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<mods:title id="83295D74AB3A04A6FEA504656A695B7A">The taxonomy and nomenclature associated with Kalanchoe adelae (Crassulaceae subfam. Kalanchooideae)</mods:title>
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<mods:date id="86177E7BF222CA64D2C5AA01FA9B4A22">2024</mods:date>
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<treatment id="038C8793FFB7012B709854FAFC48059E" ID-DOI="http://doi.org/10.5281/zenodo.14516022" ID-Zenodo-Dep="14516022" LSID="urn:lsid:plazi:treatment:038C8793FFB7012B709854FAFC48059E" httpUri="http://treatment.plazi.org/id/038C8793FFB7012B709854FAFC48059E" lastPageId="3" lastPageNumber="36" pageId="1" pageNumber="34" scope_family="Crassulaceae">
<subSubSection id="C33F650EFFB70129709854FAFDD205AC" box="[136,626,1940,1967]" pageId="1" pageNumber="34" type="nomenclature">
<paragraph id="8B9A3685FFB70129709854FAFDD205AC" blockId="1.[136,626,1940,1967]" box="[136,626,1940,1967]" pageId="1" pageNumber="34">
<heading id="D0D281E9FFB70129709854FAFDD205AC" box="[136,626,1940,1967]" fontSize="11" level="2" pageId="1" pageNumber="34" reason="3">
<treatmentCitationGroup id="AB3511ABFFB70129709854FAFDD205AC" box="[136,626,1940,1967]" pageId="1" pageNumber="34">
<treatmentCitation id="0A841094FFB70129709854FAFD8C05AC" author="Hamet, R." box="[136,556,1940,1967]" page="878" pageId="1" pageNumber="34" year="1907">
<taxonomicName id="4C254D06FFB70129709854FAFD8C05AC" ID-CoL="3QYFP" authority="Hamet (1907: 878" authorityName="R. Hamet" authorityYear="1907" box="[136,556,1940,1967]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="1" pageNumber="34" phylum="Tracheophyta" rank="species" species="adelae">
<emphasis id="B951EA97FFB70129709854FAFEF705AD" bold="true" box="[136,343,1940,1967]" italics="true" pageId="1" pageNumber="34">Kalanchoe adelae</emphasis>
<bibRefCitation id="EFB44B74FFB70129714D54FBFD8C05AC" author="Hamet, R." box="[349,556,1940,1967]" pageId="1" pageNumber="34" pagination="869 - 900" refId="ref3796" refString="Hamet, R. (1907) Monographie du genre Kalanchoe. (A suivre. [To be continued]) Bulletin de l'Herbier Boissier 2 me ser. 7 (11): 869 - 900. [https: // www. biodiversitylibrary. org / item / 105267 # page / 939 / mode / 1 up]" type="journal article" year="1907">Hamet (1907: 878</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 882).
</treatmentCitationGroup>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33F650EFFB7012B70AD54D7FC48059E" lastPageId="3" lastPageNumber="36" pageId="1" pageNumber="34" type="materials_examined">
<paragraph id="8B9A3685FFB7012970AD54D7FD760A18" blockId="1.[136,1452,1976,2074]" pageId="1" pageNumber="34">
<emphasis id="B951EA97FFB7012970AD54D7FF5105D0" box="[189,241,1977,2002]" italics="true" pageId="1" pageNumber="34">Type</emphasis>
:—[
<collectingCountry id="F3327615FFB70129710954D6FDBA05D0" box="[281,538,1976,2002]" name="Comoros" pageId="1" pageNumber="34">COMORO ISLANDS</collectingCountry>
.
<collectingRegion id="49E1F867FFB70129723354D6FD7B05D0" box="[547,731,1976,2002]" country="Comoros" name="Grande Comore" pageId="1" pageNumber="34">Grande Comore</collectingRegion>
], undated, likely
<date id="FF9B1045FFB70129738754D6FBA605D0" box="[919,1030,1976,2002]" pageId="1" pageNumber="34" value="1886-07">July 1886</date>
, [Mr
<emphasis id="B951EA97FFB70129745154D7FB2705D0" box="[1089,1159,1977,2002]" italics="true" pageId="1" pageNumber="34">
<collectionCode id="ED34AE40FFB70129745154D7FBF305D0" box="[1089,1107,1977,2002]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="1" pageNumber="34" type="Herbarium">L</collectionCode>
.
<collectionCode id="ED34AE40FFB70129744754D7FBC705D0" box="[1111,1127,1977,2002]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="1" pageNumber="34" type="Herbarium">J</collectionCode>
.
<collectionCode id="ED34AE40FFB70129747A54D7FB2305D0" box="[1130,1155,1977,2002]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="1" pageNumber="34" type="Herbarium">H</collectionCode>
.
</emphasis>
]
<emphasis id="B951EA97FFB70129748654D6FA9605D0" box="[1174,1334,1976,2002]" italics="true" pageId="1" pageNumber="34">Humblot 1570</emphasis>
(
<typeStatus id="549E8827FFB70129755354D6FA0C05D0" box="[1347,1452,1976,2002]" pageId="1" pageNumber="34" type="lectotype">lectotype</typeStatus>
Herb.
<collectionCode id="ED34AE40FFB7012970DF54B3FF7E05F5" box="[207,222,2013,2039]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="1" pageNumber="34" type="Herbarium">P</collectionCode>
barcodeP00431000 [image of the specimen accessible online at http://coldb.mnhn.fr/catalognumber/mnhn/p/ p00431000]),
<emphasis id="B951EA97FFB7012971375B6EFDD40A18" bold="true" box="[295,628,2048,2074]" pageId="1" pageNumber="34">
here designated as
<typeStatus id="549E8827FFB7012972195B6EFDD40A18" box="[521,628,2048,2074]" pageId="1" pageNumber="34" type="lectotype">lectotype</typeStatus>
</emphasis>
(
<figureCitation id="131E2A00FFB7012972935B6EFD640A18" box="[643,708,2048,2074]" captionStart="FIGURE 1" captionStartId="2.[136,229,1953,1975]" captionTargetBox="[233,1354,190,1929]" captionTargetId="figure-20@2.[233,1354,190,1929]" captionTargetPageId="2" captionText="FIGURE 1. Lectotype, [L.J.H.] Humblot 1570 (Herb. P barcode P00431000 [image of the specimen accessible online at http://coldb. mnhn.fr/catalognumber/mnhn/p/p00431000]), of the name Kalanchoe adelae designated in this paper: “[COMORO ISLANDS. Grande Comore], undated, likely July 1886”. Copyright of the Muséum national d’Histoire naturelle (MNHN—Paris, France), and reproduced with their permission, which is gratefully acknowledged." figureDoi="http://doi.org/10.5281/zenodo.14516026" httpUri="https://zenodo.org/record/14516026/files/figure.png" pageId="1" pageNumber="34">Fig. 1</figureCitation>
).
</paragraph>
<caption id="DF5A660DFFB4012A709854CFFD0C0A21" ID-DOI="http://doi.org/10.5281/zenodo.14516026" ID-Zenodo-Dep="14516026" httpUri="https://zenodo.org/record/14516026/files/figure.png" pageId="2" pageNumber="35" startId="2.[136,229,1953,1975]" targetBox="[233,1354,190,1929]" targetPageId="2" targetType="figure">
<paragraph id="8B9A3685FFB4012A709854CFFD0C0A21" blockId="2.[136,1452,1953,2083]" pageId="2" pageNumber="35">
<emphasis id="B951EA97FFB4012A709854CFFEA005B4" bold="true" box="[136,256,1953,1975]" pageId="2" pageNumber="35">FIGURE 1.</emphasis>
Lectotype, [
<emphasis id="B951EA97FFB4012A716E54CCFE1A05B5" box="[382,442,1954,1975]" italics="true" pageId="2" pageNumber="35">L.J.H.</emphasis>
]
<emphasis id="B951EA97FFB4012A71D954CFFDF505B4" box="[457,597,1953,1974]" italics="true" pageId="2" pageNumber="35">Humblot 1570</emphasis>
(Herb. P barcode P00431000 [image of the specimen accessible online at http://coldb. mnhn.fr/catalognumber/mnhn/p/p00431000]), of the name
<taxonomicName id="4C254D06FFB4012A72D454ABFCD405D8" authorityName="R. Hamet" authorityYear="1907" box="[708,884,1989,2010]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="2" pageNumber="35" phylum="Tracheophyta" rank="species" species="adelae">
<emphasis id="B951EA97FFB4012A72D454ABFCD405D8" box="[708,884,1989,2010]" italics="true" pageId="2" pageNumber="35">Kalanchoe adelae</emphasis>
</taxonomicName>
designated in this paper: “[COMORO ISLANDS. Grande Comore], undated, likely July 1886”. Copyright of the
<emphasis id="B951EA97FFB4012A72B25487FBB105FC" box="[674,1041,2025,2046]" italics="true" pageId="2" pageNumber="35">Muséum national d’Histoire naturelle</emphasis>
(MNHN—Paris, France), and reproduced with their permission, which is gratefully acknowledged.
</paragraph>
</caption>
<paragraph id="8B9A3685FFB5012B70AD53F1FBE302BA" blockId="3.[136,1452,158,1949]" box="[189,1091,158,185]" pageId="3" pageNumber="36">
<emphasis id="B951EA97FFB5012B70AD53F1FE0102BA" box="[189,417,159,184]" italics="true" pageId="3" pageNumber="36">Homotypic synonym</emphasis>
:—
<taxonomicName id="4C254D06FFB5012B71D553F0FD0802BA" box="[453,680,158,184]" class="Magnoliopsida" family="Crassulaceae" genus="Bryophyllum" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="36" phylum="Tracheophyta" rank="species" species="adelae">
<emphasis id="B951EA97FFB5012B71D553F0FD0802BA" box="[453,680,158,184]" italics="true" pageId="3" pageNumber="36">Bryophyllum adelae</emphasis>
</taxonomicName>
(Raym.-Hamet)
<bibRefCitation id="EFB44B74FFB5012B737653F1FB9F02BA" author="Berger, A." box="[870,1087,158,185]" pageId="3" pageNumber="36" pagination="402 - 412" refId="ref3386" refString="Berger, A. (1930) Crassulaceae. Unterfam. II. Kalanchoideae [sic, Kalanchooideae]. 6. Kalanchoe Adanson. 7. Kitchingia Baker. 8. Bryophyllum Salisb. In: Engler, A. &amp; Prantl, K. (Eds.) Die naturlichen Pflanzenfamilien, edn 2, 18 a. Verlag von Wilhelm Engelmann, Leipzig, pp. 378 - 386, 402 - 412." type="book chapter" year="1930">Berger (1930: 412)</bibRefCitation>
.
</paragraph>
<paragraph id="8B9A3685FFB5012B70AD53ADFD590303" blockId="3.[136,1452,158,1949]" pageId="3" pageNumber="36">
<emphasis id="B951EA97FFB5012B70AD53ADFE0B02DE" box="[189,427,195,220]" italics="true" pageId="3" pageNumber="36">Heterotypic synonym</emphasis>
:—
<taxonomicName id="4C254D06FFB5012B71DF53ACFC1402DF" authority="Tulasne (1857: 150)" authorityName="Tulasne" authorityPageNumber="150" authorityYear="1857" box="[463,948,194,221]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="36" phylum="Tracheophyta" rank="species" species="floribunda">
<emphasis id="B951EA97FFB5012B71DF53ACFD6602DE" box="[463,710,194,220]" italics="true" pageId="3" pageNumber="36">Kalanchoe floribunda</emphasis>
<bibRefCitation id="EFB44B74FFB5012B72DE53ACFC1402DF" author="Tulasne, L. - R." box="[718,948,194,221]" pageId="3" pageNumber="36" pagination="44 - 163" refId="ref5292" refString="Tulasne, L. - R. [' Edmond'] (1857) Florae madagascariensis. Fragmenta scripsit collectave digessit. Fragmentum alterum (1). Passifloreas, Homalineas, Chailletieas, Hippocrateaceas, Celastrineas, Ilicineas, Rhamneas, Pittosporeas, Hamamelideas, Crassulaceas et Saxifragaceas exhibens. [Crassulaceae. I. Bryophyllum. II. Calanchoe (sic, Kalanchoe) on pp. 147 - 150.] Annales des Sciences Naturelles Botanique ser. 4 8: 44 - 163. [https: // www. biodiversitylibrary. org / item / 129817 # page / 151 / mode / 1 up]" type="journal article" year="1857">Tulasne (1857: 150)</bibRefCitation>
</taxonomicName>
, non [
<taxonomicName id="4C254D06FFB5012B73ED53ADFB3502DE" authorityName="Tulasne" authorityYear="1857" box="[1021,1173,194,220]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="36" phylum="Tracheophyta" rank="species" species="floribunda">
<emphasis id="B951EA97FFB5012B73ED53ADFB3502DE" box="[1021,1173,194,220]" italics="true" pageId="3" pageNumber="36">K. floribunda</emphasis>
</taxonomicName>
]
<bibRefCitation id="EFB44B74FFB5012B74B753ACFEAE0303" author="Wight, R. &amp; Walker-Arnot, G. A." pageId="3" pageNumber="36" refId="ref5565" refString="Wight, R. &amp; Walker-Arnot, G. A. (1834) Prodromus florae peninsulae indiae orientalis: containing abridged descriptions of the plants found in the Peninsula of British India, arranged according to the natural system. Vol. I. Parbury, Allen, &amp; Co., London, 480 pp. https: // doi. org / 10.5962 / bhl. title. 252" type="book" year="1834">Wight &amp; Walker-Arnot (1834: 359)</bibRefCitation>
,
<emphasis id="B951EA97FFB5012B71095389FE2A0302" box="[281,394,230,256]" italics="true" pageId="3" pageNumber="36">nom. illeg</emphasis>
. (
<bibRefCitation id="EFB44B74FFB5012B718F5388FDDA0302" author="Turland, N. J. &amp; Wiersema, J. H. &amp; Barrie, F. R. &amp; Greuter, W. &amp; Hawksworth, D. L. &amp; Herendeen, P. S. &amp; Knapp, S. &amp; Kusber, W. - H. &amp; Li, D. - Z. &amp; Marhold, K. &amp; May, T. W. &amp; McNeill, J. &amp; Monro, A. M. &amp; Prado, J. &amp; Price, M. J. &amp; Smith, G. F." box="[415,634,230,257]" pageId="3" pageNumber="36" refId="ref5403" refString="Turland, N. J., Wiersema, J. H., Barrie, F. R., Greuter, W., Hawksworth, D. L., Herendeen, P. S., Knapp, S., Kusber, W. - H., Li, D. - Z., Marhold, K., May, T. W., McNeill, J., Monro, A. M., Prado, J., Price, M. J. &amp; Smith, G. F. (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Koeltz Botanical Books, Glashutten. [Regnum Vegetabile 159], 254 pp. https: // doi. org / 10.12705 / Code. 2018" type="book" year="2018">
Turland
<emphasis id="B951EA97FFB5012B71EF5389FD910302" box="[511,561,230,256]" italics="true" pageId="3" pageNumber="36">et al</emphasis>
. 2018
</bibRefCitation>
: Art. 53.1).
</paragraph>
<paragraph id="8B9A3685FFB5012B70AD5265FC03036E" blockId="3.[136,1452,158,1949]" pageId="3" pageNumber="36">
<emphasis id="B951EA97FFB5012B70AD5265FF510326" box="[189,241,267,292]" italics="true" pageId="3" pageNumber="36">Type</emphasis>
:—[
<collectingCountry id="F3327615FFB5012B71095264FDBD0326" box="[281,541,266,292]" name="Comoros" pageId="3" pageNumber="36">COMORO ISLANDS</collectingCountry>
.] Iles Comores, Angazija, 1847–1852,
<emphasis id="B951EA97FFB5012B73D85265FC470326" box="[968,999,267,292]" italics="true" pageId="3" pageNumber="36">
<collectionCode id="ED34AE40FFB5012B73D85265FC430326" box="[968,995,267,292]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="3" pageNumber="36" type="Herbarium">M</collectionCode>
.
</emphasis>
[
<emphasis id="B951EA97FFB5012B73E65264FB110326" box="[1014,1201,266,292]" italics="true" pageId="3" pageNumber="36">Louis-Hyacinthe</emphasis>
]
<emphasis id="B951EA97FFB5012B74D05265FA8F0326" box="[1216,1327,267,292]" italics="true" pageId="3" pageNumber="36">Boivin s.n</emphasis>
. (
<typeStatus id="549E8827FFB5012B75535264FA0C0326" box="[1347,1452,266,292]" pageId="3" pageNumber="36" type="lectotype">lectotype</typeStatus>
Herb.
<collectionCode id="ED34AE40FFB5012B70DE5241FF7D034B" box="[206,221,303,329]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="36" type="Herbarium">P</collectionCode>
barcode
<collectionCode id="ED34AE40FFB5012B71535240FEF2034A" box="[323,338,302,328]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="36" type="Herbarium">P</collectionCode>
00431004 [image of the specimen accessible online at http://coldb.mnhn.fr/catalognumber/mnhn/p/ p00431004]), designated by
<bibRefCitation id="EFB44B74FFB5012B71D8523CFCE1036E" author="Smith, G. F. &amp; Figueiredo, E." box="[456,833,338,364]" pageId="3" pageNumber="36" pagination="275 - 284" refId="ref4861" refString="Smith, G. F. &amp; Figueiredo, E. (2023 b) Names validly published in Kalanchoe (Crassulaceae subfam. Kalanchooideae) by Louis-Rene ' Edmond' Tulasne in 1857, with reference to names proposed in sched. by Hilsenberg and Bojer for two Malagasy species, and with notes on the type of the name K. tubiflora. Phytotaxa 584 (4): 275 - 284. https: // doi. org / 10.11646 / phytotaxa. 584.4.4" type="journal article" year="2023">Smith &amp; Figueiredo (2023b: 280)</bibRefCitation>
(
<figureCitation id="131E2A00FFB5012B7340523CFC36036E" box="[848,918,338,364]" captionStart="FIGURE 2" captionStartId="4.[136,229,1900,1922]" captionTargetBox="[238,1349,190,1876]" captionTargetId="figure-20@4.[238,1349,190,1876]" captionTargetPageId="4" captionText="FIGURE 2. Lectotype, M. [Mr] [Louis-Hyacinthe] Boivin s.n. (Herb. P barcode P00431004 [image of the specimen accessible online at http://coldb.mnhn.fr/catalognumber/mnhn/p/p00431004]), of the name Kalanchoe floribunda Tulasne (1857: 150), non Wight &amp; Walker- Arnot (1834: 359), nom. illeg. (Turland et al. 2018:Art. 53.1), designated by Smith &amp; Figueiredo (2023: 280): “[COMORO ISLANDS.] Iles Comores, Angazija, 1847–1852”. Copyright of the Muséum national d’Histoire naturelle (MNHN—Paris, France), and reproduced with their permission, which is gratefully acknowledged." figureDoi="http://doi.org/10.5281/zenodo.14516028" httpUri="https://zenodo.org/record/14516028/files/figure.png" pageId="3" pageNumber="36">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8B9A3685FFB5012B70AD5218FE1D03FF" blockId="3.[136,1452,158,1949]" pageId="3" pageNumber="36">
<emphasis id="B951EA97FFB5012B70AD5218FDED0392" box="[189,589,374,400]" italics="true" pageId="3" pageNumber="36">
Nomenclatural notes on the
<typeStatus id="549E8827FFB5012B72105219FD8F0392" box="[512,559,375,400]" pageId="3" pageNumber="36">type</typeStatus>
of
</emphasis>
<taxonomicName id="4C254D06FFB5012B72455218FCBD0392" authorityName="R. Hamet" authorityYear="1907" box="[597,797,374,400]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="36" phylum="Tracheophyta" rank="species" species="adelae">Kalanchoe adelae</taxonomicName>
:—When treating
<taxonomicName id="4C254D06FFB5012B73FA5219FBF70392" authorityName="R. Hamet" authorityYear="1907" box="[1002,1111,374,400]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="36" phylum="Tracheophyta" rank="species" species="adelae">
<emphasis id="B951EA97FFB5012B73FA5219FBF70392" box="[1002,1111,374,400]" italics="true" pageId="3" pageNumber="36">K. adelae</emphasis>
</taxonomicName>
, the only material that
<bibRefCitation id="EFB44B74FFB5012B75715219FF5E03B6" author="Hamet, R." pageId="3" pageNumber="36" pagination="17 - 48" refId="ref3858" refString="Hamet, R. (1908) Monographie du genre Kalanchoe (Suite et fin. [Continuation and end], with Errata on p. 17). Bulletin de l'Herbier Boissier 2 me ser. 8 (1): 17 - 48. [https: // www. biodiversitylibrary. org / item / 104945 # page / 31 / mode / 1 up]" type="journal article" year="1908">Hamet (1908: 26)</bibRefCitation>
cited was “[
<collectionCode id="ED34AE40FFB5012B719C52F4FE0A03B6" box="[396,426,410,436]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="3" pageNumber="36" type="Herbarium">M</collectionCode>
. [Mr] Humblot,
<superScript id="7C509BCDFFB5012B727552F5FDDB03AA" attach="left" box="[613,635,410,437]" fontSize="6" pageId="3" pageNumber="36">no</superScript>
1570!]”, “
<collectingCountry id="F3327615FFB5012B72E352F4FCDF03B6" box="[755,895,410,436]" name="Madagascar" pageId="3" pageNumber="36">Madagascar</collectingCountry>
” [sic, it should have been “
<collectingCountry id="F3327615FFB5012B74A652F4FAD403B6" box="[1206,1396,410,436]" name="Comoros" pageId="3" pageNumber="36">Comoro Islands</collectingCountry>
”]. At least
<specimenCount id="9D23FD0CFFB5012B70D152D0FECA03DA" box="[193,362,446,472]" count="4" pageId="3" pageNumber="36" type="generic">four specimens</specimenCount>
that were all collected by [
<emphasis id="B951EA97FFB5012B728252D1FD7803DA" box="[658,728,447,472]" italics="true" pageId="3" pageNumber="36">
<collectionCode id="ED34AE40FFB5012B728252D1FD0403DA" box="[658,676,447,472]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="3" pageNumber="36" type="Herbarium">L</collectionCode>
.
<collectionCode id="ED34AE40FFB5012B72B852D1FD1803DA" box="[680,696,447,472]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="3" pageNumber="36" type="Herbarium">J</collectionCode>
.
<collectionCode id="ED34AE40FFB5012B72AB52D1FD7403DA" box="[699,724,447,472]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="3" pageNumber="36" type="Herbarium">H</collectionCode>
.
</emphasis>
]
<emphasis id="B951EA97FFB5012B72F652D0FCE803DA" box="[742,840,446,472]" italics="true" pageId="3" pageNumber="36">Humblot</emphasis>
under his collecting number “
<emphasis id="B951EA97FFB5012B748552D0FB6D03DA" box="[1173,1229,446,472]" italics="true" pageId="3" pageNumber="36">1570</emphasis>
”, as cited by
<bibRefCitation id="EFB44B74FFB5012B757152D1FEA003FE" author="Hamet, R." pageId="3" pageNumber="36" pagination="17 - 48" refId="ref3858" refString="Hamet, R. (1908) Monographie du genre Kalanchoe (Suite et fin. [Continuation and end], with Errata on p. 17). Bulletin de l'Herbier Boissier 2 me ser. 8 (1): 17 - 48. [https: // www. biodiversitylibrary. org / item / 104945 # page / 31 / mode / 1 up]" type="journal article" year="1908">Hamet (1908: 26)</bibRefCitation>
, exist at Herb.
<collectionCode id="ED34AE40FFB5012B71B9528DFE1903FF" box="[425,441,483,509]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="36" type="Herbarium">P</collectionCode>
.
</paragraph>
<paragraph id="8B9A3685FFB5012B70AD5168FEEA006A" blockId="3.[136,1452,158,1949]" pageId="3" pageNumber="36">
These
<specimenCount id="9D23FD0CFFB5012B71185168FE140022" box="[264,436,518,544]" count="4" pageId="3" pageNumber="36" type="generic">four specimens</specimenCount>
—all of them
<typeStatus id="549E8827FFB5012B72425169FD140023" box="[594,692,519,545]" pageId="3" pageNumber="36" type="syntype">syntypes</typeStatus>
(see
<bibRefCitation id="EFB44B74FFB5012B72E05168FC6C0022" author="Turland, N. J. &amp; Wiersema, J. H. &amp; Barrie, F. R. &amp; Greuter, W. &amp; Hawksworth, D. L. &amp; Herendeen, P. S. &amp; Knapp, S. &amp; Kusber, W. - H. &amp; Li, D. - Z. &amp; Marhold, K. &amp; May, T. W. &amp; McNeill, J. &amp; Monro, A. M. &amp; Prado, J. &amp; Price, M. J. &amp; Smith, G. F." box="[752,972,518,545]" pageId="3" pageNumber="36" refId="ref5403" refString="Turland, N. J., Wiersema, J. H., Barrie, F. R., Greuter, W., Hawksworth, D. L., Herendeen, P. S., Knapp, S., Kusber, W. - H., Li, D. - Z., Marhold, K., May, T. W., McNeill, J., Monro, A. M., Prado, J., Price, M. J. &amp; Smith, G. F. (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Koeltz Botanical Books, Glashutten. [Regnum Vegetabile 159], 254 pp. https: // doi. org / 10.12705 / Code. 2018" type="book" year="2018">
Turland
<emphasis id="B951EA97FFB5012B73405169FC220022" box="[848,898,518,544]" italics="true" pageId="3" pageNumber="36">et al</emphasis>
. 2018
</bibRefCitation>
: Art. 9.6 and Note 1 under Art. 40)—were mounted on four different sheets. These specimens can be identified as follows with reference to the various labels attached to them:
</paragraph>
<paragraph id="8B9A3685FFB5012B70AD511CFEAF011E" blockId="3.[136,1452,158,1949]" pageId="3" pageNumber="36">
1.
<collectionCode id="ED34AE40FFB5012B70CB511CFF4E008E" box="[219,238,626,652]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="3" pageNumber="36" type="Herbarium">A</collectionCode>
printed label headed “Herbier
<collectionCode id="ED34AE40FFB5012B7248511DFDCC008F" box="[600,620,627,653]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="3" pageNumber="36" type="Herbarium">E</collectionCode>
. Drake” that was stamped “Humblot. –
<collectingCountry id="F3327615FFB5012B7450511CFB6A008E" box="[1088,1226,626,652]" name="Comoros" pageId="3" pageNumber="36">COMORES</collectingCountry>
” and completed by hand as “1570
<taxonomicName id="4C254D06FFB5012B712051F8FDD200B2" authority="R. Hamet" authorityName="R. Hamet" authorityYear="1907" box="[304,626,662,689]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="36" phylum="Tracheophyta" rank="species" species="adelae">Kalanchoe Adelae R. Hamet</taxonomicName>
” with the date “
<date id="FF9B1045FFB5012B733651F8FC0B00B2" box="[806,939,662,688]" pageId="3" pageNumber="36" value="1886-07">juillet 1886</date>
” added in the same handwriting. An undated, handwritten determinavit slip that was signed by Henri Humbert, stating: “=
<taxonomicName id="4C254D06FFB5012B73F451D5FB0F00D6" authority="Tul." authorityName="Tulasne" authorityYear="1857" box="[996,1199,698,725]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="36" phylum="Tracheophyta" rank="species" species="floribunda">K. floribunda Tul.</taxonomicName>
non
<collectionCode id="ED34AE40FFB5012B74F651D5FAA200D7" box="[1254,1282,699,725]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="3" pageNumber="36" type="Herbarium">W</collectionCode>
. et Arn” is also attached to the mounting board. [The preserved material is of a flowering specimen that lacks leaves.] Herb.
<collectionCode id="ED34AE40FFB5012B752E51B1FAED00FB" box="[1342,1357,735,761]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="36" type="Herbarium">P</collectionCode>
barcode
<collectionCode id="ED34AE40FFB5012B7098506CFF37011E" box="[136,151,770,796]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="36" type="Herbarium">P</collectionCode>
00430999.
</paragraph>
<paragraph id="8B9A3685FFB5012B70AD5048FECF01F6" blockId="3.[136,1452,158,1949]" pageId="3" pageNumber="36">
2.
<collectionCode id="ED34AE40FFB5012B70CA5048FF4D0142" box="[218,237,806,832]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="3" pageNumber="36" type="Herbarium">A</collectionCode>
printed label headed “
<collectionCode id="ED34AE40FFB5012B71E15048FDE10142" box="[497,577,806,832]" pageId="3" pageNumber="36">HERB</collectionCode>
.
<collectionCode id="ED34AE40FFB5012B725E5048FD2E0142" box="[590,654,806,832]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13677" lsid="urn:lsid:biocol.org:col:13677" name="Muskingum College" pageId="3" pageNumber="36" type="Herbarium">MUS</collectionCode>
.
<collectionCode id="ED34AE40FFB5012B728B5048FD4B0142" box="[667,747,806,832]" pageId="3" pageNumber="36">PARIS</collectionCode>
” that was completed by hand as “
<superScript id="7C509BCDFFB5012B74645048FB320136" attach="left" box="[1140,1170,806,832]" fontSize="6" pageId="3" pageNumber="36">No</superScript>
1570
<taxonomicName id="4C254D06FFB5012B74CD5048FEA00167" authority="R. Hamet" authorityName="R. Hamet" authorityYear="1907" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="36" phylum="Tracheophyta" rank="species" species="adelae">Kalanchoe Adelae R. Hamet</taxonomicName>
[.]
<collectionCode id="ED34AE40FFB5012B710C5025FE920167" box="[284,306,843,869]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="36">R</collectionCode>
. Hamet determinavit[.]
<collectionCode id="ED34AE40FFB5012B72585025FDC40167" box="[584,612,843,869]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="3" pageNumber="36" type="Herbarium">M</collectionCode>
. Humblot”.
<collectionCode id="ED34AE40FFB5012B72E35024FCA60166" box="[755,774,842,868]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="3" pageNumber="36" type="Herbarium">A</collectionCode>
small piece of paper attached to the specimen carries the handwritten number “1570”.
<collectionCode id="ED34AE40FFB5012B71DA5000FE7D018A" box="[458,477,878,904]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="3" pageNumber="36" type="Herbarium">A</collectionCode>
handwritten determinavit slip dated
<date id="FF9B1045FFB5012B73605000FB86018A" box="[880,1062,878,904]" pageId="3" pageNumber="36" value="1923-02-02">2 February 1923</date>
that was signed by Henri Humbert, stating: “=
<taxonomicName id="4C254D06FFB5012B711550FDFE6F01AE" authority="Tul." authorityName="Tulasne" authorityYear="1857" box="[261,463,914,941]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="36" phylum="Tracheophyta" rank="species" species="floribunda">K. floribunda Tul.</taxonomicName>
(non
<collectionCode id="ED34AE40FFB5012B721F50FDFD8B01AF" box="[527,555,915,941]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="3" pageNumber="36" type="Herbarium">W</collectionCode>
. et Arn.)” is also attached to the mounting board. [The preserved material is of a flowering specimen with leaves, as well as, presumably, plant fragments enclosed in a folded paper capsule.] Herb.
<collectionCode id="ED34AE40FFB5012B758D50D9FA0C01D3" box="[1437,1452,951,977]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="36" type="Herbarium">P</collectionCode>
barcode
<collectionCode id="ED34AE40FFB5012B70F850B4FF5701F6" box="[232,247,986,1012]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="36" type="Herbarium">P</collectionCode>
00431000.
</paragraph>
<paragraph id="8B9A3685FFB5012B70AD5090FEAF06CE" blockId="3.[136,1452,158,1949]" pageId="3" pageNumber="36">
3.
<collectionCode id="ED34AE40FFB5012B70C75090FF4A061A" box="[215,234,1022,1048]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="3" pageNumber="36" type="Herbarium">A</collectionCode>
printed label headed “Herbier
<collectionCode id="ED34AE40FFB5012B72515091FDF5061B" box="[577,597,1023,1049]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="3" pageNumber="36" type="Herbarium">E</collectionCode>
. Drake” that was stamped “Humblot. –
<collectingCountry id="F3327615FFB5012B741E5090FB38061A" box="[1038,1176,1022,1048]" name="Comoros" pageId="3" pageNumber="36">COMORES</collectingCountry>
” and completed by hand as “
<taxonomicName id="4C254D06FFB5012B70A3574CFE5E063E" authority="R. Hamet" authorityName="R. Hamet" authorityYear="1907" box="[179,510,1058,1085]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="36" phylum="Tracheophyta" rank="species" species="adelae">Kalanchoe Adelae R. Hamet</taxonomicName>
” with the date “
<date id="FF9B1045FFB5012B72AA574CFCE1063E" box="[698,833,1058,1084]" pageId="3" pageNumber="36" value="1886-07">juillet 1886</date>
” added in the same handwriting. The number “1570” handwritten on a small piece of paper is attached to the specimen.
<collectionCode id="ED34AE40FFB5012B739C5728FC3F0662" box="[908,927,1094,1120]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="3" pageNumber="36" type="Herbarium">A</collectionCode>
handwritten determinavit slip dated
<date id="FF9B1045FFB5012B75565728FF600686" pageId="3" pageNumber="36" value="1933-02">February 1933</date>
that was signed by Henri Humbert, stating: “=
<taxonomicName id="4C254D06FFB5012B72FD5704FB940687" authority="R. Hamet" authorityName="R. Hamet" authorityYear="1907" box="[749,1076,1130,1157]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="36" phylum="Tracheophyta" rank="species" species="adelae">Kalanchoe Adelae R. Hamet</taxonomicName>
=
<taxonomicName id="4C254D06FFB5012B74485705FA880686" authority="Tul." authorityName="Tulasne" authorityYear="1857" box="[1112,1320,1130,1157]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="36" phylum="Tracheophyta" rank="species" species="floribunda">K. floribunda Tul.</taxonomicName>
(non
<collectionCode id="ED34AE40FFB5012B757E5705FA2A0687" box="[1390,1418,1131,1157]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="3" pageNumber="36" type="Herbarium">W</collectionCode>
. et Arn.)” is also attached to the mounting board. [Leaf material only is attached to the mounting board.] Herb.
<collectionCode id="ED34AE40FFB5012B752D57E1FAEC06AB" box="[1341,1356,1167,1193]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="36" type="Herbarium">P</collectionCode>
barcode
<collectionCode id="ED34AE40FFB5012B709857DCFF3806CE" box="[136,152,1202,1228]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="36" type="Herbarium">P</collectionCode>
00431001.
</paragraph>
<paragraph id="8B9A3685FFB5012B70AD57B8FB0C0782" blockId="3.[136,1452,158,1949]" pageId="3" pageNumber="36">
4.
<collectionCode id="ED34AE40FFB5012B70CC57B8FF4F06F2" box="[220,239,1238,1264]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="3" pageNumber="36" type="Herbarium">A</collectionCode>
printed label of which the heading is obscured by leaf material, stating “
<taxonomicName id="4C254D06FFB5012B744157B8FA0106F2" authority="R. Hamet" authorityName="R. Hamet" authorityYear="1907" box="[1105,1441,1238,1265]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="36" phylum="Tracheophyta" rank="species" species="adelae">Kalanchoe Adelae R. Hamet</taxonomicName>
” and signed by
<collectionCode id="ED34AE40FFB5012B71205795FEE60717" box="[304,326,1275,1301]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="36">R</collectionCode>
. Hamet, with “
<collectingCountry id="F3327615FFB5012B71E45794FDDB0716" box="[500,635,1274,1300]" name="Madagascar" pageId="3" pageNumber="36">Madagascar</collectingCountry>
” [crossed out] and replaced by “Grande Comore”. The number “1570” handwritten on a small piece of paper is attached to the specimen. An undated, handwritten determinavit slip that was signed by Henri Humbert, stating: “=
<taxonomicName id="4C254D06FFB5012B7222562DFD5E075E" authority="Tul." authorityName="Tulasne" authorityYear="1857" box="[562,766,1346,1373]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="36" phylum="Tracheophyta" rank="species" species="floribunda">K. floribunda Tul.</taxonomicName>
non
<collectionCode id="ED34AE40FFB5012B7325562DFCF1075F" box="[821,849,1347,1373]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="3" pageNumber="36" type="Herbarium">W</collectionCode>
. et Arn.” is also attached to the mounting board. [The preserved material is of a flowering specimen with a few leaves.] Herb.
<collectionCode id="ED34AE40FFB5012B73BF5609FC1D0783" box="[943,957,1383,1409]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="36" type="Herbarium">P</collectionCode>
barcode
<collectionCode id="ED34AE40FFB5012B74345608FB930782" box="[1060,1075,1382,1408]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="36" type="Herbarium">P</collectionCode>
00431002.
</paragraph>
<paragraph id="8B9A3685FFB5012B70AD56E5FC550413" blockId="3.[136,1452,158,1949]" pageId="3" pageNumber="36">
<bibRefCitation id="EFB44B74FFB5012B70AD56E5FEDD07A6" author="Dorr, L. J." box="[189,381,1418,1445]" pageId="3" pageNumber="36" refId="ref3682" refString="Dorr, L. J. (1997) Plant collectors in Madagascar and the Comoro Islands. A biographical and bibliographical guide to individuals and groups who have collected herbarium material of algae, bryophytes, fungi, lichens, and vascular plants in Madagascar and the Comoro Islands. The Trustees, Royal Botanic Gardens, Kew, 524 pp." type="book" year="1997">Dorr (1997: 128)</bibRefCitation>
recorded that the herbarium of Emmanuel Drake del Castillo (1855–1904) includes collections made by, inter alia, Humblot, which explains the printed labels headed “Herbier
<collectionCode id="ED34AE40FFB5012B743556C1FB9907CB" box="[1061,1081,1455,1481]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="3" pageNumber="36" type="Herbarium">E</collectionCode>
. Drake” attached to some of the Humblot specimens. Other Humblot specimens were apparently sent directly to Herb.
<collectionCode id="ED34AE40FFB5012B744756BDFBC707EF" box="[1111,1127,1491,1517]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="36" type="Herbarium">P</collectionCode>
. Drake’s contributions to the taxonomy of
<taxonomicName id="4C254D06FFB5012B710D5698FE360412" authorityName="Adanson" authorityYear="1763" box="[285,406,1526,1552]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="36" phylum="Tracheophyta" rank="genus">
<emphasis id="B951EA97FFB5012B710D5698FE360412" box="[285,406,1526,1552]" italics="true" pageId="3" pageNumber="36">Kalanchoe</emphasis>
</taxonomicName>
were treated in
<bibRefCitation id="EFB44B74FFB5012B725B5698FC510413" author="Smith, G. F. &amp; Figueiredo, E." box="[587,1009,1526,1553]" pageId="3" pageNumber="36" pagination="158 - 166" refId="ref4732" refString="Smith, G. F. &amp; Figueiredo, E. (2022) Nomenclatural and taxonomic notes on the names published in Kalanchoe (Crassulaceae subfam. Kalanchooideae) by Drake del Castillo in 1903. Phytotaxa 547 (2): 158 - 166. https: // doi. org / 10.11646 / phytotaxa. 547.2.3" type="journal article" year="2022">Smith &amp; Figueiredo (2022: 158–165)</bibRefCitation>
.
</paragraph>
<paragraph id="8B9A3685FFB5012B70AD5574FBA7047E" blockId="3.[136,1452,158,1949]" pageId="3" pageNumber="36">
The four accessions enumerated as “1.” to “4.”, above, were all clearly given the number [
<emphasis id="B951EA97FFB5012B74BC5575FB520436" box="[1196,1266,1563,1588]" italics="true" pageId="3" pageNumber="36">
<collectionCode id="ED34AE40FFB5012B74BC5575FB1E0436" box="[1196,1214,1563,1588]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="3" pageNumber="36" type="Herbarium">L</collectionCode>
.
<collectionCode id="ED34AE40FFB5012B74D25575FB720436" box="[1218,1234,1563,1588]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="3" pageNumber="36" type="Herbarium">J</collectionCode>
.
<collectionCode id="ED34AE40FFB5012B74C55575FB4E0436" box="[1237,1262,1563,1588]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="3" pageNumber="36" type="Herbarium">H</collectionCode>
.
</emphasis>
]
<emphasis id="B951EA97FFB5012B75115574FAC30436" box="[1281,1379,1562,1588]" italics="true" pageId="3" pageNumber="36">Humblot</emphasis>
]
<emphasis id="B951EA97FFB5012B75635574FA0B0436" box="[1395,1451,1562,1588]" italics="true" pageId="3" pageNumber="36">1570</emphasis>
and all four qualify as original material because they were obviously available to Raymond-Hamet at Herb.
<collectionCode id="ED34AE40FFB5012B75735551FAD2045B" box="[1379,1394,1599,1625]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="36" type="Herbarium">P</collectionCode>
well before he published the name
<taxonomicName id="4C254D06FFB5012B71C9550DFDE4047E" authorityName="R. Hamet" authorityYear="1907" box="[473,580,1634,1660]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="36" phylum="Tracheophyta" rank="species" species="adelae">
<emphasis id="B951EA97FFB5012B71C9550DFDE4047E" box="[473,580,1634,1660]" italics="true" pageId="3" pageNumber="36">K. adelae</emphasis>
</taxonomicName>
in 1907 and treated the species in 1908.
</paragraph>
<paragraph id="8B9A3685FFB5012B70AD55E8FB0704C6" blockId="3.[136,1452,158,1949]" pageId="3" pageNumber="36">
Of these
<specimenCount id="9D23FD0CFFB5012B713255E8FE6D04A2" box="[290,461,1670,1696]" count="4" pageId="3" pageNumber="36" type="generic">four specimens</specimenCount>
the [
<emphasis id="B951EA97FFB5012B721655E9FDEC04A2" box="[518,588,1671,1696]" italics="true" pageId="3" pageNumber="36">
<collectionCode id="ED34AE40FFB5012B721655E9FDB804A2" box="[518,536,1671,1696]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="3" pageNumber="36" type="Herbarium">L</collectionCode>
.
<collectionCode id="ED34AE40FFB5012B720C55E9FD8C04A2" box="[540,556,1671,1696]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="3" pageNumber="36" type="Herbarium">J</collectionCode>
.
<collectionCode id="ED34AE40FFB5012B723F55E9FDE804A2" box="[559,584,1671,1696]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="3" pageNumber="36" type="Herbarium">H</collectionCode>
.
</emphasis>
]
<emphasis id="B951EA97FFB5012B724C55E8FD5C04A2" box="[604,764,1670,1696]" italics="true" pageId="3" pageNumber="36">Humblot 1570</emphasis>
specimen—the most complete one—that can be identified as Herb.
<collectionCode id="ED34AE40FFB5012B70DE55C5FF7D04C7" box="[206,221,1707,1733]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="36" type="Herbarium">P</collectionCode>
barcode
<collectionCode id="ED34AE40FFB5012B715355C4FEF204C6" box="[323,338,1706,1732]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="36" type="Herbarium">P</collectionCode>
00431000 is designated here as the
<typeStatus id="549E8827FFB5012B72F155C4FCE804C6" box="[737,840,1706,1732]" pageId="3" pageNumber="36" type="lectotype">lectotype</typeStatus>
of the name
<taxonomicName id="4C254D06FFB5012B73CA55C5FBE504C6" authorityName="R. Hamet" authorityYear="1907" box="[986,1093,1706,1732]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="36" phylum="Tracheophyta" rank="species" species="adelae">
<emphasis id="B951EA97FFB5012B73CA55C5FBE504C6" box="[986,1093,1706,1732]" italics="true" pageId="3" pageNumber="36">K. adelae</emphasis>
</taxonomicName>
(
<figureCitation id="131E2A00FFB5012B744455C4FB3604C6" box="[1108,1174,1706,1732]" captionStart="FIGURE 1" captionStartId="2.[136,229,1953,1975]" captionTargetBox="[233,1354,190,1929]" captionTargetId="figure-20@2.[233,1354,190,1929]" captionTargetPageId="2" captionText="FIGURE 1. Lectotype, [L.J.H.] Humblot 1570 (Herb. P barcode P00431000 [image of the specimen accessible online at http://coldb. mnhn.fr/catalognumber/mnhn/p/p00431000]), of the name Kalanchoe adelae designated in this paper: “[COMORO ISLANDS. Grande Comore], undated, likely July 1886”. Copyright of the Muséum national d’Histoire naturelle (MNHN—Paris, France), and reproduced with their permission, which is gratefully acknowledged." figureDoi="http://doi.org/10.5281/zenodo.14516026" httpUri="https://zenodo.org/record/14516026/files/figure.png" pageId="3" pageNumber="36">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8B9A3685FFB5012B70AD55A0FF5F057B" blockId="3.[136,1452,158,1949]" pageId="3" pageNumber="36">
<bibRefCitation id="EFB44B74FFB5012B70AD55A0FE6104EA" author="Descoings, B." box="[189,449,1742,1768]" pageId="3" pageNumber="36" pagination="143 - 181" refId="ref3645" refString="Descoings, B. (2003) Kalanchoe. In: Eggli, U. (Ed.) Illustrated handbook of succulent plants: Crassulaceae. Springer-Verlag, Berlin, pp. 143 - 181." type="book chapter" year="2003">Descoings (2003: 144)</bibRefCitation>
stated that the “
<emphasis id="B951EA97FFB5012B726F55A0FD3A04EA" bold="true" box="[639,666,1742,1768]" pageId="3" pageNumber="36">
<collectionCode id="ED34AE40FFB5012B726F55A0FD3304EA" box="[639,659,1742,1768]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="36">T</collectionCode>
:
</emphasis>
” of the name
<taxonomicName id="4C254D06FFB5012B735155A1FC0F04EA" authorityName="R. Hamet" authorityYear="1907" box="[833,943,1742,1768]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="36" phylum="Tracheophyta" rank="species" species="adelae">
<emphasis id="B951EA97FFB5012B735155A1FC0F04EA" box="[833,943,1742,1768]" italics="true" pageId="3" pageNumber="36">K. adelae</emphasis>
</taxonomicName>
is “
<collectingCountry id="F3327615FFB5012B73CD55A0FB3404EA" box="[989,1172,1742,1768]" name="Comoros" pageId="3" pageNumber="36">Comoro Islands</collectingCountry>
(
<emphasis id="B951EA97FFB5012B74B655A1FB4F04EA" box="[1190,1263,1743,1768]" italics="true" pageId="3" pageNumber="36">Boivin</emphasis>
s.n. [
<collectionCode id="ED34AE40FFB5012B752B55A0FAEF04EA" box="[1339,1359,1742,1768]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="36" type="Herbarium">P</collectionCode>
])”. This attempted typification of the name
<taxonomicName id="4C254D06FFB5012B720D559DFD2B050E" authorityName="R. Hamet" authorityYear="1907" box="[541,651,1778,1804]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="36" phylum="Tracheophyta" rank="species" species="adelae">
<emphasis id="B951EA97FFB5012B720D559DFD2B050E" box="[541,651,1778,1804]" italics="true" pageId="3" pageNumber="36">K. adelae</emphasis>
</taxonomicName>
on non-Humblot material was not effective. The abbreviation “
<emphasis id="B951EA97FFB5012B7579559CFA23050E" bold="true" box="[1385,1411,1778,1804]" pageId="3" pageNumber="36">
<collectionCode id="ED34AE40FFB5012B7579559CFADD050E" box="[1385,1405,1778,1804]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="36">T</collectionCode>
:
</emphasis>
” is defined simply as “(nomenclatural) type” in
<bibRefCitation id="EFB44B74FFB5012B72965478FCB00532" author="Eggli, U." box="[646,784,1814,1840]" pageId="3" pageNumber="36" refId="ref3745" refString="Eggli, U. (Ed.) (2003) Illustrated handbook of succulent plants: Crassulaceae. Springer-Verlag, Berlin &amp; Heidelberg, 458 pp + plates I - XLVIII. https: // doi. org / 10.1007 / 978 - 3 - 642 - 55874 - 0" type="book" year="2003">Eggli (2003</bibRefCitation>
: X). Rather, the specimen [
<emphasis id="B951EA97FFB5012B745A5478FAA50532" box="[1098,1285,1814,1840]" italics="true" pageId="3" pageNumber="36">Louis-Hyacinthe</emphasis>
]
<emphasis id="B951EA97FFB5012B75075479FA290532" box="[1303,1417,1815,1840]" italics="true" pageId="3" pageNumber="36">Boivin s.n</emphasis>
. is the
<typeStatus id="549E8827FFB5012B70A15454FEB80556" box="[177,280,1850,1876]" pageId="3" pageNumber="36" type="lectotype">lectotype</typeStatus>
of the name
<taxonomicName id="4C254D06FFB5012B71BA5455FC8B0557" authority="Tulasne (1857: 150)" authorityName="Tulasne" authorityPageNumber="150" authorityYear="1857" box="[426,811,1850,1877]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="36" phylum="Tracheophyta" rank="species" species="floribunda">
<emphasis id="B951EA97FFB5012B71BA5455FDE00556" box="[426,576,1850,1876]" italics="true" pageId="3" pageNumber="36">K. floribunda</emphasis>
<bibRefCitation id="EFB44B74FFB5012B72575454FC8B0557" author="Tulasne, L. - R." box="[583,811,1850,1877]" pageId="3" pageNumber="36" pagination="44 - 163" refId="ref5292" refString="Tulasne, L. - R. [' Edmond'] (1857) Florae madagascariensis. Fragmenta scripsit collectave digessit. Fragmentum alterum (1). Passifloreas, Homalineas, Chailletieas, Hippocrateaceas, Celastrineas, Ilicineas, Rhamneas, Pittosporeas, Hamamelideas, Crassulaceas et Saxifragaceas exhibens. [Crassulaceae. I. Bryophyllum. II. Calanchoe (sic, Kalanchoe) on pp. 147 - 150.] Annales des Sciences Naturelles Botanique ser. 4 8: 44 - 163. [https: // www. biodiversitylibrary. org / item / 129817 # page / 151 / mode / 1 up]" type="journal article" year="1857">Tulasne (1857: 150)</bibRefCitation>
</taxonomicName>
, non [
<taxonomicName id="4C254D06FFB5012B73615455FBA80556" authorityName="Tulasne" authorityYear="1857" box="[881,1032,1850,1876]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="36" phylum="Tracheophyta" rank="species" species="floribunda">
<emphasis id="B951EA97FFB5012B73615455FBA80556" box="[881,1032,1850,1876]" italics="true" pageId="3" pageNumber="36">K. floribunda</emphasis>
</taxonomicName>
]
<bibRefCitation id="EFB44B74FFB5012B74085454FA070557" author="Wight, R. &amp; Walker-Arnot, G. A." box="[1048,1447,1850,1877]" pageId="3" pageNumber="36" refId="ref5565" refString="Wight, R. &amp; Walker-Arnot, G. A. (1834) Prodromus florae peninsulae indiae orientalis: containing abridged descriptions of the plants found in the Peninsula of British India, arranged according to the natural system. Vol. I. Parbury, Allen, &amp; Co., London, 480 pp. https: // doi. org / 10.5962 / bhl. title. 252" type="book" year="1834">Wight &amp; Walker-Arnot (1834: 359)</bibRefCitation>
,
<emphasis id="B951EA97FFB5012B70985431FF58057A" box="[136,248,1886,1912]" italics="true" pageId="3" pageNumber="36">nom. illeg</emphasis>
.
</paragraph>
<paragraph id="8B9A3685FFB5012B70AD54EDFC48059E" blockId="3.[136,1452,158,1949]" box="[189,1000,1922,1949]" pageId="3" pageNumber="36">
<emphasis id="B951EA97FFB5012B70AD54EDFECE059E" box="[189,366,1923,1948]" italics="true" pageId="3" pageNumber="36">Taxonomic note</emphasis>
:—The name
<taxonomicName id="4C254D06FFB5012B721854EDFDD4059E" authorityName="R. Hamet" authorityYear="1907" box="[520,628,1922,1948]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="36" phylum="Tracheophyta" rank="species" species="adelae">
<emphasis id="B951EA97FFB5012B721854EDFDD4059E" box="[520,628,1922,1948]" italics="true" pageId="3" pageNumber="36">K. adelae</emphasis>
</taxonomicName>
is in use for an accepted species.
</paragraph>
</subSubSection>
</treatment>
</document>

341
data/03/8C/87/038C87BE1701817A4C9FF9C6FF707CBF.xml
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@ -1,268 +1,279 @@
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<mods:affiliation id="100087E2DDF5B27E4EF7CFC23E3966D6">Charles University, Faculty of ºcience, Department of Zoology, Viničná 7, 12800 Prague 2, Czech Republic &amp; Museum of Eastern Bohemia in Hradec Králové, Eliščino nábřeží 465, 500 03 Hradec Králové 3, Czech Republic</mods:affiliation>
<mods:nameIdentifier id="B9885B163A78134B35862FDB63908B6B" type="email">j.prazak@muzeumhk.cz</mods:nameIdentifier>
<mods:namePart id="E3086D035257B794750D8E9F5DF61290">Yamamoto, Shûhei</mods:namePart>
<mods:affiliation id="1DF28B3D5C5A4F6F67DFE545B7CB9E64">The Hokkaido University Museum, Hokkaido University, Kita 10, Nishi 8, Kita-ku, Sapporo 060 - 0810, Japan</mods:affiliation>
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<mods:namePart id="D90C0F79477A801D7818566FB39124C7">Yamamoto</mods:namePart>
<mods:affiliation id="F86A57A9E4E04C2CF3490FF17FEE125C">Ŋe Hokkaido University Museum, Hokkaido University, Kita 10, Nishi 8, Kita-ku, ºapporo 060 - 0810, Japan</mods:affiliation>
<mods:namePart id="914F98C3C72FA6E9658D51E2A1988A8E">Lackner, Tomáš</mods:namePart>
<mods:affiliation id="4B70C937979459A3777E4FE4E415F99A">Department of Environmental Systems Science, ETH Zürich, Weinbergstrasse 56, 8092 Zürich, Switzerland</mods:affiliation>
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<mods:namePart id="A944211E2466C26DB7A0F78EC44F08EC">Lackner, Tomáš</mods:namePart>
<mods:affiliation id="289B806E3A20604C23BA0D0EE27A774E">Department of Environmental ºystems ºcience, ETH Zürich, Weinbergstrasse 56, 8092 Zürich, ºwitzerland</mods:affiliation>
<mods:namePart id="DC71F1A729761BB0C722727BB3AC34E1">Fikáček, Martin</mods:namePart>
<mods:affiliation id="ABC53E65F85816BACBD1FFE825E37120">Department of Biological Sciences, National Sun Yat-sen University, No. 70, Lienhai Rd., Kaohsiung 80424, Taiwan &amp; Department of Entomology, National Museum, Cirkusová 1740 CZ- 19300 Praha 9, Czech Republic</mods:affiliation>
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<mods:namePart id="0D08A7F588F7A46DF51963D22C97A04A">Fikáček, Martin</mods:namePart>
<mods:affiliation id="F0B58DE9D56F4B098A0682CC14233CFA">Department of Biological ºciences, National ºun Yat-sen University, No. 70, Lienhai Rd., Kaohsiung 80424, Taiwan &amp; Department of Entomology, National Museum, Cirkusová 1740 CZ- 19300 Praha 9, Czech Republic</mods:affiliation>
<mods:namePart id="4D82BF5A26692A6BDB3EA7D3DDD49E1F">Prokop, Jakub</mods:namePart>
<mods:affiliation id="57D0D99D3E55FDEF29B091FC7AD99A05">Charles University, Faculty of Science, Department of Zoology, Viničná 7, 12800 Prague 2, Czech Republic &amp; Department of Plant and Environmental Sciences, Clemson University, Clemson, SC 29634, United States</mods:affiliation>
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<mods:namePart id="C1385F84CA470BE40C302E62FDCE5511">Prokop, Jakub</mods:namePart>
<mods:affiliation id="42BD7A16B4884089ACA4E36F509A89B4">Charles University, Faculty of ºcience, Department of Zoology, Viničná 7, 12800 Prague 2, Czech Republic</mods:affiliation>
<mods:namePart id="3BA0E34A68A529F5C168BE3E0FBC3F13">Caterino, Michael S</mods:namePart>
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<mods:affiliation id="8A45A9F7D45239AF64656CFCF3D98742">Department of Plant and Environmental ºciences, Clemson University, Clemson, ºC 29634, United ºtates</mods:affiliation>
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<mods:date id="822F01019E102E76D9AD2855920DF309">2024</mods:date>
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<mods:title id="DE3E4410079E0D53F3851971384A4A5A">zlae 137</mods:title>
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<treatment id="038C87BE1701817A4C9FF9C6FF707CBF" ID-DOI="http://doi.org/10.5281/zenodo.14424556" ID-Zenodo-Dep="14424556" LSID="urn:lsid:plazi:treatment:038C87BE1701817A4C9FF9C6FF707CBF" httpUri="http://treatment.plazi.org/id/038C87BE1701817A4C9FF9C6FF707CBF" lastPageId="12" pageId="9">
<subSubSection id="C33F65231701817F4C9FF9C6FAB67D8F" box="[967,1302,1638,1665]" pageId="9" type="nomenclature">
<paragraph id="8B9A36A81701817F4C9FF9C6FAB67D8F" blockId="9.[967,1302,1638,1665]" box="[967,1302,1638,1665]" pageId="9">
<heading id="D0D281C41701817F4C9FF9C6FAB67D8F" bold="true" box="[967,1302,1638,1665]" centered="true" fontSize="11" level="2" pageId="9" reason="2">
<emphasis id="B951EABA1701817F4C9FF9C6FAB67D8F" bold="true" box="[967,1302,1638,1665]" pageId="9">
<taxonomicName id="4C254D2B1701817F4C9FF9C6FB1A7D8F" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[967,1210,1638,1665]" class="Insecta" family="Histeridae" genus="Palaeabraeus" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="glabrus" status="sp. nov.">
<emphasis id="B951EABA1701817F4C9FF9C6FB1A7D8F" bold="true" box="[967,1210,1638,1665]" italics="true" pageId="9">Palaeabraeus glabrus</emphasis>
<treatment id="038C87BE1701817A4C9FF9C6FF707CBF" ID-DOI="http://doi.org/10.5281/zenodo.14424556" ID-Zenodo-Dep="14424556" LSID="urn:lsid:plazi:treatment:038C87BE1701817A4C9FF9C6FF707CBF" httpUri="http://treatment.plazi.org/id/038C87BE1701817A4C9FF9C6FF707CBF" lastPageId="12" lastPageNumber="13" pageId="9" pageNumber="10">
<subSubSection id="C33F65231701817F4C9FF9C6FAB67D8F" box="[967,1302,1638,1665]" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="8B9A36A81701817F4C9FF9C6FAB67D8F" blockId="9.[967,1302,1638,1665]" box="[967,1302,1638,1665]" pageId="9" pageNumber="10">
<heading id="D0D281C41701817F4C9FF9C6FAB67D8F" bold="true" box="[967,1302,1638,1665]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="10" reason="2">
<emphasis id="B951EABA1701817F4C9FF9C6FAB67D8F" bold="true" box="[967,1302,1638,1665]" pageId="9" pageNumber="10">
<taxonomicName id="4C254D2B1701817F4C9FF9C6FB1A7D8F" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[967,1210,1638,1665]" class="Insecta" family="Histeridae" genus="Palaeabraeus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="glabrus" status="sp. nov.">
<emphasis id="B951EABA1701817F4C9FF9C6FB1A7D8F" bold="true" box="[967,1210,1638,1665]" italics="true" pageId="9" pageNumber="10">Palaeabraeus glabrus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C11701817F4BE7F9C7FAB67D8F" box="[1215,1302,1639,1665]" pageId="9" rank="species">sp. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="A26257C11701817F4BE7F9C7FAB67D8F" box="[1215,1302,1639,1665]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33F65231701817A4B1FF905FF707CBF" lastPageId="12" pageId="9" type="description">
<paragraph id="8B9A36A81701817F4B1FF905FB367DB1" blockId="9.[1095,1174,1701,1727]" box="[1095,1174,1701,1727]" pageId="9">
<emphasis id="B951EABA1701817F4B1FF905FB367DB1" bold="true" box="[1095,1174,1701,1727]" pageId="9">
<subSubSection id="C33F65231701817F4B1FF905FB367DB1" box="[1095,1174,1701,1727]" pageId="9" pageNumber="10" type="description">
<paragraph id="8B9A36A81701817F4B1FF905FB367DB1" blockId="9.[1095,1174,1701,1727]" box="[1095,1174,1701,1727]" pageId="9" pageNumber="10">
<emphasis id="B951EABA1701817F4B1FF905FB367DB1" bold="true" box="[1095,1174,1701,1727]" pageId="9" pageNumber="10">
(
<figureCitation id="131E2A2D1701817F4B0AF905FB2B7DB1" box="[1106,1163,1701,1727]" captionStart="Figure 8" captionStartId="11.[113,178,1498,1522]" captionTargetBox="[114,1458,144,1471]" captionTargetId="figure-177@11.[114,1458,144,1471]" captionTargetPageId="11" captionText="Figure 8. Palaeabraeus glabrus gen.et sp. nov.. A, dorsal view.B, ventral view. C, mouthparts lateral view.D, head, prosternal process and mesoventrite ventral view.E, frontal view.F, lateral view.Abbreviations:ac, antennal cavity; apm, anterior margin of prosternal process; ees, elytral epipleural stria; ga, galea; lp, labial palpus; lps, lateral stria on prosternal process; ls, labral setae; mms, marginal mesoventral stria; mp, maxillary palpus; pf, prosternal fissures for receiving antennal funicle; pms, postmesocoxal stria; ppm, posterior margin of prosternal process." pageId="9">Fig. 8</figureCitation>
<figureCitation id="131E2A2D1701817F4B0AF905FB2B7DB1" box="[1106,1163,1701,1727]" captionStart="Figure 8" captionStartId="11.[113,178,1498,1522]" captionTargetBox="[114,1458,144,1471]" captionTargetId="figure-177@11.[114,1458,144,1471]" captionTargetPageId="11" captionText="Figure 8. Palaeabraeus glabrus gen.et sp. nov.. A, dorsal view.B, ventral view. C, mouthparts lateral view.D, head, prosternal process and mesoventrite ventral view.E, frontal view.F, lateral view.Abbreviations:ac, antennal cavity; apm, anterior margin of prosternal process; ees, elytral epipleural stria; ga, galea; lp, labial palpus; lps, lateral stria on prosternal process; ls, labral setae; mms, marginal mesoventral stria; mp, maxillary palpus; pf, prosternal fissures for receiving antennal funicle; pms, postmesocoxal stria; ppm, posterior margin of prosternal process." pageId="9" pageNumber="10">Fig. 8</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8B9A36A81701817F4C72F946FA9A7C33" blockId="9.[810,1460,1766,1853]" pageId="9">
<emphasis id="B951EABA1701817F4C72F946FC267DF0" box="[810,902,1766,1790]" italics="true" pageId="9">Material:</emphasis>
<collectingRegion id="49E1F84A1701817F4CD5F946FC777DF0" box="[909,983,1766,1790]" country="Myanmar" name="Kachin" pageId="9">Kachin</collectingRegion>
</subSubSection>
<subSubSection id="C33F65231701817F4C72F946FA9A7C33" pageId="9" pageNumber="10" type="materials_examined">
<paragraph id="8B9A36A81701817F4C72F946FA9A7C33" blockId="9.[810,1460,1766,1853]" pageId="9" pageNumber="10">
<emphasis id="B951EABA1701817F4C72F946FC267DF0" box="[810,902,1766,1790]" italics="true" pageId="9" pageNumber="10">Material:</emphasis>
<materialsCitation id="3B4D3CF51701817F4CD5F946FA957C33" collectionCode="HUM" location="Kachin" pageId="9" pageNumber="10" specimenCode="SEHU-0000121213" specimenCount="1" stateProvince="Kachin">
<collectingRegion id="49E1F84A1701817F4CD5F946FC777DF0" box="[909,983,1766,1790]" country="Myanmar" name="Kachin" pageId="9" pageNumber="10">Kachin</collectingRegion>
amber (
<emphasis id="B951EABA1701817F4B69F947FB9A7DF0" box="[1073,1082,1767,1790]" italics="true" pageId="9">c</emphasis>
<emphasis id="B951EABA1701817F4B69F947FB9A7DF0" box="[1073,1082,1767,1790]" italics="true" pageId="9" pageNumber="10">c</emphasis>
. 99 Mya). Amber piece dimensions: 5 × 2 ×
<quantity id="4CDD9B4D1701817F4C25F8A5FC187C13" box="[893,952,1797,1821]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="9" unit="mm" value="2.0">2 mm</quantity>
. Amber clear with several insect fragments and air bubbles. Deposited at HUM (ºEHU-0000121213).
<quantity id="4CDD9B4D1701817F4C25F8A5FC187C13" box="[893,952,1797,1821]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="10" unit="mm" value="2.0">2 mm</quantity>
. Amber clear with several insect fragments and air bubbles. Deposited at
<collectionCode id="ED34AE6D1701817F4B54F885FBEB7C32" box="[1036,1099,1829,1852]" pageId="9" pageNumber="10">HUM</collectionCode>
(
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)
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.
</paragraph>
<paragraph id="8B9A36A81701817A4C72F8FFFDEE7931" blockId="9.[810,1459,1886,1973]" lastBlockId="12.[129,778,144,575]" lastPageId="12" pageId="9">
<emphasis id="B951EABA1701817F4C72F8FFFC1C7C78" box="[810,956,1887,1910]" italics="true" pageId="9">Measurements:</emphasis>
</subSubSection>
<subSubSection id="C33F65231701817A4C72F8FFFDEE7931" lastPageId="12" lastPageNumber="13" pageId="9" pageNumber="10" type="description">
<paragraph id="8B9A36A81701817A4C72F8FFFDEE7931" blockId="9.[810,1459,1886,1973]" lastBlockId="12.[129,778,144,575]" lastPageId="12" lastPageNumber="13" pageId="9" pageNumber="10">
<emphasis id="B951EABA1701817F4C72F8FFFC1C7C78" box="[810,956,1887,1910]" italics="true" pageId="9" pageNumber="10">Measurements:</emphasis>
Head width:
<quantity id="4CDD9B4D1701817F4B0AF8FEFB0D7C7B" box="[1106,1197,1886,1910]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="9" unit="mm" value="0.23">0.23 mm</quantity>
<quantity id="4CDD9B4D1701817F4B0AF8FEFB0D7C7B" box="[1106,1197,1886,1910]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="9" pageNumber="10" unit="mm" value="0.23">0.23 mm</quantity>
; width between anterior pronotal angles:
<quantity id="4CDD9B4D1701817F4C86F8DDFB997C9B" box="[990,1081,1917,1941]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="9" unit="mm" value="0.25">0.25 mm</quantity>
<quantity id="4CDD9B4D1701817F4C86F8DDFB997C9B" box="[990,1081,1917,1941]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="9" pageNumber="10" unit="mm" value="0.25">0.25 mm</quantity>
; width between posterior pronotal angles:
<quantity id="4CDD9B4D1701817F4CDAF83CFC6F7CBA" box="[898,975,1948,1972]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="9" unit="mm" value="0.5">0.5 mm</quantity>
<quantity id="4CDD9B4D1701817F4CDAF83CFC6F7CBA" box="[898,975,1948,1972]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="10" unit="mm" value="0.5">0.5 mm</quantity>
; pronotal length:
<quantity id="4CDD9B4D1701817F4BF1F83DFAA47CBA" box="[1193,1284,1949,1973]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="9" unit="mm" value="0.23">0.23 mm</quantity>
<quantity id="4CDD9B4D1701817F4BF1F83DFAA47CBA" box="[1193,1284,1949,1973]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="9" pageNumber="10" unit="mm" value="0.23">0.23 mm</quantity>
; sutural elytral length:
<quantity id="4CDD9B4D1702817C4F8CF990FE827D49" box="[212,290,1584,1608]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="10" unit="mm" value="0.6">0.6 mm</quantity>
<quantity id="4CDD9B4D1702817C4F8CF990FE827D49" box="[212,290,1584,1608]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="11" unit="mm" value="0.6">0.6 mm</quantity>
; elytral width (in pair, across widest point):
<quantity id="4CDD9B4D1702817C4FD9F9EFFF7D7D69" box="[129,221,1615,1639]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" pageId="10" unit="mm" value="0.56">0.56 mm</quantity>
<quantity id="4CDD9B4D1702817C4FD9F9EFFF7D7D69" box="[129,221,1615,1639]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" pageId="10" pageNumber="11" unit="mm" value="0.56">0.56 mm</quantity>
.—
<emphasis id="B951EABA1702817C4FA4F9EFFE9C7D69" bold="true" box="[252,316,1615,1639]" italics="true" pageId="10">Body:</emphasis>
<emphasis id="B951EABA1702817C4FA4F9EFFE9C7D69" bold="true" box="[252,316,1615,1639]" italics="true" pageId="10" pageNumber="11">Body:</emphasis>
Oval, tapered anteriorly and posteriorly, strongly convex, impunctate, glabrous.—
<emphasis id="B951EABA1702817C4D75F9CEFDCF7D88" bold="true" box="[557,623,1646,1670]" italics="true" pageId="10">Head:</emphasis>
<emphasis id="B951EABA1702817C4D75F9CEFDCF7D88" bold="true" box="[557,623,1646,1670]" italics="true" pageId="10" pageNumber="11">Head:</emphasis>
Frons convex, weakly and sparsely punctate; frontal stria absent; narrow epistoma with fine, sparse setae; eye convex, coarsely faceted, anterior portion exposed laterally; labrum short, convex, bearing two dominant setae curving ventrad and several secondary setae; mandibles largely concealed, with a few setae on lateral margins; galeae exposed, densely setose; maxillary palps long, penultimate palpomere about third length of terminal; terminal palpomere swollen at base, tapered apically, shallowly concave on inner surface; labial palpi with apical palpomere long, slender, penultimate palpomere about half its length; antennal insertions slightly open anteriorly, under weak frontolateral bulge; scape short, weakly widening apically, curved posteriad below eye; pedicel cylindrical, almost as long as antennomeres 3 and 4 combined; antennomeres 3–8 widening slightly toward club, each with peripheral setae, 3–6 moniliform, 7 and 8 with sharper basal margins, each narrowed to apex; antennal club round, slightly tapered to apex, tomentose, without annuli.—
<emphasis id="B951EABA1702817C4A35F8ABFA627C2D" bold="true" box="[1389,1474,1803,1827]" italics="true" pageId="10">Ŋorax:</emphasis>
<emphasis id="B951EABA1702817C4A35F8ABFA627C2D" bold="true" box="[1389,1474,1803,1827]" italics="true" pageId="10" pageNumber="11">Thorax:</emphasis>
Pronotal disk convex, almost twice as wide as long, sides weakly curved, evenly tapered from base to apex; anterior emargination deep; anterior corners strongly downturned, acute; marginal pronotal stria present, distant from anterior margin, closer to lateral margins. Elytra strongly convex, apically truncate, glabrous, and impunctate, without distinct dorsal striae; epipleuron with single weak submarginal carina. Wings fully developed (protruding beyond elytra). Prosternum short, lacking distinct lobe; keel shallowly emarginate at base, with lateral striae narrowed at middle, widened to base and apex; prosternum incised on either side for reception of the antennal funicle; antennal cavities located anterolaterally, well defined by marginal ridge posteriorly and laterally. Mesoventrite rather long, more than half as long as intercoxal width, strongly developed anteriorly, with broadly acute median process inserting into base of prosternal keel; marginal mesoventral stria complete, broadly rounded; mesometaventral suture well developed; metaventrite slightly impressed along midline; postmesocoxal stria complete, strongly recurved behind coxa to mesepimeron; lateral metaventral stria not evident; metepisternum broadly exposed, tapered beneath anteriorly produced metepimeron.—
<emphasis id="B951EABA1703817D4BEDF893FB4D7C44" bold="true" box="[1205,1261,1843,1866]" italics="true" pageId="11">Legs:</emphasis>
Profemur slightly convex along inner margin, both inner and outer margins setose; protibia weakly curved, widened slightly to apex, setose and with one subapical denticle along outer margin; protarsal groove well developed; protibial spurs absent; mesofemur similar to profemur; mesotibia slender, widened slightly to apex, with fine setae along inner and outer margins, few stronger spines present near inner apical corners; metafemur and metatibia similar to those of mesothorax; tarsi 5-5-5, tarsomeres 1–4 similar in length, ultimate tarsomeres as long as tarsomeres 2–4 combined; protarsomeres 1–4 with expanded and flaưened ventral setae (probable sexual dimorphism); all tarsomeres with numerous ventral setae (not just one apicoventral pair); tarsal claws simple.—
<emphasis id="B951EABA1704817A4FBDFE09FEF17ACF" bold="true" box="[229,337,425,449]" italics="true" pageId="12">Abdomen:</emphasis>
<emphasis id="B951EABA1703817D4BEDF893FB4D7C44" bold="true" box="[1205,1261,1843,1866]" italics="true" pageId="11" pageNumber="12">Legs:</emphasis>
Profemur slightly convex along inner margin, both inner and outer margins setose; protibia weakly curved, widened slightly to apex, setose and with one subapical denticle along outer margin; protarsal groove well developed; protibial spurs absent; mesofemur similar to profemur; mesotibia slender, widened slightly to apex, with fine setae along inner and outer margins, few stronger spines present near inner apical corners; metafemur and metatibia similar to those of mesothorax; tarsi 5-5-5, tarsomeres 1–4 similar in length, ultimate tarsomeres as long as tarsomeres 2–4 combined; protarsomeres 1–4 with expanded and flattened ventral setae (probable sexual dimorphism); all tarsomeres with numerous ventral setae (not just one apicoventral pair); tarsal claws simple.—
<emphasis id="B951EABA1704817A4FBDFE09FEF17ACF" bold="true" box="[229,337,425,449]" italics="true" pageId="12" pageNumber="13">Abdomen:</emphasis>
Abdominal ventrite 1 flat between coxae, with slightly curved parallel striae from inner corner of metacoxa to ventrite 2; ventrites 2–5 short; propygidium shorter than pygidium, glabrous; pygidium slightly longer than basal width, weakly convex, apically rounded, impunctate.
</paragraph>
<caption id="DF5A66201702817C4FD9FA6CFAB07D0E" pageId="10" startId="10.[129,194,1484,1508]" targetBox="[129,1473,144,1456]" targetPageId="10" targetType="figure">
<paragraph id="8B9A36A81702817C4FD9FA6CFAB07D0E" blockId="10.[129,1382,1484,1536]" pageId="10">
<emphasis id="B951EABA1702817C4FD9FA6CFF787EEA" bold="true" box="[129,216,1484,1508]" pageId="10">Figure 7.</emphasis>
<taxonomicName id="4C254D2B1702817C4F85FA6DFE347EEA" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[221,404,1484,1508]" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="hirsutus" status="sp. nov.">
<emphasis id="B951EABA1702817C4F85FA6DFE347EEA" box="[221,404,1484,1508]" italics="true" pageId="10">Pantostictus hirsutus</emphasis>
</subSubSection>
<caption id="DF5A66201702817C4FD9FA6CFAB07D0E" pageId="10" pageNumber="11" startId="10.[129,194,1484,1508]" targetBox="[129,1473,144,1456]" targetPageId="10" targetType="figure">
<paragraph id="8B9A36A81702817C4FD9FA6CFAB07D0E" blockId="10.[129,1382,1484,1536]" pageId="10" pageNumber="11">
<emphasis id="B951EABA1702817C4FD9FA6CFF787EEA" bold="true" box="[129,216,1484,1508]" pageId="10" pageNumber="11">Figure 7.</emphasis>
<taxonomicName id="4C254D2B1702817C4F85FA6DFE347EEA" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[221,404,1484,1508]" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="hirsutus" status="sp. nov.">
<emphasis id="B951EABA1702817C4F85FA6DFE347EEA" box="[221,404,1484,1508]" italics="true" pageId="10" pageNumber="11">Pantostictus hirsutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C11702817C4EC2FA6CFE7B7EED" box="[410,475,1484,1507]" pageId="10" rank="species">sp. nov</taxonomicNameLabel>
<taxonomicNameLabel id="A26257C11702817C4EC2FA6CFE7B7EED" box="[410,475,1484,1507]" pageId="10" pageNumber="11" rank="species">sp. nov</taxonomicNameLabel>
.. A, dorsal view. B, ventral view. C, lateral view. D, elytral setation, E, dorsal view. F, metaventrite, abdomen. Abbreviations: ac, antennal club; as, antennal scape; ls, labral setae; pl, prosternal lobe; pmss, postmesocoxal stria.
</paragraph>
</caption>
<caption id="DF5A66201703817D4F29FA7AFA3D7D48" pageId="11" startId="11.[113,178,1498,1522]" targetBox="[114,1458,144,1471]" targetPageId="11" targetType="figure">
<paragraph id="8B9A36A81703817D4F29FA7AFA3D7D48" blockId="11.[113,1437,1498,1607]" pageId="11">
<emphasis id="B951EABA1703817D4F29FA7AFF687EFD" bold="true" box="[113,200,1498,1523]" pageId="11">Figure 8.</emphasis>
<taxonomicName id="4C254D2B1703817D4F95FA7BFE2B7EFD" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[205,395,1499,1523]" class="Insecta" family="Histeridae" genus="Palaeabraeus" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="glabrus" status="gen. et sp. nov">
<emphasis id="B951EABA1703817D4F95FA7BFE2B7EFD" box="[205,395,1499,1523]" italics="true" pageId="11">Palaeabraeus glabrus</emphasis>
<caption id="DF5A66201703817D4F29FA7AFA3D7D48" pageId="11" pageNumber="12" startId="11.[113,178,1498,1522]" targetBox="[114,1458,144,1471]" targetPageId="11" targetType="figure">
<paragraph id="8B9A36A81703817D4F29FA7AFA3D7D48" blockId="11.[113,1437,1498,1607]" pageId="11" pageNumber="12">
<emphasis id="B951EABA1703817D4F29FA7AFF687EFD" bold="true" box="[113,200,1498,1523]" pageId="11" pageNumber="12">Figure 8.</emphasis>
<taxonomicName id="4C254D2B1703817D4F95FA7BFE2B7EFD" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[205,395,1499,1523]" class="Insecta" family="Histeridae" genus="Palaeabraeus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="glabrus" status="gen. et sp. nov">
<emphasis id="B951EABA1703817D4F95FA7BFE2B7EFD" box="[205,395,1499,1523]" italics="true" pageId="11" pageNumber="12">Palaeabraeus glabrus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C11703817D4EC8FA7BFDB37EFC" box="[400,531,1499,1523]" pageId="11" rank="species">gen. et sp. nov</taxonomicNameLabel>
<taxonomicNameLabel id="A26257C11703817D4EC8FA7BFDB37EFC" box="[400,531,1499,1523]" pageId="11" pageNumber="12" rank="species">gen. et sp. nov</taxonomicNameLabel>
.. A, dorsal view. B, ventral view. C, mouthparts lateral view. D, head, prosternal process and mesoventrite ventral view. E, frontal view. F, lateral view. Abbreviations: ac, antennal cavity; apm, anterior margin of prosternal process; ees, elytral epipleural stria; ga, galea; lp, labial palpus; lps, lateral stria on prosternal process; ls, labral setae; mms, marginal mesoventral stria; mp, maxillary palpus; pf, prosternal fissures for receiving antennal funicle; pms, postmesocoxal stria; ppm, posterior margin of prosternal process.
</paragraph>
</caption>
<paragraph id="8B9A36A81704817A4FD9FDC1FDBB7EE8" blockId="12.[128,779,609,1510]" pageId="12">
<emphasis id="B951EABA1704817A4FD9FDC1FECA7976" box="[129,362,609,632]" italics="true" pageId="12">Taxonomic assignment:</emphasis>
Ŋe species can be assigned to the subfamily
<taxonomicName id="4C254D2B1704817A4F9DFD20FE8F7996" authorityName="W. S. MacLeay" authorityYear="1819" box="[197,303,640,664]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="subFamily" subFamily="Abraeinae">Abraeinae</taxonomicName>
<subSubSection id="C33F65231704817A4FD9FDC1FDBB7EE8" pageId="12" pageNumber="13" type="discussion">
<paragraph id="8B9A36A81704817A4FD9FDC1FDBB7EE8" blockId="12.[128,779,609,1510]" pageId="12" pageNumber="13">
<emphasis id="B951EABA1704817A4FD9FDC1FECA7976" box="[129,362,609,632]" italics="true" pageId="12" pageNumber="13">Taxonomic assignment:</emphasis>
The species can be assigned to the subfamily
<taxonomicName id="4C254D2B1704817A4F9DFD20FE8F7996" authorityName="W. S. MacLeay" authorityYear="1819" box="[197,303,640,664]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subFamily" subFamily="Abraeinae">Abraeinae</taxonomicName>
based on the lack of distinctive prosternal lobe (
<figureCitation id="131E2A2D1704817A4FD3FD3FFF5579B9" box="[139,245,671,695]" captionStart="Figure 8" captionStartId="11.[113,178,1498,1522]" captionTargetBox="[114,1458,144,1471]" captionTargetId="figure-177@11.[114,1458,144,1471]" captionTargetPageId="11" captionText="Figure 8. Palaeabraeus glabrus gen.et sp. nov.. A, dorsal view.B, ventral view. C, mouthparts lateral view.D, head, prosternal process and mesoventrite ventral view.E, frontal view.F, lateral view.Abbreviations:ac, antennal cavity; apm, anterior margin of prosternal process; ees, elytral epipleural stria; ga, galea; lp, labial palpus; lps, lateral stria on prosternal process; ls, labral setae; mms, marginal mesoventral stria; mp, maxillary palpus; pf, prosternal fissures for receiving antennal funicle; pms, postmesocoxal stria; ppm, posterior margin of prosternal process." pageId="12">Fig. 8B, D</figureCitation>
<figureCitation id="131E2A2D1704817A4FD3FD3FFF5579B9" box="[139,245,671,695]" captionStart="Figure 8" captionStartId="11.[113,178,1498,1522]" captionTargetBox="[114,1458,144,1471]" captionTargetId="figure-177@11.[114,1458,144,1471]" captionTargetPageId="11" captionText="Figure 8. Palaeabraeus glabrus gen.et sp. nov.. A, dorsal view.B, ventral view. C, mouthparts lateral view.D, head, prosternal process and mesoventrite ventral view.E, frontal view.F, lateral view.Abbreviations:ac, antennal cavity; apm, anterior margin of prosternal process; ees, elytral epipleural stria; ga, galea; lp, labial palpus; lps, lateral stria on prosternal process; ls, labral setae; mms, marginal mesoventral stria; mp, maxillary palpus; pf, prosternal fissures for receiving antennal funicle; pms, postmesocoxal stria; ppm, posterior margin of prosternal process." pageId="12" pageNumber="13">Fig. 8B, D</figureCitation>
), convex eyes (
<figureCitation id="131E2A2D1704817A4EC1FD3FFE5E79B9" box="[409,510,671,695]" captionStart="Figure 8" captionStartId="11.[113,178,1498,1522]" captionTargetBox="[114,1458,144,1471]" captionTargetId="figure-177@11.[114,1458,144,1471]" captionTargetPageId="11" captionText="Figure 8. Palaeabraeus glabrus gen.et sp. nov.. A, dorsal view.B, ventral view. C, mouthparts lateral view.D, head, prosternal process and mesoventrite ventral view.E, frontal view.F, lateral view.Abbreviations:ac, antennal cavity; apm, anterior margin of prosternal process; ees, elytral epipleural stria; ga, galea; lp, labial palpus; lps, lateral stria on prosternal process; ls, labral setae; mms, marginal mesoventral stria; mp, maxillary palpus; pf, prosternal fissures for receiving antennal funicle; pms, postmesocoxal stria; ppm, posterior margin of prosternal process." pageId="12">Fig. 8E, F</figureCitation>
<figureCitation id="131E2A2D1704817A4EC1FD3FFE5E79B9" box="[409,510,671,695]" captionStart="Figure 8" captionStartId="11.[113,178,1498,1522]" captionTargetBox="[114,1458,144,1471]" captionTargetId="figure-177@11.[114,1458,144,1471]" captionTargetPageId="11" captionText="Figure 8. Palaeabraeus glabrus gen.et sp. nov.. A, dorsal view.B, ventral view. C, mouthparts lateral view.D, head, prosternal process and mesoventrite ventral view.E, frontal view.F, lateral view.Abbreviations:ac, antennal cavity; apm, anterior margin of prosternal process; ees, elytral epipleural stria; ga, galea; lp, labial palpus; lps, lateral stria on prosternal process; ls, labral setae; mms, marginal mesoventral stria; mp, maxillary palpus; pf, prosternal fissures for receiving antennal funicle; pms, postmesocoxal stria; ppm, posterior margin of prosternal process." pageId="12" pageNumber="13">Fig. 8E, F</figureCitation>
) and fused antennal club (
<figureCitation id="131E2A2D1704817A4FD3FD1EFF5379D8" box="[139,243,702,727]" captionStart="Figure 8" captionStartId="11.[113,178,1498,1522]" captionTargetBox="[114,1458,144,1471]" captionTargetId="figure-177@11.[114,1458,144,1471]" captionTargetPageId="11" captionText="Figure 8. Palaeabraeus glabrus gen.et sp. nov.. A, dorsal view.B, ventral view. C, mouthparts lateral view.D, head, prosternal process and mesoventrite ventral view.E, frontal view.F, lateral view.Abbreviations:ac, antennal cavity; apm, anterior margin of prosternal process; ees, elytral epipleural stria; ga, galea; lp, labial palpus; lps, lateral stria on prosternal process; ls, labral setae; mms, marginal mesoventral stria; mp, maxillary palpus; pf, prosternal fissures for receiving antennal funicle; pms, postmesocoxal stria; ppm, posterior margin of prosternal process." pageId="12">Fig. 8A, B</figureCitation>
<figureCitation id="131E2A2D1704817A4FD3FD1EFF5379D8" box="[139,243,702,727]" captionStart="Figure 8" captionStartId="11.[113,178,1498,1522]" captionTargetBox="[114,1458,144,1471]" captionTargetId="figure-177@11.[114,1458,144,1471]" captionTargetPageId="11" captionText="Figure 8. Palaeabraeus glabrus gen.et sp. nov.. A, dorsal view.B, ventral view. C, mouthparts lateral view.D, head, prosternal process and mesoventrite ventral view.E, frontal view.F, lateral view.Abbreviations:ac, antennal cavity; apm, anterior margin of prosternal process; ees, elytral epipleural stria; ga, galea; lp, labial palpus; lps, lateral stria on prosternal process; ls, labral setae; mms, marginal mesoventral stria; mp, maxillary palpus; pf, prosternal fissures for receiving antennal funicle; pms, postmesocoxal stria; ppm, posterior margin of prosternal process." pageId="12" pageNumber="13">Fig. 8A, B</figureCitation>
).
<taxonomicName id="4C254D2B1704817A4E52FD1EFED479D8" authorityName="W. S. MacLeay" authorityYear="1819" box="[266,372,702,726]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="subFamily" subFamily="Abraeinae">Abraeinae</taxonomicName>
<taxonomicName id="4C254D2B1704817A4E52FD1EFED479D8" authorityName="W. S. MacLeay" authorityYear="1819" box="[266,372,702,726]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subFamily" subFamily="Abraeinae">Abraeinae</taxonomicName>
contain the following tribes:
<taxonomicName id="4C254D2B1704817A4DF1FD1EFCA579D8" authorityName="MacLeay" authorityYear="1819" box="[681,773,702,726]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="tribe" tribe="Abraeini">Abraeini</taxonomicName>
<taxonomicName id="4C254D2B1704817A4DF1FD1EFCA579D8" authorityName="MacLeay" authorityYear="1819" box="[681,773,702,726]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Abraeini">Abraeini</taxonomicName>
,
<taxonomicName id="4C254D2B1704817A4FD8FD7EFF7379F8" authorityName="Wenzel" authorityYear="1944" box="[128,211,734,758]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="tribe" tribe="Acritini">Acritini</taxonomicName>
<taxonomicName id="4C254D2B1704817A4FD8FD7EFF7379F8" authorityName="Wenzel" authorityYear="1944" box="[128,211,734,758]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Acritini">Acritini</taxonomicName>
,
<taxonomicName id="4C254D2B1704817A4F86FD7EFE2579F8" authorityName="Wenzel" authorityYear="1944" box="[222,389,734,758]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="tribe" tribe="Acritomorphini">Acritomorphini</taxonomicName>
<taxonomicName id="4C254D2B1704817A4F86FD7EFE2579F8" authorityName="Wenzel" authorityYear="1944" box="[222,389,734,758]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Acritomorphini">Acritomorphini</taxonomicName>
,
<taxonomicName id="4C254D2B1704817A4EC9FD7EFDAA79F8" authorityName="Portevin" authorityYear="1929" box="[401,522,734,758]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="tribe" tribe="Plegaderini">Plegaderini</taxonomicName>
<taxonomicName id="4C254D2B1704817A4EC9FD7EFDAA79F8" authorityName="Portevin" authorityYear="1929" box="[401,522,734,758]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Plegaderini">Plegaderini</taxonomicName>
,
<taxonomicName id="4C254D2B1704817A4D4DFD7EFDDA79F8" authorityName="Bickhardt" authorityYear="1914" box="[533,634,734,758]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="tribe" tribe="Teretriini">Teretriini</taxonomicName>
<taxonomicName id="4C254D2B1704817A4D4DFD7EFDDA79F8" authorityName="Bickhardt" authorityYear="1914" box="[533,634,734,758]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Teretriini">Teretriini</taxonomicName>
,
<taxonomicName id="4C254D2B1704817A4DDDFD7EFCA579F8" authorityName="Marseul" authorityYear="1857" box="[645,773,734,758]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="tribe" tribe="Trypanaeini">Trypanaeini</taxonomicName>
<taxonomicName id="4C254D2B1704817A4DDDFD7EFCA579F8" authorityName="Marseul" authorityYear="1857" box="[645,773,734,758]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Trypanaeini">Trypanaeini</taxonomicName>
,
<taxonomicName id="4C254D2B1704817A4FD8FD5DFF57781B" authorityName="Bickhardt" authorityYear="1913" box="[128,247,765,789]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="tribe" tribe="Trypeticini">Trypeticini</taxonomicName>
<taxonomicName id="4C254D2B1704817A4FD8FD5DFF57781B" authorityName="Bickhardt" authorityYear="1913" box="[128,247,765,789]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Trypeticini">Trypeticini</taxonomicName>
, and the extinct
<taxonomicName id="4C254D2B1704817A4EA2FD5DFD22781B" box="[506,642,765,789]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="tribe" tribe="Pantostictini">Pantostictini</taxonomicName>
<taxonomicName id="4C254D2B1704817A4EA2FD5DFD22781B" box="[506,642,765,789]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Pantostictini">Pantostictini</taxonomicName>
.
<taxonomicName id="4C254D2B1704817A4DF9FD5DFCA6781B" authorityName="Bickhardt" authorityYear="1914" box="[673,774,765,789]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="tribe" tribe="Teretriini">Teretriini</taxonomicName>
<taxonomicName id="4C254D2B1704817A4DF9FD5DFCA6781B" authorityName="Bickhardt" authorityYear="1914" box="[673,774,765,789]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Teretriini">Teretriini</taxonomicName>
,
<taxonomicName id="4C254D2B1704817A4FD8FCBCFEA1783A" authorityName="Marseul" authorityYear="1857" box="[128,257,796,820]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="tribe" tribe="Trypanaeini">Trypanaeini</taxonomicName>
<taxonomicName id="4C254D2B1704817A4FD8FCBCFEA1783A" authorityName="Marseul" authorityYear="1857" box="[128,257,796,820]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Trypanaeini">Trypanaeini</taxonomicName>
, and
<taxonomicName id="4C254D2B1704817A4E63FCBCFE10783A" authorityName="Bickhardt" authorityYear="1913" box="[315,432,796,820]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="tribe" tribe="Trypeticini">Trypeticini</taxonomicName>
<taxonomicName id="4C254D2B1704817A4E63FCBCFE10783A" authorityName="Bickhardt" authorityYear="1913" box="[315,432,796,820]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Trypeticini">Trypeticini</taxonomicName>
can be ruled out as they all represent groups with elongate cylindrical body shapes (
<bibRefCitation id="EFB44B591704817A4DE5FC9CFF15787D" author="Wenzel RL" pageId="12" pagination="51 - 151" refId="ref19054" refString="Wenzel RL. On the classification of the histerid beetles. Fieldiana Zoology 1944; 28: 51 - 151. hups: // doi. org / 10.1093 / aesa / 37.3.375" type="journal article" year="1944">Wenzel 1944</bibRefCitation>
<bibRefCitation id="EFB44B591704817A4DE5FC9CFF15787D" author="Wenzel RL" pageId="12" pageNumber="13" pagination="51 - 151" refId="ref19054" refString="Wenzel RL. On the classification of the histerid beetles. Fieldiana Zoology 1944; 28: 51 - 151. hups: // doi. org / 10.1093 / aesa / 37.3.375" type="journal article" year="1944">Wenzel 1944</bibRefCitation>
).
<taxonomicName id="4C254D2B1704817A4F8BFCFBFED8787D" authorityName="Wenzel" authorityYear="1944" box="[211,376,859,883]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="tribe" tribe="Acritomorphini">Acritomorphini</taxonomicName>
<taxonomicName id="4C254D2B1704817A4F8BFCFBFED8787D" authorityName="Wenzel" authorityYear="1944" box="[211,376,859,883]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Acritomorphini">Acritomorphini</taxonomicName>
have an elongate body shape, long prosternum, anteriorly positioned antennal cavities, and annulate antennal club (
<bibRefCitation id="EFB44B591704817A4E3FFC3AFE4F78BC" author="Wenzel RL" box="[359,495,922,946]" pageId="12" pagination="51 - 151" refId="ref19054" refString="Wenzel RL. On the classification of the histerid beetles. Fieldiana Zoology 1944; 28: 51 - 151. hups: // doi. org / 10.1093 / aesa / 37.3.375" type="journal article" year="1944">Wenzel 1944</bibRefCitation>
<bibRefCitation id="EFB44B591704817A4E3FFC3AFE4F78BC" author="Wenzel RL" box="[359,495,922,946]" pageId="12" pageNumber="13" pagination="51 - 151" refId="ref19054" refString="Wenzel RL. On the classification of the histerid beetles. Fieldiana Zoology 1944; 28: 51 - 151. hups: // doi. org / 10.1093 / aesa / 37.3.375" type="journal article" year="1944">Wenzel 1944</bibRefCitation>
,
<bibRefCitation id="EFB44B591704817A4EA4FC3AFDDB78BC" author="Mazur" box="[508,635,922,946]" pageId="12" pagination="27 - 32" refId="ref18506" refString="Mazur º. New species of histerid beetles (Coleoptera, Histeridae). Annales Zoologici 1997; 38: 27 - 32. hup: // rcin. org. pl" type="journal article" year="1997">Mazur 1997</bibRefCitation>
<bibRefCitation id="EFB44B591704817A4EA4FC3AFDDB78BC" author="Mazur" box="[508,635,922,946]" pageId="12" pageNumber="13" pagination="27 - 32" refId="ref18506" refString="Mazur º. New species of histerid beetles (Coleoptera, Histeridae). Annales Zoologici 1997; 38: 27 - 32. hup: // rcin. org. pl" type="journal article" year="1997">Mazur 1997</bibRefCitation>
).
<taxonomicName id="4C254D2B1704817A4DCBFC3AFCA978BC" authorityName="Portevin" authorityYear="1929" box="[659,777,922,946]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="tribe" tribe="Plegaderini">Plegaderini</taxonomicName>
have a flaưened body shape and pronotum with a pair of longitudinal grooves laterally, ossen with a transverse pronotal stria connecting the grooves (
<bibRefCitation id="EFB44B591704817A4ED0FC58FDB17F1E" author="Wenzel RL" box="[392,529,1016,1040]" pageId="12" pagination="51 - 151" refId="ref19054" refString="Wenzel RL. On the classification of the histerid beetles. Fieldiana Zoology 1944; 28: 51 - 151. hups: // doi. org / 10.1093 / aesa / 37.3.375" type="journal article" year="1944">Wenzel 1944</bibRefCitation>
<taxonomicName id="4C254D2B1704817A4DCBFC3AFCA978BC" authorityName="Portevin" authorityYear="1929" box="[659,777,922,946]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Plegaderini">Plegaderini</taxonomicName>
have a flattened body shape and pronotum with a pair of longitudinal grooves laterally, often with a transverse pronotal stria connecting the grooves (
<bibRefCitation id="EFB44B591704817A4ED0FC58FDB17F1E" author="Wenzel RL" box="[392,529,1016,1040]" pageId="12" pageNumber="13" pagination="51 - 151" refId="ref19054" refString="Wenzel RL. On the classification of the histerid beetles. Fieldiana Zoology 1944; 28: 51 - 151. hups: // doi. org / 10.1093 / aesa / 37.3.375" type="journal article" year="1944">Wenzel 1944</bibRefCitation>
).
<taxonomicName id="4C254D2B1704817A4D71FC58FD0E7F1E" box="[553,686,1016,1040]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="tribe" tribe="Pantostictini">Pantostictini</taxonomicName>
likewise have a slightly flaưened and rugosely punctuate body surface (
<bibRefCitation id="EFB44B591704817A4FD3FB96FE2E7F41" author="Poinar G &amp; Brown AE" box="[139,398,1078,1103]" pageId="12" pagination="38 - 46" refId="ref18750" refString="Poinar G, Brown AE. Pantostictus burmanicus, a new genus and species of Cretaceous beetles (Coleoptera: Hydrophiloidea: Histeridae) in Burmese amber. Proceedings of the Entomological Society of Washington 2009; 111: 38 - 46. hups: // doi. org / 10.4289 / 0013 - 8797 - 111.1.38" type="journal article" year="2009">Poinar and Brown 2009</bibRefCitation>
<taxonomicName id="4C254D2B1704817A4D71FC58FD0E7F1E" box="[553,686,1016,1040]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Pantostictini">Pantostictini</taxonomicName>
likewise have a slightly flattened and rugosely punctuate body surface (
<bibRefCitation id="EFB44B591704817A4FD3FB96FE2E7F41" author="Poinar G &amp; Brown AE" box="[139,398,1078,1103]" pageId="12" pageNumber="13" pagination="38 - 46" refId="ref18750" refString="Poinar G, Brown AE. Pantostictus burmanicus, a new genus and species of Cretaceous beetles (Coleoptera: Hydrophiloidea: Histeridae) in Burmese amber. Proceedings of the Entomological Society of Washington 2009; 111: 38 - 46. hups: // doi. org / 10.4289 / 0013 - 8797 - 111.1.38" type="journal article" year="2009">Poinar and Brown 2009</bibRefCitation>
).
<taxonomicName id="4C254D2B1704817A4EF0FB96FE587F40" authorityName="Wenzel" authorityYear="1944" box="[424,504,1078,1102]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="tribe" tribe="Acritini">Acritini</taxonomicName>
<taxonomicName id="4C254D2B1704817A4EF0FB96FE587F40" authorityName="Wenzel" authorityYear="1944" box="[424,504,1078,1102]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Acritini">Acritini</taxonomicName>
have their tarsal formula 5-5-4 (
<bibRefCitation id="EFB44B591704817A4F94FBF6FDE07F60" author="Kryzhanovskij OL &amp; Reichardt AN" box="[204,576,1110,1134]" pageId="12" refId="ref18111" refString="Kryzhanovskij OL, Reichardt AN. Zhuki Nadsemejstva Histeroidea (semejstva Sphaeritidae, Histeridae, Synteliidae) [in Russian]. Nauka, ed., Fauna ºººR, 1976." type="book" year="1976">Kryzhanovskij and Reichardt 1976</bibRefCitation>
). Ŋerefore, based on the round and convex body shape, fused antennal clubs and the 5-5-5 tarsal formula (
<figureCitation id="131E2A2D1704817A4ED0FB34FE4B7FA2" box="[392,491,1172,1197]" captionStart="Figure 8" captionStartId="11.[113,178,1498,1522]" captionTargetBox="[114,1458,144,1471]" captionTargetId="figure-177@11.[114,1458,144,1471]" captionTargetPageId="11" captionText="Figure 8. Palaeabraeus glabrus gen.et sp. nov.. A, dorsal view.B, ventral view. C, mouthparts lateral view.D, head, prosternal process and mesoventrite ventral view.E, frontal view.F, lateral view.Abbreviations:ac, antennal cavity; apm, anterior margin of prosternal process; ees, elytral epipleural stria; ga, galea; lp, labial palpus; lps, lateral stria on prosternal process; ls, labral setae; mms, marginal mesoventral stria; mp, maxillary palpus; pf, prosternal fissures for receiving antennal funicle; pms, postmesocoxal stria; ppm, posterior margin of prosternal process." pageId="12">Fig. 8B, F</figureCitation>
<bibRefCitation id="EFB44B591704817A4F94FBF6FDE07F60" author="Kryzhanovskij OL &amp; Reichardt AN" box="[204,576,1110,1134]" pageId="12" pageNumber="13" refId="ref18111" refString="Kryzhanovskij OL, Reichardt AN. Zhuki Nadsemejstva Histeroidea (semejstva Sphaeritidae, Histeridae, Synteliidae) [in Russian]. Nauka, ed., Fauna ºººR, 1976." type="book" year="1976">Kryzhanovskij and Reichardt 1976</bibRefCitation>
). Therefore, based on the round and convex body shape, fused antennal clubs and the 5-5-5 tarsal formula (
<figureCitation id="131E2A2D1704817A4ED0FB34FE4B7FA2" box="[392,491,1172,1197]" captionStart="Figure 8" captionStartId="11.[113,178,1498,1522]" captionTargetBox="[114,1458,144,1471]" captionTargetId="figure-177@11.[114,1458,144,1471]" captionTargetPageId="11" captionText="Figure 8. Palaeabraeus glabrus gen.et sp. nov.. A, dorsal view.B, ventral view. C, mouthparts lateral view.D, head, prosternal process and mesoventrite ventral view.E, frontal view.F, lateral view.Abbreviations:ac, antennal cavity; apm, anterior margin of prosternal process; ees, elytral epipleural stria; ga, galea; lp, labial palpus; lps, lateral stria on prosternal process; ls, labral setae; mms, marginal mesoventral stria; mp, maxillary palpus; pf, prosternal fissures for receiving antennal funicle; pms, postmesocoxal stria; ppm, posterior margin of prosternal process." pageId="12" pageNumber="13">Fig. 8B, F</figureCitation>
) the species may be placed in the
<taxonomicName id="4C254D2B1704817A4F9AFB14FEBC7FC2" authorityName="MacLeay" authorityYear="1819" box="[194,284,1204,1228]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="tribe" tribe="Abraeini">Abraeini</taxonomicName>
<taxonomicName id="4C254D2B1704817A4F9AFB14FEBC7FC2" authorityName="MacLeay" authorityYear="1819" box="[194,284,1204,1228]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Abraeini">Abraeini</taxonomicName>
tribe. Other characters supporting the position of the species within
<taxonomicName id="4C254D2B1704817A4E05FB73FE177FE5" authorityName="MacLeay" authorityYear="1819" box="[349,439,1235,1259]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="tribe" tribe="Abraeini">Abraeini</taxonomicName>
<taxonomicName id="4C254D2B1704817A4E05FB73FE177FE5" authorityName="MacLeay" authorityYear="1819" box="[349,439,1235,1259]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Abraeini">Abraeini</taxonomicName>
are the presence and position of antennal cavities (
<figureCitation id="131E2A2D1704817A4E1BFB52FE2D7E04" box="[323,397,1266,1290]" captionStart="Figure 8" captionStartId="11.[113,178,1498,1522]" captionTargetBox="[114,1458,144,1471]" captionTargetId="figure-177@11.[114,1458,144,1471]" captionTargetPageId="11" captionText="Figure 8. Palaeabraeus glabrus gen.et sp. nov.. A, dorsal view.B, ventral view. C, mouthparts lateral view.D, head, prosternal process and mesoventrite ventral view.E, frontal view.F, lateral view.Abbreviations:ac, antennal cavity; apm, anterior margin of prosternal process; ees, elytral epipleural stria; ga, galea; lp, labial palpus; lps, lateral stria on prosternal process; ls, labral setae; mms, marginal mesoventral stria; mp, maxillary palpus; pf, prosternal fissures for receiving antennal funicle; pms, postmesocoxal stria; ppm, posterior margin of prosternal process." pageId="12">Fig. 8F</figureCitation>
<figureCitation id="131E2A2D1704817A4E1BFB52FE2D7E04" box="[323,397,1266,1290]" captionStart="Figure 8" captionStartId="11.[113,178,1498,1522]" captionTargetBox="[114,1458,144,1471]" captionTargetId="figure-177@11.[114,1458,144,1471]" captionTargetPageId="11" captionText="Figure 8. Palaeabraeus glabrus gen.et sp. nov.. A, dorsal view.B, ventral view. C, mouthparts lateral view.D, head, prosternal process and mesoventrite ventral view.E, frontal view.F, lateral view.Abbreviations:ac, antennal cavity; apm, anterior margin of prosternal process; ees, elytral epipleural stria; ga, galea; lp, labial palpus; lps, lateral stria on prosternal process; ls, labral setae; mms, marginal mesoventral stria; mp, maxillary palpus; pf, prosternal fissures for receiving antennal funicle; pms, postmesocoxal stria; ppm, posterior margin of prosternal process." pageId="12" pageNumber="13">Fig. 8F</figureCitation>
), emarginate basal pronotal margin receiving anterior projection of mesoventrite (
<figureCitation id="131E2A2D1704817A4DDCFAB2FD587E27" box="[644,760,1298,1322]" captionStart="Figure 8" captionStartId="11.[113,178,1498,1522]" captionTargetBox="[114,1458,144,1471]" captionTargetId="figure-177@11.[114,1458,144,1471]" captionTargetPageId="11" captionText="Figure 8. Palaeabraeus glabrus gen.et sp. nov.. A, dorsal view.B, ventral view. C, mouthparts lateral view.D, head, prosternal process and mesoventrite ventral view.E, frontal view.F, lateral view.Abbreviations:ac, antennal cavity; apm, anterior margin of prosternal process; ees, elytral epipleural stria; ga, galea; lp, labial palpus; lps, lateral stria on prosternal process; ls, labral setae; mms, marginal mesoventral stria; mp, maxillary palpus; pf, prosternal fissures for receiving antennal funicle; pms, postmesocoxal stria; ppm, posterior margin of prosternal process." pageId="12">Fig. 8B, D</figureCitation>
<figureCitation id="131E2A2D1704817A4DDCFAB2FD587E27" box="[644,760,1298,1322]" captionStart="Figure 8" captionStartId="11.[113,178,1498,1522]" captionTargetBox="[114,1458,144,1471]" captionTargetId="figure-177@11.[114,1458,144,1471]" captionTargetPageId="11" captionText="Figure 8. Palaeabraeus glabrus gen.et sp. nov.. A, dorsal view.B, ventral view. C, mouthparts lateral view.D, head, prosternal process and mesoventrite ventral view.E, frontal view.F, lateral view.Abbreviations:ac, antennal cavity; apm, anterior margin of prosternal process; ees, elytral epipleural stria; ga, galea; lp, labial palpus; lps, lateral stria on prosternal process; ls, labral setae; mms, marginal mesoventral stria; mp, maxillary palpus; pf, prosternal fissures for receiving antennal funicle; pms, postmesocoxal stria; ppm, posterior margin of prosternal process." pageId="12" pageNumber="13">Fig. 8B, D</figureCitation>
). Close relation to
<taxonomicName id="4C254D2B1704817A4E6EFA91FE307E47" authorityName="MacLeay" authorityYear="1819" box="[310,400,1329,1353]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="tribe" tribe="Abraeini">Abraeini</taxonomicName>
was also indicated by all performed phylogenetic analyses. Ŋe main differences seưing this species apart from recent
<taxonomicName id="4C254D2B1704817A4ECEFAD0FE507E86" authorityName="MacLeay" authorityYear="1819" box="[406,496,1392,1416]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="tribe" tribe="Abraeini">Abraeini</taxonomicName>
<taxonomicName id="4C254D2B1704817A4E6EFA91FE307E47" authorityName="MacLeay" authorityYear="1819" box="[310,400,1329,1353]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Abraeini">Abraeini</taxonomicName>
was also indicated by all performed phylogenetic analyses. The main differences setting this species apart from recent
<taxonomicName id="4C254D2B1704817A4ECEFAD0FE507E86" authorityName="MacLeay" authorityYear="1819" box="[406,496,1392,1416]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Abraeini">Abraeini</taxonomicName>
genera (
<taxonomicName id="4C254D2B1704817A4D3AFAD0FCA57E86" authority="Leach" authorityName="Leach" authorityYear="1817" box="[610,773,1392,1416]" class="Insecta" family="Histeridae" genus="Abraeus" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA1704817A4D3AFAD0FD137E86" box="[610,691,1392,1416]" italics="true" pageId="12">Abraeus</emphasis>
<taxonomicName id="4C254D2B1704817A4D3AFAD0FCA57E86" authority="Leach" authorityName="Leach" authorityYear="1817" box="[610,773,1392,1416]" class="Insecta" family="Histeridae" genus="Abraeus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA1704817A4D3AFAD0FD137E86" box="[610,691,1392,1416]" italics="true" pageId="12" pageNumber="13">Abraeus</emphasis>
Leach
</taxonomicName>
,
<taxonomicName id="4C254D2B1704817A4FD8FA2FFEEE7EA8" authority="Moro" authorityName="Moro" authorityYear="1957" box="[128,334,1423,1447]" class="Insecta" family="Histeridae" genus="Spelaeabraeus" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA1704817A4FD8FA2FFEAC7EA9" box="[128,268,1423,1447]" italics="true" pageId="12">Spelaeabraeus</emphasis>
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<emphasis id="B951EABA1704817A4FD8FA2FFEAC7EA9" box="[128,268,1423,1447]" italics="true" pageId="12" pageNumber="13">Spelaeabraeus</emphasis>
Moro
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, and
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<emphasis id="B951EABA1704817A4EDAFA2FFDAA7EA9" box="[386,522,1423,1447]" italics="true" pageId="12">Chaetabraeus</emphasis>
<taxonomicName id="4C254D2B1704817A4EDAFA2FFDCD7EA9" authority="Portevin" authorityName="Portevin" authorityYear="1929" box="[386,621,1423,1447]" class="Insecta" family="Histeridae" genus="Chaetabraeus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA1704817A4EDAFA2FFDAA7EA9" box="[386,522,1423,1447]" italics="true" pageId="12" pageNumber="13">Chaetabraeus</emphasis>
Portevin
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) are: oval body shape, relatively long prosternal process, absence of punctuation, and only slightly dilated protibia.
</paragraph>
<paragraph id="8B9A36A81704817A4FD8F9A8FE787C56" blockId="12.[128,778,1543,1880]" pageId="12">
<emphasis id="B951EABA1704817A4FD8F9A8FF467D11" box="[128,230,1544,1567]" italics="true" pageId="12">Diagnosis:</emphasis>
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<subSubSection id="C33F65231704817A4FD8F9A8FE787C56" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="8B9A36A81704817A4FD8F9A8FE787C56" blockId="12.[128,778,1543,1880]" pageId="12" pageNumber="13">
<emphasis id="B951EABA1704817A4FD8F9A8FF467D11" box="[128,230,1544,1567]" italics="true" pageId="12" pageNumber="13">Diagnosis:</emphasis>
Absence of dorsal striae and punctuation. Marginal mesoventral stria complete, broadly rounded; mesometaventral suture well developed; metaventrite slightly impressed along midline; postmesocoxal stria complete, strongly recurved behind coxa to mesepimeron; lateral metaventral stria not evident. Only
<taxonomicName id="4C254D2B1704817A4E59F904FE7E7DB5" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[257,478,1700,1724]" class="Insecta" family="Histeridae" genus="Pseudacritus" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="species" species="extinctus">
<emphasis id="B951EABA1704817A4E59F904FE7E7DB5" box="[257,478,1700,1724]" italics="true" pageId="12">Pseudacritus extinctus</emphasis>
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<emphasis id="B951EABA1704817A4E59F904FE7E7DB5" box="[257,478,1700,1724]" italics="true" pageId="12" pageNumber="13">Pseudacritus extinctus</emphasis>
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, described in this study, is similar in general body shape and some characters to this species.
<taxonomicName id="4C254D2B1704817A4FE9F942FEDD7DF4" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[177,381,1762,1786]" class="Insecta" family="Histeridae" genus="Palaeabraeus" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="species" species="glabrus">
<emphasis id="B951EABA1704817A4FE9F942FEDD7DF4" box="[177,381,1762,1786]" italics="true" pageId="12">Palaeabraeus glabrus</emphasis>
<taxonomicName id="4C254D2B1704817A4FE9F942FEDD7DF4" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[177,381,1762,1786]" class="Insecta" family="Histeridae" genus="Palaeabraeus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="glabrus">
<emphasis id="B951EABA1704817A4FE9F942FEDD7DF4" box="[177,381,1762,1786]" italics="true" pageId="12" pageNumber="13">Palaeabraeus glabrus</emphasis>
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can be distinguished by the position of antennal cavities (mediolaterally as opposed to
<taxonomicName id="4C254D2B1704817A4D30F8A2FD757C17" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[616,725,1794,1817]" class="Insecta" family="Histeridae" genus="Pseudacritus" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="species" species="extinctus">
<emphasis id="B951EABA1704817A4D30F8A2FD757C17" box="[616,725,1794,1817]" italics="true" pageId="12">P. extinctus</emphasis>
<taxonomicName id="4C254D2B1704817A4D30F8A2FD757C17" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[616,725,1794,1817]" class="Insecta" family="Histeridae" genus="Pseudacritus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="extinctus">
<emphasis id="B951EABA1704817A4D30F8A2FD757C17" box="[616,725,1794,1817]" italics="true" pageId="12" pageNumber="13">P. extinctus</emphasis>
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with anteriorly positioned cavities), absence of antescutellar stria and complete absence of punctuation.
</paragraph>
<paragraph id="8B9A36A81704817A4FD8F8DAFF707CBF" blockId="12.[128,777,1914,1969]" pageId="12">
<emphasis id="B951EABA1704817A4FD8F8DAFF507C9C" box="[128,240,1914,1938]" italics="true" pageId="12">Etymology:</emphasis>
</subSubSection>
<subSubSection id="C33F65231704817A4FD8F8DAFF707CBF" pageId="12" pageNumber="13" type="etymology">
<paragraph id="8B9A36A81704817A4FD8F8DAFF707CBF" blockId="12.[128,777,1914,1969]" pageId="12" pageNumber="13">
<emphasis id="B951EABA1704817A4FD8F8DAFF507C9C" box="[128,240,1914,1938]" italics="true" pageId="12" pageNumber="13">Etymology:</emphasis>
Latin ‘glabrus’ refers to the smooth cuticle of the species.
</paragraph>
</subSubSection>

