -255,7 +269,13 @@ elongate, 1.3 times longer than wide, anterior margin significantly narrower tha
</paragraph>
<paragraph id="8BCA3682FFACD677FF6CFB4AC11E2ADD" blockId="2.[151,1437,295,1905]" pageId="2" pageNumber="346">
<emphasis id="B901EA90FFACD677FF6CFB4AC2D12BFD" box="[199,484,1195,1221]" italics="true" pageId="2" pageNumber="346">Abdomen and aedeagus.</emphasis>
Abdominal tergite VIII (Fig. 5A) nearly semilune, lateral angles poorly defined. Abdominal ventrite VIII (Fig. 5B) with 2 slightly protruding tips on basal margin near middle. Ventral plate of abdominal segment IX (Fig. 5D) elongate without membranous stripe, basal part extremely narrow, then suddenly became wider, widens gradually to apex and constricted near apex. Aedeagus (
Abdominal tergite VIII (
<figureCitation id="134E2A07FFACD677FCA5FB4AC05F2BFE" box="[782,874,1195,1222]" captionStart="FIGURE 5" captionStartId="7.[151,250,1799,1823]" captionTargetBox="[157,1434,185,1769]" captionTargetId="figure-17@7.[151,1436,181,1775]" captionTargetPageId="7" captionText="FIGURE 5. Male abdominal tergite VIII (A, E, I, M), abdominal ventrite VIII (B, F, J, N) and abdominal segment IX (CD, G H, KL, OP) of Neolucanus species. AH, N. horizontalis Xin, Zhong &amp; Qi, new species AD, holotype (Lvchun, Yunnan); EH, paratype (Jinping, Yunnan). IL, N. fuscus Didier, 1926 (Ziyuan, Guangxi). MP, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, C, E, G, I, K, M, O, dorsal view; B, D, F, H, J, L, N, P, ventral view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14756558" httpUri="https://zenodo.org/record/14756558/files/figure.png" pageId="2" pageNumber="346">Fig. 5A</figureCitation>
) nearly semilune, lateral angles poorly defined. Abdominal ventrite VIII (
<figureCitation id="134E2A07FFACD677FE6BFB2EC12C2BD2" box="[448,537,1231,1258]" captionStart="FIGURE 5" captionStartId="7.[151,250,1799,1823]" captionTargetBox="[157,1434,185,1769]" captionTargetId="figure-17@7.[151,1436,181,1775]" captionTargetPageId="7" captionText="FIGURE 5. Male abdominal tergite VIII (A, E, I, M), abdominal ventrite VIII (B, F, J, N) and abdominal segment IX (CD, G H, KL, OP) of Neolucanus species. AH, N. horizontalis Xin, Zhong &amp; Qi, new species AD, holotype (Lvchun, Yunnan); EH, paratype (Jinping, Yunnan). IL, N. fuscus Didier, 1926 (Ziyuan, Guangxi). MP, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, C, E, G, I, K, M, O, dorsal view; B, D, F, H, J, L, N, P, ventral view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14756558" httpUri="https://zenodo.org/record/14756558/files/figure.png" pageId="2" pageNumber="346">Fig. 5B</figureCitation>
) with 2 slightly protruding tips on basal margin near middle. Ventral plate of abdominal segment IX (
<figureCitation id="134E2A07FFACD677FE0CFB12C2CA2A36" box="[423,511,1267,1294]" captionStart="FIGURE 5" captionStartId="7.[151,250,1799,1823]" captionTargetBox="[157,1434,185,1769]" captionTargetId="figure-17@7.[151,1436,181,1775]" captionTargetPageId="7" captionText="FIGURE 5. Male abdominal tergite VIII (A, E, I, M), abdominal ventrite VIII (B, F, J, N) and abdominal segment IX (CD, G H, KL, OP) of Neolucanus species. AH, N. horizontalis Xin, Zhong &amp; Qi, new species AD, holotype (Lvchun, Yunnan); EH, paratype (Jinping, Yunnan). IL, N. fuscus Didier, 1926 (Ziyuan, Guangxi). MP, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, C, E, G, I, K, M, O, dorsal view; B, D, F, H, J, L, N, P, ventral view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14756558" httpUri="https://zenodo.org/record/14756558/files/figure.png" pageId="2" pageNumber="346">Fig. 5D</figureCitation>
) elongate without membranous stripe, basal part extremely narrow, then suddenly became wider, widens gradually to apex and constricted near apex. Aedeagus (
<figureCitation id="134E2A07FFACD677FBB8FAF6C7912A09" box="[1043,1188,1303,1330]" captionStart="FIGURE 6" captionStartId="8.[151,250,1909,1933]" captionTargetBox="[180,1408,181,1879]" captionTargetId="figure-18@8.[174,1414,181,1885]" captionTargetPageId="8" captionText="FIGURE 6. Aedeagus (not including flagellum) of Neolucanus species. AB, EF, IJ, N. horizontalis Xin, Zhong &amp; Qi, new species A, E, I, holotype (Lvchun, Yunnan); B, F, J, paratype (Jinping, Yunnan). C, G, K, N. fuscus Didier, 1926 (Ziyuan, Guangxi). D, H, L, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). AD, dorsal view; EH, ventral view; IL, lateral view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14735143" httpUri="https://zenodo.org/record/14735143/files/figure.png" pageId="2" pageNumber="346">Fig. 6A, E, I</figureCitation>
) in dorsal view about 3.2 times longer than wide. Basal piece elongated, weakly longer than parameres in dorsal view, with a significant and relatively slender pair of sclerotized dorsal plates, and almost as wide as paramere; ventral plate at apical end of basal piece membranous. Paramere relatively narrow, with a small inner curvature at base, apex of paramere curved outward. Penis about 0.9 times length of paramere in ventral view. Flagellum long, about 1.6 times longer than aedeagus, apex of flagellum simple.
