treatments-xml/data/D3/2F/57/D32F57BDA41F5A6D96915338F8693DCC.xml

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<mods:title id="508116626BD73B0D43AC36A2480294EA">Diversity underfoot of agromyzids (Agromyzidae, Diptera) mining thalli of liverworts and hornworts</mods:title>
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<mods:namePart id="25C96E6EA820A5EEAF5BBFAC260E0173">Kato, Makoto</mods:namePart>
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<mods:namePart id="D53531FE4AA588C6DCE8BF8FFBBA21C0">Yamamori, Luna</mods:namePart>
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<mods:namePart id="AB2D387D8E58C3ED84E7559A8ADA49E7">Imada, Yume</mods:namePart>
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<treatment id="D32F57BDA41F5A6D96915338F8693DCC" ID-GBIF-Taxon="205344542" LSID="urn:lsid:zoobank.org:act:C62852AE-213D-4143-AB0A-E8472CFFD2AC" httpUri="http://treatment.plazi.org/id/D32F57BDA41F5A6D96915338F8693DCC" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection id="FFA6F6465156561EABE94CD104DAED40" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="1D77DA8ABC3075848675F36C04528002" pageId="0" pageNumber="1">
15. 
<taxonomicName id="42CEB06BD658CE543D0D800566CEE3F9" ID-CoL="BDJLH" LSID="https://zoobank.org/C62852AE-213D-4143-AB0A-E8472CFFD2AC" authority="Kato" authorityName="Kato" authorityYear="2022" class="Insecta" family="Agromyzidae" genus="Phytoliriomyza" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Phytoliriomyza falcata" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="falcata" status="sp. nov.">Phytoliriomyza falcata Kato</taxonomicName>
<taxonomicNameLabel id="06B3BA79D99E15A8700AE304C1EC695A" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="0BD901A287D148BE9D7BFA89AD9DBAA0" pageId="0" pageNumber="1" type="description">
<paragraph id="8425CA0502C8797F36C05A804D25588B" pageId="0" pageNumber="1">
<figureCitation id="016EE8F2B8B99232E27CE82A2D453DEC" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Phytoliriomyza falcata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 18) dorsal G-L male genitalia (G-K, at type locality L at Tatsuyama) G whole genitalia, ventral H phallic complex, ventral I, J, L epandrium (I anterior J anterior-ventral L ventral) K ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure29" httpUri="https://binary.pensoft.net/fig/775402" pageId="0" pageNumber="1">Figs 29</figureCitation>
<figureCitation id="55F8BAE3FEA2FCA37CFAEB059B70B64A" captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Female morphology and larval ecology of Phytoliriomyza falcata sp. nov. A, B female postabdomen A oviscape and spermatheca B tergite 10 C puparium D-F mined thalli of Reboulia hemisphaerica orientalis (D, E at type locality F Yasui-keikoku), arrows indicating puparia." figureDoi="10.3897/zookeys.1133.94530.figure30" httpUri="https://binary.pensoft.net/fig/775403" pageId="0" pageNumber="1">, 30</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="7E0BB142074D960E9121EB2F0F17392B" pageId="0" pageNumber="1" type="material examined">
<paragraph id="287A00810967CC09889FF8DCB3092384" pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph id="024953493257C572EF48DB5EBB2DCF4C" pageId="0" pageNumber="1">
<emphasis id="18C7AEB67B3FC71C26F59B92A3400F37" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
<emphasis id="487AD27C49D88F40C079FFFE48FEE237" bold="true" pageId="0" pageNumber="1">:</emphasis>
Japan: 1♂ (MK-AG-a19), Kanna-gawa, Nakatsugawa, Chichibu, Saitama Pref. (
<geoCoordinate id="1D1ABA0A8AD1B831F0160DBEF375C425" degrees="36.0044" direction="north" orientation="latitude" precision="5" value="36.0044">36.0044°N</geoCoordinate>
, 
<geoCoordinate id="77F5D892C0C26764FE3AE6768B1BE93E" degrees="138.8108" direction="east" orientation="longitude" precision="5" value="138.8108">138.8108°E</geoCoordinate>
, 760 m asl), 14-XI-2010 (as larva), emerged on 23-IV-2011, NSMT-I-Dip 31972. 
<emphasis id="3296D55889C160707EE658781B73CD20" bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
<emphasis id="620679F4663762F4B712C60157021ED3" bold="true" pageId="0" pageNumber="1">:</emphasis>
Japan: 1♂2♀ (MK-AG-423, 807, a18), same data as holotype, emerged on 28-23-IV-2011, NSMT-I-Dip 31973-31975; 1♀ (MK-AG-a376), Ukawa, Tango, Kyotango, Kyoto Pref., 5-III-2021 (as larva), emerged on 12-IV-2021, NSMT-I-Dip 31976; 1♂ (MK-AG-a287), Seya-gawa, Miyazu, Kyoto Pref., 18-IV-2013 (as larva), emerged on 25-IV-2013, NSMT-I-Dip 31977; 1♂ (MK-AG-803), Ryugakyo, Yamashiro, Miyoshi, Tokushima Pref., 1-II-2014 (as larva), emerged on 25-IV-2014, NSMT-I-Dip 31978; 1♀ (MK-AG-444), Yasui-keikoku, Niyodogawa, Agawa, Kochi Pref., 27-II-2011 (as larva), emerged on 28-IV-2011, NSMT-I-Dip 31979; 1♀ (MK-AG-795), Chiromo, Toyotama, Tsushima, Nagasaki Pref., 11-XI-2011 (as larva), emerged on 28-IV-2012, NSMT-I-Dip 31980.
