treatments-xml/data/ED/18/87/ED1887CBFFD1FFDFFCF8B191B254638A.xml

<document id="CEC83BFC9135982F1BC94FCD613F7167" ID-DOI="10.12651/JSR.2024.13.2.185" ID-ISSN="2713-8615" ID-Zenodo-Dep="13344859" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722550107885" checkinUser="felipe" docAuthor="Lee, Il-Hoi Kim and Taekjun" docDate="2024" docId="ED1887CBFFD1FFDFFCF8B191B254638A" docLanguage="en" docName="JSpeciesRes.13.2.185-254.pdf" docOrigin="Journal of Species Research 13 (2)" docStyle="DocumentStyle:4D398BFF45FF8CA9E08F546BB06AE372.2:JSpeciesRes.2012-.journal_article" docStyleId="4D398BFF45FF8CA9E08F546BB06AE372" docStyleName="JSpeciesRes.2012-.journal_article" docStyleVersion="2" docTitle="Asterocheres nodulosus Lee, 2024, n. sp." docType="treatment" docVersion="3" lastPageNumber="236" masterDocId="1121FFB3FFFEFFEBFFD1B25AB0306275" masterDocTitle="Copepods of the genus Asterocheres Boeck, 1859 (Siphonostomatoida: Asterocheridae) from Korean waters" masterLastPageNumber="254" masterPageNumber="185" pageNumber="232" updateTime="1724091014337" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="F05F2F4C5366D36F2C37A353D11041D9">Copepods of the genus Asterocheres Boeck, 1859 (Siphonostomatoida: Asterocheridae) from Korean waters</mods:title>
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<mods:namePart id="EDD8D3797FDA92B1ABF4DD7BDD5881C1">Lee, Il-Hoi Kim and Taekjun</mods:namePart>
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<mods:title id="2DED021365E3B945E7DFE3398F60B338">Journal of Species Research</mods:title>
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<mods:date id="FECAD1EE7742F4CB72923B5EE54DF8C2">2024</mods:date>
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<paragraph id="650E36DDFFD1FFC4FCF8B191B5386196" blockId="47.[809,1414,971,1059]" box="[809,1288,971,995]" pageId="47" pageNumber="232">
<heading id="3E4681B1FFD1FFC4FCF8B191B5386196" bold="true" box="[809,1288,971,995]" fontSize="10" level="2" pageId="47" pageNumber="232" reason="4">
<emphasis id="57C5EACFFFD1FFC4FCF8B191B5386196" bold="true" box="[809,1288,971,995]" pageId="47" pageNumber="232">
<taxonomicName id="A2B14D5EFFD1FFC4FCF8B191B41F6196" authorityName="Lee" authorityYear="2024" box="[809,1071,971,995]" class="Copepoda" family="Asterocheridae" genus="Asterocheres" kingdom="Animalia" order="Siphonostomatoida" pageId="47" pageNumber="232" phylum="Arthropoda" rank="species" species="nodulosus" status="sp. nov.">
<emphasis id="57C5EACFFFD1FFC4FCF8B191B41F6196" bold="true" box="[809,1071,971,995]" italics="true" pageId="47" pageNumber="232">Asterocheres nodulosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CF657B4FFD1FFC4FBE4B191B4416196" box="[1077,1137,971,995]" pageId="47" pageNumber="232" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="FD8A2A58FFD1FFC4FBACB191B5306196" box="[1149,1280,971,995]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="48.[174,207,1922,1944]" captionStartId-1="49.[144,177,1592,1614]" captionStartId-2="50.[174,207,1926,1948]" captionTargetBox-0="[276,1347,267,1863]" captionTargetBox-1="[265,1295,260,1551]" captionTargetBox-2="[268,1350,259,1877]" captionTargetId-0="figure-13@48.[192,1447,254,1892]" captionTargetId-1="figure-167@49.[175,1413,254,1561]" captionTargetId-2="figure-13@50.[171,1447,254,1892]" captionTargetPageId-0="48" captionTargetPageId-1="49" captionTargetPageId-2="50" captionText-0="Fig. 30. Asterocheres nodulosus n. sp., female. A, habitus, dorsal; B, urosome, dorsal; C, right caudal ramus, ventral; D, rostrum; E, antennule; F, antenna; G, oral siphon; H, mandible; I, maxillule; J, maxilla; K, maxilliped. Scale bars: A, 0.1 mm; B, E, G, H, 0.05 mm; C, D, F, I-K, 0.02 mm." captionText-1="Fig. 31. Asterocheres nodulosus n. sp., female. A, leg 1; B, leg 2; C, endopod of leg 3; D, leg 4; E, exopod of leg 5; F, left side of genital double-somite, dorsal. Scale bars: A-D, 0.05mm; E, F, 0.02mm." captionText-2="Fig. 32. Asterocheres nodulosus n. sp., male. A, habitus, dorsal; B, urosome, ventral; C, antennule; D, maxilliped; E, endopod of leg 2; F, endopod of leg 3; G, exopod of leg 5; H, leg 6. Scale bars:A, 0.1mm; B, 0.05 mm; C-H, 0.02 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13344921" figureDoi-1="http://doi.org/10.5281/zenodo.13344923" figureDoi-2="http://doi.org/10.5281/zenodo.13344925" httpUri-0="https://zenodo.org/record/13344921/files/figure.png" httpUri-1="https://zenodo.org/record/13344923/files/figure.png" httpUri-2="https://zenodo.org/record/13344925/files/figure.png" pageId="47" pageNumber="232">Figs. 30-32</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="650E36DDFFD1FFC4FCF8B1B1B5B66676" blockId="47.[809,1414,971,1059]" box="[809,1414,1003,1027]" pageId="47" pageNumber="232">https://zoobank.org/urn:lsid:zoobank.org:act:C3D1431</paragraph>
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<subSubSection id="2DAB6556FFD1FFC4FCEEB651B4EB6656" box="[831,1243,1035,1059]" pageId="47" pageNumber="232" type="description">
<paragraph id="650E36DDFFD1FFC4FCEEB651B4EB6656" blockId="47.[809,1414,971,1059]" box="[831,1243,1035,1059]" pageId="47" pageNumber="232">D-8821-4AE6-AF00-F40D8D5787CE</paragraph>
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<subSubSection id="2DAB6556FFD1FFC4FCF8B616B5336713" pageId="47" pageNumber="232" type="materials_examined">
<paragraph id="650E36DDFFD1FFC4FCF8B616B5336713" blockId="47.[809,1416,1098,1995]" pageId="47" pageNumber="232">
<emphasis id="57C5EACFFFD1FFC4FCF8B616B3EC6611" bold="true" box="[809,988,1100,1124]" pageId="47" pageNumber="232">Type material.</emphasis>
<materialsCitation id="D5D93C80FFD1FFC4FC36B616B41E6770" collectingDate="2018-08-29" collectionCode="SCUBA" collectorName="J. Lee." elevation="20" latitude="37.540276" location="Ulleung Island" longLatPrecision="19" longitude="130.84721" pageId="47" pageNumber="232" specimenCode="NIBRIV0000909865, NIBRIV0000909866" specimenCount="23" specimenCount-female="17" specimenCount-male="6" typeStatus="holotype">
<typeStatus id="BA0A887FFFD1FFC4FC36B616B4606611" box="[999,1104,1100,1124]" pageId="47" pageNumber="232" type="holotype">Holotype</typeStatus>
(
<specimenCount id="73B7FD54FFD1FFC4FBB0B610B44C6610" box="[1121,1148,1098,1125]" count="1" pageId="47" pageNumber="232" type="female">♀</specimenCount>
