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<document id="4E2D379426F4F1A7D3732204AA94BC4D" ID-CLB-Dataset="25217" ID-DOI="10.11646/zootaxa.4686.3.7" ID-GBIF-Dataset="f8824fbc-a420-4e93-937e-c3bde707227e" ID-ISSN="1175-5326" ID-Zenodo-Dep="3491614" ID-ZooBank="55E65352-A39B-4FC6-84F0-FA659B25A5C9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1571210711216" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2019" docId="6269CD49FFEEFFA6FF1AFC6033CF5C97" docLanguage="en" docName="zootaxa.4686.3.7.pdf" docOrigin="Zootaxa 4686 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Euconnus (Borneoconnus) sabahanus Franz 1992" docType="treatment" docVersion="7" lastPageNumber="427" masterDocId="9E50B531FFEBFFAEFF8DFFC7317B5D1C" masterDocTitle="Taxonomy of ‘ Euconnus complex’. Part XIX. Status of subgenus Borneoconnus Franz, with review of species (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="428" masterPageNumber="419" pageNumber="424" updateTime="1698759263074" updateUser="ExternalLinkService">
<mods:mods id="D1423B1221105EC6A9120C9A989468E2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="AB9C9DE62D0A3FBB8810084D8C614794">Taxonomy of ‘ Euconnus complex’. Part XIX. Status of subgenus Borneoconnus Franz, with review of species (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="1AA44D9F61093666CA4F946C5BAF4BB3">Jałoszyński, Paweł</mods:namePart>
</mods:name>
<mods:typeOfResource id="2A25086CCA1B2005E5A1225DBFA3322F">text</mods:typeOfResource>
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<mods:date id="75E80FA4569297A4940D94F8CF733E6E">2019</mods:date>
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<mods:number id="14302C91FB56825C059CD35350B039BC">2019-10-16</mods:number>
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<mods:number id="8A7EA69D300503C35DC072854F0850A4">4686</mods:number>
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<mods:classification id="888CA1B634517958957CC109259C7ABE">journal article</mods:classification>
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<mods:identifier id="0171A55B98F207508E39B1810731EFF8" type="DOI">10.11646/zootaxa.4686.3.7</mods:identifier>
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</mods:mods>
<treatment id="6269CD49FFEEFFA6FF1AFC6033CF5C97" ID-GBIF-Taxon="159721474" LSID="urn:lsid:plazi:treatment:6269CD49FFEEFFA6FF1AFC6033CF5C97" httpUri="http://treatment.plazi.org/id/6269CD49FFEEFFA6FF1AFC6033CF5C97" lastPageId="8" lastPageNumber="427" pageId="5" pageNumber="424">
<subSubSection id="A2DA2FD4FFEEFFABFF1AFC6033C75EDD" box="[151,700,934,961]" pageId="5" pageNumber="424" type="nomenclature">
<paragraph id="EA7F7C5FFFEEFFABFF1AFC6033C75EDD" blockId="5.[151,700,934,997]" box="[151,700,934,961]" pageId="5" pageNumber="424">
<heading id="B137CB33FFEEFFABFF1AFC6033C75EDD" bold="true" box="[151,700,934,961]" fontSize="11" level="1" pageId="5" pageNumber="424" reason="1">
<taxonomicName id="2DC007DCFFEEFFABFF1AFC6033C75EDD" authority="Franz" authorityName="Franz" authorityYear="1992" box="[151,700,934,961]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="424" phylum="Arthropoda" rank="species" species="sabahanus" subGenus="Borneoconnus">
<emphasis id="D8B4A04DFFEEFFABFF1AFC6033C75EDD" bold="true" box="[151,700,934,961]" pageId="5" pageNumber="424">
<emphasis id="D8B4A04DFFEEFFABFF1AFC6030685EDD" bold="true" box="[151,275,935,961]" italics="true" pageId="5" pageNumber="424">Euconnus</emphasis>
(
<emphasis id="D8B4A04DFFEEFFABFEA9FC6030AD5EDD" bold="true" box="[292,470,935,961]" italics="true" pageId="5" pageNumber="424">Borneoconnus</emphasis>
)
