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<document id="6BC3C5D692FDC2E6A278C7E10192AE13" ID-CLB-Dataset="32367" ID-DOI="10.11646/zootaxa.4306.4.3" ID-GBIF-Dataset="6abc8130-1b65-4df2-b6aa-ef3e2779ef6d" ID-ISSN="1175-5326" ID-Zenodo-Dep="845117" ID-ZooBank="28CF30B0-3477-4DBD-8883-5FA903464C00" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1503038923636" checkinUser="plazi" docAuthor="Roza, André Silva, Quintino, Hingrid Yara Souza, Mermudes, José Ricardo Miras &amp; Silveira, Luiz Felipe Lima Da" docDate="2017" docId="B61787F2DE2A7D0309A3D67BFD9CF871" docLanguage="en" docName="zootaxa.4306.4.3.pdf" docOrigin="Zootaxa 4306 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Akamboja monteirorum Roza &amp; Quintino &amp; Mermudes &amp; Silveira 2017, sp. nov." docType="treatment" docUuid="D4A334CE-4D84-4DB9-808F-739B809DBB47" docUuidSource="ZooBank" docVersion="8" lastPageNumber="515" masterDocId="4A2EFF8ADE247D0D0934D458FFA5FFE2" masterDocTitle="Akamboja gen. nov., a new genus of railroad-worm beetle endemic to the Atlantic Rainforest, with five new species (Coleoptera: Phengodidae, Mastinocerinae)" masterLastPageNumber="523" masterPageNumber="501" pageNumber="515" updateTime="1698466867872" updateUser="ExternalLinkService">
<mods:mods id="5A9166AE4EDA94E5AF7E1858079EE5AC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4EDD9A261F6CC1B5A03401386FA9D3C6">
<mods:title id="1FC75B2EA028E37F50585887482FBF0E">Akamboja gen. nov., a new genus of railroad-worm beetle endemic to the Atlantic Rainforest, with five new species (Coleoptera: Phengodidae, Mastinocerinae)</mods:title>
</mods:titleInfo>
<mods:name id="008931B523B2093C42E89461947B790D" type="personal">
<mods:role id="F7E9874BA075F2ACBF252BCB27193CEE">
<mods:roleTerm id="6DAA5B9EF728AC2F2219D370BC9EF823">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E101D5C6CC8337C0749E99F7A20C2196">Roza, André Silva</mods:namePart>
</mods:name>
<mods:name id="872806B855A9B9448645F2D67C91616F" type="personal">
<mods:role id="859EB06725044DE5C5B1F1CE1A65D096">
<mods:roleTerm id="B7AE0C5A3B984F4BCF2954818EFD847E">Author</mods:roleTerm>
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<mods:namePart id="73385BC3611144BD32897ABA3EF470E7">Quintino, Hingrid Yara Souza</mods:namePart>
</mods:name>
<mods:name id="8CF482610B082DA141D0196EEA426690" type="personal">
<mods:role id="F0950CB6EA4521DBD1E4477B35AA1B22">
<mods:roleTerm id="7D87C95D8B057DF1B7677EB4AE7FB3B6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7D368C8B5379F21171A14EC6C62440C9">Mermudes, José Ricardo Miras</mods:namePart>
</mods:name>
<mods:name id="F414CBFA920AD449E5D5CAF9CCEAF785" type="personal">
<mods:role id="42C780DE3B250793459E2EDD8622B9A5">
<mods:roleTerm id="093D24976C8DA2893B12D168899CAA63">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0190AC35CA6C41E53DD2411748DB2438">Silveira, Luiz Felipe Lima Da</mods:namePart>
</mods:name>
<mods:typeOfResource id="94ED1E1950BE9C1C84484F1038B5605B">text</mods:typeOfResource>
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<mods:titleInfo id="8033C4EA7062C710D9B3A7BDD7EDA693">
<mods:title id="CCDC7AD8442C8247A6C2159D0EF55A59">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="809C7D2F79CE377C884CC74165825C4D">
<mods:date id="C95076B8D2591F3C84A4E52B52A6779D">2017</mods:date>
<mods:detail id="A61AD3359911B236F2312C39158F9481" type="volume">
<mods:number id="E4E278D4260E3384B01F28215C978076">4306</mods:number>
</mods:detail>
<mods:detail id="64637B0E4A3AB3EA9ED06138C6C44D06" type="issue">
<mods:number id="73435B7ADF74DE9F05A290A6249EC566">4</mods:number>
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<mods:classification id="F00C20CC16C0A15B7C319ECA74E9D1BF">journal article</mods:classification>
<mods:identifier id="DDA9906A157CE82C809DF7D0F9119CCC" type="CLB-Dataset">32367</mods:identifier>
<mods:identifier id="D25D5A7B76E33E1E8FFD1216FD173763" type="DOI">10.11646/zootaxa.4306.4.3</mods:identifier>
<mods:identifier id="11A94DA03A802E2BCA5225BB5C9FA1DF" type="GBIF-Dataset">6abc8130-1b65-4df2-b6aa-ef3e2779ef6d</mods:identifier>
