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<document id="D938FF190E2584C967677D50D51719BA" ID-DOI="10.5281/zenodo.12766004" ID-ISSN="1989-6581" ID-Zenodo-Dep="12766004" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721246549113" checkinUser="felipe" docAuthor="Fa ̇ ndez, Eduardo I. &amp; Rider, David A." docDate="2016" docId="ED6FFF21D020A402FF66FB5DB452AA61" docLanguage="es" docName="ArqEntomol.15.383-389.pdf" docOrigin="Arquivos Entomolóxicos 15" docStyle="DocumentStyle:DF23C849EA72F80F6DF49FA5382909D5.2:ArqEntomol.2008-.journal_article.1column" docStyleId="DF23C849EA72F80F6DF49FA5382909D5" docStyleName="ArqEntomol.2008-.journal_article.1column" docStyleVersion="2" docTitle="Acledra (Acledra) albocostata subsp. lueri ndez &amp; Rider, 2016, n. ssp." docType="treatment" docVersion="2" lastPageNumber="386" masterDocId="11568759D022A401FFD0FFF9B010AE58" masterDocTitle="Notas sobre el complejo de Acledra (Acledra) albocostata (Spinola, 1852) (Hemiptera: Heteroptera: Pentatomidae: Pentatominae)." masterLastPageNumber="389" masterPageNumber="383" pageNumber="385" updateTime="1721253630782" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="F1655188A72BDD5DC86AF70735495F54" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6386BE7888A6AE2FAACF1302A9CE98E6">
<mods:title id="3359A0F1F9B1047344DCE59B4B014DFE">Notas sobre el complejo de Acledra (Acledra) albocostata (Spinola, 1852) (Hemiptera: Heteroptera: Pentatomidae: Pentatominae).</mods:title>
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<mods:name id="08C36404F37FC8E22D29CDB81695AF1B" type="personal">
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<mods:namePart id="4A27A7A234D4FAEF2559D195464BCB5C">Fa ̇ ndez, Eduardo I.</mods:namePart>
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<mods:namePart id="8828402D5D2D51C5F821729ACEEA60E4">Rider, David A.</mods:namePart>
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<mods:typeOfResource id="CD0A9B7C39CCC8C82FA25C50B27DE528">text</mods:typeOfResource>
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<mods:title id="2CA88C133449D5AFB84EEF36B785B88C">Arquivos Entomolóxicos</mods:title>
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<mods:part id="910B66E18CFA4E6A5F24A914C4B0705A">
<mods:date id="27FB34C39C3A4BDC253F92DF65AB24C3">2016</mods:date>
<mods:detail id="5A59043806C3D3D32EF01FD5910212E7" type="pubDate">
<mods:number id="9BA5EF52D48FFE199FE1CA705D9EDDFE">2016-06-30</mods:number>
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<mods:number id="50F73B7085B49CA0EA3D183F60401205">15</mods:number>
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<mods:classification id="6A635FAB0AB80A94F1BD01B4B9889FE4">journal article</mods:classification>
<mods:identifier id="8B2F33E86FA350E377BF77DACAE7F801" type="DOI">10.5281/zenodo.12766004</mods:identifier>
<mods:identifier id="5CB2CCA3497A413C3CF0D57848965ADE" type="ISSN">1989-6581</mods:identifier>
<mods:identifier id="6E2A579D8F79DAF46CC51AB6B16A3B29" type="Zenodo-Dep">12766004</mods:identifier>
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<treatment id="ED6FFF21D020A402FF66FB5DB452AA61" ID-DOI="http://doi.org/10.5281/zenodo.12766040" ID-Zenodo-Dep="12766040" LSID="urn:lsid:plazi:treatment:ED6FFF21D020A402FF66FB5DB452AA61" httpUri="http://treatment.plazi.org/id/ED6FFF21D020A402FF66FB5DB452AA61" lastPageId="3" lastPageNumber="386" pageId="2" pageNumber="385">
<subSubSection id="2DDC1DBCD020A403FF66FB5DB4C8AAAA" pageId="2" pageNumber="385" type="nomenclature">
<paragraph id="65794E37D020A403FF66FB5DB3EFAAE5" blockId="2.[182,1023,1188,1213]" box="[182,1023,1188,1213]" pageId="2" pageNumber="385">
<heading id="3E31F95BD020A403FF66FB5DB3EFAAE5" bold="true" box="[182,1023,1188,1213]" fontSize="10" level="2" pageId="2" pageNumber="385" reason="6">
<emphasis id="57B29225D020A403FF66FB5DB3EFAAE5" bold="true" box="[182,1023,1188,1213]" pageId="2" pageNumber="385">
