<documentid="9116C8C24BEA58F23537CD84CD2F6B99"ID-CLB-Dataset="269786"ID-DOI="10.11646/zootaxa.5346.4.4"ID-GBIF-Dataset="f631dec3-14e8-4dab-b340-df57d7cc8c5a"ID-ISSN="1175-5326"ID-Zenodo-Dep="8390441"ID-ZooBank="3D23DF3F-6B66-4C60-BE3D-A98EA43F01C5"IM.bibliography_approvedBy="diego"IM.illustrations_approvedBy="felipe"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="felipe"IM.taxonomicNames_approvedBy="felipe"IM.treatments_approvedBy="diego"checkinTime="1695985560601"checkinUser="plazi"docAuthor="Tanasevitch, Andrei V."docDate="2023"docId="03A9879D977DFF9FFF62F6C4D7D87AA9"docLanguage="en"docName="zootaxa.5346.4.4.pdf"docOrigin="Zootaxa 5346 (4)"docSource="http://dx.doi.org/10.11646/zootaxa.5346.4.4"docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article"docStyleId="647186512141C8FC8976D5BCC54AEB7D"docStyleName="Zootaxa.2013-.journal_article"docStyleVersion="9"docTitle="Savignia ericola Tanasevitch 2023, new species"docType="treatment"docVersion="6"lastPageNumber="437"masterDocId="FF90FFE59772FF8EFFF5F05FD6617B78"masterDocTitle="Survey of the Ethiopian linyphiid spider fauna. I. Subfamily Erigoninae (Arachnida, Araneae, Linyphiidae)"masterLastPageNumber="442"masterPageNumber="420"pageNumber="435"updateTime="1732480545820"updateUser="ExternalLinkService"zenodo-license-document="CLOSED">
<mods:titleid="960600E398915E3135529153376E6C42">Survey of the Ethiopian linyphiid spider fauna. I. Subfamily Erigoninae (Arachnida, Araneae, Linyphiidae)</mods:title>
<figureCitationid="133B2A0E977DFF81FF62F6E0D68A7DA1"box="[151,235,1727,1753]"captionStart="FIGURE 11"captionStartId="16.[151,250,1697,1721]"captionTargetBox="[158,1429,604,1667]"captionTargetId="figure-245@16.[151,1436,597,1673]"captionTargetPageId="16"captionText="FIGURE 11. Photographs of male holotype (A–D) of Savignia ericola n. sp. A, habitus, dorsal view; B, C, same, lateral and anterolateral view, respectively; D, same, frontal view. Scale bar: 0.5mm."figureDoi="http://doi.org/10.5281/zenodo.8390477"httpUri="https://zenodo.org/record/8390477/files/figure.png"pageId="15"pageNumber="435">Figs 11</figureCitation>
,
<figureCitationid="133B2A0E977DFF81FF02F6E0D7727DA1"box="[247,275,1727,1753]"captionStart="FIGURE 12"captionStartId="17.[151,250,1440,1464]"captionTargetBox="[167,1433,489,1401]"captionTargetId="figure-134@17.[151,1436,489,1417]"captionTargetPageId="17"captionText="FIGURE 12. Details of male palpal structure (A–G) of holotype of Savignia ericola n. sp. A, left palp, retrolateral view; B, tibia, dorsal view; C, tegulum, distal suprategular apophysis and embolic division, prolateral view; D, distal suprategular apophysis, lateral view; E, embolic division, mesal view; F, apex of median radical process; G, apex of embolus. Scale bars: A–E, 0.1mm; G, F, not to scale."figureDoi="http://doi.org/10.5281/zenodo.8390479"httpUri="https://zenodo.org/record/8390479/files/figure.png"pageId="15"pageNumber="435">12</figureCitation>
<materialsCitationid="3B683CD6977DFF81FEBBF758D2827C11"ID-GBIF-Occurrence="4417449301"collectingDate="2022-10-30"collectionCode="ZMMU"collectorName="Arsi Mountains N. P. & A. Tanasevitch"country="Ethiopia"county="Asela Zone"elevation="3503"latitude="7.81944"location="Arsi Mountains"longLatPrecision="1"longitude="39.35429"municipality="Asela"pageId="15"pageNumber="435"specimenCount="1"stateProvince="Oromiya"typeStatus="holotype">
