<documentid="FF1EA3E1D1E46E58BF1E93BD5D8618EA"ID-DOI="10.11646/zootaxa.5514.5.1"ID-ISSN="1175-5326"ID-Zenodo-Dep="13914727"ID-ZooBank="615D8E3D-9216-4CCC-9A6D-A6AFDA4C6FE3"IM.bibliography_approvedBy="juliana"IM.illustrations_approvedBy="juliana"IM.materialsCitations_approvedBy="juliana"IM.metadata_approvedBy="juliana"IM.tables_approvedBy="juliana"IM.taxonomicNames_approvedBy="felipe"IM.treatmentCitations_approvedBy="felipe"IM.treatments_approvedBy="juliana"checkinTime="1728581305616"checkinUser="plazi"docAuthor="Barroso, Aylin Alegre & Pérez-González, Abel"docDate="2024"docId="03DB4C6DFFCCFFAB24FBF874FEC363B1"docLanguage="en"docName="zootaxa.5514.5.1.pdf"docOrigin="Zootaxa 5514 (5)"docSource="http://dx.doi.org/10.11646/zootaxa.5514.5.1"docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article"docStyleId="647186512141C8FC8976D5BCC54AEB7D"docStyleName="Zootaxa.2013-.journal_article"docStyleVersion="9"docTitle="Caribbiantes barbai Barroso & Pérez-González, 2024, sp. nov."docType="treatment"docUuid="06118480-8423-403D-8B64-336E3F6C73E6"docUuidSource="ZooBank"docVersion="5"lastPageNumber="424"masterDocId="FFE23415FFDCFFBC246CFF7AFFB56078"masterDocTitle="Two new species of Caribbiantes, with the redescription of the type species and a review of male genital patterns in Antillean Stenostygninae (Opiliones: Laniatores: Biantidae)"masterLastPageNumber="430"masterPageNumber="401"pageNumber="417"updateTime="1728650480329"updateUser="ExternalLinkService"zenodo-license-document="CLOSED">
<mods:titleid="D81874031E657C87645B313C373D8630">Two new species of Caribbiantes, with the redescription of the type species and a review of male genital patterns in Antillean Stenostygninae (Opiliones: Laniatores: Biantidae)</mods:title>
<mods:affiliationid="DE9863588E804BE392FDBE23F5CB3600">Museum of Comparative Zoology, Department of Organismic and Evolutionary Biology, Harvard University, 26 Oxford Street, Cambridge, MA, 02138, USA</mods:affiliation>
<mods:affiliationid="E65487F535BDB3FAC5F909CEC249BFE8">Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ”, Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Av. Ángel Gallardo 470, C 1405 DJR, Buenos Aires, Argentina</mods:affiliation>
<taxonomicNameid="4C7286F8FFCCFFAC24FBF874FE326751"authorityName="Barroso & Pérez-González"authorityYear="2024"box="[151,391,1806,1833]"class="Arachnida"family="Biantidae"genus="Caribbiantes"kingdom="Animalia"order="Opiliones"pageId="16"pageNumber="417"phylum="Arthropoda"rank="species"species="barbai"status="sp. nov.">
<figureCitationid="1349E1FEFFCCFFAC24F3F82DFEA26709"box="[159,279,1879,1905]"captionStart-0="FIGURE 9"captionStart-1="FIGURE 10"captionStart-2="FIGURE 11"captionStartId-0="17.[151,250,1757,1781]"captionStartId-1="18.[151,250,890,914]"captionStartId-2="21.[151,250,1761,1785]"captionTargetBox-0="[158,1436,355,1724]"captionTargetBox-1="[151,1435,181,866]"captionTargetBox-2="[163,1432,191,1737]"captionTargetId-0="figure-89@17.[151,1436,345,1734]"captionTargetId-1="figure-19@18.[151,1435,181,866]"captionTargetId-2="figure-18@21.[155,1432,181,1737]"captionTargetPageId-0="17"captionTargetPageId-1="18"captionTargetPageId-2="21"captionText-0="FIGURE 9. Caribbiantes barbai sp. nov., habitus of male holotype (CZACC 3.4380): A–B. Dorsal view; C–D. Lateral view. Scale bars = 1 mm."captionText-1="FIGURE 10. Glandular structure of male metatarsus III in Caribbiantes barbai sp. nov. A. Light microscope photo, holotype male (CZACC 3. 4380), general view of the whole metatarsus III (divided into four quarters to facilitate comparisons); B–E. Scanning electron micrographs, male (CZACC 3.3392); B. View of the glandular swelling, dashed line delineates the form of the ventral, buttonhole-like groove; C. Close-up of the groove delimited in B, note the depth of the groove; D. Closer zoom into the groove, white arrow shows a seta (long paint brush, basally striated and twisted), small arrowheads indicate glandular openings (pores) regularly arranged around the setae; E. Detail of the integument and pores (arrowheads) of the ventral groove."captionText-2="FIGURE 11. Caribbiantes barbai sp. nov., entire (C) anddistal part (A–B, D–J) of penis,A–F.Drawings with optical microscope, male holotype (CZACC 3.4380): A. Dorsal view; B. Dorsal view, expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view, expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm. G–J. Scanning electron micrographs (male CZACC 3.3392): G. Dorsal view, expanded; H. Apical view, expanded; I. Left lateral view, expanded; J. Ventral view, expanded.Abbreviations: AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae."figureDoi-0="http://doi.org/10.5281/zenodo.13914747"figureDoi-1="http://doi.org/10.5281/zenodo.13914749"figureDoi-2="http://doi.org/10.5281/zenodo.13914751"httpUri-0="https://zenodo.org/record/13914747/files/figure.png"httpUri-1="https://zenodo.org/record/13914749/files/figure.png"httpUri-2="https://zenodo.org/record/13914751/files/figure.png"pageId="16"pageNumber="417">Figs. 9–11</figureCitation>
