188 lines
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188 lines
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<document id="4BA05FC1F11BA457CEA33E4412D217A6" ID-DOI="10.1206/350.1" ID-ISSN="0003-0090" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721339324786" checkinUser="felipe" docAuthor="Gaffney, Eugene S., Meylan, Peter A., Wood, Roger C., Simons, Elwyn & De Almeida Campos, Diogenes" docDate="2011" docId="C95DDC2BFFA75E52FFABA5F19C8AD1A7" docLanguage="en" docName="BulAmeMusNatHis.2011.B350.1-237.pdf" docOrigin="Bulletin of the American Museum of Natural History 2011 (350)" docSource="http://www.bioone.org/doi/abs/10.1206/350.1" docStyle="DocumentStyle:915933466F796C9C739DF4DB6B8DCFA6.9:BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleId="915933466F796C9C739DF4DB6B8DCFA6" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleVersion="9" docTitle="Dacquemys paleomorpha Williams 1954" docType="treatment" docVersion="1" lastPageNumber="34" masterDocId="3564A453FF855E70FFD9A1399E7CD533" masterDocTitle="Evolution Of The Side-Necked Turtles: The Family Podocnemididae" masterLastPageNumber="237" masterPageNumber="1" pageNumber="34" updateTime="1721367619560" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-4.0">
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<mods:title id="2B8D3C57C61EE60C6E73F4309189F75C">Evolution Of The Side-Necked Turtles: The Family Podocnemididae</mods:title>
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<mods:namePart id="D21D541E2A5688161947E55DD3CF7A5D">Gaffney, Eugene S.</mods:namePart>
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<mods:namePart id="998DAFC551BAF0ED7CED0239B708D962">Meylan, Peter A.</mods:namePart>
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<mods:namePart id="2AADBB7C02367202A3DA0DC18C5D9ADC">Wood, Roger C.</mods:namePart>
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<mods:namePart id="FAC2D2DEF6606581042941FE0561E857">Simons, Elwyn</mods:namePart>
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<mods:namePart id="8B2BE4F24AE733DDD769F999C6A022A0">De Almeida Campos, Diogenes</mods:namePart>
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<mods:title id="21FF9D208D07E65B7B532E75AB33D2EF">Bulletin of the American Museum of Natural History</mods:title>
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<mods:date id="360E73FA678907B559A09543E1FD8D6D">2011</mods:date>
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<mods:number id="7BB1912CE9FAC69B5B964D5039517914">2011-04-29</mods:number>
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<mods:number id="8F91DA8A8FD301C1964E80EDEF7EA59C">2011</mods:number>
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<mods:number id="F5B2F434FF6A8B22E494DE53C7E3B69F">350</mods:number>
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<mods:url id="FC58F5400254600836DB60E3C122EAD1">http://www.bioone.org/doi/abs/10.1206/350.1</mods:url>
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<mods:classification id="C70F3450CB3015AD45FB5F88927C114B">journal article</mods:classification>
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<mods:identifier id="547E20D7BDA2942ECE65882F796F1CE2" type="DOI">10.1206/350.1</mods:identifier>
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<mods:identifier id="3423BB344647A1C79555C74E0DE1B4E0" type="ISSN">0003-0090</mods:identifier>
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<treatment id="C95DDC2BFFA75E52FFABA5F19C8AD1A7" LSID="urn:lsid:plazi:treatment:C95DDC2BFFA75E52FFABA5F19C8AD1A7" httpUri="http://treatment.plazi.org/id/C95DDC2BFFA75E52FFABA5F19C8AD1A7" lastPageNumber="34" pageId="34" pageNumber="34">
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<subSubSection id="09EE3EB6FFA75E52FFABA5F19C2AD1EE" box="[114,598,1222,1245]" pageId="34" pageNumber="34" type="nomenclature">
