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<document ID-DOI="http://dx.doi.org/10.3897/BDJ.12.e118110" ID-Pensoft-Pub="1314-2828-12-e118110" ID-Pensoft-UUID="57B0CE31B4055266A115FC1275D70C79" ModsDocID="1314-2828-12-e118110" checkinTime="1712322014339" checkinUser="pensoft" docAuthor="Xian, Chunlan, Leong, Chi Man, Luo, Jiuyang, Jia, Fenglong, Han, Hongxiang &amp; Xie, Qiang" docDate="2024" docId="CB732E05E055555DA8042AAF14CA49D8" docLanguage="en" docName="BiodivDatJour 12: e118110" docOrigin="Biodiversity Data Journal 12" docPubDate="2024-04-05" docSource="http://dx.doi.org/10.3897/BDJ.12.e118110" docTitle="Tetramorium tsushimae Emery 1925" docType="treatment" docVersion="1" id="57B0CE31B4055266A115FC1275D70C79" lastPageNumber="118110" masterDocId="57B0CE31B4055266A115FC1275D70C79" masterDocTitle="Diversity pattern of insects from Macao based on an updated species checklist after 25 years" masterLastPageNumber="118110" masterPageNumber="118110" pageNumber="118110" updateTime="1712322014339" updateUser="pensoft">
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<mods:namePart id="A90348FF418BC216E77D4DD17C372B59">Leong, Chi Man</mods:namePart>
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<mods:date id="B0E0D274992079A5E53C93BEC7B21460">2024</mods:date>
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<taxonomicName id="88808EFBA7D1F2C437CFF30CFE39A38C" ID-CoL="55PV4" LSID="CB732E05-E055-555D-A804-2AAF14CA49D8" authority="Emery, 1925" authorityName="Emery" authorityYear="1925" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tetramorium tsushimae" order="Hymenoptera" pageId="0" pageNumber="118110" phylum="Arthropoda" rank="species" species="tsushimae">Tetramorium tsushimae Emery, 1925</taxonomicName>
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<paragraph id="C87D14620B724891691D1FE7F6B1E019" pageId="0" pageNumber="118110">Notes</paragraph>
<paragraph id="C0F735609CAD59F4EFF65A63D746F8CC" pageId="0" pageNumber="118110">
<bibRefCitation id="53782AA735809A1E1684A136DB634E78" author="Grimaldi, D." journalOrPublisher="Cambridge University Press, New York" pageId="0" pageNumber="118110" refId="B10982430" refString="Guenard B S Ant diversity, from local to global scales: Effects of environmental conditions, community structure and biological invasions https://www.proquest.com/dissertations-theses/ant-diversity-local-global-scales-effects/docview/1035140127/se-2 2023-12-19T00:00:00+02:00" year="2005">Guenard (2011)</bibRefCitation>
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<mods:title id="887D7A314299A765C11BDF0206EFD659">Species plantarum: exhibentes plantas rite cognitas, ad genera relatas, cum differentiis specificis, nominibus trivialibus, synonymis selectis, locis natalibus, secundum systema sexuale digestas</mods:title>
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<taxonomicName id="72076EEA546CDB7639D9D54633E3D9AA" authorityName="Linnaeus" authorityYear="1753" class="Magnoliopsida" family="Resedaceae" genus="Reseda" kingdom="Plantae" order="Capparales" phylum="Tracheophyta" rank="species" species="glauca" status="spec. nov.">Reseda glauca</taxonomicName>
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<paragraph id="DA2FACDFC90DB153739410B084FF95E2" pageNumber="449">3. Reseda foliis linearibus basi dentatis, floribus tetragynis.</paragraph>
<paragraph id="D21BE7738B169CA1D48A723AD25A0B97" pageNumber="449">
Reseda linariae foliis.
