171 lines
19 KiB
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171 lines
19 KiB
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<document id="F3A454A07754FFBD64FC632973F36EE4" ID-CLB-Dataset="31886" ID-DOI="10.11646/zootaxa.4329.3.3" ID-GBIF-Dataset="f1889f03-d51c-4de1-a1b5-339de952df0a" ID-ISSN="1175-5326" ID-Zenodo-Dep="1002542" ID-ZooBank="178F0C4B-Bff6-41B1-8Ba5-59Aea19F12F4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507185220440" checkinUser="plazi" docAuthor="Russell-Smith, Anthony" docDate="2017" docId="705F87EA497BFF9B6585FEC4FD55FE73" docLanguage="en" docName="zootaxa.4329.3.3.pdf" docOrigin="Zootaxa 4329 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Setaphis mccowani Russell-Smith 2017, sp. n." docType="treatment" docVersion="7" lastPageNumber="248" masterDocId="8C66FF924971FF906512FF88FFA4FFFB" masterDocTitle="New species and new records of ground spiders (Araneae: Gnaphosidae) from Cyprus" masterLastPageNumber="255" masterPageNumber="237" pageNumber="247" updateTime="1698469338266" updateUser="ExternalLinkService">
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<mods:mods id="27A3A1A0115D78436059697F5756EEB9" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="F00594CE5C1FD3C03C7BB09A4E2FE710">
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<mods:title id="FBAB949456CDCF1E17A6742BECB36A37">New species and new records of ground spiders (Araneae: Gnaphosidae) from Cyprus</mods:title>
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</mods:titleInfo>
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<mods:name id="F593A5DAC0B10D16B4F22169CDF35942" type="personal">
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<mods:roleTerm id="6D7E2B40DD764053D6143194D7D411C0">Author</mods:roleTerm>
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<mods:namePart id="7DFCE2EE065C40DFD06E52F6D6C0310A">Russell-Smith, Anthony</mods:namePart>
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<mods:typeOfResource id="BC73931773E2B89B64913ECDB71A0DF2">text</mods:typeOfResource>
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<mods:title id="E4D366E9882605228105776BA65486F9">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="E47561DE891A37A64A8C34570D44AE1E">
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<mods:date id="379C2990A3D6FC56F7FA3E3644DA1344">2017</mods:date>
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<mods:detail id="ABDA24E4AEB51DA2BD7A903225678E3C" type="pubDate">
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<mods:number id="8E3C2C44248A7B84C6F9F9B3792FF734">2017-10-05</mods:number>
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<mods:number id="10C92552F2F0FAF3E903DE73CADC5516">4329</mods:number>
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<mods:detail id="05B411788815BF654CCFF250B61DDA32" type="issue">
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<mods:number id="71C07A896D53F2330ECEFB7457525A85">3</mods:number>
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<mods:classification id="FD94EF53EB3A524CD24CDCBBF69BAF44">journal article</mods:classification>
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<mods:identifier id="A95E65ED685C725013A4C8767B8D99EA" type="CLB-Dataset">31886</mods:identifier>
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<mods:identifier id="D1FA9D143C897D633C570D9ED8EF0886" type="DOI">10.11646/zootaxa.4329.3.3</mods:identifier>
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<mods:identifier id="9ADE8F36FE6F2D036CB7794CFBEFFAA7" type="GBIF-Dataset">f1889f03-d51c-4de1-a1b5-339de952df0a</mods:identifier>
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</mods:mods>
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<treatment id="705F87EA497BFF9B6585FEC4FD55FE73" ID-DOI="http://doi.org/10.5281/zenodo.6051693" ID-GBIF-Taxon="135111949" ID-Zenodo-Dep="6051693" LSID="urn:lsid:plazi:treatment:705F87EA497BFF9B6585FEC4FD55FE73" httpUri="http://treatment.plazi.org/id/705F87EA497BFF9B6585FEC4FD55FE73" lastPageId="11" lastPageNumber="248" pageId="10" pageNumber="247">
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<subSubSection id="B0EC6577497BFF9A6585FEC4FEB0FE7C" pageId="10" pageNumber="247" type="nomenclature">
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<paragraph id="F84936FC497BFF9A6585FEC4FE6CFE9D" blockId="10.[151,456,332,391]" box="[151,456,332,358]" pageId="10" pageNumber="247">
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<heading id="A3018190497BFF9A6585FEC4FE6CFE9D" bold="true" box="[151,456,332,358]" fontSize="11" level="1" pageId="10" pageNumber="247" reason="1">
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<emphasis id="CA82EAEE497BFF9A6585FEC4FE6CFE9D" bold="true" box="[151,456,332,358]" pageId="10" pageNumber="247">
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<taxonomicName id="3FF64D7F497BFF9A6585FEC4FEDAFE9D" ID-CoL="94HPR" authority="Russell-Smith, 2017" authorityName="Russell-Smith" authorityYear="2017" box="[151,382,332,358]" class="Arachnida" family="Gnaphosidae" genus="Setaphis" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="247" phylum="Arthropoda" rank="species" species="mccowani" status="sp. nov.">