<emphasis id="D8B4A04DFFEEFFABFE6AFC6133125EDD" bold="true" box="[487,617,934,961]" italics="true" pageId="5" pageNumber="424">sabahanus</emphasis>
Franz
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2DA2FD4FFEEFFABFF1AFC0D30005EF8" box="[151,379,970,997]" pageId="5" pageNumber="424" type="description">
<paragraph id="EA7F7C5FFFEEFFABFF1AFC0D30005EF8" blockId="5.[151,700,934,997]" box="[151,379,970,997]" pageId="5" pageNumber="424">
(
<figureCitation id="72FB60DAFFEEFFABFF12FC0D305B5EF9" box="[159,288,970,997]" captionStart="FIGURES 1–7" captionStartId="4.[151,264,1224,1249]" captionTargetBox="[151,1433,180,1201]" captionTargetId="figure@4.[151,1436,174,1205]" captionTargetPageId="4" captionText="FIGURES 1–7. Dorsal habitus (1–4) and original sets of holotype labels (5–7) of Euconnus (Borneoconnus) spp. Euconnus laticlava Franz, male (1, 5); E. eremita Franz, male (2, 6); E. sabahanus, male (3, 7) and female (4)." figureDoi="http://doi.org/10.5281/zenodo.3491616" httpUri="https://zenodo.org/record/3491616/files/figure.png" pageId="5" pageNumber="424">Figs 3–4, 7</figureCitation>
,
<figureCitation id="72FB60DAFFEEFFABFEA1FC0D30085EF8" box="[300,371,970,996]" captionStart="FIGURES 12–23" captionStartId="7.[151,264,1752,1777]" captionTargetBox="[172,1426,187,1720]" captionTargetId="figure@7.[151,1436,174,1733]" captionText="FIGURES 12–23. Aedeagus in ventral (12, 14, 16, 18, 20, 22) and lateral (13, 15, 17, 19, 21, 23) views. Euconnus laticlava Franz (12–15); E. eremita Franz (16–19); E. sabahanus (20–23)." figureDoi="http://doi.org/10.5281/zenodo.3492109" httpUri="https://zenodo.org/record/3492109/files/figure.png" pageId="5" pageNumber="424">20–23</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A2DA2FD4FFEEFFABFF1AFBC83231593B" box="[151,842,1038,1063]" pageId="5" pageNumber="424" type="reference_group">
<paragraph id="EA7F7C5FFFEEFFABFF1AFBC83231593B" blockId="5.[151,842,1038,1063]" box="[151,842,1038,1063]" pageId="5" pageNumber="424">
<treatmentCitationGroup id="CAD05B71FFEEFFABFF1AFBC83231593B" box="[151,842,1038,1063]" pageId="5" pageNumber="424">
<taxonomicName id="2DC007DCFFEEFFABFF1AFBC833B7593B" authority="Franz, 1992: 902" authorityName="Franz" authorityPageNumber="902" authorityYear="1992" box="[151,716,1038,1063]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="424" phylum="Arthropoda" rank="species" species="sabahanus" subGenus="Borneoconnus">
<emphasis id="D8B4A04DFFEEFFABFF1AFBC83180593A" box="[151,251,1039,1062]" italics="true" pageId="5" pageNumber="424">Euconnus</emphasis>
(
<emphasis id="D8B4A04DFFEEFFABFE87FBC830E7593A" box="[266,412,1039,1062]" italics="true" pageId="5" pageNumber="424">Borneoconnus</emphasis>
)
<emphasis id="D8B4A04DFFEEFFABFE26FBC9336C593A" box="[427,535,1038,1062]" italics="true" pageId="5" pageNumber="424">sabahanus</emphasis>
<treatmentCitation id="6B615A4EFFEEFFABFD93FBC833B7593B" author="Franz, H." box="[542,716,1039,1063]" page="902" pageId="5" pageNumber="424" year="1992">
<bibRefCitation id="8E5101AEFFEEFFABFD93FBC833B7593B" author="Franz, H." box="[542,716,1039,1063]" pageId="5" pageNumber="424" pagination="859 - 953" refId="ref5142" refString="Franz, H. (1992) Monographie der Scydmaenidae (Coleoptera) von Sabah (NO-Borneo). Revue suisse de Zoologie, 99 (4), 859 - 953. https: // doi. org / 10.5962 / bhl. part. 79857" type="journal article" year="1992">Franz, 1992: 902</bibRefCitation>
</treatmentCitation>
</taxonomicName>
, figs 49–50.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A2DA2FD4FFEEFFABFF1AFB9235B758AF" pageId="5" pageNumber="424" type="materials_examined">
