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<document id= "E8AC9A46264EE29F02505E890F8DAA9C" ID-CLB-Dataset= "25883" ID-DOI= "http://dx.doi.org/10.3897/zookeys.867.28728" ID-GBIF-Dataset= "bff0bb63-a934-4fa2-ae76-50c4db24d98d" ID-PMC= "PMC6685932" ID-Pensoft-Pub= "1313-2970-867-139" ID-Pensoft-UUID= "729147F7504F521DB514FA3017CB4CD3" ID-PubMed= "31406481" ID-ZooBank= "9765595E1A204418B5C6AC4B6F587CE5" ModsDocID= "1313-2970-867-139" checkinTime= "1564648495948" checkinUser= "pensoft" docAuthor= "Goldstein, Paul Z. & Zilli, Alberto" docDate= "2019" docId= "8DEC755AE41C5E8E99A65011D0AF8368" docLanguage= "en" docName= "ZooKeys 867: 139-160" docOrigin= "ZooKeys 867" docSource= "http://dx.doi.org/10.3897/zookeys.867.28728" docTitle= "Phuphena Walker 1858" docType= "treatment" docVersion= "6" lastPageNumber= "139" masterDocId= "414CFF867946142B7D60E5509002FFFB" masterDocTitle= "Thraumata, a new genus from South America with description of a new species from Peru (Lepidoptera, Noctuidae)" masterLastPageNumber= "160" masterPageNumber= "139" pageNumber= "139" updateTime= "1732786731291" updateUser= "ExternalLinkService" >
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<mods:title id= "CBBC094A0CAF2507ADCE1B5B476FE16E" > Thraumata, a new genus from South America with description of a new species from Peru (Lepidoptera, Noctuidae)</mods:title>
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<mods:namePart id= "568347959B98E739C583591A6E845424" > Goldstein, Paul Z.</mods:namePart>
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<mods:namePart id= "567F42914085D19B1D88668BFB3DB1C0" > Zilli, Alberto</mods:namePart>
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<mods:title id= "BF537775BE2ED3123749352C6FEE350F" > ZooKeys</mods:title>
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<treatment id= "8DEC755AE41C5E8E99A65011D0AF8368" ID-GBIF-Taxon= "159293017" LSID= "urn:lsid:plazi:treatment:8DEC755AE41C5E8E99A65011D0AF8368" httpUri= "http://treatment.plazi.org/id/8DEC755AE41C5E8E99A65011D0AF8368" lastPageNumber= "139" pageId= "0" pageNumber= "139" scope_family= "Noctuidae" scope_order= "Lepidoptera" >
<subSubSection id= "2E19A9FDCE4E8AE8C3CD1A5EEE57278C" pageId= "0" pageNumber= "139" type= "nomenclature" >
<paragraph id= "7B054C6A85B479F1142ADFC06130BA70" pageId= "0" pageNumber= "139" >
<taxonomicName id= "318D8D3DA0FF2BF3AE8D0B713A32030E" ID-CoL= "92JXN" LSID= "8dec755a-e41c-5e8e-99a6-5011d0af8368" authority= "Walker, 1858" authorityName= "Walker" authorityYear= "1858" class= "Insecta" family= "Noctuidae" genus= "Phuphena" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Phuphena" order= "Lepidoptera" pageId= "0" pageNumber= "139" phylum= "Arthropoda" rank= "genus" > Phuphena Walker, 1858</taxonomicName>
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</paragraph>
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<subSubSection id= "8D7965096112AD8901B248B116E88ADC" pageId= "0" pageNumber= "139" type= "type species" >
<paragraph id= "04B9AAD2AAC6FE4B1988C245F7E68C61" pageId= "0" pageNumber= "139" > Type species.</paragraph>
<paragraph id= "0055D9B3B6CA9F7C5064757C8F0C7630" pageId= "0" pageNumber= "139" >
<taxonomicName id= "DB3027662537F6DEE18DE1FCDAF6FE8B" class= "Insecta" family= "Noctuidae" genus= "Phuphena" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Phuphena fusipennis" order= "Lepidoptera" pageId= "0" pageNumber= "139" phylum= "Arthropoda" rank= "species" species= "fusipennis" >
<emphasis id= "E6D8799EBC79F1558C48A2B4B4866528" italics= "true" pageId= "0" pageNumber= "139" > Phuphena fusipennis</emphasis>
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Walker, 1858 by monotypy.