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<emphasis id="CA82EAEE497BFF9A6585FEC4FEDAFE9D" bold="true" box="[151,382,332,358]" italics="true" pageId="10" pageNumber="247">Setaphis mccowani</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="D1B15795497BFF9A6494FEC4FE6CFE9D" box="[390,456,332,358]" pageId="10" pageNumber="247" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="F84936FC497BFF9A6585FEE6FEB0FE7C" blockId="10.[151,456,332,391]" box="[151,276,366,391]" pageId="10" pageNumber="247">
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<figureCitation id="60CD2A79497BFF9A6585FEE6FEB0FE7C" box="[151,276,366,391]" captionStart="FIGURES 27 – 28" captionStartId="10.[151,264,1919,1941]" captionTargetBox="[172,1430,1321,1880]" captionTargetId="figure@10.[151,1436,1309,1897]" captionTargetPageId="10" captionText="FIGURES 27 – 28. Setaphis mccowani sp. n. holotype. 27. Epigyne. 28. Vulva. S 1, S 2: ventral and dorsal chambers of bipartite spermathecae." httpUri="https://zenodo.org/record/1002558/files/figure.png" pageId="10" pageNumber="247">Figs 27–28</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="B0EC6577497BFF9A6585FE3EFEBFFD78" pageId="10" pageNumber="247" type="diagnosis">
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<paragraph id="F84936FC497BFF9A6585FE3EFEBFFD78" blockId="10.[151,1437,438,1256]" pageId="10" pageNumber="247">
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<emphasis id="CA82EAEE497BFF9A6585FE3EFEB5FE34" bold="true" box="[151,273,438,463]" pageId="10" pageNumber="247">Diagnosis.</emphasis>
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By having metatarsal preening combs and characteristically coiled copulatory ducts, this species has the distinctive characters of the genus
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<taxonomicName id="3FF64D7F497BFF9A6726FE53FD37FE08" box="[564,659,475,499]" class="Arachnida" family="Gnaphosidae" genus="Setaphis" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="247" phylum="Arthropoda" rank="genus">
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<emphasis id="CA82EAEE497BFF9A6726FE53FD37FE08" box="[564,659,475,499]" italics="true" pageId="10" pageNumber="247">Setaphis</emphasis>
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</taxonomicName>
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. It is distinguished by the wide, open epigynal cavity formed by separated anterior margins and by a medially pointed (V-shaped) posterior margin and by the characteristically coiled copulatory ducts located only in the posterior part of the epigynal cavity and ending to bipartite bean shaped spermathecae. None of the currently known species of the genus share this combination of characters (but see comments).
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</paragraph>
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</subSubSection>
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<subSubSection id="B0EC6577497BFF9A65D5FD06FAEBFD5C" box="[199,1359,654,679]" pageId="10" pageNumber="247" type="materials_examined">
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<paragraph id="F84936FC497BFF9A65D5FD06FAEBFD5C" blockId="10.[151,1437,438,1256]" box="[199,1359,654,679]" pageId="10" pageNumber="247">
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<materialsCitation id="489E3CA1497BFF9A65D5FD06FAEBFD5C" ID-GBIF-Occurrence="1661822831" box="[199,1359,654,679]" collectingDate="2009-07-01" collectorName="D. McCowan." country="Cyprus" location="Agridi" municipality="Dali" pageId="10" pageNumber="247" specimenCount="1" typeStatus="holotype">
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<emphasis id="CA82EAEE497BFF9A65D5FD06FE99FD5C" bold="true" box="[199,317,654,679]" pageId="10" pageNumber="247">
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<typeStatus id="274D885E497BFF9A65D5FD06FE9CFD5C" box="[199,312,654,679]" pageId="10" pageNumber="247" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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female,
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<collectingCountry id="80E1766C497BFF9A648CFD07FE56FD5C" box="[414,498,655,679]" name="Cyprus" pageId="10" pageNumber="247">Cyprus</collectingCountry>
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,
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<location id="FD296027497BFF9A64EFFD06FDE2FD5C" LSID="urn:lsid:plazi:treatment:705F87EA497BFF9B6585FEC4FD55FE73:FD296027497BFF9A64EFFD06FDE2FD5C" box="[509,582,654,679]" country="Cyprus" municipality="Dali" name="Agridi" pageId="10" pageNumber="247">Agridi</location>
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near
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<collectingMunicipality id="182DAC86497BFF9A6796FD06FD13FD5C" box="[644,695,654,679]" pageId="10" pageNumber="247">Dali</collectingMunicipality>
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, grass in lawn of a garden,
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<collectingDate id="9C0CE9D4497BFF9A66FBFD07FBD1FD5C" box="[1001,1141,655,679]" pageId="10" pageNumber="247" value="2009-07-01">
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<date id="8C48103C497BFF9A66FBFD07FBD1FD5C" box="[1001,1141,655,679]" pageId="10" pageNumber="247" value="2009-07-01">01.VII.2009</date>
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</collectingDate>
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, leg.
