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<document id="A8B1C4DAA7ACE4C30D21E63AFF383009" ID-DOI="10.5070/P9371050859" ID-ISSN="0031-0298" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726019095505" checkinUser="felipe" docAuthor="Marsh, Adam D. &amp; Parker, William G." docDate="2022" docId="038C8F41F824096A6288D2ADFB2D8F15" docLanguage="en" docName="PaleoBios.37.50859.pdf" docOrigin="PaleoBios 37" docStyle="DocumentStyle:2D14AFDE223D5FD9EB1703D6A63DEF6C.1:PaleoBios.2018-2021.journal_article" docStyleId="2D14AFDE223D5FD9EB1703D6A63DEF6C" docStyleName="PaleoBios.2018-2021.journal_article" docStyleVersion="1" docTitle="Dinosauria OWEN 1842" docType="treatment" docVersion="1" lastPageNumber="13" masterDocId="FFB5F739F82A09646206D72CFFC88C26" masterDocTitle="New dinosauromorph specimens from Petrified Forest National Park and a global biostratigraphic review of Triassic dinosauromorph body fossils" masterLastPageNumber="56" masterPageNumber="1" pageNumber="13" updateTime="1726078387551" updateUser="felipe" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:title id="3DD870008865C1251E4947C9C36AF67D">New dinosauromorph specimens from Petrified Forest National Park and a global biostratigraphic review of Triassic dinosauromorph body fossils</mods:title>
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<mods:namePart id="A9260D1F7ACB6BAFE0803947A29170EE">Marsh, Adam D.</mods:namePart>
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<mods:namePart id="7C63045C52F76C42188EE9491080C6DD">Parker, William G.</mods:namePart>
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<mods:typeOfResource id="BBE98BC208494066F805ADA137AAD4BC">text</mods:typeOfResource>
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<mods:title id="588A6565484911EBE84F32EA11786D88">PaleoBios</mods:title>
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<mods:date id="778179C022F403B4DE38E12DCA03EF18">2022</mods:date>
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<mods:number id="585F7417E64090D58BDDB5506CD63BE9">37</mods:number>
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<treatment id="038C8F41F824096A6288D2ADFB2D8F15" LSID="urn:lsid:plazi:treatment:038C8F41F824096A6288D2ADFB2D8F15" httpUri="http://treatment.plazi.org/id/038C8F41F824096A6288D2ADFB2D8F15" lastPageNumber="13" pageId="14" pageNumber="13">
<subSubSection id="C33F6DDCF824096A6288D2ADFCCA89BF" box="[142,770,1409,1433]" pageId="14" pageNumber="13" type="nomenclature">
<paragraph id="8B9A3E57F824096A6288D2ADFCCA89BF" blockId="14.[142,770,1409,1471]" box="[142,770,1409,1433]" pageId="14" pageNumber="13">
<heading id="D0D2893BF824096A6288D2ADFCCA89BF" allCaps="true" box="[142,770,1409,1433]" centered="true" fontSize="10" level="2" pageId="14" pageNumber="13" reason="2">
<taxonomicName id="4C2545D4F824096A6288D2ADFD4089BF" authority="OWEN, 1842 SENSU PADIAN AND" authorityName="OWEN" authorityYear="1842" box="[142,648,1409,1433]" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="14" pageNumber="13" phylum="Chordata" rank="order">
DINOSAURIA OWEN, 1842
<emphasis id="B951E245F824096A63BDD2ADFE3589BE" box="[443,509,1409,1432]" italics="true" pageId="14" pageNumber="13">SENSU</emphasis>
PADIAN AND
</taxonomicName>
MAY, 1993
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33F6DDCF824096A6397D28BFE368999" box="[401,510,1447,1471]" pageId="14" pageNumber="13" type="description">
<paragraph id="8B9A3E57F824096A6397D28BFE368999" blockId="14.[142,770,1409,1471]" box="[401,510,1447,1471]" pageId="14" pageNumber="13">
