Revision of the rhinoceros beetle genus Oryctophileurus Kolbe with description of a new species, the male of O. varicosus Prell, and notes on biogeography (Scarabaeoidea, Dynastinae, Phileurini)AuthorPerger, RobertAuthorGrossi, Paschoal CoelhotextZooKeys2013346116http://dx.doi.org/10.3897/zookeys.346.6114journal articlehttp://dx.doi.org/10.3897/zookeys.346.61141313-2970-346-1Oryctophileurus Kolbe, 1878Type species.Phileurus nasicornis
Burmeister, 1847 (original combination) (Fig. 3).
Species of
Oryctophileurus
are distinguished from other
Phileurini
by a combination of the following apomorphic characters: a single cephalic horn with lateral carina, pronotal cavity with ocellate punctures and two teeth or tubercles close to the anterior pronotal margin, teeth vertically positioned at about the same level as the outer eye margins.
Taxonomy
and discussion.
The following characters were cited by
Endroedi
(1985)
for
Oryctophileurus
,
Amblyodus
,
Microphileurus
Kolbe,
Metaphileurus
Kolbe,
Trioplus
Burmeister, and
Goniophileurus
Kolbe and are here not considered as generic: outer side of mandible tridentate; antenna 10-jointed; elytra with punctate rows; proleg sexually monomorphic, protarsus not thickened in male.
Oryctophileurus
and
Amblyodus
are distinguished from the other mentioned genera by large horns, a distinctly developed and posteriorly carinate pronotal cavity and four external teeth on the protibia (
Endroedi
1985
;
Ide 1998
). However, in
Amblyodus
the two cephalic horns lack the lateral carina, the punctures in the pronotal cavity are predominately fused to short, transverse ridges, and the teeth close to the anterior pronotal margin are absent. Except for the horn number, these generic differences between
Oryctophileurus
and
Amblyodus
have not been recognized by previous workers.
Based
on the combination of apomorphic characters of
Oryctophileurus
that lacks in
Amblyodus
we suggest retaining the genus
Oryctophileurus
.
Key to species of
Oryctophileurus