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<document id="6690E386CD041993E6B5C8C93D5AE505" ID-CLB-Dataset="51371" ID-DOI="10.5281/zenodo.156420" ID-GBIF-Dataset="76c096e3-d27c-4abc-a504-a62ae8c7ba92" ID-ISSN="1175­5326" ID-Zenodo-Dep="156420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459586556453" checkinUser="plazi" docAuthor="Gusarov, Vladimir I." docDate="2003" docId="62088784C317C74DE12E72B7FED2FE66" docLanguage="en" docName="zt00142.pdf" docOrigin="Zootaxa 142" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Seeversiella brevipennis Gusarov, sp. n." docType="treatment" docVersion="10" lastPageNumber="62" masterDocId="9E31FFFCC32EC770E0267411FFBAFFC8" masterDocTitle="A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="102" masterPageNumber="1" pageNumber="58" updateTime="1698203169024" updateUser="ExternalLinkService">
<mods:mods id="C77CDD7956A30B635F9A192DF9036FAB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DA180343A2B948F573C634B4D61F1B24">
<mods:title id="2F04BCC81938C116CA5C4FDB180A847D">A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
</mods:titleInfo>
<mods:name id="D12828C5B5F3E5D9ED514D20A0115731" type="personal">
<mods:role id="2EEC12E2F6BB006C17A4BEA915AD4ADC">
<mods:roleTerm id="BD813E1D44A6ACEB00360CEF046A24F2">Author</mods:roleTerm>
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<mods:namePart id="5EA46DE9C9654DCC953E3FCDC514A639">Gusarov, Vladimir I.</mods:namePart>
</mods:name>
<mods:typeOfResource id="8274D08214CBD8C9619863B36EE1C20D">text</mods:typeOfResource>
<mods:relatedItem id="D8346D5C9208F96C825E48F11CB27BA9" type="host">
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<mods:title id="69434D7484B0FAC09FBF03F067CC00D6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="61310F7D5EF4B48A37F355B8D9E49C3C">
<mods:date id="41624747C4BBB8403E51448E2AB6D987">2003</mods:date>
<mods:detail id="BF58F5D3BBC166286529914E30C3A36A" type="volume">
<mods:number id="1ABB7EAD95629C9C997DA43BCEC6F4D8">142</mods:number>
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<mods:classification id="E2D8472A4F30FEAADE553B603E365D09">journal article</mods:classification>
<mods:identifier id="5318948D98A8FE2C9EB1F630567ECD6D" type="CLB-Dataset">51371</mods:identifier>
<mods:identifier id="9D4A0798955A3508E5AE03F9B08AD673" type="DOI">10.5281/zenodo.156420</mods:identifier>
<mods:identifier id="7B03E1A2AD7B24AFFD287A9E7491C4A9" type="GBIF-Dataset">76c096e3-d27c-4abc-a504-a62ae8c7ba92</mods:identifier>
<mods:identifier id="E9EB306D86685BBCFA955F7CEFB77374" type="ISSN">1175­5326</mods:identifier>
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<treatment id="62088784C317C74DE12E72B7FED2FE66" ID-DOI="http://doi.org/10.5281/zenodo.6274670" ID-Zenodo-Dep="6274670" LSID="urn:lsid:plazi:treatment:62088784C317C74DE12E72B7FED2FE66" httpUri="http://treatment.plazi.org/id/62088784C317C74DE12E72B7FED2FE66" lastPageId="61" lastPageNumber="62" pageId="57" pageNumber="58">
<subSubSection id="A2BB6519C317C749E12E72B7FC62F908" box="[264,984,1702,1729]" pageId="57" pageNumber="58" type="key">
<paragraph id="EA1E3692C317C749E12E72B7FC62F908" blockId="57.[264,984,1702,1729]" box="[264,984,1702,1729]" pageId="57" pageNumber="58">
<heading id="B15681FEC317C749E12E72B7FC62F908" bold="true" box="[264,984,1702,1729]" fontSize="11" level="1" pageId="57" pageNumber="58" reason="1">
<keyLead id="515095A7C317C749E12E72B7FC62F908" box="[264,984,1702,1729]" pageId="57" pageNumber="58">
<emphasis id="D8D5EA80C317C749E12E72B7FC62F908" bold="true" box="[264,984,1702,1729]" pageId="57" pageNumber="58">
<keyStep id="51552E37C317C749E12E72B7FC62F908" box="[264,984,1702,1729]" pageId="57" pageNumber="58">
<key id="9E3061A3C317C749E12E72B7FC62F908" box="[264,984,1702,1729]" pageId="57" pageNumber="58">
16.
