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248 lines
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.929.49987" ID-GBIF-Dataset="97c79ed9-33fd-49b8-90c5-95a159839db2" ID-PMC="PMC7192954" ID-Pensoft-Pub="1313-2970-929-1" ID-Pensoft-UUID="EFB92B457B1D56A1996483C5EECFAAF0" ID-PubMed="32377146" ID-ZooBank="4FCA3A27–973C49F7–8264–6A34D279EC5C" ModsDocID="1313-2970-929-1" checkinTime="1587627147117" checkinUser="pensoft" docAuthor="Gittenberger, Edmund, Leda, Pema, Wangchuk, Jigme, Gyeltshen, Choki & Bjoern Stelbrink," docDate="2020" docId="C16E201EF7AA5B7BB2E1D9D2C4050BD6" docLanguage="en" docName="ZooKeys 929: 1-17" docOrigin="ZooKeys 929" docSource="http://dx.doi.org/10.3897/zookeys.929.49987" docTitle="Erhaia jannei Gittenberger & Stelbrink 2020, sp. nov." docType="treatment" docUuid="3CEAF442-E7C8-4699-B291-882EE902D7D7" docUuidSource="ZooBank" docVersion="4" id="EFB92B457B1D56A1996483C5EECFAAF0" lastPageNumber="4" masterDocId="EFB92B457B1D56A1996483C5EECFAAF0" masterDocTitle="The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya" masterLastPageNumber="17" masterPageNumber="1" pageNumber="4" updateTime="1668168753491" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Gittenberger, Edmund</mods:namePart>
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<mods:affiliation>Naturalis Biodiversity Center, P. O. Box 9517, NL- 2300 RA Leiden, The Netherlands</mods:affiliation>
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<mods:nameIdentifier type="email">egittenberger@yahoo.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Leda, Pema</mods:namePart>
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<mods:affiliation>National Biodiversity Centre, Serbithang, Thimphu, Bhutan</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wangchuk, Jigme</mods:namePart>
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<mods:affiliation>Ugyen Wangchuck Institute for Conservation and Environmental Research, Bumthang, Bhutan</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-3503-4369</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Gyeltshen, Choki</mods:namePart>
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<mods:affiliation>National Biodiversity Centre, Serbithang, Thimphu, Bhutan</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-6790-6278</mods:nameIdentifier>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bjoern Stelbrink,</mods:namePart>
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<mods:affiliation>Department of Animal Ecology & Systematics, Justus Liebig University, Heinrich-Buff-Ring 26 - 32 IFZ, D- 35392 Giessen, Germany & Zoological Institute, Department of Environmental Sciences, University of Basel, Vesalgasse 1, 4051 Basel, Switzerland</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-7471-4992</mods:nameIdentifier>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>929</mods:number>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>17</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.929.49987</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.929.49987</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-929-1</mods:identifier>
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<mods:identifier type="ZooBank">4FCA3A27-973C49F7-8264-6A34D279EC5C</mods:identifier>
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<treatment ID-GBIF-Taxon="163735476" LSID="urn:lsid:zoobank.org:act:3CEAF442-E7C8-4699-B291-882EE902D7D7" httpUri="http://treatment.plazi.org/id/C16E201EF7AA5B7BB2E1D9D2C4050BD6" lastPageNumber="4" pageId="3" pageNumber="4">
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<subSubSection pageId="3" pageNumber="4" type="nomenclature">
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<paragraph pageId="3" pageNumber="4">
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<taxonomicName LSID="C16E201E-F7AA-5B7B-B2E1-D9D2C4050BD6" authority="Gittenberger & Stelbrink" authorityName="Gittenberger & Stelbrink" authorityYear="2020" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia jannei" order="Littorinimorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="jannei" status="sp. nov.">Erhaia jannei Gittenberger & Stelbrink</taxonomicName>