, 
<specimenCode id="35179EA6FFD1FFC4FB57B616B5496611" box="[1158,1401,1100,1124]" collectionCode="NIBRIV" pageId="47" pageNumber="232">NIBRIV0000909865</specimenCode>
), intact 
<typeStatus id="BA0A887FFFD1FFC4FCA3B637B3EE66F0" box="[882,990,1133,1157]" pageId="47" pageNumber="232" type="paratype">paratypes</typeStatus>
(
<specimenCount id="73B7FD54FFD1FFC4FC3FB636B40966F0" box="[1006,1081,1130,1157]" count="15" pageId="47" pageNumber="232" type="female">15♀♀</specimenCount>
, 
<specimenCount id="73B7FD54FFD1FFC4FB92B636B44F66F0" box="[1091,1151,1130,1157]" count="5" pageId="47" pageNumber="232" type="male">5♂♂</specimenCount>
, 
<specimenCode id="35179EA6FFD1FFC4FB59B636B54A66F1" box="[1160,1402,1132,1156]" collectionCode="NIBRIV" pageId="47" pageNumber="232">NIBRIV0000909866</specimenCode>
), and dissected 
<typeStatus id="BA0A887FFFD1FFC4FC16B6D7B40066D0" box="[967,1072,1165,1189]" pageId="47" pageNumber="232" type="paratype">paratypes</typeStatus>
(
<specimenCount id="73B7FD54FFD1FFC4FB90B6D7B45866D0" box="[1089,1128,1162,1189]" count="1" pageId="47" pageNumber="232" type="female">1♀</specimenCount>
, 
<specimenCount id="73B7FD54FFD1FFC4FBA1B6D7B4A966D0" box="[1136,1177,1162,1189]" count="1" pageId="47" pageNumber="232" type="male">1♂</specimenCount>
) from mixed species of sponges, 
<location id="606E6006FFD1FFC4FC7FB6F7B47E66B0" LSID="urn:lsid:plazi:treatment:ED1887CBFFD1FFDFFCF8B191B254638A:606E6006FFD1FFC4FC7FB6F7B47E66B0" box="[942,1102,1197,1221]" latitude="37.540276" longLatPrecision="19" longitude="130.84721" name="Ulleung Island" pageId="47" pageNumber="232">Ulleung Island</location>
(approximately 
<geoCoordinate id="0085501AFFD1FFC4FAD0B6F7B5B266B1" box="[1281,1410,1195,1221]" degrees="37" direction="north" minutes="32" orientation="latitude" pageId="47" pageNumber="232" precision="15" seconds="25" value="37.540276">
37°32 
<emphasis id="57C5EACFFFD1FFC4FA90B6F1B57766B0" bold="true" box="[1345,1351,1195,1221]" italics="true" pageId="47" pageNumber="232">ʹ</emphasis>
25 
<emphasis id="57C5EACFFFD1FFC4FAB3B6F1B55C66B0" bold="true" box="[1378,1388,1195,1221]" italics="true" pageId="47" pageNumber="232">ʺ</emphasis>
N
</geoCoordinate>
, 
<geoCoordinate id="0085501AFFD1FFC4FCF8B697B3866690" box="[809,950,1227,1253]" degrees="130" direction="east" minutes="50" orientation="longitude" pageId="47" pageNumber="232" precision="15" seconds="50" value="130.84721">
130°50 
<emphasis id="57C5EACFFFD1FFC4FCA9B691B34E6690" bold="true" box="[888,894,1227,1253]" italics="true" pageId="47" pageNumber="232">ʹ</emphasis>
50 
<emphasis id="57C5EACFFFD1FFC4FC48B691B3936690" bold="true" box="[921,931,1227,1253]" italics="true" pageId="47" pageNumber="232">ʺ</emphasis>
E
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), depth about 
<quantity id="A2499B38FFD1FFC4FB89B697B4A06691" box="[1112,1168,1229,1253]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="47" pageNumber="232" unit="m" value="20.0">
<elevation id="EE9CD1EEFFD1FFC4FB89B697B4A06691" box="[1112,1168,1229,1253]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="47" pageNumber="232" unit="m" value="20.0">20 m</elevation>
</quantity>
, 
<collectionCode id="03A0AE18FFD1FFC4FB4FB697B4C76690" box="[1182,1271,1229,1253]" pageId="47" pageNumber="232">SCUBA</collectionCode>
diving, coll. 
<collectorName id="C844530BFFD1FFC4FCF8B6B4B3436770" box="[809,883,1262,1285]" pageId="47" pageNumber="232">J. Lee.</collectorName>
<date id="110F101DFFD1FFC4FCADB6B7B41E6770" box="[892,1070,1261,1285]" pageId="47" pageNumber="232" value="2018-08-29">
<collectingDate id="014BE9F5FFD1FFC4FCADB6B7B41E6770" box="[892,1070,1261,1285]" pageId="47" pageNumber="232" value="2018-08-29">29 August 2018</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="D5D93C80FFD1FFC4FBEAB6B7B4CF6713" collectionCode="NIBR" collectorName="Dissected &amp; I. - H. Kim" country="South Korea" location="National Institute of Biological Resources" pageId="47" pageNumber="232" specimenCount="3" stateProvince="Incheon" typeStatus="holotype">
<typeStatus id="BA0A887FFFD1FFC4FBEAB6B7B4AF6770" box="[1083,1183,1261,1285]" pageId="47" pageNumber="232" type="holotype">Holotype</typeStatus>
and intact 
<typeStatus id="BA0A887FFFD1FFC4FACFB6B4B5B56773" box="[1310,1413,1262,1286]" pageId="47" pageNumber="232" type="paratype">paratypes</typeStatus>
have been deposited in the 
<location id="606E6006FFD1FFC4FB83B757B3F56730" LSID="urn:lsid:plazi:treatment:ED1887CBFFD1FFDFFCF8B191B254638A:606E6006FFD1FFC4FB83B757B3F56730" country="South Korea" name="National Institute of Biological Resources" pageId="47" pageNumber="232" stateProvince="Incheon">National Institute of Biological Resources</location>
(
<collectionCode id="03A0AE18FFD1FFC4FC03B774B4276733" box="[978,1047,1326,1350]" country="South Korea" httpUri="http://grbio.org/cool/r4iq-k5jx" name="National Institute of Biological Resources" pageId="47" pageNumber="232">NIBR</collectionCode>
), 
<collectingRegion id="A775F83FFFD1FFC4FBFBB774B4B36733" box="[1066,1155,1326,1350]" country="South Korea" name="Incheon" pageId="47" pageNumber="232">Incheon</collectingRegion>
, 
<collectingCountry id="1DA6764DFFD1FFC4FB5EB774B4E46730" box="[1167,1236,1326,1349]" name="South Korea" pageId="47" pageNumber="232">Korea</collectingCountry>
. 
<collectorName id="C844530BFFD1FFC4FB0EB774B5776733" box="[1247,1351,1326,1350]" pageId="47" pageNumber="232">Dissected</collectorName>
<typeStatus id="BA0A887FFFD1FFC4FA9EB774B3516712" pageId="47" pageNumber="232" type="paratype">paratypes</typeStatus>
are kept in the collection of 
<collectorName id="C844530BFFD1FFC4FB42B715B4CF6713" box="[1171,1279,1358,1382]" pageId="47" pageNumber="232">I.-H. Kim</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="2DAB6556FFD1FFD8FCF8B735B4AA66F1" lastPageId="51" lastPageNumber="236" pageId="47" pageNumber="232" type="etymology">
<paragraph id="650E36DDFFD1FFC4FCF8B735B4A267B2" blockId="47.[809,1416,1098,1995]" pageId="47" pageNumber="232">
<emphasis id="57C5EACFFFD1FFC4FCF8B735B39967F2" bold="true" box="[809,937,1391,1415]" pageId="47" pageNumber="232">Etymology.</emphasis>
The specific name refers to the nodule-like proximal expansion of the medio-distal seta of the third endopodal segment of male leg 2.