168
data/03/8C/87/038C87BE1701817F4CB4FBCEFC417D49.xml
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@ -1,120 +1,118 @@
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<heading id="D0D281C41701817F4CB4FBCEFB517F87" bold="true" box="[1004,1265,1134,1161]" centered="true" fontSize="11" level="2" pageId="9" reason="2">
<emphasis id="B951EABA1701817F4CB4FBCEFB517F87" bold="true" box="[1004,1265,1134,1161]" pageId="9">
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<emphasis id="B951EABA1701817F4CB4FBCEFB247F87" bold="true" box="[1004,1156,1134,1161]" italics="true" pageId="9">Palaeabraeus</emphasis>
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<heading id="D0D281C41701817F4CB4FBCEFB517F87" bold="true" box="[1004,1265,1134,1161]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="10" reason="2">
<emphasis id="B951EABA1701817F4CB4FBCEFB517F87" bold="true" box="[1004,1265,1134,1161]" pageId="9" pageNumber="10">
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<emphasis id="B951EABA1701817F4CB4FBCEFB247F87" bold="true" box="[1004,1156,1134,1161]" italics="true" pageId="9" pageNumber="10">Palaeabraeus</emphasis>
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<taxonomicNameLabel id="A26257C11701817F4BD1FBCFFB517F87" box="[1161,1265,1135,1161]" pageId="9" pageNumber="10" rank="genus">gen. nov.</taxonomicNameLabel>
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<emphasis id="B951EABA1701817F4C72FB03FBD57FB5" box="[810,1141,1187,1211]" italics="true" pageId="9">
<typeStatus id="549E880A1701817F4C72FB03FCFA7FB4" box="[810,858,1187,1210]" pageId="9">Type</typeStatus>
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<typeStatus id="549E880A1701817F4C72FB03FCFA7FB4" box="[810,858,1187,1210]" pageId="9" pageNumber="10">Type</typeStatus>
species.
<taxonomicName id="4C254D2B1701817F4CF0FB03FBD57FB5" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[936,1141,1187,1211]" class="Insecta" family="Histeridae" genus="Palaeabraeus" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="glabrus" status="sp. nov.">Palaeabraeus glabrus</taxonomicName>
<taxonomicName id="4C254D2B1701817F4CF0FB03FBD57FB5" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[936,1141,1187,1211]" class="Insecta" family="Histeridae" genus="Palaeabraeus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="glabrus" status="sp. nov.">Palaeabraeus glabrus</taxonomicName>
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<taxonomicNameLabel id="A26257C11701817F4B22FB03FB617FB4" box="[1146,1217,1187,1210]" pageId="9" rank="species">sp. nov</taxonomicNameLabel>
<taxonomicNameLabel id="A26257C11701817F4B22FB03FB617FB4" box="[1146,1217,1187,1210]" pageId="9" pageNumber="10" rank="species">sp. nov</taxonomicNameLabel>
..
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<emphasis id="B951EABA1701817F4C72FB7DFC307FFA" box="[810,912,1245,1268]" italics="true" pageId="9">Diagnosis:</emphasis>
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<paragraph id="8B9A36A81701817F4C72FB7DFAFA7EC1" blockId="9.[810,1459,1244,1488]" pageId="9" pageNumber="10">
<emphasis id="B951EABA1701817F4C72FB7DFC307FFA" box="[810,912,1245,1268]" italics="true" pageId="9" pageNumber="10">Diagnosis:</emphasis>
Body oval, strongly dorsally convex. Head without frontal stria, short labrum with a pair of prominent setae. Body surface glabrous, without punctuation. Widely open antennal cavities present at the anterolateral position on the prosternal hypomeron. Mesoventrite anteriorly with a round apex inserted into the base of the prosternal keel, with a curved postcoxal striae and complete lateral striae. Metaventrite long, with postmesocoxal striae. Legs simple, tarsal formula 5-5-5.
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<emphasis id="B951EABA1701817F4C72FA51FC397D07" box="[810,921,1521,1545]" italics="true" pageId="9">Etymology:</emphasis>
Ŋe genus is considered as an ancient representative of the
<taxonomicName id="4C254D2B1701817F4C28F9B0FC687D26" authorityName="MacLeay" authorityYear="1819" box="[880,968,1552,1576]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="tribe" tribe="Abraeini">Abraeini</taxonomicName>
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<paragraph id="8B9A36A81701817F4C72FA51FC417D49" blockId="9.[810,1459,1521,1607]" pageId="9" pageNumber="10">
<emphasis id="B951EABA1701817F4C72FA51FC397D07" box="[810,921,1521,1545]" italics="true" pageId="9" pageNumber="10">Etymology:</emphasis>
The genus is considered as an ancient representative of the
<taxonomicName id="4C254D2B1701817F4C28F9B0FC687D26" authorityName="MacLeay" authorityYear="1819" box="[880,968,1552,1576]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="tribe" tribe="Abraeini">Abraeini</taxonomicName>
tribe and the generic name refers to this taxonomic placement.
</paragraph>
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295
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<mods:namePart id="427F9751CA56FCF236E1E00E486ECCAD">Prokop, Jakub</mods:namePart>
<mods:affiliation id="CC87BF2C7FB41374ED83BBA169DAB3D0">Charles University, Faculty of ºcience, Department of Zoology, Viničná 7, 12800 Prague 2, Czech Republic</mods:affiliation>
<mods:namePart id="F847AC31A7C750DDB732C914B565606B">Caterino, Michael S</mods:namePart>
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<mods:namePart id="964549B8516FA78A636074FF045DF9FF">Michael</mods:namePart>
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<mods:namePart id="92F43F65E91052C3A6EA828E9CCE8B3A">Caterino</mods:namePart>
<mods:affiliation id="94FB8F182B808BB2EB7A033978F2E4C3">Department of Plant and Environmental ºciences, Clemson University, Clemson, ºC 29634, United ºtates</mods:affiliation>
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<mods:title id="A4BF08D44E08768F06CD2BDB7D5DF875">Zoological Journal of the Linnean Society</mods:title>
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<mods:title id="9EAFB51E711B27A3DE9DC41922ED27F8">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="BFB4EEDA90F9DE366B90BA4DDFC338DA">
<mods:date id="8E683BEDA73F971953F33C0A2C49F7CB">2024</mods:date>
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<mods:number id="B86995A456BBF04346BA8D72F6B05490">2024-11-20</mods:number>
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<mods:date id="A3D1DD4F2D49824063A63115DA9D182A">2024</mods:date>
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<mods:title id="946FA29C4619C58454A0CB21388AD5E6">zlae 137</mods:title>
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<mods:detail id="11E5489EEA63D15583BE883A13C9FB86" type="volume">
<mods:number id="0C8A88537D0D04D6C338B92AACE2EB61">202</mods:number>
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<mods:number id="9BC5CA2574E400709388ACC56AF55EBB">3</mods:number>
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<mods:number id="C722233A5235582304A48F6CD8737AFA">202</mods:number>
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<mods:number id="4560D0108C7F51731174CBC9769D981E">3</mods:number>
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<mods:url id="7001C2C8A0B6F61273F9E66565EA57D8">https://doi.org/10.1093/zoolinnean/zlae137</mods:url>
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<mods:identifier id="9FF88DDFA375A0E5E1C337A0BEE97440" type="ISSN">0024-4082</mods:identifier>
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<treatment id="038C87BE1701817F4E4AFE09FC6078E5" ID-DOI="http://doi.org/10.5281/zenodo.14424550" ID-Zenodo-Dep="14424550" LSID="urn:lsid:plazi:treatment:038C87BE1701817F4E4AFE09FC6078E5" httpUri="http://treatment.plazi.org/id/038C87BE1701817F4E4AFE09FC6078E5" pageId="9">
<subSubSection id="C33F65231701817F4E4AFE09FDF97ACA" box="[274,601,425,452]" pageId="9" type="nomenclature">
<paragraph id="8B9A36A81701817F4E4AFE09FDF97ACA" blockId="9.[274,601,425,452]" box="[274,601,425,452]" pageId="9">
<heading id="D0D281C41701817F4E4AFE09FDF97ACA" bold="true" box="[274,601,425,452]" centered="true" fontSize="11" level="2" pageId="9" reason="2">
<emphasis id="B951EABA1701817F4E4AFE09FDF97ACA" bold="true" box="[274,601,425,452]" pageId="9">
<taxonomicName id="4C254D2B1701817F4E4AFE09FE5D7ACA" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[274,509,425,452]" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="hirsutus" status="sp. nov.">
<emphasis id="B951EABA1701817F4E4AFE09FE5D7ACA" bold="true" box="[274,509,425,452]" italics="true" pageId="9">Pantostictus hirsutus</emphasis>
<treatment id="038C87BE1701817F4E4AFE09FC6078E5" ID-DOI="http://doi.org/10.5281/zenodo.14424550" ID-Zenodo-Dep="14424550" LSID="urn:lsid:plazi:treatment:038C87BE1701817F4E4AFE09FC6078E5" httpUri="http://treatment.plazi.org/id/038C87BE1701817F4E4AFE09FC6078E5" lastPageNumber="10" pageId="9" pageNumber="10">
<subSubSection id="C33F65231701817F4E4AFE09FDF97ACA" box="[274,601,425,452]" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="8B9A36A81701817F4E4AFE09FDF97ACA" blockId="9.[274,601,425,452]" box="[274,601,425,452]" pageId="9" pageNumber="10">
<heading id="D0D281C41701817F4E4AFE09FDF97ACA" bold="true" box="[274,601,425,452]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="10" reason="2">
<emphasis id="B951EABA1701817F4E4AFE09FDF97ACA" bold="true" box="[274,601,425,452]" pageId="9" pageNumber="10">
<taxonomicName id="4C254D2B1701817F4E4AFE09FE5D7ACA" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[274,509,425,452]" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="hirsutus" status="sp. nov.">
<emphasis id="B951EABA1701817F4E4AFE09FE5D7ACA" bold="true" box="[274,509,425,452]" italics="true" pageId="9" pageNumber="10">Pantostictus hirsutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C11701817F4D5AFE0AFDF97ACA" box="[514,601,426,452]" pageId="9" rank="species">sp. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="A26257C11701817F4D5AFE0AFDF97ACA" box="[514,601,426,452]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33F65231701817F4ED7FE7BFC6078E5" pageId="9" type="description">
<paragraph id="8B9A36A81701817F4ED7FE7BFE7D7AFB" blockId="9.[399,477,475,502]" box="[399,477,475,502]" pageId="9">
<emphasis id="B951EABA1701817F4ED7FE7BFE7D7AFB" bold="true" box="[399,477,475,502]" pageId="9">
<subSubSection id="C33F65231701817F4ED7FE7BFC6078E5" pageId="9" pageNumber="10" type="description">
<paragraph id="8B9A36A81701817F4ED7FE7BFE7D7AFB" blockId="9.[399,477,475,502]" box="[399,477,475,502]" pageId="9" pageNumber="10">
<emphasis id="B951EABA1701817F4ED7FE7BFE7D7AFB" bold="true" box="[399,477,475,502]" pageId="9" pageNumber="10">
(
<figureCitation id="131E2A2D1701817F4EC1FE7BFE727AF8" box="[409,466,475,502]" captionStart="Figure 7" captionStartId="10.[129,194,1484,1508]" captionTargetBox="[129,1473,144,1456]" captionTargetId="figure-220@10.[129,1473,144,1456]" captionTargetPageId="10" captionText="Figure 7. Pantostictus hirsutus sp. nov.. A, dorsal view. B, ventral view.C, lateral view. D, elytral setation, E, dorsal view. F, metaventrite, abdomen.Abbreviations:ac, antennal club; as, antennal scape; ls, labral setae; pl, prosternal lobe; pmss, postmesocoxal stria." pageId="9">Fig. 7</figureCitation>
<figureCitation id="131E2A2D1701817F4EC1FE7BFE727AF8" box="[409,466,475,502]" captionStart="Figure 7" captionStartId="10.[129,194,1484,1508]" captionTargetBox="[129,1473,144,1456]" captionTargetId="figure-220@10.[129,1473,144,1456]" captionTargetPageId="10" captionText="Figure 7. Pantostictus hirsutus sp. nov.. A, dorsal view. B, ventral view.C, lateral view. D, elytral setation, E, dorsal view. F, metaventrite, abdomen.Abbreviations:ac, antennal club; as, antennal scape; ls, labral setae; pl, prosternal lobe; pmss, postmesocoxal stria." pageId="9" pageNumber="10">Fig. 7</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8B9A36A81701817F4F29FDBCFDCD797D" blockId="9.[113,763,540,627]" pageId="9">
<emphasis id="B951EABA1701817F4F29FDBCFF6D793A" box="[113,205,540,564]" italics="true" pageId="9">Material:</emphasis>
<collectingRegion id="49E1F84A1701817F4F8BFDBCFEBD793A" box="[211,285,540,564]" country="Myanmar" name="Kachin" pageId="9">Kachin</collectingRegion>
<paragraph id="8B9A36A81701817F4F29FDBCFDCD797D" blockId="9.[113,763,540,627]" pageId="9" pageNumber="10">
<emphasis id="B951EABA1701817F4F29FDBCFF6D793A" box="[113,205,540,564]" italics="true" pageId="9" pageNumber="10">Material:</emphasis>
<materialsCitation id="3B4D3CF51701817F4F8BFDBCFDC9797D" collectionCode="HUM" location="Kachin" pageId="9" pageNumber="10" specimenCode="SEHU-121212" specimenCount="1" stateProvince="Kachin">
<collectingRegion id="49E1F84A1701817F4F8BFDBCFEBD793A" box="[211,285,540,564]" country="Myanmar" name="Kachin" pageId="9" pageNumber="10">Kachin</collectingRegion>
amber (c
<emphasis id="B951EABA1701817F4E26FDBDFE27793A" box="[382,391,541,564]" italics="true" pageId="9">c</emphasis>
<emphasis id="B951EABA1701817F4E26FDBDFE27793A" box="[382,391,541,564]" italics="true" pageId="9" pageNumber="10">c</emphasis>
. 99 Mya). Amber piece dimensions: 12 × 7 ×
<quantity id="4CDD9B4D1701817F4F8AFD9BFEAD795D" box="[210,269,571,595]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="9" unit="mm" value="1.0">1 mm</quantity>
. Amber clear with several insect fragments and air bubbles. Deposited at HUM (ºEHU-121212).
<quantity id="4CDD9B4D1701817F4F8AFD9BFEAD795D" box="[210,269,571,595]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="10" unit="mm" value="1.0">1 mm</quantity>
. Amber clear with several insect fragments and air bubbles. Deposited at
<collectionCode id="ED34AE6D1701817F4E2BFDFBFE12797C" box="[371,434,603,626]" pageId="9" pageNumber="10">HUM</collectionCode>
(
<specimenCode id="DB839ED31701817F4EE7FDFBFDC1797D" box="[447,609,603,627]" pageId="9" pageNumber="10">SEHU-121212</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B9A36A81701817F4F29FD35FD127D98" blockId="9.[113,763,660,1686]" pageId="9">
<emphasis id="B951EABA1701817F4F29FD35FEA479A2" box="[113,260,661,684]" italics="true" pageId="9">Measurements:</emphasis>
<paragraph id="8B9A36A81701817F4F29FD35FD127D98" blockId="9.[113,763,660,1686]" pageId="9" pageNumber="10">
<emphasis id="B951EABA1701817F4F29FD35FEA479A2" box="[113,260,661,684]" italics="true" pageId="9" pageNumber="10">Measurements:</emphasis>
Head width:
<quantity id="4CDD9B4D1701817F4EC1FD34FE5479A5" box="[409,500,660,684]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="9" unit="mm" value="0.25">0.25 mm</quantity>
<quantity id="4CDD9B4D1701817F4EC1FD34FE5479A5" box="[409,500,660,684]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="9" pageNumber="10" unit="mm" value="0.25">0.25 mm</quantity>
; width between anterior pronotal angles:
<quantity id="4CDD9B4D1701817F4E7DFD13FE2079C5" box="[293,384,691,715]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="9" unit="mm" value="0.35">0.35 mm</quantity>
<quantity id="4CDD9B4D1701817F4E7DFD13FE2079C5" box="[293,384,691,715]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="9" pageNumber="10" unit="mm" value="0.35">0.35 mm</quantity>
; width between posterior pronotal angles:
<quantity id="4CDD9B4D1701817F4F9FFD72FE8279E4" box="[199,290,722,746]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="9" unit="mm" value="0.55">0.55 mm</quantity>
<quantity id="4CDD9B4D1701817F4F9FFD72FE8279E4" box="[199,290,722,746]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="9" pageNumber="10" unit="mm" value="0.55">0.55 mm</quantity>
; pronotal length:
<quantity id="4CDD9B4D1701817F4EADFD72FDEF79E4" box="[501,591,722,746]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="9" unit="mm" value="0.35">0.35 mm</quantity>
<quantity id="4CDD9B4D1701817F4EADFD72FDEF79E4" box="[501,591,722,746]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="9" pageNumber="10" unit="mm" value="0.35">0.35 mm</quantity>
; sutural elytral length:
<quantity id="4CDD9B4D1701817F4F9BFD52FEBE7807" box="[195,286,754,778]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="9" unit="mm" value="0.35">0.35 mm</quantity>
<quantity id="4CDD9B4D1701817F4F9BFD52FEBE7807" box="[195,286,754,778]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="9" pageNumber="10" unit="mm" value="0.35">0.35 mm</quantity>
; elytral width (in pair, across widest point):
<quantity id="4CDD9B4D1701817F4F29FCB1FF6E7827" box="[113,206,785,809]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="9" unit="mm" value="0.65">0.65 mm</quantity>
<quantity id="4CDD9B4D1701817F4F29FCB1FF6E7827" box="[113,206,785,809]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="9" pageNumber="10" unit="mm" value="0.65">0.65 mm</quantity>
.—
<emphasis id="B951EABA1701817F4FB5FCB1FE8D7827" bold="true" box="[237,301,785,809]" italics="true" pageId="9">Body:</emphasis>
<emphasis id="B951EABA1701817F4FB5FCB1FE8D7827" bold="true" box="[237,301,785,809]" italics="true" pageId="9" pageNumber="10">Body:</emphasis>
Oval with a densely and coarsely punctuated cuticle. Dorsal surface with sparse short setation.—
<emphasis id="B951EABA1701817F4D21FC90FD1A7846" bold="true" box="[633,698,816,840]" italics="true" pageId="9">Head:</emphasis>
Frons and clypeus flat, coarsely punctuate, without striae. Labrum short and wide, rounded apically, plurisetose. Less mandible appearing bidentate. Antennal club flat and triangular (most likely deformed).—
<emphasis id="B951EABA1701817F4E19FC0EFE3578C8" bold="true" box="[321,405,942,966]" italics="true" pageId="9">Ŋorax:</emphasis>
Pronotal disc coarsely punctuate, broadly rounded basally, anterior angles pointed. Lateral margin finely crenulate. ºcutellar shield hidden. Elytra expanded laterally in basal third, surface of elytral disc coarsely punctuate and finely setose, without striae. Epipleura punctate, but not striate. Prosternum short and wide. Prosternal process widely rectangular anteriorly, coarsely punctuated with crenulate margin and lateral fissures for antennal passage. Antennal clubs rest in shallow cavities located in anterior prosternal angles. Prosternal lobe not distinct. Meso- and metaventrite coarsely punctuated. Metaventrite with recurved postcoxal striae.—
<emphasis id="B951EABA1701817F4F29FAA7FF097E10" bold="true" box="[113,169,1287,1310]" italics="true" pageId="9">Legs:</emphasis>
Profemur flaưened, with a groove for protibial reception. Protibia slightly expanding apically, outer margin rounded in apical half, bearing few short spines; protibial apex with a small tibial spur. Protarsomeres short, terminal as long as 3–4 combined. Middle leg similar to front leg. Metafemur swollen with setae dorsally, metatibia slightly expanded apically, with several small setae along outer margin increasing in size apically and with a tibial spur. Tarsomeres short, terminal tarsomere as long as tarsomeres 3–4 combined; tarsomeres 1–4 bearing single pair of apicoventral spines.—
<emphasis id="B951EABA1701817F4E25F980FE4A7D36" bold="true" box="[381,490,1568,1592]" italics="true" pageId="9">Abdomen:</emphasis>
<emphasis id="B951EABA1701817F4D21FC90FD1A7846" bold="true" box="[633,698,816,840]" italics="true" pageId="9" pageNumber="10">Head:</emphasis>
Frons and clypeus flat, coarsely punctuate, without striae. Labrum short and wide, rounded apically, plurisetose. Left mandible appearing bidentate. Antennal club flat and triangular (most likely deformed).—
<emphasis id="B951EABA1701817F4E19FC0EFE3578C8" bold="true" box="[321,405,942,966]" italics="true" pageId="9" pageNumber="10">Thorax:</emphasis>
Pronotal disc coarsely punctuate, broadly rounded basally, anterior angles pointed. Lateral margin finely crenulate. Scutellar shield hidden. Elytra expanded laterally in basal third, surface of elytral disc coarsely punctuate and finely setose, without striae. Epipleura punctate, but not striate. Prosternum short and wide. Prosternal process widely rectangular anteriorly, coarsely punctuated with crenulate margin and lateral fissures for antennal passage. Antennal clubs rest in shallow cavities located in anterior prosternal angles. Prosternal lobe not distinct. Meso- and metaventrite coarsely punctuated. Metaventrite with recurved postcoxal striae.—
<emphasis id="B951EABA1701817F4F29FAA7FF097E10" bold="true" box="[113,169,1287,1310]" italics="true" pageId="9" pageNumber="10">Legs:</emphasis>
Profemur flattened, with a groove for protibial reception. Protibia slightly expanding apically, outer margin rounded in apical half, bearing few short spines; protibial apex with a small tibial spur. Protarsomeres short, terminal as long as 3–4 combined. Middle leg similar to front leg. Metafemur swollen with setae dorsally, metatibia slightly expanded apically, with several small setae along outer margin increasing in size apically and with a tibial spur. Tarsomeres short, terminal tarsomere as long as tarsomeres 3–4 combined; tarsomeres 1–4 bearing single pair of apicoventral spines.—
<emphasis id="B951EABA1701817F4E25F980FE4A7D36" bold="true" box="[381,490,1568,1592]" italics="true" pageId="9" pageNumber="10">Abdomen:</emphasis>
First abdominal ventrite punctuated with the same coarseness as metaventrite. Other visible abdominal sclerites (incl. pygidium) with finer punctuation. Propygidium short and wide, largely concealed by elytra.
</paragraph>
<paragraph id="8B9A36A81701817F4F29F961FBB87AEF" blockId="9.[113,763,1728,1972]" lastBlockId="9.[810,1463,144,481]" pageId="9">
<emphasis id="B951EABA1701817F4F29F961FEF77DD6" box="[113,343,1729,1752]" italics="true" pageId="9">Taxonomic assignment:</emphasis>
<paragraph id="8B9A36A81701817F4F29F961FBB87AEF" blockId="9.[113,763,1728,1972]" lastBlockId="9.[810,1463,144,481]" pageId="9" pageNumber="10">
<emphasis id="B951EABA1701817F4F29F961FEF77DD6" box="[113,343,1729,1752]" italics="true" pageId="9" pageNumber="10">Taxonomic assignment:</emphasis>
We assign this new species to the genus
<taxonomicName id="4C254D2B1701817F4F29F940FF457DF9" authorityName="Poinar and Brown" authorityYear="2009" box="[113,229,1760,1783]" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA1701817F4F29F940FF457DF9" box="[113,229,1760,1783]" italics="true" pageId="9">Pantostictus</emphasis>
<taxonomicName id="4C254D2B1701817F4F29F940FF457DF9" authorityName="Poinar and Brown" authorityYear="2009" box="[113,229,1760,1783]" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA1701817F4F29F940FF457DF9" box="[113,229,1760,1783]" italics="true" pageId="9" pageNumber="10">Pantostictus</emphasis>
</taxonomicName>
(in the monotypic Pantosticini;
<bibRefCitation id="EFB44B591701817F4D19F940FD4F7DF6" author="Zhou Y &amp; Ren D" box="[577,751,1760,1784]" pageId="9" pagination="521 - 39" refId="ref19290" refString="Zhou Y, Caterino Mº, Ren D et al. Phylogeny and evolution of Mesozoic and extant lineages of Histeridae (Coleoptera), with discovery of a new subfamily Antigracilinae from the Lower Cretaceous. Cladistics 2020; 36: 521 - 39. hups: // doi. org / 10.1111 / cla. 12418" type="journal article" year="2020">
<bibRefCitation id="EFB44B591701817F4D19F940FD4F7DF6" author="Zhou Y &amp; Ren D" box="[577,751,1760,1784]" pageId="9" pageNumber="10" pagination="521 - 39" refId="ref19290" refString="Zhou Y, Caterino Mº, Ren D et al. Phylogeny and evolution of Mesozoic and extant lineages of Histeridae (Coleoptera), with discovery of a new subfamily Antigracilinae from the Lower Cretaceous. Cladistics 2020; 36: 521 - 39. hups: // doi. org / 10.1111 / cla. 12418" type="journal article" year="2020">
Zhou
<emphasis id="B951EABA1701817F4DDAF940FD147DF6" box="[642,692,1760,1784]" italics="true" pageId="9">et al.</emphasis>
<emphasis id="B951EABA1701817F4DDAF940FD147DF6" box="[642,692,1760,1784]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
2020
</bibRefCitation>
) based on the ventrally flat and oval body shape, broad and rectangular apex of prosternal process, and the coarse punctuation. Ŋe genus was further characterized by
<bibRefCitation id="EFB44B591701817F4D6AF89EFF1C7C7B" author="Poinar G &amp; Brown AE" pageId="9" pagination="38 - 46" refId="ref18750" refString="Poinar G, Brown AE. Pantostictus burmanicus, a new genus and species of Cretaceous beetles (Coleoptera: Hydrophiloidea: Histeridae) in Burmese amber. Proceedings of the Entomological Society of Washington 2009; 111: 38 - 46. hups: // doi. org / 10.4289 / 0013 - 8797 - 111.1.38" type="journal article" year="2009">Poinar and Brown (2009)</bibRefCitation>
) based on the ventrally flat and oval body shape, broad and rectangular apex of prosternal process, and the coarse punctuation. The genus was further characterized by
<bibRefCitation id="EFB44B591701817F4D6AF89EFF1C7C7B" author="Poinar G &amp; Brown AE" pageId="9" pageNumber="10" pagination="38 - 46" refId="ref18750" refString="Poinar G, Brown AE. Pantostictus burmanicus, a new genus and species of Cretaceous beetles (Coleoptera: Hydrophiloidea: Histeridae) in Burmese amber. Proceedings of the Entomological Society of Washington 2009; 111: 38 - 46. hups: // doi. org / 10.4289 / 0013 - 8797 - 111.1.38" type="journal article" year="2009">Poinar and Brown (2009)</bibRefCitation>
as having an acute, triangular-shaped antennal club, and by lacking postmesocoxal and ventral abdominal striae. Despite appearing somewhat similar in the sole specimen of this new species, we believe that the odd antennal club shape is a postmortem deformity caused by collapse of a poorly sclerotized structure, as its exact shape varies considerably among available specimens (compare, e.g. Poinar and Brown’s figs 2 vs. 3). Further, although the tribal separation of
<taxonomicName id="4C254D2B1701817F4A6AFEADFA137A2B" box="[1330,1459,269,293]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="tribe" tribe="Pantostictini">Pantostictini</taxonomicName>
<taxonomicName id="4C254D2B1701817F4A6AFEADFA137A2B" box="[1330,1459,269,293]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="tribe" tribe="Pantostictini">Pantostictini</taxonomicName>
from
<taxonomicName id="4C254D2B1701817F4C3EFE8CFC7A7A4A" authorityName="Portevin" authorityYear="1929" box="[870,986,300,324]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="tribe" tribe="Plegaderini">Plegaderini</taxonomicName>
<taxonomicName id="4C254D2B1701817F4C3EFE8CFC7A7A4A" authorityName="Portevin" authorityYear="1929" box="[870,986,300,324]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="tribe" tribe="Plegaderini">Plegaderini</taxonomicName>
is well justified on several characters (
<bibRefCitation id="EFB44B591701817F4A22FE8CFC357A6A" author="Zhou Y &amp; Ren D" pageId="9" pagination="521 - 39" refId="ref19290" refString="Zhou Y, Caterino Mº, Ren D et al. Phylogeny and evolution of Mesozoic and extant lineages of Histeridae (Coleoptera), with discovery of a new subfamily Antigracilinae from the Lower Cretaceous. Cladistics 2020; 36: 521 - 39. hups: // doi. org / 10.1111 / cla. 12418" type="journal article" year="2020">
<bibRefCitation id="EFB44B591701817F4A22FE8CFC357A6A" author="Zhou Y &amp; Ren D" pageId="9" pageNumber="10" pagination="521 - 39" refId="ref19290" refString="Zhou Y, Caterino Mº, Ren D et al. Phylogeny and evolution of Mesozoic and extant lineages of Histeridae (Coleoptera), with discovery of a new subfamily Antigracilinae from the Lower Cretaceous. Cladistics 2020; 36: 521 - 39. hups: // doi. org / 10.1111 / cla. 12418" type="journal article" year="2020">
Zhou
<emphasis id="B951EABA1701817F4C72FEECFCFB7A6A" box="[810,859,332,356]" italics="true" pageId="9">et al.</emphasis>
<emphasis id="B951EABA1701817F4C72FEECFCFB7A6A" box="[810,859,332,356]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
2020
</bibRefCitation>
), the alleged absence of postcoxal striae (on the lateral metaventrite and on abdominal ventrite 1) should be disregarded. Ŋeir absence is difficult to assess in the specimens of
<taxonomicName id="4C254D2B1701817F4AFAFE2BFC3C7ACC" authorityName="Poinar and Brown" authorityYear="2009" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="burmanicus">
<emphasis id="B951EABA1701817F4AFAFE2BFC3C7ACC" italics="true" pageId="9">P. burmanicus</emphasis>
), the alleged absence of postcoxal striae (on the lateral metaventrite and on abdominal ventrite 1) should be disregarded. Their absence is difficult to assess in the specimens of
<taxonomicName id="4C254D2B1701817F4AFAFE2BFC3C7ACC" authorityName="Poinar and Brown" authorityYear="2009" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="burmanicus">
<emphasis id="B951EABA1701817F4AFAFE2BFC3C7ACC" italics="true" pageId="9" pageNumber="10">P. burmanicus</emphasis>
</taxonomicName>
, and the postmesocoxal striae, at least, are visible in
<taxonomicName id="4C254D2B1701817F4C72FE6AFC2D7AEF" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[810,909,457,481]" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="hirsutus">
<emphasis id="B951EABA1701817F4C72FE6AFC2D7AEF" box="[810,909,457,481]" italics="true" pageId="9">P. hirsutus</emphasis>
<taxonomicName id="4C254D2B1701817F4C72FE6AFC2D7AEF" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[810,909,457,481]" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="hirsutus">
<emphasis id="B951EABA1701817F4C72FE6AFC2D7AEF" box="[810,909,457,481]" italics="true" pageId="9" pageNumber="10">P. hirsutus</emphasis>
</taxonomicName>
(see
<figureCitation id="131E2A2D1701817F4C9AFE69FBA87AEE" box="[962,1032,457,481]" captionStart="Figure 7" captionStartId="10.[129,194,1484,1508]" captionTargetBox="[129,1473,144,1456]" captionTargetId="figure-220@10.[129,1473,144,1456]" captionTargetPageId="10" captionText="Figure 7. Pantostictus hirsutus sp. nov.. A, dorsal view. B, ventral view.C, lateral view. D, elytral setation, E, dorsal view. F, metaventrite, abdomen.Abbreviations:ac, antennal club; as, antennal scape; ls, labral setae; pl, prosternal lobe; pmss, postmesocoxal stria." pageId="9">Fig. 7F</figureCitation>
<figureCitation id="131E2A2D1701817F4C9AFE69FBA87AEE" box="[962,1032,457,481]" captionStart="Figure 7" captionStartId="10.[129,194,1484,1508]" captionTargetBox="[129,1473,144,1456]" captionTargetId="figure-220@10.[129,1473,144,1456]" captionTargetPageId="10" captionText="Figure 7. Pantostictus hirsutus sp. nov.. A, dorsal view. B, ventral view.C, lateral view. D, elytral setation, E, dorsal view. F, metaventrite, abdomen.Abbreviations:ac, antennal club; as, antennal scape; ls, labral setae; pl, prosternal lobe; pmss, postmesocoxal stria." pageId="9" pageNumber="10">Fig. 7F</figureCitation>
).
</paragraph>
<paragraph id="8B9A36A81701817F4C72FDA2FCCF789C" blockId="9.[810,1463,514,914]" pageId="9">
<emphasis id="B951EABA1701817F4C72FDA2FBA07914" box="[810,1024,514,538]" italics="true" pageId="9">Differential diagnosis:</emphasis>
Ŋe species appears to be closely related to
<taxonomicName id="4C254D2B1701817F4C1EFD82FA967934" authority="Poinar and Brown, 2009" authorityName="Poinar and Brown" authorityYear="2009" box="[838,1334,546,570]" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="burmanicus">
<emphasis id="B951EABA1701817F4C1EFD82FB937934" box="[838,1075,546,570]" italics="true" pageId="9">Pantostictus burmanicus</emphasis>
<bibRefCitation id="EFB44B591701817F4B61FD82FA967934" author="Poinar G &amp; Brown AE" box="[1081,1334,546,570]" pageId="9" pagination="38 - 46" refId="ref18750" refString="Poinar G, Brown AE. Pantostictus burmanicus, a new genus and species of Cretaceous beetles (Coleoptera: Hydrophiloidea: Histeridae) in Burmese amber. Proceedings of the Entomological Society of Washington 2009; 111: 38 - 46. hups: // doi. org / 10.4289 / 0013 - 8797 - 111.1.38" type="journal article" year="2009">Poinar and Brown, 2009</bibRefCitation>
<paragraph id="8B9A36A81701817F4C72FDA2FCCF789C" blockId="9.[810,1463,514,914]" pageId="9" pageNumber="10">
<emphasis id="B951EABA1701817F4C72FDA2FBA07914" box="[810,1024,514,538]" italics="true" pageId="9" pageNumber="10">Differential diagnosis:</emphasis>
The species appears to be closely related to
<taxonomicName id="4C254D2B1701817F4C1EFD82FA967934" authority="Poinar and Brown, 2009" authorityName="Poinar and Brown" authorityYear="2009" box="[838,1334,546,570]" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="burmanicus">
<emphasis id="B951EABA1701817F4C1EFD82FB937934" box="[838,1075,546,570]" italics="true" pageId="9" pageNumber="10">Pantostictus burmanicus</emphasis>
<bibRefCitation id="EFB44B591701817F4B61FD82FA967934" author="Poinar G &amp; Brown AE" box="[1081,1334,546,570]" pageId="9" pageNumber="10" pagination="38 - 46" refId="ref18750" refString="Poinar G, Brown AE. Pantostictus burmanicus, a new genus and species of Cretaceous beetles (Coleoptera: Hydrophiloidea: Histeridae) in Burmese amber. Proceedings of the Entomological Society of Washington 2009; 111: 38 - 46. hups: // doi. org / 10.4289 / 0013 - 8797 - 111.1.38" type="journal article" year="2009">Poinar and Brown, 2009</bibRefCitation>
</taxonomicName>
. It is similar in overall appearance, but there are significant differential characters. Ŋe elytral punctuation is coarser in
<taxonomicName id="4C254D2B1701817F4BACFDC1FADE7976" authorityName="Poinar and Brown" authorityYear="2009" box="[1268,1406,608,632]" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="burmanicus">
<emphasis id="B951EABA1701817F4BACFDC1FADE7976" box="[1268,1406,608,632]" italics="true" pageId="9">P. burmanicus</emphasis>
. It is similar in overall appearance, but there are significant differential characters. The elytral punctuation is coarser in
<taxonomicName id="4C254D2B1701817F4BACFDC1FADE7976" authorityName="Poinar and Brown" authorityYear="2009" box="[1268,1406,608,632]" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="burmanicus">
<emphasis id="B951EABA1701817F4BACFDC1FADE7976" box="[1268,1406,608,632]" italics="true" pageId="9" pageNumber="10">P. burmanicus</emphasis>
</taxonomicName>
. Ŋe lateral pronotal margins are finely crenulate in
<taxonomicName id="4C254D2B1701817F4A4CFD20FADA7996" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[1300,1402,640,664]" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="hirsutus">
<emphasis id="B951EABA1701817F4A4CFD20FADA7996" box="[1300,1402,640,664]" italics="true" pageId="9">P. hirsutus</emphasis>
. The lateral pronotal margins are finely crenulate in
<taxonomicName id="4C254D2B1701817F4A4CFD20FADA7996" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[1300,1402,640,664]" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="hirsutus">
<emphasis id="B951EABA1701817F4A4CFD20FADA7996" box="[1300,1402,640,664]" italics="true" pageId="9" pageNumber="10">P. hirsutus</emphasis>
</taxonomicName>
(
<figureCitation id="131E2A2D1701817F4AD5FD20FCE679B8" captionStart="Figure 7" captionStartId="10.[129,194,1484,1508]" captionTargetBox="[129,1473,144,1456]" captionTargetId="figure-220@10.[129,1473,144,1456]" captionTargetPageId="10" captionText="Figure 7. Pantostictus hirsutus sp. nov.. A, dorsal view. B, ventral view.C, lateral view. D, elytral setation, E, dorsal view. F, metaventrite, abdomen.Abbreviations:ac, antennal club; as, antennal scape; ls, labral setae; pl, prosternal lobe; pmss, postmesocoxal stria." pageId="9">Fig. 7E</figureCitation>
<figureCitation id="131E2A2D1701817F4AD5FD20FCE679B8" captionStart="Figure 7" captionStartId="10.[129,194,1484,1508]" captionTargetBox="[129,1473,144,1456]" captionTargetId="figure-220@10.[129,1473,144,1456]" captionTargetPageId="10" captionText="Figure 7. Pantostictus hirsutus sp. nov.. A, dorsal view. B, ventral view.C, lateral view. D, elytral setation, E, dorsal view. F, metaventrite, abdomen.Abbreviations:ac, antennal club; as, antennal scape; ls, labral setae; pl, prosternal lobe; pmss, postmesocoxal stria." pageId="9" pageNumber="10">Fig. 7E</figureCitation>
), whereas in
<taxonomicName id="4C254D2B1701817F4C95FD00FBF579B9" authorityName="Poinar and Brown" authorityYear="2009" box="[973,1109,671,695]" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="burmanicus">
<emphasis id="B951EABA1701817F4C95FD00FBF579B9" box="[973,1109,671,695]" italics="true" pageId="9">P. burmanicus</emphasis>
<taxonomicName id="4C254D2B1701817F4C95FD00FBF579B9" authorityName="Poinar and Brown" authorityYear="2009" box="[973,1109,671,695]" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="burmanicus">
<emphasis id="B951EABA1701817F4C95FD00FBF579B9" box="[973,1109,671,695]" italics="true" pageId="9" pageNumber="10">P. burmanicus</emphasis>
</taxonomicName>
they are smooth. In
<taxonomicName id="4C254D2B1701817F4A7CFD00FA2779B9" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[1316,1415,671,695]" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="hirsutus">
<emphasis id="B951EABA1701817F4A7CFD00FA2779B9" box="[1316,1415,671,695]" italics="true" pageId="9">P. hirsutus</emphasis>
<taxonomicName id="4C254D2B1701817F4A7CFD00FA2779B9" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[1316,1415,671,695]" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="hirsutus">
<emphasis id="B951EABA1701817F4A7CFD00FA2779B9" box="[1316,1415,671,695]" italics="true" pageId="9" pageNumber="10">P. hirsutus</emphasis>
</taxonomicName>
, the pronotum is wider basally and lateral margins are simply rounded and parallel-sided in the posterior half (
<figureCitation id="131E2A2D1701817F4BEEFD7DFB5E79F8" box="[1206,1278,733,758]" captionStart="Figure 7" captionStartId="10.[129,194,1484,1508]" captionTargetBox="[129,1473,144,1456]" captionTargetId="figure-220@10.[129,1473,144,1456]" captionTargetPageId="10" captionText="Figure 7. Pantostictus hirsutus sp. nov.. A, dorsal view. B, ventral view.C, lateral view. D, elytral setation, E, dorsal view. F, metaventrite, abdomen.Abbreviations:ac, antennal club; as, antennal scape; ls, labral setae; pl, prosternal lobe; pmss, postmesocoxal stria." pageId="9">Fig. 7A</figureCitation>
<figureCitation id="131E2A2D1701817F4BEEFD7DFB5E79F8" box="[1206,1278,733,758]" captionStart="Figure 7" captionStartId="10.[129,194,1484,1508]" captionTargetBox="[129,1473,144,1456]" captionTargetId="figure-220@10.[129,1473,144,1456]" captionTargetPageId="10" captionText="Figure 7. Pantostictus hirsutus sp. nov.. A, dorsal view. B, ventral view.C, lateral view. D, elytral setation, E, dorsal view. F, metaventrite, abdomen.Abbreviations:ac, antennal club; as, antennal scape; ls, labral setae; pl, prosternal lobe; pmss, postmesocoxal stria." pageId="9" pageNumber="10">Fig. 7A</figureCitation>
), while the lateral pronotal margins of
<taxonomicName id="4C254D2B1701817F4B59FD5EFB2C781B" authorityName="Poinar and Brown" authorityYear="2009" box="[1025,1164,765,789]" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="burmanicus">
<emphasis id="B951EABA1701817F4B59FD5EFB2C781B" box="[1025,1164,765,789]" italics="true" pageId="9">P. burmanicus</emphasis>
<taxonomicName id="4C254D2B1701817F4B59FD5EFB2C781B" authorityName="Poinar and Brown" authorityYear="2009" box="[1025,1164,765,789]" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="burmanicus">
<emphasis id="B951EABA1701817F4B59FD5EFB2C781B" box="[1025,1164,765,789]" italics="true" pageId="9" pageNumber="10">P. burmanicus</emphasis>
</taxonomicName>
are sinuate. Furthermore,
<taxonomicName id="4C254D2B1701817F4AFAFD5EFCD7783A" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="hirsutus">
<emphasis id="B951EABA1701817F4AFAFD5EFCD7783A" italics="true" pageId="9">P. hirsutus</emphasis>
<taxonomicName id="4C254D2B1701817F4AFAFD5EFCD7783A" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="hirsutus">
<emphasis id="B951EABA1701817F4AFAFD5EFCD7783A" italics="true" pageId="9" pageNumber="10">P. hirsutus</emphasis>
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displays a setose dorsal cuticle (
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). ºetation presence in
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<emphasis id="B951EABA1701817F4C26FC9CFBA9785D" box="[894,1033,827,851]" italics="true" pageId="9">P. burmanicus</emphasis>
<figureCitation id="131E2A2D1701817F4B9BFCBCFAAE783A" box="[1219,1294,796,820]" captionStart="Figure 7" captionStartId="10.[129,194,1484,1508]" captionTargetBox="[129,1473,144,1456]" captionTargetId="figure-220@10.[129,1473,144,1456]" captionTargetPageId="10" captionText="Figure 7. Pantostictus hirsutus sp. nov.. A, dorsal view. B, ventral view.C, lateral view. D, elytral setation, E, dorsal view. F, metaventrite, abdomen.Abbreviations:ac, antennal club; as, antennal scape; ls, labral setae; pl, prosternal lobe; pmss, postmesocoxal stria." pageId="9" pageNumber="10">Fig. 7D</figureCitation>
). Setation presence in
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<emphasis id="B951EABA1701817F4C26FC9CFBA9785D" box="[894,1033,827,851]" italics="true" pageId="9" pageNumber="10">P. burmanicus</emphasis>
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cannot be determined based on the original description and published photographs (
<bibRefCitation id="EFB44B591701817F4BA4FCFBFCFE789C" author="Poinar G &amp; Brown AE" pageId="9" pagination="38 - 46" refId="ref18750" refString="Poinar G, Brown AE. Pantostictus burmanicus, a new genus and species of Cretaceous beetles (Coleoptera: Hydrophiloidea: Histeridae) in Burmese amber. Proceedings of the Entomological Society of Washington 2009; 111: 38 - 46. hups: // doi. org / 10.4289 / 0013 - 8797 - 111.1.38" type="journal article" year="2009">Poinar and Brown 2009</bibRefCitation>
<bibRefCitation id="EFB44B591701817F4BA4FCFBFCFE789C" author="Poinar G &amp; Brown AE" pageId="9" pageNumber="10" pagination="38 - 46" refId="ref18750" refString="Poinar G, Brown AE. Pantostictus burmanicus, a new genus and species of Cretaceous beetles (Coleoptera: Hydrophiloidea: Histeridae) in Burmese amber. Proceedings of the Entomological Society of Washington 2009; 111: 38 - 46. hups: // doi. org / 10.4289 / 0013 - 8797 - 111.1.38" type="journal article" year="2009">Poinar and Brown 2009</bibRefCitation>
).
</paragraph>
<paragraph id="8B9A36A81701817F4C72FC13FC6078E5" blockId="9.[810,1459,947,1003]" pageId="9">
<emphasis id="B951EABA1701817F4C72FC13FC3978C5" box="[810,921,947,971]" italics="true" pageId="9">Etymology:</emphasis>
Ŋe name of this species was chosen to point out its hirsute cuticle.
<paragraph id="8B9A36A81701817F4C72FC13FC6078E5" blockId="9.[810,1459,947,1003]" pageId="9" pageNumber="10">
<emphasis id="B951EABA1701817F4C72FC13FC3978C5" box="[810,921,947,971]" italics="true" pageId="9" pageNumber="10">Etymology:</emphasis>
The name of this species was chosen to point out its hirsute cuticle.
</paragraph>
</subSubSection>
</treatment>

291
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<subSubSection id="C33F65231704817A4C8AFDC6FA8A798F" box="[978,1322,614,641]" pageId="12" type="nomenclature">
<paragraph id="8B9A36A81704817A4C8AFDC6FA8A798F" blockId="12.[978,1322,614,641]" box="[978,1322,614,641]" pageId="12">
<heading id="D0D281C41704817A4C8AFDC6FA8A798F" bold="true" box="[978,1322,614,641]" centered="true" fontSize="11" level="2" pageId="12" reason="2">
<emphasis id="B951EABA1704817A4C8AFDC6FA8A798F" bold="true" box="[978,1322,614,641]" pageId="12">
<taxonomicName id="4C254D2B1704817A4C8AFDC6FB6D798E" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[978,1229,614,641]" class="Insecta" family="Histeridae" genus="Pseudacritus" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="species" species="extinctus" status="sp. nov.">
<emphasis id="B951EABA1704817A4C8AFDC6FB6D798E" bold="true" box="[978,1229,614,641]" italics="true" pageId="12">Pseudacritus extinctus</emphasis>
<treatment id="038C87BE170481794C8AFDC6FD5B7BA9" LSID="urn:lsid:plazi:treatment:038C87BE170481794C8AFDC6FD5B7BA9" httpUri="http://treatment.plazi.org/id/038C87BE170481794C8AFDC6FD5B7BA9" lastPageId="15" lastPageNumber="16" pageId="12" pageNumber="13">
<subSubSection id="C33F65231704817A4C8AFDC6FA8A798F" box="[978,1322,614,641]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="8B9A36A81704817A4C8AFDC6FA8A798F" blockId="12.[978,1322,614,641]" box="[978,1322,614,641]" pageId="12" pageNumber="13">
<heading id="D0D281C41704817A4C8AFDC6FA8A798F" bold="true" box="[978,1322,614,641]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="13" reason="2">
<emphasis id="B951EABA1704817A4C8AFDC6FA8A798F" bold="true" box="[978,1322,614,641]" pageId="12" pageNumber="13">
<taxonomicName id="4C254D2B1704817A4C8AFDC6FB6D798E" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[978,1229,614,641]" class="Insecta" family="Histeridae" genus="Pseudacritus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="extinctus" status="sp. nov.">
<emphasis id="B951EABA1704817A4C8AFDC6FB6D798E" bold="true" box="[978,1229,614,641]" italics="true" pageId="12" pageNumber="13">Pseudacritus extinctus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C11704817A4B8AFDC7FA8A798F" box="[1234,1322,615,641]" pageId="12" rank="species">sp. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="A26257C11704817A4B8AFDC7FA8A798F" box="[1234,1322,615,641]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33F6523170481794B0FFD35FD5B7BA9" lastPageId="15" pageId="12" type="description">
<paragraph id="8B9A36A81704817A4B0FFD35FB0579A1" blockId="12.[1111,1189,661,688]" box="[1111,1189,661,688]" pageId="12">
<emphasis id="B951EABA1704817A4B0FFD35FB0579A1" bold="true" box="[1111,1189,661,688]" pageId="12">
<subSubSection id="C33F65231704817A4B0FFD35FB0579A1" box="[1111,1189,661,688]" pageId="12" pageNumber="13" type="description">
<paragraph id="8B9A36A81704817A4B0FFD35FB0579A1" blockId="12.[1111,1189,661,688]" box="[1111,1189,661,688]" pageId="12" pageNumber="13">
<emphasis id="B951EABA1704817A4B0FFD35FB0579A1" bold="true" box="[1111,1189,661,688]" pageId="12" pageNumber="13">
(
<figureCitation id="131E2A2D1704817A4B39FD36FB3A79BE" box="[1121,1178,662,688]" captionStart="Figure 9" captionStartId="13.[112,177,1501,1525]" captionTargetBox="[114,1458,144,1473]" captionTargetId="figure-201@13.[114,1458,144,1473]" captionTargetPageId="13" captionText="Figure 9. Pseudacritus extinctus gen.et sp. nov.. A, dorsal view.B, ventral view. C, lateroventral view.D, antenna. E, frontal view. F, pronotal base. Abbreviations:aca, antennal cavity; acl, antennal club; as, antennal scape; ass, antescutellar stria; lab, labrum; mp, maxillary palpus; sd, scutellar disc." pageId="12">Fig. 9</figureCitation>
<figureCitation id="131E2A2D1704817A4B39FD36FB3A79BE" box="[1121,1178,662,688]" captionStart="Figure 9" captionStartId="13.[112,177,1501,1525]" captionTargetBox="[114,1458,144,1473]" captionTargetId="figure-201@13.[114,1458,144,1473]" captionTargetPageId="13" captionText="Figure 9. Pseudacritus extinctus gen.et sp. nov.. A, dorsal view.B, ventral view. C, lateroventral view.D, antenna. E, frontal view. F, pronotal base. Abbreviations:aca, antennal cavity; acl, antennal club; as, antennal scape; ass, antescutellar stria; lab, labrum; mp, maxillary palpus; sd, scutellar disc." pageId="12" pageNumber="13">Fig. 9</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8B9A36A81704817A4C61FD76FB467842" blockId="12.[825,1475,726,844]" pageId="12">
<emphasis id="B951EABA1704817A4C61FD76FC3579E0" box="[825,917,726,750]" italics="true" pageId="12">Material:</emphasis>
<collectingRegion id="49E1F84A1704817A4CC4FD76FC4679E0" box="[924,998,726,750]" country="Myanmar" name="Kachin" pageId="12">Kachin</collectingRegion>
</subSubSection>
<subSubSection id="C33F65231704817A4C61FD76FB467842" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="8B9A36A81704817A4C61FD76FB467842" blockId="12.[825,1475,726,844]" pageId="12" pageNumber="13">
<emphasis id="B951EABA1704817A4C61FD76FC3579E0" box="[825,917,726,750]" italics="true" pageId="12" pageNumber="13">Material:</emphasis>
<materialsCitation id="3B4D3CF51704817A4CC4FD76FB417842" collectionCode="MEBHK" location="Kachin" pageId="12" pageNumber="13" specimenCode="P1423/E" specimenCount="1" stateProvince="Kachin">
<collectingRegion id="49E1F84A1704817A4CC4FD76FC4679E0" box="[924,998,726,750]" country="Myanmar" name="Kachin" pageId="12" pageNumber="13">Kachin</collectingRegion>
amber (
<emphasis id="B951EABA1704817A4B18FD77FBE979E0" box="[1088,1097,727,750]" italics="true" pageId="12">c</emphasis>
<emphasis id="B951EABA1704817A4B18FD77FBE979E0" box="[1088,1097,727,750]" italics="true" pageId="12" pageNumber="13">c</emphasis>
. 99 Mya). Amber piece dimensions: 16 × 5 ×
<quantity id="4CDD9B4D1704817A4CC2FD56FC757803" box="[922,981,758,782]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="12" unit="mm" value="2.0">2 mm</quantity>
<quantity id="4CDD9B4D1704817A4CC2FD56FC757803" box="[922,981,758,782]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="13" unit="mm" value="2.0">2 mm</quantity>
. Amber clear with a large
<taxonomicName id="4C254D2B1704817A4BA2FD56FA297800" box="[1274,1417,758,782]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Hymenoptera" pageId="12" phylum="Tracheophyta" rank="order">Hymenoptera</taxonomicName>
<taxonomicName id="4C254D2B1704817A4BA2FD56FA297800" box="[1274,1417,758,782]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="order">Hymenoptera</taxonomicName>
fragment, a single specimen of
<taxonomicName id="4C254D2B1704817A4B3CFCB5FB4E7823" authorityName="Latreille" authorityYear="1802" box="[1124,1262,789,813]" class="Insecta" family="Trogossitidae" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="family">Trogossitidae</taxonomicName>
and large pieces of debris. Deposited at MEBHK (P1423/E).
<taxonomicName id="4C254D2B1704817A4B3CFCB5FB4E7823" authorityName="Latreille" authorityYear="1802" box="[1124,1262,789,813]" class="Insecta" family="Trogossitidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="family">Trogossitidae</taxonomicName>
and large pieces of debris. Deposited at
<collectionCode id="ED34AE6D1704817A4B52FC95FBC67842" box="[1034,1126,821,844]" pageId="12" pageNumber="13">MEBHK</collectionCode>
(
<specimenCode id="DB839ED31704817A4B2CFC94FB797842" box="[1140,1241,820,844]" pageId="12" pageNumber="13">P1423/E</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B9A36A81704817B4C61FCCEFE667DDD" blockId="12.[825,1475,878,1967]" lastBlockId="13.[113,762,1629,1747]" lastPageId="13" pageId="12">
<emphasis id="B951EABA1704817A4C61FCCEFC6C788B" box="[825,972,878,901]" italics="true" pageId="12">Measurements:</emphasis>
</subSubSection>
<subSubSection id="C33F65231704817B4C61FCCEFE667DDD" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="description">
<paragraph id="8B9A36A81704817B4C61FCCEFE667DDD" blockId="12.[825,1475,878,1967]" lastBlockId="13.[113,762,1629,1747]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<emphasis id="B951EABA1704817A4C61FCCEFC6C788B" box="[825,972,878,901]" italics="true" pageId="12" pageNumber="13">Measurements:</emphasis>
Head width:
<quantity id="4CDD9B4D1704817A4B30FCCEFB15788B" box="[1128,1205,878,902]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="12" unit="mm" value="0.3">0.3 mm</quantity>
<quantity id="4CDD9B4D1704817A4B30FCCEFB15788B" box="[1128,1205,878,902]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="13" unit="mm" value="0.3">0.3 mm</quantity>
; width between anterior pronotal angles:
<quantity id="4CDD9B4D1704817A4CB5FC2DFBE878AA" box="[1005,1096,909,933]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="12" unit="mm" value="0.34">0.34 mm</quantity>
<quantity id="4CDD9B4D1704817A4CB5FC2DFBE878AA" box="[1005,1096,909,933]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="12" pageNumber="13" unit="mm" value="0.34">0.34 mm</quantity>
; width between posterior pronotal angles:
<quantity id="4CDD9B4D1704817A4CC9FC0CFC4C78CA" box="[913,1004,940,964]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="12" unit="mm" value="0.68">0.68 mm</quantity>
<quantity id="4CDD9B4D1704817A4CC9FC0CFC4C78CA" box="[913,1004,940,964]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="12" pageNumber="13" unit="mm" value="0.68">0.68 mm</quantity>
; pronotal length:
<quantity id="4CDD9B4D1704817A4B9DFC0CFAB378CA" box="[1221,1299,940,964]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="12" unit="mm" value="0.4">0.4 mm</quantity>
<quantity id="4CDD9B4D1704817A4B9DFC0CFAB378CA" box="[1221,1299,940,964]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="12" pageNumber="13" unit="mm" value="0.4">0.4 mm</quantity>
; sutural elytral length:
<quantity id="4CDD9B4D1704817A4CD3FC6CFC4678ED" box="[907,998,972,996]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="12" unit="mm" value="0.48">0.48 mm</quantity>
<quantity id="4CDD9B4D1704817A4CD3FC6CFC4678ED" box="[907,998,972,996]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="12" pageNumber="13" unit="mm" value="0.48">0.48 mm</quantity>
; elytral width (in pair, across widest point):
<quantity id="4CDD9B4D1704817A4C61FC4BFC297F0C" box="[825,905,1003,1027]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="12" unit="mm" value="0.7">0.7 mm</quantity>
<quantity id="4CDD9B4D1704817A4C61FC4BFC297F0C" box="[825,905,1003,1027]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="12" pageNumber="13" unit="mm" value="0.7">0.7 mm</quantity>
.—
<emphasis id="B951EABA1704817A4CF0FC4BFC487F0D" bold="true" box="[936,1000,1003,1027]" italics="true" pageId="12">Body:</emphasis>
<emphasis id="B951EABA1704817A4CF0FC4BFC487F0D" bold="true" box="[936,1000,1003,1027]" italics="true" pageId="12" pageNumber="13">Body:</emphasis>
Round and dorsoventrally convex, brown in colour.—
<emphasis id="B951EABA1704817A4CC3FBAAFC7C7F2C" bold="true" box="[923,988,1034,1058]" italics="true" pageId="12">Head:</emphasis>
Without frontal stria. Frons smooth, slightly convex. Clypeus basally constricted, widening anteriorly. Labial as well as maxillary palpi exposed, the laưer rather short, wide. Labrum with a transverse carinate median ridge, with a pair of prominent setae and several shorter setae. Terminal maxillary palpomere oval and pointed. Terminal labial palpomere short, pointed. Eyes large, convex. Antennal scape thickened, as long as antennomeres 2–7 combined. Pedicel approximately third scape’s length, wider basally. Club large, oval, flaưened, with two distinct annuli.—
<emphasis id="B951EABA1704817A4B7EFA84FBDA7E32" bold="true" box="[1062,1146,1316,1340]" italics="true" pageId="12">Ŋorax:</emphasis>
Pronotal disc almost smooth, with widely isolated small punctures and with a complete, punctuate antescutellar stria. Anterior corners acute. Marginal stria present. Lateral pronotal stria carinate, slightly expanding over pronotal margin medially and gradually distanced from pronotal margin basally. ºcutellar shield small, triangular. Elytra dorsally with four weak costae. Elytral epipleuron concave, epipleural stria obscured. Elytral humeri with four short distinct costate striae. Prosternal keel short and wide, largely unobservable, laterally incised for the antennal passage. Round and relatively well-developed antennal cavities located at the anteriormost part of the prosternal hypomeron. Hypomeron excavate, creating a depression shaped to receive the profemur and protibia. When retracted the protibial would probably conceal the antennal club. Mesoventrite obscured by legs and fracturing. Metaventrite very finely punctuated, with a median depression, which may be an artefact. Postmesocoxal stria complete, recurved along mesocoxa. Lateral metaventral stria complete, continuous on to metepisternum. Metepisternum exposed.—
<emphasis id="B951EABA1704817A4A4FF8F8FAEE7C61" bold="true" box="[1303,1358,1880,1903]" italics="true" pageId="12">Legs:</emphasis>
All femora and tibiae thin, laterally flaưened. Tarsal formula 5-5-5, ultimate tarsomeres as long as the preceding two.—
<emphasis id="B951EABA1704817A4A41F836FA267CA0" bold="true" box="[1305,1414,1942,1966]" italics="true" pageId="12">Abdomen:</emphasis>
<emphasis id="B951EABA1704817A4CC3FBAAFC7C7F2C" bold="true" box="[923,988,1034,1058]" italics="true" pageId="12" pageNumber="13">Head:</emphasis>
Without frontal stria. Frons smooth, slightly convex. Clypeus basally constricted, widening anteriorly. Labial as well as maxillary palpi exposed, the latter rather short, wide. Labrum with a transverse carinate median ridge, with a pair of prominent setae and several shorter setae. Terminal maxillary palpomere oval and pointed. Terminal labial palpomere short, pointed. Eyes large, convex. Antennal scape thickened, as long as antennomeres 2–7 combined. Pedicel approximately third scape’s length, wider basally. Club large, oval, flattened, with two distinct annuli.—
<emphasis id="B951EABA1704817A4B7EFA84FBDA7E32" bold="true" box="[1062,1146,1316,1340]" italics="true" pageId="12" pageNumber="13">Thorax:</emphasis>
Pronotal disc almost smooth, with widely isolated small punctures and with a complete, punctuate antescutellar stria. Anterior corners acute. Marginal stria present. Lateral pronotal stria carinate, slightly expanding over pronotal margin medially and gradually distanced from pronotal margin basally. Scutellar shield small, triangular. Elytra dorsally with four weak costae. Elytral epipleuron concave, epipleural stria obscured. Elytral humeri with four short distinct costate striae. Prosternal keel short and wide, largely unobservable, laterally incised for the antennal passage. Round and relatively well-developed antennal cavities located at the anteriormost part of the prosternal hypomeron. Hypomeron excavate, creating a depression shaped to receive the profemur and protibia. When retracted the protibial would probably conceal the antennal club. Mesoventrite obscured by legs and fracturing. Metaventrite very finely punctuated, with a median depression, which may be an artefact. Postmesocoxal stria complete, recurved along mesocoxa. Lateral metaventral stria complete, continuous on to metepisternum. Metepisternum exposed.—
<emphasis id="B951EABA1704817A4A4FF8F8FAEE7C61" bold="true" box="[1303,1358,1880,1903]" italics="true" pageId="12" pageNumber="13">Legs:</emphasis>
All femora and tibiae thin, laterally flattened. Tarsal formula 5-5-5, ultimate tarsomeres as long as the preceding two.—
<emphasis id="B951EABA1704817A4A41F836FA267CA0" bold="true" box="[1305,1414,1942,1966]" italics="true" pageId="12" pageNumber="13">Abdomen:</emphasis>
First abdominal ventrite wide, with thin and complete postmetacoxal striae. Pygidium triangular, laterobasally with depressions, apex with a single short groove medially. Propygidium short and broad, with irregular depressions.
</paragraph>
<caption id="DF5A66201705817B4F28FA7DFF3D7D23" pageId="13" startId="13.[112,177,1501,1525]" targetBox="[114,1458,144,1473]" targetPageId="13" targetType="figure">
<paragraph id="8B9A36A81705817B4F28FA7DFF3D7D23" blockId="13.[112,1457,1501,1581]" pageId="13">
<emphasis id="B951EABA1705817B4F28FA7DFF687EFB" bold="true" box="[112,200,1501,1525]" pageId="13">Figure 9.</emphasis>
<taxonomicName id="4C254D2B1705817B4F95FA7DFE337EFB" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[205,403,1501,1525]" class="Insecta" family="Histeridae" genus="Pseudacritus" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="species" species="extinctus" status="gen. et sp. nov">
<emphasis id="B951EABA1705817B4F95FA7DFE337EFB" box="[205,403,1501,1525]" italics="true" pageId="13">Pseudacritus extinctus</emphasis>
</subSubSection>
<caption id="DF5A66201705817B4F28FA7DFF3D7D23" pageId="13" pageNumber="14" startId="13.[112,177,1501,1525]" targetBox="[114,1458,144,1473]" targetPageId="13" targetType="figure">
<paragraph id="8B9A36A81705817B4F28FA7DFF3D7D23" blockId="13.[112,1457,1501,1581]" pageId="13" pageNumber="14">
<emphasis id="B951EABA1705817B4F28FA7DFF687EFB" bold="true" box="[112,200,1501,1525]" pageId="13" pageNumber="14">Figure 9.</emphasis>
<taxonomicName id="4C254D2B1705817B4F95FA7DFE337EFB" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[205,403,1501,1525]" class="Insecta" family="Histeridae" genus="Pseudacritus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="extinctus" status="gen. et sp. nov">
<emphasis id="B951EABA1705817B4F95FA7DFE337EFB" box="[205,403,1501,1525]" italics="true" pageId="13" pageNumber="14">Pseudacritus extinctus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C11705817B4EC0FA7EFDBB7EFB" box="[408,539,1502,1526]" pageId="13" rank="species">gen. et sp. nov</taxonomicNameLabel>
<taxonomicNameLabel id="A26257C11705817B4EC0FA7EFDBB7EFB" box="[408,539,1502,1526]" pageId="13" pageNumber="14" rank="species">gen. et sp. nov</taxonomicNameLabel>
.. A, dorsal view. B, ventral view. C, lateroventral view. D, antenna. E, frontal view. F, pronotal base. Abbreviations: aca, antennal cavity; acl, antennal club; as, antennal scape; ass, antescutellar stria; lab, labrum; mp, maxillary palpus; sd, scutellar disc.
</paragraph>
</caption>
<paragraph id="8B9A36A81705817B4F29F955FAB27C5E" blockId="13.[113,763,1781,1961]" lastBlockId="13.[810,1460,1629,1872]" pageId="13">
<emphasis id="B951EABA1705817B4F29F955FEF17C02" box="[113,337,1781,1804]" italics="true" pageId="13">Taxonomic assignment:</emphasis>
Ŋis species can be tentatively placed in the
<taxonomicName id="4C254D2B1705817B4F29F8B4FF777C22" authorityName="W. S. MacLeay" authorityYear="1819" box="[113,215,1812,1836]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="subFamily" subFamily="Abraeinae">Abraeinae</taxonomicName>
<subSubSection id="C33F65231705817B4F29F955FAB27C5E" pageId="13" pageNumber="14" type="discussion">
<paragraph id="8B9A36A81705817B4F29F955FAB27C5E" blockId="13.[113,763,1781,1961]" lastBlockId="13.[810,1460,1629,1872]" pageId="13" pageNumber="14">
<emphasis id="B951EABA1705817B4F29F955FEF17C02" box="[113,337,1781,1804]" italics="true" pageId="13" pageNumber="14">Taxonomic assignment:</emphasis>
This species can be tentatively placed in the
<taxonomicName id="4C254D2B1705817B4F29F8B4FF777C22" authorityName="W. S. MacLeay" authorityYear="1819" box="[113,215,1812,1836]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subFamily" subFamily="Abraeinae">Abraeinae</taxonomicName>
subfamily, based on the absence of a distinct pronotal lobe, small body size of round shape, and strongly convex eyes (
<figureCitation id="131E2A2D1705817B4F24F8F3FF667C65" box="[124,198,1875,1899]" captionStart="Figure 9" captionStartId="13.[112,177,1501,1525]" captionTargetBox="[114,1458,144,1473]" captionTargetId="figure-201@13.[114,1458,144,1473]" captionTargetPageId="13" captionText="Figure 9. Pseudacritus extinctus gen.et sp. nov.. A, dorsal view.B, ventral view. C, lateroventral view.D, antenna. E, frontal view. F, pronotal base. Abbreviations:aca, antennal cavity; acl, antennal club; as, antennal scape; ass, antescutellar stria; lab, labrum; mp, maxillary palpus; sd, scutellar disc." pageId="13">Fig. 9A</figureCitation>
). Ŋis position was also supported by all phylogenetic analyses. Ŋe species bears characters strikingly similar to the extant
<taxonomicName id="4C254D2B1705817B4FEFF831FEA97CA7" authorityName="Wenzel" authorityYear="1944" box="[183,265,1937,1961]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="tribe" tribe="Acritini">Acritini</taxonomicName>
<figureCitation id="131E2A2D1705817B4F24F8F3FF667C65" box="[124,198,1875,1899]" captionStart="Figure 9" captionStartId="13.[112,177,1501,1525]" captionTargetBox="[114,1458,144,1473]" captionTargetId="figure-201@13.[114,1458,144,1473]" captionTargetPageId="13" captionText="Figure 9. Pseudacritus extinctus gen.et sp. nov.. A, dorsal view.B, ventral view. C, lateroventral view.D, antenna. E, frontal view. F, pronotal base. Abbreviations:aca, antennal cavity; acl, antennal club; as, antennal scape; ass, antescutellar stria; lab, labrum; mp, maxillary palpus; sd, scutellar disc." pageId="13" pageNumber="14">Fig. 9A</figureCitation>
). This position was also supported by all phylogenetic analyses. The species bears characters strikingly similar to the extant
<taxonomicName id="4C254D2B1705817B4FEFF831FEA97CA7" authorityName="Wenzel" authorityYear="1944" box="[183,265,1937,1961]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="tribe" tribe="Acritini">Acritini</taxonomicName>
: position of the antennal cavity at the frontmost part of prosternal hypomeron (
<figureCitation id="131E2A2D1705817B4B34F9FDFB177D7B" box="[1132,1207,1629,1653]" captionStart="Figure 9" captionStartId="13.[112,177,1501,1525]" captionTargetBox="[114,1458,144,1473]" captionTargetId="figure-201@13.[114,1458,144,1473]" captionTargetPageId="13" captionText="Figure 9. Pseudacritus extinctus gen.et sp. nov.. A, dorsal view.B, ventral view. C, lateroventral view.D, antenna. E, frontal view. F, pronotal base. Abbreviations:aca, antennal cavity; acl, antennal club; as, antennal scape; ass, antescutellar stria; lab, labrum; mp, maxillary palpus; sd, scutellar disc." pageId="13">Fig. 9C</figureCitation>
<figureCitation id="131E2A2D1705817B4B34F9FDFB177D7B" box="[1132,1207,1629,1653]" captionStart="Figure 9" captionStartId="13.[112,177,1501,1525]" captionTargetBox="[114,1458,144,1473]" captionTargetId="figure-201@13.[114,1458,144,1473]" captionTargetPageId="13" captionText="Figure 9. Pseudacritus extinctus gen.et sp. nov.. A, dorsal view.B, ventral view. C, lateroventral view.D, antenna. E, frontal view. F, pronotal base. Abbreviations:aca, antennal cavity; acl, antennal club; as, antennal scape; ass, antescutellar stria; lab, labrum; mp, maxillary palpus; sd, scutellar disc." pageId="13" pageNumber="14">Fig. 9C</figureCitation>
) and the presence of an antescutellar stria on the pronotal disc (
<figureCitation id="131E2A2D1705817B4B95F9DDFA9A7D9A" box="[1229,1338,1661,1685]" captionStart="Figure 9" captionStartId="13.[112,177,1501,1525]" captionTargetBox="[114,1458,144,1473]" captionTargetId="figure-201@13.[114,1458,144,1473]" captionTargetPageId="13" captionText="Figure 9. Pseudacritus extinctus gen.et sp. nov.. A, dorsal view.B, ventral view. C, lateroventral view.D, antenna. E, frontal view. F, pronotal base. Abbreviations:aca, antennal cavity; acl, antennal club; as, antennal scape; ass, antescutellar stria; lab, labrum; mp, maxillary palpus; sd, scutellar disc." pageId="13">Fig. 9A, D</figureCitation>
<figureCitation id="131E2A2D1705817B4B95F9DDFA9A7D9A" box="[1229,1338,1661,1685]" captionStart="Figure 9" captionStartId="13.[112,177,1501,1525]" captionTargetBox="[114,1458,144,1473]" captionTargetId="figure-201@13.[114,1458,144,1473]" captionTargetPageId="13" captionText="Figure 9. Pseudacritus extinctus gen.et sp. nov.. A, dorsal view.B, ventral view. C, lateroventral view.D, antenna. E, frontal view. F, pronotal base. Abbreviations:aca, antennal cavity; acl, antennal club; as, antennal scape; ass, antescutellar stria; lab, labrum; mp, maxillary palpus; sd, scutellar disc." pageId="13" pageNumber="14">Fig. 9A, D</figureCitation>
). However, it lacks the key defining feature of
<taxonomicName id="4C254D2B1705817B4BC9F93CFB407DBA" authorityName="Wenzel" authorityYear="1944" box="[1169,1248,1692,1716]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="tribe" tribe="Acritini">Acritini</taxonomicName>
<taxonomicName id="4C254D2B1705817B4BC9F93CFB407DBA" authorityName="Wenzel" authorityYear="1944" box="[1169,1248,1692,1716]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="tribe" tribe="Acritini">Acritini</taxonomicName>
, the four-segmented metatarsi. Moreover, the species exhibits a plesiomorphic annulate antennal club (
<figureCitation id="131E2A2D1705817B4CBEF97BFB8E7DFD" box="[998,1070,1755,1779]" captionStart="Figure 9" captionStartId="13.[112,177,1501,1525]" captionTargetBox="[114,1458,144,1473]" captionTargetId="figure-201@13.[114,1458,144,1473]" captionTargetPageId="13" captionText="Figure 9. Pseudacritus extinctus gen.et sp. nov.. A, dorsal view.B, ventral view. C, lateroventral view.D, antenna. E, frontal view. F, pronotal base. Abbreviations:aca, antennal cavity; acl, antennal club; as, antennal scape; ass, antescutellar stria; lab, labrum; mp, maxillary palpus; sd, scutellar disc." pageId="13">Fig. 9C</figureCitation>
<figureCitation id="131E2A2D1705817B4CBEF97BFB8E7DFD" box="[998,1070,1755,1779]" captionStart="Figure 9" captionStartId="13.[112,177,1501,1525]" captionTargetBox="[114,1458,144,1473]" captionTargetId="figure-201@13.[114,1458,144,1473]" captionTargetPageId="13" captionText="Figure 9. Pseudacritus extinctus gen.et sp. nov.. A, dorsal view.B, ventral view. C, lateroventral view.D, antenna. E, frontal view. F, pronotal base. Abbreviations:aca, antennal cavity; acl, antennal club; as, antennal scape; ass, antescutellar stria; lab, labrum; mp, maxillary palpus; sd, scutellar disc." pageId="13" pageNumber="14">Fig. 9C</figureCitation>
), as opposed to most recent
<taxonomicName id="4C254D2B1705817B4A1FF97BFA0F7DFD" authorityName="W. S. MacLeay" authorityYear="1819" box="[1351,1455,1755,1779]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="subFamily" subFamily="Abraeinae">Abraeinae</taxonomicName>
<taxonomicName id="4C254D2B1705817B4A1FF97BFA0F7DFD" authorityName="W. S. MacLeay" authorityYear="1819" box="[1351,1455,1755,1779]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subFamily" subFamily="Abraeinae">Abraeinae</taxonomicName>
, except
<taxonomicName id="4C254D2B1705817B4C20F95AFBB87C1C" authorityName="Wenzel" authorityYear="1944" box="[888,1048,1786,1810]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="tribe" tribe="Acritomorphini">Acritomorphini</taxonomicName>
<taxonomicName id="4C254D2B1705817B4C20F95AFBB87C1C" authorityName="Wenzel" authorityYear="1944" box="[888,1048,1786,1810]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="tribe" tribe="Acritomorphini">Acritomorphini</taxonomicName>
(
<bibRefCitation id="EFB44B591705817B4B77F95AFB1A7C1C" author="Wenzel RL" box="[1071,1210,1786,1810]" pageId="13" pagination="51 - 151" refId="ref19054" refString="Wenzel RL. On the classification of the histerid beetles. Fieldiana Zoology 1944; 28: 51 - 151. hups: // doi. org / 10.1093 / aesa / 37.3.375" type="journal article" year="1944">Wenzel 1944</bibRefCitation>
<bibRefCitation id="EFB44B591705817B4B77F95AFB1A7C1C" author="Wenzel RL" box="[1071,1210,1786,1810]" pageId="13" pageNumber="14" pagination="51 - 151" refId="ref19054" refString="Wenzel RL. On the classification of the histerid beetles. Fieldiana Zoology 1944; 28: 51 - 151. hups: // doi. org / 10.1093 / aesa / 37.3.375" type="journal article" year="1944">Wenzel 1944</bibRefCitation>
,
<bibRefCitation id="EFB44B591705817B4B94F95AFAEE7C1C" author="Mazur" box="[1228,1358,1786,1810]" pageId="13" pagination="27 - 32" refId="ref18506" refString="Mazur º. New species of histerid beetles (Coleoptera, Histeridae). Annales Zoologici 1997; 38: 27 - 32. hup: // rcin. org. pl" type="journal article" year="1997">Mazur 1997</bibRefCitation>
<bibRefCitation id="EFB44B591705817B4B94F95AFAEE7C1C" author="Mazur" box="[1228,1358,1786,1810]" pageId="13" pageNumber="14" pagination="27 - 32" refId="ref18506" refString="Mazur º. New species of histerid beetles (Coleoptera, Histeridae). Annales Zoologici 1997; 38: 27 - 32. hup: // rcin. org. pl" type="journal article" year="1997">Mazur 1997</bibRefCitation>
). Recent
<taxonomicName id="4C254D2B1705817B4C72F8B9FC6C7C3F" authorityName="Wenzel" authorityYear="1944" box="[810,972,1817,1841]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="tribe" tribe="Acritomorphini">Acritomorphini</taxonomicName>
<taxonomicName id="4C254D2B1705817B4C72F8B9FC6C7C3F" authorityName="Wenzel" authorityYear="1944" box="[810,972,1817,1841]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="tribe" tribe="Acritomorphini">Acritomorphini</taxonomicName>
, however, have an oblong, elongated body shape. Hence, the placement within a tribe is uncertain.
</paragraph>
<paragraph id="8B9A36A8170581784C72F8D3FA9A7CBA" blockId="13.[810,1459,1906,1961]" lastBlockId="14.[825,1475,1885,1972]" lastPageId="14" pageId="13">
<emphasis id="B951EABA1705817B4C72F8D3FC2A7C84" box="[810,906,1907,1930]" italics="true" pageId="13">Diagnosis:</emphasis>
Pronotum with antescutellar stria and small widely separated punctures.Elytra with four low costae and without an epipleural stria. Postmesocoxal stria complete, recurved along mesocoxa. Lateral metaventral stria complete, continuous onto metepisternum. Ŋe species can be distinguished from
<taxonomicName id="4C254D2B170681784EB6F83CFD0D7CBA" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[494,685,1948,1972]" class="Insecta" family="Histeridae" genus="Palaeabraeus" kingdom="Animalia" order="Coleoptera" pageId="14" phylum="Arthropoda" rank="species" species="glabrus">
<emphasis id="B951EABA170681784EB6F83CFD0D7CBA" box="[494,685,1948,1972]" italics="true" pageId="14">Palaeabraeus glabrus</emphasis>
</subSubSection>
<subSubSection id="C33F6523170581784C72F8D3FA9A7CBA" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="8B9A36A8170581784C72F8D3FA9A7CBA" blockId="13.[810,1459,1906,1961]" lastBlockId="14.[825,1475,1885,1972]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<emphasis id="B951EABA1705817B4C72F8D3FC2A7C84" box="[810,906,1907,1930]" italics="true" pageId="13" pageNumber="14">Diagnosis:</emphasis>
Pronotum with antescutellar stria and small widely separated punctures.Elytra with four low costae and without an epipleural stria. Postmesocoxal stria complete, recurved along mesocoxa. Lateral metaventral stria complete, continuous onto metepisternum. The species can be distinguished from
<taxonomicName id="4C254D2B170681784EB6F83CFD0D7CBA" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[494,685,1948,1972]" class="Insecta" family="Histeridae" genus="Palaeabraeus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="glabrus">
<emphasis id="B951EABA170681784EB6F83CFD0D7CBA" box="[494,685,1948,1972]" italics="true" pageId="14" pageNumber="15">Palaeabraeus glabrus</emphasis>
</taxonomicName>
(the only similar abraeine reported from Burmese ambers) by the presence of antescutellar stria, the anterior position of antennal cavities, presence of elytral costae, and absence of elytral epipleural stria.
</paragraph>
<caption id="DF5A6620170681784FD9F88BFC007C4D" box="[129,928,1835,1859]" pageId="14" startId="14.[129,194,1835,1859]" targetBox="[141,1461,143,1807]" targetPageId="14" targetType="figure">
<paragraph id="8B9A36A8170681784FD9F88BFC007C4D" blockId="14.[129,928,1835,1859]" box="[129,928,1835,1859]" pageId="14">
<emphasis id="B951EABA170681784FD9F88BFF457C4D" bold="true" box="[129,229,1835,1859]" pageId="14">Figure 10.</emphasis>
<taxonomicName id="4C254D2B170681784FB1F88BFE3F7C4D" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[233,415,1835,1859]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" phylum="Arthropoda" rank="species" species="planatus" status="sp. nov.">
<emphasis id="B951EABA170681784FB1F88BFE3F7C4D" box="[233,415,1835,1859]" italics="true" pageId="14">Phasmister planatus</emphasis>
</subSubSection>
<caption id="DF5A6620170681784FD9F88BFC007C4D" box="[129,928,1835,1859]" pageId="14" pageNumber="15" startId="14.[129,194,1835,1859]" targetBox="[141,1461,143,1807]" targetPageId="14" targetType="figure">
<paragraph id="8B9A36A8170681784FD9F88BFC007C4D" blockId="14.[129,928,1835,1859]" box="[129,928,1835,1859]" pageId="14" pageNumber="15">
<emphasis id="B951EABA170681784FD9F88BFF457C4D" bold="true" box="[129,229,1835,1859]" pageId="14" pageNumber="15">Figure 10.</emphasis>
<taxonomicName id="4C254D2B170681784FB1F88BFE3F7C4D" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[233,415,1835,1859]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="planatus" status="sp. nov.">
<emphasis id="B951EABA170681784FB1F88BFE3F7C4D" box="[233,415,1835,1859]" italics="true" pageId="14" pageNumber="15">Phasmister planatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C1170681784EFCF88BFE4A7C4C" box="[420,490,1835,1858]" pageId="14" rank="species">sp. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="A26257C1170681784EFCF88BFE4A7C4C" box="[420,490,1835,1858]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
(A–C) and
<taxonomicName id="4C254D2B170681784D04F88BFCB07C4D" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[604,784,1835,1859]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" phylum="Arthropoda" rank="species" species="kraliceki" status="sp. nov.">
<emphasis id="B951EABA170681784D04F88BFCB07C4D" box="[604,784,1835,1859]" italics="true" pageId="14">Phasmister kraliceki</emphasis>
<taxonomicName id="4C254D2B170681784D04F88BFCB07C4D" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[604,784,1835,1859]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="kraliceki" status="sp. nov.">
<emphasis id="B951EABA170681784D04F88BFCB07C4D" box="[604,784,1835,1859]" italics="true" pageId="14" pageNumber="15">Phasmister kraliceki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C1170681784C4CF88BFCFB7C4C" box="[788,859,1835,1858]" pageId="14" rank="species">sp. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="A26257C1170681784C4CF88BFCFB7C4C" box="[788,859,1835,1858]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
(D, E).
</paragraph>
</caption>
<paragraph id="8B9A36A8170781794F29FF30FD5B7BA9" blockId="15.[113,763,144,168]" box="[113,763,144,168]" pageId="15">
<emphasis id="B951EABA170781794F29FF30FF417BA6" box="[113,225,144,168]" italics="true" pageId="15">Etymology:</emphasis>
Ŋe specific name signifies the species’ extinct status.
<subSubSection id="C33F6523170781794F29FF30FD5B7BA9" box="[113,763,144,168]" pageId="15" pageNumber="16" type="etymology">
<paragraph id="8B9A36A8170781794F29FF30FD5B7BA9" blockId="15.[113,763,144,168]" box="[113,763,144,168]" pageId="15" pageNumber="16">
<emphasis id="B951EABA170781794F29FF30FF417BA6" box="[113,225,144,168]" italics="true" pageId="15" pageNumber="16">Etymology:</emphasis>
The specific name signifies the species’ extinct status.
</paragraph>
</subSubSection>
</treatment>

174
data/03/8C/87/038C87BE1704817A4CA7FF4EFAE97948.xml
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@ -1,125 +1,123 @@
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<mods:title id="72C16129B9C08B9058539E0BAF3DB7D9">A bonanza of Cretaceous fossils provides insights into the evolution of antennal protection in clown beetles (Coleoptera: Histeridae)</mods:title>
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<emphasis id="B951EABA1704817A4CA7FF4EFB307A07" bold="true" box="[1023,1168,238,265]" italics="true" pageId="12" pageNumber="13">Pseudacritus</emphasis>
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species.
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..
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ºmall, round and dorsoventrally convex body. Head without frontal stria, convex eyes, labrum with a pair of prominent setae. Antennal club annulate. Antennal cavities present at the anteriormost hypomeral angle. Pronotal disc with an antescutellar stria. Tarsal formula 5-5-5.
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<emphasis id="B951EABA1704817A4C61FEFAFC3F7A7F" box="[825,927,346,369]" italics="true" pageId="12" pageNumber="13">Diagnosis:</emphasis>
Small, round and dorsoventrally convex body. Head without frontal stria, convex eyes, labrum with a pair of prominent setae. Antennal club annulate. Antennal cavities present at the anteriormost hypomeral angle. Pronotal disc with an antescutellar stria. Tarsal formula 5-5-5.
</paragraph>
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<emphasis id="B951EABA1704817A4C61FDB0FC087926" box="[825,936,528,552]" italics="true" pageId="12">Etymology:</emphasis>
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<emphasis id="B951EABA1704817A4C61FDB0FC087926" box="[825,936,528,552]" italics="true" pageId="12" pageNumber="13">Etymology:</emphasis>
Generic name
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<emphasis id="B951EABA1704817A4B16FDB0FB697926" box="[1102,1225,528,552]" italics="true" pageId="12">Pseudacritus</emphasis>
<taxonomicName id="4C254D2B1704817A4B16FDB0FB697926" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[1102,1225,528,552]" class="Insecta" family="Histeridae" genus="Pseudacritus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA1704817A4B16FDB0FB697926" box="[1102,1225,528,552]" italics="true" pageId="12" pageNumber="13">Pseudacritus</emphasis>
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(masculine) points out the striking resemblance to the recent genus
<taxonomicName id="4C254D2B1704817A4BA5FD8FFAE37948" box="[1277,1347,559,582]" class="Insecta" family="Histeridae" genus="Acritus" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA1704817A4BA5FD8FFAE37948" box="[1277,1347,559,582]" italics="true" pageId="12">Acritus</emphasis>
<taxonomicName id="4C254D2B1704817A4BA5FD8FFAE37948" box="[1277,1347,559,582]" class="Insecta" family="Histeridae" genus="Acritus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA1704817A4BA5FD8FFAE37948" box="[1277,1347,559,582]" italics="true" pageId="12" pageNumber="13">Acritus</emphasis>
</taxonomicName>
.
</paragraph>

311
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<taxonomicName id="4C254D2B170781794C9CFE52FB1D7902" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[964,1213,498,525]" class="Insecta" family="Histeridae" genus="Prodigister" kingdom="Animalia" order="Coleoptera" pageId="15" phylum="Arthropoda" rank="species" species="tricostatus" status="sp. nov.">
<emphasis id="B951EABA170781794C9CFE52FB1D7902" bold="true" box="[964,1213,498,525]" italics="true" pageId="15">Prodigister tricostatus</emphasis>
<treatment id="038C87BE170781674C9CFE52FE227CBF" LSID="urn:lsid:plazi:treatment:038C87BE170781674C9CFE52FE227CBF" httpUri="http://treatment.plazi.org/id/038C87BE170781674C9CFE52FE227CBF" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16">
<subSubSection id="C33F6523170781794C9CFE52FAB97902" box="[964,1305,498,525]" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="8B9A36A8170781794C9CFE52FAB97902" blockId="15.[964,1305,498,525]" box="[964,1305,498,525]" pageId="15" pageNumber="16">
<heading id="D0D281C4170781794C9CFE52FAB97902" bold="true" box="[964,1305,498,525]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="16" reason="2">
<emphasis id="B951EABA170781794C9CFE52FAB97902" bold="true" box="[964,1305,498,525]" pageId="15" pageNumber="16">
<taxonomicName id="4C254D2B170781794C9CFE52FB1D7902" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[964,1213,498,525]" class="Insecta" family="Histeridae" genus="Prodigister" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="tricostatus" status="sp. nov.">
<emphasis id="B951EABA170781794C9CFE52FB1D7902" bold="true" box="[964,1213,498,525]" italics="true" pageId="15" pageNumber="16">Prodigister tricostatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C1170781794B9AFE52FAB97902" box="[1218,1305,498,524]" pageId="15" rank="species">sp. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="A26257C1170781794B9AFE52FAB97902" box="[1218,1305,498,524]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33F6523170781674B18FD81FE227CBF" lastPageId="17" pageId="15" type="description">
<paragraph id="8B9A36A8170781794B18FD81FB3D7935" blockId="15.[1088,1181,545,571]" box="[1088,1181,545,571]" pageId="15">
<emphasis id="B951EABA170781794B18FD81FB3D7935" bold="true" box="[1088,1181,545,571]" pageId="15">
<subSubSection id="C33F6523170781794B18FD81FB3D7935" box="[1088,1181,545,571]" pageId="15" pageNumber="16" type="description">
<paragraph id="8B9A36A8170781794B18FD81FB3D7935" blockId="15.[1088,1181,545,571]" box="[1088,1181,545,571]" pageId="15" pageNumber="16">
<emphasis id="B951EABA170781794B18FD81FB3D7935" bold="true" box="[1088,1181,545,571]" pageId="15" pageNumber="16">
(
<figureCitation id="131E2A2D170781794B13FD81FB327935" box="[1099,1170,545,571]" captionStart="Figure 11" captionStartId="16.[129,194,1836,1860]" captionTargetBox="[213,1389,144,1808]" captionTargetId="figure-8@16.[213,1389,144,1808]" captionTargetPageId="16" captionText="Figure 11. Prodigister tricostatus sp. nov.. A, dorsal view.B, lateroventral view.C, lateral view. D, elytra.E, frontal view.F, mouthparts. Abbreviations:C1–C3, costae 1–3; ds, dorsal setae; gal, galea; lab, labrum; man, mandible; mp, maxillary palpus; mt, mesotibia; pt, protibia." pageId="15">Fig. 11</figureCitation>
<figureCitation id="131E2A2D170781794B13FD81FB327935" box="[1099,1170,545,571]" captionStart="Figure 11" captionStartId="16.[129,194,1836,1860]" captionTargetBox="[213,1389,144,1808]" captionTargetId="figure-8@16.[213,1389,144,1808]" captionTargetPageId="16" captionText="Figure 11. Prodigister tricostatus sp. nov.. A, dorsal view.B, lateroventral view.C, lateral view. D, elytra.E, frontal view.F, mouthparts. Abbreviations:C1–C3, costae 1–3; ds, dorsal setae; gal, galea; lab, labrum; man, mandible; mp, maxillary palpus; mt, mesotibia; pt, protibia." pageId="15" pageNumber="16">Fig. 11</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8B9A36A8170781794C72FDC2FAF979B7" blockId="15.[810,1460,610,697]" pageId="15">
<emphasis id="B951EABA170781794C72FDC2FC267974" box="[810,902,610,634]" italics="true" pageId="15">Material:</emphasis>
<collectingRegion id="49E1F84A170781794CD5FDC2FC777974" box="[909,983,610,634]" country="Myanmar" name="Kachin" pageId="15">Kachin</collectingRegion>
</subSubSection>
<subSubSection id="C33F6523170781794C72FDC2FAF979B7" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="8B9A36A8170781794C72FDC2FAF979B7" blockId="15.[810,1460,610,697]" pageId="15" pageNumber="16">
<emphasis id="B951EABA170781794C72FDC2FC267974" box="[810,902,610,634]" italics="true" pageId="15" pageNumber="16">Material:</emphasis>
<materialsCitation id="3B4D3CF5170781794CD5FDC2FAF579B7" collectionCode="HUM" location="Kachin" pageId="15" pageNumber="16" specimenCode="SEHU-0000121220" specimenCount="1" stateProvince="Kachin">
<collectingRegion id="49E1F84A170781794CD5FDC2FC777974" box="[909,983,610,634]" country="Myanmar" name="Kachin" pageId="15" pageNumber="16">Kachin</collectingRegion>
amber (
<emphasis id="B951EABA170781794B69FDC3FB9A7974" box="[1073,1082,611,634]" italics="true" pageId="15">c</emphasis>
<emphasis id="B951EABA170781794B69FDC3FB9A7974" box="[1073,1082,611,634]" italics="true" pageId="15" pageNumber="16">c</emphasis>
. 99 Mya). Amber piece dimensions: 15 × 8 ×
<quantity id="4CDD9B4D170781794CD2FD21FC667997" box="[906,966,641,665]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="15" unit="mm" value="2.0">2 mm</quantity>
. Amber clear with several insect fragments and air bubbles. Deposited at HUM (ºEHU-0000121220).
<quantity id="4CDD9B4D170781794CD2FD21FC667997" box="[906,966,641,665]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="16" unit="mm" value="2.0">2 mm</quantity>
. Amber clear with several insect fragments and air bubbles. Deposited at
<collectionCode id="ED34AE6D170781794B74FD01FBCB79B6" box="[1068,1131,673,696]" pageId="15" pageNumber="16">HUM</collectionCode>
(
<specimenCode id="DB839ED3170781794B2FFD01FAEE79B7" box="[1143,1358,673,697]" pageId="15" pageNumber="16">SEHU-0000121220</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B9A36A8170781674C72FD7BFD9F7973" blockId="15.[810,1460,730,1976]" lastBlockId="17.[113,763,144,638]" lastPageId="17" pageId="15">
<emphasis id="B951EABA170781794C72FD7BFC0179FC" box="[810,929,731,754]" italics="true" pageId="15">Dimensions.</emphasis>
</subSubSection>
<subSubSection id="C33F6523170781674C72FD7BFD9F7973" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16" type="description">
<paragraph id="8B9A36A8170781674C72FD7BFD9F7973" blockId="15.[810,1460,730,1976]" lastBlockId="17.[113,763,144,638]" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16">
<emphasis id="B951EABA170781794C72FD7BFC0179FC" box="[810,929,731,754]" italics="true" pageId="15" pageNumber="16">Dimensions.</emphasis>
Head width:
<quantity id="4CDD9B4D170781794B12FD7AFB3879FF" box="[1098,1176,730,754]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="15" unit="mm" value="0.4">0.4 mm</quantity>
<quantity id="4CDD9B4D170781794B12FD7AFB3879FF" box="[1098,1176,730,754]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="15" pageNumber="16" unit="mm" value="0.4">0.4 mm</quantity>
; width between anterior pronotal angles:
<quantity id="4CDD9B4D170781794C86FD59FB99781F" box="[990,1081,761,785]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="15" unit="mm" value="0.42">0.42 mm</quantity>
<quantity id="4CDD9B4D170781794C86FD59FB99781F" box="[990,1081,761,785]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="15" pageNumber="16" unit="mm" value="0.42">0.42 mm</quantity>
; width between posterior pronotal angles:
<quantity id="4CDD9B4D170781794CDCFCB9FC72783E" box="[900,978,793,817]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="15" unit="mm" value="0.7">0.7 mm</quantity>
<quantity id="4CDD9B4D170781794CDCFCB9FC72783E" box="[900,978,793,817]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="15" pageNumber="16" unit="mm" value="0.7">0.7 mm</quantity>
; pronotal length:
<quantity id="4CDD9B4D170781794BEAFCB8FB5F783E" box="[1202,1279,792,816]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="15" unit="mm" value="0.5">0.5 mm</quantity>
<quantity id="4CDD9B4D170781794BEAFCB8FB5F783E" box="[1202,1279,792,816]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="16" unit="mm" value="0.5">0.5 mm</quantity>
; sutural elytral length:
<quantity id="4CDD9B4D170781794C24FC98FC777841" box="[892,983,824,848]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" pageId="15" unit="mm" value="0.95">0.95 mm</quantity>
<quantity id="4CDD9B4D170781794C24FC98FC777841" box="[892,983,824,848]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" pageId="15" pageNumber="16" unit="mm" value="0.95">0.95 mm</quantity>
; elytral width (in pair, across widest point):
<quantity id="4CDD9B4D170781794C72FCF7FC277861" box="[810,903,855,879]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="15" unit="mm" value="0.75">0.75 mm</quantity>
<quantity id="4CDD9B4D170781794C72FCF7FC277861" box="[810,903,855,879]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="15" pageNumber="16" unit="mm" value="0.75">0.75 mm</quantity>
.—
<emphasis id="B951EABA170781794CFEFCF7FC467861" bold="true" box="[934,998,855,879]" italics="true" pageId="15">Body:</emphasis>
Elongate, subparallel-sided, almost rectangular in shape. Pronotum irregularly sculptured. Elytra with several very prominent costae. Dorsal cuticle with fine setation. Ŋorax and abdomen strongly compressed (deformed) laterally.—
<emphasis id="B951EABA170781794CAEFC74FB9778E2" bold="true" box="[1014,1079,980,1004]" italics="true" pageId="15">Head:</emphasis>
Frons and clypeus with sparse setae. Frontoclypeal suture absent. Frontal stria complete, weakly carinate. Labrum long (width: length ratio is 3: 2), plurisetose and punctuate. Mandibles with several prominent setae on outer margin and a blunt subapical tooth, prostheca well developed. Terminal maxillary palpomere elongate and pointed, as long as palpomeres 2–3 combined. ºcape straight and parallel-sided with several setae apically. Pedicel short and bulbous, as long as antennomeres 3–4 combined, wider basally than distally. Antennomere 3 short (about half of antennomere 4 length). Antennomeres 3–8 becoming wider apically. Antennal club round and flaưened, setose, without annuli.—
<emphasis id="B951EABA170781794BAAFA8DFAE77E4B" bold="true" box="[1266,1351,1325,1349]" italics="true" pageId="15">Ŋorax:</emphasis>
Pronotum with irregular sculpturing, mostly convex medially, with a pair of large depressions posterolaterally and one mediobasally, elevations on either side aligned with longitudinal elytral ridges; pronotal sides margined, sinuate, widening posterad, narrowed anterad; marginal pronotal striae not evident. Basal pronotal margin rounded. Marginal prosternal stria complete. Pronotum sparsely covered with fine, short setae. ºcutellar shield not visible. Elytra with fine punctuation and vestiture (setae 5× as long as in pronotum). Dorsal elytral striation not visible. Epipleural stria complete. Outer elytral margin carinate and weakly elevated. Medioanterior part strongly depressed. ºutural area strongly elevated forming a keel that is sharp on anterior two-thirds, thence gradually diminishing posterad; each elytron with strong lateral longitudinal costa, rather sharp in anterior half of elytra, becoming lower and broader apically; elytral apices obscured by cracks and evidently distorted. Elytral epipleural stria complete. Ventral body surface strongly compressed, creating a (probably artificial) keel. Prosternal lobe short and broad, punctuate. Ŋe presence and character of antennal cavity cannot be determined due to the ventral deformation. Prosternal keel without apparent striation or punctuation. Mesoventrite strongly compressed ventrally, striation not visible. Metaventrite finely punctuate, lateral metaventral stria visible on anterior third.—
<emphasis id="B951EABA171981674F9DFF4EFF5C7A0B" bold="true" box="[197,252,238,261]" italics="true" pageId="17">Legs:</emphasis>
Profemur very wide, flaưened. Protibia strongly dilated with denticles along its outer margin. Protarsal groove present. Protibial spur not developed. Terminal protarsomere as long as protarsomeres 1–4 combined. Mesofemur flaưened, broad basally and becoming narrower apically. Mesotibia approximately half of the mesofemoral length, slightly dilated with thick setae along the outer margin, diminishing in size basally. Terminal mesotarsomere as long as mesotarsomeres 2–4 combined. Metafemur broad. Metatibia narrow with only two to three thick setae apically. Metatarsus identical to mesotarsus.—
<emphasis id="B951EABA171981674F29FD87FF7E7931" bold="true" box="[113,222,551,575]" italics="true" pageId="17">Abdomen:</emphasis>
<emphasis id="B951EABA170781794CFEFCF7FC467861" bold="true" box="[934,998,855,879]" italics="true" pageId="15" pageNumber="16">Body:</emphasis>
Elongate, subparallel-sided, almost rectangular in shape. Pronotum irregularly sculptured. Elytra with several very prominent costae. Dorsal cuticle with fine setation. Thorax and abdomen strongly compressed (deformed) laterally.—
<emphasis id="B951EABA170781794CAEFC74FB9778E2" bold="true" box="[1014,1079,980,1004]" italics="true" pageId="15" pageNumber="16">Head:</emphasis>
Frons and clypeus with sparse setae. Frontoclypeal suture absent. Frontal stria complete, weakly carinate. Labrum long (width: length ratio is 3: 2), plurisetose and punctuate. Mandibles with several prominent setae on outer margin and a blunt subapical tooth, prostheca well developed. Terminal maxillary palpomere elongate and pointed, as long as palpomeres 2–3 combined. Scape straight and parallel-sided with several setae apically. Pedicel short and bulbous, as long as antennomeres 3–4 combined, wider basally than distally. Antennomere 3 short (about half of antennomere 4 length). Antennomeres 3–8 becoming wider apically. Antennal club round and flattened, setose, without annuli.—
<emphasis id="B951EABA170781794BAAFA8DFAE77E4B" bold="true" box="[1266,1351,1325,1349]" italics="true" pageId="15" pageNumber="16">Thorax:</emphasis>
Pronotum with irregular sculpturing, mostly convex medially, with a pair of large depressions posterolaterally and one mediobasally, elevations on either side aligned with longitudinal elytral ridges; pronotal sides margined, sinuate, widening posterad, narrowed anterad; marginal pronotal striae not evident. Basal pronotal margin rounded. Marginal prosternal stria complete. Pronotum sparsely covered with fine, short setae. Scutellar shield not visible. Elytra with fine punctuation and vestiture (setae 5× as long as in pronotum). Dorsal elytral striation not visible. Epipleural stria complete. Outer elytral margin carinate and weakly elevated. Medioanterior part strongly depressed. Sutural area strongly elevated forming a keel that is sharp on anterior two-thirds, thence gradually diminishing posterad; each elytron with strong lateral longitudinal costa, rather sharp in anterior half of elytra, becoming lower and broader apically; elytral apices obscured by cracks and evidently distorted. Elytral epipleural stria complete. Ventral body surface strongly compressed, creating a (probably artificial) keel. Prosternal lobe short and broad, punctuate. The presence and character of antennal cavity cannot be determined due to the ventral deformation. Prosternal keel without apparent striation or punctuation. Mesoventrite strongly compressed ventrally, striation not visible. Metaventrite finely punctuate, lateral metaventral stria visible on anterior third.—
<emphasis id="B951EABA171981674F9DFF4EFF5C7A0B" bold="true" box="[197,252,238,261]" italics="true" pageId="17" pageNumber="18">Legs:</emphasis>
Profemur very wide, flattened. Protibia strongly dilated with denticles along its outer margin. Protarsal groove present. Protibial spur not developed. Terminal protarsomere as long as protarsomeres 1–4 combined. Mesofemur flattened, broad basally and becoming narrower apically. Mesotibia approximately half of the mesofemoral length, slightly dilated with thick setae along the outer margin, diminishing in size basally. Terminal mesotarsomere as long as mesotarsomeres 2–4 combined. Metafemur broad. Metatibia narrow with only two to three thick setae apically. Metatarsus identical to mesotarsus.—
<emphasis id="B951EABA171981674F29FD87FF7E7931" bold="true" box="[113,222,551,575]" italics="true" pageId="17" pageNumber="18">Abdomen:</emphasis>
First visible abdominal sternite long and wide, finely punctuate. Rest of the abdomen hardly observable due to cracks and contamination. Pygidium weakly costate.
</paragraph>
<caption id="DF5A6620171881664FD9F88CFA087C6E" pageId="16" startId="16.[129,194,1836,1860]" targetBox="[213,1389,144,1808]" targetPageId="16" targetType="figure">
<paragraph id="8B9A36A8171881664FD9F88CFA087C6E" blockId="16.[129,1448,1836,1888]" pageId="16">
<emphasis id="B951EABA171881664FD9F88CFF457C4A" bold="true" box="[129,229,1836,1860]" pageId="16">Figure 11.</emphasis>
<taxonomicName id="4C254D2B171881664FB1F88CFE0B7C4A" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[233,427,1836,1860]" class="Insecta" family="Histeridae" genus="Prodigister" kingdom="Animalia" order="Coleoptera" pageId="16" phylum="Arthropoda" rank="species" species="tricostatus" status="sp. nov.">
<emphasis id="B951EABA171881664FB1F88CFE0B7C4A" box="[233,427,1836,1860]" italics="true" pageId="16">Prodigister tricostatus</emphasis>
</subSubSection>
<caption id="DF5A6620171881664FD9F88CFA087C6E" pageId="16" pageNumber="17" startId="16.[129,194,1836,1860]" targetBox="[213,1389,144,1808]" targetPageId="16" targetType="figure">
<paragraph id="8B9A36A8171881664FD9F88CFA087C6E" blockId="16.[129,1448,1836,1888]" pageId="16" pageNumber="17">
<emphasis id="B951EABA171881664FD9F88CFF457C4A" bold="true" box="[129,229,1836,1860]" pageId="16" pageNumber="17">Figure 11.</emphasis>
<taxonomicName id="4C254D2B171881664FB1F88CFE0B7C4A" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[233,427,1836,1860]" class="Insecta" family="Histeridae" genus="Prodigister" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="tricostatus" status="sp. nov.">
<emphasis id="B951EABA171881664FB1F88CFE0B7C4A" box="[233,427,1836,1860]" italics="true" pageId="16" pageNumber="17">Prodigister tricostatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C1171881664EE8F88CFE517C4D" box="[432,497,1836,1859]" pageId="16" rank="species">sp. nov</taxonomicNameLabel>
<taxonomicNameLabel id="A26257C1171881664EE8F88CFE517C4D" box="[432,497,1836,1859]" pageId="16" pageNumber="17" rank="species">sp. nov</taxonomicNameLabel>
.. A, dorsal view. B, lateroventral view. C, lateral view. D, elytra. E, frontal view. F, mouthparts. Abbreviations: C1–C3, costae 1–3; ds, dorsal setae; gal, galea; lab, labrum; man, mandible; mp, maxillary palpus; mt, mesotibia; pt, protibia.
</paragraph>
</caption>
<paragraph id="8B9A36A8171981674F29FD00FF7F7D2A" blockId="17.[113,763,671,1572]" pageId="17">
<emphasis id="B951EABA171981674F29FD00FEFF79B9" box="[113,351,672,695]" italics="true" pageId="17">Taxonomic assignment:</emphasis>
Few characters that would serve as diagnostic at higher levels are clearly visible in this specimen. Furthermore, it is considerably deformed by lateral compression, especially ventrally, and sternal characters are severely altered. Ŋe strongly convex pronotum and costate elytra, being reasonably bilaterally symmetrical, show no signs of being artefacts of deformation, and distinguish this taxon from any known histerids, extinct or extant. Ŋese are not similar to costae in onthophiline genera, and this genus could not be placed there. Ŋe antennal club is not easily visible, but it is sufficiently clear that it lacks distinct annuli, as are seen in a number of other Cretaceous ‘basal grade’ histeroids. Ŋe strongly expanded protibiae are a remarkably common feature among Cretaceous taxa, seen in
<taxonomicName id="4C254D2B171981674E41FB96FEC87F40" authorityName="Simon Prazak &amp; Lackner" authorityYear="2023" box="[281,360,1078,1102]" class="Insecta" family="Histeridae" genus="Olexum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA171981674E41FB96FEC87F40" box="[281,360,1078,1102]" italics="true" pageId="17">Olexum</emphasis>
<subSubSection id="C33F6523171981674F29FD00FF7F7D2A" pageId="17" pageNumber="18" type="discussion">
<paragraph id="8B9A36A8171981674F29FD00FF7F7D2A" blockId="17.[113,763,671,1572]" pageId="17" pageNumber="18">
<emphasis id="B951EABA171981674F29FD00FEFF79B9" box="[113,351,672,695]" italics="true" pageId="17" pageNumber="18">Taxonomic assignment:</emphasis>
Few characters that would serve as diagnostic at higher levels are clearly visible in this specimen. Furthermore, it is considerably deformed by lateral compression, especially ventrally, and sternal characters are severely altered. The strongly convex pronotum and costate elytra, being reasonably bilaterally symmetrical, show no signs of being artefacts of deformation, and distinguish this taxon from any known histerids, extinct or extant. These are not similar to costae in onthophiline genera, and this genus could not be placed there. The antennal club is not easily visible, but it is sufficiently clear that it lacks distinct annuli, as are seen in a number of other Cretaceous ‘basal grade’ histeroids. The strongly expanded protibiae are a remarkably common feature among Cretaceous taxa, seen in
<taxonomicName id="4C254D2B171981674E41FB96FEC87F40" authorityName="Simon Prazak &amp; Lackner" authorityYear="2023" box="[281,360,1078,1102]" class="Insecta" family="Histeridae" genus="Olexum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA171981674E41FB96FEC87F40" box="[281,360,1078,1102]" italics="true" pageId="17" pageNumber="18">Olexum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D2B171981674E27FB96FE687F40" authorityName="Caterino" authorityYear="2021" box="[383,456,1078,1102]" class="Insecta" family="Histeridae" genus="Yethiha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA171981674E27FB96FE687F40" box="[383,456,1078,1102]" italics="true" pageId="17">Yethiha</emphasis>
<taxonomicName id="4C254D2B171981674E27FB96FE687F40" authorityName="Caterino" authorityYear="2021" box="[383,456,1078,1102]" class="Insecta" family="Histeridae" genus="Yethiha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA171981674E27FB96FE687F40" box="[383,456,1078,1102]" italics="true" pageId="17" pageNumber="18">Yethiha</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D2B171981674E87FB97FD9A7F40" authorityName="Caterino" authorityYear="2021" box="[479,570,1079,1102]" class="Insecta" family="Histeridae" genus="Druantia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA171981674E87FB97FD9A7F40" box="[479,570,1079,1102]" italics="true" pageId="17">Druantia</emphasis>
<taxonomicName id="4C254D2B171981674E87FB97FD9A7F40" authorityName="Caterino" authorityYear="2021" box="[479,570,1079,1102]" class="Insecta" family="Histeridae" genus="Druantia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA171981674E87FB97FD9A7F40" box="[479,570,1079,1102]" italics="true" pageId="17" pageNumber="18">Druantia</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D2B171981674D09FB96FD1C7F40" authorityName="Simon Prazak &amp; Lackner" authorityYear="2023" box="[593,700,1078,1102]" class="Insecta" family="Histeridae" genus="Platycretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA171981674D09FB96FD1C7F40" box="[593,700,1078,1102]" italics="true" pageId="17">Platycretus</emphasis>
<taxonomicName id="4C254D2B171981674D09FB96FD1C7F40" authorityName="Simon Prazak &amp; Lackner" authorityYear="2023" box="[593,700,1078,1102]" class="Insecta" family="Histeridae" genus="Platycretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA171981674D09FB96FD1C7F40" box="[593,700,1078,1102]" italics="true" pageId="17" pageNumber="18">Platycretus</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C254D2B171981674F29FBF6FF567F63" box="[113,246,1110,1133]" class="Insecta" family="Histeridae" genus="Promyrmister" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA171981674F29FBF6FF567F63" box="[113,246,1110,1133]" italics="true" pageId="17">Promyrmister</emphasis>
<taxonomicName id="4C254D2B171981674F29FBF6FF567F63" box="[113,246,1110,1133]" class="Insecta" family="Histeridae" genus="Promyrmister" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA171981674F29FBF6FF567F63" box="[113,246,1110,1133]" italics="true" pageId="17" pageNumber="18">Promyrmister</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFB44B59171981674E5EFBF6FE0C7F60" author="Zhou YL &amp; Slipinski A &amp; Ren D" box="[262,428,1110,1134]" pageId="17" refId="ref19248" refString="Zhou YL, Slipinski A, Ren D et al. A Mesozoic clown beetle myrmecophile (Coleoptera: Histeridae). eLife 2019; 8: e 44985. hups: // doi. org / 10.7554 / eLife. 44985" type="journal volume" year="2019">
<bibRefCitation id="EFB44B59171981674E5EFBF6FE0C7F60" author="Zhou YL &amp; Slipinski A &amp; Ren D" box="[262,428,1110,1134]" pageId="17" pageNumber="18" refId="ref19248" refString="Zhou YL, Slipinski A, Ren D et al. A Mesozoic clown beetle myrmecophile (Coleoptera: Histeridae). eLife 2019; 8: e 44985. hups: // doi. org / 10.7554 / eLife. 44985" type="journal volume" year="2019">
Zhou
<emphasis id="B951EABA171981674E1CFBF6FED37F60" box="[324,371,1110,1134]" italics="true" pageId="17">et al.</emphasis>
<emphasis id="B951EABA171981674E1CFBF6FED37F60" box="[324,371,1110,1134]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2019
</bibRefCitation>
,
<bibRefCitation id="EFB44B59171981674EEFFBF6FDEA7F60" author="Caterino" box="[439,586,1110,1134]" pageId="17" pagination="211 - 21" refId="ref17393" refString="Caterino Mº. New fossil histerid species from Cretaceous Burmese amber (Coleoptera: Histeridae). Ne Coleopterists Bulletin 2021; 75: 211 - 21." type="journal article" year="2021">Caterino 2021</bibRefCitation>
, ºimon-Pražák
<emphasis id="B951EABA171981674DB1FBF6FF2A7F83" italics="true" pageId="17">et al.</emphasis>
<bibRefCitation id="EFB44B59171981674EEFFBF6FDEA7F60" author="Caterino" box="[439,586,1110,1134]" pageId="17" pageNumber="18" pagination="211 - 21" refId="ref17393" refString="Caterino Mº. New fossil histerid species from Cretaceous Burmese amber (Coleoptera: Histeridae). Ne Coleopterists Bulletin 2021; 75: 211 - 21." type="journal article" year="2021">Caterino 2021</bibRefCitation>
, Simon-Pražák
<emphasis id="B951EABA171981674DB1FBF6FF2A7F83" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2023b). Of these, the species bears some characters linking it to
<taxonomicName id="4C254D2B171981674FFFFB34FEB27FA2" authorityName="Simon Prazak &amp; Lackner" authorityYear="2023" box="[167,274,1172,1196]" class="Insecta" family="Histeridae" genus="Platycretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA171981674FFFFB34FEB27FA2" box="[167,274,1172,1196]" italics="true" pageId="17">Platycretus</emphasis>
<taxonomicName id="4C254D2B171981674FFFFB34FEB27FA2" authorityName="Simon Prazak &amp; Lackner" authorityYear="2023" box="[167,274,1172,1196]" class="Insecta" family="Histeridae" genus="Platycretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA171981674FFFFB34FEB27FA2" box="[167,274,1172,1196]" italics="true" pageId="17" pageNumber="18">Platycretus</emphasis>
</taxonomicName>
: very wide and flat profemur, and mesofemur longer than mesotibia (ºimon-Pražák
<emphasis id="B951EABA171981674D5BFB14FD967FC2" box="[515,566,1204,1228]" italics="true" pageId="17">et al.</emphasis>
: very wide and flat profemur, and mesofemur longer than mesotibia (Simon-Pražák
<emphasis id="B951EABA171981674D5BFB14FD967FC2" box="[515,566,1204,1228]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2023b). However, the plurisetose labrum, as well as fused antennal club, would suggest a closer relationship to
<taxonomicName id="4C254D2B171981674EEFFB52FDF97E04" authorityName="Reitter" authorityYear="1909" box="[439,601,1266,1290]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="17" phylum="Arthropoda" rank="subFamily" subFamily="Dendrophilinae">Dendrophilinae</taxonomicName>
<taxonomicName id="4C254D2B171981674EEFFB52FDF97E04" authorityName="Reitter" authorityYear="1909" box="[439,601,1266,1290]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="subFamily" subFamily="Dendrophilinae">Dendrophilinae</taxonomicName>
taxa—
<taxonomicName id="4C254D2B171981674DFEFB52FD557E04" authorityName="Simon Prazak &amp; Lackner" authorityYear="2023" box="[678,757,1266,1290]" class="Insecta" family="Histeridae" genus="Olexum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA171981674DFEFB52FD557E04" box="[678,757,1266,1290]" italics="true" pageId="17">Olexum</emphasis>
<taxonomicName id="4C254D2B171981674DFEFB52FD557E04" authorityName="Simon Prazak &amp; Lackner" authorityYear="2023" box="[678,757,1266,1290]" class="Insecta" family="Histeridae" genus="Olexum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA171981674DFEFB52FD557E04" box="[678,757,1266,1290]" italics="true" pageId="17" pageNumber="18">Olexum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D2B171981674F29FAB2FF1A7E24" authorityName="Caterino" authorityYear="2021" box="[113,186,1298,1322]" class="Insecta" family="Histeridae" genus="Yethiha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA171981674F29FAB2FF1A7E24" box="[113,186,1298,1322]" italics="true" pageId="17">Yethiha</emphasis>
<taxonomicName id="4C254D2B171981674F29FAB2FF1A7E24" authorityName="Caterino" authorityYear="2021" box="[113,186,1298,1322]" class="Insecta" family="Histeridae" genus="Yethiha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA171981674F29FAB2FF1A7E24" box="[113,186,1298,1322]" italics="true" pageId="17" pageNumber="18">Yethiha</emphasis>
</taxonomicName>
, or
<taxonomicName id="4C254D2B171981674F87FAB2FE9A7E27" authorityName="Caterino" authorityYear="2021" box="[223,314,1298,1321]" class="Insecta" family="Histeridae" genus="Druantia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA171981674F87FAB2FE9A7E27" box="[223,314,1298,1321]" italics="true" pageId="17">Druantia</emphasis>
<taxonomicName id="4C254D2B171981674F87FAB2FE9A7E27" authorityName="Caterino" authorityYear="2021" box="[223,314,1298,1321]" class="Insecta" family="Histeridae" genus="Druantia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA171981674F87FAB2FE9A7E27" box="[223,314,1298,1321]" italics="true" pageId="17" pageNumber="18">Druantia</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFB44B59171981674E11FAB2FE7C7E24" author="Caterino" box="[329,476,1298,1322]" pageId="17" pagination="211 - 21" refId="ref17393" refString="Caterino Mº. New fossil histerid species from Cretaceous Burmese amber (Coleoptera: Histeridae). Ne Coleopterists Bulletin 2021; 75: 211 - 21." type="journal article" year="2021">Caterino 2021</bibRefCitation>
, ºimon-Pražák
<emphasis id="B951EABA171981674D20FAB2FD067E24" box="[632,678,1298,1322]" italics="true" pageId="17">et al.</emphasis>
2023b). Ŋe position of this genus could not be sufficiently resolved via phylogenetic analyses either, as it nested in various positions in
<taxonomicName id="4C254D2B171981674F29FAD0FF7C7E86" box="[113,220,1392,1416]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="17" phylum="Arthropoda" rank="family">Histeridae</taxonomicName>
<bibRefCitation id="EFB44B59171981674E11FAB2FE7C7E24" author="Caterino" box="[329,476,1298,1322]" pageId="17" pageNumber="18" pagination="211 - 21" refId="ref17393" refString="Caterino Mº. New fossil histerid species from Cretaceous Burmese amber (Coleoptera: Histeridae). Ne Coleopterists Bulletin 2021; 75: 211 - 21." type="journal article" year="2021">Caterino 2021</bibRefCitation>
, Simon-Pražák
<emphasis id="B951EABA171981674D20FAB2FD067E24" box="[632,678,1298,1322]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2023b). The position of this genus could not be sufficiently resolved via phylogenetic analyses either, as it nested in various positions in
<taxonomicName id="4C254D2B171981674F29FAD0FF7C7E86" box="[113,220,1392,1416]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="family">Histeridae</taxonomicName>
outside the
<taxonomicName id="4C254D2B171981674E38FAD0FE4F7E86" authorityName="Zhou, Yu-Lingxi, Caterino, Dong Ren &amp; Ślipiński" authorityYear="2020" box="[352,495,1392,1416]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="17" phylum="Arthropoda" rank="subFamily" subFamily="Antigracilinae">Antigracilinae</taxonomicName>
<taxonomicName id="4C254D2B171981674E38FAD0FE4F7E86" authorityName="Zhou, Yu-Lingxi, Caterino, Dong Ren &amp; Ślipiński" authorityYear="2020" box="[352,495,1392,1416]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="subFamily" subFamily="Antigracilinae">Antigracilinae</taxonomicName>
and
<taxonomicName id="4C254D2B171981674D7DFAD0FD2D7E86" authorityName="W. S. MacLeay" authorityYear="1819" box="[549,653,1392,1416]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="17" phylum="Arthropoda" rank="subFamily" subFamily="Abraeinae">Abraeinae</taxonomicName>
clades. As some of the key characters for higher taxonomic assignment are not visible due to deformation, we leave this genus unplaced to a higher taxon for now, and hope that beưer preserved specimens materialize to reveal more phylogenetically significant characters.
<taxonomicName id="4C254D2B171981674D7DFAD0FD2D7E86" authorityName="W. S. MacLeay" authorityYear="1819" box="[549,653,1392,1416]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="subFamily" subFamily="Abraeinae">Abraeinae</taxonomicName>
clades. As some of the key characters for higher taxonomic assignment are not visible due to deformation, we leave this genus unplaced to a higher taxon for now, and hope that better preserved specimens materialize to reveal more phylogenetically significant characters.
</paragraph>
<paragraph id="8B9A36A8171981674F29F9E6FEC47C56" blockId="17.[113,762,1606,1880]" pageId="17">
<emphasis id="B951EABA171981674F29F9E6FF777D53" box="[113,215,1606,1629]" italics="true" pageId="17">Diagnosis:</emphasis>
Prolonged, parallel-sided body shape with several elytral dorsal costae. Dorsal cuticle setose. Labrum long and multisetose. Head without a frontoclypeal suture, frontal stria complete. Antennal club fused, without annuli. Basal pronotal margin widely rounded. Wide and flat profemur. Protibia strongly dilated with denticles. Mesotibia narrow and spinose, metatibia narrow and almost smooth. Ŋe species is unlike any known recent or extinct
<taxonomicName id="4C254D2B171981674E26F881FE497C37" box="[382,489,1825,1849]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="17" phylum="Arthropoda" rank="family">Histeridae</taxonomicName>
</subSubSection>
<subSubSection id="C33F6523171981674F29F9E6FEC47C56" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="8B9A36A8171981674F29F9E6FEC47C56" blockId="17.[113,762,1606,1880]" pageId="17" pageNumber="18">
<emphasis id="B951EABA171981674F29F9E6FF777D53" box="[113,215,1606,1629]" italics="true" pageId="17" pageNumber="18">Diagnosis:</emphasis>
Prolonged, parallel-sided body shape with several elytral dorsal costae. Dorsal cuticle setose. Labrum long and multisetose. Head without a frontoclypeal suture, frontal stria complete. Antennal club fused, without annuli. Basal pronotal margin widely rounded. Wide and flat profemur. Protibia strongly dilated with denticles. Mesotibia narrow and spinose, metatibia narrow and almost smooth. The species is unlike any known recent or extinct
<taxonomicName id="4C254D2B171981674E26F881FE497C37" box="[382,489,1825,1849]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="family">Histeridae</taxonomicName>
species and is easily, unequivocally identifiable.
</paragraph>
<paragraph id="8B9A36A8171981674F29F8DAFE227CBF" blockId="17.[113,762,1914,1969]" pageId="17">
<emphasis id="B951EABA171981674F29F8DAFF417C9C" box="[113,225,1914,1938]" italics="true" pageId="17">Etymology:</emphasis>
Ŋe species name ‘tricostatus’ refers to the three dorsal costae on the elytra.
</subSubSection>
<subSubSection id="C33F6523171981674F29F8DAFE227CBF" pageId="17" pageNumber="18" type="etymology">
<paragraph id="8B9A36A8171981674F29F8DAFE227CBF" blockId="17.[113,762,1914,1969]" pageId="17" pageNumber="18">
<emphasis id="B951EABA171981674F29F8DAFF417C9C" box="[113,225,1914,1938]" italics="true" pageId="17" pageNumber="18">Etymology:</emphasis>
The species name ‘tricostatus’ refers to the three dorsal costae on the elytra.
</paragraph>
</subSubSection>
</treatment>

164
data/03/8C/87/038C87BE170781794E19F8C0FA0F7AB3.xml
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@ -1,116 +1,116 @@
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<emphasis id="B951EABA170781794E19F8C0FE1D7C75" bold="true" box="[321,445,1888,1915]" italics="true" pageId="15">Prodigister</emphasis>
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<emphasis id="B951EABA170781794E19F8C0FE1D7C75" bold="true" box="[321,445,1888,1915]" italics="true" pageId="15" pageNumber="16">Prodigister</emphasis>
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<taxonomicNameLabel id="A26257C1170781794E9BF8C0FD8A7C74" box="[451,554,1888,1914]" pageId="15" pageNumber="16" rank="genus">gen. nov.</taxonomicNameLabel>
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species.
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..
<emphasis id="B951EABA170781794C72FF31FC307BA6" box="[810,912,145,168]" italics="true" pageId="15">Diagnosis:</emphasis>
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<emphasis id="B951EABA170781794C72FF31FC307BA6" box="[810,912,145,168]" italics="true" pageId="15" pageNumber="16">Diagnosis:</emphasis>
Prolonged, parallel-sided body shape with several elytral dorsal costae. Labrum long and multisetose. Head without a frontoclypeal suture, frontal stria complete. Antennal club fused, without annuli. Basal pronotal margin widely rounded. Wide and flat profemur. Protibia strongly dilated with denticles. Mesotibia narrow and spinose, metatibia narrow and almost smooth.
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<emphasis id="B951EABA170781794C72FE25FC397A93" box="[810,921,389,413]" italics="true" pageId="15">Etymology:</emphasis>
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<emphasis id="B951EABA170781794C72FE25FC397A93" box="[810,921,389,413]" italics="true" pageId="15" pageNumber="16">Etymology:</emphasis>
Generic name, masculine, is derived from the Latin adjective ‘prodigiosus’ meaning bizarre, remarkable or amazing.
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243
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<mods:title id="3E4F331F40FD7B4392FC76718753D83E">A bonanza of Cretaceous fossils provides insights into the evolution of antennal protection in clown beetles (Coleoptera: Histeridae)</mods:title>
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(
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<figureCitation id="131E2A2D170781794E21FBB7FE527F3F" box="[377,498,1047,1073]" captionStart="Figure 10" captionStartId="14.[129,194,1835,1859]" captionTargetBox="[141,1461,143,1807]" captionTargetId="figure-60@14.[141,1461,143,1807]" captionTargetPageId="14" captionText="Figure 10. Phasmister planatus sp. nov. (A–C) and Phasmister kraliceki sp. nov.(D, E)." pageId="15" pageNumber="16">Fig. 10A–C</figureCitation>
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<emphasis id="B951EABA170781794F29FBF8FF6D7F7E" box="[113,205,1112,1136]" italics="true" pageId="15" pageNumber="16">Material:</emphasis>
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<collectingRegion id="49E1F84A170781794F8DFBF8FEBF7F7E" box="[213,287,1112,1136]" country="Myanmar" name="Kachin" pageId="15" pageNumber="16">Kachin</collectingRegion>
amber (
<emphasis id="B951EABA170781794E20FBF9FE217F7E" box="[376,385,1113,1136]" italics="true" pageId="15">c</emphasis>
<emphasis id="B951EABA170781794E20FBF9FE217F7E" box="[376,385,1113,1136]" italics="true" pageId="15" pageNumber="16">c</emphasis>
. 99 Mya). Amber piece dimensions: 9 × 5 ×
<quantity id="4CDD9B4D170781794F9EFBD7FEB37F81" box="[198,275,1143,1167]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="15" unit="mm" value="1.5">1.5 mm</quantity>
. Amber clear with several insect fragments and air bubbles. Deposited at HUM (ºEHU-0000121216).
</paragraph>
<paragraph id="8B9A36A8170781794F29FB71FD707E4B" blockId="15.[113,763,1232,1350]" pageId="15">
<emphasis id="B951EABA170781794F29FB71FF487FE6" box="[113,232,1233,1256]" italics="true" pageId="15">Dimensions.</emphasis>
Head width:
<quantity id="4CDD9B4D170781794ED3FB70FE467FE9" box="[395,486,1232,1256]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="15" unit="mm" value="0.48">0.48 mm</quantity>
; width between anterior pronotal angles:
<quantity id="4CDD9B4D170781794E45FB4FFECB7E09" box="[285,363,1263,1287]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="15" unit="mm" value="0.6">0.6 mm</quantity>
; width between posterior pronotal angles:
<quantity id="4CDD9B4D170781794FF1FAAFFF577E28" box="[169,247,1295,1319]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="15" unit="mm" value="1.1">1.1 mm</quantity>
; pronotal length:
<quantity id="4CDD9B4D170781794EE2FAAFFDA87E28" box="[442,520,1295,1319]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="15" unit="mm" value="0.6">0.6 mm</quantity>
; sutural elytral length:
<quantity id="4CDD9B4D170781794F29FA8EFF107E4B" box="[113,176,1326,1350]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="15" unit="mm" value="1.0">1 mm</quantity>
; elytral width (in pair, across widest point):
<quantity id="4CDD9B4D170781794D37FA8EFD6D7E4B" box="[623,717,1326,1350]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="15" unit="mm" value="1.25">1.25 mm</quantity>
<quantity id="4CDD9B4D170781794F9EFBD7FEB37F81" box="[198,275,1143,1167]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="15" pageNumber="16" unit="mm" value="1.5">1.5 mm</quantity>
. Amber clear with several insect fragments and air bubbles. Deposited at
<collectionCode id="ED34AE6D170781794E2DFB37FE147FA0" box="[373,436,1175,1198]" pageId="15" pageNumber="16">HUM</collectionCode>
(
<specimenCode id="DB839ED3170781794E98FB37FD377FA1" box="[448,663,1175,1199]" pageId="15" pageNumber="16">SEHU-0000121216</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B9A36A8170781794F29FAC8FD537DB7" blockId="15.[113,763,1383,1721]" pageId="15">
<emphasis id="B951EABA170781794F29FAC8FEEF7E71" box="[113,335,1383,1407]" italics="true" pageId="15">Diagnostic description.</emphasis>
Ŋe state of pronotal carinae is almost identical to
<taxonomicName id="4C254D2B170781794FB3FA27FD587E91" authority="Caterino and Yamamoto, 2023" authorityName="Caterino and Yamamoto" authorityYear="2023" box="[235,760,1415,1439]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="B951EABA170781794FB3FA27FE157E91" box="[235,437,1415,1439]" italics="true" pageId="15">Phasmister parallelus</emphasis>
</subSubSection>
<subSubSection id="C33F6523170781794F29FB71FD707E4B" pageId="15" pageNumber="16" type="description">
<paragraph id="8B9A36A8170781794F29FB71FD707E4B" blockId="15.[113,763,1232,1350]" pageId="15" pageNumber="16">
<emphasis id="B951EABA170781794F29FB71FF487FE6" box="[113,232,1233,1256]" italics="true" pageId="15" pageNumber="16">Dimensions.</emphasis>
Head width:
<quantity id="4CDD9B4D170781794ED3FB70FE467FE9" box="[395,486,1232,1256]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="15" pageNumber="16" unit="mm" value="0.48">0.48 mm</quantity>
; width between anterior pronotal angles:
<quantity id="4CDD9B4D170781794E45FB4FFECB7E09" box="[285,363,1263,1287]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="15" pageNumber="16" unit="mm" value="0.6">0.6 mm</quantity>
; width between posterior pronotal angles:
<quantity id="4CDD9B4D170781794FF1FAAFFF577E28" box="[169,247,1295,1319]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="15" pageNumber="16" unit="mm" value="1.1">1.1 mm</quantity>
; pronotal length:
<quantity id="4CDD9B4D170781794EE2FAAFFDA87E28" box="[442,520,1295,1319]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="15" pageNumber="16" unit="mm" value="0.6">0.6 mm</quantity>
; sutural elytral length:
<quantity id="4CDD9B4D170781794F29FA8EFF107E4B" box="[113,176,1326,1350]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="16" unit="mm" value="1.0">1 mm</quantity>
; elytral width (in pair, across widest point):
<quantity id="4CDD9B4D170781794D37FA8EFD6D7E4B" box="[623,717,1326,1350]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="15" pageNumber="16" unit="mm" value="1.25">1.25 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F6523170781794F29FAC8FD537DB7" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="8B9A36A8170781794F29FAC8FD537DB7" blockId="15.[113,763,1383,1721]" pageId="15" pageNumber="16">
<emphasis id="B951EABA170781794F29FAC8FEEF7E71" box="[113,335,1383,1407]" italics="true" pageId="15" pageNumber="16">Diagnostic description.</emphasis>
The state of pronotal carinae is almost identical to
<taxonomicName id="4C254D2B170781794FB3FA27FD587E91" authority="Caterino and Yamamoto, 2023" authorityName="Caterino and Yamamoto" authorityYear="2023" box="[235,760,1415,1439]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="B951EABA170781794FB3FA27FE157E91" box="[235,437,1415,1439]" italics="true" pageId="15" pageNumber="16">Phasmister parallelus</emphasis>
Caterino and Yamamoto, 2023
</taxonomicName>
. However,
<taxonomicName id="4C254D2B170781794F83FA07FEE77EB0" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[219,327,1446,1470]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" phylum="Arthropoda" rank="species" species="planatus">
<emphasis id="B951EABA170781794F83FA07FEE77EB0" box="[219,327,1446,1470]" italics="true" pageId="15">P. planatus</emphasis>
<taxonomicName id="4C254D2B170781794F83FA07FEE77EB0" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[219,327,1446,1470]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="planatus">
<emphasis id="B951EABA170781794F83FA07FEE77EB0" box="[219,327,1446,1470]" italics="true" pageId="15" pageNumber="16">P. planatus</emphasis>
</taxonomicName>
bears a significantly more flaưened body shape and several differential characteristics are linked with that— elytra are straight and flat (whereas in
<taxonomicName id="4C254D2B170781794EB3FA45FDFB7EF3" authorityName="Caterino and Yamamoto" authorityYear="2023" box="[491,603,1509,1533]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="B951EABA170781794EB3FA45FDFB7EF3" box="[491,603,1509,1533]" italics="true" pageId="15">P. parallelus</emphasis>
bears a significantly more flattened body shape and several differential characteristics are linked with that— elytra are straight and flat (whereas in
<taxonomicName id="4C254D2B170781794EB3FA45FDFB7EF3" authorityName="Caterino and Yamamoto" authorityYear="2023" box="[491,603,1509,1533]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="B951EABA170781794EB3FA45FDFB7EF3" box="[491,603,1509,1533]" italics="true" pageId="15" pageNumber="16">P. parallelus</emphasis>
</taxonomicName>
they are convex and curved downwards apically), spaces between first dorsal carina and upper epipleural carina and between the upper and lower epipleural carinae are narrower than in
<taxonomicName id="4C254D2B170781794D7DF9E3FD347D55" authorityName="Caterino and Yamamoto" authorityYear="2023" box="[549,660,1603,1627]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="B951EABA170781794D7DF9E3FD347D55" box="[549,660,1603,1627]" italics="true" pageId="15">P.parallelus</emphasis>
<taxonomicName id="4C254D2B170781794D7DF9E3FD347D55" authorityName="Caterino and Yamamoto" authorityYear="2023" box="[549,660,1603,1627]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="B951EABA170781794D7DF9E3FD347D55" box="[549,660,1603,1627]" italics="true" pageId="15" pageNumber="16">P.parallelus</emphasis>
</taxonomicName>
. Ventrally,
<taxonomicName id="4C254D2B170781794F29F9C3FF437D74" authorityName="Caterino and Yamamoto" authorityYear="2023" box="[113,227,1634,1658]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="B951EABA170781794F29F9C3FF437D74" box="[113,227,1634,1658]" italics="true" pageId="15">P. parallelus</emphasis>
<taxonomicName id="4C254D2B170781794F29F9C3FF437D74" authorityName="Caterino and Yamamoto" authorityYear="2023" box="[113,227,1634,1658]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="B951EABA170781794F29F9C3FF437D74" box="[113,227,1634,1658]" italics="true" pageId="15" pageNumber="16">P. parallelus</emphasis>
</taxonomicName>
has a long impunctate channel along the metaventral midline. In
<taxonomicName id="4C254D2B170781794FBDF922FEED7D97" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[229,333,1665,1689]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" phylum="Arthropoda" rank="species" species="planatus">
<emphasis id="B951EABA170781794FBDF922FEED7D97" box="[229,333,1665,1689]" italics="true" pageId="15">P. planatus</emphasis>
<taxonomicName id="4C254D2B170781794FBDF922FEED7D97" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[229,333,1665,1689]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="planatus">
<emphasis id="B951EABA170781794FBDF922FEED7D97" box="[229,333,1665,1689]" italics="true" pageId="15" pageNumber="16">P. planatus</emphasis>
</taxonomicName>
the entire metaventral disc is covered with punctures and displays a finely carinate median metaventral stria.
</paragraph>
<paragraph id="8B9A36A8170781794F29F97AFF4B7C1F" blockId="15.[113,762,1754,1809]" pageId="15">
<emphasis id="B951EABA170781794F29F97AFF417DFC" box="[113,225,1754,1778]" italics="true" pageId="15">Etymology:</emphasis>
Ŋe species name was chosen to signify its flaưened body shape.
</subSubSection>
<subSubSection id="C33F6523170781794F29F97AFF4B7C1F" pageId="15" pageNumber="16" type="etymology">
<paragraph id="8B9A36A8170781794F29F97AFF4B7C1F" blockId="15.[113,762,1754,1809]" pageId="15" pageNumber="16">
<emphasis id="B951EABA170781794F29F97AFF417DFC" box="[113,225,1754,1778]" italics="true" pageId="15" pageNumber="16">Etymology:</emphasis>
The species name was chosen to signify its flattened body shape.
</paragraph>
</subSubSection>
</treatment>