</paragraph>
</subSubSection>
<subSubSection id="5D1C1A57D6CF47F57EBEE020422F6E07" pageId="0" pageNumber="1" type="other material">
<paragraph id="0E75626DD95A268ACA642373FBE6769C" pageId="0" pageNumber="1">Other material.</paragraph>
<paragraph id="0D73E04654D29F72E511059A01E3479F" pageId="0" pageNumber="1">Japan: 17♂22♀, Kanna-gawa, Nakatsugawa, Chichibu, Saitama Pref., 14-XI-2010 (as larva), emerged on 14-IV-2011; 6♂11♀, Oochi-gawa, Chichibu, Saitama Pref., 13-III-2017 (as larva), emerged on 25-28-IV-2011; 1♂2♀, Ukawa, Tango, Kyotango, Kyoto Pref., 5-III-2021 (as larva), emerged on 9-17-IV-2021; 1♀, Doro-kyo, Totsugawa-mura, Nara Pref., 29-III-2019 (as larva), emerged on 3-IV-2019; 1♂4♀, Tazukawa-keikoku, Katsuura, Tokushima Pref., 30-III-2021 (as larva), emerged on 23-30-IV-2021; 2♂1♀, Kurase-keikoku, Tanbara, Saijo, Ehime Pref., 2-II-2014 (as larva), emerged on 20-26-IV-2014.</paragraph>
</subSubSection>
<subSubSection id="DC78197429E067E48CDC3BB6F58002F1" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="C76AE2A3F97BE7096F092CB84D33CCC2" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="3E10D72A733781967CAE33F5FF633DE3" pageId="0" pageNumber="1">
A medium-sized yellow species (wing length 1.6-2.0 mm) having subshiny brown scutum with an oval yellow pattern extending from the mid-posterior margin to the scutellum, a yellow 1st flagellomere, yellow maxillary palpus, yellow halteres, and yellow legs. Male epandrium inner-laterally with a long hypertrophied arm which apically bears a dark, long, apically flattened, obliquely truncated tubercle-like seta. Larva mines the thallus of 
<taxonomicName id="30757AC2BD1D2962587132DE1D0FD828" class="Marchantiopsida" family="Aytoniaceae" genus="Reboulia" kingdom="Plantae" lsidName="Reboulia hemisphaerica subsp. orientalis" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="subSpecies" species="hemisphaerica" subSpecies="orientalis">
<emphasis id="68EE741E5C823E6CE61A9E9400758FDF" italics="true" pageId="0" pageNumber="1">Reboulia hemisphaerica orientalis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="9CFF86F2F0D090B1D345055AC2C18F93" pageId="0" pageNumber="1" type="description">
<paragraph id="4082F82E5A5ADFF94FC5A595347F78F7" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="14C6CEBF5850ACC6693FD91858E283D9" pageId="0" pageNumber="1">
<emphasis id="905B4C441A4B4DE801EE4C3317FD9B02" bold="true" pageId="0" pageNumber="1">Adult male</emphasis>
(Fig. 
<figureCitation id="71932F78A71F316BEB67BD63E168324C" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Phytoliriomyza falcata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 18) dorsal G-L male genitalia (G-K, at type locality L at Tatsuyama) G whole genitalia, ventral H phallic complex, ventral I, J, L epandrium (I anterior J anterior-ventral L ventral) K ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure29" httpUri="https://binary.pensoft.net/fig/775402" pageId="0" pageNumber="1">29A-E</figureCitation>
).
</paragraph>
<paragraph id="5A2FE5E9EFE638F62EE71314A7857D43" pageId="0" pageNumber="1">
<emphasis id="78A539025F42AB732352FA71981D6921" bold="true" italics="true" pageId="0" pageNumber="1">Head</emphasis>
<emphasis id="6D474837D91D6EE1BA393EB2EFE5BAF7" bold="true" pageId="0" pageNumber="1">:</emphasis>
Head entirely yellow, with ocellar tubercle brown, and back of head dark brown (Fig. 
<figureCitation id="3A6143157645E53F82A7A36ED703F93F" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Phytoliriomyza falcata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 18) dorsal G-L male genitalia (G-K, at type locality L at Tatsuyama) G whole genitalia, ventral H phallic complex, ventral I, J, L epandrium (I anterior J anterior-ventral L ventral) K ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure29" httpUri="https://binary.pensoft.net/fig/775402" pageId="0" pageNumber="1">29C</figureCitation>
). Antenna porrect, first flagellomere black, pedicel yellow and scape light yellow. Arista subbasal, brown, pubescent. Frons with brownish pruinosity. Face, gena, parafacial and postgena yellow. Proboscis normal, yellow; palpus yellow, cylindrical (Fig. 
<figureCitation id="07FA2DE7D33906EED13AED2759670C6B" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Phytoliriomyza falcata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 18) dorsal G-L male genitalia (G-K, at type locality L at Tatsuyama) G whole genitalia, ventral H phallic complex, ventral I, J, L epandrium (I anterior J anterior-ventral L ventral) K ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure29" httpUri="https://binary.pensoft.net/fig/775402" pageId="0" pageNumber="1">29C</figureCitation>
). 
<emphasis id="48E4B6472AD17E85F02A029BB04D0976" bold="true" italics="true" pageId="0" pageNumber="1">Chaetotaxy</emphasis>
<emphasis id="F9CF04247C47061F96D3E66A7040CB78" bold="true" pageId="0" pageNumber="1">:</emphasis>
Front orbitals three pairs; one ori directed inward; two ors directed upward (Fig. 
<figureCitation id="087A900F68BCC5E027E25DAD70EE1B27" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Phytoliriomyza falcata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 18) dorsal G-L male genitalia (G-K, at type locality L at Tatsuyama) G whole genitalia, ventral H phallic complex, ventral I, J, L epandrium (I anterior J anterior-ventral L ventral) K ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure29" httpUri="https://binary.pensoft.net/fig/775402" pageId="0" pageNumber="1">29B</figureCitation>
). Orbital setulae minute and erect, in a single row.
