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<document id="DEC2559739CD9C01B63092BDE74A26F0" ID-DOI="10.1649/0010-065X-72.1.97" ID-ISSN="1938-4394" ID-Zenodo-Dep="4836701" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622211867469" checkinUser="carolina" docAuthor="Miller, Kelly B. &amp; Wolfe, G. William" docDate="2018" docId="03A1932F092CFF9A81D1CD36FE37FCF4" docLanguage="en" docName="TheColeopBull.72.1.113-117.pdf" docOrigin="The Coleopterists Bulletin 72 (1)" docSource="http://dx.doi.org/10.1649/0010-065x-72.1.97" docStyle="DocumentStyle:6D94AA43905A5ECAD3A4A280834A9134.5:TheColeopBull.2017-.journal_article.0cover" docStyleId="6D94AA43905A5ECAD3A4A280834A9134" docStyleName="TheColeopBull.2017-.journal_article.0cover" docStyleVersion="5" docTitle="Desmopachria lloydi Miller and Wolfe 2018, new species" docType="treatment" docVersion="4" lastPageNumber="106" masterDocId="FF98EB570924FF938321CE69FFB6FFCE" masterDocTitle="Nine New Species in the Desmopachria nitida Species-Group of the Neotropical Genus Desmopachria Babington, 1841 (Coleoptera: Adephaga: Dytiscidae: Hydroporinae: Hyphydrini)" masterLastPageNumber="112" masterPageNumber="97" pageNumber="105" updateTime="1698949393783" updateUser="plazi">
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<mods:title id="7FFD2A0CED12F14DAD907EF753260D1C">Nine New Species in the Desmopachria nitida Species-Group of the Neotropical Genus Desmopachria Babington, 1841 (Coleoptera: Adephaga: Dytiscidae: Hydroporinae: Hyphydrini)</mods:title>
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<mods:namePart id="DD6E88B1FC682C11DE0E848582E91EF4">Miller, Kelly B.</mods:namePart>
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<mods:namePart id="4C270C4A662ECD985BFABCE6C7F4ACED">Wolfe, G. William</mods:namePart>
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<mods:title id="695372810AD99DA40FF9CCEC9B990B9D">The Coleopterists Bulletin</mods:title>
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<mods:date id="75FA3C7B58EAC908F23630146C4E7969">2018</mods:date>
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<mods:number id="C5DD73CDD8F1C2ECFC1A39A0A8FF2DA2">2018-03-23</mods:number>
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<mods:number id="8C08FEBA569E846EC62F0A140285B657">72</mods:number>
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<mods:identifier id="B5DA4719938AB9012CAD529CE344BAFB" type="DOI">10.1649/0010-065X-72.1.97</mods:identifier>
<mods:identifier id="C122B14BB978C4E586440E784FC0A13A" type="ISSN">1938-4394</mods:identifier>
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<treatment id="03A1932F092CFF9A81D1CD36FE37FCF4" ID-DOI="http://doi.org/10.5281/zenodo.4836734" ID-Zenodo-Dep="4836734" LSID="urn:lsid:zoobank.org:act:38EE2BE0-8A32-4ADB-9F9F-F03B257A5449" httpUri="http://treatment.plazi.org/id/03A1932F092CFF9A81D1CD36FE37FCF4" lastPageId="9" lastPageNumber="106" pageId="8" pageNumber="105">
<subSubSection id="C31271B2092CFF9B81D1CD36FBBCFC10" pageId="8" pageNumber="105" type="nomenclature">
<paragraph id="8BB72239092CFF9B81D1CD36FC46FC41" blockId="8.[721,1178,863,991]" pageId="8" pageNumber="105">
<heading id="D0FF9555092CFF9B81D1CD36FC46FC41" centered="true" fontSize="9" level="2" pageId="8" pageNumber="105" reason="2">
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<taxonomicName id="4C0859BA092CFF9B81D1CD36FBC1FCBA" ID-CoL="8LZVF" authority="Miller and Wolfe" authorityName="Miller and Wolfe" authorityYear="2018" box="[752,1143,863,884]" class="Insecta" family="Dytiscidae" genus="Desmopachria" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="105" phylum="Arthropoda" rank="species" species="lloydi" status="sp. nov.">
<emphasis id="B97CFE2B092CFF9B81D1CD36FC08FCBA" bold="true" box="[752,958,863,884]" italics="true" pageId="8" pageNumber="105">Desmopachria lloydi</emphasis>
Miller and Wolfe
</taxonomicName>
,
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</emphasis>
