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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1108.85156" ID-Pensoft-Pub="1313-2970-1108-89" ID-Pensoft-UUID="4C588406675B5E2F8199E4C9DF13E485" ID-ZooBank="DB9FDD0F5FDF4B028E0B003EAE84EC91" ModsDocID="1313-2970-1108-89" checkinTime="1656059516731" checkinUser="pensoft" docAuthor="Zhao, Yi, Guo, Wan-Ru, Golovatch, Sergei I. &amp; Liu, Wei-Xin" docDate="2022" docId="CB523FEB7A2554609F1B4086B3618D6B" docLanguage="en" docName="ZooKeys 1108: 89-118" docOrigin="ZooKeys 1108" docPubDate="2022-06-23" docSource="http://dx.doi.org/10.3897/zookeys.1108.85156" docTitle="Glyphiulus conuliformis Zhao &amp; Liu 2022, sp. nov." docType="treatment" docUuid="7FCC4E95-5E67-450E-8003-D5F41C71E5B1" docUuidSource="ZooBank" docVersion="1" id="4C588406675B5E2F8199E4C9DF13E485" lastPageNumber="89" masterDocId="4C588406675B5E2F8199E4C9DF13E485" masterDocTitle="Revision of the javanicus species group of the millipede genus Glyphiulus Gervais, 1847, with descriptions of five new species from China (Diplopoda, Spirostreptida, Cambalopsidae)" masterLastPageNumber="118" masterPageNumber="89" pageNumber="89" updateTime="1656059516731" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the javanicus species group of the millipede genus Glyphiulus Gervais, 1847, with descriptions of five new species from China (Diplopoda, Spirostreptida, Cambalopsidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhao, Yi</mods:namePart>
<mods:affiliation>College of Plant Protection, South China Agricultural University, Guangzhou 510642, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Guo, Wan-Ru</mods:namePart>
<mods:affiliation>College of Plant Protection, South China Agricultural University, Guangzhou 510642, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Golovatch, Sergei I.</mods:namePart>
<mods:affiliation>Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071, Russia</mods:affiliation>
<mods:nameIdentifier type="email">sgolovatch@yandex.ru</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liu, Wei-Xin</mods:namePart>
<mods:affiliation>College of Plant Protection, South China Agricultural University, Guangzhou 510642, China</mods:affiliation>
<mods:nameIdentifier type="email">da2000wei@163.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-06-23</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1108</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>89</mods:start>
<mods:end>118</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1108.85156</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1108.85156</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1108-89</mods:identifier>
<mods:identifier type="ZooBank">DB9FDD0F5FDF4B028E0B003EAE84EC91</mods:identifier>
<mods:identifier type="Pensoft-UUID">4C588406675B5E2F8199E4C9DF13E485</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:zoobank.org:act:7FCC4E95-5E67-450E-8003-D5F41C71E5B1" httpUri="http://treatment.plazi.org/id/CB523FEB7A2554609F1B4086B3618D6B" lastPageNumber="89" pageId="0" pageNumber="89">
<subSubSection pageId="0" pageNumber="89" type="nomenclature">
<paragraph pageId="0" pageNumber="89">
<taxonomicName LSID="http://zoobank.org/7FCC4E95-5E67-450E-8003-D5F41C71E5B1" authority="Zhao &amp; Liu" authorityName="Zhao &amp; Liu" authorityYear="2022" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphiulus conuliformis" order="Spirostreptida" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="conuliformis" status="sp. nov.">Glyphiulus conuliformis Zhao &amp; Liu</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="89">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="89" type="description">
<paragraph pageId="0" pageNumber="89">
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Pictures of live animals A G. sinuatoprocessus Zhao &amp; Liu, sp. nov. from Cave Niubi Yan B G. scutatus Zhao &amp; Liu, sp. nov. from Cave Bianfu Dong C G. portaliformis Zhao &amp; Liu, sp. nov. from Cave Baiyan Dong D G. xiniudong Zhao &amp; Liu, sp. nov. from Cave Xiniu Dong E G. conuliformis Zhao &amp; Liu, sp. nov. from Cave Yanzi Dong." figureDoi="10.3897/zookeys.1108.85156.figure5" httpUri="https://binary.pensoft.net/fig/705255" pageId="0" pageNumber="89">Figs 5E</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Glyphiulus conuliformis Zhao &amp; Liu, sp. nov., ♂ paratype A, B anterior body rings, dorsal and lateral views C, D mid-body rings, dorsal and lateral views, respectively E, F posterior body rings, dorsal and ventral views, respectively. Scale bars: 1 mm. Abbreviation: ot: ozoporiferous tubercle." figureDoi="10.3897/zookeys.1108.85156.figure8" httpUri="https://binary.pensoft.net/fig/705258" pageId="0" pageNumber="89">, 8</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Glyphiulus conuliformis Zhao &amp; Liu, sp. nov., ♂ paratype A head and collum B right antenna, oral view C gnathochilarium D mid-leg and claw E leg 3, caudal view F leg 1, frontal view G ♀ paratype, vulvae H penes I anterior gonopods, caudal view J, K posterior gonopods, frontal and caudal views, respectively. Scale bars: 0.5 mm (A-E), 0.2 mm (F-K)." figureDoi="10.3897/zookeys.1108.85156.figure9" httpUri="https://binary.pensoft.net/fig/705259" pageId="0" pageNumber="89">, 9</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="89" type="type material">
<paragraph pageId="0" pageNumber="89">Type material.</paragraph>
<paragraph pageId="0" pageNumber="89">
<emphasis bold="true" italics="true" pageId="0" pageNumber="89">Holotype</emphasis>
♂ (SCAU G24), China, Guangdong Province, Yangjiang City, Yangchun, Cave Yanzi Dong,
<geoCoordinate degrees="22" direction="north" minutes="5" orientation="latitude" precision="925" value="22.083334">22°5'N</geoCoordinate>
,
<geoCoordinate degrees="111" direction="east" minutes="36" orientation="longitude" precision="15" seconds="50" value="111.613884">111°36'50&quot;E</geoCoordinate>
, 400 m alt., 2016-X-29, leg. Tian Mingyi, Chen Mengzhen &amp; Wang Dianmei.
<emphasis bold="true" italics="true" pageId="0" pageNumber="89">Paratypes</emphasis>
: 2 ♂, 26 ♀ (SCAU G24), same data as the holotype.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="89" type="etymology">
<paragraph pageId="0" pageNumber="89">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="89">To emphasise the metatergal anterior tubercles being very sharp and coniform.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="89" type="diagnosis">
<paragraph pageId="0" pageNumber="89">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="89">
Differs from congeners of the
<taxonomicName baseAuthorityName="Pocock" baseAuthorityYear="1895" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphiulus formosus" order="Spirostreptida" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="formosus">
<emphasis italics="true" pageId="0" pageNumber="89">Glyphiulus formosus</emphasis>
</taxonomicName>
-group by the conical shape of the anterior tubercles of metaterga and by the first segment of the telopodite being significantly enlarged in ♂ legs 1, combined with the anterior gonopod process being slender, finger-shaped and curved inwards distally. Based on molecular evidence,
<taxonomicName genus="G." lsidName="G. conuliformis" pageId="0" pageNumber="89" rank="species" species="conuliformis">
<emphasis italics="true" pageId="0" pageNumber="89">G. conuliformis</emphasis>
</taxonomicName>
Zhao &amp; Liu, sp. nov. differs from all other
<taxonomicName authorityName="Gervais" authorityYear="1847" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphiulus" order="Spirostreptida" pageId="0" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="89">Glyphiulus</emphasis>
</taxonomicName>
species analysed in a&gt; 17.0% uncorrected p-distance of the COI barcoding gene.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="89" type="description">
<paragraph pageId="0" pageNumber="89">Description.</paragraph>
<paragraph pageId="0" pageNumber="89">
Length ca. 32.5-37.0 (♂) or 47.0-58.0 mm (♀), mid-body rings round in cross-section, their width and height similar, 1.7-2.0 mm (♂) or 2.2-3.0 mm (♀). Body with 48-53 (♂) or 57-67 (♀) podous + 2-4 apodous rings + telson. Colouration dark brownish, head and legs yellowish (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Pictures of live animals A G. sinuatoprocessus Zhao &amp; Liu, sp. nov. from Cave Niubi Yan B G. scutatus Zhao &amp; Liu, sp. nov. from Cave Bianfu Dong C G. portaliformis Zhao &amp; Liu, sp. nov. from Cave Baiyan Dong D G. xiniudong Zhao &amp; Liu, sp. nov. from Cave Xiniu Dong E G. conuliformis Zhao &amp; Liu, sp. nov. from Cave Yanzi Dong." figureDoi="10.3897/zookeys.1108.85156.figure5" httpUri="https://binary.pensoft.net/fig/705255" pageId="0" pageNumber="89">5E</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="89">
<emphasis bold="true" italics="true" pageId="0" pageNumber="89">Head</emphasis>
surface smooth. Labrum with four teeth anteromedially (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Glyphiulus conuliformis Zhao &amp; Liu, sp. nov., ♂ paratype A head and collum B right antenna, oral view C gnathochilarium D mid-leg and claw E leg 3, caudal view F leg 1, frontal view G ♀ paratype, vulvae H penes I anterior gonopods, caudal view J, K posterior gonopods, frontal and caudal views, respectively. Scale bars: 0.5 mm (A-E), 0.2 mm (F-K)." figureDoi="10.3897/zookeys.1108.85156.figure9" httpUri="https://binary.pensoft.net/fig/705259" pageId="0" pageNumber="89">9C</figureCitation>
). Ocellaria blackish, with 12-22 ommatidia arranged in 2-3 irregular linear rows (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Glyphiulus conuliformis Zhao &amp; Liu, sp. nov., ♂ paratype A head and collum B right antenna, oral view C gnathochilarium D mid-leg and claw E leg 3, caudal view F leg 1, frontal view G ♀ paratype, vulvae H penes I anterior gonopods, caudal view J, K posterior gonopods, frontal and caudal views, respectively. Scale bars: 0.5 mm (A-E), 0.2 mm (F-K)." figureDoi="10.3897/zookeys.1108.85156.figure9" httpUri="https://binary.pensoft.net/fig/705259" pageId="0" pageNumber="89">9A</figureCitation>
). Antennae slender, slightly clavate, reaching back to middle of rings 2 and 3; in length, antennomeres 5&gt; 3&gt; 2 ≈ 4&gt; 6&gt; 1&gt; 7. Antennomeres 5-7 each with a distodorsal field or corolla of bacilliform sensilla (sensory bacilli). Antennomere 7 with four sensory cones (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Glyphiulus conuliformis Zhao &amp; Liu, sp. nov., ♂ paratype A head and collum B right antenna, oral view C gnathochilarium D mid-leg and claw E leg 3, caudal view F leg 1, frontal view G ♀ paratype, vulvae H penes I anterior gonopods, caudal view J, K posterior gonopods, frontal and caudal views, respectively. Scale bars: 0.5 mm (A-E), 0.2 mm (F-K)." figureDoi="10.3897/zookeys.1108.85156.figure9" httpUri="https://binary.pensoft.net/fig/705259" pageId="0" pageNumber="89">9B</figureCitation>
). Gnathochilarium with a separate promentum, polytrichous on promentum and mentum, lamellae linguales each with 6-7 setae (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Glyphiulus conuliformis Zhao &amp; Liu, sp. nov., ♂ paratype A head and collum B right antenna, oral view C gnathochilarium D mid-leg and claw E leg 3, caudal view F leg 1, frontal view G ♀ paratype, vulvae H penes I anterior gonopods, caudal view J, K posterior gonopods, frontal and caudal views, respectively. Scale bars: 0.5 mm (A-E), 0.2 mm (F-K)." figureDoi="10.3897/zookeys.1108.85156.figure9" httpUri="https://binary.pensoft.net/fig/705259" pageId="0" pageNumber="89">9C</figureCitation>
). Mandible not dissected.
</paragraph>
<paragraph pageId="0" pageNumber="89">
<emphasis bold="true" italics="true" pageId="0" pageNumber="89">Collum</emphasis>
: crests complete and evident; carinotaxy formula I-III+P+M (Figs
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Glyphiulus conuliformis Zhao &amp; Liu, sp. nov., ♂ paratype A, B anterior body rings, dorsal and lateral views C, D mid-body rings, dorsal and lateral views, respectively E, F posterior body rings, dorsal and ventral views, respectively. Scale bars: 1 mm. Abbreviation: ot: ozoporiferous tubercle." figureDoi="10.3897/zookeys.1108.85156.figure8" httpUri="https://binary.pensoft.net/fig/705258" pageId="0" pageNumber="89">8A, B</figureCitation>
). Following metaterga strongly crested, anterior tubercles very sharp and coniform; carinotaxy formula 2/2+I/i+3/3 (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Glyphiulus conuliformis Zhao &amp; Liu, sp. nov., ♂ paratype A, B anterior body rings, dorsal and lateral views C, D mid-body rings, dorsal and lateral views, respectively E, F posterior body rings, dorsal and ventral views, respectively. Scale bars: 1 mm. Abbreviation: ot: ozoporiferous tubercle." figureDoi="10.3897/zookeys.1108.85156.figure8" httpUri="https://binary.pensoft.net/fig/705258" pageId="0" pageNumber="89">8</figureCitation>
). Ozoporiferous tubercles very large, coniform (Figs
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Glyphiulus conuliformis Zhao &amp; Liu, sp. nov., ♂ paratype A, B anterior body rings, dorsal and lateral views C, D mid-body rings, dorsal and lateral views, respectively E, F posterior body rings, dorsal and ventral views, respectively. Scale bars: 1 mm. Abbreviation: ot: ozoporiferous tubercle." figureDoi="10.3897/zookeys.1108.85156.figure8" httpUri="https://binary.pensoft.net/fig/705258" pageId="0" pageNumber="89">8C, D</figureCitation>
). Tegument delicately alveolate-areolate, fine longitudinal striations in front of stricture. Rings 2 and 3 with long pleural flaps.
</paragraph>
<caption doi="10.3897/zookeys.1108.85156.figure8" httpUri="https://binary.pensoft.net/fig/705258" pageId="0" pageNumber="89" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="89">
<emphasis bold="true" pageId="0" pageNumber="89">Figure 8.</emphasis>
<taxonomicName authorityName="Zhao &amp; Liu" authorityYear="2022" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphiulus conuliformis" order="Spirostreptida" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="conuliformis">
<emphasis italics="true" pageId="0" pageNumber="89">Glyphiulus conuliformis</emphasis>
</taxonomicName>
Zhao &amp; Liu, sp. nov., ♂ paratype
<emphasis bold="true" pageId="0" pageNumber="89">A, B</emphasis>
anterior body rings, dorsal and lateral views
<emphasis bold="true" pageId="0" pageNumber="89">C, D</emphasis>
mid-body rings, dorsal and lateral views, respectively
<emphasis bold="true" pageId="0" pageNumber="89">E, F</emphasis>
posterior body rings, dorsal and ventral views, respectively. Scale bars: 1 mm. Abbreviation: ot: ozoporiferous tubercle.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="89">
<emphasis bold="true" italics="true" pageId="0" pageNumber="89">Epiproct</emphasis>
simple, caudal edge with a very low central protrusion, dorsally with an obvious sharp tubercle (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Glyphiulus conuliformis Zhao &amp; Liu, sp. nov., ♂ paratype A, B anterior body rings, dorsal and lateral views C, D mid-body rings, dorsal and lateral views, respectively E, F posterior body rings, dorsal and ventral views, respectively. Scale bars: 1 mm. Abbreviation: ot: ozoporiferous tubercle." figureDoi="10.3897/zookeys.1108.85156.figure8" httpUri="https://binary.pensoft.net/fig/705258" pageId="0" pageNumber="89">8E</figureCitation>
). Paraprocts regularly convex, each with several irregular rows of setae (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Glyphiulus conuliformis Zhao &amp; Liu, sp. nov., ♂ paratype A, B anterior body rings, dorsal and lateral views C, D mid-body rings, dorsal and lateral views, respectively E, F posterior body rings, dorsal and ventral views, respectively. Scale bars: 1 mm. Abbreviation: ot: ozoporiferous tubercle." figureDoi="10.3897/zookeys.1108.85156.figure8" httpUri="https://binary.pensoft.net/fig/705258" pageId="0" pageNumber="89">8F</figureCitation>
). Hypoproct transversely bean-shaped, with 3+3 small setae.
</paragraph>
<paragraph pageId="0" pageNumber="89">
<emphasis bold="true" italics="true" pageId="0" pageNumber="89">Legs</emphasis>
slender, about 1.3 times as long as mid-body height; claw with a small accessory spine at base, about 1/6 as long as claw (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Glyphiulus conuliformis Zhao &amp; Liu, sp. nov., ♂ paratype A head and collum B right antenna, oral view C gnathochilarium D mid-leg and claw E leg 3, caudal view F leg 1, frontal view G ♀ paratype, vulvae H penes I anterior gonopods, caudal view J, K posterior gonopods, frontal and caudal views, respectively. Scale bars: 0.5 mm (A-E), 0.2 mm (F-K)." figureDoi="10.3897/zookeys.1108.85156.figure9" httpUri="https://binary.pensoft.net/fig/705259" pageId="0" pageNumber="89">9D</figureCitation>
). ♂ legs 1 very strongly reduced, represented by a sternum showing a pair of small, unfused, paramedian, subunciform prongs directed forward, with about 4-5+4-5 long setae at base; flanked by strongly separated, rudimentary, 2-segmented, asymmetric leg vestiges, first segment significantly enlarged (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Glyphiulus conuliformis Zhao &amp; Liu, sp. nov., ♂ paratype A head and collum B right antenna, oral view C gnathochilarium D mid-leg and claw E leg 3, caudal view F leg 1, frontal view G ♀ paratype, vulvae H penes I anterior gonopods, caudal view J, K posterior gonopods, frontal and caudal views, respectively. Scale bars: 0.5 mm (A-E), 0.2 mm (F-K)." figureDoi="10.3897/zookeys.1108.85156.figure9" httpUri="https://binary.pensoft.net/fig/705259" pageId="0" pageNumber="89">9F</figureCitation>
). ♂ legs 2 slightly hypertrophied, coxae large; penes rather small, much shorter than coxae, oblong-subtrapeziform, each with 1 or 2 strong setae distolaterally (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Glyphiulus conuliformis Zhao &amp; Liu, sp. nov., ♂ paratype A head and collum B right antenna, oral view C gnathochilarium D mid-leg and claw E leg 3, caudal view F leg 1, frontal view G ♀ paratype, vulvae H penes I anterior gonopods, caudal view J, K posterior gonopods, frontal and caudal views, respectively. Scale bars: 0.5 mm (A-E), 0.2 mm (F-K)." figureDoi="10.3897/zookeys.1108.85156.figure9" httpUri="https://binary.pensoft.net/fig/705259" pageId="0" pageNumber="89">9H</figureCitation>
). ♂ legs 3 modified through coxae being especially slender and elongate (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Glyphiulus conuliformis Zhao &amp; Liu, sp. nov., ♂ paratype A head and collum B right antenna, oral view C gnathochilarium D mid-leg and claw E leg 3, caudal view F leg 1, frontal view G ♀ paratype, vulvae H penes I anterior gonopods, caudal view J, K posterior gonopods, frontal and caudal views, respectively. Scale bars: 0.5 mm (A-E), 0.2 mm (F-K)." figureDoi="10.3897/zookeys.1108.85156.figure9" httpUri="https://binary.pensoft.net/fig/705259" pageId="0" pageNumber="89">9E</figureCitation>
). ♂ femora 6 and 7 normal, without modifications.
</paragraph>
<caption doi="10.3897/zookeys.1108.85156.figure9" httpUri="https://binary.pensoft.net/fig/705259" pageId="0" pageNumber="89" start="Figure 9" startId="F9">
<paragraph pageId="0" pageNumber="89">
<emphasis bold="true" pageId="0" pageNumber="89">Figure 9.</emphasis>
<taxonomicName authorityName="Zhao &amp; Liu" authorityYear="2022" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphiulus conuliformis" order="Spirostreptida" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="conuliformis">
<emphasis italics="true" pageId="0" pageNumber="89">Glyphiulus conuliformis</emphasis>
</taxonomicName>
Zhao &amp; Liu, sp. nov., ♂ paratype
<emphasis bold="true" pageId="0" pageNumber="89">A</emphasis>
head and collum
<emphasis bold="true" pageId="0" pageNumber="89">B</emphasis>
right antenna, oral view
<emphasis bold="true" pageId="0" pageNumber="89">C</emphasis>
gnathochilarium
<emphasis bold="true" pageId="0" pageNumber="89">D</emphasis>
mid-leg and claw
<emphasis bold="true" pageId="0" pageNumber="89">E</emphasis>
leg 3, caudal view
<emphasis bold="true" pageId="0" pageNumber="89">F</emphasis>
leg 1, frontal view
<emphasis bold="true" pageId="0" pageNumber="89">G</emphasis>
♀ paratype, vulvae
<emphasis bold="true" pageId="0" pageNumber="89">H</emphasis>
penes
<emphasis bold="true" pageId="0" pageNumber="89">I</emphasis>
anterior gonopods, caudal view
<emphasis bold="true" pageId="0" pageNumber="89">J, K</emphasis>
posterior gonopods, frontal and caudal views, respectively. Scale bars: 0.5 mm (
<emphasis bold="true" pageId="0" pageNumber="89">A-E</emphasis>
), 0.2 mm (
<emphasis bold="true" pageId="0" pageNumber="89">F-K</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="89">
<emphasis bold="true" italics="true" pageId="0" pageNumber="89">Anterior gonopods</emphasis>
(Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Glyphiulus conuliformis Zhao &amp; Liu, sp. nov., ♂ paratype A head and collum B right antenna, oral view C gnathochilarium D mid-leg and claw E leg 3, caudal view F leg 1, frontal view G ♀ paratype, vulvae H penes I anterior gonopods, caudal view J, K posterior gonopods, frontal and caudal views, respectively. Scale bars: 0.5 mm (A-E), 0.2 mm (F-K)." figureDoi="10.3897/zookeys.1108.85156.figure9" httpUri="https://binary.pensoft.net/fig/705259" pageId="0" pageNumber="89">9I</figureCitation>
) with a broad and plate-shaped coxosternum supplied with about 14-16 microsetae near distal margin; apicomesal process of coxite high, slender and digitiform, curved inwards distally. Telopodite very large and stout, coiled, 1-segmented, lateral in position, almost parallel to lateral corner of coxite, with a field of 8-16 microspinules at base and five strong setae apically.
</paragraph>
<paragraph pageId="0" pageNumber="89">
<emphasis bold="true" italics="true" pageId="0" pageNumber="89">Posterior gonopods</emphasis>
(Figs
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Glyphiulus conuliformis Zhao &amp; Liu, sp. nov., ♂ paratype A head and collum B right antenna, oral view C gnathochilarium D mid-leg and claw E leg 3, caudal view F leg 1, frontal view G ♀ paratype, vulvae H penes I anterior gonopods, caudal view J, K posterior gonopods, frontal and caudal views, respectively. Scale bars: 0.5 mm (A-E), 0.2 mm (F-K)." figureDoi="10.3897/zookeys.1108.85156.figure9" httpUri="https://binary.pensoft.net/fig/705259" pageId="0" pageNumber="89">9J, K</figureCitation>
) compact. Coxite subtrapezoid, with a longitudinal field of 22-24 basolateral microspinules in frontal view and a slanted field of 32-36 median microspinules in caudal view; apical and mesal parts of coxite with dense, strong and curved setae. Lamelliform lobe membranous, with a rather short and broad flagellum.
</paragraph>
<paragraph pageId="0" pageNumber="89">
<emphasis bold="true" italics="true" pageId="0" pageNumber="89">Vulvae</emphasis>
very simple, bare, M-shaped (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Glyphiulus conuliformis Zhao &amp; Liu, sp. nov., ♂ paratype A head and collum B right antenna, oral view C gnathochilarium D mid-leg and claw E leg 3, caudal view F leg 1, frontal view G ♀ paratype, vulvae H penes I anterior gonopods, caudal view J, K posterior gonopods, frontal and caudal views, respectively. Scale bars: 0.5 mm (A-E), 0.2 mm (F-K)." figureDoi="10.3897/zookeys.1108.85156.figure9" httpUri="https://binary.pensoft.net/fig/705259" pageId="0" pageNumber="89">9G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="89" type="remarks">
<paragraph pageId="0" pageNumber="89">Remark.</paragraph>
<paragraph pageId="0" pageNumber="89">In the absence of direct troglomorphic traits, this species can only be considered as troglophilic at most.</paragraph>
</subSubSection>
</treatment>
</document>

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<paragraph id="4344E72D9D45CA52FF7CF888FE36F8B2" blockId="31.[151,486,1803,1865]" box="[151,469,1803,1829]" pageId="31" pageNumber="32">
<heading id="180C50419D45CA52FF7CF888FE36F8B2" bold="true" box="[151,469,1803,1829]" fontSize="11" level="1" pageId="31" pageNumber="32" reason="1">
<taxonomicName id="84FB9CAE9D45CA52FF7CF888FE36F8B2" ID-CoL="GZYH" authority="Evans, 1955" authorityName=", Reserva Duque, Manaus" authorityYear="1955" box="[151,469,1803,1829]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="focus">
<emphasis id="718F3B3F9D45CA52FF7CF888FE36F8B2" bold="true" box="[151,469,1803,1829]" pageId="31" pageNumber="32">
<emphasis id="718F3B3F9D45CA52FF7CF888FEDBF8B2" bold="true" box="[151,312,1803,1829]" italics="true" pageId="31" pageNumber="32">Artines focus</emphasis>
<bibRefCitation id="276A9ADC9D45CA52FED4F888FE36F8B2" author="Evans, W. H." box="[319,469,1803,1829]" pageId="31" pageNumber="32" refId="ref40325" refString="Evans, W. H. (1955) A Catalogue of the American Hesperiidae indicating the Classification and Nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. British Museum, London, 499 pp." type="book" year="1955">Evans, 1955</bibRefCitation>
</emphasis>
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</paragraph>
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<subSubSection id="0BE1B4A69D45CA52FF7CF8ACFE05F8DE" box="[151,486,1838,1865]" pageId="31" pageNumber="32" type="description">
<paragraph id="4344E72D9D45CA52FF7CF8ACFE05F8DE" blockId="31.[151,486,1803,1865]" box="[151,486,1838,1865]" pageId="31" pageNumber="32">
<figureCitation id="DBC0FBA89D45CA52FF7CF8ACFF0FF8DE" box="[151,236,1839,1865]" captionStart="FIGURES 14–19" captionStartId="10.[151,264,1406,1431]" captionTargetBox="[155,1435,175,1386]" captionTargetId="figure@10.[151,1436,174,1394]" captionTargetPageId="10" captionText="FIGURES 14–19. Species of the “aquilina group” (14–18) and “bipunctata group” (19) in dorsal and ventral views. 14. Artines aquilina, A–B: male, Joinville, Santa Catarina, Brazil (DZ 41.321), C–D: female from same locality of male (DZ 41.220). 15. Artines angelica sp. nov., A–B: holotype male, C–D: allotype female. 16. Artines focus, A–B: male, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.233), C–D: female, Pacaraima, Roraima, Brazil (DZ 41.333). 17. Artines cofus sp. nov., A–B: holotype male, C–D: allotype female. 18. Artines donia sp. nov., A–B: holotype male. 19. Artines bipunctata, A–B: male, Jardim Zoológico, Brasília, Distrito Federal, Brazil (DZ 30.961), C–D: female, Córrego Bananal, Brasília, Distrito Federal, Brazil (DZ 41.224). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241045/files/figure.png" pageId="31" pageNumber="32">Figs 16</figureCitation>
,
<figureCitation id="DBC0FBA89D45CA52FF13F8ACFEF5F8DE" box="[248,278,1839,1865]" captionStart="FIGURES 21–26" captionStartId="22.[151,264,1840,1865]" captionTargetBox="[151,1436,922,1828]" captionTargetId="figure@22.[151,1436,922,1828]" captionText="FIGURES 21–26. Male brands (no scales) on dorsal forewing of the “aquilina group” (21–25) and “bipunctata group” species (26). 21. Artines aquilina, Joinville, Santa catarina, Brazil, (DZ 41.341). 22. Artines angelica sp. nov., paratype, Fazenda Ter- rasse, Santa Luzia, Maranhão, Brazil (DZ 41.273) 23. Artines focus, Ilha de Maracá, Roraima, Brazil (DZ 41.233). 24. Artines cofus sp. nov., paratype, Santa Rosa do Purus, Acre, Brazil (DZ 41.303). 25. Artines donia sp. nov., paratype, Santa Rita do Araguaia, Goiás, Brazil (OM 66.000). 26. Artines bipunctata, Ponte Funda, Vianópilis, Goiás, Brazil (DZ 41.264). Scale bar = 1 mm." httpUri="https://zenodo.org/record/3406270/files/figure.png" pageId="31" pageNumber="32">23</figureCitation>
, 38, 51, 63, 75, 80
</paragraph>
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<paragraph id="4344E72D9D45CA6DFF7CF8F7FE3BFEBB" blockId="31.[151,1437,1906,2023]" lastBlockId="32.[158,1437,152,330]" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
<taxonomicName id="84FB9CAE9D45CA52FF7CF8F7FE4FF81C" ID-CoL="GZYH" authority="Evans, 1955" authorityName=", Reserva Duque, Manaus" authorityYear="1955" box="[151,428,1906,1931]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="focus">
<emphasis id="718F3B3F9D45CA52FF7CF8F7FEFCF81C" box="[151,287,1907,1931]" italics="true" pageId="31" pageNumber="32">Artines focus</emphasis>
<bibRefCitation id="276A9ADC9D45CA52FEC2F8F7FE4FF81C" author="Evans, W. H." box="[297,428,1906,1931]" pageId="31" pageNumber="32" refId="ref40325" refString="Evans, W. H. (1955) A Catalogue of the American Hesperiidae indicating the Classification and Nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. British Museum, London, 499 pp." type="book" year="1955">Evans, 1955</bibRefCitation>
</taxonomicName>
.
<emphasis id="718F3B3F9D45CA52FE52F8F1FD6EF81C" bold="true" box="[441,653,1906,1931]" pageId="31" pageNumber="32">Cat. Amer. Hesp. 4</emphasis>
, p. 107, pl. 60 (male gen.); [holo]
<typeStatus id="9C40598F9D45CA52FC1CF8F1FBC6F81C" box="[1015,1061,1906,1931]" pageId="31" pageNumber="32">type</typeStatus>
male, Roraima, British Guyana, H. Whiteley
<emphasis id="718F3B3F9D45CA52FEC7F811FEA8F83D" box="[300,331,1938,1962]" italics="true" pageId="31" pageNumber="32">leg</emphasis>
.; BM(NH). —Lamas*, 1981.
<emphasis id="718F3B3F9D45CA52FD6DF811FC95F83D" bold="true" box="[646,886,1937,1962]" pageId="31" pageNumber="32">Rev. Soc. Mex. Lep. 6</emphasis>
: 37.—
<bibRefCitation id="276A9ADC9D45CA52FC57F812FBB2F83D" author="Bridges, C. A." box="[956,1105,1937,1962]" pageId="31" pageNumber="32" refId="ref38436" refString="Bridges, C. A. (1983) Lepidoptera: Hesperiidae. Notes on species-group names. Vol. 1 - 5. Author, Urbana, Author, 277 pp. [2 pp., Vol. 1: 129 pp., Vol. 2: 41 pp., Vol. 3: 62 pp., Vol. 4: 30 pp., Vol. 5: 13 pp.]" type="book" year="1983">Bridges, 1983</bibRefCitation>
.
<emphasis id="718F3B3F9D45CA52FBB7F811FB04F83D" bold="true" box="[1116,1255,1937,1962]" pageId="31" pageNumber="32">Lep. Hesp. 1</emphasis>
, p. 44;
<emphasis id="718F3B3F9D45CA52FAD2F812FAA5F83D" bold="true" box="[1337,1350,1937,1962]" pageId="31" pageNumber="32">2</emphasis>
, p. 3.— Bridges, 1988.
<emphasis id="718F3B3F9D45CA52FE8EF833FE0EF85E" bold="true" box="[357,493,1968,1993]" pageId="31" pageNumber="32">Cat. Hesp. 1</emphasis>
, p. 70;
<emphasis id="718F3B3F9D45CA52FDD4F833FDAFF85E" bold="true" box="[575,588,1968,1993]" pageId="31" pageNumber="32">2</emphasis>
, p. 5.—O.
<bibRefCitation id="276A9ADC9D45CA52FD29F833FC0BF85E" author="Mielke, O. H. H. &amp; Casagrande, M. M." box="[706,1000,1968,1993]" pageId="31" pageNumber="32" pagination="175 - 210" refId="ref43250" refString="Mielke, O. H. H. &amp; Casagrande, M. M. (1992) Lepidoptera: Papilionoidea e Hesperioidea coletados na Ilha de Maraca, Alto Alegre, Roraima, parte do Projeto Maraca, com uma lista complementar de Hesperiidae de Roraima. Acta Amazonica, Manaus, 21, 175 - 210. https: // doi. org / 10.1590 / 1809 - 43921991211210" type="journal article" year="1992">Mielke &amp; Casagrande, 1992</bibRefCitation>
.
<emphasis id="718F3B3F9D45CA52FC18F833FB70F85E" bold="true" box="[1011,1171,1968,1993]" pageId="31" pageNumber="32">Acta Amaz. 21</emphasis>
: 199.—Lamas*, 1994,
<emphasis id="718F3B3F9D45CA52FA6CF833FA7FF85F" bold="true" box="[1415,1436,1968,1992]" italics="true" pageId="31" pageNumber="32">in</emphasis>
Foster
<emphasis id="718F3B3F9D45CA52FEE4F853FEDDF870" bold="true" box="[271,318,1998,2023]" italics="true" pageId="31" pageNumber="32">et al</emphasis>
.
<emphasis id="718F3B3F9D45CA52FEA7F84DFDACF870" bold="true" box="[332,591,1998,2023]" pageId="31" pageNumber="32">RAP Working Papers 6</emphasis>
: 175.—
<bibRefCitation id="276A9ADC9D45CA52FD49F84DFCDBF870" author="Bridges, C. A." box="[674,824,1998,2023]" pageId="31" pageNumber="32" refId="ref38783" refString="Bridges, C. A. (1994) Catalogue of the family-group, genus-group and species-group names of the Hesperioidea (Lepidoptera) of the world. Vols. 1 - 13 &amp; Apps. 1 - 3. Author, Urbana, 3. [11] + 598 pp. [11 pp., Vol. 1: 1 pp., Vol. 2: 1 pp., Vol. 3: 1 pp., Vol. 4: 36 pp., Vol. 5: 9 pp., Vol. 6: 20 pp., Vol. 7: 6 pp., Vol. 8: 246 pp., Vol. 9: 76 pp., Vol. 10: 90 pp., Vol. 11: 71 pp., Vol. 12: 28 pp., Vol. 13: 9 pp., App. 1: 1 pp., App. 2: 2 pp., App. 3: 1 pp.]" type="book" year="1994">Bridges, 1994</bibRefCitation>
.
<emphasis id="718F3B3F9D45CA6DFCA8F84DFE84FF26" bold="true" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">Cat. Fam.-Group, Gen.-Group, Sp. Group Nam. Hesp. (Lep.) World 8</emphasis>
, p. 84;
<emphasis id="718F3B3F9D7ACA6DFE5EFF1BFE21FF26" bold="true" box="[437,450,152,177]" pageId="32" pageNumber="33">9</emphasis>
, p. 6.—Robbins
<emphasis id="718F3B3F9D7ACA6DFD84FF1AFD7FFF26" bold="true" box="[623,668,152,177]" italics="true" pageId="32" pageNumber="33">et al</emphasis>
.*, 1996,
<emphasis id="718F3B3F9D7ACA6DFD10FF1AFCF3FF26" bold="true" box="[763,784,153,177]" italics="true" pageId="32" pageNumber="33">in</emphasis>
Wilson &amp; Sandoval.
<taxonomicName id="84FB9CAE9D7ACA6DFC00FF1AFBCFFF26" box="[1003,1068,153,177]" class="Reptilia" genus="Manu" higherTaxonomySource="CoL" kingdom="Animalia" order="Dinosauria" pageId="32" pageNumber="48" phylum="Chordata" rank="genus">
<emphasis id="718F3B3F9D7ACA6DFC00FF1AFBCFFF26" bold="true" box="[1003,1068,153,177]" pageId="32" pageNumber="33">Manu</emphasis>
</taxonomicName>
, p. 248.—O. Mielke &amp; Casagrande, 1998,
<emphasis id="718F3B3F9D7ACA6DFEEDFF34FEF8FF58" bold="true" box="[262,283,183,207]" italics="true" pageId="32" pageNumber="33">in</emphasis>
Milliken &amp; Ratter.
<taxonomicName id="84FB9CAE9D7ACA6DFE0AFF34FDD6FF47" box="[481,565,183,208]" class="Arachnida" family="Theraphosidae" genus="Maraca" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="718F3B3F9D7ACA6DFE0AFF34FDD6FF47" bold="true" box="[481,565,183,208]" pageId="32" pageNumber="33">Maracá</emphasis>
</taxonomicName>
, p. 477.—
<bibRefCitation id="276A9ADC9D7ACA6DFD4BFF34FCA5FF58" author="Garrouste, R." box="[672,838,183,207]" pageId="32" pageNumber="33" pagination="139 - 142" refId="ref40750" refString="Garrouste, R. (2003) Premiere contribution entomologique a la connaissance de Tumuc-Humac-Les lepidopteres du Mitaraka. In: Lacomme, D. &amp; Manil, L. (Eds.), Lepidopteres de Guyane. Lepidopteristes parisiense, Paris, pp. 139 - 142." type="book chapter" year="2003">Garrouste, 2003</bibRefCitation>
,
<emphasis id="718F3B3F9D7ACA6DFCBBFF34FC86FF58" bold="true" box="[848,869,183,207]" italics="true" pageId="32" pageNumber="33">in</emphasis>
Lacomme &amp; Manil.
<emphasis id="718F3B3F9D7ACA6DFBD2FF34FB27FF47" bold="true" box="[1081,1220,183,208]" pageId="32" pageNumber="33">
Lép.
<collectingCountry id="3BECA7BD9D7ACA6DFB9BFF34FB27FF47" box="[1136,1220,183,208]" name="French Guiana" pageId="32" pageNumber="33">Guyane</collectingCountry>
</emphasis>
, p. 141.—O.
<bibRefCitation id="276A9ADC9D7ACA6DFAA5FF34FF1EFF79" author="Mielke, O. H. H." pageId="32" pageNumber="33" pagination="3 - 11" refId="ref43459" refString="Mielke, O. H. H. (2004) Hesperiidae. In: Lamas, G. (Ed.), Checklist: Part 4 A, Hesperioidea-Papilionoidea. In: Heppner, J. (Ed.), Atlas of Neotropical Lepidoptera 5 A. Scientific Publishers, Association for Tropical Lepidoptera, Gainesville, pp. 3 - 11 + 25 - 86." type="book chapter" year="2004">Mielke, 2004</bibRefCitation>
. Hesperioidea, p. 62,
<emphasis id="718F3B3F9D7ACA6DFE0FFF55FE1AFF79" bold="true" box="[484,505,214,238]" italics="true" pageId="32" pageNumber="33">in</emphasis>
Lamas (ed.).
<emphasis id="718F3B3F9D7ACA6DFD65FF56FB75FF79" bold="true" box="[654,1174,213,238]" pageId="32" pageNumber="33">Checklist: Part 4A, Hesperioidea-Papilionoidea</emphasis>
,
<emphasis id="718F3B3F9D7ACA6DFB4DFF55FB58FF79" bold="true" box="[1190,1211,214,238]" italics="true" pageId="32" pageNumber="33">in</emphasis>
Heppner (ed.).
<emphasis id="718F3B3F9D7ACA6DFA8FFF56FE62FE9A" bold="true" pageId="32" pageNumber="33">
<taxonomicName id="84FB9CAE9D7ACA6DFA8FFF56FEBFFE9A" authority="Neotrop. Lep." authorityName="Neotrop. Lep." class="Insecta" genus="Atlas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">Atlas Neotrop. Lep.</taxonomicName>
5A
</emphasis>
.—O.
<bibRefCitation id="276A9ADC9D7ACA6DFE54FF77FDABFE9A" author="Mielke, O. H. H." box="[447,584,244,269]" pageId="32" pageNumber="33" pagination="775 - 1055" refId="ref43529" refString="Mielke, O. H. H. (2005) Catalogue of the American Hesperioidea: Hesperiidae (Lepidoptera) 4. Hesperiinae 1. Adlerodea- Lychnuchus. Sociedade Brasileira de Zoologia, Curitiba. pp. 775 - 1055." type="book chapter" year="2005">Mielke, 2005</bibRefCitation>
.
<emphasis id="718F3B3F9D7ACA6DFDBAFF77FC88FE9A" bold="true" box="[593,875,244,269]" pageId="32" pageNumber="33">Cat. Amer. Hesperioidea 4</emphasis>
, p. 833.—Garrouste, 2007,
<emphasis id="718F3B3F9D7ACA6DFB6FFF76FB7AFE9A" bold="true" box="[1156,1177,245,269]" italics="true" pageId="32" pageNumber="33">in</emphasis>
Lacomme &amp; Manil.
<emphasis id="718F3B3F9D7ACA6DFA87FF77FED3FEBB" bold="true" pageId="32" pageNumber="33">
Lép.
<collectingCountry id="3BECA7BD9D7ACA6DFF2CFE90FEFEFEBB" box="[199,285,275,300]" name="Guyana" pageId="32" pageNumber="33">Guyana</collectingCountry>
2
</emphasis>
,
<emphasis id="718F3B3F9D7ACA6DFED6FE90FE71FEBB" bold="true" box="[317,402,275,300]" pageId="32" pageNumber="33">Rhop. 1</emphasis>
, p. 64.
</paragraph>
<paragraph id="4344E72D9D7ACA6DFF75FEB1FD29FEDD" blockId="32.[158,1437,152,330]" box="[158,714,305,330]" pageId="32" pageNumber="33">
(no genus)
<taxonomicName id="84FB9CAE9D7ACA6DFEE4FEB1FEA7FEDD" ID-CoL="GZYH" authorityName=", Reserva Duque, Manaus" authorityYear="1955" box="[271,324,306,330]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="focus">
<emphasis id="718F3B3F9D7ACA6DFEE4FEB1FEA7FEDD" box="[271,324,306,330]" italics="true" pageId="32" pageNumber="33">focus</emphasis>
</taxonomicName>
;
<bibRefCitation id="276A9ADC9D7ACA6DFEB9FEB2FE3EFEDD" author="Beattie, J. R." box="[338,477,305,330]" pageId="32" pageNumber="33" refId="ref38340" refString="Beattie, J. R. (1976) The Rhopalocera Directory. JB Indexes, Berkeley, xiv + 365 pp." type="book" year="1976">Beattie, 1976</bibRefCitation>
.
<emphasis id="718F3B3F9D7ACA6DFE0DFEB1FD92FEDD" bold="true" box="[486,625,305,330]" pageId="32" pageNumber="33">Rhop. Direct</emphasis>
., p. 147.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D7ACA6DFF7CFEFBFD84FE4D" pageId="32" pageNumber="33" type="description">
<paragraph id="4344E72D9D7ACA6DFF7CFEFBFD84FE4D" blockId="32.[151,1437,376,2011]" pageId="32" pageNumber="33">
<emphasis id="718F3B3F9D7ACA6DFF7CFEFBFE95FE04" bold="true" box="[151,374,376,403]" pageId="32" pageNumber="33">Taxonomic history</emphasis>
.
<taxonomicName id="84FB9CAE9D7ACA6DFE6FFEFAFDF4FE04" authorityName=", Reserva Duque, Manaus" authorityYear="1955" box="[388,535,377,403]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="focus">
<emphasis id="718F3B3F9D7ACA6DFE6FFEFAFDF4FE04" box="[388,535,377,403]" italics="true" pageId="32" pageNumber="33">Artines focus</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="276A9ADC9D7ACA6DFD0CFEFAFC9EFE04" author="Evans, W. H." box="[743,893,376,403]" pageId="32" pageNumber="33" refId="ref40325" refString="Evans, W. H. (1955) A Catalogue of the American Hesperiidae indicating the Classification and Nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. British Museum, London, 499 pp." type="book" year="1955">Evans (1955)</bibRefCitation>
based in five males and five females from
<collectingCountry id="3BECA7BD9D7ACA6DFA88FEFAFF5DFE20" name="Guyana" pageId="32" pageNumber="33">Guyana</collectingCountry>
and
<collectingRegion id="813F29CF9D7ACA6DFF1FFE1FFEB4FE21" box="[244,343,412,438]" country="Brazil" name="Roraima" pageId="32" pageNumber="33">Roraima</collectingRegion>
, Pará (Santarém) and
<collectingRegion id="813F29CF9D7ACA6DFDA5FE1FFD3EFE21" box="[590,733,412,438]" country="Brazil" name="Pernambuco" pageId="32" pageNumber="33">Pernambuco</collectingRegion>
,
<collectingCountry id="3BECA7BD9D7ACA6DFD0CFE1FFCCCFE21" box="[743,815,412,438]" name="Brazil" pageId="32" pageNumber="33">Brazil</collectingCountry>
. The other authors mentioned this species in taxonomic comments, faunistic studies and catalogs.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D7ACA6DFF2CFE60FD8DFD1B" pageId="32" pageNumber="33" type="diagnosis">
<paragraph id="4344E72D9D7ACA6DFF2CFE60FD8DFD1B" blockId="32.[151,1437,376,2011]" pageId="32" pageNumber="33">
<emphasis id="718F3B3F9D7ACA6DFF2CFE60FED9FE69" bold="true" box="[199,314,483,510]" pageId="32" pageNumber="33">Diagnosis</emphasis>
.
<taxonomicName id="84FB9CAE9D7ACA6DFEA1FE67FE3DFE69" authorityName=", Reserva Duque, Manaus" authorityYear="1955" box="[330,478,484,510]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="focus">
<emphasis id="718F3B3F9D7ACA6DFEA1FE67FE3DFE69" box="[330,478,484,510]" italics="true" pageId="32" pageNumber="33">Artines focus</emphasis>
</taxonomicName>
differs from the other species of the “
<taxonomicName id="84FB9CAE9D7ACA6DFC7EFE67FC11FE69" authorityName="Plotz" authorityYear="1883" box="[917,1010,484,510]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="aquilina">
<emphasis id="718F3B3F9D7ACA6DFC7EFE67FC11FE69" box="[917,1010,484,510]" italics="true" pageId="32" pageNumber="33">aquilina</emphasis>
</taxonomicName>
group” in the following combination of characters: ventral forewing without spot in CuA
<subScript id="DF7FE5689D7ACA6DFD0BFD9AFD0BFDB0" attach="left" box="[736,744,537,551]" fontSize="6" pageId="32" pageNumber="33">2</subScript>
–2A; uncus distally wide, with a ventral projection; aedeagus proximal portion narrow; opening of ejaculatory bulb separated from proximal margin of aedeagus by short distance; lamella postvaginalis without proximal plate, distal sclerotized plate without median notch on proximal margin and with distal margin approximately straight.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D7ACA6DFF2CFD16FBFAF977" pageId="32" pageNumber="33" type="description">
<paragraph id="4344E72D9D7ACA6DFF2CFD16FB25FD26" blockId="32.[151,1437,376,2011]" box="[199,1222,661,689]" pageId="32" pageNumber="33">
<emphasis id="718F3B3F9D7ACA6DFF2CFD16FE8EFD27" bold="true" box="[199,365,661,688]" pageId="32" pageNumber="33">Redescription</emphasis>
. Male (
<figureCitation id="DBC0FBA89D7ACA6DFE2FFD15FDF6FD26" box="[452,533,662,689]" captionStart="FIGURES 14–19" captionStartId="10.[151,264,1406,1431]" captionTargetBox="[155,1435,175,1386]" captionTargetId="figure@10.[151,1436,174,1394]" captionTargetPageId="10" captionText="FIGURES 14–19. Species of the “aquilina group” (14–18) and “bipunctata group” (19) in dorsal and ventral views. 14. Artines aquilina, A–B: male, Joinville, Santa Catarina, Brazil (DZ 41.321), C–D: female from same locality of male (DZ 41.220). 15. Artines angelica sp. nov., A–B: holotype male, C–D: allotype female. 16. Artines focus, A–B: male, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.233), C–D: female, Pacaraima, Roraima, Brazil (DZ 41.333). 17. Artines cofus sp. nov., A–B: holotype male, C–D: allotype female. 18. Artines donia sp. nov., A–B: holotype male. 19. Artines bipunctata, A–B: male, Jardim Zoológico, Brasília, Distrito Federal, Brazil (DZ 30.961), C–D: female, Córrego Bananal, Brasília, Distrito Federal, Brazil (DZ 41.224). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241045/files/figure.png" pageId="32" pageNumber="33">Fig. 16</figureCitation>
A–B).
<emphasis id="718F3B3F9D7ACA6DFD8AFD15FD7DFD27" box="[609,670,662,688]" italics="true" pageId="32" pageNumber="33">Head</emphasis>
: antenna with nudum bearing 12 to 13 segments.
</paragraph>
<paragraph id="4344E72D9D7ACA6DFF2CFD38FDF3FD42" blockId="32.[151,1437,376,2011]" box="[199,528,697,725]" pageId="32" pageNumber="33">
<emphasis id="718F3B3F9D7ACA6DFF2CFD38FE62FD43" box="[199,385,698,724]" italics="true" pageId="32" pageNumber="33">Forewing length</emphasis>
:
<quantity id="84034AC89D7ACA6DFE7BFD3AFDEFFD42" box="[400,524,697,725]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="32" pageNumber="33" unit="mm" value="13.5" valueMax="15.0" valueMin="12.0">12–15 mm</quantity>
.
</paragraph>
<paragraph id="4344E72D9D7ACA6DFF2CFD5DFD6DFCA8" blockId="32.[151,1437,376,2011]" pageId="32" pageNumber="33">
<emphasis id="718F3B3F9D7ACA6DFF2CFD5DFE63FD6F" box="[199,384,734,760]" italics="true" pageId="32" pageNumber="33">Dorsal forewing</emphasis>
(
<figureCitation id="DBC0FBA89D7ACA6DFE7BFD5DFE19FD6E" box="[400,506,734,761]" captionStart="FIGURES 14–19" captionStartId="10.[151,264,1406,1431]" captionTargetBox="[155,1435,175,1386]" captionTargetId="figure@10.[151,1436,174,1394]" captionTargetPageId="10" captionText="FIGURES 14–19. Species of the “aquilina group” (14–18) and “bipunctata group” (19) in dorsal and ventral views. 14. Artines aquilina, A–B: male, Joinville, Santa Catarina, Brazil (DZ 41.321), C–D: female from same locality of male (DZ 41.220). 15. Artines angelica sp. nov., A–B: holotype male, C–D: allotype female. 16. Artines focus, A–B: male, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.233), C–D: female, Pacaraima, Roraima, Brazil (DZ 41.333). 17. Artines cofus sp. nov., A–B: holotype male, C–D: allotype female. 18. Artines donia sp. nov., A–B: holotype male. 19. Artines bipunctata, A–B: male, Jardim Zoológico, Brasília, Distrito Federal, Brazil (DZ 30.961), C–D: female, Córrego Bananal, Brasília, Distrito Federal, Brazil (DZ 41.224). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241045/files/figure.png" pageId="32" pageNumber="33">Fig. 16A</figureCitation>
): one to three apical spots, spot in R
<subScript id="DF7FE5689D7ACA6DFC79FD6CFC79FD6A" attach="left" box="[914,922,751,765]" fontSize="6" pageId="32" pageNumber="33">3</subScript>
–R
<subScript id="DF7FE5689D7ACA6DFC50FD6CFC20FD6A" attach="right" box="[955,963,751,765]" fontSize="6" pageId="32" pageNumber="33">4</subScript>
, when present, represented by few scales; discal spots as in
<taxonomicName id="84FB9CAE9D7ACA6DFEB9FC81FE2FFC8C" authorityName="Plotz" authorityYear="1883" box="[338,460,769,796]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="aquilina">
<emphasis id="718F3B3F9D7ACA6DFEB9FC81FE2FFC8C" box="[338,460,769,796]" italics="true" pageId="32" pageNumber="33">A. aquilina</emphasis>
</taxonomicName>
; a white and yellow spot in CuA
<subScript id="DF7FE5689D7ACA6DFCC5FC90FCD5FCB6" attach="left" box="[814,822,787,801]" fontSize="6" pageId="32" pageNumber="33">2</subScript>
–2A; brand with lower projection long, upper projection with narrow base and pointed apex (
<figureCitation id="DBC0FBA89D7ACA6DFDC5FCA6FD61FCA8" box="[558,642,805,831]" captionStart="FIGURES 21–26" captionStartId="22.[151,264,1840,1865]" captionTargetBox="[151,1436,922,1828]" captionTargetId="figure@22.[151,1436,922,1828]" captionText="FIGURES 21–26. Male brands (no scales) on dorsal forewing of the “aquilina group” (21–25) and “bipunctata group” species (26). 21. Artines aquilina, Joinville, Santa catarina, Brazil, (DZ 41.341). 22. Artines angelica sp. nov., paratype, Fazenda Ter- rasse, Santa Luzia, Maranhão, Brazil (DZ 41.273) 23. Artines focus, Ilha de Maracá, Roraima, Brazil (DZ 41.233). 24. Artines cofus sp. nov., paratype, Santa Rosa do Purus, Acre, Brazil (DZ 41.303). 25. Artines donia sp. nov., paratype, Santa Rita do Araguaia, Goiás, Brazil (OM 66.000). 26. Artines bipunctata, Ponte Funda, Vianópilis, Goiás, Brazil (DZ 41.264). Scale bar = 1 mm." httpUri="https://zenodo.org/record/3406270/files/figure.png" pageId="32" pageNumber="33">Fig. 23</figureCitation>
).
</paragraph>
<paragraph id="4344E72D9D7ACA6DFF2CFCCAFD3CFC3C" blockId="32.[151,1437,376,2011]" pageId="32" pageNumber="33">
<emphasis id="718F3B3F9D7ACA6DFF2CFCCAFE64FCF4" box="[199,391,841,867]" italics="true" pageId="32" pageNumber="33">Ventral forewing</emphasis>
(
<figureCitation id="DBC0FBA89D7ACA6DFE7CFCCAFE1CFCF3" box="[407,511,841,868]" captionStart="FIGURES 14–19" captionStartId="10.[151,264,1406,1431]" captionTargetBox="[155,1435,175,1386]" captionTargetId="figure@10.[151,1436,174,1394]" captionTargetPageId="10" captionText="FIGURES 14–19. Species of the “aquilina group” (14–18) and “bipunctata group” (19) in dorsal and ventral views. 14. Artines aquilina, A–B: male, Joinville, Santa Catarina, Brazil (DZ 41.321), C–D: female from same locality of male (DZ 41.220). 15. Artines angelica sp. nov., A–B: holotype male, C–D: allotype female. 16. Artines focus, A–B: male, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.233), C–D: female, Pacaraima, Roraima, Brazil (DZ 41.333). 17. Artines cofus sp. nov., A–B: holotype male, C–D: allotype female. 18. Artines donia sp. nov., A–B: holotype male. 19. Artines bipunctata, A–B: male, Jardim Zoológico, Brasília, Distrito Federal, Brazil (DZ 30.961), C–D: female, Córrego Bananal, Brasília, Distrito Federal, Brazil (DZ 41.224). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241045/files/figure.png" pageId="32" pageNumber="33">Fig. 16B</figureCitation>
): costal area with weak violet aspect; three to four apical spots, spot in R
<subScript id="DF7FE5689D7ACA6DFAD4FCD9FAA4FCFF" attach="left" box="[1343,1351,858,872]" fontSize="6" pageId="32" pageNumber="33">2</subScript>
–R
<subScript id="DF7FE5689D7ACA6DFA83FCD9FA93FCFF" attach="left" box="[1384,1392,858,872]" fontSize="6" pageId="32" pageNumber="33">3</subScript>
not conspicuous and shifted to base; discal spots as on dorsal surface; submarginal band conspicuous and not contiguous in the middle of the cells; without spot in CuA
<subScript id="DF7FE5689D7ACA6DFD74FC21FD44FC27" attach="left" box="[671,679,930,944]" fontSize="6" pageId="32" pageNumber="33">2</subScript>
–2A.
</paragraph>
<paragraph id="4344E72D9D7ACA6DFF2CFC37FBACFC65" blockId="32.[151,1437,376,2011]" pageId="32" pageNumber="33">
<emphasis id="718F3B3F9D7ACA6DFF2CFC37FE72FC59" box="[199,401,948,974]" italics="true" pageId="32" pageNumber="33">Ventral hind wing</emphasis>
(
<figureCitation id="DBC0FBA89D7ACA6DFE4AFC37FDE4FC58" box="[417,519,948,975]" captionStart="FIGURES 14–19" captionStartId="10.[151,264,1406,1431]" captionTargetBox="[155,1435,175,1386]" captionTargetId="figure@10.[151,1436,174,1394]" captionTargetPageId="10" captionText="FIGURES 14–19. Species of the “aquilina group” (14–18) and “bipunctata group” (19) in dorsal and ventral views. 14. Artines aquilina, A–B: male, Joinville, Santa Catarina, Brazil (DZ 41.321), C–D: female from same locality of male (DZ 41.220). 15. Artines angelica sp. nov., A–B: holotype male, C–D: allotype female. 16. Artines focus, A–B: male, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.233), C–D: female, Pacaraima, Roraima, Brazil (DZ 41.333). 17. Artines cofus sp. nov., A–B: holotype male, C–D: allotype female. 18. Artines donia sp. nov., A–B: holotype male. 19. Artines bipunctata, A–B: male, Jardim Zoológico, Brasília, Distrito Federal, Brazil (DZ 30.961), C–D: female, Córrego Bananal, Brasília, Distrito Federal, Brazil (DZ 41.224). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241045/files/figure.png" pageId="32" pageNumber="33">Fig. 16B</figureCitation>
): apical portion of discal cell densely covered by orange scales; central black spot quadrate to rectangular; postdiscal band yellow, limited distally by not wavy margin.
</paragraph>
<paragraph id="4344E72D9D7ACA6DFF2CFC7FFA81FA71" blockId="32.[151,1437,376,2011]" pageId="32" pageNumber="33">
<emphasis id="718F3B3F9D7ACA6DFF2CFC7FFED2FB81" box="[199,305,1020,1046]" italics="true" pageId="32" pageNumber="33">Genitalia</emphasis>
(
<figureCitation id="DBC0FBA89D7ACA6DFED4FC78FE70FB81" box="[319,403,1019,1046]" captionStart="FIGURES 38–39" captionStartId="29.[151,264,1720,1745]" captionTargetBox="[161,1436,431,1681]" captionTargetId="figure@29.[151,1436,450,1693]" captionTargetPageId="29" captionText="FIGURES 38–39. Male genitalia of the “aquilina group” species. 38. Artines focus, Reserva Duque, Manaus, Amazonas, Brazil (DZ 41.192). 39. Artines cofus sp. nov., paratype, Tambopata, Puerto Maldonado Madre de Díos, Peru (MUSM). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241055/files/figure.png" pageId="32" pageNumber="33">Figs 38</figureCitation>
A–M, 51): tegumen longer than wide, distally trapezoid from 1/2 of its extension (
<figureCitation id="DBC0FBA89D7ACA6DFAC2FC78FA6CFB81" box="[1321,1423,1019,1046]" captionStart="FIGURES 38–39" captionStartId="29.[151,264,1720,1745]" captionTargetBox="[161,1436,431,1681]" captionTargetId="figure@29.[151,1436,450,1693]" captionTargetPageId="29" captionText="FIGURES 38–39. Male genitalia of the “aquilina group” species. 38. Artines focus, Reserva Duque, Manaus, Amazonas, Brazil (DZ 41.192). 39. Artines cofus sp. nov., paratype, Tambopata, Puerto Maldonado Madre de Díos, Peru (MUSM). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241055/files/figure.png" pageId="32" pageNumber="33">Fig. 38C</figureCitation>
), in lateral view rectangular, dorsal margin strongly convex (
<figureCitation id="DBC0FBA89D7ACA6DFCCEFB9CFC6FFBAE" box="[805,908,1055,1081]" captionStart="FIGURES 38–39" captionStartId="29.[151,264,1720,1745]" captionTargetBox="[161,1436,431,1681]" captionTargetId="figure@29.[151,1436,450,1693]" captionTargetPageId="29" captionText="FIGURES 38–39. Male genitalia of the “aquilina group” species. 38. Artines focus, Reserva Duque, Manaus, Amazonas, Brazil (DZ 41.192). 39. Artines cofus sp. nov., paratype, Tambopata, Puerto Maldonado Madre de Díos, Peru (MUSM). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241055/files/figure.png" pageId="32" pageNumber="33">Fig. 38A</figureCitation>
). Saccus, in ventral view, narrower on proximal 1/2, anterior portion narrower than tegumen, proximal margin straight or convex (
<figureCitation id="DBC0FBA89D7ACA6DFBAFFBC0FB4DFBCA" box="[1092,1198,1091,1117]" captionStart="FIGURES 38–39" captionStartId="29.[151,264,1720,1745]" captionTargetBox="[161,1436,431,1681]" captionTargetId="figure@29.[151,1436,450,1693]" captionTargetPageId="29" captionText="FIGURES 38–39. Male genitalia of the “aquilina group” species. 38. Artines focus, Reserva Duque, Manaus, Amazonas, Brazil (DZ 41.192). 39. Artines cofus sp. nov., paratype, Tambopata, Puerto Maldonado Madre de Díos, Peru (MUSM). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241055/files/figure.png" pageId="32" pageNumber="33">Fig. 38B</figureCitation>
). Uncus with ventral projection, in dorsal view with base as wide as 1/2 width of tegumen, proximal 1/2 with lateral margin concave, distal 1/2 narrower, tip rounded to slightly bifid (
<figureCitation id="DBC0FBA89D7ACA6DFD23FB09FCD3FB32" box="[712,816,1162,1189]" captionStart="FIGURES 38–39" captionStartId="29.[151,264,1720,1745]" captionTargetBox="[161,1436,431,1681]" captionTargetId="figure@29.[151,1436,450,1693]" captionTargetPageId="29" captionText="FIGURES 38–39. Male genitalia of the “aquilina group” species. 38. Artines focus, Reserva Duque, Manaus, Amazonas, Brazil (DZ 41.192). 39. Artines cofus sp. nov., paratype, Tambopata, Puerto Maldonado Madre de Díos, Peru (MUSM). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241055/files/figure.png" pageId="32" pageNumber="33">Fig. 38C</figureCitation>
), in lateral view, strongly curved ventrally (
<figureCitation id="DBC0FBA89D7ACA6DFACDFB09FA73FB32" box="[1318,1424,1162,1189]" captionStart="FIGURES 38–39" captionStartId="29.[151,264,1720,1745]" captionTargetBox="[161,1436,431,1681]" captionTargetId="figure@29.[151,1436,450,1693]" captionTargetPageId="29" captionText="FIGURES 38–39. Male genitalia of the “aquilina group” species. 38. Artines focus, Reserva Duque, Manaus, Amazonas, Brazil (DZ 41.192). 39. Artines cofus sp. nov., paratype, Tambopata, Puerto Maldonado Madre de Díos, Peru (MUSM). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241055/files/figure.png" pageId="32" pageNumber="33">Fig. 38A</figureCitation>
). Gnathos reaching distal 1/2 of uncus, arms parallel and slightly divergent at apex in ventral view (
<figureCitation id="DBC0FBA89D7ACA6DFAC8FB2DFA6CFB5F" box="[1315,1423,1198,1224]" captionStart="FIGURES 38–39" captionStartId="29.[151,264,1720,1745]" captionTargetBox="[161,1436,431,1681]" captionTargetId="figure@29.[151,1436,450,1693]" captionTargetPageId="29" captionText="FIGURES 38–39. Male genitalia of the “aquilina group” species. 38. Artines focus, Reserva Duque, Manaus, Amazonas, Brazil (DZ 41.192). 39. Artines cofus sp. nov., paratype, Tambopata, Puerto Maldonado Madre de Díos, Peru (MUSM). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241055/files/figure.png" pageId="32" pageNumber="33">Fig. 38D</figureCitation>
), tips slightly curved dorsally, directed to distal 1/2 of uncus (
<figureCitation id="DBC0FBA89D7ACA6DFCBEFB51FC22FB7B" box="[853,961,1234,1260]" captionStart="FIGURES 38–39" captionStartId="29.[151,264,1720,1745]" captionTargetBox="[161,1436,431,1681]" captionTargetId="figure@29.[151,1436,450,1693]" captionTargetPageId="29" captionText="FIGURES 38–39. Male genitalia of the “aquilina group” species. 38. Artines focus, Reserva Duque, Manaus, Amazonas, Brazil (DZ 41.192). 39. Artines cofus sp. nov., paratype, Tambopata, Puerto Maldonado Madre de Díos, Peru (MUSM). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241055/files/figure.png" pageId="32" pageNumber="33">Fig. 38A</figureCitation>
). Valva twice longer than wide; sacculus reduced; costa narrow and triangular, in dorsal view as in
<taxonomicName id="84FB9CAE9D7ACA6DFCF3FB75FC70FA87" authorityName="Plotz" authorityYear="1883" box="[792,915,1270,1296]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="aquilina">
<emphasis id="718F3B3F9D7ACA6DFCF3FB75FC70FA87" box="[792,915,1270,1296]" italics="true" pageId="32" pageNumber="33">A. aquilina</emphasis>
</taxonomicName>
, however less projected inwards and separated from ampulla by short distance; harpe longer than 1/2 valva, uniformly narrower than in the other species of the group; ampulla as wide as 1/2 harpe, dorsally protruding; distal projection of the valva slightly curved inwards and dorsally (
<figureCitation id="DBC0FBA89D7ACA6DFEE8FAE2FEB6FAEC" box="[259,341,1377,1403]" captionStart="FIGURES 38–39" captionStartId="29.[151,264,1720,1745]" captionTargetBox="[161,1436,431,1681]" captionTargetId="figure@29.[151,1436,450,1693]" captionTargetPageId="29" captionText="FIGURES 38–39. Male genitalia of the “aquilina group” species. 38. Artines focus, Reserva Duque, Manaus, Amazonas, Brazil (DZ 41.192). 39. Artines cofus sp. nov., paratype, Tambopata, Puerto Maldonado Madre de Díos, Peru (MUSM). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241055/files/figure.png" pageId="32" pageNumber="33">Fig. 38</figureCitation>
E–F). Aedeagus narrower on proximal portion; opening of ejaculatory bulb ovoid and separated from proximal margin of aedeagus by short distance; ventral projection bifid, narrow at base, diverging at an angle less than 45 degrees and with distal portion asymmetrically bifid (
<figureCitation id="DBC0FBA89D7ACA6DFC95FA2BFC33FA54" box="[894,976,1448,1475]" captionStart="FIGURES 38–39" captionStartId="29.[151,264,1720,1745]" captionTargetBox="[161,1436,431,1681]" captionTargetId="figure@29.[151,1436,450,1693]" captionTargetPageId="29" captionText="FIGURES 38–39. Male genitalia of the “aquilina group” species. 38. Artines focus, Reserva Duque, Manaus, Amazonas, Brazil (DZ 41.192). 39. Artines cofus sp. nov., paratype, Tambopata, Puerto Maldonado Madre de Díos, Peru (MUSM). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241055/files/figure.png" pageId="32" pageNumber="33">Fig. 38</figureCitation>
G–J). Cornutus with spines as setae from base to median portion of vesica (
<figureCitation id="DBC0FBA89D7ACA6DFDFAFA4FFD62FA71" box="[529,641,1484,1510]" captionStart="FIGURES 38–39" captionStartId="29.[151,264,1720,1745]" captionTargetBox="[161,1436,431,1681]" captionTargetId="figure@29.[151,1436,450,1693]" captionTargetPageId="29" captionText="FIGURES 38–39. Male genitalia of the “aquilina group” species. 38. Artines focus, Reserva Duque, Manaus, Amazonas, Brazil (DZ 41.192). 39. Artines cofus sp. nov., paratype, Tambopata, Puerto Maldonado Madre de Díos, Peru (MUSM). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241055/files/figure.png" pageId="32" pageNumber="33">Figs 38M</figureCitation>
,
<figureCitation id="DBC0FBA89D7ACA6DFD66FA48FD4FFA71" box="[653,684,1483,1510]" captionStart="FIGURES 49–53" captionStartId="34.[151,264,895,920]" captionTargetBox="[155,1421,182,878]" captionTargetId="figure@34.[151,1436,174,883]" captionTargetPageId="34" captionText="FIGURES 49–53. Ventral view of vesica and cornutus of the “aquilina group” species. 49. Artines aquilina. 50. Artines angelica sp. nov.. 51. Artines focus. 52. Artines cofus sp. nov.. 53. Artines donia sp. nov.. Scale bar = 0,2 mm." httpUri="https://zenodo.org/record/3241057/files/figure.png" pageId="32" pageNumber="33">51</figureCitation>
). Fultura inferior, in lateral view, widened distally (
<figureCitation id="DBC0FBA89D7ACA6DFB1BFA4FFAB6FA71" box="[1264,1365,1484,1510]" captionStart="FIGURES 38–39" captionStartId="29.[151,264,1720,1745]" captionTargetBox="[161,1436,431,1681]" captionTargetId="figure@29.[151,1436,450,1693]" captionTargetPageId="29" captionText="FIGURES 38–39. Male genitalia of the “aquilina group” species. 38. Artines focus, Reserva Duque, Manaus, Amazonas, Brazil (DZ 41.192). 39. Artines cofus sp. nov., paratype, Tambopata, Puerto Maldonado Madre de Díos, Peru (MUSM). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241055/files/figure.png" pageId="32" pageNumber="33">Fig. 38L</figureCitation>
).
</paragraph>
<paragraph id="4344E72D9D7ACA6DFF2CFA73FC7CF99D" blockId="32.[151,1437,376,2011]" box="[199,927,1520,1547]" pageId="32" pageNumber="33">
Female (
<figureCitation id="DBC0FBA89D7ACA6DFEC2FA73FE9BF99C" box="[297,376,1520,1547]" captionStart="FIGURES 14–19" captionStartId="10.[151,264,1406,1431]" captionTargetBox="[155,1435,175,1386]" captionTargetId="figure@10.[151,1436,174,1394]" captionTargetPageId="10" captionText="FIGURES 14–19. Species of the “aquilina group” (14–18) and “bipunctata group” (19) in dorsal and ventral views. 14. Artines aquilina, A–B: male, Joinville, Santa Catarina, Brazil (DZ 41.321), C–D: female from same locality of male (DZ 41.220). 15. Artines angelica sp. nov., A–B: holotype male, C–D: allotype female. 16. Artines focus, A–B: male, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.233), C–D: female, Pacaraima, Roraima, Brazil (DZ 41.333). 17. Artines cofus sp. nov., A–B: holotype male, C–D: allotype female. 18. Artines donia sp. nov., A–B: holotype male. 19. Artines bipunctata, A–B: male, Jardim Zoológico, Brasília, Distrito Federal, Brazil (DZ 30.961), C–D: female, Córrego Bananal, Brasília, Distrito Federal, Brazil (DZ 41.224). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241045/files/figure.png" pageId="32" pageNumber="33">Fig. 16</figureCitation>
C–D).
<emphasis id="718F3B3F9D7ACA6DFE2EFA72FD9CF99D" box="[453,639,1520,1546]" italics="true" pageId="32" pageNumber="33">Forewing length</emphasis>
:
<quantity id="84034AC89D7ACA6DFD65FA73FD03F99D" box="[654,736,1520,1546]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="32" pageNumber="33" unit="mm" value="14.0">14 mm</quantity>
. Similar to male.
</paragraph>
<paragraph id="4344E72D9D7ACA6DFF2CF997FCCCF9E1" blockId="32.[151,1437,376,2011]" pageId="32" pageNumber="33">
<emphasis id="718F3B3F9D7ACA6DFF2CF997FED2F9B9" box="[199,305,1556,1582]" italics="true" pageId="32" pageNumber="33">Genitalia</emphasis>
(
<figureCitation id="DBC0FBA89D7ACA6DFEABF990FE76F9BA" box="[320,405,1555,1581]" captionStart="FIGURES 61–65" captionStartId="36.[151,264,653,678]" captionTargetBox="[151,1435,178,637]" captionTargetId="figure@36.[151,1436,174,642]" captionTargetPageId="36" captionText="FIGURES 61–65. Female genitalia of the “aquilina group” and “bipunctata group” species. 61. Artines aquilina, Cananéia, São Paulo, Brazil (OM 49.831). 62. Artines angelica sp. nov., paratype, Tambopata, Puerto Maldonado, Madre de Díos, Peru (DZ 41.222) 63. Artines focus, Pacaraima, Roraima, Brazil (DZ 41.333). 64. Artines cofus sp. nov. paratype, Tambopata, Puerto Maldonado, Madre de Díos, Peru (DZ 41. 293). 65. Artines bipunctata, Reserva IBGE, Brasília, Distrito Federal, Brazil (DZ 41.165). For reference of the letters see legend of the figure 54. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241061/files/figure.png" pageId="32" pageNumber="33">Figs 63</figureCitation>
A–B, 75): lamella antevaginalis narrow; lamella postvaginalis without proximal plate, distal plate sclerotized, without sclerotized median notch on proximal margin, distal margin approximately straight. Ostium on proximal 1/2 of sterigma; ductus bursae narrow.
</paragraph>
<paragraph id="4344E72D9D7ACA6DFF2CF9FDFBFAF977" blockId="32.[151,1437,376,2011]" pageId="32" pageNumber="33">
<emphasis id="718F3B3F9D7ACA6DFF2CF9FDFED9F90E" bold="true" box="[199,314,1662,1689]" pageId="32" pageNumber="33">Variation</emphasis>
. Some specimens from Reserva Duque, Manaus, Amazonas,
<collectingCountry id="3BECA7BD9D7ACA6DFC11F9FCFBDDF90E" box="[1018,1086,1663,1689]" name="Brazil" pageId="32" pageNumber="33">Brazil</collectingCountry>
and from
<collectingCountry id="3BECA7BD9D7ACA6DFB5CF9FCFABCF90E" box="[1207,1375,1663,1689]" name="French Guiana" pageId="32" pageNumber="33">French Guiana</collectingCountry>
have vestigial spot in CuA
<subScript id="DF7FE5689D7ACA6DFE6AF937FE6AF955" attach="left" box="[385,393,1716,1730]" fontSize="6" pageId="32" pageNumber="33">2</subScript>
–2A on ventral forewing. The male genitalia of the specimens from
<collectingCountry id="3BECA7BD9D7ACA6DFB9DF921FAF8F92B" box="[1142,1307,1698,1724]" name="French Guiana" pageId="32" pageNumber="33">French Guiana</collectingCountry>
have uncus wider and ventral projection more developed, with distal end dilated (
<figureCitation id="DBC0FBA89D7ACA6DFC4FF945FBEFF977" box="[932,1036,1734,1760]" captionStart="FIGURES 38–39" captionStartId="29.[151,264,1720,1745]" captionTargetBox="[161,1436,431,1681]" captionTargetId="figure@29.[151,1436,450,1693]" captionTargetPageId="29" captionText="FIGURES 38–39. Male genitalia of the “aquilina group” species. 38. Artines focus, Reserva Duque, Manaus, Amazonas, Brazil (DZ 41.192). 39. Artines cofus sp. nov., paratype, Tambopata, Puerto Maldonado Madre de Díos, Peru (MUSM). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241055/files/figure.png" pageId="32" pageNumber="33">Fig. 38D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D7ACA6DFF2CF96AFCA7F84C" pageId="32" pageNumber="33" type="discussion">
<paragraph id="4344E72D9D7ACA6DFF2CF96AFB5AF8DC" blockId="32.[151,1437,376,2011]" pageId="32" pageNumber="33">
<emphasis id="718F3B3F9D7ACA6DFF2CF96AFEABF893" bold="true" box="[199,328,1769,1796]" pageId="32" pageNumber="33">Comments</emphasis>
.
<taxonomicName id="84FB9CAE9D7ACA6DFEB3F969FE0EF893" authorityName=", Reserva Duque, Manaus" authorityYear="1955" box="[344,493,1770,1796]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="focus">
<emphasis id="718F3B3F9D7ACA6DFEB3F969FE0EF893" box="[344,493,1770,1796]" italics="true" pageId="32" pageNumber="33">Artines focus</emphasis>
</taxonomicName>
is easily recognized for being the only species of the genus that has uncus with a ventral projection. It is sympatric with
<taxonomicName id="84FB9CAE9D7ACA6DFDB8F88DFD27F8BF" box="[595,708,1806,1832]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D7ACA6DFDB8F88DFD27F8BF" box="[595,708,1806,1832]" italics="true" pageId="32" pageNumber="33">A. aepitus</emphasis>
</taxonomicName>
in the northern Amazon Rainforest, from which it can be easily differentiated by the broad discal spots on forewing and by the presence of brand in the males.
</paragraph>
<paragraph id="4344E72D9D7ACA6DFF2CF8D6FB3BF8F8" blockId="32.[151,1437,376,2011]" box="[199,1240,1876,1904]" pageId="32" pageNumber="33">
The record of
<bibRefCitation id="276A9ADC9D7ACA6DFE8CF8D5FE1FF8F8" author="Evans, W. H." box="[359,508,1876,1904]" pageId="32" pageNumber="33" refId="ref40325" refString="Evans, W. H. (1955) A Catalogue of the American Hesperiidae indicating the Classification and Nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. British Museum, London, 499 pp." type="book" year="1955">Evans (1955)</bibRefCitation>
for
<collectingRegion id="813F29CF9D7ACA6DFDC0F8D6FD59F8F8" box="[555,698,1877,1903]" country="Brazil" name="Pernambuco" pageId="32" pageNumber="33">Pernambuco</collectingRegion>
,
<collectingCountry id="3BECA7BD9D7ACA6DFD2EF8D6FCEAF8F8" box="[709,777,1877,1903]" name="Brazil" pageId="32" pageNumber="33">Brazil</collectingCountry>
is probably from a mislabeled specimen.
</paragraph>
<paragraph id="4344E72D9D7ACA6DFF2CF8FBFCA7F84C" blockId="32.[151,1437,376,2011]" pageId="32" pageNumber="33">
<emphasis id="718F3B3F9D7ACA6DFF2CF8FBFE0AF804" bold="true" box="[199,489,1912,1939]" pageId="32" pageNumber="33">Geographic distribution</emphasis>
(
<figureCitation id="DBC0FBA89D7ACA6DFE13F8FAFDAEF804" box="[504,589,1913,1939]" captionStart="FIGURE 80" captionStartId="43.[151,250,930,955]" captionTargetBox="[151,1436,187,905]" captionTargetId="figure@43.[151,1436,187,905]" captionTargetPageId="43" captionText="FIGURE 80. Geographic distribution of the “aquilina group” and “bipunctata group” species. The symbols with white inferior half represent records from literature." httpUri="https://zenodo.org/record/3241071/files/figure.png" pageId="32" pageNumber="33">Fig. 80</figureCitation>
). The species occurs in the
<collectingCountry id="3BECA7BD9D7ACA6DFC6FF8FAFC36F804" box="[900,981,1913,1939]" name="Guyana" pageId="32" pageNumber="33">Guiana</collectingCountry>
Shield. VENEZUELA—
<emphasis id="718F3B3F9D7ACA6DFB1FF8FAFA84F804" box="[1268,1383,1913,1939]" italics="true" pageId="32" pageNumber="33">Amazonas</emphasis>
: Yavita. FRENCH GUIANA—
<emphasis id="718F3B3F9D7ACA6DFE38F81EFD0CF820" box="[467,751,1949,1975]" italics="true" pageId="32" pageNumber="33">Saint-Laurent-du-Maroni</emphasis>
: Régina.
<emphasis id="718F3B3F9D7ACA6DFCB4F81EFC22F820" box="[863,961,1949,1975]" italics="true" pageId="32" pageNumber="33">Cayenne</emphasis>
; Mitaraka (
<bibRefCitation id="276A9ADC9D7ACA6DFBACF81EFB18F820" author="Garrouste, R." box="[1095,1275,1949,1975]" pageId="32" pageNumber="33" pagination="139 - 142" refId="ref40750" refString="Garrouste, R. (2003) Premiere contribution entomologique a la connaissance de Tumuc-Humac-Les lepidopteres du Mitaraka. In: Lacomme, D. &amp; Manil, L. (Eds.), Lepidopteres de Guyane. Lepidopteristes parisiense, Paris, pp. 139 - 142." type="book chapter" year="2003">Garrouste 2003</bibRefCitation>
, 2007). BRA- ZIL—
<emphasis id="718F3B3F9D7ACA6DFF34F842FEA2F84D" box="[223,321,1985,2010]" italics="true" pageId="32" pageNumber="33">Roraima</emphasis>
: Alto Alegre, Pacaraima.
<emphasis id="718F3B3F9D7ACA6DFD89F842FD36F84C" box="[610,725,1985,2011]" italics="true" pageId="32" pageNumber="33">Amazonas</emphasis>
: Manaus.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:namePart id="7BABEBCDF4DED5B3A4BF6A8AC04E99C1">Dolibaina, Diego Rodrigo</mods:namePart>
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<mods:name id="957DD9DC5DD4A983BA6FA389702D5472" type="personal">
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<mods:namePart id="1DD917A181FBDE44ECD5DF2D8DE196DC">Carneiro, Eduardo</mods:namePart>
</mods:name>
<mods:name id="EF5266738A2075B5F1DB5AFFCBFF2063" type="personal">
<mods:role id="672C253B5EB4EBB42B0A56D0E88D379A">
<mods:roleTerm id="BC1EB11E993055F3FED5867155B65A9E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9F5D0CDF70972C4E9A55E050C0F05962">Mielke, Olaf Hermann Hendrik</mods:namePart>
</mods:name>
<mods:name id="35E5ACAD944E3F709F3EEF3688B92ECA" type="personal">
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</mods:role>
<mods:namePart id="685ACA2EECF5D1CCC094E2CAAF5658E9">Casagrande, Mirna Martins</mods:namePart>
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<mods:typeOfResource id="F78876FA25B9E926DC3F49A6AEE47E27">text</mods:typeOfResource>
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<mods:title id="5DFE266B869DF84B5533230E11A8A74F">Zootaxa</mods:title>
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<mods:part id="D6D0277F47B4FD8C6CAC826D2B5B3EBA">
<mods:date id="41E6E875137BD3580A265C6835B6430D">2019</mods:date>
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<mods:number id="BE321401ED155819DC6F7CB49186690A">2019-06-07</mods:number>
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<mods:classification id="E28C7A1DADE240CAA15BF2318EDA7E4A">journal article</mods:classification>
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<mods:identifier id="F7FE9104060492A926C3F641D7D1B5D9" type="DOI">10.11646/zootaxa.4614.1.1</mods:identifier>
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<treatment id="CB52563B9D48CA5EFF7CF926FCD2F848" ID-DOI="http://doi.org/10.5281/zenodo.5935856" ID-GBIF-Taxon="157436014" ID-Zenodo-Dep="5935856" LSID="urn:lsid:zoobank.org:act:4E9AB23B-A26F-4961-A2E6-4B8790B57A5A" httpUri="http://treatment.plazi.org/id/CB52563B9D48CA5EFF7CF926FCD2F848" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
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<paragraph id="4344E72D9D48CA5FFF7CF926FD2FF928" blockId="18.[151,974,1701,1799]" box="[151,716,1701,1728]" pageId="18" pageNumber="19">
<heading id="180C50419D48CA5FFF7CF926FD2FF928" bold="true" box="[151,716,1701,1728]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
<emphasis id="718F3B3F9D48CA5FFF7CF926FD2FF928" bold="true" box="[151,716,1701,1728]" pageId="18" pageNumber="19">
<taxonomicName id="84FB9CAE9D48CA5FFF7CF926FD84F928" authority="Medeiros &amp; O. Mielke" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[151,615,1701,1728]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bamba" status="sp. nov.">
<emphasis id="718F3B3F9D48CA5FFF7CF926FEABF957" bold="true" box="[151,328,1701,1728]" italics="true" pageId="18" pageNumber="19">Artines bamba</emphasis>
Medeiros &amp; O. Mielke
</taxonomicName>
<taxonomicNameLabel id="6ABC86449D48CA5FFD85F926FD2FF928" box="[622,716,1701,1727]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="0BE1B4A69D48CA5FFF7CF94AFC2DF890" pageId="18" pageNumber="19" type="description">
<paragraph id="4344E72D9D48CA5FFF7CF94AFE5FF974" blockId="18.[151,974,1701,1799]" box="[151,444,1736,1763]" pageId="18" pageNumber="19">
<figureCitation id="DBC0FBA89D48CA5FFF7CF94AFF0FF974" box="[151,236,1737,1763]" captionStart="FIGURES 9–13" captionStartId="7.[151,264,1180,1205]" captionTargetBox="[152,1436,180,1162]" captionTargetId="figure@7.[151,1436,174,1169]" captionTargetPageId="7" captionText="FIGURES 9–13. Species of the “aepitus group” in dorsal and ventral views. 9. Artines maya, A–B: holotype male. 10. Artines litoralis sp. nov., A–B: holotype male, C–D: allotype female. 11. Artines rica, A–B: male, Turrialba, Cartago, Costa Rica (DZ 41.890), C–D: female from same locality of male (DZ 41.840). 12. Artines liege sp. nov., A–B: holotype male, C–D: allotype female. 13. Artines bamba sp. nov., A–B: holotype male, C–D: tentative female, Moyobamba, San Martin, Peru (OM 79.413). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241043/files/figure.png" pageId="18" pageNumber="19">Figs 13</figureCitation>
,
<figureCitation id="DBC0FBA89D48CA5FFF13F94BFEF5F974" box="[248,278,1736,1763]" captionStart="FIGURES 33–35" captionStartId="25.[151,264,1776,1801]" captionTargetBox="[151,1436,187,1743]" captionTargetId="figure@25.[151,1436,181,1750]" captionTargetPageId="25" captionText="FIGURES 33–35. Male genitalia of the “aepitus group” species. 33. Artines rica, Turrialba, Cartago, Costa Rica (DZ 4.061). 34. Artines liege sp. nov., holotype. 35. Artines bamba sp. nov., paratype, Pakitza, Parque Nacional del Manu, Madre de Díos, Peru (DZ 9.728). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241051/files/figure.png" pageId="18" pageNumber="19">35</figureCitation>
,
<figureCitation id="DBC0FBA89D48CA5FFEC9F94AFEA3F974" box="[290,320,1737,1763]" captionStart="FIGURES 42–48" captionStartId="33.[151,264,1892,1917]" captionTargetBox="[151,1431,625,1874]" captionTargetId="figure@33.[151,1436,625,1880]" captionText="FIGURES 42–48. Dorsal view of vesica and cornutus of the &quot;aepitus group&quot; species. 42. Artines aepitus. 43. Artines panama sp. nov.. 44. Artines delfos sp. nov.. 45. Artines litoralis sp. nov.. 46. Artines rica. 47. Artines liege sp. nov. 48. Artines bamba sp. nov. Scale bar = 0,2 mm." httpUri="https://zenodo.org/record/3406274/files/figure.png" pageId="18" pageNumber="19">48</figureCitation>
, 60, 72, 78
</paragraph>
<paragraph id="4344E72D9D48CA5FFF7CF96FFC2DF890" blockId="18.[151,974,1701,1799]" box="[151,974,1772,1799]" pageId="18" pageNumber="19">
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urn:lsid:zoobank.org:act:
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<subSubSection id="0BE1B4A69D48CA5FFF7CF8CAFA61F847" pageId="18" pageNumber="19" type="diagnosis">
<paragraph id="4344E72D9D48CA5FFF7CF8CAFA61F847" blockId="18.[151,1436,1865,2037]" pageId="18" pageNumber="19">
<emphasis id="718F3B3F9D48CA5FFF7CF8CAFEE9F8F3" bold="true" box="[151,266,1865,1892]" pageId="18" pageNumber="19">Diagnosis</emphasis>
.
<taxonomicName id="84FB9CAE9D48CA5FFEFCF8C8FE5AF8F3" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[279,441,1866,1893]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bamba" status="sp. nov.">
<emphasis id="718F3B3F9D48CA5FFEFCF8C8FE5AF8F3" box="[279,441,1866,1893]" italics="true" pageId="18" pageNumber="19">Artines bamba</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D48CA5FFE55F8C9FDF4F8F3" bold="true" box="[446,535,1866,1892]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="6ABC86449D48CA5FFE55F8C9FDF4F8F3" box="[446,535,1866,1892]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from the other species of the “
<taxonomicName id="84FB9CAE9D48CA5FFC58F8C8FBE7F8F3" baseAuthorityName="Geyer" baseAuthorityYear="1832" box="[947,1028,1867,1892]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D48CA5FFC58F8C8FBE7F8F3" box="[947,1028,1867,1892]" italics="true" pageId="18" pageNumber="19">aepitus</emphasis>
</taxonomicName>
group” in the following combination of characters: harpe with ventral process narrow and cylindrical, strongly arched inwards and distally pointed, with long and thick setae on subapical portion; cornutus with long and robust spines, curved at base and directed anteriorly; proximo-lateral projections of lamella postvaginalis developed; ductus bursae with sclerotized plates.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D48CA5EFF2CF85AFBDFFB84" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="description">
<paragraph id="4344E72D9D48CA5FFF2CF85AFB89F862" blockId="18.[151,1436,1865,2037]" box="[199,1130,2009,2037]" pageId="18" pageNumber="19">
<emphasis id="718F3B3F9D48CA5FFF2CF85AFEB2F863" bold="true" box="[199,337,2009,2036]" pageId="18" pageNumber="19">Description</emphasis>
. Male (
<figureCitation id="DBC0FBA89D48CA5FFE43F859FE1BF863" box="[424,504,2010,2036]" captionStart="FIGURES 9–13" captionStartId="7.[151,264,1180,1205]" captionTargetBox="[152,1436,180,1162]" captionTargetId="figure@7.[151,1436,174,1169]" captionTargetPageId="7" captionText="FIGURES 9–13. Species of the “aepitus group” in dorsal and ventral views. 9. Artines maya, A–B: holotype male. 10. Artines litoralis sp. nov., A–B: holotype male, C–D: allotype female. 11. Artines rica, A–B: male, Turrialba, Cartago, Costa Rica (DZ 41.890), C–D: female from same locality of male (DZ 41.840). 12. Artines liege sp. nov., A–B: holotype male, C–D: allotype female. 13. Artines bamba sp. nov., A–B: holotype male, C–D: tentative female, Moyobamba, San Martin, Peru (OM 79.413). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241043/files/figure.png" pageId="18" pageNumber="19">Fig. 13</figureCitation>
A–B).
<emphasis id="718F3B3F9D48CA5FFDAEF859FD61F863" box="[581,642,2010,2036]" italics="true" pageId="18" pageNumber="19">Head</emphasis>
: antenna with nudum bearing 12 segments.
</paragraph>
<paragraph id="4344E72D9D49CA5EFF2CFF19FDF3FF24" blockId="19.[151,1437,152,2015]" box="[199,528,152,179]" pageId="19" pageNumber="20">
<emphasis id="718F3B3F9D49CA5EFF2CFF19FE62FF24" box="[199,385,153,179]" italics="true" pageId="19" pageNumber="20">Forewing length</emphasis>
:
<quantity id="84034AC89D49CA5EFE7BFF1BFDEFFF24" box="[400,524,152,179]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="19" pageNumber="20" unit="mm" value="13.5" valueMax="15.0" valueMin="12.0">12–15 mm</quantity>
.
</paragraph>
<paragraph id="4344E72D9D49CA5EFF2CFF3EFD83FF6C" blockId="19.[151,1437,152,2015]" pageId="19" pageNumber="20">
<emphasis id="718F3B3F9D49CA5EFF2CFF3EFE9CFF40" box="[199,383,189,215]" italics="true" pageId="19" pageNumber="20">Dorsal forewing</emphasis>
(
<figureCitation id="DBC0FBA89D49CA5EFE67FF3EFE17FF40" box="[396,500,189,215]" captionStart="FIGURES 9–13" captionStartId="7.[151,264,1180,1205]" captionTargetBox="[152,1436,180,1162]" captionTargetId="figure@7.[151,1436,174,1169]" captionTargetPageId="7" captionText="FIGURES 9–13. Species of the “aepitus group” in dorsal and ventral views. 9. Artines maya, A–B: holotype male. 10. Artines litoralis sp. nov., A–B: holotype male, C–D: allotype female. 11. Artines rica, A–B: male, Turrialba, Cartago, Costa Rica (DZ 41.890), C–D: female from same locality of male (DZ 41.840). 12. Artines liege sp. nov., A–B: holotype male, C–D: allotype female. 13. Artines bamba sp. nov., A–B: holotype male, C–D: tentative female, Moyobamba, San Martin, Peru (OM 79.413). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241043/files/figure.png" pageId="19" pageNumber="20">Fig. 13A</figureCitation>
): two or three apical spots (absent in most specimens); discal spots absent or reduced to a few scales; spot in CuA
<subScript id="DF7FE5689D49CA5EFE3AFF71FE3AFE97" attach="left" box="[465,473,242,256]" fontSize="6" pageId="19" pageNumber="20">2</subScript>
–2A absent.
</paragraph>
<paragraph id="4344E72D9D49CA5EFF2CFE86FD21FE88" blockId="19.[151,1437,152,2015]" box="[199,706,261,287]" pageId="19" pageNumber="20">
<emphasis id="718F3B3F9D49CA5EFF2CFE86FE65FE88" box="[199,390,261,287]" italics="true" pageId="19" pageNumber="20">Ventral forewing</emphasis>
(
<figureCitation id="DBC0FBA89D49CA5EFE7EFE86FE1FFE88" box="[405,508,261,287]" captionStart="FIGURES 9–13" captionStartId="7.[151,264,1180,1205]" captionTargetBox="[152,1436,180,1162]" captionTargetId="figure@7.[151,1436,174,1169]" captionTargetPageId="7" captionText="FIGURES 9–13. Species of the “aepitus group” in dorsal and ventral views. 9. Artines maya, A–B: holotype male. 10. Artines litoralis sp. nov., A–B: holotype male, C–D: allotype female. 11. Artines rica, A–B: male, Turrialba, Cartago, Costa Rica (DZ 41.890), C–D: female from same locality of male (DZ 41.840). 12. Artines liege sp. nov., A–B: holotype male, C–D: allotype female. 13. Artines bamba sp. nov., A–B: holotype male, C–D: tentative female, Moyobamba, San Martin, Peru (OM 79.413). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241043/files/figure.png" pageId="19" pageNumber="20">Fig. 13B</figureCitation>
): as in
<taxonomicName id="84FB9CAE9D49CA5EFDA0FE86FD58FE88" baseAuthorityName="Geyer" baseAuthorityYear="1832" box="[587,699,261,287]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D49CA5EFDA0FE86FD58FE88" box="[587,699,261,287]" italics="true" pageId="19" pageNumber="20">A. aepitus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="4344E72D9D49CA5EFF2CFEAAFD6DFEF0" blockId="19.[151,1437,152,2015]" pageId="19" pageNumber="20">
<emphasis id="718F3B3F9D49CA5EFF2CFEAAFE6CFED4" box="[199,399,297,323]" italics="true" pageId="19" pageNumber="20">Ventral hind wing</emphasis>
(
<figureCitation id="DBC0FBA89D49CA5EFE76FEAAFDE0FED4" box="[413,515,297,323]" captionStart="FIGURES 9–13" captionStartId="7.[151,264,1180,1205]" captionTargetBox="[152,1436,180,1162]" captionTargetId="figure@7.[151,1436,174,1169]" captionTargetPageId="7" captionText="FIGURES 9–13. Species of the “aepitus group” in dorsal and ventral views. 9. Artines maya, A–B: holotype male. 10. Artines litoralis sp. nov., A–B: holotype male, C–D: allotype female. 11. Artines rica, A–B: male, Turrialba, Cartago, Costa Rica (DZ 41.890), C–D: female from same locality of male (DZ 41.840). 12. Artines liege sp. nov., A–B: holotype male, C–D: allotype female. 13. Artines bamba sp. nov., A–B: holotype male, C–D: tentative female, Moyobamba, San Martin, Peru (OM 79.413). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241043/files/figure.png" pageId="19" pageNumber="20">Fig. 13B</figureCitation>
): apex of discal cell with few yellow scales; central black spot quadrate; blue discal spots in M
<subScript id="DF7FE5689D49CA5EFEE6FEDDFEF6FEFB" attach="left" box="[269,277,350,364]" fontSize="6" pageId="19" pageNumber="20">3</subScript>
–CuA
<subScript id="DF7FE5689D49CA5EFEB3FEDDFE83FEFB" attach="left" box="[344,352,350,364]" fontSize="6" pageId="19" pageNumber="20">1</subScript>
and CuA
<subScript id="DF7FE5689D49CA5EFE27FEDDFE37FEFB" attach="left" box="[460,468,350,364]" fontSize="6" pageId="19" pageNumber="20">1</subScript>
–CuA
<subScript id="DF7FE5689D49CA5EFDFCFEDDFDFCFEFB" attach="left" box="[535,543,350,364]" fontSize="6" pageId="19" pageNumber="20">2</subScript>
quadrate.
</paragraph>
<paragraph id="4344E72D9D49CA5EFF2CFEF2FB9CFD64" blockId="19.[151,1437,152,2015]" pageId="19" pageNumber="20">
<emphasis id="718F3B3F9D49CA5EFF2CFEF2FED2FE1C" box="[199,305,369,395]" italics="true" pageId="19" pageNumber="20">Genitalia</emphasis>
(
<figureCitation id="DBC0FBA89D49CA5EFEAAFEF2FE75FE1C" box="[321,406,368,395]" captionStart="FIGURES 33–35" captionStartId="25.[151,264,1776,1801]" captionTargetBox="[151,1436,187,1743]" captionTargetId="figure@25.[151,1436,181,1750]" captionTargetPageId="25" captionText="FIGURES 33–35. Male genitalia of the “aepitus group” species. 33. Artines rica, Turrialba, Cartago, Costa Rica (DZ 4.061). 34. Artines liege sp. nov., holotype. 35. Artines bamba sp. nov., paratype, Pakitza, Parque Nacional del Manu, Madre de Díos, Peru (DZ 9.728). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241051/files/figure.png" pageId="19" pageNumber="20">Figs 35</figureCitation>
A–L, 48): tegumen more or less quadrate, with proximal margin straight and distal margin not conspicuous (
<figureCitation id="DBC0FBA89D49CA5EFEB2FE16FE5DFE38" box="[345,446,404,431]" captionStart="FIGURES 33–35" captionStartId="25.[151,264,1776,1801]" captionTargetBox="[151,1436,187,1743]" captionTargetId="figure@25.[151,1436,181,1750]" captionTargetPageId="25" captionText="FIGURES 33–35. Male genitalia of the “aepitus group” species. 33. Artines rica, Turrialba, Cartago, Costa Rica (DZ 4.061). 34. Artines liege sp. nov., holotype. 35. Artines bamba sp. nov., paratype, Pakitza, Parque Nacional del Manu, Madre de Díos, Peru (DZ 9.728). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241051/files/figure.png" pageId="19" pageNumber="20">Fig. 35C</figureCitation>
), in lateral view shorter than length of gnathos; ventral arm of tegumen longer than dorsal arm of saccus (
<figureCitation id="DBC0FBA89D49CA5EFEAFFE3AFE4DFE44" box="[324,430,440,467]" captionStart="FIGURES 33–35" captionStartId="25.[151,264,1776,1801]" captionTargetBox="[151,1436,187,1743]" captionTargetId="figure@25.[151,1436,181,1750]" captionTargetPageId="25" captionText="FIGURES 33–35. Male genitalia of the “aepitus group” species. 33. Artines rica, Turrialba, Cartago, Costa Rica (DZ 4.061). 34. Artines liege sp. nov., holotype. 35. Artines bamba sp. nov., paratype, Pakitza, Parque Nacional del Manu, Madre de Díos, Peru (DZ 9.728). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241051/files/figure.png" pageId="19" pageNumber="20">Fig. 35A</figureCitation>
). Saccus as long as tegumen+uncus, base as long as lateral margin (
<figureCitation id="DBC0FBA89D49CA5EFB58FE3AFAF8FE44" box="[1203,1307,440,467]" captionStart="FIGURES 33–35" captionStartId="25.[151,264,1776,1801]" captionTargetBox="[151,1436,187,1743]" captionTargetId="figure@25.[151,1436,181,1750]" captionTargetPageId="25" captionText="FIGURES 33–35. Male genitalia of the “aepitus group” species. 33. Artines rica, Turrialba, Cartago, Costa Rica (DZ 4.061). 34. Artines liege sp. nov., holotype. 35. Artines bamba sp. nov., paratype, Pakitza, Parque Nacional del Manu, Madre de Díos, Peru (DZ 9.728). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241051/files/figure.png" pageId="19" pageNumber="20">Fig. 35B</figureCitation>
). Uncus, in dorsal view, twice longer than wide, longer than tegumen, without disto-lateral projections, distal margin rounded (
<figureCitation id="DBC0FBA89D49CA5EFF4BFD82FEE6FD8C" box="[160,261,512,539]" captionStart="FIGURES 33–35" captionStartId="25.[151,264,1776,1801]" captionTargetBox="[151,1436,187,1743]" captionTargetId="figure@25.[151,1436,181,1750]" captionTargetPageId="25" captionText="FIGURES 33–35. Male genitalia of the “aepitus group” species. 33. Artines rica, Turrialba, Cartago, Costa Rica (DZ 4.061). 34. Artines liege sp. nov., holotype. 35. Artines bamba sp. nov., paratype, Pakitza, Parque Nacional del Manu, Madre de Díos, Peru (DZ 9.728). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241051/files/figure.png" pageId="19" pageNumber="20">Fig. 35C</figureCitation>
). Gnathos as in the other species of the group (
<figureCitation id="DBC0FBA89D49CA5EFCE6FD82FC75FD8C" box="[781,918,512,539]" captionStart="FIGURES 33–35" captionStartId="25.[151,264,1776,1801]" captionTargetBox="[151,1436,187,1743]" captionTargetId="figure@25.[151,1436,181,1750]" captionTargetPageId="25" captionText="FIGURES 33–35. Male genitalia of the “aepitus group” species. 33. Artines rica, Turrialba, Cartago, Costa Rica (DZ 4.061). 34. Artines liege sp. nov., holotype. 35. Artines bamba sp. nov., paratype, Pakitza, Parque Nacional del Manu, Madre de Díos, Peru (DZ 9.728). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241051/files/figure.png" pageId="19" pageNumber="20">Fig. 35A, D</figureCitation>
). Valva three times longer than wide; sacculus elongate and about 1/2 width of valva; costa narrow and contiguous with ampulla; ampulla triangular and shorter than costa; ventral process of harpe narrow, elongate, cylindrical and strongly arched inwards, ventral margin slightly sinuous and distal end pointed, with subapical setae thick and long; dorsal process of harpe on median portion, rounded in dorsal view, wider than ventral process, apex surpassing dorsal margin of valva (
<figureCitation id="DBC0FBA89D49CA5EFBC2FD12FB9BFD3C" box="[1065,1144,656,683]" captionStart="FIGURES 33–35" captionStartId="25.[151,264,1776,1801]" captionTargetBox="[151,1436,187,1743]" captionTargetId="figure@25.[151,1436,181,1750]" captionTargetPageId="25" captionText="FIGURES 33–35. Male genitalia of the “aepitus group” species. 33. Artines rica, Turrialba, Cartago, Costa Rica (DZ 4.061). 34. Artines liege sp. nov., holotype. 35. Artines bamba sp. nov., paratype, Pakitza, Parque Nacional del Manu, Madre de Díos, Peru (DZ 9.728). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241051/files/figure.png" pageId="19" pageNumber="20">Fig. 35</figureCitation>
E–F). Aedeagus as long as valva, slightly curved dorsally; ventral projection distally round and strongly curved ventrally (
<figureCitation id="DBC0FBA89D49CA5EFB42FD36FB1BFD58" box="[1193,1272,692,719]" captionStart="FIGURES 33–35" captionStartId="25.[151,264,1776,1801]" captionTargetBox="[151,1436,187,1743]" captionTargetId="figure@25.[151,1436,181,1750]" captionTargetPageId="25" captionText="FIGURES 33–35. Male genitalia of the “aepitus group” species. 33. Artines rica, Turrialba, Cartago, Costa Rica (DZ 4.061). 34. Artines liege sp. nov., holotype. 35. Artines bamba sp. nov., paratype, Pakitza, Parque Nacional del Manu, Madre de Díos, Peru (DZ 9.728). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241051/files/figure.png" pageId="19" pageNumber="20">Fig. 35</figureCitation>
G–J); posterior end of cornutus wide and triangular, with spines thick, long and curved anteriorly (
<figureCitation id="DBC0FBA89D49CA5EFBCBFD5AFB91FD64" box="[1056,1138,729,755]" captionStart="FIGURES 42–48" captionStartId="33.[151,264,1892,1917]" captionTargetBox="[151,1431,625,1874]" captionTargetId="figure@33.[151,1436,625,1880]" captionText="FIGURES 42–48. Dorsal view of vesica and cornutus of the &quot;aepitus group&quot; species. 42. Artines aepitus. 43. Artines panama sp. nov.. 44. Artines delfos sp. nov.. 45. Artines litoralis sp. nov.. 46. Artines rica. 47. Artines liege sp. nov. 48. Artines bamba sp. nov. Scale bar = 0,2 mm." httpUri="https://zenodo.org/record/3406274/files/figure.png" pageId="19" pageNumber="20">Fig. 48</figureCitation>
).
