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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.912.47719" ID-GBIF-Dataset="26396469-500e-42b8-acba-f6a22f652923" ID-PMC="PMC7040049" ID-Pensoft-Pub="1313-2970-912-65" ID-Pensoft-UUID="D1FDF644344C5B5B83E6024904E40A05" ID-PubMed="32123500" ID-ZooBank="4F361E38B2684B70944DD6B6FCC66542" ModsDocID="1313-2970-912-65" checkinTime="1581958640118" checkinUser="pensoft" docAuthor="Lee, Chi-Feng &amp; Bezdek, Jan" docDate="2020" docId="B7F772033BB957AEB10DE4D09DFB7BB6" docLanguage="en" docName="ZooKeys 912: 65-124" docOrigin="ZooKeys 912" docSource="http://dx.doi.org/10.3897/zookeys.912.47719" docTitle="Pseudotheopea clypealis Lee &amp; Bezděk 2020, comb. nov." docType="treatment" docVersion="4" id="D1FDF644344C5B5B83E6024904E40A05" lastPageNumber="65" masterDocId="D1FDF644344C5B5B83E6024904E40A05" masterDocTitle="Revision of the Theopea genus group (Coleoptera, Chrysomelidae, Galerucinae), part III: Descriptions of two new genera and nine new species" masterLastPageNumber="124" masterPageNumber="65" pageNumber="65" updateTime="1668168482312" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the Theopea genus group (Coleoptera, Chrysomelidae, Galerucinae), part III: Descriptions of two new genera and nine new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lee, Chi-Feng</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bezdek, Jan</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>912</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>65</mods:start>
<mods:end>124</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.912.47719</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.912.47719</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-912-65</mods:identifier>
<mods:identifier type="ZooBank">4F361E38B2684B70944DD6B6FCC66542</mods:identifier>
<mods:identifier type="Pensoft-UUID">D1FDF644344C5B5B83E6024904E40A05</mods:identifier>
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<treatment ID-GBIF-Taxon="162147248" LSID="urn:lsid:plazi:treatment:B7F772033BB957AEB10DE4D09DFB7BB6" httpUri="http://treatment.plazi.org/id/B7F772033BB957AEB10DE4D09DFB7BB6" lastPageNumber="65" pageId="0" pageNumber="65">
<subSubSection pageId="0" pageNumber="65" type="nomenclature">
<paragraph pageId="0" pageNumber="65">
<taxonomicName LSID="B7F77203-3BB9-57AE-B10D-E4D09DFB7BB6" authority="(Medvedev, 2015)" authorityName="Lee &amp; Bezděk" authorityYear="2020" baseAuthorityName="Medvedev" baseAuthorityYear="2015" class="Insecta" family="Chrysomelidae" genus="Pseudotheopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudotheopea clypealis" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="clypealis" status="comb. nov.">Pseudotheopea clypealis (Medvedev, 2015)</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="65">comb. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of Pseudotheopea boreri sp. nov. and P. clypealis. A P. boreri sp. nov., male, dorsal view B Same, ventral view C P. boreri, female, dorsal view D P. clypealis, male, dorsal view E Same, ventral view F P. clypealis, female, dorsal view." figureDoi="10.3897/zookeys.912.47719.figure8" httpUri="https://binary.pensoft.net/fig/380663" pageId="0" pageNumber="65">Figs 8D-F</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Heads of males of Pseudotheopea boreri sp. nov. and P. clypealis. A P. boreri sp. nov., dorsofrontal view B Same, front view C P. clypealis, dorsofrontal view D Same, front view." figureDoi="10.3897/zookeys.912.47719.figure9" httpUri="https://binary.pensoft.net/fig/380664" pageId="0" pageNumber="65">; 9C, D</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Diagnostic characters of Pseudotheopea clypealis. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure11" httpUri="https://binary.pensoft.net/fig/380666" pageId="0" pageNumber="65">; 11</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="reference_group">
