treatments-xml/data/70/27/87/702787A3FFD2FF7139D1FBC68D23BE1D.xml

<document id="CED6A644EC1A4D30DF85BD32D6D9516A" ID-CLB-Dataset="39278" ID-DOI="10.11646/zootaxa.4057.4.5" ID-GBIF-Dataset="c616e8c5-ff95-4a41-aeb0-b4deb492ec0c" ID-ISSN="1175-5326" ID-Zenodo-Dep="235113" ID-ZooBank="C787D40D-D3AE-4330-88F5-15C55C6CEF32" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461357467198" checkinUser="plazi" docAuthor="Garraffoni, André R. S. &amp; Melchior, Marina P." docDate="2015" docId="702787A3FFD2FF7139D1FBC68D23BE1D" docLanguage="en" docName="zt04057p568.pdf" docOrigin="Zootaxa 4057 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Heterolepidoderma mariae Garraffoni &amp; Melchior, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="558" masterDocId="8C1EFFDBFFD6FF793946FFD78E19BF01" masterDocTitle="New species and new records of freshwater Heterolepidoderma (Gastrotricha: Chaetonotidae) from Brazil with an identification key to the genus" masterLastPageNumber="568" masterPageNumber="551" pageNumber="555" updateTime="1698661571713" updateUser="ExternalLinkService">
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<mods:title id="8703AAF0FFA37408EC9DE539F468CF9F">New species and new records of freshwater Heterolepidoderma (Gastrotricha: Chaetonotidae) from Brazil with an identification key to the genus</mods:title>
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<mods:namePart id="906536A91A26E4E1670439A51FC4E408">Garraffoni, André R. S.</mods:namePart>
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<mods:namePart id="7F8E0A1415242FB05327312C0A011AE8">Melchior, Marina P.</mods:namePart>
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<treatment id="702787A3FFD2FF7139D1FBC68D23BE1D" ID-DOI="http://doi.org/10.5281/zenodo.6116108" ID-GBIF-Taxon="119647512" ID-Zenodo-Dep="6116108" LSID="urn:lsid:plazi:treatment:702787A3FFD2FF7139D1FBC68D23BE1D" httpUri="http://treatment.plazi.org/id/702787A3FFD2FF7139D1FBC68D23BE1D" lastPageId="8" lastPageNumber="558" pageId="4" pageNumber="555">
<subSubSection id="B094653EFFD2FF7D39D1FBC68F7EBB4D" pageId="4" pageNumber="555" type="nomenclature">
<paragraph id="F83136B5FFD2FF7D39D1FBC68C59BB2A" blockId="4.[151,576,1041,1101]" box="[151,576,1041,1067]" pageId="4" pageNumber="555">
<heading id="A37981D9FFD2FF7D39D1FBC68C59BB2A" bold="true" box="[151,576,1041,1067]" fontSize="11" level="1" pageId="4" pageNumber="555" reason="1">
<emphasis id="CAFAEAA7FFD2FF7D39D1FBC68C59BB2A" bold="true" box="[151,576,1041,1067]" pageId="4" pageNumber="555">
<taxonomicName id="3F8E4D36FFD2FF7D39D1FBC68FC5BB2A" ID-CoL="6LSRY" box="[151,476,1041,1067]" family="Chaetonotidae" genus="Heterolepidoderma" kingdom="Animalia" order="Chaetonotida" pageId="4" pageNumber="555" phylum="Gastrotricha" rank="species" species="mariae" status="sp. nov.">
<emphasis id="CAFAEAA7FFD2FF7D39D1FBC68FC5BB2A" bold="true" box="[151,476,1041,1067]" italics="true" pageId="4" pageNumber="555">Heterolepidoderma mariae</emphasis>
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<taxonomicNameLabel id="D1C957DCFFD2FF7D38A5FBC68C59BB2A" box="[483,576,1041,1067]" pageId="4" pageNumber="555" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
<paragraph id="F83136B5FFD2FF7D39D1FBE38F7EBB4D" blockId="4.[151,576,1041,1101]" box="[151,359,1076,1101]" pageId="4" pageNumber="555">
(
<figureCitation id="60B52A30FFD2FF7D39D9FBE38F15BB4C" box="[159,268,1076,1101]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStartId-0="5.[151,250,1518,1540]" captionStartId-1="6.[151,250,1843,1865]" captionStartId-2="7.[151,250,1430,1452]" captionStartId-3="8.[151,250,1023,1045]" captionTargetBox-0="[237,1376,750,1472]" captionTargetBox-1="[453,1132,481,1820]" captionTargetBox-2="[150,1436,193,1407]" captionTargetBox-3="[150,1436,339,1001]" captionTargetId-0="figure@5.[210,1396,748,1480]" captionTargetId-1="figure@6.[453,1134,481,1821]" captionTargetId-2="figure@7.[130,1457,193,1408]" captionTargetId-3="figure@8.[150,1436,339,1001]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionText-0="FIGURE 3. Schematic drawings of Heterolepidoderma mariae sp. nov. A) Dorsal view. B) Ventral view. The scales are not in scale. Scale bars: 25 Μm." captionText-1="FIGURE 4. Heterolepidoderma mariae sp. nov. holotype. DIC photomicrographs: A) Habitus. B) Dorsal view of distribution and shape of scales. C) Ventral view showing the paired keeled scales at the base of the furca. D) Ventral view showing the interciliary scales. E) Ventral view showing the connection between the parallel rows of cilia. Scale bars: 22 Μm." captionText-2="FIGURE 5. Heterolepidoderma mariae sp. nov. paratype. DIC photomicrographs: A) Dorsal view of distribution and shape of scales. B) Ventral view of distribution and shape of scales and cilia. White arrow indicates hypostomium. Scale bars: 22 Μm." captionText-3="FIGURE 6. Heterolepidoderma mariae sp. nov. SEM photomicrographs: A) Ventrolateral view of anterior end of the trunk. B) Ventrolateral view of anterior end of the trunk. White arrow indicates the connection between the parallel rows of cilia. C) Close-up of dorsal region of anterior end of the trunk, showing the keeled scales. D) Dorsal view of distribution and shape of scales. E) Close-up of dorsal region of the trunk, showing the keeled scales. F) Close-up of dorsal region of posterior end, showing the keeled scales. Scale bars: A, B, 5 Μm; C, 4 Μm; D, 10 Μm; E, F, 2 Μm." httpUri-0="https://zenodo.org/record/235116/files/figure.png" httpUri-1="https://zenodo.org/record/235117/files/figure.png" httpUri-2="https://zenodo.org/record/235118/files/figure.png" httpUri-3="https://zenodo.org/record/235119/files/figure.png" pageId="4" pageNumber="555">Figs. 3–6</figureCitation>
; 
<tableCitation id="B50C030EFFD2FF7D3851FBE38F47BB4D" box="[279,350,1076,1101]" captionStart="TABLE 2" captionStartId="5.[151,239,152,175]" captionTargetBox="[159,1308,235,670]" captionTargetPageId="5" captionText="TABLE 2. Morphometrical features of Heterolepidoderma mariae sp. nov. X = mean; N = number of specimens measured." httpUri="http://table.plazi.org/id/ACF1663DFFD3FF7C39D1FF4F8F1DBFCE" pageId="4" pageNumber="555" tableUuid="ACF1663DFFD3FF7C39D1FF4F8F1DBFCE">Tab. 2</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B094653EFFD2FF7D39D1FBAB8AFCBA91" pageId="4" pageNumber="555" type="materials_examined">
<paragraph id="F83136B5FFD2FF7D39D1FBAB8CB7BA48" blockId="4.[151,1437,1148,1965]" pageId="4" pageNumber="555">
<emphasis id="CAFAEAA7FFD2FF7D39D1FBAB8F5DBB94" bold="true" box="[151,324,1148,1173]" pageId="4" pageNumber="555">
<typeStatus id="27358817FFD2FF7D39D1FBAB8ECBBB94" box="[151,210,1148,1173]" pageId="4" pageNumber="555">Type</typeStatus>
material.
