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<document id="31732CA37BB5530EA79DBC92F333CE8F" ID-CLB-Dataset="54548" ID-DOI="10.11646/zootaxa.5222.5.1" ID-GBIF-Dataset="a2aef7c8-09e1-48f1-983e-00132055a5a2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7471736" ID-ZooBank="F4983B58-7301-4A3F-AC08-3E5120450EA1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1671694447742" checkinUser="plazi" docAuthor="Evenhuis, Neal L. &amp; Gharali, Babak" docDate="2022" docId="A1541B07260EFFB5FF60FC451A06F8CE" docLanguage="en" docName="zootaxa.5222.5.1.pdf" docOrigin="Zootaxa 5222 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Empidideicus (Anomaloptilus) rinhai Evenhuis &amp; Gharali 2022, n. sp." docType="treatment" docVersion="7" lastPageNumber="423" masterDocId="5D6D637F261FFFA3FFF7FFEB1E24FFF4" masterDocTitle="The subgenus Empidideicus (Anomaloptilus) (Diptera: Mythicomyiidae) in Madagascar" masterLastPageNumber="426" masterPageNumber="401" pageNumber="418" updateTime="1699413895303" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="A5A1814E81B9B6D5115633A2D82E4F08" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="18B54B3E15F5B7B9175A1F23D891D412">
<mods:title id="DDBB7D6141CA8A0A46EBBEBCE6E33230">The subgenus Empidideicus (Anomaloptilus) (Diptera: Mythicomyiidae) in Madagascar</mods:title>
</mods:titleInfo>
<mods:name id="D6A3DDF6EB6BBAF7F24DEACF08397DED" type="personal">
<mods:role id="C240C70D0C02459C9F3D45846245390E">
<mods:roleTerm id="4170E13347D4B05B73FC8AA2B601B4A3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="80D8C4C666ACA75DA69A1615186A4170">Evenhuis, Neal L.</mods:namePart>
<mods:nameIdentifier id="E310DF71E32F7C75045FFE7CB6F4AF78" type="ORCID">0000-0002-1314-755X</mods:nameIdentifier>
<mods:affiliation id="FD3354B547DA0FC1C170CE15B9B92CC4">J. Linsley Gressitt Center for Research in Entomology, Bishop Museum, 1525 Bernice Street, Honolulu, Hawaii 96817 - 2704, USA neale @ bishopmuseum. org; https: // orcid. org / 0000 - 0002 - 1314 - 755 X</mods:affiliation>
<mods:nameIdentifier id="B616D19958C0E7BBE31AF9E4E184CA9B" type="email">neale@bishopmuseum.org</mods:nameIdentifier>
</mods:name>
<mods:name id="5EA874D61968AAA4C15BB3F1543BCFC1" type="personal">
<mods:role id="2843FE239648BE6A4E989AEC2647858A">
<mods:roleTerm id="73E9706BF9BE8F810E9116AA93FC5CBF">Author</mods:roleTerm>
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<mods:namePart id="13B23418DC92E250E80AE62D8A301EAA">Gharali, Babak</mods:namePart>
<mods:nameIdentifier id="BDEB8FC4F2559E1B46B3813F11675DCD" type="ORCID">0000-0003-2490-156X</mods:nameIdentifier>
<mods:affiliation id="9FACFEE32A35B398554A74D26B578110">Plant Protection Research Department, Qazvin Agricultural and Natural Resources Research and Education Center, AREEO, Qazvin, Iran. bgaharaei @ yahoo. com; https: // orcid. org / 0000 - 0003 - 2490 - 156 X</mods:affiliation>
<mods:nameIdentifier id="32E387591D041C9D7201CE9F36AB9371" type="email">bgaharaei@yahoo.com</mods:nameIdentifier>
</mods:name>
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<mods:title id="23F5EBCF30E7E9017C3C510F43BC1077">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="006F23597625EF97C4CFBACDB263E048">
<mods:date id="D7E7C0F331EC0AB8ED1E8B87DF73A401">2022</mods:date>
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<mods:number id="4586A85A769C9A6D8C3D4F9286B7D2E4">2022-12-22</mods:number>
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<mods:number id="6D824C54D4C5730AAAC0F59CC8EA7E20">5222</mods:number>
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<mods:classification id="C54172AD719F6B862947FAEA826B9A3D">journal article</mods:classification>
<mods:identifier id="4254B755D737DD22D0543315F426BF79" type="CLB-Dataset">54548</mods:identifier>
<mods:identifier id="9594768F8ED4E42CDAC827A7FF56F61A" type="DOI">10.11646/zootaxa.5222.5.1</mods:identifier>
<mods:identifier id="FEC543FB0991F376D99CC4ED232796F3" type="GBIF-Dataset">a2aef7c8-09e1-48f1-983e-00132055a5a2</mods:identifier>
<mods:identifier id="7675C0A8CFD816D8CF3441D9E6B7ED2D" type="ISSN">1175-5326</mods:identifier>
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<treatment id="A1541B07260EFFB5FF60FC451A06F8CE" ID-DOI="http://doi.org/10.5281/zenodo.7471800" ID-GBIF-Taxon="207927034" ID-Zenodo-Dep="7471800" LSID="urn:lsid:plazi:treatment:A1541B07260EFFB5FF60FC451A06F8CE" httpUri="http://treatment.plazi.org/id/A1541B07260EFFB5FF60FC451A06F8CE" lastPageId="22" lastPageNumber="423" pageId="17" pageNumber="418">
<subSubSection id="61E7F99A260EFFB2FF60FC451D94FC3D" box="[151,944,942,969]" pageId="17" pageNumber="418" type="nomenclature">
<paragraph id="2942AA11260EFFB2FF60FC451D94FC3D" blockId="17.[151,944,942,1005]" box="[151,944,942,969]" pageId="17" pageNumber="418">
<heading id="720A1D7D260EFFB2FF60FC451D94FC3D" bold="true" box="[151,944,942,969]" fontSize="11" level="1" pageId="17" pageNumber="418" reason="1">
<emphasis id="1B897603260EFFB2FF60FC451D94FC3D" bold="true" box="[151,944,942,969]" pageId="17" pageNumber="418">
<taxonomicName id="EEFDD192260EFFB2FF60FC451D45FC3C" authority="Evenhuis &amp; Gharali" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[151,865,942,969]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="rinhai" status="sp. nov." subGenus="Anomaloptilus">
<emphasis id="1B897603260EFFB2FF60FC451C7FFC3D" bold="true" box="[151,603,942,969]" italics="true" pageId="17" pageNumber="418">Empidideicus (Anomaloptilus) rinhai</emphasis>
Evenhuis &amp; Gharali
</taxonomicName>
,
<taxonomicNameLabel id="00BACB78260EFFB2FC9AFC441D94FC3D" box="[877,944,943,969]" pageId="17" pageNumber="418" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61E7F99A260EFFB2FF60FC381EC9FB1D" pageId="17" pageNumber="418" type="description">
<paragraph id="2942AA11260EFFB2FF60FC381C69FC19" blockId="17.[151,944,942,1005]" box="[151,589,978,1005]" pageId="17" pageNumber="418">
(
<figureCitation id="B1C6B694260EFFB2FF68FC381F04FC19" box="[159,288,978,1005]" captionStart="FIGURES 11–16" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[158,1429,181,1920]" captionTargetId="figure-17@5.[158,1429,181,1920]" captionTargetPageId="5" captionText="FIGURES 11–16. Malagasy Empidideicus (Anomaloptilus), habitus, lateral view. 11, E. (A.) regius, n. sp., male; 12, E. (A.) regius, n. sp., female; 13, E. (A.) rhinoclypeatus Evenhuis, male; 14, E. (A.) rhinoclypeatus Evenhuis, female; 15, E. (A.) rinhai, n. sp., male; 16, E. (A.) rinhai, n. sp., female." figureDoi="http://doi.org/10.5281/zenodo.7471748" httpUri="https://zenodo.org/record/7471748/files/figure.png" pageId="17" pageNumber="418">Figs 15, 16</figureCitation>
,
<figureCitation id="B1C6B694260EFFB2FEDBFC381F6EFC19" box="[300,330,979,1005]" captionStart="FIGURES 17–19" captionStartId="8.[151,264,1390,1414]" captionTargetBox="[151,1435,741,1367]" captionTargetId="figure-245@8.[151,1435,741,1367]" captionTargetPageId="8" captionText="FIGURES 17–19. Malagasy Empidideicus (Anomaloptilus), heads. 17, E. (A.) andriambahomananii, n. sp., male, oblique view; 18, E. (A.) irwini, n. sp., female, dorsal view; 19, E. (A.) rinhai, n. sp., male, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7471750" httpUri="https://zenodo.org/record/7471750/files/figure.png" pageId="17" pageNumber="418">19</figureCitation>
,
<figureCitation id="B1C6B694260EFFB2FEA1FC381FBAFC19" box="[342,414,979,1005]" captionStart="FIGURES 34–39" captionStartId="14.[151,264,1264,1288]" captionTargetBox="[151,1435,181,1240]" captionTargetId="figure-17@14.[151,1435,181,1240]" captionTargetPageId="14" captionText="FIGURES 34–39. Malagasy Empidideicus (Anomaloptilus), thoraces, dorsal view. 34, E. (A.) regius, n. sp., male; 35, E. (A.) regius, n. sp., female; 36, E. (A.) rhinoclypeatus Evenhuis, male; 37, E. (A.) rhinoclypeatus Evenhuis, female; 38, E. (A.) rinhai, n. sp., male; 39, E. (A.) rinhai, n. sp., female." figureDoi="http://doi.org/10.5281/zenodo.7471760" httpUri="https://zenodo.org/record/7471760/files/figure.png" pageId="17" pageNumber="418">38, 39</figureCitation>
,
<figureCitation id="B1C6B694260EFFB2FE5DFC381FECFC19" box="[426,456,979,1005]" captionStart="FIGURES 40–47" captionStartId="15.[151,264,1607,1631]" captionTargetBox="[170,1416,309,1582]" captionTargetId="figure-81@15.[170,1416,309,1584]" captionTargetPageId="15" captionText="FIGURES 40–47. Malagasy Empidideicus (Anomaloptilus), wings. 40, E. (A.) andriambahomananii, n. sp.; 41, E. (A.) contrasticus, n. sp.; 42, E. (A.) hauseri, n. sp.; 43, E. (A.) irwini, n. sp.; 44, E. (A.) nanus, n. sp.; 45, E. (A.) regius, n. sp.; 46, E. (A.) rhinoclypeatus Evenhuis; 47, E. (A.) rinhai, n. sp." figureDoi="http://doi.org/10.5281/zenodo.7471764" httpUri="https://zenodo.org/record/7471764/files/figure.png" pageId="17" pageNumber="418">47</figureCitation>
,
<figureCitation id="B1C6B694260EFFB2FE23FC381C38FC19" box="[468,540,979,1005]" captionStart="FIGURES 59–62" captionStartId="19.[151,264,694,718]" captionTargetBox="[152,1435,181,670]" captionTargetId="figure-17@19.[152,1435,181,670]" captionTargetPageId="19" captionText="FIGURES 59–62. Malagasy Empidideicus (Anomaloptilus), abdomens, dorsal view. 59, E. (A.) rhinoclypeatus Evenhuis, male; 60, E. (A.) rhinoclypeatus Evenhuis, female; 61, E. (A.) rinhai, n. sp., male; 62, E. (A.) rinhai, n. sp., female." figureDoi="http://doi.org/10.5281/zenodo.7471774" httpUri="https://zenodo.org/record/7471774/files/figure.png" pageId="17" pageNumber="418">61, 62</figureCitation>
,
<figureCitation id="B1C6B694260EFFB2FDDFFC381C61FC19" box="[552,581,979,1005]" captionStart="FIGURES 77" captionStartId="23.[152,270,1257,1281]" captionTargetBox="[160,1433,187,1228]" captionTargetId="figure-17@23.[151,1436,181,1233]" captionTargetPageId="23" captionText="FIGURES 77. Empidideicus (Anomaloptilus) rinhai, n. sp., male genitalia, A, lateral view, B, gonocoxae, ventral view, C, phallic complex, dorsal view, D, apex of dorsoventral epandrial process, detail, E, gonostylus, detail." figureDoi="http://doi.org/10.5281/zenodo.7471798" httpUri="https://zenodo.org/record/7471798/files/figure.png" pageId="17" pageNumber="418">77</figureCitation>
