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<document id= "9080234BA9687A3EF355705950600B48" ID-CLB-Dataset= "34103" ID-DOI= "http://dx.doi.org/10.3897/zookeys.346.6114" ID-GBIF-Dataset= "892cdf91-8f73-426f-9143-b21e3d4df75d" ID-PMC= "PMC3821061" ID-Pensoft-Pub= "1313-2970-346-1" ID-PubMed= "24223483" ModsDocAuthor= "" ModsDocDate= "2013" ModsDocID= "1313-2970-346-1" ModsDocOrigin= "ZooKeys 346" ModsDocTitle= "Revision of the rhinoceros beetle genus Oryctophileurus Kolbe with description of a new species, the male of O. varicosus Prell, and notes on biogeography (Scarabaeoidea, Dynastinae, Phileurini)" checkinTime= "1451246756994" checkinUser= "pensoft" docAuthor= "Perger, Robert & Grossi, Paschoal Coelho" docDate= "2013" docId= "07A4CA6B5A67BAF0B6C869F89EEFD56F" docLanguage= "en" docName= "ZooKeys 346: 1-16" docOrigin= "ZooKeys 346" docSource= "http://dx.doi.org/10.3897/zookeys.346.6114" docTitle= "Oryctophileurus Kolbe 1878" docType= "treatment" docVersion= "5" lastPageNumber= "4" masterDocId= "562FFFC4FFE7E0630407FF9EFF8CFFC5" masterDocTitle= "Revision of the rhinoceros beetle genus Oryctophileurus Kolbe with description of a new species, the male of O. varicosus Prell, and notes on biogeography (Scarabaeoidea, Dynastinae, Phileurini)" masterLastPageNumber= "16" masterPageNumber= "1" pageNumber= "2" updateTime= "1732733777238" updateUser= "ExternalLinkService" >
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<mods:title id= "215450F6700E18482B61F4AC166D4AAC" > Revision of the rhinoceros beetle genus Oryctophileurus Kolbe with description of a new species, the male of O. varicosus Prell, and notes on biogeography (Scarabaeoidea, Dynastinae, Phileurini)</mods:title>
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<mods:namePart id= "47DC659C556DE4835F00B1D8D9D8AF67" > Perger, Robert</mods:namePart>
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<mods:namePart id= "ACDDC852109BC7CCCE6A06D370393E3A" > Grossi, Paschoal Coelho</mods:namePart>
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<treatment id= "07A4CA6B5A67BAF0B6C869F89EEFD56F" ID-GBIF-Taxon= "152049038" LSID= "urn:lsid:plazi:treatment:07A4CA6B5A67BAF0B6C869F89EEFD56F" httpUri= "http://treatment.plazi.org/id/07A4CA6B5A67BAF0B6C869F89EEFD56F" lastPageId= "4" lastPageNumber= "4" pageId= "1" pageNumber= "2" scope_subFamily= "Dynastinae" scope_superFamily= "Scarabaeoidea" scope_tribe= "Phileurini" >
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<paragraph id= "C567D0AC35012E5C1EEB3BE74AD27DF1" pageId= "1" pageNumber= "2" >
<taxonomicName id= "2BC30A7C23D61294CED31687E86BD320" ID-CoL= "63GZP" LSID= "http://species-id.net/wiki/Oryctophileurus" authority= "Kolbe, 1878" authorityName= "Kolbe" authorityYear= "1878" class= "Insecta" family= "Scarabaeidae" genus= "Oryctophileurus" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Oryctophileurus" order= "Coleoptera" pageId= "1" pageNumber= "2" phylum= "Arthropoda" rank= "genus" > Oryctophileurus Kolbe, 1878</taxonomicName>
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</paragraph>
</subSubSection>
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<subSubSection id= "C5D8883050F094E79DD2B8A904EAA9BE" pageId= "1" pageNumber= "2" type= "type species" >
<paragraph id= "DCF9A4284669A39D3B77C69139C3DF77" pageId= "1" pageNumber= "2" > Type species.</paragraph>
<paragraph id= "717BA25B90AC7976F583C5257BD05FCA" pageId= "1" pageNumber= "2" >
<taxonomicName id= "D7F47D9BD731110331B5769559B05D96" class= "Insecta" family= "Scarabaeidae" genus= "Phileurus" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Phileurus nasicornis" order= "Coleoptera" pageId= "1" pageNumber= "2" phylum= "Arthropoda" rank= "species" species= "nasicornis" > Phileurus nasicornis</taxonomicName>
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Burmeister, 1847 (original combination) (Fig. 3).