</emphasis>
<typeStatus id="27358817FFD2FF7D380DFBAB8FAEBB94" box="[331,439,1148,1173]" pageId="4" pageNumber="555" type="holotype">Holotype</typeStatus>
. Photographs of one specimen (adult), collected from an urban lagoon on 
<date id="8C301075FFD2FF7D3DA8FBAB8B62BB94" box="[1262,1403,1148,1173]" pageId="4" pageNumber="555" value="2015-06-26">26/Jun/2015</date>
, in floating vegetation with 
<taxonomicName id="3F8E4D36FFD2FF7D38FCFB778C21BBB9" box="[442,568,1184,1208]" class="Liliopsida" family="Pontederiaceae" genus="Eichhornia" kingdom="Plantae" order="Commelinales" pageId="4" pageNumber="555" phylum="Tracheophyta" rank="genus">
<emphasis id="CAFAEAA7FFD2FF7D38FCFB778C21BBB9" box="[442,568,1184,1208]" italics="true" pageId="4" pageNumber="555">Eichhornia</emphasis>
</taxonomicName>
sp. in the city of Paulínia, 
<collectingCountry id="80997625FFD2FF7D3ACAFB778DCABBB8" box="[908,979,1184,1209]" name="Brazil" pageId="4" pageNumber="555">Brazil</collectingCountry>
, available at the Museum of Zoology, University of Campinas, 
<collectingCountry id="80997625FFD2FF7D388CFB138C08BBDC" box="[458,529,1220,1245]" name="Brazil" pageId="4" pageNumber="555">Brazil</collectingCountry>
, under accession number 
<collectionCode id="9E9FAE70FFD2FF7D3A13FB138D87BBDD" box="[853,926,1220,1244]" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="4" pageNumber="555">ZUEC</collectionCode>
<collectionCode id="9E9FAE70FFD2FF7D3AEBFB138DF1BBDD" box="[941,1000,1220,1244]" pageId="4" pageNumber="555">GCH</collectionCode>
19. 
<typeStatus id="27358817FFD2FF7D3D6DFB128A85BBDC" box="[1067,1180,1221,1245]" pageId="4" pageNumber="555" type="paratype">Paratypes</typeStatus>
. Photographs of five specimens (adults), collected from an urban lagoon on 
<date id="8C301075FFD2FF7D3A43FB3F8D83BA00" box="[773,922,1256,1281]" pageId="4" pageNumber="555" value="2015-03-03">03/Mar/2015</date>
, in floating vegetation with 
<taxonomicName id="3F8E4D36FFD2FF7D3D9EFB3F8B4FBA01" box="[1240,1366,1256,1280]" class="Liliopsida" family="Pontederiaceae" genus="Eichhornia" kingdom="Plantae" order="Commelinales" pageId="4" pageNumber="555" phylum="Tracheophyta" rank="genus">
<emphasis id="CAFAEAA7FFD2FF7D3D9EFB3F8B4FBA01" box="[1240,1366,1256,1280]" italics="true" pageId="4" pageNumber="555">Eichhornia</emphasis>
</taxonomicName>
sp. in the city of Paulínia, 
<collectingCountry id="80997625FFD2FF7D383BFADB8FDDBA24" box="[381,452,1292,1317]" name="Brazil" pageId="4" pageNumber="555">Brazil</collectingCountry>
, available at the Museum of Zoology, University of Campinas, 
<collectingCountry id="80997625FFD2FF7D3DD0FADB8AC4BA24" box="[1174,1245,1292,1317]" name="Brazil" pageId="4" pageNumber="555">Brazil</collectingCountry>
, under accession numbers 
<collectionCode id="9E9FAE70FFD2FF7D39B8FAE78F5EBA49" box="[254,327,1328,1352]" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="4" pageNumber="555">ZUEC</collectionCode>
<collectionCode id="9E9FAE70FFD2FF7D3808FAE78F90BA49" box="[334,393,1328,1352]" pageId="4" pageNumber="555">GCH</collectionCode>
20, 21, 22, 23, 24 and 25.
</paragraph>
<paragraph id="F83136B5FFD2FF7D3981FA838AFCBA91" blockId="4.[151,1437,1148,1965]" pageId="4" pageNumber="555">
<emphasis id="CAFAEAA7FFD2FF7D3981FA838FA2BA6C" bold="true" box="[199,443,1364,1389]" pageId="4" pageNumber="555">Additional material.</emphasis>
Three specimens were examined alive with a compound microscope and are no longer extant (collected at Broa Dam, city of São Carlos, 
<collectingCountry id="80997625FFD2FF7D3B8AFAAF8D0CBA90" box="[716,789,1400,1425]" name="Brazil" pageId="4" pageNumber="555">Brazil</collectingCountry>
); two specimens were prepared for SEM.
</paragraph>
</subSubSection>
<subSubSection id="B094653EFFD2FF7D3981FA4B8CE5B968" pageId="4" pageNumber="555" type="etymology">
<paragraph id="F83136B5FFD2FF7D3981FA4B8CFEBAFC" blockId="4.[151,1437,1148,1965]" pageId="4" pageNumber="555">
<emphasis id="CAFAEAA7FFD2FF7D3981FA4B8F57BAB4" bold="true" box="[199,334,1436,1461]" pageId="4" pageNumber="555">Etymology.</emphasis>
The specific epithet honors Dr. Maria Balsamo, who has greatly contributed to our understanding of freshwater gastrotrichs. She was also very kind and helpful to the first author (ARSG) when he first started studying this tiny, funny-looking worms 7 years ago.