)
</paragraph>
<paragraph id="2942AA11260EFFB2FF60FBF01EC9FB1D" blockId="17.[151,1437,1051,1977]" pageId="17" pageNumber="418">
<emphasis id="1B897603260EFFB2FF60FBF01EC0FBC1" bold="true" box="[151,228,1051,1077]" pageId="17" pageNumber="418">Types.</emphasis>
<materialsCitation id="9995A04C260EFFB2FF1CFBF01FBAFBC1" ID-GBIF-Occurrence="4003029353" box="[235,414,1051,1079]" collectingDate="2002-12-09" collectingDateMax="2003-05-11" collectingDateMin="2002-12-09" collectionCode="ANGAP, R" collectorName="M. E. Irwin &amp; F. Parker &amp; M. Irwin &amp; Harin'Hala" country="Madagascar" county="Cape Ste. Marie Special Reserve" elevation="840" latitude="-25.587667" location="Tulear Province" longLatPrecision="12" longitude="45.163" municipality="Tulear Province" pageId="17" pageNumber="418" specimenCode="CASLOT 024735, CASLOT 024734, CASLOT 024746" specimenCount="3" specimenCount-female="1" typeStatus="holotype">
<emphasis id="1B897603260EFFB2FF1CFBF01F76FBC1" box="[235,338,1051,1077]" italics="true" pageId="17" pageNumber="418">
<typeStatus id="F64614B3260EFFB2FF1CFBF01F76FBC1" box="[235,338,1051,1077]" pageId="17" pageNumber="418" type="holotype">Holotype</typeStatus>
</emphasis>
♂ and
</materialsCitation>
<materialsCitation id="9995A04C260EFFB2FE50FBF71F5FFB89" ID-GBIF-Occurrence="4003029320" collectingDate="2002-12-09" collectingDateMax="2003-05-11" collectingDateMin="2002-12-09" collectionCode="ANGAP, R" collectorName="M. E. Irwin &amp; F. Parker &amp; M. Irwin &amp; Harin'Hala" country="Madagascar" county="Cape Ste. Marie Special Reserve" elevation="840" latitude="-25.587667" location="Tulear Province" longLatPrecision="12" longitude="45.163" municipality="Tulear Province" pageId="17" pageNumber="418" specimenCode="CASLOT 024735, CASLOT 024734, CASLOT 024746" specimenCount="3" specimenCount-female="1" typeStatus="paratype">
1♂,
<specimenCount id="3FFB6198260EFFB2FE2FFBF01FDFFBC1" box="[472,507,1051,1077]" pageId="17" pageNumber="418" type="female">1♀</specimenCount>
<emphasis id="1B897603260EFFB2FDF4FBF01C55FBC0" box="[515,625,1051,1076]" italics="true" pageId="17" pageNumber="418">
<typeStatus id="F64614B3260EFFB2FDF4FBF01C55FBC0" box="[515,625,1051,1076]" pageId="17" pageNumber="418" type="paratype">paratypes</typeStatus>
</emphasis>
from
<collectingCountry id="51EAEA81260EFFB2FD40FBF01DA7FBC1" box="[695,899,1051,1077]" name="Madagascar" pageId="17" pageNumber="418">MADAGASCAR</collectingCountry>
:
<location id="2C22FCCA260EFFB2FC79FBF01A61FBC1" LSID="urn:lsid:plazi:treatment:A1541B07260EFFB5FF60FC451A06F8CE:2C22FCCA260EFFB2FC79FBF01A61FBC1" box="[910,1093,1051,1077]" country="Madagascar" county="Cape Ste. Marie Special Reserve" latitude="-25.587667" longLatPrecision="12" longitude="45.163" municipality="Tulear Province" name="Tulear Province" pageId="17" pageNumber="418">Tulear Province</location>
:
<location id="2C22FCCA260EFFB2FBA7FBF01B44FBC1" LSID="urn:lsid:plazi:treatment:A1541B07260EFFB5FF60FC451A06F8CE:2C22FCCA260EFFB2FBA7FBF01B44FBC1" box="[1104,1376,1051,1077]" country="Madagascar" county="Cape Ste. Marie Special Reserve" latitude="-25.587667" longLatPrecision="12" longitude="45.163" municipality="Tulear Province" name="Zombitse National Park" pageId="17" pageNumber="418">Zombitse National Park</location>
, near
<collectionCode id="4FEC32D4260EFFB2FF60FBD41ED2FBAD" box="[151,246,1087,1113]" pageId="17" pageNumber="418">ANGAP</collectionCode>
<location id="2C22FCCA260EFFB2FF0AFBD41F63FBAD" LSID="urn:lsid:plazi:treatment:A1541B07260EFFB5FF60FC451A06F8CE:2C22FCCA260EFFB2FF0AFBD41F63FBAD" box="[253,327,1087,1113]" country="Madagascar" county="Cape Ste. Marie Special Reserve" latitude="-25.587667" longLatPrecision="12" longitude="45.163" municipality="Tulear Province" name="Office" pageId="17" pageNumber="418">Office</location>
,
<quantity id="EE0507F4260EFFB2FEA5FBD41FBEFBAD" box="[338,410,1087,1113]" metricMagnitude="2" metricUnit="m" metricValue="8.4" pageId="17" pageNumber="418" unit="m" value="840.0">
<elevation id="A2D04D22260EFFB2FEA5FBD41FBEFBAD" box="[338,410,1087,1113]" metricMagnitude="2" metricUnit="m" metricValue="8.4" pageId="17" pageNumber="418" unit="m" value="840.0">840 m</elevation>
</quantity>
,
<date id="5D438CD1260EFFB2FE52FBD41C44FBAD" box="[421,608,1087,1113]" pageId="17" pageNumber="418" value="2002-12-18" valueMax="2002-12-23" valueMin="2002-12-18">
<collectingDate id="4D077539260EFFB2FE52FBD41C44FBAD" box="[421,608,1087,1113]" pageId="17" pageNumber="418" value="2002-12-18" valueMax="2002-12-23" valueMin="2002-12-18">18–23 Dec 2002</collectingDate>
</date>
,
<collectorName id="8408CFC7260EFFB2FD9CFBD41CCCFBAD" box="[619,744,1087,1113]" pageId="17" pageNumber="418">M.E. Irwin</collectorName>
,
<collectorName id="8408CFC7260EFFB2FD03FBD41D73FBAD" box="[756,855,1087,1113]" pageId="17" pageNumber="418">F. Parker</collectorName>
&amp;
<collectionCode id="4FEC32D4260EFFB2FC8DFBD41DB4FBAD" box="[890,912,1087,1113]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="418">R</collectionCode>
.
<location id="2C22FCCA260EFFB2FC6DFBD41A3FFBAD" LSID="urn:lsid:plazi:treatment:A1541B07260EFFB5FF60FC451A06F8CE:2C22FCCA260EFFB2FC6DFBD41A3FFBAD" box="[922,1051,1087,1113]" country="Madagascar" county="Cape Ste. Marie Special Reserve" latitude="-25.587667" longLatPrecision="12" longitude="45.163" municipality="Tulear Province" name="Harin'Hala" pageId="17" pageNumber="418">Harin’Hala</location>
,
<location id="2C22FCCA260EFFB2FBD1FBD41A5BFBAD" LSID="urn:lsid:plazi:treatment:A1541B07260EFFB5FF60FC451A06F8CE:2C22FCCA260EFFB2FBD1FBD41A5BFBAD" box="[1062,1151,1087,1113]" country="Madagascar" county="Cape Ste. Marie Special Reserve" latitude="-25.587667" longLatPrecision="12" longitude="45.163" municipality="Tulear Province" name="Malaise" pageId="17" pageNumber="418">Malaise</location>
in deciduous spiny forest (
<specimenCode id="795B026A260EFFB2FF68FB881F4BFB89" box="[159,367,1122,1149]" collectionCode="CASLOT" pageId="17" pageNumber="418">CASLOT 024735</specimenCode>
).
</materialsCitation>
<materialsCitation id="9995A04C260EFFB2FE74FB881ED8FB31" ID-GBIF-Occurrence="4003029307" collectingDate="2002-12-09" collectingDateMax="2003-05-11" collectingDateMin="2002-12-09" collectionCode="ANGAP, R" collectorName="M. E. Irwin &amp; F. Parker &amp; M. Irwin &amp; Harin'Hala" country="Madagascar" county="Cape Ste. Marie Special Reserve" elevation="840" latitude="-25.587667" location="Tulear Province" longLatPrecision="12" longitude="45.163" municipality="Tulear Province" pageId="17" pageNumber="418" specimenCode="CASLOT 024735, CASLOT 024734, CASLOT 024746" specimenCount="3" specimenCount-female="1" typeStatus="paratype">
<emphasis id="1B897603260EFFB2FE74FB881C18FB88" box="[387,572,1123,1149]" italics="true" pageId="17" pageNumber="418">
<location id="2C22FCCA260EFFB2FE74FB881FE0FB89" LSID="urn:lsid:plazi:treatment:A1541B07260EFFB5FF60FC451A06F8CE:2C22FCCA260EFFB2FE74FB881FE0FB89" box="[387,452,1123,1149]" country="Madagascar" county="Cape Ste. Marie Special Reserve" latitude="-25.587667" longLatPrecision="12" longitude="45.163" municipality="Tulear Province" name="Other" pageId="17" pageNumber="418">Other</location>
<typeStatus id="F64614B3260EFFB2FE39FB881C18FB88" box="[462,572,1123,1148]" pageId="17" pageNumber="418" type="paratype">paratypes</typeStatus>
</emphasis>
:
<collectingCountry id="51EAEA81260EFFB2FDBAFB881D3DFB89" box="[589,793,1123,1149]" name="Madagascar" pageId="17" pageNumber="418">MADAGASCAR</collectingCountry>
: 2♂,
<collectingMunicipality id="C926306B260EFFB2FCAFFB881A35FB89" box="[856,1041,1123,1149]" pageId="17" pageNumber="418">Tulear Province</collectingMunicipality>
:
<collectingCounty id="C023D29D260EFFB2FBEAFB881BBCFB89" box="[1053,1432,1123,1149]" pageId="17" pageNumber="418">Cape Ste. Marie Special Reserve</collectingCounty>
,
<geoCoordinate id="4CC9CCD6260EFFB2FF60FB631F32FB57" box="[151,278,1160,1187]" degrees="25" direction="south" minutes="35.26" orientation="latitude" pageId="17" pageNumber="418" precision="9" value="-25.587667">25°35.26′S</geoCoordinate>
,
<geoCoordinate id="4CC9CCD6260EFFB2FED7FB631F84FB57" box="[288,416,1160,1187]" degrees="45" direction="east" minutes="09.78" orientation="longitude" pageId="17" pageNumber="418" precision="9" value="45.163">45°09.78′E</geoCoordinate>
,
<quantity id="EE0507F4260EFFB2FE5DFB6C1FC0FB55" box="[426,484,1159,1185]" metricMagnitude="1" metricUnit="m" metricValue="3.7" pageId="17" pageNumber="418" unit="m" value="37.0">
<elevation id="A2D04D22260EFFB2FE5DFB6C1FC0FB55" box="[426,484,1159,1185]" metricMagnitude="1" metricUnit="m" metricValue="3.7" pageId="17" pageNumber="418" unit="m" value="37.0">37 m</elevation>
</quantity>
,
<date id="5D438CD1260EFFB2FE18FB6D1CBFFB55" box="[495,667,1158,1185]" pageId="17" pageNumber="418" value="2002-12-09" valueMax="2002-12-15" valueMin="2002-12-09">
<collectingDate id="4D077539260EFFB2FE18FB6D1CBFFB55" box="[495,667,1158,1185]" pageId="17" pageNumber="418" value="2002-12-09" valueMax="2002-12-15" valueMin="2002-12-09">9–15 Dec 2002</collectingDate>
</date>
,
<collectorName id="8408CFC7260EFFB2FD52FB6C1D2EFB55" box="[677,778,1159,1185]" pageId="17" pageNumber="418">M. Irwin</collectorName>
,
<collectorName id="8408CFC7260EFFB2FCE2FB6C1D5EFB55" box="[789,890,1159,1185]" pageId="17" pageNumber="418">F. Parker</collectorName>
,
<collectionCode id="4FEC32D4260EFFB2FC73FB6C1DBEFB55" box="[900,922,1159,1185]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="418">R</collectionCode>
.
