Scolopendromorpha of New Guinea and adjacent islands (Myriapoda, Chilopoda)
Author
Schileyko, Arkady A.
Author
Stoev, Pavel E.
text
Zootaxa
2016
4147
3
247
280
journal article
10.11646/zootaxa.4147.3.3
bf244739-2235-4e92-a4db-a36440fc1ac8
1175-5326
264843
6238B25B-787F-4F50-BFAE-03CD33D0F699
Subgenus
Trigonocryptops
Verhoeff, 1906
Type-species.
Cryptops gigas
Kraepelin, 1903
(by subsequent designation of
Attems, 1930
).
Range.
Caribbean Islands
:
Cuba
.
S America
:
Peru
, SE
Brazil
(
São Paulo
State,
Rio
de Janeiro
State, Minas Geras State). Europe:
Spain
. Africa: Algeria, Morocco, Ivory Coast, Guinea, Tanzania, Somalia,
Yemen
(
Sokotra Island
)
, Gabon, Benin, Cameroon, Congo.
SE Asia
:
India
(Nagpur)
, Vietnam.
Pacific:
Australia
(
Queensland
;
Western Australia
,
Roe Plains
), Sumba
,
East Timor, New
Guinea
Island
, Solomon Islands, New Caledonia, New Zealand, Fiji.
We add to this list
Argentina
(
Buenos Aires
,
Sierra de la Ventana Pcia
, Cueva del Torro).
Remarks.
Attems (1930)
wrote the first general key for subgenus
Trigonocryptops
, which at the time contained 8 species; the most recent key to the subgenus is the one of
Demange (1968)
, which comprises 10 (sub-)species. According to
Bonato
et al.
(2016)
,
Trigonocryptops
includes 24 species, but
Lewis (2005)
synonymised to it the subgenus
Paratrigonocryptops
Demange, 1963 with its single species
C. (P.) quadrisulcatus
Demange, 1963
.
Murienne
et al.
(2011)
transferred to this subgenus
C. (C.) pictus
Ribaut, 1923
and
Ázara & Ferreira (2013)
described
C. (T.) iporangensis
and
C. (T.) hephaestus
. Thus, at the moment
Trigonocryptops
should include 28 species.
Although the number of species of
Trigonocryptops
has increased more than in three times since Attems’ (1930), the recent papers concerning this taxon contain neither new identification keys, nor new subgeneric diagnosis. Both
Edgecombe (2005)
and
Ázara & Ferreira (2013)
repeated the combined data of Verhoeff (1906) and
Attems (1930)
, but it should be corrected according to recent knowledge. For example at p. 235 Attems stated, that head of
Trigonocryptops
covers the anterior margin of tergite 1, however both studied New Guinean representatives of this subgenus clearly show posterior margin of head capsule covered by tergite 1 (see below; also p.
102 in
Würmli (1972))
. Also Attems (p. 235) noted “zweiteilig” (=bipartite) katopleure (as in additional specimens of both
C. (T.) iheringi
(Brölemann, 1902)
and
C. (T.) sarasini
Ribaut, 1923
), but both studied New Guinean exemplars have this sclerite definitely not divided vertically (
Fig. 50
). As for the lateral projections of the anterior corners of the endosternite (which should be diagnostic for this subgenus) the similar small projections are visible in
C. (С.) anomalans.
The latter shows well-developed sternal transverse thickening between the coxae of legs (Fig. 49), reducing the distance between
Trigonocryptops
and the nominal subgenus.
We also do not regard the following characters as diagnostic: coloration, bi- vs monopartite tarsus of legs, shape of spiracles and paired distal spinose processes of prefemur, femur, tibia and tarsus 1 of ultimate legs. All these features vary widely (sometimes even intraspecifically) in both
Cryptops
s.str.
and
C.
(
Trigonocryptops
), so they have not been included in our new diagnosis. Summing up: all characters listed in
Diagnosis
below are shared by other species of
Cryptops
s.str.
(see also
Lewis (2005: 123))
and none of them is shared by all representatives of
Trigonocryptops
. Thus we regard the subgeneric status of the latter as questionable, confirming the recent statement of Lewis (in press) who has considered
Cryptops
and
Trigonocryptops
not “as clear cut as current research indicates”.
Murienne
et al.
(2011)
at page 70 called the “rhomboid setose field delimited by sutures on the clypeus” and “trigonal sutures on anterior sternites“ to be the “unique apomorphies of
C.
(
Trigonocryptops
)”. However, the recent concept of this subgenus includes some species which have no sternal trigonal sutures; also clypeal setose plates are shared at least by
C. (С.)
nepalensis
and
C. (С.) anomalans
(see above; Figs 45, 48 respectively). The latter species has also well-developed sternal transverse thickening which are one of characteristic features of
Trigonocryptops
. Thus, we believe that the recent concept of
Trigonocryptops
must be reconsidered and only species having anterior sternites with complete trigonal sutures and clypeus with setose plate(s) should be assigned to
Trigonocryptops
.
New diagnosis.
Cephalic plate usually with paramedian sutures—complete or (more rarely) incomplete, its posterior margin covers (more rarely is covered by) the anterior margin of tergite 1.
Clypeus with 1 or 2 (large anterior and very small posterior) setose plates which are limited by sutures and bear 2–10 setae (“setose clypeal plates” sensu
Lewis (2005))
. Labrum with 1 or (more rarely) 3 teeth.
Tergite 1 mainly with anterior transverse suture, often with (complete or incomplete) paramedian sutures. Sternites with well-developed transverse thickening between the coxae. Endosternites clearly delimited anteriorly (often by the very characteristic trigonal sutures). Katopleure mainly bipartite.
Femur of ultimate legs mainly with 1 well-developed saw tooth.