<documentID-DOI="http://dx.doi.org/10.3897/zookeys.1150.93760"ID-GBIF-Dataset="ef137bea-e9d6-406b-b016-95acd7758286"ID-PMC="PMC10208737"ID-Pensoft-Pub="1313-2970-1150-1"ID-Pensoft-UUID="2D0F861C78665B9BABB241437CA5ED53"ID-PubMed="37234100"ID-ZooBank="A38C5FB69F664F858788AAA53D21704D"ModsDocID="1313-2970-1150-1"checkinTime="1677664009133"checkinUser="pensoft"docAuthor="Crews, Sarah C."docDate="2023"docId="EEF96E2FB4855B4797AFEA970D4D32B4"docLanguage="en"docName="ZooKeys 1150: 1-189"docOrigin="ZooKeys 1150"docPubDate="2023-02-27"docSource="http://dx.doi.org/10.3897/zookeys.1150.93760"docTitle="Karaops larapinta Crews 2023, sp. nov."docType="treatment"docUuid="3C4B9AAD-46DD-444C-9F69-1F5672622022"docUuidSource="ZooBank"docVersion="4"id="2D0F861C78665B9BABB241437CA5ED53"lastPageNumber="1"masterDocId="2D0F861C78665B9BABB241437CA5ED53"masterDocTitle="But wait, there's more! Descriptions of new species and undescribed sexes of flattie spiders (Araneae, Selenopidae, Karaops) from Australia"masterLastPageNumber="189"masterPageNumber="1"pageNumber="1"updateTime="1732488240156"updateUser="ExternalLinkService">
<mods:titleid="B2E9B18806A0B99A61C8EE02E8BA0ED1">But wait, there's more! Descriptions of new species and undescribed sexes of flattie spiders (Araneae, Selenopidae, Karaops) from Australia</mods:title>
<mods:affiliationid="23BE610266CBC280A1A6A94E10238889">California Academy of Sciences, Department of Entomology, 55 Music Concourse Drive, San Francisco, CA, 94118, USA</mods:affiliation>
<figureCitationid="23D413D24A51E58629369EDCAF389F45"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Species and habitats of the Central Desert species group A Karaops kwartatuma sp. nov., holotype male, Ormiston Gorge, West MacDonnell Range National Park, Northern Territory (sel _ 1093, MAGNT A 004858) B Karaops deserticola, female holotype, Mount Lindsay, South Australia (SAM N 199350) C Karaops larapinta sp. nov., holotype female, Finke River, Northern Territory (QMS 110852) D Karaops larapinta sp. nov., paratype female, Finke River, Northern Territory (QMS 110852) E Habitat of Karaops sp. from Watarrka National Park, Northern Territory F Karaops sp., King's Canyon Rim Walk, Watarrka National Park, Northern Territory G habitat of Karaops kwartatuma sp. nov., Ormiston Gorge, Northern Territory, Australia. Scale bars: 1 mm."figureDoi="10.3897/zookeys.1150.93760.figure2"httpUri="https://binary.pensoft.net/fig/816382"pageId="0"pageNumber="1">Figs 2C, D</figureCitation>
<figureCitationid="AB2AD5CD09362D09EF8E7FD2759D55A2"captionStart="Figure 3"captionStartId="F5"captionText="Figure 3. Epigynes and endogynes of members of the Central Desert species group A Karaops larapinta sp. nov., holotype female, epigyne, Finke River, Northern Territory (QMS 110852) B same, endogyne C Karaops mparntwe sp. nov., holotype female, epigyne, Alice Springs, Northern Territory (AMT AA 10.852), pink = copulatory openings, blue = median field, yellow = lateral lobes / epigynal plate D same, endogyne, yellow = accessory bulbs, blue = spermathecae, red = fertilization ducts, green = posterodorsal fold E same, epigyne F same, endogyne. Scale bars: 0.5 mm."figureDoi="10.3897/zookeys.1150.93760.figure3"httpUri="https://binary.pensoft.net/fig/816385"pageId="0"pageNumber="1">, 3A, B</figureCitation>