</paragraph>
<paragraph id="650E36DDFFD1FFDAFCF8B795B16C651D" blockId="47.[809,1416,1098,1995]" lastBlockId="49.[144,749,1711,1994]" lastPageId="49" lastPageNumber="234" pageId="47" pageNumber="232">
<emphasis id="57C5EACFFFD1FFC4FCF8B795B3B36792" bold="true" box="[809,899,1487,1511]" pageId="47" pageNumber="232">Female.</emphasis>
Body form (
<figureCitation id="FD8A2A58FFD1FFC4FBDAB795B45D6792" box="[1035,1133,1487,1511]" captionStart="Fig" captionStartId="48.[174,207,1922,1944]" captionTargetBox="[276,1347,267,1863]" captionTargetId="figure-13@48.[192,1447,254,1892]" captionTargetPageId="48" captionText="Fig. 30. Asterocheres nodulosus n. sp., female. A, habitus, dorsal; B, urosome, dorsal; C, right caudal ramus, ventral; D, rostrum; E, antennule; F, antenna; G, oral siphon; H, mandible; I, maxillule; J, maxilla; K, maxilliped. Scale bars: A, 0.1 mm; B, E, G, H, 0.05 mm; C, D, F, I-K, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.13344921" httpUri="https://zenodo.org/record/13344921/files/figure.png" pageId="47" pageNumber="232">Fig. 30A</figureCitation>
) as usual for genus. Body length 793 μm in figured and described specimen. Other 4 measured specimens 695, 730, 755, and 840 μm, respectively. Prosome 549 μm long. Cephalothorax wider than long (364 × 436 μm). All prosomal somites with rounded posterolateral corners. Urosome (
<figureCitation id="FD8A2A58FFD1FFC4FAC4B42AB54D64FC" box="[1301,1405,1648,1673]" captionStart="Fig" captionStartId="48.[174,207,1922,1944]" captionTargetBox="[276,1347,267,1863]" captionTargetId="figure-13@48.[192,1447,254,1892]" captionTargetPageId="48" captionText="Fig. 30. Asterocheres nodulosus n. sp., female. A, habitus, dorsal; B, urosome, dorsal; C, right caudal ramus, ventral; D, rostrum; E, antennule; F, antenna; G, oral siphon; H, mandible; I, maxillule; J, maxilla; K, maxilliped. Scale bars: A, 0.1 mm; B, E, G, H, 0.05 mm; C, D, F, I-K, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.13344921" httpUri="https://zenodo.org/record/13344921/files/figure.png" pageId="47" pageNumber="232">Fig. 30B</figureCitation>
) 4-segmented. Fifth pedigerous somite wider than genital double-somite, with spinules on lateral surfaces. Genital double-somite slightly wider than long (108 × 110 μm); genital apertures positioned dorsolaterally at 47% region of double-somite length; postgenital lateral margin bearing 2 groups of spinules consisting of 13 or 14 thin spinules of anterior group and 4 thicker spinules of posterior group (
<figureCitation id="FD8A2A58FFD1FFC4FC0DB529B40D65FE" box="[988,1085,1907,1931]" captionStart="Fig" captionStartId="49.[144,177,1592,1614]" captionTargetBox="[265,1295,260,1551]" captionTargetId="figure-167@49.[175,1413,254,1561]" captionTargetPageId="49" captionText="Fig. 31. Asterocheres nodulosus n. sp., female. A, leg 1; B, leg 2; C, endopod of leg 3; D, leg 4; E, exopod of leg 5; F, left side of genital double-somite, dorsal. Scale bars: A-D, 0.05mm; E, F, 0.02mm." figureDoi="http://doi.org/10.5281/zenodo.13344923" httpUri="https://zenodo.org/record/13344923/files/figure.png" pageId="47" pageNumber="232">Fig. 31F</figureCitation>
). Two free abdominal somites 36× 66 and 38 × 58 μm, respectively. Genital double-somite and first free abdominal somite bearing row of minute spinules along posterodorsal margin. Anal somite with row of spinules along posteroventral margin (
<figureCitation id="FD8A2A58FFCFFFDAFD10B495B0F0657D" captionStart="Fig" captionStartId="48.[174,207,1922,1944]" captionTargetBox="[276,1347,267,1863]" captionTargetId="figure-13@48.[192,1447,254,1892]" captionTargetPageId="48" captionText="Fig. 30. Asterocheres nodulosus n. sp., female. A, habitus, dorsal; B, urosome, dorsal; C, right caudal ramus, ventral; D, rostrum; E, antennule; F, antenna; G, oral siphon; H, mandible; I, maxillule; J, maxilla; K, maxilliped. Scale bars: A, 0.1 mm; B, E, G, H, 0.05 mm; C, D, F, I-K, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.13344921" httpUri="https://zenodo.org/record/13344921/files/figure.png" pageId="49" pageNumber="234">Fig. 30C</figureCitation>
). Caudal ramus 1.08 times longer than wide (28× 26 μm), armed with 6 setae and ornamented with few scales on outer and inner margins and row of spinules on posteroventral margin.
</paragraph>
<caption id="31CE6655FFCEFFDBFF7FB5D8B10365B8" ID-DOI="http://doi.org/10.5281/zenodo.13344921" ID-Zenodo-Dep="13344921" httpUri="https://zenodo.org/record/13344921/files/figure.png" pageId="48" pageNumber="233" startId="48.[174,207,1922,1944]" targetBox="[276,1347,267,1863]" targetPageId="48" targetType="figure">
<paragraph id="650E36DDFFCEFFDBFF7FB5D8B10365B8" blockId="48.[174,1444,1922,1997]" pageId="48" pageNumber="233">
<emphasis id="57C5EACFFFCEFFDBFF7FB5D8B0C565E2" bold="true" box="[174,245,1922,1944]" pageId="48" pageNumber="233">Fig. 30.</emphasis>
<taxonomicName id="A2B14D5EFFCEFFDBFF2DB5D8B1E365E2" authorityName="Lee" authorityYear="2024" box="[252,467,1922,1943]" class="Copepoda" family="Asterocheridae" genus="Asterocheres" kingdom="Animalia" order="Siphonostomatoida" pageId="48" pageNumber="233" phylum="Arthropoda" rank="species" species="nodulosus" status="sp. nov.">
<emphasis id="57C5EACFFFCEFFDBFF2DB5D8B1E365E2" box="[252,467,1922,1943]" italics="true" pageId="48" pageNumber="233">Asterocheres nodulosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CF657B4FFCEFFDBFE08B5D9B23C65ED" box="[473,524,1923,1944]" pageId="48" pageNumber="233" rank="species">n. sp.</taxonomicNameLabel>
, female. A, habitus, dorsal; B, urosome, dorsal; C, right caudal ramus, ventral; D, rostrum; E, antennule; F, antenna; G, oral siphon; H, mandible; I, maxillule; J, maxilla; K, maxilliped. Scale bars: A, 0.1 mm; B, E, G, H, 0.05 mm; C, D, F, I- K, 0.02 mm.
</paragraph>
</caption>
<caption id="31CE6655FFCFFFDAFF41B462B2EE641D" ID-DOI="http://doi.org/10.5281/zenodo.13344923" ID-Zenodo-Dep="13344923" httpUri="https://zenodo.org/record/13344923/files/figure.png" pageId="49" pageNumber="234" startId="49.[144,177,1592,1614]" targetBox="[265,1295,260,1551]" targetPageId="49" targetType="figure">
<paragraph id="650E36DDFFCFFFDAFF41B462B2EE641D" blockId="49.[144,1414,1592,1641]" pageId="49" pageNumber="234">
<emphasis id="57C5EACFFFCFFFDAFF41B462B0E86438" bold="true" box="[144,216,1592,1614]" pageId="49" pageNumber="234">Fig. 31.</emphasis>
<taxonomicName id="A2B14D5EFFCFFFDAFF0EB462B1866438" authorityName="Lee" authorityYear="2024" box="[223,438,1592,1613]" class="Copepoda" family="Asterocheridae" genus="Asterocheres" kingdom="Animalia" order="Siphonostomatoida" pageId="49" pageNumber="234" phylum="Arthropoda" rank="species" species="nodulosus" status="sp. nov.">
<emphasis id="57C5EACFFFCFFFDAFF0EB462B1866438" box="[223,438,1592,1613]" italics="true" pageId="49" pageNumber="234">Asterocheres nodulosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CF657B4FFCFFFDAFE6CB463B1C1643B" box="[445,497,1593,1614]" pageId="49" pageNumber="234" rank="species">n. sp.</taxonomicNameLabel>
, female. A, leg 1; B, leg 2; C, endopod of leg 3; D, leg 4; E, exopod of leg 5; F, left side of genital double-somite, dorsal. Scale bars: A- D, 0.05 mm; E, F, 0.02 mm.
</paragraph>
</caption>
<paragraph id="650E36DDFFCFFFD8FF77B52BB0C163AA" blockId="49.[144,749,1711,1994]" lastBlockId="51.[144,750,261,1996]" lastPageId="51" lastPageNumber="236" pageId="49" pageNumber="234">
Rostrum (
<figureCitation id="FD8A2A58FFCFFFDAFEC0B52BB14565FC" box="[273,373,1905,1929]" captionStart="Fig" captionStartId="48.[174,207,1922,1944]" captionTargetBox="[276,1347,267,1863]" captionTargetId="figure-13@48.[192,1447,254,1892]" captionTargetPageId="48" captionText="Fig. 30. Asterocheres nodulosus n. sp., female. A, habitus, dorsal; B, urosome, dorsal; C, right caudal ramus, ventral; D, rostrum; E, antennule; F, antenna; G, oral siphon; H, mandible; I, maxillule; J, maxilla; K, maxilliped. Scale bars: A, 0.1 mm; B, E, G, H, 0.05 mm; C, D, F, I-K, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.13344921" httpUri="https://zenodo.org/record/13344921/files/figure.png" pageId="49" pageNumber="234">Fig. 30D</figureCitation>
) as tapering ridge between antennules, without defined distal apex. Antennule (
<figureCitation id="FD8A2A58FFCFFFDAFD54B5CBB2D565DC" box="[645,741,1937,1961]" captionStart="Fig" captionStartId="48.[174,207,1922,1944]" captionTargetBox="[276,1347,267,1863]" captionTargetId="figure-13@48.[192,1447,254,1892]" captionTargetPageId="48" captionText="Fig. 30. Asterocheres nodulosus n. sp., female. A, habitus, dorsal; B, urosome, dorsal; C, right caudal ramus, ventral; D, rostrum; E, antennule; F, antenna; G, oral siphon; H, mandible; I, maxillule; J, maxilla; K, maxilliped. Scale bars: A, 0.1 mm; B, E, G, H, 0.05 mm; C, D, F, I-K, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.13344921" httpUri="https://zenodo.org/record/13344921/files/figure.png" pageId="49" pageNumber="234">Fig. 30E</figureCitation>
) 320 μm long, 21-segmented; 9th, 18th, 20th, and terminal segments with 7 setae, 2 setae + aesthetasc, 4 setae, and 7 setae, respectively; other segments each with 2 setae. First segment with several spinules on anterior margin; some of setae on anterior segments pinnate; one of setae on 9th and 19th segments with crack-like transverse lines; some of setae on proximal segments truncated and tipped with minute setule. Antenna (
<figureCitation id="FD8A2A58FFCFFFDAFA8BB52BB36465DC" captionStart="Fig" captionStartId="48.[174,207,1922,1944]" captionTargetBox="[276,1347,267,1863]" captionTargetId="figure-13@48.[192,1447,254,1892]" captionTargetPageId="48" captionText="Fig. 30. Asterocheres nodulosus n. sp., female. A, habitus, dorsal; B, urosome, dorsal; C, right caudal ramus, ventral; D, rostrum; E, antennule; F, antenna; G, oral siphon; H, mandible; I, maxillule; J, maxilla; K, maxilliped. Scale bars: A, 0.1 mm; B, E, G, H, 0.05 mm; C, D, F, I-K, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.13344921" httpUri="https://zenodo.org/record/13344921/files/figure.png" pageId="49" pageNumber="234">Fig. 30F</figureCitation>
) consisting of coxa, basis, small exopod, and 3-segmented endopod; coxa with several spinules on outer margin; basis with longitudinal row of needle-like minute spinules near outer margin and few spinules on inner margin; first endopodal segment 57 μm long, with setules on outer margin; small second endopodal segment bearing 1 seta; third endopodal segment bearing 2 setae, 1 spinule-like, minute seta; terminal claw spiniform, 54 μm long.