<paragraph pageId="0" pageNumber="65">
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Theopea clypealis" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="clypealis">Theopea clypealis</taxonomicName>
Medvedev, 2015: 72 (South Vietnam)
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="materials_examined">
<paragraph pageId="0" pageNumber="65">Type.</paragraph>
<paragraph pageId="0" pageNumber="65">Holotype ♂ (LMCM, based on photographs): &quot;Vietnam Dongnai Pr. / Cat Tien V.99 / leg. A. Polilov [p, w] // HOLOTYPUS / Theopea / clypealis / L. Medvedev [p, r]&quot;.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="materials_examined">
<paragraph pageId="0" pageNumber="65">Other specimens examined.</paragraph>
<paragraph pageId="0" pageNumber="65">VIETNAM. Kien Giang: 2♂♂, 3♀♀ (TARI), Phu Quoc island, 12-14.IV.2013, leg. Y.-T. Wang; 2♂♂, 1♀ (NMNS), same island, Ding Ba Rd. + Banh Dan Rd., 14.IV.2013, leg. M.-L. Jeng.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="redescription">
<paragraph pageId="0" pageNumber="65">Redescription.</paragraph>
<paragraph pageId="0" pageNumber="65">
Length 5.9-6.8 mm, width 2.3-2.6 mm. Body color (Figs
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of Pseudotheopea boreri sp. nov. and P. clypealis. A P. boreri sp. nov., male, dorsal view B Same, ventral view C P. boreri, female, dorsal view D P. clypealis, male, dorsal view E Same, ventral view F P. clypealis, female, dorsal view." figureDoi="10.3897/zookeys.912.47719.figure8" httpUri="https://binary.pensoft.net/fig/380663" pageId="0" pageNumber="65">8D-F</figureCitation>
,
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Diagnostic characters of Pseudotheopea clypealis. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure11" httpUri="https://binary.pensoft.net/fig/380666" pageId="0" pageNumber="65">11</figureCitation>
) golden green, but antennae, mouth parts, and legs yellowish brown, two or three apical antennomeres darker. Frontoclypeus (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Heads of males of Pseudotheopea boreri sp. nov. and P. clypealis. A P. boreri sp. nov., dorsofrontal view B Same, front view C P. clypealis, dorsofrontal view D Same, front view." figureDoi="10.3897/zookeys.912.47719.figure9" httpUri="https://binary.pensoft.net/fig/380664" pageId="0" pageNumber="65">9C, D</figureCitation>
) transverse and deeply excavated between eyes in males, concavity 0.5
<normalizedToken originalValue="×">x</normalizedToken>
as wide as interspace between eyes, anteriorly narrowed, margined with long hair-like setae and with one erect process at center, margined with hair-like setae; baso-lateral angles covered by rounded membranous sclerites. Antennae filiform in males, (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Diagnostic characters of Pseudotheopea clypealis. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure11" httpUri="https://binary.pensoft.net/fig/380666" pageId="0" pageNumber="65">11A</figureCitation>
), antennomeres III-IX slightly curved, length ratios of antennomeres I-XI 1.0: 0.3: 0.7: 0.8: 0.8: 0.8: 0.8: 0.7: 0.7: 0.6: 0.7, length to width ratios of antennomeres I-XI 3.6: 2.0: 3.2: 3.8: 3.8: 3.8: 4.2: 4.3: 3.9: 4.4: 5.3; filiform in females (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Diagnostic characters of Pseudotheopea clypealis. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure11" httpUri="https://binary.pensoft.net/fig/380666" pageId="0" pageNumber="65">11B</figureCitation>
), similar to males, length ratios of antennomeres I-XI 1.0: 0.3: 0.6: 0.8: 0.8: 0.7: 0.7: 0.7: 0.6: 0.6: 0.7, length to width ratios of antennomeres I-XI 4.0: 2.0: 3.3: 4.8: 4.9: 4.4: 4.8: 4.7: 4.3: 3.9: 4.5. Elytra elongate, parallel-sided, 1.8