<figureCitationid="2022032651DD3F6B7599B8A62B6E8CFD"captionStart="Figure 5"captionStartId="F7"captionText="Figure 5. Species from the Central Desert species group A Karaops kwartatuma sp. nov., holotype male, Ormiston Gorge, West MacDonnell Range National Park, Northern Territory (MAGNT A 004858) B same, holotype male, face showing black marks near eyes and " mustache " C Karaops larapinta sp. nov., holotype, epigyne, caudal, Finke River, Northern Territory (QMS 110852) D Karaops sp. from Watarrka National Park, Northern Territory. Scale bars: 0.5 mm."figureDoi="10.3897/zookeys.1150.93760.figure5"httpUri="https://binary.pensoft.net/fig/816387"pageId="0"pageNumber="1">, 5C</figureCitation>
<figureCitationid="6F20940BEFE2DB3C4DC4B25812E35121"captionStart="Map 1"captionStartId="F3"captionText="Map 1. Map of Australia showing areas that are expanded in subsequent figures. CD = Central Desert group, D = Karaops dawara group, F = Karaops francesae group, K = Kimberley group, UA = species that are not assigned to a group, PG = Pilbara-Gascoyne group."figureDoi="10.3897/zookeys.1150.93760.map1"httpUri="https://binary.pensoft.net/fig/816383"pageId="0"pageNumber="1">, Maps 1</figureCitation>
<figureCitationid="256B13CCBBB2BB98293EF8520DC3F7B6"captionStart="Map 2"captionStartId="F4"captionText="Map 2. Species of the Central Desert species group; orange = juvenile from Morgan Range, light purple = Karaops deserticola, teal = Karaops ngarutjaranya, red = juveniles from Watarrka, blue = Karaops larapinta sp. nov., pink = Karaops kwartatuma sp. nov., yellow = Karaops pilkingtoni, x = Karaops mparntwe sp. nov., purple = Karaops vadlaadambara, green = Karaops manaayn. Juveniles are treated as distinct species based on molecular data."figureDoi="10.3897/zookeys.1150.93760.map2"httpUri="https://binary.pensoft.net/fig/816384"pageId="0"pageNumber="1">, 2</figureCitation>
<figureCitationid="074E6134782418BB5CAC7A83BF27C6EC"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Species and habitats of the Central Desert species group A Karaops kwartatuma sp. nov., holotype male, Ormiston Gorge, West MacDonnell Range National Park, Northern Territory (sel _ 1093, MAGNT A 004858) B Karaops deserticola, female holotype, Mount Lindsay, South Australia (SAM N 199350) C Karaops larapinta sp. nov., holotype female, Finke River, Northern Territory (QMS 110852) D Karaops larapinta sp. nov., paratype female, Finke River, Northern Territory (QMS 110852) E Habitat of Karaops sp. from Watarrka National Park, Northern Territory F Karaops sp., King's Canyon Rim Walk, Watarrka National Park, Northern Territory G habitat of Karaops kwartatuma sp. nov., Ormiston Gorge, Northern Territory, Australia. Scale bars: 1 mm."figureDoi="10.3897/zookeys.1150.93760.figure2"httpUri="https://binary.pensoft.net/fig/816382"pageId="0"pageNumber="1">2C</figureCitation>
) is similar to other species of the Central Desert group by the median lobe that is unsclerotized posteriorly, the small copulatory ducts, and the large accessory bulbs (Fig.