<collectorName id="8408CFC7260EFFB2FC53FB6C1A01FB55" box="[932,1061,1159,1185]" pageId="17" pageNumber="418">Harin’Hala</collectorName>
,
<collectorName id="8408CFC7260EFFB2FBD8FB6C1AACFB55" box="[1071,1160,1159,1185]" pageId="17" pageNumber="418">Malaise</collectorName>
in spiny bush (
<specimenCode id="795B026A260EFFB2FAD9FB6C1ED4FB31" collectionCode="CASLOT" pageId="17" pageNumber="418">CASLOT 024734</specimenCode>
);
</materialsCitation>
<materialsCitation id="9995A04C260EFFB2FEF4FB471DEAFB31" ID-GBIF-Occurrence="4003029322" box="[259,974,1195,1223]" collectingDate="2002-12-09" collectingDateMax="2003-05-11" collectingDateMin="2002-12-09" collectionCode="ANGAP, R" collectorName="M. E. Irwin &amp; F. Parker &amp; Harin'Hala" country="Madagascar" county="Cape Ste. Marie Special Reserve" elevation="840" latitude="-25.587667" location="Tulear Province" longLatPrecision="12" longitude="45.163" municipality="Tulear Province" pageId="17" pageNumber="418" specimenCode="CASLOT 024735, CASLOT 024734, CASLOT 024746" specimenCount="3" specimenCount-female="1" typeStatus="holotype">
1♂, same data except
<date id="5D438CD1260EFFB2FE0DFB401CCDFB31" box="[506,745,1195,1221]" pageId="17" pageNumber="418" value="2003-04-30" valueMax="2003-05-11" valueMin="2003-04-30">
<collectingDate id="4D077539260EFFB2FE0DFB401CCDFB31" box="[506,745,1195,1221]" pageId="17" pageNumber="418" value="2003-04-30" valueMax="2003-05-11" valueMin="2003-04-30">30 Apr–11 May 2003</collectingDate>
</date>
(
<specimenCode id="795B026A260EFFB2FD00FB401DE1FB31" box="[759,965,1195,1221]" collectionCode="CASLOT" pageId="17" pageNumber="418">CASLOT 024746</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="9995A04C260EFFB2FC2FFB401ECCFB1D" ID-GBIF-Occurrence="4003029310" collectionCode="CAS, BPBM" country="Madagascar" pageId="17" pageNumber="418" specimenCount="2" typeStatus="holotype">
<typeStatus id="F64614B3260EFFB2FC2FFB401A64FB31" box="[984,1088,1195,1221]" pageId="17" pageNumber="418" type="holotype">Holotype</typeStatus>
in
<collectionCode id="4FEC32D4260EFFB2FB94FB401AB8FB31" box="[1123,1180,1195,1221]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="17" pageNumber="418" type="Museum">CAS</collectionCode>
,
<typeStatus id="F64614B3260EFFB2FB5FFB401B37FB31" box="[1192,1299,1195,1221]" pageId="17" pageNumber="418" type="paratype">paratypes</typeStatus>
in
<collectionCode id="4FEC32D4260EFFB2FAC1FB401B48FB31" box="[1334,1388,1195,1221]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="17" pageNumber="418" type="Museum">CAS</collectionCode>
and
<collectionCode id="4FEC32D4260EFFB2FF60FB241ECCFB1D" box="[151,232,1231,1257]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="17" pageNumber="418" type="Museum">BPBM</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="61E7F99A260EFFB2FF30FB191AB3FA8D" pageId="17" pageNumber="418" type="diagnosis">
<paragraph id="2942AA11260EFFB2FF30FB191AB3FA8D" blockId="17.[151,1437,1051,1977]" pageId="17" pageNumber="418">
<emphasis id="1B897603260EFFB2FF30FB191F66FAF8" bold="true" box="[199,322,1266,1292]" pageId="17" pageNumber="418">Diagnosis.</emphasis>
Most similar to
<emphasis id="1B897603260EFFB2FDF6FB181C3DFAF8" box="[513,537,1267,1292]" italics="true" pageId="17" pageNumber="418">E.</emphasis>
<taxonomicName id="EEFDD192260EFFB2FDD6FB181C8DFAF9" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[545,681,1267,1293]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="hauseri" status="sp. nov." subGenus="Anomaloptilus">
(
<emphasis id="1B897603260EFFB2FDDCFB181C18FAF9" box="[555,572,1267,1293]" italics="true" pageId="17" pageNumber="418">A</emphasis>
.)
<emphasis id="1B897603260EFFB2FDA2FB181C8DFAF9" box="[597,681,1267,1293]" italics="true" pageId="17" pageNumber="418">hauseri</emphasis>
</taxonomicName>
<emphasis id="1B897603260EFFB2FD46FB181CD5FAF9" bold="true" box="[689,753,1267,1293]" pageId="17" pageNumber="418">
<taxonomicNameLabel id="00BACB78260EFFB2FD46FB181CD5FAF9" box="[689,753,1267,1293]" pageId="17" pageNumber="418" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
based on having yellow marks in the prescutellar area. It is eastly separated from it based on size (
<emphasis id="1B897603260EFFB2FDBEFAFC1C45FAC4" box="[585,609,1303,1328]" italics="true" pageId="17" pageNumber="418">E.</emphasis>
<taxonomicName id="EEFDD192260EFFB2FD9FFAFC1CF9FAC5" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[616,733,1303,1329]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="rinhai" status="sp. nov." subGenus="Anomaloptilus">
(
<emphasis id="1B897603260EFFB2FD86FAFC1CA6FAC5" box="[625,642,1303,1329]" italics="true" pageId="17" pageNumber="418">A</emphasis>
.)
<emphasis id="1B897603260EFFB2FD6EFAFC1CF9FAC5" box="[665,733,1303,1329]" italics="true" pageId="17" pageNumber="418">rinhai</emphasis>
</taxonomicName>
<emphasis id="1B897603260EFFB2FD12FAFC1D07FAC5" bold="true" box="[741,803,1303,1329]" pageId="17" pageNumber="418">
<taxonomicNameLabel id="00BACB78260EFFB2FD12FAFC1D07FAC5" box="[741,803,1303,1329]" pageId="17" pageNumber="418" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is ca.
<quantity id="EE0507F4260EFFB2FC9CFAFC1D8FFAC5" box="[875,939,1303,1329]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="418" unit="mm" value="1.0">1 mm</quantity>
long; whereas
<emphasis id="1B897603260EFFB2FBA2FAFC1A49FAC4" box="[1109,1133,1303,1328]" italics="true" pageId="17" pageNumber="418">E.</emphasis>
<taxonomicName id="EEFDD192260EFFB2FB83FAFC1ADDFAC5" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[1140,1273,1303,1329]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="hauseri" status="sp. nov." subGenus="Anomaloptilus">
(
<emphasis id="1B897603260EFFB2FB8AFAFC1AAAFAC5" box="[1149,1166,1303,1329]" italics="true" pageId="17" pageNumber="418">A</emphasis>
.)
<emphasis id="1B897603260EFFB2FB52FAFC1ADDFAC5" box="[1189,1273,1303,1329]" italics="true" pageId="17" pageNumber="418">hauseri</emphasis>
</taxonomicName>
<emphasis id="1B897603260EFFB2FAF7FAFC1B1BFAC5" bold="true" box="[1280,1343,1303,1329]" pageId="17" pageNumber="418">
<taxonomicNameLabel id="00BACB78260EFFB2FAF7FAFC1B1BFAC5" box="[1280,1343,1303,1329]" pageId="17" pageNumber="418" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is twice that). It is also separated by the lack of coalesced yellow color in the prescutellar area (broadly yellow in
<emphasis id="1B897603260EFFB2FAA6FAD01B4DFAA0" box="[1361,1385,1339,1364]" italics="true" pageId="17" pageNumber="418">E.</emphasis>
<taxonomicName id="EEFDD192260EFFB2FA86FAD01ECFFA8D" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="hauseri" status="sp. nov." subGenus="Anomaloptilus">
(
<emphasis id="1B897603260EFFB2FA8CFAD01BA8FAA1" box="[1403,1420,1339,1365]" italics="true" pageId="17" pageNumber="418">A</emphasis>
.)
<emphasis id="1B897603260EFFB2FF60FAB41ECFFA8D" box="[151,235,1375,1401]" italics="true" pageId="17" pageNumber="418">hauseri</emphasis>
</taxonomicName>
<emphasis id="1B897603260EFFB2FF05FAB41F15FA8D" bold="true" box="[242,305,1375,1401]" pageId="17" pageNumber="418">
<taxonomicNameLabel id="00BACB78260EFFB2FF05FAB41F15FA8D" box="[242,305,1375,1401]" pageId="17" pageNumber="418" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
) and the open anal cell in the wing (anal cell closed in
<emphasis id="1B897603260EFFB2FC6CFAB41D90FA8C" box="[923,948,1375,1400]" italics="true" pageId="17" pageNumber="418">E.</emphasis>
<taxonomicName id="EEFDD192260EFFB2FC4DFAB41A64FA8D" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[954,1088,1375,1401]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="hauseri" status="sp. nov." subGenus="Anomaloptilus">
(
<emphasis id="1B897603260EFFB2FC33FAB41DF1FA8D" box="[964,981,1375,1401]" italics="true" pageId="17" pageNumber="418">A</emphasis>
.)
<emphasis id="1B897603260EFFB2FC1BFAB41A64FA8D" box="[1004,1088,1375,1401]" italics="true" pageId="17" pageNumber="418">hauseri</emphasis>
</taxonomicName>
<emphasis id="1B897603260EFFB2FBB0FAB41AA2FA8D" bold="true" box="[1095,1158,1375,1401]" pageId="17" pageNumber="418">
<taxonomicNameLabel id="00BACB78260EFFB2FBB0FAB41AA2FA8D" box="[1095,1158,1375,1401]" pageId="17" pageNumber="418" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="61E7F99A260EFFB5FF30FA691FCEF93A" lastPageId="22" lastPageNumber="423" pageId="17" pageNumber="418" type="description">
<paragraph id="2942AA11260EFFB2FF30FA691F75FA68" blockId="17.[151,1437,1051,1977]" box="[199,337,1410,1436]" pageId="17" pageNumber="418">
<emphasis id="1B897603260EFFB2FF30FA691F75FA68" bold="true" box="[199,337,1410,1436]" pageId="17" pageNumber="418">Description</emphasis>
</paragraph>
<paragraph id="2942AA11260EFFB2FF30FA4C1C5AF949" blockId="17.[151,1437,1051,1977]" pageId="17" pageNumber="418">
<emphasis id="1B897603260EFFB2FF30FA4C1F20FA35" bold="true" box="[199,260,1447,1473]" pageId="17" pageNumber="418">Male</emphasis>
(
<figureCitation id="B1C6B694260EFFB2FEE5FA4C1F41FA35" box="[274,357,1446,1473]" captionStart="FIGURES 11–16" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[158,1429,181,1920]" captionTargetId="figure-17@5.[158,1429,181,1920]" captionTargetPageId="5" captionText="FIGURES 11–16. Malagasy Empidideicus (Anomaloptilus), habitus, lateral view. 11, E. (A.) regius, n. sp., male; 12, E. (A.) regius, n. sp., female; 13, E. (A.) rhinoclypeatus Evenhuis, male; 14, E. (A.) rhinoclypeatus Evenhuis, female; 15, E. (A.) rinhai, n. sp., male; 16, E. (A.) rinhai, n. sp., female." figureDoi="http://doi.org/10.5281/zenodo.7471748" httpUri="https://zenodo.org/record/7471748/files/figure.png" pageId="17" pageNumber="418">Fig. 15</figureCitation>
). Length:
<quantity id="EE0507F4260EFFB2FE21FA4C1C5FFA35" box="[470,635,1447,1473]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="17" pageNumber="418" unit="mm" value="1.25" valueMax="1.3" valueMin="1.2">1.20–1.30 mm</quantity>
. Wing: 1.00–
<quantity id="EE0507F4260EFFB2FCF8FA4C1D51FA35" box="[783,885,1447,1473]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="17" pageNumber="418" unit="mm" value="1.4">1.40 mm</quantity>
.