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<paragraph id="8BB72239092CFF9B81DECDFDFBBCFC10" blockId="8.[721,1178,863,991]" pageId="8" pageNumber="105">
Zoobank.org/
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urn:lsid:zoobank.org:act:
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(
<figureCitation id="13333EBC092CFF9B8048CDA0FC68FC10" box="[873,990,969,991]" captionStart="Figs" captionStartId="2.[197,236,1466,1485]" captionTargetBox="[325,1043,194,1444]" captionTargetId="figure-0@2.[322,1044,194,1451]" captionTargetPageId="2" captionText="Figs. 1–32. Male genitalia of Desmopachria species in the D. nitida species-group. D. curseenae: Median lobe – 1) Right lateral aspect; 2) Ventral aspect; Right lateral lobe – 3) Right lateral aspect; 4) Ventral aspect. D. delongi: Median lobe – 5) Right lateral aspect; 6) Ventral aspect; Right lateral lobe – 7) Right lateral aspect; 8) Ventral aspect. D. gingerae: Median lobe – 9) Right lateral aspect; 10) Ventral aspect; 11) Right lateral lobe, right lateral aspect. D. gyrationi: Median lobe – 12) Right lateral aspect; 13) Ventral aspect; 14) Right lateral lobe, right lateral aspect. D. hardyae: Median lobe – 15) Right lateral aspect; 16) Ventral aspect; 17) Right lateral lobe, right lateral aspect. D. kemptonae: Median lobe – 18) Right lateral aspect; 19) Ventral aspect; Right lateral lobe – 20) Right lateral aspect; 21) Ventral aspect. D. lloydi: 22) Median lobe, ventral aspect; 23) Apices of median lobe, anterior aspect; Right lateral lobe – 24) Right lateral aspect; 25) Ventral aspect. D. singhae: Median lobe – 26) Right lateral aspect; 27) Ventral aspect; 28) Right lateral lobe, right lateral aspect. D. vohrae: Median lobe – 29) Right lateral aspect; 30) Ventral aspect; Right lateral lobe – 31) Right lateral aspect; 32) Ventral aspect." figureDoi="http://doi.org/10.5281/zenodo.4836705" httpUri="https://zenodo.org/record/4836705/files/figure.png" pageId="8" pageNumber="105">Figs. 22–25</figureCitation>
,
<figureCitation id="13333EBC092CFF9B80CBCDA0FBB5FC10" box="[1002,1027,969,990]" captionStart="Figs" captionStartId="13.[175,214,1279,1298]" captionTargetBox="[151,1173,194,1263]" captionTargetId="figure-175@13.[151,1173,194,1263]" captionTargetPageId="13" captionText="Figs. 71–72. Distribution of Desmopachria species in the D. nitida species-group. 71) D. curseenae, D. singhae, and D. vohrae; 72) D. lloydi." figureDoi="http://doi.org/10.5281/zenodo.4836725" httpUri="https://zenodo.org/record/4836725/files/figure.png" pageId="8" pageNumber="105">72</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31271B2092CFF9B81FACD93FCE7FBE4" pageId="8" pageNumber="105" type="materials_examined">
<paragraph id="8BB72239092CFF9B81FACD93FCE7FBE4" blockId="8.[707,1193,1018,1730]" pageId="8" pageNumber="105">
<materialsCitation id="3B602864092CFF9B81FACD93FCFBFBE4" collectorName="El Porvenir Station NE San Borja" country="Bolivia" location="El Porvenir Station" pageId="8" pageNumber="105" specimenCount="1" stateProvince="Beni" typeStatus="holotype">
<emphasis id="B97CFE2B092CFF9B81FACD93FCDFFBC1" bold="true" box="[731,873,1018,1039]" pageId="8" pageNumber="105">
<typeStatus id="54B39C9B092CFF9B81FACD93FCBBFBC1" box="[731,781,1018,1039]" pageId="8" pageNumber="105">Type</typeStatus>
Locality.
</emphasis>
<collectingCountry id="F31F62A9092CFF9B804FCD93FC00FBC1" box="[878,950,1018,1039]" name="Bolivia" pageId="8" pageNumber="105">Bolivia</collectingCountry>
,
<collectingRegion id="49CCECDB092CFF9B809FCD93FC5BFBC1" box="[958,1005,1018,1039]" country="Bolivia" name="Beni" pageId="8" pageNumber="105">Beni</collectingRegion>
,
<collectorName id="26FD47EF092CFF9B80D7CD93FCFBFBE4" pageId="8" pageNumber="105">
<location id="8ED774E2092CFF9B80D7CD93FB1FFBC1" LSID="urn:lsid:plazi:treatment:03A1932F092CFF9A81D1CD36FE37FCF4:8ED774E2092CFF9B80D7CD93FB1FFBC1" box="[1014,1193,1018,1039]" country="Bolivia" name="El Porvenir Station" pageId="8" pageNumber="105" stateProvince="Beni">El Porvenir Station</location>
NE San Borja
</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31271B2092CFF9B81FACA46FCB4FA50" pageId="8" pageNumber="105" type="diagnosis">