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<taxonomicNameLabel pageId="3" pageNumber="4">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figures 2–10" captionStartId="F2" captionText="Figures 2 - 10. Erhaia jannei sp. nov., holotype (2 H 2.25 mm) and paratype (3 H 2.02 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia pelkiae sp. nov., holotype (4 H 1.86 mm) and paratype (5 H 1.86 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia wangchuki Gittenberger, Sherub & Stelbrink, 2017. Shells from the type locality (6 H 1.98 mm) (7 H 2.03 mm); NBCB 1072. Bhutan, district Wangdue Phodrang, Gangchhu, Gangzetem brooklet, 2883 m a. s. l. Photos by J. Goud 8 Erhaia banepaensis Nesemann & S. Sharma, 2007, holotype (H 1.95 mm); NHMW 103319. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 9 Erhaia nainitalensis Davis & Rao, 1997, holotype of E. chandeshwariensis Nesemann & S. Sharma, 2007 (H 1.94 mm); NHMW 103315. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 10 Erhaia sugurensis Nesemann, Shah & Tachamo, 2007, holotype (H 1.95 mm); NHMW 104172. Nepal, Central Zone, Lalitpur district, Godawari, upper reaches of Sugure Khola forest stream, 1700 m a. s. l. NHMW." httpUri="https://binary.pensoft.net/fig/402717" pageId="3" pageNumber="4">Figs 2</figureCitation>
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<figureCitation captionStart="Figures 2–10" captionStartId="F2" captionText="Figures 2 - 10. Erhaia jannei sp. nov., holotype (2 H 2.25 mm) and paratype (3 H 2.02 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia pelkiae sp. nov., holotype (4 H 1.86 mm) and paratype (5 H 1.86 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia wangchuki Gittenberger, Sherub & Stelbrink, 2017. Shells from the type locality (6 H 1.98 mm) (7 H 2.03 mm); NBCB 1072. Bhutan, district Wangdue Phodrang, Gangchhu, Gangzetem brooklet, 2883 m a. s. l. Photos by J. Goud 8 Erhaia banepaensis Nesemann & S. Sharma, 2007, holotype (H 1.95 mm); NHMW 103319. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 9 Erhaia nainitalensis Davis & Rao, 1997, holotype of E. chandeshwariensis Nesemann & S. Sharma, 2007 (H 1.94 mm); NHMW 103315. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 10 Erhaia sugurensis Nesemann, Shah & Tachamo, 2007, holotype (H 1.95 mm); NHMW 104172. Nepal, Central Zone, Lalitpur district, Godawari, upper reaches of Sugure Khola forest stream, 1700 m a. s. l. NHMW." httpUri="https://binary.pensoft.net/fig/402717" pageId="3" pageNumber="4">, 3</figureCitation>
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<figureCitation captionStart="Figure 11" captionStartId="F3" captionText="Figure 11. Type locality of both Erhaia jannei and E. pelkiae; Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photo by EG." httpUri="https://binary.pensoft.net/fig/402718" pageId="3" pageNumber="4">, 11</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="4" type="reference_group">
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<paragraph pageId="3" pageNumber="4">
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<taxonomicName authorityName="Davis & Kuo" authorityYear="1985" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia" order="Littorinimorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="genus">Erhaia</taxonomicName>
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sp.
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<bibRefCitation DOI="https://doi.org/10.3897/zookeys.679.13326" author="Gittenberger, E" journalOrPublisher="ZooKeys" pageId="14" pageNumber="15" pagination="21 - 28" refId="B7" refString="Gittenberger, E, Sherub, S, Stelbrink, B, 2017a. Erhaia Davis & Kuo (Gastropoda, Rissooidea, Amnicolidae) also in Bhutan. ZooKeys 679: 21 - 28, DOI: https://doi.org/10.3897/zookeys.679.13326" title="Erhaia Davis & Kuo (Gastropoda, Rissooidea, Amnicolidae) also in Bhutan." url="https://doi.org/10.3897/zookeys.679.13326" volume="679" year="2017 a">Gittenberger et al. 2017a</bibRefCitation>
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: 25, fig. 3; 2017c: 900, 903, fig. 8.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="4" type="materials_examined">
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<paragraph pageId="3" pageNumber="4">Material examined.</paragraph>
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<paragraph pageId="3" pageNumber="4">
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<emphasis bold="true" pageId="3" pageNumber="4">
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<emphasis italics="true" pageId="3" pageNumber="4">Holotype</emphasis>
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.
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</emphasis>
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(Fig.
