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<mods:title id="988B78C9081975332C0DD7F258698B01">Biogeography and evolution of Central American cloud forest salamanders (Caudata: Plethodontidae: Cryptotriton), with the description of a new species</mods:title>
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<taxonomicName id="F8B64D66FFB7FFF3BE30B102FB68973A" ID-CoL="323D6" authority="Rovito &amp; Vásquez-Almazán &amp; Papenfuss &amp; Parra-Olea &amp; Wake, 2015" authorityName="Rovito &amp; Vásquez-Almazán &amp; Papenfuss &amp; Parra-Olea &amp; Wake" authorityYear="2015" box="[858,1205,1384,1405]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="7" pageNumber="166" phylum="Chordata" rank="species" species="xucaneborum" status="sp. nov.">
<emphasis id="0DC2EAF7FFB7FFF3BE30B102FB68973A" box="[858,1205,1384,1405]" italics="true" pageId="7" pageNumber="157">CRYPTOTRITON XUCANEBORUM</emphasis>
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, 
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<paragraph id="3F0936E5FFB7FFF3BE55B1C6FAB39783" blockId="7.[806,1422,1422,1903]" box="[831,1390,1452,1474]" pageId="7" pageNumber="157">Salamandra Escondida de la Sierra de Xucaneb</paragraph>
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<figureCitation id="A78D2A60FFB7FFF3BE55B1A1FC3197A0" box="[831,1004,1483,1505]" captionStart="Figure 6" captionStartId="11.[144,223,1443,1462]" captionTargetBox="[367,1199,195,1413]" captionTargetId="figure-180@11.[367,1199,195,1413]" captionTargetPageId="11" captionText="Figure 6. (A) Holotype of Cryptotriton xucaneborum in life. (B) Ventral view of holotype showing dark grey ventral coloration. (C) Ventral view of an individual of C. veraepacis (USAC 1920) showing lighter grey ventral coloration. (D) View of type locality of C. xucaneborum, showing small forest fragment surrounded by agricultural land. (E) Habitat where holotype of C. xucaneborum was collected." figureDoi="http://doi.org/10.5281/zenodo.10107831" httpUri="https://zenodo.org/record/10107831/files/figure.png" pageId="7" pageNumber="157">Figures 6A, 6C</figureCitation>
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<taxonomicName id="F8B64D66FFB7FFF3BE55B180FA5097BE" ID-CoL="323D3" authority="McCranie and Rovito, 2014" authorityName="McCranie and Rovito" authorityYear="2014" box="[831,1421,1514,1536]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="7" pageNumber="157" phylum="Chordata" rank="species" species="veraepacis">
<emphasis id="0DC2EAF7FFB7FFF3BE55B180FB9497BE" box="[831,1097,1514,1535]" italics="true" pageId="7" pageNumber="157">Cryptotriton veraepacis</emphasis>
<bibRefCitation id="5B274B14FFB7FFF3B938B180FA5097BE" author="McCranie JR &amp; Rovito SM" box="[1106,1421,1514,1536]" pageId="7" pageNumber="157" pagination="61 - 70" refId="ref10945" refString="McCranie JR, Rovito SM. 2014. New species of salaman- der (Caudata: Plethodontidae: Cryptotriton) from Quebrada Cataguana, Francisco Morazan, Honduras, with comments on the taxonomic status of Cryptotriton wakei. Zootaxa 3795: 61 - 70." type="journal article" year="2014">McCranie and Rovito, 2014</bibRefCitation>
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(part)
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<emphasis id="0DC2EAF7FFB7FFF3BE55B24DFB9F947D" box="[831,1090,1575,1596]" italics="true" pageId="7" pageNumber="157">Cryptotriton veraepacis</emphasis>
<bibRefCitation id="5B274B14FFB7FFF3B923B24DFA50947D" author="Vasquez-Almazan CR &amp; Rovito SM &amp; Good DA &amp; Wake DB" box="[1097,1421,1575,1597]" pageId="7" pageNumber="157" pagination="313 - 319" refId="ref11764" refString="Vasquez-Almazan CR, Rovito SM, Good DA, Wake DB. 2009. A new species of Cryptotriton (Caudata: Plethodontidae) from eastern Guatemala. Copeia 2009: 313 - 319." type="journal article" year="2009">
Vásquez-Almazán 
<emphasis id="0DC2EAF7FFB7FFF3B870B24DFA94947D" box="[1306,1353,1575,1596]" italics="true" pageId="7" pageNumber="157">et al</emphasis>
. 2009
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(part)
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<taxonomicName id="F8B64D66FFB7FFF3BE55B20EFA9A9438" ID-CoL="323D3" authority="Lynch and Wake, 1978" authorityName="Lynch and Wake" authorityYear="1978" box="[831,1351,1636,1658]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="7" pageNumber="157" phylum="Chordata" rank="species" species="veraepacis">
<emphasis id="0DC2EAF7FFB7FFF3BE55B20EFB9C943B" box="[831,1089,1636,1658]" italics="true" pageId="7" pageNumber="157">Cryptotriton veraepacis</emphasis>
<bibRefCitation id="5B274B14FFB7FFF3B922B20EFA9A9438" author="Lynch JF &amp; Wake DB" box="[1096,1351,1636,1658]" pageId="7" pageNumber="157" pagination="1 - 22" refId="ref10853" refString="Lynch JF, Wake DB. 1978. A new species of Chiropterotriton (Amphibia: Caudata) from Baja Verapaz, Guatemala, with comments on relationships among Central American members of the genus. Contributions in Science (Los Angeles) 1978: 1 - 22." type="journal article" year="1978">Lynch and Wake, 1978</bibRefCitation>
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(part)
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– 
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. Ruano-Fajardo, and 
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,
</paragraph>
<paragraph id="3F0936E5FFB8FFFCBDB7B1A9FB8C97A3" blockId="8.[221,1105,1475,1506]" box="[221,1105,1475,1506]" pageId="8" pageNumber="158">
<emphasis id="0DC2EAF7FFB8FFFCBDB7B1A9FB8C97A3" box="[221,1105,1475,1506]" italics="true" pageId="8" pageNumber="158">
<taxonomicName id="F8B64D66FFB8FFFCBDB7B1A9FDB197A3" baseAuthorityName="Papenfuss &amp; Wake" baseAuthorityYear="1987" box="[221,620,1475,1506]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="158" phylum="Chordata" rank="species" species="alvarezdeltoroi">Cryptotriton alvarezdeltoroi</taxonomicName>
<taxonomicName id="F8B64D66FFB8FFFCBFAAB1A9FB8C97A3" authorityName="Vasquez-Almazan, Rovito, Good, and Wake" authorityYear="2009" box="[704,1105,1475,1506]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="158" phylum="Chordata" rank="species" species="sierraminensis">Cryptotriton sierraminensis</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="3F0936E5FFB8FFFCBDB7B191FBC5945B" blockId="8.[221,1090,1531,1663]" box="[221,1048,1531,1562]" pageId="8" pageNumber="158">
<emphasis id="0DC2EAF7FFB8FFFCBDB7B191FBC5945B" box="[221,1048,1531,1562]" italics="true" pageId="8" pageNumber="158">
<taxonomicName id="F8B64D66FFB8FFFCBDB7B191FDCF945B" baseAuthorityName="Campbell &amp; Smith" baseAuthorityYear="1998" box="[221,530,1531,1562]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="158" phylum="Chordata" rank="species" species="monzoni">Cryptotriton monzoni</taxonomicName>
<taxonomicName id="F8B64D66FFB8FFFCBFAAB191FBC5945B" authorityName="Lynch and Wake" authorityYear="1978" box="[704,1048,1531,1562]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="158" phylum="Chordata" rank="species" species="veraepacis">Cryptotriton veraepacis</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="3F0936E5FFB8FFFCBDB7B247FB9F940D" blockId="8.[221,1090,1531,1663]" box="[221,1090,1581,1612]" pageId="8" pageNumber="158">
<emphasis id="0DC2EAF7FFB8FFFCBDB7B247FB9F940D" box="[221,1090,1581,1612]" italics="true" pageId="8" pageNumber="158">
<taxonomicName id="F8B64D66FFB8FFFCBDB7B247FE21940D" baseAuthorityName="Dunn" baseAuthorityYear="1924" box="[221,508,1581,1612]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="158" phylum="Chordata" rank="species" species="nasalis">Cryptotriton nasalis</taxonomicName>
<taxonomicName id="F8B64D66FFB8FFFCBFAAB247FB9F940D" authorityName="Rovito &amp; Vásquez-Almazán &amp; Papenfuss &amp; Parra-Olea &amp; Wake" authorityYear="2015" box="[704,1090,1581,1612]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="158" phylum="Chordata" rank="species" species="xucaneborum">Cryptotriton xucaneborum</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="3F0936E5FFB8FFFCBDB7B20AFBD6943E" blockId="8.[221,1090,1531,1663]" box="[221,1035,1632,1663]" pageId="8" pageNumber="158">
<emphasis id="0DC2EAF7FFB8FFFCBDB7B20AFBD6943E" box="[221,1035,1632,1663]" italics="true" pageId="8" pageNumber="158">
<taxonomicName id="F8B64D66FFB8FFFCBDB7B20AFDF0943E" authorityName="McCranie and Rovito" authorityYear="2014" box="[221,557,1632,1663]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="158" phylum="Chordata" rank="species" species="necopinus">Cryptotriton necopinus</taxonomicName>
<taxonomicName id="F8B64D66FFB8FFFCBFAAB20AFC7F943E" box="[704,930,1632,1663]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="158" phylum="Chordata" rank="species" species="undetermined">Cryptotriton sp.</taxonomicName>
Bethel
</emphasis>
</paragraph>
<caption id="6BC9666DFFB8FFFCBDCEB2CAFDCF94AF" pageId="8" pageNumber="158" startId="8.[164,243,1696,1715]">
<paragraph id="3F0936E5FFB8FFFCBDCEB2CAFDCF94AF" blockId="8.[164,1442,1696,1774]" pageId="8" pageNumber="158">
<emphasis id="0DC2EAF7FFB8FFFCBDCEB2CAFED194F2" bold="true" box="[164,268,1696,1715]" pageId="8" pageNumber="158">Figure 4.</emphasis>
Haplotype networks from TCS analysis of nuclear loci. Size of circle is proportionate to the frequency of each haplotype. Black dots represent inferred haplotypes not observed in sequence data, and lines connect haplotypes separated by a single mutational step.