<materialsCitationid="3B1AF726FFCCFFAD254BF8E5FC8A60C9"collectingDate="2009-01-15"collectionCode="PNAH"collectorName="R. Barba"country="Cuba"county="Baracoa Municipality"elevation="500"lastPageId="17"lastPageNumber="418"latitude="20.4575"location="Monte Iberia"longLatPrecision="21"longitude="-74.72639"municipality="Alejandro de Humboldt National Park"pageId="16"pageNumber="417"specimenCode="CZACC 3.4380"specimenCount="1"specimenCount-male="1"stateProvince="Guantanamo province"typeStatus="holotype">
<specimenCodeid="DBD45500FFCCFFAC25B2F8E5FD2167C1"box="[478,660,1951,1977]"collectionCode="CZACC"country="Cuba"httpUri="http://grbio.org/cool/b49z-cz3a"name="Coleccion Zoologia, Academia de Ciencias de Cuba"pageId="16"pageNumber="417">CZACC 3.4380</specimenCode>
<locationid="8EADABA0FFCCFFAC27FFF8E5FB9367C1"LSID="urn:lsid:plazi:treatment:03DB4C6DFFCCFFAB24FBF874FEC363B1:8EADABA0FFCCFFAC27FFF8E5FB9367C1"box="[915,1062,1951,1977]"country="Cuba"county="Baracoa Municipality"latitude="20.4575"longLatPrecision="21"longitude="-74.72639"municipality="Alejandro de Humboldt National Park"name="Monte Iberia"pageId="16"pageNumber="417"stateProvince="Guantanamo province">Monte Iberia</location>
,
<locationid="8EADABA0FFCCFFAC2043F8E5FF7F67A5"LSID="urn:lsid:plazi:treatment:03DB4C6DFFCCFFAB24FBF874FEC363B1:8EADABA0FFCCFFAC2043F8E5FF7F67A5"country="Cuba"county="Baracoa Municipality"latitude="20.4575"longLatPrecision="21"longitude="-74.72639"municipality="Alejandro de Humboldt National Park"name="Alejandro de Humboldt National Park"pageId="16"pageNumber="417"stateProvince="Guantanamo province">Alejandro de Humboldt National Park</location>
<materialsCitationid="3B1AF726FFCDFFAD2725FFEDFE8560AD"collectingDate="2009-01-15"collectionCode="CZACC"collectorName="R. Barba"country="Cuba"county="Baracoa Municipality"elevation="500"latitude="20.4575"location="Monte Iberia"longLatPrecision="21"longitude="-74.72639"municipality="Alejandro de Humboldt National Park"pageId="17"pageNumber="418"specimenCode="CZACC 3.4380"specimenCount="1"specimenCount-female="1"stateProvince="Guantanamo"typeStatus="paratype">
<specimenCodeid="DBD45500FFCDFFAD206EFFEDFB0F60C9"box="[1026,1210,151,177]"collectionCode="CZACC"country="Cuba"httpUri="http://grbio.org/cool/b49z-cz3a"name="Coleccion Zoologia, Academia de Ciencias de Cuba"pageId="17"pageNumber="418">CZACC 3.4380</specimenCode>
<materialsCitationid="3B1AF726FFCDFFAD2550FFC1FCDF60AD"box="[316,874,187,213]"collectingDate="2009-01-15"collectionCode="CZACC"collectorName="R. Barba"country="Cuba"county="Baracoa Municipality"elevation="500"latitude="20.4575"location="Trail"longLatPrecision="21"longitude="-74.72639"municipality="Alejandro de Humboldt National Park"pageId="17"pageNumber="418"specimenCode="CZACC 3.3391"specimenCount="1"specimenCount-male="1"stateProvince="Guantanamo"typeStatus="paratype">
<specimenCodeid="DBD45500FFCDFFAD251AFFC1FD9A60AD"box="[374,559,187,213]"collectionCode="CZACC"country="Cuba"httpUri="http://grbio.org/cool/b49z-cz3a"name="Coleccion Zoologia, Academia de Ciencias de Cuba"pageId="17"pageNumber="418">CZACC 3.3391</specimenCode>
<specimenCodeid="DBD45500FFCDFFAD27DCFFC1FBDD60AD"box="[944,1128,187,213]"collectionCode="CZACC"country="Cuba"httpUri="http://grbio.org/cool/b49z-cz3a"name="Coleccion Zoologia, Academia de Ciencias de Cuba"pageId="17"pageNumber="418">CZACC 3.3393</specimenCode>
<specimenCodeid="DBD45500FFCDFFAD27B4FFA5FB266081"box="[984,1171,223,249]"collectionCode="CZACC"country="Cuba"httpUri="http://grbio.org/cool/b49z-cz3a"name="Coleccion Zoologia, Academia de Ciencias de Cuba"pageId="17"pageNumber="418">CZACC 3.3392</specimenCode>
<taxonomicNameid="4C7286F8FFCDFFAD2577F9A7FE5F668D"authorityName="Barroso & Pérez-González"authorityYear="2024"box="[283,490,1757,1781]"class="Arachnida"family="Biantidae"genus="Caribbiantes"kingdom="Animalia"order="Opiliones"pageId="17"pageNumber="418"phylum="Arthropoda"rank="species"species="barbai"status="sp. nov.">
<taxonomicNameid="4C7286F8FFCEFFAE2699FC00FC7063EA"authorityName="Barroso & Pérez-González"authorityYear="2024"box="[757,965,890,914]"class="Arachnida"family="Biantidae"genus="Caribbiantes"kingdom="Animalia"order="Opiliones"pageId="18"pageNumber="419"phylum="Arthropoda"rank="species"species="barbai"status="sp. nov.">
A. Light microscope photo, holotype male (CZACC 3. 4380), general view of the whole metatarsus III (divided into four quarters to facilitate comparisons); B–E. Scanning electron micrographs, male (CZACC 3.3392); B. View of the glandular swelling, dashed line delineates the form of the ventral, buttonhole-like groove; C. Close-up of the groove delimited in B, note the depth of the groove; D. Closer zoom into the groove, white arrow shows a seta (long paint brush, basally striated and twisted), small arrowheads indicate glandular openings (pores) regularly arranged around the setae; E. Detail of the integument and pores (arrowheads) of the ventral groove.
