treatments-xml/data/C1/6E/20/C16E201EF7AA5B7BB2E1D9D2C4050BD6.xml

<document ID-CLB-Dataset="22854" ID-DOI="http://dx.doi.org/10.3897/zookeys.929.49987" ID-GBIF-Dataset="97c79ed9-33fd-49b8-90c5-95a159839db2" ID-PMC="PMC7192954" ID-Pensoft-Pub="1313-2970-929-1" ID-Pensoft-UUID="EFB92B457B1D56A1996483C5EECFAAF0" ID-PubMed="32377146" ID-ZooBank="4FCA3A27–973C49F7–8264–6A34D279EC5C" ModsDocID="1313-2970-929-1" checkinTime="1587627147117" checkinUser="pensoft" docAuthor="Gittenberger, Edmund, Leda, Pema, Wangchuk, Jigme, Gyeltshen, Choki &amp; Bjoern Stelbrink," docDate="2020" docId="C16E201EF7AA5B7BB2E1D9D2C4050BD6" docLanguage="en" docName="ZooKeys 929: 1-17" docOrigin="ZooKeys 929" docSource="http://dx.doi.org/10.3897/zookeys.929.49987" docTitle="Erhaia jannei Gittenberger &amp; Stelbrink 2020, sp. nov." docType="treatment" docUuid="3CEAF442-E7C8-4699-B291-882EE902D7D7" docUuidSource="ZooBank" docVersion="6" id="EFB92B457B1D56A1996483C5EECFAAF0" lastPageNumber="4" masterDocId="EFB92B457B1D56A1996483C5EECFAAF0" masterDocTitle="The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya" masterLastPageNumber="17" masterPageNumber="1" pageNumber="4" updateTime="1732748779243" updateUser="ExternalLinkService">
<mods:mods id="E9A2C5C0BAAA983D427A4424706D8FF6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C522CDEA31BB2D7302DA94B9E2B713BB">
<mods:title id="DF1F28C20BAA8C70671897DFC301A842">The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya</mods:title>
</mods:titleInfo>
<mods:name id="6F7502657BB7D9F3691ECB254F1EF3D1" type="personal">
<mods:role id="55CEAC393289DE9A8F963481C583278C">
<mods:roleTerm id="BD55BA850A6BCB8DE2EE12F6936CEB61">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D2453354077A6B11328DEF12903BC918">Gittenberger, Edmund</mods:namePart>
<mods:affiliation id="FA1AB67CD8DBB2DC3411659A01305012">Naturalis Biodiversity Center, P. O. Box 9517, NL- 2300 RA Leiden, The Netherlands</mods:affiliation>
<mods:nameIdentifier id="A34AF5CDE629CF46AEB12A2CA2D62069" type="email">egittenberger@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:name id="ADB5EBAF5AADC7B854FE0AA77FD06969" type="personal">
<mods:role id="A149FDC68E18CDE027AE2F8DEEDC2CA3">
<mods:roleTerm id="6482A050A0C0229E3C3570CEAC45C40F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E8C11E601FAB3CAF8A3F599EA503BEF9">Leda, Pema</mods:namePart>
<mods:affiliation id="4EA0E22C0AE08675F361613FB44E26B0">National Biodiversity Centre, Serbithang, Thimphu, Bhutan</mods:affiliation>
</mods:name>
<mods:name id="58C1F802A6CC997E1092F1F5466DA137" type="personal">
<mods:role id="9FC78DAE17A8579B7139127C8DF5B568">
<mods:roleTerm id="DE61DF54E43FA2A5DFECE9C1BAC7A956">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D8A95A15237CBD4CF126DFA96C8510C8">Wangchuk, Jigme</mods:namePart>
<mods:affiliation id="D63FBBF4469090819890D1336C4A2730">Ugyen Wangchuck Institute for Conservation and Environmental Research, Bumthang, Bhutan</mods:affiliation>
<mods:nameIdentifier id="62D09FE027B12A63C1ACA94693DB7D6D" type="orcid">https://orcid.org/0000-0003-3503-4369</mods:nameIdentifier>
</mods:name>
<mods:name id="0D0733D443AACCB8F0EEB2D9AAA4593C" type="personal">
<mods:role id="2809CE69CAD2CA60C29E246AEB25612D">
<mods:roleTerm id="27FBB8578410444CB511B7E4D5C1D56B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0E45C5BA530F8FD40C0A9D61EFF8A2D4">Gyeltshen, Choki</mods:namePart>