<emphasis id="1B897603260EFFB2FC88FA4C1D98FA35" box="[895,956,1447,1473]" italics="true" pageId="17" pageNumber="418">Head</emphasis>
(
<figureCitation id="B1C6B694260EFFB2FC3DFA4C1A39FA35" box="[970,1053,1447,1473]" captionStart="FIGURES 17–19" captionStartId="8.[151,264,1390,1414]" captionTargetBox="[151,1435,741,1367]" captionTargetId="figure-245@8.[151,1435,741,1367]" captionTargetPageId="8" captionText="FIGURES 17–19. Malagasy Empidideicus (Anomaloptilus), heads. 17, E. (A.) andriambahomananii, n. sp., male, oblique view; 18, E. (A.) irwini, n. sp., female, dorsal view; 19, E. (A.) rinhai, n. sp., male, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7471750" httpUri="https://zenodo.org/record/7471750/files/figure.png" pageId="17" pageNumber="418">Fig. 19</figureCitation>
): longer than high; eyes dichoptic, separated at vertex by 1.7× distance between lateral ocelli, tapering to narrow face, width about equal to 3 ommatidia; occiput and postgena black; front slightly depressed medially, white; face gray; antennae (cf.
<figureCitation id="B1C6B694260EFFB2FB36FA041B37F9FD" box="[1217,1299,1519,1545]" captionStart="FIGURES 23–24" captionStartId="10.[151,264,1309,1333]" captionTargetBox="[151,1435,181,1284]" captionTargetId="figure-17@10.[151,1435,181,1285]" captionTargetPageId="10" captionText="FIGURES 23–24. Malagasy Empidideicus (Anomaloptilus), antennae. 23, E. (A.) andriambahomananii, n. sp.; 24, E. (A.) nanus, n. sp." figureDoi="http://doi.org/10.5281/zenodo.7471754" httpUri="https://zenodo.org/record/7471754/files/figure.png" pageId="17" pageNumber="418">Fig. 23</figureCitation>
) with scape and pedicel yellow, flagellomeres dark brown; scape short, subtrapezoidal; pedicel subconical, wider than long; first flagellomere lanceolate, length 3.5× width; second flagellomere one-half length of first flagellomere, with minute transparent apical style; mentum white; clypeus brown, length one-half of oral margin length; proboscis brown, slightly longer than head length; labrum sclerotized, stiff, pointed apically, length subequal to clypeus, yellow basally, brown apically; palpus not evident.
</paragraph>
<paragraph id="2942AA11260EFFB2FF30F92C1B6DF885" blockId="17.[151,1437,1051,1977]" pageId="17" pageNumber="418">
<emphasis id="1B897603260EFFB2FF30F92C1F3CF915" box="[199,280,1735,1761]" italics="true" pageId="17" pageNumber="418">Thorax</emphasis>
(
<figureCitation id="B1C6B694260EFFB2FED0F92C1F5FF915" box="[295,379,1735,1761]" captionStart="FIGURES 34–39" captionStartId="14.[151,264,1264,1288]" captionTargetBox="[151,1435,181,1240]" captionTargetId="figure-17@14.[151,1435,181,1240]" captionTargetPageId="14" captionText="FIGURES 34–39. Malagasy Empidideicus (Anomaloptilus), thoraces, dorsal view. 34, E. (A.) regius, n. sp., male; 35, E. (A.) regius, n. sp., female; 36, E. (A.) rhinoclypeatus Evenhuis, male; 37, E. (A.) rhinoclypeatus Evenhuis, female; 38, E. (A.) rinhai, n. sp., male; 39, E. (A.) rinhai, n. sp., female." figureDoi="http://doi.org/10.5281/zenodo.7471760" httpUri="https://zenodo.org/record/7471760/files/figure.png" pageId="17" pageNumber="418">Fig. 38</figureCitation>
). Mesonotum matte black, with white markings in following areas: thin admedian longitudinal vittae not reaching scutellum, leaving two small white marks in prescutellar area and thin connection to vittae; dorsolaterally from humeral callus to postalar callus and two large interhumeral marks; scutellum white, dark brown below along posterior margin; pleura brown with white color on following: posterior half of anepisternum and anepimeron, upper half of meron; upper margin of katepisternum; propleuron white; halter and knob white.
</paragraph>
<paragraph id="2942AA11260EFFB2FF30F8901A2FF84D" blockId="17.[151,1437,1051,1977]" pageId="17" pageNumber="418">
<emphasis id="1B897603260EFFB2FF30F8901ED5F860" box="[199,241,1915,1940]" italics="true" pageId="17" pageNumber="418">Leg</emphasis>
. Coxae yellow, brown basally, mid and hind coxae with much more brown than fore coxa; femora and tibiae yellow with some brown basally; tarsi 3–5 pale brown, remainder of leg white.
</paragraph>
<caption id="7D82FA99260DFFB1FF60F88A1FA9F811" ID-DOI="http://doi.org/10.5281/zenodo.7471772" ID-Zenodo-Dep="7471772" httpUri="https://zenodo.org/record/7471772/files/figure.png" pageId="18" pageNumber="419" startId="18.[151,264,1889,1913]" targetBox="[239,1348,181,1865]" targetPageId="18">
<paragraph id="2942AA11260DFFB1FF60F88A1FA9F811" blockId="18.[151,1436,1889,2022]" pageId="18" pageNumber="419">
<emphasis id="1B897603260DFFB1FF60F88A1F6AF88D" bold="true" box="[151,334,1889,1913]" pageId="18" pageNumber="419">FIGURES 50–58</emphasis>
. Malagasy
<taxonomicName id="EEFDD192260DFFB1FE31F88A1CD9F88E" box="[454,765,1889,1914]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="419" phylum="Arthropoda" rank="subGenus" subGenus="Anomaloptilus">
<emphasis id="1B897603260DFFB1FE31F88A1C74F88D" box="[454,592,1889,1913]" italics="true" pageId="18" pageNumber="419">Empidideicus</emphasis>
(
<emphasis id="1B897603260DFFB1FD97F88A1CD0F88D" box="[608,756,1889,1913]" italics="true" pageId="18" pageNumber="419">Anomaloptilus</emphasis>
)
</taxonomicName>
, abdomens, dorsal view.
<emphasis id="1B897603260DFFB1FBF7F88A1A3DF88D" bold="true" box="[1024,1049,1889,1913]" pageId="18" pageNumber="419">50</emphasis>
,
<emphasis id="1B897603260DFFB1FBD0F8891A12F88D" box="[1063,1078,1890,1913]" italics="true" pageId="18" pageNumber="419">E</emphasis>
.
<taxonomicName id="EEFDD192260DFFB1FBB2F8891B6BF88D" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[1093,1359,1889,1914]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="andriambahomananii" status="sp. nov." subGenus="Anomaloptilus">
(
<emphasis id="1B897603260DFFB1FBBAF8891A78F88D" box="[1101,1116,1890,1913]" italics="true" pageId="18" pageNumber="419">A</emphasis>
.)
<emphasis id="1B897603260DFFB1FB84F88A1B6BF88D" box="[1139,1359,1889,1913]" italics="true" pageId="18" pageNumber="419">andriambahomananii</emphasis>
</taxonomicName>
,
<emphasis id="1B897603260DFFB1FAABF8891BB2F88E" bold="true" box="[1372,1430,1890,1914]" pageId="18" pageNumber="419">
<taxonomicNameLabel id="00BACB78260DFFB1FAABF8891BB2F88E" box="[1372,1430,1890,1914]" pageId="18" pageNumber="419" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, female;
<emphasis id="1B897603260DFFB1FF1EF86E1F26F869" bold="true" box="[233,258,1925,1949]" pageId="18" pageNumber="419">51</emphasis>
,
<emphasis id="1B897603260DFFB1FEF9F86D1F39F869" box="[270,285,1926,1949]" italics="true" pageId="18" pageNumber="419">E</emphasis>
.
<taxonomicName id="EEFDD192260DFFB1FEDDF86D1FF6F869" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[298,466,1926,1950]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="contrasticus" status="sp. nov." subGenus="Anomaloptilus">
(
<emphasis id="1B897603260DFFB1FEC5F86D1F65F869" box="[306,321,1926,1949]" italics="true" pageId="18" pageNumber="419">A</emphasis>
.)
<emphasis id="1B897603260DFFB1FEA1F86D1FF6F869" box="[342,466,1926,1949]" italics="true" pageId="18" pageNumber="419">contrasticus</emphasis>
</taxonomicName>
,
<emphasis id="1B897603260DFFB1FE29F86D1C33F86A" bold="true" box="[478,535,1926,1950]" pageId="18" pageNumber="419">
<taxonomicNameLabel id="00BACB78260DFFB1FE29F86D1C33F86A" box="[478,535,1926,1950]" pageId="18" pageNumber="419" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male;
<emphasis id="1B897603260DFFB1FD96F86E1C5EF869" bold="true" box="[609,634,1925,1949]" pageId="18" pageNumber="419">52</emphasis>
,
<emphasis id="1B897603260DFFB1FD71F86D1CB1F869" box="[646,661,1926,1949]" italics="true" pageId="18" pageNumber="419">E</emphasis>
.
<taxonomicName id="EEFDD192260DFFB1FD55F86D1D6EF869" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[674,842,1926,1950]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="contrasticus" status="sp. nov." subGenus="Anomaloptilus">
(
<emphasis id="1B897603260DFFB1FD5DF86D1C9DF869" box="[682,697,1926,1949]" italics="true" pageId="18" pageNumber="419">A</emphasis>
.)
<emphasis id="1B897603260DFFB1FD39F86D1D6EF869" box="[718,842,1926,1949]" italics="true" pageId="18" pageNumber="419">contrasticus</emphasis>
</taxonomicName>
,
<emphasis id="1B897603260DFFB1FCA1F86D1DABF86A" bold="true" box="[854,911,1926,1950]" pageId="18" pageNumber="419">
<taxonomicNameLabel id="00BACB78260DFFB1FCA1F86D1DABF86A" box="[854,911,1926,1950]" pageId="18" pageNumber="419" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, female;
<emphasis id="1B897603260DFFB1FC1BF86E1A21F869" bold="true" box="[1004,1029,1925,1949]" pageId="18" pageNumber="419">53</emphasis>
,
<emphasis id="1B897603260DFFB1FBE5F86D1A05F869" box="[1042,1057,1926,1949]" italics="true" pageId="18" pageNumber="419">E</emphasis>
.
<taxonomicName id="EEFDD192260DFFB1FBDAF86D1A82F869" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[1069,1190,1925,1950]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="hauseri" status="sp. nov." subGenus="Anomaloptilus">
(
<emphasis id="1B897603260DFFB1FBC1F86D1A61F869" box="[1078,1093,1926,1949]" italics="true" pageId="18" pageNumber="419">A</emphasis>
.)
<emphasis id="1B897603260DFFB1FBADF86E1A82F869" box="[1114,1190,1925,1949]" italics="true" pageId="18" pageNumber="419">hauseri</emphasis>
</taxonomicName>
,
<emphasis id="1B897603260DFFB1FB45F86D1ACFF86A" bold="true" box="[1202,1259,1926,1950]" pageId="18" pageNumber="419">
<taxonomicNameLabel id="00BACB78260DFFB1FB45F86D1ACFF86A" box="[1202,1259,1926,1950]" pageId="18" pageNumber="419" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male;
<emphasis id="1B897603260DFFB1FAC2F86E1B6AF869" bold="true" box="[1333,1358,1925,1949]" pageId="18" pageNumber="419">54</emphasis>
,
<emphasis id="1B897603260DFFB1FAADF86D1B4DF869" box="[1370,1385,1926,1949]" italics="true" pageId="18" pageNumber="419">E</emphasis>
.