<taxonomicNameid="4C7286F8FFCEFFAE25D7FA7FFD866567"authorityName="Barroso & Pérez-González"authorityYear="2024"box="[443,563,1285,1311]"class="Arachnida"family="Biantidae"genus="Caribbiantes"kingdom="Animalia"order="Opiliones"pageId="18"pageNumber="419"phylum="Arthropoda"rank="species"species="obtusus">
<taxonomicNameid="4C7286F8FFCEFFAE24FBFA53FEBA653B"authorityName="Barroso & Pérez-González"authorityYear="2024"box="[151,271,1321,1347]"class="Arachnida"family="Biantidae"genus="Caribbiantes"kingdom="Animalia"order="Opiliones"pageId="18"pageNumber="419"phylum="Arthropoda"rank="species"species="obtusus">
<taxonomicNameid="4C7286F8FFCEFFAE2612FA37FD42651F"authorityName="Barroso & Pérez-González"authorityYear="2024"box="[638,759,1357,1383]"class="Arachnida"family="Biantidae"genus="Caribbiantes"kingdom="Animalia"order="Opiliones"pageId="18"pageNumber="419"phylum="Arthropoda"rank="species"species="obtusus">
<taxonomicNameid="4C7286F8FFCEFFAE21EFFA0BFF5865D7"authorityName="Barroso & Pérez-González"authorityYear="2024"class="Arachnida"family="Biantidae"genus="Caribbiantes"kingdom="Animalia"order="Opiliones"pageId="18"pageNumber="419"phylum="Arthropoda"rank="species"species="obtusus">
<taxonomicNameid="4C7286F8FFCEFFAE24FBFAC3FEBA65AB"authorityName="Barroso & Pérez-González"authorityYear="2024"box="[151,271,1465,1491]"class="Arachnida"family="Biantidae"genus="Caribbiantes"kingdom="Animalia"order="Opiliones"pageId="18"pageNumber="419"phylum="Arthropoda"rank="species"species="obtusus">C. obtusus</taxonomicName>
,
</emphasis>
in which femora are unarmed. Swollen region of metatarsus III of males is located in the third (subdistal) quarter of the segment but it reaches the second and fourth quarters too (
<figureCitationid="1349E1FEFFCEFFAE27BEFAA7FB8C658F"box="[978,1081,1501,1527]"captionStart="FIGURE 10"captionStartId="18.[151,250,890,914]"captionTargetBox="[151,1435,181,866]"captionTargetId="figure-19@18.[151,1435,181,866]"captionTargetPageId="18"captionText="FIGURE 10. Glandular structure of male metatarsus III in Caribbiantes barbai sp. nov. A. Light microscope photo, holotype male (CZACC 3. 4380), general view of the whole metatarsus III (divided into four quarters to facilitate comparisons); B–E. Scanning electron micrographs, male (CZACC 3.3392); B. View of the glandular swelling, dashed line delineates the form of the ventral, buttonhole-like groove; C. Close-up of the groove delimited in B, note the depth of the groove; D. Closer zoom into the groove, white arrow shows a seta (long paint brush, basally striated and twisted), small arrowheads indicate glandular openings (pores) regularly arranged around the setae; E. Detail of the integument and pores (arrowheads) of the ventral groove."figureDoi="http://doi.org/10.5281/zenodo.13914749"httpUri="https://zenodo.org/record/13914749/files/figure.png"pageId="18"pageNumber="419">Fig. 10A</figureCitation>
<taxonomicNameid="4C7286F8FFCEFFAE2521F97BFE736663"authorityName="Barroso & Pérez-González"authorityYear="2024"box="[333,454,1537,1563]"class="Arachnida"family="Biantidae"genus="Caribbiantes"kingdom="Animalia"order="Opiliones"pageId="18"pageNumber="419"phylum="Arthropoda"rank="species"species="obtusus">C. obtusus</taxonomicName>
,
</emphasis>
centered between the third and fourth distal quarter, and not reaching the second basal quarter) (
<figureCitationid="1349E1FEFFCEFFAE2493F95FFED16647"box="[255,356,1573,1599]"captionStart="FIGURE 3"captionStartId="6.[151,250,890,914]"captionTargetBox="[151,1435,181,866]"captionTargetId="figure-19@6.[151,1435,181,866]"captionTargetPageId="6"captionText="FIGURE 3. Glandular structure of male metatarsus III in Caribbiantes cubanus. A. Light microscope photo, male (CZACC 3.4372), general view of the whole metatarsus III (divided into four quarters to facilitate comparisons); B–E., Scanning electron micrographs, male (CZACC 3.3266): B. View of the glandular swelling, dashed line delineates the form of the ventral, buttonhole-like groove; C. Close-up of the groove delimited in B, note its depth; D. Closer zoom into the groove, white arrow indicates a seta (long paint brush, basally striated and twisted), small arrowheads point to glandular openings (pores) irregularly arranged around the setae; E. Detail of the integument of the ventral groove."figureDoi="http://doi.org/10.5281/zenodo.13914733"httpUri="https://zenodo.org/record/13914733/files/figure.png"pageId="18"pageNumber="419">Figs. 3A</figureCitation>
;
<figureCitationid="1349E1FEFFCEFFAE251CF95FFE236647"box="[368,406,1573,1599]"captionStart="FIGURE 7"captionStartId="14.[151,250,1774,1798]"captionTargetBox="[151,1435,1065,1750]"captionTargetId="figure-443@14.[151,1435,1065,1750]"captionTargetPageId="14"captionText="FIGURE 7. Glandular structure of male metatarsus III in Caribbiantes obtusus sp. nov. (male CZACC 3.4379). A. Light microscope photo, general view of the whole metatarsus III (divided into four quarters to facilitate comparisons); B–E. Scanning electron micrographs: B. View of the glandular swelling, dashed line delineates the form of the ventral, buttonhole-like groove; C. Close-up of the groove delimited in B, note the depth of the groove; D. Closer zoom into the groove, arrows show setae (long paint brush, basally striated and twisted); E. Integument of the ventral groove, white arrowheads indicate glandular openings (pores) irregularly arranged around the setae."figureDoi="http://doi.org/10.5281/zenodo.13914743"httpUri="https://zenodo.org/record/13914743/files/figure.png"pageId="18"pageNumber="419">7A</figureCitation>
). The ventral groove may have 26–32 brush-like setae. Tarsal formula 7(3):12–15(4–6):8:8.