</paragraph>
<caption id="D1DF399974162B20456369E0E4D71851" doi="10.3897/zookeys.1133.94530.figure29" httpUri="https://binary.pensoft.net/fig/775402" pageId="0" pageNumber="1" start="Figure 29" startId="F29">
<paragraph id="1E1627C59EF4BEC25FE4A258C0B7C574" pageId="0" pageNumber="1">
<emphasis id="7E8AAACAB572C12149F2FCB0303A6C4E" bold="true" pageId="0" pageNumber="1">Figure 29.</emphasis>
<taxonomicName id="075872AC5D5B5794E117765D9A2E74EF" authorityName="Kato" authorityYear="2022" class="Insecta" family="Dendrocerotaceae" genus="Phytoliriomyza" kingdom="Animalia" lsidName="Phytoliriomyza falcata" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="falcata">
<emphasis id="010E0C1A30473982B23585571B83676C" italics="true" pageId="0" pageNumber="1">Phytoliriomyza falcata</emphasis>
</taxonomicName>
sp. nov. 
<emphasis id="12712E38F1758A78D95EC710A26820FF" bold="true" pageId="0" pageNumber="1">A-E</emphasis>
holotype male 
<emphasis id="04154FDE59800F9C588CEEC449DEE5FE" bold="true" pageId="0" pageNumber="1">A</emphasis>
habitus 
<emphasis id="C3E458CACF170DCD1CD3FFA36B8A7B67" bold="true" pageId="0" pageNumber="1">B</emphasis>
lateral 
<emphasis id="53F672E2B34F49A13BB39DAF9F4FD5F1" bold="true" pageId="0" pageNumber="1">C</emphasis>
frontal 
<emphasis id="A4C6A8BA632D5EB791FD8D08AC1AB752" bold="true" pageId="0" pageNumber="1">D</emphasis>
dorsal 
<emphasis id="0ED2E6AAF9B8CF2DE699B3805467E04D" bold="true" pageId="0" pageNumber="1">E</emphasis>
posterior 
<emphasis id="4DD44183F891997A12D74650143C5E10" bold="true" pageId="0" pageNumber="1">F</emphasis>
paratype female (MK-AG-a18) dorsal 
<emphasis id="3EA896E88EC8440E28E1270D18EA366A" bold="true" pageId="0" pageNumber="1">G-L</emphasis>
male genitalia (
<emphasis id="EA374BA7C0CB18682E64E768BA17E48A" bold="true" pageId="0" pageNumber="1">G-K</emphasis>
, at type locality 
<emphasis id="990764A167A0196D069BA75462E6CE62" bold="true" pageId="0" pageNumber="1">L</emphasis>
at Tatsuyama) 
<emphasis id="C4437712D8D41B3A94400596293818D8" bold="true" pageId="0" pageNumber="1">G</emphasis>
whole genitalia, ventral 
<emphasis id="BA59F5520BC844BBF10AA6E2485A0757" bold="true" pageId="0" pageNumber="1">H</emphasis>
phallic complex, ventral 
<emphasis id="B6882D4D3C462DC5D5D16C0553AB617C" bold="true" pageId="0" pageNumber="1">I, J, L</emphasis>
epandrium (
<emphasis id="FBF7E6665D84BE76F3724BBF3D002FDE" bold="true" pageId="0" pageNumber="1">I</emphasis>
anterior 
<emphasis id="3E30EC6F2C07EF190A05E94F39CB7F3E" bold="true" pageId="0" pageNumber="1">J</emphasis>
anterior-ventral 
<emphasis id="4217FDEA500D89DB010C6CD8295CBF91" bold="true" pageId="0" pageNumber="1">L</emphasis>
ventral) 
<emphasis id="2BB983D2CD611A0120A7D70F5DBBC2B2" bold="true" pageId="0" pageNumber="1">K</emphasis>
ejaculatory apodeme, lateral.
</paragraph>
</caption>
<paragraph id="5E8D3183F0CD4678D0F905F38FED54BF" pageId="0" pageNumber="1">
<emphasis id="2EFF2B920691EFE8F243E47057A1E658" bold="true" italics="true" pageId="0" pageNumber="1">Thorax</emphasis>
<emphasis id="BB9AAFC6DDCE672559A8ECC71F8CFE6C" bold="true" pageId="0" pageNumber="1">:</emphasis>
Thorax subshiny, with a brown medial stripe on anterior 2/3, and a pair of adjacent wide brown bands on anterior 7/8 (Fig. 
<figureCitation id="9034BF37D96795DD1A82C4C7027FE1A1" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Phytoliriomyza falcata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 18) dorsal G-L male genitalia (G-K, at type locality L at Tatsuyama) G whole genitalia, ventral H phallic complex, ventral I, J, L epandrium (I anterior J anterior-ventral L ventral) K ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure29" httpUri="https://binary.pensoft.net/fig/775402" pageId="0" pageNumber="1">29D</figureCitation>
). Scutellum and subscutellum yellow. Mediotergite brown, anatergite and katatergite yellow. Pleuron largely yellow; propleuron with small brown patch on mid-anterior corner; anepisternum with two small spots on anterior and posterior corners; anepimeron with an oblique narrow brown patch on anterior corner; katepisternum and meron with brown patches on venter (Fig. 
<figureCitation id="02932888DFBCBDA3E3F1E331E20D2F52" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Phytoliriomyza falcata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 18) dorsal G-L male genitalia (G-K, at type locality L at Tatsuyama) G whole genitalia, ventral H phallic complex, ventral I, J, L epandrium (I anterior J anterior-ventral L ventral) K ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure29" httpUri="https://binary.pensoft.net/fig/775402" pageId="0" pageNumber="1">29B</figureCitation>
). Haltere yellow. Calypter margin and hairs gray. Leg segments yellow; tibia and tarsus darker (Fig. 