<mods:identifier id="AC794CD62724DFF6A40E3194640557C7" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B6B726645B5F9BB8FFCFC802A22CB712" type="Zenodo-Dep">845117</mods:identifier>
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<treatment id="B61787F2DE2A7D0309A3D67BFD9CF871" ID-DOI="http://doi.org/10.5281/zenodo.6025180" ID-GBIF-Taxon="132785598" ID-Zenodo-Dep="6025180" LSID="urn:lsid:zoobank.org:act:D4A334CE-4D84-4DB9-808F-739B809DBB47" httpUri="http://treatment.plazi.org/id/B61787F2DE2A7D0309A3D67BFD9CF871" lastPageNumber="515" pageId="14" pageNumber="515">
<subSubSection id="76A4656FDE2A7D0309A3D67BFEDCFDBD" pageId="14" pageNumber="515" type="nomenclature">
<paragraph id="3E0136E4DE2A7D0309A3D67BFBA4FDDC" blockId="14.[151,1025,547,607]" box="[151,1025,547,574]" pageId="14" pageNumber="515">
<heading id="65498188DE2A7D0309A3D67BFBA4FDDC" bold="true" box="[151,1025,547,574]" fontSize="11" level="1" pageId="14" pageNumber="515" reason="1">
<emphasis id="0CCAEAF6DE2A7D0309A3D67BFBA4FDDC" bold="true" box="[151,1025,547,574]" pageId="14" pageNumber="515">
<taxonomicName id="F9BE4D67DE2A7D0309A3D67BFE1FFDDF" authority="Roza &amp; Quintino &amp; Mermudes &amp; Silveira, 2017" authorityName="Roza &amp; Quintino &amp; Mermudes &amp; Silveira" authorityYear="2017" box="[151,442,547,573]" class="Insecta" family="Phengodidae" genus="Akamboja" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="species" species="monteirorum" status="sp. nov.">
<emphasis id="0CCAEAF6DE2A7D0309A3D67BFE1FFDDF" bold="true" box="[151,442,547,573]" italics="true" pageId="14" pageNumber="515">Akamboja monteirorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="17F9578DDE2A7D0308F5D67CFDBBFDDC" box="[449,542,548,574]" pageId="14" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
Roza, Quintino, Mermudes &amp; Silveira
</emphasis>
</heading>
</paragraph>
<paragraph id="3E0136E4DE2A7D0309A3D61EFEDCFDBD" blockId="14.[151,1025,547,607]" box="[151,377,582,607]" pageId="14" pageNumber="515">
(
<figureCitation id="A6852A61DE2A7D0309ABD61EFEBBFDBD" box="[159,286,582,607]" captionStart="FIGURES 23 – 30" captionStartId="13.[151,264,1737,1760]" captionTargetBox="[183,1417,218,1706]" captionTargetId="figure@13.[150,1436,193,1716]" captionTargetPageId="13" captionText="FIGURES 23 – 30. Akamboja cleidae sp. nov. 23 habitus, dorsal. 24 habitus, lateral. 25 head and pronotum, lateral. 26 head, dorsal. Akamboja monteirorum sp. nov., 27 habitus, dorsal. 28 habitus, lateral. 29 head and pronotum, lateral. 30 head, dorsal." httpUri="https://zenodo.org/record/845137/files/figure.png" pageId="14" pageNumber="515">Figs 27–30</figureCitation>
and
<figureCitation id="A6852A61DE2A7D030860D61FFED4FDBD" box="[340,369,583,607]" captionStart="FIGURES 41 – 43" captionStartId="18.[151,264,1871,1894]" captionTargetBox="[408,1173,193,1820]" captionTargetId="figure@18.[408,1183,193,1843]" captionTargetPageId="18" captionText="FIGURES 41 – 43. Map of Akamboja gen. nov. species distribution. Red dots mark the regions of occurrence of the species. Blue highlight marks the Serra do Mar mountain range high altitudinal areas. Yellow highlight marks Serra da Mantiqueira mountain range high altitudinal areas. Green highlight marks Tijuca Massif high altitudinal areas. The regions limits are approximated based on maps of project radambrasil (1983). 41 Akamboja minimum sp. nov. distribution. 42 Akamboja cleidae sp. nov. distribution. 43 Akamboja monteirorum sp. nov. distribution." httpUri="https://zenodo.org/record/845141/files/figure.png" pageId="14" pageNumber="515">43</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="76A4656FDE2A7D0309A3D6D6FD50FD0D" pageId="14" pageNumber="515" type="etymology">
<paragraph id="3E0136E4DE2A7D0309A3D6D6FD50FD0D" blockId="14.[151,1437,654,1939]" pageId="14" pageNumber="515">
<emphasis id="0CCAEAF6DE2A7D0309A3D6D6FEBBFD45" bold="true" box="[151,286,654,679]" pageId="14" pageNumber="515">Etymology.</emphasis>
The name is in honor to the Monteiro family, especially our dear friends, Drs. Ricardo and Margarete. Their enthusiasm and contributions to Neotropical insect ecology are noteworthy. They also helped the last author to collect specimens of three species. Genitive, plural.