<taxonomicName id="2DA14D11C317C749E11272B6FD70F908" ID-CoL="6YBCM" authority="Gusarov" authorityName="Gusarov" box="[308,714,1702,1729]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="brevipennis" status="sp. nov.">
<emphasis id="D8D5EA80C317C749E11272B6FDEDF909" bold="true" box="[308,599,1703,1729]" italics="true" pageId="57" pageNumber="58">Seeversiella brevipennis</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC317C749E2F072B7FCA3F908" box="[726,793,1702,1728]" pageId="57" pageNumber="58" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C317C749E30E72B7FC6AF908" box="[808,976,1702,1728]" captionStart-0="FIGURES 222 ­ 225" captionStart-1="FIGURES 226 ­ 229" captionStart-2="FIGURES 230 ­ 234" captionStartId-0="58.[264,382,1219,1243]" captionStartId-1="59.[264,383,1174,1198]" captionStartId-2="60.[264,382,1729,1753]" captionTargetBox-0="[284,1303,333,1185]" captionTargetBox-1="[307,1283,459,1141]" captionTargetBox-2="[294,1292,824,1693]" captionTargetId-0="figure@58.[273,1314,320,1192]" captionTargetId-1="figure@59.[296,1291,449,1147]" captionTargetId-2="figure@60.[281,1305,809,1702]" captionTargetPageId-0="58" captionTargetPageId-1="59" captionTargetPageId-2="60" captionText-0="FIGURES 222 ­ 225. Abdominal segment 8 of Seeversiella brevipennis Gusarov, sp. n. (222 ­ 223, holotype; 224 ­ 225, paratype, 5.9 km N Cerro Punta, Panama). 222 – male tergum 8; 223 – male sternum 8; 224 – female tergum 8; 225 – female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 226 ­ 229. Aedeagus of Seeversiella brevipennis Gusarov, sp. n. (holotype). 226 – median lobe, parameral view; 227 – apex of median lobe, parameral view; 228 – median lobe, lateral view; 229 – apex of median lobe, lateral view. Scale bar 0.2 mm (226, 228), 0.1 mm (227, 229)." captionText-2="FIGURES 230 ­ 234. Genitalia of Seeversiella brevipennis Gusarov, sp. n. (230 ­ 233, holotype; 234, paratype, 5.9 km N Cerro Punta, Panama). 230 ­ 231 – details of internal sac retracted into median lobe, lateral view; 232 – details of internal sac retracted into median lobe, abparameral view; 233 – apex of left paramere; 234 – spermatheca. Scale bar 0.2 mm (230 ­ 232), 0.1 mm (233 ­ 234)." httpUri-0="https://zenodo.org/record/156458/files/figure.png" httpUri-1="https://zenodo.org/record/156459/files/figure.png" httpUri-2="https://zenodo.org/record/156460/files/figure.png" pageId="57" pageNumber="58">Figs. 222­234</figureCitation>
)
</key>
</keyStep>
</emphasis>
</keyLead>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C317C74AE12E72E0FB6CFA58" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58" type="materials_examined">
<paragraph id="EA1E3692C317C749E12E72E0FCC8F894" blockId="57.[264,1323,1777,1884]" pageId="57" pageNumber="58">
<emphasis id="D8D5EA80C317C749E12E72E0FE04F8C3" bold="true" box="[264,446,1777,1803]" pageId="57" pageNumber="58">
<typeStatus id="351A8830C317C749E12E72E0FEF9F8C3" box="[264,323,1777,1803]" pageId="57" pageNumber="58">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C317C749E1F372E3FDFFF8C4" box="[469,581,1778,1804]" pageId="57" pageNumber="58" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C317C749E25E72E0FC31F8C3" bold="true" box="[632,907,1777,1803]" pageId="57" pageNumber="58">
<collectingCountry id="92B67602C317C749E25E72E0FD47F8C3" box="[632,765,1777,1803]" name="Panama" pageId="57" pageNumber="58">PANAMA</collectingCountry>
: Chiriquí:
</emphasis>
<quantity id="2D599B77C317C749E3B972E3FC4EF8C4" box="[927,1012,1778,1804]" metricMagnitude="3" metricUnit="m" metricValue="5.9" pageId="57" pageNumber="58" unit="km" value="5.9">5.9 km</quantity>
N Cerro Punta, Par. Nac. Volcán Barú,
<geoCoordinate id="8F955055C317C749E181730BFE43F8FC" box="[423,505,1818,1844]" direction="north" orientation="latitude" pageId="57" pageNumber="58" precision="925" value="8.866667">8°52'N</geoCoordinate>
[the latitude on the label (
<geoCoordinate id="8F955055C317C749E30D730BFC22F8FC" box="[811,920,1818,1844]" direction="north" orientation="latitude" pageId="57" pageNumber="58" precision="15" value="8.366667">8°22'0&quot;N</geoCoordinate>
) is incorrect]
<geoCoordinate id="8F955055C317C749E41A730BFB07F8FC" box="[1084,1213,1818,1844]" direction="west" orientation="longitude" pageId="57" pageNumber="58" precision="15" value="-82.566666">82°34'0&quot;W</geoCoordinate>
,
<quantity id="2D599B77C317C749E4EF730BFA9EF8FC" box="[1225,1316,1818,1844]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="57" pageNumber="58" unit="m" value="2150.0">2150 m</quantity>
, alder forest litter (R.Anderson),
<date id="9E1F1052C317C749E2A77353FD44F894" box="[641,766,1858,1884]" pageId="57" pageNumber="58" value="1995-06-14">14.vi.1995</date>
(
<collectionCode id="8CB0AE57C317C749E3287353FCDCF894" box="[782,870,1858,1884]" pageId="57" pageNumber="58">KSEM</collectionCode>
).