<figureCitationid="7E2C1AF173D7F2DFFB389B54462D70C0"captionStart="Figure 3"captionStartId="F5"captionText="Figure 3. Epigynes and endogynes of members of the Central Desert species group A Karaops larapinta sp. nov., holotype female, epigyne, Finke River, Northern Territory (QMS 110852) B same, endogyne C Karaops mparntwe sp. nov., holotype female, epigyne, Alice Springs, Northern Territory (AMT AA 10.852), pink = copulatory openings, blue = median field, yellow = lateral lobes / epigynal plate D same, endogyne, yellow = accessory bulbs, blue = spermathecae, red = fertilization ducts, green = posterodorsal fold E same, epigyne F same, endogyne. Scale bars: 0.5 mm."figureDoi="10.3897/zookeys.1150.93760.figure3"httpUri="https://binary.pensoft.net/fig/816385"pageId="0"pageNumber="1">3A, B</figureCitation>
<bibRefCitationid="523C3651390A1D55AD9912487BCF916E"DOI="https://doi.org/10.3897/zookeys.99.723"author="Crews, SC"journalOrPublisher="ZooKeys"pageId="0"pageNumber="1"pagination="1 - 103"refId="B14"refString="Crews, SC, Harvey, MS, 2011. The spider family Selenopidae (Arachnida, Araneae) in Australasia and the Oriental Region. ZooKeys 99: 1 - 103, DOI: https://doi.org/10.3897/zookeys.99.723"title="The spider family Selenopidae (Arachnida, Araneae) in Australasia and the Oriental Region."url="https://doi.org/10.3897/zookeys.99.723"volume="99"year="2011">Crews and Harvey 2011</bibRefCitation>
<figureCitationid="7DB5B04B4264611B0AE46A5F200E1542"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Species and habitats of the Central Desert species group A Karaops kwartatuma sp. nov., holotype male, Ormiston Gorge, West MacDonnell Range National Park, Northern Territory (sel _ 1093, MAGNT A 004858) B Karaops deserticola, female holotype, Mount Lindsay, South Australia (SAM N 199350) C Karaops larapinta sp. nov., holotype female, Finke River, Northern Territory (QMS 110852) D Karaops larapinta sp. nov., paratype female, Finke River, Northern Territory (QMS 110852) E Habitat of Karaops sp. from Watarrka National Park, Northern Territory F Karaops sp., King's Canyon Rim Walk, Watarrka National Park, Northern Territory G habitat of Karaops kwartatuma sp. nov., Ormiston Gorge, Northern Territory, Australia. Scale bars: 1 mm."figureDoi="10.3897/zookeys.1150.93760.figure2"httpUri="https://binary.pensoft.net/fig/816382"pageId="0"pageNumber="1">2A, B</figureCitation>
<bibRefCitationid="4B89B2B3E0FF91E19BD97319647854A7"DOI="https://doi.org/10.3897/zookeys.99.723"author="Crews, SC"journalOrPublisher="ZooKeys"pageId="0"pageNumber="1"pagination="1 - 103"refId="B14"refString="Crews, SC, Harvey, MS, 2011. The spider family Selenopidae (Arachnida, Araneae) in Australasia and the Oriental Region. ZooKeys 99: 1 - 103, DOI: https://doi.org/10.3897/zookeys.99.723"title="The spider family Selenopidae (Arachnida, Araneae) in Australasia and the Oriental Region."url="https://doi.org/10.3897/zookeys.99.723"volume="99"year="2011">Crews and Harvey 2011</bibRefCitation>
<figureCitationid="39E540A3F525A106AC0031EBFDEC5C80"captionStart="Figure 3"captionStartId="F5"captionText="Figure 3. Epigynes and endogynes of members of the Central Desert species group A Karaops larapinta sp. nov., holotype female, epigyne, Finke River, Northern Territory (QMS 110852) B same, endogyne C Karaops mparntwe sp. nov., holotype female, epigyne, Alice Springs, Northern Territory (AMT AA 10.852), pink = copulatory openings, blue = median field, yellow = lateral lobes / epigynal plate D same, endogyne, yellow = accessory bulbs, blue = spermathecae, red = fertilization ducts, green = posterodorsal fold E same, epigyne F same, endogyne. Scale bars: 0.5 mm."figureDoi="10.3897/zookeys.1150.93760.figure3"httpUri="https://binary.pensoft.net/fig/816385"pageId="0"pageNumber="1">3A, B</figureCitation>