<taxonomicNameid="4C7286F8FFCEFFAE24FBF933FEC9661B"authorityName="Barroso & Pérez-González"authorityYear="2024"box="[151,380,1609,1635]"class="Arachnida"family="Biantidae"genus="Caribbiantes"kingdom="Animalia"order="Opiliones"pageId="18"pageNumber="419"phylum="Arthropoda"rank="species"species="barbai"status="sp. nov.">
<taxonomicNameid="4C7286F8FFCEFFAE2046F933FB14661B"authorityName="Barroso & Pérez-González"authorityYear="2024"box="[1066,1185,1609,1635]"class="Arachnida"family="Biantidae"genus="Caribbiantes"kingdom="Animalia"order="Opiliones"pageId="18"pageNumber="419"phylum="Arthropoda"rank="species"species="obtusus">
<taxonomicNameid="4C7286F8FFCEFFAE27EEF9EBFC4D66D3"authorityName="Barroso & Pérez-González"authorityYear="2024"box="[898,1016,1681,1707]"class="Arachnida"family="Biantidae"genus="Caribbiantes"kingdom="Animalia"order="Opiliones"pageId="18"pageNumber="419"phylum="Arthropoda"rank="species"species="obtusus">
<figureCitationid="1349E1FEFFCEFFAE206AF9EBFB1866D3"box="[1030,1197,1680,1707]"captionStart="FIGURE 5"captionStartId="10.[151,250,1441,1465]"captionTargetBox="[190,1407,181,1416]"captionTargetId="figure-19@10.[180,1407,181,1416]"captionTargetPageId="10"captionText="FIGURE 5. Caribbiantes cubanus, entire (C) and distal part (A–B, D–I) of penis,A–F. Drawings with optical microscope (male CZACC 3.3379): A. Dorsal view; B. Dorsal view expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm; G–I. Scanning electron micrographs (male CZACC 3.3266): G. Dorsal view; H. Left lateral view; I. Ventral view.Abbreviations:AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae."figureDoi="http://doi.org/10.5281/zenodo.13914737"httpUri="https://zenodo.org/record/13914737/files/figure.png"pageId="18"pageNumber="419">Figs. 5A, G, H</figureCitation>
;
<figureCitationid="1349E1FEFFCEFFAE20D5F9EBFB4866D3"box="[1209,1277,1681,1707]"captionStart="FIGURE 8"captionStartId="15.[151,250,1835,1859]"captionTargetBox="[197,1401,189,1812]"captionTargetId="figure-18@15.[186,1401,181,1812]"captionTargetPageId="15"captionText="FIGURE 8. Caribbiantes obtusus sp. nov., entire (C) and distal part (A–B, D–J) of penis. A–F: Drawings with optical microscope, male holotype (CZACC 3.3384), A. Dorsal view; B. Dorsal view, expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm. G–J. Scanning electron microscopy male CZACC 3.4379). G. Dorsal view; H. Apical view (apical portion of a conductor colored in pink); I. Ventrolateral view; J. Ventral view. Abbreviations: AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae."figureDoi="http://doi.org/10.5281/zenodo.13914745"httpUri="https://zenodo.org/record/13914745/files/figure.png"pageId="18"pageNumber="419">8A, G</figureCitation>
<taxonomicNameid="4C7286F8FFCEFFAE214BF9CFFA2866B7"authorityName="Barroso & Pérez-González"authorityYear="2024"box="[1319,1437,1717,1743]"class="Arachnida"family="Biantidae"genus="Caribbiantes"kingdom="Animalia"order="Opiliones"pageId="18"pageNumber="419"phylum="Arthropoda"rank="species"species="obtusus">
<figureCitationid="1349E1FEFFCEFFAE24F3F9A3FEB8668B"box="[159,269,1753,1779]"captionStart="FIGURE 11"captionStartId="21.[151,250,1761,1785]"captionTargetBox="[163,1432,191,1737]"captionTargetId="figure-18@21.[155,1432,181,1737]"captionTargetPageId="21"captionText="FIGURE 11. Caribbiantes barbai sp. nov., entire (C) anddistal part (A–B, D–J) of penis,A–F.Drawings with optical microscope, male holotype (CZACC 3.4380): A. Dorsal view; B. Dorsal view, expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view, expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm. G–J. Scanning electron micrographs (male CZACC 3.3392): G. Dorsal view, expanded; H. Apical view, expanded; I. Left lateral view, expanded; J. Ventral view, expanded.Abbreviations: AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae."figureDoi="http://doi.org/10.5281/zenodo.13914751"httpUri="https://zenodo.org/record/13914751/files/figure.png"pageId="18"pageNumber="419">Figs. 11E</figureCitation>
;
<figureCitationid="1349E1FEFFCEFFAE2574F9A2FE8F668B"box="[280,314,1752,1779]"captionStart="FIGURE 5"captionStartId="10.[151,250,1441,1465]"captionTargetBox="[190,1407,181,1416]"captionTargetId="figure-19@10.[180,1407,181,1416]"captionTargetPageId="10"captionText="FIGURE 5. Caribbiantes cubanus, entire (C) and distal part (A–B, D–I) of penis,A–F. Drawings with optical microscope (male CZACC 3.3379): A. Dorsal view; B. Dorsal view expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm; G–I. Scanning electron micrographs (male CZACC 3.3266): G. Dorsal view; H. Left lateral view; I. Ventral view.Abbreviations:AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae."figureDoi="http://doi.org/10.5281/zenodo.13914737"httpUri="https://zenodo.org/record/13914737/files/figure.png"pageId="18"pageNumber="419">5E</figureCitation>
;
<figureCitationid="1349E1FEFFCEFFAE2529F9A3FEDD668B"box="[325,360,1753,1779]"captionStart="FIGURE 8"captionStartId="15.[151,250,1835,1859]"captionTargetBox="[197,1401,189,1812]"captionTargetId="figure-18@15.[186,1401,181,1812]"captionTargetPageId="15"captionText="FIGURE 8. Caribbiantes obtusus sp. nov., entire (C) and distal part (A–B, D–J) of penis. A–F: Drawings with optical microscope, male holotype (CZACC 3.3384), A. Dorsal view; B. Dorsal view, expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm. G–J. Scanning electron microscopy male CZACC 3.4379). G. Dorsal view; H. Apical view (apical portion of a conductor colored in pink); I. Ventrolateral view; J. Ventral view. Abbreviations: AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae."figureDoi="http://doi.org/10.5281/zenodo.13914745"httpUri="https://zenodo.org/record/13914745/files/figure.png"pageId="18"pageNumber="419">8E</figureCitation>
). The two conductors, fused at the base, bear apical laminar lobes, ventrally projected (
<figureCitationid="1349E1FEFFCEFFAE2147F9A3FF0E676F"captionStart="FIGURE 11"captionStartId="21.[151,250,1761,1785]"captionTargetBox="[163,1432,191,1737]"captionTargetId="figure-18@21.[155,1432,181,1737]"captionTargetPageId="21"captionText="FIGURE 11. Caribbiantes barbai sp. nov., entire (C) anddistal part (A–B, D–J) of penis,A–F.Drawings with optical microscope, male holotype (CZACC 3.4380): A. Dorsal view; B. Dorsal view, expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view, expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm. G–J. Scanning electron micrographs (male CZACC 3.3392): G. Dorsal view, expanded; H. Apical view, expanded; I. Left lateral view, expanded; J. Ventral view, expanded.Abbreviations: AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae."figureDoi="http://doi.org/10.5281/zenodo.13914751"httpUri="https://zenodo.org/record/13914751/files/figure.png"pageId="18"pageNumber="419">Figs. 11E, I, J</figureCitation>