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; disc with dense, coarse punctures, arranged into longitudinal rows, with one longitudinal ridge between two longitudinal rows of punctures with convex, with distinct and indistinct ridges intertwined. Tarsomeres I of front legs swollen in males; subparallel in females. Aedeagus (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Diagnostic characters of Pseudotheopea clypealis. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure11" httpUri="https://binary.pensoft.net/fig/380666" pageId="0" pageNumber="65">11C-E</figureCitation>
) extremely slender, 10.0
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; apex with shallow notch, both apices equal in length; tectum elongate, from apical 1/10 to basal 2/5; almost straight but moderately curved at basal 1/5 in lateral view, apically curved, angular at apical 1/3; triangular sclerites small; internal sac with elongate, endophallic sclerite complex, 0.6
<normalizedToken originalValue="×">x</normalizedToken>
as long as aedeagus, composed of two sclerites, apical piece (4.0
<normalizedToken originalValue="×">x</normalizedToken>
) much longer than basal piece, one dorsal sclerite slender, 0.3
<normalizedToken originalValue="×">x</normalizedToken>
as long as endophallic sclerite; ventral sclerites absent but one additional pair of hook-like sclerites present. Gonocoxae (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Diagnostic characters of Pseudotheopea clypealis. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure11" httpUri="https://binary.pensoft.net/fig/380666" pageId="0" pageNumber="65">11G</figureCitation>
) elongate, both gonocoxae fused from basal 1/3 to apical 1/4; apices convergent and narrowly rounded, each gonocoxa with seven setae along lateral margin from apex to apical 1/6; with one pair of short lateral processes at basal 2/5. Ventrite VIII (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Diagnostic characters of Pseudotheopea clypealis. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure11" httpUri="https://binary.pensoft.net/fig/380666" pageId="0" pageNumber="65">11F</figureCitation>
) elongate and well sclerotized; disc with several long setae at sides and near apical margin, and with dense, short setae along apical margin; spiculum extremely slender. Receptacle of spermatheca (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Diagnostic characters of Pseudotheopea clypealis. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure11" httpUri="https://binary.pensoft.net/fig/380666" pageId="0" pageNumber="65">11H</figureCitation>
) tightly joined with pump, pump slender and strongly curved; proximal spermathecal duct deeply inserted into receptacle, narrow and short.
</paragraph>
<caption doi="10.3897/zookeys.912.47719.figure11" httpUri="https://binary.pensoft.net/fig/380666" pageId="0" pageNumber="65" start="Figure 11" startId="F11">
<paragraph pageId="0" pageNumber="65">
<emphasis bold="true" pageId="0" pageNumber="65">Figure 11.</emphasis>
Diagnostic characters of
<taxonomicName authorityName="Lee &amp; Bezděk" authorityYear="2020" baseAuthorityName="Medvedev" baseAuthorityYear="2015" class="Insecta" family="Chrysomelidae" genus="Pseudotheopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudotheopea clypealis" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="clypealis">
<emphasis italics="true" pageId="0" pageNumber="65">Pseudotheopea clypealis</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="0" pageNumber="65">A</emphasis>
Antenna, male
<emphasis bold="true" pageId="0" pageNumber="65">B</emphasis>
Antenna, female
<emphasis bold="true" pageId="0" pageNumber="65">C</emphasis>
Aedeagus, dorsal view
<emphasis bold="true" pageId="0" pageNumber="65">D</emphasis>
Aedeagus, lateral view
<emphasis bold="true" pageId="0" pageNumber="65">E</emphasis>
Aedeagus, ventral view
<emphasis bold="true" pageId="0" pageNumber="65">F</emphasis>
Abdominal ventrite VIII
<emphasis bold="true" pageId="0" pageNumber="65">G</emphasis>
Gonocoxae
<emphasis bold="true" pageId="0" pageNumber="65">H</emphasis>
