316 lines
35 KiB
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316 lines
35 KiB
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<document id="2D1ABAC4307F49B7BABE18306F6BA008" ID="10.11646/zootaxa.4147.3.3" ID-DOI="10.11646/zootaxa.4147.3.3" ID-GBIF-Dataset="bf244739-2235-4e92-a4db-a36440fc1ac8" ID-ISSN="1175-5326" ID-Zenodo-Dep="264843" ID-ZooBank="6238B25B-787F-4F50-BFAE-03CD33D0F699" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1470292933281" checkinUser="plazi" docAuthor="Schileyko, Arkady A. & Stoev, Pavel E." docDate="2016" docId="03FD32670E5BB019FF5CF8C6BF81F8B4" docLanguage="en" docName="zootaxa.4147.3.3.pdf" docOrigin="Zootaxa 4147 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trigonocryptops Verhoeff 1906" docType="treatment" docVersion="13" lastPageNumber="267" masterDocId="FFC44A1F0E48B00DFFCBFF9EBC17FFF9" masterDocTitle="Scolopendromorpha of New Guinea and adjacent islands (Myriapoda, Chilopoda)" masterLastPageNumber="280" masterPageNumber="247" pageNumber="266" updateTime="1698366899779" updateUser="plazi">
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<mods:mods id="8A6014D2D06C5147F5F8B2B47B7F097B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="C2D6FB23BA6FC46D41BD09215BC5D264">
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<mods:title id="5238C4202867149817FADE4337C68BB7">Scolopendromorpha of New Guinea and adjacent islands (Myriapoda, Chilopoda)</mods:title>
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</mods:titleInfo>
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<mods:name id="33856CD6EB01B59A6C0232428F77B627" type="personal">
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<mods:role id="D95AABF15CB106F394FC1058CB0AD4F8">
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<mods:roleTerm id="925185B1CF8969F0DF05E180C0FDC1B2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="64675FBFE22C61A69F2EA6A94FAB7B3C">Schileyko, Arkady A.</mods:namePart>
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</mods:name>
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<mods:name id="3F5B8E525D5880946FF5F915FA646E63" type="personal">
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<mods:role id="AB45505122D4DDEF94F42FA095974FDA">
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<mods:roleTerm id="C5DE746545D94AD2C60E500A50A6C21B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="76306D369C2985D7CB46839ED74C2BCD">Stoev, Pavel E.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="3BAE6F36F67839E664009EB5D7CE5592">text</mods:typeOfResource>
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<mods:relatedItem id="E2E44EBCB5D5AB27ED66EAFDF64FA05B" type="host">
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<mods:titleInfo id="B7463951A33C936D1D2ACC801901E7A0">
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<mods:title id="7ED0116EC57ED58FEEE109D504156D2D">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="A96E281B711C5D7793AC77F26F8C7E56">
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<mods:date id="A36C4C15F4D7B8E7CCF18A891C68DCBF">2016</mods:date>
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<mods:detail id="276DAA7E1500B3389C4FD1F6F1D4570E" type="volume">
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<mods:number id="C192C93EFED684959C24C011D636683B">4147</mods:number>
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</mods:detail>
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<mods:detail id="72DCF78A7C78F36AA5372575CB4D264F" type="issue">
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<mods:number id="88FAED44F92E1D1E9F67B914CFD743B8">3</mods:number>
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</mods:detail>
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<mods:extent id="4C02F624552B79437C77A4232C52689F" unit="page">
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<mods:start id="18C4F1DD9EF110D165557CB61BCFB3D5">247</mods:start>
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<mods:end id="DBC7A6B387FAD11D136D51983502C53A">280</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="A611602CD16958A028E86A3892F54DC7">journal article</mods:classification>
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<mods:identifier id="424197D7FCCB99BB1BF725B38005C39F" type="DOI">10.11646/zootaxa.4147.3.3</mods:identifier>
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<mods:identifier id="45B97255442D3613F274F457511C82B9" type="GBIF-Dataset">bf244739-2235-4e92-a4db-a36440fc1ac8</mods:identifier>
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<mods:identifier id="16C72F1C3C1D832E9C81BF32324A5516" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="C60F4599895ABF6835FB316CE129179F" type="Zenodo-Dep">264843</mods:identifier>
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<mods:identifier id="66ABCEB9C33A1C6DD7D93F9CB4BAF309" type="ZooBank">6238B25B-787F-4F50-BFAE-03CD33D0F699</mods:identifier>
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</mods:mods>
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<treatment id="03FD32670E5BB019FF5CF8C6BF81F8B4" ID-DOI="http://doi.org/10.5281/zenodo.6077656" ID-GBIF-Taxon="123514155" ID-Zenodo-Dep="6077656" LSID="urn:lsid:plazi:treatment:03FD32670E5BB019FF5CF8C6BF81F8B4" httpUri="http://treatment.plazi.org/id/03FD32670E5BB019FF5CF8C6BF81F8B4" lastPageId="20" lastPageNumber="267" pageId="19" pageNumber="266">
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<subSubSection id="C34ED0FA0E5BB01EFF5CF8C6B829F84D" pageId="19" pageNumber="266" type="nomenclature">
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<paragraph id="8BEB83710E5BB01EFF5CF8C6BE8EF88B" blockId="19.[151,665,1880,1906]" box="[151,665,1880,1906]" pageId="19" pageNumber="266">
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<heading id="D0A3341D0E5BB01EFF5CF8C6BE8EF88B" bold="true" box="[151,665,1880,1906]" fontSize="11" level="1" pageId="19" pageNumber="266" reason="1">
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<emphasis id="B9205F630E5BB01EFF5CF8C6BE8EF88B" bold="true" box="[151,665,1880,1906]" pageId="19" pageNumber="266">
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Subgenus
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<taxonomicName id="4C54F8F20E5BB01EFEDCF8C6BE8EF88B" authority="Verhoeff, 1906" authorityName="Verhoeff" authorityYear="1906" box="[279,665,1880,1906]" class="Chilopoda" family="Cryptopidae" genus="Trigonocryptops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="266" phylum="Arthropoda" rank="genus">
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<emphasis id="B9205F630E5BB01EFEDCF8C6BDCBF88B" bold="true" box="[279,476,1880,1906]" italics="true" pageId="19" pageNumber="266">Trigonocryptops</emphasis>
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Verhoeff, 1906
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BEB83710E5BB01EFF5CF800B829F84D" blockId="19.[151,1086,1950,1973]" box="[151,1086,1950,1973]" pageId="19" pageNumber="266">
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Type-species.