<taxonomicName id="A2C635B4D020A403FF66FB5DB26CAAE5" authorityName="Fȧndez &amp; Rider" authorityYear="2016" box="[182,636,1188,1213]" class="Insecta" family="Pentatomidae" genus="Acledra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="385" phylum="Arthropoda" rank="subSpecies" species="albocostata" status="subsp. nov." subGenus="Acledra" subSpecies="lueri">
<emphasis id="57B29225D020A403FF66FB5DB26CAAE5" bold="true" box="[182,636,1188,1213]" italics="true" pageId="2" pageNumber="385">Acledra (Acledra) albocostata lueri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C812F5ED020A403FD55FB5DB2C1AAE5" box="[645,721,1188,1213]" pageId="2" pageNumber="385" rank="subSpecies">n. ssp.</taxonomicNameLabel>
(
<figureCitation id="FDFD52B2D020A403FD30FB5DB365AAE5" box="[736,885,1188,1213]" captionStart="Figs" captionStartId="5.[168,215,1887,1907]" captionTargetBox="[148,1445,612,1846]" captionTargetId="figure-114@5.[148,1445,610,1847]" captionTargetPageId="5" captionText="Figs. 1-9.- Acledra (Acledra) albocostata. 1.- A. (A.) albocostata albocostata, habitus. 2.- A. (A.) albocostata carinata, habitus. 3.- A. (A.) albocostata lueri, habitus. 4.- A. (A.) albocostata albocostata, pigóforo en vista dorsal. 5.- A. (A.) albocostata carinata, pigóforo en vista dorsal. 6.- A. (A.) albocostata lueri, pigóforo en vista dorsal. 7.- A. (A.) albocostata albocostata, pigóforo en vista ventral. 8.- A. (A.) albocostata carinata, pigóforo en vista ventral. 9.- A. (A.) albocostata lueri, pigóforo en vista ventral." figureDoi="http://doi.org/10.5281/zenodo.12766006" httpUri="https://zenodo.org/record/12766006/files/figure.png" pageId="2" pageNumber="385">Figs. 3, 6, 9</figureCitation>
,
<figureCitation id="FDFD52B2D020A403FC52FB5DB3E7AAE5" box="[898,1015,1188,1213]" captionStart="Figs" captionStartId="6.[198,245,1751,1771]" captionTargetBox="[198,1398,219,1701]" captionTargetId="figure-18@6.[198,1408,219,1701]" captionTargetPageId="6" captionText="Figs. 10-14.- Acledra (Acledra) albocostata. 10.- A. (A.) albocostata albocostata, pronoto. 11.- A. (A.) albocostata carinata, pronoto. 12.- A. (A.) albocostata lueri, pronoto. 13.- A. (A.) albocostata lueri, terminalia femenina. 14.- Distribucion de A. (A) albocostata: área azul, A. (A.) albocostata albocostata; círculos rojos, A. (A.) albocostata carinata; cuadrados verdes, A. (A.) albocostata lueri." figureDoi="http://doi.org/10.5281/zenodo.12766008" httpUri="https://zenodo.org/record/12766008/files/figure.png" pageId="2" pageNumber="385">12, 13, 14</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="65794E37D020A403FF3EFB21B4C8AAAA" blockId="2.[182,1444,1240,2047]" box="[238,1240,1240,1267]" pageId="2" pageNumber="385">
<emphasis id="57B29225D020A403FF3EFB21B4C8AAAA" bold="true" box="[238,1240,1240,1267]" pageId="2" pageNumber="385">
Descripción: Cuerpo alargado, ocreáceo con puntuación densa y concolora (
<figureCitation id="FDFD52B2D020A403FB52FB20B4DAAAAA" box="[1154,1226,1241,1266]" captionStart="Figs" captionStartId="5.[168,215,1887,1907]" captionTargetBox="[148,1445,612,1846]" captionTargetId="figure-114@5.[148,1445,610,1847]" captionTargetPageId="5" captionText="Figs. 1-9.- Acledra (Acledra) albocostata. 1.- A. (A.) albocostata albocostata, habitus. 2.- A. (A.) albocostata carinata, habitus. 3.- A. (A.) albocostata lueri, habitus. 4.- A. (A.) albocostata albocostata, pigóforo en vista dorsal. 5.- A. (A.) albocostata carinata, pigóforo en vista dorsal. 6.- A. (A.) albocostata lueri, pigóforo en vista dorsal. 7.- A. (A.) albocostata albocostata, pigóforo en vista ventral. 8.- A. (A.) albocostata carinata, pigóforo en vista ventral. 9.- A. (A.) albocostata lueri, pigóforo en vista ventral." figureDoi="http://doi.org/10.5281/zenodo.12766006" httpUri="https://zenodo.org/record/12766006/files/figure.png" pageId="2" pageNumber="385">Fig. 3</figureCitation>
).