</paragraph>
<paragraph id="4344E72D9D49CA5EFF2CFD7EFC3DFC80" blockId="19.[151,1437,152,2015]" box="[199,990,764,791]" pageId="19" pageNumber="20">
Female (
<figureCitation id="DBC0FBA89D49CA5EFEC2FD7EFE9BFC80" box="[297,376,765,791]" captionStart="FIGURES 9–13" captionStartId="7.[151,264,1180,1205]" captionTargetBox="[152,1436,180,1162]" captionTargetId="figure@7.[151,1436,174,1169]" captionTargetPageId="7" captionText="FIGURES 9–13. Species of the “aepitus group” in dorsal and ventral views. 9. Artines maya, A–B: holotype male. 10. Artines litoralis sp. nov., A–B: holotype male, C–D: allotype female. 11. Artines rica, A–B: male, Turrialba, Cartago, Costa Rica (DZ 41.890), C–D: female from same locality of male (DZ 41.840). 12. Artines liege sp. nov., A–B: holotype male, C–D: allotype female. 13. Artines bamba sp. nov., A–B: holotype male, C–D: tentative female, Moyobamba, San Martin, Peru (OM 79.413). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241043/files/figure.png" pageId="19" pageNumber="20">Fig. 13</figureCitation>
C–D).
<emphasis id="718F3B3F9D49CA5EFE2EFD7DFD9CFC80" box="[453,639,765,791]" italics="true" pageId="19" pageNumber="20">Forewing length</emphasis>
:
<quantity id="84034AC89D49CA5EFD65FD7FFCFCFC80" box="[654,799,764,791]" metricMagnitude="-2" metricUnit="m" metricValue="1.375" metricValueMax="1.4" metricValueMin="1.35" pageId="19" pageNumber="20" unit="mm" value="13.75" valueMax="14.0" valueMin="13.5">13.5–14 mm</quantity>
. Similar to male.
</paragraph>
<paragraph id="4344E72D9D49CA5EFF2CFCA2FCBCFC30" blockId="19.[151,1437,152,2015]" pageId="19" pageNumber="20">
<emphasis id="718F3B3F9D49CA5EFF2CFCA2FED2FCAC" box="[199,305,801,827]" italics="true" pageId="19" pageNumber="20">Genitalia</emphasis>
(
<figureCitation id="DBC0FBA89D49CA5EFEADFCA2FE42FCAC" box="[326,417,801,827]" captionStart="FIGURES 54–60" captionStartId="35.[151,264,1807,1832]" captionTargetBox="[156,1436,575,1772]" captionTargetId="figure@35.[151,1436,589,1795]" captionTargetPageId="35" captionText="FIGURES 54–60. Female genitalia of the “aepitus group” species. 54. Artines aepitus, Jusepín, Monagas, Venezuela (OM 30. 125). 55. Artines panama sp. nov., allotype. 56. Artines solange sp. nov. (tentative female), Sumaúma, Maranhão, Brazil (DZ 41.800). 57. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.820). 58. Artines rica, Turrialba, Cartago, Costa Rica (DZ 41.840). 59. Artines liege sp. nov., paratype, Parque do Gama, Brasília, Distrito Federal, Brazil (DZ 41.830). 60. Artines bamba sp. nov. (tentative female), Moyobamba, San Martin, Peru (OM 81.229). A) lateral view of tergum VIII, sterigma and papilla analis; B) ventral view of esterigma. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241059/files/figure.png" pageId="19" pageNumber="20">Figs 60</figureCitation>
A–B, 72): tergum VIII with complete spiracular opening. Sterigma triangular; lamella antevaginalis with proximo-ventral folds narrow; lamella postvaginalis without lateral fold, distal projections as in
<taxonomicName id="84FB9CAE9D49CA5EFF7CFCEAFEEAFC14" baseAuthorityName="Geyer" baseAuthorityYear="1832" box="[151,265,873,899]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D49CA5EFF7CFCEAFEEAFC14" box="[151,265,873,899]" italics="true" pageId="19" pageNumber="20">A. aepitus</emphasis>
</taxonomicName>
, proximo-lateral projections developed and bilobed. Ostium narrow surrounded by membranous areas; ductus bursae with sclerotized plates near insertion of sterigma.
</paragraph>
<paragraph id="4344E72D9D49CA5EFF2CFC33FBDFFB84" blockId="19.[151,1437,152,2015]" pageId="19" pageNumber="20">
<emphasis id="718F3B3F9D49CA5EFF2CFC33FED9FC5C" bold="true" box="[199,314,944,971]" pageId="19" pageNumber="20">Variation</emphasis>
. As in most species of the “
<taxonomicName id="84FB9CAE9D49CA5EFD84FC31FD23FC5C" baseAuthorityName="Geyer" baseAuthorityYear="1832" box="[623,704,946,971]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D49CA5EFD84FC31FD23FC5C" box="[623,704,946,971]" italics="true" pageId="19" pageNumber="20">aepitus</emphasis>
</taxonomicName>
group”, the variation is in the shape of the forewing marks. These marks are absent in most individuals analyzed and only the
<typeStatus id="9C40598F9D49CA5EFCDFFC56FC75FC78" box="[820,918,981,1007]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
has three apical spots and two male
<typeStatus id="9C40598F9D49CA5EFADAFC56FA7FFC78" box="[1329,1436,981,1007]" pageId="19" pageNumber="20" type="paratype">paratypes</typeStatus>
have conspicuous discal spots. The marks are present in females from
<collectingRegion id="813F29CF9D49CA5EFC47FC7AFC06FB84" box="[940,997,1017,1043]" country="Brazil" name="Acre" pageId="19" pageNumber="20">Acre</collectingRegion>
,
<collectingCountry id="3BECA7BD9D49CA5EFC1BFC7AFBDBFB84" box="[1008,1080,1017,1043]" name="Brazil" pageId="19" pageNumber="20">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D49CA5EFF2CFB9FFA6CFAEC" pageId="19" pageNumber="20" type="discussion">
<paragraph id="4344E72D9D49CA5EFF2CFB9FFD8AFB34" blockId="19.[151,1437,152,2015]" pageId="19" pageNumber="20">
<emphasis id="718F3B3F9D49CA5EFF2CFB9FFEABFBA0" bold="true" box="[199,328,1052,1079]" pageId="19" pageNumber="20">Comments</emphasis>
. Species similar to
<taxonomicName id="84FB9CAE9D49CA5EFDC8FB9EFD8CFBA0" authorityName=", S. R. Steinhauser &amp; G. T. Austin" authorityYear="1993" box="[547,623,1053,1079]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="rica">
<emphasis id="718F3B3F9D49CA5EFDC8FB9EFD8CFBA0" box="[547,623,1053,1079]" italics="true" pageId="19" pageNumber="20">A. rica</emphasis>
</taxonomicName>
and
<taxonomicName id="84FB9CAE9D49CA5EFD4EFB9EFD19FBA0" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[677,762,1053,1079]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="liege" status="sp. nov.">
<emphasis id="718F3B3F9D49CA5EFD4EFB9EFD19FBA0" box="[677,762,1053,1079]" italics="true" pageId="19" pageNumber="20">A. liege</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6ABC86449D49CA5EFCEAFB9EFCBFFBA0" box="[769,860,1053,1079]" pageId="19" pageNumber="20" rank="species">
<emphasis id="718F3B3F9D49CA5EFCEAFB9EFCB6FBA0" bold="true" box="[769,853,1053,1079]" pageId="19" pageNumber="20">sp. nov</emphasis>
.
</taxonomicNameLabel>
in the genitalia, differing in the cornutus with long, robust spines directed anteriorly and ventral projection of harpe more elongated, distally pointed, strongly arched inwards, with setae on subapical portion. This species is unique among the “
<taxonomicName id="84FB9CAE9D49CA5EFC35FBE5FBCCFBE8" baseAuthorityName="Geyer" baseAuthorityYear="1832" box="[990,1071,1126,1151]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D49CA5EFC35FBE5FBCCFBE8" box="[990,1071,1126,1151]" italics="true" pageId="19" pageNumber="20">aepitus</emphasis>
</taxonomicName>
group” with the sclerotized plate at base of ductus bursae (
<figureCitation id="DBC0FBA89D49CA5EFE5AFB0AFDE5FB34" box="[433,518,1161,1187]" captionStart="FIGURES 54–60" captionStartId="35.[151,264,1807,1832]" captionTargetBox="[156,1436,575,1772]" captionTargetId="figure@35.[151,1436,589,1795]" captionTargetPageId="35" captionText="FIGURES 54–60. Female genitalia of the “aepitus group” species. 54. Artines aepitus, Jusepín, Monagas, Venezuela (OM 30. 125). 55. Artines panama sp. nov., allotype. 56. Artines solange sp. nov. (tentative female), Sumaúma, Maranhão, Brazil (DZ 41.800). 57. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.820). 58. Artines rica, Turrialba, Cartago, Costa Rica (DZ 41.840). 59. Artines liege sp. nov., paratype, Parque do Gama, Brasília, Distrito Federal, Brazil (DZ 41.830). 60. Artines bamba sp. nov. (tentative female), Moyobamba, San Martin, Peru (OM 81.229). A) lateral view of tergum VIII, sterigma and papilla analis; B) ventral view of esterigma. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241059/files/figure.png" pageId="19" pageNumber="20">Figs 60</figureCitation>
and
<figureCitation id="DBC0FBA89D49CA5EFDD6FB0AFDBEFB34" box="[573,605,1161,1187]" captionStart="FIGURES 66–72" captionStartId="37.[151,264,1734,1759]" captionTargetBox="[160,1433,189,1717]" captionTargetId="figure@37.[151,1436,174,1723]" captionTargetPageId="37" captionText="FIGURES 66–72. Female genitalia of the “aepitus group” species (ventral view of sterigma and bursa copulatrix). 66. Artines aepitus, Jusepín, Monagas, Venezuela (OM 30. 125). 67. Artines panama sp. nov., allotype. 68. Artines solange sp. nov. (tentative female), Sumaúma, Maranhão, Brazil (DZ 41.800). 69. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.820). 70. Artines rica, Turrialba, Cartago, Costa Rica (DZ 41.840). 71. Artines liege sp. nov., paratype, Parque do Gama, Brasília, Distrito Federal, Brazil (DZ 41.830). 72. Artines bamba sp. nov. (tentative female), Moyobamba, San Martin, Peru (OM 81.229). Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241063/files/figure.png" pageId="19" pageNumber="20">72</figureCitation>
).
</paragraph>
<paragraph id="4344E72D9D49CA5EFF2CFB2EFE13FAA4" blockId="19.[151,1437,152,2015]" pageId="19" pageNumber="20">The association of the female in this species was made based on the absence of apical and discal spots on dorsal forewing (see variation above) and distribution. However, in view of the fact that species are similar and different species can potentially occur in the same locality, the association is tentative and for this reason the females were not included in the type-series.</paragraph>
<paragraph id="4344E72D9D49CA5EFF2CFABFFA6CFAEC" blockId="19.[151,1437,152,2015]" pageId="19" pageNumber="20">
<emphasis id="718F3B3F9D49CA5EFF2CFABFFE04FAC0" bold="true" box="[199,487,1340,1367]" pageId="19" pageNumber="20">Geographic distribution</emphasis>
(
<figureCitation id="DBC0FBA89D49CA5EFE1FFABEFDA4FAC0" box="[500,583,1341,1367]" captionStart="FIGURE 78" captionStartId="42.[151,250,910,935]" captionTargetBox="[151,1436,174,898]" captionTargetId="figure@42.[151,1436,174,898]" captionTargetPageId="42" captionText="FIGURE 78. Geographic distribution of the “aepitus group” species. The symbols with white inferior half represent records from literature." httpUri="https://zenodo.org/record/3241067/files/figure.png" pageId="19" pageNumber="20">Fig. 78</figureCitation>
). The species occurs in the western Amazon. BRAZIL—
<emphasis id="718F3B3F9D49CA5EFB52FABEFACFFAC0" box="[1209,1324,1341,1367]" italics="true" pageId="19" pageNumber="20">
<collectingRegion id="813F29CF9D49CA5EFB52FABEFACFFAC0" box="[1209,1324,1341,1367]" country="Venezuela" name="Amazonas" pageId="19" pageNumber="20">Amazonas</collectingRegion>
</emphasis>
:
<collectingRegion id="813F29CF9D49CA5EFAD1FABEFF4EFAEC" country="Brazil" name="Sao Paulo" pageId="19" pageNumber="20">São Paulo</collectingRegion>
de Olivença.
<emphasis id="718F3B3F9D49CA5EFEA3FAE2FE9FFAEC" box="[328,380,1377,1403]" italics="true" pageId="19" pageNumber="20">
<collectingRegion id="813F29CF9D49CA5EFEA3FAE2FE9FFAEC" box="[328,380,1377,1403]" country="Brazil" name="Acre" pageId="19" pageNumber="20">Acre</collectingRegion>
</emphasis>
: Senador Guiomard, Porto
<collectingRegion id="813F29CF9D49CA5EFD44FAE2FD0AFAEC" box="[687,745,1377,1403]" country="Brazil" name="Acre" pageId="19" pageNumber="20">Acre</collectingRegion>
. PERU—
<emphasis id="718F3B3F9D49CA5EFCBCFAE2FBE3FAEC" box="[855,1024,1377,1403]" italics="true" pageId="19" pageNumber="20">
<collectingRegion id="813F29CF9D49CA5EFCBCFAE2FBE3FAEC" box="[855,1024,1377,1403]" country="Peru" name="Madre de Dios" pageId="19" pageNumber="20">Madre de Díos</collectingRegion>
</emphasis>
.
<emphasis id="718F3B3F9D49CA5EFBE6FAE2FB6FFAEC" box="[1037,1164,1377,1403]" italics="true" pageId="19" pageNumber="20">
<collectingRegion id="813F29CF9D49CA5EFBE6FAE2FB6FFAEC" box="[1037,1164,1377,1403]" country="Peru" name="San Martin" pageId="19" pageNumber="20">San Martin</collectingRegion>
</emphasis>
: Juanjui, Moyobamba.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D49CA5EFF2CFA06FBC9FA08" box="[199,1066,1412,1439]" pageId="19" pageNumber="20" type="description">
<paragraph id="4344E72D9D49CA5EFF2CFA06FBC9FA08" blockId="19.[151,1437,152,2015]" box="[199,1066,1412,1439]" pageId="19" pageNumber="20">
<emphasis id="718F3B3F9D49CA5EFF2CFA06FE3AFA08" bold="true" box="[199,473,1412,1439]" pageId="19" pageNumber="20">Temporal distribution.</emphasis>
This species occurs in July, September and October.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D49CA5EFF2CFA2AFE2AFA54" box="[199,457,1449,1475]" pageId="19" pageNumber="20" type="biology_ecology">
<paragraph id="4344E72D9D49CA5EFF2CFA2AFE2AFA54" blockId="19.[151,1437,152,2015]" box="[199,457,1449,1475]" pageId="19" pageNumber="20">
<emphasis id="718F3B3F9D49CA5EFF2CFA2AFEA7FA54" bold="true" box="[199,324,1449,1475]" pageId="19" pageNumber="20">Host plant</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D49CA5EFF2CFA4EFA94FA70" box="[199,1399,1485,1511]" pageId="19" pageNumber="20" type="etymology">
<paragraph id="4344E72D9D49CA5EFF2CFA4EFA94FA70" blockId="19.[151,1437,152,2015]" box="[199,1399,1485,1511]" pageId="19" pageNumber="20">
<emphasis id="718F3B3F9D49CA5EFF2CFA4EFEABFA70" bold="true" box="[199,328,1485,1511]" pageId="19" pageNumber="20">Etymology</emphasis>
. The term &quot;
<taxonomicName id="84FB9CAE9D49CA5EFE24FA4EFDF8FA70" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[463,539,1485,1511]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="bamba">
<emphasis id="718F3B3F9D49CA5EFE24FA4EFDF8FA70" box="[463,539,1485,1511]" italics="true" pageId="19" pageNumber="20">bamba</emphasis>
</taxonomicName>
&quot; is the termination of the word Moyobamba, the
<typeStatus id="9C40598F9D49CA5EFBAEFA4EFB96FA70" box="[1093,1141,1485,1511]" pageId="19" pageNumber="20">type</typeStatus>
locality of the species.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D49CA5EFF2CFA72FCD2F848" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="4344E72D9D49CA5EFF2CFA72FE88F9C4" blockId="19.[151,1437,152,2015]" pageId="19" pageNumber="20">
<emphasis id="718F3B3F9D49CA5EFF2CFA72FE92F99C" bold="true" box="[199,369,1520,1547]" pageId="19" pageNumber="20">Type material</emphasis>
.
<materialsCitation id="F393ED709D49CA5EFE6BFA72FBB4F99C" ID-GBIF-Occurrence="2248478866" box="[384,1111,1521,1547]" collectionCode="DZUP" pageId="19" pageNumber="20" specimenCount="1" typeStatus="holotype">
<typeStatus id="9C40598F9D49CA5EFE6BFA72FE0BF99C" box="[384,488,1521,1547]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
male deposited at the
<collectionCode id="25EA7FE89D49CA5EFD03FA72FCD2F99C" box="[744,817,1521,1547]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="19" pageNumber="20" type="University or college">DZUP</collectionCode>
with the following labels
</materialsCitation>
: /
<materialsCitation id="F393ED709D49CA5EFB99FA72FC46F9B8" ID-GBIF-Occurrence="2248478803" collectingDate="2015-10" collectorName="J. Pintado" country="Peru" location="Moyobamba" pageId="19" pageNumber="20" specimenCount="1" stateProvince="San Martin" typeStatus="holotype">
<typeStatus id="9C40598F9D49CA5EFB99FA72FAFBF99C" box="[1138,1304,1521,1547]" pageId="19" pageNumber="20" type="holotype">HOLOTYPUS</typeStatus>
/
<collectingCountry id="3BECA7BD9D49CA5EFADBFA72FA85F99C" box="[1328,1382,1521,1547]" name="Peru" pageId="19" pageNumber="20">Peru</collectingCountry>
,
<collectingRegion id="813F29CF9D49CA5EFA99FA72FF05F9B8" country="Peru" name="San Martin" pageId="19" pageNumber="20">San Martin</collectingRegion>
,
<location id="4624B1F69D49CA5EFF1FF996FE67F9B8" LSID="urn:lsid:plazi:treatment:CB52563B9D48CA5EFF7CF926FCD2F848:4624B1F69D49CA5EFF1FF996FE67F9B8" box="[244,388,1557,1583]" country="Peru" name="Moyobamba" pageId="19" pageNumber="20" stateProvince="San Martin">Moyobamba</location>
, Calzada.
<date id="3745C1ED9D49CA5EFE16F996FDB0F9B8" box="[509,595,1556,1583]" pageId="19" pageNumber="20" value="2015-10">
<collectingDate id="270138059D49CA5EFE16F996FDB0F9B8" box="[509,595,1556,1583]" pageId="19" pageNumber="20" value="2015-10">X.2015</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D49CA5EFD8AF996FD31F9B8" box="[609,722,1557,1583]" pageId="19" pageNumber="20">J. Pintado</collectorName>
<emphasis id="718F3B3F9D49CA5EFD30F996FCE6F9B8" box="[731,773,1557,1583]" italics="true" pageId="19" pageNumber="20">leg.</emphasis>
/ DZ 20. 673
</materialsCitation>
/
<typeStatus id="9C40598F9D49CA5EFC2BF996FBD6F9B8" box="[960,1077,1557,1583]" pageId="19" pageNumber="20" type="holotype">Holotypus</typeStatus>
<taxonomicName id="84FB9CAE9D49CA5EFBD5F996FF05F9C4" authority="Medeiros &amp; O. Mielke" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="bamba">
<emphasis id="718F3B3F9D49CA5EFBD5F996FB06F9B8" box="[1086,1253,1557,1583]" italics="true" pageId="19" pageNumber="20">Artines bamba</emphasis>
Medeiros &amp; O. Mielke
</taxonomicName>
det. 2018 /.
</paragraph>
<paragraph id="4344E72D9D49CA5EFF2CF9DEFC5EF8BC" blockId="19.[151,1437,152,2015]" pageId="19" pageNumber="20">
<materialsCitation id="F393ED709D49CA5EFF2CF9DEFBF2F90C" ID-GBIF-Occurrence="2248478806" collectionCode="OM" collectorName="Olivenca" elevation="340" location="Parque Nacional" pageId="19" pageNumber="20" specimenCode="OM 9.115" specimenCount="2" specimenCount-male="2" stateProvince="Amazonas" typeStatus="paratype">
<typeStatus id="9C40598F9D49CA5EFF2CF9DEFEDBF9E0" box="[199,312,1629,1655]" pageId="19" pageNumber="20" type="paratype">Paratypes</typeStatus>
: BRAZIL—
<emphasis id="718F3B3F9D49CA5EFE23F9DEFDD8F9E0" box="[456,571,1629,1655]" italics="true" pageId="19" pageNumber="20">
<collectingRegion id="813F29CF9D49CA5EFE23F9DEFDD8F9E0" box="[456,571,1629,1655]" country="Venezuela" name="Amazonas" pageId="19" pageNumber="20">Amazonas</collectingRegion>
</emphasis>
:
<emphasis id="718F3B3F9D49CA5EFDA6F9DFFC80F9E0" bold="true" box="[589,867,1628,1655]" pageId="19" pageNumber="20">
<collectingRegion id="813F29CF9D49CA5EFDA6F9DFFD24F9E0" box="[589,711,1628,1655]" country="Brazil" name="Sao Paulo" pageId="19" pageNumber="20">São Paulo</collectingRegion>
de
<collectorName id="EE0E82FB9D49CA5EFD12F9DFFC80F9E0" box="[761,867,1628,1655]" pageId="19" pageNumber="20">Olivença</collectorName>
</emphasis>
,
<specimenCount id="55FD2CA49D49CA5EFC9FF9DEFC25F9E0" box="[884,966,1629,1655]" pageId="19" pageNumber="20" type="male">1 male</specimenCount>
, no data, no collector,
<specimenCode id="135D4F569D49CA5EFB3EF9DEFAA8F9E0" box="[1237,1355,1628,1655]" collectionCode="OM" country="New Zealand" name="Otago Museum" pageId="19" pageNumber="20">OM 9.115</specimenCode>
(
<collectionCode id="25EA7FE89D49CA5EFAB5F9DEFA72F9E0" box="[1374,1425,1629,1655]" country="New Zealand" name="Otago Museum" pageId="19" pageNumber="20">OM</collectionCode>
). PERU—
<emphasis id="718F3B3F9D49CA5EFF10F902FE44F90C" box="[251,423,1665,1691]" italics="true" pageId="19" pageNumber="20">
<collectingRegion id="813F29CF9D49CA5EFF10F902FE44F90C" box="[251,423,1665,1691]" country="Peru" name="Madre de Dios" pageId="19" pageNumber="20">Madre de Díos</collectingRegion>
</emphasis>
: (
<location id="4624B1F69D49CA5EFE54F902FD9EF90C" LSID="urn:lsid:plazi:treatment:CB52563B9D48CA5EFF7CF926FCD2F848:4624B1F69D49CA5EFE54F902FD9EF90C" box="[447,637,1665,1691]" name="Parque Nacional" pageId="19" pageNumber="20" stateProvince="Amazonas">Parque Nacional</location>
del
<taxonomicName id="84FB9CAE9D49CA5EFD59F902FCB8F90B" authority=", Pakitza" authorityName="Pakitza" box="[690,859,1665,1692]" class="Reptilia" genus="Manu" higherTaxonomySource="CoL" kingdom="Animalia" order="Dinosauria" pageId="19" pageNumber="20" phylum="Chordata" rank="genus">Manu, Pakitza</taxonomicName>
),
<quantity id="84034AC89D49CA5EFC9BF902FC50F90C" box="[880,947,1665,1691]" metricMagnitude="2" metricUnit="m" metricValue="3.4" pageId="19" pageNumber="20" unit="m" value="340.0">
<elevation id="C8D6001E9D49CA5EFC9BF902FC50F90C" box="[880,947,1665,1691]" metricMagnitude="2" metricUnit="m" metricValue="3.4" pageId="19" pageNumber="20" unit="m" value="340.0">340m</elevation>
</quantity>
,
<specimenCount id="55FD2CA49D49CA5EFC2BF902FBF2F90C" box="[960,1041,1665,1691]" pageId="19" pageNumber="20" type="male">1 male</specimenCount>
</materialsCitation>
,
<date id="3745C1ED9D49CA5EFBF4F902FB6AF90C" box="[1055,1161,1665,1691]" pageId="19" pageNumber="20" value="1991-10-02">2.X.1991</date>
,
<specimenCount id="55FD2CA49D49CA5EFB7CF902FB0BF90C" box="[1175,1256,1665,1691]" pageId="19" pageNumber="20" type="male">1 male</specimenCount>
,
<date id="3745C1ED9D49CA5EFB1DF902FA8DF90C" box="[1270,1390,1665,1691]" pageId="19" pageNumber="20" value="1991-10-10">10.X.1991</date>
, M. Casagrande
<emphasis id="718F3B3F9D49CA5EFECBF926FEAAF928" box="[288,329,1701,1727]" italics="true" pageId="19" pageNumber="20">leg.</emphasis>
, DZ 39.901, DZ 39.895,
<specimenCount id="55FD2CA49D49CA5EFD8BF926FD4EF928" box="[608,685,1701,1727]" pageId="19" pageNumber="20" type="male">1 male</specimenCount>
,
<materialsCitation id="F393ED709D49CA5EFD5CF926FAE8F928" ID-GBIF-Occurrence="2248478805" box="[695,1291,1701,1727]" collectingDate="1991-10-20" collectionCode="DZUP" collectorName="O Mielke" pageId="19" pageNumber="20" specimenCount="1" specimenCount-male="1" stateProvince="Amazonas" typeStatus="paratype">
<date id="3745C1ED9D49CA5EFD5CF926FCCCF928" box="[695,815,1701,1727]" pageId="19" pageNumber="20" value="1991-10-20">
<collectingDate id="270138059D49CA5EFD5CF926FCCCF928" box="[695,815,1701,1727]" pageId="19" pageNumber="20" value="1991-10-20">20.X.1991</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D49CA5EFCD1F926FC47F928" box="[826,932,1701,1727]" pageId="19" pageNumber="20">O Mielke</collectorName>
<emphasis id="718F3B3F9D49CA5EFC41F926FC2FF928" box="[938,972,1701,1727]" italics="true" pageId="19" pageNumber="20">leg</emphasis>
., DZ 9.728 (|
<collectionCode id="25EA7FE89D49CA5EFBB1F926FB4FF928" box="[1114,1196,1701,1727]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="19" pageNumber="20" type="University or college">DZUP</collectionCode>
),
<specimenCount id="55FD2CA49D49CA5EFB55F926FAE8F928" box="[1214,1291,1701,1727]" pageId="19" pageNumber="20" type="male">1 male</specimenCount>
</materialsCitation>
,
<materialsCitation id="F393ED709D49CA5EFAFDF926FCB7F974" ID-GBIF-Occurrence="2248478797" collectingDate="1991-09-29" collectionCode="MUSM" collectorName="R. Robbins" elevation="200" location="Juanjui" pageId="19" pageNumber="20" specimenCount="1" specimenCount-male="1" stateProvince="San Martin" typeStatus="paratype">
<date id="3745C1ED9D49CA5EFAFDF926FA74F928" box="[1302,1431,1701,1727]" pageId="19" pageNumber="20" value="1991-09-29">
<collectingDate id="270138059D49CA5EFAFDF926FA74F928" box="[1302,1431,1701,1727]" pageId="19" pageNumber="20" value="1991-09-29">29.IX.1991</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D49CA5EFF7CF94AFEF7F974" box="[151,276,1737,1763]" pageId="19" pageNumber="20">R. Robbins</collectorName>
<emphasis id="718F3B3F9D49CA5EFEF1F94AFEA7F974" box="[282,324,1737,1763]" italics="true" pageId="19" pageNumber="20">leg.</emphasis>
, (
<collectionCode id="25EA7FE89D49CA5EFEB3F94AFE57F974" box="[344,436,1737,1763]" pageId="19" pageNumber="20">MUSM</collectionCode>
).
<emphasis id="718F3B3F9D49CA5EFE2DF94AFDA7F974" box="[454,580,1737,1763]" italics="true" pageId="19" pageNumber="20">
<collectingRegion id="813F29CF9D49CA5EFE2DF94AFDA7F974" box="[454,580,1737,1763]" country="Peru" name="San Martin" pageId="19" pageNumber="20">San Martin</collectingRegion>
</emphasis>
:
<emphasis id="718F3B3F9D49CA5EFDBAF94BFD4EF974" bold="true" box="[593,685,1736,1763]" pageId="19" pageNumber="20">
<location id="4624B1F69D49CA5EFDBAF94BFD4EF974" LSID="urn:lsid:plazi:treatment:CB52563B9D48CA5EFF7CF926FCD2F848:4624B1F69D49CA5EFDBAF94BFD4EF974" box="[593,685,1736,1763]" name="Juanjui" pageId="19" pageNumber="20" stateProvince="San Martin">Juanjui</location>
</emphasis>
,
<quantity id="84034AC89D49CA5EFD51F94AFD1EF974" box="[698,765,1737,1763]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="20" unit="m" value="200.0">
<elevation id="C8D6001E9D49CA5EFD51F94AFD1EF974" box="[698,765,1737,1763]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="20" unit="m" value="200.0">200m</elevation>
</quantity>
,
<specimenCount id="55FD2CA49D49CA5EFCECF94AFCB7F974" box="[775,852,1737,1763]" pageId="19" pageNumber="20" type="male">1 male</specimenCount>
</materialsCitation>
,
<materialsCitation id="F393ED709D49CA5EFCB5F94BFE5AF890" ID-GBIF-Occurrence="2248478878" collectingDate="2002-09-15" collectionCode="OM" location="Calzada" municipality="Moyobamba" pageId="19" pageNumber="20" specimenCode="OM 60.174" specimenCount="1" specimenCount-male="1" stateProvince="San Martin" typeStatus="paratype">
<date id="3745C1ED9D49CA5EFCB5F94BFC03F974" box="[862,992,1736,1763]" pageId="19" pageNumber="20" value="2002-09-15">
<collectingDate id="270138059D49CA5EFCB5F94BFC03F974" box="[862,992,1736,1763]" pageId="19" pageNumber="20" value="2002-09-15">15.IX.2002</collectingDate>
</date>
, no collector,
<specimenCode id="135D4F569D49CA5EFB90F94AFB18F974" box="[1147,1275,1737,1763]" collectionCode="OM" country="New Zealand" name="Otago Museum" pageId="19" pageNumber="20">OM 60.174</specimenCode>
(
<collectionCode id="25EA7FE89D49CA5EFAE2F94AFAD8F974" box="[1289,1339,1737,1763]" country="New Zealand" name="Otago Museum" pageId="19" pageNumber="20">OM</collectionCode>
);
<emphasis id="718F3B3F9D49CA5EFAA5F94AFF0AF890" bold="true" pageId="19" pageNumber="20">
<collectingMunicipality id="A3207D579D49CA5EFAA5F94AFF0AF890" pageId="19" pageNumber="20">Moyobamba</collectingMunicipality>
</emphasis>
(
<location id="4624B1F69D49CA5EFF13F96EFEBBF89F" LSID="urn:lsid:plazi:treatment:CB52563B9D48CA5EFF7CF926FCD2F848:4624B1F69D49CA5EFF13F96EFEBBF89F" box="[248,344,1773,1800]" municipality="Moyobamba" name="Calzada" pageId="19" pageNumber="20" stateProvince="San Martin">Calzada</location>
),
<specimenCount id="55FD2CA49D49CA5EFE80F96EFE5AF890" box="[363,441,1773,1799]" pageId="19" pageNumber="20" type="male">1 male</specimenCount>
</materialsCitation>
,
<materialsCitation id="F393ED709D49CA5EFE2FF96EFC68F890" ID-GBIF-Occurrence="2248478883" box="[452,907,1772,1799]" collectingDate="2015-10" collectionCode="OM" collectorName="J. Pintado" pageId="19" pageNumber="20" specimenCode="OM 79.553, 850" specimenCount="1" stateProvince="San Martin" typeStatus="paratype">
<date id="3745C1ED9D49CA5EFE2FF96EFDF9F890" box="[452,538,1772,1799]" pageId="19" pageNumber="20" value="2015-10">
<collectingDate id="270138059D49CA5EFE2FF96EFDF9F890" box="[452,538,1772,1799]" pageId="19" pageNumber="20" value="2015-10">X.2015</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D49CA5EFDCEF96EFD77F890" box="[549,660,1773,1799]" pageId="19" pageNumber="20">J. Pintado</collectorName>
<emphasis id="718F3B3F9D49CA5EFD71F96EFD5FF890" box="[666,700,1773,1799]" italics="true" pageId="19" pageNumber="20">leg</emphasis>
.,
<specimenCode id="135D4F569D49CA5EFD3AF96EFCB6F890" box="[721,853,1772,1799]" collectionCode="OM" country="New Zealand" name="Otago Museum" pageId="19" pageNumber="20">OM 79.553</specimenCode>
,
<specimenCode id="135D4F569D49CA5EFC8BF96FFC68F890" box="[864,907,1772,1799]" collectionCode="OM" country="New Zealand" name="Otago Museum" pageId="19" pageNumber="20">850</specimenCode>
</materialsCitation>
<materialsCitation id="F393ED709D49CA5EFC67F96FFA7BF890" ID-GBIF-Occurrence="2248478801" box="[908,1432,1772,1799]" collectingDate="2016-10" collectionCode="OM" collectorName="J. Pintado" pageId="19" pageNumber="20" specimenCode="OM 83.226" specimenCount="1" specimenCount-male="1" stateProvince="San Martin" typeStatus="paratype">
m,
<specimenCount id="55FD2CA49D49CA5EFC45F96EFC1FF890" box="[942,1020,1773,1799]" pageId="19" pageNumber="20" type="male">1 male</specimenCount>
,
<date id="3745C1ED9D49CA5EFBECF96EFBBEF890" box="[1031,1117,1773,1799]" pageId="19" pageNumber="20" value="2016-10">
<collectingDate id="270138059D49CA5EFBECF96EFBBEF890" box="[1031,1117,1773,1799]" pageId="19" pageNumber="20" value="2016-10">X.2016</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D49CA5EFB83F96EFB34F890" box="[1128,1239,1773,1799]" pageId="19" pageNumber="20">J. Pintado</collectorName>
<emphasis id="718F3B3F9D49CA5EFB36F96EFB1CF890" box="[1245,1279,1773,1799]" italics="true" pageId="19" pageNumber="20">leg</emphasis>
.,
<specimenCode id="135D4F569D49CA5EFAFFF96EFA7BF890" box="[1300,1432,1773,1799]" collectionCode="OM" country="New Zealand" name="Otago Museum" pageId="19" pageNumber="20">OM 83.226</specimenCode>
</materialsCitation>
,
<quantity id="84034AC89D49CA5EFF7CF892FF39F8BC" box="[151,218,1809,1835]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="19" pageNumber="20" unit="m" value="800.0">800m</quantity>
,
<specimenCount id="55FD2CA49D49CA5EFF0EF892FEDDF8BC" box="[229,318,1809,1835]" pageId="19" pageNumber="20" type="male">2 males</specimenCount>
,
<materialsCitation id="F393ED709D49CA5EFEA2F892FC5BF8BC" ID-GBIF-Occurrence="2248478807" box="[329,952,1809,1835]" collectingDate="2016-10" collectionCode="OM" collectorName="J. Pintado" pageId="19" pageNumber="20" specimenCode="OM 81.199, OM 83.166" specimenCount="1" stateProvince="San Martin" typeStatus="paratype">
<date id="3745C1ED9D49CA5EFEA2F892FE7CF8BC" box="[329,415,1809,1835]" pageId="19" pageNumber="20" value="2016-10">
<collectingDate id="270138059D49CA5EFEA2F892FE7CF8BC" box="[329,415,1809,1835]" pageId="19" pageNumber="20" value="2016-10">X.2016</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D49CA5EFE40F892FDFAF8BC" box="[427,537,1809,1835]" pageId="19" pageNumber="20">J. Pintado</collectorName>
<emphasis id="718F3B3F9D49CA5EFDCBF892FDA1F8BC" box="[544,578,1809,1835]" italics="true" pageId="19" pageNumber="20">leg</emphasis>
.,
<specimenCode id="135D4F569D49CA5EFDBCF892FD38F8BC" box="[599,731,1809,1835]" collectionCode="OM" country="New Zealand" name="Otago Museum" pageId="19" pageNumber="20">OM 81.199</specimenCode>
,
<specimenCode id="135D4F569D49CA5EFD0DF892FC89F8BC" box="[742,874,1809,1835]" collectionCode="OM" country="New Zealand" name="Otago Museum" pageId="19" pageNumber="20">OM 83.166</specimenCode>
, (
<collectionCode id="25EA7FE89D49CA5EFC95F892FC52F8BC" box="[894,945,1809,1835]" country="New Zealand" name="Otago Museum" pageId="19" pageNumber="20">OM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="4344E72D9D49CA5EFF2CF8B7FCD2F848" blockId="19.[151,1437,152,2015]" pageId="19" pageNumber="20">
<emphasis id="718F3B3F9D49CA5EFF2CF8B7FE52F8D8" bold="true" box="[199,433,1844,1871]" pageId="19" pageNumber="20">Additional material</emphasis>
(all females). BRAZIL—
<emphasis id="718F3B3F9D49CA5EFD3BF8B6FCA0F8D8" box="[720,835,1845,1871]" italics="true" pageId="19" pageNumber="20">
<collectingRegion id="813F29CF9D49CA5EFD3BF8B6FCA0F8D8" box="[720,835,1845,1871]" country="Venezuela" name="Amazonas" pageId="19" pageNumber="20">Amazonas</collectingRegion>
</emphasis>
:
<emphasis id="718F3B3F9D49CA5EFCBAF8B7FBBBF8D8" bold="true" box="[849,1112,1844,1871]" pageId="19" pageNumber="20">
<collectingRegion id="813F29CF9D49CA5EFCBAF8B7FC25F8D8" box="[849,966,1844,1871]" country="Brazil" name="Sao Paulo" pageId="19" pageNumber="20">São Paulo</collectingRegion>
de Olivença
</emphasis>
,
<specimenCount id="55FD2CA49D49CA5EFB8EF8B6FB24F8D8" box="[1125,1223,1845,1871]" pageId="19" pageNumber="20" type="female">1 female</specimenCount>
, no data, no collector,
<collectionCode id="25EA7FE89D49CA5EFF28F8DAFF13F8E4" box="[195,240,1881,1907]" country="New Zealand" name="Otago Museum" pageId="19" pageNumber="20">OM</collectionCode>
9.116 (
<collectionCode id="25EA7FE89D49CA5EFEAFF8DAFE94F8E4" box="[324,375,1881,1907]" country="New Zealand" name="Otago Museum" pageId="19" pageNumber="20">OM</collectionCode>
).
<emphasis id="718F3B3F9D49CA5EFE61F8DAFE5DF8E4" box="[394,446,1881,1907]" italics="true" pageId="19" pageNumber="20">
<collectingRegion id="813F29CF9D49CA5EFE61F8DAFE5DF8E4" box="[394,446,1881,1907]" country="Brazil" name="Acre" pageId="19" pageNumber="20">Acre</collectingRegion>
</emphasis>
:
<emphasis id="718F3B3F9D49CA5EFE27F8DBFD51F8E4" bold="true" box="[460,690,1880,1907]" pageId="19" pageNumber="20">Senador Guiomard</emphasis>
(Reserva Catuaba),
<specimenCount id="55FD2CA49D49CA5EFC7CF8DAFC18F8E4" box="[919,1019,1881,1907]" pageId="19" pageNumber="20" type="female">1 female</specimenCount>
,
<date id="3745C1ED9D49CA5EFBEDF8DAFB5FF8E4" box="[1030,1212,1881,1907]" pageId="19" pageNumber="20" value="2008-07-23" valueMax="2008-07-27" valueMin="2008-07-23">23–27.VII.2008</date>
, O. Mielke &amp; Casagrande
<emphasis id="718F3B3F9D49CA5EFF01F8FEFEF7F800" box="[234,276,1917,1943]" italics="true" pageId="19" pageNumber="20">leg.</emphasis>
, DZ 43. 327 (
<collectionCode id="25EA7FE89D49CA5EFE5AF8FEFDE3F800" box="[433,512,1917,1943]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="19" pageNumber="20" type="University or college">DZUP</collectionCode>
);
<emphasis id="718F3B3F9D49CA5EFDFFF8FEFD76F800" bold="true" box="[532,661,1916,1943]" pageId="19" pageNumber="20">
Porto
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(Humaitá),
<specimenCount id="55FD2CA49D49CA5EFCF6F8FEFC62F800" box="[797,897,1917,1943]" pageId="19" pageNumber="20" type="female">1 female</specimenCount>
,
<date id="3745C1ED9D49CA5EFC67F8FEFBA1F800" box="[908,1090,1917,1943]" pageId="19" pageNumber="20" value="2008-07-28" valueMax="2008-07-31" valueMin="2008-07-28">28–31.VII.2008</date>
, O. Mielke &amp; Casagrande
<emphasis id="718F3B3F9D49CA5EFA87F8FEFA76F800" box="[1388,1429,1917,1943]" italics="true" pageId="19" pageNumber="20">leg.</emphasis>
, DZ 43.347 (
<collectionCode id="25EA7FE89D49CA5EFECAF822FE92F82C" box="[289,369,1953,1979]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="19" pageNumber="20" type="University or college">DZUP</collectionCode>
). PERU—
<emphasis id="718F3B3F9D49CA5EFE0CF822FD84F82C" box="[487,615,1953,1979]" italics="true" pageId="19" pageNumber="20">
<collectingRegion id="813F29CF9D49CA5EFE0CF822FD84F82C" box="[487,615,1953,1979]" country="Peru" name="San Martin" pageId="19" pageNumber="20">San Martin</collectingRegion>
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:
<emphasis id="718F3B3F9D49CA5EFD9CF822FCEDF82C" bold="true" box="[631,782,1953,1979]" pageId="19" pageNumber="20">Moyobamba</emphasis>
(Calzada),
<specimenCount id="55FD2CA49D49CA5EFC7AF822FC15F82C" box="[913,1014,1953,1979]" pageId="19" pageNumber="20" type="female">1 female</specimenCount>
,
<date id="3745C1ED9D49CA5EFBE9F822FBBBF82C" box="[1026,1112,1952,1979]" pageId="19" pageNumber="20" value="2015-10">X.2015</date>
, J. Pintado
<emphasis id="718F3B3F9D49CA5EFB30F822FAE6F82C" box="[1243,1285,1953,1979]" italics="true" pageId="19" pageNumber="20">leg.</emphasis>
,
<collectionCode id="25EA7FE89D49CA5EFAF8F822FAA3F82C" box="[1299,1344,1953,1979]" country="New Zealand" name="Otago Museum" pageId="19" pageNumber="20">OM</collectionCode>
79.413,
<quantity id="84034AC89D49CA5EFF7CF846FF39F848" box="[151,218,1989,2015]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="19" pageNumber="20" unit="m" value="800.0">800m</quantity>
,
<materialsCitation id="F393ED709D49CA5EFF0EF846FCCEF848" ID-GBIF-Occurrence="2248478870" box="[229,813,1989,2015]" collectingDate="2016-10" collectionCode="OM" collectorName="J. Pintado" pageId="19" pageNumber="20" specimenCode="OM 81.229" specimenCount="1" specimenCount-female="1">
<specimenCount id="55FD2CA49D49CA5EFF0EF846FEAAF848" box="[229,329,1989,2015]" pageId="19" pageNumber="20" type="female">1 female</specimenCount>
,
<date id="3745C1ED9D49CA5EFEBFF846FE49F848" box="[340,426,1989,2015]" pageId="19" pageNumber="20" value="2016-10">
<collectingDate id="270138059D49CA5EFEBFF846FE49F848" box="[340,426,1989,2015]" pageId="19" pageNumber="20" value="2016-10">X.2016</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D49CA5EFE5EF846FDC7F848" box="[437,548,1989,2015]" pageId="19" pageNumber="20">J. Pintado</collectorName>
<emphasis id="718F3B3F9D49CA5EFDC0F846FDAEF848" box="[555,589,1989,2015]" italics="true" pageId="19" pageNumber="20">leg</emphasis>
.,
<specimenCode id="135D4F569D49CA5EFD89F846FD00F848" box="[610,739,1989,2015]" collectionCode="OM" country="New Zealand" name="Otago Museum" pageId="19" pageNumber="20">OM 81.229</specimenCode>
(
<collectionCode id="25EA7FE89D49CA5EFD19F846FCC6F848" box="[754,805,1989,2015]" country="New Zealand" name="Otago Museum" pageId="19" pageNumber="20">OM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

361
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@ -0,0 +1,361 @@
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<taxonomicName id="84FB9CAE9D4ACA5DFF7CFECEFD60FEF0" authority="Steinhauser &amp; Austin, 1993" authorityName=", S. R. Steinhauser &amp; G. T. Austin" authorityYear="1993" box="[151,643,333,359]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="rica">
<emphasis id="718F3B3F9D4ACA5DFF7CFECEFD60FEF0" bold="true" box="[151,643,333,359]" pageId="16" pageNumber="17">
<emphasis id="718F3B3F9D4ACA5DFF7CFECEFEC5FEF0" bold="true" box="[151,294,333,359]" italics="true" pageId="16" pageNumber="17">Artines rica</emphasis>
<bibRefCitation id="276A9ADC9D4ACA5DFEC6FECEFD60FEF0" author="Steinhauser, S. R. &amp; Austin, G. T." box="[301,643,333,359]" pageId="16" pageNumber="17" pagination="12 - 20" refId="ref45097" refString="Steinhauser, S. R. &amp; Austin, G. T. (1993) New species of Hesperiidae from Costa Rica. Tropical Lepidopter a, 4 (Supplement 2), 12 - 20." type="journal article" year="1993">Steinhauser &amp; Austin, 1993</bibRefCitation>
</emphasis>
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<subSubSection id="0BE1B4A69D4ACA5DFF7CFEF2FE58FE1C" box="[151,443,368,395]" pageId="16" pageNumber="17" type="description">
<paragraph id="4344E72D9D4ACA5DFF7CFEF2FE58FE1C" blockId="16.[151,643,333,395]" box="[151,443,368,395]" pageId="16" pageNumber="17">
<figureCitation id="DBC0FBA89D4ACA5DFF7CFEF2FF08FE1C" box="[151,235,369,395]" captionStart="FIGURES 9–13" captionStartId="7.[151,264,1180,1205]" captionTargetBox="[152,1436,180,1162]" captionTargetId="figure@7.[151,1436,174,1169]" captionTargetPageId="7" captionText="FIGURES 9–13. Species of the “aepitus group” in dorsal and ventral views. 9. Artines maya, A–B: holotype male. 10. Artines litoralis sp. nov., A–B: holotype male, C–D: allotype female. 11. Artines rica, A–B: male, Turrialba, Cartago, Costa Rica (DZ 41.890), C–D: female from same locality of male (DZ 41.840). 12. Artines liege sp. nov., A–B: holotype male, C–D: allotype female. 13. Artines bamba sp. nov., A–B: holotype male, C–D: tentative female, Moyobamba, San Martin, Peru (OM 79.413). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241043/files/figure.png" pageId="16" pageNumber="17">Figs 11</figureCitation>
,
<figureCitation id="DBC0FBA89D4ACA5DFF1CFEF2FEF6FE1C" box="[247,277,369,395]" captionStart="FIGURES 33–35" captionStartId="25.[151,264,1776,1801]" captionTargetBox="[151,1436,187,1743]" captionTargetId="figure@25.[151,1436,181,1750]" captionTargetPageId="25" captionText="FIGURES 33–35. Male genitalia of the “aepitus group” species. 33. Artines rica, Turrialba, Cartago, Costa Rica (DZ 4.061). 34. Artines liege sp. nov., holotype. 35. Artines bamba sp. nov., paratype, Pakitza, Parque Nacional del Manu, Madre de Díos, Peru (DZ 9.728). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241051/files/figure.png" pageId="16" pageNumber="17">33</figureCitation>
,
<figureCitation id="DBC0FBA89D4ACA5DFECAFEF2FEA3FE1C" box="[289,320,369,395]" captionStart="FIGURES 42–48" captionStartId="33.[151,264,1892,1917]" captionTargetBox="[151,1431,625,1874]" captionTargetId="figure@33.[151,1436,625,1880]" captionText="FIGURES 42–48. Dorsal view of vesica and cornutus of the &quot;aepitus group&quot; species. 42. Artines aepitus. 43. Artines panama sp. nov.. 44. Artines delfos sp. nov.. 45. Artines litoralis sp. nov.. 46. Artines rica. 47. Artines liege sp. nov. 48. Artines bamba sp. nov. Scale bar = 0,2 mm." httpUri="https://zenodo.org/record/3406274/files/figure.png" pageId="16" pageNumber="17">46</figureCitation>
, 58, 70, 78
</paragraph>
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<taxonomicName id="84FB9CAE9D4ACA5DFF7CFE35FDD3FE5A" authority="Steinhauser &amp; Austin, 1993" authorityName=", S. R. Steinhauser &amp; G. T. Austin" authorityYear="1993" box="[151,560,437,462]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="rica">
<emphasis id="718F3B3F9D4ACA5DFF7CFE35FEECFE5A" box="[151,271,438,461]" italics="true" pageId="16" pageNumber="17">Artines rica</emphasis>
<bibRefCitation id="276A9ADC9D4ACA5DFEFEFE36FDD3FE5A" author="Steinhauser, S. R. &amp; Austin, G. T." box="[277,560,437,462]" pageId="16" pageNumber="17" pagination="12 - 20" refId="ref45097" refString="Steinhauser, S. R. &amp; Austin, G. T. (1993) New species of Hesperiidae from Costa Rica. Tropical Lepidopter a, 4 (Supplement 2), 12 - 20." type="journal article" year="1993">Steinhauser &amp; Austin, 1993</bibRefCitation>
</taxonomicName>
.
<emphasis id="718F3B3F9D4ACA5DFDD2FE36FD5DFE59" bold="true" box="[569,702,437,462]" pageId="16" pageNumber="17">Trop. Lep. 4</emphasis>
, Suppl. 2: 14, figs 9, 10 (male d, v), 11, 12 (female d, v), 19 (male gen.), 20 (female gen.);
<typeStatus id="9C40598F9D4ACA5DFE97FE50FE36FE7B" box="[380,469,467,492]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
male,
<date id="3745C1ED9D4ACA5DFDF1FE57FD64FE7B" box="[538,647,468,492]" pageId="16" pageNumber="17" value="1972-06-01">1-VI-1972</date>
, Turrialba, Província
<collectingRegion id="813F29CF9D4ACA5DFC8FFE57FC54FE7B" box="[868,951,468,492]" country="Costa Rica" name="Cartago" pageId="16" pageNumber="17">Cartago</collectingRegion>
,
<collectingCountry id="3BECA7BD9D4ACA5DFC2AFE57FBD1FE7B" box="[961,1074,467,492]" name="Costa Rica" pageId="16" pageNumber="17">Costa Rica</collectingCountry>
, H. L. King
<emphasis id="718F3B3F9D4ACA5DFB59FE57FB32FE7B" box="[1202,1233,468,492]" italics="true" pageId="16" pageNumber="17">leg</emphasis>
.; AME [MGCL].—
<bibRefCitation id="276A9ADC9D4ACA5DFF2CFE71FEB8FD9C" author="Bridges, C. A." box="[199,347,498,523]" pageId="16" pageNumber="17" refId="ref38783" refString="Bridges, C. A. (1994) Catalogue of the family-group, genus-group and species-group names of the Hesperioidea (Lepidoptera) of the world. Vols. 1 - 13 &amp; Apps. 1 - 3. Author, Urbana, 3. [11] + 598 pp. [11 pp., Vol. 1: 1 pp., Vol. 2: 1 pp., Vol. 3: 1 pp., Vol. 4: 36 pp., Vol. 5: 9 pp., Vol. 6: 20 pp., Vol. 7: 6 pp., Vol. 8: 246 pp., Vol. 9: 76 pp., Vol. 10: 90 pp., Vol. 11: 71 pp., Vol. 12: 28 pp., Vol. 13: 9 pp., App. 1: 1 pp., App. 2: 2 pp., App. 3: 1 pp.]" type="book" year="1994">Bridges, 1994</bibRefCitation>
.
<emphasis id="718F3B3F9D4ACA5DFE8DFE71FB88FD9C" bold="true" box="[358,1131,498,523]" pageId="16" pageNumber="17">Cat. Fam.-Group, Gen.-Group, Sp. Group Nam. Hesp. (Lep.) World 8</emphasis>
, p. 194;
<emphasis id="718F3B3F9D4ACA5DFB21FE71FB34FD9C" bold="true" box="[1226,1239,498,523]" pageId="16" pageNumber="17">9</emphasis>
, p. 6.—O.
<bibRefCitation id="276A9ADC9D4ACA5DFAA5FE71FF1EFDBE" author="Mielke, O. H. H." pageId="16" pageNumber="17" pagination="3 - 11" refId="ref43459" refString="Mielke, O. H. H. (2004) Hesperiidae. In: Lamas, G. (Ed.), Checklist: Part 4 A, Hesperioidea-Papilionoidea. In: Heppner, J. (Ed.), Atlas of Neotropical Lepidoptera 5 A. Scientific Publishers, Association for Tropical Lepidoptera, Gainesville, pp. 3 - 11 + 25 - 86." type="book chapter" year="2004">Mielke, 2004</bibRefCitation>
. Hesperioidea, p. 62,
<emphasis id="718F3B3F9D4ACA5DFE0FFD92FE1AFDBE" bold="true" box="[484,505,529,553]" italics="true" pageId="16" pageNumber="17">in</emphasis>
Lamas (ed.).
<emphasis id="718F3B3F9D4ACA5DFD65FD92FB75FDBD" bold="true" box="[654,1174,529,554]" pageId="16" pageNumber="17">Checklist: Part 4A, Hesperioidea-Papilionoidea</emphasis>
,
<emphasis id="718F3B3F9D4ACA5DFB4DFD92FB58FDBE" bold="true" box="[1190,1211,529,553]" italics="true" pageId="16" pageNumber="17">in</emphasis>
Heppner (ed.).
<emphasis id="718F3B3F9D4ACA5DFA8FFD92FE60FDDF" bold="true" pageId="16" pageNumber="17">
<taxonomicName id="84FB9CAE9D4ACA5DFA8FFD92FEBEFDDF" authority="Neotrop. Lep." authorityName="Neotrop. Lep." class="Insecta" genus="Atlas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">Atlas Neotrop. Lep.</taxonomicName>
5A
</emphasis>
.—O.
<bibRefCitation id="276A9ADC9D4ACA5DFE29FDACFDAEFDDF" author="Mielke, O. H. H." box="[450,589,559,584]" pageId="16" pageNumber="17" pagination="775 - 1055" refId="ref43529" refString="Mielke, O. H. H. (2005) Catalogue of the American Hesperioidea: Hesperiidae (Lepidoptera) 4. Hesperiinae 1. Adlerodea- Lychnuchus. Sociedade Brasileira de Zoologia, Curitiba. pp. 775 - 1055." type="book chapter" year="2005">Mielke, 2005</bibRefCitation>
.
<emphasis id="718F3B3F9D4ACA5DFDBDFDACFC97FDDF" bold="true" box="[598,884,559,584]" pageId="16" pageNumber="17">Cat. Amer. Hesperioidea 4</emphasis>
, p. 833.—
<bibRefCitation id="276A9ADC9D4ACA5DFC36FDACFB15FDDF" author="Garwood, K. &amp; Lehman, R." box="[989,1270,559,584]" pageId="16" pageNumber="17" refId="ref40900" refString="Garwood, K. &amp; Lehman, R. (2013) Butterflies of Central America. A photographic checklist of common species 3: Hesperiidae, The skippers. RiCale Publishing, Mc Allen, xii + 288 pp." type="book" year="2013">Garwood &amp; Lehman, 2013</bibRefCitation>
.
<emphasis id="718F3B3F9D4ACA5DFB14FDB3FEF4FDF0" bold="true" pageId="16" pageNumber="17">Butt. C. Amer. 3, Hesp</emphasis>
., p. 228, fig. (v).—Carneiro; O. Mielke &amp; Casagrande, 2015.
<emphasis id="718F3B3F9D4ACA5DFC62FDCCFBF8FDF0" bold="true" box="[905,1051,590,615]" pageId="16" pageNumber="17">Zootaxa 3931</emphasis>
(2): 198.—Dolibaina; A. Warren; Carneiro &amp; O. Mielke, 2015.
<emphasis id="718F3B3F9D4ACA5DFE25FDEEFD77FD11" bold="true" box="[462,660,621,646]" pageId="16" pageNumber="17">Trop. Lep. Res. 25</emphasis>
(2): 53, figs 3-4 (male d, v).
</paragraph>
<paragraph id="4344E72D9D4ACA5DFF7CFD30FF18FC81" blockId="16.[151,1437,691,1618]" pageId="16" pageNumber="17">
<emphasis id="718F3B3F9D4ACA5DFF7CFD30FE98FD59" bold="true" box="[151,379,691,718]" pageId="16" pageNumber="17">Taxonomic history.</emphasis>
<taxonomicName id="84FB9CAE9D4ACA5DFE69FD37FDE4FD5A" authorityName=", S. R. Steinhauser &amp; G. T. Austin" authorityYear="1993" box="[386,519,692,718]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="rica">
<emphasis id="718F3B3F9D4ACA5DFE69FD37FDE4FD5A" box="[386,519,692,718]" italics="true" pageId="16" pageNumber="17">Artines rica</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="276A9ADC9D4ACA5DFD3CFD37FBFBFD59" author="Steinhauser, S. R. &amp; Austin, G. T." box="[727,1048,692,718]" pageId="16" pageNumber="17" pagination="12 - 20" refId="ref45097" refString="Steinhauser, S. R. &amp; Austin, G. T. (1993) New species of Hesperiidae from Costa Rica. Tropical Lepidopter a, 4 (Supplement 2), 12 - 20." type="journal article" year="1993">Steinhauser &amp; Austin (1993)</bibRefCitation>
based in
<specimenCount id="55FD2CA49D4ACA5DFB68FD37FAF8FD59" box="[1155,1307,692,718]" pageId="16" pageNumber="17" type="generic">12 specimens</specimenCount>
(males and females) from Turrialba,
<collectingRegion id="813F29CF9D4ACA5DFE5EFD5BFDF3FD65" box="[437,528,728,754]" country="Costa Rica" name="Cartago" pageId="16" pageNumber="17">Cartago</collectingRegion>
,
<collectingCountry id="3BECA7BD9D4ACA5DFDF6FD5BFD7FFD65" box="[541,668,728,754]" name="Costa Rica" pageId="16" pageNumber="17">Costa Rica</collectingCountry>
. The other authors mentioned this species in taxonomic studies and catalogs.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D4ACA5DFF2CFC9CFDD1FC5D" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="4344E72D9D4ACA5DFF2CFC9CFDD1FC5D" blockId="16.[151,1437,691,1618]" pageId="16" pageNumber="17">
<emphasis id="718F3B3F9D4ACA5DFF2CFC9CFEA1FCAD" bold="true" box="[199,322,799,826]" pageId="16" pageNumber="17">Diagnosis.</emphasis>
<taxonomicName id="84FB9CAE9D4ACA5DFEA1FCA3FE33FCAE" authorityName=", S. R. Steinhauser &amp; G. T. Austin" authorityYear="1993" box="[330,464,800,826]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="rica">
<emphasis id="718F3B3F9D4ACA5DFEA1FCA3FE33FCAE" box="[330,464,800,826]" italics="true" pageId="16" pageNumber="17">Artines rica</emphasis>
</taxonomicName>
differs from the other species of the “
<taxonomicName id="84FB9CAE9D4ACA5DFC6FFCA3FC36FCAE" baseAuthorityName="Geyer" baseAuthorityYear="1832" box="[900,981,800,825]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D4ACA5DFC6FFCA3FC36FCAE" box="[900,981,800,825]" italics="true" pageId="16" pageNumber="17">aepitus</emphasis>
</taxonomicName>
group” in the following combination of characters: harpe with ventral process narrow and cylindrical, shorter than tegumen+uncus, with setae on apical portion; cornutus with thin and straight spines, directed posteriorly; proximo-ventral folds of lamella antevaginalis similar to
<taxonomicName id="84FB9CAE9D4ACA5DFEE3FC0FFE94FC32" baseAuthorityName="Geyer" baseAuthorityYear="1832" box="[264,375,908,934]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D4ACA5DFEE3FC0FFE94FC32" box="[264,375,908,934]" italics="true" pageId="16" pageNumber="17">A. aepitus</emphasis>
</taxonomicName>
, but less developed, distal projections of lamella postvaginalis separated by deeper cleft; distribution restricted to Central America.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D4ACA5DFF2CFC50FE5FFB85" pageId="16" pageNumber="17" type="description">
<paragraph id="4344E72D9D4ACA5DFF2CFC50FE5FFB85" blockId="16.[151,1437,691,1618]" pageId="16" pageNumber="17">
<emphasis id="718F3B3F9D4ACA5DFF2CFC50FEDBFC79" bold="true" box="[199,312,979,1006]" pageId="16" pageNumber="17">Variation</emphasis>
. According to
<bibRefCitation id="276A9ADC9D4ACA5DFE35FC57FCFEFC79" author="Steinhauser, S. R. &amp; Austin, G. T." box="[478,797,980,1006]" pageId="16" pageNumber="17" pagination="12 - 20" refId="ref45097" refString="Steinhauser, S. R. &amp; Austin, G. T. (1993) New species of Hesperiidae from Costa Rica. Tropical Lepidopter a, 4 (Supplement 2), 12 - 20." type="journal article" year="1993">Steinhauser &amp; Austin (1993)</bibRefCitation>
there is variation in the presence or absence of discal and apical marks on forewing.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D4ACA5DFF2CFB98FB4FFB7D" pageId="16" pageNumber="17" type="discussion">
<paragraph id="4344E72D9D4ACA5DFF2CFB98FAFCFB35" blockId="16.[151,1437,691,1618]" pageId="16" pageNumber="17">
<emphasis id="718F3B3F9D4ACA5DFF2CFB98FEA5FBA1" bold="true" box="[199,326,1051,1078]" pageId="16" pageNumber="17">Comments</emphasis>
.
<bibRefCitation id="276A9ADC9D4ACA5DFEBFFB9FFD6CFBA1" author="Steinhauser, S. R. &amp; Austin, G. T." box="[340,655,1052,1078]" pageId="16" pageNumber="17" pagination="12 - 20" refId="ref45097" refString="Steinhauser, S. R. &amp; Austin, G. T. (1993) New species of Hesperiidae from Costa Rica. Tropical Lepidopter a, 4 (Supplement 2), 12 - 20." type="journal article" year="1993">Steinhauser &amp; Austin (1993)</bibRefCitation>
described
<taxonomicName id="84FB9CAE9D4ACA5DFCECFB9FFC6AFBA2" authorityName=", S. R. Steinhauser &amp; G. T. Austin" authorityYear="1993" box="[775,905,1052,1078]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="rica">
<emphasis id="718F3B3F9D4ACA5DFCECFB9FFC6AFBA2" box="[775,905,1052,1078]" italics="true" pageId="16" pageNumber="17">Artines rica</emphasis>
</taxonomicName>
and mentioned some specimens from
<collectingRegion id="813F29CF9D4ACA5DFADAFB9FFA7FFBA1" box="[1329,1436,1052,1078]" country="Brazil" name="Rondonia" pageId="16" pageNumber="17">Rondônia</collectingRegion>
and Distrito Federal,
<collectingCountry id="3BECA7BD9D4ACA5DFE94FBC3FE26FBCD" box="[383,453,1088,1114]" name="Brazil" pageId="16" pageNumber="17">Brazil</collectingCountry>
. They did not designate any of them as
<typeStatus id="9C40598F9D4ACA5DFC9EFBC3FC02FBCD" box="[885,993,1088,1114]" pageId="16" pageNumber="17" type="paratype">paratypes</typeStatus>
. The identification of the specimen from Distrito Federal is dubious because its genitalia, according to the authors, is damaged. The specimens from these localities studied by us belong to
<taxonomicName id="84FB9CAE9D4ACA5DFE23FB0BFDF8FB35" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[456,539,1160,1186]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="liege" status="sp. nov.">
<emphasis id="718F3B3F9D4ACA5DFE23FB0BFDF8FB35" box="[456,539,1160,1186]" italics="true" pageId="16" pageNumber="17">A. liege</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D4ACA5DFDC9FB0BFD9AFB35" bold="true" box="[546,633,1160,1186]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="6ABC86449D4ACA5DFDC9FB0BFD9AFB35" box="[546,633,1160,1186]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Therefore,
<taxonomicName id="84FB9CAE9D4ACA5DFD13FB0BFCA0FB36" authorityName=", S. R. Steinhauser &amp; G. T. Austin" authorityYear="1993" box="[760,835,1160,1186]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="rica">
<emphasis id="718F3B3F9D4ACA5DFD13FB0BFCA0FB36" box="[760,835,1160,1186]" italics="true" pageId="16" pageNumber="17">A. rica</emphasis>
</taxonomicName>
probably does not occur in South America.
</paragraph>
<paragraph id="4344E72D9D4ACA5DFF2CFB28FB4FFB7D" blockId="16.[151,1437,691,1618]" pageId="16" pageNumber="17">
<emphasis id="718F3B3F9D4ACA5DFF2CFB28FE0BFB51" bold="true" box="[199,488,1195,1222]" pageId="16" pageNumber="17">Geographic distribution</emphasis>
(
<figureCitation id="DBC0FBA89D4ACA5DFE1CFB2FFDAFFB51" box="[503,588,1196,1222]" captionStart="FIGURE 78" captionStartId="42.[151,250,910,935]" captionTargetBox="[151,1436,174,898]" captionTargetId="figure@42.[151,1436,174,898]" captionTargetPageId="42" captionText="FIGURE 78. Geographic distribution of the “aepitus group” species. The symbols with white inferior half represent records from literature." httpUri="https://zenodo.org/record/3241067/files/figure.png" pageId="16" pageNumber="17">Fig. 78</figureCitation>
). The species occurs in forested areas of Central America. COSTA RICA—
<emphasis id="718F3B3F9D4ACA5DFF7CFB53FF17FB7D" box="[151,244,1232,1258]" italics="true" pageId="16" pageNumber="17">
<collectingRegion id="813F29CF9D4ACA5DFF7CFB53FF17FB7D" box="[151,244,1232,1258]" country="Costa Rica" name="Cartago" pageId="16" pageNumber="17">Cartago</collectingRegion>
</emphasis>
: Turrialba.
<emphasis id="718F3B3F9D4ACA5DFE93FB53FE1DFB7D" box="[376,510,1232,1258]" italics="true" pageId="16" pageNumber="17">
<collectingRegion id="813F29CF9D4ACA5DFE93FB53FE1DFB7D" box="[376,510,1232,1258]" country="Costa Rica" name="Guanacaste" pageId="16" pageNumber="17">Guanacaste</collectingRegion>
</emphasis>
:
<collectingRegion id="813F29CF9D4ACA5DFDE6FB53FD6DFB7D" box="[525,654,1232,1258]" country="Costa Rica" name="Guanacaste" pageId="16" pageNumber="17">Guanacaste</collectingRegion>
Conservation Area (
<bibRefCitation id="276A9ADC9D4ACA5DFC9FFB53FB43FB7D" author="Janzen, D. H. &amp; Hallwachs, W." box="[884,1184,1232,1258]" pageId="16" pageNumber="17" refId="ref41792" refString="Janzen, D. H. &amp; Hallwachs, W. (2009) Dynamic database for an inventory of the macrocaterpillar fauna, and its food plants and parasitoids, of Area de Conservacion Guanacaste (ACG), northwestern Costa Rica (nn-SRNP-nnnnn voucher codes). Available from: http: // janzen. sas. upenn. edu / (accessed 15 November 2018)" type="url" year="2009">Janzen &amp; Hallwachs 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D4ACA5DFF2CFB77FC22FA99" box="[199,961,1267,1294]" pageId="16" pageNumber="17" type="description">
<paragraph id="4344E72D9D4ACA5DFF2CFB77FC22FA99" blockId="16.[151,1437,691,1618]" box="[199,961,1267,1294]" pageId="16" pageNumber="17">
<emphasis id="718F3B3F9D4ACA5DFF2CFB77FE33FA99" bold="true" box="[199,464,1267,1294]" pageId="16" pageNumber="17">Temporal distribution</emphasis>
. This species occurs in April, June and July.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D4ACA5DFF2CFA9BFB6BFAA5" box="[199,1160,1304,1330]" pageId="16" pageNumber="17" type="biology_ecology">
<paragraph id="4344E72D9D4ACA5DFF2CFA9BFB6BFAA5" blockId="16.[151,1437,691,1618]" box="[199,1160,1304,1330]" pageId="16" pageNumber="17">
<emphasis id="718F3B3F9D4ACA5DFF2CFA9BFEA7FAA5" bold="true" box="[199,324,1304,1330]" pageId="16" pageNumber="17">Host plant</emphasis>
.
<taxonomicName id="84FB9CAE9D4ACA5DFEB9FA9BFE16FAA5" authorityName="Trin" authorityYear="1836" box="[338,501,1304,1330]" class="Actinopterygii" family="Olyridae" genus="Olyra" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="caudata">
<emphasis id="718F3B3F9D4ACA5DFEB9FA9BFE16FAA5" box="[338,501,1304,1330]" italics="true" pageId="16" pageNumber="17">Olyra caudata</emphasis>
</taxonomicName>
and
<taxonomicName id="84FB9CAE9D4ACA5DFDC0FA9BFCA1FAA5" box="[555,834,1304,1330]" class="Actinopterygii" family="Olyridae" genus="Olyra" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="latifolia">
<emphasis id="718F3B3F9D4ACA5DFDC0FA9BFD2EFAA5" box="[555,717,1304,1330]" italics="true" pageId="16" pageNumber="17">Olyra latifolia</emphasis>
(Poaceae)
</taxonomicName>
(
<bibRefCitation id="276A9ADC9D4ACA5DFCBBFA9BFB9FFAA5" author="Janzen, D. H. &amp; Hallwachs, W." box="[848,1148,1304,1330]" pageId="16" pageNumber="17" refId="ref41792" refString="Janzen, D. H. &amp; Hallwachs, W. (2009) Dynamic database for an inventory of the macrocaterpillar fauna, and its food plants and parasitoids, of Area de Conservacion Guanacaste (ACG), northwestern Costa Rica (nn-SRNP-nnnnn voucher codes). Available from: http: // janzen. sas. upenn. edu / (accessed 15 November 2018)" type="url" year="2009">Janzen &amp; Hallwachs 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D4ACA5DFF2CFABFFA76FAC1" box="[199,1429,1340,1366]" pageId="16" pageNumber="17" type="etymology">
<paragraph id="4344E72D9D4ACA5DFF2CFABFFA76FAC1" blockId="16.[151,1437,691,1618]" box="[199,1429,1340,1366]" pageId="16" pageNumber="17">
<emphasis id="718F3B3F9D4ACA5DFF2CFABFFEA4FAC1" bold="true" box="[199,327,1340,1366]" pageId="16" pageNumber="17">Etymology</emphasis>
. According to
<bibRefCitation id="276A9ADC9D4ACA5DFE00FABFFCCAFAC1" author="Steinhauser, S. R. &amp; Austin, G. T." box="[491,809,1340,1366]" pageId="16" pageNumber="17" pagination="12 - 20" refId="ref45097" refString="Steinhauser, S. R. &amp; Austin, G. T. (1993) New species of Hesperiidae from Costa Rica. Tropical Lepidopter a, 4 (Supplement 2), 12 - 20." type="journal article" year="1993">Steinhauser &amp; Austin (1993)</bibRefCitation>
the term &quot;
<taxonomicName id="84FB9CAE9D4ACA5DFC76FABFFC29FAC2" authorityName=", S. R. Steinhauser &amp; G. T. Austin" authorityYear="1993" box="[925,970,1340,1365]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="rica">
<emphasis id="718F3B3F9D4ACA5DFC76FABFFC29FAC2" box="[925,970,1340,1365]" italics="true" pageId="16" pageNumber="17">rica</emphasis>
</taxonomicName>
&quot; refers to the
<typeStatus id="9C40598F9D4ACA5DFB8CFABFFB74FAC1" box="[1127,1175,1340,1366]" pageId="16" pageNumber="17">type</typeStatus>
locality of the species.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D4ACA5DFF2CFAE3FB97F9C5" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="4344E72D9D4ACA5DFF2CFAE3FDD9FA71" blockId="16.[151,1437,691,1618]" pageId="16" pageNumber="17">
<emphasis id="718F3B3F9D4ACA5DFF2CFAE3FE93FAED" bold="true" box="[199,368,1375,1402]" pageId="16" pageNumber="17">Type material</emphasis>
.
<materialsCitation id="F393ED709D4ACA5DFE6AFAE3FB08FAED" ID-GBIF-Occurrence="2248478880" box="[385,1259,1376,1402]" collectionCode="MGCL, AME" pageId="16" pageNumber="17" specimenCount="1" typeStatus="holotype">
<typeStatus id="9C40598F9D4ACA5DFE6AFAE3FE0AFAED" box="[385,489,1376,1402]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
male deposited at the
<collectionCode id="25EA7FE89D4ACA5DFD1AFAE3FCA2FAED" box="[753,833,1376,1402]" pageId="16" pageNumber="17">MGCL</collectionCode>
(ex
<collectionCode id="25EA7FE89D4ACA5DFC9DFAE3FC55FAED" box="[886,950,1376,1402]" pageId="16" pageNumber="17">AME</collectionCode>
) with the following labels
</materialsCitation>
: /
<materialsCitation id="F393ED709D4ACA5DFAE3FAE3FDCEFA71" ID-GBIF-Occurrence="2248478804" collectingDate="1972-06" collectorName="H. L. King &amp; Database &amp; Allyn Museum Photo &amp; No. &amp; A. C. Allyn Ace. &amp; Specimen" country="Costa Rica" location="Turriable" pageId="16" pageNumber="17" specimenCount="1" stateProvince="Cartago" typeStatus="holotype">
<typeStatus id="9C40598F9D4ACA5DFAE3FAE3FA7FFAED" box="[1288,1436,1376,1402]" pageId="16" pageNumber="17" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="84FB9CAE9D4ACA5DFF7CFA07FD43FA09" authority=", S. R. Steinhauser &amp; G. T. Austin" authorityName=", S. R. Steinhauser &amp; G. T. Austin" authorityYear="1993" box="[151,672,1412,1438]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="rica">
<emphasis id="718F3B3F9D4ACA5DFF7CFA07FEC6FA0A" box="[151,293,1412,1438]" italics="true" pageId="16" pageNumber="17">Artines rica,</emphasis>
S. R. Steinhauser &amp; G. T. Austin
</taxonomicName>
/
<collectingCountry id="3BECA7BD9D4ACA5DFD5CFA07FCBDFA09" box="[695,862,1412,1438]" name="Costa Rica" pageId="16" pageNumber="17">COSTA RICA</collectingCountry>
:
<collectingRegion id="813F29CF9D4ACA5DFC81FA07FC13FA09" box="[874,1008,1412,1438]" country="Costa Rica" name="Cartago" pageId="16" pageNumber="17">CARTAGO</collectingRegion>
,
<location id="4624B1F69D4ACA5DFC17FA07FB80FA09" LSID="urn:lsid:plazi:treatment:CB52563B9D4ACA5DFF7CFECEFB97F9C5:4624B1F69D4ACA5DFC17FA07FB80FA09" box="[1020,1123,1412,1438]" country="Costa Rica" name="Turriable" pageId="16" pageNumber="17" stateProvince="Cartago">Turriable</location>
[sic!] l.
<date id="3745C1ED9D4ACA5DFB51FA07FAF7FA09" box="[1210,1300,1412,1438]" pageId="16" pageNumber="17" value="1972-06">
<collectingDate id="270138059D4ACA5DFB51FA07FAF7FA09" box="[1210,1300,1412,1438]" pageId="16" pageNumber="17" value="1972-06">vi.1972</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D4ACA5DFACAFA07FA7FFA09" box="[1313,1436,1412,1438]" pageId="16" pageNumber="17">H. L. King</collectorName>
/ SRS
<collectorName id="EE0E82FB9D4ACA5DFF09FA2BFEAFFA55" box="[226,332,1448,1474]" pageId="16" pageNumber="17">Database</collectorName>
, Nº 568 /
<collectorName id="EE0E82FB9D4ACA5DFE2BFA2BFD56FA55" box="[448,693,1448,1474]" pageId="16" pageNumber="17">Allyn Museum Photo</collectorName>
/
<collectorName id="EE0E82FB9D4ACA5DFD24FA2BFD1BFA55" box="[719,760,1448,1474]" pageId="16" pageNumber="17">No.</collectorName>
890626-19.20 / 890719-21, 22 /
<collectorName id="EE0E82FB9D4ACA5DFB98FA2BFAD7FA55" box="[1139,1332,1448,1474]" pageId="16" pageNumber="17">A. C. Allyn Ace.</collectorName>
1972-5 / MGCLFLMNH
<collectorName id="EE0E82FB9D4ACA5DFEBAFA4FFE5CFA71" box="[337,447,1484,1510]" pageId="16" pageNumber="17">Specimen</collectorName>
nº 47907
</materialsCitation>
/.
</paragraph>
<paragraph id="4344E72D9D4ACA5DFF2CFA6CFB97F9C5" blockId="16.[151,1437,691,1618]" pageId="16" pageNumber="17">
<emphasis id="718F3B3F9D4ACA5DFF2CFA6CFE4EF99D" bold="true" box="[199,429,1519,1546]" pageId="16" pageNumber="17">Examined material</emphasis>
. Illustration of the
<typeStatus id="9C40598F9D4ACA5DFD6FFA73FD05F99D" box="[644,742,1520,1546]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
and the following specimens: COSTA RICA—
<emphasis id="718F3B3F9D4ACA5DFB17FA73FABAF99D" box="[1276,1369,1520,1546]" italics="true" pageId="16" pageNumber="17">
<collectingRegion id="813F29CF9D4ACA5DFB17FA73FABAF99D" box="[1276,1369,1520,1546]" country="Costa Rica" name="Cartago" pageId="16" pageNumber="17">Cartago</collectingRegion>
</emphasis>
:
<emphasis id="718F3B3F9D4ACA5DFA83FA73FF3CF9B9" bold="true" pageId="16" pageNumber="17">Turrialba</emphasis>
,
<specimenCount id="55FD2CA49D4ACA5DFF07F997FEACF9B9" box="[236,335,1556,1582]" pageId="16" pageNumber="17" type="female">1 female</specimenCount>
,
<date id="3745C1ED9D4ACA5DFEB1F990FE3BF9B9" box="[346,472,1555,1582]" pageId="16" pageNumber="17" value="1971-07-05">5.VII.1971</date>
,
<materialsCitation id="F393ED709D4ACA5DFE09F997FDFCF9C6" ID-GBIF-Occurrence="2248478873" collectingDate="1973-04-15" collectingDateMax="1973-04-30" collectingDateMin="1973-04-15" collectionCode="DZUP" collectorName="Becker" elevation="600" pageId="16" pageNumber="17" specimenCount="4" specimenCount-female="1" specimenCount-male="3">
<specimenCount id="55FD2CA49D4ACA5DFE09F997FDD8F9B9" box="[482,571,1556,1582]" pageId="16" pageNumber="17" type="male">3 males</specimenCount>
,
<specimenCount id="55FD2CA49D4ACA5DFDAEF997FD4AF9B9" box="[581,681,1556,1582]" pageId="16" pageNumber="17" type="female">1 female</specimenCount>
,
<date id="3745C1ED9D4ACA5DFD58F990FCBFF9B9" box="[691,860,1555,1582]" pageId="16" pageNumber="17" value="1973-04-15" valueMax="1973-04-30" valueMin="1973-04-15">
<collectingDate id="270138059D4ACA5DFD58F990FCBFF9B9" box="[691,860,1555,1582]" pageId="16" pageNumber="17" value="1973-04-15" valueMax="1973-04-30" valueMin="1973-04-15">15–30.IV.1973</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D4ACA5DFC8CF997FC55F9B9" box="[871,950,1556,1582]" pageId="16" pageNumber="17">Becker</collectorName>
<emphasis id="718F3B3F9D4ACA5DFC57F997FC06F9B9" box="[956,997,1556,1582]" italics="true" pageId="16" pageNumber="17">leg.</emphasis>
, DZ 31.561, DZ 4.061, DZ 41.890, DZ 30.956, DZ 41.840 (
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),
<quantity id="84034AC89D4ACA5DFE37F9B4FDFCF9C6" box="[476,543,1591,1617]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="16" pageNumber="17" unit="m" value="600.0">
<elevation id="C8D6001E9D4ACA5DFE37F9B4FDFCF9C6" box="[476,543,1591,1617]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="16" pageNumber="17" unit="m" value="600.0">600m</elevation>
</quantity>
</materialsCitation>
,
<materialsCitation id="F393ED709D4ACA5DFDC1F9BBFD25F9C5" ID-GBIF-Occurrence="2248478863" box="[554,710,1592,1618]" collectionCode="VI" pageId="16" pageNumber="17" specimenCount="1" specimenCount-male="1">
<specimenCount id="55FD2CA49D4ACA5DFDC1F9BBFD9BF9C5" box="[554,632,1592,1618]" pageId="16" pageNumber="17" type="male">1 male</specimenCount>
,
<date id="3745C1ED9D4ACA5DFD68F9BBFD25F9C5" box="[643,710,1592,1618]" pageId="16" pageNumber="17" value="1973-06-29">
29.
<collectionCode id="25EA7FE89D4ACA5DFD4DF9BBFD25F9C5" box="[678,710,1592,1618]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="16" pageNumber="17">VI</collectionCode>
</date>
</materialsCitation>
.1973,
<collectorName id="EE0E82FB9D4ACA5DFCFAF9BBFC83F9C5" box="[785,864,1592,1618]" pageId="16" pageNumber="17">Becker</collectorName>
<emphasis id="718F3B3F9D4ACA5DFC8CF9BBFC6AF9C5" box="[871,905,1592,1618]" italics="true" pageId="16" pageNumber="17">leg</emphasis>
., DZ 8.870 (
<collectionCode id="25EA7FE89D4ACA5DFBF2F9BBFB8AF9C5" box="[1049,1129,1592,1618]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="16" pageNumber="17" type="University or college">DZUP</collectionCode>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<paragraph id="4344E72D9D4ECA59FF7CFCC5FCEDFCF7" blockId="20.[151,782,838,865]" box="[151,782,838,865]" pageId="20" pageNumber="21">
<heading id="180C50419D4ECA59FF7CFCC5FCEDFCF7" bold="true" box="[151,782,838,865]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
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Key to males of the species of the “
<taxonomicName id="84FB9CAE9D4ECA59FDA2FCC5FD4DFCF6" ID-CoL="GZYD" baseAuthorityName="Plotz" baseAuthorityYear="1882" box="[585,686,838,865]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="aquilina">
<emphasis id="718F3B3F9D4ECA59FDA2FCC5FD4DFCF6" bold="true" box="[585,686,838,865]" italics="true" pageId="20" pageNumber="21">aquilina</emphasis>
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group”
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<paragraph id="4344E72D9D4ECA59FF7CFC08FA7FFC2A" blockId="20.[151,1437,906,1251]" pageId="20" pageNumber="21">
<keyLead id="F80A44189D4ECA59FF7CFC08FA7FFC2A" pageId="20" pageNumber="21">
1 Ventral hind wing with reduced central black spot and the postdiscal band limited distally by wavy margin (
<figureCitation id="DBC0FBA89D4ECA59FB07FC09FABCFC37" box="[1260,1375,906,928]" captionStart="FIGURES 14–19" captionStartId="10.[151,264,1406,1431]" captionTargetBox="[155,1435,175,1386]" captionTargetId="figure@10.[151,1436,174,1394]" captionTargetPageId="10" captionText="FIGURES 14–19. Species of the “aquilina group” (14–18) and “bipunctata group” (19) in dorsal and ventral views. 14. Artines aquilina, A–B: male, Joinville, Santa Catarina, Brazil (DZ 41.321), C–D: female from same locality of male (DZ 41.220). 15. Artines angelica sp. nov., A–B: holotype male, C–D: allotype female. 16. Artines focus, A–B: male, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.233), C–D: female, Pacaraima, Roraima, Brazil (DZ 41.333). 17. Artines cofus sp. nov., A–B: holotype male, C–D: allotype female. 18. Artines donia sp. nov., A–B: holotype male. 19. Artines bipunctata, A–B: male, Jardim Zoológico, Brasília, Distrito Federal, Brazil (DZ 30.961), C–D: female, Córrego Bananal, Brasília, Distrito Federal, Brazil (DZ 41.224). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241045/files/figure.png" pageId="20" pageNumber="21">Fig. 14B, D</figureCitation>
).................................................................................................
<taxonomicName id="84FB9CAE9D4ECA59FAA7FC2BFA7FFC2A" ID-CoL="GZYD" baseAuthorityName="Plotz" baseAuthorityYear="1882" box="[1356,1436,936,957]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="aquilina">
<emphasis id="718F3B3F9D4ECA59FAA7FC2BFA7FFC2A" box="[1356,1436,936,957]" italics="true" pageId="20" pageNumber="21">aquilina</emphasis>
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- Ventral hind wing with large central black spot and the postdiscal band limited distally by not wavy margin (
<figureCitation id="DBC0FBA89D4ECA59FB06FC46FAA4FC4C" box="[1261,1351,965,987]" captionStart="FIGURES 14–19" captionStartId="10.[151,264,1406,1431]" captionTargetBox="[155,1435,175,1386]" captionTargetId="figure@10.[151,1436,174,1394]" captionTargetPageId="10" captionText="FIGURES 14–19. Species of the “aquilina group” (14–18) and “bipunctata group” (19) in dorsal and ventral views. 14. Artines aquilina, A–B: male, Joinville, Santa Catarina, Brazil (DZ 41.321), C–D: female from same locality of male (DZ 41.220). 15. Artines angelica sp. nov., A–B: holotype male, C–D: allotype female. 16. Artines focus, A–B: male, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.233), C–D: female, Pacaraima, Roraima, Brazil (DZ 41.333). 17. Artines cofus sp. nov., A–B: holotype male, C–D: allotype female. 18. Artines donia sp. nov., A–B: holotype male. 19. Artines bipunctata, A–B: male, Jardim Zoológico, Brasília, Distrito Federal, Brazil (DZ 30.961), C–D: female, Córrego Bananal, Brasília, Distrito Federal, Brazil (DZ 41.224). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241045/files/figure.png" pageId="20" pageNumber="21">Figs 15B</figureCitation>
, D–18B, D)................................................................................................. 2
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<paragraph id="4344E72D9D4ECA59FF7CFB83FA7FFB82" blockId="20.[151,1437,906,1251]" box="[151,1436,1024,1046]" pageId="20" pageNumber="21">
<keyLead id="F80A44189D4ECA59FF7CFB83FA7FFB82" box="[151,1436,1024,1046]" pageId="20" pageNumber="21">
2 Uncus distally pointed (
<figureCitation id="DBC0FBA89D4ECA59FE2CFB83FDFCFB81" box="[455,543,1024,1046]" captionStart="FIGURES 36–37" captionStartId="26.[151,264,1557,1582]" captionTargetBox="[157,1435,179,1543]" captionTargetId="figure@26.[151,1436,174,1546]" captionTargetPageId="26" captionText="FIGURES 36–37. Male genitalia of the “aquilina group” species. 36. Artines aquilina, Caminho da Lagoinha, Cachoeiras de Macacú, Rio de Janeiro, Brazil (DZ 41.196). 37. Artines angelica sp. nov., holotype. M) dorsal view of vesica showing the cornuti. For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241053/files/figure.png" pageId="20" pageNumber="21">Fig. 37C</figureCitation>
); cornutus with conspicuous spines (
<figureCitation id="DBC0FBA89D4ECA59FC6BFB83FC25FB81" box="[896,966,1024,1046]" captionStart="FIGURES 49–53" captionStartId="34.[151,264,895,920]" captionTargetBox="[155,1421,182,878]" captionTargetId="figure@34.[151,1436,174,883]" captionTargetPageId="34" captionText="FIGURES 49–53. Ventral view of vesica and cornutus of the “aquilina group” species. 49. Artines aquilina. 50. Artines angelica sp. nov.. 51. Artines focus. 52. Artines cofus sp. nov.. 53. Artines donia sp. nov.. Scale bar = 0,2 mm." httpUri="https://zenodo.org/record/3241057/files/figure.png" pageId="20" pageNumber="21">Fig. 50</figureCitation>
)........................
<taxonomicName id="84FB9CAE9D4ECA59FB1CFB83FAA9FB82" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[1271,1354,1024,1045]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="angelica" status="sp. nov.">
<emphasis id="718F3B3F9D4ECA59FB1CFB83FAA9FB82" box="[1271,1354,1024,1045]" italics="true" pageId="20" pageNumber="21">angelica</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D4ECA59FABBFB83FA7FFB82" bold="true" box="[1360,1436,1024,1045]" pageId="20" pageNumber="21">
<taxonomicNameLabel id="6ABC86449D4ECA59FABBFB83FA7FFB82" box="[1360,1436,1024,1045]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
<paragraph id="4344E72D9D4ECA59FF7CFB9DFA7FFBA5" blockId="20.[151,1437,906,1251]" box="[151,1436,1053,1075]" pageId="20" pageNumber="21">
<keyLead id="F80A44189D4ECA59FF7CFB9DFA7FFBA5" box="[151,1436,1053,1075]" pageId="20" pageNumber="21">
- Uncus distally rhomboid (
<figureCitation id="DBC0FBA89D4ECA59FE37FB9EFDC6FBA4" box="[476,549,1053,1075]" captionStart="FIGURES 38–39" captionStartId="29.[151,264,1720,1745]" captionTargetBox="[161,1436,431,1681]" captionTargetId="figure@29.[151,1436,450,1693]" captionTargetPageId="29" captionText="FIGURES 38–39. Male genitalia of the “aquilina group” species. 38. Artines focus, Reserva Duque, Manaus, Amazonas, Brazil (DZ 41.192). 39. Artines cofus sp. nov., paratype, Tambopata, Puerto Maldonado Madre de Díos, Peru (MUSM). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241055/files/figure.png" pageId="20" pageNumber="21">Figs 38</figureCitation>
C–40C); cornutus without conspicuous spines (
<figureCitation id="DBC0FBA89D4ECA59FC00FB9EFBB6FBA4" box="[1003,1109,1053,1075]" captionStart="FIGURES 49–53" captionStartId="34.[151,264,895,920]" captionTargetBox="[155,1421,182,878]" captionTargetId="figure@34.[151,1436,174,883]" captionTargetPageId="34" captionText="FIGURES 49–53. Ventral view of vesica and cornutus of the “aquilina group” species. 49. Artines aquilina. 50. Artines angelica sp. nov.. 51. Artines focus. 52. Artines cofus sp. nov.. 53. Artines donia sp. nov.. Scale bar = 0,2 mm." httpUri="https://zenodo.org/record/3241057/files/figure.png" pageId="20" pageNumber="21">Fig. 51–53</figureCitation>
)......................... 3
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<keyLead id="F80A44189D4ECA59FF7CFBB8FA7EFBC7" box="[151,1437,1082,1104]" pageId="20" pageNumber="21">
3 Uncus with a ventral projection (
<figureCitation id="DBC0FBA89D4ECA59FDF5FBB9FD71FBC7" box="[542,658,1082,1104]" captionStart="FIGURES 38–39" captionStartId="29.[151,264,1720,1745]" captionTargetBox="[161,1436,431,1681]" captionTargetId="figure@29.[151,1436,450,1693]" captionTargetPageId="29" captionText="FIGURES 38–39. Male genitalia of the “aquilina group” species. 38. Artines focus, Reserva Duque, Manaus, Amazonas, Brazil (DZ 41.192). 39. Artines cofus sp. nov., paratype, Tambopata, Puerto Maldonado Madre de Díos, Peru (MUSM). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241055/files/figure.png" pageId="20" pageNumber="21">Fig. 38A, D</figureCitation>
)............................................................
<taxonomicName id="84FB9CAE9D4ECA59FA81FBB8FA7EFBC7" ID-CoL="GZYH" authorityName=", Reserva Duque, Manaus" authorityYear="1955" box="[1386,1437,1083,1104]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="focus">
<emphasis id="718F3B3F9D4ECA59FA81FBB8FA7EFBC7" box="[1386,1437,1083,1104]" italics="true" pageId="20" pageNumber="21">focus</emphasis>
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</paragraph>
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<keyLead id="F80A44189D4ECA59FF7CFBDBFA7FFBFA" box="[151,1436,1112,1134]" pageId="20" pageNumber="21">
- Uncus without a ventral projection (
<figureCitation id="DBC0FBA89D4ECA59FDD7FBDBFD7BFBFA" box="[572,664,1112,1134]" captionStart="FIGURES 38–39" captionStartId="29.[151,264,1720,1745]" captionTargetBox="[161,1436,431,1681]" captionTargetId="figure@29.[151,1436,450,1693]" captionTargetPageId="29" captionText="FIGURES 38–39. Male genitalia of the “aquilina group” species. 38. Artines focus, Reserva Duque, Manaus, Amazonas, Brazil (DZ 41.192). 39. Artines cofus sp. nov., paratype, Tambopata, Puerto Maldonado Madre de Díos, Peru (MUSM). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241055/files/figure.png" pageId="20" pageNumber="21">Figs 39A</figureCitation>
, D– 40A, D).................................................... 4
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<keyLead id="F80A44189D4ECA59FF7CFBF6FA7FFB3F" pageId="20" pageNumber="21">
4 Uncus with distal end simple (
<figureCitation id="DBC0FBA89D4ECA59FDE2FBF6FDB3FB1C" box="[521,592,1141,1163]" captionStart="FIGURES 38–39" captionStartId="29.[151,264,1720,1745]" captionTargetBox="[161,1436,431,1681]" captionTargetId="figure@29.[151,1436,450,1693]" captionTargetPageId="29" captionText="FIGURES 38–39. Male genitalia of the “aquilina group” species. 38. Artines focus, Reserva Duque, Manaus, Amazonas, Brazil (DZ 41.192). 39. Artines cofus sp. nov., paratype, Tambopata, Puerto Maldonado Madre de Díos, Peru (MUSM). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241055/files/figure.png" pageId="20" pageNumber="21">Fig. 39</figureCitation>
C–D); valva distally pointed (
<figureCitation id="DBC0FBA89D4ECA59FC9BFBF6FC24FB1C" box="[880,967,1141,1163]" captionStart="FIGURES 38–39" captionStartId="29.[151,264,1720,1745]" captionTargetBox="[161,1436,431,1681]" captionTargetId="figure@29.[151,1436,450,1693]" captionTargetPageId="29" captionText="FIGURES 38–39. Male genitalia of the “aquilina group” species. 38. Artines focus, Reserva Duque, Manaus, Amazonas, Brazil (DZ 41.192). 39. Artines cofus sp. nov., paratype, Tambopata, Puerto Maldonado Madre de Díos, Peru (MUSM). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241055/files/figure.png" pageId="20" pageNumber="21">Fig. 39E</figureCitation>
); aedeagus with proximal tip narrow (
<figureCitation id="DBC0FBA89D4ECA59FAD2FBF6FA9DFB1D" box="[1337,1406,1141,1163]" captionStart="FIGURES 38–39" captionStartId="29.[151,264,1720,1745]" captionTargetBox="[161,1436,431,1681]" captionTargetId="figure@29.[151,1436,450,1693]" captionTargetPageId="29" captionText="FIGURES 38–39. Male genitalia of the “aquilina group” species. 38. Artines focus, Reserva Duque, Manaus, Amazonas, Brazil (DZ 41.192). 39. Artines cofus sp. nov., paratype, Tambopata, Puerto Maldonado Madre de Díos, Peru (MUSM). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241055/files/figure.png" pageId="20" pageNumber="21">Fig. 39</figureCitation>
G– J)........................................................................................
<taxonomicName id="84FB9CAE9D4ECA59FAFCFB10FAA9FB3F" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[1303,1354,1171,1192]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="cofus" status="sp. nov.">
<emphasis id="718F3B3F9D4ECA59FAFCFB10FAA9FB3F" box="[1303,1354,1171,1192]" italics="true" pageId="20" pageNumber="21">cofus</emphasis>
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<taxonomicNameLabel id="6ABC86449D4ECA59FABBFB10FA7FFB3F" box="[1360,1436,1171,1192]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
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<keyLead id="F80A44189D4ECA59FF7CFB33FA7FFB75" pageId="20" pageNumber="21">
- Uncus with distal end weakly bifid (
<figureCitation id="DBC0FBA89D4ECA59FDAAFB33FD64FB51" box="[577,647,1200,1222]" captionStart="FIGURES 40–41" captionStartId="31.[151,264,1624,1649]" captionTargetBox="[154,1435,484,1599]" captionTargetId="figure@31.[151,1436,481,1612]" captionText="FIGURES 40–41. Male genitalia of the “aquilina group” (40) and “bipunctata group” (41) species. 40. Artines donia sp. nov., paratype, Santa Rita do Araguaia, Goiás, Brazil (DZ 66.000). 41. Artines bipunctata, Brasília, Distrito Federal, Brazil (DZ 41.194). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3406272/files/figure.png" pageId="20" pageNumber="21">Fig. 40</figureCitation>
C–D); valva not pointed distally (
<figureCitation id="DBC0FBA89D4ECA59FC21FB33FBC1FB51" box="[970,1058,1200,1222]" captionStart="FIGURES 40–41" captionStartId="31.[151,264,1624,1649]" captionTargetBox="[154,1435,484,1599]" captionTargetId="figure@31.[151,1436,481,1612]" captionText="FIGURES 40–41. Male genitalia of the “aquilina group” (40) and “bipunctata group” (41) species. 40. Artines donia sp. nov., paratype, Santa Rita do Araguaia, Goiás, Brazil (DZ 66.000). 41. Artines bipunctata, Brasília, Distrito Federal, Brazil (DZ 41.194). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3406272/files/figure.png" pageId="20" pageNumber="21">Fig. 40E</figureCitation>
); Aedeagus uniformly wide (
<figureCitation id="DBC0FBA89D4ECA59FAD1FB33FA9CFB52" box="[1338,1407,1200,1222]" captionStart="FIGURES 40–41" captionStartId="31.[151,264,1624,1649]" captionTargetBox="[154,1435,484,1599]" captionTargetId="figure@31.[151,1436,481,1612]" captionText="FIGURES 40–41. Male genitalia of the “aquilina group” (40) and “bipunctata group” (41) species. 40. Artines donia sp. nov., paratype, Santa Rita do Araguaia, Goiás, Brazil (DZ 66.000). 41. Artines bipunctata, Brasília, Distrito Federal, Brazil (DZ 41.194). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3406272/files/figure.png" pageId="20" pageNumber="21">Fig. 40</figureCitation>
G– J)......................................................................................