), although they are much more rounded and dilated than in
<taxonomicNameid="4C7286F8FFCEFFAE214AF987FA28676F"authorityName="Barroso & Pérez-González"authorityYear="2024"box="[1318,1437,1789,1815]"class="Arachnida"family="Biantidae"genus="Caribbiantes"kingdom="Animalia"order="Opiliones"pageId="18"pageNumber="419"phylum="Arthropoda"rank="species"species="obtusus">
<figureCitationid="1349E1FEFFCEFFAE24F3F85BFEB66743"box="[159,259,1824,1851]"captionStart="FIGURE 5"captionStartId="10.[151,250,1441,1465]"captionTargetBox="[190,1407,181,1416]"captionTargetId="figure-19@10.[180,1407,181,1416]"captionTargetPageId="10"captionText="FIGURE 5. Caribbiantes cubanus, entire (C) and distal part (A–B, D–I) of penis,A–F. Drawings with optical microscope (male CZACC 3.3379): A. Dorsal view; B. Dorsal view expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm; G–I. Scanning electron micrographs (male CZACC 3.3266): G. Dorsal view; H. Left lateral view; I. Ventral view.Abbreviations:AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae."figureDoi="http://doi.org/10.5281/zenodo.13914737"httpUri="https://zenodo.org/record/13914737/files/figure.png"pageId="18"pageNumber="419">Figs. 5E</figureCitation>
;
<figureCitationid="1349E1FEFFCEFFAE257DF85BFEED6743"box="[273,344,1825,1851]"captionStart="FIGURE 8"captionStartId="15.[151,250,1835,1859]"captionTargetBox="[197,1401,189,1812]"captionTargetId="figure-18@15.[186,1401,181,1812]"captionTargetPageId="15"captionText="FIGURE 8. Caribbiantes obtusus sp. nov., entire (C) and distal part (A–B, D–J) of penis. A–F: Drawings with optical microscope, male holotype (CZACC 3.3384), A. Dorsal view; B. Dorsal view, expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm. G–J. Scanning electron microscopy male CZACC 3.4379). G. Dorsal view; H. Apical view (apical portion of a conductor colored in pink); I. Ventrolateral view; J. Ventral view. Abbreviations: AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae."figureDoi="http://doi.org/10.5281/zenodo.13914745"httpUri="https://zenodo.org/record/13914745/files/figure.png"pageId="18"pageNumber="419">8E, H</figureCitation>
<taxonomicNameid="4C7286F8FFCEFFAE2595F83FFDC56727"authorityName="Barroso & Pérez-González"authorityYear="2024"box="[505,624,1861,1887]"class="Arachnida"family="Biantidae"genus="Caribbiantes"kingdom="Animalia"order="Opiliones"pageId="18"pageNumber="419"phylum="Arthropoda"rank="species"species="obtusus">
<figureCitationid="1349E1FEFFCEFFAE2613F83FFD566727"box="[639,739,1860,1887]"captionStart="FIGURE 5"captionStartId="10.[151,250,1441,1465]"captionTargetBox="[190,1407,181,1416]"captionTargetId="figure-19@10.[180,1407,181,1416]"captionTargetPageId="10"captionText="FIGURE 5. Caribbiantes cubanus, entire (C) and distal part (A–B, D–I) of penis,A–F. Drawings with optical microscope (male CZACC 3.3379): A. Dorsal view; B. Dorsal view expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm; G–I. Scanning electron micrographs (male CZACC 3.3266): G. Dorsal view; H. Left lateral view; I. Ventral view.Abbreviations:AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae."figureDoi="http://doi.org/10.5281/zenodo.13914737"httpUri="https://zenodo.org/record/13914737/files/figure.png"pageId="18"pageNumber="419">Figs. 5D</figureCitation>
;
<figureCitationid="1349E1FEFFCEFFAE2683F83FFCA06727"box="[751,789,1861,1887]"captionStart="FIGURE 8"captionStartId="15.[151,250,1835,1859]"captionTargetBox="[197,1401,189,1812]"captionTargetId="figure-18@15.[186,1401,181,1812]"captionTargetPageId="15"captionText="FIGURE 8. Caribbiantes obtusus sp. nov., entire (C) and distal part (A–B, D–J) of penis. A–F: Drawings with optical microscope, male holotype (CZACC 3.3384), A. Dorsal view; B. Dorsal view, expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm. G–J. Scanning electron microscopy male CZACC 3.4379). G. Dorsal view; H. Apical view (apical portion of a conductor colored in pink); I. Ventrolateral view; J. Ventral view. Abbreviations: AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae."figureDoi="http://doi.org/10.5281/zenodo.13914745"httpUri="https://zenodo.org/record/13914745/files/figure.png"pageId="18"pageNumber="419">8D</figureCitation>
<tableCitationid="C6F0C8C0FFCEFFAE2515F8CBFE7967B3"box="[377,460,1969,1995]"captionStart="TABLE 3"captionStartId="19.[152,244,151,175]"captionTargetPageId="19"captionText="TABLE 3. Measurements (in mm) of the male holotype (H) and paratypes (P) of Caribbiantes barbai sp. nov.."httpUri="http://table.plazi.org/id/DF0DADF3FFCFFFAF24F4FFEDFA8B60D7"pageId="18"pageNumber="419"tableUuid="DF0DADF3FFCFFFAF24F4FFEDFA8B60D7">Table 3</tableCitation>
. Measurements (in mm) of the male holotype (H) and paratypes (P) of
<taxonomicNameid="4C7286F8FFCFFFAF206EFFEDFB6E60D7"authorityName="Barroso & Pérez-González"authorityYear="2024"box="[1026,1243,151,175]"class="Arachnida"family="Biantidae"genus="Caribbiantes"kingdom="Animalia"order="Opiliones"pageId="19"pageNumber="420"phylum="Arthropoda"rank="species"species="barbai"status="sp. nov.">
). Four well-defined mesotergal areas, densely granulous, with wide sulci between them. Area I medially constricted, divided into triangular halves by a short and shallow median groove, each half bearing two slightly taller tubercles (
Coxae covered by setiferous granules. Coxa I with an anterior row of small setiferous tubercles. Coxae III–IV with an anterior row of small tubercles that form small bridges with the posterior margin of the preceding coxa. Free sternites with a row of small tubercles. Genital operculum short and narrow.
Basichelicerite with robust and well-marked bulla. Hand strongly swollen (hypertelic), with scattered setiferous granules, and longer setae towards the distal border (
). Fixed finger distally with five minute teeth, movable finger with a strong, blunt basal tubercle, followed by a smooth gap, a “dome-like” serrated curvature of the internal finger edge and distally 7–8 small teeth. Cheliceral fingers curved, leaving a wide basal gap when closed.