<figureCitation id="131E22D2F824096A6397D28BFE368999" box="[401,510,1447,1471]" captionStart="Figure 6" captionStartId="15.[112,179,1765,1787]" captionTargetBox="[229,1376,1229,1735]" captionTargetId="figure-45@15.[706,902,1480,1735]" captionTargetPageId="15" captionText="Figure 6. A–D. Proximal ends of right and left dinosaur femora, PEFO 44475 in proximal (A, C) and anterior view (B, D). E, F. Proximal end of left dinosaur femur, PEFO 34583 in proximal (E) and anterior (F) view. G, H. Proximal end of left dinosaur femur, PEFO 34863 in proximal (G) and anterior (H) view.I, J. Distal end of right saurischian femur, PEFO 21660/UCMP 126751 in distal (I) and posterior (J) view. K, L. Distal end of right saurischian femur, PEFO 33956 in distal (K) and posterior (L) view. M, N. Distal end of right saurischian femur, PEFO 44469 in distal (M) and posterior (N) view. O, P. Distal end of left saurischian femur, PEFO 34562 in distal (O) and posterior (P) view. Q, R. Distal end of left saurischian femur, PEFO 44474 in distal (Q) and posterior (R) view. S, T. Distal end of left saurischian femur, PEFO 42987/UWBM 117640 in distal (S) and posterior (T) view. Abbreviations: at, anterior trochanter; ctf, crista tibiofibularis; g, groove; lc, lateral condyle; mc, medial condyle; ve, ventral emargination. Scale bars=1 cm, arrows point in anterior direction." pageId="14" pageNumber="13">FIG. 6A–H</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33F6DDCF824096A6296D2FFFD958AB8" pageId="14" pageNumber="13" type="materials_examined">
<paragraph id="8B9A3E57F824096A6296D2FFFD958AB8" blockId="14.[112,800,1491,1939]" pageId="14" pageNumber="13">
<emphasis id="B951E245F824096A6296D2FFFDAB89CB" bold="true" box="[144,611,1491,1518]" pageId="14" pageNumber="13">Referred specimens and localities</emphasis>
—
<collectionCode id="ED34A692F824096A6079D2F8FD0C89C8" box="[639,708,1492,1518]" country="0" httpUri="http://grbio.org/cool/7x98-r3em" name="Petrified Forest" pageId="14" pageNumber="13">PEFO</collectionCode>
44475 (
<figureCitation id="131E22D2F824096A627FD2DBFF3E8A36" box="[121,246,1526,1553]" captionStart="Figure 6" captionStartId="15.[112,179,1765,1787]" captionTargetBox="[229,1376,1229,1735]" captionTargetId="figure-45@15.[706,902,1480,1735]" captionTargetPageId="15" captionText="Figure 6. A–D. Proximal ends of right and left dinosaur femora, PEFO 44475 in proximal (A, C) and anterior view (B, D). E, F. Proximal end of left dinosaur femur, PEFO 34583 in proximal (E) and anterior (F) view. G, H. Proximal end of left dinosaur femur, PEFO 34863 in proximal (G) and anterior (H) view.I, J. Distal end of right saurischian femur, PEFO 21660/UCMP 126751 in distal (I) and posterior (J) view. K, L. Distal end of right saurischian femur, PEFO 33956 in distal (K) and posterior (L) view. M, N. Distal end of right saurischian femur, PEFO 44469 in distal (M) and posterior (N) view. O, P. Distal end of left saurischian femur, PEFO 34562 in distal (O) and posterior (P) view. Q, R. Distal end of left saurischian femur, PEFO 44474 in distal (Q) and posterior (R) view. S, T. Distal end of left saurischian femur, PEFO 42987/UWBM 117640 in distal (S) and posterior (T) view. Abbreviations: at, anterior trochanter; ctf, crista tibiofibularis; g, groove; lc, lateral condyle; mc, medial condyle; ve, ventral emargination. Scale bars=1 cm, arrows point in anterior direction." pageId="14" pageNumber="13">Fig. 6A–D</figureCitation>
), proximal ends of left and right femora,
<collectionCode id="ED34A692F824096A60E8D2DBFCE88A37" box="[750,800,1527,1553]" pageId="14" pageNumber="13">PFV</collectionCode>
376: Scour Sandstone,
<collectionCode id="ED34A692F824096A6391D136FE1B8A12" box="[407,467,1562,1588]" pageId="14" pageNumber="13">PFM</collectionCode>
;
<collectionCode id="ED34A692F824096A63E4D136FDEF8A12" box="[482,551,1562,1588]" country="0" httpUri="http://grbio.org/cool/7x98-r3em" name="Petrified Forest" pageId="14" pageNumber="13">PEFO</collectionCode>
34583 (