</paragraph>
<paragraph id="F83136B5FFD2FF7D3981F9DF8CE5B968" blockId="4.[151,1437,1148,1965]" pageId="4" pageNumber="555">
<emphasis id="CAFAEAA7FFD2FF7D3981F9DF8F8EB920" bold="true" box="[199,407,1544,1569]" pageId="4" pageNumber="555">Autapomorphies.</emphasis>
Dorsal body surface covered by keeled scales of three different shapes, on the head a thin line of cilia connecting the two lomgitudinal bands of ventral cilia, interciliary area with elliptical keeled scales with short spines, with pharyngeal field portion naked.
</paragraph>
</subSubSection>
<subSubSection id="B094653EFFD2FF7D3981F9A38C94B8AC" pageId="4" pageNumber="555" type="diagnosis">
<paragraph id="F83136B5FFD2FF7D3981F9A38C94B8AC" blockId="4.[151,1437,1148,1965]" pageId="4" pageNumber="555">
<emphasis id="CAFAEAA7FFD2FF7D3981F9A38F58B98C" bold="true" box="[199,321,1652,1677]" pageId="4" pageNumber="555">Diagnosis.</emphasis>
Small species, 129–151 Μm in total length. Body width 9–11 Μm, 19–25 Μm, 27–31 Μm, 16–17 Μm and 11–14 Μm at head, neck, trunk, base of furca and adhesive tubes, respectively. Three-lobed head with developed cephalion, pleurae and hypostomium. Anterior sensory bristles anchored by papillae and posterior sensory bristles anchored by double-keeled scales. Straight furca with rather thin adhesive tubes pointing slightly outward at distal end. Dorsal body surface covered by keeled scales of three different shapes, distributed into 20– 22 longitudinal rows of 34–38 scales each. Columns converge on the narrow parts of the body. Ventral interciliary area with round keeled scales, pharyngeal field portion naked, and one pair of large keeled scales. Ventral ciliation in two longitudinal bands, connected in the anterior part of the body. Subterminal mouth. Pharynx 35–38 Μm in length, with anterior and posterior swellings.
</paragraph>
</subSubSection>
<caption id="ACF1663DFFD3FF7C39D1FF4F8F1DBFCE" ID-Table-UUID="ACF1663DFFD3FF7C39D1FF4F8F1DBFCE" httpUri="http://table.plazi.org/id/ACF1663DFFD3FF7C39D1FF4F8F1DBFCE" pageId="5" pageNumber="556" targetBox="[159,1308,235,670]" targetIsTable="true" targetPageId="5">
<paragraph id="F83136B5FFD3FF7C39D1FF4F8F1DBFCE" blockId="5.[151,1436,151,207]" pageId="5" pageNumber="556">
<emphasis id="CAFAEAA7FFD3FF7C39D1FF4F8F09BFAE" bold="true" box="[151,272,152,175]" pageId="5" pageNumber="556">TABLE 2.</emphasis>
Morphometrical features of 
<taxonomicName id="3F8E4D36FFD3FF7C3B1BFF408D9CBFAE" box="[605,901,151,175]" family="Chaetonotidae" genus="Heterolepidoderma" kingdom="Animalia" order="Chaetonotida" pageId="5" pageNumber="556" phylum="Gastrotricha" rank="species" species="mariae" status="sp. nov.">
<emphasis id="CAFAEAA7FFD3FF7C3B1BFF408D9CBFAE" box="[605,901,151,175]" italics="true" pageId="5" pageNumber="556">Heterolepidoderma mariae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D1C957DCFFD3FF7C3AD2FF4F8DE9BFAE" box="[916,1008,152,175]" pageId="5" pageNumber="556" rank="species">
<emphasis id="CAFAEAA7FFD3FF7C3AD2FF4F8DE9BFAE" bold="true" box="[916,1008,152,175]" pageId="5" pageNumber="556">sp. nov.</emphasis>
</taxonomicNameLabel>
X = mean; N=number of specimens measured.
</paragraph>
</caption>
<paragraph id="F83136B5FFD3FF7C39D9FF3C8B0EBD9F" pageId="5" pageNumber="556">
<table id="8A8EC415FFD3008639D9FF3C8B05BD9F" box="[159,1308,235,670]" gridcols="4" gridrows="11" pageId="5" pageNumber="556">
<tr id="46BE34F7FFD3008639D9FF3C8B05BE01" box="[159,1308,235,256]" gridrow="0" pageId="5" pageNumber="556">
<th id="056F5D8BFFD3008639D9FF3C8C75BE01" box="[159,620,235,256]" gridcol="0" gridrow="0" pageId="5" pageNumber="556">Features</th>
<th id="056F5D8BFFD300863AC0FF3C8A1EBE01" box="[902,1031,235,256]" gridcol="1" gridrow="0" pageId="5" pageNumber="556">Range</th>
<th id="056F5D8BFFD300863D29FF3C8AB1BE01" box="[1135,1192,235,256]" gridcol="2" gridrow="0" pageId="5" pageNumber="556">X</th>
<th id="056F5D8BFFD300863C4CFF3C8B05BE01" box="[1290,1308,235,256]" gridcol="3" gridrow="0" pageId="5" pageNumber="556">N</th>
</tr>
<tr id="46BE34F7FFD3008639D9FEC38B05BE2B" box="[159,1308,276,298]" gridrow="1" pageId="5" pageNumber="556">
<th id="056F5D8BFFD3008639D9FEC38C75BE2B" box="[159,620,276,298]" gridcol="0" gridrow="1" pageId="5" pageNumber="556">Body length</th>
<td id="056F5D8BFFD300863AC0FEC38A1EBE2B" box="[902,1031,276,298]" gridcol="1" gridrow="1" pageId="5" pageNumber="556">129–151 µm</td>
<td id="056F5D8BFFD300863D29FEC38AB1BE2B" box="[1135,1192,276,298]" gridcol="2" gridrow="1" pageId="5" pageNumber="556">138.4</td>
<td id="056F5D8BFFD300863C4CFEC38B05BE2B" box="[1290,1308,276,298]" gridcol="3" gridrow="1" pageId="5" pageNumber="556">5</td>
</tr>
<tr id="46BE34F7FFD3008639D9FEEA8B05BE52" box="[159,1308,317,339]" gridrow="2" pageId="5" pageNumber="556">
<th id="056F5D8BFFD3008639D9FEEA8C75BE52" box="[159,620,317,339]" gridcol="0" gridrow="2" pageId="5" pageNumber="556">Length of adhesive tubes</th>
<td id="056F5D8BFFD300863AC0FEEA8A1EBE52" box="[902,1031,317,339]" gridcol="1" gridrow="2" pageId="5" pageNumber="556">11–14 µm</td>
<td id="056F5D8BFFD300863D29FEEA8AB1BE52" box="[1135,1192,317,339]" gridcol="2" gridrow="2" pageId="5" pageNumber="556">12.6</td>
<td id="056F5D8BFFD300863C4CFEEA8B05BE52" box="[1290,1308,317,339]" gridcol="3" gridrow="2" pageId="5" pageNumber="556">5</td>