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="2DDC1DBCD020A402FF3EFB07B483AC69" lastPageId="3" lastPageNumber="386" pageId="2" pageNumber="385" type="description">
<paragraph id="65794E37D020A403FF3EFB07B476ABDE" blockId="2.[182,1444,1240,2047]" pageId="2" pageNumber="385">
<emphasis id="57B29225D020A403FF3EFB07B476ABDE" bold="true" pageId="2" pageNumber="385">Cabeza: más larga que ancha; ojos prominentes; ocelos ubicados detrás de los ojos, no contiguos con el pronoto; paraclípeos gruesos, levemente pasando el anteclípeo, no contiguos; tubérculo antenífero con pequeño diente lateral, antenas unicolores (ocre a rojizo), primer antenómero no sobrepasa el final de la cabeza; rostrum alargado, alcanzando las metacoxas.</emphasis>
</paragraph>
<paragraph id="65794E37D020A403FF3EFA6BB134A88D" blockId="2.[182,1444,1240,2047]" pageId="2" pageNumber="385">
<emphasis id="57B29225D020A403FF3EFA6BB134A88D" bold="true" pageId="2" pageNumber="385">
Tórax: Pronoto densamente puntuado (
<figureCitation id="FDFD52B2D020A403FD00FA6BB338ABF3" box="[720,808,1426,1451]" captionStart="Figs" captionStartId="6.[198,245,1751,1771]" captionTargetBox="[198,1398,219,1701]" captionTargetId="figure-18@6.[198,1408,219,1701]" captionTargetPageId="6" captionText="Figs. 10-14.- Acledra (Acledra) albocostata. 10.- A. (A.) albocostata albocostata, pronoto. 11.- A. (A.) albocostata carinata, pronoto. 12.- A. (A.) albocostata lueri, pronoto. 13.- A. (A.) albocostata lueri, terminalia femenina. 14.- Distribucion de A. (A) albocostata: área azul, A. (A.) albocostata albocostata; círculos rojos, A. (A.) albocostata carinata; cuadrados verdes, A. (A.) albocostata lueri." figureDoi="http://doi.org/10.5281/zenodo.12766008" httpUri="https://zenodo.org/record/12766008/files/figure.png" pageId="2" pageNumber="385">Fig. 13</figureCitation>
), sin parches o líneas impuntuadas, lados cóncavos, ángulos anterolaterales crenulados, ángulos posterolaterales redondeados, cicatrices inmaculadas; escutelo triangular (isósceles), ápice redondeado, y una gruesa carena marfileña a lo largo de la línea media, ángulos basales con dos manchas marfileñas gruesas y puntuadas; hemiélitros sobrepasando el final del abdomen,
<emphasis id="57B29225D020A403FE72F9DEB1E5A818" bold="true" box="[418,501,1575,1600]" italics="true" pageId="2" pageNumber="385">corium</emphasis>
con una carena marfileña a lo largo de la vena radial seguido de un pequeño parche impuntuado entre la vena radial y el borde del
<emphasis id="57B29225D020A403FC88F9B5B3BAA83D" bold="true" box="[856,938,1612,1637]" italics="true" pageId="2" pageNumber="385">corium</emphasis>
, membrana ahumada con ṁltiples venas logitudinales; peritrema ostiolar corto (alcanzando 1/3 de la metapleura), evaporatoria rugosa pasando levemente la mitad de la metapleura; patas cubiertas de pilosidad color caramelo, tibias sulcadas.
</emphasis>
</paragraph>
<paragraph id="65794E37D020A402FF3EF918B234AF3C" blockId="2.[182,1444,1240,2047]" lastBlockId="3.[182,1443,219,356]" lastPageId="3" lastPageNumber="386" pageId="2" pageNumber="385">
<emphasis id="57B29225D020A402FF3EF918B234AF3C" bold="true" lastPageId="3" lastPageNumber="386" pageId="2" pageNumber="385">
Abdomen: base del abdomen sin espina o tubérculo, peritrema espiracular color caramelo, conexiva aserrada. Genitalia femenina: espermateca alargada y oblonga, bastón esclerotizado grueso con la base ensanchada, ápice agudo precedido de una dilatación circular, dilatación del ducto espermatecal escasamente inflada, rodeando el bastón esclerotizado, reborde proximal de la bomba ovalado y plano, reborde distal de la bomba subcónico, bomba espermatecal ovalada; primeras gonocoxas fuertemente convexas posteriormente (
<figureCitation id="FDFD52B2D020A403FCE1F862B399A9EC" box="[817,905,1947,1972]" captionStart="Figs" captionStartId="6.[198,245,1751,1771]" captionTargetBox="[198,1398,219,1701]" captionTargetId="figure-18@6.[198,1408,219,1701]" captionTargetPageId="6" captionText="Figs. 10-14.- Acledra (Acledra) albocostata. 10.- A. (A.) albocostata albocostata, pronoto. 11.- A. (A.) albocostata carinata, pronoto. 12.- A. (A.) albocostata lueri, pronoto. 13.- A. (A.) albocostata lueri, terminalia femenina. 14.- Distribucion de A. (A) albocostata: área azul, A. (A.) albocostata albocostata; círculos rojos, A. (A.) albocostata carinata; cuadrados verdes, A. (A.) albocostata lueri." figureDoi="http://doi.org/10.5281/zenodo.12766008" httpUri="https://zenodo.org/record/12766008/files/figure.png" pageId="2" pageNumber="385">Fig. 13</figureCitation>