<figureCitation id="131E22D2F824096A6092D136FCDA8A12" box="[660,786,1562,1588]" captionStart="Figure 6" captionStartId="15.[112,179,1765,1787]" captionTargetBox="[229,1376,1229,1735]" captionTargetId="figure-45@15.[706,902,1480,1735]" captionTargetPageId="15" captionText="Figure 6. A–D. Proximal ends of right and left dinosaur femora, PEFO 44475 in proximal (A, C) and anterior view (B, D). E, F. Proximal end of left dinosaur femur, PEFO 34583 in proximal (E) and anterior (F) view. G, H. Proximal end of left dinosaur femur, PEFO 34863 in proximal (G) and anterior (H) view.I, J. Distal end of right saurischian femur, PEFO 21660/UCMP 126751 in distal (I) and posterior (J) view. K, L. Distal end of right saurischian femur, PEFO 33956 in distal (K) and posterior (L) view. M, N. Distal end of right saurischian femur, PEFO 44469 in distal (M) and posterior (N) view. O, P. Distal end of left saurischian femur, PEFO 34562 in distal (O) and posterior (P) view. Q, R. Distal end of left saurischian femur, PEFO 44474 in distal (Q) and posterior (R) view. S, T. Distal end of left saurischian femur, PEFO 42987/UWBM 117640 in distal (S) and posterior (T) view. Abbreviations: at, anterior trochanter; ctf, crista tibiofibularis; g, groove; lc, lateral condyle; mc, medial condyle; ve, ventral emargination. Scale bars=1 cm, arrows point in anterior direction." pageId="14" pageNumber="13">Fig. 6E, F</figureCitation>
), proximal end of left femur,
<collectionCode id="ED34A692F824096A63C5D112FE3D8A7E" box="[451,501,1598,1624]" pageId="14" pageNumber="13">PFV</collectionCode>
332: Chinde Valley Pin- nacle,
<collectionCode id="ED34A692F824096A62BAD14DFF308A5D" box="[188,248,1633,1659]" pageId="14" pageNumber="13">PFM</collectionCode>
;
<collectionCode id="ED34A692F824096A6307D14CFE8D8A5C" box="[257,325,1632,1658]" country="0" httpUri="http://grbio.org/cool/7x98-r3em" name="Petrified Forest" pageId="14" pageNumber="13">PEFO</collectionCode>
34863 (
<figureCitation id="131E22D2F824096A63AED14CFDE88A5C" box="[424,544,1632,1658]" captionStart="Figure 6" captionStartId="15.[112,179,1765,1787]" captionTargetBox="[229,1376,1229,1735]" captionTargetId="figure-45@15.[706,902,1480,1735]" captionTargetPageId="15" captionText="Figure 6. A–D. Proximal ends of right and left dinosaur femora, PEFO 44475 in proximal (A, C) and anterior view (B, D). E, F. Proximal end of left dinosaur femur, PEFO 34583 in proximal (E) and anterior (F) view. G, H. Proximal end of left dinosaur femur, PEFO 34863 in proximal (G) and anterior (H) view.I, J. Distal end of right saurischian femur, PEFO 21660/UCMP 126751 in distal (I) and posterior (J) view. K, L. Distal end of right saurischian femur, PEFO 33956 in distal (K) and posterior (L) view. M, N. Distal end of right saurischian femur, PEFO 44469 in distal (M) and posterior (N) view. O, P. Distal end of left saurischian femur, PEFO 34562 in distal (O) and posterior (P) view. Q, R. Distal end of left saurischian femur, PEFO 44474 in distal (Q) and posterior (R) view. S, T. Distal end of left saurischian femur, PEFO 42987/UWBM 117640 in distal (S) and posterior (T) view. Abbreviations: at, anterior trochanter; ctf, crista tibiofibularis; g, groove; lc, lateral condyle; mc, medial condyle; ve, ventral emargination. Scale bars=1 cm, arrows point in anterior direction." pageId="14" pageNumber="13">Fig. 6G, H</figureCitation>
), proximal end of left femur,
<collectionCode id="ED34A692F824096A62C3D1A8FF3F8AB8" box="[197,247,1668,1694]" pageId="14" pageNumber="13">PFV</collectionCode>
215: Zuni Well Mound,
<collectionCode id="ED34A692F824096A6018D1A8FD918AB8" box="[542,601,1668,1694]" pageId="14" pageNumber="13">PFM</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F6DDCF824096A6296D18AFB2D8F15" pageId="14" pageNumber="13" type="description">
<paragraph id="8B9A3E57F824096A6296D18AFCE88BB4" blockId="14.[112,800,1491,1939]" pageId="14" pageNumber="13">