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<taxonomicName id="4C54F8F20E5BB01EFEECF801BE7CF84D" authority="Kraepelin, 1903" authorityName="Kraepelin" authorityYear="1903" box="[295,619,1950,1973]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="266" phylum="Arthropoda" rank="species" species="gigas">
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<emphasis id="B9205F630E5BB01EFEECF801BDD6F84C" box="[295,449,1951,1973]" italics="true" pageId="19" pageNumber="266">Cryptops gigas</emphasis>
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<bibRefCitation id="EFC5FE800E5BB01EFE03F800BE7CF84D" author="Kraepelin" box="[456,619,1950,1972]" pageId="19" pageNumber="266" refString="Kraepelin, K. (1903) Revision der Scolopendriden. Mitteilungen aus dem Narturhistorischen Museum, Hamburg, 20, 1 - 276." type="journal article" year="1903">Kraepelin, 1903</bibRefCitation>
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</taxonomicName>
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(by subsequent designation of
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<bibRefCitation id="EFC5FE800E5BB01EFC6DF801B824F84D" author="Attems" box="[934,1075,1950,1972]" pageId="19" pageNumber="266" refString="Attems, C. (1930) Myriopoda. 2. Scolopendromorpha. Das Tierreich 54. Walter de Gruyter, Berlin, pp. 1 - 308." type="journal article" year="1930">Attems, 1930</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C34ED0FA0E5CB019FF5CFF09B8B9FEB9" pageId="20" pageNumber="267" type="description">
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<paragraph id="8BEB83710E5CB019FF5CFF09B8B9FEB9" blockId="20.[151,1437,151,1869]" pageId="20" pageNumber="267">
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<materialsCitation id="3B3C892C0E5CB019FF5CFF09BE1DFF49" ID-GBIF-Occurrence="1290989920" box="[151,522,151,176]" country="Cuba" location="Range. Caribbean Islands" pageId="20" pageNumber="267" specimenCount="1">
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<location id="8E8BD5AA0E5CB019FF5CFF09BDD7FF49" LSID="urn:lsid:plazi:treatment:03FD32670E5BB019FF5CF8C6BF81F8B4:8E8BD5AA0E5CB019FF5CFF09BDD7FF49" box="[151,448,151,176]" country="Cuba" name="Range. Caribbean Islands" pageId="20" pageNumber="267">
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<emphasis id="B9205F630E5CB019FF5CFF09BCFDFF49" bold="true" box="[151,234,151,176]" pageId="20" pageNumber="267">Range.</emphasis>
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Caribbean Islands
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</location>
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:
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<collectingCountry id="F343C3E10E5CB019FE00FF09BE1DFF49" box="[459,522,151,176]" name="Cuba" pageId="20" pageNumber="267">Cuba</collectingCountry>
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</materialsCitation>
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.