<paragraph id="EA7F7C5FFFEEFFABFF1AFB92339A583F" blockId="5.[151,1437,1108,2035]" pageId="5" pageNumber="424">
<emphasis id="D8B4A04DFFEEFFABFF1AFB9230DF5973" bold="true" box="[151,420,1108,1135]" pageId="5" pageNumber="424">Type material studied.</emphasis>
<materialsCitation id="5AA87602FFEEFFABFE26FB92307859C7" ID-GBIF-Occurrence="2426457913" collectingDate="1987-05-16" collectionCode="MHNG" country="Malaysia" determinerName="H. Franz" elevation="1600" location="SABAH" pageId="5" pageNumber="424" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="holotype">
<emphasis id="D8B4A04DFFEEFFABFE26FB9233905973" bold="true" box="[427,747,1108,1135]" pageId="5" pageNumber="424">
<typeStatus id="357BC2FDFFEEFFABFE26FB9233635973" box="[427,536,1109,1135]" pageId="5" pageNumber="424" type="holotype">Holotype</typeStatus>
[E
<collectingCountry id="92D73CCFFFEEFFABFDCEFB93339B5973" box="[579,736,1108,1135]" name="Malaysia" pageId="5" pageNumber="424">MALAYSIA</collectingCountry>
]:
</emphasis>
<specimenCount id="FCC6B7D6FFEEFFABFD7EFB9232705973" box="[755,779,1109,1135]" pageId="5" pageNumber="424" type="male">♂</specimenCount>
: three labels (
<figureCitation id="72FB60DAFFEEFFABFC2AFB9232965973" box="[935,1005,1109,1135]" captionStart="FIGURES 1–7" captionStartId="4.[151,264,1224,1249]" captionTargetBox="[151,1433,180,1201]" captionTargetId="figure@4.[151,1436,174,1205]" captionTargetPageId="4" captionText="FIGURES 1–7. Dorsal habitus (1–4) and original sets of holotype labels (5–7) of Euconnus (Borneoconnus) spp. Euconnus laticlava Franz, male (1, 5); E. eremita Franz, male (2, 6); E. sabahanus, male (3, 7) and female (4)." figureDoi="http://doi.org/10.5281/zenodo.3491616" httpUri="https://zenodo.org/record/3491616/files/figure.png" pageId="5" pageNumber="424">Fig. 7</figureCitation>
): “
<collectingRegion id="2804B2BDFFEEFFABFB81FB9235155973" box="[1036,1134,1109,1135]" country="Malaysia" name="Sabah" pageId="5" pageNumber="424">SABAH</collectingRegion>
/ Crocker Ra. /
<quantity id="2D38D1BAFFEEFFABFAAEFB9331D4598F" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.65" metricValueMin="1.55" pageId="5" pageNumber="424" unit="m" value="1600.0" valueMax="1650.0" valueMin="1550.0">
<elevation id="61ED9B6CFFEEFFABFAAEFB9331D4598F" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.65" metricValueMin="1.55" pageId="5" pageNumber="424" unit="m" value="1600.0" valueMax="1650.0" valueMin="1550.0">1550-1650 m</elevation>
</quantity>
,
<date id="9E7E5A9FFFEEFFABFF31FBBE3054598F" box="[188,303,1145,1171]" pageId="5" pageNumber="424" value="1987-05-16">
<collectingDate id="8E3AA377FFEEFFABFF31FBBE3054598F" box="[188,303,1145,1171]" pageId="5" pageNumber="424" value="1987-05-16">16.V.1987</collectingDate>
</date>
/ Burckhardt—Löbl” with “27a” on the reverse side [white, printed; reverse handwritten], “
<emphasis id="D8B4A04DFFEEFFABFACDFBBD3422598F" box="[1344,1369,1146,1171]" italics="true" pageId="5" pageNumber="424">E.</emphasis>
(
<emphasis id="D8B4A04DFFEEFFABFAE4FBBD307459AB" italics="true" pageId="5" pageNumber="424">Bor- neoconnus</emphasis>
) /
<taxonomicName id="2DC007DCFFEEFFABFEB9FB5A30D759AB" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[308,428,1181,1207]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="424" phylum="Arthropoda" rank="species" species="sabahanus" status="sp. nov.">
<emphasis id="D8B4A04DFFEEFFABFEB9FB5A30D759AB" box="[308,428,1181,1207]" italics="true" pageId="5" pageNumber="424">sabahanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C3871D36FFEEFFABFE38FB5A309159AB" box="[437,490,1181,1207]" pageId="5" pageNumber="424" rank="species">n.sp.</taxonomicNameLabel>
/ det.
