<documentid="8CD19FC246E1E7532546DA578C1C429A"ID-CLB-Dataset="304561"ID-DOI="10.5252/zoosystema-2024v46a24"ID-GBIF-Dataset="17ba5769-29c5-47f9-ba89-262a202fb056"ID-ISSN="1638-9387"ID-Zenodo-Dep="13941216"ID-ZooBank="7AB59C63-1DF0-43B2-B45A-301164A7BAA9"IM.bibliography_approvedBy="jonas"IM.illustrations_approvedBy="jonas"IM.materialsCitations_approvedBy="jonas"IM.metadata_approvedBy="jonas"IM.tables_approvedBy="jonas"IM.taxonomicNames_approvedBy="jonas"IM.treatmentCitations_approvedBy="jonas"IM.treatments_approvedBy="jonas"checkinTime="1727858212238"checkinUser="plazi"docAuthor="Piacentini, Luis Norberto, Marco, Yuri Marusik & Isaia, Marco"docDate="2024"docId="03E8D8349821FFC5FC2773B3FE673586"docLanguage="en"docName="zoosystema.2024.46.24.pdf"docOrigin="Zoosystema 46 (24)"docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a24.pdf"docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.9:Zoosystema.2020-.journal_article.type1"docStyleId="5CC576F744449C1860B4E72ABD884631"docStyleName="Zoosystema.2020-.journal_article.type1"docStyleVersion="9"docTitle="Vesubia Simon 1909"docType="treatment"docVersion="5"lastPageNumber="620"masterDocId="FFD1A04C9825FFC0FFF17046FFF23128"masterDocTitle="Unraveling the monotypy of Vesubia Simon, 1909 and its relationships to Alopecosa Simon, 1885 (Araneae, Lycosidae)"masterLastPageNumber="629"masterPageNumber="617"pageNumber="619"updateTime="1729090974912"updateUser="ExternalLinkService"zenodo-license-document="CC0-1.0"zenodo-license-figures="CC0-1.0"zenodo-license-treatments="UNSPECIFIED">
<mods:titleid="DC94E1519CA0265272090227726E814D">Unraveling the monotypy of Vesubia Simon, 1909 and its relationships to Alopecosa Simon, 1885 (Araneae, Lycosidae)</mods:title>
<mods:affiliationid="2999537367790BB86289377CBC9A06F9">Institute for Biological Problems of the North FEB RAS, Portovaya str., 18, Magadan 685000 (Russia) & Altai State University, Lenina Pr., 61, Barnaul (Russia) & Department of Zoology & Entomology, University of the Free State, 205 Nelson Mandela Drive, Park West Bloemfontein (South Africa)</mods:affiliation>
<mods:affiliationid="A4D86BB855D74FD9E57919B59222E68A">Laboratorio di Ecologia, Ecosistemi terrestri, Dipartimento di Scienze della Vita e Biologia dei Sistemi, Università di Torino, Via Accademia Albertina, 13, 10123 Torino (Italy) & National Biodiversity Future Center, Palermo 90133 (Italy)</mods:affiliation>
<bibRefCitationid="EFD014D39821FFC4FB8873B2FAF43527"author="SIMON E."box="[1145,1286,1012,1039]"pageId="4"pageNumber="619"pagination="335 - 449"refId="ref7279"refString="SIMON E. 1909. - Arachnides recueillis par L. Fea sur la cote occidentale d'Afrique. 2 e partie. Annali del Museo Civico di Storia Naturale di Genova 44: 335 - 449."type="journal article"year="1909">Simon, 1909</bibRefCitation>
<bibRefCitationid="EFD014D39821FFC4FC8B7478FBC2357E"author="SIMON E."box="[890,1072,1086,1110]"pageId="4"pageNumber="619"pagination="335 - 449"refId="ref7279"refString="SIMON E. 1909. - Arachnides recueillis par L. Fea sur la cote occidentale d'Afrique. 2 e partie. Annali del Museo Civico di Storia Naturale di Genova 44: 335 - 449."type="journal article"year="1909">Simon, 1909: 402</bibRefCitation>
<bibRefCitationid="EFD014D39821FFC4FAF0743EFA7635B8"author="SIMON E."box="[1281,1412,1144,1168]"pageId="4"pageNumber="619"pagination="82 - 91"refId="ref7222"refString="SIMON E. 1881. - Arachnides nouveaux ou rares de la faune francaise. Bulletin de la Societe zoologique de France 6 (3 - 4): 82 - 91."type="journal article"year="1881">Simon, 1881</bibRefCitation>
<bibRefCitationid="EFD014D39821FFC4FAD574F3FA5535E5"author="SIMON E."box="[1316,1447,1205,1229]"pageId="4"pageNumber="619"pagination="82 - 91"refId="ref7222"refString="SIMON E. 1881. - Arachnides nouveaux ou rares de la faune francaise. Bulletin de la Societe zoologique de France 6 (3 - 4): 82 - 91."type="journal article"year="1881">Simon, 1881</bibRefCitation>
)
</taxonomicName>
was described based on a female specimen collected in high alpine habitat near Saint-Martin-Vésubie (