Based on the formula of chaetotaxy (2.2.1.1) and trichobothriotaxy (Tm I–III present, TmIV absent), the modified carapace, as well as the hypertrophied distal suprategular apophysis, the new species is evidently a member of the
<bibRefCitationid="EF914B7A977DFF81FD50F780D53D7C81"author="Millidge, A. F."box="[677,860,2015,2041]"pageId="15"pageNumber="435"pagination="1 - 60"refId="ref11443"refString="Millidge, A. F. (1977) The conformation of the male palpal organs of linyphiid spiders, and its application to the taxonomic and phylogenetic analysis of the family (Araneae: Linyphiidae). Bulletin of the British Arachnological Society, 4, 1 - 60."type="journal article"year="1977">Millidge (1977)</bibRefCitation>
. Among the genera of this group, the new species seems most appropriate to be provisionally assigned to
<bibRefCitationid="EF914B7A9762FF9EFC91F0C8D5947BC9"author="Simon, E."box="[868,1013,151,177]"pageId="16"pageNumber="436"pagination="180 - 885"refId="ref11731"refString="Simon, E. (1884) Les arachnides de France. Tome cinquieme, deuxieme et troisieme partie. Roret, Paris, pp. 180 - 885, pls. 26 - 27."type="book chapter"year="1884">Simon, 1884</bibRefCitation>
</taxonomicName>
, as certain resemble is observed in the structure of the embolic division, namely, the peculiar shape of the embolus, as well as by the presence of a narrow process at its base (MRP in
<figureCitationid="133B2A0E9762FF9EFE24F080D4547B81"box="[465,565,223,249]"captionStart="FIGURE 12"captionStartId="17.[151,250,1440,1464]"captionTargetBox="[167,1433,489,1401]"captionTargetId="figure-134@17.[151,1436,489,1417]"captionTargetPageId="17"captionText="FIGURE 12. Details of male palpal structure (A–G) of holotype of Savignia ericola n. sp. A, left palp, retrolateral view; B, tibia, dorsal view; C, tegulum, distal suprategular apophysis and embolic division, prolateral view; D, distal suprategular apophysis, lateral view; E, embolic division, mesal view; F, apex of median radical process; G, apex of embolus. Scale bars: A–E, 0.1mm; G, F, not to scale."figureDoi="http://doi.org/10.5281/zenodo.8390479"httpUri="https://zenodo.org/record/8390479/files/figure.png"pageId="16"pageNumber="436">Fig. 12E</figureCitation>
). The shape of the embolus is similar to that in many Asian congeners, e.g.
<bibRefCitationid="EF914B7A9762FF9EFEE4F15CD7FF7A65"author="Eskov, K. Y."box="[273,414,259,285]"pageId="16"pageNumber="436"pagination="140 - 144"refId="ref10791"refString="Eskov, K. Y. (1991) A spider genus Savignya (s. str.) (Aranei, Linyphiidae) in the fauna of the Far East and central Asia. Zoologicheskii Zhurnal, 70 (5), 140 - 144."type="journal article"year="1991">Eskov, 1991</bibRefCitation>
</taxonomicName>
,
<taxonomicNameid="4C004D089762FF9EFE5EF15CD5CE7A65"authority="Marusik, Koponen & Danilov, 2001"authorityName="Marusik, Koponen & Danilov"authorityYear="2001"box="[427,943,259,285]"class="Arachnida"family="Linyphiidae"genus="Savignia"kingdom="Animalia"order="Araneae"pageId="16"pageNumber="436"phylum="Arthropoda"rank="species"species="eskovi">
<bibRefCitationid="EF914B7A9762FF9EFDE0F15CD5CE7A65"author="Marusik, Y. M. & Koponen, S. & Danilov, S. N."box="[533,943,259,285]"pageId="16"pageNumber="436"pagination="83 - 92"refId="ref11279"refString="Marusik, Y. M., Koponen, S. & Danilov, S. N. (2001) Taxonomic and faunistic notes on linyphiids of Transbaikalia and south Siberia (Araneae, Linyphiidae). Bulletin of the British Arachnological Society, 12, 83 - 92."type="journal article"year="2001">Marusik, Koponen & Danilov, 2001</bibRefCitation>