<emphasis id="7B0B7E5F74537B371CB7FAB9D18CC541" italics="true">Bauh. pin. 100. prodr. 41.</emphasis>
<emphasis id="F427C4AC1A03E1DD7710A14900F97750" italics="true">Burs. IV.88.</emphasis>
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Reseda, linariae folio, glaucum pyrenaicum, flore stamineo.
<emphasis id="8C148B1802285DAE23EFB1D9C6AEE46A" italics="true">Moris. hist. 3. p.601. s.15. t.1. f.4.</emphasis>
</paragraph>
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Reseda, linariae folio, pyrenaica, glauco folio.
<emphasis id="3B3F447F70BC2E1D0AC112D1BA667C19" italics="true">Pluk. alm. 317. t.107. f.2.</emphasis>
<emphasis id="BDB916EFFD25B8D972B5C804147C3C25" italics="true">Raj. suppl. 511.</emphasis>
</paragraph>
<paragraph id="13122B985A1561E06710D2F29E9168CF" pageNumber="449">
Luteola pumila pyrenaica, linariae folio.
<emphasis id="1C48A45112A61C98CBF81870B49CB09F" italics="true">Tournef. inst. 424.</emphasis>
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<paragraph id="DDB3D8A8A410266E117BE65D7B3F0E88" pageNumber="449">
<emphasis id="2818405C877CFECE9B2A7284841B92F1" italics="true">Habitat in</emphasis>
Pyrenaeis
<emphasis id="E4853518556B694A754E1052D07FB01D" italics="true">.</emphasis>
</paragraph>
</subSubSection>
</treatment>
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Genus
<taxonomicName LSID="cb73d722-d695-5abd-9cae-dc6c69062b3a" authority="Blanford, 1863" authorityName="Blanford" authorityYear="1863" class="Gastropoda" family="Camaenidae" genus="Ganesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ganesella" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="genus">Ganesella Blanford, 1863</taxonomicName>
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<taxonomicName class="Gastropoda" family="Camaenidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix (Ganesella)" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="subGenus" subGenus="Ganesella">Helix (Ganesella)</taxonomicName>
W.T. Blanford, 1863: 86.
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<taxonomicName class="Gastropoda" family="Camaenidae" genus="Trochomorphoides" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Trochomorphoides" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="genus">Trochomorphoides</taxonomicName>
Nevill, 1878: 80. Type species:
<taxonomicName class="Gastropoda" family="Camaenidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix acris" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="acris">Helix acris</taxonomicName>
Benson, 1859, by original designation.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<taxonomicName class="Gastropoda" family="Camaenidae" genus="Darwininitium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Darwininitium" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="genus">Darwininitium</taxonomicName>
Budha &amp; Mordan in
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.175.2755" author="Budha, PB" journalOrPublisher="ZooKeys" pageId="0" pageNumber="51" pagination="19 - 26" refId="B12" refString="Budha, PB, Mordan, PB, Naggs, F, Backeljau, T, 2012. Darwininitium - a new fully pseudosigmurethrous orthurethran genus from Nepal (Gastropoda: Pulmonata: Cerastidae). . ZooKeys 175: 19 - 26" title="Darwininitium - a new fully pseudosigmurethrous orthurethran genus from Nepal (Gastropoda: Pulmonata: Cerastidae)." url="https://doi.org/10.3897/zookeys.175.2755" volume="175" year="2012">Budha et al. 2012</bibRefCitation>
: 21. Type species:
<taxonomicName class="Gastropoda" family="Camaenidae" genus="Darwininitium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Darwininitium shiwalikianum" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="shiwalikianum">Darwininitium shiwalikianum</taxonomicName>
Budha &amp; Mordan in
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.175.2755" author="Budha, PB" journalOrPublisher="ZooKeys" pageId="0" pageNumber="51" pagination="19 - 26" refId="B12" refString="Budha, PB, Mordan, PB, Naggs, F, Backeljau, T, 2012. Darwininitium - a new fully pseudosigmurethrous orthurethran genus from Nepal (Gastropoda: Pulmonata: Cerastidae). . ZooKeys 175: 19 - 26" title="Darwininitium - a new fully pseudosigmurethrous orthurethran genus from Nepal (Gastropoda: Pulmonata: Cerastidae)." url="https://doi.org/10.3897/zookeys.175.2755" volume="175" year="2012">Budha et al. 2012</bibRefCitation>