<taxonomicName id="EEFDD192260DFFB1FA81F86D1EC7F835" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="hauseri" status="sp. nov." subGenus="Anomaloptilus">
(
<emphasis id="1B897603260DFFB1FA89F86D1BA9F869" box="[1406,1421,1926,1949]" italics="true" pageId="18" pageNumber="419">A</emphasis>
.)
<emphasis id="1B897603260DFFB1FF60F8421EC7F835" box="[151,227,1961,1985]" italics="true" pageId="18" pageNumber="419">hauseri</emphasis>
</taxonomicName>
,
<emphasis id="1B897603260DFFB1FF07F8411F0EF836" bold="true" box="[240,298,1962,1986]" pageId="18" pageNumber="419">
<taxonomicNameLabel id="00BACB78260DFFB1FF07F8411F0EF836" box="[240,298,1962,1986]" pageId="18" pageNumber="419" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, female;
<emphasis id="1B897603260DFFB1FE7DF8411F87F836" bold="true" box="[394,419,1962,1986]" pageId="18" pageNumber="419">55</emphasis>
,
<emphasis id="1B897603260DFFB1FE46F8411FE4F835" box="[433,448,1962,1985]" italics="true" pageId="18" pageNumber="419">E</emphasis>
.
<taxonomicName id="EEFDD192260DFFB1FE3AF8411C1CF835" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[461,568,1962,1986]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="irwini" status="sp. nov." subGenus="Anomaloptilus">
(
<emphasis id="1B897603260DFFB1FE21F8411FC1F835" box="[470,485,1962,1985]" italics="true" pageId="18" pageNumber="419">A</emphasis>
.)
<emphasis id="1B897603260DFFB1FE0CF8411C1CF835" box="[507,568,1962,1985]" italics="true" pageId="18" pageNumber="419">irwini</emphasis>
</taxonomicName>
,
<emphasis id="1B897603260DFFB1FDB2F8411C5BF836" bold="true" box="[581,639,1962,1986]" pageId="18" pageNumber="419">
<taxonomicNameLabel id="00BACB78260DFFB1FDB2F8411C5BF836" box="[581,639,1962,1986]" pageId="18" pageNumber="419" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male;
<emphasis id="1B897603260DFFB1FD3CF8421CC0F835" bold="true" box="[715,740,1961,1985]" pageId="18" pageNumber="419">56</emphasis>
,
<emphasis id="1B897603260DFFB1FD06F8411D24F835" box="[753,768,1962,1985]" italics="true" pageId="18" pageNumber="419">E</emphasis>
.
<taxonomicName id="EEFDD192260DFFB1FCF9F8411D5DF835" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[782,889,1962,1986]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="irwini" status="sp. nov." subGenus="Anomaloptilus">
(
<emphasis id="1B897603260DFFB1FCE0F8411D02F835" box="[791,806,1962,1985]" italics="true" pageId="18" pageNumber="419">A</emphasis>
.)
<emphasis id="1B897603260DFFB1FCCBF8411D5DF835" box="[828,889,1962,1985]" italics="true" pageId="18" pageNumber="419">irwini</emphasis>
</taxonomicName>
,
<emphasis id="1B897603260DFFB1FC72F8411D9BF836" bold="true" box="[901,959,1962,1986]" pageId="18" pageNumber="419">
<taxonomicNameLabel id="00BACB78260DFFB1FC72F8411D9BF836" box="[901,959,1962,1986]" pageId="18" pageNumber="419" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, female;
<emphasis id="1B897603260DFFB1FBE8F8411A1CF836" bold="true" box="[1055,1080,1962,1986]" pageId="18" pageNumber="419">57</emphasis>
,
<emphasis id="1B897603260DFFB1FBB1F8411A71F835" box="[1094,1109,1962,1985]" italics="true" pageId="18" pageNumber="419">E</emphasis>
.
<taxonomicName id="EEFDD192260DFFB1FB95F8411AE9F835" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[1122,1229,1962,1986]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="nanus" status="sp. nov." subGenus="Anomaloptilus">
(
<emphasis id="1B897603260DFFB1FB9CF8411A5EF835" box="[1131,1146,1962,1985]" italics="true" pageId="18" pageNumber="419">A</emphasis>
.)
<emphasis id="1B897603260DFFB1FB67F8411AE9F835" box="[1168,1229,1962,1985]" italics="true" pageId="18" pageNumber="419">nanus</emphasis>
</taxonomicName>
,
<emphasis id="1B897603260DFFB1FB2DF8411B30F836" bold="true" box="[1242,1300,1962,1986]" pageId="18" pageNumber="419">
<taxonomicNameLabel id="00BACB78260DFFB1FB2DF8411B30F836" box="[1242,1300,1962,1986]" pageId="18" pageNumber="419" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male;
<emphasis id="1B897603260DFFB1FA97F8421B5DF835" bold="true" box="[1376,1401,1961,1985]" pageId="18" pageNumber="419">58</emphasis>
,
<emphasis id="1B897603260DFFB1FA71F8411BB1F835" box="[1414,1429,1962,1985]" italics="true" pageId="18" pageNumber="419">E</emphasis>
.
<taxonomicName id="EEFDD192260DFFB1FF60F8251F26F811" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[151,258,1998,2022]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="regius" status="sp. nov." subGenus="Anomaloptilus">
(
<emphasis id="1B897603260DFFB1FF57F8251E8BF811" box="[160,175,1998,2021]" italics="true" pageId="18" pageNumber="419">A</emphasis>
.)
<emphasis id="1B897603260DFFB1FF33F8251F26F811" box="[196,258,1998,2021]" italics="true" pageId="18" pageNumber="419">regius</emphasis>
</taxonomicName>
,
<emphasis id="1B897603260DFFB1FEF8F8251F6CF812" bold="true" box="[271,328,1998,2022]" pageId="18" pageNumber="419">
<taxonomicNameLabel id="00BACB78260DFFB1FEF8F8251F6CF812" box="[271,328,1998,2022]" pageId="18" pageNumber="419" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male.
</paragraph>
</caption>
<caption id="7D82FA99260CFFB0FF60FD5D1ACEFD06" ID-DOI="http://doi.org/10.5281/zenodo.7471774" ID-Zenodo-Dep="7471774" httpUri="https://zenodo.org/record/7471774/files/figure.png" pageId="19" pageNumber="420" startId="19.[151,264,694,718]" targetBox="[152,1435,181,670]" targetPageId="19">
<paragraph id="2942AA11260CFFB0FF60FD5D1ACEFD06" blockId="19.[151,1436,694,754]" pageId="19" pageNumber="420">
<emphasis id="1B897603260CFFB0FF60FD5D1F69FD3A" bold="true" box="[151,333,694,718]" pageId="19" pageNumber="420">FIGURES 59–62</emphasis>
. Malagasy
<taxonomicName id="EEFDD192260CFFB0FE37FD5D1CD2FD3A" box="[448,758,694,718]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="420" phylum="Arthropoda" rank="subGenus" subGenus="Anomaloptilus">
<emphasis id="1B897603260CFFB0FE37FD5D1C6EFD3A" box="[448,586,694,718]" italics="true" pageId="19" pageNumber="420">Empidideicus</emphasis>
(
<emphasis id="1B897603260CFFB0FDAFFD5D1CC8FD3A" box="[600,748,694,718]" italics="true" pageId="19" pageNumber="420">Anomaloptilus</emphasis>
)
</taxonomicName>
, abdomens, dorsal view.
<emphasis id="1B897603260CFFB0FC05FD5D1A2FFD3A" bold="true" box="[1010,1035,694,718]" pageId="19" pageNumber="420">59</emphasis>
,
<taxonomicName id="EEFDD192260CFFB0FBE0FD5C1B7FFD3A" authority="Evenhuis" authorityName="Evenhuis" box="[1047,1371,694,718]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="403" phylum="Arthropoda" rank="species" species="rhinoclypeatus" subGenus="Anomaloptilus">
<emphasis id="1B897603260CFFB0FBE0FD5C1A02FD3A" box="[1047,1062,695,718]" italics="true" pageId="19" pageNumber="420">E</emphasis>
. (
<emphasis id="1B897603260CFFB0FBCCFD5D1A6EFD39" box="[1083,1098,694,717]" italics="true" pageId="19" pageNumber="420">A</emphasis>
.)
<emphasis id="1B897603260CFFB0FBA9FD5D1AD0FD3A" box="[1118,1268,694,718]" italics="true" pageId="19" pageNumber="420">rhinoclypeatus</emphasis>
Evenhuis
</taxonomicName>
, male;
<emphasis id="1B897603260CFFB0FF60FD311E94FD06" bold="true" box="[151,176,730,754]" pageId="19" pageNumber="420">60</emphasis>
,
<taxonomicName id="EEFDD192260CFFB0FF4AFD301C20FD06" authority="Evenhuis" authorityName="Evenhuis" box="[189,516,730,754]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="403" phylum="Arthropoda" rank="species" species="rhinoclypeatus" subGenus="Anomaloptilus">
<emphasis id="1B897603260CFFB0FF4AFD301EE8FD06" box="[189,204,731,754]" italics="true" pageId="19" pageNumber="420">E</emphasis>
. (
<emphasis id="1B897603260CFFB0FF15FD311ED5FD05" box="[226,241,730,753]" italics="true" pageId="19" pageNumber="420">A</emphasis>
.)
<emphasis id="1B897603260CFFB0FEF1FD311FB8FD06" box="[262,412,730,754]" italics="true" pageId="19" pageNumber="420">rhinoclypeatus</emphasis>
Evenhuis
</taxonomicName>
, female;
<emphasis id="1B897603260CFFB0FD97FD311C5DFD06" bold="true" box="[608,633,730,754]" pageId="19" pageNumber="420">61</emphasis>
,
<emphasis id="1B897603260CFFB0FD71FD301CB1FD06" box="[646,661,731,754]" italics="true" pageId="19" pageNumber="420">E</emphasis>
.
<taxonomicName id="EEFDD192260CFFB0FD55FD311D29FD06" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[674,781,730,754]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="420" phylum="Arthropoda" rank="species" species="rinhai" status="sp. nov." subGenus="Anomaloptilus">
(
<emphasis id="1B897603260CFFB0FD5CFD311C9EFD05" box="[683,698,730,753]" italics="true" pageId="19" pageNumber="420">A</emphasis>
.)
<emphasis id="1B897603260CFFB0FD38FD311D29FD06" box="[719,781,730,754]" italics="true" pageId="19" pageNumber="420">rinhai</emphasis>
</taxonomicName>
,
<emphasis id="1B897603260CFFB0FCEDFD311D77FD06" bold="true" box="[794,851,730,754]" pageId="19" pageNumber="420">
<taxonomicNameLabel id="00BACB78260CFFB0FCEDFD311D77FD06" box="[794,851,730,754]" pageId="19" pageNumber="420" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male;
<emphasis id="1B897603260CFFB0FC69FD311D93FD06" bold="true" box="[926,951,730,754]" pageId="19" pageNumber="420">62</emphasis>
,
<emphasis id="1B897603260CFFB0FC33FD301DF7FD06" box="[964,979,731,754]" italics="true" pageId="19" pageNumber="420">E</emphasis>
.
<taxonomicName id="EEFDD192260CFFB0FC17FD311A6FFD06" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[992,1099,730,754]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="420" phylum="Arthropoda" rank="species" species="rinhai" status="sp. nov." subGenus="Anomaloptilus">
(
<emphasis id="1B897603260CFFB0FC1EFD311DDCFD05" box="[1001,1016,730,753]" italics="true" pageId="19" pageNumber="420">A</emphasis>
.)