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<materialsCitation id="3B3C892C0E5CB019FDDEFF06BD9FFF2C" ID-GBIF-Occurrence="1290989923" collectorName="S America" country="Peru" location="Sao Paulo" pageId="20" pageNumber="267" specimenCount="1" stateProvince="Sao Paulo">
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<collectorName id="26A1E6A70E5CB019FDDEFF06BE85FF49" box="[533,658,151,176]" pageId="20" pageNumber="267">S America</collectorName>
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:
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<collectingCountry id="F343C3E10E5CB019FD56FF06BEC4FF56" box="[669,723,152,175]" name="Peru" pageId="20" pageNumber="267">Peru</collectingCountry>
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, SE
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<collectingCountry id="F343C3E10E5CB019FCCCFF09BF5CFF49" box="[775,843,151,176]" name="Brazil" pageId="20" pageNumber="267">Brazil</collectingCountry>
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(
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<collectingRegion id="49904D930E5CB019FC90FF06BFD9FF49" box="[859,974,151,176]" country="Brazil" name="Sao Paulo" pageId="20" pageNumber="267">São Paulo</collectingRegion>
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State,
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<collectingRegion id="49904D930E5CB019FBD7FF09B8A8FF49" box="[1052,1215,151,176]" country="Brazil" name="Rio de Janeiro" pageId="20" pageNumber="267">
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<collectingRegion id="49904D930E5CB019FBD7FF09B853FF49" box="[1052,1092,151,176]" country="Gambia" name="Upper River" pageId="20" pageNumber="267">Rio</collectingRegion>
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de Janeiro
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</collectingRegion>
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State, Minas Geras State). Europe:
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<collectingCountry id="F343C3E10E5CB019FE8EFF22BD9FFF2C" box="[325,392,188,213]" name="Spain" pageId="20" pageNumber="267">Spain</collectingCountry>
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</materialsCitation>
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. Africa: Algeria, Morocco, Ivory Coast, Guinea, Tanzania, Somalia,
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<materialsCitation id="3B3C892C0E5CB019FB40FF23B98FFF2C" ID-GBIF-Occurrence="1290989922" box="[1163,1432,188,213]" country="Yemen" location="Sokotra Island" pageId="20" pageNumber="267" specimenCount="1">
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<collectingCountry id="F343C3E10E5CB019FB40FF23B8CEFF2D" box="[1163,1241,189,212]" name="Yemen" pageId="20" pageNumber="267">Yemen</collectingCountry>
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(
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<location id="8E8BD5AA0E5CB019FB2CFF22B987FF2C" LSID="urn:lsid:plazi:treatment:03FD32670E5BB019FF5CF8C6BF81F8B4:8E8BD5AA0E5CB019FB2CFF22B987FF2C" box="[1255,1424,188,213]" country="Yemen" name="Sokotra Island" pageId="20" pageNumber="267">Sokotra Island</location>
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)
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</materialsCitation>
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, Gabon, Benin, Cameroon, Congo.
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<materialsCitation id="3B3C892C0E5CB019FDE6FF7EBF5AFF01" ID-GBIF-Occurrence="1290989914" box="[557,845,223,248]" collectorName="SE Asia" country="India" location="India" pageId="20" pageNumber="267" specimenCount="1">
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<collectorName id="26A1E6A70E5CB019FDE6FF7EBE86FF01" box="[557,657,223,248]" pageId="20" pageNumber="267">SE Asia</collectorName>
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:
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<collectingCountry id="F343C3E10E5CB019FD54FF41BECFFF01" box="[671,728,223,248]" name="India" pageId="20" pageNumber="267">India</collectingCountry>
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(Nagpur)
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</materialsCitation>
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, Vietnam.
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<materialsCitation id="3B3C892C0E5CB019FC05FF41BDE7FEE4" ID-GBIF-Occurrence="1290989913" country="Australia" location="Roe Plains" pageId="20" pageNumber="267" specimenCount="1" stateProvince="Queensland">
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Pacific:
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<collectingCountry id="F343C3E10E5CB019FBE4FF41B881FF01" box="[1071,1174,223,248]" name="Australia" pageId="20" pageNumber="267">Australia</collectingCountry>
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(
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<collectingRegion id="49904D930E5CB019FB63FF41B926FF01" box="[1192,1329,223,248]" country="Australia" name="Queensland" pageId="20" pageNumber="267">Queensland</collectingRegion>
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;
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<collectingRegion id="49904D930E5CB019FA8AFF7EBD16FEE4" country="Australia" name="Western Australia" pageId="20" pageNumber="267">Western Australia</collectingRegion>
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,
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<location id="8E8BD5AA0E5CB019FEC6FE9BBD9CFEE4" LSID="urn:lsid:plazi:treatment:03FD32670E5BB019FF5CF8C6BF81F8B4:8E8BD5AA0E5CB019FEC6FE9BBD9CFEE4" box="[269,395,260,285]" country="Australia" name="Roe Plains" pageId="20" pageNumber="267" stateProvince="Queensland">Roe Plains</location>
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), Sumba
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</materialsCitation>
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,
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<materialsCitation id="3B3C892C0E5CB019FE36FE9BBF71FEE4" ID-GBIF-Occurrence="1290989930" box="[509,870,260,285]" country="Guinea-Bissau" location="East" pageId="20" pageNumber="267" specimenCount="1" stateProvince="East">
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East Timor, New
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<collectingCountry id="F343C3E10E5CB019FD0DFE9ABF00FEE4" box="[710,791,260,285]" name="Guinea-Bissau" pageId="20" pageNumber="267">Guinea</collectingCountry>
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Island
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</materialsCitation>
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, Solomon Islands, New Caledonia, New Zealand, Fiji.