</paragraph>
@ -306,9 +326,9 @@ Baba,
</bibRefCitation>
</taxonomicName>
in general appearance, but the following set of characters could help to identify precisely (
<tableCitation id="C6F70339FFACD677FEA0F90AC26A283D" box="[267,351,1771,1797]" captionStart="TABLE 1" captionStartId="14.[151,239,728,752]" captionText="TABLE 1. Male main differences of external characters of Neolucanus horizontalis Xin, Zhong &amp; Qi, new species and its related species." pageId="2" pageNumber="346">Table 1</tableCitation>
<tableCitation id="C6F70339FFACD677FEA0F90AC26A283D" box="[267,351,1771,1797]" captionStart="TABLE 1" captionStartId="14.[151,239,728,752]" captionTargetPageId="14" captionText="TABLE 1. Male main differences of external characters of Neolucanus horizontalis Xin, Zhong &amp; Qi, new species and its related species." httpUri="http://table.plazi.org/id/DF0A660AFFA0D67BFF3CFD39C26E2C2C" pageId="2" pageNumber="346" tableUuid="DF0A660AFFA0D67BFF3CFD39C26E2C2C">Table 1</tableCitation>
,
<tableCitation id="C6F70339FFACD677FEC6F90AC2F6283D" box="[365,451,1771,1797]" captionStart="TABLE 2" captionStartId="15.[151,239,151,175]" captionText="TABLE 2. Main differences of male abdomen and aedeagus of Neolucanus horizontalis Xin, Zhong &amp; Qi, new species and its related species." pageId="2" pageNumber="346">Table 2</tableCitation>
<tableCitation id="C6F70339FFACD677FEC6F90AC2F6283D" box="[365,451,1771,1797]" captionStart="TABLE 2" captionStartId="15.[151,239,151,175]" captionTargetPageId="15" captionText="TABLE 2. Main differences of male abdomen and aedeagus of Neolucanus horizontalis Xin, Zhong &amp; Qi, new species and its related species." httpUri="http://table.plazi.org/id/DF0A660AFFA1D67AFF3CFF76C2BD2FEB" pageId="2" pageNumber="346" tableUuid="DF0A660AFFA1D67AFF3CFF76C2BD2FEB">Table 2</tableCitation>
).
</paragraph>
</subSubSection>
@ -318,7 +338,7 @@ in general appearance, but the following set of characters could help to identif
This new species is named after the morphological features of its horizontal mandible when viewed in lateral view.
</paragraph>
</subSubSection>
<subSubSection id="C36F6509FFACD670FF6CF8B6C74D28C1" lastPageId="5" lastPageNumber="349" pageId="2" pageNumber="346" type="distribution">
<subSubSection id="C36F6509FFACD677FF6CF8B6C1142849" box="[199,545,1879,1905]" pageId="2" pageNumber="346" type="distribution">
<paragraph id="8BCA3682FFACD677FF6CF8B6C1142849" blockId="2.[151,1437,295,1905]" box="[199,545,1879,1905]" pageId="2" pageNumber="346">
<emphasis id="B901EA90FFACD677FF6CF8B6C2542849" bold="true" box="[199,353,1879,1905]" pageId="2" pageNumber="346">Distribution.</emphasis>
<collectingCountry id="F3627612FFACD677FECCF8B6C29F2849" box="[359,426,1879,1905]" name="China" pageId="2" pageNumber="346">China</collectingCountry>
@ -326,6 +346,7 @@ This new species is named after the morphological features of its horizontal man
<collectingRegion id="49B1F860FFACD677FE12F8B6C1202849" box="[441,533,1879,1905]" country="China" name="Yunnan" pageId="2" pageNumber="346">Yunnan</collectingRegion>
).
</paragraph>
</subSubSection>
<caption id="DF0A660AFFADD676FF3CF895C7AC28C0" ID-DOI="http://doi.org/10.5281/zenodo.14735137" ID-Zenodo-Dep="14735137" httpUri="https://zenodo.org/record/14735137/files/figure.png" pageId="3" pageNumber="347" startId="3.[151,250,1908,1932]" targetBox="[312,1275,185,1877]" targetPageId="3" targetType="figure">
<paragraph id="8BCA3682FFADD676FF3CF895C7AC28C0" blockId="3.[151,1436,1908,2041]" pageId="3" pageNumber="347">
<emphasis id="B901EA90FFADD676FF3CF895C22028B4" bold="true" box="[151,277,1908,1932]" pageId="3" pageNumber="347">FIGURE 1.</emphasis>
@ -420,6 +441,7 @@ dorsal view;
ventral view. Scale bar = 10.0 mm.