</paragraph>
</caption>
<caption id="6BC9666DFFB9FFFDBDFAB71DFF0D9641" ID-DOI="http://doi.org/10.5281/zenodo.10107829" ID-Zenodo-Dep="10107829" httpUri="https://zenodo.org/record/10107829/files/figure.png" pageId="9" pageNumber="159" startId="9.[144,223,887,906]" targetBox="[144,1423,197,858]" targetPageId="9">
<paragraph id="3F0936E5FFB9FFFDBDFAB71DFF0D9641" blockId="9.[144,1422,887,1024]" pageId="9" pageNumber="159">
<emphasis id="0DC2EAF7FFB9FFFDBDFAB71DFF2591CA" bold="true" box="[144,248,887,907]" pageId="9" pageNumber="159">Figure 5.</emphasis>
Results of LAGRANGE analysis of geographic range evolution of 
<taxonomicName id="F8B64D66FFB9FFFDBEA1B712FB9191CA" authorityName="Garcia-Paris &amp; Wake" authorityYear="2000" box="[971,1100,888,907]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="159" phylum="Chordata" rank="genus">
<emphasis id="0DC2EAF7FFB9FFFDBEA1B712FB9191CA" box="[971,1100,888,907]" italics="true" pageId="9" pageNumber="159">Cryptotriton</emphasis>
</taxonomicName>
. Results for other genera are not shown. Pies at nodes are colored by the relative probability of each geographic range combination, and squares at tips show the areas currently occupied by each species. Each node has a pie representing the range of the ancestor prior to the speciation event and two pies representing the range of each daughter species immediately after the speciation event.
</paragraph>
</caption>
<paragraph id="3F0936E5FFB9FFFDBDFAB075FF379630" blockId="9.[144,759,1055,1903]" pageId="9" pageNumber="159">
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<collectingCountry id="47A17675FFB9FFFDBDFAB075FED09674" box="[144,269,1055,1077]" name="Guatemala" pageId="9" pageNumber="159">Guatemala</collectingCountry>
, 
<collectingRegion id="FD72F807FFB9FFFDBC7DB075FE749675" box="[279,425,1055,1077]" country="Guatemala" name="Alta Verapaz" pageId="9" pageNumber="159">Alta Verapaz</collectingRegion>
, 
<collectingMunicipality id="DF6DAC9FFFB9FFFDBCDEB075FDA69674" box="[436,635,1055,1077]" pageId="9" pageNumber="159">Municipio Tucurú</collectingMunicipality>
, 
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, 
<geoCoordinate id="5A825022FFB9FFFDBC68B057FE509613" box="[258,397,1085,1107]" degrees="15.3558" direction="north" orientation="latitude" pageId="9" pageNumber="159" precision="5" value="15.3558">15.3558 °N</geoCoordinate>
, 
<geoCoordinate id="5A825022FFB9FFFDBCF4B057FDC39612" box="[414,542,1085,1107]" degrees="90.0775" direction="west" orientation="longitude" pageId="9" pageNumber="159" precision="5" value="-90.0775">90.0775°W</geoCoordinate>
(
<specimenCode id="6F109E9EFFB9FFFDBF58B057FD519613" box="[562,652,1085,1106]" collectionCode="WGS" pageId="9" pageNumber="159">WGS84</specimenCode>
datum), 
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<elevation id="B49BD1D6FFB9FFFDBDFAB036FF3B9630" box="[144,230,1116,1138]" metricMagnitude="3" metricUnit="m" metricValue="2.045" pageId="9" pageNumber="159" unit="m" value="2045.0">2045 m</elevation>
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</materialsCitation>
.
</paragraph>
<paragraph id="3F0936E5FFB9FFFDBDC2B010FD2A96CE" blockId="9.[144,759,1055,1903]" box="[168,759,1146,1168]" pageId="9" pageNumber="159">
<emphasis id="0DC2EAF7FFB9FFFDBDC2B010FE7996CE" bold="true" box="[168,420,1146,1168]" italics="true" pageId="9" pageNumber="159">Referred specimens.</emphasis>
– 
<collectionCode id="59A7AE20FFB9FFFDBCAAB010FDDB96CE" box="[448,518,1146,1167]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="9" pageNumber="159">ANSP</collectionCode>
28198, 
<collectionCode id="59A7AE20FFB9FFFDBF34B010FD7996CE" box="[606,676,1146,1167]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="9" pageNumber="159">ANSP</collectionCode>
28199.
</paragraph>
</subSubSection>
<subSubSection id="77AC656EFFB9FFFDBDC2B0F3FE649573" pageId="9" pageNumber="159" type="diagnosis">
<paragraph id="3F0936E5FFB9FFFDBDC2B0F3FE649573" blockId="9.[144,759,1055,1903]" pageId="9" pageNumber="159">
<emphasis id="0DC2EAF7FFB9FFFDBDC2B0F3FEF096EF" bold="true" box="[168,301,1177,1198]" italics="true" pageId="9" pageNumber="159">Diagnosis.</emphasis>
– Distinguished by its very dark grey ventral coloration from 
<taxonomicName id="F8B64D66FFB9FFFDBCDBB0D2FD2A968C" authorityName="Vasquez-Almazan, Rovito, Good, and Wake" authorityYear="2009" box="[433,759,1208,1229]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="159" phylum="Chordata" rank="species" species="sierraminensis">
<emphasis id="0DC2EAF7FFB9FFFDBCDBB0D2FD2A968C" box="[433,759,1208,1229]" italics="true" pageId="9" pageNumber="159">Cryptotriton sierraminensis</emphasis>
</taxonomicName>
(yellow) and 
<taxonomicName id="F8B64D66FFB9FFFDBC49B0BCFE6496AD" authorityName="Lynch and Wake" authorityYear="1978" box="[291,441,1238,1260]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="159" phylum="Chordata" rank="species" species="veraepacis">
<emphasis id="0DC2EAF7FFB9FFFDBC49B0BCFE6496AD" box="[291,441,1238,1260]" italics="true" pageId="9" pageNumber="159">C. veraepacis</emphasis>
</taxonomicName>
(lighter grey); gular coloration darker than in 
<taxonomicName id="F8B64D66FFB9FFFDBC04B09FFE23974B" authorityName="Lynch and Wake" authorityYear="1978" box="[366,510,1269,1290]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="159" phylum="Chordata" rank="species" species="veraepacis">
<emphasis id="0DC2EAF7FFB9FFFDBC04B09FFE23974B" box="[366,510,1269,1290]" italics="true" pageId="9" pageNumber="159">C. veraepacis</emphasis>
</taxonomicName>
, with no yellow (some yellow in 
<taxonomicName id="F8B64D66FFB9FFFDBC6BB17EFE159768" authorityName="Vasquez-Almazan, Rovito, Good, and Wake" authorityYear="2009" box="[257,456,1300,1321]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="159" phylum="Chordata" rank="species" species="sierraminensis">
<emphasis id="0DC2EAF7FFB9FFFDBC6BB17EFE159768" box="[257,456,1300,1321]" italics="true" pageId="9" pageNumber="159">C. sierraminensis</emphasis>
</taxonomicName>
) (
<bibRefCitation id="5B274B14FFB9FFFDBC8AB179FF0D9709" author="Vasquez-Almazan CR &amp; Rovito SM &amp; Good DA &amp; Wake DB" pageId="9" pageNumber="159" pagination="313 - 319" refId="ref11764" refString="Vasquez-Almazan CR, Rovito SM, Good DA, Wake DB. 2009. A new species of Cryptotriton (Caudata: Plethodontidae) from eastern Guatemala. Copeia 2009: 313 - 319." type="journal article" year="2009">
Vásquez-Almazán 
<emphasis id="0DC2EAF7FFB9FFFDBFD2B17EFD359769" box="[696,744,1299,1321]" italics="true" pageId="9" pageNumber="159">et al</emphasis>
., 2009
</bibRefCitation>
). Distinguished from 
<taxonomicName id="F8B64D66FFB9FFFDBF66B158FD6C9709" authorityName="Lynch and Wake" authorityYear="1978" box="[524,689,1330,1352]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="159" phylum="Chordata" rank="species" species="veraepacis">
<emphasis id="0DC2EAF7FFB9FFFDBF66B158FD6C9709" box="[524,689,1330,1352]" italics="true" pageId="9" pageNumber="159">C. veraepacis</emphasis>
</taxonomicName>
and 
<taxonomicName id="F8B64D66FFB9FFFDBDFAB13BFE8A9727" authorityName="Vasquez-Almazan, Rovito, Good, and Wake" authorityYear="2009" box="[144,343,1361,1382]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="159" phylum="Chordata" rank="species" species="sierraminensis">
<emphasis id="0DC2EAF7FFB9FFFDBDFAB13BFE8A9727" box="[144,343,1361,1382]" italics="true" pageId="9" pageNumber="159">C. sierraminensis</emphasis>
</taxonomicName>
by lack of a tibial spur, and from 
<taxonomicName id="F8B64D66FFB9FFFDBDFAB105FEBD97C4" authorityName="Vasquez-Almazan, Rovito, Good, and Wake" authorityYear="2009" box="[144,352,1391,1413]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="159" phylum="Chordata" rank="species" species="sierraminensis">
<emphasis id="0DC2EAF7FFB9FFFDBDFAB105FEBD97C4" box="[144,352,1391,1413]" italics="true" pageId="9" pageNumber="159">C. sierraminensis</emphasis>
</taxonomicName>
by having prefrontal pierced by nasolacrimal duct, rather than evacuated along anterior margin and by the lack of a postorbital vomerine process (
