<documentid="34918F69424EDA4D56FB8640A328B36F"ID-CLB-Dataset="264199"ID-DOI="10.11646/zootaxa.5332.1.1"ID-GBIF-Dataset="66e0ec51-d494-43d4-965e-a2cd1462ef54"ID-ISSN="1175-5326"ID-Zenodo-Dep="8261502"ID-ZooBank="424F7439-4095-46A5-93E3-C4130E3B6D9A"IM.bibliography_approvedBy="juliana"IM.illustrations_approvedBy="juliana"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="felipe"IM.taxonomicNames_approvedBy="felipe"IM.treatmentCitations_approvedBy="felipe"IM.treatments_approvedBy="juliana"checkinTime="1692350092877"checkinUser="plazi"docAuthor="POORANI, J."docDate="2023"docId="03C7416214334658BDDF32B3FD36FDFA"docLanguage="en"docName="zootaxa.5332.1.1.pdf"docOrigin="Zootaxa 5332 (1)"docSource="http://dx.doi.org/10.11646/zootaxa.5332.1.1"docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph"docStyleId="5EBBA59367AD13919D70D935FA04F6A3"docStyleName="Zootaxa.2013-.monograph"docStyleVersion="19"docTitle="Coelophora saucia"docType="treatment"docVersion="10"lastPageNumber="112"masterDocId="FFFE391A14584637BD48367CFFCDFF9B"masterDocTitle="An illustrated guide to lady beetles (Coleoptera: Coccinellidae) of the Indian Subcontinent. Part 1. Tribe Coccinellini"masterLastPageNumber="307"masterPageNumber="1"pageNumber="108"updateTime="1732647533218"updateUser="ExternalLinkService">
<mods:titleid="DD57F638D6F5D1F00FB332869D9F759D">An illustrated guide to lady beetles (Coleoptera: Coccinellidae) of the Indian Subcontinent. Part 1. Tribe Coccinellini</mods:title>
<figureCitationid="1355ECF11433465CBDD7328FFF3BFA96"box="[159,246,1267,1293]"captionStart="FIGURE 73"captionStartId="109.[151,250,1943,1967]"captionTargetBox="[170,1418,187,1916]"captionTargetId="figure-15@109.[160,1427,181,1920]"captionTargetPageId="109"captionText="FIGURE 73. Coelophora saucia (Mulsant): a. adult, dorsal view; b. abdomen, male; c. terminal abdominal ventrites, female; d–g. male genitalia: d. tegmen, lateral view; e. tegmen, ventral view; f. penis; g. penis apex; h. spermatheca."figureDoi="http://doi.org/10.5281/zenodo.8261741"httpUri="https://zenodo.org/record/8261741/files/figure.png"pageId="107"pageNumber="108">Figs 73</figureCitation>
,
<figureCitationid="1355ECF11433465CBC4A328FFED2FA96"box="[258,287,1267,1293]"captionStart="FIGURE 74"captionStartId="110.[151,250,1943,1967]"captionTargetBox="[233,1354,181,1919]"captionTargetId="figure-20@110.[233,1354,181,1919]"captionTargetPageId="110"captionText="FIGURE 74. Coelophora saucia (Mulsant), life stages: a, b. larva; c–h. adult, variants."figureDoi="http://doi.org/10.5281/zenodo.8261743"httpUri="https://zenodo.org/record/8261743/files/figure.png"pageId="107"pageNumber="108">74</figureCitation>
<bibRefCitationid="EFFF8D851433465CBC783344FE3EFACB"author="Mulsant, E."box="[304,499,1335,1360]"pageId="107"pageNumber="108"pagination="1 - 1104"refId="ref117842"refString="Mulsant, E. (1850) Species des Coleopteres Trimeres Securipalpes. Annales des Sciences Physiques et Naturelles, d'Agriculture et d'Industrie, publiees par la Societe nationale d'Agriculture, etc., de Lyon, Deuxieme Serie, 2, 1 - 1104. https: // doi. org / 10.5962 / bhl. title. 8953"type="journal article"year="1850">Mulsant, 1850: 380</bibRefCitation>
<treatmentCitationid="0ACFD6651433465CBEFC3344FBAEFACB"author="Gordon, R. D."box="[948,1123,1336,1360]"page="18"pageId="107"pageNumber="108"year="1987">