</paragraph>
</caption>
<caption id="DF0A660AFFABD670FF3CF895C74D28C1" ID-DOI="http://doi.org/10.5281/zenodo.14756551" ID-Zenodo-Dep="14756551" httpUri="https://zenodo.org/record/14756551/files/figure.png" pageId="5" pageNumber="349" startId="5.[151,250,1908,1932]" targetBox="[225,1363,187,1879]" targetPageId="5">
<paragraph id="8BCA3682FFABD670FF3CF895C74D28C1" blockId="5.[151,1437,1908,2041]" pageId="5" pageNumber="349">
<emphasis id="B901EA90FFABD670FF3CF895C22628B4" bold="true" box="[151,275,1908,1932]" pageId="5" pageNumber="349">FIGURE 3.</emphasis>
Male fore-head
@ -492,7 +514,7 @@ dorsal view;
<emphasis id="B901EA90FFABD670FC39F800C0C028C1" bold="true" box="[914,1013,2017,2041]" pageId="5" pageNumber="349">B, F, J, N</emphasis>
lateral view.
</paragraph>
</subSubSection>
</caption>
<caption id="DF0A660AFFA8D673FF3CF9CFC221298A" ID-DOI="http://doi.org/10.5281/zenodo.14735141" ID-Zenodo-Dep="14735141" httpUri="https://zenodo.org/record/14735141/files/figure.png" pageId="6" pageNumber="350" startId="6.[151,250,1582,1606]" targetBox="[158,1429,188,1551]" targetPageId="6" targetType="figure">
<paragraph id="8BCA3682FFA8D673FF3CF9CFC221298A" blockId="6.[151,1437,1582,1714]" pageId="6" pageNumber="350">
<emphasis id="B901EA90FFA8D673FF3CF9CFC227297E" bold="true" box="[151,274,1582,1606]" pageId="6" pageNumber="350">FIGURE 4.</emphasis>
@ -551,7 +573,7 @@ lateral view.
This species can be attracted to artificial light.
</paragraph>
</subSubSection>
<subSubSection id="C36F6509FFA8D672FF6CF8C8C62A28B3" lastPageId="7" lastPageNumber="351" pageId="6" pageNumber="350" type="discussion">
<subSubSection id="C36F6509FFA8D673FF6CF8C8C6B0285F" pageId="6" pageNumber="350" type="discussion">
<paragraph id="8BCA3682FFA8D673FF6CF8C8C6B0285F" blockId="6.[151,1436,1796,1895]" pageId="6" pageNumber="350">
<emphasis id="B901EA90FFA8D673FF6CF8C8C20E287B" bold="true" box="[199,315,1833,1859]" pageId="6" pageNumber="350">Remarks.</emphasis>
Many species within the
@ -562,66 +584,6 @@ Many species within the
</taxonomicName>
group exhibit individuals with elytra featuring prominent yellowish areas. Therefore, we hypothesize that similar individuals will also be found within the newly discovered species.
</paragraph>
<paragraph id="8BCA3682FFA9D672FF3CF8E6C62A28B3" blockId="7.[151,1437,1799,1931]" pageId="7" pageNumber="351">
<emphasis id="B901EA90FFA9D672FF3CF8E6C2272827" bold="true" box="[151,274,1799,1823]" pageId="7" pageNumber="351">FIGURE 5.</emphasis>
Male abdominal tergite VIII (
<emphasis id="B901EA90FFA9D672FD94F8E6C1932827" bold="true" box="[575,678,1799,1823]" pageId="7" pageNumber="351">A, E, I, M</emphasis>
), abdominal ventrite VIII (
<emphasis id="B901EA90FFA9D672FC1DF8E6C7202827" bold="true" box="[950,1045,1799,1823]" pageId="7" pageNumber="351">B, F, J, N</emphasis>
) and abdominal segment IX (
<emphasis id="B901EA90FFA9D672FA94F8E6C213287B" bold="true" pageId="7" pageNumber="351">CD, G H, KL, OP</emphasis>
) of
<taxonomicName id="4C754D01FFA9D672FEFBF8CAC103287B" box="[336,566,1835,1859]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="351" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="B901EA90FFA9D672FEFBF8CAC2FD287B" box="[336,456,1835,1859]" italics="true" pageId="7" pageNumber="351">Neolucanus</emphasis>
species.
<emphasis id="B901EA90FFA9D672FD8FF8CAC103287B" bold="true" box="[548,566,1835,1859]" pageId="7" pageNumber="351">A</emphasis>
</taxonomicName>
<emphasis id="B901EA90FFA9D672FD9CF8CAC168287B" bold="true" box="[567,605,1835,1859]" pageId="7" pageNumber="351">H,</emphasis>
<taxonomicName id="4C754D01FFA9D672FDCFF8CDC086287B" authority="Xin, Zhong &amp; Qi" authorityName="Xin, Zhong &amp; Qi" authorityYear="2025" box="[612,947,1835,1859]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="351" phylum="Arthropoda" rank="species" species="horizontalis" status="new species">
<emphasis id="B901EA90FFA9D672FDCFF8CDC140287B" box="[612,629,1836,1859]" italics="true" pageId="7" pageNumber="351">N</emphasis>
.