<bibRefCitation id="5B274B14FFB9FFFDBC6BB1A1FD5897A0" author="Vasquez-Almazan CR &amp; Rovito SM &amp; Good DA &amp; Wake DB" box="[257,645,1483,1505]" pageId="9" pageNumber="159" pagination="313 - 319" refId="ref11764" refString="Vasquez-Almazan CR, Rovito SM, Good DA, Wake DB. 2009. A new species of Cryptotriton (Caudata: Plethodontidae) from eastern Guatemala. Copeia 2009: 313 - 319." type="journal article" year="2009">
Vásquez-Almazán 
<emphasis id="0DC2EAF7FFB9FFFDBC9BB1A6FDFA97A1" box="[497,551,1483,1505]" italics="true" pageId="9" pageNumber="159">et al</emphasis>
., 2009
</bibRefCitation>
). Distinguished from 
<taxonomicName id="F8B64D66FFB9FFFDBC59B180FE1E97BE" authorityName="McCranie and Rovito" authorityYear="2014" box="[307,451,1514,1535]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="159" phylum="Chordata" rank="species" species="necopinus">
<emphasis id="0DC2EAF7FFB9FFFDBC59B180FE1E97BE" box="[307,451,1514,1535]" italics="true" pageId="9" pageNumber="159">C. necopinus</emphasis>
</taxonomicName>
by having prefrontal processes of the premaxillary that arise fused and separate distally, rather than arising separately (
<bibRefCitation id="5B274B14FFB9FFFDBF02B24DFEC4941A" author="McCranie JR &amp; Rovito SM" pageId="9" pageNumber="159" pagination="61 - 70" refId="ref10945" refString="McCranie JR, Rovito SM. 2014. New species of salaman- der (Caudata: Plethodontidae: Cryptotriton) from Quebrada Cataguana, Francisco Morazan, Honduras, with comments on the taxonomic status of Cryptotriton wakei. Zootaxa 3795: 61 - 70." type="journal article" year="2014">McCranie &amp; Rovito, 2014</bibRefCitation>
), and by having a longer tail, more rounded nostrils, a wider foot, and fewer maxillary and vomerine teeth (
<tableCitation id="7234035EFFB9FFFDBDBCB2E8FEF894D8" box="[214,293,1666,1689]" captionStart="Table 1" captionStartId="10.[144,208,402,423]" captionText="Table 1. Means, standard deviations, and ranges for specimens measured for morphological comparisons. Total number of specimens examined is given with species name; number noted for each measurement when different from total" pageId="9" pageNumber="159">Table 1</tableCitation>
). Distinguished from 
<taxonomicName id="F8B64D66FFB9FFFDBF7DB2E9FD5E94D6" baseAuthorityName="Dunn" baseAuthorityYear="1924" box="[535,643,1666,1688]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="159" phylum="Chordata" rank="species" species="nasalis">
<emphasis id="0DC2EAF7FFB9FFFDBF7DB2E9FD5E94D6" box="[535,643,1666,1688]" italics="true" pageId="9" pageNumber="159">C. nasalis</emphasis>
</taxonomicName>
by having a prefrontal bone pierced by the nasolacrimal duct, rather than evacuated along the anterior margin (
<bibRefCitation id="5B274B14FFB9FFFDBFD9B2AAFEEF94B5" author="Lynch JF &amp; Wake DB" pageId="9" pageNumber="159" pagination="1 - 22" refId="ref10853" refString="Lynch JF, Wake DB. 1978. A new species of Chiropterotriton (Amphibia: Caudata) from Baja Verapaz, Guatemala, with comments on relationships among Central American members of the genus. Contributions in Science (Los Angeles) 1978: 1 - 22." type="journal article" year="1978">
Lynch &amp; 
<collectingCountry id="47A17675FFB9FFFDBDC7B2B4FF3294B5" box="[173,239,1758,1780]" name="Wake Island" pageId="9" pageNumber="159">Wake</collectingCountry>
, 1978
</bibRefCitation>
) and smaller nostrils (
<tableCitation id="7234035EFFB9FFFDBF56B2B4FD5294B5" box="[572,655,1758,1780]" captionStart="Table 1" captionStartId="10.[144,208,402,423]" captionText="Table 1. Means, standard deviations, and ranges for specimens measured for morphological comparisons. Total number of specimens examined is given with species name; number noted for each measurement when different from total" pageId="9" pageNumber="159">Table 1</tableCitation>
). Distinguished from 
<taxonomicName id="F8B64D66FFB9FFFDBC45B297FE259553" baseAuthorityName="Papenfuss &amp; Wake" baseAuthorityYear="1987" box="[303,504,1789,1810]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="159" phylum="Chordata" rank="species" species="alvarezdeltoroi">
<emphasis id="0DC2EAF7FFB9FFFDBC45B297FE259553" box="[303,504,1789,1810]" italics="true" pageId="9" pageNumber="159">C. alvarezdeltoroi</emphasis>
</taxonomicName>
by its larger size and smaller nostrils (
<tableCitation id="7234035EFFB9FFFDBC3CB371FE749573" box="[342,425,1819,1842]" captionStart="Table 1" captionStartId="10.[144,208,402,423]" captionText="Table 1. Means, standard deviations, and ranges for specimens measured for morphological comparisons. Total number of specimens examined is given with species name; number noted for each measurement when different from total" pageId="9" pageNumber="159">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="77AC656EFFB9FFF8BDC2B350FA7F906D" lastPageId="12" lastPageNumber="162" pageId="9" pageNumber="159" type="description">
<paragraph id="3F0936E5FFB9FFFDBDC2B350FC5697BE" blockId="9.[144,759,1055,1903]" lastBlockId="9.[806,1422,1054,1902]" pageId="9" pageNumber="159">
<emphasis id="0DC2EAF7FFB9FFFDBDC2B350FE0D9511" bold="true" box="[168,464,1850,1872]" italics="true" pageId="9" pageNumber="159">
Description of 
<typeStatus id="E00D8847FFB9FFFDBC0AB350FE169511" box="[352,459,1850,1872]" pageId="9" pageNumber="159" type="holotype">holotype</typeStatus>
.
</emphasis>
– A relatively small adult male (SVL = 
<quantity id="F84E9B00FFB9FFFDBC46B332FE48952F" box="[300,405,1880,1902]" metricMagnitude="-2" metricUnit="m" metricValue="2.57" pageId="9" pageNumber="159" unit="mm" value="25.7">25.7 mm</quantity>
). Head moderately sized, ap- proximately same width as body, snout truncate. Eyes extend slightly beyond margin of jaw. Paratoid glands large, relatively well defined. Nostrils large (maximum nostril diameter = 
<quantity id="F84E9B00FFB9FFFDBE90B010FB8C96D1" box="[1018,1105,1146,1168]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="159" unit="mm" value="0.6">0.6 mm</quantity>
). Well developed hands and feet with little webbing between digits; tips of digits rounded. Limbs relatively long; adpressed limbs separated by approximately two costal folds. Tail longer than body (
<quantity id="F84E9B00FFB9FFFDBECDB09FFBD6974B" box="[935,1035,1269,1290]" metricMagnitude="-2" metricUnit="m" metricValue="3.7399999999999998" pageId="9" pageNumber="159" unit="mm" value="37.4">37.4 mm</quantity>
), approximately rounded in crosssection, constricted at base, gradually tapering to a point. Nasolabial protuberances well developed and wide but not long, extend slightly beyond margin of lip. Large, elliptical mental hedonic gland present. Maxillary teeth (38) small and relatively numerous; premaxillary teeth (3) large, pierce upper lip. Relatively few vomerine teeth (8), arranged in line extending to internal margin of choanae.
</paragraph>
<paragraph id="3F0936E5FFB9FFFFBE55B262FD5494D9" blockId="9.[806,1422,1054,1902]" lastBlockId="11.[144,759,1605,1903]" lastPageId="11" lastPageNumber="161" pageId="9" pageNumber="159">
<emphasis id="0DC2EAF7FFB9FFFDBE55B262FC5D947D" bold="true" italics="true" pageId="9" pageNumber="159">
Measurements of the 
<typeStatus id="E00D8847FFB9FFFDB92BB26DFB7A945C" box="[1089,1191,1543,1565]" pageId="9" pageNumber="159" type="holotype">holotype</typeStatus>
(in mm) and tooth counts.
</emphasis>
– Head width 3.6; snout to gular fold (head length) 5.0; head depth at posterior angle of jaw 2.2; eyelid length 1.8; eyelid width 0.7; anterior rim of orbit to snout 0.9; horizontal orbit diameter 2.2; interorbital distance 1.3; snout to forelimb 7.7; distance separating external nares 0.8; snout projection beyond mandible 0.7; snout to posterior angle of vent (SVL) 25.7; snout to anterior angle of vent 33.7; axilla to groin 13.0; tail length 37.4; tail width at base 1.8; tail depth at base 2.3; forelimb length 5.4; hind limb length 6.5; width of right manus 1.4; width of right pes 2.3; length of longest toe 1.2; length of shortest toe 0.1; maximum nostril diameter 0.6; shoulder width 2.5. Maxillary teeth 38, premaxillary teeth 3, vomerine teeth 8.