<determinerName id="1FA3909BFFEEFFABFDB4FB5A33DA59AB" box="[569,673,1181,1207]" pageId="5" pageNumber="424">H. Franz</determinerName>
” [white, handwritten and printed], “
<typeStatus id="357BC2FDFFEEFFABFBCFFB5A35C759AB" box="[1090,1212,1181,1207]" pageId="5" pageNumber="424" type="holotype">Holotypus</typeStatus>
” [red, handwritten] (
<collectionCode id="8CD1E49AFFEEFFABFF12FB06318059C7" box="[159,251,1217,1243]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="5" pageNumber="424" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="5AA87602FFEEFFABFE9DFB06342059C7" ID-GBIF-Occurrence="2426457910" box="[272,1371,1217,1243]" collectingDate="1987-05-16" collectionCode="MHNG" country="Malaysia" determinerName="H. Franz" elevation="1600" location="SABAH" pageId="5" pageNumber="424" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="paratype">
<emphasis id="D8B4A04DFFEEFFABFE9DFB0630EA59C7" bold="true" box="[272,401,1217,1243]" pageId="5" pageNumber="424">
<typeStatus id="357BC2FDFFEEFFABFE9DFB0630F759C7" box="[272,396,1217,1243]" pageId="5" pageNumber="424" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="FCC6B7D6FFEEFFABFE16FB0630C959C7" box="[411,434,1217,1243]" pageId="5" pageNumber="424" type="male">♂</specimenCount>
, same data as for holotypes, except for a yellow “
<typeStatus id="357BC2FDFFEEFFABFC75FB0635E759C7" box="[1016,1180,1217,1243]" pageId="5" pageNumber="424" type="paratype">PARATYPUS</typeStatus>
” label (
<collectionCode id="8CD1E49AFFEEFFABFB7AFB06342859C7" box="[1271,1363,1217,1243]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="5" pageNumber="424" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="5AA87602FFEEFFABFAE5FB06300459E3" ID-GBIF-Occurrence="2426457909" collectionCode="NHMW" country="Malaysia" pageId="5" pageNumber="424" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="paratype">
<specimenCount id="FCC6B7D6FFEEFFABFAE5FB0634EC59C7" box="[1384,1431,1217,1243]" pageId="5" pageNumber="424" type="male">1 ♂</specimenCount>
, sama data (
<collectionCode id="8CD1E49AFFEEFFABFE98FB22300C59E3" box="[277,375,1253,1279]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="424" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
. Additionally
<materialsCitation id="5AA87602FFEEFFABFD91FB2232BE59E3" ID-GBIF-Occurrence="2426457912" box="[540,965,1253,1279]" collectionCode="NHMW" country="Malaysia" pageId="5" pageNumber="424" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Sabah" typeStatus="paratype">
<specimenCount id="FCC6B7D6FFEEFFABFD91FB22333D59E3" box="[540,582,1253,1279]" pageId="5" pageNumber="424" type="male">1 ♂</specimenCount>
and
<specimenCount id="FCC6B7D6FFEEFFABFDF1FB2233C059E3" box="[636,699,1253,1279]" pageId="5" pageNumber="424" type="female">2 ♀♀</specimenCount>
were studied (
<collectionCode id="8CD1E49AFFEEFFABFCD3FB2232C659E3" box="[862,957,1253,1279]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="424" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
with the same collecting data, but lacking identification labels; they presumably are
<typeStatus id="357BC2FDFFEEFFABFDE3FACE33A6583F" box="[622,733,1289,1315]" pageId="5" pageNumber="424" type="paratype">paratypes</typeStatus>
.
</paragraph>
<paragraph id="EA7F7C5FFFEEFFABFF4AFAEB35B758AF" blockId="5.[151,1437,1108,2035]" pageId="5" pageNumber="424">
<emphasis id="D8B4A04DFFEEFFABFF4AFAEB30DB585B" bold="true" box="[199,416,1324,1351]" pageId="5" pageNumber="424">Revised diagnosis.</emphasis>
BL of male
<quantity id="2D38D1BAFFEEFFABFDA4FAEA33B5585B" box="[553,718,1325,1351]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="5" pageNumber="424" unit="mm" value="1.25" valueMax="1.3" valueMin="1.2">1.20–1.30 mm</quantity>
; antennomere VIII in male slightly transverse, IX less than twice as broad as long and less than twice as broad as antennomere XI, convex dorsally and ventrally; pronotum broadest at base, strongly narrowing anterad; ventral apical plate of aedeagus subtrapezoidal with weakly rounded sides, its apex forming gradually tapered, small subtriangular lobe with its apical portion directed laterad.