221
data/03/8C/87/038C87BE170781794E4EFEB6FF5A78CD.xml
View file

@ -1,147 +1,156 @@
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<mods:role id="97BB0C7E6B2934BD06C18F8CC6A09805">
<mods:roleTerm id="0DA1BA4D428746F550FA9214E0F34AF3">Author</mods:roleTerm>
<mods:name id="5A42189D66AFF42BC6DA4EA65343E264" type="personal">
<mods:role id="A72A97A2652B53165EA102F736EAED57">
<mods:roleTerm id="84F22785A4CC997BCDCFC39A36A42CDA">Author</mods:roleTerm>
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<mods:namePart id="CD2C19F7FC52B6862F40EF13871FAAC3">Jan</mods:namePart>
<mods:namePart id="C8F188954FFE5ECD9745EB25D0A9DE49">Simon-Pražák, Jan</mods:namePart>
<mods:affiliation id="849243D1B1A23BA4A8C81B8E340B38A5">Charles University, Faculty of Science, Department of Zoology, Viničná 7, 12800 Prague 2, Czech Republic &amp; Museum of Eastern Bohemia in Hradec Králové, Eliščino nábřeží 465, 500 03 Hradec Králové 3, Czech Republic</mods:affiliation>
<mods:nameIdentifier id="5DB74C703693431E8DB57ACCAC558948" type="email">j.prazak@muzeumhk.cz</mods:nameIdentifier>
</mods:name>
<mods:name id="792EFC283251EA909E59C4EB1FCC99F9" type="personal">
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<mods:name id="018D1E185482741DC08CD6402ECCF051" type="personal">
<mods:role id="F0B690C99B5AEB6F17622C9F20C0BD7C">
<mods:roleTerm id="0FFBF2D5DCC3E939F799A92D38840B6C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="46B8662F966802A0BD4EC23E84E6065D">ºimon-Pražák</mods:namePart>
<mods:affiliation id="C914140C744628F4BD97773388E318ED">Charles University, Faculty of ºcience, Department of Zoology, Viničná 7, 12800 Prague 2, Czech Republic &amp; Museum of Eastern Bohemia in Hradec Králové, Eliščino nábřeží 465, 500 03 Hradec Králové 3, Czech Republic</mods:affiliation>
<mods:nameIdentifier id="106B79E4EEA9F4629CB3A3EF33BBFA81" type="email">j.prazak@muzeumhk.cz</mods:nameIdentifier>
<mods:namePart id="DD06F8588369ED47755FCE922CF63C96">Yamamoto, Shûhei</mods:namePart>
<mods:affiliation id="E2C1B713EA2A471AC3D06A93C1B46A56">The Hokkaido University Museum, Hokkaido University, Kita 10, Nishi 8, Kita-ku, Sapporo 060 - 0810, Japan</mods:affiliation>
</mods:name>
<mods:name id="49B87CCB205D9E05C67032265A87A67B" type="personal">
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<mods:role id="CA3B044B5671AD0947E1FAE259B757B2">
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<mods:namePart id="3A17D327C69388B7DB1F19597B3237A1">Yamamoto</mods:namePart>
<mods:affiliation id="733A6EC7E39184CC431AC22505B4F23B">Ŋe Hokkaido University Museum, Hokkaido University, Kita 10, Nishi 8, Kita-ku, ºapporo 060 - 0810, Japan</mods:affiliation>
<mods:namePart id="6761F876D0F24C10E580B1A69F1E3C64">Lackner, Tomáš</mods:namePart>
<mods:affiliation id="42F6460169E20967185B52D5E5B6D0C7">Department of Environmental Systems Science, ETH Zürich, Weinbergstrasse 56, 8092 Zürich, Switzerland</mods:affiliation>
</mods:name>
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<mods:namePart id="E4CF665AA27DCDBFF14D7878AFF4E0D6">Lackner, Tomáš</mods:namePart>
<mods:affiliation id="490F0C1164D43F070EFB7B82C42A3576">Department of Environmental ºystems ºcience, ETH Zürich, Weinbergstrasse 56, 8092 Zürich, ºwitzerland</mods:affiliation>
<mods:namePart id="48DF774D2B746EE014508C2E2BA57111">Fikáček, Martin</mods:namePart>
<mods:affiliation id="78B47918D8D1B155517D66657C34F55F">Department of Biological Sciences, National Sun Yat-sen University, No. 70, Lienhai Rd., Kaohsiung 80424, Taiwan &amp; Department of Entomology, National Museum, Cirkusová 1740 CZ- 19300 Praha 9, Czech Republic</mods:affiliation>
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<mods:namePart id="8CEE128ACE6354F49C0E65ABD29E6AAA">Fikáček, Martin</mods:namePart>
<mods:affiliation id="446A83FF4E678BD139C4E610944ECF76">Department of Biological ºciences, National ºun Yat-sen University, No. 70, Lienhai Rd., Kaohsiung 80424, Taiwan &amp; Department of Entomology, National Museum, Cirkusová 1740 CZ- 19300 Praha 9, Czech Republic</mods:affiliation>
<mods:namePart id="883C874DD8663D5FCF0AAED243FBE628">Prokop, Jakub</mods:namePart>
<mods:affiliation id="81A1E1435075008AE0A9DB529C1E7852">Charles University, Faculty of Science, Department of Zoology, Viničná 7, 12800 Prague 2, Czech Republic &amp; Department of Plant and Environmental Sciences, Clemson University, Clemson, SC 29634, United States</mods:affiliation>
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<mods:namePart id="DDABC0F795A8BB6A3838F6F416925FE1">Prokop, Jakub</mods:namePart>
<mods:affiliation id="09EA0A984C44BAD4D22331C3046CC3CB">Charles University, Faculty of ºcience, Department of Zoology, Viničná 7, 12800 Prague 2, Czech Republic</mods:affiliation>
<mods:namePart id="C80A2FE811DCDDE49D861789367B3CA2">Caterino, Michael S</mods:namePart>
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<mods:namePart id="52FB01C2311C6F5AAC3E7D548659F3F5">Caterino</mods:namePart>
<mods:affiliation id="A966327FC6176484A7C84010EE9408EB">Department of Plant and Environmental ºciences, Clemson University, Clemson, ºC 29634, United ºtates</mods:affiliation>
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<mods:title id="7B0F1D826E93B6238A52782AC03B6DF0">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="36AD94A1F4A71FAF0CDD626A89981F47">
<mods:date id="613B28874984B992AFBB8B99EADF9BEB">2024</mods:date>
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<mods:title id="9D6A94EAC19BF43CE7E5686120DFD904">zlae 137</mods:title>
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<treatment id="038C87BE170781794E4EFEB6FF5A78CD" ID-DOI="http://doi.org/10.5281/zenodo.14424565" ID-Zenodo-Dep="14424565" LSID="urn:lsid:plazi:treatment:038C87BE170781794E4EFEB6FF5A78CD" httpUri="http://treatment.plazi.org/id/038C87BE170781794E4EFEB6FF5A78CD" pageId="15">
<subSubSection id="C33F6523170781794E4EFEB6FDF67A3F" box="[278,598,278,305]" pageId="15" type="nomenclature">
<paragraph id="8B9A36A8170781794E4EFEB6FDF67A3F" blockId="15.[278,598,278,305]" box="[278,598,278,305]" pageId="15">
<heading id="D0D281C4170781794E4EFEB6FDF67A3F" bold="true" box="[278,598,278,305]" centered="true" fontSize="11" level="2" pageId="15" reason="2">
<emphasis id="B951EABA170781794E4EFEB6FDF67A3F" bold="true" box="[278,598,278,305]" pageId="15">
<taxonomicName id="4C254D2B170781794E4EFEB6FE5A7A3F" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[278,506,278,305]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" phylum="Arthropoda" rank="species" species="kraliceki" status="sp. nov.">
<emphasis id="B951EABA170781794E4EFEB6FE5A7A3F" bold="true" box="[278,506,278,305]" italics="true" pageId="15">Phasmister kraliceki</emphasis>
<treatment id="038C87BE170781794E4EFEB6FF5A78CD" ID-DOI="http://doi.org/10.5281/zenodo.14424565" ID-Zenodo-Dep="14424565" LSID="urn:lsid:plazi:treatment:038C87BE170781794E4EFEB6FF5A78CD" httpUri="http://treatment.plazi.org/id/038C87BE170781794E4EFEB6FF5A78CD" lastPageNumber="16" pageId="15" pageNumber="16">
<subSubSection id="C33F6523170781794E4EFEB6FDF67A3F" box="[278,598,278,305]" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="8B9A36A8170781794E4EFEB6FDF67A3F" blockId="15.[278,598,278,305]" box="[278,598,278,305]" pageId="15" pageNumber="16">
<heading id="D0D281C4170781794E4EFEB6FDF67A3F" bold="true" box="[278,598,278,305]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="16" reason="2">
<emphasis id="B951EABA170781794E4EFEB6FDF67A3F" bold="true" box="[278,598,278,305]" pageId="15" pageNumber="16">
<taxonomicName id="4C254D2B170781794E4EFEB6FE5A7A3F" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[278,506,278,305]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="kraliceki" status="sp. nov.">
<emphasis id="B951EABA170781794E4EFEB6FE5A7A3F" bold="true" box="[278,506,278,305]" italics="true" pageId="15" pageNumber="16">Phasmister kraliceki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C1170781794EA7FEB7FDF67A3F" box="[511,598,279,305]" pageId="15" rank="species">sp. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="A26257C1170781794EA7FEB7FDF67A3F" box="[511,598,279,305]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C33F6523170781794E2AFEE6FF5A78CD" pageId="15" type="description">
<paragraph id="8B9A36A8170781794E2AFEE6FE5A7A6E" blockId="15.[370,506,326,352]" box="[370,506,326,352]" pageId="15">
<emphasis id="B951EABA170781794E2AFEE6FE5A7A6E" bold="true" box="[370,506,326,352]" pageId="15">
<subSubSection id="C33F6523170781794E2AFEE6FE5A7A6E" box="[370,506,326,352]" pageId="15" pageNumber="16" type="description">
<paragraph id="8B9A36A8170781794E2AFEE6FE5A7A6E" blockId="15.[370,506,326,352]" box="[370,506,326,352]" pageId="15" pageNumber="16">
<emphasis id="B951EABA170781794E2AFEE6FE5A7A6E" bold="true" box="[370,506,326,352]" pageId="15" pageNumber="16">
(
<figureCitation id="131E2A2D170781794E25FEE6FE4F7A6E" box="[381,495,326,352]" captionStart="Figure 10" captionStartId="14.[129,194,1835,1859]" captionTargetBox="[141,1461,143,1807]" captionTargetId="figure-60@14.[141,1461,143,1807]" captionTargetPageId="14" captionText="Figure 10. Phasmister planatus sp. nov. (A–C) and Phasmister kraliceki sp. nov.(D, E)." pageId="15">Fig. 10D, E</figureCitation>
<figureCitation id="131E2A2D170781794E25FEE6FE4F7A6E" box="[381,495,326,352]" captionStart="Figure 10" captionStartId="14.[129,194,1835,1859]" captionTargetBox="[141,1461,143,1807]" captionTargetId="figure-60@14.[141,1461,143,1807]" captionTargetPageId="14" captionText="Figure 10. Phasmister planatus sp. nov. (A–C) and Phasmister kraliceki sp. nov.(D, E)." pageId="15" pageNumber="16">Fig. 10D, E</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8B9A36A8170781794F29FE27FDEE7AD3" blockId="15.[113,763,391,477]" pageId="15">
<emphasis id="B951EABA170781794F29FE27FF6D7A91" box="[113,205,391,415]" italics="true" pageId="15">Material:</emphasis>
<collectingRegion id="49E1F84A170781794F8DFE27FEBF7A91" box="[213,287,391,415]" country="Myanmar" name="Kachin" pageId="15">Kachin</collectingRegion>
</subSubSection>
<subSubSection id="C33F6523170781794F29FE27FDEE7AD3" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="8B9A36A8170781794F29FE27FDEE7AD3" blockId="15.[113,763,391,477]" pageId="15" pageNumber="16">
<emphasis id="B951EABA170781794F29FE27FF6D7A91" box="[113,205,391,415]" italics="true" pageId="15" pageNumber="16">Material:</emphasis>
<materialsCitation id="3B4D3CF5170781794F8DFE27FDEA7AD3" collectionCode="MEBHK" location="Kachin" pageId="15" pageNumber="16" specimenCode="P1424/E" specimenCount="1" stateProvince="Kachin">
<collectingRegion id="49E1F84A170781794F8DFE27FEBF7A91" box="[213,287,391,415]" country="Myanmar" name="Kachin" pageId="15" pageNumber="16">Kachin</collectingRegion>
amber (
<emphasis id="B951EABA170781794E20FE27FE217A90" box="[376,385,391,414]" italics="true" pageId="15">c</emphasis>
<emphasis id="B951EABA170781794E20FE27FE217A90" box="[376,385,391,414]" italics="true" pageId="15" pageNumber="16">c</emphasis>
. 99 Mya). Amber piece dimensions: 11 × 9 ×
<quantity id="4CDD9B4D170781794F89FE06FEBF7AB3" box="[209,287,422,446]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="15" unit="mm" value="1.5">1.5 mm</quantity>
. Amber clear with several insect fragments and air bubbles. Deposited at MEBHK (P1424/E).
</paragraph>
<paragraph id="8B9A36A8170781794F29FE5FFD43797A" blockId="15.[113,763,511,629]" pageId="15">
<emphasis id="B951EABA170781794F29FE5FFF487918" box="[113,232,511,534]" italics="true" pageId="15">Dimensions.</emphasis>
Head width:
<quantity id="4CDD9B4D170781794ED3FE5FFE467918" box="[395,486,511,535]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="15" unit="mm" value="0.42">0.42 mm</quantity>
; width between anterior pronotal angles:
<quantity id="4CDD9B4D170781794E7DFDBEFE20793B" box="[293,384,542,566]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="15" unit="mm" value="0.55">0.55 mm</quantity>
; width between posterior pronotal angles:
<quantity id="4CDD9B4D170781794FE6FD9DFEAC795B" box="[190,268,573,597]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="15" unit="mm" value="0.9">0.9 mm</quantity>
; pronotal length:
<quantity id="4CDD9B4D170781794E9DFD9DFDB2795B" box="[453,530,573,597]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="15" unit="mm" value="0.5">0.5 mm</quantity>
; sutural elytral length:
<quantity id="4CDD9B4D170781794F2AFDFDFF62797A" box="[114,194,605,629]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="15" unit="mm" value="0.8">0.8 mm</quantity>
; elytral width (in pair, across widest point):
<quantity id="4CDD9B4D170781794DD9FDFCFD7F797A" box="[641,735,604,628]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="15" unit="mm" value="1.15">1.15 mm</quantity>
<quantity id="4CDD9B4D170781794F89FE06FEBF7AB3" box="[209,287,422,446]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="15" pageNumber="16" unit="mm" value="1.5">1.5 mm</quantity>
. Amber clear with several insect fragments and air bubbles. Deposited at
<collectionCode id="ED34AE6D170781794E2BFE66FE6F7AD3" box="[371,463,454,477]" pageId="15" pageNumber="16">MEBHK</collectionCode>
(
<specimenCode id="DB839ED3170781794E84FE65FDE27AD3" box="[476,578,453,477]" pageId="15" pageNumber="16">P1424/E</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B9A36A8170781794F29FD37FEC17864" blockId="15.[113,763,662,874]" pageId="15">
<emphasis id="B951EABA170781794F29FD37FEF579A0" box="[113,341,662,686]" italics="true" pageId="15">Diagnostic description.</emphasis>
Ŋe species is very similar to the recently described
<taxonomicName id="4C254D2B170781794E47FD15FF0779E3" authority="Caterino and Yamamoto, 2023" authorityName="Caterino and Yamamoto" authorityYear="2023" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" phylum="Arthropoda" rank="species" species="hkamticus">
<emphasis id="B951EABA170781794E47FD15FE5479C3" box="[287,500,693,717]" italics="true" pageId="15">Phasmister hkamticus</emphasis>
</subSubSection>
<subSubSection id="C33F6523170781794F29FE5FFD43797A" pageId="15" pageNumber="16" type="description">
<paragraph id="8B9A36A8170781794F29FE5FFD43797A" blockId="15.[113,763,511,629]" pageId="15" pageNumber="16">
<emphasis id="B951EABA170781794F29FE5FFF487918" box="[113,232,511,534]" italics="true" pageId="15" pageNumber="16">Dimensions.</emphasis>
Head width:
<quantity id="4CDD9B4D170781794ED3FE5FFE467918" box="[395,486,511,535]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="15" pageNumber="16" unit="mm" value="0.42">0.42 mm</quantity>
; width between anterior pronotal angles:
<quantity id="4CDD9B4D170781794E7DFDBEFE20793B" box="[293,384,542,566]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="15" pageNumber="16" unit="mm" value="0.55">0.55 mm</quantity>
; width between posterior pronotal angles:
<quantity id="4CDD9B4D170781794FE6FD9DFEAC795B" box="[190,268,573,597]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="15" pageNumber="16" unit="mm" value="0.9">0.9 mm</quantity>
; pronotal length:
<quantity id="4CDD9B4D170781794E9DFD9DFDB2795B" box="[453,530,573,597]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="16" unit="mm" value="0.5">0.5 mm</quantity>
; sutural elytral length:
<quantity id="4CDD9B4D170781794F2AFDFDFF62797A" box="[114,194,605,629]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="15" pageNumber="16" unit="mm" value="0.8">0.8 mm</quantity>
; elytral width (in pair, across widest point):
<quantity id="4CDD9B4D170781794DD9FDFCFD7F797A" box="[641,735,604,628]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="15" pageNumber="16" unit="mm" value="1.15">1.15 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F6523170781794F29FD37FEC17864" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="8B9A36A8170781794F29FD37FEC17864" blockId="15.[113,763,662,874]" pageId="15" pageNumber="16">
<emphasis id="B951EABA170781794F29FD37FEF579A0" box="[113,341,662,686]" italics="true" pageId="15" pageNumber="16">Diagnostic description.</emphasis>
The species is very similar to the recently described
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Caterino and Yamamoto, 2023
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. However, it differs in several characters by which it can be distinguished. Pronotal disc lacks a short, curved costa in posterolateral corners; pronotal base more sinuate; anterior pronotal margin more broadly rounded, not crenulate and more emarginate above head.
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<emphasis id="B951EABA170781794F29FC2BFF4178AD" box="[113,225,907,931]" italics="true" pageId="15">Etymology:</emphasis>
Named asser Ivo Králíček, a good friend and biology tutor of J.º.P..
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<emphasis id="B951EABA170781794F29FC2BFF4178AD" box="[113,225,907,931]" italics="true" pageId="15" pageNumber="16">Etymology:</emphasis>
Named after Ivo Králíček, a good friend and biology tutor of J.S.P..
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327
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<emphasis id="B951EABA170D81734CEAFCFAFB6F787B" bold="true" box="[946,1231,858,885]" italics="true" pageId="5" pageNumber="6">Cretomalus tibiodentatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C1170D81734B8CFCFBFA8B787B" box="[1236,1323,859,885]" pageId="5" rank="species">sp. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="A26257C1170D81734B8CFCFBFA8B787B" box="[1236,1323,859,885]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33F6523170D81714B1FFC2CFF7B7E6D" lastPageId="7" pageId="5" type="description">
<paragraph id="8B9A36A8170D81734B1FFC2CFB3678A8" blockId="5.[1095,1174,908,935]" box="[1095,1174,908,935]" pageId="5">
<emphasis id="B951EABA170D81734B1FFC2CFB3678A8" bold="true" box="[1095,1174,908,935]" pageId="5">
<subSubSection id="C33F6523170D81734B1FFC2CFB3678A8" box="[1095,1174,908,935]" pageId="5" pageNumber="6" type="description">
<paragraph id="8B9A36A8170D81734B1FFC2CFB3678A8" blockId="5.[1095,1174,908,935]" box="[1095,1174,908,935]" pageId="5" pageNumber="6">
<emphasis id="B951EABA170D81734B1FFC2CFB3678A8" bold="true" box="[1095,1174,908,935]" pageId="5" pageNumber="6">
(
<figureCitation id="131E2A2D170D81734B0AFC2CFB2B78A9" box="[1106,1163,908,935]" captionStart="Figure 5" captionStartId="6.[129,194,1370,1394]" captionTargetBox="[129,1473,144,1342]" captionTargetId="figure-262@6.[129,1473,144,1342]" captionTargetPageId="6" captionText="Figure 5. Cretomalus tibiodentatus gen. et sp. nov. A, dorsal view.B, ventral view; C, postmetacoxal area. D, postmesocoxal area. E, front leg and antenna. Abbreviations: fs, frontal stria; las, lateral stria on first abdominal ventrite; lms, lateral metaventral stria; pl, prosternal lobe; pms, prosternomesoventral stria; pmss, postmesocoxal stria; pmts, postmetacoxal stria." pageId="5">Fig. 5</figureCitation>
<figureCitation id="131E2A2D170D81734B0AFC2CFB2B78A9" box="[1106,1163,908,935]" captionStart="Figure 5" captionStartId="6.[129,194,1370,1394]" captionTargetBox="[129,1473,144,1342]" captionTargetId="figure-262@6.[129,1473,144,1342]" captionTargetPageId="6" captionText="Figure 5. Cretomalus tibiodentatus gen. et sp. nov. A, dorsal view.B, ventral view; C, postmetacoxal area. D, postmesocoxal area. E, front leg and antenna. Abbreviations: fs, frontal stria; las, lateral stria on first abdominal ventrite; lms, lateral metaventral stria; pl, prosternal lobe; pms, prosternomesoventral stria; pmss, postmesocoxal stria; pmts, postmetacoxal stria." pageId="5" pageNumber="6">Fig. 5</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8B9A36A8170D81734C72FC68FAF97F11" blockId="5.[810,1460,968,1055]" pageId="5">
<emphasis id="B951EABA170D81734C72FC68FC2678EE" box="[810,902,968,992]" italics="true" pageId="5">Material:</emphasis>
<collectingRegion id="49E1F84A170D81734CD5FC68FC7778EE" box="[909,983,968,992]" country="Myanmar" name="Kachin" pageId="5">Kachin</collectingRegion>
</subSubSection>
<subSubSection id="C33F6523170D81734C72FC68FAF97F11" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="8B9A36A8170D81734C72FC68FAF97F11" blockId="5.[810,1460,968,1055]" pageId="5" pageNumber="6">
<emphasis id="B951EABA170D81734C72FC68FC2678EE" box="[810,902,968,992]" italics="true" pageId="5" pageNumber="6">Material:</emphasis>
<materialsCitation id="3B4D3CF5170D81734CD5FC68FAF57F11" collectionCode="HUM" location="Kachin" pageId="5" pageNumber="6" specimenCode="SEHU-0000121215" specimenCount="1" stateProvince="Kachin">
<collectingRegion id="49E1F84A170D81734CD5FC68FC7778EE" box="[909,983,968,992]" country="Myanmar" name="Kachin" pageId="5" pageNumber="6">Kachin</collectingRegion>
amber (
<emphasis id="B951EABA170D81734B69FC69FB9A78EE" box="[1073,1082,969,992]" italics="true" pageId="5">c</emphasis>
<emphasis id="B951EABA170D81734B69FC69FB9A78EE" box="[1073,1082,969,992]" italics="true" pageId="5" pageNumber="6">c</emphasis>
. 99 Mya). Amber piece dimensions: 13 × 8 ×
<quantity id="4CDD9B4D170D81734CD2FC47FC7878F1" box="[906,984,999,1023]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="5" unit="mm" value="1.5">1.5 mm</quantity>
. Amber clear with several insect fragments and air bubbles. Deposited at HUM (ºEHU-0000121215).
<quantity id="4CDD9B4D170D81734CD2FC47FC7878F1" box="[906,984,999,1023]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="6" unit="mm" value="1.5">1.5 mm</quantity>
. Amber clear with several insect fragments and air bubbles. Deposited at
<collectionCode id="ED34AE6D170D81734B74FBA7FBCB7F10" box="[1068,1131,1031,1054]" pageId="5" pageNumber="6">HUM</collectionCode>
(
<specimenCode id="DB839ED3170D81734B2FFBA7FAEE7F11" box="[1143,1358,1031,1055]" pageId="5" pageNumber="6">SEHU-0000121215</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B9A36A8170D81704C72FBE1FA9B7DE3" blockId="5.[810,1461,1088,1958]" lastBlockId="6.[825,1475,1498,1773]" lastPageId="6" pageId="5">
<emphasis id="B951EABA170D81734C72FBE1FC187F56" box="[810,952,1089,1112]" italics="true" pageId="5">Measurements:</emphasis>
</subSubSection>
<subSubSection id="C33F6523170D81704C72FBE1FA9B7DE3" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="description">
<paragraph id="8B9A36A8170D81704C72FBE1FA9B7DE3" blockId="5.[810,1461,1088,1958]" lastBlockId="6.[825,1475,1498,1773]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<emphasis id="B951EABA170D81734C72FBE1FC187F56" box="[810,952,1089,1112]" italics="true" pageId="5" pageNumber="6">Measurements:</emphasis>
Head width:
<quantity id="4CDD9B4D170D81734B0BFBE0FB0A7F56" box="[1107,1194,1088,1112]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="5" unit="mm" value="0.32">0.32 mm</quantity>
<quantity id="4CDD9B4D170D81734B0BFBE0FB0A7F56" box="[1107,1194,1088,1112]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="5" pageNumber="6" unit="mm" value="0.32">0.32 mm</quantity>
; width between anterior pronotal angles:
<quantity id="4CDD9B4D170D81734C8AFBFFFB8A7F79" box="[978,1066,1119,1143]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="5" unit="mm" value="0.35">0.35 mm</quantity>
<quantity id="4CDD9B4D170D81734C8AFBFFFB8A7F79" box="[978,1066,1119,1143]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="5" pageNumber="6" unit="mm" value="0.35">0.35 mm</quantity>
; width between posterior pronotal angles:
<quantity id="4CDD9B4D170D81734C39FBDFFC197F98" box="[865,953,1151,1175]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="5" unit="mm" value="0.55">0.55 mm</quantity>
<quantity id="4CDD9B4D170D81734C39FBDFFC197F98" box="[865,953,1151,1175]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="5" pageNumber="6" unit="mm" value="0.55">0.55 mm</quantity>
; pronotal length:
<quantity id="4CDD9B4D170D81734B22FBDFFB657F98" box="[1146,1221,1151,1175]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="5" unit="mm" value="0.5">0.5 mm</quantity>
<quantity id="4CDD9B4D170D81734B22FBDFFB657F98" box="[1146,1221,1151,1175]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="6" unit="mm" value="0.5">0.5 mm</quantity>
; sutural elytral length:
<quantity id="4CDD9B4D170D81734C72FB3EFC227FB8" box="[810,898,1182,1206]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" pageId="5" unit="mm" value="0.73">0.73 mm</quantity>
<quantity id="4CDD9B4D170D81734C72FB3EFC227FB8" box="[810,898,1182,1206]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" pageId="5" pageNumber="6" unit="mm" value="0.73">0.73 mm</quantity>
; elytral width (in pair, across widest point):
<quantity id="4CDD9B4D170D81734A62FB3EFA327FB8" box="[1338,1426,1182,1206]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="5" unit="mm" value="0.53">0.53 mm</quantity>
<quantity id="4CDD9B4D170D81734A62FB3EFA327FB8" box="[1338,1426,1182,1206]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="5" pageNumber="6" unit="mm" value="0.53">0.53 mm</quantity>
.—
<emphasis id="B951EABA170D81734C72FB1DFCC07FDB" bold="true" box="[810,864,1213,1237]" italics="true" pageId="5">Body</emphasis>
elongate, rectangular, parallel-sided, and flaưened. Cuticle sparsely punctuate dorsally, smooth ventrally. Colour black to chestnut brown.—
<emphasis id="B951EABA170D81734CBFFB5CFB877E1A" bold="true" box="[999,1063,1276,1300]" italics="true" pageId="5">Head:</emphasis>
Frons weakly convex, finely punctuate, punctures separated by three times their diameter. Frontoclypeal suture not evident; frontal stria thin, complete.Labrum short, trapezoidal, with several pairs of prominent setae.Mouthparts retracted. Eyes not visible from above.ºcape short, expanded apically, pedicel simple, as long as antennomeres 3–4 combined. Antennomeres 3–8 wide and short (width: length ratio 1.5: 1), almost rectangular. Antennal club round, dorsoventrally flaưened, annuli well visible.—
<emphasis id="B951EABA170D81734CD1FA57FC7C7D01" bold="true" box="[905,988,1527,1551]" italics="true" pageId="5">Ŋorax:</emphasis>
Pronotum as wide as long (width: length ratio 1: 1), median portion of disk flat, sides curved ventrad. Anterior emarginationdeep,anterioranglesacute,convergentanteriad.Basal pronotal margin rounded. Marginal pronotal stria complete along lateral and anterior margins. Pronotum punctuate, punctures separated by two to three times their diameter.ºcutellum not visible (obscured by bubbles). Elytra long and narrow, punctuation density as in pronotum. ºutural area slightly depressed (possibly deformed postmortem), then elevated medially forming a keel. Dorsal elytral striation absent; elytral epipleuron obscured. Prosternal lobe deflexed, long, wide, rounded anteriorly. Prosternal keel with fine punctuation.Prosternalstriationindeterminableduetoairbubbles. Basal prosternal margin rounded, projecting posteriorly. Antennal cavities vaguely defined and widely open, located posterolaterally (covered by the front leg when retracted). Mesoventrite very short and wide (1: 3 ratio). Anterior margin curved posteriorly to fit the basal prosternal margin. Punctuation not apparent. Marginal mesoventral stria complete, continuous with lateral metaventral stria. Transverse mesoventral stria not visible. Metaventrite large, weakly concave, slightly elevated anterad metacoxae, without apparent punctuation. Lateral metaventral striae present and extending posterolaterad on at least anterior half. Postmesocoxal striae present and extending posteriorly. ºides of meso- and metaventrites (and abdominal ventrites) ascending laterad coxae, oblique.—
<emphasis id="B951EABA170E81704FB3F8B4FE817C25" bold="true" box="[235,289,1812,1835]" italics="true" pageId="6">Legs:</emphasis>
Profemur flaưened and broad. Protibia broad and strongly curved, withsix to seven small, symmetrical teeth along the outer margin. Protarsal groove weakly developed. ºmall inner and curved median protibial spurs present. Protarsomere 5 as long as protarsomeres 1–4 combined. Protarsal claws short. Mesofemur narrower than profemur, mesotibia smooth, spur present. Terminal mesotarsomere as long as mesotarsomeres 3–4 combined. Hindleg equivalent to middle leg.—
<emphasis id="B951EABA170E81704B04F9B8FB667D3E" bold="true" box="[1116,1222,1560,1584]" italics="true" pageId="6">Abdomen:</emphasis>
First visible abdominal sternite large, intercoxal disc almost square-shaped, with very fine, sparse punctuation (punctures separated by at least 10 times their diameter). Lateral striae strongly impressed, complete, running posterolaterad. Postmetacoxal striae present, curved along metacoxae. Other sternites with identical punctuation paưern. Pygidium strongly convex, apex subtruncate, rugose.
<emphasis id="B951EABA170D81734C72FB1DFCC07FDB" bold="true" box="[810,864,1213,1237]" italics="true" pageId="5" pageNumber="6">Body</emphasis>
elongate, rectangular, parallel-sided, and flattened. Cuticle sparsely punctuate dorsally, smooth ventrally. Colour black to chestnut brown.—
<emphasis id="B951EABA170D81734CBFFB5CFB877E1A" bold="true" box="[999,1063,1276,1300]" italics="true" pageId="5" pageNumber="6">Head:</emphasis>
Frons weakly convex, finely punctuate, punctures separated by three times their diameter. Frontoclypeal suture not evident; frontal stria thin, complete.Labrum short, trapezoidal, with several pairs of prominent setae.Mouthparts retracted. Eyes not visible from above.Scape short, expanded apically, pedicel simple, as long as antennomeres 3–4 combined. Antennomeres 3–8 wide and short (width: length ratio 1.5: 1), almost rectangular. Antennal club round, dorsoventrally flattened, annuli well visible.—
<emphasis id="B951EABA170D81734CD1FA57FC7C7D01" bold="true" box="[905,988,1527,1551]" italics="true" pageId="5" pageNumber="6">Thorax:</emphasis>
Pronotum as wide as long (width: length ratio 1: 1), median portion of disk flat, sides curved ventrad. Anterior emarginationdeep,anterioranglesacute,convergentanteriad.Basal pronotal margin rounded. Marginal pronotal stria complete along lateral and anterior margins. Pronotum punctuate, punctures separated by two to three times their diameter.Scutellum not visible (obscured by bubbles). Elytra long and narrow, punctuation density as in pronotum. Sutural area slightly depressed (possibly deformed postmortem), then elevated medially forming a keel. Dorsal elytral striation absent; elytral epipleuron obscured. Prosternal lobe deflexed, long, wide, rounded anteriorly. Prosternal keel with fine punctuation.Prosternalstriationindeterminableduetoairbubbles. Basal prosternal margin rounded, projecting posteriorly. Antennal cavities vaguely defined and widely open, located posterolaterally (covered by the front leg when retracted). Mesoventrite very short and wide (1: 3 ratio). Anterior margin curved posteriorly to fit the basal prosternal margin. Punctuation not apparent. Marginal mesoventral stria complete, continuous with lateral metaventral stria. Transverse mesoventral stria not visible. Metaventrite large, weakly concave, slightly elevated anterad metacoxae, without apparent punctuation. Lateral metaventral striae present and extending posterolaterad on at least anterior half. Postmesocoxal striae present and extending posteriorly. Sides of meso- and metaventrites (and abdominal ventrites) ascending laterad coxae, oblique.—
<emphasis id="B951EABA170E81704FB3F8B4FE817C25" bold="true" box="[235,289,1812,1835]" italics="true" pageId="6" pageNumber="7">Legs:</emphasis>
Profemur flattened and broad. Protibia broad and strongly curved, withsix to seven small, symmetrical teeth along the outer margin. Protarsal groove weakly developed. Small inner and curved median protibial spurs present. Protarsomere 5 as long as protarsomeres 1–4 combined. Protarsal claws short. Mesofemur narrower than profemur, mesotibia smooth, spur present. Terminal mesotarsomere as long as mesotarsomeres 3–4 combined. Hindleg equivalent to middle leg.—
<emphasis id="B951EABA170E81704B04F9B8FB667D3E" bold="true" box="[1116,1222,1560,1584]" italics="true" pageId="6" pageNumber="7">Abdomen:</emphasis>
First visible abdominal sternite large, intercoxal disc almost square-shaped, with very fine, sparse punctuation (punctures separated by at least 10 times their diameter). Lateral striae strongly impressed, complete, running posterolaterad. Postmetacoxal striae present, curved along metacoxae. Other sternites with identical punctuation pattern. Pygidium strongly convex, apex subtruncate, rugose.
</paragraph>
<caption id="DF5A6620170E81704FD9FAFAFC217EA4" pageId="6" startId="6.[129,194,1370,1394]" targetBox="[129,1473,144,1342]" targetPageId="6" targetType="figure">
<paragraph id="8B9A36A8170E81704FD9FAFAFC217EA4" blockId="6.[129,1453,1370,1450]" pageId="6">
<emphasis id="B951EABA170E81704FD9FAFAFF787E7C" bold="true" box="[129,216,1370,1394]" pageId="6">Figure 5.</emphasis>
<taxonomicName id="4C254D2B170E81704F85FAFAFE1E7E7C" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[221,446,1370,1394]" class="Insecta" family="Histeridae" genus="Cretomalus" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="tibiodentatus" status="gen. et sp. nov.">
<emphasis id="B951EABA170E81704F85FAFAFE1E7E7C" box="[221,446,1370,1394]" italics="true" pageId="6">Cretomalus tibiodentatus</emphasis>
</subSubSection>
<caption id="DF5A6620170E81704FD9FAFAFC217EA4" pageId="6" pageNumber="7" startId="6.[129,194,1370,1394]" targetBox="[129,1473,144,1342]" targetPageId="6" targetType="figure">
<paragraph id="8B9A36A8170E81704FD9FAFAFC217EA4" blockId="6.[129,1453,1370,1450]" pageId="6" pageNumber="7">
<emphasis id="B951EABA170E81704FD9FAFAFF787E7C" bold="true" box="[129,216,1370,1394]" pageId="6" pageNumber="7">Figure 5.</emphasis>
<taxonomicName id="4C254D2B170E81704F85FAFAFE1E7E7C" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[221,446,1370,1394]" class="Insecta" family="Histeridae" genus="Cretomalus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="tibiodentatus" status="gen. et sp. nov.">
<emphasis id="B951EABA170E81704F85FAFAFE1E7E7C" box="[221,446,1370,1394]" italics="true" pageId="6" pageNumber="7">Cretomalus tibiodentatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C1170E81704E9BFAFBFDEC7E7C" box="[451,588,1371,1395]" pageId="6" rank="species">gen. et sp. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="A26257C1170E81704E9BFAFBFDEC7E7C" box="[451,588,1371,1395]" pageId="6" pageNumber="7" rank="species">gen. et sp. nov.</taxonomicNameLabel>
A, dorsal view. B, ventral view; C, postmetacoxal area. D, postmesocoxal area. E, front leg and antenna. Abbreviations: fs, frontal stria; las, lateral stria on first abdominal ventrite; lms, lateral metaventral stria; pl, prosternal lobe; pms, prosternomesoventral stria; pmss, postmesocoxal stria; pmts, postmetacoxal stria.
</paragraph>
</caption>
<paragraph id="8B9A36A8170E81714C61F8B2FE487950" blockId="6.[825,1474,1809,1958]" lastBlockId="7.[113,763,144,607]" lastPageId="7" pageId="6">
<emphasis id="B951EABA170E81704C61F8B2FB807C27" box="[825,1056,1810,1833]" italics="true" pageId="6">Taxonomic assignment:</emphasis>
<subSubSection id="C33F6523170E81714C61F8B2FE487950" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="discussion">
<paragraph id="8B9A36A8170E81714C61F8B2FE487950" blockId="6.[825,1474,1809,1958]" lastBlockId="7.[113,763,144,607]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<emphasis id="B951EABA170E81704C61F8B2FB807C27" box="[825,1056,1810,1833]" italics="true" pageId="6" pageNumber="7">Taxonomic assignment:</emphasis>
We place this species in the
<taxonomicName id="4C254D2B170E81704A0BF8B1FA627C27" authorityName="Reitter" authorityYear="1909" box="[1363,1474,1809,1833]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="tribe" tribe="Paromalini">Paromalini</taxonomicName>
<taxonomicName id="4C254D2B170E81704A0BF8B1FA627C27" authorityName="Reitter" authorityYear="1909" box="[1363,1474,1809,1833]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="tribe" tribe="Paromalini">Paromalini</taxonomicName>
tribe based on the combination of the following characters (
<bibRefCitation id="EFB44B59170E81704C1BF8F1FC1E7C66" author="Ohara M" box="[835,958,1872,1900]" pageId="6" pagination="1 - 283" refId="ref18649" refString="Ohara M. A Revision of the ºuperfamily Histeroidea of Japan [Coleoptera]. Insecta Matsumurana 1994; 51: 1 - 283." type="journal article" year="1994">Ôhara 1994</bibRefCitation>
<bibRefCitation id="EFB44B59170E81704C1BF8F1FC1E7C66" author="Ohara M" box="[835,958,1872,1900]" pageId="6" pageNumber="7" pagination="1 - 283" refId="ref18649" refString="Ohara M. A Revision of the ºuperfamily Histeroidea of Japan [Coleoptera]. Insecta Matsumurana 1994; 51: 1 - 283." type="journal article" year="1994">Ôhara 1994</bibRefCitation>
): broad, weakly convex frons (
<figureCitation id="131E2A2D170E81704BA6F8F0FAE97C66" box="[1278,1353,1872,1896]" captionStart="Figure 5" captionStartId="6.[129,194,1370,1394]" captionTargetBox="[129,1473,144,1342]" captionTargetId="figure-262@6.[129,1473,144,1342]" captionTargetPageId="6" captionText="Figure 5. Cretomalus tibiodentatus gen. et sp. nov. A, dorsal view.B, ventral view; C, postmetacoxal area. D, postmesocoxal area. E, front leg and antenna. Abbreviations: fs, frontal stria; las, lateral stria on first abdominal ventrite; lms, lateral metaventral stria; pl, prosternal lobe; pms, prosternomesoventral stria; pmss, postmesocoxal stria; pmts, postmetacoxal stria." pageId="6">Fig. 5A</figureCitation>
), short and wide labrum with several pairs of prominent setae, short and wide prosternal lobe, vaguely defined antennal cavities located at prosternal hypomeron, basal prosternal margin rounded and projecting posteriorly, wide, curving protibia with a laterally directed median apical spur. Ŋe position of this genus in
<taxonomicName id="4C254D2B170F81714F29FF4EFF407A08" authorityName="Reitter" authorityYear="1909" box="[113,224,238,262]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="tribe" tribe="Paromalini">Paromalini</taxonomicName>
<figureCitation id="131E2A2D170E81704BA6F8F0FAE97C66" box="[1278,1353,1872,1896]" captionStart="Figure 5" captionStartId="6.[129,194,1370,1394]" captionTargetBox="[129,1473,144,1342]" captionTargetId="figure-262@6.[129,1473,144,1342]" captionTargetPageId="6" captionText="Figure 5. Cretomalus tibiodentatus gen. et sp. nov. A, dorsal view.B, ventral view; C, postmetacoxal area. D, postmesocoxal area. E, front leg and antenna. Abbreviations: fs, frontal stria; las, lateral stria on first abdominal ventrite; lms, lateral metaventral stria; pl, prosternal lobe; pms, prosternomesoventral stria; pmss, postmesocoxal stria; pmts, postmetacoxal stria." pageId="6" pageNumber="7">Fig. 5A</figureCitation>
), short and wide labrum with several pairs of prominent setae, short and wide prosternal lobe, vaguely defined antennal cavities located at prosternal hypomeron, basal prosternal margin rounded and projecting posteriorly, wide, curving protibia with a laterally directed median apical spur. The position of this genus in
<taxonomicName id="4C254D2B170F81714F29FF4EFF407A08" authorityName="Reitter" authorityYear="1909" box="[113,224,238,262]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="tribe" tribe="Paromalini">Paromalini</taxonomicName>
is also supported by all phylogenetic analyses performed in this study. Within extant
<taxonomicName id="4C254D2B170F81714EA9FEADFDC27A2B" authorityName="Reitter" authorityYear="1909" box="[497,610,269,293]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="tribe" tribe="Paromalini">Paromalini</taxonomicName>
<taxonomicName id="4C254D2B170F81714EA9FEADFDC27A2B" authorityName="Reitter" authorityYear="1909" box="[497,610,269,293]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="tribe" tribe="Paromalini">Paromalini</taxonomicName>
, possession of dorsal elytral striae in several genera (e.g.
<taxonomicName id="4C254D2B170F81714D4DFE8DFD6F7A4B" authority="Marseul" authorityName="Marseul" authorityYear="1855" box="[533,719,301,325]" class="Insecta" family="Histeridae" genus="Carcinops" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA170F81714D4DFE8DFDD77A4A" box="[533,631,301,324]" italics="true" pageId="7">Carcinops</emphasis>
<taxonomicName id="4C254D2B170F81714D4DFE8DFD6F7A4B" authority="Marseul" authorityName="Marseul" authorityYear="1855" box="[533,719,301,325]" class="Insecta" family="Histeridae" genus="Carcinops" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA170F81714D4DFE8DFDD77A4A" box="[533,631,301,324]" italics="true" pageId="7" pageNumber="8">Carcinops</emphasis>
Marseul
</taxonomicName>
and
<taxonomicName id="4C254D2B170F81714F29FEECFE9A7A6A" authority="Bickhardt" authorityName="Bickhardt" authorityYear="1921" box="[113,314,332,356]" class="Insecta" family="Histeridae" genus="Diplostix" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA170F81714F29FEECFF6A7A6A" box="[113,202,332,356]" italics="true" pageId="7">Diplostix</emphasis>
<taxonomicName id="4C254D2B170F81714F29FEECFE9A7A6A" authority="Bickhardt" authorityName="Bickhardt" authorityYear="1921" box="[113,314,332,356]" class="Insecta" family="Histeridae" genus="Diplostix" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA170F81714F29FEECFF6A7A6A" box="[113,202,332,356]" italics="true" pageId="7" pageNumber="8">Diplostix</emphasis>
Bickhardt
</taxonomicName>
) is probably plesiomorphic, while their absence is apomorphic in others (
<taxonomicName id="4C254D2B170F81714EF3FECBFDD47A8D" authority="Erichson" authorityName="Erichson" authorityYear="1834" box="[427,628,363,387]" class="Insecta" family="Histeridae" genus="Paromalus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA170F81714EF3FECBFDB37A8D" box="[427,531,363,387]" italics="true" pageId="7">Paromalus</emphasis>
<taxonomicName id="4C254D2B170F81714EF3FECBFDD47A8D" authority="Erichson" authorityName="Erichson" authorityYear="1834" box="[427,628,363,387]" class="Insecta" family="Histeridae" genus="Paromalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA170F81714EF3FECBFDB37A8D" box="[427,531,363,387]" italics="true" pageId="7" pageNumber="8">Paromalus</emphasis>
Erichson
</taxonomicName>
,
<taxonomicName id="4C254D2B170F81714D23FECBFF6D7AAD" authority="Cooman" authorityName="Cooman" authorityYear="1948" class="Insecta" family="Histeridae" genus="Platylomalus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA170F81714D23FECBFD5A7A8D" box="[635,762,363,387]" italics="true" pageId="7">Platylomalus</emphasis>
<taxonomicName id="4C254D2B170F81714D23FECBFF6D7AAD" authority="Cooman" authorityName="Cooman" authorityYear="1948" class="Insecta" family="Histeridae" genus="Platylomalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA170F81714D23FECBFD5A7A8D" box="[635,762,363,387]" italics="true" pageId="7" pageNumber="8">Platylomalus</emphasis>
Cooman
</taxonomicName>
) (
<bibRefCitation id="EFB44B59170F81714FB2FE2BFE3D7AAD" author="Zhou Y &amp; Ren D" box="[234,413,395,419]" pageId="7" pagination="521 - 39" refId="ref19290" refString="Zhou Y, Caterino Mº, Ren D et al. Phylogeny and evolution of Mesozoic and extant lineages of Histeridae (Coleoptera), with discovery of a new subfamily Antigracilinae from the Lower Cretaceous. Cladistics 2020; 36: 521 - 39. hups: // doi. org / 10.1111 / cla. 12418" type="journal article" year="2020">
<bibRefCitation id="EFB44B59170F81714FB2FE2BFE3D7AAD" author="Zhou Y &amp; Ren D" box="[234,413,395,419]" pageId="7" pageNumber="8" pagination="521 - 39" refId="ref19290" refString="Zhou Y, Caterino Mº, Ren D et al. Phylogeny and evolution of Mesozoic and extant lineages of Histeridae (Coleoptera), with discovery of a new subfamily Antigracilinae from the Lower Cretaceous. Cladistics 2020; 36: 521 - 39. hups: // doi. org / 10.1111 / cla. 12418" type="journal article" year="2020">
Zhou
<emphasis id="B951EABA170F81714E74FE2BFEC07AAD" box="[300,352,395,419]" italics="true" pageId="7">et al.</emphasis>
<emphasis id="B951EABA170F81714E74FE2BFEC07AAD" box="[300,352,395,419]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
2020
</bibRefCitation>
).
<taxonomicName id="4C254D2B170F81714EE0FE2BFD107AAD" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[440,688,395,419]" class="Insecta" family="Histeridae" genus="Cretomalus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="tibiodentatus">
<emphasis id="B951EABA170F81714EE0FE2BFD107AAD" box="[440,688,395,419]" italics="true" pageId="7">Cretomalus tibiodentatus</emphasis>
<taxonomicName id="4C254D2B170F81714EE0FE2BFD107AAD" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[440,688,395,419]" class="Insecta" family="Histeridae" genus="Cretomalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="tibiodentatus">
<emphasis id="B951EABA170F81714EE0FE2BFD107AAD" box="[440,688,395,419]" italics="true" pageId="7" pageNumber="8">Cretomalus tibiodentatus</emphasis>
</taxonomicName>
would appear to be more closely related to the laưer, particularly to
<taxonomicName id="4C254D2B170F81714F29FE69FF797AEF" authorityName="Erichson" authorityYear="1834" box="[113,217,457,481]" class="Insecta" family="Histeridae" genus="Paromalus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA170F81714F29FE69FF797AEF" box="[113,217,457,481]" italics="true" pageId="7">Paromalus</emphasis>
would appear to be more closely related to the latter, particularly to
<taxonomicName id="4C254D2B170F81714F29FE69FF797AEF" authorityName="Erichson" authorityYear="1834" box="[113,217,457,481]" class="Insecta" family="Histeridae" genus="Paromalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA170F81714F29FE69FF797AEF" box="[113,217,457,481]" italics="true" pageId="7" pageNumber="8">Paromalus</emphasis>
</taxonomicName>
, with which it shares several characters: absence of prosternal striation and complete frontal stria (
<figureCitation id="131E2A2D170F81714D0AFE49FD3C790E" box="[594,668,488,513]" captionStart="Figure 6" captionStartId="8.[129,194,1677,1701]" captionTargetBox="[129,1473,144,1650]" captionTargetId="figure-114@8.[129,1473,144,1650]" captionTargetPageId="8" captionText="Figure 6. Anapleus punctulatus sp.nov.. A, dorsal view.B, elytral epipleuron. C, ventral view.D, frontal view.Abbreviations:ees, elytral epipleural striae; lps, lateral pronotal stria; mes, marginal elytral stria; mps, marginal prosternal stria; ms, metaventral stria." pageId="7">Fig. 6A</figureCitation>
<figureCitation id="131E2A2D170F81714D0AFE49FD3C790E" box="[594,668,488,513]" captionStart="Figure 6" captionStartId="8.[129,194,1677,1701]" captionTargetBox="[129,1473,144,1650]" captionTargetId="figure-114@8.[129,1473,144,1650]" captionTargetPageId="8" captionText="Figure 6. Anapleus punctulatus sp.nov.. A, dorsal view.B, elytral epipleuron. C, ventral view.D, frontal view.Abbreviations:ees, elytral epipleural striae; lps, lateral pronotal stria; mes, marginal elytral stria; mps, marginal prosternal stria; ms, metaventral stria." pageId="7" pageNumber="8">Fig. 6A</figureCitation>
) (
<bibRefCitation id="EFB44B59170F81714DE0FE49FEAE792E" author="Zhang YJ &amp; Zhou HZ" pageId="7" pagination="1 - 40" refId="ref19205" refString="Zhang YJ, Zhou HZ. Taxonomy of the tribe Paromalini Reiuer (Coleoptera: Histeridae, Dendrophilinae) from China. Zootaxa 2007; 1544: 1 - 40. hups: // doi. org / 10.11646 / zootaxa. 1544.1.1" type="journal article" year="2007">Zhang and Zhou 2007</bibRefCitation>
<bibRefCitation id="EFB44B59170F81714DE0FE49FEAE792E" author="Zhang YJ &amp; Zhou HZ" pageId="7" pageNumber="8" pagination="1 - 40" refId="ref19205" refString="Zhang YJ, Zhou HZ. Taxonomy of the tribe Paromalini Reiuer (Coleoptera: Histeridae, Dendrophilinae) from China. Zootaxa 2007; 1544: 1 - 40. hups: // doi. org / 10.11646 / zootaxa. 1544.1.1" type="journal article" year="2007">Zhang and Zhou 2007</bibRefCitation>
).
<taxonomicName id="4C254D2B170F81714E7CFDA8FDB6792E" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[292,534,520,544]" class="Insecta" family="Histeridae" genus="Cretomalus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="tibiodentatus">
<emphasis id="B951EABA170F81714E7CFDA8FDB6792E" box="[292,534,520,544]" italics="true" pageId="7">Cretomalus tibiodentatus</emphasis>
<taxonomicName id="4C254D2B170F81714E7CFDA8FDB6792E" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[292,534,520,544]" class="Insecta" family="Histeridae" genus="Cretomalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="tibiodentatus">
<emphasis id="B951EABA170F81714E7CFDA8FDB6792E" box="[292,534,520,544]" italics="true" pageId="7" pageNumber="8">Cretomalus tibiodentatus</emphasis>
</taxonomicName>
is significantly smaller than all
<taxonomicName id="4C254D2B170F81714F90FD87FE287931" box="[200,392,551,575]" class="Insecta" family="Histeridae" genus="Paromalus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B951EABA170F81714F90FD87FE907931" box="[200,304,551,575]" italics="true" pageId="7">Paromalus</emphasis>
<taxonomicName id="4C254D2B170F81714F90FD87FE287931" box="[200,392,551,575]" class="Insecta" family="Histeridae" genus="Paromalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B951EABA170F81714F90FD87FE907931" box="[200,304,551,575]" italics="true" pageId="7" pageNumber="8">Paromalus</emphasis>
species.
</taxonomicName>
Ŋe serrated protibial margin (
<figureCitation id="131E2A2D170F81714D8CFD87FF2D7950" captionStart="Figure 5" captionStartId="6.[129,194,1370,1394]" captionTargetBox="[129,1473,144,1342]" captionTargetId="figure-262@6.[129,1473,144,1342]" captionTargetPageId="6" captionText="Figure 5. Cretomalus tibiodentatus gen. et sp. nov. A, dorsal view.B, ventral view; C, postmetacoxal area. D, postmesocoxal area. E, front leg and antenna. Abbreviations: fs, frontal stria; las, lateral stria on first abdominal ventrite; lms, lateral metaventral stria; pl, prosternal lobe; pms, prosternomesoventral stria; pmss, postmesocoxal stria; pmts, postmetacoxal stria." pageId="7">Fig. 5E</figureCitation>
The serrated protibial margin (
<figureCitation id="131E2A2D170F81714D8CFD87FF2D7950" captionStart="Figure 5" captionStartId="6.[129,194,1370,1394]" captionTargetBox="[129,1473,144,1342]" captionTargetId="figure-262@6.[129,1473,144,1342]" captionTargetPageId="6" captionText="Figure 5. Cretomalus tibiodentatus gen. et sp. nov. A, dorsal view.B, ventral view; C, postmetacoxal area. D, postmesocoxal area. E, front leg and antenna. Abbreviations: fs, frontal stria; las, lateral stria on first abdominal ventrite; lms, lateral metaventral stria; pl, prosternal lobe; pms, prosternomesoventral stria; pmss, postmesocoxal stria; pmts, postmetacoxal stria." pageId="7" pageNumber="8">Fig. 5E</figureCitation>
) is unique among the entire tribe.
</paragraph>
<paragraph id="8B9A36A8170F81714F29FD21FF697E05" blockId="7.[113,764,640,1291]" pageId="7">
<emphasis id="B951EABA170F81714F29FD21FF777996" box="[113,215,641,664]" italics="true" pageId="7">Diagnosis:</emphasis>
Frons weakly convex and finely punctuate, punctures separated by three times their diameter. Pronotum relatively long, punctuated with punctures separated by two to three times their diameter. Elytra without striation, punctuation with the same paưern as in pronotum. Prosternal keel with fine punctuation. Marginal mesoventral stria complete, continuous with lateral metaventral stria. Transverse mesoventral stria not visible. Lateral metaventral striae present, extending posterolaterad on at least anterior half. Postmesocoxal striae present, running posteriorly. Protibia wide and strongly curved, with six to seven small, symmetrical teeth along the outer margin. Protarsal groove weakly developed. ºmall inner and curved median protibial spurs present. ºimilarly shaped
<taxonomicName id="4C254D2B170F81714E9DFC58FD907F1E" box="[453,560,1016,1040]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="family">Histeridae</taxonomicName>
</subSubSection>
<subSubSection id="C33F6523170F81714F29FD21FF697E05" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="8B9A36A8170F81714F29FD21FF697E05" blockId="7.[113,764,640,1291]" pageId="7" pageNumber="8">
<emphasis id="B951EABA170F81714F29FD21FF777996" box="[113,215,641,664]" italics="true" pageId="7" pageNumber="8">Diagnosis:</emphasis>
Frons weakly convex and finely punctuate, punctures separated by three times their diameter. Pronotum relatively long, punctuated with punctures separated by two to three times their diameter. Elytra without striation, punctuation with the same pattern as in pronotum. Prosternal keel with fine punctuation. Marginal mesoventral stria complete, continuous with lateral metaventral stria. Transverse mesoventral stria not visible. Lateral metaventral striae present, extending posterolaterad on at least anterior half. Postmesocoxal striae present, running posteriorly. Protibia wide and strongly curved, with six to seven small, symmetrical teeth along the outer margin. Protarsal groove weakly developed. Small inner and curved median protibial spurs present. Similarly shaped
<taxonomicName id="4C254D2B170F81714E9DFC58FD907F1E" box="[453,560,1016,1040]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="family">Histeridae</taxonomicName>
found in Burmese amber are
<taxonomicName id="4C254D2B170F81714FB2FBB7FE687F21" authorityName="Simon Prazak &amp; Lackner" authorityYear="2023" box="[234,456,1047,1071]" class="Insecta" family="Histeridae" genus="Platycretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="muscularis">
<emphasis id="B951EABA170F81714FB2FBB7FE687F21" box="[234,456,1047,1071]" italics="true" pageId="7">Platycretus muscularis</emphasis>
<taxonomicName id="4C254D2B170F81714FB2FBB7FE687F21" authorityName="Simon Prazak &amp; Lackner" authorityYear="2023" box="[234,456,1047,1071]" class="Insecta" family="Histeridae" genus="Platycretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="muscularis">
<emphasis id="B951EABA170F81714FB2FBB7FE687F21" box="[234,456,1047,1071]" italics="true" pageId="7" pageNumber="8">Platycretus muscularis</emphasis>
</taxonomicName>
ºimon-Pražák and
<bibRefCitation id="EFB44B59170F81714DFAFBB7FF077F41" author="Lackner T &amp; Zhang YM &amp; Kindler C" pageId="7" pagination="48 - 71" refId="ref18368" refString="Lackner T, Zhang YM, Kindler C et al. ºaprininae: phylogeny, biogeography and a new classification of the subfamily (Coleoptera: Histeridae). Systematic Entomology 2023; 49: 48 - 71. hups: // doi. org / 10.1111 / syen. 12606" type="journal article" year="2023">Lackner, 2023</bibRefCitation>
Simon-Pražák and
<bibRefCitation id="EFB44B59170F81714DFAFBB7FF077F41" author="Lackner T &amp; Zhang YM &amp; Kindler C" pageId="7" pageNumber="8" pagination="48 - 71" refId="ref18368" refString="Lackner T, Zhang YM, Kindler C et al. ºaprininae: phylogeny, biogeography and a new classification of the subfamily (Coleoptera: Histeridae). Systematic Entomology 2023; 49: 48 - 71. hups: // doi. org / 10.1111 / syen. 12606" type="journal article" year="2023">Lackner, 2023</bibRefCitation>
,
<taxonomicName id="4C254D2B170F81714FECFB96FE317F40" authorityName="Simon Prazak &amp; Lackner" authorityYear="2023" box="[180,401,1078,1102]" class="Insecta" family="Histeridae" genus="Olexum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="complanatum">
<emphasis id="B951EABA170F81714FECFB96FE317F40" box="[180,401,1078,1102]" italics="true" pageId="7">Olexum complanatum</emphasis>
<taxonomicName id="4C254D2B170F81714FECFB96FE317F40" authorityName="Simon Prazak &amp; Lackner" authorityYear="2023" box="[180,401,1078,1102]" class="Insecta" family="Histeridae" genus="Olexum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="complanatum">
<emphasis id="B951EABA170F81714FECFB96FE317F40" box="[180,401,1078,1102]" italics="true" pageId="7" pageNumber="8">Olexum complanatum</emphasis>
</taxonomicName>
ºimon-Pražák and
<bibRefCitation id="EFB44B59170F81714D38FB96FD567F41" author="Lackner T &amp; Zhang YM &amp; Kindler C" box="[608,758,1078,1103]" pageId="7" pagination="48 - 71" refId="ref18368" refString="Lackner T, Zhang YM, Kindler C et al. ºaprininae: phylogeny, biogeography and a new classification of the subfamily (Coleoptera: Histeridae). Systematic Entomology 2023; 49: 48 - 71. hups: // doi. org / 10.1111 / syen. 12606" type="journal article" year="2023">Lackner, 2023</bibRefCitation>
Simon-Pražák and
<bibRefCitation id="EFB44B59170F81714D38FB96FD567F41" author="Lackner T &amp; Zhang YM &amp; Kindler C" box="[608,758,1078,1103]" pageId="7" pageNumber="8" pagination="48 - 71" refId="ref18368" refString="Lackner T, Zhang YM, Kindler C et al. ºaprininae: phylogeny, biogeography and a new classification of the subfamily (Coleoptera: Histeridae). Systematic Entomology 2023; 49: 48 - 71. hups: // doi. org / 10.1111 / syen. 12606" type="journal article" year="2023">Lackner, 2023</bibRefCitation>
, and
<taxonomicName id="4C254D2B170F81714FF9FBF6FE567F60" authority="Caterino, 2021" authorityName="Caterino" authorityYear="2021" box="[161,502,1110,1134]" class="Insecta" family="Histeridae" genus="Druantia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="aeterna">
<emphasis id="B951EABA170F81714FF9FBF6FEED7F63" box="[161,333,1110,1133]" italics="true" pageId="7">Druantia aeterna</emphasis>
<bibRefCitation id="EFB44B59170F81714E0FFBF6FE567F60" author="Caterino" box="[343,502,1110,1134]" pageId="7" pagination="211 - 21" refId="ref17393" refString="Caterino Mº. New fossil histerid species from Cretaceous Burmese amber (Coleoptera: Histeridae). Ne Coleopterists Bulletin 2021; 75: 211 - 21." type="journal article" year="2021">Caterino, 2021</bibRefCitation>
<taxonomicName id="4C254D2B170F81714FF9FBF6FE567F60" authority="Caterino, 2021" authorityName="Caterino" authorityYear="2021" box="[161,502,1110,1134]" class="Insecta" family="Histeridae" genus="Druantia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="aeterna">
<emphasis id="B951EABA170F81714FF9FBF6FEED7F63" box="[161,333,1110,1133]" italics="true" pageId="7" pageNumber="8">Druantia aeterna</emphasis>
<bibRefCitation id="EFB44B59170F81714E0FFBF6FE567F60" author="Caterino" box="[343,502,1110,1134]" pageId="7" pageNumber="8" pagination="211 - 21" refId="ref17393" refString="Caterino Mº. New fossil histerid species from Cretaceous Burmese amber (Coleoptera: Histeridae). Ne Coleopterists Bulletin 2021; 75: 211 - 21." type="journal article" year="2021">Caterino, 2021</bibRefCitation>
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.
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<emphasis id="B951EABA170F81714D5BFBF6FD5A7F60" box="[515,762,1110,1134]" italics="true" pageId="7">Cretomalus tibiodentatus</emphasis>
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<emphasis id="B951EABA170F81714D5BFBF6FD5A7F60" box="[515,762,1110,1134]" italics="true" pageId="7" pageNumber="8">Cretomalus tibiodentatus</emphasis>
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can be distinguished from all of them by combination of following characters: long and narrow body (length: width 5: 2), shape of the prosterno-mesoventral suture (with a curved anteriorly pointed projection) and the state of regularly toothed protibia.
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<emphasis id="B951EABA170F81714F29FA8CFF417E4A" box="[113,225,1324,1348]" italics="true" pageId="7">Etymology:</emphasis>
Ŋe specific name refers to the unique protibial dentation.
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<subSubSection id="C33F6523170F81714F29FA8CFF7B7E6D" pageId="7" pageNumber="8" type="etymology">
<paragraph id="8B9A36A8170F81714F29FA8CFF7B7E6D" blockId="7.[113,763,1324,1379]" pageId="7" pageNumber="8">
<emphasis id="B951EABA170F81714F29FA8CFF417E4A" box="[113,225,1324,1348]" italics="true" pageId="7" pageNumber="8">Etymology:</emphasis>
The specific name refers to the unique protibial dentation.
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</treatment>

180
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<heading id="D0D281C4170D81734CAFFEBDFB467A36" bold="true" box="[1015,1254,285,312]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="6" reason="2">
<emphasis id="B951EABA170D81734CAFFEBDFB467A36" bold="true" box="[1015,1254,285,312]" pageId="5" pageNumber="6">
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<emphasis id="B951EABA170D81734CAFFEBDFBD97A36" bold="true" box="[1015,1145,285,312]" italics="true" pageId="5" pageNumber="6">Cretomalus</emphasis>
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<taxonomicNameLabel id="A26257C1170D81734B26FEBEFB467A36" box="[1150,1254,286,312]" pageId="5" rank="genus">gen. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="A26257C1170D81734B26FEBEFB467A36" box="[1150,1254,286,312]" pageId="5" pageNumber="6" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C33F6523170D81734C72FEFFFAE2783C" pageId="5" type="materials_examined">
<paragraph id="8B9A36A8170D81734C72FEFFFB547A78" blockId="5.[810,1268,350,374]" box="[810,1268,350,374]" pageId="5">
<emphasis id="B951EABA170D81734C72FEFFFB3E7A78" box="[810,1182,350,374]" italics="true" pageId="5">
<typeStatus id="549E880A170D81734C72FEFFFCFA7A78" box="[810,858,351,374]" pageId="5">Type</typeStatus>
<subSubSection id="C33F6523170D81734C72FEFFFB547A78" box="[810,1268,350,374]" pageId="5" pageNumber="6" type="type_taxon">
<paragraph id="8B9A36A8170D81734C72FEFFFB547A78" blockId="5.[810,1268,350,374]" box="[810,1268,350,374]" pageId="5" pageNumber="6">
<emphasis id="B951EABA170D81734C72FEFFFB3E7A78" box="[810,1182,350,374]" italics="true" pageId="5" pageNumber="6">
<typeStatus id="549E880A170D81734C72FEFFFCFA7A78" box="[810,858,351,374]" pageId="5" pageNumber="6">Type</typeStatus>
species:
<taxonomicName id="4C254D2B170D81734CF3FEFEFB3E7A78" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[939,1182,350,374]" class="Insecta" family="Histeridae" genus="Cretomalus" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="tibiodentatus" status="sp. nov.">Cretomalus tibiodentatus</taxonomicName>
<taxonomicName id="4C254D2B170D81734CF3FEFEFB3E7A78" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[939,1182,350,374]" class="Insecta" family="Histeridae" genus="Cretomalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="tibiodentatus" status="sp. nov.">Cretomalus tibiodentatus</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A26257C1170D81734BFBFEFFFB4A7A78" box="[1187,1258,351,374]" pageId="5" rank="species">sp. nov</taxonomicNameLabel>
<taxonomicNameLabel id="A26257C1170D81734BFBFEFFFB4A7A78" box="[1187,1258,351,374]" pageId="5" pageNumber="6" rank="species">sp. nov</taxonomicNameLabel>
..
</paragraph>
<paragraph id="8B9A36A8170D81734C72FE29FB6079B4" blockId="5.[810,1460,392,698]" pageId="5">
<emphasis id="B951EABA170D81734C72FE29FC307AAE" box="[810,912,393,416]" italics="true" pageId="5">Diagnosis:</emphasis>
Prolonged, rectangular, parallel-sided, and flaưened body shape. Frontoclypeal area slightly convex with a complete and thin frontal stria. Labrum short and trapezoidal. Antennae with a flat and round club with two visible annuli. Astriate elytra. Basal prosternal margin rounded, projecting posteriorly. Antennal cavities are vaguely defined and widely open, located posterolaterally (covered by the front leg when retracted). Mesoventrite very short and wide (1: 3 ratio). Anterior margin curved inwards to fit the basal prosternal margin. Outer protibial margin serrated with symmetrical teeth.
</subSubSection>
<subSubSection id="C33F6523170D81734C72FE29FB6079B4" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="8B9A36A8170D81734C72FE29FB6079B4" blockId="5.[810,1460,392,698]" pageId="5" pageNumber="6">
<emphasis id="B951EABA170D81734C72FE29FC307AAE" box="[810,912,393,416]" italics="true" pageId="5" pageNumber="6">Diagnosis:</emphasis>
Prolonged, rectangular, parallel-sided, and flattened body shape. Frontoclypeal area slightly convex with a complete and thin frontal stria. Labrum short and trapezoidal. Antennae with a flat and round club with two visible annuli. Astriate elytra. Basal prosternal margin rounded, projecting posteriorly. Antennal cavities are vaguely defined and widely open, located posterolaterally (covered by the front leg when retracted). Mesoventrite very short and wide (1: 3 ratio). Anterior margin curved inwards to fit the basal prosternal margin. Outer protibial margin serrated with symmetrical teeth.
</paragraph>
<paragraph id="8B9A36A8170D81734C72FD7CFAE2783C" blockId="5.[810,1459,732,818]" pageId="5">
<emphasis id="B951EABA170D81734C72FD7CFC3979FA" box="[810,921,732,756]" italics="true" pageId="5">Etymology:</emphasis>
</subSubSection>
<subSubSection id="C33F6523170D81734C72FD7CFAE2783C" pageId="5" pageNumber="6" type="etymology">
<paragraph id="8B9A36A8170D81734C72FD7CFAE2783C" blockId="5.[810,1459,732,818]" pageId="5" pageNumber="6">
<emphasis id="B951EABA170D81734C72FD7CFC3979FA" box="[810,921,732,756]" italics="true" pageId="5" pageNumber="6">Etymology:</emphasis>
‘Creto-’ signifies the Cretaceous origin, ‘-malus’ is shared with other
<taxonomicName id="4C254D2B170D81734CB1FD5BFBF8781D" authorityName="Reitter" authorityYear="1909" box="[1001,1112,763,787]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="tribe" tribe="Paromalini">Paromalini</taxonomicName>
<taxonomicName id="4C254D2B170D81734CB1FD5BFBF8781D" authorityName="Reitter" authorityYear="1909" box="[1001,1112,763,787]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="tribe" tribe="Paromalini">Paromalini</taxonomicName>
genera to which it is related (e.g.
<taxonomicName id="4C254D2B170D81734C72FCBAFC31783C" authorityName="Cooman" authorityYear="1937" box="[810,913,794,818]" class="Insecta" family="Histeridae" genus="Eulomalus" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA170D81734C72FCBAFC31783C" box="[810,913,794,818]" italics="true" pageId="5">Eulomalus</emphasis>
<taxonomicName id="4C254D2B170D81734C72FCBAFC31783C" authorityName="Cooman" authorityYear="1937" box="[810,913,794,818]" class="Insecta" family="Histeridae" genus="Eulomalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA170D81734C72FCBAFC31783C" box="[810,913,794,818]" italics="true" pageId="5" pageNumber="6">Eulomalus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D2B170D81734CC4FCBAFBA4783C" authorityName="Erichson" authorityYear="1834" box="[924,1028,794,818]" class="Insecta" family="Histeridae" genus="Paromalus" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA170D81734CC4FCBAFBA4783C" box="[924,1028,794,818]" italics="true" pageId="5">Paromalus</emphasis>
<taxonomicName id="4C254D2B170D81734CC4FCBAFBA4783C" authorityName="Erichson" authorityYear="1834" box="[924,1028,794,818]" class="Insecta" family="Histeridae" genus="Paromalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA170D81734CC4FCBAFBA4783C" box="[924,1028,794,818]" italics="true" pageId="5" pageNumber="6">Paromalus</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C254D2B170D81734B61FCBAFB1E783C" authorityName="Schmidt" authorityYear="1897" box="[1081,1214,794,818]" class="Insecta" family="Histeridae" genus="Pachylomalus" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA170D81734B61FCBAFB1E783C" box="[1081,1214,794,818]" italics="true" pageId="5">Pachylomalus</emphasis>
<taxonomicName id="4C254D2B170D81734B61FCBAFB1E783C" authorityName="Schmidt" authorityYear="1897" box="[1081,1214,794,818]" class="Insecta" family="Histeridae" genus="Pachylomalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA170D81734B61FCBAFB1E783C" box="[1081,1214,794,818]" italics="true" pageId="5" pageNumber="6">Pachylomalus</emphasis>
</taxonomicName>
), masculine.
</paragraph>

290
data/03/8C/87/038C87BE170F817F4E55F9B8FD277A0F.xml
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@ -1,218 +1,230 @@
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<mods:namePart id="FED5891E9C6DC4D25424935B553814CB">Caterino, Michael S</mods:namePart>
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<mods:name id="3EF814EC18391BA2B418AA92F36E9551" type="personal">
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<mods:namePart id="E6BD7E9769E1E2B0ADE2ECFDF993B016">Michael</mods:namePart>
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<mods:name id="7611BA83D2853784CEA48AF19E1EBD2D" type="personal">
<mods:role id="D981D64F10644AE95B99D182D01547CA">
<mods:roleTerm id="6B255865E2BDE24BC3B626EACC512302">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EA3E1691464243379CFAE9EF249F056A">Caterino</mods:namePart>
<mods:affiliation id="8F4D52D886A899AC201317B2B3C9DD0A">Department of Plant and Environmental ºciences, Clemson University, Clemson, ºC 29634, United ºtates</mods:affiliation>
</mods:name>
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<mods:title id="10F2DD3B901E0303A38DAE3E48F05C72">Zoological Journal of the Linnean Society</mods:title>
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<mods:title id="8827A1549A9929B4C5F9908C9F421A99">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="6E69CFCF82677075BB0C7B31A926B0A9">2024</mods:date>
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<mods:title id="FE4AC822837D2F32D8A7E44A98859CCE">zlae 137</mods:title>
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<mods:number id="EF03B003A55F13DA807A39249FED15EF">202</mods:number>
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<mods:number id="07B18B79862C616BB9844B6915AFAB43">2024-11-20</mods:number>
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<treatment id="038C87BE170F817F4E55F9B8FD277A0F" ID-DOI="http://doi.org/10.5281/zenodo.14424548" ID-Zenodo-Dep="14424548" LSID="urn:lsid:plazi:treatment:038C87BE170F817F4E55F9B8FD277A0F" httpUri="http://treatment.plazi.org/id/038C87BE170F817F4E55F9B8FD277A0F" lastPageId="9" pageId="7">
<subSubSection id="C33F6523170F81714E55F9B8FDFE7D3D" box="[269,606,1560,1587]" pageId="7" type="nomenclature">
<paragraph id="8B9A36A8170F81714E55F9B8FDFE7D3D" blockId="7.[269,606,1560,1587]" box="[269,606,1560,1587]" pageId="7">
<heading id="D0D281C4170F81714E55F9B8FDFE7D3D" bold="true" box="[269,606,1560,1587]" centered="true" fontSize="11" level="2" pageId="7" reason="2">
<emphasis id="B951EABA170F81714E55F9B8FDFE7D3D" bold="true" box="[269,606,1560,1587]" pageId="7">
<taxonomicName id="4C254D2B170F81714E55F9B8FDA27D3D" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[269,514,1560,1587]" class="Insecta" family="Histeridae" genus="Anapleus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="punctulatus" status="sp. nov.">
<emphasis id="B951EABA170F81714E55F9B8FDA27D3D" bold="true" box="[269,514,1560,1587]" italics="true" pageId="7">Anapleus punctulatus</emphasis>
<treatment id="038C87BE170F817F4E55F9B8FD277A0F" ID-DOI="http://doi.org/10.5281/zenodo.14424548" ID-Zenodo-Dep="14424548" LSID="urn:lsid:plazi:treatment:038C87BE170F817F4E55F9B8FD277A0F" httpUri="http://treatment.plazi.org/id/038C87BE170F817F4E55F9B8FD277A0F" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8">
<subSubSection id="C33F6523170F81714E55F9B8FDFE7D3D" box="[269,606,1560,1587]" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="8B9A36A8170F81714E55F9B8FDFE7D3D" blockId="7.[269,606,1560,1587]" box="[269,606,1560,1587]" pageId="7" pageNumber="8">
<heading id="D0D281C4170F81714E55F9B8FDFE7D3D" bold="true" box="[269,606,1560,1587]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="8" reason="2">
<emphasis id="B951EABA170F81714E55F9B8FDFE7D3D" bold="true" box="[269,606,1560,1587]" pageId="7" pageNumber="8">
<taxonomicName id="4C254D2B170F81714E55F9B8FDA27D3D" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[269,514,1560,1587]" class="Insecta" family="Histeridae" genus="Anapleus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="punctulatus" status="sp. nov.">
<emphasis id="B951EABA170F81714E55F9B8FDA27D3D" bold="true" box="[269,514,1560,1587]" italics="true" pageId="7" pageNumber="8">Anapleus punctulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C1170F81714D5FF9B9FDFE7D3D" box="[519,606,1561,1587]" pageId="7" rank="species">sp. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="A26257C1170F81714D5FF9B9FDFE7D3D" box="[519,606,1561,1587]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33F6523170F817F4ED7F9EAFD277A0F" lastPageId="9" pageId="7" type="description">
<paragraph id="8B9A36A8170F81714ED7F9EAFE7D7D6A" blockId="7.[399,477,1610,1637]" box="[399,477,1610,1637]" pageId="7">
<emphasis id="B951EABA170F81714ED7F9EAFE7D7D6A" bold="true" box="[399,477,1610,1637]" pageId="7">
<subSubSection id="C33F6523170F81714ED7F9EAFE7D7D6A" box="[399,477,1610,1637]" pageId="7" pageNumber="8" type="description">
<paragraph id="8B9A36A8170F81714ED7F9EAFE7D7D6A" blockId="7.[399,477,1610,1637]" box="[399,477,1610,1637]" pageId="7" pageNumber="8">
<emphasis id="B951EABA170F81714ED7F9EAFE7D7D6A" bold="true" box="[399,477,1610,1637]" pageId="7" pageNumber="8">
(
<figureCitation id="131E2A2D170F81714EC1F9EBFE727D6B" box="[409,466,1611,1637]" captionStart="Figure 6" captionStartId="8.[129,194,1677,1701]" captionTargetBox="[129,1473,144,1650]" captionTargetId="figure-114@8.[129,1473,144,1650]" captionTargetPageId="8" captionText="Figure 6. Anapleus punctulatus sp.nov.. A, dorsal view.B, elytral epipleuron. C, ventral view.D, frontal view.Abbreviations:ees, elytral epipleural striae; lps, lateral pronotal stria; mes, marginal elytral stria; mps, marginal prosternal stria; ms, metaventral stria." pageId="7">Fig. 6</figureCitation>
<figureCitation id="131E2A2D170F81714EC1F9EBFE727D6B" box="[409,466,1611,1637]" captionStart="Figure 6" captionStartId="8.[129,194,1677,1701]" captionTargetBox="[129,1473,144,1650]" captionTargetId="figure-114@8.[129,1473,144,1650]" captionTargetPageId="8" captionText="Figure 6. Anapleus punctulatus sp.nov.. A, dorsal view.B, elytral epipleuron. C, ventral view.D, frontal view.Abbreviations:ees, elytral epipleural striae; lps, lateral pronotal stria; mes, marginal elytral stria; mps, marginal prosternal stria; ms, metaventral stria." pageId="7" pageNumber="8">Fig. 6</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8B9A36A8170F81714F29F92BFD017DEC" blockId="7.[113,763,1675,1762]" pageId="7">
<emphasis id="B951EABA170F81714F29F92BFF6D7DAD" box="[113,205,1675,1699]" italics="true" pageId="7">Material:</emphasis>
Hkamti amber (
<emphasis id="B951EABA170F81714E2EF92CFEDF7DAD" box="[374,383,1676,1699]" italics="true" pageId="7">c</emphasis>
</subSubSection>
<subSubSection id="C33F6523170F81714F29F92BFD017DEC" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="8B9A36A8170F81714F29F92BFD017DEC" blockId="7.[113,763,1675,1762]" pageId="7" pageNumber="8">
<emphasis id="B951EABA170F81714F29F92BFF6D7DAD" box="[113,205,1675,1699]" italics="true" pageId="7" pageNumber="8">Material:</emphasis>
<materialsCitation id="3B4D3CF5170F81714F8AF92BFD3C7DEC" collectionCode="HUM" location="Hkamti" municipality="Hkamti" pageId="7" pageNumber="8" specimenCode="SEHU-0000121214" specimenCount="1">
<collectingMunicipality id="6BFEACD2170F81714F8AF92BFE817DAD" box="[210,289,1675,1699]" pageId="7" pageNumber="8">Hkamti</collectingMunicipality>
amber (
<emphasis id="B951EABA170F81714E2EF92CFEDF7DAD" box="[374,383,1676,1699]" italics="true" pageId="7" pageNumber="8">c</emphasis>
. 110 Mya). Amber piece dimensions: 14 × 8 ×
<quantity id="4CDD9B4D170F81714F8AF90BFEAD7DCC" box="[210,269,1707,1731]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="7" unit="mm" value="1.0">1 mm</quantity>
. Amber clear with several insect fragments and air bubbles. Deposited at HUM (ºEHU-0000121214).
<quantity id="4CDD9B4D170F81714F8AF90BFEAD7DCC" box="[210,269,1707,1731]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="8" unit="mm" value="1.0">1 mm</quantity>
. Amber clear with several insect fragments and air bubbles. Deposited at
<collectionCode id="ED34AE6D170F81714E2BF96AFE127DEF" box="[371,434,1738,1761]" pageId="7" pageNumber="8">HUM</collectionCode>
(
<specimenCode id="DB839ED3170F81714EE6F96AFD357DEC" box="[446,661,1738,1762]" pageId="7" pageNumber="8">SEHU-0000121214</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B9A36A8170F81714F29F8A4FC507DC8" blockId="7.[113,763,1795,1976]" lastBlockId="7.[810,1460,144,1734]" pageId="7">
<emphasis id="B951EABA170F81714F29F8A4FEA57C15" box="[113,261,1796,1819]" italics="true" pageId="7">Measurements:</emphasis>
</subSubSection>
<subSubSection id="C33F6523170F81714F29F8A4FC507DC8" pageId="7" pageNumber="8" type="description">
<paragraph id="8B9A36A8170F81714F29F8A4FC507DC8" blockId="7.[113,763,1795,1976]" lastBlockId="7.[810,1460,144,1734]" pageId="7" pageNumber="8">
<emphasis id="B951EABA170F81714F29F8A4FEA57C15" box="[113,261,1796,1819]" italics="true" pageId="7" pageNumber="8">Measurements:</emphasis>
Head width:
<quantity id="4CDD9B4D170F81714EC7F8A3FE4E7C15" box="[415,494,1795,1819]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="7" unit="mm" value="0.2">0.2 mm</quantity>
<quantity id="4CDD9B4D170F81714EC7F8A3FE4E7C15" box="[415,494,1795,1819]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="8" unit="mm" value="0.2">0.2 mm</quantity>
; width between anterior pronotal angles:
<quantity id="4CDD9B4D170F81714E41F882FED37C34" box="[281,371,1826,1850]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="7" unit="mm" value="0.25">0.25 mm</quantity>
<quantity id="4CDD9B4D170F81714E41F882FED37C34" box="[281,371,1826,1850]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="8" unit="mm" value="0.25">0.25 mm</quantity>
; width between posterior pronotal angles:
<quantity id="4CDD9B4D170F81714FFFF8E2FF557C57" box="[167,245,1858,1882]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="7" unit="mm" value="0.6">0.6 mm</quantity>
<quantity id="4CDD9B4D170F81714FFFF8E2FF557C57" box="[167,245,1858,1882]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="8" unit="mm" value="0.6">0.6 mm</quantity>
; pronotal length:
<quantity id="4CDD9B4D170F81714EEAF8E2FDAE7C57" box="[434,526,1858,1882]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="7" unit="mm" value="0.31">0.31 mm</quantity>
<quantity id="4CDD9B4D170F81714EEAF8E2FDAE7C57" box="[434,526,1858,1882]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="7" pageNumber="8" unit="mm" value="0.31">0.31 mm</quantity>
; sutural elytral length:
<quantity id="4CDD9B4D170F81714F2BF8C1FF6C7C77" box="[115,204,1889,1913]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="7" unit="mm" value="0.65">0.65 mm</quantity>
<quantity id="4CDD9B4D170F81714F2BF8C1FF6C7C77" box="[115,204,1889,1913]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="7" pageNumber="8" unit="mm" value="0.65">0.65 mm</quantity>
; elytral width (in pair, across widest point):
<quantity id="4CDD9B4D170F81714DD2F8C1FD777C77" box="[650,727,1889,1913]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="7" unit="mm" value="0.8">0.8 mm</quantity>
<quantity id="4CDD9B4D170F81714DD2F8C1FD777C77" box="[650,727,1889,1913]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="8" unit="mm" value="0.8">0.8 mm</quantity>
.—
<emphasis id="B951EABA170F81714F29F820FF127C96" bold="true" box="[113,178,1920,1944]" italics="true" pageId="7">Body:</emphasis>
<emphasis id="B951EABA170F81714F29F820FF127C96" bold="true" box="[113,178,1920,1944]" italics="true" pageId="7" pageNumber="8">Body:</emphasis>
Rounded widest at humeri, anteriorly narrowed. Dorsal and ventral surface with dense and rugose punctuation. Lateral body margin elevated dorsally. Cuticle black, legs chestnut brown.—
<emphasis id="B951EABA170F81714CD6FF0FFC707BC9" bold="true" box="[910,976,175,199]" italics="true" pageId="7">Head:</emphasis>
Frons and clypeus punctuate (punctures separated by two to three times their diameter). Frontoclypeal area with a round concavity. Frontal and frontoclypeal striae absent. Labrum trapezoidal with several setae. Mandibles robust, with outer surfaces punctate; Head retracted, other mouthparts not observable. Antennal insertions rather broadly open to front; ºcape short, slender, only weakly widened to apex; Pedicel slender, mostly hidden; Antennomeres 3–8 simple, rounded. Antennal club round, with distinct annuli and several prominent setae.—
<emphasis id="B951EABA170F81714B73FE69FB207AEF" bold="true" box="[1067,1152,457,481]" italics="true" pageId="7">Ŋorax:</emphasis>
Pronotum convex medially. Entire pronotal disc covered with dense and rugose punctuation (punctures separated by their diameter). Basal margin rounded, slightly angularly projecting posteriorly. Pronotum bi-margined laterally, strongest lateral carina aligned with marginal elytral carina; outer marginal pronotal stria similarly carinate, narrowing and descending below inner marginal in anterior half, overlapped and invisible from above in the anterior third. ºcutellar shield obscured. Elytra with the same punctuation paưern as pronotal disc. Elytral disc slightly convex, with two weakly elevated longitudinal carinae located in medial part. Elytra carinate along the outer margin and truncate posteriorly. ºutural area elevated in posterior third forming a keel. Marginal elytral stria carinate, deeply impressed. Elytral epipleuron with two carinate striae. Prosternal lobe very short and broad with dense punctuation, with incised furrows laterally for passage of the antennal funicle; Prosternal keel broad, similarly punctate to prosternal lobe, with lateral carinae weakly converging anterad, abruptly narrowing at junction with prosternal lobe; basal prosternal margin weakly and broadly emarginate. No striation visible on prosternum. Antennal cavities widely open, located posterolaterally on prosternal sides. Mesoventrite wide, densely and coarsely punctuated. Anterior margin outwardly curved along prosternal margin. Metaventrite with a pair of lateral metaventral striae present in anterior two-thirds, extending laterally to near metepisternum. Postmesocoxal striae curved along mesocoxae. Punctures on metaventrite become sparser and larger posteriorly.—
<emphasis id="B951EABA170F81714B74FAB7FBC47E20" bold="true" box="[1068,1124,1303,1326]" italics="true" pageId="7">Legs:</emphasis>
<emphasis id="B951EABA170F81714CD6FF0FFC707BC9" bold="true" box="[910,976,175,199]" italics="true" pageId="7" pageNumber="8">Head:</emphasis>
Frons and clypeus punctuate (punctures separated by two to three times their diameter). Frontoclypeal area with a round concavity. Frontal and frontoclypeal striae absent. Labrum trapezoidal with several setae. Mandibles robust, with outer surfaces punctate; Head retracted, other mouthparts not observable. Antennal insertions rather broadly open to front; Scape short, slender, only weakly widened to apex; Pedicel slender, mostly hidden; Antennomeres 3–8 simple, rounded. Antennal club round, with distinct annuli and several prominent setae.—
<emphasis id="B951EABA170F81714B73FE69FB207AEF" bold="true" box="[1067,1152,457,481]" italics="true" pageId="7" pageNumber="8">Thorax:</emphasis>
Pronotum convex medially. Entire pronotal disc covered with dense and rugose punctuation (punctures separated by their diameter). Basal margin rounded, slightly angularly projecting posteriorly. Pronotum bi-margined laterally, strongest lateral carina aligned with marginal elytral carina; outer marginal pronotal stria similarly carinate, narrowing and descending below inner marginal in anterior half, overlapped and invisible from above in the anterior third. Scutellar shield obscured. Elytra with the same punctuation pattern as pronotal disc. Elytral disc slightly convex, with two weakly elevated longitudinal carinae located in medial part. Elytra carinate along the outer margin and truncate posteriorly. Sutural area elevated in posterior third forming a keel. Marginal elytral stria carinate, deeply impressed. Elytral epipleuron with two carinate striae. Prosternal lobe very short and broad with dense punctuation, with incised furrows laterally for passage of the antennal funicle; Prosternal keel broad, similarly punctate to prosternal lobe, with lateral carinae weakly converging anterad, abruptly narrowing at junction with prosternal lobe; basal prosternal margin weakly and broadly emarginate. No striation visible on prosternum. Antennal cavities widely open, located posterolaterally on prosternal sides. Mesoventrite wide, densely and coarsely punctuated. Anterior margin outwardly curved along prosternal margin. Metaventrite with a pair of lateral metaventral striae present in anterior two-thirds, extending laterally to near metepisternum. Postmesocoxal striae curved along mesocoxae. Punctures on metaventrite become sparser and larger posteriorly.—
<emphasis id="B951EABA170F81714B74FAB7FBC47E20" bold="true" box="[1068,1124,1303,1326]" italics="true" pageId="7" pageNumber="8">Legs:</emphasis>
Profemur wide and flat, with a groove for protibial reception. Protibia curved with a row of short setae along the outer margin. Protibial margins slightly thickened, creating a shallow protibial groove just mediad the outer margin. Protibial spur absent. Mesofemur narrow, mesotibia very slender, with a row of short setae along inner and outer margin. Hind leg identical to middle leg. Tarsi 5-5- 5, with ultimate tarsomere similar in length to the combined tarsomeres 2–4; tarsomeres 2–4 bearing a single pair of apical setae ventrally.—
<emphasis id="B951EABA170F81714B54F990FBDA7D46" bold="true" box="[1036,1146,1584,1608]" italics="true" pageId="7">Abdomen:</emphasis>
<emphasis id="B951EABA170F81714B54F990FBDA7D46" bold="true" box="[1036,1146,1584,1608]" italics="true" pageId="7" pageNumber="8">Abdomen:</emphasis>
All abdominal ventrites with dense punctuation (but finer than on thoracical ventrites). First ventrite with a pair of parallel lateral striae and a pair of curved postmetacoxal striae. Propygidium and pygidium coarsely and densely punctured.
</paragraph>
<paragraph id="8B9A36A8170F817E4C72F944FE707C27" blockId="7.[810,1459,1763,1975]" lastBlockId="8.[129,778,1778,1833]" lastPageId="8" pageId="7">
<emphasis id="B951EABA170F81714C72F944FBB37DF5" box="[810,1043,1764,1787]" italics="true" pageId="7">Taxonomic assignment:</emphasis>
</subSubSection>
<subSubSection id="C33F6523170F817E4C72F944FE707C27" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="discussion">
<paragraph id="8B9A36A8170F817E4C72F944FE707C27" blockId="7.[810,1459,1763,1975]" lastBlockId="8.[129,778,1778,1833]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<emphasis id="B951EABA170F81714C72F944FBB37DF5" box="[810,1043,1764,1787]" italics="true" pageId="7" pageNumber="8">Taxonomic assignment:</emphasis>
We place this species within the tribe
<taxonomicName id="4C254D2B170F81714C72F8A2FC2D7C14" authorityName="Olexa" authorityYear="1982" box="[810,909,1794,1818]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="tribe" tribe="Anapleini">Anapleini</taxonomicName>
<taxonomicName id="4C254D2B170F81714C72F8A2FC2D7C14" authorityName="Olexa" authorityYear="1982" box="[810,909,1794,1818]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="tribe" tribe="Anapleini">Anapleini</taxonomicName>
based on the morphology of the protibia, as defined by
<bibRefCitation id="EFB44B59170F81714C12F881FC787C34" author="Olexa A" box="[842,984,1825,1850]" pageId="7" pagination="37 - 45" refId="ref18674" refString="Olexa A. Revision der palaarktischen arten der gauung Anapleus (Coleoptera, Histeridae). Acta Entomologica Bohemoslovaca 1982; 79: 37 - 45." type="journal article" year="1982">Olexa (1982)</bibRefCitation>
<bibRefCitation id="EFB44B59170F81714C12F881FC787C34" author="Olexa A" box="[842,984,1825,1850]" pageId="7" pageNumber="8" pagination="37 - 45" refId="ref18674" refString="Olexa A. Revision der palaarktischen arten der gauung Anapleus (Coleoptera, Histeridae). Acta Entomologica Bohemoslovaca 1982; 79: 37 - 45." type="journal article" year="1982">Olexa (1982)</bibRefCitation>
: tarsal groove of protibia formed by marginal thickening, row of setae along the outer margin (
<figureCitation id="131E2A2D170F81714A09F8E1FA027C57" box="[1361,1442,1857,1881]" captionStart="Figure 6" captionStartId="8.[129,194,1677,1701]" captionTargetBox="[129,1473,144,1650]" captionTargetId="figure-114@8.[129,1473,144,1650]" captionTargetPageId="8" captionText="Figure 6. Anapleus punctulatus sp.nov.. A, dorsal view.B, elytral epipleuron. C, ventral view.D, frontal view.Abbreviations:ees, elytral epipleural striae; lps, lateral pronotal stria; mes, marginal elytral stria; mps, marginal prosternal stria; ms, metaventral stria." pageId="7">Fig. 6D</figureCitation>
<figureCitation id="131E2A2D170F81714A09F8E1FA027C57" box="[1361,1442,1857,1881]" captionStart="Figure 6" captionStartId="8.[129,194,1677,1701]" captionTargetBox="[129,1473,144,1650]" captionTargetId="figure-114@8.[129,1473,144,1650]" captionTargetPageId="8" captionText="Figure 6. Anapleus punctulatus sp.nov.. A, dorsal view.B, elytral epipleuron. C, ventral view.D, frontal view.Abbreviations:ees, elytral epipleural striae; lps, lateral pronotal stria; mes, marginal elytral stria; mps, marginal prosternal stria; ms, metaventral stria." pageId="7" pageNumber="8">Fig. 6D</figureCitation>
). Other characters typical for
<taxonomicName id="4C254D2B170F81714B0EF8C0FB197C76" authorityName="Olexa" authorityYear="1982" box="[1110,1209,1888,1912]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="tribe" tribe="Anapleini">Anapleini</taxonomicName>
<taxonomicName id="4C254D2B170F81714B0EF8C0FB197C76" authorityName="Olexa" authorityYear="1982" box="[1110,1209,1888,1912]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="tribe" tribe="Anapleini">Anapleini</taxonomicName>
include weakly defined and widely open antennal cavities and two elytral epipleural striae (
<figureCitation id="131E2A2D170F81714C2DF83FFC607CB9" box="[885,960,1951,1975]" captionStart="Figure 6" captionStartId="8.[129,194,1677,1701]" captionTargetBox="[129,1473,144,1650]" captionTargetId="figure-114@8.[129,1473,144,1650]" captionTargetPageId="8" captionText="Figure 6. Anapleus punctulatus sp.nov.. A, dorsal view.B, elytral epipleuron. C, ventral view.D, frontal view.Abbreviations:ees, elytral epipleural striae; lps, lateral pronotal stria; mes, marginal elytral stria; mps, marginal prosternal stria; ms, metaventral stria." pageId="7">Fig. 6B</figureCitation>
). Ŋis species is furthermore assignable to the genus
<taxonomicName id="4C254D2B1700817E4F9FF952FE087C04" authority="Horn, 1873" authorityName="G. H. Horn" authorityYear="1873" box="[199,424,1778,1802]" class="Insecta" family="Histeridae" genus="Anapleus" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA1700817E4F9FF952FE817C04" box="[199,289,1778,1802]" italics="true" pageId="8">Anapleus</emphasis>
<figureCitation id="131E2A2D170F81714C2DF83FFC607CB9" box="[885,960,1951,1975]" captionStart="Figure 6" captionStartId="8.[129,194,1677,1701]" captionTargetBox="[129,1473,144,1650]" captionTargetId="figure-114@8.[129,1473,144,1650]" captionTargetPageId="8" captionText="Figure 6. Anapleus punctulatus sp.nov.. A, dorsal view.B, elytral epipleuron. C, ventral view.D, frontal view.Abbreviations:ees, elytral epipleural striae; lps, lateral pronotal stria; mes, marginal elytral stria; mps, marginal prosternal stria; ms, metaventral stria." pageId="7" pageNumber="8">Fig. 6B</figureCitation>
). This species is furthermore assignable to the genus
<taxonomicName id="4C254D2B1700817E4F9FF952FE087C04" authority="Horn, 1873" authorityName="G. H. Horn" authorityYear="1873" box="[199,424,1778,1802]" class="Insecta" family="Histeridae" genus="Anapleus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA1700817E4F9FF952FE817C04" box="[199,289,1778,1802]" italics="true" pageId="8" pageNumber="9">Anapleus</emphasis>
Horn, 1873
</taxonomicName>
with which it shares the simple frontoclypeal area without striae.
</paragraph>
<caption id="DF5A66201700817E4FD9F92DFB5A7DCC" pageId="8" startId="8.[129,194,1677,1701]" targetBox="[129,1473,144,1650]" targetPageId="8" targetType="figure">
<paragraph id="8B9A36A81700817E4FD9F92DFB5A7DCC" blockId="8.[129,1383,1677,1730]" pageId="8">
<emphasis id="B951EABA1700817E4FD9F92DFF787DA8" bold="true" box="[129,216,1677,1702]" pageId="8">Figure 6.</emphasis>
<taxonomicName id="4C254D2B1700817E4F85F92EFE3F7DA8" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[221,415,1678,1702]" class="Insecta" family="Histeridae" genus="Anapleus" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="punctulatus" status="sp. nov.">
<emphasis id="B951EABA1700817E4F85F92EFE3F7DA8" box="[221,415,1678,1702]" italics="true" pageId="8">Anapleus punctulatus</emphasis>
</subSubSection>
<caption id="DF5A66201700817E4FD9F92DFB5A7DCC" pageId="8" pageNumber="9" startId="8.[129,194,1677,1701]" targetBox="[129,1473,144,1650]" targetPageId="8" targetType="figure">
<paragraph id="8B9A36A81700817E4FD9F92DFB5A7DCC" blockId="8.[129,1383,1677,1730]" pageId="8" pageNumber="9">
<emphasis id="B951EABA1700817E4FD9F92DFF787DA8" bold="true" box="[129,216,1677,1702]" pageId="8" pageNumber="9">Figure 6.</emphasis>
<taxonomicName id="4C254D2B1700817E4F85F92EFE3F7DA8" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[221,415,1678,1702]" class="Insecta" family="Histeridae" genus="Anapleus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="punctulatus" status="sp. nov.">
<emphasis id="B951EABA1700817E4F85F92EFE3F7DA8" box="[221,415,1678,1702]" italics="true" pageId="8" pageNumber="9">Anapleus punctulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C11700817E4EFBF92EFE447DAB" box="[419,484,1678,1701]" pageId="8" rank="species">sp. nov</taxonomicNameLabel>
<taxonomicNameLabel id="A26257C11700817E4EFBF92EFE447DAB" box="[419,484,1678,1701]" pageId="8" pageNumber="9" rank="species">sp. nov</taxonomicNameLabel>
.. A, dorsal view. B, elytral epipleuron. C, ventral view. D, frontal view. Abbreviations: ees, elytral epipleural striae; lps, lateral pronotal stria; mes, marginal elytral stria; mps, marginal prosternal stria; ms, metaventral stria.
</paragraph>
</caption>
<paragraph id="8B9A36A81700817F4FD9F8F0FE0C7BC9" blockId="8.[129,778,1872,1959]" lastBlockId="9.[113,762,144,200]" lastPageId="9" pageId="8">
<emphasis id="B951EABA1700817E4FD9F8F0FEF87C66" box="[129,344,1872,1896]" italics="true" pageId="8">Differential diagnosis:</emphasis>
<subSubSection id="C33F65231700817F4FD9F8F0FE0C7BC9" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="8B9A36A81700817F4FD9F8F0FE0C7BC9" blockId="8.[129,778,1872,1959]" lastBlockId="9.[113,762,144,200]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<emphasis id="B951EABA1700817E4FD9F8F0FEF87C66" box="[129,344,1872,1896]" italics="true" pageId="8" pageNumber="9">Differential diagnosis:</emphasis>
Two species of
<taxonomicName id="4C254D2B1700817E4D51F8F0FDCC7C66" authorityName="Olexa" authorityYear="1982" box="[521,620,1872,1896]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="tribe" tribe="Anapleini">Anapleini</taxonomicName>
<taxonomicName id="4C254D2B1700817E4D51F8F0FDCC7C66" authorityName="Olexa" authorityYear="1982" box="[521,620,1872,1896]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="tribe" tribe="Anapleini">Anapleini</taxonomicName>
have been described from the
<collectingRegion id="49E1F84A1700817E4E75F8CFFED77C89" box="[301,375,1903,1927]" country="Myanmar" name="Kachin" pageId="8">Kachin</collectingRegion>
<collectingRegion id="49E1F84A1700817E4E75F8CFFED77C89" box="[301,375,1903,1927]" country="Myanmar" name="Kachin" pageId="8" pageNumber="9">Kachin</collectingRegion>
amber so far:
<taxonomicName id="4C254D2B1700817E4D5DF8CFFF527CA9" authority="Jiang et al., 2022" authorityName="Jiang" authorityYear="2022" class="Insecta" family="Histeridae" genus="Anapleus" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="kachinensis">
<emphasis id="B951EABA1700817E4D5DF8CFFD727C89" box="[517,722,1903,1927]" italics="true" pageId="8">Anapleus kachinensis</emphasis>
<bibRefCitation id="EFB44B591700817E4D8EF8CFFF527CA9" author="Jiang R &amp; Chen X" pageId="8" refId="ref17908" refString="Jiang R, Caterino Mº, Chen X. Discovery of the genus Anapleus Horn, 1873 from Cretaceous Kachin Amber (Coleoptera: Histeridae). Insects 2022; 13: 746. hups: // doi. org / 10.3390 / insects 13080746" type="journal volume" year="2022">
<taxonomicName id="4C254D2B1700817E4D5DF8CFFF527CA9" authority="Jiang et al., 2022" authorityName="Jiang" authorityYear="2022" class="Insecta" family="Histeridae" genus="Anapleus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="kachinensis">
<emphasis id="B951EABA1700817E4D5DF8CFFD727C89" box="[517,722,1903,1927]" italics="true" pageId="8" pageNumber="9">Anapleus kachinensis</emphasis>
<bibRefCitation id="EFB44B591700817E4D8EF8CFFF527CA9" author="Jiang R &amp; Chen X" pageId="8" pageNumber="9" refId="ref17908" refString="Jiang R, Caterino Mº, Chen X. Discovery of the genus Anapleus Horn, 1873 from Cretaceous Kachin Amber (Coleoptera: Histeridae). Insects 2022; 13: 746. hups: // doi. org / 10.3390 / insects 13080746" type="journal volume" year="2022">
Jiang
<emphasis id="B951EABA1700817E4FD9F830FF0B7CA9" box="[129,171,1935,1959]" italics="true" pageId="8">et al</emphasis>
<emphasis id="B951EABA1700817E4FD9F830FF0B7CA9" box="[129,171,1935,1959]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
., 2022
</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C254D2B1700817E4E7DF82FFE467CA9" authorityName="Simon Prazak &amp; Lackner" authorityYear="2023" box="[293,486,1935,1959]" class="Insecta" family="Histeridae" genus="Cretanapleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="seideli">
<emphasis id="B951EABA1700817E4E7DF82FFE467CA9" box="[293,486,1935,1959]" italics="true" pageId="8">Cretanapleus seideli</emphasis>
<taxonomicName id="4C254D2B1700817E4E7DF82FFE467CA9" authorityName="Simon Prazak &amp; Lackner" authorityYear="2023" box="[293,486,1935,1959]" class="Insecta" family="Histeridae" genus="Cretanapleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="seideli">
<emphasis id="B951EABA1700817E4E7DF82FFE467CA9" box="[293,486,1935,1959]" italics="true" pageId="8" pageNumber="9">Cretanapleus seideli</emphasis>
</taxonomicName>
ºimon-Pražák and
<bibRefCitation id="EFB44B591700817E4DE9F82FFCCF7C04" author="Lackner T &amp; Zhang YM &amp; Kindler C" box="[689,879,1778,1959]" pageId="8" pagination="48 - 71" refId="ref18368" refString="Lackner T, Zhang YM, Kindler C et al. ºaprininae: phylogeny, biogeography and a new classification of the subfamily (Coleoptera: Histeridae). Systematic Entomology 2023; 49: 48 - 71. hups: // doi. org / 10.1111 / syen. 12606" type="journal article" year="2023">Lackner, 2023</bibRefCitation>
Simon-Pražák and
<bibRefCitation id="EFB44B591700817E4DE9F82FFCCF7C04" author="Lackner T &amp; Zhang YM &amp; Kindler C" box="[689,879,1778,1959]" pageId="8" pageNumber="9" pagination="48 - 71" refId="ref18368" refString="Lackner T, Zhang YM, Kindler C et al. ºaprininae: phylogeny, biogeography and a new classification of the subfamily (Coleoptera: Histeridae). Systematic Entomology 2023; 49: 48 - 71. hups: // doi. org / 10.1111 / syen. 12606" type="journal article" year="2023">Lackner, 2023</bibRefCitation>
.
<taxonomicName id="4C254D2B1700817E4C26F952FBE47C04" authorityName="Simon Prazak &amp; Lackner" authorityYear="2023" box="[894,1092,1778,1802]" class="Insecta" family="Histeridae" genus="Cretanapleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="seideli">
<emphasis id="B951EABA1700817E4C26F952FBE47C04" box="[894,1092,1778,1802]" italics="true" pageId="8">Cretanapleus seideli</emphasis>
<taxonomicName id="4C254D2B1700817E4C26F952FBE47C04" authorityName="Simon Prazak &amp; Lackner" authorityYear="2023" box="[894,1092,1778,1802]" class="Insecta" family="Histeridae" genus="Cretanapleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="seideli">
<emphasis id="B951EABA1700817E4C26F952FBE47C04" box="[894,1092,1778,1802]" italics="true" pageId="8" pageNumber="9">Cretanapleus seideli</emphasis>
</taxonomicName>
can be easily distinguished by the presence of a strongly carinate frontal stria, as well as dorsally explanate body margin.
<taxonomicName id="4C254D2B1700817E4B76F890FAA07C46" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[1070,1280,1840,1864]" class="Insecta" family="Histeridae" genus="Anapleus" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="punctulatus" status="sp. nov.">
<emphasis id="B951EABA1700817E4B76F890FAA07C46" box="[1070,1280,1840,1864]" italics="true" pageId="8">Anapleus punctulatus</emphasis>
<taxonomicName id="4C254D2B1700817E4B76F890FAA07C46" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[1070,1280,1840,1864]" class="Insecta" family="Histeridae" genus="Anapleus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="punctulatus" status="sp. nov.">
<emphasis id="B951EABA1700817E4B76F890FAA07C46" box="[1070,1280,1840,1864]" italics="true" pageId="8" pageNumber="9">Anapleus punctulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C11700817E4A5DF891FAF27C46" box="[1285,1362,1841,1864]" pageId="8" rank="species">sp. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="A26257C11700817E4A5DF891FAF27C46" box="[1285,1362,1841,1864]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
can be differentiated from
<taxonomicName id="4C254D2B1700817E4CB3F8EFFB1C7C69" box="[1003,1212,1871,1895]" class="Insecta" family="Histeridae" genus="Anapleus" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="kachinensis">
<emphasis id="B951EABA1700817E4CB3F8EFFB1C7C69" box="[1003,1212,1871,1895]" italics="true" pageId="8">Anapleus kachinensis</emphasis>
<taxonomicName id="4C254D2B1700817E4CB3F8EFFB1C7C69" box="[1003,1212,1871,1895]" class="Insecta" family="Histeridae" genus="Anapleus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="kachinensis">
<emphasis id="B951EABA1700817E4CB3F8EFFB1C7C69" box="[1003,1212,1871,1895]" italics="true" pageId="8" pageNumber="9">Anapleus kachinensis</emphasis>
</taxonomicName>
by the more rugose and dense body punctuation, by the presence of carinate marginal pronotal and marginal elytral striae (
<figureCitation id="131E2A2D1700817E4B91F82EFAB77CA8" box="[1225,1303,1934,1958]" captionStart="Figure 6" captionStartId="8.[129,194,1677,1701]" captionTargetBox="[129,1473,144,1650]" captionTargetId="figure-114@8.[129,1473,144,1650]" captionTargetPageId="8" captionText="Figure 6. Anapleus punctulatus sp.nov.. A, dorsal view.B, elytral epipleuron. C, ventral view.D, frontal view.Abbreviations:ees, elytral epipleural striae; lps, lateral pronotal stria; mes, marginal elytral stria; mps, marginal prosternal stria; ms, metaventral stria." pageId="8">Fig. 6A</figureCitation>
<figureCitation id="131E2A2D1700817E4B91F82EFAB77CA8" box="[1225,1303,1934,1958]" captionStart="Figure 6" captionStartId="8.[129,194,1677,1701]" captionTargetBox="[129,1473,144,1650]" captionTargetId="figure-114@8.[129,1473,144,1650]" captionTargetPageId="8" captionText="Figure 6. Anapleus punctulatus sp.nov.. A, dorsal view.B, elytral epipleuron. C, ventral view.D, frontal view.Abbreviations:ees, elytral epipleural striae; lps, lateral pronotal stria; mes, marginal elytral stria; mps, marginal prosternal stria; ms, metaventral stria." pageId="8" pageNumber="9">Fig. 6A</figureCitation>
), presence of a distinct lateral metaventral stria (
<figureCitation id="131E2A2D1701817F4E9FFF30FDB07BA6" box="[455,528,144,168]" captionStart="Figure 6" captionStartId="8.[129,194,1677,1701]" captionTargetBox="[129,1473,144,1650]" captionTargetId="figure-114@8.[129,1473,144,1650]" captionTargetPageId="8" captionText="Figure 6. Anapleus punctulatus sp.nov.. A, dorsal view.B, elytral epipleuron. C, ventral view.D, frontal view.Abbreviations:ees, elytral epipleural striae; lps, lateral pronotal stria; mes, marginal elytral stria; mps, marginal prosternal stria; ms, metaventral stria." pageId="9">Fig. 6B</figureCitation>
<figureCitation id="131E2A2D1701817F4E9FFF30FDB07BA6" box="[455,528,144,168]" captionStart="Figure 6" captionStartId="8.[129,194,1677,1701]" captionTargetBox="[129,1473,144,1650]" captionTargetId="figure-114@8.[129,1473,144,1650]" captionTargetPageId="8" captionText="Figure 6. Anapleus punctulatus sp.nov.. A, dorsal view.B, elytral epipleuron. C, ventral view.D, frontal view.Abbreviations:ees, elytral epipleural striae; lps, lateral pronotal stria; mes, marginal elytral stria; mps, marginal prosternal stria; ms, metaventral stria." pageId="9" pageNumber="10">Fig. 6B</figureCitation>
), and the absence of a strong protibial spur (
<figureCitation id="131E2A2D1701817F4E09FF0FFE3B7BC9" box="[337,411,175,199]" captionStart="Figure 6" captionStartId="8.[129,194,1677,1701]" captionTargetBox="[129,1473,144,1650]" captionTargetId="figure-114@8.[129,1473,144,1650]" captionTargetPageId="8" captionText="Figure 6. Anapleus punctulatus sp.nov.. A, dorsal view.B, elytral epipleuron. C, ventral view.D, frontal view.Abbreviations:ees, elytral epipleural striae; lps, lateral pronotal stria; mes, marginal elytral stria; mps, marginal prosternal stria; ms, metaventral stria." pageId="9">Fig. 6D</figureCitation>
<figureCitation id="131E2A2D1701817F4E09FF0FFE3B7BC9" box="[337,411,175,199]" captionStart="Figure 6" captionStartId="8.[129,194,1677,1701]" captionTargetBox="[129,1473,144,1650]" captionTargetId="figure-114@8.[129,1473,144,1650]" captionTargetPageId="8" captionText="Figure 6. Anapleus punctulatus sp.nov.. A, dorsal view.B, elytral epipleuron. C, ventral view.D, frontal view.Abbreviations:ees, elytral epipleural striae; lps, lateral pronotal stria; mes, marginal elytral stria; mps, marginal prosternal stria; ms, metaventral stria." pageId="9" pageNumber="10">Fig. 6D</figureCitation>
).
</paragraph>
<paragraph id="8B9A36A81701817F4F29FF49FD277A0F" blockId="9.[113,647,233,257]" box="[113,647,233,257]" pageId="9">
<emphasis id="B951EABA1701817F4F29FF49FF417A0F" box="[113,225,233,257]" italics="true" pageId="9">Etymology:</emphasis>
Named asser its coarsely punctate cuticle.
</subSubSection>
<subSubSection id="C33F65231701817F4F29FF49FD277A0F" box="[113,647,233,257]" pageId="9" pageNumber="10" type="etymology">
<paragraph id="8B9A36A81701817F4F29FF49FD277A0F" blockId="9.[113,647,233,257]" box="[113,647,233,257]" pageId="9" pageNumber="10">
<emphasis id="B951EABA1701817F4F29FF49FF417A0F" box="[113,225,233,257]" italics="true" pageId="9" pageNumber="10">Etymology:</emphasis>
Named after its coarsely punctate cuticle.
</paragraph>
</subSubSection>
</treatment>

283
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<subSubSection id="C33F6523171981674C86FDE6FB5F796F" box="[990,1279,582,609]" pageId="17" type="nomenclature">
<paragraph id="8B9A36A8171981674C86FDE6FB5F796F" blockId="17.[990,1279,582,609]" box="[990,1279,582,609]" pageId="17">
<heading id="D0D281C4171981674C86FDE6FB5F796F" bold="true" box="[990,1279,582,609]" centered="true" fontSize="11" level="2" pageId="17" reason="2">
<emphasis id="B951EABA171981674C86FDE6FB5F796F" bold="true" box="[990,1279,582,609]" pageId="17">
<taxonomicName id="4C254D2B171981674C86FDE6FB03796F" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[990,1187,582,609]" class="Insecta" family="Histeridae" genus="Placatister" kingdom="Animalia" order="Coleoptera" pageId="17" phylum="Arthropoda" rank="species" species="cascus" status="sp. nov.">
<emphasis id="B951EABA171981674C86FDE6FB03796F" bold="true" box="[990,1187,582,609]" italics="true" pageId="17">Placatister cascus</emphasis>
<treatment id="038C87BE171981654C86FDE6FD4C7C92" LSID="urn:lsid:plazi:treatment:038C87BE171981654C86FDE6FD4C7C92" httpUri="http://treatment.plazi.org/id/038C87BE171981654C86FDE6FD4C7C92" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18">
<subSubSection id="C33F6523171981674C86FDE6FB5F796F" box="[990,1279,582,609]" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="8B9A36A8171981674C86FDE6FB5F796F" blockId="17.[990,1279,582,609]" box="[990,1279,582,609]" pageId="17" pageNumber="18">
<heading id="D0D281C4171981674C86FDE6FB5F796F" bold="true" box="[990,1279,582,609]" centered="true" fontSize="11" level="2" pageId="17" pageNumber="18" reason="2">
<emphasis id="B951EABA171981674C86FDE6FB5F796F" bold="true" box="[990,1279,582,609]" pageId="17" pageNumber="18">
<taxonomicName id="4C254D2B171981674C86FDE6FB03796F" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[990,1187,582,609]" class="Insecta" family="Histeridae" genus="Placatister" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="cascus" status="sp. nov.">
<emphasis id="B951EABA171981674C86FDE6FB03796F" bold="true" box="[990,1187,582,609]" italics="true" pageId="17" pageNumber="18">Placatister cascus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C1171981674BF0FDE7FB5F796F" box="[1192,1279,583,609]" pageId="17" rank="species">sp. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="A26257C1171981674BF0FDE7FB5F796F" box="[1192,1279,583,609]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33F6523171981654B18FDD5FD4C7C92" lastPageId="19" pageId="17" type="description">
<paragraph id="8B9A36A8171981674B18FDD5FB3D7981" blockId="17.[1088,1181,629,656]" box="[1088,1181,629,656]" pageId="17">
<emphasis id="B951EABA171981674B18FDD5FB3D7981" bold="true" box="[1088,1181,629,656]" pageId="17">
<subSubSection id="C33F6523171981674B18FDD5FB3D7981" box="[1088,1181,629,656]" pageId="17" pageNumber="18" type="description">
<paragraph id="8B9A36A8171981674B18FDD5FB3D7981" blockId="17.[1088,1181,629,656]" box="[1088,1181,629,656]" pageId="17" pageNumber="18">
<emphasis id="B951EABA171981674B18FDD5FB3D7981" bold="true" box="[1088,1181,629,656]" pageId="17" pageNumber="18">
(
<figureCitation id="131E2A2D171981674B13FDD6FB32799E" box="[1099,1170,630,656]" captionStart="Figure 12" captionStartId="18.[129,194,1836,1860]" captionTargetBox="[175,1428,144,1808]" captionTargetId="figure-8@18.[175,1428,144,1808]" captionTargetPageId="18" captionText="Figure 12. Placatister cascus gen.et sp. nov.. A, dorsal view.B, ventral view. C, lateral view.D, antennal club. E, mouthparts. F, frontal view.G, prosternal keel.H, antenna.Abbreviations:ees, elytral epipleural stria; fs, frontal stria; pf, prosternal foveae." pageId="17">Fig. 12</figureCitation>
<figureCitation id="131E2A2D171981674B13FDD6FB32799E" box="[1099,1170,630,656]" captionStart="Figure 12" captionStartId="18.[129,194,1836,1860]" captionTargetBox="[175,1428,144,1808]" captionTargetId="figure-8@18.[175,1428,144,1808]" captionTargetPageId="18" captionText="Figure 12. Placatister cascus gen.et sp. nov.. A, dorsal view.B, ventral view. C, lateral view.D, antennal club. E, mouthparts. F, frontal view.G, prosternal keel.H, antenna.Abbreviations:ees, elytral epipleural stria; fs, frontal stria; pf, prosternal foveae." pageId="17" pageNumber="18">Fig. 12</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8B9A36A8171981674C72FD16FB477803" blockId="17.[810,1460,694,781]" pageId="17">
<emphasis id="B951EABA171981674C72FD16FC2679C0" box="[810,902,694,718]" italics="true" pageId="17">Material:</emphasis>
<collectingRegion id="49E1F84A171981674CD5FD16FC7779C0" box="[909,983,694,718]" country="Myanmar" name="Kachin" pageId="17">Kachin</collectingRegion>
</subSubSection>
<subSubSection id="C33F6523171981674C72FD16FB477803" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="8B9A36A8171981674C72FD16FB477803" blockId="17.[810,1460,694,781]" pageId="17" pageNumber="18">
<emphasis id="B951EABA171981674C72FD16FC2679C0" box="[810,902,694,718]" italics="true" pageId="17" pageNumber="18">Material:</emphasis>
<materialsCitation id="3B4D3CF5171981674CD5FD16FB437803" collectionCode="MEBHK" location="Kachin" pageId="17" pageNumber="18" specimenCode="P1425/E" specimenCount="1" stateProvince="Kachin">
<collectingRegion id="49E1F84A171981674CD5FD16FC7779C0" box="[909,983,694,718]" country="Myanmar" name="Kachin" pageId="17" pageNumber="18">Kachin</collectingRegion>
amber (
<emphasis id="B951EABA171981674B69FD17FB9A79C0" box="[1073,1082,695,718]" italics="true" pageId="17">c</emphasis>
<emphasis id="B951EABA171981674B69FD17FB9A79C0" box="[1073,1082,695,718]" italics="true" pageId="17" pageNumber="18">c</emphasis>
. 99 Mya). Amber piece dimensions: 9 × 5 ×
<quantity id="4CDD9B4D171981674C25FD76FC1879E3" box="[893,952,726,750]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="17" unit="mm" value="1.0">1 mm</quantity>
. Amber clear with several insect fragments and air bubbles. Deposited at MEBHK (P1425/E).
<quantity id="4CDD9B4D171981674C25FD76FC1879E3" box="[893,952,726,750]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="18" unit="mm" value="1.0">1 mm</quantity>
. Amber clear with several insect fragments and air bubbles. Deposited at
<collectionCode id="ED34AE6D171981674B54FD55FBC87802" box="[1036,1128,757,780]" pageId="17" pageNumber="18">MEBHK</collectionCode>
(
<specimenCode id="DB839ED3171981674B2EFD55FB7B7803" box="[1142,1243,757,781]" pageId="17" pageNumber="18">P1425/E</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B9A36A8171981654C72FC8FFF097876" blockId="17.[809,1460,814,1966]" lastBlockId="19.[113,763,144,888]" lastPageId="19" pageId="17">
<emphasis id="B951EABA171981674C72FC8FFC1C7848" box="[810,956,815,838]" italics="true" pageId="17">Measurements:</emphasis>
</subSubSection>
<subSubSection id="C33F6523171981654C72FC8FFF097876" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18" type="description">
<paragraph id="8B9A36A8171981654C72FC8FFF097876" blockId="17.[809,1460,814,1966]" lastBlockId="19.[113,763,144,888]" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18">
<emphasis id="B951EABA171981674C72FC8FFC1C7848" box="[810,956,815,838]" italics="true" pageId="17" pageNumber="18">Measurements:</emphasis>
Head width:
<quantity id="4CDD9B4D171981674B0AFC8EFB0D7848" box="[1106,1197,814,838]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="17" unit="mm" value="0.36">0.36 mm</quantity>
<quantity id="4CDD9B4D171981674B0AFC8EFB0D7848" box="[1106,1197,814,838]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="17" pageNumber="18" unit="mm" value="0.36">0.36 mm</quantity>
; width between anterior pronotal angles:
<quantity id="4CDD9B4D171981674C86FCEDFB99786B" box="[990,1081,845,869]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" pageId="17" unit="mm" value="0.56">0.56 mm</quantity>
<quantity id="4CDD9B4D171981674C86FCEDFB99786B" box="[990,1081,845,869]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" pageId="17" pageNumber="18" unit="mm" value="0.56">0.56 mm</quantity>
; width between posterior pronotal angles:
<quantity id="4CDD9B4D171981674C27FCCDFC7A788A" box="[895,986,877,901]" metricMagnitude="-4" metricUnit="m" metricValue="9.2" pageId="17" unit="mm" value="0.92">0.92 mm</quantity>
<quantity id="4CDD9B4D171981674C27FCCDFC7A788A" box="[895,986,877,901]" metricMagnitude="-4" metricUnit="m" metricValue="9.2" pageId="17" pageNumber="18" unit="mm" value="0.92">0.92 mm</quantity>
; pronotal length:
<quantity id="4CDD9B4D171981674BF5FCCDFAA8788A" box="[1197,1288,877,901]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="17" unit="mm" value="0.58">0.58 mm</quantity>
<quantity id="4CDD9B4D171981674BF5FCCDFAA8788A" box="[1197,1288,877,901]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="17" pageNumber="18" unit="mm" value="0.58">0.58 mm</quantity>
; sutural elytral length:
<quantity id="4CDD9B4D171981674C26FC2CFC6B78AA" box="[894,971,908,932]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="17" unit="mm" value="0.8">0.8 mm</quantity>
<quantity id="4CDD9B4D171981674C26FC2CFC6B78AA" box="[894,971,908,932]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="17" pageNumber="18" unit="mm" value="0.8">0.8 mm</quantity>
; elytral width (in pair, across widest point):
<quantity id="4CDD9B4D171981674C72FC0CFCDA78CD" box="[810,890,940,964]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="17" unit="mm" value="1.2">1.2 mm</quantity>
<quantity id="4CDD9B4D171981674C72FC0CFCDA78CD" box="[810,890,940,964]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="17" pageNumber="18" unit="mm" value="1.2">1.2 mm</quantity>
.—
<emphasis id="B951EABA171981674CC1FC0BFC7978CD" bold="true" box="[921,985,939,963]" italics="true" pageId="17">Body:</emphasis>
Round, strongly dorsoventrally flaưened (probably partially compressed). Cuticle regularly punctuate. Colour reddish brown to black.—
<emphasis id="B951EABA171981674BCCFC4AFB757F0C" bold="true" box="[1172,1237,1002,1026]" italics="true" pageId="17">Head:</emphasis>
Frons flat, wide and punctuate, strongly, squarely margined along inner edge of each eye, with a fine stria along their inner edges continuous with a complete, transverse frontal stria. Frontoclypeal suture not visible. Epistoma constricted, medially ridged, sides receding below broad frontal shelf. Labrum trapezoidal (width: length ratio 5: 1), emarginate anteriorly, with a pair of short setae at anterolateral corners, lacking punctuation. Mandibles wide and thin, basally with weakly dentate incisor edge, and a long, acute tip. Terminal maxillary palpomere wide and round; other ventral mouthparts damaged and/or missing. Antenna inserted under frontal shelf, strongly dorsoventrally flaưened, possibly partially compressed. ºcape short, widened, and curved to apex. Pedicel wide, subcylindrical, as long as antennomeres 3–5 combined. Antennomeres 3–7 similar in shape and size, short, wider than long (width: length ratio 3: 1). Antennomere 8 adjacent to the antennal club, 2.5× wider than antennomere 7. Antennomeres 9–11 form a wide, flat, and round antennal club with well-defined sutures.—
<emphasis id="B951EABA171981674CCCF9BEFC497D38" bold="true" box="[916,1001,1566,1590]" italics="true" pageId="17">Ŋorax:</emphasis>
Pronotal disc flat. Anterior margin widely emarginate, with fine marginal pronotal stria; not evident on lateral or basal margins. Basal margin broadly rounded, almost angularly projecting. ºurface of pronotal disc densely punctuate, punctures separated by 1–2× their diameter. ºcutellar shield not visible, obscured by bubbles. Elytra truncate, without dorsal striation, epipleural stria complete; elytral disc appears weakly depressed in humeral corners (possibly due to damage). Elytral punctuation with the same paưern as in pronotum. Prosternal lobe rather short, subtruncate anteriorly, set off from keel by pair of deep, setose foveae, weakly conjoined at their bases; prosternal keel flat, short, abruptly descending at front into two large approximate prosternal foveae, with fine longitudinal striae; finely punctate between; sides of prosternum widely incised for passage of antennal funicle, with deep channel reaching middle of base of prosternal keel; sides of prosternal keel appear concave. Mesoventrite wide and short, obscured by bubbles, probably finely punctuated. Meso-metaventral suture visible as a fine line. Metaventrite punctuate, with lateral metaventral stria directed posterolaterally to hind coxae. Other striation not visible. Metepisternum broadly exposed, rectangular.—
<emphasis id="B951EABA171B81654D9AFECCFD5A7A8D" bold="true" box="[706,762,364,387]" italics="true" pageId="19">Legs:</emphasis>
Profemur flaưed, slightly widened. Protibia narrow and smooth, without dentation, few longer setae present along inner apical edge, without tibial spur. Tarsal groove present as an impressed line on the outer edge (along the entire protibial length). Tarsi 5-5-5; protarsomere 5 as long as protarsomeres 2–4 combined. Claws well developed, about half the length of protarsomere 5. Mesofemur swollen, mesotibia narrow. Ŋree prominent setae can be seen in the median part of the less mesotibial outer margin. Mesotarsus identical to protarsus. Metafemur slender, slightly curved and swollen in the median part. Metatibia slender, with a couple stronger spines at outer apical corner; metatarsus identical to protarsus.—
<emphasis id="B951EABA171B81654E1AFD43FE0F79F5" bold="true" box="[322,431,739,763]" italics="true" pageId="19">Abdomen:</emphasis>
<emphasis id="B951EABA171981674CC1FC0BFC7978CD" bold="true" box="[921,985,939,963]" italics="true" pageId="17" pageNumber="18">Body:</emphasis>
Round, strongly dorsoventrally flattened (probably partially compressed). Cuticle regularly punctuate. Colour reddish brown to black.—
<emphasis id="B951EABA171981674BCCFC4AFB757F0C" bold="true" box="[1172,1237,1002,1026]" italics="true" pageId="17" pageNumber="18">Head:</emphasis>
Frons flat, wide and punctuate, strongly, squarely margined along inner edge of each eye, with a fine stria along their inner edges continuous with a complete, transverse frontal stria. Frontoclypeal suture not visible. Epistoma constricted, medially ridged, sides receding below broad frontal shelf. Labrum trapezoidal (width: length ratio 5: 1), emarginate anteriorly, with a pair of short setae at anterolateral corners, lacking punctuation. Mandibles wide and thin, basally with weakly dentate incisor edge, and a long, acute tip. Terminal maxillary palpomere wide and round; other ventral mouthparts damaged and/or missing. Antenna inserted under frontal shelf, strongly dorsoventrally flattened, possibly partially compressed. Scape short, widened, and curved to apex. Pedicel wide, subcylindrical, as long as antennomeres 3–5 combined. Antennomeres 3–7 similar in shape and size, short, wider than long (width: length ratio 3: 1). Antennomere 8 adjacent to the antennal club, 2.5× wider than antennomere 7. Antennomeres 9–11 form a wide, flat, and round antennal club with well-defined sutures.—
<emphasis id="B951EABA171981674CCCF9BEFC497D38" bold="true" box="[916,1001,1566,1590]" italics="true" pageId="17" pageNumber="18">Thorax:</emphasis>
Pronotal disc flat. Anterior margin widely emarginate, with fine marginal pronotal stria; not evident on lateral or basal margins. Basal margin broadly rounded, almost angularly projecting. Surface of pronotal disc densely punctuate, punctures separated by 1–2× their diameter. Scutellar shield not visible, obscured by bubbles. Elytra truncate, without dorsal striation, epipleural stria complete; elytral disc appears weakly depressed in humeral corners (possibly due to damage). Elytral punctuation with the same pattern as in pronotum. Prosternal lobe rather short, subtruncate anteriorly, set off from keel by pair of deep, setose foveae, weakly conjoined at their bases; prosternal keel flat, short, abruptly descending at front into two large approximate prosternal foveae, with fine longitudinal striae; finely punctate between; sides of prosternum widely incised for passage of antennal funicle, with deep channel reaching middle of base of prosternal keel; sides of prosternal keel appear concave. Mesoventrite wide and short, obscured by bubbles, probably finely punctuated. Meso-metaventral suture visible as a fine line. Metaventrite punctuate, with lateral metaventral stria directed posterolaterally to hind coxae. Other striation not visible. Metepisternum broadly exposed, rectangular.—
<emphasis id="B951EABA171B81654D9AFECCFD5A7A8D" bold="true" box="[706,762,364,387]" italics="true" pageId="19" pageNumber="20">Legs:</emphasis>
Profemur flatted, slightly widened. Protibia narrow and smooth, without dentation, few longer setae present along inner apical edge, without tibial spur. Tarsal groove present as an impressed line on the outer edge (along the entire protibial length). Tarsi 5-5-5; protarsomere 5 as long as protarsomeres 2–4 combined. Claws well developed, about half the length of protarsomere 5. Mesofemur swollen, mesotibia narrow. Three prominent setae can be seen in the median part of the left mesotibial outer margin. Mesotarsus identical to protarsus. Metafemur slender, slightly curved and swollen in the median part. Metatibia slender, with a couple stronger spines at outer apical corner; metatarsus identical to protarsus.—
<emphasis id="B951EABA171B81654E1AFD43FE0F79F5" bold="true" box="[322,431,739,763]" italics="true" pageId="19" pageNumber="20">Abdomen:</emphasis>
First visible abdominal sternite without visible striation. All visible abdominal sclerites finely punctuated. Propygidium very short, almost entirely covered by elytra. Pygidium weakly convex, about two-thirds as long as wide.
</paragraph>
<caption id="DF5A6620171A81644FD9F88CFBD07C6E" pageId="18" startId="18.[129,194,1836,1860]" targetBox="[175,1428,144,1808]" targetPageId="18" targetType="figure">
<paragraph id="8B9A36A8171A81644FD9F88CFBD07C6E" blockId="18.[129,1441,1836,1888]" pageId="18">
<emphasis id="B951EABA171A81644FD9F88CFF457C4A" bold="true" box="[129,229,1836,1860]" pageId="18">Figure 12.</emphasis>
<taxonomicName id="4C254D2B171A81644FB1F88CFE237C4A" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[233,387,1836,1860]" class="Insecta" family="Histeridae" genus="Placatister" kingdom="Animalia" order="Coleoptera" pageId="18" phylum="Arthropoda" rank="species" species="cascus" status="gen. et sp. nov">
<emphasis id="B951EABA171A81644FB1F88CFE237C4A" box="[233,387,1836,1860]" italics="true" pageId="18">Placatister cascus</emphasis>
</subSubSection>
<caption id="DF5A6620171A81644FD9F88CFBD07C6E" pageId="18" pageNumber="19" startId="18.[129,194,1836,1860]" targetBox="[175,1428,144,1808]" targetPageId="18" targetType="figure">
<paragraph id="8B9A36A8171A81644FD9F88CFBD07C6E" blockId="18.[129,1441,1836,1888]" pageId="18" pageNumber="19">
<emphasis id="B951EABA171A81644FD9F88CFF457C4A" bold="true" box="[129,229,1836,1860]" pageId="18" pageNumber="19">Figure 12.</emphasis>
<taxonomicName id="4C254D2B171A81644FB1F88CFE237C4A" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[233,387,1836,1860]" class="Insecta" family="Histeridae" genus="Placatister" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="cascus" status="gen. et sp. nov">
<emphasis id="B951EABA171A81644FB1F88CFE237C4A" box="[233,387,1836,1860]" italics="true" pageId="18" pageNumber="19">Placatister cascus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C1171A81644ED0F88DFDAC7C4A" box="[392,524,1837,1861]" pageId="18" rank="species">gen. et sp. nov</taxonomicNameLabel>
<taxonomicNameLabel id="A26257C1171A81644ED0F88DFDAC7C4A" box="[392,524,1837,1861]" pageId="18" pageNumber="19" rank="species">gen. et sp. nov</taxonomicNameLabel>
.. A, dorsal view. B, ventral view. C, lateral view. D, antennal club. E, mouthparts. F, frontal view. G, prosternal keel. H, antenna. Abbreviations: ees, elytral epipleural stria; fs, frontal stria; pf, prosternal foveae.
</paragraph>
</caption>
<paragraph id="8B9A36A8171B81654F29FC3EFEF37D26" blockId="19.[113,763,925,1576]" pageId="19">
<emphasis id="B951EABA171B81654F29FC3EFEC178BB" box="[113,353,926,949]" italics="true" pageId="19">Taxonomic assignment:</emphasis>
Ŋe species displays several highly plesiomorphic characters, such as the absence of a prosternal lobe and antennal cavity, as well as in the shape of the antennal club with distinct segments (
<figureCitation id="131E2A2D171B81654EFCFC5BFE5E7F1D" box="[420,510,1019,1043]" captionStart="Figure 12" captionStartId="18.[129,194,1836,1860]" captionTargetBox="[175,1428,144,1808]" captionTargetId="figure-8@18.[175,1428,144,1808]" captionTargetPageId="18" captionText="Figure 12. Placatister cascus gen.et sp. nov.. A, dorsal view.B, ventral view. C, lateral view.D, antennal club. E, mouthparts. F, frontal view.G, prosternal keel.H, antenna.Abbreviations:ees, elytral epipleural stria; fs, frontal stria; pf, prosternal foveae." pageId="19">Fig. 12D</figureCitation>
). Ŋe flaưened antenna with wide and short antennomeres 3–7 is especially unique among the entire
<taxonomicName id="4C254D2B171B81654E6BFB9AFE3E7F5C" box="[307,414,1082,1106]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="19" phylum="Arthropoda" rank="family">Histeridae</taxonomicName>
<subSubSection id="C33F6523171B81654F29FC3EFEF37D26" pageId="19" pageNumber="20" type="discussion">
<paragraph id="8B9A36A8171B81654F29FC3EFEF37D26" blockId="19.[113,763,925,1576]" pageId="19" pageNumber="20">
<emphasis id="B951EABA171B81654F29FC3EFEC178BB" box="[113,353,926,949]" italics="true" pageId="19" pageNumber="20">Taxonomic assignment:</emphasis>
The species displays several highly plesiomorphic characters, such as the absence of a prosternal lobe and antennal cavity, as well as in the shape of the antennal club with distinct segments (
<figureCitation id="131E2A2D171B81654EFCFC5BFE5E7F1D" box="[420,510,1019,1043]" captionStart="Figure 12" captionStartId="18.[129,194,1836,1860]" captionTargetBox="[175,1428,144,1808]" captionTargetId="figure-8@18.[175,1428,144,1808]" captionTargetPageId="18" captionText="Figure 12. Placatister cascus gen.et sp. nov.. A, dorsal view.B, ventral view. C, lateral view.D, antennal club. E, mouthparts. F, frontal view.G, prosternal keel.H, antenna.Abbreviations:ees, elytral epipleural stria; fs, frontal stria; pf, prosternal foveae." pageId="19" pageNumber="20">Fig. 12D</figureCitation>
). The flattened antenna with wide and short antennomeres 3–7 is especially unique among the entire
<taxonomicName id="4C254D2B171B81654E6BFB9AFE3E7F5C" box="[307,414,1082,1106]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="family">Histeridae</taxonomicName>
family. Prosternal structure, and particularly the prosternal foveae (
<figureCitation id="131E2A2D171B81654EB9FBF9FDFC7F7F" box="[481,604,1113,1137]" captionStart="Figure 12" captionStartId="18.[129,194,1836,1860]" captionTargetBox="[175,1428,144,1808]" captionTargetId="figure-8@18.[175,1428,144,1808]" captionTargetPageId="18" captionText="Figure 12. Placatister cascus gen.et sp. nov.. A, dorsal view.B, ventral view. C, lateral view.D, antennal club. E, mouthparts. F, frontal view.G, prosternal keel.H, antenna.Abbreviations:ees, elytral epipleural stria; fs, frontal stria; pf, prosternal foveae." pageId="19">Fig. 12B, G</figureCitation>
), are plausibly similar to those of some ºaprininae (e.g.
<taxonomicName id="4C254D2B171B81654D4FFBD9FD2B7F9E" authorityName="Marseul" authorityYear="1862" box="[535,651,1145,1168]" class="Insecta" family="Histeridae" genus="Tomogenius" kingdom="Animalia" order="Coleoptera" pageId="19" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA171B81654D4FFBD9FD2B7F9E" box="[535,651,1145,1168]" italics="true" pageId="19">Tomogenius</emphasis>
<figureCitation id="131E2A2D171B81654EB9FBF9FDFC7F7F" box="[481,604,1113,1137]" captionStart="Figure 12" captionStartId="18.[129,194,1836,1860]" captionTargetBox="[175,1428,144,1808]" captionTargetId="figure-8@18.[175,1428,144,1808]" captionTargetPageId="18" captionText="Figure 12. Placatister cascus gen.et sp. nov.. A, dorsal view.B, ventral view. C, lateral view.D, antennal club. E, mouthparts. F, frontal view.G, prosternal keel.H, antenna.Abbreviations:ees, elytral epipleural stria; fs, frontal stria; pf, prosternal foveae." pageId="19" pageNumber="20">Fig. 12B, G</figureCitation>
), are plausibly similar to those of some Saprininae (e.g.
<taxonomicName id="4C254D2B171B81654D4FFBD9FD2B7F9E" authorityName="Marseul" authorityYear="1862" box="[535,651,1145,1168]" class="Insecta" family="Histeridae" genus="Tomogenius" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA171B81654D4FFBD9FD2B7F9E" box="[535,651,1145,1168]" italics="true" pageId="19" pageNumber="20">Tomogenius</emphasis>
</taxonomicName>
) (
<bibRefCitation id="EFB44B59171B81654DFFFBD8FF057FBE" author="Lackner T" pageId="19" pagination="1 - 254" refId="ref18143" refString="Lackner T. Review of the Palaearctic genera of ºaprininae (Coleoptera: Histeridae). Acta Entomologica Musei Nationalis Pragae 2010; 50: 1 - 254." type="journal article" year="2010">Lackner 2010</bibRefCitation>
). Nevertheless, the homology of these structures is highly unlikely. Ŋe presence (if true) of an antennal receptacle along the sides of the prosternal keel would also be similar to that of ºaprininae. However, other characters, such as the distinctly sutured antennal club (
<figureCitation id="131E2A2D171B81654E1DFAB5FE3C7E23" box="[325,412,1301,1325]" captionStart="Figure 12" captionStartId="18.[129,194,1836,1860]" captionTargetBox="[175,1428,144,1808]" captionTargetId="figure-8@18.[175,1428,144,1808]" captionTargetPageId="18" captionText="Figure 12. Placatister cascus gen.et sp. nov.. A, dorsal view.B, ventral view. C, lateral view.D, antennal club. E, mouthparts. F, frontal view.G, prosternal keel.H, antenna.Abbreviations:ees, elytral epipleural stria; fs, frontal stria; pf, prosternal foveae." pageId="19">Fig. 12D</figureCitation>
), flaưened body shape (
<figureCitation id="131E2A2D171B81654DCBFAB5FD497E23" box="[659,745,1301,1325]" captionStart="Figure 12" captionStartId="18.[129,194,1836,1860]" captionTargetBox="[175,1428,144,1808]" captionTargetId="figure-8@18.[175,1428,144,1808]" captionTargetPageId="18" captionText="Figure 12. Placatister cascus gen.et sp. nov.. A, dorsal view.B, ventral view. C, lateral view.D, antennal club. E, mouthparts. F, frontal view.G, prosternal keel.H, antenna.Abbreviations:ees, elytral epipleural stria; fs, frontal stria; pf, prosternal foveae." pageId="19">Fig. 12C</figureCitation>
<bibRefCitation id="EFB44B59171B81654DFFFBD8FF057FBE" author="Lackner T" pageId="19" pageNumber="20" pagination="1 - 254" refId="ref18143" refString="Lackner T. Review of the Palaearctic genera of ºaprininae (Coleoptera: Histeridae). Acta Entomologica Musei Nationalis Pragae 2010; 50: 1 - 254." type="journal article" year="2010">Lackner 2010</bibRefCitation>
). Nevertheless, the homology of these structures is highly unlikely. The presence (if true) of an antennal receptacle along the sides of the prosternal keel would also be similar to that of Saprininae. However, other characters, such as the distinctly sutured antennal club (
<figureCitation id="131E2A2D171B81654E1DFAB5FE3C7E23" box="[325,412,1301,1325]" captionStart="Figure 12" captionStartId="18.[129,194,1836,1860]" captionTargetBox="[175,1428,144,1808]" captionTargetId="figure-8@18.[175,1428,144,1808]" captionTargetPageId="18" captionText="Figure 12. Placatister cascus gen.et sp. nov.. A, dorsal view.B, ventral view. C, lateral view.D, antennal club. E, mouthparts. F, frontal view.G, prosternal keel.H, antenna.Abbreviations:ees, elytral epipleural stria; fs, frontal stria; pf, prosternal foveae." pageId="19" pageNumber="20">Fig. 12D</figureCitation>
), flattened body shape (
<figureCitation id="131E2A2D171B81654DCBFAB5FD497E23" box="[659,745,1301,1325]" captionStart="Figure 12" captionStartId="18.[129,194,1836,1860]" captionTargetBox="[175,1428,144,1808]" captionTargetId="figure-8@18.[175,1428,144,1808]" captionTargetPageId="18" captionText="Figure 12. Placatister cascus gen.et sp. nov.. A, dorsal view.B, ventral view. C, lateral view.D, antennal club. E, mouthparts. F, frontal view.G, prosternal keel.H, antenna.Abbreviations:ees, elytral epipleural stria; fs, frontal stria; pf, prosternal foveae." pageId="19" pageNumber="20">Fig. 12C</figureCitation>
), and simple leg shape with slender tibiae without denticles (
<figureCitation id="131E2A2D171B81654D8CFA94FF3A7E62" captionStart="Figure 12" captionStartId="18.[129,194,1836,1860]" captionTargetBox="[175,1428,144,1808]" captionTargetId="figure-8@18.[175,1428,144,1808]" captionTargetPageId="18" captionText="Figure 12. Placatister cascus gen.et sp. nov.. A, dorsal view.B, ventral view. C, lateral view.D, antennal club. E, mouthparts. F, frontal view.G, prosternal keel.H, antenna.Abbreviations:ees, elytral epipleural stria; fs, frontal stria; pf, prosternal foveae." pageId="19">Fig. 12B</figureCitation>
), would be incompatible with this placement. Ŋe phylogenetic analyses did not resolve the position of this taxon, as it nested in various positions throughout the
<taxonomicName id="4C254D2B171B81654D75FA32FD387EA4" box="[557,664,1426,1450]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="19" phylum="Arthropoda" rank="family">Histeridae</taxonomicName>
tree (but outside the ºaprininae and Histeromorphae clades). Ŋis species most likely represents a primitive stem lineage of
<taxonomicName id="4C254D2B171B81654DD2FA71FD567EE7" box="[650,758,1489,1513]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="19" phylum="Arthropoda" rank="family">Histeridae</taxonomicName>
<figureCitation id="131E2A2D171B81654D8CFA94FF3A7E62" captionStart="Figure 12" captionStartId="18.[129,194,1836,1860]" captionTargetBox="[175,1428,144,1808]" captionTargetId="figure-8@18.[175,1428,144,1808]" captionTargetPageId="18" captionText="Figure 12. Placatister cascus gen.et sp. nov.. A, dorsal view.B, ventral view. C, lateral view.D, antennal club. E, mouthparts. F, frontal view.G, prosternal keel.H, antenna.Abbreviations:ees, elytral epipleural stria; fs, frontal stria; pf, prosternal foveae." pageId="19" pageNumber="20">Fig. 12B</figureCitation>
), would be incompatible with this placement. The phylogenetic analyses did not resolve the position of this taxon, as it nested in various positions throughout the
<taxonomicName id="4C254D2B171B81654D75FA32FD387EA4" box="[557,664,1426,1450]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="family">Histeridae</taxonomicName>
tree (but outside the Saprininae and Histeromorphae clades). This species most likely represents a primitive stem lineage of
<taxonomicName id="4C254D2B171B81654DD2FA71FD567EE7" box="[650,758,1489,1513]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="family">Histeridae</taxonomicName>
, but a subfamily or tribal assignment is impossible based on the observable characters.
</paragraph>
<paragraph id="8B9A36A8171B81654F29F9EDFE477C4E" blockId="19.[113,762,1612,1856]" pageId="19">
<emphasis id="B951EABA171B81654F29F9EDFF777D6A" box="[113,215,1613,1636]" italics="true" pageId="19">Diagnosis:</emphasis>
Head with a complete frontal stria. Antennomeres short and wide, antennae dorsoventrally flaưened. Prosternal process with a pair of setose foveae in apical part and with concave sides. Cuticle dorsally and ventrally punctuate. Legs narrow and smooth. Ŋe species may remotely resemble the flat
<taxonomicName id="4C254D2B171B81654DF3F96AFF577C0F" authorityName="Simon Prazak &amp; Lackner" authorityYear="2023" class="Insecta" family="Histeridae" genus="Olexum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" phylum="Arthropoda" rank="species" species="complanatum">
<emphasis id="B951EABA171B81654DF3F96AFF577C0F" italics="true" pageId="19">Olexum complanatum</emphasis>
</subSubSection>
<subSubSection id="C33F6523171B81654F29F9EDFE477C4E" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="8B9A36A8171B81654F29F9EDFE477C4E" blockId="19.[113,762,1612,1856]" pageId="19" pageNumber="20">
<emphasis id="B951EABA171B81654F29F9EDFF777D6A" box="[113,215,1613,1636]" italics="true" pageId="19" pageNumber="20">Diagnosis:</emphasis>
Head with a complete frontal stria. Antennomeres short and wide, antennae dorsoventrally flattened. Prosternal process with a pair of setose foveae in apical part and with concave sides. Cuticle dorsally and ventrally punctuate. Legs narrow and smooth. The species may remotely resemble the flat
<taxonomicName id="4C254D2B171B81654DF3F96AFF577C0F" authorityName="Simon Prazak &amp; Lackner" authorityYear="2023" class="Insecta" family="Histeridae" genus="Olexum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="complanatum">
<emphasis id="B951EABA171B81654DF3F96AFF577C0F" italics="true" pageId="19" pageNumber="20">Olexum complanatum</emphasis>
</taxonomicName>
or the round-bodied species of
<taxonomicName id="4C254D2B171B81654D19F949FD3B7C0F" authorityName="G. H. Horn" authorityYear="1873" box="[577,667,1769,1793]" class="Insecta" family="Histeridae" genus="Anapleus" kingdom="Animalia" order="Coleoptera" pageId="19" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA171B81654D19F949FD3B7C0F" box="[577,667,1769,1793]" italics="true" pageId="19">Anapleus</emphasis>
<taxonomicName id="4C254D2B171B81654D19F949FD3B7C0F" authorityName="G. H. Horn" authorityYear="1873" box="[577,667,1769,1793]" class="Insecta" family="Histeridae" genus="Anapleus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA171B81654D19F949FD3B7C0F" box="[577,667,1769,1793]" italics="true" pageId="19" pageNumber="20">Anapleus</emphasis>
</taxonomicName>
. It is distinguished by the wide and segmented antennal club and the wide and short segmented flagellum.
</paragraph>
<paragraph id="8B9A36A8171B81654F29F8C4FD4C7C92" blockId="19.[113,762,1892,1948]" pageId="19">
<emphasis id="B951EABA171B81654F29F8C4FF417C72" box="[113,225,1892,1916]" italics="true" pageId="19">Etymology:</emphasis>
Ŋis species represents a plesiomorphic set of characters, and the species’ name ‘cascus’ means ancient/primitive.
</subSubSection>
<subSubSection id="C33F6523171B81654F29F8C4FD4C7C92" pageId="19" pageNumber="20" type="etymology">
<paragraph id="8B9A36A8171B81654F29F8C4FD4C7C92" blockId="19.[113,762,1892,1948]" pageId="19" pageNumber="20">
<emphasis id="B951EABA171B81654F29F8C4FF417C72" box="[113,225,1892,1916]" italics="true" pageId="19" pageNumber="20">Etymology:</emphasis>
This species represents a plesiomorphic set of characters, and the species’ name ‘cascus’ means ancient/primitive.
</paragraph>
</subSubSection>
</treatment>