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<materialsCitation id="3B3C892C0E5CB019FF07FEB6B8B9FEB9" ID-GBIF-Occurrence="1290989929" box="[204,1198,295,320]" collectorName="Sierra de la Ventana Pcia" country="Argentina" location="Buenos Aires" pageId="20" pageNumber="267" specimenCount="1" stateProvince="Buenos Aires">
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We add to this list
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<collectingCountry id="F343C3E10E5CB019FE57FEB9BE1CFEB9" box="[412,523,295,320]" name="Argentina" pageId="20" pageNumber="267">Argentina</collectingCountry>
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(
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<collectingRegion id="49904D930E5CB019FDD1FEB6BEA1FEB9" box="[538,694,295,320]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="20" pageNumber="267">Buenos Aires</collectingRegion>
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,
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<collectorName id="26A1E6A70E5CB019FD0AFEB9BFCBFEB9" box="[705,988,295,320]" pageId="20" pageNumber="267">Sierra de la Ventana Pcia</collectorName>
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, Cueva del Torro).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34ED0FA0E5CB019FF0CFED2BD64F9D5" pageId="20" pageNumber="267" type="discussion">
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<paragraph id="8BEB83710E5CB019FF0CFED2BD7BFDC4" blockId="20.[151,1437,151,1869]" pageId="20" pageNumber="267">
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<emphasis id="B9205F630E5CB019FF0CFED2BD2CFE9C" bold="true" box="[199,315,332,357]" pageId="20" pageNumber="267">Remarks.</emphasis>
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<bibRefCitation id="EFC5FE800E5CB019FE82FED2BDE4FE9D" author="Attems" box="[329,499,332,356]" pageId="20" pageNumber="267" refString="Attems, C. (1930) Myriopoda. 2. Scolopendromorpha. Das Tierreich 54. Walter de Gruyter, Berlin, pp. 1 - 308." type="journal article" year="1930">Attems (1930)</bibRefCitation>
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wrote the first general key for subgenus
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<taxonomicName id="4C54F8F20E5CB019FC3DFED3B8B8FE9D" box="[1014,1199,333,356]" class="Chilopoda" family="Cryptopidae" genus="Trigonocryptops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="267" phylum="Arthropoda" rank="genus">
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<emphasis id="B9205F630E5CB019FC3DFED3B8B8FE9D" box="[1014,1199,333,356]" italics="true" pageId="20" pageNumber="267">Trigonocryptops</emphasis>
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</taxonomicName>
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, which at the time contained 8 species; the most recent key to the subgenus is the one of
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<bibRefCitation id="EFC5FE800E5CB019FC2AFEEEB8B3FE71" author="Demange" box="[993,1188,367,392]" pageId="20" pageNumber="267" refString="Demange, J. - M. (1968) Myriapodes Chilopodes du Gabon. Biologia Gabonica, 4, 281 - 294." type="journal article" year="1968">Demange (1968)</bibRefCitation>
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, which comprises 10 (sub-)species. According to
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<bibRefCitation id="EFC5FE800E5CB019FE26FE0BBEFBFE54" box="[493,748,404,429]" pageId="20" pageNumber="267" refString="Bonato, L., Chagas Junior, A., Edgecombe, G., Lewis J., Minelli A., Pereira L., Shelley, R., Stoev, P. & Zapparoli, M. (2016) ChiloBase 2.0 - A World Catalogue of Centipedes (Chilopoda). Available from: http: // chilobase. biologia. unipd. it (Accessed 2 Aug. 2016)" type="journal article">
|
|||
|
|
Bonato
|
|||
|
|
<emphasis id="B9205F630E5CB019FD86FE0BBE99FE54" box="[589,654,404,429]" italics="true" pageId="20" pageNumber="267">et al.</emphasis>
|
|||
|
|
(2016)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C54F8F20E5CB019FCCBFE0BBFAEFE55" box="[768,953,405,428]" class="Chilopoda" family="Cryptopidae" genus="Trigonocryptops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="267" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B9205F630E5CB019FCCBFE0BBFAEFE55" box="[768,953,405,428]" italics="true" pageId="20" pageNumber="267">Trigonocryptops</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
includes 24 species, but
|
|||
|
|
<bibRefCitation id="EFC5FE800E5CB019FB36FE0AB98BFE54" author="Lewis" box="[1277,1436,404,429]" pageId="20" pageNumber="267" refString="Lewis, J. G. I. (2005) A reassessment of Cryptops loveridgei Lawrence and C. kivuensis Lawrence, with observations on the subgenera Trigonocryptops Verhoeff and Paratrigonocryptops Demange (Chilopoda, Scolopendromorpha). African Invertebrates, 46, 115 - 124." type="journal article" year="2005">Lewis (2005)</bibRefCitation>
|
|||
|
|
synonymised to it the subgenus
|
|||
|
|
<emphasis id="B9205F630E5CB019FE35FE27BEFEFE29" box="[510,745,441,464]" italics="true" pageId="20" pageNumber="267">Paratrigonocryptops</emphasis>
|
|||
|
|
Demange, 1963 with its single species
|
|||
|
|
<taxonomicName id="4C54F8F20E5CB019FB6DFE26BD58FE0C" authority="Demange, 1963" authorityName="Demange" authorityYear="1963" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="267" phylum="Arthropoda" rank="species" species="quadrisulcatus" subGenus="P.">
|
|||
|
|
<emphasis id="B9205F630E5CB019FB6DFE26B98CFE29" box="[1190,1435,439,464]" italics="true" pageId="20" pageNumber="267">C. (P.) quadrisulcatus</emphasis>
|
|||
|
|
Demange, 1963
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