<bibRefCitationid="EF914B7A9762FF9EFBE8F15CD2CA7A65"author="Eskov, K. Y."box="[1053,1195,259,285]"pageId="16"pageNumber="436"pagination="13 - 39"refId="ref10745"refString="Eskov, K. Y. (1988) The spider genera Savignya Blackwall, Diplocephalus Bertkau and Archaraeoncus Tanasevitch (Aranei, Linyphiidae) in the fauna of Siberia and the Soviet Far East. Folia Entomologica Hungarica, 49, 13 - 39."type="journal article"year="1988">Eskov, 1988</bibRefCitation>
<bibRefCitationid="EF914B7A9762FF9EFAFFF15CD3F97A65"author="Eskov, K. Y."box="[1290,1432,259,285]"pageId="16"pageNumber="436"pagination="13 - 39"refId="ref10745"refString="Eskov, K. Y. (1988) The spider genera Savignya Blackwall, Diplocephalus Bertkau and Archaraeoncus Tanasevitch (Aranei, Linyphiidae) in the fauna of Siberia and the Soviet Far East. Folia Entomologica Hungarica, 49, 13 - 39."type="journal article"year="1988">Eskov, 1988</bibRefCitation>
</taxonomicName>
, etc., all of which also show a median radical process in the embolic division. However, the new species stands apart among other congeners in lacking of a mesal outgrowth on the distal suprategular apophysis (MOSA in
<bibRefCitationid="EF914B7A9762FF9EFAE2F114D71E7AF1"author="Tanasevitch, A. V. & Trilikauskas, L. A."pageId="16"pageNumber="436"pagination="269 - 270"refId="ref11767"refString="Tanasevitch, A. V. & Trilikauskas, L. A. (2006) A new species of the genus Savignia Blackwall from the Khabarovsk Province, Russian Far East (Aranei: Linyphiidae: Erigoninae). Arthropoda Selecta, 14 (3), 269 - 270. [for 2005]"type="journal article"year="2006">Tanasevitch & Trilikauskas 2006</bibRefCitation>
, figs 2, 4), a significant generic synapomorphy. Besides this,
(namely, with a bisected ventral plate), then the placement of the new species into the genus will become justified. At the same time, it is possible that the new species represents a new genus of the
. Total length 1.83. Carapace slightly modified as shown in
<figureCitationid="133B2A0E9762FF9EFB3BF754D33A7C5D"box="[1230,1371,1803,1829]"captionStart="FIGURE 11"captionStartId="16.[151,250,1697,1721]"captionTargetBox="[158,1429,604,1667]"captionTargetId="figure-245@16.[151,1436,597,1673]"captionTargetPageId="16"captionText="FIGURE 11. Photographs of male holotype (A–D) of Savignia ericola n. sp. A, habitus, dorsal view; B, C, same, lateral and anterolateral view, respectively; D, same, frontal view. Scale bar: 0.5mm."figureDoi="http://doi.org/10.5281/zenodo.8390477"httpUri="https://zenodo.org/record/8390477/files/figure.png"pageId="16"pageNumber="436">Figs 11A–C</figureCitation>
, 1.03 long, 0.63 wide, pale greyish brown to greyish yellow. Anterior part of carapace somewhat protruded anteriorly, eyes relatively small. Chelicerae 0.22 long, a mastidion absent. Legs yellow. Leg I 2.06 long (0.63 + 0.22 + 0.48 + 0.40 + 0.33), IV 2.19 long (0.63 + 0.20 +0.58 + 0.48 + 0.30). Chaetotaxy 2.2.1.1, spines about as long as 0.5–1 diameter of corresponding leg segment. Metatarsi I–III with a trichobothrium each. TmI 0.41. Palp (