166
data/03/8C/87/038C87BE171981674CA4FF1FFC0E7929.xml
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@ -1,118 +1,116 @@
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<mods:affiliation id="495291321C493E27376039C0C4AB1BFF">Charles University, Faculty of Science, Department of Zoology, Viničná 7, 12800 Prague 2, Czech Republic &amp; Museum of Eastern Bohemia in Hradec Králové, Eliščino nábřeží 465, 500 03 Hradec Králové 3, Czech Republic</mods:affiliation>
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<heading id="D0D281C4171981674CA4FF1FFB417BD7" bold="true" box="[1020,1249,191,218]" centered="true" fontSize="11" level="2" pageId="17" reason="2">
<emphasis id="B951EABA171981674CA4FF1FFB417BD7" bold="true" box="[1020,1249,191,218]" pageId="17">
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<emphasis id="B951EABA171981674CA4FF1FFBD57BD4" bold="true" box="[1020,1141,191,218]" italics="true" pageId="17">Placatister</emphasis>
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<emphasis id="B951EABA171981674CA4FF1FFBD57BD4" bold="true" box="[1020,1141,191,218]" italics="true" pageId="17" pageNumber="18">Placatister</emphasis>
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<typeStatus id="549E880A171981674C72FEA0FCFA7A19" box="[810,858,256,279]" pageId="17" pageNumber="18">Type</typeStatus>
species.
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..
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<emphasis id="B951EABA171981674C72FE9AFC307A5F" box="[810,912,314,337]" italics="true" pageId="17">Diagnosis:</emphasis>
Round and flat body shape. Head with a complete frontal stria. Antennomeres short and wide, antennae dorsoventrally flaưened. Antennal club segmented. Antennal cavities occluded. Prosternal process with a pair of setose foveae in apical part and with concave sides. Legs narrow and smooth.
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<emphasis id="B951EABA171981674C72FE9AFC307A5F" box="[810,912,314,337]" italics="true" pageId="17" pageNumber="18">Diagnosis:</emphasis>
Round and flat body shape. Head with a complete frontal stria. Antennomeres short and wide, antennae dorsoventrally flattened. Antennal club segmented. Antennal cavities occluded. Prosternal process with a pair of setose foveae in apical part and with concave sides. Legs narrow and smooth.
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<emphasis id="B951EABA171981674C72FE50FC397906" box="[810,921,496,520]" italics="true" pageId="17">Etymology:</emphasis>
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<emphasis id="B951EABA171981674C72FE50FC397906" box="[810,921,496,520]" italics="true" pageId="17" pageNumber="18">Etymology:</emphasis>
Masculine, derived from the Czech word ‘placatý’ meaning flat.
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289
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<mods:namePart id="6C5A3F9509C7236332901769E7F52B31">ºimon-Pražák</mods:namePart>
<mods:affiliation id="9C4DEAB783EB80617B24FF1925D76B0D">Charles University, Faculty of ºcience, Department of Zoology, Viničná 7, 12800 Prague 2, Czech Republic &amp; Museum of Eastern Bohemia in Hradec Králové, Eliščino nábřeží 465, 500 03 Hradec Králové 3, Czech Republic</mods:affiliation>
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<mods:namePart id="7B16151942B3FFFCD704069CD62A6A0C">Yamamoto, Shûhei</mods:namePart>
<mods:affiliation id="10A31C9D34DBF68076498A7179003F52">The Hokkaido University Museum, Hokkaido University, Kita 10, Nishi 8, Kita-ku, Sapporo 060 - 0810, Japan</mods:affiliation>
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<mods:namePart id="8FEECDA0043CBE8D4E1FEFD0955D3CA8">Yamamoto</mods:namePart>
<mods:affiliation id="6B4CF1CE3EFF63C4B137A598F7730B93">Ŋe Hokkaido University Museum, Hokkaido University, Kita 10, Nishi 8, Kita-ku, ºapporo 060 - 0810, Japan</mods:affiliation>
<mods:namePart id="D3A087D135CD04BBBFD494B8F18A993B">Lackner, Tomáš</mods:namePart>
<mods:affiliation id="9F94A9AED9B12A8EB36CE976F2FDAD97">Department of Environmental Systems Science, ETH Zürich, Weinbergstrasse 56, 8092 Zürich, Switzerland</mods:affiliation>
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<mods:namePart id="E36C137FC67242BA18FF75EC7D6743B3">Lackner, Tomáš</mods:namePart>
<mods:affiliation id="6F802A6CC77AC204BD0EB47CC78324DF">Department of Environmental ºystems ºcience, ETH Zürich, Weinbergstrasse 56, 8092 Zürich, ºwitzerland</mods:affiliation>
<mods:namePart id="9DEE9F5AE7EC085906765DFD30508BF3">Fikáček, Martin</mods:namePart>
<mods:affiliation id="8EF9933F3D119A3830DF395D84069BED">Department of Biological Sciences, National Sun Yat-sen University, No. 70, Lienhai Rd., Kaohsiung 80424, Taiwan &amp; Department of Entomology, National Museum, Cirkusová 1740 CZ- 19300 Praha 9, Czech Republic</mods:affiliation>
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<mods:namePart id="67FD9F177218AE43E1A60AE56E57B668">Fikáček, Martin</mods:namePart>
<mods:affiliation id="4CC7B46AD0C876D9FA17877A7576FF8A">Department of Biological ºciences, National ºun Yat-sen University, No. 70, Lienhai Rd., Kaohsiung 80424, Taiwan &amp; Department of Entomology, National Museum, Cirkusová 1740 CZ- 19300 Praha 9, Czech Republic</mods:affiliation>
<mods:namePart id="5403381F60F9CEAD1850DA6511BF2A78">Prokop, Jakub</mods:namePart>
<mods:affiliation id="9BD5D87C31B07537A81407A93DC9E031">Charles University, Faculty of Science, Department of Zoology, Viničná 7, 12800 Prague 2, Czech Republic &amp; Department of Plant and Environmental Sciences, Clemson University, Clemson, SC 29634, United States</mods:affiliation>
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<mods:namePart id="BD2FB4BBCD7237BD4E1FC87F8B132755">Prokop, Jakub</mods:namePart>
<mods:affiliation id="FB4A25CD4E0B370E0D26D6FE0CC4E54C">Charles University, Faculty of ºcience, Department of Zoology, Viničná 7, 12800 Prague 2, Czech Republic</mods:affiliation>
<mods:namePart id="7A1E9C4A4BC84FD6D79DB786CEE1A9B7">Caterino, Michael S</mods:namePart>
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<mods:affiliation id="6C2A32DBFA29818567D97A2D4B2C4696">Department of Plant and Environmental ºciences, Clemson University, Clemson, ºC 29634, United ºtates</mods:affiliation>
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<mods:date id="D7F32122CF0D2AF319E0153778D9DF76">2024</mods:date>
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<mods:title id="120222131C40883F00817F54050954C0">zlae 137</mods:title>
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<treatment id="038C87BE171B81634C90FD52FC0F7BC9" LSID="urn:lsid:plazi:treatment:038C87BE171B81634C90FD52FC0F7BC9" httpUri="http://treatment.plazi.org/id/038C87BE171B81634C90FD52FC0F7BC9" lastPageId="21" pageId="19">
<subSubSection id="C33F6523171B81654C90FD52FAB57803" box="[968,1301,754,781]" pageId="19" type="nomenclature">
<paragraph id="8B9A36A8171B81654C90FD52FAB57803" blockId="19.[968,1301,754,781]" box="[968,1301,754,781]" pageId="19">
<heading id="D0D281C4171B81654C90FD52FAB57803" bold="true" box="[968,1301,754,781]" centered="true" fontSize="11" level="2" pageId="19" reason="2">
<emphasis id="B951EABA171B81654C90FD52FAB57803" bold="true" box="[968,1301,754,781]" pageId="19">
<taxonomicName id="4C254D2B171B81654C90FD52FB187803" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[968,1208,754,781]" class="Insecta" family="Histeridae" genus="Miculissima" kingdom="Animalia" order="Coleoptera" pageId="19" phylum="Arthropoda" rank="species" species="excavata" status="sp. nov.">
<emphasis id="B951EABA171B81654C90FD52FB187803" bold="true" box="[968,1208,754,781]" italics="true" pageId="19">Miculissima excavata</emphasis>
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<subSubSection id="C33F6523171B81654C90FD52FAB57803" box="[968,1301,754,781]" pageId="19" pageNumber="20" type="nomenclature">
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<heading id="D0D281C4171B81654C90FD52FAB57803" bold="true" box="[968,1301,754,781]" centered="true" fontSize="11" level="2" pageId="19" pageNumber="20" reason="2">
<emphasis id="B951EABA171B81654C90FD52FAB57803" bold="true" box="[968,1301,754,781]" pageId="19" pageNumber="20">
<taxonomicName id="4C254D2B171B81654C90FD52FB187803" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[968,1208,754,781]" class="Insecta" family="Histeridae" genus="Miculissima" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="excavata" status="sp. nov.">
<emphasis id="B951EABA171B81654C90FD52FB187803" bold="true" box="[968,1208,754,781]" italics="true" pageId="19" pageNumber="20">Miculissima excavata</emphasis>
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<taxonomicNameLabel id="A26257C1171B81654BE6FD53FAB57803" box="[1214,1301,755,781]" pageId="19" rank="species">sp. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="A26257C1171B81654BE6FD53FAB57803" box="[1214,1301,755,781]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
</paragraph>
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<subSubSection id="C33F6523171B81634B18FC81FC0F7BC9" lastPageId="21" pageId="19" type="description">
<paragraph id="8B9A36A8171B81654B18FC81FB3D7835" blockId="19.[1088,1181,801,828]" box="[1088,1181,801,828]" pageId="19">
<emphasis id="B951EABA171B81654B18FC81FB3D7835" bold="true" box="[1088,1181,801,828]" pageId="19">
<subSubSection id="C33F6523171B81654B18FC81FB3D7835" box="[1088,1181,801,828]" pageId="19" pageNumber="20" type="description">
<paragraph id="8B9A36A8171B81654B18FC81FB3D7835" blockId="19.[1088,1181,801,828]" box="[1088,1181,801,828]" pageId="19" pageNumber="20">
<emphasis id="B951EABA171B81654B18FC81FB3D7835" bold="true" box="[1088,1181,801,828]" pageId="19" pageNumber="20">
(
<figureCitation id="131E2A2D171B81654B13FC82FB327832" box="[1099,1170,802,828]" captionStart="Figure 13" captionStartId="20.[129,194,1346,1370]" captionTargetBox="[129,1473,144,1319]" captionTargetId="figure-333@20.[129,1473,144,1319]" captionTargetPageId="20" captionText="Figure 13. Miculissima excavata gen. et sp.nov.. A, dorsal view. B, frontal view.C, ventral view.D, lateral view. E, lateral view. Abbreviations:aca, antennal cavity; acl, antennal club; ga, galea; lp, labial palpus; ls, labral setae; mp, maxillary palpus; pl, prosternal lobe." pageId="19">Fig. 13</figureCitation>
<figureCitation id="131E2A2D171B81654B13FC82FB327832" box="[1099,1170,802,828]" captionStart="Figure 13" captionStartId="20.[129,194,1346,1370]" captionTargetBox="[129,1473,144,1319]" captionTargetId="figure-333@20.[129,1473,144,1319]" captionTargetPageId="20" captionText="Figure 13. Miculissima excavata gen. et sp.nov.. A, dorsal view. B, frontal view.C, ventral view.D, lateral view. E, lateral view. Abbreviations:aca, antennal cavity; acl, antennal club; ga, galea; lp, labial palpus; ls, labral setae; mp, maxillary palpus; pl, prosternal lobe." pageId="19" pageNumber="20">Fig. 13</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8B9A36A8171B81654C72FCC2FAF978B7" blockId="19.[810,1460,866,953]" pageId="19">
<emphasis id="B951EABA171B81654C72FCC2FC267874" box="[810,902,866,890]" italics="true" pageId="19">Material:</emphasis>
<collectingRegion id="49E1F84A171B81654CD5FCC2FC777874" box="[909,983,866,890]" country="Myanmar" name="Kachin" pageId="19">Kachin</collectingRegion>
</subSubSection>
<subSubSection id="C33F6523171B81654C72FCC2FAF978B7" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="8B9A36A8171B81654C72FCC2FAF978B7" blockId="19.[810,1460,866,953]" pageId="19" pageNumber="20">
<emphasis id="B951EABA171B81654C72FCC2FC267874" box="[810,902,866,890]" italics="true" pageId="19" pageNumber="20">Material:</emphasis>
<materialsCitation id="3B4D3CF5171B81654CD5FCC2FAF578B7" collectionCode="HUM" location="Kachin" pageId="19" pageNumber="20" specimenCode="SEHU-0000121230" specimenCount="1" stateProvince="Kachin">
<collectingRegion id="49E1F84A171B81654CD5FCC2FC777874" box="[909,983,866,890]" country="Myanmar" name="Kachin" pageId="19" pageNumber="20">Kachin</collectingRegion>
amber (
<emphasis id="B951EABA171B81654B69FCC3FB9A7874" box="[1073,1082,867,890]" italics="true" pageId="19">c</emphasis>
<emphasis id="B951EABA171B81654B69FCC3FB9A7874" box="[1073,1082,867,890]" italics="true" pageId="19" pageNumber="20">c</emphasis>
. 99 Mya). Amber piece dimensions: 9 × 25 ×
<quantity id="4CDD9B4D171B81654CD2FC22FC667897" box="[906,966,898,922]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="19" unit="mm" value="2.0">2 mm</quantity>
. Amber clear with several insect fragments and air bubbles. Deposited at HUM (ºEHU-0000121230).
<quantity id="4CDD9B4D171B81654CD2FC22FC667897" box="[906,966,898,922]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="20" unit="mm" value="2.0">2 mm</quantity>
. Amber clear with several insect fragments and air bubbles. Deposited at
<collectionCode id="ED34AE6D171B81654B74FC01FBCB78B6" box="[1068,1131,929,952]" pageId="19" pageNumber="20">HUM</collectionCode>
(
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)
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.
</paragraph>
<paragraph id="8B9A36A8171B81624C72FC7BFACC7D97" blockId="19.[810,1460,986,1950]" lastBlockId="20.[825,1475,1446,1689]" lastPageId="20" pageId="19">
<emphasis id="B951EABA171B81654C72FC7BFC1C78FC" box="[810,956,987,1010]" italics="true" pageId="19">Measurements:</emphasis>
</subSubSection>
<subSubSection id="C33F6523171B81624C72FC7BFACC7D97" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="description">
<paragraph id="8B9A36A8171B81624C72FC7BFACC7D97" blockId="19.[810,1460,986,1950]" lastBlockId="20.[825,1475,1446,1689]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
<emphasis id="B951EABA171B81654C72FC7BFC1C78FC" box="[810,956,987,1010]" italics="true" pageId="19" pageNumber="20">Measurements:</emphasis>
Head width:
<quantity id="4CDD9B4D171B81654B0AFC7AFB0D78FC" box="[1106,1197,986,1010]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="19" unit="mm" value="0.24">0.24 mm</quantity>
<quantity id="4CDD9B4D171B81654B0AFC7AFB0D78FC" box="[1106,1197,986,1010]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="19" pageNumber="20" unit="mm" value="0.24">0.24 mm</quantity>
; width between anterior pronotal angles:
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<quantity id="4CDD9B4D171B81654C8AFC5AFB8C7F1F" box="[978,1068,1018,1042]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="19" pageNumber="20" unit="mm" value="0.26">0.26 mm</quantity>
; width between posterior pronotal angles:
<quantity id="4CDD9B4D171B81654C06FBB9FC197F3E" box="[862,953,1049,1073]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" pageId="19" unit="mm" value="0.46">0.46 mm</quantity>
<quantity id="4CDD9B4D171B81654C06FBB9FC197F3E" box="[862,953,1049,1073]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" pageId="19" pageNumber="20" unit="mm" value="0.46">0.46 mm</quantity>
; pronotal length:
<quantity id="4CDD9B4D171B81654B29FBB9FB6D7F3E" box="[1137,1229,1049,1073]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="19" unit="mm" value="0.22">0.22 mm</quantity>
<quantity id="4CDD9B4D171B81654B29FBB9FB6D7F3E" box="[1137,1229,1049,1073]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="19" pageNumber="20" unit="mm" value="0.22">0.22 mm</quantity>
; sutural elytral length:
<quantity id="4CDD9B4D171B81654C73FB98FCD87F5E" box="[811,888,1080,1104]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="19" unit="mm" value="0.6">0.6 mm</quantity>
<quantity id="4CDD9B4D171B81654C73FB98FCD87F5E" box="[811,888,1080,1104]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="19" pageNumber="20" unit="mm" value="0.6">0.6 mm</quantity>
; elytral width (in pair, across widest point):
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<quantity id="4CDD9B4D171B81654A6FFB98FA327F5E" box="[1335,1426,1080,1104]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" pageId="19" pageNumber="20" unit="mm" value="0.54">0.54 mm</quantity>
.—
<emphasis id="B951EABA171B81654C72FBF7FCCA7F61" bold="true" box="[810,874,1111,1135]" italics="true" pageId="19">Body:</emphasis>
<emphasis id="B951EABA171B81654C72FBF7FCCA7F61" bold="true" box="[810,874,1111,1135]" italics="true" pageId="19" pageNumber="20">Body:</emphasis>
Oval and elongate. Dorsal cuticle glabrous with irregular longitudinaldeformationsintherightside.Colourreddishbrown to black.—
<emphasis id="B951EABA171B81654CC4FB36FC7D7FA0" bold="true" box="[924,989,1174,1198]" italics="true" pageId="19">Head:</emphasis>
<emphasis id="B951EABA171B81654CC4FB36FC7D7FA0" bold="true" box="[924,989,1174,1198]" italics="true" pageId="19" pageNumber="20">Head:</emphasis>
Frons smooth, flat, and wide, slightly elevated above antennal bases. Frontal stria absent. Clypeus convex, elevated along midline, truncate apically. Labrum short and wide, rounded apically, with a prominent seta on each side. Mandibles mostly retracted. Terminal maxillary palpomere swollen at base, pointed apically, slightly concave on inner margin, 4× as long as maxillary palpomere 3. Maxillary palpomere 2 twice as long as palpomere 3. Terminal labial palpomere oblong, widest medially. Antennal scape simple, slightly curved. Pedicel slender, as long as antennomeres 3–4; Antennomeres 3–8 equal in shape and length, simple square shaped, widening slightly from 3 to 8. Antennal club round, undivided, but with two setose, outwardly arcuate annuli. Eyes convex, coarsely faceted, well visible from above.—
<emphasis id="B951EABA171B81654CDFF98DFC7B7D4B" bold="true" box="[903,987,1581,1605]" italics="true" pageId="19">Ŋorax:</emphasis>
Pronotal disc smooth, slightly convex. Basal pronotal margin widely angular. Posterolateral pronotal corners curved and pointed posteriorly. Right-half of pronotal disc is irregularly deformed. Marginal pronotal stria present, complete. Lateral pronotal stria present, marginal bead elevated. ºcutellar shield tiny, triangular. Elytra smooth, elongate, slightly convex, with very fine, scratch-like striae at outer base. Right elytron irregularly deformed. Epipleural stria not apparent. Wings present (visible extending beyond elytra). Prosternum short and wide, with a very short prosternal lobe; prosternum incised at sides for passage of antennal funicle; antennal cavity widely open beneath (prosternal alae absent). Prosternal keel weakly emarginate at base, obscured by debris. Mesoventrite smooth and relatively narrow (length: width ratio 1: 2), weakly produced anteriorly, without striae. Mesometaventral suture visible as a thin line. Metaventrite slightly concave medially. Lateral metaventral stria present in anterior third of the metaventral disc, curved laterally extending to metepisternum, which is narrow, but exposed over its entire length.—
<emphasis id="B951EABA171C81624E1EF9C3FEDD7D74" bold="true" box="[326,381,1635,1658]" italics="true" pageId="20">Legs:</emphasis>
<emphasis id="B951EABA171B81654CDFF98DFC7B7D4B" bold="true" box="[903,987,1581,1605]" italics="true" pageId="19" pageNumber="20">Thorax:</emphasis>
Pronotal disc smooth, slightly convex. Basal pronotal margin widely angular. Posterolateral pronotal corners curved and pointed posteriorly. Right-half of pronotal disc is irregularly deformed. Marginal pronotal stria present, complete. Lateral pronotal stria present, marginal bead elevated. Scutellar shield tiny, triangular. Elytra smooth, elongate, slightly convex, with very fine, scratch-like striae at outer base. Right elytron irregularly deformed. Epipleural stria not apparent. Wings present (visible extending beyond elytra). Prosternum short and wide, with a very short prosternal lobe; prosternum incised at sides for passage of antennal funicle; antennal cavity widely open beneath (prosternal alae absent). Prosternal keel weakly emarginate at base, obscured by debris. Mesoventrite smooth and relatively narrow (length: width ratio 1: 2), weakly produced anteriorly, without striae. Mesometaventral suture visible as a thin line. Metaventrite slightly concave medially. Lateral metaventral stria present in anterior third of the metaventral disc, curved laterally extending to metepisternum, which is narrow, but exposed over its entire length.—
<emphasis id="B951EABA171C81624E1EF9C3FEDD7D74" bold="true" box="[326,381,1635,1658]" italics="true" pageId="20" pageNumber="21">Legs:</emphasis>
Profemur carinate, with a row of tiny setae along inner margin and a simple groove for protibial reception. Protibia dilated in apical two-thirds, with a simple tarsal groove, with several minute setae apically on inner margin; outer protibial margin smooth, but with a couple spines near the widest part of tibia. Protibia apical margin oblique, lacking apical spurs; Mesofemur slender, swollen medially. Mesotibia widened (slightly less than protibia), gradually expanding apically with a simple tarsal groove and a tiny tibial spur. Meso- and metatarsi similar to protarsus. Metafemur long and slender. Metatibia very slightly widened to apex, with minute setae in apical half of inner margin and a small tibial spur. Tarsi 5-5-5, all basal tarsomeres bearing a prominent pair of apical setae beneath, some with sparse secondary setae as well; protarsus simple, terminal tarsomere as long as tarsomeres 2–4 combined. Mesotarsus similar in shape. Metatarsomere 1 shortened, half of the length of metatarsomere 2.—
<emphasis id="B951EABA171C81624BF4F9E2FAB97D54" bold="true" box="[1196,1305,1602,1626]" italics="true" pageId="20">Abdomen:</emphasis>
<emphasis id="B951EABA171C81624BF4F9E2FAB97D54" bold="true" box="[1196,1305,1602,1626]" italics="true" pageId="20" pageNumber="21">Abdomen:</emphasis>
First abdominal ventrite smooth, with two widely separated parallel lateral striae, posterior margin widely emarginate. Pygidium smooth.
</paragraph>
<caption id="DF5A6620171C81624FD9FAE2FB6D7E78" pageId="20" startId="20.[129,194,1346,1370]" targetBox="[129,1473,144,1319]" targetPageId="20" targetType="figure">
<paragraph id="8B9A36A8171C81624FD9FAE2FB6D7E78" blockId="20.[129,1473,1346,1399]" pageId="20">
<emphasis id="B951EABA171C81624FD9FAE2FF457E54" bold="true" box="[129,229,1346,1370]" pageId="20">Figure 13.</emphasis>
<taxonomicName id="4C254D2B171C81624FB1FAE2FE097E54" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[233,425,1346,1370]" class="Insecta" family="Histeridae" genus="Miculissima" kingdom="Animalia" order="Coleoptera" pageId="20" phylum="Arthropoda" rank="species" species="excavata" status="gen. et sp. nov">
<emphasis id="B951EABA171C81624FB1FAE2FE097E54" box="[233,425,1346,1370]" italics="true" pageId="20">Miculissima excavata</emphasis>
</subSubSection>
<caption id="DF5A6620171C81624FD9FAE2FB6D7E78" pageId="20" pageNumber="21" startId="20.[129,194,1346,1370]" targetBox="[129,1473,144,1319]" targetPageId="20" targetType="figure">
<paragraph id="8B9A36A8171C81624FD9FAE2FB6D7E78" blockId="20.[129,1473,1346,1399]" pageId="20" pageNumber="21">
<emphasis id="B951EABA171C81624FD9FAE2FF457E54" bold="true" box="[129,229,1346,1370]" pageId="20" pageNumber="21">Figure 13.</emphasis>
<taxonomicName id="4C254D2B171C81624FB1FAE2FE097E54" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[233,425,1346,1370]" class="Insecta" family="Histeridae" genus="Miculissima" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="excavata" status="gen. et sp. nov">
<emphasis id="B951EABA171C81624FB1FAE2FE097E54" box="[233,425,1346,1370]" italics="true" pageId="20" pageNumber="21">Miculissima excavata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C1171C81624EF6FAE3FD917E54" box="[430,561,1347,1371]" pageId="20" rank="species">gen. et sp. nov</taxonomicNameLabel>
<taxonomicNameLabel id="A26257C1171C81624EF6FAE3FD917E54" box="[430,561,1347,1371]" pageId="20" pageNumber="21" rank="species">gen. et sp. nov</taxonomicNameLabel>
.. A, dorsal view. B, frontal view. C, ventral view. D, lateral view. E, lateral view. Abbreviations: aca, antennal cavity; acl, antennal club; ga, galea; lp, labial palpus; ls, labral setae; mp, maxillary palpus; pl, prosternal lobe.
</paragraph>
</caption>
<paragraph id="8B9A36A8171C81634C61F91CFD377A08" blockId="20.[825,1474,1723,1966]" lastBlockId="21.[113,763,144,262]" lastPageId="21" pageId="20">
<emphasis id="B951EABA171C81624C61F91CFBBF7DDD" box="[825,1055,1724,1747]" italics="true" pageId="20">Taxonomic assignment:</emphasis>
Ŋe absence of a distinct pronotal lobe, minute body size, and simple legs would suggest a relation to
<taxonomicName id="4C254D2B171C81624C61F959FBE67C1C" authority="(Wenzel 1944)" baseAuthorityName="Wenzel" baseAuthorityYear="1944" box="[825,1094,1785,1810]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="20" phylum="Arthropoda" rank="subFamily" subFamily="Abraeinae">
<subSubSection id="C33F6523171C81634C61F91CFD377A08" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="discussion">
<paragraph id="8B9A36A8171C81634C61F91CFD377A08" blockId="20.[825,1474,1723,1966]" lastBlockId="21.[113,763,144,262]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
<emphasis id="B951EABA171C81624C61F91CFBBF7DDD" box="[825,1055,1724,1747]" italics="true" pageId="20" pageNumber="21">Taxonomic assignment:</emphasis>
The absence of a distinct pronotal lobe, minute body size, and simple legs would suggest a relation to
<taxonomicName id="4C254D2B171C81624C61F959FBE67C1C" authority="(Wenzel 1944)" baseAuthorityName="Wenzel" baseAuthorityYear="1944" box="[825,1094,1785,1810]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="subFamily" subFamily="Abraeinae">
Abraeinae (
<bibRefCitation id="EFB44B59171C81624CEBF959FB9A7C1C" author="Wenzel RL" box="[947,1082,1785,1810]" pageId="20" pagination="51 - 151" refId="ref19054" refString="Wenzel RL. On the classification of the histerid beetles. Fieldiana Zoology 1944; 28: 51 - 151. hups: // doi. org / 10.1093 / aesa / 37.3.375" type="journal article" year="1944">Wenzel 1944</bibRefCitation>
<bibRefCitation id="EFB44B59171C81624CEBF959FB9A7C1C" author="Wenzel RL" box="[947,1082,1785,1810]" pageId="20" pageNumber="21" pagination="51 - 151" refId="ref19054" refString="Wenzel RL. On the classification of the histerid beetles. Fieldiana Zoology 1944; 28: 51 - 151. hups: // doi. org / 10.1093 / aesa / 37.3.375" type="journal article" year="1944">Wenzel 1944</bibRefCitation>
)
</taxonomicName>
. Labrum with two prominent lateral setae (
<figureCitation id="131E2A2D171C81624CD9F8B9FC7D7C3F" box="[897,989,1817,1841]" captionStart="Figure 13" captionStartId="20.[129,194,1346,1370]" captionTargetBox="[129,1473,144,1319]" captionTargetId="figure-333@20.[129,1473,144,1319]" captionTargetPageId="20" captionText="Figure 13. Miculissima excavata gen. et sp.nov.. A, dorsal view. B, frontal view.C, ventral view.D, lateral view. E, lateral view. Abbreviations:aca, antennal cavity; acl, antennal club; ga, galea; lp, labial palpus; ls, labral setae; mp, maxillary palpus; pl, prosternal lobe." pageId="20">Fig. 13D</figureCitation>
<figureCitation id="131E2A2D171C81624CD9F8B9FC7D7C3F" box="[897,989,1817,1841]" captionStart="Figure 13" captionStartId="20.[129,194,1346,1370]" captionTargetBox="[129,1473,144,1319]" captionTargetId="figure-333@20.[129,1473,144,1319]" captionTargetPageId="20" captionText="Figure 13. Miculissima excavata gen. et sp.nov.. A, dorsal view. B, frontal view.C, ventral view.D, lateral view. E, lateral view. Abbreviations:aca, antennal cavity; acl, antennal club; ga, galea; lp, labial palpus; ls, labral setae; mp, maxillary palpus; pl, prosternal lobe." pageId="20" pageNumber="21">Fig. 13D</figureCitation>
) and convex eyes (
<figureCitation id="131E2A2D171C81624BECF8B9FAAD7C3F" box="[1204,1293,1817,1841]" captionStart="Figure 13" captionStartId="20.[129,194,1346,1370]" captionTargetBox="[129,1473,144,1319]" captionTargetId="figure-333@20.[129,1473,144,1319]" captionTargetPageId="20" captionText="Figure 13. Miculissima excavata gen. et sp.nov.. A, dorsal view. B, frontal view.C, ventral view.D, lateral view. E, lateral view. Abbreviations:aca, antennal cavity; acl, antennal club; ga, galea; lp, labial palpus; ls, labral setae; mp, maxillary palpus; pl, prosternal lobe." pageId="20">Fig. 13E</figureCitation>
) are also typical among the subfamily. Ŋe species exhibits a distinctly annulate antennal club (
<figureCitation id="131E2A2D171C81624B52F8F7FBC67C7E" box="[1034,1126,1879,1904]" captionStart="Figure 13" captionStartId="20.[129,194,1346,1370]" captionTargetBox="[129,1473,144,1319]" captionTargetId="figure-333@20.[129,1473,144,1319]" captionTargetPageId="20" captionText="Figure 13. Miculissima excavata gen. et sp.nov.. A, dorsal view. B, frontal view.C, ventral view.D, lateral view. E, lateral view. Abbreviations:aca, antennal cavity; acl, antennal club; ga, galea; lp, labial palpus; ls, labral setae; mp, maxillary palpus; pl, prosternal lobe." pageId="20">Fig. 13C</figureCitation>
<figureCitation id="131E2A2D171C81624BECF8B9FAAD7C3F" box="[1204,1293,1817,1841]" captionStart="Figure 13" captionStartId="20.[129,194,1346,1370]" captionTargetBox="[129,1473,144,1319]" captionTargetId="figure-333@20.[129,1473,144,1319]" captionTargetPageId="20" captionText="Figure 13. Miculissima excavata gen. et sp.nov.. A, dorsal view. B, frontal view.C, ventral view.D, lateral view. E, lateral view. Abbreviations:aca, antennal cavity; acl, antennal club; ga, galea; lp, labial palpus; ls, labral setae; mp, maxillary palpus; pl, prosternal lobe." pageId="20" pageNumber="21">Fig. 13E</figureCitation>
) are also typical among the subfamily. The species exhibits a distinctly annulate antennal club (
<figureCitation id="131E2A2D171C81624B52F8F7FBC67C7E" box="[1034,1126,1879,1904]" captionStart="Figure 13" captionStartId="20.[129,194,1346,1370]" captionTargetBox="[129,1473,144,1319]" captionTargetId="figure-333@20.[129,1473,144,1319]" captionTargetPageId="20" captionText="Figure 13. Miculissima excavata gen. et sp.nov.. A, dorsal view. B, frontal view.C, ventral view.D, lateral view. E, lateral view. Abbreviations:aca, antennal cavity; acl, antennal club; ga, galea; lp, labial palpus; ls, labral setae; mp, maxillary palpus; pl, prosternal lobe." pageId="20" pageNumber="21">Fig. 13C</figureCitation>
)—a plesiomorphy found within
<taxonomicName id="4C254D2B171C81624C61F8D7FC017C81" baseAuthorityName="Wenzel" baseAuthorityYear="1944" box="[825,929,1911,1935]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="20" phylum="Arthropoda" rank="subFamily" subFamily="Abraeinae">Abraeinae</taxonomicName>
<taxonomicName id="4C254D2B171C81624C61F8D7FC017C81" baseAuthorityName="Wenzel" baseAuthorityYear="1944" box="[825,929,1911,1935]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="subFamily" subFamily="Abraeinae">Abraeinae</taxonomicName>
only in
<taxonomicName id="4C254D2B171C81624CABF8D7FB347C81" authorityName="Wenzel" authorityYear="1944" box="[1011,1172,1911,1935]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="20" phylum="Arthropoda" rank="tribe" tribe="Acritomorphini">Acritomorphini</taxonomicName>
<taxonomicName id="4C254D2B171C81624CABF8D7FB347C81" authorityName="Wenzel" authorityYear="1944" box="[1011,1172,1911,1935]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="tribe" tribe="Acritomorphini">Acritomorphini</taxonomicName>
(
<bibRefCitation id="EFB44B59171C81624BFDF8D7FA8A7C81" author="Wenzel RL" box="[1189,1322,1911,1935]" pageId="20" pagination="51 - 151" refId="ref19054" refString="Wenzel RL. On the classification of the histerid beetles. Fieldiana Zoology 1944; 28: 51 - 151. hups: // doi. org / 10.1093 / aesa / 37.3.375" type="journal article" year="1944">Wenzel 1944</bibRefCitation>
<bibRefCitation id="EFB44B59171C81624BFDF8D7FA8A7C81" author="Wenzel RL" box="[1189,1322,1911,1935]" pageId="20" pageNumber="21" pagination="51 - 151" refId="ref19054" refString="Wenzel RL. On the classification of the histerid beetles. Fieldiana Zoology 1944; 28: 51 - 151. hups: // doi. org / 10.1093 / aesa / 37.3.375" type="journal article" year="1944">Wenzel 1944</bibRefCitation>
,
<bibRefCitation id="EFB44B59171C81624A6DF8D7FA117C81" author="Mazur" box="[1333,1457,1911,1935]" pageId="20" pagination="27 - 32" refId="ref18506" refString="Mazur º. New species of histerid beetles (Coleoptera, Histeridae). Annales Zoologici 1997; 38: 27 - 32. hup: // rcin. org. pl" type="journal article" year="1997">Mazur 1997</bibRefCitation>
). Ŋe excavate and sculptured pronotal hypomeron (
<figureCitation id="131E2A2D171C81624A1DF836FA177CA0" box="[1349,1463,1942,1966]" captionStart="Figure 13" captionStartId="20.[129,194,1346,1370]" captionTargetBox="[129,1473,144,1319]" captionTargetId="figure-333@20.[129,1473,144,1319]" captionTargetPageId="20" captionText="Figure 13. Miculissima excavata gen. et sp.nov.. A, dorsal view. B, frontal view.C, ventral view.D, lateral view. E, lateral view. Abbreviations:aca, antennal cavity; acl, antennal club; ga, galea; lp, labial palpus; ls, labral setae; mp, maxillary palpus; pl, prosternal lobe." pageId="20">Fig. 13D, E</figureCitation>
<bibRefCitation id="EFB44B59171C81624A6DF8D7FA117C81" author="Mazur" box="[1333,1457,1911,1935]" pageId="20" pageNumber="21" pagination="27 - 32" refId="ref18506" refString="Mazur º. New species of histerid beetles (Coleoptera, Histeridae). Annales Zoologici 1997; 38: 27 - 32. hup: // rcin. org. pl" type="journal article" year="1997">Mazur 1997</bibRefCitation>
). The excavate and sculptured pronotal hypomeron (
<figureCitation id="131E2A2D171C81624A1DF836FA177CA0" box="[1349,1463,1942,1966]" captionStart="Figure 13" captionStartId="20.[129,194,1346,1370]" captionTargetBox="[129,1473,144,1319]" captionTargetId="figure-333@20.[129,1473,144,1319]" captionTargetPageId="20" captionText="Figure 13. Miculissima excavata gen. et sp.nov.. A, dorsal view. B, frontal view.C, ventral view.D, lateral view. E, lateral view. Abbreviations:aca, antennal cavity; acl, antennal club; ga, galea; lp, labial palpus; ls, labral setae; mp, maxillary palpus; pl, prosternal lobe." pageId="20" pageNumber="21">Fig. 13D, E</figureCitation>
) is unlike any in recent
<taxonomicName id="4C254D2B171D81634E17FF30FE1A7BA6" box="[335,442,144,168]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="21" phylum="Arthropoda" rank="family">Histeridae</taxonomicName>
taxa. Ŋe phylogenetic analyses did not resolve the position of this taxon and placed it either as sister to all
<taxonomicName id="4C254D2B171D81634FB6FF6FFEF67BE9" baseAuthorityName="Wenzel" baseAuthorityYear="1944" box="[238,342,207,231]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="21" phylum="Arthropoda" rank="subFamily" subFamily="Abraeinae">Abraeinae</taxonomicName>
<taxonomicName id="4C254D2B171D81634E17FF30FE1A7BA6" box="[335,442,144,168]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="family">Histeridae</taxonomicName>
taxa. The phylogenetic analyses did not resolve the position of this taxon and placed it either as sister to all
<taxonomicName id="4C254D2B171D81634FB6FF6FFEF67BE9" baseAuthorityName="Wenzel" baseAuthorityYear="1944" box="[238,342,207,231]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subFamily" subFamily="Abraeinae">Abraeinae</taxonomicName>
or to all
<taxonomicName id="4C254D2B171D81634EE3FF6FFD867BE9" box="[443,550,207,231]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="21" phylum="Arthropoda" rank="family">Histeridae</taxonomicName>
<taxonomicName id="4C254D2B171D81634EE3FF6FFD867BE9" box="[443,550,207,231]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="family">Histeridae</taxonomicName>
excl.
<taxonomicName id="4C254D2B171D81634D3CFF6FFD6C7BE9" baseAuthorityName="Wenzel" baseAuthorityYear="1944" box="[612,716,207,231]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="21" phylum="Arthropoda" rank="subFamily" subFamily="Abraeinae">Abraeinae</taxonomicName>
<taxonomicName id="4C254D2B171D81634D3CFF6FFD6C7BE9" baseAuthorityName="Wenzel" baseAuthorityYear="1944" box="[612,716,207,231]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subFamily" subFamily="Abraeinae">Abraeinae</taxonomicName>
and
<taxonomicName id="4C254D2B171D81634F29FF4EFEA27A08" authorityName="Zhou, Yu-Lingxi, Caterino, Dong Ren &amp; Ślipiński" authorityYear="2020" box="[113,258,238,262]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="21" phylum="Arthropoda" rank="subFamily" subFamily="Antigracilinae">Antigracilinae</taxonomicName>
. Ŋus, a subfamily placement is unclear.
<taxonomicName id="4C254D2B171D81634F29FF4EFEA27A08" authorityName="Zhou, Yu-Lingxi, Caterino, Dong Ren &amp; Ślipiński" authorityYear="2020" box="[113,258,238,262]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subFamily" subFamily="Antigracilinae">Antigracilinae</taxonomicName>
. Thus, a subfamily placement is unclear.
</paragraph>
<paragraph id="8B9A36A8171D81634F29FE88FF507977" blockId="21.[113,763,295,633]" pageId="21">
<emphasis id="B951EABA171D81634F29FE88FF777A31" box="[113,215,296,319]" italics="true" pageId="21">Diagnosis:</emphasis>
Glabrous surface. Frons flat, wide, clypeus convex, labrum short and wide, with a pair of prominent setae laterally. Eyes convex. Pronotum widely angular posteriorly, with complete marginal stria and a lateral pronotal stria. Prosternal hypomeron widely excavate and irregularly structured, with a weakly defined antennal cavity. Tibiae simple, slightly widened, with several spines. Ŋe species may resemble small abraeines found in Burmese amber—herein described
<taxonomicName id="4C254D2B171D81634D22FDA3FF187934" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" class="Insecta" family="Histeridae" genus="Palaeabraeus" kingdom="Animalia" order="Coleoptera" pageId="21" phylum="Arthropoda" rank="species" species="glabrus">
<emphasis id="B951EABA171D81634D22FDA3FF187934" italics="true" pageId="21">Palaeabraeus glabrus</emphasis>
</subSubSection>
<subSubSection id="C33F6523171D81634F29FE88FF507977" pageId="21" pageNumber="22" type="diagnosis">
<paragraph id="8B9A36A8171D81634F29FE88FF507977" blockId="21.[113,763,295,633]" pageId="21" pageNumber="22">
<emphasis id="B951EABA171D81634F29FE88FF777A31" box="[113,215,296,319]" italics="true" pageId="21" pageNumber="22">Diagnosis:</emphasis>
Glabrous surface. Frons flat, wide, clypeus convex, labrum short and wide, with a pair of prominent setae laterally. Eyes convex. Pronotum widely angular posteriorly, with complete marginal stria and a lateral pronotal stria. Prosternal hypomeron widely excavate and irregularly structured, with a weakly defined antennal cavity. Tibiae simple, slightly widened, with several spines. The species may resemble small abraeines found in Burmese amber—herein described
<taxonomicName id="4C254D2B171D81634D22FDA3FF187934" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" class="Insecta" family="Histeridae" genus="Palaeabraeus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="glabrus">
<emphasis id="B951EABA171D81634D22FDA3FF187934" italics="true" pageId="21" pageNumber="22">Palaeabraeus glabrus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C254D2B171D81634FB1FD82FE1E7934" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[233,446,546,570]" class="Insecta" family="Histeridae" genus="Pseudacritus" kingdom="Animalia" order="Coleoptera" pageId="21" phylum="Arthropoda" rank="species" species="extinctus">
<emphasis id="B951EABA171D81634FB1FD82FE1E7934" box="[233,446,546,570]" italics="true" pageId="21">Pseudacritus extinctus</emphasis>
<taxonomicName id="4C254D2B171D81634FB1FD82FE1E7934" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[233,446,546,570]" class="Insecta" family="Histeridae" genus="Pseudacritus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="extinctus">
<emphasis id="B951EABA171D81634FB1FD82FE1E7934" box="[233,446,546,570]" italics="true" pageId="21" pageNumber="22">Pseudacritus extinctus</emphasis>
</taxonomicName>
. It is distinguishable from both by the more prolonged body shape and the strongly excavated hypomeron.
</paragraph>
<paragraph id="8B9A36A8171D81634F29FD3AFF5079DF" blockId="21.[113,762,666,721]" pageId="21">
<emphasis id="B951EABA171D81634F29FD3AFF4179BC" box="[113,225,666,690]" italics="true" pageId="21">Etymology:</emphasis>
Ŋe Latin adjective ‘excavata’ refers to the excavate hypomeron.
</subSubSection>
<subSubSection id="C33F6523171D81634F29FD3AFF5079DF" pageId="21" pageNumber="22" type="etymology">
<paragraph id="8B9A36A8171D81634F29FD3AFF5079DF" blockId="21.[113,762,666,721]" pageId="21" pageNumber="22">
<emphasis id="B951EABA171D81634F29FD3AFF4179BC" box="[113,225,666,690]" italics="true" pageId="21" pageNumber="22">Etymology:</emphasis>
The Latin adjective ‘excavata’ refers to the excavate hypomeron.
</paragraph>
<paragraph id="8B9A36A8171D81634E0CFD54FDB87800" blockId="21.[340,536,756,782]" box="[340,536,756,782]" pageId="21">
<emphasis id="B951EABA171D81634E0CFD54FDB87800" bold="true" box="[340,536,756,782]" pageId="21">Additional reports</emphasis>
</subSubSection>
<subSubSection id="C33F6523171D81634E0CFD54FC0F7BC9" pageId="21" pageNumber="22" type="discussion">
<paragraph id="8B9A36A8171D81634E0CFD54FDB87800" blockId="21.[340,536,756,782]" box="[340,536,756,782]" pageId="21" pageNumber="22">
<emphasis id="B951EABA171D81634E0CFD54FDB87800" bold="true" box="[340,536,756,782]" pageId="21" pageNumber="22">Additional reports</emphasis>
</paragraph>
<paragraph id="8B9A36A8171D81634F29FCBBFC0F7BC9" blockId="21.[113,762,795,850]" lastBlockId="21.[810,1459,144,199]" pageId="21">ºome additional newly discovered specimens have shed light on characters poorly or not at all visible in previously published reports, or have expanded the spatial and temporal range in the fossil record:</paragraph>
<paragraph id="8B9A36A8171D81634F29FCBBFC0F7BC9" blockId="21.[113,762,795,850]" lastBlockId="21.[810,1459,144,199]" pageId="21" pageNumber="22">Some additional newly discovered specimens have shed light on characters poorly or not at all visible in previously published reports, or have expanded the spatial and temporal range in the fossil record:</paragraph>
</subSubSection>
</treatment>
</document>

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<heading id="D0D281C4171B81654CACFF6EFB497BE7" bold="true" box="[1012,1257,206,233]" centered="true" fontSize="11" level="2" pageId="19" reason="2">
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<emphasis id="B951EABA171B81654CACFF6EFBDC7BE7" bold="true" box="[1012,1148,206,233]" italics="true" pageId="19">Miculissima</emphasis>
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species.
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..
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<emphasis id="B951EABA171B81654C72FEE9FC307A6E" box="[810,912,329,352]" italics="true" pageId="19">Diagnosis:</emphasis>
Tiny body size, body oval-shaped and glabrous. Frons flat, wide, clypeus convex, labrum short and wide, with a pair of prominent setae laterally. Antennal club round with two distinct annuli. Eyes convex. Pronotum widely angular posteriorly, with complete marginal stria and a lateral pronotal stria. ºcutellar shield tiny, triangular. Prosternal hypomeron widely excavate and irregularly structured, with a weakly defined antennal cavity. Prosternum with fissures for antennal passage. Prosternal process short, without prosternal lobe, slightly emarginate posteriorly. Tibiae simple, slightly widened, with several spines.
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<emphasis id="B951EABA171B81654C72FEE9FC307A6E" box="[810,912,329,352]" italics="true" pageId="19" pageNumber="20">Diagnosis:</emphasis>
Tiny body size, body oval-shaped and glabrous. Frons flat, wide, clypeus convex, labrum short and wide, with a pair of prominent setae laterally. Antennal club round with two distinct annuli. Eyes convex. Pronotum widely angular posteriorly, with complete marginal stria and a lateral pronotal stria. Scutellar shield tiny, triangular. Prosternal hypomeron widely excavate and irregularly structured, with a weakly defined antennal cavity. Prosternum with fissures for antennal passage. Prosternal process short, without prosternal lobe, slightly emarginate posteriorly. Tibiae simple, slightly widened, with several spines.
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<paragraph id="8B9A36A8171B81654C72FD3CFB3879DD" blockId="19.[810,1459,668,723]" pageId="19">
<emphasis id="B951EABA171B81654C72FD3CFC3979BA" box="[810,921,668,692]" italics="true" pageId="19">Etymology:</emphasis>
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<paragraph id="8B9A36A8171B81654C72FD3CFB3879DD" blockId="19.[810,1459,668,723]" pageId="19" pageNumber="20">
<emphasis id="B951EABA171B81654C72FD3CFC3979BA" box="[810,921,668,692]" italics="true" pageId="19" pageNumber="20">Etymology:</emphasis>
Feminine, derived from the Latin noun ‘mica’ which translates as crumb, grain or morsel.
</paragraph>
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169
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<emphasis id="B951EABA171D81634C2AFF4AFB8A7A0B" bold="true" box="[882,1066,234,261]" italics="true" pageId="21">Amplectister tenax</emphasis>
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Caterino and
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<figureCitation id="131E2A2D171D81634B6BFE81FB097A32" box="[1075,1193,289,316]" captionStart="Figure 14" captionStartId="21.[113,178,1926,1950]" captionTargetBox="[114,1458,920,1898]" captionTargetId="figure-333@21.[114,1458,920,1898]" captionTargetPageId="21" captionText="Figure 14. Additional structural details.A, Amplectister tenax. B, Carinumerus sp. C, Phasmister hkamticus. D, Abdominal cavity of Amplectister tenax. E, pronotum of Carinumerus sp.Abbreviations:as, abdominal setae; po, pronotal opening." pageId="21" pageNumber="22">Fig. 14A, D</figureCitation>
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Ŋe first abdominal ventrite is more clearly visible in this specimen (
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The first abdominal ventrite is more clearly visible in this specimen (
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:
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). Ŋe steep lateral flanges are partially formed by a very dense aggregation of long setae stuck together (
<figureCitation id="131E2A2D171D81634C6DFE65FC2E7AD3" box="[821,910,453,477]" captionStart="Figure 14" captionStartId="21.[113,178,1926,1950]" captionTargetBox="[114,1458,920,1898]" captionTargetId="figure-333@21.[114,1458,920,1898]" captionTargetPageId="21" captionText="Figure 14. Additional structural details.A, Amplectister tenax. B, Carinumerus sp. C, Phasmister hkamticus. D, Abdominal cavity of Amplectister tenax. E, pronotum of Carinumerus sp.Abbreviations:as, abdominal setae; po, pronotal opening." pageId="21">Fig. 14D</figureCitation>
<typeStatus id="549E880A171D81634C8CFE27FB967A91" box="[980,1078,391,415]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
). The steep lateral flanges are partially formed by a very dense aggregation of long setae stuck together (
<figureCitation id="131E2A2D171D81634C6DFE65FC2E7AD3" box="[821,910,453,477]" captionStart="Figure 14" captionStartId="21.[113,178,1926,1950]" captionTargetBox="[114,1458,920,1898]" captionTargetId="figure-333@21.[114,1458,920,1898]" captionTargetPageId="21" captionText="Figure 14. Additional structural details.A, Amplectister tenax. B, Carinumerus sp. C, Phasmister hkamticus. D, Abdominal cavity of Amplectister tenax. E, pronotum of Carinumerus sp.Abbreviations:as, abdominal setae; po, pronotal opening." pageId="21" pageNumber="22">Fig. 14D</figureCitation>
), whereas in the original description setae were mentioned as occurring only distally on these ridges (Caterino and
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.
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).
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<mods:namePart id="616791ACC4929EEFA2F1DD91A94125B3">Rai, Santosh Kumar</mods:namePart>
<mods:nameIdentifier id="F0E742DF440811092A03862B6165880A" type="ORCID">0000-0003-0821-2879</mods:nameIdentifier>
<mods:affiliation id="D2B604D001502F40E2A35BA2186B6F93">Taxonomy and Biodiversity Laboratory, Department of Botany, Sikkim University, 6 mile, Samdur, Gangtok, Sikkim (737102), India</mods:affiliation>
<mods:nameIdentifier id="3259598A8487C01AFF3C150B84BC9438" type="email">skrai@cus.ac.in</mods:nameIdentifier>
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<mods:role id="D43CDAAE467ED0B8FDDD0C9CB771C760">
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<mods:namePart id="88CC39717A1B94E1F525B0C59F3A9CD0">Kumar, Devendra</mods:namePart>
<mods:nameIdentifier id="908CD5A569F90EC94AD2989800CCC3D1" type="ORCID">0000-0001-7184-5455</mods:nameIdentifier>
<mods:affiliation id="518751FC8D503A1557A1E67A09EB6CD0">G. B. Pant National Institute of Himalayan Environment (GBPNIHE), North-East Regional Centre, Chandranagar, Itanagar (791113), Arunachal Pradesh, India</mods:affiliation>
<mods:nameIdentifier id="F79EA99BF7481BDB1C837111D8F9FB90" type="email">devendrawii@gmail.com</mods:nameIdentifier>
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<mods:namePart id="38D2BB14F6F880ACF0B1FA50623B4BF5">Rawat, Sandeep</mods:namePart>
<mods:nameIdentifier id="7A8661F7253BAABF41296FD5771CB322" type="ORCID">0000-0001-7991-2878</mods:nameIdentifier>
<mods:affiliation id="A1DA5DCC03D1BAE23455084803D03228">G. B. Pant National Institute of Himalayan Environment (GBPNIHE), Sikkim Regional Centre, Pangthang (737101), Gangtok, Sikkim, India</mods:affiliation>
<mods:nameIdentifier id="99005830617A93A0362D333C772E431C" type="email">Sandeep_rawat15@rediffmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="78A4E4586D153908A1F09BFBDA9DD2D4">text</mods:typeOfResource>
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<mods:titleInfo id="6FF78D1A34BC51993D010624E0FD7B36">
<mods:title id="E543727C10F8657D1DDBFC9A70F350DD">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="447013168355A7F3131395E4D55FE546">
<mods:date id="A988BE02C3D4EEE773E0A7AD906F3DD9">2024</mods:date>
<mods:detail id="D4120AECAF12C2E298567082EC038198" type="pubDate">
<mods:number id="970B40A6F9222E1AAAE3492E51626F79">2024-08-15</mods:number>
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<mods:detail id="6B51A0F55DF6896E8DEE29ADAA1B38D2" type="volume">
<mods:number id="D20CDF0E670130F7A847D5D7DBD76924">663</mods:number>
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<mods:number id="9458608AB4404DB02C697C6BF4CC04A1">1</mods:number>
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<mods:start id="F782341A94C4464E9EBF90F1FD255364">26</mods:start>
<mods:end id="046BA6CF10E4CFA2592846D827E92609">32</mods:end>
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<mods:location id="FC1BC82F149AA00E633C01E8E5939CE1">
<mods:url id="0931F2AEB56947B2F3886C7BE2179FD4">https://doi.org/10.11646/phytotaxa.663.1.3</mods:url>
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<mods:identifier id="E569481F3EE4A49DFDDEB0DF20BF5405" type="DOI">10.11646/phytotaxa.663.1.3</mods:identifier>
<mods:identifier id="CD67E216ABF9FAF02A24A44799CEB59A" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="B12AD476CE1934EA1D14CA501457E884" type="Zenodo-Dep">14515997</mods:identifier>
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<treatment id="03A25F44B81AFFF36480E6445BD0FF5C" ID-DOI="http://doi.org/10.5281/zenodo.14515999" ID-Zenodo-Dep="14515999" LSID="urn:lsid:plazi:treatment:03A25F44B81AFFF36480E6445BD0FF5C" httpUri="http://treatment.plazi.org/id/03A25F44B81AFFF36480E6445BD0FF5C" lastPageId="4" lastPageNumber="30" pageId="1" pageNumber="27">
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<taxonomicName id="4C0B95D1B81AFFF66480E64459EAFBA9" authority="M. Khanal" authorityName="M. Khanal" authorityYear="2024" box="[136,466,1044,1071]" class="Liliopsida" family="Orchidaceae" genus="Gastrodia" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="27" phylum="Tracheophyta" rank="species" species="indica" status="sp. nov.">
<emphasis id="B97F3240B81AFFF66480E6445971FBA8" bold="true" box="[136,329,1044,1071]" italics="true" pageId="1" pageNumber="27">Gastrodia indica</emphasis>
M. Khanal
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<emphasis id="B97F3240B81AFFF665D6E6445A17FBA9" bold="true" box="[478,559,1044,1070]" italics="true" pageId="1" pageNumber="27">sp. nov</emphasis>
.
</taxonomicNameLabel>
(
<figureCitation id="1330F2D7B81AFFF6664DE6445A89FBA9" box="[581,689,1044,1070]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[136,229,1384,1406]" captionStartId-1="3.[136,229,1952,1974]" captionTargetBox-0="[151,1436,187,1356]" captionTargetBox-1="[169,1418,190,1928]" captionTargetId-0="figure-18@2.[151,1436,187,1356]" captionTargetId-1="figure-24@3.[169,1418,190,1928]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. A. Habit photograph of the plant in flowering stage; B. Close up photos of a single flower; C. Fruiting stages." captionText-1="FIGURE 2. Photoplate of Gastrodia indica. A. Habit; B. Close up photo of rhizome; C–E. Flowers in different view; F. Floral bract; G1–G2. Opened up and spread perianth tube (Dorsal &amp; ventral view); H1–H3. Labellum in different view; I1. Column with labellum (Different view); I2 –I3. Column (Front and side view); J1 –J2. Anther Cap (Dorsal &amp; ventral view)" figureDoi-0="http://doi.org/10.5281/zenodo.14516001" figureDoi-1="http://doi.org/10.5281/zenodo.14516003" httpUri-0="https://zenodo.org/record/14516001/files/figure.png" httpUri-1="https://zenodo.org/record/14516003/files/figure.png" pageId="1" pageNumber="27">Figs. 1–2</figureCitation>
)
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Type:—
<taxonomicName id="4C0B95D1B81AFFF664D2E60F591BFBF3" box="[218,291,1119,1140]" class="Liliopsida" family="Orchidaceae" genus="India" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="32" phylum="Tracheophyta" rank="genus">INDIA</taxonomicName>
.
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<materialsCitation id="3B63E40FB81AFFF66538E63058C1FB1E" collectionCode="M, CAL" elevation="2015" latitude="27.365046" location="Pangthang" longLatPrecision="1" longitude="88.56835" municipality="Gangtok" pageId="1" pageNumber="27" specimenCount="1" stateProvince="Sikkim" typeStatus="holotype">
<collectingRegion id="49CF20B0B81AFFF66538E6305941FBF1" box="[304,377,1120,1142]" country="India" name="Sikkim" pageId="1" pageNumber="27">Sikkim</collectingRegion>
:
<collectingMunicipality id="6BD07428B81AFFF6658DE63059E2FBF1" box="[389,474,1120,1142]" pageId="1" pageNumber="27">Gangtok</collectingMunicipality>
,
<location id="8ED4B889B81AFFF665EEE6305A76FBF1" LSID="urn:lsid:plazi:treatment:03A25F44B81AFFF36480E6445BD0FF5C:8ED4B889B81AFFF665EEE6305A76FBF1" box="[486,590,1120,1142]" latitude="27.365046" longLatPrecision="1" longitude="88.56835" municipality="Gangtok" name="Pangthang" pageId="1" pageNumber="27" stateProvince="Sikkim">Pangthang</location>
,
<quantity id="4CF343B7B81AFFF66653E6305A91FBF2" box="[603,681,1120,1142]" metricMagnitude="3" metricUnit="m" metricValue="2.0149999999999997" pageId="1" pageNumber="27" unit="m" value="2015.0">
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</quantity>
, 21.09.202 3,
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<collectionCode id="ED1A7697B81AFFF66080E6305CA7FBF2" box="[1160,1183,1120,1141]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="1" pageNumber="27" type="Herbarium">M</collectionCode>
. Khanal
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90345a [
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<emphasis id="B97F3240B81AFFF6650AE6D45951FB1E" box="[258,361,1156,1177]" italics="true" pageId="1" pageNumber="27">
<collectionCode id="ED1A7697B81AFFF6650AE6D45921FB1E" box="[258,281,1156,1177]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="1" pageNumber="27" type="Herbarium">M</collectionCode>
. Khanal
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90345b [
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<collectionCode id="ED1A7697B81AFFF66619E6D45A6DFB1E" box="[529,597,1156,1177]" country="India" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12697" lsid="urn:lsid:biocol.org:col:12697" name="Botanical Survey of India, Sikkim Himalayan Circle" pageId="1" pageNumber="27" type="Herbarium">BSHC</collectionCode>
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;
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23.09.202 3,
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<collectionCode id="ED1A7697B81AFFF666D5E6D45ACCFB1E" box="[733,756,1156,1177]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="1" pageNumber="27" type="Herbarium">M</collectionCode>
. Khanal
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90346 [
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<collectingRegion id="49CF20B0B81AFFF667E5E6D45C0AFB1D" box="[1005,1074,1156,1178]" country="India" name="Sikkim" pageId="1" pageNumber="27">Sikkim</collectingRegion>
University Herbarium]
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.
</paragraph>
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<subSubSection id="C311BDD9B81AFFF66480E69859A9F87D" pageId="1" pageNumber="27" type="description">
<paragraph id="8BB4EE52B81AFFF66480E69859A9F87D" blockId="1.[136,1452,1224,2042]" pageId="1" pageNumber="27">
<emphasis id="B97F3240B81AFFF66480E6985924FB65" bold="true" box="[136,284,1224,1250]" pageId="1" pageNumber="27">Description:</emphasis>
Holomycotrophic terrestrial herb,
<quantity id="4CF343B7B81AFFF66694E6985AD8FB64" box="[668,736,1224,1251]" metricMagnitude="-1" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="27" unit="cm" value="45.0">45 cm</quantity>
to more than
<quantity id="4CF343B7B81AFFF6677FE6985B98FB64" box="[887,928,1224,1251]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="27" unit="m" value="1.0">1 m</quantity>
tall. Rhizome 4.2–5.5 ×
<quantity id="4CF343B7B81AFFF660B8E6985D22FB64" box="[1200,1306,1224,1251]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="1" pageNumber="27" unit="cm" value="1.15" valueMax="1.3" valueMin="1.0">1–1.3 cm</quantity>
, horizontally placed, brownish, tuberous, fleshy, cylindrical, sparsely to densely pubescent with minute triangular scales with ciliate margins. Roots
<quantity id="4CF343B7B81AFFF66533E7405997FAAC" box="[315,431,1296,1323]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="1" pageNumber="27" unit="mm" value="1.05" valueMax="1.1" valueMin="1.0">1–1.1 mm</quantity>
dia., slender, filliform, few, arising from the base of the rhizome, glabrous. Inflorescence racemose, up to
<quantity id="4CF343B7B81AFFF66548E76459BDFAC8" box="[320,389,1332,1359]" metricMagnitude="-1" metricUnit="m" metricValue="9.4" pageId="1" pageNumber="27" unit="cm" value="94.0">94 cm</quantity>
tall, erect, glabrous; peduncle
<quantity id="4CF343B7B81AFFF666D7E7645B76FAC8" box="[735,846,1332,1359]" metricMagnitude="-1" metricUnit="m" metricValue="4.75" metricValueMax="6.0" metricValueMin="3.5" pageId="1" pageNumber="27" unit="cm" value="47.5" valueMax="60.0" valueMin="35.0">35–60 cm</quantity>
tall, erect, stout, 4–7 nodded towards base, each node enclosed with single membranous sheathing bract, sheathing bracts 0.5–0.6 ×
<quantity id="4CF343B7B81AFFF667E5E7085C60FAF4" box="[1005,1112,1368,1395]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="1" pageNumber="27" unit="cm" value="0.95" valueMax="1.0" valueMin="0.9">0.9–1 cm</quantity>
, glabrous, 7–10 veined; rachis
<quantity id="4CF343B7B81AFFF66480E72C58C3FA10" box="[136,251,1404,1431]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="1" pageNumber="27" unit="cm" value="20.0" valueMax="30.0" valueMin="10.0">10–30 cm</quantity>
tall, with laxly arranged 7–22 flowers. Floral bracts 0.6–0.7 ×
<quantity id="4CF343B7B81AFFF667E9E72C5C5CFA10" box="[993,1124,1404,1431]" metricMagnitude="-3" metricUnit="m" metricValue="3" metricValueMax="4.0" metricValueMin="2.0" pageId="1" pageNumber="27" unit="cm" value="0.3" valueMax="0.4" valueMin="0.2">0.2–0.4 cm</quantity>
, brownish, ovate-lanceolate, apex acute, entire, slightly cymbiform, 3-veined, glabrous, caducous. Flowers
<quantity id="4CF343B7B81AFFF66010E7F05CA0FA3C" box="[1048,1176,1440,1467]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.1" metricValueMin="1.7" pageId="1" pageNumber="27" unit="cm" value="1.9" valueMax="2.1" valueMin="1.7">1.7–2.1 cm</quantity>
long, erect, completely cleistogamous, brownish, urceolate, sparsely verrucose throughout the outer surface, slightly recurved, foul smelling when dissected; sepals brownish, petals whitish-brown, labellum &amp; column white. Pedicel and ovary 0.8–1.1 × 0.2–
<quantity id="4CF343B7B81AFFF66480E45C58D1F9A0" box="[136,233,1548,1575]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="27" unit="cm" value="0.25">0.25 cm</quantity>
, brownish, ovary portion slightly ribbed, pedicel not elongating during capsule maturation. Dorsal sepal 0.8–1.1 ×
<quantity id="4CF343B7B81AFFF664F4E46059B3F9CC" box="[252,395,1584,1611]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="1" pageNumber="27" unit="cm" value="0.375" valueMax="0.4" valueMin="0.35">0.35–0.4 cm</quantity>
, obtuse, undulate to entire, slightly incurved and thickened, slightly verrucose, little broarder than the lateral sepals, connate 4/5
<superScript id="7C7E431AB81AFFF6660EE4045A2BF9E5" attach="left" box="[518,531,1620,1634]" fontSize="6" pageId="1" pageNumber="27">th</superScript>
to petals, glabrous. Lateral sepals 0.8–1.1 ×
<quantity id="4CF343B7B81AFFF6600AE4045CB9F9E8" box="[1026,1153,1620,1647]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="1" pageNumber="27" unit="cm" value="0.25" valueMax="0.3" valueMin="0.2">0.2–0.3 cm</quantity>
, obtuse, entire to undulate, connate with each other for more than 3/4
<superScript id="7C7E431AB81AFFF66661E4285A4EF901" attach="left" box="[617,630,1656,1670]" fontSize="6" pageId="1" pageNumber="27">th</superScript>
of their length, slightly thickened, sparsely verrucose outside, glabrous. Petals ca. 0.1 ×
<quantity id="4CF343B7B81AFFF6653EE4CC598DF930" box="[310,437,1692,1719]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="1" pageNumber="27" unit="cm" value="0.15" valueMax="0.2" valueMin="0.1">0.1–0.2 cm</quantity>
, ovate to oblong with crisped margins, obtuse, membranous. Labellum 0.35–0.4 ×
<quantity id="4CF343B7B81AFFF66141E4CC5895F95C" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="1" pageNumber="27" unit="cm" value="0.175" valueMax="0.2" valueMin="0.15">0.15–0.2 cm</quantity>
, ovate, obtuse, glabrous, margins crisped to irregularly denticulate and slightly reflexed inwards, 5-nerved, base with short stalk attached to the base of coulmn; disc with 4 keels, a pair of centrally placed whitish keel arising from the hypochile which raised abruptly towards apex and other pair of shorter adjacent keel placed laterally &amp; base of the hypochile with a pair of whitish, globose subsessile calli. Column 0.4–0.45 ×
<quantity id="4CF343B7B81AFFF667FCE57C5CBAF8C0" box="[1012,1154,1836,1863]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="1" pageNumber="27" unit="cm" value="0.11" valueMax="0.12" valueMin="0.1">0.1–0.12 cm</quantity>
, slightly bent, cymbiform, glabrous; columnar wings slightly broader at the middle gradually narrowing towards the truncate apex and slightly curved inwards terminating into a pair of shortly pointed stelidia which is parallel to the anther cap; column foot absent. Rostellum absent. Stigma glossy, placed at the base of the column. Anther cap 2-chambered, dome shaped, 0.6–0.7 ×
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. Pollinia 2, globose, mealy to granular, whitish. Capsule 0.8–1.3 ×
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, attached to a non-elongating pedicel.
</paragraph>
</subSubSection>
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<emphasis id="B97F3240B819FFF56480E73858C6FAFA" bold="true" box="[136,254,1384,1406]" pageId="2" pageNumber="28">FIGURE 1.</emphasis>
A. Habit photograph of the plant in flowering stage; B. Close up photos of a single flower; C. Fruiting stages.
</paragraph>
</caption>
<subSubSection id="C311BDD9B819FFF564B5E7FC5B58F989" pageId="2" pageNumber="28" type="etymology">
<paragraph id="8BB4EE52B819FFF564B5E7FC5ADEFA6D" blockId="2.[136,1452,1452,1910]" pageId="2" pageNumber="28">
<emphasis id="B97F3240B819FFF564B5E7FC5970FA41" bold="true" box="[189,328,1452,1478]" pageId="2" pageNumber="28">Etymology:</emphasis>
The new species is named after the country of discovery—
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, as it is the first cleistogamous species of the genus
<taxonomicName id="4C0B95D1B819FFF56578E78059D9FA6D" authorityName="R. Brown" authorityYear="1810" box="[368,481,1488,1514]" class="Liliopsida" family="Orchidaceae" genus="Gastrodia" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="28" phylum="Tracheophyta" rank="genus">
<emphasis id="B97F3240B819FFF56578E78059D9FA6D" box="[368,481,1488,1514]" italics="true" pageId="2" pageNumber="28">Gastrodia</emphasis>
</taxonomicName>
discovered from
<taxonomicName id="4C0B95D1B819FFF566ADE7805ADAFA6D" box="[677,738,1488,1514]" class="Liliopsida" family="Orchidaceae" genus="India" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="32" phylum="Tracheophyta" rank="genus">India</taxonomicName>
.
</paragraph>
<paragraph id="8BB4EE52B819FFF564B5E7A45B58F989" blockId="2.[136,1452,1452,1910]" box="[189,864,1524,1551]" pageId="2" pageNumber="28">
<emphasis id="B97F3240B819FFF564B5E7A4597BF989" bold="true" box="[189,323,1524,1550]" pageId="2" pageNumber="28">Phenology:</emphasis>
Flowering and fruiting—September to October.
</paragraph>
</subSubSection>
<subSubSection id="C311BDD9B819FFF364B5E4485BD0FF5C" lastPageId="4" lastPageNumber="30" pageId="2" pageNumber="28" type="distribution">
<paragraph id="8BB4EE52B819FFF564B5E4485AB7F9B4" blockId="2.[136,1452,1452,1910]" box="[189,655,1560,1587]" pageId="2" pageNumber="28">
<emphasis id="B97F3240B819FFF564B5E4485961F9B5" bold="true" box="[189,345,1560,1586]" pageId="2" pageNumber="28">Distribution:</emphasis>
<taxonomicName id="4C0B95D1B819FFF56568E4485A3DF9B5" box="[352,517,1560,1586]" class="Liliopsida" family="Orchidaceae" genus="India" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="28" phylum="Tracheophyta" rank="subGenus" subGenus="Sikkim">India (Sikkim)</taxonomicName>
[
<emphasis id="B97F3240B819FFF5661CE4485A46F9B5" bold="true" box="[532,638,1560,1586]" pageId="2" pageNumber="28">Endemic</emphasis>
].
</paragraph>
<paragraph id="8BB4EE52B819FFF564B5E46C5967F919" blockId="2.[136,1452,1452,1910]" pageId="2" pageNumber="28">
<emphasis id="B97F3240B819FFF564B5E46C591BF9D1" bold="true" box="[189,291,1596,1622]" pageId="2" pageNumber="28">Habitat:</emphasis>
Holomycotroph, found growing on dense rotten leaf litter in temperate deciduous oak forest associated with
<taxonomicName id="4C0B95D1B819FFF564C9E43059ADF9FD" box="[193,405,1632,1658]" class="Magnoliopsida" family="Magnoliaceae" genus="Magnolia" kingdom="Plantae" order="Magnoliales" pageId="2" pageNumber="28" phylum="Tracheophyta" rank="species" species="doltsopa">
<emphasis id="B97F3240B819FFF564C9E43059ADF9FD" box="[193,405,1632,1658]" italics="true" pageId="2" pageNumber="28">Magnolia doltsopa</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0B95D1B819FFF565ACE4315A76F9FD" box="[420,590,1632,1659]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" order="Sapindales" pageId="2" pageNumber="28" phylum="Tracheophyta" rank="species" species="campbelli">
<emphasis id="B97F3240B819FFF565ACE4315A76F9FD" box="[420,590,1632,1659]" italics="true" pageId="2" pageNumber="28">Acer campbelli</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0B95D1B819FFF56655E4315B17F9FD" box="[605,815,1632,1659]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="28" phylum="Tracheophyta" rank="species" species="lamellosa">
<emphasis id="B97F3240B819FFF56655E4315B17F9FD" box="[605,815,1632,1659]" italics="true" pageId="2" pageNumber="28">Quercus lamellosa</emphasis>
</taxonomicName>
&amp;
<taxonomicName id="4C0B95D1B819FFF5675BE4305C4DF9FD" box="[851,1141,1632,1658]" class="Magnoliopsida" family="Fagaceae" genus="Lithocarpus" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="28" phylum="Tracheophyta" rank="species" species="pachyphyllus">
<emphasis id="B97F3240B819FFF5675BE4305C4DF9FD" box="[851,1141,1632,1658]" italics="true" pageId="2" pageNumber="28">Lithocarpus pachyphyllus</emphasis>
</taxonomicName>
; elevation of
<quantity id="4CF343B7B819FFF56118E4305D94F9FC" box="[1296,1452,1632,1659]" metricMagnitude="3" metricUnit="m" metricValue="2.025" metricValueMax="2.1" metricValueMin="1.95" pageId="2" pageNumber="28" unit="m" value="2025.0" valueMax="2100.0" valueMin="1950.0">1950–2100 m</quantity>
above msl. (Fig-1).
</paragraph>
<paragraph id="8BB4EE52B819FFF564B5E4F858D9F8F1" blockId="2.[136,1452,1452,1910]" pageId="2" pageNumber="28">
<emphasis id="B97F3240B819FFF564B5E4F8598AF945" bold="true" box="[189,434,1704,1730]" pageId="2" pageNumber="28">Conservation status:</emphasis>
Less than 100 individuals in flowering and fruiting stages were observed in the
<typeStatus id="54B050F0B819FFF5612AE4F95D6AF944" box="[1314,1362,1705,1731]" pageId="2" pageNumber="28">type</typeStatus>
habitat. As the
<typeStatus id="54B050F0B819FFF564D6E49D5936F960" box="[222,270,1741,1767]" pageId="2" pageNumber="28">type</typeStatus>
habitat is a long term ecological monitoring site, hence a limited anthropogenic disturbance probably resulted in a luxuriant growth of this extremely elusive species. Similar habitats in and around the
<typeStatus id="54B050F0B819FFF560E9E4A15D29F88C" box="[1249,1297,1777,1803]" pageId="2" pageNumber="28">type</typeStatus>
locality were surveyed but no population could be traced. It can be assumed that the new species being a holomycotroph is extremely sensitive to any anthropogenic pressure, hence any disturbance or changes in habitat could result in the loss of the species.
</paragraph>
<caption id="DF74BEDAB818FFF46480E5F05C70F878" ID-DOI="http://doi.org/10.5281/zenodo.14516003" ID-Zenodo-Dep="14516003" httpUri="https://zenodo.org/record/14516003/files/figure.png" pageId="3" pageNumber="29" startId="3.[136,229,1952,1974]" targetBox="[169,1418,190,1928]" targetPageId="3" targetType="figure">
<paragraph id="8BB4EE52B818FFF46480E5F05C70F878" blockId="3.[136,1452,1952,2047]" pageId="3" pageNumber="29">
<emphasis id="B97F3240B818FFF46480E5F05938F832" bold="true" box="[136,256,1952,1974]" pageId="3" pageNumber="29">FIGURE 2.</emphasis>
Photoplate of
<taxonomicName id="4C0B95D1B818FFF4659AE5F15A0EF831" authorityName="M. Khanal" authorityYear="2024" box="[402,566,1953,1974]" class="Liliopsida" family="Orchidaceae" genus="Gastrodia" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="29" phylum="Tracheophyta" rank="species" species="indica">
<emphasis id="B97F3240B818FFF4659AE5F15A0EF831" box="[402,566,1953,1974]" italics="true" pageId="3" pageNumber="29">Gastrodia indica</emphasis>
</taxonomicName>
. A. Habit; B. Close up photo of rhizome; C–E. Flowers in different view; F. Floral bract; G1–G2. Opened up and spread perianth tube (Dorsal &amp; ventral view); H1–H3. Labellum in different view; I1. Column with labellum (Different view); I2 –I3. Column (Front and side view); J1 –J2. Anther Cap (Dorsal &amp; ventral view)
</paragraph>
</caption>
<paragraph id="8BB4EE52B81FFFF36480E2CF5BD0FF5C" blockId="4.[136,1451,159,219]" pageId="4" pageNumber="30">
<emphasis id="B97F3240B81FFFF36480E2CF5957FF30" bold="true" box="[136,367,159,183]" pageId="4" pageNumber="30">Species comparison:</emphasis>
<emphasis id="B97F3240B81FFFF3657CE2CF5BD0FF5C" italics="true" pageId="4" pageNumber="30">
<taxonomicName id="4C0B95D1B81FFFF3657CE2CF5A56FF30" box="[372,622,159,183]" class="Liliopsida" family="Orchidaceae" genus="Gastrodia" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="30" phylum="Tracheophyta" rank="subSpecies" species="indica" subSpecies="shows">Gastrodia indica shows</taxonomicName>
morphological affinities with
<taxonomicName id="4C0B95D1B81FFFF367A5E2F05C3CFF30" authorityName="Hooker" authorityYear="1890" box="[941,1028,159,183]" class="Liliopsida" family="Orchidaceae" genus="Gastrodia" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="30" phylum="Tracheophyta" rank="species" species="exilis">G. exilis</taxonomicName>
and
<taxonomicName id="4C0B95D1B81FFFF3603FE2F05C81FF30" authorityName="King &amp; Pantling" authorityYear="1896" box="[1079,1209,159,183]" class="Liliopsida" family="Orchidaceae" genus="Gastrodia" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="30" phylum="Tracheophyta" rank="species" species="dyeriana">G. dyeriana</taxonomicName>
. Detailed comparisons between them have been provided in the table below to aid in easy identification.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