<paragraph id="8BB72239092CFF9B81FACA46FCB4FA50" blockId="8.[707,1193,1018,1730]" pageId="8" pageNumber="105">
<emphasis id="B97CFE2B092CFF9B81FACA46FCF4FB8A" bold="true" box="[731,834,1071,1092]" pageId="8" pageNumber="105">Diagnosis.</emphasis>
This species has exceptional male genitalia (
<figureCitation id="13333EBC092CFF9B8007CA23FC28FB91" box="[806,926,1097,1119]" captionStart="Figs" captionStartId="2.[197,236,1466,1485]" captionTargetBox="[325,1043,194,1444]" captionTargetId="figure-0@2.[322,1044,194,1451]" captionTargetPageId="2" captionText="Figs. 1–32. Male genitalia of Desmopachria species in the D. nitida species-group. D. curseenae: Median lobe – 1) Right lateral aspect; 2) Ventral aspect; Right lateral lobe – 3) Right lateral aspect; 4) Ventral aspect. D. delongi: Median lobe – 5) Right lateral aspect; 6) Ventral aspect; Right lateral lobe – 7) Right lateral aspect; 8) Ventral aspect. D. gingerae: Median lobe – 9) Right lateral aspect; 10) Ventral aspect; 11) Right lateral lobe, right lateral aspect. D. gyrationi: Median lobe – 12) Right lateral aspect; 13) Ventral aspect; 14) Right lateral lobe, right lateral aspect. D. hardyae: Median lobe – 15) Right lateral aspect; 16) Ventral aspect; 17) Right lateral lobe, right lateral aspect. D. kemptonae: Median lobe – 18) Right lateral aspect; 19) Ventral aspect; Right lateral lobe – 20) Right lateral aspect; 21) Ventral aspect. D. lloydi: 22) Median lobe, ventral aspect; 23) Apices of median lobe, anterior aspect; Right lateral lobe – 24) Right lateral aspect; 25) Ventral aspect. D. singhae: Median lobe – 26) Right lateral aspect; 27) Ventral aspect; 28) Right lateral lobe, right lateral aspect. D. vohrae: Median lobe – 29) Right lateral aspect; 30) Ventral aspect; Right lateral lobe – 31) Right lateral aspect; 32) Ventral aspect." figureDoi="http://doi.org/10.5281/zenodo.4836705" httpUri="https://zenodo.org/record/4836705/files/figure.png" pageId="8" pageNumber="105">Figs. 22–25</figureCitation>
). The male median lobe is broad laterally and deeply bifid with the lateral rami distinctly asymmetrical with each ramus irregularly curved and twisted (
<figureCitation id="13333EBC092CFF9B80D3CAF0FBD9FB60" box="[1010,1135,1177,1198]" captionStart="Figs" captionStartId="2.[197,236,1466,1485]" captionTargetBox="[325,1043,194,1444]" captionTargetId="figure-0@2.[322,1044,194,1451]" captionTargetPageId="2" captionText="Figs. 1–32. Male genitalia of Desmopachria species in the D. nitida species-group. D. curseenae: Median lobe – 1) Right lateral aspect; 2) Ventral aspect; Right lateral lobe – 3) Right lateral aspect; 4) Ventral aspect. D. delongi: Median lobe – 5) Right lateral aspect; 6) Ventral aspect; Right lateral lobe – 7) Right lateral aspect; 8) Ventral aspect. D. gingerae: Median lobe – 9) Right lateral aspect; 10) Ventral aspect; 11) Right lateral lobe, right lateral aspect. D. gyrationi: Median lobe – 12) Right lateral aspect; 13) Ventral aspect; 14) Right lateral lobe, right lateral aspect. D. hardyae: Median lobe – 15) Right lateral aspect; 16) Ventral aspect; 17) Right lateral lobe, right lateral aspect. D. kemptonae: Median lobe – 18) Right lateral aspect; 19) Ventral aspect; Right lateral lobe – 20) Right lateral aspect; 21) Ventral aspect. D. lloydi: 22) Median lobe, ventral aspect; 23) Apices of median lobe, anterior aspect; Right lateral lobe – 24) Right lateral aspect; 25) Ventral aspect. D. singhae: Median lobe – 26) Right lateral aspect; 27) Ventral aspect; 28) Right lateral lobe, right lateral aspect. D. vohrae: Median lobe – 29) Right lateral aspect; 30) Ventral aspect; Right lateral lobe – 31) Right lateral aspect; 32) Ventral aspect." figureDoi="http://doi.org/10.5281/zenodo.4836705" httpUri="https://zenodo.org/record/4836705/files/figure.png" pageId="8" pageNumber="105">Figs. 22, 23</figureCitation>
). The lateral lobes are bilaterally symmetrical with each lobe bifid (