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<paragraph id="414B6D3DFFA75E52FFABA5F19C2AD1EE" blockId="34.[114,598,1222,1245]" box="[114,598,1222,1245]" pageId="34" pageNumber="34">
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<heading id="1A03DA51FFA75E52FFABA5F19C2AD1EE" box="[114,598,1222,1245]" centered="true" fontSize="9" level="2" pageId="34" pageNumber="34" reason="2">
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<taxonomicName id="86F416BEFFA75E52FFABA5F19C2AD1EE" authority="Williams, 1954 b" authorityName="Williams" authorityYear="1954" box="[114,598,1222,1245]" class="Reptilia" family="Podocnemididae" genus="Dacquemys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="34" pageNumber="34" phylum="Chordata" rank="species" species="paleomorpha">
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<emphasis id="7380B12FFFA75E52FFABA5F19FECD1EE" box="[114,400,1222,1245]" italics="true" pageId="34" pageNumber="34">Dacquemys paleomorpha</emphasis>
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<bibRefCitation id="256510CCFFA75E52FE40A5FE9C2AD1EE" author="Williams, E. E." box="[409,598,1223,1245]" pageId="34" pageNumber="34" pagination="1 - 9" refId="ref121605" refString="Williams, E. E. 1954 b. New or redescribed pelomedusid skulls from the Tertiary of Africa and Asia (Testudines, Pelomedusidae) 1. Dacquemys paleomorpha, new genus, new species from the Lower Oligocene of the Fayum, Egypt. Breviora, 35: 1 - 9." type="journal article" year="1954">Williams, 1954b</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="09EE3EB6FFA75E52FFA3A5CA9D54D7C8" pageId="34" pageNumber="34" type="materials_examined">
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<paragraph id="414B6D3DFFA75E52FFA3A5CA9C42D0AF" blockId="34.[93,620,1267,1759]" pageId="34" pageNumber="34">
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<typeStatus id="9E4FD39FFFA75E52FFA3A5CA9EC5D03A" box="[122,185,1267,1291]" pageId="34" pageNumber="34">TYPE</typeStatus>
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</smallCapsWord>
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<smallCapsWord id="47ADFBE1FFA75E52FF1FA5CA9F41D039" baselines="1285,1286" box="[198,317,1267,1291]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Specimen" pageId="34" pageNumber="34">SPECIMEN</smallCapsWord>
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: SMNS 12645, a nearly complete skull without lower jaws, lacking parts of the left side and the central basicranial region (Dacque´, 1912: pl. 2, figs. 6–8;
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<bibRefCitation id="256510CCFFA75E52FF4DA4539F47D04C" author="Gaffney, E. S." box="[148,315,1386,1408]" pageId="34" pageNumber="34" pagination="65 - 376" refId="ref116172" refString="Gaffney, E. S. 1979. Comparative cranial morphology of recent and fossil turtles. Bulletin of the American Museum of Natural History 164 (2): 65 - 376." type="journal article" year="1979">Gaffney, 1979</bibRefCitation>
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: fig. 128;
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<bibRefCitation id="256510CCFFA75E52FE70A4539C14D0B3" author="Williams, E. E." box="[425,616,1386,1408]" pageId="34" pageNumber="34" pagination="1 - 9" refId="ref121605" refString="Williams, E. E. 1954 b. New or redescribed pelomedusid skulls from the Tertiary of Africa and Asia (Testudines, Pelomedusidae) 1. Dacquemys paleomorpha, new genus, new species from the Lower Oligocene of the Fayum, Egypt. Breviora, 35: 1 - 9." type="journal article" year="1954">Williams, 1954b</bibRefCitation>
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: pl. 1; Gaffney et al., 2002: figs. 2 and 3).