</paragraph>
</subSubSection>
<subSubSection id="A2DA2FD4FFEEFFA6FF4AFA7B33685C7B" lastPageId="8" lastPageNumber="427" pageId="5" pageNumber="424" type="description">
<paragraph id="EA7F7C5FFFEEFFABFF4AFA7B35FE58E7" blockId="5.[151,1437,1108,2035]" pageId="5" pageNumber="424">
<emphasis id="D8B4A04DFFEEFFABFF4AFA7B300E58CB" bold="true" box="[199,373,1468,1495]" pageId="5" pageNumber="424">Redescription.</emphasis>
Body of male (
<figureCitation id="72FB60DAFFEEFFABFDBFFA7A330358CB" box="[562,632,1469,1495]" captionStart="FIGURES 1–7" captionStartId="4.[151,264,1224,1249]" captionTargetBox="[151,1433,180,1201]" captionTargetId="figure@4.[151,1436,174,1205]" captionTargetPageId="4" captionText="FIGURES 1–7. Dorsal habitus (1–4) and original sets of holotype labels (5–7) of Euconnus (Borneoconnus) spp. Euconnus laticlava Franz, male (1, 5); E. eremita Franz, male (2, 6); E. sabahanus, male (3, 7) and female (4)." figureDoi="http://doi.org/10.5281/zenodo.3491616" httpUri="https://zenodo.org/record/3491616/files/figure.png" pageId="5" pageNumber="424">Fig. 3</figureCitation>
) and female (
<figureCitation id="72FB60DAFFEEFFABFC93FA7A321E58CB" box="[798,869,1469,1495]" captionStart="FIGURES 1–7" captionStartId="4.[151,264,1224,1249]" captionTargetBox="[151,1433,180,1201]" captionTargetId="figure@4.[151,1436,174,1205]" captionTargetPageId="4" captionText="FIGURES 1–7. Dorsal habitus (1–4) and original sets of holotype labels (5–7) of Euconnus (Borneoconnus) spp. Euconnus laticlava Franz, male (1, 5); E. eremita Franz, male (2, 6); E. sabahanus, male (3, 7) and female (4)." figureDoi="http://doi.org/10.5281/zenodo.3491616" httpUri="https://zenodo.org/record/3491616/files/figure.png" pageId="5" pageNumber="424">Fig. 4</figureCitation>
) moderately slender and moderately convex, BL
<quantity id="2D38D1BAFFEEFFABFF1AFA26304658E7" box="[151,317,1505,1531]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="5" pageNumber="424" unit="mm" value="1.25" valueMax="1.3" valueMin="1.2">1.20–1.30 mm</quantity>
; pigmentation uniformly dark brown; vestiture slightly lighter than cuticle.
</paragraph>
<paragraph id="EA7F7C5FFFEEFFABFF4AF9C2333C5A07" blockId="5.[151,1437,1108,2035]" pageId="5" pageNumber="424">
Head elongate rhomboidal, broadest at eyes, HL
<quantity id="2D38D1BAFFEEFFABFD6FF9C332FE5B03" box="[738,901,1540,1567]" metricMagnitude="-4" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="5" pageNumber="424" unit="mm" value="0.265" valueMax="0.28" valueMin="0.25">0.25–0.28 mm</quantity>
, HW
<quantity id="2D38D1BAFFEEFFABFC4EF9C235135B03" box="[963,1128,1541,1567]" metricMagnitude="-4" metricUnit="m" metricValue="2.15" metricValueMax="2.3" metricValueMin="2.0" pageId="5" pageNumber="424" unit="mm" value="0.215" valueMax="0.23" valueMin="0.2">0.20–0.23 mm</quantity>
; vertex and frons confluent, convex; supraantennal tubercles indistinct; eyes small, weakly convex; tempora in dorsal view about 3 × as long as eyes. Punctures on vertex and frons small and inconspicuous; setae short, moderately dense, suberect; tempora and vertex with long and dense bristles. Antennae short, AnL
<quantity id="2D38D1BAFFEEFFABFC82F9B7321B5B97" box="[783,864,1648,1675]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="424" unit="mm" value="50.0">50 mm</quantity>
; antennomeres I–II each strongly elongate, III–VIII each slightly transverse, IX modified, strongly transverse, asymmetrical, distinctly, but not strongly projecting laterad, convex dorsally and ventrally, entirely setose, X slightly modified, distinctly transverse, asymmetrical, with broad basal mesal projection, entirely setose; XI distinctly narrower and longer than X, nearly symmetrical, indistinctly elongate, with subconical apex.