<emphasis id="1B897603260CFFB0FBFAFD311A6FFD06" box="[1037,1099,730,754]" italics="true" pageId="19" pageNumber="420">rinhai</emphasis>
</taxonomicName>
,
<emphasis id="1B897603260CFFB0FBAFFD311AB5FD06" bold="true" box="[1112,1169,730,754]" pageId="19" pageNumber="420">
<taxonomicNameLabel id="00BACB78260CFFB0FBAFFD311AB5FD06" box="[1112,1169,730,754]" pageId="19" pageNumber="420" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, female.
</paragraph>
</caption>
<caption id="7D82FA99260CFFB0FF6FF9D61C43F911" ID-DOI="http://doi.org/10.5281/zenodo.7471778" ID-Zenodo-Dep="7471778" httpUri="https://zenodo.org/record/7471778/files/figure.png" pageId="19" pageNumber="420" startId="19.[152,270,1597,1621]" targetBox="[156,1432,784,1567]" targetPageId="19">
<paragraph id="2942AA11260CFFB0FF6FF9D61C43F911" blockId="19.[151,1437,1597,1765]" pageId="19" pageNumber="420">
<emphasis id="1B897603260CFFB0FF6FF9D61F76F9A1" bold="true" box="[152,338,1597,1621]" pageId="19" pageNumber="420">FIGURES 63–64</emphasis>
.
<taxonomicName id="EEFDD192260CFFB0FEA9F9D61D56F9A1" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[350,882,1597,1621]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="420" phylum="Arthropoda" rank="species" species="andriambahomananii" status="sp. nov." subGenus="Anomaloptilus">
<emphasis id="1B897603260CFFB0FEA9F9D61FCCF9A1" box="[350,488,1597,1621]" italics="true" pageId="19" pageNumber="420">Empidideicus</emphasis>
(
<emphasis id="1B897603260CFFB0FE02F9D61CADF9A1" box="[501,649,1597,1621]" italics="true" pageId="19" pageNumber="420">Anomaloptilus</emphasis>
)
<emphasis id="1B897603260CFFB0FD61F9D61D56F9A1" box="[662,882,1597,1621]" italics="true" pageId="19" pageNumber="420">andriambahomananii</emphasis>
</taxonomicName>
,
<emphasis id="1B897603260CFFB0FC8AF9D61D91F9A1" bold="true" box="[893,949,1597,1621]" pageId="19" pageNumber="420">
<taxonomicNameLabel id="00BACB78260CFFB0FC8AF9D61D91F9A1" box="[893,949,1597,1621]" pageId="19" pageNumber="420" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, genitalia.
<emphasis id="1B897603260CFFB0FBD4F9D61A18F9A1" bold="true" box="[1059,1084,1597,1621]" pageId="19" pageNumber="420">63</emphasis>
, male,
<emphasis id="1B897603260CFFB0FB73F9D61AB2F9A1" bold="true" box="[1156,1174,1597,1621]" pageId="19" pageNumber="420">A</emphasis>
. gonocoxae, ventral view,
<emphasis id="1B897603260CFFB0FF60F98A1E8CF98D" bold="true" box="[151,168,1633,1657]" pageId="19" pageNumber="420">B</emphasis>
, Epandrium, dorsal view,
<emphasis id="1B897603260CFFB0FE4BF98A1FEAF98D" bold="true" box="[444,462,1633,1657]" pageId="19" pageNumber="420">C</emphasis>
, phallic complex, dorsal view;
<emphasis id="1B897603260CFFB0FCEFF98A1D15F98D" bold="true" box="[792,817,1633,1657]" pageId="19" pageNumber="420">64</emphasis>
, female. Abbreviations: ad = apical spermathecal duct; adb = aedeagal bulb; aed = aedeagus; ap = aedeagal apodeme; cer = cercus; ep = epandrium; fur = furca [= genital fork]; gcx = gonocoxa; gst = gonostylus; hyp = hypandrium; lat = lateral arms of parameral sheath; par = parameral sheath; pmp = sperm pump; ram= lateral rami; spm = spermatheca.
</paragraph>
</caption>
<paragraph id="2942AA11260CFFB0FF30F8FF1D9EF849" blockId="19.[151,1436,1811,1982]" pageId="19" pageNumber="420">
<emphasis id="1B897603260CFFB0FF30F8FF1F25F8D9" box="[199,257,1812,1837]" italics="true" pageId="19" pageNumber="420">Wing</emphasis>
(
<figureCitation id="B1C6B694260CFFB0FEE5F8F81F4DF8DA" box="[274,361,1811,1838]" captionStart="FIGURES 40–47" captionStartId="15.[151,264,1607,1631]" captionTargetBox="[170,1416,309,1582]" captionTargetId="figure-81@15.[170,1416,309,1584]" captionTargetPageId="15" captionText="FIGURES 40–47. Malagasy Empidideicus (Anomaloptilus), wings. 40, E. (A.) andriambahomananii, n. sp.; 41, E. (A.) contrasticus, n. sp.; 42, E. (A.) hauseri, n. sp.; 43, E. (A.) irwini, n. sp.; 44, E. (A.) nanus, n. sp.; 45, E. (A.) regius, n. sp.; 46, E. (A.) rhinoclypeatus Evenhuis; 47, E. (A.) rinhai, n. sp." figureDoi="http://doi.org/10.5281/zenodo.7471764" httpUri="https://zenodo.org/record/7471764/files/figure.png" pageId="19" pageNumber="420">Fig. 47</figureCitation>
). Elongate, length 2.7× width; hyaline; veins brown; costa ends just beyond end of R
<subScript id="B579A854260CFFB0FAB6F8CD1B7FF8C0" attach="left" box="[1345,1371,1830,1844]" fontSize="6" pageId="19" pageNumber="420">4+5</subScript>
; vein Sc incomplete; Rs evanescent at connection with R
<subScript id="B579A854260CFFB0FD20F8A11CFBF8AC" attach="left" box="[727,735,1866,1880]" fontSize="6" pageId="19" pageNumber="420">1</subScript>
; R
<subScript id="B579A854260CFFB0FCF6F8A11D3FF8AC" attach="left" box="[769,795,1866,1880]" fontSize="6" pageId="19" pageNumber="420">4+5</subScript>
slightly bowed; vein M
<subScript id="B579A854260CFFB0FBD3F8A11A08F8AC" attach="left" box="[1060,1068,1866,1880]" fontSize="6" pageId="19" pageNumber="420">1</subScript>
curved to wing margin; M
<subScript id="B579A854260CFFB0FAACF8A11B47F8AC" attach="left" box="[1371,1379,1866,1880]" fontSize="6" pageId="19" pageNumber="420">2</subScript>
very slightly curved to wing margin; cell dm closed by apical crossvein; crossvein subequal in width to crossvein r-m; A
<subScript id="B579A854260CFFB0FA63F8851BB8F888" attach="left" box="[1428,1436,1902,1916]" fontSize="6" pageId="19" pageNumber="420">1</subScript>
effaced, straight to wing margin; anal cell narrowly open at wing margin; fringe of hair on posterior margin of wing well shortest at wing tip becoming longer and more sparse toward base.
</paragraph>
<caption id="7D82FA99260BFFB7FF6FFC041A1EFBDF" ID-DOI="http://doi.org/10.5281/zenodo.7471780" ID-Zenodo-Dep="7471780" httpUri="https://zenodo.org/record/7471780/files/figure.png" pageId="20" pageNumber="421" startId="20.[152,270,1007,1031]" targetBox="[156,1433,181,978]" targetPageId="20">
<paragraph id="2942AA11260BFFB7FF6FFC041A1EFBDF" blockId="20.[151,1437,1007,1067]" pageId="20" pageNumber="421">
<emphasis id="1B897603260BFFB7FF6FFC041F70FBF3" bold="true" box="[152,340,1007,1031]" pageId="20" pageNumber="421">FIGURES 65–66</emphasis>
.
<taxonomicName id="EEFDD192260BFFB7FE96FC041D3CFBF3" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[353,792,1007,1031]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="species" species="contrasticus" status="sp. nov." subGenus="Anomaloptilus">
<emphasis id="1B897603260BFFB7FE96FC041FCFFBF3" box="[353,491,1007,1031]" italics="true" pageId="20" pageNumber="421">Empidideicus</emphasis>
(
<emphasis id="1B897603260BFFB7FE0DFC041CAAFBF3" box="[506,654,1007,1031]" italics="true" pageId="20" pageNumber="421">Anomaloptilus</emphasis>
)
<emphasis id="1B897603260BFFB7FD6BFC1B1D3CFBF3" box="[668,792,1008,1031]" italics="true" pageId="20" pageNumber="421">contrasticus</emphasis>
</taxonomicName>
,
<emphasis id="1B897603260BFFB7FCD2FC041D7BFBF3" bold="true" box="[805,863,1007,1031]" pageId="20" pageNumber="421">
<taxonomicNameLabel id="00BACB78260BFFB7FCD2FC041D7BFBF3" box="[805,863,1007,1031]" pageId="20" pageNumber="421" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, genitalia.
<emphasis id="1B897603260BFFB7FC38FC041DCCFBF3" bold="true" box="[975,1000,1007,1031]" pageId="20" pageNumber="421">65</emphasis>
, male,
<emphasis id="1B897603260BFFB7FBC3FC041A62FBF3" bold="true" box="[1076,1094,1007,1031]" pageId="20" pageNumber="421">A</emphasis>
, lateral view,
<emphasis id="1B897603260BFFB7FB22FC041AC8FBF3" bold="true" box="[1237,1260,1007,1031]" pageId="20" pageNumber="421">B,</emphasis>
phallic complex, dorsal view,
<emphasis id="1B897603260BFFB7FEEFFBF81F0EFBDF" bold="true" box="[280,298,1043,1067]" pageId="20" pageNumber="421">C</emphasis>
, gonocoxae, ventral view;
<emphasis id="1B897603260BFFB7FDCBFBF81C71FBDF" bold="true" box="[572,597,1043,1067]" pageId="20" pageNumber="421">66</emphasis>
, female. Abbreviation: ap = aedeagal apodeme.
</paragraph>
</caption>
<caption id="7D82FA99260BFFB7FF6FF9041CC9F8BB" ID-DOI="http://doi.org/10.5281/zenodo.7471784" ID-Zenodo-Dep="7471784" httpUri="https://zenodo.org/record/7471784/files/figure.png" pageId="20" pageNumber="421" startId="20.[152,270,1775,1799]" targetBox="[161,1430,1091,1747]" targetPageId="20">
<paragraph id="2942AA11260BFFB7FF6FF9041CC9F8BB" blockId="20.[151,1437,1775,1871]" pageId="20" pageNumber="421">
<emphasis id="1B897603260BFFB7FF6FF9041F72F8F3" bold="true" box="[152,342,1775,1799]" pageId="20" pageNumber="421">FIGURES 67–68</emphasis>
.
<taxonomicName id="EEFDD192260BFFB7FE93F9041CCBF8F3" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[356,751,1775,1799]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="species" species="hauseri" status="sp. nov." subGenus="Anomaloptilus">
<emphasis id="1B897603260BFFB7FE93F9041FCAF8F3" box="[356,494,1775,1799]" italics="true" pageId="20" pageNumber="421">Empidideicus</emphasis>
(
<emphasis id="1B897603260BFFB7FE09F9041CB6F8F3" box="[510,658,1775,1799]" italics="true" pageId="20" pageNumber="421">Anomaloptilus</emphasis>
)
<emphasis id="1B897603260BFFB7FD54F9041CCBF8F3" box="[675,751,1775,1799]" italics="true" pageId="20" pageNumber="421">hauseri</emphasis>
</taxonomicName>
,
<emphasis id="1B897603260BFFB7FD0AF9041D1CF8F3" bold="true" box="[765,824,1775,1799]" pageId="20" pageNumber="421">
<taxonomicNameLabel id="00BACB78260BFFB7FD0AF9041D1CF8F3" box="[765,824,1775,1799]" pageId="20" pageNumber="421" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, genitalia.