</paragraph>
</subSubSection>
<subSubSection id="76A4656FDE2A7D0309F3D6A2FDD1FC9D" pageId="14" pageNumber="515" type="diagnosis">
<paragraph id="3E0136E4DE2A7D0309F3D6A2FE29FCF1" blockId="14.[151,1437,654,1939]" box="[199,396,762,787]" pageId="14" pageNumber="515">
<emphasis id="0CCAEAF6DE2A7D0309F3D6A2FE29FCF1" bold="true" box="[199,396,762,787]" pageId="14" pageNumber="515">Diagnosis. Male.</emphasis>
</paragraph>
<paragraph id="3E0136E4DE2A7D0309F3D747FDD1FC9D" blockId="14.[151,1437,654,1939]" pageId="14" pageNumber="515">
Eyes occupying two thirds of head width, in lateral view (
<figureCitation id="A6852A61DE2A7D030A52D746FC18FCD5" box="[870,957,798,823]" captionStart="FIGURES 23 – 30" captionStartId="13.[151,264,1737,1760]" captionTargetBox="[183,1417,218,1706]" captionTargetId="figure@13.[150,1436,193,1716]" captionTargetPageId="13" captionText="FIGURES 23 – 30. Akamboja cleidae sp. nov. 23 habitus, dorsal. 24 habitus, lateral. 25 head and pronotum, lateral. 26 head, dorsal. Akamboja monteirorum sp. nov., 27 habitus, dorsal. 28 habitus, lateral. 29 head and pronotum, lateral. 30 head, dorsal." httpUri="https://zenodo.org/record/845137/files/figure.png" pageId="14" pageNumber="515">Fig. 29</figureCitation>
). Apical maxillary palpomere 3.5x longer than subapical (
<figureCitation id="A6852A61DE2A7D030872D71AFE3FFCB9" box="[326,410,834,859]" captionStart="FIGURES 23 – 30" captionStartId="13.[151,264,1737,1760]" captionTargetBox="[183,1417,218,1706]" captionTargetId="figure@13.[150,1436,193,1716]" captionTargetPageId="13" captionText="FIGURES 23 – 30. Akamboja cleidae sp. nov. 23 habitus, dorsal. 24 habitus, lateral. 25 head and pronotum, lateral. 26 head, dorsal. Akamboja monteirorum sp. nov., 27 habitus, dorsal. 28 habitus, lateral. 29 head and pronotum, lateral. 30 head, dorsal." httpUri="https://zenodo.org/record/845137/files/figure.png" pageId="14" pageNumber="515">Fig. 29</figureCitation>
). Vertex occupying? of head in dorsal view (
<figureCitation id="A6852A61DE2A7D030AA6D71AFC43FCB9" box="[914,998,834,859]" captionStart="FIGURES 23 – 30" captionStartId="13.[151,264,1737,1760]" captionTargetBox="[183,1417,218,1706]" captionTargetId="figure@13.[150,1436,193,1716]" captionTargetPageId="13" captionText="FIGURES 23 – 30. Akamboja cleidae sp. nov. 23 habitus, dorsal. 24 habitus, lateral. 25 head and pronotum, lateral. 26 head, dorsal. Akamboja monteirorum sp. nov., 27 habitus, dorsal. 28 habitus, lateral. 29 head and pronotum, lateral. 30 head, dorsal." httpUri="https://zenodo.org/record/845137/files/figure.png" pageId="14" pageNumber="515">Fig. 30</figureCitation>
). Elytron reaching the posterior margin of the fourth abdominal segment (
<figureCitation id="A6852A61DE2A7D030B20D73EFDCDFC9D" box="[532,616,870,895]" captionStart="FIGURES 23 – 30" captionStartId="13.[151,264,1737,1760]" captionTargetBox="[183,1417,218,1706]" captionTargetId="figure@13.[150,1436,193,1716]" captionTargetPageId="13" captionText="FIGURES 23 – 30. Akamboja cleidae sp. nov. 23 habitus, dorsal. 24 habitus, lateral. 25 head and pronotum, lateral. 26 head, dorsal. Akamboja monteirorum sp. nov., 27 habitus, dorsal. 28 habitus, lateral. 29 head and pronotum, lateral. 30 head, dorsal." httpUri="https://zenodo.org/record/845137/files/figure.png" pageId="14" pageNumber="515">Fig. 27</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="76A4656FDE2A7D0309F3D7D2FDF4FACD" pageId="14" pageNumber="515" type="description">
<paragraph id="3E0136E4DE2A7D0309F3D7D2FE05FC41" blockId="14.[151,1437,654,1939]" box="[199,416,906,931]" pageId="14" pageNumber="515">
<emphasis id="0CCAEAF6DE2A7D0309F3D7D2FE05FC41" bold="true" box="[199,416,906,931]" pageId="14" pageNumber="515">Description, male.</emphasis>
</paragraph>
<paragraph id="3E0136E4DE2A7D0309F3D7F6FC07FBD1" blockId="14.[151,1437,654,1939]" pageId="14" pageNumber="515">
<emphasis id="0CCAEAF6DE2A7D0309F3D7F6FED3FC25" bold="true" box="[199,374,942,967]" pageId="14" pageNumber="515">Measurements</emphasis>
(n=5): Total length:
<quantity id="F9469B01DE2A7D030B5FD7F6FD50FC24" box="[619,757,942,967]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" metricValueMax="4.5" metricValueMin="4.1" pageId="14" pageNumber="515" unit="mm" value="4.3" valueMax="4.5" valueMin="4.1">4.1–4.5 mm</quantity>
(aver.
<quantity id="F9469B01DE2A7D030A73D7F7FC01FC25" box="[839,932,943,967]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="14" pageNumber="515" unit="mm" value="4.3">4.3 mm</quantity>
). Head length:
<quantity id="F9469B01DE2A7D030D63D7F6FB44FC24" box="[1111,1249,942,967]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="14" pageNumber="515" unit="mm" value="0.6" valueMax="0.7" valueMin="0.5">0.5–0.7 mm</quantity>
(aver.
<quantity id="F9469B01DE2A7D030C07D7F6FA35FC25" box="[1331,1424,942,967]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="515" unit="mm" value="0.6">0.6 mm</quantity>
). Head width:
<quantity id="F9469B01DE2A7D03081AD78AFE1CFC08" box="[302,441,978,1003]" metricMagnitude="-4" metricUnit="m" metricValue="6.499999999999999" metricValueMax="7.0" metricValueMin="6.0" pageId="14" pageNumber="515" unit="mm" value="0.6499999999999999" valueMax="0.7" valueMin="0.6">0.6–0.7 mm</quantity>
(aver.
<quantity id="F9469B01DE2A7D030B39D78AFDCFFC09" box="[525,618,978,1003]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="515" unit="mm" value="0.6">0.6 mm</quantity>
). Pronotum length:
<quantity id="F9469B01DE2A7D030A66D78AFC79FC08" box="[850,988,978,1003]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="14" pageNumber="515" unit="mm" value="0.55" valueMax="0.6" valueMin="0.5">0.5–0.6 mm</quantity>
(aver.