<taxonomicName id="84FB9CAE9D4ECA59FAF8FB4EFAA9FB75" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[1299,1354,1229,1250]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="donia" status="sp. nov.">
<emphasis id="718F3B3F9D4ECA59FAF8FB4EFAA9FB75" box="[1299,1354,1229,1250]" italics="true" pageId="20" pageNumber="21">donia</emphasis>
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<emphasis id="718F3B3F9D4ECA59FABBFB4EFA7FFB75" bold="true" box="[1360,1436,1229,1250]" pageId="20" pageNumber="21">
<taxonomicNameLabel id="6ABC86449D4ECA59FABBFB4EFA7FFB75" box="[1360,1436,1229,1250]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
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“
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<figureCitation id="DBC0FBA89D4ECA59FF7CFF3DFF18FF4F" box="[151,251,190,216]" captionStart-0="FIGURES 1–2" captionStart-1="FIGURES 3–4" captionStartId-0="2.[151,264,2012,2037]" captionStartId-1="5.[151,264,712,737]" captionTargetBox-0="[153,1433,1143,1971]" captionTargetBox-1="[153,1432,238,691]" captionTargetId-0="figure@2.[151,1436,1127,1998]" captionTargetId-1="figure@5.[151,1436,237,693]" captionTargetPageId-0="2" captionTargetPageId-1="5" captionText-0="FIGURES 1–2. Color pattern of ventral wings of Artines: 1. Artines bamba sp. nov.; 2. Artines aquilina." captionText-1="FIGURES 3–4. Androconial organs present in the species of Artines. 3. Brand (with scales) on dorsal forewing of Artines aquilina (absent in “aepitus group”). 4. Tuft of elongated scales along 2A of the dorsal hind wing of Artines liege sp. nov. (presente in all species of Artines)." httpUri-0="https://zenodo.org/record/3241037/files/figure.png" httpUri-1="https://zenodo.org/record/3241039/files/figure.png" pageId="20" pageNumber="21">Figs 2–3</figureCitation>
,
<figureCitation id="DBC0FBA89D4ECA59FEEDFF3DFEACFF4F" box="[262,335,190,216]" captionStart="FIGURES 14–19" captionStartId="10.[151,264,1406,1431]" captionTargetBox="[155,1435,175,1386]" captionTargetId="figure@10.[151,1436,174,1394]" captionTargetPageId="10" captionText="FIGURES 14–19. Species of the “aquilina group” (14–18) and “bipunctata group” (19) in dorsal and ventral views. 14. Artines aquilina, A–B: male, Joinville, Santa Catarina, Brazil (DZ 41.321), C–D: female from same locality of male (DZ 41.220). 15. Artines angelica sp. nov., A–B: holotype male, C–D: allotype female. 16. Artines focus, A–B: male, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.233), C–D: female, Pacaraima, Roraima, Brazil (DZ 41.333). 17. Artines cofus sp. nov., A–B: holotype male, C–D: allotype female. 18. Artines donia sp. nov., A–B: holotype male. 19. Artines bipunctata, A–B: male, Jardim Zoológico, Brasília, Distrito Federal, Brazil (DZ 30.961), C–D: female, Córrego Bananal, Brasília, Distrito Federal, Brazil (DZ 41.224). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241045/files/figure.png" pageId="20" pageNumber="21">14–18</figureCitation>
,
<figureCitation id="DBC0FBA89D4ECA59FEB1FF3DFE9AFF4F" box="[346,377,190,216]" captionStart="FIGURE 20" captionStartId="22.[151,250,847,872]" captionTargetBox="[157,1436,448,824]" captionTargetId="figure@22.[151,1436,445,835]" captionText="FIGURE 20. Variation in the color pattern of the ventral forewing of Artines aquilina. A: Joinville, Santa Catarina, Brazil; B: Lençóis, Bahia, Brazil and C: Caranavi, La Paz, Bolivia. Scale bar = 1 cm." httpUri="https://zenodo.org/record/3406268/files/figure.png" pageId="20" pageNumber="21">20</figureCitation>
,
<figureCitation id="DBC0FBA89D4ECA59FE6FFF3EFE2EFF4F" box="[388,461,189,216]" captionStart="FIGURES 21–26" captionStartId="22.[151,264,1840,1865]" captionTargetBox="[151,1436,922,1828]" captionTargetId="figure@22.[151,1436,922,1828]" captionText="FIGURES 21–26. Male brands (no scales) on dorsal forewing of the “aquilina group” (21–25) and “bipunctata group” species (26). 21. Artines aquilina, Joinville, Santa catarina, Brazil, (DZ 41.341). 22. Artines angelica sp. nov., paratype, Fazenda Ter- rasse, Santa Luzia, Maranhão, Brazil (DZ 41.273) 23. Artines focus, Ilha de Maracá, Roraima, Brazil (DZ 41.233). 24. Artines cofus sp. nov., paratype, Santa Rosa do Purus, Acre, Brazil (DZ 41.303). 25. Artines donia sp. nov., paratype, Santa Rita do Araguaia, Goiás, Brazil (OM 66.000). 26. Artines bipunctata, Ponte Funda, Vianópilis, Goiás, Brazil (DZ 41.264). Scale bar = 1 mm." httpUri="https://zenodo.org/record/3406270/files/figure.png" pageId="20" pageNumber="21">21–25</figureCitation>
,
<figureCitation id="DBC0FBA89D4ECA59FE33FF3DFDC2FF4F" box="[472,545,190,216]" captionStart-0="FIGURES 36–37" captionStart-1="FIGURES 38–39" captionStart-2="FIGURES 40–41" captionStartId-0="26.[151,264,1557,1582]" captionStartId-1="29.[151,264,1720,1745]" captionStartId-2="31.[151,264,1624,1649]" captionTargetBox-0="[157,1435,179,1543]" captionTargetBox-1="[161,1436,431,1681]" captionTargetBox-2="[154,1435,484,1599]" captionTargetId-0="figure@26.[151,1436,174,1546]" captionTargetId-1="figure@29.[151,1436,450,1693]" captionTargetId-2="figure@31.[151,1436,481,1612]" captionTargetPageId-0="26" captionTargetPageId-1="29" captionText-0="FIGURES 36–37. Male genitalia of the “aquilina group” species. 36. Artines aquilina, Caminho da Lagoinha, Cachoeiras de Macacú, Rio de Janeiro, Brazil (DZ 41.196). 37. Artines angelica sp. nov., holotype. M) dorsal view of vesica showing the cornuti. For reference of the letters see legend of the figure 27. Scale bar = 1 mm." captionText-1="FIGURES 38–39. Male genitalia of the “aquilina group” species. 38. Artines focus, Reserva Duque, Manaus, Amazonas, Brazil (DZ 41.192). 39. Artines cofus sp. nov., paratype, Tambopata, Puerto Maldonado Madre de Díos, Peru (MUSM). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." captionText-2="FIGURES 40–41. Male genitalia of the “aquilina group” (40) and “bipunctata group” (41) species. 40. Artines donia sp. nov., paratype, Santa Rita do Araguaia, Goiás, Brazil (DZ 66.000). 41. Artines bipunctata, Brasília, Distrito Federal, Brazil (DZ 41.194). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri-0="https://zenodo.org/record/3241053/files/figure.png" httpUri-1="https://zenodo.org/record/3241055/files/figure.png" httpUri-2="https://zenodo.org/record/3406272/files/figure.png" pageId="20" pageNumber="21">36–40</figureCitation>
, 49–53, 61–64, 73–76, 80
</paragraph>
<paragraph id="4344E72D9D4ECA59FF7CFE86FD13FD3B" blockId="20.[151,1437,261,756]" pageId="20" pageNumber="21">
<emphasis id="718F3B3F9D4ECA59FF7CFE86FEC2FEB7" bold="true" box="[151,289,261,288]" pageId="20" pageNumber="21">Description</emphasis>
.
<emphasis id="718F3B3F9D4ECA59FEDBFE85FE89FEB7" box="[304,362,262,288]" italics="true" pageId="20" pageNumber="21">Male</emphasis>
and
<emphasis id="718F3B3F9D4ECA59FE49FE85FE0EFEB7" box="[418,493,262,288]" italics="true" pageId="20" pageNumber="21">female</emphasis>
: dorsal forewing with semi-hyaline and large discal spots; ventral forewing with or without white spot in distal 1/2 of CuA
<subScript id="DF7FE5689D4ECA59FDAAFEB8FDAAFEDE" attach="left" box="[577,585,315,329]" fontSize="6" pageId="20" pageNumber="21">2</subScript>
–2A; ventral hind wing with discal area yellow to orange, with a central black spot and a yellow postdiscal band limited distally by ground stripe, wavy or not.
<emphasis id="718F3B3F9D4ECA59FBCDFECDFB83FEFF" box="[1062,1120,334,360]" italics="true" pageId="20" pageNumber="21">Male</emphasis>
: brand present; saccus wide and long, shape variable; uncus long and narrow; arms of gnathos with pointed apexes; valva with distal projection composed of harpe and ampulla, with triangular aspect from 1/2 its length; harpe simple and fused with ampulla; aedeagus without disto-dorsal projection; ventral projection of aedeagus wide, distally bifid or trifid, diverging ventrally at angle of approximately 45 degrees with aedeagus; cornutus as a moderately sclerotized plate over most of vesica and with spines of variable thickness and length; fultura inferior arched in lateral view, proximal end simple.
<emphasis id="718F3B3F9D4ECA59FF04FDA5FEA0FDD7" box="[239,323,550,576]" italics="true" pageId="20" pageNumber="21">Female</emphasis>
: externally similar to male; region between seventh sternite and lamella antevaginalis membranous; sterigma partly sclerotized, rectangular and joined with tergum by the lateral plate (
<figureCitation id="DBC0FBA89D4ECA59FBDEFDC9FB7FFDF2" box="[1077,1180,586,613]" captionStart="FIGURES 61–65" captionStartId="36.[151,264,653,678]" captionTargetBox="[151,1435,178,637]" captionTargetId="figure@36.[151,1436,174,642]" captionTargetPageId="36" captionText="FIGURES 61–65. Female genitalia of the “aquilina group” and “bipunctata group” species. 61. Artines aquilina, Cananéia, São Paulo, Brazil (OM 49.831). 62. Artines angelica sp. nov., paratype, Tambopata, Puerto Maldonado, Madre de Díos, Peru (DZ 41.222) 63. Artines focus, Pacaraima, Roraima, Brazil (DZ 41.333). 64. Artines cofus sp. nov. paratype, Tambopata, Puerto Maldonado, Madre de Díos, Peru (DZ 41. 293). 65. Artines bipunctata, Reserva IBGE, Brasília, Distrito Federal, Brazil (DZ 41.165). For reference of the letters see legend of the figure 54. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241061/files/figure.png" pageId="20" pageNumber="21">Fig. 61A</figureCitation>
); lamella postvaginalis with one or two sclerotized plates (
<figureCitation id="DBC0FBA89D4ECA59FDC8FDEDFD68FD1E" box="[547,651,622,649]" captionStart="FIGURES 61–65" captionStartId="36.[151,264,653,678]" captionTargetBox="[151,1435,178,637]" captionTargetId="figure@36.[151,1436,174,642]" captionTargetPageId="36" captionText="FIGURES 61–65. Female genitalia of the “aquilina group” and “bipunctata group” species. 61. Artines aquilina, Cananéia, São Paulo, Brazil (OM 49.831). 62. Artines angelica sp. nov., paratype, Tambopata, Puerto Maldonado, Madre de Díos, Peru (DZ 41.222) 63. Artines focus, Pacaraima, Roraima, Brazil (DZ 41.333). 64. Artines cofus sp. nov. paratype, Tambopata, Puerto Maldonado, Madre de Díos, Peru (DZ 41. 293). 65. Artines bipunctata, Reserva IBGE, Brasília, Distrito Federal, Brazil (DZ 41.165). For reference of the letters see legend of the figure 54. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241061/files/figure.png" pageId="20" pageNumber="21">Fig. 62B</figureCitation>
); lateral fold of lamella postvaginalis membranous (
<figureCitation id="DBC0FBA89D4ECA59FB3CFDEDFADDFD1E" box="[1239,1342,622,649]" captionStart="FIGURES 61–65" captionStartId="36.[151,264,653,678]" captionTargetBox="[151,1435,178,637]" captionTargetId="figure@36.[151,1436,174,642]" captionTargetPageId="36" captionText="FIGURES 61–65. Female genitalia of the “aquilina group” and “bipunctata group” species. 61. Artines aquilina, Cananéia, São Paulo, Brazil (OM 49.831). 62. Artines angelica sp. nov., paratype, Tambopata, Puerto Maldonado, Madre de Díos, Peru (DZ 41.222) 63. Artines focus, Pacaraima, Roraima, Brazil (DZ 41.333). 64. Artines cofus sp. nov. paratype, Tambopata, Puerto Maldonado, Madre de Díos, Peru (DZ 41. 293). 65. Artines bipunctata, Reserva IBGE, Brasília, Distrito Federal, Brazil (DZ 41.165). For reference of the letters see legend of the figure 54. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241061/files/figure.png" pageId="20" pageNumber="21">Fig. 61B</figureCitation>
); ductus bursae with basal portion sclerotized or membranous.
</paragraph>
<paragraph id="4344E72D9D4ECA59FF2CFD35FB1AFD63" blockId="20.[151,1437,261,756]" pageId="20" pageNumber="21">
<emphasis id="718F3B3F9D4ECA59FF2CFD35FE74FD47" bold="true" box="[199,407,693,720]" pageId="20" pageNumber="21">Included species:</emphasis>
<taxonomicName id="84FB9CAE9D4ECA59FE4AFD35FD18FD47" authority="(Plotz, 1882)" baseAuthorityName="Plotz" baseAuthorityYear="1882" box="[417,763,694,720]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="aquilina">
<emphasis id="718F3B3F9D4ECA59FE4AFD35FDBAFD47" box="[417,601,694,720]" italics="true" pageId="20" pageNumber="21">Artines aquilina</emphasis>
(
<bibRefCitation id="276A9ADC9D4ECA59FD80FD35FD10FD47" author="Plotz, C." box="[619,755,694,720]" pageId="20" pageNumber="21" pagination="26 - 64" refId="ref44660" refString="Plotz, C. (1882) Die Hesperiinen-Gattung Hesperia Aut. und ihre Arten. Stettiner entomologische Zeitung, 44 (1 / 3), 26 - 64. https: // doi. org / 10.1002 / mmnd. 47918820306" type="journal article" year="1882">Plötz, 1882</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="84FB9CAE9D4ECA59FCE1FD35FBDAFD47" authority="Medeiros" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[778,1081,694,720]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="angelica" status="sp. nov.">
<emphasis id="718F3B3F9D4ECA59FCE1FD35FC26FD47" box="[778,965,694,720]" italics="true" pageId="20" pageNumber="21">Artines angelica</emphasis>
Medeiros
</taxonomicName>
<emphasis id="718F3B3F9D4ECA59FBA8FD35FB43FD47" bold="true" box="[1091,1184,694,720]" pageId="20" pageNumber="21">
<taxonomicNameLabel id="6ABC86449D4ECA59FBA8FD35FB43FD47" box="[1091,1184,694,720]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="84FB9CAE9D4ECA59FB5AFD35FF31FD63" authority="Evans, 1955" authorityName=", Reserva Duque, Manaus" authorityYear="1955" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="focus">
<emphasis id="718F3B3F9D4ECA59FB5AFD35FAA4FD47" box="[1201,1351,694,720]" italics="true" pageId="20" pageNumber="21">Artines focus</emphasis>
<bibRefCitation id="276A9ADC9D4ECA59FABAFD35FF31FD63" author="Evans, W. H." pageId="20" pageNumber="21" refId="ref40325" refString="Evans, W. H. (1955) A Catalogue of the American Hesperiidae indicating the Classification and Nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. British Museum, London, 499 pp." type="book" year="1955">Evans, 1955</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="84FB9CAE9D4ECA59FF36FD59FD90FD63" authority="Medeiros &amp; Dolibaina" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[221,627,730,756]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="cofus" status="sp. nov.">
<emphasis id="718F3B3F9D4ECA59FF36FD59FE93FD63" box="[221,368,730,756]" italics="true" pageId="20" pageNumber="21">Artines cofus</emphasis>
Medeiros &amp; Dolibaina
</taxonomicName>
<emphasis id="718F3B3F9D4ECA59FD91FD59FD37FD63" bold="true" box="[634,724,730,756]" pageId="20" pageNumber="21">
<taxonomicNameLabel id="6ABC86449D4ECA59FD91FD59FD37FD63" box="[634,724,730,756]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="84FB9CAE9D4ECA59FCE1FD59FB7BFD63" authority="Medeiros &amp; Carneiro" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[778,1176,730,756]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="donia" status="sp. nov.">
<emphasis id="718F3B3F9D4ECA59FCE1FD59FC41FD63" box="[778,930,730,756]" italics="true" pageId="20" pageNumber="21">Artines donia</emphasis>
Medeiros &amp; Carneiro
</taxonomicName>
<emphasis id="718F3B3F9D4ECA59FB74FD59FB1AFD63" bold="true" box="[1183,1273,730,756]" pageId="20" pageNumber="21">
<taxonomicNameLabel id="6ABC86449D4ECA59FB74FD59FB1AFD63" box="[1183,1273,730,756]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

417
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<treatment id="CB52563B9D50CA46FF7CF941FC98F848" ID-GBIF-Taxon="157436008" LSID="urn:lsid:zoobank.org:act:0AAE3E9D-3557-48E6-AB04-392D7E4EB494" httpUri="http://treatment.plazi.org/id/CB52563B9D50CA46FF7CF941FC98F848" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<subSubSection id="0BE1B4A69D50CA47FF7CF941FD31F94B" box="[151,722,1729,1756]" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="4344E72D9D50CA47FF7CF941FD31F94B" blockId="10.[151,973,1729,1828]" box="[151,722,1729,1756]" pageId="10" pageNumber="11">
<heading id="180C50419D50CA47FF7CF941FD31F94B" bold="true" box="[151,722,1729,1756]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
<emphasis id="718F3B3F9D50CA47FF7CF941FD31F94B" bold="true" box="[151,722,1729,1756]" pageId="10" pageNumber="11">
<taxonomicName id="84FB9CAE9D50CA47FF7CF941FD8DF94C" authority="Medeiros &amp; Carneiro" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[151,622,1729,1756]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="panama" status="sp. nov.">
<emphasis id="718F3B3F9D50CA47FF7CF941FEBAF94B" bold="true" box="[151,345,1730,1756]" italics="true" pageId="10" pageNumber="11">Artines panama</emphasis>
Medeiros &amp; Carneiro
</taxonomicName>
<taxonomicNameLabel id="6ABC86449D50CA47FD9FF941FD31F94B" box="[628,722,1730,1756]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D50CA47FF7CF966FC2EF8B3" pageId="10" pageNumber="11" type="description">
<paragraph id="4344E72D9D50CA47FF7CF966FE4DF897" blockId="10.[151,973,1729,1828]" box="[151,430,1765,1792]" pageId="10" pageNumber="11">
<figureCitation id="DBC0FBA89D50CA47FF7CF966FF3DF968" box="[151,222,1765,1791]" captionStart="FIGURES 5–8" captionStartId="6.[151,264,1064,1089]" captionTargetBox="[154,1432,269,1044]" captionTargetId="figure@6.[151,1436,265,1052]" captionTargetPageId="6" captionText="FIGURES 5–8. Species of the “aepitus group” in dorsal and ventral views. 5. Artines aepitus, A–B: male, Cueva de Las Pavas, Tingo Maria, Huánuco, Peru (OM 80.822), C–D: female, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.790). 6. Artines panama sp. nov., A–B: holotype male, C–D: allotype female. 7. Artines solange sp. nov., A–B: holotype male, C–D: tentative female, Sumaúma, Maranhão, Brazil (DZ 41.800). 8. Artines delfos sp. nov., A–B: holotype male. Scale bar = 1 cm." httpUri="https://zenodo.org/record/3241041/files/figure.png" pageId="10" pageNumber="11">Figs 6</figureCitation>
,
<figureCitation id="DBC0FBA89D50CA47FF01F965FEEBF897" box="[234,264,1766,1792]" captionStart="FIGURES 27–29" captionStartId="23.[151,264,1797,1822]" captionTargetBox="[153,1436,174,1777]" captionTargetId="figure@23.[151,1436,174,1786]" captionTargetPageId="23" captionText="FIGURES 27–29. Male genitalia of the “aepitus group” species. 27. Artines aepitus, Jusepín, Monagas, Venezuela (OM 29.417). 28. Artines panama sp. nov., holotype. 29. Artines solange sp. nov., holotype. A) lateral view of tegumen, saccus, uncus and gnathos. B) ventral view of saccus. C) dorsal view of tegumen, uncus and gnathos. D) ventral view of tegumen, uncus and gnathos. E) interior view of right valva. F) dorsal view of right valva. G) dorsal view of aedeagus and vesica. H) left lateral view of aedeagus. I) ventral view of aedeagus. J) right lateral view of aedeagus. K) dorsal view of fultura inferior. L) lateral view of fultura inferior. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241047/files/figure.png" pageId="10" pageNumber="11">28</figureCitation>
,
<figureCitation id="DBC0FBA89D50CA47FEFFF965FED1F897" box="[276,306,1766,1792]" captionStart="FIGURES 42–48" captionStartId="33.[151,264,1892,1917]" captionTargetBox="[151,1431,625,1874]" captionTargetId="figure@33.[151,1436,625,1880]" captionText="FIGURES 42–48. Dorsal view of vesica and cornutus of the &quot;aepitus group&quot; species. 42. Artines aepitus. 43. Artines panama sp. nov.. 44. Artines delfos sp. nov.. 45. Artines litoralis sp. nov.. 46. Artines rica. 47. Artines liege sp. nov. 48. Artines bamba sp. nov. Scale bar = 0,2 mm." httpUri="https://zenodo.org/record/3406274/files/figure.png" pageId="10" pageNumber="11">43</figureCitation>
, 55, 67, 78
</paragraph>
<paragraph id="4344E72D9D50CA47FF7CF88AFC2EF8B3" blockId="10.[151,973,1729,1828]" box="[151,973,1801,1828]" pageId="10" pageNumber="11">
<uri id="376AEB2F9D50CA47FF7CF88AFC2EF8B3" box="[151,973,1801,1828]" pageId="10" pageNumber="11">
urn:lsid:zoobank.org:act:
<uuid id="375DDDF89D50CA47FE42F88AFC2EF8B3" box="[425,973,1801,1828]" pageId="10" pageNumber="11">0AAE3E9D-3557-48E6-AB04-392D7E4EB494</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D50CA46FF7CF8E0FF0AFF40" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="reference_group">
<paragraph id="4344E72D9D50CA47FF7CF8E0FC09F82E" blockId="10.[151,1437,1890,1977]" pageId="10" pageNumber="11">
<taxonomicName id="84FB9CAE9D50CA47FF7CF8E0FECBF8EC" ID-CoL="67SFD" authorityName="Godman" authorityYear="1901" box="[151,296,1891,1915]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="atizies">
<emphasis id="718F3B3F9D50CA47FF7CF8E0FECBF8EC" box="[151,296,1891,1915]" italics="true" pageId="10" pageNumber="11">Artines atizies</emphasis>
</taxonomicName>
(misid.);
<bibRefCitation id="276A9ADC9D50CA47FE67F8E1FDC5F8EC" author="Godman, F. D." box="[396,550,1890,1915]" pageId="10" pageNumber="11" pagination="142 - 457" refId="ref41039" refString="Godman, F. D. (1901) Hesperiidae [parte]. In: Godman, F. D. &amp; Salvin, O. (1879 - 1901), Biologia Centrali-Americana, Insecta, Lepidoptera Rhopalocera. Vol. 2. pp. 142 - 457, pls. 91 - 112]" type="journal article" year="1901">Godman, 1901</bibRefCitation>
,
<emphasis id="718F3B3F9D50CA47FDDAF8E0FDA5F8EC" bold="true" box="[561,582,1891,1915]" italics="true" pageId="10" pageNumber="11">in</emphasis>
Godman &amp; Salvin.
<emphasis id="718F3B3F9D50CA47FCFFF8E1FB95F8EC" bold="true" box="[788,1142,1890,1915]" pageId="10" pageNumber="11">Biol. Centr.-Amer., Lep.-Rhop. 2</emphasis>
, p. 608.—
<bibRefCitation id="276A9ADC9D50CA47FB0AF8E1FA7AF8EC" author="Huntington, E. I." box="[1249,1433,1890,1915]" pageId="10" pageNumber="11" pagination="191 - 230" refId="ref41699" refString="Huntington, E. I. (1932) A list of the Rhopalocera of Barro Colorado Island, Canal Zone, Panama, with descriptions of two new species of Theclinae. Bulletin of the American Museum of Natural History, 63 (3), 191 - 230." type="journal article" year="1932">Huntington, 1932</bibRefCitation>
.
<emphasis id="718F3B3F9D50CA47FF2CF801FDE8F80D" bold="true" box="[199,523,1921,1946]" pageId="10" pageNumber="11">Bull. Amer. Mus. Nat. Hist. 63</emphasis>
: 230.
<taxonomicName id="84FB9CAE9D50CA47FF7CF822FED3F82F" ID-CoL="GZYC" baseAuthorityName="Geyer" baseAuthorityYear="1832" box="[151,304,1953,1976]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D50CA47FF7CF822FED3F82F" box="[151,304,1953,1976]" italics="true" pageId="10" pageNumber="11">Artines aepitus</emphasis>
</taxonomicName>
(misid.);
<bibRefCitation id="276A9ADC9D50CA47FE78F822FDB2F82E" author="Basset, Y. &amp; Barrios, H. &amp; Segar, S. &amp; Srygley, R. B. &amp; Aiello, A. &amp; Warren, A. D. &amp; Delgado, F. &amp; Coronado, J. &amp; Lezcano, J. &amp; Arizala, S. &amp; Rivera, M. &amp; Perez, F. &amp; Bobadilla, R. &amp; Lopez, Y. &amp; Ramirez, A. J." box="[403,593,1952,1977]" pageId="10" pageNumber="11" pagination="0136623" refId="ref38212" refString="Basset, Y., Barrios, H., Segar, S., Srygley, R. B., Aiello, A., Warren, A. D., Delgado, F., Coronado, J., Lezcano, J., Arizala, S., Rivera, M., Perez, F., Bobadilla, R., Lopez, Y. &amp; Ramirez, A. J. (2015) The butterflies of Barro Colorado Island, Panama: Local extinction since the 1930 s. PLoS ONE, 10 (8), e 0136623, 1 - 22. https: // doi. org / 10.1371 / journal. pone. 0136623" type="journal article" year="2015">
Basset
<emphasis id="718F3B3F9D50CA47FE37F822FDEBF82E" bold="true" box="[476,520,1952,1977]" italics="true" pageId="10" pageNumber="11">et al</emphasis>
., 2015
</bibRefCitation>
.
<emphasis id="718F3B3F9D50CA47FDB1F823FD11F82E" bold="true" box="[602,754,1952,1977]" pageId="10" pageNumber="11">PLoS ONE 10</emphasis>
(8) (e0136623), S1 App.
</paragraph>
<paragraph id="4344E72D9D51CA46FF7CFF1BFF0AFF40" blockId="11.[151,1437,152,2015]" pageId="11" pageNumber="12">
<emphasis id="718F3B3F9D51CA46FF7CFF1BFE9BFF24" bold="true" box="[151,376,152,179]" pageId="11" pageNumber="12">Taxonomic history</emphasis>
. This species was treated as
<taxonomicName id="84FB9CAE9D51CA46FD23FF1AFCD2FF25" ID-CoL="67SFD" authorityName="Godman" authorityYear="1901" box="[712,817,153,179]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="atizies">
<emphasis id="718F3B3F9D51CA46FD23FF1AFCD2FF25" box="[712,817,153,179]" italics="true" pageId="11" pageNumber="12">A. atizies</emphasis>
</taxonomicName>
by
<bibRefCitation id="276A9ADC9D51CA46FC8BFF1BFBD0FF24" author="Huntington, E. I." box="[864,1075,152,179]" pageId="11" pageNumber="12" pagination="191 - 230" refId="ref41699" refString="Huntington, E. I. (1932) A list of the Rhopalocera of Barro Colorado Island, Canal Zone, Panama, with descriptions of two new species of Theclinae. Bulletin of the American Museum of Natural History, 63 (3), 191 - 230." type="journal article" year="1932">Huntington (1932)</bibRefCitation>
and
<taxonomicName id="84FB9CAE9D51CA46FB85FF1AFB02FF25" ID-CoL="GZYC" baseAuthorityName="Geyer" baseAuthorityYear="1832" box="[1134,1249,153,179]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D51CA46FB85FF1AFB02FF25" box="[1134,1249,153,179]" italics="true" pageId="11" pageNumber="12">A. aepitus</emphasis>
</taxonomicName>
by
<bibRefCitation id="276A9ADC9D51CA46FAE4FF1AFF06FF40" author="Basset, Y. &amp; Barrios, H. &amp; Segar, S. &amp; Srygley, R. B. &amp; Aiello, A. &amp; Warren, A. D. &amp; Delgado, F. &amp; Coronado, J. &amp; Lezcano, J. &amp; Arizala, S. &amp; Rivera, M. &amp; Perez, F. &amp; Bobadilla, R. &amp; Lopez, Y. &amp; Ramirez, A. J." pageId="11" pageNumber="12" pagination="0136623" refId="ref38212" refString="Basset, Y., Barrios, H., Segar, S., Srygley, R. B., Aiello, A., Warren, A. D., Delgado, F., Coronado, J., Lezcano, J., Arizala, S., Rivera, M., Perez, F., Bobadilla, R., Lopez, Y. &amp; Ramirez, A. J. (2015) The butterflies of Barro Colorado Island, Panama: Local extinction since the 1930 s. PLoS ONE, 10 (8), e 0136623, 1 - 22. https: // doi. org / 10.1371 / journal. pone. 0136623" type="journal article" year="2015">
Basset
<emphasis id="718F3B3F9D51CA46FA89FF1AFA76FF25" box="[1378,1429,152,178]" italics="true" pageId="11" pageNumber="12">et al</emphasis>
. (2015)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D51CA46FF2CFF63FE24FE1D" pageId="11" pageNumber="12" type="diagnosis">
<paragraph id="4344E72D9D51CA46FF2CFF63FE24FE1D" blockId="11.[151,1437,152,2015]" pageId="11" pageNumber="12">
<emphasis id="718F3B3F9D51CA46FF2CFF63FED9FF6C" bold="true" box="[199,314,224,251]" pageId="11" pageNumber="12">Diagnosis</emphasis>
.
<taxonomicName id="84FB9CAE9D51CA46FEA3FF62FE1AFF6D" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[328,505,225,251]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="panama" status="sp. nov.">
<emphasis id="718F3B3F9D51CA46FEA3FF62FE1AFF6D" box="[328,505,225,251]" italics="true" pageId="11" pageNumber="12">Artines panama</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D51CA46FDEBFF62FDBAFF6C" bold="true" box="[512,601,225,251]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="6ABC86449D51CA46FDEBFF62FDBAFF6C" box="[512,601,225,251]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from the other species of the “
<taxonomicName id="84FB9CAE9D51CA46FC14FF62FBB3FF6D" baseAuthorityName="Geyer" baseAuthorityYear="1832" box="[1023,1104,225,250]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D51CA46FC14FF62FBB3FF6D" box="[1023,1104,225,250]" italics="true" pageId="11" pageNumber="12">aepitus</emphasis>
</taxonomicName>
group” in the following combination of characters: harpe with ventral process wide, slightly shorter than tegumen+uncus and with distal 1/3 dilated and rounded; dorsal process of harpe on median portion; sterigma narrower at base; lamella antevaginalis with proximo-ventral folds more or less rectangular; lamella postvaginalis with proximo-lateral projections weakly developed and not bilobed.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D51CA46FF2CFE17FD39FAA5" pageId="11" pageNumber="12" type="description">
<paragraph id="4344E72D9D51CA46FF2CFE17FBB4FE38" blockId="11.[151,1437,152,2015]" box="[199,1111,404,431]" pageId="11" pageNumber="12">
<emphasis id="718F3B3F9D51CA46FF2CFE17FEB2FE38" bold="true" box="[199,337,404,431]" pageId="11" pageNumber="12">Description</emphasis>
. Male (
<figureCitation id="DBC0FBA89D51CA46FE43FE17FE08FE38" box="[424,491,404,431]" captionStart="FIGURES 5–8" captionStartId="6.[151,264,1064,1089]" captionTargetBox="[154,1432,269,1044]" captionTargetId="figure@6.[151,1436,265,1052]" captionTargetPageId="6" captionText="FIGURES 5–8. Species of the “aepitus group” in dorsal and ventral views. 5. Artines aepitus, A–B: male, Cueva de Las Pavas, Tingo Maria, Huánuco, Peru (OM 80.822), C–D: female, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.790). 6. Artines panama sp. nov., A–B: holotype male, C–D: allotype female. 7. Artines solange sp. nov., A–B: holotype male, C–D: tentative female, Sumaúma, Maranhão, Brazil (DZ 41.800). 8. Artines delfos sp. nov., A–B: holotype male. Scale bar = 1 cm." httpUri="https://zenodo.org/record/3241041/files/figure.png" pageId="11" pageNumber="12">Fig. 6</figureCitation>
A-B).
<emphasis id="718F3B3F9D51CA46FDD9FE17FD8CFE39" box="[562,623,404,430]" italics="true" pageId="11" pageNumber="12">Head</emphasis>
: antenna with nudum bearing 12 segments.
</paragraph>
<paragraph id="4344E72D9D51CA46FF2CFE3AFE06FE44" blockId="11.[151,1437,152,2015]" box="[199,485,440,467]" pageId="11" pageNumber="12">
<emphasis id="718F3B3F9D51CA46FF2CFE3AFE63FE45" box="[199,384,440,466]" italics="true" pageId="11" pageNumber="12">Forewing length</emphasis>
:
<quantity id="84034AC89D51CA46FE64FE3AFE02FE44" box="[399,481,441,467]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="11" pageNumber="12" unit="mm" value="13.0">13 mm</quantity>
.
</paragraph>
<paragraph id="4344E72D9D51CA46FF2CFE5FFE80FD8C" blockId="11.[151,1437,152,2015]" pageId="11" pageNumber="12">
<emphasis id="718F3B3F9D51CA46FF2CFE5FFE9CFE60" box="[199,383,476,503]" italics="true" pageId="11" pageNumber="12">Dorsal forewing</emphasis>
(
<figureCitation id="DBC0FBA89D51CA46FE64FE5FFE09FE60" box="[399,490,476,503]" captionStart="FIGURES 5–8" captionStartId="6.[151,264,1064,1089]" captionTargetBox="[154,1432,269,1044]" captionTargetId="figure@6.[151,1436,265,1052]" captionTargetPageId="6" captionText="FIGURES 5–8. Species of the “aepitus group” in dorsal and ventral views. 5. Artines aepitus, A–B: male, Cueva de Las Pavas, Tingo Maria, Huánuco, Peru (OM 80.822), C–D: female, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.790). 6. Artines panama sp. nov., A–B: holotype male, C–D: allotype female. 7. Artines solange sp. nov., A–B: holotype male, C–D: tentative female, Sumaúma, Maranhão, Brazil (DZ 41.800). 8. Artines delfos sp. nov., A–B: holotype male. Scale bar = 1 cm." httpUri="https://zenodo.org/record/3241041/files/figure.png" pageId="11" pageNumber="12">Fig. 6A</figureCitation>
): three apical spots; discal spots as in
<emphasis id="718F3B3F9D51CA46FC7DFE5EFBEEFE61" box="[918,1037,477,503]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="84FB9CAE9D51CA46FC7DFE5EFBEAFE61" baseAuthorityName="Geyer" baseAuthorityYear="1832" box="[918,1033,477,503]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="aepitus">A. aepitus</taxonomicName>
,
</emphasis>
but less conspicuous; white spot in CuA
<subScript id="DF7FE5689D51CA46FF27FD91FF37FDB7" attach="left" box="[204,212,530,544]" fontSize="6" pageId="11" pageNumber="12">2</subScript>
–2A present.
</paragraph>
<paragraph id="4344E72D9D51CA46FF2CFDA7FC04FDA9" blockId="11.[151,1437,152,2015]" box="[199,999,548,575]" pageId="11" pageNumber="12">
<emphasis id="718F3B3F9D51CA46FF2CFDA7FE61FDA8" box="[199,386,548,575]" italics="true" pageId="11" pageNumber="12">Ventral forewing</emphasis>
(
<figureCitation id="DBC0FBA89D51CA46FE7AFDA7FE09FDA8" box="[401,490,548,575]" captionStart="FIGURES 5–8" captionStartId="6.[151,264,1064,1089]" captionTargetBox="[154,1432,269,1044]" captionTargetId="figure@6.[151,1436,265,1052]" captionTargetPageId="6" captionText="FIGURES 5–8. Species of the “aepitus group” in dorsal and ventral views. 5. Artines aepitus, A–B: male, Cueva de Las Pavas, Tingo Maria, Huánuco, Peru (OM 80.822), C–D: female, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.790). 6. Artines panama sp. nov., A–B: holotype male, C–D: allotype female. 7. Artines solange sp. nov., A–B: holotype male, C–D: tentative female, Sumaúma, Maranhão, Brazil (DZ 41.800). 8. Artines delfos sp. nov., A–B: holotype male. Scale bar = 1 cm." httpUri="https://zenodo.org/record/3241041/files/figure.png" pageId="11" pageNumber="12">Fig. 6B</figureCitation>
): as in
<taxonomicName id="84FB9CAE9D51CA46FDD2FDA6FD4AFDA9" baseAuthorityName="Geyer" baseAuthorityYear="1832" box="[569,681,549,575]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D51CA46FDD2FDA6FD4AFDA9" box="[569,681,549,575]" italics="true" pageId="11" pageNumber="12">A. aepitus</emphasis>
</taxonomicName>
, except for paler coloration.
</paragraph>
<paragraph id="4344E72D9D51CA46FF2CFDCBFE69FD11" blockId="11.[151,1437,152,2015]" pageId="11" pageNumber="12">
<emphasis id="718F3B3F9D51CA46FF2CFDCBFE6DFDF5" box="[199,398,584,610]" italics="true" pageId="11" pageNumber="12">Ventral hind wing</emphasis>
(
<figureCitation id="DBC0FBA89D51CA46FE75FDCBFE14FDF4" box="[414,503,584,611]" captionStart="FIGURES 5–8" captionStartId="6.[151,264,1064,1089]" captionTargetBox="[154,1432,269,1044]" captionTargetId="figure@6.[151,1436,265,1052]" captionTargetPageId="6" captionText="FIGURES 5–8. Species of the “aepitus group” in dorsal and ventral views. 5. Artines aepitus, A–B: male, Cueva de Las Pavas, Tingo Maria, Huánuco, Peru (OM 80.822), C–D: female, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.790). 6. Artines panama sp. nov., A–B: holotype male, C–D: allotype female. 7. Artines solange sp. nov., A–B: holotype male, C–D: tentative female, Sumaúma, Maranhão, Brazil (DZ 41.800). 8. Artines delfos sp. nov., A–B: holotype male. Scale bar = 1 cm." httpUri="https://zenodo.org/record/3241041/files/figure.png" pageId="11" pageNumber="12">Fig. 6B</figureCitation>
): apical portion of discal cell with few yellow scales; central black spot rectangular and longer than wide.
</paragraph>
<paragraph id="4344E72D9D51CA46FF2CFD13FBC0FBA0" blockId="11.[151,1437,152,2015]" pageId="11" pageNumber="12">
<emphasis id="718F3B3F9D51CA46FF2CFD13FED2FD3D" box="[199,305,656,682]" italics="true" pageId="11" pageNumber="12">Genitalia</emphasis>
(
<figureCitation id="DBC0FBA89D51CA46FED4FD13FE77FD3C" box="[319,404,656,683]" captionStart="FIGURES 27–29" captionStartId="23.[151,264,1797,1822]" captionTargetBox="[153,1436,174,1777]" captionTargetId="figure@23.[151,1436,174,1786]" captionTargetPageId="23" captionText="FIGURES 27–29. Male genitalia of the “aepitus group” species. 27. Artines aepitus, Jusepín, Monagas, Venezuela (OM 29.417). 28. Artines panama sp. nov., holotype. 29. Artines solange sp. nov., holotype. A) lateral view of tegumen, saccus, uncus and gnathos. B) ventral view of saccus. C) dorsal view of tegumen, uncus and gnathos. D) ventral view of tegumen, uncus and gnathos. E) interior view of right valva. F) dorsal view of right valva. G) dorsal view of aedeagus and vesica. H) left lateral view of aedeagus. I) ventral view of aedeagus. J) right lateral view of aedeagus. K) dorsal view of fultura inferior. L) lateral view of fultura inferior. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241047/files/figure.png" pageId="11" pageNumber="12">Figs 28</figureCitation>
A–L, 43): tegumen more or less rectangular, with proximal margin straight and distal margin convex and angled (
<figureCitation id="DBC0FBA89D51CA46FE97FD37FE00FD58" box="[380,483,692,719]" captionStart="FIGURES 27–29" captionStartId="23.[151,264,1797,1822]" captionTargetBox="[153,1436,174,1777]" captionTargetId="figure@23.[151,1436,174,1786]" captionTargetPageId="23" captionText="FIGURES 27–29. Male genitalia of the “aepitus group” species. 27. Artines aepitus, Jusepín, Monagas, Venezuela (OM 29.417). 28. Artines panama sp. nov., holotype. 29. Artines solange sp. nov., holotype. A) lateral view of tegumen, saccus, uncus and gnathos. B) ventral view of saccus. C) dorsal view of tegumen, uncus and gnathos. D) ventral view of tegumen, uncus and gnathos. E) interior view of right valva. F) dorsal view of right valva. G) dorsal view of aedeagus and vesica. H) left lateral view of aedeagus. I) ventral view of aedeagus. J) right lateral view of aedeagus. K) dorsal view of fultura inferior. L) lateral view of fultura inferior. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241047/files/figure.png" pageId="11" pageNumber="12">Fig. 28C</figureCitation>
), in lateral view, shorter than length of gnathos; ventral arm of tegumen longer than dorsal arm of saccus (
<figureCitation id="DBC0FBA89D51CA46FE79FD5BFE1EFD64" box="[402,509,728,755]" captionStart="FIGURES 27–29" captionStartId="23.[151,264,1797,1822]" captionTargetBox="[153,1436,174,1777]" captionTargetId="figure@23.[151,1436,174,1786]" captionTargetPageId="23" captionText="FIGURES 27–29. Male genitalia of the “aepitus group” species. 27. Artines aepitus, Jusepín, Monagas, Venezuela (OM 29.417). 28. Artines panama sp. nov., holotype. 29. Artines solange sp. nov., holotype. A) lateral view of tegumen, saccus, uncus and gnathos. B) ventral view of saccus. C) dorsal view of tegumen, uncus and gnathos. D) ventral view of tegumen, uncus and gnathos. E) interior view of right valva. F) dorsal view of right valva. G) dorsal view of aedeagus and vesica. H) left lateral view of aedeagus. I) ventral view of aedeagus. J) right lateral view of aedeagus. K) dorsal view of fultura inferior. L) lateral view of fultura inferior. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241047/files/figure.png" pageId="11" pageNumber="12">Fig. 28A</figureCitation>
). Saccus shorter than tegumen+uncus, with base approximately as wide as lateral margin (
<figureCitation id="DBC0FBA89D51CA46FF1CFD7FFE82FC80" box="[247,353,764,791]" captionStart="FIGURES 27–29" captionStartId="23.[151,264,1797,1822]" captionTargetBox="[153,1436,174,1777]" captionTargetId="figure@23.[151,1436,174,1786]" captionTargetPageId="23" captionText="FIGURES 27–29. Male genitalia of the “aepitus group” species. 27. Artines aepitus, Jusepín, Monagas, Venezuela (OM 29.417). 28. Artines panama sp. nov., holotype. 29. Artines solange sp. nov., holotype. A) lateral view of tegumen, saccus, uncus and gnathos. B) ventral view of saccus. C) dorsal view of tegumen, uncus and gnathos. D) ventral view of tegumen, uncus and gnathos. E) interior view of right valva. F) dorsal view of right valva. G) dorsal view of aedeagus and vesica. H) left lateral view of aedeagus. I) ventral view of aedeagus. J) right lateral view of aedeagus. K) dorsal view of fultura inferior. L) lateral view of fultura inferior. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241047/files/figure.png" pageId="11" pageNumber="12">Fig. 28B</figureCitation>
). Uncus, in dorsal view, rectangular, about one and a half times longer than wide, shorter than tegumen, without disto-lateral projections, distal margin rounded (
<figureCitation id="DBC0FBA89D51CA46FC94FCA3FC05FCAC" box="[895,998,800,827]" captionStart="FIGURES 27–29" captionStartId="23.[151,264,1797,1822]" captionTargetBox="[153,1436,174,1777]" captionTargetId="figure@23.[151,1436,174,1786]" captionTargetPageId="23" captionText="FIGURES 27–29. Male genitalia of the “aepitus group” species. 27. Artines aepitus, Jusepín, Monagas, Venezuela (OM 29.417). 28. Artines panama sp. nov., holotype. 29. Artines solange sp. nov., holotype. A) lateral view of tegumen, saccus, uncus and gnathos. B) ventral view of saccus. C) dorsal view of tegumen, uncus and gnathos. D) ventral view of tegumen, uncus and gnathos. E) interior view of right valva. F) dorsal view of right valva. G) dorsal view of aedeagus and vesica. H) left lateral view of aedeagus. I) ventral view of aedeagus. J) right lateral view of aedeagus. K) dorsal view of fultura inferior. L) lateral view of fultura inferior. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241047/files/figure.png" pageId="11" pageNumber="12">Fig. 28C</figureCitation>
). Gnathos as in the other species of the group (
<figureCitation id="DBC0FBA89D51CA46FF0CFCC7FE73FCC8" box="[231,400,836,863]" captionStart="FIGURES 27–29" captionStartId="23.[151,264,1797,1822]" captionTargetBox="[153,1436,174,1777]" captionTargetId="figure@23.[151,1436,174,1786]" captionTargetPageId="23" captionText="FIGURES 27–29. Male genitalia of the “aepitus group” species. 27. Artines aepitus, Jusepín, Monagas, Venezuela (OM 29.417). 28. Artines panama sp. nov., holotype. 29. Artines solange sp. nov., holotype. A) lateral view of tegumen, saccus, uncus and gnathos. B) ventral view of saccus. C) dorsal view of tegumen, uncus and gnathos. D) ventral view of tegumen, uncus and gnathos. E) interior view of right valva. F) dorsal view of right valva. G) dorsal view of aedeagus and vesica. H) left lateral view of aedeagus. I) ventral view of aedeagus. J) right lateral view of aedeagus. K) dorsal view of fultura inferior. L) lateral view of fultura inferior. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241047/files/figure.png" pageId="11" pageNumber="12">Figs 28A, 28D</figureCitation>
). Valva about three times longer than wide; sacculus triangular, more than 1/2 width of valva; costa narrow and not contiguous with ampulla; ampulla elongated, shorter than costa, distally directed ventrally; ventral process of harpe wide, slightly shorter than tegumen+uncus, distal 1/3 dilated and rounded, with apical setae thick and long (setae lost during preparation), in dorsal view narrow and slightly curved; dorsal process of harpe over median portion, triangular in dorsal view, narrower than ventral process (
<figureCitation id="DBC0FBA89D51CA46FBF2FC57FB8FFC78" box="[1049,1132,980,1007]" captionStart="FIGURES 27–29" captionStartId="23.[151,264,1797,1822]" captionTargetBox="[153,1436,174,1777]" captionTargetId="figure@23.[151,1436,174,1786]" captionTargetPageId="23" captionText="FIGURES 27–29. Male genitalia of the “aepitus group” species. 27. Artines aepitus, Jusepín, Monagas, Venezuela (OM 29.417). 28. Artines panama sp. nov., holotype. 29. Artines solange sp. nov., holotype. A) lateral view of tegumen, saccus, uncus and gnathos. B) ventral view of saccus. C) dorsal view of tegumen, uncus and gnathos. D) ventral view of tegumen, uncus and gnathos. E) interior view of right valva. F) dorsal view of right valva. G) dorsal view of aedeagus and vesica. H) left lateral view of aedeagus. I) ventral view of aedeagus. J) right lateral view of aedeagus. K) dorsal view of fultura inferior. L) lateral view of fultura inferior. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241047/files/figure.png" pageId="11" pageNumber="12">Fig. 28</figureCitation>
E–F). Aedeagus as long as valva, moderately curved dorsally; ventral projection distally wide and weakly curved ventrally (
<figureCitation id="DBC0FBA89D51CA46FB3DFC7BFACBFB84" box="[1238,1320,1016,1043]" captionStart="FIGURES 27–29" captionStartId="23.[151,264,1797,1822]" captionTargetBox="[153,1436,174,1777]" captionTargetId="figure@23.[151,1436,174,1786]" captionTargetPageId="23" captionText="FIGURES 27–29. Male genitalia of the “aepitus group” species. 27. Artines aepitus, Jusepín, Monagas, Venezuela (OM 29.417). 28. Artines panama sp. nov., holotype. 29. Artines solange sp. nov., holotype. A) lateral view of tegumen, saccus, uncus and gnathos. B) ventral view of saccus. C) dorsal view of tegumen, uncus and gnathos. D) ventral view of tegumen, uncus and gnathos. E) interior view of right valva. F) dorsal view of right valva. G) dorsal view of aedeagus and vesica. H) left lateral view of aedeagus. I) ventral view of aedeagus. J) right lateral view of aedeagus. K) dorsal view of fultura inferior. L) lateral view of fultura inferior. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241047/files/figure.png" pageId="11" pageNumber="12">Fig. 28</figureCitation>
G–J); posterior end of cornutus wide and triangular, with thin and straight spines (
<figureCitation id="DBC0FBA89D51CA46FC29FB9FFBF5FBA0" box="[962,1046,1052,1079]" captionStart="FIGURES 42–48" captionStartId="33.[151,264,1892,1917]" captionTargetBox="[151,1431,625,1874]" captionTargetId="figure@33.[151,1436,625,1880]" captionText="FIGURES 42–48. Dorsal view of vesica and cornutus of the &quot;aepitus group&quot; species. 42. Artines aepitus. 43. Artines panama sp. nov.. 44. Artines delfos sp. nov.. 45. Artines litoralis sp. nov.. 46. Artines rica. 47. Artines liege sp. nov. 48. Artines bamba sp. nov. Scale bar = 0,2 mm." httpUri="https://zenodo.org/record/3406274/files/figure.png" pageId="11" pageNumber="12">Fig. 43</figureCitation>
).
</paragraph>
<paragraph id="4344E72D9D51CA46FF2CFBC3FE91FBE9" blockId="11.[151,1437,152,2015]" pageId="11" pageNumber="12">
Female (
<figureCitation id="DBC0FBA89D51CA46FEC2FBC3FE88FBCC" box="[297,363,1088,1115]" captionStart="FIGURES 5–8" captionStartId="6.[151,264,1064,1089]" captionTargetBox="[154,1432,269,1044]" captionTargetId="figure@6.[151,1436,265,1052]" captionTargetPageId="6" captionText="FIGURES 5–8. Species of the “aepitus group” in dorsal and ventral views. 5. Artines aepitus, A–B: male, Cueva de Las Pavas, Tingo Maria, Huánuco, Peru (OM 80.822), C–D: female, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.790). 6. Artines panama sp. nov., A–B: holotype male, C–D: allotype female. 7. Artines solange sp. nov., A–B: holotype male, C–D: tentative female, Sumaúma, Maranhão, Brazil (DZ 41.800). 8. Artines delfos sp. nov., A–B: holotype male. Scale bar = 1 cm." httpUri="https://zenodo.org/record/3241041/files/figure.png" pageId="11" pageNumber="12">Fig. 6</figureCitation>
C–D).
<emphasis id="718F3B3F9D51CA46FE5CFBC2FD93FBCD" box="[439,624,1088,1114]" italics="true" pageId="11" pageNumber="12">Forewing length</emphasis>
:
<quantity id="84034AC89D51CA46FD94FBC2FD18FBCC" box="[639,763,1089,1115]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="11" pageNumber="12" unit="mm" value="12.5" valueMax="13.0" valueMin="12.0">12–13 mm</quantity>
. Similar to male, except for the absence of spot in CuA
<subScript id="DF7FE5689D51CA46FA8EFBD1FA8EFBF7" attach="left" box="[1381,1389,1106,1120]" fontSize="6" pageId="11" pageNumber="12">2</subScript>
–2A on dorsal forewing.
</paragraph>
<paragraph id="4344E72D9D51CA46FF2CFB0BFB7FFA99" blockId="11.[151,1437,152,2015]" pageId="11" pageNumber="12">
<emphasis id="718F3B3F9D51CA46FF2CFB0BFED2FB35" box="[199,305,1160,1186]" italics="true" pageId="11" pageNumber="12">Genitalia</emphasis>
(
<figureCitation id="DBC0FBA89D51CA46FED4FB0BFE70FB34" box="[319,403,1160,1187]" captionStart="FIGURES 54–60" captionStartId="35.[151,264,1807,1832]" captionTargetBox="[156,1436,575,1772]" captionTargetId="figure@35.[151,1436,589,1795]" captionTargetPageId="35" captionText="FIGURES 54–60. Female genitalia of the “aepitus group” species. 54. Artines aepitus, Jusepín, Monagas, Venezuela (OM 30. 125). 55. Artines panama sp. nov., allotype. 56. Artines solange sp. nov. (tentative female), Sumaúma, Maranhão, Brazil (DZ 41.800). 57. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.820). 58. Artines rica, Turrialba, Cartago, Costa Rica (DZ 41.840). 59. Artines liege sp. nov., paratype, Parque do Gama, Brasília, Distrito Federal, Brazil (DZ 41.830). 60. Artines bamba sp. nov. (tentative female), Moyobamba, San Martin, Peru (OM 81.229). A) lateral view of tergum VIII, sterigma and papilla analis; B) ventral view of esterigma. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241059/files/figure.png" pageId="11" pageNumber="12">Figs 55</figureCitation>
A–B, 67): tergum VIII with spiracular opening incomplete. Sterigma rectangular; lamella antevaginalis with proximo-ventral folds more or less rectangular; lamella postvaginalis without lateral fold and with distal projections narrow, separated medially by a relatively deep indentation, proximo-lateral projections weakly developed and not bilobed. Ostium as in
<taxonomicName id="84FB9CAE9D51CA46FD89FB76FD31FA99" baseAuthorityName="Geyer" baseAuthorityYear="1832" box="[610,722,1269,1295]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D51CA46FD89FB76FD31FA99" box="[610,722,1269,1295]" italics="true" pageId="11" pageNumber="12">A. aepitus</emphasis>
</taxonomicName>
; ductus bursae without sclerotized plate.
</paragraph>
<paragraph id="4344E72D9D51CA46FF2CFA9BFD39FAA5" blockId="11.[151,1437,152,2015]" box="[199,730,1304,1331]" pageId="11" pageNumber="12">
<emphasis id="718F3B3F9D51CA46FF2CFA9BFEDBFAA4" bold="true" box="[199,312,1304,1331]" pageId="11" pageNumber="12">Variation</emphasis>
. No relevant variation was observed.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D51CA46FF2CFABFFD2BF9C4" pageId="11" pageNumber="12" type="discussion">
<paragraph id="4344E72D9D51CA46FF2CFABFFA72F99C" blockId="11.[151,1437,152,2015]" pageId="11" pageNumber="12">
<emphasis id="718F3B3F9D51CA46FF2CFABFFEABFAC0" bold="true" box="[199,328,1340,1367]" pageId="11" pageNumber="12">Comments</emphasis>
. This species is well characterized by the male and female genitalia. It is also the only species of the genus that has been recorded from
<collectingCountry id="3BECA7BD9D51CA46FDA7FAE2FD46FAEC" box="[588,677,1377,1403]" name="Panama" pageId="11" pageNumber="12">Panama</collectingCountry>
(
<figureCitation id="DBC0FBA89D51CA46FD5EFAE3FCE9FAEC" box="[693,778,1376,1403]" captionStart="FIGURE 79" captionStartId="42.[151,250,1782,1807]" captionTargetBox="[151,1436,1021,1770]" captionTargetId="figure@42.[151,1436,1021,1770]" captionTargetPageId="42" captionText="FIGURE 79. Geographic distribution of the “aepitus group” species. The symbols with white inferior half represent records from literature." httpUri="https://zenodo.org/record/3241069/files/figure.png" pageId="11" pageNumber="12">Fig. 79</figureCitation>
). For this reason, all literature references to
<taxonomicName id="84FB9CAE9D51CA46FB17FAE2FAAEFAEC" authorityName="Godman" authorityYear="1901" box="[1276,1357,1377,1403]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="718F3B3F9D51CA46FB17FAE2FAAEFAEC" box="[1276,1357,1377,1403]" italics="true" pageId="11" pageNumber="12">Artines</emphasis>
</taxonomicName>
in that country are here interpreted as referring to
<taxonomicName id="84FB9CAE9D51CA46FD6EFA06FCE3FA09" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[645,768,1413,1439]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="panama" status="sp. nov.">
<emphasis id="718F3B3F9D51CA46FD6EFA06FCE3FA09" box="[645,768,1413,1439]" italics="true" pageId="11" pageNumber="12">A. panama</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D51CA46FCE2FA06FC87FA08" bold="true" box="[777,868,1413,1439]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="6ABC86449D51CA46FCE2FA06FC87FA08" box="[777,868,1413,1439]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<bibRefCitation id="276A9ADC9D51CA46FC9FFA07FBD9FA08" author="Huntington, E. I." box="[884,1082,1412,1439]" pageId="11" pageNumber="12" pagination="191 - 230" refId="ref41699" refString="Huntington, E. I. (1932) A list of the Rhopalocera of Barro Colorado Island, Canal Zone, Panama, with descriptions of two new species of Theclinae. Bulletin of the American Museum of Natural History, 63 (3), 191 - 230." type="journal article" year="1932">Huntington 1932</bibRefCitation>
;
<bibRefCitation id="276A9ADC9D51CA46FBACFA06FAFAFA08" author="Basset, Y. &amp; Barrios, H. &amp; Segar, S. &amp; Srygley, R. B. &amp; Aiello, A. &amp; Warren, A. D. &amp; Delgado, F. &amp; Coronado, J. &amp; Lezcano, J. &amp; Arizala, S. &amp; Rivera, M. &amp; Perez, F. &amp; Bobadilla, R. &amp; Lopez, Y. &amp; Ramirez, A. J." box="[1095,1305,1412,1439]" pageId="11" pageNumber="12" pagination="0136623" refId="ref38212" refString="Basset, Y., Barrios, H., Segar, S., Srygley, R. B., Aiello, A., Warren, A. D., Delgado, F., Coronado, J., Lezcano, J., Arizala, S., Rivera, M., Perez, F., Bobadilla, R., Lopez, Y. &amp; Ramirez, A. J. (2015) The butterflies of Barro Colorado Island, Panama: Local extinction since the 1930 s. PLoS ONE, 10 (8), e 0136623, 1 - 22. https: // doi. org / 10.1371 / journal. pone. 0136623" type="journal article" year="2015">
Basset
<emphasis id="718F3B3F9D51CA46FB72FA06FB30FA09" box="[1177,1235,1412,1438]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2015
</bibRefCitation>
). Given the great similarity with
<taxonomicName id="84FB9CAE9D51CA46FE68FA2AFE17FA55" baseAuthorityName="Geyer" baseAuthorityYear="1832" box="[387,500,1449,1475]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D51CA46FE68FA2AFE17FA55" box="[387,500,1449,1475]" italics="true" pageId="11" pageNumber="12">A. aepitus</emphasis>
</taxonomicName>
, particularly in the female genitalia,
<bibRefCitation id="276A9ADC9D51CA46FC7EFA2BFBABFA54" author="Godman, F. D." box="[917,1096,1448,1475]" pageId="11" pageNumber="12" pagination="142 - 457" refId="ref41039" refString="Godman, F. D. (1901) Hesperiidae [parte]. In: Godman, F. D. &amp; Salvin, O. (1879 - 1901), Biologia Centrali-Americana, Insecta, Lepidoptera Rhopalocera. Vol. 2. pp. 142 - 457, pls. 91 - 112]" type="journal article" year="1901">Godman (1901)</bibRefCitation>
included two specimens from
<collectingCountry id="3BECA7BD9D51CA46FF7CFA4EFF13FA70" box="[151,240,1485,1511]" name="Panama" pageId="11" pageNumber="12">Panama</collectingCountry>
in the description of
<taxonomicName id="84FB9CAE9D51CA46FE00FA4EFDB7FA71" authorityName="Godman" authorityYear="1901" box="[491,596,1485,1511]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="atizies">
<emphasis id="718F3B3F9D51CA46FE00FA4EFDB7FA71" box="[491,596,1485,1511]" italics="true" pageId="11" pageNumber="12">A. atizies</emphasis>
</taxonomicName>
. According to
<bibRefCitation id="276A9ADC9D51CA46FCEAFA4FFC56FA70" author="Godman, F. D." box="[769,949,1484,1511]" pageId="11" pageNumber="12" pagination="142 - 457" refId="ref41039" refString="Godman, F. D. (1901) Hesperiidae [parte]. In: Godman, F. D. &amp; Salvin, O. (1879 - 1901), Biologia Centrali-Americana, Insecta, Lepidoptera Rhopalocera. Vol. 2. pp. 142 - 457, pls. 91 - 112]" type="journal article" year="1901">Godman (1901)</bibRefCitation>
those specimens were not well preserved. Given that
<taxonomicName id="84FB9CAE9D51CA46FEFFFA72FE67F99D" baseAuthorityName="Geyer" baseAuthorityYear="1832" box="[276,388,1521,1547]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D51CA46FEFFFA72FE67F99D" box="[276,388,1521,1547]" italics="true" pageId="11" pageNumber="12">A. aepitus</emphasis>
</taxonomicName>
does not occur in Central America, those specimens probably belong to
<taxonomicName id="84FB9CAE9D51CA46FB5BFA72FACAF99D" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[1200,1321,1521,1547]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="panama" status="sp. nov.">
<emphasis id="718F3B3F9D51CA46FB5BFA72FACAF99D" box="[1200,1321,1521,1547]" italics="true" pageId="11" pageNumber="12">A. panama</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D51CA46FADBFA72FA69F99C" bold="true" box="[1328,1418,1521,1547]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="6ABC86449D51CA46FADBFA72FA69F99C" box="[1328,1418,1521,1547]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
.
</paragraph>
<paragraph id="4344E72D9D51CA46FF2CF997FD2BF9C4" blockId="11.[151,1437,152,2015]" pageId="11" pageNumber="12">
<emphasis id="718F3B3F9D51CA46FF2CF997FE0BF9B8" bold="true" box="[199,488,1556,1583]" pageId="11" pageNumber="12">Geographic distribution</emphasis>
(
<figureCitation id="DBC0FBA89D51CA46FE1CF997FDAFF9B8" box="[503,588,1556,1583]" captionStart="FIGURE 78" captionStartId="42.[151,250,910,935]" captionTargetBox="[151,1436,174,898]" captionTargetId="figure@42.[151,1436,174,898]" captionTargetPageId="42" captionText="FIGURE 78. Geographic distribution of the “aepitus group” species. The symbols with white inferior half represent records from literature." httpUri="https://zenodo.org/record/3241067/files/figure.png" pageId="11" pageNumber="12">Fig. 78</figureCitation>
). The species occurs in forested areas of
<collectingCountry id="3BECA7BD9D51CA46FBFCF996FB90F9B8" box="[1047,1139,1557,1583]" name="Panama" pageId="11" pageNumber="12">Panama</collectingCountry>
. PANAMA—
<emphasis id="718F3B3F9D51CA46FAFAF997FA77F9B9" box="[1297,1428,1556,1582]" italics="true" pageId="11" pageNumber="12">Canal Zone</emphasis>
: Farfan, Barro Colorado Island (
<bibRefCitation id="276A9ADC9D51CA46FE1CF9BBFD5FF9C4" author="Huntington, E. I." box="[503,700,1592,1619]" pageId="11" pageNumber="12" pagination="191 - 230" refId="ref41699" refString="Huntington, E. I. (1932) A list of the Rhopalocera of Barro Colorado Island, Canal Zone, Panama, with descriptions of two new species of Theclinae. Bulletin of the American Museum of Natural History, 63 (3), 191 - 230." type="journal article" year="1932">Huntington 1932</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D51CA46FF2CF9DEFE93F90C" pageId="11" pageNumber="12" type="description">
<paragraph id="4344E72D9D51CA46FF2CF9DEFE93F90C" blockId="11.[151,1437,152,2015]" pageId="11" pageNumber="12">
<emphasis id="718F3B3F9D51CA46FF2CF9DEFE30F9E0" bold="true" box="[199,467,1628,1655]" pageId="11" pageNumber="12">Temporal distribution</emphasis>
. This species occurs in January. There is also in the literature record for November (
<bibRefCitation id="276A9ADC9D51CA46FF74F903FE80F90C" author="Huntington, E. I." box="[159,355,1664,1691]" pageId="11" pageNumber="12" pagination="191 - 230" refId="ref41699" refString="Huntington, E. I. (1932) A list of the Rhopalocera of Barro Colorado Island, Canal Zone, Panama, with descriptions of two new species of Theclinae. Bulletin of the American Museum of Natural History, 63 (3), 191 - 230." type="journal article" year="1932">Huntington 1932</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D51CA46FF2CF926FE2AF929" box="[199,457,1700,1727]" pageId="11" pageNumber="12" type="biology_ecology">
<paragraph id="4344E72D9D51CA46FF2CF926FE2AF929" blockId="11.[151,1437,152,2015]" box="[199,457,1700,1727]" pageId="11" pageNumber="12">
<emphasis id="718F3B3F9D51CA46FF2CF926FEA7F928" bold="true" box="[199,324,1701,1727]" pageId="11" pageNumber="12">Host plant</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D51CA46FF2CF94AFC54F975" box="[199,951,1736,1763]" pageId="11" pageNumber="12" type="etymology">
<paragraph id="4344E72D9D51CA46FF2CF94AFC54F975" blockId="11.[151,1437,152,2015]" box="[199,951,1736,1763]" pageId="11" pageNumber="12">
<emphasis id="718F3B3F9D51CA46FF2CF94AFEABF974" bold="true" box="[199,328,1737,1763]" pageId="11" pageNumber="12">Etymology</emphasis>
. This species is named in reference to the
<typeStatus id="9C40598F9D51CA46FCCFF94AFCB7F974" box="[804,852,1737,1763]" pageId="11" pageNumber="12">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D51CA46FF2CF96EFC98F848" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="4344E72D9D51CA46FF2CF96EFD8EF8D8" blockId="11.[151,1437,152,2015]" pageId="11" pageNumber="12">
<emphasis id="718F3B3F9D51CA46FF2CF96EFE96F890" bold="true" box="[199,373,1772,1799]" pageId="11" pageNumber="12">Type material</emphasis>
.
<typeStatus id="9C40598F9D51CA46FE63F96FFE13F891" box="[392,496,1772,1798]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
male deposited at the OM–DZUP with the following labels: /
<materialsCitation id="F393ED709D51CA46FB09F96EFB56F8BC" ID-GBIF-Occurrence="2248478819" collectingDate="1968-01-13" collectionCode="OM" collectorName="Farfan, C &amp; Brown" country="Panama" location="Panama" pageId="11" pageNumber="12" specimenCode="OM 29.165" specimenCount="1" typeStatus="holotype">
<typeStatus id="9C40598F9D51CA46FB09F96EFA6BF890" box="[1250,1416,1773,1799]" pageId="11" pageNumber="12" type="holotype">HOLOTYPUS</typeStatus>
/
<date id="3745C1ED9D51CA46FF7CF893FEE7F8BD" box="[151,260,1808,1834]" pageId="11" pageNumber="12" value="1968-01-13">
<collectingDate id="270138059D51CA46FF7CF893FEE7F8BD" box="[151,260,1808,1834]" pageId="11" pageNumber="12" value="1968-01-13">13.I.1968</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D51CA46FEF9F893FE9DF8BC" box="[274,382,1808,1835]" pageId="11" pageNumber="12">Farfan, C</collectorName>
[anal] Zone,
<collectingCountry id="3BECA7BD9D51CA46FDFAF892FD8EF8BC" box="[529,621,1809,1835]" name="Panama" pageId="11" pageNumber="12">Panama</collectingCountry>
,
<collectorName id="EE0E82FB9D51CA46FD92F892FD26F8BC" box="[633,709,1809,1835]" pageId="11" pageNumber="12">Brown</collectorName>
<emphasis id="718F3B3F9D51CA46FD25F893FD13F8BD" box="[718,752,1808,1834]" italics="true" pageId="11" pageNumber="12">leg</emphasis>
. / gen. prep. Mielke 1992 /
<specimenCode id="135D4F569D51CA46FBD9F892FB56F8BC" box="[1074,1205,1808,1835]" collectionCode="OM" country="New Zealand" name="Otago Museum" pageId="11" pageNumber="12">OM 29.165</specimenCode>
</materialsCitation>
/
<typeStatus id="9C40598F9D51CA46FB25F893FAA0F8BD" box="[1230,1347,1808,1834]" pageId="11" pageNumber="12" type="holotype">Holotypus</typeStatus>
<taxonomicName id="84FB9CAE9D51CA46FAA0F892FE0BF8D9" authority="Medeiros &amp; Carneiro" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="panama">
<emphasis id="718F3B3F9D51CA46FAA0F892FF12F8D9" italics="true" pageId="11" pageNumber="12">Artines panama</emphasis>
Medeiros &amp; Carneiro
</taxonomicName>
det. 2018 /.
</paragraph>
<paragraph id="4344E72D9D51CA46FF2CF8DBFD26F82C" blockId="11.[151,1437,152,2015]" pageId="11" pageNumber="12">
<typeStatus id="9C40598F9D51CA46FF2CF8DBFECAF8E5" box="[199,297,1880,1906]" pageId="11" pageNumber="12" type="allotype">Allotype</typeStatus>
female deposited at the OM–DZUP with the following labels: /
<materialsCitation id="F393ED709D51CA46FBF8F8DAFAF9F800" ID-GBIF-Occurrence="2248478830" collectingDate="1968-01-13" collectionCode="OM" collectorName="Farfan, C &amp; Brown &amp; A. Medeiros" country="Panama" location="Panama" pageId="11" pageNumber="12" specimenCode="OM 30.179" specimenCount="1" typeStatus="allotype">
<typeStatus id="9C40598F9D51CA46FBF8F8DAFB55F8E4" box="[1043,1206,1881,1907]" pageId="11" pageNumber="12" type="allotype">ALLOTYPUS</typeStatus>
/
<date id="3745C1ED9D51CA46FB3BF8DBFADDF8E5" box="[1232,1342,1880,1906]" pageId="11" pageNumber="12" value="1968-01-13">
<collectingDate id="270138059D51CA46FB3BF8DBFADDF8E5" box="[1232,1342,1880,1906]" pageId="11" pageNumber="12" value="1968-01-13">13.I.1968</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D51CA46FAA7F8DBFF49F800" pageId="11" pageNumber="12">Farfan, C</collectorName>
[anal] Zone,
<collectingCountry id="3BECA7BD9D51CA46FED2F8FEFE76F800" box="[313,405,1917,1943]" name="Panama" pageId="11" pageNumber="12">Panama</collectingCountry>
,
<collectorName id="EE0E82FB9D51CA46FE74F8FEFE08F800" box="[415,491,1917,1943]" pageId="11" pageNumber="12">Brown</collectorName>
<emphasis id="718F3B3F9D51CA46FE19F8FFFDF7F801" box="[498,532,1916,1942]" italics="true" pageId="11" pageNumber="12">leg</emphasis>
. / gen. prep. Mielke 1992 / gen. prep.
<collectorName id="EE0E82FB9D51CA46FC50F8FEFBA4F801" box="[955,1095,1916,1943]" pageId="11" pageNumber="12">A. Medeiros</collectorName>
2018 /
<specimenCode id="135D4F569D51CA46FB71F8FEFAF9F800" box="[1178,1306,1917,1943]" collectionCode="OM" country="New Zealand" name="Otago Museum" pageId="11" pageNumber="12">OM 30.179</specimenCode>
</materialsCitation>
/
<typeStatus id="9C40598F9D51CA46FAC5F8FFFA7FF801" box="[1326,1436,1916,1942]" pageId="11" pageNumber="12" type="allotype">Allotypus</typeStatus>
<taxonomicName id="84FB9CAE9D51CA46FF7CF822FDA3F82D" authority="Medeiros &amp; Carneiro" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[151,576,1952,1979]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="panama">
<emphasis id="718F3B3F9D51CA46FF7CF822FEAAF82D" box="[151,329,1953,1979]" italics="true" pageId="11" pageNumber="12">Artines panama</emphasis>
Medeiros &amp; Carneiro
</taxonomicName>
det. 2018 /.
</paragraph>
<paragraph id="4344E72D9D51CA46FF2CF846FC98F848" blockId="11.[151,1437,152,2015]" box="[199,891,1988,2015]" pageId="11" pageNumber="12">
<materialsCitation id="F393ED709D51CA46FF2CF846FC94F848" ID-GBIF-Occurrence="2248478885" box="[199,887,1988,2015]" collectingDate="1968-01-13" collectionCode="OM" collectorName="Farfan, C &amp; Brown" country="Panama" location="Panama" pageId="11" pageNumber="12" specimenCode="OM 30.083" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="9C40598F9D51CA46FF2CF846FECEF848" box="[199,301,1989,2015]" pageId="11" pageNumber="12" type="paratype">Paratype</typeStatus>
:
<specimenCount id="55FD2CA49D51CA46FED3F846FE7FF849" box="[312,412,1988,2015]" pageId="11" pageNumber="12" type="female">1 female</specimenCount>
, same data as holotype,
<specimenCode id="135D4F569D51CA46FD47F846FCCEF848" box="[684,813,1989,2015]" collectionCode="OM" country="New Zealand" name="Otago Museum" pageId="11" pageNumber="12">OM 30.083</specimenCode>
(
<collectionCode id="25EA7FE89D51CA46FCD7F846FC8CF848" box="[828,879,1989,2015]" country="New Zealand" name="Otago Museum" pageId="11" pageNumber="12">OM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

788
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<subSubSection id="0BE1B4A69D54CA43FF7CFB0AFD35FB34" box="[151,726,1160,1187]" pageId="14" pageNumber="15" type="nomenclature">
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<emphasis id="718F3B3F9D54CA43FF7CFB0AFD35FB34" bold="true" box="[151,726,1160,1187]" pageId="14" pageNumber="15">
<taxonomicName id="84FB9CAE9D54CA43FF7CFB0AFD92FB35" authority="Medeiros &amp; Dolibaina" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[151,625,1160,1187]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="litoralis" status="sp. nov.">
<emphasis id="718F3B3F9D54CA43FF7CFB0AFEB7FB34" bold="true" box="[151,340,1160,1187]" italics="true" pageId="14" pageNumber="15">Artines litoralis</emphasis>
Medeiros &amp; Dolibaina
</taxonomicName>
<taxonomicNameLabel id="6ABC86449D54CA43FD93FB0AFD35FB34" box="[632,726,1161,1187]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D54CA43FF7CFB2FFBD5FAF6" pageId="14" pageNumber="15" type="description">
<paragraph id="4344E72D9D54CA43FF7CFB2FFE5FFB50" blockId="14.[151,945,1160,1258]" box="[151,444,1196,1223]" pageId="14" pageNumber="15">
<figureCitation id="DBC0FBA89D54CA43FF7CFB2FFF0FFB50" box="[151,236,1196,1223]" captionStart="FIGURES 9–13" captionStartId="7.[151,264,1180,1205]" captionTargetBox="[152,1436,180,1162]" captionTargetId="figure@7.[151,1436,174,1169]" captionTargetPageId="7" captionText="FIGURES 9–13. Species of the “aepitus group” in dorsal and ventral views. 9. Artines maya, A–B: holotype male. 10. Artines litoralis sp. nov., A–B: holotype male, C–D: allotype female. 11. Artines rica, A–B: male, Turrialba, Cartago, Costa Rica (DZ 41.890), C–D: female from same locality of male (DZ 41.840). 12. Artines liege sp. nov., A–B: holotype male, C–D: allotype female. 13. Artines bamba sp. nov., A–B: holotype male, C–D: tentative female, Moyobamba, San Martin, Peru (OM 79.413). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241043/files/figure.png" pageId="14" pageNumber="15">Figs 10</figureCitation>
,
<figureCitation id="DBC0FBA89D54CA43FF13FB2EFEF5FB50" box="[248,278,1197,1223]" captionStart="FIGURES 30–32" captionStartId="24.[151,264,1772,1797]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure@24.[151,1436,181,1746]" captionTargetPageId="24" captionText="FIGURES 30–32. Male genitalia of the “aepitus group” species. 30. Artines delfos sp. nov., holotype. 31. Artines maya, holotype. 32. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.215). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241049/files/figure.png" pageId="14" pageNumber="15">32</figureCitation>
,
<figureCitation id="DBC0FBA89D54CA43FEC9FB2FFEA3FB50" box="[290,320,1196,1223]" captionStart="FIGURES 42–48" captionStartId="33.[151,264,1892,1917]" captionTargetBox="[151,1431,625,1874]" captionTargetId="figure@33.[151,1436,625,1880]" captionText="FIGURES 42–48. Dorsal view of vesica and cornutus of the &quot;aepitus group&quot; species. 42. Artines aepitus. 43. Artines panama sp. nov.. 44. Artines delfos sp. nov.. 45. Artines litoralis sp. nov.. 46. Artines rica. 47. Artines liege sp. nov. 48. Artines bamba sp. nov. Scale bar = 0,2 mm." httpUri="https://zenodo.org/record/3406274/files/figure.png" pageId="14" pageNumber="15">45</figureCitation>
, 57, 69, 79
</paragraph>
<paragraph id="4344E72D9D54CA43FF7CFB53FC52FB7D" blockId="14.[151,945,1160,1258]" box="[151,945,1232,1258]" pageId="14" pageNumber="15">
<uri id="376AEB2F9D54CA43FF7CFB53FC52FB7D" box="[151,945,1232,1258]" pageId="14" pageNumber="15">
urn:lsid:zoobank.org:act:
<uuid id="375DDDF89D54CA43FE43FB53FC52FB7D" box="[424,945,1232,1258]" pageId="14" pageNumber="15">29D87F1A-B981-411A-B007-873037944723</uuid>
</uri>
</paragraph>
<paragraph id="4344E72D9D54CA43FF7CFAA9FC6DFAD5" blockId="14.[151,1436,1321,1438]" box="[151,910,1321,1346]" pageId="14" pageNumber="15">
<taxonomicName id="84FB9CAE9D54CA43FF7CFAA9FECBFAD5" authorityName="Godman" authorityYear="1901" box="[151,296,1322,1346]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="atizies">
<emphasis id="718F3B3F9D54CA43FF7CFAA9FECBFAD5" box="[151,296,1322,1346]" italics="true" pageId="14" pageNumber="15">Artines atizies</emphasis>
</taxonomicName>
(misid.);
<bibRefCitation id="276A9ADC9D54CA43FE60FAAAFDC1FAD5" author="Cardoso, A. S." box="[395,546,1321,1346]" pageId="14" pageNumber="15" pagination="427 - 436" refId="ref39169" refString="Cardoso, A. S. (1949) Lepidopteros de Alagoas. Revista de Entomologia, 20 (1 / 3), 427 - 436." type="journal article" year="1949">Cardoso, 1949</bibRefCitation>
.
<emphasis id="718F3B3F9D54CA43FDC0FAA9FD6CFAD5" bold="true" box="[555,655,1322,1346]" pageId="14" pageNumber="15">Rev. Ent.</emphasis>
,
<collectingRegion id="813F29CF9D54CA43FD70FAAAFCD3FAD5" box="[667,816,1321,1346]" country="Brazil" name="Rio de Janeiro" pageId="14" pageNumber="15">Rio de Janeiro</collectingRegion>
,
<emphasis id="718F3B3F9D54CA43FCD1FAAAFCB0FAD5" bold="true" box="[826,851,1321,1346]" pageId="14" pageNumber="15">20</emphasis>
: 432.
</paragraph>
<paragraph id="4344E72D9D54CA43FF7CFACAFBD5FAF6" blockId="14.[151,1436,1321,1438]" box="[151,1078,1352,1377]" pageId="14" pageNumber="15">
<taxonomicName id="84FB9CAE9D54CA43FF7CFACAFE9CFAF7" authority="aepitus" box="[151,383,1353,1376]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="subSpecies" species="aepitus" subSpecies="aepitus">
<emphasis id="718F3B3F9D54CA43FF7CFACAFE9CFAF7" box="[151,383,1353,1376]" italics="true" pageId="14" pageNumber="15">Artines aepitus aepitus</emphasis>
</taxonomicName>
(misid.); Kesselring &amp; Ebert, [1982].
<emphasis id="718F3B3F9D54CA43FCE8FACAFC1FFAF6" bold="true" box="[771,1020,1352,1377]" pageId="14" pageNumber="15">Rev. Nordestina Biol. 2</emphasis>
: 108.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D54CA43FF7CFAEBFF06F9B8" pageId="14" pageNumber="15" type="reference_group">
<paragraph id="4344E72D9D54CA43FF7CFAEBFC05FA09" blockId="14.[151,1436,1321,1438]" pageId="14" pageNumber="15">
<taxonomicName id="84FB9CAE9D54CA43FF7CFAEBFECCFAE8" ID-CoL="GZYC" baseAuthorityName="Geyer" baseAuthorityYear="1832" box="[151,303,1384,1407]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D54CA43FF7CFAEBFECCFAE8" box="[151,303,1384,1407]" italics="true" pageId="14" pageNumber="15">Artines aepitus</emphasis>
</taxonomicName>
(misid.); K.
<bibRefCitation id="276A9ADC9D54CA43FE5BFAEBFD7FFAE8" author="Brown Jr., K. S. &amp; Freitas, A. V. L." box="[432,668,1383,1408]" pageId="14" pageNumber="15" pagination="71 - 116" refId="ref39087" refString="Brown Jr., K. S. &amp; Freitas, A. V. L. (2000) Diversidade de Lepidoptera em Santa Teresa, Espirito Santo. Boletim do Museu de Biologia Mello Leitao, New Series, 11 / 12, 71 - 116" type="journal article" year="2000">Brown &amp; Freitas, 2000</bibRefCitation>
.
<emphasis id="718F3B3F9D54CA43FD4EFAEBFBE1FA17" bold="true" box="[677,1026,1383,1408]" pageId="14" pageNumber="15">Bol. Mus. Biol. Mello Leitão, n. s</emphasis>
., Sta. Teresa,
<emphasis id="718F3B3F9D54CA43FB64FAE4FB24FA17" bold="true" box="[1167,1223,1383,1408]" pageId="14" pageNumber="15">11/12</emphasis>
: 112.—
<bibRefCitation id="276A9ADC9D54CA43FAFDFAE4FF1FFA09" author="Freitas, A. V. L. &amp; Brown Jr., K. S. &amp; Mielke, O. H. H. &amp; Santos, J. P. &amp; Vasconcellos-Neto, J." pageId="14" pageNumber="15" pagination="317 - 328" refId="ref40575" refString="Freitas, A. V. L., Brown Jr., K. S., Mielke, O. H. H., Santos, J. P. &amp; Vasconcellos-Neto, J. (2016) Borboletas da Reserva Natural Vale, Linhares / ES. In: Rolim, S. G., Menezes, L. F. T. &amp; Srbek-Araujo, A. C. (Eds.), Floresta atlantica de tabuleiro: Diversidade e endemismos na Reserva Natural Vale. Editora Rupestre, Belo Horizonte, pp. 317 - 328." type="book chapter" year="2016">
Freitas
<emphasis id="718F3B3F9D54CA43FA8FFAEBFA73FA17" bold="true" box="[1380,1424,1383,1408]" italics="true" pageId="14" pageNumber="15">et al</emphasis>
., 2016
</bibRefCitation>
,
<emphasis id="718F3B3F9D54CA43FEECFA05FEFFFA09" bold="true" box="[263,284,1414,1438]" italics="true" pageId="14" pageNumber="15">in</emphasis>
Rolim; Menezes &amp; Srbek-Araújo.
<emphasis id="718F3B3F9D54CA43FD6BFA05FC71FA09" bold="true" box="[640,914,1413,1438]" pageId="14" pageNumber="15">Floresta Atl. de Tabuleiro</emphasis>
, p. 323.
</paragraph>
<paragraph id="4344E72D9D54CA43FF7CFA4FFF06F9B8" blockId="14.[151,1437,1484,2014]" pageId="14" pageNumber="15">
<emphasis id="718F3B3F9D54CA43FF7CFA4FFE95FA70" bold="true" box="[151,374,1484,1511]" pageId="14" pageNumber="15">Taxonomic history</emphasis>
. This species was previously treated as
<taxonomicName id="84FB9CAE9D54CA43FCC7FA4EFBA7FA70" ID-CoL="67SFD" authority="(Cardoso 1949)" baseAuthorityName="Cardoso" baseAuthorityYear="1949" box="[812,1092,1484,1511]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="atizies">
<emphasis id="718F3B3F9D54CA43FCC7FA4EFC71FA71" box="[812,914,1485,1511]" italics="true" pageId="14" pageNumber="15">A. atizies</emphasis>
(
<bibRefCitation id="276A9ADC9D54CA43FC75FA4FFBDFFA70" author="Cardoso, A. S." box="[926,1084,1484,1511]" pageId="14" pageNumber="15" pagination="427 - 436" refId="ref39169" refString="Cardoso, A. S. (1949) Lepidopteros de Alagoas. Revista de Entomologia, 20 (1 / 3), 427 - 436." type="journal article" year="1949">Cardoso 1949</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="84FB9CAE9D54CA43FBBAFA4EFAF5FA71" ID-CoL="GZYC" authority="aepitus" box="[1105,1302,1485,1511]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="subSpecies" species="aepitus" subSpecies="aepitus">
<emphasis id="718F3B3F9D54CA43FBBAFA4EFAF5FA71" box="[1105,1302,1485,1511]" italics="true" pageId="14" pageNumber="15">A. aepitus aepitus</emphasis>
</taxonomicName>
(
<bibRefCitation id="276A9ADC9D54CA43FAC8FA4FFEA6F99C" author="Kesselring, J. &amp; Ebert, H." pageId="14" pageNumber="15" pagination="105 - 118" refId="ref42042" refString="Kesselring, J. &amp; Ebert, H. (1982) Relacao das borboletas encontradas na &quot; Mata do Buraquinho &quot;, Joao Pessoa, Estado da Paraiba, Brasil. Revista Nordestina de Biologia, 2 (1 / 2), 105 - 118." type="journal article" year="1982">Kesselring &amp; Ebert [1982]</bibRefCitation>
) and
<taxonomicName id="84FB9CAE9D54CA43FE6EFA72FE15F99D" ID-CoL="GZYC" baseAuthorityName="Geyer" baseAuthorityYear="1832" box="[389,502,1521,1547]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D54CA43FE6EFA72FE15F99D" box="[389,502,1521,1547]" italics="true" pageId="14" pageNumber="15">A. aepitus</emphasis>
</taxonomicName>
(
<bibRefCitation id="276A9ADC9D54CA43FDEEFA72FCC7F99C" author="Brown Jr., K. S. &amp; Freitas, A. V. L." box="[517,804,1520,1547]" pageId="14" pageNumber="15" pagination="71 - 116" refId="ref39087" refString="Brown Jr., K. S. &amp; Freitas, A. V. L. (2000) Diversidade de Lepidoptera em Santa Teresa, Espirito Santo. Boletim do Museu de Biologia Mello Leitao, New Series, 11 / 12, 71 - 116" type="journal article" year="2000">Brown Jr &amp; Freitas 2000</bibRefCitation>
;
<bibRefCitation id="276A9ADC9D54CA43FCC4FA73FC1DF99C" author="Freitas, A. V. L. &amp; Brown Jr., K. S. &amp; Mielke, O. H. H. &amp; Santos, J. P. &amp; Vasconcellos-Neto, J." box="[815,1022,1520,1547]" pageId="14" pageNumber="15" pagination="317 - 328" refId="ref40575" refString="Freitas, A. V. L., Brown Jr., K. S., Mielke, O. H. H., Santos, J. P. &amp; Vasconcellos-Neto, J. (2016) Borboletas da Reserva Natural Vale, Linhares / ES. In: Rolim, S. G., Menezes, L. F. T. &amp; Srbek-Araujo, A. C. (Eds.), Floresta atlantica de tabuleiro: Diversidade e endemismos na Reserva Natural Vale. Editora Rupestre, Belo Horizonte, pp. 317 - 328." type="book chapter" year="2016">
Freitas
<emphasis id="718F3B3F9D54CA43FC68FA72FC5FF99D" box="[899,956,1520,1546]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
2016
</bibRefCitation>
) in studies conducted in the Atlantic Forest.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D54CA43FF2CF9BBFCBDF928" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="4344E72D9D54CA43FF2CF9BBFCBDF928" blockId="14.[151,1437,1484,2014]" pageId="14" pageNumber="15">
<emphasis id="718F3B3F9D54CA43FF2CF9BBFED9F9C4" bold="true" box="[199,314,1592,1619]" pageId="14" pageNumber="15">Diagnosis</emphasis>
.
<taxonomicName id="84FB9CAE9D54CA43FEACF9BAFE14F9C5" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[327,503,1592,1619]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="litoralis" status="sp. nov.">
<emphasis id="718F3B3F9D54CA43FEACF9BAFE14F9C5" box="[327,503,1592,1619]" italics="true" pageId="14" pageNumber="15">Artines litoralis</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D54CA43FE16F9BAFDB4F9C4" bold="true" box="[509,599,1593,1619]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="6ABC86449D54CA43FE16F9BAFDB4F9C4" box="[509,599,1593,1619]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is externally similar to the other species of the “
<taxonomicName id="84FB9CAE9D54CA43FB86F9BAFB5DF9C5" baseAuthorityName="Geyer" baseAuthorityYear="1832" box="[1133,1214,1593,1618]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D54CA43FB86F9BAFB5DF9C5" box="[1133,1214,1593,1618]" italics="true" pageId="14" pageNumber="15">aepitus</emphasis>
</taxonomicName>
group”, differing in the following combination of characters: harpe with ventral process triangular and shorter than tegumen; aedeagus longer than valva; proximo-lateral projections of lamella postvaginalis triangular and not bilobed; occurrence restricted to the Brazilian coast, from
<collectingRegion id="813F29CF9D54CA43FDD4F927FD70F929" box="[575,659,1700,1726]" country="Brazil" name="Paraiba" pageId="14" pageNumber="15">Paraíba</collectingRegion>
to
<collectingRegion id="813F29CF9D54CA43FD5CF927FCBAF928" box="[695,857,1700,1727]" country="Brazil" name="Espirito Santo" pageId="14" pageNumber="15">Espírito Santo</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D54CA42FF2CF94BFEA6FCC8" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="description">
<paragraph id="4344E72D9D54CA43FF2CF94BFBD3F974" blockId="14.[151,1437,1484,2014]" box="[199,1072,1736,1763]" pageId="14" pageNumber="15">
<emphasis id="718F3B3F9D54CA43FF2CF94BFEB2F974" bold="true" box="[199,337,1736,1763]" pageId="14" pageNumber="15">Description</emphasis>
. Male (10A–B).
<emphasis id="718F3B3F9D54CA43FDF9F94BFDACF975" box="[530,591,1736,1762]" italics="true" pageId="14" pageNumber="15">Head</emphasis>
: antenna with nudum bearing12 segments.
</paragraph>
<paragraph id="4344E72D9D54CA43FF2CF96EFDF3F890" blockId="14.[151,1437,1484,2014]" box="[199,528,1772,1799]" pageId="14" pageNumber="15">
<emphasis id="718F3B3F9D54CA43FF2CF96EFE62F891" box="[199,385,1772,1798]" italics="true" pageId="14" pageNumber="15">Forewing length</emphasis>
:
<quantity id="84034AC89D54CA43FE7BF96FFDEFF890" box="[400,524,1772,1799]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="14" pageNumber="15" unit="mm" value="13.5" valueMax="15.0" valueMin="12.0">12–15 mm</quantity>
.
</paragraph>
<paragraph id="4344E72D9D54CA43FF2CF893FDDDF8D9" blockId="14.[151,1437,1484,2014]" pageId="14" pageNumber="15">
<emphasis id="718F3B3F9D54CA43FF2CF893FE61F8BC" box="[199,386,1808,1835]" italics="true" pageId="14" pageNumber="15">Dorsal forewing</emphasis>
(
<figureCitation id="DBC0FBA89D54CA43FE7FF893FE1CF8BC" box="[404,511,1808,1835]" captionStart="FIGURES 9–13" captionStartId="7.[151,264,1180,1205]" captionTargetBox="[152,1436,180,1162]" captionTargetId="figure@7.[151,1436,174,1169]" captionTargetPageId="7" captionText="FIGURES 9–13. Species of the “aepitus group” in dorsal and ventral views. 9. Artines maya, A–B: holotype male. 10. Artines litoralis sp. nov., A–B: holotype male, C–D: allotype female. 11. Artines rica, A–B: male, Turrialba, Cartago, Costa Rica (DZ 41.890), C–D: female from same locality of male (DZ 41.840). 12. Artines liege sp. nov., A–B: holotype male, C–D: allotype female. 13. Artines bamba sp. nov., A–B: holotype male, C–D: tentative female, Moyobamba, San Martin, Peru (OM 79.413). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241043/files/figure.png" pageId="14" pageNumber="15">Fig. 10A</figureCitation>
): apical spots generally absent; discal spots reduced or absent; spot in CuA
<subScript id="DF7FE5689D54CA43FA8FF8A1FA8FF8A7" attach="left" box="[1380,1388,1826,1840]" fontSize="6" pageId="14" pageNumber="15">2</subScript>
–2A absent or represented by a few scales.
</paragraph>
<paragraph id="4344E72D9D54CA43FF2CF8DBFF12F801" blockId="14.[151,1437,1484,2014]" pageId="14" pageNumber="15">
<emphasis id="718F3B3F9D54CA43FF2CF8DBFE64F8E4" box="[199,391,1880,1907]" italics="true" pageId="14" pageNumber="15">Ventral forewing</emphasis>
(
<figureCitation id="DBC0FBA89D54CA43FE7CF8DBFE1CF8E4" box="[407,511,1880,1907]" captionStart="FIGURES 9–13" captionStartId="7.[151,264,1180,1205]" captionTargetBox="[152,1436,180,1162]" captionTargetId="figure@7.[151,1436,174,1169]" captionTargetPageId="7" captionText="FIGURES 9–13. Species of the “aepitus group” in dorsal and ventral views. 9. Artines maya, A–B: holotype male. 10. Artines litoralis sp. nov., A–B: holotype male, C–D: allotype female. 11. Artines rica, A–B: male, Turrialba, Cartago, Costa Rica (DZ 41.890), C–D: female from same locality of male (DZ 41.840). 12. Artines liege sp. nov., A–B: holotype male, C–D: allotype female. 13. Artines bamba sp. nov., A–B: holotype male, C–D: tentative female, Moyobamba, San Martin, Peru (OM 79.413). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241043/files/figure.png" pageId="14" pageNumber="15">Fig. 10B</figureCitation>
): three reduced blue to white apical spots; discal spots as in the other species, but smaller.
</paragraph>
<paragraph id="4344E72D9D54CA43FF2CF823FEADF849" blockId="14.[151,1437,1484,2014]" pageId="14" pageNumber="15">
<emphasis id="718F3B3F9D54CA43FF2CF823FE6DF82D" box="[199,398,1952,1978]" italics="true" pageId="14" pageNumber="15">Ventral hind wing</emphasis>
(
<figureCitation id="DBC0FBA89D54CA43FE76F823FDE1F82C" box="[413,514,1952,1979]" captionStart="FIGURES 9–13" captionStartId="7.[151,264,1180,1205]" captionTargetBox="[152,1436,180,1162]" captionTargetId="figure@7.[151,1436,174,1169]" captionTargetPageId="7" captionText="FIGURES 9–13. Species of the “aepitus group” in dorsal and ventral views. 9. Artines maya, A–B: holotype male. 10. Artines litoralis sp. nov., A–B: holotype male, C–D: allotype female. 11. Artines rica, A–B: male, Turrialba, Cartago, Costa Rica (DZ 41.890), C–D: female from same locality of male (DZ 41.840). 12. Artines liege sp. nov., A–B: holotype male, C–D: allotype female. 13. Artines bamba sp. nov., A–B: holotype male, C–D: tentative female, Moyobamba, San Martin, Peru (OM 79.413). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241043/files/figure.png" pageId="14" pageNumber="15">Fig. 10B</figureCitation>
): apical portion of discal cell with few yellow scales; central black spot rectangular, wider than long.
</paragraph>
<paragraph id="4344E72D9D55CA42FF2CFF1AFD95FD8C" blockId="15.[151,1437,152,2015]" pageId="15" pageNumber="16">
<emphasis id="718F3B3F9D55CA42FF2CFF1AFED2FF24" box="[199,305,153,179]" italics="true" pageId="15" pageNumber="16">Genitalia</emphasis>
(
<figureCitation id="DBC0FBA89D55CA42FEAAFF1AFE74FF24" box="[321,407,153,179]" captionStart="FIGURES 30–32" captionStartId="24.[151,264,1772,1797]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure@24.[151,1436,181,1746]" captionTargetPageId="24" captionText="FIGURES 30–32. Male genitalia of the “aepitus group” species. 30. Artines delfos sp. nov., holotype. 31. Artines maya, holotype. 32. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.215). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241049/files/figure.png" pageId="15" pageNumber="16">Figs 32</figureCitation>
A–L, 45): tegumen more or less quadrate, with proximal margin straight and distal margin convex, slightly rounded (
<figureCitation id="DBC0FBA89D55CA42FE56FF3EFDC7FF40" box="[445,548,189,215]" captionStart="FIGURES 30–32" captionStartId="24.[151,264,1772,1797]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure@24.[151,1436,181,1746]" captionTargetPageId="24" captionText="FIGURES 30–32. Male genitalia of the “aepitus group” species. 30. Artines delfos sp. nov., holotype. 31. Artines maya, holotype. 32. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.215). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241049/files/figure.png" pageId="15" pageNumber="16">Fig. 32C</figureCitation>
), in lateral view, shorter than length of gnathos; ventral arm of tegumen longer than dorsal arm of saccus (
<figureCitation id="DBC0FBA89D55CA42FE54FF62FDC5FF6C" box="[447,550,225,251]" captionStart="FIGURES 30–32" captionStartId="24.[151,264,1772,1797]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure@24.[151,1436,181,1746]" captionTargetPageId="24" captionText="FIGURES 30–32. Male genitalia of the “aepitus group” species. 30. Artines delfos sp. nov., holotype. 31. Artines maya, holotype. 32. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.215). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241049/files/figure.png" pageId="15" pageNumber="16">Fig. 32A</figureCitation>
). Saccus as long as tegumen+uncus, base shorter than lateral margin (
<figureCitation id="DBC0FBA89D55CA42FAC1FF62FA73FF6C" box="[1322,1424,225,251]" captionStart="FIGURES 30–32" captionStartId="24.[151,264,1772,1797]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure@24.[151,1436,181,1746]" captionTargetPageId="24" captionText="FIGURES 30–32. Male genitalia of the “aepitus group” species. 30. Artines delfos sp. nov., holotype. 31. Artines maya, holotype. 32. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.215). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241049/files/figure.png" pageId="15" pageNumber="16">Fig. 32B</figureCitation>
). Uncus, in dorsal view, rectangular, twice longer than wide, as long as tegumen and without disto-lateral projections, distal margin rounded (
<figureCitation id="DBC0FBA89D55CA42FE4BFEAAFDEAFED4" box="[416,521,297,323]" captionStart="FIGURES 30–32" captionStartId="24.[151,264,1772,1797]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure@24.[151,1436,181,1746]" captionTargetPageId="24" captionText="FIGURES 30–32. Male genitalia of the “aepitus group” species. 30. Artines delfos sp. nov., holotype. 31. Artines maya, holotype. 32. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.215). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241049/files/figure.png" pageId="15" pageNumber="16">Fig. 32C</figureCitation>
). Gnathos as in the other species of the group (
<figureCitation id="DBC0FBA89D55CA42FBC9FEAAFB53FED4" box="[1058,1200,297,323]" captionStart="FIGURES 30–32" captionStartId="24.[151,264,1772,1797]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure@24.[151,1436,181,1746]" captionTargetPageId="24" captionText="FIGURES 30–32. Male genitalia of the “aepitus group” species. 30. Artines delfos sp. nov., holotype. 31. Artines maya, holotype. 32. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.215). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241049/files/figure.png" pageId="15" pageNumber="16">Fig. 32A, D</figureCitation>
). Valva twice longer than wide; sacculus triangular reaching 1/2 width of valva; costa narrow and contiguous with ampulla; ampulla elongated and shorter than costa; ventral process of harpe short and triangular, shorter than tegumen and slightly longer than dorsal process, with apical setae thin and long; dorsal process of harpe on distal 1/2, rounded in dorsal view and wider than ventral process (
<figureCitation id="DBC0FBA89D55CA42FDDCFE3AFD64FE44" box="[567,647,441,467]" captionStart="FIGURES 30–32" captionStartId="24.[151,264,1772,1797]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure@24.[151,1436,181,1746]" captionTargetPageId="24" captionText="FIGURES 30–32. Male genitalia of the “aepitus group” species. 30. Artines delfos sp. nov., holotype. 31. Artines maya, holotype. 32. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.215). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241049/files/figure.png" pageId="15" pageNumber="16">Fig. 32</figureCitation>
E–F). Aedeagus longer than valva, moderately curved dorsally; ventral projection distally wide, in lateral view slightly curved ventrally (
<figureCitation id="DBC0FBA89D55CA42FC84FE5EFC23FE60" box="[879,960,477,503]" captionStart="FIGURES 30–32" captionStartId="24.[151,264,1772,1797]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure@24.[151,1436,181,1746]" captionTargetPageId="24" captionText="FIGURES 30–32. Male genitalia of the “aepitus group” species. 30. Artines delfos sp. nov., holotype. 31. Artines maya, holotype. 32. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.215). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241049/files/figure.png" pageId="15" pageNumber="16">Fig. 32</figureCitation>
G–J); cornutus similar to
<taxonomicName id="84FB9CAE9D55CA42FB31FE5EFADFFE60" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[1242,1340,477,503]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="delfos" status="sp. nov.">
<emphasis id="718F3B3F9D55CA42FB31FE5EFADFFE60" box="[1242,1340,477,503]" italics="true" pageId="15" pageNumber="16">A. delfos</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D55CA42FAA9FE5EFA7FFE60" bold="true" box="[1346,1436,477,503]" pageId="15" pageNumber="16">
<taxonomicNameLabel id="6ABC86449D55CA42FAA9FE5EFA7FFE60" box="[1346,1436,477,503]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
but narrower and less corrugated (
<figureCitation id="DBC0FBA89D55CA42FDFEFD82FD89FD8C" box="[533,618,512,539]" captionStart="FIGURES 42–48" captionStartId="33.[151,264,1892,1917]" captionTargetBox="[151,1431,625,1874]" captionTargetId="figure@33.[151,1436,625,1880]" captionText="FIGURES 42–48. Dorsal view of vesica and cornutus of the &quot;aepitus group&quot; species. 42. Artines aepitus. 43. Artines panama sp. nov.. 44. Artines delfos sp. nov.. 45. Artines litoralis sp. nov.. 46. Artines rica. 47. Artines liege sp. nov. 48. Artines bamba sp. nov. Scale bar = 0,2 mm." httpUri="https://zenodo.org/record/3406274/files/figure.png" pageId="15" pageNumber="16">Fig. 45</figureCitation>
).