), sección caudal de cada gonocoxito recta; segundas gonocoxas ovaladas y poco conspicuas; paraterguitos 8 subromboidales con el ápice romo; paraterguitos 9 subtriangulares, ápice romo y lados exteriores con una leve concavidad. Genitalia masculina: Pigóforo subrectangular (
<figureCitation id="FDFD52B2D021A402FDA4FF25B2FAAEAD" box="[628,746,220,245]" captionStart="Figs" captionStartId="5.[168,215,1887,1907]" captionTargetBox="[148,1445,612,1846]" captionTargetId="figure-114@5.[148,1445,610,1847]" captionTargetPageId="5" captionText="Figs. 1-9.- Acledra (Acledra) albocostata. 1.- A. (A.) albocostata albocostata, habitus. 2.- A. (A.) albocostata carinata, habitus. 3.- A. (A.) albocostata lueri, habitus. 4.- A. (A.) albocostata albocostata, pigóforo en vista dorsal. 5.- A. (A.) albocostata carinata, pigóforo en vista dorsal. 6.- A. (A.) albocostata lueri, pigóforo en vista dorsal. 7.- A. (A.) albocostata albocostata, pigóforo en vista ventral. 8.- A. (A.) albocostata carinata, pigóforo en vista ventral. 9.- A. (A.) albocostata lueri, pigóforo en vista ventral." figureDoi="http://doi.org/10.5281/zenodo.12766006" httpUri="https://zenodo.org/record/12766006/files/figure.png" pageId="3" pageNumber="386">Figs. 6, 9</figureCitation>
), vista ventral con el ápice bilobulado, en vista dorsal se aprecia un puente apical con dos lóbulos gruesos y una fóvea media apreciable en vista caudal, parámeros con forma de bastón, lóbulo basal más delgado que el apical, lóbulo apical con un diente amplio redondeado en la base.
</emphasis>
</paragraph>
<paragraph id="65794E37D021A402FF3EFE7AB51DAFC5" blockId="3.[182,1444,387,562]" box="[238,1293,387,413]" pageId="3" pageNumber="386">
<materialsCitation id="D5AE446AD021A402FF3EFE7AB519AFC5" box="[238,1289,387,413]" collectingDate="2014-02-28" collectionCode="HRCC" country="Chile" location="Region de Aysen" pageId="3" pageNumber="386" specimenCount="1" specimenCount-male="1" stateProvince="Aysen" typeStatus="Holotipo">
<emphasis id="57B29225D021A402FF3EFE7AB484AFC4" bold="true" box="[238,1172,387,413]" pageId="3" pageNumber="386">
<typeStatus id="BA7DF095D021A402FF3EFE7AB14AAFC4" box="[238,346,387,412]" pageId="3" pageNumber="386" type="holotype">Holotipo</typeStatus>
:
<collectingCountry id="1DD10EA7D021A402FEB6FE7AB1B6AFC5" box="[358,422,387,413]" name="Chile" pageId="3" pageNumber="386">Chile</collectingCountry>
,
<collectingRegion id="A70280D5D021A402FE62FE7AB290AFC5" box="[434,640,387,413]" country="Chile" name="Aysen" pageId="3" pageNumber="386">Región de Aysén</collectingRegion>
,
<collectingRegion id="A70280D5D021A402FD5CFE7AB301AFC4" box="[652,785,387,412]" country="Chile" name="Balmaceda" pageId="3" pageNumber="386">Balmaceda</collectingRegion>
, 27/
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<collectingDate id="013C911FD021A402FC98FE7AB3CFAFC4" box="[840,991,387,412]" pageId="3" pageNumber="386" value="2014-02-28">28-II-2014</collectingDate>
</date>
, leg. A.
<collectingRegion id="A70280D5D021A402FB90FE7AB46BAFC5" box="[1088,1147,387,413]" country="Chile" name="Luer" pageId="3" pageNumber="386">Lüer</collectingRegion>
,
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</emphasis>
♂
<emphasis id="57B29225D021A402FB7EFE7AB519AFC5" bold="true" box="[1198,1289,387,413]" pageId="3" pageNumber="386">
(
<collectionCode id="03D7D6F2D021A402FB65FE7AB512AFC5" box="[1205,1282,387,413]" pageId="3" pageNumber="386">HRCC</collectionCode>
)
</emphasis>
</materialsCitation>
<emphasis id="57B29225D021A402FAD9FE7AB51DAFC5" bold="true" box="[1289,1293,387,413]" pageId="3" pageNumber="386">.</emphasis>
</paragraph>
<paragraph id="65794E37D021A402FF3EFE51B483AC69" blockId="3.[182,1444,387,562]" pageId="3" pageNumber="386">
<materialsCitation id="D5AE446AD021A402FF3EFE51B58FAF9A" box="[238,1439,424,450]" collectingDate="1968-02-09" collectionCode="DARC, IPUM, HRCC" country="Argentina" location="Epuyen" pageId="3" pageNumber="386" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Chubut" typeStatus="Paratipos">
<emphasis id="57B29225D021A402FF3EFE51B38DAF99" bold="true" box="[238,925,424,450]" pageId="3" pageNumber="386">
<typeStatus id="BA7DF095D021A402FF3EFE51B175AF99" box="[238,357,424,449]" pageId="3" pageNumber="386" type="paratype">Paratipos</typeStatus>
:
<collectingCountry id="1DD10EA7D021A402FEADFE51B1EAAF99" box="[381,506,424,449]" name="Argentina" pageId="3" pageNumber="386">Argentina</collectingCountry>
,