<mods:affiliation id="7E774A1C90C0B191C357BA71117F1309">National Biodiversity Centre, Serbithang, Thimphu, Bhutan</mods:affiliation>
<mods:nameIdentifier id="811089934BCDE10F76F6F2D1EAD7B97E" type="orcid">https://orcid.org/0000-0001-6790-6278</mods:nameIdentifier>
</mods:name>
<mods:name id="AD8C087EF4AD7AE7DF642ABC833B82C2" type="personal">
<mods:role id="717C02A3EF57203000D3CE4186147877">
<mods:roleTerm id="6E89E1C602EC034F7466EC465A4AE1B1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="759AC2F5A5BA8B8542C45F21281AE4BC">Bjoern Stelbrink,</mods:namePart>
<mods:affiliation id="7999F91AB27B89DA9F900FF62455B484">Department of Animal Ecology &amp; Systematics, Justus Liebig University, Heinrich-Buff-Ring 26 - 32 IFZ, D- 35392 Giessen, Germany &amp; Zoological Institute, Department of Environmental Sciences, University of Basel, Vesalgasse 1, 4051 Basel, Switzerland</mods:affiliation>
<mods:nameIdentifier id="F072029929BC333975991D40B21EA10B" type="orcid">https://orcid.org/0000-0002-7471-4992</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="7D5C971140B1F4DD1BDD902B2E9A17BB">text</mods:typeOfResource>
<mods:relatedItem id="6B8049732502015EBB9C57B68429BD82" type="host">
<mods:titleInfo id="7487BA0B8B4B2609E82273BCF50B51F0">
<mods:title id="8AB67B4654501148A9149548E794F7DC">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="E2899B2E532AB44CC04EFCF8E056D53D">
<mods:date id="44053AF984F3D33A449E0B3540DAB93B">2020</mods:date>
<mods:detail id="C031BD25D8A6288CC304C98ED3D29CAF" type="volume">
<mods:number id="4361DA00FC4AB96BF834AF82A9DD5E87">929</mods:number>
</mods:detail>
<mods:extent id="2157598882901A32DC6D08734C4D12D2" unit="page">
<mods:start id="A452490A1A7C6B5B2C0CABB1D5819862">1</mods:start>
<mods:end id="DC9341FC80D2C88BCF4E2AB210C0EE4D">17</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="6A04BED84946BC7D843034F75DB3EFB2">
<mods:url id="BDE98A0C9BB70DFDF568B43BB5E2B196">http://dx.doi.org/10.3897/zookeys.929.49987</mods:url>
</mods:location>
<mods:classification id="5EDD0A4987F5E33AF9DB0ED9DCD2BBB9">journal article</mods:classification>
<mods:identifier id="0D838EADA0D3CA131016119975AABAB5" type="DOI">http://dx.doi.org/10.3897/zookeys.929.49987</mods:identifier>
<mods:identifier id="449AC4A32584AD3F499E3D45737E16C8" type="Pensoft-Pub">1313-2970-929-1</mods:identifier>
<mods:identifier id="7370307A1E921F55C23397AD1BA71020" type="ZooBank">4FCA3A27-973C49F7-8264-6A34D279EC5C</mods:identifier>
<mods:identifier id="7DE5D90179DBAE266F45AB1E9BC59E82" type="Pensoft-UUID">EFB92B457B1D56A1996483C5EECFAAF0</mods:identifier>
</mods:mods>
<treatment id="C16E201EF7AA5B7BB2E1D9D2C4050BD6" ID-GBIF-Taxon="163735476" LSID="urn:lsid:zoobank.org:act:3CEAF442-E7C8-4699-B291-882EE902D7D7" httpUri="http://treatment.plazi.org/id/C16E201EF7AA5B7BB2E1D9D2C4050BD6" lastPageNumber="4" pageId="3" pageNumber="4" scope_class="Gastropoda" scope_family="Amnicolidae" scope_superFamily="Rissooidea">
<subSubSection id="92604205515F8592BF8C87D221792EC6" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="EB6882E45A470A01AB56CB7D58C9507B" pageId="3" pageNumber="4">
<taxonomicName id="4282CC9C858450981700F1DD5AED58A5" ID-CoL="7R8DD" LSID="C16E201E-F7AA-5B7B-B2E1-D9D2C4050BD6" authority="Gittenberger &amp; Stelbrink" authorityName="Gittenberger &amp; Stelbrink" authorityYear="2020" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia jannei" order="Littorinimorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="jannei" status="sp. nov.">Erhaia jannei Gittenberger &amp; Stelbrink</taxonomicName>