</paragraph>
<paragraph id="EA7F7C5FFFEEFFABFF4AF8E2357F5AEF" blockId="5.[151,1437,1108,2035]" pageId="5" pageNumber="424">
Pronotum in dorsal view subtrapezoidal, broadest at base and strongly narrowing anteriorly; PL
<quantity id="2D38D1BAFFEEFFABFA8DF8E2341C5A23" box="[1280,1383,1829,1855]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="424" unit="mm" value="0.3">0.30 mm</quantity>
, PW
<quantity id="2D38D1BAFFEEFFABFF1AF88F30475A7F" box="[151,316,1864,1891]" metricMagnitude="-4" metricUnit="m" metricValue="2.55" metricValueMax="2.6" metricValueMin="2.5" pageId="5" pageNumber="424" unit="mm" value="0.255" valueMax="0.26" valueMin="0.25">0.25–0.26 mm</quantity>
; anterior margin nearly straight, sides nearly straight, posterior corners obtuse-angled; posterior margin straight at middle and bent anterad at each side in front of humerus; pronotal base with two pairs of small and shallow pits (outer pits barely discernible), area between median pits forming slightly elevated, diffuse longitudinal carina; sublateral carinae indistinct under light microscope. Punctures on pronotal disc fine, inconspicuous; setae short, moderately dense, suberect, sides of pronotum with dense, long bristles.
</paragraph>
<caption id="BEBF2CD7FFEDFFA8FF1AF8CE32635AE5" ID-DOI="http://doi.org/10.5281/zenodo.3491618" ID-Zenodo-Dep="3491618" httpUri="https://zenodo.org/record/3491618/files/figure.png" pageId="6" pageNumber="425" startId="6.[151,264,1801,1826]" targetBox="[187,1400,166,1782]" targetPageId="6">
<paragraph id="EA7F7C5FFFEDFFA8FF1AF8CE32635AE5" blockId="6.[151,1436,1801,2042]" pageId="6" pageNumber="425">
<emphasis id="D8B4A04DFFEDFFA8FF1AF8CE303D5A3E" bold="true" box="[151,326,1801,1826]" pageId="6" pageNumber="425">FIGURES 8–11.</emphasis>
Morphological structures of
<taxonomicName id="2DC007DCFFEDFFA8FDE3F8CC32085A3D" authority="Franz" authorityName="Franz" box="[622,883,1802,1826]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="425" phylum="Arthropoda" rank="species" species="laticlava">
<emphasis id="D8B4A04DFFEDFFA8FDE3F8CC324A5A3E" box="[622,817,1802,1826]" italics="true" pageId="6" pageNumber="425">Euconnus laticlava</emphasis>
Franz
</taxonomicName>
, male. Head, prothorax and elytral base in dorsal view (8); head, prothorax and anterior portion of mesothorax in ventral view (9); right antennal clava in ventral view (10); pterothorax in ventral view (11). Abbreviations: abp, antebasal pit; an 9–11, antennomere IX–XI; bef, basal elytral fovea; bs, basistipes; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; lp2–3, labial palpomere II–III; md, mandible; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp1–4, maxillary palpomere I–IV; nss, notosternal suture; occ, occipital constriction; pcr, procoxal rest; ppf, palpifer; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum; vt, vertex.
</paragraph>
</caption>
<caption id="BEBF2CD7FFECFFA9FF1AF91F325C5A09" ID-DOI="http://doi.org/10.5281/zenodo.3492109" ID-Zenodo-Dep="3492109" httpUri="https://zenodo.org/record/3492109/files/figure.png" pageId="7" pageNumber="426" startId="7.[151,264,1752,1777]" targetBox="[172,1426,187,1720]" targetPageId="7">
<paragraph id="EA7F7C5FFFECFFA9FF1AF91F325C5A09" blockId="7.[151,1437,1752,1813]" pageId="7" pageNumber="426">
<emphasis id="D8B4A04DFFECFFA9FF1AF91F302E5BED" bold="true" box="[151,341,1752,1777]" pageId="7" pageNumber="426">FIGURES 12–23.</emphasis>
Aedeagus in ventral (12, 14, 16, 18, 20, 22) and lateral (13, 15, 17, 19, 21, 23) views.