, by original designation. New synonym.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="type species">
<paragraph pageId="0" pageNumber="51">Type species.</paragraph>
<paragraph pageId="0" pageNumber="51">
<taxonomicName class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix capitium" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="capitium">
<emphasis italics="true" pageId="0" pageNumber="51">Helix capitium</emphasis>
</taxonomicName>
Benson, 1848 by subsequent designation (
<bibRefCitation author="Pilsbry, HA" journalOrPublisher="The Academy of Natural Sciences of Philadelphia, PA" pageId="0" pageNumber="51" refId="B48" refString="Pilsbry, HA, 1895. Manual of Conchology, Structural and Systematic, with Illustrations of the Species, Series 2 volume 9. . The Academy of Natural Sciences of Philadelphia, PA" title="Manual of Conchology, Structural and Systematic, with Illustrations of the Species, Series 2 volume 9." year="1895">Pilsbry 1895</bibRefCitation>
: 168).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="description">
<paragraph pageId="0" pageNumber="51">Description.</paragraph>
<paragraph pageId="0" pageNumber="51">Shell more or less trochoid, moderately thin to solid, with 4-6 convex whorls. Last whorl rounded to angular, with or without a peripheral keel, a little descending in front. Colour light monochrome or with a few dark bands and/or spots and streaks. Embryonic shell smooth. Post apical whorls with irregular, thin, radial ridges and spiral lines (smooth below peripheral angle or keel). Aperture widely ovate, moderately oblique with variously reflected margins. Umbilicus narrow, but open, rarely closed. Shell height ranged from 4.5 to 25.0 mm and shell width ranged from 5.5 to 27.0 mm.</paragraph>
<paragraph pageId="0" pageNumber="51">Genitalia typical of camaenids, without accessory organs on penis or vagina. Penis short to long, with small to large penial verge, but no penial appendix; epiphallus and flagellum short.</paragraph>
<paragraph pageId="0" pageNumber="51">Radular teeth triangular to spatulate, central tooth unicuspid, lateral and marginal teeth tricuspid.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="remarks">
<paragraph pageId="0" pageNumber="51">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="51">
The genus
<taxonomicName class="Gastropoda" family="Camaenidae" genus="Ganesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ganesella" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="51">Ganesella</emphasis>
</taxonomicName>
s.s. differs from
<taxonomicName class="Gastropoda" family="Camaenidae" genus="Satsuma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Satsuma" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="51">Satsuma</emphasis>
</taxonomicName>
s.s by having a penial verge, but no penial appendix. In contrast,
<taxonomicName class="Gastropoda" family="Camaenidae" genus="Satsuma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Satsuma" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="51">Satsuma</emphasis>
</taxonomicName>
has a short to long penial appendix on the distal part of the penis, but lacks a penial verge. In addition,
<taxonomicName class="Gastropoda" family="Camaenidae" genus="Satsuma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Satsuma" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="51">Satsuma</emphasis>
</taxonomicName>
occurs from Japan to Taiwan and southern China, while
<taxonomicName class="Gastropoda" family="Camaenidae" genus="Ganesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ganesella" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="51">Ganesella</emphasis>
</taxonomicName>
occurs from South to Southeast Asia, including Japan (
<bibRefCitation author="Azuma, M" journalOrPublisher="Hoikusha, Japan" pageId="0" pageNumber="51" refId="B5" refString="Azuma, M, 1995. Colored illustrations of the land snails of Japan. . Hoikusha, Japan" title="Colored illustrations of the land snails of Japan." year="1995">Azuma 1995</bibRefCitation>
,