</paragraph>
<paragraph id="4344E72D9D55CA42FF2CFDA6FE91FDF4" blockId="15.[151,1437,152,2015]" pageId="15" pageNumber="16">
Female (
<figureCitation id="DBC0FBA89D55CA42FEC2FDA6FE99FDA8" box="[297,378,549,575]" captionStart="FIGURES 9–13" captionStartId="7.[151,264,1180,1205]" captionTargetBox="[152,1436,180,1162]" captionTargetId="figure@7.[151,1436,174,1169]" captionTargetPageId="7" captionText="FIGURES 9–13. Species of the “aepitus group” in dorsal and ventral views. 9. Artines maya, A–B: holotype male. 10. Artines litoralis sp. nov., A–B: holotype male, C–D: allotype female. 11. Artines rica, A–B: male, Turrialba, Cartago, Costa Rica (DZ 41.890), C–D: female from same locality of male (DZ 41.840). 12. Artines liege sp. nov., A–B: holotype male, C–D: allotype female. 13. Artines bamba sp. nov., A–B: holotype male, C–D: tentative female, Moyobamba, San Martin, Peru (OM 79.413). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241043/files/figure.png" pageId="15" pageNumber="16">Fig. 10</figureCitation>
C–D).
<emphasis id="718F3B3F9D55CA42FE2CFDA5FD62FDA8" box="[455,641,549,575]" italics="true" pageId="15" pageNumber="16">Forewing length</emphasis>
:
<quantity id="84034AC89D55CA42FD7AFDA6FCEEFDA8" box="[657,781,549,575]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="15" pageNumber="16" unit="mm" value="13.5" valueMax="14.0" valueMin="13.0">13–14 mm</quantity>
. Similar to male, differing in the more conspicuous marks on dorsal forewing.
</paragraph>
<paragraph id="4344E72D9D55CA42FF2CFDEEFF03FC80" blockId="15.[151,1437,152,2015]" pageId="15" pageNumber="16">
<emphasis id="718F3B3F9D55CA42FF2CFDEEFED2FD10" box="[199,305,621,647]" italics="true" pageId="15" pageNumber="16">Genitalia</emphasis>
(
<figureCitation id="DBC0FBA89D55CA42FEADFDEEFE43FD10" box="[326,416,620,647]" captionStart="FIGURES 54–60" captionStartId="35.[151,264,1807,1832]" captionTargetBox="[156,1436,575,1772]" captionTargetId="figure@35.[151,1436,589,1795]" captionTargetPageId="35" captionText="FIGURES 54–60. Female genitalia of the “aepitus group” species. 54. Artines aepitus, Jusepín, Monagas, Venezuela (OM 30. 125). 55. Artines panama sp. nov., allotype. 56. Artines solange sp. nov. (tentative female), Sumaúma, Maranhão, Brazil (DZ 41.800). 57. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.820). 58. Artines rica, Turrialba, Cartago, Costa Rica (DZ 41.840). 59. Artines liege sp. nov., paratype, Parque do Gama, Brasília, Distrito Federal, Brazil (DZ 41.830). 60. Artines bamba sp. nov. (tentative female), Moyobamba, San Martin, Peru (OM 81.229). A) lateral view of tergum VIII, sterigma and papilla analis; B) ventral view of esterigma. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241059/files/figure.png" pageId="15" pageNumber="16">Figs 57</figureCitation>
A–B, 69): tergum VIII with complete spiracular opening. Sterigma triangular; lamella antevaginalis with proximo-ventral folds elongated; lamella postvaginalis without lateral fold, with distal projections wider than in
<taxonomicName id="84FB9CAE9D55CA42FED1FD36FE4EFD58" baseAuthorityName="Geyer" baseAuthorityYear="1832" box="[314,429,693,719]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D55CA42FED1FD36FE4EFD58" box="[314,429,693,719]" italics="true" pageId="15" pageNumber="16">A. aepitus</emphasis>
</taxonomicName>
and separated medially by narrow and shallow indentation, proximo-lateral projections triangular and not bilobed. Ostium narrow, surrounded by membranous areas; ductus bursae without sclerotized plates.
</paragraph>
<paragraph id="4344E72D9D55CA42FF2CFCA3FEA6FCC8" blockId="15.[151,1437,152,2015]" pageId="15" pageNumber="16">
<emphasis id="718F3B3F9D55CA42FF2CFCA3FED9FCAC" bold="true" box="[199,314,800,827]" pageId="15" pageNumber="16">Variation</emphasis>
. The central black spot on ventral hind wing is longer than wide in specimens from
<collectingRegion id="813F29CF9D55CA42FB0AFCA2FA8EFCAC" box="[1249,1389,801,827]" country="Brazil" name="Pernambuco" pageId="15" pageNumber="16">Pernambuco</collectingRegion>
and
<collectingRegion id="813F29CF9D55CA42FF7CFCC6FF0DFCC8" box="[151,238,837,863]" country="Brazil" name="Paraiba" pageId="15" pageNumber="16">Paraíba</collectingRegion>
,
<collectingCountry id="3BECA7BD9D55CA42FF12FCC6FEA2FCC8" box="[249,321,837,863]" name="Brazil" pageId="15" pageNumber="16">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D55CA42FF2CFCEBFBC6FBCC" pageId="15" pageNumber="16" type="discussion">
<paragraph id="4344E72D9D55CA42FF2CFCEBFB66FC78" blockId="15.[151,1437,152,2015]" pageId="15" pageNumber="16">
<emphasis id="718F3B3F9D55CA42FF2CFCEBFEABFC14" bold="true" box="[199,328,872,899]" pageId="15" pageNumber="16">Comments</emphasis>
.
<taxonomicName id="84FB9CAE9D55CA42FEBEFCEAFDE6FC14" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[341,517,873,899]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="litoralis" status="sp. nov.">
<emphasis id="718F3B3F9D55CA42FEBEFCEAFDE6FC14" box="[341,517,873,899]" italics="true" pageId="15" pageNumber="16">Artines litoralis</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D55CA42FDE0FCEAFD86FC14" bold="true" box="[523,613,873,899]" pageId="15" pageNumber="16">
<taxonomicNameLabel id="6ABC86449D55CA42FDE0FCEAFD86FC14" box="[523,613,873,899]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is the only species of the “
<taxonomicName id="84FB9CAE9D55CA42FC66FCE9FC3DFC14" baseAuthorityName="Geyer" baseAuthorityYear="1832" box="[909,990,874,899]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D55CA42FC66FCE9FC3DFC14" box="[909,990,874,899]" italics="true" pageId="15" pageNumber="16">aepitus</emphasis>
</taxonomicName>
group” which occurs in the low areas of the Brazilian coast, penetrating inland along the Rio Doce Valley, in the states of
<collectingRegion id="813F29CF9D55CA42FBC2FC0EFB2AFC30" box="[1065,1225,909,935]" country="Brazil" name="Espirito Santo" pageId="15" pageNumber="16">Espírito Santo</collectingRegion>
and
<collectingRegion id="813F29CF9D55CA42FB14FC0EFA7AFC30" box="[1279,1433,909,935]" country="Brazil" name="Minas Gerais" pageId="15" pageNumber="16">Minas Gerais</collectingRegion>
. The farthest record from the coast is Paraopeba,
<collectingRegion id="813F29CF9D55CA42FD28FC32FCBDFC5C" box="[707,862,945,971]" country="Brazil" name="Minas Gerais" pageId="15" pageNumber="16">Minas Gerais</collectingRegion>
. There the species is potentially sympatric with
<taxonomicName id="84FB9CAE9D55CA42FA6FFC32FF2EFC78" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="liege" status="sp. nov.">
<emphasis id="718F3B3F9D55CA42FA6FFC32FF2EFC78" italics="true" pageId="15" pageNumber="16">A. liege</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D55CA42FF3EFC56FED3FC78" bold="true" box="[213,304,981,1007]" pageId="15" pageNumber="16">
<taxonomicNameLabel id="6ABC86449D55CA42FF3EFC56FED3FC78" box="[213,304,981,1007]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, from which it can be easily distinguished by the male and female genitalia.
</paragraph>
<paragraph id="4344E72D9D55CA42FF2CFC7BFBC6FBCC" blockId="15.[151,1437,152,2015]" pageId="15" pageNumber="16">
<emphasis id="718F3B3F9D55CA42FF2CFC7BFE0AFB84" bold="true" box="[199,489,1016,1043]" pageId="15" pageNumber="16">Geographic distribution</emphasis>
. (
<figureCitation id="DBC0FBA89D55CA42FE14FC7AFDB6FB84" box="[511,597,1017,1043]" captionStart="FIGURE 79" captionStartId="42.[151,250,1782,1807]" captionTargetBox="[151,1436,1021,1770]" captionTargetId="figure@42.[151,1436,1021,1770]" captionTargetPageId="42" captionText="FIGURE 79. Geographic distribution of the “aepitus group” species. The symbols with white inferior half represent records from literature." httpUri="https://zenodo.org/record/3241069/files/figure.png" pageId="15" pageNumber="16">Fig. 79</figureCitation>
). The species is restricted to the Atlantic Forest. BRAZIL—
<emphasis id="718F3B3F9D55CA42FB15FC7AFAB9FB84" box="[1278,1370,1017,1043]" italics="true" pageId="15" pageNumber="16">
<collectingRegion id="813F29CF9D55CA42FB15FC7AFAB9FB84" box="[1278,1370,1017,1043]" country="Brazil" name="Paraiba" pageId="15" pageNumber="16">Paraíba</collectingRegion>
</emphasis>
: João Pessoa.
<emphasis id="718F3B3F9D55CA42FF1BFB9EFE9CFBA0" box="[240,383,1053,1079]" italics="true" pageId="15" pageNumber="16">
<collectingRegion id="813F29CF9D55CA42FF1BFB9EFE9CFBA0" box="[240,383,1053,1079]" country="Brazil" name="Pernambuco" pageId="15" pageNumber="16">Pernambuco</collectingRegion>
</emphasis>
: Camaragibe, Recife, São Lourenço da Mata.
<emphasis id="718F3B3F9D55CA42FC96FB9EFC3AFBA0" box="[893,985,1053,1079]" italics="true" pageId="15" pageNumber="16">
<collectingRegion id="813F29CF9D55CA42FC96FB9EFC3AFBA0" box="[893,985,1053,1079]" country="Brazil" name="Alagoas" pageId="15" pageNumber="16">Alagoas</collectingRegion>
</emphasis>
: Maceió.
<emphasis id="718F3B3F9D55CA42FBACFB9EFB04FBA0" box="[1095,1255,1053,1079]" italics="true" pageId="15" pageNumber="16">
<collectingRegion id="813F29CF9D55CA42FBACFB9EFB04FBA0" box="[1095,1255,1053,1079]" country="Brazil" name="Espirito Santo" pageId="15" pageNumber="16">Espírito Santo</collectingRegion>
</emphasis>
: Baixo Guandu, Conceição da Barra, Itaguaçu, Linhares, Santa Teresa (
<bibRefCitation id="276A9ADC9D55CA42FD17FBC2FBFAFBCC" author="Brown Jr., K. S. &amp; Freitas, A. V. L." box="[764,1049,1089,1115]" pageId="15" pageNumber="16" pagination="71 - 116" refId="ref39087" refString="Brown Jr., K. S. &amp; Freitas, A. V. L. (2000) Diversidade de Lepidoptera em Santa Teresa, Espirito Santo. Boletim do Museu de Biologia Mello Leitao, New Series, 11 / 12, 71 - 116" type="journal article" year="2000">Brown Jr &amp; Freitas 2000</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D55CA42FF2CFBE6FB76FBE8" box="[199,1173,1124,1151]" pageId="15" pageNumber="16" type="description">
<paragraph id="4344E72D9D55CA42FF2CFBE6FB76FBE8" blockId="15.[151,1437,152,2015]" box="[199,1173,1124,1151]" pageId="15" pageNumber="16">
<emphasis id="718F3B3F9D55CA42FF2CFBE6FE31FBE8" bold="true" box="[199,466,1124,1151]" pageId="15" pageNumber="16">Temporal distribution</emphasis>
. This species occurs throughout the year, except in November.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D55CA42FF2CFB0AFE2AFB34" box="[199,457,1161,1187]" pageId="15" pageNumber="16" type="biology_ecology">
<paragraph id="4344E72D9D55CA42FF2CFB0AFE2AFB34" blockId="15.[151,1437,152,2015]" box="[199,457,1161,1187]" pageId="15" pageNumber="16">
<emphasis id="718F3B3F9D55CA42FF2CFB0AFEA7FB34" bold="true" box="[199,324,1161,1187]" pageId="15" pageNumber="16">Host plant</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D55CA42FF2CFB2EFB20FB50" box="[199,1219,1197,1223]" pageId="15" pageNumber="16" type="etymology">
<paragraph id="4344E72D9D55CA42FF2CFB2EFB20FB50" blockId="15.[151,1437,152,2015]" box="[199,1219,1197,1223]" pageId="15" pageNumber="16">
<emphasis id="718F3B3F9D55CA42FF2CFB2EFEABFB50" bold="true" box="[199,328,1197,1223]" pageId="15" pageNumber="16">Etymology</emphasis>
. This species is named in reference of its distribution along the Brazilian coast.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D55CA5DFF2CFB52FB99FF6C" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="4344E72D9D55CA42FF2CFB52FEF6FAA4" blockId="15.[151,1437,152,2015]" pageId="15" pageNumber="16">
<emphasis id="718F3B3F9D55CA42FF2CFB52FE92FB7C" bold="true" box="[199,369,1232,1259]" pageId="15" pageNumber="16">Type material</emphasis>
.
<materialsCitation id="F393ED709D55CA42FE6BFB52FBB9FB7C" ID-GBIF-Occurrence="2248478848" box="[384,1114,1233,1259]" collectionCode="DZUP" pageId="15" pageNumber="16" specimenCount="1" typeStatus="holotype">
<typeStatus id="9C40598F9D55CA42FE6BFB52FE0BFB7C" box="[384,488,1233,1259]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
male deposited at the
<collectionCode id="25EA7FE89D55CA42FD02FB52FCD1FB7C" box="[745,818,1233,1259]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="15" pageNumber="16" type="University or college">DZUP</collectionCode>
with the following labels
</materialsCitation>
: /
<materialsCitation id="F393ED709D55CA42FB9EFB52FCB3FA98" ID-GBIF-Occurrence="2248478838" collectingDate="1972-05-17" collectorName="C. Elias" country="Brazil" location="Linhares" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Espirito Santo" typeStatus="holotype">
<typeStatus id="9C40598F9D55CA42FB9EFB52FAF8FB7C" box="[1141,1307,1233,1259]" pageId="15" pageNumber="16" type="holotype">HOLOTYPUS</typeStatus>
/
<location id="4624B1F69D55CA42FAD8FB52FA7BFB7C" LSID="urn:lsid:plazi:treatment:CB52563B9D54CA5DFF7CFB0AFB99FF6C:4624B1F69D55CA42FAD8FB52FA7BFB7C" box="[1331,1432,1233,1259]" country="Brazil" name="Linhares" pageId="15" pageNumber="16" stateProvince="Espirito Santo">Linhares</location>
,
<collectingRegion id="813F29CF9D55CA42FF7CFB76FED8FA98" box="[151,315,1269,1295]" country="Brazil" name="Espirito Santo" pageId="15" pageNumber="16">Espírito Santo</collectingRegion>
,
<collectingCountry id="3BECA7BD9D55CA42FEACFB76FE6DFA98" box="[327,398,1269,1295]" name="Brazil" pageId="15" pageNumber="16">Brazil</collectingCountry>
,
<date id="3745C1ED9D55CA42FE71FB76FDF0FA98" box="[410,531,1269,1295]" pageId="15" pageNumber="16" value="1972-05-17">
<collectingDate id="270138059D55CA42FE71FB76FDF0FA98" box="[410,531,1269,1295]" pageId="15" pageNumber="16" value="1972-05-17">17.V.1972</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D55CA42FDCBFB76FD9AFA98" box="[544,633,1269,1295]" pageId="15" pageNumber="16">C. Elias</collectorName>
[
<emphasis id="718F3B3F9D55CA42FD60FB76FD57FA98" box="[651,692,1269,1295]" italics="true" pageId="15" pageNumber="16">leg.</emphasis>
] / DZ 41.870
</materialsCitation>
/
<typeStatus id="9C40598F9D55CA42FC83FB76FC3EFA98" box="[872,989,1269,1295]" pageId="15" pageNumber="16" type="holotype">Holotypus</typeStatus>
<taxonomicName id="84FB9CAE9D55CA42FC0FFB76FA7FFA98" authority="Medeiros &amp; Dolibaina" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[996,1436,1269,1295]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="litoralis">
<emphasis id="718F3B3F9D55CA42FC0FFB76FB75FA98" box="[996,1174,1269,1295]" italics="true" pageId="15" pageNumber="16">Artines litoralis</emphasis>
Medeiros &amp; Dolibaina
</taxonomicName>
det. 2018 /.
</paragraph>
<paragraph id="4344E72D9D55CA42FF2CFABEFE4AFA08" blockId="15.[151,1437,152,2015]" pageId="15" pageNumber="16">
<materialsCitation id="F393ED709D55CA42FF2CFABEFC56FAC0" ID-GBIF-Occurrence="2248478846" box="[199,949,1341,1367]" collectionCode="DZUP" pageId="15" pageNumber="16" specimenCount="1" typeStatus="allotype">
<typeStatus id="9C40598F9D55CA42FF2CFABEFECAFAC0" box="[199,297,1341,1367]" pageId="15" pageNumber="16" type="allotype">Allotype</typeStatus>
female deposited at the
<collectionCode id="25EA7FE89D55CA42FDA9FABEFD68FAC0" box="[578,651,1341,1367]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="15" pageNumber="16" type="University or college">DZUP</collectionCode>
with the following labels
</materialsCitation>
: /
<materialsCitation id="F393ED709D55CA42FC3AFABEFB85FAEC" ID-GBIF-Occurrence="2248478835" collectingDate="1994-03-08" collectingDateMax="1994-03-09" collectingDateMin="1994-03-08" collectorName="Marlieria, M &amp; Mielke" country="Brazil" elevation="350" location="Est" municipality="Rio Doce" pageId="15" pageNumber="16" specimenCount="1" typeStatus="allotype">
<typeStatus id="9C40598F9D55CA42FC3AFABEFB97FAC0" box="[977,1140,1341,1367]" pageId="15" pageNumber="16" type="allotype">ALLOTYPUS</typeStatus>
/
<date id="3745C1ED9D55CA42FB65FABEFAFFFAC0" box="[1166,1308,1341,1367]" pageId="15" pageNumber="16" value="1994-03-08" valueMax="1994-03-09" valueMin="1994-03-08">
<collectingDate id="270138059D55CA42FB65FABEFAFFFAC0" box="[1166,1308,1341,1367]" pageId="15" pageNumber="16" value="1994-03-08" valueMax="1994-03-09" valueMin="1994-03-08">8–9.III.1994</collectingDate>
</date>
, P[ar]q[ue]
<location id="4624B1F69D55CA42FF7CFAE3FF5EFAEC" LSID="urn:lsid:plazi:treatment:CB52563B9D54CA5DFF7CFB0AFB99FF6C:4624B1F69D55CA42FF7CFAE3FF5EFAEC" box="[151,189,1376,1403]" country="Brazil" municipality="Rio Doce" name="Est" pageId="15" pageNumber="16">Est</location>
[adual]
<collectingMunicipality id="A3207D579D55CA42FEFBFAE2FE9FFAEC" box="[272,380,1377,1403]" pageId="15" pageNumber="16">Rio Doce</collectingMunicipality>
,
<collectorName id="EE0E82FB9D55CA42FE6EFAE2FDF6FAEC" box="[389,533,1376,1403]" pageId="15" pageNumber="16">Marliéria, M</collectorName>
[inas] G[erais,
<collectingCountry id="3BECA7BD9D55CA42FD5CFAE3FCE2FAEC" box="[695,769,1376,1403]" name="Brazil" pageId="15" pageNumber="16">Brazil</collectingCountry>
],
<quantity id="84034AC89D55CA42FCFAFAE3FCB7FAEC" box="[785,852,1376,1403]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="15" pageNumber="16" unit="m" value="350.0">
<elevation id="C8D6001E9D55CA42FCFAFAE3FCB7FAEC" box="[785,852,1376,1403]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="15" pageNumber="16" unit="m" value="350.0">350m</elevation>
</quantity>
,
<collectorName id="EE0E82FB9D55CA42FCB5FAE2FC4EFAEC" box="[862,941,1377,1403]" pageId="15" pageNumber="16">Mielke</collectorName>
<emphasis id="718F3B3F9D55CA42FC59FAE2FC37FAEC" box="[946,980,1377,1403]" italics="true" pageId="15" pageNumber="16">leg</emphasis>
. / DZ 41.880
</materialsCitation>
/
<typeStatus id="9C40598F9D55CA42FB92FAE2FB04FAEC" box="[1145,1255,1377,1403]" pageId="15" pageNumber="16" type="allotype">Allotypus</typeStatus>
<taxonomicName id="84FB9CAE9D55CA42FB06FAE2FE70FA08" authority="Medeiros &amp; Dolibaina" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="litoralis">
<emphasis id="718F3B3F9D55CA42FB06FAE2FA7EFAEC" box="[1261,1437,1377,1403]" italics="true" pageId="15" pageNumber="16">Artines litoralis</emphasis>
Medeiros &amp; Dolibaina
</taxonomicName>
/.
</paragraph>
<paragraph id="4344E72D9D55CA5DFF2CFA2AFB99FF6C" blockId="15.[151,1437,152,2015]" lastBlockId="16.[151,1436,153,251]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<materialsCitation id="F393ED709D55CA42FF2CFA2AFCDCFA54" ID-GBIF-Occurrence="2248478893" box="[199,831,1449,1475]" location="Joao Pessoa" pageId="15" pageNumber="16" specimenCount="1" specimenCount-female="1" stateProvince="Paraiba" typeStatus="paratype">
<typeStatus id="9C40598F9D55CA42FF2CFA2AFEDBFA54" box="[199,312,1449,1475]" pageId="15" pageNumber="16" type="paratype">Paratypes</typeStatus>
: BRAZIL—
<emphasis id="718F3B3F9D55CA42FE2CFA2AFDC0FA54" box="[455,547,1449,1475]" italics="true" pageId="15" pageNumber="16">
<collectingRegion id="813F29CF9D55CA42FE2CFA2AFDC0FA54" box="[455,547,1449,1475]" country="Brazil" name="Paraiba" pageId="15" pageNumber="16">Paraíba</collectingRegion>
</emphasis>
:
<emphasis id="718F3B3F9D55CA42FDDEFA2AFD25FA54" bold="true" box="[565,710,1449,1475]" pageId="15" pageNumber="16">
<location id="4624B1F69D55CA42FDDEFA2AFD25FA54" LSID="urn:lsid:plazi:treatment:CB52563B9D54CA5DFF7CFB0AFB99FF6C:4624B1F69D55CA42FDDEFA2AFD25FA54" box="[565,710,1449,1475]" name="Joao Pessoa" pageId="15" pageNumber="16" stateProvince="Paraiba">João Pessoa</location>
</emphasis>
,
<specimenCount id="55FD2CA49D55CA42FD3CFA2AFCDCFA54" box="[727,831,1449,1475]" pageId="15" pageNumber="16" type="female">1 female</specimenCount>
</materialsCitation>
,
<date id="3745C1ED9D55CA42FCA6FA2AFC3AFA54" box="[845,985,1448,1475]" pageId="15" pageNumber="16" value="1954-07-24">24.VII.1954</date>
, Kesselring
<emphasis id="718F3B3F9D55CA42FB82FA2AFB68FA54" box="[1129,1163,1449,1475]" italics="true" pageId="15" pageNumber="16">leg</emphasis>
., DZ 41.750,
<specimenCount id="55FD2CA49D55CA42FADAFA2AFA7BFA54" box="[1329,1432,1449,1475]" pageId="15" pageNumber="16" type="female">1 female</specimenCount>
,
<date id="3745C1ED9D55CA42FF7CFA4EFEEFFA70" box="[151,268,1485,1511]" pageId="15" pageNumber="16" value="1992-06-04">
4.
<collectionCode id="25EA7FE89D55CA42FF47FA4EFF2FFA70" box="[172,204,1485,1511]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="15" pageNumber="16">VI</collectionCode>
<materialsCitation id="F393ED709D55CA42FF27FA4EFEEFFA70" ID-GBIF-Occurrence="2248478897" box="[204,268,1485,1511]" collectionCode="DZUP" collectorName="Tangerini" location="Camaragibe" pageId="15" pageNumber="16" specimenCount="1" specimenCount-female="1" stateProvince="Pernambuco" typeStatus="paratype">.1992</materialsCitation>
</date>
,
<collectorName id="EE0E82FB9D55CA42FEF0FA4EFE6BFA70" box="[283,392,1485,1511]" pageId="15" pageNumber="16">Tangerini</collectorName>
<emphasis id="718F3B3F9D55CA42FE7FFA4EFE55FA70" box="[404,438,1485,1511]" italics="true" pageId="15" pageNumber="16">leg</emphasis>
., DZ 41.809 (
<collectionCode id="25EA7FE89D55CA42FD89FA4EFD51FA70" box="[610,690,1485,1511]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="15" pageNumber="16" type="University or college">DZUP</collectionCode>
).
<emphasis id="718F3B3F9D55CA42FD23FA4EFCB4FA70" box="[712,855,1485,1511]" italics="true" pageId="15" pageNumber="16">
<collectingRegion id="813F29CF9D55CA42FD23FA4EFCB4FA70" box="[712,855,1485,1511]" country="Brazil" name="Pernambuco" pageId="15" pageNumber="16">Pernambuco</collectingRegion>
</emphasis>
:
<emphasis id="718F3B3F9D55CA42FC80FA4FFC1CFA70" bold="true" box="[875,1023,1484,1511]" pageId="15" pageNumber="16">
<location id="4624B1F69D55CA42FC80FA4FFC1CFA70" LSID="urn:lsid:plazi:treatment:CB52563B9D54CA5DFF7CFB0AFB99FF6C:4624B1F69D55CA42FC80FA4FFC1CFA70" box="[875,1023,1484,1511]" name="Camaragibe" pageId="15" pageNumber="16" stateProvince="Pernambuco">Camaragibe</location>
</emphasis>
,
<specimenCount id="55FD2CA49D55CA42FBF9FA4EFB99FA70" box="[1042,1146,1485,1511]" pageId="15" pageNumber="16" type="female">1 female</specimenCount>
,
<date id="3745C1ED9D55CA42FB61FA4EFAFCFA70" box="[1162,1311,1484,1511]" pageId="15" pageNumber="16" value="1957-08-18">18.VIII.1957</date>
,
<specimenCount id="55FD2CA49D55CA42FAC4FA4EFA7BFA70" box="[1327,1432,1485,1511]" pageId="15" pageNumber="16" type="female">1 female</specimenCount>
,
<date id="3745C1ED9D55CA42FF7CFA72FEFAF99C" box="[151,281,1520,1547]" pageId="15" pageNumber="16" value="1958-04-27">27.IV.1958</date>
,
<specimenCount id="55FD2CA49D55CA42FEC2FA72FE71F99C" box="[297,402,1521,1547]" pageId="15" pageNumber="16" type="female">1 female</specimenCount>
,
<date id="3745C1ED9D55CA42FE4AFA72FDFDF99C" box="[417,542,1520,1547]" pageId="15" pageNumber="16" value="1958-07-07">7.VII.1958</date>
,
<specimenCount id="55FD2CA49D55CA42FDC4FA72FD74F99C" box="[559,663,1521,1547]" pageId="15" pageNumber="16" type="female">1 female</specimenCount>
, 18.XIII.1958,
<specimenCount id="55FD2CA49D55CA42FCA0FA72FC5CF99C" box="[843,959,1521,1547]" pageId="15" pageNumber="16" type="female">2 females</specimenCount>
,
<date id="3745C1ED9D55CA42FC25FA72FBA0F99C" box="[974,1091,1520,1547]" pageId="15" pageNumber="16" value="1959-09-07">7.IX.1959</date>
,
<specimenCount id="55FD2CA49D55CA42FBB9FA72FB53F99C" box="[1106,1200,1521,1547]" pageId="15" pageNumber="16" type="male">2 males</specimenCount>
,
<date id="3745C1ED9D55CA42FB2BFA72FAFCF99C" box="[1216,1311,1521,1547]" pageId="15" pageNumber="16" value="1960-01-02">2.I.1960</date>
,
<specimenCount id="55FD2CA49D55CA42FADBFA72FA7BF99C" box="[1328,1432,1521,1547]" pageId="15" pageNumber="16" type="female">1 female</specimenCount>
,
<materialsCitation id="F393ED709D55CA42FF7CF996FD64F9C4" ID-GBIF-Occurrence="2248478843" collectingDate="1960-10-29" collectionCode="DZUP" collectorName="H. Ebert" location="Recife" pageId="15" pageNumber="16" specimenCount="1" specimenCount-male="1" stateProvince="Pernambuco" typeStatus="paratype">
<date id="3745C1ED9D55CA42FF7CF996FEF3F9B8" box="[151,272,1557,1583]" pageId="15" pageNumber="16" value="1960-10-29">
<collectingDate id="270138059D55CA42FF7CF996FEF3F9B8" box="[151,272,1557,1583]" pageId="15" pageNumber="16" value="1960-10-29">29.X.1960</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D55CA42FEF7F996FE9FF9B8" box="[284,380,1557,1583]" pageId="15" pageNumber="16">H. Ebert</collectorName>
<emphasis id="718F3B3F9D55CA42FE68F996FE46F9B8" box="[387,421,1557,1583]" italics="true" pageId="15" pageNumber="16">leg</emphasis>
., DZ 39.886, DZ 39.903, DZ 39.892, DZ 39.887, DZ 39.890, DZ 39.899, DZ 41.780, DZ 39.884, DZ 39.902 (
<collectionCode id="25EA7FE89D55CA42FE97F9BAFE28F9C4" box="[380,459,1593,1619]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="15" pageNumber="16" type="University or college">DZUP</collectionCode>
);
<emphasis id="718F3B3F9D55CA42FE34F9BBFDC9F9C4" bold="true" box="[479,554,1592,1619]" pageId="15" pageNumber="16">
<location id="4624B1F69D55CA42FE34F9BBFDC9F9C4" LSID="urn:lsid:plazi:treatment:CB52563B9D54CA5DFF7CFB0AFB99FF6C:4624B1F69D55CA42FE34F9BBFDC9F9C4" box="[479,554,1592,1619]" name="Recife" pageId="15" pageNumber="16" stateProvince="Pernambuco">Recife</location>
</emphasis>
,
<specimenCount id="55FD2CA49D55CA42FDD2F9BAFD64F9C4" box="[569,647,1593,1619]" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
</materialsCitation>
,
<date id="3745C1ED9D55CA42FD78F9BAFD1DF9C4" box="[659,766,1592,1619]" pageId="15" pageNumber="16" value="1959-05-03">3.V.1959</date>
,
<specimenCount id="55FD2CA49D55CA42FCE1F9BAFCBBF9C4" box="[778,856,1593,1619]" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
,
<materialsCitation id="F393ED709D55CA42FC8FF9BAFD5EF9E0" ID-GBIF-Occurrence="2248478840" collectingDate="1961-04-01" collectionCode="DZUP, OM" collectorName="H. Ebert" location="Sao Lourenco da Mata" pageId="15" pageNumber="16" specimenCode="OM 10.342" specimenCount="1" specimenCount-female="1" stateProvince="Pernambuco" typeStatus="paratype">
<date id="3745C1ED9D55CA42FC8FF9BAFC3BF9C4" box="[868,984,1593,1619]" pageId="15" pageNumber="16" value="1961-04-01">
<collectingDate id="270138059D55CA42FC8FF9BAFC3BF9C4" box="[868,984,1593,1619]" pageId="15" pageNumber="16" value="1961-04-01">1.IV.1961</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D55CA42FC0FF9BAFBA7F9C4" box="[996,1092,1593,1619]" pageId="15" pageNumber="16">H. Ebert</collectorName>
<emphasis id="718F3B3F9D55CA42FBA7F9BAFB96F9C4" box="[1100,1141,1593,1619]" italics="true" pageId="15" pageNumber="16">leg.</emphasis>
, DZ 39.893 (
<collectionCode id="25EA7FE89D55CA42FAE6F9BAFABFF9C4" box="[1293,1372,1593,1619]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="15" pageNumber="16" type="University or college">DZUP</collectionCode>
),
<specimenCode id="135D4F569D55CA42FA84F9BAFF07F9E0" collectionCode="OM" country="New Zealand" name="Otago Museum" pageId="15" pageNumber="16">OM 10.342</specimenCode>
(
<collectionCode id="25EA7FE89D55CA42FF1FF9DEFEC5F9E0" box="[244,294,1629,1655]" country="New Zealand" name="Otago Museum" pageId="15" pageNumber="16">OM</collectionCode>
);
<emphasis id="718F3B3F9D55CA42FED1F9DFFDA8F9E0" bold="true" box="[314,587,1628,1655]" pageId="15" pageNumber="16">
<location id="4624B1F69D55CA42FED1F9DFFDA8F9E0" LSID="urn:lsid:plazi:treatment:CB52563B9D54CA5DFF7CFB0AFB99FF6C:4624B1F69D55CA42FED1F9DFFDA8F9E0" box="[314,587,1628,1655]" name="Sao Lourenco da Mata" pageId="15" pageNumber="16" stateProvince="Pernambuco">São Lourenço da Mata</location>
</emphasis>
,
<specimenCount id="55FD2CA49D55CA42FDB2F9DEFD5EF9E0" box="[601,701,1629,1655]" pageId="15" pageNumber="16" type="female">1 female</specimenCount>
</materialsCitation>
,
<date id="3745C1ED9D55CA42FD22F9DEFCB7F9E0" box="[713,852,1629,1655]" pageId="15" pageNumber="16" value="1972-07-26">26.VII.1972</date>
, Kesselring
<emphasis id="718F3B3F9D55CA42FC35F9DEFBE3F9E0" box="[990,1024,1629,1655]" italics="true" pageId="15" pageNumber="16">leg</emphasis>
., DZ 39.896,
<specimenCount id="55FD2CA49D55CA42FB76F9DEFAE2F9E0" box="[1181,1281,1629,1655]" pageId="15" pageNumber="16" type="female">1 female</specimenCount>
,
<materialsCitation id="F393ED709D55CA42FAE7F9DEFC86F90C" ID-GBIF-Occurrence="2248478856" collectingDate="1961-12-27" collectionCode="DZUP" collectorName="H. Ebert" location="Maceio" pageId="15" pageNumber="16" specimenCount="1" specimenCount-female="1" stateProvince="Alagoas" typeStatus="paratype">
<date id="3745C1ED9D55CA42FAE7F9DEFA74F9E0" box="[1292,1431,1629,1655]" pageId="15" pageNumber="16" value="1961-12-27">
<collectingDate id="270138059D55CA42FAE7F9DEFA74F9E0" box="[1292,1431,1629,1655]" pageId="15" pageNumber="16" value="1961-12-27">27.XII.1961</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D55CA42FF7CF902FF1AF90C" box="[151,249,1665,1691]" pageId="15" pageNumber="16">H. Ebert</collectorName>
<emphasis id="718F3B3F9D55CA42FEEAF902FEC8F90C" box="[257,299,1665,1691]" italics="true" pageId="15" pageNumber="16">leg.</emphasis>
, DZ 39.900 (
<collectionCode id="25EA7FE89D55CA42FE2CF902FDF4F90C" box="[455,535,1665,1691]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="15" pageNumber="16" type="University or college">DZUP</collectionCode>
).
<emphasis id="718F3B3F9D55CA42FDC0F902FD64F90C" box="[555,647,1665,1691]" italics="true" pageId="15" pageNumber="16">
<collectingRegion id="813F29CF9D55CA42FDC0F902FD64F90C" box="[555,647,1665,1691]" country="Brazil" name="Alagoas" pageId="15" pageNumber="16">Alagoas</collectingRegion>
</emphasis>
:
<emphasis id="718F3B3F9D55CA42FD73F903FD0CF90C" bold="true" box="[664,751,1664,1691]" pageId="15" pageNumber="16">
<location id="4624B1F69D55CA42FD73F903FD0CF90C" LSID="urn:lsid:plazi:treatment:CB52563B9D54CA5DFF7CFB0AFB99FF6C:4624B1F69D55CA42FD73F903FD0CF90C" box="[664,751,1664,1691]" name="Maceio" pageId="15" pageNumber="16" stateProvince="Alagoas">Maceió</location>
</emphasis>
,
<specimenCount id="55FD2CA49D55CA42FD14F902FC86F90C" box="[767,869,1665,1691]" pageId="15" pageNumber="16" type="female">1 female</specimenCount>
</materialsCitation>
,
<materialsCitation id="F393ED709D55CA42FC98F902FDA8F928" ID-GBIF-Occurrence="2248478847" collectingDate="1973-08-11" collectionCode="DZUP" collectorName="Cardoso" location="Baixo Guandu" pageId="15" pageNumber="16" specimenCount="1" specimenCount-male="1" stateProvince="Espirito Santo" typeStatus="paratype">
<date id="3745C1ED9D55CA42FC98F902FBEBF90C" box="[883,1032,1665,1691]" pageId="15" pageNumber="16" value="1973-08-11">
<collectingDate id="270138059D55CA42FC98F902FBEBF90C" box="[883,1032,1665,1691]" pageId="15" pageNumber="16" value="1973-08-11">11.VIII.1973</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D55CA42FBFEF902FB91F90C" box="[1045,1138,1665,1691]" pageId="15" pageNumber="16">Cardoso</collectorName>
<emphasis id="718F3B3F9D55CA42FB90F902FB7EF90C" box="[1147,1181,1665,1691]" italics="true" pageId="15" pageNumber="16">leg</emphasis>
., DZ 41.770 (
<collectionCode id="25EA7FE89D55CA42FAAAF902FA72F90C" box="[1345,1425,1665,1691]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="15" pageNumber="16" type="University or college">DZUP</collectionCode>
).
<emphasis id="718F3B3F9D55CA42FF7CF925FED4F928" box="[151,311,1701,1728]" italics="true" pageId="15" pageNumber="16">
<collectingRegion id="813F29CF9D55CA42FF7CF925FED4F928" box="[151,311,1701,1728]" country="Brazil" name="Espirito Santo" pageId="15" pageNumber="16">Espírito Santo</collectingRegion>
</emphasis>
:
<emphasis id="718F3B3F9D55CA42FEAEF927FE12F928" bold="true" box="[325,497,1700,1727]" pageId="15" pageNumber="16">
<location id="4624B1F69D55CA42FEAEF927FE12F928" LSID="urn:lsid:plazi:treatment:CB52563B9D54CA5DFF7CFB0AFB99FF6C:4624B1F69D55CA42FEAEF927FE12F928" box="[325,497,1700,1727]" name="Baixo Guandu" pageId="15" pageNumber="16" stateProvince="Espirito Santo">Baixo Guandu</location>
</emphasis>
,
<specimenCount id="55FD2CA49D55CA42FE15F926FDA8F928" box="[510,587,1701,1727]" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
</materialsCitation>
,
<date id="3745C1ED9D55CA42FDBEF927FD34F928" box="[597,727,1700,1727]" pageId="15" pageNumber="16" value="1970-04-15">15.IV.1970</date>
,
<specimenCount id="55FD2CA49D55CA42FD0AF926FCA7F928" box="[737,836,1701,1727]" pageId="15" pageNumber="16" type="female">1 female</specimenCount>
,
<date id="3745C1ED9D55CA42FCA5F927FC33F928" box="[846,976,1700,1727]" pageId="15" pageNumber="16" value="1970-04-25">25.IV.1970</date>
,
<specimenCount id="55FD2CA49D55CA42FC31F926FBC4F928" box="[986,1063,1701,1727]" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
,
<specimenCount id="55FD2CA49D55CA42FBDAF926FB77F928" box="[1073,1172,1701,1727]" pageId="15" pageNumber="16" type="female">1 female</specimenCount>
,
<date id="3745C1ED9D55CA42FB75F926FAEBF928" box="[1182,1288,1701,1727]" pageId="15" pageNumber="16" value="1971-10-04">4.X.1971</date>
,
<specimenCount id="55FD2CA49D55CA42FAF8F926FA95F928" box="[1299,1398,1701,1727]" pageId="15" pageNumber="16" type="female">1 female</specimenCount>
,
<materialsCitation id="F393ED709D55CA42FA6BF926FE05F890" ID-GBIF-Occurrence="2248478887" collectingDate="1971-04-01" collectingDateMax="1971-04-10" collectingDateMin="1971-04-01" collectionCode="DZUP" collectorName="C. &amp; C. T. Elias" location="Conceicao da Barra" pageId="15" pageNumber="16" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Espirito Santo" typeStatus="paratype">
<date id="3745C1ED9D55CA42FA6BF926FEFAF974" pageId="15" pageNumber="16" value="1971-04-01" valueMax="1971-04-10" valueMin="1971-04-01">
<collectingDate id="270138059D55CA42FA6BF926FEFAF974" pageId="15" pageNumber="16" value="1971-04-01" valueMax="1971-04-10" valueMin="1971-04-01">1– 10.IV.1971</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D55CA42FECEF94AFE39F974" box="[293,474,1737,1763]" pageId="15" pageNumber="16">C. &amp; C. T. Elias</collectorName>
<emphasis id="718F3B3F9D55CA42FE0AF94AFDE8F974" box="[481,523,1737,1763]" italics="true" pageId="15" pageNumber="16">leg.</emphasis>
, DZ 41.891, DZ 39.885, DZ 41.779, DZ 41.889, DZ 41.839 (
<collectionCode id="25EA7FE89D55CA42FB54F94AFAEDF974" box="[1215,1294,1737,1763]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="15" pageNumber="16" type="University or college">DZUP</collectionCode>
);
<emphasis id="718F3B3F9D55CA42FACAF94BFEEBF890" bold="true" pageId="15" pageNumber="16">
<location id="4624B1F69D55CA42FACAF94BFEEBF890" LSID="urn:lsid:plazi:treatment:CB52563B9D54CA5DFF7CFB0AFB99FF6C:4624B1F69D55CA42FACAF94BFEEBF890" name="Conceicao da Barra" pageId="15" pageNumber="16" stateProvince="Espirito Santo">Conceição da Barra</location>
</emphasis>
,
<specimenCount id="55FD2CA49D55CA42FEF1F96EFE8EF890" box="[282,365,1773,1799]" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
,
<specimenCount id="55FD2CA49D55CA42FE96F96EFE05F890" box="[381,486,1773,1799]" pageId="15" pageNumber="16" type="female">1 female</specimenCount>
</materialsCitation>
,
<date id="3745C1ED9D55CA42FE1EF96EFD9BF890" box="[501,632,1773,1799]" pageId="15" pageNumber="16" value="1968-04-10">10.IV.1968</date>
,
<specimenCount id="55FD2CA49D55CA42FD63F96EFD39F890" box="[648,730,1773,1799]" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
,
<date id="3745C1ED9D55CA42FD01F96EFC8FF890" box="[746,876,1773,1799]" pageId="15" pageNumber="16" value="1969-04-12">12.IV.1969</date>
,
<specimenCount id="55FD2CA49D55CA42FC97F96EFC2CF890" box="[892,975,1773,1799]" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
,
<date id="3745C1ED9D55CA42FC34F96FFBA9F890" box="[991,1098,1772,1799]" pageId="15" pageNumber="16" value="1969-05-05">5.V.1969</date>
,
<specimenCount id="55FD2CA49D55CA42FBB1F96EFB4FF890" box="[1114,1196,1773,1799]" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
,
<date id="3745C1ED9D55CA42FB57F96EFAD6F890" box="[1212,1333,1773,1799]" pageId="15" pageNumber="16" value="1969-05-14">14.V.1969</date>
,
<specimenCount id="55FD2CA49D55CA42FAAEF96EFA7BF890" box="[1349,1432,1773,1799]" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
,
<date id="3745C1ED9D55CA42FF7CF893FEE8F8BC" box="[151,267,1808,1835]" pageId="15" pageNumber="16" value="1969-06-05">
5.
<collectionCode id="25EA7FE89D55CA42FF47F892FF2FF8BC" box="[172,204,1809,1835]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="15" pageNumber="16">VI</collectionCode>
.1969
</date>
,
<specimenCount id="55FD2CA49D55CA42FEF2F892FE8AF8BC" box="[281,361,1809,1835]" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
,
<date id="3745C1ED9D55CA42FE9DF892FE1BF8BC" box="[374,504,1809,1835]" pageId="15" pageNumber="16" value="1969-06-18">
18.
<collectionCode id="25EA7FE89D55CA42FE72F892FE5AF8BC" box="[409,441,1809,1835]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="15" pageNumber="16">VI</collectionCode>
.1969
</date>
,
<specimenCount id="55FD2CA49D55CA42FDEDF892FD88F8BC" box="[518,619,1809,1835]" pageId="15" pageNumber="16" type="female">1 female</specimenCount>
,
<date id="3745C1ED9D55CA42FD92F892FD1CF8BC" box="[633,767,1809,1835]" pageId="15" pageNumber="16" value="1969-08-01">1.VIII.1969</date>
,
<specimenCount id="55FD2CA49D55CA42FCE6F892FC8BF8BC" box="[781,872,1809,1835]" pageId="15" pageNumber="16" type="male">3 males</specimenCount>
,
<specimenCount id="55FD2CA49D55CA42FC9EF892FC38F8BC" box="[885,987,1809,1835]" pageId="15" pageNumber="16" type="female">1 female</specimenCount>
,
<materialsCitation id="F393ED709D55CA42FC03F892FD32F8E4" ID-GBIF-Occurrence="2248478844" collectingDate="1969-09-10" collectionCode="DZUP" collectorName="C. &amp; C. T. Elias" location="Itaguacu" pageId="15" pageNumber="16" specimenCount="1" specimenCount-male="1" stateProvince="Espirito Santo" typeStatus="paratype">
<date id="3745C1ED9D55CA42FC03F892FB89F8BC" box="[1000,1130,1809,1835]" pageId="15" pageNumber="16" value="1969-09-10">
<collectingDate id="270138059D55CA42FC03F892FB89F8BC" box="[1000,1130,1809,1835]" pageId="15" pageNumber="16" value="1969-09-10">10.IX.1969</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D55CA42FB93F892FAD6F8BC" box="[1144,1333,1809,1835]" pageId="15" pageNumber="16">C. &amp; C. T. Elias</collectorName>
<emphasis id="718F3B3F9D55CA42FAD5F892FA83F8BC" box="[1342,1376,1809,1835]" italics="true" pageId="15" pageNumber="16">leg</emphasis>
., DZ 41.902, DZ 41.829, DZ 4.062, DZ 41.801, DZ 41.751, DZ 41.912, DZ 41.761, DZ 39.888, DZ 41.771, DZ 41.781, DZ 41.791, DZ 39.891 (
<collectionCode id="25EA7FE89D55CA42FE43F8DAFE14F8E4" box="[424,503,1881,1907]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="15" pageNumber="16" type="University or college">DZUP</collectionCode>
);
<emphasis id="718F3B3F9D55CA42FDE0F8DAFD96F8E4" bold="true" box="[523,629,1881,1907]" pageId="15" pageNumber="16">
<location id="4624B1F69D55CA42FDE0F8DAFD96F8E4" LSID="urn:lsid:plazi:treatment:CB52563B9D54CA5DFF7CFB0AFB99FF6C:4624B1F69D55CA42FDE0F8DAFD96F8E4" box="[523,629,1881,1907]" name="Itaguacu" pageId="15" pageNumber="16" stateProvince="Espirito Santo">Itaguaçu</location>
</emphasis>
,
<specimenCount id="55FD2CA49D55CA42FD68F8DAFD32F8E4" box="[643,721,1881,1907]" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
</materialsCitation>
,
<date id="3745C1ED9D55CA42FD36F8DAFCBCF8E4" box="[733,863,1881,1907]" pageId="15" pageNumber="16" value="1970-04-18">18.IV.1970</date>
,
<specimenCount id="55FD2CA49D55CA42FC81F8DAFC5AF8E4" box="[874,953,1881,1907]" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
,
<date id="3745C1ED9D55CA42FC2FF8DBFBA4F8E4" box="[964,1095,1880,1907]" pageId="15" pageNumber="16" value="1970-06-15">
15.
<collectionCode id="25EA7FE89D55CA42FC0CF8DAFBE4F8E4" box="[999,1031,1881,1907]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="15" pageNumber="16">VI</collectionCode>
.1970
</date>
,
<specimenCount id="55FD2CA49D55CA42FBB9F8DAFB43F8E4" box="[1106,1184,1881,1907]" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
,
<materialsCitation id="F393ED709D55CA42FB47F8DAFC61F800" ID-GBIF-Occurrence="2248478855" collectingDate="1970-07-29" collectionCode="DZUP" collectorName="C. Elias" location="Linhares" pageId="15" pageNumber="16" specimenCount="1" specimenCount-male="1" stateProvince="Espirito Santo" typeStatus="paratype">
<date id="3745C1ED9D55CA42FB47F8DAFADBF8E4" box="[1196,1336,1881,1907]" pageId="15" pageNumber="16" value="1970-07-29">
<collectingDate id="270138059D55CA42FB47F8DAFADBF8E4" box="[1196,1336,1881,1907]" pageId="15" pageNumber="16" value="1970-07-29">29.VII.1970</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D55CA42FAA8F8DAFA7FF8E4" box="[1347,1436,1881,1907]" pageId="15" pageNumber="16">C. Elias</collectorName>
<emphasis id="718F3B3F9D55CA42FF7CF8FEFF22F800" box="[151,193,1917,1943]" italics="true" pageId="15" pageNumber="16">leg.</emphasis>
, DZ 39.883, DZ 39.880, DZ 41.759 (
<collectionCode id="25EA7FE89D55CA42FDB7F8FEFD48F800" box="[604,683,1917,1943]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="15" pageNumber="16" type="University or college">DZUP</collectionCode>
);
<emphasis id="718F3B3F9D55CA42FD56F8FFFCCBF800" bold="true" box="[701,808,1916,1943]" pageId="15" pageNumber="16">
<location id="4624B1F69D55CA42FD56F8FFFCCBF800" LSID="urn:lsid:plazi:treatment:CB52563B9D54CA5DFF7CFB0AFB99FF6C:4624B1F69D55CA42FD56F8FFFCCBF800" box="[701,808,1916,1943]" name="Linhares" pageId="15" pageNumber="16" stateProvince="Espirito Santo">Linhares</location>
</emphasis>
,
<specimenCount id="55FD2CA49D55CA42FCDEF8FEFC61F800" box="[821,898,1917,1943]" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
</materialsCitation>
,
<date id="3745C1ED9D55CA42FC67F8FEFBE0F800" box="[908,1027,1917,1943]" pageId="15" pageNumber="16" value="1970-04-01">
IV.1970,
<specimenCount id="55FD2CA49D55CA42FC1EF8FEFBE0F800" box="[1013,1027,1917,1943]" pageId="15" pageNumber="16" type="male">1</specimenCount>
</date>
male,
<date id="3745C1ED9D55CA42FBA7F8FEFB21F800" box="[1100,1218,1917,1943]" pageId="15" pageNumber="16" value="1972-02-11">11.II.1972</date>
,
<specimenCount id="55FD2CA49D55CA42FB26F8FEFAF9F800" box="[1229,1306,1917,1943]" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
,
<date id="3745C1ED9D55CA42FACFF8FEFA7BF800" box="[1316,1432,1917,1943]" pageId="15" pageNumber="16" value="1972-04-04">4.IV.1972</date>
,
<specimenCount id="55FD2CA49D55CA42FF7CF822FF1DF82C" box="[151,254,1953,1979]" pageId="15" pageNumber="16" type="female">1 female</specimenCount>
,
<date id="3745C1ED9D55CA42FEE7F822FE66F82C" box="[268,389,1953,1979]" pageId="15" pageNumber="16" value="1972-05-10">10.V.1972</date>
,
<specimenCount id="55FD2CA49D55CA42FE78F822FE07F82C" box="[403,484,1953,1979]" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
,
<date id="3745C1ED9D55CA42FE19F822FD88F82C" box="[498,619,1953,1979]" pageId="15" pageNumber="16" value="1972-05-17">17.V.1972</date>
,
<specimenCount id="55FD2CA49D55CA42FD91F822FD29F82C" box="[634,714,1953,1979]" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
,
<date id="3745C1ED9D55CA42FD32F822FCB8F82C" box="[729,859,1953,1979]" pageId="15" pageNumber="16" value="1972-06-22">
22.
<collectionCode id="25EA7FE89D55CA42FD17F822FCFFF82C" box="[764,796,1953,1979]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="15" pageNumber="16">VI</collectionCode>
.1972
</date>
,
<specimenCount id="55FD2CA49D55CA42FC82F822FC59F82C" box="[873,954,1953,1979]" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
,
<date id="3745C1ED9D55CA42FC23F822FBA8F82C" box="[968,1099,1953,1979]" pageId="15" pageNumber="16" value="1972-09-20">20.IX.1972</date>
,
<specimenCount id="55FD2CA49D55CA42FBB2F822FB49F82C" box="[1113,1194,1953,1979]" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
,
<date id="3745C1ED9D55CA42FB53F822FADBF82C" box="[1208,1336,1953,1979]" pageId="15" pageNumber="16" value="1973-03-22">22.III.1973</date>
,
<specimenCount id="55FD2CA49D55CA42FAACF822FA7BF82C" box="[1351,1432,1953,1979]" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
,
<date id="3745C1ED9D55CA42FF7CF846FEFAF848" box="[151,281,1989,2015]" pageId="15" pageNumber="16" value="1973-04-27">27.IV.1973</date>
,
<specimenCount id="55FD2CA49D55CA42FECDF846FE96F848" box="[294,373,1989,2015]" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
, 16–22.
<date id="3745C1ED9D55CA42FE25F846FDDDF848" box="[462,574,1989,2015]" pageId="15" pageNumber="16" value="1973-05-01">
V.1973,
<specimenCount id="55FD2CA49D55CA42FDDBF846FDDDF848" box="[560,574,1989,2015]" pageId="15" pageNumber="16" type="male">1</specimenCount>
</date>
male, 1–9.
<date id="3745C1ED9D55CA42FD57F846FCD5F848" box="[700,822,1989,2015]" pageId="15" pageNumber="16" value="1973-06-01">
<collectionCode id="25EA7FE89D55CA42FD57F846FD3FF848" box="[700,732,1989,2015]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="15" pageNumber="16">VI</collectionCode>
.1973,
<specimenCount id="55FD2CA49D55CA42FCC3F846FCD5F848" box="[808,822,1989,2015]" pageId="15" pageNumber="16" type="male">1</specimenCount>
</date>
male, 1–8.
<date id="3745C1ED9D55CA42FC44F846FBCEF848" box="[943,1069,1989,2015]" pageId="15" pageNumber="16" value="1973-09-01">
IX.1973,
<specimenCount id="55FD2CA49D55CA42FBF4F846FBCEF848" box="[1055,1069,1989,2015]" pageId="15" pageNumber="16" type="male">1</specimenCount>
</date>
male, 17–22.
<date id="3745C1ED9D55CA42FB2DF846FAA2F848" box="[1222,1345,1989,2015]" pageId="15" pageNumber="16" value="1973-09-01">
IX.1973,
<specimenCount id="55FD2CA49D55CA42FAD8F846FAA2F848" box="[1331,1345,1989,2015]" pageId="15" pageNumber="16" type="female">1</specimenCount>
</date>
female, 1–8.
<date id="3745C1ED9D4ACA5DFF23FF1AFED4FF24" box="[200,311,153,179]" pageId="16" pageNumber="17" value="1973-10-02">
X.1973,
<specimenCount id="55FD2CA49D4ACA5DFEC2FF1AFED4FF24" box="[297,311,153,179]" pageId="16" pageNumber="17" type="male">2</specimenCount>
</date>
males,
<date id="3745C1ED9D4ACA5DFE60FF1AFE15FF24" box="[395,502,153,179]" pageId="16" pageNumber="17" value="1974-05-02">2.V.1974</date>
,
<specimenCount id="55FD2CA49D4ACA5DFDEAFF1AFDADFF24" box="[513,590,153,179]" pageId="16" pageNumber="17" type="male">1 male</specimenCount>
, 2–7.
<date id="3745C1ED9D4ACA5DFD6CFF1AFCE1FF24" box="[647,770,153,179]" pageId="16" pageNumber="17" value="1974-09-01">
IX.1974,
<specimenCount id="55FD2CA49D4ACA5DFD1FFF1AFCE1FF24" box="[756,770,153,179]" pageId="16" pageNumber="17" type="male">1</specimenCount>
</date>
male,
<date id="3745C1ED9D4ACA5DFCA0FF1AFC1BFF24" box="[843,1016,153,179]" pageId="16" pageNumber="17" value="1974-09-23" valueMax="1974-09-30" valueMin="1974-09-23">23–30.IX.1974</date>
, C. Elias
<emphasis id="718F3B3F9D4ACA5DFB8BFF1AFB61FF24" box="[1120,1154,153,179]" italics="true" pageId="16" pageNumber="17">leg</emphasis>
., DZ 41.841, DZ 41.911, DZ 41.851, DZ 41.760, DZ 39.881, DZ 41.821, DZ 41.205, DZ 41.861, DZ 41.811, DZ 41.215, DZ 41.882, DZ 41.769, DZ 41.831, DZ 41.820, DZ 41.901, DZ 41.881, DZ 41.871, DZ 41.810 (
<collectionCode id="25EA7FE89D4ACA5DFBF4FF62FB8CFF6C" box="[1055,1135,225,251]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="16" pageNumber="17" type="University or college">DZUP</collectionCode>
).
</paragraph>
</subSubSection>
</treatment>
</document>

314
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<document id="B2D64BF9A7211E3DB5DCE203F8C12903" ID-CLB-Dataset="26561" ID-DOI="10.11646/zootaxa.4614.1.1" ID-GBIF-Dataset="8e7650ed-66fc-4b2a-b55f-11603d3d2166" ID-ISSN="1175-5326" ID-Zenodo-Dep="3241035" ID-ZooBank="4D7A4B93-7C52-4EF9-92A7-46CA81A1DEC7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1559896932110" checkinUser="plazi" docAuthor="Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik &amp; Casagrande, Mirna Martins" docDate="2019" docId="CB52563B9D56CA41FF7CFF1AFCC6F83A" docLanguage="en" docName="zootaxa.4614.1.1.pdf" docOrigin="Zootaxa 4614 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Artines solange Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande 2019, sp. nov." docType="treatment" docUuid="0B439081-EF4F-41EB-A728-8B7781217566" docUuidSource="ZooBank" docVersion="12" lastPageNumber="13" masterDocId="376B2E439D5ACA4DFFEBFF83FFE3FF97" masterDocTitle="Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species" masterLastPageNumber="49" masterPageNumber="1" pageNumber="13" updateTime="1698734344369" updateUser="ExternalLinkService">
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<mods:title id="C39B71496A07F842A0354B8641309E45">Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species</mods:title>
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<mods:namePart id="D12E888AE659AB42DE51AAFAD5B4FE1E">Medeiros, Adalberto Dantas De</mods:namePart>
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<taxonomicName id="84FB9CAE9D56CA41FF7CFF1AFD91FF24" authority="Medeiros &amp; O. Mielke" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[151,626,153,180]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="solange" status="sp. nov.">
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Medeiros &amp; O. Mielke
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<taxonomicNameLabel id="6ABC86449D56CA41FD91FF19FD34FF23" box="[634,727,154,180]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="0BE1B4A69D56CA41FF7CFF3EFC50FF6B" pageId="12" pageNumber="13" type="description">
<paragraph id="4344E72D9D56CA41FF7CFF3EFE67FF40" blockId="12.[151,947,153,252]" box="[151,388,189,216]" pageId="12" pageNumber="13">
<figureCitation id="DBC0FBA89D56CA41FF7CFF3EFF3DFF4F" box="[151,222,189,216]" captionStart="FIGURES 5–8" captionStartId="6.[151,264,1064,1089]" captionTargetBox="[154,1432,269,1044]" captionTargetId="figure@6.[151,1436,265,1052]" captionTargetPageId="6" captionText="FIGURES 5–8. Species of the “aepitus group” in dorsal and ventral views. 5. Artines aepitus, A–B: male, Cueva de Las Pavas, Tingo Maria, Huánuco, Peru (OM 80.822), C–D: female, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.790). 6. Artines panama sp. nov., A–B: holotype male, C–D: allotype female. 7. Artines solange sp. nov., A–B: holotype male, C–D: tentative female, Sumaúma, Maranhão, Brazil (DZ 41.800). 8. Artines delfos sp. nov., A–B: holotype male. Scale bar = 1 cm." httpUri="https://zenodo.org/record/3241041/files/figure.png" pageId="12" pageNumber="13">Figs 7</figureCitation>
,
<figureCitation id="DBC0FBA89D56CA41FF01FF3EFEEBFF40" box="[234,264,189,215]" captionStart="FIGURES 27–29" captionStartId="23.[151,264,1797,1822]" captionTargetBox="[153,1436,174,1777]" captionTargetId="figure@23.[151,1436,174,1786]" captionTargetPageId="23" captionText="FIGURES 27–29. Male genitalia of the “aepitus group” species. 27. Artines aepitus, Jusepín, Monagas, Venezuela (OM 29.417). 28. Artines panama sp. nov., holotype. 29. Artines solange sp. nov., holotype. A) lateral view of tegumen, saccus, uncus and gnathos. B) ventral view of saccus. C) dorsal view of tegumen, uncus and gnathos. D) ventral view of tegumen, uncus and gnathos. E) interior view of right valva. F) dorsal view of right valva. G) dorsal view of aedeagus and vesica. H) left lateral view of aedeagus. I) ventral view of aedeagus. J) right lateral view of aedeagus. K) dorsal view of fultura inferior. L) lateral view of fultura inferior. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241047/files/figure.png" pageId="12" pageNumber="13">29</figureCitation>
,
<figureCitation id="DBC0FBA89D56CA41FEFFFF3EFED1FF4F" box="[276,306,189,216]" captionStart="FIGURES 54–60" captionStartId="35.[151,264,1807,1832]" captionTargetBox="[156,1436,575,1772]" captionTargetId="figure@35.[151,1436,589,1795]" captionTargetPageId="35" captionText="FIGURES 54–60. Female genitalia of the “aepitus group” species. 54. Artines aepitus, Jusepín, Monagas, Venezuela (OM 30. 125). 55. Artines panama sp. nov., allotype. 56. Artines solange sp. nov. (tentative female), Sumaúma, Maranhão, Brazil (DZ 41.800). 57. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.820). 58. Artines rica, Turrialba, Cartago, Costa Rica (DZ 41.840). 59. Artines liege sp. nov., paratype, Parque do Gama, Brasília, Distrito Federal, Brazil (DZ 41.830). 60. Artines bamba sp. nov. (tentative female), Moyobamba, San Martin, Peru (OM 81.229). A) lateral view of tergum VIII, sterigma and papilla analis; B) ventral view of esterigma. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241059/files/figure.png" pageId="12" pageNumber="13">56</figureCitation>
,
<figureCitation id="DBC0FBA89D56CA41FED5FF3EFEBFFF40" box="[318,348,189,215]" captionStart="FIGURES 66–72" captionStartId="37.[151,264,1734,1759]" captionTargetBox="[160,1433,189,1717]" captionTargetId="figure@37.[151,1436,174,1723]" captionTargetPageId="37" captionText="FIGURES 66–72. Female genitalia of the “aepitus group” species (ventral view of sterigma and bursa copulatrix). 66. Artines aepitus, Jusepín, Monagas, Venezuela (OM 30. 125). 67. Artines panama sp. nov., allotype. 68. Artines solange sp. nov. (tentative female), Sumaúma, Maranhão, Brazil (DZ 41.800). 69. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.820). 70. Artines rica, Turrialba, Cartago, Costa Rica (DZ 41.840). 71. Artines liege sp. nov., paratype, Parque do Gama, Brasília, Distrito Federal, Brazil (DZ 41.830). 72. Artines bamba sp. nov. (tentative female), Moyobamba, San Martin, Peru (OM 81.229). Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241063/files/figure.png" pageId="12" pageNumber="13">68</figureCitation>
,
<figureCitation id="DBC0FBA89D56CA41FE83FF3EFE67FF40" box="[360,388,189,215]" captionStart="FIGURE 78" captionStartId="42.[151,250,910,935]" captionTargetBox="[151,1436,174,898]" captionTargetId="figure@42.[151,1436,174,898]" captionTargetPageId="42" captionText="FIGURE 78. Geographic distribution of the “aepitus group” species. The symbols with white inferior half represent records from literature." httpUri="https://zenodo.org/record/3241067/files/figure.png" pageId="12" pageNumber="13">78</figureCitation>
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<subSubSection id="0BE1B4A69D56CA41FF7CFEBDFBB6FD9A" pageId="12" pageNumber="13" type="diagnosis">
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<emphasis id="718F3B3F9D56CA41FF7CFEBDFEE9FECE" bold="true" box="[151,266,318,345]" pageId="12" pageNumber="13">Diagnosis</emphasis>
.
<taxonomicName id="84FB9CAE9D56CA41FEFDFEBCFE21FECE" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[278,450,319,345]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="solange" status="sp. nov.">
<emphasis id="718F3B3F9D56CA41FEFDFEBCFE21FECE" box="[278,450,319,345]" italics="true" pageId="12" pageNumber="13">Artines solange</emphasis>
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<taxonomicNameLabel id="6ABC86449D56CA41FE2CFEBCFDFCFECE" box="[455,543,319,345]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
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differs from the other species of the “
<taxonomicName id="84FB9CAE9D56CA41FC5DFEC3FBE4FECE" baseAuthorityName="Geyer" baseAuthorityYear="1832" box="[950,1031,320,345]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D56CA41FC5DFEC3FBE4FECE" box="[950,1031,320,345]" italics="true" pageId="12" pageNumber="13">aepitus</emphasis>
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group” in the following combination of characters: ventral forewing with distal portion of discal cell densely covered with orange scales; uncus with disto-lateral lobes; ventral process of harpe wide and longer than tegumen+uncus; dorsal process of harpe absent; sterigma more or less rectangular; lamella postvaginalis with lateral sclerotized folds and large distal projections; ostium rounded, surrounded by sclerotized area.
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<emphasis id="718F3B3F9D56CA41FF2CFE71FEB2FD9A" bold="true" box="[199,337,498,525]" pageId="12" pageNumber="13">Descripiton</emphasis>
. Male (
<figureCitation id="DBC0FBA89D56CA41FE43FE70FE08FD9A" box="[424,491,499,525]" captionStart="FIGURES 5–8" captionStartId="6.[151,264,1064,1089]" captionTargetBox="[154,1432,269,1044]" captionTargetId="figure@6.[151,1436,265,1052]" captionTargetPageId="6" captionText="FIGURES 5–8. Species of the “aepitus group” in dorsal and ventral views. 5. Artines aepitus, A–B: male, Cueva de Las Pavas, Tingo Maria, Huánuco, Peru (OM 80.822), C–D: female, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.790). 6. Artines panama sp. nov., A–B: holotype male, C–D: allotype female. 7. Artines solange sp. nov., A–B: holotype male, C–D: tentative female, Sumaúma, Maranhão, Brazil (DZ 41.800). 8. Artines delfos sp. nov., A–B: holotype male. Scale bar = 1 cm." httpUri="https://zenodo.org/record/3241041/files/figure.png" pageId="12" pageNumber="13">Fig. 7</figureCitation>
A–B).
<emphasis id="718F3B3F9D56CA41FDDCFE70FD97FD9A" box="[567,628,499,525]" italics="true" pageId="12" pageNumber="13">Head</emphasis>
: antenna with nudum bearing12 segments.
</paragraph>
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<subSubSection id="0BE1B4A69D56CA41FF2CFD9BFD3EFB4A" pageId="12" pageNumber="13" type="description">
<paragraph id="4344E72D9D56CA41FF2CFD9BFE05FDA6" blockId="12.[151,1437,318,1965]" box="[199,486,535,561]" pageId="12" pageNumber="13">
<emphasis id="718F3B3F9D56CA41FF2CFD9BFE62FDA6" box="[199,385,535,561]" italics="true" pageId="12" pageNumber="13">Forewing length</emphasis>
:
<quantity id="84034AC89D56CA41FE7BFD94FE01FDA6" box="[400,482,535,561]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="12" pageNumber="13" unit="mm" value="14.0">14 mm</quantity>
.
</paragraph>
<paragraph id="4344E72D9D56CA41FF2CFDB8FA8FFDC2" blockId="12.[151,1437,318,1965]" box="[199,1388,571,602]" pageId="12" pageNumber="13">
<emphasis id="718F3B3F9D56CA41FF2CFDB8FE63FDC2" box="[199,384,571,597]" italics="true" pageId="12" pageNumber="13">Dorsal forewing</emphasis>
(
<figureCitation id="DBC0FBA89D56CA41FE64FDB8FE09FDC2" box="[399,490,571,597]" captionStart="FIGURES 5–8" captionStartId="6.[151,264,1064,1089]" captionTargetBox="[154,1432,269,1044]" captionTargetId="figure@6.[151,1436,265,1052]" captionTargetPageId="6" captionText="FIGURES 5–8. Species of the “aepitus group” in dorsal and ventral views. 5. Artines aepitus, A–B: male, Cueva de Las Pavas, Tingo Maria, Huánuco, Peru (OM 80.822), C–D: female, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.790). 6. Artines panama sp. nov., A–B: holotype male, C–D: allotype female. 7. Artines solange sp. nov., A–B: holotype male, C–D: tentative female, Sumaúma, Maranhão, Brazil (DZ 41.800). 8. Artines delfos sp. nov., A–B: holotype male. Scale bar = 1 cm." httpUri="https://zenodo.org/record/3241041/files/figure.png" pageId="12" pageNumber="13">Fig. 7A</figureCitation>
): three apical spots; discal spots reduced to few scales; spot in CuA
<subScript id="DF7FE5689D56CA41FB35FDCFFB05FDCD" attach="left" box="[1246,1254,588,602]" fontSize="6" pageId="12" pageNumber="13">2</subScript>
–2A absent.
</paragraph>
<paragraph id="4344E72D9D56CA41FF2CFDDCFD57FDEE" blockId="12.[151,1437,318,1965]" box="[199,692,607,633]" pageId="12" pageNumber="13">
<emphasis id="718F3B3F9D56CA41FF2CFDDCFE65FDEE" box="[199,390,607,633]" italics="true" pageId="12" pageNumber="13">Ventral forewing</emphasis>
(
<figureCitation id="DBC0FBA89D56CA41FE7EFDDCFE0DFDEE" box="[405,494,607,633]" captionStart="FIGURES 5–8" captionStartId="6.[151,264,1064,1089]" captionTargetBox="[154,1432,269,1044]" captionTargetId="figure@6.[151,1436,265,1052]" captionTargetPageId="6" captionText="FIGURES 5–8. Species of the “aepitus group” in dorsal and ventral views. 5. Artines aepitus, A–B: male, Cueva de Las Pavas, Tingo Maria, Huánuco, Peru (OM 80.822), C–D: female, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.790). 6. Artines panama sp. nov., A–B: holotype male, C–D: allotype female. 7. Artines solange sp. nov., A–B: holotype male, C–D: tentative female, Sumaúma, Maranhão, Brazil (DZ 41.800). 8. Artines delfos sp. nov., A–B: holotype male. Scale bar = 1 cm." httpUri="https://zenodo.org/record/3241041/files/figure.png" pageId="12" pageNumber="13">Fig. 7B</figureCitation>
): as in
<taxonomicName id="84FB9CAE9D56CA41FDD6FDDCFD4EFDEE" baseAuthorityName="Geyer" baseAuthorityYear="1832" box="[573,685,607,633]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D56CA41FDD6FDDCFD4EFDEE" box="[573,685,607,633]" italics="true" pageId="12" pageNumber="13">A. aepitus</emphasis>
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.
</paragraph>
<paragraph id="4344E72D9D56CA41FF2CFD00FCE3FD56" blockId="12.[151,1437,318,1965]" pageId="12" pageNumber="13">
<emphasis id="718F3B3F9D56CA41FF2CFD00FE77FD0A" box="[199,404,643,669]" italics="true" pageId="12" pageNumber="13">Ventral hind wing</emphasis>
(
<figureCitation id="DBC0FBA89D56CA41FE4EFD00FE1CFD0A" box="[421,511,643,669]" captionStart="FIGURES 5–8" captionStartId="6.[151,264,1064,1089]" captionTargetBox="[154,1432,269,1044]" captionTargetId="figure@6.[151,1436,265,1052]" captionTargetPageId="6" captionText="FIGURES 5–8. Species of the “aepitus group” in dorsal and ventral views. 5. Artines aepitus, A–B: male, Cueva de Las Pavas, Tingo Maria, Huánuco, Peru (OM 80.822), C–D: female, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.790). 6. Artines panama sp. nov., A–B: holotype male, C–D: allotype female. 7. Artines solange sp. nov., A–B: holotype male, C–D: tentative female, Sumaúma, Maranhão, Brazil (DZ 41.800). 8. Artines delfos sp. nov., A–B: holotype male. Scale bar = 1 cm." httpUri="https://zenodo.org/record/3241041/files/figure.png" pageId="12" pageNumber="13">Fig. 7B</figureCitation>
): apical portion of discal cell densely covered by orange to yellow scales; central black spot as wide as long; blue discal spots elongated.
</paragraph>
<paragraph id="4344E72D9D56CA41FF2CFD48FD96FB92" blockId="12.[151,1437,318,1965]" pageId="12" pageNumber="13">
<emphasis id="718F3B3F9D56CA41FF2CFD48FED2FD72" box="[199,305,715,741]" italics="true" pageId="12" pageNumber="13">Genitalia</emphasis>
(
<figureCitation id="DBC0FBA89D56CA41FEA9FD48FE75FD72" box="[322,406,715,741]" captionStart="FIGURES 27–29" captionStartId="23.[151,264,1797,1822]" captionTargetBox="[153,1436,174,1777]" captionTargetId="figure@23.[151,1436,174,1786]" captionTargetPageId="23" captionText="FIGURES 27–29. Male genitalia of the “aepitus group” species. 27. Artines aepitus, Jusepín, Monagas, Venezuela (OM 29.417). 28. Artines panama sp. nov., holotype. 29. Artines solange sp. nov., holotype. A) lateral view of tegumen, saccus, uncus and gnathos. B) ventral view of saccus. C) dorsal view of tegumen, uncus and gnathos. D) ventral view of tegumen, uncus and gnathos. E) interior view of right valva. F) dorsal view of right valva. G) dorsal view of aedeagus and vesica. H) left lateral view of aedeagus. I) ventral view of aedeagus. J) right lateral view of aedeagus. K) dorsal view of fultura inferior. L) lateral view of fultura inferior. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241047/files/figure.png" pageId="12" pageNumber="13">Fig. 29</figureCitation>
A–L): tegumen more or less rectangular, with proximal margin concave and distal margin convex and strongly angled (
<figureCitation id="DBC0FBA89D56CA41FE0EFD6CFDADFC9E" box="[485,590,751,777]" captionStart="FIGURES 27–29" captionStartId="23.[151,264,1797,1822]" captionTargetBox="[153,1436,174,1777]" captionTargetId="figure@23.[151,1436,174,1786]" captionTargetPageId="23" captionText="FIGURES 27–29. Male genitalia of the “aepitus group” species. 27. Artines aepitus, Jusepín, Monagas, Venezuela (OM 29.417). 28. Artines panama sp. nov., holotype. 29. Artines solange sp. nov., holotype. A) lateral view of tegumen, saccus, uncus and gnathos. B) ventral view of saccus. C) dorsal view of tegumen, uncus and gnathos. D) ventral view of tegumen, uncus and gnathos. E) interior view of right valva. F) dorsal view of right valva. G) dorsal view of aedeagus and vesica. H) left lateral view of aedeagus. I) ventral view of aedeagus. J) right lateral view of aedeagus. K) dorsal view of fultura inferior. L) lateral view of fultura inferior. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241047/files/figure.png" pageId="12" pageNumber="13">Fig. 29C</figureCitation>
), in lateral view larger than length of gnathos; ventral arm of tegumen as long as dorsal arm of saccus (
<figureCitation id="DBC0FBA89D56CA41FE0CFC90FDB3FCBA" box="[487,592,787,813]" captionStart="FIGURES 27–29" captionStartId="23.[151,264,1797,1822]" captionTargetBox="[153,1436,174,1777]" captionTargetId="figure@23.[151,1436,174,1786]" captionTargetPageId="23" captionText="FIGURES 27–29. Male genitalia of the “aepitus group” species. 27. Artines aepitus, Jusepín, Monagas, Venezuela (OM 29.417). 28. Artines panama sp. nov., holotype. 29. Artines solange sp. nov., holotype. A) lateral view of tegumen, saccus, uncus and gnathos. B) ventral view of saccus. C) dorsal view of tegumen, uncus and gnathos. D) ventral view of tegumen, uncus and gnathos. E) interior view of right valva. F) dorsal view of right valva. G) dorsal view of aedeagus and vesica. H) left lateral view of aedeagus. I) ventral view of aedeagus. J) right lateral view of aedeagus. K) dorsal view of fultura inferior. L) lateral view of fultura inferior. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241047/files/figure.png" pageId="12" pageNumber="13">Fig. 29A</figureCitation>
). Saccus shorter than tegumen+uncus, base larger than lateral margin (
<figureCitation id="DBC0FBA89D56CA41FA84FC90FF2AFCC6" captionStart="FIGURES 27–29" captionStartId="23.[151,264,1797,1822]" captionTargetBox="[153,1436,174,1777]" captionTargetId="figure@23.[151,1436,174,1786]" captionTargetPageId="23" captionText="FIGURES 27–29. Male genitalia of the “aepitus group” species. 27. Artines aepitus, Jusepín, Monagas, Venezuela (OM 29.417). 28. Artines panama sp. nov., holotype. 29. Artines solange sp. nov., holotype. A) lateral view of tegumen, saccus, uncus and gnathos. B) ventral view of saccus. C) dorsal view of tegumen, uncus and gnathos. D) ventral view of tegumen, uncus and gnathos. E) interior view of right valva. F) dorsal view of right valva. G) dorsal view of aedeagus and vesica. H) left lateral view of aedeagus. I) ventral view of aedeagus. J) right lateral view of aedeagus. K) dorsal view of fultura inferior. L) lateral view of fultura inferior. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241047/files/figure.png" pageId="12" pageNumber="13">Fig. 29B</figureCitation>
). Uncus, in dorsal view, quadrate, shorter than tegumen and with two ventrally curved disto-lateral lobes (
<figureCitation id="DBC0FBA89D56CA41FA84FCB4FF29FCE2" captionStart="FIGURES 27–29" captionStartId="23.[151,264,1797,1822]" captionTargetBox="[153,1436,174,1777]" captionTargetId="figure@23.[151,1436,174,1786]" captionTargetPageId="23" captionText="FIGURES 27–29. Male genitalia of the “aepitus group” species. 27. Artines aepitus, Jusepín, Monagas, Venezuela (OM 29.417). 28. Artines panama sp. nov., holotype. 29. Artines solange sp. nov., holotype. A) lateral view of tegumen, saccus, uncus and gnathos. B) ventral view of saccus. C) dorsal view of tegumen, uncus and gnathos. D) ventral view of tegumen, uncus and gnathos. E) interior view of right valva. F) dorsal view of right valva. G) dorsal view of aedeagus and vesica. H) left lateral view of aedeagus. I) ventral view of aedeagus. J) right lateral view of aedeagus. K) dorsal view of fultura inferior. L) lateral view of fultura inferior. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241047/files/figure.png" pageId="12" pageNumber="13">Fig. 29C</figureCitation>
). Gnathos as in the other species of the group (
<figureCitation id="DBC0FBA89D56CA41FD13FCD8FC69FCE2" box="[760,906,859,885]" captionStart="FIGURES 27–29" captionStartId="23.[151,264,1797,1822]" captionTargetBox="[153,1436,174,1777]" captionTargetId="figure@23.[151,1436,174,1786]" captionTargetPageId="23" captionText="FIGURES 27–29. Male genitalia of the “aepitus group” species. 27. Artines aepitus, Jusepín, Monagas, Venezuela (OM 29.417). 28. Artines panama sp. nov., holotype. 29. Artines solange sp. nov., holotype. A) lateral view of tegumen, saccus, uncus and gnathos. B) ventral view of saccus. C) dorsal view of tegumen, uncus and gnathos. D) ventral view of tegumen, uncus and gnathos. E) interior view of right valva. F) dorsal view of right valva. G) dorsal view of aedeagus and vesica. H) left lateral view of aedeagus. I) ventral view of aedeagus. J) right lateral view of aedeagus. K) dorsal view of fultura inferior. L) lateral view of fultura inferior. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241047/files/figure.png" pageId="12" pageNumber="13">Fig. 29A, D</figureCitation>
). Valva approximately four times longer than wide; sacculus elongated, about 1/3 of width of valva; costa rectangular and not contiguous with ampulla; ampulla rectangular, longer than costa and directed distally; ventral process of harpe wide, longer than tegumen+uncus, distal 1/3 not dilated and approximately rounded, with apical setae thick and long; dorsal process absent (
<figureCitation id="DBC0FBA89D56CA41FA84FC44FF57FB92" captionStart="FIGURES 27–29" captionStartId="23.[151,264,1797,1822]" captionTargetBox="[153,1436,174,1777]" captionTargetId="figure@23.[151,1436,174,1786]" captionTargetPageId="23" captionText="FIGURES 27–29. Male genitalia of the “aepitus group” species. 27. Artines aepitus, Jusepín, Monagas, Venezuela (OM 29.417). 28. Artines panama sp. nov., holotype. 29. Artines solange sp. nov., holotype. A) lateral view of tegumen, saccus, uncus and gnathos. B) ventral view of saccus. C) dorsal view of tegumen, uncus and gnathos. D) ventral view of tegumen, uncus and gnathos. E) interior view of right valva. F) dorsal view of right valva. G) dorsal view of aedeagus and vesica. H) left lateral view of aedeagus. I) ventral view of aedeagus. J) right lateral view of aedeagus. K) dorsal view of fultura inferior. L) lateral view of fultura inferior. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241047/files/figure.png" pageId="12" pageNumber="13">Fig. 29</figureCitation>
E–F). Aedeagus lost during preparation.
</paragraph>
<paragraph id="4344E72D9D56CA41FF2CFB8CFC72FBBE" blockId="12.[151,1437,318,1965]" box="[199,913,1039,1065]" pageId="12" pageNumber="13">
Female (
<figureCitation id="DBC0FBA89D56CA41FEC2FB8CFE88FBBE" box="[297,363,1039,1065]" captionStart="FIGURES 5–8" captionStartId="6.[151,264,1064,1089]" captionTargetBox="[154,1432,269,1044]" captionTargetId="figure@6.[151,1436,265,1052]" captionTargetPageId="6" captionText="FIGURES 5–8. Species of the “aepitus group” in dorsal and ventral views. 5. Artines aepitus, A–B: male, Cueva de Las Pavas, Tingo Maria, Huánuco, Peru (OM 80.822), C–D: female, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.790). 6. Artines panama sp. nov., A–B: holotype male, C–D: allotype female. 7. Artines solange sp. nov., A–B: holotype male, C–D: tentative female, Sumaúma, Maranhão, Brazil (DZ 41.800). 8. Artines delfos sp. nov., A–B: holotype male. Scale bar = 1 cm." httpUri="https://zenodo.org/record/3241041/files/figure.png" pageId="12" pageNumber="13">Fig. 7</figureCitation>
C–D).
<emphasis id="718F3B3F9D56CA41FE5CFB93FD92FBBE" box="[439,625,1039,1065]" italics="true" pageId="12" pageNumber="13">Forewing length</emphasis>
:
<quantity id="84034AC89D56CA41FD6BFB8CFD31FBBE" box="[640,722,1039,1065]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="12" pageNumber="13" unit="mm" value="13.0">13 mm</quantity>
. Similar to male.
</paragraph>
<paragraph id="4344E72D9D56CA41FF2CFBB0FBCEFB2E" blockId="12.[151,1437,318,1965]" pageId="12" pageNumber="13">
<emphasis id="718F3B3F9D56CA41FF2CFBB0FED2FBDA" box="[199,305,1075,1101]" italics="true" pageId="12" pageNumber="13">Genitalia</emphasis>
(
<figureCitation id="DBC0FBA89D56CA41FED6FBB0FE73FBDA" box="[317,400,1074,1101]" captionStart="FIGURES 54–60" captionStartId="35.[151,264,1807,1832]" captionTargetBox="[156,1436,575,1772]" captionTargetId="figure@35.[151,1436,589,1795]" captionTargetPageId="35" captionText="FIGURES 54–60. Female genitalia of the “aepitus group” species. 54. Artines aepitus, Jusepín, Monagas, Venezuela (OM 30. 125). 55. Artines panama sp. nov., allotype. 56. Artines solange sp. nov. (tentative female), Sumaúma, Maranhão, Brazil (DZ 41.800). 57. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.820). 58. Artines rica, Turrialba, Cartago, Costa Rica (DZ 41.840). 59. Artines liege sp. nov., paratype, Parque do Gama, Brasília, Distrito Federal, Brazil (DZ 41.830). 60. Artines bamba sp. nov. (tentative female), Moyobamba, San Martin, Peru (OM 81.229). A) lateral view of tergum VIII, sterigma and papilla analis; B) ventral view of esterigma. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241059/files/figure.png" pageId="12" pageNumber="13">Figs 56</figureCitation>
A–B, 68): tergum VIII as in
<taxonomicName id="84FB9CAE9D56CA41FD55FBB0FCCFFBDA" baseAuthorityName="Geyer" baseAuthorityYear="1832" box="[702,812,1075,1101]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D56CA41FD55FBB0FCCFFBDA" box="[702,812,1075,1101]" italics="true" pageId="12" pageNumber="13">A. aepitus</emphasis>
</taxonomicName>
. Sterigma more or less rectangular; lamella antevaginalis without proximo-ventral folds; lamella postvaginalis with lateral fold, distal projections wide and rounded, medially separated by shallow and narrow indentation, proximo-lateral projections weakly developed and not bilobed. Ostium rounded, surrounded by sclerotized area; ductus bursae without sclerotized plates.
</paragraph>
<paragraph id="4344E72D9D56CA41FF2CFB41FD3EFB4A" blockId="12.[151,1437,318,1965]" box="[199,733,1218,1245]" pageId="12" pageNumber="13">
<emphasis id="718F3B3F9D56CA41FF2CFB41FED9FB4A" bold="true" box="[199,314,1218,1245]" pageId="12" pageNumber="13">Variation</emphasis>
. No relevant variation was observed.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D56CA41FF2CFB65FD23F9D2" pageId="12" pageNumber="13" type="discussion">
<paragraph id="4344E72D9D56CA41FF2CFB65FD6FFAFA" blockId="12.[151,1437,318,1965]" pageId="12" pageNumber="13">
<emphasis id="718F3B3F9D56CA41FF2CFB65FEABFA96" bold="true" box="[199,328,1254,1281]" pageId="12" pageNumber="13">Comments</emphasis>
.
<taxonomicName id="84FB9CAE9D56CA41FEBCFB64FDE4FA96" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[343,519,1255,1281]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="solange" status="sp. nov.">
<emphasis id="718F3B3F9D56CA41FEBCFB64FDE4FA96" box="[343,519,1255,1281]" italics="true" pageId="12" pageNumber="13">Artines solange</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D56CA41FDE4FB64FD88FA96" bold="true" box="[527,619,1255,1281]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="6ABC86449D56CA41FDE4FB64FD88FA96" box="[527,619,1255,1281]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is one of the rarest or the least collected species of the “
<taxonomicName id="84FB9CAE9D56CA41FB04FB6BFAA3FA96" baseAuthorityName="Geyer" baseAuthorityYear="1832" box="[1263,1344,1256,1281]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D56CA41FB04FB6BFAA3FA96" box="[1263,1344,1256,1281]" italics="true" pageId="12" pageNumber="13">aepitus</emphasis>
</taxonomicName>
group”. It is also one of the easiest to recognize based on the male and female genitalia. Even though the aedeagus was lost during preparation, before that the general pattern of the “
<taxonomicName id="84FB9CAE9D56CA41FCDAFAB3FC61FADE" baseAuthorityName="Geyer" baseAuthorityYear="1832" box="[817,898,1328,1353]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D56CA41FCDAFAB3FC61FADE" box="[817,898,1328,1353]" italics="true" pageId="12" pageNumber="13">aepitus</emphasis>
</taxonomicName>
group” species were observed, particularly the narrow, simple and elongate ventral process.
</paragraph>
<paragraph id="4344E72D9D56CA41FF24FAF4FADCFA6A" blockId="12.[151,1437,318,1965]" pageId="12" pageNumber="13">
The association with the female was based on the color of the ventral hind wing, such as the distal portion of the discal cell densely covered with orange scales and central black spot as wide as long, besides collecting locality. However, in view of the fact that
<taxonomicName id="84FB9CAE9D56CA41FDE4FA3CFD83FA4E" authorityName="Godman" authorityYear="1901" box="[527,608,1471,1497]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="718F3B3F9D56CA41FDE4FA3CFD83FA4E" box="[527,608,1471,1497]" italics="true" pageId="12" pageNumber="13">Artines</emphasis>
</taxonomicName>
species are similar and different species can potentially occur in the same locality, the association is tentative and for this reason the only female was not included in the type-series.
</paragraph>
<paragraph id="4344E72D9D56CA41FF2CF985FD23F9D2" blockId="12.[151,1437,318,1965]" pageId="12" pageNumber="13">
<emphasis id="718F3B3F9D56CA41FF2CF985FE0FF9B6" bold="true" box="[199,492,1542,1569]" pageId="12" pageNumber="13">Geographic distribution</emphasis>
(
<figureCitation id="DBC0FBA89D56CA41FE15F984FDB5F9B6" box="[510,598,1543,1569]" captionStart="FIGURE 78" captionStartId="42.[151,250,910,935]" captionTargetBox="[151,1436,174,898]" captionTargetId="figure@42.[151,1436,174,898]" captionTargetPageId="42" captionText="FIGURE 78. Geographic distribution of the “aepitus group” species. The symbols with white inferior half represent records from literature." httpUri="https://zenodo.org/record/3241067/files/figure.png" pageId="12" pageNumber="13">Fig. 78</figureCitation>
). The species is known only from east of Brazilian Amazon Rainforest. BRAZIL—
<emphasis id="718F3B3F9D56CA41FEF3F9A8FE6DF9D2" box="[280,398,1579,1605]" italics="true" pageId="12" pageNumber="13">
<collectingRegion id="813F29CF9D56CA41FEF3F9A8FE6DF9D2" box="[280,398,1579,1605]" country="Brazil" name="Maranhao" pageId="12" pageNumber="13">Maranhão</collectingRegion>
</emphasis>
: Imperatriz and Sumaúma.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D56CA41FF2CF9CCFC78F9FE" box="[199,923,1614,1641]" pageId="12" pageNumber="13" type="description">
<paragraph id="4344E72D9D56CA41FF2CF9CCFC78F9FE" blockId="12.[151,1437,318,1965]" box="[199,923,1614,1641]" pageId="12" pageNumber="13">
<emphasis id="718F3B3F9D56CA41FF2CF9CCFE31F9FE" bold="true" box="[199,466,1614,1641]" pageId="12" pageNumber="13">Temporal distribution</emphasis>
. This species occurs in July and August.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D56CA41FF2CF9F0FE2AF91A" box="[199,457,1651,1677]" pageId="12" pageNumber="13" type="biology_ecology">
<paragraph id="4344E72D9D56CA41FF2CF9F0FE2AF91A" blockId="12.[151,1437,318,1965]" box="[199,457,1651,1677]" pageId="12" pageNumber="13">
<emphasis id="718F3B3F9D56CA41FF2CF9F0FEA7F91A" bold="true" box="[199,324,1651,1677]" pageId="12" pageNumber="13">Host plant</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D56CA41FF2CF914FC54F96E" pageId="12" pageNumber="13" type="etymology">
<paragraph id="4344E72D9D56CA41FF2CF914FC54F96E" blockId="12.[151,1437,318,1965]" pageId="12" pageNumber="13">
<emphasis id="718F3B3F9D56CA41FF2CF914FEABF926" bold="true" box="[199,328,1687,1713]" pageId="12" pageNumber="13">Etymology</emphasis>
. The name of the new species is in honor to Dr. Solange Maria Kerpel, who has been contributing to the study of butterflies in the Brazilian northeast, in addition to her incentive to the first author in getting start the study on the taxonomy of butterflies. The name is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D56CA41FF2CF880FCC6F83A" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="4344E72D9D56CA41FF2CF880FC08F8F2" blockId="12.[151,1437,318,1965]" pageId="12" pageNumber="13">
<emphasis id="718F3B3F9D56CA41FF2CF880FE8CF88A" bold="true" box="[199,367,1794,1821]" pageId="12" pageNumber="13">Type material</emphasis>
.
<materialsCitation id="F393ED709D56CA41FE97F880FBACF88A" ID-GBIF-Occurrence="2248478881" box="[380,1103,1795,1821]" collectionCode="DZUP" pageId="12" pageNumber="13" specimenCount="1" typeStatus="holotype">
<typeStatus id="9C40598F9D56CA41FE97F880FE07F88A" box="[380,484,1795,1821]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
male deposited at the
<collectionCode id="25EA7FE89D56CA41FD30F880FCC7F88A" box="[731,804,1795,1821]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="12" pageNumber="13" type="University or college">DZUP</collectionCode>
with the followings labels
</materialsCitation>
: /
<materialsCitation id="F393ED709D56CA41FB8DF880FEFCF8F2" ID-GBIF-Occurrence="2248478864" collectingDate="1974-07-15" collectorName="Imperatriz, MA &amp; Exc &amp; Dep &amp; Zool &amp; A. Medeiros" country="Brazil" location="Brazil" pageId="12" pageNumber="13" specimenCount="1" typeStatus="holotype">
<typeStatus id="9C40598F9D56CA41FB8DF880FAEFF88A" box="[1126,1292,1795,1821]" pageId="12" pageNumber="13" type="holotype">HOLOTYPUS</typeStatus>
/
<collectorName id="EE0E82FB9D56CA41FACAF880FF25F8D6" pageId="12" pageNumber="13">Imperatriz, MA</collectorName>
[ranhão,
<collectingCountry id="3BECA7BD9D56CA41FEC3F8A5FE91F8D6" box="[296,370,1830,1857]" name="Brazil" pageId="12" pageNumber="13">Brazil</collectingCountry>
],
<date id="3745C1ED9D56CA41FE68F8A5FDECF8D6" box="[387,527,1830,1857]" pageId="12" pageNumber="13" value="1974-07-15">
<collectingDate id="270138059D56CA41FE68F8A5FDECF8D6" box="[387,527,1830,1857]" pageId="12" pageNumber="13" value="1974-07-15">15.VII.1974</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D56CA41FDF1F8A5FDABF8D6" box="[538,584,1830,1857]" pageId="12" pageNumber="13">Exc</collectorName>
[ursão]
<collectorName id="EE0E82FB9D56CA41FD70F8A5FD2EF8D6" box="[667,717,1830,1857]" pageId="12" pageNumber="13">Dep</collectorName>
[ar]t[ament]o [de]
<collectorName id="EE0E82FB9D56CA41FC71F8A5FC31F8D6" box="[922,978,1830,1857]" pageId="12" pageNumber="13">Zool</collectorName>
[ogia]
<emphasis id="718F3B3F9D56CA41FBF3F8A4FBD9F8D6" box="[1048,1082,1831,1857]" italics="true" pageId="12" pageNumber="13">leg</emphasis>
., / gen. prep.
<collectorName id="EE0E82FB9D56CA41FB39F8A4FABDF8D6" box="[1234,1374,1831,1857]" pageId="12" pageNumber="13">A. Medeiros</collectorName>
2018 / DZ 41.889
</materialsCitation>
/
<typeStatus id="9C40598F9D56CA41FEDEF8C8FE49F8F2" box="[309,426,1867,1893]" pageId="12" pageNumber="13" type="holotype">Holotypus</typeStatus>
<taxonomicName id="84FB9CAE9D56CA41FE5AF8C8FC86F8F2" authority="Medeiros &amp; O. Mielke" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[433,869,1867,1893]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="solange">
<emphasis id="718F3B3F9D56CA41FE5AF8C8FD83F8F2" box="[433,608,1867,1893]" italics="true" pageId="12" pageNumber="13">Artines solange</emphasis>
Medeiros &amp; O. Mielke
</taxonomicName>
det. 2018 /.
</paragraph>
<paragraph id="4344E72D9D56CA41FF2CF8EDFCC6F83A" blockId="12.[151,1437,318,1965]" pageId="12" pageNumber="13">
<emphasis id="718F3B3F9D56CA41FF2CF8EDFE50F81E" bold="true" box="[199,435,1902,1929]" pageId="12" pageNumber="13">Additional material</emphasis>
(only female). BRAZIL—
<emphasis id="718F3B3F9D56CA41FD0AF8ECFCB4F81E" box="[737,855,1903,1929]" italics="true" pageId="12" pageNumber="13">
<collectingRegion id="813F29CF9D56CA41FD0AF8ECFCB4F81E" box="[737,855,1903,1929]" country="Brazil" name="Maranhao" pageId="12" pageNumber="13">Maranhão</collectingRegion>
</emphasis>
:
<emphasis id="718F3B3F9D56CA41FC8DF8EDFC3CF81E" bold="true" box="[870,991,1902,1929]" pageId="12" pageNumber="13">Sumaúma</emphasis>
(formerly a district of Montes Altos),
<specimenCount id="55FD2CA49D56CA41FA65F8ECFF05F83A" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
,
<date id="3745C1ED9D56CA41FF1AF810FE65F83A" box="[241,390,1939,1965]" pageId="12" pageNumber="13" value="1974-08-10">10.VIII.1974</date>
, O. Mielke
<emphasis id="718F3B3F9D56CA41FDE1F810FDCFF83A" box="[522,556,1939,1965]" italics="true" pageId="12" pageNumber="13">leg</emphasis>
., DZ 41.800 (
<collectionCode id="25EA7FE89D56CA41FD22F810FCFAF83A" box="[713,793,1939,1965]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="12" pageNumber="13" type="University or college">DZUP</collectionCode>
).
</paragraph>
</subSubSection>
</treatment>
</document>

273
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<mods:title id="0650027D7789292AA8CF3C7407026016">Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species</mods:title>
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<heading id="180C50419D57CA40FF7CFF1AFD3FFF23" bold="true" box="[151,732,153,180]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<emphasis id="718F3B3F9D57CA40FF7CFF1AFD3FFF23" bold="true" box="[151,732,153,180]" pageId="13" pageNumber="14">
<taxonomicName id="84FB9CAE9D57CA40FF7CFF1AFD94FF24" authority="Medeiros &amp; Casagrande" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[151,631,153,180]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="delfos" status="sp. nov.">
<emphasis id="718F3B3F9D57CA40FF7CFF1AFEDCFF23" bold="true" box="[151,319,153,180]" italics="true" pageId="13" pageNumber="14">Artines delfos</emphasis>
Medeiros &amp; Casagrande
</taxonomicName>
<taxonomicNameLabel id="6ABC86449D57CA40FD94FF19FD3FFF23" box="[639,732,154,180]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="0BE1B4A69D57CA40FF7CFF3EFC3DFF6C" pageId="13" pageNumber="14" type="description">
<paragraph id="4344E72D9D57CA40FF7CFF3EFEB9FF40" blockId="13.[151,990,153,251]" box="[151,346,189,215]" pageId="13" pageNumber="14">
<figureCitation id="DBC0FBA89D57CA40FF7CFF3EFF3EFF40" box="[151,221,189,215]" captionStart="FIGURES 5–8" captionStartId="6.[151,264,1064,1089]" captionTargetBox="[154,1432,269,1044]" captionTargetId="figure@6.[151,1436,265,1052]" captionTargetPageId="6" captionText="FIGURES 5–8. Species of the “aepitus group” in dorsal and ventral views. 5. Artines aepitus, A–B: male, Cueva de Las Pavas, Tingo Maria, Huánuco, Peru (OM 80.822), C–D: female, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.790). 6. Artines panama sp. nov., A–B: holotype male, C–D: allotype female. 7. Artines solange sp. nov., A–B: holotype male, C–D: tentative female, Sumaúma, Maranhão, Brazil (DZ 41.800). 8. Artines delfos sp. nov., A–B: holotype male. Scale bar = 1 cm." httpUri="https://zenodo.org/record/3241041/files/figure.png" pageId="13" pageNumber="14">Figs 8</figureCitation>
,
<figureCitation id="DBC0FBA89D57CA40FF01FF3EFEEBFF40" box="[234,264,189,215]" captionStart="FIGURES 30–32" captionStartId="24.[151,264,1772,1797]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure@24.[151,1436,181,1746]" captionTargetPageId="24" captionText="FIGURES 30–32. Male genitalia of the “aepitus group” species. 30. Artines delfos sp. nov., holotype. 31. Artines maya, holotype. 32. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.215). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241049/files/figure.png" pageId="13" pageNumber="14">30</figureCitation>
,
<figureCitation id="DBC0FBA89D57CA40FEFFFF3EFED0FF40" box="[276,307,189,215]" captionStart="FIGURES 42–48" captionStartId="33.[151,264,1892,1917]" captionTargetBox="[151,1431,625,1874]" captionTargetId="figure@33.[151,1436,625,1880]" captionText="FIGURES 42–48. Dorsal view of vesica and cornutus of the &quot;aepitus group&quot; species. 42. Artines aepitus. 43. Artines panama sp. nov.. 44. Artines delfos sp. nov.. 45. Artines litoralis sp. nov.. 46. Artines rica. 47. Artines liege sp. nov. 48. Artines bamba sp. nov. Scale bar = 0,2 mm." httpUri="https://zenodo.org/record/3406274/files/figure.png" pageId="13" pageNumber="14">44</figureCitation>
, 78
</paragraph>
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<subSubSection id="0BE1B4A69D57CA40FF7CFEBDFB74FE36" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="4344E72D9D57CA40FF7CFEBDFB74FE36" blockId="13.[151,1437,318,1569]" pageId="13" pageNumber="14">
<emphasis id="718F3B3F9D57CA40FF7CFEBDFEE9FECE" bold="true" box="[151,266,318,345]" pageId="13" pageNumber="14">Diagnosis</emphasis>
.