<bibRefCitationid="EFFF8D851433465CBEFC3344FBAEFACB"author="Gordon, R. D."box="[948,1123,1336,1360]"pageId="107"pageNumber="108"pagination="1 - 46"refId="ref113038"refString="Gordon, R. D. (1987) A catalogue of the Crotch collection of Coccinellidae (Coleoptera). Occasional Papers on Systematic Entomology, 3, 1 - 46."type="journal article"year="1987">Gordon 1987: 18</bibRefCitation>
</treatmentCitation>
;
<treatmentCitationid="0ACFD6651433465CB9253344FAF4FACB"author="Ren, S. X. & Wang, X. M. & Pang, H. & Peng, Z. Q. & Zeng, T."box="[1133,1337,1336,1360]"page="208"pageId="107"pageNumber="108"year="2009">
<bibRefCitationid="EFFF8D851433465CB9253344FAF4FACB"author="Ren, S. X. & Wang, X. M. & Pang, H. & Peng, Z. Q. & Zeng, T."box="[1133,1337,1336,1360]"pageId="107"pageNumber="108"refId="ref120760"refString="Ren, S. X., Wang, X. M., Pang, H., Peng, Z. Q. & Zeng, T. (2009) Colored pictorial handbook of ladybird beetles in China. Science Press, Beijing, 336 pp. [in Chinese]"type="book"year="2009">
<bibRefCitationid="EFFF8D851433465CB8093344FA50FACB"author="Yu, G."box="[1345,1437,1336,1360]"pageId="107"pageNumber="108"refId="ref124811"refString="Yu, G. (2010) Chinese Lady Beetles (The Subfamily Coccinellinae). China Scientific Press, Beijing, 180 pp. [in Chinese]"type="book"year="2010">Yu 2010:</bibRefCitation>
<bibRefCitationid="EFFF8D851433465CBC2E3309FDFDFA16"author="Poorani, J."box="[358,560,1397,1421]"pageId="107"pageNumber="108"pagination="307 - 383"refId="ref118931"refString="Poorani, J. (2002 a) An annotated checklist of the Coccinellidae (excluding Epilachninae) of the Indian subregion. Oriental Insects, 36, 307 - 383. https: // doi. org / 10.1080 / 00305316.2002.10417335"type="journal article"year="2002">Poorani 2002a: 329</bibRefCitation>
<bibRefCitationid="EFFF8D851433465CBEBF33E8FA8DFA37"author="Iablokoff-Khnzorian, S. M."box="[1015,1344,1428,1452]"pageId="107"pageNumber="108"refId="ref114042"refString="Iablokoff-Khnzorian, S. M. (1982) Les Coccinelles Coleopteres-Coccinellidae. Tribu Coccinellini des regions Palearctique et Orientale. Boubee, Paris, 568 pp. [in French]"type="book"year="1982">Iablokoff-Khnzorian 1982: 249</bibRefCitation>
;
<bibRefCitationid="EFFF8D851433465CB80733E8FA51FA37"author="Gordon, R. D."box="[1359,1436,1428,1452]"pageId="107"pageNumber="108"pagination="1 - 46"refId="ref113038"refString="Gordon, R. D. (1987) A catalogue of the Crotch collection of Coccinellidae (Coleoptera). Occasional Papers on Systematic Entomology, 3, 1 - 46."type="journal article"year="1987">Gordon</bibRefCitation>
<bibRefCitationid="EFFF8D851433465CBC1A33ADFDD1FA72"author="Mulsant, E."box="[338,540,1489,1513]"pageId="107"pageNumber="108"pagination="1 - 1104"refId="ref117842"refString="Mulsant, E. (1850) Species des Coleopteres Trimeres Securipalpes. Annales des Sciences Physiques et Naturelles, d'Agriculture et d'Industrie, publiees par la Societe nationale d'Agriculture, etc., de Lyon, Deuxieme Serie, 2, 1 - 1104. https: // doi. org / 10.5962 / bhl. title. 8953"type="journal article"year="1850">Mulsant, 1850: 381</bibRefCitation>
<bibRefCitationid="EFFF8D851433465CBCF0338CFD97F99C"author="Pope, R. D."box="[440,602,1519,1544]"pageId="107"pageNumber="108"pagination="633 - 735"refId="ref118886"refString="Pope, R. D. (1989) A revision of the Australian Coccinellidae (Coleoptera). Part I. Subfamily Coccinellinae. Invertebrate Taxonomy, 3, 633 - 735. https: // doi. org / 10.1071 / IT 9880633"type="journal article"year="1989">Pope 1989: 359</bibRefCitation>