<taxonomicNameLabel id="925E0FC804D4766620742CC0A529881F" pageId="3" pageNumber="4">sp. nov.</taxonomicNameLabel>
<figureCitation id="BE236ECE900B484C92B82916A6EB1400" captionStart="Figures 2–10" captionStartId="F2" captionText="Figures 2 - 10. Erhaia jannei sp. nov., holotype (2 H 2.25 mm) and paratype (3 H 2.02 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia pelkiae sp. nov., holotype (4 H 1.86 mm) and paratype (5 H 1.86 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia wangchuki Gittenberger, Sherub &amp; Stelbrink, 2017. Shells from the type locality (6 H 1.98 mm) (7 H 2.03 mm); NBCB 1072. Bhutan, district Wangdue Phodrang, Gangchhu, Gangzetem brooklet, 2883 m a. s. l. Photos by J. Goud 8 Erhaia banepaensis Nesemann &amp; S. Sharma, 2007, holotype (H 1.95 mm); NHMW 103319. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 9 Erhaia nainitalensis Davis &amp; Rao, 1997, holotype of E. chandeshwariensis Nesemann &amp; S. Sharma, 2007 (H 1.94 mm); NHMW 103315. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 10 Erhaia sugurensis Nesemann, Shah &amp; Tachamo, 2007, holotype (H 1.95 mm); NHMW 104172. Nepal, Central Zone, Lalitpur district, Godawari, upper reaches of Sugure Khola forest stream, 1700 m a. s. l. NHMW." httpUri="https://binary.pensoft.net/fig/402717" pageId="3" pageNumber="4">Figs 2</figureCitation>
<figureCitation id="1955F86189FF072792D894822B981E34" captionStart="Figures 2–10" captionStartId="F2" captionText="Figures 2 - 10. Erhaia jannei sp. nov., holotype (2 H 2.25 mm) and paratype (3 H 2.02 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia pelkiae sp. nov., holotype (4 H 1.86 mm) and paratype (5 H 1.86 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia wangchuki Gittenberger, Sherub &amp; Stelbrink, 2017. Shells from the type locality (6 H 1.98 mm) (7 H 2.03 mm); NBCB 1072. Bhutan, district Wangdue Phodrang, Gangchhu, Gangzetem brooklet, 2883 m a. s. l. Photos by J. Goud 8 Erhaia banepaensis Nesemann &amp; S. Sharma, 2007, holotype (H 1.95 mm); NHMW 103319. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 9 Erhaia nainitalensis Davis &amp; Rao, 1997, holotype of E. chandeshwariensis Nesemann &amp; S. Sharma, 2007 (H 1.94 mm); NHMW 103315. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 10 Erhaia sugurensis Nesemann, Shah &amp; Tachamo, 2007, holotype (H 1.95 mm); NHMW 104172. Nepal, Central Zone, Lalitpur district, Godawari, upper reaches of Sugure Khola forest stream, 1700 m a. s. l. NHMW." httpUri="https://binary.pensoft.net/fig/402717" pageId="3" pageNumber="4">, 3</figureCitation>
<figureCitation id="C51538985F08D683553A3F9E4BF15337" captionStart="Figure 11" captionStartId="F3" captionText="Figure 11. Type locality of both Erhaia jannei and E. pelkiae; Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photo by EG." httpUri="https://binary.pensoft.net/fig/402718" pageId="3" pageNumber="4">, 11</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="E73FC505CE21FB6CC2B8B45E384AB049" pageId="3" pageNumber="4" type="reference_group">
<paragraph id="A3E5C3D863DBA4173F3EEB3E5D803136" pageId="3" pageNumber="4">
<taxonomicName id="D3965954B383DEEB84380B0384A2BCC0" authorityName="Davis &amp; Kuo" authorityYear="1985" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia" order="Littorinimorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="genus">Erhaia</taxonomicName>
sp. 