<figureCitation id="8D829D123F32F1FC9B0B1CFA0F6790DA" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Phytoliriomyza falcata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 18) dorsal G-L male genitalia (G-K, at type locality L at Tatsuyama) G whole genitalia, ventral H phallic complex, ventral I, J, L epandrium (I anterior J anterior-ventral L ventral) K ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure29" httpUri="https://binary.pensoft.net/fig/775402" pageId="0" pageNumber="1">29B</figureCitation>
). 
<emphasis id="FE67F7B6525C50F6E42F1C2D748D1BA5" bold="true" italics="true" pageId="0" pageNumber="1">Chaetotaxy</emphasis>
<emphasis id="CE6805833967DCCEF79D7E45BE4F3AF2" bold="true" pageId="0" pageNumber="1">:</emphasis>
Scutum with 1+3 dorsocentrals, shortened anteriorly (Fig. 
<figureCitation id="F7F21E67CE22740FF5A8EDF7B8E43A76" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Phytoliriomyza falcata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 18) dorsal G-L male genitalia (G-K, at type locality L at Tatsuyama) G whole genitalia, ventral H phallic complex, ventral I, J, L epandrium (I anterior J anterior-ventral L ventral) K ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure29" httpUri="https://binary.pensoft.net/fig/775402" pageId="0" pageNumber="1">29D</figureCitation>
). Acrostichal setulae 6-8 pairs in two rows. 
<emphasis id="DE6374BE712380D3476D62B4AB278A0E" bold="true" italics="true" pageId="0" pageNumber="1">Wing</emphasis>
: Wing length 1.6-1.9 mm, costa reaching M1 (Fig. 
<figureCitation id="97AD47EB64AA2EDE15262F0D7CE64619" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Phytoliriomyza falcata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 18) dorsal G-L male genitalia (G-K, at type locality L at Tatsuyama) G whole genitalia, ventral H phallic complex, ventral I, J, L epandrium (I anterior J anterior-ventral L ventral) K ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure29" httpUri="https://binary.pensoft.net/fig/775402" pageId="0" pageNumber="1">29A</figureCitation>
). Length of ultimate section of vein M4 divided by penultimate section 1.5-1.8.
</paragraph>
<paragraph id="E5A4991B0F4D44F8CA54C29C58F238F8" pageId="0" pageNumber="1">
<emphasis id="9871B01924A7629B5E9CF0F3907EA319" bold="true" italics="true" pageId="0" pageNumber="1">Abdomen</emphasis>
<emphasis id="AE60FE75E3834393A67484C6EBD1AF46" bold="true" pageId="0" pageNumber="1">:</emphasis>
Abdomen dorsally subshiny yellow (Fig. 
<figureCitation id="FF6CF51629816A622FAC01BC8C838BA1" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Phytoliriomyza falcata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 18) dorsal G-L male genitalia (G-K, at type locality L at Tatsuyama) G whole genitalia, ventral H phallic complex, ventral I, J, L epandrium (I anterior J anterior-ventral L ventral) K ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure29" httpUri="https://binary.pensoft.net/fig/775402" pageId="0" pageNumber="1">29E</figureCitation>
). 
<emphasis id="11AD71B51C08928D2E41105B2941D3F2" bold="true" italics="true" pageId="0" pageNumber="1">Genitalia</emphasis>
<emphasis id="9D63582FE77E03622F34DABC95E2A517" bold="true" pageId="0" pageNumber="1">:</emphasis>
(Fig. 
<figureCitation id="EF5031045A51922CA2CB7EEE4534F8D9" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Phytoliriomyza falcata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 18) dorsal G-L male genitalia (G-K, at type locality L at Tatsuyama) G whole genitalia, ventral H phallic complex, ventral I, J, L epandrium (I anterior J anterior-ventral L ventral) K ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure29" httpUri="https://binary.pensoft.net/fig/775402" pageId="0" pageNumber="1">29G-L</figureCitation>
) Epandrium dark brown, rounded apically; inner-anterior margin with two short tubercle-like setae; inner-lateral surface with a basally enlarged, extremely extended, ventrally curved arm, bearing an apically flattened, obliquely truncated, tubercle-like seta borne (Fig. 
<figureCitation id="02083FC6482CE429DE61C9E29773F089" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Phytoliriomyza falcata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 18) dorsal G-L male genitalia (G-K, at type locality L at Tatsuyama) G whole genitalia, ventral H phallic complex, ventral I, J, L epandrium (I anterior J anterior-ventral L ventral) K ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure29" httpUri="https://binary.pensoft.net/fig/775402" pageId="0" pageNumber="1">29I, J, L</figureCitation>
). Surstylus long, narrow, setose apically. Cercus narrow, setose. Subepandrial sclerite consisting of a pair of dorsal and ventral arms; ventral arm narrow, extended, plate-like, with hooked apex and a long basal seta; dorsal arm also plate-like but shorter, with a seta basally (Fig. 
<figureCitation id="CA85E7BBA9ECCE379E9C4D1D3083596C" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Phytoliriomyza falcata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 18) dorsal G-L male genitalia (G-K, at type locality L at Tatsuyama) G whole genitalia, ventral H phallic complex, ventral I, J, L epandrium (I anterior J anterior-ventral L ventral) K ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure29" httpUri="https://binary.pensoft.net/fig/775402" pageId="0" pageNumber="1">29I</figureCitation>
). Hypandrium thin, slightly sclerotized along outer margin (Fig. 