<collectingRegion id="A70280D5D021A402FDC1FE51B27CAF9A" box="[529,620,424,450]" country="Argentina" name="Chubut" pageId="3" pageNumber="386">Chubut</collectingRegion>
,
<location id="601918ECD021A402FD53FE51B2CEAF99" LSID="urn:lsid:plazi:treatment:ED6FFF21D020A402FF66FB5DB452AA61:601918ECD021A402FD53FE51B2CEAF99" box="[643,734,424,449]" country="Argentina" name="Epuyen" pageId="3" pageNumber="386" stateProvince="Chubut">Epuyen</location>
,
<date id="117868F7D021A402FD25FE51B369AF99" box="[757,889,424,449]" pageId="3" pageNumber="386" value="1968-02-09">
<collectingDate id="013C911FD021A402FD25FE51B369AF99" box="[757,889,424,449]" pageId="3" pageNumber="386" value="1968-02-09">9-II-1968</collectingDate>
</date>
,
<specimenCount id="73C085BED021A402FC41FE51B38DAF99" box="[913,925,424,449]" count="1" pageId="3" pageNumber="386" type="male">1</specimenCount>
</emphasis>
♂,
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<emphasis id="57B29225D021A402FC1AFE51B3C6AF99" bold="true" box="[970,982,424,449]" pageId="3" pageNumber="386">1</emphasis>
♀
</specimenCount>
<emphasis id="57B29225D021A402FC2DFE51B4F1AF99" bold="true" box="[1021,1249,424,450]" pageId="3" pageNumber="386">
(
<collectionCode id="03D7D6F2D021A402FBD4FE51B441AF9A" box="[1028,1105,424,450]" pageId="3" pageNumber="386">DARC</collectionCode>
;
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),
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♂,
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<emphasis id="57B29225D021A402FAC0FE51B50CAF99" bold="true" box="[1296,1308,424,449]" pageId="3" pageNumber="386">1</emphasis>
♀
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(
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)
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</materialsCitation>
<emphasis id="57B29225D021A402FA4FFE51B5B3AF9A" bold="true" box="[1439,1443,424,450]" pageId="3" pageNumber="386">;</emphasis>
<materialsCitation id="D5AE446AD021A402FF66FE37B3C7AFBF" box="[182,983,461,487]" collectionCode="DARC" collectorName="El Bolson &amp; A. Kovacs" country="Argentina" location="Rio Negro" pageId="3" pageNumber="386" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="Paratipos">
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<collectingCountry id="1DD10EA7D021A402FF66FE37B124AFBF" box="[182,308,462,487]" name="Argentina" pageId="3" pageNumber="386">Argentina</collectingCountry>
,
<location id="601918ECD021A402FE93FE34B1D4AFBF" LSID="urn:lsid:plazi:treatment:ED6FFF21D020A402FF66FB5DB452AA61:601918ECD021A402FE93FE34B1D4AFBF" box="[323,452,461,487]" country="Argentina" name="Rio Negro" pageId="3" pageNumber="386">Río Negro</location>
,
<collectorName id="C8332BE1D021A402FE04FE37B259AFBF" box="[468,585,461,487]" pageId="3" pageNumber="386">El Bolsón</collectorName>
, leg.
<collectorName id="C8332BE1D021A402FD5FFE37B31BAFBF" box="[655,779,462,487]" pageId="3" pageNumber="386">A. Kovacs</collectorName>
,
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</emphasis>
♂,
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<emphasis id="57B29225D021A402FC9DFE37B349AFBF" bold="true" box="[845,857,462,487]" pageId="3" pageNumber="386">1</emphasis>
♀
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(
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)
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<materialsCitation id="D5AE446AD021A402FC37FE37B2E6AC54" collectingDate="1985-09-05" collectionCode="USNM" collectorName="L. Pena" country="Argentina" county="Rio Negro" location="El Maiten" municipality="Bariloche" pageId="3" pageNumber="386" specimenCount="1" specimenCount-male="1" typeStatus="Paratipos">
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<collectingCountry id="1DD10EA7D021A402FC37FE37B475AFBF" box="[999,1125,462,487]" name="Argentina" pageId="3" pageNumber="386">Argentina</collectingCountry>
,
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,
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,
<location id="601918ECD021A402FA5BFE37B11FAC54" LSID="urn:lsid:plazi:treatment:ED6FFF21D020A402FF66FB5DB452AA61:601918ECD021A402FA5BFE37B11FAC54" country="Argentina" county="Rio Negro" municipality="Bariloche" name="El Maiten" pageId="3" pageNumber="386">El Maiten</location>
,
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<collectingDate id="013C911FD021A402FEF0FE0AB1B8AC54" box="[288,424,499,524]" pageId="3" pageNumber="386" value="1985-09-05">5-IX-1985</collectingDate>
</date>
, leg.