<bibRefCitation id="339AB60D8767E199D1AD273CB937A101" DOI="https://doi.org/10.3897/zookeys.679.13326" author="Gittenberger, E" journalOrPublisher="ZooKeys" pageId="14" pageNumber="15" pagination="21 - 28" refId="B7" refString="Gittenberger, E, Sherub, S, Stelbrink, B, 2017a. Erhaia Davis &amp; Kuo (Gastropoda, Rissooidea, Amnicolidae) also in Bhutan. ZooKeys 679: 21 - 28, DOI: https://doi.org/10.3897/zookeys.679.13326" title="Erhaia Davis &amp; Kuo (Gastropoda, Rissooidea, Amnicolidae) also in Bhutan." url="https://doi.org/10.3897/zookeys.679.13326" volume="679" year="2017 a">Gittenberger et al. 2017a</bibRefCitation>
: 25, fig. 3; 2017c: 900, 903, fig. 8.
</paragraph>
</subSubSection>
<subSubSection id="C131B099BBA8FD6270EBC0E75766BFEB" pageId="3" pageNumber="4" type="materials_examined">
<paragraph id="BFE845D9F1C27468A497882C7747C64C" pageId="3" pageNumber="4">Material examined.</paragraph>
<paragraph id="943F1C3FB858BD94D9D91FB186FFEB07" pageId="3" pageNumber="4">
<emphasis id="46A33D2195B2DAAF507E3CC9EA1FE96B" bold="true" pageId="3" pageNumber="4">
<emphasis id="9FDE8F89A46CA52429BE124C0DB71964" italics="true" pageId="3" pageNumber="4">Holotype</emphasis>
.
</emphasis>
(Fig. 
<figureCitation id="772FD5E4F7A821347C959B9CC57DFE52" captionStart="Figures 2–10" captionStartId="F2" captionText="Figures 2 - 10. Erhaia jannei sp. nov., holotype (2 H 2.25 mm) and paratype (3 H 2.02 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia pelkiae sp. nov., holotype (4 H 1.86 mm) and paratype (5 H 1.86 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia wangchuki Gittenberger, Sherub &amp; Stelbrink, 2017. Shells from the type locality (6 H 1.98 mm) (7 H 2.03 mm); NBCB 1072. Bhutan, district Wangdue Phodrang, Gangchhu, Gangzetem brooklet, 2883 m a. s. l. Photos by J. Goud 8 Erhaia banepaensis Nesemann &amp; S. Sharma, 2007, holotype (H 1.95 mm); NHMW 103319. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 9 Erhaia nainitalensis Davis &amp; Rao, 1997, holotype of E. chandeshwariensis Nesemann &amp; S. Sharma, 2007 (H 1.94 mm); NHMW 103315. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 10 Erhaia sugurensis Nesemann, Shah &amp; Tachamo, 2007, holotype (H 1.95 mm); NHMW 104172. Nepal, Central Zone, Lalitpur district, Godawari, upper reaches of Sugure Khola forest stream, 1700 m a. s. l. NHMW." httpUri="https://binary.pensoft.net/fig/402717" pageId="3" pageNumber="4">2</figureCitation>
) Bhutan • District Thimphu, c. 5 km E of Chhuzom, W of Geneykha; in brooklet with a prayer wheel along the road; 2750 m a.s.l.; 
<geoCoordinate id="8539745D2DE98AD0D9D98CA0E8F70C00" degrees="27" direction="north" minutes="18" orientation="latitude" precision="15" seconds="43" value="27.311943">27°18'43&quot;N</geoCoordinate>
, 
<geoCoordinate id="6C9163347E114272EBDCD61C64554667" degrees="89" direction="east" minutes="36" orientation="longitude" precision="15" seconds="10" value="89.602776">89°36'10&quot;E</geoCoordinate>
; E. Gittenberger, Choki Gyeltshen &amp; Pema Leda leg. 25.X.2018; NBCB 1057. 
<emphasis id="547F70957E432BD91937ABC6D5DB709E" bold="true" pageId="3" pageNumber="4">
<emphasis id="B86410EAE82F97C321367774C239AEF1" italics="true" pageId="3" pageNumber="4">Paratypes</emphasis>
.