<figureCitationid="133B2A0E9762FF9EFB0AF7C4D3F07CCD"box="[1279,1425,1947,1973]"captionStart="FIGURE 12"captionStartId="17.[151,250,1440,1464]"captionTargetBox="[167,1433,489,1401]"captionTargetId="figure-134@17.[151,1436,489,1417]"captionTargetPageId="17"captionText="FIGURE 12. Details of male palpal structure (A–G) of holotype of Savignia ericola n. sp. A, left palp, retrolateral view; B, tibia, dorsal view; C, tegulum, distal suprategular apophysis and embolic division, prolateral view; D, distal suprategular apophysis, lateral view; E, embolic division, mesal view; F, apex of median radical process; G, apex of embolus. Scale bars: A–E, 0.1mm; G, F, not to scale."figureDoi="http://doi.org/10.5281/zenodo.8390479"httpUri="https://zenodo.org/record/8390479/files/figure.png"pageId="16"pageNumber="436">Figs 12A–G</figureCitation>
): Patella slightly elongated. Tibia with a dorsal, conical, apically rounded outgrowth and a small projection bearing three long, stout and slightly curved setae. Paracymbium J-shaped. Distal suprategular apophysis very large, massive, ending with a strong, straight spike. Radix rounded proximally, with a membranous tissue near middle, a flat, subtrapeziform, anterior apophysis (ARA in
<figureCitationid="133B2A0E9763FF9FFD61F080D49B7B81"box="[660,762,223,249]"captionStart="FIGURE 12"captionStartId="17.[151,250,1440,1464]"captionTargetBox="[167,1433,489,1401]"captionTargetId="figure-134@17.[151,1436,489,1417]"captionTargetPageId="17"captionText="FIGURE 12. Details of male palpal structure (A–G) of holotype of Savignia ericola n. sp. A, left palp, retrolateral view; B, tibia, dorsal view; C, tegulum, distal suprategular apophysis and embolic division, prolateral view; D, distal suprategular apophysis, lateral view; E, embolic division, mesal view; F, apex of median radical process; G, apex of embolus. Scale bars: A–E, 0.1mm; G, F, not to scale."figureDoi="http://doi.org/10.5281/zenodo.8390479"httpUri="https://zenodo.org/record/8390479/files/figure.png"pageId="17"pageNumber="437">Fig. 12E</figureCitation>
), and an relatively long, narrow, apically unciform process (
<figureCitationid="133B2A0E9763FF9FFF6AF15CD7427A65"box="[159,291,259,285]"captionStart="FIGURE 12"captionStartId="17.[151,250,1440,1464]"captionTargetBox="[167,1433,489,1401]"captionTargetId="figure-134@17.[151,1436,489,1417]"captionTargetPageId="17"captionText="FIGURE 12. Details of male palpal structure (A–G) of holotype of Savignia ericola n. sp. A, left palp, retrolateral view; B, tibia, dorsal view; C, tegulum, distal suprategular apophysis and embolic division, prolateral view; D, distal suprategular apophysis, lateral view; E, embolic division, mesal view; F, apex of median radical process; G, apex of embolus. Scale bars: A–E, 0.1mm; G, F, not to scale."figureDoi="http://doi.org/10.5281/zenodo.8390479"httpUri="https://zenodo.org/record/8390479/files/figure.png"pageId="17"pageNumber="437">Figs 12E, F</figureCitation>
) situated at base of embolus. Embolus starting from middle of radix as a narrow stripe, sharply curved in middle part to gradually taper thereafter into a thin and curved tip. Opisthosoma 0.90 long, 0.58 wide, pale grey (
<figureCitationid="133B2A0E9763FF9FFF6AF114D74D7A1D"box="[159,300,331,357]"captionStart="FIGURE 11"captionStartId="16.[151,250,1697,1721]"captionTargetBox="[158,1429,604,1667]"captionTargetId="figure-245@16.[151,1436,597,1673]"captionTargetPageId="16"captionText="FIGURE 11. Photographs of male holotype (A–D) of Savignia ericola n. sp. A, habitus, dorsal view; B, C, same, lateral and anterolateral view, respectively; D, same, frontal view. Scale bar: 0.5mm."figureDoi="http://doi.org/10.5281/zenodo.8390477"httpUri="https://zenodo.org/record/8390477/files/figure.png"pageId="17"pageNumber="437">Figs 11A–C</figureCitation>