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,
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</emphasis>
♂
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(
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)
</emphasis>
</materialsCitation>
<emphasis id="57B29225D021A402FD26FE0BB2EBAC54" bold="true" box="[758,763,498,524]" pageId="3" pageNumber="386">;</emphasis>
<materialsCitation id="D5AE446AD021A402FCD8FE0AB49EAC69" collectingDate="1986-01-22" collectionCode="AMNH" collectorName="Schuh &amp; Muddy" country="Argentina" location="12 km E of Esquivel" pageId="3" pageNumber="386" specimenCount="1" specimenCount-female="1" stateProvince="Chubut" typeStatus="Paratipos">
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<collectingCountry id="1DD10EA7D021A402FCD8FE0AB396AC54" box="[776,902,499,524]" name="Argentina" pageId="3" pageNumber="386">Argentina</collectingCountry>
,
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,
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<quantity id="A23EE3D2D021A402FBD3FE0AB45FAC54" box="[1027,1103,499,524]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="386" unit="km" value="12.0">12 km</quantity>
E of Esquivel
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,
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<collectingDate id="013C911FD021A402FACAFE0AB5B0AC54" box="[1306,1440,499,524]" pageId="3" pageNumber="386" value="1986-01-22">22-I-1986</collectingDate>
</date>
,
<collectorName id="C8332BE1D021A402FF66FDE1B113AC6A" box="[182,259,536,562]" pageId="3" pageNumber="386">Schuh</collectorName>
&amp; Platnick leg.
<collectorName id="C8332BE1D021A402FE6FFDE1B202AC69" box="[447,530,536,561]" pageId="3" pageNumber="386">Muddy</collectorName>
shore of pond w/scattered vegetation
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♀
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(
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)
</emphasis>
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<emphasis id="57B29225D021A402FB5EFDE1B483AC69" bold="true" box="[1166,1171,536,561]" pageId="3" pageNumber="386">.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="2DDC1DBCD021A402FF3EFDB6B452AA61" pageId="3" pageNumber="386" type="diagnosis">
<paragraph id="65794E37D021A402FF3EFDB6B563AD84" blockId="3.[182,1444,591,988]" pageId="3" pageNumber="386">
<emphasis id="57B29225D021A402FF3EFDB6B563AD84" bold="true" pageId="3" pageNumber="386">
Diagnosis diferencial: Esta subespecie puede reconocerse dentro del complejo de
<taxonomicName id="A2C635B4D021A402FB20FDB6B5B3AC30" baseAuthorityName="Spinola" baseAuthorityYear="1852" box="[1264,1443,591,616]" class="Insecta" family="Pentatomidae" genus="Acledra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="albocostata">
<emphasis id="57B29225D021A402FB20FDB6B5B3AC30" bold="true" box="[1264,1443,591,616]" italics="true" pageId="3" pageNumber="386">A. albocostata</emphasis>
</taxonomicName>
por las siguientes características: de
<taxonomicName id="A2C635B4D021A402FD61FD8DB45AACD5" baseAuthorityName="Spinola" baseAuthorityYear="1852" box="[689,1098,628,653]" class="Insecta" family="Pentatomidae" genus="Acledra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="subSpecies" species="albocostata" subGenus="Acledra" subSpecies="albocostata">
<emphasis id="57B29225D021A402FD61FD8DB45AACD5" bold="true" box="[689,1098,628,653]" italics="true" pageId="3" pageNumber="386">A. (A.) albocostata albocostata</emphasis>
</taxonomicName>
se diferencia por poseer callosidades laterales del escutelo más reducidas, también en la genitalia masculina, los lóbulos del puente superior del pigóforo son mucho más amplios en
<taxonomicName id="A2C635B4D021A402FCBDFD46B487AC80" authorityName="Fȧndez &amp; Rider" authorityYear="2016" box="[877,1175,703,728]" class="Insecta" family="Pentatomidae" genus="Acledra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="subSpecies" species="albocostata" subGenus="Acledra" subSpecies="lueri">
<emphasis id="57B29225D021A402FCBDFD46B487AC80" bold="true" box="[877,1175,703,728]" italics="true" pageId="3" pageNumber="386">A. (A.) albocostata lueri</emphasis>
</taxonomicName>
, carácter mucho más notorio en vista ventral. De
<taxonomicName id="A2C635B4D021A402FDF8FD1DB39DACA5" authorityName="Breddin" authorityYear="1914" baseAuthorityName="Breddin" baseAuthorityYear="1914" box="[552,909,740,765]" class="Insecta" family="Pentatomidae" genus="Acledra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="subSpecies" species="albocostata" subGenus="Acledra" subSpecies="carinata">
<emphasis id="57B29225D021A402FDF8FD1DB39DACA5" bold="true" box="[552,909,740,765]" italics="true" pageId="3" pageNumber="386">A. (A.) albocostata carinata</emphasis>
</taxonomicName>
se distingue por poseer un escutelo más ensanchado en el ápice, pronoto más compacto, en la genitalia femenina los paraterguitos 8 y 9 son más compactos, y en la genitalia masculina los lóbulos el puente superior del pigóforo son mucho más amplios. Fuera del grupo de
<taxonomicName id="A2C635B4D021A402FDCCFCAAB2C1AD34" baseAuthorityName="Spinola" baseAuthorityYear="1852" box="[540,721,851,876]" class="Insecta" family="Pentatomidae" genus="Acledra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="albocostata">
A.