sp. nov., holotype female, epigyne, Alice Springs, Northern Territory (AMT AA 10.852), pink = copulatory openings, blue = median field, yellow = lateral lobes/epigynal plate
(holotype). Total length 7.49. Carapace: length 2.45, width 3.69. Chelicerae: promargin with three teeth, retromargin with two teeth. Eyes: AER slightly recurved, PER recurved; diameters AME 0.18, ALE 0.18, PME 0.29, PLE 0.36; interdistances AME-PME 0.04, PME-ALE 0.10, ALE-PLE 0.20, PME-PME 1.20, ALE-ALE 1.62, AME-AME 0.59, PLE-PLE 1.85. Sternum: length 1.67, width 1.89. Abdomen length 5.04, width 4.03. Color: Carapace: orange-yellow, with two dark markings lateroposterior to eye region, three marks each on lateral margins, faded, most setae worn off. Chelicerae: yellowish, paturon with a longitudinal curved mark frontally, setae white, long anteriorly. Maxillae: yellowish white. Labium: gray, pale distally. Sternum: yellowish. Abdomen: dorsally reddish orange, two dark spots at anterior margin, two laterally just posterior to these, two lateromedially posterior to these, two small ones just posterior to previous, two dark marks extend to lateral edges, connect to jagged markings lateromedially, two dark, wavy lines posterolaterally; ventrally grayish. Legs: pale yellow-orange, Cx with dusky mark prolaterally, Fm with dusky mark prolaterally, another medially, forming annulation, Pt with dark mark ventrally, Ti with dark mark ventrally at Pt-Ti joint, with annulation closer to Mt, dark annulation on Ti at Ti-Mt joint and halfway between that and Mt joint, dark annulation on Mt at Ti joint and at Mt-Ta joint, Ta tip not dark (may be faded); spines dark basally, pale distally; spination leg I Fm d 1-1-1, pr 1-1-0, Ti v 2-2-2-2-2-2, Mt v 2-2-2-2; leg II Fm d 1-1-1, Ti v 2-2-2-2-2-2, Mt v 2-2-2-2; leg III Fm d 1-1-1; leg IV Fm d 1-1-1, pr 0-0-1, rl 0-0-1; leg formula 4321; measurements leg I 13.86 (3.85, 1.65, 3.83, 3.24, 1.3); leg II 15.73 (4.95, 1.65, 4.13, 3.60, 1.40); leg III 16.13 (5.23, 1.40, 4.28, 3.70, 1.53); leg IV 16.36 (5.50, 1.48, 4.26, 3.71, 1.40). Palp: spination Fm d 0-1-3; 3.30 (1.05, 0.60, 0.75, 0.90); basally dusky, Ta dusky basodorsally; claw with eight teeth. Epigyne: EP triangular; MF with long, triangular depression, lobe posteriorly unsclerotized (Figs
<figureCitationid="7FF6264E7FA2F6002A628090AC664DC6"captionStart="Figure 3"captionStartId="F5"captionText="Figure 3. Epigynes and endogynes of members of the Central Desert species group A Karaops larapinta sp. nov., holotype female, epigyne, Finke River, Northern Territory (QMS 110852) B same, endogyne C Karaops mparntwe sp. nov., holotype female, epigyne, Alice Springs, Northern Territory (AMT AA 10.852), pink = copulatory openings, blue = median field, yellow = lateral lobes / epigynal plate D same, endogyne, yellow = accessory bulbs, blue = spermathecae, red = fertilization ducts, green = posterodorsal fold E same, epigyne F same, endogyne. Scale bars: 0.5 mm."figureDoi="10.3897/zookeys.1150.93760.figure3"httpUri="https://binary.pensoft.net/fig/816385"pageId="0"pageNumber="1">3A, B</figureCitation>
