<documentID-CLB-Dataset="62490"ID-DOI="http://dx.doi.org/10.3897/zookeys.1103.84015"ID-GBIF-Dataset="1b1f4bf3-acd9-43a4-bf0a-a93e80d70da6"ID-PMC="PMC9848779"ID-Pensoft-Pub="1313-2970-1103-123"ID-Pensoft-UUID="A820088C785659F18D1C8E0F14FC642D"ID-PubMed="36761788"ID-ZooBank="58199DE7FAD743E1A029DC969C6E9BFC"ModsDocID="1313-2970-1103-123"checkinTime="1654168745574"checkinUser="pensoft"docAuthor="Qiu, Lu & Kundrata, Robin"docDate="2022"docId="93A30EAF26ED555896D7C9A1E5BC09A3"docLanguage="en"docName="ZooKeys 1103: 123-138"docOrigin="ZooKeys 1103"docPubDate="2022-06-01"docSource="http://dx.doi.org/10.3897/zookeys.1103.84015"docTitle="Pseudocsikia formosana"docType="treatment"docVersion="5"id="A820088C785659F18D1C8E0F14FC642D"lastPageNumber="123"masterDocId="A820088C785659F18D1C8E0F14FC642D"masterDocTitle="Descriptions of two new flightless species of Pseudocsikia Schimmel & Platia (Coleoptera, Elateridae, Dimini) from Taiwan Island, China, with a definition of the formosana species-group"masterLastPageNumber="138"masterPageNumber="123"pageNumber="123"updateTime="1732803380616"updateUser="ExternalLinkService">
<mods:titleid="40BDB7279A0C8EC2A2B138F1B4E343F3">Descriptions of two new flightless species of Pseudocsikia Schimmel & Platia (Coleoptera, Elateridae, Dimini) from Taiwan Island, China, with a definition of the formosana species-group</mods:title>
<mods:affiliationid="63164FA02DC380D381DADE5BD6EC0DBA">Engineering Research Center for Forest and Grassland Disaster Prevention and Reduction, Mianyang Normal University, Mianxing West Road, 621000, Mianyang, China</mods:affiliation>
<mods:affiliationid="EC8D45D554E6C9F7F9A02959E29FB0B7">Department of Zoology, Faculty of Science, Palacky University, 17. listopadu 50, 77146, Olomouc, Czech Republic</mods:affiliation>
<figureCitationid="5DCDF1BBE96F68BC4E8177BAF9641138"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Habitus and characters of Pseudocsikia formosana (Ohira, 1972) A-H male holotype A habitus, dorsal view B habitus, ventral view C habitus, lateral view D labels E anterior protrusion of pronotum (indicated by an arrow), dorsal view F anterior protrusion of hypomeron (indicated by an arrow), ventral view G posterior angle of pronotum, dorsal view H abdominal tip, ventral view (arrow indicates the apex of median lobe) I, J paratype of an unknown sex I habitus, dorsal view J labels. Scale bars: 1.0 mm (A-C, I); E-H not to scale. All photos provided by Jeremy Frank (BPBM)."figureDoi="10.3897/zookeys.1103.84015.figure2"httpUri="https://binary.pensoft.net/fig/691081"pageId="0"pageNumber="123">Fig. 2A-J</figureCitation>
<bibRefCitationid="86AFBB5BEC56B7EBCB7B27F10DDE97F4"author="Bouwer, R"journalOrPublisher="Entomologische Zeitschrift mit Insektenborse"pageId="0"pageNumber="123"pagination="53 - 58"refId="B1"refString="Bouwer, R, 1991. Beschreibung einer neuen Csikia -Art von Sumatra (Coleoptera: Elateridae). Entomologische Zeitschrift mit Insektenborse 101: 53 - 58"title="Beschreibung einer neuen Csikia - Art von Sumatra (Coleoptera: Elateridae)."volume="101"year="1991">Bouwer 1991</bibRefCitation>
<bibRefCitationid="38508F5B940FA00677ADE14BA1375DA1"author="Jiang, S-H"journalOrPublisher="Annual of the National Taiwan Museum"pageId="0"pageNumber="123"refId="B4"refString="Jiang, S-H, 1993. A catalogue of the Chinese Elateridae. Beijing Agricultural University Publishing House, Beijing, 136-162."title="A catalogue of the Chinese Elateridae. Beijing Agricultural University Publishing House, Beijing, 136 - 162."year="1993">Jiang 1993</bibRefCitation>