</paragraph>
<caption id="BEDE661AC314C74AE12E70D2FBEAFAE8" httpUri="https://zenodo.org/record/156458/files/figure.png" pageId="58" pageNumber="59" targetBox="[284,1303,333,1185]" targetPageId="58">
<paragraph id="EA1E3692C314C74AE12E70D2FBEAFAE8" blockId="58.[264,1323,1219,1312]" pageId="58" pageNumber="59">
<emphasis id="D8D5EA80C314C74AE12E70D2FE5CFB13" bold="true" box="[264,486,1219,1243]" pageId="58" pageNumber="59">FIGURES 222­225.</emphasis>
Abdominal segment 8 of
<taxonomicName id="2DA14D11C314C74AE32C70D5FBC8FB12" authority="Gusarov" authorityName="Gusarov" box="[778,1138,1219,1243]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="brevipennis" status="sp. nov.">
<emphasis id="D8D5EA80C314C74AE32C70D5FBB4FB13" box="[778,1038,1220,1243]" italics="true" pageId="58" pageNumber="59">Seeversiella brevipennis</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C314C74AE45B70D2FB00FB13" bold="true" box="[1149,1210,1219,1243]" pageId="58" pageNumber="59">
<taxonomicNameLabel id="C3E657FBC314C74AE45B70D2FB00FB13" box="[1149,1210,1219,1243]" pageId="58" pageNumber="59" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(222­223, holotype; 224­225, paratype, 5.9 km N Cerro Punta, Panama). 222 – male tergum 8; 223 – male sternum 8; 224 – female tergum 8; 225 – female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<paragraph id="EA1E3692C314C74AE11E7167FB6CFA58" blockId="58.[264,1324,1397,1904]" box="[312,1238,1397,1424]" pageId="58" pageNumber="59">
<typeStatus id="351A8830C314C74AE11E7167FE14FA58" box="[312,430,1398,1424]" pageId="58" pageNumber="59" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C314C74AE1E67164FD74FA47" bold="true" box="[448,718,1397,1423]" pageId="58" pageNumber="59">
<collectingCountry id="92B67602C314C74AE1E67164FDFFFA47" box="[448,581,1397,1423]" name="Panama" pageId="58" pageNumber="59">PANAMA</collectingCountry>
: Chiriquí:
</emphasis>
,, same data as the
<typeStatus id="351A8830C314C74AE3DD7167FBD8FA58" box="[1019,1122,1398,1424]" pageId="58" pageNumber="59" type="holotype">holotype</typeStatus>
(
<collectionCode id="8CB0AE57C314C74AE4547167FB70FA58" box="[1138,1226,1398,1424]" pageId="58" pageNumber="59">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C314C74AE11E718CFCA0F930" pageId="58" pageNumber="59" type="diagnosis">
<paragraph id="EA1E3692C314C74AE11E718CFBF2F948" blockId="58.[264,1324,1397,1904]" pageId="58" pageNumber="59">
<emphasis id="D8D5EA80C314C74AE11E718CFE02FA7F" bold="true" box="[312,440,1437,1463]" pageId="58" pageNumber="59">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C314C74AE1EC718FFD50FA7F" box="[458,746,1438,1463]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="D8D5EA80C314C74AE1EC718FFD50FA7F" box="[458,746,1438,1463]" italics="true" pageId="58" pageNumber="59">Seeversiella brevipennis</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C314C74AE4DE718FFED7FA17" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C314C74AE4DE718FFED7FA17" italics="true" pageId="58" pageNumber="59">Seeversiella</emphasis>
</taxonomicName>
by having brown or reddish brown body with yellowish brown elytra, abdominal segments 3­4, legs and mouthparts; large eyes (temple length to eye length ratio 1.6­2.0); glossy pronotum with weak isodiametric microsculpture and asperate punctation; elytra as long as pronotum; wings partially reduced; tergum 8 with three pairs of macrosetae; the distinct shape of aedeagus (