<figureCitation id="13333EBC092CFF9B801ACAA6FC09FB2A" box="[827,959,1231,1252]" captionStart="Figs" captionStartId="2.[197,236,1466,1485]" captionTargetBox="[325,1043,194,1444]" captionTargetId="figure-0@2.[322,1044,194,1451]" captionTargetPageId="2" captionText="Figs. 1–32. Male genitalia of Desmopachria species in the D. nitida species-group. D. curseenae: Median lobe – 1) Right lateral aspect; 2) Ventral aspect; Right lateral lobe – 3) Right lateral aspect; 4) Ventral aspect. D. delongi: Median lobe – 5) Right lateral aspect; 6) Ventral aspect; Right lateral lobe – 7) Right lateral aspect; 8) Ventral aspect. D. gingerae: Median lobe – 9) Right lateral aspect; 10) Ventral aspect; 11) Right lateral lobe, right lateral aspect. D. gyrationi: Median lobe – 12) Right lateral aspect; 13) Ventral aspect; 14) Right lateral lobe, right lateral aspect. D. hardyae: Median lobe – 15) Right lateral aspect; 16) Ventral aspect; 17) Right lateral lobe, right lateral aspect. D. kemptonae: Median lobe – 18) Right lateral aspect; 19) Ventral aspect; Right lateral lobe – 20) Right lateral aspect; 21) Ventral aspect. D. lloydi: 22) Median lobe, ventral aspect; 23) Apices of median lobe, anterior aspect; Right lateral lobe – 24) Right lateral aspect; 25) Ventral aspect. D. singhae: Median lobe – 26) Right lateral aspect; 27) Ventral aspect; 28) Right lateral lobe, right lateral aspect. D. vohrae: Median lobe – 29) Right lateral aspect; 30) Ventral aspect; Right lateral lobe – 31) Right lateral aspect; 32) Ventral aspect." figureDoi="http://doi.org/10.5281/zenodo.4836705" httpUri="https://zenodo.org/record/4836705/files/figure.png" pageId="8" pageNumber="105">Figs. 24, 25</figureCitation>
). The dorsal ramus is abruptly curved dorsad, and the ventral ramus is evenly curved ventrad (
<figureCitation id="13333EBC092CFF9B80E1CB6DFBA7FAD7" box="[960,1041,1284,1305]" captionStart="Figs" captionStartId="2.[197,236,1466,1485]" captionTargetBox="[325,1043,194,1444]" captionTargetId="figure-0@2.[322,1044,194,1451]" captionTargetPageId="2" captionText="Figs. 1–32. Male genitalia of Desmopachria species in the D. nitida species-group. D. curseenae: Median lobe – 1) Right lateral aspect; 2) Ventral aspect; Right lateral lobe – 3) Right lateral aspect; 4) Ventral aspect. D. delongi: Median lobe – 5) Right lateral aspect; 6) Ventral aspect; Right lateral lobe – 7) Right lateral aspect; 8) Ventral aspect. D. gingerae: Median lobe – 9) Right lateral aspect; 10) Ventral aspect; 11) Right lateral lobe, right lateral aspect. D. gyrationi: Median lobe – 12) Right lateral aspect; 13) Ventral aspect; 14) Right lateral lobe, right lateral aspect. D. hardyae: Median lobe – 15) Right lateral aspect; 16) Ventral aspect; 17) Right lateral lobe, right lateral aspect. D. kemptonae: Median lobe – 18) Right lateral aspect; 19) Ventral aspect; Right lateral lobe – 20) Right lateral aspect; 21) Ventral aspect. D. lloydi: 22) Median lobe, ventral aspect; 23) Apices of median lobe, anterior aspect; Right lateral lobe – 24) Right lateral aspect; 25) Ventral aspect. D. singhae: Median lobe – 26) Right lateral aspect; 27) Ventral aspect; 28) Right lateral lobe, right lateral aspect. D. vohrae: Median lobe – 29) Right lateral aspect; 30) Ventral aspect; Right lateral lobe – 31) Right lateral aspect; 32) Ventral aspect." figureDoi="http://doi.org/10.5281/zenodo.4836705" httpUri="https://zenodo.org/record/4836705/files/figure.png" pageId="8" pageNumber="105">Fig. 24</figureCitation>
). This species is pale with the head and pronotum evenly yellow and the elytron evenly brown. The broad, asymmetrical, highly modified male median lobe is completely unique within the
<taxonomicName id="4C0859BA092CFF9B80D7CB07FBE7FA4D" authorityName="Babington" authorityYear="1841" box="[1014,1105,1389,1411]" class="Insecta" family="Dytiscidae" genus="Desmopachria" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="105" phylum="Arthropoda" rank="species" species="nitida">
<emphasis id="B97CFE2B092CFF9B80D7CB07FBE7FA4D" box="[1014,1105,1389,1411]" italics="true" pageId="8" pageNumber="105">D. nitida</emphasis>
</taxonomicName>
speciesgroup.