</paragraph>
<caption id="31CE6655FFCCFFD9FF7FB5DCB40065C3" ID-DOI="http://doi.org/10.5281/zenodo.13344925" ID-Zenodo-Dep="13344925" httpUri="https://zenodo.org/record/13344925/files/figure.png" pageId="50" pageNumber="235" startId="50.[174,207,1926,1948]" targetBox="[268,1350,259,1877]" targetPageId="50" targetType="figure">
<paragraph id="650E36DDFFCCFFD9FF7FB5DCB40065C3" blockId="50.[174,1444,1926,1975]" pageId="50" pageNumber="235">
<emphasis id="57C5EACFFFCCFFD9FF7FB5DCB0C665EE" bold="true" box="[174,246,1926,1948]" pageId="50" pageNumber="235">Fig. 32.</emphasis>
<taxonomicName id="A2B14D5EFFCCFFD9FF2CB5DCB1E565EE" authorityName="Lee" authorityYear="2024" box="[253,469,1926,1947]" class="Copepoda" family="Asterocheridae" genus="Asterocheres" kingdom="Animalia" order="Siphonostomatoida" pageId="50" pageNumber="235" phylum="Arthropoda" rank="species" species="nodulosus" status="sp. nov.">
<emphasis id="57C5EACFFFCCFFD9FF2CB5DCB1E565EE" box="[253,469,1926,1947]" italics="true" pageId="50" pageNumber="235">Asterocheres nodulosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CF657B4FFCCFFD9FE0DB5DDB23F65E9" box="[476,527,1927,1948]" pageId="50" pageNumber="235" rank="species">n. sp.</taxonomicNameLabel>
, male. A, habitus, dorsal; B, urosome, ventral; C, antennule; D, maxilliped; E, endopod of leg 2; F, endopod of leg 3; G, exopod of leg 5; H, leg 6. Scale bars: A, 0.1 mm; B, 0.05 mm; C- H, 0.02 mm.
</paragraph>
</caption>
<paragraph id="650E36DDFFCDFFD8FF77B3BDB2186690" blockId="51.[144,750,261,1996]" pageId="51" pageNumber="236">
Oral siphon (
<figureCitation id="FD8A2A58FFCDFFD8FEE3B3BDB1A4638A" box="[306,404,487,511]" captionStart="Fig" captionStartId="48.[174,207,1922,1944]" captionTargetBox="[276,1347,267,1863]" captionTargetId="figure-13@48.[192,1447,254,1892]" captionTargetPageId="48" captionText="Fig. 30. Asterocheres nodulosus n. sp., female. A, habitus, dorsal; B, urosome, dorsal; C, right caudal ramus, ventral; D, rostrum; E, antennule; F, antenna; G, oral siphon; H, mandible; I, maxillule; J, maxilla; K, maxilliped. Scale bars: A, 0.1 mm; B, E, G, H, 0.05 mm; C, D, F, I-K, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.13344921" httpUri="https://zenodo.org/record/13344921/files/figure.png" pageId="51" pageNumber="236">Fig. 30G</figureCitation>
) 197 × 65 μm, extending slightly over insertions of maxillipeds. Mandible (
<figureCitation id="FD8A2A58FFCDFFD8FD51B05DB2D4606A" box="[640,740,519,543]" captionStart="Fig" captionStartId="48.[174,207,1922,1944]" captionTargetBox="[276,1347,267,1863]" captionTargetId="figure-13@48.[192,1447,254,1892]" captionTargetPageId="48" captionText="Fig. 30. Asterocheres nodulosus n. sp., female. A, habitus, dorsal; B, urosome, dorsal; C, right caudal ramus, ventral; D, rostrum; E, antennule; F, antenna; G, oral siphon; H, mandible; I, maxillule; J, maxilla; K, maxilliped. Scale bars: A, 0.1 mm; B, E, G, H, 0.05 mm; C, D, F, I-K, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.13344921" httpUri="https://zenodo.org/record/13344921/files/figure.png" pageId="51" pageNumber="236">Fig. 30H</figureCitation>
) consisting of coxal stylet and palp; stylet 180 μm long, bearing row of 12 teeth at distal region; palp 1-segment- ed, 56 μm long, tipped with 1 long (159 μm long) and 1 shorter (68 μm long) setae, both setae weakly pinnate; palp plus longer distal seta 1.19 times longer than stylet. Maxillule (
<figureCitation id="FD8A2A58FFCDFFD8FEE9B093B1A56094" box="[312,405,713,737]" captionStart="Fig" captionStartId="48.[174,207,1922,1944]" captionTargetBox="[276,1347,267,1863]" captionTargetId="figure-13@48.[192,1447,254,1892]" captionTargetPageId="48" captionText="Fig. 30. Asterocheres nodulosus n. sp., female. A, habitus, dorsal; B, urosome, dorsal; C, right caudal ramus, ventral; D, rostrum; E, antennule; F, antenna; G, oral siphon; H, mandible; I, maxillule; J, maxilla; K, maxilliped. Scale bars: A, 0.1 mm; B, E, G, H, 0.05 mm; C, D, F, I-K, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.13344921" httpUri="https://zenodo.org/record/13344921/files/figure.png" pageId="51" pageNumber="236">Fig. 30I</figureCitation>
) bilobed; inner lobe about 59 μm long, tipped with 4 large and 1 rudimentary setae (lengths of 4 large setae 83, 85, 75, and 50 μm, respectively, from inner to outer); outer lobe 21 μm long tipped with 4 setae, lengths of these 4 setae 65, 65, 45, and 25 μm, respectively. Maxilla (
<figureCitation id="FD8A2A58FFCDFFD8FE35B130B27461F7" box="[484,580,874,898]" captionStart="Fig" captionStartId="48.[174,207,1922,1944]" captionTargetBox="[276,1347,267,1863]" captionTargetId="figure-13@48.[192,1447,254,1892]" captionTargetPageId="48" captionText="Fig. 30. Asterocheres nodulosus n. sp., female. A, habitus, dorsal; B, urosome, dorsal; C, right caudal ramus, ventral; D, rostrum; E, antennule; F, antenna; G, oral siphon; H, mandible; I, maxillule; J, maxilla; K, maxilliped. Scale bars: A, 0.1 mm; B, E, G, H, 0.05 mm; C, D, F, I-K, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.13344921" httpUri="https://zenodo.org/record/13344921/files/figure.png" pageId="51" pageNumber="236">Fig. 30J</figureCitation>
) 2-segmented; proximal segment (syncoxa) bearing excretory tube and row of fine spinules at proximal region; distal segment (basis) as long as proximal segment bearing 1 group of setules at distal third and row of minute spinules along distal third. Maxilliped (
<figureCitation id="FD8A2A58FFCDFFD8FE70B651B2376651" box="[417,519,1035,1060]" captionStart="Fig" captionStartId="48.[174,207,1922,1944]" captionTargetBox="[276,1347,267,1863]" captionTargetId="figure-13@48.[192,1447,254,1892]" captionTargetPageId="48" captionText="Fig. 30. Asterocheres nodulosus n. sp., female. A, habitus, dorsal; B, urosome, dorsal; C, right caudal ramus, ventral; D, rostrum; E, antennule; F, antenna; G, oral siphon; H, mandible; I, maxillule; J, maxilla; K, maxilliped. Scale bars: A, 0.1 mm; B, E, G, H, 0.05 mm; C, D, F, I-K, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.13344921" httpUri="https://zenodo.org/record/13344921/files/figure.png" pageId="51" pageNumber="236">Fig. 30K</figureCitation>
) consisting of coxa, basis, and 4-segmented endopod; coxa with 1 small seta subdistally on inner margin and several minute spinules at outer distal corner; basis unarmed but with minute spinules on outer margin; endopodal segments with 2, 1, 1, and 1 setae, respectively. Terminal segment 35 μm long and terminal claw of 61 μm long.