<figureCitation id="729A2A17C314C74AE26D7277FD4AF948" box="[587,752,1638,1664]" captionStart-0="FIGURES 226 ­ 229" captionStart-1="FIGURES 230 ­ 234" captionStartId-0="59.[264,383,1174,1198]" captionStartId-1="60.[264,382,1729,1753]" captionTargetBox-0="[307,1283,459,1141]" captionTargetBox-1="[294,1292,824,1693]" captionTargetId-0="figure@59.[296,1291,449,1147]" captionTargetId-1="figure@60.[281,1305,809,1702]" captionTargetPageId-0="59" captionTargetPageId-1="60" captionText-0="FIGURES 226 ­ 229. Aedeagus of Seeversiella brevipennis Gusarov, sp. n. (holotype). 226 – median lobe, parameral view; 227 – apex of median lobe, parameral view; 228 – median lobe, lateral view; 229 – apex of median lobe, lateral view. Scale bar 0.2 mm (226, 228), 0.1 mm (227, 229)." captionText-1="FIGURES 230 ­ 234. Genitalia of Seeversiella brevipennis Gusarov, sp. n. (230 ­ 233, holotype; 234, paratype, 5.9 km N Cerro Punta, Panama). 230 ­ 231 – details of internal sac retracted into median lobe, lateral view; 232 – details of internal sac retracted into median lobe, abparameral view; 233 – apex of left paramere; 234 – spermatheca. Scale bar 0.2 mm (230 ­ 232), 0.1 mm (233 ­ 234)." httpUri-0="https://zenodo.org/record/156459/files/figure.png" httpUri-1="https://zenodo.org/record/156460/files/figure.png" pageId="58" pageNumber="59">Figs. 226­233</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C314C74AE3F27277FB86F948" box="[980,1084,1638,1664]" captionStart="FIGURES 230 ­ 234" captionStartId="60.[264,382,1729,1753]" captionTargetBox="[294,1292,824,1693]" captionTargetId="figure@60.[281,1305,809,1702]" captionTargetPageId="60" captionText="FIGURES 230 ­ 234. Genitalia of Seeversiella brevipennis Gusarov, sp. n. (230 ­ 233, holotype; 234, paratype, 5.9 km N Cerro Punta, Panama). 230 ­ 231 – details of internal sac retracted into median lobe, lateral view; 232 – details of internal sac retracted into median lobe, abparameral view; 233 – apex of left paramere; 234 – spermatheca. Scale bar 0.2 mm (230 ­ 232), 0.1 mm (233 ­ 234)." httpUri="https://zenodo.org/record/156460/files/figure.png" pageId="58" pageNumber="59">Fig. 234</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C314C74AE11E729FFCA0F930" blockId="58.[264,1324,1397,1904]" pageId="58" pageNumber="59">
<taxonomicName id="2DA14D11C314C74AE11E729FFDEDF96F" box="[312,599,1678,1703]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="D8D5EA80C314C74AE11E729FFDEDF96F" box="[312,599,1678,1703]" italics="true" pageId="58" pageNumber="59">Seeversiella brevipennis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C314C74AE2DF729FFC18F96F" box="[761,930,1678,1703]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C314C74AE2DF729FFC18F96F" box="[761,930,1678,1703]" italics="true" pageId="58" pageNumber="59">S. scabricollis</emphasis>
</taxonomicName>
in having glossy pronotum with asperate punctation; elytra as long as pronotum; apex of median lobe in parameral view constricted basally (
<figureCitation id="729A2A17C314C74AE1D572CFFD20F930" box="[499,666,1758,1784]" captionStart="FIGURES 226 ­ 229" captionStartId="59.[264,383,1174,1198]" captionTargetBox="[307,1283,459,1141]" captionTargetId="figure@59.[296,1291,449,1147]" captionTargetPageId="59" captionText="FIGURES 226 ­ 229. Aedeagus of Seeversiella brevipennis Gusarov, sp. n. (holotype). 226 – median lobe, parameral view; 227 – apex of median lobe, parameral view; 228 – median lobe, lateral view; 229 – apex of median lobe, lateral view. Scale bar 0.2 mm (226, 228), 0.1 mm (227, 229)." httpUri="https://zenodo.org/record/156459/files/figure.png" pageId="58" pageNumber="59">Figs. 226­227</figureCitation>
; 194­196).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C314C74CE11E7317FDC4FD0E" lastPageId="60" lastPageNumber="61" pageId="58" pageNumber="59" type="description">
<paragraph id="EA1E3692C314C74AE11E7317FE56F8B8" blockId="58.[264,1324,1397,1904]" pageId="58" pageNumber="59">
<taxonomicName id="2DA14D11C314C74AE11E7317FDEFF8D7" box="[312,597,1798,1823]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="D8D5EA80C314C74AE11E7317FDEFF8D7" box="[312,597,1798,1823]" italics="true" pageId="58" pageNumber="59">Seeversiella brevipennis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C314C74AE2D77317FC16F8D7" box="[753,940,1798,1823]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="furcativentris">