<taxonomicName id="84FB9CAE9D57CA40FEFCFEBCFE52FECE" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[279,433,319,345]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="delfos" status="sp. nov.">
<emphasis id="718F3B3F9D57CA40FEFCFEBCFE52FECE" box="[279,433,319,345]" italics="true" pageId="13" pageNumber="14">Artines delfos</emphasis>
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<emphasis id="718F3B3F9D57CA40FE5DFEBCFDF3FECE" bold="true" box="[438,528,319,345]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="6ABC86449D57CA40FE5DFEBCFDF3FECE" box="[438,528,319,345]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from the other species of the “
<taxonomicName id="84FB9CAE9D57CA40FC5BFEC3FBE2FECE" baseAuthorityName="Geyer" baseAuthorityYear="1832" box="[944,1025,320,345]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D57CA40FC5BFEC3FBE2FECE" box="[944,1025,320,345]" italics="true" pageId="13" pageNumber="14">aepitus</emphasis>
</taxonomicName>
group” in the following combination of characters: valva three times longer than wide; harpe with ventral process wide, shorter than tegumen+uncus, distal 1/3 angular; dorsal process of harpe over proximal 1/2 and distally directed ventrally.
</paragraph>
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<emphasis id="718F3B3F9D57CA40FF2CFE29FEB2FE52" bold="true" box="[199,337,426,453]" pageId="13" pageNumber="14">Description</emphasis>
. Male (
<figureCitation id="DBC0FBA89D57CA40FE43FE28FE09FE52" box="[424,490,427,453]" captionStart="FIGURES 5–8" captionStartId="6.[151,264,1064,1089]" captionTargetBox="[154,1432,269,1044]" captionTargetId="figure@6.[151,1436,265,1052]" captionTargetPageId="6" captionText="FIGURES 5–8. Species of the “aepitus group” in dorsal and ventral views. 5. Artines aepitus, A–B: male, Cueva de Las Pavas, Tingo Maria, Huánuco, Peru (OM 80.822), C–D: female, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.790). 6. Artines panama sp. nov., A–B: holotype male, C–D: allotype female. 7. Artines solange sp. nov., A–B: holotype male, C–D: tentative female, Sumaúma, Maranhão, Brazil (DZ 41.800). 8. Artines delfos sp. nov., A–B: holotype male. Scale bar = 1 cm." httpUri="https://zenodo.org/record/3241041/files/figure.png" pageId="13" pageNumber="14">Fig. 8</figureCitation>
A–B).
<emphasis id="718F3B3F9D57CA40FDDCFE28FD97FE52" box="[567,628,427,453]" italics="true" pageId="13" pageNumber="14">Head</emphasis>
: antenna with nudum bearing 12 segments.
</paragraph>
<paragraph id="4344E72D9D57CA40FF2CFE53FE05FE7E" blockId="13.[151,1437,318,1569]" box="[199,486,463,489]" pageId="13" pageNumber="14">
<emphasis id="718F3B3F9D57CA40FF2CFE53FE62FE7E" box="[199,385,463,489]" italics="true" pageId="13" pageNumber="14">Forewing length</emphasis>
:
<quantity id="84034AC89D57CA40FE7BFE4CFE01FE7E" box="[400,482,463,489]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="13" pageNumber="14" unit="mm" value="14.0">14 mm</quantity>
.
</paragraph>
<paragraph id="4344E72D9D57CA40FF2CFE70FE94FDA6" blockId="13.[151,1437,318,1569]" pageId="13" pageNumber="14">
<emphasis id="718F3B3F9D57CA40FF2CFE70FE9CFD9A" box="[199,383,499,525]" italics="true" pageId="13" pageNumber="14">Dorsal forewing</emphasis>
(
<figureCitation id="DBC0FBA89D57CA40FE66FE70FE0BFD9A" box="[397,488,499,525]" captionStart="FIGURES 5–8" captionStartId="6.[151,264,1064,1089]" captionTargetBox="[154,1432,269,1044]" captionTargetId="figure@6.[151,1436,265,1052]" captionTargetPageId="6" captionText="FIGURES 5–8. Species of the “aepitus group” in dorsal and ventral views. 5. Artines aepitus, A–B: male, Cueva de Las Pavas, Tingo Maria, Huánuco, Peru (OM 80.822), C–D: female, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.790). 6. Artines panama sp. nov., A–B: holotype male, C–D: allotype female. 7. Artines solange sp. nov., A–B: holotype male, C–D: tentative female, Sumaúma, Maranhão, Brazil (DZ 41.800). 8. Artines delfos sp. nov., A–B: holotype male. Scale bar = 1 cm." httpUri="https://zenodo.org/record/3241041/files/figure.png" pageId="13" pageNumber="14">Fig. 8A</figureCitation>
): two apical spots reduced to few scales; discal spot in M
<subScript id="DF7FE5689D57CA40FB8AFD87FB8AFD85" attach="left" box="[1121,1129,516,530]" fontSize="6" pageId="13" pageNumber="14">3</subScript>
–CuA
<subScript id="DF7FE5689D57CA40FB47FD87FB57FD85" attach="left" box="[1196,1204,516,530]" fontSize="6" pageId="13" pageNumber="14">1</subScript>
slightly arched; spot in CuA
<subScript id="DF7FE5689D57CA40FF02FDABFF12FDA1" attach="left" box="[233,241,552,566]" fontSize="6" pageId="13" pageNumber="14">2</subScript>
–2A absent.
</paragraph>
<paragraph id="4344E72D9D57CA40FF2CFDB8FA98FDC2" blockId="13.[151,1437,318,1569]" box="[199,1403,571,597]" pageId="13" pageNumber="14">
<emphasis id="718F3B3F9D57CA40FF2CFDB8FE65FDC2" box="[199,390,571,597]" italics="true" pageId="13" pageNumber="14">Ventral forewing</emphasis>
(
<figureCitation id="DBC0FBA89D57CA40FE7EFDB8FE0DFDC2" box="[405,494,571,597]" captionStart="FIGURES 5–8" captionStartId="6.[151,264,1064,1089]" captionTargetBox="[154,1432,269,1044]" captionTargetId="figure@6.[151,1436,265,1052]" captionTargetPageId="6" captionText="FIGURES 5–8. Species of the “aepitus group” in dorsal and ventral views. 5. Artines aepitus, A–B: male, Cueva de Las Pavas, Tingo Maria, Huánuco, Peru (OM 80.822), C–D: female, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.790). 6. Artines panama sp. nov., A–B: holotype male, C–D: allotype female. 7. Artines solange sp. nov., A–B: holotype male, C–D: tentative female, Sumaúma, Maranhão, Brazil (DZ 41.800). 8. Artines delfos sp. nov., A–B: holotype male. Scale bar = 1 cm." httpUri="https://zenodo.org/record/3241041/files/figure.png" pageId="13" pageNumber="14">Fig. 8B</figureCitation>
): a row of four blue apical spots directed toward median portion of outer margin.
</paragraph>
<paragraph id="4344E72D9D57CA40FF2CFDDCFEB8FD0A" blockId="13.[151,1437,318,1569]" pageId="13" pageNumber="14">
<emphasis id="718F3B3F9D57CA40FF2CFDDCFE73FDEE" box="[199,400,607,633]" italics="true" pageId="13" pageNumber="14">Ventral hind wing</emphasis>
(
<figureCitation id="DBC0FBA89D57CA40FE74FDDCFE14FDEE" box="[415,503,607,633]" captionStart="FIGURES 5–8" captionStartId="6.[151,264,1064,1089]" captionTargetBox="[154,1432,269,1044]" captionTargetId="figure@6.[151,1436,265,1052]" captionTargetPageId="6" captionText="FIGURES 5–8. Species of the “aepitus group” in dorsal and ventral views. 5. Artines aepitus, A–B: male, Cueva de Las Pavas, Tingo Maria, Huánuco, Peru (OM 80.822), C–D: female, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.790). 6. Artines panama sp. nov., A–B: holotype male, C–D: allotype female. 7. Artines solange sp. nov., A–B: holotype male, C–D: tentative female, Sumaúma, Maranhão, Brazil (DZ 41.800). 8. Artines delfos sp. nov., A–B: holotype male. Scale bar = 1 cm." httpUri="https://zenodo.org/record/3241041/files/figure.png" pageId="13" pageNumber="14">Fig. 8B</figureCitation>
): apical portion of discal cell with few yellow scales; central black spot rectangular, longer than wide.
</paragraph>
<paragraph id="4344E72D9D57CA40FF2CFD24FE55FBBE" blockId="13.[151,1437,318,1569]" pageId="13" pageNumber="14">
<emphasis id="718F3B3F9D57CA40FF2CFD24FED2FD56" box="[199,305,679,705]" italics="true" pageId="13" pageNumber="14">Genitalia</emphasis>
(
<figureCitation id="DBC0FBA89D57CA40FEA9FD24FE7BFD56" box="[322,408,679,705]" captionStart="FIGURES 30–32" captionStartId="24.[151,264,1772,1797]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure@24.[151,1436,181,1746]" captionTargetPageId="24" captionText="FIGURES 30–32. Male genitalia of the “aepitus group” species. 30. Artines delfos sp. nov., holotype. 31. Artines maya, holotype. 32. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.215). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241049/files/figure.png" pageId="13" pageNumber="14">Figs 30</figureCitation>
A–L, 44): tegumen more or less rectangular, with proximal margin concave and the distal margin not conspicuous (
<figureCitation id="DBC0FBA89D57CA40FE5DFD48FDFDFD72" box="[438,542,715,741]" captionStart="FIGURES 30–32" captionStartId="24.[151,264,1772,1797]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure@24.[151,1436,181,1746]" captionTargetPageId="24" captionText="FIGURES 30–32. Male genitalia of the “aepitus group” species. 30. Artines delfos sp. nov., holotype. 31. Artines maya, holotype. 32. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.215). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241049/files/figure.png" pageId="13" pageNumber="14">Fig. 30C</figureCitation>
), in lateral view shorter than length of gnathos; ventral arm of tegumen longer than dorsal arm of saccus (
<figureCitation id="DBC0FBA89D57CA40FE20FD6CFDD6FC9E" box="[459,565,751,777]" captionStart="FIGURES 30–32" captionStartId="24.[151,264,1772,1797]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure@24.[151,1436,181,1746]" captionTargetPageId="24" captionText="FIGURES 30–32. Male genitalia of the “aepitus group” species. 30. Artines delfos sp. nov., holotype. 31. Artines maya, holotype. 32. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.215). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241049/files/figure.png" pageId="13" pageNumber="14">Fig. 30A</figureCitation>
). Saccus shorter than tegumen+uncus, base same size as lateral margin (
<figureCitation id="DBC0FBA89D57CA40FA84FD6CFF29FCBA" captionStart="FIGURES 30–32" captionStartId="24.[151,264,1772,1797]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure@24.[151,1436,181,1746]" captionTargetPageId="24" captionText="FIGURES 30–32. Male genitalia of the “aepitus group” species. 30. Artines delfos sp. nov., holotype. 31. Artines maya, holotype. 32. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.215). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241049/files/figure.png" pageId="13" pageNumber="14">Fig. 30B</figureCitation>
). Uncus, in dorsal view, rectangular, about twice longer than wide, longer than tegumen, without disto-lateral projections, distal margin rounded (
<figureCitation id="DBC0FBA89D57CA40FDC5FCB4FD74FCC6" box="[558,663,823,849]" captionStart="FIGURES 30–32" captionStartId="24.[151,264,1772,1797]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure@24.[151,1436,181,1746]" captionTargetPageId="24" captionText="FIGURES 30–32. Male genitalia of the “aepitus group” species. 30. Artines delfos sp. nov., holotype. 31. Artines maya, holotype. 32. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.215). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241049/files/figure.png" pageId="13" pageNumber="14">Fig. 30C</figureCitation>
). Gnathos as in the other species of the group (
<figureCitation id="DBC0FBA89D57CA40FB5DFCB4FAA7FCC6" box="[1206,1348,823,849]" captionStart="FIGURES 30–32" captionStartId="24.[151,264,1772,1797]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure@24.[151,1436,181,1746]" captionTargetPageId="24" captionText="FIGURES 30–32. Male genitalia of the “aepitus group” species. 30. Artines delfos sp. nov., holotype. 31. Artines maya, holotype. 32. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.215). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241049/files/figure.png" pageId="13" pageNumber="14">Fig. 30A, D</figureCitation>
). Valva three times longer than wide; sacculus triangular and more than 1/2 width of valva; costa narrow and contiguous with ampulla; ampulla irregular and shorter than costa; ventral process of harpe wide, as long as tegumen, distal 1/3 angular, with apical setae thick and long; dorsal process of harpe on proximal 1/2, narrower than ventral process, with apex over dorsal margin of ventral process (
<figureCitation id="DBC0FBA89D57CA40FD58FC44FCE0FC76" box="[691,771,967,993]" captionStart="FIGURES 30–32" captionStartId="24.[151,264,1772,1797]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure@24.[151,1436,181,1746]" captionTargetPageId="24" captionText="FIGURES 30–32. Male genitalia of the “aepitus group” species. 30. Artines delfos sp. nov., holotype. 31. Artines maya, holotype. 32. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.215). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241049/files/figure.png" pageId="13" pageNumber="14">Fig. 30</figureCitation>
E–F). Aedeagus shorter than valva, slightly curved dorsally; ventral projection as in
<taxonomicName id="84FB9CAE9D57CA40FE42FC68FDFFFB92" baseAuthorityName="Geyer" baseAuthorityYear="1832" box="[425,540,1003,1029]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D57CA40FE42FC68FDFFFB92" box="[425,540,1003,1029]" italics="true" pageId="13" pageNumber="14">A. aepitus</emphasis>
</taxonomicName>
(
<figureCitation id="DBC0FBA89D57CA40FDC6FC68FD62FB92" box="[557,641,1003,1029]" captionStart="FIGURES 30–32" captionStartId="24.[151,264,1772,1797]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure@24.[151,1436,181,1746]" captionTargetPageId="24" captionText="FIGURES 30–32. Male genitalia of the “aepitus group” species. 30. Artines delfos sp. nov., holotype. 31. Artines maya, holotype. 32. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.215). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241049/files/figure.png" pageId="13" pageNumber="14">Fig. 30</figureCitation>
G–J); posterior end of cornutus triangular, with distal spines thin and curved laterally (
<figureCitation id="DBC0FBA89D57CA40FEBEFB8CFE49FBBE" box="[341,426,1039,1065]" captionStart="FIGURES 42–48" captionStartId="33.[151,264,1892,1917]" captionTargetBox="[151,1431,625,1874]" captionTargetId="figure@33.[151,1436,625,1880]" captionText="FIGURES 42–48. Dorsal view of vesica and cornutus of the &quot;aepitus group&quot; species. 42. Artines aepitus. 43. Artines panama sp. nov.. 44. Artines delfos sp. nov.. 45. Artines litoralis sp. nov.. 46. Artines rica. 47. Artines liege sp. nov. 48. Artines bamba sp. nov. Scale bar = 0,2 mm." httpUri="https://zenodo.org/record/3406274/files/figure.png" pageId="13" pageNumber="14">Fig. 44</figureCitation>
).
</paragraph>
<paragraph id="4344E72D9D57CA40FF2CFBB0FE7DFBDA" blockId="13.[151,1437,318,1569]" box="[199,414,1075,1101]" pageId="13" pageNumber="14">Female. Unknown.</paragraph>
<paragraph id="4344E72D9D57CA40FF2CFBD5FE05FBE6" blockId="13.[151,1437,318,1569]" box="[199,486,1110,1137]" pageId="13" pageNumber="14">
<emphasis id="718F3B3F9D57CA40FF2CFBD5FED9FBE6" bold="true" box="[199,314,1110,1137]" pageId="13" pageNumber="14">Variation</emphasis>
. Not observed.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D57CA40FF2CFBF9FF0AFA96" pageId="13" pageNumber="14" type="discussion">
<paragraph id="4344E72D9D57CA40FF2CFBF9FB40FB2E" blockId="13.[151,1437,318,1569]" pageId="13" pageNumber="14">
<emphasis id="718F3B3F9D57CA40FF2CFBF9FEABFB02" bold="true" box="[199,328,1146,1173]" pageId="13" pageNumber="14">Comments</emphasis>
.
<taxonomicName id="84FB9CAE9D57CA40FEB3FBF8FE17FB02" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[344,500,1147,1173]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="delfos" status="sp. nov.">
<emphasis id="718F3B3F9D57CA40FEB3FBF8FE17FB02" box="[344,500,1147,1173]" italics="true" pageId="13" pageNumber="14">Artines delfos</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D57CA40FE17FBF8FDBBFB02" bold="true" box="[508,600,1147,1173]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="6ABC86449D57CA40FE17FBF8FDBBFB02" box="[508,600,1147,1173]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is apparently rare or rarely collected. The male genitalia are similar to
<taxonomicName id="84FB9CAE9D57CA40FA6FFBF8FF27FB2E" authorityName=", S. R. Steinhauser &amp; G. T. Austin" authorityYear="1993" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="rica">
<emphasis id="718F3B3F9D57CA40FA6FFBF8FF27FB2E" italics="true" pageId="13" pageNumber="14">A. rica</emphasis>
</taxonomicName>
and
<taxonomicName id="84FB9CAE9D57CA40FF10FB1CFEB3FB2E" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[251,336,1183,1209]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="liege" status="sp. nov.">
<emphasis id="718F3B3F9D57CA40FF10FB1CFEB3FB2E" box="[251,336,1183,1209]" italics="true" pageId="13" pageNumber="14">A. liege</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D57CA40FEBCFB1CFE51FB2E" bold="true" box="[343,434,1183,1209]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="6ABC86449D57CA40FEBCFB1CFE51FB2E" box="[343,434,1183,1209]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, differing in the ventral process of harpe wider and not cylindrical.
</paragraph>
<paragraph id="4344E72D9D57CA40FF2CFB41FF0AFA96" blockId="13.[151,1437,318,1569]" pageId="13" pageNumber="14">
<emphasis id="718F3B3F9D57CA40FF2CFB41FE09FB4A" bold="true" box="[199,490,1218,1245]" pageId="13" pageNumber="14">Geographic distribution</emphasis>
(
<figureCitation id="DBC0FBA89D57CA40FE11FB40FDACFB4A" box="[506,591,1219,1245]" captionStart="FIGURE 78" captionStartId="42.[151,250,910,935]" captionTargetBox="[151,1436,174,898]" captionTargetId="figure@42.[151,1436,174,898]" captionTargetPageId="42" captionText="FIGURE 78. Geographic distribution of the “aepitus group” species. The symbols with white inferior half represent records from literature." httpUri="https://zenodo.org/record/3241067/files/figure.png" pageId="13" pageNumber="14">Fig. 78</figureCitation>
). The species is known only from the
<typeStatus id="9C40598F9D57CA40FBEFFB40FBD7FB4A" box="[1028,1076,1219,1245]" pageId="13" pageNumber="14">type</typeStatus>
locality. BRAZIL—
<emphasis id="718F3B3F9D57CA40FACAFB40FA77FB4A" box="[1313,1428,1219,1245]" italics="true" pageId="13" pageNumber="14">
<collectingRegion id="813F29CF9D57CA40FACAFB40FA77FB4A" box="[1313,1428,1219,1245]" country="Venezuela" name="Amazonas" pageId="13" pageNumber="14">Amazonas</collectingRegion>
</emphasis>
: Maués.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D57CA40FF2CFA88FCA9FAB2" box="[199,842,1290,1317]" pageId="13" pageNumber="14" type="description">
<paragraph id="4344E72D9D57CA40FF2CFA88FCA9FAB2" blockId="13.[151,1437,318,1569]" box="[199,842,1290,1317]" pageId="13" pageNumber="14">
<emphasis id="718F3B3F9D57CA40FF2CFA88FE31FAB2" bold="true" box="[199,466,1290,1317]" pageId="13" pageNumber="14">Temporal distribution</emphasis>
. This species occurs in February.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D57CA40FF2CFAACFE2AFADE" box="[199,457,1327,1353]" pageId="13" pageNumber="14" type="biology_ecology">
<paragraph id="4344E72D9D57CA40FF2CFAACFE2AFADE" blockId="13.[151,1437,318,1569]" box="[199,457,1327,1353]" pageId="13" pageNumber="14">
<emphasis id="718F3B3F9D57CA40FF2CFAACFEA7FADE" bold="true" box="[199,324,1327,1353]" pageId="13" pageNumber="14">Host plant</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D57CA40FF2CFAD0FE03FA22" pageId="13" pageNumber="14" type="etymology">
<paragraph id="4344E72D9D57CA40FF2CFAD0FE03FA22" blockId="13.[151,1437,318,1569]" pageId="13" pageNumber="14">
<emphasis id="718F3B3F9D57CA40FF2CFAD0FEABFAFA" bold="true" box="[199,328,1363,1389]" pageId="13" pageNumber="14">Etymology</emphasis>
. Delfos is a Greek city that the ancient Greeks believed to be the center of the universe. The name of the species is in reference to its central geographic distribution in relation to the other species of
<taxonomicName id="84FB9CAE9D57CA40FAFBFAF4FA82FA06" authorityName="Godman" authorityYear="1901" box="[1296,1377,1399,1425]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="718F3B3F9D57CA40FAFBFAF4FA82FA06" box="[1296,1377,1399,1425]" italics="true" pageId="13" pageNumber="14">Artines</emphasis>
</taxonomicName>
. The name is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D57CA40FF2CFA3CFCA9F9B6" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="4344E72D9D57CA40FF2CFA3CFCA9F9B6" blockId="13.[151,1437,318,1569]" pageId="13" pageNumber="14">
<emphasis id="718F3B3F9D57CA40FF2CFA3CFE91FA4E" bold="true" box="[199,370,1470,1497]" pageId="13" pageNumber="14">Type material</emphasis>
.
<typeStatus id="9C40598F9D57CA40FE68FA3CFE08FA4E" box="[387,491,1471,1497]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
male deposited at the OM–DZUP with the following labels: /
<materialsCitation id="F393ED709D57CA40FB54FA3CFA68FA6A" ID-GBIF-Occurrence="2248478850" collectingDate="2008-03-01" collectingDateMax="2008-03-15" collectingDateMin="2008-03-01" collectionCode="OM" collectorName="A. Medeiros" country="Brazil" location="Rio Preto" pageId="13" pageNumber="14" specimenCode="OM 70.129" specimenCount="1" typeStatus="holotype">
<typeStatus id="9C40598F9D57CA40FB54FA3CFA86FA4E" box="[1215,1381,1471,1497]" pageId="13" pageNumber="14" type="holotype">HOLOTYPUS</typeStatus>
/
<date id="3745C1ED9D57CA40FA6BFA3CFEFBFA6A" pageId="13" pageNumber="14" value="2008-03-01" valueMax="2008-03-15" valueMin="2008-03-01">
<collectingDate id="270138059D57CA40FA6BFA3CFEFBFA6A" pageId="13" pageNumber="14" value="2008-03-01" valueMax="2008-03-15" valueMin="2008-03-01">1– 15.III.2008</collectingDate>
</date>
,
<location id="4624B1F69D57CA40FECDFA60FE75FA6A" LSID="urn:lsid:plazi:treatment:CB52563B9D57CA40FF7CFF1AFCA9F9B6:4624B1F69D57CA40FECDFA60FE75FA6A" box="[294,406,1507,1533]" country="Brazil" name="Rio Preto" pageId="13" pageNumber="14">Rio Preto</location>
, Maués, Amazonas, [
<collectingCountry id="3BECA7BD9D57CA40FD67FA61FD3BFA6A" box="[652,728,1506,1533]" name="Brazil" pageId="13" pageNumber="14">Brazil</collectingCountry>
], [no collector] / gen. prep.
<collectorName id="EE0E82FB9D57CA40FBF7FA60FB48FA6A" box="[1052,1195,1507,1533]" pageId="13" pageNumber="14">A. Medeiros</collectorName>
2018 /
<specimenCode id="135D4F569D57CA40FAECFA60FA68FA6A" box="[1287,1419,1507,1533]" collectionCode="OM" country="New Zealand" name="Otago Museum" pageId="13" pageNumber="14">OM 70.129</specimenCode>
</materialsCitation>
/
<typeStatus id="9C40598F9D57CA40FF7CF984FEEFF9B6" box="[151,268,1543,1569]" pageId="13" pageNumber="14" type="holotype">Holotypus</typeStatus>
<taxonomicName id="84FB9CAE9D57CA40FEF8F984FD26F9B6" authority="Medeiros &amp; Casagrande" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[275,709,1543,1569]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="delfos">
<emphasis id="718F3B3F9D57CA40FEF8F984FE4DF9B6" box="[275,430,1543,1569]" italics="true" pageId="13" pageNumber="14">Artines delfos</emphasis>
Medeiros &amp; Casagrande
</taxonomicName>
det. 2018 /.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="0C5E9F4E2C7BE6C9DA5ED561B5B807E3">Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species</mods:title>
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<subSubSection id="0BE1B4A69D57CA40FF7CF9F0FD4EF91A" box="[151,685,1651,1677]" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="4344E72D9D57CA40FF7CF9F0FD4EF91A" blockId="13.[151,685,1651,1713]" box="[151,685,1651,1677]" pageId="13" pageNumber="14">
<heading id="180C50419D57CA40FF7CF9F0FD4EF91A" bold="true" box="[151,685,1651,1677]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<taxonomicName id="84FB9CAE9D57CA40FF7CF9F0FD4EF91A" authority="A. Warren &amp; Dolibaina, 2015" authorityName="A. Warren &amp; Dolibaina" authorityYear="2015" box="[151,685,1651,1677]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="maya">
<emphasis id="718F3B3F9D57CA40FF7CF9F0FD4EF91A" bold="true" box="[151,685,1651,1677]" pageId="13" pageNumber="14">
<emphasis id="718F3B3F9D57CA40FF7CF9F0FEDBF91A" bold="true" box="[151,312,1651,1677]" italics="true" pageId="13" pageNumber="14">Artines maya</emphasis>
A. Warren &amp;
<bibRefCitation id="276A9ADC9D57CA40FE02F9F0FD4EF91A" author="Dolibaina, D. R. &amp; Warren, A. D. &amp; Carneiro, E. &amp; Mielke, O. H. H." box="[489,685,1651,1677]" pageId="13" pageNumber="14" pagination="52 - 59" refId="ref39956" refString="Dolibaina, D. R., Warren, A. D., Carneiro, E. &amp; Mielke, O. H. H. (2015) Three new species of Artines Godman, 1901 from Central and South America (Hesperiidae: Hesperiinae: Moncini). Tropical Lepidoptera Research, 25 (2), 52 - 59." type="journal article" year="2015">Dolibaina, 2015</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D57CA40FF7CF914FD27F8F9" pageId="13" pageNumber="14" type="description">
<paragraph id="4344E72D9D57CA40FF7CF914FED3F926" blockId="13.[151,685,1651,1713]" box="[151,304,1687,1713]" pageId="13" pageNumber="14">
<figureCitation id="DBC0FBA89D57CA40FF7CF914FF3DF926" box="[151,222,1687,1713]" captionStart="FIGURES 9–13" captionStartId="7.[151,264,1180,1205]" captionTargetBox="[152,1436,180,1162]" captionTargetId="figure@7.[151,1436,174,1169]" captionTargetPageId="7" captionText="FIGURES 9–13. Species of the “aepitus group” in dorsal and ventral views. 9. Artines maya, A–B: holotype male. 10. Artines litoralis sp. nov., A–B: holotype male, C–D: allotype female. 11. Artines rica, A–B: male, Turrialba, Cartago, Costa Rica (DZ 41.890), C–D: female from same locality of male (DZ 41.840). 12. Artines liege sp. nov., A–B: holotype male, C–D: allotype female. 13. Artines bamba sp. nov., A–B: holotype male, C–D: tentative female, Moyobamba, San Martin, Peru (OM 79.413). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241043/files/figure.png" pageId="13" pageNumber="14">Figs 9</figureCitation>
,
<figureCitation id="DBC0FBA89D57CA40FF01F914FEEBF926" box="[234,264,1687,1713]" captionStart="FIGURES 30–32" captionStartId="24.[151,264,1772,1797]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure@24.[151,1436,181,1746]" captionTargetPageId="24" captionText="FIGURES 30–32. Male genitalia of the “aepitus group” species. 30. Artines delfos sp. nov., holotype. 31. Artines maya, holotype. 32. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.215). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241049/files/figure.png" pageId="13" pageNumber="14">31</figureCitation>
,
<figureCitation id="DBC0FBA89D57CA40FEFFF914FED3F926" box="[276,304,1687,1713]" captionStart="FIGURE 78" captionStartId="42.[151,250,910,935]" captionTargetBox="[151,1436,174,898]" captionTargetId="figure@42.[151,1436,174,898]" captionTargetPageId="42" captionText="FIGURE 78. Geographic distribution of the “aepitus group” species. The symbols with white inferior half represent records from literature." httpUri="https://zenodo.org/record/3241067/files/figure.png" pageId="13" pageNumber="14">78</figureCitation>
</paragraph>
<paragraph id="4344E72D9D57CA40FF7CF958FD27F8F9" blockId="13.[151,1437,1754,1963]" pageId="13" pageNumber="14">
<taxonomicName id="84FB9CAE9D57CA40FF7CF958FECCF964" baseAuthorityName="Geyer" baseAuthorityYear="1832" box="[151,303,1755,1779]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D57CA40FF7CF958FECCF964" box="[151,303,1755,1779]" italics="true" pageId="13" pageNumber="14">Artines aepitus</emphasis>
</taxonomicName>
(misid.); J. de la Maza E. &amp; R. G. de la Maza E., 1985.
<emphasis id="718F3B3F9D57CA40FC8EF958FBAFF964" bold="true" box="[869,1100,1754,1779]" pageId="13" pageNumber="14">Rev. Soc. Mex. Lep. 9</emphasis>
: 43.—Llorente-Bousquets; Luis- Martínez &amp; Vargas-Fernández, 1990.
<emphasis id="718F3B3F9D57CA40FDAEF979FCC1F885" bold="true" box="[581,802,1786,1810]" pageId="13" pageNumber="14">Publ. esp. Mus. Zool</emphasis>
.,
<collectingRegion id="813F29CF9D57CA40FCDDF97AFC65F885" box="[822,902,1785,1810]" country="Mexico" name="Mexico" pageId="13" pageNumber="14">México</collectingRegion>
,
<emphasis id="718F3B3F9D57CA40FC7BF97AFC7EF885" bold="true" box="[912,925,1785,1810]" pageId="13" pageNumber="14">1</emphasis>
: 35.—J. de la Maza E.; White &amp; R. G. de la Maza E., 1991.
<emphasis id="718F3B3F9D57CA40FEC3F89BFDFAF8A6" bold="true" box="[296,537,1816,1841]" pageId="13" pageNumber="14">Rev. Soc. Mex. Lep. 14</emphasis>
: 29.—R. G. de la Maza E. &amp; J. de la Maza E., 1993.
<emphasis id="718F3B3F9D57CA40FBC6F89BFB34F8A6" bold="true" box="[1069,1239,1816,1841]" pageId="13" pageNumber="14">
Marip.
<collectingRegion id="813F29CF9D57CA40FB95F89BFB34F8A6" box="[1150,1239,1816,1841]" country="Mexico" name="Chiapas" pageId="13" pageNumber="14">Chiapas</collectingRegion>
</emphasis>
, p. 206.—
<bibRefCitation id="276A9ADC9D57CA40FAD4F89BFEC0F8D8" author="Austin, G. T. &amp; Mendez, C. &amp; Launer, A. E." pageId="13" pageNumber="14" pagination="8 - 18" refId="ref38142" refString="Austin, G. T., Mendez, C. &amp; Launer, A. E. (1999) A preliminary checklist of Guatemalan butterflies: Hesperiidae (Lepidoptera: Hesperioidea). Tropical Lepidoptera, 9 (Supplement 2), 8 - 18." type="journal article" year="1999">
Austin
<emphasis id="718F3B3F9D57CA40FA61F89AFF02F8D8" bold="true" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
, 1999
</bibRefCitation>
.
<emphasis id="718F3B3F9D57CA40FEC7F8B4FE55F8D8" bold="true" box="[300,438,1846,1871]" pageId="13" pageNumber="14">Trop. Lep. 9,</emphasis>
Suppl. 2: 13.—J. de la Maza &amp; R. G. de la Maza, 2015,
<emphasis id="718F3B3F9D57CA40FC02F8B4FC1DF8D8" bold="true" box="[1001,1022,1847,1871]" italics="true" pageId="13" pageNumber="14">in</emphasis>
Carabias; J. de la Maza &amp; Cadena (eds.).
<emphasis id="718F3B3F9D57CA40FF2CF8D6FD92F8F9" bold="true" box="[199,625,1877,1902]" pageId="13" pageNumber="14">Conserv. desarr. sust. Selva Lacandona</emphasis>
, p. 648.
</paragraph>
</subSubSection>
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<paragraph id="4344E72D9D57CA40FF7CF8F6FB92F83C" blockId="13.[151,1437,1754,1963]" pageId="13" pageNumber="14">
<taxonomicName id="84FB9CAE9D57CA40FF7CF8F6FDBFF81A" authority="A. Warren &amp; Dolibaina, 2015" authorityName="A. Warren &amp; Dolibaina" authorityYear="2015" box="[151,604,1908,1933]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="maya">
<emphasis id="718F3B3F9D57CA40FF7CF8F6FEFCF81B" box="[151,287,1909,1932]" italics="true" pageId="13" pageNumber="14">Artines maya</emphasis>
A. Warren &amp;
<bibRefCitation id="276A9ADC9D57CA40FE5FF8F7FDBFF81A" author="Dolibaina, D. R. &amp; Warren, A. D. &amp; Carneiro, E. &amp; Mielke, O. H. H." box="[436,604,1908,1933]" pageId="13" pageNumber="14" pagination="52 - 59" refId="ref39956" refString="Dolibaina, D. R., Warren, A. D., Carneiro, E. &amp; Mielke, O. H. H. (2015) Three new species of Artines Godman, 1901 from Central and South America (Hesperiidae: Hesperiinae: Moncini). Tropical Lepidoptera Research, 25 (2), 52 - 59." type="journal article" year="2015">Dolibaina, 2015</bibRefCitation>
</taxonomicName>
.
<emphasis id="718F3B3F9D57CA40FD8CF8F7FCD1F81A" bold="true" box="[615,818,1908,1933]" pageId="13" pageNumber="14">Trop. Lep. Res. 25</emphasis>
(2): 52, figs 1-2 (male d, v), 21 (male gen.);
<typeStatus id="9C40598F9D57CA40FAEFF8F7FABEF81A" box="[1284,1373,1908,1933]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
male,
<date id="3745C1ED9D57CA40FF2CF810FECCF83C" box="[199,303,1939,1963]" pageId="13" pageNumber="14" value="1997-03-04">4.III.1997</date>
, circa 5 Km W. Hill Bank, Orangewalk district,
<collectingCountry id="3BECA7BD9D57CA40FCFCF811FCB9F83C" box="[791,858,1938,1963]" name="Belize" pageId="13" pageNumber="14">Belize</collectingCountry>
, Valerie Giles
<emphasis id="718F3B3F9D57CA40FC05F810FBEEF83C" box="[1006,1037,1939,1963]" italics="true" pageId="13" pageNumber="14">leg</emphasis>
.; MGCL.
</paragraph>
<paragraph id="4344E72D9D57CA43FF7CF85AFD2AFF41" blockId="13.[151,1436,2009,2036]" lastBlockId="14.[151,1437,152,1079]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<emphasis id="718F3B3F9D57CA40FF7CF85AFE96F863" bold="true" box="[151,373,2009,2036]" pageId="13" pageNumber="14">Taxonomic history</emphasis>
.
<taxonomicName id="84FB9CAE9D57CA40FE68F859FDF4F864" authorityName="A. Warren &amp; Dolibaina" authorityYear="2015" box="[387,535,2010,2036]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="maya">
<emphasis id="718F3B3F9D57CA40FE68F859FDF4F864" box="[387,535,2010,2036]" italics="true" pageId="13" pageNumber="14">Artines maya</emphasis>
</taxonomicName>
was originally treated as
<taxonomicName id="84FB9CAE9D57CA40FCDFF859FC40F864" ID-CoL="GZYC" baseAuthorityName="Geyer" baseAuthorityYear="1832" box="[820,931,2010,2036]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D57CA40FCDFF859FC40F864" box="[820,931,2010,2036]" italics="true" pageId="13" pageNumber="14">A. aepitus</emphasis>
</taxonomicName>
in studies conducted in Central America (see catalog). The description of the species was based on three males, one each from Orangewalk,
<collectingCountry id="3BECA7BD9D54CA43FB3EFF1BFAC3FF25" box="[1237,1312,152,178]" name="Belize" pageId="14" pageNumber="15">Belize</collectingCountry>
; Lacantún,
<collectingRegion id="813F29CF9D54CA43FF7CFF3FFF12FF41" box="[151,241,188,214]" country="Mexico" name="Chiapas" pageId="14" pageNumber="15">Chiapas</collectingRegion>
,
<collectingCountry id="3BECA7BD9D54CA43FF14FF3FFEB6FF41" box="[255,341,188,214]" name="Mexico" pageId="14" pageNumber="15">Mexico</collectingCountry>
and Morales,
<collectingRegion id="813F29CF9D54CA43FE1EFF3FFDDFFF41" box="[501,572,188,214]" country="Guatemala" name="Izabal" pageId="14" pageNumber="15">Izabal</collectingRegion>
,
<collectingCountry id="3BECA7BD9D54CA43FDA3FF3FFD26FF41" box="[584,709,188,214]" name="Guatemala" pageId="14" pageNumber="15">Guatemala</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D54CA43FF2CFF63FE7DFE61" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="4344E72D9D54CA43FF2CFF63FC9DFE44" blockId="14.[151,1437,152,1079]" pageId="14" pageNumber="15">
<emphasis id="718F3B3F9D54CA43FF2CFF63FED9FF6C" bold="true" box="[199,314,224,251]" pageId="14" pageNumber="15">Diagnosis</emphasis>
.
<taxonomicName id="84FB9CAE9D54CA43FEA1FF62FE02FF6D" authorityName="A. Warren &amp; Dolibaina" authorityYear="2015" box="[330,481,225,251]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="maya">
<emphasis id="718F3B3F9D54CA43FEA1FF62FE02FF6D" box="[330,481,225,251]" italics="true" pageId="14" pageNumber="15">Artines maya</emphasis>
</taxonomicName>
is very similar to the other species of the “
<taxonomicName id="84FB9CAE9D54CA43FC3AFF62FBC1FF6D" baseAuthorityName="Geyer" baseAuthorityYear="1832" box="[977,1058,225,250]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D54CA43FC3AFF62FBC1FF6D" box="[977,1058,225,250]" italics="true" pageId="14" pageNumber="15">aepitus</emphasis>
</taxonomicName>
group”, but its distribution is restricted to Central America where it is potentially sympatric with
<taxonomicName id="84FB9CAE9D54CA43FC90FE86FC2BFE89" authorityName=", S. R. Steinhauser &amp; G. T. Austin" authorityYear="1993" box="[891,968,261,287]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="rica">
<emphasis id="718F3B3F9D54CA43FC90FE86FC2BFE89" box="[891,968,261,287]" italics="true" pageId="14" pageNumber="15">A. rica</emphasis>
</taxonomicName>
and
<taxonomicName id="84FB9CAE9D54CA43FBEBFE86FB9AFE89" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[1024,1145,261,287]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="panama" status="sp. nov.">
<emphasis id="718F3B3F9D54CA43FBEBFE86FB9AFE89" box="[1024,1145,261,287]" italics="true" pageId="14" pageNumber="15">A. panama</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D54CA43FB6AFE86FB38FE88" bold="true" box="[1153,1243,261,287]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="6ABC86449D54CA43FB6AFE86FB38FE88" box="[1153,1243,261,287]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and from which can be easily distinguished by being relatively smaller; discal area of ventral hind wing with the central black spot wider than long; harpe with ventral process triangular and shorter than tegumen and aedeagus longer than valva. These characters are only shared with
<taxonomicName id="84FB9CAE9D54CA43FDA9FEF2FD59FE1D" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[578,698,368,395]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="litoralis" status="sp. nov.">
<emphasis id="718F3B3F9D54CA43FDA9FEF2FD59FE1D" box="[578,698,368,395]" italics="true" pageId="14" pageNumber="15">A. litoralis</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D54CA43FD2AFEF2FCF9FE1C" bold="true" box="[705,794,369,395]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="6ABC86449D54CA43FD2AFEF2FCF9FE1C" box="[705,794,369,395]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, from which it differs by the tegumen shorter than uncus; ampulla distally curved ventrally and ventral process of harpe shorter than dorsal. Besides that,
<taxonomicName id="84FB9CAE9D54CA43FB2FFE16FADFFE39" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[1220,1340,404,431]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="litoralis" status="sp. nov.">
<emphasis id="718F3B3F9D54CA43FB2FFE16FADFFE39" box="[1220,1340,404,431]" italics="true" pageId="14" pageNumber="15">A. litoralis</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D54CA43FAA8FE16FA7FFE38" bold="true" box="[1347,1436,405,431]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="6ABC86449D54CA43FAA8FE16FA7FFE38" box="[1347,1436,405,431]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
occurs only on the Brazilian coast, from
<collectingRegion id="813F29CF9D54CA43FDB4FE3BFD50FE45" box="[607,691,440,466]" country="Brazil" name="Paraiba" pageId="14" pageNumber="15">Paraíba</collectingRegion>
to
<collectingRegion id="813F29CF9D54CA43FD3CFE3BFC9AFE44" box="[727,889,440,467]" country="Brazil" name="Espirito Santo" pageId="14" pageNumber="15">Espírito Santo</collectingRegion>
.
</paragraph>
<paragraph id="4344E72D9D54CA43FF2CFE5FFE7DFE61" blockId="14.[151,1437,152,1079]" box="[199,414,476,502]" pageId="14" pageNumber="15">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D54CA43FF2CFD83FE00FD8D" box="[199,483,512,539]" pageId="14" pageNumber="15" type="description">
<paragraph id="4344E72D9D54CA43FF2CFD83FE00FD8D" blockId="14.[151,1437,152,1079]" box="[199,483,512,539]" pageId="14" pageNumber="15">
<emphasis id="718F3B3F9D54CA43FF2CFD83FEDBFD8C" bold="true" box="[199,312,512,539]" pageId="14" pageNumber="15">Variation</emphasis>
. Not observed.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D54CA43FF2CFDA7FAEDFD58" pageId="14" pageNumber="15" type="discussion">
<paragraph id="4344E72D9D54CA43FF2CFDA7FBD8FD11" blockId="14.[151,1437,152,1079]" pageId="14" pageNumber="15">
<emphasis id="718F3B3F9D54CA43FF2CFDA7FEABFDA8" bold="true" box="[199,328,548,575]" pageId="14" pageNumber="15">Comments</emphasis>
.
<taxonomicName id="84FB9CAE9D54CA43FEBCFDA6FE0DFDA9" authorityName="A. Warren &amp; Dolibaina" authorityYear="2015" box="[343,494,549,575]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="maya">
<emphasis id="718F3B3F9D54CA43FEBCFDA6FE0DFDA9" box="[343,494,549,575]" italics="true" pageId="14" pageNumber="15">Artines maya</emphasis>
</taxonomicName>
is one of the smallest species of the genus. Although it shares great morphological similarity with
<taxonomicName id="84FB9CAE9D54CA43FEAEFDCAFE5CFDF5" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[325,447,584,611]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="litoralis" status="sp. nov.">
<emphasis id="718F3B3F9D54CA43FEAEFDCAFE5CFDF5" box="[325,447,584,611]" italics="true" pageId="14" pageNumber="15">A. litoralis</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D54CA43FE2CFDCAFDC1FDF4" bold="true" box="[455,546,585,611]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="6ABC86449D54CA43FE2CFDCAFDC1FDF4" box="[455,546,585,611]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, these two species are widely separated geographically and after relatively intense sampling between their areas of occurrence, no intermediate form was found.
</paragraph>
<paragraph id="4344E72D9D54CA43FF2CFD13FAEDFD58" blockId="14.[151,1437,152,1079]" pageId="14" pageNumber="15">
<emphasis id="718F3B3F9D54CA43FF2CFD13FE04FD3C" bold="true" box="[199,487,656,683]" pageId="14" pageNumber="15">Geographic distribution</emphasis>
(
<figureCitation id="DBC0FBA89D54CA43FE1EFD13FDABFD3C" box="[501,584,656,683]" captionStart="FIGURE 78" captionStartId="42.[151,250,910,935]" captionTargetBox="[151,1436,174,898]" captionTargetId="figure@42.[151,1436,174,898]" captionTargetPageId="42" captionText="FIGURE 78. Geographic distribution of the “aepitus group” species. The symbols with white inferior half represent records from literature." httpUri="https://zenodo.org/record/3241067/files/figure.png" pageId="14" pageNumber="15">Fig. 78</figureCitation>
). The species is known from lowland forest of Central America (
<bibRefCitation id="276A9ADC9D54CA43FAF9FD13FF1BFD58" author="Dolibaina, D. R. &amp; Warren, A. D. &amp; Carneiro, E. &amp; Mielke, O. H. H." pageId="14" pageNumber="15" pagination="52 - 59" refId="ref39956" refString="Dolibaina, D. R., Warren, A. D., Carneiro, E. &amp; Mielke, O. H. H. (2015) Three new species of Artines Godman, 1901 from Central and South America (Hesperiidae: Hesperiinae: Moncini). Tropical Lepidoptera Research, 25 (2), 52 - 59." type="journal article" year="2015">
Dolibaina
<emphasis id="718F3B3F9D54CA43FA63FD12FF57FD59" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
2015
</bibRefCitation>
). MEXICO—
<emphasis id="718F3B3F9D54CA43FE7EFD37FE11FD59" box="[405,498,692,718]" italics="true" pageId="14" pageNumber="15">
<collectingRegion id="813F29CF9D54CA43FE7EFD37FE11FD59" box="[405,498,692,718]" country="Mexico" name="Chiapas" pageId="14" pageNumber="15">Chiapas</collectingRegion>
</emphasis>
: Lacantún. GUATEMALA—
<emphasis id="718F3B3F9D54CA43FCAAFD37FC64FD59" box="[833,903,692,718]" italics="true" pageId="14" pageNumber="15">
<collectingRegion id="813F29CF9D54CA43FCAAFD37FC64FD59" box="[833,903,692,718]" country="Guatemala" name="Izabal" pageId="14" pageNumber="15">Izabal</collectingRegion>
</emphasis>
: Morales. BELIZE—
<emphasis id="718F3B3F9D54CA43FB97FD37FAE5FD59" box="[1148,1286,692,718]" italics="true" pageId="14" pageNumber="15">Orangewalk</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D54CA43FF2CFD5AFAC2FD64" box="[199,1313,728,755]" pageId="14" pageNumber="15" type="description">
<paragraph id="4344E72D9D54CA43FF2CFD5AFAC2FD64" blockId="14.[151,1437,152,1079]" box="[199,1313,728,755]" pageId="14" pageNumber="15">
<emphasis id="718F3B3F9D54CA43FF2CFD5AFE33FD64" bold="true" box="[199,464,728,755]" pageId="14" pageNumber="15">Temporal distribution</emphasis>
. This species occurs in March, May and September (
<bibRefCitation id="276A9ADC9D54CA43FBC9FD5BFAF6FD64" author="Dolibaina, D. R. &amp; Warren, A. D. &amp; Carneiro, E. &amp; Mielke, O. H. H." box="[1058,1301,728,755]" pageId="14" pageNumber="15" pagination="52 - 59" refId="ref39956" refString="Dolibaina, D. R., Warren, A. D., Carneiro, E. &amp; Mielke, O. H. H. (2015) Three new species of Artines Godman, 1901 from Central and South America (Hesperiidae: Hesperiinae: Moncini). Tropical Lepidoptera Research, 25 (2), 52 - 59." type="journal article" year="2015">
Dolibaina
<emphasis id="718F3B3F9D54CA43FB72FD5AFB31FD65" box="[1177,1234,728,754]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
2015
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D54CA43FF2CFD7EFE2AFC81" box="[199,457,764,791]" pageId="14" pageNumber="15" type="biology_ecology">
<paragraph id="4344E72D9D54CA43FF2CFD7EFE2AFC81" blockId="14.[151,1437,152,1079]" box="[199,457,764,791]" pageId="14" pageNumber="15">
<emphasis id="718F3B3F9D54CA43FF2CFD7EFEA7FC80" bold="true" box="[199,324,765,791]" pageId="14" pageNumber="15">Host plant</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D54CA43FF2CFCA2FCC3FCC8" pageId="14" pageNumber="15" type="etymology">
<paragraph id="4344E72D9D54CA43FF2CFCA2FCC3FCC8" blockId="14.[151,1437,152,1079]" pageId="14" pageNumber="15">
<emphasis id="718F3B3F9D54CA43FF2CFCA2FEABFCAC" bold="true" box="[199,328,801,827]" pageId="14" pageNumber="15">Etymology</emphasis>
. According to
<bibRefCitation id="276A9ADC9D54CA43FE12FCA3FCE2FCAC" author="Dolibaina, D. R. &amp; Warren, A. D. &amp; Carneiro, E. &amp; Mielke, O. H. H." box="[505,769,800,827]" pageId="14" pageNumber="15" pagination="52 - 59" refId="ref39956" refString="Dolibaina, D. R., Warren, A. D., Carneiro, E. &amp; Mielke, O. H. H. (2015) Three new species of Artines Godman, 1901 from Central and South America (Hesperiidae: Hesperiinae: Moncini). Tropical Lepidoptera Research, 25 (2), 52 - 59." type="journal article" year="2015">
Dolibaina
<emphasis id="718F3B3F9D54CA43FD99FCA2FD4EFCAD" box="[626,685,800,826]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
(2015)
</bibRefCitation>
this species was named in reference to the Maya people, former inhabitants of the region where the species occurs.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D54CA43FF2CFCEAFDCEFBA0" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="4344E72D9D54CA43FF2CFCEAFC22FC5C" blockId="14.[151,1437,152,1079]" pageId="14" pageNumber="15">
<emphasis id="718F3B3F9D54CA43FF2CFCEAFE93FC14" bold="true" box="[199,368,872,899]" pageId="14" pageNumber="15">Type material</emphasis>
.
<materialsCitation id="F393ED709D54CA43FE6AFCEBFB8DFC15" ID-GBIF-Occurrence="2248478851" box="[385,1134,872,899]" collectionCode="MGCL" pageId="14" pageNumber="15" specimenCount="1" typeStatus="holotype">
<typeStatus id="9C40598F9D54CA43FE6AFCEBFE0AFC15" box="[385,489,872,898]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
male deposited at the
<collectionCode id="25EA7FE89D54CA43FD1AFCEAFCA2FC14" box="[753,833,873,899]" pageId="14" pageNumber="15">MGCL</collectionCode>
with the following labels
</materialsCitation>
: /
<materialsCitation id="F393ED709D54CA43FB67FCEAFEE8FC5C" ID-GBIF-Occurrence="2248478892" collectingDate="1997-03-04" collectorName="Km W. Hill Bank &amp; Valerie Giles &amp; Genitalia Vial &amp; Andrew D. Warren" country="Belize" location="Orangewalk" pageId="14" pageNumber="15" specimenCount="1" typeStatus="holotype">
<typeStatus id="9C40598F9D54CA43FB67FCEAFAD1FC14" box="[1164,1330,873,899]" pageId="14" pageNumber="15" type="holotype">HOLOTYPUS</typeStatus>
/
<collectingCountry id="3BECA7BD9D54CA43FAA6FCEBFA7AFC15" box="[1357,1433,872,898]" name="Belize" pageId="14" pageNumber="15">Belize</collectingCountry>
:
<location id="4624B1F69D54CA43FF7CFC0FFEC3FC31" LSID="urn:lsid:plazi:treatment:CB52563B9D57CA43FF7CF9F0FDCEFBA0:4624B1F69D54CA43FF7CFC0FFEC3FC31" box="[151,288,908,934]" country="Belize" name="Orangewalk" pageId="14" pageNumber="15">Orangewalk</location>
dist[rict]., c[irc]a. 5
<collectorName id="EE0E82FB9D54CA43FDECFC0EFD2EFC31" box="[519,717,908,935]" pageId="14" pageNumber="15">Km W. Hill Bank</collectorName>
,
<date id="3745C1ED9D54CA43FD33FC0EFCBFFC30" box="[728,860,909,935]" pageId="14" pageNumber="15" value="1997-03-04">
<collectingDate id="270138059D54CA43FD33FC0EFCBFFC30" box="[728,860,909,935]" pageId="14" pageNumber="15" value="1997-03-04">Mar 4-1997</collectingDate>
</date>
<collectorName id="EE0E82FB9D54CA43FC88FC0FFC17FC31" box="[867,1012,908,934]" pageId="14" pageNumber="15">Valerie Giles</collectorName>
[
<emphasis id="718F3B3F9D54CA43FBEFFC0FFBC5FC31" box="[1028,1062,908,934]" italics="true" pageId="14" pageNumber="15">leg</emphasis>
.] /
<collectorName id="EE0E82FB9D54CA43FBA7FC0FFB04FC31" box="[1100,1255,908,934]" pageId="14" pageNumber="15">Genitalia Vial</collectorName>
#98-72
<collectorName id="EE0E82FB9D54CA43FAA9FC0FFEE8FC5C" pageId="14" pageNumber="15">Andrew D. Warren</collectorName>
</materialsCitation>
/
<typeStatus id="9C40598F9D54CA43FECBFC33FE76FC5D" box="[288,405,944,970]" pageId="14" pageNumber="15" type="holotype">Holotypus</typeStatus>
<taxonomicName id="84FB9CAE9D54CA43FE77FC32FCD8FC5D" authority="A. Warren &amp; Dolibaina" authorityName="A. Warren &amp; Dolibaina" authorityYear="2015" box="[412,827,944,971]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="maya">
<emphasis id="718F3B3F9D54CA43FE77FC32FDD2FC5D" box="[412,561,945,971]" italics="true" pageId="14" pageNumber="15">Artines maya</emphasis>
A. Warren &amp; Dolibaina
</taxonomicName>
det. 2015 /.
</paragraph>
<paragraph id="4344E72D9D54CA43FF2CFC57FDCEFBA0" blockId="14.[151,1437,152,1079]" pageId="14" pageNumber="15">
<emphasis id="718F3B3F9D54CA43FF2CFC57FE52FC78" bold="true" box="[199,433,980,1007]" pageId="14" pageNumber="15">Examined material</emphasis>
. The
<typeStatus id="9C40598F9D54CA43FE12FC57FDB8FC79" box="[505,603,980,1006]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
and the following two
<typeStatus id="9C40598F9D54CA43FC9FFC56FC07FC78" box="[884,996,981,1007]" pageId="14" pageNumber="15" type="paratype">paratypes</typeStatus>
: MEXICO—
<emphasis id="718F3B3F9D54CA43FB97FC57FB3AFC79" box="[1148,1241,980,1006]" italics="true" pageId="14" pageNumber="15">
<collectingRegion id="813F29CF9D54CA43FB97FC57FB3AFC79" box="[1148,1241,980,1006]" country="Mexico" name="Chiapas" pageId="14" pageNumber="15">Chiapas</collectingRegion>
</emphasis>
:
<emphasis id="718F3B3F9D54CA43FB07FC57FABCFC78" bold="true" box="[1260,1375,980,1007]" pageId="14" pageNumber="15">Lacantún</emphasis>
(Río Lacantún, Chajul),
<quantity id="84034AC89D54CA43FE9AFC7AFE50FB84" box="[369,435,1017,1043]" metricMagnitude="2" metricUnit="m" metricValue="1.48" pageId="14" pageNumber="15" unit="m" value="148.0">148m</quantity>
,
<specimenCount id="55FD2CA49D54CA43FE54FC7AFDEDFB85" box="[447,526,1016,1043]" pageId="14" pageNumber="15" type="male">1 male</specimenCount>
,
<date id="3745C1ED9D54CA43FDF1FC7AFD6FFB84" box="[538,652,1017,1043]" pageId="14" pageNumber="15" value="1981-05-27">27.V.1981</date>
(
<collectionCode id="25EA7FE89D54CA43FD77FC7AFD16FB84" box="[668,757,1017,1043]" pageId="14" pageNumber="15">MAZA</collectionCode>
). GUATEMALA—
<emphasis id="718F3B3F9D54CA43FC39FC7BFBFBFB85" box="[978,1048,1016,1042]" italics="true" pageId="14" pageNumber="15">
<collectingRegion id="813F29CF9D54CA43FC39FC7BFBFBFB85" box="[978,1048,1016,1042]" country="Guatemala" name="Izabal" pageId="14" pageNumber="15">Izabal</collectingRegion>
</emphasis>
:
<emphasis id="718F3B3F9D54CA43FBC3FC7AFB69FB84" bold="true" box="[1064,1162,1017,1043]" pageId="14" pageNumber="15">Morales</emphasis>
(Cayuga),
<quantity id="84034AC89D54CA43FAE2FC7BFADEFB84" box="[1289,1341,1016,1043]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="14" pageNumber="15" unit="m" value="35.0">35m</quantity>
,
<specimenCount id="55FD2CA49D54CA43FAA2FC7AFA7BFB85" box="[1353,1432,1016,1043]" pageId="14" pageNumber="15" type="male">1 male</specimenCount>
, IX, Schaus &amp; Barnes
<emphasis id="718F3B3F9D54CA43FE60FB9FFE57FBA1" box="[395,436,1052,1078]" italics="true" pageId="14" pageNumber="15">leg.</emphasis>
, (
<collectionCode id="25EA7FE89D54CA43FE21FB9EFDC2FBA0" box="[458,545,1053,1079]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="15" type="Museum">USNM</collectionCode>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="C0EFD676B6906C9D1E10E7AAE3B5D060">Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species</mods:title>
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Key to the males of the “
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<emphasis id="718F3B3F9D5CCA4BFE20FB7CFDC2FA8E" bold="true" box="[459,545,1279,1305]" italics="true" pageId="6" pageNumber="7">aepitus</emphasis>
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group”
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1 Uncus with disto-lateral lobes (
<figureCitation id="DBC0FBA89D5CCA4BFDE5FAC7FD87FACD" box="[526,612,1348,1370]" captionStart="FIGURES 27–29" captionStartId="23.[151,264,1797,1822]" captionTargetBox="[153,1436,174,1777]" captionTargetId="figure@23.[151,1436,174,1786]" captionTargetPageId="23" captionText="FIGURES 27–29. Male genitalia of the “aepitus group” species. 27. Artines aepitus, Jusepín, Monagas, Venezuela (OM 29.417). 28. Artines panama sp. nov., holotype. 29. Artines solange sp. nov., holotype. A) lateral view of tegumen, saccus, uncus and gnathos. B) ventral view of saccus. C) dorsal view of tegumen, uncus and gnathos. D) ventral view of tegumen, uncus and gnathos. E) interior view of right valva. F) dorsal view of right valva. G) dorsal view of aedeagus and vesica. H) left lateral view of aedeagus. I) ventral view of aedeagus. J) right lateral view of aedeagus. K) dorsal view of fultura inferior. L) lateral view of fultura inferior. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241047/files/figure.png" pageId="6" pageNumber="7">Fig. 29C</figureCitation>
).......................................................
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<emphasis id="718F3B3F9D5CCA4BFB14FAC7FAA9FACE" box="[1279,1354,1348,1369]" italics="true" pageId="6" pageNumber="7">solange</emphasis>
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- Uncus without disto-lateral lobes (
<figureCitation id="DBC0FBA89D5CCA4BFDC6FAE2FD96FAE1" box="[557,629,1377,1399]" captionStart="FIGURES 27–29" captionStartId="23.[151,264,1797,1822]" captionTargetBox="[153,1436,174,1777]" captionTargetId="figure@23.[151,1436,174,1786]" captionTargetPageId="23" captionText="FIGURES 27–29. Male genitalia of the “aepitus group” species. 27. Artines aepitus, Jusepín, Monagas, Venezuela (OM 29.417). 28. Artines panama sp. nov., holotype. 29. Artines solange sp. nov., holotype. A) lateral view of tegumen, saccus, uncus and gnathos. B) ventral view of saccus. C) dorsal view of tegumen, uncus and gnathos. D) ventral view of tegumen, uncus and gnathos. E) interior view of right valva. F) dorsal view of right valva. G) dorsal view of aedeagus and vesica. H) left lateral view of aedeagus. I) ventral view of aedeagus. J) right lateral view of aedeagus. K) dorsal view of fultura inferior. L) lateral view of fultura inferior. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241047/files/figure.png" pageId="6" pageNumber="7">Figs 27</figureCitation>
C–28C, 30C–35C)................................................... 2
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2 Harpe with ventral process broad, non-cylindrical and longer than tegumen (
<figureCitation id="DBC0FBA89D5CCA4BFC2BFAFDFBEBFA03" box="[960,1032,1406,1428]" captionStart="FIGURES 27–29" captionStartId="23.[151,264,1797,1822]" captionTargetBox="[153,1436,174,1777]" captionTargetId="figure@23.[151,1436,174,1786]" captionTargetPageId="23" captionText="FIGURES 27–29. Male genitalia of the “aepitus group” species. 27. Artines aepitus, Jusepín, Monagas, Venezuela (OM 29.417). 28. Artines panama sp. nov., holotype. 29. Artines solange sp. nov., holotype. A) lateral view of tegumen, saccus, uncus and gnathos. B) ventral view of saccus. C) dorsal view of tegumen, uncus and gnathos. D) ventral view of tegumen, uncus and gnathos. E) interior view of right valva. F) dorsal view of right valva. G) dorsal view of aedeagus and vesica. H) left lateral view of aedeagus. I) ventral view of aedeagus. J) right lateral view of aedeagus. K) dorsal view of fultura inferior. L) lateral view of fultura inferior. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241047/files/figure.png" pageId="6" pageNumber="7">Figs 27</figureCitation>
E–30E).......................... 3
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- Harpe with ventral process narrow and cylindrical (
<figureCitation id="DBC0FBA89D5CCA4BFD32FA1FFCC1FA26" box="[729,802,1435,1458]" captionStart="FIGURES 33–35" captionStartId="25.[151,264,1776,1801]" captionTargetBox="[151,1436,187,1743]" captionTargetId="figure@25.[151,1436,181,1750]" captionTargetPageId="25" captionText="FIGURES 33–35. Male genitalia of the “aepitus group” species. 33. Artines rica, Turrialba, Cartago, Costa Rica (DZ 4.061). 34. Artines liege sp. nov., holotype. 35. Artines bamba sp. nov., paratype, Pakitza, Parque Nacional del Manu, Madre de Díos, Peru (DZ 9.728). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241051/files/figure.png" pageId="6" pageNumber="7">Figs 33</figureCitation>
E–35E) or short and triangular, in this case shorter than tegumen (
<figureCitation id="DBC0FBA89D5CCA4BFF02FA3AFECCFA58" box="[233,303,1465,1487]" captionStart="FIGURES 30–32" captionStartId="24.[151,264,1772,1797]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure@24.[151,1436,181,1746]" captionTargetPageId="24" captionText="FIGURES 30–32. Male genitalia of the “aepitus group” species. 30. Artines delfos sp. nov., holotype. 31. Artines maya, holotype. 32. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.215). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241049/files/figure.png" pageId="6" pageNumber="7">Figs 31</figureCitation>
E–32E)....................................................................................... 5
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3 Harpe with dorsal process on proximal margin; ventral process longer than tegumen+uncus, with distal end angular (
<figureCitation id="DBC0FBA89D5CCA4BFA9EFA55FEE9F99D" captionStart="FIGURES 27–29" captionStartId="23.[151,264,1797,1822]" captionTargetBox="[153,1436,174,1777]" captionTargetId="figure@23.[151,1436,174,1786]" captionTargetPageId="23" captionText="FIGURES 27–29. Male genitalia of the “aepitus group” species. 27. Artines aepitus, Jusepín, Monagas, Venezuela (OM 29.417). 28. Artines panama sp. nov., holotype. 29. Artines solange sp. nov., holotype. A) lateral view of tegumen, saccus, uncus and gnathos. B) ventral view of saccus. C) dorsal view of tegumen, uncus and gnathos. D) ventral view of tegumen, uncus and gnathos. E) interior view of right valva. F) dorsal view of right valva. G) dorsal view of aedeagus and vesica. H) left lateral view of aedeagus. I) ventral view of aedeagus. J) right lateral view of aedeagus. K) dorsal view of fultura inferior. L) lateral view of fultura inferior. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241047/files/figure.png" pageId="6" pageNumber="7">Fig. 27E</figureCitation>
)..........................................................................................
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- Harpe with the dorsal process away from the proximal margin; ventral process shorter than tegumen+uncus, with angular or rounded distal end (
<figureCitation id="DBC0FBA89D5CCA4BFE4BF9ADFE1AF9D4" box="[416,505,1582,1604]" captionStart="FIGURES 27–29" captionStartId="23.[151,264,1797,1822]" captionTargetBox="[153,1436,174,1777]" captionTargetId="figure@23.[151,1436,174,1786]" captionTargetPageId="23" captionText="FIGURES 27–29. Male genitalia of the “aepitus group” species. 27. Artines aepitus, Jusepín, Monagas, Venezuela (OM 29.417). 28. Artines panama sp. nov., holotype. 29. Artines solange sp. nov., holotype. A) lateral view of tegumen, saccus, uncus and gnathos. B) ventral view of saccus. C) dorsal view of tegumen, uncus and gnathos. D) ventral view of tegumen, uncus and gnathos. E) interior view of right valva. F) dorsal view of right valva. G) dorsal view of aedeagus and vesica. H) left lateral view of aedeagus. I) ventral view of aedeagus. J) right lateral view of aedeagus. K) dorsal view of fultura inferior. L) lateral view of fultura inferior. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241047/files/figure.png" pageId="6" pageNumber="7">Figs 28E</figureCitation>
,
<figureCitation id="DBC0FBA89D5CCA4BFDE8F9ADFDC8F9D3" box="[515,555,1582,1604]" captionStart="FIGURES 30–32" captionStartId="24.[151,264,1772,1797]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure@24.[151,1436,181,1746]" captionTargetPageId="24" captionText="FIGURES 30–32. Male genitalia of the “aepitus group” species. 30. Artines delfos sp. nov., holotype. 31. Artines maya, holotype. 32. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.215). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241049/files/figure.png" pageId="6" pageNumber="7">30E</figureCitation>
).......................................................................
<emphasis id="718F3B3F9D5CCA4BFA7BF9ADFA7FF9D4" bold="true" box="[1424,1436,1582,1603]" pageId="6" pageNumber="7">4</emphasis>
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<paragraph id="4344E72D9D5CCA4BFF7CF9CFFA7FF9F6" blockId="6.[151,1437,1348,1956]" box="[151,1436,1612,1634]" pageId="6" pageNumber="7">
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4 Harpe with distal end of ventral process dilated and rounded (
<figureCitation id="DBC0FBA89D5CCA4BFCD9F9CFFC64F9F5" box="[818,903,1612,1634]" captionStart="FIGURES 27–29" captionStartId="23.[151,264,1797,1822]" captionTargetBox="[153,1436,174,1777]" captionTargetId="figure@23.[151,1436,174,1786]" captionTargetPageId="23" captionText="FIGURES 27–29. Male genitalia of the “aepitus group” species. 27. Artines aepitus, Jusepín, Monagas, Venezuela (OM 29.417). 28. Artines panama sp. nov., holotype. 29. Artines solange sp. nov., holotype. A) lateral view of tegumen, saccus, uncus and gnathos. B) ventral view of saccus. C) dorsal view of tegumen, uncus and gnathos. D) ventral view of tegumen, uncus and gnathos. E) interior view of right valva. F) dorsal view of right valva. G) dorsal view of aedeagus and vesica. H) left lateral view of aedeagus. I) ventral view of aedeagus. J) right lateral view of aedeagus. K) dorsal view of fultura inferior. L) lateral view of fultura inferior. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241047/files/figure.png" pageId="6" pageNumber="7">Fig. 28E</figureCitation>
)..............................
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<emphasis id="718F3B3F9D5CCA4BFB17F9CFFAAAF9F6" box="[1276,1353,1612,1633]" italics="true" pageId="6" pageNumber="7">panama</emphasis>
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<taxonomicNameLabel id="6ABC86449D5CCA4BFABBF9CFFA7FF9F6" box="[1360,1436,1612,1633]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
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<keyLead id="F80A44189D5CCA4BFF7CF9EAFA7FF9E9" box="[151,1436,1641,1663]" pageId="6" pageNumber="7">
- Harpe with distal end of ventral process angled (
<figureCitation id="DBC0FBA89D5CCA4BFD5FF9EAFCEBF9E8" box="[692,776,1641,1663]" captionStart="FIGURES 30–32" captionStartId="24.[151,264,1772,1797]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure@24.[151,1436,181,1746]" captionTargetPageId="24" captionText="FIGURES 30–32. Male genitalia of the “aepitus group” species. 30. Artines delfos sp. nov., holotype. 31. Artines maya, holotype. 32. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.215). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241049/files/figure.png" pageId="6" pageNumber="7">Fig. 30E</figureCitation>
)..........................................
<taxonomicName id="84FB9CAE9D5CCA4BFAFBF9EAFAAAF9E9" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[1296,1353,1641,1662]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="delfos" status="sp. nov.">
<emphasis id="718F3B3F9D5CCA4BFAFBF9EAFAAAF9E9" box="[1296,1353,1641,1662]" italics="true" pageId="6" pageNumber="7">delfos</emphasis>
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<emphasis id="718F3B3F9D5CCA4BFABBF9EAFA7FF9E9" bold="true" box="[1360,1436,1641,1662]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="6ABC86449D5CCA4BFABBF9EAFA7FF9E9" box="[1360,1436,1641,1662]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
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<keyStep id="F80FFF889D5CCA4BFF7CF905FA7FF92E" pageId="6" pageNumber="7">
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<keyLead id="F80A44189D5CCA4BFF7CF905FA7FF90B" box="[151,1436,1670,1692]" pageId="6" pageNumber="7">
5 Harpe with long and cylindrical ventral process (
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E–35E)................................................ 6
</keyLead>
</paragraph>
<paragraph id="4344E72D9D5CCA4BFF7CF927FA7FF92E" blockId="6.[151,1437,1348,1956]" box="[151,1436,1700,1722]" pageId="6" pageNumber="7">
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- Harpe with short and triangular ventral process (
<figureCitation id="DBC0FBA89D5CCA4BFD59F927FD1AF92D" box="[690,761,1700,1722]" captionStart="FIGURES 30–32" captionStartId="24.[151,264,1772,1797]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure@24.[151,1436,181,1746]" captionTargetPageId="24" captionText="FIGURES 30–32. Male genitalia of the “aepitus group” species. 30. Artines delfos sp. nov., holotype. 31. Artines maya, holotype. 32. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.215). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241049/files/figure.png" pageId="6" pageNumber="7">Figs 31</figureCitation>
E–32E)................................................ 7
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<keyStep id="F80FFF889D5CCA4BFF7CF942FA7FF964" pageId="6" pageNumber="7">
<paragraph id="4344E72D9D5CCA4BFF7CF942FA7FF941" blockId="6.[151,1437,1348,1956]" box="[151,1436,1729,1751]" pageId="6" pageNumber="7">
<keyLead id="F80A44189D5CCA4BFF7CF942FA7FF941" box="[151,1436,1729,1751]" pageId="6" pageNumber="7">
6 Harpe with ventral process strongly arched in dorsal view, with bristles on subapical portion (
<figureCitation id="DBC0FBA89D5CCA4BFB8FF942FB54F940" box="[1124,1207,1729,1751]" captionStart="FIGURES 33–35" captionStartId="25.[151,264,1776,1801]" captionTargetBox="[151,1436,187,1743]" captionTargetId="figure@25.[151,1436,181,1750]" captionTargetPageId="25" captionText="FIGURES 33–35. Male genitalia of the “aepitus group” species. 33. Artines rica, Turrialba, Cartago, Costa Rica (DZ 4.061). 34. Artines liege sp. nov., holotype. 35. Artines bamba sp. nov., paratype, Pakitza, Parque Nacional del Manu, Madre de Díos, Peru (DZ 9.728). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241051/files/figure.png" pageId="6" pageNumber="7">Fig. 35F</figureCitation>
)......
<taxonomicName id="84FB9CAE9D5CCA4BFAECF942FAABF941" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[1287,1352,1729,1750]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bamba" status="sp. nov.">
<emphasis id="718F3B3F9D5CCA4BFAECF942FAABF941" box="[1287,1352,1729,1750]" italics="true" pageId="6" pageNumber="7">bamba</emphasis>
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<emphasis id="718F3B3F9D5CCA4BFAA5F942FA7FF941" bold="true" box="[1358,1436,1729,1750]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="6ABC86449D5CCA4BFAA5F942FA7FF941" box="[1358,1436,1729,1750]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
<paragraph id="4344E72D9D5CCA4BFF7CF95CFA7FF964" blockId="6.[151,1437,1348,1956]" box="[151,1436,1758,1780]" pageId="6" pageNumber="7">
<keyLead id="F80A44189D5CCA4BFF7CF95CFA7FF964" box="[151,1436,1758,1780]" pageId="6" pageNumber="7">
- Harpe with ventral process slightly arched in dorsal view, with bristles on apical portion (
<figureCitation id="DBC0FBA89D5CCA4BFBD7F95DFB60F963" box="[1084,1155,1758,1780]" captionStart="FIGURES 33–35" captionStartId="25.[151,264,1776,1801]" captionTargetBox="[151,1436,187,1743]" captionTargetId="figure@25.[151,1436,181,1750]" captionTargetPageId="25" captionText="FIGURES 33–35. Male genitalia of the “aepitus group” species. 33. Artines rica, Turrialba, Cartago, Costa Rica (DZ 4.061). 34. Artines liege sp. nov., holotype. 35. Artines bamba sp. nov., paratype, Pakitza, Parque Nacional del Manu, Madre de Díos, Peru (DZ 9.728). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241051/files/figure.png" pageId="6" pageNumber="7">Figs 33</figureCitation>
F–34F)................ 8
</keyLead>
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<paragraph id="4344E72D9D5CCA4BFF7CF97FFA7FF886" blockId="6.[151,1437,1348,1956]" box="[151,1436,1788,1810]" pageId="6" pageNumber="7">
<keyLead id="F80A44189D5CCA4BFF7CF97FFA7FF886" box="[151,1436,1788,1810]" pageId="6" pageNumber="7">
7 Ampulla distally curved ventrally (
<figureCitation id="DBC0FBA89D5CCA4BFDD9F97FFD6AF885" box="[562,649,1788,1810]" captionStart="FIGURES 30–32" captionStartId="24.[151,264,1772,1797]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure@24.[151,1436,181,1746]" captionTargetPageId="24" captionText="FIGURES 30–32. Male genitalia of the “aepitus group” species. 30. Artines delfos sp. nov., holotype. 31. Artines maya, holotype. 32. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.215). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241049/files/figure.png" pageId="6" pageNumber="7">Fig. 31E</figureCitation>
); distribution restricted to Central America (
<figureCitation id="DBC0FBA89D5CCA4BFBD9F97FFB9BF885" box="[1074,1144,1788,1810]" captionStart="FIGURE 78" captionStartId="42.[151,250,910,935]" captionTargetBox="[151,1436,174,898]" captionTargetId="figure@42.[151,1436,174,898]" captionTargetPageId="42" captionText="FIGURE 78. Geographic distribution of the “aepitus group” species. The symbols with white inferior half represent records from literature." httpUri="https://zenodo.org/record/3241067/files/figure.png" pageId="6" pageNumber="7">Fig. 78</figureCitation>
)...................
<taxonomicName id="84FB9CAE9D5CCA4BFA83F97FFA7FF886" authorityName="A. Warren &amp; Dolibaina" authorityYear="2015" box="[1384,1436,1788,1809]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="maya">
<emphasis id="718F3B3F9D5CCA4BFA83F97FFA7FF886" box="[1384,1436,1788,1809]" italics="true" pageId="6" pageNumber="7">maya</emphasis>
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<paragraph id="4344E72D9D5CCA4BFF7CF89AFA7FF8B9" blockId="6.[151,1437,1348,1956]" box="[151,1436,1817,1839]" pageId="6" pageNumber="7">
<keyLead id="F80A44189D5CCA4BFF7CF89AFA7FF8B9" box="[151,1436,1817,1839]" pageId="6" pageNumber="7">
- Ampulla straight (
<figureCitation id="DBC0FBA89D5CCA4BFE7AF89AFE0BF8B8" box="[401,488,1817,1839]" captionStart="FIGURES 30–32" captionStartId="24.[151,264,1772,1797]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure@24.[151,1436,181,1746]" captionTargetPageId="24" captionText="FIGURES 30–32. Male genitalia of the “aepitus group” species. 30. Artines delfos sp. nov., holotype. 31. Artines maya, holotype. 32. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.215). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241049/files/figure.png" pageId="6" pageNumber="7">Fig. 32E</figureCitation>
); distribution restricted to the Brazilian coast (
<figureCitation id="DBC0FBA89D5CCA4BFC4DF89AFC08F8B8" box="[934,1003,1817,1839]" captionStart="FIGURE 79" captionStartId="42.[151,250,1782,1807]" captionTargetBox="[151,1436,1021,1770]" captionTargetId="figure@42.[151,1436,1021,1770]" captionTargetPageId="42" captionText="FIGURE 79. Geographic distribution of the “aepitus group” species. The symbols with white inferior half represent records from literature." httpUri="https://zenodo.org/record/3241069/files/figure.png" pageId="6" pageNumber="7">Fig. 79</figureCitation>
).....................
<taxonomicName id="84FB9CAE9D5CCA4BFB16F89AFAAAF8B9" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[1277,1353,1817,1838]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="litoralis" status="sp. nov.">
<emphasis id="718F3B3F9D5CCA4BFB16F89AFAAAF8B9" box="[1277,1353,1817,1838]" italics="true" pageId="6" pageNumber="7">litoralis</emphasis>
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<emphasis id="718F3B3F9D5CCA4BFABBF89AFA7FF8B9" bold="true" box="[1360,1436,1817,1838]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="6ABC86449D5CCA4BFABBF89AFA7FF8B9" box="[1360,1436,1817,1838]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
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<keyLead id="F80A44189D5CCA4BFF7CF8B5FA7FF8FE" pageId="6" pageNumber="7">
8 Cornutus as a triangular plate, containing thick spines directed laterally (
<figureCitation id="DBC0FBA89D5CCA4BFC7DF8B5FC3EF8DB" box="[918,989,1846,1868]" captionStart="FIGURES 42–48" captionStartId="33.[151,264,1892,1917]" captionTargetBox="[151,1431,625,1874]" captionTargetId="figure@33.[151,1436,625,1880]" captionText="FIGURES 42–48. Dorsal view of vesica and cornutus of the &quot;aepitus group&quot; species. 42. Artines aepitus. 43. Artines panama sp. nov.. 44. Artines delfos sp. nov.. 45. Artines litoralis sp. nov.. 46. Artines rica. 47. Artines liege sp. nov. 48. Artines bamba sp. nov. Scale bar = 0,2 mm." httpUri="https://zenodo.org/record/3406274/files/figure.png" pageId="6" pageNumber="7">Fig. 47</figureCitation>
); distribution restricted to South America (
<figureCitation id="DBC0FBA89D5CCA4BFA9EF8B5FF18F8FD" captionStart="FIGURE 79" captionStartId="42.[151,250,1782,1807]" captionTargetBox="[151,1436,1021,1770]" captionTargetId="figure@42.[151,1436,1021,1770]" captionTargetPageId="42" captionText="FIGURE 79. Geographic distribution of the “aepitus group” species. The symbols with white inferior half represent records from literature." httpUri="https://zenodo.org/record/3241069/files/figure.png" pageId="6" pageNumber="7">Fig. 79</figureCitation>
).......................................................................................
<taxonomicName id="84FB9CAE9D5CCA4BFAF1F8D7FAAAF8FE" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[1306,1353,1876,1897]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="liege" status="sp. nov.">
<emphasis id="718F3B3F9D5CCA4BFAF1F8D7FAAAF8FE" box="[1306,1353,1876,1897]" italics="true" pageId="6" pageNumber="7">liege</emphasis>
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<emphasis id="718F3B3F9D5CCA4BFAA5F8D7FA7FF8FE" bold="true" box="[1358,1436,1876,1897]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="6ABC86449D5CCA4BFAA5F8D7FA7FF8FE" box="[1358,1436,1876,1897]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
<paragraph id="4344E72D9D5CCA4BFF7CF8F2FA7EF833" blockId="6.[151,1437,1348,1956]" pageId="6" pageNumber="7">
<keyLead id="F80A44189D5CCA4BFF7CF8F2FA7EF833" pageId="6" pageNumber="7">
- Cornutus as an elongated plate, containing fine spines directed posteriorly (
<figureCitation id="DBC0FBA89D5CCA4BFC5CF8F2FC1CF810" box="[951,1023,1905,1927]" captionStart="FIGURES 42–48" captionStartId="33.[151,264,1892,1917]" captionTargetBox="[151,1431,625,1874]" captionTargetId="figure@33.[151,1436,625,1880]" captionText="FIGURES 42–48. Dorsal view of vesica and cornutus of the &quot;aepitus group&quot; species. 42. Artines aepitus. 43. Artines panama sp. nov.. 44. Artines delfos sp. nov.. 45. Artines litoralis sp. nov.. 46. Artines rica. 47. Artines liege sp. nov. 48. Artines bamba sp. nov. Scale bar = 0,2 mm." httpUri="https://zenodo.org/record/3406274/files/figure.png" pageId="6" pageNumber="7">Fig. 46</figureCitation>
); distribution restricted to Central America (
<figureCitation id="DBC0FBA89D5CCA4BFF02F80DFECCF833" box="[233,303,1934,1956]" captionStart="FIGURE 78" captionStartId="42.[151,250,910,935]" captionTargetBox="[151,1436,174,898]" captionTargetId="figure@42.[151,1436,174,898]" captionTargetPageId="42" captionText="FIGURE 78. Geographic distribution of the “aepitus group” species. The symbols with white inferior half represent records from literature." httpUri="https://zenodo.org/record/3241067/files/figure.png" pageId="6" pageNumber="7">Fig. 78</figureCitation>
)..........................................................................................
<taxonomicName id="84FB9CAE9D5CCA4BFA9DF80CFA7EF833" authorityName=", S. R. Steinhauser &amp; G. T. Austin" authorityYear="1993" box="[1398,1437,1935,1956]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="rica">
<emphasis id="718F3B3F9D5CCA4BFA9DF80CFA7EF833" box="[1398,1437,1935,1956]" italics="true" pageId="6" pageNumber="7">rica</emphasis>
</taxonomicName>
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<heading id="180C50419D5FCA48FF7CFCF8FDB1FC02" bold="true" box="[151,594,891,917]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
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Key to the species groups of
<taxonomicName id="84FB9CAE9D5FCA48FE11FCF8FDB1FC02" ID-CoL="922ZR" authorityName="Godman" authorityYear="1901" box="[506,594,891,917]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="718F3B3F9D5FCA48FE11FCF8FDB1FC02" bold="true" box="[506,594,891,917]" italics="true" pageId="5" pageNumber="6">Artines</emphasis>
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1 Forewing with opaque and reduced discal spots, rarely absent (
<figureCitation id="DBC0FBA89D5FCA48FCD9FC3CFC77FC42" box="[818,916,959,981]" captionStart-0="FIGURES 5–8" captionStart-1="FIGURES 9–13" captionStartId-0="6.[151,264,1064,1089]" captionStartId-1="7.[151,264,1180,1205]" captionTargetBox-0="[154,1432,269,1044]" captionTargetBox-1="[152,1436,180,1162]" captionTargetId-0="figure@6.[151,1436,265,1052]" captionTargetId-1="figure@7.[151,1436,174,1169]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURES 5–8. Species of the “aepitus group” in dorsal and ventral views. 5. Artines aepitus, A–B: male, Cueva de Las Pavas, Tingo Maria, Huánuco, Peru (OM 80.822), C–D: female, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.790). 6. Artines panama sp. nov., A–B: holotype male, C–D: allotype female. 7. Artines solange sp. nov., A–B: holotype male, C–D: tentative female, Sumaúma, Maranhão, Brazil (DZ 41.800). 8. Artines delfos sp. nov., A–B: holotype male. Scale bar = 1 cm." captionText-1="FIGURES 9–13. Species of the “aepitus group” in dorsal and ventral views. 9. Artines maya, A–B: holotype male. 10. Artines litoralis sp. nov., A–B: holotype male, C–D: allotype female. 11. Artines rica, A–B: male, Turrialba, Cartago, Costa Rica (DZ 41.890), C–D: female from same locality of male (DZ 41.840). 12. Artines liege sp. nov., A–B: holotype male, C–D: allotype female. 13. Artines bamba sp. nov., A–B: holotype male, C–D: tentative female, Moyobamba, San Martin, Peru (OM 79.413). Scale bar = 1cm." httpUri-0="https://zenodo.org/record/3241041/files/figure.png" httpUri-1="https://zenodo.org/record/3241043/files/figure.png" pageId="5" pageNumber="6">Figs 5–13</figureCitation>
); male without brand; female with region between seventh sternite and lamella antevaginalis sclerotized, forming a rounded plate that covers the ostium ventrally; lamella postvaginalis completely sclerotized (
<figureCitation id="DBC0FBA89D5FCA48FE10FC79FDA7FB87" box="[507,580,1018,1040]" captionStart="FIGURES 54–60" captionStartId="35.[151,264,1807,1832]" captionTargetBox="[156,1436,575,1772]" captionTargetId="figure@35.[151,1436,589,1795]" captionTargetPageId="35" captionText="FIGURES 54–60. Female genitalia of the “aepitus group” species. 54. Artines aepitus, Jusepín, Monagas, Venezuela (OM 30. 125). 55. Artines panama sp. nov., allotype. 56. Artines solange sp. nov. (tentative female), Sumaúma, Maranhão, Brazil (DZ 41.800). 57. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.820). 58. Artines rica, Turrialba, Cartago, Costa Rica (DZ 41.840). 59. Artines liege sp. nov., paratype, Parque do Gama, Brasília, Distrito Federal, Brazil (DZ 41.830). 60. Artines bamba sp. nov. (tentative female), Moyobamba, San Martin, Peru (OM 81.229). A) lateral view of tergum VIII, sterigma and papilla analis; B) ventral view of esterigma. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241059/files/figure.png" pageId="5" pageNumber="6">Figs 54</figureCitation>
B–60B)................................................... “
<taxonomicName id="84FB9CAE9D5FCA48FAE2FC78FAADFB87" ID-CoL="98KNF" authorityName="Geyer" authorityYear="1832" box="[1289,1358,1019,1040]" class="Insecta" family="Hesperiidae" genus="Thracides" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D5FCA48FAE2FC78FAADFB87" box="[1289,1358,1019,1040]" italics="true" pageId="5" pageNumber="6">aepitus</emphasis>
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Group”
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</paragraph>
<paragraph id="4344E72D9D5FCA48FF7CFB9BFA7FFBDD" blockId="5.[151,1437,959,1246]" pageId="5" pageNumber="6">
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- Forewing with large semi-hyaline discal spots, always present (
<figureCitation id="DBC0FBA89D5FCA48FCD4FB94FC4DFBB9" box="[831,942,1047,1070]" captionStart="FIGURES 14–19" captionStartId="10.[151,264,1406,1431]" captionTargetBox="[155,1435,175,1386]" captionTargetId="figure@10.[151,1436,174,1394]" captionTargetPageId="10" captionText="FIGURES 14–19. Species of the “aquilina group” (14–18) and “bipunctata group” (19) in dorsal and ventral views. 14. Artines aquilina, A–B: male, Joinville, Santa Catarina, Brazil (DZ 41.321), C–D: female from same locality of male (DZ 41.220). 15. Artines angelica sp. nov., A–B: holotype male, C–D: allotype female. 16. Artines focus, A–B: male, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.233), C–D: female, Pacaraima, Roraima, Brazil (DZ 41.333). 17. Artines cofus sp. nov., A–B: holotype male, C–D: allotype female. 18. Artines donia sp. nov., A–B: holotype male. 19. Artines bipunctata, A–B: male, Jardim Zoológico, Brasília, Distrito Federal, Brazil (DZ 30.961), C–D: female, Córrego Bananal, Brasília, Distrito Federal, Brazil (DZ 41.224). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241045/files/figure.png" pageId="5" pageNumber="6">Figs 14–19</figureCitation>
); male with brand (
<figureCitation id="DBC0FBA89D5FCA48FB8CFB94FB34FBBA" box="[1127,1239,1047,1069]" captionStart="FIGURES 21–26" captionStartId="22.[151,264,1840,1865]" captionTargetBox="[151,1436,922,1828]" captionTargetId="figure@22.[151,1436,922,1828]" captionText="FIGURES 21–26. Male brands (no scales) on dorsal forewing of the “aquilina group” (21–25) and “bipunctata group” species (26). 21. Artines aquilina, Joinville, Santa catarina, Brazil, (DZ 41.341). 22. Artines angelica sp. nov., paratype, Fazenda Ter- rasse, Santa Luzia, Maranhão, Brazil (DZ 41.273) 23. Artines focus, Ilha de Maracá, Roraima, Brazil (DZ 41.233). 24. Artines cofus sp. nov., paratype, Santa Rosa do Purus, Acre, Brazil (DZ 41.303). 25. Artines donia sp. nov., paratype, Santa Rita do Araguaia, Goiás, Brazil (OM 66.000). 26. Artines bipunctata, Ponte Funda, Vianópilis, Goiás, Brazil (DZ 41.264). Scale bar = 1 mm." httpUri="https://zenodo.org/record/3406270/files/figure.png" pageId="5" pageNumber="6">Figs 21–26</figureCitation>
); female with region between seventh sternite and lamella antevaginalis membranous; lamella postvaginalis partially sclerotized (
<figureCitation id="DBC0FBA89D5FCA48FB19FBB6FAD9FBDC" box="[1266,1338,1077,1099]" captionStart="FIGURES 61–65" captionStartId="36.[151,264,653,678]" captionTargetBox="[151,1435,178,637]" captionTargetId="figure@36.[151,1436,174,642]" captionTargetPageId="36" captionText="FIGURES 61–65. Female genitalia of the “aquilina group” and “bipunctata group” species. 61. Artines aquilina, Cananéia, São Paulo, Brazil (OM 49.831). 62. Artines angelica sp. nov., paratype, Tambopata, Puerto Maldonado, Madre de Díos, Peru (DZ 41.222) 63. Artines focus, Pacaraima, Roraima, Brazil (DZ 41.333). 64. Artines cofus sp. nov. paratype, Tambopata, Puerto Maldonado, Madre de Díos, Peru (DZ 41. 293). 65. Artines bipunctata, Reserva IBGE, Brasília, Distrito Federal, Brazil (DZ 41.165). For reference of the letters see legend of the figure 54. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241061/files/figure.png" pageId="5" pageNumber="6">Figs 61</figureCitation>
B–65B) 2
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2 Ventral hind wing with central black spot on discal area (
<figureCitation id="DBC0FBA89D5FCA48FCE3FBD1FC9AFBFF" box="[776,889,1106,1128]" captionStart="FIGURES 14–19" captionStartId="10.[151,264,1406,1431]" captionTargetBox="[155,1435,175,1386]" captionTargetId="figure@10.[151,1436,174,1394]" captionTargetPageId="10" captionText="FIGURES 14–19. Species of the “aquilina group” (14–18) and “bipunctata group” (19) in dorsal and ventral views. 14. Artines aquilina, A–B: male, Joinville, Santa Catarina, Brazil (DZ 41.321), C–D: female from same locality of male (DZ 41.220). 15. Artines angelica sp. nov., A–B: holotype male, C–D: allotype female. 16. Artines focus, A–B: male, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.233), C–D: female, Pacaraima, Roraima, Brazil (DZ 41.333). 17. Artines cofus sp. nov., A–B: holotype male, C–D: allotype female. 18. Artines donia sp. nov., A–B: holotype male. 19. Artines bipunctata, A–B: male, Jardim Zoológico, Brasília, Distrito Federal, Brazil (DZ 30.961), C–D: female, Córrego Bananal, Brasília, Distrito Federal, Brazil (DZ 41.224). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241045/files/figure.png" pageId="5" pageNumber="6">Figs 14–18</figureCitation>
); valva distally projected with triangular aspect from 1/2 its length; harpe and ampulla fused (
<figureCitation id="DBC0FBA89D5FCA48FDD4FBECFD6BFB12" box="[575,648,1135,1157]" captionStart="FIGURES 36–37" captionStartId="26.[151,264,1557,1582]" captionTargetBox="[157,1435,179,1543]" captionTargetId="figure@26.[151,1436,174,1546]" captionTargetPageId="26" captionText="FIGURES 36–37. Male genitalia of the “aquilina group” species. 36. Artines aquilina, Caminho da Lagoinha, Cachoeiras de Macacú, Rio de Janeiro, Brazil (DZ 41.196). 37. Artines angelica sp. nov., holotype. M) dorsal view of vesica showing the cornuti. For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241053/files/figure.png" pageId="5" pageNumber="6">Figs 36</figureCitation>
E–40E); female with lamella postvaginalis containing lateral folds (
<figureCitation id="DBC0FBA89D5FCA48FAFAFBECFA9CFB12" box="[1297,1407,1135,1157]" captionStart="FIGURES 61–65" captionStartId="36.[151,264,653,678]" captionTargetBox="[151,1435,178,637]" captionTargetId="figure@36.[151,1436,174,642]" captionTargetPageId="36" captionText="FIGURES 61–65. Female genitalia of the “aquilina group” and “bipunctata group” species. 61. Artines aquilina, Cananéia, São Paulo, Brazil (OM 49.831). 62. Artines angelica sp. nov., paratype, Tambopata, Puerto Maldonado, Madre de Díos, Peru (DZ 41.222) 63. Artines focus, Pacaraima, Roraima, Brazil (DZ 41.333). 64. Artines cofus sp. nov. paratype, Tambopata, Puerto Maldonado, Madre de Díos, Peru (DZ 41. 293). 65. Artines bipunctata, Reserva IBGE, Brasília, Distrito Federal, Brazil (DZ 41.165). For reference of the letters see legend of the figure 54. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241061/files/figure.png" pageId="5" pageNumber="6">Figs 61–64</figureCitation>
)........................................................................................“
<taxonomicName id="84FB9CAE9D5FCA48FAEFFB0EFAB7FB35" ID-CoL="GZYD" baseAuthorityName="Plotz" baseAuthorityYear="1882" box="[1284,1364,1165,1186]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="aquilina">
<emphasis id="718F3B3F9D5FCA48FAEFFB0EFAB7FB35" box="[1284,1364,1165,1186]" italics="true" pageId="5" pageNumber="6">aquilina</emphasis>
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- Ventral hind wing without central black spot on discal area (
<figureCitation id="DBC0FBA89D5FCA48FCCBFB29FC84FB57" box="[800,871,1194,1216]" captionStart="FIGURES 14–19" captionStartId="10.[151,264,1406,1431]" captionTargetBox="[155,1435,175,1386]" captionTargetId="figure@10.[151,1436,174,1394]" captionTargetPageId="10" captionText="FIGURES 14–19. Species of the “aquilina group” (14–18) and “bipunctata group” (19) in dorsal and ventral views. 14. Artines aquilina, A–B: male, Joinville, Santa Catarina, Brazil (DZ 41.321), C–D: female from same locality of male (DZ 41.220). 15. Artines angelica sp. nov., A–B: holotype male, C–D: allotype female. 16. Artines focus, A–B: male, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.233), C–D: female, Pacaraima, Roraima, Brazil (DZ 41.333). 17. Artines cofus sp. nov., A–B: holotype male, C–D: allotype female. 18. Artines donia sp. nov., A–B: holotype male. 19. Artines bipunctata, A–B: male, Jardim Zoológico, Brasília, Distrito Federal, Brazil (DZ 30.961), C–D: female, Córrego Bananal, Brasília, Distrito Federal, Brazil (DZ 41.224). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241045/files/figure.png" pageId="5" pageNumber="6">Fig. 19</figureCitation>
); valva distally rounded; harpe and ampulla not fused (
<figureCitation id="DBC0FBA89D5FCA48FA9EFB29FEEFFB49" captionStart="FIGURES 40–41" captionStartId="31.[151,264,1624,1649]" captionTargetBox="[154,1435,484,1599]" captionTargetId="figure@31.[151,1436,481,1612]" captionText="FIGURES 40–41. Male genitalia of the “aquilina group” (40) and “bipunctata group” (41) species. 40. Artines donia sp. nov., paratype, Santa Rita do Araguaia, Goiás, Brazil (DZ 66.000). 41. Artines bipunctata, Brasília, Distrito Federal, Brazil (DZ 41.194). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3406272/files/figure.png" pageId="5" pageNumber="6">Fig. 41E</figureCitation>
); female with lamella postvaginalis without lateral folds (
<figureCitation id="DBC0FBA89D5FCA48FCDEFB44FC68FB4A" box="[821,907,1223,1245]" captionStart="FIGURES 61–65" captionStartId="36.[151,264,653,678]" captionTargetBox="[151,1435,178,637]" captionTargetId="figure@36.[151,1436,174,642]" captionTargetPageId="36" captionText="FIGURES 61–65. Female genitalia of the “aquilina group” and “bipunctata group” species. 61. Artines aquilina, Cananéia, São Paulo, Brazil (OM 49.831). 62. Artines angelica sp. nov., paratype, Tambopata, Puerto Maldonado, Madre de Díos, Peru (DZ 41.222) 63. Artines focus, Pacaraima, Roraima, Brazil (DZ 41.333). 64. Artines cofus sp. nov. paratype, Tambopata, Puerto Maldonado, Madre de Díos, Peru (DZ 41. 293). 65. Artines bipunctata, Reserva IBGE, Brasília, Distrito Federal, Brazil (DZ 41.165). For reference of the letters see legend of the figure 54. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241061/files/figure.png" pageId="5" pageNumber="6">Fig. 65B</figureCitation>
)............................“
<taxonomicName id="84FB9CAE9D5FCA48FB06FB44FAB7FB4B" ID-CoL="GZYF" authorityName="O. Mielke" authorityYear="1968" box="[1261,1364,1223,1244]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bipunctata">
<emphasis id="718F3B3F9D5FCA48FB06FB44FAB7FB4B" box="[1261,1364,1223,1244]" italics="true" pageId="5" pageNumber="6">bipunctata</emphasis>
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<mods:roleTerm id="1F862B4DBF21E9C2C9608018B800C774">Author</mods:roleTerm>
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<mods:namePart id="F8F30671F2682822A23AECA331961D34">Carneiro, Eduardo</mods:namePart>
</mods:name>
<mods:name id="210518C3DC891CE633AA90DC3AEE2F89" type="personal">
<mods:role id="F32B22E9E455678E370C3C068ABA1C05">
<mods:roleTerm id="BAD7F8E7999AF37D0ED06C9B6D130CAA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9F518883BF4286AEA6587EC1FD43E51D">Mielke, Olaf Hermann Hendrik</mods:namePart>
</mods:name>
<mods:name id="941992557F469FAEC5001B13B052F36E" type="personal">
<mods:role id="48F7C27270B55F8818303ED7E2403A4A">
<mods:roleTerm id="9022CFCD253B60A2276F94C2B8F112AD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="54364BB168F6C7B7FCCBBBEC3939D35A">Casagrande, Mirna Martins</mods:namePart>
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<mods:typeOfResource id="02D7D4F53C7890538B1D88A6DB98E4B9">text</mods:typeOfResource>
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<mods:title id="8939A4BF6B65C32961DA2088DFAD5CA6">Zootaxa</mods:title>
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<mods:part id="AC0D5F9CE5E08AD1138EAFA821617F18">
<mods:date id="FCB175489D8C55EEEAB6E3AC0ED5CE01">2019</mods:date>
<mods:detail id="2EC5CB1E75E78C245FD88C1B55633C2A" type="pubDate">
<mods:number id="2DD96CAD88524D01981C84A72B8FEFC9">2019-06-07</mods:number>
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<mods:detail id="34793D90D5251AF747DB6A72F2FA53D5" type="volume">
<mods:number id="E65954FAE308E62C0F29BE4784AB7C27">4614</mods:number>
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<mods:number id="8A697254B55F24B7C804C5CE2C242A29">1</mods:number>
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<mods:classification id="D7F8650B8F0185B56C6A79B52DA3BD5E">journal article</mods:classification>
<mods:identifier id="25790E2B6BD9A1B07BC68347C047EB6B" type="CLB-Dataset">26561</mods:identifier>
<mods:identifier id="5C1E1215486B67501CCB4FAFAE500CCA" type="DOI">10.11646/zootaxa.4614.1.1</mods:identifier>
<mods:identifier id="74C5F1C0C082301E40E9A2DFB712BC61" type="GBIF-Dataset">8e7650ed-66fc-4b2a-b55f-11603d3d2166</mods:identifier>
<mods:identifier id="1540B84CD081D8C54C132A0C76CD85C2" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="259EB4C03433AD34B8785883BA92ED35" type="Zenodo-Dep">3241035</mods:identifier>
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<treatment id="CB52563B9D5FCA4BFF7CFAB3FAE6FF6C" ID-GBIF-Taxon="157435994" LSID="urn:lsid:plazi:treatment:CB52563B9D5FCA4BFF7CFAB3FAE6FF6C" httpUri="http://treatment.plazi.org/id/CB52563B9D5FCA4BFF7CFAB3FAE6FF6C" lastPageId="6" lastPageNumber="6" pageId="5" pageNumber="6">
<subSubSection id="0BE1B4A69D5FCA48FF7CFAB3FEBEFADD" box="[151,349,1328,1354]" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="4344E72D9D5FCA48FF7CFAB3FEBEFADD" blockId="5.[151,711,1328,1390]" box="[151,349,1328,1354]" pageId="5" pageNumber="6">
<heading id="180C50419D5FCA48FF7CFAB3FEBEFADD" bold="true" box="[151,349,1328,1354]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
<emphasis id="718F3B3F9D5FCA48FF7CFAB3FEBEFADD" bold="true" box="[151,349,1328,1354]" pageId="5" pageNumber="6">
“
<taxonomicName id="84FB9CAE9D5FCA48FF4DFAB3FF1FFADD" ID-CoL="98KNF" authorityName="Geyer" authorityYear="1832" box="[166,252,1328,1354]" class="Insecta" family="Hesperiidae" genus="Thracides" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D5FCA48FF4DFAB3FF1FFADD" bold="true" box="[166,252,1328,1354]" italics="true" pageId="5" pageNumber="6">aepitus</emphasis>
</taxonomicName>
group”
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D5FCA4BFF7CFAD7FAE6FF6C" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="description">
<paragraph id="4344E72D9D5FCA48FF7CFAD7FD24FAF9" blockId="5.[151,711,1328,1390]" box="[151,711,1363,1390]" pageId="5" pageNumber="6">
<figureCitation id="DBC0FBA89D5FCA48FF7CFAD7FF3FFAF9" box="[151,220,1364,1390]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,2012,2037]" captionTargetBox="[153,1433,1143,1971]" captionTargetId="figure@2.[151,1436,1127,1998]" captionTargetPageId="2" captionText="FIGURES 1–2. Color pattern of ventral wings of Artines: 1. Artines bamba sp. nov.; 2. Artines aquilina." httpUri="https://zenodo.org/record/3241037/files/figure.png" pageId="5" pageNumber="6">Figs 1</figureCitation>
,
<figureCitation id="DBC0FBA89D5FCA48FF01FAD7FEC6FAF9" box="[234,293,1364,1390]" captionStart-0="FIGURES 3–4" captionStart-1="FIGURES 5–8" captionStart-2="FIGURES 9–13" captionStartId-0="5.[151,264,712,737]" captionStartId-1="6.[151,264,1064,1089]" captionStartId-2="7.[151,264,1180,1205]" captionTargetBox-0="[153,1432,238,691]" captionTargetBox-1="[154,1432,269,1044]" captionTargetBox-2="[152,1436,180,1162]" captionTargetId-0="figure@5.[151,1436,237,693]" captionTargetId-1="figure@6.[151,1436,265,1052]" captionTargetId-2="figure@7.[151,1436,174,1169]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURES 3–4. Androconial organs present in the species of Artines. 3. Brand (with scales) on dorsal forewing of Artines aquilina (absent in “aepitus group”). 4. Tuft of elongated scales along 2A of the dorsal hind wing of Artines liege sp. nov. (presente in all species of Artines)." captionText-1="FIGURES 5–8. Species of the “aepitus group” in dorsal and ventral views. 5. Artines aepitus, A–B: male, Cueva de Las Pavas, Tingo Maria, Huánuco, Peru (OM 80.822), C–D: female, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.790). 6. Artines panama sp. nov., A–B: holotype male, C–D: allotype female. 7. Artines solange sp. nov., A–B: holotype male, C–D: tentative female, Sumaúma, Maranhão, Brazil (DZ 41.800). 8. Artines delfos sp. nov., A–B: holotype male. Scale bar = 1 cm." captionText-2="FIGURES 9–13. Species of the “aepitus group” in dorsal and ventral views. 9. Artines maya, A–B: holotype male. 10. Artines litoralis sp. nov., A–B: holotype male, C–D: allotype female. 11. Artines rica, A–B: male, Turrialba, Cartago, Costa Rica (DZ 41.890), C–D: female from same locality of male (DZ 41.840). 12. Artines liege sp. nov., A–B: holotype male, C–D: allotype female. 13. Artines bamba sp. nov., A–B: holotype male, C–D: tentative female, Moyobamba, San Martin, Peru (OM 79.413). Scale bar = 1cm." httpUri-0="https://zenodo.org/record/3241039/files/figure.png" httpUri-1="https://zenodo.org/record/3241041/files/figure.png" httpUri-2="https://zenodo.org/record/3241043/files/figure.png" pageId="5" pageNumber="6">4–13</figureCitation>
,
<figureCitation id="DBC0FBA89D5FCA48FEDBFAD0FE9AFAF9" box="[304,377,1363,1390]" captionStart-0="FIGURES 27–29" captionStart-1="FIGURES 30–32" captionStart-2="FIGURES 33–35" captionStartId-0="23.[151,264,1797,1822]" captionStartId-1="24.[151,264,1772,1797]" captionStartId-2="25.[151,264,1776,1801]" captionTargetBox-0="[153,1436,174,1777]" captionTargetBox-1="[154,1436,183,1733]" captionTargetBox-2="[151,1436,187,1743]" captionTargetId-0="figure@23.[151,1436,174,1786]" captionTargetId-1="figure@24.[151,1436,181,1746]" captionTargetId-2="figure@25.[151,1436,181,1750]" captionTargetPageId-0="23" captionTargetPageId-1="24" captionTargetPageId-2="25" captionText-0="FIGURES 27–29. Male genitalia of the “aepitus group” species. 27. Artines aepitus, Jusepín, Monagas, Venezuela (OM 29.417). 28. Artines panama sp. nov., holotype. 29. Artines solange sp. nov., holotype. A) lateral view of tegumen, saccus, uncus and gnathos. B) ventral view of saccus. C) dorsal view of tegumen, uncus and gnathos. D) ventral view of tegumen, uncus and gnathos. E) interior view of right valva. F) dorsal view of right valva. G) dorsal view of aedeagus and vesica. H) left lateral view of aedeagus. I) ventral view of aedeagus. J) right lateral view of aedeagus. K) dorsal view of fultura inferior. L) lateral view of fultura inferior. Scale bar = 1 mm." captionText-1="FIGURES 30–32. Male genitalia of the “aepitus group” species. 30. Artines delfos sp. nov., holotype. 31. Artines maya, holotype. 32. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.215). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." captionText-2="FIGURES 33–35. Male genitalia of the “aepitus group” species. 33. Artines rica, Turrialba, Cartago, Costa Rica (DZ 4.061). 34. Artines liege sp. nov., holotype. 35. Artines bamba sp. nov., paratype, Pakitza, Parque Nacional del Manu, Madre de Díos, Peru (DZ 9.728). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri-0="https://zenodo.org/record/3241047/files/figure.png" httpUri-1="https://zenodo.org/record/3241049/files/figure.png" httpUri-2="https://zenodo.org/record/3241051/files/figure.png" pageId="5" pageNumber="6">27–35</figureCitation>
,
<figureCitation id="DBC0FBA89D5FCA48FE6FFAD7FE2EFAF9" box="[388,461,1364,1390]" captionStart="FIGURES 42–48" captionStartId="33.[151,264,1892,1917]" captionTargetBox="[151,1431,625,1874]" captionTargetId="figure@33.[151,1436,625,1880]" captionText="FIGURES 42–48. Dorsal view of vesica and cornutus of the &quot;aepitus group&quot; species. 42. Artines aepitus. 43. Artines panama sp. nov.. 44. Artines delfos sp. nov.. 45. Artines litoralis sp. nov.. 46. Artines rica. 47. Artines liege sp. nov. 48. Artines bamba sp. nov. Scale bar = 0,2 mm." httpUri="https://zenodo.org/record/3406274/files/figure.png" pageId="5" pageNumber="6">42–48</figureCitation>
, 54–60, 66–72, 78–79
</paragraph>
<paragraph id="4344E72D9D5FCA48FF7CFA18FDB9F845" blockId="5.[151,1437,1435,2039]" pageId="5" pageNumber="6">
<emphasis id="718F3B3F9D5FCA48FF7CFA18FEC2FA21" bold="true" box="[151,289,1435,1462]" pageId="5" pageNumber="6">Description</emphasis>
.