300
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<mods:title id="0E2132CC150B63C2FBDA096A9E23252A">Curcuma ungmensis (Zingiberaceae): a new species from Nagaland, North-East India</mods:title>
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<mods:affiliation id="F72F43A5E5A86B73CC7DDA9B92BD78DF">KSCSTE-Malabar Botanical Garden and Institute for Plant Sciences, Calicut- 673014, Kerala, India &amp; Department of Botany, Fazl Ali College, Mokokchung- 798 601, Nagaland, India &amp; KSCSTE-Malabar Botanical Garden and Institute for Plant Sciences, Calicut- 673014, Kerala, India</mods:affiliation>
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<taxonomicName id="4C614D34FF80D604FF7DFABDA2B4E52B" authority="M.Sabu, Hareesh and P.Tiatemsu" authorityName="M. Sabu, Hareesh and P. Tiatemsu" authorityYear="2024" box="[136,755,1378,1404]" class="Liliopsida" family="Zingiberaceae" genus="Curcuma" kingdom="Plantae" order="Zingiberales" pageId="1" pageNumber="91" phylum="Tracheophyta" rank="species" species="ungmensis" status="sp. nov.">
<emphasis id="B915EAA5FF80D604FF7DFABDA13FE52B" bold="true" box="[136,376,1378,1404]" italics="true" pageId="1" pageNumber="91">Curcuma ungmensis</emphasis>
M.Sabu, Hareesh and P.Tiatemsu
</taxonomicName>
<emphasis id="B915EAA5FF80D604FD0CFABCA309E52B" box="[761,846,1379,1404]" italics="true" pageId="1" pageNumber="91">
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</emphasis>
(
<figureCitation id="135A2A32FF80D604FCA8FABDA3DAE52B" box="[861,925,1378,1404]" captionStart="FIGURE 1" captionStartId="2.[136,234,1991,2013]" captionTargetBox="[287,1300,530,1963]" captionTargetId="figure-192@2.[287,1300,530,1963]" captionTargetPageId="2" captionText="FIGURE 1. Curcuma ungmensis sp. nov. A. Habit; B. Inflorescence; C. Rhizome; D. C.S. of rhizome; E. Fertile bract; F. Coma bract; G. Bracteole; H. Flower; I. Calyx; J. Dorsal corolla lobe; K1 &amp; K2. Lateral corolla lobes; L1 &amp; L2. Lateral staminodes; M. Labellum; N. Anther lateral-view; O. Anther front-view; P. Ovary with epigynous glands; Q. Ovary. (Photos by M. Sabu &amp; V.S. Hareesh)." figureDoi="http://doi.org/10.5281/zenodo.14515945" httpUri="https://zenodo.org/record/14515945/files/figure.png" pageId="1" pageNumber="91">Fig. 1</figureCitation>
)
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<subSubSection id="C37B653CFF80D604FF7DFA71A1FAE65C" pageId="1" pageNumber="91" type="reference_group">
<paragraph id="8BDE36B7FF80D604FF7DFA71A1FAE65C" blockId="1.[136,1452,1453,1619]" pageId="1" pageNumber="91">
<taxonomicName id="4C614D34FF80D604FF7DFA71A14FE594" authorityName="M. Sabu, Hareesh and P. Tiatemsu" authorityYear="2024" box="[136,264,1454,1475]" class="Liliopsida" family="Zingiberaceae" genus="Curcuma" kingdom="Plantae" order="Zingiberales" pageId="1" pageNumber="91" phylum="Tracheophyta" rank="species" species="ungmensis">
<emphasis id="B915EAA5FF80D604FF7DFA71A14FE594" box="[136,264,1454,1475]" italics="true" pageId="1" pageNumber="91">C. ungmensis</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C614D34FF80D604FE71FA71A2E0E594" ID-CoL="32JZQ" authority="Blatter (1931: 357)" authorityName="Blatter" authorityPageNumber="357" authorityYear="1931" box="[388,679,1453,1475]" class="Liliopsida" family="Zingiberaceae" genus="Curcuma" kingdom="Plantae" order="Zingiberales" pageId="1" pageNumber="91" phylum="Tracheophyta" rank="species" species="inodora">
<emphasis id="B915EAA5FF80D604FE71FA71A1ACE595" box="[388,491,1453,1475]" italics="true" pageId="1" pageNumber="91">C. inodora</emphasis>
<bibRefCitation id="EFF04B46FF80D604FE05FA72A2E0E594" author="Blatter, E." box="[496,679,1453,1475]" pageId="1" pageNumber="91" refId="ref2593" refString="Blatter, E. (1931) Curcuma inodora. Journal and Proceedings of the Asiatic Society of Bengal 26: 357. [Calcutta]" type="journal volume" year="1931">Blatter (1931: 357)</bibRefCitation>
</taxonomicName>
in the general habit, white bracts with purple tinge at the tip, purple corolla lobes and yellow flowers but differs from the latter by its large branched rhizome (vs. small unbranched, ovoid), and flowers up to
<quantity id="4C999B52FF80D604FA9FFA0EA5ECE5B0" box="[1386,1451,1489,1511]" metricMagnitude="-2" metricUnit="m" metricValue="5.9" pageId="1" pageNumber="91" unit="cm" value="5.9">5.9 cm</quantity>
long (vs. up to
<quantity id="4C999B52FF80D604FEBBFA29A13AE65C" box="[334,381,1526,1547]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="91" unit="cm" value="4.0">4 cm</quantity>
long).
</paragraph>
</subSubSection>
<subSubSection id="C37B653CFF80D606FF7DF9C5A4AFE08B" lastPageId="3" lastPageNumber="93" pageId="1" pageNumber="91" type="materials_examined">
<paragraph id="8BDE36B7FF80D604FF7DF9C5A4FEE604" blockId="1.[136,1452,1453,1619]" pageId="1" pageNumber="91">
<emphasis id="B915EAA5FF80D604FF7DF9C5A0FDE678" bold="true" box="[136,186,1562,1583]" pageId="1" pageNumber="91">Type</emphasis>
:—
<materialsCitation id="3B093CEAFF80D604FF22F9C5A4F2E604" collectingDate="2023-09-03" collectionCode="M, V, S, P, MBGH" country="India" county="Mokokchung district" elevation="1173" latitude="26.304167" location="Ungma Village" longLatPrecision="20" longitude="94.50861" municipality="Tuensang" pageId="1" pageNumber="91" specimenCount="2" stateProvince="Nagaland" typeStatus="holotype">
<collectingCountry id="F3767627FF80D604FF22F9C5A167E678" box="[215,288,1562,1583]" name="India" pageId="1" pageNumber="91">INDIA</collectingCountry>
.
<collectingRegion id="49A5F855FF80D604FEDCF9C6A1CEE678" box="[297,393,1561,1583]" country="India" name="Nagaland" pageId="1" pageNumber="91">Nagaland</collectingRegion>
:
<collectingCounty id="62BF4E3BFF80D604FE67F9C6A21AE678" box="[402,605,1561,1583]" pageId="1" pageNumber="91">Mokokchung district</collectingCounty>
, three kms towards
<collectingMunicipality id="6BBAACCDFF80D604FCE8F9C5A33BE678" box="[797,892,1562,1583]" pageId="1" pageNumber="91">Tuensang</collectingMunicipality>
,
<location id="8EBE606CFF80D604FC73F9C5A45CE678" LSID="urn:lsid:plazi:treatment:03C887A1FF80D606FF7DFABDA512E2AF:8EBE606CFF80D604FC73F9C5A45CE678" box="[902,1051,1561,1583]" country="India" county="Mokokchung district" latitude="26.304167" longLatPrecision="20" longitude="94.50861" municipality="Tuensang" name="Ungma Village" pageId="1" pageNumber="91" stateProvince="Nagaland">Ungma Village</location>
,
<geoCoordinate id="EE555070FF80D604FBD1F9C6A4EFE678" box="[1060,1192,1561,1583]" degrees="26" direction="north" minutes="18" orientation="latitude" pageId="1" pageNumber="91" precision="15" seconds="15" value="26.304167">26° 18' 15'' N</geoCoordinate>
,
<geoCoordinate id="EE555070FF80D604FB46F9C5A572E678" box="[1203,1333,1562,1583]" degrees="94" direction="east" minutes="30" orientation="longitude" pageId="1" pageNumber="91" precision="15" seconds="31" value="94.50861">94° 30' 31'' E</geoCoordinate>
,
<quantity id="4C999B52FF80D604FACAF9C5A5CEE678" box="[1343,1417,1562,1583]" metricMagnitude="3" metricUnit="m" metricValue="1.173" pageId="1" pageNumber="91" unit="m" value="1173.0">
<elevation id="004CD184FF80D604FACAF9C5A5CEE678" box="[1343,1417,1562,1583]" metricMagnitude="3" metricUnit="m" metricValue="1.173" pageId="1" pageNumber="91" unit="m" value="1173.0">1173 m</elevation>
</quantity>
,
<date id="FFDF1077FF80D604FA66F9C5A11BE604" pageId="1" pageNumber="91" value="2023-09-03">
<collectingDate id="EF9BE99FFF80D604FA66F9C5A11BE604" pageId="1" pageNumber="91" value="2023-09-03">03 September 2023</collectingDate>
</date>
,
<emphasis id="B915EAA5FF80D604FE93F9E1A317E604" box="[358,848,1597,1619]" italics="true" pageId="1" pageNumber="91">
<collectionCode id="ED70AE72FF80D604FE93F9E1A13AE604" box="[358,381,1598,1619]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="1" pageNumber="91" type="Herbarium">M</collectionCode>
. Sabu,
<collectionCode id="ED70AE72FF80D604FE37F9E1A195E604" box="[450,466,1598,1619]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="91" type="Museum">V</collectionCode>
.
<collectionCode id="ED70AE72FF80D604FE20F9E1A1A5E604" box="[469,482,1598,1619]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="1" pageNumber="91" type="Herbarium">S</collectionCode>
. Hareesh &amp;
<collectionCode id="ED70AE72FF80D604FDAEF9E1A22DE604" box="[603,618,1598,1619]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="1" pageNumber="91" type="Herbarium">P</collectionCode>
.Tiatemsu 19599
<collectionCode id="ED70AE72FF80D604FCFEF9E1A317E604" box="[779,848,1598,1619]" pageId="1" pageNumber="91">MBGH</collectionCode>
</emphasis>
(
<typeStatus id="54DA8815FF80D604FCA8F9E2A3F4E604" box="[861,947,1597,1619]" pageId="1" pageNumber="91" type="holotype">holotype</typeStatus>
<collectionCode id="ED70AE72FF80D604FC4DF9E1A444E604" box="[952,1027,1598,1619]" pageId="1" pageNumber="91">MBGH</collectionCode>
;
<typeStatus id="54DA8815FF80D604FBF8F9E2A415E604" box="[1037,1106,1597,1619]" pageId="1" pageNumber="91" type="isotype">isotype</typeStatus>
<collectingRegion id="49A5F855FF80D604FBA2F9E1A4E9E604" box="[1111,1198,1598,1619]" country="India" name="Assam" pageId="1" pageNumber="91">ASSAM</collectingRegion>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BDE36B7FF80D607FF7DF95DA26AE1AA" blockId="1.[136,1452,1666,2089]" lastBlockId="2.[136,1452,158,509]" lastPageId="2" lastPageNumber="92" pageId="1" pageNumber="91">
Rhizomatous herb,
<quantity id="4C999B52FF80D604FE95F95DA18AE6CB" box="[352,461,1666,1693]" metricMagnitude="-1" metricUnit="m" metricValue="7.75" metricValueMax="9.0" metricValueMin="6.5" pageId="1" pageNumber="91" unit="cm" value="77.5" valueMax="90.0" valueMin="65.0">65–90 cm</quantity>
tall. Rhizome with lateral branches, 12–13.5 ×
<quantity id="4C999B52FF80D604FC22F95DA46BE6CB" box="[983,1068,1666,1693]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="1" pageNumber="91" unit="cm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 cm</quantity>
, deeply buried in the ground, older rhizomes brown externally, younger ones white, glabrous, sheathed by papery scales, white or pale yellow internally, faintly aromatic, fleshy roots end in ovoid root tubers. Leafy shoot
<quantity id="4C999B52FF80D604FC80F915A3A3E6B3" box="[885,996,1738,1765]" metricMagnitude="-1" metricUnit="m" metricValue="5.75" metricValueMax="8.0" metricValueMin="3.5" pageId="1" pageNumber="91" unit="cm" value="57.5" valueMax="80.0" valueMin="35.0">35–80 cm</quantity>
long, leaves
<quantity id="4C999B52FF80D604FB82F915A4FAE6B3" box="[1143,1213,1738,1764]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="7.62" metricValueMin="5.08" pageId="1" pageNumber="91" unit="in" value="2.5" valueMax="3.0" valueMin="2.0">2–3 in</quantity>
number. Pseudostem
<quantity id="4C999B52FF80D604FF7DF931A0B0E75F" box="[136,247,1774,1800]" metricMagnitude="-1" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="1" pageNumber="91" unit="cm" value="11.5" valueMax="13.0" valueMin="10.0">10–13 cm</quantity>
long, deeply buried in the soil, formed by leaf sheaths and 2–3 sheathing bracts; ligule absent or very short; petiole
<quantity id="4C999B52FF80D604FF29F8CDA10CE77B" box="[220,331,1810,1837]" metricMagnitude="-1" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="1" pageNumber="91" unit="cm" value="27.5" valueMax="30.0" valueMin="25.0">25–30 cm</quantity>
long, green, glabrous; lamina ovate-elliptical, 38–42 ×
<quantity id="4C999B52FF80D604FC4AF8CDA475E77B" box="[959,1074,1810,1836]" metricMagnitude="-1" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="1" pageNumber="91" unit="cm" value="17.0" valueMax="18.0" valueMin="16.0">16–18 cm</quantity>
, apex acuminate, margin hyaline, translucent white, shortly hairy, base acute to acuminate, adaxially light green, prominent veins quite closely arranged, c.
<quantity id="4C999B52FF80D604FF53F885A0ACE723" box="[166,235,1882,1909]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="91" unit="mm" value="5.0">5 mm</quantity>
apart, abaxially pale green, puberulent, midrib green, glabrous. Inflorescence central. Peduncle 11–23 × 0.8–1.0 cm, white, glabrous, hidden in pseudostem. Spike 13–19 ×
<quantity id="4C999B52FF80D604FC6FF8A1A3B4E7CF" box="[922,1011,1918,1945]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="1" pageNumber="91" unit="cm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 cm</quantity>
, 16–23 fertile bracts and 10–13 coma bracts. Fertile bracts ovate-elliptical, 3–3.5 ×
<quantity id="4C999B52FF80D604FD78F87DA2BEE7EB" box="[653,761,1954,1980]" metricMagnitude="-2" metricUnit="m" metricValue="2.85" metricValueMax="3.0" metricValueMin="2.7" pageId="1" pageNumber="91" unit="cm" value="2.85" valueMax="3.0" valueMin="2.7">2.7–3 cm</quantity>
, lower half fused with the axis, white with light purple color towards tip, glabrous on both surfaces, slightly acuminate at tip. Coma bracts more linear than fertile ones, obovatelanceolate, 4–4.2 ×
<quantity id="4C999B52FF80D604FE96F835A1A5E853" box="[355,482,2026,2053]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="1" pageNumber="91" unit="cm" value="1.55" valueMax="1.6" valueMin="1.5">1.5–1.6 cm</quantity>
, white with purple tinge towards margin and tip, glabrous, tip slightly acute. Cincinnus 3-flowered. Bracteole single per flower, 1.2–2 ×
<quantity id="4C999B52FF80D604FD4FF7D1A37AE87F" box="[698,829,2062,2089]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="1" pageNumber="91" unit="cm" value="0.6" valueMax="0.7" valueMin="0.5">0.5–0.7 cm</quantity>
, hyaline, translucent white, glabrous, apex mucronate. Flowers 5.5–5.9 ×
<quantity id="4C999B52FF83D607FEACFF41A19FE0EE" box="[345,472,158,185]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="2" pageNumber="92" unit="cm" value="1.5" valueMax="1.6" valueMin="1.4">1.4–1.6 cm</quantity>
, almost double the size of floral bracts. Calyx
<quantity id="4C999B52FF83D607FC2CFF41A417E0EE" box="[985,1104,158,185]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="2" pageNumber="92" unit="mm" value="10.5" valueMax="11.0" valueMin="10.0">10–11 mm</quantity>
long, 3-toothed,
<quantity id="4C999B52FF83D607FAF8FF41A52DE0EE" box="[1293,1386,158,185]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="2" pageNumber="92" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
deep, white, pubescent outside. Corolla tube
<quantity id="4C999B52FF83D607FDB6FF1DA2EAE08A" box="[579,685,194,221]" metricMagnitude="-2" metricUnit="m" metricValue="4.15" metricValueMax="4.3" metricValueMin="4.0" pageId="2" pageNumber="92" unit="cm" value="4.15" valueMax="4.3" valueMin="4.0">4–4.3 cm</quantity>
long, yellow, sometimes tinged with pink or yellow towards lobes, glabrous; dorsal corolla lobe broadly triangular, 1.8–2 ×
<quantity id="4C999B52FF83D607FD0FFF39A323E156" box="[762,868,230,257]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="2" pageNumber="92" unit="cm" value="1.1" valueMax="1.2" valueMin="1.0">1–1.2 cm</quantity>
, mucronate at apex, mucro
<quantity id="4C999B52FF83D607FB60FF39A4B6E156" box="[1173,1265,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="2" pageNumber="92" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
long, white with a pink tinge towards tip, glabrous; lateral corolla lobes 1.1–1.2 ×
<quantity id="4C999B52FF83D607FC8DFED5A3BDE172" box="[888,1018,266,293]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="2" pageNumber="92" unit="cm" value="0.85" valueMax="0.9" valueMin="0.8">0.8–0.9 cm</quantity>
, glabrous, purple, apex slightly acute. Lateral staminodes oblong-lanceolate, apex obtuse, 1.6–1.8 ×
<quantity id="4C999B52FF83D607FCC0FEF1A3F4E11E" box="[821,947,302,329]" metricMagnitude="-3" metricUnit="m" metricValue="6.499999999999999" metricValueMax="7.0" metricValueMin="6.0" pageId="2" pageNumber="92" unit="cm" value="0.6499999999999999" valueMax="0.7" valueMin="0.6">0.6–0.7 cm</quantity>
, usually pale yellow with white towards base. Labellum 2.4–2.5 ×
<quantity id="4C999B52FF83D607FE99FE8DA1ABE13A" box="[364,492,338,365]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="2" pageNumber="92" unit="cm" value="1.4" valueMax="1.5" valueMin="1.3">1.3–1.5 cm</quantity>
, tip emarginate, split
<quantity id="4C999B52FF83D607FD29FE8DA37EE13A" box="[732,825,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="2" pageNumber="92" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
deep, deep yellow band at middle, light yellow towards base and margins. Anther spurred, anther thecae whitish,
<quantity id="4C999B52FF83D607FCF2FEA9A323E1C6" box="[775,868,374,401]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="2" pageNumber="92" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6–7 mm</quantity>
long; filament
<quantity id="4C999B52FF83D607FBFBFEA9A42CE1C6" box="[1038,1131,374,401]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="2" pageNumber="92" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">6–8 mm</quantity>
long, yellow,
<quantity id="4C999B52FF83D607FAFCFEA9A522E1C6" box="[1289,1381,374,401]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="2" pageNumber="92" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
broad at base,
<emphasis id="B915EAA5FF83D607FF11FE44A0B0E1E3" box="[228,247,411,436]" italics="true" pageId="2" pageNumber="92">c.</emphasis>
<quantity id="4C999B52FF83D607FF0AFE45A106E1E2" box="[255,321,410,437]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="92" unit="mm" value="2.0">2 mm</quantity>
broad at upper part; spur
<quantity id="4C999B52FF83D607FD93FE45A29FE1E2" box="[614,728,410,437]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="2" pageNumber="92" unit="mm" value="3.25" valueMax="3.5" valueMin="3.0">3–3.5 mm</quantity>
long, white, pointing forward, glabrous. Ovary trilocular, 4–5 ×
<quantity id="4C999B52FF83D607FF7DFE61A0BCE18E" box="[136,251,446,473]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="2" pageNumber="92" unit="mm" value="2.75" valueMax="3.0" valueMin="2.5">2.5–3 mm</quantity>
, densely hairy; stigma appressed within lobes,
<quantity id="4C999B52FF83D607FCF0FE61A326E18E" box="[773,865,446,473]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="2" pageNumber="92" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
long, white, mouth hairy. Epigynous glands 2, 2.5–3 ×
<quantity id="4C999B52FF83D607FF6AFE3DA16FE1AA" box="[159,296,482,509]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="2" pageNumber="92" unit="mm" value="0.6" valueMax="0.7" valueMin="0.5">0.5–0.7 mm</quantity>
, yellow. Fruit not seen.
</paragraph>
<caption id="DF1E663FFF83D607FF7DF818A577E872" ID-DOI="http://doi.org/10.5281/zenodo.14515945" ID-Zenodo-Dep="14515945" httpUri="https://zenodo.org/record/14515945/files/figure.png" pageId="2" pageNumber="92" startId="2.[136,234,1991,2013]" targetBox="[287,1300,530,1963]" targetPageId="2" targetType="figure">
<paragraph id="8BDE36B7FF83D607FF7DF818A577E872" blockId="2.[136,1452,1991,2085]" pageId="2" pageNumber="92">
<emphasis id="B915EAA5FF83D607FF7DF818A143E78B" bold="true" box="[136,260,1991,2013]" pageId="2" pageNumber="92">FIGURE 1.</emphasis>
<emphasis id="B915EAA5FF83D607FEFEF817A266E78A" box="[267,545,1992,2013]" italics="true" pageId="2" pageNumber="92">
<taxonomicName id="4C614D34FF83D607FEFEF817A188E78A" authorityName="Sabu &amp; Hareesh &amp; Tiatemsu" authorityYear="2024" box="[267,463,1992,2013]" class="Liliopsida" family="Zingiberaceae" genus="Curcuma" kingdom="Plantae" order="Zingiberales" pageId="2" pageNumber="92" phylum="Tracheophyta" rank="species" species="ungmensis" status="sp. nov.">Curcuma ungmensis</taxonomicName>
<taxonomicNameLabel id="A22657DEFF83D607FE22F817A266E78A" box="[471,545,1992,2013]" pageId="2" pageNumber="92" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A. Habit; B. Inflorescence; C. Rhizome; D. C.S. of rhizome; E. Fertile bract; F. Coma bract; G. Bracteole; H. Flower; I. Calyx; J. Dorsal corolla lobe; K1 &amp; K2. Lateral corolla lobes; L1 &amp; L2. Lateral staminodes; M. Labellum; N. Anther lateral-view; O. Anther front-view; P. Ovary with epigynous glands; Q. Ovary. (Photos by M. Sabu &amp; V.S. Hareesh).
</paragraph>
</caption>
<paragraph id="8BDE36B7FF82D606FF48FF41A4AFE08B" blockId="3.[136,1452,158,761]" pageId="3" pageNumber="93">
<emphasis id="B915EAA5FF82D606FF48FF41A172E0EE" bold="true" box="[189,309,158,185]" italics="true" pageId="3" pageNumber="93">Flowering</emphasis>
:—Flowering starts in the rainy season of August and lasts up to October and the plants enter dormancy in January. Flowers open in the morning and last a single day. Inflorescence centrally amidst leaves.
</paragraph>
</subSubSection>
<subSubSection id="C37B653CFF82D606FF48FF39A3BEE277" pageId="3" pageNumber="93" type="distribution">
<paragraph id="8BDE36B7FF82D606FF48FF39A53AE18F" blockId="3.[136,1452,158,761]" pageId="3" pageNumber="93">
<emphasis id="B915EAA5FF82D606FF48FF39A2ADE157" bold="true" box="[189,746,230,257]" italics="true" pageId="3" pageNumber="93">Distribution, conservation and biotic association</emphasis>
:—The plant was found along roadside in the Ungma Village of Mokokchung district of
<collectingRegion id="49A5F855FF82D606FE63FED5A243E173" box="[406,516,266,292]" country="India" name="Nagaland" pageId="3" pageNumber="93">Nagaland</collectingRegion>
,
<emphasis id="B915EAA5FF82D606FDFAFED4A264E173" box="[527,547,267,292]" italics="true" pageId="3" pageNumber="93">ie</emphasis>
., about three kms. from the Mokokchung Town towards Tuensang. So far known only from the
<typeStatus id="54DA8815FF82D606FEDFFEF0A11DE11E" box="[298,346,303,329]" pageId="3" pageNumber="93">type</typeStatus>
locality and adjacent areas on east facing slopes. Preliminary observations have shown that more than 1000 individuals are present in this locality. This species faces a severe threat from various human activities such as road widening, construction of buildings, urbanization and natural calamities. The dominant tree species in that area is
<taxonomicName id="4C614D34FF82D606FF57FE44A272E1E2" authority="Murray (1784: 858)" authorityName="Murray" authorityPageNumber="858" authorityYear="1784" box="[162,565,410,437]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="3" pageNumber="93" phylum="Tracheophyta" rank="species" species="serrata">
<emphasis id="B915EAA5FF82D606FF57FE44A111E1E3" box="[162,342,411,437]" italics="true" pageId="3" pageNumber="93">Quercus serrata</emphasis>
Murray (1784: 858)
</taxonomicName>
with occasional mix of
<taxonomicName id="4C614D34FF82D606FCCCFE44A486E1E3" authority="Linnaeus (1753: 692)" authorityName="Linnaeus" authorityPageNumber="692" authorityYear="1753" box="[825,1217,410,437]" class="Magnoliopsida" family="Malvaceae" genus="Urena" kingdom="Plantae" order="Malvales" pageId="3" pageNumber="93" phylum="Tracheophyta" rank="species" species="lobata">
<emphasis id="B915EAA5FF82D606FCCCFE44A38AE1E3" box="[825,973,410,436]" italics="true" pageId="3" pageNumber="93">Urena lobata</emphasis>
<bibRefCitation id="EFF04B46FF82D606FC24FE45A486E1E3" author="Linnaeus, C." box="[977,1217,410,437]" pageId="3" pageNumber="93" refId="ref3025" refString="Linnaeus, C. (1753) Species Plantarum, ed. 1. Holmiae [Stockholm]: Impensis Laurentii Salvii." type="journal volume" year="1753">Linnaeus (1753: 692)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C614D34FF82D606FB3FFE45A118E18F" authority="Linnaeus (1753: 3)" authorityName="Linnaeus" authorityPageNumber="3" authorityYear="1753" class="Liliopsida" family="Zingiberaceae" genus="Kaempferia" kingdom="Plantae" order="Zingiberales" pageId="3" pageNumber="93" phylum="Tracheophyta" rank="species" species="rotunda">
<emphasis id="B915EAA5FF82D606FB3FFE45A5EBE1E3" box="[1226,1452,410,436]" italics="true" pageId="3" pageNumber="93">Kaempferia rotunda</emphasis>
<bibRefCitation id="EFF04B46FF82D606FF7DFE61A118E18F" author="Linnaeus, C." box="[136,351,446,473]" pageId="3" pageNumber="93" refId="ref3025" refString="Linnaeus, C. (1753) Species Plantarum, ed. 1. Holmiae [Stockholm]: Impensis Laurentii Salvii." type="journal volume" year="1753">Linnaeus (1753: 3)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C614D34FF82D606FE9EFE61A2BAE18F" authority="Linnaeus (1753: 832)" authorityName="Linnaeus" authorityPageNumber="832" authorityYear="1753" box="[363,765,446,473]" class="Magnoliopsida" family="Asteraceae" genus="Bidens" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="93" phylum="Tracheophyta" rank="species" species="pilosa">
<emphasis id="B915EAA5FF82D606FE9EFE61A245E18F" box="[363,514,446,472]" italics="true" pageId="3" pageNumber="93">Bidens pilosa</emphasis>
<bibRefCitation id="EFF04B46FF82D606FDFFFE61A2BAE18F" author="Linnaeus, C." box="[522,765,446,473]" pageId="3" pageNumber="93" refId="ref3025" refString="Linnaeus, C. (1753) Species Plantarum, ed. 1. Holmiae [Stockholm]: Impensis Laurentii Salvii." type="journal volume" year="1753">Linnaeus (1753: 832)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C614D34FF82D606FCFCFE61A3E4E18E" box="[777,931,446,473]" class="Liliopsida" family="Araceae" genus="Colocasia" kingdom="Plantae" order="Alismatales" pageId="3" pageNumber="93" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B915EAA5FF82D606FCFCFE61A33CE18F" box="[777,891,446,472]" italics="true" pageId="3" pageNumber="93">Colocasia</emphasis>
sp.
</taxonomicName>
, etc. forms a dominant ground vegetation.
</paragraph>
<paragraph id="8BDE36B7FF82D606FF48FE3CA3BEE277" blockId="3.[136,1452,158,761]" pageId="3" pageNumber="93">
<emphasis id="B915EAA5FF82D606FF48FE3CA0B4E1AA" bold="true" box="[189,243,483,509]" italics="true" pageId="3" pageNumber="93">Uses</emphasis>
:—The attractive colour of the inflorescence makes this a good candidate as cut flower. It can be introduced in gardens as an ornamental plant for ground cover after proper domestication.
</paragraph>
</subSubSection>
<subSubSection id="C37B653CFF82D606FF48FDF5A512E2AF" pageId="3" pageNumber="93" type="etymology">
<paragraph id="8BDE36B7FF82D606FF48FDF5A175E23F" blockId="3.[136,1452,158,761]" pageId="3" pageNumber="93">
<emphasis id="B915EAA5FF82D606FF48FDF5A17FE212" bold="true" box="[189,312,554,581]" italics="true" pageId="3" pageNumber="93">Etymology</emphasis>
:—The specific epithet
<taxonomicName id="4C614D34FF82D606FDB4FDF5A286E213" authorityName="Sabu &amp; Hareesh &amp; Tiatemsu" authorityYear="2024" box="[577,705,554,580]" class="Liliopsida" family="Zingiberaceae" genus="Curcuma" kingdom="Plantae" order="Zingiberales" pageId="3" pageNumber="93" phylum="Tracheophyta" rank="species" species="ungmensis">
‘
<emphasis id="B915EAA5FF82D606FDBEFDF4A286E213" box="[587,705,555,580]" italics="true" pageId="3" pageNumber="93">ungmensis</emphasis>
</taxonomicName>
’ refers to the name of the village Ungma where the
<typeStatus id="54DA8815FF82D606FAF8FDF4A57AE212" box="[1293,1341,555,581]" pageId="3" pageNumber="93">type</typeStatus>
materials were collected.
</paragraph>
<paragraph id="8BDE36B7FF82D606FF48FDACA512E2AF" blockId="3.[136,1452,158,761]" pageId="3" pageNumber="93">
<emphasis id="B915EAA5FF82D606FF48FDACA0B9E2DA" bold="true" box="[189,254,627,653]" italics="true" pageId="3" pageNumber="93">Notes</emphasis>
:—The new species
<taxonomicName id="4C614D34FF82D606FE1FFDACA2C3E2DB" authorityName="Sabu &amp; Hareesh &amp; Tiatemsu" authorityYear="2024" box="[490,644,627,653]" class="Liliopsida" family="Zingiberaceae" genus="Curcuma" kingdom="Plantae" order="Zingiberales" pageId="3" pageNumber="93" phylum="Tracheophyta" rank="species" species="ungmensis">
<emphasis id="B915EAA5FF82D606FE1FFDACA2C3E2DB" box="[490,644,627,653]" italics="true" pageId="3" pageNumber="93">C. ungmensis</emphasis>
</taxonomicName>
is most allied to
<taxonomicName id="4C614D34FF82D606FCA4FDACA4F2E2DA" authority="Blatter (1931: 357)" authorityName="Blatter" authorityPageNumber="357" authorityYear="1931" box="[849,1205,626,653]" class="Liliopsida" family="Zingiberaceae" genus="Curcuma" kingdom="Plantae" order="Zingiberales" pageId="3" pageNumber="93" phylum="Tracheophyta" rank="species" species="inodora">
<emphasis id="B915EAA5FF82D606FCA4FDACA38AE2DB" box="[849,973,626,653]" italics="true" pageId="3" pageNumber="93">C. inodora</emphasis>
<bibRefCitation id="EFF04B46FF82D606FC22FDADA4F2E2DA" author="Blatter, E." box="[983,1205,626,653]" pageId="3" pageNumber="93" refId="ref2593" refString="Blatter, E. (1931) Curcuma inodora. Journal and Proceedings of the Asiatic Society of Bengal 26: 357. [Calcutta]" type="journal volume" year="1931">Blatter (1931: 357)</bibRefCitation>
</taxonomicName>
, a species commonly distributed along the Western Ghats in Goa,
<collectingRegion id="49A5F855FF82D606FD84FD49A2BAE2E7" box="[625,765,662,688]" country="India" name="Maharashtra" pageId="3" pageNumber="93">Maharashtra</collectingRegion>
and
<collectingRegion id="49A5F855FF82D606FCC4FD49A3EFE2E7" box="[817,936,662,688]" country="India" name="Karnataka" pageId="3" pageNumber="93">Karnataka</collectingRegion>
. The most prominent difference between these two species lies in the flowers and rhizomes. The rhizome in
<taxonomicName id="4C614D34FF82D606FCC7FD64A38EE283" authorityName="Sabu &amp; Hareesh &amp; Tiatemsu" authorityYear="2024" box="[818,969,699,725]" class="Liliopsida" family="Zingiberaceae" genus="Curcuma" kingdom="Plantae" order="Zingiberales" pageId="3" pageNumber="93" phylum="Tracheophyta" rank="species" species="ungmensis">
<emphasis id="B915EAA5FF82D606FCC7FD64A38EE283" box="[818,969,699,725]" italics="true" pageId="3" pageNumber="93">C. ungmensis</emphasis>
</taxonomicName>
is branched and much larger against small, unbranched and conical rhizome in the latter. A detailed comparison of these two species are given in
<tableCitation id="C6E3030CFF82D606FB0EFD01A508E2AF" box="[1275,1359,734,760]" captionStart="TABLE 1" captionStartId="3.[137,229,808,832]" captionText="TABLE 1. Comparison of morphological characters of C. ungmensis and C. inodora." pageId="3" pageNumber="93">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
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<taxonomicName id="4C614D37FF985050FED66999FD7DFEE0" authority="W.H. Chen, Y.F. Han &amp; J.D. Liang" authorityName="W. H. Chen, Y. F. Han &amp; J. D. Liang" authorityYear="2024" box="[136,747,244,271]" class="Sordariomycetes" family="Bionectriaceae" genus="Ovicillium" kingdom="Fungi" order="Hypocreales" pageId="3" pageNumber="198" phylum="Ascomycota" rank="species" species="sinense" status="sp. nov.">
<emphasis id="B915EAA6FF985050FED66999FEC8FEE0" bold="true" box="[136,350,244,271]" italics="true" pageId="3" pageNumber="198">Ovicillium sinense</emphasis>
W.H. Chen, Y.F. Han &amp; J.D. Liang
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,
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(
<figureCitation id="135A2A31FF985050FD046999FC0BFEE0" box="[858,925,244,270]" captionStart="FIGURE 2" captionStartId="3.[136,234,1134,1156]" captionTargetBox="[208,1378,403,1110]" captionTargetId="figure-43@3.[208,1378,403,1110]" captionTargetPageId="3" captionText="FIGURE 2. Ovicillium sinense. A. Infected pupa. B–C. Culture on PDA, showing the top (B) and the underside (C). D–J. Phialides and conidia formed on PDA. Scale bars: B, C = 10 mm, D–J = 10 μm." pageId="3" pageNumber="198">Fig. 2</figureCitation>
)
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<paragraph id="8BDE36B4FF985050FED66875FECDFEDD" blockId="3.[136,347,280,307]" box="[136,347,280,307]" pageId="3" pageNumber="198">
<emphasis id="B915EAA6FF985050FED66875FE96FEDC" box="[136,256,280,306]" italics="true" pageId="3" pageNumber="198">MycoBank</emphasis>
854182
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<emphasis id="B915EAA6FF985050FED6680DFEB8FE94" bold="true" box="[136,302,352,378]" pageId="3" pageNumber="198">Etymology:—</emphasis>
Referring to
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, its country of origin.
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</subSubSection>
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<paragraph id="8BDE36B4FF985050FED66D03FC94FB46" blockId="3.[136,1452,1134,1193]" pageId="3" pageNumber="198">
<emphasis id="B915EAA6FF985050FED66D03FE95FB6D" bold="true" box="[136,259,1134,1156]" pageId="3" pageNumber="198">FIGURE 2.</emphasis>
<taxonomicName id="4C614D37FF985050FF576D02FE2AFB6A" authorityName="W. H. Chen, Y. F. Han &amp; J. D. Liang" authorityYear="2024" box="[265,444,1135,1156]" class="Sordariomycetes" family="Bionectriaceae" genus="Ovicillium" kingdom="Fungi" order="Hypocreales" pageId="3" pageNumber="198" phylum="Ascomycota" rank="species" species="sinense">
<emphasis id="B915EAA6FF985050FF576D02FE2AFB6A" box="[265,444,1135,1156]" italics="true" pageId="3" pageNumber="198">Ovicillium sinense</emphasis>
</taxonomicName>
. A. Infected pupa. B–C. Culture on PDA, showing the top (B) and the underside (C). D–J. Phialides and conidia formed on PDA. Scale bars: B, C = 10 mm, D–J = 10 μm.
</paragraph>
</caption>
<subSubSection id="C37B653FFF985050FEE36DBAFE6DF9FF" pageId="3" pageNumber="198" type="materials_examined">
<paragraph id="8BDE36B4FF985050FEE36DBAFD48FAD7" blockId="3.[136,1452,1239,1697]" pageId="3" pageNumber="198">
<emphasis id="B915EAA6FF985050FEE36DBAFE88FB1F" bold="true" box="[189,286,1239,1265]" pageId="3" pageNumber="198">Type:—</emphasis>
<collectingCountry id="F3767624FF985050FF436DBAFEEFFB1F" box="[285,377,1239,1265]" name="China" pageId="3" pageNumber="198">CHINA</collectingCountry>
.
<collectingRegion id="49A5F856FF985050FFDC6DBAFE71FB1F" box="[386,487,1239,1265]" country="China" name="Guizhou" pageId="3" pageNumber="198">Guizhou</collectingRegion>
: Qiannan Buyei and Miao Autonomous Prefecture, Sandu County, Yaorenshan National Forest Park (
<geoCoordinate id="EE555073FF985050FF426D96FE51FAF8" box="[284,455,1275,1302]" degrees="25" direction="north" minutes="57" orientation="latitude" pageId="3" pageNumber="198" precision="1" seconds="22.21" value="25.956171">25°57'22.21&quot;N</geoCoordinate>
,
<geoCoordinate id="EE555073FF985050FF8A6D96FD1FFAF8" box="[468,649,1275,1302]" degrees="107" direction="east" minutes="57" orientation="longitude" pageId="3" pageNumber="198" precision="1" seconds="54.69" value="107.96519">107°57'54.69&quot;E</geoCoordinate>
), on a pupa (Lepidoptera),
<date id="FFDF1074FF985050FD9D6D96FB35FAFB" box="[963,1187,1275,1301]" pageId="3" pageNumber="198" value="2021-09-20">20 September 2021</date>
, Wanhao Chen,
<collectionCode id="ED70AE71FF985050FB016D96FA3DFAFB" box="[1375,1451,1275,1301]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13167" lsid="urn:lsid:biocol.org:col:13167" name="Guizhou Agricultural College" pageId="3" pageNumber="198" type="Herbarium">GZAC</collectionCode>
SD0970 (
<typeStatus id="54DA8816FF985050FEAD6C72FECDFAD7" box="[243,347,1311,1337]" pageId="3" pageNumber="198" type="holotype">holotype</typeStatus>
), ex-type living culture, SD09701.
</paragraph>
<paragraph id="8BDE36B4FF985050FEE36C2EFE6DF9FF" blockId="3.[136,1452,1239,1697]" pageId="3" pageNumber="198">
Colonies on
<collectionCode id="ED70AE71FF985050FF186C2EFE17FAB3" box="[326,385,1347,1373]" country="Sri Lanka" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13944" lsid="urn:lsid:biocol.org:col:13944" name="Royal Botanic Gardens" pageId="3" pageNumber="198" type="Herbarium">PDA</collectionCode>
, attaining a diameter of
<quantity id="4C999B51FF985050FCDA6C2EFD6DFAB0" box="[644,763,1347,1374]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="3" pageNumber="198" unit="mm" value="19.0" valueMax="20.0" valueMin="18.0">18–20 mm</quantity>
after 14 days at 25 °C, white, consisting of a basal felt, floccose hyphal overgrowth; reverse yellowish. Hyphae septate, hyaline, smooth-walled, 1.4–2.8 μm wide. Conidiophores hyaline, smooth-walled, with single phialide or whorls of 2–5 phialides or verticillium-like directly from hyphae, 17.0–21.7 × 2.3–3.0 μm. Phialides cylindrical, somewhat inflated base, 16.2–25.8 × 1.7–2.4 μm, tapering to a thin neck. Conidia hyaline, smooth-walled, globose to ovoid, 2.1–2.9 × 1.1–1.7 μm, aggregated in large globose to subglobose heads. Sexual state not observed.
</paragraph>
</subSubSection>
<subSubSection id="C37B653FFF985050FEE36F76FD0CF94F" pageId="3" pageNumber="198" type="biology_ecology">
<paragraph id="8BDE36B4FF985050FEE36F76FE6DF9DB" blockId="3.[136,1452,1239,1697]" box="[189,507,1563,1590]" pageId="3" pageNumber="198">
<emphasis id="B915EAA6FF985050FEE36F76FE8CF9DB" bold="true" box="[189,282,1563,1589]" pageId="3" pageNumber="198">Host:—</emphasis>
Pupa (Lepidoptera).
</paragraph>
<paragraph id="8BDE36B4FF985050FEE36F52FD0CF94F" blockId="3.[136,1452,1239,1697]" pageId="3" pageNumber="198">
<emphasis id="B915EAA6FF985050FEE36F52FDA5F9B7" bold="true" box="[189,563,1599,1625]" pageId="3" pageNumber="198">Additional strain examined:—</emphasis>
<collectingCountry id="F3767624FF985050FC6D6F52FD19F9B7" box="[563,655,1599,1625]" name="China" pageId="3" pageNumber="198">CHINA</collectingCountry>
.
<collectingRegion id="49A5F856FF985050FCC06F52FC95F9B7" box="[670,771,1599,1625]" country="China" name="Guizhou" pageId="3" pageNumber="198">Guizhou</collectingRegion>
: Qiannan Buyei and Miao Autonomous Prefecture, Sandu County, Yaorenshan National Forest Park (
<geoCoordinate id="EE555073FF985050FC336F0EFC8EF990" box="[621,792,1635,1662]" degrees="25" direction="north" minutes="57" orientation="latitude" pageId="3" pageNumber="198" precision="1" seconds="22.21" value="25.956171">25°57'22.21&quot;N</geoCoordinate>
,
<geoCoordinate id="EE555073FF985050FD7A6F0EFC4CF990" box="[804,986,1635,1662]" degrees="107" direction="east" minutes="57" orientation="longitude" pageId="3" pageNumber="198" precision="1" seconds="54.69" value="107.96519">107°57'54.69&quot;E</geoCoordinate>
), on a pupa (Lepidoptera),
<date id="FFDF1074FF985050FB516F0EFF55F94F" pageId="3" pageNumber="198" value="2021-09-20">20 September 2021</date>
, Wanhao Chen, SD09702 (living culture).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="AECF252938CE9414CA3D98B74766BC63">Fritillaria shahuensis (Liliaceae), a new species from western Iran</mods:title>
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<subSubSection id="C3620283987AFFA1BA90FC89BA319AEC" box="[136,836,884,911]" pageId="1" pageNumber="280" type="nomenclature">
<paragraph id="8BC75108987AFFA1BA90FC89BA319AEC" blockId="1.[136,836,884,911]" box="[136,836,884,911]" pageId="1" pageNumber="280">
<heading id="D08FE664987AFFA1BA90FC89BA319AEC" box="[136,836,884,911]" fontSize="11" level="2" pageId="1" pageNumber="280" reason="3">
<taxonomicName id="4C782A8B987AFFA1BA90FC89B8899AEC" authority="M. Advay" authorityName="M. Advay" authorityYear="2024" box="[136,508,884,911]" class="Liliopsida" family="Liliaceae" genus="Fritillaria" kingdom="Plantae" order="Liliales" pageId="1" pageNumber="280" phylum="Tracheophyta" rank="species" species="shahuensis" status="sp. nov.">
<emphasis id="B90C8D1A987AFFA1BA90FC89B8F69AED" bold="true" box="[136,387,884,911]" italics="true" pageId="1" pageNumber="280">Fritillaria shahuensis</emphasis>
M. Advay
</taxonomicName>
,
<emphasis id="B90C8D1A987AFFA1B81FFC88BB299AEC" box="[519,604,885,910]" italics="true" pageId="1" pageNumber="280">
<taxonomicNameLabel id="A23F3061987AFFA1B81FFC88BB299AEC" box="[519,604,885,910]" pageId="1" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13434D8D987AFFA1B872FC89BBD89AED" box="[618,685,884,911]" captionStart="FIGURE 1" captionStartId="1.[136,229,1873,1895]" captionTargetBox="[303,1285,949,1836]" captionTargetId="figure-299@1.[303,1285,935,1848]" captionTargetPageId="1" captionText="FIGURE 1. Distribution of F. shahuensis (pink rectangles), F. atrolineata (light blue squares), F. chlorantha (black circles), F. assyriaca (black triangles) and F. kordestanica (blue circles) in Iran." figureDoi="http://doi.org/10.5281/zenodo.14516096" httpUri="https://zenodo.org/record/14516096/files/figure.png" pageId="1" pageNumber="280">Figs 1</figureCitation>
,
<figureCitation id="13434D8D987AFFA1B8ABFC89BBB39AED" box="[691,710,884,911]" captionStart="FIGURE 4" captionStartId="4.[136,229,1589,1611]" captionText="FIGURE 4. Habitat of Fritillaria shahuensis. Photo by Bahman Sharifi." pageId="1" pageNumber="280">4</figureCitation>
,5,
<tableCitation id="C6FA64B3987AFFA1B8F0FC89BA4C9AEC" box="[744,825,884,910]" captionStart="TABLE 1" captionStartId="5.[136,216,159,180]" captionText="TABLE 1. A comparison of selected characters differing among Fritillaria shahuensis, F. atrolineata, F. chlorantha, F. assyriaca and F." pageId="1" pageNumber="280">Table 1</tableCitation>
)
</heading>
</paragraph>
</subSubSection>
<caption id="DF070180987AFFA1BA90F8ACBBC39EE8" ID-DOI="http://doi.org/10.5281/zenodo.14516096" ID-Zenodo-Dep="14516096" httpUri="https://zenodo.org/record/14516096/files/figure.png" pageId="1" pageNumber="280" startId="1.[136,229,1873,1895]" targetBox="[303,1285,949,1836]" targetPageId="1" targetType="figure">
<paragraph id="8BC75108987AFFA1BA90F8ACBBC39EE8" blockId="1.[136,1451,1873,1931]" pageId="1" pageNumber="280">
<emphasis id="B90C8D1A987AFFA1BA90F8ACB98D9E04" bold="true" box="[136,248,1873,1895]" pageId="1" pageNumber="280">FIGURE 1</emphasis>
. Distribution of
<taxonomicName id="4C782A8B987AFFA1BB80F8AFBB6C9E04" authorityName="M. Advay" authorityYear="2024" box="[408,537,1873,1895]" class="Liliopsida" family="Liliaceae" genus="Fritillaria" kingdom="Plantae" order="Liliales" pageId="1" pageNumber="280" phylum="Tracheophyta" rank="species" species="shahuensis">
<emphasis id="B90C8D1A987AFFA1BB80F8AFBB6C9E04" box="[408,537,1873,1895]" italics="true" pageId="1" pageNumber="280">F. shahuensis</emphasis>
</taxonomicName>
(pink rectangles),
<taxonomicName id="4C782A8B987AFFA1B8D6F8AFBA259E04" authorityName="Khaniki" authorityYear="1997" box="[718,848,1873,1895]" class="Liliopsida" family="Liliaceae" genus="Fritillaria" kingdom="Plantae" order="Liliales" pageId="1" pageNumber="280" phylum="Tracheophyta" rank="species" species="atrolineata">
<emphasis id="B90C8D1A987AFFA1B8D6F8AFBA259E04" box="[718,848,1873,1895]" italics="true" pageId="1" pageNumber="280">F. atrolineata</emphasis>
</taxonomicName>
(light blue squares),
<taxonomicName id="4C782A8B987AFFA1BE03F8AFBDE99E04" authorityName="Haussknecht &amp; Bornmuller" authorityYear="1905" box="[1051,1180,1873,1895]" class="Liliopsida" family="Liliaceae" genus="Fritillaria" kingdom="Plantae" order="Liliales" pageId="1" pageNumber="280" phylum="Tracheophyta" rank="species" species="chlorantha">
<emphasis id="B90C8D1A987AFFA1BE03F8AFBDE99E04" box="[1051,1180,1873,1895]" italics="true" pageId="1" pageNumber="280">F. chlorantha</emphasis>
</taxonomicName>
(black circles),
<taxonomicName id="4C782A8B987AFFA1BF20F8AFBCDE9E05" authorityName="Baker" authorityYear="1874" box="[1336,1451,1874,1895]" class="Liliopsida" family="Liliaceae" genus="Fritillaria" kingdom="Plantae" order="Liliales" pageId="1" pageNumber="280" phylum="Tracheophyta" rank="species" species="assyriaca">
<emphasis id="B90C8D1A987AFFA1BF20F8AFBCDE9E05" box="[1336,1451,1874,1895]" italics="true" pageId="1" pageNumber="280">F. assyriaca</emphasis>
</taxonomicName>
(black triangles) and
<taxonomicName id="4C782A8B987AFFA1BB4BF88BB8939EE8" box="[339,486,1909,1931]" class="Liliopsida" family="Liliaceae" genus="Fritillaria" kingdom="Plantae" order="Liliales" pageId="1" pageNumber="280" phylum="Tracheophyta" rank="species" species="kordestanica">
<emphasis id="B90C8D1A987AFFA1BB4BF88BB8939EE8" box="[339,486,1909,1931]" italics="true" pageId="1" pageNumber="280">F. kordestanica</emphasis>
</taxonomicName>
(blue circles) in Iran.
</paragraph>
</caption>
<subSubSection id="C3620283987AFFA3BA90F840B8CD98F3" lastPageId="3" lastPageNumber="282" pageId="1" pageNumber="280" type="materials_examined">
<paragraph id="8BC75108987AFFA1BA90F840BB029178" blockId="1.[136,1452,1981,2075]" pageId="1" pageNumber="280">
<materialsCitation id="3B105B55987AFFA1BA90F840BB069178" collectingDate="2013-04-24" collectionCode="N, E, HKS, TUH" country="Iran" county="Type" elevation="1460" location="Paveh" municipality="Marivan" pageId="1" pageNumber="280" specimenCount="2" stateProvince="Kordestan" typeStatus="holotype">
<collectingCounty id="62A62984987AFFA1BA90F840B9CE9EB0" box="[136,187,1981,2002]" pageId="1" pageNumber="280">Type</collectingCounty>
:—
<collectingCountry id="F36F1198987AFFA1BACFF840B8619EB0" box="[215,276,1981,2002]" name="Iran" pageId="1" pageNumber="280">IRAN</collectingCountry>
.
<collectingRegion id="49BC9FEA987AFFA1BB06F840B8A89EB1" box="[286,477,1981,2003]" country="Iran" name="Kordestan" pageId="1" pageNumber="280">Kurdistan Province</collectingRegion>
:
<collectingMunicipality id="6BA3CB72987AFFA1BBF0F840BB4C9EB1" box="[488,569,1981,2003]" pageId="1" pageNumber="280">Marivan</collectingMunicipality>
to
<location id="8EA707D3987AFFA1B842F840BBE29EB1" LSID="urn:lsid:plazi:treatment:03D1E01E987AFFA6BA90FC89BAF89B42:8EA707D3987AFFA1B842F840BBE29EB1" box="[602,663,1981,2003]" country="Iran" county="Type" municipality="Marivan" name="Paveh" pageId="1" pageNumber="280" stateProvince="Kordestan">Paveh</location>
,
<location id="8EA707D3987AFFA1B8B9F840BB8F9EB0" LSID="urn:lsid:plazi:treatment:03D1E01E987AFFA6BA90FC89BAF89B42:8EA707D3987AFFA1B8B9F840BB8F9EB0" box="[673,762,1981,2002]" country="Iran" county="Type" municipality="Marivan" name="Avroman" pageId="1" pageNumber="280" stateProvince="Kordestan">Avroman</location>
(=Hawraman) region, north-west slope of the
<location id="8EA707D3987AFFA1BED9F840BC109EB1" LSID="urn:lsid:plazi:treatment:03D1E01E987AFFA6BA90FC89BAF89B42:8EA707D3987AFFA1BED9F840BC109EB1" box="[1217,1381,1981,2003]" country="Iran" county="Type" municipality="Marivan" name="Shahu Mountain" pageId="1" pageNumber="280" stateProvince="Kordestan">Shahu Mountain</location>
,
<location id="8EA707D3987AFFA1BF77F840B8709E95" LSID="urn:lsid:plazi:treatment:03D1E01E987AFFA6BA90FC89BAF89B42:8EA707D3987AFFA1BF77F840B8709E95" country="Iran" county="Type" municipality="Marivan" name="Novin Village" pageId="1" pageNumber="280" stateProvince="Kordestan">Novin Village</location>
, 351105°
<collectionCode id="ED69C9CD987AFFA1BB71F81CB8099E94" box="[361,380,2017,2038]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="1" pageNumber="280" type="Herbarium">N</collectionCode>
, 462149°
<collectionCode id="ED69C9CD987AFFA1BBF9F81CB8879E94" box="[481,498,2017,2038]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="1" pageNumber="280" type="Herbarium">E</collectionCode>
, in
<taxonomicName id="4C782A8B987AFFA1B800F81CBB1D9E94" box="[536,616,2017,2038]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="1" pageNumber="280" phylum="Tracheophyta" rank="genus">
<emphasis id="B90C8D1A987AFFA1B800F81CBB1D9E94" box="[536,616,2017,2038]" italics="true" pageId="1" pageNumber="280">Quercus</emphasis>
</taxonomicName>
forest, mountain steppe, subalpine pasture and grassy ledges,
<quantity id="4C80FCED987AFFA1BED5F81CBC6F9E94" box="[1229,1306,2017,2039]" metricMagnitude="3" metricUnit="m" metricValue="1.46" pageId="1" pageNumber="280" unit="m" value="1460.0">
<elevation id="0055B63B987AFFA1BED5F81CBC6F9E94" box="[1229,1306,2017,2039]" metricMagnitude="3" metricUnit="m" metricValue="1.46" pageId="1" pageNumber="280" unit="m" value="1460.0">1460 m</elevation>
</quantity>
,
<date id="FFC677C8987AFFA1BF3DF81CBCD29E94" box="[1317,1447,2017,2038]" pageId="1" pageNumber="280" value="2013-04-24">
<collectingDate id="EF828E20987AFFA1BF3DF81CBCD29E94" box="[1317,1447,2017,2038]" pageId="1" pageNumber="280" value="2013-04-24">24 Apr. 2013</collectingDate>
</date>
,
<emphasis id="B90C8D1A987AFFA1BAA5F7F8B84C9178" box="[189,313,2053,2074]" italics="true" pageId="1" pageNumber="280">Advay 11385</emphasis>
(
<typeStatus id="54C3EFAA987AFFA1BB5EF7F8B8EA9179" box="[326,415,2053,2075]" pageId="1" pageNumber="280" type="holotype">holotype</typeStatus>
:
<collectionCode id="ED69C9CD987AFFA1BBB0F7F8B8AE9178" box="[424,475,2053,2074]" country="Iran" httpUri="http://grbio.org/cool/yamd-d1t3" name="Research Center of Agricultural and Natural Resources Kurdistan Province" pageId="1" pageNumber="280" type="Herbarium">HKS</collectionCode>
;
<typeStatus id="54C3EFAA987AFFA1BBFDF7F8BB5B9179" box="[485,558,2053,2075]" pageId="1" pageNumber="280" type="isotype">isotype</typeStatus>
:
<collectionCode id="ED69C9CD987AFFA1B82EF7F8BB199178" box="[566,620,2053,2074]" country="Iran" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13098" lsid="urn:lsid:biocol.org:col:13098" name="Tehran University" pageId="1" pageNumber="280" type="Herbarium">TUH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC751089879FFA2BA90FF62BAE0999E" blockId="2.[136,1452,159,253]" pageId="2" pageNumber="281">
The new species is like
<taxonomicName id="4C782A8B9879FFA2BB73FF5DB88799D6" authorityName="Khaniki" authorityYear="1997" box="[363,498,159,181]" class="Liliopsida" family="Liliaceae" genus="Fritillaria" kingdom="Plantae" order="Liliales" pageId="2" pageNumber="281" phylum="Tracheophyta" rank="species" species="atrolineata">
<emphasis id="B90C8D1A9879FFA2BB73FF5DB80F99D7" box="[363,378,160,181]" italics="true" pageId="2" pageNumber="281">F</emphasis>
.
<emphasis id="B90C8D1A9879FFA2BB9EFF62B88799D6" box="[390,498,159,180]" italics="true" pageId="2" pageNumber="281">atrolineata</emphasis>
</taxonomicName>
<emphasis id="B90C8D1A9879FFA2BBEAFF62BB8E99D7" box="[498,763,159,181]" italics="true" pageId="2" pageNumber="281">
,
<taxonomicName id="4C782A8B9879FFA2BBE3FF5DBB0A99D6" authorityName="Haussknecht &amp; Bornmuller" authorityYear="1905" box="[507,639,159,181]" class="Liliopsida" family="Liliaceae" genus="Fritillaria" kingdom="Plantae" order="Liliales" pageId="2" pageNumber="281" phylum="Tracheophyta" rank="species" species="chlorantha">F. chlorantha</taxonomicName>
,
<taxonomicName id="4C782A8B9879FFA2B890FF5DBB8E99D7" authorityName="Baker" authorityYear="1874" box="[648,763,160,181]" class="Liliopsida" family="Liliaceae" genus="Fritillaria" kingdom="Plantae" order="Liliales" pageId="2" pageNumber="281" phylum="Tracheophyta" rank="species" species="assyriaca">F. assyriaca</taxonomicName>
</emphasis>
and
<taxonomicName id="4C782A8B9879FFA2B931FF5DBACE99D6" box="[809,955,159,181]" class="Liliopsida" family="Liliaceae" genus="Fritillaria" kingdom="Plantae" order="Liliales" pageId="2" pageNumber="281" phylum="Tracheophyta" rank="species" species="kordestanica">
<emphasis id="B90C8D1A9879FFA2B931FF5DBACE99D6" box="[809,955,159,181]" italics="true" pageId="2" pageNumber="281">F. kordestanica</emphasis>
</taxonomicName>
in its narrowly campanulate flowers and habit (
<figureCitation id="13434D8D9879FFA2BF9CFF62B9BE99BA" captionStart="FIGURE 4" captionStartId="4.[136,229,1589,1611]" captionText="FIGURE 4. Habitat of Fritillaria shahuensis. Photo by Bahman Sharifi." pageId="2" pageNumber="281">Fig. 4</figureCitation>
,
<tableCitation id="C6FA64B39879FFA2BACDFF3EB86999BA" box="[213,284,195,217]" captionStart="TABLE 1" captionStartId="5.[136,216,159,180]" captionText="TABLE 1. A comparison of selected characters differing among Fritillaria shahuensis, F. atrolineata, F. chlorantha, F. assyriaca and F." pageId="2" pageNumber="281">Table 1</tableCitation>
) but differs from these species in number and size of leaves and colour, perigone colour, size of tepal segments, filament colour, nectary shape and colour, style surface and branched style (
<tableCitation id="C6FA64B39879FFA2B927FF1ABAF2999E" box="[831,903,231,253]" captionStart="TABLE 1" captionStartId="5.[136,216,159,180]" captionText="TABLE 1. A comparison of selected characters differing among Fritillaria shahuensis, F. atrolineata, F. chlorantha, F. assyriaca and F." pageId="2" pageNumber="281">Table 1</tableCitation>
).
</paragraph>
<caption id="DF0701809879FFA2BA90F8DEBA109E5B" ID-DOI="http://doi.org/10.5281/zenodo.14516100" ID-Zenodo-Dep="14516100" box="[136,869,1827,1849]" httpUri="https://zenodo.org/record/14516100/files/figure.png" pageId="2" pageNumber="281" startId="2.[136,229,1827,1849]" targetBox="[227,1359,279,1803]" targetPageId="2" targetType="figure">
<paragraph id="8BC751089879FFA2BA90F8DEBA109E5B" blockId="2.[136,869,1827,1849]" box="[136,869,1827,1849]" pageId="2" pageNumber="281">
<emphasis id="B90C8D1A9879FFA2BA90F8DEB9829E5A" bold="true" box="[136,247,1827,1849]" pageId="2" pageNumber="281">FIGURE 2</emphasis>
.
<taxonomicName id="4C782A8B9879FFA2BB1BF8DEBB459E5B" authority="A. Floral" authorityName="A. Floral" box="[259,560,1827,1849]" class="Liliopsida" family="Liliaceae" genus="Fritillaria" kingdom="Plantae" order="Liliales" pageId="2" pageNumber="281" phylum="Tracheophyta" rank="species" species="shahuensis">
<emphasis id="B90C8D1A9879FFA2BB1BF8DEB8A19E5A" box="[259,468,1827,1848]" italics="true" pageId="2" pageNumber="281">Fritillaria shahuensis</emphasis>
A. Floral
</taxonomicName>
details. B–E. Habit and habitat.
</paragraph>
</caption>
<paragraph id="8BC751089879FFA3BA90F89AB8CD98F3" blockId="2.[136,1452,1895,2066]" lastBlockId="3.[136,1452,158,401]" lastPageId="3" lastPageNumber="282" pageId="2" pageNumber="281">
Bulb-bearing herbs, bulbs globose to ovoid,
<quantity id="4C80FCED9879FFA2B89FF89ABA739EE0" box="[647,774,1895,1922]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" metricValueMax="3.5" metricValueMin="1.3" pageId="2" pageNumber="281" unit="cm" value="2.4" valueMax="3.5" valueMin="1.3">1.3–3.5 cm</quantity>
in diameter, with 1–2 bulblets. Stem
<quantity id="4C80FCED9879FFA2BEA0F89ABC589EE0" box="[1208,1325,1895,1922]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="3.5" metricValueMin="1.5" pageId="2" pageNumber="281" unit="cm" value="25.0" valueMax="35.0" valueMin="15.0">15–35 cm</quantity>
,
<quantity id="4C80FCED9879FFA2BF22F89ABCDE9EE0" box="[1338,1451,1895,1922]" metricMagnitude="-1" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="2" pageNumber="281" unit="cm" value="22.5" valueMax="30.0" valueMin="15.0">15–30 cm</quantity>
above ground, sometimes to
<quantity id="4C80FCED9879FFA2BBCAF876BB6C9EC4" box="[466,537,1931,1958]" metricMagnitude="-1" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="281" unit="cm" value="35.0">35 cm</quantity>
in fruiting stage, glabrous. Leaves 4–7 per stem, rarely up to 16 leaves, sessile, alternate, glaucous; the lowest 6.5–16.0 ×
<quantity id="4C80FCED9879FFA2B866F852BA769EA8" box="[638,771,1967,1994]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.5" metricValueMin="0.7" pageId="2" pageNumber="281" unit="cm" value="1.1" valueMax="1.5" valueMin="0.7">0.7–1.5 cm</quantity>
, oblong, lanceolate, acute. Bracts 3.5–12.0 ×
<quantity id="4C80FCED9879FFA2BF3AF852BCD29EA8" box="[1314,1447,1967,1994]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="2" pageNumber="281" unit="cm" value="0.4" valueMax="0.5" valueMin="0.3">0.3–0.5 cm</quantity>
; narrowly oblanceolate, acute, twisted. Flower 1–3 (–4), narrowly campanulate; perianth segments yellow-greenish, yellow-whitish, green to grayish brownish, rarely reddish purplish or blackish outside, yellow brownish, rarely purplish brownish inside, outer ones 16.0–22.0 × 3.5–5.0 mm, oblong-lanceolate, subacute to obtuse, sometimes recurved, ciliate-tufted at apex, inner ones 15.0–23.0 × 4.5–5.0 mm, lanceolate, subacute to obtuse. Nectaries 3.5–4.5 × 0.5– 1.0 mm, linear, black, rarely greenish,
<quantity id="4C80FCED9878FFA3B85AFF1BBBF09863" box="[578,645,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="282" unit="mm" value="1.0">1 mm</quantity>
above the base tepal. Filaments
<quantity id="4C80FCED9878FFA3B9E3FF1BBD299863" box="[1019,1116,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="3" pageNumber="282" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 mm</quantity>
, white, yellowish or greenish in upper part slender, cylindric, papillose; anthers
<quantity id="4C80FCED9878FFA3B8CCFEF7BA379847" box="[724,834,266,293]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="11.0" metricValueMin="4.0" pageId="3" pageNumber="282" unit="mm" value="7.5" valueMax="11.0" valueMin="4.0">4–11 mm</quantity>
before dehiscence, ellipsoid, yellow, rarely greenish, basifixed. Ovary
<quantity id="4C80FCED9878FFA3BB57FED3B8DB982B" box="[335,430,302,329]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="3" pageNumber="282" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">6–8 mm</quantity>
long; style 6.0–8.5 ×
<quantity id="4C80FCED9878FFA3B8A8FED3BA4E982B" box="[688,827,302,329]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="3" pageNumber="282" unit="mm" value="2.0" valueMax="2.5" valueMin="1.5">1.5–2.5 mm</quantity>
, stout, cylindrical, greenish, entire, undivided at apex, densely papillose. Capsule 2.5–4.5 ×
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, obovoid-oblong, cuneate, not winged; seeds 5.0–6.0 × 5.0–
<quantity id="4C80FCED9878FFA3BF48FEAFBCD2980F" box="[1360,1447,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="3" pageNumber="282" unit="mm" value="5.5">5.5 mm</quantity>
, obovate, brown (
<figureCitation id="13434D8D9878FFA3BB5DFE8BB8FB98F2" box="[325,398,374,400]" captionStart="FIGURE 4" captionStartId="4.[136,229,1589,1611]" captionText="FIGURE 4. Habitat of Fritillaria shahuensis. Photo by Bahman Sharifi." pageId="3" pageNumber="282">Figs 4</figureCitation>
, 5).
</paragraph>
</subSubSection>
<caption id="DF0701809878FFA3BA90F817B88D9146" ID-DOI="http://doi.org/10.5281/zenodo.14516103" ID-Zenodo-Dep="14516103" httpUri="https://zenodo.org/record/14516103/files/figure.png" pageId="3" pageNumber="282" startId="3.[136,229,2026,2048]" targetBox="[247,1343,425,2001]" targetPageId="3" targetType="figure">
<paragraph id="8BC751089878FFA3BA90F817B88D9146" blockId="3.[136,1451,2026,2084]" pageId="3" pageNumber="282">
<emphasis id="B90C8D1A9878FFA3BA90F817B98C9E9D" bold="true" box="[136,249,2026,2048]" pageId="3" pageNumber="282">FIGURE 3</emphasis>
.
<taxonomicName id="4C782A8B9878FFA3BB1FF817B8A49E9D" box="[263,465,2026,2047]" class="Liliopsida" family="Liliaceae" genus="Fritillaria" kingdom="Plantae" order="Liliales" pageId="3" pageNumber="282" phylum="Tracheophyta" rank="species" species="shahuensi">
<emphasis id="B90C8D1A9878FFA3BB1FF817B8A49E9D" box="[263,465,2026,2047]" italics="true" pageId="3" pageNumber="282">Fritillaria shahuensi</emphasis>
</taxonomicName>
s, A, B. Habit. C, D. inner and outer surfaces of inner and outer tepals, pistil and stamens. E. Floral details. F. Seeds. G. Capsule. H. Bulb.
</paragraph>
</caption>
<caption id="DF070180987FFFA4BA90F9C8BA349F29" box="[136,833,1589,1611]" pageId="4" pageNumber="283" startId="4.[136,229,1589,1611]" targetType="figure">
<paragraph id="8BC75108987FFFA4BA90F9C8BA349F29" blockId="4.[136,833,1589,1611]" box="[136,833,1589,1611]" pageId="4" pageNumber="283">
<emphasis id="B90C8D1A987FFFA4BA90F9C8B9829F28" bold="true" box="[136,247,1589,1611]" pageId="4" pageNumber="283">FIGURE 4</emphasis>
. Habitat of
<emphasis id="B90C8D1A987FFFA4BB72F9C8BB379F28" box="[362,578,1589,1610]" italics="true" pageId="4" pageNumber="283">
<taxonomicName id="4C782A8B987FFFA4BB72F9C8BB489F28" box="[362,573,1589,1610]" class="Liliopsida" family="Liliaceae" genus="Fritillaria" kingdom="Plantae" order="Liliales" pageId="4" pageNumber="283" phylum="Tracheophyta" rank="species" species="shahuensis">Fritillaria shahuensis</taxonomicName>
.
</emphasis>
Photo by Bahman Sharifi.
</paragraph>
</caption>
<subSubSection id="C3620283987FFFA4BAA5F987BA009FF6" box="[189,885,1658,1684]" pageId="4" pageNumber="283" type="etymology">
<paragraph id="8BC75108987FFFA4BAA5F987BA009FF6" blockId="4.[136,1452,1658,1972]" box="[189,885,1658,1684]" pageId="4" pageNumber="283">
<emphasis id="B90C8D1A987FFFA4BAA5F987B8119FF6" bold="true" box="[189,356,1658,1684]" pageId="4" pageNumber="283">Etymology:—</emphasis>
Referring to the
<typeStatus id="54C3EFAA987FFFA4B804F987BB399FF6" box="[540,588,1658,1684]" pageId="4" pageNumber="283">type</typeStatus>
locality, Shahu Mountain.
</paragraph>
</subSubSection>
<subSubSection id="C3620283987FFFA6BAA5F960BAF89B42" lastPageId="6" lastPageNumber="285" pageId="4" pageNumber="283" type="distribution">
<paragraph id="8BC75108987FFFA4BAA5F960BB069E62" blockId="4.[136,1452,1658,1972]" pageId="4" pageNumber="283">
<emphasis id="B90C8D1A987FFFA4BAA5F960BBFB9FDA" bold="true" box="[189,654,1693,1720]" pageId="4" pageNumber="283">Distribution, habitat and phenology:—</emphasis>
Known also from
<collectingRegion id="49BC9FEA987FFFA4B940F963BA939FDA" box="[856,998,1694,1720]" country="Iran" name="Kermanshah" pageId="4" pageNumber="283">Kermanshah</collectingRegion>
and
<collectingRegion id="49BC9FEA987FFFA4BE03F963BD089FDA" box="[1051,1149,1694,1720]" country="Iran" name="Lorestan" pageId="4" pageNumber="283">Lorestan</collectingRegion>
Provinces. It is an element of the Irano-Turanian floristic region and grows in
<taxonomicName id="4C782A8B987FFFA4B8A6F93FBA699FBE" box="[702,796,1730,1756]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="283" phylum="Tracheophyta" rank="genus">
<emphasis id="B90C8D1A987FFFA4B8A6F93FBA699FBE" box="[702,796,1730,1756]" italics="true" pageId="4" pageNumber="283">Quercus</emphasis>
</taxonomicName>
forest, mountain steppes and grassy ledges,
<quantity id="4C80FCED987FFFA4BF12F93FBCDD9FBE" box="[1290,1448,1730,1756]" metricMagnitude="3" metricUnit="m" metricValue="1.8" metricValueMax="2.3" metricValueMin="1.3" pageId="4" pageNumber="283" unit="m" value="1800.0" valueMax="2300.0" valueMin="1300.0">1300–2300 m</quantity>
. Flowering March–April, fruiting June–July.
</paragraph>
<paragraph id="8BC75108987FFFA4BAA5F8F4BDF39ED6" blockId="4.[136,1452,1658,1972]" pageId="4" pageNumber="283">
<emphasis id="B90C8D1A987FFFA4BAA5F8F4BB2A9E46" bold="true" box="[189,607,1801,1828]" pageId="4" pageNumber="283">Species conservation assessment:—</emphasis>
Extent of occurrence (EOO) is less than
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, and the area of occupancy (AOO) is less than
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(criterion B1, B2); number of locations is less than five (a), with an estimated continuing decline (a, b) of occurrence (i), of occupancy (ii), quality of habitat (iii), and number of subpopulations (iv). Its populations are threatened by grazing. The developing capsules are usually grazed before maturation. This new species is assessed to be considered as endangered, EN B1ab (i, iv) B2ab (ii, iii, iv)” (IUCN 2019).
</paragraph>
<caption id="DF070180987EFFA5BA90FF62BCD899D7" box="[136,1453,159,181]" pageId="5" pageNumber="284" startId="5.[136,216,159,180]" targetType="table">
<paragraph id="8BC75108987EFFA5BA90FF62BCD899D7" blockId="5.[136,1453,159,217]" box="[136,1453,159,181]" pageId="5" pageNumber="284">
<emphasis id="B90C8D1A987EFFA5BA90FF62B99F99D6" bold="true" box="[136,234,159,180]" pageId="5" pageNumber="284">TABLE 1</emphasis>
. A comparison of selected characters differing among
<emphasis id="B90C8D1A987EFFA5B8E3FF62BC1F99D7" box="[763,1386,159,181]" italics="true" pageId="5" pageNumber="284">
<taxonomicName id="4C782A8B987EFFA5B8E3FF62BABA99D6" box="[763,975,159,180]" class="Liliopsida" family="Liliaceae" genus="Fritillaria" kingdom="Plantae" order="Liliales" pageId="5" pageNumber="284" phylum="Tracheophyta" rank="species" species="shahuensis">Fritillaria shahuensis</taxonomicName>
,
<taxonomicName id="4C782A8B987EFFA5B9C1FF5DBD2899D6" authorityName="Khaniki" authorityYear="1997" box="[985,1117,159,181]" class="Liliopsida" family="Liliaceae" genus="Fritillaria" kingdom="Plantae" order="Liliales" pageId="5" pageNumber="284" phylum="Tracheophyta" rank="species" species="atrolineata">F. atrolineata</taxonomicName>
,
<taxonomicName id="4C782A8B987EFFA5BE70FF5DBD9999D6" authorityName="Haussknecht &amp; Bornmuller" authorityYear="1905" box="[1128,1260,159,181]" class="Liliopsida" family="Liliaceae" genus="Fritillaria" kingdom="Plantae" order="Liliales" pageId="5" pageNumber="284" phylum="Tracheophyta" rank="species" species="chlorantha">F. chlorantha</taxonomicName>
,
<taxonomicName id="4C782A8B987EFFA5BEEEFF5DBC1F99D7" authorityName="Baker" authorityYear="1874" box="[1270,1386,160,181]" class="Liliopsida" family="Liliaceae" genus="Fritillaria" kingdom="Plantae" order="Liliales" pageId="5" pageNumber="284" phylum="Tracheophyta" rank="species" species="assyriaca">F. assyriaca</taxonomicName>
</emphasis>
and F.
</paragraph>
</caption>
<paragraph id="8BC75108987EFFA5BA90FF3EB87C99BB" blockId="5.[136,1453,159,217]" box="[136,265,195,217]" pageId="5" pageNumber="284">
<taxonomicName id="4C782A8B987EFFA5BA90FF3EB87699BA" box="[136,259,195,216]" class="Liliopsida" family="Liliaceae" genus="Fritillaria" kingdom="Plantae" order="Liliales" pageId="5" pageNumber="284" phylum="Tracheophyta" rank="species" species="kordestanica">
<emphasis id="B90C8D1A987EFFA5BA90FF3EB87699BA" box="[136,259,195,216]" italics="true" pageId="5" pageNumber="284">kordestanica</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC75108987EFFA5BA95FF17BCE0917A" pageId="5" pageNumber="284">
<table id="F978A3A8987E005FBA95FF17BCD2917B" box="[141,1447,234,2073]" gridcols="6" gridrows="17" pageId="5" pageNumber="284">
<tr id="3548534A987E005FBA95FF17BCD2999F" box="[141,1447,234,253]" gridrow="0" pageId="5" pageNumber="284">
<th id="76993A36987E005FBA95FF17B863999F" box="[141,278,234,253]" gridcol="0" gridrow="0" pageId="5" pageNumber="284">
<emphasis id="B90C8D1A987EFFA5BA95FF17B986999F" bold="true" box="[141,243,234,253]" pageId="5" pageNumber="284">Characters</emphasis>
</th>
<th id="76993A36987E005FBB5AFF17BB47999F" box="[322,562,234,253]" gridcol="1" gridrow="0" pageId="5" pageNumber="284">
<taxonomicName id="4C782A8B987EFFA5BB5AFF17B8CF999F" box="[322,442,234,253]" class="Liliopsida" family="Liliaceae" genus="Fritillaria" kingdom="Plantae" order="Liliales" pageId="5" pageNumber="284" phylum="Tracheophyta" rank="species" species="shahuensis">
<emphasis id="B90C8D1A987EFFA5BB5AFF17B8CF999F" bold="true" box="[322,442,234,253]" italics="true" pageId="5" pageNumber="284">F. shahuensis</emphasis>
</taxonomicName>
</th>
<th id="76993A36987E005FB822FF17BA6C999F" box="[570,793,234,253]" gridcol="2" gridrow="0" pageId="5" pageNumber="284">
<taxonomicName id="4C782A8B987EFFA5B822FF17BBC5999F" authorityName="Khaniki" authorityYear="1997" box="[570,688,234,253]" class="Liliopsida" family="Liliaceae" genus="Fritillaria" kingdom="Plantae" order="Liliales" pageId="5" pageNumber="284" phylum="Tracheophyta" rank="species" species="atrolineata">
<emphasis id="B90C8D1A987EFFA5B822FF17BBC5999F" bold="true" box="[570,688,234,253]" italics="true" pageId="5" pageNumber="284">F. atrolineata</emphasis>
</taxonomicName>
</th>
<th id="76993A36987E005FB93AFF17BA80999F" box="[802,1013,234,253]" gridcol="3" gridrow="0" pageId="5" pageNumber="284">
<taxonomicName id="4C782A8B987EFFA5B93AFF17BAEC999F" authorityName="Haussknecht &amp; Bornmuller" authorityYear="1905" box="[802,921,234,253]" class="Liliopsida" family="Liliaceae" genus="Fritillaria" kingdom="Plantae" order="Liliales" pageId="5" pageNumber="284" phylum="Tracheophyta" rank="species" species="chlorantha">
<emphasis id="B90C8D1A987EFFA5B93AFF17BAEC999F" bold="true" box="[802,921,234,253]" italics="true" pageId="5" pageNumber="284">F. chlorantha</emphasis>
</taxonomicName>
</th>
<th id="76993A36987E005FB9E6FF17BDA8999F" box="[1022,1245,234,253]" gridcol="4" gridrow="0" pageId="5" pageNumber="284">
<taxonomicName id="4C782A8B987EFFA5B9E6FF17BD13999F" authorityName="Baker" authorityYear="1874" box="[1022,1126,234,253]" class="Liliopsida" family="Liliaceae" genus="Fritillaria" kingdom="Plantae" order="Liliales" pageId="5" pageNumber="284" phylum="Tracheophyta" rank="species" species="assyriaca">
<emphasis id="B90C8D1A987EFFA5B9E6FF17BD13999F" bold="true" box="[1022,1126,234,253]" italics="true" pageId="5" pageNumber="284">F. assyriaca</emphasis>
</taxonomicName>
</th>
<th id="76993A36987E005FBEFEFF17BCD2999F" box="[1254,1447,234,253]" gridcol="5" gridrow="0" pageId="5" pageNumber="284">
<taxonomicName id="4C782A8B987EFFA5BEFEFF17BC18999F" authority="Bulb" authorityName="Bulb" box="[1254,1389,234,253]" class="Liliopsida" family="Liliaceae" genus="Fritillaria" kingdom="Plantae" order="Liliales" pageId="5" pageNumber="284" phylum="Tracheophyta" rank="species" species="kordestanica">
<emphasis id="B90C8D1A987EFFA5BEFEFF17BC18999F" bold="true" box="[1254,1389,234,253]" italics="true" pageId="5" pageNumber="284">F. kordestanica</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="3548534A987E005FBA95FEF2BCD29820" box="[141,1447,271,322]" gridrow="1" pageId="5" pageNumber="284">
<th id="76993A36987E005FBA95FEF2B8639820" box="[141,278,271,322]" gridcol="0" gridrow="1" pageId="5" pageNumber="284">Bulb diam.</th>
<td id="76993A36987E005FBB5AFEF2BB479820" box="[322,562,271,322]" gridcol="1" gridrow="1" pageId="5" pageNumber="284">subglobose, ovoid, 15–35× 10–23 mm,</td>
<td id="76993A36987E005FB822FEF2BA6C9820" box="[570,793,271,322]" gridcol="2" gridrow="1" pageId="5" pageNumber="284">globose-subglobose, 7–11 × 7–10 mm</td>
<td id="76993A36987E005FB93AFEF2BA809820" box="[802,1013,271,322]" gridcol="3" gridrow="1" pageId="5" pageNumber="284">10–20 mm</td>
<td id="76993A36987E005FB9E6FEF2BDA89820" box="[1022,1245,271,322]" gridcol="4" gridrow="1" pageId="5" pageNumber="284">10–30 mm</td>
<td id="76993A36987E005FBEFEFEF2BCD29820" box="[1254,1447,271,322]" gridcol="5" gridrow="1" pageId="5" pageNumber="284">10–36 mm</td>
</tr>
<tr id="3548534A987E005FBA95FEA9BCD298E7" box="[141,1447,340,389]" gridrow="2" pageId="5" pageNumber="284">
<th id="76993A36987E005FBA95FEA9B86398E7" box="[141,278,340,389]" gridcol="0" gridrow="2" pageId="5" pageNumber="284">Stolon</th>
<td id="76993A36987E005FBB5AFEA9BB4798E7" box="[322,562,340,389]" gridcol="1" gridrow="2" pageId="5" pageNumber="284">1–2 bulbils</td>
<td id="76993A36987E005FB822FEA9BA6C98E7" box="[570,793,340,389]" gridcol="2" gridrow="2" pageId="5" pageNumber="284">without bulbils or stolons</td>
<td id="76993A36987E005FB93AFEA9BA8098E7" box="[802,1013,340,389]" gridcol="3" gridrow="2" pageId="5" pageNumber="284">without bulbils or stolons</td>
<td id="76993A36987E005FB9E6FEA9BDA898E7" box="[1022,1245,340,389]" gridcol="4" gridrow="2" pageId="5" pageNumber="284">1–2 bulbils</td>
<td id="76993A36987E005FBEFEFEA9BCD298E7" box="[1254,1447,340,389]" gridcol="5" gridrow="2" pageId="5" pageNumber="284">2–3 bulbils</td>
</tr>
<tr id="3548534A987E005FBA95FE65BCD2988A" box="[141,1447,408,488]" gridrow="3" pageId="5" pageNumber="284">
<th id="76993A36987E005FBA95FE65B863988A" box="[141,278,408,488]" gridcol="0" gridrow="3" pageId="5" pageNumber="284">Stem</th>
<td id="76993A36987E005FBB5AFE65BB47988A" box="[322,562,408,488]" gridcol="1" gridrow="3" pageId="5" pageNumber="284">15–35 (–40) cm, stout, greenish, brownish, smooth</td>
<td id="76993A36987E005FB822FE65BA6C988A" box="[570,793,408,488]" gridcol="2" gridrow="3" pageId="5" pageNumber="284">20–25cm above the bulb, longer in fruit, smooth.</td>
<td id="76993A36987E005FB93AFE65BA80988A" box="[802,1013,408,488]" gridcol="3" gridrow="3" pageId="5" pageNumber="284">15–23cm</td>
<td id="76993A36987E005FB9E6FE65BDA8988A" box="[1022,1245,408,488]" gridcol="4" gridrow="3" pageId="5" pageNumber="284">up to 25 cm (elongating to 35 cm in fruit), stout, brownish, often papillose</td>
<td id="76993A36987E005FBEFEFE65BCD2988A" box="[1254,1447,408,488]" gridcol="5" gridrow="3" pageId="5" pageNumber="284">10–26 cm, smooth</td>
</tr>
<tr id="3548534A987E005FBA95FE07BCD29B4F" box="[141,1447,506,557]" gridrow="4" pageId="5" pageNumber="284">
<th id="76993A36987E005FBA95FE07B8639B4F" box="[141,278,506,557]" gridcol="0" gridrow="4" pageId="5" pageNumber="284">Leaves</th>
<td id="76993A36987E005FBB5AFE07BB479B4F" box="[322,562,506,557]" gridcol="1" gridrow="4" pageId="5" pageNumber="284">mostly 4–7 (–16), glaucous, sometimes canaliculate</td>
<td id="76993A36987E005FB822FE07BA6C9B4F" box="[570,793,506,557]" gridcol="2" gridrow="4" pageId="5" pageNumber="284">4–5, glaucous</td>
<td id="76993A36987E005FB93AFE07BA809B4F" box="[802,1013,506,557]" gridcol="3" gridrow="4" pageId="5" pageNumber="284">(4–) 5–6 (–10), shining green</td>
<td id="76993A36987E005FB9E6FE07BDA89B4F" box="[1022,1245,506,557]" gridcol="4" gridrow="4" pageId="5" pageNumber="284">4–7 (–12), glaucous</td>
<td id="76993A36987E005FBEFEFE07BCD29B4F" box="[1254,1447,506,557]" gridcol="5" gridrow="4" pageId="5" pageNumber="284">(3–) 4 (–5), shining green</td>
</tr>
<tr id="3548534A987E005FBA95FDC2BCD29B13" box="[141,1447,575,625]" gridrow="5" pageId="5" pageNumber="284" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<td id="76993A36987E005FB822FDC2BA6C9B13" box="[570,793,575,625]" gridcol="2" gridrow="5" pageId="5" pageNumber="284">2–3 usually in a subternate</td>
</tr>
<tr id="3548534A987E005FBA95FD81BCD29A4A" box="[141,1447,636,808]" gridrow="6" pageId="5" pageNumber="284">
<th id="76993A36987E005FBA95FD81B8639A4A" box="[141,278,636,808]" gridcol="0" gridrow="6" pageId="5" pageNumber="284">Lowest leaves</th>
<td id="76993A36987E005FBB5AFD81BB479A4A" box="[322,562,636,808]" gridcol="1" gridrow="6" pageId="5" pageNumber="284">3.5–16 × 0.7–1.5 cm, oblong, lanceolate, acute</td>
<td id="76993A36987E005FB822FD81BA6C9A4A" box="[570,793,636,808]" gridcol="2" gridrow="6" pageId="5" pageNumber="284">(subverticillate) or subopposite position, the rest alternate, the lowest 9 × 1–1.5cm, usually 8–9 times as long as wide, oblanceolate</td>
<td id="76993A36987E005FB93AFD81BA809A4A" box="[802,1013,636,808]" gridcol="3" gridrow="6" pageId="5" pageNumber="284">6–12×1.2–3cm, broadly ovate, lanceolate to linear</td>
<td id="76993A36987E005FB9E6FD81BDA89A4A" box="[1022,1245,636,808]" gridcol="4" gridrow="6" pageId="5" pageNumber="284">8–12 × 0.5–2 cm, canaliculate, linear, ovate- lanceolate, acute</td>
<td id="76993A36987E005FBEFEFD81BCD29A4A" box="[1254,1447,636,808]" gridcol="5" gridrow="6" pageId="5" pageNumber="284">7–12 × 1–2 cm, lanceolate to oblanceolate, acute</td>
</tr>
<tr id="3548534A987E005FBA95FCC6BCD29AEE" box="[141,1447,827,908]" gridrow="7" pageId="5" pageNumber="284">
<th id="76993A36987E005FBA95FCC6B8639AEE" box="[141,278,827,908]" gridcol="0" gridrow="7" pageId="5" pageNumber="284">Flowers</th>
<td id="76993A36987E005FBB5AFCC6BB479AEE" box="[322,562,827,908]" gridcol="1" gridrow="7" pageId="5" pageNumber="284">1–3 (–4), narrowly campanulate</td>
<td id="76993A36987E005FB822FCC6BA6C9AEE" box="[570,793,827,908]" gridcol="2" gridrow="7" pageId="5" pageNumber="284">1 (2), narrowly campanulate</td>
<td id="76993A36987E005FB93AFCC6BA809AEE" box="[802,1013,827,908]" gridcol="3" gridrow="7" pageId="5" pageNumber="284">1–3, narrowly campanulate</td>
<td id="76993A36987E005FB9E6FCC6BDA89AEE" box="[1022,1245,827,908]" gridcol="4" gridrow="7" pageId="5" pageNumber="284">1–2 (–5), narrow campanulate</td>
<td id="76993A36987E005FBEFEFCC6BCD29AEE" box="[1254,1447,827,908]" gridcol="5" gridrow="7" pageId="5" pageNumber="284">solitary, narrowly campanulate, recurved at the mouth</td>
</tr>
<tr id="3548534A987E005FBA95FC63BCD29DC4" box="[141,1447,926,1190]" gridrow="8" pageId="5" pageNumber="284">
<th id="76993A36987E005FBA95FC63B8639DC4" box="[141,278,926,1190]" gridcol="0" gridrow="8" pageId="5" pageNumber="284">Perigone colour</th>
<td id="76993A36987E005FBB5AFC63BB479DC4" box="[322,562,926,1190]" gridcol="1" gridrow="8" pageId="5" pageNumber="284">yellow-greenish, yellow- whitish, green to grayish- brownish, rarely reddish- purplish or blackish outside, yellow-brownish, rarely purplish-brownish inside</td>
<td id="76993A36987E005FB822FC63BA6C9DC4" box="[570,793,926,1190]" gridcol="2" gridrow="8" pageId="5" pageNumber="284">perianth segments usually yellowish-green to green, outside more yellowish towards margins, sometimes sparsely brown-dotted, inside tinged or sparsely dotted, pale reddish-brown towards margins</td>
<td id="76993A36987E005FB93AFC63BA809DC4" box="[802,1013,926,1190]" gridcol="3" gridrow="8" pageId="5" pageNumber="284">yellow-greenish, rarely purplish-brownish</td>
<td id="76993A36987E005FB9E6FC63BDA89DC4" box="[1022,1245,926,1190]" gridcol="4" gridrow="8" pageId="5" pageNumber="284">dusky purplish-grey outside and inside segments, often with narrowly median yellowish or greenish stripe in outside and inside segments</td>
<td id="76993A36987E005FBEFEFC63BCD29DC4" box="[1254,1447,926,1190]" gridcol="5" gridrow="8" pageId="5" pageNumber="284">outside purplish- grey, rarely with green stripe, inside greenish-yellow or purplish</td>
</tr>
<tr id="3548534A987E005FBA95FB45BCD29C6B" box="[141,1447,1208,1289]" gridrow="9" pageId="5" pageNumber="284">
<th id="76993A36987E005FBA95FB45B8639C6B" box="[141,278,1208,1289]" gridcol="0" gridrow="9" pageId="5" pageNumber="284">Outer segments</th>
<td id="76993A36987E005FBB5AFB45BB479C6B" box="[322,562,1208,1289]" gridcol="1" gridrow="9" pageId="5" pageNumber="284">16–22 × 3.5–5 mm, oblong-lanceolate, subacute to obtuse</td>
<td id="76993A36987E005FB822FB45BA6C9C6B" box="[570,793,1208,1289]" gridcol="2" gridrow="9" pageId="5" pageNumber="284">20 × 6 mm, oblanceolate, obtuse</td>
<td id="76993A36987E005FB93AFB45BA809C6B" box="[802,1013,1208,1289]" gridcol="3" gridrow="9" pageId="5" pageNumber="284">20–26 × 4–6 mm, ovate-lanceolate, acute to obtuse</td>
<td id="76993A36987E005FB9E6FB45BDA89C6B" box="[1022,1245,1208,1289]" gridcol="4" gridrow="9" pageId="5" pageNumber="284">15–28 × 4–6 mm, ovate or ovate-lanceolate, obtuse or acute</td>
<td id="76993A36987E005FBEFEFB45BCD29C6B" box="[1254,1447,1208,1289]" gridcol="5" gridrow="9" pageId="5" pageNumber="284">20–26 × 6–7 mm, ovate-lanceolate, acute to obtuse</td>
</tr>
<tr id="3548534A987E005FBA95FAE6BCD29C09" box="[141,1447,1307,1387]" gridrow="10" pageId="5" pageNumber="284">
<th id="76993A36987E005FBA95FAE6B8639C09" box="[141,278,1307,1387]" gridcol="0" gridrow="10" pageId="5" pageNumber="284">Inner segments</th>
<td id="76993A36987E005FBB5AFAE6BB479C09" box="[322,562,1307,1387]" gridcol="1" gridrow="10" pageId="5" pageNumber="284">15–23 × 4.5–5 mm, lanceolate, subacute to obtuse</td>
<td id="76993A36987E005FB822FAE6BA6C9C09" box="[570,793,1307,1387]" gridcol="2" gridrow="10" pageId="5" pageNumber="284">inner ones 8 mm wide, oblanceolate, obtuse, fascia obscure.</td>
<td id="76993A36987E005FB93AFAE6BA809C09" box="[802,1013,1307,1387]" gridcol="3" gridrow="10" pageId="5" pageNumber="284">21–25 × 4–6 mm</td>
<td id="76993A36987E005FB9E6FAE6BDA89C09" box="[1022,1245,1307,1387]" gridcol="4" gridrow="10" pageId="5" pageNumber="284">15–28 × 6–12 mm, obovate, obtuse</td>
<td id="76993A36987E005FBEFEFAE6BCD29C09" box="[1254,1447,1307,1387]" gridcol="5" gridrow="10" pageId="5" pageNumber="284">19–25 × 5–7.5 mm, ovate-lanceolate, obtuse</td>
</tr>
<tr id="3548534A987E005FBA95FA83BCD29CAC" box="[141,1447,1406,1486]" gridrow="11" pageId="5" pageNumber="284">
<th id="76993A36987E005FBA95FA83B8639CAC" box="[141,278,1406,1486]" gridcol="0" gridrow="11" pageId="5" pageNumber="284">Nectary</th>
<td id="76993A36987E005FBB5AFA83BB479CAC" box="[322,562,1406,1486]" gridcol="1" gridrow="11" pageId="5" pageNumber="284">3.5–4.5 × 0.5–1 mm, linear, 1 mm above the base tepal, black rarely greenish</td>
<td id="76993A36987E005FB822FA83BA6C9CAC" box="[570,793,1406,1486]" gridcol="2" gridrow="11" pageId="5" pageNumber="284">linear, 4–6 × 0.5–l mm, placed 0.5–l mm above base of tepals, black</td>
<td id="76993A36987E005FB93AFA83BA809CAC" box="[802,1013,1406,1486]" gridcol="3" gridrow="11" pageId="5" pageNumber="284">3–4 × 1mm, green, linear-lanceolate</td>
<td id="76993A36987E005FB9E6FA83BDA89CAC" box="[1022,1245,1406,1486]" gridcol="4" gridrow="11" pageId="5" pageNumber="284">2–4 × 1.3 mm, green, linear-lanceolate</td>
<td id="76993A36987E005FBEFEFA83BCD29CAC" box="[1254,1447,1406,1486]" gridcol="5" gridrow="11" pageId="5" pageNumber="284">4–6.1 × 1–1.4 mm, yellow-greenish, linear-lanceolate</td>
</tr>
<tr id="3548534A987E005FBA95FA1CBCD29F53" box="[141,1447,1505,1585]" gridrow="12" pageId="5" pageNumber="284">
<th id="76993A36987E005FBA95FA1CB8639F53" box="[141,278,1505,1585]" gridcol="0" gridrow="12" pageId="5" pageNumber="284">Filaments (mm)</th>
<td id="76993A36987E005FBB5AFA1CBB479F53" box="[322,562,1505,1585]" gridcol="1" gridrow="12" pageId="5" pageNumber="284">5–7 mm, white, yellowish and greenish at up part, slender, cylindric, papillose</td>
<td id="76993A36987E005FB822FA1CBA6C9F53" box="[570,793,1505,1585]" gridcol="2" gridrow="12" pageId="5" pageNumber="284">7 mm, yellow, densely papillose in upper part.</td>
<td id="76993A36987E005FB93AFA1CBA809F53" box="[802,1013,1505,1585]" gridcol="3" gridrow="12" pageId="5" pageNumber="284">6–8 mm,, densely papillose, yellow</td>
<td id="76993A36987E005FB9E6FA1CBDA89F53" box="[1022,1245,1505,1585]" gridcol="4" gridrow="12" pageId="5" pageNumber="284">6–11 mm, yellow, enlarged to base, all papillose</td>
<td id="76993A36987E005FBEFEFA1CBCD29F53" box="[1254,1447,1505,1585]" gridcol="5" gridrow="12" pageId="5" pageNumber="284">whitish yellow</td>
</tr>
<tr id="3548534A987E005FBA95F9B9BCD29FF6" box="[141,1447,1604,1684]" gridrow="13" pageId="5" pageNumber="284">
<th id="76993A36987E005FBA95F9B9B8639FF6" box="[141,278,1604,1684]" gridcol="0" gridrow="13" pageId="5" pageNumber="284">Anthers (mm)</th>
<td id="76993A36987E005FBB5AF9B9BB479FF6" box="[322,562,1604,1684]" gridcol="1" gridrow="13" pageId="5" pageNumber="284">4–11 mm, ellipsoid, yellow, rarely greenish</td>
<td id="76993A36987E005FB822F9B9BA6C9FF6" box="[570,793,1604,1684]" gridcol="2" gridrow="13" pageId="5" pageNumber="284">6–7 mm long before dehiscence, ellipsoid, yellow, basifixed.</td>
<td id="76993A36987E005FB93AF9B9BA809FF6" box="[802,1013,1604,1684]" gridcol="3" gridrow="13" pageId="5" pageNumber="284">7–8 mm long before dehiscence,</td>
<td id="76993A36987E005FB9E6F9B9BDA89FF6" box="[1022,1245,1604,1684]" gridcol="4" gridrow="13" pageId="5" pageNumber="284">7–9 mm, ellipsoid, yellow</td>
<td id="76993A36987E005FBEFEF9B9BCD29FF6" box="[1254,1447,1604,1684]" gridcol="5" gridrow="13" pageId="5" pageNumber="284">5.5–8.3, oblong, yellow</td>
</tr>
<tr id="3548534A987E005FBA95F95BBCD29E31" box="[141,1447,1702,1875]" gridrow="14" pageId="5" pageNumber="284">
<th id="76993A36987E005FBA95F95BB8639E31" box="[141,278,1702,1875]" gridcol="0" gridrow="14" pageId="5" pageNumber="284">Style (mm)</th>
<td id="76993A36987E005FBB5AF95BBB479E31" box="[322,562,1702,1875]" gridcol="1" gridrow="14" pageId="5" pageNumber="284">6–8.5 × 1.5–2.5 mm, stout, greenish, cylindrical, densely papillose, entire</td>
<td id="76993A36987E005FB822F95BBA6C9E31" box="[570,793,1702,1875]" gridcol="2" gridrow="14" pageId="5" pageNumber="284">5–7 mm, stout, greenish- yellow, densely papillose, entire or very slightly 3-lobed, apex of stigma lobes papillose and with a central hole.</td>
<td id="76993A36987E005FB93AF95BBA809E31" box="[802,1013,1702,1875]" gridcol="3" gridrow="14" pageId="5" pageNumber="284">5–10 mm, stout, greenish-yellow, densely papillose, slightly 3-lobed, 1–1.5 mm branched</td>
<td id="76993A36987E005FB9E6F95BBDA89E31" box="[1022,1245,1702,1875]" gridcol="4" gridrow="14" pageId="5" pageNumber="284">5–10 × 1.5–2 mm, unbranched or lobes 0.5–1 mm, papillose</td>
<td id="76993A36987E005FBEFEF95BBCD29E31" box="[1254,1447,1702,1875]" gridcol="5" gridrow="14" pageId="5" pageNumber="284">5–8.8 × 2–2.6, papillose</td>
</tr>
<tr id="3548534A987E005FBA95F898BCD29EB7" box="[141,1447,1893,2005]" gridrow="15" pageId="5" pageNumber="284">
<th id="76993A36987E005FBA95F898B8639EB7" box="[141,278,1893,2005]" gridcol="0" gridrow="15" pageId="5" pageNumber="284">Capsule (mm)</th>
<td id="76993A36987E005FBB5AF898BB479EB7" box="[322,562,1893,2005]" gridcol="1" gridrow="15" pageId="5" pageNumber="284">25–45 × 17–21 obovoid, tapering towards the base, obtuse, not winged</td>
<td id="76993A36987E005FB822F898BA6C9EB7" box="[570,793,1893,2005]" gridcol="2" gridrow="15" pageId="5" pageNumber="284">25–30 × 15, obovoid, tapering towards the base, obtuse, not winged</td>
<td id="76993A36987E005FB93AF898BA809EB7" box="[802,1013,1893,2005]" gridcol="3" gridrow="15" pageId="5" pageNumber="284">45–60 × 20–27, obovoid-oblong tapering towards the base, obtuse, not winged</td>
<td id="76993A36987E005FB9E6F898BDA89EB7" box="[1022,1245,1893,2005]" gridcol="4" gridrow="15" pageId="5" pageNumber="284">11–38 × 11–17 mm, obovoid</td>
<td id="76993A36987E005FBEFEF898BCD29EB7" box="[1254,1447,1893,2005]" gridcol="5" gridrow="15" pageId="5" pageNumber="284">13.6–20.7 × 15.7– 16.7 mm, obovoid, tapering towards the base, rounded</td>
</tr>
<tr id="3548534A987E005FBA95F81ABCD2917B" box="[141,1447,2023,2073]" gridrow="16" pageId="5" pageNumber="284">
<th id="76993A36987E005FBA95F81AB863917B" box="[141,278,2023,2073]" gridcol="0" gridrow="16" pageId="5" pageNumber="284">Seeds (mm)</th>
<td id="76993A36987E005FBB5AF81ABB47917B" box="[322,562,2023,2073]" gridcol="1" gridrow="16" pageId="5" pageNumber="284">5–6 × 5–5.5, obovate, coat brown</td>
<td id="76993A36987E005FB822F81ABA6C917B" box="[570,793,2023,2073]" gridcol="2" gridrow="16" pageId="5" pageNumber="284">flat, ovate in outline, 4-5 long</td>
<td id="76993A36987E005FB93AF81ABA80917B" box="[802,1013,2023,2073]" gridcol="3" gridrow="16" pageId="5" pageNumber="284">6–9.5 × 5–7.5, obovate, coat brown</td>
<td id="76993A36987E005FB9E6F81ABDA8917B" box="[1022,1245,2023,2073]" gridcol="4" gridrow="16" pageId="5" pageNumber="284">4–7.5 × 3–6.5 cm, testa pale brown</td>
<td id="76993A36987E005FBEFEF81ABCD2917B" box="[1254,1447,2023,2073]" gridcol="5" gridrow="16" pageId="5" pageNumber="284">4.5–6.7 × 3.6–5.7, flat, triangular-ovate</td>
</tr>
</table>
</paragraph>
<paragraph id="8BC75108987DFFA6BAA5FF63BAB5980F" blockId="6.[136,1452,158,545]" pageId="6" pageNumber="285">
<emphasis id="B90C8D1A987DFFA6BAA5FF63B85399DA" bold="true" box="[189,294,158,184]" pageId="6" pageNumber="285">Notes:—</emphasis>
We compared
<taxonomicName id="4C782A8B987DFFA6BBD2FF62BB1799DA" box="[458,610,158,184]" class="Liliopsida" family="Liliaceae" genus="Fritillaria" kingdom="Plantae" order="Liliales" pageId="6" pageNumber="285" phylum="Tracheophyta" rank="species" species="shahuensis">
<emphasis id="B90C8D1A987DFFA6BBD2FF62BB1799DA" box="[458,610,158,184]" italics="true" pageId="6" pageNumber="285">F. shahuensis</emphasis>
</taxonomicName>
with
<emphasis id="B90C8D1A987DFFA6B8BCFF62BD0999DA" box="[676,1148,158,184]" italics="true" pageId="6" pageNumber="285">
<taxonomicName id="4C782A8B987DFFA6B8BCFF62BA4A99DA" authorityName="Khaniki" authorityYear="1997" box="[676,831,158,184]" class="Liliopsida" family="Liliaceae" genus="Fritillaria" kingdom="Plantae" order="Liliales" pageId="6" pageNumber="285" phylum="Tracheophyta" rank="species" species="atrolineata">F. atrolineata</taxonomicName>
,
<taxonomicName id="4C782A8B987DFFA6B954FF62BA9299DA" authorityName="Haussknecht &amp; Bornmuller" authorityYear="1905" box="[844,999,158,184]" class="Liliopsida" family="Liliaceae" genus="Fritillaria" kingdom="Plantae" order="Liliales" pageId="6" pageNumber="285" phylum="Tracheophyta" rank="species" species="chlorantha">F. chlorantha</taxonomicName>
,
<taxonomicName id="4C782A8B987DFFA6B9ECFF62BD0999DA" authorityName="Baker" authorityYear="1874" box="[1012,1148,159,184]" class="Liliopsida" family="Liliaceae" genus="Fritillaria" kingdom="Plantae" order="Liliales" pageId="6" pageNumber="285" phylum="Tracheophyta" rank="species" species="assyriaca">F. assyriaca</taxonomicName>
</emphasis>
and
<taxonomicName id="4C782A8B987DFFA6BEAEFF62BC1699DA" box="[1206,1379,158,184]" class="Liliopsida" family="Liliaceae" genus="Fritillaria" kingdom="Plantae" order="Liliales" pageId="6" pageNumber="285" phylum="Tracheophyta" rank="species" species="kordestanica">
<emphasis id="B90C8D1A987DFFA6BEAEFF62BC1699DA" box="[1206,1379,158,184]" italics="true" pageId="6" pageNumber="285">F. kordestanica</emphasis>
</taxonomicName>
in the
<taxonomicName id="4C782A8B987DFFA6BA90FF3EB86099BE" box="[136,277,195,220]" class="Liliopsida" family="Liliaceae" genus="Fritillaria" kingdom="Plantae" order="Liliales" pageId="6" pageNumber="285" phylum="Tracheophyta" rank="species" species="caucasica">
<emphasis id="B90C8D1A987DFFA6BA90FF3EB86099BE" box="[136,277,195,220]" italics="true" pageId="6" pageNumber="285">F. caucasica</emphasis>
</taxonomicName>
group (
<tableCitation id="C6FA64B3987DFFA6BB76FF3FB8B699BE" box="[366,451,194,220]" captionStart="TABLE 1" captionStartId="5.[136,216,159,180]" captionText="TABLE 1. A comparison of selected characters differing among Fritillaria shahuensis, F. atrolineata, F. chlorantha, F. assyriaca and F." pageId="6" pageNumber="285">Table 1</tableCitation>
). They are similar in terms of general facies, but
<taxonomicName id="4C782A8B987DFFA6B9E0FF3FBD9899BE" box="[1016,1261,194,220]" class="Liliopsida" family="Liliaceae" genus="Fritillaria" kingdom="Plantae" order="Liliales" pageId="6" pageNumber="285" phylum="Tracheophyta" rank="species" species="shahuensis">
<emphasis id="B90C8D1A987DFFA6B9E0FF3FBD9899BE" box="[1016,1261,194,220]" italics="true" pageId="6" pageNumber="285">Fritillaria shahuensis</emphasis>
</taxonomicName>
differs in colour and size of its lowest leaves and size, shape and colour of perigone segments. The size of leaves (especially lowest leaves) and outside and inside colour of the tepals are usually variable in
<emphasis id="B90C8D1A987DFFA6B9D8FEF7BD4D9846" box="[960,1080,266,292]" italics="true" pageId="6" pageNumber="285">
<taxonomicName id="4C782A8B987DFFA6B9D8FEF7BD419846" authorityName="Linnaeus" authorityYear="1753" box="[960,1076,266,292]" class="Liliopsida" family="Liliaceae" genus="Fritillaria" kingdom="Plantae" order="Liliales" pageId="6" pageNumber="285" phylum="Tracheophyta" rank="genus">Fritillaria</taxonomicName>
,
</emphasis>
particularly in this group. The
<taxonomicName id="4C782A8B987DFFA6BF8FFEF6B98D982A" class="Liliopsida" family="Liliaceae" genus="Fritillaria" kingdom="Plantae" order="Liliales" pageId="6" pageNumber="285" phylum="Tracheophyta" rank="species" species="caucasica">
<emphasis id="B90C8D1A987DFFA6BF8FFEF6B98D982A" italics="true" pageId="6" pageNumber="285">F. caucasica</emphasis>
</taxonomicName>
group is diverse in
<collectingCountry id="F36F1198987DFFA6BBCFFED3BB52982A" box="[471,551,302,328]" name="Turkey" pageId="6" pageNumber="285">Turkey</collectingCountry>
and
<collectingCountry id="F36F1198987DFFA6B846FED2BBFA982B" box="[606,655,303,329]" name="Iran" pageId="6" pageNumber="285">Iran</collectingCountry>
. Although these species are generally widely circumscribed, especially
<emphasis id="B90C8D1A987DFFA6BA90FEAEB863980E" box="[136,278,339,364]" italics="true" pageId="6" pageNumber="285">
<taxonomicName id="4C782A8B987DFFA6BA90FEAEB867980E" authorityName="Baker" authorityYear="1874" box="[136,274,339,364]" class="Liliopsida" family="Liliaceae" genus="Fritillaria" kingdom="Plantae" order="Liliales" pageId="6" pageNumber="285" phylum="Tracheophyta" rank="species" species="assyriaca">F. assyriaca</taxonomicName>
,
</emphasis>
the distinctive floral characteristic of this species stands out.
</paragraph>
<paragraph id="8BC75108987DFFA6BAA5FE8BBAF89B42" blockId="6.[136,1452,158,545]" pageId="6" pageNumber="285">
<emphasis id="B90C8D1A987DFFA6BAA5FE8BBB3698F2" bold="true" box="[189,579,374,400]" pageId="6" pageNumber="285">Selected specimens examined:—</emphasis>
<materialsCitation id="3B105B55987DFFA6B85BFE8BBD1098D6" collectingDate="2018-04-23" collectionCode="HKS" country="Iran" county="Avroman" elevation="1450" location="Novin Village" municipality="Shahu Mountain" pageId="6" pageNumber="285" specimenCount="1" stateProvince="Kordestan">
<collectingCountry id="F36F1198987DFFA6B85BFE8BBBFE98F2" box="[579,651,374,400]" name="Iran" pageId="6" pageNumber="285">IRAN</collectingCountry>
.
<collectingRegion id="49BC9FEA987DFFA6B88FFE8BBA0298F2" box="[663,887,374,400]" country="Iran" name="Kordestan" pageId="6" pageNumber="285">Kurdistan Province</collectingRegion>
:
<collectingCounty id="62A62984987DFFA6B999FE8BBA9C98F2" box="[897,1001,374,400]" pageId="6" pageNumber="285">Avroman</collectingCounty>
(=Hawraman) region, north-west slope of the
<collectingMunicipality id="6BA3CB72987DFFA6BAC8FE67B8F898D6" box="[208,397,410,436]" pageId="6" pageNumber="285">Shahu Mountain</collectingMunicipality>
,
<location id="8EA707D3987DFFA6BB80FE67BB4C98D6" LSID="urn:lsid:plazi:treatment:03D1E01E987AFFA6BA90FC89BAF89B42:8EA707D3987DFFA6BB80FE67BB4C98D6" box="[408,569,410,436]" country="Iran" county="Avroman" municipality="Shahu Mountain" name="Novin Village" pageId="6" pageNumber="285" stateProvince="Kordestan">Novin Village</location>
,
<elevation id="0055B63B987DFFA6B85CFE67BBA398D6" box="[580,726,410,437]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="6" pageNumber="285" unit="m" value="1450.0">
<quantity id="4C80FCED987DFFA6B85CFE67BBEC98D7" box="[580,665,410,437]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="6" pageNumber="285" unit="m" value="1450.0">1450 m</quantity>
a.s.l.
</elevation>
,
<date id="FFC677C8987DFFA6B8FAFE67BA0798D6" box="[738,882,410,436]" pageId="6" pageNumber="285" value="2018-04-23">
<collectingDate id="EF828E20987DFFA6B8FAFE67BA0798D6" box="[738,882,410,436]" pageId="6" pageNumber="285" value="2018-04-23">23 Apr 2018</collectingDate>
</date>
,
<emphasis id="B90C8D1A987DFFA6B965FE67BD6598D6" box="[893,1040,410,436]" italics="true" pageId="6" pageNumber="285">Advay 13375</emphasis>
(
<collectionCode id="ED69C9CD987DFFA6BE07FE67BD2998D6" box="[1055,1116,410,436]" country="Iran" httpUri="http://grbio.org/cool/yamd-d1t3" name="Research Center of Agricultural and Natural Resources Kurdistan Province" pageId="6" pageNumber="285" type="Herbarium">HKS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B105B55987DFFA6BE69FE67BAFC9B42" collectingDate="2013-05-20" collectingDateMax="2024-04-26" collectingDateMin="2013-05-20" collectionCode="TUH, HKS" country="Iran" county="Marivan" elevation="2100" location="Zarravah Mountain" municipality="Paveh" pageId="6" pageNumber="285" specimenCount="1" stateProvince="Kermanshah">
<collectingCounty id="62A62984987DFFA6BE69FE67BDA598D6" box="[1137,1232,410,436]" pageId="6" pageNumber="285">Marivan</collectingCounty>
to
<collectingMunicipality id="6BA3CB72987DFFA6BEECFE67BC4E98D6" box="[1268,1339,410,436]" pageId="6" pageNumber="285">Paveh</collectingMunicipality>
,
<location id="8EA707D3987DFFA6BF5EFE67B98D98BA" LSID="urn:lsid:plazi:treatment:03D1E01E987AFFA6BA90FC89BAF89B42:8EA707D3987DFFA6BF5EFE67B98D98BA" country="Iran" county="Marivan" municipality="Paveh" name="Zarravah Mountain" pageId="6" pageNumber="285" stateProvince="Kermanshah">Zarravah Mountain</location>
,
<location id="8EA707D3987DFFA6BB1AFE42B8A998BA" LSID="urn:lsid:plazi:treatment:03D1E01E987AFFA6BA90FC89BAF89B42:8EA707D3987DFFA6BB1AFE42B8A998BA" box="[258,476,446,473]" country="Iran" county="Marivan" municipality="Paveh" name="Daymayaw Village" pageId="6" pageNumber="285" stateProvince="Kermanshah">Daymayaw Village</location>
,
<quantity id="4C80FCED987DFFA6BBFEFE43BB4998BB" box="[486,572,446,473]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="6" pageNumber="285" unit="m" value="2100.0">
<elevation id="0055B63B987DFFA6BBFEFE43BB4998BB" box="[486,572,446,473]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="6" pageNumber="285" unit="m" value="2100.0">2100 m</elevation>
</quantity>
,
<date id="FFC677C8987DFFA6B85FFE43BBA098BA" box="[583,725,446,472]" pageId="6" pageNumber="285" value="2024-04-26">
<collectingDate id="EF828E20987DFFA6B85FFE43BBA098BA" box="[583,725,446,472]" pageId="6" pageNumber="285" value="2024-04-26">26 Apr 2024</collectingDate>
</date>
,
<emphasis id="B90C8D1A987DFFA6B8F8FE43BA0798BA" box="[736,882,446,472]" italics="true" pageId="6" pageNumber="285">
<location id="8EA707D3987DFFA6B8F8FE43BA5398BA" LSID="urn:lsid:plazi:treatment:03D1E01E987AFFA6BA90FC89BAF89B42:8EA707D3987DFFA6B8F8FE43BA5398BA" box="[736,806,446,472]" country="Iran" county="Marivan" municipality="Paveh" name="Advay" pageId="6" pageNumber="285" stateProvince="Kermanshah">Advay</location>
48835
</emphasis>
(
<collectionCode id="ED69C9CD987DFFA6B999FE43BAB498BA" box="[897,961,446,472]" country="Iran" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13098" lsid="urn:lsid:biocol.org:col:13098" name="Tehran University" pageId="6" pageNumber="285" type="Herbarium">TUH</collectionCode>
).
<collectingRegion id="49BC9FEA987DFFA6B9CAFE43BDBB98BA" box="[978,1230,446,472]" country="Iran" name="Kermanshah" pageId="6" pageNumber="285">Kermanshah Province</collectingRegion>
: Hawraman, Paveh,
<location id="8EA707D3987DFFA6BA90FE1FB821989E" LSID="urn:lsid:plazi:treatment:03D1E01E987AFFA6BA90FC89BAF89B42:8EA707D3987DFFA6BA90FE1FB821989E" box="[136,340,482,508]" country="Iran" county="Marivan" municipality="Paveh" name="Khanegha Village" pageId="6" pageNumber="285" stateProvince="Kermanshah">Khanegha Village</location>
,
<location id="8EA707D3987DFFA6BB46FE1FBB6E989E" LSID="urn:lsid:plazi:treatment:03D1E01E987AFFA6BA90FC89BAF89B42:8EA707D3987DFFA6BB46FE1FBB6E989E" box="[350,539,482,508]" country="Iran" county="Marivan" municipality="Paveh" name="Shahu Mountain" pageId="6" pageNumber="285" stateProvince="Kermanshah">Shahu Mountain</location>
,
<quantity id="4C80FCED987DFFA6B83DFE1FBB09989F" box="[549,636,482,509]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="6" pageNumber="285" unit="m" value="1350.0">
<elevation id="0055B63B987DFFA6B83DFE1FBB09989F" box="[549,636,482,509]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="6" pageNumber="285" unit="m" value="1350.0">1350 m</elevation>
</quantity>
,
<date id="FFC677C8987DFFA6B89FFE1FBA6A989E" box="[647,799,482,509]" pageId="6" pageNumber="285" value="2023-03-30">
<collectingDate id="EF828E20987DFFA6B89FFE1FBA6A989E" box="[647,799,482,509]" pageId="6" pageNumber="285" value="2023-03-30">30 Mar. 2023</collectingDate>
</date>
,
<emphasis id="B90C8D1A987DFFA6B932FE1FBAC9989E" box="[810,956,482,508]" italics="true" pageId="6" pageNumber="285">Advay 48652</emphasis>
(
<collectionCode id="ED69C9CD987DFFA6B9D3FE1FBD7E989E" box="[971,1035,482,508]" country="Iran" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13098" lsid="urn:lsid:biocol.org:col:13098" name="Tehran University" pageId="6" pageNumber="285" type="Herbarium">TUH</collectionCode>
).
<collectingRegion id="49BC9FEA987DFFA6BE05FE1EBD98989E" box="[1053,1261,482,509]" country="Iran" name="Lorestan" pageId="6" pageNumber="285">Lorestan Province</collectingRegion>
: Brujerd,
<location id="8EA707D3987DFFA6BF42FE1FB9A29B42" LSID="urn:lsid:plazi:treatment:03D1E01E987AFFA6BA90FC89BAF89B42:8EA707D3987DFFA6BF42FE1FB9A29B42" country="Iran" county="Marivan" municipality="Paveh" name="Chenar village" pageId="6" pageNumber="285" stateProvince="Kermanshah">Chenar village</location>
,
<location id="8EA707D3987DFFA6BAFAFDFBB8DA9B42" LSID="urn:lsid:plazi:treatment:03D1E01E987AFFA6BA90FC89BAF89B42:8EA707D3987DFFA6BAFAFDFBB8DA9B42" box="[226,431,518,544]" country="Iran" county="Marivan" municipality="Paveh" name="Garrein Mountain" pageId="6" pageNumber="285" stateProvince="Kermanshah">Garrein Mountain</location>
, 1710,
<date id="FFC677C8987DFFA6B818FDFBBBED9B42" box="[512,664,518,545]" pageId="6" pageNumber="285" value="2013-05-20">
<collectingDate id="EF828E20987DFFA6B818FDFBBBED9B42" box="[512,664,518,545]" pageId="6" pageNumber="285" value="2013-05-20">20 May 2013</collectingDate>
</date>
,
<emphasis id="B90C8D1A987DFFA6B8BBFDFBBA419B42" box="[675,820,518,544]" italics="true" pageId="6" pageNumber="285">Advay 11542</emphasis>
(
<collectionCode id="ED69C9CD987DFFA6B95BFDFBBAF59B42" box="[835,896,518,544]" country="Iran" httpUri="http://grbio.org/cool/yamd-d1t3" name="Research Center of Agricultural and Natural Resources Kurdistan Province" pageId="6" pageNumber="285" type="Herbarium">HKS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