<quantity id="F9469B01DE2A7D030D04D78AFB39FC09" box="[1072,1180,978,1003]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="14" pageNumber="515" unit="mm" value="0.55">0.55 mm</quantity>
). Pronotum maximum length:
<quantity id="F9469B01DE2A7D0309D9D7AEFED3FBEC" box="[237,374,1014,1039]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="14" pageNumber="515" unit="mm" value="0.55" valueMax="0.6" valueMin="0.5">0.5–0.6 mm</quantity>
(aver.
<quantity id="F9469B01DE2A7D0308F3D7AEFD94FBED" box="[455,561,1014,1039]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="14" pageNumber="515" unit="mm" value="0.55">0.55 mm</quantity>
). Elytron length:
<quantity id="F9469B01DE2A7D030BCDD7AEFC27FBEC" box="[761,898,1014,1039]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="14" pageNumber="515" unit="mm" value="1.55" valueMax="1.6" valueMin="1.5">1.5–1.6 mm</quantity>
(aver.
<quantity id="F9469B01DE2A7D030AE6D7AEFB99FBED" box="[978,1084,1014,1039]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" pageId="14" pageNumber="515" unit="mm" value="1.55">1.55 mm</quantity>
). Elytron maximum width:
<quantity id="F9469B01DE2A7D030C4DD7AFFF66FBD0" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="515" unit="mm" value="0.4">0.4 mm</quantity>
(aver.
<quantity id="F9469B01DE2A7D03083BD043FECCFBD1" box="[271,361,1051,1075]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="515" unit="mm" value="0.4">0.4 mm</quantity>
). Elytral spot length:
<quantity id="F9469B01DE2A7D030B6ED043FD44FBD0" box="[602,737,1051,1075]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="14" pageNumber="515" unit="mm" value="0.25" valueMax="0.3" valueMin="0.2">0.2–0.3 mm</quantity>
(aver.
<quantity id="F9469B01DE2A7D030A1AD042FC33FBD1" box="[814,918,1050,1075]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="14" pageNumber="515" unit="mm" value="0.25">0.25 mm</quantity>
).
</paragraph>
<paragraph id="3E0136E4DE2A7D0309F3D066FC9FFB21" blockId="14.[151,1437,654,1939]" pageId="14" pageNumber="515">
<emphasis id="0CCAEAF6DE2A7D0309F3D066FEC1FBB5" bold="true" box="[199,356,1086,1111]" pageId="14" pageNumber="515">Morphology:</emphasis>
Eyes occupying two thirds of head width, in lateral view (
<figureCitation id="A6852A61DE2A7D030AC9D066FBF7FBB5" box="[1021,1106,1086,1111]" captionStart="FIGURES 23 – 30" captionStartId="13.[151,264,1737,1760]" captionTargetBox="[183,1417,218,1706]" captionTargetId="figure@13.[150,1436,193,1716]" captionTargetPageId="13" captionText="FIGURES 23 – 30. Akamboja cleidae sp. nov. 23 habitus, dorsal. 24 habitus, lateral. 25 head and pronotum, lateral. 26 head, dorsal. Akamboja monteirorum sp. nov., 27 habitus, dorsal. 28 habitus, lateral. 29 head and pronotum, lateral. 30 head, dorsal." httpUri="https://zenodo.org/record/845137/files/figure.png" pageId="14" pageNumber="515">Fig. 29</figureCitation>
). Vertex occupying? of head in dorsal view (
<figureCitation id="A6852A61DE2A7D03087ED03AFE04FB99" box="[330,417,1122,1147]" captionStart="FIGURES 23 – 30" captionStartId="13.[151,264,1737,1760]" captionTargetBox="[183,1417,218,1706]" captionTargetId="figure@13.[150,1436,193,1716]" captionTargetPageId="13" captionText="FIGURES 23 – 30. Akamboja cleidae sp. nov. 23 habitus, dorsal. 24 habitus, lateral. 25 head and pronotum, lateral. 26 head, dorsal. Akamboja monteirorum sp. nov., 27 habitus, dorsal. 28 habitus, lateral. 29 head and pronotum, lateral. 30 head, dorsal." httpUri="https://zenodo.org/record/845137/files/figure.png" pageId="14" pageNumber="515">Fig. 30</figureCitation>
). Apical maxillary palpomere 3.5x longer than subapical (
<figureCitation id="A6852A61DE2A7D030D0FD03AFB37FB99" box="[1083,1170,1122,1147]" captionStart="FIGURES 23 – 30" captionStartId="13.[151,264,1737,1760]" captionTargetBox="[183,1417,218,1706]" captionTargetId="figure@13.[150,1436,193,1716]" captionTargetPageId="13" captionText="FIGURES 23 – 30. Akamboja cleidae sp. nov. 23 habitus, dorsal. 24 habitus, lateral. 25 head and pronotum, lateral. 26 head, dorsal. Akamboja monteirorum sp. nov., 27 habitus, dorsal. 28 habitus, lateral. 29 head and pronotum, lateral. 30 head, dorsal." httpUri="https://zenodo.org/record/845137/files/figure.png" pageId="14" pageNumber="515">Fig. 30</figureCitation>
). Head as wide as long (