<emphasis id="B951E245F824096A6296D18AFD738AE6" bold="true" box="[144,699,1702,1728]" pageId="14" pageNumber="13">Description and rationale for assignment</emphasis>
—These proximal ends of femora are highly fragmentary and weathered; each of them is broken proximal to the fourth trochanter. Each preserves a mound-like anterior trochanter that remains attached to the shaft proximally (Nesbitt 2011:308-1) (
<figureCitation id="131E22D2F824096A638DD079FE208B49" box="[395,488,1877,1903]" captionStart="Figure 6" captionStartId="15.[112,179,1765,1787]" captionTargetBox="[229,1376,1229,1735]" captionTargetId="figure-45@15.[706,902,1480,1735]" captionTargetPageId="15" captionText="Figure 6. A–D. Proximal ends of right and left dinosaur femora, PEFO 44475 in proximal (A, C) and anterior view (B, D). E, F. Proximal end of left dinosaur femur, PEFO 34583 in proximal (E) and anterior (F) view. G, H. Proximal end of left dinosaur femur, PEFO 34863 in proximal (G) and anterior (H) view.I, J. Distal end of right saurischian femur, PEFO 21660/UCMP 126751 in distal (I) and posterior (J) view. K, L. Distal end of right saurischian femur, PEFO 33956 in distal (K) and posterior (L) view. M, N. Distal end of right saurischian femur, PEFO 44469 in distal (M) and posterior (N) view. O, P. Distal end of left saurischian femur, PEFO 34562 in distal (O) and posterior (P) view. Q, R. Distal end of left saurischian femur, PEFO 44474 in distal (Q) and posterior (R) view. S, T. Distal end of left saurischian femur, PEFO 42987/UWBM 117640 in distal (S) and posterior (T) view. Abbreviations: at, anterior trochanter; ctf, crista tibiofibularis; g, groove; lc, lateral condyle; mc, medial condyle; ve, ventral emargination. Scale bars=1 cm, arrows point in anterior direction." pageId="14" pageNumber="13">Fig. 6D</figureCitation>
), which is a feature present in Dinosauriformes, e.g.,
<taxonomicName id="4C2545D4F824096A63D0D05BFD138BB5" box="[470,731,1911,1939]" class="Reptilia" genus="Marasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="13" phylum="Chordata" rank="species" species="lilloensis">
<emphasis id="B951E245F824096A63D0D05BFD138BB5" box="[470,731,1911,1939]" italics="true" pageId="14" pageNumber="13">Marasuchus lilloensis</emphasis>
</taxonomicName>
(PVL
</paragraph>
<paragraph id="8B9A3E57F824096A6146D7EFFB2D8F15" blockId="14.[832,1521,193,820]" pageId="14" pageNumber="13">
3871, Sereno and Arcucci 1994: fig. 8b),
<taxonomicName id="4C2545D4F824096A6763D7EDFC578D24" family="Silesauridae" genus="Asilisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="13" phylum="Chordata" rank="species" species="kongwe">
<emphasis id="B951E245F824096A6763D7EDFC578D24" italics="true" pageId="14" pageNumber="13">Asilisaurus kongwe</emphasis>
</taxonomicName>
(NMT RB159, Nesbitt et al. 2019: fig. 42d), and
<taxonomicName id="4C2545D4F824096A6146D624FBF48D02" box="[832,1084,264,292]" class="Reptilia" genus="Buriolestes" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="13" phylum="Chordata" rank="species" species="schultzi">
<emphasis id="B951E245F824096A6146D624FBF48D02" box="[832,1084,264,292]" italics="true" pageId="14" pageNumber="13">Buriolestes schultzi</emphasis>
</taxonomicName>
(ULBRA-PVT280, Cabreira et al. 2016: fig. 1n). Unlike the smooth transition between the ventromedial margin of the femoral head and the shaft (Nesbitt 2011:304-0) found in early dinosauromorphs, e.g.,
<taxonomicName id="4C2545D4F824096A617FD6BAFAC98D97" authority="Romer" authorityName="Romer" authorityYear="1971" box="[889,1281,405,434]" class="Reptilia" family="Lagerpetonidae" genus="Lagerpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="13" phylum="Chordata" rank="species" species="chanarensis">
<emphasis id="B951E245F824096A617FD6BAFB6B8D97" box="[889,1187,405,434]" italics="true" pageId="14" pageNumber="13">Lagerpeton chanarensis</emphasis>
Romer
</taxonomicName>