<bibRefCitation id="EFC5FE800E5CB019FE95FE42BE71FE0D" box="[350,614,476,501]" pageId="20" pageNumber="267" refString="Murienne, J., Edgecombe, G. D. & Giribet, G. (2011) Comparative phylogeography of the centipedes Cryptops pictus and C. niuensis (Chilopoda) in New Caledonia, Fiji and Vanuatu. Organism Diversity and Evolution, 11, 61 - 74. http: // dx. doi. org / 10.1007 / s 13127 - 011 - 0041 - 7" type="journal article">
|
|||
|
|
Murienne
|
|||
|
|
<emphasis id="B9205F630E5CB019FE1EFE43BE05FE0C" box="[469,530,476,501]" italics="true" pageId="20" pageNumber="267">et al.</emphasis>
|
|||
|
|
(2011)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
transferred to this subgenus
|
|||
|
|
<taxonomicName id="4C54F8F20E5CB019FC77FE43B8EDFE0D" authority="Ribaut, 1923" authorityName="Ribaut" authorityYear="1923" box="[956,1274,476,501]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="267" phylum="Arthropoda" rank="species" species="pictus" subGenus="Cryptops">
|
|||
|
|
<emphasis id="B9205F630E5CB019FC77FE43B84CFE0D" box="[956,1115,476,500]" italics="true" pageId="20" pageNumber="267">C. (C.) pictus</emphasis>
|
|||
|
|
Ribaut, 1923
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<bibRefCitation id="EFC5FE800E5CB019FAF3FE42BD56FDE1" author="Azara" pageId="20" pageNumber="267" refString="de Azara, L. N. & Ferreira, R. L. (2013) The first troglobitic Cryptops (Trigonocryptops) (Chilopoda: Scolopendromorpha) from South America and the description of a non-troglobitic species from Brazil. Zootaxa, 3709 (5), 432 - 444. http: // dx. doi. org / 10.11646 / zootaxa. 3709.5.2" type="journal article" year="2013">Ázara & Ferreira (2013)</bibRefCitation>
|
|||
|
|
described
|
|||
|
|
<taxonomicName id="4C54F8F20E5CB019FE72FD9EBE8FFDE1" box="[441,664,512,536]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="267" phylum="Arthropoda" rank="species" species="iporangensis" subGenus="Trigonocryptops">
|
|||
|
|
<emphasis id="B9205F630E5CB019FE72FD9EBE8FFDE1" box="[441,664,512,536]" italics="true" pageId="20" pageNumber="267">C. (T.) iporangensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C54F8F20E5CB019FD06FD9EBF80FDE1" box="[717,919,511,536]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="267" phylum="Arthropoda" rank="species" species="hephaestus" subGenus="Trigonocryptops">
|
|||
|
|
<emphasis id="B9205F630E5CB019FD06FD9EBF80FDE1" box="[717,919,511,536]" italics="true" pageId="20" pageNumber="267">C. (T.) hephaestus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. Thus, at the moment
|
|||
|
|
<taxonomicName id="4C54F8F20E5CB019FB5AFD9FB95DFDE1" box="[1169,1354,513,536]" class="Chilopoda" family="Cryptopidae" genus="Trigonocryptops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="267" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B9205F630E5CB019FB5AFD9FB95DFDE1" box="[1169,1354,513,536]" italics="true" pageId="20" pageNumber="267">Trigonocryptops</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
should include 28 species.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BEB83710E5CB019FF0CFDD9B868FC14" blockId="20.[151,1437,151,1869]" pageId="20" pageNumber="267">
|
|||
|
|
Although the number of species of
|
|||
|
|
<taxonomicName id="4C54F8F20E5CB019FDA2FDD7BF35FD99" box="[617,802,585,608]" class="Chilopoda" family="Cryptopidae" genus="Trigonocryptops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="267" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B9205F630E5CB019FDA2FDD7BF35FD99" box="[617,802,585,608]" italics="true" pageId="20" pageNumber="267">Trigonocryptops</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
has increased more than in three times since Attems’ (1930), the recent papers concerning this taxon contain neither new identification keys, nor new subgeneric diagnosis. Both
|
|||
|
|
<bibRefCitation id="EFC5FE800E5CB019FE9BFD11BE3FFD51" author="Edgecombe" box="[336,552,655,680]" pageId="20" pageNumber="267" refString="Edgecombe, G. (2005) A troglomorphic species of the centipede Cryptops (Trigonocryptops) (Chilopoda: Scolopendromorpha) from Western Australia. Records of the Western Australian Museum, 22, 315 - 323." type="journal article" year="2005">Edgecombe (2005)</bibRefCitation>
|
|||
|
|
and
|
|||
|
|
<bibRefCitation id="EFC5FE800E5CB019FDA8FD11BF6DFD51" author="Azara" box="[611,890,655,680]" pageId="20" pageNumber="267" refString="de Azara, L. N. & Ferreira, R. L. (2013) The first troglobitic Cryptops (Trigonocryptops) (Chilopoda: Scolopendromorpha) from South America and the description of a non-troglobitic species from Brazil. Zootaxa, 3709 (5), 432 - 444. http: // dx. doi. org / 10.11646 / zootaxa. 3709.5.2" type="journal article" year="2013">Ázara & Ferreira (2013)</bibRefCitation>
|
|||
|
|
repeated the combined data of Verhoeff (1906) and
|
|||
|
|
<bibRefCitation id="EFC5FE800E5CB019FF0DFD2ABD7CFD35" author="Attems" box="[198,363,692,716]" pageId="20" pageNumber="267" refString="Attems, C. (1930) Myriopoda. 2. Scolopendromorpha. Das Tierreich 54. Walter de Gruyter, Berlin, pp. 1 - 308." type="journal article" year="1930">Attems (1930)</bibRefCitation>
|
|||
|
|
, but it should be corrected according to recent knowledge. For example at p. 235 Attems stated, that head of
|
|||
|
|
<taxonomicName id="4C54F8F20E5CB019FEF2FD47BDE5FD09" box="[313,498,729,752]" class="Chilopoda" family="Cryptopidae" genus="Trigonocryptops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="267" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B9205F630E5CB019FEF2FD47BDE5FD09" box="[313,498,729,752]" italics="true" pageId="20" pageNumber="267">Trigonocryptops</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
covers the anterior margin of tergite 1, however both studied New Guinean representatives of this subgenus clearly show posterior margin of head capsule covered by tergite 1 (see below; also p.