<bibRefCitation pageId="0" pageNumber="51" refId="B54">Schileyko 2003</bibRefCitation>
,
<bibRefCitation DOI="https://doi.org/10.1111/j.1096-3642.2008.00415.x" author="Wu, S-H" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="51" pagination="437 - 493" refId="B69" refString="Wu, S-H, Hwang, C-C, Lin, Y-S, 2008. Systematic revision of the arboreal snail Satsumaalbida species complex (Mollusca: Camaenidae) with description of 14 new species from Taiwan. . Zoological Journal of the Linnean Society 154: 437 - 493" title="Systematic revision of the arboreal snail Satsumaalbida species complex (Mollusca: Camaenidae) with description of 14 new species from Taiwan." url="https://doi.org/10.1111/j.1096-3642.2008.00415.x" volume="154" year="2008">Wu et al. 2008</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="51">
<taxonomicName class="Gastropoda" family="Camaenidae" genus="Darwininitium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Darwininitium shiwalikianum" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="shiwalikianum">
<emphasis italics="true" pageId="0" pageNumber="51">Darwininitium shiwalikianum</emphasis>
</taxonomicName>
Budha &amp; Mordan, 2012, the type species of the monotypic genus
<taxonomicName class="Gastropoda" family="Camaenidae" genus="Darwininitium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Darwininitium" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="51">Darwininitium</emphasis>
</taxonomicName>
Budha &amp; Mordan, 2012, appears conchologically identical to
<taxonomicName class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix capitium" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="capitium">
<emphasis italics="true" pageId="0" pageNumber="51">Helix capitium</emphasis>
</taxonomicName>
, the type species of
<taxonomicName class="Gastropoda" family="Camaenidae" genus="Ganesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ganesella" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="51">Ganesella</emphasis>
</taxonomicName>
(see also
<bibRefCitation pageId="0" pageNumber="51" refId="B13">Budha et al. 2016</bibRefCitation>
). Therefore, the genus
<taxonomicName class="Gastropoda" family="Camaenidae" genus="Darwininitium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Darwininitium" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="51">Darwininitium</emphasis>
</taxonomicName>
, which was originally described from central Nepal, is here tentatively regarded as a junior subjective synonym of the camaenid genus
<taxonomicName class="Gastropoda" family="Camaenidae" genus="Ganesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ganesella" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="51">Ganesella</emphasis>
</taxonomicName>
. If this is confirmed, then
<taxonomicName class="Gastropoda" family="Camaenidae" genus="Darwininitium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Darwininitium" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="51">Darwininitium</emphasis>
</taxonomicName>
was erroneously assigned to the family
<taxonomicName family="Cerastidae" lsidName="" pageId="0" pageNumber="51" rank="family">Cerastidae</taxonomicName>
and as such, its pallial system with a long kidney and s-shape ureter would point to a sigmurethrous condition and not to a case of pseudosigmurethry within the
<taxonomicName lsidName="Orthurethra" pageId="0" pageNumber="51" rank="suborder" suborder="Orthurethra">Orthurethra</taxonomicName>
(sensu
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.2888" author="Solem, A" journalOrPublisher="Fieldiana: Zoology" pageId="0" pageNumber="51" pagination="1 - 238" refId="B57" refString="Solem, A, 1959. Systematics of the land and fresh-water Mollusca of the New Hebrides. . Fieldiana: Zoology 43: 1 - 238" title="Systematics of the land and fresh-water Mollusca of the New Hebrides." url="https://doi.org/10.5962/bhl.title.2888" volume="43" year="1959">Solem 1959</bibRefCitation>
and see also
<bibRefCitation pageId="0" pageNumber="51" refId="B13">Budha et al. 2016</bibRefCitation>
). These taxonomic conclusions appear to be supported by DNA sequence data (
<bibRefCitation pageId="0" pageNumber="51" refId="B13">Budha et al. 2016</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>