<figureCitationid="11D3B6B70585EC7FCA07D3C331F5F69B"captionStart="Figure 5"captionStartId="F7"captionText="Figure 5. Species from the Central Desert species group A Karaops kwartatuma sp. nov., holotype male, Ormiston Gorge, West MacDonnell Range National Park, Northern Territory (MAGNT A 004858) B same, holotype male, face showing black marks near eyes and " mustache " C Karaops larapinta sp. nov., holotype, epigyne, caudal, Finke River, Northern Territory (QMS 110852) D Karaops sp. from Watarrka National Park, Northern Territory. Scale bars: 0.5 mm."figureDoi="10.3897/zookeys.1150.93760.figure5"httpUri="https://binary.pensoft.net/fig/816387"pageId="0"pageNumber="1">5C</figureCitation>
<figureCitationid="76BF73CAECC316396E19CD93690F3335"captionStart="Figure 3"captionStartId="F5"captionText="Figure 3. Epigynes and endogynes of members of the Central Desert species group A Karaops larapinta sp. nov., holotype female, epigyne, Finke River, Northern Territory (QMS 110852) B same, endogyne C Karaops mparntwe sp. nov., holotype female, epigyne, Alice Springs, Northern Territory (AMT AA 10.852), pink = copulatory openings, blue = median field, yellow = lateral lobes / epigynal plate D same, endogyne, yellow = accessory bulbs, blue = spermathecae, red = fertilization ducts, green = posterodorsal fold E same, epigyne F same, endogyne. Scale bars: 0.5 mm."figureDoi="10.3897/zookeys.1150.93760.figure3"httpUri="https://binary.pensoft.net/fig/816385"pageId="0"pageNumber="1">3A</figureCitation>
<figureCitationid="9E6010D03748C236E83832AC97B4748C"captionStart="Figure 5"captionStartId="F7"captionText="Figure 5. Species from the Central Desert species group A Karaops kwartatuma sp. nov., holotype male, Ormiston Gorge, West MacDonnell Range National Park, Northern Territory (MAGNT A 004858) B same, holotype male, face showing black marks near eyes and " mustache " C Karaops larapinta sp. nov., holotype, epigyne, caudal, Finke River, Northern Territory (QMS 110852) D Karaops sp. from Watarrka National Park, Northern Territory. Scale bars: 0.5 mm."figureDoi="10.3897/zookeys.1150.93760.figure5"httpUri="https://binary.pensoft.net/fig/816387"pageId="0"pageNumber="1">5C</figureCitation>
<figureCitationid="B334A0CB816E71872EC9480E4B506038"captionStart="Figure 3"captionStartId="F5"captionText="Figure 3. Epigynes and endogynes of members of the Central Desert species group A Karaops larapinta sp. nov., holotype female, epigyne, Finke River, Northern Territory (QMS 110852) B same, endogyne C Karaops mparntwe sp. nov., holotype female, epigyne, Alice Springs, Northern Territory (AMT AA 10.852), pink = copulatory openings, blue = median field, yellow = lateral lobes / epigynal plate D same, endogyne, yellow = accessory bulbs, blue = spermathecae, red = fertilization ducts, green = posterodorsal fold E same, epigyne F same, endogyne. Scale bars: 0.5 mm."figureDoi="10.3897/zookeys.1150.93760.figure3"httpUri="https://binary.pensoft.net/fig/816385"pageId="0"pageNumber="1">3B</figureCitation>
<taxonomicNameid="CEF4A192047CD17CB896AFDAF0A0346D"authorityName="Crews & Harvey"authorityYear="2011"class="Arachnida"family="Selenopidae"genus="Karaops"higherTaxonomySource="CoL"kingdom="Animalia"lsidName="Karaops"order="Araneae"pageId="0"pageNumber="1"phylum="Arthropoda"rank="genus">