</tr>
<tr id="46BE34F7FFD3008639D9FEB18B05BE7D" box="[159,1308,358,380]" gridrow="3" pageId="5" pageNumber="556">
<th id="056F5D8BFFD3008639D9FEB18C75BE7D" box="[159,620,358,380]" gridcol="0" gridrow="3" pageId="5" pageNumber="556">Pharynx length</th>
<td id="056F5D8BFFD300863AC0FEB18A1EBE7D" box="[902,1031,358,380]" gridcol="1" gridrow="3" pageId="5" pageNumber="556">35–38 µm</td>
<td id="056F5D8BFFD300863D29FEB18AB1BE7D" box="[1135,1192,358,380]" gridcol="2" gridrow="3" pageId="5" pageNumber="556">36.8</td>
<td id="056F5D8BFFD300863C4CFEB18B05BE7D" box="[1290,1308,358,380]" gridcol="3" gridrow="3" pageId="5" pageNumber="556">5</td>
</tr>
<tr id="46BE34F7FFD3008639D9FE478B05BEA7" box="[159,1308,400,422]" gridrow="4" pageId="5" pageNumber="556">
<th id="056F5D8BFFD3008639D9FE478C75BEA7" box="[159,620,400,422]" gridcol="0" gridrow="4" pageId="5" pageNumber="556">Diameter of mouth ring</th>
<td id="056F5D8BFFD300863AC0FE478A1EBEA7" box="[902,1031,400,422]" gridcol="1" gridrow="4" pageId="5" pageNumber="556">4 –6.4 µm</td>
<td id="056F5D8BFFD300863D29FE478AB1BEA7" box="[1135,1192,400,422]" gridcol="2" gridrow="4" pageId="5" pageNumber="556">5.2</td>
<td id="056F5D8BFFD300863C4CFE478B05BEA7" box="[1290,1308,400,422]" gridcol="3" gridrow="4" pageId="5" pageNumber="556">5</td>
</tr>
<tr id="46BE34F7FFD3008639D9FE6E8B05BECE" box="[159,1308,441,463]" gridrow="5" pageId="5" pageNumber="556">
<th id="056F5D8BFFD3008639D9FE6E8C75BECE" box="[159,620,441,463]" gridcol="0" gridrow="5" pageId="5" pageNumber="556">Cephalion length</th>
<td id="056F5D8BFFD300863AC0FE6E8A1EBECE" box="[902,1031,441,463]" gridcol="1" gridrow="5" pageId="5" pageNumber="556">9–11 µm</td>
<td id="056F5D8BFFD300863D29FE6E8AB1BECE" box="[1135,1192,441,463]" gridcol="2" gridrow="5" pageId="5" pageNumber="556">9.8</td>
<td id="056F5D8BFFD300863C4CFE6E8B05BECE" box="[1290,1308,441,463]" gridcol="3" gridrow="5" pageId="5" pageNumber="556">4</td>
</tr>
<tr id="46BE34F7FFD3008639D9FE358B05BEF9" box="[159,1308,482,504]" gridrow="6" pageId="5" pageNumber="556">
<th id="056F5D8BFFD3008639D9FE358C75BEF9" box="[159,620,482,504]" gridcol="0" gridrow="6" pageId="5" pageNumber="556">Cephalion width</th>
<td id="056F5D8BFFD300863AC0FE358A1EBEF9" box="[902,1031,482,504]" gridcol="1" gridrow="6" pageId="5" pageNumber="556">22–23 µm</td>
<td id="056F5D8BFFD300863D29FE358AB1BEF9" box="[1135,1192,482,504]" gridcol="2" gridrow="6" pageId="5" pageNumber="556">22.3</td>
<td id="056F5D8BFFD300863C4CFE358B05BEF9" box="[1290,1308,482,504]" gridcol="3" gridrow="6" pageId="5" pageNumber="556">3</td>
</tr>
<tr id="46BE34F7FFD3008639D9FDDB8B05BD23" box="[159,1308,524,546]" gridrow="7" pageId="5" pageNumber="556">
<th id="056F5D8BFFD3008639D9FDDB8C75BD23" box="[159,620,524,546]" gridcol="0" gridrow="7" pageId="5" pageNumber="556">Total number of longitudinal rows of scales</th>
<td id="056F5D8BFFD300863AC0FDDB8A1EBD23" box="[902,1031,524,546]" gridcol="1" gridrow="7" pageId="5" pageNumber="556">19–22</td>
<td id="056F5D8BFFD300863D29FDDB8AB1BD23" box="[1135,1192,524,546]" gridcol="2" gridrow="7" pageId="5" pageNumber="556">21</td>
<td id="056F5D8BFFD300863C4CFDDB8B05BD23" box="[1290,1308,524,546]" gridcol="3" gridrow="7" pageId="5" pageNumber="556">3</td>
</tr>
<tr id="46BE34F7FFD3008639D9FDE28B05BD4A" box="[159,1308,565,587]" gridrow="8" pageId="5" pageNumber="556">
<th id="056F5D8BFFD3008639D9FDE28C75BD4A" box="[159,620,565,587]" gridcol="0" gridrow="8" pageId="5" pageNumber="556">Number of scales in a single longitudinal row</th>
<td id="056F5D8BFFD300863AC0FDE28A1EBD4A" box="[902,1031,565,587]" gridcol="1" gridrow="8" pageId="5" pageNumber="556">27–30</td>
<td id="056F5D8BFFD300863D29FDE28AB1BD4A" box="[1135,1192,565,587]" gridcol="2" gridrow="8" pageId="5" pageNumber="556">28.5</td>
<td id="056F5D8BFFD300863C4CFDE28B05BD4A" box="[1290,1308,565,587]" gridcol="3" gridrow="8" pageId="5" pageNumber="556">4</td>
</tr>
<tr id="46BE34F7FFD3008639D9FD898B05BD75" box="[159,1308,606,628]" gridrow="9" pageId="5" pageNumber="556">
<th id="056F5D8BFFD3008639D9FD898C75BD75" box="[159,620,606,628]" gridcol="0" gridrow="9" pageId="5" pageNumber="556">Length of neck scales</th>
<td id="056F5D8BFFD300863AC0FD898A1EBD75" box="[902,1031,606,628]" gridcol="1" gridrow="9" pageId="5" pageNumber="556">2–3.5 µm</td>
<td id="056F5D8BFFD300863D29FD898AB1BD75" box="[1135,1192,606,628]" gridcol="2" gridrow="9" pageId="5" pageNumber="556">2.8</td>
<td id="056F5D8BFFD300863C4CFD898B05BD75" box="[1290,1308,606,628]" gridcol="3" gridrow="9" pageId="5" pageNumber="556">6</td>
</tr>
<tr id="46BE34F7FFD3008639D9FD5F8B05BD9F" box="[159,1308,648,670]" gridrow="10" pageId="5" pageNumber="556">
<th id="056F5D8BFFD3008639D9FD5F8C75BD9F" box="[159,620,648,670]" gridcol="0" gridrow="10" pageId="5" pageNumber="556">Length of trunk scales</th>
<td id="056F5D8BFFD300863AC0FD5F8A1EBD9F" box="[902,1031,648,670]" gridcol="1" gridrow="10" pageId="5" pageNumber="556">4–6 µm</td>
<td id="056F5D8BFFD300863D29FD5F8AB1BD9F" box="[1135,1192,648,670]" gridcol="2" gridrow="10" pageId="5" pageNumber="556">5.1</td>
<td id="056F5D8BFFD300863C4CFD5F8B05BD9F" box="[1290,1308,648,670]" gridcol="3" gridrow="10" pageId="5" pageNumber="556">6</td>
</tr>
</table>
</paragraph>