<figureCitation id="2F9C2C946D6232D149920F7EAEFE3DCC" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Phytoliriomyza falcata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 18) dorsal G-L male genitalia (G-K, at type locality L at Tatsuyama) G whole genitalia, ventral H phallic complex, ventral I, J, L epandrium (I anterior J anterior-ventral L ventral) K ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure29" httpUri="https://binary.pensoft.net/fig/775402" pageId="0" pageNumber="1">29G</figureCitation>
). Postgonite bare and goose barnacle-shaped, with sideward pointed apex (Fig. 
<figureCitation id="2EE71AAE1CF8E005213C25FD34D6D3F3" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Phytoliriomyza falcata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 18) dorsal G-L male genitalia (G-K, at type locality L at Tatsuyama) G whole genitalia, ventral H phallic complex, ventral I, J, L epandrium (I anterior J anterior-ventral L ventral) K ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure29" httpUri="https://binary.pensoft.net/fig/775402" pageId="0" pageNumber="1">29H</figureCitation>
). Phallophorus with deep incision below, articulated with phallapodeme, fused to epiphallus (Fig. 
<figureCitation id="73E20F5535E4821459BACDBBAD342F8E" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Phytoliriomyza falcata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 18) dorsal G-L male genitalia (G-K, at type locality L at Tatsuyama) G whole genitalia, ventral H phallic complex, ventral I, J, L epandrium (I anterior J anterior-ventral L ventral) K ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure29" httpUri="https://binary.pensoft.net/fig/775402" pageId="0" pageNumber="1">29H</figureCitation>
). Basiphallus with a pair of expanded lightly sclerotized lateral plates. Hypophallus broad, membranous, and bilaterally asymmetrical; right margin sclerotized; medially with a pair of dark fused sclerites (Fig. 
<figureCitation id="99C4569C680497ADFBA603FA53554E68" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Phytoliriomyza falcata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 18) dorsal G-L male genitalia (G-K, at type locality L at Tatsuyama) G whole genitalia, ventral H phallic complex, ventral I, J, L epandrium (I anterior J anterior-ventral L ventral) K ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure29" httpUri="https://binary.pensoft.net/fig/775402" pageId="0" pageNumber="1">29H</figureCitation>
). Paraphalli lobate, lightly sclerotized; diverging and angled anteroventrally (Fig. 
<figureCitation id="0A09F4A075D84EDB87F1BD0EED7277C7" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Phytoliriomyza falcata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 18) dorsal G-L male genitalia (G-K, at type locality L at Tatsuyama) G whole genitalia, ventral H phallic complex, ventral I, J, L epandrium (I anterior J anterior-ventral L ventral) K ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure29" httpUri="https://binary.pensoft.net/fig/775402" pageId="0" pageNumber="1">29H</figureCitation>
). Mesophallus dark, cylindrical, widest subbasally, as long as distiphallus (Fig. 
<figureCitation id="0AB398490E4B3413C7FF4760C5F4CBEC" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Phytoliriomyza falcata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 18) dorsal G-L male genitalia (G-K, at type locality L at Tatsuyama) G whole genitalia, ventral H phallic complex, ventral I, J, L epandrium (I anterior J anterior-ventral L ventral) K ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure29" httpUri="https://binary.pensoft.net/fig/775402" pageId="0" pageNumber="1">29H</figureCitation>
). Distiphallus comprising one pair of stout tubules; basal half composed of lateral dark slender sclerite and weaker medial region; covered by membrane bearing four pairs of minute arrowhead-like lateral sclerites; distal half cylindrical, dorsally and laterally pigmented, with truncated, shortly flared apex (Fig. 
<figureCitation id="18A0A5D5C51CA6F912051C04AC8B4F77" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Phytoliriomyza falcata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 18) dorsal G-L male genitalia (G-K, at type locality L at Tatsuyama) G whole genitalia, ventral H phallic complex, ventral I, J, L epandrium (I anterior J anterior-ventral L ventral) K ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure29" httpUri="https://binary.pensoft.net/fig/775402" pageId="0" pageNumber="1">29H</figureCitation>
). Ejaculatory apodeme pale and fan-shaped with broad stalk; base wide to one side; sperm pump clear (Fig. 
<figureCitation id="1B98FFBF422E1AC08ED65A8E2FEAB4B6" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Phytoliriomyza falcata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 18) dorsal G-L male genitalia (G-K, at type locality L at Tatsuyama) G whole genitalia, ventral H phallic complex, ventral I, J, L epandrium (I anterior J anterior-ventral L ventral) K ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure29" httpUri="https://binary.pensoft.net/fig/775402" pageId="0" pageNumber="1">29K</figureCitation>
).
</paragraph>
<paragraph id="5AE0B4E0DB174253BBA77215D034A311" pageId="0" pageNumber="1">
<emphasis id="5A7602C60D9B6A95777705A62162432F" bold="true" pageId="0" pageNumber="1">Female</emphasis>
(Fig. 
<figureCitation id="F24C2BB1BD006F6A6924EE7CB4326480" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Phytoliriomyza falcata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 18) dorsal G-L male genitalia (G-K, at type locality L at Tatsuyama) G whole genitalia, ventral H phallic complex, ventral I, J, L epandrium (I anterior J anterior-ventral L ventral) K ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure29" httpUri="https://binary.pensoft.net/fig/775402" pageId="0" pageNumber="1">29F</figureCitation>
). Similar to male, but larger, the lateral bands on scutum more grayish and more pruinose. Wing length 1.8-2.0 mm. 
<emphasis id="021A77A6A767EA6870515ACF235DA730" bold="true" italics="true" pageId="0" pageNumber="1">Postabdomen</emphasis>
<emphasis id="6E9411281AA41482FC6E63EBC0C6B05A" bold="true" pageId="0" pageNumber="1">:</emphasis>
(Fig. 