</paragraph>
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<paragraph id= "CAEE4C81736839257622FB7491CFC59E" pageId= "1" pageNumber= "2" >
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Species of
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<taxonomicName id= "1B3AB57EB3BC475895424A1792C39EBC" class= "Insecta" family= "Scarabaeidae" genus= "Oryctophileurus" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Oryctophileurus" order= "Coleoptera" pageId= "1" pageNumber= "2" phylum= "Arthropoda" rank= "genus" > Oryctophileurus</taxonomicName>
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are distinguished from other
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<taxonomicName id= "54F1C8C32A8D3919312C7245121C27B6" lsidName= "" pageId= "1" pageNumber= "2" rank= "tribe" tribe= "Phileurini" > Phileurini</taxonomicName>
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by a combination of the following apomorphic characters: a single cephalic horn with lateral carina, pronotal cavity with ocellate punctures and two teeth or tubercles close to the anterior pronotal margin, teeth vertically positioned at about the same level as the outer eye margins.
</paragraph>
</subSubSection>
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<subSubSection id= "EE2F5CF8EE1C3155CB076A353EB23F95" lastPageId= "3" lastPageNumber= "4" pageId= "2" pageNumber= "3" type= "taxonomy and discussion" >
<paragraph id= "D47E4BC7EE707EFCD3B461E9535AE59E" pageId= "2" pageNumber= "3" >
<pageBreakToken id= "6DF72B40C626999772A4D9D78F1ADDD7" pageId= "2" pageNumber= "3" start= "start" > Taxonomy</pageBreakToken>
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and discussion.
</paragraph>
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<paragraph id= "6315610CAB3FC0FCF7EBF9EAD7C8C726" pageId= "2" pageNumber= "3" >
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The following characters were cited by
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<bibRefCitation id= "C63FAC7664DB2D172E7EE5D18C262FFE" author= "Endrodi, S" journalOrPublisher= "Dr W. Junk, Dordrecht, Netherlands" pageId= "13" pageNumber= "14" title= "The Dynastinae of the World." year= "1985" >
<normalizedToken id= "6963DD75ED17ADEE269555FF46810664" originalValue= "Endrödi" > Endroedi</normalizedToken>
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(1985)
</bibRefCitation>
for
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<taxonomicName id= "6E0384486AEAFABF03AC0BA94A6876A3" class= "Insecta" family= "Scarabaeidae" genus= "Oryctophileurus" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Oryctophileurus" order= "Coleoptera" pageId= "2" pageNumber= "3" phylum= "Arthropoda" rank= "genus" > Oryctophileurus</taxonomicName>
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,
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<taxonomicName id= "9707E57C94D342A938C8A9F5B0398202" class= "Insecta" family= "Scarabaeidae" genus= "Amblyodus" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Amblyodus" order= "Coleoptera" pageId= "2" pageNumber= "3" phylum= "Arthropoda" rank= "genus" > Amblyodus</taxonomicName>
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,
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<taxonomicName id= "22AE6FBFFC0C0762140609F1592AA021" class= "Insecta" family= "Scarabaeidae" genus= "Microphileurus" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Microphileurus" order= "Coleoptera" pageId= "2" pageNumber= "3" phylum= "Arthropoda" rank= "genus" > Microphileurus</taxonomicName>
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Kolbe,
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<taxonomicName id= "076EECCFF7A6E76F458E2B30A09C1716" class= "Insecta" family= "Scarabaeidae" genus= "Metaphileurus" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Metaphileurus" order= "Coleoptera" pageId= "2" pageNumber= "3" phylum= "Arthropoda" rank= "genus" > Metaphileurus</taxonomicName>
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Kolbe,
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<taxonomicName id= "C7DF9FA055D4C06F75C3DD97EC1835A4" class= "Insecta" family= "Scarabaeidae" genus= "Trioplus" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Trioplus" order= "Coleoptera" pageId= "2" pageNumber= "3" phylum= "Arthropoda" rank= "genus" > Trioplus</taxonomicName>
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Burmeister, and
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<taxonomicName id= "0579F8653859B05C31BC86A58322F05D" class= "Insecta" family= "Scarabaeidae" genus= "Goniophileurus" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Goniophileurus" order= "Coleoptera" pageId= "2" pageNumber= "3" phylum= "Arthropoda" rank= "genus" > Goniophileurus</taxonomicName>
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Kolbe and are here not considered as generic: outer side of mandible tridentate; antenna 10-jointed; elytra with punctate rows; proleg sexually monomorphic, protarsus not thickened in male.