181
data/03/DA/87/03DA879DFFF2FFE255DEFBFAFC34F883.xml Normal file
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@ -0,0 +1,181 @@
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<mods:role id="4B163F2B8006B14561C45B6BFDF75945">
<mods:roleTerm id="08668970B8C548C29A38E652723322D8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BA61A2446E61490C6C96EAC5751A9805">Cai, Zhuo-Yu</mods:namePart>
<mods:nameIdentifier id="B6AD89ED198127DE6D81D129ED9FB18F" type="ORCID">0000-0001-9288-0882</mods:nameIdentifier>
<mods:affiliation id="6BA8A6190FFD07063B392ED68624CB61">Co-Innovation Center for Sustainable Forestry in Southern China, Nanjing Forestry University, Nanjing 210037, China &amp; Guangdong Provincial Key Laboratory of Digital Botanical Garden, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou, 510650, China &amp; Bamboo Research Institute, Nanjing Forestry University, Nanjing 210037, China</mods:affiliation>
<mods:nameIdentifier id="194C56BE3637ACA8125599DBCD80FEAB" type="email">caizhuoyu@njfu.edu.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="F63C8953209D0794BA53DB6F5DCB303C" type="personal">
<mods:role id="025393E7F10CCD567080993C0EC06052">
<mods:roleTerm id="C47E88DC321E18C4F814A8EE53DB765B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="75BFC353709731403E8F84AE08A4D863">Widjaja, Elizabeth Anita</mods:namePart>
<mods:nameIdentifier id="A798A394598CCBED13B0C27B17B69B1F" type="ORCID">0000-0003-1746-0567</mods:nameIdentifier>
<mods:affiliation id="484E51F9B1E829864C8C037B9B68FCE1">Environmental Bamboo Foundation, Jl. Moh. Yamin IX No. 15, Sumerta Kelod, Denpasar Timur, Denpasar, Bali, 80239, Indonesia</mods:affiliation>
<mods:nameIdentifier id="03B1E62E92646CA349A68BC251482AD6" type="email">eawidjaja3003@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="635E7FBDF136658D400896287BB80569" type="personal">
<mods:role id="BDB1B44AA8EBD9FCD5C483219757B59D">
<mods:roleTerm id="F6CC36F9DA3600519A5E34BD9C196AC3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="537DC3BFBCBB7F2FA3F44541DC3384DB">Xia, Nian-He</mods:namePart>
<mods:nameIdentifier id="21D528C9B10DC389224348F9F52EC581" type="ORCID">0000-0001-9852-7393</mods:nameIdentifier>
<mods:affiliation id="C7C1BF17AC67E87E08C8F731B1D7DE47">Guangdong Provincial Key Laboratory of Digital Botanical Garden, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou, 510650, China &amp; Key Laboratory of National Forestry and Grassland Administration on Plant Conservation and Utilization in Southern China, Guangzhou 510650, China &amp; South China National Botanical Garden, Guangzhou, 510650, China</mods:affiliation>
<mods:nameIdentifier id="DBFFC2FFE4E8BD9F0999E9842DE1D8B9" type="email">nhxia@scbg.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="4A35C442FB44B179FACD9BE5065F6191" type="personal">
<mods:role id="2707D4D0BD3F877E687B7B0EE476F5C9">
<mods:roleTerm id="C571347E0261C99B85A31A3A39EE4D8C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D2CDF73512B6D733E69E1C64997C263A">Wong, Khoon Meng</mods:namePart>
<mods:nameIdentifier id="BB388A6F7D934C2E6FD7F85EAEF0B9FD" type="ORCID">0000-0002-1950-786X</mods:nameIdentifier>
<mods:affiliation id="5C490DFF44A03C62ABEC827C7DA2C8AD">Herbarium, Singapore Botanic Gardens, National Parks Board, Singapore 259569, Singapore</mods:affiliation>
<mods:nameIdentifier id="18502B72C5B1688C4C8111B735C412F2" type="email">wkm2000@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="B083769EDC2B238CF15F296FA3778DE7">text</mods:typeOfResource>
<mods:relatedItem id="E8DD691748F117B271A6CC3AF4CA62B6" type="host">
<mods:titleInfo id="DC8EB9A08E2F7D31CB2DFF6B560402E5">
<mods:title id="163D27E012679C288C609236ADD33751">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="F387D3E08C9FA5BB052BEE9B9551150D">
<mods:date id="976AE02671A50939CBC8959181F530A2">2024</mods:date>
<mods:detail id="B39A6C6958A06B482A7EEAFDF9D03D6D" type="pubDate">
<mods:number id="BCE0EE38138B4E87DBA5218194C7840B">2024-08-20</mods:number>
</mods:detail>
<mods:detail id="B3088C0F86404E3D118769327F9F4017" type="volume">
<mods:number id="810466AF1D2C84E527DEFFAE2FFD02B0">663</mods:number>
</mods:detail>
<mods:detail id="1E35ABBE22D959BA851B633BFAC2AC82" type="issue">
<mods:number id="0617E3F519E3799921B0B5CF48666F49">2</mods:number>
</mods:detail>
<mods:extent id="4FB52E75AF0E30D5E3FB5035352C8C0E" unit="page">
<mods:start id="AAA101FBCD83B4DB7FA9A0B98556308F">101</mods:start>
<mods:end id="CB5F9C02DD752DB2CF8FB3FCF7BEEB96">105</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="5D6C7D2B3BB278BFB16831E52B6CAD32">
<mods:url id="6568316086DFAF6842E92BCD7F1B2D11">https://doi.org/10.11646/phytotaxa.663.2.6</mods:url>
</mods:location>
<mods:classification id="19FB298A7563755F001DD4628BBEDE6D">journal article</mods:classification>
<mods:identifier id="9E7D9BDE8546F3C4AB051CD31D6F4E51" type="DOI">10.11646/phytotaxa.663.2.6</mods:identifier>
<mods:identifier id="B686370EFD7A2F6814D23E46D86EE627" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="7DE0F3E7666219D4D95869B00A0B91CE" type="Zenodo-Dep">14515951</mods:identifier>
</mods:mods>
<treatment id="03DA879DFFF2FFE255DEFBFAFC34F883" ID-DOI="http://doi.org/10.5281/zenodo.14515953" ID-Zenodo-Dep="14515953" LSID="urn:lsid:plazi:treatment:03DA879DFFF2FFE255DEFBFAFC34F883" httpUri="http://treatment.plazi.org/id/03DA879DFFF2FFE255DEFBFAFC34F883" lastPageNumber="102" pageId="1" pageNumber="102">
<subSubSection id="C3696500FFF2FFE255DEFBFAFC62FB53" box="[136,973,1140,1167]" pageId="1" pageNumber="102" type="nomenclature">
<paragraph id="8BCC368BFFF2FFE255DEFBFAFC62FB53" blockId="1.[136,973,1140,1167]" box="[136,973,1140,1167]" pageId="1" pageNumber="102">
<heading id="D08481E7FFF2FFE255DEFBFAFC62FB53" box="[136,973,1140,1167]" fontSize="11" level="2" pageId="1" pageNumber="102" reason="3">
<taxonomicName id="4C734D08FFF2FFE255DEFBFAFD1DFB53" authority="(Widjaja) Z. Y. Cai &amp; Widjaja" authorityName="Z. Y. Cai &amp; Widjaja" authorityYear="2024" baseAuthorityName="Widjaja, Reinwardtia" baseAuthorityYear="2024" box="[136,690,1140,1167]" class="Liliopsida" family="Poaceae" genus="Gigantochloa" kingdom="Plantae" order="Poales" pageId="1" pageNumber="102" phylum="Tracheophyta" rank="species" species="lako" status="comb. nov.">
<emphasis id="B907EA99FFF2FFE255DEFBFAFEF1FB52" bold="true" box="[136,350,1140,1167]" italics="true" pageId="1" pageNumber="102">Gigantochloa lako</emphasis>
(Widjaja) Z.Y. Cai &amp; Widjaja
</taxonomicName>
,
<emphasis id="B907EA99FFF2FFE257EAFBFAFC9AFB53" box="[700,821,1140,1166]" italics="true" pageId="1" pageNumber="102">
<taxonomicNameLabel id="A23457E2FFF2FFE257EAFBFAFC9AFB53" box="[700,821,1140,1166]" pageId="1" pageNumber="102" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13482A0EFFF2FFE25612FBFAFC3EFB53" box="[836,913,1140,1166]" captionStart="FIGURE 1" captionStartId="2.[136,229,2005,2027]" captionTargetBox="[161,1427,187,1977]" captionTargetId="figure-35@2.[161,1427,187,1977]" captionTargetPageId="2" captionText="FIGURE 1. The flowering collection of Gigantochloa lako (Joseph Lai SING 2016-070). Scale bar = 10 cm." figureDoi="http://doi.org/10.5281/zenodo.14515955" httpUri="https://zenodo.org/record/14515955/files/figure.png" pageId="1" pageNumber="102">Figs. 1</figureCitation>
&amp;
<figureCitation id="13482A0EFFF2FFE256E3FBFAFC6BFB53" box="[949,964,1140,1166]" captionStart="FIGURE 2" captionStartId="3.[136,229,1953,1975]" captionTargetBox="[171,1416,190,1928]" captionTargetId="figure-20@3.[171,1416,190,1928]" captionTargetPageId="3" captionText="FIGURE 2. Floral morphology of Gigantochloa lako. A. pseudospikelet; B. palea with stamens and pistil; C. terminal vestigial floret; D. left: 1—keeled prophyll, right: 2—keeled prophyll; E, F. gemmiferous bracts; G. lemma; H. palea; I. anthers; J. apices of anthers; K. stamens and a pistil showing the filament tube; L. pistil without stigma. Scale bars: J = 0.5 mm; the rest = 1 mm. Based on Joseph Lai SING 2016-070. Photos by Zhuo-Yu Cai." figureDoi="http://doi.org/10.5281/zenodo.14515957" httpUri="https://zenodo.org/record/14515957/files/figure.png" pageId="1" pageNumber="102">2</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3696500FFF2FFE255DEFB31FCC6FB08" box="[136,873,1215,1238]" pageId="1" pageNumber="102" type="reference_group">
<paragraph id="8BCC368BFFF2FFE255DEFB31FCC6FB08" blockId="1.[136,1452,1215,1346]" box="[136,873,1215,1238]" pageId="1" pageNumber="102">
<emphasis id="B907EA99FFF2FFE255DEFB31FF5AFB08" bold="true" box="[136,245,1215,1237]" pageId="1" pageNumber="102">Basionym:</emphasis>
<taxonomicName id="4C734D08FFF2FFE255ADFB4EFDF9FB0B" ID-CoL="KJK6" authority="Widjaja, Reinwardtia" authorityName="Widjaja, Reinwardtia" authorityYear="1997" box="[251,598,1216,1238]" class="Liliopsida" family="Poaceae" genus="Bambusa" kingdom="Plantae" order="Poales" pageId="1" pageNumber="102" phylum="Tracheophyta" rank="species" species="lako">
<emphasis id="B907EA99FFF2FFE255ADFB4EFE2BFB08" box="[251,388,1216,1237]" italics="true" pageId="1" pageNumber="102">Bambusa lako</emphasis>
Widjaja, Reinwardtia
</taxonomicName>
11(2): 61–63,
<figureCitation id="13482A0EFFF2FFE257B2FB4EFCB3FB08" box="[740,796,1216,1238]" captionStart="FIGURE 1" captionStartId="2.[136,229,2005,2027]" captionTargetBox="[161,1427,187,1977]" captionTargetId="figure-35@2.[161,1427,187,1977]" captionTargetPageId="2" captionText="FIGURE 1. The flowering collection of Gigantochloa lako (Joseph Lai SING 2016-070). Scale bar = 10 cm." figureDoi="http://doi.org/10.5281/zenodo.14515955" httpUri="https://zenodo.org/record/14515955/files/figure.png" pageId="1" pageNumber="102">Fig. 1</figureCitation>
(1997).
</paragraph>
</subSubSection>
<subSubSection id="C3696500FFF2FFE255DEFB6AFC34F883" pageId="1" pageNumber="102" type="materials_examined">
<paragraph id="8BCC368BFFF2FFE255DEFB6AFC25FB24" blockId="1.[136,1452,1215,1346]" box="[136,906,1252,1274]" pageId="1" pageNumber="102">
<emphasis id="B907EA99FFF2FFE255DEFB6AFF6DFB24" bold="true" box="[136,194,1252,1273]" pageId="1" pageNumber="102">Type:</emphasis>
<materialsCitation id="3B1B3CD6FFF2FFE2559EFB6AFC2AFB24" box="[200,901,1252,1274]" collectionCode="BO, K, L" country="United States of America" location="Viqueque" pageId="1" pageNumber="102" specimenCount="2" stateProvince="Viqueque" typeStatus="holotype">
East Timor,
<collectingRegion id="49B7F869FFF2FFE2546BFB6AFE34FB27" box="[317,411,1252,1274]" country="Timor-Leste" name="Viqueque" pageId="1" pageNumber="102">Viqueque</collectingRegion>
,
<emphasis id="B907EA99FFF2FFE254F3FB6AFD8AFB24" box="[421,549,1252,1273]" italics="true" pageId="1" pageNumber="102">Widjaja 6535</emphasis>
(
<typeStatus id="54C88829FFF2FFE25765FB6AFD3FFB27" box="[563,656,1252,1274]" pageId="1" pageNumber="102" type="holotype">Holotype</typeStatus>
:
<collectionCode id="ED62AE4EFFF2FFE257CFFB6AFD15FB24" box="[665,698,1252,1273]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15186" name="Herbarium Bogoriense" pageId="1" pageNumber="102" type="Herbarium">BO</collectionCode>
;
<typeStatus id="54C88829FFF2FFE25791FB6AFCB4FB24" box="[711,795,1252,1273]" pageId="1" pageNumber="102" type="isotype">Isotypes</typeStatus>
:
<collectionCode id="ED62AE4EFFF2FFE25672FB6AFC98FB24" box="[804,823,1252,1273]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="1" pageNumber="102" type="Herbarium">K</collectionCode>
,
<collectionCode id="ED62AE4EFFF2FFE25617FB6AFCFFFB24" box="[833,848,1252,1273]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="1" pageNumber="102" type="Herbarium">L</collectionCode>
,
<collectingCountry id="F364761BFFF2FFE2560AFB6AFCD1FB24" box="[860,894,1252,1273]" name="United States of America" pageId="1" pageNumber="102">US</collectingCountry>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BCC368BFFF2FFE255DEFA89FE22FA9C" blockId="1.[136,1452,1215,1346]" pageId="1" pageNumber="102">
<emphasis id="B907EA99FFF2FFE255DEFA89FE60FAC0" bold="true" box="[136,463,1287,1309]" pageId="1" pageNumber="102">Additional specimen examined:</emphasis>
<collectingCountry id="F364761BFFF2FFE25481FA86FD94FAC3" box="[471,571,1288,1310]" name="Singapore" pageId="1" pageNumber="102">Singapore</collectingCountry>
, Pulau Ubin, Butterfly Hill, cultivated,
<date id="FFCD104BFFF2FFE25695FA86FBE5FAC3" box="[963,1098,1288,1310]" pageId="1" pageNumber="102" value="2016-03-29">29 Mar. 2016</date>
,
<emphasis id="B907EA99FFF2FFE25101FA86FACCFAC0" box="[1111,1379,1288,1309]" italics="true" pageId="1" pageNumber="102">
Joseph Lai
<collectionCode id="ED62AE4EFFF2FFE2519CFA86FB50FAC0" box="[1226,1279,1288,1309]" country="Singapore" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15640" lsid="urn:lsid:biocol.org:col:15640" name="Singapore Botanic Gardens" pageId="1" pageNumber="102" type="Herbarium">SING</collectionCode>
2016-070
</emphasis>
(
<collectionCode id="ED62AE4EFFF2FFE25024FA86FA03FAC0" box="[1394,1452,1288,1309]" country="Singapore" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15640" lsid="urn:lsid:biocol.org:col:15640" name="Singapore Botanic Gardens" pageId="1" pageNumber="102" type="Herbarium">SING</collectionCode>
[Barcode: 0194772]).
</paragraph>
<paragraph id="8BCC368BFFF2FFE255DEFAFEFBEFF913" blockId="1.[136,1452,1392,1887]" pageId="1" pageNumber="102">
<emphasis id="B907EA99FFF2FFE255DEFAFEFE7CFA57" bold="true" box="[136,467,1392,1418]" pageId="1" pageNumber="102">Supplementary description:</emphasis>
Pseudospikelets fasciculate at each node of flowering branches, sessile, basally subtended by several gemmiferous bracts,
<quantity id="4C8B9B6EFFF2FFE254B8FA1AFDC7FA72" box="[494,616,1428,1455]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="1" pageNumber="102" unit="mm" value="16.5" valueMax="18.0" valueMin="15.0">15–18 mm</quantity>
long; fertile florets 3, terminal floret vestigial, with only a narrow lemma; prophylls 1–2, 1–2—keeled,
<quantity id="4C8B9B6EFFF2FFE2549FFA36FDE1FA0E" box="[457,590,1464,1491]" metricMagnitude="-3" metricUnit="m" metricValue="4.85" metricValueMax="5.4" metricValueMin="4.3" pageId="1" pageNumber="102" unit="mm" value="4.85" valueMax="5.4" valueMin="4.3">4.3–5.4 mm</quantity>
long, glabrous, keels ciliolate; gemmiferous bracts 3–4, ovate,
<quantity id="4C8B9B6EFFF2FFE25052FA36FAC1FA0E" box="[1284,1390,1464,1491]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="1" pageNumber="102" unit="mm" value="8.5" valueMax="12.0" valueMin="5.0">5–12 mm</quantity>
long, both surfaces glabrous, abaxial surface sometimes sparsely pubescent at the apex, margin ciliate above the middle, 11–21—veined, apex acute, mucronate to aristulate; rachilla segments extremely short; glumes absent; lemma ovate,
<quantity id="4C8B9B6EFFF2FFE255DEF9AAFEABF9E2" box="[136,260,1572,1599]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="1" pageNumber="102" unit="mm" value="13.5" valueMax="15.0" valueMin="12.0">12–15 mm</quantity>
long, both surfaces glabrous, abaxial surface sometimes sparsely pubescent at the apex, margin ciliate above the middle, 21–23—veined, apex acute, mucronate, calluses inconspicuous; palea ca.
<quantity id="4C8B9B6EFFF2FFE251CEF9C6FB53F9BE" box="[1176,1276,1608,1635]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="1" pageNumber="102" unit="mm" value="10.5">10.5 mm</quantity>
long, glabrous, 2—keeled, keels apically ciliate, 3—veined between keels, each side 1—veined, apex retuse; lodicules absent; stamens 6, filaments connate into a tube, anthers ca.
<quantity id="4C8B9B6EFFF2FFE257D7F91EFD76F976" box="[641,729,1680,1707]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="1" pageNumber="102" unit="mm" value="6.5">6.5 mm</quantity>
long, apiculate, the apical extension bearing tiny hairs; style 1, long, ending in a single plumose stigma, ovary with a hairy top. Caryopsis unknown.
</paragraph>
<paragraph id="8BCC368BFFF2FFE255EBF956FC34F883" blockId="1.[136,1452,1392,1887]" pageId="1" pageNumber="102">
<emphasis id="B907EA99FFF2FFE255EBF956FEA6F92F" bold="true" box="[189,265,1752,1778]" pageId="1" pageNumber="102">Notes:</emphasis>
For the first time,
<taxonomicName id="4C734D08FFF2FFE254BCF956FD11F92F" authorityName="Z. Y. Cai &amp; Widjaja" authorityYear="2024" baseAuthorityName="Widjaja, Reinwardtia" baseAuthorityYear="2024" box="[490,702,1752,1778]" class="Liliopsida" family="Poaceae" genus="Gigantochloa" kingdom="Plantae" order="Poales" pageId="1" pageNumber="102" phylum="Tracheophyta" rank="species" species="lako">
<emphasis id="B907EA99FFF2FFE254BCF956FD11F92F" box="[490,702,1752,1778]" italics="true" pageId="1" pageNumber="102">Gigantochloa lako</emphasis>
</taxonomicName>
is found in Ende,
<collectingCountry id="F364761BFFF2FFE256CDF956FC4BF92F" box="[923,996,1752,1778]" name="Indonesia" pageId="1" pageNumber="102">Flores</collectingCountry>
,
<collectingCountry id="F364761BFFF2FFE256A4F956FBF0F92F" box="[1010,1119,1752,1778]" name="Indonesia" pageId="1" pageNumber="102">Indonesia</collectingCountry>
which has never been noted previously. Besides
<collectingCountry id="F364761BFFF2FFE25427F972FE15F8CB" box="[369,442,1788,1814]" name="Indonesia" pageId="1" pageNumber="102">Flores</collectingCountry>
, this species is also found abundantly in West Timor. In
<collectingCountry id="F364761BFFF2FFE2510AF972FB63F8CB" box="[1116,1228,1788,1814]" name="Indonesia" pageId="1" pageNumber="102">Indonesia</collectingCountry>
, this species is not as commonly cultivated as Javanese Black Bamboo (
<taxonomicName id="4C734D08FFF2FFE257B9F8AEFC0EF8E7" authorityName="Widjaja" authorityYear="1987" box="[751,929,1824,1850]" class="Liliopsida" family="Poaceae" genus="Gigantochloa" kingdom="Plantae" order="Poales" pageId="1" pageNumber="102" phylum="Tracheophyta" rank="species" species="atroviolacea">
<emphasis id="B907EA99FFF2FFE257B9F8AEFC0EF8E7" box="[751,929,1824,1850]" italics="true" pageId="1" pageNumber="102">G. atroviolacea</emphasis>
</taxonomicName>
), although it also holds excellent ornamental horticulture values in many Southeast Asian countries (Widjaja 2019).
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="E5E4537434CD9F88E0DC93255776DE8F" ID-DOI="10.11646/phytotaxa.662.3.3" ID-ISSN="1179-3163" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1734537013249" checkinUser="felipe" docAuthor="Jiménez, Marco M., Iturralde, Gabriel A., Mendoza, Johny, Ocupa-Horna, Luis &amp; Garzón-Suárez, Henry X." docDate="2024" docId="03E0F10DF24A00235C94FAB69A973992" docLanguage="en" docName="phytotaxa.662.3.239-250.pdf" docOrigin="Phytotaxa 662 (3)" docSource="https://doi.org/10.11646/phytotaxa.662.3.3" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Meriania judithiae M. M. Jimenez &amp; H. Garzon 2024, sp. nov." docType="treatment" docVersion="1" lastPageNumber="249" masterDocId="FFD98975F24B00295C1CFFFA9F00386F" masterDocTitle="A new species of Meriania (Melastomataceae: Merianieae) from southeastern Ecuador" masterLastPageNumber="250" masterPageNumber="239" pageNumber="240" updateTime="1734537081990" updateUser="GgImagineBatch" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="4F143520517744B98770AD0F76E0CFFA">A new species of Meriania (Melastomataceae: Merianieae) from southeastern Ecuador</mods:title>
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<mods:namePart id="DE9608ADBB2C7535A61941B87620DAFE">Jiménez, Marco M.</mods:namePart>
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<mods:affiliation id="FBB9A0A030BABCBA99EDB4E9B611C63B">Grupo de Investigación en Medio Ambiente y Salud (BIOMAS), Ingeniería en Agroindustria, Facultad de Ingenierías y Ciencias Aplicadas, Universidad de Las Américas, Vía a Nayón, Quito 170124, Ecuador</mods:affiliation>
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<mods:namePart id="DD90F4EEB61F2ADAA7A4D23D9E9626F0">Iturralde, Gabriel A.</mods:namePart>
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<mods:affiliation id="E77556BBD3D13FB467F61ABC5FFBB923">Grupo de Investigación en Medio Ambiente y Salud (BIOMAS), Ingeniería en Agroindustria, Facultad de Ingenierías y Ciencias Aplicadas, Universidad de Las Américas, Vía a Nayón, Quito 170124, Ecuador</mods:affiliation>
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<mods:namePart id="344B23D0A77CAF845FA8EFADBBC1F4CD">Mendoza, Johny</mods:namePart>
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<mods:affiliation id="03A51ABF0EBB5AC16D7CCE9F80D81F5C">Amichi Lounge, vía Gualaquiza - San Juan Bosco, E 45 s / n., Morona Santiago, Ecuador</mods:affiliation>
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<mods:namePart id="D8EE979D120ACA3EB999686E5219DC9A">Ocupa-Horna, Luis</mods:namePart>
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<mods:affiliation id="D29B8CA91315BBC44CBC67F45AFB1E0B">Departamento de Orquideología, Centro de Investigación en Biología Tropical y Conservación - CINBIOTYC, Cal. Nestor Martos Mza. e Lote. 16 A. H. Almirante Miguel Grau II, Piura, Peru &amp; Departamento de Gimnospermas y Monocotiledóneas, Museo de Historia Natural, Universidad Nacional Mayor de San Marcos, Av. Arenales 1256, Jesús María, Lima, Peru &amp; Grupo Científico Calaway Dodson: Investigación y Conservación de Orquídeas del Ecuador, Quito, 170510, Pichincha, Ecuador</mods:affiliation>
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<mods:namePart id="6A09471876B19A038F5EE4F83FBFF0F6">Garzón-Suárez, Henry X.</mods:namePart>
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<mods:affiliation id="5F5FDBF9B05E43DD124D37B2DC8017F4">Grupo Científico Calaway Dodson: Investigación y Conservación de Orquídeas del Ecuador, Quito, 170510, Pichincha, Ecuador &amp; Jungle Dave’s Science Foundation, San Juan Bosco, Ecuador &amp; Herbario HUTPL, Departamento de Ciencias Biológicas, Universidad Técnica Particular de Loja, San Cayetano Alto s / n 11 - 01 - 608, Loja, Ecuador</mods:affiliation>
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<mods:title id="C708D56062E51F8DEAA29FA3805C15E0">Phytotaxa</mods:title>
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<mods:date id="AB4A7AF800CDA39092A3D093B25C0B74">2024</mods:date>
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<taxonomicName id="4C493B98F24A00285C94FAB69D983D08" ID-CoL="CKJLL" authority="M.M.Jimenez &amp; H.Garzon" authorityName="M. M. Jimenez &amp; H. Garzon" authorityYear="2024" box="[136,664,1356,1383]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="240" phylum="Tracheophyta" rank="species" species="judithiae" status="sp. nov.">
<emphasis id="B93D9C09F24A00285C94FAB69E613D08" bold="true" box="[136,353,1356,1383]" italics="true" pageId="1" pageNumber="240">Meriania judithiae</emphasis>
M.M.Jiménez &amp; H.Garzón
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,
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<emphasis id="B93D9C09F24A00285EBFFAB79DF23D09" box="[675,754,1357,1382]" italics="true" pageId="1" pageNumber="240">sp. nov</emphasis>
.
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(
<figureCitation id="13725C9EF24A00285F15FAB69C563D08" box="[777,854,1356,1383]" captionStart="FIGURE 1" captionStartId="3.[136,229,1921,1943]" captionTargetBox="[183,1404,314,1894]" captionTargetId="figure-104@3.[183,1404,314,1894]" captionTargetPageId="3" captionText="FIGURE 1. Meriania judithiae. A. Branch with inflorescence. B. Flower with a close-up of the indumentum on the hypanthium (B1). C. Petals, abaxial (left) and adaxial (right) views. D. Flower without petals. E. Hypanthium and calyx with style. F. Ovary with style. G. Longitudinal section of the calyx, hypanthium and ovary. H. Calyx, hypanthium and ovary, top view. I. Stamen. J.Anther with connective appendage viewed from the top. K. Close-up of the connective appendage. L. Mature fruit. Prepared by L. Ocupa-Horna from photos by H. Garzón-Suárez and Nelson Espinosa-Ortega (B1) from the holotype." pageId="1" pageNumber="240">Figs. 1</figureCitation>
,
<figureCitation id="13725C9EF24A00285F78FAB79C743D08" box="[868,884,1357,1383]" captionStart="FIGURE 2" captionStartId="4.[136,229,1865,1887]" captionTargetBox="[150,1434,209,1832]" captionTargetId="figure-27@4.[138,1448,190,1841]" captionTargetPageId="4" captionText="FIGURE 2. Meriania judithiae. A. Habit. B. Type of the inflorescences. C. Flower with floral bud. D. Hypanthium and calyx showing stamens and style, longitudinal section. E. Pedicel, ovary and style. F. Stamens, lateral and upper views. G. Calyx, hypanthium and ovary, top view. Drawing by J. Mendoza from the holotype." pageId="1" pageNumber="240">2</figureCitation>
,
<figureCitation id="13725C9EF24A00285F9CFAB79C903D08" box="[896,912,1357,1383]" captionStart="FIGURE 4" captionStartId="6.[136,229,1877,1899]" captionTargetBox="[155,1432,194,1849]" captionTargetId="figure-27@6.[151,1436,190,1853]" captionTargetPageId="6" captionText="FIGURE 4. Meriania judithiae in situ. A. Habitat of the species. B. Terminal branches with inflorescences. C and D. Detail of the inflorescences. Photographs by H. Garzón-Suárez." pageId="1" pageNumber="240">4</figureCitation>
,
<figureCitation id="13725C9EF24A00285F80FAB69CBE3D08" box="[924,958,1356,1383]" captionStart="FIGURE 5" captionStartId="7.[136,229,1877,1899]" captionTargetBox="[155,1432,194,1849]" captionTargetId="figure-22@7.[151,1436,190,1853]" captionTargetPageId="7" captionText="FIGURE 5. Meriania species with 5-nerved leaves and campanulate corollas. A. Meriania judithiae M.M. Jiménez &amp; H. Garzón based on the type (Ecuador, Morona Santiago); B. M. aff. cuneifolia Wurdack based on M. Jiménez 1803 (Ecuador, Morona Santiago); C. M. bongarana Rob. Fern., R. Goldenb. &amp; Michelang. based on the type (Peru, Amazonas); D. M. dimorphanthera Wurdack based on H. Mendoza 16675 (Colombia, Cundinamarca). Photos: by Henry Garzón-Suárez (A, B), Robin Fernandez-Hilario (C) and Humberto Mendoza-Cifuentes (D)." pageId="1" pageNumber="240">5A</figureCitation>
–
<figureCitation id="13725C9EF24A00285FD2FAB79CDC3D08" box="[974,988,1357,1383]" captionStart="FIGURE 7" captionStartId="8.[136,229,1993,2015]" captionTargetBox="[222,1366,1163,1969]" captionTargetId="figure-107@8.[222,1366,1163,1969]" captionTargetPageId="8" captionText="FIGURE 7. Comparison of the holotype specimens of Meriania judithiae (left) and M. cuneifolia subsp. cuneifolia (right). Holotype specimen of Meriania cuneifolia Gleason, collected in Ecuador, Morona Santiago, “Río Santa Bárbara near Río Tintas” by J.A. Steyermark in 1943. Reproduced with permission from the New York Botanical Garden Herbarium." pageId="1" pageNumber="240">7</figureCitation>
)
</heading>
</paragraph>
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<emphasis id="B93D9C09F24A00285C94FA6D9FBA3DC3" bold="true" box="[136,186,1431,1452]" pageId="1" pageNumber="240">Type</emphasis>
:—
<collectingCountry id="F35E008BF24A00285CC5FA6D9E2B3DC2" box="[217,299,1431,1453]" name="Ecuador" pageId="1" pageNumber="240">Ecuador</collectingCountry>
.
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: San Juan Bosco. San Juan Bosco, Jungle Dave’s sendero al Cerro
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(fl.),
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. Garzón &amp;
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. Jiménez 216
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(
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:
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14847!).
</paragraph>
</subSubSection>
<subSubSection id="C3531390F24A00285C94FA279E573EEF" pageId="1" pageNumber="240" type="diagnosis">
<paragraph id="8BF6401BF24A00285C94FA279E573EEF" blockId="1.[136,1452,1431,1664]" pageId="1" pageNumber="240">
<emphasis id="B93D9C09F24A00285C94FA279FF13D9C" bold="true" box="[136,241,1501,1523]" pageId="1" pageNumber="240">Diagnosis.</emphasis>
This species is most similar to
<taxonomicName id="4C493B98F24A00285E3AFA249C913D9D" baseAuthorityName="Gleason" baseAuthorityYear="1947" box="[550,913,1501,1523]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="240" phylum="Tracheophyta" rank="subSpecies" species="cuneifolia" subSpecies="cuneifolia">
<emphasis id="B93D9C09F24A00285E3AFA249DE63D9D" box="[550,742,1501,1523]" italics="true" pageId="1" pageNumber="240">Meriania cuneifolia</emphasis>
subsp.
<emphasis id="B93D9C09F24A00285F2DFA279C913D9D" box="[817,913,1501,1522]" italics="true" pageId="1" pageNumber="240">cuneifolia</emphasis>
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but differs by the sparse furfuraceous indumentum (
<emphasis id="B93D9C09F24A0028598DFA249AAC3D9C" box="[1425,1452,1502,1523]" italics="true" pageId="1" pageNumber="240">vs.</emphasis>
hirsute), the much shorter petioles 0.3–1.0 cm long (
<emphasis id="B93D9C09F24A00285EDEF9FB9DDD3E79" box="[706,733,1537,1558]" italics="true" pageId="1" pageNumber="240">vs.</emphasis>
2.0–3.0 cm long), the leaf blades with attenuate bases extending into the petiole as wings (
<emphasis id="B93D9C09F24A00285D75F9DE9E833E56" box="[361,387,1572,1593]" italics="true" pageId="1" pageNumber="240">vs.</emphasis>
long-cuneate), the shorter calyx lobes
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long (
<emphasis id="B93D9C09F24A00285FADF9DE9CCC3E56" box="[945,972,1572,1593]" italics="true" pageId="1" pageNumber="240">vs.</emphasis>
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long) each with a conical projection that does not surpass the apex of the lobe (
<emphasis id="B93D9C09F24A00285E31F9BD9D483E33" box="[557,584,1607,1628]" italics="true" pageId="1" pageNumber="240">vs.</emphasis>
distinct claw-shaped projections extending further the lobes), and the isomorphic stamens (
<emphasis id="B93D9C09F24A00285CD9F9909FE03E10" box="[197,224,1642,1663]" italics="true" pageId="1" pageNumber="240">vs.</emphasis>
dimorphic).
</paragraph>
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<subSubSection id="C3531390F24A002B5C94F9569E733CEA" lastPageId="2" lastPageNumber="241" pageId="1" pageNumber="240" type="description">
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<emphasis id="B93D9C09F24A00285C94F9569E1A3EA9" bold="true" box="[136,282,1708,1734]" pageId="1" pageNumber="240">Description.</emphasis>
<emphasis id="B93D9C09F24A00285D39F9569E683EA9" box="[293,360,1708,1734]" italics="true" pageId="1" pageNumber="240">Shrub</emphasis>
up to
<quantity id="4CB1EDFEF24A00285DA2F9579EEE3EA8" box="[446,494,1709,1735]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="240" unit="m" value="4.0">4 m</quantity>
tall; indumentum consisting of a combination of very short, brownish, caducous trichomes,
<quantity id="4CB1EDFEF24A00285D19F92A9E8D3E85" box="[261,397,1744,1770]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="9.0" metricValueMin="2.0" pageId="1" pageNumber="240" unit="mm" value="0.55" valueMax="0.9" valueMin="0.2">0.2–0.9 mm</quantity>
long, elongated hairs with greatly roughened (substellate) base and dendritic hairs with short axis and few-moderate number of terete arms, these arranged sparsely on young branches, petioles; or with a stellate, scaly base, sparsely on the underside of leaf blades, densely on inflorescences, hypanthia and calyces.
<emphasis id="B93D9C09F24A002858E5F8ED9AAC3F5F" box="[1273,1452,1814,1840]" italics="true" pageId="1" pageNumber="240">Young branches</emphasis>
terete,
<quantity id="4CB1EDFEF24A00285CC8F8C39E5B3F3B" box="[212,347,1849,1876]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.8" metricValueMin="3.2" pageId="1" pageNumber="240" unit="mm" value="4.5" valueMax="5.8" valueMin="3.2">3.2–5.8 mm</quantity>
in diameter, furfuraceous becoming glabrous with age, nodes with faint interpetiolar lines, without flaps.
<emphasis id="B93D9C09F24A00285CCDF8A79E1F3F19" box="[209,287,1885,1910]" italics="true" pageId="1" pageNumber="240">Leaves</emphasis>
opposite, anisopyllous, decussate; petioles very short, terete, 0.3–1.0 cm long, furfuraceous; blades coriaceous, large leaves 15.5–27.1 ×
<quantity id="4CB1EDFEF24A00285E26F87A9DBE3FF5" box="[570,702,1920,1946]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" metricValueMax="8.2" metricValueMin="3.8" pageId="1" pageNumber="240" unit="cm" value="6.0" valueMax="8.2" valueMin="3.8">3.8–8.2 cm</quantity>
, small leaves (35–60% smaller) 6.1–17.7 × 1.2–5.0 cm, elliptic, apex long-acuminate, base attenuate continuing down the petiole as slightly revolute wings; margin crenulate at the apical half, venation acrodromous and suprabasal, with 2 pairs of lateral (secondaries) nerves, first pair diverging
<quantity id="4CB1EDFEF24A00285C94F8109FF4306B" box="[136,244,2026,2052]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" metricValueMax="4.1" metricValueMin="0.7" pageId="1" pageNumber="240" unit="mm" value="24.0" valueMax="41.0" valueMin="7.0">7–41 mm</quantity>
from the base of the blade, second pair diverging
<quantity id="4CB1EDFEF24A00285F32F8139CA8306B" box="[814,936,2025,2052]" metricMagnitude="-2" metricUnit="m" metricValue="4.75" metricValueMax="8.0" metricValueMin="1.5" pageId="1" pageNumber="240" unit="mm" value="47.5" valueMax="80.0" valueMin="15.0">15–80 mm</quantity>
from the base of the blade, an additional pair of faint submarginal veins running up to leaf apex, tertiary (transversal nerves) 40–60 on each side of the primary veins, percurrent,
<quantity id="4CB1EDFEF249002B5D4CFF649EAC38D6" box="[336,428,158,185]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="7.0" metricValueMin="2.0" pageId="2" pageNumber="241" unit="mm" value="4.5" valueMax="7.0" valueMin="2.0">2–7 mm</quantity>
distant from each other; midvein, secondary and tertiary veins strongly impressed adaxially, salient abaxially, adaxial surface bullate, dark green, shiny, glabrous, olive-green when dry; abaxial surface greenish dirty white when dry, puberulous.
<emphasis id="B93D9C09F249002B5E10FF1C9DAD396F" box="[524,685,230,256]" italics="true" pageId="2" pageNumber="241">Inflorescences</emphasis>
terminal or rarely pseudolateral (overtopped by developing axillary bud), erect,
<quantity id="4CB1EDFEF249002B5D09FEF09E95394A" box="[277,405,266,293]" metricMagnitude="-2" metricUnit="m" metricValue="5.199999999999999" metricValueMax="6.1" metricValueMin="4.3" pageId="2" pageNumber="241" unit="cm" value="5.199999999999999" valueMax="6.1" valueMin="4.3">4.3–6.1 cm</quantity>
long, few-flowered to multiflorous (18–130 flowers) panicles, consisting of 4–6 levels of branching nodes; axis densely pubescent, becoming glabrate in certain parts.
<emphasis id="B93D9C09F249002B580FFED49B7B3927" box="[1043,1147,302,328]" italics="true" pageId="2" pageNumber="241">Peduncle</emphasis>
when present
<quantity id="4CB1EDFEF249002B5936FED49AAB3926" box="[1322,1451,302,329]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.5" metricValueMin="1.2" pageId="2" pageNumber="241" unit="cm" value="1.85" valueMax="2.5" valueMin="1.2">1.2–2.5 cm</quantity>
long, terete.
<emphasis id="B93D9C09F249002B5D0BFEA99E873903" box="[279,391,339,364]" italics="true" pageId="2" pageNumber="241">Main axis</emphasis>
<quantity id="4CB1EDFEF249002B5D93FEA89D0E3902" box="[399,526,338,365]" metricMagnitude="-2" metricUnit="m" metricValue="5.199999999999999" metricValueMax="6.1" metricValueMin="4.3" pageId="2" pageNumber="241" unit="cm" value="5.199999999999999" valueMax="6.1" valueMin="4.3">4.3–6.1 cm</quantity>
long, quadrangular, the proximal node with 1 pair of basal paraclades, the supraproximal with 1 pair of paraclades, the subdistal and distal with an umbel of 4–8 flowers arising from more or less the same point on the pedicel.
<emphasis id="B93D9C09F249002B5DA2FE609D3E39DB" box="[446,574,410,436]" italics="true" pageId="2" pageNumber="241">Paraclades</emphasis>
simple to branching,
<quantity id="4CB1EDFEF249002B5F25FE609CB839DA" box="[825,952,410,437]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.2" metricValueMin="1.3" pageId="2" pageNumber="241" unit="cm" value="1.75" valueMax="2.2" valueMin="1.3">1.3–2.2 cm</quantity>
long; flowers in regular pendulous, umbels of 4–8 flowers at the end.
<emphasis id="B93D9C09F249002B5DACFE459EF939B7" box="[432,505,447,472]" italics="true" pageId="2" pageNumber="241">Bracts</emphasis>
foliaceous, persistent, 14.0–19.6 × 4.0–
<quantity id="4CB1EDFEF249002B5FA2FE449B0C39B6" box="[958,1036,446,473]" metricMagnitude="-2" metricUnit="m" metricValue="6.1" pageId="2" pageNumber="241" unit="cm" value="6.1">6.1 cm</quantity>
, petioles
<quantity id="4CB1EDFEF249002B586AFE449BF439B6" box="[1142,1268,446,473]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="11.0" metricValueMin="7.0" pageId="2" pageNumber="241" unit="cm" value="0.9" valueMax="1.1" valueMin="0.7">0.7–1.1 cm</quantity>
long, shape and indumentum similar to principal leaves; sometimes with one pair of additional bracts on proximal nodes of the main axis, 6.1–12.4 ×
<quantity id="4CB1EDFEF249002B5D23FDFC9EBD3A4E" box="[319,445,518,545]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" metricValueMax="3.5" metricValueMin="1.2" pageId="2" pageNumber="241" unit="cm" value="2.35" valueMax="3.5" valueMin="1.2">1.2–3.5 cm</quantity>
, petioles ca.
<quantity id="4CB1EDFEF249002B5E55FDFC9D943A4E" box="[585,660,518,545]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="241" unit="cm" value="0.7">0.7 cm</quantity>
long.
<emphasis id="B93D9C09F249002B5EC4FDFC9C503A4F" box="[728,848,518,544]" italics="true" pageId="2" pageNumber="241">Bracteoles</emphasis>
absent.
<emphasis id="B93D9C09F249002B5FB5FDFC9B053A4F" box="[937,1029,518,544]" italics="true" pageId="2" pageNumber="241">Flowers</emphasis>
2.3–2.4 ×
<quantity id="4CB1EDFEF249002B5865FDFC9BC73A4E" box="[1145,1223,518,545]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="241" unit="cm" value="0.9">0.9 cm</quantity>
, 5–merous, pendant, with campanulate corollas.
<emphasis id="B93D9C09F249002B5DDFFDD09D223A2B" box="[451,546,554,580]" italics="true" pageId="2" pageNumber="241">Pedicels</emphasis>
<quantity id="4CB1EDFEF249002B5E37FDD09DB53A2A" box="[555,693,554,581]" metricMagnitude="-3" metricUnit="m" metricValue="5.449999999999999" metricValueMax="6.1" metricValueMin="4.8" pageId="2" pageNumber="241" unit="mm" value="5.449999999999999" valueMax="6.1" valueMin="4.8">4.8–6.1 mm</quantity>
long, straight to slightly arcuate, puberulent.
<emphasis id="B93D9C09F249002B58DFFDD09A4D3A2B" box="[1219,1357,554,580]" italics="true" pageId="2" pageNumber="241">Hypanthium</emphasis>
3.4–3.9 ×
<quantity id="4CB1EDFEF249002B5CBDFDB49E2D3A06" box="[161,301,590,617]" metricMagnitude="-3" metricUnit="m" metricValue="3.35" metricValueMax="3.6" metricValueMin="3.1" pageId="2" pageNumber="241" unit="mm" value="3.35" valueMax="3.6" valueMin="3.1">3.1–3.6 mm</quantity>
, cupuliform, slightly costate, outer surface puberulent, inner surface glabrous; torus glabrous.
<emphasis id="B93D9C09F249002B5976FDB49AAB3A07" box="[1386,1451,590,616]" italics="true" pageId="2" pageNumber="241">Calyx</emphasis>
opening regularly, outer surface puberulent, inner surface glabrous; tube
<quantity id="4CB1EDFEF249002B5FD5FD889B523AE2" box="[969,1106,626,653]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="2" pageNumber="241" unit="mm" value="1.0" valueMax="1.5" valueMin="0.5">0.5–1.5 mm</quantity>
long; lobes
<quantity id="4CB1EDFEF249002B58FDFD889A6A3AE2" box="[1249,1386,626,653]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="13.0" metricValueMin="5.0" pageId="2" pageNumber="241" unit="mm" value="0.9" valueMax="1.3" valueMin="0.5">0.5–1.3 mm</quantity>
high, 2.5–3.0 mm wide, subtruncate, margin sinuate, each lobe with a conical projection,
<quantity id="4CB1EDFEF249002B5855FD6C9BD23ADE" box="[1097,1234,662,689]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="2" pageNumber="241" unit="mm" value="0.45" valueMax="0.5" valueMin="0.4">0.4–0.5 mm</quantity>
long free from the apex of the lobe and not surpasses it.
<taxonomicName id="4C493B98F249002B5E3AFD409D7C3ABB" box="[550,636,698,724]" class="Gastropoda" family="Cymbuliidae" genus="Corolla" kingdom="Animalia" order="Pteropoda" pageId="2" pageNumber="241" phylum="Mollusca" rank="genus">
<emphasis id="B93D9C09F249002B5E3AFD409D7C3ABB" box="[550,636,698,724]" italics="true" pageId="2" pageNumber="241">Corolla</emphasis>
</taxonomicName>
campanulate, petals pinkish orange, obscurely multiveined in the adaxial surface, glabrous, concave, 11.1–11.3 ×
<quantity id="4CB1EDFEF249002B5E4FFD249DF93A96" box="[595,761,734,761]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" metricValueMax="1.14" metricValueMin="1.02" pageId="2" pageNumber="241" unit="mm" value="10.8" valueMax="11.4" valueMin="10.2">10.2–11.4 mm</quantity>
, 2.0–
<quantity id="4CB1EDFEF249002B5F24FD249C8F3A96" box="[824,911,734,761]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="241" unit="mm" value="2.4">2.4 mm</quantity>
wide at base, transversely obovate and slightly asymmetric, apex rotund, margin entire, somewhat revolute, base rotund to truncate, concave.
<emphasis id="B93D9C09F249002B5886FCF99BF73B72" box="[1178,1271,771,797]" italics="true" pageId="2" pageNumber="241">Stamens</emphasis>
10, isomorphic, all bent to one side at anthesis giving the flower a zygomorphic appearance; filaments
<quantity id="4CB1EDFEF249002B5825FCDC9BBE3B2E" box="[1081,1214,806,833]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" metricValueMax="5.8" metricValueMin="4.4" pageId="2" pageNumber="241" unit="mm" value="5.1" valueMax="5.8" valueMin="4.4">4.4–5.8 mm</quantity>
long, reddish-purple, paler to the base, flat, glabrous; connectives prolonged below the thecae, reddish-purple, glabrous, with one dorsal conic, crown-shaped appendage,
<quantity id="4CB1EDFEF249002B5E19FC949D5E3BE6" box="[517,606,878,905]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="241" unit="mm" value="0.2">0.2 mm</quantity>
long, acute, red to red-purple consisting of short, several blunt, irregular projections; anther
<quantity id="4CB1EDFEF249002B5D42FC689EE43BC2" box="[350,484,914,941]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" metricValueMax="5.8" metricValueMin="5.4" pageId="2" pageNumber="241" unit="mm" value="5.6" valueMax="5.8" valueMin="5.4">5.4–5.8 mm</quantity>
long, lanceolate, dark purple, glabrous, straight, opening by one dorsally inclined pore, thecae surface slightly corrugated.
<emphasis id="B93D9C09F249002B5E13FC4D9D553BBE" box="[527,597,951,977]" italics="true" pageId="2" pageNumber="241">Ovary</emphasis>
5-locular, superior, free, 3.3 ×
<quantity id="4CB1EDFEF249002B5FADFC4C9B093BBE" box="[945,1033,950,977]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="2" pageNumber="241" unit="mm" value="1.9">1.9 mm</quantity>
, oblong, purple, glabrous, truncate to the apex, whitish to the base, not exceeding the hypanthium length, separated
<quantity id="4CB1EDFEF249002B5FF4FC209B3D3B9A" box="[1000,1085,986,1013]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="241" unit="mm" value="0.5">0.5 mm</quantity>
from the sinuate, glabrous torus; style
<quantity id="4CB1EDFEF249002B5CD9FC049E6A3C76" box="[197,362,1022,1049]" metricMagnitude="-2" metricUnit="m" metricValue="1.47" metricValueMax="1.53" metricValueMin="1.41" pageId="2" pageNumber="241" unit="mm" value="14.7" valueMax="15.3" valueMin="14.1">14.1–15.3 mm</quantity>
long, red, glabrous, slightly incurved at the apex and opposite to the anthers at anthesis; stigma punctiform and minutely papillate,
<quantity id="4CB1EDFEF249002B5E07FBD89D733C52" box="[539,627,1058,1085]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="241" unit="mm" value="0.5">0.5 mm</quantity>
wide, purple.
<emphasis id="B93D9C09F249002B5F06FBD99C5E3C53" box="[794,862,1059,1084]" italics="true" pageId="2" pageNumber="241">Fruits</emphasis>
capsular, with persistent sepals and slightly costate hypanthium, mature ovary 4.0–4.7 ×
<quantity id="4CB1EDFEF249002B5E37FBBC9DB43C0E" box="[555,692,1094,1121]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="4.9" metricValueMin="4.1" pageId="2" pageNumber="241" unit="mm" value="4.5" valueMax="4.9" valueMin="4.1">4.1–4.9 mm</quantity>
, spheroid, sepals
<quantity id="4CB1EDFEF249002B5F67FBBC9B023C0E" box="[891,1026,1094,1121]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="2" pageNumber="241" unit="mm" value="1.7" valueMax="1.8" valueMin="1.6">1.6–1.8 mm</quantity>
high; fruiting pedicels 28.0–
<quantity id="4CB1EDFEF249002B5954FBBC9AAC3C0E" box="[1352,1452,1094,1121]" metricMagnitude="-2" metricUnit="m" metricValue="2.98" pageId="2" pageNumber="241" unit="mm" value="29.8">29.8 mm</quantity>
long.
<emphasis id="B93D9C09F249002B5CD4FB909E083CEB" box="[200,264,1130,1156]" italics="true" pageId="2" pageNumber="241">Seeds</emphasis>
not seen.
</paragraph>
</subSubSection>
<subSubSection id="C3531390F249002B5CA1FB749EC53E87" pageId="2" pageNumber="241" type="distribution">
<paragraph id="8BF6401BF249002B5CA1FB749E0F3D83" blockId="2.[136,1452,158,2093]" pageId="2" pageNumber="241">
<emphasis id="B93D9C09F249002B5CA1FB749EF93CC7" bold="true" box="[189,505,1166,1193]" pageId="2" pageNumber="241">
<emphasis id="B93D9C09F249002B5CA1FB749EDD3CC6" bold="true" box="[189,477,1166,1193]" italics="true" pageId="2" pageNumber="241">Distribution and habitat:</emphasis>
—
</emphasis>
A population of approximately 8 individuals of
<taxonomicName id="4C493B98F249002B5811FB759BE03CC7" authorityName="M. M. Jimenez &amp; H. Garzon" authorityYear="2024" box="[1037,1248,1166,1192]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="241" phylum="Tracheophyta" rank="species" species="judithiae">
<emphasis id="B93D9C09F249002B5811FB759BE03CC7" box="[1037,1248,1166,1192]" italics="true" pageId="2" pageNumber="241">Meriania judithiae</emphasis>
</taxonomicName>
has been found in the evergreen lower montane forests at the Southeastern Cordillera of the Andes, coded as BsBn02 according to the Ecosystem Classification System from Continental
<collectingCountry id="F35E008BF249002B5EA3FB2C9C1C3C9F" box="[703,796,1238,1264]" name="Ecuador" pageId="2" pageNumber="241">Ecuador</collectingCountry>
(Ministerio del Ambiente del
<collectingCountry id="F35E008BF249002B5878FB2C9BC33C9F" box="[1124,1219,1238,1264]" name="Ecuador" pageId="2" pageNumber="241">Ecuador</collectingCountry>
, 2013). This presents a complex hilly topography consisting of steep slopes, valleys, and ravines, with an average annual rainfall shy of
<quantity id="4CB1EDFEF249002B596FFB009FB43D56" metricMagnitude="0" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="241" unit="mm" value="2600.0">2600 mm</quantity>
(
<bibRefCitation id="EFD83DEAF249002B5CD9FAE49E843D57" author="Baez, S. &amp; Santiana, J. &amp; Guevara, J." box="[197,388,1310,1336]" pageId="2" pageNumber="241" pagination="121 - 123" refId="ref5091" refString="Baez, S., Santiana, J. &amp; Guevara, J. (2013) BsBn 02. Bosque siempreverde montano bajo del Sur de la Cordillera Oriental de los Andes. Ministerio del Ambiente del Ecuador. Sistema de Clasificacion de los Ecosistemas del Ecuador Continental de los Andes. Ministerio del Ambiente del Ecuador, Quito, pp. 121 - 123." type="book chapter" year="2013">
Báez
<emphasis id="B93D9C09F249002B5D1AFAE59E3F3D57" box="[262,319,1310,1336]" italics="true" pageId="2" pageNumber="241">et al.</emphasis>
2013
</bibRefCitation>
). The population of the new species is located at about
<quantity id="4CB1EDFEF249002B5FE2FAE49B543D56" box="[1022,1108,1310,1337]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="241" unit="m" value="1800.0">1800 m</quantity>
a.s.l., within a private reserve near San Juan Bosco in the southern part of
<collectingRegion id="498D8EF9F249002B5E63FAB99CB03D33" box="[639,944,1346,1373]" country="Ecuador" name="Morona-Santiago" pageId="2" pageNumber="241">Morona Santiago Province</collectingRegion>
, southeastern
<collectingCountry id="F35E008BF249002B584CFAB89BAD3D33" box="[1104,1197,1346,1372]" name="Ecuador" pageId="2" pageNumber="241">Ecuador</collectingCountry>
(
<figureCitation id="13725C9EF249002B58A2FAB89A0D3D33" box="[1214,1293,1346,1372]" captionStart="FIGURE 3" captionStartId="5.[136,229,1907,1929]" captionTargetBox="[150,1438,202,1870]" captionTargetId="figure-22@5.[138,1449,190,1882]" captionTargetPageId="5" captionText="FIGURE 3. Distribution of Meriania judithiae in southeastern Ecuador. Prepared by Henry X. Garzón-Suárez using ArcGIS." pageId="2" pageNumber="241">Figs. 3</figureCitation>
,
<figureCitation id="13725C9EF249002B5907FAB89A2C3D33" box="[1307,1324,1346,1372]" captionStart="FIGURE 4" captionStartId="6.[136,229,1877,1899]" captionTargetBox="[155,1432,194,1849]" captionTargetId="figure-27@6.[151,1436,190,1853]" captionTargetPageId="6" captionText="FIGURE 4. Meriania judithiae in situ. A. Habitat of the species. B. Terminal branches with inflorescences. C and D. Detail of the inflorescences. Photographs by H. Garzón-Suárez." pageId="2" pageNumber="241">4</figureCitation>
). The local flora is represented by
<taxonomicName id="4C493B98F249002B5D96FA9C9B313DEF" authority="(Vellozo 1829: 176) Martius (1843: 38)" authorityName="Martius" authorityPageNumber="38" authorityYear="1843" baseAuthorityName="Vellozo" baseAuthorityPageNumber="176" baseAuthorityYear="1829" box="[394,1073,1382,1408]" class="Magnoliopsida" family="Meliaceae" genus="Cabralea" kingdom="Plantae" order="Sapindales" pageId="2" pageNumber="241" phylum="Tracheophyta" rank="species" species="canjerana">
<emphasis id="B93D9C09F249002B5D96FA9C9D6E3DEF" box="[394,622,1382,1408]" italics="true" pageId="2" pageNumber="241">Cabralea canjerana</emphasis>
(Vellozo 1829: 176)
<bibRefCitation id="EFD83DEAF249002B5F41FA9C9B313DEF" author="Martius, C. F. P. von" box="[861,1073,1382,1408]" pageId="2" pageNumber="241" pagination="37 - 38" refId="ref5800" refString="Martius, C. F. P. von (1843) Amara. Amara cum principio adstringente. Meliaceae. In: Missouri Botanical Garden (Org.) Systema materiae medicae vegetabilis brasiliensis. Fleischer, Lipsiae, pp. 37 - 38. https: // doi. org / 10.5962 / bhl. title. 9541" type="book chapter" year="1843">Martius (1843: 38)</bibRefCitation>
</taxonomicName>
(
<taxonomicName id="4C493B98F249002B585CFA9C9BBA3DEF" box="[1088,1210,1382,1408]" class="Magnoliopsida" family="Meliaceae" kingdom="Plantae" order="Sapindales" pageId="2" pageNumber="241" phylum="Tracheophyta" rank="family">Meliaceae</taxonomicName>
);
<taxonomicName id="4C493B98F249002B58D1FA9C9E763DCB" authority="Wurdack (1979: 347)" authorityName="Wurdack" authorityPageNumber="347" authorityYear="1979" class="Magnoliopsida" family="Melastomataceae" genus="Blakea" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="241" phylum="Tracheophyta" rank="species" species="subvaginata">
<emphasis id="B93D9C09F249002B58D1FA9C9AAB3DEF" box="[1229,1451,1382,1408]" italics="true" pageId="2" pageNumber="241">Blakea subvaginata</emphasis>
<bibRefCitation id="EFD83DEAF249002B5C94FA709E763DCB" author="Wurdack, J. J." box="[136,374,1418,1444]" pageId="2" pageNumber="241" pagination="347 - 348" refId="ref6704" refString="Wurdack, J. J. (1979) Certamen Melastomataceis XXX. Phytologia 43: 347 - 348." type="journal article" year="1979">Wurdack (1979: 347)</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C493B98F249002B5DB7FA719C643DCA" authority="H. Mendoza (2021: 85)" authorityName="H. Mendoza" authorityPageNumber="85" authorityYear="2021" box="[427,868,1418,1445]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="241" phylum="Tracheophyta" rank="species" species="neillii">
<emphasis id="B93D9C09F249002B5DB7FA719D5A3DCB" box="[427,602,1418,1444]" italics="true" pageId="2" pageNumber="241">Meriania neillii</emphasis>
H. Mendoza (2021: 85)
</taxonomicName>
(
<taxonomicName id="4C493B98F249002B5F6EFA709B3C3DCB" authorityName="Jussieu" authorityYear="1789" box="[882,1084,1418,1444]" class="Magnoliopsida" family="Melastomataceae" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="241" phylum="Tracheophyta" rank="family">Melastomataceae</taxonomicName>
);
<taxonomicName id="4C493B98F249002B5852FA719E5E3DA7" authority="A. C. Smith (1950: 360)" authorityName="A. C. Smith" authorityPageNumber="360" authorityYear="1950" class="Magnoliopsida" family="Ericaceae" genus="Ceratostema" kingdom="Plantae" order="Ericales" pageId="2" pageNumber="241" phylum="Tracheophyta" rank="species" species="charianthum">
<emphasis id="B93D9C09F249002B5852FA719A723DCB" box="[1102,1394,1418,1445]" italics="true" pageId="2" pageNumber="241">Ceratostema charianthum</emphasis>
A.C.
<bibRefCitation id="EFD83DEAF249002B5C94FA549E5E3DA7" author="Smith, A. C." box="[136,350,1454,1481]" pageId="2" pageNumber="241" pagination="317 - 393" refId="ref6262" refString="Smith, A. C. (1950) Studies of South American plants, XII. Contributions from the United States National Herbarium 29: 317 - 393." type="journal article" year="1950">Smith (1950: 360)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C493B98F249002B5D70FA559D253DA6" box="[364,549,1455,1481]" class="Magnoliopsida" family="Ericaceae" genus="Ceratostema" kingdom="Plantae" order="Ericales" pageId="2" pageNumber="241" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B93D9C09F249002B5D70FA559EFB3DA6" box="[364,507,1455,1481]" italics="true" pageId="2" pageNumber="241">Ceratostema</emphasis>
sp.
</taxonomicName>
and
<taxonomicName id="4C493B98F249002B5E7DFA559B2B3DA6" authority="Wilbur &amp; Luteyn" authorityName="Wilbur &amp; Luteyn" box="[609,1067,1454,1481]" class="Magnoliopsida" family="Ericaceae" genus="Disterigma" kingdom="Plantae" order="Ericales" pageId="2" pageNumber="241" phylum="Tracheophyta" rank="species" species="utleyorum">
<emphasis id="B93D9C09F249002B5E7DFA559C5B3DA7" box="[609,859,1454,1480]" italics="true" pageId="2" pageNumber="241">Disterigma utleyorum</emphasis>
Wilbur &amp; Luteyn
</taxonomicName>
in
<bibRefCitation id="EFD83DEAF249002B5849FA559AAC3DA6" author="Luteyn, J. L. &amp; Wilbur, R. L." box="[1109,1452,1454,1481]" pageId="2" pageNumber="241" pagination="255 - 276" refId="ref5752" refString="Luteyn, J. L. &amp; Wilbur, R. L. (1977) New genera and species of Ericaceae (Vaccinieae) from Costa Rica and Panama. Brittonia 29: 255 - 276. https: // doi. org / 10.2307 / 2806199" type="journal article" year="1977">Luteyn &amp; Wilbur (1977: 259)</bibRefCitation>
(
<taxonomicName id="4C493B98F249002B5C8DFA289E033D83" box="[145,259,1490,1516]" class="Magnoliopsida" family="Ericaceae" kingdom="Plantae" order="Ericales" pageId="2" pageNumber="241" phylum="Tracheophyta" rank="family">Ericaceae</taxonomicName>
).
</paragraph>
<paragraph id="8BF6401BF249002B5CA1FA0C9EC53E87" blockId="2.[136,1452,158,2093]" pageId="2" pageNumber="241">
<emphasis id="B93D9C09F249002B5CA1FA0C9D9E3E7F" box="[189,670,1526,1553]" italics="true" pageId="2" pageNumber="241">
<emphasis id="B93D9C09F249002B5CA1FA0C9EAC3E7E" bold="true" box="[189,428,1526,1553]" italics="true" pageId="2" pageNumber="241">Conservation status:</emphasis>
—
<taxonomicName id="4C493B98F249002B5DD5FA0D9D9E3E7F" authorityName="M. M. Jimenez &amp; H. Garzon" authorityYear="2024" box="[457,670,1526,1552]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="241" phylum="Tracheophyta" rank="species" species="judithiae">Meriania judithiae</taxonomicName>
</emphasis>
is known only from the Siete Iglesias Municipal Conservation Area (ACMSI)
<typeStatus id="54F2FEB9F249002B5CE5F9E19E293E5A" box="[249,297,1563,1589]" pageId="2" pageNumber="241">type</typeStatus>
locality near San Juan Bosco. At present, only a few individual living specimens are known. The regions in the eastern foothills of the Andes are acutely threatened by deforestation carried out for cattle pastures, mining activities, slash-and-burn agriculture and infrastructure works. Nonetheless, the new species might be protected because it grows within or near the ACMSI. For a better assessment of its conservation, additional data on its distribution and abundance are required. We recommend
<taxonomicName id="4C493B98F249002B5E53F9519C223EAB" authorityName="M. M. Jimenez &amp; H. Garzon" authorityYear="2024" box="[591,802,1706,1732]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="241" phylum="Tracheophyta" rank="species" species="judithiae">
<emphasis id="B93D9C09F249002B5E53F9519C223EAB" box="[591,802,1706,1732]" italics="true" pageId="2" pageNumber="241">Meriania judithiae</emphasis>
</taxonomicName>
be characterized as Data Defficient (DD) according to the IUCN red list (
<bibRefCitation id="EFD83DEAF249002B5D33F9349EB93E87" author="IUCN" box="[303,441,1742,1768]" pageId="2" pageNumber="241" refId="ref5621" refString="IUCN (2024) Guidelines for Using the IUCN Red List Categories and Criteria. Version 16. Prepared by the Standards and Petitions Committee. Available from: http: // www. iucnredlist. org / documents / RedListGuidelines. pdf (accessed 11 July 2024)" type="book" year="2024">IUCN 2024</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3531390F24900235CA1F9089A973992" lastPageId="10" lastPageNumber="249" pageId="2" pageNumber="241" type="etymology">
<paragraph id="8BF6401BF249002B5CA1F9089FEC3F3B" blockId="2.[136,1452,158,2093]" pageId="2" pageNumber="241">
<emphasis id="B93D9C09F249002B5CA1F9089E423F62" bold="true" box="[189,322,1778,1805]" italics="true" pageId="2" pageNumber="241">Etymology:</emphasis>
—The new species is named after Judith A. Steck, mother of David R. Goucher who has been actively supporting botanical research, environmental education and conservation in the south of
<collectingRegion id="498D8EF9F249002B586BF8ED9AA73F5F" box="[1143,1447,1814,1841]" country="Ecuador" name="Morona-Santiago" pageId="2" pageNumber="241">Morona Santiago province</collectingRegion>
,
<collectingCountry id="F35E008BF249002B5C94F8C09FE83F3B" box="[136,232,1850,1876]" name="Ecuador" pageId="2" pageNumber="241">Ecuador</collectingCountry>
.
</paragraph>
<paragraph id="8BF6401BF249002A5CA1F8A49A1B394B" blockId="2.[136,1452,158,2093]" lastBlockId="3.[136,1452,158,293]" lastPageId="3" lastPageNumber="242" pageId="2" pageNumber="241">
<emphasis id="B93D9C09F249002B5CA1F8A49DB53F17" box="[189,693,1886,1913]" italics="true" pageId="2" pageNumber="241">
<emphasis id="B93D9C09F249002B5CA1F8A49EC53F16" bold="true" box="[189,453,1886,1913]" italics="true" pageId="2" pageNumber="241">Taxonomic discussion:</emphasis>
—
<taxonomicName id="4C493B98F249002B5DFDF8A59DB53F17" authorityName="M. M. Jimenez &amp; H. Garzon" authorityYear="2024" box="[481,693,1886,1912]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="241" phylum="Tracheophyta" rank="species" species="judithiae">Meriania judithiae</taxonomicName>
</emphasis>
belongs to a group of species distinguished by the campanulate, reddish-orange corollas and the lobed calyx, usually with claw-shaped dorsal projections (
<bibRefCitation id="EFD83DEAF249002B5842F8789AA33FF3" author="Fernandez-Hilario, R. &amp; Goldenberg, R. &amp; Michelangeli, F. A." box="[1118,1443,1922,1948]" pageId="2" pageNumber="241" pagination="1 - 16" refId="ref5291" refString="Fernandez-Hilario, R., Goldenberg, R. &amp; Michelangeli, F. A. (2021) Two new species and two new country records for Meriania (Melastomataceae) from northern Peru. Nordic Journal of Botany 39: 1 - 16. https: // doi. org / 10.1111 / njb. 02969" type="journal article" year="2021">
Fernandez-Hilario
<emphasis id="B93D9C09F249002B5933F8799A653FF3" box="[1327,1381,1922,1948]" italics="true" pageId="2" pageNumber="241">et al.</emphasis>
2021
</bibRefCitation>
) (
<figureCitation id="13725C9EF249002B5C8CF85C9FD33FAE" box="[144,211,1958,1985]" captionStart="FIGURE 5" captionStartId="7.[136,229,1877,1899]" captionTargetBox="[155,1432,194,1849]" captionTargetId="figure-22@7.[151,1436,190,1853]" captionTargetPageId="7" captionText="FIGURE 5. Meriania species with 5-nerved leaves and campanulate corollas. A. Meriania judithiae M.M. Jiménez &amp; H. Garzón based on the type (Ecuador, Morona Santiago); B. M. aff. cuneifolia Wurdack based on M. Jiménez 1803 (Ecuador, Morona Santiago); C. M. bongarana Rob. Fern., R. Goldenb. &amp; Michelang. based on the type (Peru, Amazonas); D. M. dimorphanthera Wurdack based on H. Mendoza 16675 (Colombia, Cundinamarca). Photos: by Henry Garzón-Suárez (A, B), Robin Fernandez-Hilario (C) and Humberto Mendoza-Cifuentes (D)." pageId="2" pageNumber="241">Fig. 5</figureCitation>
). The new species is distinguished from other members of the group by the attenuate base of the leaves continuing down the petiole as wings and the presence of conical projections in the lobes of the calyx (
<figureCitation id="13725C9EF249002B5895F8309BD83F8B" box="[1161,1240,1994,2020]" captionStart="FIGURE 6" captionStartId="8.[136,229,1043,1065]" captionTargetBox="[221,1366,190,1018]" captionTargetId="figure-27@8.[221,1366,190,1018]" captionTargetPageId="8" captionText="FIGURE 6. Leaves of Meriania judithiae with diagnostic details. A. Adaxial view with a close-up of the apical, bullate surface (A1) and base of the leaf (A2). B. Abaxial view with a close-up of the apical surface (B1) and basal portion of the leaf (B2) with trichome (B3). Photos by Henry X. Garzón-Suárez and Nelson Espinosa-Ortega (B3)." pageId="2" pageNumber="241">Figs. 6</figureCitation>
). It is most similar to
<taxonomicName id="4C493B98F249002B5CB9F8159EFC3067" baseAuthorityName="Gleason" baseAuthorityYear="1947" box="[165,508,2030,2056]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="241" phylum="Tracheophyta" rank="subSpecies" species="cuneifolia" subSpecies="cuneifolia">
<emphasis id="B93D9C09F249002B5CB9F8159E393067" box="[165,313,2030,2056]" italics="true" pageId="2" pageNumber="241">M. cuneifolia</emphasis>
subsp.
<emphasis id="B93D9C09F249002B5D92F8149EFC3067" box="[398,508,2030,2056]" italics="true" pageId="2" pageNumber="241">cuneifolia</emphasis>
</taxonomicName>
by the distant suprabasal venation and gradually tapering base of the leaves (
<figureCitation id="13725C9EF249002B5945F8149A9D3067" box="[1369,1437,2030,2056]" captionStart="FIGURE 7" captionStartId="8.[136,229,1993,2015]" captionTargetBox="[222,1366,1163,1969]" captionTargetId="figure-107@8.[222,1366,1163,1969]" captionTargetPageId="8" captionText="FIGURE 7. Comparison of the holotype specimens of Meriania judithiae (left) and M. cuneifolia subsp. cuneifolia (right). Holotype specimen of Meriania cuneifolia Gleason, collected in Ecuador, Morona Santiago, “Río Santa Bárbara near Río Tintas” by J.A. Steyermark in 1943. Reproduced with permission from the New York Botanical Garden Herbarium." pageId="2" pageNumber="241">Fig. 7</figureCitation>
); nevertheless,
<taxonomicName id="4C493B98F249002B5D03F7E99EA73043" authorityName="M. M. Jimenez &amp; H. Garzon" authorityYear="2024" box="[287,423,2066,2092]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="241" phylum="Tracheophyta" rank="species" species="judithiae">
<emphasis id="B93D9C09F249002B5D03F7E99EA73043" box="[287,423,2066,2092]" italics="true" pageId="2" pageNumber="241">M. judithiae</emphasis>
</taxonomicName>
can be distinguished from the latter by the elliptic, long-acuminate leaves (
<emphasis id="B93D9C09F249002B58E8F7E99A123043" box="[1268,1298,2067,2092]" italics="true" pageId="2" pageNumber="241">vs.</emphasis>
oblanceolate, abruptly acuminate), adaxially bullate (
<emphasis id="B93D9C09F248002A5E5FFF659D6138D7" box="[579,609,159,184]" italics="true" pageId="3" pageNumber="242">vs.</emphasis>
flat) foliar surface, the shorter hypanthium (3.4–3.9
<emphasis id="B93D9C09F248002A58A4FF659BD638D7" box="[1208,1238,159,184]" italics="true" pageId="3" pageNumber="242">vs.</emphasis>
<quantity id="4CB1EDFEF248002A58C3FF649A3638D6" box="[1247,1334,158,185]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="3" pageNumber="242" unit="mm" value="5.2">5.2 mm</quantity>
long), the calyx lobes that are not ruptured to the torus (
<emphasis id="B93D9C09F248002A5E60FF399D9A38B3" box="[636,666,195,220]" italics="true" pageId="3" pageNumber="242">vs.</emphasis>
irregular ruptured to the torus), the shorter petals (11.1–11.3
<emphasis id="B93D9C09F248002A5926FF399A5838B3" box="[1338,1368,195,220]" italics="true" pageId="3" pageNumber="242">vs.</emphasis>
<quantity id="4CB1EDFEF248002A5942FF389AAC38B2" box="[1374,1452,194,221]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="242" unit="mm" value="13.0">13 mm</quantity>
long), the isomorphic (
<emphasis id="B93D9C09F248002A5D9BFF1D9EA5396F" box="[391,421,231,256]" italics="true" pageId="3" pageNumber="242">vs.</emphasis>
dimorphic) stamens with shorter filaments no longer than
<quantity id="4CB1EDFEF248002A582EFF1C9BB7396E" box="[1074,1207,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" metricValueMax="5.8" metricValueMin="4.4" pageId="3" pageNumber="242" unit="mm" value="5.1" valueMax="5.8" valueMin="4.4">4.4–5.8 mm</quantity>
long (
<emphasis id="B93D9C09F248002A591DFF1D9A1F396F" box="[1281,1311,231,256]" italics="true" pageId="3" pageNumber="242">vs.</emphasis>
<quantity id="4CB1EDFEF248002A593AFF1C9AAC396E" box="[1318,1452,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="7.45" metricValueMax="8.5" metricValueMin="6.4" pageId="3" pageNumber="242" unit="mm" value="7.45" valueMax="8.5" valueMin="6.4">6.4–8.5 mm</quantity>
long), and the shorter thecae of up to
<quantity id="4CB1EDFEF248002A5E3FFEF09DA7394A" box="[547,679,266,293]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" metricValueMax="5.8" metricValueMin="5.4" pageId="3" pageNumber="242" unit="mm" value="5.6" valueMax="5.8" valueMin="5.4">5.4–5.8 mm</quantity>
long (
<emphasis id="B93D9C09F248002A5EF2FEF19C0C394B" box="[750,780,267,292]" italics="true" pageId="3" pageNumber="242">vs.</emphasis>
thecae up to
<quantity id="4CB1EDFEF248002A5F82FEF09B22394A" box="[926,1058,266,293]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" metricValueMax="8.3" metricValueMin="5.3" pageId="3" pageNumber="242" unit="mm" value="6.8" valueMax="8.3" valueMin="5.3">5.3–8.3 mm</quantity>
long) (
<bibRefCitation id="EFD83DEAF248002A586DFEF09A0F394B" author="Gleason, H. A." box="[1137,1295,266,292]" pageId="3" pageNumber="242" pagination="294 - 306" refId="ref5528" refString="Gleason, H. A. (1947) New and noteworthy melastomes, chiefly Ecuadorian. Phytologia 2: 294 - 306." type="journal article" year="1947">Gleason 1947</bibRefCitation>
).
</paragraph>
<caption id="DF361093F248002A5C94F87B9C3A3047" pageId="3" pageNumber="242" startId="3.[136,229,1921,1943]" targetBox="[183,1404,314,1894]" targetPageId="3" targetType="figure">
<paragraph id="8BF6401BF248002A5C94F87B9C3A3047" blockId="3.[136,1452,1921,2088]" pageId="3" pageNumber="242">
<emphasis id="B93D9C09F248002A5C94F87B9FFF3FF8" bold="true" box="[136,255,1921,1943]" pageId="3" pageNumber="242">FIGURE 1.</emphasis>
<emphasis id="B93D9C09F248002A5D1AF8789EC43FF8" box="[262,452,1922,1943]" italics="true" pageId="3" pageNumber="242">
<taxonomicName id="4C493B98F248002A5D1AF8789EC03FF8" authorityName="M. M. Jimenez &amp; H. Garzon" authorityYear="2024" box="[262,448,1922,1943]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="3" pageNumber="242" phylum="Tracheophyta" rank="species" species="judithiae">Meriania judithiae</taxonomicName>
.
</emphasis>
<emphasis id="B93D9C09F248002A5DD6F87B9EE13FF9" bold="true" box="[458,481,1921,1942]" pageId="3" pageNumber="242">A.</emphasis>
Branch with inflorescence.
<emphasis id="B93D9C09F248002A5EE9F8789C0B3FF8" bold="true" box="[757,779,1922,1943]" pageId="3" pageNumber="242">B.</emphasis>
Flower with a close-up of the indumentum on the hypanthium (B1).
<emphasis id="B93D9C09F248002A5C94F85F9FA03FD4" bold="true" box="[136,160,1957,1979]" pageId="3" pageNumber="242">C.</emphasis>
Petals, abaxial (left) and adaxial (right) views.
<emphasis id="B93D9C09F248002A5E75F85C9D813FD4" bold="true" box="[617,641,1958,1979]" pageId="3" pageNumber="242">D.</emphasis>
Flower without petals.
<emphasis id="B93D9C09F248002A5F79F85C9C7B3FD4" bold="true" box="[869,891,1958,1979]" pageId="3" pageNumber="242">E.</emphasis>
Hypanthium and calyx with style.
<emphasis id="B93D9C09F248002A58D1F85C9BDF3FD4" bold="true" box="[1229,1247,1958,1979]" pageId="3" pageNumber="242">F.</emphasis>
Ovary with style.
<emphasis id="B93D9C09F248002A598FF85F9AAC3FD4" bold="true" box="[1427,1452,1957,1979]" pageId="3" pageNumber="242">G.</emphasis>
Longitudinal section of the calyx, hypanthium and ovary.
<emphasis id="B93D9C09F248002A5EB2F8309DC73FB0" bold="true" box="[686,711,1994,2015]" pageId="3" pageNumber="242">H.</emphasis>
Calyx, hypanthium and ovary, top view.
<emphasis id="B93D9C09F248002A5852F8309B5D3FB0" bold="true" box="[1102,1117,1994,2015]" pageId="3" pageNumber="242">I.</emphasis>
Stamen.
<emphasis id="B93D9C09F248002A58A9F8309BC83FB0" bold="true" box="[1205,1224,1994,2015]" pageId="3" pageNumber="242">J.</emphasis>
Anther with connective appendage viewed from the top.
<emphasis id="B93D9C09F248002A5DD8F8149EDD306C" bold="true" box="[452,477,2030,2051]" pageId="3" pageNumber="242">K.</emphasis>
Close-up of the connective appendage.
<emphasis id="B93D9C09F248002A5F42F8149C74306C" bold="true" box="[862,884,2030,2051]" pageId="3" pageNumber="242">L.</emphasis>
Mature fruit. Prepared by L. Ocupa-Horna from photos by
<emphasis id="B93D9C09F248002A5C94F7E89FA13048" bold="true" box="[136,161,2066,2087]" pageId="3" pageNumber="242">H.</emphasis>
Garzón-Suárez and Nelson Espinosa-Ortega (B1) from the holotype.
</paragraph>
</caption>
<caption id="DF361093F24F002D5C94F8B39D863FC8" pageId="4" pageNumber="243" startId="4.[136,229,1865,1887]" targetBox="[150,1434,209,1832]" targetPageId="4" targetType="figure">
<paragraph id="8BF6401BF24F002D5C94F8B39D863FC8" blockId="4.[136,1452,1865,1959]" pageId="4" pageNumber="243">
<emphasis id="B93D9C09F24F002D5C94F8B39FFF3F31" bold="true" box="[136,255,1865,1887]" pageId="4" pageNumber="243">FIGURE 2.</emphasis>
<taxonomicName id="4C493B98F24F002D5D1AF8B09EBB3F31" authorityName="M. M. Jimenez &amp; H. Garzon" authorityYear="2024" box="[262,443,1865,1887]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="4" pageNumber="243" phylum="Tracheophyta" rank="species" species="judithiae">
<emphasis id="B93D9C09F24F002D5D1AF8B09EBB3F31" box="[262,443,1865,1887]" italics="true" pageId="4" pageNumber="243">Meriania judithiae</emphasis>
</taxonomicName>
. A. Habit. B. Type of the inflorescences. C. Flower with floral bud. D. Hypanthium and calyx showing stamens and style, longitudinal section. E. Pedicel, ovary and style. F. Stamens, lateral and upper views. G. Calyx, hypanthium and ovary, top view. Drawing by J. Mendoza from the holotype.
</paragraph>
</caption>
<caption id="DF361093F24E002C5C94F8899A3C3FE6" box="[136,1340,1907,1929]" pageId="5" pageNumber="244" startId="5.[136,229,1907,1929]" targetBox="[150,1438,202,1870]" targetPageId="5" targetType="figure">
<paragraph id="8BF6401BF24E002C5C94F8899A3C3FE6" blockId="5.[136,1340,1907,1929]" box="[136,1340,1907,1929]" pageId="5" pageNumber="244">
<emphasis id="B93D9C09F24E002C5C94F8899FFE3FE7" bold="true" box="[136,254,1907,1929]" pageId="5" pageNumber="244">FIGURE 3.</emphasis>
Distribution of
<taxonomicName id="4C493B98F24E002C5D8BF88E9D4C3FE7" authorityName="M. M. Jimenez &amp; H. Garzon" authorityYear="2024" box="[407,588,1907,1929]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="5" pageNumber="244" phylum="Tracheophyta" rank="species" species="judithiae">
<emphasis id="B93D9C09F24E002C5D8BF88E9D4C3FE7" box="[407,588,1907,1929]" italics="true" pageId="5" pageNumber="244">Meriania judithiae</emphasis>
</taxonomicName>
in southeastern Ecuador. Prepared by Henry X. Garzón-Suárez using ArcGIS.
</paragraph>
</caption>
<caption id="DF361093F24D002F5C94F8AF9D6E3FE1" pageId="6" pageNumber="245" startId="6.[136,229,1877,1899]" targetBox="[155,1432,194,1849]" targetPageId="6" targetType="figure">
<paragraph id="8BF6401BF24D002F5C94F8AF9D6E3FE1" blockId="6.[136,1451,1877,1935]" pageId="6" pageNumber="245">
<emphasis id="B93D9C09F24D002F5C94F8AF9E013F05" bold="true" box="[136,257,1877,1899]" pageId="6" pageNumber="245">FIGURE 4.</emphasis>
<emphasis id="B93D9C09F24D002F5D15F8AC9D093F04" box="[265,521,1877,1899]" italics="true" pageId="6" pageNumber="245">
<taxonomicName id="4C493B98F24D002F5D15F8AC9EC13F05" authorityName="M. M. Jimenez &amp; H. Garzon" authorityYear="2024" box="[265,449,1877,1899]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="6" pageNumber="245" phylum="Tracheophyta" rank="species" species="judithiae">Meriania judithiae</taxonomicName>
in situ
</emphasis>
.
<emphasis id="B93D9C09F24D002F5E0BF8AF9D2F3F05" bold="true" box="[535,559,1877,1898]" pageId="6" pageNumber="245">A.</emphasis>
Habitat of the species.
<emphasis id="B93D9C09F24D002F5F03F8AC9C353F04" bold="true" box="[799,821,1878,1899]" pageId="6" pageNumber="245">B.</emphasis>
Terminal branches with inflorescences.
<emphasis id="B93D9C09F24D002F58DBF8AF9BD83F04" bold="true" box="[1223,1240,1877,1899]" pageId="6" pageNumber="245">C</emphasis>
and
<emphasis id="B93D9C09F24D002F5911F8AC9A253F04" bold="true" box="[1293,1317,1878,1899]" pageId="6" pageNumber="245">D.</emphasis>
Detail of the inflorescences. Photographs by H. Garzón-Suárez.
</paragraph>
</caption>
<caption id="DF361093F24C002E5C94F8AF9E723F94" pageId="7" pageNumber="246" startId="7.[136,229,1877,1899]" targetBox="[155,1432,194,1849]" targetPageId="7" targetType="figure">
<paragraph id="8BF6401BF24C002E5C94F8AF9E723F94" blockId="7.[136,1452,1877,2043]" pageId="7" pageNumber="246">
<emphasis id="B93D9C09F24C002E5C94F8AF9FFE3F04" bold="true" box="[136,254,1877,1899]" pageId="7" pageNumber="246">FIGURE 5.</emphasis>
<taxonomicName id="4C493B98F24C002E5D19F8AC9EAA3F04" box="[261,426,1877,1899]" pageId="7" pageNumber="246">
<emphasis id="B93D9C09F24C002E5D19F8AC9E5E3F04" box="[261,350,1878,1899]" italics="true" pageId="7" pageNumber="246">Meriania</emphasis>
species
</taxonomicName>
with 5-nerved leaves and campanulate corollas.
<emphasis id="B93D9C09F24C002E5F9FF8AF9C9B3F05" bold="true" box="[899,923,1877,1898]" pageId="7" pageNumber="246">A.</emphasis>
<taxonomicName id="4C493B98F24C002E5FBDF8AC9A6E3F05" authority="M. M. Jimenez &amp; H. Garzon" authorityName="M. M. Jimenez &amp; H. Garzon" authorityYear="2024" box="[929,1390,1877,1899]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="246" phylum="Tracheophyta" rank="species" species="judithiae">
<emphasis id="B93D9C09F24C002E5FBDF8AC9B563F05" box="[929,1110,1877,1899]" italics="true" pageId="7" pageNumber="246">Meriania judithiae</emphasis>
M.M. Jiménez &amp; H. Garzón
</taxonomicName>
based on the type (Ecuador, Morona Santiago);
<emphasis id="B93D9C09F24C002E5E34F8809D3E3FE0" bold="true" box="[552,574,1914,1935]" pageId="7" pageNumber="246">B.</emphasis>
<taxonomicName id="4C493B98F24C002E5E5BF8809C523FE0" authority="Wurdack" authorityName="Wurdack" box="[583,850,1913,1935]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" isUncertain="true" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="246" phylum="Tracheophyta" rank="species" species="cuneifolia">
<emphasis id="B93D9C09F24C002E5E5BF8809D623FE0" box="[583,610,1914,1935]" italics="true" pageId="7" pageNumber="246">M.</emphasis>
aff.
<emphasis id="B93D9C09F24C002E5E8FF8839DF33FE1" box="[659,755,1913,1934]" italics="true" pageId="7" pageNumber="246">cuneifolia</emphasis>
Wurdack
</taxonomicName>
based on
<emphasis id="B93D9C09F24C002E5FA0F8809B653FE0" box="[956,1125,1914,1935]" italics="true" pageId="7" pageNumber="246">M. Jiménez 1803</emphasis>
(Ecuador, Morona Santiago);
<emphasis id="B93D9C09F24C002E5988F8839AAB3FE0" bold="true" box="[1428,1451,1913,1935]" pageId="7" pageNumber="246">C.</emphasis>
<taxonomicName id="4C493B98F24C002E5C94F8649D983FDC" authority="Rob. Fern., R. Goldenb. &amp; Michelang." authorityName="Rob. Fern., R. Goldenb. &amp; Michelang." box="[136,664,1949,1971]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="246" phylum="Tracheophyta" rank="species" species="bongarana">
<emphasis id="B93D9C09F24C002E5C94F8649E133FDD" box="[136,275,1949,1971]" italics="true" pageId="7" pageNumber="246">M. bongarana</emphasis>
Rob. Fern., R. Goldenb. &amp; Michelang.
</taxonomicName>
based on the type (Peru, Amazonas);
<emphasis id="B93D9C09F24C002E580DF8649B283FDC" bold="true" box="[1041,1064,1950,1971]" pageId="7" pageNumber="246">D.</emphasis>
<taxonomicName id="4C493B98F24C002E582CF8649A4D3FDC" authority="Wurdack" authorityName="Wurdack" authorityYear="1957" box="[1072,1357,1949,1971]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="246" phylum="Tracheophyta" rank="species" species="dimorphanthera">
<emphasis id="B93D9C09F24C002E582CF8649BEE3FDD" box="[1072,1262,1949,1971]" italics="true" pageId="7" pageNumber="246">M. dimorphanthera</emphasis>
Wurdack
</taxonomicName>
based on
<emphasis id="B93D9C09F24C002E5C94F8389E443FB9" box="[136,324,1985,2007]" italics="true" pageId="7" pageNumber="246">H. Mendoza 16675</emphasis>
(Colombia, Cundinamarca). Photos: by Henry Garzón-Suárez (A, B), Robin Fernandez-Hilario (C) and Humberto Mendoza-Cifuentes (D).
</paragraph>
</caption>
<caption id="DF361093F24300215C94FBE99C2D3C1F" pageId="8" pageNumber="247" startId="8.[136,229,1043,1065]" targetBox="[221,1366,190,1018]" targetPageId="8" targetType="figure">
<paragraph id="8BF6401BF24300215C94FBE99C2D3C1F" blockId="8.[136,1452,1043,1137]" pageId="8" pageNumber="247">
<emphasis id="B93D9C09F24300215C94FBE99FFD3C47" bold="true" box="[136,253,1043,1065]" pageId="8" pageNumber="247">FIGURE 6.</emphasis>
Leaves of
<taxonomicName id="4C493B98F24300215D7BFBEE9D1B3C47" authorityName="M. M. Jimenez &amp; H. Garzon" authorityYear="2024" box="[359,539,1043,1065]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="8" pageNumber="247" phylum="Tracheophyta" rank="species" species="judithiae">
<emphasis id="B93D9C09F24300215D7BFBEE9D1B3C47" box="[359,539,1043,1065]" italics="true" pageId="8" pageNumber="247">Meriania judithiae</emphasis>
</taxonomicName>
with diagnostic details.
<emphasis id="B93D9C09F24300215F1AFBE99C1D3C47" bold="true" box="[774,797,1043,1064]" pageId="8" pageNumber="247">A.</emphasis>
Adaxial view with a close-up of the apical, bullate surface (A1) and base of the leaf (A2).
<emphasis id="B93D9C09F24300215D43FBCD9E753C23" bold="true" box="[351,373,1079,1100]" pageId="8" pageNumber="247">B.</emphasis>
Abaxial view with a close-up of the apical surface (B1) and basal portion of the leaf (B2) with trichome (B3). Photos by Henry X. Garzón-Suárez and Nelson Espinosa-Ortega (B3).
</paragraph>
</caption>
<caption id="DF361093F24300215C94F8339CD43048" pageId="8" pageNumber="247" startId="8.[136,229,1993,2015]" targetBox="[222,1366,1163,1969]" targetPageId="8" targetType="figure">
<paragraph id="8BF6401BF24300215C94F8339CD43048" blockId="8.[136,1452,1993,2087]" pageId="8" pageNumber="247">
<emphasis id="B93D9C09F24300215C94F8339E003FB1" bold="true" box="[136,256,1993,2015]" pageId="8" pageNumber="247">FIGURE 7.</emphasis>
Comparison of the holotype specimens of
<taxonomicName id="4C493B98F24300215EB0F8309C633FB1" authorityName="M. M. Jimenez &amp; H. Garzon" authorityYear="2024" box="[684,867,1993,2015]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="8" pageNumber="247" phylum="Tracheophyta" rank="species" species="judithiae">
<emphasis id="B93D9C09F24300215EB0F8309C633FB1" box="[684,867,1993,2015]" italics="true" pageId="8" pageNumber="247">Meriania judithiae</emphasis>
</taxonomicName>
(left) and
<taxonomicName id="4C493B98F24300215FD3F8309BFE3FB1" baseAuthorityName="Gleason" baseAuthorityYear="1947" box="[975,1278,1993,2015]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="8" pageNumber="247" phylum="Tracheophyta" rank="subSpecies" species="cuneifolia" subSpecies="cuneifolia">
<emphasis id="B93D9C09F24300215FD3F8309B513FB1" box="[975,1105,1993,2015]" italics="true" pageId="8" pageNumber="247">M. cuneifolia</emphasis>
subsp.
<emphasis id="B93D9C09F24300215882F8339BFE3FB1" box="[1182,1278,1993,2014]" italics="true" pageId="8" pageNumber="247">cuneifolia</emphasis>
</taxonomicName>
(right). Holotype specimen of
<taxonomicName id="4C493B98F24300215D1CF8149D14306C" authority="Gleason" authorityName="Gleason" authorityYear="1947" box="[256,532,2029,2051]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="8" pageNumber="247" phylum="Tracheophyta" rank="species" species="cuneifolia">
<emphasis id="B93D9C09F24300215D1CF8149EBE306D" box="[256,446,2029,2051]" italics="true" pageId="8" pageNumber="247">Meriania cuneifolia</emphasis>
Gleason
</taxonomicName>
, collected in Ecuador, Morona Santiago, “Río Santa Bárbara near Río Tintas” by J.A. Steyermark in 1943. Reproduced with permission from the New York Botanical Garden Herbarium.
</paragraph>
</caption>
<paragraph id="8BF6401BF24200205CA1FF649E7839DB" blockId="9.[136,1452,158,724]" pageId="9" pageNumber="248">
Both species have been reported in the southeastern Ecuadorian Andes and have not been found growing sympatrically.
<taxonomicName id="4C493B98F24200205D2CFF399D0738B3" authorityName="M. M. Jimenez &amp; H. Garzon" authorityYear="2024" box="[304,519,194,220]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="9" pageNumber="248" phylum="Tracheophyta" rank="species" species="judithiae">
<emphasis id="B93D9C09F24200205D2CFF399D0738B3" box="[304,519,194,220]" italics="true" pageId="9" pageNumber="248">Meriania judithiae</emphasis>
</taxonomicName>
grows at elevations around
<quantity id="4CB1EDFEF24200205F49FF389CAE38B2" box="[853,942,194,221]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="9" pageNumber="248" unit="m" value="1800.0">1800 m</quantity>
where it is restricted to the lower montane forests near the township of San Juan Bosco in
<collectingRegion id="498D8EF9F24200205EDCFF1D9CF8396F" box="[704,1016,230,257]" country="Ecuador" name="Morona-Santiago" pageId="9" pageNumber="248">Morona Santiago Province</collectingRegion>
. On the other hand, according to the specimens of
<emphasis id="B93D9C09F24200205D39FEF19D0E394B" box="[293,526,266,292]" italics="true" pageId="9" pageNumber="248">J. Steyermark 53543</emphasis>
(NY-00547958) mentioned by Wurdack (1980) and
<emphasis id="B93D9C09F24200205879FEF19AAB394B" box="[1125,1451,266,293]" italics="true" pageId="9" pageNumber="248">C. Chimbo &amp; C. Chamba 89</emphasis>
(MO-1183106),
<taxonomicName id="4C493B98F24200205D5FFED59DA53927" baseAuthorityName="Gleason" baseAuthorityYear="1947" box="[323,677,302,328]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="9" pageNumber="248" phylum="Tracheophyta" rank="subSpecies" species="cuneifolia" subSpecies="cuneifolia">
<emphasis id="B93D9C09F24200205D5FFED59EDB3927" box="[323,475,302,328]" italics="true" pageId="9" pageNumber="248">M. cuneifolia</emphasis>
subsp.
<emphasis id="B93D9C09F24200205E29FED49DA53927" box="[565,677,302,328]" italics="true" pageId="9" pageNumber="248">cuneifolia</emphasis>
</taxonomicName>
inhabits higher elevations from
<quantity id="4CB1EDFEF24200205802FED49BD73926" box="[1054,1239,302,329]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="9" pageNumber="248" unit="m" value="2150.0" valueMax="2200.0" valueMin="2100.0">2100 to 2200 m</quantity>
where it grows in wet montane forests of the
<collectingRegion id="498D8EF9F24200205DDCFEA99D833903" box="[448,643,338,365]" country="Ecuador" name="Morona-Santiago" pageId="9" pageNumber="248">Morona Santiago</collectingRegion>
and
<collectingRegion id="498D8EF9F24200205EA0FEA99C8F3903" box="[700,911,338,365]" country="Ecuador" name="Zamora-Chinchipe" pageId="9" pageNumber="248">Zamora Chinchipe</collectingRegion>
provinces. Considering the different ecological preferences, it is improbable there will be consistent gene flow between the two locations despite both being present in southeast
<collectingCountry id="F35E008BF24200205D08FE609E7439DB" box="[276,372,410,436]" name="Ecuador" pageId="9" pageNumber="248">Ecuador</collectingCountry>
.
</paragraph>
<paragraph id="8BF6401BF24200205CA1FE449EED3ABB" blockId="9.[136,1452,158,724]" pageId="9" pageNumber="248">
The new species is also similar to the Peruvian
<taxonomicName id="4C493B98F24200205EC4FE459CC339B7" box="[728,963,446,472]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="9" pageNumber="248" phylum="Tracheophyta" rank="species" species="bongarana">
<emphasis id="B93D9C09F24200205EC4FE459CC339B7" box="[728,963,446,472]" italics="true" pageId="9" pageNumber="248">Meriania bongarana</emphasis>
</taxonomicName>
and the Colombian
<taxonomicName id="4C493B98F242002058B5FE459A8539B7" authorityName="Wurdack" authorityYear="1957" box="[1193,1413,446,472]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="9" pageNumber="248" phylum="Tracheophyta" rank="species" species="dimorphanthera">
<emphasis id="B93D9C09F242002058B5FE459A8539B7" box="[1193,1413,446,472]" italics="true" pageId="9" pageNumber="248">M. dimorphanthera</emphasis>
</taxonomicName>
, as they are also characterized by leaves mostly elliptic, anisophyllous, acuminate, with two pairs of secondaries (lateral nerves) and the acrodromous and suprabasal venation. However,
<taxonomicName id="4C493B98F24200205F64FDFD9B043A4F" authorityName="M. M. Jimenez &amp; H. Garzon" authorityYear="2024" box="[888,1028,518,544]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="9" pageNumber="248" phylum="Tracheophyta" rank="species" species="judithiae">
<emphasis id="B93D9C09F24200205F64FDFD9B043A4F" box="[888,1028,518,544]" italics="true" pageId="9" pageNumber="248">M. judithiae</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="4C493B98F24200205914FDFD9AAB3A4F" box="[1288,1451,518,544]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="9" pageNumber="248" phylum="Tracheophyta" rank="species" species="bongarana">
<emphasis id="B93D9C09F24200205914FDFD9AAB3A4F" box="[1288,1451,518,544]" italics="true" pageId="9" pageNumber="248">M. bongarana</emphasis>
</taxonomicName>
by the attenuate leaf base (
<emphasis id="B93D9C09F24200205DA6FDD19ED83A2B" box="[442,472,555,580]" italics="true" pageId="9" pageNumber="248">vs.</emphasis>
cuneate), the bullate adaxial surface of the leaves (
<emphasis id="B93D9C09F2420020583DFDD19B3F3A2B" box="[1057,1087,555,580]" italics="true" pageId="9" pageNumber="248">vs.</emphasis>
flat), the shorter hypanthium of
<quantity id="4CB1EDFEF24200205C94FDB49E103A06" box="[136,272,590,617]" metricMagnitude="-3" metricUnit="m" metricValue="3.65" metricValueMax="3.9" metricValueMin="3.4" pageId="9" pageNumber="248" unit="mm" value="3.65" valueMax="3.9" valueMin="3.4">3.4–3.9 mm</quantity>
long (
<emphasis id="B93D9C09F24200205D40FDB59E7A3A07" box="[348,378,591,616]" italics="true" pageId="9" pageNumber="248">vs.</emphasis>
6.5–7.0 mm long), the calyx with subtruncate lobes (
<emphasis id="B93D9C09F24200205FC0FDB59CFB3A07" box="[988,1019,591,616]" italics="true" pageId="9" pageNumber="248">vs.</emphasis>
acute), each with a conical projection (
<emphasis id="B93D9C09F24200205C8DFD899FB03AE3" box="[145,176,627,652]" italics="true" pageId="9" pageNumber="248">vs.</emphasis>
claw-shaped dorsal projection), the reddish-purple petals (
<emphasis id="B93D9C09F24200205F4EFD899C713AE3" box="[850,881,627,652]" italics="true" pageId="9" pageNumber="248">vs.</emphasis>
pink), the stamens with shorter filaments
<quantity id="4CB1EDFEF2420020594BFD889FB43ADE" metricMagnitude="-3" metricUnit="m" metricValue="5.6" metricValueMax="5.8" metricValueMin="5.4" pageId="9" pageNumber="248" unit="mm" value="5.6" valueMax="5.8" valueMin="5.4">5.4–5.8 mm</quantity>
long (
<emphasis id="B93D9C09F24200205D1FFD6D9E213ADF" box="[259,289,663,688]" italics="true" pageId="9" pageNumber="248">vs.</emphasis>
8.0–
<quantity id="4CB1EDFEF24200205D41FD6C9EB53ADE" box="[349,437,662,689]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="9" pageNumber="248" unit="mm" value="8.5">8.5 mm</quantity>
long), and the red, shorter style
<quantity id="4CB1EDFEF24200205F28FD6C9CDA3ADE" box="[820,986,662,689]" metricMagnitude="-2" metricUnit="m" metricValue="1.47" metricValueMax="1.53" metricValueMin="1.41" pageId="9" pageNumber="248" unit="mm" value="14.7" valueMax="15.3" valueMin="14.1">14.1–15.3 mm</quantity>
long (
<emphasis id="B93D9C09F24200205835FD6D9B473ADF" box="[1065,1095,663,688]" italics="true" pageId="9" pageNumber="248">vs.</emphasis>
magenta, 25.5–27.0 mm long) (
<bibRefCitation id="EFD83DEAF24200205C93FD409EE23ABB" author="Fernandez-Hilario, R. &amp; Gonzales, R. &amp; Villanueva-Espinoza, R. &amp; Lajo, L. &amp; Sato, A. A. W. &amp; Paredes-Burneo, D. &amp; Pillaca-Huacre, L. &amp; Michelangeli, F. A. &amp; Goldenberg, R." box="[143,482,698,724]" pageId="9" pageNumber="248" pagination="39 - 74" refId="ref5349" refString="Fernandez-Hilario, R., Gonzales, R. del P. R., Villanueva-Espinoza, R., Lajo, L., Sato, A. A. W., Paredes-Burneo, D., Pillaca-Huacre, L., Michelangeli, F. A. &amp; Goldenberg, R. (2022) Nine new species and a new country record for Meriania (Melastomataceae) from Peru. Willdenowia 52: 39 - 74. https: // doi. org / 10.3372 / wi. 52.52103" type="journal article" year="2022">
Fernandez-Hilario
<emphasis id="B93D9C09F24200205D7AFD419E9E3ABB" box="[358,414,698,724]" italics="true" pageId="9" pageNumber="248">et al.</emphasis>
2022
</bibRefCitation>
).
</paragraph>
<caption id="DF361093F24200205C94FCFC9D023B50" pageId="9" pageNumber="248" startId="9.[136,216,774,795]" targetType="table">
<paragraph id="8BF6401BF24200205C94FCFC9D023B50" blockId="9.[136,1451,774,832]" pageId="9" pageNumber="248">
<emphasis id="B93D9C09F24200205C94FCFC9FEF3B74" bold="true" box="[136,239,774,795]" pageId="9" pageNumber="248">TABLE 1.</emphasis>
Summary of principal differences between
<taxonomicName id="4C493B98F24200205E91FCFC9C363B74" box="[653,822,774,795]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="9" pageNumber="248" phylum="Tracheophyta" rank="species" species="judithia">
<emphasis id="B93D9C09F24200205E91FCFC9C363B74" box="[653,822,774,795]" italics="true" pageId="9" pageNumber="248">Meriania judithia</emphasis>
</taxonomicName>
e,
<taxonomicName id="4C493B98F24200205F57FCFC9B713B74" baseAuthorityName="Gleason" baseAuthorityYear="1947" box="[843,1137,774,796]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="9" pageNumber="248" phylum="Tracheophyta" rank="subSpecies" species="cuneifolia" subSpecies="cuneifolia">
<emphasis id="B93D9C09F24200205F57FCFC9CCA3B74" box="[843,970,774,795]" italics="true" pageId="9" pageNumber="248">M. cuneifolia</emphasis>
subsp.
<emphasis id="B93D9C09F2420020580DFCFC9B713B74" box="[1041,1137,774,795]" italics="true" pageId="9" pageNumber="248">cuneifolia</emphasis>
</taxonomicName>
,
<taxonomicName id="4C493B98F24200205860FCFC9AA53B74" authorityName="Wurdack" authorityYear="1976" box="[1148,1445,774,796]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="9" pageNumber="248" phylum="Tracheophyta" rank="subSpecies" species="cuneifolia" subSpecies="subandina">
<emphasis id="B93D9C09F24200205860FCFC9BFB3B74" box="[1148,1275,774,795]" italics="true" pageId="9" pageNumber="248">M. cuneifolia</emphasis>
subsp.
<emphasis id="B93D9C09F2420020595DFCFC9AA53B74" box="[1345,1445,774,795]" italics="true" pageId="9" pageNumber="248">subandina</emphasis>
</taxonomicName>
,
<taxonomicName id="4C493B98F24200205C94FCD09E113B50" box="[136,273,810,831]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="9" pageNumber="248" phylum="Tracheophyta" rank="species" species="bongarana">
<emphasis id="B93D9C09F24200205C94FCD09E113B50" box="[136,273,810,831]" italics="true" pageId="9" pageNumber="248">M. bongarana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C493B98F24200205D5CFCD09EFC3B50" authorityName="Wurdack" authorityYear="1957" box="[320,508,810,831]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="9" pageNumber="248" phylum="Tracheophyta" rank="species" species="dimorphanthera">
<emphasis id="B93D9C09F24200205D5CFCD09EFC3B50" box="[320,508,810,831]" italics="true" pageId="9" pageNumber="248">M. dimorphanthera</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BF6401BF24200205C90FC9F9A733FB5" pageId="9" pageNumber="248">
<table id="F949B2BBF242FFD65C90FCAC9AA73FB4" box="[140,1447,854,2011]" gridcols="5" gridrows="7" pageId="9" pageNumber="248">
<tr id="35794259F242FFD65C90FCAC9AA73BE9" box="[140,1447,854,902]" gridrow="0" pageId="9" pageNumber="248">
<th id="76A82B25F242FFD65C90FCAC9FFF3BE9" box="[140,255,854,902]" gridcol="0" gridrow="0" pageId="9" pageNumber="248">Species</th>
<th id="76A82B25F242FFD65D3CFCAC9EF83BE9" box="[288,504,854,902]" gridcol="1" gridrow="0" pageId="9" pageNumber="248">
<taxonomicName id="4C493B98F24200205D3CFC9F9E893B17" authorityName="M. M. Jimenez &amp; H. Garzon" authorityYear="2024" box="[288,393,869,888]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="9" pageNumber="248" phylum="Tracheophyta" rank="species" species="judithiae">
<emphasis id="B93D9C09F24200205D3CFC9F9E893B17" box="[288,393,869,888]" italics="true" pageId="9" pageNumber="248">M. judithiae</emphasis>
</taxonomicName>
</th>
<th id="76A82B25F242FFD65E19FCAC9CDC3BE9" box="[517,988,854,902]" gridcol="2" gridrow="0" pageId="9" pageNumber="248">
<taxonomicName id="4C493B98F24200205E19FCAD9D773B06" authorityName="Gleason" authorityYear="1947" box="[517,631,854,873]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="9" pageNumber="248" phylum="Tracheophyta" rank="species" species="cuneifolia">
<emphasis id="B93D9C09F24200205E19FCAD9D773B06" box="[517,631,854,873]" italics="true" pageId="9" pageNumber="248">M. cuneifolia</emphasis>
</taxonomicName>
subsp.
<taxonomicName id="4C493B98F24200205EEEFCAD9DE43BE9" authority="(Gleason 1947)" baseAuthorityName="Gleason" baseAuthorityYear="1947" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="9" pageNumber="248" phylum="Tracheophyta" rank="subSpecies" species="cuneifolia" subSpecies="cuneifolia">
<emphasis id="B93D9C09F24200205EEEFCAD9C633B06" box="[754,867,854,873]" italics="true" pageId="9" pageNumber="248">M. cuneifolia</emphasis>
subsp.
<emphasis id="B93D9C09F24200205E19FC899D5A3BE9" box="[517,602,883,902]" italics="true" pageId="9" pageNumber="248">cuneifolia</emphasis>
(
<bibRefCitation id="EFD83DEAF24200205E7AFC899DDE3BE9" author="Gleason, H. A." box="[614,734,883,902]" pageId="9" pageNumber="248" pagination="294 - 306" refId="ref5528" refString="Gleason, H. A. (1947) New and noteworthy melastomes, chiefly Ecuadorian. Phytologia 2: 294 - 306." type="journal article" year="1947">Gleason 1947</bibRefCitation>
)
</taxonomicName>
<emphasis id="B93D9C09F24200205EEEFC899C4B3BE9" box="[754,843,883,902]" italics="true" pageId="9" pageNumber="248">subandina</emphasis>
(
<bibRefCitation id="EFD83DEAF24200205F4AFC899CD63BE9" author="Wurdack, J. J." box="[854,982,883,902]" pageId="9" pageNumber="248" pagination="1 - 13" refId="ref6670" refString="Wurdack, J. J. (1976) Certamen Melastomataceis XXV. Phytologia 35: 1 - 13. https: // doi. org / 10.5962 / bhl. part. 2606" type="journal article" year="1976">Wurdack 1976</bibRefCitation>
)
</th>
<th id="76A82B25F242FFD65FF5FCAC9BBE3BE9" box="[1001,1214,854,902]" gridcol="3" gridrow="0" pageId="9" pageNumber="248">
<taxonomicName id="4C493B98F24200205FF5FCAD9BB13BE9" authority="(Fernandez et al. 2022)" baseAuthorityName="Fernandez" baseAuthorityYear="2022" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="9" pageNumber="248" phylum="Tracheophyta" rank="species" species="bongarana">
<emphasis id="B93D9C09F24200205FF5FCAD9B643B06" box="[1001,1124,854,873]" italics="true" pageId="9" pageNumber="248">M. bongarana</emphasis>
(Fernandez
<emphasis id="B93D9C09F24200205853FC8E9B7B3BE9" box="[1103,1147,883,902]" italics="true" pageId="9" pageNumber="248">et al.</emphasis>
2022)
</taxonomicName>
</th>
<th id="76A82B25F242FFD658D0FCAC9AA73BE9" box="[1228,1447,854,902]" gridcol="4" gridrow="0" pageId="9" pageNumber="248">
<taxonomicName id="4C493B98F242002058D0FCAD9A573BE9" authority="(Wurdack 1957) Leaves Anisophyllous" authorityName="Leaves Anisophyllous" baseAuthorityName="Wurdack" baseAuthorityYear="1957" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="9" pageNumber="248" phylum="Tracheophyta" rank="species" species="dimorphanthera">
<emphasis id="B93D9C09F242002058D0FCAD9A733B06" box="[1228,1395,854,873]" italics="true" pageId="9" pageNumber="248">M. dimorphanthera</emphasis>
(
<bibRefCitation id="EFD83DEAF242002058CDFC899A523BE9" author="Wurdack, J. J." box="[1233,1362,883,902]" pageId="9" pageNumber="248" pagination="101 - 109" refId="ref6598" refString="Wurdack, J. J. (1957) Certamen Melastomataceis - IV. Brittonia 9: 101 - 109. https: // doi. org / 10.2307 / 2804774" type="journal article" year="1957">Wurdack 1957</bibRefCitation>
)
</taxonomicName>
</th>
</tr>
<tr id="35794259F242FFD65C90FC579AA73CBB" box="[140,1447,941,1236]" gridrow="1" pageId="9" pageNumber="248">
<th id="76A82B25F242FFD65C90FC579FFF3CBB" box="[140,255,941,1236]" gridcol="0" gridrow="1" pageId="9" pageNumber="248">Leaves</th>
<td id="76A82B25F242FFD65D3CFC579EF83CBB" box="[288,504,941,1236]" gridcol="1" gridrow="1" pageId="9" pageNumber="248">Anisophyllous, petioles 0.3–1.0 cm long; blades 6.1–27.1 × 1.2–8.2 cm, elliptic, apex long- acuminate, base cuneate- decurrent; venation suprabasal, abaxial surface puberulous</td>
<td id="76A82B25F242FFD65E19FC579CDC3CBB" box="[517,988,941,1236]" gridcol="2" gridrow="1" pageId="9" pageNumber="248">Anisophyllous, petioles Anisophyllous, petioles 2.0–3.0 cm long; blades 0.7–0.9 cm long; blades 10–13 × 2.5–4.5 cm, 15–23 × 5–10 cm, elliptic, oblanceolate, apex apex acuminate, base abruptly and sharply attenuate-decurrent; acuminate, base long- venation suprabasal, cuneate; venation abaxial surface sparsely suprabasal, abaxial surface stellate-setulose softly cinereous</td>
<td id="76A82B25F242FFD65FF5FC579BBE3CBB" box="[1001,1214,941,1236]" gridcol="3" gridrow="1" pageId="9" pageNumber="248">Isophyllous; petioles 1.2–1.5 cm long; blades 7.5–9.5 × 3.2–4.3 cm, elliptic, apex acuminate, base acute; venation suprabasal, abaxial surface densely pubescent</td>
<td id="76A82B25F242FFD658D0FC579AA73CBB" box="[1228,1447,941,1236]" gridcol="4" gridrow="1" pageId="9" pageNumber="248">Anisophyllous, petioles 2.0–9.0 cm long; blades 7.0–24 × 3.2–14.0 cm, elliptic to oblong-elliptic; apex obtuse to shortly acuminate, base obtuse to slightly rounded; venation basal or slightly suprabasal, abaxial surface densely rufous</td>
</tr>
<tr id="35794259F242FFD65C90FAFE9AA73D3A" box="[140,1447,1284,1365]" gridrow="2" pageId="9" pageNumber="248">
<th id="76A82B25F242FFD65C90FAFE9FFF3D3A" box="[140,255,1284,1365]" gridcol="0" gridrow="2" pageId="9" pageNumber="248">Inflorescence</th>
<td id="76A82B25F242FFD65D3CFAFE9EF83D3A" box="[288,504,1284,1365]" gridcol="1" gridrow="2" pageId="9" pageNumber="248">Multiflorous, 4.3–6.1 cm long, 4–8 flowered umbels</td>
<td id="76A82B25F242FFD65E19FAFE9CDC3D3A" box="[517,988,1284,1365]" gridcol="2" gridrow="2" pageId="9" pageNumber="248">Submultiflorous, 6.0–7.9 Few-flowered, 1.7–2.7 cm cm long, 3–9 flowered long, 3–4 flowered umbels umbels</td>
<td id="76A82B25F242FFD65FF5FAFE9BBE3D3A" box="[1001,1214,1284,1365]" gridcol="3" gridrow="2" pageId="9" pageNumber="248">Submultiflorous, 7.5–8.2 cm long, 4–5 flowered umbels</td>
<td id="76A82B25F242FFD658D0FAFE9AA73D3A" box="[1228,1447,1284,1365]" gridcol="4" gridrow="2" pageId="9" pageNumber="248">Multiflorous, 5.0–15.0 cm long, with umbels with more than 3 flowers</td>
</tr>
<tr id="35794259F242FFD65C90FA7F9AA73E5D" box="[140,1447,1413,1586]" gridrow="3" pageId="9" pageNumber="248">
<th id="76A82B25F242FFD65C90FA7F9FFF3E5D" box="[140,255,1413,1586]" gridcol="0" gridrow="3" pageId="9" pageNumber="248">Hypanthium and calyx</th>
<td id="76A82B25F242FFD65D3CFA7F9EF83E5D" box="[288,504,1413,1586]" gridcol="1" gridrow="3" pageId="9" pageNumber="248">Cupuliform, 3.4–3.9 mm long, calyx lobes 0.5–1.3 mm long, subtruncate, thickened to the apex, not extending beyond the lobes</td>
<td id="76A82B25F242FFD65E19FA7F9CDC3E5D" box="[517,988,1413,1586]" gridcol="2" gridrow="3" pageId="9" pageNumber="248">Campanulate, 5.2 mm Obconic, 7.3 mm long, long, calyx lobes 8.5 mm calyx lobes 0.8–1.1 mm long, acute with claw- long, acute, subtruncate, shaped projections 6.0–7.5 claw-shaped projections mm long 2.3–3.0 mm long</td>
<td id="76A82B25F242FFD65FF5FA7F9BBE3E5D" box="[1001,1214,1413,1586]" gridcol="3" gridrow="3" pageId="9" pageNumber="248">Campanulate, 6.5–7.0 mm long, calyx lobes 2.0–2.5 × 5–7 mm, acute with claw-shaped projections 2.0–2.5 mm long</td>
<td id="76A82B25F242FFD658D0FA7F9AA73E5D" box="[1228,1447,1413,1586]" gridcol="4" gridrow="3" pageId="9" pageNumber="248">Cupuliform, 3.5–6.0 mm long, calyx lobes broadly triangular with dorsal teeth non-visible or exceeding the length of the lobe 0.9–2.5 mm long</td>
</tr>
<tr id="35794259F242FFD65C90F9989AA73EFA" box="[140,1447,1634,1685]" gridrow="4" pageId="9" pageNumber="248">
<th id="76A82B25F242FFD65C90F9989FFF3EFA" box="[140,255,1634,1685]" gridcol="0" gridrow="4" pageId="9" pageNumber="248">
<taxonomicName id="4C493B98F24200205C90F9889FCC3EEA" authority="Petals" authorityName="Petals" box="[140,204,1650,1669]" class="Gastropoda" family="Cymbuliidae" genus="Corolla" kingdom="Animalia" order="Pteropoda" pageId="9" pageNumber="248" phylum="Mollusca" rank="genus">Corolla</taxonomicName>
</th>
<td id="76A82B25F242FFD65D3CF9989CDC3EFA" box="[288,988,1634,1685]" colspan="2" colspanRight="1" gridcol="1" gridrow="4" pageId="9" pageNumber="248">Petals 11.1–11.3 mm long Petals 13 mm long Petals 13.1–13.4 mm long</td>
<td id="76A82B25F242FFD65FF5F9989BBE3EFA" box="[1001,1214,1634,1685]" gridcol="3" gridrow="4" pageId="9" pageNumber="248">Petals 14–15.5 × 13–16 mm long</td>
<td id="76A82B25F242FFD658D0F9989AA73EFA" box="[1228,1447,1634,1685]" gridcol="4" gridrow="4" pageId="9" pageNumber="248">Petals 12.0–16.0 mm long</td>
</tr>
<tr id="35794259F242FFD65C90F9329AA73FFB" box="[140,1447,1736,1940]" gridrow="5" pageId="9" pageNumber="248">
<th id="76A82B25F242FFD65C90F9329FFF3FFB" box="[140,255,1736,1940]" gridcol="0" gridrow="5" pageId="9" pageNumber="248">Stamens</th>
<td id="76A82B25F242FFD65D3CF9329EF83FFB" box="[288,504,1736,1940]" gridcol="1" gridrow="5" pageId="9" pageNumber="248">Isomorphic, filaments 4.4–5.8 mm long, connectives prolonged below the thecae, the dorso-basal appendage crown-shaped, thecae 5.4–5.8 mm long</td>
<td id="76A82B25F242FFD65E19F9329BBE3FFB" box="[517,1214,1736,1940]" colspan="2" colspanRight="1" gridcol="2" gridrow="5" pageId="9" pageNumber="248">Dimorphic, filaments 6.4– Isomorphic, filaments Dimorphic, filaments 8.0– 8.5 mm long, connectives 8.0–8.5 mm long, 9.5 mm long, connectives prolonged below the connectives prolonged prolonged below the thecae, thecae, the dorso-basal below the thecae, the the dorso-basal appendage appendage broadly conic, dorso-basal appendage crown-shaped, thecae obtuse or rounded, thecae slightly crown-shaped, 7.6–9.7 mm long 5.3–8.3 mm long thecae 8.5–9.0 mm long</td>
<td id="76A82B25F242FFD658D0F9329AA73FFB" box="[1228,1447,1736,1940]" gridcol="4" gridrow="5" pageId="9" pageNumber="248">Dimorphic, filaments 6.0–10.0 mm long, connectives prolonged below the thecae, the dorso-basal appendage dentiform, thecae 4.8–8.0 mm long</td>
</tr>
<tr id="35794259F242FFD65C90F83D9AA73FB4" box="[140,1447,1991,2011]" gridrow="6" pageId="9" pageNumber="248">
<th id="76A82B25F242FFD65C90F83D9FFF3FB4" box="[140,255,1991,2011]" gridcol="0" gridrow="6" pageId="9" pageNumber="248">Style</th>
<td id="76A82B25F242FFD65D3CF83D9EF83FB4" box="[288,504,1991,2011]" gridcol="1" gridrow="6" pageId="9" pageNumber="248">14.1–15.3 mm long</td>
<td id="76A82B25F242FFD65E19F83D9CDC3FB4" box="[517,988,1991,2011]" gridcol="2" gridrow="6" pageId="9" pageNumber="248">21 mm long 21.5–22.4 mm long</td>
<td id="76A82B25F242FFD65FF5F83D9BBE3FB4" box="[1001,1214,1991,2011]" gridcol="3" gridrow="6" pageId="9" pageNumber="248">25.5–27.0 mm long</td>
<td id="76A82B25F242FFD658D0F83D9AA73FB4" box="[1228,1447,1991,2011]" gridcol="4" gridrow="6" pageId="9" pageNumber="248">14.0–23.0 mm long</td>
</tr>
</table>
</paragraph>
<paragraph id="8BF6401BF24100235CA1FF649CC339DB" blockId="10.[136,1452,158,509]" pageId="10" pageNumber="249">
Finally,
<taxonomicName id="4C493B98F24100235D00FF659EF238D7" authorityName="M. M. Jimenez &amp; H. Garzon" authorityYear="2024" box="[284,498,158,184]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="10" pageNumber="249" phylum="Tracheophyta" rank="species" species="judithiae">
<emphasis id="B93D9C09F24100235D00FF659EF238D7" box="[284,498,158,184]" italics="true" pageId="10" pageNumber="249">Meriania judithiae</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="4C493B98F24100235EE4FF659CD738D7" authorityName="Leaves Anisophyllous" baseAuthorityName="Wurdack" baseAuthorityYear="1957" box="[760,983,158,184]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="10" pageNumber="249" phylum="Tracheophyta" rank="species" species="dimorphanthera">
<emphasis id="B93D9C09F24100235EE4FF659CD738D7" box="[760,983,158,184]" italics="true" pageId="10" pageNumber="249">M. dimorphanthera</emphasis>
</taxonomicName>
by the narrower, bullate leaf blades (
<emphasis id="B93D9C09F24100235991FF659AAB38D7" box="[1421,1451,159,184]" italics="true" pageId="10" pageNumber="249">vs.</emphasis>
smooth) with the base being attenuate (
<emphasis id="B93D9C09F24100235E5BFF399D6538B3" box="[583,613,195,220]" italics="true" pageId="10" pageNumber="249">vs.</emphasis>
obtuse to rounded), the shorter hypanthium
<quantity id="4CB1EDFEF24100235842FF389BE638B2" box="[1118,1254,194,221]" metricMagnitude="-3" metricUnit="m" metricValue="3.65" metricValueMax="3.9" metricValueMin="3.4" pageId="10" pageNumber="249" unit="mm" value="3.65" valueMax="3.9" valueMin="3.4">3.4–3.9 mm</quantity>
long (
<emphasis id="B93D9C09F2410023592DFF399A4838B3" box="[1329,1352,195,220]" italics="true" pageId="10" pageNumber="249">vs</emphasis>
. 3.5–6.0 mm long), the calyx lobes being subtruncate, not ruptured to the torus (
<emphasis id="B93D9C09F24100235FA0FF1D9CDA396F" box="[956,986,231,256]" italics="true" pageId="10" pageNumber="249">vs.</emphasis>
broadly triangular, subcalyptrate) with a conical projection in the apex (
<emphasis id="B93D9C09F24100235DFCFEF19EFF394B" box="[480,511,267,292]" italics="true" pageId="10" pageNumber="249">vs.</emphasis>
dorsal teeth exceeding or not the lobe), the isomorphic stamens (
<emphasis id="B93D9C09F241002358CEFEF19BF1394B" box="[1234,1265,267,292]" italics="true" pageId="10" pageNumber="249">vs.</emphasis>
dimorphic) with shorter filaments
<quantity id="4CB1EDFEF24100235D52FED49ED73926" box="[334,471,302,329]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" metricValueMax="5.8" metricValueMin="4.4" pageId="10" pageNumber="249" unit="mm" value="5.1" valueMax="5.8" valueMin="4.4">4.4–5.8 mm</quantity>
long (
<emphasis id="B93D9C09F24100235E3EFED59D403927" box="[546,576,303,328]" italics="true" pageId="10" pageNumber="249">vs.</emphasis>
6.0–
<quantity id="4CB1EDFEF24100235E67FED49DCA3926" box="[635,714,302,329]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="249" unit="mm" value="10.0">10 mm</quantity>
long), the dorsal appendage of the connective crown-shaped (
<emphasis id="B93D9C09F24100235991FED59AAB3927" box="[1421,1451,303,328]" italics="true" pageId="10" pageNumber="249">vs.</emphasis>
dentiform), the shorter thecae
<quantity id="4CB1EDFEF24100235DC2FEA89D663902" box="[478,614,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" metricValueMax="5.8" metricValueMin="5.4" pageId="10" pageNumber="249" unit="mm" value="5.6" valueMax="5.8" valueMin="5.4">5.4–5.8 mm</quantity>
long (
<emphasis id="B93D9C09F24100235EADFEA99DCF3903" box="[689,719,339,364]" italics="true" pageId="10" pageNumber="249">vs.</emphasis>
4.8–8.0 mm long), and the shorter style
<quantity id="4CB1EDFEF24100235882FEA89A423902" box="[1182,1346,338,365]" metricMagnitude="-2" metricUnit="m" metricValue="1.47" metricValueMax="1.53" metricValueMin="1.41" pageId="10" pageNumber="249" unit="mm" value="14.7" valueMax="15.3" valueMin="14.1">14.1–15.3 mm</quantity>
long (
<emphasis id="B93D9C09F24100235991FEA99AAB3903" box="[1421,1451,339,364]" italics="true" pageId="10" pageNumber="249">vs.</emphasis>
14.0–23.0 mm long) (
<bibRefCitation id="EFD83DEAF24100235D63FE8C9DA239FF" author="Mendoza-Cifuentes, H." box="[383,674,374,400]" pageId="10" pageNumber="249" refId="ref5860" refString="Mendoza-Cifuentes, H. (2021) Taxonomic review of the genus Meriania (Melastomataceae) in Colombia. Acta Botanica Mexicana 128: e 1734. https: // doi. org / 10.21829 / abm 128.2021.1734" type="journal volume" year="2021">Mendoza-Cifuentes 2021</bibRefCitation>
). A summary comparing these morphological characters between
<taxonomicName id="4C493B98F24100235991FE8D9FEC39DB" authorityName="M. M. Jimenez &amp; H. Garzon" authorityYear="2024" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="10" pageNumber="249" phylum="Tracheophyta" rank="species" species="judithiae">
<emphasis id="B93D9C09F24100235991FE8D9FEC39DB" italics="true" pageId="10" pageNumber="249">M. judithiae</emphasis>
</taxonomicName>
and the other species as mentioned above is provided in
<tableCitation id="C6CB75A0F24100235F75FE609CBD39DB" box="[873,957,410,436]" captionStart="TABLE 1" captionStartId="9.[136,216,774,795]" captionText="TABLE 1. Summary of principal differences between Meriania judithiae, M.cuneifolia subsp. cuneifolia, M. cuneifolia subsp. subandina, M. bongarana and M. dimorphanthera." pageId="10" pageNumber="249">Table 1</tableCitation>
.
</paragraph>
<paragraph id="8BF6401BF24100235CA1FE449A973992" blockId="10.[136,1452,158,509]" pageId="10" pageNumber="249">
<emphasis id="B93D9C09F24100235CA1FE449DB839B7" bold="true" box="[189,696,446,472]" pageId="10" pageNumber="249">
Additional specimen examined (
<typeStatus id="54F2FEB9F24100235E25FE449DAB39B7" box="[569,683,446,472]" pageId="10" pageNumber="249" type="paratype">paratype</typeStatus>
):
</emphasis>
—
<collectingCountry id="F35E008BF24100235EC8FE449C5B39B7" box="[724,859,446,472]" name="Ecuador" pageId="10" pageNumber="249">ECUADOR</collectingCountry>
.
<collectingRegion id="498D8EF9F24100235F75FE459B2E39B7" box="[873,1070,446,473]" country="Ecuador" name="Morona-Santiago" pageId="10" pageNumber="249">Morona Santiago</collectingRegion>
: San Juan Bosco. San Juan Bosco, Jungle Dave’s sendero al Cerro PDA,
<quantity id="4CB1EDFEF24100235E2FFE189D8A3992" box="[563,650,482,509]" metricMagnitude="3" metricUnit="m" metricValue="1.815" pageId="10" pageNumber="249" unit="m" value="1815.0">1815 m</quantity>
,
<date id="FFF766DBF24100235E8AFE189C2E3993" box="[662,814,482,508]" pageId="10" pageNumber="249" value="2023-07-22">22 Julio 2023</date>
(fl.),
<emphasis id="B93D9C09F24100235F72FE199BC63993" box="[878,1222,482,509]" italics="true" pageId="10" pageNumber="249">M. Jiménez &amp; H. Garzón 1810</emphasis>
(HUTPL 15146!).
</paragraph>
</subSubSection>
</treatment>
</document>

199
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@ -0,0 +1,199 @@
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<mods:title id="17CACA9E5B2B14E24A4C2153EDF8BDA6">Podoscypha ahuensis sp. nov. (Polyporales, Basidiomycota) evidenced by the morphological characters and phylogenetic analyses</mods:title>
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<paragraph id="8BE436E57172FFA263F6A30DFBC2FA38" blockId="4.[136,1057,1396,1423]" box="[136,1057,1396,1423]" pageId="4" pageNumber="266">
<heading id="D0AC81897172FFA263F6A30DFBC2FA38" box="[136,1057,1396,1423]" fontSize="11" level="2" pageId="4" pageNumber="266" reason="3">
<taxonomicName id="4C5B4D667172FFA263F6A30DFCDCFA38" authority="S. Shen, C.W. Zhang, and B. W. Zhang" authorityName="S. Shen, C. W. Zhang, and B. W. Zhang" authorityYear="2024" box="[136,831,1396,1423]" class="Agaricomycetes" family="Meruliaceae" genus="Podoscypha" kingdom="Fungi" order="Polyporales" pageId="4" pageNumber="266" phylum="Basidiomycota" rank="species" species="ahuensis" status="sp. nov.">
<emphasis id="B92FEAF77172FFA263F6A30DFE63FA39" bold="true" box="[136,384,1396,1423]" italics="true" pageId="4" pageNumber="266">Podoscypha ahuensis</emphasis>
S. Shen, C.W. Zhang, and B. W. Zhang
</taxonomicName>
,
<emphasis id="B92FEAF77172FFA26035A30CFC43FA38" box="[843,928,1397,1422]" italics="true" pageId="4" pageNumber="266">
<taxonomicNameLabel id="A21C578C7172FFA26035A30CFC43FA38" box="[843,928,1397,1422]" pageId="4" pageNumber="266" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13602A607172FFA260D0A30DFC1EFA38" box="[942,1021,1396,1422]" captionStart="FIGURE 3" captionStartId="5.[136,229,542,564]" captionText="FIGURE 3. Basidiomata of Podoscypha ahuensis (holotype, AHU 73). Scale bars: 1 cm." pageId="4" pageNumber="266">Figs. 3</figureCitation>
,
<figureCitation id="13602A607172FFA26774A30DFBFAFA38" box="[1034,1049,1396,1422]" captionStart="FIGURE 4" captionStartId="5.[136,229,1510,1532]" captionTargetBox="[574,1034,603,1482]" captionTargetId="figure-44@5.[553,1034,590,1486]" captionTargetPageId="5" captionText="FIGURE 4. Microscopic structures of Podoscypha ahuensis (drawn from the holotype AHU 73). a. Basidiospores. b. Basidia and basidioles. c. Gloeocystidia. d. Pilocystidia. e. Hyphae. f. Caulocystidia. Bars: a–f 10 μm." pageId="4" pageNumber="266">4</figureCitation>
)
</heading>
</paragraph>
<paragraph id="8BE436E57172FFA263F6A3E1FE27FA05" blockId="4.[136,452,1432,1459]" box="[136,452,1432,1459]" pageId="4" pageNumber="266">MycoBank no.: MB 853381</paragraph>
</subSubSection>
<subSubSection id="C341656E7172FFA363F6A399FE55F7AF" lastPageId="5" lastPageNumber="267" pageId="4" pageNumber="266" type="diagnosis">
<paragraph id="8BE436E57172FFA263F6A399FD12F9F4" blockId="4.[136,1452,1504,1998]" pageId="4" pageNumber="266">
<emphasis id="B92FEAF77172FFA263F6A399FDE0FA4C" box="[136,515,1504,1530]" italics="true" pageId="4" pageNumber="266">
Diagnosis—
<taxonomicName id="4C5B4D667172FFA2626AA399FDE0FA4C" authorityName="S. Shen, C. W. Zhang, and B. W. Zhang" authorityYear="2024" box="[276,515,1504,1530]" class="Agaricomycetes" family="Meruliaceae" genus="Podoscypha" kingdom="Fungi" order="Polyporales" pageId="4" pageNumber="266" phylum="Basidiomycota" rank="species" species="ahuensis">Podoscypha ahuensis</taxonomicName>
</emphasis>
is characterized by its smooth, salmon pink, flesh pink to pink pileal surface, distinct swollen gloeocystidia, presence of pilocystidia and caulocystidia, small, thin-walled, ellipsoid to broadly ellipsoid basidiospores, on angiosperms from subtropical
<collectingCountry id="F34C76757172FFA261D9A051FD0DF9F4" box="[679,750,1576,1602]" name="China" pageId="4" pageNumber="266">China</collectingCountry>
.
</paragraph>
<paragraph id="8BE436E57172FFA263C3A034FC60F93C" blockId="4.[136,1452,1504,1998]" pageId="4" pageNumber="266">
<emphasis id="B92FEAF77172FFA263C3A034FF12F9D0" box="[189,241,1613,1638]" italics="true" pageId="4" pageNumber="266">Type</emphasis>
.—
<emphasis id="B92FEAF77172FFA2626BA035FEBDF9D0" bold="true" box="[277,350,1612,1638]" pageId="4" pageNumber="266">
<collectingCountry id="F34C76757172FFA2626BA035FEBDF9D0" box="[277,350,1612,1638]" name="China" pageId="4" pageNumber="266">China</collectingCountry>
</emphasis>
.
<collectingRegion id="499FF8077172FFA26217A035FDF9F9D0" box="[361,538,1612,1638]" country="China" name="Anhui" pageId="4" pageNumber="266">Anhui Province</collectingRegion>
, Hefei, Shushan District,
<collectingRegion id="499FF8077172FFA2604DA035FC9AF9D0" box="[819,889,1612,1638]" country="China" name="Anhui" pageId="4" pageNumber="266">Anhui</collectingRegion>
University, Qingyuan Campus, on the dead branch of angiosperm,
<date id="FFE510257172FFA26249A009FDF7F93C" box="[311,532,1648,1674]" pageId="4" pageNumber="266" value="2023-09-22">22 September 2023</date>
, AHU 337 (
<typeStatus id="54E088477172FFA261E4A009FCE3F93C" box="[666,768,1648,1674]" pageId="4" pageNumber="266" type="holotype">holotype</typeStatus>
, AHUBM).
</paragraph>
<paragraph id="8BE436E57172FFA263C3A0EDFDE3F940" blockId="4.[136,1452,1504,1998]" pageId="4" pageNumber="266">
<emphasis id="B92FEAF77172FFA263C3A0EDFED6F918" box="[189,309,1684,1710]" italics="true" pageId="4" pageNumber="266">Etymology</emphasis>
.—
<taxonomicName id="4C5B4D667172FFA26226A0EDFDE7F918" authority="(Lat.)" baseAuthorityName="Lat." box="[344,516,1684,1710]" class="Agaricomycetes" family="Meruliaceae" genus="Podoscypha" kingdom="Fungi" order="Polyporales" pageId="4" pageNumber="266" phylum="Basidiomycota" rank="species" species="ahuensis">
<emphasis id="B92FEAF77172FFA26226A0EDFE59F918" box="[344,442,1684,1710]" italics="true" pageId="4" pageNumber="266">ahuensis</emphasis>
(Lat.)
</taxonomicName>
: The new species is named after AHU, the abbreviation for
<collectingRegion id="499FF8077172FFA267E7A0EDFB3CF918" box="[1177,1247,1684,1710]" country="China" name="Anhui" pageId="4" pageNumber="266">Anhui</collectingRegion>
University, where the samples of this species were discovered and collected.And ‘ahuensis’ is an adjective, referring to the locality
<collectingRegion id="499FF8077172FFA2661BA0C1FA48F964" box="[1381,1451,1720,1746]" country="China" name="Anhui" pageId="4" pageNumber="266">Anhui</collectingRegion>
University of the
<typeStatus id="54E088477172FFA26233A0A4FE9EF941" box="[333,381,1757,1783]" pageId="4" pageNumber="266">type</typeStatus>
specimens.
</paragraph>
<paragraph id="8BE436E57172FFA263C3A179FBA0F878" blockId="4.[136,1452,1504,1998]" pageId="4" pageNumber="266">
<emphasis id="B92FEAF77172FFA263C3A179FEB0F8AC" box="[189,339,1792,1818]" italics="true" pageId="4" pageNumber="266">Basidiomata.</emphasis>
—Basidiocarps annual, gregarious, without odor or taste and corky when fresh, becoming hard corky upon drying, Pilei rod-shaped, spathulate or fan shaped when young, then the two sides of the fan are gradually joining together and it became a complete cupuliform, or a notched cupuliform when mature, stipe up to
<quantity id="4CA39B007172FFA26667A131FA8FF8D5" box="[1305,1388,1864,1891]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="4" pageNumber="266" unit="cm" value="2.0" valueMax="3.0" valueMin="1.0">1–3 cm</quantity>
high,
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in diameter, 250–450 µm thick. Basidiocarps surface smooth, salmon (6A4) to flesh pink (8A3/9A4) when fresh, buff yellowish (4A4) upon drying. Pileal surface glabrous, concentrically and radially zonate obviously on the dry specimen. Sterile margin distinct, thinner than middle, color is lighter and whiter.
</paragraph>
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<paragraph id="8BE436E57173FFA363F6A467FC01FD82" blockId="5.[136,994,542,565]" box="[136,994,542,565]" pageId="5" pageNumber="267">
<emphasis id="B92FEAF77173FFA363F6A467FF14FD82" bold="true" box="[136,247,542,564]" pageId="5" pageNumber="267">FIGURE 3</emphasis>
. Basidiomata of
<taxonomicName id="4C5B4D667173FFA362E5A466FD88FD82" box="[411,619,543,564]" class="Agaricomycetes" family="Meruliaceae" genus="Podoscypha" kingdom="Fungi" order="Polyporales" pageId="5" pageNumber="267" phylum="Basidiomycota" rank="species" species="ahuensis">
<emphasis id="B92FEAF77173FFA362E5A466FD88FD82" box="[411,619,543,564]" italics="true" pageId="5" pageNumber="267">Podoscypha ahuensis</emphasis>
</taxonomicName>
(holotype, AHU 73). Scale bars: 1 cm.
</paragraph>
</caption>
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<paragraph id="8BE436E57173FFA363F6A39FFC07F996" blockId="5.[136,1452,1510,1569]" pageId="5" pageNumber="267">
<emphasis id="B92FEAF77173FFA363F6A39FFF18FA4A" bold="true" box="[136,251,1510,1532]" pageId="5" pageNumber="267">FIGURE 4</emphasis>
. Microscopic structures of
<taxonomicName id="4C5B4D667173FFA3616DA39EFD05FA4A" box="[531,742,1511,1532]" class="Agaricomycetes" family="Meruliaceae" genus="Podoscypha" kingdom="Fungi" order="Polyporales" pageId="5" pageNumber="267" phylum="Basidiomycota" rank="species" species="ahuensis">
<emphasis id="B92FEAF77173FFA3616DA39EFD05FA4A" box="[531,742,1511,1532]" italics="true" pageId="5" pageNumber="267">Podoscypha ahuensis</emphasis>
</taxonomicName>
(drawn from the holotype AHU 73). a. Basidiospores. b. Basidia and basidioles. c. Gloeocystidia. d. Pilocystidia. e. Hyphae. f. Caulocystidia. Bars: a–f 10 μm.
</paragraph>
</caption>
<paragraph id="8BE436E57173FFA363C3A036FB99F907" blockId="5.[136,1452,1615,2073]" pageId="5" pageNumber="267">
<emphasis id="B92FEAF77173FFA363C3A036FE43F9DF" box="[189,416,1615,1641]" italics="true" pageId="5" pageNumber="267">Hyphal structure.—</emphasis>
Hyphal system dimitic; generative hyphae with clamps, 2–4.5 μm wide, hyaline, thin-to slightly thick-walled, rarely branched; skeletal hyphae colorless, 2.5–6 μm wide, thick-walled, unbranched. Hyphae in the stipe trama similar to that in the pileal context. IKI–, CB–; tissues unchanged in KOH.
</paragraph>
<paragraph id="8BE436E57173FFA363C3A0C5FF3AF81B" blockId="5.[136,1452,1615,2073]" pageId="5" pageNumber="267">
<taxonomicName id="4C5B4D667173FFA363C3A0C5FED5F963" box="[189,310,1724,1749]" genus="Hymenium" higherTaxonomySource="GBIF" kingdom="Fungi" pageId="5" pageNumber="267" rank="genus">
<emphasis id="B92FEAF77173FFA363C3A0C5FED5F963" box="[189,310,1724,1749]" italics="true" pageId="5" pageNumber="267">Hymenium</emphasis>
</taxonomicName>
<emphasis id="B92FEAF77173FFA36249A0C2FEB0F963" bold="true" box="[311,339,1723,1749]" pageId="5" pageNumber="267">—</emphasis>
Basidia clavate to cylindrical, with 4 sterigmata and a basal clamp, 23–35 × 4–6 µm, basidioles dominant, in shape similar to basidia, but slightly smaller. Gloeocystidia common, thin walled, cylindrical to subcylindrical, usually swollen distinctly in the middle and with obtuse apex, 25–35 × 6–10 µm; Pilocystidia hyaline, clavate to cylindrical, swollen near the base, apical narrowing, 25–55 × 5–11 µm; Caulocystidia almost cylindrical, more or less constricted, 40–100 × 6–10 µm. Basidiospores ellipsoid to broadly ellipsoid, hyaline, thin-walled, smooth, IKI–, CB–, (4–)4.2–6.5(–7) × 3–4.5(–5) µm, L = 4.98 µm, W = 3.65 µm, Q = 1.63–2.22, Qm = 1.36 ± 0.15 (n = 120/4).
</paragraph>
<paragraph id="8BE436E57173FFA363C3A1CEFE55F7AF" blockId="5.[136,1452,1615,2073]" pageId="5" pageNumber="267">
<emphasis id="B92FEAF77173FFA363C3A1CEFDA2F867" box="[189,577,1975,2001]" italics="true" pageId="5" pageNumber="267">Additional specimens examined.—</emphasis>
<materialsCitation id="3B333CB87173FFA3613FA1CEFCD4F843" collectingDate="2023-07-22" collectionCode="AHU" country="China" county="Hefei" location="University" municipality="Shushan District" pageId="5" pageNumber="267" specimenCode="AHU 73" specimenCount="1" stateProvince="Anhui">
<emphasis id="B92FEAF77173FFA3613FA1CEFD69F867" bold="true" box="[577,650,1975,2001]" pageId="5" pageNumber="267">
<collectingCountry id="F34C76757173FFA3613FA1CEFD69F867" box="[577,650,1975,2001]" name="China" pageId="5" pageNumber="267">China</collectingCountry>
</emphasis>
.
<collectingRegion id="499FF8077173FFA361E6A1CEFCA8F867" box="[664,843,1975,2001]" country="China" name="Anhui" pageId="5" pageNumber="267">Anhui Province</collectingRegion>
,
<collectingCounty id="62854E697173FFA36026A1CEFC7AF867" box="[856,921,1975,2001]" pageId="5" pageNumber="267">Hefei</collectingCounty>
,
<collectingMunicipality id="6B80AC9F7173FFA360DAA1CEFB81F867" box="[932,1122,1975,2001]" pageId="5" pageNumber="267">Shushan District</collectingMunicipality>
,
<collectingRegion id="499FF8077173FFA36712A1CEFB51F867" box="[1132,1202,1975,2001]" country="China" name="Anhui" pageId="5" pageNumber="267">Anhui</collectingRegion>
<location id="8E84603E7173FFA367C4A1CEFAD2F867" LSID="urn:lsid:plazi:treatment:03F287F37172FFA363F6A30DFE55F7AF:8E84603E7173FFA367C4A1CEFAD2F867" box="[1210,1329,1975,2001]" country="China" county="Hefei" municipality="Shushan District" name="University" pageId="5" pageNumber="267" stateProvince="Anhui">University</location>
,
<location id="8E84603E7173FFA36643A1CEFF04F843" LSID="urn:lsid:plazi:treatment:03F287F37172FFA363F6A30DFE55F7AF:8E84603E7173FFA36643A1CEFF04F843" country="China" county="Hefei" municipality="Shushan District" name="Qingyuan Campus" pageId="5" pageNumber="267" stateProvince="Anhui">Qingyuan Campus</location>
, on the stump of angiosperm,
<date id="FFE510257173FFA36146A1A2FD28F843" box="[568,715,2011,2037]" pageId="5" pageNumber="267" value="2023-07-22">
<collectingDate id="EFA1E9CD7173FFA36146A1A2FD28F843" box="[568,715,2011,2037]" pageId="5" pageNumber="267" value="2023-07-22">22 July 2023</collectingDate>
</date>
,
<specimenCode id="DBFD9E9E7173FFA361ABA1A2FCD4F843" box="[725,823,2011,2037]" collectionCode="AHU" country="Japan" httpUri="http://grbio.org/cool/yffg-d8ah" name="AHU Culture Collection" pageId="5" pageNumber="267">AHU 73</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B333CB87173FFA3603DA1A5FE51F7AF" collectingDate="2023-09-22" collectionCode="AHUBM" country="China" pageId="5" pageNumber="267" specimenCode="AHU 338, AHU 339" specimenCount="1">
on the stump of angiosperm,
<date id="FFE510257173FFA367F7A1A2FA86F843" box="[1161,1381,2011,2037]" pageId="5" pageNumber="267" value="2023-09-22">
<collectingDate id="EFA1E9CD7173FFA367F7A1A2FA86F843" box="[1161,1381,2011,2037]" pageId="5" pageNumber="267" value="2023-09-22">22 September 2023</collectingDate>
</date>
,
<specimenCode id="DBFD9E9E7173FFA36611A1A2FF56F7AF" collectionCode="AHU" country="Japan" httpUri="http://grbio.org/cool/yffg-d8ah" name="AHU Culture Collection" pageId="5" pageNumber="267">AHU 338</specimenCode>
,
<specimenCode id="DBFD9E9E7173FFA363C1A186FECFF7AF" box="[191,300,2047,2073]" collectionCode="AHU" country="Japan" httpUri="http://grbio.org/cool/yffg-d8ah" name="AHU Culture Collection" pageId="5" pageNumber="267">AHU 339</specimenCode>
(
<collectionCode id="ED4AAE207173FFA36245A186FE49F7AF" box="[315,426,2047,2073]" pageId="5" pageNumber="267">AHUBM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