</paragraph>
<paragraph id="650E36DDFFCDFFD8FF77B6B4B1246452" blockId="51.[144,750,261,1996]" pageId="51" pageNumber="236">
Legs 1 (
<figureCitation id="FD8A2A58FFCDFFD8FED1B6B7B1576770" box="[256,359,1261,1285]" captionStart="Fig" captionStartId="49.[144,177,1592,1614]" captionTargetBox="[265,1295,260,1551]" captionTargetId="figure-167@49.[175,1413,254,1561]" captionTargetPageId="49" captionText="Fig. 31. Asterocheres nodulosus n. sp., female. A, leg 1; B, leg 2; C, endopod of leg 3; D, leg 4; E, exopod of leg 5; F, left side of genital double-somite, dorsal. Scale bars: A-D, 0.05mm; E, F, 0.02mm." figureDoi="http://doi.org/10.5281/zenodo.13344923" httpUri="https://zenodo.org/record/13344923/files/figure.png" pageId="51" pageNumber="236">Fig. 31A</figureCitation>
), 2 (
<figureCitation id="FD8A2A58FFCDFFD8FE48B6B7B1CE6770" box="[409,510,1261,1285]" captionStart="Fig" captionStartId="49.[144,177,1592,1614]" captionTargetBox="[265,1295,260,1551]" captionTargetId="figure-167@49.[175,1413,254,1561]" captionTargetPageId="49" captionText="Fig. 31. Asterocheres nodulosus n. sp., female. A, leg 1; B, leg 2; C, endopod of leg 3; D, leg 4; E, exopod of leg 5; F, left side of genital double-somite, dorsal. Scale bars: A-D, 0.05mm; E, F, 0.02mm." figureDoi="http://doi.org/10.5281/zenodo.13344923" httpUri="https://zenodo.org/record/13344923/files/figure.png" pageId="51" pageNumber="236">Fig. 31B</figureCitation>
), 3, and 4 (
<figureCitation id="FD8A2A58FFCDFFD8FDAEB6B7B2D46770" box="[639,740,1261,1285]" captionStart="Fig" captionStartId="49.[144,177,1592,1614]" captionTargetBox="[265,1295,260,1551]" captionTargetId="figure-167@49.[175,1413,254,1561]" captionTargetPageId="49" captionText="Fig. 31. Asterocheres nodulosus n. sp., female. A, leg 1; B, leg 2; C, endopod of leg 3; D, leg 4; E, exopod of leg 5; F, left side of genital double-somite, dorsal. Scale bars: A-D, 0.05mm; E, F, 0.02mm." figureDoi="http://doi.org/10.5281/zenodo.13344923" httpUri="https://zenodo.org/record/13344923/files/figure.png" pageId="51" pageNumber="236">Fig. 31D</figureCitation>
) with 3-segmented rami. Leg 3 similar to leg 2 except bearing 1 spine and 5 setae on third endopodal segment (
<figureCitation id="FD8A2A58FFCDFFD8FF03B714B1056713" box="[210,309,1358,1382]" captionStart="Fig" captionStartId="49.[144,177,1592,1614]" captionTargetBox="[265,1295,260,1551]" captionTargetId="figure-167@49.[175,1413,254,1561]" captionTargetPageId="49" captionText="Fig. 31. Asterocheres nodulosus n. sp., female. A, leg 1; B, leg 2; C, endopod of leg 3; D, leg 4; E, exopod of leg 5; F, left side of genital double-somite, dorsal. Scale bars: A-D, 0.05mm; E, F, 0.02mm." figureDoi="http://doi.org/10.5281/zenodo.13344923" httpUri="https://zenodo.org/record/13344923/files/figure.png" pageId="51" pageNumber="236">Fig. 31C</figureCitation>
). Intercoxal plate of legs 1-3 with small spinules, but smooth in leg 4. Inner coxal seta of leg 1 small but pinnate, that of leg 4 naked, rudimentary. Out- er seta on basis of leg 1 elongated, that of leg 4 short. Inner distal margin of basis of leg 1 bearing spinules. Armature formula for legs 1-4 same as that of 
<taxonomicName id="A2B14D5EFFCDFFD8FD4EB7AAB0E46452" authorityName="Lee" authorityYear="2024" class="Copepoda" family="Asterocheridae" genus="Asterocheres" kingdom="Animalia" order="Siphonostomatoida" pageId="51" pageNumber="236" phylum="Arthropoda" rank="species" species="processus" status="n. sp.">
<emphasis id="57C5EACFFFCDFFD8FD4EB7AAB0E46452" italics="true" pageId="51" pageNumber="236">A. processus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CF657B4FFCDFFD8FF0BB44AB1246452" box="[218,276,1552,1575]" pageId="51" pageNumber="236" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
<paragraph id="650E36DDFFCDFFD8FF77B46AB14A653F" blockId="51.[144,750,261,1996]" pageId="51" pageNumber="236">
Leg 5 (
<figureCitation id="FD8A2A58FFCDFFD8FF23B46AB162643D" box="[242,338,1584,1608]" captionStart="Fig" captionStartId="48.[174,207,1922,1944]" captionTargetBox="[276,1347,267,1863]" captionTargetId="figure-13@48.[192,1447,254,1892]" captionTargetPageId="48" captionText="Fig. 30. Asterocheres nodulosus n. sp., female. A, habitus, dorsal; B, urosome, dorsal; C, right caudal ramus, ventral; D, rostrum; E, antennule; F, antenna; G, oral siphon; H, mandible; I, maxillule; J, maxilla; K, maxilliped. Scale bars: A, 0.1 mm; B, E, G, H, 0.05 mm; C, D, F, I-K, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.13344921" httpUri="https://zenodo.org/record/13344921/files/figure.png" pageId="51" pageNumber="236">Fig. 30B</figureCitation>
) represented by 1 dorsolateral seta on fifth pedigerous somite and free exopod; exopodal segment (
<figureCitation id="FD8A2A58FFCDFFD8FF04B42AB10964FC" box="[213,313,1648,1673]" captionStart="Fig" captionStartId="49.[144,177,1592,1614]" captionTargetBox="[265,1295,260,1551]" captionTargetId="figure-167@49.[175,1413,254,1561]" captionTargetPageId="49" captionText="Fig. 31. Asterocheres nodulosus n. sp., female. A, leg 1; B, leg 2; C, endopod of leg 3; D, leg 4; E, exopod of leg 5; F, left side of genital double-somite, dorsal. Scale bars: A-D, 0.05mm; E, F, 0.02mm." figureDoi="http://doi.org/10.5281/zenodo.13344923" httpUri="https://zenodo.org/record/13344923/files/figure.png" pageId="51" pageNumber="236">Fig. 31E</figureCitation>
) 2.14 times longer than wide (47 × 22 μm), bearing spinules on convex inner and outer margins and armed with 3 unequal setae; lengths of these setae 58, 38, and 22 μm, respectively, from outer to inner; longer outermost seta pinnate, other setae naked. Leg 6 (
<figureCitation id="FD8A2A58FFCDFFD8FF49B548B0C6655F" box="[152,246,1810,1834]" captionStart="Fig" captionStartId="49.[144,177,1592,1614]" captionTargetBox="[265,1295,260,1551]" captionTargetId="figure-167@49.[175,1413,254,1561]" captionTargetPageId="49" captionText="Fig. 31. Asterocheres nodulosus n. sp., female. A, leg 1; B, leg 2; C, endopod of leg 3; D, leg 4; E, exopod of leg 5; F, left side of genital double-somite, dorsal. Scale bars: A-D, 0.05mm; E, F, 0.02mm." figureDoi="http://doi.org/10.5281/zenodo.13344923" httpUri="https://zenodo.org/record/13344923/files/figure.png" pageId="51" pageNumber="236">Fig. 31F</figureCitation>
) represented by 1 pinnate seta and 1 denticle on genital operculum.