<figureCitation id="A6852A61DE2A7D0309ABD0DEFF51FB7D" box="[159,244,1158,1183]" captionStart="FIGURES 23 – 30" captionStartId="13.[151,264,1737,1760]" captionTargetBox="[183,1417,218,1706]" captionTargetId="figure@13.[150,1436,193,1716]" captionTargetPageId="13" captionText="FIGURES 23 – 30. Akamboja cleidae sp. nov. 23 habitus, dorsal. 24 habitus, lateral. 25 head and pronotum, lateral. 26 head, dorsal. Akamboja monteirorum sp. nov., 27 habitus, dorsal. 28 habitus, lateral. 29 head and pronotum, lateral. 30 head, dorsal." httpUri="https://zenodo.org/record/845137/files/figure.png" pageId="14" pageNumber="515">Fig. 30</figureCitation>
). Elytron 3.5x longer than wide (
<figureCitation id="A6852A61DE2A7D030B5DD0DEFD18FB7D" box="[617,701,1158,1183]" captionStart="FIGURES 23 – 30" captionStartId="13.[151,264,1737,1760]" captionTargetBox="[183,1417,218,1706]" captionTargetId="figure@13.[150,1436,193,1716]" captionTargetPageId="13" captionText="FIGURES 23 – 30. Akamboja cleidae sp. nov. 23 habitus, dorsal. 24 habitus, lateral. 25 head and pronotum, lateral. 26 head, dorsal. Akamboja monteirorum sp. nov., 27 habitus, dorsal. 28 habitus, lateral. 29 head and pronotum, lateral. 30 head, dorsal." httpUri="https://zenodo.org/record/845137/files/figure.png" pageId="14" pageNumber="515">Fig. 27</figureCitation>
), reaching the posterior margin of the fourth abdominal segment, elytron apical spot occupying a fifth of the elytron (
<figureCitation id="A6852A61DE2A7D030BEDD0F2FC8BFB21" box="[729,814,1194,1219]" captionStart="FIGURES 23 – 30" captionStartId="13.[151,264,1737,1760]" captionTargetBox="[183,1417,218,1706]" captionTargetId="figure@13.[150,1436,193,1716]" captionTargetPageId="13" captionText="FIGURES 23 – 30. Akamboja cleidae sp. nov. 23 habitus, dorsal. 24 habitus, lateral. 25 head and pronotum, lateral. 26 head, dorsal. Akamboja monteirorum sp. nov., 27 habitus, dorsal. 28 habitus, lateral. 29 head and pronotum, lateral. 30 head, dorsal." httpUri="https://zenodo.org/record/845137/files/figure.png" pageId="14" pageNumber="515">Fig. 27</figureCitation>
).
</paragraph>
<paragraph id="3E0136E4DE2A7D0309F3D096FC36FAE9" blockId="14.[151,1437,654,1939]" pageId="14" pageNumber="515">
<emphasis id="0CCAEAF6DE2A7D0309F3D096FEB5FB05" bold="true" box="[199,272,1230,1255]" pageId="14" pageNumber="515">Color.</emphasis>
Body overall light brown. Pronotum dark to light brown. Elytron almost black to light brown, apex pale yellow, sometimes light brown. Wing venation black to light brown.
</paragraph>
<paragraph id="3E0136E4DE2A7D0309F3D14EFDF4FACD" blockId="14.[151,1437,654,1939]" box="[199,593,1302,1327]" pageId="14" pageNumber="515">
<emphasis id="0CCAEAF6DE2A7D0309F3D14EFE79FACD" bold="true" box="[199,476,1302,1327]" pageId="14" pageNumber="515">Immature and females.</emphasis>
Unknown
</paragraph>
</subSubSection>
<subSubSection id="76A4656FDE2A7D0309F3D162FD6EFA79" pageId="14" pageNumber="515" type="biology_ecology">
<paragraph id="3E0136E4DE2A7D0309F3D162FD6EFA79" blockId="14.[151,1437,654,1939]" pageId="14" pageNumber="515">
<emphasis id="0CCAEAF6DE2A7D0309F3D162FE57FAB1" bold="true" box="[199,498,1338,1363]" pageId="14" pageNumber="515">Biology and distribution.</emphasis>
<taxonomicName id="F9BE4D67DE2A7D0308CDD162FCA1FAB1" authority="Roza &amp; Quintino &amp; Mermudes &amp; Silveira, 2017" authorityName="Roza &amp; Quintino &amp; Mermudes &amp; Silveira" authorityYear="2017" box="[505,772,1338,1363]" class="Insecta" family="Phengodidae" genus="Akamboja" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="species" species="monteirorum" status="sp. nov.">
<emphasis id="0CCAEAF6DE2A7D0308CDD162FCA1FAB1" box="[505,772,1338,1363]" italics="true" pageId="14" pageNumber="515">Akamboja monteirorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="17F9578DDE2A7D030A3FD162FCC0FAB1" box="[779,869,1338,1363]" pageId="14" pageNumber="515" rank="species">
<emphasis id="0CCAEAF6DE2A7D030A3FD162FCC0FAB1" bold="true" box="[779,869,1338,1363]" pageId="14" pageNumber="515">sp. nov.</emphasis>
</taxonomicNameLabel>
seems to occur only during spring and summer. It inhabits high altitudinal areas of Itatiaia, on the Itatiaia massif, in the Serra da Mantiqueira mountain range, from
<quantity id="F9469B01DE2A7D0309A3D1DBFEEEFA78" box="[151,331,1411,1435]" metricMagnitude="3" metricUnit="m" metricValue="1.361" metricValueMax="1.4419999999999997" metricValueMin="1.28" pageId="14" pageNumber="515" unit="m" value="1361.0" valueMax="1442.0" valueMin="1280.0">
<date id="4A001024DE2A7D0309A3D1DBFE8EFA79" box="[151,299,1411,1435]" pageId="14" pageNumber="515" value="1280" valueMax="1442">1280 to 1442</date>
m
</quantity>
. It is found on the montane forest.