<emphasis id="D8D5EA80C314C74AE2D77317FC16F8D7" box="[753,940,1798,1823]" italics="true" pageId="58" pageNumber="59">S. furcativentris</emphasis>
</taxonomicName>
in having glossy pronotum with asperate punctation; elytra as long as pronotum; larger median lobe with longer apex (
<figureCitation id="729A2A17C314C74AE4D7733FFED7F8B8" captionStart="FIGURES 226 ­ 229" captionStartId="59.[264,383,1174,1198]" captionTargetBox="[307,1283,459,1141]" captionTargetId="figure@59.[296,1291,449,1147]" captionTargetPageId="59" captionText="FIGURES 226 ­ 229. Aedeagus of Seeversiella brevipennis Gusarov, sp. n. (holotype). 226 – median lobe, parameral view; 227 – apex of median lobe, parameral view; 228 – median lobe, lateral view; 229 – apex of median lobe, lateral view. Scale bar 0.2 mm (226, 228), 0.1 mm (227, 229)." httpUri="https://zenodo.org/record/156459/files/figure.png" pageId="58" pageNumber="59">Figs. 226­229</figureCitation>
; 212­215).
</paragraph>
<paragraph id="EA1E3692C315C74BE11E750DFD3BFE4E" blockId="59.[264,1323,284,390]" pageId="59" pageNumber="60">
<taxonomicName id="2DA14D11C315C74BE11E750DFDE1FEFD" box="[312,603,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="D8D5EA80C315C74BE11E750DFDE1FEFD" box="[312,603,284,309]" italics="true" pageId="59" pageNumber="60">Seeversiella brevipennis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C315C74BE32F750DFC35FEFD" box="[777,911,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="D8D5EA80C315C74BE32F750DFC35FEFD" box="[777,911,284,309]" italics="true" pageId="59" pageNumber="60">S. brunnea</emphasis>
</taxonomicName>
in having glossy pronotum with asperate punctation; elytra as long as pronotum; straight apex of median lobe (in lateral view) (
<figureCitation id="729A2A17C315C74BE17D757DFDB8FE4E" box="[347,514,364,390]" captionStart="FIGURES 226 ­ 229" captionStartId="59.[264,383,1174,1198]" captionTargetBox="[307,1283,459,1141]" captionTargetId="figure@59.[296,1291,449,1147]" captionTargetPageId="59" captionText="FIGURES 226 ­ 229. Aedeagus of Seeversiella brevipennis Gusarov, sp. n. (holotype). 226 – median lobe, parameral view; 227 – apex of median lobe, parameral view; 228 – median lobe, lateral view; 229 – apex of median lobe, lateral view. Scale bar 0.2 mm (226, 228), 0.1 mm (227, 229)." httpUri="https://zenodo.org/record/156459/files/figure.png" pageId="59" pageNumber="60">Figs. 228­229</figureCitation>
; 241­242).
</paragraph>
<caption id="BEDE661AC315C74BE12E7087FEFAFADD" httpUri="https://zenodo.org/record/156459/files/figure.png" pageId="59" pageNumber="60" targetBox="[307,1283,459,1141]" targetPageId="59">
<paragraph id="EA1E3692C315C74BE12E7087FEFAFADD" blockId="59.[264,1323,1174,1301]" pageId="59" pageNumber="60">
<emphasis id="D8D5EA80C315C74BE12E7087FE59FB66" bold="true" box="[264,483,1174,1198]" pageId="59" pageNumber="60">FIGURES 226­229</emphasis>
. Aedeagus of
<taxonomicName id="2DA14D11C315C74BE2B37087FBB9FB65" authority="Gusarov" authorityName="Gusarov" box="[661,1027,1174,1197]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="brevipennis" status="sp. nov.">
<emphasis id="D8D5EA80C315C74BE2B37087FC26FB65" box="[661,924,1174,1197]" italics="true" pageId="59" pageNumber="60">Seeversiella brevipennis</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC315C74BE4347087FBE9FB66" box="[1042,1107,1174,1198]" pageId="59" pageNumber="60" rank="species">
<emphasis id="D8D5EA80C315C74BE4347087FBE9FB66" bold="true" box="[1042,1107,1174,1198]" pageId="59" pageNumber="60">sp. n.</emphasis>
</taxonomicNameLabel>
(holotype). 226 – median lobe, parameral view; 227 – apex of median lobe, parameral view; 228 – median lobe, lateral view; 229 – apex of median lobe, lateral view. Scale bar 0.2 mm (226, 228), 0.1 mm (227, 229).