</paragraph>
<paragraph id="650E36DDFFCDFFD8FF41B509B395632B" blockId="51.[144,750,261,1996]" lastBlockId="51.[809,1416,261,1995]" pageId="51" pageNumber="236">
<emphasis id="57C5EACFFFCDFFD8FF41B509B0E2651E" bold="true" box="[144,210,1875,1899]" pageId="51" pageNumber="236">Male.</emphasis>
Body (
<figureCitation id="FD8A2A58FFCDFFD8FEF7B508B1BE651F" box="[294,398,1874,1898]" captionStart="Fig" captionStartId="50.[174,207,1926,1948]" captionTargetBox="[268,1350,259,1877]" captionTargetId="figure-13@50.[171,1447,254,1892]" captionTargetPageId="50" captionText="Fig. 32. Asterocheres nodulosus n. sp., male. A, habitus, dorsal; B, urosome, ventral; C, antennule; D, maxilliped; E, endopod of leg 2; F, endopod of leg 3; G, exopod of leg 5; H, leg 6. Scale bars:A, 0.1mm; B, 0.05 mm; C-H, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.13344925" httpUri="https://zenodo.org/record/13344925/files/figure.png" pageId="51" pageNumber="236">Fig. 32A</figureCitation>
) narrower than that of female. Body length 655 μm. Prosome 430 × 297 μm. Urosome (
<figureCitation id="FD8A2A58FFCDFFD8FF49B5C9B0C865DE" box="[152,248,1939,1963]" captionStart="Fig" captionStartId="50.[174,207,1926,1948]" captionTargetBox="[268,1350,259,1877]" captionTargetId="figure-13@50.[171,1447,254,1892]" captionTargetPageId="50" captionText="Fig. 32. Asterocheres nodulosus n. sp., male. A, habitus, dorsal; B, urosome, ventral; C, antennule; D, maxilliped; E, endopod of leg 2; F, endopod of leg 3; G, exopod of leg 5; H, leg 6. Scale bars:A, 0.1mm; B, 0.05 mm; C-H, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.13344925" httpUri="https://zenodo.org/record/13344925/files/figure.png" pageId="51" pageNumber="236">Fig. 32B</figureCitation>
) 5-segmented. Fifth pedigerous somite 85 μm wide. Genital somite globular, 91 × 120 μm, bearing spinules on lateral surfaces. Three abdominal somites 22 × 56, 21 × 51, 30 × 50 μm, respectively. Caudal ramus 22 × 23 μm.
</paragraph>
<paragraph id="650E36DDFFCDFFD8FCEEB33CB4226015" blockId="51.[809,1416,261,1995]" pageId="51" pageNumber="236">
Rostrum as in female. Antennule (
<figureCitation id="FD8A2A58FFCDFFD8FB6FB33CB512630B" box="[1214,1314,358,382]" captionStart="Fig" captionStartId="50.[174,207,1926,1948]" captionTargetBox="[268,1350,259,1877]" captionTargetId="figure-13@50.[171,1447,254,1892]" captionTargetPageId="50" captionText="Fig. 32. Asterocheres nodulosus n. sp., male. A, habitus, dorsal; B, urosome, ventral; C, antennule; D, maxilliped; E, endopod of leg 2; F, endopod of leg 3; G, exopod of leg 5; H, leg 6. Scale bars:A, 0.1mm; B, 0.05 mm; C-H, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.13344925" httpUri="https://zenodo.org/record/13344925/files/figure.png" pageId="51" pageNumber="236">Fig. 32C</figureCitation>
) 18-segmented, with weak geniculation between 15th and 16th segments and strong geniculation between 16th and 17th segments; 9th, 13th, 17th, and terminal segments with 7 setae, 2 setae+ aesthetasc, 2 setae + aesthetasc, and 10 setae, respectively; other segment with 2 setae each; first segment with setules and spinules on anterior margin. Antenna as in female.
</paragraph>
<paragraph id="650E36DDFFCDFFD8FCEEB032B40360B5" blockId="51.[809,1416,261,1995]" pageId="51" pageNumber="236">
Oral siphon, mandible, maxillule, maxilla as in female. Maxilliped (
<figureCitation id="FD8A2A58FFCDFFD8FC7CB0D2B42260D5" box="[941,1042,648,672]" captionStart="Fig" captionStartId="50.[174,207,1926,1948]" captionTargetBox="[268,1350,259,1877]" captionTargetId="figure-13@50.[171,1447,254,1892]" captionTargetPageId="50" captionText="Fig. 32. Asterocheres nodulosus n. sp., male. A, habitus, dorsal; B, urosome, ventral; C, antennule; D, maxilliped; E, endopod of leg 2; F, endopod of leg 3; G, exopod of leg 5; H, leg 6. Scale bars:A, 0.1mm; B, 0.05 mm; C-H, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.13344925" httpUri="https://zenodo.org/record/13344925/files/figure.png" pageId="51" pageNumber="236">Fig. 32D</figureCitation>
) with beak-like proximal process on inner margin of basis.
</paragraph>
<paragraph id="650E36DDFFCDFFD8FCEEB093B53F61D7" blockId="51.[809,1416,261,1995]" pageId="51" pageNumber="236">
Legs 1 and 4 as in female. Endopods of legs 2 and 3 different from those of female. Third endopodal segment of leg 2 (
<figureCitation id="FD8A2A58FFCDFFD8FC1CB153B41F6154" box="[973,1071,777,801]" captionStart="Fig" captionStartId="50.[174,207,1926,1948]" captionTargetBox="[268,1350,259,1877]" captionTargetId="figure-13@50.[171,1447,254,1892]" captionTargetPageId="50" captionText="Fig. 32. Asterocheres nodulosus n. sp., male. A, habitus, dorsal; B, urosome, ventral; C, antennule; D, maxilliped; E, endopod of leg 2; F, endopod of leg 3; G, exopod of leg 5; H, leg 6. Scale bars:A, 0.1mm; B, 0.05 mm; C-H, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.13344925" httpUri="https://zenodo.org/record/13344925/files/figure.png" pageId="51" pageNumber="236">Fig. 32E</figureCitation>
) with mediodistal seta bearing nodule-like proximal expansion and outer distal seta reduced in size. Third endopodal segment of leg 3 (
<figureCitation id="FD8A2A58FFCDFFD8FA8BB110B36461F7" captionStart="Fig" captionStartId="50.[174,207,1926,1948]" captionTargetBox="[268,1350,259,1877]" captionTargetId="figure-13@50.[171,1447,254,1892]" captionTargetPageId="50" captionText="Fig. 32. Asterocheres nodulosus n. sp., male. A, habitus, dorsal; B, urosome, ventral; C, antennule; D, maxilliped; E, endopod of leg 2; F, endopod of leg 3; G, exopod of leg 5; H, leg 6. Scale bars:A, 0.1mm; B, 0.05 mm; C-H, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.13344925" httpUri="https://zenodo.org/record/13344925/files/figure.png" pageId="51" pageNumber="236">Fig. 32F</figureCitation>
) with hook-like, curved outer distal process and 2 small denticles medial to outer distal process.
</paragraph>
<paragraph id="650E36DDFFCDFFD8FCEEB1F1B4AA66F1" blockId="51.[809,1416,261,1995]" pageId="51" pageNumber="236">
Exopodal segment of leg 5 (
<figureCitation id="FD8A2A58FFCDFFD8FBA1B1F1B4E361B6" box="[1136,1235,939,963]" captionStart="Fig" captionStartId="50.[174,207,1926,1948]" captionTargetBox="[268,1350,259,1877]" captionTargetId="figure-13@50.[171,1447,254,1892]" captionTargetPageId="50" captionText="Fig. 32. Asterocheres nodulosus n. sp., male. A, habitus, dorsal; B, urosome, ventral; C, antennule; D, maxilliped; E, endopod of leg 2; F, endopod of leg 3; G, exopod of leg 5; H, leg 6. Scale bars:A, 0.1mm; B, 0.05 mm; C-H, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.13344925" httpUri="https://zenodo.org/record/13344925/files/figure.png" pageId="51" pageNumber="236">Fig. 32G</figureCitation>
) 2.17 times longer than wide (26 × 12 μm), shaped and ornamented as that of female, bearing 2 pinnate distal setae and 1 naked mediodistal seta; lengths of these setae 43, 34, and 21μm long, respectively, from outer to inner. Leg 6 (
<figureCitation id="FD8A2A58FFCDFFD8FA8BB676B3696611" captionStart="Fig" captionStartId="50.[174,207,1926,1948]" captionTargetBox="[268,1350,259,1877]" captionTargetId="figure-13@50.[171,1447,254,1892]" captionTargetPageId="50" captionText="Fig. 32. Asterocheres nodulosus n. sp., male. A, habitus, dorsal; B, urosome, ventral; C, antennule; D, maxilliped; E, endopod of leg 2; F, endopod of leg 3; G, exopod of leg 5; H, leg 6. Scale bars:A, 0.1mm; B, 0.05 mm; C-H, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.13344925" httpUri="https://zenodo.org/record/13344925/files/figure.png" pageId="51" pageNumber="236">Fig. 32H</figureCitation>
) consisting of 2 unequal setae on spinulose outer distal region of genital operculum.