<figureCitation id="D22C6FE3BD2944C63F666332595CCFAC" captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Female morphology and larval ecology of Phytoliriomyza falcata sp. nov. A, B female postabdomen A oviscape and spermatheca B tergite 10 C puparium D-F mined thalli of Reboulia hemisphaerica orientalis (D, E at type locality F Yasui-keikoku), arrows indicating puparia." figureDoi="10.3897/zookeys.1133.94530.figure30" httpUri="https://binary.pensoft.net/fig/775403" pageId="0" pageNumber="1">30A, B</figureCitation>
) Oviscape dark brown, setigerous (Fig. 
<figureCitation id="EA8C24E7A4E640049301FADB41BF10A4" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Phytoliriomyza falcata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 18) dorsal G-L male genitalia (G-K, at type locality L at Tatsuyama) G whole genitalia, ventral H phallic complex, ventral I, J, L epandrium (I anterior J anterior-ventral L ventral) K ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure29" httpUri="https://binary.pensoft.net/fig/775402" pageId="0" pageNumber="1">29A</figureCitation>
). Tergite 10 trifurcate, laterally uniting narrow pleural sclerites (Fig. 
<figureCitation id="7443FD73B189AF59EE8752920A98FDF8" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Phytoliriomyza falcata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 18) dorsal G-L male genitalia (G-K, at type locality L at Tatsuyama) G whole genitalia, ventral H phallic complex, ventral I, J, L epandrium (I anterior J anterior-ventral L ventral) K ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure29" httpUri="https://binary.pensoft.net/fig/775402" pageId="0" pageNumber="1">29B</figureCitation>
). Each cercus with two stout, apical, trichoid sensilla, 
<normalizedToken id="E0C47FD2E5317048439592A8BA839A99" originalValue="½">1/2</normalizedToken>
length of cercus (Fig. 
<figureCitation id="B264D4D37077292C3E0975480E1E87FA" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Phytoliriomyza falcata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 18) dorsal G-L male genitalia (G-K, at type locality L at Tatsuyama) G whole genitalia, ventral H phallic complex, ventral I, J, L epandrium (I anterior J anterior-ventral L ventral) K ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure29" httpUri="https://binary.pensoft.net/fig/775402" pageId="0" pageNumber="1">29B</figureCitation>
). Spermathecae semi-orbicular, with truncate proximal ends (Fig. 
<figureCitation id="2C5A5C820BC7611526BD6836EE84DECA" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Phytoliriomyza falcata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 18) dorsal G-L male genitalia (G-K, at type locality L at Tatsuyama) G whole genitalia, ventral H phallic complex, ventral I, J, L epandrium (I anterior J anterior-ventral L ventral) K ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure29" httpUri="https://binary.pensoft.net/fig/775402" pageId="0" pageNumber="1">29A</figureCitation>
).
</paragraph>
<caption id="952DB15A2FE56232CDA37356BA21D974" doi="10.3897/zookeys.1133.94530.figure30" httpUri="https://binary.pensoft.net/fig/775403" pageId="0" pageNumber="1" start="Figure 30" startId="F30">
<paragraph id="4A34F5088F8C598265060CE6EC208A10" pageId="0" pageNumber="1">
<emphasis id="839246DF5D6130E5F0C42D48904E2F50" bold="true" pageId="0" pageNumber="1">Figure 30.</emphasis>
Female morphology and larval ecology of 
<taxonomicName id="D038E249AA4F907F3536102D339B81EA" authorityName="Kato" authorityYear="2022" class="Insecta" family="Dendrocerotaceae" genus="Phytoliriomyza" kingdom="Animalia" lsidName="Phytoliriomyza falcata" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="falcata">
<emphasis id="C0611C57C0C9EBEFBAF8684EF7339575" italics="true" pageId="0" pageNumber="1">Phytoliriomyza falcata</emphasis>
</taxonomicName>
sp. nov. 
<emphasis id="96D8716BCE09305B9EFD38A848CC1F16" bold="true" pageId="0" pageNumber="1">A, B</emphasis>
female postabdomen 
<emphasis id="BC76CA7B499FAA68DD9362E7895E2C5C" bold="true" pageId="0" pageNumber="1">A</emphasis>
oviscape and spermatheca 
<emphasis id="1DCAB1198AD80F019CDDCA1D1BE8F374" bold="true" pageId="0" pageNumber="1">B</emphasis>
tergite 10 
<emphasis id="7F67E7EBD0615E8653170B37C5812534" bold="true" pageId="0" pageNumber="1">C</emphasis>
puparium 
<emphasis id="D3F7BC9F64766BB39F03C78A585469E6" bold="true" pageId="0" pageNumber="1">D-F</emphasis>
mined thalli of 
<taxonomicName id="DDD79AD24FD63BA3BD3906970CE67D6D" class="Marchantiopsida" family="Aytoniaceae" genus="Reboulia" kingdom="Plantae" lsidName="Reboulia hemisphaerica subsp. orientalis" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="subSpecies" species="hemisphaerica" subSpecies="orientalis">
<emphasis id="AB5E16CEF3D3E19C32C9553C3BB8BC8B" italics="true" pageId="0" pageNumber="1">Reboulia hemisphaerica orientalis</emphasis>
</taxonomicName>
(
<emphasis id="0815606E1A5535277DCF89AC8817D7AD" bold="true" pageId="0" pageNumber="1">D, E</emphasis>
at type locality 
<emphasis id="4024804BF69D4EC81E2FB2BB28E1C9EE" bold="true" pageId="0" pageNumber="1">F</emphasis>
Yasui-keikoku), arrows indicating puparia.
</paragraph>
</caption>
<paragraph id="5FE4D77115EB8076C2E4B6A1B13A8B1C" pageId="0" pageNumber="1">
<emphasis id="CBE9DF6D29441FEE1B8EAB97CA2FAC16" bold="true" pageId="0" pageNumber="1">Immatures.</emphasis>
(Fig. 