</paragraph>
<caption id="6BC9666DFFBAFFFEBDFAB5F8FC0D9384" pageId="10" pageNumber="160" startId="10.[144,208,402,423]">
<paragraph id="3F0936E5FFBAFFFEBDFAB5F8FC0D9384" blockId="10.[144,1843,402,453]" pageId="10" pageNumber="160">
<emphasis id="0DC2EAF7FFBAFFFEBDFAB5F8FF3793E6" bold="true" box="[144,234,402,423]" pageId="10" pageNumber="160">Table 1.</emphasis>
Means, standard deviations, and ranges for specimens measured for morphological comparisons. Total number of specimens examined is given with species name; number noted for each measurement when different from total
</paragraph>
</caption>
<paragraph id="3F0936E5FFBAFFFEBC2EB583F91694C3" pageId="10" pageNumber="160">
<table id="4DB6C445FFBA000BBDFAB583F8EE94DD" box="[144,1843,489,1692]" gridcols="8" gridrows="17" pageId="10" pageNumber="160">
<tr id="818634A7FFBA000BBDFAB583F8EE9061" box="[144,1843,489,544]" gridrow="0" pageId="10" pageNumber="160" rowspan-0="1">
<th id="C2575DDBFFBA000BBC2EB583FE219061" box="[324,508,489,544]" gridcol="1" gridrow="0" pageId="10" pageNumber="160">
<taxonomicName id="F8B64D66FFBAFFFEBC2EB583FE2193BF" baseAuthorityName="Papenfuss &amp; Wake" baseAuthorityYear="1987" box="[324,508,489,510]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="10" pageNumber="160" phylum="Chordata" rank="species" species="alvarezdeltoroi">
<emphasis id="0DC2EAF7FFBAFFFEBC2EB583FE2193BF" box="[324,508,489,510]" italics="true" pageId="10" pageNumber="160">C. alvarezdeltoroi</emphasis>
</taxonomicName>
N = 3
</th>
<th id="C2575DDBFFBA000BBF21B583FD179061" box="[587,714,489,544]" gridcol="2" gridrow="0" pageId="10" pageNumber="160">
<taxonomicName id="F8B64D66FFBAFFFEBF21B583FD1D93BF" baseAuthorityName="Campbell &amp; Smith" baseAuthorityYear="1998" box="[587,704,489,510]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="10" pageNumber="160" phylum="Chordata" rank="species" species="monzoni">
<emphasis id="0DC2EAF7FFBAFFFEBF21B583FD1D93BF" box="[587,704,489,510]" italics="true" pageId="10" pageNumber="160">C. monzoni</emphasis>
</taxonomicName>
N = 3
</th>
<th id="C2575DDBFFBA000BBE72B583FC519061" box="[792,908,489,544]" gridcol="3" gridrow="0" pageId="10" pageNumber="160">
<taxonomicName id="F8B64D66FFBAFFFEBE72B583FC5D93BF" baseAuthorityName="Dunn" baseAuthorityYear="1924" box="[792,896,489,510]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="10" pageNumber="160" phylum="Chordata" rank="species" species="nasalis">
<emphasis id="0DC2EAF7FFBAFFFEBE72B583FC5D93BF" box="[792,896,489,510]" italics="true" pageId="10" pageNumber="160">C. nasalis</emphasis>
</taxonomicName>
N = 5
</th>
<th id="C2575DDBFFBA000BBEB0B583FB839061" box="[986,1118,489,544]" gridcol="4" gridrow="0" pageId="10" pageNumber="160">
<taxonomicName id="F8B64D66FFBAFFFEBEB1B583FB8393BF" authorityName="McCranie and Rovito" authorityYear="2014" box="[987,1118,489,510]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="10" pageNumber="160" phylum="Chordata" rank="species" species="necopinus">
<emphasis id="0DC2EAF7FFBAFFFEBEB1B583FB8393BF" box="[987,1118,489,510]" italics="true" pageId="10" pageNumber="160">C. necopinus</emphasis>
</taxonomicName>
N = 1
</th>
<th id="C2575DDBFFBA000BB9C7B583FAB99061" box="[1197,1380,489,544]" gridcol="5" gridrow="0" pageId="10" pageNumber="160">
<taxonomicName id="F8B64D66FFBAFFFEB9C7B583FAB993BF" authorityName="Vasquez-Almazan, Rovito, Good, and Wake" authorityYear="2009" box="[1197,1380,489,510]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="10" pageNumber="160" phylum="Chordata" rank="species" species="sierraminensis">
<emphasis id="0DC2EAF7FFBAFFFEB9C7B583FAB993BF" box="[1197,1380,489,510]" italics="true" pageId="10" pageNumber="160">C. sierraminensis</emphasis>
</taxonomicName>
N = 45
</th>
<th id="C2575DDBFFBA000BB8D8B583F9E69061" box="[1458,1595,489,544]" gridcol="6" gridrow="0" pageId="10" pageNumber="160">
<taxonomicName id="F8B64D66FFBAFFFEB8D9B583F9E693BF" authorityName="Lynch and Wake" authorityYear="1978" box="[1459,1595,489,510]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="10" pageNumber="160" phylum="Chordata" rank="species" species="veraepacis">
<emphasis id="0DC2EAF7FFBAFFFEB8D9B583F9E693BF" box="[1459,1595,489,510]" italics="true" pageId="10" pageNumber="160">C. veraepacis</emphasis>
</taxonomicName>
N = 18
</th>
<th id="C2575DDBFFBA000BBBE0B583F8EE9061" box="[1674,1843,489,544]" gridcol="7" gridrow="0" pageId="10" pageNumber="160">
<taxonomicName id="F8B64D66FFBAFFFEBBE0B583F8EE93BF" authorityName="Rovito &amp; Vásquez-Almazán &amp; Papenfuss &amp; Parra-Olea &amp; Wake" authorityYear="2015" box="[1674,1843,489,510]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="10" pageNumber="160" phylum="Chordata" rank="species" species="xucaneborum">
<emphasis id="0DC2EAF7FFBAFFFEBBE0B583F8EE93BF" box="[1674,1843,489,510]" italics="true" pageId="10" pageNumber="160">C. xucaneborum</emphasis>
</taxonomicName>
N = 5
</th>
</tr>
<tr id="818634A7FFBA000BBDFAB62FF8EE9035" box="[144,1843,581,628]" gridrow="1" pageId="10" pageNumber="160">
<th id="C2575DDBFFBA000BBDFAB62FFF289035" box="[144,245,581,628]" gridcol="0" gridrow="1" pageId="10" pageNumber="160">SVL</th>
<td id="C2575DDBFFBA000BBC2EB62FFE219035" box="[324,508,581,628]" gridcol="1" gridrow="1" pageId="10" pageNumber="160">24.4 ± 3.81 (20.0–26.6)</td>
<td id="C2575DDBFFBA000BBF21B62FFD179035" box="[587,714,581,628]" gridcol="2" gridrow="1" pageId="10" pageNumber="160">22.6 ± 0.52 (22.3–23.2)</td>
<td id="C2575DDBFFBA000BBE72B62FFC519035" box="[792,908,581,628]" gridcol="3" gridrow="1" pageId="10" pageNumber="160">27.8 ± 1.58 (25.6–29.5)</td>
<td id="C2575DDBFFBA000BBEB0B62FFB839035" box="[986,1118,581,628]" gridcol="4" gridrow="1" pageId="10" pageNumber="160">26.4</td>
<td id="C2575DDBFFBA000BB9C7B62FFAB99035" box="[1197,1380,581,628]" gridcol="5" gridrow="1" pageId="10" pageNumber="160">27.8 ± 3.15 (22.2–35.4)</td>
<td id="C2575DDBFFBA000BB8D8B62FF9E69035" box="[1458,1595,581,628]" gridcol="6" gridrow="1" pageId="10" pageNumber="160">28.1 ± 2.19 (25.2–32.2)</td>
<td id="C2575DDBFFBA000BBBE0B62FF8EE9035" box="[1674,1843,581,628]" gridcol="7" gridrow="1" pageId="10" pageNumber="160">27.8 ± 2.83 (24.9–32.0)</td>
</tr>
<tr id="818634A7FFBA000BBDFAB613F8EE9083" box="[144,1843,633,706]" gridrow="2" pageId="10" pageNumber="160">
<th id="C2575DDBFFBA000BBDFAB613FF289083" box="[144,245,633,706]" gridcol="0" gridrow="2" pageId="10" pageNumber="160">TL</th>
<td id="C2575DDBFFBA000BBC2EB613FE219083" box="[324,508,633,706]" gridcol="1" gridrow="2" pageId="10" pageNumber="160">28.2 ± 3.57 (25.1–32.1)</td>
<td id="C2575DDBFFBA000BBF21B613FD179083" box="[587,714,633,706]" gridcol="2" gridrow="2" pageId="10" pageNumber="160">24.8 ± 1.89 (23.1–26.7)</td>
<td id="C2575DDBFFBA000BBE72B613FC519083" box="[792,908,633,706]" gridcol="3" gridrow="2" pageId="10" pageNumber="160">31.9 ± 1.26 (30.6–34.0)</td>
<td id="C2575DDBFFBA000BBEB0B613FB839083" box="[986,1118,633,706]" gridcol="4" gridrow="2" pageId="10" pageNumber="160">26</td>
<td id="C2575DDBFFBA000BB9C7B613FAB99083" box="[1197,1380,633,706]" gridcol="5" gridrow="2" pageId="10" pageNumber="160">30.0 ± 7.04 (14.9–43.7) N = 39</td>
<td id="C2575DDBFFBA000BB8D8B613F9E69083" box="[1458,1595,633,706]" gridcol="6" gridrow="2" pageId="10" pageNumber="160">37.9 ± 3.94 (33.4–44.9)</td>
<td id="C2575DDBFFBA000BBBE0B613F8EE9083" box="[1674,1843,633,706]" gridcol="7" gridrow="2" pageId="10" pageNumber="160">33.7 ± 3.38 (28.8–37.4)</td>
</tr>
<tr id="818634A7FFBA000BBDFAB6ADF8EE9151" box="[144,1843,711,784]" gridrow="3" pageId="10" pageNumber="160">
<th id="C2575DDBFFBA000BBDFAB6ADFF289151" box="[144,245,711,784]" gridcol="0" gridrow="3" pageId="10" pageNumber="160">AG</th>
<td id="C2575DDBFFBA000BBC2EB6ADFE219151" box="[324,508,711,784]" gridcol="1" gridrow="3" pageId="10" pageNumber="160">13.4 + 2.51 (10.5–15.2)</td>