<caption id="ACF1663DFFD3FF7C39D1FA398F8AB923" httpUri="https://zenodo.org/record/235116/files/figure.png" pageId="5" pageNumber="556" targetBox="[237,1376,750,1472]" targetPageId="5">
<paragraph id="F83136B5FFD3FF7C39D1FA398F8AB923" blockId="5.[151,1436,1518,1570]" pageId="5" pageNumber="556">
<emphasis id="CAFAEAA7FFD3FF7C39D1FA398F0BB902" bold="true" box="[151,274,1518,1540]" pageId="5" pageNumber="556">FIGURE 3.</emphasis>
Schematic drawings of 
<taxonomicName id="3F8E4D36FFD3FF7C3B41FA398D02B905" box="[519,795,1518,1540]" family="Chaetonotidae" genus="Heterolepidoderma" kingdom="Animalia" order="Chaetonotida" pageId="5" pageNumber="556" phylum="Gastrotricha" rank="species" species="mariae" status="sp. nov.">
<emphasis id="CAFAEAA7FFD3FF7C3B41FA398D02B905" box="[519,795,1518,1540]" italics="true" pageId="5" pageNumber="556">Heterolepidoderma mariae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D1C957DCFFD3FF7C3A67FA398D6BB902" box="[801,882,1518,1539]" pageId="5" pageNumber="556" rank="species">
<emphasis id="CAFAEAA7FFD3FF7C3A67FA398D6BB902" bold="true" box="[801,882,1518,1539]" pageId="5" pageNumber="556">sp. nov.</emphasis>
</taxonomicNameLabel>
A) Dorsal view. B) Ventral view. The scales are not in scale. Scale bars: 25 Μm.
</paragraph>
</caption>
<subSubSection id="B094653EFFD3FF7E3981F99C8F31B82A" lastPageId="7" lastPageNumber="558" pageId="5" pageNumber="556" type="description">
<paragraph id="F83136B5FFD3FF7C3981F99C8C20B9F5" blockId="5.[151,1437,1611,1996]" pageId="5" pageNumber="556">
<emphasis id="CAFAEAA7FFD3FF7C3981F99C8F40B965" bold="true" box="[199,345,1611,1636]" pageId="5" pageNumber="556">Description.</emphasis>
Description based on 
<typeStatus id="27358817FFD3FF7C3B12F99C8CA3B965" box="[596,698,1611,1636]" pageId="5" pageNumber="556" type="holotype">holotype</typeStatus>
. 
<taxonomicName id="3F8E4D36FFD3FF7C3B82F99C8DECB962" box="[708,1013,1611,1635]" family="Chaetonotidae" genus="Heterolepidoderma" kingdom="Animalia" order="Chaetonotida" pageId="5" pageNumber="556" phylum="Gastrotricha" rank="species" species="mariae" status="sp. nov.">
<emphasis id="CAFAEAA7FFD3FF7C3B82F99C8DECB962" box="[708,1013,1611,1635]" italics="true" pageId="5" pageNumber="556">Heterolepidoderma mariae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D1C957DCFFD3FF7C3ABBF99C8A4FB965" box="[1021,1110,1611,1636]" pageId="5" pageNumber="556" rank="species">
<emphasis id="CAFAEAA7FFD3FF7C3ABBF99C8A4FB965" bold="true" box="[1021,1110,1611,1636]" pageId="5" pageNumber="556">sp. nov.</emphasis>
</taxonomicNameLabel>
is a small-sized species, 138 Μm in total body length (
<figureCitation id="60B52A30FFD3FF7C38EAF9B88C0BB986" box="[428,530,1647,1672]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="2.[151,250,1914,1936]" captionStartId-1="5.[151,250,1518,1540]" captionStartId-2="6.[151,250,1843,1865]" captionTargetBox-0="[170,1416,205,1892]" captionTargetBox-1="[237,1376,750,1472]" captionTargetBox-2="[453,1132,481,1820]" captionTargetId-0="figure@2.[170,1417,193,1892]" captionTargetId-1="figure@5.[210,1396,748,1480]" captionTargetId-2="figure@6.[453,1134,481,1821]" captionTargetPageId-0="2" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 2. Photos of some sampling sites for the present study. A) Biribiri State Park, Diamantina, State of Minas Gerais. B) Sempre-Vivas Federal Park, Diamantina, State of Minas Gerais. C – D) urban lagoon, Paulínia, State of São Paulo." captionText-1="FIGURE 3. Schematic drawings of Heterolepidoderma mariae sp. nov. A) Dorsal view. B) Ventral view. The scales are not in scale. Scale bars: 25 Μm." captionText-2="FIGURE 4. Heterolepidoderma mariae sp. nov. holotype. DIC photomicrographs: A) Habitus. B) Dorsal view of distribution and shape of scales. C) Ventral view showing the paired keeled scales at the base of the furca. D) Ventral view showing the interciliary scales. E) Ventral view showing the connection between the parallel rows of cilia. Scale bars: 22 Μm." httpUri-0="https://zenodo.org/record/235115/files/figure.png" httpUri-1="https://zenodo.org/record/235116/files/figure.png" httpUri-2="https://zenodo.org/record/235117/files/figure.png" pageId="5" pageNumber="556">Figs 2–4</figureCitation>
). Head is three-lobed, epipleurae more developed than hypopleurae. Cephalion 9 Μm in width. Hypostomium 9.5 Μm in width, well developed without anterior teeth, upper part straight, lower part concave (
<figureCitation id="60B52A30FFD3FF7C3841F9608F43B9CE" box="[263,346,1719,1744]" captionStart="FIGURE 4" captionStartId="6.[151,250,1843,1865]" captionTargetBox="[453,1132,481,1820]" captionTargetId="figure@6.[453,1134,481,1821]" captionTargetPageId="6" captionText="FIGURE 4. Heterolepidoderma mariae sp. nov. holotype. DIC photomicrographs: A) Habitus. B) Dorsal view of distribution and shape of scales. C) Ventral view showing the paired keeled scales at the base of the furca. D) Ventral view showing the interciliary scales. E) Ventral view showing the connection between the parallel rows of cilia. Scale bars: 22 Μm." httpUri="https://zenodo.org/record/235117/files/figure.png" pageId="5" pageNumber="556">Figs. 4</figureCitation>
E; 5B; 6A–B). One pair of cephalic ciliary tufts with two different sizes of cilia: 8, 15 Μm, respectively. Ocellar granules absent.