<figureCitation id="8C0A458B34DD8EE0F1B00A384C3FEDB1" captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Female morphology and larval ecology of Phytoliriomyza falcata sp. nov. A, B female postabdomen A oviscape and spermatheca B tergite 10 C puparium D-F mined thalli of Reboulia hemisphaerica orientalis (D, E at type locality F Yasui-keikoku), arrows indicating puparia." figureDoi="10.3897/zookeys.1133.94530.figure30" httpUri="https://binary.pensoft.net/fig/775403" pageId="0" pageNumber="1">30D-F</figureCitation>
) Puparium internal, slender, and pale brown, with anterior spiracles just protruded from epidermis of mined thallus.
</paragraph>
</subSubSection>
<subSubSection id="6AECA2B11EF11B7F1309DD8921BB264F" pageId="0" pageNumber="1" type="etymology">
<paragraph id="8EC4751772AF342087CF232034EFBD18" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="95F33B63B46C54BAEF04620E2FD66DE9" pageId="0" pageNumber="1">
The specific name (
<emphasis id="3B5A31005F7E4F6A2868E3820265A729" italics="true" pageId="0" pageNumber="1">falcata</emphasis>
= sickle-shaped) refers to the sickle-shaped tubercle-like seta on the male epandrium.
</paragraph>
</subSubSection>
<subSubSection id="677593BE896DD36728BCA45CFBC0F921" pageId="0" pageNumber="1" type="japanese name">
<paragraph id="5CA38B0D316A5FFBF24F0B1CC1D7BC77" pageId="0" pageNumber="1">Japanese name.</paragraph>
<paragraph id="5E2CC170762223723115A9E19A384BE9" pageId="0" pageNumber="1">Naginata-jingasagoke-hamoguribae.</paragraph>
</subSubSection>
<subSubSection id="CAE280E74777A00FE6BBD86A4623EB84" pageId="0" pageNumber="1" type="host plant">
<paragraph id="257BAB6DACA75379E87ABE4A2B31BDCA" pageId="0" pageNumber="1">Host plant.</paragraph>
<paragraph id="EDF538E862E3073AAD9779C6A100F5A1" pageId="0" pageNumber="1">
<taxonomicName id="68D44FB01BE6D523445D579B9AEF8E68" class="Marchantiopsida" family="Aytoniaceae" genus="Reboulia" kingdom="Plantae" lsidName="Reboulia hemisphaerica subsp. orientalis" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="subSpecies" species="hemisphaerica" subSpecies="orientalis">
<emphasis id="80A6FBFA6B75E33112D86C0803CEC9F6" italics="true" pageId="0" pageNumber="1">Reboulia hemisphaerica orientalis</emphasis>
</taxonomicName>
(
<taxonomicName id="67CCAA37BF6A4F6BAEABC0C0CE5EA53A" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="family">Aytoniaceae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="279A4F6C8CB08DA1E6F02CE8B3F14A50" pageId="0" pageNumber="1" type="mine">
<paragraph id="F2F033637B85E60A0EE47817E4428F88" pageId="0" pageNumber="1">Mine.</paragraph>
<paragraph id="7EF036D416D4B67DD0672AB0477AC59E" pageId="0" pageNumber="1">
Larvae construct linear-blotch mines in the thallus, and pupate in the mines (Fig. 
<figureCitation id="A5FF6F14B966DBF5BE26172898D309C2" captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Female morphology and larval ecology of Phytoliriomyza falcata sp. nov. A, B female postabdomen A oviscape and spermatheca B tergite 10 C puparium D-F mined thalli of Reboulia hemisphaerica orientalis (D, E at type locality F Yasui-keikoku), arrows indicating puparia." figureDoi="10.3897/zookeys.1133.94530.figure30" httpUri="https://binary.pensoft.net/fig/775403" pageId="0" pageNumber="1">30E-I</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="197EB8C8FF90BECF73B43EE07AD818FE" pageId="0" pageNumber="1" type="biological notes">
<paragraph id="1AB8ADD445F62C9B5CD797B28AFBAA6B" pageId="0" pageNumber="1">Biological notes.</paragraph>
<paragraph id="1F05CD69F4CD0C20DE722AA7C300ACFB" pageId="0" pageNumber="1">
The habitats of this species are rocky cliffs in warm temperate evergreen forests. This species is sympatric with 
<taxonomicName id="AC290B905198562AA81B20E6D2D5F6CE" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. argentifasciata" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="argentifasciata">
<emphasis id="2D4CF34A24368BEF36C927FA95A06D60" italics="true" pageId="0" pageNumber="1">P. argentifasciata</emphasis>
</taxonomicName>
in some localities. Our rearing records suggest that it is bivoltine, and that adults emerge from overwintered pupae in spring.
</paragraph>
</subSubSection>
<subSubSection id="47CC817670438B61CB4A83370CF6DF6F" pageId="0" pageNumber="1" type="distribution">
<paragraph id="E352194E528776D193FFA0764FB5778C" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="30F6C8979981718EC826BE23B502C91C" pageId="0" pageNumber="1">
Japan: Honshu, Shikoku, Tsushima Island (Fig. 
<figureCitation id="EEDD9740054FB08CD4EC6BABC5A25907" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Locality records of three Phytoliriomyza species associated with Reboulia and Wiesnerella spp.: P. falcata, P. aratriformis, P. rebouliae and P. wiesnerellae." figureDoi="10.3897/zookeys.1133.94530.figure31" httpUri="https://binary.pensoft.net/fig/775404" pageId="0" pageNumber="1">31</figureCitation>
).