|
|||
|
|
<quantity id="4CAC2E940E5CB019FF72FCBEBD11FCC1" box="[185,262,799,824]" metricMagnitude="0" metricUnit="m" metricValue="2.5907999999999998" pageId="20" pageNumber="267" unit="in" value="102.0">102 in</quantity>
|
|||
|
|
<bibRefCitation id="EFC5FE800E5CB019FED8FC81BDD8FCC1" author="Wurmli" box="[275,463,799,824]" pageId="20" pageNumber="267" refString="Wurmli, M. (1972) Chilopoda von Sumba und Flores I. Scolopendromorpha. Verhandlungen der Naturforschenden Gesellschaft in Basel, 82, 88 - 104." type="journal article" year="1972">Würmli (1972))</bibRefCitation>
|
|||
|
|
. Also Attems (p. 235) noted “zweiteilig” (=bipartite) katopleure (as in additional specimens of both
|
|||
|
|
<taxonomicName id="4C54F8F20E5CB019FEB9FCDBBF16FCA5" authority="Brolemann, 1902" authorityName="Brolemann" authorityYear="1902" box="[370,769,836,861]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="267" phylum="Arthropoda" rank="species" species="iheringi" subGenus="Trigonocryptops">
|
|||
|
|
<emphasis id="B9205F630E5CB019FEB9FCDBBE08FCA4" box="[370,543,836,861]" italics="true" pageId="20" pageNumber="267">C. (T.) iheringi</emphasis>
|
|||
|
|
(Brölemann, 1902)
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C54F8F20E5CB019FCF6FCDBB89BFCA5" authority="Ribaut, 1923" authorityName="Ribaut" authorityYear="1923" box="[829,1164,836,861]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="267" phylum="Arthropoda" rank="species" species="sarasini" subGenus="Trigonocryptops">
|
|||
|
|
<emphasis id="B9205F630E5CB019FCF6FCDBBFFCFCA5" box="[829,1003,836,860]" italics="true" pageId="20" pageNumber="267">C. (T.) sarasini</emphasis>
|
|||
|
|
Ribaut, 1923
|
|||
|
|
</taxonomicName>
|
|||
|
|
), but both studied New Guinean exemplars have this sclerite definitely not divided vertically (
|
|||
|
|
<figureCitation id="136F9FF40E5CB019FC67FCF9B817FC79" box="[940,1024,871,896]" captionStart="FIGURES 50 – 52" captionStartId="18.[151,264,1861,1883]" captionText="FIGURES 50 – 52. Cryptops (T.) spinipes Pocock, 1891." pageId="20" pageNumber="267">Fig. 50</figureCitation>
|
|||
|
|
). As for the lateral projections of the anterior corners of the endosternite (which should be diagnostic for this subgenus) the similar small projections are visible in
|
|||
|
|
<emphasis id="B9205F630E5CB019FEC3FC2EBDF6FC31" box="[264,481,943,968]" italics="true" pageId="20" pageNumber="267">C. (С.) anomalans.</emphasis>
|
|||
|
|
The latter shows well-developed sternal transverse thickening between the coxae of legs (Fig. 49), reducing the distance between
|
|||
|
|
<taxonomicName id="4C54F8F20E5CB019FD5FFC4BBF5AFC15" box="[660,845,981,1004]" class="Chilopoda" family="Cryptopidae" genus="Trigonocryptops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="267" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B9205F630E5CB019FD5FFC4BBF5AFC15" box="[660,845,981,1004]" italics="true" pageId="20" pageNumber="267">Trigonocryptops</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and the nominal subgenus.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BEB83710E5CB019FF0CFC66BD64FAF4" blockId="20.[151,1437,151,1869]" pageId="20" pageNumber="267">
|
|||
|
|
We also do not regard the following characters as diagnostic: coloration, bi- vs monopartite tarsus of legs, shape of spiracles and paired distal spinose processes of prefemur, femur, tibia and tarsus 1 of ultimate legs. All these features vary widely (sometimes even intraspecifically) in both
|
|||
|
|
<taxonomicName id="4C54F8F20E5CB019FC60FBDEB806FBAE" box="[939,1041,1088,1111]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="267" phylum="Arthropoda" rank="genus" sensu="stricto">
|
|||
|
|
<emphasis id="B9205F630E5CB019FC60FBDEB806FBAE" box="[939,1041,1088,1111]" italics="true" pageId="20" pageNumber="267">Cryptops</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A213E2180E5CB019FBD1FBDEB859FBAE" box="[1050,1102,1088,1111]" pageId="20" pageNumber="267" sensu="stricto">s.str.</taxonomicNameLabel>
|
|||
|
|
and
|
|||
|
|
<emphasis id="B9205F630E5CB019FB4DFBDEB8B7FBAE" box="[1158,1184,1088,1111]" italics="true" pageId="20" pageNumber="267">C.</emphasis>
|
|||
|
|
(
|
|||
|
|
<taxonomicName id="4C54F8F20E5CB019FB7AFBDFB97DFBA1" box="[1201,1386,1089,1112]" class="Chilopoda" family="Cryptopidae" genus="Trigonocryptops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="267" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B9205F630E5CB019FB7AFBDFB97DFBA1" box="[1201,1386,1089,1112]" italics="true" pageId="20" pageNumber="267">Trigonocryptops</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