<bibRefCitationid="EFD014D39821FFC4FC5A75E5FBC13493"author="SIMON E."box="[939,1075,1443,1467]"pageId="4"pageNumber="619"pagination="335 - 449"refId="ref7279"refString="SIMON E. 1909. - Arachnides recueillis par L. Fea sur la cote occidentale d'Afrique. 2 e partie. Annali del Museo Civico di Storia Naturale di Genova 44: 335 - 449."type="journal article"year="1909">Simon (1909)</bibRefCitation>
based on the presence of two retromarginal teeth, spination pattern and the eye configuration. It remained monotypic until, without specific justification,
<bibRefCitationid="EFD014D39821FFC4FAE5759EFA5C34D8"author="ROEWER C. F."box="[1300,1454,1496,1520]"pageId="4"pageNumber="619"refId="ref7151"refString="ROEWER C. F. 1955. - Katalog der Araneae von 1758 bis 1940, Vol. 2 a. Institut Royal des Sciences naturelles de Belgique, Bruxelles, 923 p."type="book"year="1955">Roewer (1955)</bibRefCitation>
<bibRefCitationid="EFD014D39821FFC4FA96764BFC923717"author="KARSCH F."pageId="4"pageNumber="619"pagination="77 - 83"refId="ref6202"refString="KARSCH F. 1880. - Mittheilung uber die von Herrn Dr. O. Finsch wahrend seiner polynesischen Reise gesammelten Myriopoden und Arachniden. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin 1880: 77 - 83."type="journal article"year="1880">Karsch, 1880</bibRefCitation>
<bibRefCitationid="EFD014D39821FFC4FC027661FB8A3717"author="KARSCH F."box="[1011,1144,1575,1599]"pageId="4"pageNumber="619"pagination="77 - 83"refId="ref6202"refString="KARSCH F. 1880. - Mittheilung uber die von Herrn Dr. O. Finsch wahrend seiner polynesischen Reise gesammelten Myriopoden und Arachniden. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin 1880: 77 - 83."type="journal article"year="1880">Karsch, 1880</bibRefCitation>
</taxonomicName>
, merely by mentioning them in his catalogue. Subsequent nomenclatural revisions (Roewer 1960;
<bibRefCitationid="EFD014D39821FFC4FCDC761AFC37375C"author="TONGIORGI P."box="[813,965,1628,1652]"pageId="4"pageNumber="619"pagination="105 - 125"refId="ref7418"refString="TONGIORGI P. 1977. - Fam. Lycosidae. In: La faune terrestre de l'ile de Sainte-Helene IV. Annales, Musee royal de l'Afrique centrale, Sciences zoologiques (Zool. - Ser. 8 °) 220: 105 - 125."type="journal article"year="1977">Tongiorgi 1977</bibRefCitation>
;
<bibRefCitationid="EFD014D39821FFC4FC20761AFB84375C"author="YU L. M. & SONG D. X."box="[977,1142,1628,1652]"pageId="4"pageNumber="619"pagination="234 - 244"refId="ref7597"refString="YU L. M. & SONG D. X. 1988 - On new species of wolf spiders from China (Araneae: Lycosidae). Acta Zootaxonomica Sinica, 13: 234 - 244."type="journal article"year="1988">Yu & Song 1988</bibRefCitation>
;
<bibRefCitationid="EFD014D39821FFC4FB73761AFAE1375C"author="LOGUNOV D. V."box="[1154,1299,1628,1652]"pageId="4"pageNumber="619"pagination="475 - 512"refId="ref6371"refString="LOGUNOV D. V. 2023. - Further notes on the fossorial wolf spiders of Middle Asia and the Near East (Aranei: Lycosidae). Arthropoda Selecta 32 (4): 475 - 512. https: // doi. 10.15298 / arthsel. 32.4.12"type="journal article"year="2023">Logunov 2023</bibRefCitation>
) have led to the current assignment of two species to the genus
<bibRefCitationid="EFD014D39821FFC4FCA776A6FBF237D0"author="TONGIORGI P."box="[854,1024,1760,1784]"pageId="4"pageNumber="619"pagination="255 - 264"refId="ref7385"refString="TONGIORGI P. 1969. - Vesubia jugorum (Simon) un ragno licoside endemico delle Alpi marittime. Atti della Societa Toscana di Scienze Naturali B 75: 255 - 264."type="journal article"year="1969">Tongiorgi (1969)</bibRefCitation>
for the first time. Tongiorgi provided observational data about the natural history of the species and discussed several hypotheses about its evolutive origins. Additionally, he provided a diagnosis for the genus based on a combination of somatic morphology characters and concluded that
. However, no formal nomenclatural act was proposed. Recently,
<bibRefCitationid="EFD014D39821FFC4FBEF77DCFB4D369A"author="LOGUNOV D. V."box="[1054,1215,1946,1970]"pageId="4"pageNumber="619"pagination="475 - 512"refId="ref6371"refString="LOGUNOV D. V. 2023. - Further notes on the fossorial wolf spiders of Middle Asia and the Near East (Aranei: Lycosidae). Arthropoda Selecta 32 (4): 475 - 512. https: // doi. 10.15298 / arthsel. 32.4.12"type="journal article"year="2023">Logunov (2023)</bibRefCitation>