<emphasis id="718F3B3F9D5FCA48FEDBFA1FFE89FA21" box="[304,362,1436,1462]" italics="true" pageId="5" pageNumber="6">Male</emphasis>
and
<emphasis id="718F3B3F9D5FCA48FE4BFA1FFE08FA21" box="[416,491,1436,1462]" italics="true" pageId="5" pageNumber="6">female</emphasis>
: forewing with opaque and reduced discal spots; ventral forewing with white, broad and diffuse spot on distal 1/2 of CuA
<subScript id="DF7FE5689D5FCA48FDABFA51FDABFA77" attach="left" box="[576,584,1490,1504]" fontSize="6" pageId="5" pageNumber="6">2</subScript>
–2A; ventral forewing with the discal area yellow, containing central black spot, postdiscal band yellow, limited distally by a dark brown not wavy stripe.
<emphasis id="718F3B3F9D5FCA48FBF2FA67FBB0FA69" box="[1049,1107,1508,1534]" italics="true" pageId="5" pageNumber="6">Male</emphasis>
: no brand; saccus triangular; uncus simple and wide, with or without disto-lateral lobes; arms of gnathos with not pointed apex; harpe and ampulla not fused; harpe bifid (except in
<taxonomicName id="84FB9CAE9D5FCA48FDAFF9AEFC22F9D1" authority="Medeiros &amp; O. Mielke" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[580,961,1580,1607]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="solange" status="sp. nov.">
<emphasis id="718F3B3F9D5FCA48FDAFF9AEFD58F9D1" box="[580,699,1580,1607]" italics="true" pageId="5" pageNumber="6">A. solange</emphasis>
Medeiros &amp; O. Mielke
</taxonomicName>
<emphasis id="718F3B3F9D5FCA48FC23F9AFFBC0F9D0" bold="true" box="[968,1059,1580,1607]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="6ABC86449D5FCA48FC23F9AFFBC0F9D0" box="[968,1059,1580,1607]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), with ventral process of variable size, wide or narrow and cylindrical, and the dorsal process with conical and strongly sclerotized spines (
<figureCitation id="DBC0FBA89D5FCA48FAC1F9D3FA6CF9FD" box="[1322,1423,1616,1642]" captionStart="FIGURES 27–29" captionStartId="23.[151,264,1797,1822]" captionTargetBox="[153,1436,174,1777]" captionTargetId="figure@23.[151,1436,174,1786]" captionTargetPageId="23" captionText="FIGURES 27–29. Male genitalia of the “aepitus group” species. 27. Artines aepitus, Jusepín, Monagas, Venezuela (OM 29.417). 28. Artines panama sp. nov., holotype. 29. Artines solange sp. nov., holotype. A) lateral view of tegumen, saccus, uncus and gnathos. B) ventral view of saccus. C) dorsal view of tegumen, uncus and gnathos. D) ventral view of tegumen, uncus and gnathos. E) interior view of right valva. F) dorsal view of right valva. G) dorsal view of aedeagus and vesica. H) left lateral view of aedeagus. I) ventral view of aedeagus. J) right lateral view of aedeagus. K) dorsal view of fultura inferior. L) lateral view of fultura inferior. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241047/files/figure.png" pageId="5" pageNumber="6">Fig. 27E</figureCitation>
); aedeagus with the disto-dorsal expansion triangular (when vesica is not everted it is difficult to observe); ventral projection of aedeagus simple, narrow and long, directed parallel to the longitudinal axis (
<figureCitation id="DBC0FBA89D5FCA48FB78F91BFB10F925" box="[1171,1267,1688,1714]" captionStart="FIGURES 27–29" captionStartId="23.[151,264,1797,1822]" captionTargetBox="[153,1436,174,1777]" captionTargetId="figure@23.[151,1436,174,1786]" captionTargetPageId="23" captionText="FIGURES 27–29. Male genitalia of the “aepitus group” species. 27. Artines aepitus, Jusepín, Monagas, Venezuela (OM 29.417). 28. Artines panama sp. nov., holotype. 29. Artines solange sp. nov., holotype. A) lateral view of tegumen, saccus, uncus and gnathos. B) ventral view of saccus. C) dorsal view of tegumen, uncus and gnathos. D) ventral view of tegumen, uncus and gnathos. E) interior view of right valva. F) dorsal view of right valva. G) dorsal view of aedeagus and vesica. H) left lateral view of aedeagus. I) ventral view of aedeagus. J) right lateral view of aedeagus. K) dorsal view of fultura inferior. L) lateral view of fultura inferior. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241047/files/figure.png" pageId="5" pageNumber="6">Fig. 27J</figureCitation>
); cornutus as a sclerotized plate on distal end of vesica, containing elongate and setae-like spines, sometimes very thick as in
<taxonomicName id="84FB9CAE9D5FCA48FA6FF93EFE06F96D" authority="Medeiros &amp; O. Mielke" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bamba" status="sp. nov.">
<emphasis id="718F3B3F9D5FCA48FA6FF93EFF00F96D" italics="true" pageId="5" pageNumber="6">A. bamba</emphasis>
Medeiros &amp; O. Mielke
</taxonomicName>
<emphasis id="718F3B3F9D5FCA48FE07F963FDA6F96D" bold="true" box="[492,581,1760,1786]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="6ABC86449D5FCA48FE07F963FDA6F96D" box="[492,581,1760,1786]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="DBC0FBA89D5FCA48FDB9F963FD36F96D" box="[594,725,1760,1786]" captionStart="FIGURES 42–48" captionStartId="33.[151,264,1892,1917]" captionTargetBox="[151,1431,625,1874]" captionTargetId="figure@33.[151,1436,625,1880]" captionText="FIGURES 42–48. Dorsal view of vesica and cornutus of the &quot;aepitus group&quot; species. 42. Artines aepitus. 43. Artines panama sp. nov.. 44. Artines delfos sp. nov.. 45. Artines litoralis sp. nov.. 46. Artines rica. 47. Artines liege sp. nov. 48. Artines bamba sp. nov. Scale bar = 0,2 mm." httpUri="https://zenodo.org/record/3406274/files/figure.png" pageId="5" pageNumber="6">Figs 42–48</figureCitation>
); fultura inferior straight in lateral view, with proximal end bifid (
<figureCitation id="DBC0FBA89D5FCA48FF74F887FF17F889" box="[159,244,1795,1822]" captionStart="FIGURES 27–29" captionStartId="23.[151,264,1797,1822]" captionTargetBox="[153,1436,174,1777]" captionTargetId="figure@23.[151,1436,174,1786]" captionTargetPageId="23" captionText="FIGURES 27–29. Male genitalia of the “aepitus group” species. 27. Artines aepitus, Jusepín, Monagas, Venezuela (OM 29.417). 28. Artines panama sp. nov., holotype. 29. Artines solange sp. nov., holotype. A) lateral view of tegumen, saccus, uncus and gnathos. B) ventral view of saccus. C) dorsal view of tegumen, uncus and gnathos. D) ventral view of tegumen, uncus and gnathos. E) interior view of right valva. F) dorsal view of right valva. G) dorsal view of aedeagus and vesica. H) left lateral view of aedeagus. I) ventral view of aedeagus. J) right lateral view of aedeagus. K) dorsal view of fultura inferior. L) lateral view of fultura inferior. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241047/files/figure.png" pageId="5" pageNumber="6">Figs 27</figureCitation>
L–35L).
<emphasis id="718F3B3F9D5FCA48FEBEF887FE4AF889" box="[341,425,1796,1822]" italics="true" pageId="5" pageNumber="6">Female</emphasis>
: externally similar to male; region between seventh sternite and lamella antevaginalis sclerotized, forming a rounded plate that covers the opening of the ostium ventrally; lamella postvaginalis completely sclerotized, ranging from more or less triangular to rectangular; lamella antevaginalis containing proximo-ventral folds (
<figureCitation id="DBC0FBA89D5FCA48FF0BF8F3FEA4F81D" box="[224,327,1903,1930]" captionStart="FIGURES 54–60" captionStartId="35.[151,264,1807,1832]" captionTargetBox="[156,1436,575,1772]" captionTargetId="figure@35.[151,1436,589,1795]" captionTargetPageId="35" captionText="FIGURES 54–60. Female genitalia of the “aepitus group” species. 54. Artines aepitus, Jusepín, Monagas, Venezuela (OM 30. 125). 55. Artines panama sp. nov., allotype. 56. Artines solange sp. nov. (tentative female), Sumaúma, Maranhão, Brazil (DZ 41.800). 57. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.820). 58. Artines rica, Turrialba, Cartago, Costa Rica (DZ 41.840). 59. Artines liege sp. nov., paratype, Parque do Gama, Brasília, Distrito Federal, Brazil (DZ 41.830). 60. Artines bamba sp. nov. (tentative female), Moyobamba, San Martin, Peru (OM 81.229). A) lateral view of tergum VIII, sterigma and papilla analis; B) ventral view of esterigma. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241059/files/figure.png" pageId="5" pageNumber="6">Fig. 54B</figureCitation>
); lamella postvaginalis bifid, with lateral fold present or absent, strongly sclerotized, proximo-lateral expansions simple or bilobed and with variable length (
<figureCitation id="DBC0FBA89D5FCA48FCA8F817FC4FF839" box="[835,940,1939,1966]" captionStart="FIGURES 54–60" captionStartId="35.[151,264,1807,1832]" captionTargetBox="[156,1436,575,1772]" captionTargetId="figure@35.[151,1436,589,1795]" captionTargetPageId="35" captionText="FIGURES 54–60. Female genitalia of the “aepitus group” species. 54. Artines aepitus, Jusepín, Monagas, Venezuela (OM 30. 125). 55. Artines panama sp. nov., allotype. 56. Artines solange sp. nov. (tentative female), Sumaúma, Maranhão, Brazil (DZ 41.800). 57. Artines litoralis sp. nov., paratype, Linhares, Espírito Santo, Brazil (DZ 41.820). 58. Artines rica, Turrialba, Cartago, Costa Rica (DZ 41.840). 59. Artines liege sp. nov., paratype, Parque do Gama, Brasília, Distrito Federal, Brazil (DZ 41.830). 60. Artines bamba sp. nov. (tentative female), Moyobamba, San Martin, Peru (OM 81.229). A) lateral view of tergum VIII, sterigma and papilla analis; B) ventral view of esterigma. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241059/files/figure.png" pageId="5" pageNumber="6">Fig. 54B</figureCitation>
); ductus bursae membranous or with small, irregular and strongly sclerotized plates.
</paragraph>
<paragraph id="4344E72D9D5FCA4BFF2CF85FFAE6FF6C" blockId="5.[151,1437,1435,2039]" lastBlockId="6.[151,1436,153,251]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<emphasis id="718F3B3F9D5FCA48FF2CF85FFE6CF861" bold="true" box="[199,399,2011,2038]" pageId="5" pageNumber="6">Included species</emphasis>
:
<taxonomicName id="84FB9CAE9D5FCA48FE48F85EFCE2F861" authority="(Geyer, 1832)" baseAuthorityName="Geyer" baseAuthorityYear="1832" box="[419,769,2012,2039]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D5FCA48FE48F85EFDB2F861" box="[419,593,2013,2039]" italics="true" pageId="5" pageNumber="6">Artines aepitus</emphasis>
(
<bibRefCitation id="276A9ADC9D5FCA48FD8EF85FFD1AF861" author="Geyer, C." box="[613,761,2012,2038]" pageId="5" pageNumber="6" pagination="1 - 48" refId="ref40941" refString="Geyer, C. (1832) In: Hubner, J. (Ed.), Zutrage zur Sammlung exotischer Schmetterlinge, bestehend in Bekanntmachung einzelner Geschlechter neuer oder seltener nichteuropaischer Arten. Vol. 4. Jacob Hubner, Augsburg, pp. 1 - 48, pls. 138 - 143." type="book chapter" year="1832">Geyer, 1832</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="84FB9CAE9D5FCA48FCFAF85EFB2DF861" authority="Medeiros &amp; Carneiro" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[785,1230,2012,2039]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="panama" status="sp. nov.">
<emphasis id="718F3B3F9D5FCA48FCFAF85EFC2BF861" box="[785,968,2013,2039]" italics="true" pageId="5" pageNumber="6">Artines panama</emphasis>
Medeiros &amp; Carneiro
</taxonomicName>
<emphasis id="718F3B3F9D5FCA48FB31F85FFADAF861" bold="true" box="[1242,1337,2012,2038]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="6ABC86449D5FCA48FB31F85FFADAF861" box="[1242,1337,2012,2038]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="718F3B3F9D5FCA4BFAA0F85EFF0DFF24" italics="true" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<taxonomicName id="84FB9CAE9D5FCA48FAA0F85EFA7FF860" authority="Medeiros &amp; O. Mielke" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[1355,1436,2013,2039]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="13" phylum="Arthropoda" rank="species" species="solange" status="sp. nov.">Artines</taxonomicName>
solange
</emphasis>
Medeiros &amp; O. Mielke
<emphasis id="718F3B3F9D5CCA4BFE1DFF1AFDACFF24" bold="true" box="[502,591,153,179]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="6ABC86449D5CCA4BFE1DFF1AFDACFF24" box="[502,591,153,179]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="84FB9CAE9D5CCA4BFDB0FF1AFBE9FF24" authority="Medeiros &amp; Casagrande" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[603,1034,153,179]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="delfos" status="sp. nov.">
<emphasis id="718F3B3F9D5CCA4BFDB0FF1AFD16FF24" box="[603,757,153,179]" italics="true" pageId="6" pageNumber="7">Artines delfos</emphasis>
Medeiros &amp; Casagrande
</taxonomicName>
<emphasis id="718F3B3F9D5CCA4BFBE4FF1AFB8BFF24" bold="true" box="[1039,1128,153,179]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="6ABC86449D5CCA4BFBE4FF1AFB8BFF24" box="[1039,1128,153,179]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="84FB9CAE9D5CCA4BFB9FFF1AFEAFFF40" authority="A. Warren &amp; Dolibaina, 2015" authorityName="A. Warren &amp; Dolibaina" authorityYear="2015" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="maya">
<emphasis id="718F3B3F9D5CCA4BFB9FFF1AFAEBFF24" box="[1140,1288,153,179]" italics="true" pageId="6" pageNumber="7">Artines maya</emphasis>
A. Warren &amp;
<bibRefCitation id="276A9ADC9D5CCA4BFF7CFF3EFEAFFF40" author="Dolibaina, D. R. &amp; Warren, A. D. &amp; Carneiro, E. &amp; Mielke, O. H. H." box="[151,332,188,215]" pageId="6" pageNumber="7" pagination="52 - 59" refId="ref39956" refString="Dolibaina, D. R., Warren, A. D., Carneiro, E. &amp; Mielke, O. H. H. (2015) Three new species of Artines Godman, 1901 from Central and South America (Hesperiidae: Hesperiinae: Moncini). Tropical Lepidoptera Research, 25 (2), 52 - 59." type="journal article" year="2015">Dolibaina, 2015</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="84FB9CAE9D5CCA4BFEB2FF3EFCEEFF40" authority="Medeiros &amp; Dolibaina" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[345,781,189,215]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="litoralis" status="sp. nov.">
<emphasis id="718F3B3F9D5CCA4BFEB2FF3EFDE9FF40" box="[345,522,189,215]" italics="true" pageId="6" pageNumber="7">Artines litoralis</emphasis>
Medeiros &amp; Dolibaina
</taxonomicName>
<emphasis id="718F3B3F9D5CCA4BFCFFFF3EFC8DFF40" bold="true" box="[788,878,189,215]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="6ABC86449D5CCA4BFCFFFF3EFC8DFF40" box="[788,878,189,215]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="84FB9CAE9D5CCA4BFC97FF3EFADDFF40" authority="Steinhauser &amp; Austin, 1993" authorityName=", S. R. Steinhauser &amp; G. T. Austin" authorityYear="1993" box="[892,1342,189,215]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="rica">
<emphasis id="718F3B3F9D5CCA4BFC97FF3EFBE2FF40" box="[892,1025,189,215]" italics="true" pageId="6" pageNumber="7">Artines rica</emphasis>
<bibRefCitation id="276A9ADC9D5CCA4BFBE3FF3EFADDFF40" author="Steinhauser, S. R. &amp; Austin, G. T." box="[1032,1342,189,215]" pageId="6" pageNumber="7" pagination="12 - 20" refId="ref45097" refString="Steinhauser, S. R. &amp; Austin, G. T. (1993) New species of Hesperiidae from Costa Rica. Tropical Lepidopter a, 4 (Supplement 2), 12 - 20." type="journal article" year="1993">Steinhauser &amp; Austin, 1993</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="84FB9CAE9D5CCA4BFAA0FF3EFD86FF6C" authority="Medeiros, O. Mielke &amp; Casagrande" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="liege" status="sp. nov.">
<emphasis id="718F3B3F9D5CCA4BFAA0FF3EFF2EFF6C" italics="true" pageId="6" pageNumber="7">Artines liege</emphasis>
Medeiros, O. Mielke &amp; Casagrande
</taxonomicName>
<emphasis id="718F3B3F9D5CCA4BFD80FF62FD26FF6C" bold="true" box="[619,709,225,251]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="6ABC86449D5CCA4BFD80FF62FD26FF6C" box="[619,709,225,251]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="84FB9CAE9D5CCA4BFD10FF62FB46FF6C" authority="Medeiros &amp; O. Mielke" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[763,1189,225,251]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bamba" status="sp. nov.">
<emphasis id="718F3B3F9D5CCA4BFD10FF62FC7CFF6C" box="[763,927,225,251]" italics="true" pageId="6" pageNumber="7">Artines bamba</emphasis>
Medeiros &amp; O. Mielke
</taxonomicName>
<emphasis id="718F3B3F9D5CCA4BFB47FF62FAE6FF6C" bold="true" box="[1196,1285,225,251]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="6ABC86449D5CCA4BFB47FF62FAE6FF6C" box="[1196,1285,225,251]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
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<paragraph id="4344E72D9D78CA6FFF7CFB8DFD59FBBF" blockId="34.[151,965,1038,1136]" box="[151,698,1038,1064]" pageId="34" pageNumber="35">
<heading id="180C50419D78CA6FFF7CFB8DFD59FBBF" bold="true" box="[151,698,1038,1064]" fontSize="11" level="1" pageId="34" pageNumber="35" reason="1">
<emphasis id="718F3B3F9D78CA6FFF7CFB8DFD59FBBF" bold="true" box="[151,698,1038,1064]" pageId="34" pageNumber="35">
<taxonomicName id="84FB9CAE9D78CA6FFF7CFB8DFDB6FBBF" authority="Medeiros &amp; Dolibaina" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[151,597,1038,1064]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="cofus" status="sp. nov.">
<emphasis id="718F3B3F9D78CA6FFF7CFB8DFEDBFBBF" bold="true" box="[151,312,1038,1064]" italics="true" pageId="34" pageNumber="35">Artines cofus</emphasis>
Medeiros &amp; Dolibaina
</taxonomicName>
<taxonomicNameLabel id="6ABC86449D78CA6FFDB6FB8DFD59FBBF" box="[605,698,1038,1064]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="0BE1B4A69D78CA6FFF7CFBB1FA91FA87" pageId="34" pageNumber="35" type="description">
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<figureCitation id="DBC0FBA89D78CA6FFF7CFBB1FF0FFBDB" box="[151,236,1074,1100]" captionStart="FIGURES 14–19" captionStartId="10.[151,264,1406,1431]" captionTargetBox="[155,1435,175,1386]" captionTargetId="figure@10.[151,1436,174,1394]" captionTargetPageId="10" captionText="FIGURES 14–19. Species of the “aquilina group” (14–18) and “bipunctata group” (19) in dorsal and ventral views. 14. Artines aquilina, A–B: male, Joinville, Santa Catarina, Brazil (DZ 41.321), C–D: female from same locality of male (DZ 41.220). 15. Artines angelica sp. nov., A–B: holotype male, C–D: allotype female. 16. Artines focus, A–B: male, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.233), C–D: female, Pacaraima, Roraima, Brazil (DZ 41.333). 17. Artines cofus sp. nov., A–B: holotype male, C–D: allotype female. 18. Artines donia sp. nov., A–B: holotype male. 19. Artines bipunctata, A–B: male, Jardim Zoológico, Brasília, Distrito Federal, Brazil (DZ 30.961), C–D: female, Córrego Bananal, Brasília, Distrito Federal, Brazil (DZ 41.224). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241045/files/figure.png" pageId="34" pageNumber="35">Figs 17</figureCitation>
,
<figureCitation id="DBC0FBA89D78CA6FFF13FBB1FEF5FBDB" box="[248,278,1074,1100]" captionStart="FIGURES 21–26" captionStartId="22.[151,264,1840,1865]" captionTargetBox="[151,1436,922,1828]" captionTargetId="figure@22.[151,1436,922,1828]" captionText="FIGURES 21–26. Male brands (no scales) on dorsal forewing of the “aquilina group” (21–25) and “bipunctata group” species (26). 21. Artines aquilina, Joinville, Santa catarina, Brazil, (DZ 41.341). 22. Artines angelica sp. nov., paratype, Fazenda Ter- rasse, Santa Luzia, Maranhão, Brazil (DZ 41.273) 23. Artines focus, Ilha de Maracá, Roraima, Brazil (DZ 41.233). 24. Artines cofus sp. nov., paratype, Santa Rosa do Purus, Acre, Brazil (DZ 41.303). 25. Artines donia sp. nov., paratype, Santa Rita do Araguaia, Goiás, Brazil (OM 66.000). 26. Artines bipunctata, Ponte Funda, Vianópilis, Goiás, Brazil (DZ 41.264). Scale bar = 1 mm." httpUri="https://zenodo.org/record/3406270/files/figure.png" pageId="34" pageNumber="35">24</figureCitation>
, 39, 52, 64, 76, 80
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<paragraph id="4344E72D9D78CA6FFF7CFB33FBA6FB5F" blockId="34.[151,1093,1199,1224]" box="[151,1093,1199,1224]" pageId="34" pageNumber="35">
<taxonomicName id="84FB9CAE9D78CA6FFF7CFB33FEFFFB5F" authorityName=", Reserva Duque, Manaus" authorityYear="1955" box="[151,284,1200,1224]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="focus">
<emphasis id="718F3B3F9D78CA6FFF7CFB33FEFFFB5F" box="[151,284,1200,1224]" italics="true" pageId="34" pageNumber="35">Artines focus</emphasis>
</taxonomicName>
(misid.); O. Mielke; Carneiro &amp; Casagrande, 2010.
<emphasis id="718F3B3F9D78CA6FFCC4FB2CFC0EFB5F" bold="true" box="[815,1005,1199,1224]" pageId="34" pageNumber="35">
<taxonomicName id="84FB9CAE9D78CA6FFCC4FB2CFC2EFB5F" authority="Neotrop." authorityName="Neotrop." box="[815,973,1199,1224]" class="Pinopsida" family="Cupressaceae" genus="Biota" higherTaxonomySource="CoL" kingdom="Plantae" order="Pinales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">Biota Neotrop.</taxonomicName>
10
</emphasis>
(4): 296.
</paragraph>
<paragraph id="4344E72D9D78CA6FFF7CFB76FA91FA87" blockId="34.[151,1437,1269,2017]" box="[151,1394,1269,1297]" pageId="34" pageNumber="35">
<emphasis id="718F3B3F9D78CA6FFF7CFB76FE8AFA87" bold="true" box="[151,361,1269,1296]" pageId="34" pageNumber="35">Taxonomic history</emphasis>
. In its only record in the literature this species was misinterpreted as
<taxonomicName id="84FB9CAE9D78CA6FFBD5FB75FA8CFA87" authority="(Mielke et al. 2010)" baseAuthorityName="Mielke et al." baseAuthorityYear="2010" box="[1086,1391,1270,1296]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="focus">
<emphasis id="718F3B3F9D78CA6FFBD5FB75FB70FA87" box="[1086,1171,1270,1296]" italics="true" pageId="34" pageNumber="35">A. focus</emphasis>
(
<bibRefCitation id="276A9ADC9D78CA6FFB4BFB75FA84FA87" author="Mielke, O. H. H. &amp; Carneiro, E. &amp; Casagrande, M. M." box="[1184,1383,1270,1296]" pageId="34" pageNumber="35" pagination="285 - 299" refId="ref43639" refString="Mielke, O. H. H., Carneiro, E. &amp; Casagrande, M. M. (2010) Lepidopterofauna (Papilionoidea e Hesperioidea) do Parque Estadual do Chandless e arredores, Acre, Brasil. Biota Neotropica, 10 (4), 285 - 299. https: // doi. org / 10.1590 / S 1676 - 06032010000400033" type="journal article" year="2010">
Mielke
<emphasis id="718F3B3F9D78CA6FFB18FB74FAC4FA87" box="[1267,1319,1270,1296]" italics="true" pageId="34" pageNumber="35">et al.</emphasis>
2010
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D78CA6FFF2CFA9AFCE0FA53" pageId="34" pageNumber="35" type="diagnosis">
<paragraph id="4344E72D9D78CA6FFF2CFA9AFCE0FA53" blockId="34.[151,1437,1269,2017]" pageId="34" pageNumber="35">
<emphasis id="718F3B3F9D78CA6FFF2CFA9AFED9FAA3" bold="true" box="[199,314,1305,1332]" pageId="34" pageNumber="35">Diagnosis</emphasis>
.
<taxonomicName id="84FB9CAE9D78CA6FFEA2FA99FE3EFAA3" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[329,477,1306,1332]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="cofus" status="sp. nov.">
<emphasis id="718F3B3F9D78CA6FFEA2FA99FE3EFAA3" box="[329,477,1306,1332]" italics="true" pageId="34" pageNumber="35">Artines cofus</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D78CA6FFE0FFA99FDDDFAA3" bold="true" box="[484,574,1306,1332]" pageId="34" pageNumber="35">
<taxonomicNameLabel id="6ABC86449D78CA6FFE0FFA99FDDDFAA3" box="[484,574,1306,1332]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from the other species of the “
<taxonomicName id="84FB9CAE9D78CA6FFC06FA99FBA9FAA3" authorityName="Plotz" authorityYear="1883" box="[1005,1098,1306,1332]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="aquilina">
<emphasis id="718F3B3F9D78CA6FFC06FA99FBA9FAA3" box="[1005,1098,1306,1332]" italics="true" pageId="34" pageNumber="35">aquilina</emphasis>
</taxonomicName>
group” in the following characters: uncus distally wide and without ventral projection; aedeagus with proximal end narrow; opening of ejaculatory bulb separated from proximal margin of aedeagus by short distance; ventral projection of aedeagus with distal portion bifid; lamella postvaginalis without proximal plate, distal plate with sclerotized median notch on proximal margin and a small median projection on distal margin.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D78CA6EFF2CFA4EFBB6FE60" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35" type="description">
<paragraph id="4344E72D9D78CA6FFF2CFA4EFB80FA7E" blockId="34.[151,1437,1269,2017]" box="[199,1123,1485,1513]" pageId="34" pageNumber="35">
<emphasis id="718F3B3F9D78CA6FFF2CFA4EFEB2FA7F" bold="true" box="[199,337,1485,1512]" pageId="34" pageNumber="35">Description</emphasis>
. Male (
<figureCitation id="DBC0FBA89D78CA6FFE43FA4DFE1BFA7F" box="[424,504,1486,1512]" captionStart="FIGURES 14–19" captionStartId="10.[151,264,1406,1431]" captionTargetBox="[155,1435,175,1386]" captionTargetId="figure@10.[151,1436,174,1394]" captionTargetPageId="10" captionText="FIGURES 14–19. Species of the “aquilina group” (14–18) and “bipunctata group” (19) in dorsal and ventral views. 14. Artines aquilina, A–B: male, Joinville, Santa Catarina, Brazil (DZ 41.321), C–D: female from same locality of male (DZ 41.220). 15. Artines angelica sp. nov., A–B: holotype male, C–D: allotype female. 16. Artines focus, A–B: male, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.233), C–D: female, Pacaraima, Roraima, Brazil (DZ 41.333). 17. Artines cofus sp. nov., A–B: holotype male, C–D: allotype female. 18. Artines donia sp. nov., A–B: holotype male. 19. Artines bipunctata, A–B: male, Jardim Zoológico, Brasília, Distrito Federal, Brazil (DZ 30.961), C–D: female, Córrego Bananal, Brasília, Distrito Federal, Brazil (DZ 41.224). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241045/files/figure.png" pageId="34" pageNumber="35">Fig. 17</figureCitation>
A–B).
<emphasis id="718F3B3F9D78CA6FFDAEFA4DFD61FA7F" box="[581,642,1486,1512]" italics="true" pageId="34" pageNumber="35">Head</emphasis>
: antenna with nudum bearing12 segments.
</paragraph>
<paragraph id="4344E72D9D78CA6FFF2CFA70FDC6F99A" blockId="34.[151,1437,1269,2017]" box="[199,549,1521,1549]" pageId="34" pageNumber="35">
<emphasis id="718F3B3F9D78CA6FFF2CFA70FE62F99B" box="[199,385,1522,1548]" italics="true" pageId="34" pageNumber="35">Forewing length</emphasis>
:
<quantity id="84034AC89D78CA6FFE7BFA72FDC2F99A" box="[400,545,1521,1549]" metricMagnitude="-2" metricUnit="m" metricValue="1.225" metricValueMax="1.3" metricValueMin="1.15" pageId="34" pageNumber="35" unit="mm" value="12.25" valueMax="13.0" valueMin="11.5">11.5–13 mm</quantity>
.
</paragraph>
<paragraph id="4344E72D9D78CA6FFF2CF995FAD1F9C3" blockId="34.[151,1437,1269,2017]" pageId="34" pageNumber="35">
<emphasis id="718F3B3F9D78CA6FFF2CF995FE9CF9A7" box="[199,383,1558,1584]" italics="true" pageId="34" pageNumber="35">Dorsal forewing</emphasis>
(
<figureCitation id="DBC0FBA89D78CA6FFE66F995FE16F9A7" box="[397,501,1558,1584]" captionStart="FIGURES 14–19" captionStartId="10.[151,264,1406,1431]" captionTargetBox="[155,1435,175,1386]" captionTargetId="figure@10.[151,1436,174,1394]" captionTargetPageId="10" captionText="FIGURES 14–19. Species of the “aquilina group” (14–18) and “bipunctata group” (19) in dorsal and ventral views. 14. Artines aquilina, A–B: male, Joinville, Santa Catarina, Brazil (DZ 41.321), C–D: female from same locality of male (DZ 41.220). 15. Artines angelica sp. nov., A–B: holotype male, C–D: allotype female. 16. Artines focus, A–B: male, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.233), C–D: female, Pacaraima, Roraima, Brazil (DZ 41.333). 17. Artines cofus sp. nov., A–B: holotype male, C–D: allotype female. 18. Artines donia sp. nov., A–B: holotype male. 19. Artines bipunctata, A–B: male, Jardim Zoológico, Brasília, Distrito Federal, Brazil (DZ 30.961), C–D: female, Córrego Bananal, Brasília, Distrito Federal, Brazil (DZ 41.224). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241045/files/figure.png" pageId="34" pageNumber="35">Fig. 17A</figureCitation>
): apical spots absent or one or two in R
<subScript id="DF7FE5689D78CA6FFC4FF9ABFC4FF9A1" attach="left" box="[932,940,1576,1590]" fontSize="6" pageId="34" pageNumber="35">3</subScript>
–R
<subScript id="DF7FE5689D78CA6FFC27F9ABFC37F9A1" attach="left" box="[972,980,1576,1590]" fontSize="6" pageId="34" pageNumber="35">4</subScript>
and R
<subScript id="DF7FE5689D78CA6FFBF0F9ABFBC0F9A1" attach="left" box="[1051,1059,1576,1590]" fontSize="6" pageId="34" pageNumber="35">4</subScript>
–R
<subScript id="DF7FE5689D78CA6FFBAFF9A4FBAFF9A2" attach="left" box="[1092,1100,1575,1589]" fontSize="6" pageId="34" pageNumber="35">5</subScript>
; discal spots reduced and spot in CuA
<subScript id="DF7FE5689D78CA6FFF02F9CFFF12F9CD" attach="left" box="[233,241,1612,1626]" fontSize="6" pageId="34" pageNumber="35">2</subScript>
–2A absent or vestigial; brand shorter, lower projection slightly longer than
<taxonomicName id="84FB9CAE9D78CA6FFBAEF9B9FB22F9C3" authorityName="Plotz" authorityYear="1883" box="[1093,1217,1594,1620]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="aquilina">
<emphasis id="718F3B3F9D78CA6FFBAEF9B9FB22F9C3" box="[1093,1217,1594,1620]" italics="true" pageId="34" pageNumber="35">A. aquilina</emphasis>
</taxonomicName>
(
<figureCitation id="DBC0FBA89D78CA6FFB3AF9B9FAC1F9C3" box="[1233,1314,1594,1620]" captionStart="FIGURES 21–26" captionStartId="22.[151,264,1840,1865]" captionTargetBox="[151,1436,922,1828]" captionTargetId="figure@22.[151,1436,922,1828]" captionText="FIGURES 21–26. Male brands (no scales) on dorsal forewing of the “aquilina group” (21–25) and “bipunctata group” species (26). 21. Artines aquilina, Joinville, Santa catarina, Brazil, (DZ 41.341). 22. Artines angelica sp. nov., paratype, Fazenda Ter- rasse, Santa Luzia, Maranhão, Brazil (DZ 41.273) 23. Artines focus, Ilha de Maracá, Roraima, Brazil (DZ 41.233). 24. Artines cofus sp. nov., paratype, Santa Rosa do Purus, Acre, Brazil (DZ 41.303). 25. Artines donia sp. nov., paratype, Santa Rita do Araguaia, Goiás, Brazil (OM 66.000). 26. Artines bipunctata, Ponte Funda, Vianópilis, Goiás, Brazil (DZ 41.264). Scale bar = 1 mm." httpUri="https://zenodo.org/record/3406270/files/figure.png" pageId="34" pageNumber="35">Fig. 24</figureCitation>
).
</paragraph>
<paragraph id="4344E72D9D78CA6FFF2CF9DDFAAAF90B" blockId="34.[151,1437,1269,2017]" pageId="34" pageNumber="35">
<emphasis id="718F3B3F9D78CA6FFF2CF9DDFE64F9EF" box="[199,391,1630,1656]" italics="true" pageId="34" pageNumber="35">Ventral forewing</emphasis>
(
<figureCitation id="DBC0FBA89D78CA6FFE7DF9DDFE1DF9EF" box="[406,510,1630,1656]" captionStart="FIGURES 14–19" captionStartId="10.[151,264,1406,1431]" captionTargetBox="[155,1435,175,1386]" captionTargetId="figure@10.[151,1436,174,1394]" captionTargetPageId="10" captionText="FIGURES 14–19. Species of the “aquilina group” (14–18) and “bipunctata group” (19) in dorsal and ventral views. 14. Artines aquilina, A–B: male, Joinville, Santa Catarina, Brazil (DZ 41.321), C–D: female from same locality of male (DZ 41.220). 15. Artines angelica sp. nov., A–B: holotype male, C–D: allotype female. 16. Artines focus, A–B: male, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.233), C–D: female, Pacaraima, Roraima, Brazil (DZ 41.333). 17. Artines cofus sp. nov., A–B: holotype male, C–D: allotype female. 18. Artines donia sp. nov., A–B: holotype male. 19. Artines bipunctata, A–B: male, Jardim Zoológico, Brasília, Distrito Federal, Brazil (DZ 30.961), C–D: female, Córrego Bananal, Brasília, Distrito Federal, Brazil (DZ 41.224). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241045/files/figure.png" pageId="34" pageNumber="35">Fig. 17B</figureCitation>
): three to four reduced apical spots; discal spots as on dorsal surface; submarginal band conspicuous and almost discontinued at the middle of cells; weak spot white and diffuse in CuA
<subScript id="DF7FE5689D78CA6FFAE1F917FAF1F935" attach="left" box="[1290,1298,1684,1698]" fontSize="6" pageId="34" pageNumber="35">2</subScript>
–2A.
</paragraph>
<paragraph id="4344E72D9D78CA6FFF2CF925FB2BF973" blockId="34.[151,1437,1269,2017]" pageId="34" pageNumber="35">
<emphasis id="718F3B3F9D78CA6FFF2CF925FE72F957" box="[199,401,1702,1728]" italics="true" pageId="34" pageNumber="35">Ventral hind wing</emphasis>
(
<figureCitation id="DBC0FBA89D78CA6FFE4AF925FDEBF957" box="[417,520,1702,1728]" captionStart="FIGURES 14–19" captionStartId="10.[151,264,1406,1431]" captionTargetBox="[155,1435,175,1386]" captionTargetId="figure@10.[151,1436,174,1394]" captionTargetPageId="10" captionText="FIGURES 14–19. Species of the “aquilina group” (14–18) and “bipunctata group” (19) in dorsal and ventral views. 14. Artines aquilina, A–B: male, Joinville, Santa Catarina, Brazil (DZ 41.321), C–D: female from same locality of male (DZ 41.220). 15. Artines angelica sp. nov., A–B: holotype male, C–D: allotype female. 16. Artines focus, A–B: male, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.233), C–D: female, Pacaraima, Roraima, Brazil (DZ 41.333). 17. Artines cofus sp. nov., A–B: holotype male, C–D: allotype female. 18. Artines donia sp. nov., A–B: holotype male. 19. Artines bipunctata, A–B: male, Jardim Zoológico, Brasília, Distrito Federal, Brazil (DZ 30.961), C–D: female, Córrego Bananal, Brasília, Distrito Federal, Brazil (DZ 41.224). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241045/files/figure.png" pageId="34" pageNumber="35">Fig. 17B</figureCitation>
): apical portion of discal cell densely covered by yellow scales; central black spot large, as wide as long; postdiscal band yellow, limited distally by a not wavy dark brown stripe.
</paragraph>
<paragraph id="4344E72D9D78CA6EFF2CF96DFAF9FED4" blockId="34.[151,1437,1269,2017]" lastBlockId="35.[151,1437,153,575]" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
<emphasis id="718F3B3F9D78CA6FFF2CF96DFED2F89F" box="[199,305,1774,1800]" italics="true" pageId="34" pageNumber="35">Genitalia</emphasis>
(
<figureCitation id="DBC0FBA89D78CA6FFEAAF96DFE74F89F" box="[321,407,1774,1800]" captionStart="FIGURES 38–39" captionStartId="29.[151,264,1720,1745]" captionTargetBox="[161,1436,431,1681]" captionTargetId="figure@29.[151,1436,450,1693]" captionTargetPageId="29" captionText="FIGURES 38–39. Male genitalia of the “aquilina group” species. 38. Artines focus, Reserva Duque, Manaus, Amazonas, Brazil (DZ 41.192). 39. Artines cofus sp. nov., paratype, Tambopata, Puerto Maldonado Madre de Díos, Peru (MUSM). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241055/files/figure.png" pageId="34" pageNumber="35">Figs 39</figureCitation>
K–M, 52): tegumen more or less quadrate, distal margin convex (
<figureCitation id="DBC0FBA89D78CA6FFB95F96DFB06F89F" box="[1150,1253,1774,1800]" captionStart="FIGURES 38–39" captionStartId="29.[151,264,1720,1745]" captionTargetBox="[161,1436,431,1681]" captionTargetId="figure@29.[151,1436,450,1693]" captionTargetPageId="29" captionText="FIGURES 38–39. Male genitalia of the “aquilina group” species. 38. Artines focus, Reserva Duque, Manaus, Amazonas, Brazil (DZ 41.192). 39. Artines cofus sp. nov., paratype, Tambopata, Puerto Maldonado Madre de Díos, Peru (MUSM). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241055/files/figure.png" pageId="34" pageNumber="35">Fig. 39C</figureCitation>
), in lateral view rectangular, dorsal margin slightly convex (
<figureCitation id="DBC0FBA89D78CA6FFD96F891FD05F8BB" box="[637,742,1810,1836]" captionStart="FIGURES 38–39" captionStartId="29.[151,264,1720,1745]" captionTargetBox="[161,1436,431,1681]" captionTargetId="figure@29.[151,1436,450,1693]" captionTargetPageId="29" captionText="FIGURES 38–39. Male genitalia of the “aquilina group” species. 38. Artines focus, Reserva Duque, Manaus, Amazonas, Brazil (DZ 41.192). 39. Artines cofus sp. nov., paratype, Tambopata, Puerto Maldonado Madre de Díos, Peru (MUSM). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241055/files/figure.png" pageId="34" pageNumber="35">Fig. 39A</figureCitation>
). Saccus narrow medially, anterior portion slightly dilated and narrower than tegumen, proximal margin rounded (
<figureCitation id="DBC0FBA89D78CA6FFD24F8B5FCD6F8C7" box="[719,821,1846,1872]" captionStart="FIGURES 38–39" captionStartId="29.[151,264,1720,1745]" captionTargetBox="[161,1436,431,1681]" captionTargetId="figure@29.[151,1436,450,1693]" captionTargetPageId="29" captionText="FIGURES 38–39. Male genitalia of the “aquilina group” species. 38. Artines focus, Reserva Duque, Manaus, Amazonas, Brazil (DZ 41.192). 39. Artines cofus sp. nov., paratype, Tambopata, Puerto Maldonado Madre de Díos, Peru (MUSM). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241055/files/figure.png" pageId="34" pageNumber="35">Fig. 39B</figureCitation>
). Uncus shorter than tegumen, proximal 1/2 with lateral margins convex, distal 1/2 narrow, lateral margins concave, distal portion rounded (
<figureCitation id="DBC0FBA89D78CA6FFBDFF8D9FB7AF8E3" box="[1076,1177,1882,1908]" captionStart="FIGURES 38–39" captionStartId="29.[151,264,1720,1745]" captionTargetBox="[161,1436,431,1681]" captionTargetId="figure@29.[151,1436,450,1693]" captionTargetPageId="29" captionText="FIGURES 38–39. Male genitalia of the “aquilina group” species. 38. Artines focus, Reserva Duque, Manaus, Amazonas, Brazil (DZ 41.192). 39. Artines cofus sp. nov., paratype, Tambopata, Puerto Maldonado Madre de Díos, Peru (MUSM). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241055/files/figure.png" pageId="34" pageNumber="35">Fig. 39C</figureCitation>
), apex curved ventrally (
<figureCitation id="DBC0FBA89D78CA6FFF4BF8FDFEE8F80F" box="[160,267,1918,1944]" captionStart="FIGURES 38–39" captionStartId="29.[151,264,1720,1745]" captionTargetBox="[161,1436,431,1681]" captionTargetId="figure@29.[151,1436,450,1693]" captionTargetPageId="29" captionText="FIGURES 38–39. Male genitalia of the “aquilina group” species. 38. Artines focus, Reserva Duque, Manaus, Amazonas, Brazil (DZ 41.192). 39. Artines cofus sp. nov., paratype, Tambopata, Puerto Maldonado Madre de Díos, Peru (MUSM). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241055/files/figure.png" pageId="34" pageNumber="35">Fig. 39A</figureCitation>
). Gnathos reaching 1/2 length of uncus, arms parallel, slightly divergent on apex in ventral view (
<figureCitation id="DBC0FBA89D78CA6FFA84F8FDFF28F82B" captionStart="FIGURES 38–39" captionStartId="29.[151,264,1720,1745]" captionTargetBox="[161,1436,431,1681]" captionTargetId="figure@29.[151,1436,450,1693]" captionTargetPageId="29" captionText="FIGURES 38–39. Male genitalia of the “aquilina group” species. 38. Artines focus, Reserva Duque, Manaus, Amazonas, Brazil (DZ 41.192). 39. Artines cofus sp. nov., paratype, Tambopata, Puerto Maldonado Madre de Díos, Peru (MUSM). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241055/files/figure.png" pageId="34" pageNumber="35">Fig. 39D</figureCitation>
), curved upwards toward distal end of uncus (
<figureCitation id="DBC0FBA89D78CA6FFD21F821FCD1F82B" box="[714,818,1954,1980]" captionStart="FIGURES 38–39" captionStartId="29.[151,264,1720,1745]" captionTargetBox="[161,1436,431,1681]" captionTargetId="figure@29.[151,1436,450,1693]" captionTargetPageId="29" captionText="FIGURES 38–39. Male genitalia of the “aquilina group” species. 38. Artines focus, Reserva Duque, Manaus, Amazonas, Brazil (DZ 41.192). 39. Artines cofus sp. nov., paratype, Tambopata, Puerto Maldonado Madre de Díos, Peru (MUSM). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241055/files/figure.png" pageId="34" pageNumber="35">Fig. 39A</figureCitation>
). Valva two and a half times longer than wide; sacculus short and triangular separated from harpe by long distance; costa narrow, triangular, short, projected inwards and separated from ampulla by distance longer than its length; harpe longer than 1/2 of valva, uniformly narrow; ampulla 1/2 width of harpe, not protruding; distal projection of harpe, in dorsal view, weakly curved inwards and with small distal spines (
<figureCitation id="DBC0FBA89D79CA6EFF03FF62FED7FF6C" box="[232,308,225,251]" captionStart="FIGURES 38–39" captionStartId="29.[151,264,1720,1745]" captionTargetBox="[161,1436,431,1681]" captionTargetId="figure@29.[151,1436,450,1693]" captionTargetPageId="29" captionText="FIGURES 38–39. Male genitalia of the “aquilina group” species. 38. Artines focus, Reserva Duque, Manaus, Amazonas, Brazil (DZ 41.192). 39. Artines cofus sp. nov., paratype, Tambopata, Puerto Maldonado Madre de Díos, Peru (MUSM). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241055/files/figure.png" pageId="35" pageNumber="36">Fig. 39</figureCitation>
E–F). Aedeagus as in
<emphasis id="718F3B3F9D79CA6EFDF5FF62FD99FF6C" box="[542,634,225,251]" italics="true" pageId="35" pageNumber="36">
<taxonomicName id="84FB9CAE9D79CA6EFDF5FF62FD95FF6C" box="[542,630,225,251]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="focus">A. focus</taxonomicName>
,
</emphasis>
differing in the opening of the ejaculatory bulb more distant from proximal margin; ventral projection with distal lobes asymmetrical (
<figureCitation id="DBC0FBA89D79CA6EFCE9FE86FCADFE88" box="[770,846,261,287]" captionStart="FIGURES 38–39" captionStartId="29.[151,264,1720,1745]" captionTargetBox="[161,1436,431,1681]" captionTargetId="figure@29.[151,1436,450,1693]" captionTargetPageId="29" captionText="FIGURES 38–39. Male genitalia of the “aquilina group” species. 38. Artines focus, Reserva Duque, Manaus, Amazonas, Brazil (DZ 41.192). 39. Artines cofus sp. nov., paratype, Tambopata, Puerto Maldonado Madre de Díos, Peru (MUSM). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241055/files/figure.png" pageId="35" pageNumber="36">Fig. 39</figureCitation>
G–J). Cornutus similar to
<taxonomicName id="84FB9CAE9D79CA6EFB8AFE86FB55FE88" box="[1121,1206,261,287]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="focus">
<emphasis id="718F3B3F9D79CA6EFB8AFE86FB55FE88" box="[1121,1206,261,287]" italics="true" pageId="35" pageNumber="36">A. focus</emphasis>
</taxonomicName>
, however, spines less conspicuous and less numerous (
<figureCitation id="DBC0FBA89D79CA6EFE1AFEAAFDB9FED4" box="[497,602,297,323]" captionStart="FIGURES 38–39" captionStartId="29.[151,264,1720,1745]" captionTargetBox="[161,1436,431,1681]" captionTargetId="figure@29.[151,1436,450,1693]" captionTargetPageId="29" captionText="FIGURES 38–39. Male genitalia of the “aquilina group” species. 38. Artines focus, Reserva Duque, Manaus, Amazonas, Brazil (DZ 41.192). 39. Artines cofus sp. nov., paratype, Tambopata, Puerto Maldonado Madre de Díos, Peru (MUSM). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241055/files/figure.png" pageId="35" pageNumber="36">Figs 39M</figureCitation>
,
<figureCitation id="DBC0FBA89D79CA6EFD8EFEABFD60FED4" box="[613,643,296,323]" captionStart="FIGURES 49–53" captionStartId="34.[151,264,895,920]" captionTargetBox="[155,1421,182,878]" captionTargetId="figure@34.[151,1436,174,883]" captionTargetPageId="34" captionText="FIGURES 49–53. Ventral view of vesica and cornutus of the “aquilina group” species. 49. Artines aquilina. 50. Artines angelica sp. nov.. 51. Artines focus. 52. Artines cofus sp. nov.. 53. Artines donia sp. nov.. Scale bar = 0,2 mm." httpUri="https://zenodo.org/record/3241057/files/figure.png" pageId="35" pageNumber="36">52</figureCitation>
). Fultura inferior, in lateral view, uniformly narrow (
<figureCitation id="DBC0FBA89D79CA6EFB45FEAAFAEEFED4" box="[1198,1293,297,323]" captionStart="FIGURES 38–39" captionStartId="29.[151,264,1720,1745]" captionTargetBox="[161,1436,431,1681]" captionTargetId="figure@29.[151,1436,450,1693]" captionTargetPageId="29" captionText="FIGURES 38–39. Male genitalia of the “aquilina group” species. 38. Artines focus, Reserva Duque, Manaus, Amazonas, Brazil (DZ 41.192). 39. Artines cofus sp. nov., paratype, Tambopata, Puerto Maldonado Madre de Díos, Peru (MUSM). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241055/files/figure.png" pageId="35" pageNumber="36">Fig. 39L</figureCitation>
).
</paragraph>
<paragraph id="4344E72D9D79CA6EFF2CFECEFC7CFEF0" blockId="35.[151,1437,153,575]" box="[199,927,333,359]" pageId="35" pageNumber="36">
Female (
<figureCitation id="DBC0FBA89D79CA6EFEC2FECEFE9BFEF0" box="[297,376,333,359]" captionStart="FIGURES 14–19" captionStartId="10.[151,264,1406,1431]" captionTargetBox="[155,1435,175,1386]" captionTargetId="figure@10.[151,1436,174,1394]" captionTargetPageId="10" captionText="FIGURES 14–19. Species of the “aquilina group” (14–18) and “bipunctata group” (19) in dorsal and ventral views. 14. Artines aquilina, A–B: male, Joinville, Santa Catarina, Brazil (DZ 41.321), C–D: female from same locality of male (DZ 41.220). 15. Artines angelica sp. nov., A–B: holotype male, C–D: allotype female. 16. Artines focus, A–B: male, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.233), C–D: female, Pacaraima, Roraima, Brazil (DZ 41.333). 17. Artines cofus sp. nov., A–B: holotype male, C–D: allotype female. 18. Artines donia sp. nov., A–B: holotype male. 19. Artines bipunctata, A–B: male, Jardim Zoológico, Brasília, Distrito Federal, Brazil (DZ 30.961), C–D: female, Córrego Bananal, Brasília, Distrito Federal, Brazil (DZ 41.224). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241045/files/figure.png" pageId="35" pageNumber="36">Fig. 17</figureCitation>
C–D).
<emphasis id="718F3B3F9D79CA6EFE2EFECDFD9CFEF0" box="[453,639,333,359]" italics="true" pageId="35" pageNumber="36">Forewing length</emphasis>
:
<quantity id="84034AC89D79CA6EFD65FECEFD03FEF0" box="[654,736,333,359]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="35" pageNumber="36" unit="mm" value="12.0">12 mm</quantity>
. Similar to male.
</paragraph>
<paragraph id="4344E72D9D79CA6EFF2CFEF2FD54FE44" blockId="35.[151,1437,153,575]" pageId="35" pageNumber="36">
<emphasis id="718F3B3F9D79CA6EFF2CFEF2FED2FE1C" box="[199,305,369,395]" italics="true" pageId="35" pageNumber="36">Genitalia</emphasis>
(
<figureCitation id="DBC0FBA89D79CA6EFEABFEF2FE75FE1C" box="[320,406,369,395]" captionStart="FIGURES 61–65" captionStartId="36.[151,264,653,678]" captionTargetBox="[151,1435,178,637]" captionTargetId="figure@36.[151,1436,174,642]" captionTargetPageId="36" captionText="FIGURES 61–65. Female genitalia of the “aquilina group” and “bipunctata group” species. 61. Artines aquilina, Cananéia, São Paulo, Brazil (OM 49.831). 62. Artines angelica sp. nov., paratype, Tambopata, Puerto Maldonado, Madre de Díos, Peru (DZ 41.222) 63. Artines focus, Pacaraima, Roraima, Brazil (DZ 41.333). 64. Artines cofus sp. nov. paratype, Tambopata, Puerto Maldonado, Madre de Díos, Peru (DZ 41. 293). 65. Artines bipunctata, Reserva IBGE, Brasília, Distrito Federal, Brazil (DZ 41.165). For reference of the letters see legend of the figure 54. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241061/files/figure.png" pageId="35" pageNumber="36">Figs 64</figureCitation>
A–B, 76): lamella antevaginalis narrow; lamella postvaginalis without proximal plate, distal plate with sclerotized median notch on proximal margin, distal margin with small median projection. Ostium on proximal 1/2 do sterigma; ductus bursae narrow.
</paragraph>
<paragraph id="4344E72D9D79CA6EFF2CFE5FFBB6FE60" blockId="35.[151,1437,153,575]" box="[199,1109,476,503]" pageId="35" pageNumber="36">
<emphasis id="718F3B3F9D79CA6EFF2CFE5FFED9FE60" bold="true" box="[199,314,476,503]" pageId="35" pageNumber="36">Variation</emphasis>
. The central black spot on ventral hind wing is sometimes rectangular.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D79CA69FF2CFD83FF3CFC51" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36" type="discussion">
<paragraph id="4344E72D9D79CA6EFF2CFD83FC1CFDA8" blockId="35.[151,1437,153,575]" pageId="35" pageNumber="36">
<emphasis id="718F3B3F9D79CA6EFF2CFD83FEABFD8C" bold="true" box="[199,328,512,539]" pageId="35" pageNumber="36">Comments</emphasis>
. Wing pattern, male and female genitalia as in
<emphasis id="718F3B3F9D79CA6EFCBCFD82FC5BFD8C" box="[855,952,513,539]" italics="true" pageId="35" pageNumber="36">
<taxonomicName id="84FB9CAE9D79CA6EFCBCFD82FC57FD8C" box="[855,948,513,539]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="focus">A. focus</taxonomicName>
,
</emphasis>
differing in the smaller size, reduced white spots on forewing, absence of ventral projection of uncus and narrower valva.
</paragraph>
<caption id="1784B7A59D79CA6EFF7CF88CFC37F84C" httpUri="https://zenodo.org/record/3241059/files/figure.png" pageId="35" pageNumber="36" startId="35.[151,264,1807,1832]" targetBox="[156,1436,575,1772]" targetPageId="35">
<paragraph id="4344E72D9D79CA6EFF7CF88CFC37F84C" blockId="35.[151,1437,1807,2012]" pageId="35" pageNumber="36">
<emphasis id="718F3B3F9D79CA6EFF7CF88CFEB3F8BF" bold="true" box="[151,336,1807,1832]" pageId="35" pageNumber="36">FIGURES 54–60</emphasis>
. Female genitalia of the “
<taxonomicName id="84FB9CAE9D79CA6EFD8EF892FD4DF8BF" box="[613,686,1809,1832]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D79CA6EFD8EF892FD4DF8BF" box="[613,686,1809,1832]" italics="true" pageId="35" pageNumber="36">aepitus</emphasis>
</taxonomicName>
group” species.
<emphasis id="718F3B3F9D79CA6EFCB4F88CFC9CF8BF" bold="true" box="[863,895,1807,1832]" pageId="35" pageNumber="36">54.</emphasis>
<taxonomicName id="84FB9CAE9D79CA6EFC63F893FB61F8BF" authority=", Jusepin" authorityName="Jusepin" box="[904,1154,1808,1832]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D79CA6EFC63F893FBC0F8BF" box="[904,1059,1808,1832]" italics="true" pageId="35" pageNumber="36">Artines aepitus</emphasis>
, Jusepín
</taxonomicName>
, Monagas, Venezuela (OM 30. 125).
<emphasis id="718F3B3F9D79CA6EFF10F8B7FEF8F8DB" bold="true" box="[251,283,1844,1868]" pageId="35" pageNumber="36">55.</emphasis>
<taxonomicName id="84FB9CAE9D79CA6EFEC9F8B7FE26F8DB" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[290,453,1844,1868]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="panama" status="sp. nov.">
<emphasis id="718F3B3F9D79CA6EFEC9F8B7FE26F8DB" box="[290,453,1844,1868]" italics="true" pageId="35" pageNumber="36">Artines panama</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D79CA6EFE27F8B7FDFDF8DB" bold="true" box="[460,542,1844,1868]" pageId="35" pageNumber="36">
<taxonomicNameLabel id="6ABC86449D79CA6EFE27F8B7FDFDF8DB" box="[460,542,1844,1868]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, allotype.
<emphasis id="718F3B3F9D79CA6EFD67F8B0FD4FF8DB" bold="true" box="[652,684,1843,1868]" pageId="35" pageNumber="36">56.</emphasis>
<taxonomicName id="84FB9CAE9D79CA6EFD58F8B7FCB0F8DB" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[691,851,1844,1868]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="solange" status="sp. nov.">
<emphasis id="718F3B3F9D79CA6EFD58F8B7FCB0F8DB" box="[691,851,1844,1868]" italics="true" pageId="35" pageNumber="36">Artines solange</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D79CA6EFCB1F8B7FC4FF8DB" bold="true" box="[858,940,1844,1868]" pageId="35" pageNumber="36">
<taxonomicNameLabel id="6ABC86449D79CA6EFCB1F8B7FC4FF8DB" box="[858,940,1844,1868]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(tentative female), Sumaúma, Maranhão, Brazil (DZ 41.800).
<emphasis id="718F3B3F9D79CA6EFECFF8DBFEA7F8E7" bold="true" box="[292,324,1880,1904]" pageId="35" pageNumber="36">57.</emphasis>
<taxonomicName id="84FB9CAE9D79CA6EFEA0F8DBFE08F8E7" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[331,491,1880,1904]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="litoralis" status="sp. nov.">
<emphasis id="718F3B3F9D79CA6EFEA0F8DBFE08F8E7" box="[331,491,1880,1904]" italics="true" pageId="35" pageNumber="36">Artines litoralis</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D79CA6EFE19F8DBFDA0F8E7" bold="true" box="[498,579,1880,1904]" pageId="35" pageNumber="36">
<taxonomicNameLabel id="6ABC86449D79CA6EFE19F8DBFDA0F8E7" box="[498,579,1880,1904]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype, Linhares, Espírito Santo, Brazil (DZ 41.820).
<emphasis id="718F3B3F9D79CA6EFB67F8D4FB4FF8E7" bold="true" box="[1164,1196,1879,1904]" pageId="35" pageNumber="36">58.</emphasis>
<taxonomicName id="84FB9CAE9D79CA6EFB58F8DBFA7AF8E7" authority=", Turrialba" authorityName="Turrialba" box="[1203,1433,1879,1904]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="rica">
<emphasis id="718F3B3F9D79CA6EFB58F8DBFACFF8E7" box="[1203,1324,1880,1904]" italics="true" pageId="35" pageNumber="36">Artines rica</emphasis>
, Turrialba
</taxonomicName>
, Cartago, Costa Rica (DZ 41.840).
<emphasis id="718F3B3F9D79CA6EFE13F8F8FDFBF803" bold="true" box="[504,536,1915,1940]" pageId="35" pageNumber="36">59.</emphasis>
<taxonomicName id="84FB9CAE9D79CA6EFDF4F8FFFD43F803" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[543,672,1916,1940]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="liege" status="sp. nov.">
<emphasis id="718F3B3F9D79CA6EFDF4F8FFFD43F803" box="[543,672,1916,1940]" italics="true" pageId="35" pageNumber="36">Artines liege</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D79CA6EFD4CF8FFFD1AF803" bold="true" box="[679,761,1916,1940]" pageId="35" pageNumber="36">
<taxonomicNameLabel id="6ABC86449D79CA6EFD4CF8FFFD1AF803" box="[679,761,1916,1940]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype, Parque do Gama, Brasília, Distrito Federal, Brazil (DZ 41.830).
<emphasis id="718F3B3F9D79CA6EFF19F81CFEF1F82F" bold="true" box="[242,274,1951,1976]" pageId="35" pageNumber="36">60.</emphasis>
<taxonomicName id="84FB9CAE9D79CA6EFEF3F823FE4FF82F" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[280,428,1952,1976]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="bamba" status="sp. nov.">
<emphasis id="718F3B3F9D79CA6EFEF3F823FE4FF82F" box="[280,428,1952,1976]" italics="true" pageId="35" pageNumber="36">Artines bamba</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D79CA6EFE59F823FDE6F82F" bold="true" box="[434,517,1952,1976]" pageId="35" pageNumber="36">
<taxonomicNameLabel id="6ABC86449D79CA6EFE59F823FDE6F82F" box="[434,517,1952,1976]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(tentative female), Moyobamba, San Martin, Peru (OM 81.229).
<emphasis id="718F3B3F9D79CA6EFB70F81CFB4EF82F" bold="true" box="[1179,1197,1951,1976]" pageId="35" pageNumber="36">A</emphasis>
) lateral view of tergum VIII, sterigma and papilla analis;
<emphasis id="718F3B3F9D79CA6EFE07F847FE1EF84B" bold="true" box="[492,509,1988,2012]" pageId="35" pageNumber="36">B</emphasis>
) ventral view of esterigma. Scale bar = 1 mm.
</paragraph>
</caption>
<caption id="1784B7A59D7ECA69FF7CFD0EFC3AFCA2" httpUri="https://zenodo.org/record/3241061/files/figure.png" pageId="36" pageNumber="37" startId="36.[151,264,653,678]" targetBox="[151,1435,178,637]" targetPageId="36">
<paragraph id="4344E72D9D7ECA69FF7CFD0EFC3AFCA2" blockId="36.[151,1437,653,822]" pageId="36" pageNumber="37">
<emphasis id="718F3B3F9D7ECA69FF7CFD0EFEACFD31" bold="true" box="[151,335,653,678]" pageId="36" pageNumber="37">FIGURES 61–65</emphasis>
. Female genitalia of the “
<taxonomicName id="84FB9CAE9D7ECA69FDB4FD0DFD50FD31" authorityName="Plotz" authorityYear="1883" box="[607,691,654,678]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="aquilina">
<emphasis id="718F3B3F9D7ECA69FDB4FD0DFD50FD31" box="[607,691,654,678]" italics="true" pageId="36" pageNumber="37">aquilina</emphasis>
</taxonomicName>
group” and “
<taxonomicName id="84FB9CAE9D7ECA69FCA9FD0DFC4DFD31" authorityName="O. Mielke" authorityYear="1968" box="[834,942,654,678]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="bipunctata">
<emphasis id="718F3B3F9D7ECA69FCA9FD0DFC4DFD31" box="[834,942,654,678]" italics="true" pageId="36" pageNumber="37">bipunctata</emphasis>
</taxonomicName>
group” species.
<emphasis id="718F3B3F9D7ECA69FBB7FD0EFB9FFD31" bold="true" box="[1116,1148,653,678]" pageId="36" pageNumber="37">61.</emphasis>
<taxonomicName id="84FB9CAE9D7ECA69FB6FFD0DFA7AFD31" authority=", Cananeia" authorityName="Cananeia" box="[1156,1433,653,678]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="aquilina">
<emphasis id="718F3B3F9D7ECA69FB6FFD0DFACAFD31" box="[1156,1321,654,678]" italics="true" pageId="36" pageNumber="37">Artines aquilina</emphasis>
, Cananéia
</taxonomicName>
, São Paulo, Brazil (OM 49.831).
<emphasis id="718F3B3F9D7ECA69FE0EFD32FDE6FD5D" bold="true" box="[485,517,689,714]" pageId="36" pageNumber="37">62.</emphasis>
<taxonomicName id="84FB9CAE9D7ECA69FDE7FD31FD57FD5D" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[524,692,690,714]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="angelica" status="sp. nov.">
<emphasis id="718F3B3F9D7ECA69FDE7FD31FD57FD5D" box="[524,692,690,714]" italics="true" pageId="36" pageNumber="37">Artines angelica</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D7ECA69FD50FD31FCEEFD5D" bold="true" box="[699,781,690,714]" pageId="36" pageNumber="37">
<taxonomicNameLabel id="6ABC86449D7ECA69FD50FD31FCEEFD5D" box="[699,781,690,714]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype, Tambopata, Puerto Maldonado, Madre de Díos, Peru (DZ 41.222)
<emphasis id="718F3B3F9D7ECA69FEF0FD56FED8FD79" bold="true" box="[283,315,725,750]" pageId="36" pageNumber="37">63.</emphasis>
<taxonomicName id="84FB9CAE9D7ECA69FEABFD55FDDFFD79" authority=", Pacaraima" authorityName="Pacaraima" box="[320,572,725,750]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="focus">
<emphasis id="718F3B3F9D7ECA69FEABFD55FE28FD79" box="[320,459,726,750]" italics="true" pageId="36" pageNumber="37">Artines focus,</emphasis>
Pacaraima
</taxonomicName>
, Roraima, Brazil (DZ 41.333).
<emphasis id="718F3B3F9D7ECA69FC9DFD56FC75FD79" bold="true" box="[886,918,725,750]" pageId="36" pageNumber="37">64.</emphasis>
<taxonomicName id="84FB9CAE9D7ECA69FC77FD55FBFCFD79" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[924,1055,726,750]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="cofus" status="sp. nov.">
<emphasis id="718F3B3F9D7ECA69FC77FD55FBFCFD79" box="[924,1055,726,750]" italics="true" pageId="36" pageNumber="37">Artines cofus</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D7ECA69FBCEFD55FB96FD79" bold="true" box="[1061,1141,726,750]" pageId="36" pageNumber="37">
<taxonomicNameLabel id="6ABC86449D7ECA69FBCEFD55FB96FD79" box="[1061,1141,726,750]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
paratype, Tambopata, Puerto Maldonado, Madre de Díos, Peru (DZ 41. 293).
<emphasis id="718F3B3F9D7ECA69FD67FD7AFD48FC85" bold="true" box="[652,683,761,786]" pageId="36" pageNumber="37">65.</emphasis>
<taxonomicName id="84FB9CAE9D7ECA69FD58FD79FC33FC86" authority=", Reserva" authorityName="Reserva" box="[691,976,762,786]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="bipunctata">
<emphasis id="718F3B3F9D7ECA69FD58FD79FD1FFC86" box="[691,764,762,785]" italics="true" pageId="36" pageNumber="37">Artines</emphasis>
<emphasis id="718F3B3F9D7ECA69FCEFFD79FC94FC85" box="[772,887,762,786]" italics="true" pageId="36" pageNumber="37">bipunctata,</emphasis>
Reserva
</taxonomicName>
IBGE, Brasília, Distrito Federal, Brazil (DZ 41.165). For reference of the letters see legend of the figure 54. Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="4344E72D9D7ECA69FF2CFCE0FF3CFC51" blockId="36.[151,1436,867,1399]" pageId="36" pageNumber="37">
<emphasis id="718F3B3F9D7ECA69FF2CFCE0FE0BFCE9" bold="true" box="[199,488,867,894]" pageId="36" pageNumber="37">Geographic distribution</emphasis>
(
<figureCitation id="DBC0FBA89D7ECA69FE1CFCE7FDAFFCE9" box="[503,588,868,894]" captionStart="FIGURE 80" captionStartId="43.[151,250,930,955]" captionTargetBox="[151,1436,187,905]" captionTargetId="figure@43.[151,1436,187,905]" captionTargetPageId="43" captionText="FIGURE 80. Geographic distribution of the “aquilina group” and “bipunctata group” species. The symbols with white inferior half represent records from literature." httpUri="https://zenodo.org/record/3241071/files/figure.png" pageId="36" pageNumber="37">Fig. 80</figureCitation>
). This species occurs in the western Amazon Rainforest. PERU—
<emphasis id="718F3B3F9D7ECA69FADAFCE7FF2FFC34" italics="true" pageId="36" pageNumber="37">
<collectingRegion id="813F29CF9D7ECA69FADAFCE7FF2FFC34" country="Peru" name="Madre de Dios" pageId="36" pageNumber="37">Madre de Díos</collectingRegion>
</emphasis>
: Puerto Maldonado.
<emphasis id="718F3B3F9D7ECA69FE50FC0AFDD8FC35" box="[443,571,905,931]" italics="true" pageId="36" pageNumber="37">
<collectingRegion id="813F29CF9D7ECA69FE50FC0AFDD8FC35" box="[443,571,905,931]" country="Peru" name="San Martin" pageId="36" pageNumber="37">San Martin</collectingRegion>
</emphasis>
: Sacanche. BRAZIL—
<emphasis id="718F3B3F9D7ECA69FCAFFC0AFC9BFC34" box="[836,888,905,931]" italics="true" pageId="36" pageNumber="37">
<collectingRegion id="813F29CF9D7ECA69FCAFFC0AFC9BFC34" box="[836,888,905,931]" country="Brazil" name="Acre" pageId="36" pageNumber="37">Acre</collectingRegion>
</emphasis>
:
<collectingRegion id="813F29CF9D7ECA69FC6CFC0BFBE6FC34" box="[903,1029,904,931]" country="Guatemala" name="Santa Rosa" pageId="36" pageNumber="37">Santa Rosa</collectingRegion>
do Purus.
<emphasis id="718F3B3F9D7ECA69FB94FC0BFB0FFC35" box="[1151,1260,904,930]" italics="true" pageId="36" pageNumber="37">
<collectingRegion id="813F29CF9D7ECA69FB94FC0BFB0FFC35" box="[1151,1260,904,930]" country="Brazil" name="Rondonia" pageId="36" pageNumber="37">Rondônia</collectingRegion>
</emphasis>
: Ouro Preto do Oeste.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D7ECA69FF2CFC53FBEDFC7D" box="[199,1038,975,1003]" pageId="36" pageNumber="37" type="description">
<paragraph id="4344E72D9D7ECA69FF2CFC53FBEDFC7D" blockId="36.[151,1436,867,1399]" box="[199,1038,975,1003]" pageId="36" pageNumber="37">
<emphasis id="718F3B3F9D7ECA69FF2CFC53FE31FC7D" bold="true" box="[199,466,975,1002]" pageId="36" pageNumber="37">Temporal distribution</emphasis>
. This species occurs between August and October.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D7ECA69FF2CFC77FE2AFB99" box="[199,457,1012,1039]" pageId="36" pageNumber="37" type="biology_ecology">
<paragraph id="4344E72D9D7ECA69FF2CFC77FE2AFB99" blockId="36.[151,1436,867,1399]" box="[199,457,1012,1039]" pageId="36" pageNumber="37">
<emphasis id="718F3B3F9D7ECA69FF2CFC77FEA7FB99" bold="true" box="[199,324,1012,1038]" pageId="36" pageNumber="37">Host plant</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D7ECA69FF2CFB9BFB6FFBA4" box="[199,1164,1048,1075]" pageId="36" pageNumber="37" type="etymology">
<paragraph id="4344E72D9D7ECA69FF2CFB9BFB6FFBA4" blockId="36.[151,1436,867,1399]" box="[199,1164,1048,1075]" pageId="36" pageNumber="37">
<emphasis id="718F3B3F9D7ECA69FF2CFB9BFEABFBA5" bold="true" box="[199,328,1048,1074]" pageId="36" pageNumber="37">Etymology</emphasis>
. The name is an anagram of
<emphasis id="718F3B3F9D7ECA69FD66FB9BFD2CFBA5" box="[653,719,1048,1074]" italics="true" pageId="36" pageNumber="37">
<taxonomicName id="84FB9CAE9D7ECA69FD66FB9BFD28FBA5" box="[653,715,1048,1074]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="focus">focus</taxonomicName>
,
</emphasis>
the closest species of
<taxonomicName id="84FB9CAE9D7ECA69FC22FB9AFBC0FBA5" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[969,1059,1048,1075]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="cofus" status="sp. nov.">
<emphasis id="718F3B3F9D7ECA69FC22FB9AFBC0FBA5" box="[969,1059,1048,1075]" italics="true" pageId="36" pageNumber="37">A. cofus</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D7ECA69FBC1FB9BFB66FBA5" bold="true" box="[1066,1157,1048,1074]" pageId="36" pageNumber="37">
<taxonomicNameLabel id="6ABC86449D7ECA69FBC1FB9BFB66FBA5" box="[1066,1157,1048,1074]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D7ECA69FF2CFBBFFB23FAE1" pageId="36" pageNumber="37" type="materials_examined">
<paragraph id="4344E72D9D7ECA69FF2CFBBFFC2FFB09" blockId="36.[151,1436,867,1399]" pageId="36" pageNumber="37">
<emphasis id="718F3B3F9D7ECA69FF2CFBBFFE93FBC1" bold="true" box="[199,368,1083,1110]" pageId="36" pageNumber="37">Type material</emphasis>
.
<materialsCitation id="F393ED709D7ECA69FE95FBBFFBAFFBC1" ID-GBIF-Occurrence="2248478868" box="[382,1100,1084,1111]" collectionCode="DZUP" pageId="36" pageNumber="37" specimenCount="1" typeStatus="holotype">
<typeStatus id="9C40598F9D7ECA69FE95FBBFFE05FBC1" box="[382,486,1084,1110]" pageId="36" pageNumber="37" type="holotype">Holotype</typeStatus>
male deposited at the
<collectionCode id="25EA7FE89D7ECA69FD0BFBBEFCCAFBC0" box="[736,809,1085,1111]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="36" pageNumber="37" type="University or college">DZUP</collectionCode>
with the following labels
</materialsCitation>
: /
<materialsCitation id="F393ED709D7ECA69FB8EFBBFFEFCFB09" ID-GBIF-Occurrence="2248478891" collectingDate="1987-08-17" collectingDateMax="1987-08-31" collectingDateMin="1987-08-17" collectorName="Ouro Preto do Oeste, RO &amp; C. Elias &amp; Projeto Polo Noroeste &amp; A. Medeiros" country="Brazil" location="Ouro Preto do Oeste" pageId="36" pageNumber="37" specimenCount="1" typeStatus="holotype">
<typeStatus id="9C40598F9D7ECA69FB8EFBBFFAE8FBC1" box="[1125,1291,1084,1110]" pageId="36" pageNumber="37" type="holotype">HOLOTYPUS</typeStatus>
/
<collectorName id="EE0E82FB9D7ECA69FACAFBBFFECCFBED" pageId="36" pageNumber="37">
<location id="4624B1F69D7ECA69FACAFBBFFF1EFBED" LSID="urn:lsid:plazi:treatment:CB52563B9D78CA69FF7CFB8DFB23FAE1:4624B1F69D7ECA69FACAFBBFFF1EFBED" country="Brazil" name="Ouro Preto do Oeste" pageId="36" pageNumber="37">Ouro Preto do Oeste</location>
, RO
</collectorName>
[ndônia,
<collectingCountry id="3BECA7BD9D7ECA69FE7AFBDCFE38FBED" box="[401,475,1119,1146]" name="Brazil" pageId="36" pageNumber="37">Brazil</collectingCountry>
],
<date id="3745C1ED9D7ECA69FE07FBE3FD48FBED" box="[492,683,1120,1146]" pageId="36" pageNumber="37" value="1987-08-17" valueMax="1987-08-31" valueMin="1987-08-17">
<collectingDate id="270138059D7ECA69FE07FBE3FD48FBED" box="[492,683,1120,1146]" pageId="36" pageNumber="37" value="1987-08-17" valueMax="1987-08-31" valueMin="1987-08-17">17–31.VIII.1987</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D7ECA69FD5DFBE3FCEDFBED" box="[694,782,1120,1146]" pageId="36" pageNumber="37">C. Elias</collectorName>
<emphasis id="718F3B3F9D7ECA69FCFFFBE3FCD5FBED" box="[788,822,1120,1146]" italics="true" pageId="36" pageNumber="37">leg</emphasis>
. /
<collectorName id="EE0E82FB9D7ECA69FCB9FBE3FBABFBEC" box="[850,1096,1120,1147]" pageId="36" pageNumber="37">Projeto Polo Noroeste</collectorName>
/ gen. prep.
<collectorName id="EE0E82FB9D7ECA69FB39FBE3FABDFBED" box="[1234,1374,1120,1146]" pageId="36" pageNumber="37">A. Medeiros</collectorName>
2018 / DZ 41.203
</materialsCitation>
/
<typeStatus id="9C40598F9D7ECA69FEDEFB07FE49FB09" box="[309,426,1156,1182]" pageId="36" pageNumber="37" type="holotype">Holotypus</typeStatus>
<taxonomicName id="84FB9CAE9D7ECA69FE5AFB06FCA5FB09" authority="Medeiros &amp; Dolibaina" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[433,838,1156,1183]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="cofus">
<emphasis id="718F3B3F9D7ECA69FE5AFB06FDA7FB09" box="[433,580,1156,1183]" italics="true" pageId="36" pageNumber="37">Artines cofus</emphasis>
Medeiros &amp; Dolibaina
</taxonomicName>
det. 2018 /.
</paragraph>
<paragraph id="4344E72D9D7ECA69FF2CFB2BFC84FA9D" blockId="36.[151,1436,867,1399]" pageId="36" pageNumber="37">
<materialsCitation id="F393ED709D7ECA69FF2CFB2BFC26FB55" ID-GBIF-Occurrence="2248478849" box="[199,965,1192,1219]" collectionCode="DZUP" pageId="36" pageNumber="37" specimenCount="1" typeStatus="allotype">
<typeStatus id="9C40598F9D7ECA69FF2CFB2BFECAFB55" box="[199,297,1192,1218]" pageId="36" pageNumber="37" type="allotype">Allotype</typeStatus>
female deposited at the
<collectionCode id="25EA7FE89D7ECA69FDA0FB2AFD77FB54" box="[587,660,1193,1219]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="36" pageNumber="37" type="University or college">DZUP</collectionCode>
with the following labels
</materialsCitation>
: /
<materialsCitation id="F393ED709D7ECA69FC0EFB2BFF22FA9D" ID-GBIF-Occurrence="2248478852" collectingDate="1987-08-17" collectingDateMax="1987-08-31" collectingDateMin="1987-08-17" collectorName="Ouro Preto do Oeste, RO &amp; C. Elias &amp; Projeto Polo Noroeste &amp; A. Medeiros" country="Brazil" location="Ouro Preto do Oeste" pageId="36" pageNumber="37" specimenCount="1" typeStatus="allotype">
<typeStatus id="9C40598F9D7ECA69FC0EFB2BFB6BFB55" box="[997,1160,1192,1218]" pageId="36" pageNumber="37" type="allotype">ALLOTYPUS</typeStatus>
/
<collectorName id="EE0E82FB9D7ECA69FB4EFB2BFF5CFB71" pageId="36" pageNumber="37">
<location id="4624B1F69D7ECA69FB4EFB2BFA7BFB55" LSID="urn:lsid:plazi:treatment:CB52563B9D78CA69FF7CFB8DFB23FAE1:4624B1F69D7ECA69FB4EFB2BFA7BFB55" box="[1189,1432,1192,1219]" country="Brazil" name="Ouro Preto do Oeste" pageId="36" pageNumber="37">Ouro Preto do Oeste</location>
, RO
</collectorName>
[ndônia,
<collectingCountry id="3BECA7BD9D7ECA69FEC9FB48FE8FFB71" box="[290,364,1227,1254]" name="Brazil" pageId="36" pageNumber="37">Brazil</collectingCountry>
],
<date id="3745C1ED9D7ECA69FE95FB4FFDDFFB71" box="[382,572,1228,1254]" pageId="36" pageNumber="37" value="1987-08-17" valueMax="1987-08-31" valueMin="1987-08-17">
<collectingDate id="270138059D7ECA69FE95FB4FFDDFFB71" box="[382,572,1228,1254]" pageId="36" pageNumber="37" value="1987-08-17" valueMax="1987-08-31" valueMin="1987-08-17">17–31.VIII.1987</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D7ECA69FDA3FB4FFD42FB71" box="[584,673,1228,1254]" pageId="36" pageNumber="37">C. Elias</collectorName>
<emphasis id="718F3B3F9D7ECA69FD43FB4FFD29FB71" box="[680,714,1228,1254]" italics="true" pageId="36" pageNumber="37">leg</emphasis>
. /
<collectorName id="EE0E82FB9D7ECA69FD0CFB4FFC3CFB70" box="[743,991,1228,1255]" pageId="36" pageNumber="37">Projeto Polo Noroeste</collectorName>
/ gen. prep.
<collectorName id="EE0E82FB9D7ECA69FB80FB4FFB1BFB71" box="[1131,1272,1228,1254]" pageId="36" pageNumber="37">A. Medeiros</collectorName>
2018 / DZ 41. 252
</materialsCitation>
/
<typeStatus id="9C40598F9D7ECA69FF3CFB73FEA6FA9D" box="[215,325,1264,1290]" pageId="36" pageNumber="37" type="allotype">Allotypus</typeStatus>
<taxonomicName id="84FB9CAE9D7ECA69FEA7FB72FD01FA9D" authority="Medeiros &amp; Dolibaina" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[332,738,1264,1291]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="cofus">
<emphasis id="718F3B3F9D7ECA69FEA7FB72FE3CFA9D" box="[332,479,1264,1291]" italics="true" pageId="36" pageNumber="37">Artines cofus</emphasis>
Medeiros &amp; Dolibaina
</taxonomicName>
det. 2018 /.
</paragraph>
<paragraph id="4344E72D9D7ECA69FF2CFA96FB23FAE1" blockId="36.[151,1436,867,1399]" pageId="36" pageNumber="37">
<materialsCitation id="F393ED709D7ECA69FF2CFA96FB37FAC5" ID-GBIF-Occurrence="2248478877" collectingDate="1991-10-25" collectingDateMax="2012-09-05" collectingDateMin="1991-10-25" collectionCode="MUSM, OM" collectorName="R Robbins &amp; Sacanche &amp; C. Calderon" elevation="300" location="Puerto Maldonado" pageId="36" pageNumber="37" specimenCode="OM 73.640" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Madre de Dios" typeStatus="paratype">
<typeStatus id="9C40598F9D7ECA69FF2CFA96FEDBFAB8" box="[199,312,1301,1327]" pageId="36" pageNumber="37" type="paratype">Paratypes</typeStatus>
: PERU—
<emphasis id="718F3B3F9D7ECA69FE43FA97FDB7FAB8" box="[424,596,1300,1327]" italics="true" pageId="36" pageNumber="37">
<collectingRegion id="813F29CF9D7ECA69FE43FA97FDB7FAB8" box="[424,596,1300,1327]" country="Peru" name="Madre de Dios" pageId="36" pageNumber="37">Madre de Díos</collectingRegion>
</emphasis>
:
<emphasis id="718F3B3F9D7ECA69FD8EFA97FCA4FAB9" bold="true" box="[613,839,1300,1326]" pageId="36" pageNumber="37">
<location id="4624B1F69D7ECA69FD8EFA97FCA4FAB9" LSID="urn:lsid:plazi:treatment:CB52563B9D78CA69FF7CFB8DFB23FAE1:4624B1F69D7ECA69FD8EFA97FCA4FAB9" box="[613,839,1300,1326]" name="Puerto Maldonado" pageId="36" pageNumber="37" stateProvince="Madre de Dios">Puerto Maldonado</location>
</emphasis>
(Tambopata),
<quantity id="84034AC89D7ECA69FC04FA97FBD1FAB9" box="[1007,1074,1300,1326]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="36" pageNumber="37" unit="m" value="300.0">
<elevation id="C8D6001E9D7ECA69FC04FA97FBD1FAB9" box="[1007,1074,1300,1326]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="36" pageNumber="37" unit="m" value="300.0">300m</elevation>
</quantity>
,
<specimenCount id="55FD2CA49D7ECA69FBD4FA97FB6CFAB9" box="[1087,1167,1300,1326]" pageId="36" pageNumber="37" type="male">1 male</specimenCount>
,
<date id="3745C1ED9D7ECA69FB76FA90FAF6FAB9" box="[1181,1301,1299,1327]" pageId="36" pageNumber="37" value="1991-10-25">
<collectingDate id="270138059D7ECA69FB76FA90FAF6FAB9" box="[1181,1301,1299,1327]" pageId="36" pageNumber="37" value="1991-10-25">25.X.1991</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D7ECA69FAC8FA96FA7FFAB9" box="[1315,1436,1300,1327]" pageId="36" pageNumber="37">R Robbins</collectorName>
<emphasis id="718F3B3F9D7ECA69FF7CFABBFF22FAC5" box="[151,193,1336,1362]" italics="true" pageId="36" pageNumber="37">leg.</emphasis>
(
<collectionCode id="25EA7FE89D7ECA69FF3BFABBFECFFAC5" box="[208,300,1336,1362]" pageId="36" pageNumber="37">MUSM</collectionCode>
);
<emphasis id="718F3B3F9D7ECA69FEABFABAFE23FAC5" box="[320,448,1337,1363]" italics="true" pageId="36" pageNumber="37">
<collectingRegion id="813F29CF9D7ECA69FEABFABAFE23FAC5" box="[320,448,1337,1363]" country="Peru" name="San Martin" pageId="36" pageNumber="37">San Martin</collectingRegion>
</emphasis>
:
<emphasis id="718F3B3F9D7ECA69FE24FAB4FDDCFAC5" bold="true" box="[463,575,1335,1362]" pageId="36" pageNumber="37">
<collectorName id="EE0E82FB9D7ECA69FE24FAB4FDDCFAC5" box="[463,575,1335,1362]" pageId="36" pageNumber="37">Sacanche</collectorName>
</emphasis>
,
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,
<date id="3745C1ED9D7ECA69FD54FAB4FCD0FAC5" box="[703,819,1335,1362]" pageId="36" pageNumber="37" value="2012-09-05">
<collectingDate id="270138059D7ECA69FD54FAB4FCD0FAC5" box="[703,819,1335,1362]" pageId="36" pageNumber="37" value="2012-09-05">5.IX.2012</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D7ECA69FCD4FABBFC2BFAC5" box="[831,968,1336,1362]" pageId="36" pageNumber="37">C. Calderon</collectorName>
<emphasis id="718F3B3F9D7ECA69FC24FABBFC1BFAC5" box="[975,1016,1336,1362]" italics="true" pageId="36" pageNumber="37">leg.</emphasis>
,
<specimenCode id="135D4F569D7ECA69FBECFABBFB6AFAC5" box="[1031,1161,1336,1362]" collectionCode="OM" country="New Zealand" name="Otago Museum" pageId="36" pageNumber="37">OM 73.640</specimenCode>
(
<collectionCode id="25EA7FE89D7ECA69FB72FABBFB2FFAC5" box="[1177,1228,1336,1362]" country="New Zealand" name="Otago Museum" pageId="36" pageNumber="37">OM</collectionCode>
)
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BRAZIL—
<emphasis id="718F3B3F9D7ECA69FA8AFABAFA76FAC4" box="[1377,1429,1337,1363]" italics="true" pageId="36" pageNumber="37">
<collectingRegion id="813F29CF9D7ECA69FA8AFABAFA76FAC4" box="[1377,1429,1337,1363]" country="Brazil" name="Acre" pageId="36" pageNumber="37">Acre</collectingRegion>
</emphasis>
:
<emphasis id="718F3B3F9D7ECA69FF7CFAD8FE72FAE1" bold="true" box="[151,401,1371,1398]" pageId="36" pageNumber="37">
<collectingRegion id="813F29CF9D7ECA69FF7CFAD8FEFDFAE1" box="[151,286,1371,1398]" country="Guatemala" name="Santa Rosa" pageId="36" pageNumber="37">Santa Rosa</collectingRegion>
do
<location id="4624B1F69D7ECA69FEA2FADFFE72FAE1" LSID="urn:lsid:plazi:treatment:CB52563B9D78CA69FF7CFB8DFB23FAE1:4624B1F69D7ECA69FEA2FADFFE72FAE1" box="[329,401,1372,1398]" name="Purus" pageId="36" pageNumber="37" stateProvince="Acre">Purus</location>
</emphasis>
,
<specimenCount id="55FD2CA49D7ECA69FE74FADFFE0EFAE1" box="[415,493,1372,1398]" pageId="36" pageNumber="37" type="male">1 male</specimenCount>
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<date id="3745C1ED9D7ECA69FE13FADFFD78FAE1" box="[504,667,1372,1398]" pageId="36" pageNumber="37" value="2008-08-06" valueMax="2008-08-08" valueMin="2008-08-06">
<collectingDate id="270138059D7ECA69FE13FADFFD78FAE1" box="[504,667,1372,1398]" pageId="36" pageNumber="37" value="2008-08-06" valueMax="2008-08-08" valueMin="2008-08-06">6–8.VIII.2008</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D7ECA69FD4CFADFFCFBFAE1" box="[679,792,1372,1398]" pageId="36" pageNumber="37">O. Mielke</collectorName>
&amp;
<collectorName id="EE0E82FB9D7ECA69FCD7FADFFC7DFAE1" box="[828,926,1372,1398]" pageId="36" pageNumber="37">Carneiro</collectorName>
<emphasis id="718F3B3F9D7ECA69FC4EFADFFC24FAE1" box="[933,967,1372,1398]" italics="true" pageId="36" pageNumber="37">leg</emphasis>
., DZ 41.303 (
<collectionCode id="25EA7FE89D7ECA69FB8FFADFFB57FAE1" box="[1124,1204,1372,1398]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="36" pageNumber="37" type="University or college">DZUP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
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681
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<heading id="180C50419D7DCA6AFF7CFCA2FDA7FCAD" bold="true" box="[151,580,800,827]" fontSize="11" level="1" pageId="39" pageNumber="40" reason="1">
<taxonomicName id="84FB9CAE9D7DCA6AFF7CFCA2FDA7FCAD" ID-CoL="GZYF" authority="O. Mielke, 1968" authorityName="O. Mielke" authorityYear="1968" box="[151,580,800,827]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="bipunctata">
<emphasis id="718F3B3F9D7DCA6AFF7CFCA2FDA7FCAD" bold="true" box="[151,580,800,827]" pageId="39" pageNumber="40">
<emphasis id="718F3B3F9D7DCA6AFF7CFCA2FE9BFCAC" bold="true" box="[151,376,800,827]" italics="true" pageId="39" pageNumber="40">Artines bipunctata</emphasis>
O.
<bibRefCitation id="276A9ADC9D7DCA6AFE4FFCA3FDA7FCAD" author="Mielke, O. H. H." box="[420,580,800,826]" pageId="39" pageNumber="40" pagination="447 - 455" refId="ref43084" refString="Mielke, O. H. H. (1968) Lepidoptera do Planalto Central brasileiro. V: Novas especies de Hesperiidae e anotacoes sobre outras especies conhecidas. Revista Brasileira de Biologia, 28 (4), 447 - 455." type="journal article" year="1968">Mielke, 1968</bibRefCitation>
</emphasis>
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<paragraph id="4344E72D9D7DCA6AFF7CFCC7FE5FFCC8" blockId="39.[151,580,800,863]" box="[151,444,836,863]" pageId="39" pageNumber="40">
<figureCitation id="DBC0FBA89D7DCA6AFF7CFCC7FF0FFCC8" box="[151,236,836,863]" captionStart="FIGURES 14–19" captionStartId="10.[151,264,1406,1431]" captionTargetBox="[155,1435,175,1386]" captionTargetId="figure@10.[151,1436,174,1394]" captionTargetPageId="10" captionText="FIGURES 14–19. Species of the “aquilina group” (14–18) and “bipunctata group” (19) in dorsal and ventral views. 14. Artines aquilina, A–B: male, Joinville, Santa Catarina, Brazil (DZ 41.321), C–D: female from same locality of male (DZ 41.220). 15. Artines angelica sp. nov., A–B: holotype male, C–D: allotype female. 16. Artines focus, A–B: male, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.233), C–D: female, Pacaraima, Roraima, Brazil (DZ 41.333). 17. Artines cofus sp. nov., A–B: holotype male, C–D: allotype female. 18. Artines donia sp. nov., A–B: holotype male. 19. Artines bipunctata, A–B: male, Jardim Zoológico, Brasília, Distrito Federal, Brazil (DZ 30.961), C–D: female, Córrego Bananal, Brasília, Distrito Federal, Brazil (DZ 41.224). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241045/files/figure.png" pageId="39" pageNumber="40">Figs 19</figureCitation>
,
<figureCitation id="DBC0FBA89D7DCA6AFF13FCC7FEF5FCC9" box="[248,278,836,862]" captionStart="FIGURES 21–26" captionStartId="22.[151,264,1840,1865]" captionTargetBox="[151,1436,922,1828]" captionTargetId="figure@22.[151,1436,922,1828]" captionText="FIGURES 21–26. Male brands (no scales) on dorsal forewing of the “aquilina group” (21–25) and “bipunctata group” species (26). 21. Artines aquilina, Joinville, Santa catarina, Brazil, (DZ 41.341). 22. Artines angelica sp. nov., paratype, Fazenda Ter- rasse, Santa Luzia, Maranhão, Brazil (DZ 41.273) 23. Artines focus, Ilha de Maracá, Roraima, Brazil (DZ 41.233). 24. Artines cofus sp. nov., paratype, Santa Rosa do Purus, Acre, Brazil (DZ 41.303). 25. Artines donia sp. nov., paratype, Santa Rita do Araguaia, Goiás, Brazil (OM 66.000). 26. Artines bipunctata, Ponte Funda, Vianópilis, Goiás, Brazil (DZ 41.264). Scale bar = 1 mm." httpUri="https://zenodo.org/record/3406270/files/figure.png" pageId="39" pageNumber="40">26</figureCitation>
,
<figureCitation id="DBC0FBA89D7DCA6AFEC9FCC6FEA3FCC8" box="[290,320,837,863]" captionStart="FIGURES 40–41" captionStartId="31.[151,264,1624,1649]" captionTargetBox="[154,1435,484,1599]" captionTargetId="figure@31.[151,1436,481,1612]" captionText="FIGURES 40–41. Male genitalia of the “aquilina group” (40) and “bipunctata group” (41) species. 40. Artines donia sp. nov., paratype, Santa Rita do Araguaia, Goiás, Brazil (DZ 66.000). 41. Artines bipunctata, Brasília, Distrito Federal, Brazil (DZ 41.194). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3406272/files/figure.png" pageId="39" pageNumber="40">41</figureCitation>
, 65, 77, 80
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<taxonomicName id="84FB9CAE9D7DCA6AFF7CFC0AFDE8FC36" ID-CoL="GZYF" authority="O. Mielke, 1968" authorityName="O. Mielke" authorityYear="1968" box="[151,523,904,929]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="bipunctata">
<emphasis id="718F3B3F9D7DCA6AFF7CFC0AFEB6FC36" box="[151,341,905,929]" italics="true" pageId="39" pageNumber="40">Artines bipunctata</emphasis>
O.