447
data/41/38/E5/4138E526FFFCFFEF19EA4E73FB215043.xml Normal file
View file

@ -0,0 +1,447 @@
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<paragraph id="C92E5430FFFCFFEB19EA4E73FC0F5C4D" blockId="1.[136,969,1043,1071]" box="[136,969,1043,1071]" pageId="1" pageNumber="70">
<heading id="9266E35CFFFCFFEB19EA4E73FC0F5C4D" bold="true" box="[136,969,1043,1071]" fontSize="11" level="1" pageId="1" pageNumber="70" reason="1">
<emphasis id="FBE58822FFFCFFEB19EA4E73FC0F5C4D" bold="true" box="[136,969,1043,1071]" pageId="1" pageNumber="70">
<taxonomicName id="0E912FB3FFFCFFEB19EA4E73FCCE5C4D" authority="Nadruz, Magno &amp; Theofilo" authorityName="Nadruz, Magno &amp; Theofilo" authorityYear="2024" box="[136,776,1043,1071]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="1" pageNumber="70" phylum="Tracheophyta" rank="species" species="argyrophyllum" status="sp. nov.">
<emphasis id="FBE58822FFFCFFEB19EA4E73FE7D5C4C" bold="true" box="[136,443,1044,1071]" italics="true" pageId="1" pageNumber="70">Anthurium argyrophyllum</emphasis>
Nadruz, Magno &amp; Theófilo
</taxonomicName>
,
<emphasis id="FBE58822FFFCFFEB1A774E73FCAD5C4D" bold="true" box="[789,875,1044,1070]" italics="true" pageId="1" pageNumber="70">
<taxonomicNameLabel id="E0D63559FFFCFFEB1A774E73FCAD5C4D" box="[789,875,1044,1070]" pageId="1" pageNumber="70" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="51AA48B5FFFCFFEB1A194E73FC795C4D" box="[891,959,1044,1070]" captionStart="FIGURE 1" captionStartId="2.[136,229,1567,1589]" captionTargetBox="[313,1287,663,1529]" captionTargetId="figure-214@2.[299,1287,641,1543]" captionTargetPageId="2" captionText="FIGURE 1. Anthurium argyrophyllum. A. Habit. B–C. Mid petiole in cross section. D. Proximal midrib in cross section. E. Mid midrib in cross section. F. Distal midrib in cross section. G–H. Mid peduncle in cross section. I. Inflorescence." figureDoi="http://doi.org/10.5281/zenodo.14515965" httpUri="https://zenodo.org/record/14515965/files/figure.png" pageId="1" pageNumber="70">Fig. 1</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="818B07BBFFFCFFEB19EA4E07FD495D4A" pageId="1" pageNumber="70" type="reference_group">
<paragraph id="C92E5430FFFCFFEB19EA4E07FD495D4A" blockId="1.[136,1452,1120,1393]" pageId="1" pageNumber="70">
<emphasis id="FBE58822FFFCFFEB19EA4E07FD495D4A" italics="true" pageId="1" pageNumber="70">
<taxonomicName id="0E912FB3FFFCFFEB19EA4E07FE455C16" authorityName="Nadruz, Magno &amp; Theofilo" authorityYear="2024" box="[136,387,1120,1141]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="1" pageNumber="70" phylum="Tracheophyta" rank="species" species="argyrophyllum">Anthurium argyrophyllum</taxonomicName>
is very similar to
<taxonomicName id="0E912FB3FFFCFFEB1B544E07FBEF5C16" ID-CoL="67HN9" authority="Valadares &amp; Sakuragui (2015: 81)" authorityName="Valadares &amp; Sakuragui" authorityPageNumber="81" authorityYear="2015" box="[566,1065,1120,1141]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="1" pageNumber="70" phylum="Tracheophyta" rank="species" species="cachoeirense">
A. cachoeirense
<bibRefCitation id="AD0029C1FFFCFFEB1BB54E07FBEF5C16" author="Valadares, R. T. &amp; Sakuragui, C. M." box="[727,1065,1120,1141]" pageId="1" pageNumber="70" pagination="80 - 86" refId="ref4768" refString="Valadares, R. T. &amp; Sakuragui, C. M. (2015) A new species of Anthurium Schott (Araceae) in a seasonal semideciduous forest in Espirito Santo State, Brazil. Phytotaxa 217: 80 - 86. https: // doi. org / 10.11646 / phytotaxa. 217.1.7" type="journal article" year="2015">Valadares &amp; Sakuragui (2015: 81)</bibRefCitation>
</taxonomicName>
but differs in having a leaf blade ovate with an adaxial surface shiny moss green coloured and abaxial surface pale semi-glossy olive-green (vs. leaf blade elliptic to lanceolate with adaxial surface shiny dark green and abaxial surface light green coloured), primary lateral veins 11 to 16 on both sides (vs. 28–31 on both sides), peduncle with
<quantity id="0E69F9D5FFFCFFEB1B194EABFD375C82" box="[635,753,1228,1249]" metricMagnitude="-1" metricUnit="m" metricValue="1.055" metricValueMax="1.45" metricValueMin="0.6599999999999999" pageId="1" pageNumber="70" unit="cm" value="10.55" valueMax="14.5" valueMin="6.6">6.6–14.5 cm</quantity>
long, curved in fruiting (vs. peduncle with
<quantity id="0E69F9D5FFFCFFEB1DEC4EABFAD65C82" box="[1166,1296,1228,1249]" metricMagnitude="-1" metricUnit="m" metricValue="2.71" metricValueMax="3.11" metricValueMin="2.31" pageId="1" pageNumber="70" unit="cm" value="27.1" valueMax="31.1" valueMin="23.1">23.1–31.1 cm</quantity>
long, spreading in fruiting), spathe with
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long, chartaceous (vs. spathe with
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long, membranaceous) and stigma emergent and purplish (vs. stigma not emergent and greenish).
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="818B07BBFFFCFFEF19EA4F5FFB215043" lastPageId="5" lastPageNumber="74" pageId="1" pageNumber="70" type="materials_examined">
<paragraph id="C92E5430FFFCFFEB19EA4F5FFF325D12" blockId="1.[136,1452,1120,1393]" pageId="1" pageNumber="70">
<emphasis id="FBE58822FFFCFFEB19EA4F5FFF045D2E" bold="true" box="[136,194,1336,1357]" pageId="1" pageNumber="70">Type:</emphasis>
—
<collectingCountry id="B18614A0FFFCFFEB19B94F5FFEF25D2E" box="[219,308,1336,1357]" name="Brazil" pageId="1" pageNumber="70">BRAZIL</collectingCountry>
.
<collectingRegion id="0B559AD2FFFCFFEB185C4F5FFEBC5D2D" box="[318,378,1336,1358]" country="Brazil" name="Bahia" pageId="1" pageNumber="70">Bahia</collectingRegion>
: Santa Luzia, próximo a fazenda do Sr. Eugenio,
<date id="BD2F72F0FFFCFFEB1A024F5FFC795D2E" box="[864,959,1336,1357]" pageId="1" pageNumber="70" value="2019-01-29">
29.
<collectionCode id="AF80CCF5FFFCFFEB1A1F4F5FFC435D2E" box="[893,901,1336,1357]" country="Romania" lsid="urn:lsid:biocol.org:col:14415" name="&amp;quot;Alexandru Ioan Cuza&amp;quot; University" pageId="1" pageNumber="70" type="Herbarium">I</collectionCode>
.2019
</date>
, fl.,
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<collectionCode id="AF80CCF5FFFCFFEB1A8E4F5FFBC55D2E" box="[1004,1027,1336,1357]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="1" pageNumber="70" type="Herbarium">M</collectionCode>
.
<collectionCode id="AF80CCF5FFFCFFEB1D644F5FFBD45D2E" box="[1030,1042,1336,1357]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="70">T</collectionCode>
.
<collectionCode id="AF80CCF5FFFCFFEB1D774F5FFBEC5D2E" box="[1045,1066,1336,1357]" country="Denmark" name="University of Copenhagen" pageId="1" pageNumber="70" type="Herbarium">C</collectionCode>
. Lacerda &amp;
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.
<collectionCode id="AF80CCF5FFFCFFEB1DD44F5FFB0E5D2E" box="[1206,1224,1336,1357]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="1" pageNumber="70" type="Herbarium">E</collectionCode>
. Santos 1165
</emphasis>
(
<typeStatus id="162AEA92FFFCFFEB1C344F5FFA6A5D2D" box="[1366,1452,1336,1358]" pageId="1" pageNumber="70" type="holotype">holotype</typeStatus>
<collectionCode id="AF80CCF5FFFCFFEB19DF4F3BFF225D12" box="[189,228,1372,1393]" country="Brazil" lsid="urn:lsid:biocol.org:col:15528" name="Jardim Botânico do Rio de Janeiro" pageId="1" pageNumber="70" type="Herbarium">RB</collectionCode>
!).
</paragraph>
<paragraph id="C92E5430FFFCFFE819EA4FC7FE5D5A0B" blockId="1.[136,1452,1440,2079]" lastBlockId="2.[136,1452,158,617]" lastPageId="2" lastPageNumber="71" pageId="1" pageNumber="70">
Rupicolous herb, stem up to
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long, erect internodes very short; prophylls and cataphylls 3.0–4.0 cm long, greenish when young, chestnut to straw colored when old, forming a fibrous mass. Leaves not peltate, sheath
<quantity id="0E69F9D5FFFCFFEB1C354FA3FF6B5E60" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.1" metricValueMin="2.9" pageId="1" pageNumber="70" unit="cm" value="3.0" valueMax="3.1" valueMin="2.9">2.9–3.1 cm</quantity>
, not ligulate; petiole erect, 7.3–23.5 ×
<quantity id="0E69F9D5FFFCFFEB1B004F8FFD385E60" box="[610,766,1512,1539]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" metricValueMax="9.9" metricValueMin="5.5" pageId="1" pageNumber="70" unit="cm" value="0.77" valueMax="0.99" valueMin="0.55">0.55–0.99 cm</quantity>
, petiole:peduncle ratio 2.5:1.0, greenish, U-shaped, flattened to canaliculate adaxially with acute to obtuse margins, rounded abaxially, covered with raphides, without glandular punctations; geniculum 0.9–2.8 ×
<quantity id="0E69F9D5FFFCFFEB1B7A4C57FD5B5E28" box="[536,669,1584,1611]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="1" pageNumber="70" unit="cm" value="0.8" valueMax="0.9" valueMin="0.7">0.7–0.9 cm</quantity>
, clear matte green when fresh, drying blacker than rest of petiole, flattened to slightly concave with obtuse margins adaxially, rounded abaxially, without glandular punctations; leaf blade erect to spreading, ovate, 41.0–41.8 ×
<quantity id="0E69F9D5FFFCFFEB1B5D4C1FFD1A5EF0" box="[575,732,1656,1683]" metricMagnitude="-1" metricUnit="m" metricValue="1.62" metricValueMax="1.66" metricValueMin="1.58" pageId="1" pageNumber="70" unit="cm" value="16.2" valueMax="16.6" valueMin="15.8">15.8–16.6 cm</quantity>
, chartaceous when fresh and dried, apex acute-acuminate, base obtuse-truncate to obtuse-cordate, adaxial surface shiny moss green colour, with flat greenish margin, without covered with raphides, abaxial surface pale semi-glossy olive-green, drying green-greynish in both faces, without glandular punctations, posterior lobes up to
<quantity id="0E69F9D5FFFCFFEB1B774C83FDA25E9C" box="[533,612,1764,1791]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="70" unit="cm" value="1.5">1.5 cm</quantity>
long, not imbricate, rounded to obtuse at apex, sinus triangular without denudation; midrib greenish and lighter than blade adaxially, flattened at base, rounded to obtuse and prominent at apex adaxially, rounded and prominent abaxially; basal veins absent to 1, reaching the margin up to 1/8 of the length of the leaf blade; primary lateral veins 11 to 16 on both sides, obscured adaxially when fresh, slightly evident and darker than the lamina abaxially when fresh, arched, forming an angle of 30–43° at the junction with the midrib, interprimary veins distinct from primary lateral veins, collective vein arising at the base of the blade or up to 5.0–
<quantity id="0E69F9D5FFFCFFEB1C704DFFFA995FD0" box="[1298,1375,1944,1971]" metricMagnitude="-2" metricUnit="m" metricValue="6.8" pageId="1" pageNumber="70" unit="cm" value="6.8">6.8 cm</quantity>
above it,
<quantity id="0E69F9D5FFFCFFEB19CA4DDBFEE15FB4" box="[168,295,1980,2007]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.5" metricValueMin="0.8" pageId="1" pageNumber="70" unit="cm" value="1.15" valueMax="1.5" valueMin="0.8">0.8–1.5 cm</quantity>
from the margin. Inflorescence with peduncle greenish, erect up to anthesis, curved in fructification, terete to 1–2 ribbed, 6.6–14.5 ×
<quantity id="0E69F9D5FFFCFFEB18894D87FD405F98" box="[491,646,2016,2043]" metricMagnitude="-3" metricUnit="m" metricValue="6.65" metricValueMax="9.1" metricValueMin="4.2" pageId="1" pageNumber="70" unit="cm" value="0.665" valueMax="0.91" valueMin="0.42">0.42–0.91 cm</quantity>
, covered with raphides, glandular punctations absent; geniculum absent; spathe flat, ovate, 8.6–17.9 ×
<quantity id="0E69F9D5FFFCFFEB18874263FD95507C" box="[485,595,2052,2079]" metricMagnitude="-2" metricUnit="m" metricValue="4.55" metricValueMax="6.0" metricValueMin="3.1" pageId="1" pageNumber="70" unit="cm" value="4.55" valueMax="6.0" valueMin="3.1">3.1–6 cm</quantity>
, chartaceous, greenish to greenish with reddish to vinaceous nuances, not decurrent, acute at apex, erect in pre-anthesis, perpendicular to spreading in anthesis, erect in fructification, pustules absent, both surfaces densely covered with white speckles, forming a obtuse to acute angle with peduncle, decurrent
<quantity id="0E69F9D5FFFFFFE819EA4A81FEC05962" box="[136,262,230,257]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.6" metricValueMin="0.6" pageId="2" pageNumber="71" unit="cm" value="1.1" valueMax="1.6" valueMin="0.6">0.6–1.6 cm</quantity>
long; spadix stipitate, tapered, 3.8–13.7 ×
<quantity id="0E69F9D5FFFFFFE81B8A4A81FC425962" box="[744,900,230,257]" metricMagnitude="-2" metricUnit="m" metricValue="1.085" metricValueMax="1.41" metricValueMin="0.76" pageId="2" pageNumber="71" unit="cm" value="1.085" valueMax="1.41" valueMin="0.76">0.76–1.41 cm</quantity>
, light green color pre-anthesis, greenish-purplish in anthesis, gray-brown in post-anthesis, stipite
<quantity id="0E69F9D5FFFFFFE81BC64B6DFCE45946" box="[676,802,266,293]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.7" metricValueMin="0.3" pageId="2" pageNumber="71" unit="cm" value="1.0" valueMax="1.7" valueMin="0.3">0.3–1.7 cm</quantity>
long; 7 flowers visible per principal spiral, 11–12 visible per secondary spiral. Flowers rhombic, tepals greenish in pre-anthesis and anthesis, brownish post-anthesis, dorsally acute, internally convex; lateral tepals 1.74–1.85 ×
<quantity id="0E69F9D5FFFFFFE81BB84B35FC45590E" box="[730,899,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="1.855" metricValueMax="1.92" metricValueMin="1.79" pageId="2" pageNumber="71" unit="mm" value="1.855" valueMax="1.92" valueMin="1.79">1.79–1.92 mm</quantity>
; anterior/posterior tepals 1.61–1.95 ×
<quantity id="0E69F9D5FFFFFFE81C594B35FF7159F2" metricMagnitude="-3" metricUnit="m" metricValue="1.13" metricValueMax="1.14" metricValueMin="1.12" pageId="2" pageNumber="71" unit="mm" value="1.13" valueMax="1.14" valueMin="1.12">1.12–1.14 mm</quantity>
; stamens emerging in a scattered manner, laterals first followed quickly by alternates, filaments flattened, striated, 2.5–2.8 × 1.0–
<quantity id="0E69F9D5FFFFFFE818574BFDFE5659D6" box="[309,400,410,437]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="71" unit="mm" value="1.1">1.1 mm</quantity>
; anthers ovate, dorsifixed, extrorse, 0.6–0.8 ×
<quantity id="0E69F9D5FFFFFFE81AD64BFDFBD759D6" box="[948,1041,410,437]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="71" unit="mm" value="0.9">0.9 mm</quantity>
; polen yellowish to whitish; pistils purplish, oblong to elliptic, not emergent; stigma emergent, rounded, purplish; ovary bilocular, 3.2–3.5 ×
<quantity id="0E69F9D5FFFFFFE81C354BD9FF70599E" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.8" metricValueMin="1.3" pageId="2" pageNumber="71" unit="mm" value="2.05" valueMax="2.8" valueMin="1.3">1.3–2.8 mm</quantity>
, 1 ovule per locule, with apical placentation, funicle without trichomes. Infructescence with berries greenish to greenish-purplish at apex, translucent to whitish at base, oblong, 0.6–0.8 ×
<quantity id="0E69F9D5FFFFFFE81A884861FB545A42" box="[1002,1170,518,545]" metricMagnitude="-4" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="2" pageNumber="71" unit="mm" value="0.525" valueMax="0.55" valueMin="0.5">0.50–0.55 mm</quantity>
; mesocarp mucilaginous with elongate raphides; seeds cream color to yellowish, oblong in outline with convex faces, 0.35–0.40 ×
<quantity id="0E69F9D5FFFFFFE81C59484DFF705A0A" metricMagnitude="-4" metricUnit="m" metricValue="2.45" metricValueMax="2.6" metricValueMin="2.3" pageId="2" pageNumber="71" unit="mm" value="0.245" valueMax="0.26" valueMin="0.23">0.23–0.26 mm</quantity>
, with rough surface.
</paragraph>
<caption id="9DEE04B8FFFFFFE819EA4C78FB885E39" ID-DOI="http://doi.org/10.5281/zenodo.14515965" ID-Zenodo-Dep="14515965" httpUri="https://zenodo.org/record/14515965/files/figure.png" pageId="2" pageNumber="71" startId="2.[136,229,1567,1589]" targetBox="[313,1287,663,1529]" targetPageId="2" targetType="figure">
<paragraph id="C92E5430FFFFFFE819EA4C78FB885E39" blockId="2.[136,1452,1567,1626]" pageId="2" pageNumber="71">
<emphasis id="FBE58822FFFFFFE819EA4C78FF3B5E57" bold="true" box="[136,253,1567,1589]" pageId="2" pageNumber="71">FIGURE 1.</emphasis>
<taxonomicName id="0E912FB3FFFFFFE818604C47FE3D5E56" authorityName="Nadruz, Magno &amp; Theofilo" authorityYear="2024" box="[258,507,1568,1589]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="2" pageNumber="71" phylum="Tracheophyta" rank="species" species="argyrophyllum">
<emphasis id="FBE58822FFFFFFE818604C47FE3D5E56" box="[258,507,1568,1589]" italics="true" pageId="2" pageNumber="71">Anthurium argyrophyllum</emphasis>
</taxonomicName>
.
<emphasis id="FBE58822FFFFFFE81B654C78FDD85E57" bold="true" box="[519,542,1567,1588]" pageId="2" pageNumber="71">A.</emphasis>
Habit. B–C. Mid petiole in cross section. D. Proximal midrib in cross section. E. Mid midrib in cross section. F. Distal midrib in cross section. G–H. Mid peduncle in cross section. I. Inflorescence.
</paragraph>
</caption>
<paragraph id="C92E5430FFFFFFE819DF4CEFFA885F51" blockId="2.[136,1452,1672,2059]" pageId="2" pageNumber="71">
<emphasis id="FBE58822FFFFFFE819DF4CEFFE2E5EC1" bold="true" box="[189,488,1672,1698]" pageId="2" pageNumber="71">Habitat and distribution:</emphasis>
—
<taxonomicName id="0E912FB3FFFFFFE81B664CEFFCE35EC1" authorityName="Nadruz, Magno &amp; Theofilo" authorityYear="2024" box="[516,805,1672,1698]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="2" pageNumber="71" phylum="Tracheophyta" rank="species" species="argyrophyllum">
<emphasis id="FBE58822FFFFFFE81B664CEFFCE35EC1" box="[516,805,1672,1698]" italics="true" pageId="2" pageNumber="71">Anthurium argyrophyllum</emphasis>
</taxonomicName>
is a typically rupicolous species with occurrence restricted to rocky outcrops associated with rainforests currently used for cocoa crop (cabruca). Its distribution is restricted to forests located between the Rio Jequitinhonha and the Rio de Contas (
<figureCitation id="51AA48B5FFFFFFE81AFB4CB7FC185E89" box="[921,990,1744,1770]" captionStart="FIGURE 2" captionStartId="3.[136,229,990,1012]" captionTargetBox="[250,1337,196,960]" captionTargetId="figure-20@3.[244,1343,190,966]" captionTargetPageId="3" captionText="FIGURE 2. Map showing the geographic distribution of Anthurium argyrophyllum, Anthurium cachoeirense, Anthurium luschnathianum and Anthurium sagrilloanum in Brazil." figureDoi="http://doi.org/10.5281/zenodo.14515967" httpUri="https://zenodo.org/record/14515967/files/figure.png" pageId="2" pageNumber="71">Fig. 2</figureCitation>
), although it has no confirmed records in conservation units, its occurrence is indicated in the Lapão Environmental Protection Area (
<collectionCode id="AF80CCF5FFFFFFE81DED4C93FB5A5F6D" box="[1167,1180,1780,1806]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="2" pageNumber="71" type="Herbarium">J</collectionCode>
.
<collectionCode id="AF80CCF5FFFFFFE81DC24C93FB735F6D" box="[1184,1205,1780,1806]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="2" pageNumber="71" type="Herbarium">E</collectionCode>
. Santos
<emphasis id="FBE58822FFFFFFE81C714C92FA5A5F6D" box="[1299,1436,1781,1806]" italics="true" pageId="2" pageNumber="71">pers. comm.</emphasis>
). The measured size for the
<typeStatus id="162AEA92FFFFFFE818D24D7EFE265F50" box="[432,480,1817,1843]" pageId="2" pageNumber="71">type</typeStatus>
population is ca. of 600 individuals, including well-established young people.
</paragraph>
<paragraph id="C92E5430FFFFFFE819DF4D5BFB0C5FFD" blockId="2.[136,1452,1672,2059]" pageId="2" pageNumber="71">
<emphasis id="FBE58822FFFFFFE819DF4D5BFE725F35" bold="true" box="[189,436,1852,1878]" pageId="2" pageNumber="71">Conservation status:</emphasis>
—The species has little collection effort about the amount of forest fragments between the Rio Jequitinhonha and the Rio de Contas. As a result, the species is considered Data Deficient (
<collectionCode id="AF80CCF5FFFFFFE81DD94D07FB205F19" box="[1211,1254,1888,1914]" country="India" lsid="urn:lsid:biocol.org:col:13033" name="Forest Research Institute, Indian Council of Forestry Research and Education" pageId="2" pageNumber="71" type="Herbarium">DD</collectionCode>
) according to the International Union for Conservation of Nature (
<bibRefCitation id="AD0029C1FFFFFFE81BCA4DE3FCE95FFD" author="IUCN &amp; Species Survival Commission, IUCN" box="[680,815,1924,1950]" pageId="2" pageNumber="71" refId="ref3916" refString="IUCN (2012) IUCN Red List Categories and Criteria, Version 3.1. IUCN Species Survival Commission, IUCN, Gland, Switzerland, Cambridge, United Kingdom, 32 pp." type="book" year="2012">IUCN 2012</bibRefCitation>
) until more information is available.
</paragraph>
<paragraph id="C92E5430FFFFFFE819DF4DCFFF0A5F85" blockId="2.[136,1452,1672,2059]" pageId="2" pageNumber="71">
<emphasis id="FBE58822FFFFFFE819DF4DCFFEFC5FA1" bold="true" box="[189,314,1960,1986]" pageId="2" pageNumber="71">Eponymy:</emphasis>
—The specific epithet
<taxonomicName id="0E912FB3FFFFFFE81B294DCFFCC65FA1" authorityName="Nadruz, Magno &amp; Theofilo" authorityYear="2024" box="[587,768,1960,1986]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="2" pageNumber="71" phylum="Tracheophyta" rank="species" species="argyrophyllum">“argyrophyllum</taxonomicName>
” is given by the greenish-silver characteristic of the leaf blade.
</paragraph>
<paragraph id="C92E5430FFFFFFE819DF4D97FBD45069" blockId="2.[136,1452,1672,2059]" box="[189,1042,2032,2059]" pageId="2" pageNumber="71">
<emphasis id="FBE58822FFFFFFE819DF4D97FE855069" bold="true" box="[189,323,2032,2058]" pageId="2" pageNumber="71">Phenology:</emphasis>
—Flowering in January and October. Infructescence in October.
</paragraph>
<caption id="9DEE04B8FFFEFFE919EA49B9FE385C7A" ID-DOI="http://doi.org/10.5281/zenodo.14515967" ID-Zenodo-Dep="14515967" httpUri="https://zenodo.org/record/14515967/files/figure.png" pageId="3" pageNumber="72" startId="3.[136,229,990,1012]" targetBox="[250,1337,196,960]" targetPageId="3" targetType="figure">
<paragraph id="C92E5430FFFEFFE919EA49B9FE385C7A" blockId="3.[136,1451,990,1049]" pageId="3" pageNumber="72">
<emphasis id="FBE58822FFFEFFE919EA49B9FF315B90" bold="true" box="[136,247,990,1012]" pageId="3" pageNumber="72">FIGURE 2</emphasis>
. Map showing the geographic distribution of
<taxonomicName id="0E912FB3FFFEFFE91BCE49B8FC635B97" authorityName="Nadruz, Magno &amp; Theofilo" authorityYear="2024" box="[684,933,991,1012]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="3" pageNumber="72" phylum="Tracheophyta" rank="species" species="argyrophyllum">
<emphasis id="FBE58822FFFEFFE91BCE49B8FC635B97" box="[684,933,991,1012]" italics="true" pageId="3" pageNumber="72">Anthurium argyrophyllum</emphasis>
</taxonomicName>
,
<taxonomicName id="0E912FB3FFFEFFE91AD249B8FB5D5B97" authorityName="Valadares &amp; Sakuragui" authorityYear="2015" box="[944,1179,991,1012]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="3" pageNumber="72" phylum="Tracheophyta" rank="species" species="cachoeirense">
<emphasis id="FBE58822FFFEFFE91AD249B8FB5D5B97" box="[944,1179,991,1012]" italics="true" pageId="3" pageNumber="72">Anthurium cachoeirense</emphasis>
</taxonomicName>
,
<taxonomicName id="0E912FB3FFFEFFE91DC449B8FA6D5B97" authorityName="Kunth" authorityYear="1841" box="[1190,1451,991,1012]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="3" pageNumber="72" phylum="Tracheophyta" rank="species" species="luschnathianum">
<emphasis id="FBE58822FFFEFFE91DC449B8FA6D5B97" box="[1190,1451,991,1012]" italics="true" pageId="3" pageNumber="72">Anthurium luschnathianum</emphasis>
</taxonomicName>
and
<taxonomicName id="0E912FB3FFFEFFE919D34E64FE5B5C7B" box="[177,413,1027,1048]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="3" pageNumber="72" phylum="Tracheophyta" rank="species" species="sagrilloanum">
<emphasis id="FBE58822FFFEFFE919D34E64FE5B5C7B" box="[177,413,1027,1048]" italics="true" pageId="3" pageNumber="72">Anthurium sagrilloanum</emphasis>
</taxonomicName>
in Brazil.
</paragraph>
</caption>
<caption id="9DEE04B8FFFEFFE919EA4DB2FEB55073" ID-DOI="http://doi.org/10.5281/zenodo.14515969" ID-Zenodo-Dep="14515969" httpUri="https://zenodo.org/record/14515969/files/figure.png" pageId="3" pageNumber="72" startId="3.[136,229,2005,2027]" targetBox="[350,1235,1086,1969]" targetPageId="3" targetType="figure">
<paragraph id="C92E5430FFFEFFE919EA4DB2FEB55073" blockId="3.[136,1452,2005,2064]" pageId="3" pageNumber="72">
<emphasis id="FBE58822FFFEFFE919EA4DB2FF395F89" bold="true" box="[136,255,2005,2027]" pageId="3" pageNumber="72">FIGURE 3.</emphasis>
<taxonomicName id="0E912FB3FFFEFFE918644DB1FDC75F88" authorityName="Nadruz, Magno &amp; Theofilo" authorityYear="2024" box="[262,513,2006,2027]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="3" pageNumber="72" phylum="Tracheophyta" rank="species" species="argyrophyllum">
<emphasis id="FBE58822FFFEFFE918644DB1FDC75F88" box="[262,513,2006,2027]" italics="true" pageId="3" pageNumber="72">Anthurium argyrophyllum</emphasis>
</taxonomicName>
. A. Habit. B. Prophylls and cataphylls forming a fibrous mass. C. Spadix in female anthesis. D. Spadix in male anthesis.
</paragraph>
</caption>
<caption id="9DEE04B8FFF9FFEE19EA4E5AFB475C30" ID-DOI="http://doi.org/10.5281/zenodo.14515971" ID-Zenodo-Dep="14515971" box="[136,1153,1085,1107]" httpUri="https://zenodo.org/record/14515971/files/figure.png" pageId="4" pageNumber="73" startId="4.[136,229,1085,1107]" targetBox="[368,1217,202,1049]" targetPageId="4" targetType="figure">
<paragraph id="C92E5430FFF9FFEE19EA4E5AFB475C30" blockId="4.[136,1153,1085,1107]" box="[136,1153,1085,1107]" pageId="4" pageNumber="73">
<emphasis id="FBE58822FFF9FFEE19EA4E5AFF385C31" bold="true" box="[136,254,1085,1107]" pageId="4" pageNumber="73">FIGURE 4.</emphasis>
<taxonomicName id="0E912FB3FFF9FFEE18614E5AFE3B5C31" authorityName="Nadruz, Magno &amp; Theofilo" authorityYear="2024" box="[259,509,1085,1106]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="4" pageNumber="73" phylum="Tracheophyta" rank="species" species="argyrophyllum">
<emphasis id="FBE58822FFF9FFEE18614E5AFE3B5C31" box="[259,509,1085,1106]" italics="true" pageId="4" pageNumber="73">Anthurium argyrophyllum</emphasis>
</taxonomicName>
. A. Habit. B. Spadix in male anthesis. C. Detals stipite. D. Berries.
</paragraph>
</caption>
<caption id="9DEE04B8FFF9FFEE19EA4D8BFEDD5045" ID-DOI="http://doi.org/10.5281/zenodo.14515977" ID-Zenodo-Dep="14515977" httpUri="https://zenodo.org/record/14515977/files/figure.png" pageId="4" pageNumber="73" startId="4.[136,229,2028,2050]" targetBox="[366,1218,1141,1996]" targetPageId="4" targetType="figure">
<paragraph id="C92E5430FFF9FFEE19EA4D8BFEDD5045" blockId="4.[136,1452,2028,2086]" pageId="4" pageNumber="73">
<emphasis id="FBE58822FFF9FFEE19EA4D8BFEC55062" bold="true" box="[136,259,2028,2050]" pageId="4" pageNumber="73">FIGURE 5.</emphasis>
<taxonomicName id="0E912FB3FFF9FFEE186D4D8BFDC85062" authorityName="Nadruz, Magno &amp; Theofilo" authorityYear="2024" box="[271,526,2028,2049]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="4" pageNumber="73" phylum="Tracheophyta" rank="species" species="argyrophyllum">
<emphasis id="FBE58822FFF9FFEE186D4D8BFDC85062" box="[271,526,2028,2049]" italics="true" pageId="4" pageNumber="73">Anthurium argyrophyllum</emphasis>
</taxonomicName>
. A. Spadix in pre-anthesis. B. Spadix in female anthesis. C. Spadix in male anthesis. D. Infructescence.
</paragraph>
</caption>
<caption id="9DEE04B8FFF8FFEF19EA4E06FC745C14" box="[136,946,1121,1143]" pageId="5" pageNumber="74" startId="5.[136,229,1121,1143]" targetType="figure">
<paragraph id="C92E5430FFF8FFEF19EA4E06FC745C14" blockId="5.[136,946,1121,1143]" box="[136,946,1121,1143]" pageId="5" pageNumber="74">
<emphasis id="FBE58822FFF8FFEF19EA4E06FF3B5C15" bold="true" box="[136,253,1121,1143]" pageId="5" pageNumber="74">FIGURE 6.</emphasis>
<taxonomicName id="0E912FB3FFF8FFEF18614E06FE3B5C15" authorityName="Nadruz, Magno &amp; Theofilo" authorityYear="2024" box="[259,509,1121,1142]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="5" pageNumber="74" phylum="Tracheophyta" rank="species" species="argyrophyllum">
<emphasis id="FBE58822FFF8FFEF18614E06FE3B5C15" box="[259,509,1121,1142]" italics="true" pageId="5" pageNumber="74">Anthurium argyrophyllum</emphasis>
</taxonomicName>
. A. Habitat. B. Mature plant. C. Young plant.
</paragraph>
</caption>
<paragraph id="C92E5430FFF8FFEF19DF4EC1FC8A5D6B" blockId="5.[136,1452,1190,2080]" pageId="5" pageNumber="74">
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<emphasis id="FBE58822FFF8FFEF19DF4EC1FEF95CA3" bold="true" box="[189,319,1190,1216]" pageId="5" pageNumber="74">
<typeStatus id="162AEA92FFF8FFEF19DF4EC1FEFF5CA3" box="[189,313,1190,1216]" pageId="5" pageNumber="74" type="paratype">Paratypes</typeStatus>
:
</emphasis>
—
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.
<collectingRegion id="0B559AD2FFF8FFEF18AF4EC1FDD55CA3" box="[461,531,1190,1216]" country="Brazil" name="Bahia" pageId="5" pageNumber="74">Bahia</collectingRegion>
:
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, fazenda do senhor
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(cultivated),
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,
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. Magno &amp;
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.
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. Santos 01
</emphasis>
(
<collectionCode id="AF80CCF5FFF8FFEF183A4EADFE665C87" box="[344,416,1226,1252]" country="Brazil" httpUri="http://grbio.org/cool/7078-dpb4" name="Federal University of Espírito Santo" pageId="5" pageNumber="74" type="Herbarium">VIES</collectionCode>
!)
</materialsCitation>
;
<materialsCitation id="79F95E6DFFF8FFEF18DB4EADFE9C5D6B" collectingDate="2023-10-07" collectionCode="A, J, E, VIES" country="Brazil" elevation="190" location="Lapao" municipality="Santa Luzia" pageId="5" pageNumber="74" specimenCount="1" typeStatus="paratype">
<collectingMunicipality id="294ACE4AFFF8FFEF18DB4EADFD845C87" box="[441,578,1226,1252]" pageId="5" pageNumber="74">Santa Luzia</collectingMunicipality>
, aos arredores da APA do
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,
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<elevation id="42BCB303FFF8FFEF1AA24EADFBCC5C87" box="[960,1034,1226,1252]" metricMagnitude="2" metricUnit="m" metricValue="1.9" pageId="5" pageNumber="74" unit="m" value="190.0">190 m</elevation>
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,
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<collectingDate id="AD6B8B18FFF8FFEF1D754EADFB115C87" box="[1047,1239,1226,1252]" pageId="5" pageNumber="74" value="2023-10-07">07 October 2023</collectingDate>
</date>
,
<emphasis id="FBE58822FFF8FFEF1D814EADFF305D6B" italics="true" pageId="5" pageNumber="74">
<collectionCode id="AF80CCF5FFF8FFEF1D814EADFB315C87" box="[1251,1271,1226,1252]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="5" pageNumber="74" type="Herbarium">A</collectionCode>
. Magno &amp;
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.
<collectionCode id="AF80CCF5FFF8FFEF1CF14EADFA6E5C80" box="[1427,1448,1226,1251]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="5" pageNumber="74" type="Herbarium">E</collectionCode>
. Santos 18
</emphasis>
(
<collectionCode id="AF80CCF5FFF8FFEF18674E89FE8B5D6B" box="[261,333,1262,1288]" country="Brazil" httpUri="http://grbio.org/cool/7078-dpb4" name="Federal University of Espírito Santo" pageId="5" pageNumber="74" type="Herbarium">VIES</collectionCode>
!)
</materialsCitation>
;
<materialsCitation id="79F95E6DFFF8FFEF18044E89FC8E5D6B" box="[358,840,1262,1288]" collectingDate="2023-10-07" collectionCode="A, J, E, VIES" country="Brazil" elevation="190" location="Lapao" municipality="Santa Luzia" pageId="5" pageNumber="74" specimenCount="1" typeStatus="paratype">
<emphasis id="FBE58822FFF8FFEF18044E89FE5A5D6B" box="[358,412,1262,1288]" italics="true" pageId="5" pageNumber="74">idem</emphasis>
,
<emphasis id="FBE58822FFF8FFEF18C84E89FD225D6B" box="[426,740,1262,1288]" italics="true" pageId="5" pageNumber="74">
<collectionCode id="AF80CCF5FFF8FFEF18C84E89FE785D6B" box="[426,446,1262,1288]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="5" pageNumber="74" type="Herbarium">A</collectionCode>
. Magno &amp;
<collectionCode id="AF80CCF5FFF8FFEF1B5E4E89FD8A5D64" box="[572,588,1262,1287]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="5" pageNumber="74" type="Herbarium">J</collectionCode>
.
<collectionCode id="AF80CCF5FFF8FFEF1B354E89FDAA5D64" box="[599,620,1262,1287]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="5" pageNumber="74" type="Herbarium">E</collectionCode>
. Santos 19
</emphasis>
(
<collectionCode id="AF80CCF5FFF8FFEF1B914E89FCFD5D6B" box="[755,827,1262,1288]" country="Brazil" httpUri="http://grbio.org/cool/7078-dpb4" name="Federal University of Espírito Santo" pageId="5" pageNumber="74" type="Herbarium">VIES</collectionCode>
!)
</materialsCitation>
.
</paragraph>
<paragraph id="C92E5430FFF8FFEF19DF4F75FE0C5E4B" blockId="5.[136,1452,1190,2080]" pageId="5" pageNumber="74">
<emphasis id="FBE58822FFF8FFEF19DF4F75FE145D48" bold="true" box="[189,466,1297,1324]" pageId="5" pageNumber="74">Features and affinities:</emphasis>
—Specimens found in nature have predominantly spreading leaves (
<figureCitation id="51AA48B5FFF8FFEF1DAC4F75FAD45D4F" box="[1230,1298,1298,1324]" captionStart="FIGURE 6" captionStartId="5.[136,229,1121,1143]" captionText="FIGURE 6. Anthurium argyrophyllum. A. Habitat. B. Mature plant. C. Young plant." pageId="5" pageNumber="74">Fig. 6</figureCitation>
), but when in cultivation their leaves are erect (
<figureCitation id="51AA48B5FFF8FFEF18994F51FD9D5D33" box="[507,603,1334,1360]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="3.[136,229,2005,2027]" captionStartId-1="4.[136,229,1085,1107]" captionTargetBox-0="[350,1235,1086,1969]" captionTargetBox-1="[368,1217,202,1049]" captionTargetId-0="figure-45@3.[340,1247,1074,1981]" captionTargetId-1="figure-21@4.[358,1229,190,1061]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 3. Anthurium argyrophyllum. A. Habit. B. Prophylls and cataphylls forming a fibrous mass. C. Spadix in female anthesis. D. Spadix in male anthesis." captionText-1="FIGURE 4. Anthurium argyrophyllum. A. Habit. B. Spadix in male anthesis. C. Detals stipite. D. Berries." figureDoi-0="http://doi.org/10.5281/zenodo.14515969" figureDoi-1="http://doi.org/10.5281/zenodo.14515971" httpUri-0="https://zenodo.org/record/14515969/files/figure.png" httpUri-1="https://zenodo.org/record/14515971/files/figure.png" pageId="5" pageNumber="74">Fig. 3–4</figureCitation>
) and can be confused with
<taxonomicName id="0E912FB3FFF8FFEF1AE94F51FA6A5D33" authority="Theofilo &amp; T. F. Sagrillo" authorityName="Theofilo &amp; T. F. Sagrillo" box="[907,1452,1334,1360]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="5" pageNumber="74" phylum="Tracheophyta" rank="species" species="sagrilloanum">
<emphasis id="FBE58822FFF8FFEF1AE94F51FB585D33" box="[907,1182,1334,1360]" italics="true" pageId="5" pageNumber="74">Anthurium sagrilloanum</emphasis>
Theófilo &amp; T.F.Sagrillo
</taxonomicName>
in
<bibRefCitation id="AD0029C1FFF8FFEF19C14F3DFE015D17" author="Valadares, R. T. &amp; Sagrillo, T. F. &amp; Sakuragui, C. M." box="[163,455,1370,1396]" pageId="5" pageNumber="74" pagination="60 - 63" refId="ref4880" refString="Valadares, R. T., Sagrillo, T. F. &amp; Sakuragui, C. M. (2020) Anthurium sagrilloanum: a new species of Anthurium (Araceae) from Southeastern Brazil. Novon 28: 60 - 63. https: // doi. org / 10.3417 / 2019424" type="journal article" year="2020">
Valadares
<emphasis id="FBE58822FFF8FFEF18744F3DFE8B5D17" box="[278,333,1370,1396]" italics="true" pageId="5" pageNumber="74">et al.</emphasis>
(2020: 60)
</bibRefCitation>
differing from this in having prophylls and cataphylls chestnut to straw colored when old, forming fibrous mass (vs. chestnut or brownish to greenish when old, persistent and entire at stem apex, deciduous at stem base), petiole 2.4–2.6 × larger than the peduncle (vs. usually smaller), basal veins absent to 1 (vs. basal veins 2 to 4), primary lateral veins 11 to 16 on both sides (vs. 14–24 on both sides), peduncle curved in fruiting (
<figureCitation id="51AA48B5FFF8FFEF1C4C4FA1FAB75D83" box="[1326,1393,1477,1504]" captionStart="FIGURE 5" captionStartId="4.[136,229,2028,2050]" captionTargetBox="[366,1218,1141,1996]" captionTargetId="figure-41@4.[358,1229,1133,2004]" captionTargetPageId="4" captionText="FIGURE 5. Anthurium argyrophyllum. A. Spadix in pre-anthesis. B. Spadix in female anthesis. C. Spadix in male anthesis. D. Infructescence." figureDoi="http://doi.org/10.5281/zenodo.14515977" httpUri="https://zenodo.org/record/14515977/files/figure.png" pageId="5" pageNumber="74">Fig. 5</figureCitation>
) (vs. peduncle spreading in fruiting) spathe flat (vs. naviculate), spadix stipitate and greenish-purplish in anthesis (vs. spadix sessile, purplish in anthesis).
</paragraph>
<paragraph id="C92E5430FFF8FFEF19DF4C55FC305EDB" blockId="5.[136,1452,1190,2080]" pageId="5" pageNumber="74">
Also noteworthy is its morphological proximity to
<taxonomicName id="0E912FB3FFF8FFEF1A664C55FACF5E2F" authority="Kunth (1841: 73)" authorityName="Kunth" authorityPageNumber="73" authorityYear="1841" box="[772,1289,1586,1612]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="5" pageNumber="74" phylum="Tracheophyta" rank="species" species="luschnathianum">
<emphasis id="FBE58822FFF8FFEF1A664C55FBFE5E2F" box="[772,1080,1586,1612]" italics="true" pageId="5" pageNumber="74">Anthurium luschnathianum</emphasis>
<bibRefCitation id="AD0029C1FFF8FFEF1D224C55FACF5E2F" author="Kunth, C. S." box="[1088,1289,1586,1612]" pageId="5" pageNumber="74" refId="ref4039" refString="Kunth, C. S. (1841) Enumeratio Plantarum Omnium hucusque Cognitarum 3. Sumtibus J. g. Cottae, Stutgardiae and Tubingae, 644 pp." type="book" year="1841">Kunth (1841: 73)</bibRefCitation>
</taxonomicName>
, distinguished by its erect stem (vs. decumbent stem with erect apex), prophylls and cataphylls forming a fibrous mass when old (vs. little decomposed to deciduous towards the base of the stem), peduncle
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long, curved during fruiting (vs. peduncle
<quantity id="0E69F9D5FFF8FFEF19964CFAFE4B5EDB" box="[244,397,1693,1720]" metricMagnitude="-1" metricUnit="m" metricValue="4.845" metricValueMax="6.54" metricValueMin="3.15" pageId="5" pageNumber="74" unit="cm" value="48.45" valueMax="65.4" valueMin="31.5">31.5–65.4 cm</quantity>
long, not curved), spathe ovate (vs. spathe lanceolate).
</paragraph>
<paragraph id="C92E5430FFF8FFEF19DF4CA5FB215043" blockId="5.[136,1452,1190,2080]" pageId="5" pageNumber="74">
<taxonomicName id="0E912FB3FFF8FFEF19DF4CA5FE265EBF" authorityName="Nadruz, Magno &amp; Theofilo" authorityYear="2024" box="[189,480,1730,1756]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="5" pageNumber="74" phylum="Tracheophyta" rank="species" species="argyrophyllum">
<emphasis id="FBE58822FFF8FFEF19DF4CA5FE265EBF" box="[189,480,1730,1756]" italics="true" pageId="5" pageNumber="74">Anthurium argyrophyllum</emphasis>
</taxonomicName>
is provisionally included in the
<taxonomicName id="0E912FB3FFF8FFEF1A254CA5FB125EBF" authority="Engl." authorityName="Engl." box="[839,1236,1730,1756]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="5" pageNumber="74" phylum="Tracheophyta" rank="section" section="Urospadix">
<emphasis id="FBE58822FFF8FFEF1A254CA5FC795EBF" box="[839,959,1730,1756]" italics="true" pageId="5" pageNumber="74">Anthurium</emphasis>
section
<emphasis id="FBE58822FFF8FFEF1D7E4CA5FB575EBF" box="[1052,1169,1730,1756]" italics="true" pageId="5" pageNumber="74">Urospadix</emphasis>
Engl.
</taxonomicName>
because it presents characteristics very close to species with a curved peduncle during fruiting (e.g.
<taxonomicName id="0E912FB3FFF8FFEF1D7E4C81FB0D5F63" authorityName="Barroso" authorityYear="1957" box="[1052,1227,1766,1792]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="5" pageNumber="74" phylum="Tracheophyta" rank="species" species="cleistanthum">
<emphasis id="FBE58822FFF8FFEF1D7E4C81FB0D5F63" box="[1052,1227,1766,1792]" italics="true" pageId="5" pageNumber="74">A. cleistanthum</emphasis>
</taxonomicName>
,
<taxonomicName id="0E912FB3FFF8FFEF1DB94C81FAB75E9C" box="[1243,1393,1766,1792]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="5" pageNumber="74" phylum="Tracheophyta" rank="species" species="coriaceum">
<emphasis id="FBE58822FFF8FFEF1DB94C81FAB75E9C" box="[1243,1393,1766,1792]" italics="true" pageId="5" pageNumber="74">A. coriaceum</emphasis>
</taxonomicName>
) and is restricted to eastern
<collectingCountry id="B18614A0FFF8FFEF18F34D6DFE1F5F47" box="[401,473,1802,1828]" name="Brazil" pageId="5" pageNumber="74">Brazil</collectingCountry>
. The polymorphism of the base of the leaf blade raises questions about its sectional positioning. For the first author (
<collectionCode id="AF80CCF5FFF8FFEF18974D49FD895F2B" box="[501,591,1838,1864]" pageId="5" pageNumber="74">MANC</collectionCode>
), the species should be included in a new section comprising all species with leaf blades containing a chordate base and variations. For the third author (
<collectionCode id="AF80CCF5FFF8FFEF1A844D35FBE65F0F" box="[998,1056,1874,1900]" pageId="5" pageNumber="74">RTV</collectionCode>
) this hypothesis can be supported, but restricted to species with a strictly chordate base and a non-curved peduncle during fruiting (e.g.
<taxonomicName id="0E912FB3FFF8FFEF1C564D11FD4E5FD7" authority="Theofilo, L. Kollmann &amp; Sakur." authorityName="Theofilo, L. Kollmann &amp; Sakur." class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="5" pageNumber="74" phylum="Tracheophyta" rank="species" species="marcusianum">
<emphasis id="FBE58822FFF8FFEF1C564D11FED95FD0" italics="true" pageId="5" pageNumber="74">Anthurium marcusianum</emphasis>
Theófilo, L.Kollmann &amp; Sakur.
</taxonomicName>
in
<bibRefCitation id="AD0029C1FFF8FFEF1BC94DFDFC0E5FD7" author="Valadares, R. T. &amp; Kollmann, L. J. C. &amp; Sakuragui, C. M." box="[683,968,1946,1972]" pageId="5" pageNumber="74" pagination="3 - 7" refId="ref4824" refString="Valadares, R. T., Kollmann, L. J. C. &amp; Sakuragui, C. M. (2019) New species of Anthurium (Araceae) with cordate leaves from Southeastern Brazil. Novon 27: 3 - 7. https: // doi. org / 10.3417 / 2018270" type="journal article" year="2019">
Valadares
<emphasis id="FBE58822FFF8FFEF1A7D4DFDFC9E5FD7" box="[799,856,1946,1972]" italics="true" pageId="5" pageNumber="74">et al.</emphasis>
(2019: 3)
</bibRefCitation>
,
<taxonomicName id="0E912FB3FFF8FFEF1AB64DFDFA6D5FD7" authority="K. Koch &amp; Lauche" authorityName="K. Koch &amp; Lauche" box="[980,1451,1946,1972]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="5" pageNumber="74" phylum="Tracheophyta" rank="species" species="augustinum">
<emphasis id="FBE58822FFF8FFEF1AB64DFDFB105FD0" box="[980,1238,1946,1972]" italics="true" pageId="5" pageNumber="74">Anthurium augustinum</emphasis>
K.Koch &amp; Lauche
</taxonomicName>
in
<bibRefCitation id="AD0029C1FFF8FFEF19C44DD9FE935FBB" author="Koch, K." box="[166,341,1981,2008]" pageId="5" pageNumber="74" pagination="2 - 8" refId="ref4015" refString="Koch, K. (1855) Aroideae novae et criticae. Index seminum in Horto Botanico Berolinensi 1855: 2 - 8." type="journal article" year="1855">Koch (1855: 7)</bibRefCitation>
,
<taxonomicName id="0E912FB3FFF8FFEF18004DD9FCB95FBB" authority="Barroso (1970: 1))" authorityName="Barroso" authorityPageNumber="1" authorityYear="1970" box="[354,895,1982,2008]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="5" pageNumber="74" phylum="Tracheophyta" rank="species" species="xanthophylloides">
<emphasis id="FBE58822FFF8FFEF18004DD9FD675FBB" box="[354,673,1982,2008]" italics="true" pageId="5" pageNumber="74">Anthurium xanthophylloides</emphasis>
<bibRefCitation id="AD0029C1FFF8FFEF1BC84DD9FCB95FBB" author="Barroso, G. M." box="[682,895,1982,2008]" pageId="5" pageNumber="74" pagination="1 - 4" refId="ref3055" refString="Barroso, G. M. (1970) Especies novas de Anthurium Schott. Loefgrenia 46: 1 - 4." type="book chapter" year="1970">Barroso (1970: 1))</bibRefCitation>
</taxonomicName>
. In any case, it is complicated to assume that the evolutionary history of this lineage constitutes a disjunction from the
<taxonomicName id="0E912FB3FFF8FFEF1AF84D85FF785043" authority="Schott (1860: 528)" authorityName="Schott" authorityPageNumber="528" authorityYear="1860" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="5" pageNumber="74" phylum="Tracheophyta" rank="section" section="Belolonchium">
<emphasis id="FBE58822FFF8FFEF1AF84D85FBD45F9F" box="[922,1042,2018,2044]" italics="true" pageId="5" pageNumber="74">Anthurium</emphasis>
section
<emphasis id="FBE58822FFF8FFEF1D124D85FACA5F9F" box="[1136,1292,2018,2044]" italics="true" pageId="5" pageNumber="74">Belolonchium</emphasis>
<bibRefCitation id="AD0029C1FFF8FFEF1C714D85FF785043" author="Schott, H. W." pageId="5" pageNumber="74" refId="ref4539" refString="Schott, H. W. (1860) Prodromus Systematis Aroidearum. Typis Congregationis Mechitheristicae, Vindobae, 602 pp." type="book" year="1860">Schott (1860: 528)</bibRefCitation>
</taxonomicName>
, an argument supported by previous molecular studies (
<bibRefCitation id="AD0029C1FFF8FFEF1A4C4261FC135043" author="Temponi, L. G." box="[814,981,2054,2080]" pageId="5" pageNumber="74" refId="ref4714" refString="Temponi, L. G. (2006) Sistematica de Anthurium sect. Urospadix (Araceae). Ph. D. Thesis, Instituto de Biociencias, Universidade de Sao Paulo, Sao Paulo. https: // doi. org / 10.11606 / T. 41.2007. tde- 21082007 - 143011" type="book" year="2006">Temponi 2006</bibRefCitation>
,
<bibRefCitation id="AD0029C1FFF8FFEF1A824261FB1A5043" author="Carlsen, M. &amp; Croat, T. B." box="[992,1244,2054,2080]" pageId="5" pageNumber="74" pagination="576 - 588" refId="ref3264" refString="Carlsen, M. &amp; Croat, T. B. (2013) A Molecular Phylogeny of the Species-Rich Neotropical Genus Anthurium (Araceae) Based on Combined Chloroplast and Nuclear DNA. Systematic Botany 38: 576 - 588. https: // doi. org / 10.1600 / 036364413 X 670287" type="journal article" year="2013">Carlsen &amp; Croat 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="8FC3FD2797F969F74FD389307A5AD136">Cirsium wulongense (Asteraceae, Cardueae), a new species from Chongqing, China</mods:title>
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<mods:namePart id="ABB89487FBBA5FD3B54C984EC420C315">Jin, Zi-Chao</mods:namePart>
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<mods:affiliation id="582391454A714AF34A851B7854E7409B">Plant Science Research Center, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou, Guangdong 510650, China</mods:affiliation>
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<mods:namePart id="3C7C632B8F8B7C0E1BDC3B434C8F4007">Chen, Feng</mods:namePart>
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<mods:affiliation id="C0F0E6C08C3DBB1B68D9CEEBFB69196E">Chongqing Museum of Natural History, Beibei, Chongqing 400799, China</mods:affiliation>
<mods:nameIdentifier id="81A437731A505FE830991EDE2130E2DA" type="email">fengchen408@163.com</mods:nameIdentifier>
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<mods:namePart id="E145FB3A0CA94553CE0D5C16ED3522CA">Chen, You-Sheng</mods:namePart>
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<mods:affiliation id="718633AD810BDDAA3846D57F47A94947">Plant Science Research Center, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou, Guangdong 510650, China &amp; Key Laboratory of National Forestry and Grassland Administration on Plant Conservation and Utilization in Southern China, Guangzhou 510650, China</mods:affiliation>
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<subSubSection id="5BA1A0EAFFC3E15064AA8FA59E23FA46" box="[136,811,1460,1487]" pageId="3" pageNumber="204" type="nomenclature">
<paragraph id="1304F361FFC3E15064AA8FA59E23FA46" blockId="3.[136,811,1460,1487]" box="[136,811,1460,1487]" pageId="3" pageNumber="204">
<heading id="484C440DFFC3E15064AA8FA59E23FA46" box="[136,811,1460,1487]" fontSize="11" level="2" pageId="3" pageNumber="204" reason="3">
<taxonomicName id="D4BB88E2FFC3E15064AA8FA59F64FA46" ID-CoL="CJYMQ" authority="Z.C. Jin &amp; Y.S. Chen" authorityName="Z. C. Jin &amp; Y. S. Chen" authorityYear="2024" box="[136,620,1460,1487]" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="204" phylum="Tracheophyta" rank="species" species="wulongense" status="sp. nov.">
<emphasis id="21CF2F73FFC3E15064AA8FA59C7DFA47" bold="true" box="[136,373,1460,1487]" italics="true" pageId="3" pageNumber="204">Cirsium wulongense</emphasis>
Z.C. Jin &amp; Y.S. Chen
</taxonomicName>
,
<emphasis id="21CF2F73FFC3E15066558FA59FD8FA46" bold="true" box="[631,720,1460,1486]" pageId="3" pageNumber="204">
<taxonomicNameLabel id="3AFC9208FFC3E15066558FA59FD8FA46" box="[631,720,1460,1486]" pageId="3" pageNumber="204" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="8B80EFE4FFC3E15066C28FA59E2AFA46" box="[736,802,1460,1486]" captionStart="FIGURE 3" captionStartId="4.[136,229,1953,1975]" captionTargetBox="[178,1409,190,1928]" captionTargetId="figure-27@4.[178,1409,190,1928]" captionTargetPageId="4" captionText="FIGURE 3. Cirsium wulongense in the wild (Xiannü Shan, Wulong district, Chongqing, China). A. Habitat. B. Stem. C. Root. D. Basal leaves. E. Cauline leaves. F, G. Capitula. H. Longitudinal section of capitulum. I, J. Florets. K. Phyllaries. L. Achenes. Photographed by Feng Chen, based on F. Chen, X.B. Shi &amp; F.F. Xu XLS21-095 (IBSC)." pageId="3" pageNumber="204">Fig. 3</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5BA1A0EAFFC3E15064AA8FE79F9BF9BC" pageId="3" pageNumber="204" type="materials_examined">
<paragraph id="1304F361FFC3E15064AA8FE79F9BF9BC" blockId="3.[136,1452,1525,2092]" pageId="3" pageNumber="204">
<emphasis id="21CF2F73FFC3E15064AA8FE79DC4F998" bold="true" box="[136,204,1526,1552]" pageId="3" pageNumber="204">Type:</emphasis>
—
<collectingCountry id="6BACB3F1FFC3E15064CB8FE49C25F987" box="[233,301,1525,1551]" name="China" pageId="3" pageNumber="204">China</collectingCountry>
,
<collectingRegion id="D17F3D83FFC3E150651B8FE49CB0F987" box="[313,440,1525,1551]" country="China" name="Chongqing" pageId="3" pageNumber="204">Chongqing</collectingRegion>
, Wulong district, Xiannü Shan,
<date id="6705D5A1FFC3E150673F8FE79ED9F998" box="[797,977,1526,1552]" pageId="3" pageNumber="204" value="2021-08-14">14 August 2021</date>
, elev.
<quantity id="D4435E84FFC3E15060358FE79966F998" box="[1047,1134,1526,1552]" metricMagnitude="3" metricUnit="m" metricValue="1.78" pageId="3" pageNumber="204" unit="m" value="1780.0">1780 m</quantity>
,
<emphasis id="21CF2F73FFC3E15060588FE79C0EF9BC" italics="true" pageId="3" pageNumber="204">F. Chen, X.B. Shi &amp; F.F. Xu XLS21-095</emphasis>
(
<typeStatus id="CC004DC3FFC3E15065378C089C71F9BB" box="[277,377,1561,1587]" pageId="3" pageNumber="204" type="holotype">holotype</typeStatus>
IBSC,
<typeStatus id="CC004DC3FFC3E15065EE8C089F20F9BB" box="[460,552,1561,1587]" pageId="3" pageNumber="204" type="isotype">isotypes</typeStatus>
CQNM).
</paragraph>
</subSubSection>
<subSubSection id="5BA1A0EAFFC3E156649F8C2C9FE2FF30" lastPageId="5" lastPageNumber="206" pageId="3" pageNumber="204" type="description">
<paragraph id="1304F361FFC3E150649F8C2C9F7BF7A3" blockId="3.[136,1452,1525,2092]" pageId="3" pageNumber="204">
<emphasis id="21CF2F73FFC3E150649F8C2C9C59F9DF" bold="true" box="[189,337,1597,1623]" pageId="3" pageNumber="204">Description:</emphasis>
—Herbs
<quantity id="D4435E84FFC3E150659A8C2C9F3EF9D0" box="[440,566,1597,1624]" metricMagnitude="-1" metricUnit="m" metricValue="9.0" metricValueMax="15.0" metricValueMin="3.0" pageId="3" pageNumber="204" unit="cm" value="90.0" valueMax="150.0" valueMin="30.0">30–150 cm</quantity>
tall, perennial. Stems erect, ribbed, pubescent with densely long multicellular hairs. Basal and lower cauline leaves with petioles to
<quantity id="D4435E84FFC3E15066D58C739E37F9F4" box="[759,831,1634,1660]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="204" unit="cm" value="10.0">10 cm</quantity>
long, fringed with triangular teeth and with an apical spine ca.
<quantity id="D4435E84FFC3E15064DA8C949C50F928" box="[248,344,1669,1696]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="3" pageNumber="204" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 mm</quantity>
long; leaf blade elliptic, 15–25 ×
<quantity id="D4435E84FFC3E15066C98C949E5AF928" box="[747,850,1669,1696]" metricMagnitude="-1" metricUnit="m" metricValue="1.15" metricValueMax="1.5" metricValueMin="0.8" pageId="3" pageNumber="204" unit="cm" value="11.5" valueMax="15.0" valueMin="8.0">8–15 cm</quantity>
, pinnatipartite or pinnatisect; segments 6–8 pairs, ± narrowly elliptic or triangular, with numerous unequal triangular teeth laterally and with an apical spine ca.
<quantity id="D4435E84FFC3E150614A8CBB98A4F94C" box="[1384,1452,1706,1732]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="204" unit="mm" value="4.0">4 mm</quantity>
long. Cauline leaves sessile, auriculate amplexicaul, elliptic, ovate-elliptic, pinnatilobate to pinnatipartite; segments ca 5 pairs, triangular, with 1–3 unequal triangular teeth laterally and with an apical spine
<quantity id="D4435E84FFC3E150606A8CE399ACF884" box="[1096,1188,1778,1804]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="3" pageNumber="204" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity>
long; terminal segment largest. All leaves discolorous, abaxially grayish white and densely tomentose and abaxially densely pubescent with crispate multicellular hairs along veins, adaxially green, pubescent with crispate multicellular hairs. Capitula sparsely racemose, nodding. Involucre campanulate,
<quantity id="D4435E84FFC3E15066A18D4F9FD1F8F0" box="[643,729,1886,1912]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="3" pageNumber="204" unit="cm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 cm</quantity>
in diam., sparsely cobwebby. Phyllaries in ca. 7 rows, margin entire, lacking marginal spinules, wings, and scarious appendage, imbricate, outer phyllaries much shorter than inner ones; outer and middle phyllaries lanceolate,
<quantity id="D4435E84FFC3E15066B08DB49FF4F848" box="[658,764,1957,1984]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="3" pageNumber="204" unit="cm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 cm</quantity>
long, basal portion linear, apical portion subulate, patent to reflexed, and narrowed into a spine
<quantity id="D4435E84FFC3E15066018DDB9F6EF86C" box="[547,614,1994,2020]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="204" unit="mm" value="3.0">3 mm</quantity>
long; inner phyllaries lanceolate to linear-lanceolate, ca.
<quantity id="D4435E84FFC3E15060D68DDB9824F86C" box="[1268,1324,1994,2020]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="204" unit="cm" value="2.0">2 cm</quantity>
long, apex acuminate. Florets bisexual. Corolla white, ca.
<quantity id="D4435E84FFC3E150668B8DFC9FF0F780" box="[681,760,2029,2056]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="204" unit="cm" value="1.6">1.6 cm</quantity>
long, tube ca.
<quantity id="D4435E84FFC3E150678B8DFC9EF0F780" box="[937,1016,2029,2056]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="204" unit="cm" value="0.6">0.6 cm</quantity>
long. Achene brown, ca.
<quantity id="D4435E84FFC3E15061078DFF9861F780" box="[1317,1385,2030,2056]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="204" unit="mm" value="4.0">4 mm</quantity>
long. Pappus bristles dirty white, ca.
<quantity id="D4435E84FFC3E15065C582039F3BF7A4" box="[487,563,2066,2092]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="3" pageNumber="204" unit="cm" value="1.7">1.7 cm</quantity>
long.
</paragraph>
<caption id="47C4A3E9FFC4E15764AA8DB09E29F877" pageId="4" pageNumber="205" startId="4.[136,229,1953,1975]" targetBox="[178,1409,190,1928]" targetPageId="4" targetType="figure">
<paragraph id="1304F361FFC4E15764AA8DB09E29F877" blockId="4.[136,1451,1953,2047]" pageId="4" pageNumber="205">
<emphasis id="21CF2F73FFC4E15764AA8DB09DF6F83E" bold="true" box="[136,254,1953,1975]" pageId="4" pageNumber="205">FIGURE 3.</emphasis>
<taxonomicName id="D4BB88E2FFC4E15765268DB09CC1F83E" authorityName="Z. C. Jin &amp; Y. S. Chen" authorityYear="2024" box="[260,457,1953,1974]" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="205" phylum="Tracheophyta" rank="species" species="wulongense">
<emphasis id="21CF2F73FFC4E15765268DB09CC1F83E" box="[260,457,1953,1974]" italics="true" pageId="4" pageNumber="205">Cirsium wulongense</emphasis>
</taxonomicName>
in the wild (Xiannü Shan, Wulong district, Chongqing, China). A. Habitat. B. Stem. C. Root. D. Basal leaves. E. Cauline leaves. F, G. Capitula. H. Longitudinal section of capitulum. I, J. Florets. K. Phyllaries. L. Achenes. Photographed by Feng Chen, based on
<emphasis id="21CF2F73FFC4E15765758DFB9FD9F876" box="[343,721,2025,2047]" italics="true" pageId="4" pageNumber="205">F. Chen, X.B. Shi &amp; F.F. Xu XLS21-095</emphasis>
(IBSC).
</paragraph>
</caption>
<paragraph id="1304F361FFC5E156649F8A8F9FE2FF30" blockId="5.[136,1452,158,401]" box="[189,746,158,185]" pageId="5" pageNumber="206">
<emphasis id="21CF2F73FFC5E156649F8A8F9C4BFF30" bold="true" box="[189,323,158,184]" pageId="5" pageNumber="206">Phenology:</emphasis>
—Flowering from August to October.
</paragraph>
</subSubSection>
<subSubSection id="5BA1A0EAFFC5E156649F8AD39E5BFE88" pageId="5" pageNumber="206" type="etymology">
<paragraph id="1304F361FFC5E156649F8AD39E5BFE88" blockId="5.[136,1452,158,401]" pageId="5" pageNumber="206">
<emphasis id="21CF2F73FFC5E156649F8AD39C40FF54" bold="true" box="[189,328,194,220]" pageId="5" pageNumber="206">Etymology:</emphasis>
—The specific epithet
<taxonomicName id="D4BB88E2FFC5E156667E8AD39FE5FF54" authorityName="Z. C. Jin &amp; Y. S. Chen" authorityYear="2024" box="[604,749,194,220]" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="206" phylum="Tracheophyta" rank="species" species="wulongense">
“
<emphasis id="21CF2F73FFC5E156664B8AD39FE5FF54" box="[617,749,194,220]" italics="true" pageId="5" pageNumber="206">wulongense</emphasis>
</taxonomicName>
” is derived from the
<typeStatus id="CC004DC3FFC5E15667DC8AD29926FF55" box="[1022,1070,195,221]" pageId="5" pageNumber="206">type</typeStatus>
locality, i.e. Wulong district in
<collectingRegion id="D17F3D83FFC5E15664AA8AF79C0FFE88" box="[136,263,230,256]" country="China" name="Chongqing" pageId="5" pageNumber="206">Chongqing</collectingRegion>
,
<collectingCountry id="6BACB3F1FFC5E15665318AF79C52FE88" box="[275,346,230,256]" name="China" pageId="5" pageNumber="206">China</collectingCountry>
. The Chinese name is “
<emphasis id="21CF2F73FFC5E15666428AF69FBCFE89" bold="true" box="[608,692,231,257]" pageId="5" pageNumber="206">ẑĸfi</emphasis>
” (wu long ji).
</paragraph>
</subSubSection>
<subSubSection id="5BA1A0EAFFC5E156649F8B1B9E6EF9B2" pageId="5" pageNumber="206" type="distribution">
<paragraph id="1304F361FFC5E156649F8B1B981DFE19" blockId="5.[136,1452,158,401]" pageId="5" pageNumber="206">
<emphasis id="21CF2F73FFC5E156649F8B1B9CE1FEAC" bold="true" box="[189,489,266,292]" pageId="5" pageNumber="206">Distribution and habitat:</emphasis>
—
<taxonomicName id="D4BB88E2FFC5E15666278B1A9FE2FEAC" authorityName="Z. C. Jin &amp; Y. S. Chen" authorityYear="2024" box="[517,746,266,293]" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="206" phylum="Tracheophyta" rank="species" species="wulongense">
<emphasis id="21CF2F73FFC5E15666278B1A9FE2FEAC" box="[517,746,266,293]" italics="true" pageId="5" pageNumber="206">Cirsium wulongense</emphasis>
</taxonomicName>
is currently only known from two localities (
<figureCitation id="8B80EFE4FFC5E15660C08B1B982FFEAC" box="[1250,1319,266,292]" captionStart="FIGURE 4" captionStartId="5.[136,229,1391,1413]" captionTargetBox="[196,1392,431,1361]" captionTargetId="figure-126@5.[190,1398,425,1367]" captionTargetPageId="5" captionText="FIGURE 4. Distribution of Cirsium wulongense (red circle)." pageId="5" pageNumber="206">Fig. 4</figureCitation>
). One is the
<typeStatus id="CC004DC3FFC5E15664AA8B3E9DB0FEC1" box="[136,184,303,329]" pageId="5" pageNumber="206">type</typeStatus>
locality, i.e. Xiannü Shan in Wulong district,
<collectingRegion id="D17F3D83FFC5E15666928B3F9E27FEC0" box="[688,815,302,328]" country="China" name="Chongqing" pageId="5" pageNumber="206">Chongqing</collectingRegion>
,
<collectingCountry id="6BACB3F1FFC5E15667188B3F9E89FEC0" box="[826,897,302,328]" name="China" pageId="5" pageNumber="206">China</collectingCountry>
. From the Plant Photo Bank of
<collectingCountry id="6BACB3F1FFC5E15660FB8B3F9814FEC0" box="[1241,1308,302,328]" name="China" pageId="5" pageNumber="206">China</collectingCountry>
(http://ppbc. iplant.cn/tu/2837783), we found that this species also occurs in Zunyi city in
<collectingRegion id="D17F3D83FFC5E15667D08B4399C8FEE4" box="[1010,1216,338,364]" country="China" name="Guizhou" pageId="5" pageNumber="206">Guizhou province</collectingRegion>
. These two localities are very close to each other. It grows in grassy areas at the forest edge at altitudes of 1600–1800 meters.
</paragraph>
<caption id="47C4A3E9FFC5E15664AA8F7E9FDBFA0E" box="[136,723,1391,1414]" pageId="5" pageNumber="206" startId="5.[136,229,1391,1413]" targetBox="[196,1392,431,1361]" targetPageId="5" targetType="figure">
<paragraph id="1304F361FFC5E15664AA8F7E9FDBFA0E" blockId="5.[136,723,1391,1414]" box="[136,723,1391,1414]" pageId="5" pageNumber="206">
<emphasis id="21CF2F73FFC5E15664AA8F7E9DF6FA0C" bold="true" box="[136,254,1391,1413]" pageId="5" pageNumber="206">FIGURE 4.</emphasis>
Distribution of
<taxonomicName id="D4BB88E2FFC5E15665B58F619F54FA0D" authorityName="Z. C. Jin &amp; Y. S. Chen" authorityYear="2024" box="[407,604,1392,1413]" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="206" phylum="Tracheophyta" rank="species" species="wulongense">
<emphasis id="21CF2F73FFC5E15665B58F619F54FA0D" box="[407,604,1392,1413]" italics="true" pageId="5" pageNumber="206">Cirsium wulongense</emphasis>
</taxonomicName>
(red circle).
</paragraph>
</caption>
<paragraph id="1304F361FFC5E156649F8FA59E95FA7A" blockId="5.[136,1452,1460,1594]" pageId="5" pageNumber="206">
<emphasis id="21CF2F73FFC5E156649F8FA59FCDFA46" bold="true" box="[189,709,1460,1486]" pageId="5" pageNumber="206">
Additional specimens examined:—
<collectingCountry id="6BACB3F1FFC5E156664A8FA59FCDFA46" box="[616,709,1460,1486]" name="China" pageId="5" pageNumber="206">CHINA</collectingCountry>
</emphasis>
,
<emphasis id="21CF2F73FFC5E15666FB8FA59E55FA46" bold="true" box="[729,861,1460,1486]" pageId="5" pageNumber="206">
<collectingRegion id="D17F3D83FFC5E15666FB8FA59E55FA46" box="[729,861,1460,1486]" country="China" name="Chongqing" pageId="5" pageNumber="206">Chongqing</collectingRegion>
</emphasis>
, Wulong district, Xiannü Shan, elev.
<quantity id="D4435E84FFC5E15661008FA59876FA46" box="[1314,1406,1460,1487]" metricMagnitude="3" metricUnit="m" metricValue="1.6509999999999998" pageId="5" pageNumber="206" unit="m" value="1651.0">1651 m</quantity>
,
<date id="6705D5A1FFC5E15661AD8FA59C13FA7A" pageId="5" pageNumber="206" value="2021-08-14">14 August 2021</date>
,
<emphasis id="21CF2F73FFC5E15665048FC89FD5FA7A" box="[294,733,1496,1522]" italics="true" pageId="5" pageNumber="206">F. Chen, X.B. Shi &amp; F.F. Xu XLS21-093</emphasis>
(CBNM, IBSC).
</paragraph>
<paragraph id="1304F361FFC5E156649F8FED9E6EF9B2" blockId="5.[136,1452,1460,1594]" pageId="5" pageNumber="206">
<emphasis id="21CF2F73FFC5E156649F8FED9C13F99E" bold="true" box="[189,283,1532,1558]" pageId="5" pageNumber="206">Note:—</emphasis>
Because its phyllaries lack marginal spinules, wings, and scarious appendage and its leaves abaxially lack spinules, the new species should belong to
<taxonomicName id="D4BB88E2FFC5E15666488C319E57F9B2" authority="sect. Cirsium" box="[618,863,1568,1594]" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="206" phylum="Tracheophyta" rank="section" section="Cirsium">
<emphasis id="21CF2F73FFC5E15666488C319FCCF9B2" box="[618,708,1568,1594]" italics="true" pageId="5" pageNumber="206">Cirsium</emphasis>
sect.
<emphasis id="21CF2F73FFC5E15667278C319E57F9B2" box="[773,863,1568,1594]" italics="true" pageId="5" pageNumber="206">Cirsium</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

227
data/A8/1D/87/A81D87B1710DFFD6FF06FE5EFBFA8A70.xml Normal file
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@ -0,0 +1,227 @@
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<mods:title id="5C02F2E40940FF68144F4D58D14060DE">A newly described Lomatogonium madhmaheshwariansis is a synonym of Swertia alpina</mods:title>
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<mods:namePart id="70F726920E7D0FD9760ACD5525CA3B35">Kumar, Aashish</mods:namePart>
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<mods:affiliation id="4A68BD00E18FD47166A0D191EA564264">Village and post Basera, Muzaffarnagar (Uttar Pradesh) - 251310</mods:affiliation>
<mods:nameIdentifier id="1423BEF4819749C370C6D63062513CFB" type="email">aashishdeshwal848@gmail.com</mods:nameIdentifier>
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<mods:namePart id="6E2FFF9525C5B72AE8046482D6B94197">Singh, Harsh</mods:namePart>
<mods:nameIdentifier id="0A7DCB8BEDE43E62ADB85E3CD94950FE" type="ORCID">0000-0002-9416-895X</mods:nameIdentifier>
<mods:affiliation id="4F32FC0904B765F3A0FE5E2FCF1B6A52">Department of Botany, Centre for Advanced Studies in Botany, North-Eastern Hill University, Shillong- 793022, Meghalaya, INDIA</mods:affiliation>
<mods:nameIdentifier id="07D2B5335A16A910A072A1A062C30019" type="email">harshchamlegi@gmail.com</mods:nameIdentifier>
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<mods:date id="EF06680F0734528DDA65CFB05B49716D">2024</mods:date>
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<treatment id="A81D87B1710DFFD6FF06FE5EFBFA8A70" LSID="urn:lsid:plazi:treatment:A81D87B1710DFFD6FF06FE5EFBFA8A70" httpUri="http://treatment.plazi.org/id/A81D87B1710DFFD6FF06FE5EFBFA8A70" lastPageNumber="56" pageId="1" pageNumber="56">
<subSubSection id="68AE652C710DFFD6FF06FE5EFC918E15" box="[136,839,500,527]" pageId="1" pageNumber="56" type="nomenclature">
<paragraph id="200B36A7710DFFD6FF06FE5EFC918E15" blockId="1.[136,839,500,527]" box="[136,839,500,527]" pageId="1" pageNumber="56">
<heading id="7B4381CB710DFFD6FF06FE5EFC918E15" box="[136,839,500,527]" fontSize="11" level="2" pageId="1" pageNumber="56" reason="3">
<taxonomicName id="E7B44D24710DFFD6FF06FE5EFD268E15" ID-CoL="53JNC" authority="U. C. Bhattach. &amp; S. Agrawal (1981: 47)" authorityName="U. C. Bhattach. &amp; S. Agrawal" authorityPageNumber="47" authorityYear="1981" box="[136,752,500,527]" class="Magnoliopsida" family="Gentianaceae" genus="Swertia" kingdom="Plantae" order="Gentianales" pageId="1" pageNumber="56" phylum="Tracheophyta" rank="species" species="alpina">
<emphasis id="12C0EAB5710DFFD6FF06FE5EFEF98E14" bold="true" box="[136,303,500,527]" italics="true" pageId="1" pageNumber="56">Swertia alpina</emphasis>
U.C.Bhattach. &amp; S.Agrawal (1981: 47)
</taxonomicName>
.
<figureCitation id="B88F2A22710DFFD6FD75FE5EFC918E15" box="[763,839,500,526]" captionStart="FIGURE 1" captionStartId="2.[136,229,1928,1950]" captionTargetBox="[151,1435,187,1900]" captionTargetId="figure-20@2.[151,1435,187,1900]" captionTargetPageId="2" captionText="FIGURE 1. Swertia alpina U.C.Bhattach. &amp; S.Agarwal: A. habitat and habit; B. leaves pattern; C., &amp; D. flower." figureDoi="http://doi.org/10.5281/zenodo.14516018" httpUri="https://zenodo.org/record/14516018/files/figure.png" pageId="1" pageNumber="56">Figs. 1</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="68AE652C710DFFD6FF06FDEAFC838EFE" pageId="1" pageNumber="56" type="materials_examined">
<paragraph id="200B36A7710DFFD6FF06FDEAFA7D8E4E" blockId="1.[136,1452,575,742]" box="[136,1451,575,598]" pageId="1" pageNumber="56">
<typeStatus id="FF0F8805710DFFD6FF06FDEAFF6D8E4D" box="[136,187,576,598]" pageId="1" pageNumber="56">Type</typeStatus>
:—
<collectingCountry id="58A37637710DFFD6FF59FDEAFEDC8E4D" box="[215,266,576,598]" name="India" pageId="1" pageNumber="56">India</collectingCountry>
,
<collectingRegion id="E270F845710DFFD6FE9AFDEAFE5B8E4D" box="[276,397,576,598]" country="India" name="Uttarakhand" pageId="1" pageNumber="56">Uttarakhand</collectingRegion>
, Kedarnath glacial valley, Chamoli Garhwal, North West Himalaya,
<quantity id="E74C9B42710DFFD6FBACFDEAFBBD8E4E" box="[1058,1131,576,597]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="56" unit="m" value="4000.0">4000 m</quantity>
,
<date id="540A1067710DFFD6FBF3FDEAFAD98E4D" box="[1149,1295,575,598]" pageId="1" pageNumber="56" value="1958-09-27">
27
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Sept. 1958
</date>
,
<emphasis id="12C0EAB5710DFFD6FA97FDEAFA7D8E4E" box="[1305,1451,576,597]" italics="true" pageId="1" pageNumber="56">
<collectionCode id="46A5AE62710DFFD6FA97FDEAFAE78E4E" box="[1305,1329,576,597]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="1" pageNumber="56" type="Herbarium">M</collectionCode>
.
<collectionCode id="46A5AE62710DFFD6FABBFDEAFA908E4E" box="[1333,1350,576,597]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="1" pageNumber="56" type="Herbarium">A</collectionCode>
. Rau 8701
</emphasis>
</paragraph>
<paragraph id="200B36A7710DFFD6FF33FDCEFA7A8EDA" blockId="1.[136,1452,575,742]" pageId="1" pageNumber="56">
(
<collectionCode id="46A5AE62710DFFD6FF4BFDCEFF2E8E62" box="[197,248,612,633]" country="India" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12696" lsid="urn:lsid:biocol.org:col:12696" name="Botanical Survey of India, Northern Regional Centre" pageId="1" pageNumber="56" type="Herbarium">BSD</collectionCode>
).
<emphasis id="12C0EAB5710DFFD6FF06FD22FDD68E86" box="[136,512,648,669]" italics="true" pageId="1" pageNumber="56">
=Lomatognium
<taxonomicName id="E7B44D24710DFFD6FEAAFD22FDD68E86" authorityName="Sunit Singh, Bikarma Singh, Sajan Thakur &amp; Jagat Prakash Mehta" authorityYear="2023" box="[292,512,648,669]" class="Magnoliopsida" family="Gentianaceae" genus="Lomatogonium" kingdom="Plantae" order="Gentianales" pageId="1" pageNumber="56" phylum="Tracheophyta" rank="species" species="madhmaheshwariansis">madhmaheshwariansis</taxonomicName>
</emphasis>
(2023: 5)
<emphasis id="12C0EAB5710DFFD6FDEAFD22FD638E86" bold="true" box="[612,693,648,669]" italics="true" pageId="1" pageNumber="56">syn.nov.</emphasis>
<typeStatus id="FF0F8805710DFFD6FF06FD06FF6D8ED9" box="[136,187,684,706]" pageId="1" pageNumber="56">Type</typeStatus>
:—
<collectingCountry id="58A37637710DFFD6FF59FD06FEDC8ED9" box="[215,266,684,706]" name="India" pageId="1" pageNumber="56">India</collectingCountry>
, Himalayan Region,
<collectingRegion id="E270F845710DFFD6FE58FD06FD998ED9" box="[470,591,684,706]" country="India" name="Uttarakhand" pageId="1" pageNumber="56">Uttarakhand</collectingRegion>
, Rudraprayag district, Budha-Madhmaheshwar valley,
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,
<geoCoordinate id="45805060710DFFD6FA82FD00FA718EDA" box="[1292,1447,682,705]" degrees="79" direction="east" minutes="12" orientation="longitude" pageId="1" pageNumber="56" precision="1" seconds="40.183" value="79.21116">79°12′40.183″E</geoCoordinate>
,
</paragraph>
<paragraph id="200B36A7710DFFD6FF33FD7AFC838EFE" blockId="1.[136,1452,575,742]" box="[189,853,719,742]" pageId="1" pageNumber="56">
<quantity id="E74C9B42710DFFD6FF33FD7AFED08EFE" box="[189,262,720,742]" metricMagnitude="3" metricUnit="m" metricValue="3.485" pageId="1" pageNumber="56" unit="m" value="3485.0">3485 m</quantity>
<emphasis id="12C0EAB5710DFFD6FE82FD7AFEEC8EFE" box="[268,314,720,741]" italics="true" pageId="1" pageNumber="56">a.s.l.</emphasis>
,
<date id="540A1067710DFFD6FEC8FD7AFE0E8EFE" box="[326,472,719,742]" pageId="1" pageNumber="56" value="2021-09-15">
15
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Sept. 2021
</date>
,
<emphasis id="12C0EAB5710DFFD6FE6CFD7AFD748EFE" box="[482,674,720,741]" italics="true" pageId="1" pageNumber="56">Singh et al. 334 810</emphasis>
(
<typeStatus id="FF0F8805710DFFD6FD21FD7AFCDE8EFD" box="[687,776,720,742]" pageId="1" pageNumber="56" type="holotype">holotype</typeStatus>
:
<collectionCode id="46A5AE62710DFFD6FC9FFD7AFC9C8EFE" box="[785,842,720,741]" country="India" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15400" lsid="urn:lsid:biocol.org:col:15400" name="National Botanical Research Institute" pageId="1" pageNumber="56" type="Herbarium">LWG</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="68AE652C710DFFD6FF06FCBEFEEF88A1" pageId="1" pageNumber="56" type="description">
<paragraph id="200B36A7710DFFD6FF06FCBEFC458869" blockId="1.[136,1452,788,1643]" pageId="1" pageNumber="56">
<emphasis id="12C0EAB5710DFFD6FF06FCBEFEC48F35" bold="true" box="[136,274,788,814]" pageId="1" pageNumber="56">Description</emphasis>
:—Annual,
<quantity id="E74C9B42710DFFD6FE16FCBEFDDF8F34" box="[408,521,788,815]" metricMagnitude="-1" metricUnit="m" metricValue="3.25" metricValueMax="5.0" metricValueMin="1.5" pageId="1" pageNumber="56" unit="cm" value="32.5" valueMax="50.0" valueMin="15.0">15–50 cm</quantity>
long, herb. Stem erect, tetragonal, glabrous. Cauline leaves opposite, exstipulate, sessile, glabrous; middle ones on main stem, ovate-cordate to ovate-lanceolate, apices acute. Inflorescence axillary to terminal cymes of 2–6 clusters, lax in fruiting; penducles
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long, strict; bracts linear, sessile, acute; flowers purple in buds,
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long, pentamerous, pedicellate; pedicels
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long, terminal one longest; calyx lobed, 5, glabrous, persistent; lobes 4–6 ×
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, linear-lanceolate, acuminate, margins serrulate; corolla rotate, persistent longer than the calyx; lobes 5, 6–8 ×
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, ovate-lanceolate, acute; glands 2, on either side of the base of each lobe, minute, oblong-ovoid with hairy or sub-glabrous margins. Stamens 5, epipetalous; filaments filiform,
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long; anthers bluish, oblong, c.
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. Ovary oblong-elliptic, sessile, 4–5 ×
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; style reduced; stigma 2 or bifid, hemispheric or reniform, papillose. Capsule oblong, sessile, 0.8–1.2 ×
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, subincluded when ripe. Seeds numerous, brownish-black, sub-globose, c. 1 ×
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; testa alveolate.
</paragraph>
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<emphasis id="12C0EAB5710DFFD6FF33FBD6FE07888D" bold="true" box="[189,465,1148,1174]" pageId="1" pageNumber="56">Flowering and Fruiting</emphasis>
:—Flowering in the month of August–September while fruiting in the month of September last–November.
</paragraph>
</subSubSection>
<subSubSection id="68AE652C710DFFD6FF33FB6EFBFA8A70" pageId="1" pageNumber="56" type="distribution">
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<emphasis id="12C0EAB5710DFFD6FF33FB6EFE9988C5" bold="true" box="[189,335,1220,1246]" pageId="1" pageNumber="56">Distribution</emphasis>
:—So far, the species is known in
<collectingRegion id="E270F845710DFFD6FD41FB6EFC8F88C5" box="[719,857,1220,1246]" country="India" name="Uttarakhand" pageId="1" pageNumber="56">Uttarakhand</collectingRegion>
at an elevation of
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.
</paragraph>
<paragraph id="200B36A7710DFFD6FF33FB42FF6B893D" blockId="1.[136,1452,788,1643]" pageId="1" pageNumber="56">
<emphasis id="12C0EAB5710DFFD6FF33FB42FECF8919" bold="true" box="[189,281,1256,1282]" pageId="1" pageNumber="56">Habitat</emphasis>
:—
<taxonomicName id="E7B44D24710DFFD6FEB3FB42FE358919" authorityName="U. C. Bhattach. &amp; S. Agrawal" authorityYear="1981" box="[317,483,1256,1282]" class="Magnoliopsida" family="Gentianaceae" genus="Swertia" kingdom="Plantae" order="Gentianales" pageId="1" pageNumber="56" phylum="Tracheophyta" rank="species" species="alpina">
<emphasis id="12C0EAB5710DFFD6FEB3FB42FE358919" box="[317,483,1256,1282]" italics="true" pageId="1" pageNumber="56">Swertia alpina</emphasis>
</taxonomicName>
is growing in alpine pastures of mountain open slopes at elevations of
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a.s.l.
</paragraph>
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<emphasis id="12C0EAB5710DFFD6FF33FA9AFED68951" bold="true" box="[189,256,1328,1354]" pageId="1" pageNumber="56">Notes</emphasis>
:—
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<emphasis id="12C0EAB5710DFFD6FEAAFA9AFE1C8951" box="[292,458,1328,1354]" italics="true" pageId="1" pageNumber="56">Swertia alpina</emphasis>
</taxonomicName>
is well known species found in alpine pastures of an altitude of
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<emphasis id="12C0EAB5710DFFD6FAD8FA9AFA5D8951" box="[1366,1419,1328,1354]" italics="true" pageId="1" pageNumber="56">a.s.l.</emphasis>
of the Himalayas. In 1981, U.C.Bhattacharya and Sunita Agrawal was described it in the Journal Bulletin Botanical Society, Bengal based on earlier collected plant materials of M.A.Rau (collection number 8701) from glacial valley of Kedarnath at an altitude of
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in 1958. The species epithet
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‘
<emphasis id="12C0EAB5710DFFD6FCDEFA36FC4E89AD" box="[848,920,1436,1462]" italics="true" pageId="1" pageNumber="56">alpina</emphasis>
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’ meaning alpine was named after its occurrence in alpine pastures of
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. Bhattacharya &amp; Agrawal compared
<taxonomicName id="E7B44D24710DFFD6FC28FA6AFBDA89C1" authorityName="U. C. Bhattach. &amp; S. Agrawal" authorityYear="1981" box="[934,1036,1472,1498]" class="Magnoliopsida" family="Gentianaceae" genus="Swertia" kingdom="Plantae" order="Gentianales" pageId="1" pageNumber="56" phylum="Tracheophyta" rank="species" species="alpina">
<emphasis id="12C0EAB5710DFFD6FC28FA6AFBDA89C1" box="[934,1036,1472,1498]" italics="true" pageId="1" pageNumber="56">S. alpina</emphasis>
</taxonomicName>
with
<taxonomicName id="E7B44D24710DFFD6FBC3FA6AFF7B89E5" authority="Edgeworth (1846: 86)" authorityName="Edgeworth" authorityPageNumber="86" authorityYear="1846" class="Magnoliopsida" family="Gentianaceae" genus="Swertia" kingdom="Plantae" order="Gentianales" pageId="1" pageNumber="56" phylum="Tracheophyta" rank="species" species="tetragona">
<emphasis id="12C0EAB5710DFFD6FBC3FA6AFB0189C1" box="[1101,1239,1472,1498]" italics="true" pageId="1" pageNumber="56">S. tetragona</emphasis>
<bibRefCitation id="44254B56710DFFD6FB51FA6AFF7B89E5" author="Edgeworth, M." pageId="1" pageNumber="56" refId="ref2032" refString="Edgeworth, M. (1846) London Mr. Edgeworth on some unpublished Species Plants from North - Western India. Transactions of the Linnean Society of London 20 (1): 86. https: // doi. org / 10.1111 / j. 1096 - 3642.1846. tb 00410. x" type="journal volume" year="1846">Edgeworth (1846: 86)</bibRefCitation>
</taxonomicName>
and is differentiated due to its ovate-cordate to ovate-lanceolate, sessile, margins not revolute, 3–5 nerved. (
<emphasis id="12C0EAB5710DFFD6FA03FA4FFA7D89E5" box="[1421,1451,1509,1534]" italics="true" pageId="1" pageNumber="56">vs.</emphasis>
linear lanceolate, subsessile, margins slightly revolute, 1–3 nerved.), seeds brownish-black
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in size (
<emphasis id="12C0EAB5710DFFD6FAC9F9A3FAB38A39" box="[1351,1381,1545,1570]" italics="true" pageId="1" pageNumber="56">vs.</emphasis>
seeds reddish-brown, less than
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in size.
<taxonomicName id="E7B44D24710DFFD6FDC8F987FC2F8A5D" authorityName="Sunit Singh, Bikarma Singh, Sajan Thakur &amp; Jagat Prakash Mehta" authorityYear="2023" box="[582,1017,1580,1606]" class="Magnoliopsida" family="Gentianaceae" genus="Lomatogonium" kingdom="Plantae" order="Gentianales" pageId="1" pageNumber="56" phylum="Tracheophyta" rank="species" species="madhmaheshwariansis">
<emphasis id="12C0EAB5710DFFD6FDC8F987FC2F8A5D" box="[582,1017,1580,1606]" italics="true" pageId="1" pageNumber="56">Lomatogonium madhmaheshwariansis</emphasis>
</taxonomicName>
which is newly described species was misidentified due to the fact that it has all the characters conspecific with
<taxonomicName id="E7B44D24710DFFD6FC4FF9FAFBF38A71" authorityName="U. C. Bhattach. &amp; S. Agrawal" authorityYear="1981" box="[961,1061,1616,1642]" class="Magnoliopsida" family="Gentianaceae" genus="Swertia" kingdom="Plantae" order="Gentianales" pageId="1" pageNumber="56" phylum="Tracheophyta" rank="species" species="alpina">
<emphasis id="12C0EAB5710DFFD6FC4FF9FAFBF38A71" box="[961,1061,1616,1642]" italics="true" pageId="1" pageNumber="56">S. alpina</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

262
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<emphasis id="7AAF13127D6DF124FF01F98402A8F9FC" bold="true" box="[136,470,1552,1579]" italics="true" pageId="1" pageNumber="50">Pseudostaurosira katharinae</emphasis>
Van de Vijver &amp; C.E.Wetzel
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</emphasis>
(
<figureCitation id="D0E0D3857D6DF124FC1EF9840774F9FD" box="[919,1034,1552,1578]" captionStart="FIGURES 1–25" captionStartId="2.[136,248,1848,1870]" captionTargetBox="[193,1394,187,1821]" captionTargetId="figure-25@2.[193,1394,187,1821]" captionTargetPageId="2" captionText="FIGURES 1–25. Pseudostaurosira katharinae Van de Vijver &amp; C.E.Wetzel sp. nov. LM and SEM taken from the holotype sample (BR- 4850, Bosbeek, Maaseik, Belgium). 1–2. LM pictures of two connected frustules in girdle view. 3–17. LM pictures of valves in valve face view in decreasing length. 18–19 SEM external view of a complete frustule in girdle view showing the girdle structure, the large mantle with the mantle plaques and the apical depression (white arrow). 20–22. SEM external view three valves showing the large sternum, the spine structure, the large apical pore fields and the areolae decreasing in diameter towards the sternum. Note the circular series of small siliceous papillae around the areolae and the spines. 23. SEM external detail of the valve apex with the apical pore field. 24. SEM external detail of the areolae and the spines. 25. SEM internal view of a complete valve. Scale bars: 10 μm (1–17); 1 μm (18–25)." figureDoi="http://doi.org/10.5281/zenodo.14515991" httpUri="https://zenodo.org/record/14515991/files/figure.png" pageId="1" pageNumber="50">Figs 1–17</figureCitation>
LM, 18–27 SEM)
</emphasis>
</heading>
</paragraph>
<paragraph id="4864CF007D6DF126FF01F9CC02E6FED6" blockId="1.[136,1452,1624,2083]" lastBlockId="3.[136,1452,158,1038]" lastPageId="3" lastPageNumber="52" pageId="1" pageNumber="50">
Frustules rectangular in girdle view, solitary or in pairs (
<figureCitation id="D0E0D3857D6DF124FC87F9CC0008F9A5" box="[782,886,1624,1650]" captionStart="FIGURES 1–25" captionStartId="2.[136,248,1848,1870]" captionTargetBox="[193,1394,187,1821]" captionTargetId="figure-25@2.[193,1394,187,1821]" captionTargetPageId="2" captionText="FIGURES 1–25. Pseudostaurosira katharinae Van de Vijver &amp; C.E.Wetzel sp. nov. LM and SEM taken from the holotype sample (BR- 4850, Bosbeek, Maaseik, Belgium). 1–2. LM pictures of two connected frustules in girdle view. 3–17. LM pictures of valves in valve face view in decreasing length. 18–19 SEM external view of a complete frustule in girdle view showing the girdle structure, the large mantle with the mantle plaques and the apical depression (white arrow). 20–22. SEM external view three valves showing the large sternum, the spine structure, the large apical pore fields and the areolae decreasing in diameter towards the sternum. Note the circular series of small siliceous papillae around the areolae and the spines. 23. SEM external detail of the valve apex with the apical pore field. 24. SEM external detail of the areolae and the spines. 25. SEM internal view of a complete valve. Scale bars: 10 μm (1–17); 1 μm (18–25)." figureDoi="http://doi.org/10.5281/zenodo.14515991" httpUri="https://zenodo.org/record/14515991/files/figure.png" pageId="1" pageNumber="50">Figs 1–2</figureCitation>
). Ribbon-like colonies so far not observed. Valve mantle broad, with a large hyaline abvalvar part. Mantle plaques small but distinctly present on the mantle edge (
<figureCitation id="D0E0D3857D6DF124FAF3F9E803ACF96D" captionStart="FIGURES 1–25" captionStartId="2.[136,248,1848,1870]" captionTargetBox="[193,1394,187,1821]" captionTargetId="figure-25@2.[193,1394,187,1821]" captionTargetPageId="2" captionText="FIGURES 1–25. Pseudostaurosira katharinae Van de Vijver &amp; C.E.Wetzel sp. nov. LM and SEM taken from the holotype sample (BR- 4850, Bosbeek, Maaseik, Belgium). 1–2. LM pictures of two connected frustules in girdle view. 3–17. LM pictures of valves in valve face view in decreasing length. 18–19 SEM external view of a complete frustule in girdle view showing the girdle structure, the large mantle with the mantle plaques and the apical depression (white arrow). 20–22. SEM external view three valves showing the large sternum, the spine structure, the large apical pore fields and the areolae decreasing in diameter towards the sternum. Note the circular series of small siliceous papillae around the areolae and the spines. 23. SEM external detail of the valve apex with the apical pore field. 24. SEM external detail of the areolae and the spines. 25. SEM internal view of a complete valve. Scale bars: 10 μm (1–17); 1 μm (18–25)." figureDoi="http://doi.org/10.5281/zenodo.14515991" httpUri="https://zenodo.org/record/14515991/files/figure.png" pageId="1" pageNumber="50">Figs 18–19</figureCitation>
). Girdle composed of several plain, open, ligulate copulae, each bearing a thin siliceous ridge (
<figureCitation id="D0E0D3857D6DF124FA93F93406E1F96D" box="[1306,1439,1696,1722]" captionStart="FIGURES 1–25" captionStartId="2.[136,248,1848,1870]" captionTargetBox="[193,1394,187,1821]" captionTargetId="figure-25@2.[193,1394,187,1821]" captionTargetPageId="2" captionText="FIGURES 1–25. Pseudostaurosira katharinae Van de Vijver &amp; C.E.Wetzel sp. nov. LM and SEM taken from the holotype sample (BR- 4850, Bosbeek, Maaseik, Belgium). 1–2. LM pictures of two connected frustules in girdle view. 3–17. LM pictures of valves in valve face view in decreasing length. 18–19 SEM external view of a complete frustule in girdle view showing the girdle structure, the large mantle with the mantle plaques and the apical depression (white arrow). 20–22. SEM external view three valves showing the large sternum, the spine structure, the large apical pore fields and the areolae decreasing in diameter towards the sternum. Note the circular series of small siliceous papillae around the areolae and the spines. 23. SEM external detail of the valve apex with the apical pore field. 24. SEM external detail of the areolae and the spines. 25. SEM internal view of a complete valve. Scale bars: 10 μm (1–17); 1 μm (18–25)." figureDoi="http://doi.org/10.5281/zenodo.14515991" httpUri="https://zenodo.org/record/14515991/files/figure.png" pageId="1" pageNumber="50">Figs 18–19</figureCitation>
). Valvocopula largest, plain (
<figureCitation id="D0E0D3857D6DF124FE33F9500143F909" box="[442,573,1732,1758]" captionStart="FIGURES 1–25" captionStartId="2.[136,248,1848,1870]" captionTargetBox="[193,1394,187,1821]" captionTargetId="figure-25@2.[193,1394,187,1821]" captionTargetPageId="2" captionText="FIGURES 1–25. Pseudostaurosira katharinae Van de Vijver &amp; C.E.Wetzel sp. nov. LM and SEM taken from the holotype sample (BR- 4850, Bosbeek, Maaseik, Belgium). 1–2. LM pictures of two connected frustules in girdle view. 3–17. LM pictures of valves in valve face view in decreasing length. 18–19 SEM external view of a complete frustule in girdle view showing the girdle structure, the large mantle with the mantle plaques and the apical depression (white arrow). 20–22. SEM external view three valves showing the large sternum, the spine structure, the large apical pore fields and the areolae decreasing in diameter towards the sternum. Note the circular series of small siliceous papillae around the areolae and the spines. 23. SEM external detail of the valve apex with the apical pore field. 24. SEM external detail of the areolae and the spines. 25. SEM internal view of a complete valve. Scale bars: 10 μm (1–17); 1 μm (18–25)." figureDoi="http://doi.org/10.5281/zenodo.14515991" httpUri="https://zenodo.org/record/14515991/files/figure.png" pageId="1" pageNumber="50">Figs 18–19</figureCitation>
). Most copulae fracted (
<figureCitation id="D0E0D3857D6DF124FCC5F95000B5F909" box="[844,971,1732,1758]" captionStart="FIGURES 1–25" captionStartId="2.[136,248,1848,1870]" captionTargetBox="[193,1394,187,1821]" captionTargetId="figure-25@2.[193,1394,187,1821]" captionTargetPageId="2" captionText="FIGURES 1–25. Pseudostaurosira katharinae Van de Vijver &amp; C.E.Wetzel sp. nov. LM and SEM taken from the holotype sample (BR- 4850, Bosbeek, Maaseik, Belgium). 1–2. LM pictures of two connected frustules in girdle view. 3–17. LM pictures of valves in valve face view in decreasing length. 18–19 SEM external view of a complete frustule in girdle view showing the girdle structure, the large mantle with the mantle plaques and the apical depression (white arrow). 20–22. SEM external view three valves showing the large sternum, the spine structure, the large apical pore fields and the areolae decreasing in diameter towards the sternum. Note the circular series of small siliceous papillae around the areolae and the spines. 23. SEM external detail of the valve apex with the apical pore field. 24. SEM external detail of the areolae and the spines. 25. SEM internal view of a complete valve. Scale bars: 10 μm (1–17); 1 μm (18–25)." figureDoi="http://doi.org/10.5281/zenodo.14515991" httpUri="https://zenodo.org/record/14515991/files/figure.png" pageId="1" pageNumber="50">Figs 18, 19</figureCitation>
&amp;
<figureCitation id="D0E0D3857D6DF124FC66F9500773F909" box="[1007,1037,1732,1758]" captionStart="FIGURES 26–27" captionStartId="3.[136,248,1691,1713]" captionTargetBox="[151,1436,1062,1667]" captionTargetId="figure-456@3.[151,1436,1062,1667]" captionTargetPageId="3" captionText="FIGURES 26–27. Pseudostaurosira katharinae Van de Vijver &amp; C.E.Wetzel sp. nov. Specimens from France (Mare à Goriaux). SEM external view of a complete frustules in girdle view. Fig. 26: Detail showing the fracted girdle bands (white arrow); mantle plates of reduced size. Fig. 27: Open valvocopulae with fimbriate girdle elements. Serrate spines. Scale bars: 4 μm (26); 5 μm (27)." figureDoi="http://doi.org/10.5281/zenodo.14515993" httpUri="https://zenodo.org/record/14515993/files/figure.png" pageId="1" pageNumber="50">26</figureCitation>
, arrows). Valves lanceolate to linearlanceolate in larger specimens, becoming more elliptic-lanceolate to elliptic in smallest valves (
<figureCitation id="D0E0D3857D6DF124FB54F97C062BF8D5" box="[1245,1365,1768,1794]" captionStart="FIGURES 1–25" captionStartId="2.[136,248,1848,1870]" captionTargetBox="[193,1394,187,1821]" captionTargetId="figure-25@2.[193,1394,187,1821]" captionTargetPageId="2" captionText="FIGURES 1–25. Pseudostaurosira katharinae Van de Vijver &amp; C.E.Wetzel sp. nov. LM and SEM taken from the holotype sample (BR- 4850, Bosbeek, Maaseik, Belgium). 1–2. LM pictures of two connected frustules in girdle view. 3–17. LM pictures of valves in valve face view in decreasing length. 18–19 SEM external view of a complete frustule in girdle view showing the girdle structure, the large mantle with the mantle plaques and the apical depression (white arrow). 20–22. SEM external view three valves showing the large sternum, the spine structure, the large apical pore fields and the areolae decreasing in diameter towards the sternum. Note the circular series of small siliceous papillae around the areolae and the spines. 23. SEM external detail of the valve apex with the apical pore field. 24. SEM external detail of the areolae and the spines. 25. SEM internal view of a complete valve. Scale bars: 10 μm (1–17); 1 μm (18–25)." figureDoi="http://doi.org/10.5281/zenodo.14515991" httpUri="https://zenodo.org/record/14515991/files/figure.png" pageId="1" pageNumber="50">Figs 3–17</figureCitation>
). Valve margins convex to weakly convex, not linear. Apices broadly to cuneately rounded, rarely weakly protracted. Usually, one apex slightly depressed (
<figureCitation id="D0E0D3857D6DF124FE50F8A40151F89D" box="[473,559,1840,1866]" captionStart="FIGURES 1–25" captionStartId="2.[136,248,1848,1870]" captionTargetBox="[193,1394,187,1821]" captionTargetId="figure-25@2.[193,1394,187,1821]" captionTargetPageId="2" captionText="FIGURES 1–25. Pseudostaurosira katharinae Van de Vijver &amp; C.E.Wetzel sp. nov. LM and SEM taken from the holotype sample (BR- 4850, Bosbeek, Maaseik, Belgium). 1–2. LM pictures of two connected frustules in girdle view. 3–17. LM pictures of valves in valve face view in decreasing length. 18–19 SEM external view of a complete frustule in girdle view showing the girdle structure, the large mantle with the mantle plaques and the apical depression (white arrow). 20–22. SEM external view three valves showing the large sternum, the spine structure, the large apical pore fields and the areolae decreasing in diameter towards the sternum. Note the circular series of small siliceous papillae around the areolae and the spines. 23. SEM external detail of the valve apex with the apical pore field. 24. SEM external detail of the areolae and the spines. 25. SEM internal view of a complete valve. Scale bars: 10 μm (1–17); 1 μm (18–25)." figureDoi="http://doi.org/10.5281/zenodo.14515991" httpUri="https://zenodo.org/record/14515991/files/figure.png" pageId="1" pageNumber="50">Fig. 19</figureCitation>
, white arrow). Valve dimensions (n=30): valve length 9–15 μm, valve width 3.5–4.5 μm. Sternum broad, lanceolate, max width 1.2–1.8 µm. Occasionally sternum narrow but abruptly widening forming a rounded central area (
<figureCitation id="D0E0D3857D6DF124FE70F8EC0100F845" box="[505,638,1912,1938]" captionStart="FIGURES 1–25" captionStartId="2.[136,248,1848,1870]" captionTargetBox="[193,1394,187,1821]" captionTargetId="figure-25@2.[193,1394,187,1821]" captionTargetPageId="2" captionText="FIGURES 1–25. Pseudostaurosira katharinae Van de Vijver &amp; C.E.Wetzel sp. nov. LM and SEM taken from the holotype sample (BR- 4850, Bosbeek, Maaseik, Belgium). 1–2. LM pictures of two connected frustules in girdle view. 3–17. LM pictures of valves in valve face view in decreasing length. 18–19 SEM external view of a complete frustule in girdle view showing the girdle structure, the large mantle with the mantle plaques and the apical depression (white arrow). 20–22. SEM external view three valves showing the large sternum, the spine structure, the large apical pore fields and the areolae decreasing in diameter towards the sternum. Note the circular series of small siliceous papillae around the areolae and the spines. 23. SEM external detail of the valve apex with the apical pore field. 24. SEM external detail of the areolae and the spines. 25. SEM internal view of a complete valve. Scale bars: 10 μm (1–17); 1 μm (18–25)." figureDoi="http://doi.org/10.5281/zenodo.14515991" httpUri="https://zenodo.org/record/14515991/files/figure.png" pageId="1" pageNumber="50">Figs 11, 13</figureCitation>
). Striae uniseriate, weakly radiate, irregularly shortened throughout the entire valve length,
<quantity id="8F2362E57D6DF124FEE0F80802B3F861" box="[361,461,1948,1974]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" metricValueMax="4.064" metricValueMin="3.556" pageId="1" pageNumber="50" unit="in" value="15.0" valueMax="16.0" valueMin="14.0">14–16 in</quantity>
10 μm, composed of 1–3 areolae on valve face (
<figureCitation id="D0E0D3857D6DF124FC7EF8080705F861" box="[1015,1147,1948,1974]" captionStart="FIGURES 1–25" captionStartId="2.[136,248,1848,1870]" captionTargetBox="[193,1394,187,1821]" captionTargetId="figure-25@2.[193,1394,187,1821]" captionTargetPageId="2" captionText="FIGURES 1–25. Pseudostaurosira katharinae Van de Vijver &amp; C.E.Wetzel sp. nov. LM and SEM taken from the holotype sample (BR- 4850, Bosbeek, Maaseik, Belgium). 1–2. LM pictures of two connected frustules in girdle view. 3–17. LM pictures of valves in valve face view in decreasing length. 18–19 SEM external view of a complete frustule in girdle view showing the girdle structure, the large mantle with the mantle plaques and the apical depression (white arrow). 20–22. SEM external view three valves showing the large sternum, the spine structure, the large apical pore fields and the areolae decreasing in diameter towards the sternum. Note the circular series of small siliceous papillae around the areolae and the spines. 23. SEM external detail of the valve apex with the apical pore field. 24. SEM external detail of the areolae and the spines. 25. SEM internal view of a complete valve. Scale bars: 10 μm (1–17); 1 μm (18–25)." figureDoi="http://doi.org/10.5281/zenodo.14515991" httpUri="https://zenodo.org/record/14515991/files/figure.png" pageId="1" pageNumber="50">Figs 20–22</figureCitation>
), and 1–3 areolae on valve mantle (
<figureCitation id="D0E0D3857D6DF124FF61F85402E3F80D" box="[232,413,1984,2010]" captionStart="FIGURES 1–25" captionStartId="2.[136,248,1848,1870]" captionTargetBox="[193,1394,187,1821]" captionTargetId="figure-25@2.[193,1394,187,1821]" captionTargetPageId="2" captionText="FIGURES 1–25. Pseudostaurosira katharinae Van de Vijver &amp; C.E.Wetzel sp. nov. LM and SEM taken from the holotype sample (BR- 4850, Bosbeek, Maaseik, Belgium). 1–2. LM pictures of two connected frustules in girdle view. 3–17. LM pictures of valves in valve face view in decreasing length. 18–19 SEM external view of a complete frustule in girdle view showing the girdle structure, the large mantle with the mantle plaques and the apical depression (white arrow). 20–22. SEM external view three valves showing the large sternum, the spine structure, the large apical pore fields and the areolae decreasing in diameter towards the sternum. Note the circular series of small siliceous papillae around the areolae and the spines. 23. SEM external detail of the valve apex with the apical pore field. 24. SEM external detail of the areolae and the spines. 25. SEM internal view of a complete valve. Scale bars: 10 μm (1–17); 1 μm (18–25)." figureDoi="http://doi.org/10.5281/zenodo.14515991" httpUri="https://zenodo.org/record/14515991/files/figure.png" pageId="1" pageNumber="50">Figs 18–19, 21</figureCitation>
). Areolae at valve face/mantle junction markedly larger, decreasing in diameter towards sternum (
<figureCitation id="D0E0D3857D6DF124FF79F870020CF829" box="[240,370,2020,2046]" captionStart="FIGURES 1–25" captionStartId="2.[136,248,1848,1870]" captionTargetBox="[193,1394,187,1821]" captionTargetId="figure-25@2.[193,1394,187,1821]" captionTargetPageId="2" captionText="FIGURES 1–25. Pseudostaurosira katharinae Van de Vijver &amp; C.E.Wetzel sp. nov. LM and SEM taken from the holotype sample (BR- 4850, Bosbeek, Maaseik, Belgium). 1–2. LM pictures of two connected frustules in girdle view. 3–17. LM pictures of valves in valve face view in decreasing length. 18–19 SEM external view of a complete frustule in girdle view showing the girdle structure, the large mantle with the mantle plaques and the apical depression (white arrow). 20–22. SEM external view three valves showing the large sternum, the spine structure, the large apical pore fields and the areolae decreasing in diameter towards the sternum. Note the circular series of small siliceous papillae around the areolae and the spines. 23. SEM external detail of the valve apex with the apical pore field. 24. SEM external detail of the areolae and the spines. 25. SEM internal view of a complete valve. Scale bars: 10 μm (1–17); 1 μm (18–25)." figureDoi="http://doi.org/10.5281/zenodo.14515991" httpUri="https://zenodo.org/record/14515991/files/figure.png" pageId="1" pageNumber="50">Figs 20–22</figureCitation>
). Mantle areolae separated from those on valve face by relatively long spines (
<figureCitation id="D0E0D3857D6DF124FB46F8700631F828" box="[1231,1359,2020,2047]" captionStart="FIGURES 1–25" captionStartId="2.[136,248,1848,1870]" captionTargetBox="[193,1394,187,1821]" captionTargetId="figure-25@2.[193,1394,187,1821]" captionTargetPageId="2" captionText="FIGURES 1–25. Pseudostaurosira katharinae Van de Vijver &amp; C.E.Wetzel sp. nov. LM and SEM taken from the holotype sample (BR- 4850, Bosbeek, Maaseik, Belgium). 1–2. LM pictures of two connected frustules in girdle view. 3–17. LM pictures of valves in valve face view in decreasing length. 18–19 SEM external view of a complete frustule in girdle view showing the girdle structure, the large mantle with the mantle plaques and the apical depression (white arrow). 20–22. SEM external view three valves showing the large sternum, the spine structure, the large apical pore fields and the areolae decreasing in diameter towards the sternum. Note the circular series of small siliceous papillae around the areolae and the spines. 23. SEM external detail of the valve apex with the apical pore field. 24. SEM external detail of the areolae and the spines. 25. SEM internal view of a complete valve. Scale bars: 10 μm (1–17); 1 μm (18–25)." figureDoi="http://doi.org/10.5281/zenodo.14515991" httpUri="https://zenodo.org/record/14515991/files/figure.png" pageId="1" pageNumber="50">Figs 22, 25</figureCitation>
). Spines and areolae on both mantle and valve face surrounded by small siliceous papillae, arranged in circles (
<figureCitation id="D0E0D3857D6DF124FA90F79C06E1F7F5" box="[1305,1439,2056,2082]" captionStart="FIGURES 1–25" captionStartId="2.[136,248,1848,1870]" captionTargetBox="[193,1394,187,1821]" captionTargetId="figure-25@2.[193,1394,187,1821]" captionTargetPageId="2" captionText="FIGURES 1–25. Pseudostaurosira katharinae Van de Vijver &amp; C.E.Wetzel sp. nov. LM and SEM taken from the holotype sample (BR- 4850, Bosbeek, Maaseik, Belgium). 1–2. LM pictures of two connected frustules in girdle view. 3–17. LM pictures of valves in valve face view in decreasing length. 18–19 SEM external view of a complete frustule in girdle view showing the girdle structure, the large mantle with the mantle plaques and the apical depression (white arrow). 20–22. SEM external view three valves showing the large sternum, the spine structure, the large apical pore fields and the areolae decreasing in diameter towards the sternum. Note the circular series of small siliceous papillae around the areolae and the spines. 23. SEM external detail of the valve apex with the apical pore field. 24. SEM external detail of the areolae and the spines. 25. SEM internal view of a complete valve. Scale bars: 10 μm (1–17); 1 μm (18–25)." figureDoi="http://doi.org/10.5281/zenodo.14515991" httpUri="https://zenodo.org/record/14515991/files/figure.png" pageId="1" pageNumber="50">Figs 21, 24</figureCitation>
). Apical pore fields present on both apices, large, composed of several rows of pores located in linear depressions between raised small siliceous papillae (
<figureCitation id="D0E0D3857D6FF126FDDCFF5601D2FF0B" box="[597,684,194,220]" captionStart="FIGURES 1–25" captionStartId="2.[136,248,1848,1870]" captionTargetBox="[193,1394,187,1821]" captionTargetId="figure-25@2.[193,1394,187,1821]" captionTargetPageId="2" captionText="FIGURES 1–25. Pseudostaurosira katharinae Van de Vijver &amp; C.E.Wetzel sp. nov. LM and SEM taken from the holotype sample (BR- 4850, Bosbeek, Maaseik, Belgium). 1–2. LM pictures of two connected frustules in girdle view. 3–17. LM pictures of valves in valve face view in decreasing length. 18–19 SEM external view of a complete frustule in girdle view showing the girdle structure, the large mantle with the mantle plaques and the apical depression (white arrow). 20–22. SEM external view three valves showing the large sternum, the spine structure, the large apical pore fields and the areolae decreasing in diameter towards the sternum. Note the circular series of small siliceous papillae around the areolae and the spines. 23. SEM external detail of the valve apex with the apical pore field. 24. SEM external detail of the areolae and the spines. 25. SEM internal view of a complete valve. Scale bars: 10 μm (1–17); 1 μm (18–25)." figureDoi="http://doi.org/10.5281/zenodo.14515991" httpUri="https://zenodo.org/record/14515991/files/figure.png" pageId="3" pageNumber="52">Fig. 23</figureCitation>
). Internally, areolae located each in a single depression covered by rounded volae (
<figureCitation id="D0E0D3857D6FF126FEBEFF7202F5FED6" box="[311,395,230,257]" captionStart="FIGURES 1–25" captionStartId="2.[136,248,1848,1870]" captionTargetBox="[193,1394,187,1821]" captionTargetId="figure-25@2.[193,1394,187,1821]" captionTargetPageId="2" captionText="FIGURES 1–25. Pseudostaurosira katharinae Van de Vijver &amp; C.E.Wetzel sp. nov. LM and SEM taken from the holotype sample (BR- 4850, Bosbeek, Maaseik, Belgium). 1–2. LM pictures of two connected frustules in girdle view. 3–17. LM pictures of valves in valve face view in decreasing length. 18–19 SEM external view of a complete frustule in girdle view showing the girdle structure, the large mantle with the mantle plaques and the apical depression (white arrow). 20–22. SEM external view three valves showing the large sternum, the spine structure, the large apical pore fields and the areolae decreasing in diameter towards the sternum. Note the circular series of small siliceous papillae around the areolae and the spines. 23. SEM external detail of the valve apex with the apical pore field. 24. SEM external detail of the areolae and the spines. 25. SEM internal view of a complete valve. Scale bars: 10 μm (1–17); 1 μm (18–25)." figureDoi="http://doi.org/10.5281/zenodo.14515991" httpUri="https://zenodo.org/record/14515991/files/figure.png" pageId="3" pageNumber="52">Fig. 25</figureCitation>
).
</paragraph>
</subSubSection>
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<paragraph id="4864CF007D6EF127FF01F8AC066FF7F1" blockId="2.[136,1451,1848,2086]" pageId="2" pageNumber="51">
<emphasis id="7AAF13127D6EF127FF01F8AC024BF89A" bold="true" box="[136,309,1848,1870]" pageId="2" pageNumber="51">FIGURES 1–25.</emphasis>
<taxonomicName id="8FDBB4837D6EF127FEB2F8AC0018F899" authority="Van de Vijver &amp; C.E.Wetzel" authorityName="Van de Vijver &amp; C. E. Wetzel" authorityYear="2024" box="[315,870,1848,1870]" class="Bacillariophyceae" family="Fragilariaceae" genus="Pseudostaurosira" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fragilariales" pageId="2" pageNumber="51" phylum="Ochrophyta" rank="species" species="katharinae" status="sp. nov.">
<emphasis id="7AAF13127D6EF127FEB2F8AC012FF89A" box="[315,593,1848,1869]" italics="true" pageId="2" pageNumber="51">Pseudostaurosira katharinae</emphasis>
Van de Vijver &amp; C.E.Wetzel
</taxonomicName>
<emphasis id="7AAF13127D6EF127FCE5F8AD00C8F899" box="[876,950,1849,1870]" italics="true" pageId="2" pageNumber="51">
<taxonomicNameLabel id="619CAE697D6EF127FCE5F8AD00C8F899" box="[876,950,1849,1870]" pageId="2" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
LM and SEM taken from the holotype sample (BR- 4850, Bosbeek, Maaseik, Belgium). 1–2. LM pictures of two connected frustules in girdle view. 3–17. LM pictures of valves in valve face view in decreasing length. 18–19 SEM external view of a complete frustule in girdle view showing the girdle structure, the large mantle with the mantle plaques and the apical depression (white arrow). 20–22. SEM external view three valves showing the large sternum, the spine structure, the large apical pore fields and the areolae decreasing in diameter towards the sternum. Note the circular series of small siliceous papillae around the areolae and the spines. 23. SEM external detail of the valve apex with the apical pore field. 24. SEM external detail of the areolae and the spines. 25. SEM internal view of a complete valve. Scale bars: 10 μm (1–17); 1 μm (18–25).
</paragraph>
</caption>
<subSubSection id="00C19C8B7D6FF126FF34FE9E0034FE58" pageId="3" pageNumber="52" type="materials_examined">
<paragraph id="4864CF007D6FF126FF34FE9E070CFEBC" blockId="3.[136,1452,158,1038]" pageId="3" pageNumber="52">
<emphasis id="7AAF13127D6FF126FF34FE9E0260FEF3" bold="true" box="[189,286,266,292]" pageId="3" pageNumber="52">Type:—</emphasis>
<collectingCountry id="30CC8F907D6FF126FE94FE9E02DEFEF3" box="[285,416,266,292]" name="Belgium" pageId="3" pageNumber="52">BELGIUM</collectingCountry>
.
<collectingRegion id="8A1F01E27D6FF126FE3AFE9E0168FEF3" box="[435,534,266,292]" country="Belgium" name="Vlaanderen" pageId="3" pageNumber="52">Flanders</collectingRegion>
, Province of Limburg, Maaseik, Bosbeek, sample
<collectionCode id="2ECA57C57D6FF126FBF6FE9E07BCFEF3" box="[1151,1218,266,292]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15304" lsid="urn:lsid:biocol.org:col:15304" name="Algonquin Provincial Park" pageId="3" pageNumber="52" type="Herbarium">APM</collectionCode>
21-91,
<geoCoordinate id="2DEFA9C77D6FF126FA9AFE9D06D9FEF3" box="[1299,1447,265,292]" degrees="51" direction="north" minutes="5.6348" orientation="latitude" pageId="3" pageNumber="52" precision="1" value="51.093914">51°5.6348’N</geoCoordinate>
,
<geoCoordinate id="2DEFA9C77D6FF126FF01FEB90277FE9F" box="[136,265,301,328]" degrees="5" direction="east" minutes="45.894" orientation="longitude" pageId="3" pageNumber="52" precision="1" value="5.7649">5°45.894’E</geoCoordinate>
, coll. date
<date id="3C65E9C07D6FF126FE18FEB90153FE90" box="[401,557,301,328]" pageId="3" pageNumber="52" value="2021-06-25">25 Jun. 2021</date>
, leg. Vlaamse Milieu maatschappij (
<collectionCode id="2ECA57C57D6FF126FC53FEB9075DFE90" box="[986,1059,301,327]" country="USA" httpUri="http://grbio.org/cool/xptj-nxev" name="Vanderbilt Marine Museum" pageId="3" pageNumber="52">VMM</collectionCode>
) (
<typeStatus id="976071A27D6FF126FBB4FEB907DFFE90" box="[1085,1185,301,327]" pageId="3" pageNumber="52" type="holotype">holotype</typeStatus>
slide BR-4850,
<figureCitation id="D0E0D3857D6FF126FAEFFEB906D5FE9F" box="[1382,1451,301,328]" captionStart="FIGURES 1–25" captionStartId="2.[136,248,1848,1870]" captionTargetBox="[193,1394,187,1821]" captionTargetId="figure-25@2.[193,1394,187,1821]" captionTargetPageId="2" captionText="FIGURES 1–25. Pseudostaurosira katharinae Van de Vijver &amp; C.E.Wetzel sp. nov. LM and SEM taken from the holotype sample (BR- 4850, Bosbeek, Maaseik, Belgium). 1–2. LM pictures of two connected frustules in girdle view. 3–17. LM pictures of valves in valve face view in decreasing length. 18–19 SEM external view of a complete frustule in girdle view showing the girdle structure, the large mantle with the mantle plaques and the apical depression (white arrow). 20–22. SEM external view three valves showing the large sternum, the spine structure, the large apical pore fields and the areolae decreasing in diameter towards the sternum. Note the circular series of small siliceous papillae around the areolae and the spines. 23. SEM external detail of the valve apex with the apical pore field. 24. SEM external detail of the areolae and the spines. 25. SEM internal view of a complete valve. Scale bars: 10 μm (1–17); 1 μm (18–25)." figureDoi="http://doi.org/10.5281/zenodo.14515991" httpUri="https://zenodo.org/record/14515991/files/figure.png" pageId="3" pageNumber="52">Fig. 5</figureCitation>
represents the
<typeStatus id="976071A27D6FF126FEA3FEC502F1FEBC" box="[298,399,337,363]" pageId="3" pageNumber="52" type="holotype">holotype</typeStatus>
,
<typeStatus id="976071A27D6FF126FE13FEC50295FEBC" box="[410,491,337,363]" pageId="3" pageNumber="52" type="isotype">isotype</typeStatus>
slide
<quantity id="8F2362E57D6FF126FDA7FEC5010BFEBC" box="[558,629,337,363]" metricMagnitude="1" metricUnit="m" metricValue="1.13792" pageId="3" pageNumber="52" unit="in" value="448.0">448 in</quantity>
Collection University of Antwerp, Belgium).
</paragraph>
<paragraph id="4864CF007D6FF126FF34FEE00034FE58" blockId="3.[136,1452,158,1038]" box="[189,842,372,399]" pageId="3" pageNumber="52">
<emphasis id="7AAF13127D6FF126FF34FEE00280FE59" bold="true" box="[189,510,372,398]" pageId="3" pageNumber="52">PhycoBank registration:—</emphasis>
http://phycobank.org/104825.
</paragraph>
</subSubSection>
<subSubSection id="00C19C8B7D6FF126FF34FE0C063BFE66" box="[189,1349,407,434]" pageId="3" pageNumber="52" type="etymology">
<paragraph id="4864CF007D6FF126FF34FE0C063BFE66" blockId="3.[136,1452,158,1038]" box="[189,1349,407,434]" pageId="3" pageNumber="52">
<emphasis id="7AAF13127D6FF126FF34FE0C021AFE65" bold="true" box="[189,356,408,434]" pageId="3" pageNumber="52">Etymology:—</emphasis>
The new species is named after
<taxonomicName id="8FDBB4837D6FF126FD43FE0300EFFE66" authority="Wilfert" authorityName="Wilfert" box="[714,913,407,433]" class="Polyplacophora" family="Mopaliidae" genus="Katharina" kingdom="Animalia" order="Chitonida" pageId="3" pageNumber="52" phylum="Mollusca" rank="genus">Katharina Wilfert</taxonomicName>
, oldest stepdaughter of the first author.
</paragraph>
</subSubSection>
<subSubSection id="00C19C8B7D6FF126FF34FE2F063BFBD9" pageId="3" pageNumber="52" type="biology_ecology">
<paragraph id="4864CF007D6FF126FF34FE2F0278FC3E" blockId="3.[136,1452,158,1038]" pageId="3" pageNumber="52">
<emphasis id="7AAF13127D6FF126FF34FE2F0177FE02" bold="true" box="[189,521,443,469]" pageId="3" pageNumber="52">Ecology and distribution:—</emphasis>
<taxonomicName id="8FDBB4837D6FF126FD86FE2F002DFE02" authorityName="Van de Vijver &amp; C. E. Wetzel" authorityYear="2024" box="[527,851,443,469]" class="Bacillariophyceae" family="Fragilariaceae" genus="Pseudostaurosira" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fragilariales" pageId="3" pageNumber="52" phylum="Ochrophyta" rank="species" species="katharinae">
<emphasis id="7AAF13127D6FF126FD86FE2F002DFE02" box="[527,851,443,469]" italics="true" pageId="3" pageNumber="52">Pseudostaurosira katharinae</emphasis>
</taxonomicName>
was described from a small river near Maaseik, in the northeastern part of
<collectingCountry id="30CC8F907D6FF126FEE3FE4A02B0FE2F" box="[362,462,478,504]" name="Belgium" pageId="3" pageNumber="52">Belgium</collectingCountry>
. The
<typeStatus id="976071A27D6FF126FD83FE4B0144FE2E" box="[522,570,479,505]" pageId="3" pageNumber="52">type</typeStatus>
locality had an almost circumneutral pH (6.9–7.3), a moderate conductivity (&gt; 200 µS/cm), and higher sulphate (
<quantity id="8F2362E57D6FF126FD80FD960103FDCB" box="[521,637,514,540]" metricMagnitude="-5" metricUnit="kg" metricValue="2.1" metricValueMax="3.0" metricValueMin="1.2" pageId="3" pageNumber="52" unit="mg" value="21.0" valueMax="30.0" valueMin="12.0">12–30 mg</quantity>
/l) and nitrate levels (
<quantity id="8F2362E57D6FF126FCE7FD96008CFDCB" box="[878,1010,514,540]" metricMagnitude="-6" metricUnit="kg" metricValue="2.3499999999999996" metricValueMax="3.3" metricValueMin="1.4" pageId="3" pageNumber="52" unit="mg" value="2.3499999999999996" valueMax="3.3" valueMin="1.4">1.4–3.3 mg</quantity>
/l). The associated flora was dominated by several small-celled araphid species such as, apart from
<taxonomicName id="8FDBB4837D6FF126FCC9FDB200A6FDE8" authorityName="Van de Vijver &amp; C. E. Wetzel" authorityYear="2024" box="[832,984,549,575]" class="Bacillariophyceae" family="Fragilariaceae" genus="Pseudostaurosira" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fragilariales" pageId="3" pageNumber="52" phylum="Ochrophyta" rank="species" species="katharinae">
<emphasis id="7AAF13127D6FF126FCC9FDB200A6FDE8" box="[832,984,549,575]" italics="true" pageId="3" pageNumber="52">P. katharinae</emphasis>
</taxonomicName>
, also larger populations of
<taxonomicName id="8FDBB4837D6FF126FA92FDB1012AFDB4" authority="Van de Vijver &amp; V. Peeters" authorityName="Van de Vijver &amp; V. Peeters" class="Bacillariophyceae" family="Fragilariaceae" genus="Staurosirella" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fragilariales" pageId="3" pageNumber="52" phylum="Ochrophyta" rank="species" species="marginostriata">
<emphasis id="7AAF13127D6FF126FA92FDB10251FDB4" italics="true" pageId="3" pageNumber="52">Staurosirella marginostriata</emphasis>
Van de Vijver &amp; V.Peeters
</taxonomicName>
(in
<bibRefCitation id="2C4AB2F17D6FF126FDF7FDDD00B7FDB4" author="Van de Vijver, B. &amp; Peeters, V. &amp; Ballings, P. &amp; de Haan, M." box="[638,969,585,611]" pageId="3" pageNumber="52" pagination="139 - 160" refId="ref4775" refString="Van de Vijver, B., Peeters, V., Ballings, P. &amp; de Haan, M. (2024) Five new species in the genus Staurosirella (Bacillariophyta) from European freshwater habitats. PhytoKeys 242: 139 - 160. https: // doi. org / 10.3897 / phytokeys. 242.122458" type="journal article" year="2024">
Van de Vijver
<emphasis id="7AAF13127D6FF126FC93FDDE002FFDB4" box="[794,849,585,611]" italics="true" pageId="3" pageNumber="52">et al.</emphasis>
2024: 142
</bibRefCitation>
),
<taxonomicName id="8FDBB4837D6FF126FC55FDDD071FFDB4" box="[988,1121,585,611]" class="Bacillariophyceae" family="Fragilariaceae" genus="Staurosirella" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fragilariales" pageId="3" pageNumber="52" phylum="Ochrophyta" rank="species" species="stoksiana">
<emphasis id="7AAF13127D6FF126FC55FDDD071FFDB4" box="[988,1121,585,611]" italics="true" pageId="3" pageNumber="52">S. stoksiana</emphasis>
</taxonomicName>
,
<taxonomicName id="8FDBB4837D6FF126FBE7FDDD06D2FDB4" box="[1134,1452,585,611]" class="Bacillariophyceae" family="Fragilariaceae" genus="Pseudostaurosira" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fragilariales" pageId="3" pageNumber="52" phylum="Ochrophyta" rank="species" species="alvareziae">
<emphasis id="7AAF13127D6FF126FBE7FDDD06D2FDB4" box="[1134,1452,585,611]" italics="true" pageId="3" pageNumber="52">Pseudostaurosira alvareziae</emphasis>
</taxonomicName>
and
<taxonomicName id="8FDBB4837D6FF126FF3FFDF9022CFD51" authorityName="D.M.Williams &amp; Round" authorityYear="1987" box="[182,338,620,646]" class="Bacillariophyceae" family="Fragilariaceae" genus="Pseudostaurosira" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fragilariales" pageId="3" pageNumber="52" phylum="Ochrophyta" rank="species" species="brevistriata">
<emphasis id="7AAF13127D6FF126FF3FFDF9022CFD51" box="[182,338,620,646]" italics="true" pageId="3" pageNumber="52">P. brevistriata</emphasis>
</taxonomicName>
. Other dominant species in the sample included
<taxonomicName id="8FDBB4837D6FF126FCE1FDF80729FD51" authorityName="Simonsen" authorityYear="1979" box="[872,1111,620,646]" class="Bacillariophyceae" family="Aulacoseiraceae" genus="Aulacoseira" higherTaxonomySource="GBIF" kingdom="Chromista" order="Aulacoseirales" pageId="3" pageNumber="52" phylum="Ochrophyta" rank="species" species="ambigua">
<emphasis id="7AAF13127D6FF126FCE1FDF80729FD51" box="[872,1111,620,646]" italics="true" pageId="3" pageNumber="52">Aulacoseira ambigua</emphasis>
</taxonomicName>
(Grunow in
<bibRefCitation id="2C4AB2F17D6FF126FB6BFDF906D6FD51" author="Van Heurck, H." box="[1250,1448,620,647]" pageId="3" pageNumber="52" refId="ref4834" refString="Van Heurck, H. (1882) Synopsis des Diatomees de Belgique. Atlas. Ducaju et Cie, Anvers, pp. pls LXXVIII - CXXXII." type="book" year="1882">Van Heurck 1882</bibRefCitation>
: pl. 88, figs 12–15)
<bibRefCitation id="2C4AB2F17D6FF126FEEEFD040127FD7C" author="Simonsen, R." box="[359,601,656,683]" pageId="3" pageNumber="52" pagination="9 - 71" refId="ref4589" refString="Simonsen, R. (1979) The diatom system: ideas on phylogeny. Bacillaria 2: 9 - 71." type="journal article" year="1979">Simonsen (1979: 56)</bibRefCitation>
,
<taxonomicName id="8FDBB4837D6FF126FDEEFD040675FD7C" authority="(Ehrenberg 1843 b: 415) Simonsen (1979: 58)" authorityName="Simonsen" authorityPageNumber="58" authorityYear="1979" baseAuthorityName="Ehrenberg" baseAuthorityPageNumber="415" baseAuthorityYear="1843" box="[615,1291,656,683]" class="Bacillariophyceae" family="Aulacoseiraceae" genus="Aulacoseira" higherTaxonomySource="GBIF" kingdom="Chromista" order="Aulacoseirales" pageId="3" pageNumber="52" phylum="Ochrophyta" rank="species" species="granulata">
<emphasis id="7AAF13127D6FF126FDEEFD040189FD7D" box="[615,759,656,682]" italics="true" pageId="3" pageNumber="52">A. granulata</emphasis>
(
<bibRefCitation id="2C4AB2F17D6FF126FC83FD040774FD7C" author="Ehrenberg, C. G." box="[778,1034,656,683]" pageId="3" pageNumber="52" pagination="291 - 445" refId="ref3667" refString="Ehrenberg, C. G. (1843 b) Verbreitung und Einfluss des mikroskopischen Lebens in Sud- und Nord-Amerika. Abhandlungen der Koniglichen Akademie der Wissenschaften zu Berlin 1841: 291 - 445." type="journal article" year="1843">Ehrenberg 1843b: 415</bibRefCitation>
)
<bibRefCitation id="2C4AB2F17D6FF126FB92FD040675FD7C" author="Simonsen, R." box="[1051,1291,656,683]" pageId="3" pageNumber="52" pagination="9 - 71" refId="ref4589" refString="Simonsen, R. (1979) The diatom system: ideas on phylogeny. Bacillaria 2: 9 - 71." type="journal article" year="1979">Simonsen (1979: 58)</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="8FDBB4837D6FF126FACFFD040176FD19" authority="Kutzing (1844: 95)" authorityName="Kutzing" authorityPageNumber="95" authorityYear="1844" class="Bacillariophyceae" family="Naviculaceae" genus="Navicula" kingdom="Chromista" order="Naviculales" pageId="3" pageNumber="52" phylum="Ochrophyta" rank="species" species="cryptocephala">
<emphasis id="7AAF13127D6FF126FACFFD040256FD1A" italics="true" pageId="3" pageNumber="52">Navicula cryptocephala</emphasis>
<bibRefCitation id="2C4AB2F17D6FF126FEA6FD270176FD19" author="Kutzing, F. T." box="[303,520,691,718]" pageId="3" pageNumber="52" refId="ref3983" refString="Kutzing, F. T. (1844) Die kieselschaligen Bacillarien oder Diatomeen. W. Kohne, Nordhausen, 144 pp. https: // doi. org / 10.5962 / bhl. title. 64360" type="book" year="1844">Kützing (1844: 95)</bibRefCitation>
</taxonomicName>
, pointing to more meso-eutrophic, alkaline conditions (Lange-Bertalot
<emphasis id="7AAF13127D6FF126FAADFD200623FD1A" box="[1316,1373,691,717]" italics="true" pageId="3" pageNumber="52">et al.</emphasis>
2017). The species has also been found in a French lake (“
<emphasis id="7AAF13127D6FF126FD47FD4C00FAFD26" box="[718,900,727,753]" italics="true" pageId="3" pageNumber="52">Mare à Goriaux</emphasis>
”) (
<figureCitation id="D0E0D3857D6FF126FC23FD43073BFD26" box="[938,1093,727,753]" captionStart="FIGURES 26–27" captionStartId="3.[136,248,1691,1713]" captionTargetBox="[151,1436,1062,1667]" captionTargetId="figure-456@3.[151,1436,1062,1667]" captionTargetPageId="3" captionText="FIGURES 26–27. Pseudostaurosira katharinae Van de Vijver &amp; C.E.Wetzel sp. nov. Specimens from France (Mare à Goriaux). SEM external view of a complete frustules in girdle view. Fig. 26: Detail showing the fracted girdle bands (white arrow); mantle plates of reduced size. Fig. 27: Open valvocopulae with fimbriate girdle elements. Serrate spines. Scale bars: 4 μm (26); 5 μm (27)." figureDoi="http://doi.org/10.5281/zenodo.14515993" httpUri="https://zenodo.org/record/14515993/files/figure.png" pageId="3" pageNumber="52">Figs 26 &amp; 27</figureCitation>
). The Mare à Goriaux, an intraforest pond, is located within the perimeter of the Regional Natural Park of the Scarpe and Escaut Plains, at the southern edge of the Raismes, St-Amand, and Wallers state forest (
<bibRefCitation id="2C4AB2F17D6FF126FCFCFC8A0745FCEF" author="Marchyllie, M." box="[885,1083,798,824]" pageId="3" pageNumber="52" refId="ref4088" refString="Marchyllie, M. (1992) Reserve biologique domaniale de la Mare a Goriaux: etude des roselieres en vue de leur restrauration et de leur gestion. 101 pp." type="book" year="1992">Marchyllie, 1992</bibRefCitation>
). The pond, formed from mining subsidence caused by the exploitation of the Vicoigne and Arenberg pits, was originally a wetland due to groundwater outcrops. The exploitation of the Arenberg pit caused the most significant collapses, initially creating three ponds that merged around the 1930s to
<taxonomicName id="8FDBB4837D6FF126FE5AFC1C014BFC75" box="[467,565,904,930]" form="the" pageId="3" pageNumber="52" rank="form">form the</taxonomicName>
current Mare à Goriaux (
<bibRefCitation id="2C4AB2F17D6FF126FCD6FC1C0756FC75" author="Marchyllie, M." box="[863,1064,904,930]" pageId="3" pageNumber="52" refId="ref4088" refString="Marchyllie, M. (1992) Reserve biologique domaniale de la Mare a Goriaux: etude des roselieres en vue de leur restrauration et de leur gestion. 101 pp." type="book" year="1992">Marchyllie, 1992</bibRefCitation>
). The high mineral content of the pond’s water is largely due to SO
<subScript id="D45FCD457D6FF126FE7EFC2A0281FC1B" attach="left" box="[503,511,958,972]" fontSize="6" pageId="3" pageNumber="52">4</subScript>
<superScript id="BFAE62487D6FF126FE76FC38017AFC6D" attach="right" box="[511,516,940,954]" fontSize="6" pageId="3" pageNumber="52">-</superScript>
ions (
<quantity id="8F2362E57D6FF126FDCEFC3F01B1FC12" box="[583,719,939,966]" metricMagnitude="-4" metricUnit="kg" metricValue="3.53" metricValueMax="7.0" metricValueMin="0.06" pageId="3" pageNumber="52" unit="mg" value="353.0" valueMax="700.0" valueMin="6.0">6 to 700 mg</quantity>
/l),
<superScript id="BFAE62487D6FF126FD78FC3F006FFC6D" attach="left" box="[753,785,939,965]" fontSize="6" pageId="3" pageNumber="52">Cl-</superScript>
ions (
<quantity id="8F2362E57D6FF126FCDDFC3F00D2FC12" box="[852,940,939,966]" metricMagnitude="-4" metricUnit="kg" metricValue="1.45" pageId="3" pageNumber="52" unit="mg" value="145.0">145 mg</quantity>
/l), as well as Na
<superScript id="BFAE62487D6FF126FBE9FC380717FC6D" attach="left" box="[1120,1129,940,954]" fontSize="6" pageId="3" pageNumber="52">+</superScript>
ions (
<quantity id="8F2362E57D6FF126FB25FC3F067CFC12" box="[1196,1282,939,966]" metricMagnitude="-4" metricUnit="kg" metricValue="5.6" pageId="3" pageNumber="52" unit="mg" value="560.0">560 mg</quantity>
/l) and Ca
<superScript id="BFAE62487D6FF126FAE4FC380608FC6D" attach="left" box="[1389,1398,940,954]" fontSize="6" pageId="3" pageNumber="52">+</superScript>
ions (
<quantity id="8F2362E57D6FF126FF19FC5B0396FC3E" box="[144,232,975,1001]" metricMagnitude="-4" metricUnit="kg" metricValue="1.1" pageId="3" pageNumber="52" unit="mg" value="110.0">110 mg</quantity>
/l).
</paragraph>
<paragraph id="4864CF007D6FF126FF34FC60063BFBD9" blockId="3.[136,1452,158,1038]" box="[189,1349,1011,1038]" pageId="3" pageNumber="52">It is likely that the species is more widespread than currently known due to confusion with similar taxa.</paragraph>
</subSubSection>
</treatment>
</document>