genera by the combination of the following characters: presence of two retromarginal teeth, a posterior epigynal pocket, a wide embolus, an apical projection of the palea and the shape of the median apophysis, with a dorsal concavity but without a dorsal channel as in other
<bibRefCitationid="EFD014D39820FFC5FEE4711AFE48305C"author="KRONESTEDT T."box="[277,442,348,372]"pageId="5"pageNumber="620"pagination="203 - 225"refId="ref6301"refString="KRONESTEDT T. 1990. - Separation of two species standing as Alopecosa aculeata (Clerck) by morphological, behavioural and ecological characters, with remarks on related species in the pulverulenta group (Araneae, Lycosidae). Zoologica Scripta 19 (2): 203 - 225. https: // doi. org / 10.1111 / j. 1463 - 6409.1990. tb 00256. x"type="journal article"year="1990">Kronestedt 1990</bibRefCitation>
<figureCitationid="137A75A79820FFC5FE877250FE433318"box="[374,433,534,560]"captionStart="FIG"captionStartId="4.[133,144,713,731]"captionTargetBox="[132,1456,213,670]"captionTargetId="figure-724@4.[132,1455,214,670]"captionTargetPageId="4"captionText="FIG. 1. — Live specimens of Vesubia jugorum (Simon, 1881): A, female carrying spiderlings, in the nearby of Refugio Remondino, Valdieri (Alpi Marittime, Prov- ince of Cuneo, Italy); B, female and habitat, Passo Tesina, Sant’Anna di Vinadio (Alpi Marittime, Province of Cuneo, Italy). Photos: A, Francesco Tomasinelli; B, Emanuele Biggi."figureDoi="http://doi.org/10.5281/zenodo.13941218"httpUri="https://zenodo.org/record/13941218/files/figure.png"pageId="5"pageNumber="620">Fig. 1</figureCitation>
). Sternum, labium and maxillae reddish brown. Chelicerae dark brown with three promarginal and two retromarginal teeth. Abdomen uniform dark brown in fixed specimens, in live specimens a distinguishable cardiac mark followed by a series of paler marks (
<figureCitationid="137A75A79820FFC5FD5272D0FD043398"box="[675,758,662,688]"captionStart="FIG"captionStartId="4.[133,144,713,731]"captionTargetBox="[132,1456,213,670]"captionTargetId="figure-724@4.[132,1455,214,670]"captionTargetPageId="4"captionText="FIG. 1. — Live specimens of Vesubia jugorum (Simon, 1881): A, female carrying spiderlings, in the nearby of Refugio Remondino, Valdieri (Alpi Marittime, Prov- ince of Cuneo, Italy); B, female and habitat, Passo Tesina, Sant’Anna di Vinadio (Alpi Marittime, Province of Cuneo, Italy). Photos: A, Francesco Tomasinelli; B, Emanuele Biggi."figureDoi="http://doi.org/10.5281/zenodo.13941218"httpUri="https://zenodo.org/record/13941218/files/figure.png"pageId="5"pageNumber="620">Fig. 1B</figureCitation>
<figureCitationid="137A75A79820FFC5FE117293FDC333D8"box="[480,561,725,752]"captionStart="FIG"captionStartId="4.[133,144,713,731]"captionTargetBox="[132,1456,213,670]"captionTargetId="figure-724@4.[132,1455,214,670]"captionTargetPageId="4"captionText="FIG. 1. — Live specimens of Vesubia jugorum (Simon, 1881): A, female carrying spiderlings, in the nearby of Refugio Remondino, Valdieri (Alpi Marittime, Prov- ince of Cuneo, Italy); B, female and habitat, Passo Tesina, Sant’Anna di Vinadio (Alpi Marittime, Province of Cuneo, Italy). Photos: A, Francesco Tomasinelli; B, Emanuele Biggi."figureDoi="http://doi.org/10.5281/zenodo.13941218"httpUri="https://zenodo.org/record/13941218/files/figure.png"pageId="5"pageNumber="620">Fig. 1B</figureCitation>
<figureCitationid="137A75A79820FFC5FD297472FD0E3566"box="[728,764,1076,1102]"captionStart="FIG"captionStartId="6.[132,143,1981,1999]"captionTargetBox="[132,1455,1301,1936]"captionTargetId="figure-31@6.[132,1455,1301,1936]"captionTargetPageId="6"captionText="FIG. 3. — Type material of Vesubia jugorum (Simon, 1881) from Simon’s collection (MNHN): A, ventral view of the lectotype with its original label; B, ventral view of the epigyne from additional material accompanied by the original label. Not to scale."figureDoi="http://doi.org/10.5281/zenodo.13941222"httpUri="https://zenodo.org/record/13941222/files/figure.png"pageId="5"pageNumber="620">3A</figureCitation>