<taxonomicName id="2DC007DCFFECFFA9FB55F91D31AA5A09" authority="Franz" authorityName="Franz" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="426" phylum="Arthropoda" rank="species" species="laticlava">
<emphasis id="D8B4A04DFFECFFA9FB55F91D34E65BED" box="[1240,1437,1753,1777]" italics="true" pageId="7" pageNumber="426">Euconnus laticlava</emphasis>
Franz
</taxonomicName>
(12–15);
<taxonomicName id="2DC007DCFFECFFA9FEB8F93930A55A09" authority="Franz" authorityName="Franz" box="[309,478,1790,1813]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="426" phylum="Arthropoda" rank="species" species="eremita">
<emphasis id="D8B4A04DFFECFFA9FEB8F93930E65A09" box="[309,413,1790,1813]" italics="true" pageId="7" pageNumber="426">E. eremita</emphasis>
Franz
</taxonomicName>
(16–19);
<taxonomicName id="2DC007DCFFECFFA9FDCCF93933B25A09" authorityName="Jałoszyński" authorityYear="2019" box="[577,713,1789,1813]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="426" phylum="Arthropoda" rank="species" species="sabahanus">
<emphasis id="D8B4A04DFFECFFA9FDCCF93933B25A09" box="[577,713,1789,1813]" italics="true" pageId="7" pageNumber="426">E. sabahanus</emphasis>
</taxonomicName>
(20–23).
</paragraph>
</caption>
<paragraph id="EA7F7C5FFFECFFA9FF4AF88434F25AB9" blockId="7.[151,1437,1859,2030]" pageId="7" pageNumber="426">
Elytra together suboval and slightly flattened, broadest distinctly in front of middle, EL
<quantity id="2D38D1BAFFECFFA9FB31F88434185A42" box="[1212,1379,1859,1886]" metricMagnitude="-4" metricUnit="m" metricValue="6.8999999999999995" metricValueMax="7.3" metricValueMin="6.5" pageId="7" pageNumber="426" unit="mm" value="0.69" valueMax="0.73" valueMin="0.65">0.65–0.73 mm</quantity>
, EW
<quantity id="2D38D1BAFFECFFA9FF1AF8A030465A9E" box="[151,317,1895,1922]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" metricValueMax="5.0" metricValueMin="4.8" pageId="7" pageNumber="426" unit="mm" value="0.49" valueMax="0.5" valueMin="0.48">0.48–0.50 mm</quantity>
, EI 1.30–1.53; basal impressions shallow but distinct, humeral calli well-defined; apices separately rounded. Punctures on elytral disc diffuse, superficial, inconspicuous; setae moderately long and dense, suberect.
</paragraph>
<paragraph id="EA7F7C5FFFECFFA9FF4AF87733A15AD5" blockId="7.[151,1437,1859,2030]" box="[199,730,1967,1994]" pageId="7" pageNumber="426">Legs moderately long and slender; unmodified.</paragraph>
<paragraph id="EA7F7C5FFFECFFA6FF4AF81432ED5DE7" blockId="7.[151,1437,1859,2030]" lastBlockId="8.[151,1436,153,395]" lastPageId="8" lastPageNumber="427" pageId="7" pageNumber="426">
Aedeagus (
<figureCitation id="72FB60DAFFECFFA9FEC9F81430BF5AF2" box="[324,452,2003,2030]" captionStart="FIGURES 12–23" captionStartId="7.[151,264,1752,1777]" captionTargetBox="[172,1426,187,1720]" captionTargetId="figure@7.[151,1436,174,1733]" captionText="FIGURES 12–23. Aedeagus in ventral (12, 14, 16, 18, 20, 22) and lateral (13, 15, 17, 19, 21, 23) views. Euconnus laticlava Franz (12–15); E. eremita Franz (16–19); E. sabahanus (20–23)." figureDoi="http://doi.org/10.5281/zenodo.3492109" httpUri="https://zenodo.org/record/3492109/files/figure.png" pageId="7" pageNumber="426">Figs 20–23</figureCitation>
) stout, AeL
<quantity id="2D38D1BAFFECFFA9FDC0F81333C85AF2" box="[589,691,2004,2030]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="7" pageNumber="426" unit="mm" value="0.18">0.18 mm</quantity>
, median lobe in ventral view with ventral apical plate subtrapezoi- dal with slightly rounded sides, its apex gradually tapered to form a short, subtriangular rounded apical lobe shifted to the left and with its tip directed laterad; parameral apices reaching distal third of ventral apical plate, each with 4 long apical setae and one short, additionally with one subapical seta.