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</paragraph>
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<paragraph id="414B6D3DFFA75E52FFA3A49A9A71D434" blockId="34.[93,620,1267,1759]" lastBlockId="34.[667,1194,213,1172]" pageId="34" pageNumber="34">
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<typeStatus id="9E4FD39FFFA75E52FFA3A49A9EC5D08A" box="[122,185,1443,1467]" pageId="34" pageNumber="34">TYPE</typeStatus>
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</smallCapsWord>
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<smallCapsWord id="47ADFBE1FFA75E52FF1FA49A9F42D089" baselines="1461,1462" box="[198,318,1443,1467]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Locality" pageId="34" pageNumber="34">LOCALITY</smallCapsWord>
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<smallCapsWord id="47ADFBE1FFA75E52FE92A4909F03D089" baselines="1462" box="[331,383,1449,1466]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="and" pageId="34" pageNumber="34">AND</smallCapsWord>
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<smallCapsWord id="47ADFBE1FFA75E52FE55A49A9F81D089" baselines="1461,1462" box="[396,509,1443,1467]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Horizon" pageId="34" pageNumber="34">HORIZON</smallCapsWord>
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: Dacqué (1912) gave the locality at first in reference to a shell: ‘‘Unteroligocäne Fluviomarinestuffe nördlich der Birket Qerun; nordlich non Tamieh; ostlich vom Schweinforthplateau. Schädel: aus derselben stufe bei Dimêh. Alles Fayum’’ (ibid., p. 310).
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<bibRefCitation id="256510CCFFA75E52FE45A76D9C1BD359" author="Williams, E. E." box="[412,615,1620,1642]" pageId="34" pageNumber="34" pagination="1 - 9" refId="ref121605" refString="Williams, E. E. 1954 b. New or redescribed pelomedusid skulls from the Tertiary of Africa and Asia (Testudines, Pelomedusidae) 1. Dacquemys paleomorpha, new genus, new species from the Lower Oligocene of the Fayum, Egypt. Breviora, 35: 1 - 9." type="journal article" year="1954">Williams (1954b</bibRefCitation>
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, repeated in
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<bibRefCitation id="256510CCFFA75E52FF28A7489FE0D3B4" author="Gaffney, E. S." box="[241,412,1649,1671]" pageId="34" pageNumber="34" pagination="65 - 376" refId="ref116172" refString="Gaffney, E. S. 1979. Comparative cranial morphology of recent and fossil turtles. Bulletin of the American Museum of Natural History 164 (2): 65 - 376." type="journal article" year="1979">Gaffney, 1979</bibRefCitation>
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) has the locality as ‘‘Diieh’’ which seems to be a misspelling of ‘‘Dimêh.’’ The reference to the skull ‘‘from the same sediment near Dimêh’’ presumably means the Qasr el-Sagha or Birket Qarun Formations of late Eocene age.
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</paragraph>
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<paragraph id="414B6D3DFFA75E52FD61A0379A2CD4E6" blockId="34.[667,1194,213,1172]" pageId="34" pageNumber="34">
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<smallCapsWord id="47ADFBE1FFA75E52FC9CA0379DB9D417" baselines="288,288" box="[837,965,270,294]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Material" pageId="34" pageNumber="34">MATERIAL</smallCapsWord>
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:
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<collectionCode id="27E5F5F8FFA75E52FC01A0299A6FD416" box="[984,1043,272,293]" pageId="34" pageNumber="34">DPC</collectionCode>
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5986 (Gaffney et al., 2002: figs. 4, 5), partial skull lacking anterior part of palate and some of left side. Field No. 86-292, collected by Alex van Nievelt. B-4 (lower Jebel Qatrani Fm., 1/
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<quantity id="860CC0D8FFA75E52FD77A09B9C86D48B" box="[686,762,418,440]" metricMagnitude="3" metricUnit="m" metricValue="6.437376" pageId="34" pageNumber="34" unit="mi" value="4.0">4 mile</quantity>
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southeast of A),
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<collectionCode id="27E5F5F8FFA75E52FC0DA09B9A4DD484" box="[980,1073,418,439]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="34" pageNumber="34">AMNH</collectionCode>
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quarry B area, early Oligocene (
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<bibRefCitation id="256510CCFFA75E52FC7AA0869A38D4E6" author="Seiffert, E. R." box="[931,1092,447,469]" pageId="34" pageNumber="34" pagination="5000 - 5005" refId="ref120383" refString="Seiffert, E. R. 2006. Revised age estimates for the later Paleogene mammal faunas of Egypt and Oman. Proceedings of the National Academy of Sciences (USA) 103 (13): 5000 - 5005." type="journal article" year="2006">Seiffert, 2006</bibRefCitation>
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).