<emphasis id="57B29225D021A402FD91FCAAB2C1AD34" bold="true" box="[577,721,851,876]" italics="true" pageId="3" pageNumber="386">albocostata</emphasis>
</taxonomicName>
, esta subespecie tiene similitudes con
<taxonomicName id="A2C635B4D021A402FB13FCAAB18CADC9" authority="Fa" authorityName="Fa" class="Insecta" family="Pentatomidae" genus="Acledra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="bichromocornis" subGenus="Acledra">
<emphasis id="57B29225D021A402FB13FCAAB162ADCA" bold="true" italics="true" pageId="3" pageNumber="386">Acledra (Acledra) bichromocornis</emphasis>
Fa
</taxonomicName>
̇ndez,
<bibRefCitation id="015733C6D021A402FE21FC80B317ADCA" author="Rider, D. A. &amp; Carvajal, M. A." box="[497,775,888,914]" pageId="3" pageNumber="386" pagination="127 - 134" refId="ref3590" refString="Fa ˙ ndez, E. I.; Rider, D. A. &amp; Carvajal, M. A. 2014. A new species of Acledra s. str. (Hemiptera: Heteroptera: Pentatomidae) from the highlands of Argentina and Bolivia, with a checklist and key to the species of the nominate subgenus. Zootaxa, 3900 (1): 127 - 134." type="journal article" year="2014">Rider &amp; Carvajal, 2014</bibRefCitation>
y
<taxonomicName id="A2C635B4D021A402FCF7FC80B4EEADCA" authority="Berg, 1878" authorityName="Berg" authorityYear="1878" box="[807,1278,889,914]" class="Insecta" family="Pentatomidae" genus="Acledra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="gregalis" subGenus="Acledra">
<emphasis id="57B29225D021A402FCF7FC80B47EADCA" bold="true" box="[807,1134,889,914]" italics="true" pageId="3" pageNumber="386">Acledra (Acledra) gregalis</emphasis>
Berg, 1878
</taxonomicName>
de las cuales se distingue fácilmente por poseer una carena marfileña central a lo largo el escutelo; mientras que en
<emphasis id="57B29225D021A402FF0AFC3AB1FFAD84" bold="true" box="[218,495,963,988]" italics="true" pageId="3" pageNumber="386">
A. (A).
<taxonomicName id="A2C635B4D021A402FEE2FC3AB1FFAD84" box="[306,495,963,988]" class="Insecta" family="Pentatomidae" genus="Acledra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="bichromocornis">bichromocornis</taxonomicName>
</emphasis>
ésta está ausente, y
<emphasis id="57B29225D021A402FD28FC3AB3C5AD84" bold="true" box="[760,981,963,988]" italics="true" pageId="3" pageNumber="386">
en
<taxonomicName id="A2C635B4D021A402FCCCFC3AB3C5AD84" authorityName="Berg" authorityYear="1878" box="[796,981,963,988]" class="Insecta" family="Pentatomidae" genus="Acledra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="gregalis" subGenus="Acledra">A. (A.) gregalis</taxonomicName>
</emphasis>
ésta se restringe a la zona basal.
</emphasis>
</paragraph>
<paragraph id="65794E37D021A402FF3EFC03B452AA61" blockId="3.[182,1444,1018,1081]" pageId="3" pageNumber="386">
<emphasis id="57B29225D021A402FF3EFC03B452AA61" bold="true" pageId="3" pageNumber="386">
Etimología: Dedicamos esta subespecie a Alfredo Lüer, en reconocimiento a sus valiosas recolecciones de heterópteros, incluyendo el
<typeStatus id="BA7DF095D021A402FD35FBD9B358AA61" box="[741,840,1056,1081]" pageId="3" pageNumber="386" type="holotype">holotipo</typeStatus>
de esta subespecie.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

96
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<emphasis id="57B29225D023A400FF40FC5BB39BADE4" bold="true" box="[144,907,930,956]" pageId="1" pageNumber="384">
Complejo de
<taxonomicName id="A2C635B4D023A400FEE6FC5AB39BADE4" authority="(Spinola, 1852)" baseAuthorityName="Spinola" baseAuthorityYear="1852" box="[310,907,930,956]" class="Insecta" family="Pentatomidae" genus="Acledra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="species" species="albocostata" subGenus="Acledra">
<emphasis id="57B29225D023A400FEE6FC5AB2A9ADE4" bold="true" box="[310,697,931,956]" italics="true" pageId="1" pageNumber="384">Acledra (Acledra) albocostata</emphasis>
(Spinola, 1852)
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="2DDC1DBCD023A400FF3EFC31B179AABE" pageId="1" pageNumber="384" type="description">
<paragraph id="65794E37D023A400FF3EFC31B179AABE" blockId="1.[144,1444,930,1254]" pageId="1" pageNumber="384">
<emphasis id="57B29225D023A400FF3EFC31B179AABE" bold="true" pageId="1" pageNumber="384">
Cuando el subgénero