</emphasis>
(Fig. 
<figureCitation id="1AFAE2F9A2E0CAEFEE3C184A87B9C1F5" captionStart="Figures 2–10" captionStartId="F2" captionText="Figures 2 - 10. Erhaia jannei sp. nov., holotype (2 H 2.25 mm) and paratype (3 H 2.02 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia pelkiae sp. nov., holotype (4 H 1.86 mm) and paratype (5 H 1.86 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia wangchuki Gittenberger, Sherub &amp; Stelbrink, 2017. Shells from the type locality (6 H 1.98 mm) (7 H 2.03 mm); NBCB 1072. Bhutan, district Wangdue Phodrang, Gangchhu, Gangzetem brooklet, 2883 m a. s. l. Photos by J. Goud 8 Erhaia banepaensis Nesemann &amp; S. Sharma, 2007, holotype (H 1.95 mm); NHMW 103319. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 9 Erhaia nainitalensis Davis &amp; Rao, 1997, holotype of E. chandeshwariensis Nesemann &amp; S. Sharma, 2007 (H 1.94 mm); NHMW 103315. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 10 Erhaia sugurensis Nesemann, Shah &amp; Tachamo, 2007, holotype (H 1.95 mm); NHMW 104172. Nepal, Central Zone, Lalitpur district, Godawari, upper reaches of Sugure Khola forest stream, 1700 m a. s. l. NHMW." httpUri="https://binary.pensoft.net/fig/402717" pageId="3" pageNumber="4">3</figureCitation>
) 6 shells and 2 animals in ethanol 70%; same data as for holotype; NBCB 1058.
</paragraph>
</subSubSection>
<subSubSection id="BA85E4DFDD94E12DDD0CB0447E3D1ABC" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="A6F67E0BA6DE092B9F19C6970AD6C750" pageId="3" pageNumber="4">Diagnosis.</paragraph>
<paragraph id="BFDA0A3CCD9506D7B7E7CCC56D872DCD" pageId="3" pageNumber="4">Shell large for the genus (H&gt; 2mm), ovoid, with a relatively large aperture.</paragraph>
</subSubSection>
<subSubSection id="A8BD93E16297234141DB6959F65B0625" pageId="3" pageNumber="4" type="shell">
<paragraph id="50395E95332C66F7E04E423C6C32FDC7" pageId="3" pageNumber="4">Shell.</paragraph>
<paragraph id="3332869E030EDA22E5F6A89192FCF276" pageId="3" pageNumber="4">
Shell obliquely ovoid, with 
<normalizedToken id="038262AE311685D5C5A8A0108845C157" originalValue="3½">31/2</normalizedToken>
-4 convex, shouldered whorls that are separated by a deep suture; clearly higher than broad; yellowish brown with fine irregular growth lines and some blackish brown periostracal ridges, one of which runs from the apertural columellar border into the umbilicus. Peristome not reflected. Parietal, columellar and a short part of the adjoining basal apertural border thickened by a whitish callus. Most specimens with a continuous peristome and a narrow umbilical chink. Protoconch encrusted in all specimens; teleoconch without spiral sculpture.
</paragraph>
</subSubSection>
<subSubSection id="7AEBE6AF6466EBECC013F8950A3E3C03" pageId="3" pageNumber="4" type="description">
<paragraph id="7204A959CB4B10DFA7E2B7E930120EC2" pageId="3" pageNumber="4">Measurements.</paragraph>
<paragraph id="918467D1E17B1E55753E7D3D50089ADB" pageId="3" pageNumber="4">
(
<emphasis id="94767C182E74E72D2E8F7452289E2EE1" italics="true" pageId="3" pageNumber="4">N</emphasis>
= 9): H 2.2-2.4 mm, B 1.5-1.6 mm. Holotype 2.2 
<normalizedToken id="CA4EC39EE9B613D01394BEF021371365" originalValue="×">x</normalizedToken>
1.6 mm.
</paragraph>
</subSubSection>
<subSubSection id="95D63898DF1C24459B4AE1BEBD244E4D" pageId="3" pageNumber="4" type="ecology">
<paragraph id="39B90BC3D793C18EAFB561E77D4B5EBB" pageId="3" pageNumber="4">Ecology</paragraph>
<paragraph id="47F9325240F63EA021BABF2706BB490E" pageId="3" pageNumber="4">
(Fig. 