</paragraph>
<paragraph id="EA7F7C5FFFE3FFA6FF4AFEC233685C7B" blockId="8.[151,1436,153,395]" pageId="8" pageNumber="427">
<emphasis id="D8B4A04DFFE3FFA6FF4AFEC2305E5C03" bold="true" box="[199,293,261,287]" pageId="8" pageNumber="427">Female.</emphasis>
Similar to male (
<figureCitation id="72FB60DAFFE3FFA6FE6EFEC2335E5C03" box="[483,549,261,287]" captionStart="FIGURES 1–7" captionStartId="4.[151,264,1224,1249]" captionTargetBox="[151,1433,180,1201]" captionTargetId="figure@4.[151,1436,174,1205]" captionTargetPageId="4" captionText="FIGURES 1–7. Dorsal habitus (1–4) and original sets of holotype labels (5–7) of Euconnus (Borneoconnus) spp. Euconnus laticlava Franz, male (1, 5); E. eremita Franz, male (2, 6); E. sabahanus, male (3, 7) and female (4)." figureDoi="http://doi.org/10.5281/zenodo.3491616" httpUri="https://zenodo.org/record/3491616/files/figure.png" pageId="8" pageNumber="427">Fig. 4</figureCitation>
) but with unmodified antennal clava and much smaller eyes; BL
<quantity id="2D38D1BAFFE3FFA6FB7EFEC234E35C03" box="[1267,1432,261,287]" metricMagnitude="-3" metricUnit="m" metricValue="1.345" metricValueMax="1.38" metricValueMin="1.31" pageId="8" pageNumber="427" unit="mm" value="1.345" valueMax="1.38" valueMin="1.31">1.31–1.38 mm</quantity>
; HL
<quantity id="2D38D1BAFFE3FFA6FF4EFEEE30125C5F" box="[195,361,297,323]" metricMagnitude="-4" metricUnit="m" metricValue="2.85" metricValueMax="2.9" metricValueMin="2.8" pageId="8" pageNumber="427" unit="mm" value="0.285" valueMax="0.29" valueMin="0.28">0.28–0.29 mm</quantity>
, HW
<quantity id="2D38D1BAFFE3FFA6FE21FEEF332A5C5F" box="[428,593,296,323]" metricMagnitude="-4" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="8" pageNumber="427" unit="mm" value="0.245" valueMax="0.25" valueMin="0.24">0.24–0.25 mm</quantity>
, AnL
<quantity id="2D38D1BAFFE3FFA6FD1AFEEE33855C5F" box="[663,766,297,323]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="8" pageNumber="427" unit="mm" value="0.48">0.48 mm</quantity>
; PL
<quantity id="2D38D1BAFFE3FFA6FCBFFEEF32A35C5F" box="[818,984,296,323]" metricMagnitude="-4" metricUnit="m" metricValue="3.3999999999999995" metricValueMax="3.5" metricValueMin="3.3" pageId="8" pageNumber="427" unit="mm" value="0.33999999999999997" valueMax="0.35" valueMin="0.33">0.33–0.35 mm</quantity>
, PW
<quantity id="2D38D1BAFFE3FFA6FB9BFEEE35C65C5F" box="[1046,1213,297,323]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" metricValueMax="3.0" metricValueMin="2.8" pageId="8" pageNumber="427" unit="mm" value="0.29" valueMax="0.3" valueMin="0.28">0.28–0.30 mm</quantity>
; EL
<quantity id="2D38D1BAFFE3FFA6FB7FFEEF34E35C5F" box="[1266,1432,296,323]" metricMagnitude="-4" metricUnit="m" metricValue="7.25" metricValueMax="7.5" metricValueMin="7.0" pageId="8" pageNumber="427" unit="mm" value="0.725" valueMax="0.75" valueMin="0.7">0.70–0.75 mm</quantity>
, EW
<quantity id="2D38D1BAFFE3FFA6FF44FE8B30155C7B" box="[201,366,332,359]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" metricValueMax="5.5" metricValueMin="5.3" pageId="8" pageNumber="427" unit="mm" value="0.54" valueMax="0.55" valueMin="0.53">0.53–0.55 mm</quantity>
, EI 1.33–1.36.
</paragraph>
</subSubSection>
<subSubSection id="A2DA2FD4FFE3FFA6FF4AFEB733CF5C97" box="[199,692,368,395]" pageId="8" pageNumber="427" type="distribution">
<paragraph id="EA7F7C5FFFE3FFA6FF4AFEB733CF5C97" blockId="8.[151,1436,153,395]" box="[199,692,368,395]" pageId="8" pageNumber="427">
<emphasis id="D8B4A04DFFE3FFA6FF4AFEB7301A5C97" bold="true" box="[199,353,368,395]" pageId="8" pageNumber="427">Distribution.</emphasis>
E
<collectingCountry id="92D73CCFFFE3FFA6FE0DFEB630925C97" box="[384,489,369,395]" name="Malaysia" pageId="8" pageNumber="427">Malaysia</collectingCountry>
, northern Borneo.
</paragraph>
</subSubSection>
</treatment>
</document>
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