: Elongated, with large spines concentrated on tibia and tarsus. Coxa elongated, dorsally with one small ectoproximal acute tubercle, a group of four mesoproximal tubercles on a small protuberance, and one small distal tubercle; ventrally, a group of 3–4 proximal tubercles, followed by a distal setiferous tubercle. Trochanter dorsally granulous, ventrally with a proximal setiferous tubercle, followed by one subdistal setiferous tubercle and one distal granule. Femur with six spaced small setiferous granules on the dorsal surface, and four spaced small setiferous granules on the ventral surface. Patella dilated on its distal third, dorsally granulous and with a strong mesodistal spine. Tibia and tarsus dorsally granulous, more densely in tibia. Both segments ventrally armed with strong spines as follows: tibia ectal IIIi (1<2=3>4), tibia mesal IIII (1<2<3<4); tarsus ectal IiIi ((1>3)>2 =4)), tarsus mesal IiIi ((1>3)>2=4)). Tibia also with a ventral row of 4–5 small medial setiferous tubercles. The ectodistal spine on the tibia is very close to the preceding one, giving the appearance of being fused at their bases; the mesal spines on the tibia are equidistant from each other, resulting in similar-sized gaps in the spine series, as in
<taxonomicNameid="4C7286F8FFC8FFA824FBFBFBFEBA64E2"authorityName="Barroso & Pérez-González"authorityYear="2024"box="[151,271,1152,1179]"class="Arachnida"family="Biantidae"genus="Caribbiantes"kingdom="Animalia"order="Opiliones"pageId="20"pageNumber="421"phylum="Arthropoda"rank="species"species="obtusus"status="sp. nov.">
Covered by fine setiferous granules. Coxa IV covered by setiferous granules. Trochanters II–IV dorsally with granules. Femora II–IV with a distal spiniform apophysis (
). Patella IV with a medial dorsodistal small acute process. Metatarsus III spindle-shaped, with the swollen sector located in the third (subdistal) quarter of the segment but also reaching the second and fourth quarters (
<figureCitationid="1349E1FEFFC8FFA82720FA49FC016535"box="[844,948,1331,1357]"captionStart="FIGURE 10"captionStartId="18.[151,250,890,914]"captionTargetBox="[151,1435,181,866]"captionTargetId="figure-19@18.[151,1435,181,866]"captionTargetPageId="18"captionText="FIGURE 10. Glandular structure of male metatarsus III in Caribbiantes barbai sp. nov. A. Light microscope photo, holotype male (CZACC 3. 4380), general view of the whole metatarsus III (divided into four quarters to facilitate comparisons); B–E. Scanning electron micrographs, male (CZACC 3.3392); B. View of the glandular swelling, dashed line delineates the form of the ventral, buttonhole-like groove; C. Close-up of the groove delimited in B, note the depth of the groove; D. Closer zoom into the groove, white arrow shows a seta (long paint brush, basally striated and twisted), small arrowheads indicate glandular openings (pores) regularly arranged around the setae; E. Detail of the integument and pores (arrowheads) of the ventral groove."figureDoi="http://doi.org/10.5281/zenodo.13914749"httpUri="https://zenodo.org/record/13914749/files/figure.png"pageId="20"pageNumber="421">Fig. 10A</figureCitation>
). Ventral surface of the swollen region with wide, deep groove, buttonhole-shaped and bearing 31 long setae (
<figureCitationid="1349E1FEFFC8FFA827E7FA2DFB946509"box="[907,1057,1367,1393]"captionStart="FIGURE 10"captionStartId="18.[151,250,890,914]"captionTargetBox="[151,1435,181,866]"captionTargetId="figure-19@18.[151,1435,181,866]"captionTargetPageId="18"captionText="FIGURE 10. Glandular structure of male metatarsus III in Caribbiantes barbai sp. nov. A. Light microscope photo, holotype male (CZACC 3. 4380), general view of the whole metatarsus III (divided into four quarters to facilitate comparisons); B–E. Scanning electron micrographs, male (CZACC 3.3392); B. View of the glandular swelling, dashed line delineates the form of the ventral, buttonhole-like groove; C. Close-up of the groove delimited in B, note the depth of the groove; D. Closer zoom into the groove, white arrow shows a seta (long paint brush, basally striated and twisted), small arrowheads indicate glandular openings (pores) regularly arranged around the setae; E. Detail of the integument and pores (arrowheads) of the ventral groove."figureDoi="http://doi.org/10.5281/zenodo.13914749"httpUri="https://zenodo.org/record/13914749/files/figure.png"pageId="20"pageNumber="421">Figs. 10B, C</figureCitation>
); setal shafts basally striated and twisted, distally enlarged in the shape of long paint brushes (lanceolate-shape) (distally dilated as in
<figureCitationid="1349E1FEFFC8FFA824F3FAE4FE8565C0"box="[159,304,1438,1464]"captionStart="FIGURE 10"captionStartId="18.[151,250,890,914]"captionTargetBox="[151,1435,181,866]"captionTargetId="figure-19@18.[151,1435,181,866]"captionTargetPageId="18"captionText="FIGURE 10. Glandular structure of male metatarsus III in Caribbiantes barbai sp. nov. A. Light microscope photo, holotype male (CZACC 3. 4380), general view of the whole metatarsus III (divided into four quarters to facilitate comparisons); B–E. Scanning electron micrographs, male (CZACC 3.3392); B. View of the glandular swelling, dashed line delineates the form of the ventral, buttonhole-like groove; C. Close-up of the groove delimited in B, note the depth of the groove; D. Closer zoom into the groove, white arrow shows a seta (long paint brush, basally striated and twisted), small arrowheads indicate glandular openings (pores) regularly arranged around the setae; E. Detail of the integument and pores (arrowheads) of the ventral groove."figureDoi="http://doi.org/10.5281/zenodo.13914749"httpUri="https://zenodo.org/record/13914749/files/figure.png"pageId="20"pageNumber="421">Figs. 10D, E</figureCitation>
). Setae longitudinally arranged in two parallel alternating rows (
<figureCitationid="1349E1FEFFC8FFA82795FAE4FBEA65C0"box="[1017,1119,1438,1464]"captionStart="FIGURE 10"captionStartId="18.[151,250,890,914]"captionTargetBox="[151,1435,181,866]"captionTargetId="figure-19@18.[151,1435,181,866]"captionTargetPageId="18"captionText="FIGURE 10. Glandular structure of male metatarsus III in Caribbiantes barbai sp. nov. A. Light microscope photo, holotype male (CZACC 3. 4380), general view of the whole metatarsus III (divided into four quarters to facilitate comparisons); B–E. Scanning electron micrographs, male (CZACC 3.3392); B. View of the glandular swelling, dashed line delineates the form of the ventral, buttonhole-like groove; C. Close-up of the groove delimited in B, note the depth of the groove; D. Closer zoom into the groove, white arrow shows a seta (long paint brush, basally striated and twisted), small arrowheads indicate glandular openings (pores) regularly arranged around the setae; E. Detail of the integument and pores (arrowheads) of the ventral groove."figureDoi="http://doi.org/10.5281/zenodo.13914749"httpUri="https://zenodo.org/record/13914749/files/figure.png"pageId="20"pageNumber="421">Fig. 10C</figureCitation>