<figureCitationid="3461E37365DDCD273D1E8ABC08359634"captionStart="Map 2"captionStartId="F4"captionText="Map 2. Species of the Central Desert species group; orange = juvenile from Morgan Range, light purple = Karaops deserticola, teal = Karaops ngarutjaranya, red = juveniles from Watarrka, blue = Karaops larapinta sp. nov., pink = Karaops kwartatuma sp. nov., yellow = Karaops pilkingtoni, x = Karaops mparntwe sp. nov., purple = Karaops vadlaadambara, green = Karaops manaayn. Juveniles are treated as distinct species based on molecular data."figureDoi="10.3897/zookeys.1150.93760.map2"httpUri="https://binary.pensoft.net/fig/816384"pageId="0"pageNumber="1">2</figureCitation>
<figureCitationid="60DDBCF10467C7C08CC1C6ACA5497F27"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Species and habitats of the Central Desert species group A Karaops kwartatuma sp. nov., holotype male, Ormiston Gorge, West MacDonnell Range National Park, Northern Territory (sel _ 1093, MAGNT A 004858) B Karaops deserticola, female holotype, Mount Lindsay, South Australia (SAM N 199350) C Karaops larapinta sp. nov., holotype female, Finke River, Northern Territory (QMS 110852) D Karaops larapinta sp. nov., paratype female, Finke River, Northern Territory (QMS 110852) E Habitat of Karaops sp. from Watarrka National Park, Northern Territory F Karaops sp., King's Canyon Rim Walk, Watarrka National Park, Northern Territory G habitat of Karaops kwartatuma sp. nov., Ormiston Gorge, Northern Territory, Australia. Scale bars: 1 mm."figureDoi="10.3897/zookeys.1150.93760.figure2"httpUri="https://binary.pensoft.net/fig/816382"pageId="0"pageNumber="1">2C, D</figureCitation>
) is found in either the Finke or Henbury subregion of the Finke Bioregion (exact coordinates unavailable). This bioregion comprises arid sandplains, major rivers, valleys, mulga (
<bibRefCitationid="CEEED2C752A42D1AFD553398B340B141"author="Bastin, G"journalOrPublisher="Cunninghamia"pageId="0"pageNumber="1"publicationUrl="https://www.dcceew.gov.au/environment/land/publications/acris-rangelands-2008-taking-pulse"refId="B4"refString="Bastin, G, 2008. Rangelands 2008 - Taking the Pulse, published on behalf of the ACRIS Management Committee by the National Land & Water Resources Audit, Canberra. https://www.dcceew.gov.au/environment/land/publications/acris-rangelands-2008-taking-pulse"title="Rangelands 2008 - Taking the Pulse, published on behalf of the ACRIS Management Committee by the National Land & Water Resources Audit, Canberra."url="https://www.dcceew.gov.au/environment/land/publications/acris-rangelands-2008-taking-pulse"year="2008">Bastin 2008</bibRefCitation>
<bibRefCitationid="091D42EE777764BCCB9182BAD2595820"author="Bastin, G"journalOrPublisher="Cunninghamia"pageId="0"pageNumber="1"publicationUrl="https://www.dcceew.gov.au/environment/land/publications/acris-rangelands-2008-taking-pulse"refId="B4"refString="Bastin, G, 2008. Rangelands 2008 - Taking the Pulse, published on behalf of the ACRIS Management Committee by the National Land & Water Resources Audit, Canberra. https://www.dcceew.gov.au/environment/land/publications/acris-rangelands-2008-taking-pulse"title="Rangelands 2008 - Taking the Pulse, published on behalf of the ACRIS Management Committee by the National Land & Water Resources Audit, Canberra."url="https://www.dcceew.gov.au/environment/land/publications/acris-rangelands-2008-taking-pulse"year="2008">Bastin 2008</bibRefCitation>