), so they have not been included in our new diagnosis. Summing up: all characters listed in
|
|||
|
|
<emphasis id="B9205F630E5CB019FBAEFBFAB8CFFB84" bold="true" box="[1125,1240,1124,1149]" pageId="20" pageNumber="267">Diagnosis</emphasis>
|
|||
|
|
below are shared by other species of
|
|||
|
|
<taxonomicName id="4C54F8F20E5CB019FEBBFB16BDC0FB66" box="[368,471,1160,1183]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="267" phylum="Arthropoda" rank="genus" sensu="stricto">
|
|||
|
|
<emphasis id="B9205F630E5CB019FEBBFB16BDC0FB66" box="[368,471,1160,1183]" italics="true" pageId="20" pageNumber="267">Cryptops</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A213E2180E5CB019FE16FB17BE07FB59" box="[477,528,1161,1184]" pageId="20" pageNumber="267" sensu="stricto">s.str.</taxonomicNameLabel>
|
|||
|
|
(see also
|
|||
|
|
<bibRefCitation id="EFC5FE800E5CB019FDB4FB19BF40FB59" author="Lewis" box="[639,855,1159,1184]" pageId="20" pageNumber="267" refString="Lewis, J. G. I. (2005) A reassessment of Cryptops loveridgei Lawrence and C. kivuensis Lawrence, with observations on the subgenera Trigonocryptops Verhoeff and Paratrigonocryptops Demange (Chilopoda, Scolopendromorpha). African Invertebrates, 46, 115 - 124." type="journal article" year="2005">Lewis (2005: 123))</bibRefCitation>
|
|||
|
|
and none of them is shared by all representatives of
|
|||
|
|
<taxonomicName id="4C54F8F20E5CB019FF5CFB33BD47FB3D" box="[151,336,1197,1220]" class="Chilopoda" family="Cryptopidae" genus="Trigonocryptops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="267" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B9205F630E5CB019FF5CFB33BD47FB3D" box="[151,336,1197,1220]" italics="true" pageId="20" pageNumber="267">Trigonocryptops</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. Thus we regard the subgeneric status of the latter as questionable, confirming the recent statement of Lewis (in press) who has considered
|
|||
|
|
<taxonomicName id="4C54F8F20E5CB019FD25FB4EBF42FB1E" box="[750,853,1232,1255]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="267" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B9205F630E5CB019FD25FB4EBF42FB1E" box="[750,853,1232,1255]" italics="true" pageId="20" pageNumber="267">Cryptops</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C54F8F20E5CB019FC5FFB4FB85AFB11" box="[916,1101,1233,1256]" class="Chilopoda" family="Cryptopidae" genus="Trigonocryptops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="267" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B9205F630E5CB019FC5FFB4FB85AFB11" box="[916,1101,1233,1256]" italics="true" pageId="20" pageNumber="267">Trigonocryptops</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
not “as clear cut as current research indicates”.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BEB83710E5CB019FF0CFA89BD64F9D5" blockId="20.[151,1437,151,1869]" pageId="20" pageNumber="267">
|
|||
|
|
<bibRefCitation id="EFC5FE800E5CB019FF0CFA89BDD0FAC9" box="[199,455,1303,1328]" pageId="20" pageNumber="267" refString="Murienne, J., Edgecombe, G. D. & Giribet, G. (2011) Comparative phylogeography of the centipedes Cryptops pictus and C. niuensis (Chilopoda) in New Caledonia, Fiji and Vanuatu. Organism Diversity and Evolution, 11, 61 - 74. http: // dx. doi. org / 10.1007 / s 13127 - 011 - 0041 - 7" type="journal article">
|
|||
|
|
Murienne
|
|||
|
|
<emphasis id="B9205F630E5CB019FEF7FA87BD62FAC9" box="[316,373,1303,1328]" italics="true" pageId="20" pageNumber="267">et al.</emphasis>
|
|||
|
|
(2011)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
at page 70 called the “rhomboid setose field delimited by sutures on the clypeus” and “trigonal sutures on anterior sternites“ to be the “unique apomorphies of
|
|||
|
|
<emphasis id="B9205F630E5CB019FC38FAA3B81AFAAD" box="[1011,1037,1341,1364]" italics="true" pageId="20" pageNumber="267">C.</emphasis>
|
|||
|
|
(
|
|||
|
|
<taxonomicName id="4C54F8F20E5CB019FBEAFAA3B8CDFAAD" box="[1057,1242,1341,1364]" class="Chilopoda" family="Cryptopidae" genus="Trigonocryptops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="267" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B9205F630E5CB019FBEAFAA3B8CDFAAD" box="[1057,1242,1341,1364]" italics="true" pageId="20" pageNumber="267">Trigonocryptops</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
)”. However, the recent concept of this subgenus includes some species which have no sternal trigonal sutures; also clypeal setose plates are shared at least by
|
|||
|
|
<emphasis id="B9205F630E5CB019FE10FA1BBEBEFA64" box="[475,681,1412,1437]" italics="true" pageId="20" pageNumber="267">
|
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C. (С.)