<td id="C2575DDBFFBA000BBF21B6ADFD179151" box="[587,714,711,784]" gridcol="2" gridrow="3" pageId="10" pageNumber="160">12.3 ± 0.71 (11.8–12.8) N = 2</td>
<td id="C2575DDBFFBA000BBE72B6ADFC519151" box="[792,908,711,784]" gridcol="3" gridrow="3" pageId="10" pageNumber="160">14.7 ± 1.14 (12.8–15.7)</td>
<td id="C2575DDBFFBA000BBEB0B6ADFB839151" box="[986,1118,711,784]" gridcol="4" gridrow="3" pageId="10" pageNumber="160">13.7</td>
<td id="C2575DDBFFBA000BB9C7B6ADFAB99151" box="[1197,1380,711,784]" gridcol="5" gridrow="3" pageId="10" pageNumber="160">14.6 ± 1.62 (11.0–19.4)</td>
<td id="C2575DDBFFBA000BB8D8B6ADF9E69151" box="[1458,1595,711,784]" gridcol="6" gridrow="3" pageId="10" pageNumber="160">14.6 ± 1.18 (13.0–16.8)</td>
<td id="C2575DDBFFBA000BBBE0B6ADF8EE9151" box="[1674,1843,711,784]" gridcol="7" gridrow="3" pageId="10" pageNumber="160">15.1 ± 2.33 (13.0–18.7)</td>
</tr>
<tr id="818634A7FFBA000BBDFAB77FF8EE9105" box="[144,1843,789,836]" gridrow="4" pageId="10" pageNumber="160">
<th id="C2575DDBFFBA000BBDFAB77FFF289105" box="[144,245,789,836]" gridcol="0" gridrow="4" pageId="10" pageNumber="160">SG</th>
<td id="C2575DDBFFBA000BBC2EB77FFE219105" box="[324,508,789,836]" gridcol="1" gridrow="4" pageId="10" pageNumber="160">4.7 ± 0.79 (3.8–5.3)</td>
<td id="C2575DDBFFBA000BBF21B77FFD179105" box="[587,714,789,836]" gridcol="2" gridrow="4" pageId="10" pageNumber="160">4.3 ± 0.40 (4.1–4.8)</td>
<td id="C2575DDBFFBA000BBE72B77FFC519105" box="[792,908,789,836]" gridcol="3" gridrow="4" pageId="10" pageNumber="160">5.6 ± 0.2 (5.4–5.8)</td>
<td id="C2575DDBFFBA000BBEB0B77FFB839105" box="[986,1118,789,836]" gridcol="4" gridrow="4" pageId="10" pageNumber="160">5.3</td>
<td id="C2575DDBFFBA000BB9C7B77FFAB99105" box="[1197,1380,789,836]" gridcol="5" gridrow="4" pageId="10" pageNumber="160">5.6 ± 0.56 (4.2–6.7)</td>
<td id="C2575DDBFFBA000BB8D8B77FF9E69105" box="[1458,1595,789,836]" gridcol="6" gridrow="4" pageId="10" pageNumber="160">5.8 ± 0.49 (4.8–6.5)</td>
<td id="C2575DDBFFBA000BBBE0B77FF8EE9105" box="[1674,1843,789,836]" gridcol="7" gridrow="4" pageId="10" pageNumber="160">5.5 ± 0.57 (5.0–6.4)</td>
</tr>
<tr id="818634A7FFBA000BBDFAB723F8EE9139" box="[144,1843,841,888]" gridrow="5" pageId="10" pageNumber="160">
<th id="C2575DDBFFBA000BBDFAB723FF289139" box="[144,245,841,888]" gridcol="0" gridrow="5" pageId="10" pageNumber="160">HW</th>
<td id="C2575DDBFFBA000BBC2EB723FE219139" box="[324,508,841,888]" gridcol="1" gridrow="5" pageId="10" pageNumber="160">3.7 ± 0.44 (3.2–4.0)</td>
<td id="C2575DDBFFBA000BBF21B723FD179139" box="[587,714,841,888]" gridcol="2" gridrow="5" pageId="10" pageNumber="160">3.3 ± 0.06 (3.2–3.3)</td>
<td id="C2575DDBFFBA000BBE72B723FC519139" box="[792,908,841,888]" gridcol="3" gridrow="5" pageId="10" pageNumber="160">4.2 ± 0.24 (3.9–4.5)</td>
<td id="C2575DDBFFBA000BBEB0B723FB839139" box="[986,1118,841,888]" gridcol="4" gridrow="5" pageId="10" pageNumber="160">3.4</td>
<td id="C2575DDBFFBA000BB9C7B723FAB99139" box="[1197,1380,841,888]" gridcol="5" gridrow="5" pageId="10" pageNumber="160">3.9 ± 0.37 (3.2–4.8)</td>
<td id="C2575DDBFFBA000BB8D8B723F9E69139" box="[1458,1595,841,888]" gridcol="6" gridrow="5" pageId="10" pageNumber="160">4.1 ± 0.4 (3.4–4.7)</td>
<td id="C2575DDBFFBA000BBBE0B723F8EE9139" box="[1674,1843,841,888]" gridcol="7" gridrow="5" pageId="10" pageNumber="160">3.8 ± 0.39 (3.5–4.4)</td>
</tr>
<tr id="818634A7FFBA000BBDFAB716F8EE91EA" box="[144,1843,892,939]" gridrow="6" pageId="10" pageNumber="160">
<th id="C2575DDBFFBA000BBDFAB716FF2891EA" box="[144,245,892,939]" gridcol="0" gridrow="6" pageId="10" pageNumber="160">FLL</th>
<td id="C2575DDBFFBA000BBC2EB716FE2191EA" box="[324,508,892,939]" gridcol="1" gridrow="6" pageId="10" pageNumber="160">4.8 ± 1.17 (3.5–5.8)</td>
<td id="C2575DDBFFBA000BBF21B716FD1791EA" box="[587,714,892,939]" gridcol="2" gridrow="6" pageId="10" pageNumber="160">4.6 ± 0.35 (4.2–4.9)</td>
<td id="C2575DDBFFBA000BBE72B716FC5191EA" box="[792,908,892,939]" gridcol="3" gridrow="6" pageId="10" pageNumber="160">5.9 ± 0.53 (5.4–6.8)</td>
<td id="C2575DDBFFBA000BBEB0B716FB8391EA" box="[986,1118,892,939]" gridcol="4" gridrow="6" pageId="10" pageNumber="160">5.3</td>
<td id="C2575DDBFFBA000BB9C7B716FAB991EA" box="[1197,1380,892,939]" gridcol="5" gridrow="6" pageId="10" pageNumber="160">5.4 ± 0.65 (3.9–6.9)</td>
<td id="C2575DDBFFBA000BB8D8B716F9E691EA" box="[1458,1595,892,939]" gridcol="6" gridrow="6" pageId="10" pageNumber="160">5.6 ± 0.69 (3.7–6.7)</td>
<td id="C2575DDBFFBA000BBBE0B716F8EE91EA" box="[1674,1843,892,939]" gridcol="7" gridrow="6" pageId="10" pageNumber="160">5.7 ± 0.74 (5.1–7.0)</td>
</tr>
<tr id="818634A7FFBA000BBDFAB7DAF8EE919E" box="[144,1843,944,991]" gridrow="7" pageId="10" pageNumber="160">
<th id="C2575DDBFFBA000BBDFAB7DAFF28919E" box="[144,245,944,991]" gridcol="0" gridrow="7" pageId="10" pageNumber="160">HLL</th>
<td id="C2575DDBFFBA000BBC2EB7DAFE21919E" box="[324,508,944,991]" gridcol="1" gridrow="7" pageId="10" pageNumber="160">5.5 ± 1.08 (4.3–6.3)</td>
<td id="C2575DDBFFBA000BBF21B7DAFD17919E" box="[587,714,944,991]" gridcol="2" gridrow="7" pageId="10" pageNumber="160">5.1 ± 0.31 (4.8−5.4)</td>
<td id="C2575DDBFFBA000BBE72B7DAFC51919E" box="[792,908,944,991]" gridcol="3" gridrow="7" pageId="10" pageNumber="160">6.8 ± 0.69 (5.9–7.5)</td>
<td id="C2575DDBFFBA000BBEB0B7DAFB83919E" box="[986,1118,944,991]" gridcol="4" gridrow="7" pageId="10" pageNumber="160">6.8</td>
<td id="C2575DDBFFBA000BB9C7B7DAFAB9919E" box="[1197,1380,944,991]" gridcol="5" gridrow="7" pageId="10" pageNumber="160">6.0 ± 0.91 (4.2–8.8)</td>
<td id="C2575DDBFFBA000BB8D8B7DAF9E6919E" box="[1458,1595,944,991]" gridcol="6" gridrow="7" pageId="10" pageNumber="160">6.6 ± 0.81 (4.9–8.3)</td>
<td id="C2575DDBFFBA000BBBE0B7DAF8EE919E" box="[1674,1843,944,991]" gridcol="7" gridrow="7" pageId="10" pageNumber="160">6.8 ± 0.7 (6.0–7.9)</td>
</tr>
<tr id="818634A7FFBA000BBDFAB78EF8EE966C" box="[144,1843,996,1069]" gridrow="8" pageId="10" pageNumber="160">
<th id="C2575DDBFFBA000BBDFAB78EFF28966C" box="[144,245,996,1069]" gridcol="0" gridrow="8" pageId="10" pageNumber="160">RFW</th>
<td id="C2575DDBFFBA000BBC2EB78EFE21966C" box="[324,508,996,1069]" gridcol="1" gridrow="8" pageId="10" pageNumber="160">1.9 ± 0.33 (1.5–2.1)</td>
<td id="C2575DDBFFBA000BBF21B78EFD17966C" box="[587,714,996,1069]" gridcol="2" gridrow="8" pageId="10" pageNumber="160">1.5 ± 0.1 (1.4–1.6)</td>
<td id="C2575DDBFFBA000BBE72B78EFC51966C" box="[792,908,996,1069]" gridcol="3" gridrow="8" pageId="10" pageNumber="160">2.4 ± 0.11 (2.3–2.6)</td>
<td id="C2575DDBFFBA000BBEB0B78EFB83966C" box="[986,1118,996,1069]" gridcol="4" gridrow="8" pageId="10" pageNumber="160">1.3</td>
<td id="C2575DDBFFBA000BB9C7B78EFAB9966C" box="[1197,1380,996,1069]" gridcol="5" gridrow="8" pageId="10" pageNumber="160">2.4 ± 0.46 (1.5–3.6) N = 44</td>
<td id="C2575DDBFFBA000BB8D8B78EF9E6966C" box="[1458,1595,996,1069]" gridcol="6" gridrow="8" pageId="10" pageNumber="160">2.8 ± 0.45 (2.0–3.4)</td>
<td id="C2575DDBFFBA000BBBE0B78EF8EE966C" box="[1674,1843,996,1069]" gridcol="7" gridrow="8" pageId="10" pageNumber="160">2.2 ± 0.56 (1.7–3.1)</td>
</tr>
<tr id="818634A7FFBA000BBDFAB058F8EE963A" box="[144,1843,1074,1147]" gridrow="9" pageId="10" pageNumber="160">
<th id="C2575DDBFFBA000BBDFAB058FF28963A" box="[144,245,1074,1147]" gridcol="0" gridrow="9" pageId="10" pageNumber="160">IN</th>
<td id="C2575DDBFFBA000BBC2EB058FE21963A" box="[324,508,1074,1147]" gridcol="1" gridrow="9" pageId="10" pageNumber="160">0.91 ± 0.03 (0.88–0.96)</td>
<td id="C2575DDBFFBA000BBF21B058FD17963A" box="[587,714,1074,1147]" gridcol="2" gridrow="9" pageId="10" pageNumber="160">0.94 ± 0.03 (0.92–0.96) N = 2</td>
<td id="C2575DDBFFBA000BBE72B058FC51963A" box="[792,908,1074,1147]" gridcol="3" gridrow="9" pageId="10" pageNumber="160">0.95 ± 0.12 (0.85–1.14)</td>
<td id="C2575DDBFFBA000BBEB0B058FB83963A" box="[986,1118,1074,1147]" gridcol="4" gridrow="9" pageId="10" pageNumber="160">0.3</td>