</paragraph>
<paragraph id="F83136B5FFD3FF7C3981F9288ACFB8CD" blockId="5.[151,1437,1611,1996]" pageId="5" pageNumber="556">Body width 22 Μm at the head (U10), 20 Μm at the neck (U25), 29 Μm at the trunk (U54) and 13 Μm at the base of the furca (U80). Head slightly delimited from trunk by neck constriction that gradually widens into the trunk, which reaches its greatest width around U45–50. The trunk then tapers into a straight furca, 7 Μm in length, with rather thin adhesive tubes, 13.5 Μm in length, pointing slightly outward at their distal ends. Base of furca covered dorsolaterally, laterally and ventrally with keeled scales. Anterior and posterior sensory bristles present, anterior pair at U26–27 anchored by papillae and posterior pair at U84 anchored by double-keeled scales.</paragraph>
<paragraph id="F83136B5FFD0FF7F3981FF408EF9BEAC" blockId="6.[151,1436,151,429]" pageId="6" pageNumber="557">
Dorsal body surface covered by 20–22 longitudinal rows of 34–38 keeled scales with visible edges (
<figureCitation id="60B52A30FFD0FF7F3C74FF408B66BFB1" box="[1330,1407,151,176]" captionStart="FIGURE 3" captionStartId="5.[151,250,1518,1540]" captionTargetBox="[237,1376,750,1472]" captionTargetId="figure@5.[210,1396,748,1480]" captionTargetPageId="5" captionText="FIGURE 3. Schematic drawings of Heterolepidoderma mariae sp. nov. A) Dorsal view. B) Ventral view. The scales are not in scale. Scale bars: 25 Μm." httpUri="https://zenodo.org/record/235116/files/figure.png" pageId="6" pageNumber="557">Figs. 3</figureCitation>
A; 4B; 5A; 6C–F). The median row of dorsal scales is straight, except in the narrowest part of the neck and the rearmost trunk region, where it is replaced by 4 to 6 rows of diagonally arranged scales. Three different dorsal scale shapes: a) head scales small, oval with trilobate keels, 2.3–2.5 Μm in length and 1.4–2.1 Μm in width (
<figureCitation id="60B52A30FFD0FF7F3C42FED38B49BE1D" box="[1284,1360,260,285]" captionStart="FIGURE 3" captionStartId="5.[151,250,1518,1540]" captionTargetBox="[237,1376,750,1472]" captionTargetId="figure@5.[210,1396,748,1480]" captionTargetPageId="5" captionText="FIGURE 3. Schematic drawings of Heterolepidoderma mariae sp. nov. A) Dorsal view. B) Ventral view. The scales are not in scale. Scale bars: 25 Μm." httpUri="https://zenodo.org/record/235116/files/figure.png" pageId="6" pageNumber="557">Figs. 3</figureCitation>
A; 4B; 5A; 6C); b) trunk scales elongate-oval with slight constriction in the median region, straight posterior end and trilobate keels, 4–6 Μm in length and 1.5–2 Μm in width (
<figureCitation id="60B52A30FFD0FF7F3A00FE9B8D8EBE65" box="[838,919,332,357]" captionStart="FIGURE 3" captionStartId="5.[151,250,1518,1540]" captionTargetBox="[237,1376,750,1472]" captionTargetId="figure@5.[210,1396,748,1480]" captionTargetPageId="5" captionText="FIGURE 3. Schematic drawings of Heterolepidoderma mariae sp. nov. A) Dorsal view. B) Ventral view. The scales are not in scale. Scale bars: 25 Μm." httpUri="https://zenodo.org/record/235116/files/figure.png" pageId="6" pageNumber="557">Figs. 3</figureCitation>
A; 4B; 5A; 6D–E); c) posterior body scales elongate-oval with fine keels drawing out into simple fine spines, 5 Μm in length and 2 Μm in width (
<figureCitation id="60B52A30FFD0FF7F3C76FEB88B66BE89" box="[1328,1407,367,392]" captionStart="FIGURE 3" captionStartId="5.[151,250,1518,1540]" captionTargetBox="[237,1376,750,1472]" captionTargetId="figure@5.[210,1396,748,1480]" captionTargetPageId="5" captionText="FIGURE 3. Schematic drawings of Heterolepidoderma mariae sp. nov. A) Dorsal view. B) Ventral view. The scales are not in scale. Scale bars: 25 Μm." httpUri="https://zenodo.org/record/235116/files/figure.png" pageId="6" pageNumber="557">Figs. 3</figureCitation>
A; 6D,F).
</paragraph>
<caption id="ACF1663DFFD0FF7F39D1F8E48B15B887" httpUri="https://zenodo.org/record/235117/files/figure.png" pageId="6" pageNumber="557" targetBox="[453,1132,481,1820]" targetPageId="6">
<paragraph id="F83136B5FFD0FF7F39D1F8E48B15B887" blockId="6.[151,1436,1843,1926]" pageId="6" pageNumber="557">
<emphasis id="CAFAEAA7FFD0FF7F39D1F8E48F0AB849" bold="true" box="[151,275,1843,1865]" pageId="6" pageNumber="557">FIGURE 4.</emphasis>
<taxonomicName id="3F8E4D36FFD0FF7F385CF8E48C37B848" box="[282,558,1843,1865]" family="Chaetonotidae" genus="Heterolepidoderma" kingdom="Animalia" order="Chaetonotida" pageId="6" pageNumber="557" phylum="Gastrotricha" rank="species" species="mariae" status="sp. nov.">
<emphasis id="CAFAEAA7FFD0FF7F385CF8E48C37B848" box="[282,558,1843,1865]" italics="true" pageId="6" pageNumber="557">Heterolepidoderma mariae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D1C957DCFFD0FF7F3B72F8E48C9FB849" box="[564,646,1843,1864]" pageId="6" pageNumber="557" rank="species">
<emphasis id="CAFAEAA7FFD0FF7F3B72F8E48C9FB849" bold="true" box="[564,646,1843,1864]" pageId="6" pageNumber="557">sp. nov.</emphasis>
</taxonomicNameLabel>
holotype. DIC photomicrographs: A) Habitus. B) Dorsal view of distribution and shape of scales. C) Ventral view showing the paired keeled scales at the base of the furca. D) Ventral view showing the interciliary scales. E) Ventral view showing the connection between the parallel rows of cilia. Scale bars: 22 Μm.