<emphasis id="B901EA90FFA9D672FD29F8CAC1CE287B" box="[642,763,1835,1859]" italics="true" pageId="7" pageNumber="351">horizontalis</emphasis>
Xin, Zhong &amp; Qi
</taxonomicName>
,
<emphasis id="B901EA90FFA9D672FC16F8CAC74D287B" bold="true" box="[957,1144,1835,1859]" pageId="7" pageNumber="351">
<taxonomicNameLabel id="A23257EBFFA9D672FC16F8CAC70F287B" box="[957,1082,1835,1859]" pageId="7" pageNumber="351" rank="species">new species</taxonomicNameLabel>
AD,
</emphasis>
<typeStatus id="54CE8820FFA9D672FBD5F8CAC7E2287B" box="[1150,1239,1835,1859]" pageId="7" pageNumber="351" type="holotype">holotype</typeStatus>
(Lvchun,
<collectingRegion id="49B1F860FFA9D672FA94F8CAC6A4287B" box="[1343,1425,1835,1859]" country="China" name="Yunnan" pageId="7" pageNumber="351">Yunnan</collectingRegion>
);
<emphasis id="B901EA90FFA9D672FF3CF8AEC3FA285F" bold="true" box="[151,207,1871,1895]" pageId="7" pageNumber="351">EH,</emphasis>
<typeStatus id="54CE8820FFA9D672FF7FF8B1C21E2850" box="[212,299,1872,1896]" pageId="7" pageNumber="351" type="paratype">paratype</typeStatus>
(Jinping,
<collectingRegion id="49B1F860FFA9D672FE25F8AEC2D5285F" box="[398,480,1871,1895]" country="China" name="Yunnan" pageId="7" pageNumber="351">Yunnan</collectingRegion>
).
<emphasis id="B901EA90FFA9D672FE44F8AEC128285F" bold="true" box="[495,541,1871,1895]" pageId="7" pageNumber="351">IL,</emphasis>
<taxonomicName id="4C754D01FFA9D672FD89F8B1C034285F" authority="Didier, 1926" authorityName="Didier" authorityYear="1926" box="[546,769,1871,1895]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="351" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="B901EA90FFA9D672FD89F8B1C106285F" box="[546,563,1872,1895]" italics="true" pageId="7" pageNumber="351">N</emphasis>
.
<emphasis id="B901EA90FFA9D672FD95F8AEC148285F" box="[574,637,1871,1895]" italics="true" pageId="7" pageNumber="351">fuscus</emphasis>
<bibRefCitation id="EFE44B73FFA9D672FD29F8AEC034285F" author="Didier, R." box="[642,769,1871,1895]" pageId="7" pageNumber="351" pagination="210 - 214" refId="ref7377" refString="Didier, R. (1926) Descriptions sommaires de Lucanides nouveaux, du genre Neolucanus, de la faune indo-chinoise. Bulletin de la Societe Entomologique de France, 1926, 210 - 214. https: // doi. org / 10.3406 / bsef. 1926.27736" type="journal article" year="1926">Didier, 1926</bibRefCitation>
</taxonomicName>
(Ziyuan,
<collectingRegion id="49B1F860FFA9D672FCCAF8AEC088285F" box="[865,957,1871,1895]" country="China" name="Guangxi" pageId="7" pageNumber="351">Guangxi</collectingRegion>
).
<emphasis id="B901EA90FFA9D672FC67F8AEC731285F" bold="true" box="[972,1028,1871,1895]" pageId="7" pageNumber="351">MP,</emphasis>
<taxonomicName id="4C754D01FFA9D672FBA2F8B1C6A8285F" authority="Bomans &amp; Miyashita, 1997" authorityName="Bomans &amp; Miyashita" authorityYear="1997" box="[1033,1437,1871,1895]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="351" phylum="Arthropoda" rank="species" species="brochieri">
<emphasis id="B901EA90FFA9D672FBA2F8B1C72F285F" box="[1033,1050,1872,1895]" italics="true" pageId="7" pageNumber="351">N</emphasis>
.
<emphasis id="B901EA90FFA9D672FB8EF8AEC7B7285F" box="[1061,1154,1871,1895]" italics="true" pageId="7" pageNumber="351">brochieri</emphasis>
<bibRefCitation id="EFE44B73FFA9D672FB23F8B1C6A8285F" author="Bomans, H. E. &amp; Miyashita, T." box="[1160,1437,1871,1895]" pageId="7" pageNumber="351" pagination="2 - 3" refId="ref7344" refString="Bomans, H. E. &amp; Miyashita, T. (1997) Description de trois nouvelles especes de Lucanidae du nord Birmanie. Besoiro, 4, 2 - 3." type="journal article" year="1997">Bomans &amp; Miyashita, 1997</bibRefCitation>
</taxonomicName>
(Gonshan,
<collectingRegion id="49B1F860FFA9D672FEAEF892C26228B3" box="[261,343,1907,1931]" country="China" name="Yunnan" pageId="7" pageNumber="351">Yunnan</collectingRegion>
).