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<collectorName id="5503532A497BFF9A61A2FD07FAEBFD5C" box="[1200,1359,655,679]" pageId="10" pageNumber="247">D. McCowan.</collectorName>
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="B0EC6577497BFF9A65D5FD3AFF4BFCE8" pageId="10" pageNumber="247" type="etymology">
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<paragraph id="F84936FC497BFF9A65D5FD3AFF4BFCE8" blockId="10.[151,1437,438,1256]" pageId="10" pageNumber="247">
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<emphasis id="CA82EAEE497BFF9A65D5FD3AFEEAFD30" bold="true" box="[199,334,690,715]" pageId="10" pageNumber="247">Etymology.</emphasis>
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The specific name is a noun in apposition, in honour of Duncan McCowan (
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<collectingCountry id="80E1766C497BFF9A61C2FD3BFA82FD30" box="[1232,1318,691,715]" name="Cyprus" pageId="10" pageNumber="247">Cyprus</collectingCountry>
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), who has contributed substantially to the collection of the material here presented and is the only active arachnologist in
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<collectingCountry id="80E1766C497BFF9A6585FD73FF48FCE8" box="[151,236,763,787]" name="Cyprus" pageId="10" pageNumber="247">Cyprus</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="B0EC6577497BFF9A65D5FC96FA39FB13" pageId="10" pageNumber="247" type="description">
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<paragraph id="F84936FC497BFF9A65D5FC96FA39FB13" blockId="10.[151,1437,438,1256]" pageId="10" pageNumber="247">
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<emphasis id="CA82EAEE497BFF9A65D5FC96FD92FCCC" bold="true" box="[199,566,798,823]" pageId="10" pageNumber="247">
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Description. Female (
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<typeStatus id="274D885E497BFF9A64D6FC96FD8AFCCC" box="[452,558,798,823]" pageId="10" pageNumber="247" type="holotype">holotype</typeStatus>
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)
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</emphasis>
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. Small spider of yellow colouration. All femurs and Pa/Ti of the first two legs seem darker and possibly in nature they are brown, although in the only specimen available to us the colour appears faded. Measurements: TL 3.909; CL 1.249; CW 0.914; AL 2.309. Eye sizes and interdistances: AME 0.041, PME 0.038x0.051, PLE 0.046, ALE 0.061 AME-AME 0.025 (½ of AME diameter), AME-ALE almost touching, PME- PME (min/max) 0.038/0.057, PME-PLE 0.036. All eyes pearly white with black surroundings except for AME which are black. PER slightly procurved to straight, AER recurved. Cephalothorax with widening thoracic region and fovea at posterior third of its length. Right and left chelicera differently armed: retromargin with 2-3 denticles and promargin and 3-4 teeth and 1 denticle. Labium 0.176 long, 0.185 wide at its base. Sternum 0.768 long, 0.641 wide. Legs sizes are IV>I>II>III. Leg spination: Leg I: Fe 1-2d Mt
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<date id="8C48103C497BFF9A669BFBC8FC0CFBA3" box="[905,936,1088,1112]" pageId="10" pageNumber="247">4v</date>
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; Leg II: Fe 1-2d Mt
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<date id="8C48103C497BFF9A619EFBC8FB08FBA3" box="[1164,1196,1088,1112]" pageId="10" pageNumber="247">4v</date>
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; Leg III: Fe 6d Pa 1rl Ti/Mt spinose; Leg IV: Fe 4d Ti/Mt spinose. Epigyne (
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<figureCitation id="60CD2A79497BFF9A67E6FBECFCE0FB87" box="[756,836,1124,1149]" captionStart="FIGURES 27 – 28" captionStartId="10.[151,264,1919,1941]" captionTargetBox="[172,1430,1321,1880]" captionTargetId="figure@10.[151,1436,1309,1897]" captionTargetPageId="10" captionText="FIGURES 27 – 28. Setaphis mccowani sp. n. holotype. 27. Epigyne. 28. Vulva. S 1, S 2: ventral and dorsal chambers of bipartite spermathecae." httpUri="https://zenodo.org/record/1002558/files/figure.png" pageId="10" pageNumber="247">Fig. 27</figureCitation>
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) with broad medially separated anterior margin wider than lateral epigynal margins. Lateral epigynal margins parallel and short, leaving most of epigynal cavity open, medially merging to create a median posterior point. Vulva (