</paragraph>
</caption>
<caption id="ACF1663DFFD1FF7E39D1FA418B9DBACB" httpUri="https://zenodo.org/record/235118/files/figure.png" pageId="7" pageNumber="558" targetBox="[150,1436,193,1407]" targetPageId="7">
<paragraph id="F83136B5FFD1FF7E39D1FA418B9DBACB" blockId="7.[151,1436,1430,1482]" pageId="7" pageNumber="558">
<emphasis id="CAFAEAA7FFD1FF7E39D1FA418F0BBAAA" bold="true" box="[151,274,1430,1452]" pageId="7" pageNumber="558">FIGURE 5.</emphasis>
<taxonomicName id="3F8E4D36FFD1FF7E385FFA418C35BAAD" box="[281,556,1430,1452]" family="Chaetonotidae" genus="Heterolepidoderma" kingdom="Animalia" order="Chaetonotida" pageId="7" pageNumber="558" phylum="Gastrotricha" rank="species" species="mariae" status="sp. nov.">
<emphasis id="CAFAEAA7FFD1FF7E385FFA418C35BAAD" box="[281,556,1430,1452]" italics="true" pageId="7" pageNumber="558">Heterolepidoderma mariae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D1C957DCFFD1FF7E3B74FA418C9BBAAA" box="[562,642,1430,1451]" pageId="7" pageNumber="558" rank="species">
<emphasis id="CAFAEAA7FFD1FF7E3B74FA418C9BBAAA" bold="true" box="[562,642,1430,1451]" pageId="7" pageNumber="558">sp. nov.</emphasis>
</taxonomicNameLabel>
paratype. DIC photomicrographs: A) Dorsal view of distribution and shape of scales. B) Ventral view of distribution and shape of scales and cilia. White arrow indicates hypostomium. Scale bars: 22 Μm.
</paragraph>
</caption>
<paragraph id="F83136B5FFD1FF7E3981FA258C0BB99A" blockId="7.[151,1437,1522,2014]" pageId="7" pageNumber="558">
The interciliary area is covered by 5–6 longitudinal rows of 14 elliptical keeled scales with short spines (1– 1.5Μm), in each longitudinal row, with pharyngeal field portion naked. The last posterior row (including ventral terminal scales) consists of four larger keeled scales, with two different sizes (approximately 2,5 and 5 Μm in length). Ventral ciliation appears as two longitudinal bands that merge on the head, forming a transverse band of cilia (
<figureCitation id="60B52A30FFD1FF7E3990F9558F3AB99B" box="[214,291,1666,1691]" captionStart="FIGURE 3" captionStartId="5.[151,250,1518,1540]" captionTargetBox="[237,1376,750,1472]" captionTargetId="figure@5.[210,1396,748,1480]" captionTargetPageId="5" captionText="FIGURE 3. Schematic drawings of Heterolepidoderma mariae sp. nov. A) Dorsal view. B) Ventral view. The scales are not in scale. Scale bars: 25 Μm." httpUri="https://zenodo.org/record/235116/files/figure.png" pageId="7" pageNumber="558">Figs. 3</figureCitation>
B; 4C–D; 5B; 6A,B).
</paragraph>
<paragraph id="F83136B5FFD1FF7E3981F9728CD3B9E2" blockId="7.[151,1437,1522,2014]" pageId="7" pageNumber="558">Mouth subterminal, 4–6.4 Μm in diameter; pharynx 35–38 Μm in length with anterior and posterior swellings. Anterior region of pharynx without cuticular rods.</paragraph>
<paragraph id="F83136B5FFD1FF7E3981F9398F31B82A" blockId="7.[151,1437,1522,2014]" pageId="7" pageNumber="558">Some specimens were in parthenogenetic phase, with the egg (60–67 Μm in length) positioned laterally in the trunk region.</paragraph>
</subSubSection>
<subSubSection id="B094653EFFD1FF713981F8E28D23BE1D" lastPageId="8" lastPageNumber="559" pageId="7" pageNumber="558" type="discussion">
<paragraph id="F83136B5FFD1FF713981F8E28D23BE1D" blockId="7.[151,1437,1522,2014]" lastBlockId="8.[151,1436,151,285]" lastPageId="8" lastPageNumber="559" pageId="7" pageNumber="558">
<emphasis id="CAFAEAA7FFD1FF7E3981F8E28F22B84F" bold="true" box="[199,315,1845,1870]" pageId="7" pageNumber="558">Remarks.</emphasis>
The genus 
<taxonomicName id="3F8E4D36FFD1FF7E3881F8E18CB8B84F" box="[455,673,1846,1870]" family="Chaetonotidae" genus="Heterolepidoderma" kingdom="Animalia" order="Chaetonotida" pageId="7" pageNumber="558" phylum="Gastrotricha" rank="genus">
<emphasis id="CAFAEAA7FFD1FF7E3881F8E18CB8B84F" box="[455,673,1846,1870]" italics="true" pageId="7" pageNumber="558">Heterolepidoderma</emphasis>
</taxonomicName>
includes 22 species that inhabit inland water. Among them, only 
<taxonomicName id="3F8E4D36FFD1FF7E39D1F88D8FD5B873" box="[151,460,1882,1906]" family="Chaetonotidae" genus="Heterolepidoderma" kingdom="Animalia" order="Chaetonotida" pageId="7" pageNumber="558" phylum="Gastrotricha" rank="species" species="mariae" status="sp. nov.">
<emphasis id="CAFAEAA7FFD1FF7E39D1F88D8FD5B873" box="[151,460,1882,1906]" italics="true" pageId="7" pageNumber="558">Heterolepidoderma mariae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D1C957DCFFD1FF7E389FF88D8C2EB872" box="[473,567,1882,1907]" pageId="7" pageNumber="558" rank="species">
<emphasis id="CAFAEAA7FFD1FF7E389FF88D8C2EB872" bold="true" box="[473,567,1882,1907]" pageId="7" pageNumber="558">sp. nov.</emphasis>
</taxonomicNameLabel>
, 
<emphasis id="CAFAEAA7FFD1FF7E3B0CF88C8CA2B873" box="[586,699,1883,1906]" italics="true" pageId="7" pageNumber="558">
<taxonomicName id="3F8E4D36FFD1FF7E3B0CF88C8CA1B873" box="[586,696,1883,1906]" family="Chaetonotidae" genus="Heterolepidoderma" kingdom="Animalia" order="Chaetonotida" pageId="7" pageNumber="558" phylum="Gastrotricha" rank="species" species="majus">H. majus</taxonomicName>
,
</emphasis>
<taxonomicName id="3F8E4D36FFD1FF7E3B8EF88C8A6CB873" authority="Kisielewski, 1981" authorityName="Kisielewski" authorityYear="1981" box="[712,1141,1882,1907]" family="Chaetonotidae" genus="Heterolepidoderma" kingdom="Animalia" order="Chaetonotida" pageId="7" pageNumber="558" phylum="Gastrotricha" rank="species" species="brevitubulatum">
<emphasis id="CAFAEAA7FFD1FF7E3B8EF88C8D80B873" box="[712,921,1882,1906]" italics="true" pageId="7" pageNumber="558">H. brevitubulatum</emphasis>
<bibRefCitation id="9C1F4B44FFD1FF7E3AE3F88D8A6CB873" author="Kisielewski" box="[933,1141,1882,1907]" pageId="7" pageNumber="558" refString="Kisielewski, J. (1981) Gastrotricha from raised and transitional peat bogs in Poland. Monografie Fauny Polski, 11, 1 - 143. [Polska Akademia Nauk, Warszawa]" type="journal article" year="1981">Kisielewski, 1981</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="3F8E4D36FFD1FF7E3DF3F88C8B5BB873" box="[1205,1346,1883,1906]" family="Chaetonotidae" genus="Heterolepidoderma" kingdom="Animalia" order="Chaetonotida" pageId="7" pageNumber="558" phylum="Gastrotricha" rank="species" species="jureiense">
<emphasis id="CAFAEAA7FFD1FF7E3DF3F88C8B5BB873" box="[1205,1346,1883,1906]" italics="true" pageId="7" pageNumber="558">H. jureiense</emphasis>
</taxonomicName>
have a clearly three-lobed head. 