Spermatheca.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="diagnosis">
<paragraph pageId="0" pageNumber="65">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="65">
<taxonomicName authorityName="Lee &amp; Bezděk" authorityYear="2020" baseAuthorityName="Medvedev" baseAuthorityYear="2015" class="Insecta" family="Chrysomelidae" genus="Pseudotheopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudotheopea clypealis" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="clypealis">
<emphasis italics="true" pageId="0" pageNumber="65">Pseudotheopea clypealis</emphasis>
</taxonomicName>
(Medvedev) (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of Pseudotheopea boreri sp. nov. and P. clypealis. A P. boreri sp. nov., male, dorsal view B Same, ventral view C P. boreri, female, dorsal view D P. clypealis, male, dorsal view E Same, ventral view F P. clypealis, female, dorsal view." figureDoi="10.3897/zookeys.912.47719.figure8" httpUri="https://binary.pensoft.net/fig/380663" pageId="0" pageNumber="65">8D-F</figureCitation>
),
<taxonomicName lsidName="P. boreri" pageId="0" pageNumber="65" rank="species" species="boreri">
<emphasis italics="true" pageId="0" pageNumber="65">P. boreri</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of Pseudotheopea boreri sp. nov. and P. clypealis. A P. boreri sp. nov., male, dorsal view B Same, ventral view C P. boreri, female, dorsal view D P. clypealis, male, dorsal view E Same, ventral view F P. clypealis, female, dorsal view." figureDoi="10.3897/zookeys.912.47719.figure8" httpUri="https://binary.pensoft.net/fig/380663" pageId="0" pageNumber="65">8A-C</figureCitation>
),
<taxonomicName lsidName="P. hsingtzungi" pageId="0" pageNumber="65" rank="species" species="hsingtzungi">
<emphasis italics="true" pageId="0" pageNumber="65">P. hsingtzungi</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Habitus of Pseudotheopea hsingtzungi sp. nov. and P. smaragdina. A P. hsingtzungi sp. nov., male, dorsal view B P. hsingtzungi sp. nov., female, dorsal view C Same, ventral view D P. smaragdina, male, dorsal view E P. smaragdina, female, dorsal view F Same, ventral view." figureDoi="10.3897/zookeys.912.47719.figure15" httpUri="https://binary.pensoft.net/fig/380670" pageId="0" pageNumber="65">15A-C</figureCitation>
), and
<taxonomicName lsidName="P. smaragdina" pageId="0" pageNumber="65" rank="species" species="smaragdina">
<emphasis italics="true" pageId="0" pageNumber="65">P. smaragdina</emphasis>
</taxonomicName>
(Gressitt and Kimoto) (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Habitus of Pseudotheopea hsingtzungi sp. nov. and P. smaragdina. A P. hsingtzungi sp. nov., male, dorsal view B P. hsingtzungi sp. nov., female, dorsal view C Same, ventral view D P. smaragdina, male, dorsal view E P. smaragdina, female, dorsal view F Same, ventral view." figureDoi="10.3897/zookeys.912.47719.figure15" httpUri="https://binary.pensoft.net/fig/380670" pageId="0" pageNumber="65">15D-F</figureCitation>
), are characterized by their golden green coloration. They can be identified based on their distribution:
<taxonomicName lsidName="P. boreri" pageId="0" pageNumber="65" rank="species" species="boreri">
<emphasis italics="true" pageId="0" pageNumber="65">P. boreri</emphasis>
</taxonomicName>
sp. nov. from India,
<taxonomicName lsidName="P. clypealis" pageId="0" pageNumber="65" rank="species" species="clypealis">
<emphasis italics="true" pageId="0" pageNumber="65">P. clypealis</emphasis>
</taxonomicName>
from Vietnam,
<taxonomicName lsidName="P. hsingtzungi" pageId="0" pageNumber="65" rank="species" species="hsingtzungi">
<emphasis italics="true" pageId="0" pageNumber="65">P. hsingtzungi</emphasis>
</taxonomicName>
sp. nov. from Laos, and
<taxonomicName lsidName="P. smaragdina" pageId="0" pageNumber="65" rank="species" species="smaragdina">
<emphasis italics="true" pageId="0" pageNumber="65">P. smaragdina</emphasis>
</taxonomicName>
from China.