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</paragraph>
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<paragraph id="414B6D3DFFA75E52FD61A0E49D76D7B6" blockId="34.[667,1194,213,1172]" pageId="34" pageNumber="34">
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Possibly
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<collectionCode id="27E5F5F8FFA75E52FCF7A0E79DF4D4C0" box="[814,904,478,499]" pageId="34" pageNumber="34">BMNH</collectionCode>
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R3346, a lower jaw figured by
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<bibRefCitation id="256510CCFFA75E52FCC4A0C39DA3D723" author="Williams, E. E." box="[797,991,506,528]" pageId="34" pageNumber="34" pagination="1 - 9" refId="ref121605" refString="Williams, E. E. 1954 b. New or redescribed pelomedusid skulls from the Tertiary of Africa and Asia (Testudines, Pelomedusidae) 1. Dacquemys paleomorpha, new genus, new species from the Lower Oligocene of the Fayum, Egypt. Breviora, 35: 1 - 9." type="journal article" year="1954">Williams (1954b</bibRefCitation>
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; previously identified by
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<bibRefCitation id="256510CCFFA75E52FCC4A32E9DA7D71E" author="Andrews, C. W." box="[797,987,535,557]" pageId="34" pageNumber="34" refId="ref114233" refString="Andrews, C. W. 1906. A descriptive catalog of the Tertiary Vertebrata of the Fayum, Egypt. London: Trustees of the British Museum (Natural History). 324 pp." type="book" year="1906">Andrews, 1906</bibRefCitation>
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, as
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<taxonomicName id="86F416BEFFA75E52FBFBA32E9D6DD779" authorityName="Andrews" authorityYear="1903" class="Testudines" family="Podocnemididae" genus="Podocnemis" kingdom="Animalia" order="Pleurodira" pageId="34" pageNumber="34" phylum="Chordata" rank="species" species="fajumensis">
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<emphasis id="7380B12FFFA75E52FBFBA32E9D6DD779" italics="true" pageId="34" pageNumber="34">Podocnemis fajumensis</emphasis>
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</taxonomicName>
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) that fits the triturating surface of the maxilla in
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<taxonomicName id="86F416BEFFA75E52FCACA36A9D84D75B" authorityName="Williams" authorityYear="1954" box="[885,1016,595,616]" class="Reptilia" family="Podocnemididae" genus="Dacquemys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="34" pageNumber="34" phylum="Chordata" rank="genus">
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<emphasis id="7380B12FFFA75E52FCACA36A9D84D75B" box="[885,1016,595,616]" italics="true" pageId="34" pageNumber="34">Dacquemys</emphasis>
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</taxonomicName>
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, as argued by Williams.
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</paragraph>
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<smallCapsWord id="47ADFBE1FFA75E52FCECA3B29DF8D791" baselines="669,670" box="[821,900,651,675]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Work" pageId="34" pageNumber="34">WORK</smallCapsWord>
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:
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<bibRefCitation id="256510CCFFA75E52FC4DA3B49A22D790" author="Williams, E. E." box="[916,1118,653,675]" pageId="34" pageNumber="34" pagination="1 - 9" refId="ref121605" refString="Williams, E. E. 1954 b. New or redescribed pelomedusid skulls from the Tertiary of Africa and Asia (Testudines, Pelomedusidae) 1. Dacquemys paleomorpha, new genus, new species from the Lower Oligocene of the Fayum, Egypt. Breviora, 35: 1 - 9." type="journal article" year="1954">Williams (1954b)</bibRefCitation>
|
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. Gaffney et al. (2002) provides a redescription based on
|
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|
<specimenCount id="57F2A6B4FFA75E52FCD3A3F19DCED7ED" box="[778,946,712,734]" count="2" pageId="34" pageNumber="34" type="generic">two specimens</specimenCount>
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|
and a reconstruction of the skull.