<taxonomicName id="3F8E4D36FFD1FF7E3886F8A88C55B897" box="[448,588,1919,1942]" family="Chaetonotidae" genus="Heterolepidoderma" kingdom="Animalia" order="Chaetonotida" pageId="7" pageNumber="558" phylum="Gastrotricha" rank="species" species="jureiense">
<emphasis id="CAFAEAA7FFD1FF7E3886F8A88C55B897" box="[448,588,1919,1942]" italics="true" pageId="7" pageNumber="558">H. jureiense</emphasis>
</taxonomicName>
can be excluded from further comparison as it is the only species with ventrolateral lamellae among these four species. 
<taxonomicName id="3F8E4D36FFD1FF7E3BF8F8758A53B8BB" box="[702,1098,1954,1978]" family="Chaetonotidae" genus="Heterolepidoderma" kingdom="Animalia" order="Chaetonotida" pageId="7" pageNumber="558" phylum="Gastrotricha" rank="species" species="brevitubulatum">
<emphasis id="CAFAEAA7FFD1FF7E3BF8F8758A53B8BB" box="[702,1098,1954,1978]" italics="true" pageId="7" pageNumber="558">Heterolepidoderma brevitubulatum</emphasis>
</taxonomicName>
has adhesive tubes extremely short and slightly bifurcate, whereas in 
<taxonomicName id="3F8E4D36FFD1FF7E3B18F8108CC8B8DF" box="[606,721,1991,2014]" family="Chaetonotidae" genus="Heterolepidoderma" kingdom="Animalia" order="Chaetonotida" pageId="7" pageNumber="558" phylum="Gastrotricha" rank="species" species="mariae" status="sp. nov.">
<emphasis id="CAFAEAA7FFD1FF7E3B18F8108CC8B8DF" box="[606,721,1991,2014]" italics="true" pageId="7" pageNumber="558">H. mariae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D1C957DCFFD1FF7E3B9DF8128D2FB8DF" box="[731,822,1989,2014]" pageId="7" pageNumber="558" rank="species">
<emphasis id="CAFAEAA7FFD1FF7E3B9DF8128D2FB8DF" bold="true" box="[731,822,1989,2014]" pageId="7" pageNumber="558">sp. nov.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="3F8E4D36FFD1FF7E3A37F8108DC1B8DF" box="[881,984,1991,2014]" family="Chaetonotidae" genus="Heterolepidoderma" kingdom="Animalia" order="Chaetonotida" pageId="7" pageNumber="558" phylum="Gastrotricha" rank="species" species="majus">
<emphasis id="CAFAEAA7FFD1FF7E3A37F8108DC1B8DF" box="[881,984,1991,2014]" italics="true" pageId="7" pageNumber="558">H. majus</emphasis>
</taxonomicName>
, the adhesive tubes are at least 4 times longer and not bifurcate. 
<taxonomicName id="3F8E4D36FFDEFF7138F2FF4F8CFCBFAE" box="[436,741,152,176]" family="Chaetonotidae" genus="Heterolepidoderma" kingdom="Animalia" order="Chaetonotida" pageId="8" pageNumber="559" phylum="Gastrotricha" rank="species" species="mariae" status="sp. nov.">
<emphasis id="CAFAEAA7FFDEFF7138F2FF4F8CFCBFAE" box="[436,741,152,176]" italics="true" pageId="8" pageNumber="559">Heterolepidoderma mariae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D1C957DCFFDEFF713BA8FF408D51BFB1" box="[750,840,151,176]" pageId="8" pageNumber="559" rank="species">
<emphasis id="CAFAEAA7FFDEFF713BA8FF408D51BFB1" bold="true" box="[750,840,151,176]" pageId="8" pageNumber="559">sp. nov.</emphasis>
</taxonomicNameLabel>
can be distinguished from 
<taxonomicName id="3F8E4D36FFDEFF713D3AFF4E8AFABFAE" box="[1148,1251,152,176]" family="Chaetonotidae" genus="Heterolepidoderma" kingdom="Animalia" order="Chaetonotida" pageId="8" pageNumber="559" phylum="Gastrotricha" rank="species" species="majus">
<emphasis id="CAFAEAA7FFDEFF713D3AFF4E8AFABFAE" box="[1148,1251,152,176]" italics="true" pageId="8" pageNumber="559">H. majus</emphasis>
</taxonomicName>
because it has a dorsal body surface covered by keeled scales with three different shapes, as well as a transverse band of cilia on the head connecting the two bands of longitudinal ventral cilia, whereas 
<taxonomicName id="3F8E4D36FFDEFF713AD2FF368DE0BFF6" box="[916,1017,224,248]" family="Chaetonotidae" genus="Heterolepidoderma" kingdom="Animalia" order="Chaetonotida" pageId="8" pageNumber="559" phylum="Gastrotricha" rank="species" species="majus">
<emphasis id="CAFAEAA7FFDEFF713AD2FF368DE0BFF6" box="[916,1017,224,248]" italics="true" pageId="8" pageNumber="559">H. majus</emphasis>
</taxonomicName>
has only one 
<typeStatus id="27358817FFDEFF713DD5FF378ADABFF9" box="[1171,1219,224,248]" pageId="8" pageNumber="559">type</typeStatus>
of keeled scales on its dorsal surface, and its two ventral bands of cilia separate.
</paragraph>
</subSubSection>
</treatment>
</document>