); numerous pores (glandular openings) regularly scattered around the base of the setae (
<figureCitationid="1349E1FEFFC8FFA82750FAB8FC1165A4"box="[828,932,1474,1500]"captionStart="FIGURE 10"captionStartId="18.[151,250,890,914]"captionTargetBox="[151,1435,181,866]"captionTargetId="figure-19@18.[151,1435,181,866]"captionTargetPageId="18"captionText="FIGURE 10. Glandular structure of male metatarsus III in Caribbiantes barbai sp. nov. A. Light microscope photo, holotype male (CZACC 3. 4380), general view of the whole metatarsus III (divided into four quarters to facilitate comparisons); B–E. Scanning electron micrographs, male (CZACC 3.3392); B. View of the glandular swelling, dashed line delineates the form of the ventral, buttonhole-like groove; C. Close-up of the groove delimited in B, note the depth of the groove; D. Closer zoom into the groove, white arrow shows a seta (long paint brush, basally striated and twisted), small arrowheads indicate glandular openings (pores) regularly arranged around the setae; E. Detail of the integument and pores (arrowheads) of the ventral groove."figureDoi="http://doi.org/10.5281/zenodo.13914749"httpUri="https://zenodo.org/record/13914749/files/figure.png"pageId="20"pageNumber="421">Fig. 10E</figureCitation>
); integument adjacent to setae wrinkled and with shallow grooves between pores and setae (
<figureCitationid="1349E1FEFFC8FFA826DFFA9CFCAF6678"box="[691,794,1510,1536]"captionStart="FIGURE 10"captionStartId="18.[151,250,890,914]"captionTargetBox="[151,1435,181,866]"captionTargetId="figure-19@18.[151,1435,181,866]"captionTargetPageId="18"captionText="FIGURE 10. Glandular structure of male metatarsus III in Caribbiantes barbai sp. nov. A. Light microscope photo, holotype male (CZACC 3. 4380), general view of the whole metatarsus III (divided into four quarters to facilitate comparisons); B–E. Scanning electron micrographs, male (CZACC 3.3392); B. View of the glandular swelling, dashed line delineates the form of the ventral, buttonhole-like groove; C. Close-up of the groove delimited in B, note the depth of the groove; D. Closer zoom into the groove, white arrow shows a seta (long paint brush, basally striated and twisted), small arrowheads indicate glandular openings (pores) regularly arranged around the setae; E. Detail of the integument and pores (arrowheads) of the ventral groove."figureDoi="http://doi.org/10.5281/zenodo.13914749"httpUri="https://zenodo.org/record/13914749/files/figure.png"pageId="20"pageNumber="421">Fig. 10E</figureCitation>
). Two smooth tarsal claws, perpendicular to the leg axis. Distitarsi III–IV with scopula. Tarsal formula: 7(3):13–15(5–6):8:8.
exhibits on the ventral side an apical process, wide at its base and sharp at the end (hook-shaped) (
<figureCitationid="1349E1FEFFC8FFA825F0F8E9FDF567D6"box="[412,576,1939,1966]"captionStart="FIGURE 11"captionStartId="21.[151,250,1761,1785]"captionTargetBox="[163,1432,191,1737]"captionTargetId="figure-18@21.[155,1432,181,1737]"captionTargetPageId="21"captionText="FIGURE 11. Caribbiantes barbai sp. nov., entire (C) anddistal part (A–B, D–J) of penis,A–F.Drawings with optical microscope, male holotype (CZACC 3.4380): A. Dorsal view; B. Dorsal view, expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view, expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm. G–J. Scanning electron micrographs (male CZACC 3.3392): G. Dorsal view, expanded; H. Apical view, expanded; I. Left lateral view, expanded; J. Ventral view, expanded.Abbreviations: AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae."figureDoi="http://doi.org/10.5281/zenodo.13914751"httpUri="https://zenodo.org/record/13914751/files/figure.png"pageId="20"pageNumber="421">Figs. 11D–E, I</figureCitation>
), along with five pairs of ventral setae (
<figureCitationid="1349E1FEFFC8FFA82798F8E9FBC167D6"box="[1012,1140,1939,1966]"captionStart="FIGURE 11"captionStartId="21.[151,250,1761,1785]"captionTargetBox="[163,1432,191,1737]"captionTargetId="figure-18@21.[155,1432,181,1737]"captionTargetPageId="21"captionText="FIGURE 11. Caribbiantes barbai sp. nov., entire (C) anddistal part (A–B, D–J) of penis,A–F.Drawings with optical microscope, male holotype (CZACC 3.4380): A. Dorsal view; B. Dorsal view, expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view, expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm. G–J. Scanning electron micrographs (male CZACC 3.3392): G. Dorsal view, expanded; H. Apical view, expanded; I. Left lateral view, expanded; J. Ventral view, expanded.Abbreviations: AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae."figureDoi="http://doi.org/10.5281/zenodo.13914751"httpUri="https://zenodo.org/record/13914751/files/figure.png"pageId="20"pageNumber="421">Figs. 11F, J</figureCitation>
). Apical-most pair of setae notably minute and bifurcated, subapical pair of setae longer; the three pairs of ventrolateral setae are much longer than the subapical pair of setae, broad-foliar shaped, and arranged in two oblique rows (
<figureCitationid="1349E1FEFFC8FFA82009F8A1FAAB678E"box="[1125,1310,2011,2038]"captionStart="FIGURE 11"captionStartId="21.[151,250,1761,1785]"captionTargetBox="[163,1432,191,1737]"captionTargetId="figure-18@21.[155,1432,181,1737]"captionTargetPageId="21"captionText="FIGURE 11. Caribbiantes barbai sp. nov., entire (C) anddistal part (A–B, D–J) of penis,A–F.Drawings with optical microscope, male holotype (CZACC 3.4380): A. Dorsal view; B. Dorsal view, expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view, expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm. G–J. Scanning electron micrographs (male CZACC 3.3392): G. Dorsal view, expanded; H. Apical view, expanded; I. Left lateral view, expanded; J. Ventral view, expanded.Abbreviations: AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae."figureDoi="http://doi.org/10.5281/zenodo.13914751"httpUri="https://zenodo.org/record/13914751/files/figure.png"pageId="20"pageNumber="421">Figs. 11E–F, I–J</figureCitation>
<taxonomicNameid="4C7286F8FFC9FFA9254CF99BFE596681"authorityName="Barroso & Pérez-González"authorityYear="2024"box="[288,492,1761,1785]"class="Arachnida"family="Biantidae"genus="Caribbiantes"kingdom="Animalia"order="Opiliones"pageId="21"pageNumber="422"phylum="Arthropoda"rank="species"species="barbai"status="sp. nov.">
entire (C) and distal part (A–B, D–J) of penis,A–F. Drawings with optical microscope, male holotype (CZACC 3.4380): A. Dorsal view; B. Dorsal view, expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view, expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm. G–J. Scanning electron micrographs (male CZACC 3.3392): G. Dorsal view, expanded; H. Apical view, expanded; I. Left lateral view, expanded; J. Ventral view, expanded. Abbreviations: AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae.