727
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@ -0,0 +1,727 @@
<document id="3BF251D9614B0035DF12F2BB33FFDBDB" ID-DOI="10.11646/phytotaxa.663.2.3" ID-ISSN="1179-3163" ID-Zenodo-Dep="14515919" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1734536036199" checkinUser="felipe" docAuthor="Kuethe, J. R., Bernal, Hernan Dario, Molinari, Miguel &amp; Vecchia, Maurizio" docDate="2024" docId="F109C35AFFE2FF8CFF1DFD9696C280D9" docLanguage="en" docName="phytotaxa.663.2.077-089.pdf" docOrigin="Phytotaxa 663 (2)" docSource="https://doi.org/10.11646/phytotaxa.663.2.3" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Passiflora rogitamensis Molinari, Kuethe &amp; Bernal 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="83" masterDocId="0D30BB22FFE7FF8AFF95FFD892158745" masterDocTitle="Validating the names of three species of Passiflora from the northern portion of South America" masterLastPageNumber="89" masterPageNumber="77" pageNumber="82" updateTime="1734536434291" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="36023832A5DA0C96E550D9A2E642C3F7">Validating the names of three species of Passiflora from the northern portion of South America</mods:title>
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<mods:namePart id="3C663F48B8772F76BC10A7945691A2E9">Kuethe, J. R.</mods:namePart>
<mods:affiliation id="5DBC55F693C0E099FBB1481A092B0967">School of Environment, University of Auckland, New Zealand</mods:affiliation>
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<mods:namePart id="CD6DE8197C576737C2903AB73914F7EE">Bernal, Hernan Dario</mods:namePart>
<mods:affiliation id="9B7D9ED08791D1BCC94376B1EC0E730F">Passifloras de Colombia, Independent researcher and collection holder, Bogotá, Colombia</mods:affiliation>
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<mods:namePart id="846EC9CDEA9CF925AAC5438EA1A6FB8E">Molinari, Miguel</mods:namePart>
<mods:affiliation id="FF7BF8710E9E195892AFE3C7EB2A4C5D">Universidad de los Andes, Facultad de Ciencias Forestales, Herbarium (MER), Mérida, Venezuela</mods:affiliation>
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<mods:namePart id="6F5931DF1A613406FBCADF1EB79A6950">Vecchia, Maurizio</mods:namePart>
<mods:affiliation id="406DBFD6359735EFD2E17170033C111A">School of Environment, University of Auckland, New Zealand &amp; National Italian Passiflora Collection, Societá Botanica Italiana, Ripalta Cremasca, Italy &amp; School of Environment, University of Auckland, New Zealand</mods:affiliation>
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<mods:title id="62007C58F96C08F9FFC1475556F905DA">Phytotaxa</mods:title>
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<mods:date id="555C540A46BAF9B9295EA5AA0B0C4756">2024</mods:date>
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<paragraph id="791F724CFFE2FF8FFF1DFD969198852D" blockId="5.[136,909,590,617]" box="[136,909,590,617]" pageId="5" pageNumber="82">
<heading id="2257C520FFE2FF8FFF1DFD969198852D" box="[136,909,590,617]" fontSize="11" level="2" pageId="5" pageNumber="82" reason="3">
<taxonomicName id="BEA009CFFFE2FF8FFF1DFD9690DB852D" ID-CoL="CKNCF" authority="Molinari, Kuethe &amp; Bernal" authorityName="Molinari, Kuethe &amp; Bernal" authorityYear="2024" box="[136,718,590,617]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="82" phylum="Tracheophyta" rank="species" species="rogitamensis" status="sp. nov.">
<emphasis id="4BD4AE5EFFE2FF8FFF1DFD96938D852D" bold="true" box="[136,408,590,617]" italics="true" pageId="5" pageNumber="82">Passiflora rogitamensis</emphasis>
Molinari, Kuethe &amp; Bernal
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<emphasis id="4BD4AE5EFFE2FF8FFD40FD979131852D" box="[725,804,591,616]" italics="true" pageId="5" pageNumber="82">sp. nov</emphasis>
.
</taxonomicNameLabel>
(
<figureCitation id="E19B6EC9FFE2FF8FFCAFFD969195852D" box="[826,896,590,616]" captionStart="FIGURE 4" captionStartId="7.[136,234,1883,1905]" captionTargetBox="[210,1378,190,1858]" captionTargetId="figure-19@7.[210,1378,190,1858]" captionTargetPageId="7" captionText="FIGURE 4. A–F. Passiflora rogitamensis., Lankaster Composite Dissection Plate (LCDP); A. flower in situ; B. comparison between P. rogitamensis (B1) and P. longipes (B2); C. Habit in situ; C1. Detail of the stipules; C2. Detail of the petiolar nectaries; D. Longitudinal of the flower; D1. Detail of the bracts; E. Detail of the fruits; F. Direct comparison between the leaves of P.rogitamensis (F1) and P. longipes (F2). Compiled from photographs taken by Miguel Molinari, Hernan Bernal and Christian Moliné." figureDoi="http://doi.org/10.5281/zenodo.14515933" httpUri="https://zenodo.org/record/14515933/files/figure.png" pageId="5" pageNumber="82">Fig. 4</figureCitation>
).
</heading>
</paragraph>
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<emphasis id="4BD4AE5EFFE2FF8FFF1DFD4292D785EA" bold="true" box="[136,194,666,687]" pageId="5" pageNumber="82">Type:</emphasis>
—
<materialsCitation id="C9C87811FFE2FF8FFF4EFD4291A18596" collectingDate="2019-10-04" collectionCode="COL" collectorName="Reserva Natural de la Sociedad Civil Rogitama &amp; M. Molinari" country="Colombia" elevation="2470" latitude="5.79" location="Reserva Natural de la Sociedad Civil Rogitama" longLatPrecision="15" longitude="-73.4573" municipality="Arcabuco" pageId="5" pageNumber="82" specimenCount="2" stateProvince="Boyaca" typeStatus="holotype">
<collectingCountry id="01B732DCFFE2FF8FFF4EFD42934885EA" box="[219,349,666,687]" name="Colombia" pageId="5" pageNumber="82">COLOMBIA</collectingCountry>
.
<collectingRegion id="BB64BCAEFFE2FF8FFEF3FD4293A785F5" box="[358,434,666,688]" country="Colombia" name="Boyaca" pageId="5" pageNumber="82">Boyacá</collectingRegion>
:
<collectingMunicipality id="997BE836FFE2FF8FFE2CFD42900C85F5" box="[441,537,666,688]" pageId="5" pageNumber="82">Arcabuco</collectingMunicipality>
,
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,
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,
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,
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. a.s.l.,
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, (fl., fr.)
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<collectorName id="D455179AFFE2FF8FFE0DFD6790188596" box="[408,525,702,724]" pageId="5" pageNumber="82">M. Molinari</collectorName>
s.n.
</emphasis>
(
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: JBB32921!;
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:
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)
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.
</paragraph>
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<subSubSection id="31BA21C7FFE2FF8FFF1DFD3A93008426" pageId="5" pageNumber="82" type="diagnosis">
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<emphasis id="4BD4AE5EFFE2FF8FFF1DFD3A92E685B2" bold="true" box="[136,243,738,759]" pageId="5" pageNumber="82">
<emphasis id="4BD4AE5EFFE2FF8FFF1DFD3A92FE85B2" bold="true" box="[136,235,738,759]" italics="true" pageId="5" pageNumber="82">Diagnosis</emphasis>
:
</emphasis>
—
<taxonomicName id="BEA009CFFFE2FF8FFE9EFD3A93FA85B2" authorityName="Molinari, Kuethe &amp; Bernal" authorityYear="2024" box="[267,495,738,759]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="82" phylum="Tracheophyta" rank="species" species="rogitamensis">
<emphasis id="4BD4AE5EFFE2FF8FFE9EFD3A93FA85B2" box="[267,495,738,759]" italics="true" pageId="5" pageNumber="82">Passiflora rogitamensis</emphasis>
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is similar to
<taxonomicName id="BEA009CFFFE2FF8FFDFBFD3B90C285B2" authorityName="Jussieu" authorityYear="1805" box="[622,727,738,760]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="82" phylum="Tracheophyta" rank="species" species="longipes">
<emphasis id="4BD4AE5EFFE2FF8FFDFBFD3B90C285B2" box="[622,727,738,760]" italics="true" pageId="5" pageNumber="82">P. longipes</emphasis>
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but differs in the colour of the corona (white
<emphasis id="4BD4AE5EFFE2FF8FFB06FD3B96B285BD" box="[1171,1191,739,760]" italics="true" pageId="5" pageNumber="82">vs</emphasis>
. purple), shape and size of the stipules (oblong,&gt;
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<emphasis id="4BD4AE5EFFE2FF8FFE45FCDF93F18459" box="[464,484,775,796]" italics="true" pageId="5" pageNumber="82">vs</emphasis>
. ovate, &lt;
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), and the shape of the fruit (long ellipsoid
<emphasis id="4BD4AE5EFFE2FF8FFB82FCDF963E8459" box="[1047,1067,775,796]" italics="true" pageId="5" pageNumber="82">vs</emphasis>
. ovate-ovoid). It also differs notably in the shape and size of the leaves, which are linear oblong to lanceolate (up to
<quantity id="BE58DFA9FFE2FF8FFC3BFCF291FB847A" box="[942,1006,810,832]" metricMagnitude="-1" metricUnit="m" metricValue="1.6" pageId="5" pageNumber="82" unit="cm" value="16.0">16 cm</quantity>
), but ovate to oblong-ovate (up to
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) in
<taxonomicName id="BEA009CFFFE2FF8FFA0FFCF3931B8426" authorityName="Jussieu" authorityYear="1805" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="82" phylum="Tracheophyta" rank="species" species="longipes">
<emphasis id="4BD4AE5EFFE2FF8FFA0FFCF3931B8426" italics="true" pageId="5" pageNumber="82">P. longipes</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="31BA21C7FFE2FF8FFF1DFC4A92CC8011" pageId="5" pageNumber="82" type="description">
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<emphasis id="4BD4AE5EFFE2FF8FFF1DFC4A930984E9" bold="true" box="[136,284,914,940]" pageId="5" pageNumber="82">Description:</emphasis>
—climbing vine,
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long, glabrous throughout, woody at base, younger parts occasionally somewhat glaucous.
<emphasis id="4BD4AE5EFFE2FF8FFF6DFC6F932C8494" box="[248,313,951,977]" italics="true" pageId="5" pageNumber="82">Stems</emphasis>
terete, reddish-purple to green, internodal distance
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.
<emphasis id="4BD4AE5EFFE2FF8FFC49FC6E96208495" box="[988,1077,950,976]" italics="true" pageId="5" pageNumber="82">Stipules</emphasis>
2.5–4.2 ×
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, semi-ovate or oblong, rounded or obtuse at base, sharp acute or aristulate at apex, margin entire, deep green with reddish venation.
<emphasis id="4BD4AE5EFFE2FF8FFF1DFC2692F7835D" box="[136,226,1022,1048]" italics="true" pageId="5" pageNumber="82">Petioles</emphasis>
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long, terete, bearing 4 or 6 subopposite conspicuous nectaries within the apical quarter of the petiole length, the glands stipitate, very slightly globose at the tip,
<quantity id="BE58DFA9FFE2FF8FFCE0FBFA91C78378" box="[885,978,1058,1085]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="5" pageNumber="82" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
in length, green.
<emphasis id="4BD4AE5EFFE2FF8FFB02FBFA971B8379" box="[1175,1294,1058,1084]" italics="true" pageId="5" pageNumber="82">Leaf blade</emphasis>
6–16 ×
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, linear oblong to lanceolate, obtuse or rounded to semi-rounded at base, acute to acuminate at apex, margin entire, foliar texture coriaceous to chartaceous, deep green and glabrous on both sides, often with reddish purple venation at underneath.
<emphasis id="4BD4AE5EFFE2FF8FFE86FB5693BC83ED" box="[275,425,1166,1192]" italics="true" pageId="5" pageNumber="82">Inflorescence</emphasis>
solitary, very showy, borne on long peduncles.
<emphasis id="4BD4AE5EFFE2FF8FFC52FB56962F83ED" box="[967,1082,1166,1192]" italics="true" pageId="5" pageNumber="82">Peduncles</emphasis>
<quantity id="BE58DFA9FFE2FF8FFBD7FB5696B183EC" box="[1090,1188,1166,1193]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" metricValueMax="1.4" metricValueMin="0.6" pageId="5" pageNumber="82" unit="cm" value="10.0" valueMax="14.0" valueMin="6.0">6–14 cm</quantity>
long, slender, pendent, green to reddish purple, articulate
<quantity id="BE58DFA9FFE2FF8FFD9BFB6A90628388" box="[526,631,1202,1229]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="5" pageNumber="82" unit="cm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 cm</quantity>
beneath the base of the flower.
<emphasis id="4BD4AE5EFFE2FF8FFC76FB6B96398389" box="[995,1068,1203,1228]" italics="true" pageId="5" pageNumber="82">Bracts</emphasis>
1.8–2.1 ×
<quantity id="BE58DFA9FFE2FF8FFB3DFB6A97018388" box="[1192,1300,1202,1229]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="5" pageNumber="82" unit="cm" value="1.9" valueMax="2.0" valueMin="1.8">1.8–2 cm</quantity>
, triangular to triangular-ovate, obtuse to cuneate at base, acute and mucronulate at apex, margins minutely serrated, reddish purple to light green.
<emphasis id="4BD4AE5EFFE2FF8FFEA2FB2293868251" box="[311,403,1274,1300]" italics="true" pageId="5" pageNumber="82">Flowers</emphasis>
<quantity id="BE58DFA9FFE2FF8FFE0BFB2290058250" box="[414,528,1274,1301]" metricMagnitude="-1" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="5" pageNumber="82" unit="cm" value="13.5" valueMax="14.0" valueMin="13.0">13–14 cm</quantity>
in diameter, deep fuchsia pink; hypanthium 0.7–1 ×
<quantity id="BE58DFA9FFE2FF8FFB11FB22971D8250" box="[1156,1288,1274,1301]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="2.7" metricValueMin="2.3" pageId="5" pageNumber="82" unit="cm" value="2.5" valueMax="2.7" valueMin="2.3">2.3–2.7 cm</quantity>
, campanulate, green with purple outside, whitish-yellow inside; sepals 6.0–6.5 ×
<quantity id="BE58DFA9FFE2FF8FFCE7FAC691E7827C" box="[882,1010,1310,1337]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="5" pageNumber="82" unit="cm" value="1.25" valueMax="1.3" valueMin="1.2">1.2–1.3 cm</quantity>
, linear-oblong to linear, obtuse at apex, greenish yellowish abaxially, bright deep pink adaxially,, dorsally corniculate, awn
<quantity id="BE58DFA9FFE2FF8FFBA6FA9A968B8218" box="[1075,1182,1346,1373]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="10.0" metricValueMin="4.0" pageId="5" pageNumber="82" unit="mm" value="7.0" valueMax="10.0" valueMin="4.0">4–10 mm</quantity>
long, green, thin; petals 4.7–5.3 ×
<quantity id="BE58DFA9FFE2FF8FFF6FFABE937082C4" box="[250,357,1382,1409]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="5" pageNumber="82" unit="cm" value="0.9" valueMax="1.0" valueMin="0.8">0.8–1 cm</quantity>
, subequal to the sepals, lanceolate-oblong to linear-oblong, slightly concave at apex, deep fuchsia pink on both sides, membranous; corona composed of 6–9 closely spaced filamentose series, whitish purplish to white becoming darker purple at base; outer series the shortest,
<quantity id="BE58DFA9FFE2FF8FFCECFA7691CF828C" box="[889,986,1454,1481]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="5" pageNumber="82" unit="mm" value="3.5" valueMax="5.0" valueMin="2.0">2–5 mm</quantity>
long, filiform, deep purple to lavender, outfacing; intermediate series becoming progressively longer in length,
<quantity id="BE58DFA9FFE2FF8FFC56FA0A965382A8" box="[963,1094,1490,1517]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" metricValueMax="2.1" metricValueMin="1.3" pageId="5" pageNumber="82" unit="cm" value="1.7" valueMax="2.1" valueMin="1.3">1.3–2.1 cm</quantity>
, purple at base soon becoming whitish distally, filiform, erect; inner series hidden,
<quantity id="BE58DFA9FFE2FF8FFD47FA2E91248154" box="[722,817,1526,1553]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="5" pageNumber="82" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">6–8 mm</quantity>
, inclined towards the androgynophore, pinkish lavender to pinkish; operculum
<quantity id="BE58DFA9FFE2FF8FFE05F9C293E58170" box="[400,496,1562,1589]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="5" pageNumber="82" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6–7 mm</quantity>
long, membranous, cleft nearly to base into clavate segments, greenish becoming purple distally, upper margin crenulate, terminating into dorsally attached filiform appendages,; limen whitish to whitish green, membranous, inclined upwards towards the base of the androgynophore, not spotted; androgynophore
<quantity id="BE58DFA9FFE2FF8FFF1DF95E931381E4" box="[136,262,1670,1697]" metricMagnitude="-2" metricUnit="m" metricValue="4.199999999999999" metricValueMax="4.6" metricValueMin="3.8" pageId="5" pageNumber="82" unit="cm" value="4.199999999999999" valueMax="4.6" valueMin="3.8">3.8–4.6 cm</quantity>
long, whitish to yellow speckled with pink; anthers
<quantity id="BE58DFA9FFE2FF8FFCC8F95E91AE81E4" box="[861,955,1670,1697]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="5" pageNumber="82" unit="mm" value="8.0" valueMax="9.0" valueMin="7.0">7–9 mm</quantity>
long, pale yellow; ovary 1.31.5 ×
<quantity id="BE58DFA9FFE2FF8FFAD3F95E97B281E4" box="[1350,1447,1670,1697]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="5" pageNumber="82" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 mm</quantity>
, ovoid to obovoid, glabrous, green with very pale white specks.
<emphasis id="4BD4AE5EFFE2FF8FFCCBF973918D8181" box="[862,920,1707,1732]" italics="true" pageId="5" pageNumber="82">Fruit</emphasis>
4.5–7 ×
<quantity id="BE58DFA9FFE2FF8FFB95F97296798180" box="[1024,1132,1706,1733]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="4.0" metricValueMin="2.5" pageId="5" pageNumber="82" unit="cm" value="3.25" valueMax="4.0" valueMin="2.5">2.5–4 cm</quantity>
, ellipsoid, terminating in an elongated semi-acute tip on both ends, glaucous, light green to yellowish green when mature with slightly mottled white variegation, inedible; mesocarp white; arils dark grey.
<emphasis id="4BD4AE5EFFE2FF8FFCBAF92A917A8049" box="[815,879,1778,1804]" italics="true" pageId="5" pageNumber="82">Seeds</emphasis>
black [not measured].
</paragraph>
<paragraph id="791F724CFFE2FF8FFF28F8CE92CC8011" blockId="5.[136,1452,914,2093]" pageId="5" pageNumber="82">
<emphasis id="4BD4AE5EFFE2FF8FFF28F8CE93568075" bold="true" box="[189,323,1814,1840]" pageId="5" pageNumber="82">Phenology:</emphasis>
—
<taxonomicName id="BEA009CFFFE2FF8FFECAF8CE907C8075" authorityName="Molinari, Kuethe &amp; Bernal" authorityYear="2024" box="[351,617,1814,1840]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="82" phylum="Tracheophyta" rank="species" species="rogitamensis">
<emphasis id="4BD4AE5EFFE2FF8FFECAF8CE907C8075" box="[351,617,1814,1840]" italics="true" pageId="5" pageNumber="82">Passiflora rogitamensis</emphasis>
</taxonomicName>
was found in flower in December and January and with fruit in January to March.
</paragraph>
</subSubSection>
<subSubSection id="31BA21C7FFE2FF8FFF28F88696E680D8" pageId="5" pageNumber="82" type="etymology">
<paragraph id="791F724CFFE2FF8FFF28F88696E680D8" blockId="5.[136,1452,914,2093]" pageId="5" pageNumber="82">
<emphasis id="4BD4AE5EFFE2FF8FFF28F886935D803D" bold="true" box="[189,328,1886,1912]" pageId="5" pageNumber="82">Etymology:</emphasis>
—
<taxonomicName id="BEA009CFFFE2FF8FFEF1F8869078803D" authorityName="Molinari, Kuethe &amp; Bernal" authorityYear="2024" box="[356,621,1886,1912]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="82" phylum="Tracheophyta" rank="species" species="rogitamensis">
<emphasis id="4BD4AE5EFFE2FF8FFEF1F8869078803D" box="[356,621,1886,1912]" italics="true" pageId="5" pageNumber="82">Passiflora rogitamensis</emphasis>
</taxonomicName>
is named after the Rogitama Biological Reserve in
<collectingCountry id="01B732DCFFE2FF8FFB30F8869702803D" box="[1189,1303,1886,1912]" name="Colombia" pageId="5" pageNumber="82">Colombia</collectingCountry>
. This was the location where it was first collected by Miguel Molinari who introduced the species under this name.
</paragraph>
</subSubSection>
<subSubSection id="31BA21C7FFE2FF8CFF28F87E96C280D9" lastPageId="6" lastPageNumber="83" pageId="5" pageNumber="82" type="distribution">
<paragraph id="791F724CFFE2FF8CFF28F87E96AD8799" blockId="5.[136,1452,914,2093]" lastBlockId="6.[136,1452,158,1949]" lastPageId="6" lastPageNumber="83" pageId="5" pageNumber="82">
<emphasis id="4BD4AE5EFFE2FF8FFF28F87E934C8085" bold="true" box="[189,345,1958,1984]" pageId="5" pageNumber="82">Distribution:</emphasis>
—
<taxonomicName id="BEA009CFFFE2FF8FFEE0F87E906B8085" authorityName="Molinari, Kuethe &amp; Bernal" authorityYear="2024" box="[373,638,1958,1984]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="82" phylum="Tracheophyta" rank="species" species="rogitamensis">
<emphasis id="4BD4AE5EFFE2FF8FFEE0F87E906B8085" box="[373,638,1958,1984]" italics="true" pageId="5" pageNumber="82">Passiflora rogitamensis</emphasis>
</taxonomicName>
is known only from the Departments of
<collectingRegion id="BB64BCAEFFE2FF8FFBD5F87F96818084" box="[1088,1172,1959,1985]" country="Colombia" name="Boyaca" pageId="5" pageNumber="82">Boyaca</collectingRegion>
and
<collectingRegion id="BB64BCAEFFE2FF8FFB5CF87E972E8085" box="[1225,1339,1958,1984]" country="Colombia" name="Santander" pageId="5" pageNumber="82">Santander</collectingRegion>
, northeast
<collectingCountry id="01B732DCFFE2FF8FFF1DF81292EC80A1" box="[136,249,1994,2020]" name="Colombia" pageId="5" pageNumber="82">Colombia</collectingCountry>
, where it was found near to the cities of Arcabuco and Tunja (
<figureCitation id="E19B6EC9FFE2FF8FFC29F81291EB80A1" box="[956,1022,1994,2020]" captionStart="FIGURE 1" captionStartId="1.[136,234,967,989]" captionTargetBox="[151,1436,187,940]" captionTargetId="figure-19@1.[151,1436,187,940]" captionTargetPageId="1" captionText="FIGURE 1. Map showing the allopatric distribution of the concerning new species (diamonds) and their nearest relatives (circles). Passiflora azulitensis (white); P. cocuyensis (red); P. rogitamensis (purple); Passiflora gritensis (green); P. semiciliosa (yellow); and P. longipes (blue). Map generated using both herbarium coordinates and georeferenced sightings (fieldwork supplemented with iNaturalist observations)." figureDoi="http://doi.org/10.5281/zenodo.14515923" httpUri="https://zenodo.org/record/14515923/files/figure.png" pageId="5" pageNumber="82">Fig. 1</figureCitation>
). It is commonly sighted in the dense cloud forest reserve of its namesake Reserva Rogitama Biodiversidad in
<collectingRegion id="BB64BCAEFFE2FF8FFC7CF83796DF8F4D" box="[1001,1226,2030,2057]" country="Colombia" name="Boyaca" pageId="5" pageNumber="82">Boyaca department</collectingRegion>
, where it grows at elevations between 2000 and
<quantity id="BE58DFA9FFE2FF8FFE46F7CA903D8F68" box="[467,552,2066,2093]" metricMagnitude="3" metricUnit="m" metricValue="2.75" pageId="5" pageNumber="82" unit="m" value="2750.0">2750 m</quantity>
above sea level. Another notable population was seen in the Paramo de Iguaque near to the city of Tunja. It grows as a moderate to large, tendrillate vine in the undergrowth of dense rain and cloud forest. It was reported by the collectors that this species rarely (if ever) grows in direct sunlight.
</paragraph>
<paragraph id="791F724CFFE1FF8CFF28FF3E909986D4" blockId="6.[136,1452,158,1949]" pageId="6" pageNumber="83">
<emphasis id="4BD4AE5EFFE1FF8CFF28FF3E93728645" bold="true" box="[189,359,230,256]" pageId="6" pageNumber="83">Conservation:</emphasis>
—There have been several occurrences of
<taxonomicName id="BEA009CFFFE1FF8CFCCBFF3E96788645" authorityName="Molinari, Kuethe &amp; Bernal" authorityYear="2024" box="[862,1133,230,256]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="6" pageNumber="83" phylum="Tracheophyta" rank="species" species="rogitamensis">
<emphasis id="4BD4AE5EFFE1FF8CFCCBFF3E96788645" box="[862,1133,230,256]" italics="true" pageId="6" pageNumber="83">Passiflora rogitamensis</emphasis>
</taxonomicName>
reported on public science websites such as iNaturalist or Flickr, indicating a moderate to high abundance of this species within its natural habitat. The largest population of
<taxonomicName id="BEA009CFFFE1FF8CFE3FFEF79043860D" authorityName="Molinari, Kuethe &amp; Bernal" authorityYear="2024" box="[426,598,303,328]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="6" pageNumber="83" phylum="Tracheophyta" rank="species" species="rogitamensis">
<emphasis id="4BD4AE5EFFE1FF8CFE3FFEF79043860D" box="[426,598,303,328]" italics="true" pageId="6" pageNumber="83">P. rogitamensis</emphasis>
</taxonomicName>
is seen within the Rogitama Biological Reserve, which is a protected forest reserve that should help protect this species’ longterm survival. AOO is
<quantity id="BE58DFA9FFE1FF8CFC39FE8A961A8629" box="[940,1039,338,365]" metricMagnitude="6" metricUnit="m" metricValue="3.15" pageId="6" pageNumber="83" unit="km" value="3150.0">3150 km</quantity>
<superScript id="8ED5DF04FFE1FF8CFB9BFE8A96038625" attach="left" box="[1038,1046,338,352]" fontSize="6" pageId="6" pageNumber="83">2</superScript>
, EOO is
<quantity id="BE58DFA9FFE1FF8CFBEAFE8A96D08629" box="[1151,1221,338,364]" metricMagnitude="4" metricUnit="m" metricValue="2.4" pageId="6" pageNumber="83" unit="km" value="24.0">24 km</quantity>
<superScript id="8ED5DF04FFE1FF8CFB50FE8A96D88625" attach="left" box="[1221,1229,338,352]" fontSize="6" pageId="6" pageNumber="83">2</superScript>
, scaling the species as Endangered (EN) criteria B1, B2civ and D.
</paragraph>
<paragraph id="791F724CFFE1FF8CFF28FE42967C8405" blockId="6.[136,1452,158,1949]" pageId="6" pageNumber="83">
<materialsCitation id="C9C87811FFE1FF8CFF28FE42937E869D" collectingDate="2004-10-16" collectionCode="JBB" collectorName="Lizcano" country="Colombia" elevation="2450" location="Reserva Rogitama" municipality="Arcabuco" pageId="6" pageNumber="83" specimenCount="1" stateProvince="Boyaca" typeStatus="paratype">
<emphasis id="4BD4AE5EFFE1FF8CFF28FE42932A86F1" bold="true" box="[189,319,410,436]" pageId="6" pageNumber="83">
<typeStatus id="A61BCCEEFFE1FF8CFF28FE42932C86F1" box="[189,313,410,436]" pageId="6" pageNumber="83" type="paratype">Paratypes</typeStatus>
:
</emphasis>
—
<collectingCountry id="01B732DCFFE1FF8CFEC9FE4293E186F1" box="[348,500,410,436]" name="Colombia" pageId="6" pageNumber="83">COLOMBIA</collectingCountry>
.
<collectingRegion id="BB64BCAEFFE1FF8CFD91FE42904986F1" box="[516,604,410,436]" country="Colombia" name="Boyaca" pageId="6" pageNumber="83">Boyacá</collectingRegion>
.
<collectingMunicipality id="997BE836FFE1FF8CFDFFFE4290CF86F1" box="[618,730,410,436]" pageId="6" pageNumber="83">Arcabuco</collectingMunicipality>
,
<location id="7C7F2497FFE1FF8CFD7EFE4391D786F1" LSID="urn:lsid:plazi:treatment:F109C35AFFE2FF8CFF1DFD9696C280D9:7C7F2497FFE1FF8CFD7EFE4391D786F1" box="[747,962,410,437]" country="Colombia" municipality="Arcabuco" name="Reserva Rogitama" pageId="6" pageNumber="83" stateProvince="Boyaca">Reserva Rogitama</location>
,
<quantity id="BE58DFA9FFE1FF8CFC47FE42963A86F0" box="[978,1071,410,437]" metricMagnitude="3" metricUnit="m" metricValue="2.45" pageId="6" pageNumber="83" unit="m" value="2450.0">
<elevation id="F28D957FFFE1FF8CFC47FE42963A86F0" box="[978,1071,410,437]" metricMagnitude="3" metricUnit="m" metricValue="2.45" pageId="6" pageNumber="83" unit="m" value="2450.0">2450 m</elevation>
</quantity>
,
<date id="0D1E548CFFE1FF8CFBD4FE42971986F1" box="[1089,1292,410,436]" pageId="6" pageNumber="83" value="2004-10-16">
<collectingDate id="1D5AAD64FFE1FF8CFBD4FE42971986F1" box="[1089,1292,410,436]" pageId="6" pageNumber="83" value="2004-10-16">16 October 2004</collectingDate>
</date>
,
<emphasis id="4BD4AE5EFFE1FF8CFA88FE43930A869D" italics="true" pageId="6" pageNumber="83">
G. Morales-
<collectorName id="D455179AFFE1FF8CFF1DFE6792F4869D" box="[136,225,447,472]" pageId="6" pageNumber="83">Lizcano</collectorName>
6369
</emphasis>
(
<collectionCode id="1FB1EA89FFE1FF8CFEBBFE669376869D" box="[302,355,446,472]" country="Colombia" name="Jardín Botánico José Celestino Mutis" pageId="6" pageNumber="83" type="Herbarium">JBB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C9C87811FFE1FF8CFEE3FE66965886B9" collectingDate="1996-05-11" collectionCode="COL" collectorName="J. L. Fernandez-Alonso" country="Colombia" county="Rogitama Reserve" elevation="2300" latitude="5.782222" location="Vereda Penas Blancas" longLatPrecision="21" longitude="-73.44361" municipality="Arcabuco" pageId="6" pageNumber="83" specimenCount="1" typeStatus="paratype">
<collectingCounty id="907E0AC0FFE1FF8CFEE3FE669056869C" box="[374,579,446,473]" pageId="6" pageNumber="83">Rogitama Reserve</collectingCounty>
near to
<collectingMunicipality id="997BE836FFE1FF8CFD0FFE66911F869D" box="[666,778,446,472]" pageId="6" pageNumber="83">Arcabuco</collectingMunicipality>
,
<location id="7C7F2497FFE1FF8CFC86FE66961F869D" LSID="urn:lsid:plazi:treatment:F109C35AFFE2FF8CFF1DFD9696C280D9:7C7F2497FFE1FF8CFC86FE66961F869D" box="[787,1034,446,473]" country="Colombia" county="Rogitama Reserve" latitude="5.782222" longLatPrecision="21" longitude="-73.44361" municipality="Arcabuco" name="Vereda Penas Blancas" pageId="6" pageNumber="83">Vereda Peñas Blancas</location>
,
<location id="7C7F2497FFE1FF8CFB80FE669707869D" LSID="urn:lsid:plazi:treatment:F109C35AFFE2FF8CFF1DFD9696C280D9:7C7F2497FFE1FF8CFB80FE669707869D" box="[1045,1298,446,473]" country="Colombia" county="Rogitama Reserve" latitude="5.782222" longLatPrecision="21" longitude="-73.44361" municipality="Arcabuco" name="Hacienda Las Delicias" pageId="6" pageNumber="83">Hacienda Las Delicias</location>
,
<geoCoordinate id="1C94148BFFE1FF8CFA89FE1897B3869E" box="[1308,1446,448,475]" degrees="05" direction="north" minutes="46" orientation="latitude" pageId="6" pageNumber="83" precision="15" seconds="56" value="5.782222">05°46’56″N</geoCoordinate>
,
<geoCoordinate id="1C94148BFFE1FF8CFF1DFE3C930386BA" box="[136,278,484,511]" degrees="73" direction="west" minutes="26" orientation="longitude" pageId="6" pageNumber="83" precision="15" seconds="37" value="-73.44361">73°26’37″W</geoCoordinate>
,
<quantity id="BE58DFA9FFE1FF8CFEB1FE3A939686B9" box="[292,387,482,508]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="6" pageNumber="83" unit="m" value="2300.0">
<elevation id="F28D957FFFE1FF8CFEB1FE3A939686B9" box="[292,387,482,508]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="6" pageNumber="83" unit="m" value="2300.0">2300 m</elevation>
</quantity>
.a.s.l.,
<date id="0D1E548CFFE1FF8CFE52FE3A907686B9" box="[455,611,482,509]" pageId="6" pageNumber="83" value="1996-05-11">
<collectingDate id="1D5AAD64FFE1FF8CFE52FE3A907686B9" box="[455,611,482,509]" pageId="6" pageNumber="83" value="1996-05-11">11 May 1996</collectingDate>
</date>
, (fl.)
<emphasis id="4BD4AE5EFFE1FF8CFD30FE3B91E386B9" box="[677,1014,482,508]" italics="true" pageId="6" pageNumber="83">
<collectorName id="D455179AFFE1FF8CFD30FE3B91B286B9" box="[677,935,482,508]" pageId="6" pageNumber="83">J.L. Fernández-Alonso</collectorName>
14097
</emphasis>
(
<collectionCode id="1FB1EA89FFE1FF8CFB9DFE3A965386B9" box="[1032,1094,482,508]" country="Colombia" lsid="urn:lsid:biocol.org:col:15629" name="Universidad Nacional de Colombia" pageId="6" pageNumber="83" type="Herbarium">COL</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C9C87811FFE1FF8CFBCFFE3A962D8565" collectingDate="1980-05-22" collectionCode="COL" collectorName="J. E. Valencia" country="Colombia" county="Arcabuco" elevation="2300" location="Bucaramanga." municipality="Via" pageId="6" pageNumber="83" specimenCount="1" typeStatus="paratype">
<collectingCounty id="907E0AC0FFE1FF8CFBCFFE3A96DF86B9" box="[1114,1226,482,508]" pageId="6" pageNumber="83">Arcabuco</collectingCounty>
,
<collectingMunicipality id="997BE836FFE1FF8CFB43FE3A96E886B9" box="[1238,1277,482,508]" pageId="6" pageNumber="83">Via</collectingMunicipality>
que conduce a
<location id="7C7F2497FFE1FF8CFF1DFDDF933F8564" LSID="urn:lsid:plazi:treatment:F109C35AFFE2FF8CFF1DFD9696C280D9:7C7F2497FFE1FF8CFF1DFDDF933F8564" box="[136,298,519,545]" country="Colombia" county="Arcabuco" municipality="Via" name="Bucaramanga." pageId="6" pageNumber="83">Bucaramanga.</location>
“La Cumbre”,
<quantity id="BE58DFA9FFE1FF8CFE4BFDDE902D8564" box="[478,568,518,545]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="6" pageNumber="83" unit="m" value="2300.0">
<elevation id="F28D957FFFE1FF8CFE4BFDDE902D8564" box="[478,568,518,545]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="6" pageNumber="83" unit="m" value="2300.0">2300 m</elevation>
</quantity>
,
<date id="0D1E548CFFE1FF8CFDD3FDDE90F18565" box="[582,740,518,545]" pageId="6" pageNumber="83" value="1980-05-22">
<collectingDate id="1D5AAD64FFE1FF8CFDD3FDDE90F18565" box="[582,740,518,545]" pageId="6" pageNumber="83" value="1980-05-22">22 May 1980</collectingDate>
</date>
, (fl.)
<emphasis id="4BD4AE5EFFE1FF8CFCB2FDDF91F58565" box="[807,992,518,544]" italics="true" pageId="6" pageNumber="83">
<collectorName id="D455179AFFE1FF8CFCB2FDDF91A98565" box="[807,956,518,544]" pageId="6" pageNumber="83">J.E. Valencia</collectorName>
11
</emphasis>
(
<collectionCode id="1FB1EA89FFE1FF8CFC67FDDE96258565" box="[1010,1072,518,544]" country="Colombia" lsid="urn:lsid:biocol.org:col:15629" name="Universidad Nacional de Colombia" pageId="6" pageNumber="83" type="Herbarium">COL</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C9C87811FFE1FF8CFBD3FDDE967E8501" collectingDate="1996-05-14" collectionCode="COL" collectorName="de Arcabuco &amp; J. L. Fernandez-Alonso" country="Colombia" county="Arcabuco" elevation="2600" location="Via Arcabuco-Moniquira" municipality="Via" pageId="6" pageNumber="83" specimenCount="1" typeStatus="paratype">
<emphasis id="4BD4AE5EFFE1FF8CFBD3FDDE96678565" box="[1094,1138,518,544]" italics="true" pageId="6" pageNumber="83">ibid</emphasis>
.,
<location id="7C7F2497FFE1FF8CFB1CFDDE97B28565" LSID="urn:lsid:plazi:treatment:F109C35AFFE2FF8CFF1DFD9696C280D9:7C7F2497FFE1FF8CFB1CFDDE97B28565" box="[1161,1447,518,544]" country="Colombia" name="Via Arcabuco-Moniquira" pageId="6" pageNumber="83">Via Arcabuco-Moniquira</location>
, cañon
<collectorName id="D455179AFFE1FF8CFF41FDF293728501" box="[212,359,554,580]" pageId="6" pageNumber="83">de Arcabuco</collectorName>
,
<quantity id="BE58DFA9FFE1FF8CFEE1FDF293C68501" box="[372,467,554,580]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="6" pageNumber="83" unit="m" value="2600.0">
<elevation id="F28D957FFFE1FF8CFEE1FDF293C68501" box="[372,467,554,580]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="6" pageNumber="83" unit="m" value="2600.0">2600 m</elevation>
</quantity>
.a.s.l.,
<date id="0D1E548CFFE1FF8CFD82FDF290A18501" box="[535,692,554,581]" pageId="6" pageNumber="83" value="1996-05-14">
<collectingDate id="1D5AAD64FFE1FF8CFD82FDF290A18501" box="[535,692,554,581]" pageId="6" pageNumber="83" value="1996-05-14">14 May 1996</collectingDate>
</date>
,
<emphasis id="4BD4AE5EFFE1FF8CFD57FDF396018501" box="[706,1044,554,580]" italics="true" pageId="6" pageNumber="83">
<collectorName id="D455179AFFE1FF8CFD57FDF391D18501" box="[706,964,554,580]" pageId="6" pageNumber="83">J.L. Fernández-Alonso</collectorName>
14235
</emphasis>
(
<collectionCode id="1FB1EA89FFE1FF8CFBB0FDF296768501" box="[1061,1123,554,580]" country="Colombia" lsid="urn:lsid:biocol.org:col:15629" name="Universidad Nacional de Colombia" pageId="6" pageNumber="83" type="Herbarium">COL</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C9C87811FFE1FF8CFBECFDF2903B85C9" collectingDate="1994-11-21" collectionCode="COL, MA" collectorName="J. L. Fernandez-Alonso" country="Colombia" county="Duitama" elevation="1900" latitude="5.9661107" location="El Carmen" longLatPrecision="21" longitude="-73.16583" municipality="Trayecto" pageId="6" pageNumber="83" specimenCount="1" typeStatus="paratype">
<collectingCounty id="907E0AC0FFE1FF8CFBECFDF296C98501" box="[1145,1244,554,580]" pageId="6" pageNumber="83">Duitama</collectingCounty>
,
<collectingMunicipality id="997BE836FFE1FF8CFB7FFDF397588500" box="[1258,1357,555,581]" pageId="6" pageNumber="83">Trayecto</collectingMunicipality>
entre la vereda
<location id="7C7F2497FFE1FF8CFF4FFD969341852D" LSID="urn:lsid:plazi:treatment:F109C35AFFE2FF8CFF1DFD9696C280D9:7C7F2497FFE1FF8CFF4FFD969341852D" box="[218,340,590,616]" country="Colombia" county="Duitama" latitude="5.9661107" longLatPrecision="21" longitude="-73.16583" municipality="Trayecto" name="El Carmen" pageId="6" pageNumber="83">El Carmen</location>
y
<location id="7C7F2497FFE1FF8CFEE5FD9693D7852D" LSID="urn:lsid:plazi:treatment:F109C35AFFE2FF8CFF1DFD9696C280D9:7C7F2497FFE1FF8CFEE5FD9693D7852D" box="[368,450,590,616]" country="Colombia" county="Duitama" latitude="5.9661107" longLatPrecision="21" longitude="-73.16583" municipality="Trayecto" name="Virolin" pageId="6" pageNumber="83">Virolín</location>
, páramo
<location id="7C7F2497FFE1FF8CFDBCFD979084852D" LSID="urn:lsid:plazi:treatment:F109C35AFFE2FF8CFF1DFD9696C280D9:7C7F2497FFE1FF8CFDBCFD979084852D" box="[553,657,590,617]" country="Colombia" county="Duitama" latitude="5.9661107" longLatPrecision="21" longitude="-73.16583" municipality="Trayecto" name="La Rusia" pageId="6" pageNumber="83">La Rusia</location>
,
<geoCoordinate id="1C94148BFFE1FF8CFD08FD889130852E" box="[669,805,592,619]" degrees="05" direction="north" minutes="57" orientation="latitude" pageId="6" pageNumber="83" precision="15" seconds="58" value="5.9661107">05°57′58″N</geoCoordinate>
,
<geoCoordinate id="1C94148BFFE1FF8CFCA4FD8891A9852E" box="[817,956,592,619]" degrees="73" direction="west" minutes="09" orientation="longitude" pageId="6" pageNumber="83" precision="15" seconds="57" value="-73.16583">73°09′57″W</geoCoordinate>
,
<quantity id="BE58DFA9FFE1FF8CFC5DFD969630852D" box="[968,1061,590,616]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="6" pageNumber="83" unit="m" value="1900.0">
<elevation id="F28D957FFFE1FF8CFC5DFD969630852D" box="[968,1061,590,616]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="6" pageNumber="83" unit="m" value="1900.0">1900 m</elevation>
</quantity>
.a.s.l.,
<date id="0D1E548CFFE1FF8CFBF2FD969756852D" box="[1127,1347,590,616]" pageId="6" pageNumber="83" value="1994-11-21">
<collectingDate id="1D5AAD64FFE1FF8CFBF2FD969756852D" box="[1127,1347,590,616]" pageId="6" pageNumber="83" value="1994-11-21">21 November 1994</collectingDate>
</date>
, (fl.)
<emphasis id="4BD4AE5EFFE1FF8CFA14FD9793B785C9" italics="true" pageId="6" pageNumber="83">
<collectorName id="D455179AFFE1FF8CFA14FD97934285C9" pageId="6" pageNumber="83">J.L. Fernández-Alonso</collectorName>
12097
</emphasis>
(
<collectionCode id="1FB1EA89FFE1FF8CFE3AFDAA93FE85C9" box="[431,491,626,652]" country="Colombia" lsid="urn:lsid:biocol.org:col:15629" name="Universidad Nacional de Colombia" pageId="6" pageNumber="83" type="Herbarium">COL</collectionCode>
,
<collectionCode id="1FB1EA89FFE1FF8CFE60FDAA903285C9" box="[501,551,626,652]" country="Spain" lsid="urn:lsid:biocol.org:col:15682" name="Real Jardín Botánico" pageId="6" pageNumber="83" type="Herbarium">MA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C9C87811FFE1FF8CFDADFDAA916585F5" collectingDate="2004-06-21" collectionCode="COL" collectorName="J. L. Fernandez-Alonso" country="Colombia" county="Floridablanca" elevation="1750" latitude="7.121667" location="El Doce" longLatPrecision="21" longitude="-73.07694" municipality="Tona" pageId="6" pageNumber="83" specimenCount="1" stateProvince="Santander" typeStatus="paratype">
<collectingRegion id="BB64BCAEFFE1FF8CFDADFDAA90BF85C9" box="[568,682,626,652]" country="Colombia" name="Santander" pageId="6" pageNumber="83">Santander</collectingRegion>
.
<collectingCounty id="907E0AC0FFE1FF8CFD26FDAA914585C9" box="[691,848,626,652]" pageId="6" pageNumber="83">Floridablanca</collectingCounty>
, between
<collectingMunicipality id="997BE836FFE1FF8CFC2EFDAB91E685C8" box="[955,1011,627,653]" pageId="6" pageNumber="83">Tona</collectingMunicipality>
and
<location id="7C7F2497FFE1FF8CFBB0FDAA969485C8" LSID="urn:lsid:plazi:treatment:F109C35AFFE2FF8CFF1DFD9696C280D9:7C7F2497FFE1FF8CFBB0FDAA969485C8" box="[1061,1153,626,653]" country="Colombia" county="Floridablanca" latitude="7.121667" longLatPrecision="21" longitude="-73.07694" municipality="Tona" name="El Doce" pageId="6" pageNumber="83" stateProvince="Santander">El Doce</location>
,
<geoCoordinate id="1C94148BFFE1FF8CFB1FFDAC970785CA" box="[1162,1298,628,655]" degrees="07" direction="north" minutes="07" orientation="latitude" pageId="6" pageNumber="83" precision="15" seconds="18" value="7.121667">07°07′18″N</geoCoordinate>
,
<geoCoordinate id="1C94148BFFE1FF8CFA8EFDAC97B285CA" box="[1307,1447,628,655]" degrees="73" direction="west" minutes="04" orientation="longitude" pageId="6" pageNumber="83" precision="15" seconds="37" value="-73.07694">73°04′37″W</geoCoordinate>
,
<quantity id="BE58DFA9FFE1FF8CFF1DFD4E92F085F5" box="[136,229,662,689]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="6" pageNumber="83" unit="m" value="1750.0">
<elevation id="F28D957FFFE1FF8CFF1DFD4E92F085F5" box="[136,229,662,689]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="6" pageNumber="83" unit="m" value="1750.0">1750 m</elevation>
</quantity>
.a.s.l.,
<date id="0D1E548CFFE1FF8CFEB2FD4E93D485F5" box="[295,449,662,689]" pageId="6" pageNumber="83" value="2004-06-21">
<collectingDate id="1D5AAD64FFE1FF8CFEB2FD4E93D485F5" box="[295,449,662,689]" pageId="6" pageNumber="83" value="2004-06-21">21 June 2004</collectingDate>
</date>
,
<emphasis id="4BD4AE5EFFE1FF8CFE59FD4F910E85F5" box="[460,795,662,688]" italics="true" pageId="6" pageNumber="83">
<collectorName id="D455179AFFE1FF8CFE59FD4F90D885F5" box="[460,717,662,688]" pageId="6" pageNumber="83">J.L. Fernández-Alonso</collectorName>
21527
</emphasis>
(
<collectionCode id="1FB1EA89FFE1FF8CFCBFFD4E917D85F5" box="[810,872,662,688]" country="Colombia" lsid="urn:lsid:biocol.org:col:15629" name="Universidad Nacional de Colombia" pageId="6" pageNumber="83" type="Herbarium">COL</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C9C87811FFE1FF8CFCE9FD4E96FF8591" collectingDate="1994-11-01" collectionCode="COL" collectorName="J. L. Fernandez-Alonso" country="Colombia" county="Charala" elevation="2300" latitude="5.963611" location="Virolin" longLatPrecision="21" longitude="-73.19417" municipality="Carretera El Carmen" pageId="6" pageNumber="83" specimenCount="1" typeStatus="paratype">
<collectingCounty id="907E0AC0FFE1FF8CFCE9FD4E91C385F5" box="[892,982,662,688]" pageId="6" pageNumber="83">Charalá</collectingCounty>
,
<collectingMunicipality id="997BE836FFE1FF8CFC77FD4E96DE85F5" box="[994,1227,662,688]" pageId="6" pageNumber="83">Carretera El Carmen</collectingMunicipality>
a
<location id="7C7F2497FFE1FF8CFB73FD4E972D85F5" LSID="urn:lsid:plazi:treatment:F109C35AFFE2FF8CFF1DFD9696C280D9:7C7F2497FFE1FF8CFB73FD4E972D85F5" box="[1254,1336,662,688]" country="Colombia" county="Charala" latitude="5.963611" longLatPrecision="21" longitude="-73.19417" municipality="Carretera El Carmen" name="Virolin" pageId="6" pageNumber="83">Virolin</location>
,
<location id="7C7F2497FFE1FF8CFAD1FD4E97B285F5" LSID="urn:lsid:plazi:treatment:F109C35AFFE2FF8CFF1DFD9696C280D9:7C7F2497FFE1FF8CFAD1FD4E97B285F5" box="[1348,1447,662,688]" country="Colombia" county="Charala" latitude="5.963611" longLatPrecision="21" longitude="-73.19417" municipality="Carretera El Carmen" name="Duitama" pageId="6" pageNumber="83">Duitama</location>
,
<geoCoordinate id="1C94148BFFE1FF8CFF1DFD6493058592" box="[136,272,700,727]" degrees="05" direction="north" minutes="57" orientation="latitude" pageId="6" pageNumber="83" precision="15" seconds="49" value="5.963611">05°57′49″N</geoCoordinate>
,
<geoCoordinate id="1C94148BFFE1FF8CFE8FFD6493B18592" box="[282,420,700,727]" degrees="73" direction="west" minutes="11" orientation="longitude" pageId="6" pageNumber="83" precision="15" seconds="39" value="-73.19417">73°11′39″W</geoCoordinate>
,
<quantity id="BE58DFA9FFE1FF8CFE3BFD62901D8591" box="[430,520,698,724]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="6" pageNumber="83" unit="m" value="2300.0">
<elevation id="F28D957FFFE1FF8CFE3BFD62901D8591" box="[430,520,698,724]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="6" pageNumber="83" unit="m" value="2300.0">2300 m</elevation>
</quantity>
.a.s.l.,
<date id="0D1E548CFFE1FF8CFDDCFD6291068591" box="[585,787,698,724]" pageId="6" pageNumber="83" value="1994-11-01">
<collectingDate id="1D5AAD64FFE1FF8CFDDCFD6291068591" box="[585,787,698,724]" pageId="6" pageNumber="83" value="1994-11-01">1 November 1994</collectingDate>
</date>
, (fl.)
<emphasis id="4BD4AE5EFFE1FF8CFCD8FD6396838591" box="[845,1174,698,724]" italics="true" pageId="6" pageNumber="83">
<collectorName id="D455179AFFE1FF8CFCD8FD63965E8591" box="[845,1099,698,724]" pageId="6" pageNumber="83">J.L. Fernández-Alonso</collectorName>
12286
</emphasis>
(
<collectionCode id="1FB1EA89FFE1FF8CFB31FD6296F78591" box="[1188,1250,698,724]" country="Colombia" lsid="urn:lsid:biocol.org:col:15629" name="Universidad Nacional de Colombia" pageId="6" pageNumber="83" type="Herbarium">COL</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C9C87811FFE1FF8CFB61FD62931A8459" collectingDate="1926-12-20" collectorName="E. Killip &amp; A. Smith" country="United States of America" county="Onzaga" elevation="2300" latitude="6.3166666" location="Las Vegas" longLatPrecision="1304" longitude="-72.73333" municipality="Paramo Las Vegas" pageId="6" pageNumber="83" specimenCount="1" typeStatus="paratype">
<collectingCounty id="907E0AC0FFE1FF8CFB61FD6297598591" box="[1268,1356,698,724]" pageId="6" pageNumber="83">Onzaga</collectingCounty>
,
<collectingMunicipality id="997BE836FFE1FF8CFAC3FD63931485BC" pageId="6" pageNumber="83">Paramo Las Vegas</collectingMunicipality>
, montanos de
<location id="7C7F2497FFE1FF8CFE39FD07903185BC" LSID="urn:lsid:plazi:treatment:F109C35AFFE2FF8CFF1DFD9696C280D9:7C7F2497FFE1FF8CFE39FD07903185BC" box="[428,548,735,761]" country="United States of America" county="Onzaga" latitude="6.3166666" longLatPrecision="1304" longitude="-72.73333" municipality="Paramo Las Vegas" name="Las Vegas" pageId="6" pageNumber="83">Las Vegas</location>
,
<geoCoordinate id="1C94148BFFE1FF8CFDA6FD06908385BD" box="[563,662,734,760]" degrees="06" direction="north" minutes="19" orientation="latitude" pageId="6" pageNumber="83" precision="925" value="6.3166666">06°19’N</geoCoordinate>
,
<geoCoordinate id="1C94148BFFE1FF8CFD30FD06910F85BD" box="[677,794,734,760]" degrees="072" direction="west" minutes="44" orientation="longitude" pageId="6" pageNumber="83" precision="925" value="-72.73333">072°44’W</geoCoordinate>
,
<quantity id="BE58DFA9FFE1FF8CFCBCFD06919C85BD" box="[809,905,734,760]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="6" pageNumber="83" unit="m" value="2300.0">
<elevation id="F28D957FFFE1FF8CFCBCFD06919C85BD" box="[809,905,734,760]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="6" pageNumber="83" unit="m" value="2300.0">2300 m</elevation>
</quantity>
.a.s.l.,
<date id="0D1E548CFFE1FF8CFC5BFD0696B885BD" box="[974,1197,734,760]" pageId="6" pageNumber="83" value="1926-12-20">
<collectingDate id="1D5AAD64FFE1FF8CFC5BFD0696B885BD" box="[974,1197,734,760]" pageId="6" pageNumber="83" value="1926-12-20">20 December 1926</collectingDate>
</date>
,
<emphasis id="4BD4AE5EFFE1FF8CFB29FD0792D98459" italics="true" pageId="6" pageNumber="83">
<collectorName id="D455179AFFE1FF8CFB29FD07970A85BD" box="[1212,1311,734,760]" pageId="6" pageNumber="83">E. Killip</collectorName>
&amp;
<collectorName id="D455179AFFE1FF8CFADCFD0797B985BD" box="[1353,1452,734,761]" pageId="6" pageNumber="83">A. Smith</collectorName>
15911
</emphasis>
(
<collectingCountry id="01B732DCFFE1FF8CFF4BFCDA93138459" box="[222,262,770,796]" name="United States of America" pageId="6" pageNumber="83">US</collectingCountry>
)
</materialsCitation>
;
<materialsCitation id="C9C87811FFE1FF8CFE89FCDA96708405" collectingDate="1997-11-13" collectionCode="COL" collectorName="J. J. Cadena" country="United States of America" county="Charala" elevation="2450" latitude="6.0066667" location="Sierra" longLatPrecision="21" longitude="-73.148056" municipality="Vereda Santa Helena" pageId="6" pageNumber="83" specimenCount="1" typeStatus="paratype">
<collectingCounty id="907E0AC0FFE1FF8CFE89FCDA93638459" box="[284,374,770,796]" pageId="6" pageNumber="83">Charalá</collectingCounty>
,
<collectingMunicipality id="997BE836FFE1FF8CFE16FCDA90668459" box="[387,627,770,796]" pageId="6" pageNumber="83">Vereda Santa Helena</collectingMunicipality>
, predio la
<location id="7C7F2497FFE1FF8CFD7BFCDA91218459" LSID="urn:lsid:plazi:treatment:F109C35AFFE2FF8CFF1DFD9696C280D9:7C7F2497FFE1FF8CFD7BFCDA91218459" box="[750,820,770,796]" country="United States of America" county="Charala" latitude="6.0066667" longLatPrecision="21" longitude="-73.148056" municipality="Vereda Santa Helena" name="Sierra" pageId="6" pageNumber="83">Sierra</location>
,
<location id="7C7F2497FFE1FF8CFCD4FCDA96098458" LSID="urn:lsid:plazi:treatment:F109C35AFFE2FF8CFF1DFD9696C280D9:7C7F2497FFE1FF8CFCD4FCDA96098458" box="[833,1052,770,797]" country="United States of America" county="Charala" latitude="6.0066667" longLatPrecision="21" longitude="-73.148056" municipality="Vereda Santa Helena" name="Santuario de Fauna" pageId="6" pageNumber="83">Santuario de Fauna</location>
y Flora Guanentá Alto
<location id="7C7F2497FFE1FF8CFABBFCDA97B28458" LSID="urn:lsid:plazi:treatment:F109C35AFFE2FF8CFF1DFD9696C280D9:7C7F2497FFE1FF8CFABBFCDA97B28458" box="[1326,1447,770,797]" country="United States of America" county="Charala" latitude="6.0066667" longLatPrecision="21" longitude="-73.148056" municipality="Vereda Santa Helena" name="Rio Fonce" pageId="6" pageNumber="83">Río Fonce</location>
,
<geoCoordinate id="1C94148BFFE1FF8CFF1DFCF093058406" box="[136,272,808,835]" degrees="06" direction="north" minutes="00" orientation="latitude" pageId="6" pageNumber="83" precision="15" seconds="24" value="6.0066667">06°00′24″N</geoCoordinate>
,
<geoCoordinate id="1C94148BFFE1FF8CFE8EFCF093B28406" box="[283,423,808,835]" degrees="73" direction="west" minutes="08" orientation="longitude" pageId="6" pageNumber="83" precision="15" seconds="53" value="-73.148056">73°08′53″W</geoCoordinate>
,
<quantity id="BE58DFA9FFE1FF8CFE27FCFE901B8405" box="[434,526,806,833]" metricMagnitude="3" metricUnit="m" metricValue="2.45" pageId="6" pageNumber="83" unit="m" value="2450.0">
<elevation id="F28D957FFFE1FF8CFE27FCFE901B8405" box="[434,526,806,833]" metricMagnitude="3" metricUnit="m" metricValue="2.45" pageId="6" pageNumber="83" unit="m" value="2450.0">2450 m</elevation>
</quantity>
.a.s.l.,
<date id="0D1E548CFFE1FF8CFDC5FCFE913E8405" box="[592,811,806,832]" pageId="6" pageNumber="83" value="1997-11-13">
<collectingDate id="1D5AAD64FFE1FF8CFDC5FCFE913E8405" box="[592,811,806,832]" pageId="6" pageNumber="83" value="1997-11-13">13 November 1997</collectingDate>
</date>
, (fl.)
<emphasis id="4BD4AE5EFFE1FF8CFCFDFCFF96058405" box="[872,1040,806,832]" italics="true" pageId="6" pageNumber="83">
<collectorName id="D455179AFFE1FF8CFCFDFCFF91F88405" box="[872,1005,806,832]" pageId="6" pageNumber="83">J.J. Cadena</collectorName>
83
</emphasis>
(
<collectionCode id="1FB1EA89FFE1FF8CFB8AFCFE96488405" box="[1055,1117,806,832]" country="Colombia" lsid="urn:lsid:biocol.org:col:15629" name="Universidad Nacional de Colombia" pageId="6" pageNumber="83" type="Herbarium">COL</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="791F724CFFE1FF8CFF28FC92966084CD" blockId="6.[136,1452,158,1949]" pageId="6" pageNumber="83">
<emphasis id="4BD4AE5EFFE1FF8CFF28FC9293DD8421" bold="true" box="[189,456,842,868]" pageId="6" pageNumber="83">Cultivated collections:</emphasis>
—Bagsvaerd Collection (
<collectingCountry id="01B732DCFFE1FF8CFD48FC92915E8421" box="[733,843,842,868]" name="Denmark" pageId="6" pageNumber="83">Denmark</collectingCountry>
),
<emphasis id="4BD4AE5EFFE1FF8CFCCBFC93961A8421" box="[862,1039,842,868]" italics="true" pageId="6" pageNumber="83">F.R. Stausholm;</emphasis>
Jardin Botanico
<collectingRegion id="BB64BCAEFFE1FF8CFB59FC9397088420" box="[1228,1309,843,869]" country="Colombia" name="Bogota" pageId="6" pageNumber="83">Bogota</collectingRegion>
(
<collectingCountry id="01B732DCFFE1FF8CFABEFC92978A8421" box="[1323,1439,842,868]" name="Colombia" pageId="6" pageNumber="83">Colombia</collectingCountry>
),
<emphasis id="4BD4AE5EFFE1FF8CFF1DFCB7930484CD" box="[136,273,878,905]" italics="true" pageId="6" pageNumber="83">G. Morales;</emphasis>
Passiflorahoeve (
<collectingCountry id="01B732DCFFE1FF8CFE4DFCB6907384CD" box="[472,614,878,904]" name="Netherlands" pageId="6" pageNumber="83">Netherlands</collectingCountry>
),
<emphasis id="4BD4AE5EFFE1FF8CFDEDFCB7911484CD" box="[632,769,879,905]" italics="true" pageId="6" pageNumber="83">A. Lommen;</emphasis>
RBG Kew (
<collectingCountry id="01B732DCFFE1FF8CFC18FCB691AF84CD" box="[909,954,878,904]" name="United Kingdom" pageId="6" pageNumber="83">UK</collectingCountry>
),
<emphasis id="4BD4AE5EFFE1FF8CFC5BFCB7966084CD" box="[974,1141,878,905]" italics="true" pageId="6" pageNumber="83">C. Magdalena.</emphasis>
</paragraph>
<paragraph id="791F724CFFE1FF8CFF28FC4A910B82C5" blockId="6.[136,1452,158,1949]" pageId="6" pageNumber="83">
<emphasis id="4BD4AE5EFFE1FF8CFF28FC4A931C84E9" bold="true" box="[189,265,914,940]" pageId="6" pageNumber="83">Notes:</emphasis>
—
<taxonomicName id="BEA009CFFFE1FF8CFEB3FC4A902684E9" authorityName="Molinari, Kuethe &amp; Bernal" authorityYear="2024" box="[294,563,914,940]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="6" pageNumber="83" phylum="Tracheophyta" rank="species" species="rogitamensis">
<emphasis id="4BD4AE5EFFE1FF8CFEB3FC4A902684E9" box="[294,563,914,940]" italics="true" pageId="6" pageNumber="83">Passiflora rogitamensis</emphasis>
</taxonomicName>
belongs to
<taxonomicName id="BEA009CFFFE1FF8CFD2AFC4A91E084E9" box="[703,1013,914,940]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="6" pageNumber="83" phylum="Tracheophyta" rank="subGenus" subGenus="Passiflora">
<emphasis id="4BD4AE5EFFE1FF8CFD2AFC4A912784E9" box="[703,818,914,940]" italics="true" pageId="6" pageNumber="83">Passiflora</emphasis>
subg.
<emphasis id="4BD4AE5EFFE1FF8CFC17FC4A91E084E9" box="[898,1013,914,940]" italics="true" pageId="6" pageNumber="83">Passiflora</emphasis>
</taxonomicName>
supersect.
<taxonomicName id="BEA009CFFFE1FF8CFBECFC4A90468495" authority="(Cervi) Feuillet &amp; MacDougal (2003: 38)" authorityName="Feuillet &amp; MacDougal" authorityPageNumber="38" authorityYear="2003" baseAuthorityName="Cervi" genus="Stipulata" pageId="6" pageNumber="83" rank="section" section="Kermesinae">
<emphasis id="4BD4AE5EFFE1FF8CFBECFC4A96CB84E9" box="[1145,1246,914,940]" italics="true" pageId="6" pageNumber="83">Stipulata</emphasis>
sect.
<emphasis id="4BD4AE5EFFE1FF8CFAB3FC4B97B984E9" box="[1318,1452,915,940]" italics="true" pageId="6" pageNumber="83">Kermesinae</emphasis>
(Cervi)
<bibRefCitation id="1D310FBDFFE1FF8CFF4AFC6E90468495" author="Feuillet, C. &amp; MacDougal, J. M." box="[223,595,950,976]" pageId="6" pageNumber="83" pagination="34 - 38" refId="ref8477" refString="Feuillet, C. &amp; MacDougal, J. M. (2003) A new infrageneric classification of Passiflora L. (Passifloraceae). Passiflora 13: 34 - 38. https: // doi. org / 10.2307 / 3393379" type="journal article" year="2003">Feuillet &amp; MacDougal (2003: 38)</bibRefCitation>
</taxonomicName>
for having elongated, often pendent peduncles (&gt;
<quantity id="BE58DFA9FFE1FF8CFBE9FC6E96A38495" box="[1148,1206,950,977]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="83" unit="cm" value="5.0">5 cm</quantity>
), 4–6 stipitate petiolar nectaries and flowers with complex coronal structures containing more than 4 series of filaments. Among the several species with a
<emphasis id="4BD4AE5EFFE1FF8CFEA7FC2793C7835D" box="[306,466,1022,1048]" italics="true" pageId="6" pageNumber="83">nomen nudum</emphasis>
,
<taxonomicName id="BEA009CFFFE1FF8CFE77FC27909A835D" authorityName="Molinari, Kuethe &amp; Bernal" authorityYear="2024" box="[482,655,1023,1048]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="6" pageNumber="83" phylum="Tracheophyta" rank="species" species="rogitamensis">
<emphasis id="4BD4AE5EFFE1FF8CFE77FC27909A835D" box="[482,655,1023,1048]" italics="true" pageId="6" pageNumber="83">P. rogitamensis</emphasis>
</taxonomicName>
appears to be the most used in [sub-]scientific databases (iNaturalist, biodiversity4all, minambiente.gov.co) and floral surveys (e.g.,
<bibRefCitation id="1D310FBDFFE1FF8CFCAFFBFA91CF8379" author="Ocampo, J." box="[826,986,1058,1084]" pageId="6" pageNumber="83" refId="ref9133" refString="Ocampo, J. (2018) Passionflowers of Colombia. National University of Colombia - UNAL, Poster. Available from: https: // www. researchgate. net / publication / 326366326 _ Passionflowers _ of _ Colombia (Accessed 3 September 2023)" type="url" year="2018">Ocampo 2018</bibRefCitation>
).Although, hypothetically, the application of such a name that is not validly published is unfounded, and therefore the name should not be used in any scientific writing.
<taxonomicName id="BEA009CFFFE1FF8CFF7CFBB293E083C1" authorityName="Molinari, Kuethe &amp; Bernal" authorityYear="2024" box="[233,501,1130,1156]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="6" pageNumber="83" phylum="Tracheophyta" rank="species" species="rogitamensis">
<emphasis id="4BD4AE5EFFE1FF8CFF7CFBB293E083C1" box="[233,501,1130,1156]" italics="true" pageId="6" pageNumber="83">Passiflora rogitamensis</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="BEA009CFFFE1FF8CFD5AFBB3962483C1" authority="Jussieu (1805: 111)" authorityName="Jussieu" authorityPageNumber="111" authorityYear="1805" box="[719,1073,1130,1157]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="6" pageNumber="83" phylum="Tracheophyta" rank="species" species="longipes">
<emphasis id="4BD4AE5EFFE1FF8CFD5AFBB3915E83C1" box="[719,843,1130,1156]" italics="true" pageId="6" pageNumber="83">P. longipes</emphasis>
<bibRefCitation id="1D310FBDFFE1FF8CFCC1FBB2962483C1" author="Jussieu, A. L. de" box="[852,1073,1130,1157]" pageId="6" pageNumber="83" refId="ref8745" refString="Jussieu, A. L. de. (1805) Passifloraceae. Annales du Museum National d'Histoire Naturelle 6: 111" type="journal volume" year="1805">Jussieu (1805: 111)</bibRefCitation>
</taxonomicName>
and was often identified as such during earlier collections. However, the two species can readily be identified by the difference in foliar shape. In
<emphasis id="4BD4AE5EFFE1FF8CFA02FB57930B8389" italics="true" pageId="6" pageNumber="83">
<taxonomicName id="BEA009CFFFE1FF8CFA02FB57930F8389" authorityName="Molinari, Kuethe &amp; Bernal" authorityYear="2024" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="6" pageNumber="83" phylum="Tracheophyta" rank="species" species="rogitamensis">P. rogitamensis</taxonomicName>
,
</emphasis>
the leaves are distinctly longer (up to
<quantity id="BE58DFA9FFE1FF8CFD47FB6A91028388" box="[722,791,1202,1229]" metricMagnitude="-1" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="83" unit="cm" value="16.0">16 cm</quantity>
in length), narrow oblong to lanceolate, with large oblong stipules measuring up to
<quantity id="BE58DFA9FFE1FF8CFE0BFB0E93E583B4" box="[414,496,1238,1265]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="6" pageNumber="83" unit="cm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 cm</quantity>
in length. In contrast,
<taxonomicName id="BEA009CFFFE1FF8CFD7FFB0F917183B5" authorityName="Jussieu" authorityYear="1805" box="[746,868,1238,1264]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="6" pageNumber="83" phylum="Tracheophyta" rank="species" species="longipes">
<emphasis id="4BD4AE5EFFE1FF8CFD7FFB0F917183B5" box="[746,868,1238,1264]" italics="true" pageId="6" pageNumber="83">P. longipes</emphasis>
</taxonomicName>
has ovate to oblong-ovate leaves no more than
<quantity id="BE58DFA9FFE1FF8CFAE0FB0E97BE83B4" box="[1397,1451,1238,1265]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="83" unit="cm" value="8.0">8 cm</quantity>
in length, with ovate stipules up to
<quantity id="BE58DFA9FFE1FF8CFD9CFB22902B8250" box="[521,574,1274,1301]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="83" unit="cm" value="3.0">3 cm</quantity>
in length. The corona of
<taxonomicName id="BEA009CFFFE1FF8CFCC7FB2391E88251" authorityName="Molinari, Kuethe &amp; Bernal" authorityYear="2024" box="[850,1021,1275,1300]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="6" pageNumber="83" phylum="Tracheophyta" rank="species" species="rogitamensis">
<emphasis id="4BD4AE5EFFE1FF8CFCC7FB2391E88251" box="[850,1021,1275,1300]" italics="true" pageId="6" pageNumber="83">P. rogitamensis</emphasis>
</taxonomicName>
is a very light purple to whitish pink in colour, while in
<taxonomicName id="BEA009CFFFE1FF8CFEAEFAC793A0827D" authorityName="Jussieu" authorityYear="1805" box="[315,437,1310,1336]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="6" pageNumber="83" phylum="Tracheophyta" rank="species" species="longipes">
<emphasis id="4BD4AE5EFFE1FF8CFEAEFAC793A0827D" box="[315,437,1310,1336]" italics="true" pageId="6" pageNumber="83">P. longipes</emphasis>
</taxonomicName>
the corona is dark purple to blue. There is also a strong allopatry in habitat and distribution: with
<taxonomicName id="BEA009CFFFE1FF8CFF57FA9B937B8219" authorityName="Molinari, Kuethe &amp; Bernal" authorityYear="2024" box="[194,366,1347,1372]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="6" pageNumber="83" phylum="Tracheophyta" rank="species" species="rogitamensis">
<emphasis id="4BD4AE5EFFE1FF8CFF57FA9B937B8219" box="[194,366,1347,1372]" italics="true" pageId="6" pageNumber="83">P. rogitamensis</emphasis>
</taxonomicName>
being found in south-facing hills of northern
<collectingRegion id="BB64BCAEFFE1FF8CFCE1FA9A91DD8219" box="[884,968,1346,1372]" country="Colombia" name="Boyaca" pageId="6" pageNumber="83">Boyacá</collectingRegion>
and
<collectingRegion id="BB64BCAEFFE1FF8CFB95FA9A96678219" box="[1024,1138,1346,1372]" country="Colombia" name="Santander" pageId="6" pageNumber="83">Santander</collectingRegion>
, but
<taxonomicName id="BEA009CFFFE1FF8CFB3CFA9B97318219" authorityName="Jussieu" authorityYear="1805" box="[1193,1316,1346,1372]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="6" pageNumber="83" phylum="Tracheophyta" rank="species" species="longipes">
<emphasis id="4BD4AE5EFFE1FF8CFB3CFA9B97318219" box="[1193,1316,1346,1372]" italics="true" pageId="6" pageNumber="83">P. longipes</emphasis>
</taxonomicName>
confined to the west-facing slopes of the department of
<collectingRegion id="BB64BCAEFFE1FF8CFDE6FABE910F82C5" box="[627,794,1382,1408]" country="Colombia" name="Cundinamarca" pageId="6" pageNumber="83">Cundinamarca</collectingRegion>
.
</paragraph>
<paragraph id="791F724CFFE1FF8CFF28FA5296C280D9" blockId="6.[136,1452,158,1949]" pageId="6" pageNumber="83">
There remains some assertion that
<taxonomicName id="BEA009CFFFE1FF8CFDDDFA52914682E1" authorityName="Molinari, Kuethe &amp; Bernal" authorityYear="2024" box="[584,851,1418,1444]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="6" pageNumber="83" phylum="Tracheophyta" rank="species" species="rogitamensis">
<emphasis id="4BD4AE5EFFE1FF8CFDDDFA52914682E1" box="[584,851,1418,1444]" italics="true" pageId="6" pageNumber="83">Passiflora rogitamensis</emphasis>
</taxonomicName>
should be named
<taxonomicName id="BEA009CFFFE1FF8CFBB7FA5392EB828C" authority="L. Uribe (1977: 15)" authorityName="L. Uribe" authorityPageNumber="15" authorityYear="1977" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="6" pageNumber="83" phylum="Tracheophyta" rank="variety" species="longipes" variety="oxyphylla">
<emphasis id="4BD4AE5EFFE1FF8CFBB7FA53968B82E1" box="[1058,1182,1418,1444]" italics="true" pageId="6" pageNumber="83">P. longipes</emphasis>
var.
<emphasis id="4BD4AE5EFFE1FF8CFB42FA52975182E1" box="[1239,1348,1418,1444]" italics="true" pageId="6" pageNumber="83">oxyphylla</emphasis>
L.
<bibRefCitation id="1D310FBDFFE1FF8CFAF9FA5292EB828C" author="Uribe, L. U." pageId="6" pageNumber="83" pagination="13 - 18" refId="ref9660" refString="Uribe, L. U. (1977) Sertula florae colombiae, XIV. Caldasia 12: 13 - 18." type="journal article" year="1977">Uribe (1977: 15)</bibRefCitation>
</taxonomicName>
, for the species strongly resembling the [insufficient] description and subsequent herbarium filed under this name, however.
<bibRefCitation id="1D310FBDFFE1FF8CFEAAFA0A93C582A9" author="Uribe, L. U." box="[319,464,1490,1516]" pageId="6" pageNumber="83" pagination="13 - 18" refId="ref9660" refString="Uribe, L. U. (1977) Sertula florae colombiae, XIV. Caldasia 12: 13 - 18." type="journal article" year="1977">Uribe (1977)</bibRefCitation>
allocated his own specimen collected from Subachoque,
<collectingRegion id="BB64BCAEFFE1FF8CFBC7FA0A96E082A9" box="[1106,1269,1490,1516]" country="Colombia" name="Cundinamarca" pageId="6" pageNumber="83">Cundinamarca</collectingRegion>
(COL99282) as acting
<typeStatus id="A61BCCEEFFE1FF8CFF41FA2E932C8155" box="[212,313,1526,1552]" pageId="6" pageNumber="83" type="holotype">holotype</typeStatus>
, delivering a manuscript with merely a single sentence stating the leaf to be ovate-lanceolate with an acuminate apex as the defining trait to this species (
<bibRefCitation id="1D310FBDFFE1FF8CFD55F9C2915F8171" author="Uribe, L. U." box="[704,842,1562,1588]" pageId="6" pageNumber="83" pagination="13 - 18" refId="ref9660" refString="Uribe, L. U. (1977) Sertula florae colombiae, XIV. Caldasia 12: 13 - 18." type="journal article" year="1977">Uribe, 1977</bibRefCitation>
). The
<typeStatus id="A61BCCEEFFE1FF8CFC1BF9C391FB8170" box="[910,1006,1563,1589]" pageId="6" pageNumber="83" type="paratype">paratype</typeStatus>
(COL333274), collected nearby Virolín in
<collectingRegion id="BB64BCAEFFE1FF8CFF3CF9E69315811D" box="[169,256,1598,1624]" country="Colombia" name="Boyaca" pageId="6" pageNumber="83">Boyacá</collectingRegion>
, was described as having solely narrow elliptic-lanceolate leaves with acute apices. No defining traits concerning the flower, fruit or other vegetative traits were provided. Upon studying both those specimens, however, it became overwhelmingly evident that the
<typeStatus id="A61BCCEEFFE1FF8CFD16F95E90F081E5" box="[643,741,1670,1696]" pageId="6" pageNumber="83" type="holotype">holotype</typeStatus>
and
<typeStatus id="A61BCCEEFFE1FF8CFCB7F95F919781E4" box="[802,898,1671,1697]" pageId="6" pageNumber="83" type="paratype">paratype</typeStatus>
do not correspond with the same species; with the
<typeStatus id="A61BCCEEFFE1FF8CFF20F97293028181" box="[181,279,1706,1732]" pageId="6" pageNumber="83" type="holotype">holotype</typeStatus>
featuring a typical
<taxonomicName id="BEA009CFFFE1FF8CFE68F973906E8181" authorityName="Jussieu" authorityYear="1805" box="[509,635,1706,1732]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="6" pageNumber="83" phylum="Tracheophyta" rank="species" species="longipes">
<emphasis id="4BD4AE5EFFE1FF8CFE68F973906E8181" box="[509,635,1706,1732]" italics="true" pageId="6" pageNumber="83">P. longipes</emphasis>
</taxonomicName>
found across
<collectingRegion id="BB64BCAEFFE1FF8CFCB4F97291D18181" box="[801,964,1706,1732]" country="Colombia" name="Cundinamarca" pageId="6" pageNumber="83">Cundinamarca</collectingRegion>
and the
<typeStatus id="A61BCCEEFFE1FF8CFBBBF973969B8180" box="[1070,1166,1707,1733]" pageId="6" pageNumber="83" type="paratype">paratype</typeStatus>
being consistent with
<taxonomicName id="BEA009CFFFE1FF8CFA02F973930D81AD" authorityName="Molinari, Kuethe &amp; Bernal" authorityYear="2024" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="6" pageNumber="83" phylum="Tracheophyta" rank="species" species="rogitamensis">
<emphasis id="4BD4AE5EFFE1FF8CFA02F973930D81AD" italics="true" pageId="6" pageNumber="83">P. rogitamensis</emphasis>
</taxonomicName>
found from
<collectingRegion id="BB64BCAEFFE1FF8CFE33F91793EF81AC" box="[422,506,1743,1769]" country="Colombia" name="Boyaca" pageId="6" pageNumber="83">Boyaca</collectingRegion>
up into
<collectingRegion id="BB64BCAEFFE1FF8CFDCCF91690DE81AD" box="[601,715,1742,1768]" country="Colombia" name="Santander" pageId="6" pageNumber="83">Santander</collectingRegion>
. This was supported further by recent collections in both of these areas (e.g., iNaturalist observations: 115774902 and 50654857, as well as the photos presented in this paper). For the fact that the
<typeStatus id="A61BCCEEFFE1FF8CFE80F8CE93628075" box="[277,375,1814,1840]" pageId="6" pageNumber="83" type="holotype">holotype</typeStatus>
and
<typeStatus id="A61BCCEEFFE1FF8CFE3AF8CF901A8074" box="[431,527,1815,1841]" pageId="6" pageNumber="83" type="paratype">paratype</typeStatus>
do not correspond, combined with the explicit
<typeStatus id="A61BCCEEFFE1FF8CFBB3F8CE969D8075" box="[1062,1160,1814,1840]" pageId="6" pageNumber="83" type="holotype">holotype</typeStatus>
belonging to a plant of
<emphasis id="4BD4AE5EFFE1FF8CFA02F8CF92FB8011" italics="true" pageId="6" pageNumber="83">
<taxonomicName id="BEA009CFFFE1FF8CFA02F8CF92FF8011" authorityName="Jussieu" authorityYear="1805" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="6" pageNumber="83" phylum="Tracheophyta" rank="species" species="longipes">P. longipes</taxonomicName>
,
</emphasis>
the validity of the taxon “
<taxonomicName id="BEA009CFFFE1FF8CFDB5F8E291B18011" authorityName="L. Uribe" authorityYear="1977" box="[544,932,1850,1877]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="6" pageNumber="83" phylum="Tracheophyta" rank="variety" species="longipes" variety="oxyphylla">
<emphasis id="4BD4AE5EFFE1FF8CFDB5F8E290E98011" box="[544,764,1850,1876]" italics="true" pageId="6" pageNumber="83">Passiflora longipes</emphasis>
var.
<emphasis id="4BD4AE5EFFE1FF8CFCA2F8E291B18011" box="[823,932,1850,1876]" italics="true" pageId="6" pageNumber="83">oxyphylla</emphasis>
</taxonomicName>
” can be called into question. To resolve this, either the taxon needs to be redescribed with a new type (
<typeStatus id="A61BCCEEFFE1FF8CFC9FF8869173803D" box="[778,870,1886,1912]" pageId="6" pageNumber="83" type="neotype">neotype</typeStatus>
) assigned or to segregate this conundrum and apply the popular name
<taxonomicName id="BEA009CFFFE1FF8CFEC4F85B93E880D9" authorityName="Molinari, Kuethe &amp; Bernal" authorityYear="2024" box="[337,509,1923,1948]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="6" pageNumber="83" phylum="Tracheophyta" rank="species" species="rogitamensis">
<emphasis id="4BD4AE5EFFE1FF8CFEC4F85B93E880D9" box="[337,509,1923,1948]" italics="true" pageId="6" pageNumber="83">P. rogitamensis</emphasis>
</taxonomicName>
as it has already been used in surveys and literature to this point.
</paragraph>
</subSubSection>
</treatment>
</document>

537
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<mods:title id="226EBBA2A0CD7AF4213B7F4DB6587D91">Validating the names of three species of Passiflora from the northern portion of South America</mods:title>
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<mods:namePart id="E11A5187E496440A45752906D7D3B1AB">Bernal, Hernan Dario</mods:namePart>
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<mods:namePart id="E485951D59C24A56FD885C7BC92AE667">Molinari, Miguel</mods:namePart>
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<mods:namePart id="D2BD2DD50510F7192AF85A91E2EFE426">Vecchia, Maurizio</mods:namePart>
<mods:affiliation id="44F9318FAAE1288325DEB253BC99F37B">School of Environment, University of Auckland, New Zealand &amp; National Italian Passiflora Collection, Societá Botanica Italiana, Ripalta Cremasca, Italy &amp; School of Environment, University of Auckland, New Zealand</mods:affiliation>
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<treatment id="F109C35AFFE5FF8FFF1DFD2C92C38565" ID-DOI="http://doi.org/10.5281/zenodo.14515921" ID-Zenodo-Dep="14515921" LSID="urn:lsid:plazi:treatment:F109C35AFFE5FF8FFF1DFD2C92C38565" httpUri="http://treatment.plazi.org/id/F109C35AFFE5FF8FFF1DFD2C92C38565" lastPageId="5" lastPageNumber="82" pageId="2" pageNumber="79">
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<paragraph id="791F724CFFE5FF88FF1DFD2C9172844B" blockId="2.[136,871,756,783]" box="[136,871,756,783]" pageId="2" pageNumber="79">
<heading id="2257C520FFE5FF88FF1DFD2C9172844B" box="[136,871,756,783]" fontSize="11" level="2" pageId="2" pageNumber="79" reason="3">
<taxonomicName id="BEA009CFFFE5FF88FF1DFD2C904E844B" ID-CoL="CKNC3" authority="Molinari &amp; Kuethe" authorityName="Molinari &amp; Kuethe" authorityYear="2024" box="[136,603,756,783]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="79" phylum="Tracheophyta" rank="species" species="azulitensis" status="sp. nov.">
<emphasis id="4BD4AE5EFFE5FF88FF1DFD2C9368844A" bold="true" box="[136,381,756,783]" italics="true" pageId="2" pageNumber="79">Passiflora azulitensis</emphasis>
Molinari &amp; Kuethe
</taxonomicName>
<emphasis id="4BD4AE5EFFE5FF88FDF7FD2D90A2844B" box="[610,695,757,782]" italics="true" pageId="2" pageNumber="79">
<taxonomicNameLabel id="50E71325FFE5FF88FDF7FD2D90A2844B" box="[610,695,757,782]" pageId="2" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="E19B6EC9FFE5FF88FD50FD2C9106844B" box="[709,787,756,782]" captionStart="FIGURE 2" captionStartId="3.[136,234,1037,1059]" captionTargetBox="[151,1435,190,1013]" captionTargetId="figure-19@3.[151,1435,190,1013]" captionTargetPageId="3" captionText="FIGURE 2. A–F. Comparative morfology showing leaves and flowers of the related species to Passiflora azulitensis. A, B. Passiflora azulitensis, La Azulita, Venezuela; C, D. Passiflora guayaquilensis, El Oro province, Ecuador; E, F. Passiflora adenopoda, Merida, Venezuela. Photo courtesy by Miguel Molinari (A), J.R. Kuethe (B, E, F), and Xavier Cornejo (C, D)." figureDoi="http://doi.org/10.5281/zenodo.14515929" httpUri="https://zenodo.org/record/14515929/files/figure.png" pageId="2" pageNumber="79">Figs. 2</figureCitation>
and
<figureCitation id="E19B6EC9FFE5FF88FCDCFD2C914C844B" box="[841,857,756,782]" captionStart="FIGURE 3" captionStartId="4.[136,234,1922,1944]" captionTargetBox="[151,1436,190,1898]" captionTargetId="figure-22@4.[151,1436,190,1898]" captionTargetPageId="4" captionText="FIGURE 3. A–C. Passiflora azulitensis., Lankaster Composite Dissection Plate (LCDP); A. flower in situ; longitudinal of the flower (A1), floral orientation and bracts in situ (A2), detail of the bracts (A3), detail of the reduced corona (A4); B. habit and vegetative characteristics of P. azulitensis, with detail of the stipules (B1), petiolar nectaries (B2), and foliar margin (B3); C. A mature fruit found in situ. Compiled from photographs taken by Miguel Molinari, Andrés Miguel Orellana and J.R. Kuethe." figureDoi="http://doi.org/10.5281/zenodo.14515931" httpUri="https://zenodo.org/record/14515931/files/figure.png" pageId="2" pageNumber="79">3</figureCitation>
).
</heading>
</paragraph>
</subSubSection>
<subSubSection id="31BA21C7FFE5FF88FF1DFC9891C084D8" pageId="2" pageNumber="79" type="materials_examined">
<paragraph id="791F724CFFE5FF88FF1DFC9891C084D8" blockId="2.[136,1452,832,1070]" pageId="2" pageNumber="79">
<emphasis id="4BD4AE5EFFE5FF88FF1DFC9892D78410" bold="true" box="[136,194,832,853]" pageId="2" pageNumber="79">Type:</emphasis>
—
<materialsCitation id="C9C87811FFE5FF88FF4EFC9896FF843F" collectingDate="2004-06-18" collectorName="La Azulita &amp; M. Molinari" country="Venezuela" elevation="1386" latitude="8.694317" location="La Azulita" longLatPrecision="1" longitude="-71.43816" pageId="2" pageNumber="79" specimenCount="1" stateProvince="Merida">
<collectingCountry id="01B732DCFFE5FF88FF4EFC9893798410" box="[219,364,832,853]" name="Venezuela" pageId="2" pageNumber="79">VENEZUELA</collectingCountry>
.
<collectingRegion id="BB64BCAEFFE5FF88FEE3FC9893AA8413" box="[374,447,832,854]" country="Venezuela" name="Merida" pageId="2" pageNumber="79">Mérida</collectingRegion>
: 2,
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<quantity id="BE58DFA9FFE5FF88FE4EFC98901B8413" box="[475,526,832,854]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="79" unit="km" value="2.0">2 km</quantity>
South-East of La Azulita
</locationDeviation>
, on the road from
<collectorName id="D455179AFFE5FF88FC54FC9896328413" box="[961,1063,832,854]" pageId="2" pageNumber="79">
<location id="7C7F2497FFE5FF88FC54FC9896328413" LSID="urn:lsid:plazi:treatment:F109C35AFFE5FF8FFF1DFD2C92C38565:7C7F2497FFE5FF88FC54FC9896328413" box="[961,1063,832,854]" country="Venezuela" latitude="8.694317" longLatPrecision="1" longitude="-71.43816" name="La Azulita" pageId="2" pageNumber="79" stateProvince="Merida">La Azulita</location>
</collectorName>
to
<collectingRegion id="BB64BCAEFFE5FF88FBDDFC9896858413" box="[1096,1168,832,854]" country="Venezuela" name="Merida" pageId="2" pageNumber="79">Mérida</collectingRegion>
,
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,
<geoCoordinate id="1C94148BFFE5FF88FAB0FC9897BD8413" box="[1317,1448,832,854]" degrees="71" direction="west" minutes="26.290" orientation="longitude" pageId="2" pageNumber="79" precision="1" value="-71.43816">71°26.290’W</geoCoordinate>
,
<quantity id="BE58DFA9FFE5FF88FF28FCBC931B843F" box="[189,270,868,890]" metricMagnitude="3" metricUnit="m" metricValue="1.386" pageId="2" pageNumber="79" unit="m" value="1386.0">
<elevation id="F28D957FFFE5FF88FF28FCBC931B843F" box="[189,270,868,890]" metricMagnitude="3" metricUnit="m" metricValue="1.386" pageId="2" pageNumber="79" unit="m" value="1386.0">1386 m</elevation>
</quantity>
.a.s.l.,
<date id="0D1E548CFFE5FF88FED2FCBC93D9843C" box="[327,460,868,889]" pageId="2" pageNumber="79" value="2004-06-18">
<collectingDate id="1D5AAD64FFE5FF88FED2FCBC93D9843C" box="[327,460,868,889]" pageId="2" pageNumber="79" value="2004-06-18">18 June 2004</collectingDate>
</date>
, (fl.),
<emphasis id="4BD4AE5EFFE5FF88FD9FFCBC90BC843C" box="[522,681,868,889]" italics="true" pageId="2" pageNumber="79">
<collectorName id="D455179AFFE5FF88FD9FFCBC9095843C" box="[522,640,868,889]" pageId="2" pageNumber="79">M. Molinari</collectorName>
s.n.
</emphasis>
[TYPE established from cultivated material], grown in the
</materialsCitation>
<materialsCitation id="C9C87811FFE5FF88FB64FCBC91C584D8" country="United Kingdom" location="National Collection" pageId="2" pageNumber="79" specimenCount="2" typeStatus="holotype">
<collectingCountry id="01B732DCFFE5FF88FB64FCBC9701843C" box="[1265,1300,868,889]" name="United Kingdom" pageId="2" pageNumber="79">UK</collectingCountry>
at the
<location id="7C7F2497FFE5FF88FACCFCBC933484DB" LSID="urn:lsid:plazi:treatment:F109C35AFFE5FF8FFF1DFD2C92C38565:7C7F2497FFE5FF88FACCFCBC933484DB" country="United Kingdom" name="National Collection" pageId="2" pageNumber="79">National Collection</location>
<taxonomicName id="BEA009CFFFE5FF88FEB2FC50903384D8" authority=", J. Vanderplank" authorityName="J. Vanderplank" box="[295,550,904,925]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="79" phylum="Tracheophyta" rank="genus">
<emphasis id="4BD4AE5EFFE5FF88FEB2FC50939F84D8" box="[295,394,904,925]" italics="true" pageId="2" pageNumber="79">Passiflora</emphasis>
,
<emphasis id="4BD4AE5EFFE5FF88FE03FC50903384D8" box="[406,550,904,925]" italics="true" pageId="2" pageNumber="79">J. Vanderplank</emphasis>
</taxonomicName>
(
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: NCP 2523/22A!;
<typeStatus id="A61BCCEEFFE5FF88FCA8FC50919384DB" box="[829,902,904,926]" pageId="2" pageNumber="79" type="isotype">isotype</typeStatus>
: NCP!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="31BA21C7FFE5FF88FF1DFC74934A836B" pageId="2" pageNumber="79" type="diagnosis">
<paragraph id="791F724CFFE5FF88FF1DFC74934A836B" blockId="2.[136,1452,832,1070]" pageId="2" pageNumber="79">
<emphasis id="4BD4AE5EFFE5FF88FF1DFC7492E68484" bold="true" box="[136,243,940,961]" pageId="2" pageNumber="79">
<emphasis id="4BD4AE5EFFE5FF88FF1DFC7492FE8484" bold="true" box="[136,235,940,961]" italics="true" pageId="2" pageNumber="79">Diagnosis</emphasis>
:
</emphasis>
—
<taxonomicName id="BEA009CFFFE5FF88FE9EFC7493CE8484" authorityName="Molinari &amp; Kuethe" authorityYear="2024" box="[267,475,940,961]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="79" phylum="Tracheophyta" rank="species" species="azulitensis">
<emphasis id="4BD4AE5EFFE5FF88FE9EFC7493CE8484" box="[267,475,940,961]" italics="true" pageId="2" pageNumber="79">Passiflora azulitensis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="BEA009CFFFE5FF88FDF4FC7490F38484" box="[609,742,940,961]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="79" phylum="Tracheophyta" rank="species" species="adenopoda">
<emphasis id="4BD4AE5EFFE5FF88FDF4FC7490F38484" box="[609,742,940,961]" italics="true" pageId="2" pageNumber="79">P. adenopoda</emphasis>
</taxonomicName>
and
<taxonomicName id="BEA009CFFFE5FF88FC8CFC7491998484" box="[793,908,940,961]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="79" phylum="Tracheophyta" rank="species" species="morifolia">
<emphasis id="4BD4AE5EFFE5FF88FC8CFC7491998484" box="[793,908,940,961]" italics="true" pageId="2" pageNumber="79">P. morifolia</emphasis>
</taxonomicName>
by the small (&lt;
<quantity id="BE58DFA9FFE5FF88FBB9FC7496658484" box="[1068,1136,940,962]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="79" unit="cm" value="2.5">2.5 cm</quantity>
<emphasis id="4BD4AE5EFFE5FF88FBECFC7496818484" box="[1145,1172,940,961]" italics="true" pageId="2" pageNumber="79">vs.</emphasis>
&gt;
<quantity id="BE58DFA9FFE5FF88FB3CFC7496E58484" box="[1193,1264,940,962]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="79" unit="cm" value="2.5">2.5 cm</quantity>
), completely white flowers (
<emphasis id="4BD4AE5EFFE5FF88FE80FC08932584A0" box="[277,304,976,997]" italics="true" pageId="2" pageNumber="79">vs.</emphasis>
white with purple in
<taxonomicName id="BEA009CFFFE5FF88FD90FC08909C84A0" box="[517,649,976,997]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="79" phylum="Tracheophyta" rank="species" species="adenopoda">
<emphasis id="4BD4AE5EFFE5FF88FD90FC08909C84A0" box="[517,649,976,997]" italics="true" pageId="2" pageNumber="79">P. adenopoda</emphasis>
</taxonomicName>
and
<taxonomicName id="BEA009CFFFE5FF88FD2EFC08913884A0" box="[699,813,976,997]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="79" phylum="Tracheophyta" rank="species" species="morifolia">
<emphasis id="4BD4AE5EFFE5FF88FD2EFC08913884A0" box="[699,813,976,997]" italics="true" pageId="2" pageNumber="79">P. morifolia</emphasis>
</taxonomicName>
); corona that is shorter than
<quantity id="BE58DFA9FFE5FF88FBDEFC08969684A0" box="[1099,1155,976,997]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="79" unit="mm" value="3.0">3 mm</quantity>
and recurved (
<emphasis id="4BD4AE5EFFE5FF88FA8CFC08972184A0" box="[1305,1332,976,997]" italics="true" pageId="2" pageNumber="79">vs.</emphasis>
&gt;
<quantity id="BE58DFA9FFE5FF88FADCFC08979484A0" box="[1353,1409,976,997]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="79" unit="mm" value="3.0">3 mm</quantity>
and straight), leaves with denticulate (
<emphasis id="4BD4AE5EFFE5FF88FD99FC2C9032834C" box="[524,551,1012,1033]" italics="true" pageId="2" pageNumber="79">vs.</emphasis>
serrate or crenulate) margins and deeply serrate to ciliate (
<emphasis id="4BD4AE5EFFE5FF88FBE4FC2C969E834C" box="[1137,1163,1012,1033]" italics="true" pageId="2" pageNumber="79">vs.</emphasis>
smooth or shallowly serrate) stipular margins.
</paragraph>
</subSubSection>
<subSubSection id="31BA21C7FFE5FF88FF1DFB84907A8093" pageId="2" pageNumber="79" type="description">
<paragraph id="791F724CFFE5FF88FF1DFB849101802F" blockId="2.[136,1452,1116,2079]" pageId="2" pageNumber="79">
<emphasis id="4BD4AE5EFFE5FF88FF1DFB8493098333" bold="true" box="[136,284,1116,1142]" pageId="2" pageNumber="79">Description:</emphasis>
—Climbing vine,
<quantity id="BE58DFA9FFE5FF88FE79FB8490238332" box="[492,566,1116,1143]" metricMagnitude="0" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="2" pageNumber="79" unit="m" value="5.0" valueMax="6.0" valueMin="4.0">4–6 m</quantity>
long with perennial stems, semi-woody at base, finely pubescent throughout with whitish uncinate trichomes.
<emphasis id="4BD4AE5EFFE5FF88FD91FB58905083DF" box="[516,581,1152,1178]" italics="true" pageId="2" pageNumber="79">Stems</emphasis>
terete, internodal distance
<quantity id="BE58DFA9FFE5FF88FCEEFB5891E383DE" box="[891,1014,1152,1179]" metricMagnitude="-2" metricUnit="m" metricValue="8.1" metricValueMax="12.0" metricValueMin="4.2" pageId="2" pageNumber="79" unit="cm" value="8.1" valueMax="12.0" valueMin="4.2">4.2–12 cm</quantity>
.
<emphasis id="4BD4AE5EFFE5FF88FB97FB58964E83DF" box="[1026,1115,1152,1178]" italics="true" pageId="2" pageNumber="79">Stipules</emphasis>
20–22 ×
<quantity id="BE58DFA9FFE5FF88FB5EFB58975283DE" box="[1227,1351,1152,1179]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="2" pageNumber="79" unit="mm" value="11.5" valueMax="12.0" valueMin="11.0">11–12 mm</quantity>
, oblongobovate or semi-circular, subcordate at base, caudate at apex, the margin with 9–10 denticulate or caudate teeth, ciliolate, green.
<emphasis id="4BD4AE5EFFE5FF88FEA8FB10938283A7" box="[317,407,1224,1250]" italics="true" pageId="2" pageNumber="79">Petioles</emphasis>
<quantity id="BE58DFA9FFE5FF88FE0AFB10901283A6" box="[415,519,1224,1251]" metricMagnitude="-2" metricUnit="m" metricValue="4.75" metricValueMax="5.0" metricValueMin="4.5" pageId="2" pageNumber="79" unit="cm" value="4.75" valueMax="5.0" valueMin="4.5">4.5–5 cm</quantity>
long, terete, biglandular with one opposite pair positioned
<quantity id="BE58DFA9FFE5FF88FB35FB1096E883A6" box="[1184,1277,1224,1251]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="2" pageNumber="79" unit="mm" value="6.5" valueMax="8.0" valueMin="5.0">5–8 mm</quantity>
below the base of the blade, hemispherical and sessile to long-stipitate, often cup-shaped with opening abaxially, 5–6 ×
<quantity id="BE58DFA9FFE5FF88FAA4FB3497B28242" box="[1329,1447,1260,1287]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" metricValueMax="3.0" metricValueMin="2.8" pageId="2" pageNumber="79" unit="mm" value="2.9" valueMax="3.0" valueMin="2.8">2.8–3 mm</quantity>
, green.
<emphasis id="4BD4AE5EFFE5FF88FF41FAC8935E826F" box="[212,331,1296,1322]" italics="true" pageId="2" pageNumber="79">Leaf blade</emphasis>
14–22 ×
<quantity id="BE58DFA9FFE5FF88FE20FAC89032826E" box="[437,551,1296,1323]" metricMagnitude="-1" metricUnit="m" metricValue="1.65" metricValueMax="2.0" metricValueMin="1.3" pageId="2" pageNumber="79" unit="cm" value="16.5" valueMax="20.0" valueMin="13.0">13–20 cm</quantity>
, palmately 5(–7)-lobed, lobation borne
<quantity id="BE58DFA9FFE5FF88FC76FAC89623826E" box="[995,1078,1296,1323]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="2" pageNumber="79" unit="cm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 cm</quantity>
from base, lobes 8–17 ×
<quantity id="BE58DFA9FFE5FF88FAC4FAC897B2826E" box="[1361,1447,1296,1323]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="2" pageNumber="79" unit="cm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 cm</quantity>
, oblong-ovate to lanceolate, intermediate lobe the longest, squarely cordate at base, acuminate to acute at apex, margins (dentate) denticulate or apiculate with serrulate teeth terminating into mucros; foliar texture membranous, deep green on upper surface, dullish green below.
<emphasis id="4BD4AE5EFFE5FF88FDDEFAA490F482D3" box="[587,737,1404,1430]" italics="true" pageId="2" pageNumber="79">Inflorescence</emphasis>
solitary or in pairs, inconspicuous, borne on short peduncles.
<emphasis id="4BD4AE5EFFE5FF88FF1DFA7892EE82FF" box="[136,251,1440,1466]" italics="true" pageId="2" pageNumber="79">Peduncles</emphasis>
<quantity id="BE58DFA9FFE5FF88FE90FA78934982FE" box="[261,348,1440,1467]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="2" pageNumber="79" unit="cm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 cm</quantity>
long, green, articulate 2.0–
<quantity id="BE58DFA9FFE5FF88FD08FA7890F982FE" box="[669,748,1440,1467]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="79" unit="cm" value="2.7">2.7 cm</quantity>
from the base of the flower.
<emphasis id="4BD4AE5EFFE5FF88FBD2FA79968582FF" box="[1095,1168,1441,1466]" italics="true" pageId="2" pageNumber="79">Bracts</emphasis>
3–5 ×
<quantity id="BE58DFA9FFE5FF88FB7CFA78975982FE" box="[1257,1356,1440,1467]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="2" pageNumber="79" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
, located in lower half of peduncle, acutely linear or reduced to a three-lobed spinifex structure, widely obtrullate in general outline, laciniate, green.
<emphasis id="4BD4AE5EFFE5FF88FE35FA3093E98147" box="[416,508,1512,1538]" italics="true" pageId="2" pageNumber="79">Flowers</emphasis>
<quantity id="BE58DFA9FFE5FF88FD91FA3090978146" box="[516,642,1512,1539]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="2" pageNumber="79" unit="cm" value="2.35" valueMax="2.5" valueMin="2.2">2.2–2.5 cm</quantity>
in diameter, white entirely; hypanthium
<quantity id="BE58DFA9FFE5FF88FBC7FA3096DE8146" box="[1106,1227,1512,1539]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="2" pageNumber="79" unit="mm" value="12.0" valueMax="13.0" valueMin="11.0">11–13 mm</quantity>
in diameter, deeply campanulate, green to pale green outside, whitish inside; sepals 12–15 ×
<quantity id="BE58DFA9FFE5FF88FC4BF9D4965B8162" box="[990,1102,1548,1575]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="2" pageNumber="79" unit="mm" value="9.5" valueMax="11.0" valueMin="8.0">8–11 mm</quantity>
, ovate-triangular to somewhat oblong, at base tapering to an acute to acuminate at apex, greenish abaxially, whitish adaxially, conspicuously cornate or corniculate near apex, awn
<quantity id="BE58DFA9FFE5FF88FE4DF98C9020812A" box="[472,565,1620,1647]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="2" pageNumber="79" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
long, green, thick; petals 13–16 ×
<quantity id="BE58DFA9FFE5FF88FC2DF98C9602812A" box="[952,1047,1620,1647]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="2" pageNumber="79" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6–7 mm</quantity>
, subequal to sepals, ovate-oblong to ovate, truncate at apex, pure white at both sides, membranous; coronal composed of a single denticulate series,
<quantity id="BE58DFA9FFE5FF88FAF9F9A092A181F2" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="2" pageNumber="79" unit="mm" value="2.25" valueMax="2.5" valueMin="2.0">2–2.5 mm</quantity>
long, pure white throughout, slightly recurved; operculum 2.4–3.0 mm long, membranous, recurved and cleft to the base, margin denticulate, white and unmarked; limen whitish, much reduced, not spotted; androgynophore
<quantity id="BE58DFA9FFE5FF88FAC5F91897B9819E" box="[1360,1452,1728,1755]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="2" pageNumber="79" unit="mm" value="8.5" valueMax="9.0" valueMin="8.0">8–9 mm</quantity>
long, white, not speckled; anthers ca. 4.0 mm long, orange-yellow; ovary 4–4.5 ×
<quantity id="BE58DFA9FFE5FF88FBBDF93C96A781BA" box="[1064,1202,1764,1791]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="3.8" metricValueMin="3.2" pageId="2" pageNumber="79" unit="mm" value="3.5" valueMax="3.8" valueMin="3.2">3.2–3.8 mm</quantity>
, ellipsoid, glabrous to minutely tomentose, green.
<emphasis id="4BD4AE5EFFE5FF88FE2BF8D193ED8067" box="[446,504,1801,1826]" italics="true" pageId="2" pageNumber="79">Fruit</emphasis>
5–5.2 ×
<quantity id="BE58DFA9FFE5FF88FDCFF8D090D08066" box="[602,709,1800,1827]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="2" pageNumber="79" unit="cm" value="4.25" valueMax="4.5" valueMin="4.0">4–4.5 cm</quantity>
, widely ellipsoid or sometimes obovate, rounded at apex, glabrous or finely pubescent, pink-purple when mature, inedible; mesocarp whitish with slightly greenish and purple marks; arils bright orange to reddish.
<emphasis id="4BD4AE5EFFE5FF88FE4FF888900F802F" box="[474,538,1872,1898]" italics="true" pageId="2" pageNumber="79">Seeds</emphasis>
black [not measured].
</paragraph>
<paragraph id="791F724CFFE5FF88FF28F8AC907A8093" blockId="2.[136,1452,1116,2079]" pageId="2" pageNumber="79">
<emphasis id="4BD4AE5EFFE5FF88FF28F8AC935680CB" bold="true" box="[189,323,1908,1934]" pageId="2" pageNumber="79">Phenology:</emphasis>
—This species was found in flower between June and September, and subsequent flowers produced in cultivation in
<collectingCountry id="01B732DCFFE5FF88FED6F84093A380F7" box="[323,438,1944,1970]" name="Venezuela" pageId="2" pageNumber="79">Venezuela</collectingCountry>
cohort with these months. Fruit was produced subsequently, with maturity in Venezuelan cultivation between August and September.
</paragraph>
</subSubSection>
<subSubSection id="31BA21C7FFE5FF88FF28F838930E8F5A" pageId="2" pageNumber="79" type="etymology">
<paragraph id="791F724CFFE5FF88FF28F838930E8F5A" blockId="2.[136,1452,1116,2079]" pageId="2" pageNumber="79">
<emphasis id="4BD4AE5EFFE5FF88FF28F838935D80BF" bold="true" box="[189,328,2016,2042]" pageId="2" pageNumber="79">Etymology:</emphasis>
—The epithet
<taxonomicName id="BEA009CFFFE5FF88FE7CF838907E80BF" authorityName="Molinari &amp; Kuethe" authorityYear="2024" box="[489,619,2016,2042]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="79" phylum="Tracheophyta" rank="species" species="azulitensis">
“
<emphasis id="4BD4AE5EFFE5FF88FE60F838907E80BF" box="[501,619,2016,2042]" italics="true" pageId="2" pageNumber="79">azulitensis</emphasis>
</taxonomicName>
” is chosen after the small region where the species was found; La Azulita,
<collectingRegion id="BB64BCAEFFE5FF88FF1DF7DC93028F5A" box="[136,279,2052,2079]" country="Venezuela" name="Merida" pageId="2" pageNumber="79">Merida state</collectingRegion>
.
</paragraph>
</subSubSection>
<caption id="2DDF22C4FFE4FF89FF1DFBD5964E832F" ID-DOI="http://doi.org/10.5281/zenodo.14515929" ID-Zenodo-Dep="14515929" httpUri="https://zenodo.org/record/14515929/files/figure.png" pageId="3" pageNumber="80" startId="3.[136,234,1037,1059]" targetBox="[151,1435,190,1013]" targetPageId="3" targetType="figure">
<paragraph id="791F724CFFE4FF89FF1DFBD5964E832F" blockId="3.[136,1452,1037,1131]" pageId="3" pageNumber="80">
<emphasis id="4BD4AE5EFFE4FF89FF1DFBD593208367" bold="true" box="[136,309,1037,1059]" pageId="3" pageNumber="80">FIGURE 2. A–F</emphasis>
. Comparative morfology showing leaves and flowers of the related species to
<taxonomicName id="BEA009CFFFE4FF89FBA7FBD597158367" authorityName="Molinari &amp; Kuethe" authorityYear="2024" box="[1074,1280,1037,1058]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="80" phylum="Tracheophyta" rank="species" species="azulitensis">
<emphasis id="4BD4AE5EFFE4FF89FBA7FBD597158367" box="[1074,1280,1037,1058]" italics="true" pageId="3" pageNumber="80">Passiflora azulitensis</emphasis>
</taxonomicName>
.
<emphasis id="4BD4AE5EFFE4FF89FA98FBD597298367" bold="true" box="[1293,1340,1037,1058]" pageId="3" pageNumber="80">A, B</emphasis>
.
<taxonomicName id="BEA009CFFFE4FF89FADCFBD593738302" authority=", La Azulita" authorityName="La Azulita" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="80" phylum="Tracheophyta" rank="species" species="azulitensis">
<emphasis id="4BD4AE5EFFE4FF89FADCFBD592F88303" italics="true" pageId="3" pageNumber="80">Passiflora azulitensis</emphasis>
, La Azulita
</taxonomicName>
, Venezuela;
<emphasis id="4BD4AE5EFFE4FF89FE70FBE990038303" bold="true" box="[485,534,1073,1095]" pageId="3" pageNumber="80">C, D</emphasis>
.
<taxonomicName id="BEA009CFFFE4FF89FDB0FBE991668303" authority=", El Oro" authorityName="El Oro" box="[549,883,1073,1095]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="80" phylum="Tracheophyta" rank="species" species="guayaquilensis">
<emphasis id="4BD4AE5EFFE4FF89FDB0FBE991328303" box="[549,807,1073,1094]" italics="true" pageId="3" pageNumber="80">Passiflora guayaquilensis,</emphasis>
El Oro
</taxonomicName>
province, Ecuador;
<emphasis id="4BD4AE5EFFE4FF89FBABFBE996798303" bold="true" box="[1086,1132,1073,1094]" pageId="3" pageNumber="80">E, F</emphasis>
.
<emphasis id="4BD4AE5EFFE4FF89FBEEFBE9974D8303" box="[1147,1368,1073,1094]" italics="true" pageId="3" pageNumber="80">
<taxonomicName id="BEA009CFFFE4FF89FBEEFBE997468303" box="[1147,1363,1073,1094]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="80" phylum="Tracheophyta" rank="species" species="adenopoda">Passiflora adenopoda</taxonomicName>
,
</emphasis>
Merida, Venezuela. Photo courtesy by Miguel Molinari (A), J.R. Kuethe (B, E, F), and Xavier Cornejo (C, D).
</paragraph>
</caption>
<subSubSection id="31BA21C7FFE4FF8FFF28FB4192C38565" lastPageId="5" lastPageNumber="82" pageId="3" pageNumber="80" type="distribution">
<paragraph id="791F724CFFE4FF89FF28FB4191A0822D" blockId="3.[136,1452,1177,2067]" pageId="3" pageNumber="80">
<emphasis id="4BD4AE5EFFE4FF89FF28FB41934C83F6" bold="true" box="[189,345,1177,1203]" pageId="3" pageNumber="80">Distribution:</emphasis>
—
<taxonomicName id="BEA009CFFFE4FF89FEE0FB41907383F6" box="[373,614,1177,1203]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="80" phylum="Tracheophyta" rank="species" species="azulitensis">
<emphasis id="4BD4AE5EFFE4FF89FEE0FB41907383F6" box="[373,614,1177,1203]" italics="true" pageId="3" pageNumber="80">Passiflora azulitensis</emphasis>
</taxonomicName>
can be found only within a small virgin area nearby La Azulita belonging to the
<collectingRegion id="BB64BCAEFFE4FF89FF5BFB659349839D" box="[206,348,1213,1240]" country="Venezuela" name="Merida" pageId="3" pageNumber="80">Merida state</collectingRegion>
,
<collectingCountry id="01B732DCFFE4FF89FEF2FB6593CF8392" box="[359,474,1213,1239]" name="Venezuela" pageId="3" pageNumber="80">Venezuela</collectingCountry>
(
<figureCitation id="E19B6EC9FFE4FF89FE7FFB65903E839D" box="[490,555,1213,1240]" captionStart="FIGURE 1" captionStartId="1.[136,234,967,989]" captionTargetBox="[151,1436,187,940]" captionTargetId="figure-19@1.[151,1436,187,940]" captionTargetPageId="1" captionText="FIGURE 1. Map showing the allopatric distribution of the concerning new species (diamonds) and their nearest relatives (circles). Passiflora azulitensis (white); P. cocuyensis (red); P. rogitamensis (purple); Passiflora gritensis (green); P. semiciliosa (yellow); and P. longipes (blue). Map generated using both herbarium coordinates and georeferenced sightings (fieldwork supplemented with iNaturalist observations)." figureDoi="http://doi.org/10.5281/zenodo.14515923" httpUri="https://zenodo.org/record/14515923/files/figure.png" pageId="3" pageNumber="80">Fig. 1</figureCitation>
). This rich biodiverse region is perched on the northern flanks of the
<collectingCountry id="01B732DCFFE4FF89FAADFB6597BE8392" box="[1336,1451,1213,1239]" name="Venezuela" pageId="3" pageNumber="80">Venezuela</collectingCountry>
Andes, where it grows at elevations of about
<quantity id="BE58DFA9FFE4FF89FDFBFB3A90EC83B9" box="[622,761,1250,1276]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="3" pageNumber="80" unit="m" value="850.0" valueMax="1000.0" valueMin="700.0">700–1000 m</quantity>
in dense, pre-montane rain and cloud forest. Rather contrasting from its semi-arid Mexican relatives,
<taxonomicName id="BEA009CFFFE4FF89FDBCFADD910D825A" box="[553,792,1285,1311]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="80" phylum="Tracheophyta" rank="species" species="azulitensis">
<emphasis id="4BD4AE5EFFE4FF89FDBCFADD910D825A" box="[553,792,1285,1311]" italics="true" pageId="3" pageNumber="80">Passiflora azulitensis</emphasis>
</taxonomicName>
was found in a region marked by very intense rainfall, with high levels of humidity and precipitation year-round. This species was seen growing as a robust vine up trees and large bushes, with merely the free-hanging branches giving away its presence.
</paragraph>
<paragraph id="791F724CFFE4FF89FF28FAA993658105" blockId="3.[136,1452,1177,2067]" pageId="3" pageNumber="80">
<emphasis id="4BD4AE5EFFE4FF89FF28FAA9937282CE" bold="true" box="[189,359,1393,1419]" pageId="3" pageNumber="80">Conservation:</emphasis>
—With merely two known collections both from within this small virgin area, the abundance and thus conservation for
<taxonomicName id="BEA009CFFFE4FF89FE14FA4D906682EA" box="[385,627,1429,1455]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="80" phylum="Tracheophyta" rank="species" species="azulitensis">
<emphasis id="4BD4AE5EFFE4FF89FE14FA4D906682EA" box="[385,627,1429,1455]" italics="true" pageId="3" pageNumber="80">Passiflora azulitensis</emphasis>
</taxonomicName>
is alarmingly limited. The area is not under any protection by a reserve or forest protectorate, and deforestation or urban development are a significant threat to the long term survival of this species in the wild. Although with just two collection sites there is too little geographic data available to calculate an acurate AOO and EOO (essentially scaling it as DD), the authors chose to keep the species as CR criteria B1a, C1, C2ai and D (IUCN 2023).
</paragraph>
<paragraph id="791F724CFFE4FF89FF28F99196818126" blockId="3.[136,1452,1177,2067]" box="[189,1172,1609,1636]" pageId="3" pageNumber="80">
<emphasis id="4BD4AE5EFFE4FF89FF28F99190038121" bold="true" box="[189,534,1609,1636]" pageId="3" pageNumber="80">
Additional specimens
<emphasis id="4BD4AE5EFFE4FF89FE51F99190188126" bold="true" box="[452,525,1609,1635]" italics="true" pageId="3" pageNumber="80">in situ</emphasis>
:
</emphasis>
—No additional specimens were found in other herbaria.
</paragraph>
<paragraph id="791F724CFFE4FF89FF28F9B5917C81B6" blockId="3.[136,1452,1177,2067]" pageId="3" pageNumber="80">
<emphasis id="4BD4AE5EFFE4FF89FF28F9B593D381C2" bold="true" box="[189,454,1645,1671]" pageId="3" pageNumber="80">Cultivated collections:</emphasis>
—Blumen &amp;
<taxonomicName id="BEA009CFFFE4FF89FDCFF9B590D881C2" box="[602,717,1645,1671]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="80" phylum="Tracheophyta" rank="genus">
<emphasis id="4BD4AE5EFFE4FF89FDCFF9B590D881C2" box="[602,717,1645,1671]" italics="true" pageId="3" pageNumber="80">Passiflora</emphasis>
</taxonomicName>
(
<collectingCountry id="01B732DCFFE4FF89FD4FF9B6915D81CD" box="[730,840,1646,1672]" name="Germany" pageId="3" pageNumber="80">Germany</collectingCountry>
),
<emphasis id="4BD4AE5EFFE4FF89FCCCF9B691D181C2" box="[857,964,1646,1671]" italics="true" pageId="3" pageNumber="80">M. Drews</emphasis>
; Madeira
<taxonomicName id="BEA009CFFFE4FF89FBA6F9B597B681C2" authority="Gardens, P. Mendonca" authorityName="Gardens, P. Mendonca" box="[1075,1443,1645,1671]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="80" phylum="Tracheophyta" rank="genus">
<emphasis id="4BD4AE5EFFE4FF89FBA6F9B596B381C2" box="[1075,1190,1645,1671]" italics="true" pageId="3" pageNumber="80">Passiflora</emphasis>
Gardens,
<emphasis id="4BD4AE5EFFE4FF89FA81F9B697B681C2" box="[1300,1443,1645,1671]" italics="true" pageId="3" pageNumber="80">P. Mendonca</emphasis>
</taxonomicName>
;
<taxonomicName id="BEA009CFFFE4FF89FF1DF94992EE81EE" box="[136,251,1681,1707]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="80" phylum="Tracheophyta" rank="genus">
<emphasis id="4BD4AE5EFFE4FF89FF1DF94992EE81EE" box="[136,251,1681,1707]" italics="true" pageId="3" pageNumber="80">Passiflora</emphasis>
</taxonomicName>
collection
<collectingCountry id="01B732DCFFE4FF89FEE1F94993DC81EE" box="[372,457,1681,1707]" name="Austria" pageId="3" pageNumber="80">Austria</collectingCountry>
,
<emphasis id="4BD4AE5EFFE4FF89FE46F94A902C81EE" box="[467,569,1682,1707]" italics="true" pageId="3" pageNumber="80">M. Varga</emphasis>
; Passiflorahoeve (
<collectingCountry id="01B732DCFFE4FF89FC90F949918681EE" box="[773,915,1681,1707]" name="Netherlands" pageId="3" pageNumber="80">Netherlands</collectingCountry>
),
<emphasis id="4BD4AE5EFFE4FF89FC31F94A963E81EE" box="[932,1067,1682,1707]" italics="true" pageId="3" pageNumber="80">P. Moerman</emphasis>
; Passifloratuin (
<collectingCountry id="01B732DCFFE4FF89FB75F949977B81EE" box="[1248,1390,1681,1707]" name="Netherlands" pageId="3" pageNumber="80">Netherlands</collectingCountry>
),
<emphasis id="4BD4AE5EFFE4FF89FAEAF94A92AF818A" italics="true" pageId="3" pageNumber="80">H.v. Aals</emphasis>
t; Portland Botanical Gardens (
<collectingCountry id="01B732DCFFE4FF89FD82F96E90418195" box="[535,596,1718,1744]" name="United States of America" pageId="3" pageNumber="80">USA</collectingCountry>
),
<emphasis id="4BD4AE5EFFE4FF89FDFCF96E90CC818A" box="[617,729,1718,1743]" italics="true" pageId="3" pageNumber="80">D. Pogust</emphasis>
; RBG Kew (
<collectingCountry id="01B732DCFFE4FF89FCFAF96E91898195" box="[879,924,1718,1744]" name="United Kingdom" pageId="3" pageNumber="80">UK</collectingCountry>
),
<emphasis id="4BD4AE5EFFE4FF89FC25F96E9644818A" box="[944,1105,1717,1744]" italics="true" pageId="3" pageNumber="80">C. Magdalena</emphasis>
; The Italian Collection (
<collectingCountry id="01B732DCFFE4FF89FAF2F96D978A818A" box="[1383,1439,1717,1743]" name="Italy" pageId="3" pageNumber="80">Italy</collectingCountry>
),
<emphasis id="4BD4AE5EFFE4FF89FF1DF902931181B6" box="[136,260,1753,1779]" italics="true" pageId="3" pageNumber="80">M. Vecchia</emphasis>
; National
<taxonomicName id="BEA009CFFFE4FF89FEEEF90193FB81B6" box="[379,494,1753,1779]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="80" phylum="Tracheophyta" rank="genus">
<emphasis id="4BD4AE5EFFE4FF89FEEEF90193FB81B6" box="[379,494,1753,1779]" italics="true" pageId="3" pageNumber="80">Passiflora</emphasis>
</taxonomicName>
Collection (
<collectingCountry id="01B732DCFFE4FF89FDECF90290B381B1" box="[633,678,1754,1780]" name="United Kingdom" pageId="3" pageNumber="80">UK</collectingCountry>
),
<emphasis id="4BD4AE5EFFE4FF89FD2FF902917C81B6" box="[698,873,1753,1779]" italics="true" pageId="3" pageNumber="80">J. Vanderplank.</emphasis>
</paragraph>
<paragraph id="791F724CFFE4FF89FF28F92692C48F56" blockId="3.[136,1452,1177,2067]" pageId="3" pageNumber="80">
<emphasis id="4BD4AE5EFFE4FF89FF28F926931C805D" bold="true" box="[189,265,1790,1816]" pageId="3" pageNumber="80">Notes:</emphasis>
—
<taxonomicName id="BEA009CFFFE4FF89FEB3F925900F8052" box="[294,538,1789,1815]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="80" phylum="Tracheophyta" rank="species" species="azulitensis">
<emphasis id="4BD4AE5EFFE4FF89FEB3F925900F8052" box="[294,538,1789,1815]" italics="true" pageId="3" pageNumber="80">Passiflora azulitensis</emphasis>
</taxonomicName>
belongs to
<taxonomicName id="BEA009CFFFE4FF89FD3FF92591ED8052" box="[682,1016,1789,1815]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="80" phylum="Tracheophyta" rank="subGenus" subGenus="Decaloba">
<emphasis id="4BD4AE5EFFE4FF89FD3FF92591088052" box="[682,797,1789,1815]" italics="true" pageId="3" pageNumber="80">Passiflora</emphasis>
subgen.
<emphasis id="4BD4AE5EFFE4FF89FC1EF92591ED8052" box="[907,1016,1789,1815]" italics="true" pageId="3" pageNumber="80">Decaloba</emphasis>
</taxonomicName>
supersect.
<emphasis id="4BD4AE5EFFE4FF89FBEBF92597108052" box="[1150,1285,1789,1815]" italics="true" pageId="3" pageNumber="80">Bryonioides</emphasis>
(Harms) J.M. MacDougal &amp;
<bibRefCitation id="1D310FBDFFE4FF89FEA5F8F990108079" author="Feuillet, C. &amp; MacDougal, J. M." box="[304,517,1825,1852]" pageId="3" pageNumber="80" pagination="34 - 38" refId="ref8477" refString="Feuillet, C. &amp; MacDougal, J. M. (2003) A new infrageneric classification of Passiflora L. (Passifloraceae). Passiflora 13: 34 - 38. https: // doi. org / 10.2307 / 3393379" type="journal article" year="2003">Feuillet (2003: 34)</bibRefCitation>
, for having uncinate foliar vestiture, conspicuous stipules, biglandular petioles, and a distinctive floral structure with a uniserial corona. This alliance of species is mostly confined to the semi-arid habitats of
<collectingCountry id="01B732DCFFE4FF89FF3DF8B192EB80C6" box="[168,254,1897,1923]" name="Mexico" pageId="3" pageNumber="80">Mexico</collectingCountry>
and
<collectingCountry id="01B732DCFFE4FF89FEADF8B193A180C6" box="[312,436,1897,1923]" name="Guatemala" pageId="3" pageNumber="80">Guatemala</collectingCountry>
, with a few of its members spread widely across the South American domains including
<collectingCountry id="01B732DCFFE4FF89FF1DF85592EC80E2" box="[136,249,1933,1959]" name="Colombia" pageId="3" pageNumber="80">Colombia</collectingCountry>
,
<collectingCountry id="01B732DCFFE4FF89FE91F855937680E2" box="[260,355,1933,1959]" name="Ecuador" pageId="3" pageNumber="80">Ecuador</collectingCountry>
, and
<collectingCountry id="01B732DCFFE4FF89FE08F85593F480E2" box="[413,481,1933,1959]" name="Brazil" pageId="3" pageNumber="80">Brazil</collectingCountry>
where it inhabits mostly non-Amazonian habitats like Andean or the Brazilian cerrado (
<bibRefCitation id="1D310FBDFFE4FF89FF05F86993498089" author="MacDougal, J. M." box="[144,348,1969,1996]" pageId="3" pageNumber="80" pagination="1 - 146" refId="ref8920" refString="MacDougal, J. M. (1994) Revision of Passiflora subgenus Decaloba section Pseudodysosmia (Passifloraceae). Systematic Botany Monographs 10: 1 - 146. https: // doi. org / 10.2307 / 25027834" type="journal article" year="1994">MacDougal 1994</bibRefCitation>
).
<taxonomicName id="BEA009CFFFE4FF89FEE7F8699073808E" box="[370,614,1969,1995]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="80" phylum="Tracheophyta" rank="species" species="azulitensis">
<emphasis id="4BD4AE5EFFE4FF89FEE7F8699073808E" box="[370,614,1969,1995]" italics="true" pageId="3" pageNumber="80">Passiflora azulitensis</emphasis>
</taxonomicName>
is remarkable among this group for it being confined solely to a small, tropical wet region of
<collectingCountry id="01B732DCFFE4FF89FE12F80D93E880AA" box="[391,509,2005,2031]" name="Venezuela" pageId="3" pageNumber="80">Venezuela</collectingCountry>
, where it grows in dense premontane rainforest with high levels of humidity year-round.
</paragraph>
<caption id="2DDF22C4FFE3FF8EFF1DF85A964B8F40" ID-DOI="http://doi.org/10.5281/zenodo.14515931" ID-Zenodo-Dep="14515931" httpUri="https://zenodo.org/record/14515931/files/figure.png" pageId="4" pageNumber="81" startId="4.[136,234,1922,1944]" targetBox="[151,1436,190,1898]" targetPageId="4" targetType="figure">
<paragraph id="791F724CFFE3FF8EFF1DF85A964B8F40" blockId="4.[136,1452,1922,2053]" pageId="4" pageNumber="81">
<emphasis id="4BD4AE5EFFE3FF8EFF1DF85A932F80DD" bold="true" box="[136,314,1922,1944]" pageId="4" pageNumber="81">FIGURE 3. A–C</emphasis>
.
<emphasis id="4BD4AE5EFFE3FF8EFEDDF85B900B80DD" box="[328,542,1923,1944]" italics="true" pageId="4" pageNumber="81">
<taxonomicName id="BEA009CFFFE3FF8EFEDDF85B900F80DD" box="[328,538,1923,1944]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="81" phylum="Tracheophyta" rank="species" species="azulitensis">Passiflora azulitensis</taxonomicName>
.
</emphasis>
, Lankaster Composite Dissection Plate (LCDP);
<emphasis id="4BD4AE5EFFE3FF8EFB94F85B960D80DD" bold="true" box="[1025,1048,1923,1944]" pageId="4" pageNumber="81">A.</emphasis>
flower
<emphasis id="4BD4AE5EFFE3FF8EFBF3F85B96B680DD" box="[1126,1187,1923,1944]" italics="true" pageId="4" pageNumber="81">in situ</emphasis>
; longitudinal of the flower (
<emphasis id="4BD4AE5EFFE3FF8EFF05F87F92B880F9" bold="true" box="[144,173,1959,1980]" pageId="4" pageNumber="81">A1</emphasis>
), floral orientation and bracts
<emphasis id="4BD4AE5EFFE3FF8EFE74F87F903580F9" box="[481,544,1959,1980]" italics="true" pageId="4" pageNumber="81">in situ</emphasis>
(
<emphasis id="4BD4AE5EFFE3FF8EFDA4F87F905B80F9" bold="true" box="[561,590,1959,1980]" pageId="4" pageNumber="81">A2</emphasis>
), detail of the bracts (
<emphasis id="4BD4AE5EFFE3FF8EFCA7F87F915A80F9" bold="true" box="[818,847,1959,1980]" pageId="4" pageNumber="81">A3</emphasis>
), detail of the reduced corona (
<emphasis id="4BD4AE5EFFE3FF8EFB05F87F96B880F9" bold="true" box="[1168,1197,1959,1980]" pageId="4" pageNumber="81">A4</emphasis>
);
<emphasis id="4BD4AE5EFFE3FF8EFB50F87F96CE80F9" bold="true" box="[1221,1243,1959,1980]" pageId="4" pageNumber="81">B.</emphasis>
habit and vegetative characteristics of
<emphasis id="4BD4AE5EFFE3FF8EFEA4F81393A180A5" box="[305,436,1995,2016]" italics="true" pageId="4" pageNumber="81">
<taxonomicName id="BEA009CFFFE3FF8EFEA4F81393A480A5" box="[305,433,1995,2016]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="81" phylum="Tracheophyta" rank="species" species="azulitensis">P. azulitensis</taxonomicName>
,
</emphasis>
with detail of the stipules (
<emphasis id="4BD4AE5EFFE3FF8EFD2FF81390C380A5" bold="true" box="[698,726,1995,2016]" pageId="4" pageNumber="81">B1</emphasis>
), petiolar nectaries (
<emphasis id="4BD4AE5EFFE3FF8EFC08F81391AD80A5" bold="true" box="[925,952,1995,2016]" pageId="4" pageNumber="81">B2</emphasis>
), and foliar margin (
<emphasis id="4BD4AE5EFFE3FF8EFB15F813968980A5" bold="true" box="[1152,1180,1995,2016]" pageId="4" pageNumber="81">B3</emphasis>
);
<emphasis id="4BD4AE5EFFE3FF8EFB24F81296DC80A5" bold="true" box="[1201,1225,1994,2016]" pageId="4" pageNumber="81">C.</emphasis>
A mature fruit found
<emphasis id="4BD4AE5EFFE3FF8EFA0CF81392BE8F41" italics="true" pageId="4" pageNumber="81">in situ</emphasis>
. Compiled from photographs taken by Miguel Molinari, Andrés Miguel Orellana and J.R. Kuethe.
</paragraph>
</caption>
<paragraph id="791F724CFFE2FF8FFF28FF4692C38565" blockId="5.[136,1452,158,544]" pageId="5" pageNumber="82">
Vegetatively,
<taxonomicName id="BEA009CFFFE2FF8FFEC2FF46905C87FD" box="[343,585,158,184]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="82" phylum="Tracheophyta" rank="species" species="azulitensis">
<emphasis id="4BD4AE5EFFE2FF8FFEC2FF46905C87FD" box="[343,585,158,184]" italics="true" pageId="5" pageNumber="82">Passiflora azulitensis</emphasis>
</taxonomicName>
is near identical to
<taxonomicName id="BEA009CFFFE2FF8FFCBBFF47969987FD" authority="DC. (1828: 330)" authorityName="DC." authorityPageNumber="330" authorityYear="1828" box="[814,1164,158,184]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="82" phylum="Tracheophyta" rank="species" species="adenopoda">
<emphasis id="4BD4AE5EFFE2FF8FFCBBFF4791D287FD" box="[814,967,158,184]" italics="true" pageId="5" pageNumber="82">P. adenopoda</emphasis>
DC. (1828: 330)
</taxonomicName>
which can also be found in the Venezuelan Andes but is spread widely across the northern Andean countries including
<collectingCountry id="01B732DCFFE2FF8FFB37FF1A96EA8799" box="[1186,1279,194,220]" name="Ecuador" pageId="5" pageNumber="82">Ecuador</collectingCountry>
and
<collectingCountry id="01B732DCFFE2FF8FFAA3FF1A97BD8799" box="[1334,1448,194,220]" name="Colombia" pageId="5" pageNumber="82">Colombia</collectingCountry>
. Being variable in nature, the leaves, stipules and petiolar nectaries of
<taxonomicName id="BEA009CFFFE2FF8FFC05FF3F96328645" box="[912,1063,230,256]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="82" phylum="Tracheophyta" rank="species" species="adenopoda">
<emphasis id="4BD4AE5EFFE2FF8FFC05FF3F96328645" box="[912,1063,230,256]" italics="true" pageId="5" pageNumber="82">P. adenopoda</emphasis>
</taxonomicName>
share identical shapes and sizes to
<taxonomicName id="BEA009CFFFE2FF8FFF1DFED3930D8661" box="[136,280,266,292]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="82" phylum="Tracheophyta" rank="species" species="azulitensis">
<emphasis id="4BD4AE5EFFE2FF8FFF1DFED3930D8661" box="[136,280,266,292]" italics="true" pageId="5" pageNumber="82">P. azulitensis</emphasis>
</taxonomicName>
, however, the radically different floral and habitat characteristics warrants the recognition of
<taxonomicName id="BEA009CFFFE2FF8FFA8EFED397BE8661" box="[1307,1451,266,292]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="82" phylum="Tracheophyta" rank="species" species="azulitensis">
<emphasis id="4BD4AE5EFFE2FF8FFA8EFED397BE8661" box="[1307,1451,266,292]" italics="true" pageId="5" pageNumber="82">P. azulitensis</emphasis>
</taxonomicName>
as an independent taxon. The flowers of
<taxonomicName id="BEA009CFFFE2FF8FFDCCFEF790F9860D" box="[601,748,302,328]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="82" phylum="Tracheophyta" rank="species" species="azulitensis">
<emphasis id="4BD4AE5EFFE2FF8FFDCCFEF790F9860D" box="[601,748,302,328]" italics="true" pageId="5" pageNumber="82">P. azulitensis</emphasis>
</taxonomicName>
are entirely white, upright to subhorizontal, and measure less than
<quantity id="BE58DFA9FFE2FF8FFF55FE8A93188628" box="[192,269,338,365]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="82" unit="cm" value="2.5">2.5 cm</quantity>
in diameter. In contrast, the flowers of
<taxonomicName id="BEA009CFFFE2FF8FFD58FE8B91708629" box="[717,869,338,364]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="82" phylum="Tracheophyta" rank="species" species="adenopoda">
<emphasis id="4BD4AE5EFFE2FF8FFD58FE8B91708629" box="[717,869,338,364]" italics="true" pageId="5" pageNumber="82">P. adenopoda</emphasis>
</taxonomicName>
are white with purple bands, pendent, and measure 4–5(–6) cm in diameter.
<taxonomicName id="BEA009CFFFE2FF8FFE31FEAE908286D5" box="[420,663,374,400]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="82" phylum="Tracheophyta" rank="species" species="azulitensis">
<emphasis id="4BD4AE5EFFE2FF8FFE31FEAE908286D5" box="[420,663,374,400]" italics="true" pageId="5" pageNumber="82">Passiflora azulitensis</emphasis>
</taxonomicName>
also shares some resemblance to
<taxonomicName id="BEA009CFFFE2FF8FFB88FEAF976F86D4" authority="Mast. (1872: 555)" authorityName="Mast." authorityPageNumber="555" authorityYear="1872" box="[1053,1402,374,401]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="82" phylum="Tracheophyta" rank="species" species="morifolia">
<emphasis id="4BD4AE5EFFE2FF8FFB88FEAF96B686D5" box="[1053,1187,374,400]" italics="true" pageId="5" pageNumber="82">P. morifolia</emphasis>
Mast. (1872: 555)
</taxonomicName>
and
<taxonomicName id="BEA009CFFFE2FF8FFF1DFE43908C86F1" authority="Conejo &amp; Kuethe (2022: 2)" authorityName="Conejo &amp; Kuethe" authorityPageNumber="2" authorityYear="2022" box="[136,665,410,436]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="82" phylum="Tracheophyta" rank="species" species="guayaquilensis">
<emphasis id="4BD4AE5EFFE2FF8FFF1DFE43935B86F1" box="[136,334,410,436]" italics="true" pageId="5" pageNumber="82">P. guayaquilensis</emphasis>
Conejo &amp;
<bibRefCitation id="1D310FBDFFE2FF8FFE47FE42908C86F1" author="Kuethe, J. R. &amp; Vanderplank, J." box="[466,665,410,436]" pageId="5" pageNumber="82" pagination="159 - 165" refId="ref8847" refString="Kuethe, J. R. &amp; Vanderplank, J. (2022) Passiflora mistratensis, a new species of Passiflora (Passifloraceae) from Colombia, commonly known from European cultivation. Phytotaxa 552: 159 - 165. https: // doi. org / 10.11646 / phytotaxa. 552.2.3" type="journal article" year="2022">Kuethe (2022: 2)</bibRefCitation>
</taxonomicName>
, the latter of which sharing the ubiquitously white coloration of the corolla. However, it differs by the foliaceous serrate, ciliolate stipules that measure up to
<quantity id="BE58DFA9FFE2FF8FFBFFFE6696B4869C" box="[1130,1185,446,473]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="82" unit="cm" value="2.0">2 cm</quantity>
in length: while in both
<taxonomicName id="BEA009CFFFE2FF8FFF1DFE3B931E86B9" box="[136,267,482,508]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="82" phylum="Tracheophyta" rank="species" species="morifolia">
<emphasis id="4BD4AE5EFFE2FF8FFF1DFE3B931E86B9" box="[136,267,482,508]" italics="true" pageId="5" pageNumber="82">P. morifolia</emphasis>
</taxonomicName>
and
<taxonomicName id="BEA009CFFFE2FF8FFED4FE3B901686B9" authorityName="Conejo &amp; Kuethe" authorityYear="2022" box="[321,515,482,508]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="82" phylum="Tracheophyta" rank="species" species="guayaquilensis">
<emphasis id="4BD4AE5EFFE2FF8FFED4FE3B901686B9" box="[321,515,482,508]" italics="true" pageId="5" pageNumber="82">P. guayaquilensis</emphasis>
</taxonomicName>
the stipules are ovate-caudate with entire margin and measure no more than
<quantity id="BE58DFA9FFE2FF8FFACDFE3A979A86B8" box="[1368,1423,482,509]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="82" unit="cm" value="1.0">1 cm</quantity>
in length.
</paragraph>
</subSubSection>
</treatment>
</document>