<taxonomicName id="A2C635B4D023A400FE23FC31B2B8ADB9" authority="Fa" authorityName="Fa" box="[499,680,968,993]" class="Insecta" family="Pentatomidae" genus="Neoacledra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="genus">
<emphasis id="57B29225D023A400FE23FC31B290ADB9" bold="true" box="[499,640,968,993]" italics="true" pageId="1" pageNumber="384">Neoacledra</emphasis>
Fa
</taxonomicName>
̇ndez, 2010 fue erigido, Fȧndez &amp;
<bibRefCitation id="015733C6D023A400FBB1FC31B50DADB9" author="Verdejo, L. M." box="[1121,1309,968,993]" pageId="1" pageNumber="384" pagination="77 - 82" refId="ref3422" refString="Fa ˙ ndez, E. I. &amp; Verdejo, L. M. 2010. La singular morfologia de Acledra haematopa (Spinola, 1852) dentro del genero Acledra Signoret, 1864 (Hemiptera: Heteroptera: Pentatomidae), un caso de mimetismo batesiano con descripcion de un nuevo subgenero. Boletin de la Sociedad Entomologica Aragonesa, 46: 77 - 82." type="journal article" year="2010">Verdejo (2010)</bibRefCitation>
mencionan que la falta de datos impedía establecer claramente las relaciones entre otros grupos existentes de especies. Uno de estos grupos correspondería al de
<taxonomicName id="A2C635B4D023A400FCFAFBEBB408AA73" baseAuthorityName="Spinola" baseAuthorityYear="1852" box="[810,1048,1042,1067]" class="Insecta" family="Pentatomidae" genus="Acledra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="species" species="albocostata" subGenus="Acledra">
<emphasis id="57B29225D023A400FCFAFBEBB408AA73" bold="true" box="[810,1048,1042,1067]" italics="true" pageId="1" pageNumber="384">A. (A.) albocostata</emphasis>
</taxonomicName>
, la cual estaría estrechamente relacionada con
<taxonomicName id="A2C635B4D023A400FE8AFBC1B2C4AA09" authority="Breddin, 1914" authorityName="Breddin" authorityYear="1914" box="[346,724,1080,1105]" class="Insecta" family="Pentatomidae" genus="Acledra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="species" species="carinata" subGenus="Acledra">
<emphasis id="57B29225D023A400FE8AFBC1B20BAA09" bold="true" box="[346,539,1080,1105]" italics="true" pageId="1" pageNumber="384">A. (A.) carinata</emphasis>
Breddin, 1914
</taxonomicName>
, especie de la que sólo se conoce un ejemplar hembra, lo que por años ha dificultado dilucidar si ambos taxones son sinónimos o bien forman realmente un grupo de especies cercanas. El estudio de nuevos ejemplares, incluyendo machos de
<taxonomicName id="A2C635B4D023A400FBBCFB7BB53FAAC3" authorityName="Breddin" authorityYear="1914" box="[1132,1327,1154,1179]" class="Insecta" family="Pentatomidae" genus="Acledra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="species" species="carinata" subGenus="Acledra">
<emphasis id="57B29225D023A400FBBCFB7BB53FAAC3" bold="true" box="[1132,1327,1154,1179]" italics="true" pageId="1" pageNumber="384">A. (A.) carinata</emphasis>
</taxonomicName>
y nuevas localidades, permite reconocer las características generales del complejo que queda definido de la siguiente manera:
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="2DDC1DBCD023A400FF66FAF8B4F3AB8B" pageId="1" pageNumber="384" type="diagnosis">
<paragraph id="65794E37D023A400FF66FAF8B13BAB42" blockId="1.[182,1443,1281,1492]" box="[182,299,1281,1306]" pageId="1" pageNumber="384">
<emphasis id="57B29225D023A400FF66FAF8B13BAB42" bold="true" box="[182,299,1281,1306]" pageId="1" pageNumber="384">Diagnosis</emphasis>
</paragraph>
<paragraph id="65794E37D023A400FF3EFADCB4F3AB8B" blockId="1.[182,1443,1281,1492]" pageId="1" pageNumber="384">
<emphasis id="57B29225D023A400FF3EFADCB4F3AB8B" bold="true" pageId="1" pageNumber="384">
Cuerpo elongado. Cabeza más larga que ancha, paraclípeos no contiguos, sobrepasando levemente el anteclípeo. Pronoto de lados cóncavos, sin línea media transversal, escutelo con forma de triángulo isósceles, con una carena media marfileña y una en cada lado de grosor y largo variable seġn la especie. Vena radial del
<emphasis id="57B29225D023A400FE32FA6CB225ABF6" bold="true" box="[482,565,1429,1454]" italics="true" pageId="1" pageNumber="384">corium</emphasis>
en forma de carena marfileña, membrana ahumada con ṁltiples venas transversales, conexiva aserrada, espiráculos pardo claro o concoloros con el abdomen.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>