<figureCitation id="EBDB7F227412F025D7E8230F285E2DE7" captionStart="Figure 11" captionStartId="F3" captionText="Figure 11. Type locality of both Erhaia jannei and E. pelkiae; Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photo by EG." httpUri="https://binary.pensoft.net/fig/402718" pageId="3" pageNumber="4">11</figureCitation>
). The species is uncommon on the rocks in the shaded streamlet uphill of the prayer wheel, where it occurs with 
<taxonomicName id="765119DC55EBE521338E41E4909B6131" baseAuthorityName="O. F. Müller" baseAuthorityYear="1774" class="Gastropoda" family="Lymnaeidae" genus="Galba" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Galba truncatula" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="truncatula">
<emphasis id="0162EBDD7967959B2CFD172D5A5316F7" italics="true" pageId="3" pageNumber="4">Galba truncatula</emphasis>
</taxonomicName>
(
<normalizedToken id="1D26A3E34B3AF6E81B0976FFD518A422" originalValue="Müller">Mueller</normalizedToken>
, 1774), 
<taxonomicName id="3912386B03799180395347E244274B9C" class="Gastropoda" family="Physidae" genus="Physa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Physa" order="Architaenioglossa" pageId="3" pageNumber="4" phylum="Mollusca" rank="genus">
<emphasis id="54DD996B6BBADBE2221CA877C7630DC2" italics="true" pageId="3" pageNumber="4">Physa</emphasis>
</taxonomicName>
sp. (new for Bhutan), and 
<taxonomicName id="B5137BA96BD700090DAFAF68BA9790C3" authorityName="Gittenberger &amp; Gyeltshen" authorityYear="2020" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia pelkiae" order="Littorinimorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="pelkiae">
<emphasis id="1FEAD71B8D8430881B611231DAE03A67" italics="true" pageId="3" pageNumber="4">Erhaia pelkiae</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</subSubSection>
<subSubSection id="3F73B57C9BCA456ADD11ADDA49BBB0C5" pageId="3" pageNumber="4" type="dna data">
<paragraph id="C04883E73A55B7BFDA700A9757001E4A" pageId="3" pageNumber="4">DNA data</paragraph>
<paragraph id="8A6D7D68E5FCD19FB4B1BD1595D47D9D" pageId="3" pageNumber="4">
(Fig. 
<figureCitation id="09B220FC1932F3DA2EC8E96C4A39957A" captionStart="Figure 18" captionStartId="F9" captionText="Figure 18. Maximum likelihood tree reconstructed with RAxML BlackBox (Stamatakis et al. 2008; GTR + G substitution model for each partition and 100 bootstrap replicates) based on the 16 S rRNA and COI dataset of Liu et al. (2014) and Guan et al. (2008), with new data in red. Numbers on branches denote bootstrap values&gt; 50." httpUri="https://binary.pensoft.net/fig/402724" pageId="3" pageNumber="4">18</figureCitation>
). The three individuals shared the same haplotype for both 16S rRNA (GenBank acc. no. MT239078) and COI (GenBank acc. no. MT237716). The uncorrected genetic p-distances between this species and 
<taxonomicName id="FF3410D1BD2DAE301165C8F825AE27BF" lsidName="E. wangchuki" pageId="3" pageNumber="4" rank="species" species="wangchuki">
<emphasis id="DD51F7BAC2BE9D2A845278AAA9BB83C8" italics="true" pageId="3" pageNumber="4">E. wangchuki</emphasis>
</taxonomicName>
were 0.84% for 16S rRNA and 4.87% for COI. The distances were considerably higher when compared to 
<taxonomicName id="5AFBC5E1274355CC757CC66CEBD7FD88" authorityName="Davis &amp; Kuo" authorityYear="1985" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia" order="Littorinimorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="genus">
<emphasis id="22A6D0CA8E8EF9707ACB959B86D4C3CA" italics="true" pageId="3" pageNumber="4">Erhaia</emphasis>
</taxonomicName>
sp. from China, Guangxi, viz. 2.74% for 16S rRNA and 10.25% for COI (GenBank acc. nos. KC832722 and KC832701, respectively).