(1971, PVL 4619) (Nesbitt et al. 2009b: fig. 3a), and
<taxonomicName id="4C2545D4F824096A66F4D694FA388DF2" box="[1266,1520,440,468]" class="Reptilia" family="Lagerpetonidae" genus="Dromomeron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="13" phylum="Chordata" rank="species" species="romeri">
<emphasis id="B951E245F824096A66F4D694FA388DF2" box="[1266,1520,440,468]" italics="true" pageId="14" pageNumber="13">Dromomeron romeri</emphasis>
</taxonomicName>
(GR 218, Nesbitt et al. 2009b: fig. 1a) and the ventral notch (Nesbitt 2011:304-1) present on silesaurid femora (see above), these femora exhibit a concave emargination ventral to the femoral head (Nesbitt 2011:304-2) (
<figureCitation id="131E22D2F824096A67C7D56AFCAF8EA5" captionStart="Figure 6" captionStartId="15.[112,179,1765,1787]" captionTargetBox="[229,1376,1229,1735]" captionTargetId="figure-45@15.[706,902,1480,1735]" captionTargetPageId="15" captionText="Figure 6. A–D. Proximal ends of right and left dinosaur femora, PEFO 44475 in proximal (A, C) and anterior view (B, D). E, F. Proximal end of left dinosaur femur, PEFO 34583 in proximal (E) and anterior (F) view. G, H. Proximal end of left dinosaur femur, PEFO 34863 in proximal (G) and anterior (H) view.I, J. Distal end of right saurischian femur, PEFO 21660/UCMP 126751 in distal (I) and posterior (J) view. K, L. Distal end of right saurischian femur, PEFO 33956 in distal (K) and posterior (L) view. M, N. Distal end of right saurischian femur, PEFO 44469 in distal (M) and posterior (N) view. O, P. Distal end of left saurischian femur, PEFO 34562 in distal (O) and posterior (P) view. Q, R. Distal end of left saurischian femur, PEFO 44474 in distal (Q) and posterior (R) view. S, T. Distal end of left saurischian femur, PEFO 42987/UWBM 117640 in distal (S) and posterior (T) view. Abbreviations: at, anterior trochanter; ctf, crista tibiofibularis; g, groove; lc, lateral condyle; mc, medial condyle; ve, ventral emargination. Scale bars=1 cm, arrows point in anterior direction." pageId="14" pageNumber="13">Fig. 6H</figureCitation>
), which is apomorphic for dinosaurs, e.g.,
<taxonomicName id="4C2545D4F824096A6784D544FC5C8E81" class="Reptilia" genus="Eocursor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="13" phylum="Chordata" rank="species" species="parvus">
<emphasis id="B951E245F824096A6784D544FC5C8E81" italics="true" pageId="14" pageNumber="13">Eocursor parvus</emphasis>
</taxonomicName>
(SAM-PK-K8025, Butler 2009: fig. 15d),
<taxonomicName id="4C2545D4F824096A6798D5A7FB668EEC" authority="Martinez et al." authorityName="Martinez" class="Reptilia" genus="Eodromaeus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="13" phylum="Chordata" rank="species" species="murphi">
<emphasis id="B951E245F824096A6798D5A7FC398EEC" italics="true" pageId="14" pageNumber="13">Eodromaeus murphi</emphasis>
Martínez et al.
</taxonomicName>
(2011, PVSJ 562: fig. 2k), and
<taxonomicName id="4C2545D4F824096A617FD5FDFAC08EC9" authorityName="Reig" authorityYear="1963" box="[889,1288,721,751]" class="Reptilia" family="Herrerasauridae" genus="Herrerasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="14" pageNumber="13" phylum="Chordata" rank="species" species="ischigualastensis">
<emphasis id="B951E245F824096A617FD5FDFAC08EC9" box="[889,1288,721,751]" italics="true" pageId="14" pageNumber="13">Herrerasaurus ischigualastensis</emphasis>
</taxonomicName>
(PVSJ 373, Novas 1993: fig. 7c). Thus, we assign these fragmentary proximal ends of femora to
<taxonomicName id="4C2545D4F824096A6653D435FB298F15" authorityName="OWEN" authorityYear="1842" box="[1109,1249,793,819]" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="14" pageNumber="13" phylum="Chordata" rank="order">Dinosauria</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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