</paragraph>
</subSubSection>
<subSubSection id="C31271B2092CFF9A81FACBCAFED9FE95" lastPageId="9" lastPageNumber="106" pageId="8" pageNumber="105" type="description">
<paragraph id="8BB72239092CFF9A81FACBCAFDD7FEC2" blockId="8.[707,1193,1018,1730]" lastBlockId="9.[151,639,194,826]" lastPageId="9" lastPageNumber="106" pageId="8" pageNumber="105">
<emphasis id="B97CFE2B092CFF9B81FACBCAFC47FA76" bold="true" box="[731,1009,1443,1464]" pageId="8" pageNumber="105">Description. Measurements:</emphasis>
TL =
<quantity id="4CF08FDC092CFF9B8713CBCAFB13FA76" box="[1074,1189,1443,1465]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="8" pageNumber="105" unit="mm" value="1.75" valueMax="1.8" valueMin="1.7">1.7–1.8 mm</quantity>
, GW =
<quantity id="4CF08FDC092CFF9B8030CBD7FC31FA1D" box="[785,903,1469,1491]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="8" pageNumber="105" unit="mm" value="1.15" valueMax="1.2" valueMin="1.1">1.1–1.2 mm</quantity>
, HW =
<quantity id="4CF08FDC092CFF9B80C2CBD7FBECFA1D" box="[995,1114,1469,1491]" metricMagnitude="-4" metricUnit="m" metricValue="6.499999999999999" metricValueMax="7.0" metricValueMin="6.0" pageId="8" pageNumber="105" unit="mm" value="0.6499999999999999" valueMax="0.7" valueMin="0.6">0.6–0.7 mm</quantity>
, EW =
<quantity id="4CF08FDC092CFF9B81E2CBB1FCBDFA23" box="[707,779,1496,1517]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="105" unit="mm" value="0.3">0.3 mm</quantity>
, TL/GW = 1.5, HW/EW = 2.1.
<emphasis id="B97CFE2B092CFF9B8717CBB1FB1FFA23" bold="true" box="[1078,1193,1496,1517]" pageId="8" pageNumber="105">Coloration:</emphasis>
Head and pronotum evenly yellow. Elytron evenly brown. Head appendages, legs, and ventral surfaces yellow, darker on metacoxae and abdomen.
<emphasis id="B97CFE2B092CFF9B8757C841FCA9F996" bold="true" pageId="8" pageNumber="105">Male genitalia:</emphasis>
Male median lobe highly irregular and asymmetrical. In ventral aspect very broad, asymmetrical, deeply bifid with each ramus elongate, irregularly modified (
<figureCitation id="13333EBC092CFF9B80A7C8FBFBB4F969" box="[902,1026,1682,1703]" captionStart="Figs" captionStartId="2.[197,236,1466,1485]" captionTargetBox="[325,1043,194,1444]" captionTargetId="figure-0@2.[322,1044,194,1451]" captionTargetPageId="2" captionText="Figs. 1–32. Male genitalia of Desmopachria species in the D. nitida species-group. D. curseenae: Median lobe – 1) Right lateral aspect; 2) Ventral aspect; Right lateral lobe – 3) Right lateral aspect; 4) Ventral aspect. D. delongi: Median lobe – 5) Right lateral aspect; 6) Ventral aspect; Right lateral lobe – 7) Right lateral aspect; 8) Ventral aspect. D. gingerae: Median lobe – 9) Right lateral aspect; 10) Ventral aspect; 11) Right lateral lobe, right lateral aspect. D. gyrationi: Median lobe – 12) Right lateral aspect; 13) Ventral aspect; 14) Right lateral lobe, right lateral aspect. D. hardyae: Median lobe – 15) Right lateral aspect; 16) Ventral aspect; 17) Right lateral lobe, right lateral aspect. D. kemptonae: Median lobe – 18) Right lateral aspect; 19) Ventral aspect; Right lateral lobe – 20) Right lateral aspect; 21) Ventral aspect. D. lloydi: 22) Median lobe, ventral aspect; 23) Apices of median lobe, anterior aspect; Right lateral lobe – 24) Right lateral aspect; 25) Ventral aspect. D. singhae: Median lobe – 26) Right lateral aspect; 27) Ventral aspect; 28) Right lateral lobe, right lateral aspect. D. vohrae: Median lobe – 29) Right lateral aspect; 30) Ventral aspect; Right lateral lobe – 31) Right lateral aspect; 32) Ventral aspect." figureDoi="http://doi.org/10.5281/zenodo.4836705" httpUri="https://zenodo.org/record/4836705/files/figure.png" pageId="8" pageNumber="105">Figs. 22, 23</figureCitation>
). Lateral lobe in lateral aspect broad medially, apically bifid, ventral ramus short, curved ventrad, dorsal ramus short, abruptly curved dorsad (