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<taxonomicName id= "840B8A5D38F521DA2AF40F13397B5860" class= "Insecta" family= "Scarabaeidae" genus= "Oryctophileurus" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Oryctophileurus" order= "Coleoptera" pageId= "2" pageNumber= "3" phylum= "Arthropoda" rank= "genus" > Oryctophileurus</taxonomicName>
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and
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<taxonomicName id= "32DF6083BAC2D0A3FF2BB5C670D878A8" class= "Insecta" family= "Scarabaeidae" genus= "Amblyodus" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Amblyodus" order= "Coleoptera" pageId= "2" pageNumber= "3" phylum= "Arthropoda" rank= "genus" > Amblyodus</taxonomicName>
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are distinguished from the other mentioned genera by large horns, a distinctly developed and posteriorly carinate pronotal cavity and four external teeth on the protibia (
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<bibRefCitation id= "109D0E068D87F2F9FD7CC518370C66F0" author= "Endrodi, S" journalOrPublisher= "Dr W. Junk, Dordrecht, Netherlands" pageId= "13" pageNumber= "14" title= "The Dynastinae of the World." year= "1985" >
<normalizedToken id= "BD16CE0BCB55D20D571E273C01187D1A" originalValue= "Endrödi" > Endroedi</normalizedToken>
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1985
</bibRefCitation>
;
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<bibRefCitation id= "FED3179B47F9F53B9DA9676E85AA952C" author= "Ide, S" journalOrPublisher= "Universidade de Sao Paulo, Brasil" pageId= "13" pageNumber= "14" title= "Sistematica e evolucao dos generos neotropicais de Phileurini (Coleoptera: Scarabaeidae: Dynastinae)." year= "1998" > Ide 1998</bibRefCitation>
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). However, in
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<taxonomicName id= "DA4092A2FDF974809E99182EE817B094" class= "Insecta" family= "Scarabaeidae" genus= "Amblyodus" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Amblyodus" order= "Coleoptera" pageId= "2" pageNumber= "3" phylum= "Arthropoda" rank= "genus" > Amblyodus</taxonomicName>
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the two cephalic horns lack the lateral carina, the punctures in the pronotal cavity are predominately fused to short, transverse ridges, and the teeth close to the anterior pronotal margin are absent. Except for the horn number, these generic differences between
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<taxonomicName id= "6CCB171F37BE655301066DFB0D119B42" class= "Insecta" family= "Scarabaeidae" genus= "Oryctophileurus" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Oryctophileurus" order= "Coleoptera" pageId= "2" pageNumber= "3" phylum= "Arthropoda" rank= "genus" > Oryctophileurus</taxonomicName>
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and
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<taxonomicName id= "0E2D61894EF69378536C3B5008AB4D69" class= "Insecta" family= "Scarabaeidae" genus= "Amblyodus" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Amblyodus" order= "Coleoptera" pageId= "2" pageNumber= "3" phylum= "Arthropoda" rank= "genus" > Amblyodus</taxonomicName>
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have not been recognized by previous workers.
</paragraph>
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<paragraph id= "0850416596E1E7CE041B42684BBBAB1B" pageId= "3" pageNumber= "4" >
<pageBreakToken id= "146CB1C1F452213DAE88F98095BE9BA4" pageId= "3" pageNumber= "4" start= "start" > Based</pageBreakToken>
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on the combination of apomorphic characters of
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<taxonomicName id= "85C33878CAB3EDBE98D31C676633EF7B" class= "Insecta" family= "Scarabaeidae" genus= "Oryctophileurus" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Oryctophileurus" order= "Coleoptera" pageId= "3" pageNumber= "4" phylum= "Arthropoda" rank= "genus" > Oryctophileurus</taxonomicName>
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that lacks in
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<taxonomicName id= "F727E95262F29074D6770866B25EE1C8" class= "Insecta" family= "Scarabaeidae" genus= "Amblyodus" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Amblyodus" order= "Coleoptera" pageId= "3" pageNumber= "4" phylum= "Arthropoda" rank= "genus" > Amblyodus</taxonomicName>
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we suggest retaining the genus
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<taxonomicName id= "9026A9F6316981DBC53A1F7BE48F4278" class= "Insecta" family= "Scarabaeidae" genus= "Oryctophileurus" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Oryctophileurus" order= "Coleoptera" pageId= "3" pageNumber= "4" phylum= "Arthropoda" rank= "genus" > Oryctophileurus</taxonomicName>
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.