</paragraph>
</subSubSection>
<subSubSection id="76A4656FDE2A7D0309F3D1FEFF4BF975" pageId="14" pageNumber="515" type="discussion">
<paragraph id="3E0136E4DE2A7D0309F3D1FEFF4BF975" blockId="14.[151,1437,654,1939]" pageId="14" pageNumber="515">
<emphasis id="0CCAEAF6DE2A7D0309F3D1FEFE9EFA5D" bold="true" box="[199,315,1446,1471]" pageId="14" pageNumber="515">Remarks.</emphasis>
This species resembles
<taxonomicName id="F9BE4D67DE2A7D030B60D1FEFC92FA5C" authority="Roza &amp; Quintino &amp; Mermudes &amp; Silveira, 2017" authorityName="Roza &amp; Quintino &amp; Mermudes &amp; Silveira" authorityYear="2017" box="[596,823,1446,1471]" class="Insecta" family="Phengodidae" genus="Akamboja" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="species" species="minimum" status="sp. nov.">
<emphasis id="0CCAEAF6DE2A7D030B60D1FEFC92FA5C" box="[596,823,1446,1471]" italics="true" pageId="14" pageNumber="515">Akamboja minimum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="17F9578DDE2A7D030A75D1FEFC38FA5D" box="[833,925,1446,1471]" pageId="14" pageNumber="515" rank="species">
<emphasis id="0CCAEAF6DE2A7D030A75D1FEFC38FA5D" bold="true" box="[833,925,1446,1471]" pageId="14" pageNumber="515">sp. nov.</emphasis>
</taxonomicNameLabel>
, but can be distinguished by its bigger size, elytron (the elytron of
<taxonomicName id="F9BE4D67DE2A7D0308A3D192FDDDFA01" authority="Roza &amp; Quintino &amp; Mermudes &amp; Silveira, 2017" authorityName="Roza &amp; Quintino &amp; Mermudes &amp; Silveira" authorityYear="2017" box="[407,632,1482,1507]" class="Insecta" family="Phengodidae" genus="Akamboja" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="species" species="minimum" status="sp. nov.">
<emphasis id="0CCAEAF6DE2A7D0308A3D192FDDDFA01" box="[407,632,1482,1507]" italics="true" pageId="14" pageNumber="515">Akamboja minimum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="17F9578DDE2A7D030BB4D192FD7FFA01" box="[640,730,1482,1507]" pageId="14" pageNumber="515" rank="species">
<emphasis id="0CCAEAF6DE2A7D030BB4D192FD7FFA01" bold="true" box="[640,730,1482,1507]" pageId="14" pageNumber="515">sp. nov.</emphasis>
</taxonomicNameLabel>
reaches the anterior margin of the second abdominal segment, while in
<taxonomicName id="F9BE4D67DE2A7D0309CAD1B6FDAEF9E4" authority="Roza &amp; Quintino &amp; Mermudes &amp; Silveira, 2017" authorityName="Roza &amp; Quintino &amp; Mermudes &amp; Silveira" authorityYear="2017" box="[254,523,1518,1543]" class="Insecta" family="Phengodidae" genus="Akamboja" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="species" species="monteirorum" status="sp. nov.">
<emphasis id="0CCAEAF6DE2A7D0309CAD1B6FDAEF9E4" box="[254,523,1518,1543]" italics="true" pageId="14" pageNumber="515">Akamboja monteirorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="17F9578DDE2A7D030B20D1B6FDD4F9E5" box="[532,625,1518,1543]" pageId="14" pageNumber="515" rank="species">
<emphasis id="0CCAEAF6DE2A7D030B20D1B6FDD4F9E5" bold="true" box="[532,625,1518,1543]" pageId="14" pageNumber="515">sp. nov.</emphasis>
</taxonomicNameLabel>
it reaches the posterior margin of the third), smaller vertex (bigger in
<taxonomicName id="F9BE4D67DE2A7D0309A3D24AFEDCF9C9" authority="Roza &amp; Quintino &amp; Mermudes &amp; Silveira, 2017" authorityName="Roza &amp; Quintino &amp; Mermudes &amp; Silveira" authorityYear="2017" box="[151,377,1554,1579]" class="Insecta" family="Phengodidae" genus="Akamboja" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="species" species="minimum" status="sp. nov.">
<emphasis id="0CCAEAF6DE2A7D0309A3D24AFEDCF9C9" box="[151,377,1554,1579]" italics="true" pageId="14" pageNumber="515">Akamboja minimum</emphasis>
</taxonomicName>
<emphasis id="0CCAEAF6DE2A7D0308B5D24AFE43F9C9" bold="true" box="[385,486,1554,1579]" pageId="14" pageNumber="515">
<taxonomicNameLabel id="17F9578DDE2A7D0308B5D24AFE7BF9C9" box="[385,478,1554,1579]" pageId="14" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
)
</emphasis>
and its high altitude occurrence (low altitude occurrence in
<taxonomicName id="F9BE4D67DE2A7D030DA2D24AFAD7F9C9" authority="Roza &amp; Quintino &amp; Mermudes &amp; Silveira, 2017" authorityName="Roza &amp; Quintino &amp; Mermudes &amp; Silveira" authorityYear="2017" box="[1174,1394,1554,1579]" class="Insecta" family="Phengodidae" genus="Akamboja" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="species" species="minimun" status="sp. nov.">
<emphasis id="0CCAEAF6DE2A7D030DA2D24AFAD7F9C9" box="[1174,1394,1554,1579]" italics="true" pageId="14" pageNumber="515">Akamboja minimun</emphasis>
</taxonomicName>
<emphasis id="0CCAEAF6DE2A7D030C4ED24AFF74F9AD" bold="true" pageId="14" pageNumber="515">
<taxonomicNameLabel id="17F9578DDE2A7D030C4ED24AFF6CF9AD" pageId="14" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
)
</emphasis>
.