</paragraph>
</caption>
<paragraph id="EA1E3692C315C74BE11E717AFC1BFA66" blockId="59.[264,1324,1387,1894]" pageId="59" pageNumber="60">
<emphasis id="D8D5EA80C315C74BE11E717AFE75FA4D" bold="true" box="[312,463,1387,1413]" pageId="59" pageNumber="60">Description.</emphasis>
Length 2.6­3.0 mm. Body brown or reddish brown, elytra, abdominal segments 3 and 4, legs and mouthparts yellowish brown.
</paragraph>
<paragraph id="EA1E3692C315C74BE11E71ADFE87F986" blockId="59.[264,1324,1387,1894]" pageId="59" pageNumber="60">Head surface glossy, with fine isodiametric microsculpture, fine and slightly asperate punctation, distance between punctures equals 2­3 times their diameter. Temple length to eye length ratio 1.6­2.0. Antennal article 2 as long as 3, articles 4­10 transverse (ratio 1.3­ 1.6).</paragraph>
<paragraph id="EA1E3692C315C74BE11E724DFB27F8AE" blockId="59.[264,1324,1387,1894]" pageId="59" pageNumber="60">
Pronotum slightly transverse, 1.2 times as wide as head, width
<quantity id="2D599B77C315C74BE40D724DFB6CF9BE" box="[1067,1238,1628,1654]" metricMagnitude="-4" metricUnit="m" metricValue="5.45" metricValueMax="5.6" metricValueMin="5.3" pageId="59" pageNumber="60" unit="mm" value="0.545" valueMax="0.56" valueMin="0.53">0.53­0.56 mm</quantity>
, length
<quantity id="2D599B77C315C74BE12E7295FE1FF956" box="[264,421,1668,1694]" metricMagnitude="-4" metricUnit="m" metricValue="4.85" metricValueMax="5.0" metricValueMin="4.699999999999999" pageId="59" pageNumber="60" unit="mm" value="0.485" valueMax="0.5" valueMin="0.47">0.47­0.5 mm</quantity>
, width to length ratio 1.1, surface glossy, with fine and weak isodiametric microsculpture; punctation fine and asperate, distance between punctures equal to 1­1.5 times their diameter. Elytra wider and as long (measured from humeral angle) as pronotum (pronotal length to elytral length ratio 1.0), 1.4 times wider than long, glossy, with fine and weak isodiametric microsculpture; with fine and asperate punctation, distance between punctures equal to 1­2 times their diameter. Wings partially reduced.
</paragraph>
<paragraph id="EA1E3692C312C74CE11E750DFC3CFE66" blockId="60.[264,1324,284,710]" pageId="60" pageNumber="61">
Abdominal terga glossy, with fine microsculpture consisting of transverse meshes, with fine punctation, distance between punctures equals 3­6 times their diameter on terga 3­5 and 4­7 times on tergum 7. Apical margin of tergum 7 with white palisade fringe. Tergum 8 with three pairs of macrosetae (
<figureCitation id="729A2A17C312C74CE2ED7585FCC3FE66" box="[715,889,404,430]" captionStart="FIGURES 222 ­ 225" captionStartId="58.[264,382,1219,1243]" captionTargetBox="[284,1303,333,1185]" captionTargetId="figure@58.[273,1314,320,1192]" captionTargetPageId="58" captionText="FIGURES 222 ­ 225. Abdominal segment 8 of Seeversiella brevipennis Gusarov, sp. n. (222 ­ 223, holotype; 224 ­ 225, paratype, 5.9 km N Cerro Punta, Panama). 222 – male tergum 8; 223 – male sternum 8; 224 – female tergum 8; 225 – female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156458/files/figure.png" pageId="60" pageNumber="61">Figs. 222, 224</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C312C74CE11E75ADFEFFFD86" blockId="60.[264,1324,284,710]" pageId="60" pageNumber="61">
In males posterior angles of tergum 3 projecting as spines up to 2 times as long as tergum 3 (measured medially); tergum
<quantity id="2D599B77C312C74CE2E775F5FD4AFE36" box="[705,752,484,510]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="60" pageNumber="61" unit="in" value="7.0">7 in</quantity>
front of posterior margin with medial tubercle; posterior margin of male tergum 8 with emargination delimited by short projections (