<taxonomicName authorityName="Lee &amp; Bezděk" authorityYear="2020" baseAuthorityName="Medvedev" baseAuthorityYear="2015" class="Insecta" family="Chrysomelidae" genus="Pseudotheopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudotheopea clypealis" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="clypealis">
<emphasis italics="true" pageId="0" pageNumber="65">Pseudotheopea clypealis</emphasis>
</taxonomicName>
can be separated from the others by the convex and distinct longitudinal ridges on the elytra (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of Pseudotheopea boreri sp. nov. and P. clypealis. A P. boreri sp. nov., male, dorsal view B Same, ventral view C P. boreri, female, dorsal view D P. clypealis, male, dorsal view E Same, ventral view F P. clypealis, female, dorsal view." figureDoi="10.3897/zookeys.912.47719.figure8" httpUri="https://binary.pensoft.net/fig/380663" pageId="0" pageNumber="65">8D, F</figureCitation>
) (indistinct longitudinal ridges on the elytra in others (Figs
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of Pseudotheopea boreri sp. nov. and P. clypealis. A P. boreri sp. nov., male, dorsal view B Same, ventral view C P. boreri, female, dorsal view D P. clypealis, male, dorsal view E Same, ventral view F P. clypealis, female, dorsal view." figureDoi="10.3897/zookeys.912.47719.figure8" httpUri="https://binary.pensoft.net/fig/380663" pageId="0" pageNumber="65">8A, C</figureCitation>
;
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Habitus of Pseudotheopea hsingtzungi sp. nov. and P. smaragdina. A P. hsingtzungi sp. nov., male, dorsal view B P. hsingtzungi sp. nov., female, dorsal view C Same, ventral view D P. smaragdina, male, dorsal view E P. smaragdina, female, dorsal view F Same, ventral view." figureDoi="10.3897/zookeys.912.47719.figure15" httpUri="https://binary.pensoft.net/fig/380670" pageId="0" pageNumber="65">15A, B, D, E</figureCitation>
), and the narrower anterior concavity between eyes (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Heads of males of Pseudotheopea boreri sp. nov. and P. clypealis. A P. boreri sp. nov., dorsofrontal view B Same, front view C P. clypealis, dorsofrontal view D Same, front view." figureDoi="10.3897/zookeys.912.47719.figure9" httpUri="https://binary.pensoft.net/fig/380664" pageId="0" pageNumber="65">9C, D</figureCitation>
) (broadly rounded anterior margin of concavity between eyes in others (Figs
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Heads of males of Pseudotheopea boreri sp. nov. and P. clypealis. A P. boreri sp. nov., dorsofrontal view B Same, front view C P. clypealis, dorsofrontal view D Same, front view." figureDoi="10.3897/zookeys.912.47719.figure9" httpUri="https://binary.pensoft.net/fig/380664" pageId="0" pageNumber="65">9A, B</figureCitation>
;
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Heads of males of Pseudotheopea hsingtzungi sp. nov. and P. smaragdina. A P. hsingtzungi sp. nov., dorsofrontal view B Same, front view C P. smaragdina, dorsofrontal view D Same, front view." figureDoi="10.3897/zookeys.912.47719.figure16" httpUri="https://binary.pensoft.net/fig/380671" pageId="0" pageNumber="65">16A-D</figureCitation>
). Males of
<taxonomicName lsidName="P. clypealis" pageId="0" pageNumber="65" rank="species" species="clypealis">
<emphasis italics="true" pageId="0" pageNumber="65">P. clypealis</emphasis>
</taxonomicName>
are similar to those of
<taxonomicName lsidName="P. smaragdina" pageId="0" pageNumber="65" rank="species" species="smaragdina">
<emphasis italics="true" pageId="0" pageNumber="65">P. smaragdina</emphasis>
</taxonomicName>
in possessing one additional elongate dorsal sclerite and one pair of small lateral hook-like sclerites inside the internal sac (Figs
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Diagnostic characters of Pseudotheopea clypealis. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure11" httpUri="https://binary.pensoft.net/fig/380666" pageId="0" pageNumber="65">11C, D</figureCitation>
;
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Diagnostic characters of Pseudotheopea smaragdina. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure22" httpUri="https://binary.pensoft.net/fig/380677" pageId="0" pageNumber="65">22D, E</figureCitation>
). They differ in possessing symmetrical aedeagal apices and a relatively longer apical piece (4.0
<normalizedToken originalValue="×">x</normalizedToken>
longer than basal piece, Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Diagnostic characters of Pseudotheopea clypealis. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure11" httpUri="https://binary.pensoft.net/fig/380666" pageId="0" pageNumber="65">11C, D</figureCitation>
) (asymmetrical apices and relatively shorter apical piece, as long as basal piece in
<taxonomicName lsidName="P. smaragdina" pageId="0" pageNumber="65" rank="species" species="smaragdina">
<emphasis italics="true" pageId="0" pageNumber="65">P. smaragdina</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Diagnostic characters of Pseudotheopea smaragdina. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure22" httpUri="https://binary.pensoft.net/fig/380677" pageId="0" pageNumber="65">22D, E</figureCitation>
)).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="distribution">
<paragraph pageId="0" pageNumber="65">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="65">Vietnam.</paragraph>
</subSubSection>
</treatment>
</document>
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