</paragraph>
</subSubSection>
<subSubSection id="2DAB6556FFCDFFDFFCF8B6D7B254638A" lastPageId="52" lastPageNumber="237" pageId="51" pageNumber="236" type="discussion">
<paragraph id="650E36DDFFCDFFD8FCF8B6D7B5176713" blockId="51.[809,1416,261,1995]" pageId="51" pageNumber="236">
<emphasis id="57C5EACFFFCDFFD8FCF8B6D7B3A966D0" bold="true" box="[809,921,1165,1189]" pageId="51" pageNumber="236">Remarks.</emphasis>
<taxonomicName id="A2B14D5EFFCDFFD8FC73B6D6B49466D1" authorityName="Lee" authorityYear="2024" box="[930,1188,1164,1188]" class="Copepoda" family="Asterocheridae" genus="Asterocheres" kingdom="Animalia" order="Siphonostomatoida" pageId="51" pageNumber="236" phylum="Arthropoda" rank="species" species="nodulosus" status="sp. nov.">
<emphasis id="57C5EACFFFCDFFD8FC73B6D6B49466D1" box="[930,1188,1164,1188]" italics="true" pageId="51" pageNumber="236">Asterocheres nodulosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CF657B4FFCDFFD8FB7CB6D7B4D966D1" box="[1197,1257,1165,1188]" pageId="51" pageNumber="236" rank="species">n. sp.</taxonomicNameLabel>
appears to be most similar to three congeners: 
<taxonomicName id="A2B14D5EFFCDFFD8FB58B6F7B4F866B0" authorityName="Lee" authorityYear="2024" box="[1161,1224,1197,1221]" class="Copepoda" family="Asterocheridae" genus="Asterocheres" kingdom="Animalia" order="Siphonostomatoida" pageId="51" pageNumber="236" phylum="Arthropoda" rank="species" species="fici" status="n. sp.">
<emphasis id="57C5EACFFFCDFFD8FB58B6F7B4F866B0" box="[1161,1224,1197,1221]" italics="true" pageId="51" pageNumber="236">A. fici</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CF657B4FFCDFFD8FB1FB6F7B53966B1" box="[1230,1289,1197,1220]" pageId="51" pageNumber="236" rank="species">n. sp.</taxonomicNameLabel>
, sharing 14 same character states in 17 characters with the new species, and 
<taxonomicName id="A2B14D5EFFCDFFD8FC42B6B7B4276770" authorityName="Ho" authorityYear="1984" box="[915,1047,1261,1285]" class="Copepoda" family="Asterocheridae" genus="Asterocheres" kingdom="Animalia" order="Siphonostomatoida" pageId="51" pageNumber="236" phylum="Arthropoda" rank="species" species="aesthetes">
<emphasis id="57C5EACFFFCDFFD8FC42B6B7B4276770" box="[915,1047,1261,1285]" italics="true" pageId="51" pageNumber="236">A. aesthetes</emphasis>
</taxonomicName>
and 
<taxonomicName id="A2B14D5EFFCDFFD8FB81B6B7B4FE6770" authorityName="Kim" authorityYear="2016" box="[1104,1230,1261,1285]" class="Copepoda" family="Asterocheridae" genus="Asterocheres" kingdom="Animalia" order="Siphonostomatoida" pageId="51" pageNumber="236" phylum="Arthropoda" rank="species" species="horridus">
<emphasis id="57C5EACFFFCDFFD8FB81B6B7B4FE6770" box="[1104,1230,1261,1285]" italics="true" pageId="51" pageNumber="236">A. horridus</emphasis>
</taxonomicName>
each sharing 13 same character states in 17 characters with the new species (see 
<tableCitation id="28330366FFCDFFD8FC5AB774B3EE6733" box="[907,990,1326,1350]" captionStart="Table 2" captionStartId="2.[92,145,474,499]" captionText="Table 2. A character matrix of Asterocheres species (“-” indicates missing data)." pageId="51" pageNumber="236">Table 2</tableCitation>
). These three species are distinguished from the new species by the following features:
</paragraph>
<paragraph id="650E36DDFFCDFFD8FCEEB734B41164DC" blockId="51.[809,1416,261,1995]" pageId="51" pageNumber="236">
<taxonomicName id="A2B14D5EFFCDFFD8FCEEB734B40567F3" authorityName="Ho" authorityYear="1984" box="[831,1077,1390,1414]" class="Copepoda" family="Asterocheridae" genus="Asterocheres" kingdom="Animalia" order="Siphonostomatoida" pageId="51" pageNumber="236" phylum="Arthropoda" rank="species" species="aesthetes">
<emphasis id="57C5EACFFFCDFFD8FCEEB734B40567F3" box="[831,1077,1390,1414]" italics="true" pageId="51" pageNumber="236">Asterocheres aesthetes</emphasis>
</taxonomicName>
has caudal rami of the female which are wider than long and a 2-segmented mandibular palp. 
<taxonomicName id="A2B14D5EFFCDFFD8FC48B7F5B47D67B2" authorityName="Lee" authorityYear="2024" box="[921,1101,1455,1479]" class="Copepoda" family="Asterocheridae" genus="Asterocheres" kingdom="Animalia" order="Siphonostomatoida" pageId="51" pageNumber="236" phylum="Arthropoda" rank="species" species="fici" status="sp. nov.">
<emphasis id="57C5EACFFFCDFFD8FC48B7F5B47D67B2" box="[921,1101,1455,1479]" italics="true" pageId="51" pageNumber="236">Asterocheres fici</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CF657B4FFCDFFD8FB87B7F5B4A267B3" box="[1110,1170,1455,1478]" pageId="51" pageNumber="236" rank="species">n. sp.</taxonomicNameLabel>
has the terminal claw of the female antenna which is distinctly longer than the first endopodal segment and the exopodal segment of female leg 5 which is 2.83 times longer than wide. 
<taxonomicName id="A2B14D5EFFCDFFD8FA97B44AB3EE643D" authorityName="Kim" authorityYear="2016" class="Copepoda" family="Asterocheridae" genus="Asterocheres" kingdom="Animalia" order="Siphonostomatoida" pageId="51" pageNumber="236" phylum="Arthropoda" rank="species" species="horridus">
<emphasis id="57C5EACFFFCDFFD8FA97B44AB3EE643D" italics="true" pageId="51" pageNumber="236">Asterocheres horridus</emphasis>
</taxonomicName>
has a longer oral siphon which extends to leg 1, a 2-segmented mandibular palp, and the exopodal segment of female leg 5 which is 2.90 times longer than wide (
<bibRefCitation id="01204B2CFFCDFFD8FC4CB4CBB42564DC" author="Kim, I. - H." box="[925,1045,1681,1705]" pageId="51" pageNumber="236" pagination="393 - 442" refId="ref37874" refString="Kim, I. - H. 2016. Siphonostomatoid copepods (Crustacea) mainly associated with marine invertebrates from Kore- an waters. Journal of Species Research 5 (3): 393 - 442." type="journal article" year="2016">Kim, 2016</bibRefCitation>
).