<taxonomicName id="F9BE4D67DE2A7D0309D5D26EFE49F9AC" authority="Roza &amp; Quintino &amp; Mermudes &amp; Silveira, 2017" authorityName="Roza &amp; Quintino &amp; Mermudes &amp; Silveira" authorityYear="2017" box="[225,492,1590,1615]" class="Insecta" family="Phengodidae" genus="Akamboja" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="species" species="monteirorum" status="sp. nov.">
<emphasis id="0CCAEAF6DE2A7D0309D5D26EFE49F9AC" box="[225,492,1590,1615]" italics="true" pageId="14" pageNumber="515">Akamboja monteirorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="17F9578DDE2A7D0308C0D26EFDF5F9AD" box="[500,592,1590,1615]" pageId="14" pageNumber="515" rank="species">
<emphasis id="0CCAEAF6DE2A7D0308C0D26EFDF5F9AD" bold="true" box="[500,592,1590,1615]" pageId="14" pageNumber="515">sp. nov.</emphasis>
</taxonomicNameLabel>
occurs only in montane forest (from
<quantity id="F9469B01DE2A7D030D31D26FFBEBF9AC" box="[1029,1102,1591,1615]" metricMagnitude="2" metricUnit="m" metricValue="8.3" pageId="14" pageNumber="515" unit="m" value="830.0">830 m</quantity>
to
<quantity id="F9469B01DE2A7D030D40D26FFB6BF9AD" box="[1140,1230,1591,1615]" metricMagnitude="3" metricUnit="m" metricValue="1.4419999999999997" pageId="14" pageNumber="515" unit="m" value="1442.0">1442 m</quantity>
), while
<taxonomicName id="F9BE4D67DE2A7D030C1ED26EFF5AF991" authority="Roza &amp; Quintino &amp; Mermudes &amp; Silveira, 2017" authorityName="Roza &amp; Quintino &amp; Mermudes &amp; Silveira" authorityYear="2017" class="Insecta" family="Phengodidae" genus="Akamboja" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="species" species="minimum" status="sp. nov.">
<emphasis id="0CCAEAF6DE2A7D030C1ED26EFF5AF991" italics="true" pageId="14" pageNumber="515">Akamboja minimum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="17F9578DDE2A7D030832D202FEC4F991" box="[262,353,1626,1651]" pageId="14" pageNumber="515" rank="species">
<emphasis id="0CCAEAF6DE2A7D030832D202FEC4F991" bold="true" box="[262,353,1626,1651]" pageId="14" pageNumber="515">sp. nov.</emphasis>
</taxonomicNameLabel>
occurs in lower montane forest (below
<quantity id="F9469B01DE2A7D030A16D203FCCEF991" box="[802,875,1627,1651]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="14" pageNumber="515" unit="m" value="800.0">800 m</quantity>
) and partially in the montane forest (beginning in
<quantity id="F9469B01DE2A7D0309A3D226FF47F975" box="[151,226,1662,1687]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="515" unit="m" value="600.0">600 m</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="76A4656FDE2A7D0309F3D2FAFD9CF871" pageId="14" pageNumber="515" type="materials_examined">
<paragraph id="3E0136E4DE2A7D0309F3D2FAFD9CF871" blockId="14.[151,1437,654,1939]" pageId="14" pageNumber="515">
<materialsCitation id="8ED63CB9DE2A7D0309F3D2FAFF17F93C" ID-GBIF-Occurrence="1586208318" collectionCode="PENSA, RIO" country="Brazil" location="Rio de Janeiro. Itatiaia. Parque Nacional de Itatiaia." municipality="RIO Pt" pageId="14" pageNumber="515" specimenCount="2" stateProvince="Rio de Janeiro" typeStatus="holotype">
<emphasis id="0CCAEAF6DE2A7D0309F3D2FAFD18F959" bold="true" box="[199,701,1698,1723]" pageId="14" pageNumber="515">
<typeStatus id="E1058846DE2A7D0309F3D2FAFEA7F959" box="[199,258,1698,1723]" pageId="14" pageNumber="515">Type</typeStatus>
material.
<typeStatus id="E1058846DE2A7D03084FD2FAFE4DF959" box="[379,488,1698,1723]" pageId="14" pageNumber="515" type="holotype">Holotype</typeStatus>
(male):
<collectingCountry id="46A97674DE2A7D030B7FD2FAFD1DF959" box="[587,696,1698,1723]" name="Brazil" pageId="14" pageNumber="515">BRAZIL</collectingCountry>
.
</emphasis>
<location id="3B61603FDE2A7D030BF0D2FCFAA2F959" LSID="urn:lsid:plazi:treatment:B61787F2DE2A7D0309A3D67BFD9CF871:3B61603FDE2A7D030BF0D2FCFAA2F959" box="[708,1287,1698,1723]" country="Brazil" municipality="RIO Pt" name="Rio de Janeiro. Itatiaia. Parque Nacional de Itatiaia." pageId="14" pageNumber="515" stateProvince="Rio de Janeiro">
<collectingRegion id="FC7AF806DE2A7D030BF0D2FCFCC3F959" box="[708,870,1698,1723]" country="Brazil" name="Rio de Janeiro" pageId="14" pageNumber="515">
<emphasis id="0CCAEAF6DE2A7D030BF0D2FCFCC3F959" box="[708,870,1698,1723]" italics="true" pageId="14" pageNumber="515">Rio de Janeiro</emphasis>
</collectingRegion>
. Itatiaia. Parque Nacional de Itatiaia.
</location>
<collectionCode id="58AFAE21DE2A7D030C39D2FBFAC3F959" box="[1293,1382,1699,1723]" pageId="14" pageNumber="515">PENSA</collectionCode>
<collectingMunicipality id="DE65AC9EDE2A7D030C58D2FBFF17F93C" pageId="14" pageNumber="515">
<collectionCode id="58AFAE21DE2A7D030C58D2FBFA39F959" box="[1388,1436,1699,1723]" pageId="14" pageNumber="515">RIO</collectionCode>
Pt
</collectingMunicipality>
</materialsCitation>
.