|
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</paragraph>
|
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|
|
</subSubSection>
|
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|
<subSubSection id="09EE3EB6FFA75E52FD61A2389C8AD1A7" pageId="34" pageNumber="34" type="discussion">
|
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|
<paragraph id="414B6D3DFFA75E52FD61A2389C8AD1A7" blockId="34.[667,1194,213,1172]" pageId="34" pageNumber="34">
|
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|
<smallCapsWord id="47ADFBE1FFA75E52FD61A2389D39D62B" baselines="787,788" box="[696,837,769,793]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Discussion" pageId="34" pageNumber="34">DISCUSSION</smallCapsWord>
|
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: There is no support for the earlier suggestions that the shell for
|
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<taxonomicName id="86F416BEFFA75E52FB8FA2189D0DD667" authorityName="Williams" authorityYear="1954" class="Reptilia" family="Podocnemididae" genus="Dacquemys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="34" pageNumber="34" phylum="Chordata" rank="species" species="paleomorpha">
|
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<emphasis id="7380B12FFFA75E52FB8FA2189D0DD667" italics="true" pageId="34" pageNumber="34">Dacquemys paleomorpha</emphasis>
|
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|
</taxonomicName>
|
|||
|
|
is the shell-based species,
|
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|
|
<taxonomicName id="86F416BEFFA75E52FD42A2639A1ED643" authority="Andrews, 1903" authorityName="Andrews" authorityYear="1903" box="[667,1122,858,881]" class="Testudines" family="Podocnemididae" genus="Podocnemis" kingdom="Animalia" order="Pleurodira" pageId="34" pageNumber="34" phylum="Chordata" rank="species" species="fajumensis">
|
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|
|
<emphasis id="7380B12FFFA75E52FD42A2639DD9D643" box="[667,933,858,881]" italics="true" pageId="34" pageNumber="34">Podocnemis fajumensis</emphasis>
|
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|
|
<bibRefCitation id="256510CCFFA75E52FC69A2629A1ED643" author="Andrews, C. W." box="[944,1122,859,881]" pageId="34" pageNumber="34" pagination="115 - 122" refId="ref114195" refString="Andrews, C. W. 1903. On some pleurodiran chelonians from the Eocene of the Fayum, Egypt. Annals and Magazine of Natural History Series, (7) 11: 115 - 122." type="journal article" year="1903">Andrews, 1903</bibRefCitation>
|
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|
|
</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation id="256510CCFFA75E52FBACA2629D48D6BD" author="Williams, E. E." pageId="34" pageNumber="34" pagination="1 - 9" refId="ref121605" refString="Williams, E. E. 1954 b. New or redescribed pelomedusid skulls from the Tertiary of Africa and Asia (Testudines, Pelomedusidae) 1. Dacquemys paleomorpha, new genus, new species from the Lower Oligocene of the Fayum, Egypt. Breviora, 35: 1 - 9." type="journal article" year="1954">Williams, 1954b</bibRefCitation>
|
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|
|
; Lapparent
|
|||
|
|
<bibRefCitation id="256510CCFFA75E52FC12A2419A16D6BE" author="Lapparent de Broin, F. de" box="[971,1130,888,910]" pageId="34" pageNumber="34" pagination="43 - 82" refId="ref117962" refString="Lapparent de Broin, F. de, 2000 a. African chelonians from the Jurassic to the present: phases of development and preliminary catalogue of the fossil record. Palaeontologia Africana 36: 43 - 82." type="journal article" year="2000">Broin, 2000a</bibRefCitation>