<bibRefCitation id="276A9ADC9D7DCA6AFE95FC0BFDE8FC36" author="Mielke, O. H. H." box="[382,523,904,929]" pageId="39" pageNumber="40" pagination="447 - 455" refId="ref43084" refString="Mielke, O. H. H. (1968) Lepidoptera do Planalto Central brasileiro. V: Novas especies de Hesperiidae e anotacoes sobre outras especies conhecidas. Revista Brasileira de Biologia, 28 (4), 447 - 455." type="journal article" year="1968">Mielke, 1968</bibRefCitation>
</taxonomicName>
.
<emphasis id="718F3B3F9D7DCA6AFDFDFC0AFD07FC36" bold="true" box="[534,740,904,929]" pageId="39" pageNumber="40">Rev. Bras. Biol. 28</emphasis>
: 449, figs 7-10 (male, female d, v), 18-22 (male gen.), 23 (brand), 24 (female gen.);
<typeStatus id="9C40598F9D7DCA6AFE96FC24FE35FC57" box="[381,470,935,960]" pageId="39" pageNumber="40" type="holotype">holotype</typeStatus>
male,
<date id="3745C1ED9D7DCA6AFDF7FC24FD9CFC57" box="[540,639,935,960]" pageId="39" pageNumber="40" value="1967-02-03">3-II-1967</date>
, Jardim Zoológico, Brasília, D[istrito] F[ederal],
<collectingCountry id="3BECA7BD9D7DCA6AFB93FC24FB5AFC57" box="[1144,1209,935,960]" name="Brazil" pageId="39" pageNumber="40">Brazil</collectingCountry>
; O. Mielke collection, DZUP.—
<bibRefCitation id="276A9ADC9D7DCA6AFEC1FC46FE58FC49" author="Bridges, C. A." box="[298,443,965,990]" pageId="39" pageNumber="40" refId="ref38436" refString="Bridges, C. A. (1983) Lepidoptera: Hesperiidae. Notes on species-group names. Vol. 1 - 5. Author, Urbana, Author, 277 pp. [2 pp., Vol. 1: 129 pp., Vol. 2: 41 pp., Vol. 3: 62 pp., Vol. 4: 30 pp., Vol. 5: 13 pp.]" type="book" year="1983">Bridges, 1983</bibRefCitation>
.
<emphasis id="718F3B3F9D7DCA6AFE2DFC45FDACFC49" bold="true" box="[454,591,965,990]" pageId="39" pageNumber="40">Lep. Hesp. 1</emphasis>
, p. 16;
<emphasis id="718F3B3F9D7DCA6AFD75FC46FD48FC49" bold="true" box="[670,683,965,990]" pageId="39" pageNumber="40">2</emphasis>
, p. 3.—Bridges, 1988.
<emphasis id="718F3B3F9D7DCA6AFC70FC46FBC1FC49" bold="true" box="[923,1058,965,990]" pageId="39" pageNumber="40">Cat. Hesp. 1</emphasis>
, p. 26;
<emphasis id="718F3B3F9D7DCA6AFB9AFC46FB9DFC49" bold="true" box="[1137,1150,965,990]" pageId="39" pageNumber="40">2</emphasis>
, p. 5.—
<bibRefCitation id="276A9ADC9D7DCA6AFB3BFC46FA87FC49" author="Bridges, C. A." box="[1232,1380,965,990]" pageId="39" pageNumber="40" refId="ref38783" refString="Bridges, C. A. (1994) Catalogue of the family-group, genus-group and species-group names of the Hesperioidea (Lepidoptera) of the world. Vols. 1 - 13 &amp; Apps. 1 - 3. Author, Urbana, 3. [11] + 598 pp. [11 pp., Vol. 1: 1 pp., Vol. 2: 1 pp., Vol. 3: 1 pp., Vol. 4: 36 pp., Vol. 5: 9 pp., Vol. 6: 20 pp., Vol. 7: 6 pp., Vol. 8: 246 pp., Vol. 9: 76 pp., Vol. 10: 90 pp., Vol. 11: 71 pp., Vol. 12: 28 pp., Vol. 13: 9 pp., App. 1: 1 pp., App. 2: 2 pp., App. 3: 1 pp.]" type="book" year="1994">Bridges, 1994</bibRefCitation>
.
<emphasis id="718F3B3F9D7DCA6AFA85FC46FC73FC6A" bold="true" pageId="39" pageNumber="40">Cat. Fam.-Group, Gen.-Group, Sp. Group Nam. Hesp. (Lep.) World 8</emphasis>
, p. 31;
<emphasis id="718F3B3F9D7DCA6AFC34FC67FC0FFC6A" bold="true" box="[991,1004,996,1021]" pageId="39" pageNumber="40">9</emphasis>
, p. 6.—O.
<bibRefCitation id="276A9ADC9D7DCA6AFB8BFC67FB0FFC6A" author="Mielke, O. H. H." box="[1120,1260,996,1021]" pageId="39" pageNumber="40" pagination="3 - 11" refId="ref43459" refString="Mielke, O. H. H. (2004) Hesperiidae. In: Lamas, G. (Ed.), Checklist: Part 4 A, Hesperioidea-Papilionoidea. In: Heppner, J. (Ed.), Atlas of Neotropical Lepidoptera 5 A. Scientific Publishers, Association for Tropical Lepidoptera, Gainesville, pp. 3 - 11 + 25 - 86." type="book chapter" year="2004">Mielke, 2004</bibRefCitation>
. Hesperioidea, p. 62,
<emphasis id="718F3B3F9D7DCA6AFF04FB80FEE7FB8C" bold="true" box="[239,260,1027,1051]" italics="true" pageId="39" pageNumber="40">in</emphasis>
Lamas (ed.).
<emphasis id="718F3B3F9D7DCA6AFE7DFB80FC78FB8B" bold="true" box="[406,923,1027,1052]" pageId="39" pageNumber="40">Checklist: Part 4A, Hesperioidea-Papilionoidea</emphasis>
,
<emphasis id="718F3B3F9D7DCA6AFC41FB80FC5CFB8C" bold="true" box="[938,959,1027,1051]" italics="true" pageId="39" pageNumber="40">in</emphasis>
Heppner (ed.).
<emphasis id="718F3B3F9D7DCA6AFB8EFB80FA87FB8B" bold="true" box="[1125,1380,1027,1052]" pageId="39" pageNumber="40">
<taxonomicName id="84FB9CAE9D7DCA6AFB8EFB80FADEFB8B" authority="Neotrop. Lep." authorityName="Neotrop. Lep." box="[1125,1341,1027,1052]" class="Insecta" genus="Atlas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">Atlas Neotrop. Lep.</taxonomicName>
5A
</emphasis>
.—O.
<bibRefCitation id="276A9ADC9D7DCA6AFF2CFBA2FEB2FBAD" author="Mielke, O. H. H." box="[199,337,1057,1082]" pageId="39" pageNumber="40" pagination="775 - 1055" refId="ref43529" refString="Mielke, O. H. H. (2005) Catalogue of the American Hesperioidea: Hesperiidae (Lepidoptera) 4. Hesperiinae 1. Adlerodea- Lychnuchus. Sociedade Brasileira de Zoologia, Curitiba. pp. 775 - 1055." type="book chapter" year="2005">Mielke, 2005</bibRefCitation>
.
<emphasis id="718F3B3F9D7DCA6AFEB2FBA2FD91FBAD" bold="true" box="[345,626,1057,1082]" pageId="39" pageNumber="40">Cat.Amer. Hesperioidea 4</emphasis>
, p. 833.—
<bibRefCitation id="276A9ADC9D7DCA6AFD3CFBA2FC31FBAD" author="Pinheiro, C. E. G. &amp; Emery, E. O." box="[727,978,1057,1083]" pageId="39" pageNumber="40" pagination="1 - 15" refId="ref44309" refString="Pinheiro, C. E. G. &amp; Emery, E. O. (2006) As borboletas (Lepidoptera: Papilionoidea e Hesperioidea) da Area de Protecao Ambiental do Gama e Cabeca de Veado (Distrito Federal, Brasil). Biota Neotropica, 6 (3), 1 - 15. https: // doi. org / 10.1590 / S 1676 - 06032006000300012" type="journal article" year="2006">Pinheiro &amp; Emery, 2006</bibRefCitation>
.
<emphasis id="718F3B3F9D7DCA6AFC30FBA2FB69FBAD" bold="true" box="[987,1162,1057,1082]" pageId="39" pageNumber="40">
<taxonomicName id="84FB9CAE9D7DCA6AFC30FBA2FB9BFBAD" authority="Neotrop." authorityName="Neotrop." box="[987,1144,1057,1082]" class="Pinopsida" family="Cupressaceae" genus="Biota" higherTaxonomySource="CoL" kingdom="Plantae" order="Pinales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="genus">Biota Neotrop.</taxonomicName>
6
</emphasis>
(3): 6.—
<bibRefCitation id="276A9ADC9D7DCA6AFB36FBA2FF1EFBCE" author="Pinheiro, C. E. G. &amp; Emery, E. O." pageId="39" pageNumber="40" pagination="61 - 74" refId="ref44379" refString="Pinheiro, C. E. G. &amp; Emery, E. O. (2007) As borboletas (Lepidoptera: Papilionoidea e Hesperioidea) da Estacao Ecologica do Jardim Botanico, Reserva Ecologica do IBGE e Fazenda Agua Limpa (Distrito Federal). Heringeriana, 1 (1), 61 - 74." type="journal article" year="2007">Pinheiro &amp; Emery, 2007</bibRefCitation>
.
<emphasis id="718F3B3F9D7DCA6AFEECFBC3FE4FFBCE" bold="true" box="[263,428,1088,1113]" pageId="39" pageNumber="40">Heringeriana 1</emphasis>
(1): 65.—O. Mielke; Emery &amp; Pinheiro, 2008.
<emphasis id="718F3B3F9D7DCA6AFC64FBC2FB83FBCE" bold="true" box="[911,1120,1088,1113]" pageId="39" pageNumber="40">Revta bras. Ent. 52</emphasis>
(2): 286, 287.—
<bibRefCitation id="276A9ADC9D7DCA6AFAE8FBC3FF1FFBE0" author="Pinheiro, C. E. G. &amp; Malinov, I. K. &amp; Emery, E. O. &amp; Schmidt, K." pageId="39" pageNumber="40" pagination="223 - 238" refId="ref44437" refString="Pinheiro, C. E. G., Malinov, I. K., Emery, E. O. &amp; Schmidt, K. (2010) Endemismos e conservacao de borboletas (Lepidoptera: Papilionoidea e Hesperioidea) no bioma Cerrado. In: Diniz, I. R., Marinho Filho, J., Machado, J. R. B. &amp; Cavalcanti, R. B. (Eds.), Cerrado. Conhecimento cientifico quantitativo como subsidio para acoes de conservacao. Universidade de Brasilia, Brasilia, pp. 223 - 238." type="book chapter" year="2010">
Pinheiro
<emphasis id="718F3B3F9D7DCA6AFA89FBC2FA73FBCE" bold="true" box="[1378,1424,1088,1113]" italics="true" pageId="39" pageNumber="40">et al</emphasis>
., 2010
</bibRefCitation>
,
<emphasis id="718F3B3F9D7DCA6AFEECFBDCFEFFFBE0" bold="true" box="[263,284,1119,1143]" italics="true" pageId="39" pageNumber="40">in</emphasis>
Diniz
<emphasis id="718F3B3F9D7DCA6AFE8AFBE3FE6EFBEF" bold="true" box="[353,397,1119,1144]" italics="true" pageId="39" pageNumber="40">et al</emphasis>
.
<emphasis id="718F3B3F9D7DCA6AFE71FBDCFE16FBEF" bold="true" box="[410,501,1119,1144]" pageId="39" pageNumber="40">Cerrado</emphasis>
, p. 235, 237.
</paragraph>
<paragraph id="4344E72D9D7DCA6AFF7CFBFDFD0DFB01" blockId="39.[151,1437,904,1174]" box="[151,750,1149,1174]" pageId="39" pageNumber="40">
(no genus)
<taxonomicName id="84FB9CAE9D7DCA6AFEE2FBFDFE96FB01" ID-CoL="GZYF" authorityName="O. Mielke" authorityYear="1968" box="[265,373,1150,1174]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="bipunctata">
<emphasis id="718F3B3F9D7DCA6AFEE2FBFDFE96FB01" box="[265,373,1150,1174]" italics="true" pageId="39" pageNumber="40">bipunctata</emphasis>
</taxonomicName>
;
<bibRefCitation id="276A9ADC9D7DCA6AFE68FBFEFDEEFB01" author="Beattie, J. R." box="[387,525,1149,1174]" pageId="39" pageNumber="40" refId="ref38340" refString="Beattie, J. R. (1976) The Rhopalocera Directory. JB Indexes, Berkeley, xiv + 365 pp." type="book" year="1976">Beattie, 1976</bibRefCitation>
.
<emphasis id="718F3B3F9D7DCA6AFDFCFBFDFD42FB01" bold="true" box="[535,673,1149,1174]" pageId="39" pageNumber="40">Rhop. Direct</emphasis>
., p. 96.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D7DCA6AFF7CFB47FAC7FAB1" pageId="39" pageNumber="40" type="description">
<paragraph id="4344E72D9D7DCA6AFF7CFB47FAC7FAB1" blockId="39.[151,1437,1220,2039]" pageId="39" pageNumber="40">
<emphasis id="718F3B3F9D7DCA6AFF7CFB47FE95FB48" bold="true" box="[151,374,1220,1247]" pageId="39" pageNumber="40">Taxonomic history</emphasis>
.
<taxonomicName id="84FB9CAE9D7DCA6AFE6FFB46FDB6FB49" authorityName="O. Mielke" authorityYear="1968" box="[388,597,1220,1247]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="bipunctata">
<emphasis id="718F3B3F9D7DCA6AFE6FFB46FDB6FB49" box="[388,597,1220,1247]" italics="true" pageId="39" pageNumber="40">Artines bipunctata</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="276A9ADC9D7DCA6AFCCEFB47FC24FB48" author="Mielke, O. H. H." box="[805,967,1220,1247]" pageId="39" pageNumber="40" pagination="447 - 455" refId="ref43084" refString="Mielke, O. H. H. (1968) Lepidoptera do Planalto Central brasileiro. V: Novas especies de Hesperiidae e anotacoes sobre outras especies conhecidas. Revista Brasileira de Biologia, 28 (4), 447 - 455." type="journal article" year="1968">Mielke (1968)</bibRefCitation>
based on seven specimens (one male and six females) from Jardim Zoológico, Brasília, Distrito Federal,
<collectingCountry id="3BECA7BD9D7DCA6AFCB2FB6BFC7EFA95" box="[857,925,1256,1282]" name="Brazil" pageId="39" pageNumber="40">Brazil</collectingCountry>
and one male from Barbacena,
<collectingRegion id="813F29CF9D7DCA6AFB14FB6BFA7BFA95" box="[1279,1432,1256,1282]" country="Brazil" name="Minas Gerais" pageId="39" pageNumber="40">Minas Gerais</collectingRegion>
,
<collectingCountry id="3BECA7BD9D7DCA6AFF7CFA8FFF3CFAB1" box="[151,223,1292,1318]" name="Brazil" pageId="39" pageNumber="40">Brazil</collectingCountry>
. The other authors mentioned this species in taxonomic comments, faunistic studies and catalogs.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D7DCA6AFF2CFAB3FDB2F9FC" pageId="39" pageNumber="40" type="diagnosis">
<paragraph id="4344E72D9D7DCA6AFF2CFAB3FDB2F9FC" blockId="39.[151,1437,1220,2039]" pageId="39" pageNumber="40">
<emphasis id="718F3B3F9D7DCA6AFF2CFAB3FED9FADC" bold="true" box="[199,314,1328,1355]" pageId="39" pageNumber="40">Diagnosis</emphasis>
.
<taxonomicName id="84FB9CAE9D7DCA6AFEA0FAB2FDFEFADD" authorityName="O. Mielke" authorityYear="1968" box="[331,541,1328,1355]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="bipunctata">
<emphasis id="718F3B3F9D7DCA6AFEA0FAB2FDFEFADD" box="[331,541,1328,1355]" italics="true" pageId="39" pageNumber="40">Artines bipunctata</emphasis>
</taxonomicName>
differs from other species of the genus in the following combination of characters: dorsal forewing with large and semi-hyaline discal spots; ventral hind wing with discal area pale brown, without central black spot and a violet postdiscal band; male with brand; uncus wide and distally bifid; gnathos arms distally curved inward; valva distally rounded; harpe simple and not fused with ampulla; ventral projection of aedeagus wide and parallel to aedeagus, distal end bilobed, with rounded lobes bearing numerous spines; vesica entirely membranous, with very small and dispersed spines; fultura inferior laterally straight, proximal end bifid; sterigma partially sclerotized; lamella postvaginalis bifid and without lateral folds; region between seventh sternite and lamella antevaginalis completely membranous; ductus bursae with basal portion sclerotized; ostium wide, “D”- shaped, surrounded by sclerotized area.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D7DCA65FF2CF9F7FE0BFE3C" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40" type="description">
<paragraph id="4344E72D9D7DCA6AFF2CF9F7FB74F918" blockId="39.[151,1437,1220,2039]" box="[199,1175,1652,1679]" pageId="39" pageNumber="40">
<emphasis id="718F3B3F9D7DCA6AFF2CF9F7FEB2F918" bold="true" box="[199,337,1652,1679]" pageId="39" pageNumber="40">Description</emphasis>
. Male (
<figureCitation id="DBC0FBA89D7DCA6AFE43F9F7FE1BF918" box="[424,504,1652,1679]" captionStart="FIGURES 14–19" captionStartId="10.[151,264,1406,1431]" captionTargetBox="[155,1435,175,1386]" captionTargetId="figure@10.[151,1436,174,1394]" captionTargetPageId="10" captionText="FIGURES 14–19. Species of the “aquilina group” (14–18) and “bipunctata group” (19) in dorsal and ventral views. 14. Artines aquilina, A–B: male, Joinville, Santa Catarina, Brazil (DZ 41.321), C–D: female from same locality of male (DZ 41.220). 15. Artines angelica sp. nov., A–B: holotype male, C–D: allotype female. 16. Artines focus, A–B: male, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.233), C–D: female, Pacaraima, Roraima, Brazil (DZ 41.333). 17. Artines cofus sp. nov., A–B: holotype male, C–D: allotype female. 18. Artines donia sp. nov., A–B: holotype male. 19. Artines bipunctata, A–B: male, Jardim Zoológico, Brasília, Distrito Federal, Brazil (DZ 30.961), C–D: female, Córrego Bananal, Brasília, Distrito Federal, Brazil (DZ 41.224). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241045/files/figure.png" pageId="39" pageNumber="40">Fig. 19</figureCitation>
A–B). External morphology as described by
<bibRefCitation id="276A9ADC9D7DCA6AFC05F9F7FB70F918" author="Mielke, O. H. H." box="[1006,1171,1652,1679]" pageId="39" pageNumber="40" pagination="447 - 455" refId="ref43084" refString="Mielke, O. H. H. (1968) Lepidoptera do Planalto Central brasileiro. V: Novas especies de Hesperiidae e anotacoes sobre outras especies conhecidas. Revista Brasileira de Biologia, 28 (4), 447 - 455." type="journal article" year="1968">Mielke (1968)</bibRefCitation>
.
</paragraph>
<paragraph id="4344E72D9D7DCA6AFF2CF91BFE81F860" blockId="39.[151,1437,1220,2039]" pageId="39" pageNumber="40">
<emphasis id="718F3B3F9D7DCA6AFF2CF91BFED2F925" box="[199,305,1688,1714]" italics="true" pageId="39" pageNumber="40">Genitalia</emphasis>
(
<figureCitation id="DBC0FBA89D7DCA6AFEA8F91BFE74F924" box="[323,407,1688,1715]" captionStart="FIGURES 40–41" captionStartId="31.[151,264,1624,1649]" captionTargetBox="[154,1435,484,1599]" captionTargetId="figure@31.[151,1436,481,1612]" captionText="FIGURES 40–41. Male genitalia of the “aquilina group” (40) and “bipunctata group” (41) species. 40. Artines donia sp. nov., paratype, Santa Rita do Araguaia, Goiás, Brazil (DZ 66.000). 41. Artines bipunctata, Brasília, Distrito Federal, Brazil (DZ 41.194). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3406272/files/figure.png" pageId="39" pageNumber="40">Fig. 41</figureCitation>
A–J): tegumen more or less rectangular, wider than long, with short and triangular distal median projection (
<figureCitation id="DBC0FBA89D7DCA6AFE9CF93FFE02F940" box="[375,481,1724,1751]" captionStart="FIGURES 40–41" captionStartId="31.[151,264,1624,1649]" captionTargetBox="[154,1435,484,1599]" captionTargetId="figure@31.[151,1436,481,1612]" captionText="FIGURES 40–41. Male genitalia of the “aquilina group” (40) and “bipunctata group” (41) species. 40. Artines donia sp. nov., paratype, Santa Rita do Araguaia, Goiás, Brazil (DZ 66.000). 41. Artines bipunctata, Brasília, Distrito Federal, Brazil (DZ 41.194). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3406272/files/figure.png" pageId="39" pageNumber="40">Fig. 41C</figureCitation>
). Saccus triangular, base shorter than lateral margin (
<figureCitation id="DBC0FBA89D7DCA6AFBACF93FFB52F940" box="[1095,1201,1724,1751]" captionStart="FIGURES 40–41" captionStartId="31.[151,264,1624,1649]" captionTargetBox="[154,1435,484,1599]" captionTargetId="figure@31.[151,1436,481,1612]" captionText="FIGURES 40–41. Male genitalia of the “aquilina group” (40) and “bipunctata group” (41) species. 40. Artines donia sp. nov., paratype, Santa Rita do Araguaia, Goiás, Brazil (DZ 66.000). 41. Artines bipunctata, Brasília, Distrito Federal, Brazil (DZ 41.194). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3406272/files/figure.png" pageId="39" pageNumber="40">Fig. 41B</figureCitation>
). Uncus rectangular, one and a half longer than wide, distally bifid, with rounded projections separated medially by shallow concavity (
<figureCitation id="DBC0FBA89D7DCA6AFF4BF887FEE5F888" box="[160,262,1796,1823]" captionStart="FIGURES 40–41" captionStartId="31.[151,264,1624,1649]" captionTargetBox="[154,1435,484,1599]" captionTargetId="figure@31.[151,1436,481,1612]" captionText="FIGURES 40–41. Male genitalia of the “aquilina group” (40) and “bipunctata group” (41) species. 40. Artines donia sp. nov., paratype, Santa Rita do Araguaia, Goiás, Brazil (DZ 66.000). 41. Artines bipunctata, Brasília, Distrito Federal, Brazil (DZ 41.194). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3406272/files/figure.png" pageId="39" pageNumber="40">Fig. 41C</figureCitation>
). Gnathos arms distally curved inward (
<figureCitation id="DBC0FBA89D7DCA6AFD28F887FCC8F888" box="[707,811,1796,1823]" captionStart="FIGURES 40–41" captionStartId="31.[151,264,1624,1649]" captionTargetBox="[154,1435,484,1599]" captionTargetId="figure@31.[151,1436,481,1612]" captionText="FIGURES 40–41. Male genitalia of the “aquilina group” (40) and “bipunctata group” (41) species. 40. Artines donia sp. nov., paratype, Santa Rita do Araguaia, Goiás, Brazil (DZ 66.000). 41. Artines bipunctata, Brasília, Distrito Federal, Brazil (DZ 41.194). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3406272/files/figure.png" pageId="39" pageNumber="40">Fig. 41D</figureCitation>
). Valva twice longer than wide; sacculus triangular, narrow, longer than 1/2 of valva; costa rectangular and short, near median portion of valva, distally fused to ampulla; ampulla rectangular, longer than costa; harpe simple and not fused to ampulla, distal margin rounded with arched row of short and triangular spines turned inwards (
<figureCitation id="DBC0FBA89D7DCA6AFD03F8F3FCDFF81C" box="[744,828,1904,1931]" captionStart="FIGURES 40–41" captionStartId="31.[151,264,1624,1649]" captionTargetBox="[154,1435,484,1599]" captionTargetId="figure@31.[151,1436,481,1612]" captionText="FIGURES 40–41. Male genitalia of the “aquilina group” (40) and “bipunctata group” (41) species. 40. Artines donia sp. nov., paratype, Santa Rita do Araguaia, Goiás, Brazil (DZ 66.000). 41. Artines bipunctata, Brasília, Distrito Federal, Brazil (DZ 41.194). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3406272/files/figure.png" pageId="39" pageNumber="40">Fig. 41</figureCitation>
E–F). Aedeagus thick, longer than valva; opening of ejaculatory bulb on proximal end of aedeagus; ventral projection of aedeagus wide and parallel to aedeagus, distal end bilobed, with rounded lobes bearing numerous spines (
<figureCitation id="DBC0FBA89D7DCA6AFCD0F83BFC6EF844" box="[827,909,1976,2003]" captionStart="FIGURES 40–41" captionStartId="31.[151,264,1624,1649]" captionTargetBox="[154,1435,484,1599]" captionTargetId="figure@31.[151,1436,481,1612]" captionText="FIGURES 40–41. Male genitalia of the “aquilina group” (40) and “bipunctata group” (41) species. 40. Artines donia sp. nov., paratype, Santa Rita do Araguaia, Goiás, Brazil (DZ 66.000). 41. Artines bipunctata, Brasília, Distrito Federal, Brazil (DZ 41.194). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3406272/files/figure.png" pageId="39" pageNumber="40">Fig. 41</figureCitation>
G–J). Vesica bearing small and short scattered spines (
<figureCitation id="DBC0FBA89D7DCA6AFF06F85FFEB6F860" box="[237,341,2012,2039]" captionStart="FIGURES 38–39" captionStartId="29.[151,264,1720,1745]" captionTargetBox="[161,1436,431,1681]" captionTargetId="figure@29.[151,1436,450,1693]" captionTargetPageId="29" captionText="FIGURES 38–39. Male genitalia of the “aquilina group” species. 38. Artines focus, Reserva Duque, Manaus, Amazonas, Brazil (DZ 41.192). 39. Artines cofus sp. nov., paratype, Tambopata, Puerto Maldonado Madre de Díos, Peru (MUSM). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241055/files/figure.png" pageId="39" pageNumber="40">Fig. 39G</figureCitation>
).
</paragraph>
<paragraph id="4344E72D9D72CA65FF2CFF1AFCC1FF23" blockId="40.[151,1437,153,1634]" box="[199,802,153,180]" pageId="40" pageNumber="41">
Female (
<figureCitation id="DBC0FBA89D72CA65FEC3FF1AFE92FF24" box="[296,369,153,179]" captionStart="FIGURES 14–19" captionStartId="10.[151,264,1406,1431]" captionTargetBox="[155,1435,175,1386]" captionTargetId="figure@10.[151,1436,174,1394]" captionTargetPageId="10" captionText="FIGURES 14–19. Species of the “aquilina group” (14–18) and “bipunctata group” (19) in dorsal and ventral views. 14. Artines aquilina, A–B: male, Joinville, Santa Catarina, Brazil (DZ 41.321), C–D: female from same locality of male (DZ 41.220). 15. Artines angelica sp. nov., A–B: holotype male, C–D: allotype female. 16. Artines focus, A–B: male, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.233), C–D: female, Pacaraima, Roraima, Brazil (DZ 41.333). 17. Artines cofus sp. nov., A–B: holotype male, C–D: allotype female. 18. Artines donia sp. nov., A–B: holotype male. 19. Artines bipunctata, A–B: male, Jardim Zoológico, Brasília, Distrito Federal, Brazil (DZ 30.961), C–D: female, Córrego Bananal, Brasília, Distrito Federal, Brazil (DZ 41.224). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241045/files/figure.png" pageId="40" pageNumber="41">Fig 19</figureCitation>
C–D). As described by
<bibRefCitation id="276A9ADC9D72CA65FD91FF1AFCFCFF23" author="Mielke, O. H. H." box="[634,799,153,180]" pageId="40" pageNumber="41" pagination="447 - 455" refId="ref43084" refString="Mielke, O. H. H. (1968) Lepidoptera do Planalto Central brasileiro. V: Novas especies de Hesperiidae e anotacoes sobre outras especies conhecidas. Revista Brasileira de Biologia, 28 (4), 447 - 455." type="journal article" year="1968">Mielke (1968)</bibRefCitation>
.
</paragraph>
<paragraph id="4344E72D9D72CA65FF2CFF3FFC98FED6" blockId="40.[151,1437,153,1634]" pageId="40" pageNumber="41">
<emphasis id="718F3B3F9D72CA65FF2CFF3FFED2FF41" box="[199,305,188,214]" italics="true" pageId="40" pageNumber="41">Genitalia</emphasis>
(
<figureCitation id="DBC0FBA89D72CA65FEAAFF3FFE74FF40" box="[321,407,188,215]" captionStart="FIGURES 61–65" captionStartId="36.[151,264,653,678]" captionTargetBox="[151,1435,178,637]" captionTargetId="figure@36.[151,1436,174,642]" captionTargetPageId="36" captionText="FIGURES 61–65. Female genitalia of the “aquilina group” and “bipunctata group” species. 61. Artines aquilina, Cananéia, São Paulo, Brazil (OM 49.831). 62. Artines angelica sp. nov., paratype, Tambopata, Puerto Maldonado, Madre de Díos, Peru (DZ 41.222) 63. Artines focus, Pacaraima, Roraima, Brazil (DZ 41.333). 64. Artines cofus sp. nov. paratype, Tambopata, Puerto Maldonado, Madre de Díos, Peru (DZ 41. 293). 65. Artines bipunctata, Reserva IBGE, Brasília, Distrito Federal, Brazil (DZ 41.165). For reference of the letters see legend of the figure 54. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241061/files/figure.png" pageId="40" pageNumber="41">Figs 65</figureCitation>
A–B, 77): sterigma rectangular. Tergum VIII with incomplete spiracular opening. Lamella antevaginalis wide and membranous, without proximo-ventral folds. Lamella postvaginalis with two sclerotized plates, one proximal near the ostium, the other distal with two distal lateral projections. Ostium D-shaped, surrounded by sclerotized area. Ductus bursae wide, basal portion sclerotized.
</paragraph>
<paragraph id="4344E72D9D72CA65FF2CFECAFE0BFE3C" blockId="40.[151,1437,153,1634]" pageId="40" pageNumber="41">
<emphasis id="718F3B3F9D72CA65FF2CFECAFED9FEF3" bold="true" box="[199,314,329,356]" pageId="40" pageNumber="41">Variation</emphasis>
. As mentioned by
<bibRefCitation id="276A9ADC9D72CA65FDF8FEC9FD56FEF2" author="Mielke, O. H. H." box="[531,693,330,357]" pageId="40" pageNumber="41" pagination="447 - 455" refId="ref43084" refString="Mielke, O. H. H. (1968) Lepidoptera do Planalto Central brasileiro. V: Novas especies de Hesperiidae e anotacoes sobre outras especies conhecidas. Revista Brasileira de Biologia, 28 (4), 447 - 455." type="journal article" year="1968">Mielke (1968)</bibRefCitation>
some specimens have a second spot/mark in the upper portion of CuA
<subScript id="DF7FE5689D72CA65FF27FEFCFF37FE1A" attach="left" box="[204,212,383,397]" fontSize="6" pageId="40" pageNumber="41">2</subScript>
–2A of the dorsal forewing, above the larger spot. The spiracular opening of tergum VIII of the female can be complete in some individuals.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D72CA65FF2CFE37FA60FD7D" pageId="40" pageNumber="41" type="discussion">
<paragraph id="4344E72D9D72CA65FF2CFE37FE2DFD33" blockId="40.[151,1437,153,1634]" pageId="40" pageNumber="41">
<emphasis id="718F3B3F9D72CA65FF2CFE37FEABFE58" bold="true" box="[199,328,436,463]" pageId="40" pageNumber="41">Comments</emphasis>
.
<taxonomicName id="84FB9CAE9D72CA65FEB1FE36FDCEFE58" authorityName="O. Mielke" authorityYear="1968" box="[346,557,437,463]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="bipunctata">
<emphasis id="718F3B3F9D72CA65FEB1FE36FDCEFE58" box="[346,557,437,463]" italics="true" pageId="40" pageNumber="41">Artines bipunctata</emphasis>
</taxonomicName>
shares several morphological characters with the species of the two groups considered before. For example, the presence of a brand on the male forewing and broad white and semi-hyaline spots on the forewing of both sexes are shared with the species of the “
<taxonomicName id="84FB9CAE9D72CA65FC77FE7FFC1AFD81" box="[924,1017,508,534]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="aquilina">
<emphasis id="718F3B3F9D72CA65FC77FE7FFC1AFD81" box="[924,1017,508,534]" italics="true" pageId="40" pageNumber="41">aquilina</emphasis>
</taxonomicName>
group”. On the other hand, characters of the male and female genitalia, as wide uncus, harpe and ampulla not fused, fultura inferior with base bifid, ventral projection of the aedeagus not divergent and sterigma distally bifid, are also present in the species of the “
<taxonomicName id="84FB9CAE9D72CA65FAA0FDC0FA7FFDCB" box="[1355,1436,579,604]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D72CA65FAA0FDC0FA7FFDCB" box="[1355,1436,579,604]" italics="true" pageId="40" pageNumber="41">aepitus</emphasis>
</taxonomicName>
group”. However, the numerous and unique characters mentioned in the diagnosis above justify the inclusion of
<taxonomicName id="84FB9CAE9D72CA65FA6FFDE5FEECFD34" authorityName="O. Mielke" authorityYear="1968" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="bipunctata">
<emphasis id="718F3B3F9D72CA65FA6FFDE5FEECFD34" italics="true" pageId="40" pageNumber="41">A. bipunctata</emphasis>
</taxonomicName>
in a group apart.
</paragraph>
<paragraph id="4344E72D9D72CA65FF2CFD2FFA60FD7D" blockId="40.[151,1437,153,1634]" pageId="40" pageNumber="41">
<emphasis id="718F3B3F9D72CA65FF2CFD2FFE09FD50" bold="true" box="[199,490,684,711]" pageId="40" pageNumber="41">Geographic distribution</emphasis>
(
<figureCitation id="DBC0FBA89D72CA65FE11FD2EFDB3FD50" box="[506,592,685,711]" captionStart="FIGURE 80" captionStartId="43.[151,250,930,955]" captionTargetBox="[151,1436,187,905]" captionTargetId="figure@43.[151,1436,187,905]" captionTargetPageId="43" captionText="FIGURE 80. Geographic distribution of the “aquilina group” and “bipunctata group” species. The symbols with white inferior half represent records from literature." httpUri="https://zenodo.org/record/3241071/files/figure.png" pageId="40" pageNumber="41">Fig. 80</figureCitation>
). This species is only known from islands of humid forests in the Cerrado biome. BRAZIL—
<emphasis id="718F3B3F9D72CA65FE87FD52FE4CFD7C" box="[364,431,721,747]" italics="true" pageId="40" pageNumber="41">Goiás</emphasis>
:
<collectingCountry id="3BECA7BD9D72CA65FE55FD52FDC0FD7C" box="[446,547,721,747]" name="Taiwan" pageId="40" pageNumber="41">Formosa</collectingCountry>
, Goianésia, Vianópolis.
<emphasis id="718F3B3F9D72CA65FCD8FD52FC09FD7D" box="[819,1002,720,746]" italics="true" pageId="40" pageNumber="41">Distrito Federal</emphasis>
: Brasília.
<emphasis id="718F3B3F9D72CA65FBB6FD52FB15FD7C" box="[1117,1270,721,747]" italics="true" pageId="40" pageNumber="41">Minas Gerais</emphasis>
: Barbacena.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D72CA65FF2CFD77FACFFC99" box="[199,1324,755,782]" pageId="40" pageNumber="41" type="description">
<paragraph id="4344E72D9D72CA65FF2CFD77FACFFC99" blockId="40.[151,1437,153,1634]" box="[199,1324,755,782]" pageId="40" pageNumber="41">
<emphasis id="718F3B3F9D72CA65FF2CFD77FE31FC99" bold="true" box="[199,466,755,782]" pageId="40" pageNumber="41">Temporal distribution</emphasis>
. This species occurs throughout the year, probably with several generations.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D72CA65FF2CFC9BFE2AFCA6" box="[199,457,791,818]" pageId="40" pageNumber="41" type="biology_ecology">
<paragraph id="4344E72D9D72CA65FF2CFC9BFE2AFCA6" blockId="40.[151,1437,153,1634]" box="[199,457,791,818]" pageId="40" pageNumber="41">
<emphasis id="718F3B3F9D72CA65FF2CFC9BFEA7FCA5" bold="true" box="[199,324,792,818]" pageId="40" pageNumber="41">Host plant</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D72CA65FF2CFCB8FEE7FCEF" pageId="40" pageNumber="41" type="etymology">
<paragraph id="4344E72D9D72CA65FF2CFCB8FEE7FCEF" blockId="40.[151,1437,153,1634]" pageId="40" pageNumber="41">
<emphasis id="718F3B3F9D72CA65FF2CFCB8FEABFCC2" bold="true" box="[199,328,827,853]" pageId="40" pageNumber="41">Etymology</emphasis>
. Although omitted in the original description, the name is a reference of the two white spots on the forewing.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D72CA65FF2CFC01FC45F9F5" pageId="40" pageNumber="41" type="materials_examined">
<paragraph id="4344E72D9D72CA65FF2CFC01FDB8FC74" blockId="40.[151,1437,153,1634]" pageId="40" pageNumber="41">
<emphasis id="718F3B3F9D72CA65FF2CFC01FE96FC0B" bold="true" box="[199,373,897,924]" pageId="40" pageNumber="41">Type material</emphasis>
.
<typeStatus id="9C40598F9D72CA65FE63FC01FE13FC0B" box="[392,496,898,924]" pageId="40" pageNumber="41" type="holotype">Holotype</typeStatus>
male deposited at the OM–DZUP with the following labels: /
<materialsCitation id="F393ED709D72CA65FB09FC01FDA6FC74" ID-GBIF-Occurrence="2248478886" collectingDate="1967-02-03" collectionCode="OM" collectorName="Brasilia, D &amp; Tangerini" country="Brazil" determinerName="Mielke" location="Brazil" pageId="40" pageNumber="41" specimenCount="1" typeStatus="holotype">
<typeStatus id="9C40598F9D72CA65FB09FC01FA6BFC0B" box="[1250,1416,898,924]" pageId="40" pageNumber="41" type="holotype">HOLOTYPUS</typeStatus>
/
<date id="3745C1ED9D72CA65FF7CFC26FEE3FC28" box="[151,256,933,959]" pageId="40" pageNumber="41" value="1967-02-03">
<collectingDate id="270138059D72CA65FF7CFC26FEE3FC28" box="[151,256,933,959]" pageId="40" pageNumber="41" value="1967-02-03">3.II.1967</collectingDate>
</date>
, J[ardim] Zoológico,
<collectorName id="EE0E82FB9D72CA65FE15FC26FD9FFC28" box="[510,636,932,959]" pageId="40" pageNumber="41">Brasília, D</collectorName>
[istrito] F[ederal,
<collectingCountry id="3BECA7BD9D72CA65FCA1FC27FC77FC28" box="[842,916,932,959]" name="Brazil" pageId="40" pageNumber="41">Brazil</collectingCountry>
],
<collectorName id="EE0E82FB9D72CA65FC42FC26FBF5FC28" box="[937,1046,933,959]" pageId="40" pageNumber="41">Tangerini</collectorName>
<emphasis id="718F3B3F9D72CA65FBCAFC26FBA9FC28" box="[1057,1098,933,959]" italics="true" pageId="40" pageNumber="41">leg.</emphasis>
/
<taxonomicName id="84FB9CAE9D72CA65FB83FC25FF31FC74" authority="Mielke, 1968" authorityName="O. Mielke" authorityYear="1968" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="bipunctata">
<emphasis id="718F3B3F9D72CA65FB83FC25FAD8FC28" box="[1128,1339,933,960]" italics="true" pageId="40" pageNumber="41">Artines bipunctata</emphasis>
<bibRefCitation id="276A9ADC9D72CA65FAADFC26FF31FC74" author="Mielke, O. H. H." pageId="40" pageNumber="41" pagination="447 - 455" refId="ref43084" refString="Mielke, O. H. H. (1968) Lepidoptera do Planalto Central brasileiro. V: Novas especies de Hesperiidae e anotacoes sobre outras especies conhecidas. Revista Brasileira de Biologia, 28 (4), 447 - 455." type="journal article" year="1968">Mielke, 1968</bibRefCitation>
</taxonomicName>
,
<determinerName id="B6980BE99D72CA65FF36FC4AFECFFC74" box="[221,300,969,995]" pageId="40" pageNumber="41">Mielke</determinerName>
det. 1968 / [
<collectionCode id="25EA7FE89D72CA65FE53FC4BFE09FC74" box="[440,490,968,995]" country="New Zealand" name="Otago Museum" pageId="40" pageNumber="41">OM</collectionCode>
] 10.948
</materialsCitation>
/.
</paragraph>
<paragraph id="4344E72D9D72CA65FF2CFC6FFEAAFBDA" blockId="40.[151,1437,153,1634]" pageId="40" pageNumber="41">
<typeStatus id="9C40598F9D72CA65FF2CFC6FFECAFB91" box="[199,297,1004,1030]" pageId="40" pageNumber="41" type="allotype">Allotype</typeStatus>
female deposited at the OM–DZUP with the following labels: /
<materialsCitation id="F393ED709D72CA65FBE0FC6FFEDFFBDA" ID-GBIF-Occurrence="2248478895" collectingDate="1967-02-03" collectionCode="OM" collectorName="Brasilia, D &amp; Tangerini" country="Brazil" determinerName="Mielke" location="Brazil" pageId="40" pageNumber="41" specimenCount="1" typeStatus="allotype">
<typeStatus id="9C40598F9D72CA65FBE0FC6FFB4DFB91" box="[1035,1198,1004,1030]" pageId="40" pageNumber="41" type="allotype">ALLOTYPUS</typeStatus>
/
<date id="3745C1ED9D72CA65FB2CFC6FFAD3FB91" box="[1223,1328,1004,1030]" pageId="40" pageNumber="41" value="1967-02-03">
<collectingDate id="270138059D72CA65FB2CFC6FFAD3FB91" box="[1223,1328,1004,1030]" pageId="40" pageNumber="41" value="1967-02-03">3.II.1967</collectingDate>
</date>
, J[ardim] Zoológico,
<collectorName id="EE0E82FB9D72CA65FEF3FB8CFE71FBBD" box="[280,402,1039,1066]" pageId="40" pageNumber="41">Brasília, D</collectorName>
[istrito] F[ederal,
<collectingCountry id="3BECA7BD9D72CA65FDBCFB8CFD42FBBD" box="[599,673,1039,1066]" name="Brazil" pageId="40" pageNumber="41">Brazil</collectingCountry>
],
<collectorName id="EE0E82FB9D72CA65FD58FB8CFCC3FBBE" box="[691,800,1039,1065]" pageId="40" pageNumber="41">Tangerini</collectorName>
<emphasis id="718F3B3F9D72CA65FCCDFB93FCB3FBBD" box="[806,848,1040,1066]" italics="true" pageId="40" pageNumber="41">leg.</emphasis>
/
<taxonomicName id="84FB9CAE9D72CA65FC8EFB93FB30FBBE" authority="Mielke, 1968" authorityName="O. Mielke" authorityYear="1968" box="[869,1235,1039,1066]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="bipunctata">
<emphasis id="718F3B3F9D72CA65FC8EFB93FBD6FBBD" box="[869,1077,1040,1066]" italics="true" pageId="40" pageNumber="41">Artines bipunctata</emphasis>
<bibRefCitation id="276A9ADC9D72CA65FBD0FB8CFB30FBBE" author="Mielke, O. H. H." box="[1083,1235,1039,1065]" pageId="40" pageNumber="41" pagination="447 - 455" refId="ref43084" refString="Mielke, O. H. H. (1968) Lepidoptera do Planalto Central brasileiro. V: Novas especies de Hesperiidae e anotacoes sobre outras especies conhecidas. Revista Brasileira de Biologia, 28 (4), 447 - 455." type="journal article" year="1968">Mielke, 1968</bibRefCitation>
</taxonomicName>
,
<determinerName id="B6980BE99D72CA65FB35FB8CFACEFBBE" box="[1246,1325,1039,1065]" pageId="40" pageNumber="41">Mielke</determinerName>
det. 1968 / [
<collectionCode id="25EA7FE89D72CA65FF46FBB1FF3CFBDA" box="[173,223,1074,1101]" country="New Zealand" name="Otago Museum" pageId="40" pageNumber="41">OM</collectionCode>
] 10.949
</materialsCitation>
/.
</paragraph>
<paragraph id="4344E72D9D72CA65FF2CFBD4FA6AFB2F" blockId="40.[151,1437,153,1634]" pageId="40" pageNumber="41">
<materialsCitation id="F393ED709D72CA65FF2CFBD4FC36FB03" ID-GBIF-Occurrence="2248478857" collectingDate="1967-02-03" collectionCode="OM" collectorName="N. Tangerini" elevation="1000" location="Jardim Zoologico de Brasilia" municipality="Brasilia" pageId="40" pageNumber="41" specimenCode="OM 10.950, OM 10.951, DZ 41.234, DZ 41.214" specimenCount="5" specimenCount-female="5" stateProvince="Distrito Federal" typeStatus="paratype">
<typeStatus id="9C40598F9D72CA65FF2CFBD4FEDBFBE6" box="[199,312,1111,1137]" pageId="40" pageNumber="41" type="paratype">Paratypes</typeStatus>
: BRAZIL—
<emphasis id="718F3B3F9D72CA65FE2FFBD4FD9FFBE7" box="[452,636,1110,1136]" italics="true" pageId="40" pageNumber="41">
<collectingRegion id="813F29CF9D72CA65FE2FFBD4FD9FFBE7" box="[452,636,1110,1136]" country="Mexico" name="Distrito Federal" pageId="40" pageNumber="41">Distrito Federal</collectingRegion>
</emphasis>
:
<emphasis id="718F3B3F9D72CA65FD60FBD5FD0BFBE6" bold="true" box="[651,744,1110,1137]" pageId="40" pageNumber="41">
<collectingMunicipality id="A3207D579D72CA65FD60FBD5FD0BFBE6" box="[651,744,1110,1137]" pageId="40" pageNumber="41">Brasília</collectingMunicipality>
</emphasis>
(
<location id="4624B1F69D72CA65FD1CFBD5FBA7FBE6" LSID="urn:lsid:plazi:treatment:CB52563B9D7DCA65FF7CFCA2FC45F9F5:4624B1F69D72CA65FD1CFBD5FBA7FBE6" box="[759,1092,1110,1137]" municipality="Brasilia" name="Jardim Zoologico de Brasilia" pageId="40" pageNumber="41" stateProvince="Distrito Federal">Jardim Zoológico de Brasília</location>
),
<quantity id="84034AC89D72CA65FBBCFBD4FB4BFBE6" box="[1111,1192,1111,1137]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="40" pageNumber="41" unit="m" value="1000.0">
<elevation id="C8D6001E9D72CA65FBBCFBD4FB4BFBE6" box="[1111,1192,1111,1137]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="40" pageNumber="41" unit="m" value="1000.0">1000m</elevation>
</quantity>
,
<specimenCount id="55FD2CA49D72CA65FB5FFBD5FAC0FBE7" box="[1204,1315,1110,1137]" pageId="40" pageNumber="41" type="female">5 females</specimenCount>
,
<date id="3745C1ED9D72CA65FAC4FBD5FA7BFBE7" box="[1327,1432,1110,1136]" pageId="40" pageNumber="41" value="1967-02-03">
<collectingDate id="270138059D72CA65FAC4FBD5FA7BFBE7" box="[1327,1432,1110,1136]" pageId="40" pageNumber="41" value="1967-02-03">3.II.1967</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D72CA65FF7CFBF8FEC6FB03" box="[151,293,1146,1173]" pageId="40" pageNumber="41">N. Tangerini</collectorName>
<emphasis id="718F3B3F9D72CA65FEC6FBF9FEB4FB03" box="[301,343,1146,1172]" italics="true" pageId="40" pageNumber="41">leg.</emphasis>
,
<specimenCode id="135D4F569D72CA65FE8DFBF9FE08FB03" box="[358,491,1145,1172]" collectionCode="OM" country="New Zealand" name="Otago Museum" pageId="40" pageNumber="41">OM 10.950</specimenCode>
,
<specimenCode id="135D4F569D72CA65FE1CFBF9FD99FB03" box="[503,634,1145,1172]" collectionCode="OM" country="New Zealand" name="Otago Museum" pageId="40" pageNumber="41">OM 10.951</specimenCode>
(
<collectionCode id="25EA7FE89D72CA65FD61FBF9FD5FFB03" box="[650,700,1146,1172]" country="New Zealand" name="Otago Museum" pageId="40" pageNumber="41">OM</collectionCode>
),
<specimenCode id="135D4F569D72CA65FD3AFBF8FCADFB03" box="[721,846,1146,1173]" collectionCode="OM" country="New Zealand" name="Otago Museum" pageId="40" pageNumber="41">DZ 41.234</specimenCode>
,
<specimenCode id="135D4F569D72CA65FCB0FBF8FC36FB03" box="[859,981,1146,1173]" collectionCode="OM" country="New Zealand" name="Otago Museum" pageId="40" pageNumber="41">DZ 41.214</specimenCode>
</materialsCitation>
<materialsCitation id="F393ED709D72CA65FC36FBF8FA66FB2F" ID-GBIF-Occurrence="2248478859" collectingDate="1967-01-27" collectionCode="DZUP, OM" collectorName="N. Tangerini" location="Barbacena" pageId="40" pageNumber="41" specimenCode="OM 10.947" specimenCount="1" specimenCount-male="1" stateProvince="Minas Gerais" typeStatus="paratype">
DZ 41.244 (Ex. collection
<collectorName id="EE0E82FB9D72CA65FAE5FBF8FA7FFB03" box="[1294,1436,1146,1173]" pageId="40" pageNumber="41">N. Tangerini</collectorName>
now deposited at the
<collectionCode id="25EA7FE89D72CA65FE6EFB1DFE30FB2F" box="[389,467,1182,1208]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="40" pageNumber="41" type="University or college">DZUP</collectionCode>
).
<emphasis id="718F3B3F9D72CA65FE0DFB1DFD9CFB2F" box="[486,639,1182,1208]" italics="true" pageId="40" pageNumber="41">
<collectingRegion id="813F29CF9D72CA65FE0DFB1DFD9CFB2F" box="[486,639,1182,1208]" country="Brazil" name="Minas Gerais" pageId="40" pageNumber="41">Minas Gerais</collectingRegion>
</emphasis>
:
<emphasis id="718F3B3F9D72CA65FD65FB1DFCF6FB2F" bold="true" box="[654,789,1182,1208]" pageId="40" pageNumber="41">
<location id="4624B1F69D72CA65FD65FB1DFCF2FB2F" LSID="urn:lsid:plazi:treatment:CB52563B9D7DCA65FF7CFCA2FC45F9F5:4624B1F69D72CA65FD65FB1DFCF2FB2F" box="[654,785,1182,1208]" name="Barbacena" pageId="40" pageNumber="41" stateProvince="Minas Gerais">Barbacena</location>
,
</emphasis>
<specimenCount id="55FD2CA49D72CA65FCF6FB1DFC88FB20" box="[797,875,1181,1208]" pageId="40" pageNumber="41" type="male">1 male</specimenCount>
,
<date id="3745C1ED9D72CA65FC9DFB1EFC07FB20" box="[886,996,1181,1207]" pageId="40" pageNumber="41" value="1967-01-27">
<collectingDate id="270138059D72CA65FC9DFB1EFC07FB20" box="[886,996,1181,1207]" pageId="40" pageNumber="41" value="1967-01-27">27.I.1967</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D72CA65FC04FB1DFB9FFB20" box="[1007,1148,1181,1208]" pageId="40" pageNumber="41">N. Tangerini</collectorName>
<emphasis id="718F3B3F9D72CA65FB68FB1EFB4FFB20" box="[1155,1196,1181,1207]" italics="true" pageId="40" pageNumber="41">leg.</emphasis>
,
<specimenCode id="135D4F569D72CA65FB51FB1DFAD8FB2F" box="[1210,1339,1182,1208]" collectionCode="OM" country="New Zealand" name="Otago Museum" pageId="40" pageNumber="41">OM 10.947</specimenCode>
(
<collectionCode id="25EA7FE89D72CA65FAA0FB1DFA9DFB2F" box="[1355,1406,1182,1208]" country="New Zealand" name="Otago Museum" pageId="40" pageNumber="41">OM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="4344E72D9D72CA65FF2CFB43FC45F9F5" blockId="40.[151,1437,153,1634]" pageId="40" pageNumber="41">
<emphasis id="718F3B3F9D72CA65FF2CFB43FE4EFB4C" bold="true" box="[199,429,1216,1243]" pageId="40" pageNumber="41">Examined material</emphasis>
. The
<typeStatus id="9C40598F9D72CA65FE04FB42FDFCFB4C" box="[495,543,1217,1243]" pageId="40" pageNumber="41">type</typeStatus>
material mentioned above and the following specimens: BRAZIL—
<emphasis id="718F3B3F9D72CA65FACDFB42FA8AFB4C" box="[1318,1385,1217,1243]" italics="true" pageId="40" pageNumber="41">Goiás</emphasis>
:
<emphasis id="718F3B3F9D72CA65FA93FB43FF18FB68" bold="true" pageId="40" pageNumber="41">Vianópolis</emphasis>
(Ponte Funda),
<specimenCount id="55FD2CA49D72CA65FE53FB67FDE9FB69" box="[440,522,1252,1278]" pageId="40" pageNumber="41" type="male">1 male</specimenCount>
,
<date id="3745C1ED9D72CA65FDF3FB67FD66FB69" box="[536,645,1252,1278]" pageId="40" pageNumber="41" value="1981-01-17">17.I.1981</date>
, N. Tangerini
<emphasis id="718F3B3F9D72CA65FCC5FB67FCBBFB69" box="[814,856,1252,1278]" italics="true" pageId="40" pageNumber="41">leg.</emphasis>
, DZ 41.264 (
<collectionCode id="25EA7FE89D72CA65FC1CFB67FBA5FB69" box="[1015,1094,1252,1278]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="40" pageNumber="41" type="University or college">DZUP</collectionCode>
);
<materialsCitation id="F393ED709D72CA65FBB6FB66FD6FFAB5" ID-GBIF-Occurrence="2248478861" collectingDate="1977-05-15" collectionCode="DZUP" collectorName="Itiquira &amp; Gifford" country="Taiwan" location="Formosa" pageId="40" pageNumber="41" specimenCount="1" specimenCount-male="1">
<emphasis id="718F3B3F9D72CA65FBB6FB66FB24FB68" bold="true" box="[1117,1223,1253,1279]" pageId="40" pageNumber="41">
<collectingCountry id="3BECA7BD9D72CA65FBB6FB66FB24FB68" box="[1117,1223,1253,1279]" name="Taiwan" pageId="40" pageNumber="41">Formosa</collectingCountry>
</emphasis>
(
<collectorName id="EE0E82FB9D72CA65FB33FB67FAD3FB68" box="[1240,1328,1252,1279]" pageId="40" pageNumber="41">Itiquira</collectorName>
),
<specimenCount id="55FD2CA49D72CA65FAACFB67FA7BFB69" box="[1351,1432,1252,1278]" pageId="40" pageNumber="41" type="male">1 male</specimenCount>
,
<date id="3745C1ED9D72CA65FF7CFA84FEEEFAB5" box="[151,269,1287,1314]" pageId="40" pageNumber="41" value="1977-05-15">
<collectingDate id="270138059D72CA65FF7CFA84FEEEFAB5" box="[151,269,1287,1314]" pageId="40" pageNumber="41" value="1977-05-15">15.V.1977</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D72CA65FEF3FA8BFE88FAB5" box="[280,363,1288,1314]" pageId="40" pageNumber="41">Gifford</collectorName>
<emphasis id="718F3B3F9D72CA65FE98FA8BFE7EFAB5" box="[371,413,1288,1314]" italics="true" pageId="40" pageNumber="41">leg.</emphasis>
, DZ 41.155 (
<collectionCode id="25EA7FE89D72CA65FDDFFA8BFD60FAB5" box="[564,643,1288,1314]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="40" pageNumber="41" type="University or college">DZUP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F393ED709D72CA65FD7CFA84FA82FAB5" ID-GBIF-Occurrence="2248478854" box="[663,1377,1287,1314]" collectingDate="1969-09" collectionCode="DZUP" collectorName="Goianesia &amp; H. Ebert" country="Taiwan" pageId="40" pageNumber="41" specimenCount="1" specimenCount-male="1">
<emphasis id="718F3B3F9D72CA65FD7CFA84FCEEFAB5" bold="true" box="[663,781,1287,1314]" pageId="40" pageNumber="41">
<collectorName id="EE0E82FB9D72CA65FD7CFA84FCEEFAB5" box="[663,781,1287,1314]" pageId="40" pageNumber="41">Goianésia</collectorName>
</emphasis>
,
<specimenCount id="55FD2CA49D72CA65FCF7FA8BFC88FAB5" box="[796,875,1288,1314]" pageId="40" pageNumber="41" type="male">1 male</specimenCount>
,
<date id="3745C1ED9D72CA65FC9CFA8BFC35FAB5" box="[887,982,1288,1314]" pageId="40" pageNumber="41" value="1969-09">
<collectingDate id="270138059D72CA65FC9CFA8BFC35FAB5" box="[887,982,1288,1314]" pageId="40" pageNumber="41" value="1969-09">IX.1969</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D72CA65FC09FA8BFBA2FAB5" box="[994,1089,1288,1314]" pageId="40" pageNumber="41">H. Ebert</collectorName>
<emphasis id="718F3B3F9D72CA65FBA2FA8BFB91FAB5" box="[1097,1138,1288,1314]" italics="true" pageId="40" pageNumber="41">leg.</emphasis>
, DZ 41.284 (
<collectionCode id="25EA7FE89D72CA65FAE1FA8BFAB9FAB5" box="[1290,1370,1288,1314]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="40" pageNumber="41" type="University or college">DZUP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="F393ED709D72CA65FA87FA8AFE77FAFD" ID-GBIF-Occurrence="2248478874" collectingDate="2006-03-27" collectionCode="DZUP" collectorName="Distrito Federal &amp; Brasilia &amp; Corrego Bananal &amp; E. O. Emery" country="Taiwan" location="Jardim Zoologico de Brasilia" pageId="40" pageNumber="41" specimenCount="1" specimenCount-female="1">
<emphasis id="718F3B3F9D72CA65FA87FA8AFECAFAD2" italics="true" pageId="40" pageNumber="41">
<collectorName id="EE0E82FB9D72CA65FA87FA8AFECAFAD2" pageId="40" pageNumber="41">Distrito Federal</collectorName>
</emphasis>
:
<emphasis id="718F3B3F9D72CA65FED2FAA9FE75FAD2" bold="true" box="[313,406,1322,1349]" pageId="40" pageNumber="41">
<collectorName id="EE0E82FB9D72CA65FED2FAA9FE75FAD2" box="[313,406,1322,1349]" pageId="40" pageNumber="41">Brasília</collectorName>
</emphasis>
, (
<collectorName id="EE0E82FB9D72CA65FE46FAA8FD90FAD1" box="[429,627,1323,1350]" pageId="40" pageNumber="41">Córrego Bananal</collectorName>
),
<specimenCount id="55FD2CA49D72CA65FD63FAA8FD0DFAD2" box="[648,750,1323,1349]" pageId="40" pageNumber="41" type="female">1 female</specimenCount>
,
<date id="3745C1ED9D72CA65FD11FAA8FC98FAD2" box="[762,891,1323,1349]" pageId="40" pageNumber="41" value="2006-03-27">
<collectingDate id="270138059D72CA65FD11FAA8FC98FAD2" box="[762,891,1323,1349]" pageId="40" pageNumber="41" value="2006-03-27">27.III.2006</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D72CA65FC63FAAFFBF4FAD1" box="[904,1047,1323,1350]" pageId="40" pageNumber="41">E. O. Emery</collectorName>
<emphasis id="718F3B3F9D72CA65FBF4FAA8FBAAFAD2" box="[1055,1097,1323,1349]" italics="true" pageId="40" pageNumber="41">leg.</emphasis>
, DZ 41.224 (
<collectionCode id="25EA7FE89D72CA65FB08FAA8FAD1FAD2" box="[1251,1330,1323,1349]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="40" pageNumber="41" type="University or college">DZUP</collectionCode>
), (
<location id="4624B1F69D72CA65FAA5FAA8FE6FFAFD" LSID="urn:lsid:plazi:treatment:CB52563B9D7DCA65FF7CFCA2FC45F9F5:4624B1F69D72CA65FAA5FAA8FE6FFAFD" country="Taiwan" name="Jardim Zoologico de Brasilia" pageId="40" pageNumber="41">Jardim Zoológico de Brasília</location>
)
</materialsCitation>
,
<quantity id="84034AC89D72CA65FE75FACCFE0DFAFE" box="[414,494,1359,1385]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="40" pageNumber="41" unit="m" value="1000.0">1000m</quantity>
,
<materialsCitation id="F393ED709D72CA65FE12FACCFB37FAFE" ID-GBIF-Occurrence="2248478862" box="[505,1236,1359,1385]" collectingDate="1972-06-19" collectionCode="VI, DZUP" collectorName="O. Mielke &amp; Brown" country="Taiwan" pageId="40" pageNumber="41" specimenCount="1" specimenCount-male="1">
<specimenCount id="55FD2CA49D72CA65FE12FACCFDA5FAFE" box="[505,582,1359,1385]" pageId="40" pageNumber="41" type="male">1 male</specimenCount>
,
<date id="3745C1ED9D72CA65FDBBFACCFD30FAFE" box="[592,723,1359,1385]" pageId="40" pageNumber="41" value="1972-06-19">
<collectingDate id="270138059D72CA65FDBBFACCFD30FAFE" box="[592,723,1359,1385]" pageId="40" pageNumber="41" value="1972-06-19">
19.
<collectionCode id="25EA7FE89D72CA65FD98FACCFD70FAFE" box="[627,659,1359,1385]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="40" pageNumber="41">VI</collectionCode>
.1972
</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D72CA65FD37FACCFCAEFAFE" box="[732,845,1359,1385]" pageId="40" pageNumber="41">O. Mielke</collectorName>
&amp;
<collectorName id="EE0E82FB9D72CA65FC84FACCFC58FAFE" box="[879,955,1359,1385]" pageId="40" pageNumber="41">Brown</collectorName>
<emphasis id="718F3B3F9D72CA65FC2AFACCFC00FAFE" box="[961,995,1359,1385]" italics="true" pageId="40" pageNumber="41">leg</emphasis>
., DZ 30.961 (
<collectionCode id="25EA7FE89D72CA65FB96FACCFB2FFAFE" box="[1149,1228,1359,1385]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="40" pageNumber="41" type="University or college">DZUP</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="F393ED709D72CA65FB35FACCFB7AFA1A" ID-GBIF-Occurrence="2248478894" collectingDate="1967-02-03" collectionCode="OM" collectorName="N. Tangerini" country="Taiwan" location="Parque Nacional de Brasilia" pageId="40" pageNumber="41" specimenCode="OM 10.948, OM 10.949" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="55FD2CA49D72CA65FB35FACCFAC8FAFE" box="[1246,1323,1359,1385]" pageId="40" pageNumber="41" type="male">1 male</specimenCount>
,
<specimenCount id="55FD2CA49D72CA65FADEFACCFA7BFAFE" box="[1333,1432,1359,1385]" pageId="40" pageNumber="41" type="female">1 female</specimenCount>
,
<date id="3745C1ED9D72CA65FF7CFAF1FEE3FA1B" box="[151,256,1394,1420]" pageId="40" pageNumber="41" value="1967-02-03">
<collectingDate id="270138059D72CA65FF7CFAF1FEE3FA1B" box="[151,256,1394,1420]" pageId="40" pageNumber="41" value="1967-02-03">3.II.1967</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D72CA65FEE6FAF0FE78FA1B" box="[269,411,1394,1421]" pageId="40" pageNumber="41">N. Tangerini</collectorName>
<emphasis id="718F3B3F9D72CA65FE48FAF1FE2FFA1B" box="[419,460,1394,1420]" italics="true" pageId="40" pageNumber="41">leg.</emphasis>
,
<specimenCode id="135D4F569D72CA65FE37FAF1FD82FA1B" box="[476,609,1394,1420]" collectionCode="OM" country="New Zealand" name="Otago Museum" pageId="40" pageNumber="41">OM 10.948</specimenCode>
,
<specimenCode id="135D4F569D72CA65FD86FAF1FD0CFA1B" box="[621,751,1394,1420]" collectionCode="OM" country="New Zealand" name="Otago Museum" pageId="40" pageNumber="41">OM 10.949</specimenCode>
(
<collectionCode id="25EA7FE89D72CA65FCEBFAF1FCD1FA1B" box="[768,818,1394,1420]" country="New Zealand" name="Otago Museum" pageId="40" pageNumber="41">OM</collectionCode>
), (
<location id="4624B1F69D72CA65FCA5FAF1FB72FA1A" LSID="urn:lsid:plazi:treatment:CB52563B9D7DCA65FF7CFCA2FC45F9F5:4624B1F69D72CA65FCA5FAF1FB72FA1A" box="[846,1169,1394,1421]" country="Taiwan" name="Parque Nacional de Brasilia" pageId="40" pageNumber="41">Parque Nacional de Brasília</location>
)
</materialsCitation>
,
<materialsCitation id="F393ED709D72CA65FB4DFAF1FD39FA27" ID-GBIF-Occurrence="2248478899" collectingDate="1977-03-31" collectionCode="DZUP, IBGE" collectorName="Gifford" country="Taiwan" location="Reserva" pageId="40" pageNumber="41" specimenCount="1" specimenCount-female="1">
<specimenCount id="55FD2CA49D72CA65FB4DFAF1FAE8FA1B" box="[1190,1291,1394,1420]" pageId="40" pageNumber="41" type="female">1 female</specimenCount>
,
<date id="3745C1ED9D72CA65FAFCFAF1FA7BFA1B" box="[1303,1432,1394,1420]" pageId="40" pageNumber="41" value="1977-03-31">
<collectingDate id="270138059D72CA65FAFCFAF1FA7BFA1B" box="[1303,1432,1394,1420]" pageId="40" pageNumber="41" value="1977-03-31">31.III.1977</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D72CA65FF7CFA15FF09FA27" box="[151,234,1430,1456]" pageId="40" pageNumber="41">Gifford</collectorName>
<emphasis id="718F3B3F9D72CA65FF1FFA15FEFDFA27" box="[244,286,1430,1456]" italics="true" pageId="40" pageNumber="41">leg.</emphasis>
, DZ 30.960 (
<collectionCode id="25EA7FE89D72CA65FE50FA15FDE9FA27" box="[443,522,1430,1456]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="40" pageNumber="41" type="University or college">DZUP</collectionCode>
), (
<location id="4624B1F69D72CA65FDCCFA15FD60FA27" LSID="urn:lsid:plazi:treatment:CB52563B9D7DCA65FF7CFCA2FC45F9F5:4624B1F69D72CA65FDCCFA15FD60FA27" box="[551,643,1430,1456]" country="Taiwan" name="Reserva" pageId="40" pageNumber="41">Reserva</location>
<collectionCode id="25EA7FE89D72CA65FD66FA15FD30FA27" box="[653,723,1430,1456]" country="Brazil" lsid="urn:lsid:biocol.org:col:15833" name="Reserva Ecológica do IBGE" pageId="40" pageNumber="41" type="Herbarium">IBGE</collectionCode>
)
</materialsCitation>
,
<specimenCount id="55FD2CA49D72CA65FD03FA15FCDAFA27" box="[744,825,1430,1456]" pageId="40" pageNumber="41" type="male">1 male</specimenCount>
,
<date id="3745C1ED9D72CA65FCACFA16FC54FA27" box="[839,951,1429,1456]" pageId="40" pageNumber="41" value="2002-04-05">5.IV.2002</date>
,
<specimenCount id="55FD2CA49D72CA65FC2EFA15FBCFFA27" box="[965,1068,1430,1456]" pageId="40" pageNumber="41" type="female">1 female</specimenCount>
,
<date id="3745C1ED9D72CA65FBD1FA15FB26FA27" box="[1082,1221,1430,1456]" pageId="40" pageNumber="41" value="2002-07-20">20.VII.2002</date>
,
<specimenCount id="55FD2CA49D72CA65FB38FA15FAC7FA27" box="[1235,1316,1430,1456]" pageId="40" pageNumber="41" type="male">1 male</specimenCount>
,
<specimenCount id="55FD2CA49D72CA65FAD9FA15FA7BFA27" box="[1330,1432,1430,1456]" pageId="40" pageNumber="41" type="female">1 female</specimenCount>
,
<date id="3745C1ED9D72CA65FF7CFA3AFEC0FA44" box="[151,291,1464,1491]" pageId="40" pageNumber="41" value="2005-07-20">20.VII.2005</date>
,
<materialsCitation id="F393ED709D72CA65FEC4FA3AFF10FA60" ID-GBIF-Occurrence="2248478867" collectingDate="2005-08-23" collectionCode="DZUP" collectorName="E. O. Emery" country="Taiwan" pageId="40" pageNumber="41" specimenCount="1" specimenCount-male="1">
<specimenCount id="55FD2CA49D72CA65FEC4FA3AFE9DFA44" box="[303,382,1465,1491]" pageId="40" pageNumber="41" type="male">1 male</specimenCount>
,
<date id="3745C1ED9D72CA65FE60FA3AFDC3FA44" box="[395,544,1464,1491]" pageId="40" pageNumber="41" value="2005-08-23">
<collectingDate id="270138059D72CA65FE60FA3AFDC3FA44" box="[395,544,1464,1491]" pageId="40" pageNumber="41" value="2005-08-23">23.VIII.2005</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D72CA65FDC7FA39FD58FA43" box="[556,699,1465,1492]" pageId="40" pageNumber="41">E. O. Emery</collectorName>
<emphasis id="718F3B3F9D72CA65FD28FA3AFD10FA44" box="[707,755,1465,1491]" italics="true" pageId="40" pageNumber="41">leg.,</emphasis>
DZ 41.254, DZ 41.334, DZ 41.324, DZ 41.165, DZ 41.274 (
<collectionCode id="25EA7FE89D72CA65FF74FA5EFF0FFA60" box="[159,236,1501,1527]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="40" pageNumber="41" type="University or college">DZUP</collectionCode>
)
</materialsCitation>
<materialsCitation id="F393ED709D72CA65FF10FA5EFB0DFA60" ID-GBIF-Occurrence="2248478871" box="[251,1262,1501,1527]" collectingDate="1968-09-07" collectionCode="DZUP" collectorName="H. Ebert" country="Taiwan" pageId="40" pageNumber="41" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
<specimenCount id="55FD2CA49D72CA65FF10FA5EFEA8FA60" box="[251,331,1501,1527]" pageId="40" pageNumber="41" type="male">1 male</specimenCount>
,
<specimenCount id="55FD2CA49D72CA65FEB3FA5EFE2BFA60" box="[344,456,1501,1527]" pageId="40" pageNumber="41" type="female">2 females</specimenCount>
,
<date id="3745C1ED9D72CA65FE3EFA5EFDAAFA60" box="[469,585,1501,1527]" pageId="40" pageNumber="41" value="1968-09-07">
<collectingDate id="270138059D72CA65FE3EFA5EFDAAFA60" box="[469,585,1501,1527]" pageId="40" pageNumber="41" value="1968-09-07">7.IX.1968</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D72CA65FDBDFA5EFD55FA60" box="[598,694,1501,1527]" pageId="40" pageNumber="41">H. Ebert</collectorName>
<emphasis id="718F3B3F9D72CA65FD55FA5EFD0BFA60" box="[702,744,1501,1527]" italics="true" pageId="40" pageNumber="41">leg.</emphasis>
, DZ 41.194, DZ 41.184, DZ 41.204 (
<collectionCode id="25EA7FE89D72CA65FB7DFA5EFB05FA60" box="[1174,1254,1501,1527]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="40" pageNumber="41" type="University or college">DZUP</collectionCode>
)
</materialsCitation>
.
<emphasis id="718F3B3F9D72CA65FB11FA5EFA77FA60" box="[1274,1428,1501,1527]" italics="true" pageId="40" pageNumber="41">Minas Gerais</emphasis>
:
<emphasis id="718F3B3F9D72CA65FF7CF983FEFBF98D" bold="true" box="[151,280,1536,1562]" pageId="40" pageNumber="41">Barbacena</emphasis>
,
<quantity id="84034AC89D72CA65FEC2F983FE9AF98D" box="[297,377,1536,1562]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="40" pageNumber="41" unit="m" value="1100.0">1100m</quantity>
,
<specimenCount id="55FD2CA49D72CA65FE6CF983FE3BF98D" box="[391,472,1536,1562]" pageId="40" pageNumber="41" type="male">1 male</specimenCount>
,
<date id="3745C1ED9D72CA65FE0DF983FD98F98D" box="[486,635,1535,1562]" pageId="40" pageNumber="41" value="1951-08-16">16.VIII.1951</date>
,
<specimenCount id="55FD2CA49D72CA65FD62F983FD39F98D" box="[649,730,1536,1562]" pageId="40" pageNumber="41" type="male">1 male</specimenCount>
,
<date id="3745C1ED9D72CA65FD02F983FCBFF98D" box="[745,860,1535,1562]" pageId="40" pageNumber="41" value="1951-09-04">4.IX.1951</date>
,
<specimenCount id="55FD2CA49D72CA65FC80F983FC5FF98D" box="[875,956,1536,1562]" pageId="40" pageNumber="41" type="male">1 male</specimenCount>
,
<date id="3745C1ED9D72CA65FC21F983FBA3F98D" box="[970,1088,1535,1563]" pageId="40" pageNumber="41" value="1952-05-29">29.V.1952</date>
,
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<collectingDate id="270138059D72CA65FE4CF9A7FDCAF9A9" box="[423,553,1571,1598]" pageId="40" pageNumber="41" value="1952-11-14">14.XI.1952</collectingDate>
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<emphasis id="718F3B3F9D72CA65FD71F9A7FD27F9A9" box="[666,708,1572,1598]" italics="true" pageId="40" pageNumber="41">leg.</emphasis>
, DZ 41.304, DZ 41.145, DZ 41.314, DZ 41.254, DZ 41.294, DZ 41.175 (
<collectionCode id="25EA7FE89D72CA65FF18F9CBFEA1F9F5" box="[243,322,1608,1634]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="40" pageNumber="41" type="University or college">DZUP</collectionCode>
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<materialsCitation id="F393ED709D72CA65FEBEF9CBFC41F9F5" ID-GBIF-Occurrence="2248478889" box="[341,930,1607,1634]" collectingDate="1990-05-19" collectionCode="OM" collectorName="N. Alves" country="Taiwan" pageId="40" pageNumber="41" specimenCode="OM 9.826" specimenCount="1" specimenCount-female="1">
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<collectingDate id="270138059D72CA65FE2FF9CBFDDAF9F5" box="[452,569,1608,1634]" pageId="40" pageNumber="41" value="1990-05-19">19.V.1990</collectingDate>
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<emphasis id="718F3B3F9D72CA65FD45F9CBFD3DF9F5" box="[686,734,1608,1634]" italics="true" pageId="40" pageNumber="41">leg.,</emphasis>
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101
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,
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<mods:detail id="0D2E1086FFB4104CD213C771DF7EDD06" type="volume">
<mods:number id="B9CC423CB267678190730A6DB554F2DB">4614</mods:number>
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<mods:classification id="CD21FEA8C9B0F290411019921477FE25">journal article</mods:classification>
<mods:identifier id="E54377AFC304F1D157CDB4451290C716" type="CLB-Dataset">26561</mods:identifier>
<mods:identifier id="5D4DE585D23CD1585972906A18E5E678" type="DOI">10.11646/zootaxa.4614.1.1</mods:identifier>
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<mods:identifier id="4975303EDCDA89BE424E913C18D0A108" type="Zenodo-Dep">3241035</mods:identifier>
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<treatment id="CB52563B9D7ECA6AFF7CFA4BFA86FE44" ID-GBIF-Taxon="157435998" LSID="urn:lsid:zoobank.org:act:5EA37EFE-671F-4C96-B8B5-130B435F3922" httpUri="http://treatment.plazi.org/id/CB52563B9D7ECA6AFF7CFA4BFA86FE44" lastPageId="39" lastPageNumber="40" pageId="36" pageNumber="37">
<subSubSection id="0BE1B4A69D7ECA69FF7CFA4BFD56FA75" box="[151,693,1480,1507]" pageId="36" pageNumber="37" type="nomenclature">
<paragraph id="4344E72D9D7ECA69FF7CFA4BFD56FA75" blockId="36.[151,956,1480,1578]" box="[151,693,1480,1507]" pageId="36" pageNumber="37">
<heading id="180C50419D7ECA69FF7CFA4BFD56FA75" bold="true" box="[151,693,1480,1507]" fontSize="11" level="1" pageId="36" pageNumber="37" reason="1">
<emphasis id="718F3B3F9D7ECA69FF7CFA4BFD56FA75" bold="true" box="[151,693,1480,1507]" pageId="36" pageNumber="37">
<taxonomicName id="84FB9CAE9D7ECA69FF7CFA4BFDB3FA75" authority="Medeiros &amp; Carneiro" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[151,592,1480,1507]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="donia" status="sp. nov.">
<emphasis id="718F3B3F9D7ECA69FF7CFA4BFED8FA74" bold="true" box="[151,315,1480,1507]" italics="true" pageId="36" pageNumber="37">Artines donia</emphasis>
Medeiros &amp; Carneiro
</taxonomicName>
<taxonomicNameLabel id="6ABC86449D7ECA69FDBCFA4BFD56FA75" box="[599,693,1480,1506]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D7ECA69FF7CFA6FFC5FF9BD" pageId="36" pageNumber="37" type="description">
<paragraph id="4344E72D9D7ECA69FF7CFA6FFE71F991" blockId="36.[151,956,1480,1578]" box="[151,402,1515,1542]" pageId="36" pageNumber="37">
<figureCitation id="DBC0FBA89D7ECA69FF7CFA6FFF0FF991" box="[151,236,1516,1542]" captionStart="FIGURES 14–19" captionStartId="10.[151,264,1406,1431]" captionTargetBox="[155,1435,175,1386]" captionTargetId="figure@10.[151,1436,174,1394]" captionTargetPageId="10" captionText="FIGURES 14–19. Species of the “aquilina group” (14–18) and “bipunctata group” (19) in dorsal and ventral views. 14. Artines aquilina, A–B: male, Joinville, Santa Catarina, Brazil (DZ 41.321), C–D: female from same locality of male (DZ 41.220). 15. Artines angelica sp. nov., A–B: holotype male, C–D: allotype female. 16. Artines focus, A–B: male, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.233), C–D: female, Pacaraima, Roraima, Brazil (DZ 41.333). 17. Artines cofus sp. nov., A–B: holotype male, C–D: allotype female. 18. Artines donia sp. nov., A–B: holotype male. 19. Artines bipunctata, A–B: male, Jardim Zoológico, Brasília, Distrito Federal, Brazil (DZ 30.961), C–D: female, Córrego Bananal, Brasília, Distrito Federal, Brazil (DZ 41.224). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241045/files/figure.png" pageId="36" pageNumber="37">Figs 18</figureCitation>
,
<figureCitation id="DBC0FBA89D7ECA69FF13FA68FEF5F991" box="[248,278,1515,1542]" captionStart="FIGURES 21–26" captionStartId="22.[151,264,1840,1865]" captionTargetBox="[151,1436,922,1828]" captionTargetId="figure@22.[151,1436,922,1828]" captionText="FIGURES 21–26. Male brands (no scales) on dorsal forewing of the “aquilina group” (21–25) and “bipunctata group” species (26). 21. Artines aquilina, Joinville, Santa catarina, Brazil, (DZ 41.341). 22. Artines angelica sp. nov., paratype, Fazenda Ter- rasse, Santa Luzia, Maranhão, Brazil (DZ 41.273) 23. Artines focus, Ilha de Maracá, Roraima, Brazil (DZ 41.233). 24. Artines cofus sp. nov., paratype, Santa Rosa do Purus, Acre, Brazil (DZ 41.303). 25. Artines donia sp. nov., paratype, Santa Rita do Araguaia, Goiás, Brazil (OM 66.000). 26. Artines bipunctata, Ponte Funda, Vianópilis, Goiás, Brazil (DZ 41.264). Scale bar = 1 mm." httpUri="https://zenodo.org/record/3406270/files/figure.png" pageId="36" pageNumber="37">25</figureCitation>
,
<figureCitation id="DBC0FBA89D7ECA69FEC9FA6FFEA2F991" box="[290,321,1516,1542]" captionStart="FIGURES 40–41" captionStartId="31.[151,264,1624,1649]" captionTargetBox="[154,1435,484,1599]" captionTargetId="figure@31.[151,1436,481,1612]" captionText="FIGURES 40–41. Male genitalia of the “aquilina group” (40) and “bipunctata group” (41) species. 40. Artines donia sp. nov., paratype, Santa Rita do Araguaia, Goiás, Brazil (DZ 66.000). 41. Artines bipunctata, Brasília, Distrito Federal, Brazil (DZ 41.194). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3406272/files/figure.png" pageId="36" pageNumber="37">40</figureCitation>
, 53, 80
</paragraph>
<paragraph id="4344E72D9D7ECA69FF7CF98CFC5FF9BD" blockId="36.[151,956,1480,1578]" box="[151,956,1551,1578]" pageId="36" pageNumber="37">
<uri id="376AEB2F9D7ECA69FF7CF98CFC5FF9BD" box="[151,956,1551,1578]" pageId="36" pageNumber="37">
urn:lsid:zoobank.org:act:
<uuid id="375DDDF89D7ECA69FE40F98CFC5FF9BD" box="[427,956,1551,1578]" pageId="36" pageNumber="37">5EA37EFE-671F-4C96-B8B5-130B435F3922</uuid>
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</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D7ECA69FF7CF9CAFA62F963" pageId="36" pageNumber="37" type="diagnosis">
<paragraph id="4344E72D9D7ECA69FF7CF9CAFA62F963" blockId="36.[151,1437,1609,2037]" pageId="36" pageNumber="37">
<emphasis id="718F3B3F9D7ECA69FF7CF9CAFEE9F9F3" bold="true" box="[151,266,1609,1636]" pageId="36" pageNumber="37">Diagnosis</emphasis>
.
<taxonomicName id="84FB9CAE9D7ECA69FEF7F9C9FE54F9F4" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[284,439,1609,1636]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="donia" status="sp. nov.">
<emphasis id="718F3B3F9D7ECA69FEF7F9C9FE54F9F4" box="[284,439,1609,1636]" italics="true" pageId="36" pageNumber="37">Artines donia</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D7ECA69FE2AF9C9FDFCF9F3" bold="true" box="[449,543,1610,1636]" pageId="36" pageNumber="37">
<taxonomicNameLabel id="6ABC86449D7ECA69FE2AF9C9FDFCF9F3" box="[449,543,1610,1636]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from the species of the “
<taxonomicName id="84FB9CAE9D7ECA69FC4BF9CAFC1EF9F4" box="[928,1021,1609,1635]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="aquilina">
<emphasis id="718F3B3F9D7ECA69FC4BF9CAFC1EF9F4" box="[928,1021,1609,1635]" italics="true" pageId="36" pageNumber="37">aquilina</emphasis>
</taxonomicName>
group” in the following characters: ventral hind wing with central black spot wider than long; uncus distally wide, without ventral projection, apex weakly bifid; valva distally rhomboid; aedeagus uniformly wide; ventral projection of aedeagus with distal portion bifid; cornutus similar to
<taxonomicName id="84FB9CAE9D7ECA69FE2AF935FDFDF947" box="[449,542,1718,1744]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="focus">
<emphasis id="718F3B3F9D7ECA69FE2AF935FDFDF947" box="[449,542,1718,1744]" italics="true" pageId="36" pageNumber="37">A. focus</emphasis>
</taxonomicName>
and
<taxonomicName id="84FB9CAE9D7ECA69FDB0F935FD5BF947" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[603,696,1718,1744]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="cofus" status="sp. nov.">
<emphasis id="718F3B3F9D7ECA69FDB0F935FD5BF947" box="[603,696,1718,1744]" italics="true" pageId="36" pageNumber="37">A. cofus</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D7ECA69FD29F935FCC3F947" bold="true" box="[706,800,1718,1744]" pageId="36" pageNumber="37">
<taxonomicNameLabel id="6ABC86449D7ECA69FD29F935FCC3F947" box="[706,800,1718,1744]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, differing in being more sclerotized and having spines slightly conspicuous; fultura inferior robust and laterally wider than in the other species of the “
<taxonomicName id="84FB9CAE9D7ECA69FB2CF95AFAC7F964" box="[1223,1316,1753,1779]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="aquilina">
<emphasis id="718F3B3F9D7ECA69FB2CF95AFAC7F964" box="[1223,1316,1753,1779]" italics="true" pageId="36" pageNumber="37">aquilina</emphasis>
</taxonomicName>
group”.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D7ECA6BFF2CF97EFE06F8BD" lastPageId="38" lastPageNumber="39" pageId="36" pageNumber="37" type="description">
<paragraph id="4344E72D9D7ECA69FF2CF97EFB89F88F" blockId="36.[151,1437,1609,2037]" box="[199,1130,1789,1816]" pageId="36" pageNumber="37">
<emphasis id="718F3B3F9D7ECA69FF2CF97EFEB2F88F" bold="true" box="[199,337,1789,1816]" pageId="36" pageNumber="37">Description</emphasis>
. Male (
<figureCitation id="DBC0FBA89D7ECA69FE43F97EFE1BF88F" box="[424,504,1789,1816]" captionStart="FIGURES 14–19" captionStartId="10.[151,264,1406,1431]" captionTargetBox="[155,1435,175,1386]" captionTargetId="figure@10.[151,1436,174,1394]" captionTargetPageId="10" captionText="FIGURES 14–19. Species of the “aquilina group” (14–18) and “bipunctata group” (19) in dorsal and ventral views. 14. Artines aquilina, A–B: male, Joinville, Santa Catarina, Brazil (DZ 41.321), C–D: female from same locality of male (DZ 41.220). 15. Artines angelica sp. nov., A–B: holotype male, C–D: allotype female. 16. Artines focus, A–B: male, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.233), C–D: female, Pacaraima, Roraima, Brazil (DZ 41.333). 17. Artines cofus sp. nov., A–B: holotype male, C–D: allotype female. 18. Artines donia sp. nov., A–B: holotype male. 19. Artines bipunctata, A–B: male, Jardim Zoológico, Brasília, Distrito Federal, Brazil (DZ 30.961), C–D: female, Córrego Bananal, Brasília, Distrito Federal, Brazil (DZ 41.224). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241045/files/figure.png" pageId="36" pageNumber="37">Fig. 18</figureCitation>
A–B).
<emphasis id="718F3B3F9D7ECA69FDAEF97EFD61F880" box="[581,642,1789,1815]" italics="true" pageId="36" pageNumber="37">Head</emphasis>
: antenna with nudum bearing 12 segments.
</paragraph>
<paragraph id="4344E72D9D7ECA69FF2CF8A1FE05F8AB" blockId="36.[151,1437,1609,2037]" box="[199,486,1825,1852]" pageId="36" pageNumber="37">
<emphasis id="718F3B3F9D7ECA69FF2CF8A1FE62F8AC" box="[199,385,1825,1851]" italics="true" pageId="36" pageNumber="37">Forewing length</emphasis>
:
<quantity id="84034AC89D7ECA69FE7BF8A1FE01F8AB" box="[400,482,1826,1852]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="36" pageNumber="37" unit="mm" value="14.0">14 mm</quantity>
.
</paragraph>
<paragraph id="4344E72D9D7ECA69FF2CF8C6FBEFF830" blockId="36.[151,1437,1609,2037]" pageId="36" pageNumber="37">
<emphasis id="718F3B3F9D7ECA69FF2CF8C6FE61F8F7" box="[199,386,1861,1888]" italics="true" pageId="36" pageNumber="37">Dorsal forewing</emphasis>
(
<figureCitation id="DBC0FBA89D7ECA69FE78F8C6FE1DF8F7" box="[403,510,1861,1888]" captionStart="FIGURES 14–19" captionStartId="10.[151,264,1406,1431]" captionTargetBox="[155,1435,175,1386]" captionTargetId="figure@10.[151,1436,174,1394]" captionTargetPageId="10" captionText="FIGURES 14–19. Species of the “aquilina group” (14–18) and “bipunctata group” (19) in dorsal and ventral views. 14. Artines aquilina, A–B: male, Joinville, Santa Catarina, Brazil (DZ 41.321), C–D: female from same locality of male (DZ 41.220). 15. Artines angelica sp. nov., A–B: holotype male, C–D: allotype female. 16. Artines focus, A–B: male, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.233), C–D: female, Pacaraima, Roraima, Brazil (DZ 41.333). 17. Artines cofus sp. nov., A–B: holotype male, C–D: allotype female. 18. Artines donia sp. nov., A–B: holotype male. 19. Artines bipunctata, A–B: male, Jardim Zoológico, Brasília, Distrito Federal, Brazil (DZ 30.961), C–D: female, Córrego Bananal, Brasília, Distrito Federal, Brazil (DZ 41.224). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241045/files/figure.png" pageId="36" pageNumber="37">Fig. 18A</figureCitation>
): two apical spots in R
<subScript id="DF7FE5689D7ECA69FCEDF8D4FCEDF8F2" attach="left" box="[774,782,1879,1893]" fontSize="6" pageId="36" pageNumber="37">4</subScript>
–R
<subScript id="DF7FE5689D7ECA69FCC4F8D4FCD4F8F2" attach="left" box="[815,823,1879,1893]" fontSize="6" pageId="36" pageNumber="37">5</subScript>
and R
<subScript id="DF7FE5689D7ECA69FC6FF8D4FC6FF8F2" attach="left" box="[900,908,1879,1893]" fontSize="6" pageId="36" pageNumber="37">5</subScript>
–M
<subScript id="DF7FE5689D7ECA69FC58F8D4FC58F8F2" attach="right" box="[947,955,1879,1893]" fontSize="6" pageId="36" pageNumber="37">1</subScript>
; discal spots as in
<taxonomicName id="84FB9CAE9D7ECA69FB7CF8C5FAF6F8C8" box="[1175,1301,1861,1888]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="aquilina">
<emphasis id="718F3B3F9D7ECA69FB7CF8C5FAF6F8C8" box="[1175,1301,1861,1888]" italics="true" pageId="36" pageNumber="37">A. aquilina</emphasis>
</taxonomicName>
; white spot in CuA
<subScript id="DF7FE5689D7ECA69FF01F8F8FF11F81E" attach="left" box="[234,242,1915,1929]" fontSize="6" pageId="36" pageNumber="37">2</subScript>
–2A very conspicuous, triangular; brand shorter than in other species of the “
<taxonomicName id="84FB9CAE9D7ECA69FB8BF8EAFB5EF814" box="[1120,1213,1897,1923]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="aquilina">
<emphasis id="718F3B3F9D7ECA69FB8BF8EAFB5EF814" box="[1120,1213,1897,1923]" italics="true" pageId="36" pageNumber="37">aquilina</emphasis>
</taxonomicName>
group”, with lower projection long and upper projection uniformly narrow and with apex rounded.
</paragraph>
<paragraph id="4344E72D9D7ECA69FF2CF832FD3BF867" blockId="36.[151,1437,1609,2037]" pageId="36" pageNumber="37">
<emphasis id="718F3B3F9D7ECA69FF2CF832FE65F85B" box="[199,390,1969,1996]" italics="true" pageId="36" pageNumber="37">Ventral forewing</emphasis>
(
<figureCitation id="DBC0FBA89D7ECA69FE7FF832FE18F85B" box="[404,507,1969,1996]" captionStart="FIGURES 14–19" captionStartId="10.[151,264,1406,1431]" captionTargetBox="[155,1435,175,1386]" captionTargetId="figure@10.[151,1436,174,1394]" captionTargetPageId="10" captionText="FIGURES 14–19. Species of the “aquilina group” (14–18) and “bipunctata group” (19) in dorsal and ventral views. 14. Artines aquilina, A–B: male, Joinville, Santa Catarina, Brazil (DZ 41.321), C–D: female from same locality of male (DZ 41.220). 15. Artines angelica sp. nov., A–B: holotype male, C–D: allotype female. 16. Artines focus, A–B: male, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.233), C–D: female, Pacaraima, Roraima, Brazil (DZ 41.333). 17. Artines cofus sp. nov., A–B: holotype male, C–D: allotype female. 18. Artines donia sp. nov., A–B: holotype male. 19. Artines bipunctata, A–B: male, Jardim Zoológico, Brasília, Distrito Federal, Brazil (DZ 30.961), C–D: female, Córrego Bananal, Brasília, Distrito Federal, Brazil (DZ 41.224). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241045/files/figure.png" pageId="36" pageNumber="37">Fig. 18B</figureCitation>
): apical and discal spots as in
<taxonomicName id="84FB9CAE9D7ECA69FCA3F831FC27F85C" box="[840,964,1969,1996]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="aquilina">
<emphasis id="718F3B3F9D7ECA69FCA3F831FC27F85C" box="[840,964,1969,1996]" italics="true" pageId="36" pageNumber="37">A. aquilina</emphasis>
</taxonomicName>
; submarginal band as in
<taxonomicName id="84FB9CAE9D7ECA69FB31F831FAD0F85B" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[1242,1331,1970,1996]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="cofus" status="sp. nov.">
<emphasis id="718F3B3F9D7ECA69FB31F831FAD0F85B" box="[1242,1331,1970,1996]" italics="true" pageId="36" pageNumber="37">A. cofus</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D7ECA69FAD1F831FA77F85B" bold="true" box="[1338,1428,1970,1996]" pageId="36" pageNumber="37">
<taxonomicNameLabel id="6ABC86449D7ECA69FAD1F831FA77F85B" box="[1338,1428,1970,1996]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; white spot broad and strongly marked in CuA
<subScript id="DF7FE5689D7ECA69FD73F864FD43F862" attach="left" box="[664,672,2023,2037]" fontSize="6" pageId="36" pageNumber="37">2</subScript>
–2A.
</paragraph>
<caption id="1784B7A59D7FCA68FF7CF945FDFBF805" httpUri="https://zenodo.org/record/3241063/files/figure.png" pageId="37" pageNumber="38" startId="37.[151,264,1734,1759]" targetBox="[160,1433,189,1717]" targetPageId="37">
<paragraph id="4344E72D9D7FCA68FF7CF945FDFBF805" blockId="37.[151,1437,1734,1939]" pageId="37" pageNumber="38">
<emphasis id="718F3B3F9D7FCA68FF7CF945FEACF948" bold="true" box="[151,335,1734,1759]" pageId="37" pageNumber="38">FIGURES 66–72</emphasis>
. Female genitalia of the “
<taxonomicName id="84FB9CAE9D7FCA68FDB4F94BFD4BF948" box="[607,680,1736,1759]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D7FCA68FDB4F94BFD4BF948" box="[607,680,1736,1759]" italics="true" pageId="37" pageNumber="38">aepitus</emphasis>
</taxonomicName>
group” species (ventral view of sterigma and
<taxonomicName id="84FB9CAE9D7FCA68FB63F945FB5CF948" box="[1160,1215,1734,1759]" pageId="37" pageNumber="38" rank="species" species="bursa">bursa</taxonomicName>
copulatrix).
<emphasis id="718F3B3F9D7FCA68FAAEF945FA86F948" bold="true" box="[1349,1381,1734,1759]" pageId="37" pageNumber="38">66.</emphasis>
<taxonomicName id="84FB9CAE9D7FCA68FA87F944FE84F894" authority=", Jusepin" authorityName="Jusepin" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="aepitus">
<emphasis id="718F3B3F9D7FCA68FA87F944FEE9F894" italics="true" pageId="37" pageNumber="38">Artines aepitus</emphasis>
, Jusepín
</taxonomicName>
, Monagas, Venezuela (OM 30. 125).
<emphasis id="718F3B3F9D7FCA68FD02F969FCEAF894" bold="true" box="[745,777,1770,1795]" pageId="37" pageNumber="38">67.</emphasis>
<taxonomicName id="84FB9CAE9D7FCA68FCFAF968FC57F894" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[785,948,1771,1795]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="panama" status="sp. nov.">
<emphasis id="718F3B3F9D7FCA68FCFAF968FC57F894" box="[785,948,1771,1795]" italics="true" pageId="37" pageNumber="38">Artines panama</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D7FCA68FC50F968FBEEF894" bold="true" box="[955,1037,1771,1795]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="6ABC86449D7FCA68FC50F968FBEEF894" box="[955,1037,1771,1795]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, allotype.
<emphasis id="718F3B3F9D7FCA68FB90F969FB78F894" bold="true" box="[1147,1179,1770,1795]" pageId="37" pageNumber="38">68.</emphasis>
<taxonomicName id="84FB9CAE9D7FCA68FB49F968FAA1F894" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[1186,1346,1771,1795]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="solange" status="sp. nov.">
<emphasis id="718F3B3F9D7FCA68FB49F968FAA1F894" box="[1186,1346,1771,1795]" italics="true" pageId="37" pageNumber="38">Artines solange</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D7FCA68FAA1F968FA7FF894" bold="true" box="[1354,1436,1771,1795]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="6ABC86449D7FCA68FAA1F968FA7FF894" box="[1354,1436,1771,1795]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(tentative female), Sumaúma, Maranhão, Brazil (DZ 41.800).
<emphasis id="718F3B3F9D7FCA68FCE4F88DFCCCF8B0" bold="true" box="[783,815,1806,1831]" pageId="37" pageNumber="38">69.</emphasis>
<taxonomicName id="84FB9CAE9D7FCA68FCDDF88CFC35F8B0" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[822,982,1807,1831]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="litoralis" status="sp. nov.">
<emphasis id="718F3B3F9D7FCA68FCDDF88CFC35F8B0" box="[822,982,1807,1831]" italics="true" pageId="37" pageNumber="38">Artines litoralis</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D7FCA68FC36F88CFBCDF8B0" bold="true" box="[989,1070,1807,1831]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="6ABC86449D7FCA68FC36F88CFBCDF8B0" box="[989,1070,1807,1831]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype, Linhares, Espírito Santo, Brazil (DZ 41.820).
<emphasis id="718F3B3F9D7FCA68FE8FF8B1FE67F8DC" bold="true" box="[356,388,1842,1867]" pageId="37" pageNumber="38">70.</emphasis>
<taxonomicName id="84FB9CAE9D7FCA68FE62F8B0FD8FF8DC" authority=", Turrialba" authorityName="Turrialba" box="[393,620,1842,1867]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="rica">
<emphasis id="718F3B3F9D7FCA68FE62F8B0FDE2F8DC" box="[393,513,1843,1867]" italics="true" pageId="37" pageNumber="38">Artines rica</emphasis>
, Turrialba
</taxonomicName>
, Cartago, Costa Rica (DZ 41.840).
<emphasis id="718F3B3F9D7FCA68FC25F8B1FC0DF8DC" bold="true" box="[974,1006,1842,1867]" pageId="37" pageNumber="38">71.</emphasis>
<taxonomicName id="84FB9CAE9D7FCA68FC18F8B0FB91F8DC" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[1011,1138,1843,1867]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="liege" status="sp. nov.">
<emphasis id="718F3B3F9D7FCA68FC18F8B0FB91F8DC" box="[1011,1138,1843,1867]" italics="true" pageId="37" pageNumber="38">Artines liege</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D7FCA68FB93F8B0FB2BF8DC" bold="true" box="[1144,1224,1843,1867]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="6ABC86449D7FCA68FB93F8B0FB2BF8DC" box="[1144,1224,1843,1867]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype, Parque do Gama, Brasília, Distrito Federal, Brazil (DZ 41.830).
<emphasis id="718F3B3F9D7FCA68FD50F8D5FD38F8F8" bold="true" box="[699,731,1878,1903]" pageId="37" pageNumber="38">72.</emphasis>
<taxonomicName id="84FB9CAE9D7FCA68FD0AF8D4FC95F8F8" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[737,886,1879,1903]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="bamba" status="sp. nov.">
<emphasis id="718F3B3F9D7FCA68FD0AF8D4FC95F8F8" box="[737,886,1879,1903]" italics="true" pageId="37" pageNumber="38">Artines bamba</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D7FCA68FC96F8D4FC33F8F8" bold="true" box="[893,976,1879,1903]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="6ABC86449D7FCA68FC96F8D4FC33F8F8" box="[893,976,1879,1903]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(tentative female), Moyobamba, San Martin, Peru (OM 81.229). Scale bar = 1 mm.
</paragraph>
</caption>
<caption id="1784B7A59D7CCA6BFF7CFC57FC92FBEB" httpUri="https://zenodo.org/record/3241065/files/figure.png" pageId="38" pageNumber="39" startId="38.[151,264,980,1005]" targetBox="[155,1424,182,962]" targetPageId="38">
<paragraph id="4344E72D9D7CCA6BFF7CFC57FC92FBEB" blockId="38.[151,1437,980,1149]" pageId="38" pageNumber="39">
<emphasis id="718F3B3F9D7CCA6BFF7CFC57FEB0FC7A" bold="true" box="[151,339,980,1005]" pageId="38" pageNumber="39">FIGURES 73–77.</emphasis>
Female genitalia of the “
<taxonomicName id="84FB9CAE9D7CCA6BFDBBFC56FD47FC7A" box="[592,676,981,1005]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="aquilina">
<emphasis id="718F3B3F9D7CCA6BFDBBFC56FD47FC7A" box="[592,676,981,1005]" italics="true" pageId="38" pageNumber="39">aquilina</emphasis>
</taxonomicName>
group” and “
<taxonomicName id="84FB9CAE9D7CCA6BFCC7FC56FC7BFC7A" box="[812,920,981,1005]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="bipunctata">
<emphasis id="718F3B3F9D7CCA6BFCC7FC56FC7BFC7A" box="[812,920,981,1005]" italics="true" pageId="38" pageNumber="39">bipunctata</emphasis>
</taxonomicName>
group” species (ventral view of sterigma and bursa copulatrix).