<bibRefCitationid="810C6D1A5968F27CB589514CAD4E5235"author="Suzuki, W"journalOrPublisher="Miscellaneous Reports of the Hiwa Museum for Natural History"pageId="0"pageNumber="123"pagination="1 - 348"refId="B21"refString="Suzuki, W, 1999. Catalogue of the family Elateridae (Coleoptera) of Taiwan. Miscellaneous Reports of the Hiwa Museum for Natural History 38: 1 - 348"title="Catalogue of the family Elateridae (Coleoptera) of Taiwan."volume="38"year="1999">Suzuki 1999</bibRefCitation>
<bibRefCitationid="AAE4046E2FD62BCE872184B1A21D7F58"author="Schimmel, R"journalOrPublisher="Koleopterologische Rundschau"pageId="0"pageNumber="123"pagination="245 - 259"refId="B13"refString="Schimmel, R, 1993. Neue Arten sowie eine neue Gattung der Unterfamilie Diminae Candeze, 1863 aus Ostasien und dem Balkan (Coleoptera: Elateridae). Koleopterologische Rundschau 63: 245 - 259"title="Neue Arten sowie eine neue Gattung der Unterfamilie Diminae Candeze, 1863 aus Ostasien und dem Balkan (Coleoptera: Elateridae)."volume="63"year="1993">Schimmel 1993</bibRefCitation>
<bibRefCitationid="4A6471C65B416ECD978549F34DF59AC0"author="Cate, PC"editor="Loebl, I"journalOrPublisher="Apollo Books, Stenstrup"pageId="0"pageNumber="123"pagination="89 - 209"refId="B2"refString="Cate, PC, Sanchez-Ruiz, A, Loebl, I, Smetana, A, 2007. Elateridae. In: Loebl, I, Smetana, A, Eds., Catalogue of Palaearctic Coleoptera, Volume 4. Apollo Books, Stenstrup: 89 - 209"title="Elateridae."volumeTitle="Catalogue of Palaearctic Coleoptera, Volume 4."year="2007">Cate et al. 2007</bibRefCitation>
<bibRefCitationid="099A56D5AF77E6C55606D9E99E2AB5F8"author="Ohira, H"journalOrPublisher="Pacific Insects"pageId="0"pageNumber="123"pagination="1 - 14"refId="B8"refString="Ohira, H, 1972. Elaterid-beetles from Taiwan in Bishop Museum (Coleoptera). Pacific Insects 14: 1 - 14"title="Elaterid-beetles from Taiwan in Bishop Museum (Coleoptera)."volume="14"year="1972">
<bibRefCitationid="8240489847E55960C8A44BC727B734FC"author="Ohira, H"journalOrPublisher="Pacific Insects"pageId="0"pageNumber="123"pagination="1 - 14"refId="B8"refString="Ohira, H, 1972. Elaterid-beetles from Taiwan in Bishop Museum (Coleoptera). Pacific Insects 14: 1 - 14"title="Elaterid-beetles from Taiwan in Bishop Museum (Coleoptera)."volume="14"year="1972">
<figureCitationid="EC1BF6BD8A9CB90F2F8F6BA0056444A9"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Habitus and characters of Pseudocsikia formosana (Ohira, 1972) A-H male holotype A habitus, dorsal view B habitus, ventral view C habitus, lateral view D labels E anterior protrusion of pronotum (indicated by an arrow), dorsal view F anterior protrusion of hypomeron (indicated by an arrow), ventral view G posterior angle of pronotum, dorsal view H abdominal tip, ventral view (arrow indicates the apex of median lobe) I, J paratype of an unknown sex I habitus, dorsal view J labels. Scale bars: 1.0 mm (A-C, I); E-H not to scale. All photos provided by Jeremy Frank (BPBM)."figureDoi="10.3897/zookeys.1103.84015.figure2"httpUri="https://binary.pensoft.net/fig/691081"pageId="0"pageNumber="123">2A-C, I</figureCitation>
<bibRefCitationid="00AFD4F8DCA44BA0ACF75A33B3FBF772"author="Ohira, H"journalOrPublisher="Pacific Insects"pageId="0"pageNumber="123"pagination="1 - 14"refId="B8"refString="Ohira, H, 1972. Elaterid-beetles from Taiwan in Bishop Museum (Coleoptera). Pacific Insects 14: 1 - 14"title="Elaterid-beetles from Taiwan in Bishop Museum (Coleoptera)."volume="14"year="1972">
<bibRefCitationid="930788411D588B42E9F54D2F55239881"author="Ohira, H"journalOrPublisher="Pacific Insects"pageId="0"pageNumber="123"pagination="1 - 14"refId="B8"refString="Ohira, H, 1972. Elaterid-beetles from Taiwan in Bishop Museum (Coleoptera). Pacific Insects 14: 1 - 14"title="Elaterid-beetles from Taiwan in Bishop Museum (Coleoptera)."volume="14"year="1972">
). Punctures on pronotum of moderate size, intervals between punctures mostly equal 3-5 puncture diameters, anterior protrusion of pronotum with apex laterad of center (Fig.