<emphasis id="B901EA90FFA9D672FEC3F892C16C28B3" bold="true" box="[360,601,1907,1931]" pageId="7" pageNumber="351">A, C, E, G, I, K, M, O,</emphasis>
dorsal view;
<emphasis id="B901EA90FFA9D672FD49F892C0F628B3" bold="true" box="[738,963,1907,1931]" pageId="7" pageNumber="351">B, D, F, H, J, L, N, P,</emphasis>
ventral view. Scale bar = 1.0 mm.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,163 @@
<document id="7AE44F1212C768D2A51118A268ACF5D1" ID-CLB-Dataset="307890" ID-DOI="10.11646/zootaxa.5570.2.7" ID-GBIF-Dataset="80eb3b8b-511c-4927-b6b9-25dd40ec7258" ID-ISSN="1175-5326" ID-Zenodo-Dep="14735158" ID-ZooBank="E6AD6127-22D6-45EE-8667-0E5AE80D3D76" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737737809431" checkinUser="plazi" docAuthor="Tokumori, Daigo, Watanabe, Hiromi Kayama, Tanaka, Katsuhiko, Ohara, Yasuhiko &amp; Tomikawa, Ko" docDate="2025" docId="3F2087EE1D09FFCA57D1F8E9FA62FEED" docLanguage="en" docName="zootaxa.5570.2.7.pdf" docOrigin="Zootaxa 5570 (2)" docSource="https://doi.org/10.11646/zootaxa.5570.2.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Seba Bate 1862" docType="key" docVersion="4" lastPageNumber="369" masterDocId="C319FF961D0EFFC25746FFAFFFFEFFD2" masterDocTitle="A new species of the genus Seba (Crustacea: Amphipoda: Sebidae) from a cold seep of Mariana Trench" masterLastPageNumber="370" masterPageNumber="361" pageNumber="368" updateTime="1738085340846" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="67A57FA82F5A46035EFB680110C03EE8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1C3ACFCA7AEBBD7D702A8FBA57630817">
<mods:title id="05EF4B09C4C4B553C8F0AB978897066A">A new species of the genus Seba (Crustacea: Amphipoda: Sebidae) from a cold seep of Mariana Trench</mods:title>
</mods:titleInfo>
<mods:name id="4C0FDADA5F55F88E64BA8BF5D64311E4" type="personal">
<mods:role id="B35E391B1615C770F05DCDBD3E7ADAD1">
<mods:roleTerm id="B1787ADDAD00ADE7638D7AC9AA7BD0D6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5812E0B38C69D985B8ED7601C9275403">Tokumori, Daigo</mods:namePart>
<mods:nameIdentifier id="B0AA0EA6E0F7963D14728DE248EED0BB" type="ORCID">0009-0001-2877-2737</mods:nameIdentifier>
<mods:affiliation id="E1762940D1ED86642C03DF97D371E89B">Graduate School of Humanities and Social Sciences, Hiroshima University, Higashi-Hiroshima 739 - 8524, Japan.</mods:affiliation>
<mods:nameIdentifier id="FCA6D850D22A7FAB9292ABC9736033A7" type="email">da6m6k9@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="AB50DB5D7A76B40B9A19DDD6DB587663" type="personal">
<mods:role id="FD8AAEE4CE0FF2E5DD334D185AB962B7">
<mods:roleTerm id="8996BCD8447F5BC425332F612BC51A9F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AC23B6007638D1864E0B4AE12C79B529">Watanabe, Hiromi Kayama</mods:namePart>
<mods:affiliation id="2BFFFD2ED389D9DDFD4A350B01EFF72F">X-STAR, Japan Agency for Marine-Earth Science and Technology (JAMSTEC), 2 - 15 Natsushima-cho, Yokosuka, Kanagawa 237 - 0061, Japan.</mods:affiliation>
</mods:name>
<mods:name id="BBC680685FC65A6B7E1053451C4A4AC9" type="personal">
<mods:role id="D462F77A04E2D1EBD7C261650F61C181">
<mods:roleTerm id="2B6F4292B5BA412641CDF9483ABD96A9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="98D48ED83FCEF4AA6ADEF60EC9A03860">Tanaka, Katsuhiko</mods:namePart>
<mods:affiliation id="9721FA225F38C583A63B94CCAA1F573B">Department of Marine Biology, School of Marine Science and Technology, Tokai University, 3 - 20 - 1, Orido, Shimizu, Shizuoka, Shizuoka 424 - 8610, Japan.</mods:affiliation>
</mods:name>
<mods:name id="AA89AA3C62123469F641E3A93880DD8A" type="personal">
<mods:role id="E287AF5C6AA0F824172167CD027EF8B3">
<mods:roleTerm id="5A7B94BF29E735132169E191340F9025">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="15E03589BE93992A17F2BDCBF5CD9EC7">Ohara, Yasuhiko</mods:namePart>
<mods:affiliation id="D4D7475D3E4D04BFB936123B4330E502">Hydrographic and Oceanographic Department of Japan, 3 - 1 - 1 Kasumigaseki, Chiyoda-ku, Tokyo 100 - 8932, Japan. &amp; Research Institute for Marine Geodynamics (IMG), Japan Agency for Marine-Earth Science and Technology (JAMSTEC), 2 - 15 Natsushima-cho, Yokosuka, Kanagawa 237 - 0061, Japan. &amp; Nagoya University, Furo-cho, Chikusa-ku, Nagoya 464 - 8602, Japan.</mods:affiliation>