</paragraph>
</subSubSection>
<subSubSection id="D5A37B25D0AED6DF727A923434ACB3E4" pageId="3" pageNumber="4" type="notes">
<paragraph id="C5C86CB0D783375DE984BF2C3BC54E74" pageId="3" pageNumber="4">Notes.</paragraph>
<paragraph id="182803D3ACEFC3067420F958DE0053E7" pageId="3" pageNumber="4">This species was discovered in 2012, but since only a single shell was collected then, a description was considered premature.</paragraph>
</subSubSection>
<subSubSection id="BB8541DE64D4B5C557DBF70C92242822" pageId="3" pageNumber="4" type="etymology">
<paragraph id="9284A9601EAB381D484ABA2DE6902F3A" pageId="3" pageNumber="4">Etymology.</paragraph>
<paragraph id="2CF5D5AA444945120D3F0DB30F055A5E" pageId="3" pageNumber="4">
The epithet 
<taxonomicName id="24C20F7963564BD14A0735C9D9ED9045" authorityName="Gittenberger &amp; Stelbrink" authorityYear="2020" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="jannei" order="Littorinimorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="jannei">
<emphasis id="D6A9821345881017EAD7F12FB029E0E0" italics="true" pageId="3" pageNumber="4">jannei</emphasis>
</taxonomicName>
refers to Mr Janne Clewing, the son of the last author.
</paragraph>
<caption id="16C28934F9F0C9C1C1DD060C02F79318" httpUri="https://binary.pensoft.net/fig/402717" pageId="3" pageNumber="4" start="Figures 2–10" startId="F2">
<paragraph id="760F5A68B93B002DBFD74EAC0CAE7BB2" pageId="3" pageNumber="4">
<emphasis id="B443D08F8C69A0A06FCF1A16BD8F6D06" bold="true" pageId="3" pageNumber="4">Figures 2-10.</emphasis>
<taxonomicName id="E358F53D34AE7D4DB6946D0E1D64A1CC" authorityName="Gittenberger &amp; Stelbrink" authorityYear="2020" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia jannei" order="Littorinimorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="jannei">
<emphasis id="10625B398A969CF807266634B4ABA211" italics="true" pageId="3" pageNumber="4">Erhaia jannei</emphasis>
</taxonomicName>
sp. nov., holotype (
<emphasis id="10557643648C285B6C697F471F90801D" bold="true" pageId="3" pageNumber="4">2</emphasis>
H 2.25 mm) and paratype (
<emphasis id="2D28344C98383898F5876A9BDA418DE0" bold="true" pageId="3" pageNumber="4">3</emphasis>
H 2.02 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a.s.l. Photos by J. Goud 
<taxonomicName id="91FF361062F860B6624F49BCE7F0FA76" authorityName="Gittenberger &amp; Gyeltshen" authorityYear="2020" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia pelkiae" order="Littorinimorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="pelkiae">
<emphasis id="3F7FEFBCE396BE14D9DEC6DF6640B551" italics="true" pageId="3" pageNumber="4">Erhaia pelkiae</emphasis>
</taxonomicName>
sp. nov., holotype (
<emphasis id="7D4091FEFB1B9FB9363D686243B269B9" bold="true" pageId="3" pageNumber="4">4</emphasis>
H 1.86 mm) and paratype (
<emphasis id="F781F95DE58CC4EEC2C98FACCD1F28A3" bold="true" pageId="3" pageNumber="4">5</emphasis>
H 1.86 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a.s.l. Photos by J. Goud 
<taxonomicName id="B39CB30040F0E1D60D6A4B2E51A5ECB9" authorityName="Gittenberger, Sherub &amp; Stelbrink" authorityYear="2017" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia wangchuki" order="Littorinimorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="wangchuki">
<emphasis id="EA7E67EFC7B3861BFB9CFE922B730CDF" italics="true" pageId="3" pageNumber="4">Erhaia wangchuki</emphasis>
</taxonomicName>
Gittenberger, Sherub &amp; Stelbrink, 2017. Shells from the type locality (
<emphasis id="011D7799939651D17E9050AEA7AEFA92" bold="true" pageId="3" pageNumber="4">6</emphasis>
H 1.98 mm) (