<figureCitationid="21F0CB107457B1F1D5F01B1AB0FD101F"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Habitus and characters of Pseudocsikia formosana (Ohira, 1972) A-H male holotype A habitus, dorsal view B habitus, ventral view C habitus, lateral view D labels E anterior protrusion of pronotum (indicated by an arrow), dorsal view F anterior protrusion of hypomeron (indicated by an arrow), ventral view G posterior angle of pronotum, dorsal view H abdominal tip, ventral view (arrow indicates the apex of median lobe) I, J paratype of an unknown sex I habitus, dorsal view J labels. Scale bars: 1.0 mm (A-C, I); E-H not to scale. All photos provided by Jeremy Frank (BPBM)."figureDoi="10.3897/zookeys.1103.84015.figure2"httpUri="https://binary.pensoft.net/fig/691081"pageId="0"pageNumber="123">2E, F</figureCitation>
<figureCitationid="6353927F672E332A32982F01CC714C0C"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Habitus and characters of Pseudocsikia formosana (Ohira, 1972) A-H male holotype A habitus, dorsal view B habitus, ventral view C habitus, lateral view D labels E anterior protrusion of pronotum (indicated by an arrow), dorsal view F anterior protrusion of hypomeron (indicated by an arrow), ventral view G posterior angle of pronotum, dorsal view H abdominal tip, ventral view (arrow indicates the apex of median lobe) I, J paratype of an unknown sex I habitus, dorsal view J labels. Scale bars: 1.0 mm (A-C, I); E-H not to scale. All photos provided by Jeremy Frank (BPBM)."figureDoi="10.3897/zookeys.1103.84015.figure2"httpUri="https://binary.pensoft.net/fig/691081"pageId="0"pageNumber="123">2G</figureCitation>
Aedeagus with robust median lobe, twice as long as paramere; with large acute lateral processes near midlength, apex rectangular, with large subapical spines laterally (Fig.