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<figureCitation captionStart="Figures 2–10" captionStartId="F2" captionText="Figures 2 - 10. Erhaia jannei sp. nov., holotype (2 H 2.25 mm) and paratype (3 H 2.02 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia pelkiae sp. nov., holotype (4 H 1.86 mm) and paratype (5 H 1.86 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia wangchuki Gittenberger, Sherub & Stelbrink, 2017. Shells from the type locality (6 H 1.98 mm) (7 H 2.03 mm); NBCB 1072. Bhutan, district Wangdue Phodrang, Gangchhu, Gangzetem brooklet, 2883 m a. s. l. Photos by J. Goud 8 Erhaia banepaensis Nesemann & S. Sharma, 2007, holotype (H 1.95 mm); NHMW 103319. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 9 Erhaia nainitalensis Davis & Rao, 1997, holotype of E. chandeshwariensis Nesemann & S. Sharma, 2007 (H 1.94 mm); NHMW 103315. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 10 Erhaia sugurensis Nesemann, Shah & Tachamo, 2007, holotype (H 1.95 mm); NHMW 104172. Nepal, Central Zone, Lalitpur district, Godawari, upper reaches of Sugure Khola forest stream, 1700 m a. s. l. NHMW." httpUri="https://binary.pensoft.net/fig/402717" pageId="3" pageNumber="4">2</figureCitation>
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) Bhutan • District Thimphu, c. 5 km E of Chhuzom, W of Geneykha; in brooklet with a prayer wheel along the road; 2750 m a.s.l.;
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<geoCoordinate degrees="27" direction="north" minutes="18" orientation="latitude" precision="15" seconds="43" value="27.311943">27°18'43"N</geoCoordinate>
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,
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<geoCoordinate degrees="89" direction="east" minutes="36" orientation="longitude" precision="15" seconds="10" value="89.602776">89°36'10"E</geoCoordinate>
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; E. Gittenberger, Choki Gyeltshen & Pema Leda leg. 25.X.2018; NBCB 1057.
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<emphasis bold="true" pageId="3" pageNumber="4">
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<emphasis italics="true" pageId="3" pageNumber="4">Paratypes</emphasis>
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.
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</emphasis>
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(Fig.
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<figureCitation captionStart="Figures 2–10" captionStartId="F2" captionText="Figures 2 - 10. Erhaia jannei sp. nov., holotype (2 H 2.25 mm) and paratype (3 H 2.02 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia pelkiae sp. nov., holotype (4 H 1.86 mm) and paratype (5 H 1.86 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia wangchuki Gittenberger, Sherub & Stelbrink, 2017. Shells from the type locality (6 H 1.98 mm) (7 H 2.03 mm); NBCB 1072. Bhutan, district Wangdue Phodrang, Gangchhu, Gangzetem brooklet, 2883 m a. s. l. Photos by J. Goud 8 Erhaia banepaensis Nesemann & S. Sharma, 2007, holotype (H 1.95 mm); NHMW 103319. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 9 Erhaia nainitalensis Davis & Rao, 1997, holotype of E. chandeshwariensis Nesemann & S. Sharma, 2007 (H 1.94 mm); NHMW 103315. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 10 Erhaia sugurensis Nesemann, Shah & Tachamo, 2007, holotype (H 1.95 mm); NHMW 104172. Nepal, Central Zone, Lalitpur district, Godawari, upper reaches of Sugure Khola forest stream, 1700 m a. s. l. NHMW." httpUri="https://binary.pensoft.net/fig/402717" pageId="3" pageNumber="4">3</figureCitation>
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) 6 shells and 2 animals in ethanol 70%; same data as for holotype; NBCB 1058.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="4" type="diagnosis">
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<paragraph pageId="3" pageNumber="4">Diagnosis.</paragraph>
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<paragraph pageId="3" pageNumber="4">Shell large for the genus (H> 2mm), ovoid, with a relatively large aperture.</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="4" type="shell">
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<paragraph pageId="3" pageNumber="4">Shell.</paragraph>
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<paragraph pageId="3" pageNumber="4">
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Shell obliquely ovoid, with
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<normalizedToken originalValue="3½">31/2</normalizedToken>
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-4 convex, shouldered whorls that are separated by a deep suture; clearly higher than broad; yellowish brown with fine irregular growth lines and some blackish brown periostracal ridges, one of which runs from the apertural columellar border into the umbilicus. Peristome not reflected. Parietal, columellar and a short part of the adjoining basal apertural border thickened by a whitish callus. Most specimens with a continuous peristome and a narrow umbilical chink. Protoconch encrusted in all specimens; teleoconch without spiral sculpture.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="4" type="description">
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<paragraph pageId="3" pageNumber="4">Measurements.</paragraph>
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<paragraph pageId="3" pageNumber="4">
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(
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<emphasis italics="true" pageId="3" pageNumber="4">N</emphasis>
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= 9): H 2.2-2.4 mm, B 1.5-1.6 mm. Holotype 2.2
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<normalizedToken originalValue="×">x</normalizedToken>
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1.6 mm.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="4" type="ecology">
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<paragraph pageId="3" pageNumber="4">Ecology</paragraph>
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<paragraph pageId="3" pageNumber="4">
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(Fig.