<figureCitation id="729A2A17C312C74CE4DD761DFE83FD86" captionStart="FIGURES 222 ­ 225" captionStartId="58.[264,382,1219,1243]" captionTargetBox="[284,1303,333,1185]" captionTargetId="figure@58.[273,1314,320,1192]" captionTargetPageId="58" captionText="FIGURES 222 ­ 225. Abdominal segment 8 of Seeversiella brevipennis Gusarov, sp. n. (222 ­ 223, holotype; 224 ­ 225, paratype, 5.9 km N Cerro Punta, Panama). 222 – male tergum 8; 223 – male sternum 8; 224 – female tergum 8; 225 – female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156458/files/figure.png" pageId="60" pageNumber="61">Fig. 222</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C312C74CE11E764DFD48FD56" blockId="60.[264,1324,284,710]" pageId="60" pageNumber="61">
Aedeagus as in
<figureCitation id="729A2A17C312C74CE1D4764DFD23FDBE" box="[498,665,604,630]" captionStart-0="FIGURES 226 ­ 229" captionStart-1="FIGURES 230 ­ 234" captionStartId-0="59.[264,383,1174,1198]" captionStartId-1="60.[264,382,1729,1753]" captionTargetBox-0="[307,1283,459,1141]" captionTargetBox-1="[294,1292,824,1693]" captionTargetId-0="figure@59.[296,1291,449,1147]" captionTargetId-1="figure@60.[281,1305,809,1702]" captionTargetPageId-0="59" captionTargetPageId-1="60" captionText-0="FIGURES 226 ­ 229. Aedeagus of Seeversiella brevipennis Gusarov, sp. n. (holotype). 226 – median lobe, parameral view; 227 – apex of median lobe, parameral view; 228 – median lobe, lateral view; 229 – apex of median lobe, lateral view. Scale bar 0.2 mm (226, 228), 0.1 mm (227, 229)." captionText-1="FIGURES 230 ­ 234. Genitalia of Seeversiella brevipennis Gusarov, sp. n. (230 ­ 233, holotype; 234, paratype, 5.9 km N Cerro Punta, Panama). 230 ­ 231 – details of internal sac retracted into median lobe, lateral view; 232 – details of internal sac retracted into median lobe, abparameral view; 233 – apex of left paramere; 234 – spermatheca. Scale bar 0.2 mm (230 ­ 232), 0.1 mm (233 ­ 234)." httpUri-0="https://zenodo.org/record/156459/files/figure.png" httpUri-1="https://zenodo.org/record/156460/files/figure.png" pageId="60" pageNumber="61">Figs. 226­233</figureCitation>
. Apex of paramere broad, proximal seta approximately as long as the other three setae (
<figureCitation id="729A2A17C312C74CE2587695FD5FFD56" box="[638,741,644,670]" captionStart="FIGURES 230 ­ 234" captionStartId="60.[264,382,1729,1753]" captionTargetBox="[294,1292,824,1693]" captionTargetId="figure@60.[281,1305,809,1702]" captionTargetPageId="60" captionText="FIGURES 230 ­ 234. Genitalia of Seeversiella brevipennis Gusarov, sp. n. (230 ­ 233, holotype; 234, paratype, 5.9 km N Cerro Punta, Panama). 230 ­ 231 – details of internal sac retracted into median lobe, lateral view; 232 – details of internal sac retracted into median lobe, abparameral view; 233 – apex of left paramere; 234 – spermatheca. Scale bar 0.2 mm (230 ­ 232), 0.1 mm (233 ­ 234)." httpUri="https://zenodo.org/record/156460/files/figure.png" pageId="60" pageNumber="61">Fig. 233</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C312C74CE11E76BDFDC4FD0E" blockId="60.[264,1324,284,710]" box="[312,638,684,710]" pageId="60" pageNumber="61">
Spermatheca as in
<figureCitation id="729A2A17C312C74CE23276BDFDC0FD0E" box="[532,634,684,710]" captionStart="FIGURES 230 ­ 234" captionStartId="60.[264,382,1729,1753]" captionTargetBox="[294,1292,824,1693]" captionTargetId="figure@60.[281,1305,809,1702]" captionTargetPageId="60" captionText="FIGURES 230 ­ 234. Genitalia of Seeversiella brevipennis Gusarov, sp. n. (230 ­ 233, holotype; 234, paratype, 5.9 km N Cerro Punta, Panama). 230 ­ 231 – details of internal sac retracted into median lobe, lateral view; 232 – details of internal sac retracted into median lobe, abparameral view; 233 – apex of left paramere; 234 – spermatheca. Scale bar 0.2 mm (230 ­ 232), 0.1 mm (233 ­ 234)." httpUri="https://zenodo.org/record/156460/files/figure.png" pageId="60" pageNumber="61">Fig. 234</figureCitation>
.