</mods:name>
<mods:name id="862D0F5051DC3DFD3836FA64852952B6" type="personal">
<mods:role id="CC27E12836C769E5156EC9528BD259B8">
<mods:roleTerm id="A19D5C056B39AB1E2C72CC2B584712B3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8D48D2EF437554142F4A487E85D1BF8C">Tomikawa, Ko</mods:namePart>
<mods:affiliation id="F944F2E6AA0A8F103495F8AB0804BF96">Graduate School of Humanities and Social Sciences, Hiroshima University, Higashi-Hiroshima 739 - 8524, Japan.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="3520835D56983C50B20887E861747B39">text</mods:typeOfResource>
<mods:relatedItem id="61141156D8E40B5838EA76A56D1A25EA" type="host">
<mods:titleInfo id="01197082F07581F18E7C5F6E5298767E">
<mods:title id="3F184104CB0AB890468ECB7A43F59F4A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="F35D13DCEE9D1126DAA5A58A3176E08B">
<mods:date id="62F95B06C5C51289546AE0D0B6B30915">2025</mods:date>
<mods:detail id="43DC0DCCFF9FDFFE065118F022131A3C" type="pubDate">
<mods:number id="FEBACCEC36A7798779B0C7679E410D74">2025-01-17</mods:number>
</mods:detail>
<mods:detail id="570F5D809C130F10ADB229DC33516B2A" type="volume">
<mods:number id="99CA84C8B7A93590FEDB1C268BA0FBB7">5570</mods:number>
</mods:detail>
<mods:detail id="3386AA55A613BAB5930AF81926A75C90" type="issue">
<mods:number id="174A2923A23728CAF2370456EA4EACFF">2</mods:number>
</mods:detail>
<mods:extent id="AF35E3A81012C9CD884E1AB7D179D705" unit="page">
<mods:start id="289F81ECB2181DF528EAF614CCD23246">361</mods:start>
<mods:end id="CD85BC202C1C472451D30678C9176008">370</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="C54B375B793EDFF4D580C1FD346948B3">
<mods:url id="0730C56684721266CAFE27F24A5F145D">https://doi.org/10.11646/zootaxa.5570.2.7</mods:url>
</mods:location>
<mods:classification id="80FE4BC3A57A87671CFF9300E4DFD145">journal article</mods:classification>
<mods:identifier id="74D231565E94F5A3D5F606608E023C4A" type="CLB-Dataset">307890</mods:identifier>
<mods:identifier id="D8F9F55E5911AB62537AD9721582C8D3" type="DOI">10.11646/zootaxa.5570.2.7</mods:identifier>
<mods:identifier id="15527DA349A86C6EDF5D042FFA719699" type="GBIF-Dataset">80eb3b8b-511c-4927-b6b9-25dd40ec7258</mods:identifier>
<mods:identifier id="D1ABBD0A17396230C92C565190437D5D" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A8FF8EEE5947A98CFF0B9235E467B777" type="Zenodo-Dep">14735158</mods:identifier>
<mods:identifier id="7D9706F805753F2C5117BBB60FF1FDB6" type="ZooBank">E6AD6127-22D6-45EE-8667-0E5AE80D3D76</mods:identifier>
</mods:mods>
<treatment id="3F2087EE1D09FFCA57D1F8E9FA62FEED" ID-DOI="http://doi.org/10.5281/zenodo.14756586" ID-Zenodo-Dep="14756586" LSID="urn:lsid:plazi:treatment:3F2087EE1D09FFCA57D1F8E9FA62FEED" httpUri="http://treatment.plazi.org/id/3F2087EE1D09FFCA57D1F8E9FA62FEED" lastPageId="8" lastPageNumber="369" pageId="7" pageNumber="368">
<subSubSection id="FF9365731D09FFC557D1F8E9FC59F8B2" box="[151,935,1862,1889]" pageId="7" pageNumber="368" type="nomenclature">
<paragraph id="B73636F81D09FFC557D1F8E9FC59F8B2" blockId="7.[151,935,1862,1889]" box="[151,935,1862,1889]" pageId="7" pageNumber="368">
<heading id="EC7E81941D09FFC557D1F8E9FC59F8B2" bold="true" box="[151,935,1862,1889]" fontSize="11" level="1" pageId="7" pageNumber="368" reason="1">
<emphasis id="85FDEAEA1D09FFC557D1F8E9FC59F8B2" bold="true" box="[151,935,1862,1889]" pageId="7" pageNumber="368">
Key to species of
<taxonomicName id="70894D7B1D09FFC55628F8E9FE57F8B3" ID-CoL="7PZB8" authorityName="Bate" authorityYear="1862" box="[366,425,1862,1889]" class="Malacostraca" family="Sebidae" genus="Seba" kingdom="Animalia" order="Amphipoda" pageId="7" pageNumber="368" phylum="Arthropoda" rank="genus">
<emphasis id="85FDEAEA1D09FFC55628F8E9FE57F8B3" bold="true" box="[366,425,1862,1889]" italics="true" pageId="7" pageNumber="368">Seba</emphasis>
</taxonomicName>
known from depths greater than
<quantity id="70719B1D1D09FFC5540BF8E9FC59F8B2" box="[845,935,1862,1888]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="368" unit="m" value="1000.0">1000 m</quantity>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FF9365731D09FFCA57D1F824FA62FEED" lastPageId="8" lastPageNumber="369" pageId="7" pageNumber="368" type="key">