<td id="C2575DDBFFBA000BB9C7B058FAB9963A" box="[1197,1380,1074,1147]" gridcol="5" gridrow="9" pageId="10" pageNumber="160">0.79 ± 0.10 (0.60–1.0)</td>
<td id="C2575DDBFFBA000BB8D8B058F9E6963A" box="[1458,1595,1074,1147]" gridcol="6" gridrow="9" pageId="10" pageNumber="160">0.82 ± 0.10 (0.60–1.0)</td>
<td id="C2575DDBFFBA000BBBE0B058F8EE963A" box="[1674,1843,1074,1147]" gridcol="7" gridrow="9" pageId="10" pageNumber="160">0.69 ± 0.19 (0.40–0.89)</td>
</tr>
<tr id="818634A7FFBA000BBDFAB0EAF8EE9688" box="[144,1843,1152,1225]" gridrow="10" pageId="10" pageNumber="160">
<th id="C2575DDBFFBA000BBDFAB0EAFF289688" box="[144,245,1152,1225]" gridcol="0" gridrow="10" pageId="10" pageNumber="160">T3</th>
<td id="C2575DDBFFBA000BBC2EB0EAFE219688" box="[324,508,1152,1225]" gridcol="1" gridrow="10" pageId="10" pageNumber="160">0.66 ± 0.04 (0.62–0.70) N = 2</td>
<td id="C2575DDBFFBA000BBF21B0EAFD179688" box="[587,714,1152,1225]" gridcol="2" gridrow="10" pageId="10" pageNumber="160">0.59 ± 0.07 (0.54–0.64) N = 2</td>
<td id="C2575DDBFFBA000BBE72B0EAFC519688" box="[792,908,1152,1225]" gridcol="3" gridrow="10" pageId="10" pageNumber="160">1.04 ± 0.15 (0.94–1.3)</td>
<td id="C2575DDBFFBA000BBEB0B0EAFB839688" box="[986,1118,1152,1225]" gridcol="4" gridrow="10" pageId="10" pageNumber="160">1.9</td>
<td id="C2575DDBFFBA000BB9C7B0EAFAB99688" box="[1197,1380,1152,1225]" gridcol="5" gridrow="10" pageId="10" pageNumber="160">0.90 ± 0.12 (0.60–1.1) N = 38</td>
<td id="C2575DDBFFBA000BB8D8B0EAF9E69688" box="[1458,1595,1152,1225]" gridcol="6" gridrow="10" pageId="10" pageNumber="160">0.89 ± 0.11 (0.8–1.2)</td>
<td id="C2575DDBFFBA000BBBE0B0EAF8EE9688" box="[1674,1843,1152,1225]" gridcol="7" gridrow="10" pageId="10" pageNumber="160">1.01 ± 0.37 (0.6–1.5)</td>
</tr>
<tr id="818634A7FFBA000BBDFAB0A4F8EE9756" box="[144,1843,1230,1303]" gridrow="11" pageId="10" pageNumber="160">
<th id="C2575DDBFFBA000BBDFAB0A4FF289756" box="[144,245,1230,1303]" gridcol="0" gridrow="11" pageId="10" pageNumber="160">T5</th>
<td id="C2575DDBFFBA000BBC2EB0A4FE219756" box="[324,508,1230,1303]" gridcol="1" gridrow="11" pageId="10" pageNumber="160">0.32 ± 0.02 (0.30–0.34) N = 2</td>
<td id="C2575DDBFFBA000BBF21B0A4FD179756" box="[587,714,1230,1303]" gridcol="2" gridrow="11" pageId="10" pageNumber="160">0.17 ± 0.04 (0.14–0.20) N = 2</td>
<td id="C2575DDBFFBA000BBE72B0A4FC519756" box="[792,908,1230,1303]" gridcol="3" gridrow="11" pageId="10" pageNumber="160">0.60 ± 0.08 (0.48–0.68)</td>
<td id="C2575DDBFFBA000BBEB0B0A4FB839756" box="[986,1118,1230,1303]" gridcol="4" gridrow="11" pageId="10" pageNumber="160">1.6</td>
<td id="C2575DDBFFBA000BB9C7B0A4FAB99756" box="[1197,1380,1230,1303]" gridcol="5" gridrow="11" pageId="10" pageNumber="160">0.46 ± 0.05 (0.34–0.60) N = 34</td>
<td id="C2575DDBFFBA000BB8D8B0A4F9E69756" box="[1458,1595,1230,1303]" gridcol="6" gridrow="11" pageId="10" pageNumber="160">0.44 ± 0.10 (0.3–0.6)</td>
<td id="C2575DDBFFBA000BBBE0B0A4F8EE9756" box="[1674,1843,1230,1303]" gridcol="7" gridrow="11" pageId="10" pageNumber="160">0.54 ± 0.25 (0.2–0.9)</td>
</tr>
<tr id="818634A7FFBA000BBDFAB176F8EE9725" box="[144,1843,1308,1380]" gridrow="12" pageId="10" pageNumber="160">
<th id="C2575DDBFFBA000BBDFAB176FF289725" box="[144,245,1308,1380]" gridcol="0" gridrow="12" pageId="10" pageNumber="160">NL</th>
<td id="C2575DDBFFBA000BBC2EB176FE219725" box="[324,508,1308,1380]" gridcol="1" gridrow="12" pageId="10" pageNumber="160">0.60 ± 0.07 (0.50–0.66)</td>
<td id="C2575DDBFFBA000BBF21B176FD179725" box="[587,714,1308,1380]" gridcol="2" gridrow="12" pageId="10" pageNumber="160">0.53 ± 0.04 (0.50–0.56) N = 2</td>
<td id="C2575DDBFFBA000BBE72B176FC519725" box="[792,908,1308,1380]" gridcol="3" gridrow="12" pageId="10" pageNumber="160">0.66 ± 0.01 (0.64–0.67)</td>
<td id="C2575DDBFFBA000BBEB0B176FB839725" box="[986,1118,1308,1380]" gridcol="4" gridrow="12" pageId="10" pageNumber="160">0.8</td>
<td id="C2575DDBFFBA000BB9C7B176FAB99725" box="[1197,1380,1308,1380]" gridcol="5" gridrow="12" pageId="10" pageNumber="160">0.62 ± 0.05 (0.52–0.80)</td>
<td id="C2575DDBFFBA000BB8D8B176F9E69725" box="[1458,1595,1308,1380]" gridcol="6" gridrow="12" pageId="10" pageNumber="160">0.62 ± 0.05 (0.58–0.76)</td>
<td id="C2575DDBFFBA000BBBE0B176F8EE9725" box="[1674,1843,1308,1380]" gridcol="7" gridrow="12" pageId="10" pageNumber="160">0.50 ± 0.03 (0.48–0.54) N = 3</td>
</tr>
<tr id="818634A7FFBA000BBDFAB103F8EE97F3" box="[144,1843,1385,1458]" gridrow="13" pageId="10" pageNumber="160">
<th id="C2575DDBFFBA000BBDFAB103FF2897F3" box="[144,245,1385,1458]" gridcol="0" gridrow="13" pageId="10" pageNumber="160">NW</th>
<td id="C2575DDBFFBA000BBC2EB103FE2197F3" box="[324,508,1385,1458]" gridcol="1" gridrow="13" pageId="10" pageNumber="160">0.51 ± 0.01 (0.50–0.52)</td>
<td id="C2575DDBFFBA000BBF21B103FD1797F3" box="[587,714,1385,1458]" gridcol="2" gridrow="13" pageId="10" pageNumber="160">0.45 ± 0.04 (0.42–0.48) N = 2</td>
<td id="C2575DDBFFBA000BBE72B103FC5197F3" box="[792,908,1385,1458]" gridcol="3" gridrow="13" pageId="10" pageNumber="160">0.54 ± 0.05 (0.48–0.60)</td>
<td id="C2575DDBFFBA000BBEB0B103FB8397F3" box="[986,1118,1385,1458]" gridcol="4" gridrow="13" pageId="10" pageNumber="160">0.5</td>
<td id="C2575DDBFFBA000BB9C7B103FAB997F3" box="[1197,1380,1385,1458]" gridcol="5" gridrow="13" pageId="10" pageNumber="160">0.47 ± 0.04 (0.40–0.6)</td>
<td id="C2575DDBFFBA000BB8D8B103F9E697F3" box="[1458,1595,1385,1458]" gridcol="6" gridrow="13" pageId="10" pageNumber="160">0.50 ± 0.04 (0.44–0.56)</td>
<td id="C2575DDBFFBA000BBBE0B103F8EE97F3" box="[1674,1843,1385,1458]" gridcol="7" gridrow="13" pageId="10" pageNumber="160">0.41 ± 0.06 (0.34–0.46) N = 3</td>
</tr>
<tr id="818634A7FFBA000BBDFAB1DDF8EE9441" box="[144,1843,1463,1536]" gridrow="14" pageId="10" pageNumber="160">
<th id="C2575DDBFFBA000BBDFAB1DDFF289441" box="[144,245,1463,1536]" gridcol="0" gridrow="14" pageId="10" pageNumber="160">ND</th>
<td id="C2575DDBFFBA000BBC2EB1DDFE219441" box="[324,508,1463,1536]" gridcol="1" gridrow="14" pageId="10" pageNumber="160">1.18 ± 0.16 (1.00–1.28)</td>
<td id="C2575DDBFFBA000BBF21B1DDFD179441" box="[587,714,1463,1536]" gridcol="2" gridrow="14" pageId="10" pageNumber="160">1.18 ± 0.02 (1.17–1.19) N = 2</td>
<td id="C2575DDBFFBA000BBE72B1DDFC519441" box="[792,908,1463,1536]" gridcol="3" gridrow="14" pageId="10" pageNumber="160">1.22 ± 0.12 (1.10–1.40)</td>
<td id="C2575DDBFFBA000BBEB0B1DDFB839441" box="[986,1118,1463,1536]" gridcol="4" gridrow="14" pageId="10" pageNumber="160">1.60</td>
<td id="C2575DDBFFBA000BB9C7B1DDFAB99441" box="[1197,1380,1463,1536]" gridcol="5" gridrow="14" pageId="10" pageNumber="160">1.33 ± 0.11 (1.08–1.60)</td>
<td id="C2575DDBFFBA000BB8D8B1DDF9E69441" box="[1458,1595,1463,1536]" gridcol="6" gridrow="14" pageId="10" pageNumber="160">1.26 ± 0.11 (1.11–1.50)</td>
<td id="C2575DDBFFBA000BBBE0B1DDF8EE9441" box="[1674,1843,1463,1536]" gridcol="7" gridrow="14" pageId="10" pageNumber="160">1.24 ± 0.15 (1.14–1.41) N = 3</td>
</tr>
<tr id="818634A7FFBA000BBDFAB26FF8EE940F" box="[144,1843,1541,1614]" gridrow="15" pageId="10" pageNumber="160">
<th id="C2575DDBFFBA000BBDFAB26FFF28940F" box="[144,245,1541,1614]" gridcol="0" gridrow="15" pageId="10" pageNumber="160">PMT+MT</th>
<td id="C2575DDBFFBA000BBC2EB26FFE21940F" box="[324,508,1541,1614]" gridcol="1" gridrow="15" pageId="10" pageNumber="160">44.0 ± 15.62 (26–54)</td>
<td id="C2575DDBFFBA000BBF21B26FFD17940F" box="[587,714,1541,1614]" gridcol="2" gridrow="15" pageId="10" pageNumber="160">32.7 ± 15.04 (23–50)</td>
<td id="C2575DDBFFBA000BBE72B26FFC51940F" box="[792,908,1541,1614]" gridcol="3" gridrow="15" pageId="10" pageNumber="160">53.0 ± 2.92 (49–56)</td>
<td id="C2575DDBFFBA000BBEB0B26FFB83940F" box="[986,1118,1541,1614]" gridcol="4" gridrow="15" pageId="10" pageNumber="160">59</td>