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<figureCitation captionStart="Figure 11" captionStartId="F3" captionText="Figure 11. Type locality of both Erhaia jannei and E. pelkiae; Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photo by EG." httpUri="https://binary.pensoft.net/fig/402718" pageId="3" pageNumber="4">11</figureCitation>
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). The species is uncommon on the rocks in the shaded streamlet uphill of the prayer wheel, where it occurs with
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<taxonomicName baseAuthorityName="O. F. Müller" baseAuthorityYear="1774" class="Gastropoda" family="Lymnaeidae" genus="Galba" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Galba truncatula" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="truncatula">
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<emphasis italics="true" pageId="3" pageNumber="4">Galba truncatula</emphasis>
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</taxonomicName>
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(
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<normalizedToken originalValue="Müller">Mueller</normalizedToken>
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, 1774),
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<taxonomicName class="Gastropoda" family="Physidae" genus="Physa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Physa" order="Architaenioglossa" pageId="3" pageNumber="4" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="3" pageNumber="4">Physa</emphasis>
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</taxonomicName>
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sp. (new for Bhutan), and
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<taxonomicName authorityName="Gittenberger & Gyeltshen" authorityYear="2020" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia pelkiae" order="Littorinimorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="pelkiae">
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<emphasis italics="true" pageId="3" pageNumber="4">Erhaia pelkiae</emphasis>
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</taxonomicName>
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sp. nov.
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|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="3" pageNumber="4" type="dna data">
|
|||
|
|
<paragraph pageId="3" pageNumber="4">DNA data</paragraph>
|
|||
|
|
<paragraph pageId="3" pageNumber="4">
|
|||
|
|
(Fig.
|
|||
|
|
<figureCitation captionStart="Figure 18" captionStartId="F9" captionText="Figure 18. Maximum likelihood tree reconstructed with RAxML BlackBox (Stamatakis et al. 2008; GTR + G substitution model for each partition and 100 bootstrap replicates) based on the 16 S rRNA and COI dataset of Liu et al. (2014) and Guan et al. (2008), with new data in red. Numbers on branches denote bootstrap values> 50." httpUri="https://binary.pensoft.net/fig/402724" pageId="3" pageNumber="4">18</figureCitation>
|
|||
|
|
). The three individuals shared the same haplotype for both 16S rRNA (GenBank acc. no. MT239078) and COI (GenBank acc. no. MT237716). The uncorrected genetic p-distances between this species and
|
|||
|
|
<taxonomicName lsidName="E. wangchuki" pageId="3" pageNumber="4" rank="species" species="wangchuki">
|
|||
|
|
<emphasis italics="true" pageId="3" pageNumber="4">E. wangchuki</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
were 0.84% for 16S rRNA and 4.87% for COI. The distances were considerably higher when compared to
|
|||
|
|
<taxonomicName authorityName="Davis & Kuo" authorityYear="1985" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia" order="Littorinimorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="genus">
|
|||
|
|
<emphasis italics="true" pageId="3" pageNumber="4">Erhaia</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
sp. from China, Guangxi, viz. 2.74% for 16S rRNA and 10.25% for COI (GenBank acc. nos. KC832722 and KC832701, respectively).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="3" pageNumber="4" type="notes">
|
|||
|
|
<paragraph pageId="3" pageNumber="4">Notes.</paragraph>
|
|||
|
|
<paragraph pageId="3" pageNumber="4">This species was discovered in 2012, but since only a single shell was collected then, a description was considered premature.</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="3" pageNumber="4" type="etymology">
|
|||
|
|
<paragraph pageId="3" pageNumber="4">Etymology.</paragraph>
|
|||
|
|
<paragraph pageId="3" pageNumber="4">
|
|||
|
|
The epithet
|
|||
|
|
<taxonomicName authorityName="Gittenberger & Stelbrink" authorityYear="2020" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="jannei" order="Littorinimorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="jannei">
|
|||
|
|
<emphasis italics="true" pageId="3" pageNumber="4">jannei</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
refers to Mr Janne Clewing, the son of the last author.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption httpUri="https://binary.pensoft.net/fig/402717" pageId="3" pageNumber="4" start="Figures 2–10" startId="F2">
|
|||
|
|