</paragraph>
</subSubSection>
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<subSubSection id= "36F093A89FB6B709D946AFE869D395E9" lastPageId= "4" lastPageNumber= "5" pageId= "3" pageNumber= "4" type= "key to species of oryctophileurus" >
<paragraph id= "43ED8FEA3F897D01ED811D135319B72D" pageId= "3" pageNumber= "4" >
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Key to species of
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<taxonomicName id= "8C0B8CE48B6415238D1046C2CED1C9DB" class= "Insecta" family= "Scarabaeidae" genus= "Oryctophileurus" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Oryctophileurus" order= "Coleoptera" pageId= "3" pageNumber= "4" phylum= "Arthropoda" rank= "genus" > Oryctophileurus</taxonomicName>
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</paragraph>
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<paragraph id= "D6C0E6A79A547FA5EB0F6BA27FF78878" lastPageId= "4" lastPageNumber= "5" pageId= "3" pageNumber= "4" >
<table id= "14B9E23E6B9421267D850426F79C3C74" lastPageId= "4" lastPageNumber= "5" pageId= "3" pageNumber= "4" >
<tr id= "7081E75A309318A76678A64D57348FC3" pageId= "3" pageNumber= "4" >
<td id= "17FF593C48103F2B3AC28C2753964E77" colspan= "1" pageId= "3" pageNumber= "4" rowspan= "1" > Figs 3B, DFigs 3A, C</td>
<td id= "487900A4F857D667BB676E1BF382244B" colspan= "1" pageId= "3" pageNumber= "4" rowspan= "1" >
<taxonomicName id= "0BEE43C39CE4AE4F8F055A08A0EE6A3C" class= "Insecta" family= "Scarabaeidae" genus= "Oryctophileurus" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Oryctophileurus nasicornis" order= "Coleoptera" pageId= "3" pageNumber= "4" phylum= "Arthropoda" rank= "species" species= "nasicornis" > Oryctophileurus nasicornis</taxonomicName>
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</td>
</tr>
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<tr id= "C495A2A84E602D50BC02EFB2F87B193A" pageId= "3" pageNumber= "4" >
<td id= "F47FDB2261F9DF6A85DCB1E9C020D132" colspan= "1" pageId= "3" pageNumber= "4" rowspan= "1" > Figs 4A, C</td>
<td id= "6DEA0671A8A7486A95BC380C2852F1A7" colspan= "1" pageId= "3" pageNumber= "4" rowspan= "1" >
<taxonomicName id= "8148CDD3536DE625A339FCDBDC9850E8" class= "Insecta" family= "Scarabaeidae" genus= "Oryctophileurus" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Oryctophileurus varicosus" order= "Coleoptera" pageId= "3" pageNumber= "4" phylum= "Arthropoda" rank= "species" species= "varicosus" > Oryctophileurus varicosus</taxonomicName>
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</td>
</tr>
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<tr id= "FDFD12DDA1878FDA124EC79BA89B5ECC" pageId= "3" pageNumber= "4" >
<td id= "1AD0A8B0BD929F9E3B81EC93B69FC00E" colspan= "1" pageId= "3" pageNumber= "4" rowspan= "1" > Fig. 7AFig. 6CFig. 7C</td>
<td id= "9B0CFAF96EA592DA5345E1DCAC7BAF3A" colspan= "1" pageId= "3" pageNumber= "4" rowspan= "1" >
<taxonomicName id= "C55CBFD1825D0EE7258CE5802BFFB02A" class= "Insecta" family= "Scarabaeidae" genus= "Oryctophileurus" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Oryctophileurus armicollis" order= "Coleoptera" pageId= "3" pageNumber= "4" phylum= "Arthropoda" rank= "species" species= "armicollis" > Oryctophileurus armicollis</taxonomicName>
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</td>
</tr>
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<tr id= "0156115F14DF76B11A8BB1F67BCE631B" pageId= "4" pageNumber= "5" >
<td id= "30BA8C641BCD7E7D39B87DE65DAF8456" colspan= "1" pageId= "4" pageNumber= "5" rowspan= "1" >
<pageBreakToken id= "19675088D92F6AB11FF2C45057526568" pageId= "4" pageNumber= "5" start= "start" > Fig</pageBreakToken>
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. 7BFig. 8CFig. 7D
</td>
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<td id= "7507FF881B54E6C31DEB91F8F2806CF6" colspan= "1" pageId= "4" pageNumber= "5" rowspan= "1" >
<taxonomicName id= "7B63182F6E9CA12DF3BA5530B9D06A21" class= "Insecta" family= "Scarabaeidae" genus= "Oryctophileurus" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Oryctophileurus guerrai" order= "Coleoptera" pageId= "4" pageNumber= "5" phylum= "Arthropoda" rank= "species" species= "guerrai" > Oryctophileurus guerrai</taxonomicName>
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</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>