|
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|
|
<taxonomicName id="4C54F8F20E5CB019FDF8FA1ABEBEFA64" box="[563,681,1412,1437]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="267" phylum="Arthropoda" rank="species" species="nepalensis">nepalensis</taxonomicName>
|
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|
|
</emphasis>
|
|||
|
|
and
|
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|
|
<emphasis id="B9205F630E5CB019FD28FA1BBFA2FA64" box="[739,949,1412,1437]" italics="true" pageId="20" pageNumber="267">C. (С.) anomalans</emphasis>
|
|||
|
|
(see above; Figs 45, 48 respectively). The latter species has also well-developed sternal transverse thickening which are one of characteristic features of
|
|||
|
|
<taxonomicName id="4C54F8F20E5CB019FF5CFA53BD47FA1D" box="[151,336,1485,1508]" class="Chilopoda" family="Cryptopidae" genus="Trigonocryptops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="267" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B9205F630E5CB019FF5CFA53BD47FA1D" box="[151,336,1485,1508]" italics="true" pageId="20" pageNumber="267">Trigonocryptops</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. Thus, we believe that the recent concept of
|
|||
|
|
<taxonomicName id="4C54F8F20E5CB019FCA4FA53B83FFA1D" box="[879,1064,1485,1508]" class="Chilopoda" family="Cryptopidae" genus="Trigonocryptops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="267" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B9205F630E5CB019FCA4FA53B83FFA1D" box="[879,1064,1485,1508]" italics="true" pageId="20" pageNumber="267">Trigonocryptops</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
must be reconsidered and only species having anterior sternites with complete trigonal sutures and clypeus with setose plate(s) should be assigned to
|
|||
|
|
<taxonomicName id="4C54F8F20E5CB019FF7FF98BBD7AF9D5" box="[180,365,1557,1580]" class="Chilopoda" family="Cryptopidae" genus="Trigonocryptops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="267" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B9205F630E5CB019FF7FF98BBD7AF9D5" box="[180,365,1557,1580]" italics="true" pageId="20" pageNumber="267">Trigonocryptops</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C34ED0FA0E5CB019FF0CF9A9BF81F8B4" pageId="20" pageNumber="267" type="diagnosis">
|
|||
|
|
<paragraph id="8BEB83710E5CB019FF0CF9A9B829F98D" blockId="20.[151,1437,151,1869]" pageId="20" pageNumber="267">
|
|||
|
|
<emphasis id="B9205F630E5CB019FF0CF9A9BD6CF9A9" bold="true" box="[199,379,1591,1616]" pageId="20" pageNumber="267">New diagnosis.</emphasis>
|
|||
|
|
Cephalic plate usually with paramedian sutures—complete or (more rarely) incomplete, its posterior margin covers (more rarely is covered by) the anterior margin of tergite 1.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BEB83710E5CB019FF0CF9E1B8C7F944" blockId="20.[151,1437,151,1869]" pageId="20" pageNumber="267">
|
|||
|
|
Clypeus with 1 or 2 (large anterior and very small posterior) setose plates which are limited by sutures and bear 2–10 setae (“setose clypeal plates” sensu
|
|||
|
|
<bibRefCitation id="EFC5FE800E5CB019FDA3F93ABF1CF944" author="Lewis" box="[616,779,1700,1725]" pageId="20" pageNumber="267" refString="Lewis, J. G. I. (2005) A reassessment of Cryptops loveridgei Lawrence and C. kivuensis Lawrence, with observations on the subgenera Trigonocryptops Verhoeff and Paratrigonocryptops Demange (Chilopoda, Scolopendromorpha). African Invertebrates, 46, 115 - 124." type="journal article" year="2005">Lewis (2005))</bibRefCitation>
|
|||
|
|
. Labrum with 1 or (more rarely) 3 teeth.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BEB83710E5CB019FF0CF959B813F8D1" blockId="20.[151,1437,151,1869]" pageId="20" pageNumber="267">Tergite 1 mainly with anterior transverse suture, often with (complete or incomplete) paramedian sutures. Sternites with well-developed transverse thickening between the coxae. Endosternites clearly delimited anteriorly (often by the very characteristic trigonal sutures). Katopleure mainly bipartite.</paragraph>
|
|||
|
|
<paragraph id="8BEB83710E5CB019FF0CF8ABBF81F8B4" blockId="20.[151,1437,151,1869]" box="[199,918,1844,1869]" pageId="20" pageNumber="267">Femur of ultimate legs mainly with 1 well-developed saw tooth.</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|