|
|||
|
|
). We argue below that
|
|||
|
|
<taxonomicName id="86F416BEFFA75E52FCB6A2AC9D9AD699" authorityName="Andrews" authorityYear="1903" box="[879,998,917,938]" class="Testudines" family="Podocnemididae" genus="Podocnemis" kingdom="Animalia" order="Pleurodira" pageId="34" pageNumber="34" phylum="Chordata" rank="species" species="fajumensis">
|
|||
|
|
<emphasis id="7380B12FFFA75E52FCB6A2AC9D9AD699" box="[879,998,917,938]" italics="true" pageId="34" pageNumber="34">fajumensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is the shell of a
|
|||
|
|
<taxonomicName id="86F416BEFFA75E52FD42A2889D0FD6FB" box="[667,883,945,968]" class="Reptilia" family="Podocnemididae" genus="Neochelys" kingdom="Animalia" order="Testudines" pageId="34" pageNumber="34" phylum="Chordata" rank="species" species="undetermined">
|
|||
|
|
<emphasis id="7380B12FFFA75E52FD42A2889D73D6FB" box="[667,783,945,968]" italics="true" pageId="34" pageNumber="34">Neochelys</emphasis>
|
|||
|
|
species.
|
|||
|
|
</taxonomicName>
|
|||
|
|
We have concluded that based on skull morphology,
|
|||
|
|
<taxonomicName id="86F416BEFFA75E52FC3AA2E99A1AD6D6" authorityName="Williams" authorityYear="1954" box="[995,1126,976,997]" class="Reptilia" family="Podocnemididae" genus="Dacquemys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="34" pageNumber="34" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis id="7380B12FFFA75E52FC3AA2E99A1AD6D6" box="[995,1126,976,997]" italics="true" pageId="34" pageNumber="34">Dacquemys</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is the sister taxon to UCMP 42008, which has a very unusual shell similar to that of
|
|||
|
|
<emphasis id="7380B12FFFA75E52FB81A5309D6BD10E" italics="true" pageId="34" pageNumber="34">
|
|||
|
|
<taxonomicName id="86F416BEFFA75E52FB81A5309D6FD10E" authorityName="Gaffney & Meylan & Wood & Simons & De Almeida Campos" authorityYear="2011" class="Testudines" family="Podocnemididae" genus="Albertwoodemys" higherTaxonomySource="CoL" kingdom="Animalia" order="Pleurodira" pageId="34" pageNumber="34" phylum="Chordata" rank="genus">Albertwoodemys</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</emphasis>
|
|||
|
|
The possibility exists that
|
|||
|
|
<taxonomicName id="86F416BEFFA75E52FB8FA5119CA8D169" authorityName="Williams" authorityYear="1954" class="Reptilia" family="Podocnemididae" genus="Dacquemys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="34" pageNumber="34" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis id="7380B12FFFA75E52FB8FA5119CA8D169" italics="true" pageId="34" pageNumber="34">Dacquemys</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is the skull for
|
|||
|
|
<emphasis id="7380B12FFFA75E52FC51A57D9A30D168" box="[904,1100,1092,1115]" italics="true" pageId="34" pageNumber="34">
|
|||
|
|
<taxonomicName id="86F416BEFFA75E52FC51A57D9A3BD168" authorityName="Gaffney & Meylan & Wood & Simons & De Almeida Campos" authorityYear="2011" box="[904,1095,1092,1115]" class="Testudines" family="Podocnemididae" genus="Albertwoodemys" higherTaxonomySource="CoL" kingdom="Animalia" order="Pleurodira" pageId="34" pageNumber="34" phylum="Chordata" rank="genus">Albertwoodemys</taxonomicName>
|
|||
|
|
,
|
|||
|
|
</emphasis>
|
|||
|
|
as both are known from the early Oligocene of the Fayum.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|