<key id="C31861C91D09FFCA57D1F824FA62FEED" lastPageId="8" lastPageNumber="369" pageId="7" pageNumber="368">
<keyStep id="0C7D2E5D1D09FFC557D1F824FA62F86C" pageId="7" pageNumber="368">
<paragraph id="B73636F81D09FFC557D1F824FA62F872" blockId="7.[151,1436,1931,2041]" box="[151,1436,1931,1953]" pageId="7" pageNumber="368">
<keyLead id="0C7895CD1D09FFC557D1F824FA62F872" box="[151,1436,1931,1953]" pageId="7" pageNumber="368">1 Gnathopod 2 coxa with a prominent tooth at the posterodistal corner............................................. 2</keyLead>
</paragraph>
<paragraph id="B73636F81D09FFC557D1F806FA62F86C" blockId="7.[151,1436,1931,2041]" box="[151,1436,1961,1983]" pageId="7" pageNumber="368">
<keyLead id="0C7895CD1D09FFC557D1F806FA62F86C" box="[151,1436,1961,1983]" pageId="7" pageNumber="368">- Gnathopod 2 coxa without a prominent tooth at the posterodistal corner.......................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="0C7D2E5D1D09FFCA57D1F869FA62FF18" lastPageId="8" lastPageNumber="369" pageId="7" pageNumber="368">
<paragraph id="B73636F81D09FFC557D1F869FA62F82A" blockId="7.[151,1436,1931,2041]" pageId="7" pageNumber="368">
<keyLead id="0C7895CD1D09FFC557D1F869FA62F82A" pageId="7" pageNumber="368">
2 Mandibular palp article 2 with seta; gnathopod 1 coxa with tooth at the posteroventral corner; uropod 3 proximal article of ramus with dorsal setae.............................................................
<taxonomicName id="70894D7B1D09FFC553F6F84BFAB7F82B" authorityName="Tokumori &amp; Watanabe &amp; Tanaka &amp; Ohara &amp; Tomikawa" authorityYear="2025" box="[1200,1353,2020,2041]" class="Malacostraca" family="Sebidae" genus="Seba" kingdom="Animalia" order="Amphipoda" pageId="7" pageNumber="368" phylum="Arthropoda" rank="species" species="marianaensis" status="sp. nov.">
<emphasis id="85FDEAEA1D09FFC553F6F84BFAB7F82B" box="[1200,1353,2020,2041]" italics="true" pageId="7" pageNumber="368">S. marianaensis</emphasis>
</taxonomicName>
<emphasis id="85FDEAEA1D09FFC55216F84CFA62F82A" bold="true" box="[1360,1436,2019,2040]" pageId="7" pageNumber="368">
<taxonomicNameLabel id="9ECE57911D09FFC55216F84CFA62F82A" box="[1360,1436,2019,2040]" pageId="7" pageNumber="368" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="B73636F81D06FFCA57D1FF37FA62FF18" blockId="8.[151,1436,151,320]" pageId="8" pageNumber="369">
<keyLead id="0C7895CD1D06FFCA57D1FF37FA62FF18" pageId="8" pageNumber="369">
- Mandibular palp article 2 without seta; gnathopod 1 coxa with two teeth at the posteroventral corner; uropod 3 proximal article of ramus without dorsal setae.........................................................
<taxonomicName id="70894D7B1D06FFCA53EBFF1AFA62FF18" ID-CoL="4W77M" authority="Larsen, 2007" authorityName="Larsen" authorityYear="2007" box="[1197,1436,181,202]" class="Malacostraca" family="Sebidae" genus="Seba" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="369" phylum="Arthropoda" rank="species" species="bathybia">
<emphasis id="85FDEAEA1D06FFCA53EBFF1AFAE7FF18" box="[1197,1305,181,202]" italics="true" pageId="8" pageNumber="369">S. bathybia</emphasis>
<bibRefCitation id="D3184B091D06FFCA5259FF1AFA62FF18" author="Larsen, K." box="[1311,1436,181,202]" pageId="8" pageNumber="369" pagination="1 - 26" refId="ref4069" refString="Larsen, K. (2007) Amphipoda (Crustacea; Peracarida) from the hydrothermal vent system of the Juan de Fuca Ridge, Escabana Trough and Gorda Ridge, Northeast Pacific. Zootaxa, 1445 (1), 1 - 26. https: // doi. org / 10.11646 / zootaxa. 1445.1.1" type="journal article" year="2007">Larsen, 2007</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="0C7D2E5D1D06FFCA57D1FF7DFA62FED7" pageId="8" pageNumber="369">
<paragraph id="B73636F81D06FFCA57D1FF7DFA62FF35" blockId="8.[151,1436,151,320]" box="[151,1436,210,232]" pageId="8" pageNumber="369">
<keyLead id="0C7895CD1D06FFCA57D1FF7DFA62FF35" box="[151,1436,210,232]" pageId="8" pageNumber="369">
3 Coxa 4 smaller than coxae 2 and 3; coxa 5 anterior lobe smaller than posterior one