<materialsCitation id="8ED63CB9DE2A7D030988D29FFCFFF8E1" ID-GBIF-Occurrence="1586208322" collectingDate="2013-10-12" collectionCode="DZRJ" collectorName="R. Monteiro" country="Brazil" elevation="1442" latitude="-22.4285" location="Rio de Janeiro. Itatiaia. Parque Nacional de Itatiaia" longLatPrecision="1" longitude="-44.628387" pageId="14" pageNumber="515" specimenCount="1" specimenCount-male="1" stateProvince="Rio de Janeiro" typeStatus="paratype">
3,
<geoCoordinate id="5B8A5023DE2A7D0309E3D29EFED8F93D" box="[215,381,1734,1759]" direction="south" orientation="latitude" pageId="14" pageNumber="515" precision="1" value="-22.4285">22°25’42.6’’ S</geoCoordinate>
,
<geoCoordinate id="5B8A5023DE2A7D0308BDD29FFD9CF93C" box="[393,569,1735,1759]" direction="west" orientation="longitude" pageId="14" pageNumber="515" precision="1" value="-44.628387">44°37’42,2’’ W</geoCoordinate>
,
<elevation id="B593D1D7DE2A7D030B70D29FFD3FF93C" box="[580,666,1735,1759]" metricMagnitude="3" metricUnit="m" metricValue="1.4419999999999997" pageId="14" pageNumber="515" unit="m" value="1442.0">
<quantity id="F9469B01DE2A7D030B70D29FFD3FF93C" box="[580,666,1735,1759]" metricMagnitude="3" metricUnit="m" metricValue="1.4419999999999997" pageId="14" pageNumber="515" unit="m" value="1442.0">1442 m</quantity>
</elevation>
,
<specimenCount id="28B8FD6DDE2A7D030B91D29FFD56F93D" box="[677,755,1734,1759]" pageId="14" pageNumber="515" type="male">1 male</specimenCount>
,
<collectingDate id="5A44E9CCDE2A7D030BCAD29FFCD2F93D" box="[766,887,1735,1759]" pageId="14" pageNumber="515" value="2013-10-12">
<date id="4A001024DE2A7D030BCAD29FFCD2F93D" box="[766,887,1735,1759]" pageId="14" pageNumber="515" value="2013-10-12">12.X.2013</date>
</collectingDate>
,
<collectorName id="934B5332DE2A7D030AB6D29FFBAFF93D" box="[898,1034,1734,1759]" pageId="14" pageNumber="515">R. Monteiro</collectorName>
col. (
<collectionCode id="58AFAE21DE2A7D030D7DD29FFB37F93D" box="[1097,1170,1735,1759]" pageId="14" pageNumber="515">DZRJ</collectionCode>
).
<emphasis id="0CCAEAF6DE2A7D030D90D29EFEACF8E1" bold="true" pageId="14" pageNumber="515">
<typeStatus id="E1058846DE2A7D030D90D29EFAB9F93D" box="[1188,1308,1734,1759]" pageId="14" pageNumber="515" type="paratype">Paratypes</typeStatus>
(all male):
<collectingCountry id="46A97674DE2A7D0309A3D2B2FEA0F8E1" box="[151,261,1770,1795]" name="Brazil" pageId="14" pageNumber="515">BRAZIL</collectingCountry>
.
</emphasis>
<location id="3B61603FDE2A7D030825D2B4FCFFF8E1" LSID="urn:lsid:plazi:treatment:B61787F2DE2A7D0309A3D67BFD9CF871:3B61603FDE2A7D030825D2B4FCFFF8E1" box="[273,858,1770,1795]" country="Brazil" latitude="-22.4285" longLatPrecision="1" longitude="-44.628387" name="Rio de Janeiro. Itatiaia. Parque Nacional de Itatiaia" pageId="14" pageNumber="515" stateProvince="Rio de Janeiro">
<collectingRegion id="FC7AF806DE2A7D030825D2B4FE12F8E1" box="[273,439,1770,1795]" country="Brazil" name="Rio de Janeiro" pageId="14" pageNumber="515">
<emphasis id="0CCAEAF6DE2A7D030825D2B4FE12F8E1" box="[273,439,1770,1795]" italics="true" pageId="14" pageNumber="515">Rio de Janeiro</emphasis>
</collectingRegion>
. Itatiaia. Parque Nacional de Itatiaia
</location>
</materialsCitation>
.
<materialsCitation id="8ED63CB9DE2A7D030A51D2B3FBBCF8E0" ID-GBIF-Occurrence="1586208311" box="[869,1049,1771,1795]" collectionCode="PENSA, RIO" country="Brazil" location="RIO Pt" pageId="14" pageNumber="515" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="paratype">
<collectionCode id="58AFAE21DE2A7D030A51D2B3FC1BF8E1" box="[869,958,1771,1795]" pageId="14" pageNumber="515">PENSA</collectionCode>
<location id="3B61603FDE2A7D030AF2D2B3FBBCF8E0" LSID="urn:lsid:plazi:treatment:B61787F2DE2A7D0309A3D67BFD9CF871:3B61603FDE2A7D030AF2D2B3FBBCF8E0" box="[966,1049,1771,1795]" country="Brazil" name="RIO Pt" pageId="14" pageNumber="515" stateProvince="Rio de Janeiro">
<collectionCode id="58AFAE21DE2A7D030AF2D2B3FC53F8E1" box="[966,1014,1771,1795]" pageId="14" pageNumber="515">RIO</collectionCode>
Pt
</location>
</materialsCitation>
.
<materialsCitation id="8ED63CB9DE2A7D030D10D2B3FC74F8C4" ID-GBIF-Occurrence="1586208314" collectingDate="2015-02-12" collectionCode="DZRJ, PENSA, RIO" collectorName="R. Monteiro" country="Brazil" elevation="1280" latitude="-22.433222" location="RIO Pt" longLatPrecision="1" longitude="-44.627693" pageId="14" pageNumber="515" specimenCount="1" specimenCount-male="1" stateProvince="Rio de Janeiro" typeStatus="paratype">
2,
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Setor Lago Azul, Mata,
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</paragraph>
</subSubSection>
</treatment>
</document>
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