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<figureCitation id="60CD2A79497BFF9A6670FB24FC1EFB3F" box="[866,954,1196,1221]" captionStart="FIGURES 27 – 28" captionStartId="10.[151,264,1919,1941]" captionTargetBox="[172,1430,1321,1880]" captionTargetId="figure@10.[151,1436,1309,1897]" captionTargetPageId="10" captionText="FIGURES 27 – 28. Setaphis mccowani sp. n. holotype. 27. Epigyne. 28. Vulva. S 1, S 2: ventral and dorsal chambers of bipartite spermathecae." httpUri="https://zenodo.org/record/1002558/files/figure.png" pageId="10" pageNumber="247">Fig. 28</figureCitation>
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). Spermathecae with ventral bean-shaped chambers (S
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<quantity id="3F0E9B19497BFF9A6436FB58FEE8FB13" box="[292,332,1231,1256]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="10" pageNumber="247" unit="in" value="1.0">1 in</quantity>
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<figureCitation id="60CD2A79497BFF9A6440FB47FE06FB13" box="[338,418,1231,1256]" captionStart="FIGURES 27 – 28" captionStartId="10.[151,264,1919,1941]" captionTargetBox="[172,1430,1321,1880]" captionTargetId="figure@10.[151,1436,1309,1897]" captionTargetPageId="10" captionText="FIGURES 27 – 28. Setaphis mccowani sp. n. holotype. 27. Epigyne. 28. Vulva. S 1, S 2: ventral and dorsal chambers of bipartite spermathecae." httpUri="https://zenodo.org/record/1002558/files/figure.png" pageId="10" pageNumber="247">Fig. 27</figureCitation>
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) and with dorsal round chambers (S
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<quantity id="3F0E9B19497BFF9A6627FB58FCC4FB13" box="[821,864,1231,1256]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="10" pageNumber="247" unit="in" value="2.0">2 in</quantity>
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<figureCitation id="60CD2A79497BFF9A6675FB47FC13FB13" box="[871,951,1231,1256]" captionStart="FIGURES 27 – 28" captionStartId="10.[151,264,1919,1941]" captionTargetBox="[172,1430,1321,1880]" captionTargetId="figure@10.[151,1436,1309,1897]" captionTargetPageId="10" captionText="FIGURES 27 – 28. Setaphis mccowani sp. n. holotype. 27. Epigyne. 28. Vulva. S 1, S 2: ventral and dorsal chambers of bipartite spermathecae." httpUri="https://zenodo.org/record/1002558/files/figure.png" pageId="10" pageNumber="247">Fig. 28</figureCitation>
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) leading to coiled tubular copulatory ducts.
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</paragraph>
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</subSubSection>
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<caption id="AC896674497BFF9A6585F8F7FE8CF84F" httpUri="https://zenodo.org/record/1002558/files/figure.png" pageId="10" pageNumber="247" targetBox="[172,1430,1321,1880]" targetPageId="10">
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<paragraph id="F84936FC497BFF9A6585F8F7FE8CF84F" blockId="10.[151,1436,1919,1972]" pageId="10" pageNumber="247">
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<emphasis id="CA82EAEE497BFF9A6585F8F7FEF7F86E" bold="true" box="[151,339,1919,1941]" pageId="10" pageNumber="247">FIGURES 27–28.</emphasis>
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<taxonomicName id="3FF64D7F497BFF9A644BF8F7FDB8F86E" authority="Russell-Smith, 2017" authorityName="Russell-Smith" authorityYear="2017" box="[345,540,1919,1941]" class="Arachnida" family="Gnaphosidae" genus="Setaphis" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="247" phylum="Arthropoda" rank="species" species="mccowani" status="sp. nov.">
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<emphasis id="CA82EAEE497BFF9A644BF8F7FDB8F86E" box="[345,540,1919,1941]" italics="true" pageId="10" pageNumber="247">Setaphis mccowani</emphasis>
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</taxonomicName>
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<emphasis id="CA82EAEE497BFF9A6730F8F7FDFEF86E" bold="true" box="[546,602,1919,1941]" pageId="10" pageNumber="247">
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|||
|
|
<taxonomicNameLabel id="D1B15795497BFF9A6730F8F7FDFEF86E" box="[546,602,1919,1941]" pageId="10" pageNumber="247" rank="species">sp. n.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
holotype.