<paragraph pageId="3" pageNumber="4">
|
|||
|
|
<emphasis bold="true" pageId="3" pageNumber="4">Figures 2-10.</emphasis>
|
|||
|
|
<taxonomicName authorityName="Gittenberger & Stelbrink" authorityYear="2020" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia jannei" order="Littorinimorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="jannei">
|
|||
|
|
<emphasis italics="true" pageId="3" pageNumber="4">Erhaia jannei</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
sp. nov., holotype (
|
|||
|
|
<emphasis bold="true" pageId="3" pageNumber="4">2</emphasis>
|
|||
|
|
H 2.25 mm) and paratype (
|
|||
|
|
<emphasis bold="true" pageId="3" pageNumber="4">3</emphasis>
|
|||
|
|
H 2.02 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a.s.l. Photos by J. Goud
|
|||
|
|
<taxonomicName authorityName="Gittenberger & Gyeltshen" authorityYear="2020" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia pelkiae" order="Littorinimorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="pelkiae">
|
|||
|
|
<emphasis italics="true" pageId="3" pageNumber="4">Erhaia pelkiae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
sp. nov., holotype (
|
|||
|
|
<emphasis bold="true" pageId="3" pageNumber="4">4</emphasis>
|
|||
|
|
H 1.86 mm) and paratype (
|
|||
|
|
<emphasis bold="true" pageId="3" pageNumber="4">5</emphasis>
|
|||
|
|
H 1.86 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a.s.l. Photos by J. Goud
|
|||
|
|
<taxonomicName authorityName="Gittenberger, Sherub & Stelbrink" authorityYear="2017" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia wangchuki" order="Littorinimorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="wangchuki">
|
|||
|
|
<emphasis italics="true" pageId="3" pageNumber="4">Erhaia wangchuki</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
Gittenberger, Sherub & Stelbrink, 2017. Shells from the type locality (
|
|||
|
|
<emphasis bold="true" pageId="3" pageNumber="4">6</emphasis>
|
|||
|
|
H 1.98 mm) (
|
|||
|
|
<emphasis bold="true" pageId="3" pageNumber="4">7</emphasis>
|
|||
|
|
H 2.03 mm); NBCB 1072. Bhutan, district Wangdue Phodrang, Gangchhu, Gangzetem brooklet, 2883 m a.s.l. Photos by J. Goud
|
|||
|
|
<emphasis bold="true" pageId="3" pageNumber="4">8</emphasis>
|
|||
|
|
<taxonomicName authorityName="Nesemann & S. Sharma" authorityYear="2007" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia banepaensis" order="Littorinimorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="banepaensis">
|
|||
|
|
<emphasis italics="true" pageId="3" pageNumber="4">Erhaia banepaensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
Nesemann & S. Sharma, 2007, holotype (H 1.95 mm); NHMW 103319. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW
|
|||
|
|
<emphasis bold="true" pageId="3" pageNumber="4">9</emphasis>
|
|||
|
|
<taxonomicName authorityName="Davis & Rao" authorityYear="1997" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia nainitalensis" order="Littorinimorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="nainitalensis">
|
|||
|
|
<emphasis italics="true" pageId="3" pageNumber="4">Erhaia nainitalensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
Davis & Rao, 1997, holotype of
|
|||
|
|
<taxonomicName lsidName="E. chandeshwariensis" pageId="3" pageNumber="4" rank="species" species="chandeshwariensis">
|
|||
|
|
<emphasis italics="true" pageId="3" pageNumber="4">E. chandeshwariensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
Nesemann & S. Sharma, 2007 (H 1.94 mm); NHMW 103315. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW
|
|||
|
|
<emphasis bold="true" pageId="3" pageNumber="4">10</emphasis>
|
|||
|
|
<taxonomicName authorityName="Nesemann, Shah & Tachamo" authorityYear="2007" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia sugurensis" order="Littorinimorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="sugurensis">
|
|||
|
|
<emphasis italics="true" pageId="3" pageNumber="4">Erhaia sugurensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
Nesemann, Shah & Tachamo, 2007, holotype (H 1.95 mm); NHMW 104172. Nepal, Central Zone, Lalitpur district, Godawari, upper reaches of Sugure Khola forest stream, 1700 m a.s.l. NHMW.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption httpUri="https://binary.pensoft.net/fig/402718" pageId="3" pageNumber="4" start="Figure 11" startId="F3">
|
|||
|
|
<paragraph pageId="3" pageNumber="4">
|
|||
|
|
<emphasis bold="true" pageId="3" pageNumber="4">Figure 11.</emphasis>
|
|||
|
|
Type locality of both
|
|||
|
|
<taxonomicName authorityName="Gittenberger & Stelbrink" authorityYear="2020" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia jannei" order="Littorinimorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="jannei">
|
|||
|
|
<emphasis italics="true" pageId="3" pageNumber="4">Erhaia jannei</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName lsidName="E. pelkiae" pageId="3" pageNumber="4" rank="species" species="pelkiae">
|
|||
|
|
<emphasis italics="true" pageId="3" pageNumber="4">E. pelkiae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
; Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a.s.l. Photo by EG.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|