<taxonomicNameid="4C7286F8FFCAFFAA271FF87BFC6B6760"authorityName="Barroso & Pérez-González"authorityYear="2024"box="[883,990,1792,1816]"class="Arachnida"family="Biantidae"genus="Caribbiantes"kingdom="Animalia"order="Opiliones"pageId="22"pageNumber="423"phylum="Arthropoda"rank="species"species="obtusus"status="sp. nov.">
<taxonomicNameid="4C7286F8FFCAFFAA24FBF85FFF4F6744"authorityName="Barroso & Pérez-González"authorityYear="2024"box="[151,250,1828,1852]"class="Arachnida"family="Biantidae"genus="Caribbiantes"kingdom="Animalia"order="Opiliones"pageId="22"pageNumber="423"phylum="Arthropoda"rank="species"species="barbai"status="sp. nov.">
(squares, type locality in red); B. Submontane rain forest in Alejandro de Humboldt National Park, Monte Iberia, type locality of
<taxonomicNameid="4C7286F8FFCAFFAA25EDF833FE566718"authorityName="Barroso & Pérez-González"authorityYear="2024"box="[385,483,1864,1888]"class="Arachnida"family="Biantidae"genus="Caribbiantes"kingdom="Animalia"order="Opiliones"pageId="22"pageNumber="423"phylum="Arthropoda"rank="species"species="barbai"status="sp. nov.">
<taxonomicNameid="4C7286F8FFCAFFAA24AEF817FE9867FC"authorityName="Barroso & Pérez-González"authorityYear="2024"box="[194,301,1900,1924]"class="Arachnida"family="Biantidae"genus="Caribbiantes"kingdom="Animalia"order="Opiliones"pageId="22"pageNumber="423"phylum="Arthropoda"rank="species"species="obtusus"status="sp. nov.">
): Dorsum reddish-brown, with some yellowish areas; a dark brown line borders the anterior margin and the cheliceral sockets. Brown reticulate marks anterolaterally on the carapace, leaving the median and the posterior median (next to area I) sectors of the carapace yellowish. Lateral margins with irregular brown marks. Areas I–II brown, with yellowish spots. Areas III–IV with brown reticulated marks and yellowish spots. Spiniform apophyses on areas III–IV dark brown. Posterior margin and free tergites dark brown, with yellowish spots on the tubercles. Legs brown, with a striped pattern of irregular yellowish spots. Chelicerae yellowish with brown reticle; fingers yellowish-brown. Pedipalps pale brown with yellow stripes, distal segments yellowish.
(CZACC 3.4380). Similar to male, but slightly slenderer. Free tergite III with four small tubercles between the pair of spiniform apophyses. Chelicerae small, non hypertelic, without a gap between the cheliceral fingers as in α males; fixed and movable fingers with six distal teeth. Legs III–IV with femur, patella, and tibia slightly thinner than in males; metatarsus III not enlarged. Femora II–IV with a distal spiniform apophysis, as in the male. Coloration pattern as in males. Genital operculum wider than the male. Tarsal formula: 7(3):12(4):8:8. Measurements:
<tableCitationid="C6F0C8C0FFCBFFAB2525FD11FE2B62FD"box="[329,414,619,645]"captionStart="TABLE 3"captionStartId="19.[152,244,151,175]"captionTargetPageId="19"captionText="TABLE 3. Measurements (in mm) of the male holotype (H) and paratypes (P) of Caribbiantes barbai sp. nov.."httpUri="http://table.plazi.org/id/DF0DADF3FFCFFFAF24F4FFEDFA8B60D7"pageId="23"pageNumber="424"tableUuid="DF0DADF3FFCFFFAF24F4FFEDFA8B60D7">Table 3</tableCitation>
<tableCitationid="C6F0C8C0FFCBFFAB270CFDF5FC0062D1"box="[864,949,655,681]"captionStart="TABLE 3"captionStartId="19.[152,244,151,175]"captionTargetPageId="19"captionText="TABLE 3. Measurements (in mm) of the male holotype (H) and paratypes (P) of Caribbiantes barbai sp. nov.."httpUri="http://table.plazi.org/id/DF0DADF3FFCFFFAF24F4FFEDFA8B60D7"pageId="23"pageNumber="424"tableUuid="DF0DADF3FFCFFFAF24F4FFEDFA8B60D7">Table 3</tableCitation>
. Cheliceral dentition: fixed finger with 5–7 distal teeth, movable finger with 6–8 distal teeth. In the swollen region of metatarsus III each studied male had a different number of setae (26, 31, 32). The tarsal formula varies only in leg II: 7(3):12–15(4–6):8:8. Hypertelic chelicerae in only two studied males (α males—including the
This species has the eastern-most records for the genus. It is likely a Cuban short-ranged endemic, so far restricted to three localities in Toldo and Monte
Plateaus (Holguín and Guantánamo provinces, respectively) (
<figureCitationid="1349E1FEFFCBFFAB255BFC1DFE1563F9"box="[311,416,871,897]"captionStart="FIGURE 12"captionStartId="22.[151,250,1756,1780]"captionTargetBox="[164,1423,181,1732]"captionTargetId="figure-19@22.[164,1423,181,1732]"captionTargetPageId="22"captionText="FIGURE 12. Records and habitat of Caribbiantes spp. A. Map of Cuba (general and expanded sector as indicated), showing known localities of Caribbiantes cubanus (circles, type locality in red); C. obtusus sp. nov. (triangles, type locality in red) and C. barbai sp. nov. (squares, type locality in red); B. Submontane rain forest in Alejandro de Humboldt National Park, Monte Iberia, type locality of C. barbai sp. nov.; C. Mountains of Topes de Collantes, Sancti Spíritus province, record of C. cubanus and C. obtusus sp. nov."figureDoi="http://doi.org/10.5281/zenodo.13914753"httpUri="https://zenodo.org/record/13914753/files/figure.png"pageId="23"pageNumber="424">Fig. 12A</figureCitation>
<figureCitationid="1349E1FEFFCBFFAB249BFCD5FEDC63B1"box="[247,361,943,969]"captionStart="FIGURE 12"captionStartId="22.[151,250,1756,1780]"captionTargetBox="[164,1423,181,1732]"captionTargetId="figure-19@22.[164,1423,181,1732]"captionTargetPageId="22"captionText="FIGURE 12. Records and habitat of Caribbiantes spp. A. Map of Cuba (general and expanded sector as indicated), showing known localities of Caribbiantes cubanus (circles, type locality in red); C. obtusus sp. nov. (triangles, type locality in red) and C. barbai sp. nov. (squares, type locality in red); B. Submontane rain forest in Alejandro de Humboldt National Park, Monte Iberia, type locality of C. barbai sp. nov.; C. Mountains of Topes de Collantes, Sancti Spíritus province, record of C. cubanus and C. obtusus sp. nov."figureDoi="http://doi.org/10.5281/zenodo.13914753"httpUri="https://zenodo.org/record/13914753/files/figure.png"pageId="23"pageNumber="424">Figs. 12B</figureCitation>