<taxonomicName id="70894D7B1D06FFCA5347FF7DFA62FF35" ID-CoL="7ZL4D" authority="Wang, Zhu, Sha &amp; Ren, 2019" authorityName="Wang, Zhu, Sha &amp; Ren" authorityYear="2019" box="[1025,1436,210,232]" class="Malacostraca" family="Sebidae" genus="Seba" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="369" phylum="Arthropoda" rank="species" species="longimera">
<emphasis id="85FDEAEA1D06FFCA5347FF7DFB82FF35" box="[1025,1148,210,231]" italics="true" pageId="8" pageNumber="369">S. longimera</emphasis>
<bibRefCitation id="D3184B091D06FFCA53C7FF7CFA62FF35" author="Wang, Y. &amp; Zhu, C. &amp; Sha, Z. &amp; Ren, X." box="[1153,1436,210,232]" pageId="8" pageNumber="369" pagination="141 - 149" refId="ref4768" refString="Wang, Y., Zhu, C., Sha, Z. &amp; Ren, X. (2019) Description of Seba longimera sp. nov. from hydrothermal vents in the Okinawa Trough, Northwest Pacific (Amphipoda, Amphilochoidea, Sebidae). ZooKeys, 899, 141 - 149. https: // doi. org / 10.3897 / zookeys. 899.39442" type="journal article" year="2019">Wang, Zhu, Sha &amp; Ren, 2019</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B73636F81D06FFCA57D1FF5FFA62FED7" blockId="8.[151,1436,151,320]" box="[151,1436,239,261]" pageId="8" pageNumber="369">
<keyLead id="0C7895CD1D06FFCA57D1FF5FFA62FED7" box="[151,1436,239,261]" pageId="8" pageNumber="369">- Coxa 4 larger than coxae 2 and 3; coxa 5 anterior lobe larger than posterior one.................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="0C7D2E5D1D06FFCA57D1FEA2FA62FEED" pageId="8" pageNumber="369">
<paragraph id="B73636F81D06FFCA57D1FEA2FA62FEF0" blockId="8.[151,1436,151,320]" box="[151,1436,269,291]" pageId="8" pageNumber="369">
<keyLead id="0C7895CD1D06FFCA57D1FEA2FA62FEF0" box="[151,1436,269,291]" pageId="8" pageNumber="369">
<specimenCount id="A18FFD711D06FFCA57D1FEA2FEEDFEF1" box="[151,275,269,291]" count="4" pageId="8" pageNumber="369" type="male">4 Male</specimenCount>
gnathopod 2 carpus length 1.2 times of width; uropod 3 ramus with robust setae.............
<taxonomicName id="70894D7B1D06FFCA53F1FEA2FA62FEF0" ID-CoL="4W782" authority="Shaw, 1989" authorityName="Shaw" authorityYear="1989" box="[1207,1436,269,291]" class="Malacostraca" family="Sebidae" genus="Seba" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="369" phylum="Arthropoda" rank="species" species="profunda">
<emphasis id="85FDEAEA1D06FFCA53F1FEA2FAD8FEF0" box="[1207,1318,269,290]" italics="true" pageId="8" pageNumber="369">S. profunda</emphasis>
<bibRefCitation id="D3184B091D06FFCA526AFEA2FA62FEF0" author="Shaw, P." box="[1324,1436,269,291]" pageId="8" pageNumber="369" pagination="1882 - 1890" refId="ref4544" refString="Shaw, P. (1989) New amphipods from geothermal vent sites off the west coast of Vancouver Island, British Columbia, with a reappraisal of the amphipod family Sebidae. Canadian Journal of Zoology, 67, 1882 - 1890. https: // doi. org / 10.1139 / z 89 - 269" type="journal article" year="1989">Shaw, 1989</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B73636F81D06FFCA57D1FE84FA62FEED" blockId="8.[151,1436,151,320]" box="[151,1436,298,320]" pageId="8" pageNumber="369">
<keyLead id="0C7895CD1D06FFCA57D1FE84FA62FEED" box="[151,1436,298,320]" pageId="8" pageNumber="369">
- Male gnathopod 2 carpus length 2.3 times of width; uropod 3 ramus without setae.............
<taxonomicName id="70894D7B1D06FFCA53D6FE85FA62FEED" ID-CoL="4W77L" authority="(Chevreux, 1889)" baseAuthorityName="Chevreux" baseAuthorityYear="1889" box="[1168,1436,298,320]" class="Malacostraca" family="Sebidae" genus="Seba" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="369" phylum="Arthropoda" rank="species" species="armata">
<emphasis id="85FDEAEA1D06FFCA53D6FE85FB13FE92" box="[1168,1261,298,320]" italics="true" pageId="8" pageNumber="369">S. armata</emphasis>
(
<bibRefCitation id="D3184B091D06FFCA53BFFE85FA6BFEED" author="Chevreux, E." box="[1273,1429,298,320]" pageId="8" pageNumber="369" pagination="283 - 289" refId="ref4037" refString="Chevreux, E. (1889) Amphipodes nouveaux provenant des campagnes de l'Hirondelle 1887 - 1888. Bulletin de la Societe zoologique de France, 14, 283 - 289." type="journal article" year="1889">Chevreux, 1889</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>