<figureCitation id="13333EBC092DFF9A82A2CEB5FE7AFF3F" box="[387,460,220,241]" captionStart="Figs" captionStartId="2.[197,236,1466,1485]" captionTargetBox="[325,1043,194,1444]" captionTargetId="figure-0@2.[322,1044,194,1451]" captionTargetPageId="2" captionText="Figs. 1–32. Male genitalia of Desmopachria species in the D. nitida species-group. D. curseenae: Median lobe – 1) Right lateral aspect; 2) Ventral aspect; Right lateral lobe – 3) Right lateral aspect; 4) Ventral aspect. D. delongi: Median lobe – 5) Right lateral aspect; 6) Ventral aspect; Right lateral lobe – 7) Right lateral aspect; 8) Ventral aspect. D. gingerae: Median lobe – 9) Right lateral aspect; 10) Ventral aspect; 11) Right lateral lobe, right lateral aspect. D. gyrationi: Median lobe – 12) Right lateral aspect; 13) Ventral aspect; 14) Right lateral lobe, right lateral aspect. D. hardyae: Median lobe – 15) Right lateral aspect; 16) Ventral aspect; 17) Right lateral lobe, right lateral aspect. D. kemptonae: Median lobe – 18) Right lateral aspect; 19) Ventral aspect; Right lateral lobe – 20) Right lateral aspect; 21) Ventral aspect. D. lloydi: 22) Median lobe, ventral aspect; 23) Apices of median lobe, anterior aspect; Right lateral lobe – 24) Right lateral aspect; 25) Ventral aspect. D. singhae: Median lobe – 26) Right lateral aspect; 27) Ventral aspect; 28) Right lateral lobe, right lateral aspect. D. vohrae: Median lobe – 29) Right lateral aspect; 30) Ventral aspect; Right lateral lobe – 31) Right lateral aspect; 32) Ventral aspect." figureDoi="http://doi.org/10.5281/zenodo.4836705" httpUri="https://zenodo.org/record/4836705/files/figure.png" pageId="9" pageNumber="106">Fig. 24</figureCitation>
); in ventral aspect broad, apical rami short and divergent (
<figureCitation id="13333EBC092DFF9A812FCE9EFDE3FEC2" box="[526,597,247,268]" captionStart="Figs" captionStartId="2.[197,236,1466,1485]" captionTargetBox="[325,1043,194,1444]" captionTargetId="figure-0@2.[322,1044,194,1451]" captionTargetPageId="2" captionText="Figs. 1–32. Male genitalia of Desmopachria species in the D. nitida species-group. D. curseenae: Median lobe – 1) Right lateral aspect; 2) Ventral aspect; Right lateral lobe – 3) Right lateral aspect; 4) Ventral aspect. D. delongi: Median lobe – 5) Right lateral aspect; 6) Ventral aspect; Right lateral lobe – 7) Right lateral aspect; 8) Ventral aspect. D. gingerae: Median lobe – 9) Right lateral aspect; 10) Ventral aspect; 11) Right lateral lobe, right lateral aspect. D. gyrationi: Median lobe – 12) Right lateral aspect; 13) Ventral aspect; 14) Right lateral lobe, right lateral aspect. D. hardyae: Median lobe – 15) Right lateral aspect; 16) Ventral aspect; 17) Right lateral lobe, right lateral aspect. D. kemptonae: Median lobe – 18) Right lateral aspect; 19) Ventral aspect; Right lateral lobe – 20) Right lateral aspect; 21) Ventral aspect. D. lloydi: 22) Median lobe, ventral aspect; 23) Apices of median lobe, anterior aspect; Right lateral lobe – 24) Right lateral aspect; 25) Ventral aspect. D. singhae: Median lobe – 26) Right lateral aspect; 27) Ventral aspect; 28) Right lateral lobe, right lateral aspect. D. vohrae: Median lobe – 29) Right lateral aspect; 30) Ventral aspect; Right lateral lobe – 31) Right lateral aspect; 32) Ventral aspect." figureDoi="http://doi.org/10.5281/zenodo.4836705" httpUri="https://zenodo.org/record/4836705/files/figure.png" pageId="9" pageNumber="106">Fig. 25</figureCitation>
).
</paragraph>
<paragraph id="8BB72239092DFF9A838ECF78FED9FE95" blockId="9.[151,639,194,826]" pageId="9" pageNumber="106">
<emphasis id="B97CFE2B092DFF9A838ECF78FEA3FEE8" bold="true" box="[175,277,273,294]" pageId="9" pageNumber="106">Variation.</emphasis>
Most of the specimens examined are teneral, and there is variation in intensity of the coloration of surfaces.
</paragraph>
</subSubSection>
<subSubSection id="C31271B2092DFF9A838ECF08FE0AFE5F" pageId="9" pageNumber="106" type="etymology">
<paragraph id="8BB72239092DFF9A838ECF08FE0AFE5F" blockId="9.[151,639,194,826]" pageId="9" pageNumber="106">
<emphasis id="B97CFE2B092DFF9A838ECF08FE94FEB8" bold="true" box="[175,290,353,374]" pageId="9" pageNumber="106">Etymology.</emphasis>
This species is named
<taxonomicName id="4C0859BA092DFF9A82D4CF09FD9DFEB8" authorityName="Miller and Wolfe" authorityYear="2018" box="[501,555,352,374]" class="Insecta" family="Dytiscidae" genus="Desmopachria" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="106" phylum="Arthropoda" rank="species" species="lloydi">
<emphasis id="B97CFE2B092DFF9A82D4CF09FD9DFEB8" box="[501,555,352,374]" italics="true" pageId="9" pageNumber="106">lloydi</emphasis>
</taxonomicName>
in honor of Dr. Michael S. Lloyd, MD.