<figureCitationid="1355ECF11433465CBECB304AFC2EF9CB"box="[899,995,1590,1616]"captionStart="FIGURE 73"captionStartId="109.[151,250,1943,1967]"captionTargetBox="[170,1418,187,1916]"captionTargetId="figure-15@109.[160,1427,181,1920]"captionTargetPageId="109"captionText="FIGURE 73. Coelophora saucia (Mulsant): a. adult, dorsal view; b. abdomen, male; c. terminal abdominal ventrites, female; d–g. male genitalia: d. tegmen, lateral view; e. tegmen, ventral view; f. penis; g. penis apex; h. spermatheca."figureDoi="http://doi.org/10.5281/zenodo.8261741"httpUri="https://zenodo.org/record/8261741/files/figure.png"pageId="107"pageNumber="108">Fig. 73a</figureCitation>
) circular, dorsum strongly convex and glabrous. Head black. Pronotum black except lateral one-third areas white in live specimens and yellowish in dead specimens, occasionally medially paler with only a posterior transverse black macula. Elytra shiny black, with two large, transverse blood red maculae in middle (
<figureCitationid="1355ECF11433465CBFF230DEFCF6F927"box="[698,827,1698,1724]"captionStart="FIGURE 74"captionStartId="110.[151,250,1943,1967]"captionTargetBox="[233,1354,181,1919]"captionTargetId="figure-20@110.[233,1354,181,1919]"captionTargetPageId="110"captionText="FIGURE 74. Coelophora saucia (Mulsant), life stages: a, b. larva; c–h. adult, variants."figureDoi="http://doi.org/10.5281/zenodo.8261743"httpUri="https://zenodo.org/record/8261743/files/figure.png"pageId="107"pageNumber="108">Fig. 74c, d</figureCitation>
), sometimes elytra only partially black with anterior and anterolateral areas reddish, with discal reddish spots reduced in size (
<figureCitationid="1355ECF11433465CBE9230BAFB9FF97B"box="[986,1106,1734,1760]"captionStart="FIGURE 74"captionStartId="110.[151,250,1943,1967]"captionTargetBox="[233,1354,181,1919]"captionTargetId="figure-20@110.[233,1354,181,1919]"captionTargetPageId="110"captionText="FIGURE 74. Coelophora saucia (Mulsant), life stages: a, b. larva; c–h. adult, variants."figureDoi="http://doi.org/10.5281/zenodo.8261743"httpUri="https://zenodo.org/record/8261743/files/figure.png"pageId="107"pageNumber="108">Fig. 74e, f</figureCitation>
) or almost completely black. Abdominal postcoxal line (
<figureCitationid="1355ECF11433465CBC843096FDE3F89F"box="[460,558,1770,1796]"captionStart="FIGURE 73"captionStartId="109.[151,250,1943,1967]"captionTargetBox="[170,1418,187,1916]"captionTargetId="figure-15@109.[160,1427,181,1920]"captionTargetPageId="109"captionText="FIGURE 73. Coelophora saucia (Mulsant): a. adult, dorsal view; b. abdomen, male; c. terminal abdominal ventrites, female; d–g. male genitalia: d. tegmen, lateral view; e. tegmen, ventral view; f. penis; g. penis apex; h. spermatheca."figureDoi="http://doi.org/10.5281/zenodo.8261741"httpUri="https://zenodo.org/record/8261741/files/figure.png"pageId="107"pageNumber="108">Fig. 73b</figureCitation>
) incomplete; ventrite 5 posteriorly broadly emarginate and ventrite 6 slightly more deeply but narrowly emarginate in male (
<figureCitationid="1355ECF11433465CBFFC3172FCDAF8B3"box="[692,791,1806,1832]"captionStart="FIGURE 73"captionStartId="109.[151,250,1943,1967]"captionTargetBox="[170,1418,187,1916]"captionTargetId="figure-15@109.[160,1427,181,1920]"captionTargetPageId="109"captionText="FIGURE 73. Coelophora saucia (Mulsant): a. adult, dorsal view; b. abdomen, male; c. terminal abdominal ventrites, female; d–g. male genitalia: d. tegmen, lateral view; e. tegmen, ventral view; f. penis; g. penis apex; h. spermatheca."figureDoi="http://doi.org/10.5281/zenodo.8261741"httpUri="https://zenodo.org/record/8261741/files/figure.png"pageId="107"pageNumber="108">Fig. 73b</figureCitation>