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<subSubSection id= "A95D51B8BA2892E1B165F795CD910565" pageId= "0" pageNumber= "139" type= "diagnosis" >
<paragraph id= "8F21306A64EB330BF7C135A36CB3E463" pageId= "0" pageNumber= "139" > Diagnosis.</paragraph>
<paragraph id= "04137469C676DB8E787BBB14966DB273" pageId= "0" pageNumber= "139" >
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Species of
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<taxonomicName id= "1C83B11DB98A6F6EFF3A1886D079A320" class= "Insecta" family= "Noctuidae" genus= "Phuphena" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Phuphena" order= "Lepidoptera" pageId= "0" pageNumber= "139" phylum= "Arthropoda" rank= "genus" >
<emphasis id= "6A2B9F2B747B3F1AEED766A070E690DA" italics= "true" pageId= "0" pageNumber= "139" > Phuphena</emphasis>
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</taxonomicName>
diverge with respect to wing pattern, but most share a bounded medial area. Orbicular and reniform stigmata sometimes absent or indistinct; outer forewing margin smooth in the majority of species but crenulate in
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<taxonomicName id= "12295E721BB6A2EE827C199718F9CF3E" lsidName= "cilix" pageId= "0" pageNumber= "139" rank= "species" species= "cilix" >
<emphasis id= "0AE553B20ADCD2B244D551D7C03A4EE1" italics= "true" pageId= "0" pageNumber= "139" > cilix</emphasis>
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</taxonomicName>
. Hindwing upperside coloration uniform. Male genitalia of
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<taxonomicName id= "18B7EDA9DC297CFDDDD7E067516C73DE" class= "Insecta" family= "Noctuidae" genus= "Phuphena" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Phuphena" order= "Lepidoptera" pageId= "0" pageNumber= "139" phylum= "Arthropoda" rank= "genus" >
<emphasis id= "684E47D017B2070508FE2EF2119C3885" italics= "true" pageId= "0" pageNumber= "139" > Phuphena</emphasis>
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distinct, the valvae uniquely reduced, with no evidence of clasping architecture, narrow for most of their length, and in most species swollen apically to form a knob-like cucullus, giving them a club-shaped appearance, with a corona comprising an unorganized arrangement of setae. Vinculum V-shaped, slightly truncate in some species, and extending below the base of the valves for a distance equal to or greater than the width at its broadest point. The phallus is asymmetrically sclerotized, often with a patch of scale-like cornuti, and the vesica without well-developed cornuti or diverticula, but with a corresponding ventro-distal strip of scale-like cornuti. Basal abdominal brushes present but not visibly subdivided as in
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<taxonomicName id= "04FA5A89E0AC807C647FC8491137ABB8" class= "Insecta" family= "Noctuidae" genus= "Callopistria" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Callopistria" order= "Lepidoptera" pageId= "0" pageNumber= "139" phylum= "Arthropoda" rank= "genus" >
<emphasis id= "A36308DCFEFC29FAC94039441D05A5F0" italics= "true" pageId= "0" pageNumber= "139" > Callopistria</emphasis>
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</taxonomicName>
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<normalizedToken id= "8D0EA9280D89F721DD4DCD4BA5EAE353" originalValue= "Hübner" > Huebner</normalizedToken>
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, [1821] (type species:
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<taxonomicName id= "30D29F57A11D18C5485B3097502DBDDD" class= "Insecta" family= "Noctuidae" genus= "Noctua" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Noctua juventina" order= "Lepidoptera" pageId= "0" pageNumber= "139" phylum= "Arthropoda" rank= "species" species= "juventina" >
<emphasis id= "20B55E880507C5E0B49F2E16ECD80484" italics= "true" pageId= "0" pageNumber= "139" > Noctua juventina</emphasis>
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Stoll, 1782); a smaller pair on A8, and eversible saccular hair pencils completely absent. Female genitalia simple, the corpus bursae lemon-shaped, with no appendix or signa, the ductus seminalis attached apically, to the distal (anterior) end of the corpus. This distinguishes
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<taxonomicName id= "B6C7A1E2199014A75A7E1B836CBF7977" class= "Insecta" family= "Noctuidae" genus= "Phuphena" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Phuphena" order= "Lepidoptera" pageId= "0" pageNumber= "139" phylum= "Arthropoda" rank= "genus" >
<emphasis id= "C2C4B60BD95FC83320FE7AC328498073" italics= "true" pageId= "0" pageNumber= "139" > Phuphena</emphasis>
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from all
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<taxonomicName id= "01E499E3882A3A6DF000666BB372445E" genus= "Thraumata" lsidName= "Thraumata" pageId= "0" pageNumber= "139" rank= "genus" >
<emphasis id= "77332DCDB3256B3604C3612BA7C25C7C" italics= "true" pageId= "0" pageNumber= "139" > Thraumata</emphasis>
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</taxonomicName>
spp. and from
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<taxonomicName id= "6A1A663BD39CFE13D7545DF072347B35" class= "Insecta" family= "Noctuidae" genus= "Callopistria" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Callopistria juventina" order= "Lepidoptera" pageId= "0" pageNumber= "139" phylum= "Arthropoda" rank= "species" species= "juventina" >
<emphasis id= "BE31DFB76AABCAE1670A91AC16818D9F" italics= "true" pageId= "0" pageNumber= "139" > Callopistria juventina</emphasis>
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</taxonomicName>
, in which the ductus arises from the proximal (posterior) end of the corpus, but this character varies among other
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<taxonomicName id= "1A7EBBA98684267920DD059DA50CD04E" class= "Insecta" family= "Noctuidae" genus= "Callopistria" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Callopistria" order= "Lepidoptera" pageId= "0" pageNumber= "139" phylum= "Arthropoda" rank= "genus" >
<emphasis id= "DE5144F52B0B229F95B9E0B3BF4CCD40" italics= "true" pageId= "0" pageNumber= "139" > Callopistria</emphasis>
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species.
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