<td id="C2575DDBFFBA000BB9C7B26FFAB9940F" box="[1197,1380,1541,1614]" gridcol="5" gridrow="15" pageId="10" pageNumber="160">50.8 ± 6.37 (37–61) N = 43</td>
<td id="C2575DDBFFBA000BB8D8B26FF9E6940F" box="[1458,1595,1541,1614]" gridcol="6" gridrow="15" pageId="10" pageNumber="160">48.9 ± 6.61 (35–59)</td>
<td id="C2575DDBFFBA000BBBE0B26FF8EE940F" box="[1674,1843,1541,1614]" gridcol="7" gridrow="15" pageId="10" pageNumber="160">45.8 ± 7.56 (37–57)</td>
</tr>
<tr id="818634A7FFBA000BBDFAB239F8EE94DD" box="[144,1843,1619,1692]" gridrow="16" pageId="10" pageNumber="160">
<th id="C2575DDBFFBA000BBDFAB239FF2894DD" box="[144,245,1619,1692]" gridcol="0" gridrow="16" pageId="10" pageNumber="160">VT</th>
<td id="C2575DDBFFBA000BBC2EB239FE2194DD" box="[324,508,1619,1692]" gridcol="1" gridrow="16" pageId="10" pageNumber="160">9.0 ± 1.00 (8–10)</td>
<td id="C2575DDBFFBA000BBF21B239FD1794DD" box="[587,714,1619,1692]" gridcol="2" gridrow="16" pageId="10" pageNumber="160">8.3 ± 0.58 (8–9)</td>
<td id="C2575DDBFFBA000BBE72B239FC5194DD" box="[792,908,1619,1692]" gridcol="3" gridrow="16" pageId="10" pageNumber="160">11.2 ± 1.79 (9–14)</td>
<td id="C2575DDBFFBA000BBEB0B239FB8394DD" box="[986,1118,1619,1692]" gridcol="4" gridrow="16" pageId="10" pageNumber="160">23</td>
<td id="C2575DDBFFBA000BB9C7B239FAB994DD" box="[1197,1380,1619,1692]" gridcol="5" gridrow="16" pageId="10" pageNumber="160">10.2 ± 1.98 (4–15) N = 44</td>
<td id="C2575DDBFFBA000BB8D8B239F9E694DD" box="[1458,1595,1619,1692]" gridcol="6" gridrow="16" pageId="10" pageNumber="160">9.1 ± 1.32 (8–12)</td>
<td id="C2575DDBFFBA000BBBE0B239F8EE94DD" box="[1674,1843,1619,1692]" gridcol="7" gridrow="16" pageId="10" pageNumber="160">9.0 ± 2.65 (6–13)</td>
</tr>
</table>
</paragraph>
<caption id="6BC9666DFFBBFFFFBDFAB1C9FD8F944F" ID-DOI="http://doi.org/10.5281/zenodo.10107831" ID-Zenodo-Dep="10107831" httpUri="https://zenodo.org/record/10107831/files/figure.png" pageId="11" pageNumber="161" startId="11.[144,223,1443,1462]" targetBox="[367,1199,195,1413]" targetPageId="11">
<paragraph id="3F0936E5FFBBFFFFBDFAB1C9FD8F944F" blockId="11.[144,1421,1443,1551]" pageId="11" pageNumber="161">
<emphasis id="0DC2EAF7FFBBFFFFBDFAB1C9FF2597F7" bold="true" box="[144,248,1443,1462]" pageId="11" pageNumber="161">Figure 6.</emphasis>
(A) Holotype of 
<taxonomicName id="F8B64D66FFBBFFFFBCC6B1CEFD1C97F7" authorityName="Rovito &amp; Vásquez-Almazán &amp; Papenfuss &amp; Parra-Olea &amp; Wake" authorityYear="2015" box="[428,705,1443,1463]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="11" pageNumber="161" phylum="Chordata" rank="species" species="xucaneborum">
<emphasis id="0DC2EAF7FFBBFFFFBCC6B1CEFD1C97F7" box="[428,705,1443,1463]" italics="true" pageId="11" pageNumber="161">Cryptotriton xucaneborum</emphasis>
</taxonomicName>
in life. (B) Ventral view of holotype showing dark grey ventral coloration. (C) Ventral view of an individual of 
<taxonomicName id="F8B64D66FFBBFFFFBF0FB1ABFC5C9795" authority="(USAC 1920)" baseAuthorityName="USAC" baseAuthorityYear="1920" box="[613,897,1473,1492]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="11" pageNumber="161" phylum="Chordata" rank="species" species="veraepacis">
<emphasis id="0DC2EAF7FFBBFFFFBF0FB1ABFD339792" box="[613,750,1473,1492]" italics="true" pageId="11" pageNumber="161">C. veraepacis</emphasis>
(USAC 1920)
</taxonomicName>
showing lighter grey ventral coloration. (D) View of type locality of 
<taxonomicName id="F8B64D66FFBBFFFFBC3FB1B4FE2397B0" authorityName="Rovito &amp; Vásquez-Almazán &amp; Papenfuss &amp; Parra-Olea &amp; Wake" authorityYear="2015" box="[341,510,1502,1521]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="11" pageNumber="161" phylum="Chordata" rank="species" species="xucaneborum">
<emphasis id="0DC2EAF7FFBBFFFFBC3FB1B4FE2397B0" box="[341,510,1502,1521]" italics="true" pageId="11" pageNumber="161">C. xucaneborum</emphasis>
</taxonomicName>
, showing small forest fragment surrounded by agricultural land. (E) Habitat where holotype of 
<taxonomicName id="F8B64D66FFBBFFFFBC67B196FE6B944F" authorityName="Rovito &amp; Vásquez-Almazán &amp; Papenfuss &amp; Parra-Olea &amp; Wake" authorityYear="2015" box="[269,438,1531,1551]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="11" pageNumber="161" phylum="Chordata" rank="species" species="xucaneborum">
<emphasis id="0DC2EAF7FFBBFFFFBC67B196FE6B944F" box="[269,438,1531,1551]" italics="true" pageId="11" pageNumber="161">C. xucaneborum</emphasis>
</taxonomicName>
was collected.
</paragraph>
</caption>
<paragraph id="3F0936E5FFBBFFFFBDC2B2CBFD2B952F" blockId="11.[144,759,1605,1903]" pageId="11" pageNumber="161">
<emphasis id="0DC2EAF7FFBBFFFFBDC2B2CBFF299494" bold="true" italics="true" pageId="11" pageNumber="161">
Coloration of the 
<typeStatus id="E00D8847FFBBFFFFBCF6B2CBFDD794F6" box="[412,522,1697,1719]" pageId="11" pageNumber="161" type="holotype">holotype</typeStatus>
in life (based on photos).
</emphasis>
– Numbers in parentheses refer to colours from 
<bibRefCitation id="5B274B14FFBBFFFFBDA1B2B4FEB894B5" author="Kohler G" box="[203,357,1758,1780]" pageId="11" pageNumber="161" refId="ref10724" refString="Kohler G. 2012. Color catalogue for field biologists. Offenbach: Herpeton." type="book" year="2012">Köhler (2012)</bibRefCitation>
. Dorsal surface of head Flesh Ocher (57) with Burnt Umber (48) in the centre of head from intercanthal region to just anterior to insertion of forelimbs. Dorsal surface of body Salmon Colour (58) with a suggestion of Dark Yellow Buff (54). Chevrons of Burnt
</paragraph>
<paragraph id="3F0936E5FFBBFFF8BE4CB22FFE7A93AE" blockId="11.[806,1422,1605,1903]" lastBlockId="12.[164,779,198,1902]" lastPageId="12" lastPageNumber="162" pageId="11" pageNumber="161">Umber (48) along dorsal midline from insertion of forelimbs to base of tail. Dorsal surface of tail uniform Dark Salmon Colour (58), becoming more reddish along final third of tail. Dorsal surface of limbs Light Pratt’s Rufous (71) and Salmon Colour (58) with Dark Drab (45) speckles and white flecks. Dorsal surface of feet Light Neutral Grey (297) with white flecks; toe tips pinkish. Lateral surface of head and body with same background colours as above, blending ventrally into Dark Greyish Brown (284) with numerous white flecks. Gular region mottled Light Neutral Grey (297) and Sepia (286), becoming nearly uniform Sepia just anterior to gular fold. Ventral surface of body and anterior third of tail Sepia. White flecks on gular region and ventral surface of body and tail, becoming less numerous toward ventral midline. Posterior two-thirds of tail Vinaceous (247), with some Sepia; white flecks throughout tail. Ventral surfaces of limbs Pale Neutral Grey (296) with Tawny Olive (17), small white flecks. Underside of feet Pale Neutral Grey, toe tips pinkish.</paragraph>
<paragraph id="3F0936E5FFBCFFF8BDD6B59DFE6D91DA" blockId="12.[164,779,198,1902]" pageId="12" pageNumber="162">
<emphasis id="0DC2EAF7FFBCFFF8BDD6B59DFD8C904C" bold="true" box="[188,593,503,525]" italics="true" pageId="12" pageNumber="162">
Colour of the 
<typeStatus id="E00D8847FFBCFFF8BC08B59DFE15904C" box="[354,456,503,525]" pageId="12" pageNumber="162" type="holotype">holotype</typeStatus>
in alcohol.
</emphasis>
– Dorsal surface of head Raw Umber (22), with Hair Brown (277) in the centre of head from intercanthal region to just anterior to insertion of forelimbs. Dorsal surface of body Raw Umber with Hair Brown chevrons along dorsal midline. Dorsal surface of tail Buff (5). Dorsal surface of limbs and feet mottled Buff and Raw Umber (23). Gular region mottled Drab-Grey (256) and Hair Brown, becoming nearly all brown towards gular fold. Ventral surface of body Hair Brown. Anterior third of tail Hair Brown, mottled with Buff towards tip. Underside of limbs Buff mottled with Hair Brown, underside of feet Smoke Grey (266). Pale flecks throughout ventral surface of head, body, and tail.