</paragraph>
<paragraph id="650E36DDFFCDFFDFFCEEB4EBB254638A" blockId="51.[809,1416,261,1995]" lastBlockId="52.[174,780,261,511]" lastPageId="52" lastPageNumber="237" pageId="51" pageNumber="236">
The distinctiveness of the above three species and 
<taxonomicName id="A2B14D5EFFCDFFD8FABEB4EBB3AB649C" authorityName="Lee" authorityYear="2024" class="Copepoda" family="Asterocheridae" genus="Asterocheres" kingdom="Animalia" order="Siphonostomatoida" pageId="51" pageNumber="236" phylum="Arthropoda" rank="species" species="nodulosus" status="n. sp.">
<emphasis id="57C5EACFFFCDFFD8FABEB4EBB3AB649C" italics="true" pageId="51" pageNumber="236">A. nodulosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CF657B4FFCDFFD8FC74B488B3D4649C" box="[933,996,1746,1769]" pageId="51" pageNumber="236" rank="species">n. sp.</taxonomicNameLabel>
can be confirmed by their different forms of legs 2 and 3 of the male, as follows: (1) in the third endopod of male leg 2, the outer distal process is expanded with a small tubercle in 
<taxonomicName id="A2B14D5EFFCDFFD8FB77B568B516653F" authorityName="Ho" authorityYear="1984" box="[1190,1318,1842,1866]" class="Copepoda" family="Asterocheridae" genus="Asterocheres" kingdom="Animalia" order="Siphonostomatoida" pageId="51" pageNumber="236" phylum="Arthropoda" rank="species" species="aesthetes">
<emphasis id="57C5EACFFFCDFFD8FB77B568B516653F" box="[1190,1318,1842,1866]" italics="true" pageId="51" pageNumber="236">A. aesthetes</emphasis>
</taxonomicName>
, thinned in 
<taxonomicName id="A2B14D5EFFCDFFD8FC94B509B38E651F" authorityName="Kim" authorityYear="2016" box="[837,958,1874,1898]" class="Copepoda" family="Asterocheridae" genus="Asterocheres" kingdom="Animalia" order="Siphonostomatoida" pageId="51" pageNumber="236" phylum="Arthropoda" rank="species" species="horridus">
<emphasis id="57C5EACFFFCDFFD8FC94B509B38E651F" box="[837,958,1874,1898]" italics="true" pageId="51" pageNumber="236">A. horridus</emphasis>
</taxonomicName>
, and slightly curved in 
<taxonomicName id="A2B14D5EFFCDFFD8FB6DB509B4CB651F" authorityName="Lee" authorityYear="2024" box="[1212,1275,1874,1898]" class="Copepoda" family="Asterocheridae" genus="Asterocheres" kingdom="Animalia" order="Siphonostomatoida" pageId="51" pageNumber="236" phylum="Arthropoda" rank="species" species="fici" status="n. sp.">
<emphasis id="57C5EACFFFCDFFD8FB6DB509B4CB651F" box="[1212,1275,1874,1898]" italics="true" pageId="51" pageNumber="236">A. fici</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CF657B4FFCDFFD8FAD0B509B50B651F" box="[1281,1339,1875,1898]" pageId="51" pageNumber="236" rank="species">n. sp.</taxonomicNameLabel>
and 
<taxonomicName id="A2B14D5EFFCDFFD8FABEB509B3A565FF" authorityName="Lee" authorityYear="2024" class="Copepoda" family="Asterocheridae" genus="Asterocheres" kingdom="Animalia" order="Siphonostomatoida" pageId="51" pageNumber="236" phylum="Arthropoda" rank="species" species="nodulosus" status="n. sp.">
<emphasis id="57C5EACFFFCDFFD8FABEB509B3A565FF" italics="true" pageId="51" pageNumber="236">A. nodulosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CF657B4FFCDFFD8FC4FB529B3EC65FF" box="[926,988,1907,1930]" pageId="51" pageNumber="236" rank="species">n. sp.</taxonomicNameLabel>
; (2) in the third endopodal segment of male leg 3, the outer distal process is slightly curved in 
<taxonomicName id="A2B14D5EFFCDFFD8FCFAB5E9B39C65BE" authorityName="Ho" authorityYear="1984" box="[811,940,1971,1995]" class="Copepoda" family="Asterocheridae" genus="Asterocheres" kingdom="Animalia" order="Siphonostomatoida" pageId="51" pageNumber="236" phylum="Arthropoda" rank="species" species="aesthetes">
<emphasis id="57C5EACFFFCDFFD8FCFAB5E9B39C65BE" box="[811,940,1971,1995]" italics="true" pageId="51" pageNumber="236">A. aesthetes</emphasis>
</taxonomicName>
, 
<taxonomicName id="A2B14D5EFFCDFFD8FC6DB5E9B3CD65BE" authorityName="Lee" authorityYear="2024" box="[956,1021,1971,1995]" class="Copepoda" family="Asterocheridae" genus="Asterocheres" kingdom="Animalia" order="Siphonostomatoida" pageId="51" pageNumber="236" phylum="Arthropoda" rank="species" species="fici" status="n. sp.">
<emphasis id="57C5EACFFFCDFFD8FC6DB5E9B3CD65BE" box="[956,1021,1971,1995]" italics="true" pageId="51" pageNumber="236">A. fici</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CF657B4FFCDFFD8FBD7B5EEB47365BE" box="[1030,1091,1972,1995]" pageId="51" pageNumber="236" rank="species">n. sp.</taxonomicNameLabel>
, and 
<taxonomicName id="A2B14D5EFFCDFFD8FB50B5E9B53E65BE" authorityName="Lee" authorityYear="2024" box="[1153,1294,1971,1995]" class="Copepoda" family="Asterocheridae" genus="Asterocheres" kingdom="Animalia" order="Siphonostomatoida" pageId="51" pageNumber="236" phylum="Arthropoda" rank="species" species="nodulosus" status="n. sp.">
<emphasis id="57C5EACFFFCDFFD8FB50B5E9B53E65BE" box="[1153,1294,1971,1995]" italics="true" pageId="51" pageNumber="236">A. nodulosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CF657B4FFCDFFD8FAC9B5EEB56565BE" box="[1304,1365,1972,1995]" pageId="51" pageNumber="236" rank="species">n. sp.</taxonomicNameLabel>
, but not modified in 
<taxonomicName id="A2B14D5EFFCAFFDFFE8EB35FB1EB6368" authorityName="Kim" authorityYear="2016" box="[351,475,261,285]" class="Copepoda" family="Asterocheridae" genus="Asterocheres" kingdom="Animalia" order="Siphonostomatoida" pageId="52" pageNumber="237" phylum="Arthropoda" rank="species" species="horridus">
<emphasis id="57C5EACFFFCAFFDFFE8EB35FB1EB6368" box="[351,475,261,285]" italics="true" pageId="52" pageNumber="237">A. horridus</emphasis>
</taxonomicName>
; (3) the outer distal seta of the third endopodal segment of male leg 3 is minimized in 
<taxonomicName id="A2B14D5EFFCAFFDFFF1AB31CB13B6328" authorityName="Lee" authorityYear="2024" box="[203,267,325,349]" class="Copepoda" family="Asterocheridae" genus="Asterocheres" kingdom="Animalia" order="Siphonostomatoida" pageId="52" pageNumber="237" phylum="Arthropoda" rank="species" species="fici" status="n. sp.">
<emphasis id="57C5EACFFFCAFFDFFF1AB31CB13B6328" box="[203,267,325,349]" italics="true" pageId="52" pageNumber="237">A. fici</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CF657B4FFCAFFDFFEC3B31CB17D6328" box="[274,333,326,349]" pageId="52" pageNumber="237" rank="species">n. sp.</taxonomicNameLabel>
and 
<taxonomicName id="A2B14D5EFFCAFFDFFE54B31CB2206328" authorityName="Lee" authorityYear="2024" box="[389,528,325,349]" class="Copepoda" family="Asterocheridae" genus="Asterocheres" kingdom="Animalia" order="Siphonostomatoida" pageId="52" pageNumber="237" phylum="Arthropoda" rank="species" species="nodulosus" status="n. sp.">
<emphasis id="57C5EACFFFCAFFDFFE54B31CB2206328" box="[389,528,325,349]" italics="true" pageId="52" pageNumber="237">A. nodulosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CF657B4FFCAFFDFFDC6B31CB2616328" box="[535,593,326,349]" pageId="52" pageNumber="237" rank="species">n. sp.</taxonomicNameLabel>
but normal in 
<taxonomicName id="A2B14D5EFFCAFFDFFD24B31CB120630B" authorityName="Ho" authorityYear="1984" class="Copepoda" family="Asterocheridae" genus="Asterocheres" kingdom="Animalia" order="Siphonostomatoida" pageId="52" pageNumber="237" phylum="Arthropoda" rank="species" species="aesthetes">
<emphasis id="57C5EACFFFCAFFDFFD24B31CB120630B" italics="true" pageId="52" pageNumber="237">A. aesthetes</emphasis>
</taxonomicName>
and 
<taxonomicName id="A2B14D5EFFCAFFDFFE99B33CB1F2630B" authorityName="Kim" authorityYear="2016" box="[328,450,358,382]" class="Copepoda" family="Asterocheridae" genus="Asterocheres" kingdom="Animalia" order="Siphonostomatoida" pageId="52" pageNumber="237" phylum="Arthropoda" rank="species" species="horridus">
<emphasis id="57C5EACFFFCAFFDFFE99B33CB1F2630B" box="[328,450,358,382]" italics="true" pageId="52" pageNumber="237">A. horridus</emphasis>
</taxonomicName>
; and finally, the presence of a nodule-like proximal expansion on the inner distal seta of the third endopodal segment of male leg 2 is a unique feature of 
<taxonomicName id="A2B14D5EFFCAFFDFFECAB39DB19563AB" authorityName="Lee" authorityYear="2024" box="[283,421,454,478]" class="Copepoda" family="Asterocheridae" genus="Asterocheres" kingdom="Animalia" order="Siphonostomatoida" pageId="52" pageNumber="237" phylum="Arthropoda" rank="species" species="nodulosus" status="n. sp.">
<emphasis id="57C5EACFFFCAFFDFFECAB39DB19563AB" box="[283,421,454,478]" italics="true" pageId="52" pageNumber="237">A. nodulosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CF657B4FFCAFFDFFE7DB39DB1D763AB" box="[428,487,455,478]" pageId="52" pageNumber="237" rank="species">n. sp.</taxonomicNameLabel>
, which is easily observable under the microscope without dissection.
</paragraph>
</subSubSection>
</treatment>
</document>