</paragraph>
</subSubSection>
<subSubSection id="C31271B2092DFF9A838ECFFFFDE9FE2E" pageId="9" pageNumber="106" type="distribution">
<paragraph id="8BB72239092DFF9A838ECFFFFDC8FE08" blockId="9.[151,639,194,826]" pageId="9" pageNumber="106">
<emphasis id="B97CFE2B092DFF9A838ECFFFFE86FE65" bold="true" box="[175,304,406,427]" pageId="9" pageNumber="106">Distribution.</emphasis>
The species is known only from the
<typeStatus id="54B39C9B092DFF9A83B6CFD8FF76FE08" box="[151,192,433,454]" pageId="9" pageNumber="106">type</typeStatus>
locality in northern lowland
<collectingCountry id="F31F62A9092DFF9A82F9CFD8FDA8FE08" box="[472,542,433,454]" name="Bolivia" pageId="9" pageNumber="106">Bolivia</collectingCountry>
(
<figureCitation id="13333EBC092DFF9A810ACFD8FDC5FE08" box="[555,627,433,454]" captionStart="Figs" captionStartId="13.[175,214,1279,1298]" captionTargetBox="[151,1173,194,1263]" captionTargetId="figure-175@13.[151,1173,194,1263]" captionTargetPageId="13" captionText="Figs. 71–72. Distribution of Desmopachria species in the D. nitida species-group. 71) D. curseenae, D. singhae, and D. vohrae; 72) D. lloydi." figureDoi="http://doi.org/10.5281/zenodo.4836725" httpUri="https://zenodo.org/record/4836725/files/figure.png" pageId="9" pageNumber="106">Fig. 72</figureCitation>
).
</paragraph>
<paragraph id="8BB72239092DFF9A838ECFA2FDE9FE2E" blockId="9.[151,639,194,826]" box="[175,607,459,480]" pageId="9" pageNumber="106">
<emphasis id="B97CFE2B092DFF9A838ECFA2FEB5FE2E" bold="true" box="[175,259,459,480]" pageId="9" pageNumber="106">Habitat.</emphasis>
Specimens were collected at lights.
</paragraph>
</subSubSection>
<subSubSection id="C31271B2092DFF9A838ECF8FFE37FCF4" pageId="9" pageNumber="106" type="materials_examined">
<paragraph id="8BB72239092DFF9A838ECF8FFE37FCF4" blockId="9.[151,639,194,826]" pageId="9" pageNumber="106">
<emphasis id="B97CFE2B092DFF9A838ECF8FFEFBFE35" bold="true" box="[175,333,486,507]" pageId="9" pageNumber="106">Type Material.</emphasis>
<materialsCitation id="3B602864092DFF9A827ACF8FFE08FD4E" collectingDate="1988-08-06" collectingDateMax="1988-08-09" collectingDateMin="1988-08-06" collectionCode="KUNHM" collectorName="Robert W. Brooks" country="Bolivia" location="El" pageId="9" pageNumber="106" specimenCount="2" stateProvince="Beni" typeStatus="holotype">
<typeStatus id="54B39C9B092DFF9A827ACF8FFE05FE35" box="[347,435,486,507]" pageId="9" pageNumber="106" type="holotype">Holotype</typeStatus>
in
<collectionCode id="ED19BAFC092DFF9A82C3CF8FFD88FE35" box="[482,574,486,507]" pageId="9" pageNumber="106">KUNHM</collectionCode>
, male labeled, “
<collectingCountry id="F31F62A9092DFF9A83D0CC69FEE0FDDB" box="[241,342,512,533]" name="Bolivia" pageId="9" pageNumber="106">BOLIVIA</collectingCountry>
:
<location id="8ED774E2092DFF9A827FCC69FEC5FDDB" LSID="urn:lsid:plazi:treatment:03A1932F092CFF9A81D1CD36FE37FCF4:8ED774E2092DFF9A827FCC69FEC5FDDB" box="[350,371,512,533]" country="Bolivia" name="El" pageId="9" pageNumber="106" stateProvince="Beni">El</location>
<collectingRegion id="49CCECDB092DFF9A8259CC69FE13FDDB" box="[376,421,512,533]" country="Bolivia" name="Beni" pageId="9" pageNumber="106">Beni</collectingRegion>
El Porvenir Stn.