</paragraph>
</subSubSection>
<caption id="BEDE661AC312C74CE12E72D0FB63F888" httpUri="https://zenodo.org/record/156460/files/figure.png" pageId="60" pageNumber="61" targetBox="[294,1292,824,1693]" targetPageId="60">
<paragraph id="EA1E3692C312C74CE12E72D0FB63F888" blockId="60.[264,1323,1729,1856]" pageId="60" pageNumber="61">
<emphasis id="D8D5EA80C312C74CE12E72D0FE67F911" bold="true" box="[264,477,1729,1753]" pageId="60" pageNumber="61">FIGURES 230­234</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C312C74CE25272D3FC6DF910" authority="Gusarov" authorityName="Gusarov" box="[628,983,1729,1753]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="brevipennis" status="sp. nov.">
<emphasis id="D8D5EA80C312C74CE25272D3FCCCF911" box="[628,886,1730,1753]" italics="true" pageId="60" pageNumber="61">Seeversiella brevipennis</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C312C74CE3C772D0FBA6F911" bold="true" box="[993,1052,1729,1753]" pageId="60" pageNumber="61">
<taxonomicNameLabel id="C3E657FBC312C74CE3C772D0FBA6F911" box="[993,1052,1729,1753]" pageId="60" pageNumber="61" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(230­233, holotype; 234, paratype, 5.9 km N Cerro Punta, Panama). 230­231 – details of internal sac retracted into median lobe, lateral view; 232 – details of internal sac retracted into median lobe, abparameral view; 233 – apex of left paramere; 234 – spermatheca. Scale bar 0.2 mm (230­232), 0.1 mm (233­234).
</paragraph>
</caption>
<subSubSection id="A2BB6519C313C74DE11E750AFED2FE66" pageId="61" pageNumber="62" type="distribution">
<paragraph id="EA1E3692C313C74DE11E750AFD25FE96" blockId="61.[264,1323,283,430]" pageId="61" pageNumber="62">
<emphasis id="D8D5EA80C313C74DE11E750AFE62FEFD" bold="true" box="[312,472,283,309]" pageId="61" pageNumber="62">Distribution.</emphasis>
Known from Cerro Punta,
<collectingCountry id="92B67602C313C74DE302750DFC3BFEFE" box="[804,897,284,310]" name="Panama" pageId="61" pageNumber="62">Panama</collectingCountry>
(
<figureCitation id="729A2A17C313C74DE3B4750DFC40FEFE" box="[914,1018,284,310]" captionStart="FIGURE 388" captionStartId="98.[264,368,1798,1822]" captionTargetBox="[420,1168,317,1782]" captionTargetId="figure@98.[413,1175,313,1788]" captionTargetPageId="98" captionText="FIGURE 388. Geographical distribution of Scabricollis group in Costa Rica and Panama. Isohypses 1000 m, 2000 m and 3000 m are shown. Areas above 2000 m are grey and those above 3000 m are solid black." httpUri="https://zenodo.org/record/156490/files/figure.png" pageId="61" pageNumber="62">Fig. 388</figureCitation>
). In this locality
<taxonomicName id="2DA14D11C313C74DE4EE750DFEEFFE95" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="D8D5EA80C313C74DE4EE750DFEEFFE95" italics="true" pageId="61" pageNumber="62">S. brevipennis</emphasis>
</taxonomicName>
is sympatric
<taxonomicName id="2DA14D11C313C74DE1D47555FD22FE95" box="[498,664,324,349]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C313C74DE1D47555FD22FE95" box="[498,664,324,349]" italics="true" pageId="61" pageNumber="62">S. scabricollis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="EA1E3692C313C74DE11E757AFED2FE66" blockId="61.[264,1323,283,430]" pageId="61" pageNumber="62">
<emphasis id="D8D5EA80C313C74DE11E757AFDBCFE4D" bold="true" box="[312,518,363,389]" pageId="61" pageNumber="62">Natural History.</emphasis>
<taxonomicName id="2DA14D11C313C74DE23E757DFC8DFE4D" box="[536,823,364,389]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="D8D5EA80C313C74DE23E757DFC8DFE4D" box="[536,823,364,389]" italics="true" pageId="61" pageNumber="62">Seeversiella brevipennis</emphasis>
</taxonomicName>
was collected in forest litter at altitude of
<quantity id="2D599B77C313C74DE12E7585FEDEFE66" box="[264,356,404,430]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="61" pageNumber="62" unit="m" value="2150.0">2150 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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