<emphasis id="1B897603260BFFB7FC5BF9041DE1F8F3" bold="true" box="[940,965,1775,1799]" pageId="20" pageNumber="421">67</emphasis>
, male,
<emphasis id="1B897603260BFFB7FBE3F9041A02F8F3" bold="true" box="[1044,1062,1775,1799]" pageId="20" pageNumber="421">A</emphasis>
, lateral view,
<emphasis id="1B897603260BFFB7FB4DF9041AEFF8F3" bold="true" box="[1210,1227,1775,1799]" pageId="20" pageNumber="421">B</emphasis>
, gonocoxae, ventral view,
<emphasis id="1B897603260BFFB7FF23F8F81EC2F8DF" bold="true" box="[212,230,1811,1835]" pageId="20" pageNumber="421">C</emphasis>
, phallic complex, dorsal view,
<emphasis id="1B897603260BFFB7FDDFF8F81C1EF8DF" bold="true" box="[552,570,1811,1835]" pageId="20" pageNumber="421">D</emphasis>
, epiphallic membrane, dorsal view,
<emphasis id="1B897603260BFFB7FC5AF8F81D9AF8DF" bold="true" box="[941,958,1811,1835]" pageId="20" pageNumber="421">E</emphasis>
, gonostylus, detail;
<emphasis id="1B897603260BFFB7FB79F8F81A83F8DF" bold="true" box="[1166,1191,1811,1835]" pageId="20" pageNumber="421">68</emphasis>
, female. Abbreviations: cer = cercus; mem= membrane associated with epandrium.
</paragraph>
</caption>
<paragraph id="2942AA11260BFFB7FF30F8961EF9F82B" blockId="20.[151,1436,1917,2015]" pageId="20" pageNumber="421">
<emphasis id="1B897603260BFFB7FF30F8961F15F863" box="[199,305,1917,1943]" italics="true" pageId="20" pageNumber="421">Abdomen</emphasis>
(
<figureCitation id="B1C6B694260BFFB7FEC8F8961FB4F863" box="[319,400,1917,1943]" captionStart="FIGURES 59–62" captionStartId="19.[151,264,694,718]" captionTargetBox="[152,1435,181,670]" captionTargetId="figure-17@19.[152,1435,181,670]" captionTargetPageId="19" captionText="FIGURES 59–62. Malagasy Empidideicus (Anomaloptilus), abdomens, dorsal view. 59, E. (A.) rhinoclypeatus Evenhuis, male; 60, E. (A.) rhinoclypeatus Evenhuis, female; 61, E. (A.) rinhai, n. sp., male; 62, E. (A.) rinhai, n. sp., female." figureDoi="http://doi.org/10.5281/zenodo.7471774" httpUri="https://zenodo.org/record/7471774/files/figure.png" pageId="20" pageNumber="421">Fig. 61</figureCitation>
). Tergites dark brown to black, narrowly white along posterior margin of tergites and laterally; tergites II and III with pale brown spots dorsolaterally; tergites with sparsely scattered minute brown hairs; sternites white.
</paragraph>
<paragraph id="2942AA11260AFFB6FF30FF7C1B0DFE7D" blockId="21.[151,1437,151,393]" pageId="21" pageNumber="422">
<emphasis id="1B897603260AFFB6FF30FF7C1F15FF45" box="[199,305,151,177]" italics="true" pageId="21" pageNumber="422">Genitalia</emphasis>
(
<figureCitation id="B1C6B694260AFFB6FEB5FF7C1FBCFF45" box="[322,408,151,177]" captionStart="FIGURES 77" captionStartId="23.[152,270,1257,1281]" captionTargetBox="[160,1433,187,1228]" captionTargetId="figure-17@23.[151,1436,181,1233]" captionTargetPageId="23" captionText="FIGURES 77. Empidideicus (Anomaloptilus) rinhai, n. sp., male genitalia, A, lateral view, B, gonocoxae, ventral view, C, phallic complex, dorsal view, D, apex of dorsoventral epandrial process, detail, E, gonostylus, detail." figureDoi="http://doi.org/10.5281/zenodo.7471798" httpUri="https://zenodo.org/record/7471798/files/figure.png" pageId="21" pageNumber="422">Fig. 77</figureCitation>
). Epandrium in lateral view subquadrate with long thin dorsoventral process, process well sclerotized and slightly clubbed apically (
<figureCitation id="B1C6B694260AFFB6FD85FF501CF8FF21" box="[626,732,187,213]" captionStart="FIGURES 77" captionStartId="23.[152,270,1257,1281]" captionTargetBox="[160,1433,187,1228]" captionTargetId="figure-17@23.[151,1436,181,1233]" captionTargetPageId="23" captionText="FIGURES 77. Empidideicus (Anomaloptilus) rinhai, n. sp., male genitalia, A, lateral view, B, gonocoxae, ventral view, C, phallic complex, dorsal view, D, apex of dorsoventral epandrial process, detail, E, gonostylus, detail." figureDoi="http://doi.org/10.5281/zenodo.7471798" httpUri="https://zenodo.org/record/7471798/files/figure.png" pageId="21" pageNumber="422">Fig. 77A</figureCitation>
), apex with microscopic hairs (
<figureCitation id="B1C6B694260AFFB6FBB5FF501A89FF21" box="[1090,1197,187,213]" captionStart="FIGURES 77" captionStartId="23.[152,270,1257,1281]" captionTargetBox="[160,1433,187,1228]" captionTargetId="figure-17@23.[151,1436,181,1233]" captionTargetPageId="23" captionText="FIGURES 77. Empidideicus (Anomaloptilus) rinhai, n. sp., male genitalia, A, lateral view, B, gonocoxae, ventral view, C, phallic complex, dorsal view, D, apex of dorsoventral epandrial process, detail, E, gonostylus, detail." figureDoi="http://doi.org/10.5281/zenodo.7471798" httpUri="https://zenodo.org/record/7471798/files/figure.png" pageId="21" pageNumber="422">Fig. 77D</figureCitation>
); gonocoxa in lateral view conical, tapering to apex; in ventral view with basal sclerotize band and large seta medially on each lobe (
<figureCitation id="B1C6B694260AFFB6FA98FF341EEFFEE9" captionStart="FIGURES 77" captionStartId="23.[152,270,1257,1281]" captionTargetBox="[160,1433,187,1228]" captionTargetId="figure-17@23.[151,1436,181,1233]" captionTargetPageId="23" captionText="FIGURES 77. Empidideicus (Anomaloptilus) rinhai, n. sp., male genitalia, A, lateral view, B, gonocoxae, ventral view, C, phallic complex, dorsal view, D, apex of dorsoventral epandrial process, detail, E, gonostylus, detail." figureDoi="http://doi.org/10.5281/zenodo.7471798" httpUri="https://zenodo.org/record/7471798/files/figure.png" pageId="21" pageNumber="422">Fig. 77A</figureCitation>
); gonostylus subtrapezoidal with rounded medial process, apex of medial process with microscopic hairs (
<figureCitation id="B1C6B694260AFFB6FA98FEE81EECFEB5" captionStart="FIGURES 77" captionStartId="23.[152,270,1257,1281]" captionTargetBox="[160,1433,187,1228]" captionTargetId="figure-17@23.[151,1436,181,1233]" captionTargetPageId="23" captionText="FIGURES 77. Empidideicus (Anomaloptilus) rinhai, n. sp., male genitalia, A, lateral view, B, gonocoxae, ventral view, C, phallic complex, dorsal view, D, apex of dorsoventral epandrial process, detail, E, gonostylus, detail." figureDoi="http://doi.org/10.5281/zenodo.7471798" httpUri="https://zenodo.org/record/7471798/files/figure.png" pageId="21" pageNumber="422">Fig. 77E</figureCitation>
); lateral arms of parameral sheath large, flared apically, not as long as aedeagal apodeme; aedeagal apodeme peanut-shaped, truncate ventrally, with foliate lateral rami; aedeagal bulb large, in dorsal view truncate apically; distal portion of parameral sheath cylindrical with subapical collar-like sheath over aedeagus (
<figureCitation id="B1C6B694260AFFB6FB42FE841B39FE7D" box="[1205,1309,367,393]" captionStart="FIGURES 77" captionStartId="23.[152,270,1257,1281]" captionTargetBox="[160,1433,187,1228]" captionTargetId="figure-17@23.[151,1436,181,1233]" captionTargetPageId="23" captionText="FIGURES 77. Empidideicus (Anomaloptilus) rinhai, n. sp., male genitalia, A, lateral view, B, gonocoxae, ventral view, C, phallic complex, dorsal view, D, apex of dorsoventral epandrial process, detail, E, gonostylus, detail." figureDoi="http://doi.org/10.5281/zenodo.7471798" httpUri="https://zenodo.org/record/7471798/files/figure.png" pageId="21" pageNumber="422">Fig. 77C</figureCitation>
).
</paragraph>
<caption id="7D82FA99260AFFB6FF6FFC131CFFFBC0" ID-DOI="http://doi.org/10.5281/zenodo.7471786" ID-Zenodo-Dep="7471786" httpUri="https://zenodo.org/record/7471786/files/figure.png" pageId="21" pageNumber="422" startId="21.[152,270,1016,1040]" targetBox="[156,1429,422,990]" targetPageId="21">
<paragraph id="2942AA11260AFFB6FF6FFC131CFFFBC0" blockId="21.[151,1437,1016,1076]" pageId="21" pageNumber="422">
<emphasis id="1B897603260AFFB6FF6FFC131F77FBE4" bold="true" box="[152,339,1016,1040]" pageId="21" pageNumber="422">FIGURES 69–70</emphasis>
.
<taxonomicName id="EEFDD192260AFFB6FEA8FC131CF2FBE4" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[351,726,1016,1040]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="422" phylum="Arthropoda" rank="species" species="irwini" status="sp. nov." subGenus="Anomaloptilus">
<emphasis id="1B897603260AFFB6FEA8FC131FCDFBE4" box="[351,489,1016,1040]" italics="true" pageId="21" pageNumber="422">Empidideicus</emphasis>
(
<emphasis id="1B897603260AFFB6FE00FC131CAFFBE4" box="[503,651,1016,1040]" italics="true" pageId="21" pageNumber="422">Anomaloptilus</emphasis>
)
<emphasis id="1B897603260AFFB6FD6EFC121CF2FBE4" box="[665,726,1017,1040]" italics="true" pageId="21" pageNumber="422">irwini</emphasis>
</taxonomicName>
,
<emphasis id="1B897603260AFFB6FD15FC131D3FFBE4" bold="true" box="[738,795,1016,1040]" pageId="21" pageNumber="422">
<taxonomicNameLabel id="00BACB78260AFFB6FD15FC131D3FFBE4" box="[738,795,1016,1040]" pageId="21" pageNumber="422" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, genitalia.
<emphasis id="1B897603260AFFB6FC7DFC131D87FBE4" bold="true" box="[906,931,1016,1040]" pageId="21" pageNumber="422">69</emphasis>
, male,
<emphasis id="1B897603260AFFB6FC1AFC131DDBFBE4" bold="true" box="[1005,1023,1016,1040]" pageId="21" pageNumber="422">A</emphasis>
, gonocoxae, ventral view,
<emphasis id="1B897603260AFFB6FAFAFC131B3AFBE4" bold="true" box="[1293,1310,1016,1040]" pageId="21" pageNumber="422">B</emphasis>
, epandrium, dorsal view,
<emphasis id="1B897603260AFFB6FEEFFBF71F0EFBC0" bold="true" box="[280,298,1052,1076]" pageId="21" pageNumber="422">C</emphasis>
, phallic complex, dorsal view;
<emphasis id="1B897603260AFFB6FD9EFBF71CA6FBC0" bold="true" box="[617,642,1052,1076]" pageId="21" pageNumber="422">70</emphasis>
, female.
</paragraph>
</caption>
<caption id="7D82FA99260AFFB6FF6FF84F1C7EF815" ID-DOI="http://doi.org/10.5281/zenodo.7471790" ID-Zenodo-Dep="7471790" httpUri="https://zenodo.org/record/7471790/files/figure.png" pageId="21" pageNumber="422" startId="21.[152,270,1956,1980]" targetBox="[152,1436,1100,1933]" targetPageId="21">
<paragraph id="2942AA11260AFFB6FF6FF84F1C7EF815" blockId="21.[151,1436,1956,2017]" pageId="21" pageNumber="422">
<emphasis id="1B897603260AFFB6FF6FF84F1F77F849" bold="true" box="[152,339,1956,1981]" pageId="21" pageNumber="422">FIGURES 71–72</emphasis>
.