<emphasis id="14F529D4C33240346C96559779E8C4C3" bold="true" pageId="3" pageNumber="4">7</emphasis>
H 2.03 mm); NBCB 1072. Bhutan, district Wangdue Phodrang, Gangchhu, Gangzetem brooklet, 2883 m a.s.l. Photos by J. Goud 
<emphasis id="5ED223DF751F4002189CA60ADA9BDF0F" bold="true" pageId="3" pageNumber="4">8</emphasis>
<taxonomicName id="EBE6CAA817409AC6D257473C0A2C027C" authorityName="Nesemann &amp; S. Sharma" authorityYear="2007" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia banepaensis" order="Littorinimorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="banepaensis">
<emphasis id="29C95269AF8731C515A01D0F7C56CB5E" italics="true" pageId="3" pageNumber="4">Erhaia banepaensis</emphasis>
</taxonomicName>
Nesemann &amp; S. Sharma, 2007, holotype (H 1.95 mm); NHMW 103319. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 
<emphasis id="3B3B984566890E4E168DE26449C39090" bold="true" pageId="3" pageNumber="4">9</emphasis>
<taxonomicName id="1C9B04A99B5FCE48C57093BE26E23404" authorityName="Davis &amp; Rao" authorityYear="1997" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia nainitalensis" order="Littorinimorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="nainitalensis">
<emphasis id="7F0AE3C4C0BE61B5EA796519506F06B7" italics="true" pageId="3" pageNumber="4">Erhaia nainitalensis</emphasis>
</taxonomicName>
Davis &amp; Rao, 1997, holotype of 
<taxonomicName id="2BCF611605B90E343993604D6EC60398" lsidName="E. chandeshwariensis" pageId="3" pageNumber="4" rank="species" species="chandeshwariensis">
<emphasis id="DBCBF0EFA31200746A121C70E3B4827B" italics="true" pageId="3" pageNumber="4">E. chandeshwariensis</emphasis>
</taxonomicName>
Nesemann &amp; S. Sharma, 2007 (H 1.94 mm); NHMW 103315. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 
<emphasis id="4FF18E7AE33F932EC0F38EC78CA30585" bold="true" pageId="3" pageNumber="4">10</emphasis>
<taxonomicName id="C8C531B12C256AEBF1FD8C72E177E26C" authorityName="Nesemann, Shah &amp; Tachamo" authorityYear="2007" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia sugurensis" order="Littorinimorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="sugurensis">
<emphasis id="B7AC7255D4D250490401579672320410" italics="true" pageId="3" pageNumber="4">Erhaia sugurensis</emphasis>
</taxonomicName>
Nesemann, Shah &amp; Tachamo, 2007, holotype (H 1.95 mm); NHMW 104172. Nepal, Central Zone, Lalitpur district, Godawari, upper reaches of Sugure Khola forest stream, 1700 m a.s.l. NHMW.
</paragraph>
</caption>
<caption id="7664B25F75F8B09BA986AD33C95A452F" httpUri="https://binary.pensoft.net/fig/402718" pageId="3" pageNumber="4" start="Figure 11" startId="F3">
<paragraph id="FF79842F6ABD570FAFC13942B2D8EC15" pageId="3" pageNumber="4">
<emphasis id="FE1531FEE33C24C9ADEAC1334AA095C0" bold="true" pageId="3" pageNumber="4">Figure 11.</emphasis>
Type locality of both 
<taxonomicName id="591BCD0749667622F5CE9CAB8F1634D5" authorityName="Gittenberger &amp; Stelbrink" authorityYear="2020" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia jannei" order="Littorinimorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="jannei">
<emphasis id="3412CC1F94504792F4D5AECF20269A37" italics="true" pageId="3" pageNumber="4">Erhaia jannei</emphasis>
</taxonomicName>
and 
<taxonomicName id="F4D5E0059DE45CB6C26F142AF7E7CAAA" lsidName="E. pelkiae" pageId="3" pageNumber="4" rank="species" species="pelkiae">
<emphasis id="BF90F4DEB4851EA3DCD4FB04A012665A" italics="true" pageId="3" pageNumber="4">E. pelkiae</emphasis>
</taxonomicName>
; Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a.s.l. Photo by EG.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>