<figureCitationid="73A16022F9FEFEE917F8A6B95B5774A8"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Habitus and characters of Pseudocsikia formosana (Ohira, 1972) A-H male holotype A habitus, dorsal view B habitus, ventral view C habitus, lateral view D labels E anterior protrusion of pronotum (indicated by an arrow), dorsal view F anterior protrusion of hypomeron (indicated by an arrow), ventral view G posterior angle of pronotum, dorsal view H abdominal tip, ventral view (arrow indicates the apex of median lobe) I, J paratype of an unknown sex I habitus, dorsal view J labels. Scale bars: 1.0 mm (A-C, I); E-H not to scale. All photos provided by Jeremy Frank (BPBM)."figureDoi="10.3897/zookeys.1103.84015.figure2"httpUri="https://binary.pensoft.net/fig/691081"pageId="0"pageNumber="123">2H</figureCitation>
<bibRefCitationid="A4DFB8D73CA1B2AE44F93EFE26A2A5D0"author="Ohira, H"journalOrPublisher="Pacific Insects"pageId="0"pageNumber="123"pagination="1 - 14"refId="B8"refString="Ohira, H, 1972. Elaterid-beetles from Taiwan in Bishop Museum (Coleoptera). Pacific Insects 14: 1 - 14"title="Elaterid-beetles from Taiwan in Bishop Museum (Coleoptera)."volume="14"year="1972">
and not observable in this study). Paramere stout, reaching half length of median lobe; apex rounded, slightly outwards. Phallobase subquadrate, margins thickened, medially with longitudinal thickened line, basal angles rounded (based on
<bibRefCitationid="B0D952A0140E025A0A44145B3315E283"author="Ohira, H"journalOrPublisher="Pacific Insects"pageId="0"pageNumber="123"pagination="1 - 14"refId="B8"refString="Ohira, H, 1972. Elaterid-beetles from Taiwan in Bishop Museum (Coleoptera). Pacific Insects 14: 1 - 14"title="Elaterid-beetles from Taiwan in Bishop Museum (Coleoptera)."volume="14"year="1972">
<subSubSectionid="93CA4559A91AA46789B4FDD2145767D1"pageId="0"pageNumber="123"type="checklist of pseudocsikia species from china, with notes on their type localities">
<taxonomicNameid="E3B937E5ECCD509CB626162FC5C770FC"authorityName="Schimmel & Platia"authorityYear="1991"class="Insecta"family="Elateridae"genus="Pseudocsikia"higherTaxonomySource="CoL"kingdom="Animalia"lsidName="Pseudocsikia"order="Coleoptera"pageId="0"pageNumber="123"phylum="Arthropoda"rank="genus">
<bibRefCitationid="E9C54A8F7FCB88240C620D5DB56A57F3"author="Ohira, H"journalOrPublisher="Pacific Insects"pageId="0"pageNumber="123"pagination="1 - 14"refId="B8"refString="Ohira, H, 1972. Elaterid-beetles from Taiwan in Bishop Museum (Coleoptera). Pacific Insects 14: 1 - 14"title="Elaterid-beetles from Taiwan in Bishop Museum (Coleoptera)."volume="14"year="1972">
<bibRefCitationid="D8B8B7F518902574FA4804D197E3AA05"author="Chu, Y-I"journalOrPublisher="Annual of the National Taiwan Museum"pageId="0"pageNumber="123"pagination="121 - 150"refId="B3"refString="Chu, Y-I, Yamanaka, T, 1973. A check List of the present and old name of insect collected localities in Taiwan. Annual of the National Taiwan Museum 18: 121 - 150"title="A check List of the present and old name of insect collected localities in Taiwan."volume="18"year="1973">Chu and Yamanaka 1973</bibRefCitation>
<bibRefCitationid="F7152BD987F39F04358A0DA3C3373E32"author="Schimmel, R"journalOrPublisher="Koleopterologische Rundschau"pageId="0"pageNumber="123"pagination="161 - 177"refId="B15"refString="Schimmel, R, 1996b. Neue und wenig bekannte Elateriden aus Griechenland und Ostasien (Coleoptera: Elateridae). Koleopterologische Rundschau 66: 161 - 177"title="Neue und wenig bekannte Elateriden aus Griechenland und Ostasien (Coleoptera: Elateridae)."volume="66"year="1996 b">Schimmel 1996b</bibRefCitation>
: Gaoligongshan [高黎贡山] is an extensive mountain range lying on the border of Yunnan, China and Myanmar; the exact locality information of the holotype is unknown. However, according to the original paper, the holotype was collected 100 km west of Baoshan, which is near Tengchong.
<taxonomicNameid="311284641A34BC867E0A24893C62B0D7"authorityName="Qiu & Kundrata"authorityYear="2022"class="Insecta"family="Elateridae"genus="Pseudocsikia"higherTaxonomySource="CoL"kingdom="Animalia"lsidName="Pseudocsikia choui"order="Coleoptera"pageId="0"pageNumber="123"phylum="Arthropoda"rank="species"species="choui">
<taxonomicNameid="256EE0BC102F430D13BB62BDC7395CB8"authorityName="Qiu & Kundrata"authorityYear="2022"class="Insecta"family="Elateridae"genus="Pseudocsikia"higherTaxonomySource="CoL"kingdom="Animalia"lsidName="Pseudocsikia chanjuan"order="Coleoptera"pageId="0"pageNumber="123"phylum="Arthropoda"rank="species"species="chanjuan">