<emphasis id="718F3B3F9D7CCA6BFEF9FC7BFED1FB86" bold="true" box="[274,306,1016,1041]" pageId="38" pageNumber="39">73.</emphasis>
<taxonomicName id="84FB9CAE9D7CCA6BFEDCFC7AFDABFB86" authority=", Cananeia" authorityName="Cananeia" box="[311,584,1016,1041]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="aquilina">
<emphasis id="718F3B3F9D7CCA6BFEDCFC7AFE39FB86" box="[311,474,1017,1041]" italics="true" pageId="38" pageNumber="39">Artines aquilina</emphasis>
, Cananéia
</taxonomicName>
, São Paulo, Brazil (OM 49.831).
<emphasis id="718F3B3F9D7CCA6BFC7DFC7BFC55FB86" bold="true" box="[918,950,1016,1041]" pageId="38" pageNumber="39">74.</emphasis>
<taxonomicName id="84FB9CAE9D7CCA6BFC50FC7AFB82FB86" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[955,1121,1017,1041]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="angelica" status="sp. nov.">
<emphasis id="718F3B3F9D7CCA6BFC50FC7AFB82FB86" box="[955,1121,1017,1041]" italics="true" pageId="38" pageNumber="39">Artines angelica</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D7CCA6BFB8CFC7AFB5EFB86" bold="true" box="[1127,1213,1017,1041]" pageId="38" pageNumber="39">
<taxonomicNameLabel id="6ABC86449D7CCA6BFB8CFC7AFB5AFB86" box="[1127,1209,1017,1041]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
paratype, Tambopata, Puerto Maldonado, Madre de Díos, Peru (DZ 41.222).
<emphasis id="718F3B3F9D7CCA6BFD2CFB9EFD04FBA2" bold="true" box="[711,743,1053,1077]" pageId="38" pageNumber="39">75.</emphasis>
<taxonomicName id="84FB9CAE9D7CCA6BFD05FB9EFC0FFBA2" authority=", Pacaraima" authorityName="Pacaraima" box="[750,1004,1052,1077]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="focus">
<emphasis id="718F3B3F9D7CCA6BFD05FB9EFC90FBA2" box="[750,883,1053,1077]" italics="true" pageId="38" pageNumber="39">Artines focus</emphasis>
, Pacaraima
</taxonomicName>
, Roraima, Brazil (DZ 41.333).
<emphasis id="718F3B3F9D7CCA6BFAC6FB9FFAAFFBA2" bold="true" box="[1325,1356,1052,1077]" pageId="38" pageNumber="39">76.</emphasis>
<taxonomicName id="84FB9CAE9D7CCA6BFAB8FB9EFF2FFBCE" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="cofus" status="sp. nov.">
<emphasis id="718F3B3F9D7CCA6BFAB8FB9EFF2FFBCE" italics="true" pageId="38" pageNumber="39">Artines cofus</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D7CCA6BFF3EFBC2FEC4FBCE" bold="true" box="[213,295,1089,1113]" pageId="38" pageNumber="39">
<taxonomicNameLabel id="6ABC86449D7CCA6BFF3EFBC2FEC4FBCE" box="[213,295,1089,1113]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
paratype, Tambopata, Puerto Maldonado, Madre de Díos, Peru (DZ 41. 293).
<emphasis id="718F3B3F9D7CCA6BFBBCFBC2FB94FBCE" bold="true" box="[1111,1143,1089,1113]" pageId="38" pageNumber="39">77.</emphasis>
<taxonomicName id="84FB9CAE9D7CCA6BFB95FBC2FA7EFBCF" authority=", Reserva" authorityName="Reserva" box="[1150,1437,1089,1113]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="bipunctata">
<emphasis id="718F3B3F9D7CCA6BFB95FBC2FAA0FBCE" box="[1150,1347,1089,1113]" italics="true" pageId="38" pageNumber="39">Artines bipunctata,</emphasis>
Reserva
</taxonomicName>
IBGE, Brasília, Distrito Federal, Brazil (DZ 41.165). Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="4344E72D9D7CCA6BFF2CFB28FB37FB7E" blockId="38.[151,1437,1195,2013]" pageId="38" pageNumber="39">
<emphasis id="718F3B3F9D7CCA6BFF2CFB28FE72FB52" box="[199,401,1195,1221]" italics="true" pageId="38" pageNumber="39">Ventral hind wing</emphasis>
(
<figureCitation id="DBC0FBA89D7CCA6BFE4AFB28FDEBFB52" box="[417,520,1195,1221]" captionStart="FIGURES 14–19" captionStartId="10.[151,264,1406,1431]" captionTargetBox="[155,1435,175,1386]" captionTargetId="figure@10.[151,1436,174,1394]" captionTargetPageId="10" captionText="FIGURES 14–19. Species of the “aquilina group” (14–18) and “bipunctata group” (19) in dorsal and ventral views. 14. Artines aquilina, A–B: male, Joinville, Santa Catarina, Brazil (DZ 41.321), C–D: female from same locality of male (DZ 41.220). 15. Artines angelica sp. nov., A–B: holotype male, C–D: allotype female. 16. Artines focus, A–B: male, Ilha de Maracá, Alto Alegre, Roraima, Brazil (DZ 41.233), C–D: female, Pacaraima, Roraima, Brazil (DZ 41.333). 17. Artines cofus sp. nov., A–B: holotype male, C–D: allotype female. 18. Artines donia sp. nov., A–B: holotype male. 19. Artines bipunctata, A–B: male, Jardim Zoológico, Brasília, Distrito Federal, Brazil (DZ 30.961), C–D: female, Córrego Bananal, Brasília, Distrito Federal, Brazil (DZ 41.224). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241045/files/figure.png" pageId="38" pageNumber="39">Fig. 18B</figureCitation>
): apical portion of discal cell densely covered by orange scales; central black spot rectangular, wider than long; postdiscal band yellow, limited distally by wavy dark brown stripe.
</paragraph>
<paragraph id="4344E72D9D7CCA6BFF2CFB70FAC1F92A" blockId="38.[151,1437,1195,2013]" pageId="38" pageNumber="39">
<emphasis id="718F3B3F9D7CCA6BFF2CFB70FED2FA9A" box="[199,305,1267,1293]" italics="true" pageId="38" pageNumber="39">Genitalia</emphasis>
(
<figureCitation id="DBC0FBA89D7CCA6BFED5FB70FE71FA9A" box="[318,402,1267,1293]" captionStart="FIGURES 40–41" captionStartId="31.[151,264,1624,1649]" captionTargetBox="[154,1435,484,1599]" captionTargetId="figure@31.[151,1436,481,1612]" captionText="FIGURES 40–41. Male genitalia of the “aquilina group” (40) and “bipunctata group” (41) species. 40. Artines donia sp. nov., paratype, Santa Rita do Araguaia, Goiás, Brazil (DZ 66.000). 41. Artines bipunctata, Brasília, Distrito Federal, Brazil (DZ 41.194). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3406272/files/figure.png" pageId="38" pageNumber="39">Figs 40</figureCitation>
A–L, 53): tegumen more or less oval, longer than wide (
<figureCitation id="DBC0FBA89D7CCA6BFC1CFB70FBBFFA9A" box="[1015,1116,1267,1293]" captionStart="FIGURES 40–41" captionStartId="31.[151,264,1624,1649]" captionTargetBox="[154,1435,484,1599]" captionTargetId="figure@31.[151,1436,481,1612]" captionText="FIGURES 40–41. Male genitalia of the “aquilina group” (40) and “bipunctata group” (41) species. 40. Artines donia sp. nov., paratype, Santa Rita do Araguaia, Goiás, Brazil (DZ 66.000). 41. Artines bipunctata, Brasília, Distrito Federal, Brazil (DZ 41.194). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3406272/files/figure.png" pageId="38" pageNumber="39">Fig. 40C</figureCitation>
), in lateral view, rectangular, dorsal margin straight on its entire stretch (
<figureCitation id="DBC0FBA89D7CCA6BFD63FA94FD10FAA6" box="[648,755,1303,1329]" captionStart="FIGURES 40–41" captionStartId="31.[151,264,1624,1649]" captionTargetBox="[154,1435,484,1599]" captionTargetId="figure@31.[151,1436,481,1612]" captionText="FIGURES 40–41. Male genitalia of the “aquilina group” (40) and “bipunctata group” (41) species. 40. Artines donia sp. nov., paratype, Santa Rita do Araguaia, Goiás, Brazil (DZ 66.000). 41. Artines bipunctata, Brasília, Distrito Federal, Brazil (DZ 41.194). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3406272/files/figure.png" pageId="38" pageNumber="39">Fig. 40A</figureCitation>
). Saccus narrowed medially, anterior portion narrower than tegumen, proximal margin rounded (
<figureCitation id="DBC0FBA89D7CCA6BFDAAFAB8FD48FAC2" box="[577,683,1339,1365]" captionStart="FIGURES 40–41" captionStartId="31.[151,264,1624,1649]" captionTargetBox="[154,1435,484,1599]" captionTargetId="figure@31.[151,1436,481,1612]" captionText="FIGURES 40–41. Male genitalia of the “aquilina group” (40) and “bipunctata group” (41) species. 40. Artines donia sp. nov., paratype, Santa Rita do Araguaia, Goiás, Brazil (DZ 66.000). 41. Artines bipunctata, Brasília, Distrito Federal, Brazil (DZ 41.194). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3406272/files/figure.png" pageId="38" pageNumber="39">Fig. 40B</figureCitation>
). Uncus shorter than tegumen, proximal 1/2 with lateral margins slightly concave and distally divergent, distal 1/2 narrower with strong lateral concavity, distal end weakly bifid (
<figureCitation id="DBC0FBA89D7CCA6BFF4BFA00FEE7FA0A" box="[160,260,1411,1437]" captionStart="FIGURES 40–41" captionStartId="31.[151,264,1624,1649]" captionTargetBox="[154,1435,484,1599]" captionTargetId="figure@31.[151,1436,481,1612]" captionText="FIGURES 40–41. Male genitalia of the “aquilina group” (40) and “bipunctata group” (41) species. 40. Artines donia sp. nov., paratype, Santa Rita do Araguaia, Goiás, Brazil (DZ 66.000). 41. Artines bipunctata, Brasília, Distrito Federal, Brazil (DZ 41.194). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3406272/files/figure.png" pageId="38" pageNumber="39">Fig. 40C</figureCitation>
), little curved ventrally (
<figureCitation id="DBC0FBA89D7CCA6BFDF8FA00FD99FA0A" box="[531,634,1411,1437]" captionStart="FIGURES 40–41" captionStartId="31.[151,264,1624,1649]" captionTargetBox="[154,1435,484,1599]" captionTargetId="figure@31.[151,1436,481,1612]" captionText="FIGURES 40–41. Male genitalia of the “aquilina group” (40) and “bipunctata group” (41) species. 40. Artines donia sp. nov., paratype, Santa Rita do Araguaia, Goiás, Brazil (DZ 66.000). 41. Artines bipunctata, Brasília, Distrito Federal, Brazil (DZ 41.194). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3406272/files/figure.png" pageId="38" pageNumber="39">Fig. 40A</figureCitation>
). Gnathos in ventral view reaching 1/2 of uncus, arms parallel with distal ends straight (
<figureCitation id="DBC0FBA89D7CCA6BFED8FA24FE78FA56" box="[307,411,1447,1473]" captionStart="FIGURES 40–41" captionStartId="31.[151,264,1624,1649]" captionTargetBox="[154,1435,484,1599]" captionTargetId="figure@31.[151,1436,481,1612]" captionText="FIGURES 40–41. Male genitalia of the “aquilina group” (40) and “bipunctata group” (41) species. 40. Artines donia sp. nov., paratype, Santa Rita do Araguaia, Goiás, Brazil (DZ 66.000). 41. Artines bipunctata, Brasília, Distrito Federal, Brazil (DZ 41.194). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3406272/files/figure.png" pageId="38" pageNumber="39">Fig. 40D</figureCitation>
), curved dorsally toward distal end of uncus (
<figureCitation id="DBC0FBA89D7CCA6BFC7EFA24FC1EFA56" box="[917,1021,1447,1473]" captionStart="FIGURES 40–41" captionStartId="31.[151,264,1624,1649]" captionTargetBox="[154,1435,484,1599]" captionTargetId="figure@31.[151,1436,481,1612]" captionText="FIGURES 40–41. Male genitalia of the “aquilina group” (40) and “bipunctata group” (41) species. 40. Artines donia sp. nov., paratype, Santa Rita do Araguaia, Goiás, Brazil (DZ 66.000). 41. Artines bipunctata, Brasília, Distrito Federal, Brazil (DZ 41.194). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3406272/files/figure.png" pageId="38" pageNumber="39">Fig. 40A</figureCitation>
). Valva three times longer than wide; costa narrow and triangular, laterally projected inwards, separated from ampulla by distance shorter than its length; harpe longer than 1/2 valva, widened from base; ampulla 1/2 width of median portion of harpe, dorsally protruding, slightly exceeding dorsal margin of valva; distal projection of valva wide, with thorn-like projection on dorsal-distal portion (
<figureCitation id="DBC0FBA89D7CCA6BFF12F9B4FEAFF9C6" box="[249,332,1591,1617]" captionStart="FIGURES 40–41" captionStartId="31.[151,264,1624,1649]" captionTargetBox="[154,1435,484,1599]" captionTargetId="figure@31.[151,1436,481,1612]" captionText="FIGURES 40–41. Male genitalia of the “aquilina group” (40) and “bipunctata group” (41) species. 40. Artines donia sp. nov., paratype, Santa Rita do Araguaia, Goiás, Brazil (DZ 66.000). 41. Artines bipunctata, Brasília, Distrito Federal, Brazil (DZ 41.194). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3406272/files/figure.png" pageId="38" pageNumber="39">Fig. 40</figureCitation>
E–F). Aedeagus uniformly wide, weakly curved dorsally; opening of ejaculatory bulb dorsal near proximal margin of aedeagus; ventral projection diverging from aedeagus at an angled of 45 degrees, distal portion asymmetrically bifid (
<figureCitation id="DBC0FBA89D7CCA6BFE65F9FCFE03F90E" box="[398,480,1663,1689]" captionStart="FIGURES 40–41" captionStartId="31.[151,264,1624,1649]" captionTargetBox="[154,1435,484,1599]" captionTargetId="figure@31.[151,1436,481,1612]" captionText="FIGURES 40–41. Male genitalia of the “aquilina group” (40) and “bipunctata group” (41) species. 40. Artines donia sp. nov., paratype, Santa Rita do Araguaia, Goiás, Brazil (DZ 66.000). 41. Artines bipunctata, Brasília, Distrito Federal, Brazil (DZ 41.194). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3406272/files/figure.png" pageId="38" pageNumber="39">Fig. 40</figureCitation>
G–J). Cornutus uniformly sclerotized, bearing setae-like spines at base of vesica (
<figureCitation id="DBC0FBA89D7CCA6BFA81F9FCFF2CF92A" captionStart="FIGURES 40–41" captionStartId="31.[151,264,1624,1649]" captionTargetBox="[154,1435,484,1599]" captionTargetId="figure@31.[151,1436,481,1612]" captionText="FIGURES 40–41. Male genitalia of the “aquilina group” (40) and “bipunctata group” (41) species. 40. Artines donia sp. nov., paratype, Santa Rita do Araguaia, Goiás, Brazil (DZ 66.000). 41. Artines bipunctata, Brasília, Distrito Federal, Brazil (DZ 41.194). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3406272/files/figure.png" pageId="38" pageNumber="39">Figs 40M</figureCitation>
, 53). Fultura inferior robust, in lateral view wider than in the other species of the group (
<figureCitation id="DBC0FBA89D7CCA6BFB5BF920FAF5F92A" box="[1200,1302,1699,1725]" captionStart="FIGURES 40–41" captionStartId="31.[151,264,1624,1649]" captionTargetBox="[154,1435,484,1599]" captionTargetId="figure@31.[151,1436,481,1612]" captionText="FIGURES 40–41. Male genitalia of the “aquilina group” (40) and “bipunctata group” (41) species. 40. Artines donia sp. nov., paratype, Santa Rita do Araguaia, Goiás, Brazil (DZ 66.000). 41. Artines bipunctata, Brasília, Distrito Federal, Brazil (DZ 41.194). For reference of the letters see legend of the figures 27 and 36. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3406272/files/figure.png" pageId="38" pageNumber="39">Fig. 40L</figureCitation>
).
</paragraph>
<paragraph id="4344E72D9D7CCA6BFF2CF944FE7DF976" blockId="38.[151,1437,1195,2013]" box="[199,414,1735,1761]" pageId="38" pageNumber="39">Female. Unknown.</paragraph>
<paragraph id="4344E72D9D7CCA6BFF2CF969FE06F8BD" blockId="38.[151,1437,1195,2013]" pageId="38" pageNumber="39">
<emphasis id="718F3B3F9D7CCA6BFF2CF969FED9F892" bold="true" box="[199,314,1770,1797]" pageId="38" pageNumber="39">Variation</emphasis>
. The specimen from
<collectingRegion id="813F29CF9D7CCA6BFDC1F968FD8DF892" box="[554,622,1771,1797]" country="Brazil" name="Goias" pageId="38" pageNumber="39">Goiás</collectingRegion>
,
<collectingCountry id="3BECA7BD9D7CCA6BFD91F968FD22F892" box="[634,705,1771,1797]" name="Brazil" pageId="38" pageNumber="39">Brazil</collectingCountry>
, is paler than the specimen from
<collectingRegion id="813F29CF9D7CCA6BFBDDF968FB45F892" box="[1078,1190,1771,1797]" country="Brazil" name="Rondonia" pageId="38" pageNumber="39">Rondônia</collectingRegion>
, which may be due to the fact that the wing is worn.
</paragraph>
</subSubSection>
<subSubSection id="0BE1B4A69D7CCA6AFF2CF8B1FA86FE44" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39" type="discussion">
<paragraph id="4344E72D9D7CCA6BFF2CF8B1FEAAF84A" blockId="38.[151,1437,1195,2013]" pageId="38" pageNumber="39">
<emphasis id="718F3B3F9D7CCA6BFF2CF8B1FEABF8DA" bold="true" box="[199,328,1842,1869]" pageId="38" pageNumber="39">Comments</emphasis>
. Collor pattern as in
<emphasis id="718F3B3F9D7CCA6BFDD5F8B7FCC9F8DA" box="[574,810,1843,1870]" italics="true" pageId="38" pageNumber="39">
<taxonomicName id="84FB9CAE9D7CCA6BFDD5F8B7FD7EF8DA" box="[574,669,1843,1870]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="focus">A. focus</taxonomicName>
,
<taxonomicName id="84FB9CAE9D7CCA6BFD42F8B7FCC9F8DA" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[681,810,1843,1870]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="angelica" status="sp. nov.">A. angelica</taxonomicName>
</emphasis>
<emphasis id="718F3B3F9D7CCA6BFCD8F8B0FC73F8DA" bold="true" box="[819,912,1843,1869]" pageId="38" pageNumber="39">
<taxonomicNameLabel id="6ABC86449D7CCA6BFCD8F8B0FC73F8DA" box="[819,912,1843,1869]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="84FB9CAE9D7CCA6BFC21F8B7FBC5F8DA" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[970,1062,1843,1870]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="cofus" status="sp. nov.">
<emphasis id="718F3B3F9D7CCA6BFC21F8B7FBC5F8DA" box="[970,1062,1843,1870]" italics="true" pageId="38" pageNumber="39">A. cofus</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D7CCA6BFBC4F8B0FB6FF8DA" bold="true" box="[1071,1164,1843,1869]" pageId="38" pageNumber="39">
<taxonomicNameLabel id="6ABC86449D7CCA6BFBC4F8B0FB6FF8DA" box="[1071,1164,1843,1869]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, differing in the central black spot of the ventral hind wing more reduced and wider than long. The male genitalia are similar to
<taxonomicName id="84FB9CAE9D7CCA6BFAFFF8DBFA8DF8E6" box="[1300,1390,1879,1906]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="focus">
<emphasis id="718F3B3F9D7CCA6BFAFFF8DBFA8DF8E6" box="[1300,1390,1879,1906]" italics="true" pageId="38" pageNumber="39">A. focus</emphasis>
</taxonomicName>
and
<taxonomicName id="84FB9CAE9D7CCA6BFF7CF8FFFF12F802" authority="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande, 2019" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[151,241,1915,1942]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="cofus" status="sp. nov.">
<emphasis id="718F3B3F9D7CCA6BFF7CF8FFFF12F802" box="[151,241,1915,1942]" italics="true" pageId="38" pageNumber="39">A. cofus</emphasis>
</taxonomicName>
<emphasis id="718F3B3F9D7CCA6BFF12F8F8FEB7F802" bold="true" box="[249,340,1915,1941]" pageId="38" pageNumber="39">
<taxonomicNameLabel id="6ABC86449D7CCA6BFF12F8F8FEB7F802" box="[249,340,1915,1941]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, from which it differs in the following characters: uncus slightly bifid, without ventral projection, aedeagus uniformly wide, fultura inferior robust and valva distally not pointed, with projection similar to a distally projected thorn.
</paragraph>
<paragraph id="4344E72D9D7DCA6AFF2CFF1BFDEBFE38" blockId="39.[151,1436,152,467]" pageId="39" pageNumber="40">
<emphasis id="718F3B3F9D7DCA6AFF2CFF1BFE09FF24" bold="true" box="[199,490,152,179]" pageId="39" pageNumber="40">Geographic distribution</emphasis>
(
<figureCitation id="DBC0FBA89D7DCA6AFE10FF1BFDB2FF24" box="[507,593,152,179]" captionStart="FIGURE 80" captionStartId="43.[151,250,930,955]" captionTargetBox="[151,1436,187,905]" captionTargetId="figure@43.[151,1436,187,905]" captionTargetPageId="43" captionText="FIGURE 80. Geographic distribution of the “aquilina group” and “bipunctata group” species. The symbols with white inferior half represent records from literature." httpUri="https://zenodo.org/record/3241071/files/figure.png" pageId="39" pageNumber="40">Fig. 80</figureCitation>
). It is only known from the southern Brazilian Amazon Rainforest. BRA- ZIL—
<emphasis id="718F3B3F9D7DCA6AFF34FF3FFEAFFF41" box="[223,332,188,214]" italics="true" pageId="39" pageNumber="40">
<collectingRegion id="813F29CF9D7DCA6AFF34FF3FFEAFFF41" box="[223,332,188,214]" country="Brazil" name="Rondonia" pageId="39" pageNumber="40">Rondônia</collectingRegion>
</emphasis>
: Vilhena.
<emphasis id="718F3B3F9D7DCA6AFE54FF3EFDE1FF40" box="[447,514,189,215]" italics="true" pageId="39" pageNumber="40">
<collectingRegion id="813F29CF9D7DCA6AFE54FF3EFDE1FF40" box="[447,514,189,215]" country="Brazil" name="Goias" pageId="39" pageNumber="40">Goiás</collectingRegion>
</emphasis>
: Santa Rita do Araguaia.
<emphasis id="718F3B3F9D7DCA6AFF2CFF62FE33FF6C" bold="true" box="[199,464,224,251]" pageId="39" pageNumber="40">Temporal distribution</emphasis>
. This species occurs in May and July.
<emphasis id="718F3B3F9D7DCA6AFF2CFE86FEA7FE88" bold="true" box="[199,324,261,287]" pageId="39" pageNumber="40">Host plant</emphasis>
. Unknown.
<emphasis id="718F3B3F9D7DCA6AFF2CFEAAFEABFED4" bold="true" box="[199,328,297,323]" pageId="39" pageNumber="40">Etymology</emphasis>
. The term “
<taxonomicName id="84FB9CAE9D7DCA6AFE3BFEABFDF3FED5" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[464,528,296,322]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="donia">
<emphasis id="718F3B3F9D7DCA6AFE3BFEABFDF3FED5" box="[464,528,296,322]" italics="true" pageId="39" pageNumber="40">donia</emphasis>
</taxonomicName>
” is the termination of the word “
<collectingRegion id="813F29CF9D7DCA6AFC6BFEABFC16FED5" box="[896,1013,296,322]" country="Brazil" name="Rondonia" pageId="39" pageNumber="40">Rondônia</collectingRegion>
”, the type locality of the species.
<materialsCitation id="F393ED709D7DCA6AFF2CFECEFDEBFE38" ID-GBIF-Occurrence="2336657221" collectingDate="1969-05-21" collectingDateMax="1972-07-10" collectingDateMin="1969-05-21" collectionCode="OM" collectorName="Mielke &amp; Brown" country="Brazil" determinerName="Medeiros &amp; Carneiro" location="RO" municipality="RO" pageId="39" pageNumber="40" specimenCount="1" specimenCount-male="1" stateProvince="Vilhena" typeStatus="holotype">
<emphasis id="718F3B3F9D7DCA6AFF2CFECEFE91FEF0" bold="true" box="[199,370,332,359]" pageId="39" pageNumber="40">Type material</emphasis>
. Holotype male deposited at the DZUP with the following labels: /
<typeStatus id="9C40598F9D7DCA6AFB96FECEFAC0FEF0" box="[1149,1315,333,359]" pageId="39" pageNumber="40" type="holotype">HOLOTYPUS</typeStatus>
/
<collectingRegion id="813F29CF9D7DCA6AFAD6FECFFA7AFEF1" box="[1341,1433,332,358]" pageId="39" pageNumber="40">Vilhena</collectingRegion>
,
<collectingMunicipality id="A3207D579D7DCA6AFF7CFEF3FF5CFE1D" box="[151,191,368,394]" pageId="39" pageNumber="40">RO</collectingMunicipality>
[ndônia,
<collectingCountry id="3BECA7BD9D7DCA6AFECAFEF3FE88FE1C" box="[289,363,368,395]" name="Brazil" pageId="39" pageNumber="40">Brazil</collectingCountry>
],
<date id="3745C1ED9D7DCA6AFE96FEF2FDEBFE1C" box="[381,520,369,395]" pageId="39" pageNumber="40" value="1972-07-10">
<collectingDate id="270138059D7DCA6AFE96FEF2FDEBFE1C" box="[381,520,369,395]" pageId="39" pageNumber="40" value="1972-07-10">10.VII.1972</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D7DCA6AFDF8FEF3FD81FE1D" box="[531,610,368,394]" pageId="39" pageNumber="40">Mielke</collectorName>
<emphasis id="718F3B3F9D7DCA6AFD82FEF3FD68FE1D" box="[617,651,368,394]" italics="true" pageId="39" pageNumber="40">leg</emphasis>
. / gen. prep. A. Medeiros 2017 / DZ 41.283 / Holotypus
<taxonomicName id="84FB9CAE9D7DCA6AFAEEFEF2FA7FFE1D" authority="Medeiros &amp; Carneiro" authorityName="Medeiros &amp; Dolibaina &amp; Carneiro &amp; Mielke &amp; Casagrande" authorityYear="2019" box="[1285,1436,368,395]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="37" phylum="Arthropoda" rank="species" species="donia" status="sp. nov.">
<emphasis id="718F3B3F9D7DCA6AFAEEFEF2FA7FFE1D" box="[1285,1436,368,395]" italics="true" pageId="39" pageNumber="40">Artines donia</emphasis>
</taxonomicName>
<determinerName id="B6980BE99D7DCA6AFF7CFE17FEE2FE39" box="[151,257,404,430]" pageId="39" pageNumber="40">Medeiros</determinerName>
&amp;
<determinerName id="B6980BE99D7DCA6AFECEFE17FE64FE39" box="[293,391,404,430]" pageId="39" pageNumber="40">Carneiro</determinerName>
det. 2018 /
</materialsCitation>
</paragraph>
<paragraph id="4344E72D9D7DCA6AFDE3FE16FA86FE44" blockId="39.[151,1436,152,467]" pageId="39" pageNumber="40">
.
<typeStatus id="9C40598F9D7DCA6AFF2CFE3AFECEFE44" box="[199,301,441,467]" pageId="39" pageNumber="40" type="paratype">Paratype</typeStatus>
:
<collectingCountry id="3BECA7BD9D7DCA6AFED3FE3AFE5AFE44" box="[312,441,441,467]" pageId="39" pageNumber="40">BRAZIL—</collectingCountry>
<emphasis id="718F3B3F9D7DCA6AFE52FE3AFE1FFE44" box="[441,508,441,467]" italics="true" pageId="39" pageNumber="40">
<collectingRegion id="813F29CF9D7DCA6AFE52FE3AFE1FFE44" box="[441,508,441,467]" country="Brazil" name="Goias" pageId="39" pageNumber="40">Goiás</collectingRegion>
</emphasis>
:
<emphasis id="718F3B3F9D7DCA6AFDE0FE3BFCC7FE44" bold="true" box="[523,804,440,467]" pageId="39" pageNumber="40">
<collectingMunicipality id="A3207D579D7DCA6AFDE0FE3BFCC7FE44" box="[523,804,440,467]" name="Santa Rita do Araguaia" pageId="39" pageNumber="40" stateProvince="Goias">Santa Rita do Araguaia</collectingMunicipality>
</emphasis>
,
<specimenCount id="55FD2CA49D7DCA6AFCD9FE3AFC63FE45" box="[818,896,440,467]" pageId="39" pageNumber="40" type="male">1 male</specimenCount>
,
<date id="3745C1ED9D7DCA6AFC60FE3AFBE3FE45" box="[907,1024,440,467]" pageId="39" pageNumber="40" value="1969-05-21">
<collectingDate id="270138059D7DCA6AFC60FE3AFBE3FE45" box="[907,1024,440,467]" pageId="39" pageNumber="40" value="1969-05-21">21.V.1969</collectingDate>
</date>
,
<collectorName id="EE0E82FB9D7DCA6AFBE7FE3AFBBBFE44" box="[1036,1112,441,467]" pageId="39" pageNumber="40">Brown</collectorName>
<emphasis id="718F3B3F9D7DCA6AFBB4FE3BFB6BFE45" box="[1119,1160,440,466]" italics="true" pageId="39" pageNumber="40">leg.</emphasis>
, OM 66.000 (OM).
</paragraph>
</subSubSection>
</treatment>
</document>

86
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<taxonomicName id="84FB4D04FFB4FFD91B94FE2EFDBBCDAC" authority="Lacordaire, 1866" authorityName="Lacordaire" authorityYear="1866" box="[151,540,388,414]" class="Insecta" family="Anthribidae" genus="Diastatotropis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="430" phylum="Arthropoda" rank="genus">
<emphasis id="718FEA95FFB4FFD91B94FE2EFDBBCDAC" bold="true" box="[151,540,388,414]" pageId="1" pageNumber="430">
<emphasis id="718FEA95FFB4FFD91B94FE2EFEE7CDAC" bold="true" box="[151,320,388,414]" italics="true" pageId="1" pageNumber="430">Diastatotropis</emphasis>
<bibRefCitation id="276A4B76FFB4FFD91A44FE2EFDBBCDAC" author="Lacordaire" box="[327,540,388,414]" pageId="1" pageNumber="430" refString="Lacordaire, J. T. (1866) Histoire naturelle des insectes. In: Genera des Coleopteres ou expose methodique et critique de tous les genres proposes jusqu´ici dans cet ordre d´insectes. Tome septieme contenant les families des curculionides (suite), scolytides, brenthides, anthribides et bruchides. Vol. 7. Librairie Roret, Paris, pp. 520." type="book chapter" year="1866">Lacordaire, 1866</bibRefCitation>
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<emphasis id="718FEA95FFB4FFD91B94FE60FE8ACDD1" bold="true" box="[151,301,458,483]" pageId="1" pageNumber="430">Recognition.</emphasis>
Body elongate. Head comparatively long, rostrum longer, extended apically, without deep depression in the middle, with one central longitudinal carina (or without this carina in case of
<taxonomicName id="84FB4D04FFB4FFD91F63FE45FF68CE19" authority="Waterhouse, 1882" authorityName="Waterhouse" authorityYear="1882" class="Insecta" family="Anthribidae" genus="Diastatotropis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="430" phylum="Arthropoda" rank="species" species="nitidipennis">
<emphasis id="718FEA95FFB4FFD91F63FE45FAAFCE34" box="[1120,1288,494,518]" italics="true" pageId="1" pageNumber="430">D. nitidipennis</emphasis>
Waterhouse, 1882
</taxonomicName>
and
<taxonomicName id="84FB4D04FFB4FFD91A0FFDB9FDDCCE19" authority="Waterhouse, 1882" authorityName="Waterhouse" authorityYear="1882" box="[268,635,530,555]" class="Insecta" family="Anthribidae" genus="Diastatotropis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="430" phylum="Arthropoda" rank="species" species="planifrons">
<emphasis id="718FEA95FFB4FFD91A0FFDB9FE04CE19" box="[268,419,531,555]" italics="true" pageId="1" pageNumber="430">D. planifrons</emphasis>
Waterhouse, 1882
</taxonomicName>
) and two lateral carinae. Eyes spherical or elliptical, not emarginate. Antennae of males usually reaching posterior margin of pronotum or longer, antennae of females shorter, not reaching posterior margin of pronotum. Funicle thin, antennal club 3-segmented or antennomeres continuously extended. Dorsal transverse carina of pronotum distinct. Pronotal declivity wide. Medium- or large-sized species, from
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(small specimens of
<taxonomicName id="84FB4D04FFB4FFD9190AFD09FCDCCE89" authority="Fairmaire, 1893" authorityName="Fairmaire" authorityYear="1893" box="[521,891,674,699]" class="Insecta" family="Anthribidae" genus="Diastatotropis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="430" phylum="Arthropoda" rank="species" species="rubricollis">
<emphasis id="718FEA95FFB4FFD9190AFD09FD05CE89" box="[521,674,675,699]" italics="true" pageId="1" pageNumber="430">D. rubricollis</emphasis>
(Fairmaire, 1893))
</taxonomicName>
to more than
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(
<taxonomicName id="84FB4D04FFB4FFD91F7FFD09FF74CEED" authority="Waterhouse, 1877" authorityName="Waterhouse" authorityYear="1877" class="Insecta" family="Anthribidae" genus="Diastatotropis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="430" phylum="Arthropoda" rank="species" species="olivaceus">
<emphasis id="718FEA95FFB4FFD91F7FFD09FAADCE89" box="[1148,1290,675,699]" italics="true" pageId="1" pageNumber="430">D. olivaceus</emphasis>
Waterhouse, 1877
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>

212
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<mods:title id="76502FAE7E50E5BF6BBB6569E541C925">Two new species of Diastatotropis Lacordaire (Coleoptera: Anthribidae) from east Madagascar with a key to species of the genus</mods:title>
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<emphasis id="718FEA95FFB4FFD91B94FC8FFE3ACF0D" bold="true" box="[151,413,805,831]" italics="true" pageId="1" pageNumber="430">Diastatotropis lepidus</emphasis>
Trýzna &amp; Baňař
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<taxonomicNameLabel id="6ABC57EEFFB4FFD91979FC8FFD70CF0D" box="[634,727,805,831]" pageId="1" pageNumber="430" rank="species">sp. nov.</taxonomicNameLabel>
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(
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,
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)
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<paragraph id="43443687FFB4FFD91B94FC23FBCFCF93" blockId="1.[151,1437,905,2038]" box="[151,1128,905,930]" pageId="1" pageNumber="430">
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<emphasis id="718FEA95FFB4FFD91B94FC23FE92CF90" bold="true" box="[151,309,905,930]" pageId="1" pageNumber="430">
<typeStatus id="9C408825FFB4FFD91B94FC23FF75CF90" box="[151,210,905,930]" pageId="1" pageNumber="430">Type</typeStatus>
locality.
</emphasis>
South-east
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,
<collectingRegion id="813FF865FFB4FFD9194EFC23FCAFCF90" box="[589,776,905,930]" country="Madagascar" name="Toliara" pageId="1" pageNumber="430">Toliara province</collectingRegion>
,
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,
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.
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material.
<typeStatus id="9C408825FFB4FFD91A78FC06FE4FCFF7" box="[379,488,940,965]" pageId="1" pageNumber="430" type="holotype">Holotype</typeStatus>
(female): SE
<collectingCountry id="3BEC7617FFB4FFD91989FC06FCFECFF7" box="[650,857,940,965]" name="Madagascar" pageId="1" pageNumber="430">MADAGASCAR</collectingCountry>
,
<collectingRegion id="813FF865FFB4FFD91866FC06FC68CFF6" box="[869,975,940,965]" country="Madagascar" name="Toliara" pageId="1" pageNumber="430">TOLIARA</collectingRegion>
</emphasis>
PROVINCE: ‘
<collectingCounty id="AA254E0BFFB4FFD91F5CFC06FB19CFF7" box="[1119,1214,940,965]" pageId="1" pageNumber="430">Chaines</collectingCounty>
anosyennes [Sic!] /
<collectingMunicipality id="A320ACFDFFB4FFD91B94FC7AFF44CFDB" box="[151,227,976,1001]" pageId="1" pageNumber="430">Massif</collectingMunicipality>
nord,
<quantity id="84039B62FFB4FFD91A3BFC7AFE34CFDA" box="[312,403,976,1001]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="1" pageNumber="430" unit="m" value="1050.0">
<elevation id="C8D6D1B4FFB4FFD91A3BFC7AFE34CFDA" box="[312,403,976,1001]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="1" pageNumber="430" unit="m" value="1050.0">1050 m</elevation>
</quantity>
/ moyenne
<location id="4624605CFFB4FFD9192AFC7AFD1ECFDB" LSID="urn:lsid:plazi:treatment:CB528791FFB4FFDB1B94FC8FFACFCE0F:4624605CFFB4FFD9192AFC7AFD1ECFDB" box="[553,697,976,1001]" country="Madagascar" county="Chaines" municipality="Massif" name="Ranomandry" pageId="1" pageNumber="430" stateProvince="TOLIARA PROVINCE">Ranomandry</location>
/ 11/
<date id="37451047FFB4FFD919FCFC7BFC2FCFDB" box="[767,904,977,1001]" pageId="1" pageNumber="430" value="1971-11-30">
<collectingDate id="2701E9AFFFB4FFD919FCFC7BFC2FCFDB" box="[767,904,977,1001]" pageId="1" pageNumber="430" value="1971-11-30">30-XI-1971</collectingDate>
</date>
’ [p] (
<collectionCode id="25EAAE42FFB4FFD918CCFC7BFB89CFDB" box="[975,1070,977,1001]" pageId="1" pageNumber="430">BSNPC</collectionCode>
)
</materialsCitation>
. Red label [p] HOLOTYPE /
<emphasis id="718FEA95FFB4FFD91B94FC5FFE39C83F" box="[151,414,1013,1037]" italics="true" pageId="1" pageNumber="430">
<taxonomicName id="84FB4D04FFB4FFD91B94FC5FFE9FC83E" box="[151,312,1013,1036]" class="Insecta" family="Anthribidae" genus="Diastatotropis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="430" phylum="Arthropoda" rank="genus">Diastatotropis</taxonomicName>
/
<taxonomicName id="84FB4D04FFB4FFD91A4EFC5FFE39C83F" box="[333,414,1013,1037]" class="Insecta" family="Anthribidae" genus="Diastatotropis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="430" phylum="Arthropoda" rank="species" species="lepidus" status="sp. nov.">lepidus</taxonomicName>
</emphasis>
<emphasis id="718FEA95FFB4FFD91AA6FC5EFE58C83F" bold="true" box="[421,511,1012,1037]" pageId="1" pageNumber="430">
<taxonomicNameLabel id="6ABC57EEFFB4FFD91AA6FC5EFE58C83F" box="[421,511,1012,1037]" pageId="1" pageNumber="430" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
/ M. Trýzna &amp; P. Baňař det., 2016.
</paragraph>
</subSubSection>
<subSubSection id="0BE1650CFFB4FFDB1BC4FBB2FE20CD57" lastPageId="3" lastPageNumber="432" pageId="1" pageNumber="430" type="description">
<paragraph id="43443687FFB4FFD91BC4FBB2FB7BC8F2" blockId="1.[151,1437,905,2038]" pageId="1" pageNumber="430">
<emphasis id="718FEA95FFB4FFD91BC4FBB2FD8DC803" bold="true" box="[199,554,1048,1073]" pageId="1" pageNumber="430">
Description. Female
<typeStatus id="9C408825FFB4FFD91AC0FBB2FD8DC803" box="[451,554,1048,1073]" pageId="1" pageNumber="430" type="holotype">holotype</typeStatus>
</emphasis>
<emphasis id="718FEA95FFB4FFD91937FBB3FD71C802" box="[564,726,1049,1072]" italics="true" pageId="1" pageNumber="430">Measurements</emphasis>
(in mm): Total body length—8.32. Head: total length—1.82; length of rostrum—1.07; maximum width of rostrum—1.27; length of eye—0.78; maximum width across eyes— 1.62; minimum distance between eyes—0.78. Antenna: length of segments: II—0.24, III—0.31, IV—0.22, V— 0.22, VI—0.16, VII—0.16, VIII—0.11, IX—0.38, X—0.20, XI—0.36. Pronotum: maximum length—1.76; width at carina—2.27; minimum width—1.53. Elytra: maximum length—4.32; maximum width—2.75.
</paragraph>
<paragraph id="43443687FFB4FFD91BC4FB61FB69C8D6" blockId="1.[151,1437,905,2038]" box="[199,1230,1227,1252]" pageId="1" pageNumber="430">
<emphasis id="718FEA95FFB4FFD91BC4FB61FEFFC8D6" bold="true" box="[199,344,1227,1252]" pageId="1" pageNumber="430">Colouration</emphasis>
of the cuticle of entire body dark brown, tarsomeres and antennae light brown.
</paragraph>
<paragraph id="43443687FFB4FFD91BC4FB44FEEBCB70" blockId="1.[151,1437,905,2038]" pageId="1" pageNumber="430">
<emphasis id="718FEA95FFB4FFD91BC4FB44FE9DC935" bold="true" box="[199,314,1262,1287]" pageId="1" pageNumber="430">Vestiture.</emphasis>
Head with indistinct sparse yellowish appressed setae along central longitudinal carina on rostrum and in depression above this carina. Frons with only very short and almost invisible setae. Lateral sides of rostrum almost bare, this part only with coarse sculpture. Labrum yellowish with a few longer light setae. Vertex with two stripes behind lateral carinae of rostrum and continued to inner margin of eyes, these stripes bright yellow with very dense appressed setae. Antennae with indistinct light pubescence, only with very short, soft, sparse setae. Pronotum with two wider distinct longitudinal stripes posteriorly extended to transverse pronotal carina. Pronotal declivity with very narrow stripe in central part. All stripes covered with dense yellow setae. Rest of pronotum with delicate sculpture, covered with very short dark brown appressed setae. Elytra with wide humeral part covered with bright yellow setae. Each elytron with two distinct longitudinal stripes; one stripe on the second elytral interval extended from indistinct subbasal tubercle, continuing to preapical part but not reaching apical margin of elytra; the second stripe in lateral part on the eighth elytral interval reaching the tip of elytron. Rest of elytra covered with dense brown appressed setae. Humeral angle without yellow colouration, with only sparse short brown setae. Scutellum with yellow setae. All legs covered with dense fine yellowish setae. Tibiae covered with dense indistinct yellowish setae appressed in proximal part and subdecumbent in distal part. All tibiae on dorsal side with small spot of yellow setae in proximal third. Tarsomeres with yellowish setae. Lateral parts of venter of thorax and abdominal sternites with dense pubescence with short brown setae. Pygidium covered with short dense brown appressed setae.
</paragraph>
<paragraph id="43443687FFB4FFDB1BC4F8E4FB73CD72" blockId="1.[151,1437,905,2038]" lastBlockId="3.[151,1437,151,573]" lastPageId="3" lastPageNumber="432" pageId="1" pageNumber="430">
<emphasis id="718FEA95FFB4FFD91BC4F8E4FE98CB55" bold="true" box="[199,319,1870,1895]" pageId="1" pageNumber="430">Structure.</emphasis>
Head appropriately long, rostrum noticeably extended apically, weakly depressed in the middle, with one short longitudinal carina in depression of rostrum, not reaching anterior margin of eyes, and two short indistinct lateral carinae. Ratio of rostrum length to maximum width 0.92. Eyes elliptical, not emarginate, dorsal ocular index 1.86. Ratio of maximum width across eyes to maximum width of rostrum 1.28.
<emphasis id="718FEA95FFB4FFD91FAFF810FAB1CBE3" box="[1196,1302,1978,2001]" italics="true" pageId="1" pageNumber="430">Antennae</emphasis>
shorter, not reaching to posterior margin of pronotum. Pedicel not robust, funicle thin, club somewhat widened, antennomere IX and XI longer than wide, antennomere X moderately wider than long. Pronotum slightly transverse, ratio of its length to width at carina 0.78, gradually extended anteriorly to quarter of its length, here widest (at the carina), then narrowed to distal part. Dorsal transverse carina subtly bisinuate with indistinct arch in middle. Lateral carina of pronotum well-developed but short, scarcely extending to half of pronotum, rounded at contact with dorsal transverse carina. Elytra suboval, ratio of the maximum length to maximum width of elytra 1.57.
</paragraph>
<caption id="1784660FFFB7FFDA1B94F81FFBD9CBDB" httpUri="https://zenodo.org/record/256121/files/figure.png" pageId="2" pageNumber="431" targetBox="[180,1347,199,1901]" targetPageId="2">
<paragraph id="43443687FFB7FFDA1B94F81FFBD9CBDB" blockId="2.[151,1436,1973,2025]" pageId="2" pageNumber="431">
<emphasis id="718FEA95FFB7FFDA1B94F81FFE9ECBF9" bold="true" box="[151,313,1973,1995]" pageId="2" pageNumber="431">FIGURES 1–4.</emphasis>
1, 3,
<taxonomicName id="84FB4D04FFB7FFDA1A72F81CFCAECBF9" authority="Tryzna &amp; Banar" authorityName="Tryzna &amp; Banar" box="[369,777,1973,1995]" class="Insecta" family="Anthribidae" genus="Diastatotropis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="431" phylum="Arthropoda" rank="species" species="lepidus" status="sp. nov.">
<emphasis id="718FEA95FFB7FFDA1A72F81CFDF6CBF9" box="[369,593,1973,1995]" italics="true" pageId="2" pageNumber="431">Diastatotropis lepidus</emphasis>
<emphasis id="718FEA95FFB7FFDA1954F81FFCAECBF9" bold="true" box="[599,777,1973,1995]" pageId="2" pageNumber="431">Trýzna &amp; Baňař</emphasis>
</taxonomicName>
<emphasis id="718FEA95FFB7FFDA180DF81FFCF9CBF9" bold="true" box="[782,862,1973,1995]" pageId="2" pageNumber="431">
<taxonomicNameLabel id="6ABC57EEFFB7FFDA180DF81FFCF9CBF9" box="[782,862,1973,1995]" pageId="2" pageNumber="431" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype female; 2, 4,
<taxonomicName id="84FB4D04FFB7FFDA1F4EF81CFF51CBDB" authority="Tryzna &amp; Banar" authorityName="Tryzna &amp; Banar" class="Insecta" family="Anthribidae" genus="Diastatotropis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="431" phylum="Arthropoda" rank="species" species="humeralis" status="sp. nov.">
<emphasis id="718FEA95FFB7FFDA1F4EF81CFAEECBF9" box="[1101,1353,1973,1995]" italics="true" pageId="2" pageNumber="431">Diastatotropis humeralis</emphasis>
<emphasis id="718FEA95FFB7FFDA1E4CF81FFF51CBDB" bold="true" pageId="2" pageNumber="431">Trýzna &amp; Baňař</emphasis>
</taxonomicName>
<emphasis id="718FEA95FFB7FFDA1BFFF879FEEBCBDB" bold="true" box="[252,332,2003,2025]" pageId="2" pageNumber="431">
<taxonomicNameLabel id="6ABC57EEFFB7FFDA1BFFF879FEEBCBDB" box="[252,332,2003,2025]" pageId="2" pageNumber="431" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype female; 1–2, dorsal habitus; 3–4, head, dorsal view. Scale bars in mm.
</paragraph>
</caption>
<paragraph id="43443687FFB6FFDB1BC4FEE6FE20CD57" blockId="3.[151,1437,151,573]" box="[199,391,332,357]" pageId="3" pageNumber="432">
<emphasis id="718FEA95FFB6FFDB1BC4FEE6FEACCD57" bold="true" box="[199,267,332,357]" pageId="3" pageNumber="432">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="0BE1650CFFB6FFDB1BC4FEC5FDD3CDE2" pageId="3" pageNumber="432" type="etymology">
<paragraph id="43443687FFB6FFDB1BC4FEC5FDDACD9F" blockId="3.[151,1437,151,573]" pageId="3" pageNumber="432">
<emphasis id="718FEA95FFB6FFDB1BC4FEC5FEE9CDBA" bold="true" box="[199,334,367,392]" pageId="3" pageNumber="432">Etymology.</emphasis>
Based on the Latin word
<taxonomicName id="84FB4D04FFB6FFDB199CFEDAFD57CDBA" box="[671,752,368,392]" class="Insecta" family="Anthribidae" genus="Diastatotropis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="species" species="lepidus">
<emphasis id="718FEA95FFB6FFDB199CFEDAFD57CDBA" box="[671,752,368,392]" italics="true" pageId="3" pageNumber="432">lepidus</emphasis>
</taxonomicName>
(= nice, good-looking, charming), from the unique and remarkable appearance of this new species.
</paragraph>
<paragraph id="43443687FFB6FFDB1BC4FE1DFDD3CDE2" blockId="3.[151,1437,151,573]" box="[199,628,439,464]" pageId="3" pageNumber="432">
<emphasis id="718FEA95FFB6FFDB1BC4FE1DFE50CDE2" bold="true" box="[199,503,439,464]" pageId="3" pageNumber="432">Collecting circumstances.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="0BE1650CFFB6FFDB1BC4FE76FD56CDC7" box="[199,753,476,501]" pageId="3" pageNumber="432" type="distribution">
<paragraph id="43443687FFB6FFDB1BC4FE76FD56CDC7" blockId="3.[151,1437,151,573]" box="[199,753,476,501]" pageId="3" pageNumber="432">
<emphasis id="718FEA95FFB6FFDB1BC4FE76FEC7CDC7" bold="true" box="[199,352,476,501]" pageId="3" pageNumber="432">Distribution.</emphasis>
Only known from the
<typeStatus id="9C408825FFB6FFDB1963FE77FD37CDC7" box="[608,656,477,501]" pageId="3" pageNumber="432">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="0BE1650CFFB6FFDB1BC4FE55FACFCE0F" pageId="3" pageNumber="432" type="diagnosis">
<paragraph id="43443687FFB6FFDB1BC4FE55FACFCE0F" blockId="3.[151,1437,151,573]" pageId="3" pageNumber="432">
<emphasis id="718FEA95FFB6FFDB1BC4FE55FE7FCE2A" bold="true" box="[199,472,511,536]" pageId="3" pageNumber="432">Differential diagnosis.</emphasis>
<taxonomicName id="84FB4D04FFB6FFDB1AEBFDAAFC62CE2A" authority="Tryzna &amp; Banar" authorityName="Tryzna &amp; Banar" box="[488,965,512,536]" class="Insecta" family="Anthribidae" genus="Diastatotropis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="species" species="lepidus" status="sp. nov.">
<emphasis id="718FEA95FFB6FFDB1AEBFDAAFD4CCE2A" box="[488,747,512,536]" italics="true" pageId="3" pageNumber="432">Diastatotropis lepidus</emphasis>
Trýzna &amp; Baňař
</taxonomicName>
<emphasis id="718FEA95FFB6FFDB18D5FE55FB9DCE2A" bold="true" box="[982,1082,511,536]" pageId="3" pageNumber="432">
<taxonomicNameLabel id="6ABC57EEFFB6FFDB18D5FE55FB9DCE2A" box="[982,1082,511,536]" pageId="3" pageNumber="432" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished by its characteristic colour pattern, not resembling any other species of the genus and by characters given in the key.
</paragraph>
</subSubSection>
</treatment>
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471
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<mods:title id="71378CBE9486928A6BE6BDBF1E70D2A3">Two new species of Diastatotropis Lacordaire (Coleoptera: Anthribidae) from east Madagascar with a key to species of the genus</mods:title>
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<paragraph id="43443687FFB6FFDB1B94FD25FD5CCE98" blockId="3.[151,763,655,715]" box="[151,763,655,682]" pageId="3" pageNumber="432">
<heading id="180C81EBFFB6FFDB1B94FD25FD5CCE98" bold="true" box="[151,763,655,682]" fontSize="11" level="1" pageId="3" pageNumber="432" reason="1">
<emphasis id="718FEA95FFB6FFDB1B94FD25FD5CCE98" bold="true" box="[151,763,655,682]" pageId="3" pageNumber="432">
<taxonomicName id="84FB4D04FFB6FFDB1B94FD25FD30CE98" authority="Tryzna &amp; Banar" authorityName="Tryzna &amp; Banar" box="[151,663,655,682]" class="Insecta" family="Anthribidae" genus="Diastatotropis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="species" species="humeralis" status="sp. nov.">
<emphasis id="718FEA95FFB6FFDB1B94FD25FE66CE9B" bold="true" box="[151,449,655,681]" italics="true" pageId="3" pageNumber="432">Diastatotropis humeralis</emphasis>
Trýzna &amp; Baňař
</taxonomicName>
<taxonomicNameLabel id="6ABC57EEFFB6FFDB199DFD3AFD5CCE98" box="[670,763,656,682]" pageId="3" pageNumber="432" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
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<paragraph id="43443687FFB6FFDB1B94FD18FEEDCEF9" blockId="3.[151,763,655,715]" box="[151,330,690,715]" pageId="3" pageNumber="432">
(
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,
<figureCitation id="DBC02A02FFB6FFDB1A15FD18FE82CEF9" box="[278,293,690,715]" captionStart="FIGURE 5" captionStartId="4.[151,250,1839,1861]" captionTargetBox="[415,1171,772,1779]" captionTargetId="figure@4.[415,1171,772,1780]" captionTargetPageId="4" captionText="FIGURE 5. Microhabitat of the type locality of Diastatotropis humeralis Trýzna &amp; Baňař sp. nov. in Andasibe – Mantadia NP, Analamazaotra forest. Black arrow indicates the position of siting holotype specimen." httpUri="https://zenodo.org/record/256122/files/figure.png" pageId="3" pageNumber="432">5</figureCitation>
,
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)
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<paragraph id="43443687FFB6FFDB1B94FD50FF13CF04" blockId="3.[151,1437,762,2011]" pageId="3" pageNumber="432">
<materialsCitation id="F3933CDAFFB6FFDB1B94FD50FF13CF04" ID-GBIF-Occurrence="1306553030" country="Madagascar" county="Analamazaotra forest" elevation="990" location="Tamatave province" municipality="Andasibe-Mantadia National Park" pageId="3" pageNumber="432" specimenCount="990" typeStatus="holotype">
<emphasis id="718FEA95FFB6FFDB1B94FD50FE92CF21" bold="true" box="[151,309,762,787]" pageId="3" pageNumber="432">
<typeStatus id="9C408825FFB6FFDB1B94FD50FF75CF21" box="[151,210,762,787]" pageId="3" pageNumber="432">Type</typeStatus>
locality.
</emphasis>
East
<collectingCountry id="3BEC7617FFB6FFDB1A71FD50FE5CCF21" box="[370,507,762,787]" name="Madagascar" pageId="3" pageNumber="432">Madagascar</collectingCountry>
,
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,
<collectingMunicipality id="A320ACFDFFB6FFDB19EAFD50FBCECF21" box="[745,1129,762,787]" pageId="3" pageNumber="432">Andasibe-Mantadia National Park</collectingMunicipality>
,
<collectingCounty id="AA254E0BFFB6FFDB1F77FD50FACFCF21" box="[1140,1384,762,787]" pageId="3" pageNumber="432">Analamazaotra forest</collectingCounty>
,
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.
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</materialsCitation>
</paragraph>
<paragraph id="43443687FFB6FFDB1BC4FCE8FD56CFD9" blockId="3.[151,1437,762,2011]" pageId="3" pageNumber="432">
<materialsCitation id="F3933CDAFFB6FFDB1BC4FCE8FE32CF91" ID-GBIF-Occurrence="1306553027" collectingDate="2016-02-05" collectionCode="BSNPC" collectorName="Andasibe-Mantadia N. P. &amp; M. Tryzna" country="Madagascar" county="Indri" elevation="990" latitude="-18.945526" location="Madagascar" longLatPrecision="1" longitude="48.421112" municipality="Analamazaotra forest" pageId="3" pageNumber="432" specimenCount="2" typeStatus="holotype">
<emphasis id="718FEA95FFB6FFDB1BC4FCE8FC4ACF68" bold="true" box="[199,1005,834,859]" pageId="3" pageNumber="432">
<typeStatus id="9C408825FFB6FFDB1BC4FCE8FEA5CF69" box="[199,258,834,859]" pageId="3" pageNumber="432">Type</typeStatus>
material.
<typeStatus id="9C408825FFB6FFDB1A82FCE8FE49CF69" box="[385,494,834,859]" pageId="3" pageNumber="432" type="holotype">Holotype</typeStatus>
(female): E
<collectingCountry id="3BEC7617FFB6FFDB1989FCE8FCFECF69" box="[650,857,834,859]" name="Madagascar" pageId="3" pageNumber="432">MADAGASCAR</collectingCountry>
, TAMATAVE
</emphasis>
PROVINCE: ‘
<location id="4624605CFFB6FFDB1F80FCE8FAB6CF69" LSID="urn:lsid:plazi:treatment:CB528791FFB6FFDE1B94FD25FA3EC916:4624605CFFB6FFDB1F80FCE8FAB6CF69" box="[1155,1297,834,859]" country="Madagascar" county="Indri" latitude="-18.945526" longLatPrecision="1" longitude="48.421112" municipality="Analamazaotra forest" name="Madagascar" pageId="3" pageNumber="432">Madagascar</location>
,
<date id="37451047FFB6FFDB1E1CFCE8FA22CF69" box="[1311,1413,834,859]" pageId="3" pageNumber="432" value="2016-02-05">
<collectingDate id="2701E9AFFFB6FFDB1E1CFCE8FA22CF69" box="[1311,1413,834,859]" pageId="3" pageNumber="432" value="2016-02-05">5.ii.2016</collectingDate>
</date>
, /
<collectorName id="EE0E5351FFB6FFDB1B94FCCCFE0BCF4C" box="[151,428,870,895]" pageId="3" pageNumber="432">Andasibe-Mantadia N.P.</collectorName>
, /
<collectingMunicipality id="A320ACFDFFB6FFDB1AC4FCCCFD1DCF4D" box="[455,698,870,895]" pageId="3" pageNumber="432">Analamazaotra forest</collectingMunicipality>
, / circuit ‘
<collectingCounty id="AA254E0BFFB6FFDB1824FCCCFCC5CF4D" box="[807,866,870,895]" pageId="3" pageNumber="432">Indri</collectingCounty>
2’ //
<geoCoordinate id="26CF5040FFB6FFDB189DFCCDFBEFCF4D" box="[926,1096,870,895]" direction="south" orientation="latitude" pageId="3" pageNumber="432" precision="1" value="-18.945526">S 18°56´43.9´´</geoCoordinate>
, /
<geoCoordinate id="26CF5040FFB6FFDB1F61FCCDFAAACF4D" box="[1122,1293,870,895]" direction="east" orientation="longitude" pageId="3" pageNumber="432" precision="1" value="48.421112">E 48°25´16.0´´</geoCoordinate>
, /
<elevation id="C8D6D1B4FFB6FFDB1E24FCCDFAD7CF4C" box="[1319,1392,871,895]" metricMagnitude="2" metricUnit="m" metricValue="9.9" pageId="3" pageNumber="432" unit="m" value="990.0">
<quantity id="84039B62FFB6FFDB1E24FCCDFAD7CF4C" box="[1319,1392,871,895]" metricMagnitude="2" metricUnit="m" metricValue="9.9" pageId="3" pageNumber="432" unit="m" value="990.0">990 m</quantity>
</elevation>
,
<collectorName id="EE0E5351FFB6FFDB1E7FFCCDFF42CF91" pageId="3" pageNumber="432">M. Trýzna</collectorName>
leg.’ (
<collectionCode id="25EAAE42FFB6FFDB1A2DFC21FE2ACF91" box="[302,397,907,931]" pageId="3" pageNumber="432">BSNPC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="F3933CDAFFB6FFDB1AA3FC20FD5FCFF5" ID-GBIF-Occurrence="1306553031" collectingDate="2001-12-17" collectingDateMax="2001-12-30" collectingDateMin="2001-12-17" collectionCode="MTDC" collectorName="J. Horak" country="Madagascar" location="Tamatave distr." pageId="3" pageNumber="432" specimenCount="1" typeStatus="paratype">
<emphasis id="718FEA95FFB6FFDB1AA3FC20FDD1CF91" bold="true" box="[416,630,906,931]" pageId="3" pageNumber="432">
<typeStatus id="9C408825FFB6FFDB1AA3FC20FDAACF91" box="[416,525,906,931]" pageId="3" pageNumber="432" type="paratype">Paratype</typeStatus>
(female)
</emphasis>
(without antennae and anterior legs): ‘E
<collectingCountry id="3BEC7617FFB6FFDB1F47FC20FB6CCF91" box="[1092,1227,906,931]" name="Madagascar" pageId="3" pageNumber="432">Madagascar</collectingCountry>
/
<location id="4624605CFFB6FFDB1FE1FC21FA2ACF91" LSID="urn:lsid:plazi:treatment:CB528791FFB6FFDE1B94FD25FA3EC916:4624605CFFB6FFDB1FE1FC21FA2ACF91" box="[1250,1421,906,931]" country="Madagascar" name="Tamatave distr." pageId="3" pageNumber="432">Tamatave distr.</location>
/ ANDASIBE,
<date id="37451047FFB6FFDB1A30FC04FE4FCFF5" box="[307,488,942,967]" pageId="3" pageNumber="432" value="2001-12-17" valueMax="2001-12-30" valueMin="2001-12-17">
<collectingDate id="2701E9AFFFB6FFDB1A30FC04FE4FCFF5" box="[307,488,942,967]" pageId="3" pageNumber="432" value="2001-12-17" valueMax="2001-12-30" valueMin="2001-12-17">17.–30.xii 2001</collectingDate>
</date>
,
<collectorName id="EE0E5351FFB6FFDB1AF0FC05FDF4CFF5" box="[499,595,942,967]" pageId="3" pageNumber="432">J. Horák</collectorName>
lgt.’ (
<collectionCode id="25EAAE42FFB6FFDB199BFC05FD48CFF5" box="[664,751,943,967]" pageId="3" pageNumber="432">MTDC</collectionCode>
)
</materialsCitation>
. Red label [p] HOLOTYPE / PARATYPE /
<emphasis id="718FEA95FFB6FFDB1FEFFC05FEA0CFD9" italics="true" pageId="3" pageNumber="432">
<taxonomicName id="84FB4D04FFB6FFDB1FEFFC05FA2ACFF4" box="[1260,1421,943,966]" class="Insecta" family="Anthribidae" genus="Diastatotropis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="genus">Diastatotropis</taxonomicName>
/
<taxonomicName id="84FB4D04FFB6FFDB1B94FC79FEA0CFD9" box="[151,263,979,1003]" class="Insecta" family="Anthribidae" genus="Diastatotropis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="species" species="humeralis" status="sp. nov.">humeralis</taxonomicName>
</emphasis>
<emphasis id="718FEA95FFB6FFDB1A0DFC78FEC0CFD9" bold="true" box="[270,359,978,1003]" pageId="3" pageNumber="432">
<taxonomicNameLabel id="6ABC57EEFFB6FFDB1A0DFC78FEC0CFD9" box="[270,359,978,1003]" pageId="3" pageNumber="432" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
/ M. Trýzna &amp; P. Baňař det., 2016.
</paragraph>
</subSubSection>
<subSubSection id="0BE1650CFFB6FFDC1BC4FC5CFE20CD57" lastPageId="4" lastPageNumber="433" pageId="3" pageNumber="432" type="description">
<paragraph id="43443687FFB6FFDB1BC4FC5CFB7BC8AD" blockId="3.[151,1437,762,2011]" pageId="3" pageNumber="432">
<emphasis id="718FEA95FFB6FFDB1BC4FC5CFD8DC83D" bold="true" box="[199,554,1014,1039]" pageId="3" pageNumber="432">
Description. Female
<typeStatus id="9C408825FFB6FFDB1AC0FC5CFD8DC83D" box="[451,554,1014,1039]" pageId="3" pageNumber="432" type="holotype">holotype</typeStatus>
</emphasis>
<emphasis id="718FEA95FFB6FFDB1937FC5DFD71C83C" box="[564,726,1015,1038]" italics="true" pageId="3" pageNumber="432">Measurements</emphasis>
(in mm): Total body length—9.14. Head: total length—2.04; length of rostrum—1.22; maximum width of rostrum—1.36; length of eye—0.82; maximum width across eyes— 1.67; minimum distance between eyes—0.78. Antenna: length of segments: II—0.29, III—0.34, IV—0.24, V— 0.24, VI—0.18, VII—0.20, VIII—0.13, IX—0.38, X—0.20, XI—0.33. Pronotum: maximum length—1.96; width at carina—2.53; minimum width—1.67. Elytra: maximum length—4.95; maximum width—3.22.
</paragraph>
<paragraph id="43443687FFB6FFDB1BC4FB00FC7FC939" blockId="3.[151,1437,762,2011]" pageId="3" pageNumber="432">
<emphasis id="718FEA95FFB6FFDB1BC4FB00FEFFC8F1" bold="true" box="[199,344,1194,1219]" pageId="3" pageNumber="432">Colouration</emphasis>
of the cuticle of entire body black, proximal part of gula, distal part of rostrum and mandible dark brown. Scape, pedicel and funicle light brown, antennal club dark brown. All femora bright pale brown, only distal part darker; tibiae and tarsomeres I–II dark brown, rest of tarsomeres pale.
</paragraph>
<paragraph id="43443687FFB6FFDB1BC4FABCFAA9CB14" blockId="3.[151,1437,762,2011]" pageId="3" pageNumber="432">
<emphasis id="718FEA95FFB6FFDB1BC4FABCFE9DC91D" bold="true" box="[199,314,1302,1327]" pageId="3" pageNumber="432">Vestiture.</emphasis>
Head almost bare, nearly invisible sparse yellowish appressed setae along central longitudinal carina on rostrum. Frons bare, without setae. Lateral sides of rostrum almost bare, only with coarse sculpture, the same sculpture also on vertex. Labrum yellowish with a few longer light setae. Vertex (as in
<taxonomicName id="84FB4D04FFB6FFDB1FB0FAF5FA80C944" box="[1203,1319,1374,1398]" class="Insecta" family="Anthribidae" genus="Diastatotropis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="species" species="lepidus">
<emphasis id="718FEA95FFB6FFDB1FB0FAF5FA80C944" box="[1203,1319,1374,1398]" italics="true" pageId="3" pageNumber="432">D. lepidus</emphasis>
</taxonomicName>
) with two stripes behind to lateral carinae of rostrum and continued to inner margin of eyes, these narrow stripes rusty-yellow with dense appressed setae. Antennae with indistinct light pubescence, only with very short, soft, sparse setae. Disk of pronotum with fine sculpture, covered with indistinct brown setae, lateral parts of pronotum with more distinct yellowish setae. Basal part of disc with narrow indistinct stripe, pronotal declivity with two triangular spots in lateral part and one narrow stripe in middle covered with rusty-yellow pubescence. Elytra with short dense black setae, each elytron in humeral part with one rusty-yellow spot extended from third to sixth elytral interval. Rest of elytra as well as humeral angle black, without any spots. Scutellum with short rusty-yellowish setae. All legs covered with dense setae. Ventral face of all femora with bright yellow pubescence on distal half and dark brown distal tip. Tibiae covered with dense brown setae appressed in proximal part and subdecumbent in distal part. All tibiae with yellowish stripe in proximal third interrupted in ventral part. Tarsomeres I–II with brownish setae, rest of antennomeres pale. Lateral parts of venter of thorax and abdominal sternites with dense pubescence and short dark setae. Pygidium covered with short dense brown appressed setae, yellowish stripe in central part.
</paragraph>
<paragraph id="43443687FFB6FFDC1BC4F898FDFACD72" blockId="3.[151,1437,762,2011]" lastBlockId="4.[151,1437,151,717]" lastPageId="4" lastPageNumber="433" pageId="3" pageNumber="432">
<emphasis id="718FEA95FFB6FFDB1BC4F898FE98CB79" bold="true" box="[199,319,1842,1867]" pageId="3" pageNumber="432">Structure.</emphasis>
Head moderately long, rostrum extended apically, weakly depressed in middle, with one longitudinal carina in depression of rostrum, almost reaching anterior margin of eyes, and two short distinct lateral carinae. Ratio of rostrum length to maximum width 0.89. Eyes elliptical, not emarginate, dorsal ocular index 1.75. Ratio of maximum width across eyes to maximum width of rostrum 1.23. Antennae shorter, not reaching posterior margin of pronotum. Pedicel not robust, funicle thin, club widened, antennomere IX and XI longer than wide, antennomere X slightly wider than long. Pronotum slightly transverse, ratio of its length to width at carina 0.77, gradually extending anteriorly to a quarter of its length, here widest, then narrowed to distal part. Dorsal transverse carina almost straight without arch in middle. Lateral carina of pronotum well-developed but short, scarcely reaching to half of pronotum, contact with dorsal transverse carina rounded. Elytra suboval, ratio of the maximum length to maximum width of elytra 1.54.
</paragraph>
<paragraph id="43443687FFB1FFDC1BC4FEE6FE20CD57" blockId="4.[151,1437,151,717]" box="[199,391,332,357]" pageId="4" pageNumber="433">
<emphasis id="718FEA95FFB1FFDC1BC4FEE6FEACCD57" bold="true" box="[199,267,332,357]" pageId="4" pageNumber="433">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="0BE1650CFFB1FFDC1BC4FEC5FF55CE2A" pageId="4" pageNumber="433" type="etymology">
<paragraph id="43443687FFB1FFDC1BC4FEC5FE8ACD9E" blockId="4.[151,1437,151,717]" pageId="4" pageNumber="433">
<emphasis id="718FEA95FFB1FFDC1BC4FEC5FEE9CDBA" bold="true" box="[199,334,367,392]" pageId="4" pageNumber="433">Etymology.</emphasis>
Based on the Latin word
<taxonomicName id="84FB4D04FFB1FFDC196FFEDAFD7BCDBA" box="[620,732,368,392]" class="Insecta" family="Anthribidae" genus="Diastatotropis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="433" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="718FEA95FFB1FFDC196FFEDAFD7BCDBA" box="[620,732,368,392]" italics="true" pageId="4" pageNumber="433">humeralis</emphasis>
</taxonomicName>
(= shoulder), due to the two well-marked rusty-yellow spots in humeral part.
</paragraph>
<paragraph id="43443687FFB1FFDC1BC4FE1DFF55CE2A" blockId="4.[151,1437,151,717]" pageId="4" pageNumber="433">
<emphasis id="718FEA95FFB1FFDC1BC4FE1DFE5BCDE2" bold="true" box="[199,508,439,464]" pageId="4" pageNumber="433">Collecting circumstances.</emphasis>
The
<typeStatus id="9C408825FFB1FFDC193DFE12FDC9CDE2" box="[574,622,440,464]" pageId="4" pageNumber="433">type</typeStatus>
specimen was collected by the authors inside secondary forest at an elevation of 990 metres by sweeping the trunk of an unidentified species of deciduous tree with diameter ca
<quantity id="84039B62FFB1FFDC1E54FE76FA3BCDC6" box="[1367,1436,476,500]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="433" unit="cm" value="10.0">10 cm</quantity>
(
<figureCitation id="DBC02A02FFB1FFDC1B9CFE55FF43CE2A" box="[159,228,511,536]" captionStart="FIGURE 5" captionStartId="4.[151,250,1839,1861]" captionTargetBox="[415,1171,772,1779]" captionTargetId="figure@4.[415,1171,772,1780]" captionTargetPageId="4" captionText="FIGURE 5. Microhabitat of the type locality of Diastatotropis humeralis Trýzna &amp; Baňař sp. nov. in Andasibe – Mantadia NP, Analamazaotra forest. Black arrow indicates the position of siting holotype specimen." httpUri="https://zenodo.org/record/256122/files/figure.png" pageId="4" pageNumber="433">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0BE1650CFFB1FFDC1BC4FD8EFE02CE52" pageId="4" pageNumber="433" type="distribution">
<paragraph id="43443687FFB1FFDC1BC4FD8EFE02CE52" blockId="4.[151,1437,151,717]" pageId="4" pageNumber="433">
<emphasis id="718FEA95FFB1FFDC1BC4FD8EFEC7CE0F" bold="true" box="[199,352,548,573]" pageId="4" pageNumber="433">Distribution.</emphasis>
This species is known only from its
<typeStatus id="9C408825FFB1FFDC1815FD8FFCE1CE0F" box="[790,838,549,573]" pageId="4" pageNumber="433">type</typeStatus>
locality in Andasibe National Park, Analamazaotra forest, east
<collectingCountry id="3BEC7617FFB1FFDC1A14FDEDFE06CE52" box="[279,417,583,608]" name="Madagascar" pageId="4" pageNumber="433">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="0BE1650CFFB1FFDC1BC4FDC6FEF6CEFF" pageId="4" pageNumber="433" type="diagnosis">
<paragraph id="43443687FFB1FFDC1BC4FDC6FEF6CEFF" blockId="4.[151,1437,151,717]" pageId="4" pageNumber="433">
<emphasis id="718FEA95FFB1FFDC1BC4FDC6FE69CEB7" bold="true" box="[199,462,620,645]" pageId="4" pageNumber="433">Differential diagnosis.</emphasis>
<taxonomicName id="84FB4D04FFB1FFDC1AD6FDC7FC0CCEB6" authority="Tryzna &amp; Banar" authorityName="Tryzna &amp; Banar" box="[469,939,620,644]" class="Insecta" family="Anthribidae" genus="Diastatotropis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="433" phylum="Arthropoda" rank="species" species="humeralis" status="sp. nov.">
<emphasis id="718FEA95FFB1FFDC1AD6FDC7FD4ACEB6" box="[469,749,620,644]" italics="true" pageId="4" pageNumber="433">Diastatotropis humeralis</emphasis>
Trýzna &amp; Baňař
</taxonomicName>
<emphasis id="718FEA95FFB1FFDC18B1FDC6FBAACEB7" bold="true" box="[946,1037,620,645]" pageId="4" pageNumber="433">
<taxonomicNameLabel id="6ABC57EEFFB1FFDC18B1FDC6FBAACEB7" box="[946,1037,620,645]" pageId="4" pageNumber="433" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from all other species of the genus mainly by the two unique rusty-yellow spots in the humeral part of the elytra and by characters given in the key.
</paragraph>
</subSubSection>
<caption id="1784660FFFB1FFDC1B94F885FC5ACB51" httpUri="https://zenodo.org/record/256122/files/figure.png" pageId="4" pageNumber="433" targetBox="[415,1171,772,1779]" targetPageId="4">
<paragraph id="43443687FFB1FFDC1B94F885FC5ACB51" blockId="4.[151,1436,1839,1891]" pageId="4" pageNumber="433">
<emphasis id="718FEA95FFB1FFDC1B94F885FEB5CB77" bold="true" box="[151,274,1839,1861]" pageId="4" pageNumber="433">FIGURE 5.</emphasis>
Microhabitat of the type locality of
<taxonomicName id="84FB4D04FFB1FFDC197DF89AFB96CB77" authority="Tryzna &amp; Banar" authorityName="Tryzna &amp; Banar" box="[638,1073,1839,1861]" class="Insecta" family="Anthribidae" genus="Diastatotropis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="433" phylum="Arthropoda" rank="species" species="humeralis" status="sp. nov.">
<emphasis id="718FEA95FFB1FFDC197DF89AFCDDCB77" box="[638,890,1839,1861]" italics="true" pageId="4" pageNumber="433">Diastatotropis humeralis</emphasis>
<emphasis id="718FEA95FFB1FFDC1883F885FB96CB77" bold="true" box="[896,1073,1839,1861]" pageId="4" pageNumber="433">Trýzna &amp; Baňař</emphasis>
</taxonomicName>
<emphasis id="718FEA95FFB1FFDC1F34F885FB20CB77" bold="true" box="[1079,1159,1839,1861]" pageId="4" pageNumber="433">
<taxonomicNameLabel id="6ABC57EEFFB1FFDC1F34F885FB20CB77" box="[1079,1159,1839,1861]" pageId="4" pageNumber="433" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in Andasibe–Mantadia NP, Analamazaotra forest. Black arrow indicates the position of siting holotype specimen.
</paragraph>
</caption>
<caption id="1784660FFFB0FFDD1B94F931FA23CA83" box="[151,1412,1691,1713]" httpUri="https://zenodo.org/record/256123/files/figure.png" pageId="5" pageNumber="434" targetBox="[306,1280,203,1659]" targetPageId="5">
<paragraph id="43443687FFB0FFDD1B94F931FA23CA83" blockId="5.[151,1412,1691,1713]" box="[151,1412,1691,1713]" pageId="5" pageNumber="434">
<emphasis id="718FEA95FFB0FFDD1B94F931FEB5CA83" bold="true" box="[151,274,1691,1713]" pageId="5" pageNumber="434">FIGURE 6.</emphasis>
Type localities of
<taxonomicName id="84FB4D04FFB0FFDD1ACDF936FCCFCA83" authority="Tryzna &amp; Banar" authorityName="Tryzna &amp; Banar" box="[462,872,1691,1713]" class="Insecta" family="Anthribidae" genus="Diastatotropis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="434" phylum="Arthropoda" rank="species" species="lepidus" status="sp. nov.">
<emphasis id="718FEA95FFB0FFDD1ACDF936FD08CA83" box="[462,687,1691,1713]" italics="true" pageId="5" pageNumber="434">Diastatotropis lepidus</emphasis>
<emphasis id="718FEA95FFB0FFDD19B5F931FCCFCA83" bold="true" box="[694,872,1691,1713]" pageId="5" pageNumber="434">Trýzna &amp; Baňař</emphasis>
</taxonomicName>
<emphasis id="718FEA95FFB0FFDD186DF931FC19CA83" bold="true" box="[878,958,1691,1713]" pageId="5" pageNumber="434">
<taxonomicNameLabel id="6ABC57EEFFB0FFDD186DF931FC19CA83" box="[878,958,1691,1713]" pageId="5" pageNumber="434" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="84FB4D04FFB0FFDD18F3F936FA8ACA83" authority="Tryzna &amp; Banar" authorityName="Tryzna &amp; Banar" box="[1008,1325,1691,1713]" class="Insecta" family="Anthribidae" genus="Diastatotropis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="434" phylum="Arthropoda" rank="species" species="humeralis" status="sp. nov.">
<emphasis id="718FEA95FFB0FFDD18F3F936FBAECA83" box="[1008,1033,1692,1713]" italics="true" pageId="5" pageNumber="434">D.</emphasis>
<emphasis id="718FEA95FFB0FFDD1F0CF931FBD3CA83" box="[1039,1140,1691,1713]" italics="true" pageId="5" pageNumber="434">humeralis</emphasis>
Trýzna &amp; Baňař
</taxonomicName>
<taxonomicNameLabel id="6ABC57EEFFB0FFDD1E30F931FA23CA83" box="[1331,1412,1691,1713]" pageId="5" pageNumber="434" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<subSubSection id="0BE1650CFFB0FFDE1B94F8A8FA3EC916" lastPageId="6" lastPageNumber="435" pageId="5" pageNumber="434" type="key">
<paragraph id="43443687FFB0FFDD1B94F8A8FCA9CB29" blockId="5.[151,782,1793,1820]" box="[151,782,1793,1820]" pageId="5" pageNumber="434">
<emphasis id="718FEA95FFB0FFDD1B94F8A8FE50CB2E" bold="true" box="[151,503,1793,1820]" pageId="5" pageNumber="434">
Key to
<taxonomicName id="84FB4D04FFB0FFDD1BF3F8ABFE3ECB29" box="[240,409,1793,1819]" class="Insecta" family="Anthribidae" genus="Diastatotropis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="718FEA95FFB0FFDD1BF3F8ABFE3ECB29" bold="true" box="[240,409,1793,1819]" italics="true" pageId="5" pageNumber="434">Diastatotropis</emphasis>
</taxonomicName>
species
</emphasis>
(see also
<bibRefCitation id="276A4B76FFB0FFDD1968F8ABFCA1CB29" author="Frieser" box="[619,774,1793,1819]" pageId="5" pageNumber="434" refString="Frieser, R. (1992) Zur Synonymie sowie zwei neue Arten der Gattung Diastatotropis Lacordaire. Coleoptera: Anthribidae. Acta Coleopterologica, 8 (1), 46 - 50." type="journal article" year="1992">Frieser, 1992</bibRefCitation>
)
</paragraph>
<key id="376A61B6FFB0FFDE1B94F8E2FA3EC916" lastPageId="6" lastPageNumber="435" pageId="5" pageNumber="434">
<keyStep id="F80F2E22FFB0FFDD1B94F8E2FA3CCB87" pageId="5" pageNumber="434">
<paragraph id="43443687FFB0FFDD1B94F8E2FA32CB48" blockId="5.[151,1435,1864,2032]" pageId="5" pageNumber="434">
<keyLead id="F80A95B2FFB0FFDD1B94F8E2FA32CB48" pageId="5" pageNumber="434">
1 Antennal club slender and cylindrical. Terminal segment elongate-oval, always distinctly longer than wide. Antennomere X as long as wide (or weakly wider than long in females of
<taxonomicName id="84FB4D04FFB0FFDD19EFF8CFFCC9CB48" box="[748,878,1892,1914]" class="Insecta" family="Anthribidae" genus="Diastatotropis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="434" phylum="Arthropoda" rank="species" species="tessellatus">
<emphasis id="718FEA95FFB0FFDD19EFF8CFFCC9CB48" box="[748,878,1892,1914]" italics="true" pageId="5" pageNumber="434">D. tessellatus</emphasis>
</taxonomicName>
)........................................... 2
</keyLead>
</paragraph>
<paragraph id="43443687FFB0FFDD1B94F829FA3CCB87" blockId="5.[151,1435,1864,2032]" pageId="5" pageNumber="434">
<keyLead id="F80A95B2FFB0FFDD1B94F829FA3CCB87" pageId="5" pageNumber="434">- Antennal club wide and stout. Terminal segment not longer than wide. Antennomere X approximately twice as wide as long.................................................................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="F80F2E22FFB0FFDD1B94F817FA3CCBC2" pageId="5" pageNumber="434">
<paragraph id="43443687FFB0FFDD1B94F817FA3CCBE0" blockId="5.[151,1435,1864,2032]" box="[151,1435,1981,2002]" pageId="5" pageNumber="434">
<keyLead id="F80A95B2FFB0FFDD1B94F817FA3CCBE0" box="[151,1435,1981,2002]" pageId="5" pageNumber="434">2 Sub-basal tubercle of elytra high and obvious............................................................... 3</keyLead>
</paragraph>
<paragraph id="43443687FFB0FFDD1B94F871FA3CCBC2" blockId="5.[151,1435,1864,2032]" box="[151,1435,2011,2032]" pageId="5" pageNumber="434">
<keyLead id="F80A95B2FFB0FFDD1B94F871FA3CCBC2" box="[151,1435,2011,2032]" pageId="5" pageNumber="434">- Sub-basal area of elytra flat.............................................................................. 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="F80F2E22FFB3FFDE1B94FF3DFA3CCD10" pageId="6" pageNumber="435">
<paragraph id="43443687FFB3FFDE1B94FF3DFA3CCCF8" blockId="6.[151,1436,151,1317]" pageId="6" pageNumber="435">
<keyLead id="F80A95B2FFB3FFDE1B94FF3DFA3CCCF8" pageId="6" pageNumber="435">
3 Central longitudinal carina of rostrum long, almost reaching to posterior margin of eyes. Elytral intervals II, IV and VI strongly elevated. Elytra spotted and tessellated..................................................
<taxonomicName id="84FB4D04FFB3FFDE1FB5FF1EFA3CCCF8" authority="Fairmaire" authorityName="Fairmaire" box="[1206,1435,180,202]" class="Insecta" family="Anthribidae" genus="Diastatotropis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="435" phylum="Arthropoda" rank="species" species="tessellatus">
<emphasis id="718FEA95FFB3FFDE1FB5FF1EFA9FCCF8" bold="true" box="[1206,1336,180,202]" italics="true" pageId="6" pageNumber="435">D. tessellatus</emphasis>
Fairmaire
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="43443687FFB3FFDE1B94FF78FA3CCD10" blockId="6.[151,1436,151,1317]" pageId="6" pageNumber="435">
<keyLead id="F80A95B2FFB3FFDE1B94FF78FA3CCD10" pageId="6" pageNumber="435">
- Central longitudinal carina of rostrum very short, reaching only to middle of rostrum. Elytral intervals flat. Elytra brown, each elytron with large yellow spot behind sub-basal tubercle, reaching from elytral interval I to IX and four small yellow spots on elytral interval II and VIII on median and preapical part......................................
<taxonomicName id="84FB4D04FFB3FFDE1FD2FEA6FA3CCD10" authority="Fairmaire" authorityName="Fairmaire" box="[1233,1435,268,290]" class="Insecta" family="Anthribidae" genus="Diastatotropis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="435" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="718FEA95FFB3FFDE1FD2FEA6FA9FCD10" bold="true" box="[1233,1336,268,290]" italics="true" pageId="6" pageNumber="435">D. elegans</emphasis>
Fairmaire
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="F80F2E22FFB3FFDE1B94FE80FA3BCDE0" pageId="6" pageNumber="435">
<paragraph id="43443687FFB3FFDE1B94FE80FA3BCD48" blockId="6.[151,1436,151,1317]" pageId="6" pageNumber="435">
<keyLead id="F80A95B2FFB3FFDE1B94FE80FA3BCD48" pageId="6" pageNumber="435">
4 Central longitudinal carina of rostrum very short, reaching only to middle of rostrum, depression here shallow. Head and disc of pronotum with fine sculpture. Elytra dark brown with wide humeral yellow part, each elytron with two longitudinal stripes on elytral interval II and VIII reaching to apical part..............................
<taxonomicName id="84FB4D04FFB3FFDE1F47FECEFAEFCD48" authority="Tryzna &amp; Banar" authorityName="Tryzna &amp; Banar" box="[1092,1352,356,378]" class="Insecta" family="Anthribidae" genus="Diastatotropis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="435" phylum="Arthropoda" rank="species" species="lepidus" status="sp. nov.">
<emphasis id="718FEA95FFB3FFDE1F47FECEFB0FCD48" bold="true" box="[1092,1192,356,378]" italics="true" pageId="6" pageNumber="435">D. lepidus</emphasis>
Trýzna &amp; Baňař
</taxonomicName>
<emphasis id="718FEA95FFB3FFDE1E4CFECEFA3BCD48" bold="true" box="[1359,1436,356,378]" pageId="6" pageNumber="435">
<taxonomicNameLabel id="6ABC57EEFFB3FFDE1E4CFECEFA3BCD48" box="[1359,1436,356,378]" pageId="6" pageNumber="435" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="43443687FFB3FFDE1B94FE28FA3BCDE0" blockId="6.[151,1436,151,1317]" pageId="6" pageNumber="435">
<keyLead id="F80A95B2FFB3FFDE1B94FE28FA3BCDE0" pageId="6" pageNumber="435">
- Central longitudinal carina of rostrum longer, almost reaching anterior margin of eyes, depression here deeper. Head and disc of pronotum with coarse sculpture. Elytra blackish with two unique rusty-yellow spots in the humeral part.....................................................................................
<taxonomicName id="84FB4D04FFB3FFDE1F24FE16FAEECDE0" authority="Tryzna &amp; Banar" authorityName="Tryzna &amp; Banar" box="[1063,1353,444,466]" class="Insecta" family="Anthribidae" genus="Diastatotropis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="435" phylum="Arthropoda" rank="species" species="humeralis" status="sp. nov.">
<emphasis id="718FEA95FFB3FFDE1F24FE16FB0FCDE0" bold="true" box="[1063,1192,444,466]" italics="true" pageId="6" pageNumber="435">D. humeralis</emphasis>
Trýzna &amp; Baňař
</taxonomicName>
<emphasis id="718FEA95FFB3FFDE1E4CFE16FA3BCDE0" bold="true" box="[1359,1436,444,466]" pageId="6" pageNumber="435">
<taxonomicNameLabel id="6ABC57EEFFB3FFDE1E4CFE16FA3BCDE0" box="[1359,1436,444,466]" pageId="6" pageNumber="435" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="F80F2E22FFB3FFDE1B94FE70FA3CCE3E" pageId="6" pageNumber="435">
<paragraph id="43443687FFB3FFDE1B94FE70FA3CCDDD" blockId="6.[151,1436,151,1317]" box="[151,1435,474,495]" pageId="6" pageNumber="435">
<keyLead id="F80A95B2FFB3FFDE1B94FE70FA3CCDDD" box="[151,1435,474,495]" pageId="6" pageNumber="435">5 Elytra spotted. Tibia brightly coloured..................................................................... 6</keyLead>
</paragraph>
<paragraph id="43443687FFB3FFDE1B94FE5DFA3CCE3E" blockId="6.[151,1436,151,1317]" box="[151,1435,503,524]" pageId="6" pageNumber="435">
<keyLead id="F80A95B2FFB3FFDE1B94FE5DFA3CCE3E" box="[151,1435,503,524]" pageId="6" pageNumber="435">- Elytra differently patterned, not spotted. Tibia dark coloured................................................... 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="F80F2E22FFB3FFDE1B94FDBFFA3BCEE8" pageId="6" pageNumber="435">
<paragraph id="43443687FFB3FFDE1B94FDBFFA3BCEB0" blockId="6.[151,1436,151,1317]" pageId="6" pageNumber="435">
<keyLead id="F80A95B2FFB3FFDE1B94FDBFFA3BCEB0" pageId="6" pageNumber="435">
6 Rostrum with depression in middle, sides of rostrum with coarse sculpture. Head, pronotum and elytra predominantly black, head and pronotum with two longitudinal whitish stripes. Each elytron with oblique short stripes in humeral parts, rest with numerous more or less irregular delicate whitish spots. All tibia and tarsomere I covered with whitish setae................................................................................................
<taxonomicName id="84FB4D04FFB3FFDE1FBAFDC6FA3BCEB0" authority="Lacordaire" authorityName="Lacordaire" box="[1209,1436,620,642]" class="Insecta" family="Anthribidae" genus="Diastatotropis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="435" phylum="Arthropoda" rank="species" species="irroratus">
<emphasis id="718FEA95FFB3FFDE1FBAFDC6FA89CEB0" bold="true" box="[1209,1326,620,642]" italics="true" pageId="6" pageNumber="435">D. irroratus</emphasis>
Lacordaire
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="43443687FFB3FFDE1B94FD20FA3BCEE8" blockId="6.[151,1436,151,1317]" pageId="6" pageNumber="435">
<keyLead id="F80A95B2FFB3FFDE1B94FD20FA3BCEE8" pageId="6" pageNumber="435">
- Rostrum without depression in middle, sides of rostrum with fine sculpture. Dorsum of head, pronotum and elytra predominantly yellowish, disc of pronotum with two large black spots. Each elytron with six large black spots. All tibia and tarsomere I covered with yellowish setae..........................................................
<taxonomicName id="84FB4D04FFB3FFDE1FC0FD6EFA3BCEE8" authority="Lacordaire" authorityName="Lacordaire" box="[1219,1436,708,730]" class="Insecta" family="Anthribidae" genus="Diastatotropis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="435" phylum="Arthropoda" rank="species" species="tigrinus">
<emphasis id="718FEA95FFB3FFDE1FC0FD6EFA89CEE8" bold="true" box="[1219,1326,708,730]" italics="true" pageId="6" pageNumber="435">D. tigrinus</emphasis>
Lacordaire
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="F80F2E22FFB3FFDE1B94FD48FA3CCF26" pageId="6" pageNumber="435">
<paragraph id="43443687FFB3FFDE1B94FD48FA3CCEC5" blockId="6.[151,1436,151,1317]" box="[151,1435,738,759]" pageId="6" pageNumber="435">
<keyLead id="F80A95B2FFB3FFDE1B94FD48FA3CCEC5" box="[151,1435,738,759]" pageId="6" pageNumber="435">7 At least pronotum always brightly red..................................................................... 8</keyLead>
</paragraph>
<paragraph id="43443687FFB3FFDE1B94FD55FA3CCF26" blockId="6.[151,1436,151,1317]" box="[151,1435,767,788]" pageId="6" pageNumber="435">
<keyLead id="F80A95B2FFB3FFDE1B94FD55FA3CCF26" box="[151,1435,767,788]" pageId="6" pageNumber="435">- Pronotum always dark, blackish or dark with greenish shine.................................................... 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="F80F2E22FFB3FFDE1B94FCB7FA3BCFB8" pageId="6" pageNumber="435">
<paragraph id="43443687FFB3FFDE1B94FCB7FA3CCF7D" blockId="6.[151,1436,151,1317]" pageId="6" pageNumber="435">
<keyLead id="F80A95B2FFB3FFDE1B94FCB7FA3CCF7D" pageId="6" pageNumber="435">
8 Head and elytra blackish. Pronotum red, only narrow anterior margin of pronotum blackish. Pronotum with dense sculpture...................................................................................
<taxonomicName id="84FB4D04FFB3FFDE1FB7FC90FA3CCF7D" authority="Fairmaire" authorityName="Fairmaire" box="[1204,1435,826,848]" class="Insecta" family="Anthribidae" genus="Diastatotropis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="435" phylum="Arthropoda" rank="species" species="rubricollis">
<emphasis id="718FEA95FFB3FFDE1FB7FC90FA9FCF62" bold="true" box="[1204,1336,826,848]" italics="true" pageId="6" pageNumber="435">D. rubricollis</emphasis>
Fairmaire
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="43443687FFB3FFDE1B94FCFDFA3BCFB8" blockId="6.[151,1436,151,1317]" pageId="6" pageNumber="435">
<keyLead id="F80A95B2FFB3FFDE1B94FCFDFA3BCFB8" pageId="6" pageNumber="435">
- Head, pronotum and elytra red, pronotum and elytra polished with fine pubescence. Pronotum with sparse sculpture................................................................................................
<taxonomicName id="84FB4D04FFB3FFDE1FFCFCDEFA3BCFB8" authority="Frieser" authorityName="Frieser" box="[1279,1436,884,906]" class="Insecta" family="Anthribidae" genus="Diastatotropis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="435" phylum="Arthropoda" rank="species" species="ruber">
<emphasis id="718FEA95FFB3FFDE1FFCFCDEFAF4CFB8" bold="true" box="[1279,1363,884,906]" italics="true" pageId="6" pageNumber="435">D. ruber</emphasis>
Frieser
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="F80F2E22FFB3FFDE1B94FC38FA3CCFF6" pageId="6" pageNumber="435">
<paragraph id="43443687FFB3FFDE1B94FC38FA3CCF95" blockId="6.[151,1436,151,1317]" box="[151,1435,914,935]" pageId="6" pageNumber="435">
<keyLead id="F80A95B2FFB3FFDE1B94FC38FA3CCF95" box="[151,1435,914,935]" pageId="6" pageNumber="435">9 Rostrum without carinnae.............................................................................. 10</keyLead>
</paragraph>
<paragraph id="43443687FFB3FFDE1B94FC05FA3CCFF6" blockId="6.[151,1436,151,1317]" box="[151,1435,943,964]" pageId="6" pageNumber="435">
<keyLead id="F80A95B2FFB3FFDE1B94FC05FA3CCFF6" box="[151,1435,943,964]" pageId="6" pageNumber="435">- Rostrum with three distinct carinae....................................................................... 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="F80F2E22FFB3FFDE1B94FC67FA3EC808" pageId="6" pageNumber="435">
<paragraph id="43443687FFB3FFDE1B94FC67FA3ECFCD" blockId="6.[151,1436,151,1317]" pageId="6" pageNumber="435">
<keyLead id="F80A95B2FFB3FFDE1B94FC67FA3ECFCD" pageId="6" pageNumber="435">
10 Elytra metallic, shiny. Central part of rostrum with very shallow depression. Slender species, ratio of maximum length of elytra to maximum width 2.0...........................................................
<taxonomicName id="84FB4D04FFB3FFDE1F90FC40FA3ECFCD" authority="Waterhouse" authorityName="Waterhouse" box="[1171,1433,1002,1024]" class="Insecta" family="Anthribidae" genus="Diastatotropis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="435" phylum="Arthropoda" rank="species" species="nitidipennis">
<emphasis id="718FEA95FFB3FFDE1F90FC40FA82C832" bold="true" box="[1171,1317,1002,1024]" italics="true" pageId="6" pageNumber="435">D. nitidipennis</emphasis>
Waterhouse
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="43443687FFB3FFDE1B94FBADFA3EC808" blockId="6.[151,1436,151,1317]" pageId="6" pageNumber="435">
<keyLead id="F80A95B2FFB3FFDE1B94FBADFA3EC808" pageId="6" pageNumber="435">
- Elytra dark brown to blackish, matte. Central part of rostrum flat, without depression. Stouter species, ratio of maximum length of elytra to maximum width 1.7.....................................................
<taxonomicName id="84FB4D04FFB3FFDE1FA2FB8EFA3EC808" authority="Waterhouse" authorityName="Waterhouse" box="[1185,1433,1060,1082]" class="Insecta" family="Anthribidae" genus="Diastatotropis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="435" phylum="Arthropoda" rank="species" species="planifrons">
<emphasis id="718FEA95FFB3FFDE1FA2FB8EFA82C808" bold="true" box="[1185,1317,1060,1082]" italics="true" pageId="6" pageNumber="435">D. planifrons</emphasis>
Waterhouse
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="F80F2E22FFB3FFDE1B94FBE8FA3CC846" pageId="6" pageNumber="435">
<paragraph id="43443687FFB3FFDE1B94FBE8FA3BC865" blockId="6.[151,1436,151,1317]" box="[151,1436,1090,1112]" pageId="6" pageNumber="435">
<keyLead id="F80A95B2FFB3FFDE1B94FBE8FA3BC865" box="[151,1436,1090,1112]" pageId="6" pageNumber="435">
11 Rostrum distinctly wider than long..........................................................
<taxonomicName id="84FB4D04FFB3FFDE1FE4FBE8FA3BC865" authority="Frieser" authorityName="Frieser" box="[1255,1436,1090,1112]" class="Insecta" family="Anthribidae" genus="Diastatotropis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="435" phylum="Arthropoda" rank="species" species="claviger">
<emphasis id="718FEA95FFB3FFDE1FE4FBE8FAF4C86A" bold="true" box="[1255,1363,1090,1112]" italics="true" pageId="6" pageNumber="435">D. claviger</emphasis>
Frieser
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="43443687FFB3FFDE1B94FBF5FA3CC846" blockId="6.[151,1436,151,1317]" box="[151,1435,1119,1140]" pageId="6" pageNumber="435">
<keyLead id="F80A95B2FFB3FFDE1B94FBF5FA3CC846" box="[151,1435,1119,1140]" pageId="6" pageNumber="435">- Rostrum distinctly longer than wide..................................................................... 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="F80F2E22FFB3FFDE1B94FBD7FA3EC916" pageId="6" pageNumber="435">
<paragraph id="43443687FFB3FFDE1B94FBD7FA3EC8FE" blockId="6.[151,1436,151,1317]" pageId="6" pageNumber="435">
<keyLead id="F80A95B2FFB3FFDE1B94FBD7FA3EC8FE" pageId="6" pageNumber="435">
12 Rostrum very narrowed in central part, all three longitudinal carinae of rostrum distinct. Antennae continuously extended from antennomere VI (only
<typeStatus id="9C408825FFB3FFDE1AB9FB30FDA9C89D" box="[442,526,1178,1199]" pageId="6" pageNumber="435" type="holotype">holotype</typeStatus>
of female is known). Elytra shorter, ratio of maximum length of elytra to maximum width 1.4. Head, pronotum and elytra predominantly dark with greenish shine........................
<taxonomicName id="84FB4D04FFB3FFDE1F92FB1DFA3EC8FE" authority="Waterhouse" authorityName="Waterhouse" box="[1169,1433,1207,1229]" class="Insecta" family="Anthribidae" genus="Diastatotropis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="435" phylum="Arthropoda" rank="species" species="crassicornis">
<emphasis id="718FEA95FFB3FFDE1F92FB1DFA82C8FF" bold="true" box="[1169,1317,1207,1229]" italics="true" pageId="6" pageNumber="435">D. crassicornis</emphasis>
Waterhouse
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="43443687FFB3FFDE1B94FB7FFA3EC916" blockId="6.[151,1436,151,1317]" pageId="6" pageNumber="435">
<keyLead id="F80A95B2FFB3FFDE1B94FB7FFA3EC916" pageId="6" pageNumber="435">
- Rostrum wide, sides parallel, not narrowed, lateral longitudinal carinae of rostrum indistinct. Funicle thin, antennal club threesegmented. Elytra longer, ratio of maximum length of elytra to maximum width 1.7. Head, pronotum and elytra predominantly black or dark brown, without shine....................................................
<taxonomicName id="84FB4D04FFB3FFDE1FAFFAA5FA3EC916" authority="Waterhouse" authorityName="Waterhouse" box="[1196,1433,1295,1317]" class="Insecta" family="Anthribidae" genus="Diastatotropis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="435" phylum="Arthropoda" rank="species" species="olivaceus">
<emphasis id="718FEA95FFB3FFDE1FAFFAA5FA82C917" bold="true" box="[1196,1317,1295,1317]" italics="true" pageId="6" pageNumber="435">D. olivaceus</emphasis>
Waterhouse
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>

54
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<mods:namePart id="5B2124419CFBF994DA39A0D453E14802">Paprocki, Henrique</mods:namePart>
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Genus
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This is a New World genus that comprises 22 species (
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), seven of which are present in Mexico (
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;
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