), medially produced and arcuate, respectively, in female (
<figureCitationid="1355ECF11433465CBDD7314EFECDF8D7"box="[159,256,1842,1868]"captionStart="FIGURE 73"captionStartId="109.[151,250,1943,1967]"captionTargetBox="[170,1418,187,1916]"captionTargetId="figure-15@109.[160,1427,181,1920]"captionTargetPageId="109"captionText="FIGURE 73. Coelophora saucia (Mulsant): a. adult, dorsal view; b. abdomen, male; c. terminal abdominal ventrites, female; d–g. male genitalia: d. tegmen, lateral view; e. tegmen, ventral view; f. penis; g. penis apex; h. spermatheca."figureDoi="http://doi.org/10.5281/zenodo.8261741"httpUri="https://zenodo.org/record/8261741/files/figure.png"pageId="107"pageNumber="108">Fig. 73c</figureCitation>
). Male genitalia (
<figureCitationid="1355ECF11433465CBC8C314EFD8DF8D7"box="[452,576,1842,1868]"captionStart="FIGURE 73"captionStartId="109.[151,250,1943,1967]"captionTargetBox="[170,1418,187,1916]"captionTargetId="figure-15@109.[160,1427,181,1920]"captionTargetPageId="109"captionText="FIGURE 73. Coelophora saucia (Mulsant): a. adult, dorsal view; b. abdomen, male; c. terminal abdominal ventrites, female; d–g. male genitalia: d. tegmen, lateral view; e. tegmen, ventral view; f. penis; g. penis apex; h. spermatheca."figureDoi="http://doi.org/10.5281/zenodo.8261741"httpUri="https://zenodo.org/record/8261741/files/figure.png"pageId="107"pageNumber="108">Fig. 73d–g</figureCitation>
) and spermatheca (
<figureCitationid="1355ECF11433465CBE51314EFCB4F8D7"box="[793,889,1842,1868]"captionStart="FIGURE 73"captionStartId="109.[151,250,1943,1967]"captionTargetBox="[170,1418,187,1916]"captionTargetId="figure-15@109.[160,1427,181,1920]"captionTargetPageId="109"captionText="FIGURE 73. Coelophora saucia (Mulsant): a. adult, dorsal view; b. abdomen, male; c. terminal abdominal ventrites, female; d–g. male genitalia: d. tegmen, lateral view; e. tegmen, ventral view; f. penis; g. penis apex; h. spermatheca."figureDoi="http://doi.org/10.5281/zenodo.8261741"httpUri="https://zenodo.org/record/8261741/files/figure.png"pageId="107"pageNumber="108">Fig. 73h</figureCitation>
: a. Type, dorsal view (BMNH); b, d. adult, dorsal view; c. adult, lateral view; e. adult, frontal view; f, g. female genitalia; h. spermatheca (enlarged).
: a. adult, dorsal view; b. abdomen, male; c. terminal abdominal ventrites, female; d–g. male genitalia: d. tegmen, lateral view; e. tegmen, ventral view; f. penis; g. penis apex; h. spermatheca.
, as it is more commonly distributed, particularly in northeastern states. Illustrations of the habitus, life stages and the genitalia are given by
<bibRefCitationid="EFFF8D8514374658BCD8343BFD83FDFA"author="Ren, S. X. & Wang, X. M. & Pang, H. & Peng, Z. Q. & Zeng, T."box="[400,590,583,609]"pageId="111"pageNumber="112"refId="ref120760"refString="Ren, S. X., Wang, X. M., Pang, H., Peng, Z. Q. & Zeng, T. (2009) Colored pictorial handbook of ladybird beetles in China. Science Press, Beijing, 336 pp. [in Chinese]"type="book"year="2009">
<bibRefCitationid="EFFF8D8514374658BFCB343BFD3AFDFA"author="Yu, G."box="[643,759,583,609]"pageId="111"pageNumber="112"refId="ref124811"refString="Yu, G. (2010) Chinese Lady Beetles (The Subfamily Coccinellinae). China Scientific Press, Beijing, 180 pp. [in Chinese]"type="book"year="2010">Yu (2010)</bibRefCitation>