<locationDeviation id="9A053B1B092DFF9A8164CC69FF4AFDFE" location="San Borja" pageId="9" pageNumber="106">NE of San Borja</locationDeviation>
<date id="FFB604F9092DFF9A8228CC72FE00FDFE" box="[265,438,539,560]" pageId="9" pageNumber="106" value="1988-08-06" valueMax="1988-08-09" valueMin="1988-08-06">
<collectingDate id="EFF2FD11092DFF9A8228CC72FE00FDFE" box="[265,438,539,560]" pageId="9" pageNumber="106" value="1988-08-06" valueMax="1988-08-09" valueMin="1988-08-06">6-9 August 1988</collectingDate>
</date>
/ BIOLAT SI/MAB
<collectorName id="26FD47EF092DFF9A83B6CC5CFEE0FD84" box="[151,342,565,586]" pageId="9" pageNumber="106">Robert W. Brooks</collectorName>
ex., at lights/
<typeStatus id="54B39C9B092DFF9A8121CC5CFDCBFD84" box="[512,637,565,586]" pageId="9" pageNumber="106" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C0859BA092DFF9A83B6CC26FDF9FDAB" authority="Miller and Wolfe, 2017" authorityName="Miller and Wolfe" authorityYear="2017" box="[151,591,591,613]" class="Insecta" family="Dytiscidae" genus="Desmopachria" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="106" phylum="Arthropoda" rank="species" species="lloydi">
<emphasis id="B97CFE2B092DFF9A83B6CC26FED4FDAB" box="[151,354,591,613]" italics="true" pageId="9" pageNumber="106">Desmopachria lloydi</emphasis>
Miller and Wolfe, 2017
</taxonomicName>
[red label with black line border].”
</materialsCitation>
<materialsCitation id="3B602864092DFF9A82E6CC02FECBFCF4" collectingDate="1988-07-31" collectorName="Robert W. Brooks" country="Bolivia" county="Five" location="Palm Camp" municipality="El" pageId="9" pageNumber="106" specimenCount="4" stateProvince="Beni Beni" typeStatus="paratype">
<typeStatus id="54B39C9B092DFF9A82E6CC02FD90FD4E" box="[455,550,619,640]" pageId="9" pageNumber="106" type="paratype">Paratypes</typeStatus>
:
<collectingCounty id="62D65AB5092DFF9A8113CC02FDEAFD4E" box="[562,604,619,640]" pageId="9" pageNumber="106">Five</collectingCounty>
labeled same as
<typeStatus id="54B39C9B092DFF9A8272CCECFE10FD54" box="[339,422,645,666]" pageId="9" pageNumber="106" type="holotype">holotype</typeStatus>
except each with “…
<typeStatus id="54B39C9B092DFF9A839BCCC9FE86FD7B" box="[186,304,672,693]" pageId="9" pageNumber="106" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4C0859BA092DFF9A821DCCF6FEA6FD01" authority="Miller and Wolfe, 2017" authorityName="Miller and Wolfe" authorityYear="2017" class="Insecta" family="Dytiscidae" genus="Desmopachria" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="106" phylum="Arthropoda" rank="species" species="lloydi">
<emphasis id="B97CFE2B092DFF9A821DCCF6FDBFFD7B" box="[316,521,671,693]" italics="true" pageId="9" pageNumber="106">Desmopachria lloydi</emphasis>
Miller and Wolfe, 2017
</taxonomicName>
[blue label with black line border];” two labeled, “
<collectingCountry id="F31F62A9092DFF9A8200CCBCFE30FD24" box="[289,390,725,746]" name="Bolivia" pageId="9" pageNumber="106">BOLIVIA</collectingCountry>
:
<collectingMunicipality id="6BD3B843092DFF9A82B0CCBCFE10FD24" box="[401,422,725,746]" pageId="9" pageNumber="106">El</collectingMunicipality>
<collectingRegion id="49CCECDB092DFF9A828FCCBCFDA7FD24" box="[430,529,725,746]" country="Bolivia" isEnumeration="true" name="Beni" pageId="9" pageNumber="106">Beni Beni</collectingRegion>
Stn.,
<location id="8ED774E2092DFF9A816CCCBCFF64FCCB" LSID="urn:lsid:plazi:treatment:03A1932F092CFF9A81D1CD36FE37FCF4:8ED774E2092DFF9A816CCCBCFF64FCCB" country="Bolivia" county="Five" municipality="El" name="Palm Camp" pageId="9" pageNumber="106" stateProvince="Beni Beni">Palm Camp</location>
,
<locationDeviation id="9A053B1B092DFF9A83FFCC99FE35FCCB" box="[222,387,752,773]" location="San Borja" pageId="9" pageNumber="106">NE of San Borja</locationDeviation>
,
<date id="FFB604F9092DFF9A82AFCC99FDBAFCCB" box="[398,524,752,773]" pageId="9" pageNumber="106" value="1988-07-31">
<collectingDate id="EFF2FD11092DFF9A82AFCC99FDBAFCCB" box="[398,524,752,773]" pageId="9" pageNumber="106" value="1988-07-31">31 July 1988</collectingDate>
</date>
/
<collectorName id="26FD47EF092DFF9A813ACC99FF6AFCD1" pageId="9" pageNumber="106">Robert W. Brooks</collectorName>
ex., at lights BIOLAT-SI/MAB/…” and
<typeStatus id="54B39C9B092DFF9A83B6CD4CFF5EFCF4" box="[151,232,805,826]" pageId="9" pageNumber="106" type="paratype">paratype</typeStatus>
label as above
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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