<taxonomicName id="EEFDD192260AFFB6FEA9F84E1CF0F849" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[350,724,1957,1981]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="422" phylum="Arthropoda" rank="species" species="nanus" status="sp. nov." subGenus="Anomaloptilus">
<emphasis id="1B897603260AFFB6FEA9F84E1FCCF849" box="[350,488,1957,1981]" italics="true" pageId="21" pageNumber="422">Empidideicus</emphasis>
(
<emphasis id="1B897603260AFFB6FE01F84E1CAEF849" box="[502,650,1957,1981]" italics="true" pageId="21" pageNumber="422">Anomaloptilus</emphasis>
)
<emphasis id="1B897603260AFFB6FD60F84D1CF0F849" box="[663,724,1958,1981]" italics="true" pageId="21" pageNumber="422">nanus</emphasis>
</taxonomicName>
,
<emphasis id="1B897603260AFFB6FD28F84E1D33F849" bold="true" box="[735,791,1957,1981]" pageId="21" pageNumber="422">
<taxonomicNameLabel id="00BACB78260AFFB6FD28F84E1D33F849" box="[735,791,1957,1981]" pageId="21" pageNumber="422" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, genitalia.
<emphasis id="1B897603260AFFB6FC71F84E1DBBF849" bold="true" box="[902,927,1957,1981]" pageId="21" pageNumber="422">71</emphasis>
, male,
<emphasis id="1B897603260AFFB6FC10F84E1DDDF849" bold="true" box="[999,1017,1957,1981]" pageId="21" pageNumber="422">A</emphasis>
, lateral view,
<emphasis id="1B897603260AFFB6FB72F84E1AB8F849" bold="true" box="[1157,1180,1957,1981]" pageId="21" pageNumber="422">B,</emphasis>
gonocoxae, ventral view,
<emphasis id="1B897603260AFFB6FF60F8221E8DF815" bold="true" box="[151,169,1993,2017]" pageId="21" pageNumber="422">C</emphasis>
, phallic complex, dorsal view;
<emphasis id="1B897603260AFFB6FE1FF8221C25F815" bold="true" box="[488,513,1993,2017]" pageId="21" pageNumber="422">72</emphasis>
, female.
</paragraph>
</caption>
<caption id="7D82FA992609FFB5FF6FFD001C93FCD3" ID-DOI="http://doi.org/10.5281/zenodo.7471792" ID-Zenodo-Dep="7471792" httpUri="https://zenodo.org/record/7471792/files/figure.png" pageId="22" pageNumber="423" startId="22.[152,270,747,771]" targetBox="[158,1432,187,717]" targetPageId="22">
<paragraph id="2942AA112609FFB5FF6FFD001C93FCD3" blockId="22.[151,1436,747,808]" pageId="22" pageNumber="423">
<emphasis id="1B8976032609FFB5FF6FFD001F76FCF7" bold="true" box="[152,338,747,771]" pageId="22" pageNumber="423">FIGURES 73–74</emphasis>
.
<taxonomicName id="EEFDD1922609FFB5FEA9FD001CF1FCF7" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[350,725,747,772]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="423" phylum="Arthropoda" rank="species" species="regius" status="sp. nov." subGenus="Anomaloptilus">
<emphasis id="1B8976032609FFB5FEA9FD001FCCFCF7" box="[350,488,747,771]" italics="true" pageId="22" pageNumber="423">Empidideicus</emphasis>
(
<emphasis id="1B8976032609FFB5FE02FD001CADFCF7" box="[501,649,747,771]" italics="true" pageId="22" pageNumber="423">Anomaloptilus</emphasis>
)
<emphasis id="1B8976032609FFB5FD60FD071CF1FCF7" box="[663,725,748,771]" italics="true" pageId="22" pageNumber="423">regius</emphasis>
</taxonomicName>
,
<emphasis id="1B8976032609FFB5FD16FD071D3DFCF0" bold="true" box="[737,793,748,772]" pageId="22" pageNumber="423">
<taxonomicNameLabel id="00BACB782609FFB5FD16FD071D3DFCF0" box="[737,793,748,772]" pageId="22" pageNumber="423" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, genitalia.
<emphasis id="1B8976032609FFB5FC70FD001D84FCF7" bold="true" box="[903,928,747,771]" pageId="22" pageNumber="423">73</emphasis>
, male,
<emphasis id="1B8976032609FFB5FC1FFD001DDEFCF7" bold="true" box="[1000,1018,747,771]" pageId="22" pageNumber="423">A</emphasis>
, lateral view,
<emphasis id="1B8976032609FFB5FB72FD071AB2FCF0" bold="true" box="[1157,1174,748,772]" pageId="22" pageNumber="423">B</emphasis>
, gonocoxae, ventral view,
<emphasis id="1B8976032609FFB5FF60FCE41E8DFCD3" bold="true" box="[151,169,783,807]" pageId="22" pageNumber="423">C</emphasis>
, phallic complex,
<emphasis id="1B8976032609FFB5FE92FCFB1F53FCDC" bold="true" box="[357,375,784,808]" pageId="22" pageNumber="423">D</emphasis>
, gonostylus, detail;
<emphasis id="1B8976032609FFB5FDB3FCE41C79FCD3" bold="true" box="[580,605,783,807]" pageId="22" pageNumber="423">74</emphasis>
, female.
</paragraph>
</caption>
<caption id="7D82FA992609FFB5FF6FFA7D1DF4FA26" ID-DOI="http://doi.org/10.5281/zenodo.7471796" ID-Zenodo-Dep="7471796" httpUri="https://zenodo.org/record/7471796/files/figure.png" pageId="22" pageNumber="423" startId="22.[152,270,1430,1454]" targetBox="[155,1432,838,1400]" targetPageId="22">
<paragraph id="2942AA112609FFB5FF6FFA7D1DF4FA26" blockId="22.[151,1436,1430,1490]" pageId="22" pageNumber="423">
<emphasis id="1B8976032609FFB5FF6FFA7D1F73FA5A" bold="true" box="[152,343,1430,1454]" pageId="22" pageNumber="423">FIGURES 75–76</emphasis>
.
<taxonomicName id="EEFDD1922609FFB5FE90FA7D1D8FFA5A" authority="Evenhuis" authorityName="Evenhuis" box="[359,939,1430,1454]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="423" phylum="Arthropoda" rank="species" species="rhinoclypeatus" subGenus="Anomaloptilus">
<emphasis id="1B8976032609FFB5FE90FA7D1FD5FA5A" box="[359,497,1430,1454]" italics="true" pageId="22" pageNumber="423">Empidideicus</emphasis>
(
<emphasis id="1B8976032609FFB5FDF3FA7D1CBCFA5A" box="[516,664,1430,1454]" italics="true" pageId="22" pageNumber="423">Anomaloptilus</emphasis>
)
<emphasis id="1B8976032609FFB5FD5DFA7D1D64FA5A" box="[682,832,1430,1454]" italics="true" pageId="22" pageNumber="423">rhinoclypeatus</emphasis>
Evenhuis
</taxonomicName>
, genitalia.
<emphasis id="1B8976032609FFB5FBD7FA7D1A1DFA5A" bold="true" box="[1056,1081,1430,1454]" pageId="22" pageNumber="423">75</emphasis>
, male,
<emphasis id="1B8976032609FFB5FB7CFA7D1AB9FA5A" bold="true" box="[1163,1181,1430,1454]" pageId="22" pageNumber="423">A</emphasis>
, lateral view,
<emphasis id="1B8976032609FFB5FAC1FA7D1B63FA5A" bold="true" box="[1334,1351,1430,1454]" pageId="22" pageNumber="423">B</emphasis>
, phallic complex and gonocoxae, dorsal view;
<emphasis id="1B8976032609FFB5FDEAFA511C12FA26" bold="true" box="[541,566,1466,1490]" pageId="22" pageNumber="423">76</emphasis>
, female. Abbreviation: gcx = gonocoxa.
</paragraph>
</caption>
<paragraph id="2942AA112609FFB5FF30F9EB1FCEF93A" blockId="22.[151,1437,1536,1850]" pageId="22" pageNumber="423">
<emphasis id="1B8976032609FFB5FF30F9EB1F3AF9EE" bold="true" box="[199,286,1536,1562]" pageId="22" pageNumber="423">Female</emphasis>
: As in male except as follows: Mesonotum (
<figureCitation id="B1C6B6942609FFB5FCC1F9EB1DA8F9EE" box="[822,908,1536,1562]" captionStart="FIGURES 34–39" captionStartId="14.[151,264,1264,1288]" captionTargetBox="[151,1435,181,1240]" captionTargetId="figure-17@14.[151,1435,181,1240]" captionTargetPageId="14" captionText="FIGURES 34–39. Malagasy Empidideicus (Anomaloptilus), thoraces, dorsal view. 34, E. (A.) regius, n. sp., male; 35, E. (A.) regius, n. sp., female; 36, E. (A.) rhinoclypeatus Evenhuis, male; 37, E. (A.) rhinoclypeatus Evenhuis, female; 38, E. (A.) rinhai, n. sp., male; 39, E. (A.) rinhai, n. sp., female." figureDoi="http://doi.org/10.5281/zenodo.7471760" httpUri="https://zenodo.org/record/7471760/files/figure.png" pageId="22" pageNumber="423">Fig. 39</figureCitation>
) with admedian vittae more pronounced and complete from interhumeral marks to prescutellar marks, in some specimens somewhat coalesced in prescutellar area with orangish color; scutellum all white. Abdomen (
<figureCitation id="B1C6B6942609FFB5FCF9F9A31D7AF996" box="[782,862,1608,1634]" captionStart="FIGURES 59–62" captionStartId="19.[151,264,694,718]" captionTargetBox="[152,1435,181,670]" captionTargetId="figure-17@19.[152,1435,181,670]" captionTargetPageId="19" captionText="FIGURES 59–62. Malagasy Empidideicus (Anomaloptilus), abdomens, dorsal view. 59, E. (A.) rhinoclypeatus Evenhuis, male; 60, E. (A.) rhinoclypeatus Evenhuis, female; 61, E. (A.) rinhai, n. sp., male; 62, E. (A.) rinhai, n. sp., female." figureDoi="http://doi.org/10.5281/zenodo.7471774" httpUri="https://zenodo.org/record/7471774/files/figure.png" pageId="22" pageNumber="423">Fig. 62</figureCitation>
) with brown and yellow to yellowish white pattern; tergite I with anterior brown band complete; tergites II–V with brown band restricted to medial area brown becoming narrower on succeeding segments, tergites II–V with brown dorsolateral spots in yellowish white area. Sternites white. Genitalia not dissected.
</paragraph>
</subSubSection>
<subSubSection id="61E7F99A2609FFB5FF30F9331A06F8CE" pageId="22" pageNumber="423" type="etymology">
<paragraph id="2942AA112609FFB5FF30F9331A06F8CE" blockId="22.[151,1437,1536,1850]" pageId="22" pageNumber="423">
<emphasis id="1B8976032609FFB5FF30F9331F6AF906" bold="true" box="[199,334,1752,1778]" pageId="22" pageNumber="423">Etymology.</emphasis>
Named for Rasolandalao Harin’Hala Hasinjaka (“Rin’ha”) who helped collected the
<typeStatus id="F64614B32609FFB5FACFF9321B4CF907" box="[1336,1384,1753,1779]" pageId="22" pageNumber="423">type</typeStatus>
and was instrumental in collecting thousands and thousands of microbombyliids from
<collectingCountry id="51EAEA812609FFB5FBCDF9171AE5F8E2" box="[1082,1217,1788,1814]" name="Madagascar" pageId="22" pageNumber="423">Madagascar</collectingCountry>
as part of National Science Foundation-funded the “Terrestrial arthropod inventory of
<collectingCountry id="51EAEA812609FFB5FC70F8CB1A30F8CE" box="[903,1044,1824,1850]" name="Madagascar" pageId="22" pageNumber="423">Madagascar</collectingCountry>
”.
</paragraph>
</subSubSection>
</treatment>
</document>
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