</paragraph>
<paragraph id="3F0936E5FFBCFFF8BDD6B7CEFD979768" blockId="12.[164,779,198,1902]" pageId="12" pageNumber="162">
<emphasis id="0DC2EAF7FFBCFFF8BDD6B7CEFE9A91F8" bold="true" box="[188,327,932,953]" italics="true" pageId="12" pageNumber="162">Variation –</emphasis>
The male 
<typeStatus id="E00D8847FFBCFFF8BCAAB7CFFDF991FB" box="[448,548,933,954]" pageId="12" pageNumber="162" type="paratype">paratype</typeStatus>
is roughly the same size as the 
<typeStatus id="E00D8847FFBCFFF8BC4EB7A9FE5E9198" box="[292,387,963,985]" pageId="12" pageNumber="162" type="holotype">holotype</typeStatus>
(
<quantity id="F84E9B00FFBCFFF8BCF8B7A9FE2A9199" box="[402,503,963,984]" metricMagnitude="-2" metricUnit="m" metricValue="2.4899999999999998" pageId="12" pageNumber="162" unit="mm" value="24.9">24.9 mm</quantity>
), while the 
<specimenCount id="29B0FD6CFFBCFFF8BF16B7A9FCD69198" box="[636,779,963,985]" pageId="12" pageNumber="162" type="female">three female</specimenCount>
<typeStatus id="E00D8847FFBCFFF8BDCEB788FEC891B6" box="[164,277,994,1015]" pageId="12" pageNumber="162" type="paratype">paratypes</typeStatus>
are slightly larger, reaching a maximum of 
<quantity id="F84E9B00FFBCFFF8BDCEB06AFF289654" box="[164,245,1024,1045]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="12" pageNumber="162" unit="mm" value="32.0">32 mm</quantity>
. There is some variation in maxillary teeth in the type series, with the largest female having only 37 maxillary-premaxillary teeth, while another female has a total of 57 maxillary-premaxillary teeth. Similar variation is seen in the vomerine tooth counts; the largest female has only six vomerine teeth, while the smallest male has the most vomerine teeth (13). The female 
<typeStatus id="E00D8847FFBCFFF8BD93B0BCFEB796AA" box="[249,362,1238,1259]" pageId="12" pageNumber="162" type="paratype">paratypes</typeStatus>
have nasolabial protuberances that are only very slightly developed and visible only as a slight swelling in the labial margin.
</paragraph>
<paragraph id="3F0936E5FFBCFFF8BDD6B15BFE5994D9" blockId="12.[164,779,198,1902]" pageId="12" pageNumber="162">
<emphasis id="0DC2EAF7FFBCFFF8BDD6B15BFEE69706" bold="true" box="[188,315,1329,1351]" italics="true" pageId="12" pageNumber="162">Osteology.</emphasis>
– We examined X-rays of the 
<typeStatus id="E00D8847FFBCFFF8BFC0B158FCD69709" box="[682,779,1330,1352]" pageId="12" pageNumber="162" type="holotype">holotype</typeStatus>
and a 
<typeStatus id="E00D8847FFBCFFF8BD80B13BFE909727" box="[234,333,1361,1382]" pageId="12" pageNumber="162" type="paratype">paratype</typeStatus>
(MVZ 269516). Prefrontal bones appear to be pierced by a nasolacrimal duct, as in 
<taxonomicName id="F8B64D66FFBCFFF8BF16B105FCD697C4" baseAuthorityName="USAC" baseAuthorityYear="1920" box="[636,779,1391,1413]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="12" pageNumber="162" phylum="Chordata" rank="species" species="veraepacis">
<emphasis id="0DC2EAF7FFBCFFF8BF16B105FCD697C4" box="[636,779,1391,1413]" italics="true" pageId="12" pageNumber="162">C. veraepacis</emphasis>
</taxonomicName>
and the single known specimen of 
<taxonomicName id="F8B64D66FFBCFFF8BF40B1E4FD6B97E2" authorityName="McCranie and Rovito" authorityYear="2014" box="[554,694,1422,1443]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="12" pageNumber="162" phylum="Chordata" rank="species" species="necopinus">
<emphasis id="0DC2EAF7FFBCFFF8BF40B1E4FD6B97E2" box="[554,694,1422,1443]" italics="true" pageId="12" pageNumber="162">C. necopinus</emphasis>
</taxonomicName>
(
<bibRefCitation id="5B274B14FFBCFFF8BFACB1E4FE999780" author="Lynch JF &amp; Wake DB" pageId="12" pageNumber="162" pagination="1 - 22" refId="ref10853" refString="Lynch JF, Wake DB. 1978. A new species of Chiropterotriton (Amphibia: Caudata) from Baja Verapaz, Guatemala, with comments on relationships among Central American members of the genus. Contributions in Science (Los Angeles) 1978: 1 - 22." type="journal article" year="1978">
Lynch &amp; 
<collectingCountry id="47A17675FFBCFFF8BDD5B1C6FEDD9783" box="[191,256,1452,1474]" name="Wake Island" pageId="12" pageNumber="162">Wake</collectingCountry>
, 1978
</bibRefCitation>
; 
<bibRefCitation id="5B274B14FFBCFFF8BC3AB1C6FDAD9783" author="McCranie JR &amp; Rovito SM" box="[336,624,1452,1474]" pageId="12" pageNumber="162" pagination="61 - 70" refId="ref10945" refString="McCranie JR, Rovito SM. 2014. New species of salaman- der (Caudata: Plethodontidae: Cryptotriton) from Quebrada Cataguana, Francisco Morazan, Honduras, with comments on the taxonomic status of Cryptotriton wakei. Zootaxa 3795: 61 - 70." type="journal article" year="2014">McCranie &amp; Rovito, 2014</bibRefCitation>
). Frontal processes of premaxillary bone arise fused and separate distally. Vomer lacks a postorbital process. Tibial spur absent. As in other species of 
<taxonomicName id="F8B64D66FFBCFFF8BF02B262FCDF945C" authorityName="Garcia-Paris &amp; Wake" authorityYear="2000" box="[616,770,1544,1565]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="12" pageNumber="162" phylum="Chordata" rank="genus">
<emphasis id="0DC2EAF7FFBCFFF8BF02B262FCDF945C" box="[616,770,1544,1565]" italics="true" pageId="12" pageNumber="162">Cryptotriton</emphasis>
</taxonomicName>
, septomaxillary absent and mesopodials unmineralized. Phalangeal formula for manus is 1-2-3-2 and 1-2-3- 3-2 for pes, identical to that of other species in the genus 
<taxonomicName id="F8B64D66FFBCFFF8BD9BB2E9FEA094D9" authorityName="Garcia-Paris &amp; Wake" authorityYear="2000" box="[241,381,1667,1688]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="12" pageNumber="162" phylum="Chordata" rank="genus">
<emphasis id="0DC2EAF7FFBCFFF8BD9BB2E9FEA094D9" box="[241,381,1667,1688]" italics="true" pageId="12" pageNumber="162">Cryptotriton</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="3F0936E5FFBCFFF8BDD6B2CAFA7F906D" blockId="12.[164,779,198,1902]" lastBlockId="12.[826,1442,197,556]" pageId="12" pageNumber="162">
<emphasis id="0DC2EAF7FFBCFFF8BDD6B2CAFE2694F7" bold="true" box="[188,507,1696,1718]" italics="true" pageId="12" pageNumber="162">Habitat and distribution.</emphasis>
– Known only from two localities separated by 
<quantity id="F84E9B00FFBCFFF8BCC8B2AAFE3A9494" box="[418,487,1727,1749]" metricMagnitude="4" metricUnit="m" metricValue="2.2" pageId="12" pageNumber="162" unit="km" value="22.0">22 km</quantity>
in the Sierra de Xucaneb, 
<collectingRegion id="FD72F807FFBCFFF8BDCEB2B4FEE294B5" box="[164,319,1758,1780]" country="Guatemala" name="Alta Verapaz" pageId="12" pageNumber="162">Alta Verapaz</collectingRegion>
, 
<collectingCountry id="47A17675FFBCFFF8BC24B2B4FE0D94B5" box="[334,464,1758,1780]" name="Guatemala" pageId="12" pageNumber="162">Guatemala</collectingCountry>
. The species may occur at a few other points of relatively high elevation in the Sierra de Xucaneb, but extensive deforestation in most areas has eliminated what may have once been suitable habitat for the species. The 
<typeStatus id="E00D8847FFBCFFF8BF43B332FD57952F" box="[553,650,1880,1902]" pageId="12" pageNumber="162" type="holotype">holotype</typeStatus>
was found in an arboreal bromeliad, while the 
<specimenCount id="29B0FD6CFFBCFFF8B9ACB4AFFABA929A" box="[1222,1383,197,219]" pageId="12" pageNumber="162" type="generic" typeStatus="paratypes">four paratypes</specimenCount>
from Chelemhá were found active at night on vegetation 
<quantity id="F84E9B00FFBCFFF8BE50B569FC5D9359" box="[826,896,259,280]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="12" pageNumber="162" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">1–2 m</quantity>
above the ground. The forest at Finca Volcán represents a transition from lower elevation forest to cloud forest, with relatively few bromeliads and other epiphyte growth, and is classified as subtropical rainforest (
<bibRefCitation id="5B274B14FFBCFFF8BEE4B517FB9693D2" author="Holdridge LR" box="[910,1099,381,403]" pageId="12" pageNumber="162" refId="ref10665" refString="Holdridge LR. 1967. Life zone ecology. San Jose, Costa Rica: Tropical Science Center." type="book" year="1967">Holdridge, 1967</bibRefCitation>
), while habitat at Chelemhá is lower montane rain forest (
<bibRefCitation id="5B274B14FFBCFFF8B9F5B5F6FA8793F3" author="Holdridge LR" box="[1183,1370,412,434]" pageId="12" pageNumber="162" refId="ref10665" refString="Holdridge LR. 1967. Life zone ecology. San Jose, Costa Rica: Tropical Science Center." type="book" year="1967">Holdridge, 1967</bibRefCitation>
) with typical cloud forest vegetation and extensive epiphyte cover. The only syntopic species of salamander known to occur with 
<taxonomicName id="F8B64D66FFBCFFF8B92FB592FA5C904D" authorityName="Rovito &amp; Vásquez-Almazán &amp; Papenfuss &amp; Parra-Olea &amp; Wake" authorityYear="2015" box="[1093,1409,503,525]" class="Amphibia" family="Plethodontidae" genus="Cryptotriton" kingdom="Animalia" order="Caudata" pageId="12" pageNumber="162" phylum="Chordata" rank="species" species="xucaneborum">
<emphasis id="0DC2EAF7FFBCFFF8B92FB592FA5C904D" box="[1093,1409,503,525]" italics="true" pageId="12" pageNumber="162">Cryptotriton xucaneborum</emphasis>
</taxonomicName>
is 
<taxonomicName id="F8B64D66FFBCFFF8BE50B67CFBFA906A" baseAuthorityName="Schmidt" baseAuthorityYear="1936" box="[826,1063,534,555]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="12" pageNumber="162" phylum="Chordata" rank="species" species="helmrichi">
<emphasis id="0DC2EAF7FFBCFFF8BE50B67CFBFA906A" box="[826,1063,534,555]" italics="true" pageId="12" pageNumber="162">Bolitoglossa helmrichi</emphasis>
</taxonomicName>
, which occurs at both known sites.
</paragraph>
</subSubSection>
</treatment>
</document>