</paragraph>
<caption id="D4F647A763FD8BE8730BE7CA16992CE9" doi="10.3897/zookeys.1133.94530.figure31" httpUri="https://binary.pensoft.net/fig/775404" pageId="0" pageNumber="1" start="Figure 31" startId="F31">
<paragraph id="614DC9FA912DF6129F6BC1A36D684849" pageId="0" pageNumber="1">
<emphasis id="8C5E1681AB5177A93145EC1D441877D0" bold="true" pageId="0" pageNumber="1">Figure 31.</emphasis>
Locality records of three 
<taxonomicName id="5BD89867E19B6EA4FAF3CBAC86FA7899" authorityName="Hendel" authorityYear="1931" class="Insecta" family="Dendrocerotaceae" genus="Phytoliriomyza" kingdom="Animalia" lsidName="Phytoliriomyza" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="C43BF8B061625257123E5DD9076F00DB" italics="true" pageId="0" pageNumber="1">Phytoliriomyza</emphasis>
</taxonomicName>
species associated with 
<taxonomicName id="AAF48798B96E9E7E4218DDB234A263F4" class="Marchantiopsida" family="Aytoniaceae" genus="Reboulia" kingdom="Plantae" lsidName="Reboulia" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="genus">
<emphasis id="27583201602462D088DAD6D815B3C89F" italics="true" pageId="0" pageNumber="1">Reboulia</emphasis>
</taxonomicName>
and 
<taxonomicName id="BB43B6DAF7B56BCB2174BA8C8034FAC5" authorityName="Cushman" authorityYear="1933" class="Marchantiopsida" family="Wiesnerellaceae" genus="Wiesnerella" kingdom="Plantae" lsidName="Wiesnerella" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="genus">
<emphasis id="C778A7AD7C67A273AEACE78D154044D8" italics="true" pageId="0" pageNumber="1">Wiesnerella</emphasis>
</taxonomicName>
spp.: 
<taxonomicName id="BC00EE4ABB7FB14CE6AB196C06F54C5E" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. falcata" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="falcata">
<emphasis id="AE8B5826BD4CB9009436906436FDDF5A" italics="true" pageId="0" pageNumber="1">P. falcata</emphasis>
</taxonomicName>
, 
<taxonomicName id="98881B874EBD4224BE1DC053EB252D74" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. aratriformis" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="aratriformis">
<emphasis id="28272C6F74499E009C558A5181843581" italics="true" pageId="0" pageNumber="1">P. aratriformis</emphasis>
</taxonomicName>
, 
<taxonomicName id="0D59B31A86A34F87B51C5AA4F45D0C6A" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. rebouliae" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="rebouliae">
<emphasis id="B3693A333910978F5595723F8D6DB30D" italics="true" pageId="0" pageNumber="1">P. rebouliae</emphasis>
</taxonomicName>
and 
<taxonomicName id="AF9EA141BF8E745557F3F2AC84391C81" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. wiesnerellae" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="wiesnerellae">
<emphasis id="58D761C5712FB70F01708AA91F040894" italics="true" pageId="0" pageNumber="1">P. wiesnerellae</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="604E678F7DBB3585C43F51633C46308A" pageId="0" pageNumber="1" type="remarks">
<paragraph id="45F63453D0E8FCA57B4DDF53232A9CF1" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="B57543847197D56BE2DFAB84A5ECA847" pageId="0" pageNumber="1">
This species resembles 
<taxonomicName id="0161E66D94350B3FC134050A6A659F01" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. arcus" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="arcus">
<emphasis id="40F9BBEA511934950B28F1853D7F0E23" italics="true" pageId="0" pageNumber="1">P. arcus</emphasis>
</taxonomicName>
, 
<taxonomicName id="48AED2FA547C0072BB52CB7A78557A69" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. plagiochasmatos" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="plagiochasmatos">
<emphasis id="520F6E862A4E5291FAABABB41D5B6EC4" italics="true" pageId="0" pageNumber="1">P. plagiochasmatos</emphasis>
</taxonomicName>
and 
<taxonomicName id="B23F0FE19BB0B6E2BB6BF93956545D56" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. aratriformis" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="aratriformis">
<emphasis id="BE68A18418EB8C7407B90C90E1049DE6" italics="true" pageId="0" pageNumber="1">P. aratriformis</emphasis>
</taxonomicName>
in having a pair of brown lateral bands and pale yellow mark on the scutum; it is distinguished from them by the absence of an extremely extended, forked tubercle-like seta on the subdistal margin of the male epandrium. This species is sympatric with 
<taxonomicName id="F8DA933414202C7E3B16F9F17A76213E" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. aratriformis" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="aratriformis">
<emphasis id="F738165DD40C0B7633DCAA0F48E787F8" italics="true" pageId="0" pageNumber="1">P. aratriformis</emphasis>
</taxonomicName>
on 
<taxonomicName id="9A659CB90231370754AD59DC522C323F" class="Marchantiopsida" family="Aytoniaceae" genus="Reboulia" kingdom="Plantae" lsidName="Reboulia" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="genus">
<emphasis id="6BBBEA1648F8635DA9E6E6486C327B45" italics="true" pageId="0" pageNumber="1">Reboulia</emphasis>
</taxonomicName>
, and can be distinguished from the latter by the yellow mark on the scutum; the mark is large and well defined by lateral stripes in 
<taxonomicName id="195A4C4EB9C15CE2F5AEC59FD823F676" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. falcata" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="falcata">
<emphasis id="796BC8280599011C4BA1F81405E6EDE0" italics="true" pageId="0" pageNumber="1">P. falcata</emphasis>
</taxonomicName>
but small and obscure in 
<taxonomicName id="30BA275078F5990FA0CCC5FC1C52E018" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. aratriformis" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="aratriformis">
<emphasis id="2FC494DE5DB9E0938BA090EFABCCF862" italics="true" pageId="0" pageNumber="1">P. aratriformis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>