|
|||
|
|
<emphasis id="CA82EAEE497BFF9A67D4F8F7FD42F86E" bold="true" box="[710,742,1919,1941]" pageId="10" pageNumber="247">27.</emphasis>
|
|||
|
|
Epigyne.
|
|||
|
|
<emphasis id="CA82EAEE497BFF9A665EF8F7FCC8F86E" bold="true" box="[844,876,1919,1941]" pageId="10" pageNumber="247">28.</emphasis>
|
|||
|
|
Vulva. S1, S2: ventral and dorsal chambers of bipartite spermathecae.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<subSubSection id="B0EC6577497AFF9B65D5FF1FFF59FE9F" pageId="11" pageNumber="248" type="discussion">
|
|||
|
|
<paragraph id="F84936FC497AFF9B65D5FF1FFF59FE9F" blockId="11.[151,1436,151,392]" pageId="11" pageNumber="248">
|
|||
|
|
<emphasis id="CA82EAEE497AFF9B65D5FF1FFEEBFF4B" bold="true" box="[199,335,151,176]" pageId="11" pageNumber="248">Comments.</emphasis>
|
|||
|
|
In the revision by
|
|||
|
|
<bibRefCitation id="9C674B0D497AFF9B6739FF1FFCD4FF4B" author="Platnick" box="[555,880,151,176]" pageId="11" pageNumber="248" refString="Platnick, N. I. & Murphy, J. A. (1996) A review of the zelotine ground spider genus Setaphis (Araneae, Gnaphosidae). American Museum Novitates, 3162, 1 ‾ 23." type="journal article" year="1996">Platnick and Murphy (1996)</bibRefCitation>
|
|||
|
|
, the females of all the
|
|||
|
|
<taxonomicName id="3FF64D7F497AFF9B616BFF10FB7CFF4B" box="[1145,1240,152,176]" class="Arachnida" family="Gnaphosidae" genus="Setaphis" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="248" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="CA82EAEE497AFF9B616BFF10FB7CFF4B" box="[1145,1240,152,176]" italics="true" pageId="11" pageNumber="248">Setaphis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
species recorded form (
|
|||
|
|
<emphasis id="CA82EAEE497AFF9B65CFFF35FECFFF2F" box="[221,363,188,212]" italics="true" pageId="11" pageNumber="248">grosso modo</emphasis>
|
|||
|
|
) M-shaped epigynal margins and coiled copulatory ducts which cover the whole of the epigynal plate. Our species deviates from this pattern (V-shaped epigynal frame and copulatory ducts not so expanded), but still we believe that it conforms more to this genus rather than to
|
|||
|
|
<taxonomicName id="3FF64D7F497AFF9B6661FE8CFC60FEE7" box="[883,964,260,284]" class="Arachnida" family="Gnaphosidae" genus="Zelotes" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="248" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="CA82EAEE497AFF9B6661FE8CFC60FEE7" box="[883,964,260,284]" italics="true" pageId="11" pageNumber="248">Zelotes</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
or any other zelotine. When the matching male is found, it may turn out that it belongs to a new genus, but in the absence of it, we tentatively place it to
|
|||
|
|
<taxonomicName id="3FF64D7F497AFF9B6585FEC4FF52FE9F" box="[151,246,332,356]" class="Arachnida" family="Gnaphosidae" genus="Setaphis" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="248" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="CA82EAEE497AFF9B6585FEC4FF52FE9F" box="[151,246,332,356]" italics="true" pageId="11" pageNumber="248">Setaphis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="B0EC6577497AFF9B65D5FEE7FD55FE73" box="[199,753,367,392]" pageId="11" pageNumber="248" type="distribution">
|
|||
|
|
<paragraph id="F84936FC497AFF9B65D5FEE7FD55FE73" blockId="11.[151,1436,151,392]" box="[199,753,367,392]" pageId="11" pageNumber="248">
|
|||
|
|
<emphasis id="CA82EAEE497AFF9B65D5FEE7FEC4FE73" bold="true" box="[199,352,367,392]" pageId="11" pageNumber="248">Distribution.</emphasis>
|
|||
|
|
Known only from the
|
|||
|
|
<typeStatus id="274D885E497AFF9B6772FEF8FD34FE73" box="[608,656,368,392]" pageId="11" pageNumber="248">type</typeStatus>
|
|||
|
|
locality.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|